August 2004 - FMCS-Freshwater Mollusk Conservation

Submissions for the December 2004 issue of Ellipsaria may be sent in at any time but are due by Nov. 1, 2004. 

Anyone may submit an article but you must be a member of FMCS to receive Ellipsaria. Categories for contributions 

include news, new publications, meeting announcements, current issues affecting mollusks, job postings, contributed 

articles (including ongoing research projects), abstracts, and society committee reports. Electronic submissions are 

preferred; please send submissions to the editor. 

Submissions to Ellipsaria are not peer reviewed, but are checked for content and general editing. 

Please send change of address information to the Secretary, Rita Villella.

FMCS Reports: 

2005 FMCS Symposium ............................. 2 

Genetics Workshop ..................................... 3 

June Board Meeting .................................... 4 

Committee Reports...................................... 5 

FMCS Awards............................................. 6 

Publications......................................................... 9 

Contributed Articles.......................................... 10 

2003 Mollusk Bibliography .............................. 22 

2004 Membership List ...................................... 47 

http://ellipse.inhs.uiuc.edu/FMCS/

FMCS Reports 

2005 FMCS Symposium Announced! 

St. Paul Minnesota, here we come! Plans are set for the 4th 

FMCS Symposium on May 15-19, 2005 at the Radisson 

Riverfront Hotel. We return to the Midwest in spring and 

join our Mississippi River brethren to rekindle old 

friendships, establish new relationships, and spend 3 days 

talking shell. Our theme will center on exotics and threats to 

the fauna but will be inclusive enough to draw papers in 

from all parts of the mussel spectrum. Indeed – the first call 

for papers is in this very edition of Ellipsaria…imagine that! 

WE NEED YOUR HELP! Consider how you can lend a 

hand and contact the committee chair of your choice…but 

do it! Many hands make light work and this is a great 

opportunity to make a significant contribution to the Society. 

An organizational framework with committee chairs has 

been established and is posted on the FMCS web site. Or, 

contact the general chair, Kurt Welke @ 608-273-5946 and 

we’ll find a home for your talents. 

More information including a call for papers follows in the 

next article. Other details will be included in the December 

newsletter, member mailings, and postings on the Unio 

server. 

Fire Up! St. Paul is a great town with exceptional resources 

only a stone’s throw away. This promises to be an 

outstanding meeting so grease your skids now and plan on 

Minnesota in May! 

Submitted by Kurt Welke, Symposium Chair 

FMCS 2005 Symposium 

May 15 - 19, 2005 

Radisson Riverfront Hotel 

St. Paul, Minnesota 

Minnesota is pleased to host the 2005 Symposium of the 

Freshwater Mollusk Conservation Society. The symposium 

will be held at the Radisson Riverfront Hotel in St. Paul, 

Minnesota. As time allows, we welcome you to explore the 

area’s rich natural resource, cultural, and culinary delights. 

The 2005 meeting will be co-hosted by the Upper 

Mississippi River Conservation Committee (UMRCC). 

Members of the UMRCC’s ad hoc mussel team are working 

on FMCS committees to bring a strong local flavor to the 

discussion and information exchange. As many as three 

field trips to sample local river mussel communities and to 

see the results of endangered species recovery efforts are 

being planned. 

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Theme 

“Are Your Natives Restless? Holistic 

Strategies for Conserving Freshwater 

Mollusks during Exotic Species Invasions” 

Rationale 

The invasion of exotic species into the freshwaters of North 

America and elsewhere in the world has placed additional 

stress on an already imperiled freshwater mollusk fauna. 

Vital ecosystems for preserving freshwater mollusks like the 

Mississippi, Ohio, Tennessee, and other large rivers 

unfortunately also serve as conduits for the rapid dispersal 

and colonization of exotic species, where they often have 

adverse effects on native species. Scientists from a variety 

of disciplines including ecology, fisheries biology, 

malacology, conservation biology, engineering, hydrology, 

and others are working together to help mitigate the potential 

spread and negative effects of exotic species. These folks 

need your help! The 2005 symposium will provide an 

opportunity for biologists and others to share ideas, 

exchange information, and engage in the interpersonal 

networking needed to support the conservation of our native 

freshwater molluscan resources. 

Think critically about your own research, and its association 

with other research projects, and its implications relative to 

the following interrelated topics: 

Native Species Relocation 

Artificial and Natural Refugia 

Native Species-Exotic Species Interactions 

Population Biology 

Alteration of Ecological Conditions 

Influence on Nutrient and Food Resources 

Parasites and Contaminant Flux 

...and other topics related to exotic species interactions with 

native species. 

Instructions for Authors – Call for Abstracts 

We are requesting abstracts associated with the above theme 

and others associated with the following topics: Habitat and 

Conservation, Range-wide Status and Distribution (coauthorship 

is encouraged), Life History and Ecology, 

Evolution and Phylogenetics, Outreach and Education, 

Propagation and Reproduction, Recovery, and Contaminants 

and Water Quality. 

ABSTRACTS MUST BE RECEIVED BY 

DECEMBER 17, 2004 

Instructions for Abstract Preparation and Submittal 

Submittal form: Abstracts should be submitted as an email 

attachment in Microsoft Word® or Rich Text format to Greg 

Cope (greg_cope@ncsu.edu). File name should include 

presenter’s last name and initials (e.g., jonesjm.doc). 

Acknowledgment of abstract receipt, if requested, will be 

provided by e-mail.

Limit abstracts to 300 words or less (including title, authors 

and affiliations). Abstracts with greater than 300 words will 

be edited. 

Submittal format: The abstract should contain the title in 

BOLD, CAPITAL letters, followed by the author(s), and 

address(es). Underscore the presenter's name. Skip one line 

and begin the text (see example below) including a clear 

summary of presentation including objectives, results, and 

conclusions. 

PROPAGATION OF FRESHWATER MUSSELS IN 

A CLOSED RECIRCULATING SYSTEM, Shane D. 

Hanlon 1 , Jay F. Levine 2 , Lori Gustafson 2 , and Chris 

Eads 2 . 1 U.S. Fish and Wildlife Service, Southwestern 

Virginia Field Office, 330 Cummings Street, Abingdon, 

VA 24210; 2 College of Veterinary Medicine, North 

Carolina State University, Raleigh, NC 27606. 

We renovated a preexisting wet laboratory located at 

North Carolina State University for purposes of 

propagating . . . 

At the bottom of the page, type: 

1. The name, address, telephone, fax, and e-mail of the 

presenting author 

2. Preference for Platform or Poster presentation and 

willingness (yes or no) to convert from one format to 

another 

3. Regular or Student* attendee 

*Note: All students submitting abstracts, provided they 

meet eligibility requirements (see accompanying newsletter 

article), will be judged for the best student platform or poster 

presentation, unless otherwise indicated. 

ABSTRACTS MUST BE RECEIVED BY 

DECEMBER 17, 2004 

Oral Presentation Requirements 

Not to exceed 20 minutes (15 minutes for talk and 5 minutes 

for questions and answers). Slides and LCD projector visual 

aids only (no overheads). 

Poster requirements 

The poster should be readable from 5 feet, titles from 10 

feet; and the poster should not exceed a size of 4 feet high by 

8 feet wide. Authors must be present at the designated 

poster session. 

Accommodations 

The Radisson Riverfront Hotel rate is $100 per night, single 

or double occupancy. Rooms must be booked by April 15, 

2005 to guarantee this rate. Please call 651-292-1900 or 

800-333-3333 for reservations. Check-in time is 4 p.m., and 

check-out time is 12 noon. The Radisson Riverfront is 

located along the Mississippi River on Kellogg Boulevard in 

downtown St. Paul, Minnesota. 

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*** Please plan to book your room with the Radisson 

Riverfront. FMCS meeting room rental costs are based 

upon the number of hotel rooms booked by members. We 

need to book at least 100 rooms each night to ensure 

reasonable meeting room costs. Also, please remember to 

tell the hotel receptionist you are attending the FMCS 

symposium when making your reservations so the society 

will receive credit for the rooms reserved. *** 

Travel 

The Radisson Riverfront Hotel does not provide 

complementary shuttle service from the airport. However, 

the SuperShuttle will provide transportation to and from the 

Radisson Riverfront for $12 one way or $22 round trip. Call 

612-827-7777 for reservations. The hotel parking ramp is 

$12.50/day for registered hotel guests. 

Submitted by Greg Cope, Symposium Program Chair 

Workshop Wrap-Up: Conservation 

Genetics Workshop on Imperiled 

Freshwater Mollusks and Fishes 

The Conservation Genetics Workshop on Imperiled 

Freshwater Mollusks and Fishes was a success! This twoday 

event sponsored by the FMCS, USFWS, and 

Department of Fisheries and Wildlife Sciences, Virginia 

Polytechnic Institute and State University, was held at the 

National Conservation Training Center, Shepherdstown, 

West Virginia in June 2004. The workshop provided 

resource managers and biologists with an opportunity to 

learn about conservation genetics issues as applied to 

recovery of imperiled freshwater mollusks and fishes. The 

workshop included 22 platform presentations and 17 poster 

presentations. Nationally recognized experts spoke on 

topics concerning quantitative genetics, molecular genetics, 

phylogenetics, species concepts, taxonomic analysis, cryptic 

species, hybridization, and genetic management guidelines 

for captive propagation and releases of endangered species. 

The evening poster presentation session provided 

participants with an opportunity to learn about ongoing 

genetics projects and interact with people from around the 

country in a festive and informal setting. The workshop was 

successful at bringing together conservation professionals 

with different backgrounds to share their expertise with 

natural resource managers. Biologists from National Marine 

Fisheries Service in Seattle, Washington, shared their 

experiences with conservation genetics of Pacific salmon, 

many of which directly parallel our own experiences and 

challenges with trying to protect freshwater mollusks and 

non-game fishes in other parts of the country. The program 

and abstracts can be viewed and downloaded from the 

FMCS website at http://ellipse.inhs.uiuc.edu/FMCS/ 

Additionally, the program abstracts likely will be published 

in an upcoming issue of Walkerana, the Journal of the 

Freshwater Mollusk Conservation Society. 

Approximately 110 people attended the workshop, with 

more than 30 of these registrants becoming new FMCS

members. Extra programs are available ($20.00) to those 

interested in having a bound original copy, as are t-shirts 

($15.00). The t-shirts are white and have a black and white 

event logo on the front with a small fish, and a color logo on 

the back with various fish and mussels contained in an 

underwater scene; sponsor logos also are the back. Both the 

program and t-shirts can be obtained by contacting Jess 

Jones or Dick Neves at Virginia Tech. Revenues for the 

workshop were approximately $15,000.00; once all revenues 

and expenses have been tabulated, an expense report will be 

submitted to the FMCS Treasurer for inclusion in a 

subsequent issue of Ellipsaria. 

The Organizing Committee would like to thank the staff at 

the National Conservation Training Center for coordinating 

local arrangements and making the event go smoothly. 

Robert Butler was the USFWS representative for the 

workshop, which allowed the society to receive discounts on 

facilities charges. A special thank you goes to all of the 

speakers and poster presenters, who traveled from all parts 

of the country to share their work and made the workshop 

interesting and informative. The FMCS continues to be at 

the forefront of emerging issues and promote science-based 

conservation of species, streams, and rivers. The society 

also continues to extend its hand to other institutions, 

societies, and aquatic biologists in an effort to gain new 

members and to work toward common goals to protect 

mollusks, fishes and their habitats. 

Submitted by Jess Jones 

FMCS Treasurer Election Results 

Heidi Dunn was unanimously re-elected as FMCS treasurer 

in April 2004. She will serve for 2 years. We owe Heidi 

many thanks for the work she has done and will continue to 

do for FMCS. 

Leroy Koch, Nominations Committee 

FMCS Board Meeting 

National Conservation Training Center 

Shepherdstown, West Virginia 

June 28, 2004 

Treasurer Report 

The snail workshop generated an income of $9165 

(including a $500 donation) and expenses were $8827.92. 

Jess Jones reported that the genetics workshop has generated 

an income of about $14,000 as of June 28 th , with expenses 

running about $7,000.00. A commitment was received from 

the Fish and Wildlife Dept at Virginia Tech for a $1000 

donation. There are quite a few new members this year and 

overall expenses have been minimal. Major expenses for the 

current year include the workshops, the newsletter mailings, 

and $1000 for the FMCS booth at the upcoming AFS 

meeting. Total assets of the society are about $48,119 with 

about $15,000 yet to be deposited. The board voted not to 

rejoin AIBS. This spring ended our probation period as a 

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charitable nonprofit organization. FMCS is officially 

classified as a nonprofit under section 501 (c) (3). 

Committee Reports 

Symposium Committee 

Kurt Welke held a conference call earlier in June to establish 

committees for organizing the 2005 symposium. 

Committees and committee chairs include: 

Budget (bills, accounting, solicitations, registration) – 

Heidi Dunn 

Registration (advertisement of symposium, tracking) – 

Jennifer Sauer 

Local Arrangements (hotel, catering, etc.) – 

Susan Rogers and Mark Hove 

Communications (call for papers, moderators, assist 

with program) – Ken Lubinski 

Program (abstracts, scheduling, program production) – 

Greg Cope 

Plenary (coordination of keynote speakers) – Kevin 

Cummings 

Awards/Students (announcement, judging, etc.) – 

Catherine Gatenby 

TVA has been contacted for sponsorship (hopefully $2000), 

and other sponsors are needed (including private industry 

and government). The dates for the symposium are May 15- 

19, 2005. Action item requested from the symposium 

committee: FMCS adopt a registration policy and have a 

theme discussion. So far we have waived registration fee for 

plenary speakers and for speakers attending from other 

countries. The board decided to continue the policy of 

waiving fees for symposium plenary speakers but to leave 

the decision to waive other fees to the organizers of each 

meeting. Kurt also solicited help from the board – Heidi 

will do the pre-registration form and help in getting 

sponsors. Exotics was put forth as a potential symposium 

theme. The board had no problem with this theme as long as 

the focus of the symposium remains on native freshwater 

mussels. In an effort to put a more positive spin on this 

theme, several examples were presented: areas of possible 

refugia for native mussels; invasives – can we work with 

them or against them; fighting back against invasives. The 

symposium committee needs to keep this from becoming an 

invasives conference; the program committee will play a 

large role. 

Awards Committee 

The committee did a good job outlining each award, how to 

apply, the nomination process, and selection of recipients. 

The issue of waiving fees for award recipients was put forth. 

The board approved giving lifetime achievement recipients 

lifetime membership. 

Outreach Committee 

Trade show booth update: Set up is Sunday August 22 nd , 

noon to 6 p.m.; trade show floor opens Monday August 23 rd 

from noon to 8 p.m. The trade show social is on the 23 rd 

from 5:30 to 8 p.m. and staffing of the booth is needed. The 

display will remain open on Tuesday and Wednesday and 

staffing of the booth those days is needed. There will be

shells on display, some publications including tools for 

outreach, membership brochures, etc. Kurt requested prices 

for the FMCS publications so he can print order forms to 

have at the booth. Can send T-shirts and sell for $10 a piece, 

Ohio River mussel posters for $3 apiece or $2 each or 2 for 

$5. Kurt needs folks to man the display. The board was 

asked whether FMCS could pay or offset some of the costs 

for attending the AFS meeting for those asked to make 

presentations. Several speakers will talk about mussel life 

history and introduction/background on mussels. Heidi 

suggested giving each organizer and each FMCS committee 

a pot of funds to use as they need with the caveat that each 

committee present a small proposal for spending funds to the 

board for approval. The board should retain oversight on 

how committees plan to spend funds. Committee should 

prepare a small proposal for the board to review. 

Information Exchange Committee 

Walkerana – Dr. Burch is completing the series (2 numbers 

remaining) on the subject of unionids and would like FMCS 

to postpone publishing at least until 2005. Kevin Cummings 

suggested FMCS pay for printing these 2 numbers or at least 

the unionid issue and send to the membership without 

raising dues to get things rolling. Potential printing cost is 

$3000. Paul Johnson has a supplement on the mussels and 

snails of Georgia and possibly the funds to help publish. 

The board agreed to fund the printing of the unionid number 

for Walkerana for up to $3000. The board would like to 

have Kevin submit a proposal outlining the editorial board 

structure and slate of folks for these positions for the board 

to review. Tom will get back with Kevin. 

Environmental Affairs Committee 

The Powell River is overrun with coal fines. Two black 

water events have occurred in the Clinch River recently. 

There was discussion on writing a letter to be sent through 

the committee to the congressman from this district and cc to 

the FWS field office and other environmental organizations. 

Dick Biggins would sign the letter. 

Unionid Status Committee 

The mussel atlas will be published in serial form either in or 

as a supplement to Walkerana. We have 70-80 species 

covered, with about a half dozen folks working on accounts. 

We are still looking for others to write species accounts. A 

suggestion was made to put several accounts on the FMCS 

website – this may generate interest in volunteers to develop 

additional species accounts. Kevin Roe will contact Kevin 

Cummings and the authors about publishing current 

accounts on the website. Once published will this be 

copyrighted? The suggestion was made to have John van 

Hassel check on how an organization can copyright material. 

Also need to check on copyright of the FMCS logo and the 

name of the newsletter. 

Propagation/Restoration Committee 

Registration desk is being set up Monday June 28, 2004 for 

the genetics workshop; about 103 registered to date. Poster 

session is Tuesday evening 7-9 p.m. with the social held 

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concurrently and an open bar. All participants receive a 

free t-shirt. Do we need a policy for registration fee charges 

for workshops and symposiums? Heidi will check on 

expenses and profits from the symposia and workshops. 

Should make more of an effort to get sponsors for 

workshops. 

FMCS needs to begin soliciting ideas from the membership 

for a theme for the next workshop. Ideas include: 

stream/habitat restoration with presentations on what has 

been tried to see what does and doesn’t work; health issues 

and sampling; pharmaceutical chemicals in the active form 

are showing up in streams; endocrine disruptors and how 

they apply to mollusks; human health and mussel health 

workshop. 

Nominations Committee 

Tom will contact Leroy to draft a nomination policy. An 

additional person is needed to serve on the committee. 

MOA with FWS and FMCS 

DOI employees are to get a waiver to serve as an officer in 

professional societies. This used to be signed in the regional 

office, now has to go to the director. FWS wants to 

encourage employees to be liaisons to committees and not 

serve as officers. Ethics counselor for DOI signs for USGS 

folks. MOA itself is not an issue but the waiver is. 

Change in president/president-elect terms 

Since the term for president of FMCS changed from 1 year 

to 2 years, the board needed to determine how to get our 

elections back on track. Two options were proposed: 

1. Tom (current president) serves a 2 year until May 2005. 

2. Bob Anderson (current president-elect) begins serving 

as president now and the board appoints a presidentelect; 

vote for president, president-elect, and secretary 

this fall. 

The board decided to have Tom serve a 2-year term and 

have Bob take the reigns in May 2005 at the symposium. 

Leroy will post a call for nominations for president-elect and 

secretary in Ellipsaria. Tom has appointed an executive 

assistant. 

The FMCS business meeting will be held on Tuesday 

evening. 

FMCS Committee Reports 

Awards 

The primary activity of the Awards Committee during 2004 

has been to draft formal descriptions of the three 

Professional Level Society Awards (Lifetime Achievement, 

William J. Clench Memorial Award, and Meritorious 

Service Award). The descriptions were completed and 

approved by the FMCS Executive Board in June 2004. The 

descriptions and call for nominations for these awards are 

presented elsewhere in the newsletter and are on the Awards 

Committee web page. In addition, preparations have been

made to announce the 2005 Student Travel and Best Student 

Platform and Poster awards that will be given in conjunction 

with the 2005 Symposium in St. Paul, Minnesota. Students 

and their advisors are encouraged to see the Award 

Announcements and Eligibility Requirements in this 

newsletter or on the web. For the upcoming Symposium, a 

cash award of $500 will be presented by the Society to a 

student making the best platform and poster presentation. 

Up to $3000 will be available to qualified students for 

Student Travel Awards to help defer the costs of attending 

the 2005 Symposium. It is anticipated that 10 to 30 travel 

awards of $100 to $300 (actual number and amount depends 

on the number of qualified applicants) will be presented 

during this cycle. 

Submitted by Greg Cope, Chair 

Environmental Quality and Affairs 

No report. 

Gastropod Status and Distribution 

No report. 

Guidelines and Techniques 

Nothing new to report. 

Information Exchange 

Nothing new to report. 

Mussel Status and Distribution 

The progress on the Mussel Atlas continues to move at a 

glacial pace. Any interested members are encouraged to 

sign up for a particular species by emailing the chair 

kroe@delmnh.org and begin working on a species account. 

A list of available accounts can be obtained at the FMCS 

web site or by contacting the committee chair. A proposal 

was floated at the most recent board meeting to publish 

some accounts on the web site as a means of generating 

interest and is being pursued. 

Submitted by Kevin Roe, Chair 

Outreach 

Outreach has been busy! Understatement City, friends. 

Yes, the little group of mussel apostles have been hard at it 

in preparation for our biggest venue yet…AFS 2004. If you 

have been beneath the proverbial rock, here’s the skinny. 

The American Fisheries Society annual meeting (its 134 th ) is 

August 22-26, in lovely Madison Wisconsin. Our session, 

titled "Mussels in America: shells of their former selves", 

will be held on August 24 th . Details can be had by following 

this link: http://www.afs2004madison.org/ This meeting is 

an outstanding opportunity to teach, inform and convert 

fisheries professionals to a new sensitivity towards mussels. 

These are the folks with direct jurisdiction over the rivers, 

streams, and lakes our beloved Unionids inhabit. To that 

end, we are planning the following: 

1. The FMCS Trade Booth. We have space on the 

trade show floor in a prime, highly visible location. 

We have purchased a 10' x 8' "pop-up" that highlights 

the Society, it's products, and the beauty and intrigue 

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of the fauna. The display is welcoming, tasteful, 

and will hopefully lure fish folks into a new 

appreciation. Look for pictures on the FMCS web 

site and the newsletter this fall. And…this is 

available to all members for their venues. The 

display can be easily customized to any audience. 

2. FMCS Mussel Symposium. Jeremy Tiemann has put 

together a stellar cast to spread the word. FMCS 

heavy hitters will present at the Tuesday AFS 

symposium with talks on status, life history, threats, 

recovery efforts, and how to get involved. We will 

lure (more like bribe!) participants to attend by 

raffling off original Dick Biggins' carvings. This 

promises to edify the rank and file and hopefully pull 

in some new society members. 

3. New Tools for Outreach. New contributions were 

solicited this spring and we have collated them into a 

new offering with updated contacts and resources. A 

copy will replace the current net-based .pdf files in 

the near future. This is one of the products to be 

rolled out in Madison as well. 

The April 2004 newsletter mentioned that if you can swing a 

few days in Madison to help out with the booth, we welcome 

all takers. It will have to be on your dime and it is a spendy 

meeting. However, it is a great chance to talk fish and 

mussels and interact with a group our Society needs to 

embrace. 

Submitted by Kurt Welke 

Propagation, Restoration, and Introduction 

See page 3 for workshop summary. 

Symposium 

See page 1. 

FMCS Awards 

Statement of Organization and Purpose 

FMCS Awards are divided into two categories; Professional 

Awards (3 total) and Student Awards (2 total). Each award 

category is evaluated by separate guidelines but maintains 

the intent of the Society's award process. Unless an award 

specifies otherwise, candidates may: 1) be self-nominated 

or nominated by an individual or group; 2) may be from 

North America or any other geographical area; and 3) be 

FMCS members or non-members. Recipients of these 

awards are announced every two years at the FMCS Biennial 

Symposium, during the awards ceremony. 

The FMCS Awards Committee is responsible for the 

administration and oversight of all FMCS awards, but award 

approval, unless otherwise stated, is granted by the FMCS 

Executive Board. The duties of the Awards Committee are 

to: (1) solicit nominations and to recommend recipients for 

each award given by the Society; (2) administer awards to 

students, including selecting the awardees in conjunction 

with a panel of independent judges; and (3) coordinate all 

award activities with the FMCS Treasurer and Executive

Board to ensure the timely and smooth implementation of 

award programs. 

Lifetime Achievement Award 

The Lifetime Achievement Award was established by FMCS 

in 1998 and is given every two years (in conjunction with 

the biennial symposium), if warranted, to an individual for 

singular accomplishments or long-term contributions that 

have advanced the conservation and science of freshwater 

mollusks at a national or international level. The recipient of 

this award must have had (1) activity in one or more aspects 

of freshwater mollusk research and/or conservation for a 

substantial period of time, with a recommended minimum 

guideline of 20 years and (2) made substantial contributions 

to the scientific understanding of freshwater mollusks and/or 

their conservation. 

Nomination Process: The candidate may or may not 

be an active member of FMCS. However, nominations for 

this award must be made by an active FMCS member and 

should clearly and thoroughly document the 

accomplishments of the nominee. 

Application Instructions: An application package 

containing the following materials must be submitted to the 

Chair, FMCS Awards Committee and must contain all of the 

material requested to be considered for the Award. 

1. Letter of Nomination 

2. A curriculum vitae 

3. Documentation of contributions to the field 

4. Three letters of recommendation from active 

members of the Society. The letters should explain 

how the candidate has excelled among colleagues in 

the field with regard to duration and extent of 

contributions. 

Selection of Recipient: The FMCS Awards Committee 

will receive and evaluate nomination packages for 

completeness and eligibility. Packages will be 

forwarded with recommendations to the Executive 

Board of the Society for majority approval. 

Announcement of the Award will be made at the 

Biennial Symposium. The recipient of this award will 

be granted free lifetime membership in the Society 

Completed applications must arrive by 30 November of the 

year preceding the Biennial Symposium. Send to: 

Chair, FMCS Awards Committee 

Past Lifetime Achievement Award Recipients 

Year Name 

1999 John B. Burch 

W. D. Russell-Hunter 

David H. Stansbery 

Paul W. Parmalee 

2001 Richard I. Johnson 

William H. Heard 

2003 Richard G. Biggins 

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William J. Clench Memorial Award 

The William J. Clench Memorial Award was established by 

FMCS in 2000 to recognize outstanding contributions to the 

field of malacology, including significant collections. 

William J. Clench was born on October 24, 1897 and later 

served as Curator of Mollusks at the Harvard University 

Museum of Comparative Zoology from 1926 to 1966. He 

worked toward having a well-balanced worldwide research 

collection, through exchanges, collecting trips, and acquiring 

collections from small private museums or individuals which 

could no longer afford to care for them. He remained active 

in this regard until his death on February 22, 1984. 

This award is given every two years (in conjunction with the 

biennial symposium), if warranted, to an individual for 

singular accomplishments or long-term contributions that 

have advanced the natural history and understanding of 

freshwater mollusks at an academic or non-academic level. 

The recipient of this award must have had (1) activity in one 

or more aspects of freshwater mollusks for a substantial 

period of time, with a recommended minimum guideline of 

20 years and (2) made substantial contributions to the field 

of freshwater malacology. 

Nomination Process: The candidate may or may not 

be an active member of FMCS. However, nominations for 

this award must be made by an active FMCS member and 

should clearly and thoroughly document the 

accomplishments of the nominee. 







3. Documentation of contributions to the field 

4. One letter of recommendation from an active 

member of the Society. The letter should explain 

how the candidate has excelled among colleagues in 

the field with regard to duration and extent of 

contributions. 







Biennial Symposium. 




Past William J. Clench Memorial Award Recipients 

Year Name 

2001 Herbert C. Athearn 

Frieda H. Schilling 

2003 Marian E. Havlik 

W. Henry McCullagh

Meritorious Service Award 

The Meritorious Service Award was established by FMCS in 

2000 and is given every two years (in conjunction with the 

biennial symposium), if warranted, to an individual for 

singular accomplishments or long-term contributions to the 

Society. The recipient of this award must be a past or 

present FMCS member who has performed long-term, 

exceptionally high-quality service to the Society. 

Nomination Process: The candidate must have been an 

active member of FMCS to qualify. Nominations for this 

award may be made by the candidate or by a past or present 

FMCS member. The nomination must be accompanied by 

documentation of service made to the Society. 







3. Documentation of service to the Society 

4. One letter of recommendation from an active 

member of the Society. The letter should explain 

how the candidate has excelled among colleagues 

with regard to duration and extent of service to 

FMCS. 







Biennial Symposium. 




Past Meritorious Service Award Recipients 

Year Name 

2000 Alan C. Buchanan 

Best Student Presentation Awards 

The Best Student Presentation Awards were established by 

FMCS in 2002 and are given every two years (in 

conjunction with the biennial symposium) to an individual 

for the Best Platform (oral) and Best Poster Presentations at 

the symposium. 

Eligibility Requirements: 

All students submitting abstracts, provided they meet the 

eligibility requirements, will be judged for the best student 

platform or poster presentation unless otherwise indicated on 

the abstract submittal form. 

1. Must presently be a student or graduated after March 

in the year preceding the Biennial Symposium 

2. Must be a first author and present the paper 

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3. The presentation must be from research conducted 

as a student and not as a post-graduate 

4. The FMCS encourages students to give presentations 

on preliminary stages of their research prior to any 

data having been collected; however, only 

presentations that contain data (i.e., results) will be 

considered for awards 

5. Student status must be indicated on the abstract 

Past Best Student Platform Award Recipients 

Year Name 

2003 Constance L. Rogers 

Lora L. Zimmerman 

Past Best Student Poster Award Recipients 

Year Name 

2003 Ashley S. McBride 

Student Travel Award 

The Student Travel Award was established by FMCS in 

2002 and is a monetary award given to students to facilitate 

their participation and attendance at the biennial symposium. 

Student Travel Award Criteria: 

Each applicant must 

1. be a student member of the society or 

2. have included dues for membership in their 

application for the travel award. 

Applicants must be making a platform or poster presentation 

at the biannual symposium of the society (written acceptance 

of abstract from FMCS is not required as long as an abstract 

was submitted). 

Application Materials: 

1. A completed application form 

2. A copy of the applicant’s abstract that was already 

submitted through the application submission 

process 

3. A one-page curriculum vitae of the applicant 

4. A letter of recommendation and statement of 

financial need (why travel funds are not available 

from the home institution) from the applicant’s 

research advisor (one page maximum) 

Evaluation Procedure: 

Because the number of awards is limited by available funds, 

applicants will be ranked on the basis of the following 

criteria: 

1. Completeness and timeliness of application package 

2. Quality of the abstract, and 

3. Letter of recommendation and statement of financial 

need 

Past Student Travel Award Recipients 

Year Name 

2003 Jennifer E. Buhay 

Ariel Capili 

Elizabeth A. Neal 

Josh H. Seagraves 

Daelyn A. Woolnough 

Submitted by Greg Cope

Call for 2005 FMCS 

Professional Award Nominations 

Do you know someone who has made worthwhile 

contributions to mussel conservation or to the Society either 

through donating their professional time or expertise or 

through their scientific endeavors? Consider nominating 

them for one of the FMCS Professional Awards. 

Nominations and supporting documentation are due on 30 

November 2004. See the preceding article in this newsletter 

or go to Awards Committee web site for more details. 

Contact Dr. Greg Cope, greg_cope@ncsu.edu or 

919.515.5296, for more information. 

Mississippi Mussel Guide 

Each current FMCS member has received a copy of 

"Freshwater Mussels of the Upper Mississippi River" with 

this newsletter. This booklet was produced by the 

Wisconsin Department of Natural Resources and funded in 

part by FMCS. It is a revision of the popular 1985 booklet. 

Publications 

New Guide to the Freshwater Mussels 

of the New York Metropolitan Region 

and New Jersey 

 

James (Jay) R. Cordeiro 

NatureServe, 11 Avenue de Lafayette, 5 th Fl., Boston, MA 

02111 and 

Division of Invertebrate Zoology, American Museum of 

Natural History, Central Park West at 79 th Street, New York, 

NY 10024 

jay_cordeiro@natureserve.org 

Due to the overwhelming positive response and instructional 

value to the American Museum of Natural History’s Center 

for Biodiversity and Conservation (CBC) annual 

identification workshops (butterflies and moths, dragonflies 

and damselflies, freshwater mollusks, bees), the CBC has 

developed a web handbook as a short-course to the 

freshwater mussels occurring within 75 miles of New York 

City (including southwestern Connecticut) and throughout 

New Jersey. It was based on a 2000 taxonomic workshop on 

freshwater mollusks taught by Dr. David Strayer and myself 

and is designed for naturalists, biologists, and resource 

managers working to conserve local freshwater ecosystems. 

The handbook includes a section on mussel identification 

morphology, a photo illustrated guide to 14 species plus 

9 

three species of invasive freshwater bivalves, a 

dichotomous key, conservation status assessment, habitat 

guide, glossary, bibliography of print and web resources, and 

essays on freshwater mussel taxonomy, diversity and 

distribution, biology, ecology, threats, conservation, and 

study methods, including a guide to local regulations. Keys 

and resources about other invertebrate taxa are also 

available. The handbook is generously illustrated with color 

photographs of specimens from multiple viewpoints, 

morphological shell features, county level distribution maps, 

and sample habitat photographs for each taxon. All 

information is freely available online at: 

http://cbc.amnh.org/mussel/ 

Conservation Plan for Freshwater 

Mussels of the UMR System 

The Conservation Plan for Freshwater Mussels of the Upper 

Mississippi River System (revised May 2004), prepared by 

the Mussel Ad Hoc Committee of the UMRCC, is available 

at: http://www.mississippi-river.com/umrcc/ 

This is intended to be a "living document" that will be 

revised periodically as new information becomes available 

and new strategies are developed. The authors welcome any 

comments and suggestions that would help enhance short- 

and long-term mussel conservation goals. 

Gary Wege, U.S. Fish and Wildlife Service Biologist 

gary_wege@fws.gov 

New Internet Publication: 

History of Malacology 

The American Malacological Society has recently posted 

2,400 Years of Malacology by Eugene V. Coan, Alan R. 

Kabat, and Richard E. Petit on their website: 

http://erato.acnatsci.org/ams/publications/2400_malacology.html 

This is a comprehensive catalog of biographical and 

bibliographical papers on malacologists, conchologists, 

paleontologists, and others with an interest in mollusks. At 

present, the catalog is over 600 pages and indexes over 

5,000 individuals. This catalog is a work in progress, and 

updated versions will be posted periodically. 

Thorsen, W. A., W. G. Cope, and D. Shea. 2004. 

Bioavailability of PAHs: Effects of soot carbon and 

PAH source. Environmental Science and Technology 

38 (7): 2029-2037. 

Article includes information on native freshwater (e.g., 

Elliptio complanata) and marine bivalves. Contact 

Greg Cope (greg_cope@ncsu.edu) for a reprint. 

Peeples, G.E. 2004. Dick Biggins, recovery hero. 

Endangered Species Bulletin 28(4):24-25.

Contributed Articles 

Ohio Pigtoe Host Suitability Trials 

G. Thomas Watters and Kody Kuehnl 

Department of Evolution, Ecology, and Organismal Biology, 

The Ohio State University, 1315 Kinnear Rd., Columbus, 

OH 43212 

and 

The Columbus Zoo and Aquarium Freshwater Mussel 

Conservation and Research Facility, PO Box 400, Powell, 

OH 43065 

Yokley (1972) identified Redfin Shiner as a host for the 

Ohio Pigtoe, Pleurobema cordatum (Rafinesque, 1820); 

Fuller (1974) added Bluegill. No work seems to have been 

done on that mussel species since. Specimens of the Ohio 

Pigtoe, an Ohio endangered species, were collected in 

August, 2003, in the Muskingum River at Devola, Ohio. 

They have been kept in captivity at the Columbus Zoo and 

Aquarium Freshwater Mussel Conservation and Research 

Facility since that time. On 19 May 2004 numerous (~50) 

conglutinates were released by one or more mussels. 

(Additional conglutinates were sporadically released into 

mid-June). Conglutinates were 10-15 mm in length, white, 

composed of two longitudinal bodies connected by ladderlike 

rungs. Most of the conglutinate was composed of 

unfertilized, structural eggs in which were embedded 

functional glochidia. Conglutinates were fed to fish on 19 

May. Fish were obtained from a nearby reservoir and a local 

bait store and maintained at 18-20°C in isolated AHAB 

tanks. Typically, many of the cyprinids were lost before 

completion of the tests. 

Fish that facilitated metamorphosis: 

Creek Chub, Guppy, Brook Stickleback. The encystment 

period was 27-46 days, with most juveniles collected over 

days 43-45. Juveniles newly metamorphosed had nearly 

doubled in length from their glochidial size. Brook 

stickleback was the best host; a single fish produced >40 

juveniles. Guppy was a marginal host. Neither Guppy nor 

Brook Stickleback co-occur with Ohio Pigtoe in Ohio. 

Fish that did not facilitate metamorphosis: 

Bluegill, Goldfish, Green Darter, Green Sunfish, Stonecat, 

White Crappie. 

Fuller’s identification of Bluegill as a host was not 

substantiated. 

Reference 

Fuller, S.L.H. 1974. Clams and mussels (Mollusca: 

Bivalvia). Pp. 215-273 in C.W. Hart & S.L.H. Fuller 

(eds.), Pollution Ecology of Freshwater Invertebrates. 

Academic Press, New York. 389 pp. 

10 

Yokley, P. 1972. Life history of Pleurobema cordatum 

(Rafinesque, 1820) (Bivalvia: Unionacea). Malacologia 

11: 351-364. 

A first attempt towards a compilation 

of a list of predators of freshwater 

molluscs in Israel and Palestine 

Henk K. Mienis 

National Mollusc Collections 

Dept. Evolution, Systematics & Ecology 

Hebrew University of Jerusalem 

IL-91904 Jerusalem, Israel, and 

The Zoological Museum, Tel Aviv University 

IL-69978 Tel Aviv, Israel 

mienis@netzer.org.il 

Freshwater molluscs play an important role in the food chain 

of many other animals, yet little information has been 

published about this subject in the Levant. For example, 

most records of predation on aquatic molluscs have appeared 

in the form of short faunistic notes, often in journals hardly 

available to the general public. This is a first attempt to 

summarize the published and unpublished information 

dealing with predation on freshwater molluscs in Israel and 

Palestine. Only records of prey species identified at least at 

the generic level have been included in the list. I hope 

sincerely that it will lead to a stream of additional records. 

In the near future a similar list will be published dealing with 

the parasites recorded so far from freshwater molluscs in the 

same region. These parasites exploit the molluscs often as a 

intermediate station in order to reach their final host: often a 

predator of the aquatic molluscs. 

Predators of freshwater molluscs in Israel and Palestine 

Family NERITIDAE 

Theodoxus (Neritaea) jordani jordani (Sowerby, 1836) 

Starling – Sturnus vulgaris (Mienis, unpublished). 

Theodoxus (Neritaea) karasuna (Mousson, 1874) 

Damascus barbel – Capoeta damascina (Mienis, 

unpublished). 

Theodoxus (Neritaea) michonii (Bourguignat, 1852) 

Rainbow trout – Oncorhynchus mykiss (Mienis, 

unpublished); 

Levantine dace – Pseudophoxinus kervillei (Mienis, 

unpublished); 

Josephus cichlid – Astatotilapia flavijosephi (Mienis, 

unpublished); 

Mediterranean Hooded Crow – Corvus corone 

sardonius (Aharoni, 1938). 

Family BITHYNIIDAE 

Bithynia phialensis (Conrad, 1852) 


unpublished); 


unpublished);


unpublished); 


Family THIARIDAE 

Melanoides tuberculatus (Müller, 1774) 

Leech – Helobdella punctatolineata** (Mienis, 1986a); 

Levant freshwater crab – Potamon potamios* (Mienis, 

unpublished); 


unpublished); 

Glossy ibis – Plegadis falcinellus (Mienis, 1997); 

Starling – Sturnus vulgaris (Mienis, unpublished); 


sardonius (Aharoni, 1938 as Melania). 

Family MELANOPSIIDAE 

Melanopsis buccinoidea (Olivier, 1801) 


2003); 


unpublished); 


unpublished); 

Starling – Sturnus vulgaris (Mienis, unpublished); 

Brown-necked raven – Corvus ruficollis (Aharoni, 1938 

as M. praemorsa). 

Melanopsis cerithiopsis Bourguignat, 1884 


unpublished); 


Melanopsis costata costata (Olivier, 1804) 


unpublished); 


unpublished); 


Melanopsis costata jordanica Roth, 1839 


unpublished); 

Yellow-vented bulbul – Pycnonotus xanthopygos 

(Mienis, 1994b as M. praemorsa jordanica). 

Melanopsis lampra Bourguignat, 1884 


unpublished); 

Family VALVATIDAE 

Valvata (Cincinna) saulcyi Bourguignat, 1853 


unpublished); 


Family PHYSIDAE 

Haitia acuta (Draparnaud, 1805) 

Leech – Helobdella punctatolineata** (Mienis, 1986a 

as Physella acuta); 

Banded newt – Triturus vittatus (Mienis, unpublished); 


unpublished); 

Levantina frog – Rana levantina (Mienis, 1996 as 

Physella acuta); 

Lapwing – Vanellus vanellus (Mienis, 1985 as Physella 

acuta); 

11 

Black-winged stilt – Himantopus himantopus 

(Mienis, 1994a as Physella acuta); 

Green sandpiper – Tringa ochropus (Mienis, 1986b as 

Physella acuta). 

Family PLANORBIDAE 

Bulinus (Isidora) truncatus (Audouin, 1826) 

Green Sandpiper – Tringa ochropus (Mienis, 1986b). 

Gyraulus (Gyraulus) piscinarum (Bourguignat, 1852) 


unpublished). 

Planorbella duryi (Wetherby, 1879) 

Leech – Helobdella punctatolineata** (Mienis, 1986a 

as Helisoma duryi). 

Planorbis species 

Green Sandpiper – Tringa ochropus (Cockburn, 1946). 

Family LYMNAEIDAE 

Pseudosuccinea columella (Say, 1817) 

Leech – Helobdella punctatolineata** (Mienis, 1986a); 

Moorhen – Gallinula chloropus (Mienis, 1987); 

Green sandpiper – Tringa ochropus (Mienis, 1990). 

Family UNIONIDAE 

Unio mancus eucirrus Bourguignat, 1857 

Moorhen – Gallinula chloropus (Mienis, unpublished). 

Unio terminalis delicatus Lea, 1863 

Kingfishers – either Halcyon smyrnensis or Ceryle rudis 

(Ashkenazi and Mienis, unpublished). 

Unio terminalis terminalis Bourguignat, 1852 

Moorhen – Gallinula chloropus (Mienis, unpublished); 


sardonius (Mienis, unpublished). 

Family CORBICULIDAE 

Corbicula consobrina (Cailliaud, 1823) 


unpublished). 

Corbicula fluminalis (Müller, 1774) 


unpublished). 

* So far only one species of freshwater crab has been 

reported from Israel and Palestine (Potamon potamios s.l); 

however, according to Dr. Sh. Ashkenazi (pers. com.) two 

morphological types seem to occur among these crabs. 

** Helobdella triserialis seems to be an older name for 

this North-American species (Bromley, 1994). 

Acknowledgements 

I like to thank Dr. Sh. Ashkenazi (Hebrew University of 

Jerusalem) and Mr. Shalom Hayat for donating large 

quantities of freshwater mussels that were probably predated 

upon by a species of Kingfisher to the National Mollusc 

Collection of the Hebrew University of Jerusalem. Likewise 

I wish to thank the students of Prof. Menachem Goren (Tel 

Aviv University) for regularly showing me the molluscs 

found in the intestines of freshwater fishes. 

References 

Aharoni, I., 1938. [The Ravens of Palestine.] The Magnes 

Anniversary Book, 257-266. (in Hebrew with English 

summary)

Bromley, H.J., 1994. The freshwater leeches (Annelida, 

Hirudinea) of Israel and adjacent areas. Israel Journal of 

Zoology 40:1-24. 

Cockburn, T.A., 1946. Bird parasite and crop investigations. 

Middle East Biological Scheme, Jerusalem Naturalists’ 

Club, Bulletin 14:7. 

Mienis, H.K., 1985. Lapwings feeding on Physella acuta 

near Ramla, Israel. Levantina 54:618-619. 

Mienis, H.K., 1986a. An American Leech, Helobdella 

punctatolineata, in Israel. Zoology, in the Middle East 

1:153-154. 

Mienis, H.K., 1986b. Green sandpipers feeding on 

freshwater snails near Ramla, Israel;. Levantina 64:687- 

688. 

Mienis, H.K., 1987. Moorhens Gallinula chloropus feeding 

on Pseodosuccinea columella snails in Israel. Mitt. 

Zool. Ges. Braunau 5(1-4):57-58. 

Mienis, H.K., 1990. Pseudosuccinea columella snails as 

food items of the Green sandpiper Tringa ochropus. 

Mitt. Zool. Ges. Braunau 5(9-12):187-188. 

Mienis, H.K., 1994a. Predatie op Physella acuta door 

Steltkluten in Israel. C.B. Ned. Malac. Ver. 280:119- 

120. 

Mienis, H.K., 1994b. Subfossil shells in pellets of the Barn 

owl in Israel. Malakologiai Tajekoztato 13:39. 

Mienis, H.K., 1996. Physella acuta in a faecal pellet of Rana 

levantina. The Papustyla 10(5):3. 

Mienis, H.K., 1997. A case of predation on Melanoides 

tuberculata by the Glossy ibis in Israel. The 

Conchologists’ Newsletter 140:783-784. 

Mienis, H.K., 2003. Molluscs from the excavation of Tel 

Kabri (with an appendix dealing with Crustaceans found 

at that site). Triton 7:28-37. 

Survey of the Freshwater Mussels of 

the Channelized Missouri River 

Ellet Hoke 

Midwest Malacology, Inc. 

1878 Ridgeview Circle Drive 

Manchester, MO 63021 

ellethoke@earthlink.net 

Investigations conducted at 67 sites along the channelized 

Missouri River and its floodplain lakes and sloughs in 

Missouri, Kansas, Iowa, and Nebraska primarily between 

1988 and 1990 documented the presence of 15 unionid 

species. The most common unionids collected from the 

Missouri River were Anodonta suborbiculata, Leptodea 

fragilis, Potamilus alatus, P. ohiensis, and Pyganodon 

grandis. Other species documented for the channelized 

reach of the Missouri River were: Lampsilis teres, 

Lasmigona c. complanata, Leptodea leptodon, Obliquaria 

reflexa, Obovaria olivaria, Quadrula quadrula, Toxolasma 

parvus, Truncilla donaciformis, and Utterbackia imbecillis. 

The bivalve Corbicula fluminea was found to be common in 

the reach extending from the confluence of the Missouri 

12 

River above St. Louis to about 50 miles east of Kansas 

City. This bivalve was not recovered in any other reach of 

the river. 

The location of unionid populations in the Missouri River 

almost always coincided with the presence of stable 

substrates and relatively slow currents. The most productive 

habitats were as follows: areas immediately downstream and 

along the inside bank of river bends; behind wing dams; and 

along rock lined banks where the gaps between rocks had 

filled with sediment. Mussels were rare or absent along 

banks immediately adjacent to rapid currents. 

Floodplain lakes and sloughs contained a less diverse fauna 

comprised of the following eleven species: Anodonta 

suborbiculata, Lampsilis teres, Lasmigona c. complanata, 

Leptodea fragilis, Potamilus alatus, P. ohiensis, Pyganodon 

grandis, Quadrula quadrula, Toxolasma parvus, Uniomerus 

tetralasmus, and Utterbackia imbecillis. 

A paper on this survey is nearing completion and will be 

submitted for publication in the near future. 

Problems with dating freshwater snails 

from extinct populations in Israel 


National Mollusc Collections 

Dept. Evolution, Systematics & Ecology 

Hebrew University of Jerusalem 

IL-91904 Jerusalem, Israel, and 

The Zoological Museum, Tel Aviv University 

IL-69978 Tel Aviv, Israel 


Israel is situated in an area suffering from increasing 

desertification. At many widely separated locations you can 

come across deposits showing a rich assortment of 

freshwater shells, while on the map you will look in vain for 

a spring, a stream, or any other aquatic biotope in the 

vicinity. If such shells are found within the context of a 

historical site, then the mollusc remains can be dated with 

the help of a coin, a pottery shard ,or another item to which a 

precise date can be fixed. In cases that easily datable 

artifacts are not available, then we have to look for other 

means to get a reliable date for these extinct mollusc 

populations. Such dates can be obtained by estimating the 

age with the help of radiocarbon dating techniques carried 

out at the Weizmann Institute of Science in Rehovot. In 

most cases, especially with charcoal, pieces of wood, bones, 

ostrich egg-shells, and marine molluscs, this technique 

seems to work perfectly; however, in the case of freshwater 

molluscs, we have come across some unexpected anomalies 

in the results. 

Some examples of anomalies in dating freshwater shells 

Numerous areas in the south of Israel - the Negev desert and 

the Arava Valley - show evidence of former much wetter

periods than today. At many places intensive farming with 

the help of ingenious irrigation methods have been 

discovered. Most of these farms have one thing in common: 

they were deserted towards the end of the Byzantine period 

or the beginning of the Early Islamic period (6 th -7 th Century 

C.E.). One such farm was situated near ‘En ‘Avrona in the 

Arava Valley, north of Elat. Samples taken from an 

irrigation pool, which stopped functioning during the Early 

Islamic period, contained numerous specimens of 

Melanoides tuberculatus (Müller, 1774), Melanopsis 

buccinoidea (Olivier, 1801) and what seems to be an 

undescribed species of Heleobia. Specimens belonging to 

Melanopsis were submitted for radiocarbon dating and this 

resulted in a C value of 6575+90 BP (BP=before present) 

(Segal & Carmi, 1996: 98), i.e. about 5000 years older than 

it was supposed to be. 

Near Tel Goded, in the Judean lowlands, an interesting water 

system which provided water to the ancient settlement of Bet 

Guvrin during historic times has been excavated (Sagiv et 

al., 1997). This water system consisted of a man-made pool 

and a stone-hewn canal running from Tel Goded to Bet 

Guvrin. At the bottom of the pool, more exactly in the 

south-eastern corner of it, a rich assortment of freshwater 

molluscs was discovered, showing that the pool was fed with 

running water year round (Mienis, in preparation). Today 

you will look in vain for a spring or streamlet on the map of 

that area. That part of the pool had been built during or after 

the reign of Constantius II (337-361 C.E.); a coin dating 

back to that period was found pressed into the plaster at the 

base of its inner wall. A batch of snails belonging to 

Melanopsis buccinoidea was also submitted for radiocarbon 

dating, which resulted in a C value of 5615+60 BP (Segal 

& Carmi, 1996: 91-92), i.e. about 4000 years too old. 

How can we explain these anomalies? 

According to Segal & Carmi (1996: 92) these anomalies are 

caused by the incorporation of water carbonate into the 

shells by the snails as seen from the C value. In my 

opinion it is much more likely that Melanopsis picks up old 

carbonate particles while scraping algae from the submerged 

limestone rocks and other objects in its aquatic environment. 

A part of this material is being incorporated in the shell and 

this results in a mixing of old and contemporary carbonates 

(i.e. from the time the snail actually lived) within the shell 

matrix. Since such a mixture is highly environmentally 

dependant and may vary from place to place, it is probably 

impossible to find a tool for calibrating the C values. At 

the Weizmann Institute they are now looking for a method to 

separate organic particles from freshwater shells and to 

restrict the radiocarbon dating process to these fractions in 

order to obtain more reliable dating results. 

References 

Sagiv, N., Zissu, B. & Amit, D., 1997. [A water system 

north of Bet Guvrin.] In Y. Eshel (Ed.): Studies of Judea 

and Samaria, 245-258. (in Hebrew) 

Segal, D. & Carmi, I., 1996. Rehovot radiocarbon date list 

V. Atiqot 29 (supplement):79-106. 

13 

Mussel Studies 

Steve Ahlstedt 

U. S. Geological Survey, Knoxville, TN 37921 

(865) 545-4140 x.17 

ahlstedt@usgs.gov 

Clinch and Powell Rivers, TN/VA 

Every five years since 1979, mussels in the Clinch and 

Powell Rivers are quantitatively evaluated (quadrate 

sampling). This has established long-term trend monitoring 

for mussel populations in both drainages. It was determined 

that only 12 sites would be evaluated in 2004 (6 each in both 

rivers). Sampling in the Powell River is completed. Mussel 

densities and species composition continue a downward 

trend. The Powell suffers from high concentrations of coal 

fines mined and released from settling lagoons in Virginia. 

Four of the six sites in the Clinch are finished and sites in 

Tennessee (3) show a continued upward trend in mussel 

densities and species composition. Many juvenile T & E 

species, especially E. capsaeformis and E. brevidens, were 

noted along with C. stegaria and P. plenum. Of concern is 

the amount of coal fines showing up in Tennessee from 

mining operations in Virginia. Black-water events from coal 

mining operations are a serious problem and unless it is 

stopped, the Clinch will look like the Powell. 

Powell River, TN/VA 

Gravid P. cyphyus and L. fasciola were collected from the 

Powell River for culture and propagation at Virginia’s Buller 

hatchery and Virginia Tech’s mussel culture facility. This is 

a joint effort between Virginia Department of Fish and 

Game, VPI, FWS, and USGS. Goals are to restore mussel 

populations in the Powell River. 

Cumberland Plateau – Upper Caney Fork, TN 

Survey efforts in the upper Caney Fork drainage found some 

new populations of Pleurobema gibberum and 

Venustachoncha sima. No P. fabula or L. diversa were 

found. Survey will continue this fall. 

Big South Fork Cumberland (TN), Clinch River 

(TN), Paint Rock River (AL), Mobile River Basin 

Continue assisting with the collection of gravid T & E 

mussel species for culture and propagation at Virginia 

Tech’s mussel culture facility and Tennessee Aquarium 

Research Institute’s mussel culture facility located in 

Cohutta, Georgia. Gravid E. capsaeformis, E. brevidens, 

E. walkeri, V. perpurpurea, T. cylindrellus, P. greenii, and 

L. altilus were found for culturing. 

Duck River Report and Juvenile Mussel Habitat 

Study 

Final report to The Nature Conservancy is just about 

completed. Will be starting a juvenile mussel habitat study 

related to increased flows in the Duck River. Increased 

minimum flows in the last 10 years from TVA’s Normandy

Dam has created nursery areas along the shoreline of the 

river that previously went dry. Efforts will be concentrated 

at two sites. Work is expected to begin in August or 

September and was funded by the FWS Cookeville, 

Tennessee field office. 

Recovery Plan Development for Tennessee and 

Cumberland River Basins 

Efforts are underway to develop a recovery plan for mussels 

in the Tennessee and Cumberland River basins. This is a 

joint effort at identifying the most critically rare species and 

prioritizing which species to concentrate on first that are 

recoverable or offer the best chances for success. Some 

species are at critical low levels and may not survive the 

next ten years. State and federal agencies involved with this 

process include TN, KY, VA, NC, USGS, and FWS. 

Additional new records of freshwater 

mollusks (Gastropoda & Bivalvia) 

from Santa Catarina State, Southern 

Brazil, with complementary 

observations about mollusks fauna 

from the Mampituba river basin 

A. Ignacio Agudo 

Projeto Naiade (Naiade Project) 

Centro Integral de Educação Ambiental Cachoeira (CIEAC) 

(Integral Center of Environ. Education of the Waterfall), 

Caixa Postal (P. O. Box) 010, 88010-970 Centro, 

Florianópolis, Santa Catarina – SC, Brasil 

iagudo@lycos.com 

http://www.intergate.com.br/malacologia 

Keywords: Freshwater mollusks; Gastropoda; Bivalvia; 

Continental malacological fauna survey; Mampituba river 

basin; Santa Catarina state; Southern Brazilian country 

The malacological inventory recently conducted in the Santa 

Catarina´s state continental territory (Agudo 2004b) 

included new reports of freshwater mollusks species and 

other observations from the Mampituba river basin region 

(Agudo 2004a). Seven new records of native mussels/ 

naiades and 1 freshwater snail were included from the State, 

elevating the known number of freshwater species to 47 (27 

Gastropoda and 20 Bivalvia: 15 Unionoida & 5 Veneroida) 

(Agudo 2004b). 

Systematic Species List: new records 

Class GASTROPODA 

Subclass Prosobranchia 

Family HYDROBIIDAE (1) 

Potamolithus catharinae Pilsbry, 1911 

Class PELECYPODA = BIVALVIA 

Order Unionoida 

Family MYCETOPODIDAE (2) 

Anodontites iheringi (Clessing, 1882) 

Anodontites lucidus (Orbigny, 1835) 

14 

Family HYRIIDAE (4) 

Diplodon aethiops (Lea, 1860) 

Diplodon martensi (Ihering, 1893) 

Diplodon pilsbryi Marshall, 1928 

Diplodon rhuacoicus (Orbigny, 1835) 

Order Veneroida 

Family SPHAERIIDAE (1) 

Sphaerium observationis (Pilsbryi, 1911) 

From the Mampituba River Basin, the boundary between the 

states of Santa Catarina and Rio Grande do Sul (Agudo 

2004a), only four short contributions were found in the 

literature: Pereira (1993) from the freshwater snail Chilina 

sp, and Guterres & Guterres (1997 a, b, c) from the estuarine 

snail Melampus coffeus (Linnaeus, 1758), with particular 

reference to other two local species (Guterres & Guterres 

1997 b): Heleobia piscium (= australis) (Orbigny, 1835) and 

Littorina flava King & Broderip, 1832. 

The malacological inventory also included one regional new 

species record of invasive exotic mussel or naiade: 

Class PELECYPODA = BIVALVIA 


Family CORBICULIDAE (1) 

Corbicula fluminea (Müller, 1774) 

...and one native terrestrial snail: Punctum (Toltecia) pilsbryi 

(Scott, 1957). 

For a general visual of the zoogeographical records in this 

Brazilian territory, see Agudo (2004c). 

References 

Agudo, A. Ignacio. 2004a. Preliminary report on the 

freshwater mollusks fauna of Mampituba river basin, 

Santa Catarina State, Southern Brazil. FMCS Newsletter 

Ellipsaria, 6 (1): 10-11. 

Agudo, A. Ignacio. 2004b. The freshwater mollusks fauna of 

Santa Catarina State, Southern Brazil: a comprehensive 

general synthesis. FMCS Newsletter Ellipsaria, 6 (1): 

11-12. 

Agudo, A. Ignacio. 2004c. Levantamento Atualizado – 

Espécies em Santa Catarina (Up-to-date rising – Species 

in SC). Florianópolis, SC: AVULSOS 

MALACOLÓGICOS – AM, Internet Brazilian Website: 

http://www.intergate.com.br/malacologia/levantamento/ 

levantamento.html 

Guterres, Luís Fernando dos Reis & Patrícia Macedo de 

Castro Guterres. 1997a. Comparação entre as relações 

do peso da concha/peso da parte mole e 

peso/comprimento de duas populações de Melampus 

coffeus (Linnaeus, 1758) (Pulmonata, Ellobiidae) na 

Região Sul do Brasil. Florianópolis, SC: Resumos XV 

Encontro Brasileiro de Malacologia – XV EBRAM, 

Julho 21 a 25 de 1997, p. 58. 

Guterres, Luís Fernando dos Reis & Patrícia Macedo de 

Castro Guterres. 1997b. Estudo comparativo de 

Melampus coffeus (Linnaeus, 1758) (Pulmonata, 

Ellobiidae) em duas áreas distintas no sul do Brasil. 

Florianópolis, SC: Resumos XV Encontro Brasileiro de

Malacologia – XV EBRAM, Julho 21 a 25 de 1997, p. 

59. 

Guterres, Patrícia Macedo de Castro & Luís Fernando dos 

Reis Guterres. 1997c. Crescimento de Melampus 

coffeus (Linnaeus, 1758) (Pulmonata, Ellobiidae) no 

Rio Mampituba, Torres, Rio Grande do Sul, Brasil. 

Florianópolis, SC: Resumos XV Encontro Brasileiro de 

Malacologia – XV EBRAM, Julho 21 a 25 de 1997, p. 

59. 

15 

Pereira, Paulo Antonio Cypriano. 1993. Sobre o encontro 

de formas albinas em uma populacao de Chilina Gray, 

1828 (Mollusca; Gastropoda; Chilinidae), no canhao do 

Malacara, Praia Grande, Santa Catarina. Rio de Janeiro, 

RJ: Programa e Resumos XIII Encontro Brasileiro de 

Malacologia – XIII EBRAM, Julho de 1993, p. 81. 

A review of the United States distribution of Melanoides tuberculatus 

(Müller, 1774), an exotic freshwater snail 

Tamara K. Anderson 

University of Colorado Museum 

Boulder, Colorado 

Address for Correspondence: 285 Smith Street, Lander, Wyoming 82520 

E-mail: tamander@onewest.net 

INTRODUCTION 

Melanoides tuberculatus (Müller, 1774), the red-rim melania, is native to Africa, the Middle East, and southeast Asia (Pilsbry 

and Bequaert 1927), but is now common in tropical to semi-tropical areas around the globe. The species is live-bearing and 

generally considered to be parthenogenetic, although evidence exists for sexual reproduction in some populations (Livshits and 

Fishelson 1983; Heller and Farstey 1990). M. tuberculatus is a secondary host of several parasites that affect various species of 

fish and birds. 

Melanoides tuberculatus has invaded many areas of the globe with varying effects on the local gastropods. Abbott (1973) noted 

its spread into Puerto Rico, Mexico, and central America in the 1970s. M. tuberculatus may have a competitive advantage over 

native snails due to its ability to reach high densities (Pontier et al. 1991). In Mexico, the arrival of M. tuberculatus coincided 

with decreases in native gastropod populations (Contreras-Arquieta et al. 1995, as cited in Contreras-Arquieta 1998). Several 

studies in the Caribbean also showed negative impacts on Biomphalaria glabrata (Pointier and McCullough 1989; Pointier et al. 

1989; Pointier 1993). M. tuberculatus may also have contributed to the decline in B. glabrata in Venezuela (Pontier et al. 1994). 

In Africa, however, M. tuberculatus does not appear to displace freshwater pulmonate snails (Mkoji et al. 1992). Due to the 

potential effects on the aquatic community, the distribution of this species is important to track. 

METHODS 

In July 2001, I surveyed aquatic mollusks in Bridal Veil Falls and portions of Cascade Creek, Cool Creek, and the Cheyenne 

River on The Nature Conservancy's Whitney Preserve and adjacent properties in southwestern South Dakota. An informal 

sampling design allowed baseline information to be gathered with a minimum effort. The substrate at eleven points was sampled 

using a net and sieve bucket. 

In addition to my own survey results in South Dakota, I used available literature sources to research locations of Melanoides 

tuberculatus. Through the internet, I found records of occurrence at the Florida Natural History Museum, the Philadelphia 

Academy of Natural Sciences, and the Smithsonian Institution. Database searches at the Illinois Natural History Museum, the 

Bailey-Mathews Shell Museum, and the National Museums and Galleries of Wales failed to yield any U.S. records for M. 

tuberculatus. I visited the Field Museum of Natural History in Chicago and searched their collections in person. Contacts at the 

Delaware Museum of Natural History, the University of Michigan, and the Santa Barbara Museum of Natural History provided 

records from those collections. Personal communications with others who had recent field observations resulted in additional 

location information. 

RESULTS 

Shells of Melanoides tuberculatus were collected from one point in Cascade Creek. The water temperature at the collecting site 

was 21 o C. The shells measured from 5.3 mm to 13.8 mm in length. Robert Herschler confirmed the identification of the samples 

(pers. com.). Voucher specimens were deposited at the Field Museum of Natural History in Chicago (Voucher #296306). 

The known current distribution of Melanoides tuberculatus is summarized in Figure 1 and Table 1*. The distribution discussed 

here is limited to museum records, published literature reports, and personal communications with other scientists known to be

sampling in the field. More locations are likely along the same drainages in the south (especially in Texas and Florida), but are 

not plotted unless specific evidence exists for the presence of M. tuberculatus. 

Melanoides tuberculatus has been in the United States for almost half a century, although the exact date of the first incidence is 

unclear. Records exist for locations in Arizona in the 1950s. The original Arizona population was apparently absent by the 

1970s (Murray 1971), although another population may have been present (Dundee 1974). Further investigation is necessary to 

clarify these early reports because no Arizona records appear later. Certainly the species was in Florida and Texas by the early 

1960s (Clench 1969; Murray 1964). In the 1970s, the known U.S. distribution included Florida, Texas, Louisiana, Arizona, and 

Oregon (as summarized by Dundee 1974; Dundee and Paine 1977; Burch and Tottenham 1980). Museum records and published 

monographs revealed M. tuberculatus was present in Colorado, California, and Hawaii in the 1980s. A compilation of more 

recent, unpublished sources show additional isolated populations exist throughout the western United States in Idaho, Montana, 

Wyoming, Nevada, and Utah. 

Cowie (1997) stated that M. tuberculatus likely was present in Hawaii many years before he recorded it. The lack of earlier 

records in Hawaii illustrates that first records are the result of a combination of when a species reaches an area and whether 

anyone is looking for it. It is unknown whether the increased number of known locations reflects recent expansions of the 

species or a recent increase in mollusk surveys in the western U.S.. The latter might be the case for springs in remote areas of the 

Great Basin and northern Rocky Mountains. However, recent sightings in North Carolina indicate that the range is probably still 

increasing, at least in some areas. 

DISCUSSION 

As with any invading species, it is important to understand the mechanisms that contribute to the spread of the species. Factors 

influencing the dispersal and restrictions to its expansion of Melanoides tuberculatus are discussed below. 

Method of Dispersal 

The pet trade is blamed for the introduction of these animals into Florida and Texas (Murray 1971). Escape or release from 

personal aquariums is the speculated method of dispersal to many isolated springs. The presence of tropical fish in all of the 

springs where the snails are found in Montana, Idaho, and Wyoming (D. L. Gustafson, pers. com.) supports this theory. 

Not all populations are permanent. A population in Phoenix was reported present in 1955, but not present by the 1970s (Dundee 

1974). A 1963 Oregon record appears at the University of Michigan Museum (L. Appleton, pers. com.). In recent surveys of 

Oregon springs (T. Frest, pers. com.) no Melanoides tuberculatus were located, although it is unclear if the exact location was 

revisited. The reasons for these "disappearances" are unknown, but could represent introductions in areas where temperature 

tolerances were exceeded. 

Further Expansion of Range 

Further range expansion is currently limited by temperature constraints. Most, if not all, of the western localities are in isolated 

thermal springs. The warmer water can insulate the snails from the cold winters in the area. Murray (1971) states Melanoides 

tuberculatus has a temperature tolerance of 18 to 25 o C. Mitchell and Brandt (unpublished data) suggest water temperatures 

between 18 o C and 32 o C could support populations, which indicates the potential geographical range is less restricted than 

previously thought. Global warming may also increase the suitable habitat for the species. Food availability and water velocity 

may secondarily affect the distribution of the species, at least at the local level (Laamrani et al. 1997). The malacological 

community should continue to track the range of Melanoides tuberculatus in order to determine if the species is having an impact 

on the native fauna. 

ACKNOWLEDGEMENTS 

Special thanks to T. Brandt, A. Mitchell, and D. L. Gustafson for sharing their information. G. Anderson provided field 

assistance. R. Herschler confirmed identification of South Dakota specimens. H. Chaney provided information on the Santa 

Barbara Museum collection. L. Appleton provided information on the University of Michigan collection. L. Skibinski provided 

information on the Delaware Museum of Natural History collection. J. Gerber and others at the Field Museum provided access to 

their collection. Thanks to the Philadelphia Academy of Natural Sciences and the Florida Natural History Museum for allowing 

internet access to their aquatic gastropod records. Personal communication with the following individuals provided additional 

information: B. Bosworth, W. C. Fields, Jr., R. F. McMahon, D. Smith, B. T. Watson, D. Sada, H. D. Murray, M. Vinson, and T. 

Frest. R. Dillon provided useful comments on an early draft. 

LITERATURE CITED 

Abbott, R. T. 1973. Spread of Melanoides tuberculata. The Nautilus 87: 29. 

Burch, J. B. and J. L. Tottenham. 1980. North American freshwater snails: species list, ranges and illustrations. Walkerana 1:81- 

215. 

Clench, W. J. 1969. Melanoides tuberculata (Muller) in Florida. Nautilus 83:72. 

16

Contreras-Arquieta, A. 1998. New records of the snail Melanoides tuberculata (Müller, 1774) (Gastropoda: Thiaridae) in the 

Cuatro Cienegas Basin, and its distribution in the state of Coahuila, Mexico. Southwestern Naturalist 43:283-286. 

Contreras-Arquieta, A., G. Guajardo-Martinez, and S. Contreras-Balderas. 1995. Thiara (Melanoides) tuberculata (Müller, 

1774) (Gastropoda: Thiaridae) probable impacto ecologico en Mexico. Publications Biology, FCB/UANL, Mexico, 8(1-2): 

17-24. 

Cowie, R. H. 1997. Catalog and bibliography of the non-indigenous non-marine snails and slugs of the Hawaiian Islands. Bishop 

Museum Occasional Papers 50: 1-66. 

Dundee, D. S. 1974. Catalog of introduced molluscs of eastern North America (north of Mexico). Sterkiana 55:1-37. 

Dundee, D. S. and A. Paine. 1977. Ecology of the snail Melanoides tuberculata (Muller), intermediate host of the human liver 

fluke (Opisthorchis sinensis) in New Orleans, Louisiana. The Nautilus 91:17-20. 

Frest, T. J. and E. J. Johannes. 2002. Land snail survey of the Black Hills National Forest, South Dakota and Wyoming, Summary 

Report 1991-2001. Unpublished report to the USDA Forest Service. 

Heller, J. and V. Farstey. 1990. Sexual and parthenogenetic populations of the freshwater snail Melanoides tuberculata in Israel. 

Israel Journal of Zoology 37: 75-87. 

Howells, R. G. 1992. Annotated list of introduced non-native fishes, mollusks, and plants. Texas Parks and Wildlife Management 

Data Series 78:1-19. 

Laamrani, H., K. Khallayoune, B. Delay, and J. P. Pointier. 1997. Factors affecting the distribution and abundance of two 

prosobranch snails in a thermal spring. Journal of Freshwater Ecology 12:75-79. 

Livshits, G. and L. Fishelson. 1983. Biology and reproduction of the freshwater snail Melanoides tuberculata (Gastropoda: 

Prosobranchia) in Israel. Israel Journal of Zoology 32:21-35. 

Mitchell, A. J., M. J. Salmon, D. G. Huffman, A. E. Goodwin, and T. M. Brandt. 2000. Prevalence and pathogenicity of a 

heterophyid trematode infecting the gills of an endangered fish, the fountain darter, in two central Texas spring-fed rivers. 

Journal of Aquatic Animal Health 12:283-289. 

Mkoji, G. M., B. N. Mungai, D. K. Koech, B. V. Hofkin, E. S. Loker, J. H. Kihara, and F. M. Kageni. 1992. Does the snail 

Melanoides tuberculata have a role in biological control of Biomphalaria pfeifferi and other medically important African 

pulmonates? Annals of Tropical Medicine and Parasitology 86: 201-204. 

Murray, H. D. 1964. Tarebia granifera and Melanoides tuberculatus in Texas. Bulletin of the American Malacological Union, 

1964:15-16. 

Murray, H. D. 1971. The introduction and spread of thiarids in the United States. The Biologist 53:133-135. 

Neck, R. W. 1985. Melanoides tuberculata (Thiaridae) in extreme southern Texas. Texas Conchologist 21:150-152. 

Pilsbry, H. A. and J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of 

Congo malacology. Bulletin of the American Museum of Natural History 52: 69-602. 

Pointier, J. P. 1993. The introduction of Melanoides tuberculata (Mollusca: Thiaridae) to the island of Saint Lucia (West Indies) 

and its role in the decline of Biomphalaria glabrata, the snail intermediate host of Schistosoma mansoni. Acta Tropica 

54:13-18. 

Pointier, J. P., A. Guyard, and A. Mosser. 1989. Biological control of Biomphalaria glabrata and B. straminea by the 

competitor snail Thiara tuberculata in a transmission site of schistosomiasis in Martinique, French West Indies. Annals of 

Tropical Medicine and Parasitology 83:263-269. 

Pointier, J. P., R. N. Incani, C. Balzan, P. Chroschiechowski, and S. Prypchan. 1994. Invasion of the rivers of the littoral central 

region of Venezuela by Thiara granifera and Melanoides tuberculata (Mollusca, Prosobranchia, Thiaridae) and the absence 

of Biomphalaria glabrata, snail host of Schistosoma mansoni. Nautilus 107:124-128. 

Pointier, J. P. and F. S. McCullough. 1989. Biological control of the snail hosts of Schistosoma mansoni in the Caribbean area 

using Thiara spp. Acta Tropica 46:147-155. 

Pointier, J. P., J. L. Toffart, and M. Lefevre. 1991. Life tables of freshwater snails of the genus Biomphalaria (B. glabrata, B. 

alexandrina, B. straminea) and of one of its competitors Melanoides tuberculata under laboratory conditions. Malacologia 

33:43-54. 

Roessler, M. A., G. L. Beardsley, and D. C. Tabb. 1977. New records of the introduced snail, Melanoides tuberculata (Mollusca: 

Thiaridae) in south Florida. Florida Scientist 40: 87-94. 

Russo, T. N. 1973. Discovery of the gastropod snail Melanoides (Thiara) tuberculata (Müller) in Florida. Florida Scientist 36: 

212-213. 

Wu, S.-K. 1989. Colorado freshwater mollusks. Natural History Inventory of Colorado 11: 1-117. 

*Table 1. Known records for Melanoides tuberculatus (Müller, 1774) in the United States. [Table 1 is too long to be included 

in this newsletter. Please email Tamara at tamander@onewest.net to request a copy (Excel format, ~32K)] 

17

Figure 1. Map of known occurrences of Melanoides tuberculatus (Müller, 1774) in the United States. 

Counties with occurrences are colored according to the first report of the species in that county. 

Phase I of Pennsylvania Aquatic Community Classification Complete 

Betsy Nightingale 1 , Mary Walsh 1 , D. David Homans 2 , Ryan Evans 2 , Jeremy Deeds 2 , and Emily Bond 3 

Pennsylvania Natural Heritage Program 

1 The Nature Conservancy, PA Science Office, 208 Airport Drive, Middletown, PA 17057, c-bnightin@state.pa.us 

2 Western Pennsylvania Conservancy, 209 4 th Ave., Pittsburgh, PA 15222, jdeeds@paconserve.org 

3 Michael Baker Corporation, 100 Airside Drive, Moon Township, PA 15108 

Pennsylvania Natural Heritage Program biologists have recently completed the pilot phase of a statewide classification of 

aquatic communities. In this preliminary classification, a variety of methods were used to classify stream communities of 

macroinvertebrates, fish and mussels in eight Pennsylvania watersheds. The complete Phase I report is available online 

(http://www.paconserve.org/rc/acp.html), and a summary of the mussel community classification results is provided here. 

Mussel and physical habitat data (adjacent land use, water chemistry, habitat quality, and stream hydrology) for Pennsylvania 

streams were acquired from more than 30 sources, principally from PA DEP, EPA EMAP, and The Nature Conservancy 

Freshwater Initiative. Data were also acquired from universities, museums, conservation groups, and individuals. Cluster 

Analysis, Indicator Species Analysis, and Nonmetric Multidimensional Scaling were used to classify communities of mussels, 

and Canonical Correspondence Analysis (CCA) was used to examine relationships among community types and physical 

habitat characteristics. 

18

Eight mussel community types were found in the pilot watersheds (Table 1). Four groups were specific to the Ohio River 

drainage, three specific to the Atlantic Drainage, and one (creeper community) had a small number of sites in both drainages. 

Generally, CCA did not show strong relationships between the occurrence of communities and physical habitat characteristics. 

However, some patterns were evident. The yellow lampmussel and creeper communities appeared to be weakly associated 

with increasing forest cover, and the Ohio lacustrine group showed a slight association with higher wetland coverage. There 

also appeared to be relationships between the occurrence of communities and water chemistry (alkalinity, dissolved oxygen, 

and pH), as well as stream characteristics (gradient and stream order). 

We continue to evaluate our methods as we look towards expanding our aquatic community classification statewide. We are 

still in the process of gathering feedback from reviewers, and welcome all comments and suggestions. 

Literature Cited 

Nedeau, E., M. McCullough, and B. Swartz. 2000. The freshwater mussels of Maine. Maine Department of Inland Fisheries 

and Wildlife, State House Station # 41, Augusta, Maine. 118 pp. 

Strayer, D.L. 1993. Macrohabitats of freshwater mussels (Bivalvia: Unionacea) in streams of the northern Atlantic Slope. 

Journal of North American Benthological Society 12(3): 236-246. 

Table 1. Mussel Communities found in the pilot classification. 

Mussel 

Community 

Eastern Elliptio 

community 

Yellow lampmussel 

community 

Brook floater 

community 

Ohio Backwater 

community 

Fatmucket 

community 

French Creek 

Mainstem 

community 

Ohio Lacustrine 

community 

Creeper community 

Significant Indicator Species Drainage Habitat Type 

Eastern Elliptio 

(Elliptio complanata) 

Yellow lampmussel (Lampsilis 

cariosa); Triangle floater 

(Alasmidonta undulata) 

Brook floater (Alasmidonta 

varicosa) 

Giant floater (Pyganodon 

grandis); White heelsplitter 

(Lasmigona complanata) 

Fatmucket (Lampsilis siliquoidea) 

19 

Atlantic 

Found in 2 nd to 7 th order streams across a range of 

land use types, stream orders, and temperatures 

Atlantic These species are often found in silt, sand, cobble, 

and gravel substrates in medium streams to large 

rivers (Nedeau et al. 2000). 

Atlantic Only two occurrences within the pilot watersheds; 

found in 3 rd & 4 th order streams; species may be 

associated with soft waters (Strayer 1993) 

Ohio Found in 3 rd and 4 th order streams along margins, 

backwaters, and reservoirs in sand, mud, and silt; 

both species pollution tolerant 

Ohio Found in 2 nd to 5 th order streams in sand, gravel 

and mud substrates 

15 significant species Ohio Found in 3 rd to 5 th order low-gradient streams, only 

in French Creek; shallow riffle and run areas with 

mixed cobble/gravel and sand substrates 

Spike (Elliptio dilatata); 

cylindrical papershell 

(Anodontoides ferussacianus) 

Creeper (Strophitus undulatus) 

Ohio Only found in 2 nd to 4 th order streams in the French 

Creek watershed 

Atlantic/ 

Ohio 

Strayer (1993) found this species to occur in 

hydrologically stable streams; We found only a 

weak correlation increasing forest cover.

Zoogeographical distribution on the freshwater mussels/naiades (Bivalvia: 

Unionoida & Veneroida) in Santa Catarina State's territory, Southern Brazil 

A. Ignacio Agudo 

Projeto Naiade (Naiade Project) 

Centro Integral de Educação Ambiental Cachoeira (CIEAC) (Integral Center of Environ. Education of the Waterfall), 

Caixa Postal (P. O. Box) 010, 88010-970 Centro, Florianópolis, Santa Catarina – SC, Brasil 

iagudo@lycos.com http://www.intergate.com.br/malacologia 

Keywords: Freshwater mussels – naiades; Unionoida; Veneroida; Continental malacological fauna survey; Santa Catarina 

state; Zoogeographical distribution; Southern Brazilian country. 

Starting from our first field registrations (Agudo 2002) to today with 20 registered species (18 natives, 2 invasive exotics), 

freshwater mussels/clams or naiades now represent 19% of the known mollusk continental fauna in the State of Santa Catarina 

(Agudo 2004b), and 43% of the regional freshwater representation (Agudo 2004 a, f). The distribution in the State's territory 

includes 28 hydrographical points in the Atlantic Coastal Plains (with 12 species for 16 localities) and the Uruguay River (with 

16 species for 6 localities) (Agudo 2004 b, c), with just 8 species occurring in both systems of basins (Agudo 2004 b, e). 

Bibliographical Antecedents 

1.- Sudamerican River Basins 

Parodiz & Bonetto (1963, pp. 188, 196) present the distribution of the Naiade families Mycetopodidae and Hyriidae in 

South América, including the geographical Brazilian territory of Santa Catarina´s State. 

Bonetto (1964) revised the distribution of genus Diplodon Spix, 1827 in the southern Atlantic coastal plains basin 

systems, into Brazil (State of Rio de Janeiro) and the country of Uruguay. 

2.- Atlantic Coastal Plains Basin System 

Poli et al. (1978, pp. 40-41) mention the occurrence of Leila sp from the South area of “Lagoa (Pond) of Peri”, in the 

Santa Catarina´s Island. 

Simone (1994, pp. 173-174, 180) mentions Anodontites trapesialis (Lamarck, 1819) from Joinville, Rio Itapoçu and 

"President Vargas" (?). 

Agudo (2002) relates Anodontites sp., Leila blanvilliana (Lea, 1834), Mycetopoda legumen (Martens, 1888), and 

Eupera klappenbachi Mansur & Veitenheimer, 1975 from the hidrographical portion corresponding to Atlantic 

coastal plain. 

3.- Uruguay River Basin System 

Callil & Mansur (2002, pp. 154-155) mention the exotic Corbicula largillierti (Philippi, 1844) for the Municipal 

Districts of Joaçaba & Piratuba (Rio do Peixe), Marcelino Ramos (Uruguay River), and Corbicula fluminea (Müller, 

1774) for the Municipal District of Concórdia (Jacutinga River). 

For the upper & middle sections of the Uruguay River Basin (Brazil and Argentina), 40 species are cited in the literature 

(Agudo 2004 d); of these, 14 are confirmed/included in this registration. The occurrence of Anodontites iheringi (Clessing, 

1882) and Diplodon pilsbryi Marshall, 1928 in the Santa Catarina´s territory increases the species total to 42. For the 

remainder, the occurrence of only 5 of the 14 cited species in the historical literature for the Southern Brazilian Atlantic 

Coastal Plains ( Bonetto 1964; Mansur 1970, pp. 66, 74) and the Iguazu (= Iguaçu) River Basin System in the neighboring 

State of Paraná, to the North (Morretes 1949, pp. 17-19, 25, 30; Zanardini 1965; Parodiz 1968, pp. 4, 10), previously revised in 

the specialized regional literature (Mansur 1970), are confirmed in this study (Agudo 2004 e). 

Finally, another invasive exotic species, the freshwater asiatic bivalve Limnoperna fortunei (Dunker, 1857), was recently 

profiled with potential occurrence in the territory of the State (Mansur et al 2004). Imminent entrance is awaited through the 

Uruguay & Iguazu River Basin Systems and, secondarily, in the “Itajaí River Basin” (through the Port of Itajaí, at the Itajaí- 

Açu River estuary), in the Atlantic Coastal Plain. 

Systematic Species List Regional Distribution 

Class PELECYPODA = BIVALVIA Atlantic Plains Uruguay River 

Order Unionoida 

Family MYCETOPODIDAE (8) 

Anodontites crispatus tenebricosus (Lea, 1834) X** X 

Anodontites ferrarisi (Orbigny, 1835) X 

Anodontites iheringi (Clessing, 1882) X 

Anodontites lucidus (Orbigny, 1835) X 

Anodontites trapesialis (Lamarck, 1819) X X 

20

Leila blainvilliana (Lea, 1834) X 

Monocondylaea minuana Orbigny, 1835 X 

Mycetopoda legumen (Martens, 1888) X X 

Family HIRIIDAE (7) 

Diplodon aethiops (Lea, 1860) X 

Diplodon ellipticus (Wagner in Spix, 1827) X*/*** X 

Diplodon expansus (Küster, 1856) X*/**/*** 

Diplodon hildae Ortmann, 1921 X 

Diplodon martensi (Ihering, 1893) X** X 

Diplodon pilsbryi Marshall, 1928 X 

Diplodon rhuacoicus (Orbigny, 1835) X* X 


Family CORBICULIDAE (2) 

Corbicula fluminea (Müller, 1774) X X 

Corbicula largillierti (Philippi, 1844) X X 

Family SPHAERIIDAE (3) 

Eupera klappenbachi Mansur & Veitenheimer, 1975 X 

Eupera platensis Doello-Jurado, 1921 X 

Sphaerium observationis (Pilsbryi, 1911) X 

*Bonetto (1964); **Zanardini (1965); ***Mansur (1970) 

References 

Agudo, A. Ignacio. 2002. Preliminary report about the freshwater mussels/clams (Bivalvia: Unionoida & Veneroida) of Santa 

Catarina State, Southern Brazil. FMCS Newsletter Ellipsaria 4(2):10-11. 

Agudo, A. Ignacio. 2004a. The freshwater mollusks fauna of Santa Catarina State, Southern Brazil: a comprehensive general 

synthesis. FMCS Newsletter Ellipsaria 6(1):11-12. 

Agudo, A. Ignacio. 2004b. Levantamento Atualizado – Espécies em Santa Catarina (Up-to-date rising – Species in SC). 

Florianópolis, SC: AVULSOS MALACOLÓGICOS – AM, Internet Brazilian Website: 

http://www.intergate.com.br/malacologia/levantamento/levantamento.html 

Agudo, A. Ignacio. 2004c. Projeto Naiade (Naiade Project). Florianópolis – SC: AVULSOS MALACOLÓGICOS – AM, 

Internet Brazilian WebSite: http://www.intergate.com.br/malacologia/projetonaiade/projetonaiade.html 

Agudo, A. Ignacio. 2004d. Bivalves Naiades do Medio e Alto Rio Uruguai. In: Boletim AM Especial no. 37-38A, Ano IV, 

Fevereiro-Março 2004. Florianópolis – SC: AVULSOS MALACOLÓGICOS – AM, Internet Brazilian WebSite: 

http://www.intergate.com.br/malacologia/boletins/boletins.html 

Agudo, A. Ignacio. 2004e. Bivalves Naiades do Rio Iguaçu & Estado do Paraná. In: Boletim AM no. 40, Ano IV, Maio 2004. 

Florianópolis – SC: AVULSOS MALACOLÓGICOS – AM, Internet Brazilian WebSite: 

http://www.intergate.com.br/malacologia/boletins/boletins.html 

Agudo, A. Ignacio. 2004f. Additional new records of freshwater mollusks (Gastropoda & Bivalvia) from Santa Catarina State, 

Southern Brazil, with complementary observations about mollusks fauna from the Mampituba river basin. FMCS 

Newsletter Ellipsaria 6(2):this issue 

Bonetto, Argentino A. 1964. Las espécies del género Diplodon (MOLL. UNIONACEA) en los ríos de la Pendiente Atlántica 

del Sur del Brasil. PHYSIS 24(68):323-328. 

Callil, C. T. & M. C. Dreher Mansur. 2002. Corbiculidae in the Pantanal: history of invasion in southeast and central South 

America and biometrical data. Amazoniana 17(1/2):153-167. 

Mansur, Maria Cristina Dreher. 1970. Lista dos moluscos bivalves das familias Hyriidae e Mycetopodidae para o Estado do 

Rio Grande do Sul. Iheringia (Porto Alegre), Sér. Zool. (39):33-95. 

Mansur, Maria Cristina Dreher; Cassiano Barboza Quevedo, Cíntia Pinheiro dos Santos & Cláudia Tasso Callil. 2004. 

Prováveis Vías da Introdução de Limnoperna fortunei (Dunker, 1857) (Mollusca, Bivalvia, Mytilidae) na Bacia da Laguna 

dos Patos, Rio Grande do Sul e Novos Registros de Invasão no Brasil pelas Bacias do Paraná e Paraguai (Cap. 4, pp. 35- 

38). In: Silva, Julieta Salles Vianna da & Rosa Cristina Corrêa Luz de Souza (Orgs.), Água de Lastro e Bioinvasão. Rio de 

Janeiro, RJ: Editora Interciência, 224 p. Morretes, Frederico Lange de. 1949. Ensaio de Catálogo dos Moluscos do Brasil. 

Curitiba, PR: Arquivos do Museu Paranaense, Vol. VII, art. 1, pp. 5-216. 

Parodiz, J. J. 1968. Annotated catalogue of the genus Diplodon (UNIONACEA – HYRIIDAE). Sterkiana, (30): 1-22. 

Parodiz, J. J. & A. A. Bonetto. 1963. Taxonomy and Zoogeographic relationships of the South American Naiades (Pelecypoda: 

Unionacea and Mutelacea). Malacologia 1 2):179-213. 

Poli, Carlos Rogério; Fernando Sniziek & Paulo F. de Araújo Lago. 1978. Levantamento da fauna aquática da Lagoa do Peri. 

Florianópolis, SC: CCA/UFSC – IPUF, Relatório Técnico, 68 p. 

Simone, Luiz Ricardo Lopes de. 1994. Anatomical characters and systematics of Anodontites trapesialis (Lamarck, 1819) from 

South America (Mollusca, Bivalvia, Unionoida, Muteloidea). Studies on Neotropical Fauna and Environment, 29(3):169- 

185. 

21

FMCS 2003 Freshwater Mollusk Bibliography 

Compiled by Kevin S. Cummings 

Illinois Natural History Survey, Champaign, Illinois 

This bibliography lists freshwater mollusk papers that have 

been published up to and including 2003 and that have not 

appeared in previous FMCS bibliographies. Citations are 

split into five groups: Unionoida, Sphaeriidae, Corbiculidae, 

Dreissenoidea (and other bivalves), and Gastropoda. Papers 

that list taxa from more than one category are included in 

each group. A searchable database of over 14,300 

references on freshwater mollusks is available on the web at 

http://ellipse.inhs.uiuc.edu:591/mollusk/ To insure that 

papers are cited correctly, researchers are encouraged to 

send reprints to: Kevin S. Cummings, Illinois Natural 

History Survey, 607 E. Peabody Drive, Champaign, Illinois 

61820 or ksc@inhs.uiuc.edu 

UNIONOIDA (FRESHWATER MUSSELS) 

Aboul-Dahab, H.M. 2001. Ecological and histophathological 

[histopathological] studies on two freshwater mussels 

infected by the trematode Aspidogaster niloticus with 

reference to the host-parasite relationships. Egyptian 

Journal of Zoology 36:49-65. 

Aboul-Dahab, H.M. 2001. Host-parasite interactions between 

the molluscan bivalve Caelatura aegyptiaca and a 

haplosporodian [haplosporidian] parasite. Egyptian Journal 

of Zoology 37:295-314. 

Aboul-Dahab, H.M. 2002. Cellular responses of the 

molluscan host, Unio abyssinicus infected by two new 

haplosporidian parasites with emphasis on the 

morphological characteristics of the host haemolymph. 

Egyptian Journal of Zoology 38:149-170. 

Abraszewska-Kowalczyk, A. 2002. Unionid bivalves of the 

Pilica river catchment area. Folia Malacologica 10(3):99- 

173. 

Afanasjev, S.A., B. Zdanowski, and A. Kraszewski. 2001. 

Growth and population structure of the mussel Anodonta 

woodiana (Lea, 1834) (Bivalvia, Unionidae) in the heated 

Konin Lakes system. Archiwum Rybactwa Polskiego 

9(1):123-131. 

Aldridge, D.C., and A.L. McIvor. 2003. Gill evacuation and 

release of glochidia by Unio pictorum and Unio tumidus 

(Bivalvia: Unionidae) under thermal and hypoxic stress. 

Journal of Molluscan Studies 69(1):55-59. 

Allen, G.T., S.H. Blackford, V. Tabor, and M.S. Cringan. 

2001. Metals, boron, and selenium in Neosho madtom 

habitats in the Neosho River in Kansas, U.S.A. 

Environmental Monitoring and Assessment. 61:1-21. 

Altaba, C.R. 2000. La ultima oportunidade de Margaritifera 

auricularia nuestro bivalvo de agua dulce mas amenazado. 

[The last chance for Margaritifera auricularia our most 

threatened freshwater bivalve.] Quercus 170:16-23. 

Altaba, C.R., and J. Lascurain. 2001. Una poblacio 

supervivent del genere Anodonta (Bivalvia: Unionoida) a la 

conca de la Tordera. [A surviving population of the genus 

Anodonta (Bivalvia: Unionoida) in the Tordera drainage.] 

Butlleti de la Institucio Catalana d'Historia Natural 69:71- 

76. 

22 

Altaba, C.R., and M.A. Lopez. 2001. Experimental 

demonstration of viability for the endangered giant 

pearlmussel Margaritifera auricularia (Bivalvia: 

Unionoida) in its natural habitat. Bolleti de la Societat 

d'Historia Natural de les Balears 44:15-21. 

Alvarez-Halcon, R.M., R. Araujo, and G. Delvene. 2000. 

Margaritifera auricularia, un bivalvo de agua dulce 

amenazado en Aragon. [Margaritifera auricularia, an 

endangered softwater bivalve in Aragon.] Naturaleza 

Aragonesa 5:29-37. 

Angelo, R.T., and M.S. Cringan. 2003. Rediscovery of the 

black sandshell, Ligumia recta (Bamarck, 1819), in Kansas. 

Transactions of the Kansas Academy of Science 106(3- 

4):111-113. 

Anon. 2003. Moving mussels. Wildlife in North Carolina 

67(2):41. 

Antunes, C., T. Magalhaes-Cardoso, G. Moura, D. Goncalves, 

and J. Machado. 2002. Effects of Al, Ni, Co, Zn, Cd, and 

Cu metals on the outer mantle epithelium of Anodonta 

cygnea (Unionidae). Haliotis 31:71-84. 

Araujo, R. 1999. Bivalvos de agua dulce de la peninsula 

Iberica. [Freshwater bivalves from the Iberian Peninsula.] 

Quercus 160:26-30. 

Araujo, R., and A. Ramos. 2001. Action plans for 

Margaritifera auricularia and Margaritifera margaritifera 

in Europe. Council of Europe Nature and Environment 

Series 117:1-66. 

Araujo, R., M. Quiros, and M.A. Ramos. 2003. Laboratory 

propagation and culture of juveniles of the endangered 

freshwater mussel Margaritifera auricularia (Spengler, 

1793). Journal of Conchology 38(1):53-60. 

Augspurger, T., A.E. Keller, M.C. Black, W.G. Cope, and F.J. 

Dwyer. 2003. Water quality guidance for protection of 

freshwater mussels (Unionidae) from ammonia exposure. 

Environmental Toxicology and Chemistry 22(11):2569- 

2575. 

Auti, R.G., J.M. Patwari, and S.S. Patil. 2001. Acute toxicity 

of mercury on the fresh water bivalve molluscs, Lamellidens 

corrianus (Lee) from Nandrabad Pond M.S. Dr Babasaheb 

Ambedkar Marathwada University Journal of Science 

31(8):1-5. 

Bába, K., D. Sári, and T. Tóth. 2002. Bivalve biocoenoses on 

different substrates, the 2000-year heavy metal 

contamination. Soosiana 23:37-48. 

Baer, O. 2002. Die Vorkommen von Unio pictorum (L.) in 

der Elbe bei Dresden (Bivalvia: Unionidae). [The Elbe river 

population of Unio pictorum (L.) near Dresden (Bivalvia: 

Unionidae).] Malakologische Abhandlungen-(Dresden) 

20(2):357-366. 

Baker, A.M., C. Bartlett, S.E. Bunn, K. Goudkamp, F. 

Sheldon, and J.M. Hughes. 2003. Cryptic species and 

morphological plasticity in long-lived bivalves (Unionoida: 

Hyriidae) from inland Australia. Molecular Ecology 

12(10):2707-2717. 

Baker, S.M., and J.S. Levinton. 2003. Selective feeding by 

three native North American freshwater mussels implies 

food competition with zebra mussels. Hydrobiologia 

505(1-3):97-105.

Barnhart, M.C. 2000. Fish hosts and culture of mussel 

species of special concern. Report to U.S. Fish and Wildlife 

Service and Missouri Department of Conservation 39 pp. 

Barnhart, M.C. 2000. Neosho mucket restoration project. 

Kansas Pearly Mussel Newsline 5:1-2. 

Barnhart, M.C. 2001. Fish hosts and culture of mussel 



Barnhart, M.C. 2001. Venustaconcha in the Spring River are 

something special. Kansas Pearly Mussel Newsline 6:1-2. 

Barnhart, M.C. 2002. Propagation and culture of mussel 



Barnhart, M.C. 2003. Culture and restoration of mussel 

species of concern. Report to U.S. Fish and Wildlife 


Barsiene, J., and D. Barsyte-Lovejoy. 2000. Environmental 

genotoxicity in Klaipeda port area. International Review of 

Hydrobiology 85(5-6):663-672. 

Barsiene, J., R. Bucinskiene, and K. Joksas. 2002. 

Cytogenetic damage and heavy metal bioaccumulation in 

molluscs inhabiting different sites of the Neris River. 

Ekologija 2:52-57. 

Bartsch, M.R., T.J. Newton, J.W. Allran, J.A. O'Donnell, and 

W.B. Richardson. 2003. Effects of pore-water ammonia on 

in situ survival and growth of juvenile mussels (Lampsilis 

cardium) in the St. Croix Riverway, Wisconsin, USA. 


2568. 

Baskakov, A.V., A.V. Polevshchikov, and A.D. Kharazova. 

2000. [An evaluation of the ligand specificity of 

haemagglutinins and haemolysins in the haemolymph of 

gastropod and bivalve molluscs.] Zhurnal Evolyutsionnoi 

Biokhimii i Fiziologii 36(4):281-285. 

Baskay, I., Z. Dobo, and B. Penzes. 1996. Az amuri kagylo 

okozta glochidiozis vizsgalata. [Glochidiosis caused by the 

mussel, Anodonta woodiana woodiana Lea, 1834.] 

Allattani Kozlemenyek 81:9-14. 

Beasley, C.R., L.D. Miranda, S.T.M. Alves, and C.H. 

Tagliaro. 2003. Evaluation of methods for assessing brood 

size in freshwater mussels (Hyriidae). Journal of Molluscan 

Studies 69(2):159-161. 

Beck, K., and R.J. Neves. 2003. An evaluation of selective 

feeding by three age-groups of the rainbow mussel Villosa 

iris. North American Journal of Aquaculture 65:203-209. 

Begum, A., and S.I. Khan. 2001. Abundance of Vibrio 

cholerae non-01 and Vibrio mimicus in various aquatic 

components of the pond ecosystem in Matlab, Bangladesh. 

Microbes and Environments 16(1):19-23. 

Begum, S., and A.K. Sinha. 2000. Length-weight relationship 

of Parreysia favidens of freshwater drainage systems of 

Burhi Gandak, north Bihar (India). Records of the 

Zoological Survey of India 98(3):61-66. 

Beran, L., and M. Horsak. 1999. Mollusca. Folia Facultatis 

Scientiarum Naturalium Universitatis Masarykianae 

Brunensis Biologia 101:79-87. 

Bergbauer, M. 2001. Kaltwasserfische. Der Bitterling hat ein 

aussergewoehnliches Fortpflanzungsverhalten entwickelt. 

[Cold water fish. The bitterling has developed an 

23 

extraordinary reproductive strategy.] Aquarium 

(Bornheim) 381:20-23. 

Bergman, S.M., M.E. Eberle, and B.K. Obermeyer. 2000. 

Freshwater mussels (Bivalvia: Unionoidea) in streams of 

northwestern Kansas. Prairie-Naturalist 32(1):1-15. 

Bhamre, P.R., A.E. Desai, and B.M. Deoray. 2001. Effect of 

mercuric chloride on glycogen content of the freshwater 

mussel Parreysia favidens. Pollution Research 20(1):13- 

15. 

Bigham, S. 2002. Host specificity of freshwater mussels: a 

critical factor in conservation. MS Thesis, Southwest 

Missouri State University 

Biswas, A., and S.K. Raut. 2000. Bioindicators for 

freshwater mussel, Lamellidens marginalis (Lamarck) 

farming. Pp. 38-53 in Aditya, A.K.; Haldar, P. [Eds]. 

Biodiversity and environment: Proceedings of the National 

Seminar on Environmental Biology, April 03-05, 1998. 

Daya Publishing House, Delhi. i-xiii, 1-249. 

Blaise, C., F. Gagne, M. Salazar, S. Salazar, S. Trottier, and 

P.D. Hansen. 2003. Experimentally-induced feminisation 

of freshwater mussels after long-term exposure to a 

municipal effluent. Fresenius Environmental Bulletin 

12(8):865-870. 

Blaise, C., S. Trottier, F. Gagne, C. Lallement, and P.D. 

Hansen. 2002. Immunocompetence of bivalve hemocytes 

as evaluated by a miniaturized phagocytosis assay. 

Environmental Toxicology 17(3):160-169. 

Blanchard, M., M.J. Teil, A.M. Carru, B. Garban, D. Ollivon, 

A. Chesterikoff, and M. Chevreuil. 1999. Biota 

contamination by PCBs and trace metals in the freshwater 

estuary of the River Seine (France). Hydrobiologia 

400:149-154. 

Boessneck, U. 2001. Extensiv genutzte Fischteiche als 

Refugial-Lebensraeume fuer hochgradig bestandsbedrohte 

Arten - der Juteteich bei Triebes (Lkr. Greiz). [Extensively 

used fish ponds as refuges for highly endangered species - 

the Jute pond near Triebes (Gera District).] 

Landschaftspflege und Naturschutz in Thueringen 

38(2):50-54. 

Boessneck, U. 2002. Lebendnachweise von Microcondylaea 

bonellii (A. Ferussac 1827) und Pisidium tenuilineatum 

Stelfox 1918 aus Istrien (Kroatien) (Bivalvia: Unionidae et 

Sphaeriidae). [Recent records of living Microcondylaea 

bonellii (A. Ferussac 1827) and Pisidium tenuilineatum 

Stelfox 1918 from Istria (Croatia) (Bivalvia: Unionidae et 

Sphaeriidae).] Malakologische Abhandlungen-(Dresden) 

20(2):313-317. 

Bogatov, V.V. 2000. The first of Middendorffinaia (Bivalvia, 

Unionoidea) in Okhotsk region. Zoologicheskii Zhurnal 

79(7):861-862. 

Bogatov, V.V. 2001. New data on Unioniformes of Sakhalin 

Island. Byulleten' Dal'nevostochnogo Malakologicheskogo 

Obshchestva 5:71-77. 

Bogatov, V.V., and -Y.I. Starobogatov. 2001. Anodontinae 

(Bivalvia) of the genus Beringiana. Zoologicheskii Zhurnal 

80(1):26-31. 

Bogatov, V.V., and E.M. Sayenko. 2002. The history of 

investigation of Anodontinae and Pseudanodontinae in the 

Russian Far East. Byulleten' Dal'nevostochnogo 

Malakologicheskogo Obshchestva 6:102-112.

Bogatov, V.V., E.M. Saenko, and Ya.I. Starobogatov. 2002. 

On taxonomic position of the genus Kunashiria (Bivalvia, 

Unioniformes). Zoologicheskii Zhurnal 81(5):521-528. 

Bogatov, V.V., L.A. Prozorova, and -Y.I. Starobogatov. 2003. 

The family Margaritiferidae (Mollusca: Bivalvia) in Russia. 

Ruthenica 13(1):41-52. 

Bolden, S.R., and K.M. Brown. 2002. Role of stream, 

habitat, and density in predicting translocation success in the 

threatened Louisiana pearlshell, Margaritifera hembeli 

(Conrad). Journal of the North American Benthological 

Society 21(1):89-96. 

Brim-Box,-J., R.M. Dorazio, and W.D. Liddell. 2002. 

Relationships between streambed substrate characteristics 

and freshwater mussels (Bivalvia: Unionidae) in Coastal 

Plain streams. Journal of the North American Benthological 

Society 21(2):253-260. 

Bucinskiene, R. 2001. Studies of aneugenic effects in 

molluscan tissues. Ekologija 3:23-27. 

Burch, J.B. 2003. Virginia’s James River basin project. 

Walkerana 12(29-30):109-112. 

Burch, P.R. 2003. Mollusks. [Reprinted - a preliminary list 

of the land and freshwater mollusks of the James River 

Basin, Virginia]. Walkerana 12(29-30):113-122. 

Burton, K. 2003. Sending surrogates to the rescue. 

Endangered Species Bulletin 28(2):22-23. 

Butler, R.S., and R.G. Biggins. 2003. Agency draft recovery 

plan for Cumberland elktoe, oyster mussel, Cumberland 

combshell, purple bean, and rough rabbitsfoot. U.S. Fish & 

Wildlife Service, Atlanta, Georgia. 176 pp. 

Butler, R.S., R. Kessler, and J.B. Harrel. 2003. Down by the 

Green River. Endangered Species Bulletin 28(2):20-21. 

Buttstedt, L. 2000. Ein aktuelles Vorkommen der 

Bachmuschel im suedwestlichen Sachsen-Anhalt. [The 

current distribution of the brook mussel in south west 

Saxony-Anhalt.] Naturschutz im Land Sachsen Anhalt 

37(2):57-60. 

Byrne, M. 2000. Calcium concretions in the interstitial 

tissues of the Australian freshwater mussel Hyridella 

depressa (Hyriidae). Geological Society Special 

Publication 177:329-337. 

Carpentier, A., J.M. Paillisson, L. Marion, E. Feunteun, A. 

Baisez, and C. Rigaud. 2003. Trends of a bitterling 

(Rhodeus sericeus) population in a man-made ditch 

network. Comptes Rendus Biologies 326(Suppl 1):S166- 

S173. 

Carretero, M.A. C.R. Altaba, E. Bassols, S. Giraldez, and C. 

Rosell. 2000. El plan de recuperacion de la nayade Unio 

aleroni en la comarca catalana de la Garrotxa. [A 

recuperation plan for Unio aleroni in the Catalan district of 

Garrotxa.] Quercus 170:24-30. 

Cartwright, C. 2003. Unusual use of freshwater mussel (Unio 

sp.) shells during the Early Bronze Age at Tell es-Sa'idiyeh, 

Jordan. Environmental Archaeology 8(1):85-89. 

Cencetti, E., and L. Castagnolo. 1997. Systematica e 

distribuzione di Microcondylaea compressa Menke, 1828 

(Bivalvia: Unionidae), una specie ormai rara in Italia. Nota 

breve. [Systematic and distribution of Microcondylaea 

compressa Menke, 1828 (Bivalvia: Unionidae), a rare 

species in Italy. A brief note.] Quaderni ETP 26:115-117. 

24 

Checa, A. 2000. A new model for periostracum and shell 

formation in Unionidae (Bivalvia, Mollusca). Tissue and 

Cell 32(5):405-416. 

Checa, A.G., and A. Rodriguez-Navarro. 2001. Geometrical 

and crystallographic constraints determine the selforganization 

of shell microstructures in Unionidae 

(Bivalvia: Mollusca). Proceedings Royal Society of London 

Series B Biological Sciences 268(1468):771-778. 

Chen, -Jing-Chun, and Shi, -An-Jing. 1998. Cytological 

observation on the fertilization of Anodonta woodiana 

elliptica. Journal of Fisheries of China 22(1):78-80. 

Chen, -Jing- Chun, and Shi, -An-jing. 2002. Study on 

oogenesis of Anodonta woodiana elliptica (Heude). 

Sichuan Daxue Xuebao (Ziran-Kexueban) 39(3):546-551. 

Chen, -Jinhua, and Tanaka, -Hitoshi. 2001. Jiangxiella, an 

Upper Triassic non-marine bivalve, found from southwest 

Japan. Dizhi Xuebao. 75(4):446-450. 

Cheney, M.A., K. Birdsall, and J.J. Kukor. 2001. Uptake and 

diverse effects of polycyclic aromatic hydrocarbons on the 

metabolic activity of Elliptio complanata measured by 

calorespirometry. Environmental Toxicology and 

Chemistry 20(5):1031-1036. 

Chernomaz, T.V. 2001. Ciliary activity of outliving cells of 

the glimmeral epithelium of gill and leg of Unionidae 

invaded by the trematodes Aspidogaster conchicola and 

Bucephalus polymorphus. Parazitologiya (St.-Petersburg) 

35(5):443-448. 

Chernomaz, T.V. 2002. Ciliary activity of the glimmeral 

epithelium of gill and leg Unio conus (in vitro) as testfunction 

at ecologo-physiological investigations. Visnyk 

Zhytomyrskoho Pedahohichnoho Universytetu 10:159-162. 

Chittick, B., M. Stoskopf, M. Law, R, Overstreet, and J. 

Levine. 2001. Evaluation of potential health risks to 

eastern elliptio (Elliptio complanata) (Mollusca: Bivalvia: 

Unionida: Unionidae) and implications for sympatric 

endangered freshwater mussel species. Journal of Aquatic 

Ecosystem Stress and Recovery 9(1):35-42. 

Chu, S.G., A. Covaci, S. Voorspoels, and P. Schepens. 2003. 

The distribution of octachlorostyrene (OCS) in 

environmental samples from Europe. Journal of 

Environmental Monitoring 5(4):619-625. 

Cicerello, R.R., and G.A. Schuster. 2003. A guide to the 

freshwater mussels of Kentucky. Kentucky State Nature 

Preserves Commission Scientific & Technical Series 

Number 7. 62 pp. 

Cisotto, P. 2003. Primo rinvenimento in Italia di Anodonta 

suborbiculata Say, 1831 e di Anodonta grandis Say, 1829. 

First record in Italy of Anodonta suborbiculata Say, 1831 

and Anodonta grandis Say, 1829. Societa Veneziana di 

Scienze Naturali Lavori 28:11-15. 

Cochet, G. 2000. Impact des amenagements des cours d'eau 

et des activites agrosylvicoles sur les populations francaises 

de Margaritifera margaritifera. Proposition pour une 

reconquete des cours d'eau par l'espece. [Impact of the 

planning of rivers and the agricultural and forestry activities 

on the French population of Margaritifera margaritifera. 

Proposal for a reconquest of the rivers by the species.] 

Council of Europe Environmental Encounters Series 

45:123-127.

Cope, W.G. M.C. Hove, D.L. Waller, D.J. Hornbach, M.R. 

Bartsch, L.A. Cunningham, H.L. Dunn, and A.R. 

Kapuscinski. 2003. Evaluation of relocation of unionid 

mussels to in situ refugia. Journal of Molluscan Studies 

69(1):27-34. 

Copley, L., and T.K. McCarthy. 2001 The first record of the 

monogenean gill fluke Pseudodactylogyrus bini (Kikuchi 

1929) in Ireland, with observations on other ectoparasites of 

River Erne eels. Irish Naturalists' Journal 26(11):405-413. 

Cordeiro, J.R. 2003. Case 3223. Unio ochraceus Say, 1817 

(currently Ligumia ochracea; Mollusca, Bivalvia): Proposed 

precedence of the specific name over Mytilus fluviatilis 

Gmelin, 1791. Bulletin of Zoological Nomenclature 

60(1):20-22. 

Cosgrove, P.J., and L.C. Hastie. 2001. Conservation of 

threatened freshwater pearl mussel populations: river 

management, mussel translocation and conflict resolution. 

Biological Conservation 99(2):183-190. 

Cummings, K.S., and R.V. Anderson. 2003. Freshwater 

mussels: “aquatic canaries in the coal mine.” Wetland 

Matters. Newsletter of the Wetlands Initiative 8(2):1-7. 

Danilin, I.A., B. Synzynys, G.V. Koz'min, and G.M. Rott. 

2002. Experimental substantiation of a new method for 

biotesting freshwater bodies by determining metallothionein 

contents in the organs and tissues of bivalve mollusks. 

Ekologiya (Moscow) 5:397-400. 

Das, S., and B.B. Jana. 2003. In situ cadmium reclamation by 

freshwater bivalve Lamellidens marginalis from an 

industrial pollutant-fed river canal. Chemosphere 

52(1):161-173. 

Davis, L.G. and K. Muehlenbachs. 2001. A Late Pleistocene 

to Holocene record of precipitation reflected in 

Margaritifera falcata shell [delta]18O from three 

archaeological sites in the Lower Salmon River Canyon, 

Idaho. Journal of Archaeological-Science 28(5):291-303. 

Delp, A. 2002. Rhabdocoel flatworms as predators of 

juvenile freshwater mussels. MS Thesis, Southwest 


Deng, -Dao-gui, and Li, -Rui-zhi. 2002. Histological and 

scanning electron microscopic observations on the gill of 

Cristaria plicata. Journal of Fisheries of China 26(4):301- 

306. 

Deng, -Dao-gui, and Tan, -Qi-kun. 2001. Studies on 

ultrastructure of the sperm of freshwater mussel, Cristaria 

plicata. Acta Hydrobiologica Sinica 25(5):481-485. 

Dettman, D.L., M.J. Kohn, J. Quade, F.J. Ryerson, T.P. Ojha, 

and S. Hamidullah. 2001. Seasonal stable isotope evidence 

for a strong Asian monsoon throughout the past 10.7 m.y. 

Geology (Boulder) 29(1):31-34. 

Diamond, J.M., and V.B. Serveiss. 2001. Identifying sources 

of stress to native aquatic fauna using a watershed 

ecological risk assessment framework. Environmental 

Science and Technology 35(24):4711-4718. 

Diamond, J.M., D.W. Bressler, and V.B. Serveiss. 2002. 

Assessing relationships between human land uses and the 

decline of native mussels, fish, and macroinvertebrates in 

the Clinch and Powell River watershed, USA. 


1155. 

25 

Dickson, T.L. 2000. A comparison of stream segment and 

quadrat mussel sampling techniques. Bios 71(2):42-49. 

Didonato, G.T., and A.E. Stiven. 2000. Three year variation 

in shell growth of the mussel, Elliptio waccamawensis 

(Lea), in Lake Waccamaw, a Bay Lake in North Carolina. 

Journal of the Elisha Mitchell Scientific Society 

116(4):324-333. 

Diggins, T.P., and R.J. Snyder. 2003. Three decades of 

change in the benthic macroinvertebrate community and 

water quality in the Buffalo River area of concern, 1964- 

1993. Journal of Great Lakes Research 29(4):652-663. 

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Bandoni, T. Wilke, D. Minchella, F. Lewis, and M. Knight. 

2003. Use of microsatellite variation and RAPD-PCR to 

assess genetic polymorphism in Biomphalaria glabrata 

snails from a single locale in a schistosomiasis endemic 

area. Malacologia 45(1):149-166. 

Schneck, J.L., B. Fried, and J. Sherma. 2003. Thin layer 

chromtographic analysis of neutral lipids and phospholipids 

in Helisoma trivolvis (Colorado Strain) maintained on a 

high fat diet. Veliger 46(4):325-328. 

Schreiber, E.S.G., G.P. Quinn, and P.S. Lake. 2003. 

Distribution of an alien aquatic snail in relation to flow 

variability, human activities and water quality. Freshwater 

Biology 48(6):951-961. 

Sivaramakrishna, B., and K. Radhakrishnaiah. 2000. 

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Szentesi, G., and T. Domokos. 1998. A Sr-90 izotop 

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Takaichi, S., V. Mizuhira, H. Hasegawa, T. Suzaki, M. 

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Revista de Biología Tropical 51(Suppl. 1):1-287. 

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Freshwater Mollusk Conservation Society Membership List - August 2004 

Please contact Rita Villella, FMCS secretary, with any corrections. 

Jae Abel 

Santa Clara Valley Water District 

310 Victoria Place 

Palo Alto, CA 94306 

(408) 265-2607 x 2655 

jaeabel@valleywater.org 

Steve Ahlstedt 

P.O. Box 460, 57 Deer Ridge Road 

Norris, TN 37828 

(865) 545-4140 x 17 


Brett Albanese 

Georgia Natural Heritage Program 

2117 U.S. Highway 278 S.E. 

Social Circle, GA 30677 

(770) 557-3032 

brett_albanese@dnr.state.ga.us 

John Alderman 

244 Red Gate Road 

Pittsboro, NC 27312 

(919) 542-5331 

aldermjm@mindspring.com 

Dan Allen 

Minnesota DNR 

3805 Bryant Ave. S., Apt. #3 

Minneapolis, MN 55409 

Richard Anderson 

Western Illinois Univ., Dept. Biol. Sci. 

Waggoner Hall 381 

Macomb, IL 61455 

(309) 298-2408 

r-anderson1@wiu.edu 

Robert Anderson 

USFWS 

315 South Allen Street, Suite 322 

State College, PA 16801 

(814) 234-4090 

robert_m_anderson@fws.gov 

Tamara Anderson 

285 Smith Street 

Lander, WY 82520 

(307) 335-8662 

tamander2002@yahoo.com 

Rafael Araujo 

Museo Nacional Ciencias Naturales 

Jose Gutiérreaz Abascal 2 

Madrid 28020 

Spain 

34-916111328 

rafael@mncn.csic.es 

Elizabeth Ashcraft 

P.O. Box 444 

Horn Lake, MS 38637 

(662) 393-0878 

eashcraft@astate.edu 

Herb Athearn 

Museum of Fluvatile Mollusks 

5819 Benton Pike NE 

Cleveland, TN 37323-5301 

(423) 476-4963 

James Atkinson 

Dept. Zoology 

Michigan State University 

East Lansing, MI 48824-1115 

(517) 353-2269 

atkinso9@msu.edu 

Peter Badra 

MI Natural Features Inventory 

Mason Building, P.O. Box 30444 

Lansing, MI 48909-7944 

(517) 241-4179 

badrap@michigan.gov 

Raynie Bambarger 

Louisiana State University 

383 Life Science Bldg. 

Baton Rouge, LA 70803 

(225) 578-1740 

abamba1@lsu.edu 

Lee Barclay 

USFWS 

446 Neal Street 

Cookeville, TN 38501 

(931) 528-6481 x 212 

lee_barclay@fws.gov 

Meredity Bartron 

USFWS 

PO Box 75 

Lamar, PA 16848 

(570) 726-4995 

meredith_bartron@fws.gov 

47 

Bruce Bauer 

BHE Environmental, Inc. 

7041 Maynardville Highway 

Knoxville, TN 37918 

(865) 922-4305 

bbauer@bheenvironmental.com 

Braven Beaty 

The Nature Conservancy 

146 E. Main Street 

Abingdon, VA 24210 

(540) 676-2209 

bbeaty@tnc.org 

Chris Bedel 

19 Abner Hollow Road 

Lynx, OH 45650 

(937) 544-2880 

eoa@bright.net 

David Berg 

Dept. of Zoology, Miami University 

Oxford, OH 45056 

(513) 529-3174 

bergdj@muohio.edu 

Randall Bernot 

University of Notre Dame 

107 Galvin Life Science Center 

Dept. of Biological Sciences 

Notre Dame, IN 46556-0369 

rbernot@nd.edu 

Richard Biggins 

55 Pyfrom Drive 

Swannanoa, NC 28778 

(828) 299-9128 

rgbiggins@aol.com 

Holly Blalock-Herod 

USFWS 

1601 Balboa Ave. 

Panama City, FL 32405 

(850) 769-0552 x 239 

holly_blalock_herod@fws.gov 

Juan Felipe Blanco 

Dept. of Biology 

University of Puerto Rico 

Rio Piedras Campus 

FB-241, University Station 

PO Box 23360 

San Juan, PR 00931-3360 

(787) 565-7531 

jblanco@lternet.edu

Arthur Bogan 

NC State Museum Natural Sciences 

4301 Reedy Creek Road 

Raleigh, NC 27607 

(919) 733-7450 x 753 

arthur.bogan@ncmail.net 

Susan Bolden 

Yale University 

FES - Greeley Lab 

370 Prospect Street 

New Haven, CT 06511 

(203) 432-5321 

susan_bolden@yale.edu 

William Bouthillier 

Georgia Dept. of Transportation 

3993 Aviation Circle 

Atlanta, GA 30336 

(404) 699-4309 

william.bouthillier@dot.state.ga.us 

Bonnie Bowen 

Iowa State University 

Dept. Ecology, Evolution & Organismal 

Biology 

339 Science II 

Ames, IA 50011 

(515) 294-6391 

bsbowen@iastate.edu 

Julie Boyles 

US Fish and Wildlife Service 

400 E. Main Street 

White Sulphur Springs, WV 24986 

(304) 536-1361 

julie_boyles@fws.gov 

David Braatz 

Streamside Systems 

P.O. Box 245 

Boonville, NC 27011 

(704) 682-1434 

dabljj@ix.netcom.com 

Tony Brady 

Genoa Fish Hatchery 

S 5689 State Rd 35 

Genoa, WI 54632 

(608) 689-2605 

tony_brady@fws.gov 

David Britton 

University of Texas at Arlington 

Box 19498, Biology Dept 

Arlington, TX 76019 

(817) 272-5577 

britton@uta.edu 

Mike Brittsan 

Columbus Zoo & Aquarium 

9990 Riverside Drive 

Powell, OH 43065 

(614) 72403580 

mike.brittsan@columbuszoo.org 

Joy Broach 

USACE 

P.O. Box 1070 (PM-P) 

Nashville, TN 37202-1070 

(615) 736-7956 

joy.i.broach@lrn02.usace.army.mil 

Andy Brookens 

Skelly and Loy, Inc. 

3642 Pampas Circle 

Chambersburg, PA 17201 

(717) 263-2244 

abrook@comcast.net 

Kenneth M. Brown 

Dept. of Biological Sciences, 508 LSB 

Louisiana State University 

Baton Rouge, LA 70803-1725 

(225) 578-4201 

kmbrown@lsu.edu 

Al Buchanan 

MO Dept. of Conservation 

1110 S. College Ave. 

Columbia, MO 65201 

(573) 882-9880 x 3257 

buchaa@mail.mdc.state.mo.us 

Jennifer Buhay 

Brigham Young University 

Dept. of Integrative Biology 

401 Widtsoe Building 

Provo, UT 84602 

(801) 422-9375 

crayfish@email.byu.edu 

John Burch 

Univ. of Michigan, Museum of Zoology 

1109 Geddes Ave. 

Ann Arbor, MI 48109-1079 

(734) 647-2189 

jbburch@umich.edu 

Ryan Burdick 

101 S. Shaffer St., 21D 

Athens, OH 45701 

(740) 407-2191 

rb330202@ohio.edu 

Robert Butler 

USFWS 

160 Zillicoa Street 

Asheville, NC 28801 

(828) 258-3939 x 235 

bob_butler@fws.gov 

48 

Scott Byrne 

73 Elm Street 

Dover, NJ 07801-2813 

(973) 989-5326 

scottbyrne@ems.att.com 

Holly Cabrera 

TNARI 

5385 Red Clay Road 

Cohutta, GA 30710 

Geoff Call 

US FWs 



(931) 528-6481 

geoff_call@fws.gov 

David Campbell 

University of Alabama 

425 Scientific Collections Bldg. 

Box 870345 

Tuscaloosa, AL 35487-0345 

(205) 348-5828 

amblema@bama.ua.edu 

Stephanie Carman 

Conservation Services Division 

New Mexico Dept. of Game and Fish 

P.O. Box 25112 

Santa Fe, NM 87504 

(505) 476-8092 

scarman@state.nm.us 

Joe Carney 

Dept. of Zoology 

Brandon University 

Brandon, Manitoba R7A 649 

Canada 

(204) 727-9607 

carneyj@brandonu.ca 

Stephanie Chance 

TVA 

2811 Lake City Hwy 

Lake City, TN 37769 

(865) 632-1605 

sachance@tva.gov 

Steve Chordas III 

Ohio State University 

1063 West 2nd Avenue 

Columbus, OH 43212-3612 

(614) 421-7159 

chordas.2@osu.edu

Alan Christian 

Dept. Biological Sciences 

Arkansas State University 

P.O. Box 599 

State University, AR 72467 

(870) 972-3082 

achristian@astate.edu 

Ronald Ciccerello 

KY Nature Preserves 

801 Schenkel Lane 

Frankfort, KY 40601 

(502) 573-2886 

ronald.cicerello@ky.gov 

Stephanie Clark 


Tuscaloosa, AL 35487 

(205) 348-5828 

meridolum@ozemail.com.au 

Janet Clayton 

WV Div. Natural Resources 

PO Box 67, Ward Road 

Elkins, WV 26241 

(304) 637-0245 

jclayton@dnr.state.wv.us 

Jeff Conway 

USFWS, Wolf Creek NFH 

50 Kendall Rd 

Jamestown, KY 42629 

(270) 343-3797 

jeff_conway@fws.gov 

Joyce Coombs 

University of TN - Knoxville 

8236 Wood Road 

Corryton, TN 37721-2114 

(865) 974-7229 

jcoombs@utk.edu 

W. Gregory Cope 

North Carolina State 

Dept. Environmental Toxicology 

Box 7633 


(919) 515-5296 

greg_cope@ncsu.edu 

James R. Cordeiro 

Nature Serve 

11 Avenue de Lafayette, 5th Floor 

Boston, MA 02111 

(617) 542-1908 

jay_cordeiro@natureserve.org 

Mark Cornish 

USACE Rock Island District 

P.O. Box 2004, Clock Tower Bldg. 

Rock Island, IL 61204-2004 

(309) 794-5385 

mark.a.cornish@usace.army.mil 

Peter Cosgrove 

Coilintra House, High Street 

Grantown on Spey 

Morayshire, Scotland, UK 

PH26 3EN 

01479 873151 

pearlmussel@coilintra.freeserve.co.uk 

Karen Couch 

49 Sierra Circle 

Olathe, KS 66061 

(913) 829-3981 

kcouch.mussels@juno.com 

John Crane 

P.O. Box 1633 

Plymouth, MA 02362 

(508) 747-5021 

Betty Crump 

USDA Forest Service 

PO Box 1270 

Hot Springs, AR 71902 

(501) 321-5236 

bcrump@fs.fed.us 

Kevin Cummings 

Illinois Natural History Survey 

607 E Peabody Drive 

Champaign, IL 61820 

(217) 333-1623 

ksc@inhs.uiuc.edu 

Jason Curole 

Dept. of Biology, Marine Environ. Bio. 

University of Southern California 

Los Angeles, CA 90089 

(818) 795-6486 

jcurole@usc.edu 

Andrew Currie 

USFWS, Dale Hollow NFH 

50 Fish Hatchery Rd 

Celina, TN 38551-6269 

(931) 243-2443 

andrew_currie@fws.gov 

Hua Dan 

113 College of Natural Resources 

Virginia Tech 

Blacksburg, VA 24061 

(540) 231-5927 

huad@vt.edu 

49 

Chris Davidson 

US Fish and Wildlife Service 

1500 Museum Road, Suite 105 

Conway, AR 72032 

(501) 513-4481 

chris_davidson@fws.gov 

Mike Davis 

MN DNR 

1801 S. Oak Street 

Lake City, MN 55041 

(651) 345-3331 

mike.davis@dnr.state.mn.us 

Sara Denham 

McHenry County Conservation District 

6419 Giant Oaks Road 

Wonder Lake, IL 60097 

(815) 728-8307 

sdenham@mccdistrict.org 

David Dettman 

Univ. of Arizona, Geosciences Dept. 

Gould-Simpson Bldg. 

Tucson, AZ 85721 

(520) 621-4618 

dettman@geo.arizona.edu 

Tom Dickinson 

The Catena Group, Inc. 

410 - B Millstone Drive 

Hillsborough, NC 27278 

(919) 732-1300 

tdickinson@thecatenagroup.com 

Carmen Dillon 

Marshall University 

2469 Third Avenue, Apt. 102 

Huntington, WV 25703 

(304) 617-9063 

dillon33@marshall.edu 

Rob Dillon, Jr. 

College of Charleston, Dept. of Biology 

66 George St. 

Charleston, SC 29424 

(843) 943-8087 

dillonr@cofc.edu 

Ron Dimock 

Wake Forest University 

Dept. of Biology, P.O. Box 7325 

Winston-Salem, NC 27109 

(336) 758-5567 

dimock@wfu.edu 

Barbara Dinkins 

Dinkins Biological Consulting 

3716 W. Beaver Creek Drive 

Powell, TN 37849 

(865) 938-7739 

biodink@covenantmail.net

Gerald Dinkins 

Dinkins Biological Consulting 

3716 W. Beaver Creek Drive 

Powell, TN 37849 

(865) 938-7739 

biodink@covenantmail.net 

Benjamin Dodd 

SMSU 

2524 S. Ingram Mill Road (213) 

Springfield, MO 65804 

(417) 889-3282 

bjd464s@smsu.edu 

Daniel Drennen 

US FWS 

6578 Dogwood View Parkway 

Ridgeland, MS 39213 

(601) 321-1127 

daniel_drennen@fws.gov 

Kari Duncan 

USFWS 

4401 N. Fairfax Drive 

Arlington, VA 22203 

(703) 358-2464 

kari_duncan@fws.gov 

Heidi Dunn 

Ecological Specialists, Inc. 

1417 Hoff Industrial Park 

O'Fallon, MO 63366 

(636) 281-1982 

hdunn@ecologicalspecialists.com 

Daniel Dutton 

105 Sherwood Court 


(540) 961-6510 

ddutton@vt.edu 

Stanley Dvorak 

Field Museum Natural History (volunteer) 

3512 Woodside 

Brookfield, IL 60513 

(708) 387-0687 

EBSCO Industries, Inc. 

Archives and Research Library 

New Brunswick Museum 

277 Douglas Ave 

St John, NB E2K 1E5 

Canada 

David Edds 

Emporia State University 

1200 Commercial Street 

Campus Box 4050 

Emporia, KS 66801-5087 

(316) 341-5622 

eddsdavi@emporia.edu 

Curt Elderkin 

Miami University 

212 Pearson Hall, Dept. of Zoology 


(513) 529-3180 

elderkcl@muohio.edu 

John Epifanio 


607 E. Peabody Drive 


William Ettinger 

Normandeau Associates 

23723 Woods Drive 

Lewes, DE 19958-3314 

(302) 945-3567 

wettinger@normandeau.com 

Brian Evans 

USFWS 

330 Cummings Street 

Abingdon, VA 24210 

(276) 623-1233 x 26 

brian_evans@fws.gov 

Ryan Evans 

Western PA Conservancy 

61 N. Linwood Avenue 

Pittsburgh, PA 15205 

(412) 478-0489 

revans@paconserve.org 

Mark Fagg 

TN Wildlife Resources Agency 

3030 Wildlife Way 

Morristown, TN 37814 

(423) 587-7037 x 112 

mark.fagg@state.tn.us 

Joe Ferraro 

VA Dept. Game & Inland Fisheries 

1724 Buller Hatchery Road 

Marion, VA 24354 

(276) 783-2138 

jferraro@dgif.state.va.us 

Brant Fisher 

IN Dept. Natural Resources 

Atterbury Fish & Wildlife Area 

7970 S. Rowe St., P.O. Box 3000 

Edinburgh, IN 46124 

(812) 526-5816 

bfisher@dnr.state.in.us 

James Flowers 

North Carolina State University 

P.O. Box 1459 

Pittsboro, NC 27312 

(919) 513-6404 

james_flowers@ncsu.edu 

50 

Brook Fluker 

101 Garrett Hall, Dept. of Biology 

Univ. of Louisiana at Monroe 

Monroe, LA 71209-0520 

(318) 342-3299 

fluker44@hotmail.com 

Michael Ford 

NOAA 

2725 Montlake Blvd E. 

Seattle, WA 98112 

(206) 860-5612 

mike_ford@moaa.gov 

Steve Fraley 

NC Wildlife Resources Commission 

10257 Rush Fork Road 

Clyde, NC 28721 

(828) 627-8414 

fraleysj@earthlink.net 

Paul Freeman 


2100 First Ave North, Suite 500 

Birmingham, AL 35203 

(205) 251-1155 x 110 

pfreeman@tnc.org 

Terrence Frest 

2517 NE 65th Street 

Seattle, WA 98115-7125 

(206) 527-6764 

tjfrest@earthlink.net 

John Fridell 

USFWS 



(828) 258-3939 x 225 

john_fridell@fws.gov 

Michael Gangloff 

Dept of Biological Sciences 

331 Funchess Hall 

Auburn University 

Auburn, AL 36849 

(334) 844-7345 

ganglmm@auburn.edu 

Jeff Garner 

AL Div Wildlife & Freshwater Fisheries 

350 County Road 275 

Florence, AL 35630 

(256) 767-7673 

bleufer@aol.com

Catherine Gatenby 

USFWS 

400 East Main Street 

White Sulphur Springs, WV 24986 

(304) 536-1361 

catherine_gatenby@fws.gov 

Jochen Gerber 

Division of Invertebrates 

Field Museum of Natural History 

1400 S. Lake Shore 

Chicago, IL 60605-2496 

(312) 665-7577 

jgerder@fieldmuseum.org 

Leighann Gipson 

US Army Corps of Engineers 

167 N. Main St., Room B-202 

Memphis, TN 38103 

(901) 544-4015 

leighann.c.gipson@mvm02.usace.army.mil 

Jim Godwin 

AL Natural Heritage Program 

Huntingdon College 

1500 E. Fairview Ave. 

Montgomery, AL 36106 

(334) 834-4519 

jgodwin@alnhp.org 

Larry Goldman 

USFWS 

P.O. Drawer 1190 

Daphne, AL 36526 

(251) 441-5181 

larry_goldman@fws.gov 

Cory Goldsworthy 

205 Pennebaker Hall 

North Dixie Avenue 


(981) 267-6553 

cgoldswor21@tntech.edu 

Roger Gorden 

USFWS 

Genoa NFH 

S 5689 State Highway 35 

Genoa, WI 54632 

(608) 689-2605 

roger_gorden@fws.gov 

Daniel Graf 

The Academy of Natural Sciences 

1900 Benjamin Franklin Parkway 

Philadelphia, PA 19103 

(215) 299-1132 

graf@acnatsci.org 

Lane C. Graham 

The University of Manitoba 

42 Crowson Bay 

Winnipeg Manitoba R3T 0J8 

Canada 

(204) 452-9591 

lcgrahm@cc.umanitoba.ca 

James Gray 

USFWS, Wolf Creek NFH 

50 Kendall Rd 

Jamestown, KY 42629 

(270) 343-3797 

james_gray@fws.gov 

Janet Gregory 


Univ. of Louisiana at Monroe 

Monroe, LA 71209-0520 

(318) 342-1799 

gregorj1@tribe.ulm.edu 

Jennifer Guyot 

149 Cheatham Hall 

Virginia Tech 


(540) 231-5320 

guyot@vt.edu 

Eric Hallerman 

VA Tech, Dept. Fisheries & Wildlife Sci. 


(540) 231-3257 

ehallerm@vt.edu 

Ed Hammer 

5711 Lenox Road 

Lisle, IL 60532 

(630) 353-1933 

ejhammer@sbcglobal.net 

Jeff Hard 

NOAA Fisheries, NW Fisheries Sci. Ctr 

2725 Montlake Blvd. E. 

Seattle, WA 98112 

(206) 860-3275 

jeff.hard@noaa.gov 

Willard Harman 

SUNY - Oneonta, Biol. Field Station 

5838 State Hwy 80 

Cooperstown, NY 13326 

(607) 547-8778 

harmanwn@oneonta.edu 

Fred Harris 

NC Wildlife Commission 

1721 Mail Service Center 


(919) 733-3633 

fred.harris@ncwildlife.org 

51 

John L. Harris 

12301 Pleasant Forest Drive 

Little Rock, AR 72212 

(501) 569-2281 

omibob@aol.com 

Julian R. Harrison, Ph. D. 

College of Charleston 

738 Swanson Avenue 

Charleston, SC 29412-9140 

(843) 795-1694 

harrisonj@cofc.edu 

Paul Hartfield 

USFWS 

6578 Dogwood View Parkway 

Jackson, MS 39213 

(601) 321-1125 

paul_hartfield@fws.gov 

Marian Havlik 

Malacological Consultants 

1603 Mississippi Street 

LaCrosse, WI 54601-4969 

(608) 782-7958 

havlikme@aol.com 

David Hayes 

1306 N. McGuire Avenue 

Monroe, LA 71203 

(318) 355-7306 

hayesdm9@juniata.edu 

Tom Hayes 

Pittsburgh Zoo and PPG Aquarium 

One Wild Place 


(412) 686-1803 

mqhayes@aol.com 

David Heath 

WI DNR 

3550 Mormon Coulee Road 

LaCrosse, WI 54601 

(608) 785-9993 

david.heath@dnr.state.wi.us 

Ryan Heise 

NCWRC 

1142 I-85 Service Road 

Creedmoor, NC 27522 

(919) 528-9886 

ryan.heise@ncwildlife.org 

Don Helms 

Helms & Associates 

814 North 7th Street 

Bellevue, IA 52031-9321 

(563) 872-4563 

helmsdon@cistelecom.net


National Mollusc Collection 

Dept. Evolution, Systematics & 

Ecology 

Hebrew University 

Jerusalem, Israel IL 91904 


Max Henschen 

3023 Winfield Ave. 

Indianapolis, IN 46222-1951 

(317) 926-6430 

Jeff Herod 

USFWS 

107 Highway 85N 

Niceville, FL 32578 

(850) 882-4164 x 372 

jeffrey_herod@fws.gov 

Nathaniel Hitt 


Virginia Tech 

Blacksburg, Va 24060 

(540) 231-5573 

than@vt.edu 

Michael Hoggarth 

Otterbein College 

Dept. Life & Earth Sciences 

Westerville, OH 43081 

(614) 823-1667 

mhoggarth@otterbein.edu 

Jessica Hoisington 


309 Stoddard Hall 


(513) 529-6691 

hoisinj1@muohio.edu 

Ellet Hoke 

1878 Ridgeview Circle Drive 

Manchester, MO 63021 

(636) 391-9459 

ellethoke@earthlink.net 

Daniel Hornbach 

Macalester College 

1600 Grand Ave., Provost Office 

St. Paul, MN 55105 

(651) 696-6160 

hornbach@macalester.edu 

Mark Hove 

Macalester College 

1600 Grand Ave. 


(651) 696-6827 

mark.hove@umn.edu 

Charles Howard 

Enviroscience, Inc. 

3781 Darrow Road 

Stow, OH 44224 

(330) 688-0111 

choward@enviroscienceinc.com 

Jeanette Howard 

73239 Confederated Way 

P.O. Box 638 

Pendleton, OR 97801 

(541) 966-2370 

jhowie@uclink.berkeley.edu 

Robert G. Howells 

Texas Parks & Wildlife Dept. 

Heart of the Hills Fisheries Science Ctr. 

5103 Junction Highway 

Ingram, TX 78025 

(830) 866-3356 

robert.howells@tpwd.state.tx.us 

Tantan Huang 

4700 Hillsborough Street 


(919) 513-6302 

yhuang6@unity.ncsu.edu 

Don Hubbs 


PO Box 70 

Camden, TN 38320 

(731) 584-9032 

tnmussels@aol.com 

Fred Huber 

US Forest Service 

5162 Valleypointe Parkway 

Roanoke, VA 24019 

(540) 265-5157 

fhuber@fs.fed.us 

Patrick Hubert 


13498 N. 200E Road 

Fairmount, IL 61841 

(217) 896-3025 

phubert@uiuc.edu 

Robert Hudson 

Presbyterian College, Biology Dept. 

503 S. Broad Street 

Clinton, SC 29325 

(864) 833-8448 

rhudson@mail.presby.edu 

R. Douglas Hunter 

Oakland University 

2200 N. Squirrel Rd. 

Rochester, MI 48309-4476 

(248) 370-3552 

hunter@oakland.edu 

52 

Carla Hurt 

Arizona State University 

1716 S. Sycamore 

Mesa, AZ 85202 

(480) 775-3726 

carla_asu@yahoo.com 

Will Jaeckle 

Illinois Wesleyan Univ, Dept. Biology 

P.O. Box 2900 

Bloomington, IL 61702-2900 

(309) 556-1063 

wjaeckle@iwu.edu 

Joan Jass 

Milwaukee Public Museum 

800 West Wells 

Milwaukee, WI 53233 

(414) 278-2761 

jass@mpm.edu 

John Jenkinson 

TVA 

305 Revere Avenue 

Clinton, TN 37716 

(865) 632-1513 

jjjenkinson@tva.gov 

Kurt Jirka 

Ichthyological Associates, Inc. 

50 Ludlowville Road 

Lansing, NY 14882 

(607) 533-8801 

kjjirka@yahoo.com 

Nathan Johnson 

604 McBryde Drive 


(540) 239-0876 

najohns4@vt.edu 

Paul D. Johnson 

Tennessee Aquarium Research Institute 



(706) 694-4419 

pdj@tnari.org 

Richard I. Johnson 

Museum of Comparative Zoology 

Harvard University 

Cambridge, MA 02138 

(617) 493-2468 

Jess Jones 


Virginia Tech 

Blacksburg, VA 24061-0321 

(540) 231-7241 

vtaquaculture@hotmail.com

Thomas Jones 


1 John Marshall Highway 


(304) 696-7243 

jonest@marshall.edu 

Gary Jordan 

USFWS 

P.O. Box 33726 

Raleigh, NC 27636-3726 

(919) 856-4520 x 32 

gary_jordan@fws.gov 

David Kamms 

7784 E. Linden Lane 

Parma, OH 44130 

(440) 845-7545 

dudmeister2000@yahoo.com 

Cindy Kane 

USFWS 

6669 Short Lane 

Gloucester, VA 23061 

(804) 693-6694 x 109 

cindy_kane@fws.gov 

Brian Keas 

Ohio Northern University 

Dept. of Biol. Sciences 

Ada, OH 45810 

(419) 772-2335 

b-keas@onu.edu 

Eugene Keferl 

Coastal GA Community College 

Dept. Natural Sciences & Mathematics 

3700 Altama Ave. 

Brunswick, GA 31520 

(912) 262-3089 

keferl@cgcc.edu 

Daniel Kelner 


190 5th Street East 


(651) 290-5277 

daniel.e.kelner@usace.army.mil 

John Kent 

394 Cub Creek Road 

Chapel Hill, NC 27517-6327 

(919) 933-5650 

jkent@tmug.org 

David Kesler 

Rhodes College, Biology Dept. 

2000 N. Parkway 


(901) 843-3557 

kesler@rhodes.edu 

John Michael Sawaske King 


409 Cabel Court 


(304) 419-0082 

mountaineerh2o@yahoo.com 

Tim King 

USGS 

Leetown Science Center 

11649 Leetown Road 

Kearneysville, WV 25430 

(304) 724-8340 

tim_king@usgs.gov 

Ray Kinney 

Siuslaw Soil & Water Conserv. District 

91636 West Fork Road 

Deadwood, OR 97430 

(541) 964-3981 

kennyr@casco.net 

Sheila Kirk 

USFWS 

50 Fish Hatchery Road 

Celina, TN 38551 

(931) 243-2443 

sheila_kirk@fws.gov 

Helen Elise Kitchel 

WI DNR/BER 

101 S. Webster St. 

Madison, WI 53707 

(608) 266-5248 

lisie.kitchel@dnr.state.wi.us 

Katherine Klyce 


5385 Red Clay Rd 


(706) 694-4666 

kklyce@tnari.org 

Stephen Kneeland 

5755 Nutting Hall 

Univ. of Maine 

Orono, ME 04469 

(207) 581-2921 

stephen.kneeland@umit.main 

Karen Kobey 

Three Rivers Park District 

10360 West River Rd 

Brooklyn Park, MN 55444 

(763) 424-8172 

kkobey@threeriversparkdistrict.org 

53 

Leroy Koch 

USFWS 

3761 Georgetown Road 

Frankfort, KY 40601 

(502) 695-0468 

leroy_koch@fws.gov 

Martin Kohl 

TN Dept. Environment & Conservation 

3003 Greenway Drive 


(865) 689-5732 

mkohl1@aol.com 

L. Russert Kraemer 

Dept. Biological Sciences, SCEN - 632 

Univ. of Arkansas 

Fayetteville, AR 72701 

(479) 575-3251 

rkraemer@uark.edu 

Leos Kral 

State Univ. of West Georgia, Dept. of Bio. 

Carrollton, GA 30118 

(770) 836-4546 

lkral@westga.edu 

George Krause 

900 West Spring Valley Road 

Centerville, OH 45458 

(937) 433-8619 

mmgkrause@aol.com 

Danielle Kreeger 

Academy of Natural Sciences 

1900 Ben Franklin Pkwy 

Philadelphia, PA 19103 

(215) 299-1184 

kreeger@acnatsci.org 

Eric Krueger 


1049 Morrison Drive 

PO Box 20246 


(843) 937-8807 x 18 

ekrueger@tnc.org 

Kody Kuehnl 


6685 Concourse Drive 

Columbus, OH 43229 

(614) 901-0496 

kuehnl.2@osu.edu 

Gerald Lang 

Carnegie Museum of Natural History 

22317 Highway 285 

Cochranton, PA 16314 

(814) 332-0390 

jerrylang1@hotmail.com

Susan Lanier 


PO Box 40747 

Nashville, TN 37204 

(615) 781-6543 

susan.lanier@state.tn.us 

James Layzer 

TN Coop. Fish. Res. Unit 

Tenn. Tech Univ., Box 5114 


(931) 372-3032 

jim_layzer@tntech.edu 

Jacquie Lee 

6549 Georgia Street 

Burnaby, BC V5B 1V9 Canada 

(604) 294-6199 

jacquie_lee@telus.net 

Bill Lees 

University of Richmond 

2115 Jefferson Avenue 

Richmond, VA 23223 

(804) 648-3111 

william.lees@richmond.edu 

William Lellis 

National Park Service 

UMCES Appalachian Laboratory 

301 Braddock Road 

Frostburg, MD 21532 

(301) 689-7108 

wlellis@al.umces.edu 

Jay Levine 

North Carolina State University 



(919) 513-6497 

jay_levine@ncsu.edu 

Todd Levine 


Zoology Dept., Pearson Hall 


(513) 529-3387 

levinetd@muohio.edu 

Aaron Liberty 

PO Box 10284 


(540) 998-0885 

aliberty@vt.edu 

Douglas Locy 

Aquatic Systems, Inc. 

4621 Baptist Road 


(412) 427-2758 

aquatic.systems@comcast.net 

Cynthia Loftin 

Maine Coop. Fish & Wildlife Unit 

5755 Nutting Hall 

University of Maine 

Orono, ME 04469-5755 

(207) 581-2843 

cyndy_loftin@umenfa.maine.edu 

Charles Lydeard 


Biodiversity and Systematics 

Box 870345 


(205) 348-1792 

clydeard@bama.ua.edu 

Gerry Mackie 

University of Guelph 

Dept. of Zoology 

Guelph, Ontario N1G 2W1 

Canada 

(519) 824-4120 x 5305 

gmackie@uoguelph.ca 

Paul Marangelo 


101 East Grand River Ave. 

Lansing, MI 48906 

(517) 316-2270 

pmarangelo@tnc.org 

Kevin Markham 

Environmental Services, Inc. 

524 South New Hope Road 


(919) 212-1760 x 123 

kmarkham@esinc.cc 

Scott Martin 

712 Harley Drive 


(614) 447-3600 x 2483 

smartin@cas.org 

David Martinez 

USFWS 

6315 E., 57th Place 

Tulsa, OK 74135-8122 

(918) 581-7458 x 228 

david_martinez@fws.gov 

Larry Master 

NatureServe 

11 Avenue de Lafayette, 5th Floor 

Boston, MA 02111 

(617) 542-1908 x 230 

larry_master@natureserve.org 

54 

Charles Mather 

Univ. of Science & Arts of Oklahoma 

1727 W. Alabama 

Chickasha, OK 73018 

(405) 574-1282 

facmathercm@usao.edu 

Richard Mayden 

Saint Louis University 


3507 Laclede Avenue 

St. Louis, MO 63103-2010 

(314) 977-3494 

maydenrl@slu.edu 

Christine Mayer 

Ilinois Natural History Survey 

607 E. Peabody Dr. 


(217) 244-2354 

camayer@uiuc.edu 

Katherine McCafferty 

Georgia DOT 

1000 St. Charles Ave., Apt 2 

Atlanta, GA 30306 

(770) 712-2126 

kamccafferty@yahoo.com 

James McCann 

Maryland DNR 

Natural Heritage Program 

PO Box 68 

Wye Mills, MD 21679 

(410) 827-8612 

jmccann@dnr.state.md.us 

Mary McCann 

Devine Tarbell & Associates 

970 Baxter Blvd. 

Portland, ME 04103 

(207) 775-4495 

mary.mccann@devinetarbell.com 

Stephanie McCormick 


University of Louisiana at Monroe 

Monroe, LA 71209-5020 

(318) 342-3299 

stephjmcc@yahoo.com 

Henry McCullagh 

2735 Holly Point Road East 

Orange Park, FL 32073 

(904) 264-8384 

lmcstjohns@aol.com

Daryl McGoldrick 

Environment Canada 

National Water Research Inst. 

PO Box 5050, 867 Lakeshore Road 

Burlington, Ontartio L7R 4A6 

Canada 

(905) 336-4790 

daryl.mcgoldrick@ec.gc.ca 

Stuart McGregor 

Geological Survey of Alabama 

P.O. Box 869999 


(205) 247-3629 

smcgregor@gsa.state.al.us 

Dan McGuire 

McGuire Consulting 

95607 Reata Road 

Kennewick, WA 99338 

(509) 628-8772 

Anna McIvor 

Dept. Zoology, Downing Street 

Cambridge University 

Cambridge, Cams CB2 3EJ 

UK 

alm1000@cam.ae.uk 

Robert McMahon 

Univ. of Texas at Arlington 

Honors College 

Box 19222 

Arlington, TX 76019 

(817) 272-7215 

r.mcmahon@uta.edu 

Stephen McMurray 

Missouri Dept. of Conservation 

1110 S. College Ave. 


(573) 882-9909 

stephen.mcmurray@mdc.mo.gov 

Sarah McRae 

NC Natural Heritage Program 

1601 MSC 


(919) 715-1751 

sarah.mcrae@ncmail.net 

Neil Medlin 

NC DOT 

2728 Capital Blvd., Suite 168 


(919) 715-1447 

knmedlin@dot.state.nc.us 

Timothy Merritt 

US FWS 



(931) 528-6481 

timothy_merritt@fws.gov 

Andrew Miller 

CEWES ER-EE-A 

3909 Halls Fery Road 

Vicksburg, MS 39180-6194 

(601) 634-2141 

millerA3@wes.army.mil 

Edwin Miller 

KS Dept Wildlife & Parks 

5089 Co. Road 2925 

Independence, KS 67301 

(620) 331-6820 

edm@wp.state.ks.us 

Glenn Miller 

USFWS 

2800 E. Lake Shore Dr. 

Ashland, WI 54806 

(715) 682-6186 x 210 

glenn_miller@fws.gov 

Denise Moldenhauer 

USFWS 



(828) 258-3939 x 226 

denise_moldenhauer@fws.gov 

Reverie Molina 

3939-C Marcom Street 


(919) 834-5115 

revmolina@yahoo.com 

Emy Monroe 


Zoology Dept. 


(513) 529-8183 

monroeem@muohio.edu 

William Montgomery 

66 Gorman Lane, Apt. B 

Reading, OH 45215 

(513) 769-6724 

Todd Morris 

Fisheries & Oceans Canada 

867 Lakeshore Rd. 

Burlington, Ontario L7R 4A6 

Canada 

(905) 336-4734 

morrist@dfo-mpo.gc.ca 

55 

Cheryl Morrison 

USGS 

Leetown Science Center 



(304) 724-4464 

cheryl_morrison@usgs.gov 

Patricia Morrison 

USFWS, Ohio River Islands NWR 

P.O. Box 1811 

Parkersburg, WV 26102 

(304) 422-0752 

patricia_morrison@fws.gov 

Rachel Muir 

USGS 

1967 B Villa Ridge Drive 

Reston, VA 20191 

(703) 262-0170 

damselfly@earthlink.net 

Melody Myers-Kinzie 

261 Pennwood Lane 

Brownsburg, IN 46112 

(317) 858-0677 

mmk7119@cs.com 

Ethan Nedeau 

Biodrawversity 

15 High Street 

Amherst, MA 01002 

(413) 253-6561 

ejnedeau@comcast.net 

Richard Neves 

Dept. of Fish & Wildlife 

Virginia Tech 


(540)231-5927 

mussel@vt.edu 

Teresa Newton 

USGS 

Upper Midwest Environ. Sci. Center 

2630 Fanta Reed Rd 

LaCrosse, WI 54603 

(608) 781-6217 

tnewton@usgs.gov 

Jerrie Nichols 

USGS 

1451 Green Road 

Ann Arbor, MI 48105 

(734) 214-7218 

s_jerrine_nichols@usgs.gov

Robert Nichols 

NC Wildlife Resource Commission 

1142 I-85 Service Road 

Creedmoor, NC 27522 

(919) 528-9886 

rob.nichols@ncwildlife.org 

Betsy Nightingale 


319 S. River Street 

Harrisburg, PA 17104 

(717) 234-3125 

c_bnightin@state.pa.us 

Sabrina Novak 




(706) 694-4666 

snovak@tnari.org 

Christine O'Brien 

Browns River Environmental Consultants 

279 River Road 

Underhill, VT 05489 

(802) 899-1768 

amblema@adelphia.net 

Scott O'Dee 

1687 Gypsy Lane 


(614) 891-5609 

sodee@insight.rr.com 

Brett Ostby 

Virginia Tech 



(540) 230-1042 

bostby@vt.edu 

Nick Owens 

Eastern Illinois University 

14783 E. CR. 600 N 

Charleston, IL 61920 

(217) 276-8047 

musselguy@hotmail.com 

Paul Parmalee 

Frank H. McClung Museum 

University of Tennessee 

Knoxville, TN 37996-3200 

(865) 974-2144 

pparmale@utk.edu 

Tim Pearce 

Carnegie Museum of Natural History 

4400 Forbes Avenue 


(412) 622-1916 

PearceT@Carnegiemnh.org 

Wendell Pennington 

250 McGee Lane 


(931) 526-6038 

Kathryn Perez 

Univ. of Alabama 

Box 870345 


(205) 348-5828 

perez005@bama.ua.edu 

Melissa Petty 

610 Montgomery Street 


(540) 552-3159 

mepetty@vt.edu 

John Petzing 


607 East Peabody Drive 


(217) 244-9864 

jpetzing@inhs.uiuc.edu 

Malcolm Pierson 

Alabama Power Company 

GSC #8, P.O. Box 2641 

Birmingham, AL 35291 

(205) 664-6177 

jmpierso@southernco.com 

Kyle Piller 

Dept. of Biological Sciences, SLU 

10736 

Southeastern Louisiana University 

Hammond, LA 70402 

(985) 549-2191 

kyle.piller@selu.edu 

Michael Pinder 

VA DGIF 

2206 S. Main St., Suite C 


(540) 961-8387 

mpinder@dgif.state.va.us 

Bill Posey 

AR Game & Fish Commission 

P.O. Box 6740 

Perrytown, AR 71801 

(877) 777-5580 

brposey@agfc.state.ar.us 

Gerald Pottern 

Robert J. Goldstein & Associates 

8380 Garvey Drive 


(919) 872-1174 

gpottern@RJGAcarolina.com 

56 

Jack K. Pounders 

Southeastern Divers, Inc. 

325 S. Court Street 


(256) 766-3483 

jack@sdiscuba.com 

Jeffrey Powell 

USFWS 

P.O. Drawer 1190, 1208-B Main St. 


(251) 441-5858 

jeffrey_powell@fws.gov 

Tom Proch 

PA DEP 

2721 Cedric Avenue 


(412) 343-6821 

tproch@stargate.net 

Warren Pryor 

University of St. Francis 

Biology Department 

2701 Spring Street 

Fort Wayne, IN 46808 

(260) 385-5659 

wpryor@sf.edu 

Morgan Raley 

NC State University 



(919( 618-3212 

morgan_raley@ncsu.edu 

Brenda Rashleigh 

US EPA 

960 College Station Road 

Athens, GA 30605 

(706) 355-8148 

rashleigh.brenda@epa.gov 

Joe Rathbun 

MDEQ 

3313 W. Mt. Hope Avenue, #113 

Lansing, MI 48911 

(517) 373-8868 

rathbunj@michigan.gov 

Marsha May Reimer 

Texas Parks & Wildlife Dept 

9602 Cheryl Lynn Drive 

Austin, TX 78747 

(512) 912-7062 

marsha.reimer@tpwd.state.tx.us

Bert Remley 

Third Rock Consultants, LLC. 

2514 Regency Road, Suite 104 

Lexington, KY 40503 

(859) 977-2000 

bremley@thirdrockconsultants.com 

Andy Roberts 

USFWS 

101 Park DeVille Drive, Suite A 


(573) 234-2132 

andy_roberts@fws.gov 

Jamie Roberts 

Fisheries and Wildlife Sciences 

Virginia Tech 


(540) 231-1707 

darter@vt.edu 

Angie Rodgers 


1424 Park Summit Blvd 

Apex, NC 27523 

(919) 460-7350 

angie.rodgers@earthlink.net 

Kevin Roe 

Delaware Museum of Natural History 

4840 Kennett Pike 

Wilmington, DE 19807-0937 

(302) 658-9111 

kroe@delmnh.org 

Susan Rogers 

USFWS 

687 Crimson Leaf Trail 

Eagan, MN 55123 

(612) 725-3548 x 219 

susan_rogers@fws.gov 

Bill Roston 

Box 623 

Forsuth, MO 65653 

(417) 546-2401 

roston@tri-lakes.net 

Nick Rowse 

USFWS 

4101 E. 80th Street 

Bloomington, MN 55425 

(612) 725-3548 x 210 

nick_rowse@fws.gov 

Louie Rundo 

647 Rehwinkle Road 

Sagamore Hills, OH 44067 

(216) 534-4788 

louie.rundo@lnoca.org 

Diane Rusanowsky 

Antioch New England Graduate School 

74 Miller Road 

Bethany, CT 06524-3224 

diane_rusanowsky@antiochne.edu 

Todd Sampsell 

Central Pennsylvania Conservancy 

P.O. Box 587 

Harrisburg, PA 17108 

(717) 223-0221 

sampsell_cpc@paonline.com 

Tiffany Sanders 

Arkansas State University 

P. O. 599 State University 

Jonesboro, AR 72467 

(870) 972-3082 

tiffany.sanders@astate.edu 

Tim Savidge 

The Catena Group, Inc. 

410-B Millstone Drive 


(919) 732-1300 

tsavidge@thecatenagroup.com 

Charles Saylor 

TVA 

516 Vista Drive 

Clinton, TN 37716 

(865) 403-2316 

cfsaylor@tva.gov 

Robert Schanzle 

Illinois DNR 

One Natural Resources Way 

Springfield, IL 62702-1271 

(217) 785-4863 

bschanzle@dnrmail.state.il.us 

Beth Schilling 

11931 Couch Mill Road 


(865) 691-8267 

schillingem@ornl.gov 

Frieda Schilling 

Greater St. Louis Shell Club 

3707 Lan Drive 

St. Louis, MO 63125-4415 

(314) 892-3454 

John Schmerfeld 

USFWS 

6669 Short Lane 

Glouchester, VA 23061 

(804) 693-6694 x 107 

john_schmerfeld@fws.gov 

57 

Jeffrey Schmid 

2404 Buchenhorst Road 

State College, PA 16801 

(814) 234-2102 

Kathryn Schneider 

Stuyvesant Environmental Consulting 

16 Frisbee Lane 

Stuyvesant, NY 12173 

(518) 799-3457 

fallline@berk.com 

Bob Schnelle 

Chair, Mussel Mitigation Trust 

139 East 4th Street, Room 552-A 

Cincinnati, OH 45202 

(513) 287-2239 

bschnelle@cinergy.com 

ul Schulze 

16113 ASA Drive 

Spencerville, MD 20868 

(301) 435-3636 

ps6@nih.gov 

Guenter Schuster 

Eastern Kentucky University 

Dept. Biological Sciences 

Richmond, KY 40475 

(859) 622-1016 

guenter.schuster@eku.edu 

John Schwegman 

3626 RiverPoint Lane 

Metropolis, IL 62960 

(618) 543-9429 

botany@midwest.net 

Mary Scott 

703 Gladiolus Drive #E8 

Jonesboro, AR 72404 

(870) 972-2017 

mc_scott27@yahoo.com 

Josh Seagraves 

2207 Highland 

Pocahontas, AR 72455 

(636) 281-1982 

jseagraves@ecologicalspecialists.com 

Jeremy Shiflet 

USFS/ Virginia Tech Dept. Fisheries & 

Wildlife 

13763 Turkeytown Road 

Broadway, VA 22815 

(540) 421-0432 

jeshifle@vt.edu

James Sickel 

Murray State University 

Biology Dept., BL 334 

Murray, KY 42071 

(270) 762-6326 

jim.sickel@murraystate.edu 

Jeffrey Sides 


609 King Valley Circle 

Pelham, AL 35124 

(205) 663-3312 

sides003@bama.ua.edu 

Bernard Sietman 

P.O. Box 782 

Chanhassen, MN 55317 

(651) 282-2509 

bernard.sietman@dnr.state.mn.us 

Jeff Simmons 


18 Sunrise Lane 

Candler, NC 28715 

(828) 665-4717 

jeff.simmons@earthlink.net 

Thomas Simpson 

McHenry County Conservation District 

6512 Harts Road 

Ringwood, IL 60072 

(815) 678-7644 

tsimpson@mccdistrict.org 

Chris Skelton 

Georgia College & State University 

Dept. of Biological & Environmental Sci. 

Milledgeville, GA 31061 

(478) 445-2440 

chris.skelton@gcsu.edu 

Allan K. Smith 

16661 SW Chehalem Way 

Hillsboro, OR 97123 

(503) 628-7825 

mxasmith@pacifier.com 

David R. Smith 

USGS, Leetown Science Center 



(304) 724-4467 

david_r_smith@usgs.gov 

Jamie Smith 

NC Museum of Natural Sciences 

5924 Forestville Road 


(919) 733-7450 x 762 

jamie.smith@ncmail.net 

Janice Smith 

Environment Canada 

National Water Research Institute 

PO Box 5050, 867 Lakeshore Road 

Burlington, Ontario L7R 4A6 

Canada 

(905) 336-4685 

janice.smith@ec.gc.ca 

Mark Smith 


1651 Dorset Drive 


(901) 683-7683 

mark.r.smith@mvm02.usace.army.mil 

Matthew Smith 

Environmental Services, Inc. 

524 South New Hope Road 


(919) 212-1760 

msmith@esiral.com 

Rob Southwick 

Southwick Associates, Inc. 

P.O. Box 6435 

Fernandina Beach, FL 32035 

(904) 272-9765 

rob@southwickassociates.com 

Michael Sovic 

The Ohio State University 

318 W. 12th Avenue 


(614) 292-4570 

sovic.1@osu.edu 

James Spence 


135 Oney Ave 


(304) 523-2060 

james.b.spence@lrh01.usace.army.mil 

Beverly Spurlock 

Cabell County WV Health Dept. 

6676 Merritts Creek Rd. 


(304) 736-2238 

David Stansbery 

Museum of Biological Diversity 


1315 Kinnear Rd 

Columbus, OH 43212-1192 

(614) 292-8560 

stansbery.1@osu.edu 

58 

George Stanton 

Dept of Biology 

Columbus State University 

Columbus, GA 31907 

(706) 568-2056 

stanton_george@colstate.edu 

James Steffen 

Chicago Botanic Garden 

1000 Lake Cook Road 

Glencoe, IL 60022 

(847) 835-8266 

jsteffen@chicagobotanic.org 

Bruce Stephen 

Southeast Community College 

8800 O Street 

Lincoln, NE 68510 

(402) 437-2544 

bstephen@southeast.edu 

Janet Sternburg 

MO Dept. of Conservation 

PO Box 180 

Jefferson City, MO 65101 

(573) 522-4115 

janet.sternburg@mdc.mo.us 

Philip Stevenson 

Creek Laboratory LLC 

P. O. Box 953 

Fredericksburg, VA 22404 

(540) 368-9227 

phil@creeklab.com 

Craig Stihler 

WV DNR 

P.O. Box 67, Ward Road 

Elkins, WV 26241 

(304) 637-0245 

cstihler@dnr.state.wv.us 

David Strayer 

Institute of Ecosystem Studies 

P.O. Box AB 

Millbrook, NY 12545 

(845) 677-5343 

strayerd@ecostudies.org 

Carson Stringfellow 

Columbus State University 

P.O. Box 186 

Waverly Hall, GA 31831 

(706) 571-8457 

stringfellow_carson@colstate.edu 

Gregory Styborski 

Civil & Environmental Consultants, Inc. 

333 Baldwin Road 


(412) 429-2324 

gstyborski@cecinc.com

Dale Suiter 

US FWS 

PO Box 33726 

Raleigh, NC 27636-3726 

(919) 856-4520 x 18 

dale-suiter@msn.com 

Kathryn Sukkestad 

Fort Stewart Fish & Wildlife 

1177 Frank Cochran Drive 

Ft. Stewart, GA 31314-4940 

(912) 767-1715 

sukkestadke@stewart.army.mil 

Gerald Summers 

University of Missouri 

114 Lefevre Hall 


summersg@missouri.edu 

Stacy Surgenor 

Mammoth Cave National Park 

P.O. Box 7 

Mammoth Cave, KY 42259 

(270) 758-2162 

stacy_surgener@nps.gov 

Beth Swartz 

ME Dept. Inland Fish & Wildlife 

650 State Street 

Bangor, ME 04401 

(207) 941-4476 

beth.swartz@maine.gov 

Doug Sweet 

Detroit Zoological Institute 

8450 West Ten Mile Road 

Royal Oak, MI 48068 

(313) 852-4141 

dsweet@detroitzoo.org 

Bob Szafoni 

Illinois DNR 

1660 W. Polk Street 

Charleston, IL 61920 

(217) 345-2420 

rszafoni@dnrmail.state.il.us 

Christopher Tabit 

University of West Georgia 

Biology Department 

Carrollton, GA 30118 

(770) 836-4543 

ctabit@westga.edu 

Rob Tawes 

USFWS 

P.O. Drawer 1190 


(251) 441-5830 

robert_tawes@fws.gov 

Ralph Taylor 


2 Francis Circle 

Barboursville, WV 25504 

(304) 736-1342 

snail@ezwv.com 

John Tetzloff 

Darby Creek Association 

606 Woodbury Avenue 


(614) 288-0313 

jftetzloff@aol.com 

Leslie Tewinkel 

US FWS 

1 Federal Drive 

Fort Snelling, MN 55111 

(612) 713-5164 

leslie_tewinkel@fws.gov 

Pam Thiel 

USFWS 

555 Lester Ave 

Onalaska, WI 54650 

(608) 783-8431 

pam_thiel@fws.gov 

Fred Thompson 

Florida Museum of Natural History 

P.O. Box 117800 

Gainesville, FL 32611-7800 

(352) 392-1721 x 458 

fgt@flmnh.ufl.edu 

Jeremy Tiemann 




(217) 244-4594 

jtiemann@inhs.uiuc.edu 

Richard Tippit 


212 Glenwood Drive 

Goodlettsville, TN 37202 

(615) 736-7958 

richard.n.tippit@usace.army.mil 

Sara Turner 

Purdue University 

175 Marsteller St. 

Dept. Forestry & Natural Resources 

West Lafayette, IN 47907 

(765) 494-3628 

smturner@purdue.edu 

59 

John VanHassel 

American Electric Power 

1 Riverside Plaza 


(614) 223-1249 

jhvanhassel@aep.com 

Caryn Vaughn 

Oklahoma Biological Survey 

University of Oklahoma 

111 E. Chesapeake St. 

Norman, OK 73019 

(405) 325-4034 

cvaughn@ou.edu 

Rita Villella 

USGS/ Leetown Science Center 



(304) 724-4472 

rita_villella@usgs.gov 

Susi von Oettingen 

USFWS 

70 Commercial Street, Suite 300 

Concord, NH 03301 

(603) 223-2541 

susi_vonoettingen@fws.gov 

Andrea Walther 

University of Michigan 

1109 Geddes Avenue 

Ann Arbor, MI 48109-1079 

(734) 647-2189 

awalther@umich.edu 

Doug Warmolts 

Columbus Zoo & Aquarium 

P.O. Box 400 

Powell, OH 43065-0400 

(614) 645-3524 

doug.warmolts@columbuszoo.org 

Mel Warren, Jr. 

US Forest Service, Southern Res. Station 

1000 Front Street 

Oxford, MS 38655 

(662) 234-2744 x 246 

fswarren@olemiss.edu 

Brian Watson 

VA Dept. Game & Inland Fisheries 

1143 Bell Lane 

Forest, VA 24551 

(434) 525-7522 

bwatson@dgif.state.va.us

Charles Watson, Jr. 

SoBran, Inc. c/o USEPA 

26 West Martin Luther King Drive 

Cincinnati, OH 45268 

(513) 569-7082 

procladius@aol.com 

G. Thomas Watters 


Museum of Biol. Diversity 

1315 Kinnear Rd. 


(614) 292-6170 

watters.1@osu.edu 

Gary Wege 

USFWS 

8811 Hillside Trail South 

Cottage Grove, MN 55016 

(612) 725-3548 x 207 

gary_wege@fws.gov 

Kurt Welke 

Wisconsin DNR, SCR HQ 


Fitchburg, WI 53711 

(608) 273-5946 

kurt.welke@dnr.state.wi.us 

Nevin Welte 

915 Laurel Avenue, Apt. U-3-E 


(931) 372-4028 

ntwelte21@tntech.edu 

Kara Wendeln 

Wright State University 

2562 Conwood Drive 

Beavercreek, OH 45434 

(937) 689-3681 

wendeln.4@wright.edu 

Amy R. Wethington 

Post Doctoral Research Associate 

Department of Biological Sciences 

Purdue University 

West Lafayette, IN 47907 

(765) 494-9103 

awething@purdue.edu 

Matthew White 

Ohio University 

Biological Sciences 

Athens, OH 45701 

(740) 593-2413 

whitem@ohio.edu 

Jim Williams 

USGS 

7920 NW 71st Street 

Gainesville, FL 32653 

(352) 264-3475 

jim_williams@usgs.gov 

Deborah Wills 

304 Sobotka Street 

Hartselle, AL 35640 

(256) 773-7641 

dwills@hiwaay.net 

Helpful Hints from Hoppy: 

Submitted by Steve Ahlstedt 

Doug Winford 

US FWS 



(931) 528-6481 

douglas_winford@fws.gov 

Jason Wisniewski 

1957 West Broad Street 

Apt 6E 


(931) 528-6978 

jmwisniews21@tntech.edu 

Michael Wood 

The Catena Group 

410-B Millstone Drive 


(919) 732-1300 

mwood@thecatenagroup.com 

60 

Robert Wood 

Saint Louis University 


3507 Laclede Ave. 

St. Louis, MO 63103-2010 

(314) 977-3904 

wood2@slu.edu 

Daelyn Woolnough 

Iowa State University 

1400 Gateway Hills Park Drive 

Apt. 902 

Ames, IA 50014 

(515) 292-1770 

daelyn@iastate.edu 

Paul Yokley, Jr. 

3698 Chisholm Road 


(256) 764-3780 

pyj@hiwaay.net 

Angela Zimmerman 

USFWS 

6950 Americana Parkway, Suite H 

Reynoldsburg, OH 43068 

(614) 469-6923 x 22 

angela_zimmerman@fws.gov 

Greg Zimmerman 

EnviroScience, Inc. 

3781 Darrow Road 

Stow, OH 44224 

(330) 688-0111 

gzimmerman@enviroscienceinc.com 

Lora Zimmerman 

USFWS 

176 Croghan Spur Road, Suite 200 


(843) 727-4707 

lora_zimmerman@fws.gov 

Needles, glass, nails, and spam cans will ruin your 

day…wear Kevlar gloves, kneepads, and boots. 

Don’t come crying to me!

If you are interested in joining a committee, please contact the appropriate chair. 

 

W. Gregory Cope 

North Carolina State 

Dept. Environ. & Molecular Toxicology 

Box 7633 

Raleigh, NC 27695-7633 

919-515-5296 

greg_cope@ncsu.edu 

 

Patricia Morrison 

USFWS, Ohio River Islands NWR 

P.O. Box 1811 

Parkersburg, WV 26102 

304-422-0752 

patricia_morrison@fws.gov 

 

Paul Johnson 

Tennessee Aquatic Research Institute 



706- 694-4419 Fax: 3957 

pdj@sari.org 

Guidelines and 

John Van Hassel 

American Electric Power 

1 Riverside Plaza 


614-223-1249 Fax: 1252 

jhvanhassel@aep.com 

Steve A. Ahlstedt 

USGS 

1820 Midpark Drive 


865-545-4140 x 17 Fax: 4496 


 

 

Kevin Cummings 


172 Natural Resources Building 



217-333-1623 

ksc@inhs.uiuc.edu 

 

Kevin J. Roe 

Delaware Museum of Natural History 

4840 Kennett Pike 

P.O. Box 3937 

Wilmington, DE 19807-0937 

302-658-9111 Fax: 2610 

kroe@delmnh.org 

 

Kurt Welke 

Wisconsin - DNR 



608-275-3266 

welkek@dnr.state.wi.us 

Join the UNIO Listserver 

http://my.fit.edu/~rtankers/unio.htm 

 

Jess Jones 

Virginia Tech 

606 Broce Drive 


540-231-5927 

vtaquaculture@hotmail.com 

 

 

Kurt Welke 

Wisconsin - DNR 



608-275-3266 

welkek@dnr.state.wi.us

… dedicated to the advocacy and conservation science of freshwater molluscan resources

August 2004 - FMCS-Freshwater Mollusk Conservation

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