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Perennial Ryegrass - USDA Plants Database

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SNAKE RIVER<br />

WHEATGRASS<br />

Elymus wawawaiensis<br />

J. Carlson & M. Barkworth<br />

Plant Symbol = ELWA2<br />

Contributed by: <strong>USDA</strong> NRCS Idaho State Office<br />

Snake River Wheatgrass. Dr. Tom Jones, <strong>USDA</strong>, ARS, Logan, UT<br />

Alternate Names<br />

Formerly referred to as bluebunch wheatgrass before the<br />

description of Snake River wheatgrass as a new species.<br />

Uses<br />

Grazing/rangeland/wildlife: Snake River wheatgrass is<br />

palatable to all classes of livestock and wildlife. It is a<br />

preferred feed for cattle, horses, sheep, elk, deer and<br />

antelope in spring when protein levels can be as high as<br />

20 percent. Protein levels decrease to about 4 percent as<br />

the grass matures and cures by early to mid summer.<br />

Digestible carbohydrates remain about 45 percent<br />

throughout the active growth period.<br />

Erosion control/reclamation: Snake River wheatgrass is<br />

very drought tolerant, stands are persistent once<br />

established, and it is adapted to stabilization of disturbed<br />

soils. It is very compatible with slower developing<br />

natives such as streambank wheatgrass, thickspike<br />

wheatgrass, basin wildrye, western wheatgrass, bluegrass<br />

Plant Guide<br />

species, and needlegrass species. It does not compete<br />

well with aggressive introduced grasses. Its drought<br />

tolerance, combined with an extensive root system and<br />

fair to good seedling vigor, make this species ideal for<br />

reclamation in areas receiving 10 to 20 inches annual<br />

precipitation. The cultivars ‘Discovery’ and 'Secar'<br />

compete well in areas as low as 8 inches annual rainfall.<br />

This grass can be used in urban areas where irrigation<br />

water is limited to stabilize ditch banks, dikes, and<br />

roadsides as a component of a native mix.<br />

Description<br />

General: The natural distribution of Snake River<br />

wheatgrass is limited to eastern Oregon, eastern<br />

Washington, and west central to northern Idaho, but it is<br />

widely used as a surrogate for bluebunch wheatgrass for<br />

restoration, reclamation, and rehabilitation of rangelands<br />

in the Columbia Plateau, Northern Basin and Range,<br />

Central Basin and Range, and Snake River Plain, which<br />

encompass large portions of eastern Washington, eastern<br />

Oregon, southern Idaho, Nevada, and western Utah.<br />

Snake River wheatgrass is a long-lived, perennial, coolseason<br />

native bunchgrass with an extensive root system<br />

with strong tillers. Snake River wheatgrass spreads by<br />

seed.<br />

Snake River wheatgrass can be morphologically<br />

distinguished from bluebunch wheatgrass by characters<br />

such as the ratio of spikelet length to internode length,<br />

glume shape, seedling pubescence, and seedling leaf<br />

marginal barbs (Jones et al., 1991).<br />

Snake River (left) vs. awned (center) and awnless( right) bluebunch<br />

wheatgrasses<br />

Snake River wheatgrass has a seed of considerably<br />

smaller mass than bluebunch wheatgrass. Snake River


wheatgrass is always awned, while bluebunch wheatgrass<br />

may be awned or awnless. Snake River wheatgrass is<br />

always allotetraploid (2n=28) while bluebunch<br />

wheatgrass may be diploid (2n=14) or autotetraploid<br />

(2n=28) (Carlson and Barkworth, 1997).<br />

Adaptation and Distribution<br />

Snake River wheatgrass does best on medium to coarsetextured<br />

soils, but can be found on heavy to medium to<br />

coarse-textured soils over 10 inches deep, including fairly<br />

sandy sites. It can be seeded on clayey sites. It may be<br />

found on thin, rocky sites and on very steep slopes. It<br />

tolerates weakly saline conditions. It does not grow on<br />

acidic sites. It is cold tolerant, moderately shade tolerant,<br />

and very tolerant of fire. It is intolerant of high water<br />

tables, poor drainage, and periods of extended inundation.<br />

On native sites Snake River wheatgrass is most abundant<br />

in the 8 to 20 inch annual precipitation zones. When<br />

seeded, it does best with 10 to 20 inches of precipitation<br />

(‘Discovery’ and 'Secar' have established with as little as<br />

7 inches) and it will tolerate up to 35 inches on very welldrained<br />

soils. The elevation range is from 500 feet above<br />

sea level to 10,000 feet. It is a major component of native<br />

plant communities where it is found and generally<br />

occupies 20 to 60 percent of the overall composition by<br />

weight of the plant community.<br />

For a current distribution map, consult the Plant Profile<br />

page for this species on the PLANTS Website.<br />

Status<br />

Consult the PLANTS Website and your State Department<br />

of Natural Resources for this plant’s current status (e.g.<br />

threatened or endangered species, state noxious status,<br />

and wetland indicator values).<br />

Establishment<br />

This species should be seeded with a drill at a depth of 1/2<br />

inch or less on medium-textured soils, 1/4 inch on finetextured<br />

soils and 3/4 inch or less on coarse-textured soils.<br />

The single species seeding rate recommended for this<br />

grass is 8 pounds Pure Live Seed (PLS) or about 24 PLS<br />

per square foot. If used as a component of a mix, adjust<br />

to percent of mix desired. For broadcast plantings, mined<br />

lands, and other harsh critical areas, the seeding rate<br />

should be increased to 40 to 50 PLS per square foot.<br />

Seedlings are significantly weaker than crested<br />

wheatgrass and a clean firm weed free seedbed is required<br />

for establishment (cheatgrass and other weed competition<br />

can cause stand failure).<br />

Seed in very early spring on heavy to medium- textured<br />

soils and in late fall on medium to light- textured soils.<br />

Late summer (August to mid- September) seeding is not<br />

recommended unless irrigation is available. Dormant fall<br />

seedings will pre-chill seed and improve germination.<br />

Snake River wheatgrass establishes fairly quickly for a<br />

native grass and stands should be given up to 3 years to<br />

ensure establishment. It is compatible with other native<br />

species and should be used in seeding mixtures. It should<br />

not be seeded with strongly competitive introduced<br />

species.<br />

Stands may require weed-control measures during<br />

establishment. Application of 2,4-D should not be made<br />

until plants have reached the four to six-leaf stage. Mow<br />

weeds at or prior to their bloom stage. New stands may be<br />

damaged by grasshoppers and other insects and pesticides<br />

may be needed.<br />

Management<br />

Stands of Snake River wheatgrass should not be grazed<br />

until they have firmly established (usually two to three<br />

growing seasons) and started to flower.<br />

Six inches of new growth should be attained in spring<br />

before grazing is allowed in established stands. The<br />

growing point of Snake River wheatgrass is fairly high<br />

and stands can be easily overgrazed. Spring grazing<br />

should occur no more than one out of three years and no<br />

more than 40 percent utilization should occur during rapid<br />

growth. Heavy early-spring grazing is especially<br />

damaging, and grazing should be delayed until flowers<br />

are halfway emerging from the protecting leaf. No more<br />

than 60 percent utilization should occur after seed ripens.<br />

Snake River wheatgrass is not suited to hay production<br />

and is best suited to rangeland, critical area stabilization,<br />

and cropland retirement type uses.<br />

Environmental Concerns<br />

Snake River wheatgrass is long lived and spreads<br />

primarily via seed distribution. It is not considered a<br />

"weedy" or invasive species, but can spread into adjoining<br />

vegetative communities under ideal climatic and<br />

environmental conditions. Most seedings do not spread<br />

from original plantings.<br />

It is a cross-pollinating species and is known to cross with<br />

bluebunch wheatgrass, thickspike wheatgrass, quackgrass,<br />

and bottlebrush squirreltail. These interspecific hybrids<br />

generally exhibit low fertility and do not dominate a site<br />

or crowd out non-hybrid plants.<br />

Seed Production<br />

Seed production of Snake River wheatgrass has been very<br />

successful under cultivated conditions. Row spacing of<br />

24 to 36 inches are recommended under irrigation and 36<br />

inches under dryland conditions. Seeding rates of 3 to 4<br />

PLS per acre are recommended. Cultivation and/or<br />

judicious use of appropriate herbicides will be needed to<br />

maintain rows and weed-free conditions.<br />

Seed fields are productive for three to four years.<br />

Average production of 75 to 100 pounds per acre can be<br />

expected under dryland conditions. Average production


of 250-500 pounds per acre can be expected under<br />

irrigated conditions.<br />

A Snake River wheatgrass seed production field<br />

Dr. Tom Jones, <strong>USDA</strong>, ARS, Logan, UT<br />

Harvesting is best completed by swathing, followed by<br />

combining of the cured rows. The seed heads readily<br />

shatter and require close scrutiny of maturing stands. If<br />

direct combined, harvest at 30 percent moisture and dry<br />

seed down to 12 percent moisture (if stored in bins) or 15<br />

percent moisture (if stored in sacks).<br />

Debearding will be required during processing. Seed is<br />

generally harvested from mid-July to mid-August.<br />

Cultivars, Improved, and Selected Materials (and area<br />

of origin)<br />

‘Discovery’, the second Snake River wheatgrass (Elymus<br />

wawawaiensis) plant material, was released in 2008<br />

(Jones, 2008). It was developed from four accessions that<br />

were found to have improved vigor in comparison to<br />

Secar. These accessions were crossed and seed was<br />

harvested in bulk to generate Discovery.<br />

Discovery was compared to Secar in seeded trials at two<br />

sites in the Intermountain Region and four sites in the<br />

Great Plains (Jones, 2008). Stand frequency of Discovery<br />

in the Intermountain locations for year 2 was 49.3%<br />

compared to 27.4% for Secar. Forage yield of Discovery<br />

for years 2 to 4 was 108.0% greater than Secar. No<br />

significant differences were observed at the Great Plains<br />

locations. Discovery was also compared to Secar in a<br />

transplanted trial at Millville, Utah where it produced<br />

18% greater straw weight across two densities the<br />

summer after establishment.<br />

The populations used to generate Discovery trace to<br />

materials collected in Whitman and Asotin counties in<br />

southeastern Washington and Idaho County in central<br />

Idaho.<br />

Breeder seed is maintained by the <strong>USDA</strong>-ARS Forage<br />

and Range Research Laboratory, Logan, UT, and will be<br />

made available to commercial growers through the Utah<br />

Crop Improvement Association for production of<br />

Foundation, Registered, and Certified generations.<br />

'Secar' is the original cultivar release of Snake River<br />

wheatgrass (Elymus wawawaiensis). It was originally<br />

released as a bluebunch wheatgrass, but cytological<br />

examination later determined it to be Snake River<br />

wheatgrass. The original collection site for Secar is along<br />

the Snake River Gorge near Lewiston, Idaho. The Idaho-<br />

Oregon- Montana-Wyoming AES, Washington<br />

Agriculture Research Center, and Pullman Plant Materials<br />

Center released Secar in 1980.<br />

It is one of the most drought-tolerant native perennial<br />

grasses available and can survive down to 8 inches<br />

rainfall. It is a bunchgrass with fair to good seedling vigor<br />

and establishes well under droughty conditions. While<br />

Secar is considered to be highly drought tolerant as a<br />

mature plant, drought often reduces the stand during the<br />

establishment year. 'Secar' is intended for use on<br />

rangeland for re-establishment of native-plant<br />

communities and for cropland retirement type plantings.<br />

Certified seed is available and Breeder seed is maintained<br />

by Pullman PMC. Foundation seed is available through<br />

the Washington State Crop Improvement Association.<br />

References<br />

Alderson, J. & W.C. Sharp 1994. Grass varieties in the<br />

United States. Agriculture Handbook No. 170.<br />

<strong>USDA</strong>, SCS, Washington, D.C.<br />

Carlson, J.R. & M.E. Barkworth 1997. Elymus<br />

wawawaiensis: a species hitherto confused with<br />

Pseudoroegneria spicata (Triticeae, Poaceae).<br />

Phytologia 83:312-330.<br />

Cronquist, A., A.H. Holmgren, N.H. Holmgren, J.L.<br />

Reveal, & P. K. Holmgren 1977. Intermountain<br />

Flora. Vol. 6. The New York Botanical Garden.<br />

Columbia University Press, New York, New York.<br />

Hitchcock, A. S. 1950. Manual of the grasses of the<br />

United States. <strong>USDA</strong>, Washington, DC.<br />

Jones, T.A. 2008. Notice of release of ‘Discovery’ Snake<br />

River wheatgrass. Native <strong>Plants</strong> J. 9:99-102.<br />

Jones, T.A., D.C. Nielson, and J.R. Carlson. 1991.<br />

Developing a grazing-tolerant native grass for<br />

bluebunch wheatgrass sites. Rangelands 13:147-150.<br />

Miller, R.P., J.M. Seufert, and M.R. Haferkamp. 1987.<br />

The ecology and management of bluebunch<br />

wheatgrass (Agropyron spicatum): A review. Oregon<br />

Agric. Exp. Sta. Bull. No. 669.


Morrison, K.J., and C.A. Kelley. 1981. Secar bluebunch<br />

wheatgrass. EB 0991. Cooperative Extension,<br />

Washington State University, Pullman.<br />

<strong>USDA</strong>, Forest Service 1996. Fire effects information<br />

system. Version: 000413. Rocky Mountain Research<br />

Station, Fire Sciences Laboratory, Missoula, MT.<br />

http://www.fs.fed.us/database/feis/<br />

<strong>USDA</strong>, NRCS 2000. The PLANTS database.<br />

http://plants.usda.gov National Plant Data Center,<br />

Baton Rouge, Louisiana.<br />

Prepared By<br />

Daniel G. Ogle<br />

<strong>USDA</strong>, NRCS, Idaho State Office, Boise, Idaho<br />

Mark Stannard<br />

<strong>USDA</strong>, NRCS, Plant Materials Center, Pullman,<br />

Washington<br />

Dr. Thomas A. Jones<br />

<strong>USDA</strong>, ARS, Forage and Range Laboratory, Logan, Utah<br />

Species Coordinator<br />

Daniel G. Ogle<br />

<strong>USDA</strong>, NRCS, Idaho State Office, Boise, Idaho<br />

Published September 2008<br />

Edited: 091608dgo; 091508ms; 071108taj<br />

For more information about this and other plants, please<br />

contact your local NRCS field office or Conservation<br />

District at http://www.nrcs.usda.gov/ and visit the<br />

PLANTS Web site at http://plants.usda.gov/ or the Plant<br />

Materials Program Web site http://plantmaterials.nrcs.usda.gov.<br />

PLANTS is not responsible for the content or availability<br />

of other Web sites.<br />

<strong>USDA</strong> IS AN EQUAL OPPORTUNITY PROVIDER AND EMPLOYER

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