Checklist of - Entomological Society of Canada

l*l 331:!y" 

Checklist of 

of Canada and Alaska

Checklist of 

BEETLES 

of Canada and Alaska 

Edited by 

Yves Bousquet 

Biosystematics Research Centre 

Ottawa, Ontario 

Research Branch 

Agriculture Canada 

Publication 186UE 

t99l

oMinister of Supply anrl Services Canada 1991 

Available in Canada through 

Assoeiated Bookstores 

and other borksellers 

or by mail fr,rm 

Canada Com-nunication Group 

- Publishing 

Ottawa, Canada KlA0S9 

Cat. no. A43-1861/1991E 

ISBN 0-660-13?67-4 

Canadian Cataloguing in Publication Data 

Checklist c f beetles of Canada and Alaska 

(Publication ; 18'61/E) 

Includes bibliographical references. 

Cat. no. A43. 1861i1991E 

ISBN 0-660- r3767-4 

1. Beetles-(lanada-Nomenclature. 2. Beetles- 

Alaska-Nomr:nclature. I. Bousquet, Yves. II. 

Canada. Agriculture Canada. Research Branch. 

III. Series: .?ublication (Canada. Agriculture 

Canada). English ; 1861/E. 

QI581.C4 1991 595.76 C91-099101-4 

Stafr Editor 

Frances Smith

CONTENTS 

List ofcontributors v 

Acknowledgments v 

Summary vi 

R6sum6 vi 

Synoptic table 1 

Introduction 4 

Format 4 

Subsequent revisions ofthe checklist 5 

Abbreviations and symbols 5 

ORDERCOLEOPTERA 6 

SuborderArchostemata 6 

Family Cupedidae (J.M. Campbell) 6 

Family Micromalthidae (J.M. Campbell) 6 

SuborderAdephaga 7 

Family Rhysodidae (Y. Bousquet) 7 

Family Trachypachidae (Y. Bousquet) 7 

Family Carabidae (Y. Bousquet) 8 

Family Haliplidae (R.E. Roughley) 60 

Family Amphizoidae (Y. Bousquet) 61 

Family Noteridae (R.E. Roughley) 61 

Family Dy'tiscidae (D.J. Larson and R.E. 

Roughley) 62 

Family Gyrinidae (R.E. Roughley) 72 

Suborder Polyphaga 74 

SuperfamilyStaphylinoidea 74 

Family Hydraenidae (R.E. Roughley) 74 

Family Ptiliidae (J.M. Campbell) 75 

Family Limulodidae (J.M. Campbell) 76 

Family Agyrtidae (J.M. Campbell) 77 

Family Leiodidae (S.B. Peck) 77 

Family Scydmaenidae (A. Davies) 82 

Family Micropeplidae (J.M. Campbell) 84 

Family Silphidae (J.M. Campbell) 84 

Family Staphylinidae (J.M. Campbell and A. 

Davies) 86 

Family Scaphidiidae (J.M. Campbell) 124 

Family Pselaphidae (A. Davies) 125 

SuperfamilyHydrophiloidea 130 

Family Hydrophilidae (R.E. Roughley) 130 

Family Georyssidae (R.E. Roughley) 135 

Family Sphaeritidae (R.E. Roughley) 135 

Family Histeridae (A. Davies) 135 

SuperfamilyEucinetoidea I4I 

Family Eucinetidae (J.M. Campbell) I4I 

Family Clambidae (J.M. Campbell) I4l 

Family Scirtidae (J.M. Campbell) 142 

SuperfamilyDascilloidea 143 

Family Dascillidae (J.M. Campbell) 143 

Family Rhipiceridae (J.M. Campbell) I43 

SuperfamilyScarabaeoidea 143 

Family Lucanidae (J. McNamara) 143 

Family Passalidae (J. McNamara) 145 

Family Scarabaeidae (J. McNamara) 145 

SuperfamilyByrrhoidea 158 

Family Byrrhidae (P.J. Johnson) 158 

SuperfamilyBuprestoidea 160 

Family Buprestidae (A. Davies) 160 

SuperfamilyDryopoidea 169 

Family Ptilodactylidae (L. LeSage) 169 

Family Heteroceridae (L. LeSage) 169 

Family Limnichidae (L. LeSage) 171 

Family Dryopidae (L. LeSage) 771 

Family Elmidae (L. LeSage) 172 

Family Psephenidae (L. LeSage) 174 

SuperfamilyElateroidea L74 

Family Artematopidae (Y. Bousquet) 174 

Family Elateridae (Y. Bousquet) 175 

Family Throscidae (Y. Bousquet) 186 

Family Eucnemidae (Y. Bousquet) 186 

Superfamily Cantharoidea 188 

Family Lycidae (J. McNamara) 188 

Family Phengodidae (J. McNamara) 190 

Family Lampyridae (J. McNamara) 190 

Family Cantharidae (J. McNamara) 192 

SuperfamilyDermestoidea 195 

Family Derodontidae (D.E. Bright) 195 

Family Nosodendridae (J.M. Campbell) 196 

Family Dermestidae (Y. Bousquet) 196 

SuperfamilyBostrichoidea 199 

Family Bostrichidae (J. McNamara) 199 

Family Lyctidae (J. McNamara) 20L 

Family Anobiidae (J. McNamara) 201 

Family Ptinidae (Y. Bousquet) 205 

Superfamily Lymexyloidea 206 

Family Lymexylidae (J.M. Campbell) 206 

SuperfamilyCleroidea 206 

Family Trogossitidae (Y. Bousquet) 206 

Family Cleridae (J. McNamara) 208 

Family Melyridae (D.E. Bright) 2LI 

Superfamily Cucujoidea 213 

Family Sphindidae (J.M. Campbell) 213 

Family Nitidulidae (J. McNamara) 2I4 

Family Rhizophagidae (Y. Bousquet) 2I8 

Family Cucujidae (Y. Bousquet) 219 

Family Cryptophagidae (Y. Bousquet) 22I 

Family Languriidae (J.M. Campbell) 223 

Family Erotylidae (J.M. Campbell) 225 

Family Phalacridae (J.M. Campbell) 226 

Ill

Family Cerylonidae (J.M. Campbell) 227 

Family Corylophidae (J.M. Campbell) 228 

Family Coccinellidae (J. McNamara) 229 

Family Endomychidae (J.M. Campbell) 287 

Family Lathridiidae (Y. Bousquet) 239 

Family Biphyllidae (J.M. Campbell) 247 

Family Byturidae (J.M. Campbell) 24I 

SuperfamilyTenebrionoidea 242 

Family Mycetophagidae (Y. Bousquet) 242 

Family Tetratomidae (L. LeSage) 243 

Family Ciidae (J. McNamara) 244 

Family Melandryidae (L. LeSage) 245 

Family Mordellidae (J. McNamara) 248 

Family Rhipiphoridae (J.M. Campbell) 250 

Family Colydiidae (Y. Bousquet) 251 

Family Zopheridae (J.M. Campbell) 252 

Family Tenebrionidae (Y. Bousquet and J.M. 

Campbell) 253 

Familr' Lagriidae (J.M. Campbell) 261 

Famils Cephaioidae (J.M. Campbell) 262 

Famil r Meloidae (J.M. Campbell) 269 

Famil r Oedemeridae (J.M. Campbell) 266 

Famil r' Prostomidae (J.M. Campbell) 267 

Famil Synchroidae (L. LeSage) 267 

'z 

Familr Mycteridae (J.M. Campbell) 267 

Famil,r Boridae (J.M. Campbell) 268 

Famil r Pythidae (J.M. Campbell) 268 

Famil r Pyrochroidae (J.M. Campbell) 269 

Famil,r Salpingidae (J.M. Campbell) 270 

Family Anthicidae (Y. Bousquet) 27t 

Family Pedilidae (Y. Bousquet) 274 

Family Aderidae (Y. Bousquet) 275 

Family Scraptiidae (L. LeSage) 275 

SuperfamilyChrysomeloidea 277 

Family Cerambycidae (J. McNamara) 277 

Family Bruchidae (J. McNamara) 300 

Family Chrysomelidae (L. LeSage) 3C,1 

SuperfamilyCurculionoidea 323 

Family Nemonychidae (J. McNamara) 323 

Family Anthribidae (J. McNamara) 324 

Family Ithyceridae (J. McNamara) 32:,5 

Family Attelabidae (J. McNamara) 3tl6 

Family Rhynchitidae (J. McNamara) 326 

Family Brentidae (J. McNamara) 327 

Family Apionidae (J. McNamara) 328 

Family Curculionidae (J. McNamara) 329 

Family Scolytidae (J. McNamara) 35€; 

Family Platypodidae (J. McNamara) :]65 

ORDERSTREPSIPTERA 366 

SuborderStylopidia 366 

Family Corioxenidae (S.B. Peck) 366 

Family Halictophagidae (S.B. Peck) 336 

Family Stylopidae (S.B. Peck) 367 

References 368 

Index 397 

iv

LIST OF CONTRIBUTORS 

Y. Bousquet, Biosystematics Research Centre, 

Agriculture Canada, Ottawa, Ontario K1A 0C6 

D.E. Bright, Biosystematics Research Centre, 


J.M. Campbell, Biosystematics Research Centre, 


A. Davies, Biosystematics Research Centre, 


P.J. Johnson, Department of Entomology, 

University of Wisconsin, Madison, Wisconsin 

53706 

D.J. Larson, Department of Biology, Memorial 

University of Newfoundland, St. John's, 

Newfoundland AlB 3X9 

L. LeSage, Biosystematics Research Centre, 


J. McNamara, Biosystematics Research Centre, 

Agriculture Canada, Ottawa, Ontario KlA 0C6 

S.B. Peck, Department of Biology, Carleton 

University, Ottawa, Ontario KlS 586 

R.E. Roughley, Department of Entomology, 

University of Manitoba, Winnipeg, Manitoba 

R3T 2N2 

ACKNOWLEDGMENTS 

We wish to thank the following persons for 

reviewing sections of the checklist: R.L. Aalbu 

(Tenebrionidae); R.S. Anderson (Silphidae, 

Curculionidae); F.G. Andrews (Lathridiidae); LS. 

Askevold (Chrysomelidae); F.M. Atton (Gyrinidae); 

E.U. Balsbaugh (Chrysomelidae); R. Baranowski 

(Leiodidae); W.F. Barr (Cleridae); E.C. Becker 

(Elateroidea); H.P. Brown (Dryopoidea); D. 

Chandler (Anthicidae, Pselaphidae, Melandryidae, 

Tetratomidae, Synchroidae, Scraptiidae); J. Cicero 

(Lampyridae); W. Connell (Nitidulidae); M. 

Daccordi (Chrysomelinae); J.G. Edwards 

(Synetinae); D.G. Furth (Chrysomelidae); R.E. 

Gordon (Coccinellidae): L.H. Herman 

(Staphylinidae); A.T. Howden (Curculionidae); H. 

Howden (Scarabaeidae); C. Johnson (Ptiliidae); J.B. 

Karren (Chlamisinae); J.M. Kingsolver (Bruchidae); 

J.L. Krysan (Chrysomelidae); S. Laplante (Cleridae, 

Cerambycidae); J.F. Lawrence (Melandryidae, 

Tetratomidae, Synchroidae, Scraptiidae); I. Lobl 

(Scaphiidae); W.V. Miller (Heteroceridae); A.R. 

Moldenke (Clytrinae); H.G. Nelson (Dryopoidea); 

A.F. Newton Jr. (Leiodidae, Staphylinidae); S. 

Oygur (Gyrinidae); J. Pakaluk (Corylophidae); J.D. 

Pinto (Meloidae); D. Pollock (Mycteridae, Boridae, 

Pythidae, Salpingidae); V. Puthz (Steninae); E.G. 

Riley (Cassidinae); W.T. Schultz (Eumolpinae); T.N. 

Seeno (Chrysomelidae); R.B. Selander (Meloidae); 

S.A. Slipinski (Cerylonidae); C.L. Staines 

(Hispinae); W.E. Steiner Jr. (Phalacridae); J.B. 

Stribling (Ptilodactylidae); M. Thayer 

(Staphylinidae); M.C. Thomas (Cucujidae, 

Prostomidae); C.A. Triplehorn (Tenebrionidae); R.E. 

White (Anobiidae, Chrysomelidae); D.P. Wooldridge 

(Limnichidae); D.K. Young (Scirtidae, 

Pyrochroidae); P. Zwick (Leiodidae). 

Thanks are also extended to B.F. Carr, J.L. 

Carr, and D.W. Langor, for providing several 

provincial records; to D. Beauchamp, B. Walter, L. 

Robillard, and R. Hutchinson, for typing the checklist; 

and to A. Smetana, for checking the termination 

of all species-group names.

SUMMARY 

All species and subspecies of beetles known to 

occur in C.anada and Alaska are listed along with 

their synonyms; these names are arranged alphabetically 

t,nder the respective genus-group names. 

Valid genus-group and family-group names are 

listed in p aylogenetic order. No new taxa, combinations, 

or synonyms are proposed. Species distributions 

are presented in tabular form, indicating the 

presence or absence of the taxa in each province 

and territory, in Labrador, and in Alaska. 

Introduced and holarctic species are highlighted. 

References to pertinent identifrcation keys are indicated 

under the corresponding supraspecific taxon. 

The work also includes a brief introduction for 

every family treated and an index to all supraspecifrc 

names. 

RESUME 

L'ouvrage r6pertorie toutes les espdces et sousespdces 

dt,: Col6optdres connues du Canada et de 

l'Alaska arnsi que leurs synonymes; ces noms sont 

6num6r6s par ordre alphab6tique d la suite du nom 

du groupe-genre correspondant. Les noms valides 

des group':-genre et groupe-famille sont pr6sent6s 

par ordre phylog6n6tique. Cette liste ne propose 

aucun nouveau taxon, combinaison ou synonyme. 

La r6partibion des espdces est pr6sent6e sous forme 

tabulaire, indiquant la pr6sence ou l'absl:nce des 

taxa dans chaque province et territoire, le 

Labrador et l'Alaska. On met en relief les espbces 

introduites ou holarctiques et on mentionne, sous 

le taxon suprasp6cifique correspondra.nt, les 

r6f6rences contenant des tableaux de d61;ermination 

utiles. L'ouvrage comprend aussi une courte 

introduction sur chaque famille consid6r'6e ainsi 

qu'un index des noms suprasp6cifrques. 

vi

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INTRODUCTION 

Beetles represent the most diverse group of 

living organisms on earth. More than 250 000 

species have been described, and the actual number 

may easily exceed a million. Given such staggering 

frgures, it is not surprising that Coleoptera can be 

found neally everywhere, from the tropics to the 

Arctic and from caves to the summit of high mountains. 

Coleoptera species are particularly abundant 

on vegetation, inside rotten wood, under the bark of 

dead trees, and in soil, ground litter, fungi, dung, 

carrion, rodent burrows, and nests. Several groups 

are adapted to life in fresh or brackish water. 

Although some dytiscids can venture into salt water 

and some species do complete all stages of their life 

cycle in the intertidal zone where they are periodically 

immersed, no beetles have become completely 

adapted to marine life. Beetles are important to 

humans because many species are beneficial in the 

biological control of pest arthropods and weeds or 

are major pests of stored products, fabrics, forests, 

wood prodructs, and crops. 

No comprehensive checklist of Coleoptera is 

available for Canada and Alaska. Leng's catalogue 

(1920) and its supplements (Leng and Mutchler 

1927, 1933; Blackwelder 1939a; Blackwelder and 

Blackwelder 1948) as well as the world-based 

Coleopterorum Catalogus list most of the species 

that occur in Canada and Alaska. However, these 

publications are out of date for many groups, and 

their distribution data are incomplete. Beaulne 

(1913-1929) started a checklist of the beetles of 

Canada but never completed it, and as pointed out 

by Lindroth (1966) for the ground beetles, many of 

Beaulne's distributional records are inaccurate. 

This checklist is intended to provide the correct 

names of the beetles that occur in Canada and 

Alaska, give a brief survey of the distribution of the 

taxa in the area covered, and lead the user's to the 

proper publications for the identification of the 

taxa 

the checklist of Strepsiptera has been added 

at the end simply because the group has often been 

included in the Coleoptera. 

FORMAT 

With some exceptions reflecting the author's 

personal perspective, the concept of the families 

used in this checklist follows that of Lawrence 

(1982a). N,najor changes concern the trachypachids, 

limulodids, scaphidiids, lyctids, lagriids, pedilids, 

rhynchitids, scolytids, and platypodids, which are 

treated as distinct families, and the leptinids, 

trogids, and geotrupids, which are included respectively 

within the Leiodidae and the Scarabaeidae 

for the last two groups. The superfamily concepts 

and the listing of the families follow Lawrence 

(1982a). 

The classification 

and information for each 

family are the responsibility of the respective 

authors. No new taxa, combinations, or synonyms 

are proporied. Variety and form names published 

before 1961 are given subspecifrc rank unless they 

were deemed to be synonyms or their status has 

been changed (article +5 (g) of the International 

Code of Zoological Nomenclature, 3rd ed.). 

Varieties and forms published after 1960 are 

omitted. New distributional records are listed but 

not identified as such. 

The arLthors of species-group names are listed in 

parentheses when the original combination differs 

from the one used in the checklist. To avoid confusion, 

initials are included for several authors, such 

as C.R. Sahlberg, R.F. Sahlberg. J.R. Sahlberg, J. 

MiiIIer, O.F. Miiller, W. Horn, J.E. LeConte, and O. 

Fabricius. LeConte, Horn, and Fabricius stand for 

J.L. LeConte, G. Horn, and J.C. Fabriciusr respectively. 

An asterisk [*] placed after a species-group 

name indicates that the taxon is Holarctic, a dagger 

ttl that it is introduced in North Americra, and a 

double dagger t{l that it is probably introduced in 

North America. 

AII synonyms based on material collected in 

North America north of Mexico should be included 

and are listed in italics. Synonyms of Hola:nctic and 

introduced species based on material frorri outside 

North America have been included occasionally, 

particularly when the name has been in use in 

North American literature. Misidentifications, 

spelling errors, and invalid emendations are not 

llited. The ending of all species-group names has 

been checked to conform with the gender of their 

respective genera. 

Valid genus-group names are in capital letters, 

their synonyms in italics. All synonyms crf genusgroup 

names whose type species are indigengu-s to 

North America north of Mexico should be included. 

Some of the synonyms whose type species are not 

indigenous to the continent are included also when 

the name has been used in North American literature. 

References to keys are indicated under the corresponding 

supraspecific taxon. It should be noted 

that only references including keys useful for the 

determination of all or most Canadian taxa of the 

group treated are considered. Keys published

efore the 1930s are usually out of date, and most of 

them are therefore omitted. 

An abbreviation of two letters is used to indicate 

Canadian provinces and Alaskan records (see 

section entitled "Abbreviations and symbols"). 

Labrador is considered to be a distinct geographic 

entity. The distributional citation in the checklist is 

from west to east. 

The distributional records are based on published 

records and on specimens in the Canadian 

National Collection. For some families. records 

from other collections have been included. Doubtful 

records from the literature that apparently are 

based on misidentifications or mislabeled specimens 

are rejected. 

A question mark (?) inserted in a geographic 

column indicates that the taxon was recorded in the 

area, but the record is questionable. The absence of 

a geographic entry after a species-group name indicates 

that the taxon was recorded in Canada. but 

the province was not specified. 

SUBSEQUENT REVISIONS OF THE CHECKLIST 

This publication is set as a data base that can 

be easily updated to maintain an accurate summary 

of our current knowledge of the beetles of Canada 

and Alaska. The authors intend to maintain a corrected 

version of the checklist and to republish it 

whenever we have accumulated enough changes. 

The authors would appreciate the assistance of all 

users of this checklist in calling errors and omissions 

to their attention. We would also welcome 

receiving specimens that represent new provincial 

or Alaskan records. Specialists and other users 

may send material to the Biosystematics Research 

Centre for confirmation, and with the sender's permission, 

for incorporation as voucher specimens in 

the Canadian National Collection. 

AB B REV 

*TI r.rl' O N S AND SYMB O L S 

AB Alberta 

AK Alaska 

BC British Columbia 

LB Labrador 

MB Manitoba 

NB New Brunswick 

NF Newfoundland 

NS Nova Scotia 

NT NorthwestTerritories 

ON Ontario 

PE Prince Edward Island 

PQ Quebec 

SK Saskatchewan 

YK Yukon Territorv 

H Holarctic 

Introduced in North America 

I 

* 

t 

Holarctic 

Introduced in North America 

f Probably introduced in North America 

$ No subgeneric name available

Micromalthidae 

Order Coleoptera 

Suborder Archostemata 

FamilyCUPEDIDAE reticulatedbeetles 

J.M. Campbell 

Three species of Cupedidae occur in Canada. 

This small, primitive family of beetles contains only 

30 extant species around the world, but it is well 

represented by fossils dating back to the early 

Triassic. l,ittle is known about the biology of the 

species-the larvae apparently bore into wood, and 

the adults iue collected from foliage or under bark. 

The farnily Cupedidae is well known taxonomically. 

Atk.ns (1963) revised the world species, and 

Neboiss (1984) the species of Cupes. Ponornarenko 

(1969) discussed the fossil origins of the Coleoptera 

with particular reference to the srrborder 

Archostemata and the family Cupedidae, rerviewing 

the fossil and extant species of the family. Atkins 

(1979) cataloged the North American taxa of the 

family. 

BC (1); oN (2);PQ (1) 

Subfamily CUPEDINAE 

Tribe Priacmini 

Genus PRIAC MA LeConte 

P. serrata (LeConte) 

BC 

Tribe Cupedini 

Genus CUPEII Fabricius 

C. capitatus tr abricius 

ON 

Genus TENO T{ERGA Neboiss 

T. concolor (Si estwood) 

cinerea (flay) 

oculata (Oasey) 

t r iI ineat c; (Melsheimer) 

ON PQ 

Family MICROMAL'THIDAE telephone-pole beetles 

J.M. Campbell 

This fz: mily contains only one species, which 

was descri ced from the eastern United States but 

has subserluently become established in western 

Canada, C uba, Brazil, Hawaii, Hong Kong, and 

South Afrir:a. The larvae are wood borers and are 

found in decaying oak logs and in moist, creosotetreated 

timbers and poles. The life cycle of 

Micromalthus debilis is one of the most complex 

known to occur in Coleoptera. The larvae are paedogenetic, 

producing either living larvae or a single 

egg that develops through a different larval type to 

produce an adult male (Pringle 1938). 

Atkins (1982) cataloged the species for North 

America. 

BC (1) 

Genus MICROMALTHUS LeConte 

M. debilis Le(jonte BC

Trachvoachidae 

Suborder Adephaga 

Family RIIYSODIDAE wrinkled bark beetles 

Y. Bousquet 

The family Rhysodidae includes only two and larvae live under the bark of dead trees and in 

species in Canada. The adults are cylindrical, elon- rotting wood. 

gate beetles resembling in habitus some species of Bell (1970) revised the North American species. 

the families Colydiidae or Cucujidae. Both adults eC (1);ON (1) 

Genus OMOGLYMMIUS Ganglbauer 

Key to species: Bell and Bell (1982) 

(Subgenus BOREOGLYMMIUS Bell & Bell) 

O. americanus (Castelnau) - ON 

orolas (Newman) 

e ra r at u s (Audinet-Serville) 

Genus CLINIDIUM Kirby 

Key to North American species: Bell and Bell (1975) 

(Subgenus ARCTOCLINIDIUM Bell) 

C. calcaratum LeConte - BC 

Family TRAC IIYPAC HIDAE 

Y. Bousquet 

This family includes two genera: Trachypachus, decades, however, some authors have highlighted 

which has three species in western North America many character states that suggest that 

and one species in Eurasia, and Systolosorno Solier, Trachypachus and Sysfolosoma are more closely 

which has two species in Chile. Members of related to Hydradephaga than to Carabidae (see 

Trachypachus are usually found on dry, more or Kavanaugh (1986) for a different point of view). 

Iess shaded soil, often under dead leaves. Nothing Lindroth (1961) has redescribed the two species 

is known about their feeding habits. 

that inhabit Canada. 

This group has been included within the 

Carabidae for a long time. During the last two aK (1);YK (1); NT (1); BC (2); aB (1); SK (1) 

Genus TRACHYPACHUS Motschulsky 

T. holmbergi Mannerheim 

inermis Motschulsky 

oregonus Casey 

specularis Casey 

T. gibbsii LeConte 

alticola Casey 

c alifor nicu s Motschulsky 

AK YK .\T BC AB SK 

_BC

Carabidae 

Family CARABIDAE ground beetles 

Y. Bousquet 

With 9,t6 species and subspecies in Canada and 

Alaska, thr,: family Carabidae is an important element 

of thr: Coleoptera fauna in the northern part 

of North A:nerica. In Canada and Alaska members 

of these species occur in most terrestrial habitats as 

well as in .marshes and bogs; a few species live on 

trees. Adults of most species are carnivorous or 

omnivorous,; only those of a few groups seem to be 

primarily phytophagous; larvae of most species are 

probably ca.rnivorous, but those of a few groups are 

known to be phytophagous or parasitic. 

The Carabidae of North America are here 

grouped in 36 tribes. The classification used is basically 

that of Lindroth (1961-1969), with a few modifications 

re,flecting recent taxonomic developments 

in the family. In recent years, some authc,rs haYe 

included the rhysodids within Carabidae, but based 

on larval characters I prefer to exclude thenL; on the 

other hand, some others consider the cicinclelids to 

be a distinct family, a view with which I do not 

agree. Thanks to the work of Wallis (1961) lor tiger 

beetles and Lindroth (1961-1969), the carabid fauna 

of Canada and Alaska is well known taxonomically. 

Bell (1990) recently published a key to the North 

American genera. 

AK(237); YK (164); NT (207); BC (a83); AB (386); 

SK (314); MB (350); ON (503); PQ (451); NB (2a3); 

NS (24a); PE (137); LB (.94); NF (175) 

' II (84); I 

(46) 

Tribe Cicindelini 

Genus OMUS Eschscholtz 

Leptontut Casey 

Megomus Casey 

O. audouini audouini Reiche 

aequicorr,.is Casey 

audens Casey 

borealis ()asey 

delicatuhn Casey 

distans Casey 

oregonentiis Casey 

paruulus Casey 

rugipenn,.s Casey 

tacomae ()asey 

thoracicus Casey 

uandykei W. Horn 

O. dejeani Rei:he 

foueatus l)asey 

robustus Casey 

BC 

BC 

Genus CICIN.)ELA Linn6 

(Subgenus CICINDELA s. str.) 

(Pachydetn Rivalier) 

(Tribonia Rivalier) 

C. ancocisconr nsis Harris 

carolinae Casey 

dowiana Casey 

eriensis (iasey 

C. decemnotata Say 

albertina Casey 

lantzi lJ.a.rcts 

C. depressula depressula Casey AK 

scapularis Casey 

C. duodecimgrrttata Dejean 

bucolica tlasey 

edmontor'.ensis Can:r 

hudsonic,z Casey 

proteus Birby 

C. formosa folmosa Say 

fletcheri Criddle 

luxuriosa Casey 

manitoba Lelrg 

YK BC SK 

BC 

BC AB SK 

AB 

SK 

PQ 

MB ON PQ NB NS 

PE LB 

NF

Carabidae 

C. formosa generosa Dejean 

latecincta Godd 

C. formosa gibsoni Brown 

C. fulgida westbournei Calder 

elegans Calder 

C. hirticollis athabascensis Graves 

C. hirbicollis couleensis Graves 

C. hirticollis rhodensis Calder 

nigrita Davis 

shermani Casey 

C. hirticollis shelfordi Graves 

C. lengi versuta Casey 

C. limbalis limbalis KIug 

arnoena LeConte 

awemeana Casey 

eldorensis Casey 

m il ita r is Varas-Arangua 

spreta LeConte 

C. limbata hyperborea LeConte 

C. lirnbata nympha Casey 

C. longilabris longilabris Say 

albilabris Kirby 

nouaterrae Leng 

terracensis Casey 

C. nebraskana nebraskana Casey 

calgaryana Casey 

canadensis Casey 

chamberlaini Knars 

montdna LeConte 

splssllorsis Casey 

uteana Casey 

C. oregona guttifera LeConte 

audax Casey 

oregonella Casey 

sterope Casey 

C. oregona oregona LeConte 

oualipennis Casey 

quadripennis Casey 

sonoma Casey 

C. parowana wallisi Calder 

azurea Cald,er 

C. patruela patruela Dejean 

C. pugetana Casey 

nigerrimoides Hatch 

C. purpurea audubonii LeConte 

auguralis Casey 

graminea Schaupp 

nigerrina Leng 

C. purpurea purpurea Olivier 

marginalis Fabricius 

ramosa Gistel 

C. repanda novascotiae Vaurie 

C. repanda repanda Dejean 

baltimorensis Hetbst 

d.uncaniKnats 

hoosieri Mares 

maehleri Robinson 

unijuncta Casey 

C. scutelaris criddlei Casey 

C. scutelaris lecontei Haldeman 

C. scutelaris scutelaris Say 

C. sexguttata denikei Brown 

C. sexguttata sexguttata Fabricius 

guttata Emmons 

harrisii Leng 

illinoensis Mates 

kansanal(nans 

leuettei Casey 

quadriguttata Davis 

tridens Casey 

uarians Ljungh 

uiolacea Fabricius 

BC 

AB 

AB 

AB 

SK 

SK 

SK 

bt\ 

NT_ABSKMB 

AB SK MB 

AK YK NT BC AB SK MB 

AK YK NT BC 

AB SK MB 

_BCABSKMB 

NT BC AB SK MB 

BC AB SK MB 

BC 

BC 

BC 

BC AB SK MB 

BC 

NT BC 

AB 

MB ON PQ 

lln- : 

_ONPQNBNS_LB 

SK MB ON 

oN pe 

-NS-- 

LB 

NF 

NF 

oN pe NB NS PE LB NF 

ON PQ 

ON PQ 

PQ 

PQ 

NB 

NB 

NS 

_MB 

-MBONPQ 

_ABSK _MBON 

oN PQ NB NS 

NS PE 

NS PE LB 

NF

Carabidae 

C. tranqueba'ica borealis Harris 

C. tranqueba:ica kirbyi LeOonte 

C. tranquebartca roguensis Harris 

C. tranqueba ica tranquebarica 

Herbst 

crinifron.s Casey 

horiconezsis Leng 

minor Lt:ng 

obliquat':.t Dejean 

turbulenta Casey 

u ir idula Varas-Arangua 

uulgaris Say 

uulgaris;,ninor Haris 

wichitan.a Casey 

C. tranquebar:ica vibex Horn 

(Subgenus CI CINDELIDIA Rivalier) 

C. punctulatri punctulata Olivier 

boulderensis Casey 

chihuah:ne BaLes 

fontinar,ra Casey 

jenisoni llistel 

prolixa (.)asey 

(Subgenus C' T,INDERA Westwood) 

(Plectogratr. ha Rivalier) 

C. tericola cinctipennis LeConte 

cyanella LeOonte 

C. terricola irrperfecta LeConte 

C. terricola tcrricola Say 

pusilla E ay 

sayanellt Casey 

(Subgenus EI,LIPSOPTERA Dohktourow) 

C. cuprascenr, LeConte 

amnicohL Casey 

mundukr Casey 

C. lepida Dejean 

insomnil: Casey 

C. nevadica k.nausi Leng 

YK 

NT BC 

NT_ABSK 

-BCABsx 

BC 

_AB 

Bcy 

nae 

MB ON pe Ns NS PE NF 

AB SK MB ON PQ NB 

SK 

AB SK 

AB SK 

Tribe Carabini 

SK MB 

MB 

MB ON 

MB 

PQ 

Genus CALOSOMA Weber 

Key to Norbh American species: Gidaspow (1959) 

(Subgenus CitLOSOMA s. str.) 

(Callipa'o Motschulsky) 

(S y nc alc so ma Breuning) 

C. lrigidum l.jrby 

Ieuettei Casey 

(Subgenus CTTLODREPA Motschulsky) 

C. scmlator (Fabricius.t 

C. wilcoxi Letlonte 

(Subgenus C;T LLITROPA Motschulsky) 

(Paratrooa Vacher de Lapouge) 

C. exlernum Sayt 

longipenne Dejean 

(Subgenus CI{RYSOSTIGMA Kirby) 

(Lyperostenia Vacher de Lapouge) 

(Tapinosthenes Kolbe) 

C. calidum (Fabricius) 

comes Casey 

concretum Casey 

expansu,n Casey 

laticolle Casey 

stellatur,z Casey 

BC 

SK MB ON PQ NB NS PE 

_ONPQ 

_ONPQ 

ON 

NT SK MB ON PQ NB NS NF 

10

Carabidae 

C. cancellatum Eschscholtz 

aenescens LeConte 

esuriens Casey 

praestans Casey 

rectilaterum Casey 

sagax Casey 

transuersum Casey 

C. lepidum LeConte 

C. obsoletum Say 

microstictum Casey 

C. tepidum LeConte 

caelator Casey 

cogitans Casey 

indigens Casey 

irregulare Walker 

pellar Casey 

semicupreum Casey 

(Subgenus CAILISTHENES Fischer von Waldheim) 

(Callistenia Vacher de Lapouge) 

(Isostenia Vacher de Lapouge) 

BC 

BC 

AB SK MB 

AB SK MB 

C. luxatum Say 

debile (Casey) 

diffractum (Casey) 

etcaratum (Casey) 

opacum (G6hin) 

parowanunr (Casey) 

pimelioides (Walker) 

pustulosum (Casey) 

reflexum (Casey) 

semotum (Casey) 

striatulum LeConte 

striatum Breuning 

tahoense Van Dyke 

tegulatum (Casey) 

utense (Casey) 

uiator (Casey) 

(?) zimtnermanl (LeConte) 

C. moniliatum ( Leconte) 

bicolor (Walker') 

concinnum (Casey) 

laqueatum LeConte 

uancouuericum (Csjkl) 

C. monticola I Casey) 

neuadense (Casey) 

C. subaeneum Chaudoir 

C. wilkesii (Leconre) 

Genus CARABUS Linn6 

(Subgenus CARABUS s. str.) 

(Cryocarabus Vacher de Lapouge) 

(H omoeocar abus Reitter) 

(.Lic hnocarab us Reitter) 

(Neocarabus Hatchl 

C. chamissonis Fischer AK YK 

von Waldheim 

brachyderus Wiedemann 

groenlandicus Dejean 

washingtoni Casey 

C.goryt Dejean 

clarkei Blwnenthal 

limbatus Say 

C. granulatus granulatus Linn6t 

C. granulatus hibernicus Lindrotht 

C. maeander maeander Fischer AK 

von Waldheimx 

atlanticus Vacher de Lapouge 

excatenatus Kraatz 

excostatus Kraatz 

h udsonicus Motschulsky 

NT_AB 

_BCAB 

BC SK 

BC AB SK 

SK 

NT BC AB SK 

MB ON PQ 

ON 

MB ON 

MB ON 

LB 

PQ 

_NBNSNF 

PQ NB NS PE LB NF 

11

Carabidae 

lapilayi Castelnau 

lecontei G6hin 

simoni I{eyden 

C. nemoralis O.F. Mullert 

canadensis Vacher de Lapouge 

C. vinctus W':ber 

carinatu s Dejean 

georgiae Cslki 

interruptus Say 

ligatus Getrnat 

BC AB SK oN PQ 

ON 

NB NS 

PE 

NF 

(Subgenus DIOCARABUS Reitter) 

C. taedatus a-gassii LeConte 

canadic,ts Roescke 

coloradensis Breuning 

gladiatcr Motschulsky 

montant:.cus Casey 

oregonetrcis LeConte 

patulicotlis Casey 

stocktonensis Casey 

C. taedatus taedatus Fabricius 

bacciuorus Fischer von Waldheim 

bicanaliceps Casey 

francisutnus Casey 

rainieri Van Dyke 

seriatus Wiedemann 

C. truncatico llis truncaticollis 

Eschscholtz 

ala s ken :,:i s Basilewsky 

lutshniA ianus Basilewsky 

AK 

AK 

tiK 

tT( 

NT 

BC 

SK 

MB ON PQ 

(Subgenus H 0MICARABUS G6hin) 

C. serratus Say 

canadensis LeConte 

lineato p unct atzs Dej ean 

lolumi l4otschulsky 

uegasensis Casey 

BC 

AB SK MB ON PQ NB NS PE 

NF 

(Subgenus T,\NAOCARABUS Reitter) 

C. sylvosus s.ylvosus Say 

lhermin:eri Dejean 

ON PQ 

(Subgenus MEGODONTUS Solier) 

C. vietinghol hi vietinghoffii 

Adarr:s* 

AK 

YK 

NT 

Tlibe Cychrini 

Genus SPHAIRODERUS Dejean 

S. canadensiri canadensis Chaudoir 

blancha,di Leng 

palpalis Motschulsky 

S. Iecontei Ie:ontei Dejean 

bicarinatus LeConte 

diffractt.ts Casey 

niagarensis Castelnau 

S. nitidicoliis brevoorti LeConte 

granulosus Chaudoir 

S. nitidicollis nitidicollis 

Chevlolat 

SK 

oN PQ 

MB ON PQ NB 

MB ON PQ 

PQ 

NS 

NS 

NF 

NF 

Genus SCAI'HINOTUS Dejean 

(Subgenus STIAPHINOTUS s. str.) 

(Anabol'rs Gishel') 

(Irichroa Newman) 

(Megaliridia Casey) 

S. elevatus coloradensis Van Dyke 

S. viduus (Dejean) 

ir r eg ula r i s Beutenmuller 

leonard:'.i (Harris) 

yu_ 

pe 

NB 

t2

Carabidae 

(Subgenus NOMARETUS LeConte) 

Key to species: Gidaspow (1973) 

S. bilobus (Sayt MB ON PQ NB NS 

(Subgenus PSEUDONOMARETUS Roeschke) 


S. merkelii (Horn) - BC 

idahoensis (Webb) 

S. regularis (LeContet - BC 

S. relictus (Horn) - BC 

(Subgenus STENOCANTHARUS Gistel) 

(Pemphus Motschulsky) 


S. angusticollis (Fischer AK BC 

von Waldheim) 

longipes Casey 

nigripennis Roeschke 

opacus Casey 

uelutinus M6n6tries 

(Subgenus NEOCYCHRUS Roeschke) 


S. angulatus (Harris) - BC 

rnaritimus Van Dyke 

(Subgenus BRENNUS Motschulsky) 


S. johnsoni Van Dyke - BC 

hlahowyae Perrault 

S. marginatus (Fischer AK BC AB 

von Waldheim) 

columbianus (Casey) 

confusus (Casey) 

cupripennis (Casey) 

/ollor (Roeschke) 

fulleri (Horn) 

gracilis (G6hin) 

insularis (Casey) 

montanicus (Casey) 

oregonus (Casey) 

wrangelli (Casey) 

Genus CYCHRUS Fabricius 

Key to North American species: Gidaspow (1973) 

C. rickseckeri LeConte - BC 

C. tuberculatus Harris - BC 

pustulosus Casey 

Genus LEISTUS Friilich 

Key to North American species: Erw.in (1970a) 

(Subgenus LEISTUS s. str.) 

Tlibe Nebriini 

L. fernrgineus (Linn6)f - NF 

(Subgenus NEOLEISTUS Erwin) 

L. ferruginosus Mannerheim AK BC AB 

ferrugineus Dejean 

nigropiceus Casey 

Genus PELOPHILA Dejean 

P. borealis (Paykull)* 

californica Motschulsky 

AK YK NT BC AB SK MB ON PQ - LB NF 

e s c hs c holtzi (Mannerheim) 

shermani Casey 

ulkei Horn 

P. rudis (LeConte) NT BC AB - MB ON - NF 

13

Carabidae 

Genus NEBRIA Latreille 

N. acuta acuta Lindroth 

N. arkansana edwardsi Kavanaugh 

N. brevicollis (Fabricius)t 

N. campbelli Kavanaugh 

N. charlottae Lindroth 

N. crassicorn is crassicornis 

Van fryke 

N. crassicornis intermedia Van Dvke 

N. diversa LeConte 

liuidaLeConte 

townsendi Casey 

N. frigida R.I'. Sahlberg* 

paruula J. Sahlberg 

reducta'Casey 

uiridis E:lorn 

N. gebleri cai;cadensis Kavanaugh 

N. gebleri gebleri Dejean 

melanariaHatch 

N. gouleti Kavanaugh 

N. gregaria Fischer von Waldheim 

cuneata casey 

N. gyllenhali castanipes (Kirby)* 

curtulata Casey 

(?) elias Motschulsky 

labradorica Casey 

moesta leCorrte 

prominens Casey 

N. haida Karanaugh 

N. hudsonica LeConte 

N. kincaidi k ncaidi Schwarz 

columbi'ma Casey 

N. lacustris I lcustris Casev 

expansa Casey 

N. lituyae Kavanaugh 

N. Iouiseae Kavanaugh 

N. mannerheimii Fischer 

von S'aldheim 

coruallis Casey 

hippisleyi Casey 

oregona Casey 

N. meanyi meanyi Van Dyke 

N. meanyi sylvatica Kavanaugh 

N. metallica Fischer von Waldheim 

pacifica Chaudoir 

N. nivalis gar;pesiana Kavanaugh 

N. nivalis nir alis (Paykull)* 

bifaria llIannerheim 

mollis l!:lotschulsky 

N. obliqua obliqua LeOonte 

incerta Casey 

obtusa I.,eConte 

testaceipes Casey 

texana Casey 

N. pallipes Say 

N. piperi Van Dyke 

N. sahlbergii sahlbergii Fischer 

von Waldheim 

aleuta\anDyke 

uiolacea Motschulsky 

N. schwarzi schwarzi Van Dvke 

N. suturalis l.eConte 

longula LeConte 

nimbosa Casey 

N. virescens .florn 

breuis Casey 

AK 

_v 

AK 

_BC 

AK YK I.TT _ 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

v_ 

AB 

AB 

? 

AK 

AK YK NT BC AB SK MB ON 

_BC 

_YKNTBCABSK 

AK 

BC 

AK 

AK 

-BC 

AKYK_BC 

-BC 

AK 

AKYK_BC 

AK--eC 

AKYK-BCAB 

BC 

BC 

:" 

BC 

AB 

SK 

yu_ 

MB ON 

_PQ 

_PQ 

PQ 

PQ 

PQ 

9* PQ 

ON 

PQ 

NB 

NB NS 

PE 

LB 

LB 

LB 

NF 

v 

T4

Carabidae 

Tlibe Notiophilini 

Genus NOTIOPHILUS Dum6ril 

N. aeneus (Herbst) 

porrectus Say 

N. aquaticus (Linn6)x 

hardyi Putzeys 

N. biguttatus (Fabricius)t 

N. borealis Harris* 

N. directus Casey 

lanei lJatc]l 

N. intermedius Lindroth 

N. nitens LeConte 

N. semistriatus Say 

americanus }{atris 

coloradensis Casey 

confusus LeConte 

punctatus LeConte 

N. simulator FalI 

euanescens Casey 

N. sylvaticus Eschscholtz 

AK 

AK 

AK 

AK 

YK 

YK- 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

AB 

AB 

SK 

oN PQ 

MB ON PQ 

MB ON PQ 

MB PQ 

MB ON PQ 

NB NS 

NB NS 

NB NS 

NS 

_LB 

LB NF 

LB 

LB 

NF 

NF 

NF 

Tribe Loricerini 

Genus LORICERA Latreille 

Key to species: Ball and Erwin (1969) 

(Subgenus LORICERA s. str.) 

L. decempunctata Eschscholtz 

L. pilicornis congesta Mannerheimx 

L. pilicornis pilicornis 

(Fabricius)* 

californica LeConte 

neoscotica LeConte 

semip unct ata Eschscholtz 

sierraeYanDyke 

uteana Casey 

AK 

AK 

AK YK 

_BC 

NT BC AB SK 

MB ON PQ 

NB NS 

is 

NT 

Genus OPISTHIUS Kirby 

O. richardsoni Kirby 

Tribe Opisthiini 

AK YK NT BC AB SK 

TYibe Elaphrini 

Genus DIACHEILA Motschulsky 

Arctobia C.G. Thomson 

D. arctica amoena (Faldermann)* 

americana Motschulskv 

subpolaris LeOonte 

D. polita (Faldermann)* 

AK 

YK 

NT 

LB 

Genus BLETHISA Bonelli 

HelobiumLeach 

Rhaphiona Fischer von Waldheim 

Key to species: Goulet and Smetana ( 1983) 

B. catenaria W.J. Brown* 

B. inexspectata Goulet and Smetana 

B. julii LeConte 

B. multipunctata aurata 

Fischer von Waldheimx 

hudsonica Casey 

B. oregonenis LeConte 

acutangula Chaudoir 

columbica Casey 

B. quadricollis Haldeman 

arnericana Hatris 

AKYKNT_MB 

AK 

NT_ABSKMBONPQNBNSNF 

AK YK NT BC AB SK MB ON PQ NB NS PE - 

BC 

NT BC AB SK MB ON PQ NB NS PE LB 

NF 

NF 

15

Carabidae 

Genus ELAI'HRUS Fabricius 

Key to species: Goulet (1983) 

(Subgenus ARCTELAPHRUS Semenov) 

E. lapponicus lapponicus Gyllenhal* AK YK NT BC AB - MB ON PQ - LB 

obscurit.tr I{trby 

E. lapponicus obliteratus AK 

ManrLerheim 

(Subgenus NEOELAPHRUS Hatch) 

E. cicatricosrrs LeConte PQ 

rhodear; us Casey 

E. clairuillei KJrby AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

/rosll H.ppisley 

lynni Pierce 

politus ,eCorrte 

torreyer,rsis Tanner 

E. fuliginosus Say AB - MB ON PQ 

E. olivaceus LeOonte NT BC AB SK MB ON PQ NB NS PE - NF 

(Subgenus ELAPHRUS s. str.) 

(.Trichel tphrus Semenov) 

E. americanrrs americanus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

bitubercsus Casey 

gratiost^s Mannerheim 

interme lizs Kirby 

punctat ssjmzs LeConte 

sinuatu i LeConte 

E. americanrrs syivanus Goulet BC AB 

E. californicrLs Mannerheim AK - NT BC AB SK MB ON PQ NB NS PE 

hesperh'.s Casey 

similis LeCorrte 

E. lecontei Crotch - NT BC AB SK MB ON 

deuinctt:s Casey 

spisslco''nls Casey 

E. parviceps Van Dyke* AK - NT - MB ON 

E. ruscarius Say ON PQ 

texanus Casey 

E. tubercular us Maklin* AK YK NT 

(Subgenus E LAPHROTERUS Semenov) 

(Elaphriatus Semenov) 

E. angustico lis angusticollis AX YK NT 

R.F. liahlberg* 

E. purpuranr' Hausen 

pallipes Hotr' 

AK YK NT BC AB 

Genus OMO.?HRON Latreille 

Key to North American species: Benschoter and Cook (1956) 

Tribe Omophronini 

O. americanus Dejean AB SK MB ON PQ NB NS PE - NF 

fontinal;.s Casey 

illustris Casey 

iridescens Casey 

locustri'; Casey 

lengi Casey 

proximus Chandler 

soil Kirby 

tanneri Chandler 

texanus Casey 

O. Iabiatus (Fabricius) NS 

O. ovalis Horn - BC AB SK 

concinnits Casey 

frater Casey 

gemma'Jasey 

O. robustus llorn AB ON 

breuipen.nis Casey 

decoloratus Fall 

16

Carabidae 

O. tesselatus Say 

ellipticus Casey 

lecontei Dejean 


Genus PROMECOGNATHUS Chaudoir 

P. crassus LeOonte 

contractus Casey 

corpulentus Casey 

grandiceps Casey 

Tribe Promecognathini 

BC 

Genus SCARITES Fabricius 

S. subterraneus Fabricius 

beckwithii Stephens 

denticollis Chaudoir 

interruptus (Herbst) 

spinipes (Sulzer) 

subtemeus Bonelli 

Tribe Scaritini 

ON 

Genus PASIMACHUS Bonelli 

P. elongatus LeConte AB SK 

angustulus Casey 

euanescens Casey 

pirnalis Casey 


uernicatus Casey 

Genus DYSCHIRIUS Bonelli 

Key to North American species: Bousquet (1g88) 

(Subgenus AKEPHORUS LeConte) 

D. obesus LeConte - BC 

(Subgenus DYSCHIRIUS s. str.) 

D. affinis Fall 

duplicatus Fall 

D. alticola Lindroth 

D. aratus LeConte 

D. brevispinus LeConte 

D. campicola Lindroth 

D. compactus Lindroth 

D. criddlei FaIl 

D. er5rthrocerus LeConte 

D. foigidus Mannerheim 

aeneolus LeConte 

D. globulosus (Say) 

paruus LeConte 

D. haemorrhoidalis (Dejean) 

D. hiemalis Bousquet 

D. integer LeOonte 

apicalis LeOonte 

basalis LeConte 

nigripes LeConte 

tr an s m arin u s Mannerheim 

D. interior Fall 

arizonicus Van Dyke 

D. laevifasciatus Horn 

D. larochellei Bousquet 

D. longulus LeConte 

D. montanus LeConte 

thompsoni Hatcll 

AK YK 

AK YK 

AK 

AK 

Tribe Clivinini 

-BC 

_BCAB 

_:rf 

NT BC 

-BCAB 

AB 

SK 

NT BC AB 

NT BC AB SK 

_AB 

BC AB 

BC AB 

BC AB 

SK 

SK 

MB 

MB 

-ONPQ 

MB 

MB 

ON PQ 

MB 

-oNPQ 

_ONPQ 

MB ON PQ 

-oN 

MB ON PQ 

MB ON PQ 

MB_PQ 

MB ON PQ 

NB NS 

_LB 

NB NS PE 

NB 

NS 

LB 

NF 

NF 

_NF 

_v 

L7

Carabidae 

D. nigricornis Motschulsky* AK YK 

secretus Fall 

D. pacificus .l,indroth 

D. pallipenn:s (Say) 

D. perversus Fall 

desertut Fall 

D. pilosus LrConte 

hispidu,; LeConte 

D. planatus ,indroth 

D. politus (Dejean)* AK YK 

aureolu.; Notman 

D. quadrima :ulatus Lindroth 

D. sellatus LeConte 

D. setosus LcConte AK 

alternatus Hatch 

D. sextonj Bousquet 

D. sphaerico lis tSayr 

subpunt:tatus Hatch 

D. subarcticus Lindroth AK YK 

D. tenuispin rs Lindroth 

D. timidus Lindroth 

D. truncatus LeConte AK 

NT 

_BC 

AB SK 

AB 

AB SK MB 

_BCABSKMB 

NT 

nc 

BC 

Nr sc 

AB 

AB 

AB 

AB 

AB 

MB 

MB 

SK MB 

SK MB 

sr me 

-me 

SK MB 

ON 

ON 

ON 

ON 

ON 

ON 

PQ 

PQ 

PQ 

PQ NB 

PQ NB 

PQ NB 

NS 

NS 

NS 

PE 

PE 

po 

NF 

NF 

NF 

Genus CLIVINA Latreille 

(Subgenus S 0MICLN'INA Kult) 

C. dentipes.l)ejean 

coruina Putzeys 

confusa LeConte 

fissipes Putzeys 

georgiana LeConte 

ON 

(Subgenus PARACLIVINA Kult) 

C. bipustula' a (Fabrici us) 

quadrintaculafo (Palisot de Beauvois) 

ON 

(Subgenus CLIVINA s. str.) 

C. americana Dejean 

acuducta Haldeman 

cordata Putzeys 

ludouiciana Putzeys 

C. collaris tIlerbst)t 

elongatt Randall 

randalli LeConte 

C. fossor il,inn6)t 

C. impressifrons LeConte 

C. oregona Fal) 

BC 

_BCAB 

-BC 

ON PQ 

MB ON PQ 

ON PQ 

_-ONPQ 

NB 

NS 

NS PE 

NF 

Genus SCHIZOGENIUS Putzeys 

Key to North American species: Whitehead (1972) 

(Subgenus SCHIZOGENIUS s. str.) 

S. amphibius (Haldeman) 

frontalis (LeConte) 

S. depressus LeConte 

S. ferrugineus Putzeys 

sulcatu;; (LeConte) 

S. lineolatus {Sayr 

S. litigiosus Fall 

S. sulcifrons Putzeys 

BC 

ec 

sx 

ON 

ON 

ON 

PQ 

pe 

PQ 

PQ 

NB 

TYibe Brachininr 

Genus BRA(IHINUS Weber 

Key to North American species: Erwin (19706) 

(Subgenus NEOBRACHINUS Erwin) 

B. cordicollis Dejean 

gracilis Blatchley 

- ON 

leptocerus Chaudoir 

uelox LeConte 

PQ NB 

18

Carabidae 

B. cyanipennis Say MB ON PQ NB 

cephalotes Dejean 

rejectus LeConte 

B. cyanochroaticus Erwin 

B. fumans (Fabriciust 

affinis LeConte 

- 

BC - SK MB ON PQ 

BC AB - MB ON PQ 

amplipennis Bates 

atbarae Stehr 

cyanopterus LeConte 

suffians LeConte 

tabasconus Bates 

B. janthinipennis (Dejean t - ON PQ 

pumilio LeConte 

B. medius Harris - BC - ON PQ 

minutus }J.arris 

B. ovipennis LeConte - ON PQ 

B. quadripennis Dejean - BC AB SK MB ON 

stygicornis Say 

I schern ihhii Mannerheim 

B. tenuicollis LeConte 

ballistarius LeConte 

- ON PQ 

similis LeConte 

B. texanus Chaudoir AB 

Genus METRIUS Eschscholtz 

Tribe Paussini 

M. contractus contractus - BC 

Eschscholtz 

Tribe Gehringiini 

Genus GEHRINGIA Darlinston 

G. olympica Darlington 

NT BC AB 

Tribe Broscini 

Genus BROSCUS Panzer 

B. cephalotes (Linn6tf NS PE 

Genus MISCODERA Eschscholtz 

Leiochiton Curtis 

M. arctica (Paykull)x 

americana Mannerheim 

AK YK NT BC AB SK MB ON PQ - NS PE LB NF 

hardyi Chaudoir 

Genus BROSCODERA Lindroth 

B. insignis tMannerheim) AK BC 

Genus ZACOTUS LeConte 

Z. matthewsii LeConte AK BC 

angustus Casey 

fre deric ki Nunenmacher 

subopacus Hopping 

Genus BLEMUS Dejean 

Las iotrec hu s Ganglbauer 

Tlibe Trechini 

B. discus (Fabriciustf - ON PQ NB 

Genus TRECHUS Clairville 

(Subgenus TRECHUS s. str.) 

19

Carabidae 

T. apicalis Motschulskyx 

borealis Schaeffer 

brumalts Casey 

canader'.sis Putzeys 

fuluus (",eCorte) 

micans .LeConte) 

pallescens Casey 

puritan,ts Casey 

rhodensis Casey 

T. chalybeus Dejean* 

brachycierus Jeannel 

califor n ic us Motschulsky 

tahoensi.s Casey 

T. crassiscapus Lindroth 

T. obtusus Erichsont 

T. oregonensis Hatch 

T. ovipennis Motschulsky 

T. quadristriatus (Schrank)f 

T. rubens (Fabricius)t 

T. tenuiscapr.rs Lindroth 

AK YK NT 

AK 

BC 

BC 

AB 

---ec-- -BCAB 

y_ _ BC_ _ 

_YK_BCAB 


_ :.T _ 

_ONPQ 

PQN 

NS 

NS 

eNs 

LB NF 

_:uIu 

pe- y 

Genus ASAI HIDION des Gozis 

Tachypi:.ts Dejean 

A. alaskanurn Wickham 

A. yukonens'r Wickham 

AK YK NT 

AKYK_BCAB 

Tribe Bembidiini 

Genus BEM.IIIDION Latreille 

(Subgenus CHRYSOBRACTEON Netolitzky) 

(Parabrzcteon Notman) 

B. alaskense Lindroth 

B. balli Lindroth 

B. carinula ( jhaudoir 

tubercu o/um Notman 

B. co1villenstr Lindroth 

B. foveum Motschulsky* 

bryanti Carr 

(.?) glabr iusculum Motschulsky 

grahamiHatch 

B. hesperium Fall 

binariu;n Casey 

B. inaequale inaequale Say 

arenari,r,m Dejean 

lacustre LeConte 

B. inaequale opaciceps Casey 

B. lapponicum Zetterstedt* 

pugetanumFall 

B. levettei Casey 

carrianlm Casey 

B. lorquinii (lhaudoir 

tacomar Clssy 

B. punctatostriatum Say 

(?) sigiliare Say 

stigmat:icum Dejean 

B. zephyrurr Fall 

marginosum Casey 

tristicuium Casey 

(Subgenus BRACTEON Bedel) 

B. bowditchii LeConte 

(Subgenus ODONTIUM LeConte) 

B. confusum Hay'ward 

aeneorubrum Casey 

marquettense Casey 

nitidult:m Dejean 

AK 

.AK 

AKYKNTBC_SK 

AK 

AK_NT 

---ABix 

YK NT BC AB SK 

AK YK NT 

BC 

BC AB SK 

BC AB SK 

BC 

BC 

BC 

BC 

BC 

AB 

SK 

SK 

ff,fS On 

MB 

MB ON 

MB 

PQ 

oN PQ 

oN PQ 

AB SK MB ON PQ 

NB 

PQ NB NS 

NB NS 

NB 

PE 

NF 

LB NF 

20

Carabidae 

B. coxendix Say 

snowi Casey 

unicum Casey 

uenator Casey 

uigilans Casey 

ueesi Hatch & Ortenburger 

(Subgenus PSEUDOPERYPHUS Hatch) 

(B r a cteomimus Lindroth) 

B. chalceum Dejean 

antiquum Dejean 

dilatatum (LeConte) 

B. honestum Say 

basale (LeConte) 

platyderum Chaudoir 

NT_AB 

BC 

SK MB 

AB SK MB ON PQ NB NS 

NF 

ON PQ NB 

(Subgenus OCHTHEDROMUS LeConte) 

B. americanum Dejean 

illini Casey 

B. bifossulatum (LeOonte) 

cheyennense Casey 

ferreum Casey 

nuperum Casey 

regestum Casey 

suffiatum Casey 

(Subgenus EURYTRACHELUS Motschulsky) 

(Eudromus l\trby) 

(P la ty t r a c hel us Motschulsky) 

B. interventor Lindroth AK 

B. nitidum (Kirby) AK 

edolatum Casey 

(Subgenus HYDRIUM LeConte) 

B. levigatum Say 

delawarense Casey 

BC AB SK 

-oN 

MB ON 

PQ 

PQ 

YK NT BC AB SK MB ON PQ 


AB 

(Subgenus METAILINA Motschulsky) 

B. dyschirinum LeOonte AK 

agitabile Casey 

aleneanum Casey 

atrolucens Casey 

heechelum }Iatch 

perturbatum Casey 

speculinum Casey 

B. lampros tHerbsilf 

B. properans ( Stephens tt 

YK 

BC 

BC 

AB 

PQ NB NS 

PE 

NF 

NF 

(Subgenus PI{YLA Motschulsky) 

B. obtusum Audinet-Servillet 

(Subgenus LIONEPFIA Casey) 

Key to species: Erwin and Kavanaugh (1981) 

oN PQ 

B. castum Casey AK 

brumale Casey 

nescium Casey 

serenum Casey 

uaciuum Casey 

B. disjunctum Lindroth 

B. erasum LeConte 

lasciuum Casey 

lubricum Casey 

probatum Casey 

B. lindrothellum Erwin & Kavanaugh AK 

B. lummi Erwin & Kavanaugh 

B. sequoiae Lindroth 

BC 

BC AB 

BC 

BC 

BC 

(Subgenus ACTEDIUM Motschulsky) 

B. lachnophoroides Darlington 

AB SK 

(Subgenus TRECHONEPHA Casey) 

B. iridescens I LeConte) 

BC 

AB 

2I

Carabidae 

atnicum Casey 

deceptor Casey 

fabrum Oasey 

impium Casey 

obliuiost:.m Casey 

p ar allel c'colle (Motschulsky) 

repens Casey 

uolatile t)asey 

R 

J 

B. simplex Hayrvard 

essexense Casey 

PQ 

$ 

B. kuprianov:.i Mannerheim 

adultum Casey 

b iimp r es s u m Mannerheim 

bucolicum Casey 

dilutum Casey 

funereurz LeConte 

maehlint Hayward 

ouipennt (Motschulsky) 

(Subgenus PI,ATAPHODES Ganglbauer) 

B. arcticum Lindroth* 

B. brachytho:'ax Lindrothx 

B. complanul om (Mannerheim) 

paruulutn Notman 

B. compressum Lindroth* 

B. farrarae Ilatch 

B. haruspex l.)asey 

B. incertum ( Motschulsky) 

ampliatum Casey 

blanditum Casey 

(?') breue (Motschulsky) 

improuit,um Casey 

liuidulu n Casey 

saturatum Casey 

testatun Casey 

tetr ag ly 1 

t u m (Mannerheim) 

B. manninge:rse Lindroth 

ochropu:; Lindroth 

B. occultator Notman 

B. quadrifoveolatum Mannerheim 

iller Casey 

B. rosslandicrm Lindroth 

B. viator Casey 

(Subgenus P.I,ATAPHUS Motschulsky) 

(.Melomt'lus Casey) 

(M icromelomalus Casey) 

(Trachelonepho Casey) 

B. basicorne Notman 

B. carolinense Casey 

heeneanzm Casey 

B. curtuiaturn Casey 

decrepitu.m Casey 

effetum Casey 

flebile Casey 

timefactum Casey 

B. gebleri turbatum Casey 

conflictt:m Casey 

B. gordoni Lindroth 

B. gratiosum Casey 

pallidumHatch 

B. hyperboraeorum Munster* 

B. kalumae Lindroth 

B. planatum (LeConte) 

adjutor Casey 

aequalis: (Walker) 

solutum Casey 


AK YK 

AK YK NT BC 

AK 

BC 

AK YK NT 

AK 

_BC 

AKYK?BC 

AK YK 

AK 

AK YK 

AK YK 

AK 

AK 

AK 

YK 

YK 

YK 

YK 

BC 

NT BC 

NT 

NT 

AB 

AB 

AB 

_AB 

BC AB 

BC AB 

BC 

BC AB 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

SK 

9* 

PQ NB NS 

_PQ_NS 

_PQNBNS 

AB--oNpeNnNs--Nr 

LB 

NF 

NF 

22

Carabidae 

B. planiusculum Mannerheim AK BC 

B. rufrnum Lindroth AK YK - BC .AB 

B. rusticum lenensoides Lindroth AK YK - BC 

B. nrsticum rusticum Casey - ON PQ NB NS - LB NF 

notmani Casey 

B. siemicola Casey - BC 

B. stillaguamish Hatch - BC 

B. sulcipenne hyperboroides AK YK - BC 

Lindroth 

B. sulcipenne prasinoides Lindroth - BC .AB - MB - PQ NB LB NF 

(Subgenus BLEPHAROPLATAPHUS Netolitzky) 

B. hastii C.R. Sahlbergx AK YK NT BC MB ON PQ - LB 

(Subgenus TRICHOPLATAPHUS Netolitzky) 

B. planum (Haldeman) - ON PQ NB NS 

filicorne Casey 

guexii Chatdo\r 

uulsum Casey 

B. rolandi Fall - ON PQ NB NS 

(Subgenus HIRMOPLATAPHUS Netolitzky) 

B. concolor (Kirby) AK YK NT BC .AB SK MB ON PQ NB NS NF 

longulum (LeConte) 

subaeneum (LeConte) 

B. nigrum Say - ON PQ NB NS PE 

facile Casey 

morosum Casey 

B. quadrulum LeConte 

aegrotum Casey 

- YK - BC .AB 

callidum Casey 

porrectum Casey 

tartareurn Casey 

tritum Casey 

uiridinigrum Casey 

B. recticolle LeConte 

oblatum Casey 

- BC AB 

pertinax Casey 

tetr ag onoderum Chaudoir 

umbraticola Casey 

B. salebratum (LeOonte) NT BC .AB SK MB ON PQ NB NS NF 

consessor Casey 

inopinum Casey 

mackinacense }latc}r 

purpurascens (LeConte) 

(Subgenus HYDRIOMICRUS Casey) 

B. quadratulum Notman PQ NB NS NF 

proximum Notman 

B. semistriatum (Haldeman) - ON PQ - NS 

(Subgenus BEMBIDIONETOLITZKYA Strand) 

(Daniela Netolitzky) 

B. mckinleyi carneum Lindroth - BC AB 

B. mckinleyi mckinleyi FalI AK YK 

$ 

B. lenae Csiki* AK YK NT 

intermedium Poppius 

$ 

B. commotum Casey 

seclusum Casey 

B. nebraskense LeConte 

denueranum Casey 

gouanicum Casey 

tractabile Casey 

$ 

_BCAB 

_BCAB 

B. actuosum Casey 

debilicolle Casey 

- BC AB 

urgens Casey 

23

Carabidae 

B. nevadense Illke 

uiaticun.: Casey 

B. plagiatum (Zimmermann) 

B. striola (LeConte) 

angustit:r Casey 

modulatum Casey 

shastani.cum Casey 

uancouu zri Casey 

BC 

BC 

ON 

(Subgenus PiIRYPHANES Jeannel) 

B. dauricum Motschulskyx 

B. grapii Gyl enhal* 

nitens Casey 

picipes (t{irby) 

scrutatum Casey 

seducturn Casey 

B. lacunariurn (Zimmerrnann) 

histricurn Casey 

militare Casey 

B. platynoides Hayward 

insopita:zs Casey 

merens li)asey 

optatum Casey 

sedulutr, Casey 

B. stephensi lrotch'l 

canadense Hayward 

B. texanum ( lhaudoir) 

cogitans Casey 

inquietum Casey 

B. yukonum lall* 

AK YK NT 


_BC 

AK YK NT BC 

MB 

MB ON 

-oN 

MB 

MB 

ON 

PQ 

PQ 

PQ 

PQ 

NB NS is NF 

NB NS PE NF 

$ 

B. bimaculat rm (Kirby) 

B. postremur:r Say 

B. sordidum rKirby) 

B. umiatenser Lindroth 

AK YK 

AI< YK 

AK 

NT BC 

NT BC 

AB 

AB 

SK MB 

SK MB 

ON 

-PQ 

oN PQ 

NB 

LB 

(Subgenus PIIRYPHUS Stephens) 

B. bruxellense Wesmaelt 

B. femoratunr Sturm* 

B. obscurellu m (Motschulsky)x 

albidipezne Casey 

caducunt Casey 

fuscicrus (Motschulsky) 

lucidum (Mannerheim) 

mixtum LeConte 

parowar.um Casey 

petulans Casey 

(?) subinflatum (Motschulsky) 

B. petrosum ilttuense Lindroth 

B. petrosum'letrosum Gebler* 

castaliu;n Casey 

exiguiceps Casey 

lepusculum Casey 

lucidum (LeConte) 

substric,um (LeConte) 

wenatchee Hatch 

B. poppii sch,rlleri Lindroth 

B. rupicola (lGrby) 

auersan:; Casey 

B. sejunctum sejunctum Casey 

B. sejuncturn semiaureum Fall 

fenderi llatch 

B. tetracolum Sayt 

nactum Casey 

AK YK 

AK 

AK YK 

AK 

AK 

NT BC 

AB 

Nr ec AB SX 

NT BC 

NT 

_BC 

_BC 

AB 

AB 

sx lle 

sr l.rn 

SK 

-PQNBNSPE 

NB NS 

ONPQNB_PE 

ON PQ 

oN- 

_:a 

oN PQ 

Nn lls pr 

YNS 

NB NS 

LB 

LB 

NF 

NF 

NF 

NF 

NF 

$ 

B. transverszrle Dejean 

acoman,tm casey 

animatu.m Casey 

YK_BC SK MB ON PQ 

NB NS 

LB 

NF 

24

Carabidae 

(?) erosum (Motschulsky) 

ercursum Casey 

haplogonum Chaudoir 

ma nner heimii (LeConte) 

marinicum Casey 

perspicuum (LeConte) 

sarpedon Casey 

tuolumnae Casey 

$ 

B. scopulinum (Kirby)x 

bellulum Casey 

gelidum (LeConte) 

(Subgenus LIOCOSMIUS Casey) 

B. mundum (LeConte) 

b ifa s c iat um (Motschulsky) 

hilare Casey 

(Subgenus PERYPHODES Casey) 

B. ephippigerum (LeConte) 

B. salinarium Casey 

$ 

B. constricticolle Hayward 

B. nudipenne Lindroth 

(Subgenus EUPETEDROMUS Netolitzky) 

B. graciliforme Hayward 

B. immaturum Lindroth 

B. incrematum LeConte 

arcuatum LeConte 

fortunatum Casey 

gulosum Casey 

/ezgi Notman 

mobile Casey 

nubiferum Casey 

oblectans Casey 

sernotum Casey 

(Subgenus LEUCIIYDRIUM Casey) 

B. tigrinum LeConte 

$ 

B. consimile Hayward 

B. flohri Bates 

henshawi Hayward 

B. obtusidens Fall 

B. scudderi LeConte 

spissicorne Casey 

AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

AK YK Nt 

--nc 

BC 

_BC 

_BCAB 

-BC 

_AB 

_AB 

EC AB 

BC 

SK 

SK 

sr 

MB 

MB 

,AB SK MB 

.AB SK MB 

,AB SK MB 

,AB SK MB 

-oNPQ _ONPQNBNSNF 

MB ON PQ NB NS PE LB NF 

$ 

B. dejectum Casey 

aberti}].atch 

fidele Casey 

tencenti Hatch 

B. mormon Hayward 

B. obtusangulum LeConte 

cornix Casey 

(Subgenus NOTAPHUS Stephens) 

B. aeneicolle (LeConte) 

B. castor Lindroth 

B. coloradense Hayward 

albertanum Casey 

amplipenne Casey 

imperitum Casey 

prociduum Casey 

B. cordatum (LeConte) 

placabile Casey 

B. debiliceps Casey 

_BC 

_ BC ,{B 

NT 

NT 

,AB 

SK 

SK 

MB 

MB 

BC .q,B SK MB 

_ ,A,B SK MB ON PQ NB NS 

BC .{B SK MB 

BC 

MB ON 

NF 

25

Carabidae 

B. dorsale Say 

B, graphicum Casey 

lassulutn. Casey 

B. indistinctum Dejean 

barbarae Casey 

caudex (;asey 

conse nt a neum (LeConte) 

curiosun,, Casey 

derisor C.iasey 

deuinctum Casey 

erpositurn Casey 

extricatum Casey 

formale l}asey 

franciscc:,num Casey 

ornatellt'.m Casey 

reconditum Casey 

B. insulatum (LeConte) 

caliginosum Casey 

B. intermeditLm (Kirby) 

B. nigripes (IrJrby)x 

concitatt.Lm Casey 

quadrah colle (Mannerheim) 

retracturn Casey 

B. patruele Dejean 

contristans Casey 

editum Oasey 

exclusunt Casey 

fenisex (.asey 

fraternun LeOonte 

imitator Casey 

marcidum Casey 

mediocn: Casey 

monstra,'.um Casey 

plectile Oasey 

pos ticun,'. (Haldeman) 

subexiguum Casey 

B. rapidum (LeConte) 

docile Ctsey 

fugitans Casey 

negligeni Casey 

sociale ( asey 

B. semipunct etum (Donovan)* 

accuratt'.m Casey 

prouoanitm Casey 

B. umbratum (LeConte) 

u ar iolos u m ( Motschulsky) 

B. variegaturn Say 

postfascntum Hamilton 

B. versutum LeConte 

AK 

AK 

AK 

AK 

_BCAB 

NT BC AB 

NT 

BC 

BC 

AB 

BC AB 

YK NT BC AB 

YK 

NT 

NT 

BC 

BC 

BC 

AB 

AB 

SK MB 

SK MB ON 

SK MB 

SK MB ON 

SK MB ON 

SK MB ON PQ 

PQ 

PQNB_PELB 

SK MB ON PQ NB 

SK 

AB SK 

ON 

ON PQ 

oN PQ 

NB NS 

NB 

PE 

NF 

NF 

$ 

B. constricturn (LeConte) 

ciuile Ca.sey 

festinans Casey 

uernula Casey 

B. contracturr Say 

B. viridicolle (LaFert6-S6nectdre) 

hamiferum Chaudoir 

particeps Casey 

NT 

AB 

_PQ 

SK MB ON 

NB NS 

NB NS 

PE 

PE 

NF 

R 

B. oberthueri Hayward 

(Subgenus FIJRCACAMPA Netolitzky) 

B. affine Say 

decipiens Dejean 

fallax Dejearr 

thespis Casey 


ON 

R 

J 

B. impotens Casey 

BC SK MB ON NB 

26

Carabidae 

aestiuum Casey 

frugale Casey 

gratuitum Casey 

imbelle Casey 

ind,igens Casey 

pictum (LeConte) 

pullulum Casey 

uirgatulum Casey 

B. mimus Hay'ward pellax Casey 

- SK MB ON PQ NB NS PE - NF 

rotundiceps Casey 

B. timidum (LeConte) NT BC AB SK MB ON 

oregonum Casey 

sord.idulum Chaudoir 

B. versicolor (LeConte) terracense Casey 

AK - NT BC AB SK MB ON PQ NB NS PE LB NF 

tolerans Casey 

wisconsinium Casey 

(Subgenus EMPHANES Motschulsky) 

(Omala Motschulsky) 

B. diligens Casey parabile Casey 

- BC AB SK MB 

uilescans Casey 

B. vile (LeConte.) - BC 

(Subgenus BEMBIDION s. str.) 

(Lopfto Stephenst 

B. mutatum Gemminger & Harold arillare (LeConte) 

AK YK NT BC AB SK MB ON PQ - NS NF 

uegetum Casey 

B. praecinctum LeConte alutaceum Hatch 

AB SK MB 

ueridicum Casey 

B. quadrimaculatum dubitans (LeConte) 


caseyi Letg 

crurale (LeConte) 

gregale Casey 

pugetanurn Casey 

sapphicum Casey 

tenax Casey 

B. quadrimaculatum oppositum Say AB SK MB ON PQ NB NS PE - NF 

(Subgenus SEMICAMPA Netolitzky) 

B. convexulum Hayrvard AK YK - BC AB 

nouellum Casey 

B. morulum LeConte AK YK NT BC AB SK MB NF 

browni Lindroth 

B. muscicola Hayward - SK MB ON PQ NB NS PE 

B. praticola Lindroth - BC - SK - ON PQ NB 

B. roosevelti Pic AB SK MB 

blaisdelli Casey 

concinnum Blaisdell 

p er conc innunt Blaisdell 

B. semicinctum Notman - ON PQ NB NS 

(Subgenus DIPLOCAMPA Bedel) 

(Paralopha Casey) 

B. transparens (Gebler)* AK YK NT BC AB SK MB ON pQ NB NS pE LB NF 

edmontonense Casey 

sulcatum (LeConte) 

trepidum (LeConte) 

(Subgenus TREPANEDORIS Netolitzky) 

B. acutifrons LeConte AK - NT BC AB SK MB 

m i c r o r e t ic u la t u m Hatch 

B. canadianum Casey NT - AB SK MB ON PQ 

B. concretum Casey AK YK NT BC AB SK MB ON PQ NB NS PE - NF 

congruens Casey 

habile Casey 

27

Carabidae 

B. connivens (LeConte) 

digressum Casey 

elizabetl:ae Hatch 

B. fortestriatrm (Motschulsky) 

cdutum .LeConte) 

concurre'.ns Fall 

peregrinum Casey 

utnbrati:um Casey 

B. frontale (l.eOonte) 

B. pseudocaubum Lindroth 

B. siticum Cz.sey 

adolescew Casey 

BC 

AK YK NT BC SK MB ON PQ NB NS PE NF 

_BC_SKMBONPQ 

_BCABSKMBONPQ 

_BC 

)u 

NS 

PE 

(Subgenus AIvIERIZUS Chaudoir) 

B. oblonguloiJes Lindroth 

B. oblongulurn (Mannerheim) 

crassicorne (Casey) 

heeni (C*ey') 

B. spectabile (Mannerheim) 

longicor.rc (Casey) 

B. wingatei Filand 

AK 

AK 

_BC 

BC 

BC 

oN PQ 

NB NS 

LB 

NF 

Genus PHR\PEUS Casey 

P. ricksecker (Hayward) 

rutilinut Casey 

BC 

Genus ELAP HROPUS Motschulsky 

Barytac,rys Chaudoir 

Tachyur a Motschulsky 

E. anceps (Lt.Conte) 

E. capax (Le0onte) 

E. dolosus (LeConte) 

apacheanus { Casey) 

audax (l,eConte) 

rapar (I eConte) 

E. granarius (Dejean) 

glossem'r (Casey) 

E. incurwus ( Jay) 

parallel 1s (Casey) 

rhodens'.s (Cslki) 

E. parvulus (Dejean)t 

E. saturatus (Casey) 

E. tripunctatus (Say) 

barnesi (Stehr) 

laredoanus (Casey) 

seruzs ({lasey) 

E. vernicatus, (Casey) 

laetificus (Casey) 

unionis (Csiki) 

E. vivax (LeConte) 

mendar (LeConte) 

E. xanthopur (Dejean) 

ancilla rCasey) 

famelia.Ls (Casey) 

loaipennis (Casey) 

NT 

BC 

BC 

AB 

SK 

SK 

MB ON PQ 

-oN 

-oN 

-oN 

MB ON 

PQ 

PQ 

---oNFeNeNs-- 

_ONPQNBNS 

_NS 


oN PQ 

ON 

YNS 

NB NS 

NS 

PE 

Genus TACFiYTA Kirby 

Key to species: Erwin (1975) 

T. angulata Casey 

T. falli (Hayrvard) 

T. nana inornata (Say) 

arizonica Casey 

californ;;,ca Casey 

collaris Casey 

debilicol,lis Casey 

picipes lktby 

riu ulari s (Motschulsky) 

T. nana kirb ri Casey 

YK 

BC AB 

BC 

BC 

SK MB ON PQ 

ON PQ 

BC AB ON PQ 

fu NS 

28

Carabidae 

Genus POROTACHYS Netolitzky 

P. bisulcatus tNicolaitt - ON PQ NB 

frontalis (Hayward) 

Genus POLYDERIS Motschulsky 

Microtachys Casey 

P. laevis {Sayt 

congesta (Casey) 

- ON PQ - NS 

flumenalis (Casey) 

recta (Casey) 

troglodytes (Dejean) 

unistriata (Casey) 

unistriolata (Cslki) 

Genus TACHYS Stephens 

Isotachys Casey 

T. bryanti Lindroth ,\B 

T. halophilus Lindroth ,\B SK MB 

T. vittiger LeConte - BC 

marginellus LeConte 

Genus PARATACHYS Casey 

P. edax (LeConte) 

P. oblitus {Casey) 

- BC 

appalachius (Casey) 

- ON PQ 

(Casey) 

iowensis (Casey) 

cuneatus (Casey) 

gentilis 

obliquus (Casey) 

P. proximus tSayt - ON PQ 

nubifer {Casey) 

P. rhodeanus (Caseyr - ON PQ 

P. scitulus (LeConte) 

pallescens (Casey) 

p a I I i d i u s c u I u s (C sikl) 

Genus MIOPTACIfYS Bates 

T ac hy menis Motschulsky 

- ON PQ NB 

M. flavicauda (Say) 

ma rg inicollis (Motschulsky) 

- BC r\B ON PQ 

r eflexicoll is (Motschulsky) 

Genus NOMIUS Castelnau 

Haplochile LeConte 

Tlibe Nomriini 

N. pygmaeus (Dejeant - BC AB - MB ON PQ 

Genus PSYDRUS LeConte 

Monillopatrobzs Hatch 

P. piceus LeConte - BC AB SK MB ON PQ 

punctdtus (Hatch) 

Genus PLATYPATROBUS Darlington 

Tribe Patrobini 

P. lacustris Darlington NT - l\B SK MB ON PQ NB 

Genus PATROBUS Dejean 

Geo p a t r ob u s Darlington 

N eo pat r ob u s Darlington 

P. lossifrons (Eschscholtz)x AK BC l\B 

dimorp hic us Darlington 

/uluus Mannerheim 

latiusculus Chaudoir 

long i uen I ri s Mannerheim 

29

Carabidae 

P. foveocollis (Eschscholtz)* AK YK NT BC AB SK MB ON PQ - LB NF 

ang usticollis Mannerheim 

insularis Casey 

Iaeuiceps Casey 

obt usiu sc u I us Chaudoir 

tenuis (l'eConte) 

P. lecontei Chaudoir 


- BC AB SK MB - PQ NB - NF 

grauidus Darlington 

ruftpes LeConte 

p. tongicbrnis (Say) NT BC AB SK MB ON PQ NB NS PE - NF 

americanus Dejean 

P. septentrionis Dejlan* AK YK NT BC AB SK MB ON PQ - LB NF 

hyperboreus Dejean 

Iabradorinus Casey 

minuen.; Casey 

tritus Casey 

P. stygicus Chaudoir AK YK NT BC AB SK MB ON PQ - LB NF 

Genus DIPLOUS Motschulskv 

Platidiu.s Chaudoir 

D. aterrimus, (Dejean) AK YK NT BC AB 

breuiceps (Casey) 

coloradensis (Casey) 

fulcratus (LeConte) 

reflexus (Casey) 

tenuitarsis (Casey) 

D. californic rs (Motschulsky) 

breuius;ylas (Casey) 

- BC 

lnciszs (Casey) 

Iatipenris (Casey) 

rectus (Jasey) 

sietan es (Casey) 

strenuus (Casey) 

troc han te ricus (LeConte) 

D. frlicornis iCasey) - BC 

D. rugicollis (Randall) 

longipalpis (Notman) 

PQ NB NS 

Genus PLAI]IDIOLUS Chaudoir 

Patroboidea Van Dyke 

P. vandykei Kurnakow AK YK - BC AB 

rufus (Van Dyke) 

Genus LOK\NDRUS LeConte 

Key to Ncrth American species: Allen (1972) 

Tlibe Pterostichini 

L. minor (Chaudoir) - ON 

inquietrc Casey 

L. velocipes Casey - ON 

Genus STOMIS Clairville 

S. pumicatus (Panzer)t NS 

Genus LOP}{OGLOSSUS LeConte 

L. scrutator (LeConte) - ON PQ 

canadensis (Chaudoir) 

Genus POECILUS Bonelli 

Feronio Latreille 

Leconteus Lutshnik 

(Subgenus POECILUS s. str.) 

P. chalcites (Say) - ON PQ 

micans Chaudoir 

soyi (BruII6) 

30

Carabidae 

P. corvus (LeContet AB SK MB 

P. lucublandus (Say) NT BC AB SK MB ON pe NB NS pE 

castanipes Kirby 

conue*icollis (Say) 

dilatatus LeConte 

elucens Casey 

fraternus Say 

manhattanis (Casey) 

planifer Casey 

P. scitulus LeConte - BC AB SK MB 

(Subgenus DERUS Motschulsky) 

(D er ulus Tschitsch6rine) 

P. nearcticus (Lindroth.t* NT 

Genus STEREOCERUS Kirby 

Boreobi a Tschitscherine 

S. haematopus (Dejean)* AK YK NT BC AB SK MB ON pe - LB NF 

similis I\trby 

S. rubripes (Motschulsky)* AK 

hry z hanou s kii (Lindroth) 

Genus MYAS Dejean 

Key to North American species: Allen (1980) 

M. coracinus (Say) 

- ON 

M. cyanescens Dejean - ON pe NB 

foueatus LeConte 

Genus PTEROSTICHUS Bonelli 

(Subgenus CYLINDROCHARIS Casey) 

Key to species: Barr (1971) 

P. rostratus (Newman) 

grandiceps (LeConte) 

piceatus (Casey) 

sulcatulus (Casey) 

(Subgenus HOLCIOPHORUS LeConte) 

P. lama (M6netri6st 

o/er (Dejean) 

- BC 

at e r r imu s (Motschulsky) 

cephalus (Casey) 

domitor (Casey) 

pollens (Casey) 

uancouueri (Casey) 

(Subgenus FIYPHERPES Chaudoir) 

(B r ac hy s tilu s Chaudoir) 

(Gonoderus Motschulsky) 

(H ap locoelus Chaudoir) 

- ON pe NB 

P. adoxus(Say) 

rejectus LeConte 

- ON pe NB NS 

sufflatus (Casey) 

P. a)gidus LeConLe 

anthrat (Casey) 

AK BC 

innatus (Casey) 

responsor (Casey) 

ualidus (Dejean) 

P. amethystinus Mannerheim 

jejunus Casey 

AK BC 

metlakatlae Casey 

nouellus Casey 

stoicus (Casey) 

P. castaneus (Dejean) 

brunneus (Dejean) 

AK BC 

terracensis (Casey) 

wrangelli Casey 

P. crenicollis LeConte AK BC 

pugetanus Casey 

rectilatus Casey 

31

Carabidae 

P. ecarinatur, Hatch 

P. herculane,rs Mannerheim 

scenicus Casey 

P. neobrunneus Lindroth 

P. nigrocoerrleus Van Dyke 

P. protractus LeConte 

fontinalis Casey 

inornah.ts Bland 

prouenss (Casey) 

utensis t.Casey) 

uiuax (Casey) 

P. tristis (De"iean) 

interfector (Newman) 

subarcuztus LeConte 

sustentu s LeConte 

tetriculc Casey 

zephyru; Casey 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

ON PQ NB 

NS 

(Subgenus L.DPTOFERONIA Casey) 

Key to speries: Hacker (1968) 

P. pumilus Casey 

longicol'.is (Casey) 

longicol'.is LeConte 

oregonui (Chaudoir) 

oregonis Cslki 

P. sphodrinus LeConte 

idahoanus (Casey) 

BC 

BC 

AB 

(Subgenus M ONOFERONIA Casey) 

P. diligendus (Chaudoir) 

apalachius Horn 

osculan:, Casey 

(Subgenus G A.STRELLARIUS Casey) 

P. honestus (Say) 

america ous (Castelnau) 

atronite * (Casey) 

deficien.t (Casey) 

fastiditu s (Dejean) 

scolopateus (Casey) 

(Subgenus S'IEROPUS Dejean) 

(Refonic: Casey) 

(.Sterode rus Motschulsky) 

P. circulosus Lindroth 

(Subgenus B')THRIOPTERUS Chaudoir) 

(Dysidit.s Chaudoir) 

(.Pararg;ttor Casey') 

P. adstrictus Eschscholtz+ 

angusticollis (Casey) 

commixius Chaudoir 

latebricola (Casey) 

latescans (Casey) 

laxicolli.; (Casey) 

luczotii .Dejean) 

maehlini LeConte 

oblong ius c ulus (Motschul sky) 

oblongotzotatus (Say) 

ob t u sa n,7 ulu s (Motschulsky) 

saxatilis (Casey) 

sericeus (Casey) 

seriepun ctatus Mannerheim 

sex,p unc :atus (Mannerheim) 

shastan.rs (Casey) 

P. lustrans L,eConte 

atrolucens (Casey) 

p uncticollis (Motschulsky) 

P. mutus (Say) 

carbonarius (Dejean) 

egens (Casey) 

AK YK 

ON PQ NB 


BC 

PQ 

BC MB ON PQ NB NS PE NF 

NS 

32

Carabidae 

rnlorosus (Dejean) 

picicornis (Kirby) 

politus (Harris) 

puluinatus Hausen 

stenopus Hausen 

P. oregonus LeConte 

colligatus (Walker) 

P. pensylvanicus LeConte 

NT 

BC 

BC 


NF 

(Subgenus MORPHNOSOMA Lutshnik) 

(Euferonia Casey) 

(Omaseidius Jeannel) 

P. coracinus (Newman) 

adjunctus LeConte 

erebeus Casey 

flebilis LeConte 

lacustris (Casey) 

ludibundus (Casey) 

moerens (Newrnan) 

(?) monedulus (Newman) 

roanicus (Casey) 

rufitarsis (Nicolay & Weiss) 

strigosulus (Casey) 

uenator (Casey) 

uashingtonensis (Nicolay & Weiss) 

P. Iachrymosus (Newman) 

P. melanarius (IlligerX 

P. novus Straneo 

P. relictus (Newman) 

protensus LeConte 

P. stygicus (Say) 

bisigillatus (Harris) 

(?) picipes (Newman) 

probus Casey 

quadrifer (Casey) 

rugicollis (Haldeman) 

subaequalis (Casey) 

umbonatus (Casey) 

uapidus Casey 

_BCAB 

oN PQ NB NS PE 

ON PQ NB 

SKMBONPQNBNSPE-NF 

-MBONPQ 

ON 

oN PQ 

LB 

NF 

(Subgenus ABACIDUS LeConte) 

(Peristethus LeConte) 

P. atratus (Newman) 

P. permundus (Say) 

planifer (Casey) 

ON 

ON 

(Subgenus MELANIUS Bonelli) 

(Ly p ero som us Motschulsky) 

(Lyperus Chaudoir) 

(M e tame I ani us Tschitsch6rine ) 

(Omaseus Dejean) 

P. castor Goulet & Bousquet 

P. caudicalis (Say) 

agrestis Bland 

breuibasis (Casey) 

P. corvinus (Dejean) 

aequalis (Casey) 

subpunctatus (Harris) 

tenebrosus (Chaudoir) 

P. ebeninus (Dejean) 

acutangulus (Chaudoir) 

NT BC 

NT BC 

AB 

SK MB 

AB SK MB ON 

ON PQ 

ON PQ 

ON 

PQ 

NB 

NF 

(Subgenus PSEUDOMASEUS Chaudoir) 

P. luctuosus (Dejean) 

abjectus LeConte 

confluens (Casey) 

hamatus (Harris) 

P. tenuis (Casey) 

testaceus (Casey) 

BC 

AB 

AB 

SK 

MB ON 

MB ON 

PQ NB NS PE 

PQ NB NS PE 

NF 

NF 

(Subgenus LAGARUS Chaudoir) 

(Pseudargutor Casey) 

(P se udol ag ar us Lutshnik) 

33

Carabidae 

P. Ieconteianus (Lutshnik) 

erythropus (Dejean) 

nitidus iKirby) 

BC SK MB ON PQ NB NS 

(Subgenus ARGUTOR Dejean) 

(Americ cmasezzs Csiki) 

(Micronnseus Casey) 

(.O ma se : rlu s Lutshnik) 

P. corruscultLs LeConte 

P. femoralis iKirby) 

aequicoi.Iis (Casey) 

desidiosus LeConte 

P. patruelis (Dejean) 

bicolor (Kitby) 

linearis (Mannerheim) 

P. strenuus (Panzer)f 

(Subgenus CRYOBIUS Chaudoir) 

Key to No th American species: 

P. arcticola (Chaudoir) 

I ab r ad.a rens is ( Chaudoir) 

P. auriga Ball 

P. barryorun Ball 

P. biocryus liall 

P. brevicornis (Kirby)* 

delicatus (Casey) 

e p ipl e ur alis (J. Sahlberg) 

fa 

s t idiosus (Mannerheim) 

P. bryanti (\ian Dyke) 

P. bryantoid:s Ball 

P. cacumenir; Ball 

P. caribou Brrll 

P. chipewyarr Ball 

P. empetricola (Dejean)* 

globicollis Csikt 

r ot undi :olli s Mannerheim) 

( 

r ufi coll,:. s (Mannerheim) 

P. gerstlensis Ball 

P. haftorni I.,indroth 

P. hudsonicr:.s LeConte 

P. kotzebuei Ball 

P. mandibulrryoides Ball 

P. nivalis {R F. Sahlbergt+ 

P. parasimil:s BaIl* 

P. pinguedineus (Eschscholtz)x 

alaskensis (Poppius) 

beringi (Casey) 

diplogrnus (Chaudoir) 

holmbergi (Poppius) 

incogni;lus (Poppius) 

mandibularis (Kirby) 

montanellus Poppius 

splendidus (J. Sahlberg) 

s ub nititlul u s (Poppius) 

s ub s inuos us (Chaudoir) 

washingtoni (Casey) 

P. planus (J. Sahlberg) 

blaisdeili Van Dyke 

P. riparius ( Dejean) 

breuiusculus (Casey) 

caseyi Csiki 

/ofuus (Mannerheim) 

fus coae,zeus ( Chaudoir) 

laeuilatus Notman 

patulus (Casey) 

P. similis Mannerheim* 

q uadr icollis ( Mannerheim) 

P. soperi BalI 

P. stantonersis BaIl 

AK 

Ball (1966) 

AK YK 

AK 

AK 

AK 

AK YK 

NT 

NT 

BC 

BC 

NT BC 

NT 

AB 

AB 

AB 

AK 

AI{ 

AK YK NT 

NT BC AB 

AKYK_BC 

AK 

AK 

AK YK 

AK 

AK YK 

AK 

AK YK 

AK YK 

AK 

AK YK 

AK 

AK 

NT BC 

NT BC 

I'ff - 

NT BC 

NT 

NT 

AB 

SK MB 

ON PQ 

oN PQ 


MB 

nfg ON 

_MB 

_MBON 

PQ 

pl:) 

- rg 

P6-LB 

LB 

LB 

NF 

NF 

NT 

34

Carabidae 

P. surgens LeConte AB 

P. tareumiut Ballx AK YK NT 

P. tiliaceoradix Ball AK 

P. ventricosus tEschscholtz)x AI{ YK NT 

aleutorum (Lutshnik) 

b reu icoll is (Tschitsch6rine) 

hy perboreus (Mannerheim) 

otariidinus (Casey) 

r ufi sca p us (Mannerheim) 

r ug ifer us ( Tschitsch6rine) 

s ubcaudatus ( Mannerheim) 

s ubexarat us (Mannerheim) 

u indic atus (Mannerheim) 

(Subgenus LYPEROPHERUS Motschulsky) 

Key to species: Budarin (1976) 

P. agonus Horn* AK YK NT 

P. costatus (Men6tri6s)* AK YK NT 

P. punctatissimus (Randall) - YK NT BC AB SK MB ON PQ NB NS - LB NF 

P. sublaevis (J. Sahlberg)* AI{ YK NT 

rufofemoralis Van Dyke 

P. vermiculosus (M6n6tri6s)x AK YK NT 

innuitorum (W.J. Brown) 

Genus CYCLOTRACHELUS Chaudoir 

Key to species: Freitag (1969) 

(Subgenus EVARTHRUS LeConte) 

C. sodalis sodalis (LeConte) 

corax (LeConte) 

- ON 

fatuus (LeConte) 

inflatipennis (Casey) 

I icinoide s ( Motschul sky) 

retractus (Casey) 

sulcatus (Casey) 

uagans (.LeConte') 

Genus ABAX Bonelli 

A. parallelepipedus NS NF 

(Piller & Mitterpacher)f 

aler (Villers) 

s/riolo (Fabricius) 

Genus DISAMARA Lindroth 

Pseud,amara Lindrotn 

D. arenaria (LeConte) 

Tribe Zabrini 

_ONPQNBNSPE 

Genus AMARA Bonelli 

(Subgenus CURTONOTUS Stephens) 

(Leirus Zimmermann) 

A. alpina (Paykull)* 

alaskana Cslki 

angustata J. Sahlberg 

arguta (Casey) 

borealis (Chaudoir) 

brunnipennis Dejean 

deficiens (Casey) 

inanis (Casey) 

obtusa LeConte 

rubripennis (Casey) 

subtilis (Casey) 

A. aulica (Panzer)t 

A. blanchardi Hay'ward 

A. bokori Csiki* 

sahlbergi Poppius 

A. carinata (LeOonte) 

AKYKNTBCAB-MBONPQ-LBNF 

___sc____PQIUNS__NF 

AK YK NT BC 

MB 

_BCABSKMBONPQNB 

35

Carabidae 

adstricta (Putzeys) 

catenulata (Casey) 

giluipes (Casey) 

laticollis (LeConte) 

A. hyperborea Dejean* 

canadensis (Putzeys) 

elongatr.r (LeConte) 

imperfe':ta (W.J. Brown) 

tristis (Prtzeys) 

A. lacustris l,eConte 

manitobensis (Casey) 

A. pennsylva-nica Hayward 

fuluipes (Putzeys) 

A. thoracica Hayward 

A. torrida (Panzer)* 

albertana (Casey) 

biarcuala (Casey) 

breuilat'ris (Kirby) 

breuipennis (Casey) 

consueta Fall 

cylindrica LeConte 

dura (Casey) 

holnrbergi (Putzeys) 

hudsonica Hayward 

labradarensis (Casey) 

melanollastrica Dejean 

reflexa rPrtzeys) 

rufimat a (Kirby) 

scrutat( (Casey) 

somnoh nta (Putzeys) 

(Subgenus B RADYTUS Stephens) 

(P seuda brady tzs Csiki) 

AK YK NT 

BC 


AK YK NT 

AB 

BC AB SK 

BC AB SK 

SK MB ON PQ 

MB ON 

-oN 

PQ 

NS 

NS 

LB 

MB ON PQ Ne NS pn LB 

NF 

NF 

A. apricaria Paykull)t 

putzeystana Csiki 

putzeysti Hotn 

A. avida (Sa d 

confinis Dejean 

A. browni Li rdroth 

A. colvillensi s Lindroth 

A. exarata Dejean 

curta ((:asey') 

furtiua;Jay 

grauida (Casey) 

stygiali:; (Casey) 

A. fulva (O.F. Mriller)i 

A. glacialis ( Mannerheim)* 

nainensis Casey 

A. Iatior (Kirby) 

decepta (Casey) 

humphreysi (Casey) 

laeuistri.ata (Putzeys) 

Iibera LeConte 

oregona LeConte 

relicta (Casey) 

A. schwarzi.{ayward 

septentrionalis (LeConte) 

-YK 

AK 

AK YK 

NT 

NT 

BC 

BC 

Nr sc 

_BC 

AB 

SK 

SK 

MB ON PQ 

MB ON PQ 

-ms-- 

ON 

SK 

SK MB 

_PQ 

oN PQ 

oN PQ 

YK NT AB MB ON PQ 

NB NS 

NB NS 

NB NS 

NB NS 

PE 

LB 

PE LB 

_LB 

NF 

NF 

NF 

NF 

(Subgenus PERCOSIA Zimmermann) 

Key to species: Hieke (1978) 

A. obesa (Say) 

dffinis (LeConte) 

extensa (Casey) 

latissima (Casey) 

sulcatuta (Casey) 

uentricosa (Casey) 

YK NT BC 

SK MB ON PQ NB NS 

PE 

NF 

(Subgenus CELIA Zimmermann) 

(.Isopleurus l{rtby) 

A. aurata De;jean 

euanida (Casey) 

36

Carabidae 

fragilis (Casey) 

gouanensis (Casey) 

irnitatrix Horn 

proba (Casey) 

A. bifrons (Gyllenhal)t 

A. brunnea (Gyllenhal)* 

am pl ico ll is (Mannerheim) 

A. californica Dejean 

mericana Dejean 

perspecta Casey 

pugetana (Casey) 

A. chalcea Dejean 

coruina (Casey) 

maneei (Casey) 

pinorunr (Casey) 

schotti (Casey) 

sodalis (Casey) 

A. discors Kirby 

gibba (LeConte) 

lugubris (Casey) 

merula (Casey) 

neuadica (Casey) 

obsolescens (Casey) 

paganica (Casey) 

paroudna (Casey) 

scolopax (Casey) 

spaldingi (Casey) 

sphaerops (Casey) 

wahelandi Hatcft 

A. hicksi Lindroth* 

A. idahoana (Casey) 

exlineae Minsk & Hatch 

A. musculis (Say) 

breuitarsis (Casey) 

contempta (LeConte) 

crenulata (Casey) 

curticeps (Casey) 

fluminea (Casey) 

limbalis (Casey) 

lyncea (Casey) 

minnesotana (Casey) 

nugator (Casey) 

paulula (Casey) 

scintilla (Casey) 

shoemaheri (Casey) 

uegrandis (Casey) 

A. pseudobrunnea Lindroth 

A. quenseli (Schbnherr)x 

brumalis (Casey) 

brunnescens (Casey) 

ceruicalis (Casey) 

defecta (Casey) 

definita (Casey) 

docilis (Casey) 

eldorensis (Casey) 

erplanatula (Casey) 

exposita (Casey) 

femoralis Hotn 

fontinalis (Casey) 

hornis Cslki 

i ndi s t inc ta (Mannerheim) 

laxicollis (Casey) 

mimica (Casey) 

obligata (Casey) 

piperi (.Casey) 

relucens (Mannerheim) 

remota (Zimmermann) 

remotestriata Dejean 

tahomae (Casey) 

washoeana (Casey) 

A. rubrica Haldeman 

AK 

ON 

AK YK NT BC AB SK MB ON PQ 

NT 

_BC 

_BC 

BC 

AB 

AB 

SK 

MB ON PQ 


AKl'K-BCABSKMB-PQ 

AK YK NT BC AB SK MB ON PQ NB NS pe 

ON PQ 

NS 

I\D 

LB 

LB 

LB 

NF 

NF 

37

Carabidae 

ferruginea Casey 

liquida (Casey) 

lubrica (Casey) 

lucina (Oasey') 

nigripennis (Casey) 

pallida (Casey) 

piceonitt:ns (Casey) 

politissitna (Casey) 

proditor (Casey) 

A. sinuosa (Casey) 

elusa (Casey) 

hospes (Casey) 

nitida (Yitby) 

nupta (Casey) 

subaenett (LeConte) 

subaenescens Csiki 

thoracicu (Casey) 

A. spuria Lindroth 

(Subgenus AIIARA s. str.) 

A. aenea (DeGeer)f 

deuincta Casey 

A. aeneopolit.r Casey 

A. coelebs Hayward 

leuiceps Jasey 

nebrask

Carabidae 

parallela (Casey) 

sho.stanica (Casey) 

solita (casey) 

subdepressa (Casey) 

subsimilis (Casey) 


A. impuncticollis (Say) 

(? ) a nt hracina Haldeman 

arcuata Casey 

dfficilis LeConte 

edor Notman 

otiosa Casey 

wadei Casey 

A. interstitialis Dejean* 

A. Iaevipennis Kirby 

aeneolucens Casey 

A. littoralis Mannerheim 

conuergens Casey 

fallaxLeConte 

hesperia Casey 

inepta LeConte 

heeni Casey 

lacustrina Casey 

laurana Casey 

microcephala Casey 

mystica Casey 

oodiformis Casey 

pullmani Casey 

recticollis Casey 

(?) teres Notman 

turbata Casey 

A. iunicollis Schiddtex 

carriana Casey 

marquettensis Casey 

A. patruelis Dejean* 

columbiana (Casey) 

inaequalis Kirby 

reducta (Casey) 

splendida Haldeman 

A. sanjuanensis Hatch 

(Subgenus ZEZEA Csiki) 

(Pseudotriaena Minsk & Hatch) 

(Triaena LeConte) 

A. angustata (Say) 

flebilis (Casey) 

indistincta Haldeman 

A. Iongula LeConte 

afoueolata Hayward 

alaxnoguia Minsk & Hatch 

atrichata Minsk & Hatch 

glabrata Minsk & Hatch 

profuga (Casey) 

A. pallipes Kirby 

depressa (LeConte) 

h,incaidi Minsk & Hatch 

lawrenciana (Casey) 


sherrnani (Casey) 

A. scitula Zimmermann 

irregularis (Casey) 

obliqua Casey 

prouana Casey 

uinta (Casey) 

uapida (.Casey) 

AK YK NT 

AK_NTBCABSK 


SK MB ON PQ NB NS PE NF 

ms oN 

MB ON 

AK YK BC AB SK MB ON PQ NB 

PQ 

PQ 

NB 

NB 

AK YK NT BC AB SK MB ON PQ NB 

BC 

BC 

BC 

BC 

AB 

AB 

SK MB ON PQ 


SK MB 

NS PE 

NS PE 

NS 

NS 

PE 

LB 

NF 

NF 

NF 

NF 

Tribe Harpalini 

Genus CRATACANTHUS Dejean 

C. dubius (Palisot de Beauvois) 

bisectus Casey 

AB 

SK 

39

Carabidae 

litoreus Jasey 

pensylucinicus Dejean 

Genus PIOS0MA LeConte 

P. setosum L-'Conte - BC AB SK 

breuipenne Casey 

Genus EURYDERUS LeConte 

Nothopus LeConte 

Key to species: Bail (1960) 

E. grossus (S,ryt 

arizonic,ts (Casey) 

- BC AB SK MB ON 

obtusus .Casey) 

priuatus (Casey) 

ualens ((.)asey) 

zabroides LeConte 

Genus HARF ALOBRACIIYS Tschitsch6rine 

H. leiroides t vlotschulskyrx AK - NT - A-B 

Genus HARPAI-US Latreille 

(Subgenus OPHONUS Dejean) 

H. puncticeps (Stephensti - ON PQ - NS 

H. rufibarbis t Fabricius rt PQ 

(Subgenus PSiEUDOPHONUS Motschulsky) 

Key to Norbh American species: BaIl and Anderson (1962) 

H. bicolor tFz bricius t 

admissu; Casey 

MB ON PQ - NS 

compar .,eConte 

excubant: Casey 

feroculut Casey 

nactus Casey 

H. ery.thropu r Dejean MB ON PQ - NS 

abstrusus Casey 

cupiens t)asey 

deludent Casey 

effetus Casey 

fenisex Casey 

rufopicet.ts Casey 

H. faunus Sa.r MB ON PQ 

badius Ltejean 

conuiuus LeConte 

H. longicollis LeConte 

dolosus ()asey 

- ON PQ 

haldemoni Casey 

latescanr Casey 

uagans L,eConte 

H. paralus Cusey AB SK 

spaldingi Casey 

H. pennsylva:ricus (DeGeer) 

inmixtus Casey 

- BC - SK MB ON PQ NB NS PE 

longior \:Jtby 

mormoni.cus Casey 

H. rufrpes (DeGeer)t PQ NB NS PE - NF 

(Subgenus M IGAPANGUS Casey) 

H. caliginosu; tFabricius) MB ON PQ 

dux Casey 

(Subgenus PI,ECTRAI-IDUS Casey) 

Key to species: El-Moursy (1958) 

H. erraticus Siay MB ON PQ 

caudalis Casey 

(Subgenus PIIARAJ-US Casey) 

H. indianus C siki - ON 

testaceus LeConte 

40

Carabidae 

(Subgenus IIARPAI-US s. str.) 

H. affinis (Schranktt 

aenescens Casey 

oenezs (Fabricius) 

assimilis Dejean 

canonicus Casey 

conuictor Casey 

lustralis Casey 

uiridiaeneus Palisot de Beauvois 

uiridis Say 

(Subgenus HARPALOMERUS Casey) 

H. amputatus Say 

ancillaris Casey 

bracatus Casey 

cuculus Casey 

cupreolatus Casey 

papagonalis Casey 

rotundicollis I\ttby 

stephensi l\ttby 

transuersus Casey 

(Subgenus EUHARPALOPS Casey) 

H. animosus Casey 

montanus LeConte 

montuosus Csiki 

H. egregius Casey 

alienus LeConte 

baergi Csiki 

cascadiensis Hatch 

instructus Casey 

rufimanus LeConte 

H. fraternus LeConte 

nimius Casey 

occidenlalis Chaudoir 

praestans Casey 

wadei (Casey) 

H. funerarius Csiki 

funestus LeConte 

reuersus Casey 

H. hoppingi Lindroth 

H. laticeps LeConte 

solutus Casey 

H. lewisii LeConte 

aesopus Casey 

H. viduus LeConte 

prouidens Casey 

-BCABONPQNBNSPELBNF 


_BCABSK 

AK YK NT BC AB SK MB ON PQ _ 

-BCAB 


_BC 

AK YK NT BC AB SK MB ON PQ NB NS - 

NT-ABSKMBONPQNBNS 

_ONPQ 

NS PE LB NF 

LB NF 

$ 

H. indigens Casey 

beatulus Casey 

- ON PQ NB 

piperi (Casey) 

H. ventralis LeConte AB SK MB 

electus Casey 

$ 

H. fulvilabris Mannerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

H. megacephalus LeConte MB ON 

(Subgenus CORDOHARPALUS Hatch) 

H. cordifer Notman AK BC 

w ashingtoniensls Van Dyke 

$ 

H. carbonatus LeConte NT BC AB SK 

H. fallax LeConte - ON PQ NB NS 

amiculus Casey 

recisus Casey 

H. herbivagus Say - BC AB SK MB ON PQ NB NS PE 

ossenszs Casey 

4L

Carabidae 

foueicollis LeConte 

mutabilis (Haldeman) 

H. intactus Casey 

uintanus Casey 

H. nigritarsis C.R. Sahlbergx 

curtatus Mannerhein 

proximus LeConte 

recensus Casey 

H. pleuriticus Kirby 

aequabilis Casey 

lasciuus tJasey 

liuidulus Casey 

niualis Casey 

perspicuus Casey 

placidus Casey 

pumilio Casey 

H. seclusus Casey 

parowanus Casey 

H. solitaris D,:jean 

fuliginos zs (Duftschmid) 

opicus Ctl.sey 

H. somnulentrs Dejean 

blanditu; Casey 

contactut'Casey 

pacificus FalI 

H. uteanus Cr,Lsey 

oppositut Casey 

A 

x 

H. alaskensis Lindroth 

H. innocuus LeConte 

fugitans Casey 

intactus Jasey 

persolus Casey 

_BC 

AK YK NT BC AB MB ON PQ 

AK 

AK 

BC 

AB 

-BCAB 

NT BC AB 

NT 

BC 

sc 

MB ON PQ NB NS PE LB 

SK MB ON PQ 

AB SK MB 

AB 

SK 

oN pe 

LB 

LB 

NF 

NF 

NF 

$ 

H. cautus Dej:an 

aduena I eConte 

(? ) alb ior, icus Mannerheim 

crenatell n Casey 

columbianus Casey 

(?) defix.us Walker 

oregonensis Casey 

H. ellipsis Le(lonte 

H. obnixus CzLsey 

H. ochropus FJrby 

H. opacipennis (Haldeman) 

lacustris Casey 

leuiceps Oasey 

oodioides Chaudoir 

H. plenalis Casey 

latebricoi,a Casey 

modulatus Casey 

BC 

---scTsK------- 

AK BC AB SK MB ON PQ 


NT BC 

MB ON PQ 

NB NS 

NF 

$ 

H. desertus LeConte 

captiosu:t Casey 

couleensts Hatch 

dulciculus Casey 

lucidus LeConte 

lustrans Casey 

metuens Casey 

uaciuus Oasey 

BC 

AB 

SK 

R 

Y 

H. fulgens Csiki 

nitidulus Chaudoir 

Genus IIARP ALELLUS Lindroth 

H. basilaris (Kirby) 

durescans (Casey) 

AK YK NT BC AB SK MB ON PQ NS 

ON 

42

Carabidae 

er.tensus (Walker) 

furuiculus (Casey) 

obesulus (LeConte) 

oblongus (Casey) 

renoicus (Casey) 

sejunctus (Casey) 

stocktonensis (Casey) 

subenormis (Casey) 

uaricornis (LeConte) 

uentricosus (Casey) 

Genus TRICHOTICHNUS Morawitz 

(Subgenus TRICHOTICHNUS s. str.) 

(Episcopellus Casey) 

(Pteropalus Casey) 

T. autumnalis (Say) 

nitescens (Casey) 

T. dichrous (Dejean) 

fluuialis (Casey) 

uersutulus (Casey) 

T. vulpeculus (Say) 

nigripennis (Dejean) 

Genus SELENOPHORUS Dejean 

(Subgenus SELENOPHORUS s. str.) 

(Gynandropus Dejean) 

(Hemisopalus Casey) 

S. gagatinus Dejean 

m(turus Haldeman 

uiridescens LeConte 

S. hylacis (Say) 

americanus (Dejean) 

S. opalinus (LeConte) 

iripennis LeConte 

S. pedicularius Dejean 

aereus LeConte 

puellus Putzeys 

troglodjttes Dejean 

S. planipennis LeConte 

( Subgenus CELIAMORPHUS Casey) 

S. ellipticus Dejean 

currens (Casey) 

opaculus (Casey) 

oualis Dejean 

ON 

ON 

BC AB SK MB ON 

ON PQ 

ON 

ON 

oN PQ 

ON 

ON 

PQ NB 

Genus DISCODERUS LeConte 

Selenalius Casey 

D. parallelus (Haldeman) 

amer icanus (Motschulsky) 

gener Casey 

hesperus Casey 

paruuliceps Casey 

BC 

ON 

Genus GEOPINUS LeConte 

G. incrassatus (Dejean) 

fluuiaticus Casey 

AB 

SK MB ON PQ 

Genus ANISODACTYLUS Dejean 

(Subgenus ANADAPTUS Casey) 

A. discoideus Dejean 

A. nivalis Horn 

paruulus (Casey) 

A. pitychrous LeConte 

A. sanctaecrucis (Fabricius) 

baltimoriensis (Say) 

uteanus (Casey) 

BC 

sK yu oN 

PQ 

_ABSKMB 

BC AB SK MB ON PQ 

NB 

lls 

PE 

43

Carabidae 

A. viridescenr LeConte 

chalceus LeConte 

idahoensis (Hatch) 

Iecontei Gemminger & Harold 

(Subgenus G\T{ANDROTARSUS LaFert6-S6nectbre) 

(Triplect:us LeConte) 

Key to Norl;h American species: Noonan (1973) 

A. dulcicollis LaFert6-S6nectdre) 

ellipticu: LeConte 

elongatu.; Chaudoir 

modicus (Casey) 

A. merula Ge'mar 

aethiops (Casey) 

crassus LeConte 

grauidus LeConte 

hetnpi (Casey) 

margina.lus (Casey) 

pinguis l.,eConte 

sulcipennis (Casey) 

uolcotti iCasey) 

A. ovularis {Casey} 

semirubilus (Casey) 

A. rusticus t S ry r 

oblongus (Casey) 

lrjslls Dr jean 

ON 

MB ON PQ 

-oN 

MB ON 

PQ 

NB 

NS 

PE 

(Subgenus AIIISODACTYLUS s. str.) 

(.Cephaloqyna Casey) 

A. binotatus ( Fabricius tt 

A. californicu r Dejean 

aleneanus Casey 


comes Ct.sey 


humeral,s Casey 

maestus Casey 

obsolescens Casey 

oregonus Casey 

paganict s Casey 

sinuatus Casey 

A. carbonariu s t Say t 

breuior (t laseY) 

luctuosu,:; Dejean 

rufipenn,'.s LeConte 

A. furvus LeC onte 

A. harrisii Le'lonte 

Iacertosus Casey 

A. kirbyi Lincroth 

(?) Iecontei Chaudoir 

A. melanopus (Haldeman) 

A. nigerrimus {Dejean) 

Iaticollis (Kirby) 

opacus (()asey) 

punctulatus LeOonte 

A. nigrita Dejean 

inte r p un.:t at u s (Kirby) 

A. similis Le(lonte 

incertus Jasey 

incisus C. asey 

punclico,lis Chaudoir 

semipunctdtus LeConte 

sericatus Casey 

solidus Casey 

_BC 

_BC 

BC 

BC 

BC 

BC 

AB 

AB 

SK 

SK 

-oN 

MB ON 

MB ON 

oN PQ 

PQ 

PQ 

ON 

oN PQ 

MB ON PQ 

NS- 

NB NS 

NN NS 

NB NS 

PE 

NF 

(Subgenus AI'LOCENTRUS LeConte) 

A. amaroides LeConte 

BC 

(Subgenus PS,EUDAPLOCENTRUS Noonan) 

A. laetus Dejr an 

gemmeus (Casey) 

ON 

44

Carabidae 

(Subgenus SPONGOPUS LeConte) 

A. verticalis (LeConte) 

Genus AMPHASIA Newman 

(Subgenus AMPI{ASIA s. str.) 

A. interstitialis { Say) 

fuluicollis Newman 

mollis Casey 

obscuripennis (Dejean) 

(Subgenus PSEUDAMPHASIA Casey) 

A. sericea (Harris) 

femorata (Dejean) 

Genus DICHEIRUS Mannerheim 

Key to species: Noonan (1968) 

D. piceus (Menetriest 

alutaceus Casey 


australinus Casey 

decoloratus Casey 

facilis Casey 

incidens Casey 

iteg ul ar is Motschulsky 

insularis Casey 

parallelus LeConte 

rupimontis Casey 

sodalis Casey 

uillosus Motschulsky 

BC 

SK 

SK 

MB ON PQ 

ON PQ 

oN PQ 

Genus XESTONOTUS LeConte 

X. lugubris tDejeant 

manhattanis (Casey) 

tioganus (Casey) 

Genus NOTIOBIA Perty 

(Subgenus ANISOTARSUS Chaudoir) 

(Diaphoromerus Chaudoir) 

(Eurytrichus LeConte) 

(Stilbolidus Casey) 

Key to North American species: Noonan (1973) 

N. nitidipenni s (LeConte) 

agitabilis (Casey) 

cephala (Casey) 

conspectd (Casey) 

delicata (Casey) 

tenuitarsis (Casey) 

N. picea tLeConte) 

soyi (Blatchley) 

N. terminata tSayl 

agilis (Dejean) 

angusticollis (Casey) 

conniuens (Casey) 

florid,ana (Casey) 

foueicollis (Bates) 

fuscipennis (Casey) 

hebes (Casey) 

inerrans (Casey) 

ocreata (Say) 


similis (Say) 

suboualis (Casey) 

subuirens (Casey) 

testacea (Haldeman) 

uernicata (Casey) 

oN PQ 

oN PQ 

oN PQ 

ON PQ 

NS 

NB NS 

45

Carabidae 

Genus TRIC JOCELLUS Ganglbauer 

(Subgenus T RICHOCELLUS s. str.) 

T. cognatus (Gyllenhal)x AK YX NT BC AB SK MB ON PQ NB NS PE LB NF 

axillar i s (Mannerheim) 

boreellus Casey 

conflag r at us ( Mannerheim) 

lateralis Casey 

longius

Carabidae 

B. conformis (Fall) 

tetricus (Casey) 

B. georgei Lindroth 

B. kirbyi (Horn) 

B. lugubris (LeConte) 

frosti (Fall) 

protractus (Casey) 

BC 

AB 

AB 

oN PQ 

_MBONPQ 

NB 

NS 

PE 

NF 

Genus STENOLOPHUS Stephens 

(Subgenus STENOLOPHUS s. str.) 

S. anceps LeConte 

fidelis Casey 

rolu nd i colli s Motschul sky 

S. carbonarius (Dejean) 

S. fuliginosus Dejean 

fuscipennis (LeConte) 

uersicolor Kirby 

S. fuscatus Dejean 

S. humidus Harnilton 

S. incultus Casey 

consors Casey 

debiliceps Casey 

S. limbalis LeConte 

indistinctus Motschulsky 

longitarsis Casey 

S. megacephalus Lindroth 

S. ochropezus (Say) 

conuexicollis LeConte 

floridanus Casey 

(?) gracilis Casey 

testaceicollis Casey 

S. plebejus Dejean 

(?) lugubris (Haldeman) 

riuularis Casey 

(Subgenus AGONODERUS Dej ean) 

S. comma (Fabricius) 

dorsalis (LeConte) 

gracilitarsis Casey 

obliqulus Casey 

similis (I\trby) 

S. lecontei (Chaudoir) 

plagiatus Casey 

S. lineola (Fabricius) 

furcatus (Fabricius) 

s 

S. conjunctus (Say) 

captiosus Casey 

humilis (Dejean) 

immunis (Kirby) 

misellus (Dejean) 

moquinus Casey 


S. rotundatus LeConte 

S. rotundicollis (Haldeman) 

incitatus Casey 

scitulus Casey 

Genus ACUPAJ-PUS Latreille 

(Subgenus PHILODES LeConte) 

(Goniolophus Casey) 

A. alternans (LeConte) 

testaceus (LeConte) 

A. rectangulus Chaudoir 

(Subgenus ACUPALPUS s. str.) 

A. canadensis Casey 

A. carus (LeConte) 

BC 

BC 

BC 

BC 

AB 

ON 

SK MB ON 

ON 

ON 

ON 

NS 

PQ NU NS PE N" 

PQ_ fr_ _ _ 

PQ 

NB NS 

BC SK MB ON PQ NB NS PE 

BC 

AB 

oN PQ 

_MBONPQ 

ON 

SK MB ON 


-oN 

-oN 

PQ 

PQ 

oN PQ 

oN PQ 

pe- 

_ __ NC_:"MB3Ni3NBN3Ff;_ Nil 

PE 

NF 

47

Carabidae 

curlipet nis Casey 

expertus Casey 

subrectu,.s Casey 

triuialis Casey 

A. meridiant s ll-inn6t+ - BC AB - PQ 

A. nanellus (lasey - ON PQ NB NS 

A. pumilus I indroth - ON PQ - NS 

(Subgenus T A.CHISTODES Casey) 

A. indistinct rs Dejean - ON 

obscurus (Casey) 

A. partiariu: tSayt ON PQ 

conuerg tns (Casey) 

Iyratus Casey') 

A. pauperculus Dejean ON PQ NB NS PE 

consimi'.is Dejean 

A. testaceus Dejean - ON PQ 

fusciceps (Casey) 

micros LeConte 

Genus DIPL ]CHEILA Brull6 

Rembus Dejean 

(Subgenus If IOREMBUS Jeannel) 

Tlibe Licinini 

Key to North American species: Ball (1959) 

D. assimilis rl-eConter ON PQ 

modesta Casey 

D. impressic',llis rDejeanr 

cliens C*ey 

ON PQ 

foueata Jasey 

laticollis (LeConte) 

planulala Casey 

D. major ma. or (Leconte ) - ON 

expansa Casey 

oblonga Casey 

procera Casey 

D. obtusa (LeConte) NT BC AB SK - ON PQ NB NS PE 

parallelt Casey 

D. oregona tllatcht BC AB SK MB 

D. striatopurLctata (LeConte) NT BC AB SK MB ON PQ 

alternar.s Casey 

amplipennis Casey 

breuicoli,is Casey 

D. undulata larr AB - MB 

Genus DICAELUS Bonelh 

(Subgenus P.ARADICA-ELUS Ball) 

D. dilatatus lilatatus Say PQ 

carolinensis Casey 

dejeanii Dejean & Boisduval 

planicollis LeConte 

D. elongatus Bonelli 

osieuill rnsis Casey 

- ON PQ 

debilice;ts Casey 

obscuru.; LeConte 

simplex Dejean 

D. politus Dt jean - ON PQ 


leonardli Hatis 

D. sculptilis ntricatus LeConte - ON 

ocellatus Blatchley 

D. sculptilis rpioides Ball - SK MB ON 

D. teter Bonelli ON PQ 

ouipenn'.s Casey 

(Subgenus DICAELUS s. str.) 

D. purpuratus purpuratus Bonelli * ON 

uiolaceus Bonelli 

48

Carabidae 

(Subgenus LIODICAELUS Casey) 

D. laevipennis laevipennis LeOonte AB 

eudnescens (Casey) 

Genus BADISTER Clairville 

(Subgenus BADISTER s. str.) 

B. lernrgineus Dejean - BC 

anthracinus LeConte 

B. neopulchellus Lindroth NT BC AB SK MB ON PQ - NS PE 

B. notatus Haldeman - ON PQ 


giluip.es Casey 

terminalis LeConte 

B. obtusus LeConte NT - AB SK MB ON PQ - NS 

B. pulchellus LeConte - ON 

(Subgenus BAUDIA Ragusa) 

B. grandiceps Casey - BC AB - MB ON PQ - NS 

B. transversus Casey MB ON PQ 

B. ocularis Casey - ON PQ NB NS PE 

B. parviceps Ball - BC MB ON PQ 

B. reflexus LeConte - ON 

seclusus Blatchley 

Genus OODES Bonelli 

Tribe Oodini 

O. amaroides Dejean - ON 

O. americanus Dejean - ON PQ 

fluuialis LeConte 

O. brevis Lindroth - ON 

Genus LACHNOCREPIS LeConte 

L. parallela tSayt MB ON PQ 

Genus STENOCREPIS Chaudoir 

(Subgenus STENOUS Chaudoir) 

S. cuprea tChaudoirt - ON 

leucodacty Ia (LaFert6-S6nectdre) 

Tribe Panagaeini 

Genus PANAGAEUS Latreille 

P. fasciatus Say 

-oN 

Genus CHLAXNIUS Bonelli 

Key to North American species: Bell (1960) 

(Subgenus EURYDACTYLUS LaFert6-S6nectdre) 

(Gly ptoder us LaFert6-S6nectdre) 

Tlibe Callistini 

C. tomentosus tomentosus (Say) AB SK MB ON PQ 

amplus LeConte 

lacustrinus Casey 

luctuosus Germar 

(Subgenus ANOMOGLOSSUS Chaudoir) 

C. emarginatus Say 

delectans (Casey) 

- ON PQ NB NS 

grauis (Casey) 

semotus (Casey) 

49

Carabidae 

C. pusillus Say 

elegant,ilus Dejean 

- ON 

nanulu,; (Casey) 

(Subgenus CHLAENIUS s. str.) 

C. aestivus liay 

cobaltir'.us Dejean 

- ON 

congener LeConte 

C. cordicollir, Kirby MB ON PQ NB 

C. erlrthropr.. s Germar - ON 

oblongi;cennis Casey 

rufilabris Dejean 

umbritursis Casey 

C. laticollis :iay ON 

brachyclerus Chaudoir 

breuicol.lis LeConte 

rufipes Dejean 

C. sericeus sericeus (Forster) 

peruirict.is LeConte 

- BC AB SK MB ON PQ NB NS PE - NF 

uteanus Casey 

C. solitarius Say AB 

chlorophanus Dejean 

(Subgenus C HLAXNIELLUS Reitter) 

C. brevilabr-s LeCon[e 

(.?) cons tnilis LeConte 

- ON 

texaneltus Casey 

C. impunctilrons Say 

C. nebraske.'rsis LeConte AB SK 

MB ON PQ 

sedulus Casey 

C. nemoralir Say - ON 

Iongico,lis Chaudoir 

C. pensylvarLicus pensylvanicus Say BC AB SK MB ON PQ NB NS PE - 

(?) fulgtceps Newman 

impunctifrons l\ttby 

pubescc'ns l{arlis 

NF 

uicinus Dejean 

C. tricolor Dejean 

atripen,'tis LeConte 

- BC AB SK MB ON PQ NB NS NF 

chaly be ipennis Chevrolat 

jacinto Casey 

quadricollis Kirby 

uigilant Say 

(Subgenus F,RACFM,OBUS Chaudoir) 

C. Iithophilus lithophilus Say 

sma ra g' dig e r Motschulsky 

NT BC AB SK MB ON PQ NB NS PE - NF 

uiridanus Dejean 

(Subgenus A GOSTENUS Motschulsky) 

C. alternatus Horn AK - NT BC AB SK MB ON PQ NB NS NF 

alberta,tus Casey 

C. harpalinr s Eschscholtz - BC AB 

crestonensis W.J. Browrr 

C. interrupt rs Horn - BC 

C. niger Rardall NT BC AB SK MB ON PQ NB NS PE - NF 

ludouici.anus Leng 

$ 

C. purpuric< llis Randall NT BC AB SK MB ON PQ 

frostii Carr 

punctipennis Casey 

Genus PENIIAGONICA Schmidt-Goebel 

Didetut LeConte 

Rhomb tdera Reiche 

P. picticornis Bates 

Tribe Pentagonicini 

-ONPQ 

50

Carabidae 

Tribe Perigonini 

Genus PERIGONA Castelnau 

Trechicus LeConte 

P. nigriceps tDejeant - ON PQ 

umbripennis (LeConte) 

P. pallipennis (LeConte) - ON 

Genus CALATHUS Bonelli 

Key to North American species: Ball and Ndgre (1972) 

(Subgenus CALATHUS s. str.) 

(Fuscocalathus Ndgre) 

Tribe Platynini 

C. fuscipes (Goeze)* - BC 

(Subgenus NEOCAJ-ATHUS BalI & Nbgre) 

C. gregarius (Sayt - ON PQ NB NS 

distinguendus LeConte 

C. ingratus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE - NF 

acomanus Casey 

aquilus Casey 

beringi Casey 

calator Casey 



incommodus Mannerheim 

labrad,orinus Casey 

nanulus Casey 

planifer Casey 

reductus Casey 

C. opaculus LeConte 


- ON PQ 

appalachius Casey 

obesulus Casey 

sonoricus Casey 

uentricosus Casey 

C. ruficollis grandicollis Casey - BC 

(Subgenus PROCALATHUS Jedlicka) 

C. advena(LeConte) 

binarius (Casey) 

AK BC AB SK - ON PQ - LB NF 

b reu ius cul u s (Mannerheim) 

b runne s cen s (Mannerheim) 

dzlcis (Mannerheim) 

lenis (Mannerheim) 

mollis (Eschscholtz) 

scolopax (Casey) 

Genus LAEMOSTENUS Bonelli 

Key to species: Casale (1988) 

(Subgenus LAEMOSTENUS s. str.) 

L. complanatus (Dejean )* - BC 

(Subgenus PRISTOI.IYCHUS Dej ean) 

L. terricola (Herbst)f - BC PQ NB NS PE - NF 

Genus SYNUCHUS Gyllenhal 

Pristodactyla Dejean 

Key to species: Lindroth (1956) 

S. irnpunctatus (Say) 

americanus (Dejean) 

- BC AB SK MB ON PQ NB NS PE - NF 

americanus (LeConte) 

conuexu.s (Casey) 

coruinus (LeConte) 

Genus OLISTHOPUS Dejean 

Odontonyx Stephens 

51

Carabidae 

O. micans LcConte 

O. parmatus (Say) 

cinctus )ay 

iterans t)asey 

pictus C'asey 

Genus SERI{IODA Kirby 

Agonodromius Reitter 

S. bembidioides Kirby 

cicatricosa Motschulsky 

colonica Casey 

montico,!,a Casey 

S. bogemannii (Gyllenhal)* 

inuidiosa Casey 

s t r ig icolle (Mannerheim) 

tacomae Casey 

S. obsoleta ([iay) 

consimi.e (LeConte) 

insulina Casey 

luctuosc (Dejean) 

uicina (()emminger & Harold) 

S. quadripun ctata (DeGeer)* 

octocola (Mannerheim) 

stigmost.t, (LeConte) 

Genus ANCIIOMENUS Bonelli 

Tetraleucus Casey 

A. picticornis, Newman 

Genus PSEL DANCHUS Casey 

P. aeneolus (LeConte) 

bjorkmcnae Gray & Hatch 

P. quadratus (LeConte) 

dilleyanus (Casey) 

metuen: (Casey) 

Genus ATRANUS LeConte 

A. pubescens (Dejean) 

obconia.rs (Haldeman) 

pallesce.zs Casey 

Genus RHADINE LeConte 

Comstockia Van Dyke 

R. Iindrothi llan 

Genus PARANCHUS Lindroth 

P. albipes (F rbricius)'l 

clemens (LeConte) 

Genus OXYI'SELAPHUS Chaudoir 

Anchus LeConte 

O. puncticeps (Casey) 

pusillus (LeConte) 


AK YK _ 

BC .AB 

-oNPQ _ONPQ 

NT BC AB SK MB ON PQ NB NS _ 

AK YK NT BC AB SK MB ON PQ NB NS _ 

-BCAB 

-BC 

AB SK 

-oN 

_ONPQNB 

-BC_SKMBONPQ_NSPE 

_NBNSNF 

LB NF 

LB NF 

Genus AGOI{UM Boneili 

(Subgenus E UROPHILUS Chaudoir) 

A. anchomenoides Randall 

lanellum Gray 

A. canadenser Goulet 

A. consimile (GyllenhaI)x 

fragile (Mannerheim) 

inualidum Casey 

A. darlingtorri Lindroth 

A. exaratum (Mannerheim)* 

A. galvestonicum (Casey) 

AK _ NT BC AB SK MB ON PQ NB NS PE - NF 


AK YK NT BC AB _ MB ON PQ - NS - LB NF 

AK5*rec _ _ r,rn:x:a]"]'_ 

_ _ 

52

Carabidae 

A. gratiosum (Mannerheim) 

antiquum (Notman) 

carri (.Casey) 

properans (Casey) 

ruficorne (LeConte) 

symmetricum (Casey) 

A. lutulentum (LeConte) 

adustum (Casey) 

atriceps (Casey) 

elegantulum (Casey) 

A. palustre Goulet 

A. picicornoides Lindroth 

A. retractum LeConte 

collusor (Casey) 

dil uti p e nne (Motschulsky) 

elongatulum Haldeman 

facile (Casey) 

serenum (Casey) 

A. simile Kirby 

A. sordens Kirby 

ineptum (Casey) 

frosti (Casey) 

picicorne (LeConte) 

A. superioris Lindroth 

A. thoreyi Dejeanx 

gemellum (LeConte) 

(?) lenum Dejean 

picipenne Kirby 

AK YK NT BC AB SK MB ON PQ NB NS PE 

NT BC AB SK MB ON PQ NB NS PE 

- - - scABsxrunBNiSNsNspr- 


AKYK_BCABSKMB 


LB 

NF 

oN pe Ns Ns pr LB NF 

AK-NT-AB-MBONPQNB 

AK YK NT BC AB SK MB ON PQ NB NS PE _ 

NF 

(Subgenus PLAT\' IOMICRUS Casey) 

A. bicoior (Dejean)* 

ma r g inell un'L (LeConte) 

A. fer"ruginosum (Dejean) 

erasutn (LeConte) 

marcidum (Casey) 

uadei (.Casey) 

A. nigriceps LeConte* 

fragilissimum (Casey) 

A. piceolum (LeOonte) 

d.issensum (Casey) 

fractum (Casey) 

frigidulum (Casey) 

lasciuum (Casey) 

tritum (Casey) 

AK YK 

AK 

AK 

NT BC 

_BC 

NT BC 

_BC 

AB 

AB SK 

AB SK 

AB SK 

MB ON 

MB ON 

PQ NB 

PQ 

PE 

NF 

NF 

(Subgenus AGONUM s. str.) 

(Melanagonum Casey) 

(Paragonum Casey) 

(Punctagonum Casey) 

(Taphranchus Casey) 

A. affrne Kirby 

cor6o (LeConte) 

A. albicrus Dejean 

A. belleri (Hatch) 

A. brevicolle Dejean 

frater (LeConte) 

A. corvus (LeConte) 


hyslopi Casey 

A. cupreum Dejean 

borealinum Casey 

chalceum LeConte 

crassicolle (LeConte) 

cupreolucens Casey 

esuriale Casey 

Iongulum Casey 

marquettense Casey 

oualicauda Casey 

parallelum Casey 

protractum (LeConte) 

seminitidum Kirby 

AK YK NT BC AB SK PQ NB NS PE NF 

NT 

BC 

BC 

BC 

BC 

AB 

AB 

SK 

SK 

MB ON 

-oN 

MB ON 

MB ON 

PQ 

PQ 

NB 

53

Carabidae 

A. cupripenne (Say) NT BC AB SK MB ON PQ NB NS PE - NF 

gemmeum Casey 

nitidulunt Dejean 

A. deceptivurn (LeContet NS 

A. excavatum Dejean - ON PQ NB 

ontarioni; (Casey) 

trinariunt (Casey) 

A. fidele Case, - ON PQ NB NS PE 

laeue (Let)onte) 

molestwn (LeConte) 

subinflat,tm Casey 

A. harrisii LeConte 

aethiops l)asey 

- BC AB SK MB ON PQ NB NS PE 

mordax Casey 

A. melanariun Dejean 

maurum Haldeman 

- BC AB SK MB ON PQ NB NS PE 

militare l)asey 

(?) scuteliiare (.Say) 

A. metallescer:rs (LeConte) NT BC AB SK MB ON PQ NB NS NF 

lacustre Oasey 

terracens: Casey 

A. moerens Dr jean MB ON PQ 

A. muelleri (Eerbst)f - BC - ON PQ NB NS PE - NF 

hardyi (LeConte) 

A. mutatum ((lemminger & Harold) AK BC - SK MB ON PQ NB NS PE LB NF 

dtratum t LeConte) 

A. octopunctat.um (Fabricius) - ON PQ NB NS PE 

A. placidum (fiay) - BC AB SK MB ON PQ NB NS PE LB NF 

amplior (.)asey 

aztecanu;n Casey 

citatum (iasey 

morosum Dejean 

rhodeant.m Casey 

A. propinquurr (Gemminger & Harold) AK YK NT BC AB SK MB ON PQ NB NS PE - NF 

amens Ci:tsey 

fratercuhLm (LeConte) 

humile Crcey 

insuetum Casey 

piceum LeConte 

A. quinquepu:rctatum Motschuisky* AK YK NT - AB SK MB - PQ - LB 

perforatum (LeConte) 

A. subsericeurn (Leconte) - BC AB SK 

latiusculum Casey 

suffusum Casey 

uteanum Casey 

uirid.issintum Casey 

A. tenue (Lec')nte) 

A. trigeminunr Lindroth 

- ON PQ NB NS PE 

MB ON PQ NB NS PE 

(Subgenus MICRAGONUM Casey) 

(Circinal;:dia Casey) 

A. aeruginosum Dejean - ON PQ NB NS 

undulatum (Casey) 

A. nutans (Sayr - ON PQ 

femoratun Dejean 

A. striatopunr.tatum Dejean - ON 

breuiceps (Casey) 

decipiens LeConte 

luculentum (Casey) 

A. variolatum (LeConte) 

coronadiizum (Casey) 

- BC 

(?) limba:um Motschulsky 

shastanicum (Casey) 

(Subgenus CIRCINALIA Casey) 

A. crenistriatum (LeConte) 

liticola (Casey) 

- oN PQ - NS PE 

politissimum (Casey) 

roticolle (Casey) 

statenense (Casey) 

A. punctiforme t Sayt 2 - 

54

Carabidae 

foueicolle Chaudoir 

ludouicianum (Casey) 

A. rufrpes Dejean 

rubripes (Zimmermann) 

ON 

(Subgenus STEREAGONUM Casey) 

A. errans (Say) 

erythropus Kirby 

subcordatum (LeConte) 

A. ferreum Haldeman 

ocreatum Haldeman 

quadrulum (Casey) 

solid.ulum (Casey) 

BC 

SK 

MB ON PQ 

ON 

(Subgenus STICTANCHUS Casey) 

Key to species: Liebherr (1986) 

A. decorum (Say) 

arenarium (Casey) 

californicum (Dejean) 

charmis (Bates) 

ftornii (Hausen) 

itnpictum (Casey) 

irruptum (Casey) 

luxatum (Casey) 

obscurum (LeConte) 

sitnplex (LeConte) 

solutum (Casey) 

syrdcusense (Hatch) 

tepidum (Casey) 

t e stac eonot um (Hausen) 

thoracicum (Dejean) 

uteanum (Casey) 

uinnulum (Casey) 

A. extensicolle (Say) 

clientulum (Casey) 

(? ) cy ane sce ns (Motschulsky) 

gaudens (Casey) 

lecontei LeConte 

ob s c urat utn (Chaudoir) 

proximum (Harris) 

uigilans (Casey) 

uiride (LeConte) 

BC 

SK MB ON PQ 


NS 

(Subgenus DERATANCHUS Casey) 

A. quadrimaculatum (Horn) 

AB 

Genus PLATYNUS Bonelli 

(Subgenus PLATYNUS s. str.) 

P. brunneomarginatus (Mannerheim) 

bicolor LeConte 

bicoloratus Gemminger & Harold 

cinctellus LeConte 

ma r ginat u s (M6n6tries) 

rugiceps (Mannerheim) 

tenebricosus Gemminger & Harold 

P. cincticollis (Say) 

(?) blandus Germar 

coruinus (Dejean) 

deplanatus (Chaudoir) 

(?) maculifrons (Say) 

marginalis (Haldeman) 

P. decentis (Say) 

albertanus (Casey) 

aleneanus (Casey) 

coracinus (LeConte) 

d.epressus (Haldeman) 

gagates (Dejean) 

iowanus (Casey) 

missourianus (Casey) 

oblongipennis (Casey) 

BC 

oN PQ NS 

AK NT BC AB SK MB ON PQ NB NS 

LB 

NF 

DD

Carabidae 

pacatus ( Casey) 

sinuatus (Dejean) 

turbidus (Casey) 

P. mannerhei:nii (Dejean)* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

inter s t itiali s (Motschulsky) 

octofoueo atus (Maklin) 

t 

stygicus ,-eConte 

P. opaculus L':Conte 

6oopls (Casey) 

- ON PQ 

inquisitor (Casey) 

P. tenuicollis LeConte) 

distinguendus (Casey) 

- ON PQ NB NS 

margina;:us (LeOonte) 

pennsylu znicus (Casey) 

reflexus -l,eConte 

(Subgenus PI ATYNIDIUS Casey) 

P. hypolithos Sayr 

erythrop'.ts Dejean 

- ON 

ontarien::is (Casey) 

P. ovipennis (Mannerheim) 

arachnoides (Casey) 

- BC 

maurus (Motschulsky) 

rotundip znnis (Motschulsky) 

similatut: (Casey) 

Genus LEPT( )TRACHELUS Latreille 

Sphaera':ra Say 

Tribe Ctenodactylini 

L. dorsalis (F.rbriciusj - ON 

Genus COLL. URIS DeGeer 

(Subgenus C()SNANIA Dejean) 

(.O dacan,, hella Liebke) 

Tribe Odacanthini 

C. pensylvanica (Linn6r ON PQ 

Genus TETR,,\GONODERUS Dejean 

Tlibe Masoreini 

T. lasciatus (llaldemanj ON 

undulatt'.s LeConte 

Genus LEBIA. Latreille 

Key to Nor:,h American species: Madge (1967) 

(Subgenus LOXOPEZA Chaudoir) 

Tribe Lebiini 

L. atriceps LeConte AB SK MB 

nanulinc: (Casey) 

L. atriventris Say - SK MB ON PQ 

enormis .Casey) 

L. grandis Hrntz - ON PQ 

magister (Casey) 

majuscui.a (Chaudoir) 

riuularis (Casey) 

L. tricolor Sa., - ON PQ 

(Subgenus LIMPRIAS Bonelli) 

L. divisa LeC rnte AB SK MB 

concinna LeConte 

(Subgenus LI)BIA s. str.) 

(Ap helog enia Chaudoir) 

56

Carabidae 

(D ianchomena Chaudoir) 

(Metabola Chaudoir) 

L. analis Dejean 

anchora Chevrolat 

ap p endic ulata Chaudoir 

bonellii Putzeys 

(? ) s ubfr.g ur ata Motschulsky 

L. cyanipennis Dejean 

barbarae Casey 

montanaHorn 

melaenaHatch 

ruficollis LeConte 

L. fuscata Dejean 

canonica Casey 

L. guttula LeConte 

metuens Casey 

pacifica Casey 

L. lobulata LeConte 

(? ) b r unnicol lis Motschulsky 

L. moesta LeConte 

cynica Casey 

L. ornata Say 

(?) apicalis (Haldeman) 

asheuillensis Casey 

axillaris Dejean 

brunnea Haldeman 

erythrocephala Dejean 

fl au iu entris Motschulsky 

fluuiatilis Casey 

frigida Chaudoir 

marginella Dejean 

nigripennis Dejean 

reperta Casey 

uirginica Casey 

L. perita Casey 

L. pleuritica LeConte 

L. pulchella Dejean 

rufopyga (Chaudoir) 

tahoensis Casey 

uiuida (Bates) 

L. pumila Dejean 

frugalis Casey 

illini Casey 

ludouiciana Casey 

maculicornis LeConte 

quadrata Casey 

rhodopa Schwarz 

tertiaria Casey 

L. solea Hentz 

(.? 

) fl au olineata Motschulsky 

scapularis Dejean 

websteri Casey 

L. viridis Say 

adolescens Casey 

bracata Casey 

castigata Casey 

cobaltina Casey 

(?) cyanella (Motschulsky) 

duluthiana Casey 

euoluta Casey 

histrica Casey 

incitata Casey 

magica Casey 

papago Casey 

planifera Casey 


smaragd.ula Dejean 

subaffinis Casey 

subopaca Schaeffer 

trajecta Casey 

_BC 

BC 

BC 

BC 

BC 

YK NT BC 

AB 

SK 

SK 

ON 

MB ON PQ 

-oN 

MB ON 

-oN 

_SK 

AB SK MB 

AB 

ON 

ON 

PQ 

PQ 

NB 

NS 

NB NS 

BC SK MB ON PQ NB NS 

SK 

SK 

MB ON PQ 

MB ON PQ 

NB 

NS 

NS 

PE 

LB 

NF 

57

Carabidae 

truckeens s Casey 

uermiculi za Casey 

uiridis D< jean 

L. vittata (Fabricius) 

amnicola Casey 

- BC AB SK MB ON PQ - NS 

conjunget:s LeConte 

connecta tlChaudoir) 


depicta Horn 

flauouittata Chevrolat 

furcata LeConte 

sonomae l)asey 

spraguei (}{orlr) 

tempeana Casey 

Genus DROM. US Bonelli 

(Subgenus DRIMIUS s. str.) 

(Dinodrortius Casey) 

D. piceus Dejean - BC AB - MB ON PQ NB NS 

quadricollis LeConte 

Genus APRIS''IUS Chaudorr 

A. constrictus Jasey BC AB 

A. latens (LeCrntet - SK MB ON PQ - NS 

(.?) 

fuscipt nnis Motschulsky 

A. pugetanus l)asey - BC 

A. iubsulcatus (Dejean) - ON PQ NB NS 

cordicolli, ; (LeConte) 

Genus MICRC LESTES Schmidt-Goebel 

Blechrus Motschulskv 

Bomius L eConte 

M. brevilobus Lindroth - ON 

M. curtipennis (Casey) AB SK 

M. linearis (Lt,Conte) - BC AB SK MB ON PQ - NS 

anEustus (LeConte) 

prominul rs (Casey) 

M. major Lindroth - BC 

M. nigrinus (N'[annerheim) - BC 

fretus (.Ca,sey) 

obispinus (Casey) 

M. pusio (Leconle) - ON 

Genus SYNTC,MUS Hope 

M etable tt.. s Schmidt-Goebel 

s. americanus (Dejean) AK \T( NT BC AB sK MB ON PQ NB NS PE LB NF 

borealis ( 4immermann) 

Genus AXINOPAIPUS LeConte 

Variopalpis Solier 

A. biplagiatus tDejeant - BC AB SK - ON PQ 

Genus CALLtr.IDA Dejean 

(Subgenus CAI,LEIDA s. str.) 

(Spongoloba Chaudoir) 

C. punctata LeConte MB ON PQ 

C. purpurea tsiayt IIB 

cyanipennis Chaudoir 

smaragdina Dejean 

(Subgenus PHILOPHUGA Motschulsky) 

Key to species: Larson (1969) 

C. viridis amorna (LeConte) - BC AB SK MB 

canora (.Casey) 

obscura ( lasey) 

puella (C*ey) 

58

Carabidae 

Genus TECNOPHILUS Chaudoir 


T. croceicollis peigani Larson AB SK 

Genus CYMINDIS Latreille 

(Subgenus PINACODERA Schaum) 

(Pla nes us Motschuisky) 

C. limbata Dejean - ON PQ - NS 

(?) fuscicollis (Motschulsky) 

(?) laeuigata (Motschulsky) 

C. platicollis tSayt - ON PQ 

fuscata Dejean 

uirescens Drrty 

(Subgenus CYMINDIS s. str.) 

C. americana Dejean - ON PQ 

continens Casey 

uenator Dejean 

C. borealis LeConte - BC AB SK MB ON PQ NB NS NF 

C. cribricollis Dejean - YK NT BC AB SK MB ON PQ NB NS - LB NF 

abstrusa LeConte 

acomana Casey 

alternans Casey 

alticola Casey 

coseyi Liebke 

hirbyi Casey 

marginata Kirby 

obliqua Casey 

planifera Casey 

reflexa LeConte 

rupimontis Casey 

sinuata Casey 

tardaLiebke 

C. interior Lindroth AB SK MB 

C. neglecta Haldeman AB SK MB ON PQ NB 

C. pilosa Say - ON PQ 

cribrata LeConte 

pubescens Dejean 

C. planipennis LeConte NT BC AB SK MB - PQ 

bipartita Casey 

b reu i p enni s Zimmermann 

directa Casey 

gouanica Casey 

C. unicolor Kirby AK YK NT BC AB - MB ON PQ - LB NF 

hudsonica LeConte 

parowana Casey 

Genus APENES LeConte 

(Subgenus APENES s. str.) 

(Nominus Motschulsky) 

A. sinuatus (Say) - ON 

pustulatus (Dejean) 

Tlibe Zuphiini 

Genus ZUPHIUM Latreille 

Z. americanum Dejean 

-oN 

Tfibe Galeritinr 

Genus GAJ-ERITA Fabricrus 

Galeritina Jeannel 

Galeritula Strand, 

Key to New World species: Reichardt (1967) 

(Subgenus PROGAIERITINA Jeannel) 

Key to species: BalI & Nimmo (1983) 

59

Haliplidae 

G. janus (Fablicius) 

ahasueru; Liebke 

americana (Fabricius) 

borealis Oastelnau 

caseyi Lel'g 

caseyi Lit:lske 

cordicollis Chaudoir 

cyanipeniis Dejean 

intermed o Casey 

occipitali.; Casey 

thoracica Casey 

MB ON PQ 

Family HALIPLIDAE crawling water beetles 

R.E. Roughley 

Haliplidae is a family of aquatic beetles that 

includes 3 7 species in Canada and Alaska. The 

adults are distinguished among aquatic beetles by 

their enlarged hind coxal plates suspended below 

the abdominal sterna. Adult haliplids swim slowly 

or crawl o r submerged vegetation. Members of 

Brychius are found in erosional portions of streams. 

Those of other genera use a wide variety of freshwater 

habitats but are most often found in permanent, 

clean, nutrient-rich ponds and lakes or in slow 

backwaters of streams. Larvae feed on periphyton 

or algae. Adults are primarily algophagous but may 

be facultative scavengers. 

The hr'liplids have often been placed near 

Dytiscidae in classifications. This position is maintained 

by lleutel and Roughley (1988), who, however, 

sugg(, st that haliplids represent an indepen- 

dent invasion of the water (for alternative views, 

see Kavanaugh 1986). There is no recetrt taxonomic 

revision of the North American fipecies. 

Matheson (1912) and Roberts (1913) provided the 

last comprehensive treatments. Proper species 

identification often requires examination of male 

genitalia. Therefore, previously widespread species 

are found to represent species complexes. A number 

of regional taxonomic works, such as Gordon 

and Post (1965), Matta (1976), Brigham (1982), 

Hilsenhoff and Brigham (1978), and Gunderson 

and Otremba (1988), are useful for interpreting the 

Canadian fauna. 

AK (3); YK (4); NT (9); BC (23); AB (15); liK (10); 

MB (13); ON (17); PQ (18); NB (3); NS (3); LB (2); 

NF(4) . H(1) 

Genus BRYCI'IIUS C.G. Thomson 

Key to North American species: 

B. albertanus Carr 

B. hornii Crotr:h 

B. hungerfordi Spangler 

B. pacificus Carr 

Carr (1928) 

_AB 

BC 

BC 

SK 

MB 

-oN 

Genus HAJ-IPLUS Latreille 

Key to North Arnerican species: 

(Subgenus IL.aI-IPLUS s. str.) 

H. allisonae Brigham 

H. apicalis C.(i. Thomson* 

strigatus Roberts 

H. blanchardi Roberts 

H. distinctus ''Vallis 

H. dorsomacu, atus Zimmermann 

H. falli Mank 

H. immaculicollis Harris 

americanus Aub6. 

impressus Kirby 

H. longulus LeConte 

hoppingiWallis 

H. robertsi Zirnmermann 

pallidus loberts 

H. stagninus lL,eech 

(Subgenus PARAIIAPHLUS Wallis) 

Wallis (1933) 

^K 

YI< Nr BC AB SK MB 

A]{ YK NT 

AK }'K NT 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB SK 

AB 

AB SK 

_-PQ 

MB_PQ 

MB ON PQ 

MB ON 

MB 

PQ 

NB \Tq DF 

-NF 

LB NF 

60

Noteridae 

H. borealis LeConte 

H. connexus Matheson 

H. fasciatus Aub6 

H. leopardus Roberts 

H. pantherinus Aub6 

H. triopsis Say 

(Subgenus LIAPHLUS Guignot) 

H. canadensis Wallis 

H. columbiensis Wallis 

H. cribrarius LeConte 

H. leechi carteri Leech 

H. leechi leechi Wailis 

H. nitens LeConte 

H. salinarius Wallis 

H. salmo Wallis 

H. subguttatus Crotch 

H. ungularis Wallis 

H. vancouverensis Matheson 

_BCAB 

NT BC 

-YKNTBCAB 

_BC 

_BCAB 

NT 

_BC 

_BCAB 

_ABSKMBONPQ 

-MBONPQ ON PQ 

-PQ 

oN PQ 

ON PQ 

NT BC AB 

_BC 

_BCAB 

SK MB 

:" yu 

SK MB 

ON PQ 

oN PQ 

ON 

ON PQ 

NB NS 

NB 

_NS 

_LB 

NF 

NF 

Genus PELTODYTES R6gimbart 

Cnemidotus Erichson 

P. callosus (LeConte) 

P. duodecimpunctatus (Say) 

P. edentulus (LeConte) 

P. lengi Roberts 

P. muticus (LeConte) 

P. torbulosus Roberts 

BC 

BC 

BC 

AB 

AB 

AB 

SK MB 

SK MB 

oN PQ 

ON PQ 

ON PQ 

oN PQ 

oN PQ 

Family AMPHIZOIDAE trout-stream beetles 

Y. Bousquet 

Amphizoidae is a small monogeneric family containing 

the genus Amphizoa, which has four 

species: three occur in western North America and 

one in the People's Republic of China, wesrern 

Szechwan Province. Both larvae and adults are 

found crawling over submerged rocks, debris, or 

wood in rivers, fast-flowing mountain streams, or 

waterfalls. They apparently feed on living stonefly 

nymphs. 

Kavanaugh (1986) recently revised the species 

of this family. 

aK (1); YK (2); BC (3); AB (2) 

Genus AMPHIZOA LeOonte 

Dysmalhes Mannerheim 

A. insolens LeConte 

josephi Matthews 

sah I ber g ii (Mannerheim) 

A. lecontei Matthews 

carinata Edwards 

planata Van Dyke 

A. striata Van Dyke 

AKYK_BC 

YK_BC 

BC 

Family NOTERIDAE burrowing water beetles 

R.E. Roughley 

Noteridae is a family of aquatic beetles that 

includes only two species in Canada. Adults and 

larvae are found in standing water and are associated 

with dense masses of aquatic vegetation. The 

larvae of a European species burrow in the mud and 

take air from submerged plant tissue. Most North 

American larvae crawl on the substrate or pond 

vegetation. Adults are predaceous, and larvae may 

61

Dytiscidae 

be omnivorous. A detailed study of the life history 

of any North American species is needed. 

Taxonomically, Nearctic noterids are well 

known bet:ause all genera have been revised. No 

additional species are expected for the Canadian 

fauna. TlLe classification of noterids is fairly well 

known (Beutel and Roughley 1987), and ther systematic 

position of the family near Dytiscidae is well 

supported (Beutel and Roughley 1988, Kavanaugh 

1986). 

oN (2);PQ (1) 

Genus HYDFi,OCANTHUS Say 

Key to Nevr World species: Young (1985) 

(Subgenus IIIDROCANTHUS s. str.) 

H. iricolor Say 

similato- Zimmermann 

Genus SUPIJISELLUS Crotch 

Key to North American species: Young (1979a) 

S. puncticolli; (Crotch r - 

ON PQ 

ON 

Family DYTISCIDAE predaceous diving beetles 

D.J. Larson and. R.E. Roughley 

In Car'ada and Alaska 262 species of Dytiscidae 

are known, making this the largest family of 

aquatic beetles in the region. These beetles occur in 

a wide valiety of shallow-water habitats. The richest 

fauna is in the emergent vegetation zone along 

the edges of pools, small ponds, and small, slow 

streams. .t\. number of species are characteristic of 

cold-wate: habitats, such as small seepages and 

springs, as weII as bog, forest, alpine, and arctic 

pools. Clear, gravelly streams and saline ponds 

each contain distinctive groups ofspecies. 

Adults are predators or scavengers, and all larvae 

are apparently predaceous. The range of prey 

taken probably is determined by what prey of suitable 

size , s available, but some hydroporines feed 

largely olL microcrustacea, whereas some colymbetines 

prey mainly on culicid larvae. As far as is 

known or can be inferred, aII species in the fauna 

are univo itine or semivoltine. Most species overwinter 

as adults, but time of breeding and occurrence 

of larval stages vary greatly among species. 

Represenuative larvae of most genera have been 

described, but larvae are known for only a small 

percentagr: of species. 

The feLmily is relatively well known taxonomically. 

The principal taxonomic problems include 

relating the northern Nearctic fauna with that of 

the Palaezrrctic; analyzing and interpreting patterns 

of geogral,hical variation, which in many species is 

complex; aind developing suprasp ecifrc classifrcation, 

especially within the larger genera. Identifrcation 

of species may be difficult, as differences between 

species are often subtle. There is therefore a need 

for additional analysis to provide better species definitions 

and characters for their recognition. 

Reviews and revisions of many genera hirrve been 

prepared or are in progress, but no single publicaiion 

treats the entire fauna. The regional works of 

Hatch (1953) and Larson (1975) together ccver most 

of the western and boreal species. Wallis (1973) 

and Larson and Colbo (1983) have publisled fairly 

complete faunal lists for Manito ba and 

Newfoundland, respectively. Leech and Chandler 

(1956) and D.S. White et al. (1984) provide good 

keys to adults and larvae of most North l\merican 

genera. A number of works treating the fauna of 

various states have been published in the United 

States. Some of these works are especially valuable 

in interpreting the Canadian fauna: California 

(Leech and Chandler 1956), Florida (Young 1954), 

Maine (Malcolm 1971), Nevada (La Rivers 1951), 

North Dakota (Gordon and Post 1965), North and 

South Carolina (Brigham 1982), and Utah (R.D. 

Anderson 7962). Zaitzev (1953) outlijned the 

Palaearctic distribution of Holarctic species. 

AK (92); YK (111); NT (118); BC (167); AB {153); SK 

(r24); MB (145); ON (155); pQ (147); NB (82); NS 

(77):PE (31);LB (7a); NF (78) ' H (45) 

Genus COPELATUS Erichson 

Key to No:r'th American species: Young (1963o) 

Subfamily COPELATINAE 

TYibe Copelatini 

62

Dytiscidae 

C. chewolati Aub6 - ON 

renouatus Guiglot 

C. glyphicus (Say) - oN pe - NS pE - NF 

Subfamily AGABETINAE 

Tlibe Agabetini 

Genus AGABETES Crotch 

A. acuductus (Harris) - ON pe 

Genus LACCOPHILUS Leach 

Key to North American species: Zimmerman (1970) 

Subfamily LACCOPHILINAE 

T?ibe Laccophilini 

L. biguttatus Kirbyx 

inconspicuus Fall 

AK YK NT BC AB SK MB ON pe NB LB 

stroehmi C.G. Thomson 

L. maculosus Say 

cal ifornic u s Motschulsky 

- BC AB SK MB ON pe NB NS pE 

descipiens LeConte 

fusculus Sharp 

shermaniLeech 

trunco tus Mannerheim 

L. proximus Say 

americanus Au.b6 

- ON 

confusus Sharp 

L. undatus Aub6 - ON 

Subfamily FfYDROPORINAE 

Tlibe Hydrovatini 

Genus HYDROVATUS Motschulsky 

Key to North American species: Young (1g56, 1g63b) 

H. pustulatus (Melsheimer) 

compressus Shatp 

- ON pe 

cuspidatus auct. amer. 

Genus CELINA Aub6 

Key to North American species: Young (1979b) 

Tribe Methlini 

C. hubbelli Young - ON pe 

Genus DESMOPACHRIA tsabinston 

D. convexa (Aub6) 

Tlibe Hyphydrini 


Genus BIDESSONOTUS R6gimbart 

Key to North American species: Young (1g54) 

B. inconspicuus (LeConte] 

B. pulicarius (Aub6) 

Genus LIODESSUS Guignot 

Key to North American species: Larson and Roughley (1990) 

Tlibe Bidessini 

Key to genera: Young (1967) 

- ON pe - NS 

- ON 

L. affinis (Say) AK yK NT BC AB SK MB ON pe NB NS pE - NF 

charlotti (Clark) 

emilianus (Clark) 

e ry t h r ostotn us (Mannerheim) 63

Dytiscidae 

macularis (LeOonte) 

microrei.iculatzs (Hatch) 

nanus (l\tb6) 

nigrinut (Casey) 

obscurellus (LeConte) 

L. cantralli (''loung) 

L. fl avicollis r LeConte) 

L. fuscatus (Crotch) 

.AB SK MB 

_MBONPQ 

SKMBONPQ_NS_LB 

Genus NEOCLYPEODYTES Young 

N. ornatellus (Fali) 

Genus IJVAIiiUS Guignot 

U. granarius (Aub6) 

U. lacustris (Say) 

U. subplicatt.s (Hatch) 

MB 

ON PQ NB NS 

oN______ 

Genus HYGIIOTUS Stephens 

Coelamitus C.G. Thomson 

Key to North American species: Anderson (1971, 1976, 1983) 

H. acaroides lLeconte) 

winnipe,g (Wallis) 

H. bruesi (Fzll) 

H. compar (tr all) AK 

H. dissimilis Gemminger & Haroid 

dispar (. -eConte) 

H. falli tWallis ) - 

H. farctus (L rConte) 

H. impressol unctatus (Schaller)x AK \-I( 

similis (.Kjtby) 

H. infuscatur (Sharp) - YK 

dentiger (FalI) 

H. 

H. 

H. 

oregonu.; (FalI) 

intermedi -rs iFall,1 

laccophili rus t LeContet 

lutescens ileConte) 

impress /rons Motschulsky 

infacetus (Clark) 

mediali:: (LeConte) 

AK 

H. marklini (Gyllenhal)x AK YK 

canadensis (Fa11) 

uirgo (F:rll) 

H. masculinr s (Crotchl 

fastidiotus (Fa11') 

H. nigrescen r tFallt 

H. novemlinc,atus hudsonicus (Fa11)* AK YK 

H. nubilus tI eConte) 

H. obscurepl rgiatus t Fall t 

columbi znus (W.J. Brown) 

H. patruelis ileConte) AK YK 

borealis (Fall) 

discoideus (LeConte) 

quebecensis (W.J. Brown) 

H. picatus (Iriirby) AK 

H. punctilintatus tFall t 

H. salinarius lWallist 

H. sal Balfour-Browne 

punctatus (Say) 

H. sellatus (LeConteJ 

H. semivittarus tFall) 

H. suturalis rl,eConte) AK 

ouoideur,; (LeConte) 

H. sylvanus ,Fallt 

H. tumidiver tris (Fallt 

H. turbidus (LeOonte) AK 

H. unguicularis (Crotch)* AX 

mongoli:us (Jakovlev) 

Tribe Hydroporini 

_BC 

_BC 

NT BC 

NT BC 

BC 

BC 

sc 

BC 

AB 


AB_MBONPQ 

_ONPQ 

ABSKMB_PQNB 


AB 

AB 

SK 

MB 

SK MB 

_BC 

NT_ABSKMBONPQNBNS 

_BC 

NT 

NT 

NT 

YK NT 

YK 

_NT 

YK NT 

YK 

NT 

NT 

_NT 

YK NT 

YK NT 

SK 

AB SK 

MB 

MB 

MB 

MB ON 

PQ 

-pe 

oN PQ 

PQ 



_ABSKMB 


_ABSKMBON 

BCAB_MB 


ON PQ 



BCABSKMB-PQ 

NS 

NN NS 

NB 

NB 

NS 

NS 

PE 

PE 

PE 

LB 

LB 

LB 

LB 

LB 

in 

NF 

NF 

NF 

NF 

NF 

NF 

lre 

NSpn-- LB NF 

64

Dytiscidae 

Genus FIYDROPORUS Clairville 

(Subgenus HETEROSTERNUTA Strand) 

(H eter o ster nu s Zimmermann) 

(H etero s teth u s Falkenstrom) 

Key to species: Matta and Wolfe (1981) 

H. alleghenianus Matta & Wolfe 

H. cocheconis Fall 

H. pulcher LeConte 

H. wickhami Zaitzev 

concinnus LeConte 

(Subgenus SANFILIPPODYTES Franciscolo) 

H. browni Wallis 

H. brumalis W.J. Brown 

H. compertus W.J. Brown 

H. edwardsi Wallis 

H. falsificus W.J. Brown 

H. pacil-rcus Fall 

H. planiusculus Fall 

H. terminalis Sharp 

s 

H. acadianus Balfour-Browne - 

H. persimilis Crotch 

aequus Fall 

H. paugus Fall AK 

H. rubyae Larson 

H. stagnalis Gemminger & Harold 

collaris LeConte 

(Subgenus NEOPORUS Guignot) 

(C ir cinopo ru s Guignot) 

Key to species: Fa]] (1923) and Wolfe (1984) 

H. blanchardi Sherman 

H. carolinus Fall 

H. clypealis Sharp 

anticus Shato 

H. dilatatus Fall 

H. dimidiatus Gemminger & Harold 

semirufus LeConte 

H. mellitus LeConte 

H. mixtus LeConte 

H. solitarius Sharp 

H. spurius LeConte 

peltatus Shatp 

H. sulcipennis Fall 

H. superioris Balfour-Browne - 

sericeus LeConte 

H. tennetum Wolfe 

H. undulatus Say 

anticus Sharp 

consimilis LeConte 

fasciatus Haris 

placatus Fall 

pubipennis Aub6 

p un ctati s sirn u s Attb6. 

scrutator Sharp 

uelutinus Aub6 

H. vittatus LeConte 

H. vitiosus LeConte 

(Subgenus IIIaDROPORUS s. str.) 

Key to North American species: Fall (1923) 

H. appalachius Sherman 

signatus Sharp 

H. badiellus Fall AK 

H. brevicornis Fall 

H. carri Larson 

H. columbianus Fall 

BC 

BC AB 

BC AB 

BC 

BC 

BC AB 

BC 

YK NT BC AB 

YK NT BC AB 

_NT_AB 

YK 

YK 

YK 

Nr ec 

NT BC 

_AB 

NT BC 

NT BC 

NT BC 

sx 

PQ NB 

_ONPQNBNS-LBNF 

-oNPQNB 

PQ 

y" oN pe 

- - - pq- N3 




_ONPQ 

-oNPQNBNS-LBNF 

_ONPQNBNSNF 

NB 

ON PQ NB 

ON PQ 


ON PQ NB 

_ONPQ NB 

AB SK MB ON 

AB 

AB 

AB 

AB 

AB 

AB 

SK 

SK MB 

SK MB 

SK MB 

s* ms 

-oN 

MB ON PQ 

ON 

oN PQ 

ON 

ON 

ON 

PQ 

NS 

NB NS 

DD 

rD 

pn 

PE 

PQNBNS_LBNF 

PQNBNS_LBNF 

PQ NB 

LB NF 

LB NF 

LB NF 

LB NF 

-NF 

ie 

LB 

NF 

NF 

65

Dy'tiscidae 

H. dentellus Fall AK YK NT BC AB SK MB ON PQ NB NS PE LB 

H. despectus Sharp - YK NT BC AB - MB ON PQ - NS NF 

rusticut Sharp 

H. dichrous Melsheimer - ON PQ 

discicol',is Say 

H. fatigus G rrdon BC 

H. fortis Le('onte BC 

H. funestus Fall - BC - SK 

H. fuscipennis Schaumx AK YK NT BC AB SK MB ON PQ NB LB NF 

criniticrtxis Larson 

piceus ,cittrrn 

puberut'us Mannerheim 

H. lapponunL Gyllenhal AK YK NT BC MB ON PQ - LB NF 

labradc rensis Fall 

[L larsoni NLlsson - YK NT BC AB SK MB - PQ - LB 

lL longiusculus Gemminger & Harold AK BC 

oblongus Aub6 

H. mannerhr:imi Balfour-Browne AK YK NT BC AB 

humerolis Alb6 

H. melsheimeri Fall ON PQ 

H. morio Aur6* AK YK NT BC AB SK MB ON PQ 

caligintsus LeConte 

- LB NF 

me lanor:ep ha I us Gyllenhal 

H. nigellus l{annerheim* AK YK NT BC AB SK MB ON PQ NB LB NF 

H. niger Say - ON PQ NB NS PE - NF 

latifrons Sharp 

modestus Attb6 

H. notabilis LeConte AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

arcticut C.G. Thomson 

H. obscurus Sturm* AK YK NT BC AB SK MB ON PQ NB LB NF 

hockin1i Larson 

H. occidentalis Sharp - YK BC AB SK 

H. pervicinus Fall - BC AB SK MB 

H. polaris Fr,Ll1 AK YK NT MB - PQ 

subuiret;cens Fall 

H. puberulur LeContex AK YK NT BC AB - MB ON PQ NB LB NF 

leuanderi J. Sahlberg 

H. rectus Fall AK YK NT BC AB SK MB ON PQ NB LB NF 

H. rufinasus Mannerheim AK YK NT BC AB SK MB ON PQ - NS 

H. sibiricus,,I. Sahlbergx AK YK NT 

boreus llordon 

H. signatus \{annerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

inornat'"ts Sharp 

youngi k,rdorl 

H. similaris Fall BC 

H. sinuatipe* Fall - BC 

H. striola (Gyllenhal)x AK YK NT BC AB SK MB ON PQ NB NS - LB NF 

ambiguus Aub6 

subtonsus LeConte 

uittula .Eic]l.son 

H. tademus Leech - BC AB 

H. tartaricus LeConte* AK YK NT BC AB - 

cam ina., iu s Motschulsky 

MB ON PQ 

lugubri: Motschulsky 

mel an c nolicu s Motschulsky 

H. tenebrosus LeConte 

luridiptnnis LeConte 


lutulen: us LeConte 

H. transpun-'tatus H.P. Chandler 

H. tristis (Pr,rykull)* 

- BC AB 

AK YK NT BC AB SK MB ON PQ NB NS - LB NF 

elongat:ilus Schiodte 

ru fi co p, ll u s Mannerheim 

uarians LeOonte 

Genus POTITIVIONECTES Zimmermann 

Deronectes auct. amer. 

P otarno dy t e s Zimmermann 

Key to North American species: Zimmerman and Smith (1975a) 

P. aequinoctialis (Clarkt AB 

P. deceptus Fallr - BC 

66

Dy'tiscidae 

P. depressus t Fabricius)* 

P. elegans (Panzer)8 

P. griseostriatus (DeGeer)x AK YK 

catascopium (Say) 

coloradensis (FalIt 

mathiasi (Hatch) 

parallelus (Say) 

suffusus (Sharp) 

P. rotundatus ( LeConte t 

P. spenceri (Leech) - YK 

P. striatellus (LeConte) - YK 

infaustus (Clark) 

pulcher (Motschulsky) 

NT_AB_MB 

_BCABSKMBON 

NT BC AB SK MB ON PQ NB NS 

_BCABSKMB 


pn in Nr 

ON PQ NB NS PE LB NF 

Genus OREODYTES Seidlitz 

Key to North American species: Zimmerman (1985) 

O. alaskanus (Fall) AK YK 

p r o d u c t o t r u n c at u s (Hatch) 

rainieri (Hatch) 

recticollis (Fall) 

O. angustior (Hatcht 

O. congruus (LeContet 

O. crassulus iFall; 

O. laevis (Kirby) 

duodecimlinealzs (LeConte) 

AK YK 

hortensis (Hatch) 

semiclarus (.Fall) 

yukonensis (Fall) 

O. leechi Zimmerman AK YK 

O. obesus cordillerensis Larson - YK 

O. picturatus (Horn) 

O. quadrimaculatus (Horn) 

brodei (.Gellerrnan) 

O. sanmarkii (C.R. Sahlberg)* 

riuolis (Gyllenhal) 

- YK 

O. scitulus {LeConte) 

bisulcatus (Fa],\) 

O. snoqualmie (Hatchl 

BC 

AB 

_BC 

_BCAB 

_BCAB 

NT BC AB MB ON PQ 

_BCAB 

_BC 

_BCAB 

NT 

NT 

BC AB 

BC AB 


NS 

-iuNu 

NF 

Genus LACCORNIS des Gozis 

Agaporus Zimmermann 

Key to species: Wolfe and Roughley (1990) 

L. conoideus (LeConte) - YK 

L. deltoides (Fall) 

L. latens (Fall) 

L. oblongus t Stephens tx 

L. pacificus Leech 

NT 

NT 

BC 

BC 

AB 

AB 

SK MB ON 

ON 

ON 

PQ NB 

PQ NB 

_NF 

Genus AGABINUS Crotcl' 

Key to species: Leech (1941) 

A. glabrellus (Motschulsky) 

A. sculpturellus Zimmermann 

Subfamily COLYMBETINAE 

Tribe Agabini 

Key to North American genera: Leech (1942) 

_BC 

_BC 

Genus AGABUS Leach 

Allonychus Zaitzev 

Anisornera LeConte 

Apator Semenov 

Arctodytes C.G. Thomson 

Gaurodytes C.G. Thomson 

Heteronychus Seidlitz 

Ilybiosoma C'rotch 


A. adpressus Aub6* 

solas Leech 

Fa\l (1922b), Larson (1989) 

AK YK NT BC 

67

Dytiscidae 

A. aeruginosrs Aub6 

disposit'ts Guignot 

A. ajax Fal1 

fuscipennis auct. amer. 

A. ambiguus (Say) 

fimbriatus LeConte 

reticulaLus A,e.b6 

A. antennatus Leech 

clauatus LeConte 

A. anthracin:rs Mannerherm 

scapuloris Mannerheim 

A. approximatus FaIl 

A. arcticus (['aykull)x 

reticulaius (Kirby) 

subfasctttus LeConte 

A. audeni Wl1lis 

A. austinii S.rarp 

A. bicolor (K:rby) 

mutus S.harp 

A. bifarius (F.irby)* 

hessleri Hochhuth 

A. bjorkmanare Hatch 

rectus (LeConte) 

A. canadensir Fall 

A. clavicorniu Sharp* 

uerzs W,J. Brown 

A. col)rynbus :eech 

A. confertus ,eOonte 

A. confrnis (( iyllenhal)* 

longulu: (LeConte) 

ouoideut (Crotch) 

A. coxaiis Shrrpx 

A. discolor (F.arris) 

A. discors Lelonte 

A. disintegrabus (Crotch) 

A. elongatus (Gyllenhal)* 

bryanti Jan 

A. erichsoni tlemminger & Haroldx 

lutosus t Crotch) 

nigroaer,,eus Erichson 

sobrinus Motschulsky 

A. erythropterus (Say) 

A. falli (Zimrnermann) 

semtpunctatus auct. amer. 

sharpi F all 

A. fuscipenni.s (Paykull)* 

ontarior,,is FalI 

A. gagates Arrb6 

A. hypomelar; Mannerheim 

i rregula 'is Mannerheim 

A. infuscatus Aub6 

A. inscriptus (Crotch) 

A. leptapsis (LeConte) 

A. lugens LeConte 

perplexus Sharp 

suturalis (Crotch) 

A. lutosus LeConte 

discolor LeConte 

lecontei lCrotch) 

mimusLeech 

A. margaretae Larson 

A. moestus (Ourtis)* 

borealis Sharp 

nigripalcis (J. Sahlberg) 

A. nectris Leech 

A. oblongulus Fall 

A. obtusatus (Say) 

nitidus lSay) 

A. opacus Aub6* 

gelidus )!a71 

pseudoc' tn fertus Wallis 

sharpi Jacobson 

YK 

YK 

AK YK 

AK YK 

NT BC 

_BC 

AB SK 

AB SK 

NT BC AB SK 

NT BC AB SK 

NT BC AB SK 





AK 

AK 

AK 

YK 

NT 

AK YK NT 

BC AB 

BC AB 

YK_BCABSK 

YKNTBC_SK 

YK NT 

BC 


AK YK NT BC 

AK YK NT BC 

_BC 

AK YK NT BC 

AK 

AK 

YK 

NT 

AK YK NT 

AK \'K 

BC 

BC 

BC 

AB SK 

AB SX 

AB SK 

AB 

AB 

AB 

SK MB 

SK MB 

SK MB 

-oN 

MB ON 

MB ON 

MB ON 

MB ON 

PQ 

PQ 

PQ 

ms oN pe 

MB 

MB 

MB 

MB 

SK MB 

SK 

MB ON 

-oN 

MB 

MB ON 

MB ON 

-oN 

oNoN 

PQ 

oN pe 

NB 

PQ NB NS 

PQ 

PQ 

PQ 

AKYKNTBCAB_MBONPQ 


_YK_BCABSKMBONPQ 

-BC 

-BC 

_BC 

BC 

NT BC 

AB 

AB 

AB 

AB 

SK 

SK 

_MBONPQ 

SK 

SK 

MB 

PQ 

NB 

LB 

NS PE LB NF 

NS- 

NS 

NB NS 

NB 

NS 

ON PQ NB NS 

oN PQ 

oN PQ 

ON PQ 

NB NS 

_NS 

NB NS 

f.fn NS 

NB 

LD 

PE 

LB 

LB NF 

LB 

LB 

NE 

- NS- 

ie- 

_LBNF 

_LBNF 

_LBNF 

NF 

NF 

NF 

LB NF 

68

Dytiscidae 

A. phaeopterus (Kirby) 

A. pisobius Leech 

A. punctulatus Aub6 

aeneolus (Crotch) 

A. semipunctatus (Kirby) 

stridulator Shatp 

A. semivittatus LeConte 

spilotus LeConte 

texanus Sharp 

A. seriatus (Say) 

arcfzs Melsheimer 

intersectus Ctotch 

parallelus LeConte 

striatus Aub6, 

A. smithi W.J. Brown 

A. strigulosus (Crotch) 

nanus (.LeConte) 

A. subfuscatus Sharp 

A. thomsoni (J. Sahlberg)* 

A. tristis Aub6x 

atrdtus Mannerheim 

crotchi Zaitzev 

dubizs Mannerheim 

piceus Zaitzev 

piceolus Zaitzev 

subopocus Mannerheim 

A. triton Fall 

A. vancouverensis Leech 

A. velox Leech 

A. verisimilis W.J. Brown 

A. wasastjernae (C.R. Sahlberg)* 

henaiensis Fall 

palustris Wallis 

A. zattzewi Poppius* 

hudsonicus Leech 

A. zetterstedti C.G. Thomson* 

browni Leech 

AK YK 

_YK 

NT BC AB 

_BCAB 

AB 

AK YK NT BC 

YK NT BC 

AB 


_BCAB 


AKYKNTBCABSKMB-PQ 


AK_-SCT 

AK YK NT 

_BCAB 


AK YK 

AK YK 

NT 

NT 

SK MB ON 

SK MB ON 

SK MB ON 

SK MB 

_MR 

SK MB ON 

PQ 

PQ 

PQ 

PQ 

MB ON PQ 

MB 

MB 

ON 

PQ 

pe- 

NB 

NB 

NB 

NB 

NS 

NS 

NS 

NB NS 

PE 

PE 

_LBNF 

LB NF 

LB 

LB NF 

NF 

LB NF 

LB NF 

Genus CARRI{YDRUS Fall 

C. crassipes Fall 

YK NT BC 

SK MB ON 

Genus ILYBIUS Erichson 


I. angustior (Gylienhal)* 

I. biguttulus (Germar) 

laramaeus LeConte 

L churchillensis Wallis 

I. confusus Aub6 

deniheiWallis 

I. discedens Sharp 

L fraterculus (LeConte) 

I. ignarus (LeConte) 

I. incarinatus Zimmermann 

I. oblitus Sharp 

I. picipes (Kirby)* 

L pleuriticus (LeConte) 

inuersus Sharp 

I. quadrimaculatus Aub6 

I. subaeneus (Erichson)* 

uiridianeus Crctch 

I. vittiger (Gyllenhal)* 

Larson (1987) 

AK 

f_ 

YNTBCYSKMB3NiS 

I'_ 

_MB 

MB 

ON 


--NTBCY:"YuoNpe -oNPQ 

AK "K 

Nr sc AB !x nrn 3N pe 

_ \'K NT BC AB SK MB ON PQ 

AK BC AB 


AK YK NT BC 

NB 

NB 

NB 

NB 

NS PE LB NF 

NS 

NF 

NS 

NS 

NS 

NS 

_LBNF 

__I" 

-isNr 

PE LB NF 

LB NF 

Genus COLYMBETES Clairville 

Cymatopterus Dejean 


C. dahuricus Aub6x 

obscuratus Mannerheim 

seminiger LeConte 

Tribe Colymbetini 

Zimmerman (1981) 


MB ON PQ 

69

Dytiscidae 

C. densus LeConte AK YK - BC AB SK MB 

inaequalLs Horn 

C. doiabratus (Paykull)* AK YK NT BC MB - 

drewsonl. LeConte 

PQ - LB NF 

groenlanCicus At:.b6 

margina'.is (Fabricius) 

thomsont. Sharp 

triseriatu.s I{trby 

C. exaratus LeConte 

i ncog nitr"r s Zimmerman 

- YK NT BC AB SK MB 

rugipenn.is Sharp 

C. paykulli Erichsonx - YK NT BC AB SK MB ON PQ NB NS NF 

longulus LeConte 

C. sculptilis Ftarris AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

Genus NEOS'IUTOPTERUS Balfour-Browne 

P seudo sc utop te r u s Hatch 


N. angustus (LeConte) - YK - BC AB SK MB ON PQ NB NS - LB NF 

N. hornii (Crc,tch) AK YK NT BC AB SK MB ON PQ NB LB 

coriaceut: Hotn 

Genus RHANTUS Dejean 

Key to Nor',h American species: Zimmerrnan and Smith (1975b) 

(Subgenus NI,\RTUS Zaitzev) 

(Rantogilon des Gozis) 

R. sinuatus (l,eConte) NT BC AB SK MB ON PQ - NS 

(Subgenus RIIANTUS s. str.) 

R. binotatus (Harris) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 

aequalis ll.atc]r 

assimilit (Kirby) 

diuiszs ( {ub6) 

flauogriseus Crotch 

longipes Sharp 

obscurus, Shatp 

plebeius Shatp 

R. consimilis Motschulsky NT BC AB SK MB ON PQ 

disceden; Shatp 

loslzs (LeConte) 

R. frontalis (l{arsham)* AK YK NT BC AB SK MB ON PQ 

fla u icoll t s E schscholtz 

gyllenhat i (Castelnau) 

notatus (Fabricius) 

punctatus (Hope) 

sericozs Sharp 

suturalk (Boisduval & Lacordaire) 

R. gutticollis Sayt 

dominguensis Gemminger & Harold 

- BC AB 

hoppingt.Wallis 

hubbelliHatch 

maculiccllis (Arb6) 

mexican, t s (Castelnau) 

R. suturellus (Harris) 

subopacus (Mannerheim) 

AK YK NT BC AB SK MB ON PQ - NS - LB NF 

zimmermanni WalTis 

R. wallisi Hatch AK YK NT BC AB SK MB ON PQ NB NS - LB NF 

suturellus anct. 

Genus COPTJTOMUS Say 

Key to eastern North American species: Hilsenhoff(1980) 

Tribe Coptotomini 

C. longulus LeConte - BC AB SK MB ON PQ NB NS PE 

lenticus Hilsenhoff 

C. loticus Hil;enhoff - ON PQ 

70

Dytiscidae 

Tlibe Matini 

Genus MATUS Aub6 

Key to species: Young (1953) 

M. bicarinatus (Say) 

M. ovatus ovatus Leech 

ON 

ON PQ 

Subfamily DYTISCINAE 

TYibe Dytiscini 

Genus DYTISCUS Linn6 

Leionotus l\trby 

Macrodytes C.G. Thomson 

Key to species: Roughley (1990) 

D. alaskanus Balfour-Browne 

paruulus Motschulsky 

D. circumcinctus Ahrens* 

arrrizs Mannerheim 

ooligbukiilirby 

D. cordieri Aub6 

sublimbatus LeConte 

D. dauricus Gebler* 

confluens Say 

franklinii l{trby 

frontalis Motschulsky 

uentralis Motschulsky 

uexatus Sharp 

D. fasciventris Say 

D. harrisii Kirby 

D. hatchi WaIIis 

D. hybridus Aub6 

compar (Melsheimer) 

D. marginicollis LeConte 

alb ioni c u s Motschulsky 

fu scostr iat us Motschulsky 

D. verticalis Say 

AK YK NT BC 

AK YK NT BC 

AK 

AK 

YK 

AB SK 

AB SK 

NT BC AB SK 

NT BC AB SK 

_YKNTBCABSK 


-BCABsx 

NT 

BC 

AB 

SK 

MB ON 

MB ON 

MB ON 

MB ON 

MB 

MB 

MB 

MB 

MB ON 

PQ 

PQ 

PQ 

PQ 

NB 

ONPQNBNS_LB 

ONPQNBNS_LB 

ON PQ NB 

PQ 

NB NS 

NB 

NS 

PE 

PE 

LB 

LB 

NF 

NF 

NF 

Tlibe Hydaticini 

Genus HYDATICUS Leach 

Key to North American species: Roughley and Pengelly (1982) 

(Subgenus I{YDATICUS s. str.) 

H. aruspex Clark* AK YK NT BC 

americanus Sharp 

laeuipennis C.G. Thomson 

modestus S}:'atp 

H. oiceus LeConte 



NB 

NS 

NS 

PE 

LB 

NF 

Tribe Aciliini 

Genus ACILIUS Leach 

Key to North American species: Hilsenhoff(1975) 

A. abbreviatus Mannerheim AK 

oregonensis Ctotch 

A. athabascae Larson 

A. mediatus (Say) 

maccullochii (Kirby) 

A. semisulcatus Aub6 AK YK 

A. sylvanus Hilsenhoff 

_BCAB 

_ ICT:" 

y:'T:" 

MB ON 

-oN 

MB ON 

MB ON 

PQ 

PQ 

PQ 

PQ 

NB NS 

NB NS 

PE 

LB 

NF 

Genus GRAPHODERUS Dejean 

Key to North American species: Wallis (1939) 

G. fascicollis (Harris) 

oN PQ 

7T

Gytininae 

G. liberus (Sr,Ly) 

brunnipznnis Aub6 

rugicolli.s I{ttby 

thoracic ^ts Haris 

G. manitober.sis Wallis 

G. occidentalis Horn 

G. perplexus Sharp 

elatus S tarp 

NT BC SK MB ON PQ 

_YKN,scABs^MBoNpe 


NB 

NB 

NS 

NF 

Ns po ie Nr 

Genus THERMONECTUS Dejean 

T. basilaris (]{arris) 

-oN 

Tribe Cybistrini 

Genus CYBIfITER Curtis 

C. frmbriolatrrs (Say) 

crotchi \{ilke 

oliuieri (.)totc}' 

MB ON 

NS 

Family GYRINIDAE whirligig beetles 

R.E. Roughley 

Adult lyrinids form a group of beetles wonderfully 

adalrted for life on the water surface; 36 

species ocsur in Canada and Alaska in streams, 

rivers, pords, and lakes. They can be found quite 

far from s rore but are more common closer to the 

shore. Thc adults are readily distinguished by their 

highly mo

Gyrininae 

Tribe Gyrinini 

Genus GYRINUS O.F. Miiller 

(Subgenus GYRINULUS Zaitsev) 

G. minutus Fabricius* 

(Subgenus GYRINUS s. str.) 

G. aeneolus LeConte 

G. aeratus Stephensx 

instabilis Fall 

G. affrnis Aub6 

canadensis R6gimbart 

G. analis Say 

G. aquiris LeConte 

G. bifarius Fall 

G. cavatus Atton 

G. confinis LeConte 

G. consobrinus LeConte 

G. dichrous LeConte 

G. dubius Wallis 

G. fraternus Couper 

G. frosti Fall 

G. gehringi Chamberlain 

G. hoppingi Leech 

G. impressicollis Kirby 

G. latilimbus Fall 

G. lecontei Fall 

G. maculiventris LeConte 

G. marinus Gyllenhalx 

G. marginellus FalI 

G. opacus C.R. Sahlberg* 

hoppingiLeech 

G. pectoralis LeConte 

G. piceolus Blatchley 

G. picipes Aub6 

G. pleuralis Fall 

G. plicifer LeConte 

G. pugionis Fall 

G. punctellus Ochs 

G. sayi Aub6 

lugens LeConte 

G. ventralis Kirby 

G. wallisi Fail 

AK YK NT 

AX 

-YK 

_YK 

AK 

YK 

AK YK 

AK YK 

_YK 

AK YK 

AK YK 

_YK 

YK 

BC 

AB 

NT BC AB 

NT BC AB 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

SK 

SK 

SK 

AB_ 

AB SK 

AB 

AB SK 

AB SK 

_SK 

_SK 

_:* 

AB SK 

AB SK 

AB ;K 

AB SK 

MB ON 

MB ON 

-oN 

MB ON 

MB ON 

MB ON 

MB ON 

MB 

MB ON 

MB ON 

MB 

MB ON 

-oN 

mn on 

-oN 

MB ON 

yu oN 

MB ON 

AB SK MB ON 

AB SK MB ON 

ABSK_ON 

AB 

_SKMBON 

AB SK 

AB SK MB ON 

ABSK_ON 

AB SK MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB 

NB 

NB 

NS 

NB NS 

_NS 

NB 

NB NS 

-NS 

NB NS 

-NS 

NN NS 

Tu_ 

NB 

lln - 

NB NS 

NB NS 

NB- 

PE 

PE 

pe 

pt 

LB 

LB 

LB 

:u 

LB 

LB 

LB 

NF 

NT 

NF 

NF 

NF 

NF 

NF 

NF 

NF 

73

Hydraenidae 

Suborder Polyphaga 

Superfamily Staphylinoidea 

Family FIYDRAENIDAE minute moss beetles 

R.E. Roughley 

Twenty.ssyen species of Hydraenidae are found 

in Canada and Alaska. The family includes minute 

aquatic beeitles, most of which are under 2 mm long. 

Adults are found along the edges of aquatic habitats, 

particularly streams; some also live in semiaquatic 

habitats. Adults and larvae feed on periphy'ton. 

The farnily is sometimes assigned to the superfamily 

Hyclrophiloidea; however, several features, 

such as wing venation and the absence of tire basal 

piece on the aedeagus, leave little doubt about its 

place in the superfamily Staphylinoidea. Perkins 

(1980) recently revised the Canadian and .Alaskan 

species in his monograph of the Hydraenidae of the 

Western Hemisphere. 

NT (a); BC (21);AB (10); SK (4); UB (5); ON (5);PQ 

(7);NB (3);NS (1);NF (1) 

Subfamily IIYDRAENINAE 

Tribe Hydraenini 

Genus HYDR\XNA Kugelann 

H. anguiicollil Notman 

H. atlantica F erkins 

H. circulata Perkins 

H. nigra Hatcr 

H. occidentali r Perkins 

H. pacifica Pe.:kins 

H. pennsylvar.ica Kiesenwetter 

H. petila Perk ins 

H. vandykei d'Orchynont 

NT_AB-MB 

_BCAB__ 

-BC 

_BC 

_BCAB 

_BC 

_BC 

ON 

ON 

PQ NB 

PQ IU 

PQ_ 

NS 

NF 

Genus LIMNEBIUS Leach 

Limnocharis Horn 

L. alutaceus (Casey) 

L. borealis Pelkins 

_BCAB 

_BC 

Genus GYMNOCHTHEBIUS d'Orchwnont 

G. nitidus (Le,lonte) 

Subfamily OCHTHEBIINAE 

NT 

oN PQ 

Genus OCHTII{EBIUS Leach 

(Subgenus OCHTHEBIUS s. str.) 

(Enicocerus Stephens) 

O. benefossus LeConte 

O. bisinuatus Perkins 

O. borealis Pe:rkins 

O. intermptus LeOonte 

aberti }Ia.tc}r 

O. kaszabi Janssens 

O. lecontei Perkins 

O. lineatus LeConte 

milleriHatch 

O. marinus PnLykull 

holmbergi Maklin 

O. rectus LeConte 

(Subgenus ASIOBATES C.G. Thomson) 

(H omaloc ht hebizs Kuwert) 

.(Trymoch thebizs Kuwert) 

O. brevipennir; Perkins 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB__ 

AB SK MB 

AB SK MB 

AB SK MB 

oN PQ 

ON PQ 

NB 

74

Ptiliidae 

O. cribricollis LeConte 

O. discretus LeConte 

O. hibernus Perkins 

O. mimicus W.J. Brown 

_BCABSK 

_BC 

NT 

_BCAB 

MB_ 

PQ 

Genus NEOCHTHEBIUS d'Orchymont 

N. vandykei (Knisch) 

lapid.icola (Van Dyke) 

Family PTILIIDAE feather-winged beetles 

J.M. Campbell 

Members of Ptiliidae, a family of very small beetles, 

have fringed wings with long filaments, which 

give them a feather-like appearance. The family 

comprises about 560 species around the world, of 

which 32 are known to occur in Canada and Alaska. 

Most feather-winged beetles are found in association 

with moist organic matter, such as leaf litter, 

decaying wood, and fungi. 

Ptiliidae is probably the most poorly known 

family of beetles in North America. Undoubtedly, 

when the family is revised, many additional species 

will be found, particularly in eastern Canada. 

Hatch (1957) studied the western species. Bistrom 

and Silfverberg (1979) examined many of the 

generic type species and frxed the generic names. 

Dybas and Dybas (1990) recently published a key to 

the North American genera. Many Canadian 

records require confrrmation, as the identification of 

most species has not been checked by specialists. 

aK (5);BC (16);MB (1); oN (2); PQ (10); NB (5); NS 

(1) . H(1);I(11) 

Subfamily PTILIINAE 

Genus MOTSCHULSKIUM Matthews 

M. sinuatocolle Matthews 

Genus PTENIDIUM Erichson 

P. nitidum (Heer)t 

P. pullum Maklin 

P. pusillum (Gyllenhal)t 

apicale Erichson 

terminale Holdeman 

Genus ACTIDIUM Matthews 

A. crotchianum Matthews 

BC 

_BC 

_BCPQ 

BC 

NB 

_NS 

Genus PTILIOLA Haldeman 

P. brevicollis (Matthews)t 

P. collani Mannerheim 

canadensis LeConte 

-oN 

PQ 

Genus PTILIOLUM Flach 

P. columbianum (Matthews) 

P. obscurum (Matthews) 

P. sharpi (Matthews) 

P. spencei (A]libert)t 

Genus PTILIUM Gyllenhal 

Millidium Motschulsky 

P. minutissimum (Ljungh)t 

_BC 

_BC 

_BC 

PQ 

PQ 

75

Limuiodidae 

Subfamily ACROTRICHIINAE 

Genus NEPHANES C.G. Thomson 

N. titan (Nev'man)f 

Genus PTILI IPYCNA Casey 

P. moerens (ldatthews) 

quadratr.t (Casey) 

MB ON 

PQ 

PQ 

Genus ACROTRICHIS Motschulsky 

Macdonuldium Abdullah & Abdullah 

Trichopteryx Kirby & Spence 

A. castanea (lvlatthews) 

A. cognata (Niatthews)f 

platonoffi Renkonen 

pruinosc Hatdn 

A. dilfinis (Mrtthews) 

A. fasciculari ; (Herbst)i 

A. fungi (Abdullah & Abdullah) 

A. grandicolli s (Mannerheim)f 

A. henrici (M,rtthews) 

fratercuia (Matthews) 

A. insularis (.v{aklin) 

A. intermediz (Gillmeister) 

A. josephi (M,rtthews) 

A. laticollis N aklin 

A. parallelopi peda (Matthews) 

A. sericans (Iteer)i 

A. sitkaensis (Motschulsky)* 

A. thoracica ( Waltl)t 

anthracina (Matthews) 

A. vicina (Ma bthews) 

A. volans (Mctschulsky)i 

fennica l:lenkonen 

A. xanthocer:. (Matthews) 

AK 

AK 

AK 

AK 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

PQ 

PQ 

PQ 

PQ 

Ne- 

NB 

NB 

NB 

Family LIMULODIDAE horse-shoe crab beetles 

J.M. Campbell 

Members of this small family are obligate 

guests of ants. The family comprises about 30 

species distributed in North and South America and 

Australia, lnly one of which is found in Canada. 

Recent workers have placed the limulodids in 

the family Ptiliidae (e.g. Lawrence 1982o), but I 

prefer to nraintain the group as a separate family, 

on basis of papers by Seevers and Dybas (1943) and 

Dybas (19?6). This family has not been previously 

reported in Canada, but one species of Litnulodes, 

which I have tentatively identifred as L. pa,radoxus, 

occurs in southern Ontario. Seevers antl Dybas 

(1943) revised the classification of the farnily and 

included a key to all genera and most North 

American species. 

oN (1) 

Genus LIMU -ODES Matthews 

(Subgenus LIMULODES s. str.) 

L. paradoxus Matthews ON 

76

Leiodidae 

Family AGYRTIDAE agyrtid beetles 

J.M. Campbell 

Eleven species of agyrtids are known in 

America north of Mexico, of which eight are found 

in Canada and Alaska. Adults are scavengers of 

decaying organic material. Larval feeding habits 

are probably similar to those of the adults 

(Anderson and Peck 1985). 

Until recent papers by Lawrence (1982o) and 

Lawrence and Newton (1982), species of this family 

had been included in the family Silphidae. 

However, these studies have shown that agyrtids 

are more closely related to species of the family 

Leiodidae than to Silphidae and therefore must be 

treated as a separate family. The North American 

species are well known taxonomically. The species 

of Canada and Alaska were treated bv Anderson 

and Peck (1985). 

AK (4); NT (1); BC (6);AB (3); ON (1) 

Genus NECROPHILUS Latreille 

N. hydrophiloides Gu6rin-M6neville 

a/er Motschulsky 

N. pettiti Horn 

AK 

BC 

ON 

Genus IPELATES Reitter 

Pelates }Iotn 

Pelatines Cockerell 

I. latus (Mannerheim) 

AK 

BC 

Genus AGYRTES Frrtlich 

(Subgenus AGYRTECANUS Reitter) 

A. longulus (LeConte) 

politus (Casey) 

AK 

BC 

Genus LYROSOMA Mannerheim 

L. opacum Mannerheim 

AK 

Genus PTEROLOMA Gyllenhal 

P. nebrioides W.J. Brown 

Genus APTEROLOMA Hatch 

A. caraboides (Fall) 

A. tenuicorne (LeConte) 

_BC 

_BCAB 

Family LEIODIDAE small scavenger beetles 

S.B. Peclz 

In Canada and Alaska 128 species of Leiodidae 

are known. Those grouped here have been placed 

under many family names, but all share the derived 

(but diffrcult to see) character of a nearly enclosed 

gutter that circles the antennal articulation on the 

dorsal face of segments 7,9, and 10 or 8-10 in 11- 

segmented antennae and segments 7-9 in 10-segmented 

antennae, with internal sensory vesicles in 

some segments. The higher classification here lists 

five subfamilies and follows recent advances 

reported by Lawrence (1982o), Lawrence and 

Newton (1982), and Newton (1984). 

Members of Leiodidae are, for the most part, 

scavengers, both as larvae and adults, feeding on 

various decaying organic materials in usually moist 

forest or freld habitats. The two biggest subfamilies 

are Leiodinae and Cholevinae. Members of the 

Leiodinae subfamily (= Anisotomidae, Agathidiidae), 

or round fungus beetles, are often globose and compact 

in body shape, and many species can contract 

into a ball-like form. Some species are specialized 

for digging in soil, where they feed on subterranean 

fungi; others are specialized predators of slime 

molds; and still others feed on soft fungi or are more 

generally saprophagous (Wheeler 1984). The feeding 

habits of both adult and larval Coloninae (= 

Colonidae) are not known. Members of the 

Cholevinae subfamily (= Catopidae, Leptodiridae), 

77

Leiodidae 

or small carrion beetles, are more elliptical in body 

shape and feed on carrion or dung. Some species are 

specializecl occupants of ant nests, caves (but not in 

Canada), rr the nests and burrows of vertebrates. 

Members irf the Leptininae family (= Leptinidae), or 

mammal rrest beetles and beaver beetles, are nest 

scavengers and ectoparasites on small mammal 

hosts or on beavers. 

Collec;ing Leiodidae usually requires a concentration 

on the particular habits or habitats of the 

genera or species sought. They can be collected in 

the following ways: in baited pitfall traps; by sifting 

and Berlese extraction of litter or fungi; by sweeping 

forest vegc,tation on still, warm, and humid evenings; 

and by use,, of large-area flight intercept traps. 

The leiodid fauna of Canada and Alaska is only 

partly known. Several genera have been reviewed 

and revised recently, and others are being prepared. 

No single publication treats the entire fauna, but a 

generic review of the North American fauna is 

available (Peck 1990). The largest faunal works 

(Hatch 1933, 1957; Jeannel 1936) are no longer 

completely reliable. Distributions and bionomics of 

the species are usually incompletely known, and 

many species in the northern United States may 

eventually be found in southern Canada. 

AK (36);YK (18); NT (16); BC (84);AB (39)t SK (18); 

MB (26); oN (57); PQ (53); NB (1e);NS (15); t B (7); 

NF(10) . H(6);I(1) 

Subfamily LEIODINAE 

Tribe Hydnobiini 

Genus HYDIIOBIUS Schmidt 

H. contortus Hatch 

H. crestonensis Hatch 

H. femoratus Hatch 

H. longulus l,eConte 

longidets LeConte 

H. luggeri H rtch 

H. matthewsii Crotch 

H. obtusus L,eConte 

H. pumilus L,eConte 

hiseri Hatch 

latidens LeConte 

H. simulator W.J. Brown 

H. stacesmit ri Hatch 

H. strigilatu r Horn 

H. substriatrLs LeOonte 

curuid.ens LeConte 

H. validus \[.J. Brown 

AK 

--ec-- 

-BC 

_BCAB 

BC 

_BC 

_BC 

_BC 

_BC 

AB 

SK 

ON 

PQ 

pe 

PQ 

NS 

Genus TRIAIRTHRON Markel 

T. lecontei Horn 

cedonuli'.i Schaufuss 

_BC 

Genus ANO(]DUS LeConte 

Neocyrtlsa W.J. Brown 

Key to species: Daffner (1988) 

A. fusciclavus (Fall t 

sternitu; (Hatch) 

A. insolitus tW.J. Brownt 

A. obsoletus Melsheimer) 

blandis :; ima (Zimmermann) 

A. potens tW.J. Brownt 

A. puritanus {Fallt 

A. secretus (W.J. Brown) 

A. superans Fall) 

Tribe Leiodini 

BC 

BC 

BC 

SK 

oN-- 

_ONPQ 

_ONPQ 

-oNPQ 

AB_MBON 

Genus LIONOTHUS W.J. Brown 

Key to No;th American species: Daffner (1988) 

L. forticornis Daffner 

L. ulkei W.J Brown 

-oN 

-oN 

PQ 

78

Leiodidae 

Genus CYRTUSA Erichson 

Caenocyrta W.J. Brown 

Key to North American species: Daffner (1988) 

C. subtestacea (Gyllenhal)* 

picipennis (LeConte) 

BC 

oN PQ 

Genus LIOCYRTUSA Daffner 

Key to North American species: Daffner (1988) 

L. luggeri (Hatch) 

L. nigriclavis (Hlisnikovsky)* 

Genus ECARINOSPHAERULA Hatch 

E. ecarinata (Horn) 

Genus ISOPLASTUS Horn 

I. fossor Horn 

uncus Wheeler 


_BC_ONPQ 

BC 

oN PQ 

Genus LEIODES Latreille 

L. assimilis (LeConte) 

L. bradorata W.J. Brown 

L. canadensis (W.J. Brown) 

L. collaris (LeConte) 

L. conferta (LeConte) 

L. curvata (Mannerheim) 

morula LeConte 

L. diffrcilis (Horn) 

L. dilatata Hatch 

L. grossa Hatch 

L. impersonata W.J. Brown 

L. lateritia (Mannerheim) 

L. lostine (Hatch) 

L. merkeliana (Horn) 

L. mirabilis (Hatch) 

L. paludicola (Crotch) 

L. puncticollis (C.G. Thomson) 

L. punctostriata Kirby 

indistincta (LeConte) 

laefo (Mannerheim) 

punctatostriato Horn 

punctistriata LeConte 

L. punctulata (GyIlenhal) 

L. rhaetica (Erichson) 

L. rufrpes (Gebler) 

L. serripes Hatch 

L. strigata (LeConte) 

L. tenuis Hatch 

L. triepkei (Schmidt) 

L. valida (Horn) 

yyNrBC 

AKYK_BC 

AK 

BC 

AB 

AB 

AB 

- - - ncy 

-BCAB 

AKBC_ _BCAB 

NT BC 

_BC 

-BC 

AKYK_BCAB 


SK 

SK 

SK 

MB ON PQ 

PQ 

MB 

MB ON PQ 

_ 9*PQ 

-oNpe 

nfn ON Pq 

MB ON PQ 

AKYKNTBCAB_MBONPQ 

AK 


_BCABSK 

AKBCABSK-ONPQ 

_BC 

-YKNT-AB-MBONPQ 

NTBCAB_MBONPQ 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NS 

NS 

NS 

LB NF 

LB NF 

LB NF 

Genus ZEADOLOPUS Broun 

Apheloplastus W.J. Brown 

Key to North American species: Daffer (1988) 

Z. egenus (LeConte) 

impubis (Zimmermann) 

oN PQ 

Genus AGLYPTINUS Cockerell 

A. laevis (LeConte) 

Tlibe Scotocryptini 

oN PQ 

Genus COLENIS Erichson 

C. impunctata LeConte 

Tribe Pseudoliodini 

BC MB ON PQ NB NS 

79

Leiodidae 

Tribe Agathidini 

Genus ANIS()TOMA Panzer 

Key to North American species 

A. amicaW.J. Brown 

A. basalis (Lc,Conte) 

dichroa .LeConte) 

A. blanchard' (Horn) 

A. confusa (Horn) 

interstrialis Hatch 

A. discolor (l\'lelsheimer) 

piceo (Melsheimer) 

A. errans W..1. Brown 

A. expolita S .J. Brown 

polita (LeConte) 

A. geminata r.Horn) 

A. globososa {atch 

globosa LeConte) 

A. horni Whe,eler 

A. humeralis (Fabricius)i 

A. inops W.J. Brown 

A. nevadensi:i W.J. Brown 

bicolor (..Jorn) 

Wheeler (1979) 

AK 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB 

AB SK 

AB_ 

oN _ONPQ_NS 

_9* 

-oN 

-oN 

-oN 

-oN 

MB ON 

-oN 

-oN 

MB 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB 

NB 

NS 

Genus AGAT'HIDIUM P anzer 

Key to North American species: FaIl (1933) 

A. alutaceum Fa11 

A. athabasca:rum Fali 

A. angulare Idannerheim 

A. californicu.m Horn 

A. cavisternum Fa1l 

A. columbianum Fall 

A. concinnuni Mannerheim 

A. conjuncturn W.J. Brown 

A. contiguun Fall 

A. depressun. Fall 

A. difforrnis (LeConte) 

canadense W.J. Brown 

A. effluens Mannerheim 

A. jasperanurn Fall 

A. maculosurr W.J. Brown 

A. obtusum llatch 

A. pulchrurn LeConte 

ma ndib rilatum Mannerheim 

A. rotundulu.n Mannerheim 

hincaidi Hatch 

AK 

AK 

AK 

AK 

AK 

AK 

AK 

YK 

_BC 

_BC 

-BC 

_BC 

-BC 

_BC 

_BC 

_BC 

AB_ 

AB 

-oN 

PQ 

Subfamily CATOPOCERINAE 

Genus CATOPOCERUS Motschulsky 

Homoeosoma ArsLin 

Pinodytr,'s Hotn 

Typhloleiodes Hatch 

C. cryptophagoides (Mannerheim) 

nolilzs l[otschulskv 

AK 

BC 

Subfamily COLONINAE 

Genus COLCN Herbst 

(Subgenus CiJRVIMANON Fleischer) 

C. bidentaturn (C.R. Sahlberg)* 

C. mannerheimi Szl'rnczakowski 

(Subgenus COLON s. str.) 

C. femorale I'Iatch 

C. serripoides Hatch 

C. tibiale Har,ch 

-BCABONPQ 

_BC 

_BC 

_BC 

ON 

Iu_ 

NF 

80

Leiodidae 

(Subgenus MYLOECHUS Latreille) 

C. aedeagosum Hatch 

C. celatum Horn 

C. clavatum Mannerheim 

C. cornplicatum Hatch 

C. dentatum LeConte 

C. discretum Hatch 

C. elongatum Notman 

C. inerme Mannerheim 

C. kincaidi Hatch 

C. magnicolle Mannerheim 

C. nevadense Horn 

C. pribilof Hatch 

C. pusillum Horn 

AK 

AK 

AK 

AK 

_YK 

_BC 

_BC 

_BC 

-BC 

-BC 

_BC 

-BC 

NT BC AB 

_BC 

-oN 

-oN 

MB ON 

PQ 

PQ 

pe 

NB 

LB 

NF 

Subfamily CHOLEVINAE 

Tribe Ptomaphagini 

Genus PTOMAPHAGUS Illiger 

Key to North American species: Peck (1973) 

P. breviorJeannel 

P. nevadicus Horn 

densus Hatch 

piperi Hatc}r 

thomomysiHatch 

BCAB_MB 

ON PQ 

Tribe Nemadini 

Genus NEMADUS C.G. Thomson 

N. brachyderus (LeConte) 

N. decipiens (Horn) 

N. gracilicornis Fall 

N. parasitus (LeConte) 

N. pusio (LeConte) 

--ec--- 

BC 

_MB 

_MB 

9* PQ 

ON PQ 

NS 

T?ibe Cholevini 

Genus CATOPS Paykuli 

C. alpinus sitkanus Jeannel 

C. alsiosus (Horn)x 

C. americanus Hatch 

clauicornis LeConte 

C. basilaris Say 

co d o ueri n u s Mannerheim 

spencianus (Kirby) 

C. egenus (Horn) 

C. gratiosus (Blanchard) 

C. luridipennis Mannerheimx 

b r unnip e nni s Mannerheim 

C. mathersi Hatch 

C. simplex Say 

AK YK NT BC 



AK YK NT BC 

AB 


_BCAB 


-YK_BCAB 

AK 

BC AB 

SK 

SK 

SK 

SK 

MB ON PQ 

MB ON PQ 

MB ON PQ 

MB 

SK_ONPQ 

ON 

oN PQ 

ON PQ 

NB 

NB 

NB 

NB 

NS 

NS 

NS 

NS 

NS_LBNF 

LB 

LB 

NF 

Genus SCIODREPOIDES Hatch 

Sciodrepa C.G. Thomson 

S. fumatus terminans (LeConte) 

S. watsoni hornianus (Blanchard) 

AK 

_NTBCABSKMBONPQ 


YNS 

NF 

Genus CATOPTRICHUS Murray 

C. frankenhaeuseri (Mannerheim) 

AK 

BC 

Genus PRIONOCHAXTA Horn 

P. opaca (Say) 

ON PQ NS 

81

Scydmaenidr,Le 

Subfamily LEPTININAE 

Genus LEPTINILLUS Horn 

L. validus (Horn) AK 

l.K 

NT 

MB ON PQ 

BC AB SK NB 

NF 

Genus PLAI'YPSYLLUS Ritsema 

Platy psitllus Westwood 

P. castoris Ritsema* AK 

castorinus Westwood 

BC 

MB ON PQ 

Genus LEPTINUS P.W.J. Muller 

Key to North American species: Peck (1982) 

L. occidentarnericanus Peck AK 

YK 

BC 

Family SCYDMAENIDAE 

antlike stone beetles 

A. Dauies 

Forty-eight species of Scydmaenidae have been 

recorded in Canada and Alaska. Improvements in 

the current taxonomic status and in the methods 

and frequency of collection of the species should 

greatly in:rease this number. Scydmaenidae tend 

to be common locally, possibly following distributions 

of the mites on which both the adults and larrrae 

feed. They are found chiefly in wet mosses, 

deep forer;t litter, decaying logs and tree stumps, 

tree holes, sawdust piles, and compost; some species 

are associirted with ants and caves. 

LeCorrte (1852) and Casey (1897) described 

most of the North American species of 

Scydmaenidae, but these works included rlany synonyms, 

and the higher classification was confused. 

Marsh (1957) made major improvements for the 

western North America fauna, and recently Franz 

(1985) viewed Casey's North American gemera in a 

worldwide perspective. The last publlication, 

together with recent work by Suter and Newton 

(personal communication), forms the basis of the 

present arrangement. 

AK (5); BC (12);AB (1); SK (1); MB (3); ON (2e); PQ 

(14);NB (8);NS (6);LB (1) 

Subfamily SCYDMAENINAE 

Tlibe Lophioderini 

Genus LOPF:IIODERUS Casey 

L. biformis (.iweklin) 

L. insignis Ni.arsh 

L. myops Ca;ey 

L. similis Mr.rsh 

AK 

BC 

BC 

_BC 

-BC 

Genus DRASITOPHUS Casey 

D. laevicollis Casey 

Tribe Euconnini 

BC 

Genus EUCONNUS C.G. Thomson 

(Subgenus EUCONNUS s. str.) 

E. clavipes (Say) 

pilosico Jlis (LeConte) 

E. lecontei (Sichaufuss) 

E. semiruber Casey 

E. ventralis ilasey 

AB SK 

MB ON PQ 

-oN 

MB 

_NS 

(Subgenus CONNOPHRON Casey) 

E. ater (Casr:y) 

E. brevicornis (Say) 

E. comptus (Casey) 

E. debilitans (Casey) 

E. elongatus (Casey) 

E. flavitarsis (LeConte) 

E. gaudens (Casey) 

ON 

ON 

ON 

oN PQ 

ON 

oN PQ 

ON 

NB 

NB 

NB 

82

Scydmaenidae 

E. illustris (Casey) 

E. novellus (Casey) 

E. optatus (Casey) 

E. oreophilus (Casey) 

E. pertinax (Casey) 

E. pumilus (Casey) 

E. tenebrosus (Casey) 

E. testaceipes (Casey) 

E. trifrdus (Casey) 

ON 

-PQ 

ON PQ 

ON 

ON 

ON 

ON 

ON PQ 

ON 

(Subgenus SCOPOPHUS Casey) 

E. affrnis Casey 

ON 

(Subgenus PSOMOPHUS Casey) 

(Xestophus Casey) 

E. debilis Casey 

E. fatuus (LeConte) 

E. salinator (LeConte) 

ON 

ON PQ NB 

_MBON_NB 

NS 

(Subgenus PYCNOPHUS Casey) 

(Nudatoconnus Franz) 

E. rasus (LeConte) 

ON PQ 

$ 

E. longiceps Fall 

AK 

BC 

Tribe Stenichnini 

Genus MICROSCYDMUS Saulcy & Croissandeau 

Opresus Casey 

M. misellus (LeConter 

oN PQ 

NB 

Genus STENICHNUS C.G. Thomson 

(Subgenus CYRTOSCYDMUS Motschulsky) 

S. badius (Casey) 

S. ovipennis {Casey) 

S. perforatus (Schaumr 

S. turbatus tCaseyt 

(Subgenus BRACHYCEPSIS Brendel) 

_BC 

-oN 

ON 

PQ 

_NS 

NB NS 

S. adjutor Fall 

S. pacificus (Casey) 

S. rothi (Marsh) 

S. subpunctatus (LeOonte) 

(Subgenus PARASCYDMUS Casey) 

AK 

ON PQ 

_NS_LB 

S. corpusculus (Casey) 

oN PQ 

NB 

(Subgenus TAPHROSCYDMUS Casey) 

S. californicus Motschulskv 

AK 

Tfibe Eutheiini 

Genus EUTHEIA Stephens 

E. morae Marsh 

E. scitula Meklin AK 

BC 

Genus \IERAPHIS Casey 

V. lacuna Marsh 

BC 

Tribe Leptoscydmini 

Genus LEPTOSCYDMUS Casey 

L. caseyi (Brendel) 

ON 

PQ 

Tlibe Scydmaenini 

Genus SCYDMAENUS Latreille 

Acholerops Casey 

83

Silphidae 

S. retrusus (Casey) 

S. zimmermz:.nni Schaum 

punctatus Casey 

oll 

PQ 

Family MICROPEPLIDAE micropeplid beetles 

J.M. Campbell 

Micropeplidae is a small family of beetles with 

10 species known in Canada and Alaska. They are 

found in a variety of habitats, including deciduous 

and conifr,,rous leaf litter, inside beaver lodges, in 

Carex clurnps growing in shallow water, on freshly 

cut stumps of fir and cedar and possibly associated 

with the fungus Pyrobema, which is found in forests 

after a firr:. Matthews (1970) described two species, 

M. hoogervdorni and M. hopkinsi, from Pliocene 

deposits irr western Alaska. Little is known about 

the feeding habits of the species; however, both larvae 

and adults are assumed to feed ott fungal 

spores. 

The family is well known in North .t\merica. 

Campbell (1968) revised the New World spr;cies and 

published a catalog of the North Americatr species 

(1984c). 

AK (a); YK (1); NT (1); BC (8); AB (a); SK. (2); MB 

(1); ON (5);PQ (a) ' H(1) 

Tribe Micropeplini 

Genus KALI|jSUS LeConte 

K. nitidus Lc Conte 

BC 

Genus MICROPEPLUS Latreille 

Anhenooeplus Koch 

Key to North American species: Campbell (1973b) 

M. browni Cr'Lmpbell 

M. brunneus Maklin 

M. cribratus LeConte 

M. laticollis li{eklin 

M. punctatus LeConte 

costatus Maklin 

oregonensis Hatch' 

M. robustus ()ampbell 

M. sculptus l,eConte 

obliquus LeConte 

M. smetanai Campbell 

M. tesserula Curtis* 

costatus LeConte 

costipenzis Makiin 

AK 

AK 

AK 

AK 

YK 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

AB 

AB 

SK 

sx 

-oNPQ 

-oN _ONPQ 

ON PQ 

MB ON PQ 

Family SILPHIDAE carrion beetles 

J.M. Campbell 

The ca.rrion beetles belong to a family of about 

175 species found primarily in the Northern 

Hemisphere, with a few species having ranges 

extending into South America, southeast Asia, and 

Australia. Twenty-five species are known to occur in 

Canada and Alaska. Carrion beetles are common, 

and because oftheir large size and often bright colors 

they are frequently collected. Both adults and Iarvae 

ofmost species feed on carcasses ofvertebrates; other 

species are scavengers, and some species of Aclypea 

are phy'tophagous and may be considered to be pests 

ofcultivated crops such as spinach. 

The family name Silphidae is userl in the 

restricted sense of Lawrence (I982a), Lawrence and 

Newton (1982), and Anderson and Peck (198{5). These 

authors exclude the agyrtids, treating them es a separate 

family, Agyrtidae. The Canadian and Alaskan 

species are well known taxonomically, based on a 

handbook ofthe species by Anderson and Peck (1985). 

AK (8); YK (6); NT (9); BC (13); AB (17); SK (18); 

MB (18); ON (18); PQ (15); NB (10); NS (11); PE (7); 

LB (1); NF (6) . H (4) 

84

Silphidae 

Subfamily SILPHINAE 

Genus NECRODES Leach 

Proton ec rod es Portevin 

N. surinamensis (Fabricius) - BC AB SK MB ON PQ NB NS PE - NF 

Genus THANATOPHILUS Leach 

Philas Portevin 

T. coloradensis (Wickham) AK BC 

obalskii (Portevin) 

T. lapponicus (Herbst)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF 

cal ifor ni cus (Mannerheim) 

caudatus (Say) 

tuberculatus (Germar) 

T. sagax (Mannerheim) AK YK NT BC AB SK MB ON PQ 

T. trituberculatus (Kirby) AK - NT - AB SK MB ON 

Genus OICEOPTOMA Leach 

O. inaequale tFabricius) - ON PQ 

O. noveboracense (Forster) AB SK MB ON PQ NB NS PE 

marginale (Fabricius) 

marginata Kirby 

Genus NECROPHILA Kirby & Spence 

N. americana {Linn6} MB ON PQ NB NS 

affinis (I{ttby) 

canadensis (Kirby) 

peltata (Catesby) 

terminata (KJrby) 

Genus HETEROSILPHA Portevin 

H. ramosa {Say) NT BC AB SK MB ON 

ceruaria (Mannerheim) 

Genus ACLYPEA Reitter 

Blitophaga Reitter 

A. bituberosa {Leconte) NT BC AB SK MB 

A. opaca tl-innet* AX YK NT 

Subfamily NICROPHORINAE 

Genus NICROPHORUS Fabricius 

Neonicrophorus Hatch 

N. americanus Olivier 

grandis Fabricius 

- ON pe 

uirginicus Fr6lich 

N. carolinus (Linn6J 

med.iatus Fabricius 

AB 

mysticallis Angell 

N. defodiens Mannerheim 

binotatus Portevin 

AK YK NT BC AB sK MB oN PQ NB NS pE - NF 

conuersator Walker 

hadjakensis Portevin 

lateralis Porte.rin 

mannerheimi Portevin 

plagiatus Motschulsky 

pollinctor LeConte 

N. guttula Motschulsky 

hecate Bland 

- BC AB SK 

immaculosis }{atch 

punctostriatus Pietce 

N. hybridus Hatch & Angell - BC AB SK MB ON 

N. investigator Zetterstedt* AK YK NT BC AB SK MB oN Pe - NS pE - NF 

aleuticus Gistel 

confossor LeConte 

85

Staphylinidae 

infodiens Mannerheim 

mar it imu,s Gu6rin-M6neville 

melsheimeri lltrby 

particeps Fischer von Waldheim 

pollinctor Mannerheim 

N. marginatus, Fabricius 

cordiger Portevin 

labreae Pterce 

latifrons Pierce 

mchittrictei Pierce 

m ontez u r,' ta e Matthews 

obtusiscu lellum Pier ce 

requiescalor Gistel 

N. nigrita Marrnerheim 

alphaPierce 

ruficornit Motschulsky 

N. obscurus K.rby 

melsheimeri LeOonte 

N. orbicollis S,ry 

hallii \{trby 

quadrisignatus Castelnau 

N. pustulatus Herschel 

bicolon N ewman 

fasciatus Portevin 

lordzs Mrnnerheim 

unicolor l)ortevin 

N. sayi Castel rau 

lunatus I eOonte 

luniger Harold 

lunulatus Gistel 

N. tomentosur Weber 

a ngustifosciolzs Portevin 

aurigaster Portevin 

uelutinus Fabricius 

N. vespilloider, Herbst* AK 1T< 

hebes I\trby 

pygmaeu.r; I{trby 

BC AB SK 

BC 

AB SK 

SK 

AB SK 

AB SK 

SK 

NT BC AB SK 

MB ON PQ 

MB 

MB ON PQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

NB NS 

NB NS 

NB NS 

NB NS 

NB NS 

PE 

PE 

PE 

NF 

NF 

Family STAPFIfl INIDAE rove beetles 

J.M. Campbell and A. Dauies 

The role beetles form one of the largest families 

of insects in the world, with over 32 000 described 

species. ,[n Canada, ll29 species have been 

reported, b,ut hundreds more remain to be either 

described or recorded. Most rove beetles are predators 

of oth,:r insects and live on or in the soil, in 

ground litt*:r, moss, or in decomposing organic matter. 

Some r:ove beetles prey on aphids, bark beetles, 

or other ir.sects on trees and herbaceous plants, 

whereas others are specialized feeders on pollen or 

are myceto rhagous. Rove beetles prefer moist habitats 

along t,he margins of streams, lakes, and snowfrelds, 

and in decomposing vegetation, manure, and 

carcasses. They are common in all biomes, ranging 

from tropical forests to tundra well above arctic and 

alpine timberlines. Some species are associated 

with mammal burrows, ant and termite nests, 

intertidal and coastal habitats. and saline flats: 

they can also be found in a variety of habitats 

around human dwellings. 

Several authors are currently investigating the 

higher classifrcation of the family. The system fol- 

Iowed in this checklist has been extensively modified 

from the conventional system used in earlier 

papers. The system is based primarily on a paper 

by Lawrence and Newton (1982), but will uindoubtedly 

be modifred further as more phylogene bic studies 

are completed. Adults of most genera of rove 

beetles (except the genera of Aleocharinae) can be 

identifred with the use of keys by Arnett (1968) and 

Moore and Legner (1974 and 1979) or with keys 

based on the European fauna such as those ofLohse 

(1964) and Lohse et al. (1974). Most larvae have 

not yet been described. Newton (1990) provides keys 

to the North American genera of larvae and adults 

that inhabit soil litter. Other Nearctic genera can 

86

Staohvlinidae 

be identified using the keys by Kasule (1966), based 

on the rove beetle larvae of Britain. 

Taxonomically, the North American fauna of 

many tribes and subfamilies is now well known: 

Quediini and Xantholininae (Smetana Lg7l, 1982); 

most Tachyporinae (various generic revisions by 

Campbell between 1973 and 1988); Oxyporinae 

(Campbell f969); and Pseudopsinae (Herman 1975, 

Newton 1982). The large subfamilies Steninae and 

Staphylininae are becoming well known among the 

Canadian fauna, although comprehensive revisions 

of most genera are not yet available. However, 

except for a few genera, Aleocharinae, the largest 

subfamily of Canadian rove beetles, still remains 

almost totally unknown. Although Seevers (1978) 

provides a key and a redescription ofthe genera and 

tribes of the subfamily, identification even to genus 

remains difficult or impossible without access to 

type specimens. In recent years a few genera of 

North American Aleocharinae have been revised, 

and their species can now be identified. These 

include the genera Aleochara, Myllaena, Gymnusa, 

and Deinopsls by Klimaszewski (1979, 1982, 1984) 

and the genus Falagriini by Hoebeke (1985). Ashe 

(1984,1986) revised adults and larvae ofthe genera 

of the aleocharine subtribe Gyrophaenina. Other 

subfamilies with most species poorly known in 

Canada include Paederinae and Oxytelinae, except 

for the genus Bledius, which was revised by 

Herman (7972a-f986). 

Moore and Legner (1975a) published a catalog 

of the North American Staphylinidae. 

AK (308);YK(r72); NT (155);BC (581);AB (265); SK 

(126); MB (278); ON (aa1); PQ (a60); NB (167); NS 

(195);PE (19); LB (63);NF (171) ' H (88);I (99) 

Genus EMPELUS LeConte 

E. brunnipennis (Mannerheim) 

Subfamily EMPELINAE 

BC 

Genus MEGARTHRUS Curtis 

M. americanus Sachse 

M. arcuatus Hatch 

M. atratus Maklin 

M. excisus LeConte 

M. pictus Motschulsky 

M. sinuaticollis Boisduval & 

Lacordaire* 

angulicollis Maklin 

AI{ 

AK 

Subfamily PROTEININAE 

Tribe Proteinini 

BC 

BC AB 

AK 

BC 


MB ON 

-oN 

PQ IU 

NS PE 

Genus PROTEINUS Latreille 

P. atomarius Erichsonf 

paruulus LeConte 

P. basalis Makiin 

P. brachypterus (Fabricius)* 

P. limbatus Maklin* 

maekliniFavel 

AK 

AK 

AK 

BC 

BC 

BC 

BC 

AB 

-oN 

ON 

Subfamily OMALIINAE 

Tribe Eusphalerini 

Genus EUSPHALERUM Kraatz 

Anthobium auct. 

(Subgenus EUSPHALERUM s. str.) 

E. convexum (Fauvel) 

E. fenyesi (Bernhauer) 

E. pothos (Mannerheim) 

dirn idiat um (Melsheimer) 

E. rugulosum (Meklin) 

E. segmentarium (Meklin) 

E. torquatum ( Marsham)t 

AI{ 

AK 

AK 

SC AB 

BC AB 

ON PQ 

_MBONPQ 

ON PQ 

NB NS 

NB NS 

NB NS 

NF 

NF 

NF 

NF 

(Subgenus ONIBATHUM Tottenham) 

87

Staphylinida I 

E. californicum (Fauvel) 

E. farrarae Hatch 

E. grayae Ha:ch 

E. swauki Ha.tch 

AK 

_BC 

_BC 

BC AB 

_BC 

Genus ACRLTLIA C.G. Thomson 

A. crenulata lHatch) AK 

A. tumidula (Maklin) AK 

Tribe Omaliini 

_BC 

_BC 

Genus PYCNOGLYPTA C.G. Thomson 

Key to Norbh American species: Campbell (19836) 

P. aptera Carnpbell 

P. lurida (Gy lenhal)* AK YK 

Nr--ix 

MB_PQ 

MB ON PQ 

NB 

NB 

NS_LB 

NS 

NF 

Genus ELONIUM Leach 

Acrolocha C.G. Thomson 

E. diffusum ( :'auvel,) 

E. exsculptun tMiiklin r 

E. fractum {Fauvel) 

E. helferi (St.elt 

E. leechi tHa chj 

E. minutum r Olivierl'i 

striatum (Gravenhorst) 

E. rugosum t . {atch r 

BC 

RC 

BC 

BC 

BC 

v_ 

-oN 

MB ON 

PQ 

NB 

NS 

NF 

Genus HAPALARAEA C.G. Thomson 

Phyllodr epa C.G. Thomson 

H. angustior Hatcht 

H. caseyi (Be'nhauer & Schubert) 

pacifica .Casey) 

H. floralis tPr.ykull)+ - 

H. hamata (Fauvel) 

H. humerosa tFauvel) 

H. humilis (Ntaklin) AK 

H. longula (Nl,iklin) AK 

H. megarthrcides (Fauvel) 

H. strigipennLs (Maklin) AK 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB_MBONPQ 

MB ON PQ 

___9*PQ_ 

-AB 

NS 

NS 

NF 

Genus OMALIUM Gravenhorst 

Ochthen'nus Motschulsky 

O. algarum Casey 

fucicola Jasey 

O. callosum llaklin AK 

O. excavaturr Stephenst 

O. flavidum Ilamilton 

O. foraminosum Meklinx AK YK 

O. oxyacanth;le Gravenhorstt - 

O. plagiatum Mannerheim AK 

O. repandum Erichson 

O. rivulare (Faykull)t AK 

Genus PHLOEONOMUS Heer 

(Subgenus PIILOEOSTIBA C.G. Thomson) 

P. iapponicus (Zetterstedt)E AK YK 

argus Lt.Conte 

(Subgenus PHLOEONOMUS s. str.) 

(Distemrnus LeConte) 

P. flavipennis (Maklin) AK 

P. pusillus (Gravenhorst)* AK 

laesicollis (M€iklin) 

P. suffusus (Casey) AK 

Genus MICRI\J-YMMA Westwood 

Key to species: Bdcher (1988) 

BC 

NT BC AB SK 

?- 

_BC 

NT 

BC 

BC 

AB 

AB 

-o^ 

MB ON 

-o^ 

PQ 

PQ 

PQ 

pe- 

PQ 


-oN 

PQ 

NS 

Ns-:Ne 

NF 

88

Staphylinidae 

M. brevilingue Schipdte* AK YK NT 

dicksoni Maklin 

M. marinum (Strctm)* PQ NB - NF 

stimpsoni LeConte 

Genus CORYPHIUM Stephens 

Occlephelinus Hatch 

Tribe Coryphiini 

Key to North American species: Campbell (1978c) 

C. arizonense tBernhauer, - BC 

C. brunneum (Hatch) - BC 

C. hyperboreum (Maklint* AK YK 

C. nigrum Campbell PQ - NS 

Genus SUBHAIDA Hatch 

S. ingratatHatcht AI( BC AB 

S. monticola Campbell AB 

Genus HOLOBOREAPHILUS Campbell 

H. nordenskioeldi (Maklin)x AK YK NT - MB - PQ 

nearcticus (B\ait) 

Genus BOREAPHILUS C.R. Sahlberg 

B. henningianus C.R. Sahlbergx AK YK AB SK MB ON PQ NB 

americanus Notman 

Genus BRATHINUS LeConte 

Key to species: Frank et al. (1987) 

Tribe Anthophagini 

B. nitidus LeConte PQ NB NS NF 

B. varicornis LeConte - ON PQ NB 

Genus ACIDOTA Stephens 

Key to North American species: Campbell (7982a) 

A. crenata (Fabricius)* AK BC AB SK MB ON PQ NB NS PE LB NF 

seriata LeConte 

A. quadrata (Zetterstedt)x AK YK NT BC AB - MB ON PQ - LB 

fr a nkenh ae u s eri Miiklin 

patruelis Leconte 

A. subcarinata Erichson - ON PQ NB NS NF 

Genus OLOPHRUM Erichson 

Lathrium LeConte 

Key to North American species: Campbell (1983o) 

O. boreale (Paykull)x 

helleni Scheerpeltz 

AK YK NT BC AB - MB - PQ 

nig r op ic eum Motschulsky 

quesneliHatch 

O. consimile (Gyllenhal)* 

Iimbatum Maklin 

marginatum (Kirby) 

AK YK NT BC AB SK MB ON PQ NB NS NF 

paruulum Meklin 

O. latum Maklin* 

O. obtectum Erichson 

emarginatum Erichson 

AK YK NT 

- ON PQ 

rotundicolle (Say) 

O. rotundicoile (C.R. Sahlberg)* 

conuexicolle LeConte 


conuercum Meklin 

Genus ARPEDIIIM Erichson 

Key to North American species: Campbell (19840) 

A. cribratum Fauvel - BC MB ON PQ NB NS - LB NF 

89

Staphylinidae 

angulare Fanvel 

columbier,.se Hatch 

Genus EUCNIICOSUM Reitter 

Reuelstoh,:a Hatch 

Key to Nort.r American species: Campbell (19840) 

E. brachyptenLm (Gravenhorst)* 

beringense (Van Dyke) 

E. brunnescens (J. Sahlberg)* 

hoppingi (Hatch) 

E. tenue (LeConte)* 

AK YK 

AK YK 

AK YK 

NT 

NT BC AB 

NT BC AB 

SK 

MB 

MB 

MB 

ON PQ 

oN PQ 

LB 

.B 

NF 

NF 

Genus AMPHICHROUM Kraatz 

Staphygrophis Horn 

A. floribundum LeConte 

flauicorne'Casey 

A. macuiatum (Horn) 

immaculctum Hatch 

A. maculicolle (Mannerheim) 

AK 

BC 

BC 

Genus PELEC OMALIUM Casey 

Heterops -Vlannerheim 

P. laevicolle (LeConte) 

P. puberulum tFauvel) 

P. testaceum (:Vlannerheim) 

binotatun',. Casey 

nigrwn Hatch 

AK 

_BCAB 

BC AB 

_ :a_ I'_ . _ 

Genus LESTE VA Latreille 

Zeslo Bla:kweider 

Paralestetn Casey 

Pseudolesteua Casey 

Teuales C*ey 

L. pallipes Le( jonte 

biguttata LeConte 

picescens LeConte 

ON PQ NB 

NS 

NF 

Genus UNAMIS Casey 

U. columbiensis Hatch 

U. fulvipes Fall 

Genus PHLAEOPTERUS Motschulsky 

Phloeopte,.us LeConte 

Tilea Fatvel 

P. castaneus (Oasey) 

cascadien;is Hatc}' 

P. cavicollis (Fauvel) 

breuipennis Casey 

longipalp;rs Hatch 

P. frosti Hatch 

P. fusconiger trlotschulsky 

P. houkae Hat,:h 

P. lagrandeuri Hatch 

P. loganensis llatch 

Genus VELLICA Casey 

V. longipennis Casey 

Genus OROBANUS LeConte 

O. falli Mank 

O. rufipes Cas,:y 

O. simulator LeConte 

O. tarsalis Hatch 

Genus ARTOCHIA Casey 

A. productifrorrs Casey 

AK 

AK 

AK 

AK 

AK 

_BC 

_BC 

BC 

BC 

BC AB 

_BC 

_BCAB 

_BC 

_BC 

_BC 

_BC 

BC AB 

BC AB 

BC 

BC 

90

Staphylinidae 

Genus XYLODROMUS Heer 

X. concinnus (Marsham)t AK 

Iacustris Casey 

X. depressus (Gravenhorst)t AK 

BC 

BC 

AB 

SK 

Genus ANTHOBIUM Leach 

Lathrimaeum Erichson 

A. atrocephalum (Gyllenhal)* AK 

A. clarki Hatch 

A. fimetarium (Mannerheim) AK 

A. occiduum (Casey) 

A. reflexicolle Casey 

A. sinuosum Hatch 

_BC 

_BC 

_BC 

_BC 

_BC 

AB 

Genus DEINOPTEROLOMA Jansson 

Mathrilaeum Moore 

Key to species: Smetana (19856) 

D. pictum tFauvel) 

D. subcostatum (Maklin) AK 

humerale (Casey) 

-BC 

_BC 

Genus TANYRHINUS Mannerheim 

T. singularis Mannerheim AK 

BC 

Genus TRIGONODEMUS LeConte 

T. fasciatus Leech 

T. striatus LeConte 

BC 

ON PQ 

NS 

Genus PORRHODITES Kraatz 


P. fenestralis (Zetterstedt)x AK YK 

breuicollis Maklin 

P. inflatus tHatcht 

NT BC AB 

_BCAB 

MB 

-oN 

PQ 

PQ 

LB 

Genus OROCHARES Kraatz 

Paradeliphrum Hatc}: 


O. tumidus (Hatch) 

BC 

Genus GEODROMICUS Redtenbacher 

G. brunneus (Say) 

coeszs Erichson 

G. fauveli Casey 

G. integer Casey 

G. rufrpennis Van Dyke AK 

G. schultzi Fauvel 

G. strictus Fauvel 

nigrita Farel 

G. temporalis Casey 

G. verticalis (Say) AK YK 

ouipennis LeConte 

_BC 

_BCAB 

-BC 

BC 

PQ 

o" pe 

NF 

AB MB ON PQ NB 

NF 

Genus MICROEDUS LeConte 

M. austinianus LeConte AK YK 

M. fenderi Hatch 

M. laticollis (Mannerheim) AK 

M. porterae Hatch 

M. rogersi Hatch 

Genus HAIDA Keen 

Key to species: Campbell (19786) 

H. keeni Keen AK 

_BCAB_PQ 

_BC 

_BC 

_BCAB 

_BC 

BC 

91

Staphylinidar,, 

Subfamily OLISTHAERINAE 

Genus OLIST HAXRUS Dejean 

O. megacephz,.lus (Zetterstedt)* AX 

laticeps i,eConte 

O. substriatur; (Gyllenhal)x 

nitidus I eConte 

YK NT 

YK NT 

AB 

AB 

_PQ 

oN PQ 

Subfamily PIESTINAE 

Tribe Piestini 

Genus SIAGCTNIUM Kirby & Spence 

Prognatl, us Berthold 

Key to Nort,h American species: Moore (1975) 

S. americanurn Melsheimer 

conuergeizs Sachse 

S. punctatum LeConte 

S. stacesmith: Hatch 

_BC 

AB 

AB SK 

AB SK 

MBONPQ_NS 

MBONPQ_LB 

NF 

NF 

.Genus TRIG(,NURUS Mulsant 

Key to Nor h American species 

T. crotchii Lellonte 

lecontei flharp 

T. dilaticollis Van Dyke 

T. sharpi Blat.kwelder 

T. subcostatu,' Meklin 

Blackwelder (1941) 

AK 

Tlibe Trigonurini 

BC 

BC 

BC 

Subfamily OSORIINAE 

Tribe Osoriini 

Genus HOLO IROCHUS Erichson 

Neotroch,^ts Blackwelder 

H. laevicaudus (LeConte) 

MB 

Tlibe Lispinini 

Genus CLAVILISPINUS Bernhauer 

C. prolixus (L,-.Conte) 

C. rufescens (,:Iatch) 

BC 

AB 

MB 

PQ 

Genus THOR,\COPHORUS Motschulsky 

Glyptomtt Erichson 

T. costalis (Erichson) 

MB ON PQ 

Genus RENAIDIA Motschulsky 

Eumalus Sharp 

R. canadensis Horn 

R. nigrella (Lt:Conte) 

jubilaea Motschulsky 

---ecAB- 

Tlibe Eleusinini 

_ 9*_ _ _ _ _ _ 

Subfamily OXYTELINAE 

Key to genera: Herman (1970) 

Tribe Coprophilini 

Genus DELEi\STER Erichson 

D. concolor LcConte 

D. dichrous (Ciravenhorst)t 

BC 

oN PQ 

LB 

NF 

92

Staphylinidae 

Genus COPROPHILUS Latreille 

Zonoptilus Solsky 

Key to North American species: Campbell (1979a) 

C. castoris Campbell 

C. sexualis Leech 

Genus SYNTOMIUM Curtis 

Eunonia Casey 

- 9*PQ NB 

AKBCAB_PQ 

AK_PQ _BC 

AK 

BC 

NB 

Key to North American species: Herman (I972a,1976, 1983, 1986) 

S. confragosum Meklin 

S. grahami Hatch 

S. keenianum (Casey) 

S. malkini Hatch 

Genus MITOSYNUM Campbell 

M. vockerothi Campbell 

Genus APLODERUS Stephens 

Phloeonaeus Erichson 

A. linearis LeConte 

Genus BLEDIUS Leach 

Microbledius Herman 

Psamathobledizs Herman 

B. albonotatus Miiklin 

alb idi p ennis Bernhauer 

ornatus LeConte 

B. analis LeConte 

B. annularis LeConte 

Ianguidus Casey 

B. aquilonarius Herman 

B. assimilis Casey 

B. basalis LeConte 

ignauus Casey 

misellus Casey 

B. confusus LeConte 

B. coulteri Hatch 

B. emarginatus (Say) 

troglodytes Erichson 

B. flavipennis LeConte 

agonus Casey 

cuspidatus LeConte 

nelsoniHatch 

tenuis Casey 

B. foraminosus Casey 

deceptiuus FalI 

relictus Fall 

B. fumatus LeConte . 

B. gravidus Casey 

missionensis Hatch 

B. habrus Herman 

B. ineptus Casey 

B. longipennis M,iklin 

B. mandibularis Erichson 

breuidens LeConte 

B. melanocephalus (Say) 

confinis Fall 

B. monstratus Casey 

B. mysticus Fall 

B. nardus Herman 

B. neglectus Casey 

B. nitidicoilis LeConte 

B. omega Herrnan 

B. opacifrons LeConte 

condonensis Hatc}' 

lectus Casey 

piceus Fall 

regularis Fall 

specularis Fall 

AK 

AK 

BC 

AK YK NT BC 

AK YK NT BC 

AK YK 

AK 

AB 

YK NT BC AB 

BC 

BC 

BC 

NT BC 

BC 

BC 

BC 

BC 

nc 

AB_MB 

AB SK MB 

AB 

_MB 

SK 

SK 

SK 

_ONPQ 

MB 

-oNPQ 

MB ON 

AB SK MB ON 

AB 

MB 

SK MB 

SK MB 

_MB 

_MB 

_MB 

MB 

ON PQ 

ON PQ 

oN PQ 

ON PQ 

ON 

oN PQ 

NB 

-NS--- 

NB 

NS 

LB 

)" 

NF 

NF 

93

Staphylinidr,Le 

B. opaculus LeConte 

B. philadelphicus Fall 

dickersoni Notman 

dissimit'.is Fall 

folli Wendeler 

mirlas.Notman 

B. politus Erichson 

nigriceJts Notman 

B. rotundicollis LeConte 

B. rubiginosus Erichson 

B. ruficornis LeConte 

B. semiferrugineus LeConte 

canaliailatus Notman 

B. strenuus tlasey 

arizonensis Fall 

furtiuus Casey 

B. suturalis LeConte 

luteiper nis LeConte 

medialis Fall 

oregonensis Hatch 

pleuralt.s LeConte 

B. tarandus Herman 

diuisus LeConte 

B. tau LeCorrte 

B. turgidus t.)asey 

borealis Blatchley 

bowronensis Hatch 

B. viriosus lr.erman 

B. washingtonensis Hatch 

B. zophus H rrman 

Genus OCH'|HEPHILUS Mulsant & Rey 

Ancyrol horus Kraatz 

O. biimpressus (Meklin)* 

O. columbierLsis (Hatch) 

O. planus (LeOonte) 

Genus THIN ODROMUS Kraatz 

T. corvinus (Casey) 

T. diffusus (tlasey) 

AK 

YK 

-oNPQ 

MB 

MB 

_:c_:"yuo^rF8_ 

NT 

BC 

BC 

AB 


AK YK NT BC 

AB SK MB ON 

AB 


_BC 

_YK_BC 

AK 

AKYK_BC 

AKBC_ONPQ 

BC 

SK 

SK 

SK 

SK 

ON 

MB ON PQ 

MB ON PQ 

MB ON PQ 

_9*PQ 

_NS 

NB 

NB 

NB 

PE 

_NF 

_NF 

_ _ _ _ 

NS 

NF 

NF 

NF 

_v 

_ nc _ _ _ g*PQ _ -_ _ _ _ 

Genus CARI'ELIMUS Leach 

P a rat r a,go p hlo e us Hatch 

Trogophloeus Mannerheim 

C. bilineatus Stephenst 

C. congener Jasey 

C. conjunctus (Casey) 

C. diffusus (tlasey) 

C. harneyi H.atch 

C. nanuloides Hatch 

C. obesus (Kiesenwetter)f 

fo s s ula t u s Motschulsky 

mancus (Casey) 

spectatl;.s (Casey) 

C. vancouvelensis Hatch 

C. volans (Notman) 

-BC 

_BC 

BC 

BC 

BC 

BC 

BC 

BC 

-PQ 

Genus THIN OBIUS Kiesenwetter 

Torrentornus Bietig 

T. alaskanus (Fall) 

T. frizzelli Hatch 

T. teres (Fall) 

.AK 

AI{ 

_BC 

Genus PLA1 YSTETHUS Mannerheim 

P. americamrs Erichson 

P. degener Mulsant & Reyt 

Tlibe Oxytelini 

_:'T:" 

MB ON PQ 

-oN 

94

Staphylinidae 

Genus OXYTELUS Gravenhorst 

O. fuscipennis Mannerheim 

O. Iaqueatus (Marsham)t 

O. montanus Casey 

O. nimius Casey 

O. pennsylvanicus Erichson 

O. sculptus Gravenhorstf 

AKYK-BC _BC 

T_ _ BC 

_BC 

ABSKMBONPQ_NS 

AB 

ON PQ NB 

_ONPQ 

_ONPQ 

_ONPQ 

Genus APOCELLUS Erichson 

A. sphaericollis (Say) 

amabilis Sachse 

AB 

MB ON PQ 

Genus ANOTYLUS C.G.Thomson 

Key to species: Hammond (1976) 

A. insecatus (Gravenhorst)t 

A. insignitus (Gravenhorst) 

A. niger (LeConte) 

A. nitidulus (Gravenhorst)f 

b or eal is (Motschulsky) 

punctatus (LeOonte) 

rugulosus (Say) 

A. rugosus (Fabricius)t 

A. sobrinus (LeConte) 

A. suspectus (Casey) 

A. tetracarinatus (Block)t 

_BC 

_BC 

_BC 

_BC 

_BC 

AB SK 

-oN 

MB 

MB ON 

MB 

MB 

PQ IU 

PQ 

NS 

__Iu 

Subfamily PSELJDOPSINAE 

Key to genera: Newton (1982) 

Genus PSEUDOPSIS Newman 

Key to species: Herman (1975) 

P. montoraria Herman 

P. sagitta Herman 

P. subulata Herman 

T 

_BC 

NF 

Genus ZALOBIUS LeConte 


Z. spinicollis LeConte 

BC 

Genus NANOBIUS Herman 


N. serricollis (LeConte) 

BC 

Genus ASEMOBIUS Horn 


A. caelatus Horn 

BC 

Genus CHARIfYPHUS Sharp 

Chapmania Bernhauer 

P seude le u s is Bernhauer 

Siberia Blackwelder 

Triga Farel 

Trigites Handlirsch 

Key to species: Herman (I972b) 

C. picipennis (LeConte) 

Subfamily PHLOE OCIIARINAE 

Key to genera: Smetana and Campbell (1980) 

oN PQ 

NS 

Genus VICELVA Moore & Leqner 

V. vandykei (Hatch) 

AK YK 

BC 

95

Staphylinidu e 

Genus TACFIINUS Gravenhorst 

Key to North American species: Campbell (1973o), (1983) 

(Subgenus T.ACHINUS s. str.) 

T. addendus Horn 

T. angustatus Horn - YK 

T. apterus M:iklin AK 

T. basalis Erichson* AK YK 

circumcinctus Maklin 

T. beckeri Ci,Lmpbell - YK 

T. brevipennis J. Sahlberg* AK YK 

T. canadensis Horn 

T. contortus Hatch 

uarians Hatch 

T. corticinus Gravenhorstt 

T. crotchii H.rrn 

T. dilatatus l{atch 

T. elongatus Gyllenhal* AK YK 

parallelus Hot:n 

T. fi mbriatus, Gravenhorst 

T. frigidus E ichson AI( YK 

Subfamily TACI{YPORINAE 

Tribe Tachyporini 

_BC 

_BC 

_BC 

NT 

_BC 

propinq tus Mannerherm 

T. fumipennis tSayt 

T. horni Can pbell 

T. instabilis \{tiklin* AK YK 

T. jacuticus j rcuticus Poppiusx AK l-I( NT 

nearctic'ts Campbell 

T. jacuticus r Lllrichi Campbell 

T. lanei Hatch 

T. limbatus itlelsheimer 

T. luridus Er ichson 

T. maculicoll s Maklin 

T. memnonil s Gravenhorst 

batychrous Gravenhorst 

rulzzs Sz.chse 

T. minimus (lampbell 

T. nigricornil; Mannerheim AK 

T. picipes Er chson 

T. quebecensis Robert AK YK NT 

acutus Ilatch 

T. rainieri H rtch 

T. rufipes (D rCeert+ 

T. schwarzi Ilorn 

T. scrutator tlemminger & Harold 

discoideus Melsheimer 

repandu:.s Hotn 

T. semirufus Horn AK 

T. smeLanai tlampbell 

T. stacesmitl i Campbell 

T. subterranlus t Linne rt 

T. tachyporoides Horn NT 

T. thruppi H;rtch - YK 

T. vergatus ( ampbell 

(Subgenus T.,\CHINODERUS Motschulsky) 

T. debilis Ho'n 

NT 

NT 

AB 

AB 

AB 

BC 

BC AB 

BC AB 

BC 

BC 

BC 

BC 

_AB 

AB 

AB 

BC AB 

ec- 

BC AB 

BC AB SK 

BC 

BC AB 

BC 

BC 

BC AB 

BC AB 

BC 

_MB 

ix me 

SK MB 

SK MB 

_MB 

_MB 

_MB 

ON 

ON 

PQ 

PQ 

oN pe 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

PQ 

PQ 

_NS 

PQ_NS 

PQ_NS 

NN NS 

PQ NB NS 

PQ_NS 

PQ 

PQ NB NS 

PQ NB NS 

PQ NB NS 

ON PQ 

_ONPQNBNSPE 


MB 

SX ff,fS 

_MB 

-PQNBNS 

_PQ-NS 

ON PQ NB NS 

oN pe Nn Ns 

ON PQ NB NS 

oN PQ 

LB 

TII 

LB 

NF 

NF 

NF 

NT 

NF 

- ie lrr 

_LBNF 

Genus COPROPORUS Kraatz 

Erchom'ts Motschulsky 


C. ventriculus (Say) - NT 

acuducttrs (Kirby) 

affinis (I{trby) 

flauidus (Casey) 

gibbulus (Erichson) 

maneei liicheerpeltz 

punctul

Staphylinidae 

Genus CILEA Jacquelin du Val 


C. silphoides ( Linn6tt 

BC AB oN PQ NB 

Genus TACHYPORUS Gravenhorst 


(Subgenus PAI-PORUS Campbell) 

T. nitidulus (Fabricius)* 

faber Say 

(Subgenus TACHYPORUS s. str.) 

T. abdominalis (Fabricius)* 

T. borealis Campbell 

T. browni Campbeil 

T. californicus Horn 

tehamoe Blackwelder 

T. canadensis Campbeli 

T. dimorphus Campbell 

T. dispar (Paykull)t 

chrysomelinus arct. 

T. elegans Horn 

T. flavipennis Campbell 

T. inornatus Campbell 

T. jocosus Say 

acaudus Say 

arduus Erichson 

infuscatus Bernhauer 

maculipennis LeConte 

snyderi Blackwelder 

T. lecontei Campbell 

T. maculicollis LeConte 

T. mexicanus Sharp 

alleni Blackwelder 

oregonus Blackwelder 

temacus Blackwelder 

T. nanus Erichson 

T. nimbicola Campbell 

T. ornatus Campbell 

T. pacifrcus Campbell 

T. pulchrus Blatchley 

T. rulomoides Campbell 

T. rulomus Blackwelder 

T. stacesmithi Campbell 

T. transversalis Gravenhorstf 

AK YK NT 

YK_ 

BC 



_BC 

-\'K-BC 

_BC 

_BC 

AK YK NT BC 

AK YK NT BC 

AK YK 

TY 

NT 

NT 

BC 

BC 

BC 

BC 

AB 

v 

SK 

SK 

SK 

SK 

SK 

MB ON PQ 

MB ON PQ 

MB ON PQ 

PQ 

MB ON PQ 

_ONPQ 

-ONPQ 

MB ON PQ 

MB ON PQ 

MB ON 


BCABSKMBONPQ_NS 

BCABSKMB_PQ_NSNF 

AB 

T 

AB 

SK 

-oNPQ 

MB_PQ 

PQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

_ONPQ 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NS 

NS 

NS 

-LB 

_:u 

LB 

NF 

NF 

NF 

NF 

NF 

NF 

NT 

Genus SEPEDOPHILUS Gistel 


S. basalis (Erichson) 

pulicarius (Sachse) 

setifer (Casey) 

S. bisigratus (Horn) 

S. cinctulus (Erichson) 

corticola (Casey) 

S. crassus (Gravenhorst) 

moestus (Say) 

S. ctenidialis Campbell 

S. Iittoreus (Linn6)t 

carissimus Flatch 

hnoxii (LeConte) 

S. marshami (Stephens)t 

S. opicus (Say) 

S. scriptus (Horn) 

S. testaceus (Fabricius)t 

elongatus (Blatchley) 

imbricatus (Casey) 

limuloides (Casey) 

S. velocipes (Casey) 

uirginiae (Casey) 

S. wickhami Campbeil 

ON 

BC 

ON PQ 

ON PQ 

ON 

BC AB MB ON PQ NN NS 

-ABSK 

-PQ 

-PQ 

oN PQ 

ON PQ 

ON PQ 

_NS 

NB NS 

NS 

NF 

I\TF 

97

Staphylinidr.e 

Tlibe Bolitobiini 

Genus LORDITHON C.G. Thomson 

Key to No:th American species: Campbell (1982b) 

(Subgenus LORDITHON s. str.) 

L. anticus (Forn) 

L. appalachi,rnus Campbell 

L. axillaris {,lravenhorst) 

L. cascadensis iMalkin) 

hatchi (llalkin) 

L. diffrcilis Campbell 

L. facilis {Caseyt 

L. niger (Grevenhorstt 

L. poecilus (J,{annerheim t 

L. scutellarii Campbeli 

L. thoracicus thoracicus AK YK 

(Fabricius)x 

cincticoilis (Say) 

occiduus (Casey) 

NT 

BC 

BC 

BC 

BC 

AB 

AB 

MB 

ON 

ON 

-oN 

-oN 

-oN 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB NS 

NB NS 

NB 

NB 

NB 

NS_ 

NS 

NS 

_NF 

_NF 

LB NF 

(Subgenus B OLITOBUS Tottenham) 

L. bimaculat rs (Coupen 

kelleyi l[.alkin 

L. cinctus (Gravenhorsl) 

atricautt.atus (.Say) 

gentilis iLeConte) 

L. fungicola ilampbell AK YK 

L. idahoae Hatch 

L. longiceps lleConte) AK 

elelas (Elernhauer) 

L. notabilis (:ampbell 

L. obsoletus Sayr 

sellolzs Sachse 

L. oregonus l)ampbell 

L. quaesitor Hornt 

rostratus (LeConte) 

BC 

AB 

NT BC AB SK 

AB 

_BCAB 

BC 

MB ON 

-oN 

MB ON 

-oN 

-oN 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB 

NB 

NS 

NS 

NS 

NS 

NS 

PE 

:uy 

Genus CARI'HACIS des Gozis 


C. intrusus ( lornJ 

C. nepigoner:sis (Bernhauer) - 

rz6er (1\lalkin) 

wallisi (Malkin) 

BC 

-MBONPQ 


_NF 

Genus BOLI'IOBIUS Leach 

B. analis tPaykullt* 

cingulai.us arct. 

B. kremeri tldalkin] 

NT 

_BC 

AB 

SK MB ON PQ 

NS 

Genus BRYCITPORUS Kraatz 

B. rubescens (Hatch) 

B. rufescens LeConte 

paruulu.s Casey 

rubidus Leconte 

testaceui LeConte 

B. varians (F atch) 

_BC 

_BC 

_BC 

-oN 

peNe- 

LB 

Genus MYCIITOPORUS Mannerheim 

(Subgenus ISCHNOSOMA Stephens) 

M. pictus Horn 

M. splendidus (Gravenhorst)* AK YK 

elegans Matthews 

inquisiti^ts Casey 

pallidus Mannerheim 

_BCABSK 

NT BC AB SK 

MB ON 

MB ON 

PQ 

PQ 

NB 

NS 

NS 

NF 

NF 

(Subgenus MYCETOPORUS s. str.) 

M. american rs Erichson 

lucidulus LeConte 

MB ON PQ NB 

98

Staphylinidae 

M. brunneus (Marsham) 

M. consors LeConte 

M. discalis Hatch 

M. inquisitus Casey 

M. insignitus Maklin 

M. neotomae Fall 

M. punctatissimus Hatch 

M. punctulatus Hatch 

M. rusosus Hatch 

_BC 


-BC 

AK___f:"yu9*PQIU___ 

_BCAB 

_BC 

_BC 

AK YK NT BC AB SK MB ON PQ - NS 

Subfamily TRICHOPIIYINAE 

Genus TRICHOPHYA Mannerheim 

Eumitocerus Casey 

T. piiicornis (Gyllenhal)f 

BC 

ONPQ_NS 

Subfamily HABROCERINAE 

Genus HABROCERUS Erichson 

H. capillaricornis (Gravenhorst)t 

H. magnus LeConte 

H. schwarzi Horn 

BC-ONPQ 

_AB_MBONPQ 

ON PQ 

_NF 

_NF 

Subfamily ALE O CHARINAE 

T?ibe Oxypodini 

Key to North American genera: Seevers (1978) 

Genus DEVIA Blackwelder 

Dasyglossa Kraatz 

D. congruens (Casey) 

AK_NT 

MB 

Genus GNATHUSA Fenyes 

G. caribou Lohse 

G. tenuicornis Fenyes 

AK YK NT 

_BC 

Genus OCYUSAl{raatz 

O. canadensis Lohse 

AK YK 

Genus ILYOBATES Kraalz 

Gennadota Casey 

L canadensis (Casey) 

-oN 

Genus MOLUCIBA Casey 

M. g:randipennis Casey 

_BC 

Genus OCAJ-EA Erichson 

lsoglosso Casey 

O. vancouveri Casey 

_BC 

Genus OXYPODA Mannerheim 

Hylota Casey 

O. bradorensis Lohse 

O. chillcotti Lohse 

O. demissa Casey 

O. egestosa Casey 

O. famula Casey 

O. frigida Bernhauer 

O. glenorae Casey 

O. hemingi Lohse 

O. hiemalis Casey 

O. impressa Casey 

O. irrasa Meklin 

O. lacustris Casey 

O. lassula Casey 

O. lata Lohse 

YTNT_ 

AK 

AK 

AK 

AK 

AK 

BC 

_BC 

_BC 

_BC 

BC 

_ _ 

PQ 

MB 

_ 9N_ _ 

-oN-- 

-oN-- 

MB 

99

Staphylinida e 

O. leechi Lohse 

O. madgei Lc,hse 

O. manitobae Casey 

O. nelsoni Lchse 

O. optiva Casey 

O. parafunebris Lohse 

O. regressa Clasey 

O. subpolaris Casey 

Genus PENT ANOTA Bernhauer 

P. meuseli Bernhauer* 

_YK 

AK 

AK__- 

_BC 

AK YK NT 

_BC 

AK 

AK 

MB 

MB 

Genus BRACIHYUSA Mulsant & Rey 

Tetralinz Casey 

B. americana (Fenyes) 

B. helenae (C.asey) 

AK 

_BC 

.\T 

Genus GNYI:ETA C.G. Thomson 

Euliusa Casey 

G. brevicornis Casey 

G. caerula (C.R. Sahlberg)'l 

G. manitobae Casey 

BC 

_MB 

NF 

Genus MER()NERA Sharp 

M. venustula (Erichson) 

Genus ACROTONA C.G. Thomson 

A. adjuvans Oasey 

A. onthophik Lohse 

Genus HYDTrOSMECTA C.G. Thomson 

H. pseudodior;ica Lohse 

AKYK_ 

YK 

Tribe Athetini 

-oNPQ 

_ _ _ 9"_ _ _ _ _ _ 

Genus CANASTOTA Casey 

C. canadensis; (Casey) 

-oN 

Genus DATOMICRA Mulsant & Rey 

Hilarina:. Casey 

Micromt:ta Casey 

Oligomia Casey 

Taxicere.tla Casey 

D. surgens (Casey) 

D. wrangeli (tlasey) 

AK 

BC 

Genus DIME'IROTA Mulsant & Rey 

Dalotia ()asey 

Dimetrotina Casey 

Engamotla Casey 

D. alaskana ( Oasey) 

D. altaica (Berrnhauer)* 

D. bradorensrs Lohse 

D. burwelli L,rhse 

D. campbelli -ohse 

D. caribou Lohse 

D. dempsterensis Lohse 

D. moesta Miiklin 

maeklini (Fenyes) 

D. munsteri (Bernhauer)* 

D. nearctica Lohse 

D. omissa Casey 

D. picipennis (Mannerheim)x 

carlottae (Casey) 

D. prudhoensis Lohse 

D. regissalmonis Lohse 

yv_ 

AK 

- 

BC 

_ 

AK \'K 

_YK 

-YK 

AK 

AK YK NT 

AK YK NT 

_BC 

AK 

AK 

AK 

BC 

MB 

pa:) 

DN !q 

oat 

!q 

LB 

100

Staphylinidae 

D. resplendens Casey 

D. retrusa Casey 

D. smetanai Lohse 

D. stercoris (Fenyes) 

columbica (Bernhauer) 

D. venti Lohse 

D. vicaria Casey 

AK YK 

YK 

_BC 

-BC 

NT 

_BC 

BC 

MB 

pe 

LB 

Genus PSEUDOTA Casey 

P. cornicula Casey 

P. formalis Casey 

P. irrupta Casey 

P. nanulina Casey 

P. nescia Casey 

P. vana Casey 

_BC 

_BC 

_BC 

_BC 

_BC 

_BC 

Genus AMISCHA C.G. Thomson 

Colposura Casey 

A. analis (Gravenhorst)t 

NF 

Genus GEOSTIBA C.G. Thomson 

Sibiota Casey 

G. circellaris (Gravenhorst)t 

impressula Casey 

G. parvipennis (Bernhauer) 

AB 

NF 

Genus BOREOSTIBA Lohse 

B. campbelliana Lohse 

B. hudsonica Lohse 

B. lagunae Lohse 

B. lamellifera Lohse 

B. sibirica (Maklin)* 

AK YK 

-YK----PQ----- 

_YKPQ 

AKYKNT_PQ_LB 

Genus ADOTA Casey 

Panalota Casey 

A. comparabilis (Maklin) 

A. maritima (Mannerheim) 

massettensis Casey 

-AK 

AK 

BC 

Genus ANEPSIOTA Casey 

A. quadricollis Casey 

A. terminalis Casey 

renominata Bernhauer & Scheerpeltz 

_BC 

_BC 

Genus ATHETA C.G. Thomson 

Elytrusa Casey 

Stonalota Mannerheim 

A. circulicollis Lohse 

A. granulata (Mannerheim) 

nomadica Casey 

A. martini Lohse 

AK YK 

- I't( 

NT 

PQ 

MB-PQ-LB 

Genus ATHETALIA Casey 

A. metlakatlana (Bernhauer) 

A. nimia (Casey) 

_BC 

_BC 

Genus ATHETOTA Casey 

A. atriventris Casey 

A. wickhami (Casey) 

BC 

BC 

Genus LAMIOTA Casey 

L. achromata (Casey) 

L. concessa (Casey) 

L. keeni (Casey) 

L. profecta (Casey) 

BC 

BC 

BC 

BC 

101

Staphylinidae 

Genus LIOGLUTA C.G. Thomson 

P s e udon; eg ist a Bernhauer 

L. aloconotoic.es Lohse 

L. granulosa Lohse 

L. insolens (Casey) 

L. nigropolita (Bernhauer) 

L. nitens (Maklin) 

L. trapezicollis Lohse 

L. vasta (Maklin) 

AK 

_BC 

_YK 

AKYK-BC 

_YK 

AK \.K 

PQ 

LB 

LB 

Genus MOCITA Mulsant & Rey 

M. amblystegii (Brundin tx 

M. breviuscula (Maklin) 

prudens (Casey) 

AK YK NT 

AK 

BC AB 

MB: 

Genus AI-AO.BIA C.G. Thomson 

A. sparreschr eideri (Munster)+ 

Genus SCHISiTOGLOSSA Kraatz 

S. aubeiodes .llrundin 

S. holmbergi Brundin 

Genus PSEUIfOUSIPAJ-IA Lohse 

P. microptera. Lohse 

AK 

-AK 

.AK 

AK 

YK 

YK 

Genus BORE fPHILIA Benick 

B. caseyi Loh;le 

B. caseyiana..,ohse 

B. chillcotti Lrhse 

B. eremita (R re)* 

B. fusca (C.R. Sahlberg)x 

B. gelida (J. Sahlberg) 

B. hyperborei,. (Brundin)* 

R. insecuta (F ppelsheim)* 

B. islandica (.iftaatz)x 

B. manitobensis Lohse 

B. nearctica l,ohse 

B. nomensis (Casey) 

B. subplana tJ. Sahlbergt* 

AK YK NT 

_YK 

AK-MB 

AK_NT 

AKYKNT_MB 

AK_NT 

AK YK 

AK YK NT 

AK_MB 

AK YK 

AK 

AK_NT 

MB 

MB 

MB 

PQ 

PQ- 

LB 

LB 

Genus CLUSIOTA Casey 

C. claviventris Casey 

Genus DINA-IiA-EA C.G. Thomson 

D. borealis Lchse 

D. planaris (I\Ieklin) 

AKYKPQ__-__ 

BC 

Genus BESSOBIA C.G. Thomson 

B. cryptica Lchse 

PQ 

Genus HOMIIOTUSA Casey 

H. manitobae (Casey) 

MB 

Genus MICRODOTA Mulsant & Rey 

Hilara Mrlsant & Rey 

Sonomota Casey 

M. globicollis (Bernhauer) 

M. holmbergi (Bernhauer) 

AK 

ON 

Genus PARAI)ILACRA Bernhauer 

P. glenorica ({)asey) 

BC 

Genus PHILI{YGRA Mulsant & Rey 

P. awemeana (Casey) 

P. botanicaru.m Muonaf 

MB 

N" 

L02

Staphylinidae 

P. criddlei (Casey) 

P. erudita (Casey) 

P. falcifera Lohse 

P. junii Lohse 

P. laevicollis (Meklin) 

P. leechi Lohse 

P. malleoides Lohse 

P. polaris (Bernhauer)x 

P. prognata (Casey) 

P. pseudoboreostiba Lohse 

P. ripicoloides Lohse 

P. rostrifera Lohse 

P. satanas (Bernhauer) 

P. stylifera Lohse 

P. surrufa (Casey) 

P. varula (Casey) 

_YK 

AK 

NT 

AK YK NT 

AK YK NT 

_YK 

_ 5NT 

BC 

-T 

Genus XENOTA Mulsant & Rey 

Delphota Casey 

Donesia Casey 

H alob r e cthina Bernhauer 

Nemota Casey 

Roualida Casey 

X. apposita (Casey) 

X. brumalis (Casey) 

X. cursor (Meklin) 

X. districta (Casey) 

X. fanatica (Casey) 

X. fungi (Gravenhorst)t 

X. mollicula (Casey) 

X. mordax (Casey) 

X. pallitarsis (Kirby) 

X. postulans (Casey) 

X. pratensis (Maklin) 

X. pugnans (Fenyes) 

militaris (Casey) 

X. relicta (Casey) 

X. remulsa (Casey) 

X. resplendens (Casey) 

X. rurigena (Casey) 

X. subintima (Casey) 

X. sumpta (Casey) 

X. tenuicula (Casey) 

X. wrangelica (Casey) 

AK 

AK 

AK 

AK 

_BC 

_BC 

_BC 

_BC 

_BO-_ 

_BC 

_BC 

_BC 

-BC _BC 

_BC 

_BC 

_:r_ 

_ 

MB 

MB 

v 

Genus LYPOGLOSSA Fenyes 

M eg acrotona Scheerpeltz 

L. angularis (Maklin) 

L. fenyesi (Bernhauer) 

AK 

YK 

-BC-ONPQ 

_BC 

NF 

Genus PONTOMALOTA Casey 

P. Iuctuosa Casey 

BC 

Genus STROPHOGASTRA Fenyes 

S. penicillata Fenyes 

MB 

Genus TARPHIOTA Casey 

T. debilicollis Casey 

T. fucicola (Maklin) 

T. geniculata (Maklin) 

insolita Casey 

T. seditiosa Casey 

AK 

AK 

BC 

BC 

BC 

Genus TRICHIUSA Casey 

T. columbica Casey 

T. postica Casey 

BC 

ON 

103

Staphylinidae 

Genus AJ-ECiCHARA Gravenhorst 

Key to North American species: Klimaszewski (1984) 

(Subgenus CI)PROCHARA Mulsant & Rey) 

(.Eucharina Casey) 

(Funda Blackwelder) 

Tribe Aleocharini 

A. bilineata Gyllenhalt 

anthomviae Sprague 

- BC AB - MB ON PQ - NS PE - NF 

ontarionis (Casey) 

A. bimaculata Gravenhorst 

deserticc'la (Casey) 

NT BC AB SK MB ON PQ - LB 

innocua (Casey) 


recta (Casey) 

A. suffusa (Casey)* 

acumincta (Casey) 

AK BC AB - PQ 

perturbc,ns Fenyes 

A. sulcicollis Mannerheim AK BC 

cylindre']a (Casey) 

debilicornis (Casey) 

rugosa (Jasey) 

tibialis ( Casey) 

A. verna Say': AK YK - BC AB SK MB ON PQ - NS PE LB NF 

(Subgenus XINOCHARA Mulsant & Rey) 

A. castaneipcnnis Mannerheim 

acomantl (Casey) 

AK YK - BC AB ON PQ 

concurr(,ns (Casey) 

eludens .Casey) 

glenorar.a (Casey) 

insulant. (Casey) 

mannertteimi (Casey) 

oregona Hatch 

rotundicollis (Casey) 

A. fumata Gravenhorsti 

affiuens (Casey) 

- YK - BC AB - MB ON PQ NB NS PE 

defecta (Casey) 

languidu Sachse 

A. inexspectata Klimaszewski - - PQ - NS 

A. lacertina Sharp - BC AB SK MB ON PQ NB NS 

den s iu er, t r is Bernhauer 

idonea ('t)asey) 

imbricata (Casey) 

salicola lCasey) 

A. lanuginosaL Gravenhorstf - BC AB ON PQ NF 

A. sculptivenbris (Casey) - ON PQ NB - NF 

A. tristis Gravenhorstf PQ 

(Subgenus Cr\I-OCHARA Casey) 

(.OreochcLra Casey) 

A. laramiens s (Casey) AB SK 

A. rubricalis Caseyt BC - ON 

A. rubripenn s (Casey) MB ON PQ 

rubripes Blatchley 

wallawallae Hatc}r 

A. speculicollis Bernhauer AB ON PQ 

A. villosa Mannerheimf AK BC AB 

(Subgenus AI,EOCHARA s. str.) 

A. curtula (Greze)t 

bugnioni Fa:uel 

- BC - ON PQ NB NS NF 

fuscipes (Linn6) 

A. gracilicornis Bernhauer* 

americata Casey 

NT BC - SK MB ON PQ - NS 

ellipsicoi.lis Casey 

kansana Casey 

pleuralis Casey 

A. Iata Gravr nhorst+ - ON PQ 

A. assiniboin Klimaszewski - YK - BC - SK MB ON - 

r ufo nig r z Klimaszewski 

t04

Staphylinidae 

A. tahoensis Casey 

montanica Casey 

postpicta Casey 

A. thoracica Casey 

collusor Casey 

A.' sekanai Klimaszewski 

unicolor Klimaszewskr 

AK YK 

YK NT BC AB SK MB ON 

AB 

-oN 

MB ON 

PQ 

NS 

(Subgenus ECHOCHARA Casey) 

A. ocularis Kiimaszewski 

MB ON PQ 

(Subgenus EMPLENOTA Casey) 

(Polistoma Casey) 

(Polystoma Stephens) 

A. curtidens Klimaszewskr 

A. litoralis (Miiklin) 

arenaria (Casey) 

longiceps (Casey) 

maritima (Casey) 

quadrifer (Casey) 

trilimbata (Casey) 

A. pacifica (Casey) 

_BC 

AK BC PQ 

BC 

NB 

NS 

NF 

Genus PHLOEOPORA Erichson 

P. arctica Lohse 

NT 

Genus PAROCAI-EA Bernhauer 

P. nearctica Lohse 

P. pseudobaicalica Lohse 

AK YK NT 

AKYKNT_PQ 

Tribe Hoplandriini 

Genus HOPLANDRIA Kraatz 

Key to species: G6nier (1989) 

(Subgenus HOPLANDRIA s. srr.r 

H. lateralis (Melsheimer) 

arizonica Casey 

carinata Casey 

ochracea Ktaatz 

texana Casey 

(Subgenus LOPHOMUCTER Notman) 

H. klimaszewskii G6nier 

H. Iaevicollis (Notman) 

Genus CORDALIA Jacobs 

C. obscura (Gravenhorst)f 

Genus MYRMECOCEPHALUS Macleay 

Lorinota Casey 

Stenagria Sharp 

Tlibe Falagriini 

Key to North American species: Hoebeke (1985) 

ON 

_PQ 

-PQ 

ON 

PQ 

PQ 

M. arizonicus (Casey) 

acomanus (Casey) 


M. cingulatus (LeConte) 

sinuosus Casey 

tenuicornis Casey 

M. gatineauensis Hoebeke 

BC 

oN PQ 

PQ 

NB 

NS 

Genus FAI-AGRIA Leach 

F. dissecta Erichson 

angulata Casey 

ery t hr optera Melsheimer 


NS 

105

Staphylinidae 

globosa l{elsheimer 

iowana (iasey 

ithacana Casey 

sterilis Casey 

subsimilis Casey 

tetana Casey 

F. sulcata (Paykull)t 

sicula Jekel 

AB 

ON 

PQ 

Genus ALEODORUS Say 

Chitalia Sh.ary 

A. bilobatus (llay) 

canadent:is (Casey) 

nigresceris (Casey) 

turbatus (Casey) 

A. intricatus (Casey) 

A. scutellaris iLeConte) 

granulos ts (Casey) 

illustris (Casey) 

AB 

SK 

ON 

ON 

PQ 

Genus DRUS;LLA Leach 

D. cavicollis ((lasey) AK 

Tribe Myrmedoniini 

PQ 

Genus XENO DUSA Wasmann 

Pseudolomechuso Mann 

Key to Nonh American species: Hoebeke (1976) 

X. cava (LeCo nte) 

lobata Cr,sey 

probata l)asey 

X. reflexa (W:,Llker) 

Genus ZYRAfi Stephens 

Platyusa Casey 

Z. criddlei (Ce.sey) 

Z. loricatus Casey 

BC 

ON 

SK MB ON PQ 

MB 

-oN 

Genus GYRO PIIAENA Mannerheim 

(Subgenus G\ROPHAENA s. str.) 

G. affinis C.R. Sahlbergf 


inconspir.:ua Casey 

subpunctata Casey 

G. antennalis Casey 

G. californica Casey 

G. caseyi Seevers 

G. criddlei Casey 

G. egena Casey 

exilis Ca,:ey 

G. flavicornis Melsheimer 

G. gaudens C,rsey 

G. geniculata Meklin 

G. gilvicollis Casey 

G. insolens Casey 

G. keeni Casey 

G. modesta Casey 

G. nana (Paykull)* 

perpolita Casey 

G. nanoides Seevers 

G. sculptipennis Casey 

Tlibe Gyrophaenini 

Subtribe Gyrophaenina 

Key to North American genera: Ashe (1984) 

Key to North American species: Seevers (1951) 

AK 

AK 

YK 

YK 

BC MB PQ NB 

ec----- 

BC 

BC 

AB 

AB 

_MB 

MB 

-PQ 

ON PQ 

oN PQ 

oN PQ 

NB_ 

; --- 

ON-NB 

ON PQ NB 

NB 

ON 

ON PQ 

_PQ 

106

Staphylinidae 

G. uteana Casey PQ 


G. vitrina Casey 

dttonsa Casey 

- ON PQ 

(Subgenus PHAENOGYRA Seevers) 

G. gracilis Seevers - 

2 

G. subnitens Casey MB ON 

Genus EUMICROTA Casey 

E. socia lErichsont PQ 

Genus BOLITOCHARA Mannerheim 

Ditropalia Casey 

Pleurotobia Casey 

Silusid.a Casey 

Stictalia Casey 

Venusa Casey 

B. arcuata (Casey) 

B. carlottae (Casey) 

B. densicollis (Casey) 

B. notata M:iklin 

AK 

Subtribe Bolitocharina 

_BC 

_BC 

-BC 

Genus LEPTUSA Kraatz 

Dianusa Casey 

Eucryptusa Casey 

Ulitusa Casey 

L. fontana (Casey) 

-oN 

Genus HOMALOTA Mannerheim 

H. depressiuscula Mannerheim AK 

Subtribe Homalotina 

Genus PLACUSA Erichson 

P. turbata Casey - BC 

Genus CYPHEA Fauvel 

C. wallisi Fenyes 

Genus AUTAI-IA Leach 

A. brevicornis Casey 

A. truncatula Casey 

Subtribe Cyphina 

Tlibe Autaliini 

_BC 

_BC 

MB 

TYibe Phytosini 

Key to western Norbh American species: Moore (1956) 

Genus AMBLOPUSA Casey 

A. borealis Casey 

A. brevipes Casey 

pallida Casey 

AK 

AK 

BC 

BC 

Genus DIAULOTA Casey 

D. densissima Casey AK BC 

insolita Casey 

107

Staphyiinida: 

Genus LIPAIiOCEPHALUS Maklin 

L. brevipennis Miiklin 

L. cordicollis LeConte 

Genus THIN JSA Casey 

T. fletcheri Casey 

diuergens Casey 

nigra Ct.sey 

robustulz Casey 

AK 

AK 

BC 

BC 

Tribe Gymnusini 

Key to North American species: Klimaszewski (1979) 

Genus GYMIIUSA Gravenhorst 

G. atra Casel'* 

G. brevicollis (Paykull)t 

G. campbelli Klimaszewski 

G. grandicepr; Casey 

G. konopacki. Klimaszewski 

G. pseudovariegata Klimaszewski 

G. smetanai .l(] imaszewski 

AK YK NT BC 

AK YK NT 

AK YK NT 

AKYKNT_AB 

AK YK NT 

MB ON 

-oN 

MB ON 

MB 

MB ON 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

NB 

NB 

NB 

NS 

lrs 

NS 

_LBNF 

_LB 

_ 

NF 

NF 

I,B NF 

NF 

Tribe Deinopsini 

Kev to North American species: Klimaszewski (1979) 

Genus DEIN,JPSIS Matthews 

D. canadensi,i Klimaszewski 

D. harringtorLi Casey 

D. rhadina K limaszewski 

-oN 

-oNPQ 

-oN 

NB 

NS 

NF 

Genus MYLI.AENA Erichson 

M. arcana Casey 

intmunci.a Casey 

M. audax Car;ey 

abdita Casey 

dissimuLans Casey 

molesta Casey 

uegeta Casey 

M. decreta Casey 

impellen.s Casey 

scobinel't.a Casey 

umbra Qasey 

M. fenyesi Bcrnhauer 

esuriens Casey 

M. insomnis ilasey 

M. Iudifrcans Casey 

breuiues;lis Casey 

M. procidua Casey 

M. vulpina B:rnhauer 

Genus CYPHA Leach 

Hypocyphtus Gyllenhal 

C. crotchi Horn 

Tribe Myllaenini 

Kev to North American species: Klimaszewski (1982) 

AK 

NT 

BC 

BC 

AB 

AB 

BC 

yl'BCysKyuSN 

Tlibe Oligotini 

BC 

AB 

ON PQ 

oN PQ 

PQ 

PQ 

PQ 

NB 

NB 

NB NS 

NS 

108

Staphylinidae 

Genus OXYPORUS Fabricius 

(Subgenus PSEUDOXYPORUS Nakane & Sawada) 

O. lateraiis Gravenhorst 

breuis Melsheimer 

O. occipitalis Fauvel 

O. quinquemaculatus LeConte 

(Subgenus OXYPORUS s. str.) 

O. femoralis femoralis Gravenhorst 

pulcher Ziegler 

O. kiteleyi Campbell 

O. rufipennis LeConte 

O. stygicus Say 

O. vittatus vittatus Gravenhorst 

cinclzs Gravenhorst 

dimidiatus Melsheimer 

fasciatus Melsheimer 

Subfamily OXYPORINAE 

Key to North American species: Campbeil (1969), (19780) 

_ T_:"ysKy" 

MB 

-oN 

MB ON 

Subfamily STENINAE 

Key to North American species: Casey (1884) 

ON PQ 

oN oNPQ-NS 

ON PQ 

NB NS 

PQ 

PQ_NS 

PQ_NS 

PQ NB NS 

Genus DIANOUS Samouelle 

D. chalybaeus LeConte 

c oer u les cen s Blatchley 

D. nitidulus LeConte 

zephyrus Casey 

AK 

AK YK 

NT 

BC 

AB 

SK 

oN PQ 

-PQ 

NB 

_NS 

NF 

NF 

Genus STENUS Latreille 

Laestris Melsheimer 

(Subgenus "HEMISTENUS" Motschulsky) 

S. croceatus (Casey) 

S. kryzhanovskii Puthzx 

S. niveus Fauvels 

uislo Sanderson 

S. pubescens fraternus (Casey) 

S. pubescens sandersonianus Puthz 

S. retrusus (Casey) 

S. umbratilis (Casey)x 

(Subgenus IIYPOSTENUS Rey) 

(Arezs Casey) 

S. advena (Casey) 

S. aiexanderi Puthz* 

S. annularis Erichson 

S. arculus Erichson 

S. cubensis Bernhauer 

S. dissentiens (Casey) 

hubbardi (Casey) 

leuiceps (Casey) 

lucidus (Casey) 

politulus (Casey) 

politus Sanderson 

simiolus (Casey) 

S. flavicornis Erichson 

geniculatus Say 

S. fulvicornis Stephensf 

S. insperatus Puthz 

S. lugens (Casey) 

perbreuis L. Benick 

S. pinguis (Casey) 

trajectus (Casey) 

utensis (Casey) 

S. plicipennis (Casey) 

agnatus (Casey) 

/orzs L. Benick 

- - Nr ec AB ix Yu oN PQ - - - - - 

AK I'K NT _ MB 

AK YK 

AK YK 

AK 

AK YK 


_BC 

NTBCAB-MB 

ON 

-PQ 

NT-MBONPQ 

-BCAB-MBONPQ 

_ _ _:"TSKyuSNPQ 

AK 

-oN 


BC 

BC 

BC 

AB 


SK MB ON PQ 

SK 

lrn oN 

MB 

MB 

PQ 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NS 

NS 

lls - 

NS 

_v 

LB NF 

_T" 

-Ne 

109

Staphylinidae 

S. pollens (Casey) 

patens (Casey) 

S. punctatus Erichson 

artus ((.asey) 

dilutus lCasey) 

obsoletus (Casey) 

S. quebecensis Puthz 

S. reconditus breviusculus Puthz 

S. reconditus, reconditus (Casey) 

canader.sis (Casey) 

propinquus (Casey) 

S. rossi Sanc erson 

S. varipes (Casey) 

(Subgenus P ARASTENUS Heyden) 

S. pterobrachys Gemminger & Harold 

breuipermis Maklin 

S. sibiricus J. Sahlberg* 

alpicolo Sanderson 

confusus Casey 

deducto" (Casey) 

exaspentlus L. Benick 

montan'"ts Casey 

noctiua1lus Casey 

wenzeli Prt]az 

(Subgenus SIENUS s. str.) 

S. ageus Caseyx 

rigidus Casey 

S. alveolatus Casey 

S. angustus'Jasey 

indigen.; Casey 

mendan Casey 

pettiti C.asey 

placidu.; Casey 

tutnicol,'is Casey 

S. assequens Rey* 

nanulu:, Casey 

pusio Clsey 

S. austini Cr.sey 

amplifir::atus L. Benick 

S. bilentigat rs Casey 

murphyanus Bernhauer 

S. bilineatus J. Sahlberg* 

coruus Casey 

tacomar: Casey 

S. brivioi Pu,hz 

S. callosifror.s Puthz 

S. canaliculatus Gylienhal* 

congener Maklin 

S. cariniceps Miiklin 

S. carinicollis Casey 

S. caseyi Puthz 

S. clarostigma Puthz 

S. clavicornis (Scopoli)t 

S. colon Say 

carolintts Bernhauer 

S. colonus E:ichson 

arizonae Casey 

championi Sharp 

chapini Blackwelder 

floridan,us Casey 

inermis Sharp 

integer Dasey 

interpres L. Benick 

militaris Casey 

obstrusus Casey 

suspectus Blatchley 

weiseri Betnhaner 

S. columbianus Sanderson 

AK 

AX 

YK 


-oN 

AK_NTBCABSK 

AX 

BC 

AK_NTBCAB 

AK YK 

NT 

_BC 

NT 

_BC 

NT BC 

NT BC 

NT BC 

PQ 

MB ON PQ 

MB ON PQ 


-MBONPQ 

MB 

MB 

ON 

ON 

MB ON PQ 

MB ON 

-oN 

MB ON 

MB 

MB 

ON 

NB NS 

NB 

NB ft 

AK 

AK NT BC AB MB ON PQ NS 

AK YK 

AK YK 

AK 

AK 

AK 

AK 

YK NT 

YK NT 

BC 

BC 

BC 

AB 

AB 

AB 

AB 

AB 

AB 

SK 

SK 

SK 

SK 

MB 

PQ 

PQ 

PQ 

PQ 

PQ 

oN PQ 

ON 

oN PQ 

ON PQ 

ON PQ 

_PQ 

ON 

PQ 

NB NS 

NB NS 

NS 

NB NS 

NB NS 

-NS-- 

NB NS 

NB 

NB 

NS 

PE 

PE 

NF 

NF 

LB NF 

LB NF 

NF 

NF 

LB NF 

:u 

NF 

NF 

NF 

110

Staphylinidae 

S. comrna LeConte* 

punctiger Casey 

S. coriaceus Puthz* 

S. dolosus Casey 

jejunus Casey 

uillosus Casey 

S. egenulus Puthz 

S. egenus Erichson 

S. eilipticus Casey 

S. erythropus Melsheimer 

S. exploratus Fall* 

incanoides Prt}:rz 

S. fasciculatus J. Sahlberg* 

S. femoratus Say 

S. fulvoguttatus Notman 

S. geminorum Puthz 

S. hyperboreus J. Sahlberg* 

falli Scheetpeltz 

frigid,us Fall 

S. insularis Casey 

S. intrusus Casey* 

S. jacuticus Poppius* 

S. juno Paykull* 

S. kamtschaticus Motschulskyx 

aquilonius L. Benick 

S. koebelei Puthz 

raniermontis Ptt]az 

S. Iabilis Erichsonx 

S. laccophilus Cdsey 

anastamozans Casey 

edar Notman 

rugifer Casey 

uerticosus Casey 

S. latipennis J. Sahlberg* 

S. latissimus hatchi Sanderson 

S. lustrator Erichsont 

S. mammops Casey* 

humilis Blatchley 

rulomus Blackwelder 

S. maritimus Motschulsky 

S. melanarius Stephens* 

S. melanopus (Marsham)l 

S. mendosus Puthz 

S. morio Gravenhorst* 

diues Casey 

enod.is Casey 

haplus Casey 

indistinctus Casey 

luculentus Casey 

subgriseus Casey 

terricola Casey 

S. mundulus Casey 

S. neglectus Casey 

S. occidentalis Casey 

S. parallelopipedus Mdklin 

adspector Maklin 

animatus Casey 

conuictor Casey 

S. paululus L. Benick 

campbelliPrthz 

S. pauperculus Casey 

brumalis Casey 

S. planatus Puthz 

S. pluto Casey 

S. pudicus Casey 

enodis Casey 

juuencus Casey 

minor Casey 

sibyllinus L. Benick 

S. pumilio Erichson* 

atomarius Casey 


AK-NT 

AKYKNT_ABSKMBONPQNB 

TY 

AK YK 

AK YK 

AKYK 

NT 

NT 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

AB 

AB 

AK 

BC 

AK YK NT 

NT 


AK YK 

BC 

AK YK NT 

AK_NTBCAB 

AK YK NT 

AKYK_BCAB 

AK \'K NT BC AB 

AK 

vvv_ 

BC AB 

_ 


_BC 

AK-NTBCABSKMB 

AK BC AB 

AK 

YK 

AK YK NT 

BC 

SK MB ON PQ 

-MBONPQ 

ON PQ 

SK 

sr 

SK 

SK 

SK 

- ftfS 

_MB 

s^ me 

MB ON PQ 

MB ON PQ 

-oNPQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

MB ON 

MB ON 

-pe 

oN PQ 

NSNS_ 

NB 

NB NS 

NB NS PE 

NB 

oNpeNs-- 

ON 

ON 

ON PQ NB 

PQ_ 

PQ 

-oN 

MB ON PQ 

-oN 

MB ON PQ 

LB 

LB 

LB 

LB 

NF 

NF 

NF 

Ne Ns Fe is Nr 

NF 

111

Staphylinid re 

S. pusillimus Puthz 

S. scabiosus Casey 

S. schwarzi Oasey 

S. scrupeus Casey 

S. sculptilis Casey 

S. sectator Oasey 

S. semicolon LeConte 

S. sordidus .l)uthz 

S. sphaeroprir Casey 

S. strangula.tus Casey 

S. stygicus Say 

S. tenuis Casey 

dfficilis Casey 

eriensis Casey 

ingratus Casey 

inornatus Casey 

paralle,-us Casey 

pauper Casey 

subtilis Casey 

S. venustus Casey 

S. vespertinrrs Casey 

incultu:; Casey 

luctuos;ts Casey 

S. vexatus C.asey 

S. vicinoides Puthz 

S. vicinus Crrsey 

simplet Casey 

tricuspidatus L. Benick 

S. vinnulus ilasey* 

confusoLdes Renkonen 

AK 

AK 

AK 

AK 

AK YK 

YK rVr ec AB ix 

_NTBCABSK 

BC 

_NTBCABSK 

_ *t_ y 

\'K NT BC AB SK 

_ABSK 

_BC 

_BC 

_BCAB 

_BCAB 

NT 

AB 

SK 

MB ON PQ NB 

MB ON PQ NB 

MB ON PQ NB 

MB ON PQ 

lo*peNn 

MB ON PQ 

_ONPQNB 

_ONPQNB 

MB ON PQ NB 

_ONPQ 

MB 

MB 

ON 

MB ON PQ 

NB 

NS 

NS 

NS:u_ 

NS 

NS 

NS 

LB 

NF 

v 

NF 

(Subgenus lESNUS Rey) 

(MutintLs Casey) 

S. californicrrs Casey 

S. curtus Casey 

S. formicetorurn Mannerheim+ 

borealis M,iklin 

S. gratiosus Casey 

hirsutu; Casey 

S. immargin atus Miiklin* 

monticrla Casey 

uariabi,;.is Prthz 

S. tahoensis Casey 

pugetensis Casey 

uacuus Casey 

BC 

_NT_AB_MBONPQ 

YKAB_MB_PQ 

AK YK NT BC 

BC 

AB 

-oN 

MB 

PQ 

PQ 

NB 

NS 

NS 

Genus NOR.lf ENSKIOELDIA J. Sahlberg 

Key to species: Puthz (197h) 

N. columbiana Puthz 

Subfamily EUAESTHETINAE 

_BC 

Genus STIC'IOCRANIUS LeConte 

Key to species: Puthz (7974b) 

S. puncticeps LeConte 

ON 

Genus EUAIISTHETUS Gravenhorst 

E. americanus Erichson 

E. brevipennis Casey 

E. floridae Casey 

E. ganglbaueri Bernhauer 

E. iripennis Casey 

E. laeviusculus Mannerheim 

E. pacificus Jlall 

E. pugetensis Hatch 

T_ I'- 

____ 

AK l'K NT 

AK 

_BC 

BC 

T_ 

T:" 

MB 

MB 

ON PQ NB NS 

ON PQ NB 

_NS 

oN PQ 

oNPQ-NS 

oNPQ-NS 

oNPQNBNS-- 

NF 

t12

Staphylinidae 

Subfamily LEPTOTYPHLINAE 

Genus CHIONOTYPHLUS Smetana 

C. alaskensis Smetana 

4K 

Subfamily PAEDERINAE 

Key to genera: Blackwelder (1939b) 

Tlibe Pinophilini 

Genus PAJ-AMINUS Erichson 

P. luteus Casey 

-oN 

Tlibe Paederini 

Subtribe Medonina 

Genus LITHOCHARIS Deiean 

Metaxyodonta Casey 

(Subgenus PSEUDOMEDON Mulsant & Rey) 

(Ramona Casey) 

L. capitula {Casey) - BC 

L. obsoleta tNordmann)t - BC 

L. thoracica {Casey) MB ON PQ NB 

clarescens (Casey) 

ruficollis (Casey) 

(Subgenus LITHOCHARIS s. str.) 

L. ochracea (Gravenhorst)'l 

alutacea Casey 

- BC - ON PQ NB 

quadricollis Casey 

Genus ACHENOMORPHUS Motschulsky 

(Subgenus ADEROCHARIS Sharp) 

A. corticinus tGravenhorst) MB ON PQ 

millepunctatus (Say) 

rufescens Nordmann 

Genus SUNIUS Curtis 

Medonella Casey 

Micromedon Casey 

Oligopterus Casey 

Xenocharis Bierig 

(Subgenus CAI-ODERMA Casey) 

S. rugithorax Hatch - BC 

(Subgenus IIYPOMEDON Mulsant & Rey) 

(Asteria Fauvel) 

(Hemimedon Casey) 

(Lena Casey) 

S. melanocephalus (Fabriciustf PQ 

S. peregrinus (Casey) - BC 

discolor Casey 

S. quadripennis Casey - BC 

(Subgenus TRACHYSECTUS Casey) 

S. confluentus rSay) MB ON PQ NB NS 

confluens Erichson 

Genus MEDON Stephens 

Oxymedon Casey 

(Subgenus MEDON s. str.) 

M. americanus Casey PQ 

M. fusculus (Mannerheimt+ - - PQ 

M. guppyi Hatch - BC 

113

Staphylinidr.e 

(Subgenus PLATYMEDON Casey) 

(Paramzdon Casey) 

M. capitalis Flatch 

M. convergens (Casey) 

M. insularis Casey 

M. laticollis iCasey) 

M. mcleodi llatch 

M. pugetensis Hatch 

M. shastanic,us (Casey) 

M. truncatu'r Hatch 

M. vancouve,ri (Casey) 

borealil Casey 

AK 

_BC 

-BC 

_BCAB 

_BC 

_BC 

BC 

_BC 

MB 

(Subgenus X'IEDONODONTA Casey) 

M. ctenopho:um Hatch 

_BC 

Subtribe Scopaeina 

Genus ORUII Casey 

(Subgenus FYCNORUS Casey) 

Key to North American species: Herrnan (1965) 

O. dentiger (LeConte) 

humilis (Notman) 

AK - MB ON PQ 

iowanus Casey 

(Subgenus C RUS s. str.) 

Key to North American species: Herman (1965) 

O. distinctu, Casey BC 

O. punctatu,r Casey BC 

filius C*ey 

pallidus Casey 

sonomae Casey 

Genus SCOI'AEUS Erichson 

Leptorus Casey 

Pseudo"us Casey 

(Subgenus SCOPAXOMA Casey) 

S. brunniper LeConte - BC AB 

(Subgenus SCOPAXUS s. str.) 

S. notangulrLs Casey PQ 

(Subgenus -cCOPAEOPSIS Casey) 

S. ventralis Casey) - ON 

Subtribe Stilicina 

Genus PACI{YSTILICUS Casey 

P. hanhami (Wickham) 

Genus RUG.[LUS Leach 

Stilicosoma Casey 

R. biarrnatus (LeConte) 

abbreuiellus Casey 

R. dentatus Say 

luculentus Casey 

minusculus Casey 

nigrolucens Casey 

R. fragilis (Gravenhorst)t 

dngustatus (Fourcroy) 

s cutella t us Motschulsky 

R. latiusculus (Casey) 

R. oregonus (Casey) 

R. rufrpes Germarf 

Genus MEGASTILICUS Casey 

M. formicar:us Casey 

_BCAB_MBNB 

MB ON PQ NB 

-oN 

_ONPQ_NS 

AK- - 

sc- - - il- - - - - - 

-oN 

rt4

Staphylinidae 

Genus LATHROBIUM Gravenhorst 

(Subgenus APTERAI-IUM Casey) 

L. sibiricum Fauvelx AK 

(Subgenus ABLETOBIUM Casey) 

L. shermani Fall 

(Subgenus LATHROBIUM s. str.) 

(Litolathra Casey) 

L. amplipenne Casey 

deceptiuum Casey 

L. arrnatum Say 

procerum Casey 

L. confusum LeConte 

L. divisum LeConte 

L. fauveli Duvivier 

simplex LeConte 

L. fulvipenne (Gravenhorst)t - 

p unct ulat u m Mannerheim 

L. hesperum (Casey) 

L. leconteanum Scheerpeltz 

concolor LeConte 

L. nigrolineum (Casey) 

L. obtusum Casey 

L. shoshonicum (Caseyr 

L. simile LeConte 

L. sparsellum Casey 

L. spissicorne Casey 

L. vancouveri Casey AK 

L. washingtoni Casey AK 

(Subgenus LATHROBIOMA Casey) 

L. othioides LeConte 

L. scolopaceum (Caseyt 

L. tenue LeConte 

(Subgenus LATHROLEPTA Casey) 

L. debile LeConte 

(Subgenus TETARTOPEUS Czwalina) 

(Deratopeus Casey) 

Key to North American species: Watrous (1980) 

L. angulare LeConte 

nigrescens (Casey) 

semirubrum (Casey) 

L. convolutum Watrous 

L. furvulum {Casey) 

L. Iacustre (Caseyt 

L. nigrum LeConte AK YK 

finitimum LeConte 

stibium (Casey) 

L. nitidulum LeConte 

bicolor LeConte 

bicoloratutn Bernhauer & Schubert 

paruipenne (Casey) 

L. punctulatum LeConte AK 

capitosum (Casey) 

hebes Casey 

L. rubripenne tCasey) 

agitans (Casey) 

callidwn (Casey) 

L. sollicitum Fall AK YK 

L. tetricum (Caseyt 

canoaensis (Bierig) 

nigriceps Notman 

Subtribe Lathrobiina 

-BC 

BC 

-ABSKMB 

BC 

BC 

AB 

AB 

_BC 

NT BC AB 

NT 

NT BC 

_MB 

_MB 

ON 

ON 

oN PQ 

ON PQ 

oN PQ 

-MBONPQ-NS-- 

_MBONPQNBNS 

ON PQ 

SKMBONPQ_NS 

ON PQ 

nc - - MnoNFS 

ON 

PQ 

-ONPQ 


MB ON PQ 


NT BC AB SK MB ON PQ 

AB 

ON PQ 

ON PQ 

NB NS PE 

f"NS: 

PQ NB NS 

ON_NBNS 

PE 

NB NS PE 

NB NS 

NB NS PE 

NF 

NF 

NF 

NT 

LB N" 

NF 

NF 

$ 

L. puncticolle Kirby 

MB 

115

Staphylinidr:e 

Genus LOBIiATHIUM Mulsant & Rey 

(Subgenus EULATHROBIUM Casey) 

(Lathro.iropis Casey) 

L. grande (LeConte) 

L. subseriatum (LeOonte) 

ustulatum Casey 

(Subgenus LOBRATHIUM s. str.) 

(Lathro,Siella Casey) 

(Lathro;laxis Casey) 

L. caseyianum ( Scheerpeltz) 

bipartitum Casey 

L. collare (Erichson) 

L. coloraden,;e Casey 

L. impressinLum Hatch 

L. longiuscu um (Gravenhorst)f 

L. multipunc,tum (Gravenhorst)t 

L. rubricolle (Casey) 

L. sewardi (Fall) 

(Subgenus PSEUDOLATHRA Casey) 

(Linolathra Casey) 

(Microluthra Casey) 

(.Parala::hra Casey) 

L. ambiguunr (LeConte) 

L. dimidiatum (Say) 

horni Duvivier 

rufoparlitum Fairmaire & Germain 

L. famelicunr (Casey) 

L. lineiformt, (Casey) 

L. oregonense (Casey) 

___sc___g*PQ_fr___ 

AK 

AK 

_BC 

NT BC 

_BC 

-BO _BC 

BC 

MB ON 

-ox 

_ONPQ 

-oN 

PQ 

_PQ 

9*_ 

NS 

Genus PAEI)ERUS Fabricius 

Leucopt.ederus Casey 

N eo p ae t:.le r us Blackwelder 

Paederillus Casey 

P. littorarius Gravenhorst 

pugetensis Casey 

Subtribe Paederina 

NT BC 


NF 

Subtribe Cryptobiina 

Genus HOMAEOTARSUS Hochhuth 

(Subgenus GASTROLOBIUM Casey) 

H. badius (Gravenhorst) 

H. bicolor (Gravenhorst) 

H. carolinus (Erichson) 

(Subgenus HESPEROBIUM Casey) 

H. californicus (LeConte) 

pacificus Casey 

uancout'eri Casey 

H. cinctus (Say) 

latebricola Nordmann 

H. cribratus (LeConte) 

H. pallipes (Gravenhorst) 

grauenhorsti I\trby 

H. sellatus (]-eConte) 

BC 

BC 

AB 

-oNPQ 

-oNPQ 

ON 

oN PQ 

Dr\ 

- rq 

oN PQ 

oN PQ 

NS 

Genus OCHI|HEPHILUM Stephens 

(Subgenus OCHTHEPHILUM s. str.) 

(C ry p to b ium Mannerheim) 

O. fracticorne (Paykull)+ 

brachelytratum Korge 

furcaticcrne Grimmer 

BC ON PQ NS 

116

Staphylinidae 

Subtribe Astenina 

Genus ASTENUS Dejean 

A. americanus (Casey) 

A. binotatus (Say) 

A. brevipennis (Austin) 

A. cinctus (Say) 

A. discopunctatus (Say) 

centralis Ars.stin 

A. linearis (Erichson) 

A. longiusculus (Mannerheim) 

(?') t risig natu s Boheman 

_BCAB 

BC 

AB 

-oN 

-oN 

MB ON 

-oNPQ 

PQ 

NB 

Subfamily STAPFIYLININAE 

Tribe Philonthini 

Genus NEOBISNIUS Ganglbauer 

Key to New World species: Frank (1981) 

N. jucundus (Horn t - BC AB 

N. lathrobioides (Baudi )* 

N. occidentoides Frank AB - MB 

N. senilis tHornt - BC 

N. sobrinus (Erichsont 

arduus (Sharp) 

ocreatus (Hotn) 

simulator Smetana 

N. terminalis ( LeConte) 

adustus (Sharp) 

delicatulus (Sharp) 

N. viliosulus t Stephenslt 

fuluicornis (Notman) 

ON 

ON 

ON 

PQ 

PQ 

PQ 

PQ 

NB 

NS 

I\TF 

Genus ERICHSONIUS Fauvel 

Actobius Fatvel 

Key to North American species: Frank (192b) 

E. alumnus Frank 

E. civicus Frank 

E. fraterculus (Horn) 

E. inutilis tHorn.l 

E. nanus (Horn) 

E. patella {Horn r 

AK - NT 

E. pusio tHorn ) 

E. rosellus Frank 

Genus PHILONTHUS Curtis 

P. aequalis Horn 

P. agilis {Gravenhorsilt - 

nlger Melsheimer 

P. albionicus Mannerheim AK 

P. asper Horn 

P. aurulentus Horn AK 

P. blandus ( Gravenhorst) 

paederinus Sachse 

pulchellus Melsheimer 

P. brunneus (Gravenhorst) 

breuis Melsheimer 

dimidiatus (Say) 

picatus l\ttby 

P. bucephalus Horn 

P. carbonarius (Gravenhorst)t AK 

uorlzs (GyllenhaI) 

P. caurinus Horn 

P. cephalicus Casey 

P. cephalotes (Gravenhorst)t - 

P. chlorinus Tottenham 

uiridanus Hotn 

P. cognatus Stephensf 

fu sci p enni s (Mannerheim) 

BC 

Ilrr 

BC 

BC 

BC 

BC 

BC 

BC 

_AB 

BC 

BC 

BC 

_ONPQ 

-oN 

-oNPQ 

-oNPQ _ONPQ 

-oNPQ *oNPQ 

-oNPQ 

ON 

- l,ff - 

_ yuoN 

oN PQ 

Of,l 

ON 

PQ 

PQ 

PQ 

NB 

PQ-NS 

NB 

NS 

NS 

NS 

NS 

NS 

NF 

NF 

NF 

NF 

NF 

r17

Staphylinidar,r 

P. concinnus t Gravenhorst)f 

oc hro pus ( Gravenhorst) 

P. confertoides Hatch 

P. confertus f,eOonte 

P. couleensis Hatch 

P. crestonensrs Hatch 

P. crotchi Horn 

P. cruentatus (Gmelin)t 

P. cunctans F.orn 

P. cyanipennis (Fabricius)* 

coe r ule i7:, ennis (Mannerheim) 

P. debilis (Gravenhorst)t 

P. dimidiatus C.R. Sahlbergf 

P. distans Horn 

P. duplicatus Bernhauer & Schubert 

basalis LIotn 

P. ferreipenn:s Horn 

P. fi metarius (Gravenhorst)t 

P. finitimus Cjasey 

P. flavibasis Oasey 

P. forcipatus .Flatch 

P. fraternus Fall 

P. furvus Nordmann 

ca li forn i : us Mannerheim 

comptus Haldeman 

P. fusiformis Melsheimer 

P. gaudens Tlrttenham 

P. grandicollir Horn 

P. hepaticus ltrrichson 

cinctulu:: Melsheimer 

(?) nanu;; Melsheimer 

uaricolor Boheman 

P. hudsonicurr Horn 

P. hyperboreus alaskensis Smetana 

P. instabilis }Iorn 

P. lacustrinuir Scheerpeltz 


P. Iecontei Hr.rrn 

P. leechensis Hatch 

P. lindrothi Smetana 

P. lomatus Erichson 

georgianus Sachse 

P. Iongicornis Stephenst 

P. micans (Glavenhorst)t 

P. neonatus Simetana 

P. nigritoides Smetana 

P. occidentalis Horn 

P. opacipennis Notman 

P. palliatus (Gravenhorst) 

P. perforatus Fauvel 

P. perversus Horn 

P. pettiti Horn 

P. picicornis lJorn 

P. politus (Lirrn6)t 

horrisi Melsheimer 

mandibularis Kirby 

P. protervus Casey 

P. puberulus Horn 

P. pubes Hor:r 

P. pumilio Ct;.sey 

P. quadricollis Horn 

P. quediinus Horn 

P. rainieri Hatch 

P. rectangulus Sharpt 

tetr ag onoce p halus Notman 

P. sanguinolentus (Gravenhorst)t 

P. septentrionis Fall 

P. sericinus [Iorn 

P. siegwaldii Mannerheim 

P. sordidus (Gravenhorst)t 

long i pen nis Provancher 

pachycephalus Nordman 

AK YK 

AK 

AK_NT 

_YK 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

_BC_ 

---ec- _BC 

-YK 

NT BC 

AK 

AK 

AK_ 

AK_ 

_YK 

AK 

NT 


AB 

AB 

AB 

AB 

_MB 

_ 

IYID 

ecABi*Ms-- 

_ABSKMBONPQ 

BC 

BC 

-:"y 

-sc- 

_BC 

_BC 

_BCAB 

BC_ONPQ 

BC 

BC 

BC AB 

-AB 

BC 

BC 

BCi 

BC AB 

SK MB 

-MB 

_MB 

SK MB 

SK MB 

ON 

ON 

ON 

ON 

PQ 

PQ 

PQ 

PQ 

pe 

PQ 

oN PQ 

_Fe 

_PQ 

ON PQ 

-"e 

ON PQ 

ON PQ 

NB 

NS 

PE 

NS__Nr 

NS 

NS 

NB NS 

-NS 

_NS 

-NS 

-NS 

_NS 

we Ns 

NB NS 

-LB 

-NF 

ie 

_LBNF 

NF 

NF 

NF 

NF 

NT 

v 

NF 

NF 

NF 

NF 

IU 

NF 

fu 

NI' 

NF 

\TET 

118

Staphylinidae 

P. spiniformis Hatch 

P. stacesmithi Hatch 

P. stictus Hausen 

P. subvirescens C.G. Thomson* 

P. theveneti Horn 

P. triangulum Horn 

P. umbratilis (Gravenhorst)t 

P. umbrinus (Gravenhorst) 

P. varians (Paykull)f 

P. vulgatus Casey 

_BC 

_BC 

AI( YK NT BC AB 

AK 

BC 

_BC 

-BC 

NT BC 

AK 

ON 

PQ 

PQ 

PQ 

NB 

NB 

NS 

NS 

NS 

NF 

Genus GABRONTHUS Tottenham 

G. thermarum (Aub6)t 

oN PQ 

Genus GABRIUS Stephens 

G. bidentatus (Horn) 

G. brevipennis (Horn) 

G. decipiens (Horn) 

G. downiei (Hatch) 

G. linearis (Casey) 

G. malkini (Hatch) 

G. microphthalmus (Horn) 

G. nigritulus (Gravenhorst)t 

pumilus Mannerheim 

G. oceanus (Hatch) 

G. ottawensis (Casey) 

G. picipennis (Maklin) 

G. seattlensis (Hatch) 

G. subnigritulus (Reitter)+ 

G. virilis (Horn) 

G. wawawai (Hatch) 

AK_N' 

AK__ 

AK YK 

NT 

BC AB 

BC 

BC 

BC 

BC 

BC AB 

BC 

BC AB 

BC 

BC 

BC 

BC 

SK MB 

- lle 

ON PQ 

oN pe 

ON 

SK MB ON PQ 

_PQ 

_-__ 

NB NS 

NE NS 

T\TF 

NF 

-inNr NF 

NF 

Genus CAFIUS Stephens 

(Subgenus EUREMUS Bierig) 

C. bistriatus Erichson 

bilineatus Erichson 

rufifrons Bierig 

C. luteipennis Horn 

BC 

NB 

NF 

(Subgenus BRYONOMUS Casey) 

C. canescens Miiklin 

C. seminitens Horn 

AK 

_BC 

_BC 

(Subgenus REMUS Holme) 

C. femoralis Meklin 

mutatus Gemminger & Harold 

AK 

BC 

(Subgenus PSEUDOREMUS Koch) 

C. nudusjohnsoni Fall 

BC 

Genus BELONUCHUS Nordmann 

B. rufrpennis (Fabricius) 

for mos us ( Gravenhorst) 

pallipes Melsheimer 

ON 

PQ 

Genus LAETULONTHUS Moore & Legrer 

L. Iaetulus (Say) 

Genus HESPERUS Fauvel 

H. apicialis (Say) 

haernaturus (Erichson) 

oN PQ 

ON PQ 

119

Staphylinide:e 

Tribe Staphylinini 

Genus CREOPHILUS Samouelle 

Saprophilus Streubel 

C. maxillosuii (Linn6)t 

arcticus Erichson 

uillosus (Gravenhorst) 

AK YK NT BC AB SK MB ON PQ NB NS 

PE 

LB 

NF 

Genus ONTIIOLESTES Ganglbauer 

O. cingulatus (Gravenhorst) 

O. murinus (Linn6)t 


NB NS 

_LBNF 

NF 

Genus HADIIOTES Meklin 

H. crassus (I{annerheim) 

extensu! LeConte 

AK 

BC 

Genus THINOPINUS LeConte 

Tri c hon r n t h us Motschulsky 

T. variegatus (Motschulsky) 

BC 

Genus PLATYDRACUS C.G. Thomson 

P. cinnamopterus (Gravenhorst) 

P. comes (Letlonte) 

P. exulans (Lrichson) 

P. fossator ((|ravenhorst) 

immaculatus (Horn) 

P. maculosui (Gravenhorst) 

ery thr otr ennis (Mannerheim) 

uiduatu r (Fabricius) 

P. mysticus ( 0richson) 

P. rutilicaudrr (Horn) 

P. tarsalis (N lannerheim) 

sub meta: llicus (Mannerheim) 

P. tomentosus (Gravenhorst) 

P. violaceus ( Gravenhorst) 

cupriper',nis (Melsheimer) 

P. viridanus Horn) 

P. vulpinus ( \ordmann) 

AK 

_BC 

_BC 

-oNPQ 

? 

-oN 

MB ON PQ NB 

ON 

PQ 

-ONPQ-____ 

ON 

ON PQ 

ON PQ 

ON PQ 

NS 

NS 

Genus STAP.FIYLINUS Linn6 

S. aeneocephllus DeGeerf 

S. ater Gravenhorstt 

S. badipes LcConte 

carbona',us LeConte 

S. capitatus 131and 

S. erythropterus Linn6f 

S. melanarius Heert 

S. nigrellus Ilorn 

S. pleuralis L,eConte 

S. saphyrinus LeConte 

S. winkleri Bernhauert 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

^B ix 

AB 

v- 

Io*pe 

MB ON PQ 

_ONPQ 

MB ON PQ 

_ONPQ 

NB 

NB 

NS-- 

NS 

NS 

NT 

NF 

Genus TYMF'ANOPHORUS Nordmann 

Diaphoetes Waterhouse 

T. puncticollis (Erichson) 

borealis (Hatch) 

Tlibe Xanthopygini 

BC 

AB 

oN PQ NB NF 

Genus BEERIA Hatch 

B. nematocera (Casey) 

Tribe Quediini 

Key to North American species: Smetana (1971) 

BC 

t20

Staphylinidae 

Genus HETEROTHOPS Stephens 

H. asperatus Smetana 

H. carbonatus Fall 

H. conformis Smetana 

H. fraternus Smetana 

H. fusculus LeConte 

californicus LeConte 

fumigatus LeConte 

H. marmotae Smetana 

H. minor Smetana 

H. pusio LeConte 

H. sordidus Smetana 

AB SK MB ON PQ _NS 

N,ecAB-tu3NPe-ns--wr 

_BC_ONPQ 

AK_ NT BC 

Genus QUEDIUS Stephens 

(Subgenus MEGAQUEDIUS Casey) 

BC 

AB -ll"- 

Q. campbelli Smetana 

SK oN PQ 

Q. canadensis (Casey) 

oN PQ Ne- 

Q. caseyi caseyi Scheerpeltz --Nt- 

AB SK MB oN PQ 

curtipennis (Casey) 

Q. caseyi divergens (Casey) 

_BC AB 

paruipennis Bernhauer 

uteanus (Casey) 

Q. charactus Hatch 

-BC 

Q. crescenti Hatch 

_BC 

Q. criddlei (Casey) 


hoodiHatch 

Q. erythrogaster Mannerheim 

_BC AB SK ON PQ 

neotomae Hatc}' 

Q. fulgidus (Fabricius)t 

_BC _MB 

iracundus (Say) 

Q. griffinae Hatch 

BC 

Q. incisus Hatch 

BC AB 

Q. Iimbifer Horn 

BC 

Q. mesomelinus (Marsham)t A"- 

BC AB oN PQ _NS 

Q. mordax Smetana 

_YK 

BC AB -y" 

Q. pellax Smetana 

BC 

Q. peregrinus (Gravenhorst) 

ON PQ -NS 

v_ 

siluicola Casey 

Q. pullmani Hatch 

BC AB SK MB 

Q. rainieri Hatch 

_ BC 

Q. spelaeus spelaeus Horn 

BC AB SK 

_NS 

quadriceps Casey 

Q. tahomae Hatch 

BC 

Q. terminatus Melsheimer 

Q. velox Smetana 

AK 

BC AB 

(Subgenus QUEDIUS s. str.) 

Q. curtipennis Bernhauert 

_BC 

parallelus Hatch 

Q. horni Hatch 

_BC 

Q. labradorensis insularis Smetana AK. 

Q. labradorensis labradorensis 

AK NT BC AB SK MB ON 

Smetana 

Q. lanei Hatch 

_BC 

Q. laticollis (Gravenhorst) 

ON 

marginatus Korge 

neomolochinus Korge 

Q. molochinus (Gravenhorst)t 

_PQ 

nigribasis Kotge 

Q. molochinoides molochinoides AKYK_BC 

oN PQ 

Smetana 

(Subgenus QUEDIONUCHUS Sharp) 

Q. plagiatus Mannerheimx 

Q. explanatus LeConte 

Q. manitobensis (Casey) 

(Subgenus MICROSAURUS Stephens) 

(Anastictodera Casey) 

(Quediochrus Casey) 

_BC 

_BC 

_BC 

_BC 

_BC 

pe---ie 

AI{ YK NT BC AB MB ON PQ LB 

LB 

NF 

NF 

NF 

NF 

NF 

r2l

Staphyiinida: 

laeuigati ts (Gyllenhal) 

I ongipen n is Mannerheim 

ruftpennis Maklin 

(Subgenus Dl STICHALIUS Casey) 

Q. capucinus (Gravenhorst) 

ater (Zie1let) 

bordzs Melsheimer 

inuersus (.Say) 

seeuersi Korge 

Q. frater Smetana 

Q. montivagus Smetana 

Q. nevadensis (Casey) 

Q. pediculus (Nordmann) 

Q. tetricus Smetana 

Q. transparerLs Motschulsky 

marginalis Mdklin 

pugetensis Hatc}l 

_YK 

AK 

AK 

_BCAB 

_BC 

_BCAB 

_BCAB 

_BC 

_BCAB 

ON PQ 

(Subgenus RI\PHIRUS Stephens) 

(.Quediel'.us Casey) 

Q. adnexus Snetana 

Q. aenescens Maklin 

agnatus iCasey) 

Q. breviceps (Casey) 

Q. brunnipen nis Mannerheim 

Q. densiventris (Casey) 

alutaceu; Hatch 

bonapari:ei }{atc}i' 

Q. elevatus Hatch 

Q. felimani (Zetterstedt) 

hyperboreus Erichson 

Q. ftigidus Srretana 

Q. fulvicollis r Stephens)* 

pugetant.rs (Casey) 

raphiroides Hatch 

Q. impar Smctana 

Q. nanulus (C asey) 

helenae ( Casey) 

humilis rtCasey) 

Q. oculeus (Crrsey) 

sparsus (Casey) 

Q. prostans Horn 

rupimonlis (Casey) 

Q. rusticus Snetana 

Q. seriatus H,rrn 

Q. simulator iJmetana 

Q. sublimbatr.rs Maklinx 

orbiceps ICasey) 

solitarius (Casey) 

Q. vilis SmetzLna 

(Subgenus P/.RAQUEDIUS Casey) 

AK 

BC 

AB 

AK 

BC 

AKYKNTBCABONPQ_LBNF 

AKYK-BCAB-PQNF 

_BC 


AK-NT_AB 

AKYK_BCAB 

AK 

BC AB 

_BCAB 

BC 

BC 

_BCAB_MB_PQ 

_BC 

AKYKNT_ABSKMBONPQ 

AKYKNTBCABSKMB_PQ 

BC 

SK MB PQ 

ON 

_MBONPQ_LB 

v_ 

_NF 

LB 

- in- 

_LBNF 

NF 

NF 

NF 

Q. puncticeps Horn 

BC 

Genus HEMIiIUEDIUS Horn 

H. ferox (LeC,rnte) 

ON 

PQ 

NS 

Genus ANAQIIEDIUS Casey 

A. vernix (LeConte) 

ON 

PQ 

NB 

NS 

Genus ACYL()PHORUS Nordmann 

(Subgenus AI{ACYLOPHORUS Smetana) 

A. pratensis l,eConte 

(Subgenus ACYLOPHORUS s. str.) 

A. alternans Smetana 

A. caseyi Leni; 

A. flavicollis Sachse 

A. pronus Eri,chson 

AK 

NT BC 

ON PQ 

-oN 

-oNPQ 

-oN 

MB ON PQ 

_NS 

NB NS 

NF 

NF 

r22

Staphylinidae 

Subfamily XANTHOLININAE 


Tribe Diochini 

Genus DIOCHUS Erichson 

D. schaumi Kraatz 


pallidiceps Casey 


_ONPQ 

Genus ATRECUS Jacquelin du Val 

T?ibe Othiini 

A. americanus (Casey) - ON PQ - NS 

A. macrocephalus (Nordmann) AK - NT BC AB ON PQ NB NS NF 

A. quadripennis (Caseyt - BC AB 

Genus PAROTHIUS Casey 

P. californicus (Mannerheim.t - BC 

P. punctatus Smetana - BC 

Genus GAUROPTERUS C.G. Thomson 

Tribe Xantholinini 

G. fulgidus {Fabricius)* - ON 

Genus NIIDOBIUS C.G. Thomson 

N. cephalus (Say) 

assimilis (Kirby) 

AK - NT BC AB SK MB ON PQ NB NS NF 

consentane us (Nordmann) 

flauipennis (Nordmann) 

giluipennis (Casey) 

hiesenwetteri (Sachse) 

nubipennis Casey 

p alliat us (Melsheimer) 

N. pugetanus Casey 

corticallis Casey 

- BC 

debilis Casey 

limbalis Casey 

Genus LEPTACINUS Erichson 

Leptacinodes Casey 

Xa nt ho p hius Motschul sky 

L. batychrus tGyllenhal)+ - BC MB ON PQ 

seriatus LeConte 

L. intermedius Donisthorpet - BC AB SK MB ON PQ NF 

L. pusillus (Stephens)f - ON PQ 

nigritulus LeConte 

Genus PHACOPHAILUS Coiffait 

P. parumpunctatus tGyllenhal)* - ON PQ 

cephalicus (LeConte) 

Genus STENISTODERUS Jacquelin du Val 

S. rubripennis tleConte) - ON 

Genus LITHOCIIARODES Sharo 

Nematolinus Casey 

Oligolinus Casey 

L. longicollis (LeContet - ON PQ 

Genus LINOHESPERUS Smetana 

L. borealis (Casey) - BC 

L. cuspifer Smetana - BC AB 

L. subhamaLus Smetana AB 

t23

Scaphidiidae 

Genus STIC'IOLINUS Casey 

S. flavipes (leConte) 

scolopaceus Casey 

oN PQ 

NS 

Genus HESF;EROLINUS Casey 

Leiolinus Casey 

H. parcus (LeConte) 

tarsalis (Casey) 

uteonus (Casey) 

Genus GYRCTIIYPNUS Samouelle 

G. angustatus Stephenst 

G. campbelli Smetana 

G. fracticorn:.s (O.F. Mulier)t 

obsidiar us (Melsheimer) 

_BC_ONPQ 

MB ON PQ 

_BCAB_MBONPQ 

NB NS 

NB NS 

NF 

NF' 

Genus NEOI|YPNUS Coiffait & Saiz 

N. beckeri Srretana 

N. emmesus,'Gravenhorst) 

luteiuen; ris (Casey) 

sang uinolent us (Melsheimer) 

N. fragilis (C,rsey) 

N. hamatus ( Say) 

macilenius (Casey) 

N. lineatus (lVendeler) 

linearis Blatchley) 

N. melanops .Casey) 

N. monstrosr- s Smetana 

N. obscurus (Orichson) 

oregonul: (Casey) 

protractils (Casey) 

N. picipennis (LeConte) 

---ecABix 

_BC 

NT BC AB SK 

_BC 

IYI D 

MB 

ON PQ 

oN PQ 

ON 

ON PQ 

ON 

oN PQ 

ON PQ 

oN PQ 

NS 

N" NS 

NF 

Genus OXYBLEPTES Smetana 

O. kiteieyi Slretana 

ON 

PQ NB 

Genus XANTHOLINUS Dejean 

X. linearis (Olivier)t 

R11 

NS 

Genus HYPNOGYRA Casey 

H. gularis (Lr,rConte) 

uernicaht Casey 

Genus XEST()LINUS Casey 

X. abdominalis Casey 

frontalis Hatc}: 

indianeri sis Blatchley 

ouiceps Casey 

ON 

SK MB ON PQ 

Family SCAPHIDIIDAE 

shining fungus beetles 

J.M. Campbell 

Scaphidiidae is a small family of fungus-feeding 

beetles that comprises over 1200 species around the 

world, but; only about 50 species occur in North 

America: cif these. 17 are recorded in Canada and 

Alaska. IVIost species are found in fungi, but some 

live in rottrng wood, leaf litter, and under bark. 

Althorrgh many recent authors (Lawrence 

1982a; Lawrence and Newton 1982) have placed the 

scaphidiids in the Staphylinidae family, there is no 

consensus among researchers on their proper placement. 

The family Scaphidiidae is poorly known in 

North America, and species recognition of most 

Canadian taxa is not possible. Casey (1893, 1900) 

has completed the most recent revision of the North 

American shining fungus beetles. 

aK (1); YK (1); NT (1); BC (4); AB (1); SK (2); MB 

(3); oN (13);PQ (10);NF (1) 

t24

Pselaphidae 

Tribe Scaphidiini 

Genus SCAPHIUM Kirby 

S. castanipes Kirby 

AK 

YK 

NT 

BC AB 

MB ON PQ 

Genus SCAPHIDIUM Olivrer 

S. obliteratum LeConte 

S. piceum Melsheimer 

S. quadriguttatum Say 

q uadr inota t u m Castelnau 

quadripustulatum Say 

-oN 

-oN 

_ONPQ 

T?ibe Scaphisomini 

Genus SCAPHISOMA Leach 

S. castaneum Motschulsky 

S. convexum Say 

S. lacustris Casey 

S. repandum Casey 

S. rubens Casey 

Genus TOXIDIUM LeOonte 

T. compressum Zimmermann 

T. gammaroides LeConte 

| -:'- - yuBN 

_i 

oN 

ON 

ON 

pe_ 

PQ 

pe- 

Genus BAEOCERA Erichson 

Eubaeocera Cornell 

Key to Norbh American species: Cornell (1967) 

B. apicalis LeConte 

distincta (Blatchley) 

B. congenera Casey 

speculifera Casey 

B. deflexa Casey 

B. lalsata Achard 

concolor auct. 

youngi (Corr]ell) 

B. humeralis FaIl 

B. nana Casey 

rubriuentris Casey 

__BC___ 

BC 

SK 

-oN 

MB ON 

PQ 

PQ 

D'\ 

- rq 

SK_ONPQ 

oN Fe 

NF 

Family PSELAPHIDAE 

short-winged mold beetles 

A. Dauies 

Eighty-frve species of Pselaphidae are recorded 

in Canada and Alaska: however. the members of 

this family have been poorly collected because of 

their minute size and the obscurity of their habitats. 

They are found chiefly in moist habitats, 

including the edges of bogs and marshes, under 

bark of dead trees, and especially in leaf litter and 

rotten wood; a few are myrmecophilous. Adults of 

most Pselaphidae are predatory on small invertebrates 

such as nematodes, mites, and collembolans; 

the larvae are seldom seen, and their habits are 

equally unknown. 

Jeannel (1955) divided Pselaphidae into six subfamilies, 

all of which are represented in Canada. 

J.L. LeConte, E. Brendel, and T.L. Casey described 

most North American species in the mid to late 

nineteenth century; their work was subsequently 

brought together by Bowman (1934). The present 

arrangement of the supraspecific taxa is based on 

the recent world catalog of genera by Newton and 

Chandler (1989). Chandler (1990o) also published a 

key to the North American genera. 

AK (10); YK (1); NT (3); BC (23); AB (16); SK (9); 

MB (16); ON (56); PQ (38);NB (15);NS (11); pE (2); 

LB (1);NF (3) ' l(2) 

125

Pselaphidae 

Genus MEGARAFONUS Casey 

Key to Ncrrth American species: Schuster and Marsh (1958) 

(Subgenus IIEGARAI'ONUS s. str.) 

Subfamilv FARONINAE 

Tribe Faronini 

M. ventralis Casey AK BC 

Genus SONTfMA Casey 

Rafonus Casey 

Key to North American species: Marsh and Schuster (1962) 

S. margemina Park & Wagner AK BC 

S. parviceps (Maklin) AK BC 

Subfamilv EUPLECTINAE 

Trib; Tlogastrini 

Genus CON,CPLECTUS Brendel 

Hexirherius Grigarick & Schuster 

Prorhet:ius Raffray 

C. canaliculttus (LeConte) 

syluatit.us (Raffray) 

PQ 

Genus ORO ?US Casey 

Key to Nc'rth American species: Schuster & Grigarick (1960) 

O. striatus Oasey 

Genus TETIIASCAPIIA Schuster & Marsh 

T. dentata Park & Wagner 

Genus FOVIIOSCAPHA Park & Wagner 

F. terracola Park & Wagner 

BC 

BC 

BC 

Tlibe Euplectini 

Genus THE|IIUM Casey 

Apothinus Shatp 

T. cavifrons (LeConte) 

ON 

Genus EUP],ECTUS Leach 


Wagner (1975) 

E. acomanus Casey 

E. confluens LeConte 

E. duryi Casey 

barri P;trk & Wagner 

E. elongatus Brendel 

E. karsteni (Reichenbach)t 

iowensis Casey 

E. longicollis Casey 

E. signatus tReichenbach)t 

E. silvicolus Chandler 

--s^- 

SK MB 

ON 

ON 

ON 

ON 

PQ 

PQ 

PQ NB NS 

PQ 

pa:) 

NS 

PE 

Genus PYCI{OPLECTUS Casey 

Key to Ncrrth American species: 

Wagner (1975) 

P. congener (Casey) 

tenellus Casey 

P. interruptrrs (LeConte) 

hudsonicus (Casey) 

P. linearis (LeOonte) 

irnpressiceps (Casey) 

ON 

ON 

ON 

Genus RAMECIA Casey 

Liniolis Grigarick & Schuster 

R. crinitus (tsrendel) 

PQ 

NS 

126

Pselaphidae 

Genus TRIMIOPLECTUS Brendel 

Key to North American species: Chandler (1990b) 

T. obsoletus Brendel - ON PQ 

Genus BIBLOPLECTUS Reitter 

Key to North American species: Chandler (19906) 

B. integer (LeConte) MB ON PQ NB NS 

B. ruficeps tleContet - ON 

Genus ACTIUM Casey 

Proplectus Raffray 

Key to North American species: Grigarick and Schuster ( 1971) 

A. barri Park & Wagner - BC AB 

A. retractum Casey AK BC 

hatchiPatk & Wagner 

A. wawonaensis Grigarick & Schuster - BC 

Genus ACTIASTES Casey 

Key to North American species: Grigarick and Schuster (1971) 

A. foveicollis (LeConte.l - BC - ON PQ 

costalis (Brendel) 

pennsy lu anicus (Bowman) 

A. globiferum (LeConte) PQ 

blandurn (Casey) 

Genus ACTIZONA Chandler 

A. borealis Chandler - BC AB 

Genus CUPILA Casey 

Key to North American species: Grigarick and Schuster (1968) 

(Subgenus CUPILA s. str.) 

C. clavicornis (Maklinl AK BC 

Genus MELBA Casey 

Key to eastern North American species: Chandler ( 1990b) 

(Subgenus MELBA s. str.) 

M. parvr:la tl,eConte) - ON 

Subfamily BATRISINAE 

Tlibe Batrisini 

Genus BATRISODES Reitter 

Key to western North American species: Grigarick and Schuster (1962) 

Key to eastern North American species: Park (1947) 

(Subgenus BABNORMODES Park) 

B. riparius (Sayt - ON PQ 

(Subgenus EMPINODES Park) 

B. albionicus (Aub6) AK BC 

B. denticauda (Caseyt - BC 

(Subgenus EXCAVODES Park) 

B. frontalis tl,eConte) AB - MB ON PQ 

B. globosus (LeConte) AB ON PQ NB NS 

B. scabriceps (LeConte) - ON PQ 

harringtoni (Casey) 

B. sinuatifrons (Brendelt - ON 

(Subgenus PUBIMODES Park) 

B. striatus tl,eConte) 

aterrimus (Casey) 

- ON PQ 

cephalotes (Casey) 

psotai Park 

sirnplex (LeConte) 

I27

Pselaphidae 

Genus MAC HAXRODES Brendel 

M. carinatus (Brendel) 

Subfamily GONIACERINAE 

Tribe Bythinini 

ON 

Genus TYCI{OBYTHINUS Ganslbauer 

Bythinopsis Raffray 

T. bythinioides (Brendel) 

ON PQ 

NS 

Genus RYB,I\XIS Saulcy 

Bryaxis Leach 

R. clavata tlirendel) 

brendeii (LeConte) 

R. conjunctr tLeContet 

R. mystica ( asey 

R. transversa Fall 

R. varicornir, t Brendel) 

AK - NT 

Genus BRAI.)HYGLUTA Thomson 

Nisa Ci,Lsey 

B. arguta (Casey) 

B. terebrata (Casey, 

intricatt (Casey) 

perfora o (Brendel) 

Tlibe Brachyglutini 

-AB 

AB 

SK 

ON 

PQ 

ON PQ 

-MBONPQ 

SK MB ON PQ 

ON PQ 

ON 

ON PQ 

NB 

NB NS 

NB NS 

NB 

NB 

NF 

Genus REIC HENBACHIA Leach 

Key to western North American species: Grigarick and Schuster (1967) 

R. albionica Morschulskyr AK 

americt:..na Raffray 

compar (LeConte) 

francistana Casey 

fundata Casey 

R. arcilera C asey 

R. binodiferi,L Casey AK BC AB 

R. borealis Casey AB - MB 

R. canadens.s (Brendel) 

R. congener Brendelr 

R. corporalis Casey 

R. cyiindrana tBrendel r 

R. distans C,rsey 

R. facilis Ca rey 

R. fusLicorni s Casey BC AB 

R. kapones (lrigarick & Schuster - BC AB 

R. peregrinaLor Casey MB 

R. propinqu:. rleconte) MB 

R. rubicundr. (Aubet 

R. scabra (B'endelt 

R. spatulifer Casey - \aK - BC AB SK MB 

Genus NISAXIS Casey 

N. tomentos..r tAub6, 

cincinnLtta Casey 

minuta (Brendel) 

Genus EUTIiICHITES LeOonte 

E. zonatus (llrendel t 

zimmer,nanni LeOonte 

Genus DECTTiRTHRON Brendel 


(Subgenus DECARTHRON s. str.) 

D. abnorme (LeConte) 

D. formiceti il,eConte) 

D. inusitaturn Park 

_ yl"T:"yu 

_PQ 

ONPQNB_PE 

ON 

_PQ 

oN PQ 

ON 

ON 

ON 

oN---- 

ON PQ NB NS 

ON 

ON 

ON PQ NB 

ON 

ON PQ 

ON PQ 

ON 

_PQ 

LB NF 

I28

Pselaphidae 

D. laurenticum Casey 

D. Iongulurn (LeConte) 

D. scarifrcatum Brendel 

D. stigrnosum Brendel 

ON 

oN PQ 

UN 

Tribe Tychini 

Genus LUCIFOTYCHUS Park & Wagner 

Key to western North American species: Park and Wagner (1962) 


L. cogrratus (LeConte) 

L. testaceus (Casey) 

T---BC--nneoNpeNs--- 

NF 

Genus PSELAPHUS Herbst 

(Subgenus PSELAPHUS s. str.) 

P. bellax Casey 

P. erichsoni LeConte 

P. fustifer Casey 

Subfamily PSELAPHINAE 

Tribe Pselaphini 

BC AB SK 

_AB 


-oN 


Tribe Tyrini 

Genus TYRUS Aub6 

Pytna Casey 

T. corticinus (Casey) 

T. humeralis tAub6t 

compdr LeOonte 

NT 

BC 

AB 

nfn OW Pe 

Ne-- 

Genus CEDIUS LeConte 

Key to North American species: Park (1949) 

(Subgenus SINISTROCEDIUS Park) 

C. ziegleri LeConte 

ON 

Tlibe Ceophyllini 

Genus CEOPHYLLUS LeConte 

C. monilis LeConte 

MB ON 

Tlibe Tmesiphorini 

Genus TMESIPHORUS LeConte 

T. costalis LeConte 

ON 

Tribe Ctenistini 

Genus PILOPIUS Casey 

P. piceus (LeConte) 


saginatus Casey 

ON PQ 

Genus ADRANES LeConte 

A. lecontei Brendel 

A. taylori Wickharn 

Subfamily CLAVIGERINAE 

Tribe Adranini 

_ABSK 

BC_SK 

MB ON 

t29

Hydrophilicae 

Superfamily Hydrophiloidea 

FamilyIIYDROPHILIDAE waterscavengerbeetles 

R.E. Roughley 

Hydr'tphilidae forms a large family of mostly 

aquatic t'eetles; 149 species are known within the 

Canadiar' and Alaskan borders. The family can be 

divided into two biologically different groups: one 

aquatic lnd the other terrestrial. The aquatic 

species or:cur in a wide range of habitats, but most 

prefer shallow standing water. The terrestrial 

species (subfamily Sphaeridiinae) live in most 

kinds of lecaying organic matter. Adults of most 

species, both aquatic and terrestrial, are scavengers. 

Smetana (1988) has recently revised 1;he taxonomy 

of the Canadian species, and his contribution 

was used as the basis for this checklist. 

AJ{Q0; YK (20); NT (36); BC (81);AB (65); SK (52); 

MB (6a); oN (e2); PQ (86);NB (38);NS (33); PE (2); 

H (11);I (18) 

LB (5);NF (24) ' 

Subfamily IfYDROC HINAE 

Genus HYjI'ROCHUS Leach 

H. brevitars is Knisch 

H. currani IV.J. Brown 

H. granulatns Blatchley 

ouellet, Leech 

H. neosquarnifer Hellman 

H. pseudoscuamifer D.C. Miller 

H. scabratus Mulsant 

gibbost.ts Melsheimer 

H. squamife r LeConte 

laticollis Notman 

H. subcupreus Randall 

_BCABSX 

_BC 

BC_ 

:" 

BC AB SK 

-oN 

MB ON 

MB ON 

-oN 

MB ON 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB 

Subfamily HELOPHORINAE 

Key to North American species: Smetana ( 19854) 

Genus HEL OPHORUS Fabricius 

(Subgenus ]IELOPHORUS s. str.) 

H. g'randis I lliger* 

(Subgenus ITHOPALELOPHORUS Kuwert) 

H. angustic,rllis d'Orchymont 

H. auricollisi Eschscholtzx 

angust tIus Mannerheim 

consimilis Mannerheim 

inquinttus Mannerheim 

H. browni Angusx 

H. columbia.nus McCorkle 

H. electus d'Orch).rnont 

corri d'Orchymont 

H. frosti Srr.etana 

H. furius Srnetana 

frater ,t\rr:.etana 

[I. inflectus McCorkle 

[I. lacustris LeConte 

lL ledatus rl'Orchymont 

H. leechi M,:Corkle 

H. linearis ..-eConte 

oblitus Shatp 

pallent Casey 

H. linearoioes d'Orchlnnont 

H. lineatus Say 

o b solet e s ulcat us Motschulsky 

uentratis Motschulsky 

H. marginicollis Smetana 

H. nitiduloicles d'Orchymont 

AK YK 

AK 

BC 

YK NT 

_NTBCABSK 

_NT_ABSK 

NT 

BC 

AB 

ON PQ 

SK MB ON PQ 

SK 

MB 

MB 

MB ON 

BC 

YK NT BC AB SK MB ON 

BC .AB 

BC AB SK MB 

_ABON 

_ _ _ :cy:"yuoN 


s^ 

MB ON 

-oN 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

pe 

PQ 

NB NS 

NB NS 

NB 

NU lls 

NB 

NB 

_LBNF 

NF 

NF 

NF 

130

Hydrophilidae 

H. nitidulus LeConte 

gregolis d'Orchymont 

H. oblongus LeConte* AK 

H. orchymonti Smetana 

H. orientalis Motschulskyx 

H. parasplendidus parasplendidus 

Angus 

H. sempervarians Angus AK 

H. splendidus J. Sahlberg* 

(Subgenus GEPHELOPHORUS Sharp) 

H. sibiricus (Motschulsky)x AK 

(Subgenus CYPHELOPHORUS Kuwert) 

H. tuberculatus Gyllenhal* AK 

scaber LeConte 

(Subgenus ORPHELOPHORUS d'Orchyrnont) 

H. arcticus W.J. Brown* 

YK 

YK 

NT 

NT 

BC AB 

YK NT BC AB 

_NTBCAB 

YK NT 

YK NT BC AB 

YK NT 

BC 

SK 

SK 

SK 

SK 

MB 

MB ON PQ 

-oN 

MB ON PQ 

MB 

MB ON 

AB SK MB ON PQ NS 

MB ON 

PQ 

NB 

NB 

NS 

NS 

LB NF 

,, 

Genus BEROSUS Leach 

Enoplurus Hope 

B. fraternus LeConte 

cal ifor nicus Motschulsky 

B. hatchi D.C. Miller 

B. maculosus Mannerheim 

te sselatu s Motschulsky 

B. oregonensis D.C. Miller 

B. peregrinus (Herbst) 

auritus Melsheimer 

imma c uI at u s Zimmermann 

subsignatus LeConte 

B. striatus (Say) 

B. stylifer Horn 

falcatus Shatp 

AK 

Subfamily BEROSINAE 

_BC 

NT BC 

NT 

AB 

AB 

AB 

SK 

SK 

SK 

BC AB SK 

_AB 

MB ON 

MB ON 

ON 

PQ 

PQ 

PQ 

MB ON PQ 

-oN 

Subfamily CHAETARTHRIINAE 

Genus CHA.ETARTHRIA Stephens 

Key to northwestern species: Miller (1974) 

C. atra (LeContet 

C. nigrella (LeConte) - BC 

C. pallida tl-eContet - SK 

nigriceps (LeConte) 

_PQ_NS 

,,2 

Subfamily IIYDROBIINAE 

Tlibe Hydrobiini 

Genus LACCOBIUS Erichson 

Key to Canadian species: Gentili ( 1985) 

L. agilis (RandaII) 

L. borealis Cheary 

L. carri d'Orchymont 

pacificus D.C. Miller 

L. cinereus columbianus D.C. Miller 

L. minutoides d'Orchymont 

L. reflexipenis Cheary 

reflexipenis Malcolm 

L. spangleri Cheary 

spangleri Malcolm 

L. teneralis Cheary 

L. truncatipenis D.C. Miller 

NT BC 

AK YK NT BC 

AK 

YK 

NT 

NT 

BC 

AB_MB 

AB 

AB SK 

AB SK MB 

SK 

NT AB SK 

MB 

ON PQ 

ON 

ON 

-PQ 

ON PQ 

ON PQ 

MB ON PQ 

-oN 

MB ON 

v_ 

NB 

NB 

NB 

NS 

NF 

\TIT 

Genus IfYDROBIUS Leach 

131

Hydrophili

Hydrophilidae 

E. reflexipennis (Zimmermann) PQ NB NS 

angustulus (Casey) 

Genus CYMBIODYTA Bedel 


C. acuminata Fall - YK NT BC AB SK MB ON PQ - NS 

C. blanchardi Horn - ON PQ 

C. chamberlaini Smetana - ON 

C. columbiana Leech AK BC 

C. minima Notman - BC MB ON PQ 

C. pacifica Leech - BC 

hatchi D.C. Millet 

C. semistriata (Zimmermann) - ON PQ NB 

C. toddi Spangler MB ON 

C. vindicata Fall NT BC AB SK MB ON PQ NB NS - LB NF 

Genus HELOCOMBUS Horn 

H. bifidus {LeContet - ON PQ NB 

Genus TROPISTERNUS Solier 

Subfamilv IIYDROPHILINAE 

T. columbianus W.J. Brown - BC AB SK - ON 

T. ellipticus (Leconte) - BC 

ofizis Motschulsky 

concolor Sharp 

T. glaber lHerbst) - ON PQ - NS 

T. lateralis humeralis Motschulsky - BC 

T. lateralis marginatus Motschulsky - BC AB 

binotatus Walker 

limbalis (LeConte) 

T. lateralis nimbatus (Say) AB - MB ON Pe - NS 

T. mixtus (Leconte) MB ON pQ - NS 

T. natalor d'Orchyrnont - ON PQ 

Genus HYDROPHILUS O.F. Miiller 

H. triangularis Say 

subsulcatus LeConte 

- BC AB SK - ON pe 

Genus DIBOLOCELUS Bedel 

D. ovatus (Gemminger & Haro1d) - ON pQ 

oualis (Ziegler) 

Genus HYDROCHARA Berthold 


H. brevipalpis Smetana - ON 

H. obtusata {Say) NT BC AB SK MB ON pQ 

H. simula Hilsenhoff& Tracy - ON 

Subfamily SPHAERIDIINAE 

Tlibe Sphaeridiini 


Genus PHAENONOTUM Sharp 

P. exstriatum (Say) 

dubium S}ratp 

sem ig I obos um (Zimmermann) 

Genus SPHAERIDIUM Fabricius 

S. bipustulatum Fabriciust 

S. lunatum Fabriciust 

S. scarabaeoides (Linn6)t 

crendtum Palisot de Beauvois 

-oN 

_BCABSK_ONPQNBNS 

_BCABSKMBONPQNBNSNF 

_BCABSKMBONPQNBNSNF 

133

Hydrophilidae 

Genus CERCYON Leach 

(Subgenus PROSTERCYON Smetana) 

C. roseni Knlsch 

opacellus Fall 

(Subgenus CERCYON s. str.) 

C. adumbrat us Mannerheim 

unicolor' (Motschulsky) 

C. assecla Slnetana 

C. atricapilh.Ls (Marsham)t 

mundut Melsheimer 

C. cinctus Slretana 

C. connivens Fall 

C. erraticus i]metana 

C. fimbriatus Mannerheim 

C. haemorrhoidalis (Fabricius)t 

basillaris Notman 

nigricol,t.is (Say) 

C. herceus frigidus Smetana 

C. herceus h':rceus Smetana 

C. impressus (Sturm)t 

C. indistinct,rs Horn 

C. lateralis ( Vlarsham)-i 

incrematus Notman 

C. limbatus . vlannerheim 

ful uiper n i s Mannerhei m 

C. litoralis (( iyllenhal)x 

C. luniger Ma.nnerheim 

C. marinus (j.G. Thomson* 

C. matthewsi Smetana 

C. mendax Smetana 

C. nevadanus Knisch 

C. praetextat;us (Say) 

testaceus Blatchley 

C. pygmaeus (Illiger)+ 

apicalis (Say) 

C. quisquilius (Linn6)t 

C. terminatus { Marsham;f 

nozzs llelsheimer 

C. tolfrno Habch 

C. unipunctu tus (Linn6)l 

(Subgenus DICYRTOCERCYON Ganglbauer) 

C. ustulatus ( Preyssler)* 

(Subgenus P.ARACERCYON Seidlitz) 

C. analis tPaykullt* 

falliWi:fterc 

maculalus Melsheimer 

C. minusculus Melsheimer 

Genus TECT OSTERNUM Balfour-Browne 

Genyon Smetana 

T. naviculare (Zimmermann) - 

AX 

AK 

AK YK NT 

Tribe Cercyonini 


BC 

NT BC 

_BC 

_BC 

-BC 

NT 

AK YK NT 

PQ 

PQ 

PQ 

PQ 

PQ 



BC 

_PQNB 

BC AB ON PQ NB NS 

BC 

AB 

AB 

AB 

AB 

SK 

AK 

BC 

AK 1T( NT BC AB SK 

AK_AB 

AK 

NT 

BC 

BC 

AB SK 

AB SK 

BC 

BC AB SK 

BC 

BC 

AB 

SK 

SK 

-oN 

MB ON 

-oN 

MB ON 

-oN 

MB ON 

-oN 

MB ON 

MB ON 

MB ON 

-oN 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

MB ON PQ 

MB ON PQ 

ON PQ 

NB 

NB 

NB 

NB 

NB 

NB 

NS 

NS 

NS PE NF 

NS 

NS 

NS 

NS 

NS 

NB NS 

NB 

__I" 

NT 

NF 

NF 

NF 

NF 

Genus DELI'OSTETHUS Sharo 

Ictinosternum Smetana 

D. columbier.sis ( Hatcht 

,, 

Genus MEGASTERNUM Mulsant 

M. obscurum (Marsham)t 

M. posticaturn (Mannerheim) 

pugetense Halch 

AK 

Tribe Megasternini 

_BC 

BC 

134

Histeridae 

Genus CRYPTOPLEURUM Mulsant 

C. minutum (Fabricius)t 

C. subtile Sharpf 

_BCAB 

_BCAB 

SK 

MB ON PQ 

MB ON PQ 

NB NS 

PE 

-NF 

_NF 

Family GEORYSSIDAE minute mud-loving beetles 

R.E. Roughley 

Georyssidae includes minute beetles, usually 

under 2 mm long; two species are known for the 

Nearctic region, one of which occurs in Canada. 

Adults are found in mud or sand at the edges of 

streams and are often concealed by sand grains or 

mud attached to the body; they are apparently algal 

feeders. 

BC (1);AB (1);PQ (rl 

Genus GEORYSSUS Latreille 

G. ousillus LeConte 

BC 

AB 

PQ 

Family SPHAERITIDAE false clown beetles 

R.E. Roughley 

Sphaeritidae contains only one genus, 

Sphaerites, with two species in the Holarctic 

region; one of them occurs in Canada and Alaska. 

Adults are found in dung and other decaying 

organic material (e.g. 

sap runs. 

eK (1); BC (1);AB (1) 

fungi) and on trees when the 

Genus SPHAERITES Duftschmid 

S. politus Mannerheim AK 

BC 

Family HISTERIDAE hister beetles 

A. Dauies 

In Canada and Alaska 118 species of Histeridae 

have been recorded. The adults and larvae are chiefly 

predators of other insects and their larvae. Most 

species are associated with decaying organic material, 

dung, and carrion; some are found in sandy areas, in 

mammal burrows, under bark, and in bark-beetle 

galleries; a few are myrrnecophilous. Although some 

species are associated with stored products, they most 

likely feed on other insects that occur there. 

LeConte (1845), Marseul (1853-1857, 1860-1862), 

and Horn (1873) have conducted major taxonomic 

treatments of the North American Histeridae. The 

classification used here is based on Arnett (1968), 

whereas the status of the species is based on Mazur 

(1984) and on work by Helava (personal communication). 

AK (1); YK (2); NT (2); BC (57); AB (42); SK (25); 

MB (43); oN (65);PQ (58);NB (13);NS (12);PE (3); 

NF(7) . H(1);I(10) 

Subfamily ABRAEINAE 

T?ibe Abraeini 

Genus ABRAEUS Leach 

A. bolteri LeConte BC 

135

Histeridae 

Genus PLEGADERUS Erichson 

Tribe Plegaderini 

P. comonfor.i Marseul - BC 

uergran.dis Casey 

P. consors Horn - BC 

egenus Casey 

P. fraternus Horn - BC 

P. molestus Casey - BC 

obesus Casey 

P. nitidus Horn - BC 

P. sayi Mars,eul - BC - ON PQ NB NS 

P. setulosus Ross BC 

P. transversus (Say) ON PQ NF 

Genus ACR.ITUS LeConte 

Tribe Acritini 

A. exiguus I lrichson I 

acicula ius (J.E. LeConte) 

- ON PQ 

cribriptnnis Marseul 

elliptictrc Casey 

natchez Marseul 

obliquu s (J.E. LeConte) 

subdep,'essus (Blatchley) 

A. nigricorn s lHoffman r-l' - ON PQ 

Genus AELIITES Horn 

A. politus tleContet - ON PQ 

A. simplex t,l.E. LeConter - ON PQ 

acupict ts (Marseul) 

Genus TERIITRIUS Erichson 

T. american rs J.E. LeConte 

dissonans Marseul 

picipes J.E. LeConte 

T. montanus Horn 

Tlibe Teretriini 

_BC 

-oNPQ 

Genus SAPIIINUS Erichson 

Subfamily SAPRININAE 

S. discoidalir LeConte 

amplus Casey 

- BC 

latiforn,.is Casey 

mayhewi Papp 

S. impressusr J.E. LeConte 

infaustus LeConte 

- BC AB ON 

piceus ., .8. LeConte 

S. lugens Errchson NT BC AB SK MB ON PQ 

co I iforn icus Mannerheim 

consimilis Waiker 

disting uendoides Hatch 

oregonensoides Hatch 

p se udoclete r sus Dahl gren 

sejunctoides }Jatch 

S. oregonensis J.E. LeConte 

distinguendus Marseul 

- BC AB SK MB ON PQ 

lecontecides Hatch 

p s e u do cl i s t ing u e ndu s Hatch 

p se udo s ej unc toide s Hatch 

pseudosejunctzs Hatch 

sejunctus Marseul 

spurcus LeConte 

S. pensylvar-"icus tPaykull) 

profusoi.des Hatch 

MB ON PQ 

136

Histeridae 

S. sedakovii Motschulsky* 

YK 

Genus EUSPILOTUS Lewis 

(Subgenus HESPEROSAPRINUS Wenzel) 

E. assimilis (Paykull) 

extraneus (Illiger) 

sardous (Flach) 

semisulcus (Hatch) 

simulatus (Blatchley) 

E. conformis (J.E. LeOonte) 

ouiformis (Blatchley) 

E. cribrum (Casey) 

carri (Hatch) 

E. insertus (LeConte) 

obductus (LeOonte) 

opacellus (Casey) 

uinctus (LeConte) 

BC 

BC 

AB SK 

AB SK 

MB ON PQ 

-oN 

MB 

NB 

(Subgenus NEOSAPRINUS Bickhardt) 

E. scrupularis (J.E. LeConte) 

BC 

Genus XEROSAPRINUS Wenzel 

X. fimbriatus (LeOonte) 

desertorum (Marseul) 

X. lubricus (LeConte) 

rotundiftons (Marseul) 

NT BC 

_BC 

AB SK 

AB SK 

MB 

Genus GEOMYSAPRINUS Ross 

(Subgenus PRISCOSAPRINUS Wenzel) 

G. castanipennis (Fall) 

G. cheyennensis (Casey) 

G. copei (Horn) 

G. lanei (Hatch & McGrath) 

G. laramiensis (Casey) 

G. obscurus (LeOonte) 

G. obsidianus (Casey) 

moniliatus (Casey) 

AB SK 

_BCAB 

_BCAB 

_BC 

-BCAB 

_ _ T_ 

yu_ 

-oN 

pr]) 

(Subgenus GEOMYSAPRINUS s. str.) 

G. rugosifrons (FaIl) 

MB 

Genus HYPOCACCUS C.G. Thomson 

H. aeneipunctatus (Horn) 

H. bigemmeus (LeOonte) 

H. bigener (J.E. LeConte) 

eriensis (Hatch) 

illinoensis Wolcott 

impunctellus ( Casey) 

lakensis Blatchley 

michiganensis (Hatch) 

ohioensis (Hatch) 

ontarioensis (Hatch) 

sphaeroides (J.E. LeConte) 

H. estriatus (LeConte) 

H. frtchi (Marseul) 

H. fraternus (Say) 

H. iris (Fall) 

alutiger Wenzel 

H. mancus (Say) 

H. omissus (Casey) 

H. patruelis (J.E. LeConte) 

jaueti (Marceul) 

H. seminitens (LeConte) 

-BC 

- - - :'T 3* MBor.pe 

BC 

BC 

AB 

PQ 



MB 

AB SK MB 

AB_MBONPQ 

MB ON PQ 

NS: 

NS 

PE 

NF 

Genus BAECKMANNIOLUS Reichardt 

B. dimidiatipennis (J.E. LeConte) 

deserticola (Marseul) 

desertorum (Marseul) 

palmatus (Say) 

PQ NB NS 

NF 

L37

Histeridae 

Genus NEOI'ACHYLOPUS Reichardt 

N. sulcilrons tMannerheim.l - BC 

Genus GNA'IIHONCUS Jacquelin du Val 

G. rossi Hat< h - BC 

G. rotundatus (Kugelann)f - BC AB SK MB ON PQ NF 

communis (Marseul) 

deletus (J.E. LeConte) 

nanus (Scriba) 

Genus DENI )ROPHILUS Leach 

Subfamilv DENDROPHILINAE 

Tri6e Dendrophilini 

D. punctatus (Herbsu+ - BC MB ON PQ 

punctulatus (Say) 

D. xavieri M rrseult - BC - ON 

sexstria;lus Hatc}a 

Tlibe Bacaniini 

Genus BACFNIUS LeOonte 

B. punctifon:ris (LeConte) 

_ONPQ 

Genus CARC INOPS Marseul 

Carcinus Marseul 

Tribe Paromalini 

C. pumilio (Etrichson)t 

nanus Ql.E. LeConte) 

- BC AB SK MB ON PQ NF 

q uatuordecimsfrjolzs (Stephens) 

Genus XESTIPYGE Marseul 

X. conjunctu n tSayt - ON 

X. geminaturn tJ.E. LeConte) - ON 

Genus PLATYLOMAIUS Cooman 

P. aequalis (iiayt 

affinis (,1.8. LeConte) 

- BC AB - MB ON PQ 

estriatu.t (J.E. LeConte) 

Genus PARC,MALUS Erichson 

P. bistriatus Erichson AB ON PQ NB - NF 

P. mancus C,lsey - BC AB 

P. seeversi (\Venzel; - ON 

P. teres LeConte - ON PQ NB - NF 

Genus EPIEIRUS Erichson 

Subfamilv TRIBALINAE 

E. pulicarius Erichson - ON PQ 

minor J,8. LeOonte 

E. regularis iPalisot de Beauvois) - ON 

nigrellus (Say) 

uicinus LeConte 

Genus ONTIIOPHILUS Leach 

O. deflectus .l{elava - ON 

Genus CAEF|OSTERNUS LeConte 

C. americanus U.E. LeConte) - ON 

138

Histeridae 

Genus ATHOLUS C.G. Thomson 

Peranus Lewis 

A. americanus {Paykull ) 

diffractus (Casey) 

exard.tus (J.E. LeConte) 

perplexus (LeConte) 

A. bimaculatus ( Linn6)t 

obliquus (.Say) 

A. falli tBickhardtt 

gagates Fatl 

A. sedecimstriatus { Sayl 

Subfamily HISTERINAE 

Tribe Histerini 

AB 

BC AB 

BC AB SK 

SK MB ON PQ 

-oN 

MB ON 

MB ON 

PQ 

PQ 

PQ 

Genus MARGARINOTUS Marseul 

(Subgenus MARGARINOTUS s. str.) 

M. guttifer Horn 

_AB 

(Subgenus PTOMISTER Houlbert & Monnot) 

M. brunneus (Fabricius;f 

cadau e r inu s (Ho{frnann) 

impressus (Fabricius) 

obtusatus (Harris) 

uirginiae (Casey) 

M. egregius {Casey) 

M. foedatus (J.E. LeConte) 

M. fractifrons {Caseyt 

fidelis (Casey) 

planifrons (Lewis) 

M. harrisii (Kirbyt 

repletus (J.E. LeConte) 

M. hudsonicus (Caseyt 

M. immunis (Erichson) 

M. intermptus (Palisot de Beauvois) 

M. merdarius (Hoffmann)t 

memnonius (Say) 

M. pluto (Casey) 

albertensis (Hatch) 

carri (}{atc}r) 

M. rectus (Caseyt 

M. umbrosus (Casey) 

umbratilis (Casey) 

-:nc- 

BC 

BC 

nc 

BC 

BC 

BC 


AB 

-MBONPQ 

_yuoNPQ 

MB 

ON PQ NB 

MB ON PQ 

MB ON PQ 

-oNPQ 

_ONPQ 

NB 

NB 

NB 

NS PE 

NS 

ols--- 

NS 

(Subgenus STENISTER Reichardt) 

M. obscurus (Kugelann)f 

semisculptus (LeConte) 

stercorarius (Hoffmann) 

BC 

(Subgenus PARAIISTER Bickhardt) 

M. cognatus (J.E. LeConte) 

unicus (Casey) 

M. lecontei Wenzel 

M. purpurascens (Herbst)t 

(Subgenus PROMETHISTER Kryzhanovsky) 

M. confusus Wenzel 

M. marginicollis (J.E. LeConte) 

--nc- 

SK 

MB ON PQ 

MB ON PQ 

-oN 

MB ON 

PQ 

PQ 

NB 

NS 

PE 

Genus HISTER Linn6 

H. abbreviatus Fabricius AK 

bifidus Say 


crenicollis Flach 

subhernisphaericus Palisot de Beauvois 

H. albertanus Casey 

H. curtatus J.E. LeConte 

haldemanni Marseul 

BC 

AB 

AB 


PQ 

NS 

139

Histeridae 

H. defectus .f.E. LeConte 

permixlus Zimmermann 

H. depuratol Say 

paykulliKirby 

perbreu.'.s Casey 

spretus J.E. LeConte 

BC 

AB 

SK 

SK 

-oN 

MB ON 

PQ 

PQ 

NB NS 

Genus SPILODISCUS Lewis 

S. arcuatus (Say) 

S. militaris (Horn) 

electus (Casey) 

oregonus (Casey) 

simpliupes (.Fa\l) 

S. sellatus (LeOonte) 

senneui lei (Marseul) 

S. semiruber (Casey) 

soloris t Carnochan) 

-BC 

BC 

BC 

AB 

SK 

MB 

NS 

Genus PSILr)SCELIS Marseul 

P. abnormal s Mann 

P. carri Hatch 

P. corrosa Casey 

P. planipes (LeConte) 

harrisi:J.8. LeConte) 

P. subopaca LeConte) 

YK 

_AB 

_AB 

BC AB 

SK 

MB 

MB 

MB 

ON 

PQ_NS 

Tribe Platysomatini 

Genus PLA1YSOMA Leach 

(Subgenus PLATYSOMA s. str.) 

P. aequum (l,eConte) 

P. aurelianu n (Horn) 

P. lecontei Nlarseul 

d.epress tm J.E. LeConte 

tabellurt Casey 

BC 

AB 

_MBON 

ON 

PQ 

PQ 

PQ 

(Subgenus EURYLISTER Bickhardt) 

P. carolinum (Paykull) 

cinnamomeum (A. White) 

sordidum (Say) 

oN PQ 

(Subgenus CYLISTER Cooman) 

P. coarctaturn J.E. LeConte 

P. punctigemm (LeConte) 

Genus CYLISTIX Marseu. 

-BCAB 

_BCAB 

_ yuoNPQ 

C. cylindrica (Palisot de Beauvois) 

C. deficiens Casey 

C. gracilis (J.E. LeConte) 

_ _ rsxy"giliH _ _ _ _ _ 

Tribe Hololeptini 

Genus HOLOLEPTA Paykull 

H. aequalis Say 

fossularis Say 

BC 

AB 

MB ON PQ 

Tribe Exosternini 

Genus PHEL,ISTER Marseul 

P. subrotunius (Say) 

P. vernus (Say) 

saunieri Marseul 

oN PQ 

ON 

I40

Clambidae 

Subfamily HETAERIINAE 

Tribe Hetaeriini 

Genus HETAERIUS Erichson 

H. brunneipennis (Randall) 

H. exiguus Mannerheim 

nitidus Martin 

H. tristriatus Horn 

BC 

BC 

AB 

-oN 

SK MB 

Superfamily Eucinetoidea 

Family EUCINETIDAE plate-thigh beetles 

J.M. Campbell 

Eucinetidae is a small family that contains 

about 30 species throughout the world; seven of 

them, placed in one genus, are known to occur in 

Canada. These beetles are associated with fungal 

decay and are found in leaf litter or under bark of 

fungus-infested logs. 

Horn (1880) last reviewed the species of this 

family, as part of Dascillidae; they are poorly known 

taxonomically. 

vK (1); NT (2); BC (5); eB (3); SK (3); MB (4); ON 

(a);PQ (a);NB (2);NS (2) . I (1) 

Genus EUCINETUS Germar 

E. haemorrhoidalis (Gerrnar)t 

E. infumatus LeConte 

E. morio LeConte 

E. oviformis LeConte 

E. punctulatus LeConte 

E. terminalis LeConte 

E. testaceus LeConte 

_v 

YK_ 

_NT 

BC AB 

BC 

SC AB 

BC 

BC AB 

SK 

sx 

MB 

MB 

MB 

SK MB 

oN PQ 

ON PQ 

ON PQ 

oN PQ 

NB NS 

_NS 

NB 

Family CLAMBIDAE minute beetles 

J.M. Campbell 

Clambidae forms a family of small beetles that 

are rarely collected because of their size and habitat. 

The family contains five genera and about 50 

species in the world. Only two genera comprising 

seven species are known to occur in Canada and 

Alaska, but more are likely to be found. Adults are 

usually collected during their evening flight. They 

can also be collected by sifting leaflitter and piles of 

decaying vegetation, where they probably feed on 

spores of fungi and slime molds. 

The American species were revised by Endrridy- 

Younga (1981) and are well-known taxonomically. 

AK (1); BC (3); AB (3); oN (1); PQ (6) o I1[ 

(1); I (1) 

Genus CALYPTOMERUS Redtenbacher 

C. oblongulus Mannerheim AK 

BC 

Genus CLAMBUS Fischer von Waldheim 

Sternuchus LeConte 

C. armadillo armadillo (DeGeer)t 

niger Hatch 

C. gibbulus (LeContetf, 

r adul a Endr }dy-Younga 

C. howdeni Endrddy-Younga 

BC 

AB 

ON 

PQ 

PQ 

PQ 

r47

Scirtidae 

C. pubescens Redtenbacher* 

puberu.lus LeConte 

C. smetanai Endrcidy-Younga 

C. vulneratus LeConte 

BC AB PQ 

PQ 

PQ 

Family SCIRTIDAE marsh beetles 

J.M. Campbell 

The lamily Scirtidae contains about 600 species 

throughout the world, but most of them are found in 

temperai;e zones. Six genera and 26 species are 

known tc occur in Canada and Alaska. Adults are 

collected, often in large numbers, on vegetation in 

wet habitats, particularly marshes, bogs, and 

swamps; they are attracted to light. Larvae are 

aquatic and feed on fungi, algae, diatoms, and other 

organic nratter. 

Most Canadian species are well known except 

those of the genus Cyphon, which is in need of revision. 

Horn (1880) last reviewed the species of this 

family, as part of Dascillidae. Klausnitzer (1976), 

however, reviewed several species occurring in 

Canada; he (1980) also segregated some species formerly 

assigned to Elodes to form the new genus 

Flauohelodes. 

AK (1); YK (1); NT (2); BC (10); AB (a); SK (3); MB 

(3); oN (15);PQ (15);NB (7);NS (3);NF (5) 

Genus ELCTDES Latreille 

Hemic;tphon LeConte 

Sacodt:s LeConte 

E. angusta Hatch 

E. apicalis 1,eConte 

E. emargin;r.ta Hatch 

E. maculicc'llis Horn 

_BC 

_BC 

_BC 

PQ_NS 

NF 

Genus FLAVOHELODES Klausnitzer 

F. fuscipennis (Gu6rin-M6neville) 

F. pulchella (Gu6rin-M6neville) 

F. thoracica. (Melsheimer) 

ON 

ON 

ON 

Genus MICROCARA C.G. Thomson 

M. explanat;a LeConte 

BC 

SK MB ON PQ 

NB 

Genus CYPHON Paykull 

C. brevicoll:.s (LeConte) 

C. collaris (rGu6rin-M6neville) 

C. concinnus (LeOonte) 

C. confusus W.J. Brown 

horiont Klausnitzer 

C. exiguus :1orn 

C. modestu'; (LeConte) 

C. nebulosus (LeConte) 

C. neovaria bilis Klausnitzer 

C. obscurus (Gu6rin-M6neville) 

C. ochreatus Klausnitzer 

C. punctatr:.s (LeConte) 

C. pusillus,.LeConte) 

C. rufrcollis (Say) 

C. variabilis (Thunberg) 

AK 

_NT 

YK NT 

BC 

BC 

BC 

BC 

BC 

BC 

AB- - - :a 

_ONPQ 

AB 

AB 

SK 

SK 

--pe 

-oNPQ 

_ONPQ 

_ONPQ 

yu oN PQ 

_ONPQ 

MB ON PQ 

NB 

NSfu 

Ns 

lle Ns 

NF 

NF' 

NF 

NF 

Genus PRIONOCYPHON Redtenbacher 

P. discoideus (Say) 

P. limbatus LeConte 

oN PQ 

oN PQ 

NB 

Genus SCIIITES Illiger 

S. orbiculatus (Fabricius) 

S. tibialis Gu6rin-M6neville 

AB 

oN PQ 

ON PQ 

NB 

t42

Lucanidae 

Superfamily Dascilloidea 

Family DASCILLIDAE soft-bodied plant beetles 

J.M. Campbell 

The status of Dascillidae has changed considerably 

since Horn reviewed it (1880). Its size has 

been greatly reduced by the r_emoval of species now 

included in the families Scirtidae, Eucinetidae, 

Artematopidae, and Ptilodactylidae, so that only 

one species is now known to occur in Canada. 

Fifteen genera and about 80 species are known to 

occur throughout the world. The adults are usually 

found on foliage; they feed on plants or are carnivorous. 

The larvae live in soil or are aquatic; they feed 

on vegetative matter such as roots or leaf mold. 

BC (1) 

Subfamily DASCILLINAE 

T?ibe Cneoglossini 

Genus ARAIOPIDIUS Cockerell 

Araeopus LeConte 

A. monachus (LeConte) 

BC 

Family RHIPICERIDAE cedar beetles 

J.M. Campbell 

Rhipiceridae contains five genera and about 50 

species throughout the world. Only one genus with 

one species is known to occur in southern Ontario. 

The larvae of this species live on the roots of trees, 

where they are parasitic on the nymphs of cicadas. 

The adults are often found in 

on the trunks of trees. with 

around one female. 

oN (1) 

mating congregations 

many males grouped 

Genus SANDAI-US Knoch 

S. niger Knoch 

hnochi (Gt6in-M6neville) 

rubidus Melsheimer 

r uftpennis (Latreille) 

scabricollis Haldeman 

ON 

Superfamily Scarabaeoidea 

Family LUCANIDAE stag beetles 

J. McNq.mara 

Lucanidae includes 14 species in Canada. The 

larvae of all species breed in or beneath the decaying 

wood of logs or stumps and therefore are of little 

economic importance. The adults are believed to 

feed on honeydew or sap. They lay their eggs in 

bark crevices, especially near the roots. The adults 

are attracted to light and are most commonly found 

in wooded areas. 

Howden and Lawrence (1974) published a key 

of the North American genera, and since the genera 

occurring in Canada include few species, specific 

identifrcation is relativelv easv. 

vK (1); NT (2); gC (7); AB (2); SK (2); MB (3); ON 

(8);PQ (5); NB (2);NS (3);NF (1) 

r43

Lucanidae 

Subfamilv SYNDESINAE 

Tribe Sinodendrini 

Genus SINODENDRON Hellwig 

S. rugosum Mannerheim - BC 

Tlibe Ceruchini 

Genus CERUCHUS Macleay 

C. piceus (V/eber) 

bolbi (t.lastelnau) 

uirginiznsis Casey 

C. punctatus LeConte 

C. striatus LeConte 

BC 

_BC 

MBONPQ-NS 

Genus NIC,\GUS LeConte 

Subfamily NICAGINAE 

T?ibe Nicagini 

N. obscurus tleConter ON PQ 

Genus LUC ANUS Scopoli 

(Subgenus LUCANUS s. str.) 

Subfamily LUCANINAE 

Tribe Lucanini 

L. elaphus I'abricius - ON 

carleng:i Angell 

(Subgenus I'SEUDOLUCANUS Hope & Westwood) 

L. capreolus tl.inndt 

domo l:'abricius 

- ON 

muticu.; Thunberg 

nigricephalus (Benesh) 

trigonu:.s Thunberg 

L. placidus iay 

lentus \)astelnat 

- ON 

Genus DORCUS Macleay 

Tribe Dorcini 

D. parallelu* {Sayt 

carnocl,,ani Angell 

- ON PQ 

costatu,s LeConte 

nanus Casey 

oblongus (Charpentier) 

uoeti (Schonherr) 

Genus PLA',YCEROPSIS Benesh 

P. keeni (Casey) BC 

pedicelt aris (Moilenk) 

Genus PLAIIYCERUS Geoffroy 

Key to species: Benesh (1946) 

TYibe Platycerini 

P. marginalis Casey NT BC AB SK MB NB NS 

P. oregonensis Westwood - BC 

chalybo:eus Casey 

coerulescens LeConte 

P. piceus (K,rby) - \'K NT BC AB SK MB ON PQ NB - NF 

depressus LeConte 

144

Scarabaeidae 

virescens (Fabricius) 

angustus casey 

iowanus Casey 

peregrinus Casey 

quercus (Weber) 

securidens (.Say) 

oN PQ 

NS 

Family PASSALIDAE peg and b,ess beetles 

J. McNamara 

Only one species of Passalidae occurs in 

Canada. These beetles, which live in rotten logs 

and stumps, are not economically important. The 

adults and larvae communicate with each other by 

stridulation. The adults prepare food for the larvae 

by chewing the wood and mixing it with saliva. 

Reyes-Castillo (1970) published 

American genera of this family. 

MB (1);ON (1) 

key to the 

Genus ODONTOTAINIUS Kuwert 

Passalotaenius Kuwert 

O. disjunctus (Illiger) 

cornutus (Fabricius) 

Tribe Proculini 

NtB ON 

Family SCARABAEIDAE 

scarErb beetles 

J. McNamara 

Scarabaeidae is one of the largest families of 

beetles in the world; 248 species are known to occur 

in Canada and Alaska. Beetles of this family are 

popular with collectors because of their bright colors 

and the large horns of some groups. The size of 

these beetles varies greatly, from 1.9 mm for 

Pleurophorus longulus to 15 cm for Goliathus, 

which is probably the largest insect known based on 

mass. The scarabs contain some of the most 

destructive beetles, such as the Japanese beetle 

(Popillia japonica), the Asiatic garden beetle 

(Maladera castanea), and the European chafer 

(Amphirnallon majalis). These belong to a group of 

scarabs that feed primarily on Iive plant matter as 

adults, and usually on plant roots or rotting wood as 

larvae. The other group of scarabs are mainly dung 

feeders or scavengers. They feed primarily on dung 

or decaying plant and animal matter as adults, and 

in many cases the larvae feed on the same material 

as it is supplied to them by the adults. Because of 

their economic importance, the life cycle of many 

species is fairly well known. That of the Japanese 

beetle is probably typical of the family in general. 

Mating taker; place during the summer. The eggs 

are laid in the soil in late July and early August; 

they hatch immediately, and the larvae begin to 

feed. When bhe weather turns cold the larvae burrow 

deeper into the soil, below the frost line, where 

they hibernate. In the spring when the weather is 

warmer, they move up near the surface of the 

ground and continue to feed until they pupate in 

June. The adults emerge about the first of July, 

depending on the area. Many variations occur in 

the life cycle of scarabs. The Phyllophoga species 

cycle ranges firom 2 to 4 years, depending largely on 

the latitude. 

John Cooper is currently revising the scarabs of 

Canada and Alaska, and his work is expected to be 

published as part of The Insects and Arachnids of 

Canada, a serries published by Agriculture Canada. 

The classifrcation used here is taken almost entirely 

from his manuscript. 

AK (13); YK (9); NT (17); BC (88); AB (96); SK (58); 

MB (85); ON (136); PQ (109); NB (a1); NS (a7); PE 

(11);LB (7);ltIF (20) . H (1); I (21) 

145

Scarabaeidae 

Genus CAIITHON Hoffmansegg 

Key to North American species: Robinson (1948) 

(Subgenus CANTHON s. str.) 

Subfamilv SCARABAEINAE 

Tribl Scarabaeini 

C. chalcites (Haldemanr ON 

C. pilularir:s {Linn6) - AB 

hudsonias (Forster) 

laeuis i.Dtury) 

obtusi'fens (Ziegler) 

uiridetcens Horn 

uoluen; (Fabricius) 

C. vigilans LeConte - ON 

(Subgenus BOREOCANTHON Halflter) 

C. praticok LeConte BC A-B SK MB 

C. simplex Leconte - BC AB 

(Subgenus ILAPHYROCANTHON Martinez) 

C. viridis 1l)alisot de Beauvois) 

metallicus Stutrn 

- ON 

obsoletus (.Say'1 

uiridicatus (Say) 

Genus COF'RIS O.F. Miiller 

Key to New World species: Matthews (1961) 

Tribe Coprini 

C. lricator lricator rFabricius) - ON PQ 

anagllpticus Say 

tullius (O\ivter) 

C. minutus (Drury) PQ 

ammo:l, Fabricius 

lar Fa.:ricius 

reflexu:.s Panzer 

silenzs Fabricius 

Genus ONTHOPHAGUS Latreille 

Key to Nrrth American species: Howden and Cartwright (1963) 

Tlibe Onthophagini 

O. hecate hecate (Panzer) - BC AB SK MB ON PQ NB NS 

furcicollis Dejean 

hastator (Fabricius) 

lamal)ejean 

latebrosus (Fabricius) 

obtectt:s (Palisot de Beauvois) 

soyi C;rstelnau 

scabricollis l{rtby 

O. nuchicornis (Linn6)t - BC AB - MB ON PQ NB NS NF 

r hinoc'zro s Mel sheimer 

xiphias LeConte 

O. orpheus canadensis lFabriciusr - ON PQ 

O. orpheus pseudorpheus MB 

Hou'den & Cartwright 

O. pennsyl' anicus Harold - ON PQ 

Genus MICRAIGIAJ-IA W.J. Brown 

Subfamily APHODIINAE 

Tribe Aegialiini 

Key to North American species: Gordon and Cartwright (1988) 

M. pusilla (Horn) - BC AB - MB 

I46

Scarabaeidae 

Genus AXGIALIA Latreille 

(Subgenus LEPTAIGIALIA W.J. Brown) 

A. humeralis W.J. Brown 

A. montana W.J. Brown 

A. rufescens Horn 

(Subgenus PSAMMOPORUS Thomson) 

A. criddlei W.J. Brown 

A. cylindrica (Eschscholtz) 

A. lacustris LeConte 

A. nana W.J. Brown 

A. opaca W.J. Brown 

A. terminalis W.J. Brown 

(Subgenus RHYSOTHORAX Bedel) 

(Anomalaeg ial ia W . J . Brown) 

A. spissipes LeContef 

rzlo (Fabricius) 

rufina Sllfvergerg 

(Subgenus AXGIAJ-IA s. str.) 

A. arenaria (Fabricius)t 

A. blanchardi Horn 

A. carri Gordon & Cartwrieht 

A. conferta Horn 

A. crassa LeConte 

insularis W.J. Brown 

A. opifex Horn 

_BCAB 

_BCABSK 

AK BC AB SK 

AK 

BC 


_BCAB 


_BC 

AB 

_ _ _ ecf:" 

-oN 

MB ON 

MB ON 

MB ON 

MB 

NtrD 

PQ 

PQ 

PQ 

oN PQ 

r\I\T 

NB 

PQ NB NS 

PQIU___ 

PQ 

PQ 

oN PQ 

NS 

NS 

NS 

PE 

LB 

NF 

)u 

NF 

NF 

Genus APHODIUS illiger 

Key to eastern North American species: Gordon (1983) 

A. aenictus Cooper & Gor'lon 

A. albertanus W.J. Brown 

A. aleutus Eschscholtz 

ursinus Motschulsky 

A. alternatus Horn 

inermis Robinson 

moclipsiHaLch 

A. anthracinus LeConte 

A. aquilonarius W.J. Brown 

A. badipes Melsheimer 

oblongus Say 

A. bicolor Say 

A. bidentatus Schmidt 

bidens LeConte 

A. borealis Gyllenhalx 

errons W.J. Brown 

A. brevicollis LeConte 

A. browni Hinton 

smithiW.J.Brcwn 

A. bryanti W.J. Brown 

A. carri W.J. Brown 

A. coloradensis Horn 

A. congregatus Mannerherm 

arcticus Harold 

A. consentaneus LeConte 

A. cormptor W.J. Brown 

A. cribratulus Schmidt 

cribratus LeConte 

A. criddlei W.J. Brown 

A. decipiens Horn 

A. denticulatus Haldeman 

A. distinctus (O.F. Miiller)t 

inquinatus (Herbst) 

m a eu I i pe n n i s Mel sheimer 

A. duplex LeConte 

A. edmontonus W.J. Brown 

AK 

AK \'K 

Tribe Aphodiini 

NT BC AB 

_BCAB 

-BC 

NT 

___sc 

_BC 

BC 

_ 

ec 

SK 

MB 

ON 

^AB 

BCTSKIo*pe----- 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

__9* 

_MBON 

_ y"_ 

SK 

SK 

SK MB 

SK MB 

:" yu_ 

SK MB ON 

SK MB 

PQ 

PQ NB NS LB NF 

pe 

NS 

I47

Scarabaeic ae 

A. erraticus (Linn6)t 

pensuullensis Melsheimer 

A. fimetarius (Linn6)-f 

nodifrons Randall 

A. fossor (Linn6)t 

A. granarirLs (Linn6)t 

aterr i;' n u s Melsheimer 

inutilt:s Horn 

metalticus Haldemarr 

spretu.s Haldeman 

A. guttatus Eschscholtz 

A. haemorrhoidalis (Linn6)i 

A. hamatus hamatus Say 

occidentalis Horn 

sagittarius Harold 

A. hirsutus W.J. Brown 

A. hyperboreus LeConte 

anguloris LeConte 

A. iowensis Wickham 

A. lentus Eorn 

A. leopardr.Ls Horn 

A. leptotarr;is W.J. Brown 

A. manitobensis W.J. Brown 

A. omissus omissus LeConte 

A. omissus torpidus Horn 

A. opacus l,eConte 

colum,tiensis Wickham 

A. pardaiis LeConte 

A. pectoral:s LeConte 

A. peculios rs Schmidt 

awemeanus W.J. Brown 

longitursis Fall 

tenuihr,rsis Reitter 

A. phaeopterus LeConte 

canadensis Garnett 

(?) rus;elli Hatch 

A. piceatus Robinson 

rainer', Hatch 

A. pinguell rs W.J. Brown 

A. pinguis1:Ialdeman 

ualidus Hotn 

A. prodromus (Brahm)t 

A. richards'tnae Hatch 

A. rogersi Ilatch 

A. rubeolus (Palisot de Beauvois) 

c op r on.y tn u s Mel sheimer 

A. rubripennis Horn 

A. rufrpes (.Linn6)t 

A. ruricola Melsheimer 

aureliunus Harold 

curtus Haldeman 

A. scabriceps LeConte 

A. scrofa (Fabricius)t 

A. sigmoideus Van Dyke 

A. socialis \V.J. Brown 

A. stercoros,us Melsheimer 

A. subaeneits LeConte 

gentne.i }{atch 

perpoli,tus Gordon 

A. subterraneus (Linn6)t 

A. talpoides,i W.J. Brown 

A. tenellus Say 

A. thomomJrsi W.J. Brown 

A. vittatus Say 

A. walshii }Iorn 

AK 

BC AB 

_BCAB 

AKYK_BCAB 

_BCAB 

_BCAB 

AK 

AK 

YK 

YK 

YK 

NT 

NT 

f' 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

? 

_BC 

_BC 

_:r_ 

-AB 

BC 

_AB 

BC? 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

AK I'K NT BC AB 

_BCAB 

AB 

SK 

MB 

MB 

SK MB 

_MB 

_MB 

SK MB 

SK MB 

SK MB 

SK MB 

SK MB 

_MB 

SK MB 

SK MB 

_MB 

SK MB 

SK MB 

SK MB 

-MB 

SK 

_MB 

-ms 

_MB 

_MB 

SK MB 

SK MB 

ON PQ 

ON PQ 

ON 

ON 

ON 

NB 

NB 

NS 

NS 

LB 

PQ NB NS PE LB 

PQ NB NS 

PQ 

PQ 

PQ 

ON PQ 

ON PQ 

ON PQ 

ON PQ 

ON 

oN pe 

oN PQ 

ON 

oN PQ 

ON PQ 

oN PQ 

-PQ 

oN pe 

ON PQ 

ON PQ 

oN PQ 

_NS 

NB NS 

NB NS 

NN NS 

_NS 

NB NS 

NB NS 

_NS 

NB 

NB NS 

NB NS 

_LB 

_LB 

NF 

NF 

NF 

NF 

NF 

TU 

NF 

NF 

NF 

Genus OXYOMUS Stephens 

O. silvestris (Scopoli)'l 

opacifrons }Iorn 

porcatus (Fabricius) 

BC 

PQ 

I48

Scarabaeidae 

Tlibe Eupariini 

Genus ATAENIUS Harold 

Key to North American species: Cartwright (1974) 

A. abditus {Haldeman.t 

attenuator Harold 

A. gracilis ( Melsheimer t 

A. imbricatus (Melsheimer) 

sordidus Harold 

A. spretulus (Haldemant 

cozsors Fall 

falliHinton 

A. strigatus (Sayt 

ON PQ NB 

ON PQ 

ON 

ON 

oN PQ 

Genus DIAJ-YTES Harold 

D. criddlei W.J. Brown 

D. striatulus tSayt 

cribrosus (LeConte) 

D. truncatus (Melsheimer) 

coruinus (Haldeman) 

D. ulkei Horn 

MB 

MB ON 

ON 

ON 

PQ NB 

PQ 

PQ 

NS 

Genus DIAI-YTELLUS W.J. Brown 

Key to species: Brown (1929) 

D. dialytoides tFall r 

D. humeralis (Leconte) 

-oNPQ 

-oN 

Genus PSAMMODIUS Fallen 

Key to New World species: Cartwright (1955) 

P. caelatus (LeContet 

P. laevipennis Costa+ 

aegialioides Haldeman 

P. oregonensis Cartwright 

P. sulcicollis (Illigertt 

Tribe Psammodiini 

BC 

BC 

-oN 

Dr'l * rq 

Genus RHYSSEMUS Mulsant 

Key to New World species: Gordon and Cartwright (1980) 

R. californicus Horn AB 

R. germanus tLinnet* 

puncticollis W.J. Brown 

R. neglectus W.J. Brown AB 

R. sonatus LeConte AB SK 

-oN 

MB 

pa:) 

Genus PLEUROPHORUS Mulsant 

Key to New World species: Cartwright (1948) 

P. atlanticus Cartwright 

P. caesus (Creutzer)t - BC 

ON 

ON PQ 

Subfamily OCHODAEINAE 

Genus OCHODAXUS Audinet-Serville 

Cadocera Eschscholtz 

Psephus I\ttby 

Stornphax Fischer von Waldheim 

O. luscinus Howden 

O. musculus (Say) 

americanus Westwood 

O. opacus LeConte 

O. simplex LeOonte 

_BCAB 

AB 

SK 

-oN 

MB 

MB 

t49

Scarabaeidae 

Genus BOL,IOCEROSOMA Schaeffer 

Subfamily GEOTRUPINAE 

Key to North American species: Howden (1955) 

Tribe Bolboceratini 

B. bruneri f.tawson & McColloch MB 

Genus EUC.ANTHUS Westwood 

Key to Ncrth American species: Howden (1964) 

E. greeni Rcbinson AB - MB 

E. lazarus iltabricius; - ON 

melibot us (Fabricius) 

Genus BOLTIOCERAS Kirby 

Odontaeus Fischer von Waldheim 

Key to Ncrth American species: Wallis ( 1928) 

B. falli (wallis) - SK MB ON PQ 

B. filicornis Sayt - SK MB 

B. liebecki l ilallisr - ON PQ - NS 

B. obesus (L.econte) - BC AB 

Genus GEOIRUPES Latreille 

(Subgenus (IEOTRUPES s. str.) 

Tlibe Geotrupini 

G. stercorarius (Linn6)t PQ NB - PE - NF 

(Subgenus INOPLOTRUPES Jekel) 

(M ela notr u p e s Blanchard) 

G. ball Jek:l MB ON PQ NB NS 

similis Jekel 

G. hornii Bl rnchard - ON PQ NF 

(Subgenus ()NYCHOTRUPES Jekel) 

G. semiopacus Jekel MB ON PQ 

b la chb ur nii Melsheimer 

melsheimeri Jekel 

oualipennis Jekel 

G. splendidus miarophagus Say - ' 

G. splendidrrs splendidus - ON PQ - NS 

(Fab:ricius) 

gilnichi Jekel 

mixtus Ilorn 

starki .tekel 

Genus LICI INANTHE Burmeister 

Dasyde'ra LeConte 

Key to sp':cies: Carlson (1980) 

Subfamily GLAPFIYRINAE 

L. rathvoni LeConte) 

canina Ilorn 

- BC 

edwara'siHorn 

Subfamily TROGINAE 

Genus GLARESIS Erichson 

Key to Nt,rth American species: Gordon (1970c) 

G. canadens,is W.J. Brown AB - MB 

G. medialis Gordon - BC 

Genus TRO.K Fabricius 

Omorgus Erichson 

Pseudotrox Robinson 

Key to North American species: Vaurie (1955) 

T. aequalis Say MB ON PQ NB NS 

T. atrox LeConte - BC AB SK - ON 

150

Scarabaeidae 

T. capillaris Say 

T. fascifer LeConte 

T. foveicollis Harold 

aciculeatus Robinson 

T. hamatus Robinson 

T. robinsoni Vaurie 

T. scaber (Linn6) 

T. scabrosus Palisot de Beauvois 

T. sonorae LeConte 

alternans LeConte 

lecontei Harold 

T. sordidus LeConte 

T. spinulosus spinulosus Robinson 

T. striatus Melsheimer 

T. unistriatus Palisot de Beauvois 

porcatus Say 

T. variolatus Melsheimer 

erinaceus LeOonte 

monticola Robinson 

_BC 

-ec 

NT 

AB 

AB 

AB 

AB 

SK MB 

_MBON 

SK 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

SK MB ON 

ON 

PQ NB 

PQ 

pe NB 

PQ NB 

NS- 

NS 

NS 

ID 

NF 

Subfamily MELOLONTHINAE 

T?ibe Sericini 

Genus SERICA Macleay 

Camptorhina Kirby 

Stilbolemma Harris 

S. anthracina LeConte 

crdssata Walker 

frontalis LeConte 

robusta LeConte 

ualidaHarcld. 

S. atracapilla (Kirby) 

cucullata Dawson 

S. curvata LeConte 

S. elusa Dawson 

S. georgiana Leng 

lecontei Dawson 

S. intermixta Blatchley 

S. parallela Casey 

S. sericea (Illiger) 

S. sponsa Dawson 

S. tristis LeConte 

_BC 

NT 

BC 

BC 

NT BC 

NT BC 

AB 

SK 

T:^ 

AB SK 

_SK 

MB ON 

MB 

-oN 

MB ON 

-oN 

MB ON 

-oN 

MB ON 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB NS 

NB NS PE 

NB NS PE 

Tribe Melolonthini 

Key to North American genera: Hardy (1974) 

Genus DIPLOTAXIS Kirby 

Alobus LeConte 

Diazus LeConte 

Orsonyx LeConte 

Key to species: Vaurie (1958; 1960) 

D. atlantis Fall 

nigra Caftwight 

D. brevicollis LeConte 

nigropicea (Walker) 

D. haydenii LeConte 

impressifrons Caziet 

innoxia LeConte 

leuicoxa FalI 

D. obscura LeOonte 

cornpacta Fall 

consequens (Walker) 

protninens Fall 

D. sordida (Say) 

carbonaria Burmeister 

rugosioides Schaeffer 

D. subangulata LeConte 

californica Schaeffer 

D. tenebrosa Fall 

AK 

BC 

AB 

AB 

SK 

BC AB SK MB 

BC 

BC 

ON 

ON 

PQ 

151

Scarabaeid,re 

D. tristis K rby MB ON pe NB NS pE 

c or p ul e n t a Burrneis ter 

Genus PHYLLOPHAGA Harris 

Ancylonycha Blanchard 

Endrosa LeConte 

Gymnis LeConte 

Lachnr>sterna Hope 

Triche:;thes Erichson 

Key to North American species: Luginbilt and Painter (1g53) 

P. anxia (LeConte) 

alpina (Linell) 

NT BC AB SK MB ON pe NB NS pE 

- NF 

breuiccllis (Blanchard) 

cephal,.ca (LeConte) 

dubia:.Smittr) 

inspert:ta (Smith) 

p u nc t i r::ol li s (Blanchard) 

uninot:tta (Walker) 

P. balia , Sa/r ON 

comola (Burmeister) 

P. crenulatit tFrrilichr ON 

georgicana (Gyllenhal) 

P. drakii (Kirby) AB SK MB ON pe NB NS pE 

consimilis (LeConte) 

grandi.; (Smith) 

P. ephilida 'Sayt 

burmasteri (LeConte) 

P. fervidatl'abricius) 

arcuatut (Smith) 

quercir a (Knoch) 

P. lraterna Jarris ON pQ 

cognaflt (Burmeister) 

P. fusca (Fr tlich) - BC - oN pe 

- 

feruens (Gyllenhal) 

P. futilis ( Lt.Conte) - ON PQ NB 

decidutt (LeConte) 

gibbosc,. (Burmeister) 

NS NF 

serricornis (LeConte) 

P. gracilis tllurmeisterr 

inana (LeConte) 

- ON 

uoluulo. (LeConte) 

P. hirsuta tl(nocht - ON pe 

P. hirticula Knochr - ON PQ 

P.ilicistKn,rcht - ON 

ciliata LeConte) 

fimb r ia ta (Burmeister) 

ybnesi [ianderson 

porcina (Hentz) 

P. inversa tllorn r ON 

P. lanceolat.t tSayt MB ON 

arizoncte von Bloeker 

cazieri von Bloeker 

grisian,z von Bloeker 

P. longispin,r tSmithr - ON PQ 

P. marginal s (LeConter ON PQ 

P. nitida t Lt Conte) AB - MB ON PQ 

innominata (Smith) 

limula lHorn) 

P. rugosa tl\lelsheimert AB - MB ON pQ 

P. tristis tFa,,bricius) - BC - ON 

pilosicctllis (Knoch) 

P. vilifrons tl.eConter ON pe 

hirticep,s (LeConte) 

Genus POL'IPHYLLA Harris 

MacrorLoria CroLch 

Key to Ncrth American species: Young (1988) 

P. crinita Le Conte 

incolum.is Casey 

- BC 

mystica Casey 

L52

Scarabaeidae 

ona von Bloeker 


peruersa Casey 

santarosae von Bloeker 

P. decemlineata (Say) 

castanea Casey 

com stockiana von Bloeker 

laticauda Casey 

matrona Casey 

oregona Casey 

parilis Casey 

peruersa Casey 

potsiana Casey 

reducta Casey 

ruficollis Casey 

squamotecta Casey 

P. hammondi LeConte 

acomana Casey 

bisinuata Casey 

diffusa Casey 

impigra Casey 

latifrons Casey 

molesta Casey 

oblita Casey 

oklahomensis }].atc}r 

pimalis Casey 

proba Casey 

rufescenta Tanner 

sejuncta Casey 

speciosa Casey 

squatnicauda Casey 

subuittata LeConte 

uerecunda Casey 

P. variolosa (Hentz) 

BC 

AB 

AB 

SK 

SK 

MB 

ON PQ 

Genus AMPHIMAILON Berthold 

A. majalis (Razoumowsky)f 

ON PQ 

Genus PHOBETUS LeConte 

Key to species: Hardy (1978) 

P. comatus LeConte 

leachi Banett 

Tlibe Pachydeniini 

BC 

Genus DICHELONYX Harris 

Dichelonycha lltrby 

Key to North American species: Saylor (1945) 

Key to eastern North American species: Brown (1946) 

D. albicollis (Burmeister) 

D. backii (Kirby) 

arizonica Barrett 

D. canadensis (Horn) 

D. diluta (FalI) 

D. elongatula (Schtinherr) 

D. fulgida (LeConte) 

crotchi (Horn) 

mormona (Fall) 

oregona (Van Dyke) 

D. kirbyi W.J. Brown 

D. linearis (Gyllenhal) 

hexagona (Germar) 

testacea (Kirby) 

D. subvittata (LeConte) 

D. truncata (LeConte) 

Tribe Macrodactylini 

_NTBCABSKMB 

_BCAB 

sc 

AB SK MB 

MB ON 

AB 

SK 

ON PQ NB 

oN PQ 

oN PQ 

ON PQ NB 

_:a)u 

PQ 

MB ON PQ 

NS 

NS- 

_PE 

NB NS 

lrD 

153

Scarabaeida': 

D. vicina (Fall) 

columbinna Hopping 

- BC 

deserta Hopping 

Genus MAC I,ODACTYLUS Latreille 

Stenothyax Haris 

M. subspinor,us (Fabricius) 

barbatu:,s Fitct' 

elongatus (Herbst) 

poly p ha. g us (Burmeister) 

oN PQ 

Genus HOPLIA Illiger 

Echyra Erichson 

Key to No:th American species: Hardy (1977) 

H. callipyge LeConte 

cazieri \loyer 

conuexul,a LeConte 

desertic'tla Boye'r 

humbol,.ltensis Boyet 

irrorata LeConte 

lecontei Dalla Torre 

mutata Gemminger & Harold 

oregona LeConte 

pubicollis LeConte 

utahensis Boyer 

H. Iaticollis LeConte 

H. modesta llaldeman 

barbata Blatchley 

heluola Melsheimer 

monticota Melsheimer 

singularis Burmeister 

H. sackenii LeConte 

H. trifasciate. Say 

ferrisi E:,oyet 

primarit Burmeister 

lrlsfis N.,elsheimer 

Tribe Hopliini 

BC 

BC AB 

_MB?_ 

ON 

MB ON PQ 

NB NS 

Genus ANOUIALA Samouelle 

Anomalepta Casey 

Anomalopus Casey 

Hemispilota Casey 

Oliganomala Casey 

Paranornala Casey 

Rhombanalia Casey 

Spilota Burmeister 

Key to North American species: Potts (1977) 

A. binotata {'}yllenhal t 

compacia Casey 

maculata (Castelnau) 

marginella LeConte 

unifascifia Say 

A. lucicola iF abricius) 

olrolo (llabricius) 

cinctello Burmeister 

moerens (Fabricius) 

nigritula (Castelnau) 

p hy I lope' r thoides Nonfried 

pinicola Melsheimer 

q uadr im.ac ulat a Dr apiez 

A. undulata .Vlelsheimer 


dubia BlatchTey 

Subfamily RUTELINAE 

Tribe Anomalini 

ON 

ON 

ON 

r54

Scarabaeidae 

floridana Casey 

nigropicta Casey 

saginatula Casey 

uarians Burmeister 

Genus STRIGODERMA Burmeister 

Alamona Casey 

Strigodermella Casey 

S. arboricola (Fabricius) 

Genus POPILLIA Audinet-Serville 

P. japonica Newman-i 

ON 

ON PQ 

NS 

Tribe Rutelini 

Genus PELIDNOTA Macleay 

Key to North American species: Hardy (1975) 

(Subgenus PELIDNOTA s. str.) 

(Pelidnotidia Casey) 

P. punctata (Linn6) 

breuicollis Casey 

breuis Casey 


hudsonica Casey 

lutea (Olivier) 

oblonga Casey 

pallidipes Casey 

ponderella Casey 

strenua Casey 

tarsalis Casey 

texensis Casey 

Genus COTAI-PA Burmeister 

Ciocotalpa Saylot 

C. lanigera (Linn6) 

molaris Casey 

fou Wickham 

uernicata Casey 

MB ON PQ 

SK MB ON 

Subfamily DYNASTINAE 

Key to species: Endrodi (1985) 

Tribe Cyclocephalini 

Genus CYCLOCEPHALA Latreille 

Dichromina Casey 

Ochrosidia Casey 

Spilosota Casey 

C. borealis Ar-rolv 

uillosa Burmeister 

C. longula LeConte 

abrupta (Casey) 

ambiens (Casey) 

californica Atrow 

marcida (Casey) 

modulata (Casey) 

obesula (Casey) 

oblongula (Casey) 

phasma (Casey) 

prona (Casey) 

reflexa (Casey) 

regulifrons (Casey) 

rustica (Casey) 

BC 

NS 

155

Scarabaeidae 

Tribe Pentodontinr 

Genus LIGf RUS Burrneister 

(Subgenus L,IGYRODES Casey) 

L. relictus (Say) 

clypealis (Casey) 

dawsoni (Casey) 

paruice,:s (Casey) 

quadriT ennis (Casey) 

uernicicollis (Casey) 

MB ON PQ 

NB 

(Subgenus L,IGYRUS s. str.) 

(Anagrylius Casey) 

(.Ligyretilus Casey) 

L. gibbosus 1,;ibbosus (DeGeer) 

bicorniculatus Casey 

breuiusculus Casey 

curtiper'.nis Casey 

farctus Jasey 

juuencus (Fabricius) 

lacustri; Casey 

laetulu:, Casey 

Iaticaut:ta Casey 

Iongulu; Casey 

lucubla tdus Casey 

morio (LeConte') 

parallelus Casey 

punctiutuda Casey 

remotut: Casey 

rubidus Casey 

texanus Casey 

uariolos us (Burmeister) 

uirginic'"ts Casey 

AB 

ON 

Genus APHONUS LeConte 

Key to No:th American species: 

A. castaneus (Melsheimer) 

cubiforr zis Casey 

obesus (Burmeister) 

saginat,"rc Casey 

trapezictllis Casey 

A. tridentatts (Say) 

aterrim;ts Casey 

congestt,ts Casey 

elongatus Casey 

frater L.:Conte 

modula,lus Casey 

politus l)asey 

scutella:is Casey 

Gill and Howden (1985) 

ON 

PQ 

PQ 

Tribe Oryctini 

Genus XYLC.,RYCTES Hope 

X. jamaicensis (Drury) 

americanus (Palisot de Beauvois) 

faunus \)asey 

hebes Casey 

lacustri:; Casey 


sotyrus (Fabricius) 

tenuicornutus Casey 

ON 

PQ 

156

Scarabaeidae 

Subfamily CETONIINAE 

Tlibe Cetoniini 

Key to New World genera: Hardy (1988) 

Genus EUPHORIA Burrneister 

Erirhipidia Casey 

Erirhipis Burmeister 

Euphorhipis Casey 

Euphoriaspis Casey 

Haplophoria Casey 

Isorhipina Casey 

Rhipiphorina Casey 

E. fulgida (Fabricius) 

E. inda (Linn6) 

barbata (Say) 

brunnea (Gory & Percheron) 

marilandica (Frdlich) 

nigripennis l{lages 

E. rufobrunnea Casey 

BC 

_ONPQ 


Tribe Cremastocheilini 

Genus CREMASTOCHEILUS Knoch 

Key to North American species: Potts (1945) 

(Subgenus CREMASTOCHEILUS s. str.) 

C. armatus Walker 

cribripennis Casey 

densicollis Casey 

maritimus Casey 

montanus Casey 

obliquus Casey 

pilosicollis Horn 

C. canaliculatus Kirby 

hentzi Haris 

C. castaneus Knoch 

breuisetosus Casey 

lecontei Westwood 

pocularis Casey 

C. crinitus LeConte 

bifoueatus Van Dyke 

pugetanus Casey 

C. knockii LeConte 

areolatus Casey 

crenicollis Westwood 

gracilipes Casey 

BC 

BC 

AB 

AB 

SK MB 

SK MB 

-oNPQ 

NS 

(Subgenus ANATRINODIA Casey) 

C. wheeleri LeConte 

SK MB 

Subfamily TRICHIINAE 


Tlibe Osmodermini 

Genus OSMODERMA LePeletier & Audinet-Serville 

Gymnodus Kirby 

O. eremicola (Knoch) 

O. scabra (Palisot de Beauvois) 

beauuoisii Bainbridge 

coseyi Schenkling 

cauiceps Casey 

delicatula Casey 

foueatus (Kirby) 

gracilipes Casey 


rugosa (Kirby) 

O. subplanata Casey 

montana Wickham 

AB 

-oNPQ _ONPQ 

MB ON 

NB 

NS 

NS 

t57

Byrrhidae 

Genus TRITIHIOTINUS Casey 

Tribe Trichiini 

T. affinis (C'ory & Percheron) - ON PQ 

paruuius Casey 

uentricosus Casey 

T. assimilis, (Kirby) NT BC AB SK MB ON PQ NB NS 

bistriga (Newman) 

uariabilis (Burmeister & Schaum) 

T. piger (Fabriciust ON 

drumrzondi (Gory & Percheron) 

reductus Casey 

r ot unc:,'.icol I is (Kirby) 

T. viridans tKirby) - ON 

Genus GNORIMELLA Casey 

G. maculosr tKnocht 

bigs6i, (Kirby) 

- ON PQ 

dissiniilis (Gory & Percheron) 

Superfamily Byrrhoidea 

Family BYRRHIDAE pill beetles 

P.J. Johnson 

Taxonomically, the family is reasonably well 

and Alaska by 26 of the 34 North American species, known in North America. Johnson (-t987) has 

compare(l with about 280 species around the world. treated the bionomics, larval taxonomy, and bio- 

The greatest diversity of Canadian byrrhids is geography of the North American generur, and has 

found in the montane provinces. All species in provided a key to the North American taxa (1991b). 

The lamily Byrrhidae is represented in Canada 

Canada irre bryophagous and express microhabitat 

specifrcity. Many species are associated with the 

moss Cerotodon purpureus on disturbed sandygravelly 

soils. Species of the coastal forests of 

British C.iolumbia and Alaska are closely associated aK (14); yK (9); NT (13); BC (15); AB (1:2); SK (5); 

with certain forest communities and are more MB (7); ON (10); PQ (12);NB (2);NS (a); l,B (5);NF 

polyphagous in their host mosses. (4) . H (3); I (2) 

Genus CHIiETOPHORA Stephens 

Subfamilv SYNCALYPTINAE 

C. spinosa Rossi t+ - BC - ON PQ 

Genus CURIMOPSIS Ganglbauer 

Key to North American species: Johnson (1986) 

C. albonotata (LeConter - YK AB 

grisea LeConte 

C. echinata (LeOonte) AK - NT BC AB SK PQ 

breuicr'tllis Casey 

C. moosilar.ike Johnson AK - NT - AB - PQ - LB 

C. seLulosa (Mannerheim) AK - NT 

Genus BYRRHUS Linn6 

Key to North American species: El-Moursy (1970) 

Subfamilv BYRRHINAE 

Tribl Byrrhini 

B. americanus LeConte 


MB ON PQ - NS - LB 

brunnescens Casey 

158

B1a-rhidae 

criddlei Casey 

imperitus Casey 

manitobae casey 

neglectus Casey 

B. concolor Kirby 

laramiensis Casey 

rigens Casey 

B. cyclophorus Kirby 

canterius Casey 

dfficilis Casey 

fasciatus atct. 

fuluouestitus Casey 

stolidus Casey 

torpidus Casey 

B. eximius LeConte 

callidus Casey 

centralis Casey 

explicatus Casey 

B. geminatus LeContex 

pacificus Casey 

pettiti }{otn 

B. kirbyi LeConte 

consuetus Casey 

egenus Casey 

perditus Casey 

picipes Casey 

uafer Casey 

wickhami Casey 

B. pilula (Linn6)x 

AK YK NT 

AK 

YK 

YK 

NT 

NT 

I'K 

AK NT 

NT 

BC 

BC 

AB 

AB 

SK 

SK MB ON PQ 

PQ 

MB ON PQ 

NB 

LB 

NS LB NF 

Genus PORCINOLUS Mulsant & Rey 

P. undatus (Melsheimer) 

crescentifer Casey 

glabellus Melsheimer 

hystris Casey 

murinus Fabricius 


Tlibe Simplocariini 

Genus SIMPLOCARIA Stephens 

S. elongata J.Sahlberg* - YK 

remotaW.J.Btown 

S. metallica (Sturm) 

colurnbica Casey 

subnuda Casey 

tesselata LeConte 

S. semistriata ( Fabricius)t 

NT 

NT BC AB 

BC 

MB ON 

ON 

PQ 

PQ 

NS 

NF 

NF 

Genus LIOLIGUS Casey 

L. nitidus (Motschulsky) AK 

aequabilis Casey 

heeni Casey 

striolatus Casey 

BC 

Genus LIOON Casey 

L. simplicipes (Mannerheim) AK 

puncticeps Casey 

inflatum LeConte 

speculare Casey 

BC 

Genus EXOMELLA Casey 

Key to North American species: Johnson (1985) 

E. pleuralis (Casey) AK 

BC 

159

Buprestida: 

Tlibe Pedilophorini 

Genus EUSOMALIA Casey 

E. lecontei tWickham) 

BC 

Genus LISIIEMUS Casey 

L. acuminabus (Mannerheim) 

formosus Casey 

satelles Casey 

AK 

BC 

Genus MOIIYCHUS Erichson 

M. aeneolur; (LeConte) 

lateralis (.Fall1 

albertt:. nus (W.J. Brown) 

szbseloszs (Fall) 

hesper rc Wickham 

M. oblongur; (LeConte) 

subcup,reus (Fall) 

insulsus Casey 

AK 

AK 

NT 

BC 

BC 

AB 

AB 

MB ON 

Tribe Tylicini 

Genus TYLICUS Casey 

T. subcanur, (LeConte) 

\T( NT BC 

AB 

SK 

ON PQ 

Genus CYT I LUS Erichsor 

C. alternatrLs (Say) 

nigrans Casey 

C. mimicus Casey 

longultLs Casey 

NT 

BC 

AB 


LB 

NF 

Superfamily Buprestoidea 

Family BUPRESTIDAE metallic wood-boringbeetles 

A. Dauies 

One hundred and fifty-two species of 

Buprestidae have been recorded in Canada and 

Alaska. The adults are usually found resting or 

ovipositi eg on the bark and foliage of trees and 

shrubs, and feed chiefly on pollen-rich flowers; they 

are also ;active fliers. The larvae develop mainly 

under ther bark of the host plants, which are usually 

fairly specific, although a few are leafminers, gallmakers, 

and coniferous cone borers. 

Major taxonomic treatments of the North 

America.n buprestids include those of LeConte 

(1860) and Chamberlin (1926). Bright (1987) provides 

keys and descriptions ofthe Canadian species; 

his handbook has been used as the basis for this 

checklist, with updated distributiorLal data. 

Further revision of the species in such groups as 

Anthaxiinae is required before a proper subgeneric 

arrangement can be established. It should be noted 

that the number of species recorded in the northern 

regions does not reflect the number of species present 

but rather only the lack of collecting in those 

regions. 

AK (11);YK (13); NT (18); BC (75);AB (57); SK (a1); 

MB (63); ON (99);PQ (80);NB (36);NS (22);pE (7); 

LB (4);NF (5) . H (1); I (2) 

160

Buprestidae 

Subfamily POLYCESTINAE 

Tribe Polycestini 

Genus CHRYSOPHANA LeConte 

C. placida (LeConte) 

cupreola Van Dyke 

strandi Obenberger 

BC 

AB 

Subfamily ACMAEODERINAE 

Tlibe Acmaeoderini 

Genus ACMAEODERA Eschscholtz 

A. idahoensis Barr 

A. pulchella (Herbst) 

arizonaeHotn 

flauosignata Gory 

imtnaculata Horn 

mixtaLeConte 

ornata Olivier 

uoluulus Castelnau 

A. tubulus (Fabricius) 

culta (Weber) 

geranii (Harris) 

q uattuor de cimg uttata (F abicirs) 

A. vandykei Fall 

A. variegata LeConte 

distincta Kerremans 

Iucia FalI 

BC 

AB 

-llsoN- 

ON 

Genus ANAMBODERA Barr 

A. gemina (Horn) 

nebulosa (}lorn) 

BC 

Subfamily CHALCOPHORINAE 

T?ibe Chalcophorini 

Genus CHALCOPHORA Solier 

C. flortis LeConte 

cupreola Casey 

Iaurentica Casey 

C. liberta (Germar) 

borealis (Gory & Castelnau) 

paruiceps Casey 

C. virginiensis (Drury) 

ang ulicollis (LeConte) 

- BC AB SK 

antennalis Casey 


c upr eomac ulata (Goeze) 

ingens Casey 

lacustris LeConte 

melanotum Muttkowski 

montana Casey 

nouaebo racensis (Fitch) 

obliterata Casey 

obscurata (FiLch) 

oregonensis (Filc}r) 

pallida Kerremans 


MB ON PQ 

MB ON PQ 

MB ON PQ 

NB 

NB 

NS 

Genus TEXANIA Casey 

T. campestris (Sayt 

substrigosa (Gory & Castelnau) 

ON 

161

Buprestidae 

Genus SPE CTRAJ-IA Casey 

S. gracilipes t Melsheimer) 

Subfamily BUPRESTINAE 

Tlibe Dicercini 

ON 

Genus DICERCA Eschscholtz 

Argante Gistel 

Stenuris I{ttby 

Key to North American species: Nelson (1975) 

D. callosa callosa Casey 

pertirutx Casey 

rigida Casey 

tetrica Casey 

D. callosa f rosti Nelson 

D. caudata LeOonte 

abrup;a Casey 

biangulata Casey 

cupreola Casey 

filiola Casey 

inflatula Casey 

longicu.ud.a Casey 

pisciformis Casey 

D. crassico.llis LeConte 

californica Crotch 

hesperica Casey 

D. divarica;a (Say) 

aestiuo Casey 

angus icauda Casey 

aurich alcea Melsheimer 

dabio Melsheimer 

incisa Casey 

lirnulo Casey 

nigra l)asey 

parumpunctata Melshermer 

rustict:,, Casey 

subaet,ualis Casey 

uancotlueri Casey 

D. dumolini(Castelnau & Gory) 

consobrina Melsheimer 

D. hesperoboreaiis Hatch & Beer 

D. horni Crotch 

ampliuta Casey 

D. lugubris LeConte 

auster't Casey 

lacustt'is LeConte 

morio Casey 

D. lurida (tr'abricius) 

erosa Casey 

floridae Casey 

gaudens Casey 

gracilis Casey 

indis tinc ta Mel sheimer 

indura,ta Casey 

innocua Casey 

leuette,;. Casey 

porcat,'tla Casey 

pruinasa Gory 

regula,is Casey 

sagor lasey 

seriato. Casey 

truncata Casey 

D. pectorosa LeConte 

D. pugionata (Germar) 

D. punctulata (Schtinherr) 

pinorum Casey 

transuersa Say 

D. sexualis Crotch 

D. tenebrica (Kirby) 

longipennis Casey 

prolongata LeConte 

AX YK NT BC 

AK YK 

AB 

_BCAB 

NT_ABSKMBONPQ 

BC 

_BCAB 

_BC 

NT 

_BC_SK 

AB 

SK MB ON PQ 

SK 

SK 

_BC 

NT BC AB SK 

MB ON PQ 

MB ON PQ 

MB ON 

oN PQ 

ON 

oN PQ 

NB 

PQ NB 

NS 

NB NS 


162

Buprestidae 

seuera casey 

subargentea Casey 

subcuprea Casey 

sulcatula Casey 

D. tenebrosa (Kirby) 

acerba Casey 

bifoueata LeConte 

chrysea Melsheimer 

distinguenda (Castelnau & Go.y) 

montana Casey 

mystica Casey 

stolida Casey 

wickhami Casey 

D. tuberculata (Castelnau & Go.y) 

asperata Castelnau 

hilaris LeConte 

mancaLeConte 

molitor Melsheimer 

scobina Chewolat 

AK YK NT BC AB SK MB ON PQ NB NS PE 

oN PQ 

NB NS 

LB 

Genus DESCARPENTRIESINA Leraut 

Key to North American species: Evans (1957) 

D. cyanipes (Say) 

apicilla (Obenberger) 

collaris (Obenberger) 

cupripes (Casey) 

debilis (LeConte) 

erecta (Gory) 

paruiceps (Casey) 

D. ferrea (Melsheimer) 

D. fraseri (Chamberlin) 

D. montana (Chamberlin) 

D. thureura (Say) 

costicollis (Gory) 

NT BC AB SK MB ON PQ NB 

_BCABSK 

-BC 

_BC 

MB ON PQ 

MB 

MB 

Tribe Buprestini 

Genus TRACHYKELE Marseul 

T. blondeli Marseul 

T. nimbosa Fall 

_BC 

-BC 

Genus BUPRESTIS Linn6 

Ancy lo c h ir a Eschscholtz 

Anoplis l{trby 

Gymnota Gistel 

Key to North American species: Helfer (1941) 

B. adjecta (LeConte) 

breuis Casey 

B. aurulenla Linn6 

adulans Casey 

aemula Casey 

affinis Casey 

bicostata Dejean 

chry sochlora (Philippi) 

fabulosa Casey 

lauta (.LeConLe) 

nupta Casey 

prospera Casey 

radians (LeConte) 

tacornae Casey 

uenusta Casey 

uillosa (LeConte) 

B. confluenta Say 

tesselata Casey 

B. fasciata Fabricius 

fortunata Casey 

fulgens Casey 

lherminieri Chevrolat 

saturata Casey 

YK_BC 

BC AB 

BC AB 

AB 

SK 

SK 

MB ON 

MB ON 

MB ON 

PQ 

PQ 

NB NS 

163

Buprestidae 

sexmaculata Hausmann 

sexplagiata (LeConte) 

B. intricateL Casey 

con,tortae Hopping 

murraydnae Hopping 

B. laeviventris (LeConte) 

pugeta,na Casey 

B. langi Mannerheim 

angusta Casey 

bistrinotata Casey 

callida Casey 

crenata Casey 

d.epressa Casey 

fastidt.osa Casey 

gramitea Casey 

incolumis Casey 

Iaeuicirps Casey 

medio:ris Casey 

obliqu.a Casey 

oregon.a Casey 

ornata (Walker) 

patruelis Casey 

sediticsa Casey 

uiridirnicans Casey 

B. lineata ltabricius 

douisi Nicolay & Weiss 

B. lyrata Casey 

acomana Casey 

adducta Casey 

caligitnsa Casey 

fuscat)asey 

lecontt::i Saunders 

morosd Casey 

nigricuns Casey 

B. maculat.ventris Say 

pagancrum l{trby 

rusticorum Kirby 

sexnotzta Gory & Castelnau 

B. maculipennis Gory 

deficiens Casey 

fusiformis Casey 

inconstans Melsheimer 

leporirua Casey 

reductt Casey 

scriptt:. Casey 

B. nuttalli Kirby 

altern':tns (LeConte) 

boulderensis Casey 

conici

Buprestidae 

Tribe Melanophilini 

Genus MELANOPHILA Eschscholtz 

Phaenops Dejean 

Key to North American species: Sloop (1937) 

M. abies Champlain & Knull 

o6iells Obenberger 

M. aeneola Melsheimer 

metallica Melsheimer 

M. californica Van Dyke 

M. drummondi drummondi (Kirby) AK 

guttulata Mannerheim 

umbellatarum (Kirby) 

M. fulvoguttata (Harris) 

croceosignata (Castelnau & Gory) 

decolorata (Castelnau & Gory) 

ftarrisi Obenberger 

octospilota (Castelnau & Gory) 

pacifica Obenberger 

M. gentilis LeConle 

prasina LeConte 

M. intrusa Horn 

ON 

ON 

PQ NB 

BC 

YK NT BC AB SK MB ON PQ NB 

BC 

BC 

BC 

PQ 


LB 

NF 

Genus OXYPTERIS Kirby 

Apatura Castelnau & Gory 

O. acuminata (DeGeer)* AK 

ap pendiculata (Fabricius) 

i mma c u I ata ( Mannerheim) 

longipes (Say) 

morio (Fabricius) 

O. occidentalis Obenberger 

YK 

NT 

BC 

BC 

AB 


NF 

Tribe Anthaxiini 

Genus ANTHAXIA Eschschoitz 

(Subgenus MELANTHAXIA Richter) 

A. aeneogaster Castelnau & Gory 

aerica Ctotch 

grosso Obenberger 

strigata LeConte 

A. aenescens Casey 

A. inornata (Randall) 

expdnsa LeConte 

foueicollis LeConte 

imperfecta LeConte 

A. porella Barr 

A. prasina Horn 

falsula Obenberger 

A. retifera LeConte 

( Subgenus IIAPLANTHAXIA Reitter) 

A. deleta (LeConte) 

A. pseudotsugae (Chamberlin) 

A. quercata (Fabricius) 

cuneiformis (Gory) 

A. viridicornis (Say) 

subaenea (LeConte) 

uiridifrons (Gory) 

Genus AGRILAXIA Kerremans 

A. flavimana (Gory) 

gracilis (Melsheimer) 

_BC 

_YKNTBCABSKMBONPQNBLB 

AK YK 

_BC 

_BC 

BC 

BC 

BC 

BC 

SK 

MB 

MB 

ON PQ 

ON PQ 

ON 

N" 

165

Buprestidae 

Subfamily CHRYSOBOTHRINAE 

Tribe Chrysobothrini 

Key to North American species: Fisher ( 1942) 

Genus CHRYSOBOTHRIS Eschscholtz 

Knowltonia Fisher 

Odontomus l\ttby 

C. adelpha Gemminger & Harold 

soror LeConte 

C. aeneola l,eConte 

(?) purpurata Bland 

C. azttea LeConte 

lecontet Leng 

C. beeri Barr 

C. blancharrli Horn 

C. breviloba Fall 

C. carinipernis LeConte 

C. caurina IIorn 

C. columbiana Barr 

C. cribraria Mannerheim 

calcarota Melsheimer 

f'Loricola Gory 

lalo Kerremans 

C. dentipes iGermar) 

c har acr e r is tica ( Harris) 

planattt Gory & Castelnau 

posticatis Gory & Castelnau 

rotund,collis Gory & Castelnau 

C. femorata (Olivier) 

alaban,ae Gory 

dissim,,.Iis Dejean 

fastidit,sa Gory 

insculF'ta (Herbst) 

lesueuri Gory & Castelnau 

misello LeConte 

nigrituta Gory & Castelnau 

obscurr.t LeConte 

quadri;',mpresso Gory & Castelnau 

C. monticoler Fall 

grandis Chamberlin 

C. harrisi (Flentz) 

C. laricis Van Dyke 

C. leechi Barr 

C. mali Horn 

C. neopusilla Fisher 

C. nixa Horr 

C. oregona Ohamberlin 

planon .arginafo Chamberlin 

C. orono Frost 

C. pseudots rgae Van Dyke 

C. pusilla Gory & Castelnau 

bigutta,ta Gory & Castelnau 

str ang ulata Melsheimer 

C. rugosicelrs Melsheimer 

C. scabripennis Gory & Castelnau 

proxima (Kirby) 

scabra Goty 

C. sexsignata (Say) 

germari Gory & Casteinau 

ignipes Gory & Castelnau 

serguttata (Say) 

C. sylvania Fall 

C. trinervia (Kirby) 

cicat r i cos a Motschulsky 

C. verdigripennis Frost 

C. viridiceps Melsheimer 

C. vulcanica LeConte 

canadensis Chamberlin 

_BC 

BC 

AB 

AB 

_BCAB 

_BCAB 

_BC 

_BC 

NT_ABSK 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

sc 

_BC 

AK YK NT BC 

SK 

SK 

AB SK 

ON 

ms oll 

lln Of,l 

PQ 

ON PQ 

PQ 

pe 

SK MB ON PQ 


AB 

AB-MBON 

AB 

AB 

AB SK MB 

_ _ T:'. 

MB ON 

MB ON 

PQ 

PQ 

PQ 

xr nc AB ix MB 3N i3 

nc 

AB SX 

AB SK 

AB 

oN PQ 

MB ON PQ 

MB ON PQ 

MB ON 

NB 

NB 

NB NS 

NB 

NS_ 

NS_ 

fu 

NB 

NB 

PE 

NB 

PE 

NB NS 

NB NS 

LB 

NE 

166

Buprestidae 

Subfamily AGRILINAE 

Tribe Coraebini 

Genus EUPRISTOCERUS Dey'r'olle 

(? ) R haeb oscelis LeConte 

E. cogitans (Weber) 

ignarus (Fabricius) 

ruficollis (Herbst) 

Genus AGRILUS Curtis 

A. acutipennis Mannerheim 

latebrus Castelnau & Gorv 

A. anxius Gory 

grauis LeConte 

torpidus LeConte 

A. arbuti Fisher 

manzanitae Chamberlin 

A. arcuatus (Say) 


A. atricornis Fisher 

A. aurichalceus Redtenbacherf 

A. bilineatus bilineatus (Weber) 

aurolineatus Gory 

biuittatus l{trby 

flau olineatus Mannerheim 

triuittatus Horn 

A. bilineatus carpini Knull 

A. burkei Fisher 

A. celti Knull 

A. cephalicus LeConte 

A. crataegi Frost 

A. criddlei Frost 

A. crinicornis Horn 

A. cyanescens (Ratzeburg)t 

A. defectus LeConte 

A. egenus Gory 

A. fallax Say 

impressipenni.s Uhler 

zemes Gory 

A. frosti Knull 

A. geminatus (Say) 

A. granulatus populi Fisher 

tr ic hocar p ae Chamberlin 

A. horni Kerremans 

A. imbellis Crotch 

A. juglandis Knull 

A. lacustris LeConte 

cuneus LeConte 

pubiuentris Ctotch 

A. lecontei Saunders 

subfasciatus LeOonte 

A. liragus Barter & W.J. Brown 

A. malvastri Fisher 

A. masculinus Horn 

A. nigricans Gory 

auricomus Frost 

A. obsoletoguttatus Gory 

interruptus LeConte 

subaeratus Hatris 

A. otiosus Say 

uirens Goty 

A. parvus Saunders 

pusillus Say 

A. pensus Horn 

betulae Fisher 

A. politus (Say) 

canadensis Obenberger 

cupreolus LeConte 

AK 

Tlibe furilini 

_BC 

_MBONPQ 

ON PQ NB 

BC AB SK MB ON PQ NB NS 

-ncAB--9*PQ----- 

-oN 

-BCAB-MBONPQ 

AB 

MB ON PQ NB 

-ONPQ 

_ONPQ 

MB ON PQ 

ON PQ 

_ _ _:"yu3Ni3Iu_ _ _ _ 

BC AB 

-ONPQ 

-ONPQ 

-oN 

-oNPQ 

_MBONPQ 

oN PQ 

- - - -"gN;:- 

-oN 

---:"ffisKYuoNPQNB--- 

-SKMBONPQ 

-oN 

_ONPQ 

MB ON PQ 

MB 



NF 

167

Buprestidar: 

desertus LeConte 

plumbcus LeConte 

solitar,t,us Harold 

A. pseudoccryli Fisher 

A. putilius l5ay 

A. rufrcollis (Fabricius) 

cupricollis Goty 

A. sayi Saurders 

browni Catlson & Knight 

lateralis (Say) 

A. subcinctus Gory 

A. vittaticollis (Randall) 

frenatus Gory 

A. walsinghami Crotch 

BC 

BC 

-MBONPQ oN PQ 

-MBONPQ 

_PQ 

ON 

oN PQ 

NB NS 

Subfamily TRACFTYINAE 

Tribe Pachyschelini 

Genus PACHYSCHELUS Solier 

P. confusus Wellso & Manley 

P. laevigatts (Say) 

oblong zs (Motschulsky) 

ouatus (Say) 

politus Kertemans 

punctatus (Goty) 

P. purpurer.,s (Say) 

biimpr essus Motschulsky 

AB 

-oN 

_ONPQ 

ON 

Tribe Brachyini 

Genus BRACHYS Solier 

B. aerosus l1{elsheimer 

rufescens Nicolay & Weiss 

B. aeruginosus Gory 

B. ovatus (\Veber) 

auruleztus (Kirby) 

bellporli Nicolay & Weiss 

horni }(erremans 

laeuicauda LeConte 

molestus Gory 

termin,zns Castelnau & Gory 

AB 

SK MB ON PQ 

-oNPQ _ONPQ 

Genus TAPFIROCERUS Solier 

T. gracilis (iiay) 

albog u tt at us (Mannerheim) 

grossu:; Obenberger 

texanus Kerremans 

T. nicolayi Obenberger 

T. schaefferi Nicolay & Weiss 

AB 

SK MB ON PQ 

- - ABsr- 3N- - - - - - 

168

Heteroceridae 

Superfamily Dryopoidea 

Family PTILODACTYLIDAE toed-winged beetles 

L. LeSage 

The family Ptilodactylidae, which is primarily 

tropical in distribution, comprises about 500 species 

around the world, but only three occur in Canada. 

Adults hide in the riparian vegetation or in the leaf 

litter during the day; they are easily attracted to 

light. Larvae of Ptilodactyla prefer moist or wet 

habitats, such as beaver houses or wet, decaying 

plant matter; those of Anchytarsus are truly 

aquatic, burrowing in submerged decaying wood in 

creeks (LeSage and Harper 1976a). 

Stribling's recent key (1986) is useful for identifying 

the genera. 

BC (1);MB (1);oN (2);PQ (3);PE (1) 

Genus FTILODACTYI,A Illiger 

Key to North American species: Johnson & Frey'tag (1982) 

P. serricollis (Say) 

elate r ina Gu6rin-M6neville 

- BC 

/usca Melsheimer 

P. carinata Johnson & Freytag 

MB ON PQ 

PQ 

PE 

Genus ANCIIYTARSUS Gu6rin-M6neville 

Key to North American species: Stribling (1986) 

A. bicolor (Melsheimer) 

ofer Gu6rin-M6neville 

debilis Ziegler 

ON 

PQ 

Family HETEROCERIDAE variegated mud-lovingbeetles 

L. LeSage 

Heterocerids are represented in Canada and 

Alaska by 25 species. The adults are nocturnal and 

are readily attracted to light. During the day they 

hide and make tunnels in sandy areas near water 

courses or standing water, a behavior shared by the 

Iarvae. Adults and larvae feed on zooplankton, 

diatoms, and algae. 

Pacheco (1964) proposed a drastic change in the 

traditional treatment of the family when he 

described 15 new genera and restricted the genus 

Heterocerus to the OId World. Although several of 

his new genera are questionable, his classifrcation 

has been followed for this checklist except for the 

genus Dampfius, which was recently placed in synonymy 

witlr' Heterocerus by Miller (1988b). The 

Canadian fauna is relatively well known, but new 

species are still being discovered (Miller 1988o, 

1988b). Both the arctic and the maritime fauna 

have been little surveyed. Pacheco (1978) has 

recently cataloged the North American species. 

aKQ); YK (a);NT (5); BC (e);AB (16); SK (14); MB 

(1e);ON (16);PQ (16);NB (5);PE (1) 

Genus EXPLORATOR Pacheco 

E. canadensis (Fall) 

AK 

Tlibe Auryliini 

YK NT AB SK MB 

Genus CENTURIATUS Pacheco 

C. auromicans (Kiesenwetter) 

C. compactus (Fall) 

_ -_ _ :"ffi:"MBg"i8 _ _ _ _ _ 

r69

Heteroceridae 

Genus MICIIOAUGYLES Pacheco 

M. moleculus (Fall) 

M. mundulus (Fall) 

Genus LAN'IERNARIUS Pacheco 

L. brunneus (Melsheimer) 

L. gemmatur tHornt 

L. mollinus (Kiesenwetter) - 

L. parrotus llacheco 

L. sinuosus .Pacheco 

_ABSKMB_PQ 

-AB 

Tlibe Heterocerini 

_NTBCABSKMBONPQPE 

BC AB SK 

_ABSKMBONPQ 

_NTBCABSKMBONPQNB 

YK NT BC AB SK MB 

Genus NEO F{ETEROCERUS Pacheco 

N. fatuus lKiesenwetter ) - 

N. gnatho {I,econte) 

labratus (LeConte) 

N. longilobu lus Pacheco 

N. pallidus tSayt 

uentrahs (Melsheimer) 

N. sandersoni Pacheco 

YK 

_BCABSK 

_ABSK 

MB ON 

MB ON 

MB ON 

MB 

pe 

PQ 

PQ 

NB 

NB 

Genus HETEROCERUS Fabricius 

Dampfius Pacheco 

Key to North American species: Miller (1988b) 

H. cof laris (liesenwetter) - 

H. inciertus Pacheco.) 

H. mexicanus Sharp 

H. subtilis Vt.V. Miller 

H. undatus {Melsheimer; 

H. unicus W V. Miller 

_BCAB 

SK 

AB SK 

_BCABSK 

MB ON PQ 

MB ON PQ 

-oN 

MB ON PQ 

MB ON PQ 

lvlD 

NB 

Genus PEDI TATUS Pacheco 

P. schwarzi Horn) 

P. tenuis tW V. Miller) 

_ONPQ 

MB ON 

Genus LAPSUS Pacheco 

L. tristis (Mannerheim) AK 

YK 

NT 

BC 

AB 

SK 

MB ON PQ NB 

Genus TROI'ICUS Pacheco 

Tribe Tropicini 

T. pusillus {flayt 

americanus (Sharp) 

- ON 

Iimbatus (Kiesenwetter) 

luteolus (LeConte) 

PQ 

170

Dryopidae 

FamilyLIMNICHIDAE minute marsh-lovingbeetles 

L. LeSage 

Of the three subfamilies and 15 genera of 

Limnichidae described in the Western Hemisphere, 

only two genera and four species occur in Canada. 

Adults of limnichids are usually active during the 

day near water in leaf litter; they are readily 

attracted to light. The larvae are rarely collected 

and are not represented in most collections. Paulus 

(1970) frrst described the immature stages of two 

European species: they are terrestrial, Iive in moist 

soil, and tunnel under layers of mosses and algae, 

which thev use as food. 

This group of beetles was first considered as a 

tribe of Byrrhidae, but Hinton (1939o, 1939b) elevated 

it to family rank and placed it in the superfamily 

Dryopoidea, after a study of the internal 

anatomy. The North American limnichid fauna is 

now well known after a series of recent revisions by 

Wooldridge, who also published a catalog of the 

North American species in 1986. 

eC (1);MB (1);oN (4);PQ (2) 

Subfamily LIMNICHINAE 

Tribe Limnichini 

Key to North American genera: Wooldridge (1975) 

Genus LIMNICHITES Casey 


Wooldridge (1977) 

L. huronicus Casey 

L. olivaceus (LeConte) 

uirginicus Casey 

L. punctatus (LeConte) 

austinianus Casey 

____ 

BC 

MB 

ON 

ON 

oN PQ 

Genus LUTROCHUS Erichson 


Brown (1972) 

L. laticeps Casey 

ON PQ 

Family DRYOPIDAE long-toed water beetles 

L. LeSage 

Dryopidae includes over 200 species around the 

world, but only six occur in Canada. Adults are 

essentially phytophagous, feeding on diatoms, 

algae, and decaying wood. The truly aquatic adults 

of Helichus inhabit riflles of water courses, using a 

hydrofuge plastron for breathing, as do elmids. 

Adults of Dryops are typically riparian. They can 

take flight directly from water surface when disturbed. 

Dryopid larvae are rarely collected; the few 

specimens collected were found in moist soil on 

banks of creeks where the adults live. 

The family has not been revised since Brown 

(1972) put together all the taxonomic information. 

The Canadian fauna is reasonably well known for 

Quebec and Ontario (LeSage and Harper 1975). 

Brown (1983o) recently published a catalog of the 

North American species. 

sC (2); AB (1); SK (1); MB (1); ON (2); PQ (5); NB 

(2);NS (2);PE (1);NF (1) ' I (1) 

Genus DRYOPS Olivier 

(Subgmus YRDOPS Steffan) 

D. viennensis (Heer)t 

PQ 

Genus HELICHUS Erichson 

Key to North American species: Musgrave (1935); Brown (1972) 

L77

Elmidae 

H. basalis L,:Conte 

H. columbia.rus W.J. Brown 

H. fastigiatts (Say) 

H. lithophih.Ls (Germar) 

H. striatus LeConte 

foueatus LeConte 

--ecAB 

BC 

SK 

MB 

ON 

ON 

PQ 

PQ 

PQ 

PQ 

NB NS 

NS NS 

PE_NF 

Familv ELMIDAE riffle beetles 

L. LeSage 

Elmidae comprises about 1100 species around 

the world: 32 of them occur in Canada. Elmids are 

small beetles, usually less than 3 mm long, and most 

are aquatic in both the adult and the larval stages. 

Adults breathe by means of a hydrofuge ventral plastron, 

the arvae with retractile cloacal gills. Adults 

and larver e inhabit the substratum of creeks and 

rivers, feeding on diatoms, encrusted algae detritus, 

or submer:ged decaying wood (LeSage and Harper 

t976b); they are good indicators of water quality 

(Hilsenhoff I977). Pupation occurs near water, 

under ro

Elmidae 

Genus HETERLIMNIUS Hinton 

Ceralimnius Hinton 

Key to North American species: Brown (1972) 

H. corpulentus tl,eContet - BC AB 

antennatus (Fall') 

H. koebelei (Martinj - BC AB 

Genus MICROCYLLOEPUS Hinton 

Key to North American species: La Rivers (1949) 

M. pusillus pusillus (LeConte) - BC - ON PQ NB 

browni (Hatch) 

Genus NARPUS Casey 

N. concolor (LeConter - BC AB 

solutus (W.J. Brown) 

Genus OPTIOSERVUS Sanderson 

Key to North American species: White (1978) 

O. divergens (LeConter - BC AB SK 

pecosensis (Fall) 

O. fastiditus (LeConte) NT BC AB SK MB ON PQ NB NS PE - NF 

O. quadrimaculatus tHorn) - BC AB 

O. ovalis tl-eContet - ON PQ NB NS PE LB 

ampliatus (Fa'r\) 

O. seriatus (Leconte) - BC AB 

O. trivittatus (W.J. Brown) - ON PQ NB NS NF 

Genus OULIMNIUS des Gozis 

O. latiusculus (LeOonte) - ON PQ NB NS PE - NF 

Genus ORDOBREVIA Sanderson 

O. nubifera (Fall) - BC 

Genus PROMORESIA Sanderson 


P. elegans ll-eContet PQ 

P. tardella (Fall) PQ NB NS - LB NF 

subarctica (W.J. Brown) 

Genus STENELMIS Dufour 


S. bicarinata LeConte - ON PQ - NS 

S. concinna Sanderson PQ 

S. crenata (Say) MB ON PQ NB NS PE - NF 

sordida Motschulsky 

S. mera Sanderson PQ NB NS 

S. musgravei Sanderson PQ 

S. quadrimaculata Horn - ON PQ 

blatchleyi Musgrave 

sulcata Blatchley 

S. sandersoni Musgrave - ON PQ 

tarsalis Sanderson 

S. vittipennis Zimmermann - SK MB ON PQ 

Genus MACRONYCHUS P.W.J. Miiller 

Key to North American species: Hinton (1940) 

T?ibe Macronychini 

M. glabratus Say - ON PQ NB NS 

lateralis Melsheimer 

Genus ZAITZE\rIA Champion 

Z. milleri H.P. Brown - BC 

Z. parvula (Horn t - \'K - BC AB 

columbiensis (Angell) 

173

Artematopidae 

Family PSEPHENIDAE water-penny beetles 

L. LeSage 

Psephenids form a small family of aquatic beetles; 

onl.' three species have been recorded so far in 

The taxonomy of the North American 

psephenids is relatively well known, tha.nks to the 

Canada. Their vernacular name refers to the works of Brown (1972, 1983b), Brown and Murvosh 

unique round and flat shape of the body of the lar- (I974), and Brigham (1981). 

vae, which live in riffles of creeks and rivers. The 

adults o1' Ectopria spend the daytime in the foliage 

of marginal vegetation; they are readily attracted to 

light. Tlrose of Psephenus scurry over exposed rocks 

in riflles during the day and hide at night. ON (3); PQ (2);NB (1); NS (2) 

Genus EC,fopRrALeconte 

Eurea LeConLe 

Key to Irforth American species: Brigham (1981) 

subfamily EUBRIINAE 

E. leechi Brigham - ON 

E. nervosa tMelsheimerr - ON PQ - NS 

tarsalis LeConte 

thorar:'ica (Ziegler) 

tibiali)s LeConte 

subfamily PSEPHENTNAE 

Genus ps[]pHENUS Hardeman 

Eurytr:.alpus LeConte 

Fluuit:'ola DeKay 

Key to liorth American species: Brown & Murvosh (1974) 

P. herricki lDeKayr 

lecontei (LeConte) 

- ON PQ NB NS 

t re nto nen si s Zimmermann 

Superfamily Elateroidea 

Family ARTEMA*TOPIDAE 

Y. Bousquet 

Thi-" family includes eight North American one found in North America. Many authors, includspecies, 

four of which are currently known to occur ing Arnett (1968), have placed the macropogonines 

in Canarla. Little is known about the bionomics of within Dascillidae. 

the Near:ctic species. The adults are usually found 

on foliage; the larvae of some species have been 

reared or.r mosses (Lawrence 1982q.). 

Lawrence (1982a) recognized three subfamilies NT (1); BC (3); AB (1); MB (1); ON (2); PQ (2); NB 

of Artematopidae, Macropogoninae being the only (1) 

Genus MACROPOC'ON Motschulsky 

Key to North American species: Hopping (1936) 

Subfamily MACROPO GONINAE 

M. piceus l,eConte NT BC AB - MB ON PQ NB 

dubiusW.J. Btown 

rufipes l{orn 

M. testaceipennis Motschulsky - BC 

174

Elateridae 

cribricollis W.J. Brown 

rubricollis Pic 

Genus EURYPOGON Motschulsky 

Dasypogon LeConte 

E. californicus Horn 

E. niger (Melsheimer) 

horrisi (Westwood) 

BC 

oN pe 

Family ELATERIDAE click beetles 

Y. Bousquet 

This family is well known for the ability of the 

adults to click when lying on their back. It includes 

369 species and subspecies in Canada and Alaska. 

The adults occur on foliage and flowers, under bark, 

and in rotting wood; the larvae are found in leaf litter, 

soil, moss, and rotting wood. Larvae of most species 

living in leaf litter and rotting wood are predaceous; 

those occurring in the soil feed on roots, and some are 

very injurious to crops. Adults of some species apparently 

do not feed, whereas those ofother species probably 

feed on tender plant tissues, such as leaves and 

flower buds. 

The classifrcation of the family followed here is 

Hyslop's (1917), with slight modifrcation. His classifrcation 

is preferred Stibick's (1979), as it is used more 

widely. There is no recent taxonomic revision of the 

North American species. Most species that occur in 

the areas covered in this checklist could be identifred 

using Dietrich (1945), Brooks (1961), Lane and 

Lancaster (1971), and the taxonomic reYisions listed. 

AK (53); YK (32); NT (49); BC (19a); AB (131); SK 

(88);MB (99); ON (175);PQ (165);NB (81); NS (87); 

PE (21);LB (25);NF (46) ' H (6);I (5) 

Subfamily PYROPHORINAE 

Tlibe Pyrophorini 

Genus LACON Castelnau 

Aulacon Arnett 

Diphyaulon Arr:.ett 

Lepidotus Stephens 

Zalepia Arnett 

Key to North American species: Arnett (1952) 

L. auroratus (Say) 

L. avitus (Say) 

L. discoideus (Weber) 

pennatus (Fabricius) 

L. impressicollis (Say) 

lepturus (Say) 

senilis (Germar) 

L. maculatus (LeConte) 

L. marmoratus (Fabricius) 

L. rorulentus (LeConte) 

L. sparsus (Candbze) 

_ -_ v_ 

-ec-- 

-BC 

yu 

oN PQ 

oN PQ 

ON 

ON 

ON 

oN PQ 

NB 

Genus DANOSOMA Thompson 

Key to North American species: Arnett (1952) 

D. brevicorne (LeConte) AK 

cauicolle (LeConte) 

profusutn (Candbze) 

uetusturn (Walker) 

D. obtectum (Say) 

YK 

YK 

NT 

NT 

BC 

BC 

AB 

SK 

MB ON PQ NB 


NS 

Genus ALAUS Eschscholtz 

A. melanops LeConte 

naja Canddze 

A. myops (Fabriciust 

canadensis Canddze 

A. oculatus (Linn6) 

luscus (Fabricius) 

BC 

oN PQ 

oN PQ 

]-75

Elateridae 

Genus DR.A.STERIUS Eschscholtz 

D. debilis L,eConte 

incongruus Fall 

AK YK BC AB SK MB ON PQ NB NS 

Genus AEC'LUS Eschscholtz 

A. amabilis (LeConte) 

A. mellillus comis (LeConte) 

A. mellillus marginicollis (Horn) 

A. mellillus mellillus (Say) 

dorsali.s (Say) 

NT BC AB 

sx 

-oN 

MB 

-oN 

PQ 

Genus COIT|ODERUS Eschscholtz 

C. auritus (Herbst) 

caseyi'.Leng) 

crassicollis (Melsheimer) 

dilectus (Say) 

C. bellus (Say) 

piclzs lSteinheil) 

uariegutus (Steinheil) 

AB 

SK MB ON PQ 

ON 

NS 

Tribe Pityobini 

Genus PIT)|OBIUS LeConte 

P. anguinusi LeConte 

billing.si Bland 

AR SK MB ON PQ NB 

NS 

Tlibe Athoini 

Genus LIM,INIUS Eschscholtz 

L. aeger Leilonte 

hnulli ?all 

L. agonus (fiay) 

L. anceps L:Conte 

L. aurifer LeConte 

L. auripilis (Say) 

pubico;Iis (LeConte) 

L. basillaris (Say) 

semiaeneus LeConte 

L. bicolor Van Dyke 

L. cal ifornicus t Mannerheimj 

hispidus LeConte 

occider,.talis Candbze 

L. canus LeConte 

discoiaeus LeConte 

subcosi atus Motschulsky 

L. confusus LeConte 

aenesc(:ns LeConte 

L. crotchi Horn 

L. ectypus (Say) 

L. griseus (l']alisot de Beauvois) 

hirticot lis Melsheimer 

propexits Canddze 

L. humidus Lane 

L. infuscatus Motschulsky 

L. lanei Van Dyke 

L. lecontei Lane 

consimilis Walker 

nitidicollis LeConte 

L. maculicollis Motschulsky 

discicollis Canddze 

L. nitidulus Horn 

L. pectoralir; LeConte 

L. pictus Van Dyke 

L. pilosulus Canddze 

pilosus LeConte 

L. plebejus {Say) 

m e ta I I e scens Melsheimer 

AK YK 

BC 

_BC 

_BCABSK 

BC 

BC 

AB 

_BC 

_BCAB 

_BCABSKMB 

_BCABSK 

_BC 

_BC 

NT BC 

_BC 

-BC 

MB ON PQ NB 

ON PQ 

SK MB ON PQ 

_MBONPQ 

ON PQ 

SK 

SK 

MB 

mn- 

-oN 

ON 

MB ON 

PQ 

oN PQ 

ABy:" LB NT 

oN PQ 

NB 

PQNB_ 

NS 

NS 

PE 

LB 

NF 

176

Elateridae 

L. quercinus (Sayt 

L. rudis W.J. Brown 

L. rufihumeralis Lane 

L. seminudus Van Dyke 

L. snakensis Lane 

L. stigma tHerbst) 

armus (Say) 

L. subauratus LeConte 

L. ursinus Van Dyke 

L. venabelsi Wickham 

ECAB_ 

BC_- 

BC 

BC 

AB 

SK 

MB 

Genus ELATHOUS Reitter 

E. californicus Van Dyke 

E. discalceatus (Say) 

ober ndor ffe r i ( Schwarz ) 

E. nebulosus (Van Dyke) 

BC 

BC 

ON PQ 

NS 

Genus ATHOUS Eschscholtz 

Crepidophorus Mulsant & Guillebean 

Key to eastern North American species: Becker ( 1974) 

Key to North American species: Becker (1979) 

A. acanthus (Say) 

carolinus Van Dyke 

flauangularis Hotrr 

maculicollis LeConte 

A. brightwelli {Kirby} 

arcticollis Melsheimer 

oblong icoll is Melsheimer 

A. campyloides Newmant 

affinis Coupet 

A. cucullatus {Say) 

hypoleucus Melsheimer 

p rocer icoll is Melsheimer 

stripatus Melsheimer 

A. lossularis (LeConte) 

A. imitans Fall 

A. neacanthus Becker 

A. nigropilis Motschulsky 


nugalis Fall 

orophilus Harold 

speculifer FalI 

A. orvus Becker 

A. posticus (Melsheimert 

erebusYanDyke 

A. productus (Randall) 

bipunctatus Provancher 

A. rufrfrons (Randallt 

lengiDtry 

reflexus LeConte 

A. rufrventris rectithorax Reitter 

recticollis Fall 

A. rufrventris rufiventris AK 

(Eschscholtz) 

fer r ug inos us Eschscholtz 

maritimus Van Dyke 

A. scapularis (Say) 

A. scissus LeConte 

ingens Fall 

A. sierrae varius Lane AK 

A. vittiger LeConte 

BC 

BC AB 

BC 

BC 

BC 

BC 

BC 

AB 

SK 

-oNPQ 

_ONPQ 

-oNPQ 

MB ON 

MB ON 

ON PQ 

ON PQ 

-PQ 

ON PQ 

PQ 

PQ 

PQ 

___9*PQ_____ 

NS 

NB NS 

NB NS 

_NS 

NB 

NB 

NS 

PE 

NST_ 

_ 

NF 

NF 

Genus HARMINIUS Fairmaire 

H. triundulatus (Mannerheim) AK YK 

Genus MICRATHOUS Lane 

M. brevis (Van Dyke) 

Genus DENTICOLLIS Piller & Mitterpacher 

Key to North American species: Becker (1952) 

BC 

BC 

SK ON PQ 

177

Elateridae 

D. denticor nis { Kirby t 

flauinrcus (Melsheimer) 

D. varians (Germar) AK 

uariabilis (Eschscholtz) 

YK 

NT BC 

NT BC 

AB 


NS 

NF 

Genus BERNINELSONIUS Leseigneur 

Key to s;cecies: Stibick (1976) 

B. hyperboreus (Gyllenhal)* AK 

plona,:us (Eschscholtz) 

NT 

Genus LIGMARGUS Stibick 

Key to N.orth American species: Stibick (1976) 

L. funebris (Canddze) 

L. lecontei (Leng) 

planaius (LeConte) 

AK YK 

NT 

BC 

AB 

-oN PQ NS 

Genus ASCOLIOCERUS M6quignon 

Scolio :erus Motschulsky 

Key to North American species: Stibick (1978) 

A. sanborni(Horn)x AK 

barba:tus (J.R. Sahlberg) 

YK NT AB 

MB 

PQ 

Genus MAI,GAIOSTUS Stibick 


(Subgenus MARGAIOSTUS s. str.) 

M. glacialirr (Van Dyke) AK 

M. grandic'rllis (Lecontel 

BC 

AB 

oN PQ 

Genus HYI'OLITHUS Eschscholtz 

C ry pto hy pnus Eschscholtz 

Key to n*orth American species: Stibick (1978) 

H. littoralir; Eschscholtzx AK 

Genus HYPNOIDUS Dillwyn 


H. abbreviatus (Say) AK 

silaceipes (Germar) 

H. bicolor ( Eschscholtz)* AK 

fallax (Mamer}rein) 

lacustris (LeConte) 

limbatus (Mannerheim) 

picescens (LeConte) 

pumilits (Dejean) 

scar ifi c at u s (Mannerheim) 

uestitus (Mannerheim) 

H. impress collis (Mannerheim) AK 

H. leei Stibick 

H. nocturnus (Eschscholtz) AK 

H. rivularius (Gyllenhal)* AK 

H. squalidr.s tLeContet 

YK 

YK 

BC 

NT BC 

YK NT BC AB 

_AB 

YK_BCAB 

YKNTBC_SK 

BC AB 

AB SK 

AB SK 

MB ON 

MB ON 

MB ON 

MB 

PQ 

PQ 

PQ 

PQ 

NB NS 

NB NS 

PE 

LB 

LB 

NF 

NF 

NF 

NF 

Genus OEDOSTETHUS LeConte 

O. femoralis LeConte 

AB 

SK 

MB ON 

PQ 

NB 

Genus NEGASTRIUS Thomson 

N. caurinus lHorn; 

N. choris (Sayt 

N. delumbi s tHorn) 

N. dispersus (Horn) 

N. dubius ( Horn) 

N. exiguus (Randallt 

guttatulus (Melsheimer) 

N. extricatus (Fall) AK 

N. gentilis rLeConte) 

N. manki (Fall) 

N. melsheirneri tHorn) 

N. musculus (Eschscholtz) AK 

YK NT 

BC 

_ABSK 

BC 

_ABSK 

BC AB SK 

BC AB 

BC 

MB ON 

MB 

-oN 

_PQ 

PQ 

yI' 

NB NS 

PE 

178

Elateridae 

N. obliquatulus (Melsheimer) 

N. ornatus (LeConte) 

colon (Horn) 

moerens (Hotrr) 

N. pectoralis (Say) 

futilis (LeCofte) 

inops (LeConte) 

N. restrictul us (Mannerheim.) 

N. striatulus (LeConte) 

N. tumescens LeConte 

Genus CTENICERA Latreille 

Corymbites Latreille 

C. aeripennis (Kirby) 

tincta (LeConte) 

C. angularis (LeConte) 

C. angusticollis (Mannerheim) 

fraterna (LeConte) 

spectabilis (Mannerheim) 

C. appressa (Randall) 

mirifica LeConte 

C. appropinquans (Randall) 

C. arata (LeConte) 

C. atropurpurea (Melsheimer) 

C. bipunctata (W.J, Brown) 

C. bombycina (Germar) 

C. callida (W.J. Brown) 

C. carbo LeConte 

C. caricina (Germar) 

C. comes (W.J. Brown) 

C. confluens (Gebler) 

C. conjungens (LeOonte) 

C. consors (W.J. Brown) 

C. crestonensis (W.J. Brown) 

C. cylindriformis (Herbst) 

appressifrons (Say) 

breuicornis (Say) 

p ar al lel ip ip eda ( Germar) 

submetallica (Germar) 

C. destructor (W.J. Brown) 

C. diversicolor (Eschscholtz) 

C. dorothyae Knull 

C. elongaticollis (Hamilton) 

C. fallax (Say) 

C. falsifrca (LeConte) 

C. fulvipes (Bland) 

C. funerea (W.J. Brown) 

C. furtiva (LeConte) 

monticola Horn 

C. glauca (Germar) 

simili s sima ( Motschulsky) 

C. hamata (Say) 

C. hieroglyphica (Say) 

C. hoppingi (Van Dyke) 

C. inflata (Say) 

C. insidiosa (LeConte) 

C. kendalli Kirby 

anchorago (Randall) 

C. laricis (W.J.Brown) 

C. lateralis (LeConte) 

q uadr iu ittat a (Walker) 

uittata (Walker) 

C. limoniiformis (Horn) 

C. lobata (Eschscholtz) 

telum (LeConLe) 

C. lutescens (FaIl) 

C. maura (LeConte) 

C. mediana (Germar) 

r ubid.ipennis (LeConte) 

C. mendax (LeConte) 

AK YK NT 

AK BC AB 


AK YK 

AK 

-YK 

AK 

AK 

AI{ YK 

AK YK 

AK 

AK YK 

NT 

BC 

ON 

NS 

BC AB SK MB ON PQ NS 

BC 

BC 

BC 

AB 

NT BC AB 

_BCAB 

_BCAB 

_BC 

_BC 

_BC 

-BC 

_BC 

AB 

_BCAB 

Nr sc 

NT BC 

_BC 

_BC 

NT 

_BC 

_BC 

NT 

BC 

AB 

AB 

AB 

AB 

SK 

SK MB 

SK MB 

SK MB 

SK MB 

SK 

TSK 

AB SK 

AB SK 

AB SK 

_MB 

SK MB 

SK MB 

_ONPQNBNS 


MB 

ON PQ NB 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ftfS ON 

-oN 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

pe 

p(l 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NB 

NS PE 

NS 

NS PE 

NS-- 

NS 

NS 

NS-_ 

NS 

NS 

NS 

NB NS 

-? 

_NF 

NF 

NF 

NF 

NF 

NF 

NF 

179

Elateridae 

C. moerens. tLeConte) 

C. monLanrr (W.J. Brownl 

C. morula rl,eConte) AK 

brunnipes (Bland) 

C. nebraskensis t Bland) 

C. nigricans tFall t 

C. nigricollis (Bland) AK 

C. nitidula (LeConte) AK 

C. ochreipennis (LeOonte) AK 

C. opacula {Leconte) 

serricornis (Mannerheim) 

C. propola columbiana (W.J. Brown) AK 

C. propola propola (LeConte) 

furcifer (LeConte) 

C. protracta { Leconte) 

C. pruininir tHornt 

no:rlo iHyslop) 

pruinosa (Schwarz) 

pruinosula (Schwarz) 

C. pudica ( W.J. Brown ) 

C. pulchra tLeconte) 

festiua (LeConte) 

C. pygmaert tVan Dyket 

C. pyrrhos {Herbst} 

aequa,',is (Melsheimer) 

pyrrhica (Haldeman) 

uagralLs (Melsheimer) 

C. respleniens aeraria (Randail) ? 

junAi , Schenkling) 

racinei (Chevrolal) 

uiridi.. (Germar) 

C. resplencens resplendens AK 

(Esr:hscholtz) 

C. rotundirollis r Sayt 

russicollis (Germar) 

stictico (Germar) 

C. rufopleuralis tFall t 

C. rupestri r {Germarr 

C. sagitticc,llis (Eschscholtz) 

C. semimel allica (Walker) 

C. sexualis tW.J. Brown.) 

C. silvatica Van Dyke 

C. sjaelandica (O.F. MtiLiler)t 

uiridis (Say') 

C. spinosa ' 

LeContet 

C. spf enderrs lZieglert 

C. stricklar:Ldi (W.J. Brown) AK 

C. suckleyi suckleyi (LeConte) 

C. sulcicoll:s tSay) 

parallela (Say) 

C. tarsalis Melsheimer; 

C. tristis tCandbzeJ 

C. triundulata (Randall) AK 

C. uliginostt tVan Dyket 

C. umbrico a (Eschscholtz) AK 

rzdis (Motschulsky) 

C. umbripennis (LeConte) 

gracilior (LeConte) 

n ub il ipennis (LeConte) 

C. vernalis (Hentz) 

C. vidua tS'.J. Brown) 

C. volitans (Eschscholtz) AK 

uaria (W .J. Brown) 

C. vulnerata (LeConte) 

C. watsoni W.J. Brown) 

C. weidtii t.\ngellr 

Genus OXI'GONUS LeConte 

Key to North American species: Roache (1963) 

O. montanus Schaeffer 

YK 

YK 

YK 

_BC 

_BCAB 

NT BC AB 

_BCAB 

_BC 

_BCAB 

NT BC AB 

NT BC AB 

_BC 

NT 

BC AB 

BC AB 

BC 

BC 

SK MB ON PQ 

-nneoNpe- 


:"yuoNPQ_ 


NTQ 

_BCAB 



-oN 

_BC 

_BC 

_BCAB 

_BCAB 

AB 

DF 

PE 

PE 

NS PE 

_ _ _ ABSXMBSNiSNBNS_ _ 

_NTBCABSKMBON 

BC--laeoNpo---- 

_NT 

NT? 

NT BC 

BC 

BC AB 

BC AB 

BC AB 

BC AB 

BC 

nc-- 

AB 

AB 

SK 

ON PQ 

_PQ 

_PQ 

NS 


SK 

ON 

_yuoNPQ 

ix- - - 

oN PQ 

-oNPQ 

MB ON PQ 

PQ 

NB 

NU 

NB NS 

NB NS 

NS 

NS 

NB NS 

LB 

PE LB 

LB NF 

LB NF 

LB 

LB 

NF 

NF 

NF 

NF 

180

Elateridae 

O. obesus (Say) 

acutipennis (Randall) 

MB ON 

PQ 

Genus EANUS LeConte 

P ar anom u s Kiesenwetter 

E. albertanus W.J. Brown 

E. decoratus (Mannerheim) 

paruicollis (Mannerheim) 

uagus (LeConte) 

E. estriatus (LeConte) 

subarcticus W.J. Brown 

E. granicollis (Van Dyke) 

E. maculipennis LeConte 

pictus (Canddze) 

E. striatipennis W.J. Brown 

_BCAB 


AK YK 

AK 

NT BC 

_BC 

BC 

AB 

MB ON PQ 

MB ON PQ 

ON PQ 

NB NS 

NB 

LB NF 

LB NF 

LB NF 

Genus HEMICREPIDIUS Germar 

Asaphes l\trby 

H. bilobatus (Say) 

c au ifron s (Melsheimer) 

c on s ent ane u s (LeOonte) 

mel ano p ht hahn us (Melsheimer) 

planatus (LeOonte) 

tener (LeConte) 

H. brevicollis (Candbze) 

H. carbonatus (LeConte) 

coracinus Candbze 

H. dilaticollis Motschulsky 

H. hemipodus (Say) 

aeneolus (Melsheimer) 

oereus (Melsheimer) 

corporosus (Germar) 

decoloratus (Say) 

H. memnonius (Herbst) 

baridius (Say) 

ruficornis (Kirby) 

thomasi Germar 

H. montanus Lane 

H. morio (LeConte) 

uernus Canddze 

H. pallidipennis (Mannerheim) 

oregonus (LeConte) 

H. simplex (LeConte) 

H. tumescens (LeConte) 

AK 

BC 

BC 

BC 

BC 

BC 

_BC 

_BC 

AB 

AB 

AB 

SK 

SK 

ON 

MB ON 

MB 

PQ 

ON PQ 

MB ON PQ 

NB NS 

NS 

PE 

Genus OESTODES LeConte 

O. puncticollis Horn 

O. tenuicollis (Randall) 

g r ac ilifor mis (RandalI) 

obscuripes Candbze 

Tlibe Oestodini 

AB 

SK MB 

ON PQ NB NS PE 

Genus ELATER Linn6 

Ludius Berthold 

Steatoderus Dejean 

E. abruptus Say 

coracinus (Germar) 

Genus PARALLELOSTETHUS Schwarz 

P. attenuatus (Say) 

/uscus Castelnau 

Subfamily EI"ATERINAE 

Tribe Elaterini 

Key to North American species: Roache (1961) 

oN PQ 

ON 

NS 

181

Elateridae 

Genus SERICUS Eschcholtz 

Atractooterus LeConte 

Dolerosomus Motschulsky 

Sericosomus Dejean 

S. blaisdelli (Van Dyke) 

S. honestus (Randall) 

fusiforrzis (LeConte) 

S. incongrurLs (LeConte) 

behren::i (Canddze) 

S. silaceus (tjay) 


flauipennis (Motschulsky) 

umbraticus (LeConte) 

S. viridanus (Say) 

sublucens (Randall) 

AK YK 

BC 

NT BC 

SK 

-oN 

MB ON 

-oN 

PQ 

PQ 

PQ 

ON PQ 

NB 

NB 

NS 

NS 

LB 

NF 

Tlibe Agriotini 

Genus GLY?HONYX Canddze 

Key to Ncrth American species: Smith and Balsbaugh (1984) 

G. testaceus (Melsheimer) 

ON 

Genus AGRTOTES Eschscholtz 

Pedetes l{trby 

Key to N< rth American species: 

A. apicalis l,eConte 

A. arcanus \V.J. Brown 

A. avulsus (.LeConte) 

A. collaris (LeConte) 

A. criddlei Van Dyke 

A. fer"rugineipennis (LeConte) 

A. fucosus (],eConte) 

A. hoodi Bec:ker 

A. isabellinus (Melsheimer) 

A. limosus ( LeOonte) 

A. Iineatus tLinn6X 

A. mancus (Say) 

striatutus Melsheimer 

A. oblongicollis (Melsheimer) 

A. obscurus (Linn6)t 

A. opaculus (LeConte) 

montartus LeConte 

A. pubescens Melsheimer 

A. quebecensis W.J. Brown 

A. sparsus LeConte 

A. sputator il-inn6)f 

A. stabilis (l,eConte) 

A. tardus W.J. Brown 

A. thevenetl,i Horn 

Becker (1956) 

_NT 

--sc- 

BC 

SC AB 

BC AB 

_AB 

BC 

BC AB 

BC 

_AB 

BC 

BC AB 

BC 

BC AB 

BC 

ix 

SK 

SK 

SK 

-o* 

-oN 

-oN 

MB 

MB ON 

-on 

MB ON 

MB ON 

_9^ 

PQ 

pe 

PQ 

PQ 

PQ 

PQ 

_MBONPQ 

_ yuoNPQ 

SK MB ON PQ 

____ 

v 

NB 

NE 

NB 

-lt 

NB 

_NS 

NB NS 

v 

NSpe-- 

NS--Nr 

Ns pn is Nr 

NS 

NS PE LB 

NF 

NF 

Genus AGRIOTELLA W.J. Brown 

A. bigeminata (Randall) 

A. columbiana W.J. Brown 

A. occidentzrlis W.J. Brown 

BC 

BC A3 

yu 9* PQ 

YNS 

Genus DALOPIUS Eschscholtz 

D. agnellus W.J. Brown 

D. asellus W.J. Brown 

D. brevicornis W.J. Brown 

D. cognatus W.J. Brown 

D. corvinus W.J. Brown 

D. fucatus W.J. Brown 

D. fuscipes W.J. Brown 

D. gartrelli W.J. Brown 

D. gentilis W.J. Brown 

D. inordinar;us W.J. Brown 

D. insolens W.J. Brown 

D. insolitus W.J. Brown 

- - scTsx- a*;: 

BC 

BC AB 

BC AB 

--scABSf-9*PQ 

-_l"T--oNpe 

oN PQ 

_PQ 

Iu_ 

NB NS 

-v 

r82

Elateridae 

D. insulanus W.J. Brown 

D. maritimus W.J. Brown 

D. mirabilis W.J. Brown 

D. pallidus W.J. Brown 

D. parvulus W.J. Brown 

D. spretus W.J. Brown 

D. suspectus W.J. Brown 

D. tristis W.J. Brown 

D. vagus W.J. Brown 

D. vernus W.J. Brown 

Genus AMPEDUS Dejean 

A. antracinus (LeConte) 

A. apicatus (Say) 

melanopygus Germar 

A. areolatus (Say) 

obliquus (Say) 

scitulus Germar 

A. atripennis (Horn) 

u iolaceipennis (Candbze) 

A. behrensi (Horn) 

A. brevis (Van Dyke) 

A. collaris (Say) 

A. deletus (LeConte) 

A. dimidiatus (LeConte) 

A. evansi W.J. Brown 

A. fuscatus (Melsheimer) 

A. fusculus (LeConte) 

A. impolitus (Melsheimer) 

A. laesus (LeConte) 

A. laurentinus W.J. Brown 

A. linteus (Say) 

lugubris Germar 

A. luctuosus (LeConte) 

A. melanotoides W.J. Brown 

A. melinus (LeConte) 

A. melsheimeri (Leng) 

hepaticus (Melsheimer) 

A. militaris (Harris) 

A. miniipennis (LeConte) 

A. mixtus (Herbst) 

A. moerens (LeConte) 

A. molestus (LeConte) 

A. nigricans Germar 

A. nigricollis (Herbst) 

A. nigrinus (Herbst)* 

A. oblessus (Say) 

sellatus (Leng) 

discoideus (Fabricius) 

A. occidentalis Lane 

bimaculatus (Van Dyke) 

A. pedalis Germar 

rufipes (Palisot de Beauvois) 

A. phelpsi (Horn) 

hoppingi (Van Dyke) 

A. phoenicopterus Germar 

A. protervus (LeConte) 

A. pullus Germar 

Iacustris (LeConte) 

A. pusio Germar 

luteolus (LeConte) 

A. quebecensis W.J. Brown 

A. rhodopus (LeConte) 

A. rubricollis (Herbst) 

socer (LeConte) 

uerticinus (Palisot de Beauvois) 

A. rubricus (Say) 

A. sanguinipennis (Say) 

- 

-- 

_BC 

_BC 

ABSKMB_PQ 

_BCABSKMBONPQ 

- ec - :" 

_BC 

MB- 

_ - -:"-:"ilB3Nia 

AK_ 

T_ 

AK 

NS- 

AK- 

_v 

AK YK 

Tribe Ampedini 

-BC 

NT BC AB SK nfS O* 

-oN 

_BC 

_BC 

_BC 

NT BC 

_BC 

NT_ 

Nr sc 

NT 

-BC 

AK YK NT 

T 

- 

NS- 

NS 

NS 

Nr- AB- - SNFBNn- - 


NT BC AB 

-BCMBONPQNBNS 


-oNPQ 



NT 

BC 

BC 

BC 

BC 

AB 

AB SK 

-s* 

AB SK 

AB_ 

YSK 

_SK 

AB 

AB 

AB 

SK 

MB ON 

-oN 

MB ON 

MB ON 

-oN 

MB 

-oN 

PQ 

PQ 

___ 

MB_PQ 

MB ON PQ 

MB ON PQ 

-oNPQ 

MB ON PQ 

_ONPQ 

-oN 

MB ON PQ 

_ONPQ 

PQ 

PQ 

PQ 

PQ 

---o*pe 


BC AB SK 

BC 

PQ 

PQ 

NB 

v 

NB 

NB 

NB 

NS 

NB 

NB 

\Tq 

NS 

DF 

NB NS PE 

NS_ 

NS 

NS 

NS 

_NF 

- lrr 

is N" 

iey 

_NF 

LB NF 

ie Ne 

_NF 

:"y 

-N" 

LB NF 

- - - - MB3N|8- N3- - - 

183

Elateridae 

A. sayi (Let)onter 

- ON 

A. semicinrtus rRandall) - ON PQ NS 

A. subtilis tleOonte) - BC AB SK MB ON PQ - NS 

A. uteanus (Van Dyker - SK 

A. varipilis (Van Dyket - \'K NT BC AB 

columoianus W.J. Brown 

A. vitiosus LeContet - ON PQ NB 

Genus ME(IAPENTHES Kiesenwetter 

Dolopt osomus Motschulsky 

Key to Nrrth American species: Becker (19?1) 

M. caprella tl-eContet AK BC 

califor nicus W.J. Brown 

M. gentner Lane - BC 

M. nigriventris LeConte - BC 

M. rogersi }{orn ON PQ NB 

M. solitariL.s Fall PQ 

M. stigmosus (LeConte) BC AB SK MB ON PQ NB 

M. tartareus (Lecontej - BC 

Genus ANC HASTUS LeConte 

B r ac h2,' cre p i s LeConte 

Crepiaotritus LeConte 

Elatrh,.us Horn 

Tribe Physorhini 

A. anthrax Horn.) - BC 

coruzs Van Dyke 

militaris Candbze 

A. cinereipt nnis tEschscholtzr 

hir s utul us Motschulsky 

- BC AB 

recedens LeConte 

p ilifer t.: s Motschulsky 

puberu.lus (Mannerheim) 

regularis Motschulsky 

ta nt il I t.t s Mannerheim) 

( 

Genus MEL,ANOTUS Eschscholtz 

Cratonychus Dejean 

Ctenon,ychus Stephens 

Perimecus Stephens 

Key to North American species: Quate and Thompson (1967) 

M. americanus (Herbst) 

ang u st.a tu s (Erichson) 

M. castanipes (Paykull)t 

inaequ,zlis (LeConte) 

paradaxus Melsheimer 

scrobiccllis (LeConte) 

texanu:; Canddze 

M. communis (Gyllenhal) 

pag(tnt,ts Canddze 

M. corticinus (Say) 

g I andieo lor Melsheimer 

uetulus (Erichson) 

M. decumar:Lus (Erichson) 

canadensis Candbze 

cuneo.tus (LeConte) 

incertus (LeConte) 

M. depressus (Melsheimer) 

diu ar c arinus Blatchley 

lixus Blatchley 

p ar um p unc tatas (Melsheimer) 

Tribe Melanotini 

M. hyslopi \/an Zwaluwenburg - ON PQ 

M. ignobilis Melsheimer 

secretu.s (LeConte) 

M. leonardi (LeConte) 

-oN 

MBONPQNBNSPE_NF 

_ONPQ 

rlNT 

_ONPQ_NS 

.rNT 

-oN 

MB ON PQ 

184

Elateridae 

M. longulus oregonensis (LeConte) 

franciscanus Van Dyke 

uariolatus LeConte 

M. miscellus Quate 

M. morosus Canddze 

debilis Blatc}:.ley 

longulus (LeConte) 

simulans Blatchley 

M. prasinus Blatchiey 

M. sagittarius (LeConte) 

carinus Blatchley 

M. similis (Kirby) 

flssilis (Say) 

exuberans (LeConte) 

oc hr acei p ennis (Melsheimer) 

s p henoidal is (Melsheimer) 

M. trapezoideus (LeConte) 

BC 

_SK 

-oN 

-oN 

_9* 

MB ON 

ON 

PQ 

PQ 

PQ 

NB NS 

Genus CARDIOPHORUS Eschscholtz 

C. acutus Lanchester 

C. alpestris Lanchester 

C. brunnipennis Lanchester 

C. cardisce (Say) 

dejeanii LeConte 

C. communius Lanchester 

C. convexulus LeOonte 

C. convexus (Say) 

amictus Melsheimer 

insulsus Canddze 

C. edwardsi Horn 

C. fenestratus LeConte 

larquini Candbze 

C. gagates Erichson 

laeuicollis Erichson 

C. kooskooskiensis Lanchester 

C. Iatiusculus Eschscholtz 

C. montanus Bland 

C. nevadensis Blanchard 

C. plebejus Lanchester 

C. plexus Lanchester 

C. pratensus Lanchester 

C. propinquus Lanchester 

C. pubescens Blanchard 

C. silvanus Lanchester 

C. sodalis Lanchester 

C. tenebrosus LeConte 

fuluipes LeConte 

C. tumidicollis LeConte 

C. tutus Lanchester 

Genus HORISTONOTUS Canddze 

H. curiatus (Say) 

areolatus (Erichson) 

Subfamily CARDIOPHORINAE 

_AB 

BC 

BC 

_AB 

BC 

_BC 

NT BC AB 

SK 

BC AB 

BC 

_AB 

BC 

BC 

BC 

_AB 

BC 

BC AB SK 

_AB 

_AB 

BC AB SK 

BC AB 

BC AB 

MB ON PQ 

--oNpeNe---- 

ON PQ 

MB ON 

MB 

MB 

ON 

PQ 

NB NS 

NB NS 

185

Eucnemidae: 

FamilyTHROSCIDAE throscidbeetles 

Y. Bousquet 

Throscidae includes about 30 North American 

species, of which nine have been reported in 

Canada. The adults, which look like small, stout 

elaterids, are usually found on foliage and flowers. 

The larvae occur in litter, in decaying wood, and 

under bark. 

Few authors agree on the position of the lissomines. 

Crowson (1961) included Drapetes in 

Elateridae and Lissomus in Throscidae; Burakowski 

(1975) el:vated the group to family level; Lohse 

(1979) treated the lissomines as a subfamily of 

Eucnemidae; Lawrence (1982a) listed them in 

Elateridae. I have followed Yensen (1975) in considering 

the lissomines as a subfamily of 

Throscidae. Yensen (1975) provided a key to the 

extant genera of the family and revised t;he genus 

Trixagus. 

NT (2); BC (6); AB (1); MB (1); ON (5); PQ (5); NB 

(2);NS (2);NF (1) 

Subfamily 

LISSOMINAE 

Genus DRAIIETES Dejean 

D. gerninatus (Say) 

ame r icc,nus ( Gerstaecker) 

basalis (Randal1) 

exstriatus (Say) 

oN PQ 

Subfamily THROSCINAE 

Genus TRD(AGUS Kugelann 

Throsct.Ls Latreille 

Key to North American species: Yensen (1975) 

T. carinicoll s tSchaeffer.t 

T. chevrolat {Bonvouloirj 

paruulu.s (LeConte) 

sejunctus (Hotn) 

T. mendax (.lorn) 

T. sericeus ( LeConte; 

debilis t}{orn) 

_BC 

_BC 

NT BC 

NT BC 

AB 

ON 

ON 

PQ 

PQ 

NB 

NS 

NF 

Genus AULONOTHROSCUS Horn 

A. constrictor (Say) 

A. punctatus (Bonvouloirt 

A. validus (LeConte) 

_BC 

MB ON 

_9* 

PQ 

PQ 

NB NS 

Genus PACI]IOPUS LeConte 

P. hornii (LeConte) 

fuchsi Casey 

BC 

Family EUCNEMIDAE false click beetles 

Y. Bousquet 

Thirty-two species of eucnemids have been 

reported in Canada and Alaska. The adults occur 

mainly on vegetation and under bark but are rather 

infrequently collected. The larvae bore into various 

hardwoods that are usually rotten. Like elaterids, 

adults have the clicking mechanism, but few species 

have been actually observed to click. 

The taxonomy of the North American species of 

Eucnemidae has been poorly studied, and therefore 

many changes and additions to the Canad.i.an fauna 

can be expected. Horn's revision (1886) is the latest 

one for the North American species. 

AK (1); BC (13); AB (2); SK (1); MB (3); ON (18); PQ 

(20);NB (5);NS (3);NF (1) 

186

Eucnemidae 

Genus STETHON LeOonte 

Subfamily EUCNEMINAE 

S. pectorosus LeConte - ON PQ 

Genus DELTOMETOPUS Bonvouloir 

D. amoenicornis tSay) 

clypeatus (Say) 

- ON PQ NB NS 

ereptus Bonvouloir 

D. rufipes (Melsheimert 

Genus DROMAEOLUS Kiesenwetter 

D. basalis (LeContet - BC 

D. californicus Bonvouloir - BC 

D. cylindricollis (Sayt - ON PQ 

D. harringtoni Horn MB ON PQ NB 

D. nitens Horn - BC 

Genus FORNAX Castelnau 

Isarl hrus LeConte 

Onichodon Newman 

F. badius (Melsheimer.t - ON PQ 

basalis Hotn 

nouitius (Bonvouloir) 

F. bicolor (Melsheimer) PQ 

luridus Chevrolat 

F. calcealus tSay) - ON PQ 

spretus (LeConte) 

F. canadensis W.J. Brown - ON PQ NB NS 

F. hornii Bonvouloir - ON PQ 

F. orchesides (Newman) - ON PQ 

Genus PHAENOCERUS Bonvouloir 

P. americanus Horn - BC 

Genus DIRHAGUS Latreille 

Microrhagus Dejean 

Subfamily DIRIIAGINAE 

Key to genera: Cobos (1964) 

D. pectinatus (LeConte) 

D. subsinuatus (LeConte) 

- BC - ON PQ NB 

MB ON PQ 

imperfectus (LeConte) 

oblilus (Bonvouloir) 

D. triangularis tSayt - ON PQ 

Genus ENTOMOPHTTIALMUS Bonvouloir 

E. rufiolus (LeConte) - ON PQ 

pallens Bonvouloir 

Genus SARPEDON Bonvouloir 

S. scabrosus Bonvouloir - BC - ON PQ 

Genus RHAGOMICRUS Fleutiaux 

R. humeralis { Say) PQ 

meticulos us (Bonvouloir) 

Genus ISORHIPIS Boisduval & Lacordairei 

Subfamily MELASINAE 

I. obliqua (Say) - BC - ON PQ NB NS 

Genus MELASIS Olivier 

M. rufipennis Horn - BC 

M. tsugae Hopping - BC 

187

Lycidae 

Genus EPIPHANIS Eschscholtz 

E. cornutus Eschscholtz AK BC - SK - ON PQ NB - NF 

cristatus LeConte 

Genus SCHI ZOPHILUS Bonvouloir 

S. subrufus Randall,l 

sirnplet (LeConte) 

trilobatus Bonvouloir 

Genus HYLI.S des Gozis 

H. frontosus tSayt 

canaliculatus (LeConte) 

H. terminalrs (Leconte, - BC AB SK - ON PQ 

Genus XYLOPHILUS Mannerheim 

Xylobius Latreille 

Xyloecus Dejean 

X. cylindriformis lHornt - BC 

Genus NEMATODES Berthold 

Hypocozlus Dejean 

N. atropos {l;ay} MB 

N. penetran; (Leconte) - ON PQ 

Genus ANE,,ASTES Kirby 

A. druryi Kirby 

brunne.rc (Latreille) 

- BC AB 

aroszs (Say) 

latreille'i LeConte 

Superfamily C antharoidea 

Family LYCIDAE net-winged beetles 

Twent;y-nine species of Lycidae are recorded in 

J. McNamara 

tions; his paper of 1949 includes a key to the 

Canada and Alaska. The adults occur on leaves, on Nearctic tribes. 

flowers, or under bark and are thought to be predaceous. 

The larvae are predaceous, and those of 

most species are found under bark. 

Greer, has revised the taxonomy of the North AK (3); yK (1); NT (1); BC (8); AB (5); SK (a); MB 

American species of Lycidae in a series of publica- (8); ON (25); PQ (19);NB (5);NS (5); LB (1); NF (1) 

Genus CAI-OCHROMUS Gu6rin-M6neville 

C. dimidiatr,is (LeConte) 

C. fervens LeConte 

C. perfacetus (Say) 

substritttus (LeConte) 

Subfamily LYCINAE 

Tlibe Lygistopterini 

Key to North American species: Green (1950) 

-BC 

---:"Tsx-o*pe----- 

Genus DICTYOPTERUS Mulsant 

D. aurora (Flerbst) 

coccinatus (Say) 

nigripes (Schaeffer) 

Tlibe Platerodini 

Key to North American species: Green (1951, 1953) 

AK YK NT BC AB _ MB ON PQ NB NS _ LB NF 

188

Lycidae 

D. hamatus Mannerheim 

D. simplicipes Mannerheim 

loelzs Motschulsky 

D. thoracicus (Randall) 

praefectus (Newman) 

AK 

AK 

BC 

BC AB 

BC AB 

SK 

MB ON 

PQ 

Genus EROS Newman 

E. humeralis (Fabricius) 

incestus LeConte 

obliquus (.Say) 

ON 

PQ 

Genus PLATYCIS Thomson 

Erotid,es Waterhouse 

P. sculptilis (Say) 

axill aris (Melsheimer) 

oblira (Newman) 

ON 

PQ 

Genus LOPHEROS LeConte 

L. crenatus (Germar) 

cruciatus (Randall) 

L. fraternus (Randall) 

MB ON PQ 

-oNPQ 

Genus EROPTERUS Green 

E. arculus Green 

E. trilineatus (Melsheimer) 

_ONPQ 

_ONPQ 

Genus PLATEROS Bourgeois 

Neoceletes Leng & Shoemaker 

P. batillifer Green 

P. bispiculatus Green 

P. borealis Green 

P. centralis Green 

P. fl avoscutellatus Blatchley 

P. floralis (Melsheimer) 

P. lictor (Newman) 

californicus Van Dyke 

columbiensis W.J. Brown 

crateracollis (Leng & Shoemaker) 

P. peregrinus Green 

P. subfurcatus Green 

P. timidus (LeConte) 

P. volatus Green 

-oN 

-oNPQ 

-oN 

-oN 

-oNPQ 

- ecABsxmn3Ni3 

_BC 

oN PQ 

ON 

ON 

ON PQ 

NB 

NB 

NS 

NS 

Tlibe Calopterini 


Genus CAI-OPTERON Gu6rin-M6neville 

Diagrapha Newman 

C. discrepans (Newman) 

C. reticulatum (Fabricius) 

affinis (LeConte) 

retiferum LeConte 

typicum (Newman) 

C. terminale (Say) 

apicale (LeConte) 

diuisum (Newman) 

dorsale (Newman) 

duplicatum (Haldeman) 

- - - yuSN 

PQ 

MB ON PQ 

Genus CAXNIA Newman 

Caeniella Cockerell 

C. dimidiata (Fabricius) 

scapularis Newman 

MB ON PQ 

NB NS 

Genus CELETES Newman 

C. basalis LeConte 

mystacina LeConte 

tabida LeConte 

SK 

MB ON PQ 

NB NS 

189

Lampyrrida e 

FamiIyPHENGODIDAE glowworrns 

J. McNq.mara 

Only one species of Phengodidae occurs in 

Canada. The larvae of most species are luminescent, 

hertce the vernacular name glowworms. The 

larvae fe,,ed on soft-bodied insects and other small 

organisrls. The females are larviform but have 

compound, instead of simple, eyes. Adults and larvae 

occur in leaf litter and under logs, bark, and 

debris on the ground. The males are al;tracted to 

light 

Ẇittmer .1975) revised the North American 

species of the genus Phengodes, the only genus 

occurring in Canada. 

oN (1) 

Genus PHIINGODES Illiger 

Key to North American species: Wittmer (1975) 

(Subgenus PHENGODES s. str.) 

P. plumosa tOliviert ON 

Familv LAMPYRIDAE fireflies 

J. McNamara 

Twenty-nine species of Lampyridae are 

recorded in Canada. Members of this family are 

fairly well known because of their ability to produce 

flashing light. Species may be recognized in flight 

by the color of the light as well as by the number 

and the length of flashes. This flashing behavior 

enables l;he sexes to frnd each other. All larvae and 

most adults are luminescent. The larvae are predaceous 

and feed on other insects, larvae. snails, and 

slugs. The feeding habits of the adults are not well 

known. 

Many genera of lampyrids have been revised in 

the past few decades, but a thorough revision ofthe 

family would be very useful. Green (1948) published 

a revision of the Nearctic genera. 

xT (3); BC (9);AB (3); SK (5); MB (12); ON (19); PQ 

(14);NB (8);NS (10);PE (1);NF (3) ' I (1) 

Subfamily LAMPYRINAE 

Tribe Cratomorphini 

Genus PYI|ACTOMENA Melsheimer 

Lecontea E. Olivier 


P. angulata (Say) 

flauoc:,.ncta LeConte 

P. borealis (Randall) 

P. dispersa Green 

P. linearis LeConte 

P. lucifera Melsheimer 

P. sinuata Green 

NT 

NT 

AB 

SK 

SK 

SK 

MB ON 

MB 

MB 

MB 

MB 

ON 

ON 

ON 

PQ 

PQ 

PQ 

NB 

NB 

NS 

NS 

T?ibe Photinini 

Genus PHOTINUS Castelnau 

Key to N'orth American species 

P. aquilonius Lloyd 

P. ardens LeConte 

taedif'zr LeConte 

P. consimi]is Green 

P. ignitus llall 

P. indictus (LeConte) 

ablucens Fall 

P. marginellus LeConte 

castus LeCor,:te 

Green (1956) 

MB ON 

MB ON 

-oN 

-oN 

-oN 

-oN 

PQ_NS 

PQ NB NS 

PQ 

NF 

190

Lampyridae 

P. obscurellus LeConte 

frigidus Olivier 

P. punctulatus LeConte 

P. scintillans (Sayt 

MB ON 

ON 

ON 

PQ 

Genus PHAUSIS LeConte 

Key to North American species: Fender (1966) 

P. nigra Hopping 

P. rhombica Fender 

BC 

BC 

AB- 

Genus ELLYCHNIA LeConte 

Key to western North American species: Fender (1970) 

(Subgenus ELLYCHNIA s. str.) 

E. cornrsca (Linn6) 

E. facula LeConle 

E. greeni Fender 

E. hatchi Fender 

E. lacustris LeConte 

-BC 

_BC 

_BC 

_BC 

_BC 

y" oN PQ NB NS 

NF 

Genus PYROPYGA Motschul sky 

Keyto species: Green (1961) 

P. decipiens (Har-ris) 

P. nigricans (Say) 

californica Motschulsky 

californica (Gorham) 

fene stralis (Melsheimer) 

reuersus (Gemminger) 

NrscABixnrnSNiSNENS - 

-- - 

Genus LUCIDOTA Castelnau 

RileyaE. Oliviet 

L. atra (Olivier) MB ON PQ NB NS 

laticor nis (Fabricius) 

morio (Melsheimer) 

tardaLeConte 

Genus PHOSPHAENUS Castelnau 

P. hemipterus (Fourcroy)f PQ 

NS 

Subfamily ANIYDETINAE 

Genus POLLACLASIS Newman 

Polyclasis LeConte 

P. bifaria (Say) 

oualus Newman 

ON 

PQ 

Subfamily PHOTURINAE 

Genus PHOTURIS Dejean 

Py r ec tosoma Motschulsky 

Key to North American species: Barber (1951) 

P. fairchildi Barber 

P. pennsylvanica (DeGeer) 

marginata (Panzer) 

u ittiger a (Motschulsky) 

BC_SKMBON 

PQ 

NB 

NS 

NS PE 

NF 

191

Cantharid,re 

Family CANTHARIDAE soldier beetles 

J. McNamqra 

Cantharidae includes 126 species and subspecies 

in Canarla and Alaska. The adults are common on 

goldenrc,d, milkweed, and hydrangea, where they 

feed on nectar and pollen. Most larvae are carnivorous, 

gererally preying on other soft-bodied insects; a 

few are cmnivorous, feeding on plants such as wheat, 

potatoes. celery, and other plants. They occur mostly 

beneath bark and debris. 

Some of the genera of Cantharidae are poorly 

known taxonomically, including the large genus 

Podabrus, and are in need ofrevision. Fender (1962) 

provides keys to the species of the Pacifrc Northwest. 

aK (15); vK (8); NT (13); BC (57); AB (20); SK (9); 

MB (22); ON (a8); PQ (48); NB (22); NS (rl5); PE (1); 

LB (5); NF (16) . I(2) 

Subfamily Cantharinae 

Tlibe Podabrini 

Genus PODABRUS Westwood 

Brachynotus I\irl:y 

Malthacus l\trby 

Key to Irorth American species 

P. alexand,rri Fender 

P. brevicol,is Fall 

P. cavicolli; cavicollis LeConte 

P. conspiratus Fall 

P. deceptus W.J. Brown 

P. diadema (Fabricius) 

P. excursur; excursus Fall 

P. extremus LeConte 

P. falli Hop,ping 

P. fissilis Fall 

P. flavicollis LeConte 

P. frater LeConte 

q uadr,.collis ( Motschul sky) 

P. frosti Fender 

P. fumiganus Green 

P. furtivus Fall 

P. heteronl'chus Fall 

P. instabilis Fall 

P. intrusus Green 

P. laevicoil s (Kirby) 

P. limbellur LeConte 

P. lutosus LeConte 

P. lygarius Fender 

P. melvillei Fender 

P. modestus (Say) 

P. nothoides LeConte 

P. obscurevittatus Fall 

P. pattoni LeConte 

P. perplexus W.J. Brown 

P. piniphilus (Eschscholtz) 

P. planulus Green 

P. probus FaIl 

P. protensu s LeConte 

P. pruinosts atrocervicus Fender 

P. pruinosr;.s diversipes Fall 

P. puberulus LeOonte 

P. punctatuLs LeConte 

P. puncticollis (Kirby) 

marginellus LeConte 

P. punctula.tus LeConte 

P. pustulatus Fender 

P. pygmaeus Green 

P. rossi Fender 

P. rugosulus LeConte 

p ar u icol lis (Motschulsky) 

P. scaber LeConte 

Fa]l (1927) 

AK 

AK 

AK 

AK 

TY 

AK YK 

_YK 

AK 

AK 

vy 

\.K NT 

_AB 

BC 

BC 

BC 

_BC 

NT BC 

_ IJU 

Nt sc 

_BC 

_BC 

_BC 

NT BC 

NT 

NT BC 

_BC 

_BC 

_BC 

_BC 

BC AB 

BCAB-MB 

BC AB 

BC AB 

MB 

AB SK MB 

AB SK MB 

AB ix 

AB 

AB 

MB 

MB 

MB 

MB 

MB 

MB 

ON 

oN PQ 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

ON 

NB 

_NB 

PQ NB 

PQ 

PQ NB NS 

PQ NB 

PQ NB 

pe-Ns 

PQ 

PQ NB 

PQ 

PQ_NS 

PQ-_ 

PQ NB NS 

PQ NB NS 

PQ NB 

PQ 

PQ 

NS 

PQ NB NS 

l*" 

LB 

_NF 

LB NF 

-Ne 

_Iu 

LB NF 

192

Cantharidae 

P. secretus W.J. Brown 

P. sericatus (Mannerheim) 

P. simplex Couper 

P. smithi Fender 

P. tetragonoderus Fall 

P. tomentosus (Say) 

cinereipennis Motschulsky 

rufiolus (Melsheimer) 

P. tricostatus (Say) 

atripes (Motschulsky) 

benneti (Kirby) 

AK 

AK_ 

_BC 

_BCAB 

na" o* 

-oN 

ON 

PQ 

PQ 

PQ 

IU 

NS 

Tlibe Cantharini 

Genus ANCISTROI.JYCHA Markel 

Key to North American species: McKey-Fender (1950) 

A. bilineata tSayt 

A. dentigera (LeContet 

A. neglecta tFall r 

oNPQ-NS 

oN PQ 

oN PQ 

Genus CANTIIARIS Linn6 

C y r tomo p tila Motschulsky 

Telephorus Schaeffer 

C. alticola (LeConte) 

C. aneba borealis McKey-Fender NT 

C. beta Fender 

C. consors (LeConte) 

tibialis (LeConte) 

tibiella (Gemminger) 

C. curtisi (Kirby) - YK NT 

samouellei (Kirby) 

C. impressa (LeConte) 

C. oregona tLeConte) 

C. rotundicollis Say 

C. rufa Linn6t 

C. transmarina transmarina 

(Motschulsky) 

C. transmarina scopa (LeConte) 

C. tubercu lata ( LeConte) 

armigera (Couper) 

breuicollis (LeConte) 

C. westwoodi tKirby) 

BC 

BC AB SK 

BC 

BC 

BC 

BC 

ec 

BC 

SK 

MB 

MB ON PQ NB 

-oNPQ-NS 

-oNPQNBNS 

-_-9*PQIUNS:' 

PQ 

LB 

NF 

NF 

Genus CYRTOMOPTERA Motschulsky 

C. divisa (LeConter - BC 

latiu sc ula Motschulsky 

Genus RHAXOI.IYCHA Motschul sky 

Key to North American species: McKey-Fender (1950) 

R. bilobata (McKey-Fender) MB 

R. carolina t Fabricius) 

jactata (Say) 

oN PQ 

oN PQ 

Genus RHAGONYCHA Eschscholtz 


R. alaskensis Fender 

R. anthracina Mannerheim 

R. costipennis (LeConte) 

R. excavata (LeConte) 

R. fraxini (Say) 

atra (l\trby) 

binodula Mannerheim 

nigrita (LeConte) 

R. fulva (Scopoli)t 

R. gilvipes (Gemminger) 

dichroa (LeOonte) 

flauipes (LeConte) 

AK 

AK 

AK 

BC 

BC 

-BCPQ 

NT_? 

AB 

-oN------ 

MBONPQ-NS 

MBONPQ_NSNF 

193

Cantharidaer 

R. imbecillis (LeConte) 

albolineat a (Blatchley) 

R. luteicollis (Germar) 

cinctellt (LeConte) 

R. mandibuiaris (Kirby) 

nigritu.la (LeConte) 

R. mollis mc,llis Fall 

nigriceps (LeConte) 

pusilla (LeConte) 

R. nanula (LeConte) 

R. oriflava ( LeConte) 

R. recta (Melsheimer) 

pusilla (LeConte) 

rufipes (Say) 

R. scitula (Say) 

R. septentri,rnis (Green) 

R. syivatica (Green) 

R. tantilla (l,eConte) 

pzsio (l,eConte) 

R. vilis (LeConte) 

R. walshi (LeConte) 

AK YK 

---ecAB-- 

NT BC 

AB 

-SK 

SK 

SK 

MB 

MB 

ON 

ON 

-PQ 

ON 

ON 

oN PQ 

_PQ 

ON 

ON 

_ONPQ 

MB 

NS 

NS 

NS 

NS 

LB 

NF 

NF 

NF 

NF 

Tfibe Silini 

Genus POLItrMIUS LeConte 

P. canadensis W.J. Brown 

P. laticornis (Say) 

dubjzs (Melsheimer) 

incisus LeConte 

inualid,.rs (Say) 

undulatus (LeConte) 

_ONPQ 

fuft_ 

_ _ 

Genus SILII) Charpentier 

Key to Ncrth American species: 

(Subgenus SIILIS s. str.) 

Green (1966) 

S. atra LeC

Derodontidae 

M. columbiensis Fender 

M. fragilis (LeConte) 

trdnsuersus (LeConte) 

M. fuliginosus fuliginosus LeConte 

M. fuliginosus flavicollis Fender 

M. intricatus Fender 

M. macnabi Fender 

M. medioccidens Fender 

M. niger (LeConte) 

M. oregonus Fender 

M. parvulus (LeConte) 

M. piceolus Fall 

M. piperi Fender 

M. similis Fender 

M. stacesmithi Fender 

M. terraenovae Fender 

BC 

ON PQ 

_PQ 

ON 

_PQ 

oN PQ 

ON 

NB NS 

NB 

NN 

NB 

NS 

NS 

NS_- 

NS 

T\TIT 

NT 

NF 

Subfamily CIIAULIOGNATHINAE 

Tlibe Chauliognathini 

Genus CHAULIOGNATHUS Hentz 


C. pensylvanicus (Deceer) 

bimac u la t us (Fabricius) 

MB ON PQ 

Tlibe Ichthyurini 

Genus TRYPHERUS LeConte 

Lygerus Kiesenwetter 


T. frisoni Fender 

ON 

Superfamily Dermestoidea 

FamilyDERODONTIDAE tooth-neckedfungus beetles 

D.E. Bright 

The family Derodontidae includes four genera 

and 19 species of small beetles that occur in the 

temperate parts of both the Northern and Southern 

Hemispheres. They are rarely encountered in the 

freld and are uncommon in collections. Three genera 

and eight species are found in Canada and 

Alaska. With the exception of Laricobius species, 

both larvae and adults of this family feed on various 

kinds of fungi or on the products of fungal 

metabolism. Peltastica species appear to be 

restricted to fermenting sap flows or to areas under 

bark, where bacterial or fungal fermentation takes 

place. Derodontus species feed on the fruiting bodies 

of a wide range of higher Basidiomycetes. The 

beetles prefer fresh fruiting bodies, although they 

have been recorded in rotting mushrooms, "slimy 

fungus," and "watery fungus." Franz (1958) has 

documented the feeding habits of Lqricobius erichsonii 

quite well. Adults and larvae feed on the pine 

and spruce aphids of the family Adelgidae. The preferred 

host of L. erichsonii is the balsam woolly 

aphid; the derodontid was introduced into eastern 

Canada in the 1950s for the control of this pest. 

The other species of Laricobius also feed on other 

species of Adelgidae. 

Information on adult ahd larval morphology, 

feeding behavior, life cycle, and evolution, as well as 

keys and diagnoses ofall genera and species, can be 

found in Lawrence and Hlavac (1979). Lawrence 

(1989) recently published a catalog of the North 

American species. 

AK (2); BC (5); aB (3); ON (2); PQ (1); NB (2); NS 

(3);NF(1) . I(1) 

195

Dermestida,: 

Genus PELi|ASTICA Mannerheim 

P. tubercula.ta Mannerheim 

AK 

Subfamily PELTASTICINAE 

BC 

Subfamily DERODONTINAE 

Genus LARI.COBIUS Rosenhauer 

L. erichsoni Rosenhauert 

L. laticollis Fall 

L. nigrinus Fender 

L. rubidus l,eConte 

_BC 

_BCAB 

---:cT-,oNpe 

YNS 

NB 

NF 

Genus DERCDONTUS LeConte 

D. esotericus Lawrence 

D. maculatu,s (Melsheimer) 

D. trisig:natus (Mannerheim) 

AK 

ec 

AB 

__oN__N3__ 

Family NOSODENDRIDAE wounded'tree beetles 

J.M. Campbell 

Noso

Dermestidae 

murinus LeConte 

nubilus Say 

tessellatus Hetbst 

D. carnivorus Fabriciust 

rnucoreu,s LeConte 

sobrinus LeConte 

uersicolor Castelnau 

D. fasciatus LeConte 

D. frischii Kugelannt 

D. maculatus DeGeert 

elongatus Hope 

lupinus Mannerheim 

truncatus Casey 

uulpinus Fabricius 

D. marmoratus Say 

D. talpinus Mannerheim 

D. tristis Fall 

(Subgenus DERMESTES s. str.) 

D. ater DeGeert 

D. lardarius Linn6f 

D. pulcher LeConte 

D. reductus Kalik 

D. signatus LeConte 

BC 

_ _ _ ncT 

_BC 

_BCAB_MB 


_BCAB 

--NrecAB 

_BC 

_BCAB 

SK MB 

_MB 

ONPQ_NS 

oN PQ 

oNpeI"_ 

_MBONPQ 

SK MB ON PQ 

_MB_PQ 

SK MB 

NB 

NSPE_NF 

T[ibe Attagenini 

Key to North American species: Beal (1970) 

Genus ATTAGENUS Latreille 

A. brunneus Faldermannt 

elongatulus Casey 

extricatus Casey 

A. pellio (Linn6)t 

A. schaefferi hypar Beal 

A. unicolor japonicus Reittert 


A. unicolor unicolor (Brahm)t 

megatoma (Fabricius) 

piceus (Olivier) 

-BC 

_BC 

-BC-SKMBONPQ 

NS 

PE 

NF 

Genus NO\IELSIS Casey 

Genattus Sharp 

(Subgenus PARANOVELSIS Casey) 

N. perplexa (Jayne) 

BC 

Tribe Megatomini 

Key to North American genera: BeaI (1967) 

Genus TROGODERMA Dejean 


T. glabrum (Herbst)t 

boronBeal 

T. inclusum LeConte 

aduena Casey 

brunnescens Casey 

frosti Casey 

nigrescans Casey 


T. ornatum (Say) 

pallipes Ziegler 

paruulum Casey 

procerum Casey 

pusillum LeConte 

serriferum Dalla Torre 

serrigerum Casey 

lorsole Melsheimer 

T. simplex Jayne 

ajar Casey 

pollens Casey 

BC 

BC 

AB 

-SK 

-oN 

MB ON 

ON 

PQ 

PQ 

197

Dermestidae 

T. sinistrurn FaIl AK 

T. sternale sternale Jayne 

caseyi Arrow 

simuh.tns Casey 

uariipes Casey 

T. variabilr'Balliont 

parab: Ie Beal 

scabricennis Casey 

YK NT BC AB 

BC 

_AB 

SK 

SK 

)IB ON PQ 

_ONPQ 

ON 

PE 

Genus REESA Beal 

R. vespulac (Milliront 

NT BC AB SK 

MB ON PQ 

PE 

NF 

Genus MEGATOMA Herbst 


(Subgenus MEGATOMA s. str.) 

M. pubescens (Zetterstedt) 

prolira BeaI 

BC 

(Subgenus PERIMEGATOMA Horn) 

M. ampla ('lasey) 

M. angularis Mannerheim AK 

M. cylindri:a (Kirby) AK 

impre:;sa (Casey) 

montit:ola (Casey) 

M. giffardi (Blaisdel]) AK 

M. leucochlidon Beal 

M. variegar.a tHornt 

guadalupensis (Casey) 

jaynei (Casey) 

neuad;',ca (Casey) 

BC 

YK NT BC AB 

^-ec-- BC AB 

ON 

Genus PStr UDOHADROTOMA Kalik 


P. kaliki B,ral 

P. perversa (Fall) AK 

BC 

BC AB 

Genus DETjIRTHRUS LeConte 


D. stebbinsi Beal 

BC 

Genus CR\?TORHOPALUM Gu6rin-M6neville 

Key to North American species: Beal (1985o) 

C. haemonhoidale (J.E. LeConte) 

C. triste Le Conte 

fuscicl,aue Casey 

modestum Casey 

nigriatrne LeConte 

picicorne LeConte 

-oN 

-oN 

Tribe Anthrenini 

Genus AN|IIHRENUS I.C. Schaeffer 

Key to errstern Nonh American species: 

(Subgenus ANTHRENUS s. str.) 

A. pimpinellae Fabriciust 

conspersus Casey 

lepidus LeConte 

neuad,icus Casey 

obtectus Casey 

occidens Casey 

pictus Casey 

suffusus Casey 

A. scrophu ariae (Linn6)f 

thoracicus Melsheimer 

(Subgenus NATHRENUS Casey) 

Hoebeke et a]. (1985) 

BC 

BC ON PQ NB NS 

198

Bostrichidae 

A. verbasci (Linn6)t 

caseyi Hinton 

destructor Melsheimer 

pistor Casey 

substriatus Casey 

uorax Casey 

BC 

AB 

MB ON PQ 

(Subgenus FLORILINUS Mulsant & Rey) 

A. castaneae Melsheimer 


A. museorum (Linn6)t 

ON 

ON 

PQ NB 

PQ 

NF 

(Subgenus HELOCERUS Mulsant & Rey) 

A. fuscus Oliviert 

oN PQ 

NS 

Genus ORPHILUS Erichson 

Key to North American species: Beal (1985b) 

O. ater Erichson 

O. subnitidus LeConte 

aequalis Casey 

chalybeus Casey 

Tribe Orphilini 

BC 

oN PQ 

Tribe Thylodriini 

Genus THYLODRIAS Motschulsky 

T. contractus Motschulskyt 

NT 

AB 

MB ON PQ 

Superfamily Bostrichoidea 

FamilyBOSTRICHIDAE bostrichidpowder'postbeetles 

J. McNamara 

Bostrichidae comprises 18 species in Canada 

and Alaska. It includes some of the most destructive 

beetles known. Some species cause considerable 

damage to such stored products as dried roots 

and grains. In the tropics these insects are especially 

destructive, injuring felled timber and bamboo. 

The larvae do most of the damage. Some 

species can kill weakened trees; others have the 

habit of tunneling into the gTeen shoots of plants. 

Members of Bostricidae have been widely distributed 

through commerce because of their tendency 

to bore into wood products, such as crating 

and bamboo. 

Fisher (1950) revised the North American 

species of the family. 

AK (1); vK (1); NT (3); BC (8); AB (3); SK (2); MB 

(6); ON (13); PQ (6);NB (1);NS (1) ' I (1); H (1) 

Subfamily PSOINAE 

Tlibe Polycaonini 

Genus MELA.LGUS Dejean 

E ro pioide s Gu6rin-M6n6ville 

Heterarthron Dejean 

M. confertus (LeConte) 

inciszs (LeConte) 

Genus POLYCAON Castelnau 

Alloeocnemis LeConte 

P. stoutii (LeConte) 

ouicollis (LeConte) 

BC 

BC 

199

Bostrichidac 

Genus PSOI, Herbst 

Acrepis LeConte 

Tribe Psoini 

P. quadrisig:rata tHornr - BC 

sexguttt.,.ta Lesne 

Subfamily ENDECATOMINAE 

Genus END.IICATOMUS Me11i6 

E. rugosus (Randall) 

Genus RHYZIOPERTIIA Stephens 

R. dominica ,Fabriciustt 

pusilla (Fabricius) 

Subfamily DINODERINAE 

MB ON PQ 

- BC MB ON pe 

Genus PROSTEPHANUS Lesne 

P. punctatus {Sayr - ON 

Genus STEPHANOPACIIYS Waterhouse 

S. cribratus t LeConter NT BC AB SK MB ON pe 

S. rugosus (Ctlivier) 

- ON NB 

opacus (Casey') 

porcatu! (LeConte) 

S. sobrinus lr)aseyr - BC AB 

S. substriatus (Paykull)* AK YK NT BC AB - MB ON pe - NS 

pacificu:; (Casey) 

Genus AMPI IICERUS LeConte 

(Subgenus AI\.{PHICERUS s. str.) 

Subfamily BOSTRICHINAE 

Tribe Bostrichini 

A. bicaudatu,; (Sayr 

asperico,t lis (Germar) 

NT SK MB ON 

gracilis'Jasey 

serricollt's (Germar) 

Genus LICHIINOPHANES Lesne 

L. armiger {Leconte) MB ON 

L. bicornis (M/ebert 

ON pe 

L. mutchieri Belkin - ON pe 

angustatus (Casey) 

L. truncaticollis tl-eConrer - ON 

Genus SCOBICIA Lesne 

Tribe Xyloperthini 

S. bidentata tHorn r - ON 

S. declivis (LeContet 

- BC 

Genus XYLOBIOPS Casey 

X. basilaris tSayt 

basalis ( Dejean) 

- ON 

h ume ral is (Melsheimer) 

200

Anobiidae 

Family LYCTIDAE lyctid powder'post beetles 

J. McNarnara 

Only six species of lyctids are known to occur in 

Canada. Members of this family are commonly 

called powder-post beetles because of the tendency 

of the larvae to reduce sapwood, especially of hardwoods, 

into a powdery frass. They are found more 

often in recently dried wood rather than in old 

wood. They attack lumber used for furniture, crating, 

hardwood floors, and other purposes. Second 

only to termites, Lyctidae cause the most destruction 

to wood and its products. The adult beetles 

cause further damage to wood by producing exit 

holes when they emerge. The family is worldwide 

in distribution, each region having an indigenous 

fauna plus introduced species. 

Gerberg (1957) has revised the New World 

species. 

BC (4);uB (1); oN (4);PQ (3);NB (1);NS (2) o I 

(2) 

Tribe Lyctini 

Genus LYCTUS Fabricius 

L. brunneus (Stephens)f 


L. cavicollis LeConte 

L. linearis (Goeze)f 

arillaris Melsheimer 

striatus Melsheimer 

L. opaculus LeConte 


L. planicollis LeConte 

BC 

BC 

BC 

BC 

-oN 

MB ON 

PQ 

oN PQ 

PQ 

NB 

NS 

NS 

Genus TROGOXYLON LeConte 

T. parallelopipedum (Melsheimer) 

ON 

Family ANOBIIDAE deathwatch beetles 

J. McNaruara 

Anobiids include 79 species in Canada and 

Alaska. Larvae of most species bore into dead hardwoods 

and softwoods; two species of Ernobius attack 

and destroy pine cones; one species, Anobium punctq.tum 

(furniture beetle), damages woodwork, furniture, 

bookbindings, and similar products. The beetles 

often produce a tapping sound in their burrows, 

which gives rise to the ancient superstition that they 

are a warning of death, hence the common name. 

Some species feed in the young stems or shoots of 

growing trees. Two species, Stegobium paniceum 

(drugstore beetle) and Le.sioderma serricorne 

(cigarette beetle), are serious pests of stored products, 

including drugs, tobacco, seeds, spices, cereal 

products, leather, and museum specimens. 

North American anobiids are relatively well 

known taxonomically, thanks to the work of White, 

who published a key to the genera (1971), a number 

of generic revisions, and a catalog of the North 

American species ( 1982). 

AK (4);YK (6);NT (2); BC (35);AB (18); SK (9); MB 

(21); oN @2);PQ (40);NB (8);NS (7);NF (7) o I 

(5) 

Subfamily DRYOPHILINAE 

Tlibe Eucradini 

Genus EUCRADA LeOonte 

E. humeralis (Melsheimer) 

E. robusta Van Dyke 

_BC 

-oN 

PQ 

201

Anobiidae 

Subfamily ERNOBIINAE 

Tribe Xestobiini 

Genus XESiTOBIUM Motschulsky 

Hyperisus Mulsant & Rey 

Key to North American species: White (1975o) 

X. abietis Fisher 

X. alfine LeConte 

X. gaspensrs R.E. White 

X. marginicolle (LeConter 

BC 

BC 

BC 

-oN 

PQ 

Genus UTOBIUM Fall 


U. elegans iHorn) 

U. granulal,um R.E. White 

U. marmorrtum Fisher 

White (1966, 1976) 

_ v_ 

BCAB_MBONPQ 

BCAB_PQ 

BC 

Tlibe Ernobiini 

Genus EPISERNUS Thomson 

E. trapezoideus (Fall) 

BC 

Genus ERITIOBIUS Thomson 

Philoxylon LeConte 

E. bicolor R.E.White 

E. gentilis Jlall 

E. luteipen.ris LeConte 

E. mollis (Iinn6)+ 

conu et ifrons (Melsheimer) 

E. nigrans ,lall 

E. pallitarsis Fall 

E. punctulatus (LeConte) 

E. schedli ![.J. Brown 

_BC 

_BC 

_BC 

BC 

ON 

ON 

PQ 

PQ_NS 

PQ 

NF 

NF 

NF 

Subfamily ANOBIINAE 

Tribe Euceratocerini 

Genus CTENOBIUM LeConte 

C. antennal;um LeConte 

oN PQ 

Tlibe Gastrallini 

Genus GASTRALLUS Jacquelin du Val 

G. marginipennis LeConte 

BC 

Tlibe Nicobiini 

Genus TRICHODESMA LeConte 

Ptinodzs LeConte 

T. cristata Oasey 

T. gibbosa (Say) 

T. klagesi Fall 

_ _ _:c_ 

_ _ 

ON 

ON 

TYibe Stegobiini 

Genus STETIOBIUM Motschulsky 

S. paniceum (Linn6)t AK 

obesurn (Melsheimer) 

tenuistriatum (Say) 

BC 

SK 

MB ON PQ 

NB 

Genus OLIGOMERUS Redtenbacher 


O. alternan; LeConte 

ON 

PQ 

202

Anobiidae 

O. crestonensis Hatch 

O. obtusus LeConte 

O. sericans (Melsheimer) 

BC 

lo*pe-- 

_ONPQ 

Tribe Anobiini 

Genus PLATYBREGMUS Fisher 

P. canadensis Fisher 

ON 

NS 

Genus ANOBIUM Fabricius 

A. punctatum (DeGeer)f 

pumilum LeConte 

BC 

MB_PQ 

NS 

Genus HEMICOELUS LeConte 

Cacotemnus LeConte 


H. carinatus (Say) 

errons (Melsheimer) 

linearis (LeConte) 

H. defectus (Fall) 

H. gibbicollis (LeConte) 

destructor (Fisher) 

roguensis (Hatch) 

H. pusillus (Fall) 

H. umbrosus (Fall) 

White (1976) 

AK 

AK YK 

_BC 

BC 

BC 

AB 

AB 

SK 

MB ON PQ 

yu oN PQ 

-oNPQ 

MB ON PQ 

NB 

NB 

Genus MICROBREGMA Seidlitz 

M. emarginatum emarginatum 

(Duftschmid)t 

foueatutn (Kirby) 

M. emarginatum granicolle Fall 

BC 

BC 

AB 

SK 

MB ON PQ 

Tribe Hadrobregmini 

Genus HADROBREGMUS C.G. Thomson 

Coelostethus LeConte 

H. americanus (Fall) 

H. notatus t Sayt 

H. quadrulus (Leconte) 

H. truncatus (Fall) 

BC AB 

BC 

BC 

oN PQ 

NS__- 

Genus PRIOBIUM Motschulsky 

Key to North American species: White (1975b) 

P. sericeum (Sayt 

MB ON 

PQ NB NS 

Genus DESMATOGASTER Knutson 

D. subconnata (FaIlt 

YK 

AB 

MB 

PQ 

Genus PTILINUS O.F. Mriller 

P. basalis LeConte 

P. Iobatus Casey 

serricollis Say 

P. pruinosus Casey 

P. rufrcornis Say 

bicolor Melsheimer 

Subfamily PTILININAE 

Tribe Ptilinini 

BC 

YK_BCAB_MBON 

-MBONPQ 

-ABONPQ 

Subfamily XYLETININAE 

Tribe Xyletinini 

NB 

_NS 

Genus VRILLETTA LeConte 

Pseudoxyletinus Pic 

V. decorata Van Dyke 

V. laurentina Fall 

BC 

ON 

PQ 

203

Anobiidae 

Genus EIIIRILLETTA Fall 

Oligor;wodes Fall 


E. brevis R E. White 

E. harrisii lFall) 

E. peltata (Harris) 

White (1985) 

AB 

-oN 

-oNPQ 

MB ON PQ 

Genus XYI ETINUS Latreille 

Key to N crth American species: 

X. bicolor E.E. White 

X. confusus R.E. White 

X. fucatus i,eConte 

X. lugubris LeConte 

X. rotundic'rllis R.E. White 

White (1973, 1977) 

-YK 

AB SK 

NT BC AB 

_BC_SK 

AB SK 

-BCABSK 

MB 

PQ 

MB ON PQ 

yu_:a 

NB 

NF 

Tribe Lasiodermini 

Genus LASIODERMA Stephens 


L. serricorne (Fabricius)t AK 

castanzum Melsheimer 

MB ON 

PQ 

Genus CALYMMADERUS Solier 

Eupac.lus LeConte 

C. nitidus ( LeConte) 

Subfamily DORCATOMINAE 

Tlibe Calymmaderini 

TYibe Petaliini 

Genus PETALIUM LeConte 

Key to N'rrth American species: 

P. bistriatum (Say) 

P. seriaturr Fall 

Ford (1973) 

ON 

ON PQ 

Tribe Prothecini 

Genus SCULPTOTHECA Schilsky 

S. puberula tLeConte) 

ON PQ 

Genus STAGETUS Wollaston 

Key to New Worid species: White (19756) 

S. prolundus (Leconte) 

s triat orc u n ctatzs (LeConte) 

ON PQ 

TYibe Dorcatomini 

Genus DOIi,CATOMA Herbst 


D. falli R.E, White 

D. integra ( Fall) 

D. moderata R.E. White 

D. pallicornis LeConte 

foueata R.E. White 

D. setulosa LeConte 

White (1966) 

_BC 

-SKMBONPQ 

-oN 

MB ON PQ 

oN PQ 

NB 

NF 

Genus CAENOCARA C.G. Thomson 

Tylistus LeOonte 

C. bicoior (Germar) 

C. blanchardi Fall 

C. oculaturrL (Say) 

C. scymnoides LeConte 

C. tenuipalpum Fall 

_BC 

_YKNTBCABSK 

MB ON 

_ONPQ 

MB ON PQ 

_,, 

NE 

NS 

204

Ptinidae 

Genus BYRRHODES LeConte 

B. intermedius (Leconte) PQ 

Genus TRICORYNUS Waterhouse 

C atorama Gu6rin-M6neville 

Hemiptychus LeConte 


Subfamily TRICORYNINAE 

Tribe TYicorynini 

T. borealis (LeContet MB ON 

T. confusus (Fall) PQ 

T. densus (Fall) AB 

rogersi (Hatch) 

T. nigritulus (Lecontet - ON 

T. nubilus tFallt 

schuhi (I].atc]e], 

Family PTINIDAE spider beetles 

Y. Bousquet 

Only 16 ptinid species are found in Canada and Ptinidae, the common name of which refers to 

Alaska, and all but three are introduced on this con- the fact that the adults of many species look like 

tinent. Many of these species are found in mills, small spiders, is closely related to Anobiidae. The 

warehouses, granaries, stores, museums, and hospi- species are divided into two groups that are considtals 

where they feed, in both the adult and larval ered to be subfamilies or tribes. Papp has reviewed 

stages, on a wide variety of products including the North American fauna (1962). Many species 

grain, seeds, and drugs. These species are usually that occur in Canada can be identifred with the key 

minor pests, since they rarely build up large popula- provided by Brown (1940). 

tions. Spider beetles also occur in nests of mammals, 

birds, and bees; they are probably scayengers AK (1); BC (13);AB (5); SK (6); IVIB (7); ON (11); PQ 

that feed on dry animal and vegetable matter. (11);NB (9);NS (10); PE (1); NF (5) . I (13) 

Genus GIBBIUM Scopoli 

Scotias Czenpinski 

Subfamily GIBBIINAE 

G. aequinoctiale Boieldieuf - ON PQ NB NS 

psylloides auct. 

Genus MEZIUM Curtis 

M. affrne Boieldieut - BC - SK MB ON PQ NB NS 

Genus SPHAERICUS Wollaston 

Subfamily PTININAE 

S. gibboides tBoieldieut+ - BC 

Genus TIPNUS C.G. Thomson 

Epauloecus Mulsant & Rey 

T. unicolor (Piller & - NB NS 

Mitterpacher)t 

Genus TRIGONOGENIUS Solier 

T. gJobulus Soliert - BC 

farctus LeConte 

Genus NIPTUS Boieldieu 

N. hololeucus (Faldermann)f - BC AB SK MB ON PQ NB NS PE - NF 

205

Trogossitida e 

Genus PSEI.IDEUROSTUS Heyden 

P. hilleri (Reitter)t 

alienus (W.J. Brown) 

Genus PTINUS Linn6 

(Subgenus I'TINUS s. str.) 

P. bicinctus Sturmf 

P. clavipes ['anzert 

b r unne;t. ts Duftschmid 

hirtellus Sturm 

testaceus Olivier 

P. fur (Linn6)t 

humeralis Say 

lofro Fabricius 

P. raptor Sturmt 

P. villiger (Reitter)-f 

(Subgenus CYNOPTERUS Mulsant & Rey) 

P. californicus Pic 

P. fallax Fall 

P. falli Pic 

P. ocellus W J. Brownf 

tectus aoct. 

AK 

BC AB 

ON PQ NB 

_BCABSK 

_BC 

BC 

BC 

BC 

BC 

BC 

BC 

MB 

oN PQ 

ON PQ 

_SKMBONPQ 


NB 

ON PQ NB NS NF 

MB_PQ 

-oN 

_SKMBONPQ 

NB 

NS 

NS 

NS 

NS 

NS 

NS 

_NF 

NF 

NF 

Superfamily Lymexyloidea 

FamilyLYMD(YLIDAE ship-timberbeetles 

J.M. Campbell 

Thirt;'ssvsn species of Lymexylidae are known 

around the world. Only one rare species occurs in 

Canada. .\dults are found in decaying wood, on tree 

trunks, or under bark. The Canadian species is 

probably .introduced, since it also occurs in Europe 

and was firrmerly very destructive to wooden ships. 

The classifrcation of the family is now fairly stable. 

Crorvson (1955) and Wheeler (1986) believe 

that Lymexylidae is closely related to Cleroidea and 

Cucujoidea. Arnett (1968) places the farnily near 

Lampyridae, based on wing venation. Wheeler 

(1986) revised the world genera and higher classifications, 

including a study of the phylogerny, morphology, 

and known biology of the genera. 

aB (1);ON (1);PQ (1);NB (1); NS (1);NF (1) 

Genus IIYLItrCOETUS Latreille 

Elatero;;.des J.C. Schaeffer 

Hyloceotus Melsheimer 

Subfamily FIYI ECOETINAE 

H. Iugubris lJayf AB ON PQ NB NS 

americc:nus }{aris 

NF 

Superfamily Cleroidea 

Family TROGOSSITIDAE bark-gnawing beetles 

Y. Bousquet 

Trogossitidae includes 22 species in Canada and 

Alaska. Most North American species are found 

under the bark of dead trees, in logs, and in stumps; 

a few species occur in fungi. The adults and larvae 

of many species feed on wood-eating insects; others 

feed on dercaying matter or fungi. Tenebroides mauritanicus 

is a well-known stored-product pest that 

feeds on grains and cereal products. 

Barron (1971) has revised the trogositids of 

North America. The species are divided into two 

subfamilies, Peltinae and Trogossitinae. 

AK (a);YK (a); NT (5); BC (17); aB (6); SK (6); MB 

(6); oN (12); PQ (e); NB (2); NS (2); PE (1); LB (1); 

NF(3) . H(2) 

206

Trogossitidae 

Subfamily PELTINAE 

Tribe Peltini 

Genus CAI-ITYS C.G. Thomson 

Nosodes LeOonte 

Peltidea Motschulsky 

C. minor Hatch 

C. scabra (Thunberg)* 

sercata (LeConte) 

silphides (Newman) 

_YK_BCABSK_ONPQ 

AK_NTBCAB_MBONPQ 

Genus OSTOMA Laicharting 

Gaurambe C.G. Thomson 

O. columbiana Casey 

O. ferruginea (Linn6)* 

fraterna (Randall) 

nigrina Casey 

se ptentr io nalis ( Randall) 

O. pippingskoeldi (Mannerheim) 


AKYKNTBCABSK_ONPQ 

BC 

NB 

NS 

LB 

NF 

Genus GRYNOCHARIS C.G. Thomson 

G. oregonensis (Schaeffer) 

G. quadrilineata (Melsheimer) 

marg inata (Melsheimer) 

BC 

MB ON PQ 

Genus THYMALUS Latreille 

T. marginicollis Chevrolat 

aubei L6veill6 

fulgidus Erichson 


NS 

NF 

Genus ERONYXA Reitter 

Ostomodes Reitter 

E. paliida (Motschulsky) 

dohrni (Reitter) 

lagrioides Reitter 

pilosula (Crotch) 

BC 

Subfamily TROGOSSITINAE 

Tribe Trogossitini 

Genus NEMOSOMA Latreille 

Pseudalindria Fall 

N. punctulatum Van Dyke 

punctatum Van Dyke 

BC 

Genus CORTICOTOMUS Sharp 

Parafilumis Casey 

C. caviceps (Fall) 

laeuiuentris Casey 

BC 

Genus AIRORA Reitter 

A. aequalis Reitter 

bicolor Casey 

polita Casey 

punctiuentris Casey 

A. cylindrica (Audinet-Serville) 

nlgello (Melsheimer) 

nigra (Melsheimer) 

teres (Melsheimer) 

Genus TEMNOCHILA Westwood 

T. chlorodia (Mannerheim) NT 

melanica lJ.atc}a 

prosternalis Schaeffer 

u ir idicy anea (Mannerheim) 

BC 

BC 

ON 

207

Cleridae 

Genus TEI'IEBROIDES Piller & Mitterpacher 

Trogossita Olivier 

T. americanus tKirby) 

castat teus (Melsheimer) 

nigrit ^rs 

(Horrr) 

T. collaris Sturm) 

T. corticalis (Melsheimer) AK 

dubius (Melsheimer) 

internrcdius (.Horn) 

limbalis (Melsheimer) 

T. crassico'nis (Horn) 

califo.ni"u" 11fornt 

pleurulis (.Horn) 

T. laticollir, (Horn) 

obscu:us (Hotn) 

T. mauritanicus t Linn6rf 

nitidt s (Horn) 

T. occidenlalis Fall 

T. sinuatu,r (LeConte) 

Cleridae includes 50 species and subspecies in 

Canada and Alaska. Most species are predaceous, 

the adul':s feeding on adult bark beetles and the larvae 

on eggs, larvae, and adults of wood-boring beetles 

in galleries. Several species of Enoclerus and 

Thanasimus are believed to be imoortant in the 

natural r:ontrol ofbark beetles in coniferous forests. 

Adults of Cleridae are found mostly on plants, tree 

trunks, or under bark. Some are found on flowers, 

feeding on pollen. Some species of Necrobia are 

scavengers and are economically important as pests 

of stored food, carcasses, and skins. 

ON 

AK (5); vK (4); NT (5); BC (22); AB (12); SK (10); 

MB (14); ON (32); PQ Q4; NB (11); NS [4); NF (4) 

. I(5) 

PQ 

-- - - - 

SK 

-oN 

MB ON PQ NB PE NF 

In North America the taxonomy of clerids is reasonably 

well known, but the literature is scattered. 

A revision of the family would be helpful. Barr 

(I962a) has published a key to the genera. There is 

no comprehensive work for the Canadian fauna, but 

most eastern species could be identifred by referring 

to Knull (1951) and most western species by referring 

to Barr 0962b). 

YK NT ec- sxusBNpe - 

BC 

BC 

BC 

BC 

Family CLERIDAE checkered beetles 

J. McNarnara 

Subfamily CLERINAE 

Tlibe Tillini 

Genus MONOPHYLLA Spinola 

Elasn ocerus LeConte 

Macrttelus lllug 

M. terminata (Say) 

megatoma Spinola 

ON 

Genus CYIyIATODERA Gray 

C. bicolor (Say) 

C. decipiens Fall 

C. inornatiL (Say) 

tenera LeConLe 

C. ovipennis LeConte 

pilosei.Ia LeConte 

C. undulata (Say) 

boscil (Chevrolat) 

longicollis Spinola 

uauth.ieri Spinola 

BC 

BC 

oN PQ 

ON PQ 

ON 

_-n'___ 

Tribe Phyllobaenini 

Genus PH'{LLOBAENUS Deiean 

Hydnocera Newman 

208

Cleridae 

P. dubius (Wolcott) 

P. humeralis (Say) 

cyanescens (LeConte) 

cydneus (Spinola) 

dfficilis (LeConte) 

P. lecontei (Wolcott) 

P. pallipennis (Say) 

serratus (Newman) 

P. scaber (LeConte) 

P. subfasciatus (LeConte) 

P. unifasciatus (Say) 

punctatus (Spinola) 

P. verticalis (Say) 

brachypterus (I0.ug) 

I ine a toc oll is ( Spinola) 

tenellus (.Say') 

Genus ISOHYDNOCERA E.A. Chapin 

I. curtipennis (Newman) 

longicollis (Ziegler) 

L tabida (LeConte) 

- - - ecABSfMBoNFeNs- - - - 

T- I'BC-:"MBBNiSNBws- - - 

_BC 

_BCABSKMB 

-oN 

-oNPQNB 


_ONPQ 

Tribe Thaneroclerini 

Genus ZENODOSUS Wolcott 

Z. sanguineus (Say) 

-ONPQ 

Genus PRIOCERA Kirby 

Tribe Clerini 

P. castanea (Newman) 

maculata Ziegler 

- ON 

rufescens Spinoia 

Genus OPILO Latreille 

O. domesticus (Sturmt* 

Genus THANASIMUS Latreille 

T. dubius (Fabricius) AK YK NT BC AB SK MB ON PQ NB - NF 

erythrocep halus (Spinola) 

ruficeps (Spinola) 

T. trilasciatus (Say) - ON PQ 

T. undatulus (Say) AK \T( NT BC AB SK MB ON PQ NB NS NF 

abdominalis I{rtby 

mel anocep halus Chevrolat 

nubilus (l\lug) 

Genus PLACOPTERUS Wolcott 

P oeci loc hroa Chevrolat 

P. thoracicus tOliviert MB ON PQ 

monilis (Melsheimer) 

pallipes (Wolcott) 

Genus ENOCLIIRUS Gahan 

E. barri Knull - BC 

E. coccineus coccineus (Schenkling) AB SK 

corallinus (Fall\ 

E. eximius (Mannerheim) - BC 

holosericeus (A. White) 

E. ichneumoneus (Fabricius.t - ON 

rufus (Olivier) 

E. Iaetus tKlugt - BC 

abruptus (LeConte) 

E. lecontei (Wolcott) - BC 

nigriuentris (LeConte) 

E. muttkowskii (Wolcott) - ON PQ NB 

liljebladi Wolcott 

209

Cleridae 

E. nigrifrons tSayt * ON PQ 

gerhardi Wolcott 

testaceipes Wolcott 

E. nigripes nigripes {Say) MB ON PQ NB 

dublzs (Spinola) 

incertus (LeConte) 

E. nigripes rufiventris (Spinola) - ON PQ NB 

E. rosmarus (Say) - ON 

oculatus (Spinola) 

E. schaefferi Barr - BC AB 

humer,zlis Schaeffer 

E. sphegeus (Fabriciust - BC AB 

sobrius (Walker) 

Genus TRIOHODES Herbst 

Key to Norbh American species: Foster (1976) 

T. apivorus borealis Wolcott & 

E.A. Chapin 

interruptus LeConte 

nuttalii Say 

trifasaatus Strnn 

T. nutalli tliirby) AB SK MB ON PQ 

T. ornatus lrartwegianus A. White AK YK NT BC AB 

Genus PHI OGISTOSTERNUS Wolcott 

Phyllo nenus Spinola 

Subfamilv KORYNETINAE 

Tribi Epiphloeini 

P. dislocatus {Sayt MB ON PQ NB 

distrop.hus (IArg) 

transuersalis (Spinola) 

Genus NEI(IHNEAWolcott & E.A. Chapin 

N. laticorni ; (Sayt - ON 

Genus NECTRTHOPLEURA Barr 

Key to sprecies: Barr (1976) 

Tribe Enopliini 

N. thoracicr'L ( Say r 

b imac t.iata (Melsheimer) 

- ON PQ 

nig ript: n ni s ( Spinola) 

p ennsy lu anica (Chevrolat) 

p uncto tis s im a (Chevrolat) 

Genus CHARIESSA Perty 

B r achy mor p hus Chevrolat 

C. elegans Horn - BC 

lemberti La Rivers 

C. pilosa (F)rster) - BC AB SK MB ON PQ 

cinclo lSpinola) 

margirLata (Say) 

onusta (Say) 

Genus BOSCHELLA Barr 

B. fasciata tl-eConte) - BC 

trilobata (Van Dyke) 

Genus CREGYA LeConte 

Galeruclerus Gahan 

C. oculata (Say) - ON 

marginipennis ( Spinola) 

2to

Melyridae 

Tribe Korynetini 

Genus NECROBIA Olivier 

N. ruficoilis (Fabricius)t 

N. rufrpes (DeGeer)t 

N. violacea (Linn6)t 

errons Melsheimer 

AK YK 

-BC-oNPQ 

_BCABONPQNF 


NF 

Subfamily TARSOSTENINAE 

Genus TARSOSTENUS Spinola 

T. univittatus (Rossi)f 

alb ofa sc iat us ( Melsheimer) 

BC 

Family MELYRIDAE soft-winged flower beetles 

D.E. Bright 

Melyridae includes 49 species known to occur in 

Canada. The adults commonly occur on flowers and 

evidently feed on other insects as well as on pollen. 

Larvae are primarily predators, occurring in various 

habitats but most commonly in soil, leaf litter, 

or under bark. One species, Malachius deneus,has 

been introduced in North America and causes considerable 

damage to developing wheat in the Prairie 

Provinces. 

The taxonomy of the North American melyrids 

is in poor condition, and the family is in need of 

careful study. There are no recent revisions or taxonomic 

treatments of the Nearctic species; the information 

below therefore reflects our best effort at 

achieving a list based on available literature. 

Arnett (1968) provides a key to the genera and 

Hatch (1962) a key to the species of the Pacific 

Northwest. 

YK (2); NT (3); BC (35);aB (16); SK (5); MB (6); ON 

(10);PQ (10);NB (1);NS (1);NF (1) ' I(2) 

Genus COLLOPS Erichson 

C. bipunctatus (Say) 

C. cribrosus LeConte 

cy anip en nis Motschulsky 

C. discretus Fall 

C. hirtellus LeConte 

C. quadrimaculatus (Fabricius) 

C. tricolor (Say) 

C. versatilis FalI 

C. vittatus (Say) 

hauorthi Westwood 

Subfamily MALACHIINAE 

BC 

BC 

__ftBCy:"_ 

AB 

SK 

SK MB 

- - lrr Bc AB ir nrs 

oNpe----- 

oN PQ 

ON PQ 

Genus ENDEODES LeConte 


E. collaris LeConte 

Moore & Legner (1975b) 

BC 

Genus MALACHIUS Fabricius 

Anthocomus Erichson 

Hapalorhinus LeConte 

Microlipus LeConte 

M. aeneus (Linn6)l 

M. auritus LeConte 

M. bipunctatus (Harrer)t 

M. blaisdelli Hopping 

M. erichsonii (LeConte) 

criddlei W.J. Brown 

lateralis LeConte 

M. flavilabrus (Say) 

coeruleus Randall 

-BCAB 

_BC 

-Bcms 

ON PQ 

ON PQ 

ON PQ 

NB NS NF 

2tl

Melyridae 

M. floricola Marshall - BC 

M. francisc,rnus Fall - BC 

M. horni FrLll - BC AB 

antenruatus Hopping 

M. macer fl.orn - BC 

M. mixtus l{orn - BC 

M. mirandrrs (LeConte) - BC 

M. moerens. LeConte - BC 

u n ifor in is Motschulsky 

M. nigrinui Fall - BC 

M. thevene,ii Horn - BC AB 

contorl.us Fall 

M. ulkei H

Sphindidae 

Genus TRICHOCHROUS Motschulsky 

T. alberbensis Blaisdell 

Genus AMECOCERUS Solier 

Listrus Motschulsky 

A. confusus Casey 

A. glabratus Hatch 

A. medicatus Blaisdell 

A. olSrmpianus Blaisdell 

A. plenus Casey 

A. provincialis Blaisdell 

A. quadricollis Hatch 

A. robustus Blaisdell 

A. senilis (LeConte) 

Genus DASYRHADUS FalI 

D. impressicollis Fall 

_BC 

_BC 

_BC 

_BC 

-BC 

_BC 

_BC 

BC 

AB 

AB 

AB 

Superfamily Cucuj oidea 

FamilySPHINDIDAE dry-fungusbeetles 

J.M. Campbell 

Sphindidae is a small family comprising 35 

species around the world, of which five are known to 

occur in Canada. Adults and larvae feed on spores 

of slime molds. 

Casey (1898) prepared the last revision of the 

North American sphindids. Sen Gupta and Crowson 

(1979) revised the higher classification of the family, 

which is in need of a taxonomic revision; most 

current records are unreliable. 

BC (2);AB (1); oN (3);PQ (4) 

Subfamily SPHINDINAE 

Tribe Odontosphindini 

Genus ODONTOSPHINDUS LeConte 

O. clavicornis Casey 

O. denticollis LeConte 

BC 

ON 

pc) 

Tribe Sphindini 

Genus SPHINDUS Chewolat 

Coniophagus Mink 

S. americanus LeConte 

S. trinifer Casey 

-BCABONPQ 

_ONPQ 

Genus EURYSPHINDUS LeConte 

E. hirtus LeConte 

Subfamily EURYSPHINDINAE 

PQ 

213

Nitidulidaer 

Family NITIDULIDAE sap beetles 

J. McNamara 

The lamily Nitidulidae comprises 101 species in 

Canada and Alaska. Members of this family are 

mostly saprophagous and mycetophagous. The 

majority live in decaying fruit, fermenting plant 

juice, an

Nitidulidae 

C. dimidiatus (Fabricius)* 

C. discoideus LeConte 

apicalis LeConte 

caudalis (LeConte) 

C. hemipterus (Linn6)f 

b imac ulatu s Melsheimer 

C. ligneus Murray 

decipiens l{orn 

C. lugubris Murray 

C. marginellus Motschulskyf 

nitens Fall 

C. mutilatus Erichsonf 

luridus Murray 

C. pallipennis (Say) 

pollens Sharp 

C. sayi Parsons 

AK 

AK 

BC 

BC 

BC 

AB 

AB 

AB 

SK 

MB 

-oN 

MB ON 

MB 

oN PQ 

ON 

ON 

PQ 

pe 

PQ 

Subfamily NITIDULINAE 

Genus EPURA-EA Erichson 

Epuraeanella Ctotch 

E. adumbrata Mannerheim 

E. aestiva (Linn6)t 

conu exi u sc ula Mannerheim 

E. alternans Grouvelle 

alticola Fall 

E. alternata Parsons 

E. ambigua Mannerheim 

E. avara (Randall) 

nubila LeCotte 

E. boreades Parsons 

E. corticina Erichson 

E. erichsoni Reitter 

E. flavomaculata Miiklin 

E. fulvescens Horn 

E. helvola Erichson 

castanea (Melsheimer) 

rufa Reitter 

E. horni Crotch 

duryiBlatchley 

E. labilis Erichson 

E. linearis Maklin 

E. obliquus Hatch 

E. obtusicollis Reitter 

E. ovata Horn 

E. parsonsi Connell 

E. peltoides Horn 

E. planulata Erichson 

placida Meklin 

E. populi Dodge 

E. rufa (Say) 

bodio (Melsheimer) 

rotundicollis Reitter 

E. rufida (Melsheimer) 

E. terminalis Mannerheim* 

infuscata M€iklin 

E. truncatella Mannerheim 

nigraMeklin 

ornatula Notman 

E. umbrosa Horn 

AK-NTBCAB 

AK BC AB 

_BC 

AK_- 

AK YK 

AK 

AK 

AX 

AK YK 

AK YK 

_I' 

I"_ 

NT 

_BC 

BC 

BC 

BC 

NT BC 

BC-SK 

_AB 

SK 

BC 

BC AB 

BC AB 

BC AB SK 

BC 

BC AB SK 

_AB 

BC AB 

BC AB 

AB 

AB 

SK 

ms oN 

-oNPQ 

MB ON PQ 

_ONPQ 

PQ 

MB ON PQ 

MB ON PQ 

-oNPQ 

MB ON PQ 

ON PQ 

MB ON PQ 

PQ 

PQ 

MB_PQ 

MB_PQ 

PQ 

MB ON PQ 

MB ON PQ 

MB_PQ 

_ONPQ 

PQ 

PQ 

_ONPQ 

MB ON PQ 

MB ON PQ 

NB 

NB 

NS 

NB 

NB 

NB 

PQ NB 

NS 

NS__ 

Ns__ 

LB 

NF 

NF 

NF 

NF 

Genus APHENOLIA Reitter 

A. monogama (Crotch) 

BC 

Genus STELIDOTA Erichson 

S. octomaculata (Say) 

biseriata Reitter 

ON 

PQ 

2t5

Nitidulidae 

Genus OMOSITA Erichson 

(Subgenus IJAPROBIA Ganglbauer) 

O. colon (Linn6)t - BC AB SK MB ON pe NB NS NF 

O. discoidezL (Fabricius){ AK BC AB - MB ON pe - NS NF 

inuersc: LeConte 

Genus NITIDULA Fabricius 

N. bipunctata (Linn6)t AK YK - BC AB - MB ON pe 

N. carnaria (Schaller)i BC - ON pe - NS 

N. nigra Sc.iraeffer AK yK - BC AB - MB 

N. rufipes (,inn6)t - BC AB - MB ON pe NB NS NF 

ossium l{ttby 

N. ziczac Say NT BC AB - MB 

humerr:tlis LeConte 

inorna;a Horn 

unig u t i:a t a Melsheimer 

Genus PRO METOPIA Erichson 

P. sexmaculata tSayr ON pe 

Genus LOB [OPA Erichson 

Cerophorus Castelnau 

L. setosa Hi,rold BC 

substriata (Hamilton) 

L. undulata tSayr AB - MB ON pO 

Genus SOR')NIA Erichson 

S. grisea (Linn6rt 

NF 

S. guttulata tl-eConter - BC 

Genus PHE.\OLIA Erichson 

P. grossa tF.rbricius) ON pO 

Genus THA-YCRA Erichst-rn 

Perthaiycra Horn 

Key to New World species: Howden (1961) 

T. concolor (LeConte) - YK - BC AB - MB ON pe 

T. dentata I l. Howden - BC 

T. keltoni H Howden - BC AB 

T. leechi H. Howden BC 

T. mixta H. Howden BC AB 

T. monticola H. Howden - BC 

T. murraf lHornt - BC AB 

T. parsonsi .r{. Howden - BC 

T. sinuata H. Howden - BC AB 

Genus POC,,\DIUS Erichson 

P. fulvipenn s Erichson BC 

dorsali:; Horn 

P. helvolus llrichson NT BC - SK MB 

breuiusculus Reitter 

infusca;',us Reitter 

limbatt.s Reitter 

Genus AMP.] ICROSSUS Erichson 

A. ciliatus t()liviert - ON 

unilineatus (.Say) 

Genus CYCIIRAMUS Kugelann 

Quadrifrons Blatchley 

C. adustus !.richson * ON pO 

Genus PALL,ODES Erichson 

P. pallidus tl)alisot de Beauvois) - ON 

silaceus Erichson 

216

Nitidulidae 

Genus CYLLODES Erichson 

C. biplagiatus LeConte MB 

Genus MELIGETHES Stephens 

Key to North American species: Easton (1955) 

M. atratus (Olivier)l 

M. canadensis Easton 

M. cleominis Easton 

M. dauricus Motschulsky* 

californicus Reitter 

moerens LeConte 

rufimanus LeConte 

M. nigrescens Stephens* 

seminulum LeOonte 

M. pinguis Horn 

M. saevus LeConte 

M. simplipes Easton 

Genus PITYOPHAGUS Shuckard 

Subfamily MELIGETHINAE 

Subfamily CRYPTARCHINAE 

P. rufipennis Horn - BC 

Genus CRYPTARCHA Shuckard 

(Subgenus CR\?TARCHA s. str.) 

C. ampla Erichson - BC AB - MB ON PQ 

(Subgenus LEPIARCHA Sharp) 

C. concinna Melsheimer 

bella Reitter 

- BC AB - MB ON PQ 

liturata LeConte 

piclo Melsheimer 

Genus GLISCHROCHILUS Reitter 

(Subgenus LIBRODOR Reitter) 

G. fasciatus (Olivier) - BC MB ON PQ - NS 

g eminat us (Melsheimer) 

q uadr imaculoso (Melsheimer) 

G. quadrisignatus (Say) - BC AB - MB ON PQ - NS NF 

b ip unctatu s (Melsheimer) 

bipustulatus (Melsheimer) 

canadensis W.J. Brown 

s exp u st ulatu s (Reitter) 

sirnilis (Melsheimer) 

G. sanguinolentus rubromaculatus PQ NB NS 

(Reitter) 

G. sanguinolentus sanguinolentus - BC AB ON PQ 

(Olivier) 

G. siepmanni W.J. Brown - BC AB SK MB ON PQ NB 

(Subgenus GLISCHROCHILUS s. str.) 

_BC 

AKYK_BCAB_MBONPQ 

_BCABSKMB 

_BCABSKMB 

_YK_BCAB_MBONPQ_NSPE 

AB-nrn------NF 

_ONPQ_NSPE 

G. confluentus tsayt - ON PQ 

G. lecontei W.J. Brown - BC IVIB 

G. moratus W.J. Brown - BC AB SK MB ON PQ 

G. vittatus (Say) AK BC AB - MB ON PQ 

dejeanii (Kirby) 

sepulcralis (Randall) 

Genus CYBOCEPHALUS Erichson 

C. californicus Horn 

Subfamily CYB OCEPHALINAE 

_BC 

217

Rhizophagi lae 

Family RHIZOPIIAGIDAE rhizophagid beetles 

Y. Bousquet 

Twerrty-four species and subspecies of rhizophagids 

are known to occur in Canada and Alaska. 

They are collected most commonly under the bark of 

dead trees, in rotten logs, and in decaying vegetation. 

A few species live in ant nests, and some species of 

Monotomit are occasionally found in buildings. Very 

little is known about their feeding habits. Members 

of Rhizophaginae are mostly predaceous, feeding on 

wood-boring beetles, especially scolytids; those of 

Monotominae are apparently fungus feeders. 

The taxonomy of the Nearctic species has been 

poorly studied, and most genera are in need of modern 

revision. Crowson (1955) recognized four subfamilies, 

two of which (Rhizophaginae and 

Monotominae) are found within the borders of 

Canada and Alaska. Sen Gupta (1988) recently 

reviewed the world genera of the family. 

AK (3); vK (1); sC (15);AB (7); SK (3); MB (a); ON 

(15);PQ (11);NB (a);NS (3);NF (5) . I (3) 

Subfamilv RHIZOPHAGINAE 

Genus RHIZOPHAGUS Herbst 

Key to North American species: Bousquet (1990) 

(Subgenus ;\NOMOPHAGUS Reitter) 

R. brunneuri brunneus Horn 

R. galbus Bousquet 

R. pseudobrunneus Bousquet 

_BC 

AKBCAB_MB 

AB 

ON 

PQ 

v_ 

(Subgenus ITURHIZOPHAGUS M6quignon) 

R. grouvellei M6quignon 

BC 

(Subgenus RHIZOPHAGUS s. str.) 

R. dimidiatus Mannerheirn 

testaceits M6quignon 

R. minutus minutus Mannerheim 

R. minutus rotundicollis Bousquet 

R. paralleloi:ollis Gyllenhalf 

robustt,ts Schaeffer 

R. remotus .LeConte 

luteus ,VI6qrignon 

R. sayi Sche,effer 

bipunc,tatus (Say) 

R. sculpturu.tus Mannerheim 

AKYK_BCABON 

BC 

_BC_ON 

AK 

BC AB 

BC 

-oN 

MB ON 

-oN 

PQ 

pe 

PQ 

PQ 

NS 

NS 

NS 

NF 

NF 

NF 

Subfamily MONOTOMINAE 

Genus PYCI\OTOMINA Casey 

P. cavicollis (Horn) 

ON 

PQ 

Genus MACREUROPS Casey 

M. longicoll:s (Horn) 

Genus BAC'IRIDIUM LeConte 

Crine Pascoe 

B. ephippigerum (Gu6rin-M6neville) 

B. striolaturn (Reitter) 

Genus HES PEROBAXNUS LeConte 

H. abbrevia':us (Motschulsky) 

rufipennis (LeConte) 

Genus MOITIOTOMA Herbst 

BC 

BC 

BC 

-oN _ONPQ 

M. americana Aub6 

M. arida Casey 

M. fulvipes .Melsheimer 

opaca iJiegler 

p ar allelocollis Motschulsky 

subnigra Motschulsky 

--SCAB-nfe 

ON 

ON 

PQ 

218

Cucujidae 

M. longicollis (Gyllenhal)f 

M. parallela LeConte 

M. picipes Herbstt 

foueata LeConte 

M. producta LeConte 

M. quadraria Casey 

_BCABSK 

_BC_SK 

_BCAB 

SK 

_ONPQ 

_ONPQ 

MB ON PQ 

ON 

NB 

NB 

NB 

NF 

NF 

Family CUCUJIDAE flat bark beetles 

Y. Bousquet 

The cucujids currently include 31 species and subspecies 

in Canada and Alaska. The adults are usually 

found under the bark ofdead trees, in logs, and in 

decaying plant matter. Some species apparently feed 

on fungi and decaying matter, others prey on insects. 

Six of the species occurring in Canada, Cryptolestes 

ferrugineus, C. pusillus, C. turcicus, Ahasuerus 

aduena, Oryzaephilus surinqmensls, and O. mercator, 

are associated with stored products; four ofthem are 

among the most serious pests in the country. 

Lawrence's concept (1982a) of the family is followed 

here. Many authors rank the silvanids as a 

distinct family. There is no modern taxonomic revision 

of Cucujidae, but some genera have recently 

been revised. 

AK (3);vK (1);NT (2); BC (15);AB (10); SK (9); MB 

(17); oN (26); PQ (24); NB (5); NS (a); PE (1); LB 

(1);NF (5) . H (1);I (10) 

Subfamily CUCUJINAE 

Genus CUCUJUS Fabricius 

C. clavipes clavipes Fabricius 

C. clavipes puniceus Mannerheim 

subnitens Schaeffer 

AK 

_MB 

BC AB SK MB 

oN PQ 

_NS 

NF 

Genus PEDIACUS Shuckard 

P. depressus (Herbst)f 

subglaber LeConte 

P. fuscus Erichsonx 

planus LeConte 

s ub car inatu s Mannerheim 

AK YK 

_BC 

NT BC 


NB 

LB 

Subfamily PASSANDRINAE 

Genus CATOGENUS Westwood 

C. rufus (Fabricius) 

oN PQ 

Genus PLACONOTUS Macleay 

Siluanophloeus Sharp 

Key to New World species: Thomas (1984) 

P. zimmermanni (LeConte) 

apertus (Casey) 

illustris (Casey) 

sobrinus (Casey) 

Subfamily LAEMOPHLOEINAE 

MB ON 

PQ 

Genus LAXMOPHLOEUS Deiean 

L. biguttatus (Say) - BC - SK MB ON PQ NB NS PE 

b isig nat u s Gu6rin-M6neville 

L. fasciatus Melsheimer MB ON PQ 

Genus CHARAPHLOEUS Casey 

C. adustus (LeContet MB oN PQ -NS 

C. convexul us (LeConte t ON 

2r9

Cucujidae 

Genus CRY FIOLESTES Ganglbauer 

Key to New World species: Thomas (1988) 

C. ferrugine,us (Stephens)t - BC AB SK MB ON PQ NF 

C. punctatus (Lecontet - ON 

adumbratus Casey 

extrica:lus (Casey) 

geminctus (LeConte) 

C. pusillus Sch

Cryptophagidae 

Family CRYPTOPIIAGIDAE silken fungus beetles 

Y. Bousquet 

Sixty-six species of cryptophagids are recorded 

in Canada and Alaska. These beetles are usually 

found on mold, on fungi, under bark, as well as in 

decaying vegetation and nests of social 

Hymenoptera, birds, and mammals; one species, 

Myrmedophila americana, lives in ant nests. 

Adults and larvae of most species feed on mold and 

decaying vegetation; those of Telmophilus species 

apparently feed on flower heads of certain aquatic 

plants. Several species of Cryptophagus are found 

in stored products, where they feed on mold. 

This group is poorly studied taxonomically. 

Since Casey's (1900) revision of the North American 

fauna, only the genus Cryptophagus has been 

revised (Woodroffe and Coombs 1961). Many genera 

previously included within the cryptophagids 

have been transferred recently to Languriidae. The 

reader can expect many nomenclatural changes at 

the specifrc level, as well as many new distribution 

records, when the cryptophagid groups are revised. 

The generic classification of the Cryptophaginae 

used here follows Bousquet (1989). 

AK (26); YK (1); NT (3); BC (47); AB (15); SK (7); 

MB (12); oN (1e); PQ (17); NB (2); NS (3); LB (2); 

NF (3) . H (11);I (8) 

Genus TELMATOPHILUS Heer 

T. americanus LeConte 

Subfamily TELMATOPHILINAE 


Genus ANTHEROPHAGUS Latreille 

A. convexulus LeConte 

A. ochraceus Melsheimer 

oregon us Casey 

A. pallidivestis Casey 

A. suturalis Meklin 

Subfamily CRYPTOPHAGINAE 

Key to North American genera: Bousquet (1989) 

Tlibe Cryptophagini 

- - - ec AB !r< r,re 3N 

AK 

_BC 

BC 

PQ NB NS 

PQNBNSPE_NF 

Genus HENOTIDERUS Reitter 

Crosimus Casey 

Henoticoides Hatch 

H. lorna (Hatch) 

H. obesulus (Casey) 

AK 

AK_NT 

BC 

AB ON PQ NS 

Genus HENOTICUS C.G. Thomson 

Coniophthalmo Kangas 

Glyptophorus Patk 

H. californicus (Mannerheim) 

inconspicuus (LeConte) 

H. serratus (Gyllenhal) 

denticulatus (LeConte) 

AK 

BC 

BC 

MB ON PQ 

Genus PTERYNGIUM Reitter 

P. crenatum (Gyilenhal)* 

malacum Casey 

BC 

PQ 

Genus SALEBIUS Casey 

S. octodentatus (Mriklin) 

sexdentatus Casey 

AK 

BC 

Genus CRYPTOPHAGUS Herbst 

C r y pto p hag i stes Crotch 

Key to North American species: Woodroffe and Coombs (1961) 

C. acutangulus (Gyllenhal)* AK BC 


restrictus Casey 

AB 

MB ON PQ 

NF 

227

Cryptophag idae 

C. bidentatus Maklin AK BC 

puncta tis sim us MEikIin 

C. cellaris (llcopoli)+ BC AB SK MB ON 

angustztus Casey 

rupimc,ntis Casey 

C. confertus Casey* AK - NT BC AB - PQ 

arc han. gelicus J.R. Sahlberg 

C. corticinus C.G. Thomson* AK BC - ON PQ 

depressulus Casey 

C. dentatus (Herbst)* AK BC 

fumatus (Marsham) 

q uad.ride ntat us Mannerheim 

C. difficilis tlasey - BC AB - PQ 

contractus Casey 

C. distinguendus Sturm* - BC 

heeni Casey 

C. hebes Casey AK BC 

lyraticollis Casey 

o/ioszs Casey 

C. histricus Casey BC LB 

C. lapponicLrs Gyllenhalx - BC 

ualidus Kraatz 

C. laticollis Lucast - BC AB 

af,finic Sfrrm 

cribricollis Casey 

inscitu.s Casey 

C. maximus Blake - BC 

C. obsoletur Reittert - ON 

C. peregrin rs Woodroffe & Coombs - BC 

C. pilosus C yllenhalf - BC - - MB ON 

cicatrir.osus Casey 

infuscatus Casey 

paruinodus Casey 

plectrum Casey 

C. porrectur' Casey AK 

C. quadrihamatus Mtiklinx AK 

BC 

plagiatus Poppius 

C. saginatu; Sturml AK BC - ON 

C. scanicus tlinn6)t - NF 

C. scutellat rs Newmant ON 

C. setulosus, Sturm* - BC - ON PQ - LB 

antennatus Casey 

grossutus Casey 

C. stromus Woodroffe & Coombs - SK MB 

C. subluma us Kraatzl - BC AB SK - 

C. tubercult,sus Meklinx AK BC AB 

ON PQ 

bouldetanus Casey 

breuipilis Casey 

debilis LeConLe 

instabtlis Bnrce 

lepidut Casey 

C. varus Woodroffe & Coombs - BC AB SK 

Genus MYF,MEDOPHILA Bousquet 

M. americana (LeConte) AK l'K - BC AB - MB - PQ 

blzmi iBlaisdell) 

Genus CAENOSCELIS C.G. Thomson 

Macrodea Casey 

Tribe Caenoscelini 

C. anlenna]is (Caseyt AB - MB 

C. basalis C asey PQ 

C. ferruginea tC.R. Sahlbergt* AK BC 

C. parallela Casey AK BC 

222

Languriidae 

Genus EPHISTEMUS Stephens 

(Subgenus EPHISTEMUS s. str.) 

E. globulus (Paykull)* 

apicalis LeOonte 

Subfamily ATOMARIINAE 

Tribe Atomariini 

BC 

ON 

Genus ATOMARIA Stephens 

(Subgenus ANCHICERA C.G. Thomson) 

A. aleutica Casey 

A. apicalis Erichsonf 

A. curtula Casey 

pumilio Casey 

A. ephippiata Zimmermann 

hesperica Casey 

A. fulvipennis Mannerheim 

A. kamtschatica Motschulsky* 

fallax Casey 

A. laetuia LeConte 

A. nubipennis Casey 

A. ochracea Zimmermann 


A. ovalis Casey 

A. pusilla (Paykull)* 

A. saginata Casey 

(Subgenus ATOMARIA s. str.) 

(Agathengis des Gozis) 

A. capitata (Casey) 

A. fuscicollis Mannerheim* 

A. lepidula Maklin 

A. longipennis (Casey) 

nigricollis (Casey) 

A. melanica Hatch 

A. planulata Maklin 

A. quadricollis (Casey) 

A. stricticollis (Casey) 

A. subdentata (Casey) 

A. vespertina Miiklin 

Genus HYPOCOPRUS Motschulsky 

H. tenuis Casey 

1"_ _ _ _ 

AK 

AK 

AK 

AK 

AK 

AK 

AK 

NT BC AB 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

_MB 

SK MB 

ON 

ON 

pl:) 

- - lr,fe ON Pe 

sc - - - a*:a - -- - - - 

BC 

BC 

Subfamily IfYPOC OPRINAE 

AB 

Family LANGURIIDAE 

lizard beetles 

J.M. Campbell 

The family Languriidae, as recently defined, is 

widely distributed in all faunal regions but is best 

represented in tropical areas. OnIy seven species 

are known to occur in Canada, all but one of which 

are found east of the Rocky Mountains. Larvae of 

the subfamily Languriinae (Languriidae of previous 

authors) are stem borers of which one, the clover 

stern borer, Languria mozardi, is an economic pest 

in Canada. The feeding habits and biology of the 

larvae and adults of the other subfamilies are 

poorly known, but they probably feed on decaying 

vegetation and fungal hyphae rather than on living 

plants (Lawrence and Vaurie 1983). 

Until recent papers by Sen Gupta and Crowson, 

this family was restricted to those genera now 

placed in Languriinae. Sen Gupta (1967) transferred 

Toramus and allied genera from the family 

Cryptophagidae to the subfamily Toraminae. 

223

Languriidze 

Subsequi:ntly (1968), he transferred Loberus to the 

tribe Lolrerini, both in the family Languriidae. The 

family vl,as redefined, and the higher classification 

and genr:ra of the world were revised bv Sen Gupta 

and Crorvson (1921). The North Ameiican "o".i"" 

of the family are reasonably well known, with most 

subfamilies having been revised since 19,[8 and the 

species cataloged by Lawrence and Vaurie (1983). 

BC (1); SK (1); MB (2); ON (6); PQ (5) . I (1) 

Subfamily XENOSCELINAE 

Tribe Loberini 

Genus LOI}ERUS LeConte 

L. impressr.rs LeConte 

ON 

PQ 

Subfamily TORAMINAE 

Key to species: Sen Gupta (1967) 

Tribe Toramini 

Genus TOF:AMUS Grouvelle 

Tomarus LeConte 

T. pulchellr.Ls (LeConte) 

oN PQ 

Subfamily CRYPTOPHILINAE 

Tribe Cryptophilini 

Genus CR'IIPTOPHILUS Reitter 

C. integer ( F{eer)f 

ON 

PQ 

Subfamily LANGURIINAE 

Tlibe Languriini 


Genus LAI\ GURIA Latreille 

L. angustata (Paiisot de Beauvois) 

pulchr t LeConte 

uhlerit Horn 

L. convexicollis Horn 

in t er s t,t}ialis C asey 

L. mozardi mozardi Latreille 

apicalis Motschulsky 

BC 

ON 

MB ON 

PQ 

Genus ACROPTEROXYS Gorham 

A. gracilis (Newman) 

aztecana Casey 

diuisa (}]orn) 

inornata (Randall) 

latreillei (LeConte) 

nigrice o s (Motschulsky) 

obscuro (Motschulsky) 

pertenuis Casey 

texana Schaeffer 

thoracina Casey 

SK MB ON PQ 

224

Erotylidae 

Family EROTYLIDAE pleasing fungus beetles 

J.M. Campbell 

The family Erotylidae is found in all faunal The family is well known taxonomically; Boyle 

regions of the world but primarily in moist, tropical (1956) revised the North American species, and Sen 

areas. Only 20 species are known to occur in 

Gupta (1969) worked out the higher classifrcation. 

Canada. Adults are frequently found under bark or 

under logs. Both larvae and adults feed on the eC (5);AB (6); SK (4); MB (5); ON (1a); PQ (1a); NB 

fruiting bodies of higher fungr. (2); NS (1) 

Genus DACNE Latreille 

(Subgenus DACNE s. str.) 

(Ezgis Paykull) 

D. californica (Horn) 

cephalotes Casey 

elongata Casey 

Iaticollis Casey 

uteana Casey 

D. quadrimaculata (Say) 

(Subgenus XENODACNE Boyle) 

D. picea LeConte 

Genus MEGAI-ODACNE Crotch 

M. fasciata (Fabricius) 

bifasciatus (Olivier) 

M. heros (Say) 

Subfamily DACNINAE 

_BC 

_BC 

MB ON PQ 

-oNPQ 

-oNPQ 

Genus TRIPLAX Herbst 

Subfamily TRIPLACINAE 

T. antica LeConte - BC AB SK 

mormonalis Casey 

T. californica LeConle - BC AB 

T. cuneata Boyle - BC AB 

T. dissimulator (Crotcht AB SK MB ON PQ 

carri Casey 

T. flavicollis Lacordaire - ON PQ 

confinis LeConte 

T. frosti Casey AB SK MB ON PQ NB 

T. macra LeConte MB ON PQ 

T. thoracica Say AB SK MB ON PQ NB 

antennata Casey 



latiuscula Casey 

melano p t era Lacordarre 

obliqua Casey 

Genus TRITOMA Fabricius 

Cyrtotriplar Ctotch 

T. biguttata tSay) PQ 

basalis Lacordaire 

T. humeralis Fabricius 

ornata Casey 

- ON PQ 

ruficeps LeConte 

taeniata LeConte 

uittata LeConLe 

T. mimetica (Crotch) ON PQ 

T. pulchra Say - ON PQ - NS 

basalis Melsheimer 

225

Phalacrida,r 

cincta Lacordafte 

dimidiata Lacordaire 

sellata Ktthnt 

T. sanguinipennis (Sayl 

T. unicolor Say 

unicolor Lacordaire 

Genus ISCIIYRUS Lacordaire 

Key to Ncrth American species: Skellay and Goodrich (1989) 

I. quadripunctatus quadripunctatus 

(OIi rier) 

alabartae Schaeffer 

antediLisa Mader 

oN PQ 

ON 

oN PQ 

Family PHALACRIDAE shining flower beetles 

J.M. Campbell 

The flamily Phalacridae comprises about 600 

species around the world; only five have been 

reported to occur in Canada, but many other species 

certainly occur. Adults are usually found on flowers; 

larvae feed in flower heads of Compositae or on 

spores aILd stomata of rusts and smuts. 

Casey (1916) revised the North American 

species o[ phalacrids, but this work is very difficult 

to use. Phalacridae and Ptiliidae are probably the 

most poorly known families of beetles in Canada, 

and based on existing literature, a reliable specific 

identifrcation for most genera is not possitrle. 

BC (3);MB (1); ON (3);PQ (2) 

T?ibe Phalacrini 

Genus PHIJ-ACRUS Paykull 

P. penicillabus (Say) 

BC 

Tribe Olibrini 

Genus OLLBRUS Erichson 

O. rufipes l.,eConte 

tritus ,Jasey 

O. semistri rtus LeConte 

striatulus LeConte 

Genus STII,BUS Seidlitz 

Eustilbus Shatp 

S. apicalis (Melsheimer) 

Genus LITOCHROPUS Erichson 

BC 

BC 

ON 

ON 

L. scalptus Casey 

Genus ACI-I-AMUS Sharp 

A. pugetan-rs Casey 

PQ 

MB ON PQ 

226

Cerylonidae 

Family CERil ONIDAE cerylonid beetles 

J.M. Campbell 

OnIy about 400 species of Cerylonidae, a family 

that consists of small to very small beetles, are 

known around the world, of which seven have been 

recorded in Canada and Alaska. Most adults are 

collected in leaf litter, in rotten wood, and under 

bark. Hypodacne punctata is known in the galleries 

of Camponotus ants, and Murmidius oualis occurs 

on stored products (Lawrence and Stephan 1975). 

Little is known about the feeding habits of the larvae 

and adults, although some larvae are known to 

feed on fungal hyphae or spores, and some larvae 

feed on a plasmodium of slime molds (Slipinski 

1988). 

Adults of this family are reasonably well known 

taxonomically. Sen Gupta and Crowson (1973) 

redefined the family and revised the higher classification. 

Lawrence and Stephan (1975) revised the 

North American species, and Lawrence (1982b) cataloged 

them. 

AK (1); NT (1); eC (3); AB (2); tt{B (1); ON (6); PQ 

(3);NB (1);NS (2) . I (1) 

Subfamily ELIXESTINAE 

Genus HYPODACNE LeConte 

H. ounctata LeConte 

ON 

Subfamily MURMIDIINAE 

Genus MURMIDIUS Leach 

M. ovalis (Beck)t 

Genus MYCOCERINUS Slipinski 

Mychocerus a:uct. 

M. depressus (LeConte) 

AK 

ON 

ON 

Subfamily CERYLONINAE 

Genus PHILOTHERMUS Aub6 

P. g)abriculus LeConte 

ON PQ 

Genus MYCHOCERUS Erichson 

Lapethus Casey 

M. discretus (Casey) 

BC 

Genus CERYLON Latreille 

C. castaneum Say 

clypeale Casey 

stricticutn Casey 

C. unicolor (Zieglet) NT 

angustulum LeConte 

simplex LeConte 

syluaticum Casey 

BC 

BC 

AB 

AB 

MB ON 

ON 

PQ NB NS 

PQ 

NS 

227

Corylophidae 

Family CORYLOPHIDAE minute firngus beetles 

J.M. Campbell 

Members of Corylophidae, a small family of 

about 4315 species, are found in all major faunal 

regions, but only 11 species are known to occur in 

Canada. Adults are usually collected in moldy leaf 

litter ancl piles of grass cuttings. Adults and larvae 

are thought to feed on fungal spores. 

The taxonomy of the Canadian corylophid 

species is poorly known; reliable species identifrcation 

is not possible at this time, and a number of 

species have yet to be discovered. Casey (1900) prepared 

the last revision covering the North American 

species of Corylophidae. The classificatic,n adopted 

here is based on Paulian's (1950) reclassifrcation of 

the Old World taxa. 

YK (1); NT (2); BC (7); AB (4); SK (2); MB (1); ON 

(a); PQ (3); NB (1) . I (2) 

Genus ORTHOPERUS Stephens 

O. atomus (Gyllenhal)t 

O. scutellaris LeConte 

lucidus Casey 

O. suturalis LeConte 

Genus GRC)NE\IUS Casey 

G. fuscicorrLis Casey 

Subfamily C ORYLOPHINAE 

Tribe Orthoperini 

_BC 

NT BC AB SK 

Tlibe Rypobiini 

ON 

oN PQ 

Genus SERICODERUS Steohens 

S. lateralis (Gyllenhal)t 

Subfamily SERICODERINAE 

Tribe Sericoderini 

BC 

PQ 

Subfamily SACIINAE 

Tlibe Saciini 

Genus MOLAMBA Casey 

M. biguttat,a (LeConte) 

M. fasciata (Say) 

s u b fa s : iata (Gu6rin-M6nevilIe) 

M. obesa Casey 

BC 

BC 

AB 

oN pe N" 

Genus SACIIUM LeConte 

Parmu.lus Harold 

S. lugubre l,eConte 

S. montanum Casey 

Genus ARTHROLIPS Wollaston 

A. decolor (LeConte) 

califor nicus (Matthews) 


_BC 

yuoN_ 

AB 

228

Coccinellidae 

Family COCCINELLIDAE ladybird beetles 

J. McNamara 

Coccinellidae comprises 162 species and subspecies 

in Canada and Alaska. Many species are 

introduced in North America, some of these purposely 

for biological control. Most coccinellid adults 

and larvae are beneficial, because they are predators 

of aphids, scale insects, and mites. However, in 

the absence of preferred food, many species feed on 

the immature stages of other insects. The species of 

Psyllobora feed on fungi, and the only species of 

Epilachna that occurs in Canada feeds on plants of 

the family Leguminaceae. 

The taxonomy of the North American 

Coccinellidae is well known. thanks to the recent 

monograph by Gordon (1985). Only some of the 

most recent references on coccinellids have been 

listed in the checklist. Good regional works on 

Coccinellidae include those of Chapin (1974) for 

Louisiana, Belicek (1976) for western Canada and 

Alaska, and Larochelle (1979) for Quebec. 

/J{(26); YK (31); NT (35); BC (9a);aB (83); SK (70); 

MB (56); ON (83); PQ (77);NB (37); NS (25);pE (6); 

LB (9);NF (13) . H (2); I (8) 

Subfamily STICHOLOTIDINAE 

Tribe Microweisini 

Genus MICROWEISEA Cockerell 

Epismilia Cockerell 

Pseudoweisea Schwarz 

SmiliaWeise 

Key to species: Gordon (1970o) 

M. misella (LeContet 

BC 

AB 

SK 

MB ON 

PQ NB 

NF 

Genus COCCIDOPHILUS Brethes 

Cryptoweisea Gordon 

Key to North American species: Gordon 0970a) 

C. marginatus { Leconte) 

BC 

AB 

SK 

MB ON PQ NB 

Tlibe Serangiini 

Genus DELPHASTUS Casey 

Key to species: Gordon (19706) 

D. pusillus (LeConte) 

puncticollis (LeConte) 

sonoricus Casey 

ON PQ 

Subfamily SCYMNINAE 

Tlibe Stethorini 

Genus STETHORUS Weise 

Key to New World species: Gordon and Chapin (1983) 

S. punctillum Weiset 

S. punctum punctum (LeConte) 

S. punctum picipes Casey 

6reuis Casev 

BC 

BC 

-oN 

ABSK-ONPQ-NS 

Tlibe Scymnini 

Key to North American species: Gordon (1976) 

Genus DIDION Casey 

D. longulum Casey 

occiduum (Casey) 


D. nanum (LeConte) 

D. punctatum (Meisheimer) 

YK NT BC AB 

_NTBCABSKMB 

SK 

PQ 

ON PQ 

ON PQ NB_PE NF 

229

Coccinellidae 

Genus SCYMNUS Kugelann 

(Subgenus {ICYMNUS s. str.) 

S. americar..us Mulsant 

S. apicanus pseudapicanus Gordon 

boreal;s Gordon 

S. caurinus Horn 

aluticctllis Casey 

S. fenderi Ilalkin 

S. indianensis Weise 

rustict;s Casey 

S. nebulosus LeConte 

harneli Hatch 

infuscc:tus Boheman 

phelps;'.i Crotch 

S. opaculus Horn 

S. paracanus linearis Gordon 

(Subgenus PULLUS Mulsant) 

S. abbreviai;us LeConte 

S. aquilona::ius Gordon 

S. ardelio Eorn 

apacheanus Casey 

decipeins Casey 

sanctu; Casey 

S. aridus Cr,rsey 

S. brullei Mulsant 

diuisut Casey 

laureniicus Casey 

lodi Stt}rr 

subaen:.eus Casey 

S. calaveras Casey 

saginalus Casey 

strenuus Casey 

stygicu:.s Casey 

tenuiue stis Casey 

S. carri Gordon 

S. caudalis LeConte 

agricola Casey 

indutus Casey 

natche zianus Casey 

S. cervicalir Mulsant 

S. compar ( asey 

impun:tusWingo 

uicksbt.rgicus Casey 

S. coniferarum Crotch 

S. fraternus LeConte 

dentipes FaIl 

haemo;rhous LeOonte 

S. humboldt,i Casey 

S. impexus Mulsant'l 

S. impletus Gordon 

S. iowensis Casey 

collarit Melsheimer 

melsheimeri Weise 

S. kansanus Casey 

S. lacustris LeConte 

cultratus Wingo 

S. marginic, >ll is Mannerheim 

borealis Hatch 

californicus Boheman 

desertcrum Casey 

S. postpictus Casey 

S. puncticollis LeConte 

S. rubricaudus Casey 

canteri.us Casey 

chromopygus Casey 

texanus Casey 

S. securus J. Chapin 

S. solidus Casey 

blaisde'lli Casey 

BC 

BC 

BC 

oN PQ 

- - - :t ffi 3[ nre- - - - - - - 

BC 

AB 

AB 

_ABSK 

BC AB 

BC 

BC 

_ _ _ rSf 

_BC 

_BC 

_BC 

_BC_SKMB 

YI( NT BC AB SX 

BC 

BC 

nc 

AB 

AB 

SK MB 

SK MB 

:" 

-oN 

-oN 

ON 

ON PQ 

ON PQ 

nre oN pe 

ON 

PQ 

oN PQ 

-ONPQ 

MB ON PQ 

ON 

-oNpe-- 

_ONPQ 

oN_ 

PQ 

lln 

NB 

NB 

ivs--- 

NS_- 

NS 

230

Coccineilidae 

S. suturalis Thunbergt 

S. tahoensis Casey 

S. tenebrosus Mulsant 

S. uncus Wingo 

S. utahensis Gordon 

NT 

D/r 

AB_ 

AB SK 

AB 

-oN 

MB ON 

PQ 

PQ 

Genus NEPHUS Mulsant 

(Subgenus NEPHUS s. str.) 

N. ornatus naviculatus (Casey) 

h.incaidi (Hatch) 

N. ornatus ornatus (LeConte) 

frosti (Casey) 

sanguinifer (Casey) 

AK 

BC 

AB 

SK 

MB ON PQ 

_ONPQ 

NB NS 

NF 

(Subgenus TURBOSCYMNUS Gordon) 

N. georgei (Weise) 

bisignatus (Horlr) 

AK 

YK 

NT 

BC 

AB 

ON 

PQ 

NF 

(Subgenus SCYMNOBIUS Casey) 

N. flavifrons (Melsheimer) 

bioculatus (Mulsant) 

guttiger (Mulsant) 

Iudouictanus (Casey) 

marginellus (Mulsant) 

N. intrusus (Horn) 

inops (Casey) 

N. sordidus (Horn) 

intrusoides (Hatch) 

SK 

ON PQ 

ON 

MB ON 

Genus DIOMUS Mulsant 

D. amabilis (LeConte) 

dulcis (Casey) 

emertoni (Casey) 

D. debilis (LeConte) 

aeger (Casey) 

minutissimus (Casey) 

molliculus (Casey) 

pusio (Casey) 

D. liebecki (Horn) 

adulans (Casey) 

ohioensis (Stehr) 

D. terminatus (Say) 

femoralis (LeConte) 

partitus (Casey) 

AB 

SK 

ON 

MB ON 

ON 

PQ 

PQ 

Tlibe Hyperaspini 

Genus HYPERASPIDIUS Crotch 

H. comparatus Casey 

juniperus Nunenmacher 

H. hercules Belicek 

H. insignis Casey 

H. mimus Casey 

carri Nunenmacher 

c oloradens is Nunenmacher 

H. vittigerus (LeConte) 

YK_BC 

BC 

_AB 

-AB _AB 

AB 

SK 

SK 

Genus HYPERASPIS Redtenbacher 

Oxynychus LeConte 

H. bigeminata (Randall) 

gueri Mulsant 

H. binotata (Say) 

affinis (Randall) 

Leucopsis Melsheimer 

normata (Say) 

paspalis Watson 

H. borealis Dobzhansky 

eloli Nutting 

obscura Malkin 

YK BC AB 

SK 

-oN 

MB ON 

PQ 

PQ 

NB 

NB 

237

Coccinellidae 

schuhi }{atch 

simuloides Hatch 

H. consimilis LeConte 

canadensis Dobzhansky 

H. conviva rlasey 

congressis Watson 

insolen.s Casey 

H. disconotata Mulsant 

H. dissolutzL dissoluta Crotch 

coloraclana Casey 

nupta,iasey 

H. fastidiosr Casey 

s e pte nt rion is Dobzhansky 

H. inflexa Casey 

serena Casey 

H. jasperentis Belicek 

H. lateralis Mulsant 

flammula Nunenmacher 

idoe N rnenmacher 

laeuipennis Casey 

montartica Casey 

nig rocc. uda Dobzhansky 

omissa Casey 

wellmani Nunen.macher 

H. moerens (LeConte) 

H. octavia C asey 

H. oregona )obzhansky 

lanei E atch 

H. postica I eConte 

angust tla Casey 

elliptict Casey 

essigl n{alkin 

H. pratensis LeConte 

H. proba (Say) 

trinifer Casey 

H. quadrivittata LeConte 

tetrane;tra Casey 

H. signata signata (Olivier) 

taedato. LeConte 

H. simulatrix Dobzhansky 

H. troglodyt:s Mulsant 

discretc: LeConte 

nouasct'liae E.A. Chapin 

H. undulata (Say) 

elegans Mulsant 

guttifer a Weise 

maculilbra Melsheimer 

NT BC 

BC 

_BCAB 

_BCAB 

NT_AB 

_BCAB 

SK 

SK 

PQ 

PQ 

PQ 

_ _ sx_ 3NF3 

BC AB 

BC AB SK 

BC 

AB 

AB 

AB 

BC AB 

MB 

_MBON 

SK MB 

SK 

MB ON 

SK MB ON 

SK 

SK 

SK 

_MB 

PQ 

PQ 

PQ 

pe 

NB 

NB_ 

NB 

BC SK MB ON PQ NB 

lls 

Genus BRA(IHIACANTHA Dejean 

B. albifrons iSay) 


B. decempustulata (Melsheimer) 

stellata Casey 

troglod^rtes Mulsant 

B. dentipes (Fabricius) 

separatiz Leng 

socialis Casey 

B. quadripunctata quadripunctata 

Melsheimer 

basalis Melsheimer 

confusa Mulsant 

diuersa Mulsant 

B. rotunda Ci'ordon 

B. ursina (Fabricius) 

congruens Casey 

B. uteella CzLsey 

fenyesi Leng 

lezgl Nunenmacher 

sonorana Casey 

BC 

AB 

AB 

SK 

MB 

-oN 

ON 

ON 

ON 

SK MB ON 

PQ 

PQ 

NB NS 

PQ 

PQ NB NS 

232

Coccinellidae 

Genus BRUMOIDES E.A. Chapin 

Subfamily CHILOCORINAE 

Tribe Chilocorini 

Key to genera: Chapin (1965) 

B. histrio (Fall) - BC 

paruicollis (Casey) 

B. septentrionis septentrionis AK YK NT BC AB SK MB ON 

(Weise) 

desertorutn (Casey) 

neuadensis (Leng) 

ouoideus (Casey) 

B. septentrionis davisi (Leng) - ON PQ 

Genus EXOCHOMUS Redtenbacher 

E. aethiops rBlandi AB SK 

mormonicus Casey 

E. californicus Casey - BC 

Genus CHILOCORUS Leach 

C. hexacyclus Smith AB SK 

C. stigma (Say) AB SK MB ON PQ NB NS 

biuulnerus Mulsant 

C. tricyclus Smith - BC 

Genus COCCIDULA Kugelann 

Subfamily COCCIDULINAE 

Tlibe Coccidulini 

C. lepida LeConte AK YK NT BC AB SK MB ON PQ NB 

occidentalis Horn 

reitteri Dodge 

suturalis Reitter 

szlzrolis Weise 

Genus RHYZOBIUS Stephens 

Lindorus Casey 

R. lophanthae (Blaisdell)t - BC 

Genus ANISOSTICTA Dejean 

A. bitriangularis (Say) 

irregularis Weise 

multig uttata (Randall) 

A. borealis Timberlake 

Genus MACRONAEMIA Casey 

M. episcopalis (Kirby) 

Genus PARANAEMIA Casey 

P. vittigera (Mannerheim) 

similis Casey 

Subfamily COCCINELLINAE 

Tribe Coccinellini 


AKYKNT_MB_PQ 


_rn_ 

LB NF 

Genus COLEOMEGILLA Timberlake 

Megilla Mulsant 

C. maculata lengi Timberlake 

Genus CERATOMEGILLA Crotch 

C. ulkei Crotch 

poruo (Watson) 


-oNPQ 

ON 

233

Coccinellidz,.e 

Genus HIPI'ODAMIA Dejean 

H. americana Crotch 

H. apicalis Casey 

lezgl Johnson 

nig r om a c ula ta (Nunenmacher) 

tricolor Nunenmacher 

H. arctica (lichneider)x 

H. caseyi Jc,hnson 

H. convergens Gu6rin-M6neville 

juncta Jasey 

modestz Melsheimer 

obsoleta Crotch 

pratico!.a Mulsant 

H. expurgata Casey 

H. falcigera Crotch 

albertana Ca.sey 

cottlei (.Nwenmacher) 

H. glacialis glacialis (Fabricius) 

H. glacialis lecontei Muisant 

abduce.zs Casey 

defecta Johnson 

h op p in,;1 i Nunenmacher 

macher.zie E.A. Chapin 

p se u do;1 lacialis Johnson 

H. lunatomi rculata dobzhanskyi 

E.A. Chapin 

H. moesta nLoesta LeOonte 

H. moesta b':wditchi Johnson 

H. oregonensis Crotch 

cockere.'li Johnson 

d,ispar'Jasey 

liliputo.na Casey 

puncticollis Casey 

H. parenthesis (Say) 

albomacula Fitch 

appror,:mata Fitch 

confluezta Fitch 

connatu Fitch 

discopttnctata Fitc}' 

insulata Fitch 

linearis Fitch 

liturico!.lis Fitch 

nimia I'itc}r 

permac"ifrons Fitch 

triangu.laris Fitch 

tridens (l\ttby) 

tridentifrons Fitch 

H. quindecirnmaculata Mulsant 

H. quinques ignata quinquesignata 

(KirLY) 

coccineu Casey 

leporina Mulsant 

mulsanti LeConte 

pugetar'.a Casey 

quadra,"ia Casey 

subsiru.lis Casey 

uteana Casey 

uernis Oasey 

H. quinquesignata ambigua LeConte 

obliqua Casey 

punctulata LeConte 

H. sinuata crotchi Casey 

disjunc;la Timberlake 

fontinaiis Casey 

straminea E.A. Chapin 

H. sinuata spuria LeConte 

cornpler Casey 

H. tredecimpunctata tibialis (Say) 

AK 

YK 

NT BC AB SK 

_BC 

NT _BCAB 

_BCABSK 

_YK_BCABSK 


YK 

AK YK NT BC 

AK YK 

AK YK 

MB ON 

MB 

MB 

_PQ 

PQ 

ON PQ 

PQ 


NT_ABSKMB 

BC 

_BC 

_BCABSK 

_BCAB 

NT 

_BC 

BC 

BC 

BC 

AB 

MB 

NB 


_SK_ON 


AB 

SK 

NT BC AB SK 

MB 

MB ON 

NS 

NB_PE 

LB 


234

Coccineliidae 

H. variegata (Goeze)t 

H. washingtoni Timberlake 

BC 

9*PQ_ 

Genus ANATIS Mulsant 

Key to species: Watson (1976) 

A. Iabiculata (Say) 

canadensis Provancher 

coseyl Westcott 

q uinde c i m p u n c t a ta (Oliv iet) 

A. lecontei Casey 

A. mali (Say) 

borealis Belicek 

signaticollis Mulsant 

A. rathvoni (LeConte) 

AKYKNT 

BC AB 

BC AB 

BC 

AB 

SK 

SK 

MB ON PQ 

MB ON PQ 

NB 

NB NS PE 

NF 

Genus MYZIA Mulsant 

Neomysia Casey 

Paramysia Reitter 

M. interrupta (Casey) 

coseyi (Timberlake) 

M. pullata (Say) 

montana (Casey) 

nolons (Randall) 

randalli (Casey) 

M. subvittata (Mulsant) 

hornii Crotch 

oregona (Casey) 

YK 

BC 

BC 

AB 

AB SK 

AB 

MB ON PQ 

NB NS 

LB 

Genus CAI-VIA Mulsant 

Anisocaluia Crotch 

C. quatuordecimguttata (Linn6)* 

bicordifera (Casey) 

cardisce (Randall) 

comple*a Nunenmacher 

elliptica (Casey) 

hesperica (Crotch) 

incarnata (Kirby) 


obliqua (Randall) 

postplagiata (Casey) 

quadrisignata (Casey) 

sicordi (Nunenmacher) 

simllis (Randall) 

uniformis (Casey) 


uictoriana (Casey) 

AK YK NT 

BC 


LB 

NF 

Genus ADAI,IA Mulsant 

A. bipunctata ilinn6)* 

annectans (Crotch) 

bioculata (Say) 


disjuncta (Randall) 

duplicata Leng 

frigido (Schneider) 

humeralis (Say) 

ludouicae Mulsant 

mel anop I eur a (LeConte) 

ocellata Johnson 

opthalmica Mulsant 

ornatella Casey 

ouipennis Casey 

postica Johnson 

sexpustulata Johnson 

transuersalis Casey 

AK YK NT BC 

AB 

SK 

MB ON PQ 

NB NS 

LB 

NF 

Genus COCCINELLA Linn6 

Key to North American species: Brown (7962b) 

235

Coccinellidi,'e 

C. alta W.J. Brown 

sutura'.is Casey 

C. californica Mannerheim 

melanc'collis Johnson 

C. fulgida V/atson 

C. hieroglyphica humboldtiensis 

Nunenmacher 

C. hieroglyphica kirbyi Crotch 

tricuspis Kitby 

C. hieroglyphica mannerheimi 

Muklant 

C.johnsoni Casey 

C. monticola Mulsant 


bigutta.ta Johnson 

confluenta Johnson 

impres';a Casey 

lacustris LeConte 

neuadica Casey 

postica Johnson 

sellica Johnson 

C. novemno:ata Herbst 

confluenta Fitch 

conjunt:ta Fitch 

degenet'Casey 

diuisict'llis Fitch 

francis,:ana Mulsant 

inaequulis Fitc}:' 

oregontt Casey 

paruan,,aculato Fitch 

C. prolongata prolongata Crotch 

C. septempr..nctata Linn6J 

C. transvers oguttata richardsoni 

W.J. Brown 

interrulta Fitdn 

quinquznotata I{ttby 

zetterstzdti Madet 

C. trifasciat,r perplexa Mulsant 

C. trifasciatl subversa LeConte 

barda T,eConte 

euginii Mtisant 

juliana Mulsant 

praedicta Mad.et 

C. undecimp,unctata undecimpunctata 

Linn5t 

AK 

TY 

_YK 

AK YK 

AK YK 

NT 

NT 

NT 

AX 

AK_NTBCAB 

BC AB 

BC AB 

BC 

BC 

BC 

BC 

AB 

SK 

SK 

MB ON 

MB 

ON PQ 

PQ 


BC A.B 

_BCABSKMBONPQ 



_BC 

BC 

NB NS 

MB ON PQ Nn f,IS - 

NB 

NB 


ON PQ 

NS 

NS 

NB NS PE 

_LBNF 

in Ne 

NF 

Genus PRO )YLAEA Mulsant 

P. quatuord,:cimpunctata (Linn6)t 

ON PQ 

NB 

Genus CYC.i,ONEDA Crotch 

C. munda (Say) 

C. polita Casey 

flauaT'lr'berlake 

_BCABSK 

MB ON 

PQ 

Genus OLLr\ Casey 

O. v-nigrum (Mulsant) 

abdominalis (Say) 

arizonae Casey 

fenestra'.lis Casey 

minuta Casey 

plagiaht Casey 

soyi (Clotch) 

semiluri.aris Johnson 

sobrina Casey 

BC A3 SK MB ON 

Genus NEO.FIARMONIA Crotch 

Agrabiu Casey 

Harmottiaspis Casey 

Neoharnonia Casey 

236

Endomychidae 

N. venusta venusta (Melsheimer) 

centralis Casey 

dissimila Blatchley 

fattigi B\atc}:.ley 

notulata (Mulsant) 

Genus MULSANTTNA Weise 

Pseudocleis Casey 

Key to species: Chapin (1985) 

M. hudsonica (Casey) 

M. picta (Randall) 

blanchardi (Johnson) 

c on cinnata (Melsheimer) 

contert(r (Mulsant) 

impictipennis (Weise) 

minor (Casey) 

nubilata (Casey) 

AK YK 

_YK 

ON 

NT BC AB SK MB ON PQ NB NS PE LB 

_BCABSKMBONPQNBNS 

Tribe Psylloborini 

Genus PSYLLOBORA Dejean 

P. borealis Casey 

deficiens Casey 

P. vigintimaculata (Say) AK YK NT 

obsoleta Casey 

separata Casey 

taedata LeConte 

BC 


NF 

Subfamily EPILACHNINAE 

Key to New World species: Gordon (1975) 

Tribe Epilachnini 

Genus EPILACHNA Dejean 

E. varivestis Mulsant 

ceruina Mulsant 

corrupta Mulsant 

cuprea Cockerell 

genuina Mulsant 

juncta Johnson 

maculiuentris Bland 

toweri Johnson 

oN PQ 

Family ENDOMYCHIDAE handsome ftrngus beetles 

J.M. Campbell 

The family Endomychidae is found in most 

regions of the world, but the majority of species 

occur in tropical areas. The range of 15 species 

extends northward into Canada. Adults and larvae 

may be collected on soft fungi, in leaf litter, or 

under bark with fungal growth. One species, 

Mycetaea subterranea (M. hirta in most publications), 

is a pest of stored products. Most species 

feed on fungr. 

Because of their moderately large size and often 

bright coloration, many North American species are 

taxonomically well known. However, the family has 

never been revised for North America. Strohecker 

(1953) reviewed the genera of the world and published 

a catalog of the North American species 

(1986). Hatch (1962) treated the genera and species 

that occur in the Pacific Northwest. A modern revision 

of the North American species of this family is 

needed. 

NT (1); BC (7); AB (2); SK (2); MB (5); ON (10); PQ 

(8);NB (2);NS (6);PE (1);NF (2) ' I(2) 

237

Endomychidae 

Subfamily MYCETAEINAE 

Tribe Mycetaeini 

Genus SYMBIOTES Redtenbacher 

S. duryi Blatchley 

Iacustris Casey 

oblong,ts Casey 

pilosus Casey 

waltoni Dury 

S. gibberosr;.s .(Lucas)t 

duryiWalton 

montanus Casey 

Genus MYCETAXA Stephens 

M. subtenanea (Fabricius)f 

fumata Stephens 

hirta (Ivlarcham) 

-oN 

-oN 

_BC_ONPQ_NSPE_NF 

Genus RHANIDEA Strohecker 

Rhanis LeConte 

Tribe Leiestini 

Key to North American species: Blaisdell (1931) 

R. unicolor Zieglerr 

apicalis (Melsheimer) 

- ON PQ 

haemo,"rhoidalis (Gu6rin-M6nevi11e) 

Genus STE'IHORHANIS Blaisdell 

S. borealis ]]laisdell - BC 

Genus PHYMAPHORA Newman 

P. californica Horn 

P. pulchella Newman 

- BC 

cr a s s icor nis ( Melsheimer) 

MB ON PQ - NS 

p u nc t i r:' o I I i s (Zi e gl er' ) 

Subfamily STENOTARSINAE 

TYibe Stenotarsini 

Genus DA]riAI Reiche 

D. testacea iZiegler) 

fer r ug i ne a (Gu6rin-M6neville ) 

lzleo (lvlelsheimer) 

piloso i.Melsheimer) 

_ONPQ_NS 

Genus LYC'OPERDINA Latreille 

(Subgenus (]OLGIA Mulsant) 

Subfamily ELIMORPHINAE 

Tribe Lycoperdinini 

L. fermgine a LeConte 

angulata (Say) 

- BC AB SK MB ON PQ - NS 

g lab rala Gu6rin-M6neville 

TYibe Mycetinini 

Genus APHORISTA Gorham 

A. laeta (LeConte) 

A. vittata (Fabricius) 

distincta (Say) 

lineata (Olivier) 

ouipenizis Casey 

---:"--meo*pe----- 

238

Lathridiidae 

Genus MYCETINA Mulsant 

M. idahoensis Fall 

M. perpulchra (Newman) 

q uadrig uttata (Melsheimer) 

BC 

MB ON PQ 

NB 

NS 

Subfamily ENDOMYCHINAE 

Tribe Endomychini 

Genus ENDOMYCHUS Panzer 

E. biguttatus Say 

E. limbatus (Horn) 

endomychoides (Fall) 

_NTSKMB 

BC AB 

ON PQ 

NBNS_ 

NF 

Family LATHRIDIIDAE minute brown scavenger beetles 

Y. Bousquet 

In Canada and Alaska 55 species of lathridiids 

have been recorded. These species are usually 

found on foliage, in rotten wood, under logs, and 

under the bark ofdead trees, as well as in fungi and 

nests of social Hymenoptera, birds, and mammals. 

A number of species, including Cartodere constricta 

and Aridius nodifer, are regularly found in stored 

products, especially those that are damp and moldy. 

Adults and larvae feed on fungr, particularly molds. 

Lathridiidae is another family of Cucujoidea 

that has been poorly studied in North America. 

Most of the genera, particularly those of the subfamily 

Corticariinae, are in need of modern revision. 

The generic classifrcation of the Lathridiinae used 

here follows that of Walkley (1952). 

AK (19);YK (5);NT (6); BC (3a); AB (9); SK (8); MB 

(17); ON (22); PQ (22); NB (6); NS (8); PE (1); LB 

(1);NF (a) . H (6);I (15) 

Subfamily I"ATHRIDIINAE 

Genus DIENERELLA Reitter 

Microgramme Walkley 

D. arga (Reitter)t 

D. cordicollis (Mannerheim) 

D. costulata (Reitter)t 

D. frliformis ( Gyllenhal)f 

D. filum (Aub6)t 

D. rufrcollis (Marsham)t 

p ulicar ia (Melsheimer) 

AI{ 

---ec- _BC 

oN PQ 

_MBONPQ 

SK MB 

_MBONPQ 

ON PQ 

NB 

_NS 

PE 

NF 

Genus ENICMUS C.G. Thomson 

E. aterrimus Motschulsky 

ferrugineus Belon 

opaculus LeConte 

E. cordatus Belon 

E. fictus FaIl 

E. mimus Fail 

E. nitens FalI 

E. tenuicornis LeConte 

laticollis LeConte 

AK 

AK 

-BC 

NT BC 

NT BC 

BC 

oN PQ 

AB ix ms o* Pe 

AB SK MB 

y_ yuoNpe_ _ _ _ _ 

Genus THES Semenov 

T. bergrothi (Reitter)f 

tricarinatus (W.J. Brown) 

BC 

SK 

MB 

NB NS 

Genus LATHRIDIUS Herbst 

Conithassa C.G. Thomson 

L. consimilis Mannerheim* 

par allelocollis Mannerheim 

L. crenatus LeConte 

AK 

BC 

239

Lathridiidae 

L. hirtus Gyllenhalf 

L. minutus (Linn6)+ 

porcatus Herbst 

reflextls LeConte 

L. nigritus (Fall) 

L. protensi :oliis Mannerheim 

quadr :collis Mannerheim 

sobrinus Mannerheim 

suspectus (Fall) 

AK 

_BC 

_BCAB 

_BC 

BC 

SK 

MB ON PQ 

MB 

Genus STEPHOSTETHUS LeOonte 

S. breviciavis (FalI) 

S. cinnamopterus (Mannerheim) 

S. costicollis (LeConte) 

armat:zlus (Fall) 

S. curtulus (Mannerheim) 

S. lardariuir (DeGeer)f 

S. liratus (LeConte) 

carinil'er (Fa\\') 

trilobatus (Motschulsky) 

S. montants (Fall) 

laeuior (W .J. Brown) 

AK 

AK 

AK 

AK 

AK YK 

_BC 

_BC 

BC 

NT BC 

AB 

SK 

-oNPQ 

MB ON 

MB 

PQ 

PQ 

NB NS 

wS 

NF 

Genus ADISTEMIA Fall 

A. watsoni lWollaston)t 

ON PQ 

Genus CAIi,TODERE C.G. Thomson 

Coninomus C.G. Thomson 

C. constrict,a (Gyllenhal)t 

sculpti.lis (LeConte) 

BC 

AB 

SK 

MB ON 

PQ 

NB 

NF 

Genus ARI )IUS Motschulsky 

A. nodifer ( Westwood)f 

BC 

MB ON 

PQ 

Genus COI,,TICARIA Marsham 

C. canalicu ata Mannerheim 

C. columbia. FalI 

C. dentigera LeConte 

C. dentiventris Poppius* 

C. fenestralis (Linn6)* 

deleta Mannetlr'eim 

denticulata I{trby 

kirbyi LeConte 

obtusa LeConte 

rugulo.;a LeConte 

C. ferrrrginea Marshamx 

C. Iinearis (Paykull)x 

C. orbicollis Mannerheimx 

munsteri Stand 

C. parallela Fall 

C. pubescers Gyllenhal-i 

grosso LeConte 

C. salpingoides Motschulsky 

C. serrata (Paykull)f 

octodentata (Say) 

C. serricollis LeConte 

C. spinulosa Mannerheim 

C. valida Fall 

C. varicolor Fall 

AK 

Subfamily CORTICARIINAE 

AK YK 

AX 

AK 

AK 

AK YK 

AK 

YK 

_BC 

_BC 

NT 

NT 

_BC 

_BC 

_BC 

_BC 

-nc 

_BC 

BC 

AB 

MB 

MB 

-on 

ON PQ 

ON 

PQ 

PQ- 

_ _ luIu 

Genus MEL,ANOPHTHALMA Motschulsky 

M. americana (Mannerheim) 

distinguenda auct. amer. 

morsa',LeConte) 

pusilla (Melsheimer) 

rufula (.LeConte) 

BC 

ON PQ 

240

Byturidae 

M. pumila (LeConte) 

M. villosa (Zimmermann) 

Genus CORTICARINA Reitter 

C. alberta (Fall) 

C. cavicollis (Mannerheim) 

C. exigua (Mannerheim) 

C. fuscula (Gyllenhal) 

americana auct. amer. 

compt(r (LeConte) 

expansa (LeConte) 

grata (LeConte) 

C. herbivagans (LeConte) 

leuis (LeConte) 

similata a:uct. 

_BC_ON 

_BC 

AB 

-BC-ONPQ-NS 

AK 

AK l'K NT BC 

_BC 

MB ON PQ 

Genus CORTINICARA Johnson 

C. gibbosa (Herbst)t 

Genus CORTILENA Motschulsky 

C. picta (LeConte) 

_BCABSKMBONPQNBNS 

-oNPQNBNS 

Family BIPI{YLLIDAE false skin beetles 

J.M. Campbell 

Only four species of Biphyllidae are known to This family is poorly known taxonomically, and 

occur in North America, one of which, Diplocoelus the North American species have not been revised. 

brunneus, is found in eastern Canada. Little is 

known about the habits of this species; adults are ON (1);PQ (1);NB (1) 

found in fungus or under bark. 

Genus DIPLOCOELUS Gu6rin-M6neville 

Marginus LeConte 

Tlibe Biphyllini 

D. brunneus (LeConte) - oN PQ NB 

Family BYTURIDAE fouitworm beetles 

J.M. Campbell 

Two species of Byturidae are found in North sion by Springer and Goodrich (1983) of the North 

America of which one, Byturus unicolor, the rasp- American species of the family recognized only one, 

berry fruitworm, is common and widely distributed somewhat variable species. 

across Canada. Adults ofthis species feed on leaves 

and flowers, and the larvae feed on the receptacle of 

the red raspberry, Rubus idaeus. 

Byturus unicolor has been known under a vari- aK (1); yK (1); NT (1); BC (1); AB (1); SK (1); MB 

ety of Latin and common names, but a recent revi- (1); ON (1); PQ (1); NB (1);NS (1);NF (1) 

Genus BYTURUS Latreille 

Horticola Lindemann 

Tribe Byturini 

B. unicolor Say AK YK NT BC AB SK MB ON PQ NB NS NF 

americanus Gemminger & Harold 

247

Mycetophagidae 

baker;'.Barber 


punctatus Casey 

rubi Eiarber 

sordictus Barber 

Superfamily Tenebrionoidea 

Family MYCETOPIHGIDAE hairy fungus beetles 

Y. Bousquet 

Of the 26 species of mycetophagids recorded 

from Nc,rth America, 15 are known to occur in 

Parsons (I975b) has revised the mycetophagids 

of theNearcticfauna. 

Canada. These beetles are found mainly in decaying 

vegetable matter, in fungi, and under the bark 

of rotten logs. A few species, such as Typhaea stercorea, 

are occasionally found in buildings. As the 

Latin name suggests, members of this family are BC (6);aB (4); SK (2); MB (7); ON (10); PQ (12); NB 

mycetophagous. (2);NS (1); PE (1) . I (1) 

Genus MY,IETOPHAGUS Hellwig 

(Subgenus MYCETOPHAGUS s. str.) 

M. californicus Horn - BC 

prouer,,sis Casey 

M. flexuosrrs Say MB ON PQ 

M. punctat us Say MB ON PQ 

M. serrulat,us (Caseyt MB ON PQ 

pictus (.Casey) 

quadralius Casey 

subderzressus (Casey) 

tribaheatus Casey 

(Subgenus ILENDUS Casey) 

M. distinctus Hatch - BC AB SK MB 

M. pluripunctatus LeConte AB - MB ON PQ 

(Subgenus PARILENDUS Casey) 

M. quadriguttatus P.W.J. Miillerf 

bimac ulatus Melsheimer 

- BC AB SK MB ON PQ NB 

bipust.ulatus Melsheimer 

b isig n at us Melsheirner 

uariegatus C.R. Sahlberg 

(Subgenus GRATUSUS Casey) 

M. plurigu,tatus LeConte - BC 

franciscanus Van Dyke 

M. tenuilasciatus Horn - BC AB ON PQ 

notalulus ( Casey, 

Genus LITARGUS Erichson 

(Subgenus AJ-ITARGUS Casey) 

L. balteatus LeConte 

infulatus LeConte 

- ON PQ 

tr(l,nsuersus LeConte 

(Subgenus PARALITARGUS Casey) 

L. didesmus (Sayt PQ 

asperulus Casey 

(Subgenus LITARGUS s. str.) 

L. sexpuncbatus tSay) - ON PQ 

obsolescens Casev 

242

Tetratomidae 

(Subgenus TILARGUS Casey) 

L. tetraspilotus LeConte MB ON PQ 

Genus TYPHAEA Stephens 

T. stercorea (Linn6)f - BC - ON PQ NB NS PE 

fumata (Linn6) 

Genus THRIMOLUS Casey 

T. minutus Casey PQ 

duryi Casey 

FamilyTETRATOMIDAE tetratomidbeetles 

L. LeSage 

The family Tetratomidae includes nine species on beetles, such as Blatchley (1910), Chagnon and 

in Canada and Alaska. Both adults and larvae live Robert (1962), Hatch (1965), and Arnett (1968). 

in the fruiting bodies of higher fungi and are myce- Young (1976) recently transferred the genus 

tophagous. 

Pectotoma, which had been previously placed in this 

Previously considered a subfamily of family, to Scraptiidae. The survey of the 

Melandryidae, the tetratomids were raised to family Tetratomidae of Quebec by Chantal (1985) includes 

status by Crowson (1955). This ranking is accepted genera placed here in Melandryidae. 

by most recent authors, including Miyatake (1960) 

and Lawrence (1982a). There is no recent revision 

ofthefamilyinNorthAmerica. Generaandspecies aK(1); YK(1); nC(2); AB(1);SK(1); ON(5);PQ 

can be identifred tentatively by using general books (7); NB (1);NF (1) . I (1) 

Genus TETRATOMA Fabricius 

Subfamily TETRATOMINAE 

T. concolorleConte - YK - BC AB SK 

T. truncorum LeConte - ON PQ 

Genus ABSTRULIA Casey 

A. tesselata (Melsheimer) - ON PQ 

maculata Casey 

A. variegata Casey PQ NF 

Genus EUPISENUS Casey 

E. elongatus (LeCon[e) AK BC 

Genus PISENUS Casey 

P. humeralis (Kirbyt 

concolor (Kirby) 

- ON PQ 

ruficornis (LeConte) 

Genus PENTHE Newman 

Pytocis Castelnau 

P. obliquata (Fabricius) 

P. pimelia (Fabricius) 

anglica (Gmelin) 

funerea Newman 

lesueuri (Castelnau) 

morio (Fabricius) 

Genus MALLODRYA Horn 

M. subaenea Horn 

Subfamily PENTHINAE 

-oNPQ 

-oNPQNB 

prl 

243

Ciidae 

Family CIIDAE minute tree-fungus beetles 

J. McNamarq, 

The family Ciidae, which consists of small beetles, 

comprises 29 species in Canada and Alaska. 

The larvae and adults of nearly all species feed on 

fungal h'7phae comprising the mycelia and especially 

the fruiting bodies of wood rotting Basidiomycetes. 

Several l;pecies are found under bark, in dead wood, 

or in the galleries ofbark beetles. 

Lawrence (1971) revised the North American 

species and also published a catalog of the species 

(1982c). 

eK (6); xT (6); BC (19); AB (10); SK (5); MB (14); 

ON (16);PQ (17);NB (5);NS (6);NF (6) ' I (1) 

Subfamily CIINAE 

Tlibe Ciini 

Genus CIS Latreille 

Xestocis Casey 

C. americanus Mannerheim 

forsteri Abdullah 

frosti Dury 

hatch, Lawtence 

insolerc (Casey) 

minot (Hatch) 

nitidt s (Hatch) 

oweni (.Hatch) 

serricollis Drty 

strigulosus (Hatch) 

C. angustus Hatch 

C. biarmatus Mannerheim 

C. castlei (Dury) 

C. creberrimus Melli6 

C. ephippiatus Mannerheim 

moznt'ttei (Dtry) 

C. fuscipes Melli6 

atripennis Me1li6, 

caroli:zae Casey 

cheurolatii Melli6 

dubius Melli6 

impressus Casey 

pallens Casey 

C. horriduius Casey 

mormonicus Casey 

C. hystriculus Casey 

C. levettei (Casey) 

C. maritimus (Hatch) 

C. pistorius Casey 

C. striolatus Casey 

fraternus Casey 

macilentus Casey 

C. subtilis Mel1i6 

confusus Blatchley 

julichiDrry 

wenzeli Dtry 

C. tridentatus Mannerheim 

ednae (I].atch) 

reflexus (Hatch) 

AK 

AK 

AK 

AK 

_BC 

_BC 

BC 

-nc 

NT BC 

NT 

BC 

AB 


-o* 

_:* 

-BC 

_BCABSKMBONPQ 

_BCMB 

NT-ABSKMBONPQ 

NTBCAB_MBONPQ 

BC 

AB 

SK 

MB ON 

MB ON 

PQ 

PQ 

PQ 

PQ 

_NS 

NB 

NS 

NS 

NS-- 

NF 

NF 

NF 

Genus ENNEARTHRON Melli6 

E. spencen (Hatch)t 

-BC 

Genus DOLICHOCIS Dury 

D. indistinctus Hatch 

D. manitoba Dury 

-BC 

NT BC AB 

MB ON 

PQ 

PQ 

NB- 

244

Melandryidae 

Genus ORTHOCIS Casey 

O. punctatus (Mel1i6) AK 

aterrimus Casey 

NT 

BC 

AB 

MB ON PQ 

NF 

Genus STRIGOCIS Dury 

S. opacicollis Dury 

ON 

Genus HADRAULE C.G. Thomson 

Maphoca Casey 

H. blaisdelli (Casey) 

H. elongatula (Gyllenhal)t 

Genus PLESIOCIS Casey 

BC 

NB 

P. cribrum Casey 

BC 

AB MB PQ 

Genus CERACIS Me11i6 

Key to North American species: Lawrence (1967) 

C. sallei (Melliel 

C. singularis t Duryt 

C. thoracicornis t Zieglert 

bifoueata Dtry 

laminifrons (Casey) 

mellyi (.Mel1i6) 

oblonga (Blatchley) 

picea (Casey) 

pumicdta (Meili6) 

unicolor (Casey) 

oN PQ 

- - - r,rs3Npe- - - - - 

Genus SULCACIS Dury 

S. curtula tCasey) 

criddlei (Drry) 

cylindrica (Dury) 

montana (Casey) 

nigra Dury 

sorror (Casey) 

Genus MALACOCIS Gorham 

Brachycis Casey 

M. brevicollis (Casey) 

BC AB SK MB PQ 

MB ON PQ 

NF 

Genus OCTOTEMNUS Melli6 

O. laevis Casey 

denudatus Casey 

Subfamily OROPHIINAE 

Tlibe Orophiini 

AK BC AB SK MB ON PQ NB NS 

NF 

Family MELANDRYIDAE 

false darkling beetles 

L. LeSage 

Melandryids are common throughout the world 

and include about 100 genera and 1200 species, of 

which at least 55 occur in Canada and Alaska. 

Adults live under bark, in fungi, or in flowers. 

Larvae are carnivorous or feed on decaying plant 

material; they live under bark, in dead logi, or in 

stumps; a few are found in leaf litter, but detailed 

life-history studies, such as that of Hoebeke and 

McCabe (1977), are seldom published. 

The taxonomy of the family is in a state of disarray 

at all levels. Generic keys from general books on 

beetles work inconsistently. Up-to-date revisions at 

the species level are badly needed. Chantal (1985) 

keyed out the species of Hallomeninae and 

Eustrophinae from Quebec. At this time, no expert is 

working on the family in North America. The classification 

used here corresponds to the arrangement suggested 

by J.F. Lawrence (personal communication). 

aK (4);YK (2);NT (3); BC (27); AB (10); SK (5); MB 

(e); ON (27);PQ (42); NB (11); NS (10); LB (3); NF 

(13) . H(3);I(2) 

245

Melandryidre 

Subfamily IIALLOMENINAE 

Genus HALLOMENUS Panzer 

H. binotatu,; (Quenselti PQ 

H. debilis L,:Conte PQ 

ziger Fialdeman 

H. punctulatus LeConte AK BC - ON PQ 

basalis Mannerheim 

H. scapulars Melsheimer - BC - ON PQ 

Iuridus Haldeman 

ruficor,zis Melsheimer 

H. serricornis LeConte - BC AB ON PQ 

Subfamily EUSTROPHINAE 

Genus EUS'IROPHINUS Seidlitz 

E.bicolor(Fabriciusr - ON PQ 

indistittctu s ( LeConte) 

E. confinis t LeContet - BC - SK MB ON PQ 

Genus EUSTROPHUS ll I iger 

E. tomentosus Say - BC - ON PQ 

niger Nlelsheimer 

Genus SYNSTROPHUS Seidlitz 

S. repandus t Horn t - BC MB ON PQ 

Genus HOL,OSTROPHUS Horn 

H. bifasciat rs (Sayt - ON PQ 

q uadr i:nac ulalzs (Melsheimer) 

H. discolor l{orn PQ 

H. impressi':ollis tl-eConter - BC 

Genus ORCHESIA Latreille 

Key to North American species: Lalibert6 (1966) 

Subfamilv MELANDRYINAE 

Trile Orchesiini 

O. castanea (Melsheimer) - BC MB ON PQ - NS NF 

O. cultrifonnis Laiibert6 - NT SK MB - PQ - NS 

O. ornata Horn - BC 

O. ovata La ibertti ON PQ 

Genus MIC ROSCAPHA LeConte 

M. clavicon.is LeConte PQ 

Genus LEDERIA Reitter 

Crioscctpha Flotn 

L. arctica (llorn) AK BC 

Genus EMNIESA Newman 

Tlibe Melandryini 

E. connectens Newman - ON PQ - NS - LB NF 

maculcrta LeConte 

E. labiata (,(iay) - ON PQ NB - NF 

E. stacesmi-,hi Hatch - BC 

E. testacea leeperi Malkin - BC 

Genus MET,ANDRYA Fabrici us 

NeoemtnesaHatch 

M. blackmani Hatch PQ 

M. striata Say - BC AB SK - ON PQ NB 

bicolor Melsheimer 

246

Melandryidae 

diuersa Mulsant 

excauata Haldeman 

thoracica Melsheimer 

Genus PHRYGANOPHILUS C.R. Sahlberg 

P. collaris LeConte NT BC AB - MB ON PQ NB 

Genus ZILORA Mulsant 

Key to North American species: Mank (1938) 

Tlibe Hypulini 

Z. canadensis Hausen PQ 

Z. hispida LeConte - YK - BC AB ON PQ NF 

Z.nudaProvancher 

PQ 

Z. occidentalis Mank - BC 

Genus HYPTILUS Paykul. 

H. concolor (LeConte) PQ 

Genus MICROTONUS LeConte 

M. sericans LeConte - '' ? - 

Genus SYMPHORA LeConte 

Trotomma LeConte 

S. flavicollis (Haldeman) MB ON PQ - NS 

S. rugosa (Haldeman) - ON PQ - NS 

Genus PROTHALPIA LeOonte 

P. holmbergi (Mannerheim)* AK BC 

fulrninans (LeConte) 

P. undata LeConte - ON PQ - NS NF 

Genus AMBLYCTIS LeConte 

A. praeses LeConte 

Genus XYLITA Paykull 

X. livida (C.R. Sahlberg)* 

X. laevigata (Hellenius)* 

decolorata (Randall) 

Tlibe Serropalpini 

_YK_BCAB_PQNBNS_LBNF 

-BCAB-MBONPQNB_NF 

PQ 

Genus SPILOTUS LeConte 

S. quadripustulatus (Melsheimer) PQ 

Genus RUSHIA Forel 

Carebara LeConte 

R. californica FaIl 

californica Schaeffer 

schaefferi Csiki 

_BC 

Genus SCOTOCHROIDES Mank 

S. antennatus Mank 

Genus SCOTOCHROA LeConte 

Key to species: Mank (1939b) 

S. atra LeConte 

S. basalis LeConte 

S. buprestoides (Kirby) 

Genus ENCHODES LeConte 

E. sericea (Haldeman) 

_-_:"AB__;.::il___Ni 

_BCABSKMBONPQ 

PQ 

247

Mordellida: 

Genus SEF,ROPAIPUS Hellenius 


S. coxalis Nlank 

S. obsoletur; Haldeman 

perfoueatus Mank 

S. substriatus Haldeman 

Mank (1939a) 

T_ 

_BC 

_BC 

NT BC 

MB 

ON PQ 

-PQ 

ON PQ 

NB NS 

NB 

LB 

NF 

NF 

Genus MYSTAXUS Kugelann 

M. simulator (Newman) 

trifasc iatus (Melsheimer) 

ON PQ 

NB NS 

NF 

Genus DIR,IA-EA Fabricius 

D. liturata '.LeConte) 

quadrtmacularo (Say) 

D. horiei Hatch 

BC 

AB SK 

ON 

PQ 

NB 

Genus PHLOEOTRYA Stephens 

P. vaudoueri Mulsantt 

fusca (.LeConte) 

PQ 

NS 

Genus ABDERA Stephens 

A. frrma Ha.tch 

BC 

Genus ANIIIOXYA Mulsant 

A. glaucula LeConte 

ON 

Genus OSPFfYA Illiger 

Nothut Olivier 

O. varians (LeConte) 

Subfamily OSPFIYINAE 

Tlibe Osphyini 

oN PQ 

Family MORDELLIDAE tumbling flower beetles 

J. McNamara 

Severrty species and subspecies of mordellids 

have beer recorded in Canada. The adults are small, 

humpbacked beetles, and their most distinctive feature 

is their long, pointed abdomen. They are commonly 

found on umbelliferous flowers, on which they 

feed. Larvae are carnivorous or feed on plant material. 

Sorne are found boring in rotten wood, others 

are leaf and stem miners, but very few are known. 

Liljetrlad (1945) revised the North American 

species oI'Mordellidae; Ermisch (1951) treated the 

genera of the world; and Bright (1986) pr:Lblished a 

catalog of the Nearctic species. Neverlheless, a 

modern revision would be useful. Members of 

Anaspini, which used to be included within 

Mordellidae, are now in the family Scraptirdae. 

YK (1); NT (6); BC (18); AB (10); SK (12); MB (37); 

ON (49);PQ (29);NB (14);NS (7);PE (1) 

Genus TOMOXIA Costa 

T. borealis ( LeConte) 

T. inclusa Lr:Conte 

T. lineella LeConte 

T. serval (Sz,y) 

T. triloba (Say) 

T. undulata (Melsheimer) 

Tribe Mordellini 

_ =_:"Y_;;lNifrilrs_ _ - 

Genus MORDELLA Linne 

Sphalera LeConte 

248

Mordellidae 

M. albosuturalis Liljeblad 

M. atrata atrata Melsheimer 

M. atrata lecontei Csiki 

itorata LeConte 

M. brevistylis Liljeblad 

M. grandis Liljeblad 

M. hubbsi Liljeblad 

M. marginata lineata Melsheimer 

M. marginata marginata Melsheimer 

M. melaena Germar 

M. quadripunctata (Say) 

_NT 

_NT 

NT 

BC 

BC 

BC 

BC 

BC 

BC 

BC 


-ONPQNB 

AB 

AB SK 

-o*--- 

MB ON PQ NB 

MB ON PQ 

MB ON 

NS 

NS 

Genus GLIPA LeConte 

G. bidentata (Say) 

G. octopunctata (Fabricius) 

G. oculata (Say) 

jouialis (LeConte) 

_ONPQ 

-oN 

-oN 

Genus CONALIA Mulsant & Rey 

C. melanops Ray 

ON PQ 

Tribe Mordellistenrnr 

Genus MORDELLISTENA Costa 

M. aemula LeConte 

M. aethiops Smith 

M. andreae ancilla LeConte 

uarians LeConte 

M. andreae andreae LeConte 

grammica LeOonte 

M. andreae ustulata LeConte 

M. arida Leconte 

rutila Liljeblad 

M. aspersa (Melsheimer) 

M. bicinctella LeConte 

M. cervicalis LeConte 

auricorna Helmuth 

erilis Liljeblad 

picicornis LeConte 

M. convicta LeConte 

M. discolor (Melsheimer) 

cinereofasciata Smith 

wolcotti Liljeblad 

M. divisa LeConte 

M. downesi Hatch 

M. errans Fall 

M. frosti Liljeblad 

M. fulvicollis (Melsheimer) 

M. fuscata (Melsheimer) 

M. fuscipennis (Melsheimer) 

pityptera LeConte 

M. gigas Liljeblad 

M. guttulata Helmuth 

M. hebraica LeConte 

M. impatiens LeConte 

M. incommunis Liljeblad 

M. infima LeConte 

tiara Ray 

M. liljebladi Ermisch 

egregia Liljeblad 

M. limbalis (Melsheimer) 

M. liturata (Melsheimer) 

M. lutea (Melsheimer) 

M. marginalis (Say) 

M. morula LeConte 

M. nigricans Melsheimer 

nigenima Helmuth 

M. nunenmacheri Liljeblad 

NT 

- Y 

BC 

AB 

_MB 

sK MBoN - - - - 

-- - 

SK 

MB 

ON 

ON 

'_ _:'I_ illgN;: 

_NT 

_ _ _ yuSNPQ 

BC 

BC 

BC 

BC 

MB 

MB-PQ 

-oN 

_ONPQ 

-oNPQ 

_ONPQ 

-oN 

MB ON 

MB ON 

MB 

MB ON 

PQ 

SK MB ON PQ 

ON 

ON 

-MBONPQ 

SK 

_MBONPQ 

NB 

NB 

NB 

_ft 

NB 

NS 

NS PE 

249

Rhipiphoridae 

M. ornata (trfelsheimer) 

decoreLa LeConte 

pectora,tis (LeConte) 

M. pallipes fimith 

M. picilabris Heimuth 

M. pustulatr,r (Melsheimer) 

M. quadrinobata Liljeblad 

M. scapulais (Say) 

biplagiuta Helmuth 

M. semiusta LeConte 

M. sericans .Fall 

M. sexnotata Dury 

falli Cslki 

festiua Tall 

M. smithi Dury 

M. suturella Helmuth 

M. syntaenia Liljeblad 

M. testacea.Blatchley 

M. tosta LeConte 

M. trifasciata (Say) 

amica L,eConte 

lepidula LeConte 

nigricollis Helmuth 

M. unicolor LeConte 

M. vilis (LeClonte) 

-:-ecAB 

_BCAB 

NT 

BC 

SK 

i* 

SK 

SK 

MB ON 

PQ 

MB ON 

MB 

MB ON PQ 

-oNPQ 

MB ON PQ 

MB 

MB 

-oN 

NB 

_MBON 

_MB 

_MBNB 

ON 

_MBONPQNB 


MB 

MB ON 

NS 

Genus GLIPOSTENODA Ermisch 

G. ambusta iLeConte) 

diuersa (Ray) 

BC 

ON 

PQ 

Family RHIPIPHORIDAE wedge-shaped beetles 

J.M. Campbell 

The family Rhipiphoridae contains about 400 

species, of which 10 are known to occur in Canada. 

Larvae of the subfamily Rhipiphorinae are parasitic 

on the larvae of bees and wasps; the frrst instars are 

endoparasites and the last ones ectoparasites. 

Adults are often found on flowers, where females 

lay their eggs. 

Rivnay (L929) revised the North American 

species o.l this family. However, the species of 

Rhipiphoru.s, the largest North American genus of 

the family, are poorly known and cannot be reliably 

determined. The records cited below, based on specimens 

identifred with the aid of Rivnay's revision, 

are not all reliable. When revised, a number of additional 

species and provincial records will be 

recorded. 

eC (3);AB (2); SK (2); ON (5);PQ (2) 

Genus PELECOTOMA Fischer von Waldheim 

P. flavipes Melsheimer 

Subfamily PELECOTOMINAE 

ON PQ 

Genus MACIIOSIAGON Hentz 

M. cruenturr (Germar) 

M. limbatun. (Fabricius) 

Genus RHIPIPHORUS Bosc d'Antic 

Dorthesia Say 

Myodes Latreille 

Myodites Latreille 

R. californicus (LeConte) 

R. columbiar:,us W.J. Brown 

R. fasciatus (Say) 

Subfamily RHIPIPHORINAE 

SK 

-o*- 

250

Colydiidae 

R. mutchieri Rivnay 

R. nevadicus (LeConte) 

R. walshii (LeConte) 

R. zeschii (LeConte) 

Family COLYDIIDAE cylindrical bark beetles 

Y. Bousquet 

Only 21 species of colydiids are known to occur 

in Canada. They are usually found under bark, in 

galleries of wood-boring larvae, and in fungi. 

Adults and larvae of many species are predators of 

wood-boring insects; others feed on rotten wood or 

fungr; larvae of some species are parasitic on various 

groups such as Scolytidae, Platypodidae, 

Cerambycidae, and Buprestidae. 

The North American species of colydiids are 

usually divided into two subfamilies: Colydiinae, 

which includes most of the species, and 

Bothriderinae. Stephan (1989) has recently revised 

the North American species of the family. 

NT (2);sC (10); AB (3); SK (1); MB (1); ON (10);PQ 

(5); NB (1); NF (1) ' I (1) 

Subfamily COLYDIINAE 

T?ibe Synchitini 

Genus SYNCHITA Hellwig 

S. fuliginosa Melsheimer 

nigripennis LeConte 

Genus ENDEITOMA Sharp 

E. granulata (Say) 

floridana Casey 

Genus EUCICONES Sharp 

E. marginalis (Melsheimer) 

latus Casey 

Genus BITOMA Herbst 

Synchytodes Ctotch 

B. crenata (Fabricius)t 

B. ornata (LeConte) 

B. quadriguttata (Say) 

trinotata Casey 

zimmer manni Gu6rin-M6neville 

oN PQ 

PQ 

ON 

_y__:x;:__ __ _ 

Genus NAMUNARIA Reitter 

Coxelus Horn 

N. guttulatus (LeConte) 

N. pacificus (Horn) 

BC 

ON 

Genus LASCONOTUS LeConte 

L. borealis Horn 

L. complex LeConte 

L. intricatus Kraus 

h.rausiHatc}r 

L. planipennis Kraus 

L. tuberculatus Kraus 

L. vegrandis Horn 

apicalis Casey 

schwarzi lkans 

NT 

-BC 

NT BC 

-BC _BC 

-BC 

_SK 

AB 

AB 

MB ON 

PQ 

NB_ 

NF 

Genus AULONIIIM Erichson 

Key to North American species: Kingsolver & White (1967) 

Tlibe Colydiini 

A. longum LeOonte - BC 

25t

Zopheridae 

Genus COLYDIUM Fabricrus 

C. lineola Sry - BC - ON 

longiut,culum Say 

Genus PYC NOMERUS Erichson 

Endeclus LeConte 

Penthetispa Pascoe 

Tribe Pycnomerini 

P. reflexus tSay) - ON 

nitidus (LeConte) 

Genus DERETAPHRUS Newman 

Subfamily BOTHRIDERINAE 

Tlibe Deretaphrini 

D. oregonensis Horn - BC 

Genus OXYLAEMUS Erichson 

Redistes Casey 

Tribe Teredini 

O. californir us Crotch - BC 

Genus BOT :IRIDERES Erichson 

B. cryptus Stephan 

B. geminatus (Say) 

Tribe Bothriderini 

-oN 

-oN 

Family ZOPHERIDAE 

J.M. Campbell 

The f amily Zopheridae comprises about 125 Lawrence (1979). However, a revision of t,he North 

species,

Tenebrionidae 

Family TENEBRIONIDAE darkling beetles 

Y. Bousquet and J.M. Campbell 

The family Tenebrionidae comprises over 1200 

species in North America, but more than 8O7o of 

them are restricted to the southwestern United 

States. Only 135 species occur within the Canadian 

and Alaskan borders. Many tenebrionids live in 

semiarid areas and deserts, but in Canada most 

species are found in rotten wood, in fungr, under logs 

and stones, and under the bark of dead trees. A 

number of species, such as those of Tenebrio, 

Gnatocerus, Tribolium, and Polorus, are storedproduct 

pests, some of which are the most important 

in the country. Adults and larvae feed on plants or 

plant material, including decaying vegetation, seeds, 

roots, cereals and their products, and fungr. 

The concept of the family used for the checklist 

is similar to that of the tenebrionoid branch of 

Doyen and Tschinkel (1982). Adults of all species 

have paired defensive glands, with reservoirs opening 

between the seventh and eighth sternites. The 

lagrioid lineage, which is probably the sister group 

of the tenebrionids, has been given family status; 

these species lack defensive glands. Although the 

limits of the family are now fairly stable, its higher 

classifrcation is still poorly understood. Numerous 

changes have been proposed recently in the classification 

of the family (see Doyen and Tschinkel 1982), 

and more are expected; the alleculids, for example, 

are probably polyphyletic in origin. Rolf L. Aalbu 

(personal communication) has provided the classifrcation 

used here. There is no recent revision of the 

Nearctic Tenebrionidae, and most genera that occur 

in Canada have not been revised. Identifrcation is 

often difficult, especially in large groups like 

Eleodes. 

AK (3); YK (3); NT (8); BC (69); AB (32); SK (22); 

MB (32); oN (70); PQ (63); NB (16);NS (1a); PE (5); 

LB (2); NF (7) . H (1);I (11) 

Subfamily PHRENAPATINAE 

Tribe Penetini 

Genus CLAMORIS des Gozis 

Phthora Mulsant 

C. americana (Horn) 

BC 

Subfamily TENTYRIINAE 

Tlibe Eurymetopini 

Genus STERIPHANUS Casey 

S. subopacus (Horn) 


Tribe Anepsiini 

Genus ANEPSIUS LeConte 

A. montanus Casey 

AB 

Tribe Coniontini 

Genus CONIONTIS Eschscholtz 

Coelotaxis Hotn 

Coniontellus Casey 

Coniontides Casey 

C. ovalis LeConte 

acerba Casey 


anita Casey 

arida Casey 

breuiuscula Casey 

coruina Casey 

inepta Casey 

oblita Casey 

okanagani Boddy 

parilis Casey 

BC 

253

Tenebrionldae 

punchtta Casey 

regulc,ris Casey 

sculptipennis Casey 

uteanu Casey 

uanco.zueri Casey 

weidtt Casey 

Genus COItrLUS Eschscholtz 

Coelotnorpha Casey 

Pseudocoelus Casey 

Keyto species: Doyen (1976) 

C. ciliatus Eschscholtz 

ampli,:ollis Casey 

arenarius Casey 

curtulus Casey 

debilis Casey 

latus ()asey 

longuLus Casey 

obscurus Casey 

scolopax Casey 

sternalis Casey 

Tribe Coelini 

_BC 

Genus ASIDOPSIS Casey 

Tlibe Asidini 

A. opaca tSayt AB 

A. polita (Sayt AB SK 

Genus MICjROSCHATIA Solier 

Acrosthatia WlTke 

Pycno;'totida Casey 

P. inaequa.is tl,eConte) 

diuersa (Casey) 

- , 

Genus GL\?IASIDA Casey 

G. sordida Leconre) AB 

Genus PEL,ECYPHORUS Solier 

P. densicollis Horn - BC 

corrosus Casey 

Genus PHALEROMELA Reitter 

Nelite:t LeConte 

P. globosa (LeConte) 

P. picta (Mrnnerheim) 

P. variegat r Triplehorn 

picta (Horn) 

AK 

Subfamily DIAPERINAE 

Tribe Phaleriini 

_BC 

_BCAB 

Genus SCAPHIDEMA Redtenbacher 

Key to North American species: Triplehorn (1965) 

TYibe Nitionini 

S. aeneolum (LeConte) - YK - BC AB - MB ON PQ NB LB 

Genus RHIPIDANDRUS LeConte 

Tlibe Rhipidandrini 

R. flabellicornis (Sturm) - ON PQ 

paradorus (Palisot de Beauvois) 

254

Tenebrionidae 

Tlibe Bolitophagini 

Genus BOLITOTHERUS Candbze 

Phellidius LeConte 

B. cornutus (Panzer) 

bifurcus (Fabricius) 

Genus ELEATES Casey 

E. depressus (Randall) 

tetraopes (Newman) 

E. explanatus Casey 

Genus BOLITOPHAGUS Illiger 

B. corticola Say 

_BC 

_SKMBONPQNB 

MB ON PQ NB 

_ONPQ 

Genus ADELINA Dejean 

Doliema Pascoe 

Schedarosus Reitter 

Tribe Diaperini 

Key to North American genera: Doyen (1984) 

D. plana rFabricius) - ON 

depressa (Erichson) 

lecontei (I].orn) 

scidaria (Reitter) 

Genus CYNAEUS LeConte 

C. angustus tleConte) - BC AB SK MB ON PQ 

opoczs Champion 

Genus GNATOCERUS Thunberg 

CerandriaLrcas 

Echocerus Hotn 

Sicinus Champion 

(Subgenus GNATOCERUS s. str.) 

G. cornutus (Fabricius)f - BC MB ON PQ - NS 

maxillaris (PaIisot de Beauvois) 

Genus ALPHITOPHAGUS Stephens 

Phyletes Redtenbacher 

P hy lethus Redtenbacher 


A. bifasciatus {Sayr - SK MB ON PQ 

Genus DIAPERIS O.F. Mriller 


D. maculata Olivier MB ON PQ NB 

floridana Blatchley 

hydactina Fabricius 

hydni Fabricitts 

suturalis Chevrolat 

Genus NEOMIDA Latreille 

Arrhenoplita IGrby 

Euoplus LeConte 

Hoplocephala Castelnau & Brull6 


N. bicornis (Fabricius) 

cornigera (Fabricius) 

- ON PQ - NS 

g r acilis (Motschulsky) 

uirescens (Castelnau & Brull6) 

u ir idipennis (Fabricius) 

metallica (PaIisot de Beauvois) 

Genus PLATYDEMA Castelnau & Brull6 

Histeropsis Chevrolat 


255

Tenebrionidae 

P. americanum Castelnau & Brull6 - BC AB - MB ON PQ NB 

P. excavatLm {Say) - t - ON PQ 

fraten;.utn Chevrolat 

nigrih".m Motschulsky 

paruulum Casey 

tubercllatum Castelnau & Brull6 

P. neglectun Triplehorn - BC 

P. oregonense LeConte - BC 

P. ruficorne (Sturm) - ON PQ 

analeHald.eman 

opaculum Casey 

rufum (Melsheimer) 

rufiuen.tre Castelnau & Bru116 

P. subcosta:um Castelnau & Brulle - ON PQ 

clypea,um Haldeman 

o b lo ng ulum Motschulsky 

P. teleops 'l riplehorn - ON PQ 

Genus PENTAPHYLLUS Dejean 


P. pallidus LeConte - ON PQ 

Genus CORTICEUS Piller & Mitterpacher 

Hy pop,llaeus Fabricius 


Subfamilv HIPOPHLOEINAE 

Trib'e Hypophloeini 

C. parallelr- s tMelsheimert - ON PQ 

C. praetern.issus (Fa11) AK YK NT BC AB SK MB ON PQ 

C. subopacr.s (Wallis) NT BC AB - MB ON - NF 

C. substrialus (LeConteJ - BC 

C. tenuis (t.eOonte) - BC AB ON PQ NB 

minor'Wallis) 

o c c ider: talis (Wallis ) 

C. thoracicr.s tMelsheimerJ - ON 

piliger (LeConte) 

Genus ELEODES Eschscholtz 

(Subgenus Iv{ELANELEODES Blaisdell) 

Subfamilv OPATRINAE 

Tribe Eleodini 

E. humeral.s LeConte - BC 

latiusczlus Walker 

E. obsoletus obsoletus (Say) NT - AB SK MB ON 

E. tricostatus (Say) NT - AB SK MB ? - 

alternatus Kitby 

planetus Solier: 

robustus LeOonte 

(Subgenus LITHELEODES Blaisdell) 

E. extricatus convexicollis AB SK 

Blaisdell 

E. extricatus extricatus (Say) NT BC AB SK 

E. parvulus Blaisdell - BC 

E. vandykei modificatus Blaisdell - BC 

(Subgenus PROMUS LeConte) 

E. opacus (Slayt NT - AB SK MB 

(Subgenus IILEODES s. str.) 

E. hispilabris connexus LeConte 

binotatus Walker 

- BC 

imitabilis Blaisdell 

immundus Blaisdell 

loeuis Illaisdell 

256

Tenebrionidae 

E. hispilabris hispilabris (Say) 

lecontei Gemminger 

- BC AB SK MB 

sulcatus LeConte 

E. obscurus sulcipennis Mannerheim 

aratus LeConte 

- BC 

conjunctus Walker 

conuexicollis Walker 

(Subgenus BLAPYLIS Horn) 

(Am p hido ra E schscholtz ) 

(Eleodopsis Blaisdell) 

E. indentatus Blaisdell - BC 

E. novovermculus Boddy - BC AB 

E. nunenmacheri nunenmacheri - BC 

Blaisdell 

E. pimelioides Mannerheim - BC 

oregonus Blaisdell 

patruelis Blaisdell 

subligatus LeConte 

uerruculus Blaisdell 

uiator LeConte 

E. productus Mannerheim - BC 

uariolosus Blaisdell 

E. rolundipennis LeConte - BC 

strictus LeConte 

(Subgenus METABLAPYLIS Blaisdell) 

E. nigrinus LeConte - BC 

(Subgenus CRATIDUS LeConte) 

E. osculans (LeConte) - BC 

Genus EMBAPHION Say 

E. muricatum Say AB 

concduum LeConte 

Genus EPHALUS LeConte 

E. latimanus (LeConte) 

Tlibe Opatrini 

NS 

Genus BLAPSTINUS Dejean 

Aspidius Mulsant & Rey 

Heteropus Castelnau 

Lodinus Mulsant & Rey 

B. discolor Horn 


B. intermixtus Casey 

B. metallicus (Fabricius) 

interruptus (Say) 

luridus Mulsant & Rev 

B. pratensis LeConte 

B. pulverulentus Mannerheim - BC 

c alifor nicu s Motschulsky 

B. substriatus Charnpion 

_BC 

_BC 

-BCABSKMBONPQ_NS 

-oN 


Genus GONOCEPHAIUM Chevrolat 

G. bilineatum Walkert 

latifrons (LeConte) 

_BC 

Genus ULOMA Latreille 

U. impressa Melsheimer 

U. longula LeOonte 

Tribe Lllomini 

-BC 

-oN 

257

Tenebrionidae 

T?ibe Helopini 

Genus TAIiPELA Bates 

Larnperos AIIard 

T. micans (Fabricius) - ON PQ 

Genus HELOPS Fabricius 

H. californicus Mannerheim - BC 

H. convexulus LeConte - BC 


H. perniter.rs LeConte - BC 

H. punctipennis LeConte - BC 

punctatus Gemminger 

H. regulus Blaisdell - BC AB 

Genus TRIBOLIUM Macleay 

Margu,s Dejean 

Key to species: Hinton (1948) 

(Subgenus TRIBOLIUM s. str.) 

Subfamilv TENEBRIONINAE 

rriue Tlibotiini 

T. audax Halstead - BC AB SK MB ON PQ 

T. castaneum (Herbst)* - BC AB SK MB ON PQ PE 

ferrugineum (Dejean) 

T. confusu:n Jacquelin du Val* - BC - SK MB ON PQ NF 

T. destruct:r Uyttenboogaart* - BC AB SK MB ON PQ - NS PE - NF 

T. madens Charpentiertt - NB NS 

(Subgenus APIIANOTUS LeConte) 

T. brevicor;re (LeConte.t - BC 

Genus BIL'S Mulsant 

B. estriatu,; (LeContet - BC AB - PQ 

Genus PAI,ORUS Mulsant 

Key to species: Halstead (1967) 

(Subgenus PALORUS s. str.) 

P. ratzeburgii (Wissman)t - BC - ON PQ - NS 

(Subgenus CIRCOMUS Fleischer) 

P. subdeprr:ssus (Wollastont* MB 

Genus LAT'HETICUS Waterhouse 

L. oryzae Waterhouse* MB - PQ NB 

Genus ALF'HITOBIUS Stephens 

Tribe Tenebrionini 

A. diaperinus (Panzer)f - BC AB SK MB ON PQ NB NS PE - NF 

A. laevigat.Ls Fabriciusi - BC - ON 

plcezs (Olivier) 

picipes (Panzer) 

Genus MEI|ACLISA Jacquelin du Val 

M. marginalis Horn - BC 

Genus CENTRONOPUS Solier 

M ene chide s Motschul sky 

Pyres Champion 

Scotobates Fiye 

Key to New World species: Spilman (1962) 

C. calcarat:s tFabricius) 

aeneus (DeGeer) 

- ON PQ - NS 

carolirt ensis (Palisot de Beauvois) 

reflexus (Say) 

258

Tenebrionidae 

Genus TENEBRIO Linn6 

T. molitor Linn6t 

laticollis Stephens 

T. obscurus Fabriciust 

poliens Casey 

BC 

BC 

AB SK 

AB 

MB ON PQ NB 

-oNPQ 

NS 

PE 

NF 

NF 

Genus NEATUS LeConte 

N. tenebrioides (Palisot de 

Beauvois) 

badius (Say) 

picipes auct. amer. 

BC 

MB ON PQ 

Genus IDIOBATES Casey 

D. castaneus (Knoch) 

interstitialis (Say) 

ON 

PQ 

Genus MERACANTHAKiTbv 

Falacer Castelnau 

M. contracta (Palisot de Beauvois) 

canadensis Kirby 

cuprea (Castelnatt) 

inflata (Solier) 

(?) rugosa (Castelnau) 

(?) turnida (Melsheimer) 

Tlibe Amarygmini 

ON 

Genus HYMENORUS Mulsant 

Hymenophorus Mulsant 

Key to North American species: Fall (1931) 

H. caurinus Fall 

H. discretus Casey 

H. megops Hatch 

H. molestus Fall 

H. niger (Melsheimer) 

H. obesus Casey 

H. picipennis Casey 

H. pilosus (Melsheimer) 

H. sinuatus Fall 

Subfamily ALLE CLILINAE 

Tribe Alleculini 

BC 

BC 

BC 

-oN- 

_ONPQ 

-oNPQ _ONPQ 

-oNPQ 

- :*PQ 

NB 

NB 

Genus ISOMIRA Mulsant 

Tedinus Casey 

Key to eastern North American species: Marshall (1970) 

I. comstocki Papp 

I. iowensis Casey 

I. oblongula Casey 

I. pulla (Melsheimer) 

ignora Blatc}:.ley 

I. quadristriata (Couper) 

ueluntina LeConte 

I. sericea (Say) 

tenebrosa Casey 

I. valida Schwarz 

similis Blatchley 

Genus PSEUDOCISTELA Crotch 

Chromatia LeConte 

Cisfelo Fabricius 

Telacis Poey 

P. amoena (Say) 

P. brevis (Say) 

erythroptera (Ziegler) 

Tiibe Gonoderinr 

BC 

- ow: 

_ONPQ 

-oNPQ 

MB ON PQ 

-oNPQ 

-oN 

-ONPQ 

-ONPQ 

NB NS 

_NS 

259

Tenebrionilae 

P. pectinat.r Hopping - BC 

P. pinguis l.eConle - BC 

pacific:a Hopping 

Genus CAI'NOCHROA LeConte 

C. fuliginosa (Melsheimer) - ON PQ NB 

Genus ANI)ROCHIRUS LeConte 

A. erythropus (Kirbyt - ON PQ - NS 

fuscipes (Melsheimer) 

luteipes LeConte 

Tlibe Mycetocharini 

Genus MY(IETOCHARA Berthold 

M y ceto chare s Latreille 

M y cetocharis Gyllenhal 

M y cetct p h il a Gyllenhal 

Pterna Seidlitz 

Stigmotoma LeConte 

Key to North American species: Campbell (1978€) 

M. analis (LeConte) 

dauisi Hatch 

horni ))rry 

lugubris (LeConte) 

M. bicolor (Couper) 

M. binotata (Say) 

longula (LeConte) 

margiltata (LeConte) 

M. foveata .LeConte) 

gilipes Casey 

gracilis (LeConte) 

lecontti Borchmann 

tenuis (LeConle) 

M. fraterna (Say) 

laticol is (LeConte) 

megahlps Casey 

nigerrtma Casey 

M. procera Casey 

angusta Hatch 

caseyi Hatch 

crassu]ipes Casey 

downe'.Hatch 

malkh;iHatch 

neuadensis Casey 

Genus HYI\{ENOCHARA Campbell 

Key to species: Campbell (I978d) 

H. rufipes (LeConte) 

-BC-SKMBONPQ 

_BC 

-ONPQ 

-oNPQ 

-ONPQ 

MB ON PQ 

-oNPQ 

Subfamilv COELOMETOPINAE 

Tri6e Coelometopini 

Genus MEIIINUS LeConte 

M. laevis tCtliviert - ON PQ 

Genus XYLOPINUS LeConte 

T aenoltate s Motschulsky 

X. aenescens LeConte PQ 

X. saperdioides tOlivier) - ON PQ 

anthracinus (Knoch) 

calcaratus (Palisot de Beauvois) 

ruftpes (Say) 

spinipes (Fabricius) 

Genus HAILANDRUS LeConte 

H. concolor LeConte MB ON PQ 

260

H. fulvipes (Herbst) 

femoratus (Fabricius) 

Genus COELOCNEMIS Mannerheim 

Key to species: Doyen (1973) 

C. californica Mannerheim 

barretti BlaisdelI 

bosolis Casey 

columbiana Casey 

dilat icoll is Mannerherm 

idahoensis Casey 

ouipennis Casey 

rauca Casey 

rugosa Linne\l 


utensis Casey 

Genus IPHTHIMINUS Spilman 

I. opacus (LeConte) 

I. salebrosus (Casey) 

L serratus (Mannerheim) 

serudtor (Walker) 

seruilis (Walker) 

subligatus (Walker) 

_BC 

-BC _BC 

AB 

_SX 

AB 

Lagriidae 

oN PQ 

yu oN PQ NB NS 

Genus AI-OBATES Motschulsky 

A. pensylvanicus (DeGeer) 

chrysops (Herbst) 

inermis (Mannerheim) 

sublaeuis (Palisot de Beauvois) 

MB ON 

PQ 

Genus UPIS Fabricius 

U. ceramboides (Linn6)x 

reticulata (Say) 

uariolosa (DeGeer) 

AK 

YK 

NT 

BC 

AB 

SK MB ON PQ NB PE 

LB 

NF 

Tribe Strongyliini 

Genus STRONGYLIUM Kirbv 

Reminius Casey 

Stenochia I{rtby 


S. tenuicolle (Say) 

Triplehorn and Spilman (1973) 

ON 

PQ 

Family LAGRIIDAE lagriid beetles 

J.M. Campbell 

The family Lagriidae is primarily tropical in distribution 

and comprises over 1000 species around the 

world. Only six species have been recorded in Canada. 

Adults of Statird and, Arthromacra are usually collected 

by beating the foliage of trees; larvae are 

thought to breed in decaying wood or leaf litter. Little 

is known about the biolory of the species. 

The status of this family is uncertain. In recent 

studies of the higher classification of the 

Tenebrionidae, Watt (t974), Doyen and Lawrence 

(1979), and Doyen and Tschinkel (1982) placed the 

lagriids in Tenebrionidae, usually as a subfamily. 

However, Doyen and Tschinkel (1982) suggested that 

the lagriids and related groups form the sister group 

to all other tenebrionids and possibly should be 

restored to family status. I prefer to recognize the 

lagriids as a distinct family until their relationship 

with respect to other tenebrionoid groups has been 

better established. The classification of the North 

American lagriids was reviewed by Parsons (19750), 

who includedAnaedus andParatenetus in the subfamily 

Lupropinae (tribe Heterotarsini of Tenebrionidae, 

in most publications); this subfamily more properly 

should be named Goniaderinae. Doyen and Tschinkel 

(1982) have suggested that the genus Paratenetus be 

placed in or near the tenebrionid tribe Belopini. Until 

the classification of the Lagriidae is reviewed, I have 

chosen to follow the arrangement of Parsons (1975o). 

ttlT (1);sC (1);AB (1);MB (3); oN (5); PQ (5);NB (1); 

NS (1) 

26t

Cephaloidae 

Subfamily GONIADERINAE 

Genus PAR.A.TENETUS Spinola 

Storthe,phora Mdklin 

P. crinitus lrall 

P. fuscus LeConte 

P. punctatus Spinola 

Genus ANAEDUS Blanchard 

A. brunneur; (.Zieglet) 

- - NrncAB- MB3N|8- - - - - 

-oNPQ 

ON 

Subfamily LAGRIINAE 

Genus ART.FIROMACRA Kirby 

Key to N,.'rth American species: 

A. aenea (Say) 

donacioides I\trby 

Parsons (1975a) 

MB ON PQ 

NB NS 

Genus STA IIRA Audinet-Serville 

Key to Ntrrth American species: Parsons (I975a) 

S. gagatina Melsheimer 

respler,.dens Melsheimer 

Subfamily STATIRINAE 

PQ 

Family CEPIIALOIDAE false long-hornedbeetles 

J.M. Campbell 

Cephaloidae is a small family of about 20 

species rerstricted to the cooler parts of the Holarctic 

region. l)nly nine species are known to occur in 

Canada ernd Alaska. Larvae are found in old, rotting 

logs; adults of Cephaloon are often taken on 

flowers. 

Species of this family are reasonably well 

known taxonomically. Hopping and Hopping (1934) 

and Arnett (1953) revised and reviewed, respectively, 

the species of the family, but included only 

the genus Cephaloon. Subsequently, Arnett (1968) 

transferred the subfamilies Stenotrachelinae and 

Nematoplinae from the melandryiids and pediliids, 

respectively, to Cephaloidae. Both subfarrLilies have 

been considered by some authors to be disbinct families 

(Mamayev 1973), but I have followed Lawrence 

(1982a) in accepting Arnett's treatment. 

AK (1); YK (1); NT (2); BC (5); AB (3); MB (1); ON 

(a); PQ (5);NB (1); PE (1);LB (2);NF (2) 

Subfamily CEPHALOINAE 

Genus CEPHALOON Newman 

Drachj,lis Casey 

Ichnodes Dejean 

Sponia',ium Casey 

Typitium Casey 

Key to species: Arnett (1953) 

C. bicolor Horn 

simulans (Casey) 

C. lepturoid.es Newman 

uarians Haldeman 

C. pacifrcum Van Dyke 

C. tenuicome LeConte 

piceuri }Iorn 

ornatum Casey 

uersicolor Casey 

C. ungulare LeConte 

BC 

-BC 

_BCAB 

oN PQ 

ON PQ PE LB 

NF 

262

Meloidae 

Subfamily STENOTRA.CHELINAE 

Genus STENOTRACHELUS Latreille 

S. aeneus (Fabricius) 


MB ON PQ 

LB 

NF 

Genus ANELPISTUS Horn 

A. americanus Horn 

A. canadensis Mank 

NT BC 

AB 

PQ 

Subfamily NEMATOPLINAE 

Genus NEMATOPLUS LeConte 

N. collaris LeConte 

ON 

PQ NB 

Family MELOIDAE blister beetles 

J.M. Campbell 

Meloidae comprises over 3000 species, which 

are found in most regions of the world. However, 

the majority are restricted to warmer, drier areas. 

Only 49 species and subspecies are known to occur 

in Canada and Alaska, and most of these are found 

in the semiarid areas of the Prairie Provinces and 

British Columbia. Larvae of Epicautini feed on 

grasshopper eggs, and those of aII other Canadian 

blister beetles feed on eggs, larvae, and food 

reserves of ground-nesting bees. Larvae of Meloini 

and Zonitinae are phoretic. The eggs are laid on 

flowers, the larvae hatch, and the frrst-instar larvae 

(called triungulin) attach themselves to bees that 

visit flowers and are transported to the bee's nest, 

where they drop off. Adults of Canadian meloids are 

phytophagous or feed on nectar; many are considered 

economic pests. Adults of this family produce 

cantharidin (Spanish fly), which causes blisters on 

contact with the skin and, if present in dried hay, 

has been known to be fatal to horses. Cantharidin 

is also used for medical purposes. 

Most genera of North American Meloidae are 

well known taxonomically. A few genera such as 

Tricrania have not been revised, and the large, 

important genus Epicauta is in need of additional 

study. The Canadian fauna is well known, and 

therefore few additional species are expected to be 

added to the checklist in the future. Van Dyke 

(1928a) and Arnett (1968) provide the most recent 

treatments of the family at the generic level. 

aK (3); vK (1); NT (3); sC (25); AB (26); SK (23); 

MB (20); ON (17);PQ (12);NB (3);NS (3) 

Subfamily MELOINAE 

Tribe Meloini 

Genus EURYMELOE Reitter 

E. aleuticus (Borchmann) 

AX 

Genus MELOE Linn6 


Pinto and Selander (1970) 

(Subgenus TREIODOUS Dugbs) 

M. barbarus LeConte 

BC 

(Subgenus MELOE s. str.) 

Melittophagus Kirby 

M. americanus Leach 

tnoerens LeConte 

temnus Dillon 

M. angusticollis Say 


rugipennis LeConte 

M. carbonaceus LeConte 

M. dianellus Pinto & Selander 

M. impressus Kirby 

occidentalis Van Dyke 

BC 

AB 

SK 

ON 

MB ON PQ 

---ncffisx-oNPe 


-NS 

NB 

NS 

263

Meloidae 

M. niger Kirby 

hirbyi Drllon 

opacus LeConte 

perplee:us LeConte 

M. occultus Pinto & Selander 

M. strigulos,us Mannerheim 

_BC 

AK 

BC 

M. vandykei Pinto & Selander - BC 

Genus LYTTA Fabricius 

Key to species: Selander (1960) 

(Subgenus PARALYTTA Selander) 

L. vulnerata tl,eContet 

anguicollis (Dugbs) 

- BC 

cooperi LeConte 

(Subgenus I'OREOSPASTA Horn) 

NTBCABSKMBONPQ_NS 

Tlibe Lyttini 

Subtribe Lyttina 

L. cyanipen:ris (Leconte) - BC AB 

solicis LeConte 

L. nuttalli Say AB SK MB 

fulgifera (LeConte) 

L. stygica (LeConte) 

arbore

Meloidae 

E. puncticollis Mannerheim - BC AB 

barberi Werner 

oblita LeConte 

E. sericans Leconte AB sK 

E. ventralis werner AB sK 

E. vittata (Fabriciust - ON PQ 

chapmani (Woodhouse) 

I e rnni scata (Fabricius) 

(Subgenus MACROBASIS LeConte) 

(Ap teros pasta LeConte) 

E. fabricii (LeContet AB SK MB oN PQ 


E. Ilavocinerea (Blatchleyr MB ON 

E. murina (LeConte) - ON PQ NB 

unicolor Kirby 

E. subglabra rFall I - BC AB SK MB 

Genus LINSLEYA MacSwain 

Key to species: Selander (1955) 

(Subgenus LINSLEYINA Selander) 

L. sphaericollis {Say) 

chalybea (LeConte) 

- BC AB SK MB 

c haly b e ata ( Gemrninger) 

infidelis (Fall) 

Genus NEMOGNATIIA Iiliger 

Key to North American species: Enns (1956) 

(Subgenus MEGANEMOGNATHA Enns) 

Subfamily ZONITINAE 

T?ibe Zonitini 

Subtribe Nemognathina 

N. lutea dichroa LeConte - BC AB SK MB 

N. lutea lutea LeConte AB 

N. soror l\lacSwain - BC 

(Subgenus PAURONEMOGNATIIA Enns) 

N. nemorensis Henlz - ON 

b irnac ulata Melsheimer 

(Subgenus NEMOGNATHA s. str.) 

N. piazata bicolor LeOonte AB SK MB 

discolor LeConte 

stellaris Beauregard 

texana LeConte 

Genus TRICRANIA LeConte 

Tricraniodes Wellman 

T. mur"rayi LeConte - BC 

T. sanguinipennis {Say) - ON PQ 

T. stansburl (Haldeman) NT BC 

Genus HORNIA Riley 

H. minutipennis Riley AB 

Genus ZONITIS Fabricius 

Key to North American species: Enns (1956) 

(Subgenus NEOZONITIS Enns) 

Subtribe Zonitina 

Z. bilineata Say - ON PQ 

lineata Mels}reilner 

m an d ib u laris Mel sheimer 

Z. vermiculata Schaeffer - BC 

(Subgenus PARAZONITIS Enns) 

265

Oedemeridae 

Z. sayi Wickham 

imrnactilata (Say) 

AB SK 

MB 

Genus GNAIHIUM Kirby 

Key to species: MacSwain (1952) 

G. minimurrL (Say) 

AB 

SK 

Family OEDEMERIDAE false blister beetles 

J.M. Campbell 

The family Oedemeridae comprises at least 100 

genera arrd 1500 species, distributed in all faunal 

regions. Thirteen species are known to occur in 

Canada and Alaska. The biology of most species is 

unknown. Larvae of some species live in rotten 

wood, and adults of most species are found on flowers, 

where they feed on pollen and nectar. One 

species, Nacerdes melanura (wharf borer), is an economic 

pest of pilings and ship timber. A few exotic 

species have been reported to cause blister:ing similar 

to that produced by some blister beetles,. 

Arnett (1951) revised the North American 

species of oedemerids; little information has since 

been added to our knowledge of the Clanadian 

fauna. 

AK(2); BC (10);AB (4); MB (3); ON (5); PQ (5);NB 

(1);NS (2);NF (1) ' I (1) 

Genus CAJ-OPUS Fabricius 

C. ansustus LeConte 

Subfamily CALOPODINAE 

BC 

AB 

oN PQ 

Genus XANTHOCHROA Schmidt 

X. testacea llorn 

Subfamily NACERDINAE 

BC 

Genus NACIIRDES Dejean 

Anogcotles Dejean 

N. melanura (Linn6)t 

apicalis (Say) 

BC 

MB ON PQ 

NB NS 

NF 

Subfamily OEDEMERINAE 

Tlibe Ditylini 

Genus DITYLUS Fischer von Waldheim 

D. caeruleus (Randall) 

D. gracilis L,:Conte 

uestitus LeConte 

D. quadricol)is LeConte 

consors LeConte 

AK 

AK 

BC 

BC 

AB 

_MB 

ONPQ_--__ 

Tribe Asclerini 

Genus XANIIHROCHROINA Ganglbauer 

X. bicolor (LerConte) 

BC 

Genus ASCL,ERA Stephens 

A. discolor L,eConte 

A. excavata]-eConte 

A. nigra LeConte 

A. puncticollis (Say) 

A. ruficollis (Say) 

carinatct (Newman) 

_BC 

-BCAB 

---:t--^ras 

oN pq 

ON PQ 

NS 

Genus COPIDITA LeConte 

C. quadrimaculata LeConte 

BC 

266

Mycteridae 

Family PROSTOMIDAE prostomid beetles 

J.M. Campbell 

Prostomidae is a small family that contains only 

two genera and about 20 species. Most faunal 

regions, except South America, have a few species of 

the family; only one species occurs in Canada. 

Nothing is known about the habitat of this species. 

In most North American literature Prostomis 

arnericanus has been referred to as P. mandibularis 

(Fabricius), a European species. 

BC (1) 

Genus PROSTOMIS Latreille 

P. americanus Crotch 

BC 

FamilySYNCHROIDAE synchroidbeetles 

L. LeSage 

As currently defined (Crowson 1966, Lawrence 

1982a), the family Synchroidae is reduced to two 

genera and 10 species, which were previously 

placed in the Melandryidae. Only one species of the 

genus Synchroa occurs in Canada. Both adults and 

Iarvae live under the bark ofold dead trees. 

There is no recent treatment of the North 

American species of this family. 

uB (1);oN (1);PQ (rl 

Genus SYNCHROA Newman 

S. punctata Newman 

nzurina Haldeman 

umbrina Melsheimer 

MB ON PQ 

Family MYCTERIDAE 

mycterid beetles 

J.M. Campbell 

Mycteridae includes about 160 species around 

the world, of which four are known to occur in 

Canada. Larvae live under bark, and adults are 

often found on flowers. 

Species of this family have been included in a 

variety of families, including Melandryidae (Leng 

L920), Salpingidae (Arnett 1968), and Pythidae 

(Hatch 1965). I have followed Lawrence (1982o) in 

considering mycterids to be a distinct family. The 

species are rather poorly known taxonomically, and 

a modern revision is needed badly. 

sC (2); ON (2);PQ (2) 

Genus LACCONOTUS LeConte 

L. pinicola Horn 

L. punctatus LeConte 

Genus MYCTERUS Clairville 


M. concolor LeConte 

flauipennis Horn 

M. scaber Haldeman 

Hopping (1935) 

BC 

BC 

oN PQ 

ON PQ 

267

Pythidae 

Family BORIDAE 

J.M. Campbell 

Boridae contains three genera and six species; 

only two species occur in Canada. Larvae of Boros 

occur under the bark of coniferous trees and nrobably 

feed on decayed cambium (Lawrence L9824). 

The species of Boridae have been placed in a 

variety of families such as Tenebrionidae (Leng 

1920), P;'thidae (Hatch 1965), and Salpingidae 

(Arnett 1968). Young (19856) transferred Lecontia 

from the family Pythidae to the Boridae. I have followed 

Lawrence (1982a) in recognizing Boridae as a 

separate family. 

NT (1); sC (1); AB (2); SK (2); MB (2); O.N (2); PQ 

(2);NB (1) 

Genus BOR()S Herbst 

B. unicolor Siay 

Genus LECONTIA Champion 

Crymoctes LeConte 

L. discicollis (LeConte) 

SK MB ON PQ 


Family PYTHIDAE pythid beetles 

J.M. Campbell 

The family P1'thidae forms a small group comprising 

orrly about 50 species around the world, of 

which six occur in Canada and Alaska. Members of 

the subfermily Pythinae occur primarily in the 

Holarctic region, whereas members of the other two 

subfamilies are found in Central and South 

America, .New Zealand, and Australia. Both larvae 

and adults are found under bark ofconiferous trees. 

Larvae fe':d on decaying inner bark (Pytho) or sapwood 

(Prirgnathus). 

Meml,ers of this family have previously been 

assigned to other families such as Salpingidae 

(Arnett 1968) or, conversely, Pythidae has included 

a number of taxa now transferred to other families 

such as Salpingidae, Cononotidae, and Mycteridae. 

The arrangement followed here is based on 

Lawrence (1982a) with the exception of Lecontia, 

which has been transferred to Boridae as Young 

suggests (1985b). Pollock is currently revising the 

genus Pytho, so that most species of the family will 

soon be well known taxonomically. 

AK (3); YK (3); NT (3); BC (5); AB (4); SK (3); MB 

(a);ON (a);PQ (a);NB (3);NS (a); LB (1);NF (2) 

Subfamily PYTHINAE 

Genus PYTIIO Latreille 

P. americamrs Kirby 

deplanatus Mannerheim 

P. niger Kirtry 

P. seidlitzi Blair 

P. strictus L,-.Conte 

AK YK 

5v 

NT 

BC 

AB 

NT BC AB 

ItBCy 

SK 

SK 

MB ON PQ NB 

MB ON PQ 

MB ON PQ 

NB 

NS 

NS 

NS 

LB 

NF 

Genus PRIOGNATHUS LeConte 

P. monilicornis (Randall) 

sahl b e rg i (Mannerheim) 

Genus SPHI'LMA Horn 

S. quadricollis Horn 

AK YK BC SK MB ON PQ NB NS 

BC 

NF 

268

Pyrochroidae 

Family PYROCHROIDAE fire-colored beetles 

J.M. Campbell 

Twelve species of the family Pyrochroidae are 

known to occur in Canada and Alaska. Adult 

pyrochroids are rarely collected, although adults of 

Dendroides and Neopyrochroo occasionally are 

attracted to light, and those of most species can be 

collected in flight intercept traps or Malaise traps. 

Larvae of many species are common and are among 

the most frequently collected and distinctive species 

of Coleoptera found under bark and in rotting wood. 

Young (1975) summarized the feeding habits and 

biology of pyrochroids. Larvae live in dead wood 

and are apparently fungivorous rather than predaceous 

or xylophagous, as reported by earlier 

authors. Adults rarely feed, but may be attracted at 

night to fermenting fruit or bait. Adults of 

Neopyrochroo and Schizotus ceruicalis are attracted 

to cantharidin. 

The family is well known taxonomically, thanks 

to the revision and catalog of the North American 

species by Young (1975 and 1983, respectively). 

However, in recent papers, Young has suggested 

that Ischalia be transferred to the Anthicidae 

(1985o) and that the genus Pedilus be transferred 

from Pedilidae to Pyrochroidae (1984). I prefer to 

postpone making these changes until his more 

detailed analysis of Pedilus is published. 

AK (2); NT (1); BC (5); AB (a); SK (1); MB (3); ON 

(7);PQ (7); NB (3); NS (a);NF (1) 

Subfamily ISCIIALIINAE 

Genus ISCHALIA Pascoe 

(Subgenus EUPLEURIDA LeConte) 

L costata (LeConte) 

I. vancouverensis Harrington 

ecAB--9*PQ-TS--- 

Subfamily PYROCHROINAE 

Genus SCHIZOTUS Newman 

S. cervicalis Newman 

S. fuscicollis (Dejean) 

AK 

NT BC AB 

MB ON 

PQ NB NS 

Genus NEOPYROCHROA Blair 

N. femoralis (LeConte) 

N. flabellata (Fabricius) 

-oNPQ 

-ONPQ 

Genus DENDROIDES Latreille 

D. canadensis Latreille 

blcolor (Newman) 

cyanipennis Latreilie 

D. concolor (Newman) 

D. ephemeroides (Mannerheim) 

D. marginatus Van Dyke 

pacificus Barrett 

D. picipes Horn 

D. testaceus LeConte 

T 

BC 

BC 

BC 

AB 

AB 

SK 

MB ON PQ 

_ONPQ 

MB ON pe 

NB NS 

NB NS 

NF 

269

Salpingidae 

Family SALPINGIDAE narrow-waistedbarkbeetles 

J.M. Campbell 

Salpingidae comprises 10 species in Canada and 

Alaska and about 350 species around the world. 

The adults are most often found under the bark of 

conifers, on flowers, or in detritus. Their feeding 

habits are unknown. 

Salpingidae, as currently constituted (Lawrence 

Ig82a), includes groups previously treated as distinct 

families, such as the aegialitids, dacoderids, 

elacatidsr, eurystethids, inopeplids, othniids, and 

tretothoracids. On the other hand. some of the sal- 

pingid genera were previously included in families 

such as the Pythidae and Colydiidae (Agleruus). The 

family is poorly known taxonomically, and a comprehensive 

revision of most North American genera 

is needed. 

AK (3); YK (1); NT (1); BC (7); AB (1); SK (1); MB 

(1); oN (3); PQ (2); NB (1); NS (1); LB (1); NF (2) 

. I(1) 

Subfamily SALPINGINAE 

Genus SPHAERIESTES Stephens 

Salpingus atct. 

S. alternatus (LeConte) 

S. virescens (LeCon!e) 

BC 

YK NT BC 

AB SX nrS ON pe NS - 

is Nr 

Genus VINCENZELLUS Reitter 

V. elongatus (Mannerheim) AI( 

Genus RHINOSIMUS Latreille 

Key to Ncrth American species: Blair (1932) 

R. viridiaerLeus Randall AK 

a e n e ir nt r is Mannerheim 

nitens LeConte 

R. iecontei Blair 

R. pallipes Boheman 

-BC 

_BC 

BC 

oN PQ 

NS 

Subfamily AE GIALITINAE 

Key to species: Spilman (1967) 

Genus AEC|IALITES Dejean 

Eurystethus Seidlitz 

A. californi,:us Motschulsky 

AK 

BC 

Genus ELA.CATIS Pascoe 

Othnius LeOonte 

E. lugubris (Horn) 

Subfamily OTHNIINAE 

BC 

Subfamily AGLENINAE 

Genus AGLENUS Erichson 

A. brunneus (Gyllenhal)t 

obsoler)us (Shuckard) 

ON 

270

Anthicidae 

Family ANTHICIDAE antlike flower beetles 

Y. Bousquet 

Anthicidae comprises 55 species in Canada and 

Alaska. The adults are usually found under debris 

and in leaf litter on the ground or on flowers and 

foliage; the larvae live in the soil and in decayed 

vegetation. Apparently, members of many species 

are scavengers; those of some species are known to 

be egg predators. 

The North American species of this family are 

usually grouped into two subfamilies, Notoxinae 

and Anthicinae. Thanks to the work of Werner and 

of Chandler, the taxonomy of most of the groups 

that occur in North America has been worked out. 

AK (2); YK (2); NT (5); BC (29); AB (25); SK (24); 

MB (19); ON (25); PQ (21); NB (9); NS (5); PE (1); 

NF(3) .I(3) 

Genus MECYNOTARSUS LaFert6-S6nectbre 

Key to New World species: Chandler (1977) 

M. candidus LeConte 

Subfamily NOTOXINAE 

TYibe Notoxini 

AB 

Genus NOTOXUS Fabricius 

Ceratoderus Blanchard 

Monocerus Faldermann 

Key to North Arnerican species: Chandler (1982) 

N. anchora Hentz 

N. bifasciatus (LeConte) 

N. cavicornis LeConte 

breuiusculus Fall 

obesulus Blaisdell 

N. desertus Casey 

constrictus Casey 

N. intermedius Fall 

paruidens Fa\l 

N. manitoba Chandler 

N. marginatus LeConte 

N. montanus Casey 

microcerus Casey 

similis Fall 

N. pictus Casey 

N. robustus Casey 

humboldti Casey 

N. schwarzi Horn 

N. serratus (LeConte) 

digitalis LeConte 

uandykei Blaisdell 

N. subtilis LeConte 

_NT_ABSK 

BC 

-AB 

_AB 

AB 

BC AB 

BC 

BC 

BC AB 

AB 

SK 

ix 

MB ON PQ 

MB-_ Iu_ PE 

MB_ 

ON 

SK MB ON 

Genus AMBLYDERUS LaFert6-S6nectdre 

A. granularis LeConte 

A. obesus Casey 

A. pallens LeConte 

Subfamily ANTHICINAE 

TYibe Anthicini 

ON PQ 

BC 


NB 

Genus ANTHICUS Paykull 

Nathicus Casey 

Key to North American species: Werner (1964) 

A. ancil)a Casey 

caseyi Pic 

parallelus Casey 

A. bellulus LeConte 

BC 

AB 

SK 

277

Anthicidaer 

heluin.us Casey 

minusculus Champion 

A. biguttulus LeConte 

ouicol,'.is Casey 

protectus Casey 

A. cervinusi LaFert6-S6nectbre 

bifasc;iatus Say 

b izonc'.t u s LaFert6-S6nectbre 

gilenst.s Casey 

seminotatus Casey 

termir:alis LeConte 

A. coracinu.s LeConte AK 

A. cribratus LeConte 

hecate Casey 

A. ephippium LaFert6-S6nectbre 


diffici;.is LeConte 

luteolus LeConte 

pinguescens Casey 

simiolus Casey 

A. flavicans LeConte 

A. haldemr.ni LeConte 

decorellus Casey 

q uadr iguttatus Haldeman 

sozcias Casey 

A. hastatus Casey 

solidus Casey 

subcati.uus Casey 

A. heroicus Casey 

A. lecontei Champion 

auriger Casey 

junctus Casey 

stellat,^ts Casey 

uulneratus Casey 

A. lutuient.rs Casey 

agilis Casey 

herifu.,lus Casey 

mund.ts Casey 

A. melanch olicus LaFert6-S6nectbre 

facilis Casey 

latebntns LeConte 

spretus LeConte 

A. nanus L.:Conte 

amoenus Casey 

innocens Casey 

inscitt',s Casey 

nympltn Casey 


peninsularis Casey 

uagan:; Casey 

A. nigritus Mannerheim AK 

A. praeceps Casey 

A. punctule.tus LeConte 

cephalotes Casey 

decrepitus Casey 

mercurialis Casey 

monticola Casey 

A. scabriceps LeConte 

compositus Casey 

NT 

BC 

BC 

YK NT BC AB 

BC 

BC AB 

_NTBCAB 

-NT_AB 

--ec 

YK NT 

AB SK MB ON 

:" yu 9* 

SK MB ON 

SK MB ON 

SK_ON 

BC AB SK MB 

BC 

BC 

AB 

BC AB 

AB SK 

SK 

_ONPQ 

S^ l,fn - 

ON 

PQ 

PQ 

PQ 

PQ 

PQ 

ON PQ 

PQ NB 

NB NS 

NB NS 


NE 

NF 

Genus SAPINTUS Casey 

Key to Ncrth American species: Werner (1962) 

S. fulvipes l.LaFert6-56nectdre) 

colonus Casey 

rustict:s Casey 

S. lemniscatus Werner 

S. pubescer.Ls (LaFert6-S6nectdre) 

S. pusillus (LaFert6-S6nectbre) 

binomi.natus Pic 

festinans Casey 

mollis Casey 

BC 

_MBONPQ 

ON 

oN _MBONPQ_NS 

272

Anthicidae 

Genus STRICTICOMUS Pic 

S. tobias (Marseul)i PQ 

Genus OMONADUS Mulsant & Rey 

Hemantus Casey 


O. floralis (Linn6)t - BC AB SK MB ON PQ NB - NF 

(?) basillaris (Say) 

O. formicarius (Goeze)f - BC AB SK MB ON PQ NB 

enodis (Casey) 

rixator (Casey 

scenicus (Casey) 

Genus MALPORUS Casey 

M. cinctus tsay) - SK - ON 

M. formicarius (LaFert6-S6nectbre) - SK MB ON PQ NB NS 

Genus THICANUS Casey 

T. mimus Casey - BC AB SK MB 

Genus EIJVACUSUS Casey 

E. coloradanus Casey - BC AB SK MB 

Genus VACUSUS Casey 

Key to species: Werner (1961) 

V. formicetorum (Wasmann) AB SK 

desertorum Casey 


suspectus Casey 

V. nigritulus (LeConte) 

arcanus Casey 

- BC 

Genus ISCHYROPAIPUS LaFert6-S6nectbre 

Lappus Casey 


L cochisei Werner BC 

L nitidulus (LeConte) - BC 

alacer (Casey) 

- ON PQ 

asperulus (Casey) 

cursor (Casey) 

nubilatus (Casey 

soliuagans (Casey) 

uigilans (Casey) 

I. obscurus (LaFert6-s6nectdre) ON PQ 

I. sturmi (LaFert6-S6nectbre) - ON 

e le g ans (LaFert6-S6nectbre) 

I. turgidicollis (Casey) 

animatus (Casey) 

- BC 

Genus TANARTHRUS LeOonte 

Key to species: Chandler (1975) 

(Subgenus TANARTHROPSIS Casey) 

T. salicola LeConte - SK 

densus Casey 

mormon Wickham 

nubifer Casey 

tricolor Casey 

uirginalis Wickham 

273

Pedilidae 

Family PEDILIDAE pedilid beetles 

Y. Bousquet 

Pedilidae currently includes 19 species in 

Canada; none has been reported from Alaska. The 

adults usually occur on flowers or foliage and feed 

on flowers, pollen, and nectar; they are attracted to 

light at night. The larvae live in decaying plant 

matter, uLsually in wet places. 

The group is probably closely related to 

Anthicidae, and some authors, including Crowson 

(1955) and Lawrence (1982o), have included the 

pedilids in Anthicidae. Members of Pedilidae are 

usually classified in two subfamilies, Pedilinae and 

Eurygeniinae. Young (1984) suggestecl that the 

genus Pedilus be transferred from Pedilidae to 

Pyrochroidae and placed in its own subfamily 

(Pedilinae) and that other North American pedilids 

be tranferred to Anthicidae. I prefer to wait until 

Young's revision of the genus Pedilus is published 

before incorporating these changes. 

BC (8); AB (4); SK (1); MB (2); ON (6); PQ (e); NB 

(1); NS (1) 

Genus PEDILUS Fischer von Waldheim 

Corphyra Say 

P. abnormis (Horn) 

P. canaliculatus (LeConte) 

P. cyanipen.nis Bland 

P. elegans (Hentz) 

fuluipe's Newman 

h a e m a t ho i dal i s Zi e gler 

lugubris Newman 

newmani LeConte 

ruficollis Ziegler 

P. flabellat:rs (Horn) 

P. impressrLs (Say) 

P. inconspicuus (Horn) 

P. longilobrs Fall 

P. lugubris (Say) 

imzs llewman 

inornatus (Randall) 

infumtr.tus (Hentz) 

nigricctns Ziegler 

rufithorax Newman 

P. oregonus Fall 

P. punctulatus LeConte 

funebris (Hotn) 

P. terminalis (Say) 

guttulu Newman 

Subfamily PEDILINAE 

_?AB-_ 

BC 

BC 

_ _ _ f _ _ 

_BC 

_BC 

MB 

ON 

ON 

ON 

pe----- 

PQ 

PQ 

PQ 

ON pe NB NS 

PQ 

Genus PEBGETUS Casey 

P. campanulatus (LeOonte) 

Genus STEREOPAJ-PUS LaFert6-S6nectdre 

Key to species: Abdullah (1965) 

Subfarnily EURYGENIINAE 

Tlibe Eurygeniini 

BC 

S. columbianus Hopping 

S. guttatus LeConte 

impressicollis Casey 

subalbicans Casey 

S. hirtus Hatch 

S. rufipes C:asey 

S. vestitus Say) 

bad,iipennis LeOonte 

__BC_-_-_ 

BC 

BC AB 

SK MB ON 

po 

PQ 

274

Scraptiidae 

Genus MACRATRIA Newman 

M. confusa LeConte 

Tribe Macratriini 

ON PQ 

Family ADERIDAE antlike leaf beetles 

Aderidae is a small family that includes nine 

Y. Bousquet 

Except for Werner's (1990) recent review of the 

species within the borders of Canada. Very little is eastern North American species, members of this 

known about these beetles. The adults are usually family has been little studied taxonomically. 

found on foliage and flowers; the larvae of some 

species have been found in rotten wood and leaf litter. gC (1); ON (8); PQ (8); NB (1); NS (1) ' I (1) 

Genus ELONUS Casey 

E. basalis (Leconte) - ON PQ 

princeps Casey 

E. nebulosus (LeConter - ON PQ 

Genus ADERUS Westwood 

Phomalus Casey 

A. populneus (Panzertt - BC - ON PQ 

Genus VANONUS Casey 

Tanilotes Casey 

V. calvescens Casey PQ 

V. piceus (LeConte) - ON 

congener Casey 

fusciceps Casey 

t ub er c ulifer ( Hamilton) 

V. vigilans Casey - ON PQ 

V. wickhami Casey - ON PQ NB NS 


Genus ZONANTES Casey 

Z. fasciatus (Melsheimert - ON PQ 

tricuspis Casey 

Z. pallidus Werner - ON PQ 

Family SCRAPTIIDAE scraptiid beetles 

L. LeSage 

The family Scraptiidae, which comprises about 

There is no recent treatment of the family in 

400 species around the world and 21 in Canada and North America; the Canadian fauna is poorly 

Alaska, represents a group of beetles with uncertain known except in the provinces of Quebec and 

affinities. Previously placed in Mordellidae, or British Columbia. Young (1976) transferred the 

Melandryidae, or Anthicidae, they are now consid- genus Pectotom.a, previously included in 

ered to be a separate farnily closely related to those Tetratomidae, to Scraptiini. 

families (Crowson 1955; Franciscolo 1963; Lawrence 

I982a). Adults are often found in large numbers on 

flowers, and larvae occur in decaying wood or in leaf AK (2); yK (1); gC (16); AB (5); SK (1); MB (1); ON 

litter. (8);PQ (10); NB (3);NS (2); NF (3) 

275

Scraptiidac 

Subfamily ANASPIDINAE 

Tribe Anaspidini 

Key to North American species: Liljeblad (1945) 

Genus ANIiSPIS Geo{Iroy 

A. atrata Champion 

atra LzConte 

A. duryi Liljeblad 

A. flavipennis Haldeman 

A. nigrina Csiki 

zlgro (Haldeman) 

A. rayi Hat,ch 

A. rufa Say 

dimidiata Melsheimer 

filiforrnis LeConte 

nigrice:ps LeConte 

pallescens Mannerherm 

uenlro'is Melsheimer 

A. seposita Liljeblad 

A. sericea ( \{annerheim) 

luteipennis LeConte 

AK 

AK 

YK 

_BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

ON 

ON 

PQ 

PQ 

ON PQ 

NB 

NB 

lls - 

NF 

NF 

Genus SILARIA Mulsant 

S. revelstokei Liljeblad 

BC 

Genus LARISIA Emery 

L. nigricolo r Liljeblad 

BC 

AB 

ON 

Genus NASSIPA Emery 

N. hopping Liljeblad 

BC 

Subfamily SCRAPTIINAE 

Tlibe Allopodini 

Genus ALLOPODA LeOonte 

A. californir:a Schaeffer 

A. lutea (Haldeman) 

fuscos ut ur alis (Blatchley) 

BC 

Tlibe Scraptiini 

Genus SCRAPTIA Latreille 

S. sericea t lylelsheimer) 

PQ 

Genus CAI IFA LeConte 

C. pallipennis LeConte 

C. pailipes' Melsheimert 

biimpressa (Haldeman) 

C. plagiata ( Melsheimert 

americana (Haldeman) 

C. pusilla (I{aldeman) 

BC 

MB 

ON 

ON 

ON 

PQ 

PQ 

PQ 

PQ 

NB 

NS 

NF 

Genus PECTOTOMA Hatch 

P. hoppingi Hatch 

BC 

AB 

T?ibe Pentariini 

Genus PENTARIA Mulsant 

Anthobates LeOonte 

Anthobatula Sftalrd. 


P. fuscula (l,eConte) 

P. trifasciata Melsheimer 

nubila (LeConte) 

Liljeblad (1945) 

BC 

BC AB SK 

-ONPQ 

PQ 

276

Cerambycidae 

Superfamily Chrysomeloidea 

Family CERAMBYCIDAE longhorned beetles 

J. McNamara 

Cerambycidae is represented in Canada and 

Alaska by 354 species and subspecies; it is one of 

the largest groups of wood-boring insects. These 

beetles have always been popular with collectors 

because of their bright colors and the large size of 

adults of many species. The adults generally feed 

on wood, roots, leaves, and pollen. Flower-feeding 

adults may visit many different blossoms, but some 

are host-specific; they are important plant-pollination 

agents. The larvae bore into wood and roots; 

the host wood may be living or thoroughly decayed. 

Some may even totally destroy timber for commercial 

purposes. Cerambycids play an important role 

in forest ecolory because they help in the decomposition 

ofdead and dying trees. 

Thanks to the works of Linsley (1962a, 1962b, 

1963, 1964) and Linsley and Chemsak (1972, 1976, 

1985), the North American fauna is taxonomically 

well known; only the second part of the subfamily 

Lamiinae remains to be done. Chemsak and Linsley 

(1982) also published a checklist of the North 

American cerambycids. 

AK (31); YK(27); NT (32); BC (145); AB (88); SK 

(56); MB (100); ON (20e); PQ (178); NB (68); NS 

(51);PE (11);LB (13);NF (24) . H (4);I (1) 

Genus PARANDRA Latreille 

(Subgenus NEANDRA Lameere) 

P. brunnea brunnea (Fabricius) 

ampliceps Casey 

conformis J. Thomson 

dentata J. Thomson 

grauidula Casey 

minuta J. Thomson 

purpureus (Herbst) 

q uad.r icoll is J. Thomson 

sayi J. Thomson 

P. brunnea quebecensis Abdullah 

Genus ERGATES Audinet-Serville 

Trichocnemis LeConte 

Subfamily PARANDRINAE 

Tribe Parandrini 

Subfamily PRIONINAE 

Tribe Ergatini 

oN PQ 

PQ 

E. spiculatus spiculatus (LeConte) 

californicus (A. White) 

- BC 

spiculigerus (A. White) 

Genus ORTHOSOMA Audinet-Serville 

O. brunneum (Forster) 

amplians Casey 

cy lindr icum (Fabricius ) 

cylindroides (Gmelin) 

pennsyluanicum (DeGeer) 

spadix Casey 

unicolor (Drury) 

Genus PRIONUS Geoffroy 

(Subgenus PRIONUS s. str.) 

(Prionellus Casey) 

Tlibe Prionini 

P. californicus Motschulsky AK BC 

acomanus Casey 


oN PQ 

NS 

277

Cerambycidee 

ambiguus Casey 



compar Casey 

consors Casey 

crassict>rnis LeConte 

curuatr^.s LeConte 

fissifrons Casey 

horni l.ameere 

humerc:lis Casey 

ineptus Casey 

nanus i)asey 

orbiceps Casey 

ouipenr:is Casey 

paruicollis Casey 

proxim,zns Casey 

punctui',atus Casey 

scutellaris Casey 

seniger Casey 

solidus Casey 


spiculosus Casey 

stultus Casey 

suspectus Casey 

terminulis Casey 

texanus: Casey 

uintanus Casey 

P. laticollis Druryt 

b reu ico." nis Fabricius 

densus Casey 

frosti C'.asey 

hempi Casey 

nigrescans Casey 

oblongu.s Casey 

paruus Casey 

P. lecontei I,ameere 

P. pocularis Dalman 

bicolor Casey 

curticornis LeConte 

laeuigatus Hanis 

obliqui:ornis LeConte 

prolixus Casey 

ualidiceps Casey 

(Subgenus I{EOPOLYARTHRON Semenov) 

(Riporuts Casey) 

P. imbricorrris (Linn6) 

brunneus Casey 

cuneatus Casey 

diuersus Casey 

mimus Casey 

robustu,s Casey 

(Subgenus III.{TENNAJ,IA Casey) 

P. fissicorni s Haldeman 



transuersus Casey 

BC 

oN PQ 

oll pe - 

ON 

ON 

Tribe Tragosomini 

Genus TRATIOSOMA Audinet-Serville 

T. depsariurn (Linn6)* 

harisii LeConte 

paruicc'lle Casey 

repens Casey 

sodale Casey 

spiculu.tn Casey 

NT BC SK MB ON PQ NB NS NF 

278

Cerambycidae 

Genus SPONDYLIS Fabricius 

S. upiformis Mannerheim 

basalis Casey 


laticeps LeConte 

parua Casey 

robustula Casey 

subpubescens Casey 

Subfamily SPONDYLINAE 

AK 

BC AB 

ON PQ 

Subfamily ASEMINAE 

TYibe Asemini 

Genus ARHOPAJ-US Audinet-Serville 

Criocephalum Dejean 

Criocephalus Mulsant 

A. asperatus (LeConte) 

filitarsis (Casey) 

A. foveicollis (Haldeman) 

agrestis (Kirby) 

ceruinus (Casey) 

lacustrinus (Casey) 

A. productus (LeConte) 

elongatus (Casey) 

propinquus (Casey) 

A. rusticus obsoletus (Randall) 

mexicanus (J. Thomson) 

obscurus (LeConte) 

AK YK 

NT BC 

NT 

BC 

AB SK 

AB SK 

AB 

MB ON PQ 

ON 

NB NS 

LB 

NF 

Genus MEGASEMUM Kraatz 

M. asperum (LeConte) 

impressum (Casey) 

rectipennis (Casey) 

uteanum (Casey) 

BC 

AB 

Genus ASEMUM Eschscholtz 

Liasetnum Casey 

A. australe LeConte 

c ham p lainianum (C asey) 

A. caseyi Linsley 

A. nitidum LeConte 

mokelumne Casey 

A. striatum (Linn6)8 

amputatum Casey 

atrum Eschscholtz 

breuicorne Casey 

brunneum Haldeman 

carolinum Casey 

costulatum Casey 

curtipenne Casey 

ebenutn Casey 

fuluipenne Casey 

fuscutn Haldeman 

gracilicorne Casey 

juuencum Haldeman 

moestum Haldeman 

obsoletum Haldeman 

paruicorne Casey 

pugetanum Casey 

stoctonense Casey 

substriatum Haldeman 

YK 

NT 

BC 

BC 

MB ON 

PQ 

BC AB SK PQ NB 

NF 

Genus TETROPIUM Kirby 

T. abietis FaIl 

T. cinnamopterum Kirby 

T. parvulum Casey 

alaskanumFall 

-BC 

AKYKNTBCAB_MB 

AKYKNTBCAB_MB 

ON 

ON 

PQ 

PQ 

NS-- 

LB 

NF 

279

Cerambycii.ae 

T. schwarzilnum Casey 

T. velutinurn LeConte 

boreak Casey 

columlianum Casey 

hexagonum Casey 

---ncAB--oNPQ----y 

Genus ATItr{IA Haldeman 

Tlibe Atimiini 

A. confusa confusa (Say) - ON 

frislls l{aldeman 

A. confusa corsalis LeConte - BC 

A. hoppingi Linsley - BC 

Genus OPSIMUS Mannerheim 

Subfamily CERAMBYCINAE 

Tribe Opsimini 

O. quadrilineatus Mannerheim AK BC 

biplectralis Casey 

Genus DICI)NTRUS LeConte 

Paraopsimus Champlain & Knull 

D. bluthner LeConte - BC 

Genus SMODICIIM Haldeman 

S. cucujiforrre (Say) 


cy lindr;;d.e s (Newrnan) 

TYibe Smodicini 

- ON 

Genus TYLONOTUS Haldeman 

Tlibe Hesperophanini 

T. bimacular,us Haldeman MB ON PQ 

Genus HESIIEROPHANES Mulsant 

AnoplitLm Haldeman 

H. pubescens {Haldeman) - ON PQ 

Genus EBU:RIA Audinet-Serville 

E. quadrigerninata (Say) - ON 

Genus PSYI|,ASSA Pascoe 

PseudiL,idion Casey 

Tribe Elaphidiini 

P. unicolor (.Randallt - ON PQ - NS 

p loch ionoc era (Haldeman) 

Genus STEIIOSPHENUS Haldeman 

S. notatus (Olivierr 

deflend,zs (Newman) 

- ON pQ 

discicol,:!.is (Dej ean) 

discoiderus (Sturm) 

pinorunz Casey 

Genus ENAIIIIALODES Haldeman 

Hy pe rm a ll us Lacordaire 

Romaleum A. White 

Thersalus Pascoe 

280

Cerambycidae 

E. atomarius (Drury) 

bispinosus (Pascoe) 

mary landicus (Fabricius) 

multicolor (Gmelin) 

p ulu er ule ntus (DeGeer) 

thoracicurn (Casey) 

uariegatus (Goeze) 

E. cortiphagus (Craigheac, 

E. rufulus (Haldeman) 

crinicornis (Casey) 

mancus (Casey) 

Genus PARELAPHIDION Skiles 

Key to species: Skiles (1985) 

P. incertum (Newman) 

breuicorne (Casey) 

neglectum (LeConte) 


Genus ANELAPIIUS Linsley 

E I ap hidionoides Linsley 

A. parallelus (Newman) 

orclzs (Newman) 

canadensis (Casey) 

defectus (Casey) 

densicollis (Casey) 

medialis (Casey) 

molliculus (Casey) 

oblitus (LeConte) 

p ulu erulentus (Olivier) 

A. villosus (Fabricius) 

abruptus (Casey) 

breuiusculus (Casey) 


militaris (Casey) 

p r uinos u s (Gu6rin-M6neville) 

putator (Peck) 

scuticularis (Casey) 

Genus ELAPHIDION Audinet-Serville 

Cycliopleurus Hope 

E. mucronatum (Say) 

muriaticum (Haldeman) 

muricatum Haldeman 

neb ulosum Gu6rin-M6neville 

uestitum Haldeman 

ON 

ON 

ON 

ON 

MB ON 

MB ON 

ON 

PQ 

PQ 

PQ 

Tlibe Ibidiini 

Genus HETERACHTHES Newman 

Heteracthon J. Thomson 

Key to species: Martins (1970) 

H. quadrimaculatus Haldeman 

euanescens (J. Thomson) 

- ON 

fluuiatilis Casey 

pallidus Haldeman 

PQ 

Tlibe Obriini 

Genus OBRIUM Dejean 

Diozodes Haldeman 

Phyton Newman 

O. maculatum (Olivier) 

howdeniRnrTl 

limum (Newman) 

pallidum (Say) 

O. rufulum Gahan MB ON PQ 

28r

Cerambycide.e 

Tlibe Hyboderini 

Genus HYBODERA LeOonte 

H. tuberculaua LeConte 

Genus LAMPROPTERUS Mulsant 

Callitne.llum Sftand 

Callitnus Mulsant 

Pilema.LeConte 

L. ruficollis (LeConte) 

longicol'.is (Casey) 

opacipennis (Casey) 

_BC 

_BC 

Tfibe Eumichthinr 

Genus EUMI.CHTHUS LeConte 

E. oedipus LcConte 

ater Hardy & Preece 

- BC 

funebrus (Van Dyke) 

malkini (Fisher) 

ruber H,*dy & Preece 

Genus POECIILOBRIUM Horn 

P. chalybeunr (LeConter 

gibsoni [Iopping 

- BC 

minuturn Casey 

rugosipt:nne Linell 

Genus MOLORCIIUS Fabricius 

Glaphyra Newman 

Laphyrt,. Newman 

Tlibe Molorchini 

M. bimaculal;us bimaculatus Say AB SK MB ON PQ 

affinis l.eCorrte 

obscuru:; (LeConte) 

M. longicollis LeConte - BC 

Genus CALLIMOXYS Kraatz 

Tribe Stenopterini 

C. sanguinicollis luscipennis 

(LeCcnte) 

- BC 

C. sanguinic,rllis sanguinicollis 

(Olivier) 

- ON PQ 

pinorurr;, Casey 

Genus ROSALIA Audinet-Serville 

Tlibe Rosaliini 

R. funebris l\lotschulsky AK BC 

acorrrano, Casey 

mandibitlaris Casey 

Genus PRONOCERA Motschulskv 

Gonocallus LeConte 

Tlibe Callidiini 

P. coilaris coilaris (Kirby) AK YK NT BC AB SK MB ON PQ NB - NF 

lepida (LeConte) 

Genus HYLOTRUPES Audinet-Serville 

282

Cerambycidae 

H. bajulus (Linn6)t 

bullatus Haldeman 

inaequalis Casey 

incertus Casey 

latus Casey 

minutus Casey 

nematocerus Casey 

pedicellatus Casey 

puncticollis Casey 

PQ 

Genus SEMANOTUS Mulsant 

Anocomis Casey 

Hemicallidium Casey 

S. amethystinus (Leconte) 

S. ligneus amplus (Casey) 

angustus (Casey) 

obliuiosus (Casey) 

terracensis (Casey) 

S. ligneus basalis (Casey) 

thujaeYanDyke 

S. ligneus ligneus (Fabricius) 

nicolas (A. White) 

russicus (Stephens) 

S. litigiosus (Casey) 

canadensis (Casey) 

terminatus (Casey) 

AK 

_BC 

NT 

BC 

BC 

MB ON PQ 

BC AB SK MB ON PQ NB 

NS 

Genus MERIELLUM Linsley 

Merium LeConte 

M. proteus (Kirby) 

bifossulatum (Casey) 

simile (I{trby) 

AK YK 

NT 


Genus CALLIDIUM Mulsant 

Meridion des Gozis 

C. antennatum hesperum Casey 

C. cicatricosum Mannerheim 

subopacum Swaine 

udncouuerense Van Dyke 

C. frigidum Casey 

lacustre Casey 

C. hoppingi Linsley 

C. schotti Schaeffer 

C. violaceum (Linn6)t 

janthinum LeConte 

-BC 

AKYK_BCAB 

oN PQ 

NN NS 

NF 

Genus PHYMATODES Mulsant 

Microcallidium Casey 

Phymatod,ina Casey 

Poecilium Fairmaire 

P. aeneus LeConte 

P. aereus (Newman) 

pallipes (Haldeman) 

P. amoenus (Say) 

coccineico llis (Haldeman) 

hudsonicus Casey 

P. ater LeConte 

P. decussatus decussatus (LeConte) 


harfordi Casey 

latifasciatus Hardy & Preece 


P. dimidiatus (Kirby) 

haltnii (Schon]nen) 

manner h eimii (LeConte) 

palliatus (Haldeman) 

P. fulgidus Hopping 

P. hirtellus densipennis Casey 

P. lecontei Linsley 

BC 

BC 

AK YK AB 

_BCPQ 

_BCAB 

_BC 

oN pe 

oN PQ 

BC SK MB ON PQ NB 

_NS 

283

Cerambycidae 

grandi; Casey 

obscur,ts (LeConte) 

P. maculico.lis LeConte - YK - BC PQ NB 

elonga;:us Hopping 

hard,yi (YanDyke) 

P. nitidus f.eConte - BC 

crucialis Casey 

elegant Casey 

exilis Qasey 

P. testaceus tl-innerf 

uariabilis (Linn6) 

- ON PQ 

uentralis Haldeman 

P. varjus tFabricius) 

albofasciatus (Bland) 

- ON PQ 

diffidens Casey 

latipennis Casey 

s emic ir c ular is (Blano.r 

subfast:iellus (A. White) 

uersicoJor (Gmelin) 

P. vulneratrrs (LeConte) 

concinnus Casey 

- BC 

nigrescens Hardy & Preece 

Genus PHYSOCNEMUM Haldeman 

Dulari,ts J. Thomson 

P. andreae tHaldeman) - ON 

luscum (J. Thomson) 

P. breviline,rm t Sayt MB ON PQ 

angulatum Casey 

compressipes Casey 

densun't Casey 

longitarse Casey 

pictum Casey 

P. violaceip'rnne HamilLon 

Genus ROP AJ-OPUS Mulsant 

Euryotr tera Horn 

R. sanguini.:ollis (Horn r MB ON PQ NB 

Genus ELA'IOTRYPES Fisher 

E. hoferi Fir,her AB 

Genus XYLTJCRIUS LeOonte 

X. agassizi {Leconter - BC 

Tribe Clytini 

Genus MEC'.ACYLLENE Casey 

Cyllene Newman 

M. angulifera (Casey) 

M. caryae (Gahan) 

M. decora (Olivier) 

cas teln,zui (Aurivillius) 

chara (Say) 

genitiuo (Casey) 

horni (.A,urivillius) 

infausta (LeConte) 

hansana (Casey) 

mediana (Aurivillius) 

solida (Casey) 

M. robiniae (Forster) 

angulata (Fabricius) 

flexuos'z (Fabricius) 

picta (Drtry) 

Genus GLYOOBIUS LeConte 

G. speciosus (Say) 

haiji (Gtay) 

AB SK MB 

-oN 

AB_MB 

_ONPQ 

_ONPQ 

284

Cerambycidae 

Genus CAI-LOIDES LeConte 

C. nobilis nobilis (Harris) 

MB ON PQ 

Genus SAROSESTHES J. Thomson 

S. fulminans (Fabricius) 

ON 

PQ 

Genus XYLOTRECHUS Chevrolat 

X. aceris Fisher 

X. annosus annosus (Say) 

X. annosus emotus W.J. Brown 

X. colonus (Fabricius) 

agrestis (LeConte) 

campestris (OIivier) 

te rrnina n s (Fabricius ) 

upsilon (Stephens) 

uillicus (Olivier) 

X. integer (Haldeman) 

frosli Van Dyke 

X. lengi Schaeffer 

X. longitarsis Casey 

columbianus Casey 

inflaticollis Casey 

montanicus Casey 

X. mormonus (LeConte) 

X. quadrimaculatus (Haldeman) 

X. sagittatus sagittatus (Germar) 

cylindrus Casey 

pubescens (Haldeman) 

salebrosus Casey 

X. schaefferi Schott 

X. undulatus (Say) 

fuscus (Kirby) 

interruptus (Castelnau & Go.y) 

lunulatus (Kirby) 

soyi (Castelnau & Gory) 

undatus (Kirby) 

Genus NEOCLYIUS J. Thomson 

R ho p alop achys Chevrolat 

R ho p alome ru s Chevrolat 

N. acuminatus acuminatus (Fabricius) 

americanus (Gmelin) 

aspericollis (Germar) 

erythrocep halus (Fabricius) 

N. balteatus LeOonte 

N. caprea (Say) 

eleuatus (Castelnau & Go.y) 

gibbicollis (Castelnau & Gory) 

N. conjunctus (LeConte) 

occiduus Casey 

tularensis Casey 

N. fulguratus Casey 

confusus Van Dyke 

N. mucronatus mucronatus 

(Fabricius) 

humeralis (Newman) 

luscus (Fabricius) 

tnaculosus (Gmelin) 

strictulus Casey 

N. muricatulus muricatulus (Kirby) 

&lr6yi Aurivillius 

leucozonus (Castelnau & Gory) 

longipes (Kirby) 

N. provoanus Casey 

basalis Van Dyke 

basillaris Van Dyke 

Genus CLYTUS Laicharting 

C. canadensis Hopping 

YK 

NT 

-BC 

BC 

AB 

_BC 

NT BC AB 

AB 

SK 

SK 

_ONPQ 

MB ON PQ 

MB ON PQ 

NB 

NB 

ON PQ NB NS PE 

MB 

MB ON PQ 

MB ON PQ 

NB 

MB ON PQ NB 

ON 

ON 

PQ 

NS--- 

AK Yr( Nr ec AB sx MB 3N FB - Ns pn is Nr' 

BC 

BC 


BC 

BC 

AB 

285

Cerambycirlae 

C. margini

Cerambycidae 

Genus BATYLE J. Thomson 

Batyleoma Casey 

B. suturaiis suturalis (Say) 


miniata (Germar) 

rubro (Newman) 

B. suturalis pearsalli (Bland) 

B. ignicollis ignicollis (Say) 

acotnana Casey 

punctipennis Casey 

sang uinicoll is ( Germar) 

seclusa Casey 

AB 

SK 

_ABSKMB 

_ 

^AB 

ON 

PQ 

Genus CROSSIDIUS LeConte 

Oxoplus LeConte 

C. carolinus (LeConte) 

C. discoideus discoideus (Say) 

pulchrior Bland 

C. discoideus blandi Casey 

C. pulchellus LeConte 

debilis Casey 

lanaticollis Casey 

nitidicollis Casey 

palmeri Bates 

retractus Casey 

sinuosus Casey 


C. punctatus LeConte 

crossipes Casey 

hirsuticollis Casey 


uinosus Casey 

BC 

AB 

-{B 

AB 

SK 

Genus ELYTROLEPTUS Dug6s 

E. floridanus (LeOonte) 

ON 

Tribe Holopleurini 

Genus HOLOPLEURA LeConte 

H. marginata LeConte 

helena LeConte 

BC 

Subfamily LEPTURINAE 

Tlibe Desmocerini 

Genus DESMOCERUS Audinet-Serville 

D. auripennis auripennis Chevrolat 

D. auripennis cribripennis Horn 

D. auripennis piperi Webb 

D. palliatus (Forster) 

blandus (Fabricius) 

cyaneus (Fabricius) 

elongatus Bland 

ON 

pe 

NB 

\TIT 

Genus NECYDAI-IS Linn6 

N. cavipennis LeConte 

N. diversicollis diversicolhs 

Schaeffer 

N. laevicollis LeConte 

seminiger Linsley 

N. mellita (Say) 

annericanus Haldeman 

Mbe Necydalini 

BC 

BC 

BC 

ON 

287

CerambyciJae 

Genus ULOCIIAETES LeOonte 

U. leoninur; LeConte 

BC 

Tlibe Lepturini 

Genus CEI{TRODERA LeConte 

Parapachyta Casey 

C. decolorata (Harris) 


rubidct (Haldeman) 

C. spurca (LeOonte) 

ceruin.z. (Walker) 

BC 

oN PQ 

NB NS 

Genus STEiNOCORUS Fabricius 

Anisorus Mulsant 

Eutoxotus Casey 

Toxotc,psis Casey 

ToxottLs Dejean 

S. cylindricollis (Say) 

atratus (Haldeman) 

dentip ennis (Haldeman) 

diues (Newman) 

S. fl avolineatus (LeConte) 

S. nubifer (LeConte) 

apiciu,:ntris Casey 

hesperus Casey 

lateratis (Casey) 

margi:tellus Casey 

pacifit us Casey 



rufipe,znis Casey 

spinosus (Hopping) 

tenellt..s Casey 

truncctulus Casey 

S. obtusus .LeConte) 


S. schaumii(LeConte) 

croceus (Leng) 

S. trivittatus (Say) 

S. vestitus (Haldeman) 

ater (L,eng) 

aureatus Casey 

flaccia'.us Casey 

giluicarnis Casey 


sericai.us Casey 

subpir;guis Casey 

uirgat,rc (LeConte) 

S. vittiger (Randall) 

nigrip,zs (Haldeman) 

_BC 

_BCAB 

BC 

AB 

_ABSK 

BC AB 

SK 

MB ON PQ 

MB 

oN PQ 

Genus ENCYCLOPS Newman 

E. caerulea (Say) 

pallipes Newman 

ON PQ 

Genus PYR,OTRICHUS LeConte 

P. vitticollis LeConte 

cribripennis Casey 

BC 

Genus PACHYTA Dejean 

Argaleus LeConte 

Linsle;lana Podany 

P. armata LeConte 

P. lamed lit,urata Kirby 

amerit:ana Podany 

confl ag1 r ata Motschul sky 

nitens (LeConte) 

_BC 


meoNpe: 

LB 

NF 

288

Cerambycidae 

Genus EVODINUS LeConte 

E. monticola monticola (Randall) 

carolinensis Casey 

E. monticola vancouveri Casey AK 

NT 

BC 

MB ON PQ 

NB NS 

LB 

NF 

Genus ANTHOPHYLAX LeOonte 

A. attenuatus (Haldeman) 

A. cyaneus tHaldeman) 

malac hitic u s (Haldeman) 

A. viridis LeConte 

uiridipennis Casey 

oN PQ 

oN PQ 

-PQ 

NB NS 

NB 

NS 

Genus NEANTHOPHYLAX Linsley & Chemsak 

N. mirificus (Bland) 

Denustus (Bland) 

N. tenebrosus nigrolineatus 

(Van Dyke) 

BC 

BC 

AB 

Genus RHAGIUM Fabricius 

Allorhagium Kolbe 

Hargium Samouelle 

Stenocorus Geoffroy 

R. inquisitor (Linndtx 

americanurn Podany 

boreale Casey 

californicum Casey 

canadense Podany 

cariniuentre Casey 

crosslpes Casey 

i nuesliga lor Mannerheim 

lineatus (Olivier) 

mexicanurn Casey 

montanum Casey 

paruicorne Casey 

quadricostatum Podany 

thoracicum Casey 

YK 

NT BC 

AB 


NF 

Genus SACHALINOBIA Jacobson 

Pseudopachyta Swaine & Hopping 

S. rugipennis rugipennis (Newman) 

MB ON PQ NB 

Genus GAUROTES LeConte 

G. cyanipennis tSayt 

ioze (Newman) 

lapor te i ( Gu6rin-M6neville) 

leonardi (Haldeman) 

ser u illei (Audinet-Serville) 

ON PQ 

Genus ACMAEOPS LeOonte 

A. proteus proteus (Kirby) AK 

aurora Casey 

cauicollis Casey 


gibbula LeConte 


obsoleta Casey 

parkeri Casey 

pinicola Schaeffer 

puncticeps Casey 

scutellata Casey 

sublineata (Haldeman) 

MB ON PQ NB 


LB 

Genus LEPTALIA LeConte 

L. macilenta (Mannerheim) AK 

frankenhaeuseri (Mannerheim) 

fuscicollis (LeConte) 

BC 

AB 

289

Cerambycidae 

Genus CO IITODERA Mulsant 

Acma,zopsilla Casey 

Lepta.:maeops Casey 

C. coniferae Hopping & Hopping - BC 

C. longicornis {Kirbyt - BC AB SK 

alticoia (Casey) 

6oher,' Hopping & Hopping 

dorsa,',is (LeConte) 

harneyensis Hopping & Hopping 

ligata (LeConte) 

marginalis (LeConte) 

oregonensis Hopping & Hopping 

punctiuentris (Casey) 

suturalis (Casey) 

uincta (LeConte) 

C. militari'r militaris (LeConr,eJ - BC 

C. subpilosa (Leconte) - BC AB 

intermedia (Casey) 

lupino (LeConte) 

pugetuna (Casey) 

quadriceps (Casey) 

Genus PSEIIDOGAUROTINA Plavilstshikov 

P. abdomirralis (Bland I - ON PQ 

P. cressoni cressoni (Bland) - BC AB 

Genus BR.I\CHYSOMIDA Casey 

B. atra (Leconte) 

aterriina Casey 

- BC AB 

morata Casey 

B. bivittata (Say) AB SK MB ON PQ NB NS 

fuscicttps (LeConte) 

incert r (Bland)' 

nigrip,ennis (LeConte) 

oblonga (Casey) 

uariarts (LeConte) 

Genus GN.{.THACMAEOPS Linsley & Chemsak 

G. pratens s (Laicharting)* AK YK NT BC AB SK MB ON PQ NB LB NF 

fu lu i p ennis (Mannerheim) 

longiceps (Kirby) 

s em i - marg inara (Randall) 

s t rig il ata (Fabricius) 

Genus ME'IACMAEOPS Linsley & Chemsak 

M. vittata Swederus) PQ 

direcfrz (Newman) 

pallidc (Haldeman) 

q uadr iu ittata (Schonherr) 

Genus STIIANGALIA Audinet-Serville 

Ophistomis J. Thomson 

S tr anil alina Auriviilius 

S. acuminz.ta (Olivier) - ON 

emaciata Newman 

unicolor Haldeman 

S. bicolor (:Swederus) - ON 

simulons (Casey) 

S. famelica solitaria Haldeman - ON 

S. luLeicorrris (Fabriciust - ON PQ 

carolinae (Casey) 

euersa (Casey) 

Genus BELLAMIRA LeConte 

B. scalaris (Say) - SK MB ON PQ NB NS 

coarcto.ta (Haldeman) 

290

Cerambycidae 

Genus ANALEPTURA Linsley & Chemsak 

A. lineola (Say) - ON PQ NB 

cincta (Haldeman) 

indirecta (Newman) 

lateralis (HaIdeman) 

obsoleta (Haldeman) 

Genus CHARISALIA Casey 

C. americana (Haldeman) - ON 

Genus GRAMMOPTERA Audinet-Serville 

Parallelina Casey 

G. exigua (Newman) - ON PQ 

saucia (LeConte) 

G. haematites (Newman) - SK MB ON PQ NB 

zono (Newman) 

ruficeps (LeConte) 

G. molybdica {Leconte) - BC 

G. subargentata (Kirby) AK YK NT BC AB SK MB ON PQ NB 

filicornis (Casey) 

quebecensis (Couper) 

rufi.basis (LeConte) 

similis (I\trby) 

Genus JUDOLIA Mulsant 

J. gaurotoides gaurotoides (Casey) AB SK 

hnulli (Swaine & Hopping) 

uiuaria Casey 

J. instabilis (Haldeman) - YK - BC AB SK MB 

conuexa (LeOonte) 

fl au iu entris (Schaeffer) 


trajecta Casey 

J. montivagans montivagans (Couper) AK YK NT BC AB SK MB ON PQ NB NS - LB 

seminigra Casey 

J. quadrata (LeConte) - YK NT BC - SK MB ON PQ 

Genus PYGOLEPTURA Linsley & Chemsak 

P. carbonata (LeConte) - BC 

P. nigrella nigrella (Say) AK - NT BC AB SK MB ON PQ NB LB NF 


nigrita (HaLch) 

praestans (Casey) 

serricornis (Casey) 

Genus DORCASINA Casey 

D. matthewsi (LeConte) - BC 

macrocera (Casey) 

Genus IDIOPIDONIA Swaine & Hopping 

l. pedalis (LeConte) - ON PQ NB NS 

Genus TYPOCERUS LeConte 

T. acuticauda acuticauda Casey - ON PQ NB NS 


T. balteatus Horn NT - AB 

T. confluens Casey MB 

caligans Casey 

T. lugubris tSayt - ON 

T. octonotatus (Haldeman) - ON 

arapahoe Casey 

sinuatus LeConte 

T. sparsus LeConte MB ON PQ 

T. velutinus velutinus (Olivier) MB ON PQ NB NS 

fugo* (Fabricius) 

manitobensis Swaine & Hopping 

tenuior (Kirby) 

297

Cerambycidae 

T. zebra (Olivier) 

aurige" (Newman) 

carolir,La (Weber) 

zebratus (Fabricius) 

ON 

Genus NEOALOSTERNA Podany 

N. capitata {Newman) 

N. rubida (l,eConte) 

heeni (Casey) 

BC 

_9* 

PQ NB NS 

Genus PSEUDOSTRANGAIIA Swaine & Hopping 

P. cruentat a (Haldemant 

maneii'. (Casey) 

ON 

PQ 

Genus LEPTURA Linn6 

(Subgenus LEPTURA s. str.) 

L. obliterata obliterata (Haldeman) 

idahoensis (Casey) 

perductor (Walker) 

uitiosa (LeConte) 

L. plagifera LeConLe 

L. plebeja Flandall 

L. propinquaBland AK 

minuscula (Casey) 

regularis (Casey) 

L. subhami.ta Randall 

annatfl (Haldeman) 

elegan; (Haldeman) 

interrt..pta Newman 

BC 

BC 

BC 

yuoNpeNe_ 

ON PQ NB 

NS 

(Subgenus iTENURA Dejean) 

(Mega,i.eptura Casey) 

L. emargin,rta Fabricius 

ON 

Genus STENOSTROPHIA Casey 

S. amabilis {Leconte) 

S. tribalteaba serpentina (Casey) 

BC 

BC 

Genus TRACIfYSIDA Casey 

T. aspera aspera (LeConte) AK 

parkeri (Hippisly) 

T. aspera b:revifrons (H. Howden) 

T. mutabili; (Newman) 

I ae tifi

Cerambvcidae 

(Subgenus CERATOPIDONIA Linsley & Chemsak) 

P. quadrata Hopping - BC 

Genus ORTHOLEPTURA Casey 

O. valida (Leconte) - BC 

oculea Casey 

Genus STRANGAI-EPTA Casey 

S. abbreviata (Germarr 

gulosa (Kirby) 

- ON PQ NB NS 

nitidipe nnis ( Provancher) 

saratogensis (Rat) 

semiuittata (Kirby) 

S. pubera tSay) MB ON PQ 

Genus STICTOLEPTURA Casey 

S. canadensis canadensis (Olivier) MB ON PQ NB NS PE - NF 

c innamop te r a (Haldeman) 

erythroptera (Kirby) 

te nuic or n is (Haldeman) 

S. canadensis cribripennis (LeConte) 

coccinea (LeConte) 

- BC AB SK MB 

diuisa (Casey) 

ebena (Leng) 

Genus ANASTRANGAJ-IA Casey 

A. laetifica (LeConte) - BC 

lugens (LeConte) 

A. sanguinea (LeOonte) AK YK - BC AB SK MB ON PQ NB 

apicata (Casey) 

bould.erensis (Casey) 

Genus XESTOLEFTURA Casey 

X. behrensi (LeConte) - BC 

X. crassipes (LeContet - BC 

fas ciu en tr is ( LeConte) 

muliebris Casey 

shastana Casey 

uancouueri Casey 

tanthogaster (LeConte) 

X. octonotata (Sayt - ON 

q uadr ip unctato ( Haldeman) 

stictica (Newman) 

X. tibialis (LeConte) - \'K - BC AB ON PQ - NS - LB NF 

columbica Casey 

hirtella (LeConte) 

miquelonensis (Pic) 

pictipennis Casey 

Genus LEPTUROPSIS Linsley & Chemsak 

L. biforis (Newman) - ON PQ NB 

laurentica (Casey) 

L. dolorosa (Leconte) - BC 

Genus TRIGONARTHRIS Haldeman 

Cercolia Casey 

T. minnesotana (Casey) MB ON PQ NB NS 

T. proxima (Sayt MB ON PQ - NS 

hempiana (Casey) 

subpubescens (Kirby) 

Genus ACMAEOPSOIDES Linsley & Chemsak 

A. rufulus (Haldeman) - BC AB SK MB ON PQ - NS 

Genus BRACHYLEPTURA Casey 

B. champlaini Casey - ON PQ - NS PE 

293

Cerambycidae 

B. dehiscens (LeConte) 

B. rubrica ([iay) 

erythraptera (Germar) 

B. vexatrix, Mannerheim) 

conuoluens (Casey) 

quadrii,lum (LeConte) 

AK 

BC 

BC 

ON 

PQ 

NB 

Genus STROPHIONA Casey 

S. laeta (Le(lonte) 

ostenta Casey 

S. nitens (Forster) 

bellina Casey 

quagga (Germar) 

zebra (t)livier) 

BC 

MB ON 

PQ 

NS 

Genus COS.MOSAJ-IA Casey 

C. chrysocorna (Kirby) 

aureolc,. (Casey) 

auripilis (LeConte) 

densep,'.losa Casey 

YK NT BC 


PE 

NF 

Genus PLEOTRURA Mannerheim 

P. spinicauc.a Mannerheim 

produc.la LeConte 

AK 

Subfamily LAMIINAE 

Tlibe Parmenini 

BC 

Genus MO]TIEILEMA Say 

Collapt:eryr Newman 

M ono p,'.e sa Motschulsky 

Omosc"ilon J. Thomson 

Tlibe Moneileminr 

M. annulatrLm Say AB SK 

angulatumPsota 

demissutn Casey 

hybridum Psota 

montanum Psota 

nubeculum Casey 

puncticolle Psota 

Genus HEBESTOLA Haldeman 

Cacoplt.a LeConte 

Ttibe Monochamini 

H. nebulosa Haldeman 

albata (Knu]l) 

pullato (Haldeman) 

pruinosa (LeConte) 

Genus MONOCHAMUS Dejean 

M. carolinensis (Olivier) 

dentator (Fabricius) 

minor (LeConte) 

M. clamator' latus Casey 

M. marmorirtor Kirby 

acutus (Lacordaire) 

fautor (.LeConte) 

M. mutator LeConte 

M. notatus lDrury) 

confusor l{trby 

morgani Hopping 

M. obtusus rbtusus Casey 

BC 

NT 

NT BC 

_BC 

_SKMB 

AB SK MB 

PQ 

oN PQ 

NB 


oN PQ 

oN PQ 

NB 

NB NS PE 

294

Cerambycidae 

M. scutellatus scutellatus (Say) 

resutor Kirby 

M. scutellatus oregonensis (LeConte) 


M. titillator (Fabricius) 

angusticollis Casey 

obesus Casey 

AK YK NT 

BC AB 

BC AB 

SK 


-oN 

PE LB NF 

Genus MICROGOES Casey 

M. oculatus (LeConte) 

tenuicornis Casey 

-ONPQ 

Genus GOES LeConte 

G. debilis LeConte 

G. pulcher (Haldeman) 

fuscatus IKntTl 

G. pulverulentus (Haldeman) 

laurenticus Casey 

ON PQ 

oN PQ 

ON PQ 

Genus SYNAPHAXTA J. Thomson 

Tribe Mesosini 

S. guexi (LeConte) 

annulata Casey 

- BC 


hutneralis Casey 

Tlibe Dorcaschematini 

Genus DORCASCHEMA Haldeman 

Hetoemis Haldeman 

D. nigrum (Say) 

oN PQ 

TYibe Apodasyini 

Genus EUPOGONIUS LeConte 

Eriopsilus Bates 

Phidola J. Thomson 

E. pauper LeConte 

frafiniKnull 

uestitus (Say) 

E. subarmatus (LeConte) 

E. tomentosus (Haldeman) 

piniuora Filc}r 

Genus PSENOCERUS LeConte 

Paralarica Breuning 

P. supernotatus (Say) 

funebris Casey 

picfzs (Breuning) 

pini LeConte 

frislis Casey 

_ONPQ 

_ONPQ 

_ONPQ 

MB ON PQ NB 

Genus POLIAENUS Bates 

Tlibe Pogonocherini 

P. oregonus (Leconte) - BC 

Genus ECYRUS LeOonte 

Oebaceres J. Thomson 

E. dasycerus dasycerus (Say) - ON 

exiguus (Haldeman) 

obscurus (Haldeman) 

PQ 

295

Cerambycirlae 

Genus LOI'HOPOGONIUS Linsley 

L. crinitus LeConter - BC 

Genus POCiONOCHERUS Dejean 

(Subgenus .POGONOCHERUS s. str.) 

P. penicillalus Leconte 

alashanus Schaeffer 

AK - NT BC AB - MB oN pe NB NS pE - NF 

carinatus Casey 

(Subgenus .OUPOGONOCHERUS Linsley) 

P. mixtus l:laldeman - yK NT BC AB - MB ON pe NB 

simple.x LeConte 

P. parvulus LeConte AB SK MB ON pe 

salicol'r Casey 

P. pictus Fi,ll NT BC AB 

emarginatus Casey 

fastigiutus Casey 

P. propinquus Fall - BC 

Genus ONCTIDERES Lepeletier & Audinet-Serville 

Tribe Onciderini 

O. cingulatrrs cingulatus rsayr 

pallesczns Casey 

- ON 

praecit:.tens Casey 

Genus HIPI)OPSIS Lepeletier & Audinet-serville 

Tribe Hippopsini 

H. lemniscata tFabriciusr 

lineata Gt6rin-M6neville 

- ON 

lineola la Audinet-Serville 

meinerli Aurivillius 

Tribe Acanthoderini 

Genus AXGIMORPHUS Haldeman 

Aethioltoctines J. Thomson 

A. modestus {Gyllenhal } - - MB ON pe 

decipie:zs Haldeman 

A. morrisi (lJhler) 

leucogenus (J. Thomson) 

A. quadrigit,bus tSayr - ON 

lucidus (Knull) 

Genus OPLOSIA Mulsant 

O. nubila (LeContet 

- ON pe 

Genus CANIfNURA Casey 

Tribe Acanthocinini 

Key to North American species: Dillon (1956) 

C. princeps Walkerr - BC 

uincta (Casey) 

Genus NEACANTHOCINUS Dilion 

N. obliquus,rbliquus (LeConter AB 

acornanLls (Casey) 

angulosus (Casey) 

sedulus (Casey) 

N. obliquus racificus (Casey) - BC 

N. obsoletus (Olivier) 

- ON pe 

floridar.:us (Casey) 

N. pusillus (Kirby) AK - AB SK MB ON pe NB NS 

296

Cerambycidae 

Genus CERATOGRAPHIS Gahan 

C. biguttata (LeConte) PQ 

Genus GRAPHISURUS Kirbv 

Urographis Horn 

G. despectus (LeConte) - ON PQ 

hebes (Casey) 

G. fasciatus (DeGeer) - ON PQ 

mirlus (Fabricius) 

p enn sy lu anicus ( Gmelin) 

reductus (Casey) 

Genus AMNISCUS Haldeman 

Astylopsis Casey 

A. collaris Haldeman - ON PQ 

intenuptus Haldeman 

A. macula (Say) - ON PQ NB NS 

sticticus Haldeman 

A. sexguttatus (Say) MB ON PQ 

commixtus Haldeman 

guttatus (Say) 

Genus LEPTOSTYLUS LeConte 

L. transversus transversus - ON PQ 

(Gyllenhal) 

aculiferus (Say) 

rnarg inell us (Haldeman) 

t ub erculatu s (Frolich) 

Genus STERNIDIUS LeConte 

Key to North American species: Lewis (1986) 

S. alpha(Say) - ON PQ 

cinereus (LeConte) 

coloradensis DilIon 

diuergens (Haldeman) 

lateralis (Haldeman) 

nelsonicus (Casey) 

suturalis Dillon 

uicinus (Haldeman) 

S. fascicularis (Harris) - ON PQ - NS 

dentatus (Casey) 

nigricans Dillon 

obscurellus (Casey) 

pleuralis (Casey) 

scapalis (Casey) 

testaceus (Casey) 

timid,us (Casey) 

xanthoxyli (Shimer) 

s. misellus (Leconte) MB oN PQ 

S. rusticus (LeConte) MB 

Genus ASTYLEIOPUS Dillon 

A. variegatus (Haldeman) AB SK MB ON PQ NB 

obscurus (Haldeman) 

tr ifa sciatu s ( Haldeman) 

Genus DECTES LeConte 

D. sayi Dillon & Dillon AB - MB ON 

spizosus (Say) 

Genus LEPTURGES Bates 

(Subgenus LEPTURGES s. str.) 

L. confluens (Haldeman) - ON PQ 

L. symmetricus (Haldeman) - ON PQ 

297

Cerambycidae 

Genus URGLEPTES Dillon 

U. facetus (liay) 

U. querci (Fitch) 

/risfis (Casey) 

U. signatus (LeConte) 

tenebrosus (Casey) 

-ONPQ 

_ONPQ 

-oNPQ 

Genus HYPERPLATYS Haldeman 

H. aspersa (Say) 

uigilans Casey 

H. maculata Haldeman 

amnicola Casey 

binocul.aris Casey 

cryptiat Casey 

delicatr:.t Casey 

frigida Casey 

lentigit;.osa Casey 

nigrellcr Haldeman 

robustula Casey 

uariolata Casey 

H. montana Casey 

BC 

BC 

AB 

AB 


_MBONPQNBNS 

SK MB 

NF 

Genus CYR|CINUS LeConte 

Myrmo,bmia Bates 

C. pygmaeu,r (Haldeman) 

Tlibe Cyrtinini 

ON 

Tlibe Saperdini 

Genus SAPI)RDA Fabricius 

Eutetropha Bates 

Key to species: Abdullah and Abdullah ( 1966) 

S. calcarata Say 

adspersa LeConte 

S. candida brpuncLata Hopping 

S. candida candida Fabricius 

biuittato Say 

S. concolor unicolor Felt & Joutel 

mecasotdes Casey 

S. cretata Nr)wman 

S. discoidea Fabricius 

dorsalit Haldeman 

fuscipes Say 

S. fayi Blan< 

S. horni Jou;el 

S. imitans F.:lt & Joutel 

S. lateralis F'abricius 

abbreuiata Fitch 

connecte. Felt & Joutel 

disconolata Pic 

suturalis Fitch 

transeuns Breuning 

S. mutica Say 

S. obliqua Say 

S. populnea :moesta LeConte 

S. populnea'populnea Linn6 

S. puncticoll s Say 

trigeminata Randall 

S. tridentata Olivier 

dubiosa Haldeman 

interme':lia Fitch 

rubronotata Fitc}r 

trifasciota Casey 

S. vestita Say 

atkinsoni Crrtis 

YK NT SC 

BC 

BC 

SK 

BC AB MB ON PQ 

BC AB 

SK 

SK 

AB SK 

AB 

MB ON 

-oNPQ 

MB ON PQ NB 

MB 

_ONPQ 

oN PQ 

_MB 

oN PQ 

-MBONPQ 

SK 

MB ON PQ 

MB ON PQ 

MB ON PQ 

-oNPQ 

MB ON PQ 

MB ON PQ 

NS 

NS 

NB NS 

PE 

NB NS PE 

LB 

NF 

298

Cerambycidae 

Genus OBEREA Mulsant 

Amaurostoma J. Miilier 

Isosceles Newman 

Key to species: Breuning (1960) 

O. appaiachiana Casey 

O. bimaculata (Olivier) 

affinis Haris 

basalis LeConte 

basicollis Breuning 

bicallosicollis Breuning 

bipunctata (Creutzer) 

broo&si Wallis 

delicatula Casey 

dolosa Casey 

filum Casey 

flauocephala Blatchley 

insignior Breuning 

insignis Casey 


rnontana Casey 

per s picill ata Haldeman 

tr ip unctata ( Fabricius ) 

ulmicola Chittenden 

umbra Casey 

O. canadensis Fisher 

O. cylindricollis Casey 

O. deficiens Casey 

O. delongi Knull 

anter ufi collis Breuning 

O. exilis Casey 

O. ocellata Haldeman 

plagiata Casey 

O. oculaticollis (Say) 

O. pallida Casey 

O. praelonga Casey 

O. pruinosa Casey 

O. quadricallosa LeConte 

O. schaumii LeOonte 

caseyi Plavilstshikov 

ferruginea Casey 

infrarufa Breuning 

su bcy I ind r i collis Breuni ng 

uittipennis Breuning 

wapleri Chevrolat 

O. tripunctata (Swederus) 

amabilis Haldeman 

intermedia Casey 

mandarina (Fabricius) 

rnyops Haldeman 

subdeficiens Breuning 

subexilis Breuning 

submondarina Breuning 

texana Horn 

Genus MECAS LeConte 

(Subgenus MECAS s. str.) 

M. bicallosa Martin 

Genus TETRAOPES Schiinherr 

Key to species: Chemsak (1963) 

T. annulatus LeConte 

canescens LeConte 

fontinalis Casey 

uteanus Casey 

uestitus Casey 

Tribe Phytoecini 

_BC 

-BC 

BC 

_SKMBONPQ 

AB_ 

AB SK 

Tribe Tetraopini 

AB-MB 

_ON-NB 

MB ON PQ 

-oN 

-ONPQ 

-oNPQ 

-oN 

-oN 

MB ON 

MB ON PQ 

_ONPQ 

-oN 

MB ON PQ 

oN PQ 

NB 

NB 

299

Bruchidae 

T. femoratus LeConte 

amnicc'la Casey 

atrisetosus Casey 

breuisetosus Casey 

coseyj ,\urivillius 

collaris Horn 

fortis Casey 


nigripes Casey 

oregonetnsis LeConte 

Punctiltennis Qasst 

robustt,s Casey 

ruber (.Asey 

spissicornls Casey 

uegaser,,sis Casey 

uelutin ^ts Casey 

T. quinquemaculatus Haldeman 

T. tetrophthalmus (Forster) 

humerc:Iis Casey 


torndtor (Fabricius) 

t r e dec im punc tatu s (Dr apiez) 


ON 

-MBONPQ 

Family BRUCHIDAE seed beetles 

J. McNamara 

The t ruchids comprise 19 species in Canada 

and Alasra. Their habits vary considerably, but 

they depend almost entirely on seeds for their 

development. The larvae live in seeds of legumes, 

notably l)eas and beans, and are often pests of 

stored leguminous seeds. The adults are commonly 

found in flowers and on foliage. The pea weevil 

(Bruchus pisorum) and the bean weevil 

(Acanthoscelides obtectus) are major pests of peas 

and beans, and have spread around the world. 

Borowiec (1987) published a key to the genera of 

the world, and Udayagiri and Raj Wadhi (1989) 

published a catalog of the world species. 

aK (1);YK (1); NT (2); BC (10); AB (8); SK (7); MB 

(e); ON (11); PQ (5); NB (2); NS (2); PE (1); NF (1) 

. I(6) 

Genus AMB LYCERUS Thunbere 

A. robiniae ( Fabricius) 

gleditsiae (Castiglioni) 

Subfamily AMBLYCERINAE 

AB 

Subfamily KYTORHININAE 

Genus KYTORHINUS Fischer von Waldheim 

K. prolixus (Fall) 


Genus MEG.A.CERUS Fahraeus 

Pachybruchus Pic 

(Subgenus NTEGACERUS s. str.) 

M. discoidus (Say) 

Genus CALI,OSOBRUCHUS Pic 

C. maculatus (Fabricius) 

ambiguzs (Gyllenhal) 

quadrinzaculatus (Fabricius) 

sinuatus (Fahraeus) 

Subfamily BRUCHINAE 

BC AB 

SK MB ON 

SK ON 

PQ NB NS 

300

Chrysomelidae 

Genus GIBBOBRUCHUS Pic 

Key to North American species: Whitehead and Kingsolver (1975) 

G. mimus (Sayt 

borealis (Schdnherr) 

crataegi (Fahraeus) 

Genus MEIBOMEUS Bridwell 

Key to North American species: Kingsolver and Whitehead (1976) 

M. musculus tSay) 

a lb og ut tis (Motschulsky) 

erythrocerus (Riley) 

ON 

ON 

PQ 

Genus ACANTHOSCELIDES Schilskv 

A. aureolus (Horn) 

A. calvus (Horn) 

A. fraterculus (Horn) 

A. obtectus (Say)t 

fabae (Pi'iley) 

u ar icor ni s (Motschulsky) 

A. pauperculus (LeConte) 

p ulicar iu s (Motschulsky) 

rulzs (Motschulsky) 

simplex (Motschulsky) 

A. perforatus (Horn) 

A. pullus (FalI) 

A. seminuium (Horn) 

_BC 

NT BC 

_BC 

BC 

BC 

AB SK MB 

-oN 

AB SK MB ON 

MB ON 

AB 

_MB 

AB SK MB 

PQ 

Genus BRUCHUS Linn6 

B. brachialis Fahraeust 

B. pisorum (Linn6)t 

pisi Linn6 

B. rufimanus (Boheman)t 

_BC 

_BCAB 

SK 

-oN 

MB ON 

MB ON 

PQ 

PQ 

NU NS 

PE 

NF 

Genus BRUCHIDIUS Schilsky 


B. cisti (Fabricius)t 

unicolor (Olivier) 

B. villosus (Fabricius)f 

oler (Marsham) 

Bottimer (1968) 

BC 

ON 

Family CHRYSOMELIDAE leaf beetles 

L. LeSage 

The family Chrysomelidae is one of the largest 

group of beetles in the world, with over 50 000 

known species and many additional thousands to be 

discovered, especially from Asia and the Neotropical 

region; 569 species and subspecies are listed here for 

Canada and Alaska. All known adults are phytophagous. 

Primitive plants and ferns are used 

infrequently, and conifers are barely touched; deciduous 

trees, bushes, and weeds are preferred hostplants. 

Some species are pollen eaters when in the 

adult stage. Larvae show a broader ecological diversity: 

Donaciinae are aquatic; Clytrinae live in ant 

nests, where they feed on plant detritus and on ant 

larvae or pupae; most Cryptocephalinae feed on 

decaying matter in leaf litter; Criocerinae, 

Chlamisinae, Chrysomelinae, and Cassidinae and 

some Galerucinae and Alticinae feed on leaves: those 

in remaining subfamilies are either root feeders, root 

miners, stem miners, or leaf miners. 

Chrysomelidae is usually classifred into 19 subfamilies 

(Seeno and Wilcox 1982) of which 14 occur 

in Canada. The fauna of the leaf beetles is relatively 

well known in eastern Canada (except for some 

alticine, galerucine, and cryptocephaline genera) but 

is poorly known in western Canada. The present list 

follows essentially the arrangement proposed by 

Seeno and Wilcox (1982), except where the generic 

concepts have been recently modifred. 

AJ{ (42); YK (41); NT (72); BC (181); AB (217); SK 

(174); MB (25a); ON (363); PQ (27a); NB (116); NS 

(103);PE (19); LB (1a); NF (3a) o H (9); I (38) 

301

Chrysome lidae 

Genus PL,A.TEUMARIS C.G. Thomson 

P. aurifera (LeConte) 

P. balli Askevold 

P. chalcea (Lacordaire) 

lodingi .Schaeffer) 

s hoema,ize ri ( Schaeffer) 

P. diversa (Schaeffer) 

P. dubia (Schaeffer) 

idola (Hatch) 

P. flavipes (Kirby) 

wallil;i (Schaeffer') 

P. frosti (Schaeffer) 

P. fulvipes (Lacordaire) 

P. germari (Mannerheim) 

diues (LeConte) 

fl au ip ennis ( Mannerheim) 

serricauda (Schaeffer) 

P. metallir,a (Ahrens) 

catar'rctae (Newman) 

femor alis (Kirby) 

gentitis (Leconte) 

indicu (Melsheimer) 

zaza (Melsheimer) 

parua: (Lacord.afte) 

P. nitida (tlermar) 

emarSlinata (Kirby) 

juncina (Couper) 

paciftca (Schaeffer) 

P. neomex icana (Schaeffer) 

long it:' ol I i s ( Schaeffer) 

u er mti. c ulat a ( Schaeffer) 

P. pusilla Say) 

pyritt: sa (LeConte) 

rugiftons (Newman) 

P. robusta (Schaeffer) 

P. rufa (Say) 

affinis (I\trby) 

jucunda (LeConte) 

hirbyi (Lacordaite) 

s ulcic ollis (Lacordaire) 

Subfamily DONACIINAE 

Key to New World genera: Askevold (1990) 

Tlibe Plateumarini 

AK YK NT 

_BC 

AK--nCAB-yuoN- 

AB_MB 

AB SK MB 

PQ 

PQ 

PQ 


AK_NTBCABSKMBONPQNBNSNF 


ON 


AX 

YK 

NT 

BC 

BC 

BC 

_ y:'TSKyu 

NB 

NB NS 

PE 


ff 

ON 

PQ 

SK MB ON PQ 

oN PQ 

oN PQ 

NB 

NB 

NS 

NB NS 

NB NS 

NF 

LB NF 

LB NF 

LB NF 

LB NF 

Genus DO NACIELLA Reitter 

Tribe Donaciini 

D. pubicollis ( Suffrian) 

Genus DONACIA Fabricius 

(Subgenus DONACIA s. str.) 

Key to l\lorth American species: Marx (1957) 

AB 

SK MB ON 

D. cinctico:rnis Newman 

antillarum Suffrian 

lucidl:, Lacordaite 

pulch,zlla LeConte 

rufipennis Lacordaire 

tenuis Schaeffer 

tryphera Schaeffer 

D. palmata Olivier 

angustipes Marx 

claud;icans Germar 

D. parvide.ns Schaeffer 

D. piscatrir Lacordaire 

alutacea LeOonte 

carolina Lacordaire 

conge;'Ler LeConte 

cuprea Kirby 

MB ON PQ 

ON PQ 

-oN 

MB ON PQ 

NB NS 

NB 

NS 

302

Chrysomelidae 

D. proxima Kirby 

californica LeConte 

e pisco palis Lacordaire 

D. rufescens Lacordaire 

D. texana Crotch 

minor Schaeffer 

NT BC 

AB 


ON PQ 

ON 

NF 

(Subgenus DONACIOMIMA Medvedev) 

D. biimpressa Melsheimer 

a ur ichalcea Melsheimer 

torosa Le0onte 

D. caerulea Olivier 

aequalis Say 

confusa Lacordaire 

confusa LeConte 

D. canadensis Askevold 

D. cazieri Marx 

D. confluenta Say 

D. cuprea Kirby 

curticollis Knab 

D. distincta LeConte 

occidentalis Mead 

D. fulgens LeConte 

D. hirticollis Kirby 

rudicollis Lacordaire 

D. limonia Schaeffer 

D. magnifrca LeConte 

D. megacornis Blatchley 

megaloceraWeise 

D. porosicollis Lacordaire 

D. pubescens LeConte 

D. rugosa LeConte 

D. subtilis Kunze 

aenea Ahtens 

aerea Lacord.aire 

magistrigata Mead 

quadricollis Say 

D. tuberculifrons Schaeffer 

AK 

YK 

AK YK 

NT BC 

_BC 

NT 

NT 

BC 

NT BC 

AB SK 

AB 

AB SK 

-SK 

AB SK 

_SK 

AB SK 

BC AB SK 

BC AB SK 

BCTSK 

BC AB SK 

MB ON PQ 

MB ON PQ NB 

MB ON PQ 

_ONPQ 

MB ON PQ 

-oNPQ 

MB ON PQ 

MB ON PQ NB 

MB ON PQ NB 

_ONPQ 

MB ON PQ NB 

-oN 

MB ON 

MB ON PQ 

-oN 

MB ON PQ 

BC MB ON PQ NB 

LB 

LB 

NS PE 

NS_LBNF 

NS 

NF 

Tribe Haemoniini 

Genus NEOFIAEMONIA Sz6kessy 

Key to North American species: Askevold (1988) 

N. flagellata Askevold 

N. melsheimeri (Lacordaire) 

N. minnesotensis Askevold 

N. nigricornis tKirby) 

americana (Gu6rin-M6neville) 

BC 

YSK 

MB ON 

MB ON PQ 

MB ON 

MB ON PQ 

Genus ORSODACNE Latreille 

Orsodacna Dejean 

O. atra (Ahrens) 

anneniacae Germar 

childreniKirby 

hepatica Say 

inconstans Newman 

luctuosa Lacordaire 

ruficollis Newman 

tibialis Iltrby 

tricolor Melsheimer 

triuittata Lacordaire 

uittata Say 

Subfamily ORSODACNINAE 

YK NT BC AB SK MB ON PQ NB 

NS 

303

Chrysomelidae 

Genus ZEIIGOPHORA Kunze 

(Subgenus ZEUGOPHORA s. str.) 

(Auchenia Thunberg) 

(Tararis LeConte) 

Z. abnormis (LeConte) 

hirbyi Baly 

reineckei Gtote 

Z. atraFall 

Z. consanguinea Crotch 

Z. multnomah Hatch 

Z. puberula Crotch 

Z. scutellaris Suffriant 

Z. varians Crotch 

Subfamily ZEUGOPHORINAE 

AK YK NT BC 

YK NT 

YK 

-Nr 

SK MB ON PQ 

BCABSKMB_PQ 

BC AB SK 

BC_ON 

ON 



Genus LIL IOCERIS Reitter 

(Subgenus LILIOCERIS s. str.) 

L. lilii (Scopoli)t 

lilioru,m (C.G. Thomson) 

melanocephala (Say) 

m e r di,;1 e r a (Fabricius ) 

Subfamily CRIOCERINAE 

Tlibe Criocerini 

oN PQ 

Genus CRIOCERIS O.F. Mriller 

Pleurcphora Chewolat 

C. asparagr (Linn6)t 

C. duodecirnpunctata (Linn6)f 

_BCABSKMBONPQ 

_BCABSKMBONPQ 

NB 

NB 

Genus LEIIA Fabricius 

(Subgenus LEMA s. str.) 

L. cyanella Linn6f 

L. gaspensis W.J. Brown 

L. palustris Blatchley 

L. trilinea trilinea R.E. White 

bilineo.ta Omer-Couper & Miles 

medionata Schaeffer 

medionota Schaeffer 

trilineuta atct. 

L. trivittatr,L trivittata Say 

imma< ulicollis Chevrolat 

Iecontti Clatk 

nigrou.ittata Schaeffer 

triuirgzta LeConte 

Genus OUI.,EMA des Gozis 

(Subgenus I)ULEMA s. str.) 

O. melanopus (Linn6)t 

Tribe Lemiini 

PQ 

_ONPQ 

MB ON PQ 

ON PQ 

ON PQ 

Genus ANC,TMOEA Lacordaire 

Anomoia Dejean 

Euthyna Gistel 

A. laticlavia laticlavia (Forster) 

ephipptum (Germar) 

lloridana Schaeffer 

Subfamily CLYTRINAE 

Key to New World genera: Moldenke (1981) 

Key to North American species: Moldenke (1970) 

TYibe Clytrini 

MB ON 

PQ 

304

Chrysomeiidae 

hansana Schaeffer 

nigripes (Fabricius) 

o6silo (Fabricius) 

ruficauda (Foersberg) 

Tribe Megalostomini 

Genus COLEOTHORPA Moldenke 

C. dominicana dominicana 

(Fabricius) 

C. vittigera vittigera (LeConte) 

NT_AB_MBON 

_BCABSKMB 

Tribe Babiini 

Genus BABIA Chevrolat 

Harpasta Gistel 

(Subgenus BABIA s. str.) 

B. quadriguttata quadriguttata 

(Olivier) 

biguttata Crotch 

quadriforis (Newman) 

Genus SAXINIS Lacordaire 

(Subgenus BOREOSAXINIS Moldenke) 

S. omogera omogera Lacordaire 

S. saucia saucia LeOonte 

ec 

Subfamily CRYPTOC EPHALINAE 

Tlibe Pachybrachini 

ON 

ON 

Genus PACHYBRACHIS Chevrolat 

(Subgenus PACH\aBRACHIS s. str.) 

(Pachystylus Rey) 

P. abdominalis (Say) 

P. alticola Fall 

P. atomarius (Melsheimer) 

atomus Bowditch 

infaustus Haldeman 

P. autolycus FaIl 

P. bivittatus (Say) 

albescens Suffrian 

uiduatus Suffrian 

P. calcaratus Fall 

P. coloradensis Bowditch 

seuier Bowditch 

P. discoideus Bowditch 

P. donneri Crotch 

P. hepaticus (Melsheimer) 

punctdtus Haldeman 

P. luridus (Fabricius) 

oesculi (Melsheimer) 

femoratus (Say) 

festiuus Fall 

moerens Stil, 

nigrinus Blatchley 

P. melanostictus Suffrian 

P. minor Bowditch 

P. mnigrum (Melsheimer) 

intricatus Suffrian 

P. nigricornis carbonarius 

(Haldeman) 

P. nigricornis diffrcilis Fall 

wahsatchensis Fall 

P. nigricornis nigricornis (Say) 

janus Fall 

AB SK MB 

AB 

AB-MBON 

-BCABSKPQ 

-BCABSK-ONPQ 

---AB--oN-- 

_AB 

BC AB 

-ABSKMBONPQ 

ON PQ 

_YKNTBCABSKMB 

- - - :c - - 

AB_MBON 

_BCABSKMBONPQ 

AB SK MB 

MB- pe - - - - - 

305

Chrysomelidae 

P. obsoletr.,s Suffrian 

P. othonous othonous (Say) 

rna rg r. naticoll is Randall 

P. othonous pallidipennis Suffrian 

striatus LeConte 

P. othonous sioux Balsbaugh 

P. peccans Suffrian 

P. pectoralis (Melsheimer) 

oculatus Suffrian 

sobrin.us Haideman 

stricticus Blatchley 

P. praeclarus Weise 

elegar,.s Blatchley 

P. relictus Fall 

P. spumar:us Suffrian 

robori:s Fall 

P. subfasci atus LeConte 

bigutiatus Suffrian 

P. tridens tMelsheimer) 

P. trinotatrrs (Melsheimer) 

P. vau Fal] 

P. vacillatus Fall 

-__AB 

BC 

--sx 

BC 

AB 

AB 

AB 

_MBONPQ 

_MBONPQ 

SK 

SK 

MB 

MB ON PQ 

_ONPQ 

MB ON 

-oN 

MB ON 

MB ON 

PQ 

NB 

NB 

NS--- 

NS 

Tribe Monachini 

Genus LEIIIPHANES Gistel 

Mona,:hulus Leng 

Mona:hus Chevrolat 

Key to l\'orth American species: Balsbaugh (1g66) 

L. saponat rs ( Fabricius t 

oler (IIaldeman) 

uiridanus (Falll 

MB ON PQ 

NB 

Genus CRIIFTOCEPHAIUS O.F. Mtiller 

(Subgenus CRYIvIOCEPHAJ-US s. str. ) 

(C antiiostethus Haldeman) 


C. calidus Suffrian 

C. confluer:.tus confluentus Say 

C. gibbicollis gibbicollis Haldeman 

C. guttulatus Olivier 

loufur Newman 

C. incertus Olivier 

lineoltfius Haldeman 

C. insertus Haldeman 

ellipscidalis Casey 

C. leucomelas leucomelas Suffrian 

C. mutabil: s Melsheimer 

discoideus Suffrian 

dispersus Haldeman 

C. notatus notatus Fabricius 

C. notatus quadrimaculatus Say 

C. notatus sellatus Schaeffer 

C. quadruplex Newman 

q uadr tg uttulus Suffrian 

C. sanguin collis Suffrian 

nigerrimus Crotch 

C. schreibersii Suffrian 

rugicallis Haldeman 

C. spurcus vandykei R.E. White 

C. striatulrLs LeConte 

C. tinctus LeConte 

C. trivittatrs Olivier 

lirus Newman 

uilolus Haldeman 

C. venustus cinctipennis Randall 

Tlibe Cryptocephalini 

_ _ Bcy 

_MBON 

BCY:"-o*pe 

BC 

BC 

AB SK 

_MBON 

-oN 

MB ON 

oN PQ 

ON 

ON 

PQ 

PQ 

PQ 

ON PQ NB 

SK MB ON PQ 

_MB 

_MBONPQ 

_PQ 

_PQ 

_PQ 

SK MB ON 

NB NS 

NS 

NS 

306

Chrvsomelidae 

C. venustus hamatus Melsheimer 

C. venustus ornatulus Clavareau 

ornatus Fabricius 

C. venustus simplex Haldeman 

C. venustus venustus Fabricius 

flaccidus Suffrian 

MB 

MB 

AB SX nlS 

9" 

ON 

ON 

PQ 

Genus DIACHUS LeConte 

Fabricianus Weise 

D. aeruginosus LeConte 

D. auratus (Fabricius) 

aeneus (OIiier) 

chalconotu s (Mannerheim) 

urrldis (Melsheimer) 

D. catarius (Suffrian) 

D. pallidicornis (Suffrian) 

NT BC 

-BC 

BC 

l.BsKMB 

MB 

MB ON 

pe 

ON PQ 

ON 

NS 

NBI'- 

Genus BASSAREUS Haldeman 

B. formosus (Melsheimer) 

egenus Suffrian 

guttatus Haldeman 

B. lituratus (Fabricius) 

B. Iuteipennis (Melsheimer) 

B. mammifer (Newman) 

B. pretiosus (Melsheimer) 

B. sellatus Suffrian 

(?) notatus (Kirby) 

(?) q uadris ig nolus Suffrian 

B. sulphuripennis (Melsheimer) 

-oN 

-oN 

-oN 

-oN 

MB 

MB ON 

ON 

PQ 

PQ_ 

PQ 

NS 

Genus TRIACHUS LeConte 

T. atomus (Suffrian) 

T. vacuus LeConte 

-oN 

-oN 

Ne- 

Genus NEOCHLAMISUS Karren 

Key to North American species: Karren (1972) 

N. alni (W.J. Brown) 

N. bebbianae (W.J. Brown) 

N. chamaedaphnes (W.J. Brown) 

N. comptoniae (W.J. Brown) 

N. coryli (W.J. Brown) 

N. cribripennis (LeConte) 

N. eubati (W.J. Brown) 

N. fragariae (W.J. Brown) 

N. tectus (W.J. Brown) 

Genus EXEMA Lacordaire 

Key to North American species: Karren (1966) 

E. canadensis Pierce 

E. dispar Lacordaire 

pennsyluanica Pierce 

Subfamily CHLAMISINAE 

-oN 

MB ON PQ 

-ONPQ 

-oN 

_ONPQ 

_ONPQ 

-oN 

MB ON 

-oN 

PQ 

NB 

NB 

NB 

lls 

NS 

NS 

Genus SYNETA Dejean 

Key to species: Edwards (1953) 

Key to eastern North American species: 

S. albida LeConte 

suturalis LeConte 

S. carinata Mannerheim 

S. extorris borealis W.J. Brown 

S. ferruginea (Germar) 

cosfolo (Newman) 

rubicunda Lacordaire 

tripla (Say) 

Brown (1961) 

AK 

Subfamily SYNETINAE 

BC 

BCT 

--oNpeNnNs-- 

_MBONPQNBNS 

NF 

NF 

307

Chrysomel:idae 

S. hamata Horn - BC AB SK 

S. pilosaW.J. Brown - YK NT BC AB SK MB ON PQ NB NS NF 

S. simplex simplex LeConte - BC 

S. simplex subalpina Edwards - BC AB 

Genus METACHROMA Chevrolat 

Atracli.ela Gistel 

Key to syrecies: Blake (1970) 

Subfamilv ELIMOLPINAE 

Tri6e Nodinini 

M. angustr:lum Crotch AB SK MB ON 

parall,zlum}J.orn 

M. longulun Horn AB SK 

no u e m s triat um Schaeffer 

opacipenne Fall 

Genus PAF:IA LeOonte 

Key to North American species: Wilcox (1957); Balsbaugh (1970) 

P. aterrima tOliviert PQ 

P. fragaria,r fragariae Wilcox - ON PQ 

P. pratensi s Balsbaugh PQ 

P. quadrigr.ttata LeConte AB SK MB 

salicetLWllcox 

P. quadrin< tata (Say) 

giluipts Crotch 

- ON PQ 

infusafta (LeConte) 

melan.t ra (Melsheimer) 

P. sexnoLat r tSayt MB 

P. scutellaris (Notmanr - ON 

P. thoraci*L (Melsheimert MB ON PQ 

Genus SPIITITHEROPTryTA Dei ean 

Chalc

Chrysomelidae 

R. deceptor Barber 

R. praetextus (Say) 

spiculatus Barber 

ueisei (Schaeffer) 

AB 

MB ON 

-oNPQ 

PE 

Genus CHRYSOCHUS Chevrolat 

Atymius Gistel 

Key to western North American species: Hatch (1971) 

C. auratus (Fabricius) 

C. cobaltinus LeConte 

californicus Marshall 

castaneus Marshall 

tenebricosus Marshall 

_BCABSK 

_BC 

MB ON PQ 

Tribe Adoxini 

Genus GRAPHOPS LeConte 

Heteraspis LeConte 

Key to species: Blake (1955) 

G. cupraeus ( Provancher) 

G. curtipennis curtipennis 

(Melsheimer) 

G. marcassitus (Crotch) 

G. nebulosus (LeConte) 

G. negrellus Blake 

G. nigellus Blake 

G. pubescens (Melsheimer) 

G. varians LeConte 

G. viridis Blake 

_AB 

. - _ TSK 

SK 

PQ 

-oN 

MB ON PQ 

MB 

MB 

MB ON PQ 

yu_ _ 

NS 

Genus XANTHONIA Baly 

X. decemnotata (Sayt 

X. vil losula (Melsheimer) 

plagiata (Melsheimer) 

steuensi Blake 

Genus FIDIA Walsh 

F. viticida Walsh 

lurida Lefevre 

Genus BROMIUS Chevrolat 

Adoxus Kirby 

Eumolpus Redtenbacher 

B. obscurus (Linn6)* AK 

cochlearius Say 

concinnus Weise 

leaisi Weise 

SKMBONPQ-NS 

_MBONPQ_NS 

-oN 

YK NT BC AB SK MB ON PQ NB NS PE NF 

Genus MYOCHROUS Erichson 

Key to North American species: Blake (1950) 

M. denticollis tSayt 

M. squamosus LeConte 

AB 

SK 

PQ 

Genus GLYPTOSCELIS Chevrolat 


G. albida albida LeConte - 

G. artemisiae Blake 

G. longior LeConte 

G. pubescens (Fabriciust 

pini (Say) 

hirsuta (Gmelin) 

hirta (Olivier) 

G. septentrionalis Blake 

_-Bc_- 

BC 

BC 

BC 

MB ON PQ 

309

Chrysomelirlae 

Genus TIMARCHA Latreille 

(Subgenus r\MERICANOTIMARCHA Jolivet) 

Key to New World species: Jotivet (1948) 

T. cerdo StAl 

aenea Jtolivet 

intento Jolivet 

nigra Jolivet 

T. intricata Haldeman 

adusta Jolivet 

uiolacet Jolivet 

uiridis Jollet 

Genus ZYGDGRAMMA Chewolat 

Subfamily CHRYSOMELINAE 

Key to North American species: Wilcox (1972) 

T?ibe Timarchini 

_BC 

_BC 

Tribe Chrysomelini 

Z. conjuncta conjuncta (Rogers) AB SK MB 

Z. continua LeConter AB 

Z. disrupta'Rogersr - ON 

Z. exclamationis (Fabricius) AB SK MB ON pe 

Z. suturalis casta ( Rogers) MB 

fesliuo .Fabricius) 

Z. suturalis suturalis tFabriciusr - ON pe 

pzlcro ,Fabricius) 

Genus CALI,IGRAPHA Chevrolat 

B oliog r tp ha Motschulsky 

M eta llc; g r ap ha Motschul sky 

Phyllis Gistel 

Polyspita Hope 

Key to North American species: Brown (1g45) 

(Subgenus C|.RAPHICAI-LO Monros) 

C. lunata hlbrida tSayr AB SK MB 

lateuittuta (Achard) 

C. lunata lunata (Fabricius) 

brunnea, (Thunberg) 

bowditchi (Achard) 

mediori:tpta (Achard) 

(Subgenus I IDENSOMELA Monros) 

- ON 

C. bidenticola bidenticola 

W.J. Brown 

- ON PQ 

hilaris rAchard) 

intermedia (Achard) 

similis (Rogers) 

(Subgenus COREOPSOMELA Monros) 

C. californica californica Linell - BC 

C. californica coreopsivora NT BC AB SK MB ON PQ NB - 

pE 

W.J. Brown 

elegans (Olivier) 

(Subgenus ACALLIGRAPHA Monros) 

C. praecelsis (Rogers) MB 

(Subgenus CAILIGRAPHA s. str.) 

C. alni Schaeffer - - AB SK MB ON Pe NB NS 

confinis (Kirby) 

C. alnicola V/.J. Brown - ON PQ NB NS 

C. amator S'.J. Brown - ON 

C. apicalis Notman - ON PQ 

C. confluens Schaeffer - ON PQ NB NS 

C. dolosa W.J. Brown - ON PQ 

310

Chrysomelidae 

C. igrrota W.J. Brown 

C. incisa (Rogers) 

C. knabi W.J. Brown 

C. multipunctata bigsbyana (Kirby) 

C. multipunctata multipunctata 

(SaY) 

C. ostryae W.J. Brown 

C. philadelphica (Linn6) 

decipiens (Weber) 

ruficornis (Olivier) 

C. pnirsa (StAl) 

labyrinthica Horn 

C. pruni W.J. Brown 

C. rhoda Knab 

C. rowena Knab 

C. scalaris (LeConte) 

multiguttis Stitl 

C. sigmoidea (LeConte) 

C. spiraeae {Say) 

C. tiliae W.J. Brown 

C. verrucosa (Suffriant 

C. vicina Schaeffer 

C. virginea W.J. Brown 

NT 

nc AB sx 

_AB 

BC_SK 

BC 

BC 

T 

AB SK 

MB ON PQ 

MB 

MB ON PQ 

y" 9* 

PQ 

-oNPQ 

MB ON PQ 

ON 

PQ 

-oN 

MB ON 

MB ON PQ NB 

MB ON PQ 

-oN _ON-NB 

MB ON 

-oNPQNB 

-ONPQNB 

NB NS 

NE NS 

NB NS 

PE 

Genus LEPTINOTARSA SIAI 

Myocoryna Sthl 

Polygratnma Chevrolat 

Key to North American species: Jacques (1988) 

L. decemlineata decemlineata (Say) 

albida Tower 

intermedia Towet 

melanica Tower 

minuta Towet 

multitaeniata (StA\:l 

oblongata "Iower 

pallidaTower 

rubicundaTower 

rubriuittata 'Iower 

tortuosa Tower 

BC 

AB 

SK 


PE 

Genus I"ABIDOMERA Chevrolat 

P arop simena Motschulsky 

L. trimaculata (Fabriciust 

cliuicollis (Kirby) 

rogersi LeConte 

MB ON PQ 

Genus CHRYSOLINA Motschulsky 

Ap te r osoma Motschulsky 

Chrysomela auct. 

OreinaMonros & Bechyn6 

Key to species: Brown (I962a) 

(Subgenus CHRYSOLINA s. str.) 

(S tic hosoma Motschulsky) 

C. staphylea tLinn6)t 

PQ 

NS 

NF 

(Subgenus SPHAEROMELA Bedel) 

C. varians (Schaller)t 

(Subgenus HYPERICIA Bedel) 

C. hyperici (Forster)* 

C. quadrigemina (Suffrian)t 

gemellata atct. 

gemtnata anct. 

(Subgenus CHALCOIDEA Motchulsky) 

C. frnitima W.J. Brown AK 

C. fl avomarginata fl avomarginata 

(Say) 

C. flavomarginata vidua (Rogers) 

YK 

BC 

BC_ON 

BC_ON 

BC 

AB 

AB 

NB 

NB 

NS 

NS 

311

Chrysomelirlae 

C. hudsonica W.J. Brown 

NT 

MB ON PQ 

NF 

(Subgenus I'LEUROSTICA Motschulskv) 

C. cavigera iJ.R. Sahlberg)x AK yK 

lolli Jar:obson 

(Subgenus I,RCTOLINA Kontkanen) 

C. caurina Vy'.J. Brown AK 

C. magniceps (J. Sahlberg) AK 

C. subsuicat.a (Mannerheim) AK yK 

(Subgenus I'EZOCROSITA Jacobson) 

C. auripenn:s (Say) AK 

inornalz (Rogers) 

C. basilaris rSay) - YK 

BC 

AB 

MB 

Genus GASI ]ROPHYSA Chevrolat 

Key to North American species: Jolivet (1951) 

(Subgenus CASTROPIrySA s. str.) 

(Gastro'zidea Hope) 

G. cyanea caesia Rogers 

G. cyanea c;anea Melsheimer 

G. dissimilis (Sayr 

G. lormosa t iay r 

G. polygoni tLinner+ 

caeruli1tennis (.Say) 

r uftcoll, s (Fabricius) 

NT BC AB SK 

N'T_ABSK_ 

AB SK MB 

_BCABSKMB 

ON 

ON PQ 

NB NS 

PE 

Genus PHAINDON Megerle von Miihlfeld 

Key to Nodh American species: Balsbaugh (1983) 

(Subgenus PFIAEDON s. str.) 

P. armoraciae armoraciae (Linn6)* AK YK 

americcnus Schaeffer 

P. armoraciae niger Hatch 

planus Jatch 

P. laevigatus (Duftschmid)* AK 

carri Hr*ch 

cochleariae auct. amer. 

uniform:is Fall 

uancout'erensis Hatch 

P. oviformis lleConte) AK YK 

P. prasinellus (LeConter 

punctat.us Hatch 

uandyke'iHatch 

P. viridis Melsheimer 

aeruginosus Suffrian 

dietrichi Hatch 

micr o re i: ic ula tzs Hatch 

oklahortensis }{ahch 

NT BC 

_BC 

NT BC 

AB 

NT BC AB 

_BCAB 

AB SK 


MB ON 

sK yu oN 

SK 

PQ NB NS 

PQ:tuNS--- 

MB ON PQ NB 

NF 

Genus HYDIIOTHASSA C.G. Thomson 

(Subgenus HYDROTIIASSA s. str.) 

(Erernosis des Gozis) 

H. boreela Schaeffer 

H. obliquata (Leconte) 

- YK 

H. ovalis tBlatchleyr 

H. vittata tOlivierJ* 

triuittatt (Say) 

uaripes r.LeConte) 

v:' 

NT 

AB 

AB 

SK MB 

SK MB 

ON PQ 

ON 

ON PQ 

NS 

Genus PRASOCURIS Latreille 

Helodes Paykull 

P. phellandrii (Linn6)* AK 


Genus PLAGIODERA Chevrolat 

(Subgenus PI-AGIODERA s. str.) 

(Me laso tn ida Schaeffer) 

(Pseudoiina Schaeffer) 

312

Chrysomelidae 

P. versicolora (Laicharting)t 

ar tnor aciae ( Fabricius) 

MB ON PQ NB 

Genus CHRYSOMELA Linne 

Key to New World species: Brown (1956) 

(Subgenus CHRYSOMELA s. str.) 

(Lina Dejean) 

(Melasoma Stephens) 

C. aeneicollis (Schaeffer) 

quadriguttoides Beller & Hatch 

C. blaisdelli (Van Dyke) AK 

C. confluens (Rogers) 

maculicollis Schaeffer 

C. crotchi W.J. Brown AK 

C. engelhardti (Hatch) AK 

C. falsa W.J. Brown AK 

C. invicta W.J. Brown 

C. knabi hesperia W.J. Brown 

C. knabi knabi W.J. Brown 

C. laurentia W.J. Brown 

C. lineatopunctata (Forster) 

nouaboracensis Gmelin 

obsoleta (.Say) 

s cr ip toide s ( Schaeffer) 

uariegata Olivier 

C. mainensis interna W.J. Brown 

C. mainensis littorea W.J. Brown AK 

C. mainensis mainensis Bechyn6 

C. schaefferi W.J. Brown AK 

immaculata (Schaeffer) 

C. scripta Fabricius 

C. semota W.J. Brown 

C. walshi W.J. Brown 

YK 

YK 

YK NT 

YK 

YK 

BC 

NT BC 

_BC 

AB 

NT_ABSKMB 

NT 


AB 

AB SK MB 

NT_AB 


)' 

BC AB 

BC 

_AB 

BC AB 

BC AB 

BC AB 

SK 

MB 

MB 

MB 

ON PQ NB 

ON PQ 

oN-- 

oN PQ 

oN PQ 

9* PQ 

ON 

ON PQ 

NS 

NB NS 

_LB 

_LBNF 

v 

Genus GONIOCTENA Chewolat 

Key to North American species: Brown (1942o) 

(Subgenus C.ONIOCTENA s. str.) 

(Phytodecta Kirby) 

G. americana (Schaeffer) AK 

arctica Bechyn6 

guttifera Bechyn6 

G. nivosa alberta W.J. Brown 

G. nivosa arctica (Mannerheim) AK 

G. nivosa nivosa (Suffrian) 

G. notmani (Schaeffer) 

G. occidentalis (W.J. Brown) 

YK 

NT 

BC 

-NT_AB 

YK NT BC AB 

YKNT_ABSK 

YK-BCAB 

AB SK 

MB ON 

MB 

MB ON 

PQ NB 

PQ 

LB 

LB 

Genus PHRATORA Chevrolat 


(Subgenus PHRATORA s. str.) 

(Phyllodecta l\rby) 

P. americana canadensis W.J. Brown 

P. frosti frosti W.J. Brown 

P. frosti remissa W.J. Brown AK 

P. hudsonia W.J. Brown AK 

P. kenaiensis W.J. Brown AK 

P. interstitialis Mannerheim AK 

aklauiki (Can) 

P. purpurea novaeterrae W.J. Brown 

P. purpurea purpurea W.J. Brown 

YK NT 

YK NT 

YK NT BC 

EC AB 

BC AB 

BC 

AB 

Sf 

ON PQ 

SK MB ON PQ 

sK yu oN PQ 

ff,fS ON Pe 

NB 

NN 

NS 

NS__ 

NS 

NF 

Genus ENTOMOSCELIS Chewolat 

C hry sornelop silo Strand 

Chrysomelopsis Achard 

Key to North American species: Brown (1942b) 

E. americana W.J. Brown AK YK NT BC AB SK MB 

313

Chrysomelic;.ae 

Subfamily GALERUCINAE 

Tribe Galerucini 

Key to North American species: Wilcox (1965) 

Genus DEROSPIDEA Blake 


D. brevicollis (LeContet 

ON 

Genus TRIRHABDA LeConte 


T. adela Bla-

Chrysomelidae 

Genus ERYNEPHAI-A Blake 


E. rnaritima (LeConte) 

texana Schaeffer 

E. puncticollis (Say) 

AB 

SK 

MB 

NS 

Genus MONOXIA LeConte 


M. angularis (LeConte) 

M. consputa LeConte 

M. debilis LeConte 

M. grisea Blake 

M. inornata Blake 

M. puberula Blake 

M. sordida LeConte 

Genus OPHRAELLA Wilct.rx 

Key to North American species: LeSage (1986) 

O. americana (Fabricius) 

O. arctica LeSage 

O. bilineata (Kirby) 

O. communa LeSage 

notulata anct. 

O. conferta (LeConte) 

O. cribrata (LeConte) 

O. notata (Fabricius) 

O. nuda LeSage 

O. pilosa LeSage 

NT 

-BC _BC 

_BC 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

SK 

SK 

SK 

SK 

SK 

SK 

MB 

MB 

MB 

MB 

yu 

MB 

9* PQ 

ON 

oN PQ 

ON PQ 

oN PQ 

ON PQ 

NB 

NS 

Genus GA.LERUCA O.F. Miiller 


(Subgenus GALERUCA s. str.) 

(Adimonia Laicharting) 

G. browni Blake 

G. externa (Say) 

G. rudis (LeConte) 

Blake (1945) 

Genus DIABROTICA Chewolat 

Tribe Luperini 

D. barberi Smith & Lawrence 

longicornis atct. 

D. cristata (Harris) 

atripennis (Say) 

- SK 

fossata LeConte 

D. undecimpunctata howardi Barber 

d uodec im p unclofa (Fabricius) 

D. virgifera virgifera LeConte 

filicornis Hotn 

Genus ACAI-YMMA Barber 

Key to Norbh American species: Munroe and Smith (1980) 

A. gouldi Barber 

A. vittatum (Fabricius) 

atnericanutn (Gmelin) 

melanocep halurz ( Fabrici us) 

pallipes (OIivier) 

stolatum (Gmelin) 

Genus CEROTOMA Chevrolat 

Andrector Horn 

Cerotana Bowditch 

Ceratorna Doziet 

C. trifurcata (Forster) 

MB ON 

MB ON 

PQ 

oN PQ 

ON 

_ONPQ 

MB ON PQ 

MB ON PQ 

NB 

NE NS 

315

ChrysomeliCae 

Genus PSELIDOLUPERUS Beller & Hatch 

P. lecontii (tlrotch) 

P. longulus (LeConte) 

baheri Beller & Hatch 

_BC 

-BC 

Genus SCELOLYPERUS Crotch 

S. cyanellusr (LeConte) 

S. nigrovirescens (FalI) 

S. schwarzi Horn 

S. torquatus (LeConte) 

S. varipes (i-eConte) 

concauus Beller & Hatch 

-BCAB 

_BC 

_BC 

_BCAB 

ON 

NS 

Genus PHI'LLOBROTICA Chevrolat 

P. decorata (Say) 

P. discoidea (Fabricius) 

circumdata (Say) 

P. limbata (Fabricius) 

BC 

AB 

SK 

MB 

-oN 

-oN 

PQ 

PQ 

NS 

Subfamilv ALTICINAE 

Genus BLEPHARIDA Chevrolat 

Key to North American species: Mignot (1971); Balsbaugh and Hays (1972) 

(Subgenus ULEPHARIDA s. srr.t 

(BlephuridulaWeise) 

B. rhois (Forster) AB SK MB ON 

atripennis }Iotn 

dorothea Mignot 

meticu'.osa (Olivier) 

roi (Ol vier) 

s/olido (Fabricius) 

Genus PHYLLOTRETA Chewolat 

Tanygctster Blatchley 

Key to North American species (part): Smith (1985) 

P. alberta Chittenden 

P. albionica (LeconteJ 

P. aerea Allardt 

P. armoraciae (Koch)t 

biplagiata Chittenden 

P. bipustule.ta t Fabricius t 

P. brevipen'ris Chittenden 

P. chalybeil'ennis tCrotch t 

P. conjunctrr Gentner 

P. constrict r E.H. Smith 

P. crucifera : I Goeze tt 

columl: iana Chittenden 

P. decipiens Horn 

ordina;ia Chittenden 

P. denticon is Horn 

a m pl ic cr ni s Chittenden 

aequal.is Hatch 

P. inconspicua Chittenden 

P. lewisii (Crotcht 

P. liebecki Schaeffer 

P. oblonga Chil,tenden 

P. pusilla Horn 

P. ramosoides E.H. Smith NT 

P. robusta l,eConte 

P. striolata (Fabricius)f - YK NT 

artiu ittata Chittenden 

lineolata Chittenden 

uerniccsa Chittenden 

uillolo (Fabricius) 

P. utanula li.H. Smith 

P. zimmerrnanni (Crotch)* AK - NT 

sinuatu Horn 

BC 

nc 

BC 

BC 

AB 

AB SK 

_SK 

AB SX 

AB SK 

AB SK 

AB 

ms 

MB 

MB 

MB 

ON 

oN PQ 

oN PQ 

ON PQ 

oN pe 

NB 

-NS 

NE NS 

BC-SKMB 

BC 

_MBON 

_ABSKMB 

-ABSKMBON 

BC 

_ABSKMBONPQ 


BC 

BC AB SK 

PE 

MB ON PQ NB NS PE NF 

316

Chrysomelidae 

Genus APHTHONA Chevroiat 

Cerataltica Ctotch 

A. cyparissiae (Koch)t 

A. flava Guillebeaut 

A. nigriscutis Foudrast 

AB?_ON-? 

AEr I ',' _ ') 

AB_MBON 

Genus LONGITARSUS Berthold 

(Subgenus LONGITARSUS s. str.) 

(Apterius Blatchley) 

(T einodacty la Chevrolat) 

(Thyamis Stephens) 

L. erro Horn 

L. fermgineus (Foudras)t 

L. ganglbaueri Heikertingert 

L. insolens Horn 

L. jacobaeae Waterhouset 

L. luridus (Scopoli)f 

L. melanurus (Melsheimer) 

L. pallescens Blatchley 

L. pellucidus (Foudras)t 

L. pratensis (Panzer)t 

L. rubiginosus (Foudras)f 

L. succineus Foudrasf 

L. suspectus Blatchley 

L. testaceus (Melsheimer) 

YBC 

-nc 

AB-MBONPQ 

-oNPQ 

MB 

AB 

ON PQ 

-oNPQ 

MB ON PQ 

_ONPQ 

-oN 

_ONPQ 

_ONPQ 

-oN 

_ONPQ 

NU 

NB 

NS 

NS 

PE 

PE 

NF 

v 

NT 

Genus GLYFTINA LeConte 

Balophila Ctotch 

G. abbreviata Gentner 

G. atriventris Horn 

G. brunnea Horn 

G. cyanipennis (Crotch) 

G. spuria LeConte 

lissotorques LeConte 

BC 

T 

sx lle 

_MB 

_MB 

ON 

ON 

ON PQ 

ON 

ON PQ 

Genus SYSTENA Chevrolat 

S. bitaeniata (LeConte) 

S. blanda (Melsheimer) 

ligata (.LeConte) 

ochracea (LeOonte) 

taeniata (.Say) 

S. carri Blake 

S. corni Schaeffer 

S. dimorpha Blake 

S. elongata (Fabricius) 

floridana Blatchley 

subdenea (LeConte) 

S. frontalis (Fabricius) 

S. hudsonias (Forster) 

S. marginalis (Illiger) 

oblonga (LeConte) 

_AB 

_AB 

AB 

AB_MBONPQ 

s^- 

_MB 

oN- 

_MBONPQ 

-MBONPQ ON PQ 

Genus STRABAJ-A Chevrolat 

(Subgenus STRABAI-A s. str.) 

Key to New World species: Blake (1953) 

S. rufa (Illiger) 

oN PQ 

Genus AJ-TICA O.F. Miiller 

(Subgenus ALTICA s. str.) 

(.Gr a p toder a Chevrolat) 

(Megaltica Hatc}l':' 

A. aeneola Blatchley 

aenescens Blatchley 

A. ambiens LeConte 

alni Hatris 

latiplicata Blake 

A. betulae Schaeffer 

NT BC 

-oN 

MB ON 

PQ 

PQ 

PQ 

NS 

317

ChrysomeLidae 

A. bimargirrata Say 

p licipe nnis ( Mannerheim) 

A. browni }4ohamedsaid 

potentillae W.J. Brown 

A. canadensis Gentner 

A. chalybar,ra Illiger 

uitiuota Thomas 

A. carinata Germar 

e*apta Say 

A. caurina Blake 

A. corni Woods 

A. elongatula Csiki 

elongata Gentner 

A. foliaceae LeConte 

puncti.oennis LeOonte 

A. heucherae FaiI 

A. ignita Illiger 

A. inaerata LeConte 

A. kalmiae Melsheimer 

uirida:za Schaeffer 

A. litigah lrall 

A. populi S'.J. Brown 

A. prasina LeOonte 

A. probata Fall 

A. ribis W.,1. Brown 

A. rosae Woods 

A. subplica;a LeConte 

A. sylvia Malloch 

cuprascens FalI 

torquo.ta LeConte 

A. tincta LeConte 

A. tombacina (Mannerheim) 

euicta LeConte 

shoemtkeri Schaeffer 

A. ulmi Wocds 

A. vicaria Ilorn 

A. woodsi Iiely 

Genus HOIINAJ-TICA Barber 

H. atriventris (Meisheimer) 

Genus ORT'HALTICA Crotch 

Leptot,brix Heikertinger & Csiki 

Le ptot. icha I ti co Heikertinger 

Leptot,ix Horn 

M ic re p itr ix Laboissidre 

Serraticollis R.E. White 

Key to North American species: Scherer (1974) 

O. copalina (Fabricius) 

fortico.nis (Illiger) 

Genus DEFiOCREPIS Weise 

D. erythroJ'us (Melsheimer) 

rufipes (Ctotch) 

D. oregoni .{atch 

Genus ORIISTIA Germar 

Orestirtides Hatch 

PyladiaWeise 

O. robusta LeContet 

Genus HIPPURIPHILA Foudras 


H. canadensis W.J. Brown 

H. equiseti Beller & Hatch 

H. manculzL (LeConte) 

modeeri anct. 


NT 

:-BC__ 

BC AB 

BC 

_AB 

AB SK 

SK 

BC AB SK 

-BC 


AK 

AK 

BC 

BC 

AB 

PQ 

MB ON 

MB 

-oN 

MB ON 

MB ON 

MB 

-oN 

MB 

-oN 

yu oN 

-oN 

MB ON 

_ONPQ 

-oNPQ 

l,fB ON 

MB ON 

-oN 

ON 

oN 

MB ON 

y" oN PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

Pe 

PQ 

PQ 

PQ 

PQ 

PQ 

PQ 

NB NS 

NB 

NB NS 

NB 

NU NS 

NU WS 

_MB 

-MBONPQ 

_MBON 

SK 

-NT_ABSK BC AB 

BC AB SK 

LB 

LB 

NF 

v 

318

Chrysomelidae 

Genus CREPIDODERA Chevrolat 

Key to North American species: Parry (1986) 

C. browni Parry 

C. decora Parry 

C. digra Parry 

C. heikertingeri (Lazorko) 

C. luminosa Parry 

C. nana (Say) 

c hittende ni Heikertinger 

C. populivora Parry 

C. sculpturata (Lazorko) 

C. solita Parry 

C. spenceri (Lazorko) 

C. vaga Parry 

C. violacea Melsheimer 

AK 

_ECAB 

_BCAB 

_ _ _:'_ 

NT BC AB 

NT BC AB 

_BC 

_BCAB 

s^ 

SK 

SK 

SK 

SK 

-oN 

-oNPQ 

MB ON PQ 

MB ON PQ 

_ONPQ 

MB ON PQ 

MB ON PQ 

MB ON 

MB ON PQ 

-oN 

_ONPQ 

r.rs lls 

NB NS 

NB NS 

NB 

NB NS 

NF 

T\TF 

Genus EPITRIX Foudras 

E. brevis Schwarz 

E. cucumeris (Harris) 

(?) pubescens llliget 

seminulum LeConte 

E. hirtipennis (Melsheimer) 

E. humeralis Dury 

E. subcrinita (LeConte) 

E. tuberis Gentner 

-oN 

_SKMBONPQ 

NS NS 

Genus MANTURA Stephens 

M. chrysanthemi (Koch)f 

M. floridana Crotch 

AB SK 

lrs on 

peNeNs--Nil 

Genus CHAXTOCNEMA Stephens 

(Subgenus CHAETOCNEMA s. str.) 

C. concinna (Marsham)'l 

C. confrnis Crotch 

flauicornis LeConte 

C. cribrata LeConte 

rudis LeConte 

C. cribrifrons LeConte 

C. decipiens LeConte 

C. denticulata (Illiger) 

americana Motschulsky 

s emichalchea Melshermer 

texana Ctotc}r 

C. ectypa Horn 

C. minuta Melsheimer 

C. opulenta Horn 

C. perturbata Horn 

C. protensa LeConte 

C. pulicaria Melsheimer 

(?) aeneola LeConte 

C. subcylindrica LeConte 

cylindrica LeConte 

C. subviridis LeConte 

v 

NT BC 

-nc _BC 

NT BC 

AB SK 

AB SK 

_SK 

lrs om pe Ne 

MB ON PQ NB 

_ONPQ 

-oN 

_SKMB 

MB ON PQ 

AB SK MB 

_SKMB 


AB 

ON 


AB SK MB 

_PE 

NS 

NS 

NB NS 

NF 

Genus DISONYCHA Chevrolat 


D. alternata (Illiger) 

punctigera Schaeffer 

(? ) q uinq ueu ittaro ( Say) 

D. arizonae Casey 

borealis Blake 

douisi Schaeffer 

glabrata Jacoby 

D. latifrons Schaeffer 

osferis Schaeffer 

laticollis Schaeffer 

D. latiovittata Hatch 

D. pensylvanica (Illiger) 

_BC 

BC 

BC 


MB ON 


ON PQ 

319

Chrysomel,dae 

parua Blatch\ey 

pensyiuania Stwn 

(?) s e x,t ineata (Olivier) 

D. procera Casey 

n igriutntris Schaeffer 

(?) pallipes Crctch 

(?) uici.na l\trby 

D. punctigera LeConte 

'neglecta Schaeffer 

punctipennis Schaeffer 

D. schaefferi Blake 

D. triangularis (Sayr 

montanensis Blake 

puncticollis (Kirby) 

D. uniguttr.ta { Say t 

uicina \{ttby 

D. xanthonrelas ( Dalman) 

atrella Blake 

ceruicctlis LeConte 

collaris (Illiger) 

merdiuora Melsheimer 

xanthomelaeno Gemminger & Harold 

NT BC A3 

NT AB 

SK MB ON PQ 

SK 

- - scABSf 

NT AB SK 

MB 

MB ON PQ 

MB ON PQ 

MB ON PQ 

PE 

Genus KUSICHELINA Bechyn6 


K. barberi (Blake) 

K. frmbriat.r (Forster) 

circun cincta (Crotch) 

suture'Ia (Say) 

K. gibbitanra (Say) 

K. lugens (li,eConte) 

lamprt)cydnea (B1ake) 

K. perplexzi (Blake) 

K. thoracica. (Fabricius) 

flaua (Gmelin) 

K. vians (Il iger) 

abdon',.inalis ( Olivier) 

badia Blake) 

scripti.:ollis (Say) 

NT 

_AB 

- -4,8 

BC 

NT BC 

AB 

SK MB 

SK MB 

ON 

ON 

ON 

ON 

ON PQ 

NS 

Genus CAPRAITA Bechyn6 

ChloephagaWeise 

Key to Nrrrth American species: 

C. circumditta (Randall) 

limbalis (Melsheimer) 

(?) que"cata (Crotch) 

C. scalaris t,Melsheimer) 

lobata (LeConte) 

C. sexmacuLata (Illiger) 

cretica (Jacoby) 

palliatt (RandalI) 

C. subvittata (Horn) 

C. thyamoides (Crotch) 

Blake (1927) 

AB 

SK MB 

ON 

oN PQ 

ON 

oN PQ 

ON 

NB NS 

Genus DISTIGMOPTERA Blake 


D. apicalis l3lake 

D. borealis Blake 

D. pilosa (Illiger) 

Genus DIBOLIA Latreille 

Key to N'rrth American species: 

(Subgenus IDIBOLIA s. str.) 

D. borealis tlhevrolat 

D. chelones Parr5r 

D. Iibonoti l{orn 

D. melampyri Parry 

D. penstemrrnis Parry 

D. reyheria Mignot 

Blake (1943) 

Parry (1974) 

BC AB SK 

sc 

AB 

T 

AB 

SK MB 

_MB 

-oN 

MB ON PQ 

-oNPQ 

NB 

_PQNBNS 

oNPQ-NS 

ONPQNBNS___ 

9*PQIUNS_-_ 

320

Chrysomelidae 

Genus PSYLLIODES Berthold 

P. affinis (Paykull)t 

P. chrysocephala (Linn6)t 

P. convexior LeConte 

P. cucullata (Illiger)f 

P. elegans Horn 

P. napi (Fabricius)f 

P. picina (Marsham)-l 

P. punctulata Melsheimer 

aenescens Casey 

extricata Casey 

paruicollis LeConte 

AB 

AB 

-oN 

-oN 

PQ 

_ONPQ 

_ONPQ 

MB ON PQ 

NB 

NB 

_NS 

--Ne 

pn-*" 

I\TE] 

Genus ANISOSTENAWeise 

(Subgenus ANISOSTENA s. str.) 

A. californica Van Dyke 

A. nigrita tOlivier) 

Genus SUMITROSIS Butte 

Anoplitis Chapuis 

Key to North American species: Butte (1969) 

Subfamily HISPINAE 

Tribe Chalepini 

Key to North American genera: Butte (1968o) 

_BC 

-BCMBONPQ 

S. ancoroides (Schaeffer) 

S. inaequalis (Weber) 

bazcis (Newman) 

- BC 

flauipes (Germar) 

neruosa (Hotn') 

obsoleta (Say) 

pallida (Say) 

suturalis (Fabricius) 

S. rosea (Weber) 

philemon (Newman) 

AB SK MB 

AB SK MB 

NB 

oN pe N" 

NS 


Genus ODONTOTA Chevrolat 

Key to North American species: Butte (1968c) 

O. dorsalis tThunberg) 

hanisi Crotch 

scutellaris (Olivier) 

suturalis (Harris) 

O. mundulus (Sanderson) 

O. scapularis t Oliviert 

lateralis (Say) 

Genus CHALEPUS Thunberg 

Anoplitis Kirby 

Key to North American species: Butte (1968b) 

C. bicolor (Olivier) 

rufiuentris (Suffrian) 

rufogaster Thunberg 

MB 

-oN 

oN PQ 

ON 

Genus BALIOSUS Weise 

B. nervosus (Panzer) 

marginatus (Say) 

marmoratus (Baly) 

obsoletus (Say) 

pallipes (Germar) 

quadratus (Fabricius) 

quercifoliae (Harris) 

rubellus (Schonherr) 

ruber (Weber) 

ON 

PQ 

NB 

NS 

32L

ChrysomelLdae 

Genus STEiNOPODIUS Horn 

Key to North American species: Blaisdell (19386) 

Tlibe Uroplatini 

S. lateralis lSchaeffer) AB SK 

uand.uzeei Blaisdell 

S. texanus (Schaeffer.t AB SK MB 

Genus BRACHYCORYNA Gu6rin-M6neville 

Key to North American species: Staines (1986) 

B. hardyi ((lrotcht 

confusa Hatch 

- BC AB 

horni'Weise 

melsheimeri Horn 

B. montana (Horn) AB SK MB ON 

Genus GL)?HUROPLATA llhmann 

Key to North American species: Riley (1985b) 

G. pluto (Newman) AB - MB ON PQ 

porcatd (Melsheimer) 

Genus MIC.IRORHOPALA Chevrolat 

Key to North American species: Clark (1983) 

M. excavat.r cyanea { Say } - BC AB SK MB 

M. excavatrr excavata (Olivier) MB ON PQ NB NS 

M. vittata tFabricius) - BC AB SK MB ON PQ NB 

laetuluLeConte 

M. xerene (Newman) AB SK MB ON PQ NB 

interrt.tpta Couper 

Subfamily CASSIDINAE 

Tribe Stolaini 

Key to genera: Hincks (1952) 

Genus CHINLYMORPHA Chevrolat 

C. cassidea (Fabricius) 

orgus iLichtenstein) 

cribraria (Olivier) 

geniculata Boheman 

lewisi Crotch 

s e p t e rrtde ci m p u nc t at a (S ay) 

_SKMBONPQ 

Tribe Physonotini 

Genus PHI'SONOTA Boheman 

Key to North American species: Sanderson (1948) 

(Subgenus PIfYSONOTA s. str.) 

P. helianthi tRandallt MB ON PQ 

q uinquepu nct ato (Walsh) 

sep ten,tr io nalis Boheman 

P. unipunctata (Say) - SK MB ON PQ 

Genus CASSIDA Linn6 

Key to North American species: Riley (1986b) 

(Subgenus CASSIDA s. str.) 

T?ibe Cassidini 

Key to North American genera: Riley (1986a) 

C. azurea F'abriciusf - SK 

C. flaveola Thunbergt - YK NT BC AB SK MB ON PQ 

C. rubiginosa O.F. Miiller+ AB SK MB ON PQ NB 

Genus JON'THONOTA Spaeth 

J. nigripes .Olivier) AB SK MB 

ellipsis (LeConte) 

322

Nemonychidae 

Genus AGROICONOTA Soaeth 

A. bivittata (Say) 

striolata (Boheman) 

uittula (Boheman\ 

MB ON 

PQ 

Genus PLAGIOMETRIONA Spaeth 

(Subgenus PARAMETRIONA Spaeth) 

P. clavata (Fabricius) 

diu er s icoll is ( Schaeffer) 

ON 

PQ 

NB 

Genus STRONGYLOCASSIS Hincks 

Strongylaspis Spaeth 

Key to species: Riley (1985o) 

S. atripes (LeConte) 

creolo (Sanderson & King) 

loedingi (Schaeffer) 

MB ON 

Genus DELOYALA Chewolat 

Chirida Chapuis 

D. guttata (Olivier) 

bohemani (Spaeth) 

columbica (Spaeth) 

cruciata (Fabricius) 

fuliginosa (OIivier) 

hamulata (Boheman) 

immunita (Boheman) 

immunda (Boheman) 

lucidula (Boheman) 

lugubrina (Boheman) 

p ennsy lu anic a ( Spaeth) 

signifera (Herbst) 

trabeata (Boheman) 

SK MB ON PQ 

NB 

NS 

PE 

Genus CIIARIDOTELLA Weise 

(Subgenus CHAEROCASSIS Spaeth) 

C. purpurata (Boheman) 

C. sexpunctata bicolor (Fabricius) 

auric halcea (Fabricius) 

auris plendens (Mannerheim) 

b istrip ustulata (Herbst) 

floridana (Schaeffer) 

(? ) mary landico ( Herbst) 

pallida (Herbst) 

SK 

MB ON 

-oN 

PQ 

PQ 

NB 

NS 

NS 

Superfamily Curculionoidea 

Family NEMOI{YCHIDAE 

J. McNamara 

Nemonychidae comprises eight species in 

Canada. The adults feed on the staminate flowers 

ofvarious conifers. 

The nemonychids are often included in 

Curculionidae, but their larvae show an aflinity to 

Anthribidae. Kuschel (1989) recently published a 

revision of this family for North America The list 

below follows O'Brien and Wibmer (1982). 

BC (6); AB (3);MB (1); ON (2);PQ (3); NB (1) 

323

Anthribidae 

Genus CIMBERIS des Gozis 

C. bihirsuta Hatch 

C. compta (LeConte) 

paruulus lJatch 

C. decipiens Kuschel 

C. eiongata (LeConte) 

C. pallipennis (Blatchley) 

C. pilosa (LeConte) 

Genus PITYOMACER Kuschel 

P. pix Kuschel 

Subfamily DOYDIRFIYNCHINAE 

Tribe Cimberidini 

BC 

BC 

BC 

BC 

BC 

AB 

AB 

MB ON 

-oN 

PQ 

PQ 

PQ NB 

Genus ACFI OMACER Kuschel 

A. bombifrr: ns (LeConte) 

BC 

AB 

Family ANTHRIBIDAE fungus weevils 

J. McNamara 

Anthribidae includes 18 species in Canada and 

Alaska. One of these, the coffee bean weevil 

(Araecerus fasciculatus), is introduced and may be 

establishred in warehouses; this species damages 

many stored plant materials, including seeds. 

Adults of anthribids are often collected by beating 

dead or diseased vegetation or by sweeping in 

weedy areas. The larvae are usually found in similar 

habitats. Adults feed on fungr, pollen, or even 

bark; larvae feed in plant stems, fungr, orvvoods. 

Valentine (1960) has revised the North 

American genera of Anthribidae. 

AJ{ (2); YK (2); NT (2); BC @); AB (8); S.K (6); MB 

(7); ON (12); PQ (12); NB (1);NS (1);NF (1) o fl 

(1); I (1) 

Subfamily CHORAGINAE 

T?ibe Araecerini 

Genus ARAECERUS Schrinher"r 

A. fasciculatus (DeGeer)t 

BC 

ON 

PQ 

Tlibe Choragini 

Genus EIIXENUS LeConte 

E. punctatus LeConte 

PQ 

Genus EUPARIUS Schdnherr 

Key to Norlh American species: 

E. marmoreus (Olivier) 

albifrons (Boheman) 

lunatus (Fabricius) 

Valentine (1960) 

Subfamily ANTHRIBINAE 

TYibe Eupariini 

MB ON PQ 

NB 

Genus TRICIONORHINUS Wollaston 

Anthrihulus LeConte 

Brachy tarsoides Pierce 

Key to wrstern North American species: 

T. alternatus (Say) 

T. annulatus (Carr) 

Vaientine (1971) 

Tlibe Anthribini 

_BC 

AB 

AB 

MB ON 

324

Ithyceridae 

T. griseus (LeConte) 

T. limbatus (Say) 

T. sticticus (Boheman) 

obsoletus (Fahreus) 

uariegatus (Say) 

T. tomentosus tomentosus (Say) 

6reuis (Fahreus) 

paululus (Casey) 

- - - - ffi3f reoNpe - - - - - 


ON 

Genus ORMISCUS Waterhouse 

Eusphyrus LeOonte 

Gonops LeConte 

Toxotropis LeConte 

O. saitator LeConte 

angulatus Pietce 

O. walshii (LeConte) 

Tribe Ormiscini 

Tribe Tropiderini 

oN PQ 

ON 

Genus TROPIDERES Schbnherr 

Eurymycter LeConte 

Gonotropis LeConte 

'Key to North American species: 

T. dorsalis (Thunberg)* 

gibbosus (LeConte) 

T. fasciatus (Olivier) 

bicarinatus (Pierce) 

T. latifascia (Pierce) 

T. tricarinatus (Pierce) 

Valentine (1960) 

AK 

YK 

NT 

_BC 

AB SK MB 

PQ 

SK MB ON PQ 

_ _ _ oNiS - 

NS 

NF 

Genus AI,LANDRUS LeConte 

Key to North American species: Valentine (1960) 

A. bifasciatus LeConte 

A. brevicornis Frost 

A. populi Pierce 

Tribe Allandrini 

----AB-nreoNPQ----- 

AK \'K AB SK _ ON PQ 

Family ITIIYCERIDAE 

New York weevil 

J. McNamara 

Ithyceridae includes only one species. The 

adults feed on the tender shoots and buds ofvarious 

hardwoods and occasionally on some cultivated fruit 

trees. The larvae feed on the roots of the host 

plants. 

Sanborne (1981) studied the morphology of both 

larvae and adults, as well as the biology of the 

species. 

ON (1); PQ (1) 

Genus ITHYCERUS Schonherr 

Pochyrhynchus Kirby 

L noveboracensis (Forster) 

cur culionoides (Herbst) 

p unc ta t uI us ( Fabricius) 

schoenherri (Kirby) 

oN PQ 

325

Rhynchitidrre 

FamilyATTELABIDAE leaf-rollingweevils 

J. McNamara 

Four species of Attelabidae occur in Canada. 

The females form rolls from leaves and lay a single 

egg in each roll. The larvae feed on the inner part 

of the roll. 

The classification used here follows that of 

Robert W. Hamilton (unpublished Ph.D. thesis, 

1969), who revised the North American species. 

SK (1); MB (2); ON (3); PQ (a); NB (1); NS (2) 

Genus HIMATOLABUS Jekel 

H. pubescerrs (Say) 

colora

Brentidae 

ruficollis (Germar) 

sulcifrons Gyllenhal 

Genus PSELAPHORHYNCHITES Schilsky 

Key to North American species: Hamilton (1971) 

p. cyanellus (LeConte) NT - AB sK MB ON PQ NB NS PE - NF 

P. perplexus (Blatchley) - ON PQ NB NS PE 

Genus MERHYNCHITES Sharp 


M. bicolor bicolor (Fabricius) NT BC AB SK MB oN PQ 

antennalis (Yoss) 

intermedius (Yoss) 

M. bicolor cerdonis (Voss) - BC AB 

M. bicolor cockerelli (Pierce) AB 

erythrosoma (Cockerell & Harris) 

M. bicolor nigricephalus Hamilton - BC 

M. wickhami wickhami (Cockerell) NT BC AB SK MB 

homocidae (Yoss) 

mandibularis (Yoss 

Genus HAPLORHYNCHITES Voss 


H. aeneus (Boheman) - SK MB 

consobrinus (Yoss) 

Genus AULETOBIUS Desbrochers 

Key to North American subgenera: Hamilton (1983) 

(Subgenus ALETINUS Desbrochers) 

TYibe Auletini 

A. ater (Leconte) MB oN PQ 

(Subgenus MESAULETES Voss) 

A. cassandrae (LeConte) - ON PQ NB NS NF 

albou e stitus (Blatchley) 

blatchleyiYoss 

minor (Blatchley) 

(Subgenus AULETOBIUS s. str.) 

Metopon Waterhouse 

A. congruus (Walker) - BC AB SK MB 

subcoeruleus LeOonte 

Family BRENTIDAE brentid beetles 

J. McNamara 

Only one species of Brentidae, the oak timber- into the heartwood, often causing serious injury to 

worm, occurs in Canada. The adults are found oak stock. 

under bark or in decayed wood. The larvae bore ON (1); PQ (1) 

Genus ARRHENODES Schdnherr 

T?ibe Arrhenodini 

A. minutus (Drury) - ON PQ 

brunneus (Panzer) 

lecontei (Power) 

maxillosus (Olivier) 

p aup er c ula (Blatchley) 

sallei (Power) 

s epte ntrionis (Herbst) 

327

Apionidae 

Family APIONIDAE pear-shaped weevils 

J. McNamara 

Thirty-five species of Apionidae are recorded in 

Canada and Alaska. The larvae feed on leaves and 

stems, carrsing galls, or develop within the seeds of 

various plants. The adults usually feed on leaves. 

Than}s to the work of Kissinger (1968), the taxonomy 

of t;he North and Central American species is 

well known. 

aK (3); vK (3); NT (3); BC (12); AB (20); SK (13); 

MB (15); ON (17); PQ (13); NB (2); NS (2); NF (1) 

. I(1) 

Tlibe Nanophyini 

Genus NAN{)PHYES Schonherr 

Nanoda cty lus Blatchley 

Nanodes Schtinherr 

P se udo ty c hius Blatchley 

(Subgenus NANOPHYES s. str.) 

N. canadens s W.J. Brown 

AB 

SK MB 

T?ibe Apionini 

Genus APION Herbst 

(Subgenus ALOCENTRON Schilsky) 

A. attenuatum Smith 

(Subgenus C ERATAPION Schilsky) 

BC A,E MB ON 

PQ 

A. contusum Smith 

notabile, Buchanan 

spinipes Fall 

A. extensum Smith 

(Subgenus C OELOCEPHALAPION Wagner) 

SK 

PQ 

A. carinatun. Smith 

concoloratum Srlrilh 

A. emaciipes Fall 

A. segnipes Siay 

(Subgenus E UTRICHAPION Reitter) 

ON 

ON 

ON 

A. alaskanun Fall AK 

A. cavifrons .-eConte 

A. cyanitinctum Fall 

A. huron Falt 

AK 

(Subgenus F,{ILAPION Kissinger) 

A. finitimum FaIi 

A. impeditum Fall 

A. melanarium Gerstaecker 

A. occidentale Fall 

A. pennsylvanicum Boheman 

erythrocerum Srlrith 

A. robustum Smith 

obesum Srrlith 

A. speculifenrm Fall 

(Subgenus L-APION Roudier & Tempere) 

(ftios Deville) 

A. frosti Kissinger 

A. idiastes K.,ssinger 

(Subgenus PERAPION Wagrrer) 

YK 

YK 

NT BC AB 

_BCAB 

NTBCAB_MB 

AB_MB 

_ _ BCT 

_AB 

BC AB 

AB 

AB 

SK 

SK 

SK 

SK 

-pe 

ON 

MB ON 

-oN 

mn oN 

MB 

-oNPQ 

-ONPQ 

PQ 

PQ 

PQ 

PQ 

NB 

NS 

A. punctinasrm Smith 

A. wickhami Kissinger 


_ABSK 

328

Curculionidae 

(Subgenus PSEUDAPION Schilsky) 

A. amaurum Kissinger 

A. capitone Kissinger 

A. disparatum Sharp 

nasutum Fall 

-AB_MB _ABSKMB 

_AB_MB 

(Subgenus RHOPAIAPION Schilsky) 

A. longirostre Oliviert 

BC 

ON PQ 

(Subgenus TRICHAPION Wagner) 

A. centrale Fall 

A. commodum Fall 

A. porcatum Boheman 

A. proclive LeConte 

crassinasum LeConte 

nunenmacheri Wagner 

A. reconditum Gyllenhal 

turbulentum Smith 

A. simile Kirby 

lanuginosum. Walsh 

uicinum Srrrith 

walshii Smit}r 

A. tenuirostrum Smith 

_:" 

-BC 

BC 

AB SK MB 

TSKMB 

AB SK 

AB SK 

ON 

ON 

-oN 

MB ON 

MB 

PQ 

PQ 

NB NS 

NF 

$ 

A. antennatum Smith 

AK YK 

BC 

Tlibe Aplemonini 

Genus PODAPION Riley 

P. gallicola Riley 

alemnderi Sleeper 

ON PQ 

Family CURCULIONIDAE snout beetles or weevils 

J. McNamara 

The family Curculionidae includes 609 species 

and subspecies in Canada and Alaska. AII of them 

are phytophagous. Most of the larvae live inside 

the tissues of the host plant, but some feed externally 

on the leaves. The adults feed on leaves, 

pollen, flowers, developing fruit, and fungi, and 

some burrow in wood; they do not necessarily feed 

on plants preferred by the larvae. Many weevils are 

host-specifrc and some are economic pests. 

Kissinger (1964) wrote a key to the genera. 

O'Brien and Wibmer (1982) published a checklist of 

the North American species with supplements in 

1984 and 1989, the latter under Wibmer and 

O'Brien. The classification used here follows that 

checklist. 

AK (61); YK (51); NT (55); BC (261); AB (206); SK 

(I24); MB (179); ON (312); PQ (280); NB (72); NS 

(91); PE (17); LB (11); NF (68) ' H (14); I (69) 

Subfamily BRACITYDERINAE 

Tribe Naupactini 

Genus ASYNONYCHUS Crotch 

A. cervinus (Boheman)t 

fulleri (Il.orn) 

ON 

329

Curculionidae 

Tribe Barynotini 

Genus TRI(IONOSCUTA Motschuisky 

Key to North American species: Pierce (1975) 

(Subgenus I)RIGONOSCUTA s. str.) 

T. pilosa iritlescens Pierce - BC 

Genus STAMODERES Casey 

S. lanei {Van Dyke) - BC 

Genus BAR'INOTUS Germar 

Key to Nc'rth American species: Brown (1950) 

B. moerens Fabricius)t NS 

B. obscurus (Fabricius)t - BC PQ NB NS PE - NF 

B. schoenherri (Zetterstedt)t PQ NB NS NF 

Genus POLYDRUSUS Germar 

Cyphontimus Horn 

Poly drt ts u s Schcinherr 

Key to Ncrth American species: Sleeper (1957a) 

Tribe Polydrosini 

P. americanrs (Gyllenhalt - ON PQ 

d.orsali:; (Horn) 

P. cervinus,Linn6tt PQ 

P. impressif'rons (Gyllenhal)+ - - MB ON PQ - NS 

P. sericeus ( Schallertf - ON PQ NB 

Genus PACI{YRHINUS Schrinherr 

S cy t hrt:. p u s Schdnherr 

P. californic -rs (Horn) - BC 

P. elegans(Oouper) - BC AB ON PQ - NS 

metall i c us (Desbrochers) 

P. ferrugine:s (Casey) - BC 

Genus BARYPEITHES Jacquelin du VaI & Fairmaire 

Tlibe Brachyderini 

B. pellucidus (Boheman)t - BC - ON PQ - NS PE - NF 

Genus SCIAPHILUS Schtinherr 

S. asperatus (Bonsdorff)t - BC - ON PQ NB NS NF 

Genus BRAOHYSOMUS Schonherr 

Thricol zpoide s O'Brien 

B. echinatus (Bonsdorfl)* PQ NF 

clauisetis (O'Brien) 

Genus STROPHOSOMA Billberg 

Tribe Strophosomini 

S. melanogrammum (Forster)f - BC - ON PQ - NS NF 

Genus PHILOPEDON Schtinherr 

P. plagiatunL (Schaller)t 

Tlibe Cneorhinini 

_NBNSPE-NF 

330

Curculionidae 

Tribe Sitonini 

Genus SITONA Germar 

S. californicus (Fahraeus) 

angustulus (Casey) 

apacheanus (Casey) 

cockerelli Blaisdell 

explicatus (Casey) 

ertrusus (Casey) 

hispidiceps (Casey) 

oregonensis Tanner 

prominens (Casey) 

S. cylindricollis (Fahraeus)t 

alpinensis Tanner 

S. hispidulus (Fabricius)t 

S. lepidus Gyllenhal 

S. Iineatus (Linn6)t 

S. lineellus (Bonsdorfi)t 

S. lupinus Sleeper 

S. scissifrons Say 

uittatus (Couper) 

S. tibialis (Herbst)t 

f_ 

-YK 

AK_ 

AK 

BC 

_BCABSK 

-BC _BC 

-BC 

NT BC 

_BC 

NT BC 

_BC 

AB 

AB 

AB 

AB 

sr 

SK 

MB ON PQ 

oN PQ 

MB ON PQ 

MB ON 

ln" oN pe 

_ONPQ 

NB 

NS 

NB NS 

NB NS 

NB NS 

lln 

NS 

NF 

NT 

NF 

Genus MINYOMERUS Horn 

Elissa Casey 

Pseudelissa Casey 

M. innocuus Horn 

Genus TANYMECUS Germar 

T. confusus Say 

confertus Gyllenhal 

uariabilis Fahraeus 

Genus PANDELETEIUS Schdnherr 


P. hilaris (Herbst) 

pauperculus Gyllenhal 

T[ibe Tanymecini 

AB 

NT-ABSK 

MB 

_ONPQ 

Subfamily OTIORFIYNCHINAE 

Tlibe Otiorhynchini 

Genus OTIORIIYNCHUS Germar 

B r ac hy r hinu s Latreille 

Key to North American species: Warner & Negley (1976) 

O. desertus Rosenhauerf 

O. Iigneus (Olivier)t 

O. ligustici (Linn6)t 

O. ovatus (Linn6)t 

O. porcatus (Herbst)t 

O. raucus (Fabricius)t 

O. rugifrons (Gyllenhal)f 

O. rugosostriatus (Goeze)f 

O. scaber (Linn6)t 

O. singularis (Linn6)l 

picipes (Fabricius) 

O. sulcatus (Fabricius)t 

AK YK 

AK 

NT SC AB 

-sc- 

_BC 

_BC 

SK 

NSNS- 

-Olf- 

MB ON PQ 

PQ 

-oN 

-ONPQ 

-ONPQ 

_MBON 

PQ 

PQ 

NB NS 

___ 

NB NS 

_NS 

_NS 

NB NS 

NB NS PE 

LB 

NF 

NF 

NF 

NF 

NF 

NT 

NF 

Genus SCIOPITHES Horrr 

S. obscurus Horn 


brumalis Casey 

significans Casey 

BC 

331

Curculionidae 

Genus AGRONUS Horn 

A. carri Bu

Curculionidae 

Genus APHRASTUS Say 

M icrony c h us Provancher 

A. taeniatus Say 

sulcatus (Provancher) 

Genus PHYXELIS Schdnherr 

Geoderces }Iotn 

P. rigidus (Say) 

glomerosus Boheman 

melanothrix (Kirby) 

setiferus Boheman 

Tribe Leptopiini 

ON PQ 

oN PQ NB NS PE NF 

Genus LEPIDOPHORUS Kirby 

Lophalophus LeConte 


L. inquinatus (Mannerheim) 

L. lineaticollis Kirby 

L. pumilus Buchanan 

Genus CIMBOCERA Horn 

C. pauper Horn 

Genus PLINTHODES LeConte 

P. taeniatus (LeConte) 

Buchanan (1936b) 

AI{ 

BC 

AK YK NT BC 

-BCAB 

BC 

AB 

Genus TRICHALOPHUS LeConte 

T. alternatus (Say) 

T. brunneus Van Dyke 

T. didymus (LeConte) 

constrictus (LeConte) 

T. seriatus (Mannerheim) 

T. simplex LeConte 

T. stefanssoni Leng 

_YK-BC 

AK 

AK 

AK 

AK_NT 

NT 

BC 

BC 

AB 

AB SK 

AB SA 

MB ON 

MB 

MB 

Genus LEPESOMA Motschulsky 

Amnesia Horn 

Dyslobus LeConte 

Melamomphus l{orn 

Thricomigus Horn 

Key to North American species: Van Dyke (1933) 

L. alternatum (Horn) 

L. bakeri (Van Dyke) 

L. decoratum (LeOonte) 

L. granicolle granicolle (LeConte) 

debilis (Casey) 

discors (Casey) 

sculptilis (Casey) 

L. lecontei (Casey) 

L. luteum (Horn) 

L. raucum (Horn) 

ciliatum (Pierce) 

L. tanneri (Van Dyke) 

L. ursinum (Horn) 

L. verruciferum (Casey) 

b ituberculatum (Pierce) 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

BC 

YSK 

SK 

Genus PANSCOPUS Schdnherr 

Key to North American species: Buchanan (1936o) 

(Subgenus NEOPANSCOPUS Pierce) 

P. aequalis (Horn) 

uestitus (Casey) 

(Subgenus NOCHELES LeConte) 

(Panscopidius Pierce) 

P. torpidus (LeContet 

BC 

BC 

AB SK MB 

333

Curculionidae 

(Subgenus IIIOMIDUS Casey) 

P. abruptus (Casey) - BC 

P. ovalis Pi,rrce - BC AB SK 

P. pallidus Suchanan - BC AB 

P. rugicollis Buchanan - BC 

(Subgenus I)ANSCOPUS s. str.) 

P. erinaceu,r (Say) - ON PQ 

carinat:us Pierce 

(Subgenus PARAPANSCOPUS Buchanan) 

P. maculosu s Blatchley - ON 

P. ovatipennis Buchanan - ON 

(Subgenus PlIfMAfl NUS LeConte) 

P. gemmatu.s (LeConte) - BC 

(Subgenus PSEUDOPANSCOPUS Buchanan) 

P. costatus Buchanan - BC 

Genus CON NATICHELA Anderson 

C. artemisir,Le Anderson AK YK NT 

Genus ANAMETIS Horn 

A. granulatrr (Say) - BC AB SK MB ON PQ NF 

grisea':Jorn 

Genus TRO PIPHORUS Schrinherr 

T. elevatus Herbst)t - NF 

carinatus (O.F. Mtiller) 

T. obtusus (Bonsdorff)t NS NF 

T. terricola.Newman)l PQ - NS PE - NF 

Genus OPIIRYASTES Say 

AmydrDgmus Pierce 

Eupagoderes Horn 

?ososfes Sharp 

Tribe Ophryastini 

Key to North American species: Kissinger (1970) 

O. cinerascens (Pierce) - BC 

columbianus (Van Dyke) 

O. sulcirost:ts (Say) NT - AB SK MB 

ligatus LeConte 

O. tuberosu; LeConte AB 

collaris Champion 

Genus CYR'IEPISTOMUS Marshall 

Subfamily EREMNINAE 

Tribe Cyphicerini 

C. castaneu; (Roelofs.tt - ON 

Genus CAI-(JMYCTERUS Roelofs 

C. setarius lloelofsf - ON 

Subfamily RIIYTIRRHININAE 

Genus LISTRONOTUS Jekel 

Hyperodes Jekel 

MacroP'9l{n"rY 

Key to Nc,rth American species: Henderson (1940); O'Brien (1981) 

L. alternatus (Dietz) PQ NB 

L. anthracir.us {Dietz) - ON PQ 

334

Curculionidae 

L. appendiculatus (Boheman) 

floridensis Blatchley 

impressus Van Dyke 

leucozonatus Chittenden 

L. californicus (Dietz) 

L. caudatus (Say) 

L. delumbis (Gyllenhal) 

indistinctus (Dietz) 

schwarzi (LeConte) 

solutus (Boheman) 

spurcus (Boheman) 

L. echinodori O'Brien 

L. frontalis LeConte 

L. humilis (Gyllenhal) 

subcribratus (Dietz) 

L. maculicollis (Kirby) 

mirabilis (Dietz) 

myasellus (Dietz) 

L. montanus (Dietz) 

L. oregonensis oregonensis (LeConte) 

impressifrons LeConte 

rudipennis Blatchley 

L. oregonensis tessellatus Casey 

L. porcellus (Say) 

minitnus (Blatchley) 

L. punctiger LeConte 

gracilis LeConte 

leechi Sleeper 

L. sordidus (GyIlenhal) 

disting uendu s (Gyllenhal) 


L. sparsus (Say) 

inbellis (Dietz) 

itnmundus (Boheman) 

latiusculus (Boheman) 

lineatulus (Say) 

longulus (Dietz) 

squalidus (Gyllenhal) 

L. squamiger (Say) 

inaeq uali pennis (Boheman) 

L. truncatus (Hatch) 

L. tuberosus LeConte 

L. vitticollis (Kirby) 

interp unctatulus (Dietz ) 

inter s tit ialis (Dietz) 

tenebrosus (Dietz) 

ulkei (Dietz) 

A"- 

AK YK 

BC 

-AB 

BC AB 

EC AB 

BC 

BC AB 

BC 

BC 

NT_AB 

BC 

BC 

AB 

AB 

AB 

AB 

AB 

SK MB ON 

ON 

SK MB ON 

_MB 

SK_ON 

SK MB 

_MBON 

_MBONPQ 

SK MB ON 

_MBON 

SK MB 

PQ 

prl 

pa:) 

PQ 

PQ 

PQ 

PQ 

PQ 

ON PQ 

PQ 

Pc) 

NF 

Genus LIXELLUS LeConr,e 

Anchodemus LeConte 

L. filiformis LeConte 

L. Iutulentus (Boheman) 

angustus (LeConte) 

Genus BYRSOPAGE S Schdnherr 

B. carinatus Motschuisky 

- Tf'BCfsKMBoNiS- - - - - 

AK 

Subfamily THECESTERNINAE 

Genus THECESTERNUS Say 

Lithodus Germar 

T. affinis (LeConte) 

eroszs (LeConte) 

morbillosus (LeConte) 

rectus (LeConle) 

rudis (LeConte) 

T. humeralis (Say) 

AB 

AB 

335

Curculioni lae 

Genus HYPERA Germar 

Phytonomus Sch6nherr 

H. castor (LeConte) 

H. compta (Say) 

diuersz (Gyilenhal) 

H. diversipunctata (Schrankrx AK - AB SK - ON 

H. eximia (LeConte) 

H. meles (['abricius)t 

H. nigriros iris (Fabricius)i 

uiridi.. (Provancher) 

H. postica iGyllenhal)f 

H. pubicoli.s (LeConte) 

H. punctatir (Fabricius)t 

opimus (LeOonte) 

H. quadricollis (LeConte) 

H. rumicis (Linn6)+ 

H. seriata (Mannerheim) 

H. trivittata (Say) 

setigera (LeConte) 

Genus CLtr.ONIS Dejean 

Cleont..s Sch

Curculionidae 

Genus LIXUS Fabricius 

L. caudifer LeConte 

L. concavus Say 

coloradensis Chittenden 

crassulus Notman 

rectus LeCorte 

soror Casey 

L. laramiensis Casey 

bischoffi Fall 

L. macer LeOonte 

blatchleyi Csiki 

capitatus Chittenden 

cauicollis Blatchley 

nouellus Blatchley 

L. mucidus LeConte 

morulus Blatchley 

L. parcus LeConte 

pygmaeus Casey 

L. punctinasus LeOonte 

albisetiger Chittenden 

ele phantu lus Chittenden 

planicol I is Chittenden 

L. rubellus Randall 

auctus LeConte 

L. terminalis LeConte 

blakeae Chittenden 

Tribe Lixini 

-:"--MBoNpe- 

AB 

AB 

-oN 

ON PQ 

ON 


AB_MBON 

Tlibe Rhinocyllini 

Genus RHINOCYLLUS Germar 

R. conicus (Frolich)f 

_SK_ONPQ 

Genus EMPHYASTES Mannerheim 

E. fucicola Mannerheirn 

AK 

Subfamily MOLYTINAE 

Tlibe Emphyastini 

BC 

Genus THALASSELEPHAS Egorov & Korotyaev 

Neophycocoetes O'Brien & Wibmer 

Phycocoetes LeConte 

T. testaceus (LeContet - BC 

Genus LEPYRUS Germar 

Key to North American species: Van Dyke (1928b) 

L. capucinus (Schaller) 

L. gemellus gemellus Kirby 

L. Iabradorensis Blair 

L. nordenskioeldi alternans Casey 

L. nordenskioeldi caesius Csiki 

cinereus Van Dyke 

L. nordenskioeldi canadensis Casey 

L. oregonus oregonus Casey 

perforatus Casey 

L. oregonus tessellatus Van Dyke 

L. palustris (Scopoli)x 

colon (Lint6) 

geminatus Say 

Tlibe Hylobiini 

NT 

AK YK NT BC 

NT 

AK YK 

AI{ 

AK 

NT BC AB SK 

_BC 

_BCAB 

NTBCAB_MBONPQ 

Genus IfYLOBIUS Germar 

Hypomolyx LeConte 

Key to North American species: Millers, Benjamin, and Warner (1963) 

PQ 

- - is- 

LB 

is- 

337

Curculioni*1ae 

H. congener Dalla Torre, 

Schenkling & Marshall 

confusus Ilttby 

H. pales (Ilerbst) 

macelous (Germar) 

H. pinicola (Couper) 

H. radicis Iluchanan 

H. warreni Wood 


_YKNTBC 

-BC 

AB 

AB 

AB 

SK 

oN PQ 

MB ON PQ 

MB ON PQ 

MB ON PQ 

NB 

NB 

NS 

NS 

NS 

NS 

LB 

:" 

NF 

NF 

NF 

Genus PACIIfYLOBIUS LeConte 

P. picivoru:; (Germar) 

stupitius (Boheman) 

ON 

LB 

Genus STEREMNIUS Schcinherr 

Parap,t,inthus Faust 

S. carinatus (Boheman) 

S. tuberosus Gyllenhal 

s crob ic ulatus (Mannerheim) 

AK 

AK 

Tribe Plinthini 

BC 

BC 

Genus STFI EREUS Motschulsky 

Lobosuma Buchanan 

P hilo s lrat u s Zimmerman 

Key to Nrrth American species: Zimmerman (1g64) 

S. horridus (Mannerheim) AK 

S. multitutrerculatus Buchanan AK 

S. ptinoides; (Germar)x AK 

boreaL,s Motschulsky 

fas cic u.Iatu s Motschulsky 

S. quadritu.berculatus Motschulsky AK 

_BC 

_BC 

_BCNS 

BC 

SK 

I\TIT 

Genus GASTROTAPHRUS Buchanan 

G. barberi ]luchanan 

BC 

Genus TRACHODES Germar 

T. hispidus (Linn6)t 

Subfamily ACICNEMIDINAE 

NF 

Genus PISIIODES Germar 

P. affinis Rrrndall 

czrriel Hopkins 

P. costatus Mannerheim 

P. fasciatus LeConte 

P. fiskei Hopkins 

P. nemorensis Germar 

approrimatus Hopkins 

canadensis Hopkins 

deodarae Hopkins 

P. radiatae Hopkins 

P. rotundatus LeConte 

alascensis Hopkins 

nigrae Hopktns 

P. schwarzi Hopkins 

yosemite Hopkins 

P. similis Hopkins 

utahensis Hopkins 

P. striatulus (Fabricius) 

dubius Ratdall 

froserl Hopkins 

prperi Hopkins 

P. strobi (Peck) 

AK 

AK YK 

Subfamily PISSODINAE 

_NTBCAB 

BC 

BC 

5_:cysK 

-BC 

_BCAB_MBONPQ 

BC 

BC 

BC 

AB 

ON 

PQ 

rtlB 

NS 

rasoNFeNs---- 


-PQ 

ON PQ 

olu Ns - 

_NS 

NB NS 

BC AB SK MB ON PQ NB NS NF 

NF 

NF 

NF 

338

Curculionidae 

engelmanni Hopkins 

sitchensis Hopkins 

P. terminalis Hopping 

BC 

Genus DORYIOMUS Germar 

Alycod,es Dietz 

Key to North American species: O'Brien (19706) 

D. brevicollis LeConte 

dubius (Dietz) 

fuscipes Casey 

indifferens Casey 

D. frostii Blatchley 

D. hirtus LeConte 

columbianus Sleeper 

hispidus LeConte 

nubeculinus Casey 

D. imbecillus Faust 

alaskanus Casey 

subsimilis Blatchley 

D. inaequalis Casey 

D. laticollis LeConte 

D. lecontei O'Brien 

D. leucophyllus (Motschulsky) 

adustus Motschulsky 

bajulus Farst 

longulus LeConte 

s ub sig natus (Mannerheim) 

te s sel latu s Motschulskv 

D, luridus (Mannerheim) 

cuneatulus Casey 

D. mannerheimi (Gemminger) 

uestitus (Mannerheim) 

D. marmoreus Casey 

D. mucidus (Say) 

D. parvicollis Casey 

amplus Casey 

D. rufulus (Mannerheim) 

D. squamosus LeConte 

angustatus (Dietz) 

tesselatus (WaIsh) 

D. vagenotatus Casey 

Genus ERIRHINUS Schdnherr 

E. aethiops (Fabricius)* 

morio Mannerheim 

Subfamily ERIRHININAE 

TYibe Erirhinini 

YK 

AK YK NT 

-BCAB 

AK 

AK 

AK YK NT BC 

AK YK 

AK 

v_ 

AK YK 

_1 

-YK 

AK YK 


-BCAB 

-ec 

YBC 

NT BC 

NT BC 

AB 

BC AB 

NT BC AB 

NT BC AB 

AB 

AB 

AB 

AB 

AB 

SK MB 

sK Yu 

oN 

SK 

SK 

SK 

AB SK 

AB SK 

mn 

MB 

ON PQ 

oN PQ 

_PQ 

PQ 

-PQ 

ON PQ 

_PQ 

oN PQ 

oN PQ 

MB ON PQ 

MB ON PQ 

_NS 

_NS 

NB_NF 

NS 

NS 

NF 

Genus GRYPUS Germar 

Grypidius Schiinherr 

Key to species: Cawthra (1957) 

G. brunnirostris (Fabricius) 

G. equiseti (Fabricius)* 

G. Ieechi (Cawthra) 

Genus NOTARIS Germar 

Key to North American species: Buchanan (1927) 

N. bimaculatus (Fabricius) 

wyoming ensis Chittenden 

N. flavipilosus Chittenden 

N. puncticollis (LeConte) 


-BCAB 

_YKNTBCABSKMBON 

ON 

SK MB ON 

SK MB ON 

T- NT BC AB SK MB ON 

PQ 

PQ 

PQ NB NS 

pe 

NS 

NF 

NT 

Genus PROCAS Stephens 

P. lecontei Bedel 

YK NT MB ON PQ 

339

Curculioniclae 

Genus SMICRONYX Schonhen 

Key to North American species: Arderson (1962) 

(Subgenus DESMORIS LeConte) 

Tlibe Smicronychini 

S. fulvus LrrConte AB SK - ON 

S. humilis rDietzr - SK MB 

S. incertus tDietzt MB 

rustict',s Dietz 

uitiostt:.s Dietz 

S. pinguis Jllatchley - ON 

S. sordidus LeConte MB 

montazus (Dietz) 

(Subgenus .PACFfYPHANES Dietz) 

(P ac h5, ty c hius LeConte) 

S. amoenus (Say) - SK MB 

S. centralis (Dietz t AB 

lateralis (Dietz) 

S. discoideus {LeConte) - SK MB ON 

carus lDietz) 

(Subgenus PSEUDOSMICRONYX Dietz) 

(Syner:ha Dietz) 

S. commixt rs Dietz AB SK MB 

coseyi Blatchley 

S. corniculatus (Fahraeusl - 

columl:'ianus Dietz 

ON PQ 

conniuens Casey 

fallacir>sus l{]ima 

fallar Dietz 

laungi;zosus Dietz 

spurcus Casey 

squam'.tlatus LeConte 

S. flavicans LeConte 

dietzil1ima 

- ON 

macuh:tus Dietz 

nebulosus Dietz 

ornatil:tennis Dietz 

S. squalidur Casey - ON 

cinerascens Dietz 

morio l)ietz 

S. tessellatrLs Dietz MB ON 

im p r essiros t ris Dietz 

S. utilis Bu

Curculionidae 

Genus BAGOUS Germar 

Key to North American species: Tanner (1943) 

B. americanus LeConte 

B. aLratus Blatchley 

B. bituberosus LeConte 

B. blanchardi Blatchley 

B. magister LeConte 

B. mamillatus Say 

B. nebulosus LeConte 

ochraceus Blatchley 

B. obliquus LeConte 

B. planatus LeConte 

B. restrictus LeConte 

B. tanneri O'Brien 

longirostrus Tanner 

B. transversus LeConte 

carinatus Blatchley 

Genus LISSORHOPTRUS LeConte 

Key to species: Kuschel (1951) 

L. oryzophilus Kuschel 

L. simplex tSayt 

egezus (Gyllenhal) 

Genus BRACHYBAMUS Germar 

B. electus Germar 

inceratus Boheman 

Genus NOTIODES Schtinherr 

Endalus Castelnau 

Notionomus Erichsorr 

N otiop hilus Schtinherr 

Key to North American species: Tanner (1943) 

N. aeratus (LeConte) 

N. limatulus tGyllenhal t 

N. ovalis iLeConte) 

N. punctatus (LeConte) 

Genus ONYCHYLIS LeConte 

Key to North American species: Burke (1961) 

O. nigri rostris (Boheman t 

--ec-- 

Tribe Bagoini 

_ _ _ _ 3N;:_ _ _ _ _ 

ON 

ON 

_ABONPQ 

BCABSK_NB 

BC AB SK 

_AB 

_ABONPQ 

_ONPQ 

_ONPQ 

-ONPQ 

-oN 

_ .AB 

_ABONPQ 

-ABSKMBONPQ 

BCAB_MB 

MB ON PQ 

_ONPQ 

PQ 

NB 

Genus PHYLLOTROX Schrinherr 

Euclyptus Dietz 

P. ferrugineus LeConte 

testaceus (Dietz) 

P. nubifer LeConte 

fuluipennis Sleeper 

P. rutilus (FaIl) 

Tribe Derelomini 

_BCAB_MB 

_BC 

-oN 

Genus ACAI-YPTUS Sch

Curculionitiae 

M. myrmex (Herbst) 

americ tnus (Chevrolat) 

herbstti Stunn 

myrmecodes (Say) 

PQ 

Subfamily MAGDALINAE 

Tribe Magdalini 

Genus MA(I.DALIS Germar 

M. aenescerLs LeConte 

M. alutacea LeConte 

M. armicoll s (Say) 

pallida (Say) 

M. austera austera Fall 

M. austera substriga Fall 

M. barbita (Say) 

M. cuneiforrnis Horn 

M. gentilis LeConte 

M. gracilis (LeConte) 

M. hispoides LeConte 

M. imbellis (LeConte) 

M. inconspir:ua Hor-rr 

M. lecontei ,lecepta Sleeper 

M. olyra (Horbst) 

brunnices (Gyllenhal) 

M. pandura (Say) 

M. perforatr,r Horn 

M. piceae B rchanan 

M. salicis Hcrn 

M. subtincta LeConte 

AK 

AK YK NT 

YK 

YK 

-BCAB 

BC AB 

BC AB 

BC 

BC AB 

BC 

BC AB 

BC AB 

BC 

SK 

SK 

sx 

-oN 

-oN 

MB ON 

-oN 

-oN 

MB ON 

-oN 

-oN 

-oN 

PQ 

PQ 

PQ 

PQ 

PQ 

pe 

PQ 

pe 

PQ 

PQ 

PQ 

PQ 

NF 

Tribe Laemosaccinr 

Genus LAE.MOSACCUS Schainherr 

L. nephele (Flerbst) 

plagiatus (Fabricius) 

ON 

Subfamily CURCTILIONINAE 

Genus CURCULIO Linn6 

Balani:tus Germar 

Key to Nc,rth American species: 

C. aurivestis Chittenden 

b reu inti. s us Chittenden 

C. caryae (lliorn) 

C. caryatrypes (Boheman) 

cylindricollis (Casey) 

hariolus (Casey) 

picus (Boheman) 

C. confusor,.Hamilton) 

C. iowensis lOasey) 

e*ilis Chittenden 

funicul aris Chittenden 

i6ls Chittenden 

nume n;',u s Chittenderr 

C. nasicus (fjay) 

auctus (Casey) 

ordinatus (Casey) 

C. obtusus (Bianchard) 

neocorylus Glbson 

C. pardalis (Chittenden) 

appalachius (Casey 

m ul t ifa sciatu s Chittenden 

paruicollis (Casey) 

C. proboscideus Fabricius 

cuneatus (Casey) 

nasutus (Say) 

Gibson (1969) 

BC 

_ONPQ 

-oN 

ON 

SKMBONPQ_NS 

ON 

MB ON 

MB ON 

ON 

PQ 

PQ 

PQ 

NB NS 

PE 

342

Curculionidae 

quercus (I{orrr) 

r ectiro str is ( Gyllenhal ) 

rectus (Say) 

rostrdtus (Gyllenhal) 

sparsellus (Casey) 

C. sayi (Gyllenhal) 

acuminatus (Casey) 

algonquinus (Casey) 

auriger (Casey) 

macilentus (Casey) 

mollis (Casey) 

perexilis (Casey) 

setosicornis (Casey) 

sparsus (Gyllenhai) 

strigosus (Casey) 

C. strictus (Casey) 

longipes (Casey) 

tubulatus (Casey) 

utensis (Casey) 

uirginicus (Casey) 

C. sulcatulus (Casey) 

baculi (Chittenden) 

curtus (Chittenden) 

striatus Chittenden 

C. uniformis (LeConte) 

occidentis (Casey) 

BC 

_ONPQ 

MB ON 

ON 

PQ 

NS 

Genus MACRORHOPTUS LeConte 

Key to North American species: Sleeper (1957b) 

M. hispidus Dietz 

M. sidalceae Sleeper 

Subfamily ANTHONOMINAE 

nc 

v 

SK 

Genus ANTHONOMOPSIS Dietz 

A. mixta (LeConte) 

Genus CHELONYCHUS Dietz 

C. Iongipes Dietz 

Tlibe Anthonomini 

_BCABSKMB 

_ONPQ 

Genus ANTHONOMUS Germar 

Anthomorphus Weise 

Cnemocyllus Dietz 

Paranthonomus Dietz 

Serarthrus Blatchley 

Tachypterellus Fall & Cockerell 

Tachypterus Dietz 

A. confusus Dietz 

A. consimilis Dietz 

A. consors (Dietz) 

A. convictus Gates 

A. corvulus LeConte 

A. deceptus Sleeper 

A. elongatus LeConte 

A. haematopus Boheman 

sycophantaWalsh 

A. hirtus Leconte 

A. interstitialis Dietz 

A. lecontei Burke 

uariegatus LeConte 

A. melancholicus Dietz 

murinus Dietz 

A. molochinus Dietz 

A. musculus Say 

sulcifrons LeConte 

A, nebulosus LeConte 

_BCAB 

-BC 

- - - ncAB- me3N;3Ns 

-BC 

_BCABSKMBONPQ 

-oNPQ 

-BC 

-ONPQNB 

-BCABSKMBONPQ 

-BC 

-BC-ONPQ 

_ONPQ 

_ONPQ 

PQ 

NS 

NS 

NS 

NF 

343

Curculionidae 

A. nigrinus.Boheman 

A. ochreopiltsus Dietz 

A. pictus Blatchley 

A. profundus LeConte 

uulpinz,:s Dietz 

A. quadrigil:bus (Say) 

cerasi (List) 

magna (List) 

A. quesnelensis Sleeper 

A. robustuh:.s LeConte 

A. rubidus L,eConte 

A. rubricus lJchenkling & Marshall 

rubellus Dietz 

A. rufrpes LeConte 

A. rutilus (Boheman) 

nubiloides FalI 

A. signatus lJay 

bisignalus Gyllenhal 

pallidus Dietz 

scut eI lo. t u s Gyllenhal 

A. simiolus Illatchley 

A. sphaeralciae Fall 

A. squamosr.s LeConte 

A. squamulatus Dietz 

A. subfascia;us LeConte 

latiusct;.Ius Dietz 

rantus Blatchley 

A. suturalis LeConte 

erythropterus Say 

flauicornis Boheman 

hicoriat: Pierce 

uespertinus Dietz 

A. tectus Le(lonte 

A. xanthocnr'mus Dietz 

NT 

_BC 

BC 

AB 

BC AB 

_ABSK 

BC 

BC 

AB 

SK 

MB 

-oN 

MB 

ON 

ON 

PQ 

PQ 

PQ 

PQ 

Dr\ rq 

PQ 

NB 

BC AB SK MB ON PQ NS PE 

---AB-l4e 

_ _ BCTSKMB 

BC 

SK MB 

ON 

ON 

ON 

PQ 

PQ 

NB 

NS 

NF 

Genus COC()OTORUS LeConte 

Key to No:th American species: Brown (1966) 

C. pumilae r:rajor (W.J. Brown) 

C. pumilae pumilae (W.J. Brown) 

C. scutellari,; ( LeconteJ 

prunicida (Walsh) 

_MB 

_MB 

ON 

PQ 

Genus DIETZIANUS Sleeper 

Xanthus Dietz 

D. pygmaeus. tDietzt 

MB 

Genus PSEI.IDANTHONOMUS Dietz 

Key to No;th American species: Clark (1987) 

P. crataegi (Walsh) 

brunneus Dietz 

longulus Dietz 

paruulus Dietz 

p un ct icol I is Blatchley 

P. helvolus (13oheman) 

hamamelidis Pie'rce 

incipiens Dietz 

relictus Dietz 

P. rufulus Dietz 

P. seriesetosus Dietz 

P. validus Dietz 

facetus Dietz 

YT 

Tribe Bradybatini 


ON 

_MB-PQ 

oNPQ-NS 

BCAB-MBONPQNBNS 

NS 

344

Curculionidae 

Subfamily RFIYNCIIAENINAE 

Key to North American species: Anderson (1989) 

Genus RHYNCHAENUS Clairville 

Orchestes llTiger 

R. griseus Sleeper 

R. mixtus (Blatchley) 

R. pallicornis (Say) 

puberulus (Boheman) 

uniformis (W.J. Brown) 

R. testaceus (O.F. Mriller)x 

canus (Horn) 

pallidior (Leng) 

_BC 

_BCABSKMBONPQNBNSPE 

_BCAB_MBONPQNBNSNF 

AK BC AB SK MB ON PQ NB NS NF 

Genus TACHYERGES Schtinherr 

T. ephippiatus (Say) 

T. niger (Horn) 

illinoisensis (Fall) 

paruicollis (LeConte) 

T. salicis (Linn6)x 

subhirtus (Horn) 

Genus ISOCHNUS C.G. Thomson 

L arcticus (Korotyaev)x 

L flagellum (Erichson)* 

I. rufrpes (LeConte) 

minutus (Horr') 

I. populicola (Silfverberg)J 

-BCABSKMBONPQNB 


NTBCASSKMBONPQ-NSNF 

AK_NT 

AK YK NT 

-BCAB-MBONPQ-NSNF 

_ONPQNBNS 

LB 

Subfamily PRIONOMERINAE 

Tlibe Prionomerini 

Genus ODONTOPUS Say 

O. calceatus tSay) - ON PQ 

c ar bonar iu s ( Gyllenhal) 

Tribe Piazorhinini 

Genus PIAZORHINUS Schdnherr 

P. pictus LeOonte 

P. scutellaris (Say) 

MB ON PQ 

MBONPQ_NS 

Genus ELLESCHUS Dejean 

Alyca LeConte 

Subfamilv TYCHIINAE 

Tribe-nlleschini 

E. bipunctatus (Linn6) MB ON PQ 

E. borealis Carr AK YK AB - MB 

E. ephippiatus (Say) - YK - BC AB SK MB ON PQ NF 

rufus (Say) 

E. scanicus (Paykullt AB SK - ON PQ 

Genus PROCTORUS LeConte 

Encalus LeConte 

P. armatus LeConte AK - NT BC AB SK MB ON PQ NB 

P. decipiens (LeConte) AK YK NT BC AB SK MB ON PQ NB NS 

Genus LIGNYODES Dejean 

Key to North American species: Ciark (1980) 

Tribe Lignyodini 

Key to genera: Clark (1982) 

(Subgenus LIGNYODES s. str.) 

(T hy sanocnemis LeConte) 

345

Curculionid:re 

L. bischoffi tBlatchley) ON PQ 

L. fraxini tl,eContet - SK - ON PQ 

L. helvolus tl.eConte) MB ON PO 

b alaninoide s ( Schaeffer) 

breuis (Casey) 

caseyi (IAima) 

L. horridulus (Casey) 

punctatus (Casey) 

- ON PQ 

squamtger (Casey) 

L. ocularis ('laseyt - ON 

Genus PLO(IETES LeConte 

Hamaba Casey 

Rosella Whitehead 

Key to spercies: Clark (1982) 

P. ulmi LeConte - ON PQ 

Genus SIBINIA Germar 

Key to New World species: Clark (1978) 

Tribe Tychiini 

S. maculata (LeConte) AB 

maculilbr (Hatch) 

Genus TYCIIIUS Germar 

Miccotrcgus Schdnherr 

Paraty

Curculionidae 

Genus MIARUS Schtinherr 

Cleopomiarus Pierce 

M. hispidulus LeConte 

consuetus Casey 

illini Casey 

nanus Casey 

puritanus Casey 

TYibe Mecinini 

MB ON PQ 

Genus MECINUS Germar 

M. pyraster (Herbst)t - BC 

Genus RHYSSOMATUS Schdnherr 

R. aequalis Horn 

R. Iineaticollis (Say) 

Subfamily CRYPTORITYNCHINAE 

Tribe Ithyporini 

ON 

oN PQ 

Genus CONOTRACHELUS Dejean 

Loceptes Casey 

Key to North American species: Schoof ( 1942) 

C. anaglypticus (Say) 

C. aratus (Germar) 

C. corni W.J. Brown 

C. crataegi Walsh 

C. elegans (Say) 

C. geminatus LeConte 

C. juglandis LeConte 

C. nenuphar (Herbst) 

orgula (Fabricius) 

cerasi (Peck) 

C. nivosus LeOonte 

plagiatus LeConte 

C. posticatus Boheman 

C. recessus (Casey) 

atokanus Fall 

C. schoofr Papp 

tibialis Schoof 

C. seniculus LeConte 

BC 

-AB 

-oNPQ 

-oN 

-oN 

-ONPQ 

ON PQ 

_PQ 

-MBONPQ ON PQ 

ON PQ 

_PQ 

oN PQ 

-MBONPQ 

NS 

-NF 

Genus PHELOCONUS Roelofs 

P. cribricollis (Say) 

puncticollis (Walsh) 

-oN 

Genus MICROIIYUS LeConte 

Echinaspis Blatchley 

M. setiger LeConte 

wolcotti (Blatchley) 

ON 

Genus GERSTAECKERIA Champion 

Opuntiaphila Pietce 

Philopuntia Pietce 

Key to North American species: O'Brien (1970a) 

G. basahs (LeConte) 

Tlibe Cryptorhynchini 

AB 

Genus ACALLES Schdnherr 

A. carinatus LeConte 

Genus TYLODERMA Say 

Analcis Schdnherr 

Key to North American species: Wibmer (1981) 

T. aereum (Say) 

ON 

ON PQ 

347

Curculionidae 

T. capitale \Vibmer 

T. foveolaLum tSayt 

T. minimun Blatchley 

T. nigrum Casey 

asclepiusae Sleeper 

T. oenotherare Wibmer 

Genus CRYTTORHYNCHUS Illiger 

C ry p to r hy nc hidiu s Pierce 

C. fuscatus .-eConte 

C. lapathi tl,inn6tx 

Genus APTINROMECHUS Faust 

Key to Nc,rth American species: Whitehead (19Tg) 

A. ferratus (Say) 

Genus COPIIES Champion 

Key to North American species: Sleeper ( 1955) 

C. fallax (LeConte) 

C. obtentus,'Herbst) 

ypsilon (Boheman) 

Genus EUBULUS Kirsch 

Key to species: Fiedler (1939) 

E. bisignatu; rSayr 

o b liq ue,hs ciatzs (Boheman) 

E. parochus (Herbstr 

(.?) luctt:osus (Boheman) 

BC 

AB 

AB SK MB 

ru 

_MB 

-oN 

-oN 

-oNPQ 

NB 

ON PQ 

ONPQNBNSPE_NF 

ON 

ON 

ON 

ON PQ 

oN PQ 

Genus CYLI NDROCOPTURUS Heller 

Copturt'des Casey 

Crjtrotus Casey 

C. deleoni Brrchanan 

C. longulus (LeConte) 

disperst:.ts (Casey) 

mucidu; (Casey) 

nubilat:.ts (Casey) 

obscure,'.lus (Casey) 

subcupreus (Casey) 

C. medicatusr Carr 

Subfamily ZYGOPINAE 

Key to North American genera: Sleeper (1963) 

Tribe Zygopini 

BC 

AB 

ON 

Tlibe Lechriopini 

Genus LECHRIOPS Schdnherr 

Gelus Casey 


L. californice. (LeConte) 

L. oculata (Say) 

Genus ACOf'TUS LeConte 

Homogaster Provancher 

A. suturalis i,eConte 

q ue be cens is (Provancher) 

Sleeper (1963) 

- :"- - 

^ne 

oll 

ON 

PQ 

PQ NB NS 

NS--- 

Genus PSOMUS Casey 

P. armatus (.r)ietz) 

politus ()asey 

ON PQ 

348

Curcul.ionidae 

Genus MONONYCHUS Germar 

M. vulpeculus (Fabricius) 

Subfamily CEUTORIIYNCHINAE 

Tlibe Mononychini 

ON PQ 

Tlibe Cnemogonini 

Key to genera: Colonnelli (1979) 

Genus CRAPONIUS LeConte 

C. inaequalis (Say) 

Genus CNEMOGONUS LeConte 

C. iecontei Dietz AK YK NT BC AB 

Genus ACANTHOSCELIDIUS Dalla Torre & Hustache 

Acanthoscelis Dietz 

A. acephalus (Say) 

leprosus (Boheman) 

- BC AB SK 

s ub uliro str i s (Gyllenhal) 

A. curtus tSay) 

ON 

MB ON PQ NB 

MB ON PQ 

PQ 

PE 

Genus AULEUTES Dietz 

A. epilobii (Paykull)* - YK NT BC AB SK 

cruralis (LeConte) 

A. nebulosus (LeConte) - BC - SK 

A. tenuipes (Leconte) 

MB ON PQ 

MB ON PQ 

-oNPQ 

NB 

NB 

NS 

Genus PERIGASTER Dietz 

Key to North American species: Buchanan (1931) 

P. cretura (Herbst) 

quadrispinosa (Say) 

P. liturata (Dietz) 

longirostris Buchanan 

oN PQ 

oN PQ 

NF 

Genus DIETZELLA Champion 

Coelogaster Schdnherr 

D. zimmermanni (Gyllenhai) - BC 

ON PQ 

NF 

Tribe Ceutorhynchini 

Genus CEUTORIIYNCHUS Germar 

C euthorhy nchidius J acqrelin du Val 

C. americanus Buchanan 

C. assimilis (Paykull)f 

C. bolteri Dietz 

hirticollis Dietz 

C. carteri W.J. Brown 

C. convexipennis Fall 

C. cupreus Hatch 

C. dubitans W.J. Brown 

C. echinatus Fall 

C. erysimi (Fabricius)f 

C. floralis (Paykull)f 

puberulus LeConte 

sep tent r ionalis Gyllenhal 

C. fulvotertius FaII 

C. handlordi W.J. Brown 

C. hearnei W.J. Brown 

C. invisus Fall 

C. lecontei Colonnelli 

conuexicollis LeConte 

C. munki W.J. Brown 

C. neglectus Blatchley 

C. omissus Fall 

C. opertus W.J. Brown 

AK 

TY 

)' 

T_ 

NT BC 

-sc 

BC 

BC 

BC 

sc 

BC 

18:"MB 

AB 

_SK 

T_ _ 

f 

AB 

SK 

s^ 

sc AB sx 

_ABSK 

BC 

MB 

MB 

MB 

MB 

y" 

MB 

MB 

MB 

9* 

ON 

ON 

oN- 

PQ- 

ON PQ 

oN PQ 

PQ 

PQ NB 

NS 

NS 

NS 

I\'D 

349

Curculionidae 

C. oregonersis Dietz 

C. ovipenni; Dietz 

C. pauxillur; Dietz 

C. pervestit,us FaIl 

C. punctige r Gyllenhalt 

C. querceti lGyllenhal) 

hamiltoniDietz 

pzslo Mannerheim 

C. rapae Gyllenhalt 

affiuentus Dietz 

C. squamatus LeConte 

C. squamosulus Sleeper 

sleeperi Colonnelli 

C. subpubescens LeConte 

Genus POOPHAGUS Schttnherr 

P. sisymbri:i (Fabricius)t 

Genus AMAIUS Schrinherr 

A. scortillurn (Herbst)t 

robinsoni (Blatchley) 

Genus CAL'CSIRUS Thomson 

C. medialis (LeConte) 

C. semirufu.s (LeOonte) 

C. zimmern rannii (Gyllenhal) 

_YK-BCAB_MB 

- - - l" AB sx nre- - - - - - - 

_YK_BCABSKMBONPQNBNSNF 

AK _ NT BC AB _ MB - PQ NB NS PE _ NF 

AKYK_BCABSKMBONPQ 

_BC 

_YKNT_MB 

_ONPQ 

-BCABSKMBONPQ_NSNF 

PQ 

_ONPQ 

AB ON PQ NF 

MB ON 

Genus ALLI)SIROCAIUS Colonnelli 

A. angulatu.s (LeConte) 

A. consangu.ineus (Dietz) 

A. disturbatus (Dietz) 

Genus HAT; ROPLONTUS Thomson 

H. litura (F;rbricius)f 

Genus SIR()CALODES Voss 

S. sericans (LeOonte) 

Genus NEDYUS Schdnherr 

Cidnorhinus Thomson 

N. fl avicaudis (Boheman) 

apicalis (Dietz) 

_BCAB 

_BC 

AB SK MB 

-oN 

MB 


Genus HONIIOROSOMA Frivaldszky 

Tribe Scleropterini 

H. sulcipenrre (LeConte) - YK - SK MB ON PQ 

Genus RUTIDOSOMA Stephens 

R. decipiens (LeConte) AK YK - BC AB SK MB ON PQ 

Genus ACA.LLODES LeConte 

A. saltoides Dietz PQ 

A. ventricosus LeConte MB ON 

Genus PIIYIOBIUS Schdnherr 

Hydaticus Schdnherr 

Li todacty I us Redtenbacher 

Parenthis Dietz 

Tlibe Phytobiini 

Key to New World genera: Colonnelli (1980) 

Key to genera: Colonnelli (1986) 

350

Curculionidae 

P. leucogaster (Marsham) 

griseomicans Schwarz 

my r iop hy I li (Gyllenhal) 

P. vestitus (Dietz) 

YK 

NT 

BC 

AB 

SK 

MB ON PQ 

ON 

Genus EUBRYCHIOPSIS Dietz 

E. albertana (W.J. Brown) 

NT 

BC 

AB 

SK 

Genus EUBRYCHIUS Thomson 

E. velatus (Beck) 

aquaticus Thomson 

BC 

Genus RHINONCUS Schtinherr 

C ampy lir hy nch zs Dej ean 


R. castor (Fabricius)t 

R. longulus LeConte 

R. pericarpius (Linn6)t 

umbellae (Fabricius) 

R. pyrhopus Boheman 

R. triangularis (Say) 

occidentalis Dietz 

Hoebeke & Whitehead (1980) 

_BC_ONPQ 

-BCABSKMBONPQ 

_BCABONPQ 

NTBC_SKMBONPQ 


NB 

NS 

PE 

NF 

NF 

Genus PELENOMUS Thomson 

Mecopeltus Dietz 

P. asperulus Dietz 

aeneosquamoszs (Dietz) 

P. congenialis (Dietz) 

P. fuliginosus (Dietz) 

P. gracilipes Dietz 

griseus Blatchley 

P. pusillus Dietz 

scandens (.Dietz) 

P. scoliasus (Dietz) 

P. squamipennis (Sleeper) 

P. squamosus LeConte 

im pressi uen tris Blatchley 

P. sulcicollis (Fahraeus) 

P. ventralis (Sleeper) 

\tK 

YK 

YK 

-BC 

_BC 

BC 

BC 

AB 

AB 

AB 

AB 

AB 

AB 

AB 

MB 

MB 

ON 

ON PQ 

-PQ 

_PQ 

ON 

ON 

pe- 

PQ 

NB 

Genus NEOPIIYTOBIUS Wagner 

N. cavifrons (LeConte) 

toruipes (Dietz) 

BC 

AB 

Genus BARIS Germar 

Baridius Schiinhen 

Cy p hi r h inu s Schainherr 

Orthobaris Casey 

B. canadensis Blatchley 

B. interstitialis (Say) 

agnita Casey 

anaplata Casey 

appalachia Casey 

apposita Casey 

aprica Casey 

apricoides Fall 

attonsa Casey 

austiniana Casey 

bispeculata Casey 

brachyrhina Casey 


carinulata (LeConte) 

caseyi HusLache 

cribrifera Casey 

cuneipennis Casey 

Subfamily BARIDINAE 

Tribe Baridini 

-oN 

-oNPQ 

351

Curculionidae 

demissa Casey 

dolosa Casey 

fultoni Casey 

immur,.is casey 

irrita Casey 

lauaca,za Casey 

lubrica Casey 

nephiana Casey 

oblata Casey 

oblong,tla Casey 

penins,ilae Horn 

quadr(:..ta (LeConte) 

retrusa Casey 

subaenea (LeConte) 

surrufcL Casey 

tumescens (LeConte) 

uagans Casey 

zuniana Casey 

B. lengi Hustache 

carbon,zria Blatchley 

B. stacesmit;hi Sleeper 

B. strenua (LeConte) 

c o I o r a c: ;'.e n s i s C as ey 

formali',s Casey 

longulicollis Casey 

montanica Casey 

satelles Casey 

separara Casey 

subpar'.lis Casey 

uinta (.lasey 

B. striata (Say) 

B. subsimilis Casey 

BC 

MB 

_MBON 

ON 

ON 

Genus COS.MOBARIS Casey 

C. americarra Casey 

sionilli Hayes 

squamLger Hayes 

SK 

oN PQ 

Genus AUL OBARIS LeConte 

A. pusilla (l,eConte) 

PQ 

Genus AMF'ELOGLYPIER LeConte 

A. ampeiopsis (Riley) 

ater LeConte 

ON 

Genus PSEIJDOBARIS LeConte 

P. angusta (LeConte) 

angustula LeConte 

P. nigrina (Siay) 

oN PQ 

Genus TRICIHOBARIS LeConte 

T. trinotata (Say) 

impotens Casey 

jejuniosa Casey 

plumbea (LeConte) 

ueslila (Boheman) 

ON 

Genus ORTFIORIS LeConte 

O. crotchii LeConte 

O. tenuirostris Casey 

_ _ _:"ffi_ _ _ _ _ _ _ _ _ 

Genus ONYCHOBARIS LeConte 

O. subtonsa LeConte 

O. pectorosa. LeConte 

Tlibe Madarini 

- i{B 

PQ 

352

Curculionidae 

Genus MADARELLUS Casey 

Willinkia Bondar 

M. undulatus (Say) - ON PQ 

Genus GERAEUS Pascoe 

Centrinaspis Casey 

Tribe Centrinini 

G. picumnus (Herbst) 

albotectus (Casey) 

- ON 

compactus (Casey) 

floridensis (Casey) 

lasciuus (Casey) 

oliuaceus (Gyllenhal) 

paululus (Casey) 

per scillus (Gyllenhal) 

sulor (Harris) 

Genus ODONTOCORYNUS Schdnherr 

O. denticornis (Casey) - ON 

O. scutellumalbum t$sy) - ON PQ 

Genus CENTRINOGYNA Casey 

C. canadensis Casey AB - MB 

Genus CALANDRINUS LeConte 

C. grandico)lis LeConte AB 


C. obsoletus Casey AB 

insignus Casey 

Genus CATAPASTUS Casey 

C. conspersus tl,eConte) - ON PQ 

Genus CYLINDRIDIA Casey 

C. prolixa tl-eContet MB ON PQ 

Genus DIRABIUS Casey 

(Subgenus DIRABIUS s. str.t 

D. rectirostris (LeConte) ON PQ 

tenua (Blatchley) 

Genus PROSALDIUS Oglobin 

Anacentrinus Buchanan 

P. subaequalis (Casey) MB 

Genus STETHOBARIS LeConte 

S. commixta Blatchley - ON PQ 

S. laevimargo {Championt - ON 

S. ovata (LeConte) - ON PQ 

Genus IDIOSTETHUS Casey 

l. tubulatus tSayt 

s t rig apu nc t us (Harnilton) 

- SK PQ 

Genus HAPLOSTETHOPS Casey 

H. ellipsoideus (Casey) - ON 

Genus PLOCAMUS LeConte 

Epeuchaetes Lyman 

Euchaetes LeConte 

Eunyssobia Casey 

P. echidna (LeConle) - ON 

P. hispidulus LeConte - NB 

353

Curcuiionii.ae 

Genus SPHENOPHORUS Schrtnherr 

Calandra Clairville 

Trichi:;chius LeConte 


Subfamily RIfYNCHOPHORINAE 

Tribe Sphenophorini 

S. aequalis aequalis Gyllenhal 

scirpi {Jhittenden 

- BC AB SK MB ON PQ 

uniuitia Chittenden 

S. australis australis Chittenden - ON PQ 

typhae' Chittenden 

S. callosus Oliviert - ON PQ 

jugosus Chittenden 

S. cariosus ,Oliviert NS 

cicatricosus (Say') 

flexuosus Gyllenhal 

larualis (Germar) 

sculptilis Uhler 

S. cicatristriatus Fahraeus AB SK MB 

cic at r i.,cennis Fahraeus 

ulkei llorn 

uariolc'sus LeConte 

S. costipennis Horn 

laeu ig t:.Lt u s Chittenden 

- BC AB SK MB ON PQ NB 

me do r aensis ( Satterthwait) 

S. incongruus Chittenden - ON 

elephantulus Chittenden 

S. melanoccphalus {Fabricius) - ON PQ 

nubilus Gyllenhal 

S. minimus Hart - ON 

S. mormon Chittenden - BC AB SK MB 

distich lidis Chittenden 

S. parvulus Gyllenhal - ON PQ 

pumilus Gyllenhal 

S. pertinax pertinax (Olivier) - ON - NB NS 

canaliculatus Boheman 

inter u c: I I at us Chittenden 

nasutu.s Chittenden 

penins ularis Chittenden 

sefiger Chittenden 

truncatus (Say) 

S. robustior Chittenden MB 

costifer Chittenden 

S. robustus Horn 

m uI t ih, neat us ( Satterthwait) 

- BC MB ON 

r ect i s t r iat u s Chittenden 

S. sayi Gyllenhal - BC 

s ubca r inat u s Mannerheim 

S. serratipes Chittenden AB 

S. striatipe:rnis Chittenden AB SK MB ON NS 

S. venatus ;enatus (Say) - ON PQ NB 

confusus Gyllenhal 

fallax Boheman 

immunis (Say) 

placid;ts (Say) 

S. villosiventris Chittenden PQ 

S. vomerinus LeConte AB 

baridioides llotrr 

S. zeae Walsh - ON PQ 

blatchieyi Chittenden 

Genus RHODOBAENUS LeConte 

H otnaiosty I us Chelrolat 


R. quinquepunctatus (Say) - ON PQ 

R. tredecimpunctatus tllliger) - ON PQ 

cribrarius (Fabricius) 

leptocerus (Panzer) 

q uatuor decirn punc tatus (Panzer) 

354

Curculionidae 

Genus SITOPHILUS Schonherr 

Tribe Sitophilini 

S. granarius (Linn6)f - BC MB ON PQ NF 

r emotep unctatus Gyllenhal 

S. oryzae (Linn6)t BC - ON PQ NF 

S. zeamais Motschulsky* MB ON 

Genus DIOCALANDRA Faust 

D. elongata (Roelofs)t - BC 

Genus ACAMPTUS LeConte 

A. echinus Casey 

A. rigidus LeConte 

Subfamily COSSONINAE 

Tribe Acamptini 

_ _ _ _ _ _ _ BNpe _ _ _ _ _ 

Genus HIMATIUM Wollaston 

Himatinum Cockerell 

Tfibe Cossoninr 

H. errans LeConte 

nigritulum Casey 

PQ - NS 

paruula Blatchley 

Genus COSSONUS Clairville 

Borophloeus Wollaston 

C. americanus Buchanan PQ - NS NF 

C. crenatus Horn - BC 

C. impressifrons Boheman - ON PQ 

concinnus Boheman 

C. pacificus Van Dyke - BC AB 

C. piniphilus Boheman - BC 

californicu s Motschulsky 

scrobiculatus LeConte 

C. platalea Say - ON PQ 

bohetnanniHotn 

subareatus Boheman 

C. quadricollis Van Dyke - BC AB 

Genus PENTARTHRUM Wollaston 

Tlibe Rhyncolini 

P. huttoni Wollastont PQ NF 

Genus EUOPHRYUM Broun 

E. confine (Broun)f - NF 

Genus HEXARTHRUM Wollaston 

H. thujae W.J. Brown - BC 

H' ulkei Horn - oN PQ 

Genus ELASSOPTES Horn 

E. marinus Horn - BC 

Genus STENOSCELIS Wollaston 

S. brevis (Boheman) - ON PQ 

Genus PHLOEOPHAGUS Schonherr 

P. apionides Horn - ON PQ 

P. cinadensis Van Dyke - BC AB - MB - PQ 

P. minor Horn PQ 

carinatus (Blatchley) 

355

Scolytidae 

Genus RHIINCOLUS Germar 

Xylocc.mesus Thatcher 

R. brunneu.s Mannerheim 

c alifor ni c u s Wollaston 

relictus Casey 

R. cylindricollis Wollaston 

R. knowltoni (Thatcher) 

R. macrops Buchanan 

R. oregonensis Horn 

R. pallens Oasey 

R. stacesm:thi Sleeper 

AK YK NT BC AB MB ON PQ NB NF 

-BC 

_BC 

_BC 

-BC 

_BC 

MB 

ON 

pr) 

NS 

Genus CA-I|,PHONOTUS Casey 

C. testaceus Casey 

AK 

BC 

AB 

ON 

PQ NB 

NF 

Genus DRYOPHTHORUS Germar 

D. america:rus Bedel 

corticalis Say 

Subfamily DRYOPHTHORINAE 

Tribe Dryophthorini 

MB ON PQ 

NB NS 

Family SCOLYTIDAE 

bark beetles 

J. McNamara 

In Canada and Alaska 204 species of Scolytidae 

are recorded. Most of the Canadian species attack 

forest trees, especially conifers, and may cause considerabler 

damage to timber. Both the larvae and 

adults form galleries under the bark. Some of the 

galleries may be so distinctive that the genus and, 

in some ,iases, the species that made them can be 

identifieci. 

The North American scolytids are well known 

taxonomically; Wood (1982) revised all t;he North 

American species, and Bright (1976) covered the 

Canadian and Alaskan ones. 

AK (52); YK (33); NT (31); BC (13a); AB (80); SK 

(40); MB (a7); ON (89); PQ (81); NB (53); NS (a6); 

PE (8);LB (6);NF (27) . I (5) 

Genus SCIItrRUS LeOonte 

S. annectans LeConte 

S. pubescens Swaine 

Genus HYLURGOPS LeConte 

H. porosus (LeConte) 

lecontei Swaine 

H. reticulatus Wood 

H. rugipennis pinifex (Fitch) 

H. rugipenrris rugipennis 

(Mannerheim) 

H. subcostulatus subcostulatus 

(Mannerheim) 

c r i s tat u s ( Mannerheim) 

Genus IfYLASTES Erichson 

H. gracilis LeOonte 

asper Swaine 

Iongus LeConte 

nitidus Swaine 

H. longicollis Swaine 

H. macer LeConte 

Subfamily FIYLE SININAE 

Tribe Hylastini 

AK_NTBCAB 

AKYK-BCAB 

AK 

AK 

YK 

NT 

NT 

BC 

AB 

SK 

BC 

BC AB SK 

BC AB 

BC 

BC 

_BCAB 

_BC 

ONPQ-NS 

MB ON PQ 

NB NS 

NF 

356

Scolytidae 

H. nigrinus (Mannerheim) 

yukonis Fall 

H. porculus Erichson 

canadensis Blackman 

carbonarius Fihch 

cau ernos u s ( Zimmermann) 

granosus Chapuis 

scober Swaine 

swainei Eggets 

ruebbi Swaine 

H. ruber Swaine 

H. tenuis Eichhoff 

criticus Eichhoff 

minutus Blackman 

pd.ruus Blackman 

pusillus Blackman 

AK YK 

BC 

AB 

NT BC AB 

_BC 

MB ON PQ NB 

NS 

Tribe Hylesinini 

Genus HYLASTINUS Bedel 

H. obscurus (Marsham)f 

BC 

_ONPQ 

NS 

Genus HYLESINUS Fabricius 

Leperisinus Reitter 

H. aculeatus Say 

cinereus (Swaine) 

imperialis Eichhoff 

H. californicus (Swaine) 

hoferi (B\ackman) 

H. criddiei (Swaine) 

H. fasciatus LeConte 

H. pruinosus Eichhoff 

SK 

MB ON PQ 

MB 

MB ON PQ 

-oN 

-oN 

NS 

Genus AI-NIPI{AGUS Swaine 

H y la s tinoides Spessivetsev 

A. aspericollis (LeConte) 

A. hirsutus Schedl 

AK 

_BC 

_BC 

Tribe Tomicini 

Genus PSEUDOIIYLESINUS Swaine 

P. dispar dispar Blackman 

P. dispar pullatus Blackman 

P. granulatus (LeConte) 

P. nebulosus nebulosus (LeConte) 

P. nobilis Swaine 

furnissi Blackman 

P. pini Wood 

P. sericeus (Mannerheim) 

grandis Swaine 

P. sitchensis Swaine 

P. tsugae Swaine 

heeni Blackman 

obeszs Swaine 

similis Blackman 

T 

AK 

AK 

AK 

_BC 

-BC _BC 

_BC 

-BC 

_BC 

_BC 

_BC 

_BC 

AB 

Genus HYLURGOPINUS Swaine 

H. rufrpes (Eichhoffl 

opaculus (LeConte) 

SK MB ON PQ 

NB 

NS 

Genus XYLECHINUS Chapuis 

X. americanus Blackman 

X. montanus Blackman 

Genus DENDROCTONUS Erichson 

D. bre.ricomis LeConte 

barberi Hopkins 

AK 

ecABsr-9*PQ-Tt--- 

BC 

AB 

357

Scolytidae 

D. murrayar:.ae Hopkins 

D. ponderosrLe Hopkins 

monticolae Hopkins 

D. pseudotsugae Hopkins 

D. punctatus, LeConte 

johanse.ci Swaine 

D. rufipennis (Kirby) 

borealis Hopkins 

engelmc:.nni Hopkins 

o6eszs (Mannerheim) 

piceaperda Hopkins 

similis .i,eConte 

D. simplex LeConte 

D. valens Lelonte 

AK YK 

AK YK 

AX 

_BCAB 

_BC 

_BCAB 

NT BC AB 

NT BC AB 

_ y"9*_ _ _ _ _ _ 

SK 

-oN 

MB ON 

_ NT BC AB SK MB ON PQ NB NS PE LB NF 

_NTBCAB_MBONPQNBNSNF 

PQ 

PQ 

NB 

NB NS 

NF 

NF 

Tribe Phloeotribinr 

Genus PHLOEOTRIBUS Latreille 

P. dentifrons (Blackman) 

P. lecontei Schedl 

puberulus LeConte 

P. liminaris tHarris) 

mississt pp iensis (Blackman) 

P. piceae Sweine 

---ecAB- 


SK 

_9" 

MB ON 

MB ON 

PQ NB 

PQ NB 

NS 

Genus PHL()EOSINUS Chapuis 

P. canadensis Swaine 

P. cupressi l{opkins 

nitidus Swaine 

P. dentatus ( Say) 

enieus l:llackrr.an 

graniger (EichhofD 

haagi (Eichhoff) 

P. hoferi Blar:kman 

P. hoppingi Swaine 

woodiBrig}rt 

P. keeni Blackman 

P. pini Swaine 

alaskanus Blackman 

piceae Swaine 

P. punctatus LeConte 

buckhorni Blackman 

chamberlini Blackman 

haniksu Blackman 

rubicun,lulus Swaine 

rzsli Blackman 

P. scopulorum scopulorum Swaine 

P. sequoiae llopkins 

blackmcLni Schedl 

squamo.sus Blackman 

Genus CHRAMESUS LeConte 

C. hicoriae LeConte 

lecontei (Chapuis) 

AK 

AK YK NT 

AK- 

Tribe Phloeosinini 

BC 

BC 

BC 

BC 

BC 

BC 

_BCAB 

_BC 

AB 

-oNPQNB 

ON 

MB 

PQ 

oN PQ 

Genus CHA.ETOPHLOEUS LeConte 

Pse ud oc' ry p h alus Swaine 

Renocis Casey 

C. heterodoxus (Casey) 

brittaini:. (Swaine) 

brunneu.s (Blackman) 

commix,ius (Blackman) 

criddlei (Swaine) 

/asczs (Blackman) 

Tribe Hypoborini 

BC AB SK MB 

358

Scoly'tidae 

Genus CARPHOBORUS Eichhoff 

C. andersoni Swaine 

C. carri Swaine 

C. dunni Swaine 

C. intermedius Wood 

C. ponderosae Swaine 

C. sansoni Swaine 

engelmanniWood. 

C. vandykei Bruck 

Genus POLYGRAPHUS Erichson 

Lepisotnus IKtrby 

P. convexifrons Wood 

P. rufrpennis (Kirby) 

breuicornis (Kirby) 

nigriceps (Kirby) 

saginatus Mannerheim 

AKYKNT_AB 

AKYKNT_AB 

AK 

Tbibe Polygraphini 

-BC 

BC 

- - - :"AB- 

MB 

-NS-- 

_NB 

AKYK_BCAB_PQ 


Subfamily SCOLYTINAE 

Tribe Scolytini 

Genus SCOLYTUS Geoffroy 

S. fagi Walsh 

S. laricis Blackman 

S. mali (Bechstein)t 

sulcatus LeConte 

S. monticolae Swaine 

S. multistriatus (Marsham)f 

S. muticus Say 

S. opacus Blackman 

abietis Blackman 

S. piceae (Swaine) 

S. praeceps LeConte 

S. quadrispinosus Say 

carya Riley 

coryoe Walsh 

S. rugulosus (P.W.J. Mtiller)t 

S. subscaber LeConte 

S. tsugae (Swaine) 

S. unispinosus LeOonte 

fis&ei Blackman 

sobrinus Blackman 

S. ventralis LeConte 

_ _ _:"_ _ _:x;,_ _ 

-BCAB 

_ - _:'AB_ I"3NTl"- 


___:"___oNpe__ 

-BC-oNPQ _BC 

_BCAB 

-BCAB 

BC 

NS_-_ 

_:"f" 

Genus PSEUDOTHYSANOES Blackman 

Chalcohyus Blackman 

C ry ptocle ptes Blackman 

Cryptulocleptus Wood 

P. rigidus (LeConte) 

drakeiBlackrr'an 

Genus MICRACIS LeConte 

M. swainei Blackman 

populi Swaine 

Genus HYLOCURUS Eichhoff 

M icr acisoides Blackman 

H. hirtellus LeConte 

crinitus Blackman 

Ttibe Micracini 

_BC 

_PQ 

ON 

359

Scolytidae 

Tribe Ipini 

Genus PITYOGENES Bedel 

P. carinulatus (LeConte) 

hannatus (LeConte) 

P. fossifrons (LeConte) 

P. hopkinsi Swaine 

P. plagiatus knechteli Swaine 

P. plagiatus plagiatus (LeConte) 

lecontei. Swaine 

_YK 

BC 

BC 

BC 

- SX l,fg 

AB SK 

AB SK MB 

ON 

ON 

pq 

PQ 

NB NS 

NF 

Genus PITYOKTEINES Fuchs 

Orthotomides Wood, 

P. elegans Swaine 

P. Iasiocarp'. (Swaine) 

P. minutus rSwaine) 

jasperi Swaine 

P. ornatus ( lwaine) 

P. sparsus (.LeConte) 

balsameus (LeConte) 

BC AB 

BC AB 

BC AB 

BC 

_ABSK 

MB ON PQ 

NB NS 

PE 

LB 

NF 

Genus ORTHOTOMICUS Ferrari 

O. caelatus i.EichhofD 

decretus (Eichhoff) 

Puncti7'ennis (LeConte) 

uicinus (.LeConte) 


MB ON PQ 

NB NS 

NF 

Genus IPS DeGeer 

I. borealis Swaine 

swaine. Hopping 

thomasi Hopping 

I. calligraphus (Germar) 

chloroticus Dejean 

conforntis Dejean 

eresus rlsay) 

ponderosae Swaine 

praetnorsus (Eichhoffl 

I. concinnus (Mannerheim) 

chamberlini (Swaine) 

hirsutus (Eichhoff) 

I. emarginatus (LeConte) 

I. grandicollis (Eichhofl) 

cacographus (LeConte) 

chagno;zi Swaine 

cloudcrcfti Swaine 

c r i b r icct llis ( Eichhofl) 

I. integer (E ichhofl) 

I. latidens (l.,eConte) 

guildilllackrnan 

longidens Swaine 

I. mexicanuri (Hopkins) 

radiatae Hopkins 

I. montanus (Eichhof0 

uancoutseri Swaine 

L perroti Swaine 

I. perturbatus (EichhofD 

hudsonicus (LeConte) 

interp u.nctus (EichhofD 

I. pini (Say) 

dentatus (Sturm) 

laticolli.s Swaine 

oregoni:s (Eichhoff) 

pallipes (Sturm) 

praefrictus (Eichhoff) 

rectus (LeConte') 

I. plastographus plastographus 

(LeConte) 

L tridens (Mannerheim) 

am isk iuL iensis Hopping 

AI{ YK NT BC AB SK MB ON PQ NB 

AK 

AK 

YK_ 

BC 

BC 

SK 

BC 

BC AB SK 

BC 

BC 

oN PQ 

lae oN pe 

ON PQ 




BC 

AB 

YK NT BC AB 

NS 

NS 

LB 

NF 

NF 

360

Scolytidae 

dubius Swaine 

engelmanni Swaine 

inter r u ptu s (Mannerheim) 

semirostris Hopping 

yohoensis Swaine 

I. woodi Thatcher AB 

Tribe Dryocoetini 

Genus LYMANTOR Lovendal 

L. alaskanus Wood 

L. decipiens (LeConte) 

AK 

oN pe 

Genus DRYOCOETES Eichhoff 

D. affaber (Mannerheim) 

piceoe Hopkins 

pubescens Swaine 

D. autographus (Ratzeburg) 

americanus Hopkins 

pseudotsugae Swaine 

setn ica steneus (Mannerheim) 

sep te nt r ionis ( Mannerheim) 

D. betulae Hopkins 

eichhoffi Hopkins 

I iq uidatnb ar is Hopkins 

D. caryi Hopkins 

D. confusus Swaine 

abiells Hopkins 

D. granicollis (LeConte) 

D. sechelti Swaine 

AK YK NT 

AK YK NT 

AK 

BC 

BC 

BC 

AB 

BC AB 

BC AB 

BC 

SK 

SK 

MB ON PQ 

MB ON PQ 

ON PQ 

-T 

-PQ 

NB NS 

NB NS 

NB NS 

NB 

LB 

NF 

NF 

NF 

Genus CRYPTURGUS Erichson 

C. borealis Swaine 

corrugatus Swaine 

C. pusillus (Gyllenhal)t 

dtotnus LeConte 

Tlibe Crypturgini 

AK-NTBCABSKMBONPQNBNSNF 

_SK_ONPQNBNSPE-NF 

Genus DOLURGUS Eichhoff 

D. pumilus (Mannerheim) 

AK 

BC 

Tlibe Xyloterini 

Genus XYLOTERINUS Swaine 

X. politus (Say) 

unicolor (Eichhofft 

MB ON PQ 

NB NS 

Genus TRYPODENDRON Stephens 

T. betulae Swaine 

T. lineatum (Olivier) 

biuittatum (Kirby) 

borealis Swaine 

c au ifron s (Mannerheim) 

uittigerum Eichhoff 

T. retusum (LeConte) 

T. rufrtarsis (Kirby) 

ponderosae Swaine 

NT BC AB 

AKYK_BCAB 

AKYK-BCABSKMBONPQNB 

AKYK-BCAB-MBONPQNBNS 

-MBONPQNBNSPE 


NF 

Genus XYLOSANDRUS Reitter 

X. germanus (Blandford) 

Tlibe Xyleborini 

ON 

361

Scolytidae 

Genus XYLI.BORUS Eichhoff 

Ambros;odmus Hopkins 

Anisantl.rus Ferrari 

X. affinis Eichhoff ON 

X. dispar (Fabricius) - BC - ON PQ - 

pyri (Peck) 

NS NF 

swainei (Drake) 

X. intrusus Blandford 

futcfri Hopkins 

howardt Hopkins 

- BC 

scopulorum Hopkins 

X. obesus Letlonte 

populi (Swarne) 

- ON PQ 

serratus Swaine 

X. pubescens Zimmermann - ON 

propinguus Eichhoff 

X. sal (Hopl:inst - ON PQ 

minor Swaine 

neardus Schedl 

X. xylographrs (Sayt 

canadensis Swaine 

- ON PQ 

inermis Eichhoff 

Genus XYLEBORINUS Reitter 

X. saxeseni { latzeburg) 

orbufi (llopkins) 

- BC - ON PQ 

floridem;is (Hopkins) 

Iibocedr,:. Swaine 

pecanus (Hopkins) 

quercus (Hopkins) 

lszgoe Swaine 

Genus TRYPOPHLOEUS Fairmaire 

Tlibe Cryphalini 

T. populi Hopkins - BC AB SK MB NB 

T. striatulus (Mannerheim) AK YK AB - MB - PQ - 

nitidus fiwaine 

NS NF 

punctipennis Hopkins 

T. thatcheri (Woodt - BC 

Genus PROCRYPHAIUS Hopkins 

P. mucronatrLs (LeContet AX BC AB 

idahoensis Hopkins 

populi L{opkins 

P. utahensis Hopkins 

oceris Hopkins 

AK BC AB - PQ 

solicis F.opkins 

Genus CRYPHALUS Erichson 

Taeniogi),yptes Bedel 

C. pubescens Hopkins - BC 

subconcentralis Hopkins 

C. ruficollis f-aseri Hopkins - - PQ 

balsameus Hopkins 

C. rufrcollis ruficollis Hopkins AK YK - BC AB - MB ON PQ NB NS PE - NF 

amabilis Chamberiin 

approximatus Hopkins 

canadensis Chamberlin 

coloradensis (Wood) 

grandis Chamberlin 

mainensis Blackman 

Genus HYPGTHENEMUS Westwood 

Stephanoderes Eichhoff 

H. dissimilis (Zimmermann) - ON PQ 

chapuisti (Eichhoff) 

362

Scolytidae 

H. eruditus Westwood 

asiminae Hopkins 

- ON 

citri Ebling 

euonymi Hopkins 

hamamelidis Hopkins 

juglandis Blackman 

nigripennis Hopkins 

przni Hopkins 

punctifrons Hopkins 

rumseyi Hopkins 

schwarzi (Hopkins) 

Tlibe Corthylini 

Genus PSEUDOPITYOPHTHORUS Swaine 

P. minutissimus (Zimmermann) 

P. pubipennis ( LeConte) 

sc 

_9* 

PQ 

Genus CONOPHTHORUS Hopkins 

C. banksianae McPherson 

C. coniperda (Schwarz) 

clunicus Hopkins 

taedae Hopkins 

C. ponderosae Hopkins 

contortae Hopkins 

flerilis Hopkins 

lambertianae Hopkins 

monticolae Hopkins 

scopulorum Hopkins 

C. resinosae Hopkins 

uirginianae Hopkins 

tn 

BC 

-oN 

-oN 

PQ 

oN PQ 

NB 

NS 

NB NS 

Genus PITYOPHTHORUS Eichhoff 

Key to North American species: Brieht (1981) 

(Subgenus PITYOPHTHORUS s. str.) 

(Myeleborus Blackman) 

P. absonus Blackman 

demissus Blackman 

inyoensis Big}rt 

P. alpinensis Hopping 

P. angustus Blackman 

P. aquilus Blackman 

aristatae Bright 

caelator Blackman 

P. balsaneus Blackman 

patchiBlackman 

P. bassetti Blackman 

P. biovalis Blackman 

P. boycei Swaine 

catulus (Blackman) 

iniquus (Blackman) 

siouxensis Bight 

P. briscoei Blackman 

mundus Blackman 

P. carinatus carinatus Bright 

P. carinatus monticolae Bright 

P. cariniceps LeConte 

canad,ensis Swaine 

cognatus Blackman 

P. carinulatus Swaine 

opimus Blackman 

P. cascoensis Blackman 

pilifer Schedl 

P. cavatus Bright 

P. concavus Blackman 

P. confertus Swaine 

agndtus Blackman 

burkeiBlackman 

com.Dtus Blackman 

___sc 

AK 

AK 

-NT 

YK NT 

\tK 

_BC 

NT 

BC 

sc 

? 

-AB 

BC 

AB 

AB 

AB 

AB 

AB_ 

AB SK 

__ON_NENS 

SK 

SK 

-oN 

ON 

-ONNS 

ff,fe O* 

MB 

ON 

PQ NB NS 

PQ 

PQ 

PQ 

oN PQ 

NB 

NB 

NB NS 

NB NS 

NF 

363

Scolytidae 

P. confinis LeConte BC 

P. consimilis LeConte - BC AB SK MB ON PQ - 

granulatus Swaine 

NS 

nudus Siwaine 

P. dentifrons Blackman 

P. digestus ( LeContet 

aplanatus Schedl 

- 

AB 

BC AB 

ON PQ NB NS PE - NF 

hophinsi Blackman 

idoneus Blackman 

ponderosae Blackman 

P. fuscus Bla.ckman - BC 

smithi Schedl 

P. grandis B'ackman BC 

P. hesperius Bright BC 

P. intextus Swaine - BC AB SK MB ON PQ NB NS NF 

shepardi Blackman 

lonsus lJlackman 

P. lautus Eichhoff 

natalis Blackman 

- ON PQ 

rhois Svraine 

P. murrayanae Biackman 

acceptut: Bight 

aurulentus Bight 

AK - NT BC AB - MB ON - NB 

cutleri S,waine 

depygis Blackman 

elongatt.!.s Swaine 

e:rilis Swaine 

gracilis Swaine 

tenuis Swaine 

watsoni Schedl 

P. nitidulus tMannerheim) 

atratulus (LeConte) 

AK BC AB 

punctict. llis (LeConte) 

P. nitidus Svvaine 

anceps Illackman 

aquilon,:us Bight 

AK YK NT BC AB ON PQ NB NS NF 

borealis Swaine 

uorions Schedl 

P. opaculus L,eConte 

abietis Eilacknan 

AK YK NT BC AB SK MB ON PQ NB NS NF 

albertensis Blackman 

exiguus Blackman 

pygmaetts Schedl 

P. orarius Bright BC 

P. pseudotsugae Swaine - BC 

thatcheriBight 

P. pubemlus (LeConte) - ON PQ NB NS NF 

infans Eichhoff 

P. pulchellus Eichhoff - YK NT BC AB SK MB ON PQ NB 

hirticept, LeConte 

pusio LeConte 

P. pulicarius (Zimmermann) - SK MB ON PQ NB NS 

P. ramiperda Swaine - ON PQ 

fiuazi (Blackman) 

P. recens Bri.;ht - YK BC 

P. scalptor Blackman BC 

P. scalptus Bright BC 

P. serratus Swaine BC 

P. sierrensis Bright BC 

P. toralis Wo,rd - BC AB 

collinus Bng}rt 

confusus (Bright) 

P. tuberculatus Eichhoff AK YK - BC AB 

rugicollis Swaine 

P. venustus Rlackman - BC 

artifexBlackman 

Genus GNAT'HOTRICHUS Eichhoff 

Ancyloderes Blackman 

Gnathot,"ic hoides Blackman 

364

Platypodidae 

G. materiarius (Fitch) - SK MB ON PQ NB NS 

corthyloides Eichhoff 

G. retusus (LeConte) - BC 

alni Blackman 

G. sulcatus (LeContet - BC 

aciculatus Blackman 

Genus MONARTHRUM Kirsch 

M. fasciatum (Say) - ON PQ 

gracile (Eichhoff) 

simile (Eichhofl\ 

M. mali (Firch) - oN PQ NB 

longulum Eichhoff 

M. scutellare (LeConte) - BC 

cauum (LeConte) 

obl iq uecaudatum ( Schedl) 

Genus CORTF{YLUS Erichson 

C. punctatissimus (Zimmermann) - ON PQ 

FamilyPLAIYPODIDAE pinholeborers 

J. McNamara 

One species of Platypodidae occurs in Canada. 

These beetles infest the wood of weakened trees, North American species. 

where they mine galleries. The larvae feed on the 

fungus that grows on the gallery walls. BC (1) 

Genus PLATYPUS Herbst 

P. wilsoni Swaine - BC 

Subfamily PLATYPODINAE 

TYibe Platypodini 

Wood (1979) has published a catalog of the 

365

Halictophagidae 

Order Strepsiptera twisted-winged parasites 

S.B. Peclt 

In Canada 11 species of Strepsiptera are now 

known in three of the nine families recognized 

around the world. These species are distributed in 

Southern Canada. Another 10 species occur in the 

northern tier of the United States and may eventually 

be found in Canada. Many taxa are known 

only throrrgh adult females. 

The biology of this endoparasitic group is highly 

unusual. Studies on species similar to those in 

Canada suggest the following. The frrst-instar triungulin-like 

larvae are free-living at birth but 

immedial;ely seek and parasitize immature specimens 

of lvgaeid bugs, various leafhoppers and their 

relatives, and Andreno solitary bees. The heteromorphic 

larvae mature in the host insect and 

pupate there. Adult female strepsipterans remain 

larva-like and in the host. Adult males are winged 

and live r)nly a day and an evening or two, during 

which tinre they actively search for females. After 

the female has been inseminated, eggs develop and 

hatch inside her. A thousand or more larvae may 

then leave the female, to seek new hosts and begin 

the cycle again. 

Adults may be reared from parasitized hosts. 

Males may be attracted to ultraviolet black light 

traps or may be lured by placing virgin females (in 

their hosts) in frnely screened cages. 

The systematic position of this group is becoming 

clear. It has been grouped near the similarly 

endoparasitic Rhipiphoridae in general works on 

beetles (Arnett 1968, Crowson 1981, White 1983). 

Extensive morphological studies by Kinzelbach 

(1971) and Kathirithamby (1989) suggest that 

strepsipterans are the sister group to the beetles as 

a whole. They are then ranked as a separate order 

most clearly related to the beetles (Kinzelbach 1971, 

Lawrence and Newton 1982, Kathirithamby 1989). 

Overviews of the group in North Arrrerica are 

found in Bohart (1941), Kinzelbach (1$71), and 

Luna de Carvalho and Kogan (1990). The classification 

used here follows Kathirithamby (1980). 

BC (2);AB (1); SK (3); ON (2);PQ (2);NS (1) 

Genus LOANIA Kinzelbach 

L. canadensis Kinzelbach 

Suborder Stylopidia 

Family CORIOXENIDAE 

Subfamily CORIOXENINAE 

ON 

Genus HALICTOPHAGUS Bohart 

Agalliaphagus Pterce 

Anthericomma Pierce 

Bruesia Perkins 

Cyrtocaraxenos Pierce 

Dacyrtocara Pietce 

Dioxocera Pietce 

Diozocera Pierce 

Megalechthrus Perkins 

Membracixenos Pierce 

NeocholaxPierce 

Pentac{.adocera Pierce 

Pe n log'qm mqPft i1a Pierce 

Pentaxocera Pierce 

Pentazocera Pietce 

Pentozoe Pierce 

Pseudopatella Bohaft 

Pyrilloxenos Pierce 

Family HALICTOPHAGIDAE 

Subfamily IIALICTOPHAGINAE 

366

Stylopidae 

Key to North American species: Bohart (1943) 

H. americanus Perkins 

H. bidentatus Bohart 

H. mackal Bohart 

H. omani Bohart 

_SK 

_SK 

AB 

PQ 

Genus STENOCRANOPHILUS Pierce 

Delphacircnos Pierce 

Muirixenos Pierce 

S. canadensis Kinzelbach 

SK 

Family STYLOPIDAE 

Subfamily STYLOPINAE 

Genus STYLOPS Kirby 

Katastylops Pierce 

Neostylops Pierce 

Prostylops Pierce 

S. advarians Pierce 

S. childreni Gray 

S. leechi Bohart 

S. vicinae Pierce 

-BC 

_BC 

prf 

NS 

Subfamily XENINAE 

Genus PSEUDOXENOS Saunders 

P. tigridis Pierce 

ON 

367

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396

INDEX to supraspecific taxa 

(Synonyms are in italics.) 

Abacidus LeConte 33 

Abax Bonelli 35 

Abdera Stephens 248 

Abletobium Casey 115 

Abraeinae 135 

Abraeini 135 

Abraeus Leach 135 

Abstrulia Casey 243 

Acalles Schiinherr 347 

Acalligrapha Monros 310 

Acallodes LeConte 350 

Acalymma Barber 315 

Acalyptus Schtjnherr 341 

Acamptini 355 

Acamptus LeConte 355 

Acanthocinini 296 

Acanthoderini 296 

Acanthoscelides Schilsky 301 

Acanthoscelidius Dalla Torre & Hustache 

Acanthoscelis Dietz 349 

Achenomorphus Motschulsky 113 

Acholerops Casey 83 

Acicnemidinae 338 

Acidota Stephens 89 

Aciliini 71 

Acilius Leach 71 

Acletus LeConte 212 

Aclypea Reitter 85 

Acmaeodera Eschscholtz l.6l- 

Acmaeoderinae 161 

Acmaeoderini 161 

Acmaeops LeConte 289 

Acmaeopsilla Casey 290 

Acmaeopsoides Linsley & Chemsak 293 

Acoptus LeConte 348 

Acrepis LeConte 200 

Acritini 136 

Acritus LeConte 136 

Acrolochct Thomson 88 

Acromacer Kuschel 324 

Acropteroxys Gorham 224 

Acroschatia Wilke 254 

Acrotona Thomson 100 

Acrotrichiinae 76 

Acrotrichis Motschulsky 76 

Acrulia Thomson 88 

Actedium Motschulsky 21 

Actiastes Casey 127 

Actidium Matthews 75 

Actium Casey 127 

Actizona Chandler 127 

Actobius Fauvel 117 

Acupalpus Latreille 47 

Acylamus Sharp 226 

Acylophorus Nordmann 122 

Adalia Mulsant 235 

Adelina Dejean 255 

Adephaga 7 

Aderidae 275 

Aderocharis Sharp 113 

Aderus Westwood 275 

Adimonia Laicharting 315 

Adistemia Fall 240 

Adota Casey 101 

Adoxini 309 

Adoxus Kirby 309 

Adranes LeConte 129 

Adranini 129 

Aegialia Latreille 147 

Aegialiini 146 

349 Aegialites Dejean 270 

Aegialitinae 270 

Aegomorphus Haldeman 296 

Aeletes Horn 136 

Aeolus Eschscholtz 176 

Aethiopoctines Thomson 296 

Agabetes Crotch 63 

Agabetinae 63 

Agabetini 63 

Agabini 67 

Agabinus Crotch 67 

Agabus Leach 67 

Agalliaphagus Pierce 366 

Agaporus Zimmermann 67 

Agasphaerops Horn 332 

Agathengis des Gozis 223 

Agathidini 80 

Agathidium Panzer 80 

Agleninae 270 

Aglenus Erichson 270 

Aglyptinus Cockerell 79 

Agonoderus Dejean 47 

Agonodromius Reitter 52 

Agonum Bonelli 52,53 

Agostenus MotschulskY 50 

Agrabia Casey 236 

Agrilaxia Kerremans 165 

Agrilinae 167 

Agrilini 167 

Agrilus Curtis 167 

Agriotella Brown 182 

Agriotes Eschscholtz 182 

Agriotini 182 

Agroiconota Spaeth 323 

397

Agronus Horn 332 

Agyrtecanus Reitter 77 

Agyrtes Itrdlich 77 

Agyrtidae 77 

Ahasverus des Gozis 220 

Airora Reitter 207 

Akephoms LeConte 17 

Alamona Casey 155 

Alaobia Thomson 102 

Alaus Eschscholtz 175 

Aleocharil Gravenhorst I04 

Aleocharinae 99 

Aleocharini 104 

Aleodoru,s Say 106 

Aletinus Desbrochers 327 

Alitargus Casey 242 

Allandrini 325 

Allandrus LeConte 325 

Alleculinne 259 

Alleculinii 259 

Alloeocnemis LeConte 199 

Allonychus Zaitzev 67 

AllopodaLeConte 276 

Allopodini 276 

AllorhagtumKolbe 289 

Allosirocrilus Colonnelli 350 

Alniphagus Swaine 357 

Alobates Motschulsky 26L 

Alobus LeConte 151 

Alocentron Schilsky 328 

Alphitobius Stephens 258 

Alphitophagus Stephens 255 

Altica Mtiller 317 

Alticinae 316 

Alyca Le(lonte 345 

Alycodes Dietz 339 

Amacylophorus Smetana 122 

Amalus Schdnherr 350 

Amara Bonelli 35, 38 

Amarygrrrini 259 

Amqurosromo Mtiller 299 

Amblopusa Casey 107 

Amblycerinae 300 

Amblycerus Thunberg 300 

Amblyctis LeConte 247 

Amblyderus LaFert6-S lnecldre 27 | 

Ambrosiodmus Hopkins 362 

Americanotimarcha Jolivet 310 

Americonr,asezs Csiki 34 

Amerinusr Casey 46 

Amerizus Chaudoir 28 

Amecocerus Solier 213 

Ametor Semenov 132 

Amischa Thomson 101 

Arnnesia.Florn 333 

Amniscus, Haldeman 297 

Ampedini 183 

Ampedus Dejean 183 

Ampeloglypter LeConte 352 

Amphasia Newman 45 

Amphicerus LeConte 200 

Amphichroum Kraatz 90 

Amphicrossus Erichson 216 

Amphidora Eschscholtz 257 

Amphimallon Berthold 153 

Amphivectura Blaisdell 212 

Amphizoa LeConte 61 

Amphizoidae 61 

Amydetinae 191 

Amydrogmus Pierce 334 

Anabolus Gistel 12 

Anacaena Thomson 132 

Anacentrinus Buchanan 353 

Anadaptus Casey 43 

Anaedus Blanchard 262 

Anaglyptini 286 

Anagrylius Casey 156 

Analcis Schdnherr 347 

Analeptura Linsley & Chemsak 291 

Anambodera Barr 161 

Anametis Horn 334 

Anaquedius Casey 122 

Anaspidinae 276 

Anaspidini 276 

Anaspis Geoffroy 276 

Anqstictodera Casey l2I 

Anastrangalia Casey 293 

Anatis Mulsant 235 

Anatrinodia Casey 157 

Anchastus LeConte 184 

Anchicera Thomson 223 

Anchodemus LeConte 335 

Anchomenus Bonelli 52 

Aruchus LeConte 52 

Anchytarsus Gu6rin-M6neville 169 

Ancistronycha Markel 193 

Ancylochira Eschscholtz 163 

Ancyloderes Blackman 364 

Ancylonycha Blanchard 152 

Ancyronychini I72 

Ancyronyx Erichson L72 

Ancy rop horus I{r aatz 94 

Andrector Horn 315 

Androchirus LeConte 260 

Anelaphus Linsley 281 

Anelastes Kirby 188 

Anelpistus Horn 263 

Anepsiini 253 

Anepsius LeConte 253 

Anepsiota Casey 101 

Anisa.ndrus Ferrari 362 

Anisocaluio Crotch 235 

Anisodactylus Dejean 43, 44 

Anisomera LeConte 67 

Anisorus Mulsant 288 

Anisostena Weise 321 

398

Anisosticta Dejean 233 

Anisotarsus Chaudoir 45 

Anisotoma Panzer 80 

Anisoxya Mulsant 248 

Anobiidae 201 

Anobiinae 202 

Anobiini 203 

Anobium Fabricius 203 

Anocomis Casey 283 

Anogcodes Dejean 266 

Anogdus LeConte 78 

Anomala Samouelle 154 

Anomalaegialia Brown 147 

Anomalepta Casey 154 

Anomalini 154 

Anomalonychus Saylor 264 

Anomalopus Casey 154 

Anomoea Lacordaire 304 

Anomoglossus Chaudoir 49 

Anomoia Dejean 304 

Anomophagus Reitter 218 

Anoplis Kirby 163 

Anoplitis Chapuis 321 

Anoplitis Kirby 321 

AnopliurnHaldeman 280 

Anoplotrupes Jekel 150 

Anotylus Thomson 95 

Antennalia Casey 278 

Anthaxia Eschscholtz 165 

Anthaxiini 165 

Anthericomma Pierce 366 

Antherophagus Latreille 221 

Anthicidae 271 

Anthicinae 271 

Anthicini 271 

Anthicus Paykull 271 

Anthobates LeConte 276 

Anthobatulo Strand 276 

Anthobium Leach 91 

Anthobium auct. 87 

Anthocornus Erichson 211 

Anthomarp,hus Weise 343 

Anthonominae 343 

Anthonomini 343 

Anthonomopsis Dietz 343 

Anthonomus Germar 343 

Anthophagini 89 

Anthophylax LeConte 289 

Anthrenini 198 

Anthrenus Schaeffer 198 

Anthribidae 324 

Anthribinae 324 

Anthribini 324 

Anthribulus LeConte 324 

Apator Semenov 67 

Apatura Castelnau & Gory 165 

Apelocera Chevrolat 286 

Apenes LeConte 59 

Aphanotus LeConte 258 

Aphelogenio Chaudoir 56 

Apheloplastus Brown 79 

Aphenolia Reitter 215 

Aphodiinae 146 

Aphodiini 147 

Aphodius Illiger 147 

Aphonus LeConte 156 

Aphorista Gorham 238 

Aphrastus Say 333 

Aphthona Chevrolat 317 

Apion Herbst 328 

Apionidae 328 

Apionini 328 

Aplemonini 329 

Aplocentrus LeConte 44 

Aploderus Stephens 93 

Apocellus Erichson 95 

Apodasyini 295 

Apothinus Sharp 126 

Apotomus Kirby 326 

Apristus Chaudoir 58 

Apteralium Casey 115 

Apterius Blatchley 317 

Apteroloma Hatch 77 

Apteromechus Faust 348 

Apterosoma Motschulsky 311 

Apterospasta LeConte 265 

Araecerini 324 

Araecerus Schtinherr 324 

Araeopidius Cockerell 143 

Araeopus LeConte 143 

Archostemata 6 

Arctelaphrus Semenov 16 

ArctobiaThomson 15 

Arctoclinidium Bell 7 

Arctodytes Thomson 67 

Arctolina Kontkanen 312 

Areus Casey 109 

Argaleus LeConte 288 

Argante Gistel 162 

Argutor Dejean 34 

Arhopalus Audinet-Serville 279 

Aridius Motschulsky 240 

Arpedium Erichson 89 

Arrhenodes Schiinherr 327 

Arrhenodini 327 

Arrhenopeplus Kocb. 84 

Arrhenoplita l{trby 255 

Artematopidae 174 

Arthrolips Wollaston 228 

Arthromacral{trby 262 

Artochia Casey 90 

AsaphesKirby 181 

Asaphidion des Gozis 20 

Asclera Stephens 266 

Asclerini 266 

Ascoliocerus M6quignon 178 

399

Aseminae 279 

Asemini 279 

Asemobius Horn 95 

Asemum Eschscholtz 279 

Asidini Il54 

Asidopsis Casey 254 

Asiobates Thomson 74 

Aspidius Mulsant & Rey 257 

Astenina 117 

Astenus Dejean 117 

Asteria Fauvel 113 

Astyleiopus Dillon 297 

Astylopsis Casey 297 

Asynonyt:hus Crotch 329 

Ataenius Harold 149 

Atheta Thomson 101 

Athetalia Casey 101 

Athetini 100 

Athetota Casey 101 

Athoini .176 

Atholus llhomson 139 

Athous Eschscholtz 177 

Atimia Haldeman 280 

Atimiini 280 

Atomaria Stephens 223 

Atomariirae 223 

Atomarii:ri 223 

Atrachela. Gistel 308 

Atractopterus LeConte 182 

Atranus l,eConte 52 

Atrecus Jacquelin du Val 123 

Attagenirri 197 

Attagenus Latreille 197 

Attalus Erichson 212 

Attelabidae 326 

Attelabini 326 

Attelabus;Linn6 326 

Atymius (]istel 309 

Auchenia Thunberg 304 

Augyliini 169 

AulaconArnett 175 

Auletini 327 

Auletobiu.s Desbrochers 327 

Auleutes Dietz 349 

Aulobaris LeConte 352 

Aulonium Erichson 251 

Aulonothr:oscus Horn 186 

Autalia Leach 107 

Autaliini 107 

Axinopalpus LeConte 58 

Babia Chevrolat 305 

Babiini 305 

Babnormrrdes Park 127 

Bacaniini 138 

Bacanius LeConte 138 

Bactridium LeConte 218 

Badister Clairville 49 

Baeckmanniolus Reichardt 137 

Baeocera Erichson 125 

Bagoini 341 

Bagous Germar 341 

Balaninus Germar 342 

Baliosus Weise 321 

BalophilaCrotch 317 

Baridinae 351 

Baridini 351 

Baridius Schtinherr 351 

Baris Germar 351 

Barynotini 330 

Barynotus Germar 330 

Barypeithes Jacquelin du Val & Fairmaire 

Barytachys Chaudoir 28 

Bassareus Haldeman 307 

Batrisinae 127 

Batrisini 127 

Batrisodes Reitter 127 

Batyle Thomson 287 

Batyleoma Casey 287 

Baudia Ragusa 49 

Beeria Hatch 120 

Bellamira LeConte 290 

Belonuchus Nordmann 119 

Bembidiini 20 

Bembidion Latreille 20, 27 

Bembidionetolitzkya Strand 23 

Berninelsonius Leseigneur 178 

Berosinae 131 

Berosus Leach 131 

Bessobia Thomson 102 

Bibloplectus Reitter 127 

Bidensomela Monros 310 

Bidessini 63 

Bidessonotus R6gimbart 63 

Biphyllidae 241 

Biphyllini 241 

Bitoma Herbst 251 

Bius Mulsant 258 

Blapstinus Dejean 257 

Blapylis Horn 257 

Blechrus Motschulskv 58 

Bledius Leach 93 

Blemus Dejean 19 

Blepharida Chewolat 316 

Blepharidulo Weise 316 

Blepharoplataphus Netolitzky 23 

Blethisa Bonelli 15 

Blitophaga Reitter 85 

Bolboceras Kirby 150 

Bolboceratini 150 

Bolbocerosoma Schaeffer 150 

Boliographa Motschulsky 310 

Bolitobiini 98 

Bolitobius Leach 98 

Bolitobus Tottenham 98 

Bolitochara Mannerheim 107 

330 

400

Bolitocharina I07 

Bolitophaginl 255 

Bolitophagus Illiger 255 

Bolitotherus Candbze 255 

Bomius LeConte 58 

Boreades Parsons 214 

Boreaphilus Sahlberg 89 

BoreobiaTschitsch6rine 31 

Boreocanthon Halffter 146 

Boreoglymmius Bell & BeIl 7 

Boreophilia Benick 102 

Boreosaxinis Moldenke 305 

Boreostiba Lohse 101 

Boridae 268 

Borophloeus Wollaston 355 

Boros Herbst 268 

Boschella Barr 210 

Bostrichidae 199 

Bostrichinae 200 

Bostrichini 200 

Bostrichoidea 199 

Bothrideres Erichson 252 

Bothriderinae 252 

Bothriderini 252 

Bothriopterus Chaudoir 32 

Brachiacantha Dejean 232 

Brachinini 18 

Brachinus Weber 18 

Brachybamus Germar 341 

Brachycepsis Brendel 83 

Brachycis Casey 245 

Brachycoryna Gu6rin-M6neville 

Brachycrepis LeConte 184 

Brachyderinae 329 

Brachyderini 330 

Brachygluta Thomson 128 

Brachyglutini 128 

Brachyini 168 

Brachyleptura Casey 293 

Brachylobus Chaudoir 50 

Brachymorphzs Chewolat 210 

Brachynotus Kirby 192 

Brachypnoeo Gistel 308 

Brachypterolus Grouvelle 214 

Brachypterus Kugelann 214 

Brachyrhinus Latreille 331 

Brachys Solier 168 

Brachysomida Casey 290 

Brachysomus Schijnherr 330 

Brachystilus Chaudoir 31 

Brachytarsoides Pierce 324 

Brachyusa Mulsant & Rey 100 

Bracteomimus Lindroth 21 

Bracteon Bedel 20 

Bradybatini 344 

Bradycellus Erichson 46 

Bradytus Stephens 36 

Brathinus LeConte 89 

322 

Brennus Motschulsky 13 

Brenthidae 327 

Bromius Chevrolat 309 

Brontes Fabricius 220 

Broscini 19 

Broscodera Lindroth 19 

Broscus Panzer 19 

Bruchidae 300 

Bruchidius Schilsky 301 

Bruchinae 300 

Bruchus Linn6 301 

Bruesia Perkins 366 

Brumoides Chapin 233 

Bryaxis Leach 128 

Brychius Thomson 60 

Bryonomus Casey 119 

Bryoporus lkaatz 98 

Buprestidae 160 

Buprestinae 162 

Buprestini 163 

Buprestis Linn6 163 

Buprestoidea 160 

Byrrhidae 158 

Byrrhinae 158 

Byrrhini 158 

Byrrhodes LeConte 205 

Byrrhoidea 158 

Byrrhus Linn6 158 

Byrsopages Schijmherr 335 

Bythinini 128 

Bythinopsis Raffray 128 

Byturidae 241 

Byturini 241 

Byturus Latreille 241 

CacopliaLeConte 294 

Cacoternnus LeConte 203 

Cq.docera Eschscholtz 149 

Caenia Newman 189 

Caeniella Cockerell 189 

Caenocara Thomson 204 

Caenocyrta Brown 79 

Caenoscelini 222 

Caenoscelis Thomson 222 

Caerosternus LeConte 138 

Cafius Stephens 119 

Calandra Clairville 354 

Calandrinus LeConte 353 

Calathus Bonelli 51 

Calitys Thomson 207 

Calleida Dejean 58 

Callidiini 282 

Callidium Mulsant 283 

Calligrapha Chewolat 310 

Callimellum Strand 282 

Callimoxys lkaatz 282 

Callimus Mulsant 282 

Callipara Motschulsky 10 

401

Callisten,',a Vacher de Lapouge 11 

Callisthenes Fischer von Waldheim 

Callisthiri 49 

Callitrop;r Motschulsky 10 

Calloides LeConte 285 

Callosobruchus Pic 300 

Calochara Casey 104 

Calochromus Gu6rin-M6neville 188 

Caloderma Casey 113 

Calodrepa Motschulsky 10 

Calomycterus Roelofs 334 

Calopodinae 266 

Calopterini 189 

Calopteron Gu6rin-M6neville 189 

Calopus Irabricius 266 

Calosirus Thomson 350 

Calosoma Weber 10 

Calvia Mulsant 235 

Calyrnmaderini 204 

Calymmaderus Solier 204 

Calyptomerus Redtenbacher 141 

Camptorl;.ino Kirby 151 

Campylir.hynchus Dejean 351 

Canastottr Casey 100 

Canifa LeConte 276 

Canonura Casey 296 

Cantharidae 192 

Cantharinae 192 

Cantharirri 193 

Cantharis Linn6 193 

Cantharoidea 188 

Canthon .lloffmansegg L46 

Canthosttthus Haldeman 306 

Caphalistes Motschulsky 212 

Capnochroa LeConte 260 

Capraita Bechyn6 320 

Carabidae 8 

Carabini 10 

Carabus Linn6 11 

Carcinops Marseul 138 

Carcinus.Marseul 138 

Cardiophorinae 185 

Cardiophorus Eschscholtz L85 

Carebq,ra LeConte 247 

Carpelimus Leach 94 

Carphacis des Gozis 98 

Carphoborus Eichhoff 359 

Carphonobus Casey 356 

Carpophilinae 2L4 

Carpophilus Stephens 214 

Carrhydms Fall 69 

Cartodere Thomson 240 

Cassida Linn6 322 

Cassidinae 322 

Cassidini 322 

Catapastus Casey 353 

Cateretes Herbst 214 

Cateretinae 214 

11 

Catharellus Casey 46 

Cathartosilvanus Grouvelle 220 

Catogenus Westwood 219 

Catopocerinae 80 

Catopocerus Motschulsky 80 

Catops Paykull 81 

Catoptrichus Murray 81 

Catorama Gu6rin-M6neville 205 

Cedius LeConte 129 

Celetes Newman 189 

CeliaZimmermann 36 

Celiamorphus Casey 43 

Celina Aub6 63 

Centrinapsis Casey 353 

Centrinini 353 

Centrinogyna Casey 353 

Centrodera LeConte 288 

Centronopus Solier 258 

Centuriatus Pacheco 169 

Ceophyllini 129 

Ceophyllus LeConte 129 

Cephalogyna Casey 44 

Cephaloidae 262 

Cephaloinae 262 

Cephaloon Newman 262 

Ceracis Melli6 245 

Ceralimnius Hinton 173 

Cerambycidae 277 

Cerambycinae 280 

CerandriaLucas 255 

Cerataltica Crotch 317 

Ceratapion Schilsky 328 

Ceratoderus Blanchard 271 

Ceratographis Gahan 297 

CeratomaDozier 315 

Ceratomegilla Crotch 233 

Ceratopidonia Linsley & Chemsak 293 

Cercolia Casey 293 

Cercopedius Sleeper 332 

Cercyon Leach 134 

Cercyonini 134 

Cerophorus Castelnau 216 

CerotaruaBowditch 315 

Cerotoma Chewolat 315 

Ceruchini 144 

Ceruchus MacLeay 144 

Cerylon Latreille 227 

Cerylonidae 227 

Ceryloninae 227 

Cetoniinae 157 

Cetoniini 157 

Ceutorhynchidius Jacquelin du Val 349 

Ceutorhynchinae 349 

Ceutorhynchini 349 

Ceutorhynchus Germar 349 

Chaerocassis Spaeth 323 

Chaetarthria Stephens 131 

Chaetarthriinae 131 

402

Chq.etechus Horn 332 

Chaetocnema Stephens 319 

Chaetophloeus LeConte 358 

Chaetophora Stephens 158 

Chalcoidea Motschulsky 311 

Chalcoparia Crotch 308 

Chalcophora Solier 161 

Chalcophorinae 161 

Chalcophorini 161 

Chalcohyus Blackman 359 

Chalepini 321 

Chalepus Thunberg 321 

Chapmania Bernhauer 95 

Charaphloeus Casey 219 

Charhyphus Sharp 95 

Charidotella Weise 323 

Chariessa Perty 210 

Charisalia Casey 291 

Chauliognathinae 195 

Chauliognathini 195 

Chauliognathus Hentz 195 

Chelonychus Dietz 343 

Chelymorpha Chewolat 322 

Chilocorinae 233 

Chilocorini 233 

Chilocorus Leach 233 

ChionNewman 286 

Chionotyphlus Smetana 113 

Chirida Chapuis 323 

Chitalia Sharp 106 

Chlaeniellus Reitter 50 

Chlaenius Bonelli 49, 50 

Chlamisinae 307 

ChloephagaWeise 320 

Cholevinae 81 

Cholevini 81 

Choraginae 324 

Choragini 324 

Chramesus LeConte 358 

Chromatia LeConte 259 

Chrysobothrinae 166 

Chrysobothrini 166 

Chrysobothris Eschscholtz 166 

Chrysobracteon Netolitzky 20 

Chrysochus Chevrolat 309 

Chrysodina Baly 308 

Chrysolina Motschulsky 311 

Chrysomela auct. 311 

Chrysomela Linn6 313 

Chrysomelidae 301 

Chrysomelinae 310 

Chrysomelini 310 

Chrysomeloidea 277 

Chrysomelopsilo Strand 313 

Chrysomelopsis Achard 313 

Chrysophana LeConte 161 

Chrysostigma Kirby 10 

Cicindela Linn6 8 

Cicindelidia Rivalier 10 

Cicindelini 8 

Cidnorhinus Thomson 350 

Clidae 244 

Ciinae 244 

Ciini 244 

Cilea Jacquelin du Val 97 

Cimberidini 324 

Cimberis des Gozis 324 

Cimbocera Horn 333 

Ciocotalpa Saylor 155 

Circinalia Casey 54 

CircinalidiaCasey 54 

Circinoporus Guignot 65 

Circomus Fleischer 258 

Cis Latreille 244 

Cistelct. Fabricius 259 

Clambidae 141 

Clambus Fischer von Waldheim 141 

Clamoris des Gozis 253 

Clavigerinae I29 

Clavilispinus Bernhauer 92 

Cleonidius Casey 336 

Cleoninae 336 

Cleonini 336 

Cleonis Dejean 336 

Cleonus Schdnherr 336 

Cleopomiarus Pierce 347 

Cleptelmis Sanderson 172 

Cleridae 208 

Clerinae 208 

Clerini 209 

Cleroidea 206 

Clinidium Kirby 7 

Clivina Latreille 18 

Clivinini 17 

Clusiota Casey 102 

Clytini 284 

Clytoleptus Casey 286 

Clytrinae 304 

Clytrini 304 

Clytus Laicharting 285 

Cnernidotus Erichson 61 

Cnemocyllus Dietz 343 

Cnemogonini 349 

Cnemogonus LeConte 349 

Cneoglossini 143 

Cneorhinini 330 

Coccidophilus Brethes 229 

Coccidula Kugelann 233 

Coccidulinae 233 

Coccidulini 233 

Coccinella Linn6 235 

Coccinellidae 229 

Coccinellinae 233 

Coccinellini 233 

Coccotorus LeConte 344 

Coelambus Thomson 64 

403

Coelini 254 

Coptotomini 70 

Coelocephalapion Wagner 328 

Coelocnernis Mannerheim 261 

Coelogaster Schiinherr 349 

Coelometopinae 260 

Coelometopini 260 

Coelomorpha Casey 254 

Coelostethus LeConte 203 

Coelotaxis Horn 253 

Coelus Eschscholtz 254 

Colaspis Fabricius 308 

Colastus Erichson 214 

Colenis Erichson 79 

Coleomegilla Timberlake 233 

Coleothorpa Moldenke 305 

Collapterl,'x Newman 294 

Colliuris DeGeer 56 

Collops Erichson 211 

Colon Her:bst 80 

Coloninaer 80 

Colopterus Erichson 214 

Colposura', Casey 101 

Colydiida,: 251 

Colydiinae 251 

Colydiini 251 

Colydium Fabricius 252 

Colymbet,:s Clairville 69 

Colymbet.nae 67 

Colymbet:ni 69 

Comstocle,'.a Van Dyke 52 

Conalia lVlulsant &Rey 249 

Coninomu'.s Thomson 240 

Coniocleonus Motschulsky 336 

ConiontelLus Casey 253 

Coniontides Casey 253 

Coniontini 253 

Coniontis Eschscholtz 253 

Coniophagus Mink 213 

Con iop hth:.alma Kangas 221 

Conithassa Thomson 239 

Connatichela Anderson 334 

Connophron Casey 82 

Conoderus Eschscholtz 176 

Conophthorus Hopkins 363 

Conoplectus Brendel 126 

Conotelus Erichson 214 

Conotrachelus Dejean 347 

Copelatinae 62 

Copelatini 62 

Copelatus Erichson 62 

Cophes Champion 348 

Copidita LeConte 266 

Coprini 1,46 

Copris Mtiller 146 

Coprochara Mulsant & Rey 104 

Coprophibrni 92 

Coprophilus Latreille 93 

Coproporus Kraatz gG 

Coptotomus Say 70 

Copturodes Casey 348 

Coraebini 167 

Cordalia Jacobs 105 

Cordoharpalus Hatch 41 

Coreopsomela Monros 310 

Corioxenidae 366 

Corioxeninae 366 

CorphyraSay 274 

Corthylini 363 

Corthylus Erichson 365 

Corticaria Marsham 240 

Corticariinae 240 

Corticarina Reitter 241 

Corticeus Piller & Mitterpacher 256 

Corticotomus Sharp 207 

Cortilena Motschulsky 241 

Cortinicara Johnson 241 

Cortodera Mulsant 290 

Corylophidae 228 

Corylophinae 228 

Corymbites Latreille 179 

Coryphiini 89 

Coryphium Stephens 89 

Cosmobaris Casey 352 

Cosmosalia Casey 294 

Cosnania Dejean 56 

Cossoninae 355 

Cossonini 355 

Cossonus Clairville 355 

Cotalpa Burmeister 155 

Coxelus Horn 251 

Craponius LeConte 349 

Cratacanthus Dejean 39 

Cratidus LeConte 257 

Cratomorphini 190 

Cratonychus Dejean 184 

Crerya LeConte 210 

Cremastocheilini 157 

Cremastocheilus Knoch 157 

Creniphilus Horn 132 

Crenitis Bedel 132 

CrenitulusWinters 132 

Creophilus Samouelle 120 

Crepidodera Chevrolat 319 

Crepidophorus Mulsant & Guillebean 

Crepidotritus LeConte 184 

Crine Pascoe 218 

Criocephalum Dejean 27 I 

Criocephalus Mulsant 279 

Criocerinae 304 

Criocerini 304 

Crioceris Mi.iller 304 

CrioscaphaHorn 246 

Crosimus Casey 221 

Crossidius LeConte 287 

Cryrnodes LeConte 268 

177 

404

Cryobius Chaudoir 34 

Cryocarabus Vacher de Lapouge 11 

Cryphalini 362 

Cryphalus Erichson 362 

Cryptarcha Shuckard 217 

Cryptarchinae 217 

Cryptobiina 116 

Cryptobiurn Mannerheim 116 

Cryptocephalinae 305 

Cryptocephalini 306 

Cryptocephalus Miiller 306 

Cryptocleptes Blackman 359 

Cryptohypnus Eschscholtz 17 8 

Cryptolestes Ganglbauer 220 

Cryptophagidae 221 

Cryptophaginae 221 

Cryptophagini 221 

Cry ptophagistes Crotch 221 

Cryptophagus Herbst 221 

Cryptophilinae 224 

Cryptophilini 224 

Cryptophilus Reitter 224 

Cryptopleurum Mulsant 135 

Cryptorhopalum Gu6rin-M6neville 

Cryptorhynchidius Pierce 348 

Cryptorhynchinae 347 

Cryptorhynchini 347 

Cryptorhynchus Illiger 348 

Cryptoweisea Gordon 229 

Cryptulocleplus Wood 359 

Crypturgini 361 

Crypturgus Erichson 361 

Ctenicera Latreille 179 

Ctenistini 129 

Ctenobium LeConte 202 

Ctenodactylini 56 

Ctenonychus Stephens 184 

Cucujidae 219 

Cucujinae 219 

Cucujoidea 213 

Cucujus Fabricius 219 

Cupedidae 6 

Cupedinae 6 

Cupedini 6 

Cupes Fabricius 6 

Cupila Casey 127 

Curculio Linn6 342 

Curculionidae 329 

Curculionoidea 323 

Curculioninae 342 

Curimopsis Ganglbauer 158 

Curtonotus Stephens 35 

Curvimanon Fleischer 80 

Cybister Curtis 72 

Cybistrini 72 

Cybocephalinae 2I7 

Cybocephalus Erichson 217 

Cychramus Kugelann 216 

198 

Cychrini 12 

Cychrus Fabricius 13 

Cyclinus I{trby 72 

Cycliopleurus Hope 281 

Cyclocephala Latreille 155 

Cyclocephalini 155 

Cycloneda Crotch 236 

Cyclotrachelus Chaudoir 35 

Cylindera Westwood 10 

Cylindridia Casey 353 

Cylindrocharis Casey 31 

Cylindrocopturus Heller 348 

Cylister Cooman 140 

Cylistix Marseul 140 

Cyllene Newman 284 

Cyllodes Erichson 217 

Cyllopodes Murray 2\4 

Cymatodera Gray 208 

Cymatopterus Dejean 69 

Cymbiodyta Bedel 133 

Cymindis Latreille 59 

Cynaeus LeConte 255 

Cypha Leach 108 

Cyphea Fauvel 107 

Cyphelophorus Kuwert 131 

Cyphicerini 334 

Cyphina 107 

Cyphirhinus Schiinherr 351 

Cyphomimus Horn 330 

CyphonPaykull 142 

Cyrtepistomus Marshall 334 

Cyrtinini 298 

Cyrtinus LeConte 298 

Cyrtocaraxenos Pierce 366 

Cyrtomoptera Motschulsky 193 

Cyrtomoptila Motschulsky 193 

Cyrtophorus LeConte 286 

Cyrtoscydmus Motschulsky 83 

Cyrtotriplax Crotch 225 

Cyrtusa Erichson 79 

Cytilus Erichson 160 

Dacne Latreille 225 

Dacninae 225 

Dacyrtocara Pierce 366 

Dalopius Eschscholtz 182 

Dalotia Casey 100 

Dampfius Pacheco 170 

Danae Reiche 238 

Daniela Netolitzky 23 

Danosoma Thompson 175 

Dascillidae 143 

Dascillinae 143 

Dascilloidea 143 

Dasydera LeConte 150 

Dasyglossal{raatz 99 

DasypogonLeConte 175 

Dasvrhadus Fall 213 

405

Dasy'bes Faykull 212 

Dasytinae 212 

Datomicra Mulsant & Rey 100 

Dearthrus LeConte 198 

Decarthron Brendel 128 

Dectes LeConte 297 

Deinopsini 108 

Deinopsis Matthews 108 

Deinopteroloma Jansson 91 

Deleaster Erichson 92 

Deloyala Chewolat 323 

Delphacixenos Pierce 367 

Delphastrrs Casey 229 

Delphota Jasey 103 

Deltometopus Bonvouloir 187 

Deltostethus Sharp 134 

Dendroctonus Erichson 357 

Dendroiders Latreille 269 

Dendrophagus Schrinherc 220 

Dendrophilinae 138 

Dendrophilini 138 

Dendrophilus Leach 138 

Denticollis Piller & Mitternacher 

Deratanclrus Casey 55 

Deratopeus Casey 115 

DerelomirLi 341 

Deretaphrini 252 

Deretaphrus Newman 252 

Dermestes Linn6 196. 197 

Dermestic{ae 196 

Dermestirri 196 

Dermestinus Zhantiev 196 

Dermestoidea 195 

Derocrepis Weise 318 

Derodontidae 195 

Derodontinae 196 

DerodontrLs LeConte 196 

Deronectes auct. amer. 66 

Derospidea Blake 314 

Derulus Tschitch6rine 31 

Derus Motschulsky 31 

Descarpentriesina Leraut 163 

Desmatogaster Knutson 203 

Desmocerini 287 

Desmocerrrs Audinet-Serville 287 

Desmopachria Babington 63 

Desmoris LeConte 340 

Devia Blar:kwelder 99 

DiabroticeL Chewolat 315 

Diacheila Motschulsky 15 

Diachus LeConte 307 

Diagraphct Newman 189 

Dialytellus Brown 149 

Dialytes Harold 149 

Dianchomeno Chaudoir 57 

Dianous Samouelle 109 

Diqnusa Casey 107 

Diaperinae 254 

t77 

Diaperini 255 

Diaperis Miiller 255 

Diaphoetes Waterhouse 120 

Diaphoromerus Chaudoir 45 

Diaulota Casey 107 

Diazus LeConte 151 

Dibolia Latreille 320 

Dibolocelus Bedel 133 

Dicaelus Bonelli 48 

Dicentrus LeConte 280 

Dicerca Eschscholtz 162 

Dicercini 162 

Dicheirus Mannerheim 45 

Dichelonycho Kirby 153 

Dichelonyx Harris 153 

Dichrornino Casey 155 

Dictyopterus Mulsant 188 

Dicyrtocercyon Ganglbauer 134 

Didetus LeConte 50 

Didion Casey 229 

Dienerella Reitter 239 

Dietzella Champion 349 

Dietzianus Sleeper 344 

Dimetrota Mulsant & Rey 100 

Dimetrotina Casey 100 

Dinaraea Thomson 102 

Dineutus MacLeay 72 

Dinoderinae 200 

Dinodromius Casey 58 

Diocalandra Faust 355 

Diocarabus Reitter 12 

Diochini 123 

Diochus Erichson 123 

Diomus Mulsant 231 

DioxoceraPierce 366 

Diozocera Pierce 366 

DiozodesHaldeman 281 

DiphyaulonArnett 175 

Diplocampa Bedel 27 

Diplocheila Brull6 48 

Diplocoelus Gu6rin-M6neville 241 

Diplotaxis Kirby 151 

Diplous Motschulsky 30 

Dirabius Casey 353 

Dircaea Fabricius 248 

Dirhaginae 187 

Dirhagus Latreille 187 

Disamara Lindroth 35 

Discoderus LeConte 43 

Disonycha Chevrolat 319 

Distemmus LeConte 88 

Distichalius Casey 122 

Distigmoptera Blake 320 

Ditemnus LeConte 194 

Ditropalia Casey 107 

Ditylini 266 

Ditylus Fischer von Waldheim 266 

Dolerosomus Motschulskv 182 

406

Dolichocis Dury 244 

Dolichosoma Stephens 212 

Doliema Pascoe 255 

Dolopiosomus Motschulsky 184 

Dolurgus Eichhoff 361 

Donacia Fabricius 302 

Donaciella Reitter 302 

Donaciinae 302 

Donaciini 302 

Donaciomima Medvedev 303 

DonesiaCasey 103 

Dorcaschema Haldeman 295 

Dorcaschematini 295 

Dorcasina Casey 291 

Dorcatoma Herbst 204 

Dorcatominae 204 

Dorcatomini 204 

Dorcini 144 

Dorcus Macleay 144 

Dorthesia Say 250 

Dorytomus Germar 339 

Doydirhynchinae 324 

Drachylis Casey 262 

Drapetes Dejean 186 

Drasterius Eschscholtz 176 

Drastophus Casey 82 

Dromaeolus Kiesenwetter 187 

Dromius Bonelli 58 

Drusilla Leach 106 

Dryocoetes Eichhoff 361 

Dryocoetini 361 

Dryophilinae 2Ol 

Dryophthorinae 356 

Dryophthorini 356 

Dryophthorus Germar 356 

Dryopidae 171 

Dryopoidea 169 

Dryops Olivier 171 

Dubiraphia Sanderson 172 

Dularius Thomson 284 

Dynastinae 155 

Dyschirius Bonelli 17 

Dysidius Chaudoir 32 

Dyslobus LeConte 333 

Dysmathes Mannerheim 61 

Dytiscidae 62 

Dytiscinae 71 

Dytiscini 71 

Dytiscus Linn6 71 

Eanus LeConte 181 

Eburia Audinet-Serville 280 

Ecarinosphaerula Hatch 79 

Echinaspis Blatchley 347 

Echocerus Horn 255 

Echochara Casey 105 

Echyra Erichson 154 

Ectopria LeConte 174 

Ecyrus LeConte 295 

Elacatis Pascoe 270 

Elaphidiini 280 

Elaphidion Audinet-Serville 281 

El ap hidionoide s Linsley 281 

Elaphrini 15 

Elaphropus Motschulsky 28 

Elaphrotatus Semenov 16 

Elaphroterus Semenov 16 

Elaphrus Fabricius 16 

Elasmocerus LeConte 208 

Elassoptes Horn 355 

Elater Linn6 181 

Elateridae 175 

Elaterinae 181 

Elaterini 181 

Elateroidea 174 

Elateroides Schaeffer 206 

Elathous Reitter 177 

Elatotrypes Fisher 284 

Elatrinus Horn 184 

Eleates Casey 255 

Eleodes Eschscholtz 256 

Eleodini 256 

Eleodopsis Blaisdell 257 

Eleusinini 92 

Elissa. Casey 331 

Elleschini 345 

Elleschus Dejean 345 

Ellipsoptera Dohktourow 10 

Ellychnia LeConte 191 

Elmidae 172 

Elminae 172 

Elmini 172 

Elodes Latreille 142 

Elonium Leach 88 

Elonus Casey 275 

Elytroleptus Dug6s 287 

ElytrusaCasey 101 

Embaphion Say 257 

Emmesa Newman 246 

Empelinae 87 

Empelus LeConte 87 

Emphanes Motschulsky 27 

Emphyastes Mannerheim 337 

Emphyastini 337 

Empinodes Park I27 

Emplenota Casey 105 

Enaphalodes Haldeman 280 

Encalus LeConte 345 

Enchodes LeConte 247 

Encyclops Newman 288 

Endalus Castelnau 341 

Endecatominae 200 

Endecatomus Melli6 200 

Endectus LeConte 252 

Endeitoma Sharp 251 

Endeodes LeConte 211 

407

Endomychidae 237 

Endomychinae 239 

Endomychini 239 

Endomychus Panzer 239 

Endrosq.LeConte 152 

Engamota Casey 100 

EngisPayxull 225 

Enhydrini 72 

Enicmus Thomson 239 

Enicoceru-. Stephens 74 

EnnearthronMelli6 244 

Enochrus'lhomson 132 

Enoclerus Gahan 209 

Enopliini 210 

Enoplurus Hope 131 

Entomophthalmus Bonvouloir 187 

Entomosce:Iis Chevrolat 313 

Epauloecus Mulsant & Rey 205 

Epeuchaet,zs Lyman 353 

Ephalus LeConte 257 

Ephistemtrs Stephens 223 

Epicauta Dejean 264 

Epicautina 264 

Epierus Edchson 138 

Epilachna Dejean 237 

Epilachninae 237 

Epilachnirri 237 

Epiphanis Eschscholtz 188 

EpiphloeirLi 210 

Episcopell,ts Casey 43 

Episernus Thomson 202 

Epismilia C ocker ell 229 

Epitrix Foudras 319 

Eplophorus Chevrolat 286 

Epuraea Erichson 215 

Epuraeanella Crotch 215 

Erchomus Motschulsky 96 

Eremninar: 334 

Eremosis cles Gozis 312 

Ergates Audinet-Servllle 27 7 

Ergatini 2',77 

Erichsonius Fauvel 117 

Eriopsilus Bates 295 

Erirhininae 339 

Erirhinini 339 

Erirhinus Schiinherr 339 

Erirhipidi,z Casey 157 

Erirhipis Ilurmeister 157 

Ernobiinae 202 

Ernobiini 202 

Ernobius illhomson 202 

Eronyxa Reitter 207 

Eropterus Green 189 

Eros Newrnan 189 

Erotides $Iaterhouse 189 

Erotylidae 225 

Erynephala Blake 315 

Euaesthetinae Il2 

Euaesthetus Gravenhorst 112 

EubaeoceraCornell 125 

Eubriinae 174 

Eubrychiopsis Dietz 351 

Eubrychius Thomson 351 

Eubulus Kirsch 348 

Eucanthus Westwood 150 

Euceratocerini 202 

Euchaetes LeConte 353 

Euchariruq Casey 104 

Eucicones Sharp 251 

Eucinetidae 141 

Eucinetoidea l4l 

Eucinetus Germar 141 

Euclyptus Dietz 341 

Eucnecosum Reitter 90 

Eucnemidae 186 

Eucneminae 187 

Euconnini 82 

Euconnus Thomson 82 

Eucrada LeConte 201 

Eucradini 201 

Eucryptusa Casey 107 

Euderces LeConte 286 

Eudromus l{trby 21 

Euferonia Casey 33 

Eugnamptus Schijnherr 326 

Euharpalops Casey 41 

Eulathrobium Casey 116 

Euliusa Casey 100 

Eumalus Sharp 92 

Eumichthini 282 

Eumichthus LeConte 282 

Eumicrota Casey 107 

Eumitocerus Casey 99 

Eumolpinae 308 

Eumolpini 308 

Eumolpus Redtenbacher 309 

Eumorphinae 238 

Eunonia Casey 93 

EunyssobiaCasey 353 

EuophryumBroun 355 

Eupactus LeConte 204 

Eupagoderes Horn 334 

Eupariini 149 

Eupariini 324 

Euparius Schiinherr 324 

Eupetedromus Netolitzky 25 

Euphorhipis Casey 157 

EuphoriaBurmeister 157 

Euphoriaspls Casey 157 

Eupisenus Casey 243 

Euplectinae 126 

Euplectini 126 

Euplectus Leach 126 

Eupleurida LeConte 269 

Eupogonius LeConte 295 

Eupogonocherus Linsley 296 

408

Eupristocerus Deyrolle 167 

EurealeConte 174 

Eurelymis Casey 212 

Euremus Bierig 119 

Eurhizophagus M6quignon 218 

Europhilus Chaudoir 52 

Eurydactylus LaFert6-S6nectbre 49 

Euryderus LeConte 40 

Eurygeniinae 274 

Eurygeniini 274 

Eurylister Bickhardt 140 

Eurymeloe Reitter 263 

Eurymetopini 253 

Eurymycter LeConte 325 

Euryoptera Horn 284 

Eurypalpus LeConte 174 

Eurypogon Motschulsky 175 

Eurysphindinae 213 

Eurysphindus LeConte 213 

Eurystethus Seidlitz 270 

Eurytrachelus Motschulsky 21 

Eurytrichus LeConte 45 

Eusomalia Casey 160 

Eusphalerini 87 

Eusphalerum Kraatz 87 

Eusphyrus LeConte 325 

Euspilotus Lewis 137 

Eustilbus Sharp 226 

Eustrophinae 246 

Eustrophinus Seidlitz 246 

Eustrophus Illiger 246 

Eutetrapha Bates 298 

Eutheia Stephens 83 

Eutheiini 83 

Euthyna Gistel 304 

Eutoxotus Casey 288 

Eutrichapion Reitter 328 

Eutrichites LeConte 128 

Euvacusus Casey 273 

Euvrilletta Fall 204 

Euxenus LeConte 324 

Euxestinae 227 

Evarthrus LeConte 35 

Evodinus LeConte 289 

Euoplus LeConte 255 

Evotus LeConte 332 

Excavodes Park 127 

Exema Lacordaire 307 

Exochomus Redtenbacher 233 

Exomella Casey 159 

Exopioide s Gu6rin-M6n6ville 199 

Exosternini 140 

Explorator Pacheco 169 

Fabricianus Weise 307 

Falacer Castelnau 259 

Falagria Leach 105 

Falagriini 105 

Fallapion Kissinger 328 

Faroninae 126 

Faronini 126 

Feronia Latreille 30 

Fidia Walsh 309 

Flavohelodes Klausnitzer 142 

Florilinus Mulsant & Rey 199 

FluuicolaDeKay 174 

Fornax Castelnau 187 

Foveoscapha Park & Wagner 126 

Funda Blackwelder 104 

Furcacampa Netolitzky 26 

Fuscocalathzs Ndgre 51 

Gabrius Stephens 119 

Gabronthus Tottenham 119 

Galerita Fabricius 59 

Galeritino Jeannel 59 

Galeritini 59 

Goleritula Strand 59 

Galeruca Miiller 315 

Galerucella Crotch 314 

Galerucinae 314 

Galerucini 314 

Galeruclerus Gahan 210 

Gastrallini 202 

Gastrallus Jacquelin du Val 202 

Gastrellarius Casey 32 

Gastroeideq.Hope 312 

Gastrolobium Casey 116 

Gastrophysa Chewolat 312 

Gastrotaphrus Buchanan 338 

Gaurambe Thomson 207 

Gaurodytes Thomson 67 

Gauropterus Thomson 123 

Gaurotes LeConte 289 

Gehringia Darlington 19 

Gehringiini 19 

Gelus Casey 348 

Genattus Sharp 197 

Gennadota Casey 99 

Genyon Smetana 134 

Geoderces Horn 333 

Geodercodes Casey 332 

Geodromicus Redtenbacher 91 

Geomorus Schiinherr 336 

Geomysaprinus Ross 137 

Geopatrobus Darlington 29 

Geopinus LeConte 43 

Georyssidae 135 

Georyssus Latreille 135 

Geostiba Thomson 101 

Geotrupes Latreille 150 

Geotrupinae 150 

Geotrupini 150 

Gephelophorus Sharp 131 

Geraeus Pascoe 353 

Gerstaeckeria Champion 347 

409

Gibbiinae 205 

Gibbium Scopoli 205 

Gibbobruchus Pic 301 

Glaphyra Newman 282 

Glaphyrinae 150 

Glaphyrocanthon Martinez 146 

Glaresis Erichson 150 

Glipa LeConte 249 

Glipostenoda Ermisch 250 

Glischrochilus Reitter 217 

Glycerius Casey 46 

Glycobius LeConte 284 

Glyphony>: Candbze 182 

Glyphuroplata Uhmann 322 

Glyptasida Casey 254 

Glyptina l,eConte 317 

Glyptoderus LaFert6-S6nectdre 49 

Glyptoma Erichson 92 

Glyptophorus Park 221 

Glyptoscelis Chevrolat 309 

Gnathacmaeops Linsley & Chemsak 

Gnathium Kirby 266 

Gnathoncus Jacquelin du Val 138 

Gnathospa'.sta Horn 264 

Gnathotri

Helocerus Mulsant & Rey 199 

Helocharini 132 

Helocombus Horn 133 

HelodesPaykull 312 

Helophorinae 130 

Helophorus Fabricius 130 

Helopini 258 

Helops Fabricius 258 

Hemantus Casey 273 

Hemicallidium Casey 283 

Hemicarabus G6hin 12 

Hemicoelus LeConte 203 

Hemicrepidius Germar 181 

HemicyphonLeConte 142 

Hemimedon Casey 113 

Hemiptychus LeConte 205 

Hemiquedius Horn 122 

Hemisopalus Casey 43 

Hemispilota Casey 154 

Hemistenus Motschulsky 109 

Henoticoides Hatch 221 

Henoticus Thomson 221 

Henotiderus Reitter 221 

Henous Haldeman 264 

Hesperobaenus LeConte 2I8 

Hesperobium Casey 116 

Hesperolinus Casey 124 

Hesperophanes Mulsant 280 

Hesperophanini 280 

Hesperosaprinus Wenzel 137 

Hesperus Fauvel 119 

Hetaeriinae 141 

Hetaeriini 141 

Hetaerius Erichson 141 

Heterachthes Newman 281 

Heteracthon Thomson 281 

Heterarthron Dejean 199 

Heteraspis LeConte 309 

Heterhelus Jacquelin du Val 214 

Heterlimnius Hinton 173 

Heteroceridae 169 

Heterocerini 170 

Heterocerus Fabricius 170 

Heterodromla Haldeman 220 

Heteronychus Seidlitz 67 

Heterops Mannerheim 90 

Heteropus Castelnau 257 

HeterosilphaPortevin 85 

Heterosternus Zimmermann 65 

Heterosternuta Strand 65 

Heterostethus Falkenstrom 65 

Heterothops Stephens 121 

Hetoernis Haldeman 295 

Hexarthrum Wollaston 355 

Hexirhexius Grigarick & Schuster 

Hiloro Mulsant & Rey 102 

Hilarina Casey 100 

Himatinum Cockerell 355 

126 

Himatium Wollaston 355 

Himatolabus Jekel 326 

HippodamiaDejean 234 

Hippopsini 296 

Hippopsis Lepeletier & Audinet-Serville 

Hippuriphila Foudras 318 

Hirmoplataphus Netolitzky 23 

Hispinae 321 

Hister Linn6 139 

Histeridae 135 

Histerinae 139 

Histerini 139 

Histeropsis Chevrolat 255 

Holciophorus LeConte 31 

Holoboreaphilus Campbell 89 

Hololepta Paykull 140 

Hololeptini 140 

Holopleura LeConte 287 

Holopleurini 287 

Holostrophus Horn 246 

Holotrochus Erichson 92 

Homaeotarsus Hochhuth 116 

Homalochthebius Kttwefi 7 4 

Homalostylus Chevrolat 354 

Homalota Mannerheim 107 

Homalotina 107 

Homalotusa Casey 102 

Homoeocarabus Reitter 11 

Homoeolabus Jekel 326 

Hornoeosomo Austin 80 

HomogasterProvancher 348 

Homorosoma Frivaldszky 350 

Hoplandria Kraatz 105 

Hoplandriini 105 

Hoplia Illiger 154 

Hopliini 154 

Hoplocephalo Castelnau & Brull6 255 

Hoppingiana Blaisdell 212 

Horistonotus Canddze 185 

Hormorini 332 

Hormorus Horn 332 

Hornaltica Barber 318 

Hornia Riley 265 

Horticola Lindemann 241 

Hybodera LeConte 282 

Hyboderini 282 

Hydaticini 71 

Hydaticus Leach 71 

Hydaticus Sch6nherr 350 

Hydnobiini 78 

Hydnobius Schmidt 78 

Hydnocera Newman 208 

Hydraena Kugelann 74 

Hydraenidae 74 

Hydraeninae 74 

Hydraenini 74 

Hydriomicrus Casey 23 

Hydrium LeConte 21 

296 

tr

Hydrobiinae 131 

Hydrobiini 131 

Hydrobiusr Leach 131 

Hydrocan-:hus Say 62 

HydrocharaBerthold 133 

Hydrochinae 130 

Hydrochu,s Leach 130 

Hydrogale,ruco Laboissidre 314 

Hydrophilidae 130 

Hydrophilinae 133 

Hydrophiloidea 130 

Hydrophilus Miiller 133 

Hydroporinae 63 

Hydroporini 64 

Hydroponrs Clairville 65 

HydrosmectaThomson 100 

Hydrothassa Thomson 312 

Hydrovatini 63 

Hydrovatrrs Motschulsky 63 

Hygrotus Stephens 64 

Hylastes Ilrichson 356 

Hylastini 356 

Hylastinotdes Spessivetsev 357 

Hylastinus Bedel 357 

Hylecoetirrae 206 

Hylecoetus Latreille 206 

Hylesininae 356 

Hylesinin; 357 

Hylesinus Fabricius 357 

Hylis des,]ozis 188 

Hylobiini 337 

Hylobius (]ermar 337 

Hyloceotu,:,; Melsheimer 206 

Hylocurus Eichhoff 359 

Hylota Casey 99 

Hylotrupes Audinet-Servllle 282 

Hylurgopinus Swaine 357 

Hylurgops LeOonte 356 

Hymenochara Campbell 260 

Hy menoph.orus Mulsant 259 

Hymenortrs Mulsant 259 

Hypebaeus Kiesenwett er 212 

Hypera Gr:rmar 336 

Hyperaspidius Crotch 231 

Hyperaspini 231 

Hyperaspis Redtenbacher 23I 

Hypericia Bedel 311 

Hyperinae 336 

Hyperini 336 

Hyperisus Mulsant &Rey 202 

Hypermal,t u.s Lacordaire 280 

Hyperodes Jekel 334 

Hyperplatys Haldeman 298 

Hypherpes Chaudoir 31 

Hyphydrini 63 

Hypnoryra Casey 124 

Hypnoidus Dillwyn 178 

Hypoborini 358 

Hypocaccus Thomson 137 

HypocoelusDejean 188 

Hypocoprinae 223 

Hypocoprus Motschulsky 223 

Hypocyphtus Gyllenhal 108 

Hypodacne LeConte 227 

Hypolithus Eschscholtz 178 

Hypomedon Mulsant & Rey 113 

HypomolyxleConte 337 

Hypophlaeus Fabricius 256 

Hypophloeinae 256 

Hypophloeini 256 

Hypostenus Rey 109 

Hypothenemus Westwood 362 

Hypulini 247 

Hypulus Paykull 247 

Ibidiini 281 

Ichnodes Dejean 262 

Ichthyurini 195 

Ictinosternun Smetana 134 

Idiobates Casey 259 

Idiopidonia Swaine & Hopping 291 

Idiostethus Casey 353 

Ilendus Casey 242 

Ilybiosoma Crotch 67 

Ilybius Erichson 69 

Ilyobates Kraatz 99 

Ipelates Reitter 77 

Iphthiminus Spilman 261 

Ipini 360 

Ips DeGeer 360 

Irichroa Newman 12 

Isarthrus LeConte 187 

Ischalia Pascoe 269 

Ischaliinae 269 

Ischnosoma Stephens 98 

Ischyropalpus LaFert6-S6nectdre 273 

Ischyrus Lacordafte 226 

Isochnus Thomson 345 

Isoglossa Casey 99 

Isohydnocera Chapin 209 

Isomira Mulsant 259 

Isoplastus Horn 79 

Isopleurus Kirby 36 

Isorembus Jeannel 48 

Isorhipina Casey 157 

Isorhipis Boisduval & Lacordaire 187 

Isosceles Newman 299 

Isostenia Vacher de Lapouge 11 

Isotachys Casey 29 

Ithyceridae 325 

Ithycerus Sch6nherr 325 

Ithyporini 347 

Ixapion Roudier & Tempere 328 

.Irios Deville 328 

4t2

Jonthonota Spaeth 322 

Judolia Mulsant 291 

Kalissus LeConte 84 

Katastylops Pierce 367 

Knowltonia Fisher 166 

Knulliana Linsley 286 

Korynetinae 210 

Korynetini 211 

Kuschelina Bechyn6 320 

Kytorhininae 300 

Kytorhinus Fischer von Waldheim 

Labidomera Chevrolat 311 

Laccobius Erichson 131 

Lacconotus LeConte 267 

Laccophilinae 63 

Laccophilini 63 

Laccophilus Leach 63 

Laccornis des Gozis 67 

Lachnocrepis LeConte 49 

LachnosternaHope 152 

Lacon Castelnau 175 

Laemophloeinae 219 

Laemophloeus Dejean 219 

Laemosaccini 342 

Laemosaccus Schdnherr 342 

Laemostenus Bonelli 51 

Laestris Melsheimer 109 

Laetulonthus Moore & Legner 119 

Lagarus Chaudoir 33 

Lagriidae 261 

Lagrlinae 262 

Lamiinae 294 

Lamiota Casey 101 

Lamperos Allard 258 

Lamprias Bonelli 56 

Lampropterus Mulsant 282 

Lampyridae 190 

Lampyrinae 190 

Languria Latreille 224 

Languriidae 223 

Languriinae 224 

Langurlini 224 

Lanternarius Pacheco 170 

Lapethus Casey 227 

Laphyra Newman 282 

Lappus Casey 273 

Lapsus Pacheco 170 

Lara LeConte 172 

Laricobius Rosenhauer 196 

Lafinae I72 

Larini 172 

Larisia Err,ery 276 

Lasconotus LeConte 251 

Lasioderma Stephens 204 

Lasiodermini 204 

Lasiotrechus Ganglbauer 19 

300 

Latheticus Waterhouse 258 

Lathridiidae 239 

Lathridiinae 239 

Lathridius Herbst 239 

Lathrimaeunz Erichson 91 

Lathrium LeConte 89 

Lathrobiella Casey 116 

Lathrobiina 115 

Lathrobioma Casey 115 

Lathrobium Gravenhorst 115 

Lathrolepta Casey 115 

Lathrotaxis Casey 116 

Lathrotropis Casey 116 

Lebia Latreille 56 

Lebiini 56 

Lechriopini 348 

Lechriops Schcinherr 348 

Lecontea Olivier 190 

Leconteus Lutshnik 30 

Lecontia Champion 268 

Lederia Reitter 246 

Leiestini 238 

Leiochiton Curtis 19 

Leiodes Latreille 79 

Leiodidae 77 

Leiodinae 78 

Leiodini 78 

Leiolinus Casey 124 

Leionotus Kirby 71 

Leirus Zimmermann 35 

Leistus Frrilich 13 

Lema Fabricius 304 

Lemiini 304 

Lerua Casey 7I3 

Leperisinus Reitter 357 

Lepesoma Motschulsky 333 

Lepiarcha Sharp 217 

Lepidophorus Kirby 333 

Lepidotus Stephens 175 

Lepisomus Kirby 359 

Leptacinodes Casey 123 

Leptacinus Erichson 123 

Leptacmaeops Casey 290 

Leptaegialia Brown 147 

Leptalia LeConte 289 

Leptinillus Horn 82 

Lentininae 82 

Leptinotarsa StAl 311 

Leptinus Mtiller 82 

Leptoferonia Casey 32 

Leptomus Casey 8 

Leptophloeus Casey 220 

Leptopiini 333 

Leptorus Casey 114 

Leptoscydmini 83 

Leptoscydmus Casey 83 

Leptostylus LeConte 297 

Leptothrix Heikertinger & Csiki 318 

413

Leptotrachelus Latreille 56 

Leptotrichaltica Helkertinger 3 18 

LeptotrixHorn 318 

Leptotyphlinae 113 

Leptura Linr'6 292 

Lepturges Bates 297 

Lepturinae 287 

Lepturini 288 

Lepturopsis Linsley & Chemsak 293 

Leptusa I{raatz 107 

Lepyrus Germar 337 

LestqBlackwelder 90 

Lesteva Latreille 90 

Leuchydrium Casey 25 

Leucopaed,erus Casey 116 

Lexiphaners Gistel 306 

Liaphlus ()uignot 61 

Liq,semum, Casey 279 

Librodor Reitter 217 

LichenophLanes Lesne 200 

LichnantL,e Burmeister 150 

Lichnocarabus Reitter 11 

Licinini 48 

Ligmargus Stibick 178 

Lignyodes Dejean 345 

Lignyodini 345 

Ligyrellus Casey 156 

Liryrodes Casey 156 

Ligyrus Burmeister 156 

Lilioceris Reitter 304 

Limnebiui; Leach 74 

Limnichiciae 171 

Limnichir.ae 171 

Limnichir.i 171 

Limnichites Casey 171 

Limnocharis Horn 74 

Limonius Eschscholtz 176 

Limulodes Matthews 76 

Limulodidae 76 

LinaDejean 313 

Lindorus'Casey 233 

Liniolis Grigarick & Schuster 126 

Linohespe'rus Smetana 123 

Linolathro.Casey 116 

Linsleya MacSwain 265 

Linsleyana, Podany 288 

Linsleyina Selander 265 

Liocellus Motschulsky 46 

Liocellus Tschitch6rine 46 

Liocosmius Casey 25 

Liocyrtusa Daffner 79 

Liodessus Guignot 63 

Liodicaelus Casey 49 

Liogluta Thomson 102 

Lioligus Casey 159 

Lionepha Casey 21 

Lionothus Brown 78 

Lioon Casey 159 

Liparocephalus Miiklin 108 

Lispinini 92 

Lissominae 186 

Lissorhoptrus LeConte 341 

Listemus Casey 160 

Listronotus Jekel 334 

Listrus Motschulsky 213 

Litargus Erichsor' 242 

Litheleodes Blaisdell 256 

Lithocharis Dejean 113 

Lithocharodes Sharp 123 

Lithodus Germar 335 

Litochropus Erichson 226 

Litodactylus Redtenbacher 350 

Litolethra Casey 115 

Lixellus LeConte 335 

Lixestus Reitter 336 

Lixini 337 

Lixus Fabricius 337 

Loania Kinzelbach 366 

Loberini 224 

Loberus LeConte 224 

Lobiopa Erichson 216 

Lobosoma Buchanan 338 

Lobrathium Mulsant & Rey 116 

Loceptes Casey 347 

Lodinus Mulsant &Rey 257 

Longitarsus Berthold 317 

Lopha Stephens 27 

Lophalophus LeConte 333 

Lopheros LeConte 189 

Lophioderini 82 

Lophioderus Casey 82 

Lophoglossus LeConte 30 

Lophomucter Notman 105 

Lophopogonius Linsley 296 

Lordithon Thomson 98 

Loricera Latreille 15 

Loricerini 15 

Lorinota Casey 105 

Loxandrus LeConte 30 

Loxopeza Chaudoir 56 

Lucanidae 143 

Lucaninae 144 

Lucanini 144 

Lucanus Scopoli 144 

Lucidota Castelnau 191 

Lucifotychus Park & Wagner 129 

Ludius Berthold 181 

Lumetus Zaitzev L32 

Luperini 315 

Lutrochus Erichson 171 

Lycidae 188 

Lycinae 188 

Lycoperdina Latreille 238 

Lycoperdinini 238 

Lyctidae 201 

Lvctini 201 

4L4

Lyctus Fabricius 201 

Lygerus Kiesenwetter 195 

Lygistopterini 188 

Lymantor Lovendal 361 

Lymexylidae 206 

Lymexyloidea 206 

Lyperopherus Motschulsky 35 

Lyperosomus Motschulsky 33 

Lyperostenia Vacher de Lapouge 10 

Lyperus Chaudoir 33 

Lypoglossa Fenyes 103 

Lyrosoma Mannerheim 77 

Lytta Fabricius 264 

Lyttina 264 

Lyttim 264 

Macdonaldium Abdullah & Abdullah 

Machaerodes Brendel 128 

Macratria Newman 275 

Macratriini 275 

Macreurops Casey 218 

Macrobasis LeConte 265 

Macrodactylini 153 

Macrodactylus Latreille L54 

Macrodea Casey 222 

MacrodytesThomson 71 

Macronaemia Casey 233 

Macronoxia Crotch 152 

Macronychini 173 

Macronychus Miiller 173 

Macropogon Motschulsky 174 

Macropogoninae 174 

Macrops Kirby 334 

Macrorhoptus LeConte 343 

Macrosiagon Hentz 250 

Macrotelus Klug 208 

Maculicq.uta Dillon 264 

Madarellus Casey 353 

Madarini 352 

Magdalinae 342 

Magdalini 342 

Magdalis Germar 342 

Malachiinae 2l]- 

Malachius Fabricius 211 

Malacocis Gorham 245 

Mallodrya Horn 243 

Malporus Casey 273 

Malthacus Kirby 192 

Malthinini 194 

Malthinus Latreille 194 

Malthodes Kiesenwetter 194 

Mantura Stephens 319 

Maphoca Casey 245 

Margaiostus Stibick 178 

Margarinotus Marseul 139 

Marginus LeConte 241 

Margus Dejean 258 

Masoreini 56 

76 

Mathrilaeurn Moore 91 

Matini 71 

Matus Aub6 71 

Mecas LeConte 299 

Mecinini 347 

Mecinus Germar 347 

Mecopeltus Dietz 351 

Mecynotarsus LaFert6-S6nectdre 271 

Medon Stephens 113 

MedonellaCasey 113 

Medonina 113 

Medonodonta Casey 114 

Megacerus Fahraeus 300 

Megacrotono Scheerpeltz 103 

Megacyllene Casey 284 

Megalechthrus Perkins 366 

Megaleptura Casey 292 

Megaliridia Casey 12 

Megalodacne Crotch 225 

Megalostomini 305 

MegalticaHatch 317 

Meganemognatha Enns 265 

Megapangus Casey 40 

Megapenthes Kiesenwetter 184 

Megaquedius Casey 121 

Megarafonus Casey 126 

Megarthrus Curtis 87 

Megasemum l{raatz 279 

Megasternini 134 

Megasternum Mulsant 134 

Megastilicus Casey 114 

Megatoma Herbst 198 

Megatomini 197 

Megilla Mulsant 233 

Megodontus Solier 12 

Megomus Casey 8 

Meibomeus Bridwell 301 

Melalgus Dejean 199 

Melamomphus Horn 333 

Melanagonum Casey 53 

Melandrya Fabricius 246 

Melandryidae 245 

Melandryiinae 246 

Melandryini 246 

Melaneleodes Blaisdell 256 

Melanius Bonelli 33 

Melanophila Eschscholtz L65 

Melanophilini 165 

Melanophthalma Motschulsky 240 

Melanotini 184 

Melanotrupes Blanchard 150 

Melanotus Eschscholtz 184 

Melanthaxia Richter 165 

Melasinae 187 

Melasis Olivier 187 

Melq.soma Stephens 313 

Melasomida Schaeffer 312 

Melba Casey 127 

4r5

Meligethes Stephens 217 

Meligethinae 217 

Melittophcgus Kirby 263 

Meloe Lir.n6 263 

Meloidae 263 

Meloinae 263 

Meloini 263 

MelolontkLinae 151 

Melolonthini 151 

Melomalus Casey 22 

Melyridaer 211 

Membrqctxenos Pierce 366 

Menechid'zs Motschulsky 258 

Meracantha Kirby 259 

Merhynchites Sharp 327 

Meridion des Gozis 283 

Meriellum Linsley 283 

Merinus LeConte 260 

Merium LeConte 283 

Meronera Sharp 100 

Mesauletes Voss 327 

Mesosini 295 

Metablap";'lis Blaisdell 257 

Metqbletus Schmidt-Goebel 58 

Metabola Chaudoir 57 

Metachro:na Chewolat 308 

Metaclisa Jacquelin du Val 258 

Metacmaeops Linsley & Chemsak 

Metallina Motschulsky 21 

Metallogra,pho Motschulsky 310 

Metamelanlus Tschitsch6rine 33 

Metaxyodonla Casey 113 

Methlini 63 

Methydrus Rey 132 

MetoponWaterhouse 327 

Metrius Eschscholtz 19 

Mezium Clurtis 205 

Miarus Schiinherr 347 

Miccotrogus Schajnherr 346 

Micracini 359 

Micracis l,eConte 359 

Micracisoides Blackman 359 

Micraegialia Brown 146 

Micragonum Casey 54 

Micralymma Westwood 88 

Micrathous Lane I77 

Micrepitrix Laboissidre 318 

Microaug;vles Pacheco 170 

Microbledius Herman 93 

Microbregma Seidlitz 203 

Microcallidium Casey 283 

Microcara Thomson 142 

Microclytus LeConte 286 

Microcylloepus Hinton 173 

Microdota Mulsant & Rey 102 

Microedus LeConte 91 

Microgoes Casey 295 

Micrograrnrne W alkJey 239 

290 

Microhyus LeConte 347 

MicrolathraCasey 116 

Microlestes Schmidt-Goebel 58 

Microlipus LeConte 211 

Micromalthidae 6 

Micromalthus LeConte 6 

Micromaseus Casey 34 

MicromedonCasey 113 

Micromelomalus Casey 22 

Micromota Casey 100 

Micronychus Provancher 333 

Micropeplidae 84 

Micropeplini 84 

Micropeplus Latreille 84 

Microrhagus Dejean 187 

Microrhopala Chewolat 322 

Microsaurus Stephens 121 

Microscaph a LeConte 246 

Microschatia Solier 254 

Microscydmus Saulcy & Croissandeau 

Microtachys Casey 29 

Microtonus LeConte 247 

Microweise a Cockerell 229 

Microweisini 229 

Millidium Motschulsky 75 

Minyomerus Horn 331 

Mioptachys Bates 29 

Miscodera Eschscholtz 19 

Mitosynum Campbell 93 

Mocyta Mulsant & Rey 102 

Molamba Casey 228 

Molorchini 282 

Molorchus Fabricius 282 

Moluciba Casey 99 

Molytinae 337 

Monachini 306 

Monachulus Leng 306 

Monachus Chevrolat 306 

Monarthrum Kirsch 365 

Moneilema Say 294 

Moneilemini 294 

M onillop atrob us }Iatcli' 29 

Monocerus Faldermann 27 | 

Monochamini 294 

Monochamus Dejean 294 

MonoferoniaCasey 32 

Mononychini 349 

Mononychus Germar 349 

Monophylla Spinola 208 

Monoplesa Motschulsky 294 

Monotoma Herbst 218 

Monotominae 218 

Monoxia LeConte 315 

Mordella Linn6 248 

Mordellidae 248 

Mordellini 248 

Mordellistena Costa 249 

Mordellistenini 249 

83 

416

Morphnosoma Lutshnik 33 

Morychus Erichson 160 

Motschulskium Matthews 75 

Muirixenos Pierce 367 

Mulsantina Weise 237 

Murmidiinae 227 

Murmidius Leach' 227 

Mutinus Casey 112 

Myas Dejean 31 

Mycetaea Stephens 238 

Mycetaeinae 238 

Mycetaeini 238 

Mycetina Mulsant 239 

Mycetinini 238 

Mycetochara Berthold 260 

Mycetochares Latreille 260 

Mycetocharini 260 

Mycetocharis Gyllenhal 260 

Mycetophagidae 242 

Mycetophagus Hellwig 242 

Mycetophila Gyllenhal 260 

Mycetoporus Mannerheim 98 

Mychocerus aluct. 227 

Mychocerus Erichson 227 

Mycocerinus Slipinski 227 

Mycteridae 267 

Mycterus Clairville 267 

Myeleborus Blackman 363 

Myllaena Erichson 108 

Myllaenini 108 

Myloechus Latreille 81 

Myochrous Erichson 309 

MyocorynaStel 311 

Myodes Latreille 250 

Myodites Latreille 250 

Myrmecocephalus Macleay 105 

Myrmedoniini 106 

Myrmedophila Bousquet 222 

Myrmex Sturm 341 

Myrmolamio Bates 298 

Mystaxus Kugelann 248 

MyziaMulsant 235 

Nacerdes Dejean 266 

Nacerdinae 266 

Namunaria Reitter 251 

Nanobius Herman 95 

Nanodactylus Blatchley 328 

Nanodes Schtinherr 328 

Nanophyes Schiinherr 328 

Nanophyini 328 

Narpus Casey 173 

Nartus Zaitzev 70 

Nassipa Ernery 276 

Nathicus Casey 271 

Nathrenus Casey 198 

Naupactini 329 

Nausibius Redtenbacher 220 

Neacanthocinus Dillon 296 

Neandra Lameerc 277 

Neanthophylax Linsley & Chemsak 289 

Neatus LeConte 259 

Nebria Latreille 14 

Nebriini 13 

Necrobia Olivier 211 

Necrodes Leach 85 

Necrophila Kirby & Spence 85 

Necrophilus Latreille 77 

Necydalini 287 

Necydalis Llnn6 287 

Nedyus Sch

Neophytobius Wagner 351 

Neopolya.rthron Semenov 278 

Neoporus Guignot 65 

Neopyrochroa Blair 269 

Neorthopleura Barr 210 

Neosaprinus Bickhardt 137 

Neoscutopterus Balfour-Browne 

Neostylops Pierce 367 

Neotrochus Blackwelder 92 

Neozonitis Enns 265 

Nephanes Thomson 76 

Nephus Mulsant 231 

Nicaginae 144 

Nicagini 144 

Nicagus l,eConte 144 

Nicobiini 202 

Nicrophorinae 85 

Nicrophorus Fabricius 85 

Niptus Boieldieu 205 

Nlsa Casey 128 

Nisaxis Casey 128 

Nitidula Fabricius 216 

Nitidulidae 214 

Nitidulinae 215 

Nitionini 254 

Nocheles LeConte 333 

Noda Chevrolat 308 

Nodinini 308 

Nodonota Lefbvre 308 

Nomaretus LeConte 13 

Nomaspis LeConte 264 

Nomidus Casey 334 

Nomiini 29 

Nominus Motschulsky 59 

Nomius Castelnau 29 

Nordenskioeldia Sahlberg Il2 

Nosodendridae 196 

Nosodenclron Latreille 196 

Nosodes LeConte 207 

Notaphusr Stephens 25 

Notaris Ciermar 339 

Noteridaer 61 

Nothopus LeConte 40 

Nothus Olivier 248 

Notiodes Schiinherr 341 

Notionomus Erichson 341 

Notiophilini 15 

Notiophilus Schtinherr 341 

Notiophilus Dum6ril 15 

Notobia Perty 45 

Notoxinae 271 

Notoxini 271 

Notoxus Fabricius 271 

Novelsis Casey 197 

Nudatoconnus Franz 83 

Nudobius Thomson l-23 

Oberea Mulsant 299 

70 

Obriini 281 

Obrium Dejean 281 

Ocalea Erichson 99 

Occlephelinus Hatch 89 

Ochodaeinae 149 

Ochodaeus Audinet-Serville 149 

Ochrosidia Casey 155 

Ochthebiinae 74 

Ochthebius Leach 74 

Ochthedromus LeConte 21 

Ochthephilum Stephens 116 

Ochthephilus Mulsant & Rey 94 

Ochthexenus Motschulsky 88 

Octotemnus Melli6 245 

Ocyusa l{raatz 99 

OdacanthellaLiebke 56 

Odacanthini 56 

Odontaeus Fischer von Waldheim 150 

Odontium LeConte 20 

Odontocorynus Schijnherr 353 

Odontomus Kirby 166 

Odontonyx Stephens 51 

Odontopus Say 345 

Odontosphindini 213 

Odontosphindus LeConte 213 

Odontota Chewolat 321 

Odontotaenius Kuwert 145 

OebaceresThomson 295 

Oedemeridae 266 

Oedemerinae 266 

Oedostethus LeConte 178 

Oestodes LeConte 181 

Oestodini 181 

Oiceoptoma Leach 85 

Olibrini 226 

Olibrus Erichson 226 

Oliganomalo Casey 154 

Oligolinus Casey 123 

Oligomerodes Fall 204 

Oligomerus Redtenbacher 202 

Oligomia Casey 100 

Oligopterus Casey 113 

Oligotini 108 

Olisthaerinae 92 

Olisthaerus Dejean 92 

Olisthopus Dejean 51 

Olla Casey 236 

Olophrum Erichson 89 

Omq.lq. Motschulsky 27 

Omaliinae 87 

Omaliini 88 

Omalium Gravenhorst 88 

Omq.seidius Jeannel 33 

Omaseulus Lutshnik 34 

Omuseus Dejean 33 

Omias Germar 332 

Omoglymmius Ganglbauer 7 

Omonadus Mulsant & Rev 273 

418

Omophron Latreille 16 

Ornophronini 16 

Omorgus Erichson 150 

OmoscylonThomson 294 

Omosita Erichson 216 

Omus Eschscholtz 8 

Oncideres Lepeletier & Audinet-Serville 

Onciderini 296 

Onibathum Tottenham 87 

OnichodonNewman 187 

Ontholestes Ganglbauer 120 

Onthophagini 146 

Onthophagus Latreille 146 

Onthophilus Leach 138 

Onychobaris LeConte 352 

Onychotrupes Jekel 150 

Onychylis LeConte 341 

Oodes Bonelli 49 

Oodini 49 

Opatrinae 256 

Opatrini 257 

Ophistomis Thomson 290 

Ophonus Dejean 40 

Ophraella Wilcox 315 

Ophryastes Say 334 

Ophryastini 334 

Opilo Latreille 209 

Opisthiini 15 

Opisthius Kirby 15 

Oplosia Mulsant 296 

Opresus Casey 83 

Opsimini 280 

Opsimus Mannerheim 280 

Optioservus Sanderson 173 

Opuntiaphila Pierce 347 

Orchesia Latreille 246 

Orchesiini 246 

Orchestes Illiger 345 

Ordobrevia Sanderson 173 

Oreina Monros & Bechyn6 311 

Oreocharq Casey 104 

Oreodytes Seidlitz 67 

Oreoxenus Tschitch6rine 46 

Orestia Germar 318 

Orestioides Hatch 318 

Ormiscini 325 

Ormiscus Waterhouse 325 

Orobanus LeConte 90 

Orochares Kraatz 91 

Orophiinae 245 

Orophiini 245 

Oropus Casey 126 

Orphelophorus d'Orchymont 131 

Orphilini 199 

Orphilus Erichson 199 

Orsodacna Dejean 303 

Orsodacne Latreille 303 

Orsodacninae 303 

296 

OrsonyxLeConte 151 

Orthaltica Crotch 318 

Orthoboris Casey 351 

Orthocis Casey 245 

Ortholeptura Casey 293 

Orthoperini 228 

Orthoperus Stephens 228 

Orthoris LeConte 352 

Orthosoma Audinet-Sewllle 27 7 

Orthotomicus Ferrari 360 

Orthotomides Wood 360 

Orus Casey 114 

Oryctini 156 

Oryzaephilus Ganglbauer 220 

Osmoderma LePeletier & Audinet-Serville 157 

Osmodermini 157 

Osoriinae 92 

Osoriini 92 

Osphya Illiger 248 

Osphyinae 248 

Osphyini 248 

Ostoma Laicharting 207 

Ostomodes Reitter 207 

Othiini 123 

Othniinae 270 

Othnius LeConte 270 

Otidocephalinae 341 

Otidocephalini 341 

Otidocephalzs Chevrolat 341 

Otiorhynchinae 331 

Otiorhynchini 331 

Otiorhynchus Germar 331 

Oulema des Gozis 304 

Oulimnius des Gozis 173 

Oxoplus LeConte 287 

Oxybleptes Smetana 124 

Oxygonus LeConte 180 

Oxylaemus Erichson 252 

Oxymedon Casey 113 

Oxynychus LeConte 231 

Oxyomus Stephens 148 

Oxypoda Mannerheim 99 

Oxypodini 99 

Oxyporinae 109 

Oxyporus Fabricius 109 

Oxypselaphus Chaudoir 52 

Oxypteris Kirby 165 

Oxytelinae 92 

Oxytelini 94 

Oxytelus Gravenhorst 95 

Pachybrachini 305 

Pachybrachys Chevrolat 305 

Pachybruchus Pic 300 

Pachydela Rivalier 8 

Pachydeniini 153 

Pachylobius LeConte 338 

Pachyphanes Dietz 340 

4t9

Pachyrhjnus Schcinherr SB0 

Pachy rhSt nchus l{trby 325 

Pachysctrelini 168 

Pachyschelus Solier 168 

Pachystilicus Casey 114 

Pachystylus Rey 305 

Pachyba Dejean 288 

Pachytychius LeConte 340 

Pactopus LeConte 186 

Poederillus Casey 116 

Paederina 116 

Paederinae 113 

Paederini 113 

Paederus Fabricius 116 

Palaminus Erichson 113 

Pallodes Erichson 216 

Palorus Mulsant 258 

Palporus Campbell 97 

Panagaeini 49 

Panagaeus Latreille 49 

PanalotaCasey 101 

Pandeletr:ius Schtinherr 331 

Panscopi,l,ius Pierce 333 

Panscopu.s Schiinherr 333, 334 

Parabracteon Notman 20 

Paracercrron Seidlitz 134 

Paraclivi:ra Kult 18 

Parac5rmus Thomson 132 

Paradelitrthrum Hatch gt 

Paradicaelus Ball 48 

Paradilacra Bernhauer 102 

Parafilum,is Casey 207 

Paragonu.m Casey 53 

Paralarica Breuning 295 

ParalathraCasey 116 

Paralesteua Casey g0 

Paraliaphlus Wallis 60 

Paralister Bickhardt 139 

Paralitargus Casey 242 

Parollelin.a Casey 291 

Parallelostethus Schwarz 181 

Paralopha Casey 27 

Paralytta Selander 264 

ParqmedonCasey 114 

Parametriona Spaeth 323 

ParamysiaReitter 235 

Paranaemia Casey 233 

Paranchus Lindroth 52 

Parandra Latreille 277 

Parandrinae 277 

Parandrini 277 

Paranomctla Casey 154 

Pq,ranomus Kiesenwetter 181 

Paranove].sis Casey 197 

Paranthonomus Dietz 343 

Paraopsiraus Champlain & Knull 

Parapach;yto Casey 288 

Parapanscopus Buchanan 334 

280 

Paraplinthus Faust 338 

Paraptochus Seidlitz 332 

Paraquedius Casey 122 

Parargutor Casey 32 

Parascydmus Casey 83 

Parastenus Heyden 110 

Paratachys Casey 29 

Paratenetus Spinola 262 

Paratrogophloeus Hatch 94 

Paratropa Vacher de Lapouge 10 

Paratychius Casey 346 

Parazonitis Enns 265 

Parelaphidion Skiles 281 

Parenthis Dietz 350 

Paria LeConte 308 

Parilendus Casey 242 

Parmenini 294 

Parmulus Harold 228 

ParocaleaBernhauer 105 

Paromalini 138 

Paromalus Erichson 138 

Paropsimeno Motschulsky 311 

Parothius Casey 123 

Pasimachus Bonelli 17 

Passalidae 145 

Passalotaenlus Kuwert 145 

Passandrinae 2I9 

Patrobini 29 

Patroboidea Van Dyke 30 

Patrobus Dejean 29 

Pauronemognatha Enns 265 

Paussini 19 

Pectotoma Hatch 276 

Pedetes Kirby 182 

Pediacus Shuckard 219 

Pedilldae 274 

Pedilinae 274 

Pedilophorini 160 

Pedilus Fischer von Waldheim 274 

Peditatus Pacheco 170 

Pelates }lorn 77 

Pelatines Cockerell 77 

Pelecomalium Casey 90 

Pelecotoma Fischer von Waldheim 250 

Pelecotominae 250 

Pelec;ryhorus Solier 254 

Pelenomus Thomson 351 

Pelidnota Macleay 155 

Pelidnotidia Casey 155 

Pelophila Dejean 13 

Peltastica Mannerheim 196 

Peltasticinae 196 

Peltideo Motschulsky 207 

Peltinae 207 

Peltini 207 

Peltodytes R6gimbart 61 

Pemphus Motschulsky 13 

Penetini 253 

420

Pentacladocera Pierce 366 

Pentagonica Schmidt-Goebel 50 

Pentagonicini 50 

Pentagrammaphila Pierce 366 

Pentanota Bernhauer 100 

Pentaphyllus Dejean 256 

Pentaria Mulsant 276 

Pentariini 276 

Pentarthrum Wollaston 355 

Pentaxocerq. Pierce 366 

Pentazocera Pierce 366 

Penthe Newman 243 

PenthelispaPascoe 252 

Penthinae 243 

Pentodontini 156 

Pentozoe Pierce 366 

Peranus Lewis 139 

Perapion Wagner 328 

Percosia Zimmermann 36 

Pergetus Casey 274 

Perigaster Dietz 349 

Perigona Castelnau 51 

Perigonini 51 

Perimecus Stephens 184 

Perimegatoma Horn 198 

Peristethus LeConte 33 

Peritelini 332 

Peritelinus Casey 332 

PerthalycraHorn 216 

Peryphanes Jeannel 24 

Peryphodes Casey 25 

Peryphus Stephens 24 

Petaliini 204 

Petalium LeConte 204 

Pezocrosita Jacobson 312 

Phacophallus Coiffait 123 

Phaedon Megerle von Miihlfeld 312 

Phaenocerus Bonvouloir 187 

Phaenog'yra Seevers 107 

Phaenonotum Sharp 133 

Phaenops Dejean 165 

Phalacridae 226 

Phalacrini 226 

Phalacrus Paykull 226 

Phaleriini 254 

Phaleromela Reitter 254 

Pharalus Casey 40 

Phausis LeConte 191 

Phelister Marseul 140 

Phellidius LeConte 255 

Phellopsis LeConte 252 

Pheloconus Roelofs 347 

Phengodes Illiger 190 

Phengodidae 190 

Phenolia Erichson 216 

Phidola Thomson 295 

Philq.s Portevin 85 

Philhygra Mulsant & Rey 102 

Philodes LeConte 4? 

Philonthini 117 

Philonthus Curtis 117 

Philopedon Schiinherr 330 

Philophuga Motschulsky 58 

Philopuntia Pierce 347 

Philostrq.tus Zimmerman 338 

Philothermus Attb6 227 

PhiloxylonLeConte 202 

Phlaeopterus Motschulsky 90 

Phloeocharinae 95 

Phloeonaeus Erichson 93 

Phloeonomus Heer 88 

Phloeophagus Schtinherr 355 

Phloeopora Erichson 105 

Phloeopterus LeConte 90 

Phloeosinini 358 

Phloeosinus Chapuis 358 

Phloeostiba Thomson 88 

Phloeotribini 358 

Phloeotribus Latreille 358 

Phloeotrya Stephens 248 

Phlogistosternus Wolcott 210 

Phobetus LeConte 153 

Phomalus Casey 275 

Phosphaenus Castelnau 191 

Photinini 190 

Photinus Castelnau 190 

Photurinae 191 

Photuris Dejean 191 

Phratora Chevrolat 313 

Phrenapatinae 253 

Phryganophilus Sahlberg 247 

Phrypeus Casey 28 

Phthora Mulsant 253 

Phycocoetes LeConte 337 

Phyla Motschulsky 21 

Phfi.etes Redtenbacher 255 

Phylethus Redtenbacher 255 

PhyIIis Gistel 310 

Phyllobaenini 208 

Phyllobaenus Dejean 208 

Phyllobaenus Spinola 210 

Phyllobiini 332 

Phyllobius Germar 332 

Phyllobrotica Chewolat 316 

Phyllodecta Kirby 313 

Phyllodrepa Thomson 88 

Phyllophaga Harris 152 

Phyllotreta Chevrolat 316 

Phyllotrox Schtinherr 341 

Phymaphora Newman 238 

Phymatinus LeConte 334 

Phymatodes Mulsant 283 

Phymatodina Casey 283 

Physocnemum Haldeman 284 

Physonota Boheman 322 

Physonotini 322 

427

Physorhini 184 

Phytobiini 350 

Phytobius Schonherr 350 

PhytodectaKirby 313 

Phytoecini 299 

PhytonNewman 281 

Phytonomu,s Schiinherr 336 

Phytosini 107 

Phyxelis Schtinherr 333 

Piazorhinini 345 

Piazorhinus Schiinherr 345 

Pidonia Mulsant 292 

Piestinae 92 

Piestini 92 

PilemaLeConte 282 

Pilopius Casey 129 

Pinacodera Schaum 59 

Pinodytes Florn 80 

Pinophilini 113 

Piosoma LeConte 40 

Pisenus Casey 243 

Pissodes G'ermar 338 

Pissodinae 338 

Pityobini 176 

Pityobius l,eConte 176 

Pityogenes,Bedel 360 

Pityokteines Fuchs 360 

Pityomacer Kuschel 324 

Pityophagrrs Shuckard 217 

Pityophthorus Eichhoff 363 

Placonotusr Macleay 219 

Placopterus Wolcott 209 

Placusa Erichson 107 

Plagiodera Chevrolat 312 

Plagiometriona Spaeth 323 

Planesus Motschulsky 59 

Plataphodes Ganglbauer 22 

Plataphus Motschulsky 22 

Platerodini 188 

Plateros Bourgeois 189 

Plateumarini 302 

Plateumaris Thomson 302 

Platidiolus Chaudoir 30 

Platidius Chaudoir 30 

Platybregmus Fisher 203 

Platycerini 144 

Platyceropsis Benesh 144 

Platycerus Geoffroy 144 

Platycis Thomson 189 

Platydema Castelnau & Brull6 

Platydracus Thomson 120 

Platylomaius Cooman 138 

Platymedon Casey 114 

Platynidius Casey 56 

Platynini 51 

Platynomicrus Casey 53 

Platynus Bonelli 55 

Platypatrobus Darlington 29 

255 

Platypodidae 365 

Platypodinae 365 

Platypodini 365 

Platypsyllus Ritsema 82 

Platypsyllus Westwood 82 

Platypus Herbst 365 

Platysoma Leach 140 

Platysomatini 140 

Platystethus Mannerheim 94 

Platytrachelus Motschulsky 21 

Platyusa Casey 106 

Plectographo Rivalier 10 

Plectralidus Casey 40 

Plectrura Mannerheim 294 

Plegaderini 136 

Plegaderus Erichson 136 

Plesiocis Casey 245 

Pleurophoro Chewolat 304 

Pleurophorus Mulsant 149 

Pleuropompha LeConte 264 

Pleurostica Motschulsky 312 

Pleurotobia Casey 107 

Plinthini 338 

Plinthodes LeConte 333 

Plocamus LeConte 353 

Plocetes LeConte 346 

Pocadius Erichson 216 

Podabrini 192 

Podabrus Westwood 192 

Podapion Riley 329 

Poecilium Fairmaire 283 

Poecilobrium Horn 282 

Poecilochroo Chevrolat 209 

Poecilus Bonelli 30 

Pogonocherini 295 

Pogonocherus Dejean 296 

Polemius LeOonte 194 

Poliaenus Bates 295 

Polistoma Casey 105 

Pollaclasis Newman 191 

Polycaon Castelnau 199 

Polycaonini 199 

Polycestinae 161 

Polycestini 161 

Polyclasis LeConte 191 

Polyderis Motschulsky 29 

Polydrosus Schdnherr 330 

Polydrosini 330 

Polydrusus Germar 330 

Polygrammo Chewolat 311 

Polygraphini 359 

Polygraphus Erichson 359 

Polyphaga 74 

Polyphylla Harris 152 

PolyspilaHope 310 

Polystoma Stephens 105 

PomphopoeaLeConte 264 

Pontomalota Casey 103 

422

Poophagus Schiinherr 350 

Popillia Audinet-Serville 155 

Porcinolus Mulsant l3z Rey 159 

Poreospasta Horn 2134 

Porotachys Netolitzky 29 

Porrhodites Kraatz',91 

Potamodytes Zimmermann 66 

Potamonectes Zimmermann 66 

Prasocuris Latreille 312 

Priacma LeOonte 6 

Priacmini 6 

Priobium Motschulsl

Pseudota Casey 101 

Pseudothysanoes Blackman 359 

Pseudotrtaeno Minsk & Hatch 39 

Pseudotrox Robinson 150 

Pseudotychlus Blatchley 328 

Pseudousipalia Lohse 102 

Pseudow eisea Schwarz 229 

Pseudoxenos Saunders 367 

Pseudoxyletinus Pic 203 

Pseudoxyporus Nakane & Sawada 

Psiloscelis Marseul 140 

Psoa Herbst 200 

Psoinae 199 

Psoini 2(t0 

Psomophrrs Casey 83 

Psomus Clasey 348 

Psydrus LeConte 29 

Psylliodes Berthold 321 

Psyllobora Dejean 237 

Psylloborini 237 

Psyrassa Pascoe 280 

Ptenidium Erichson 75 

Pterna Seidlitz 260 

Pterocolirrae 326 

Pterocolus Say 326 

Pteroloma Gyllenhal 77 

Pteropalus Casey 43 

Pterostichini 30 

Pterostichus Bonelli 31 

Pteryngium Reitter 221 

Ptiliidae 75 

Ptiliinae 75 

Ptilininae 203 

Ptilinini 203 

Ptilinus I\fiiller 203 

Ptiliola Haldeman 75 

Ptiliolum Flach 75 

Ptiliopycra Casey 76 

Ptilium Gyllenhal 75 

Ptilodact5'la Illiger 169 

Ptilodacty'lidae 169 

Ptinidae 205 

Ptininae 205 

Ptinodes LeConte 202 

Ptinus Linn6 206 

Ptomaphagini 81 

Ptomaphagus Illiger 81 

Ptomister Houlbert & Monnot 139 

Pubimodes Park I27 

Pullus Mulsant 230 

PunctagonumCasey 53 

Purpuricenini 286 

Purpuricenus Dejean 286 

Pycnoglyprta Thomson 88 

Pycnomerini 252 

Pycnomerus Erichson 252 

Pycnoruotida Casey 254 

Pycnophus Casey 83 

109 

Pycnoplectus Casey 126 

Pycnorus Casey 114 

Pycnotomina Casey 218 

Pygoleptura Linsley & Chemsak 291 

PyladiaWeise 318 

Pyractomena Melsheimer 190 

Pyrectosomo Motschulsky 191 

Pyres Champion 258 

Pyrilloxenos Pierce 366 

Pyrochroidae 269 

Pyrochroinae 269 

Pyrophorinae 175 

Pyrophorini 175 

Pyropyga Motschulsky 191 

Pytotrichus LeConte 288 

Pyrrhalta Joannis 314 

Pyrrocis Castelneau 243 

Py.thidae 268 

Pybhinae 268 

Py'tho Latreille 268 

Pytna Casey 129 

Quadrifrons Blatchley 216 

Quediellus Casey L22 

Quediini 120 

Quediochrus Casey 121 

Quedionuchus Sharp 121 

Quedius Stephens 121 

Rafonus Casey 126 

Ramecia Casey 126 

RamonaCasey 113 

Rantogiton des Gozis 70 

Raphirus Stephens 122 

Redistes Casey 252 

Reesa Beal 198 

Refonia Casey 32 

Reichenbachia Leach 128 

Rembus Dejean 48 

Reminius Casey 267 

Remus Holme 119 

Renardia Motschulsky 92 

Renocis Casey 358 

Reuelstohea Hatch 90 

Rhabdophorus Leftvre 308 

Rhabdopterus Lefbvre 308 

Rhadalinae 212 

Rhadine LeConte 52 

Rhaeboscelis LeConte 167 

Rhagium Fabricius 289 

Rhagomicrus Fleutiaux 187 

RhagonychaEschscholtz 193 

Rhanidea Strohecker 238 

Rhanis LeConte 238 

Rhantus Dejean 70 

Rhaphiona Fischer von Waldheim 15 

Rhaxonycha Motschulsky 193 

Rhinocyllini 337 

424

Rhinocyllus Germar 337 

Rhinoncus Schiinherr 351 

Rhinosimus Latreille 270 

Rhipiceridae 143 

Rhipidandrini 254 

Rhipidandrus LeConte 254 

Rhipiphoridae 250 

Rhipiphorino Casey 157 

Rhipiphorinae 250 

Rhipiphorus Bosc dAntic 250 

Rhizophagidae 218 

Rhizophaginae 218 

Rhizophagus Herbst 2l-8 

Rhodobaenus LeConte 354 

Rhombodera Reiche 50 

Rhombonalia Casey 154 

Rhopalapion Schilsky 329 

Rhopalelophorus Kuwert 130 

Rhopalomerus Chewol at 285 

Rhopalopaclzys Chevrolat 285 

Rhynchaeninae 345 

Rhynchaenus Clairville 345 

Rhynchitidae 326 

Rhynchitinae 326 

Rhynchitini 326 

Rhynchophorinae 354 

Rhyncolini 355 

Rhyncolus Germar 356 

Rhysodidae 7 

Rhysothorax Bedel 147 

Rhyssemus Mulsant 149 

Rhyssomatus Schijnheru 347 

Rhytirrhininae 334 

Rhyzobius Stephens 233 

Rhyzopertha Stephens 200 

RileyaOlivier 191 

Riponus Casey 278 

RomaleumWhite 280 

Ropalopus Mulsant 284 

Rosalia Audinet-Serville 282 

Rosaliini 282 

Rosella Whitehead 346 

Roualida Casey 103 

Rugilus Leach 114 

Rushia Forel 247 

Rutelini 155 

Rutelinae 154 

Rutidosoma Stephens 350 

Rybaxis Saulcy 128 

Rypobiini 228 

Sachalinobia Jacobson 289 

Saciinae 228 

Saciini 228 

Sacium LeConte 228 

Sa.codes LeConte 142 

Salebius Casey 22I 

Salpingidae 270 

Salpinginae 270 

Salpingus auct. 270 

Sandalus Knoch 143 

Sanfilippody'tes Franciscolo 65 

Saperda Fabricius 298 

Saperdini 298 

Sapintus Casey 272 

Saprininae 136 

Saprinus Erichson 136 

Saprobia Ganglbauer 216 

Saprophilus Streubel 120 

Sarosesthes Thomson 285 

Sarpedon Bonvouloir 187 

Saxinis Lacordaire 305 

Scalopterus Motschulsky 2\2 

Scaphidema Redtenbacher 254 

Scaphidiidae I24 

Scaphidiini 125 

Scaphidium Olivier 125 

Scaphinotus Dejean 12 

Scaphisoma Leach 125 

Scaphisomini 125 

Scaphium l{trby I25 

Scarabaeidae 145 

Scarabaeinae 146 

Scarabaeini 146 

Scarabaeoidea 143 

Scarites Fabricius 17 

Scaritini 17 

Scelolyperus Crotch 316 

Schedarosus Reitter 255 

Schistogloss a I{r aatz lO2 

Schizogenius Putzeys 18 

Schizophilus Bonvouloir 188 

Schizotus Newman 269 

Sciaphilus Schiinherr 330 

Scierus LeConte 356 

SciodrepaThomson 81 

Sciodrepoides Hatch 81 

Sciopithes Horn 331 

Scirtes Illiger 142 

Scirtidae 142 

Scleropterini 350 

Scobicia Lesne 200 

Scoliocerus Motschulsky 178 

Scolytidae 356 

Scolytinae 359 

Scolytini 359 

Scolytus Geoffroy 359 

Scopaeina 114 

Scopaeoma Casey 114 

Scopaeopsis Casey 114 

Scopaeus Erichson 114 

Scopophus Casey 83 

Scotias Czenpinski 205 

Scotobates Rye 258 

Scotochroa LeConte 247 

Scotochroides Mank 247 

425

Scotocryptini 79 

Scraptia Latreille 276 

Scraptiidae 275 

Scraptiinae 276 

Scraptiini 276 

Sculptotheca Schilsky 204 

Scydmaenidae 82 

Scydmaeninae 82 

Scydmaenrni 83 

Scydmaenr-rs Latreille 83 

Scymninaer 229 

Scymnini 229 

Scymnobius Casey 231 

Scymnus Kugelann 230 

Scythropus Schtlnherr 330 

Selenalius Casey 43 

Selenophorus Dejean 43 

Semanotus Mulsant 283 

Semicampa Netolitzky 27 

Semiclivina Kult 18 

Semijulistus Schilsky 212 

Sepedophilus Gistel 97 

Serangiini 229 

Serica Ma

Steninae 109 

Stenister Reichardt 139 

Stenistoderus Jacquelin du Val 123 

Stenocantharus Gistel 13 

Stenocellus Casey 46 

StenochiaKirby 261 

Stenocorus Geoffroy 289 

Stenocorus Fabricius 288 

Stenocranophilus Pierce 367 

Stenocrepis Chaudoir 49 

Stenolophus Stephens 47 

Stenopodius Horn 322 

Stenopterini 282 

Stenoscelis Wollaston 355 

Stenosphenus Haldeman 280 

Stenostrophia Casey 292 

Stenotarsinae 238 

Stenotarsini 238 

StenothoraxHarris 154 

Stenotrachelinae 263 

Stenotrachelus Latreille 263 

Stenous Chaudoir 49 

Stenura Dejean 292 

Stenuris Kirby 162 

Stenus Latreille 109, 110 

Stephanocleonus Motschulsky 336 

Stephanoderes Eichhoff 362 

Stephanopachys Waterhouse 200 

Stephostethus LeConte 240 

Stereagonum Casey 55 

Steremnius Schtinherr 338 

Stereocerus Kirby 31 

Stereopalpus LaFert6-S inectdre 27 4 

Steriphanus Casey 253 

Sternidius LeConte 297 

Sternuchus LeConte 141 

Steroderus Motschulsky 32 

Steropus Dejean 32 

Stethobaris LeConte 353 

Stethon LeConte 187 

Stethorhanis Blaisdell 238 

Stethorini 229 

Stethorus Weise 229 

Sthereus Motschulsky 338 

Sticholotidinae 229 

Stichosoma Motschulsky 311 

Stictalia Casey 107 

Stictanchus Casey 55 

Stictocranius LeConte 112 

Stictoleptura Casey 293 

Stictolinus Casey 124 

Stigrnatoma LeConte 260 

Stilbolemrna Harris 151 

Stilbolidus Casey 45 

Stilbus Seidlitz 226 

Stilicina 114 

Stilicosoma Casey 114 

Stolaini 322 

Stomis Clairville 30 

Stornphax Fischer von Waldheim 149 

Storualota Mannerheim 101 

Storthephora Miiklin 262 

Strabala Chewolat 317 

Strangalepta Casey 293 

Strangalia Audinet-Serville 290 

Strangalina Aurivillius 290 

Strepsiptera 366 

Stricticomus Pic 273 

Strigocis Dury 245 

Strigoderma Burmeister 155 

Strigodermella Casey L55 

Strongylaspis Spaeth 323 

Stronryliini 261 

Strongylium Kirby 261 

Strongylocassis Hincks 323 

Strophiona Casey 294 

Strophogastra Fenyes 103 

Strophosoma Billberg 330 

Strophosomini 330 

Stylopidae 367 

Stylopidia 366 

Stylopinae 367 

Stylops Kirby 367 

Subhaida Hatch 89 

Sulcacis Dury 245 

Sumitrosis Butte 321 

Sunius Curtis 113 

Suphisellus Crotch 62 

Symbiotes Redtenbacher 238 

Symphora LeConte 247 

Synaphaeta Thomson 295 

Syncalosomo Breuning 10 

Syncalyptinae 158 

Synchita Hellwig 251 

Synchitini 251 

Synchroa Newman 267 

Synchroidae 267 

Synchytodes Crotch 251 

Syndesinae 144 

SynerthaDietz 340 

Syneta Dejean 307 

Synetinae 307 

Synolabus Jekel 326 

Synstrophus Seidlitz 246 

Syntomium Curtis 93 

Syntomus Hope 58 

Synuchus Gyllenhal 51 

Systena Chewolat 317 

Tachinoderus Motschulsky 96 

Tachinus Gravenhorst 96 

Tachistodes Casey 48 

Tachyerges Schiinherr 345 

Tachymenis Motschulsky 29 

Tachyporinae 96 

Tachyporini 96 

427

Tachyporu.s Gravenhorst 97 

Tachypterellus FaII & Cockerell 343 

Tachypteru.s Dietz 343 

Tachypus Dejean 20 

Tachys Stephens 29 

Tachyta Kirby 28 

Tachyura Motschulsky 28 

Taenioglyptes Bedel 362 

Taenobqtes Motschulsky 260 

Tanaocarabus Reitter 12 

Tanaops LeConte 212 

Tanarthropsis Casey 273 

Tanarthrus LeConte 273 

Tanilotes Casey 275 

Tanygaster Blatchley 316 

Tanymecirri 331 

Tanymecus Germar 331 

Tanyrhinus Mannerheim 91 

Tanysphyrini 340 

Tanysphyrus Germar 340 

Taphranch.us Casey 53 

Taphrocerus Solier 168 

Taphroscydmus Casey 83 

Tapinosthenes Kolbe 10 

Taraxis LeConte 304 

Tarpela Bates 258 

Tarphiota Casey 103 

Tarsosteninae 2ll 

Tarsostenrrs Spinola 211 

Taxicerellc,. Casey 100 

Tecnophilus Chaudoir 59 

Tectosternum Balfour-Browne 134 

Tedinus Crasey 259 

Teinodactylo Chevrolat 317 

Telacis Poey 259 

Telephanus Erichson 220 

Telephorus Schaeffer 193 

Telmatophilinae 22L 

Telmatophilus Heer 221 

TemnochilaWestwood 207 

Tenebrio L,inn6 259 

Tenebrioni.dae 253 

Tenebrioninae 258 

Tenebrionini 258 

Tenebrionoidea 242 

Tenebroides Piller & Mitterpacher 208 

Tenomergil Neboiss 6 

Tentyriinae 253 

Teredini 252 

Teretriini 136 

Teretrius Erichson 136 

Tesnus Rey 112 

Tetartoperrs Czwalina 115 

Tetragonorlerus Dejean 56 

Tetraleucus Casey 52 

Tetralina (lasey 100 

Tetraopes Schtjnherr 299 

Tetraopini 299 

Tetrascapha Schuster & Marsh 126 

Tetratoma Fabricius 243 

Tetratomidae 243 

Tetratominae 243 

Tetropium Kirby 279 

Teuales Casey 90 

Texania Casey 161 

Thalasselephas Egorov & Karotyaev 

Thalycra Erichson 216 

Thanasimus Latreille 209 

Thanatophilus Leach 85 

Thaneroclerini 209 

Thecesterninae 335 

Thecesternus Say 335 

Thermonectus Dejean 72 

Thersalus Pascoe 280 

Thes Semenov 239 

Thesalia Casey 292 

Thesium Casey 126 

Thicanus Casey 213 

Thinobius Kiesenwetter 94 

Thinodromus Kraatz 94 

Thinopinus LeConte 120 

Thinusa Casey 108 

Thoracophorus Motschulsky 92 

Thricolepoides O'Brien 330 

Thricomigus Horn 333 

Thrimolus Casey 243 

Throscidae 186 

Throscinae 186 

Throscus Latreille 186 

Thyamis Stephens 317 

Thylodrias Motschulsky 199 

Thylodriini 199 

Thymalus Latreille 207 

Thy sanocnemis LeConte 345 

Tilargus Casey 243 

TileaFauvel 9O 

Tillini 208 

Tilloclytus Bates 286 

Tillomorphini 286 

Timarcha Latreille 310 

Timarchini 310 

Tipnus Thomson 205 

Tmesiphorini 129 

Tmesiphorus LeConte 129 

Tomq.rus LeConte 224 

Tomicini 357 

Tomoxia Costa 248 

Toraminae 224 

Toramim 224 

Toramus Grouvelle 224 

Torrentomus Bierig 94 

Tosastes Sharp 334 

Toxidium LeConte 125 

Toxotopsis Casey 288 

Toxotropis LeConte 325 

Toxotus Dejean 288 

337 

428

Trachelonepha Casey 22 

Trachodes Germar 338 

Trachyinae 168 

Trachykele Marseul 163 

Trachypachidae 7 

Trachypachus Motschulsky 7 

Trachyphloeini 332 

Trachyphloeus Germar 332 

Trachysectus Casey 113 

Trachysida Casey 292 

Tragidion Audinet-Serville 286 

Tragosoma Audinet-Serville 278 

Tragosomini 278 

Trechicus LeConte 51 

Trechini 19 

Trechonepha Casey 21 

Trechus Clairville 19 

Treiodous Dugds 263 

Trepanedoris Netolitzky 27 

Triachus LeConte 307 

Triaena LeConte 39 

Triarthron Markel 78 

Tribalinae 138 

Triboliini 258 

Tribolium Macleay 258 

Tribonia Rivalier 8 

Tribrachys LeConte 2I4 

Trichalophus LeConte 333 

Trichapion Wagner 329 

Trichelaphrus Semenov 16 

Trichesthes Erichson 152 

Trichiinae 157 

Trichiini 158 

Trichiotinus Casey 158 

Trichischius LeConte 354 

Trichiusa Casey 103 

Trichobaris LeConte 352 

Trichocanthus Motschulsky 120 

Trichocellus Ganglbauer 46 

Trichochrous Motschulsky 2I3 

Trichocnemis LeConte 27 7 

Trichodes Herbst 210 

Trichodesma LeConte 202 

Tricholochmaea Laboissidre 314 

Trichophya Mannerheim 99 

Trichophyinae 99 

Trichoplataphus Netolitzky 23 

Trichopteryx Kirby & Spence 76 

Trichoscelis Bates 286 

Trichotichnus Morawitz 43 

Tricoryninae 205 

Tricorynini 205 

Tricorynus Waterhouse 205 

Tricrania LeConte 265 

Tricraniodes Wellman 265 

TrigaFauvel 95 

Trigites Handlirsch 95 

Trigonarthris Haldeman 293 

Trigonodemus LeConte 91 

Trigonogenius Solier 205 

Trigonorhinus Wollaston 324 

Trigonoscuta Motschulsky 330 

Trigonurini 92 

Trigonurus Mulsant 92 

Triliarthrus Casey 46 

Trimioplectus Brendel 127 

Triplacinae 225 

Triplax Herbst 225 

Triplectrus LeConte 44 

Trirhabda LeConte 314 

Tritoma Fabricius 225 

Trixagus Kugelann 186 

Trogastrini 126 

Troginae 150 

Trogoderma Dejean 197 

Trogophloeus Nannerheim 94 

Trogossita Olivier 208 

Trogossitidae 206 

Trogossitinae 207 

Trogossitini 207 

Trogoxylon LeConte 201 

Tropicini 170 

Tropicus Pacheco 170 

Tropideres Sch0nherr 325 

Tropiderini 325 

Tropiphorus Schiinherr 334 

Tropisternus Solier 133 

TrotomrnaLeConte 247 

Trox Fabricius 150 

Trymochthebius Kuweft 7 4 

Trypherus LeConte 195 

Trypodendron Stephens 361 

Trypophloeus Fairmaire 362 

Turboscymnus Gordon 231 

Tychiinae 345 

Tychiini 346 

Tychini 129 

Tychius Germar 346 

Tychobythinus Ganglbauer l-28 

Tylicini 160 

Tylicus Casey 160 

Tylistus LeConte 204 

Tyloderma Say 347 

Tylonotus Haldeman 280 

Tympanophorus Nordmann 120 

Typhaea Stephens 243 

Typhloleiodes Hatch 80 

Typitium Casey 262 

Typocerus LeConte 291 

Tyrini 129 

TyrusAub6 129 

Uleiota Latreille 220 

Uleiotini 220 

Ulitusa Casey 107 

Ulochaetes LeConte 288 

429

Uloma Ltrtreille 257 

Ulomini 257 

Unamis Casey 90 

Upis Fabricius 261 

Urgleptes Dillon 298 

Urograph',is Horn 297 

Uroplatini 322 

Utobium FalI 202 

LJvarus Guignot 64 

Vacusus Casey 273 

Vanonus Casey 275 

Variopalpis Solier 58 

Vellica Casey 90 

Venusa Casey 107 

Veraphis Casey 83 

Vicelva l\:[oore & Legner 95 

Vincenze.llus Reitter 270 

Vrilletta LeConte 203 

WillinkiaBondar 353 

Xanthochroa Schmidt 266 

Xanthochroina Ganglbauer 266 

Xanthoga.leruca Laboissidre 314 

Xantholirrinae 123 

Xantholinini 123 

Xantholirrus Dejean 124 

Xanthonia Baly 309 

Xanthoph,ius Motschulsky 123 

Xanthopl'gini 120 

Xanthus Dietz 344 

Xeninae 367 

Xenochara Mulsant & Rey 104 

Xenocharis Bierig 113 

Xenodacne Boyle 225 

Xenodusa Wasmann 106 

Xenoscelinae 224 

Xenota Mulsant & Rey 103 

Xerosaprinus Wenzel 137 

XestipygerMarseul 138 

Xestobiini 202 

Xestobiurn Motschulsky 202 

Xestocis Casey 244 

Xestoleptura Casey 293 

Xestolinus Casey 124 

Xestonotrrs LeConte 45 

Xestophu;s Casey 83 

Xyleborini 361 

Xyleborinus Reitter 362 

Xyleborus Eichhoff 362 

Xylechinus Chapuis 357 

Xyletininae 203 

Xyletinini 203 

Xyletinus Latreille 204 

Xylita Paykull 247 

Xylobiops Casey 200 

Xylobius Latreille 188 

Xylocomesus Thatcher 356 

Xylocrius LeConte 284 

Xylodromus Heer 91 

Xyloecus Dejean 188 

Xyloperthini 200 

Xylophilus Mannerheim 188 

Xylopinus LeOonte 260 

Xyloryctes Hope 156 

Xylosandrus Reitter 361 

Xyloterini 361 

Xyloterinus Swaine 361 

Xylotrechus Chevrolat 285 

Yrdops Steffan 171 

Zabrini 35 

Zacotus LeConte 19 

ZaitzeviaChampion 173 

ZalepiaArnett 175 

Zalobius LeConte 95 

Zeadolopus Broun 79 

Zenodosus Wolcott 209 

Zeugophora Kunze 304 

Zeugophorinae 304 

ZezeaCsiki 39 

ZiloraMulsant 247 

ZonantesCasey 275 

Zonitina 265 

Zonltinae 265 

Zonitini 265 

Zonitis Fabricius 265 

Zonoptilus Solsky 93 

Zopheridae 252 

Zophefinae 252 

Zuphiini 59 

Zuphium Latreille 59 

Zygogramma Chewolat 3 10 

Zygopinae 348 

Zygopini 348 

ZyrasStephens 106 

Recycled ,pn 

,*r1. 

faper GP 

*""" 

430

Canad'd

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