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Catalog of species-group names of Recent and fossil Scaphopoda ...

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Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)AcronymNYSMNZGSPIRSPIUVPRIQMRGMRSMSAFMSAMSBMNHSHMSMFSMNHSMTTMHUCMUCMPUIMGUMTUMZCUNAMUOUPMNUSNMUWMWAMWSMYPMZINZMAZMBZMOZMUCZMUUZSIInstitutionNew York State Museum, Albany, USANew Zeal<strong>and</strong> Institute <strong>of</strong> Geological <strong>and</strong> Nuclear Science, Lower Hutt (NZ Geological Survey), NewZeal<strong>and</strong>Primary Industries <strong>and</strong> Resources <strong>of</strong> South Australia, Adelaide, AustraliaPaläontologisches Institut, Universität Wien, AustriaPaleontological Research Institution, Ithaca, New York, USAQueensl<strong>and</strong> Museum, Brisbane, AustraliaRijksmuseum van Geologie en Mineralogie, Leiden, Netherl<strong>and</strong>sRoyal Scottish Museum, Edinburgh, UKSouth African Museum, Cape Town, South AfricaSouth Australian Museum, Adelaide, AustraliaSanta Barbara Museum <strong>of</strong> Natural History, USASaito Ho-on Kai Museum, JapanNatur-Museum Senckenberg, Frankfurt, GermanyNaturhistoriska Riksmuseet, Stockholm, SwedenSakurai Museum, Tokyo, Japan; transferred to NSMTTasmanian Museum <strong>and</strong> Art Gallery, Hobart, AustraliaUniversity <strong>of</strong> Colorado Museum, Boulder, USAUniversity <strong>of</strong> California Museum <strong>of</strong> Paleontology, Berkeley, USAUniversity <strong>of</strong> Ife, Museum <strong>of</strong> Geology, NigeriaUniversity Museum Tokyo, JapanUniversity Museum <strong>of</strong> Zoology, Cambridge, UKInstituto de Geologia, Universidade Nacional Autónoma de Mexico, Mexico City, MexicoUniversity <strong>of</strong> Oregon, Eugene, USAMuseo di Paleontologica, Università di Napoli, ItalyNational Museum <strong>of</strong> Natural History, Washington, D.C., USABurke Museum <strong>of</strong> Natural History & Culture, University <strong>of</strong> Washington, Seattle, USAWestern Australian Museum, Perth, AustraliaWashington State Museum, Seattle, USAYale Peabody Museum, Yale, USAZoological Institute, Academy <strong>of</strong> Sciences, St Petersburg, RussiaZoölogisch Museum Amsterdam, Netherl<strong>and</strong>sZoologisches Museum Berlin, GermanyZoologisk Museum, Universitetet i Oslo, NorwayZoologisk Museum Copenhagen, DenmarkZoologiska Museet, Uppsala Universitet, SwedenZoological Survey <strong>of</strong> India, CalcuttaThe Cenozoic mollusk card catalogs housed bythe Department <strong>of</strong> Paleobiology, NationalMuseum <strong>of</strong> Natural History, SmithsonianInstitution (USNM), are invaluable resources thatmerit wider notice. During the middle part <strong>of</strong> the20th century, Barbara A. Bedette <strong>and</strong> DruidWilson, paleontologists associated with the USGeological Survey, maintained a card catalog,comprising 4” × 6” cards with typed informationon the Atlantic Gulf Coastal Plain <strong>species</strong> <strong>of</strong>Cenozoic mollusks from the southeastern UnitedStates (approximately, Maryl<strong>and</strong> to Texas).Meanwhile, another US Geological Survey paleontologist,Wendell P. Woodring, similarlymaintained a card catalog, comprising 3” × 5”cards with h<strong>and</strong>written information on the<strong>species</strong> <strong>of</strong> Cenozoic mollusks described from“tropical America” (the isl<strong>and</strong>s <strong>of</strong> the Caribbean;Mexico; Central America; <strong>and</strong> the northern part<strong>of</strong> South America). Both card catalogs containedentries for <strong>fossil</strong> scaphopod <strong>species</strong> described inthe 20th century that were not recorded in theZoological Record <strong>and</strong> were overlooked in the subsequentliterature. Although these card catalogshave not been updated for some years, researcherswho visit the Smithsonian collections may wishto examine these files.The catalog consists <strong>of</strong> 1965 entries. There are828 entries <strong>of</strong> <strong>Recent</strong> scaphopod <strong>species</strong>-<strong>group</strong><strong>names</strong>, 517 <strong>of</strong> which are here considered validZOOSYSTEMA • 2004 • 26 (4)553


Steiner G. & Kabat A. R.<strong>species</strong>. The entries are alphabetically ordered bytheir original binomen following the scheme:<strong>species</strong>, original genus Author, year: page. Stratigraphy<strong>of</strong> type locality. Type locality [translatedinto English <strong>and</strong>/or modern equivalent as needed].Depository <strong>of</strong> type material. Synonymy.Current taxonomic status (reference if differentfrom original combination). Geographic distribution.Bathymetric range. Remarks.Names <strong>of</strong> valid <strong>Recent</strong> <strong>species</strong> are in boldface.Names <strong>of</strong> the 816 valid <strong>fossil</strong> <strong>species</strong>-<strong>group</strong><strong>names</strong> are preceded by a dagger (†) symbol. Thesynonymies given are based upon publisheddeterminations. Where contradictory opinionshave been published, one is given preference <strong>and</strong>the alternative is noted in the Remarks.Synonymies, current status <strong>and</strong> distribution aregiven only for a few <strong>fossil</strong> taxa. Synonymies donot include misidentifications, as their completelisting is beyond the scope <strong>of</strong> this paper. We havenot attempted to resolve all transfers <strong>of</strong> syntypes<strong>and</strong> paratypes subsequent to the original descriptions,a problem caused by the fact that suchtypes were sometimes split <strong>of</strong>f from the originallots long after the original descriptions (especiallyat the museums <strong>of</strong> the United States <strong>of</strong> America);thus, types are <strong>of</strong>ten located in depositories notcited in the original publication. Additionalinformation such as modern locality <strong>names</strong>,depths in m, references cited by Linnaeus <strong>and</strong>Gmelin, etc., is given in square brackets. Boss(1988) provided full references for the <strong>of</strong>tencrypticabbreviations used by Linnaeus (1758,1767) <strong>and</strong> many <strong>of</strong> those used by Gmelin(1791). Current generic assignments are tentativefor many <strong>species</strong> due to the lack <strong>of</strong> anatomicaldata needed to distinguish certain genera.The generic assignments <strong>of</strong> <strong>Recent</strong> <strong>species</strong> thathave not been recently studied were based uponthe generic diagnoses in Scarabino (1995). Informationon bathymetric ranges can be problematic,since published records do not alwaysdifferentiate between live-collected specimens<strong>and</strong> empty shells, the latter <strong>of</strong> which can betransported to deeper depths before beingdredged. Hence, researchers interested in determiningthe bathymetric range for a given <strong>species</strong>are advised to examine the relevant specimens inmuseum collections to ascertain their taphonomicstatus.For the sake <strong>of</strong> completeness, we have includedincorrect subsequent spellings in this catalog.They refer to the corresponding valid name.Although these are not available <strong>names</strong> (ICZN1999: article 33.3), their inclusion herein willreadily allow future generations <strong>of</strong> researcherswho find these <strong>names</strong> in the older literature todetermine their status from this catalog.Table 1 lists the abbreviations <strong>of</strong> type specimendepositories taken from Kabat & Boss (1992:188-232), with some additions. The currentdepositories <strong>of</strong> type material were, in some cases,difficult to determine. Information was obtainedfrom the original descriptions, from catalogs <strong>of</strong>type specimens, on-line searches in museumdatabases, correspondence with curators, <strong>and</strong>from visits to The Natural History Museum(London), Muséum national d’Histoire naturelle(Paris), <strong>and</strong> the National Museum <strong>of</strong> NaturalHistory (Washington, D.C.). The most importantlists <strong>of</strong> types <strong>of</strong> <strong>Recent</strong> scaphopod <strong>species</strong>,among others listed by Kabat & Boss (1992: 243;1997: 354), are Turner (1955), Palmer (1958),Dance (1967), Zeidler & Macphail (1978), Giles& Gosliner (1983), Oliver (1984), Scott et al.(1990), Spamer & Bogan (1992), Wallin (1992),Kilias (1995), <strong>and</strong> Boyko & Sage (1996). Otherimportant sources <strong>of</strong> information on types <strong>of</strong><strong>fossil</strong> scaphopod <strong>species</strong> are Newton (1891),Beecher (1900), Maury (1917), Rossi Ronchetti(1955), Flügel (1959), Ludbrook (1959), Brann& Kent (1960), Hanzawa et al. (1961), Keyes(1972), Ferrero Mortara et al. (1984), <strong>and</strong> White(1998). It was not always possible to obtain informationon the current status <strong>of</strong> type material (i.e.if a lectotype had been designated) for severaltaxa. If the original publication <strong>and</strong> any relevantsubsequent publications, or correspondence withmuseum curators, did not yield information onthe status <strong>of</strong> type specimens, then the entry reads“Type material in [museum]” without greaterspecificity Due to the relocation <strong>and</strong> re-cataloging<strong>of</strong> the Mollusca collection in the NationalMuseum <strong>of</strong> Natural History (Washington,554ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)TABLE 2. — Number <strong>of</strong> <strong>species</strong>-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Recent</strong> <strong>and</strong> <strong>fossil</strong> <strong>Scaphopoda</strong> for the most prolific authors, including originaldescriptions <strong>and</strong> replacement <strong>names</strong>. *, including co-authored <strong>species</strong> <strong>names</strong>.<strong>Recent</strong> Fossil TotalAuthor No. Author No. Author No.Henderson 51 Sacco 67 Pilsbry* 99Scarabino 47 Pilsbry* 54 Sacco 67Lamprell & Healy 46 Cossmann* 42 Henderson 51Locard 45 Deshayes 29 Dall 49Pilsbry* 45 Meyer* 20 Scarabino 47Boissevain 39 Martin 19 Lamprell & Healy 46Dall 37 Gardner 18 Locard 45Watson 31 Tate 18 Cossmann* 45Sowerby* 30 Koenen 16Habe* 19 Chenu 14Jeffreys 19 Palmer C. P. 13Plate 17 Dall 12Chistikov 14 D’Orbigny 12Janssen R. 11D.C.), paratype catalog numbers for several<strong>species</strong> (especially <strong>of</strong> Henderson’s types, whichsometimes shared catalog numbers with otherlots) are not available at present.Boissevain (1906), in her monograph on theSiboga Expedition scaphopods, introduced39 <strong>species</strong>-level <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong>. Unfortunately,this monograph had a number <strong>of</strong> typographicalerrors regarding the numbers <strong>of</strong>specimens <strong>and</strong> dredging station numbers forsome <strong>of</strong> the type material; furthermore, some lectotypedesignations by subsequent authors havenot always clearly identified the precise specimenthat was to be the lectotype. Robert Moolenbeek<strong>and</strong> A. N. Van der Bijl (ZMA) have thoroughlyreviewed the Siboga type material <strong>and</strong>Boissevain’s annotated copy <strong>of</strong> her monograph.Based upon this review, they have carefully reconciledthe discrepancies in this monograph <strong>and</strong>in the subsequent literature regarding the typematerial. We have incorporated their correctionsinto our catalog.A comparison <strong>of</strong> the most prolific authors <strong>of</strong><strong>Recent</strong> <strong>species</strong> (Table 2) shows that Hendersonintroduced 52 new <strong>names</strong>; Scarabino 47 <strong>names</strong>;Lamprell & Healy 46 <strong>names</strong>; Locard <strong>and</strong> Pilsbry45 <strong>names</strong> each; Boissevain introduced 39 <strong>names</strong><strong>of</strong> <strong>Recent</strong> <strong>species</strong>-<strong>group</strong> taxa. Sacco coined 67<strong>fossil</strong> <strong>species</strong>-<strong>group</strong> <strong>names</strong>; Pilsbry <strong>and</strong> his coauthors54 <strong>names</strong>; Cossmann <strong>and</strong> his co-authors42 <strong>names</strong>.Determining the correct locations <strong>of</strong> type depositories,synonyms <strong>and</strong> validity <strong>of</strong> <strong>Recent</strong> taxa havereceived considerably greater attention than <strong>of</strong><strong>fossil</strong> taxa, primarily because <strong>of</strong> the neontologicalbackground <strong>and</strong> interests <strong>of</strong> the first author. Forthe same reason, stratigraphic ranges <strong>of</strong> <strong>fossil</strong><strong>species</strong> are not provided, beyond the informationin the original description.The taxonomic research activity in <strong>Scaphopoda</strong>,measured in <strong>names</strong> introduced per decade, variedconsiderably through time (Fig. 1). After lownumbers <strong>of</strong> <strong>species</strong> described during the late 18th<strong>and</strong> early 19th centuries, first the number <strong>of</strong>newly described <strong>fossil</strong> <strong>species</strong> increased in thewake <strong>of</strong> prospecting <strong>and</strong> mining activities in thecourse <strong>of</strong> the industrial revolution. With theonset <strong>of</strong> ocean floor sampling expeditions, thenumbers <strong>of</strong> described <strong>Recent</strong> <strong>species</strong> followedsuit. The 1890s feature the unique peak inscaphopod research with over 280 newly introduced<strong>names</strong>, most <strong>of</strong> which originated in thepublications <strong>of</strong> Pilsbry, Locard, Sacco, <strong>and</strong>Cossmann. The decline in research activity in the20th century is most conspicuous in the decades<strong>of</strong> World Wars I <strong>and</strong> II. Taxonomic activityrebounded in the second half <strong>of</strong> the past century<strong>and</strong> peaked in its final decade mainly due to theZOOSYSTEMA • 2004 • 26 (4)555


Steiner G. & Kabat A. R.1701601501401301201101009080706050403020100<strong>Recent</strong>Fossil17016015014013012011010090807060504030201001750-17591760-17691770-17791780-17891790-17991800-18091810-18191820-18291830-18391840-18491850-18591860-18691870-18791880-18891890-18991900-19091910-19191920-19291930-19391940-19491950-19591960-19691970-19791980-19891990-19992000-2004FIG. 1. — Number <strong>of</strong> <strong>species</strong>-<strong>group</strong> <strong>names</strong> introduced per decade for <strong>fossil</strong> <strong>and</strong> <strong>Recent</strong> taxa. The period from 2000 to 2004 includesthe <strong>names</strong> introduced herein.monographs <strong>of</strong> Scarabino (1995) <strong>and</strong> Lamprell& Healy (1998). With increased sampling activitiesin the tropical Indo-Pacific region, the level<strong>of</strong> descriptions <strong>of</strong> new <strong>species</strong> will likely be sustainedin the first decade <strong>of</strong> this century. So far,in the period from 2000 to 2004, 18 new <strong>names</strong>were published, including those introduced inthe present paper.The present contribution to scaphopod taxonomyis not intended as a global revision but constitutesa documentation <strong>of</strong> the status quo. Toobtain an accurate estimate <strong>of</strong> the total number<strong>of</strong> valid <strong>species</strong> <strong>and</strong> the number <strong>of</strong> <strong>species</strong> in eachgenus, it became necessary to include eighthitherto unpublished synonyms <strong>and</strong> eightnew combinations. These include several byB. Métivier (in litt.) <strong>and</strong> V. Scarabino (in litt.).The appendices <strong>of</strong> the catalog summarize the rectifications<strong>of</strong> nomenclature with the renamedhomonyms (Appendix 1) <strong>and</strong> the arguments fornew synonymies <strong>and</strong> new generic combinations(Appendix 2). The listing <strong>of</strong> valid <strong>Recent</strong> <strong>species</strong>for each genus (Appendix 3) is updated fromSteiner & Kabat (2001). Finally, Appendix 4 liststhe presumably valid <strong>fossil</strong> taxa by their stratigraphicage.This catalog is the most complete publishedcompilation <strong>of</strong> <strong>species</strong>-level <strong>names</strong> in the class<strong>Scaphopoda</strong>. However, we recognize that, giventhe abundance <strong>of</strong> this class in the <strong>Recent</strong> <strong>and</strong> <strong>fossil</strong>marine fauna, <strong>and</strong> the proliferation <strong>of</strong> publicationsthat have been overlooked by recorders, itis not possible for this catalog to be 100%complete. In particular, we suspect that there arevalid <strong>names</strong> described by Chinese <strong>and</strong> Russianpaleontologists that we have not found (<strong>and</strong> arenot recorded in the Zoological Record), becausewe did not have the time to search through all therelevant publications listed in the RussianReferativnii Zhurnal or the Chinese GushengwuxueWenzhai [Paleontological Abstracts] (seealso Bouchet & Rocroi 1992, 1993). For thatreason, we do not intend that the present catalogbe considered for inclusion in a “List <strong>of</strong> AvailableNames in Zoology” pursuant to ICZN (1999)article 79.556ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)LIST OF SPECIES-GROUP NAMESabbreviatum, Dentalium Deshayes, 1825. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.aberrans, Cadulus Whiteaves, 1887: 124, fig. 2.<strong>Recent</strong>. Dawson stn No. 20, Forward Inlet,Quatsino Sound, north-west coast <strong>of</strong> VancouverIsl<strong>and</strong> [British Columbia, Canada], 10-20 fms [18-37 m]. Syntypes NMC 555 (5). Junior synonyms:C. hepburni Dall, 1897; C. fusiformis Pilsbry &Sharp, 1898; C. nitentior Arnold, 1903 (fideShimek 1989: 234, 235). Gadila aberrans(Whiteaves, 1887) (fide Pilsbry & Sharp 1898:193). Eastern Pacific: Alaska to SouthernCalifornia. 7-365 m. Remarks: Shimek (1989: 235)erroneously designated USNM 133809 as a “lectotype<strong>of</strong> C. aberrans”. Emerson in Turgeon (1998:55) places this in Cadulus.abruptoinflatus, Cadulus (Gadila) Boissevain, 1906:75, pl. 6, fig. 65, text-fig. 39. <strong>Recent</strong>. Madura Strait[Indonesia], Siboga stn 5, 7°46’S, 114°30.5’E,330 m. Holotype ZMA 3.06.106. Gadila abruptoinflata(Boissevain, 1906) (fide Scarabino 1995:364). Indo-Pacific: Indonesia. 330 m.† abruptus, Cadulus Meyer & Aldrich, 1886: 40, pl. 2,fig. 2. Oligocene. Newton <strong>and</strong> Wautubbee,Mississippi, USA. Syntypes USNM Paleobiology638725, 638726. Dischides abruptus Meyer &Aldrich, 1886 (fide Palmer & Brann 1965: 360,361). Remark: Aldrich (1895: 4) <strong>and</strong> Pilsbry &Sharp (1898: 235) as a probable junior synonym <strong>of</strong>Cadulus subcoarctuatus (Gabb, 1860).† absconditum, Dentalium Deshayes, 1861: 212, 213,pl. 1, figs 15-17. Eocene. Paris Basin (Parnes,Chaumont, Saint-Félix <strong>and</strong> Mouchy), France.Antalis abscondita (Deshayes, 1861) (fide Le Renard1995: 175; Le Renard & Pacaud 1995: 85).abyssicola, Helonyx subfusiformis var. Monterosato,1875: 20, 21. <strong>Recent</strong>. Palermo [Sicily], 210 m.Type material not located. Junior synonym <strong>of</strong>Cadulus jeffreysi (Monterosato, 1875) (fide Pilsbry& Sharp 1898: 163). Remark: Monterosato (1878:77) elevated this to a full <strong>species</strong>, but subsequently(1880: 65) treated this as a junior synonym <strong>of</strong>Helonyx subfusiformis (M. Sars, 1865).abyssorum, Dentalium M. Sars, 1859: 52. <strong>Recent</strong>.Norwegian coast. Type material not located. Juniorsynonym <strong>of</strong> Antalis occidentalis (Stimpson, 1851)(fide Thorson & Spärck 1939: 2).† acicula, Fustiaria (Episiphon) Hodgkinson, 1974:22, 23, text-figs 7c-e, 8i-k; pl. 3, figs 2-6, 8, pl. 7,fig. 3, pl. 8, figs 3, 5a, b. Middle Eocene,Claibornian Stage, Cook Mountain Formation,Wheelock Member. Little Brazos Bluff, BrazosRiver Valley, Brazos County, Texas, USA.Holotype USNM Paleobiology 180456; paratypesUSNM Paleobiology 180451-180455, 180479,PRI 29230, ANSP 31508 (all from Scott’s localityMS-2-N, Little Brazos River).aciculatum, Dentalium Hall, 1860. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.aciculum, Dentalium Gould, 1859: 165. <strong>Recent</strong>.Coast <strong>of</strong> China, 23°50’N, 25 fms [45 m]. LectotypeUSNM 24149 (designated by Johnson 1964: 36, as“holotype”); paralectotypes ANSP 35539 (1), MCZ169034 (ex NYSM 145) (1), MCZ 216577 (1).Junior synonym: Dentalium luchuanum Dall, 1926(fide Habe 1964: 24). Dentalium aciculum Gould,1859. Indo-Pacific: China, Japan, PhilippineIsl<strong>and</strong>s, Indonesia, eastern Australia. 5-1000 m.Remark: Pilsbry & Sharp (1897: 93) placed this<strong>species</strong> in Graptacme, but V. Scarabino (in litt.5.XI.2001) has determined that this is correctlyplaced in Dentalium.aciculum, Dentalium Deshayes, 1861: 202, 203, pl. 1,figs 31, 32. Eocene. Paris Basin (Grignon <strong>and</strong>Montmirel), France. Junior homonym <strong>of</strong>Dentalium aciculum Gould, 1859. Replacementname: Dentalium acre Sharp & Pilsbry in Pilsbry &Sharp, 1898.acompsus, Cadulus (Polyschides) quadridentatusHenderson, 1920: 100, 101, pl. 17, fig. 4. <strong>Recent</strong>.Monkey River, British Honduras [Belize].Holotype ANSP 76557; paratypes ANSP 390502,USNM 314939 (not located). Junior synonym <strong>of</strong>Polyschides tetraschistus (Watson, 1879) (fideScarabino 1980: 11).† acre, Dentalium Sharp & Pilsbry in Pilsbry & Sharp,1898: 197. Eocene. Paris Basin (Grignon <strong>and</strong>Montmirel), France. Replacement name forDentalium aciculum Deshayes, 1861, a junior homonym<strong>of</strong> Dentalium aciculum Gould, 1859.† acriculum, Entalis Tate, 1887: 192, pl. 20, fig. 11.Lower Miocene. Muddy Creek, Victoria, Australia.Holotype <strong>and</strong> 8 paratypes SAM T 251. Junior synonym:Dentalium lacteolum Tate, 1899 (fideLudbrook 1959: 144). Dentalium (Laevidentalium)acriculum (Tate, 1887) (fide Ludbrook 1959: 144,145, pl. 1, fig. 2).actiniophorum, Fissidentalium Shimek, 1997: 181-190, figs 1-6. <strong>Recent</strong>. Pulse Project, stn 1809 M,34°44’N, 123°12’W, 4100-4134 m <strong>of</strong>f SouthernCalifornia. Holotype LACM 2792; paratypesLACM 2793, LACM 2809-2811, USNM 886326,BMNH 1996120-21. Fissidentalium actiniophorumShimek, 1997. Eastern Pacific: California.4100-4134 m.aculeatum, Dentalium Sowerby, 1860: 100, pl. 225,fig. 63. <strong>Recent</strong>. No information on type locality,type material <strong>and</strong> distribution. Antalis aculeata(Sowerby, 1860) (fide Pilsbry & Sharp 1897: 61).† acumen, Entalis“?” de Koninck, 1883: 216, pl. 49,fig. 22. Carboniferous. Visé, Belgium. Dentaliumacumen (de Koninck, 1883) (fide Pilsbry & Sharp1898: 229).acuminatum, Dentalium Deshayes, 1825: 369, pl. 17,figs 19, 20. Eocene. Parnes <strong>and</strong> Mouchy-le-Châtel,Paris Basin, France. Junior synonym <strong>of</strong> DentaliumZOOSYSTEMA • 2004 • 26 (4)557


Steiner G. & Kabat A. R.fissura Lamarck, 1818 (fide Deshayes 1864: 213).Fustiaria fissura (Lamarck, 1818) (fide Newton &Harris 1894: 65).acuminatum, Dentalium Brown, 1849: 243, pl. 98,fig. 74. Name for Dentalium entalis sensu J.Sowerby 1814. Tertiary. Hordwell Cliffs, Engl<strong>and</strong>,United Kingdom. Junior homonym <strong>of</strong> Dentaliumacuminatum Deshayes, 1825. Objective junior synonym<strong>of</strong> Dentalium entaloides Fleming, 1825.acuminatum, Dentalium Gardner, 1878: 62, pl. 3,figs 34-39. Cretaceous, Gaultian. Folkestone,Engl<strong>and</strong>, United Kingdom. Junior homonym <strong>of</strong>Dentalium acuminatum Deshayes, 1825 <strong>and</strong>Dentalium acuminatum Brown, 1849. Replacementname: Dentalium gardneri Sharp & Pilsbry inPilsbry & Sharp, 1898, itself a junior homonym <strong>of</strong>Entalis gardneri (Holzapfel, 1888); replacementname: Dentalium infortunatum Pilsbry & Sharp,1898.acuminatus, Cadulus “Deshayes” Angas, 1871: 97.<strong>Recent</strong>. Middle Harbour, Port Jackson, New SouthWales, Australia. Nomen nudum (fide Ludbrook1978: 86). Remark: it seems unlikely that Angas hadmisidentified an Australian <strong>species</strong> with Deshayes’name, since D. acuminatum Deshayes, 1825 doesnot resemble any <strong>species</strong> assigned to Cadulus.† acuminatus, Cadulus Tate, 1887: 194. Oyster beds<strong>of</strong> the Upper Aldinga series [Late Pliocene <strong>of</strong>Australia]. Holotype SAM T 231A. Gadila acuminata(Tate, 1887) (fide Ludbrook 1959: 147).Indo-Pacific: South Australia. 47-164 m.acus, Cadulus Dall, 1889: 432, pl. 27, fig. 11. <strong>Recent</strong>.Samana Bay, Santo Domingo, 16 fms [29 m].Lectotype USNM 95379 (designated by Henderson1920: 140); paralectotypes USNM 887454; MCZ7751, AMNH 148337 (2), AMNH 148342 (8).Gadila acus (Dall, 1889) (fide Pilsbry & Sharp1898: 191). Western Atlantic: Caribbean, Florida,Bahamas, Brazil. 20-550 m. Remark: Turner (1955:317) erroneously stated MCZ 7751 to be the “holotype”(fide Boyko & Sage 1996: 29).acus, Dentalium Eichwald, 1857. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.acus, Dentalium Cooke, 1885: 274, 275. <strong>Recent</strong>. Gulf<strong>of</strong> Suez. Holotype UMZC I.100,735. Junior homonym<strong>of</strong> Dentalium acus Eichwald, 1857.Replacement name: Dentalium cookei Sharp &Pilsbry in Pilsbry & Sharp, 1897: 29.† acutangularis, Dentalium sexangulare var. Cocconi,1874: 645. Pliocene. Castell’Arquato, Italy.acuticosta, Dentalium – Nyst 1835: 36 non Deshayes1825. Oligocene. Near Antwerp, Belgium. Nextavailable name: Dentalium kickxii Nyst, 1843.† acuticostata, Dentalium Deshayes, 1825: 357, pl. 18,fig. 3. Eocene. London, Engl<strong>and</strong>, Great Britain.Replacement name for Dentalium striatum J.Sowerby, 1814, a junior homonym <strong>of</strong> Dentaliumstriatum Born, 1778. Objective junior synonym:Dentalium bartonense Palmer, 1974 (n. syn.).acuticostatum, Dentalium (Graptacme) Plate, 1908a:352, pl. 30, fig. 37. <strong>Recent</strong>. Dar-es-Salaam,Tanzania, East Africa, Valdivia stn 244, 5°55.8’S,39°1.2’E, 50 m. Holotype ZMB 61082. Juniorhomonym <strong>of</strong> Dentalium acuticostata Deshayes,1825. Replacement name Graptacme acutistriatan. nom.acutissimum, Dentalium Watson, 1879: 514, 515[Watson 1886: 8, pl. 1, fig. 8]. <strong>Recent</strong>. North <strong>of</strong>Papua New Guinea, Challenger stn 218, 2°33’S,144°04’E, 1070 fms [1957 m]. Lectotype BMNH1887.2.9.31 (designated by Ludbrook 1954: 110,as holotype); paralectotype BMNH 1887.2.9.32.Graptacme acutissima (Watson, 1879) (fide Pilsbry& Sharp 1897: 94). Indo-Pacific: East Africa toAustralia, Japan <strong>and</strong> New Caledonia. 865-3749 m.Remarks: paralectotype BMNH 1887.2.9.33(Challenger stn 246, 36°10’N, 178°0’E, east <strong>of</strong>Japan, 2050 fms) is a different <strong>species</strong> (Scarabino1995: 243; Lamprell & Healy 1998: 98). Lamprell& Healy (1998: 97) obviously overlookedLudbrook’s lectotype designation <strong>and</strong> selected thesame specimen as lectotype.acutistriata, Graptacme n. nom. <strong>Recent</strong>. Dar-es-Salaam, Tanzania, East Africa, Valdivia stn 244,5°55.8’S, 39°1.2’E, 50 m. Holotype ZMB 61082.Replacement name for Dentalium (Graptacme) acuticostatumPlate, 1908 non D. acusticostataDeshayes, 1825. Graptacme acutistriata Steiner &Kabat, 2004. Known from type locality only.Remark: Kilias (1995: 171) noted that Plate gavethe incorrect station number, i.e. stn 247, but thecorrect coordinates.acutisulcatum, Dentalium Gurley, 1883: 7.Carboniferous, “Lower Coal Measures”. Newport,Vermillion County, Indiana, USA. Syntypes FMUC.6314 (2) (fide Forney & Nitecki 1976: 11).Junior synonym <strong>of</strong> Prodentalium obsoletum (Hall,1858), the oldest available name for Dentalium sublaeveHall in Miller, 1877, a junior homonym <strong>of</strong>Dentalium dentalis var. sublaeve Cocconi, 1874 (fideYancey 1978: 307). Objective junior synonym:Prodentalium fredericae Engeser & Riedel, 1992.acutoides, Dentalium Emerson, 1954: 183. Jurassic,Pliensbachian. Railway-cutting, Dixton West, nearGotherington, near Cheltenham, Engl<strong>and</strong>, UnitedKingdom. An unnecessary replacement name forDentalium acutum Richardson, 1906, a junior homonym<strong>of</strong> Dentalium acutum Hébert, 1849.Objective junior synonym <strong>of</strong> Dentalium richardsoniCossmann, 1907.† acutum, Dentalium Hébert, 1849: 469. Oligocene.Dego, Tertiary Piedmont Basin [Italy]. Junior synonyms:Dentalium nysti d’Orbigny, 1852 (fidePilsbry & Sharp 1898: 197); Dentalium (Entalis)acuta var. apenninica Sacco, 1897 (fide Bonci et al.2000: 206, 207). Antalis acuta (Hébert, 1849) (fideBonci et al. 2000: 206, 207, pl. 1, fig. 4). Remark:The specimens Deshayes (1864: 205, pl. 20, figs 1-4)558ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)misidentified as Dentalium acutum belong toDentalium novaki Koenen, 1892 (fide Cossmann1892: 330).acutum, Dentalium Richardson, 1906: 574, pl. 45,figs 10, 11. Jurassic, Pliensbachian. Railway-cutting,Dixton West, near Gotherington, nearCheltenham, Engl<strong>and</strong>, United Kingdom. Juniorhomonym <strong>of</strong> Dentalium acutum Hébert, 1849.Replacement <strong>names</strong>: Dentalium richardsoniCossmann, 1907; Dentalium acutoides Emerson,1954.† addicotti, Cadulus (Platyschides “?”) Emerson, 1957:990, pl. 126, fig. 2. Miocene, Temblor stage. 1 mileSW <strong>of</strong> mouth <strong>of</strong> Kern River Gorge, Kern County,California, USA. Holotype UCMP 37584; paratypesUCMP 37585-UCMP 37589.† adelaidense, Siphonodentalium (Pulsellum) Ludbrook,1956: 4, pl. 1, fig. 1. Pliocene. HindmarshBore <strong>and</strong> Dry Creek S<strong>and</strong>s, South Australia.Holotype SAM F 15141; 4 paratypes SAM.adenense, Dentalium (Dentalium) Ludbrook, 1954:97, fig. 2. <strong>Recent</strong>. Gulf <strong>of</strong> Aden, John Murray Exp.stn 28, 12°00’N, 50°38’42”E, 201 m. HolotypeBMNH 1952.3.25.124; paratypes BMNH1952.2.25.252-261, BMNH 1952.2.25.251(stn 103, 101 m), BMNH 1952.3.25.262 (stn 180,397 m), BMNH 1952.3.25.263-272 (stn 179B,275 m), BMNH 1952.3.25.46-52 (stn 185,2000 m). Dentalium adenense Ludbrook, 1954.Red Sea. 201-2000 m. Remark: BMNH1952.3.25.124 carries the label “stn 185, 2000 m”.adenensis, Entalina Ludbrook, 1954: 112, fig. 17.<strong>Recent</strong>. Gulf <strong>of</strong> Aden, John Murray Exp. stn 185,2000 m; 13°48’06”-36”N, 49°16’48”-24”E.Holotype BMNH 1952.3.25.20; paratype BMNH1952.3.25.21. Entalina adenensis Ludbrook, 1954.Known from type locality only.aegeum, Dentalium Watson, 1879: 509, 510 [Watson1886: 2, pl. 1, fig. 2]. <strong>Recent</strong>. Off London River,Kerguelen Isl<strong>and</strong>s, Challenger stn 149 (8), 48°50’S,69°18’E, 110 fms [198 m]. Holotype BMNH1887.2.9.8. Fissidentalium aegeum (Watson, 1879)(fide Pilsbry & Sharp 1897: 69). Indian Ocean. 88-2926 m.aenigmaticum, Dentalium Jordan, 1895: 264, 265,pl. 16, fig. 1. <strong>Recent</strong>. Faroe Channel, “cold area”,Triton stn 8 [60°18’N, 06°15’W, fide Warén 1980:59], 640 fms [1174 m]; <strong>of</strong>f West coast <strong>of</strong> Irel<strong>and</strong>,1000 fms [1828 m]. Syntype(?) NMW. Junior synonym<strong>of</strong> Antalis occidentalis (Stimpson, 1851) (fideFriele & Grieg 1901: 49).† aequale, Dentalium Deshayes, 1861: 205, pl. 20,figs 5-7. Eocene. Paris Basin (Cuise-la-Motte,Laversine, Laon, Osly <strong>and</strong> Cuisy-en-Almont),France. Remark: a valid <strong>species</strong> <strong>of</strong> Antalis (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 85).aequalis, Cadulus Dall, 1881: 34. <strong>Recent</strong>. Off Tortugas,Blake stn 43, 24°08’N, 82°51’W, 339 fms[618 m]. Lectotype USNM 95373 (designated byHenderson 1920: 109); paralectotypes USNM887455, MCZ 7740. Gadila aequalis (Dall, 1881)(fide Pilsbry & Sharp 1898: 170). Western Atlantic,Caribbean: Florida. 618 m. Remark: Turner (1955:318) erroneously stated that MCZ 7740 was theholotype.aequatorialis, Cadulus (Gadila) Jaeckel, 1932: 311,312, text-fig. 10. <strong>Recent</strong>. Off West Sumatra,Valdivia stn 191, 0°39.2’S, 98°52.3’E, 750 m.Lectotype ZMB 75372 (designated by Kilias 1995:171); paralectotype ZMB 75373 (stn 256, <strong>of</strong>fSomalia, 1°49’N, 45°29.5’E, 1134 m). Cadulusaequatorialis Jaeckel, 1932. Indo-Pacific: EastAfrica to Indonesia <strong>and</strong> New Caledonia. 750-1915 m.aequatorium, Dentalium (Rhabdus) Pilsbry & Sharp,1897: 112, 113, pl. 21, fig. 43. <strong>Recent</strong>. Off Manta,Ecuador, U.S. Fish Commission, stn 2792, 401 fms[720 m]. Holotype USNM 122759. Rhabdusaequatorius (Pilsbry & Sharp, 1897). Known fromtype locality only.aequicostata, Dentalium dentalis var. Foresti, 1895:252. Pliocene. Ponticello in val di Savena, nearBologna, Italy. Junior homonym <strong>of</strong> Dentaliumaequicostatum Koenen, 1892. Junior synonym <strong>of</strong>Antalis dentalis (Linnaeus, 1758). Remark: underICZN (1999) articles 31.2 <strong>and</strong> 34.2, this <strong>species</strong>name must be emended to aequicostatum in order toagree in gender with Dentalium, which renders it ajunior homonym. Its synonymy with Antalis dentalis,however, makes it unnecessary to introduce areplacement name.† aequicostatum, Dentalium Koenen, 1892: 981-983,pl. 59, figs 3, 4. Lower Oligocene. Lettorf, Calbe<strong>and</strong> Unseburg, Germany.affine, Dentalium Biondi, 1859: 120-122, pl. 1, fig. 7.<strong>Recent</strong>. Sicily, Italy. Type material not located.Junior synonym <strong>of</strong> Antalis vulgaris (Da Costa,1778) (fide Pilsbry & Sharp 1897: 42).affine, Dentalium Deshayes, 1861: 201, pl. 1, figs 12-14. Eocene. Paris Basin (Parnes), France. Juniorhomonym <strong>of</strong> Dentalium affine Biondi, 1859.Replacement <strong>names</strong>: Dentalium xiphias Pilsbry &Sharp, 1898; Dentalium desmoulinsi Le Renard,1994.affine, Dentalium Gabb, 1873: 244. Tertiary.Dominican Republic. Syntype ANSP 2711. Juniorhomonym <strong>of</strong> Dentalium affine Biondi, 1859, <strong>and</strong>Dentalium affine Deshayes, 1861. Replacementname: Dentalium gabbi Pilsbry & Sharp, 1898a.affine, Siphodentalium [sic] Gardner, 1878: 62, 63,pl. 3, figs 41-44. Cretaceous, Gaultian. Folkestone,Engl<strong>and</strong>, United Kingdom. Junior homonym <strong>of</strong>Siphonodentalium affine M. Sars, 1865; replacementname: Entalina gardneri Sharp & Pilsbry in Pilsbry& Sharp, 1898.affine, Siphonodentalium M. Sars, 1865a: 300, 301,pl. 6, figs 34, 35. <strong>Recent</strong>. L<strong>of</strong>oten Isl<strong>and</strong>s[Norway], 100-300 fms [183-548 m]. SyntypesZOOSYSTEMA • 2004 • 26 (4)559


Steiner G. & Kabat A. R.SMNH 4625 (L<strong>of</strong>oten, 200-300 fms [365-548 m]).Pulsellum affine (M. Sars, 1865) (fide Pilsbry &Sharp 1897: 140). Northern Atlantic. 70-3400 m.africanum, Dentalium Sowerby, 1903: 224, 225, pl. 5,fig. 10. <strong>Recent</strong>. Natal, South Africa, north <strong>of</strong>Umtwalumi [Mtwalume] River, 25 fms [45 m].Holotype BMNH 1903.7.27.54; paratypes BMNH1903.7.27.55-56, SAFM A5489 (A5491), ZMB59876, NMW. Graptacme africana (Sowerby,1903) (fide Scarabino 1995: 244). Indian Ocean:South Africa, Madagascar. 45-384 m.agassizi, Dentalium (Dentalium) Pilsbry & Sharp, 1897:26, 27, pl. 12, figs 90-94. <strong>Recent</strong>. Gulf <strong>of</strong> Panama,Albatross stn 3393, 7°15’N, 79°36’W, 1020 fms [1862m]. Holotype USNM 122985; paratypes USNM601273. Dentalium agassizi Pilsbry & Sharp, 1897.Eastern Pacific: California to Panama. 757-2322 m.agassizii, Cadulus Dall, 1881: 35. <strong>Recent</strong>. South <strong>of</strong>Marquesas Key, Florida, Blake stn 5, 24°05’N,82°13’W, 229 fms [418 m]. Holotype MCZ 7764.Synonym: Cadulus agassizii hatterasensis Sharp &Pilsbry in Pilsbry & Sharp, 1898 (n. syn.). Gadilaagassizii (Dall, 1881) (fide Pilsbry & Sharp 1898:168). Western Atlantic: Florida to Maine. 40-545 m. Remark: Emerson in Turgeon (1998: 55)places this in Polyschides.agile, Dentalium G. O. Sars, 1872: 31-34, pl. 3, figs 4-15 [ex M. Sars MS]. Nomen nudum in M. Sars(1869: 257 [14]). <strong>Recent</strong>. L<strong>of</strong>oten Isl<strong>and</strong>s, Norway.Syntypes ZMO 32725, 25955-56, 26034 (16),26035 (44). Name for Dentalium incertum sensuPhilippi 1844, a junior homonym <strong>of</strong> Dentaliumincertum Deshayes, 1825. Junior synonym:Dentalium fusticulus Brugnone, 1876 (fideMonterosato 1880: 64); Dentalium vagina Jeffreys,1877 (fide Pilsbry & Sharp 1897: 46). Antalis agilis(G. O. Sars, 1872) (fide Pilsbry & Sharp 1897: 46).Eastern Atlantic: Icel<strong>and</strong> <strong>and</strong> Norway to AscensionIsl<strong>and</strong>, <strong>and</strong> North Africa, Mediterranean; WesternAtlantic: Canada to Caribbean. 60-5000 m.agulhasense, Dentalium Plate, 1908a: 349, pl. 30,figs 21-23. <strong>Recent</strong>. Cap Agulhas [South Africa],Valdivia stn 95, 34°51’S, 19°37.8’E, 80 m.Lectotype ZMB 61087a (designated by Kilias 1995:172); paralectotype ZMB 61087b. Junior synonym<strong>of</strong> Dentalium strigatum Gould, 1859 (fide Barnard1974: 742).† akasakensis, Dentalium Hayasaka, 1925: 107, 108[23, 24], pl. 8 [1], figs 17, 17a. Permian.Kinshozan, Akasaka-Machi, Mino province, Japan.Holotype Inst. Geol. Paleont., Tohoku Univ. (fideHanzawa et al. 1961: 209).† alatum, Dentalium Gardner, 1878: 60, 61, pl. 3,figs 16-20. Cretaceous, Gaultian. Folkestone,Engl<strong>and</strong>, United Kingdom.† alazanum, Dentalium (Dentalium) Cooke, 1928: 9,pl. 2, figs 6, 6a. Oligocene. Alazan Clay, Rio Buenavista, west <strong>of</strong> Alazan, Vera Cruz, Mexico. SyntypeUSNM Paleobiology 352709 (1).alba, Dentalium scamnatum var. Locard, 1898: 110.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium scamnatum, a junior synonym <strong>of</strong>Fissidentalium capillosum (Jeffreys, 1877) (B.Métivier, n. syn.).albicomatus, Cadulus Dall, 1890: 295, pl. 9, fig. 8.<strong>Recent</strong>. Off Manta, Ecuador, USFC stn 2792,0°37’S, 81°W, 401 fms [731 m]. Syntypes USNM602251, USNM 87539, SBMNH 35018 (2),AMNH 146292 (1). Striocadulus albicomatus(Dall, 1890) (fide Emerson 1962: 478). EasternPacific: Ecuador, Gulf <strong>of</strong> Panama. 731-3056 m.albida, Dentalium agile var. Locard, 1898: 118.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Antalis agilis (G.O. Sars, 1872).albida, Dentalium ergasticum var. Locard, 1898: 106.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium ergasticum P. Fischer, 1883, a junior synonym<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877)(B. Métivier, n. syn.).albida, Dentalium rubescens var. Monterosato, 1875:20; 1878: 76. Nomen nudum.albida, Dentalium semivestitum var. Locard, 1898:109. <strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Fissidentalium semivestitum (P. Fischer in Locard,1898).albina, Dentalium vulgare var. Monterosato, 1884: 32.Nomen nudum without type locality or description.albina, Pseudantalis rubescens var. Monterosato, 1884:33. Nomen nudum without type locality or description.† aldrichi, Cadulus Gardner, 1933: 200, pl. 20, fig. 1.Eocene, Midway Group, Wills Point Formation.USGS stn 8245, Salado Creek, Bexar County,5.3 miles SE <strong>of</strong> San Antonio, Texas, USA.Holotype USNM Paleobiology 373066. Name forCadulus subcoarctatus sensu Aldrich 1895.† alineatum, Dentalium Stephenson, 1952: 142,pl. 34, figs 1, 2. Upper Cretaceous, Cenomanian,Woodbine Formation. Lamar County Locality 201,near old Slate Shoals, Red River, 8 miles east <strong>of</strong>Arthur City, Lamar County, Texas, USA. HolotypeUSNM Paleobiology 105603; paratypes USNMPaleobiology 105604-105606.alloschismum, Dentalium (Laevidentalium) liodon var.Pilsbry & Sharp, 1897: 108, pl. 21, figs 40-42.<strong>Recent</strong>. St Martin [Caribbean]. Holotype ANSP35552. Synonym <strong>of</strong> Fustiaria liodon (Pilsbry &Sharp, 1897) (fide Díaz 1989: 31, as Laevidentaliumliodon “forma” alloschismum; generic placement fideScarabino 1975: 185). Caribbean. 63-120 m (fideV. Scarabino, in litt. 5.XI.2001).alternans, Dentalium Chenu, 1843: pl. 4, fig. 17;1850: 1. Type locality not specified. Type materialnot located. Nomen dubium.560ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)alternans, Dentalium J. Müller, 1851: 5, pl. 3, figs 1a,1b. Cretaceous. Königsthore, Germany. Juniorhomonym <strong>of</strong> Dentalium alternans Chenu, 1843.Replacement name: Dentalium muellerianumPilsbry & Sharp, 1898.alternans, Dentalium Ryckholt, 1851: 71, 72, pl. 2,figs 45, 46. Cretaceous: Turonian. Visé, Belgium.Junior homonym <strong>of</strong> Dentalium alternans Chenu,1843, <strong>and</strong> Dentalium alternans Müller, 1851;replacement name: Dentalium confusum Sharp &Pilsbry in Pilsbry & Sharp, 1898.alternans, Dentalium Bucquoy, Dautzenberg &Dollfus, 1882: 561-564, pl. 66, figs 7-9. <strong>Recent</strong>.Roussillon [Gulf <strong>of</strong> Lion, Mediterranean coast <strong>of</strong>France]. Junior homonym <strong>of</strong> Dentalium alternansChenu, 1843. Replacement name: Dentalium inaequicostatumDautzenberg, 1891. Antalis inaequicostata(Dautzenberg, 1891) (generic placementfide Pilsbry & Sharp 1897: 52). Remark: Dentaliuminaequicostatum is itself a junior homonym <strong>of</strong>Dentalium inaequicostata Seguenza, 1879. However,we prefer not to introduce yet another name for thistaxon until its status is clear (see also Remarks forAntalis inaequicostata. Earlier replacement <strong>names</strong>turned out to be preoccupied: Dentalium dollfusiCossmann, 1899 is a junior homonym <strong>of</strong>Dentalium dollfusi Koenen, 1883. Moreover,Cossmann was unaware <strong>of</strong> Dentalium inaequicostatumDautzenberg, 1891 (fide Cossmann 1900b:186).alternans, Antale dentale var. Sangiorgi, 1926: 119,pl. 7 [1], fig. 31 [ex Berti MS]. Neogene. Ponticelladi Sàvena, near Bologna, Italy. Junior secondaryhomonym <strong>of</strong> Dentalium alternans Chenu, 1843,Dentalium alternans Müller, 1851, Dentalium alternansRyckholt, 1851, <strong>and</strong> Dentalium alternansBucquoy, Dautzenberg & Dollfus, 1882. Replacementname: Dentalium sangiorgii Emerson, 1954.Antalis sangiorgii (Emerson, 1954) (fide Caprotti1979: 237).alternatum, Dentalium Lea, 1833: 34, 35, pl. 1, fig. 2.Eocene. Claiborne, Alabama, USA. LectotypeANSP 5005 (designated by Palmer & Brann 1965:369 as “holotype”); paralectotypes ANSP 5006 to5014. Junior synonym <strong>of</strong> Dentalium thalloidesConrad, 1833 (fide Pilsbry & Sharp 1898: 218).amaliense, Dentalium (Graptacme) Henderson, 1920:71, pl. 11, figs 4, 5. <strong>Recent</strong>. St Thomas [VirginIsl<strong>and</strong>s]. Holotype <strong>and</strong> paratype USNM 250088.Junior synonym <strong>of</strong> Graptacme eborea (Conrad,1846) (fide Scarabino 1975: 185).† amalthei, Dentalium Engel, 1891: 34, pl. 3, fig. 11[as nomen nudum in Engel 1890: 42]. Jurassic:Middle Liassic. Filsbetts near Eislingen, Germany.Laevidentalium amalthei (Engel, 1891) (fide Engeser& Riedel 1992: 40, 44).ambiguum, Dentalium Chenu, 1843: pl. 3, fig. 1;1850: 1. <strong>Recent</strong>. Type locality unknown. Syntype(?)MHNG 1155/34 (fide Y. Finet in litt. 16.X.1996).Laevidentalium(?) ambiguum (Chenu, 1843) (fidePilsbry & Sharp 1897: 100). Remark: Genericplacement <strong>and</strong> systematic validity <strong>of</strong> this taxon areuncertain.americanum, Dentalium Chenu, 1843: pl. 4, figs 9,10; 1850: 1. <strong>Recent</strong>. “Côtes d’Amérique”. Syntype(?)MHNG 1155/15. Junior synonyms: Dentalium texasianumPhilippi, 1849; Dentalium picteti Pilsbry &Sharp, 1897; Dentalium texasianum rioense Henderson,1920; Dentalium texasianum cestum Henderson,1920; Dentalium rebeccaense Henderson,1920 (fide Scarabino 1975: 183). Paradentaliumamericanum (Chenu, 1843) n. comb. WesternAtlantic <strong>and</strong> Caribbean: Florida to Brazil. 5-150 m.amiantus, Cadulus Dall, 1889: 431, pl. 27, fig. 7.<strong>Recent</strong>. Off Bahia Honda, Cuba, Blake stn 19,23°03’N, 83°10’30”W, 310 fms [566 m]. LectotypeMCZ 7749 (designated by Henderson 1920:128, 129, as “the type”); paralectotypes USNM95378 (<strong>of</strong>f Cape San Antonio, Cuba, 1002 fms[1829 m]); the other type specimen cited by Dall(<strong>of</strong>f Cape Florida, 8 fms [15 m]) was not located.Gadila amianta (Dall, 1889) (fide Pilsbry & Sharp1898: 174). Western Atlantic: Caribbean. 15-1829m. Remark: Henderson (1920: 113, 114) recognizedthat the original description <strong>of</strong> Cadulus amiantusDall, 1889 encompassed two <strong>species</strong>;therefore, he described parvus for those specimens<strong>of</strong> “amiantus” that were not conspecific with amiantuss.s.amphialum, Dentalium Watson, 1879: 510 [Watson1886: 3, pl. 1, fig. 3]. <strong>Recent</strong>. Mouth <strong>of</strong> La PlataRiver, Challenger stn 323, 35°39’S, 50°47’W,1900 fms [3468 m]. Lectotype BMNH 1887.2.9.9(designated by Henderson 1920: 59 as “the type”);the second specimen mentioned by Watson was notfound. Fissidentalium amphialum (Watson, 1879)(fide Pilsbry & Sharp 1897: 71). Western Atlantic:Brazil. 1800-3740 m.amphora, Cadulus Jeffreys, 1883: 665, 666, pl. 49,figs 9-11. <strong>Recent</strong>. Porcupine 1870, Atlantic, stn 16,54°19’N, 11°50’W, 816 fms [1489 m] (fide Warén1980: 53, 57). Holotype BMNH 85.11.5.1286(fide Warén 1980: 53). Cadulus amphora Jeffreys,1883. Eastern Atlantic. 1489 m.ampullaceus, Cadulus Watson, 1879: 529 [Watson1886: 23, pl. 3, fig. 11]. <strong>Recent</strong>. Culebra Isl<strong>and</strong>,West Indies, Challenger stn 24, 18°38’30”N,65°05’30” W, 390 fms [712 m]. Holotype BMNH1887.2.9.93. Cadulus ampullaceus Watson, 1879.Western Atlantic: Caribbean. 320-3392 m.anatorum, see aratorum.† anceps, Dentalium J. Sowerby in Wetherell, 1837:135, 136, pl. 8, fig. 17. Eocene. Lower Heath,Hampstead, London, Great Britain. Antalis anceps(J. Sowerby in Wetherell, 1837) (fide Newton &Harris 1894: 66, as “Entaliopsis”).anceps, Dentalium Vinassa de Regny, 1898: 185[143], pl. 20 [7], figs 13-15 [ex Meneghini MS].ZOOSYSTEMA • 2004 • 26 (4)561


Steiner G. & Kabat A. R.Oligocene. Via degli Orti <strong>and</strong> Valle Orgagna, Italy.Junior homonym <strong>of</strong> Dentalium anceps J. Sowerby,1837. Replacement name: Dentalium vinassaiCossmann, 1912.<strong>and</strong>ersoni, Polyschides Lamprell & Healy, 1998: 170,figs 174E, 175E, 180. <strong>Recent</strong>. Off Woollongong,New South Wales, Australia, 34°27’-26’S,151°27’E, 1200 m. Holotype AMS C174896;paratypes AMS C169929 (22), AMS C173797 (2;80 km south west <strong>of</strong> Cape Martin, South Australia,37°50’-55’S, 139°15’-25’E, 1555 m), AMSC174895 (3; <strong>of</strong>f Sydney, New South Wales,33°31.33’S, 152°08-07’E, 914-907 m), AMSC169930 (1; <strong>of</strong>f Sydney, New South Wales, 33°37-39’S, 152°04.02’E, 924-896 m), AMS C169932 (9;east <strong>of</strong>f Sydney, New South Wales, 33°35-37’S,152°05’E, 1143-1106 m), AMS C169933 (3; <strong>of</strong>fNowra, New South Wales, 34°50’S, 151°15’E,841 m), AMS C169934 (5; <strong>of</strong>f Sydney, New SouthWales, 33°35’S, 152°09’E, 1097 m), AMSC169988 (7; east <strong>of</strong> Sydney, New South Wales,33°44’S, 151°57’E, 322 m). Polyschides <strong>and</strong>ersoniLamprell & Healy, 1998. Indo-Pacific: SEAustralia. 153-1555 m.† <strong>and</strong>leri, Dentalium Oppel, 1856: 93. Jurassic. NearVaihingen, Germany. Remark: Tate (1870b: 17)suggested that Dentalium <strong>and</strong>leri Oppel, 1856 “isprobably a synonym” <strong>of</strong> Dentalium portlocki Tate,1870 [Jurassic, Irel<strong>and</strong>], “but, assuming that thetwo <strong>names</strong> refer to the same <strong>species</strong>, the imperfectdiagnosis <strong>of</strong> Oppel justifies me in discarding hisname”.† <strong>and</strong>rewsi, Dentalium Cuvillier, 1933: 27, pl. 3,fig. 7. Eocene, Bartonian. Gebel Mokattam, east <strong>of</strong>Kaït-Bey, Egypt.† angelae, Prodentalium Engeser & Riedel, 1992: 41.Triassic. North <strong>of</strong> Launsdorf, Austria. Replacementname for Dentalium arctoides var. multicostataGugenberger, 1934, a junior homonym <strong>of</strong>Dentalium multicostatum Favre, 1869.† angsananum, Dentalium (s.s.) Martin, 1921: 479,pl. 60 [3], fig. 87. Miocene. Tji Angsana, nearNjalindung, Java, Indonesia. Holotype RGM 7282(fide Van den Hoek Ostende et al. 2002: 111).anguidens, Cadulus Melvill & St<strong>and</strong>en, 1898: 32,pl. 1, fig. 6. <strong>Recent</strong>. Off Madras, India. SyntypesManchester Museum (fide Trew 1987: 24). Gadilaanguidens (Melvill & St<strong>and</strong>en, 1898) (fideScarabino 1995: 364). Indo-Pacific: India, Indonesia,Japan, Red Sea. 2-732 m.† angulare, Dentalium Kaunhowen, 1897: 13, pl. 1,fig. 1. Cretaceous, Maestrichtian. Kunraed,Germany.angulati, Dentalium Quenstedt, 1852: 443. Jurassic,Liassic. Europe. Nomen nudum.† angulatum, Dentalium Buckman in Murchison,1845: 101. Jurassic, Pliensachian. Lias Marlstone,Alderton, Dumbleton <strong>and</strong> Stanway Hills, Engl<strong>and</strong>,Great Britain. Remark: Engeser & Riedel (1992: 37)concluded that “the description does not correspondwith any known scaphopod genus”.angulatum, Dentalium Vinassa de Regny, 1897: 163[ex Meneghini MS]. Tertiary. Roncà, Italy. Juniorhomonym <strong>of</strong> Dentalium angulatum Buckman inMurchison, 1845. Remarks: Vinassa de Regny consideredthe single, incomplete shell only to be a newform. We prefer not to introduce a replacementname for this dubious <strong>species</strong>.† angusticostatum, Dentalium (Antalis) Baluk, 1972:556, 557, pl. 1, figs 6-8, text-figs 5a-c. Miocene,Lower Tortonian (Badenian). Korytnica, 24 kmsouth-southwest <strong>of</strong> Kielce, Holy Cross Mountains,Pol<strong>and</strong>. Holotype <strong>and</strong> paratypes, Baluk collection.angustior, Cadulus Verco, 1911a: 211, 212, pl. 26,figs 5, 5a, b. <strong>Recent</strong>. 18 miles SE <strong>of</strong> Newl<strong>and</strong> Head,outside Backstairs Passage, South Australia, 26 fms[48 m]. Holotype SAM D 13728; possible paratypeNMW. Gadila angustior (Verco, 1911) (fideLamprell & Healy 1998: 150). South Australia. 24-667 m.angustistriatum, Paradentalium Chistikov, 1979a:110, 111, fig. 2. <strong>Recent</strong>. Vietnam, Orlik T.13,18°00’N, 109°16’E, 47 m. Holotype MMSU.Paradentalium angustistriatum Chistikov, 1979.Indo-Pacific: Vietnam. 22-151 m.† angustum, Dentalium Deshayes, 1861: 210, 211,pl. 1, figs 1-3. Eocene. Paris Basin (Grignon,Parnes, Fontenay-Saint-Père, Mouchy, Mouy <strong>and</strong>Vaudancourt), France. Remark: a valid <strong>species</strong> <strong>of</strong>Antalis (fide Le Renard 1995: 175; Le Renard &Pacaud 1995: 85).† anisicum, Dentalium Schnetzer, 1934: 32, pl. 1,figs 7-10. Middle Triassic, Anisian. Saalfelden,Austria.† annectens, Cadulus (Platyschides) Woodring, 1925:210, pl. 28, fig. 11. Miocene. Bowden, Jamaica.Holotype USNM Paleobiology 353077.annulare, Dentalium Sowerby, 1829: 199. <strong>Recent</strong>.East Indies. Types not located in BMNH. Juniorsynonym <strong>of</strong> Laevidentalium eburneum (Linnaeus,1767) (fide Pilsbry & Sharp 1897: 116).annulata, Dentalium McClell<strong>and</strong>, 1834. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.annulata, Entaliopsis Newton & Harris, 1894: 67,pl. 6, fig. 1. Eocene, London Clay. Portsmouth,Southampton <strong>and</strong> Primrose Hill, Engl<strong>and</strong>, GreatBritain. Secondary junior homonym <strong>of</strong> Dentaliumannulatum Gmelin, 1791; Dentalium annulatumCrist<strong>of</strong>ori & Jan, 1832; Dentalium annulatumS<strong>and</strong>berger, 1842; Dentalium annulatum Meyer,1886; <strong>and</strong> Entalis annulatum Tate, 1887; replacementname: Dentalium entaliopsis Pilsbry & Sharp,1898. Junior synonym <strong>of</strong> Antalis constricta (Newton& Harris, 1894) (fide Jeffery & Tracey 1997: 90, 91).† annulatum, Dentalium Gmelin, 1791: 3738 [citingGuettard (Mineral. Belustig. 4 t. 5 f. 3)]. [Fossil].Type locality not specified. Remarks: Sacco (1897:112) listed two Italian citations to this <strong>species</strong> as a562ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)potential senior synonym <strong>of</strong> Dentalium (Fustiaria)jani Hörnes, 1856. We have not found the publication<strong>of</strong> Guettard as cited by Gmelin.annulatum, Dentalium Crist<strong>of</strong>ori & Jan, 1832: 2.[Fossil]. Italy. Types not located. Junior homonym<strong>of</strong> Dentalium annulatum Gmelin, 1791. Sacco(1897: 112) listed this name together with Dentaliumannulatum Gmelin as a potential senior synonym<strong>of</strong> Dentalium (Fustiaria) jani Hörnes, 1856.We, thus, do not propose a replacement name untilthe relationships <strong>of</strong> these taxa are revised.annulatum, Dentalium Jay, 1850. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.annulatum, Dentalium S<strong>and</strong>berger & S<strong>and</strong>berger,1856: 240, pl. 26, figs 20, 20a. Nomen nudum inS<strong>and</strong>berger (1842: 399). Carboniferous. Villmar(“Stringocephalenkalk”), Germany. Junior homonym<strong>of</strong> Dentalium annulatum Gmelin, 1791;Dentalium annulatum Crist<strong>of</strong>ori & Jan, 1832.Replacement name: Dentalium annuliferum Pilsbry& Sharp, 1898.annulatum, Dentalium Meyer, 1886: 64, 65, pl. 1,fig. 1. Eocene. Claiborne, Alabama, USA. LectotypeUSNM Paleobiology 638812 (as “holotype” inPalmer & Brann 1965: 365). Junior homonym <strong>of</strong>Dentalium annulatum Gmelin, 1791; Dentaliumannulatum Crist<strong>of</strong>ori & Jan, 1832; Dentaliumannulatum S<strong>and</strong>berger & S<strong>and</strong>berger, 1856.Replacement name Dentalium parannulatumn. nom. Remark: Hodgkinson (1974: 20)concluded that this <strong>species</strong> “probably is valid”, havingrejected the conclusion <strong>of</strong> Palmer (1937: 19)that it was the juvenile stage <strong>of</strong> Dentalium minutistriatumGabb, 1860. However, neither Palmer norHodgkinson renamed this junior homonym.annulatum, Dentalium Kokay, 1966: 67, 113, pl. 10,fig. 15. Miocene, Tortonian. Bakony, Hungary.Holotype Hungarian Geological Collection, M.279.Junior homonym <strong>of</strong> Dentalium annulatum Gmelin,1791, Dentalium annulatum Crist<strong>of</strong>ori & Jan, 1832,Dentalium annulatum S<strong>and</strong>berger & S<strong>and</strong>berger,1856, Dentalium annulatum Meyer, 1886; replacementname: Dentalium bakoniense Palmer, 1974.annulatum, Entalis Tate, 1887: 191, 192, pl. 20,figs 6a, 6b. Miocene. Muddy Creek, Victoria,Australia. Holotype <strong>and</strong> 3 paratypes SAM T 250A.Junior secondary homonym <strong>of</strong> Dentalium annulatumGmelin, 1791. Replacement name: Dentaliumaustralis Sharp & Pilsbry in Pilsbry & Sharp, 1898.Laevidentalium australe (Sharp & Pilsbry in Pilsbry& Sharp, 1898) (fide Ludbrook 1959: 145).† annulatus, Cadulus Pilsbry, 1911: 168. Miocene.Bowden, Jamaica. Syntype ANSP 649. Polyschidesannulatus (Pilsbry, 1911) (fide Woodring 1925:207, 208, as Cadulus (Polyschides) annulatus).annulatus, Dentalites Krueger, 1823: 312. [Fossil].Paris, France. Remark: this taxon might correspondto Dentalium annulatum Gmelin, 1791; butKrueger did not cite Gmelin.† annuliferum, Dentalium Pilsbry & Sharp, 1898:229. Carboniferous. Villmar (“Stringocephalenkalk”),Germany. Replacement name forDentalium annulatum S<strong>and</strong>berger & S<strong>and</strong>berger,1856, a junior homonym <strong>of</strong> Dentalium annulatumGmelin, 1791.† annulopunctatum, Dentalium (Fustiaria) Cossmann& Pissarro, 1902: 180, pl. 31, fig. 39. Eocene.Fresville, Cotentin, France.† annulostriatum, Dentalium Meek & Worthen,1871: 45. Carboniferous. Danville, Illinois, USA.annulosum, see anulosum.† anomalocostata, Dentalium (Entalis) taurostriata “?”var. Sacco, 1897: 110, pl. 9, fig. 57 [as a nomen nudumin Sacco 1896: 97]. Miocene. Turin hills, Italy.Type material not mentioned by Ferrero Mortara etal. (1984: 308).† anomalum, Dentalium (Entalina) Cossmann &Peyrot, 1917: 171, 172, pl. 2, figs 19, 20. Miocene,Helvetian. Saint-Martin-de-Hinx, France.antarcticus, Cadulus dalli var. Odhner, 1931: 5-7, pl. 1,figs 5-7, 9, 11, 13, 14; pl. 2, fig. 18-21, 24, 25. <strong>Recent</strong>.Graham Region, Swedish Antarctic Exp. stn 8,64°05’S, 56°37’W, 360 m; stn 11, 65°19’S, 56°48’W,400 m. Syntypes SMNH 4196 (fide S<strong>and</strong>berg &Warén 1993: 122). Synonym <strong>of</strong> Siphonodentaliumdalli (Pilsbry & Sharp, 1898) (fide Steiner & Linse2000: 21-25). Antarctica. 130-695 m.† antarctotubulus, Cadulus (Gadila) Stilwell &Zinsmeister, 1992: 174, pl. 25, figs q, r. LowerTertiary, La Meseta Formation. IPS locality 443,Seymour Isl<strong>and</strong>, Antarctic Peninsula. HolotypeUSNM Paleobiology 441875; paratypes USNMPaleobiology 441876, 441877.antillarum, Dentalium d’Orbigny, 1853: 202, pl. 25,figs 10-13. <strong>Recent</strong>. Saint-Thomas [Virgin Isl<strong>and</strong>s].Syntypes BMNH.10.4.467 (2). Antalis antillaris(d’Orbigny, 1853) (fide Henderson 1920: 44).Western Atlantic <strong>and</strong> Caribbean: Florida to Brazil.1-300 m.† antiquum, Dentalium Goldfuss, 1841: 2, pl. 166,fig. 2. Devonian? Eifel Mountain, Germany.antiquum, Dentalium Goldfuss, 1841. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.antverpiense, Dentalium Goddeeris, 1978: 70, 72.Miocene. Belgium. Nomen nudum.anulata, Dentalium entalis var. Jeffreys, 1865: 192.<strong>Recent</strong>. British Isl<strong>and</strong>s. Type material not located.Synonym <strong>of</strong> Antalis entalis (Linnaeus, 1758).anulosum, Dentalium Brazier, 1877: 58. <strong>Recent</strong>.Princess Charlotte Bay, Queensl<strong>and</strong>, NE Australia,13 fms [24 m]. Lectotype AMS C170759 (designatedby Ponder & Stanbury 1972: 53) <strong>and</strong> 1 paralectotype.Junior synonym <strong>of</strong> Episiphon subtorquatum(Fischer, 1871) (fide Scarabino 1995: 286).Remarks: Lamprell & Healy (1998: 104) consideredanulosum to be a valid <strong>species</strong> <strong>of</strong> Omniglypta.Dentalium annulosum [Hedley 1901: 129] is anincorrect subsequent spelling.ZOOSYSTEMA • 2004 • 26 (4)563


Steiner G. & Kabat A. R.apenninica, Dentalium (Entalis) cf. acuta “?” var.Sacco, 1897: 106, 107, pl. 9, figs 14-16 [as a nomennudum in Sacco 1896: 97]. Oligocene. Dego,Piedmont Basin [Italy]. Lectotype MRSNBS.106.03.008 (designated by Bonci et al. 2000:206, 207, pl. 1, fig. 4); paralectotypes MRSNBS.106.03.009, DSTRG 1173/Sa-II-S10 (1),DSTRG 2105/Sa-II-S48 (1). Junior synonym <strong>of</strong>Antalis acuta (Hébert, 1849) (fide Bonci et al. 2000:206, 207, pl. 1, fig. 4).† apicostatum, Dentalium (Antalis) Vokes, 1939: 105,pl. 16, fig. 25. Eocene, Domengine Formation.UCMP locality 672, southern portion <strong>of</strong> the crest<strong>of</strong> Parson’s Peak, Coalinga Quadrangle, California,USA. Holotype UCMP 15747.† applanatum, Dentalium Torley, 1908: 43. Devonian.Schleddenh<strong>of</strong>es, near Iserlohn, Germany.applanatum, Dentalium (Bathoxiphus) Colman, 1958:145, fig. 12. <strong>Recent</strong>. 35 miles E <strong>of</strong> Sydney, 1463 m.Holotype AMS C22652; paratypes AMS C170645(24). Junior homonym <strong>of</strong> Dentalium applanatumTorley, 1908. Replacement name: Dentalium colmani(Palmer, 1974b: 124). Rhomboxiphus colmani(Palmer, 1974) (fide Chistikov 1983: 183).aprinum, Dentalium Linnaeus, 1767: 1263. [No referencesgiven]. <strong>Recent</strong>. Indian Ocean. Possible syntypeLS 611. Junior synonyms: Dentalium striatumBorn, 1778 partim; Dentalium striatulum Gmelin,1791; Dentalium lupinum Röding, 1798 partim;Dentalium caprinum Anton, 1838 (fide Pilsbry &Sharp 1897: 3); Dentalium interstriatum Sowerby,1860 (fide Habe 1964: 6, 7); Dentalium aprinumincolor Boissevain, 1906. Dentalium aprinumLinnaeus, 1767. Indo-Pacific: Madagascar to Philippines,New Caledonia <strong>and</strong> Australia. 5-123 m.† aquensis, Gadila Holzapfel, 1888: 179, pl. 20, fig. 8.Cretaceous. Near Vaals, Netherl<strong>and</strong>s. Cadulusaquensis (Holzapfel, 1888) (fide Pilsbry & Sharp1898: 255).† arabicum, Dentalium Cuvillier, 1933: 27, 28, pl. 3,fig. 8. Eocene, Bartonian. Gebel Giouch, east <strong>of</strong>Citadelle du Caire, Egypt.aratorum, Dentalium Cooke, 1885: 273, 274. <strong>Recent</strong>.Gulf <strong>of</strong> Suez, 10-30 fms [18-55 m]. SyntypesUMZC I.100,740. Junior synonym <strong>of</strong> Dentaliumclavus Cooke, 1885 (fide Boissevain 1906: 25, asDentalium mac<strong>and</strong>rewi). Remark: Singer (2003)removed D. clavus along with this <strong>species</strong> from thesynonymy <strong>of</strong> D. reevei Fischer, 1871. Dentaliumanatorum [Sharabati 1984: caption to pl. 1] is anincorrect subsequent spelling.† aratum, Dentalium Tate, 1887: 192, pl. 20, fig. 8.Miocene. River Murray Cliffs, near Morgan;Muddy Creek; Schnapper Point; Muloowurtie yellowclays near Ardrossan, Australia. Holotype <strong>and</strong>25 paratypes SAM T 256A. Paradentalium aratum(Tate, 1887) (fide Cotton & Ludbrook, 1938: 223,224); Dentalium (Dentalium) aratum Tate, 1887(fide Ludbrook 1959: 140).aratus, Cadulus Hedley, 1899: 551, fig. 60. <strong>Recent</strong>.Funafuti Atoll, Pacific Ocean, 60 m. Syntypes AMSC5635, AMS C5636, AMS C5638. Cadulus aratusHedley, 1899. Indo-Pacific: Madagascar to NewCaledonia. 45-460 m.† araucanum, Dentalium Philippi, 1887a: 107, pl. 12,fig. 17; 1887b: 101, 102, pl. 12, fig. 17. Tertiary.Lebu, Chile.† archiaci, Dentalium Tournouër in de Boullé, 1873:443 [17], 465 [39], pl. 6, fig. 10; 1876: 265, 275.Eocene, Auversien. Biarritz, France.† arciformis, Dentalium Conrad, 1846a: 212, pl. 1,fig. 3. Eocene. Claiborne, Alabama, USA. Juniorsynonym: Dentalium leai Meyer, 1885 (fide Pilsbry& Sharp 1898: 199).† arcotinum, Dentalium Forbes, 1846: 138, pl. 12,fig. 16. Cretaceous. Pondicherry, India. Remark:Tate (1889: 230) inadequately used this name for a<strong>species</strong> that was later described as Fustiaria wollumbillaensis(Etheridge in Jack & Etheridge, 1892)(fide Stilwell 1999: 220).† arcticus, Cadulus Dall, 1920: 30, pl. 5, fig. 8.Pliocene. USGS station 7070, Carter Creek, CamdenBay region, Arctic coast, Alaska, USA. LectotypeUSNM Paleobiology 324322 (designated byMacNeil 1957: 107, pl. 12, fig. 27, as “holotype”);paralectotypes USNM Paleobiology. Siphonodentalium“?” arcticus (Dall, 1920) (fide Emerson1957: 986). Remark: Emerson (1957: 986) alsocited USNM 324322 as the “holotype” <strong>and</strong> referedto “paratypes unnumbered”.† arctoides, Dentalium Gugenberger, 1934: 42, 43,pl. 1, fig. 14 [nomen nudum in Gugenberger 1933a:102; 1933b: 184]. Triassic. North <strong>of</strong> Launsdorf,Austria. Prodentalium arctoides (Gugenberger,1934) (fide Stiller 2001: 630).† arctum, Dentalium Pichler, 1857: 695. Triassic.Tirol, Austria. Prodentalium arctum (Pichler, 1857)(fide Stiller 2001: 630).† arcturus, Calstevenus Yancey, 1973: 1063, 1064,text-fig. 1a-1f. Lower Permian, Arcturus Group,Riepetown Formation. Buck Mountain, south <strong>of</strong> theRuby Range, White Pine County, eastern Nevada,USA. Holotype USNM Paleobiology 179130;paratypes USNM Paleobiology 179131-179135.arctus, Cadulus (Platyschides) Henderson, 1920: 124,pl. 19, fig. 19. <strong>Recent</strong>. Off Cape San Blas, Florida,USBF stn 2404, 28°44’N, 85°16’W, 60 fms [109m]. Holotype USNM 323955; paratypes USNM151830, AMNH 148348 (1). Gadila arctus (Henderson,1920). West Atlantic, Caribbean. 109 m.arcuata, Cadulus olivi var. Locard, 1898: 134, pl. 7,figs 8, 9. <strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Polyschides olivi (Scacchi, 1835).arcuata, Cadulus tumidosus var. Locard, 1898: 143.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Cadulus tumidosus Jeffreys, 1877.564ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)arcuata, Dentalium agile var. Locard, 1898: 118.<strong>Recent</strong>. Type locality not specified [North Atlantic].Type material not located. Synonym <strong>of</strong> Antalisagilis (G. O. Sars, 1872).arcuatum, Dentalium Gmelin, 1791: 3738 [citingGualtieri 1742: t. 10. f. G]. <strong>Recent</strong>. Type localitynot specified. Type material not located. Junior synonym<strong>of</strong> Dentalium elephantinum Linnaeus, 1758(fide Pilsbry & Sharp 1897: 2).† arenarium, Dentalium Römer, 1855: 13, pl. 3,fig. 16. Paleozoic? Spirifer-S<strong>and</strong>stone, near Schalke,Harz Mountains, Germany. Remark: Pilsbry &Sharp (1898: 230) suggested that this taxon “mayprove to be a Pteropod, Coleolus”.arenarium, Dentalium (Episiphon) Suter, 1907: 214,215, pl. 18, fig. 11. <strong>Recent</strong>. Port Pegasus, StewartIsl<strong>and</strong>, New Zeal<strong>and</strong>, 18 fms [33 m]. HolotypeNZGS TM1219 (fide Boreham 1959: 75) ; paratypesSAM D 16001(fide Zeidler & Macphail 1978: 83),ZMB 61911 (fide Kilias 1995: 172). Junior homonym<strong>of</strong> Dentalium arenarium Römer, 1855. Replacementname: Dentalium suteri Emerson, 1954: 185.Antalis suteri (Emerson, 1954) (fide Dell 1957: 573).arguticosta, Dentalium Brugnone, 1878: 25. Tertiary.Monte Pellegrino <strong>and</strong> Ficarazzi, Italy. Junior synonym<strong>of</strong> Antalis panorma (Chenu, 1843) (fidePilsbry & Sharp 1897: 54; Bellini 1909: 226).arietinum, Dentalium Müller, 1776. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† armillatum, Dentalium Toula, 1911: 496, pl. 31,fig. 8. Miocene. Gatun, Panama. Type material notlocated. Synonym: Dentalium armillatum proterumWoodring, 1973. Remark: Woodring (1973: 482)gave “Technische Hochschule, Vienna” as typedepository; this could not be verified.† armoricense, Siphonodentalium Cossmann, 1902:109 [159], pl. 10 [15], figs 26, 27. Eocene. Le Bois-Gouët <strong>and</strong> Arthon, France.arnaudi, Polyschides Scarabino, 1995: 338, 339,figs 141, 147b, h. <strong>Recent</strong>. West Indian Ocean, MD32 Réunion, stn DC 124, 20°52’S, 55°37’E, 40 m.Holotype MNHN; paratypes MNHN (13),USNM 890855 (1); NM (1). Polyschides arnaudiScarabino, 1995. Indian Ocean: Madagascar, LaRéunion. 40-980 m.arne, Cadulus (Platyschides) rushii Henderson, 1920:118, pl. 18, fig. 9. <strong>Recent</strong>. Off New Jersey [USA],USBF stn 2234, 30°09’N, 72°03’05”W, 810 fms[1478 m]. Holotype <strong>and</strong> 5 paratypes USNM314729. Synonym <strong>of</strong> Polyschides rushii (Pilsbry &Sharp, 1898). Western Atlantic: New Jersey,Massachusetts. 360-1796 m.† arnoensis, Polyschides Maxwell, 1992: 188, pl. 9,figs e, f. Eocene. NZGS locality GS9480 <strong>and</strong>GS9508, Waihao River, near McCulloch’s Bridge,South Canterbury, New Zeal<strong>and</strong>. Holotype NZGSTM7232; paratypes NZGS (2).arnoldi, Laevidentalium Lamprell & Healy, 1998:111, 112, figs 112B, 113D, 116. <strong>Recent</strong>. OffCairns, northern Queensl<strong>and</strong>, Australia, 17°22’S,146°48’E, 303 m. Holotype QM MO53676;paratypes QM MO17768 (189; northernQueensl<strong>and</strong>, 17°21.77’S, 146°48.5’E, 303 m), QMMO17971 (27; 17°21.06’S, 147°02.5’E, northernQueensl<strong>and</strong>, 303 m), QM MO17996 (15;17°22’S, 146°48’E, 303 m), QM MO53677 (202;17°22’S, 146°48’E, 303 m), QM MO17892 (45;17°22’S, 146°48’E, 303 m), QM MO17996 (56;northern Queensl<strong>and</strong>, 17°21.77’S, 146°48.5’E,303 m), AMS C201737 (2 lots: 229; east <strong>of</strong>Innisfail, Queensl<strong>and</strong>, 17°21.77’S, 146°48.52’E,296-303 m; many; 17°22’S, 146°48’E, 303 m),AMS C310225 (2; east <strong>of</strong> North West Isl<strong>and</strong>,Queensl<strong>and</strong>, 23°15.2’S, 152°24.1’E, 284 m).Laevidentalium arnoldi Lamprell & Healy, 1998.Indo-Pacific: NE Australia. 155-1224 m. Remark:V. Scarabino (in litt. 5.XI.2001) suggested that thestatus <strong>of</strong> L. arnoldi Lamprell & Healy, 1998 vis-àvisL. houbricki Scarabino, 1995 requires furtherresearch.† arrosus, Cadulus (Platyschides) Woodring, 1925:210, pl. 28, fig. 12. Miocene. Bowden, Jamaica.Holotype USNM Paleobiology 353078.artatus, Cadulus Locard [ex Jeffreys MS], 1897a: 4(nomen nudum in Jeffreys 1880: 317). <strong>Recent</strong>. Bay<strong>of</strong> Biscay, 1019-2651 m. Type material not located.Cadulus artatus Locard, 1897. Eastern Atlantic:Bay <strong>of</strong> Biscay to Angola. 1019-2651 m.asgum, Dentalium de Gregorio, 1890: 171, pl. 17,figs 22-24. Eocene. Claiborne, Alabama. Typematerial lost (fide Palmer & Brann 1965: 369).Junior synonym <strong>of</strong> Dentalium thalloides Conrad,1833 (fide Pilsbry & Sharp 1898: 218).† ashiyaensis, Dentalium Nagao, 1928: 88, 89, pl. 14,figs 38-44. Paleocene. Sakamizu beds (Sakamizu)<strong>and</strong> Wakita beds (Wakita), Shimago-mura, Ongagun,Chikuzen Province, Kyushu, Japan. Holotype<strong>and</strong> paratype Inst. Geol. Paleont., Tohoku Univ.(fide Hanzawa et al. 1961: 209).asperum, Dentalium Michelotti, 1847: 144, pl. 5,figs 20, 21. Miocene. Tortone, Italy. Junior synonym<strong>of</strong> Coccodentalium radula (Gmelin, 1791)(fide Sacco 1897: 111, pl. 10, figs 7-15). Dentaliumaspersum [Pilsbry & Sharp, 1898] is an incorrectsubsequent spelling.asperum, Dentalium Brunet, 1995: 53, pl. 5, fig. 7.Upper Miocene, Entrerriense Formation. BalizaPunta Flecha, Puerto Madryn, Chubut Province,Argentina. Holotype MPEF-PI-080. Junior homonym<strong>of</strong> Dentalium asperum Michelotti, 1847.Replacement name: Dentalium bruneti n. nom.aspersum, see asperum Michelotti, 1847.† asphaltodes, Dentalium (Dentalium) Beets, 1943:307, pl. 29, figs 110-112. Upper Oligocene.Waisiu, Buton Isl<strong>and</strong>, Indonesia.† astensis, Dentalium (Antale “?”) dentale var. Sacco,1897: 104, pl. 8, figs 72, 73 [as a nomen nudumin Sacco 1896: 97]. Pliocene. Astigiana, Italy.ZOOSYSTEMA • 2004 • 26 (4)565


Steiner G. & Kabat A. R.Syntypes MIGT BS.106.02.063, BS.106.02.064(fide Ferrero Mortara et al. 1984: 306).† astensis, Dentalium (Fustiaria) polita var. Sacco,1897: 112, pl. 10, fig. 24. Pliocene. Astigiano, Italy.Syntype MIGT BS.106.06.001 (fide FerreroMortara et al. 1984: 309).† atava, Dentalium (Antale “?”) taurocostatum var.Sacco, 1897: 102, pl. 8, fig. 56 [as a nomen nudumin Sacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.02.048 (fide FerreroMortara et al. 1984: 304).† atavus, Cadulus (Gadila) Finlay & Marwick, 1937:111, pl. 15, fig. 20. Eocene. Castle Hill Shaft,Matau, New Zeal<strong>and</strong>. Holotype NZGS TM4815(fide Keyes 1972: 97). Cadulus (s.l.) atavus Finlay &Marwick, 1937 (fide Beu & Maxwell 1990: 424).atlanticus, Cadulus (Gadila) Henderson, 1920: 137-139, pl. 20, fig. 2. <strong>Recent</strong>. Off Nantucket Isl<strong>and</strong>[Massachusetts, USA], USBF stn 2682, 39°38’N,70°22’W, 1004 fms [1830 m]. Holotype USNM78238; paratypes USNM 78238 (3), MCZ 186815(1), AMNH 148338 (2; USBF stn 2115, 843 fms[1538 m]). Cadulus atlanticus Henderson, 1920.Western Atlantic: Massachusetts to Delaware, USA.780-1934 m.atlantideus, Cadulus (Dischides) Nicklès, 1955: 100,101, fig. 5. <strong>Recent</strong>. Guinea, Atlantide stn 44,10°22’N, 16°22’W, 41-55 m. Syntypes ZMUC.Dischides atlantideus (Nicklès, 1955). EasternAtlantic: Guinea, Mauritania. 9-55 m.atramentum, Dentalium Schlüter, 1838: 39. “Fossil”.Nomen nudum.attenuata, Cadulus olivi var. Locard, 1898: 134.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Polyschides olivi (Scacchi, 1835).attenuata, Cadulus cyathus var. Monterosato, 1875: 21(1872: 27, as Cadulus ovulum var. attenuata, nomennudum). <strong>Recent</strong>. Ficarazzi (Palermo) [Pleistocene <strong>of</strong>Italy] <strong>and</strong> Mediterranean. Cadulus attenuatusMonterosato, 1875 (fide Di Geronimo & La Perna1997: 417, in remarks to Cadulus ovulum).Mediterranean. No bathymetric data. Remarks:introduced as a nomen nudum, the name is laterreferred to the Pliocene <strong>species</strong> Cadulus cyathus (DeCrist<strong>of</strong>ori & Jan, 1832), by Monterosato (1875:21), Pilsbry & Sharp (1898: 158) <strong>and</strong> Caprotti(1968: 78). Here, the decision <strong>of</strong> Di Geronimo &La Perna (1997: 417) is favoured, keeping C. attenuatusseparate from C. cyathus <strong>and</strong> C. ovulum.attenuata, Dentalium vulgare var. Monterosato, 1884:32. <strong>Recent</strong>. Mediterranean. Nomen nudum.† attenuatum, Dentalium Say, 1824: 154, 155, pl. 8,fig. 3. Miocene. Maryl<strong>and</strong>, USA. Type material inBMNH Palaeontology (fide Newton 1902: 304).Junior synonym: Dentalium duodecenaria Conrad,1862 (fide Pilsbry & Sharp 1898: 199).attenuatum, Dentalium Sowerby, 1860: 99, pl. 225,fig. 40. <strong>Recent</strong>. Type locality not specified. Typematerial not located. Junior homonym <strong>of</strong> Dentaliumattenuatum Say, 1824. Episiphon attenuatum(Sowerby, 1860) (fide Pilsbry & Sharp 1897: 120).Remark: Pilsbry & Sharp (1897: 120) stated thattheir Dentalium (Episiphon) longum may be a juniorsynonym <strong>of</strong> this <strong>species</strong>. Since the locality is unknownfor both <strong>species</strong>, these <strong>names</strong> are <strong>of</strong> dubious value,<strong>and</strong> we prefer not to rename the junior homonym.† aturensis, Pseudantalis Cossmann & Peyrot, 1917:179, 180, pl. 1, figs 67-72. Miocene, Burdigalian<strong>and</strong> Helvetian. Dax (Maïnot), Saint-Paul-lès-Dax,<strong>and</strong> Salies-de-Béarn, Clermont (L<strong>and</strong>es), France.austinclarki, Cadulus (Platyschides) Emerson, 1951:24-26. <strong>Recent</strong>. Santa Inez Bay, Baja California[Mexico], W around Santa Inez Point, 6-12 feet [2-4 m]. Holotype USNM 602347; paratypes USNM602347, SBMNH 35021. Gadila austinclarki(Emerson, 1951). Eastern Pacific: Mexico toPanama, Galapagos. 2-73 m.austini, Dentalium (Dentalium) Lamprell & Healy,1998: 31-33, figs 20E, 21E, 27. <strong>Recent</strong>. CullenBeach, Darwin, Northern Territory, Australia.Holotype MV F75838; paratypes MV F67466(18), MV F67474 (13; Cameron Beach, Shoal Bay,Darwin), MV F67469 (16), AMS C172543 (13;Dudley Point, on mud flats [Northern Territory,Australia]), AMS C172539 (5; east end CullenBeach, Darwin [Australia]), AMS C172540 (14;Port Darwin [Australia]), AMS C380639 (2;Mindel Beach, Darwin [Australia], low tide), WAM106-93 (1; <strong>of</strong>f Eagle Bay [Australia], 32 m).Dentalium austini Lamprell & Healy, 1998. Indo-Pacific: northern Australia. 0-32 m.australasiae, Siphonodentalium Boissevain, 1906: 64,pl. 6, fig. 68. <strong>Recent</strong>. B<strong>and</strong>a Sea, Siboga stn 211,5°40.7’S, 120°45.5’E, 1158 m. Holotype ZMA3.06.087. Siphonodentalium australasiae Boissevain,1906. Known from type locality only.† australis, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 199. Miocene. Muddy Creek,Victoria, Australia. Holotype <strong>and</strong> 3 paratypes SAMT 250A. Replacement name for Entalis annulatumTate, 1887, a junior secondary homonym <strong>of</strong> Dentaliumannulatum Gmelin, 1791. Laevidentalium australe(Sharp & Pilsbry in Pilsbry & Sharp, 1898)(fide Ludbrook 1959: 145, pl. 2, fig. 1).badenense, see badense.† badense, Dentalium Hörnes, 1856: 652, 653, pl. 50,fig. 30 [ex Partsch MS]. Miocene. Baden, Austria.Lectotype NMW 1846.37.534 (designated by Pavia1991: 147); paralectotypes NMW (10). Fissidentaliumbadense (Hörnes, 1856) (fide Pavia 1991:146-148, pl. 5, fig. 4, pl. 6, fig. 6). Remarks:Trautschold (1859: 313, pl. 6, figs 4a-4c) erroneouslyidentified a <strong>species</strong> from the Oligocenearound Aral Sea as Dentalium badense which waslater named Dentalium trautscholdi Koenen, 1868.Dentalium (Entalis) badenense “Ptsch.” [Boettger,1902: 181] is an incorrect subsequent spelling.566ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)† bakoniense, Dentalium Palmer, 1974b: 124.Miocene, Tortonian. Bakony, Hungary. HolotypeHungarian Geological Collection, M.279.Replacement name for Dentalium annulatumKokay, 1966, a junior homonym <strong>of</strong> Dentaliumannulatum Gmelin, 1791, D. annulatum Crist<strong>of</strong>ori& Jan, 1832, D. annulatum S<strong>and</strong>berger &S<strong>and</strong>berger, 1856, <strong>and</strong> D. annulatum Meyer, 1886.balanoides, Dentalium (Laevidentalium) Plate, 1908a:357, pl. 30, figs 42-44. <strong>Recent</strong>. Valdivia stn 186,03°22.1’S, 101°11.5’E, 903 m, West Sumatra.Holotype ZMB 61109. Calliodentalium balanoides(Plate, 1908) (fide Scarabino 1995: 275). Indo-Pacific: Indonesia, Philippines, New Caledonia.187-903 m.bambusa, Anulidentalium Chistikov, 1975: 19.<strong>Recent</strong>. Tonkin Bay, Vietnam, Pelamida stn 38, 72m. Lectotype ZIN 1 (designated by Chistikov1979: 113); paralectotype ZIN (Pelamida stn 2,87 m). Anulidentalium bambusa Chistikov, 1975.Indo-Pacific: Vietnam, Philippines, Indonesia. 38-270 m.banale, Dentalium (Laevidentalium) Boissevain, 1906:55, pl. 6, fig. 30. <strong>Recent</strong>. Indonesia, Timor Sea,Siboga stn 300, 10°48.6’S, 123°23.1’E, 918 m.Lectotype ZMA 3.06.068 (designated by Lamprell& Healy 1998: 114); paralectotype ZMA 3.06.069(1). Laevidentalium banale (Boissevain, 1906) (fideHabe & Kosuge 1964: 7). Indo-Pacific: Indonesiato West Australia. 0-918 m. Remarks: Scarabino(1995: 292) considered this <strong>species</strong> to be a juniorsynonym <strong>of</strong> Laevidentalium leptosceles (Watson,1879), an opinion rejected by Lamprell & Healy(1998: 184). Ludbrook (1954: 105) mentioned a“holotype” but did not indicate which <strong>of</strong> the 2 specimensshe referred to; thus, the lectotype selectionby Lamprell & Healy (1998) is considered valid.b<strong>and</strong>ata, Dentalia Perry, 1811: caption to pl. 52. Typelocality not specified. Nomen dubium. No informationon type material.† b<strong>and</strong>eli, Prodentalium Engeser & Riedel, 1992: 43,44, pl. 1, figs 1-3, text-fig. 1. Jurassic, MiddleLiassic, Pliensbachian. Hoisdorf, Schleswig-Holstein, Germany. Holotype GPIMH 2516/1;paratype GPIMH 2516/2.† bangtoupoensis, Laevidentalium Stiller, 2001: 620-623, figs 3, 5.1-5.4. Upper Anisian, Middle Triassic.Bangtoupo, NNE <strong>of</strong> Qingyan, Guizhou Province,southwestern China. Holotype GPIM B6D-1.S1.Fbt-1; paratypes GPIM B6D-1.S1.Fbt-2,GPIM B6D-1.S1.Fbt-3, GPIM B6D-1.S1.F30:32-1, GPIM B6D-1.S1.Fbt-4.barbadensis, Cadulus (Cadulus) transitorius Henderson,1920: 144, pl. 19, fig. 7. <strong>Recent</strong>. S<strong>and</strong>y Bay,Barbados, 100 fms [183 m]. Holotype USNM95375. Junior synonym <strong>of</strong> Cadulus transitoriusHenderson, 1920. Remarks: MCZ 7744 (Blake, <strong>of</strong>fBarbados, 100 fms) was labeled by Henderson asbeing a “cotype” <strong>of</strong> barbadensis, but this specimenwas not mentioned in the original description <strong>and</strong> isneither a syntype nor a paratype. Turner (1955:318) cited this specimen as a “paratype” even thoughshe recognized that it was not cited by Henderson.† barquense, Dentalium “?” Winchell, 1862: 425.Carboniferous, Huron Group. Pointe aux Barques,Michigan, USA. Remarks: Winchell (1865: 131),based upon examination <strong>of</strong> additional specimens,questioned whether this <strong>species</strong> might actually represent“corals with very large hollow axes”.† barremicum, Dentalium Cossmann, 1916: 31, pl. 2,figs 4, 5. Cretaceous, Barremian. Gisement desAugustines, near Brouzet-les-Alais, Gard, France.barremicus, Dentalium Sasonova, 1958: 131, 132,pl. 13, figs 1-3, pl. 14, fig. 2, pl. 15, fig. 2.Cretaceous, Barremian. River Kubre, near VolgaRiver, Russia. Junior homonym <strong>of</strong> Dentalium barremicumCossmann, 1916. Replacement name:Dentalium kubraense Palmer, 1974.bartletti, Dentalium (Antalis) Henderson, 1920: 55,56, pl. 8, figs 2, 7. <strong>Recent</strong>. Off Frederickstadt,St Cruz, Blake stn 130, 451 fms [825 m]. HolotypeUSNM 224986; paratypes USNM 93121 (1; <strong>of</strong>fCape Fear, North Carolina, USBF stn 2616, 17 fms[31 m]), USNM 93840 (9; between MississippiDelta <strong>and</strong> Cedar Keys, USBF stn 2398, 227 fms[141 m]), USNM 94081 (1; Off Arrowsmith Bank,Yucatan, USBF stn 2355, 399 fms [728 m]),USNM 95368 (3; NW <strong>of</strong> Tortugas, Blake stn 44,539 fms [984 m]), USNM 95369 (1; Martinique,Blake stn 211, 357 fms [652 m]), USNM 95370(1; <strong>of</strong>f Morro Light, Habana, Blake stn 100,400 fms [730 m]), USNM 95371 (11; St Vincent,West Indies, Blake stn 230, 464 fms [847 m]),USNM 314449, 314450 (2; <strong>of</strong>f Fowey Light,Florida, Eolis stn 353, 85 fms [155 m]), USNM323776 (1; <strong>of</strong>f Cape San Blas, Florida, USBFstn 2399, 196 fms [358 m]), USNM 330524 (1; <strong>of</strong>fCape Florida, Florida, USBF stn 2644, 193 fms[352 m]), AMNH 148300, 148301 (2; <strong>of</strong>fSt Vincent Isl<strong>and</strong>, Blake stn 230, 13°13’20”N,61°18’45”W, 464 fms [847 m]). Antalis bartletti(Henderson, 1920). Western Atlantic <strong>and</strong>Caribbean. 30-1000 m.bartonense, Dentalium Palmer, 1974b: 125. Eocene.London, Engl<strong>and</strong>, Great Britain. Replacementname for Dentalium striatum J. Sowerby, 1814, ajunior homonym <strong>of</strong> Dentalium striatum Born,1778. Junior objective synonym <strong>of</strong> Dentalium acuticostatumDeshayes, 1825 (n. syn.). Remarks:Palmer (1974b: 125) proposed Dentalium bartonenseas a replacement name, apparently beingunaware that Deshayes had already renamed this<strong>species</strong>. Used as a valid <strong>species</strong> <strong>of</strong> Antalis by LeRenard (1995: 175) <strong>and</strong> Le Renard & Pacaud(1995: 86), who were unaware that it was an objectivejunior synonym.† basteroti, Entalis Cossmann & Peyrot, 1917: 178,179, pl. 1, figs 42-45. Miocene, Burdigalian <strong>and</strong>ZOOSYSTEMA • 2004 • 26 (4)567


Steiner G. & Kabat A. R.Aquitanian. Mérignac (le Pontic), Dax (Maïnot),Saint-Médard (La Fontaine), Cestas, Saint-Paul-lès-Dax, Léognan (Thibaudeau), Canéjean, Martillac(Pas-de-Barreau), <strong>and</strong> Pessac (Lorient), France.† batheri, Dentalium Finlay, 1927: 521. Carboniferous.New Zeal<strong>and</strong>. Replacement name forDentalium huttoni Bather, 1905, a junior homonym<strong>of</strong> Dentalium huttoni Kirk, 1880.beachportensis, Tesseracme Cotton & Ludbrook, 1938. Nota scaphopod. See Other taxa described as <strong>Scaphopoda</strong>.bednalli, Dentalium Pilsbry & Sharp, 1898: 248.<strong>Recent</strong>. St Vincent’s Gulf, South Australia. HolotypeANSP 72554. Junior synonym <strong>of</strong> Paradentaliumintercalatum (Gould, 1859) (fide Verco1904: 135).beecheyi, Pulsellum Lamprell & Healy, 1998: 146,147, figs 150C, 151C, D, 153. <strong>Recent</strong>. 200 kmNorth <strong>of</strong> Croker Isl<strong>and</strong>, Northern Territory,Australia, 9°14’S, 132°54’E, 145 m. Holotype AMSC174890; paratypes AMS C172289 (6), AMSC172288 (2; 17°34’S, 120°22’E, c. 212 km NW <strong>of</strong>Roebuck Bay, Western Australia, 188 m), AMSC172293 (7; 9°18’S, 133°38’E, 230 km N <strong>of</strong>Goulburn Isl<strong>and</strong>, Northern Territory, 135 m),AMS C172296 (4; 9°24’S, 133°02’E, 180 km N <strong>of</strong>Croker Isl<strong>and</strong>, Northern Territory, 125 m).Pulsellum beecheyi Lamprell & Healy, 1998. Indo-Pacific: northern Australia. 18-256 m. Remark: thelatitude <strong>of</strong> the type locality in the original description,“29°14’S,” is apparently an error for 9°14’S.belcheri, Cadulus (Dischides) Pilsbry & Sharp, 1898:145, pl. 26, figs 84-87. <strong>Recent</strong>. North Pacific. HolotypeUSNM 107703. Dischides belcheri Pilsbry &Sharp, 1898. North Pacific: Japan. 0-50 m.belcheri, Dentalium Sowerby, 1860: 101, pl. 224,figs 28, 29. <strong>Recent</strong>. East Indian Archipelago[Indonesia]. Lectotype BMNH 1996077/1 (designatedby Scarabino 1995: 201, “[...] the largest(25 mm) <strong>of</strong> the 3 syntypes BMNH”); paralectotypesBMNH 1996077/2 + 3. Junior synonym <strong>of</strong>Dentalium variabile Deshayes, 1825 (fide Habe1963: 258). Remark: Tomlin (1931: 337) placedDentalium belcheri in the synonymy <strong>of</strong> Dentaliumstrigatum Gould, 1859.belcheri, Prodentalium Nassichuk & Hodgkinson,1976. Not a scaphopod. See Other taxa described as<strong>Scaphopoda</strong>.† bellulus, Cadulus Clark, 1895: 5; 1896: 72, pl. 14,fig. 6. Eocene. Woodstock, Hanoverville, Virginia,USA.† benoisti, Gadila Cossmann & Peyrot, 1917: 182,183, pl. 2, figs 1-3. Miocene. Giraudeau, France.Replacement name for Gadus ventricosus Benoist,1873, a junior homonym <strong>of</strong> Cadulus ventricosus(Bronn, 1828). Junior synonym: Gadila benoistihoutalensis (Goddeeris, 1977) (fide Janssen 1984:117, 118, pl. 42, figs 18a, 18b).berryi, Dentalium Smith & Gordon, 1948: 216, 217,pl. 3, figs 1, 4. <strong>Recent</strong>. Monterey Bay; MontereyCo., Humpback Reef [California], 40 fms [73 m].Holotype CAS 065592; paratypes SBMNH 35020,CAS 065593 (1), CAS 065594 (2), CAS 065595(1; Monterey County, <strong>of</strong>f Cabrillo Point, near bellbuoy, 20 fms [37 m]), ANSP 185227 (1; near bellbuoy, Pacific Grove, Monterey Bay, California,36°36’N, 121°56’W). Antalis berryi (Smith &Gordon, 1948) (fide Emerson in Turgeon 1988:50). Eastern Pacific, California. 37-73 m. Remark:Keen (1971: 884) listed Dentalium (Antalis) pretiosumberryi as a southern sub<strong>species</strong> <strong>of</strong> Antalis pretiosa.This indicates a possible synonymy <strong>of</strong>A. berryi (Smith & Gordon, 1948) with D. indianorumCarpenter, 1864 <strong>and</strong> A. denseliratum Pilsbry &Sharp, 1898, both <strong>names</strong> for the southern variety orsub<strong>species</strong> <strong>of</strong> A. pretiosa . Further research isrequired to decide whether or not A. denseliratum isa senior synonym <strong>of</strong> A. berryi.bicarinatum, Dentalium Deshayes, 1825: 364, pl. 18,figs 16, 17. Eocene. Near Paris, France. Junior synonym<strong>of</strong> Dentalium duplex Defrance, 1819 (fideDeshayes 1864: 203).bicarinatum, Dentalium Wanner, 1902: 124, pl. 18,figs 9, 10. Cretaceous. North <strong>of</strong> Oasis Dachel,Libya. Junior homonym <strong>of</strong> Dentalium bicarinatumDeshayes, 1825. Replacement name: Dentaliumwanneri n. nom. Remark: Wanner’s description <strong>of</strong> awell preserved specimen is detailed <strong>and</strong> referred toby Quaas (1902: 234). We, therefore, decided torename this junior homonym.bicostale, Dentalium Ryckholt, 1851: 71, pl. 2, figs 43,44. Cretaceous, Neocomian. Tournay, Belgium.Tentative junior synonym <strong>of</strong> Dentalium decussatumJ. Sowerby, 1814 (fide Ryckholt 1851).bifissum, Dentalium S. Wood, 1848: 190, pl. 20,fig. 3. Coralline Crag (Pliocene), Sutton, Engl<strong>and</strong>.Type material not located. Junior synonym <strong>of</strong>Dischides politus (S. Wood, 1842) (fide Pilsbry &Sharp 1897: 144).† bifissuratum, Dentalium Deshayes, 1850: 36, pl. 61,figs 11, 12, 14. Fossil. Locality not specified.Remark: Pilsbry & Sharp (1898: 235) erroneouslydated this name to “1864” <strong>and</strong> they therefore listedit as a junior synonym <strong>of</strong> Dischides bilabiatus(Deshayes, 1861).† bifrons, Dentalium “(?)” Tate, 1887: 192, 193,pl. 20, fig. 5. Tertiary (Miocene-Pliocene). Upperbeds at Muddy Creek, Hamilton, Victoria,Australia. Holotype <strong>and</strong> 3 paratypes SAM T 255.Dentalium (Fissidentalium) bifrons Tate, 1887 (fideTate 1899: 261); Dentalium (Antalis) bifrons Tate,1887 (fide Ludbrook 1959: 142, 143, pl. 2, fig. 2).† bilabiatus, Gadus Deshayes, 1861: 219, pl. 3,figs 22-24. Eocene. Paris Basin (Houdan, Grignon,Hérouval, Montjavoult, Rozières <strong>and</strong> Saint-Sulpice), France. Dischides bilabiatus (Deshayes,1861) (fide Pilsbry & Sharp 1898: 235; Le Renard1995: 175; Le Renard & Pacaud 1995: 86).Remark: Pilsbry & Sharp (1898: 235) erroneously568ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)listed Dentalium bifissuratum Deshayes, “1864” as ajunior synonym <strong>of</strong> this <strong>species</strong>; in fact, D. bifissuratumwas described in 1850, not 1864.billingsi, Salterella Safford, 1869. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.bimixtum, Dentalium de Gregorio, 1890: 172, pl. 17,figs 32-34. Eocene. Claiborne, Alabama. Juniorsynonym <strong>of</strong> Dentalium thalloides Conrad, 1833(fide Pilsbry & Sharp 1898: 218).† binkhorsti, Dentalium Pilsbry & Sharp, 1898: 220.Cretaceous. Belgium. Replacement name forDentalium nysti Van den Binkhorst, 1861, a juniorhomonym <strong>of</strong> Dentalium nysti d’Orbigny, 1852.† bironense, Dentalium Douvillé, 1929: 374, 375,pl. 32, fig. 1. Eocene. Biron-Marnière, France.bisexangulatum, Dentalium Sowerby, 1860: 102, pl.223, fig. 8. <strong>Recent</strong>. Java. Lectotype BMNH1993009/1 (designated by Lamprell & Healy 1998:65); paralectotypes BMNH 1993009/2-3 (2).Dentalium bisexangulatum Sowerby, 1860. Indo-Pacific: Red Sea, East Africa to Australia, Java,Philippines, Japan. 1-720 m.bisinuatum, Dentalium André, 1896: 397, pl. 17,figs 9a-c. <strong>Recent</strong>. Amboyna [Ambon, Indonesia].Holotype MHNG 1155/40 (ex M3/35). Laevidentaliumbisinuatum (André, 1896) (fide Pilsbry& Sharp 1897: 108). Known from the type localityonly. Remark: V. Scarabino (in litt. 5.XI.2001) suggestedthat the status <strong>of</strong> D. bisinuatum vis-à-visLaevidentalium eburneum (Linnaeus, 1767) requiresfurther research.bisiphonata, Fustiaria Newton, 1891: 284 [ex EdwardsMS]. Lower Eocene. London Clay, HaverstockHill, Engl<strong>and</strong>, Great Britain. Nomen nudum.Remark: Pilsbry & Sharp (1898: 200) stated that“as the <strong>species</strong> is not mentioned by Newton &Harris [1894] in their later revision <strong>of</strong> BritishEocene Scaphopods, it is probably either not validor based on material unsuitable for characterization”.† bisissura, Cadulus “(?)” Garvie, 1996: 46, 47, pl. 9,figs 3-5. Eocene, Lower Claibornian, ReklawFormation, Marquez Member. Locality 4, Bluff onRidge Creek, Texas, USA. Holotype PRI 30314;paratype PRI 30315.† bitubatum, Dentalium Meyer, 1886: 64, pl. 3, fig. 1.Eocene. Jackson, Mississippi, USA. LectotypeUSNM Paleobiology 638811 (as “holotype” inPalmer & Brann 1965: 366).† bl<strong>and</strong>um, Dentalium de Gregorio, 1890: 172,pl. 17, figs 26-31. Eocene. Claiborne, Alabama.Type material lost (fide Palmer & Brann 1965:366).† blountense, Cadulus Mansfield, 1935: 46, 47, pl. 5,fig. 10. Upper Middle Miocene. Locality 12046,Vaughan Creek, Walton County, Florida, USA.Holotype USNM Paleobiology 373159.† bocasensis, Dentalium (Dentalium) Olsson, 1922:338 [166], pl. 15, figs 2, 3. Miocene, Gatun Stage.Bocas del Toro, Costa Rica. Syntypes PRI 21116,PRI 21117 (fide Brann & Kent 1960: 317).† boettgeri, Dentalium Noetling, 1901: 251, pl. 17,fig. 4. Miocene. Kama, Burma.† boggsi, Dentalium Berry, 1926: 19, 20, figs 1, 2.Eocene, Saman shale. Negritos, Peru.boissevainae, Antalis Palmer, 1974b: 124. <strong>Recent</strong>.Indonesia, Siboga stn 159, 00°59.1’S, 129°48.8’E,411 m. Holotype (<strong>of</strong> indicum Boissevain) ZMA3.06.051. Replacement name for Dentalium entalisvar. indicum Boissevain, 1906, a junior homonym<strong>of</strong> Dentalium indicum Chenu, 1843. Junior synonym<strong>of</strong> Antalis tibana (Nomura, 1940) (fide Habe1961: 105, but not Habe 1963: 262). Remark:Scarabino (1995: 231) unnecessarily designated thesingle specimen as lectotype.boissevaini, Cadulus (Gadila) Jaeckel, 1932: 311, textfig.8. <strong>Recent</strong>. Off Zanzibar, Africa, Valdivia stn251, 1°40.6’S, 41°47.1’E, 693 m. Lectotype ZMB75359a (designated by Kilias 1995: 172); paralectotypesZMB 75359b, ZMB 75360. Junior synonym:Cadulus (Gadila) reesi Ludbrook, 1954 (fideScarabino 1995: 358). Gadila boissevainae (Jaeckel,1932) (fide Scarabino 1995: 358). Indian Ocean,Africa. 274-1280 m. Remark: since MariaBoissevain was a woman, the gender <strong>of</strong> the genetive<strong>of</strong> the <strong>species</strong> name was emended by Scarabino(1995: 358).bonei, see bouei.booceras, Cadulus Tomlin, 1926: 298, pl. 16, fig. 11.<strong>Recent</strong>. Durban Bay, South Africa; Cangella (fideBarnard 1963b: 352). Syntypes BMNH1926.12.6.3, NMW. Siphonodentalium booceras(Tomlin, 1926) (fide Scarabino 1995: 333). IndianOcean: South Africa. 89-183 m.† boonei, Dentalium Cossmann, 1924: 22, 23, pl. 2,figs 13-16, 25, 26, pl. 7, fig. 24. Jurassic, Callovian.Doux, France.borcei, see bouei.bordaensis, Cadulus (Gadila) Cotton & Godfrey,1940: 340, fig. 362. <strong>Recent</strong>. Off Cape Borda,Kangaroo Isl<strong>and</strong>, South Australia, 55 fms [100 m].Holotype SAM D 13761. Gadila bordaensis(Cotton & Godfrey, 1940) (fide Lamprell & Healy1998: 153). Indian Ocean: South Australia. 50-600 m.bordaensis, Dentalium (Episiphon) Cotton &Ludbrook, 1938: 220, 221, pl. 12, fig. 3. <strong>Recent</strong>.Cape Borda, Kangaroo Isl<strong>and</strong>, South Australia, 60fms [110 m]. Holotype SAM D 13340. Episiphonbordaensis (Cotton & Ludbrook, 1938) (fideLamprell & Healy 1998: 132). Indian Ocean:Australia. 75-400 m.† boreale, Dentalium Böhm, 1904: 53, 54, pl. 6,figs 17, 33, 36. Upper Triassic. Mount Misery, BearIsl<strong>and</strong>, Spitsbergen, Arctic Sea.† borealis, Dentalium (Entalis) badense var. Kautsky,1925: 53, 54, pl. 5, fig. 12. Miocene. Hemmoor<strong>and</strong> Basbeck, Lower Saxony, Germany.ZOOSYSTEMA • 2004 • 26 (4)569


Steiner G. & Kabat A. R.† borelliana, Gadilina Pavia, 1991: 131, 132, pl. 7,figs 4, 5, 9, 10, pl. 8, figs 5, 6. Upper Miocene.Borelli, near Turin, Italy. Holotype <strong>and</strong> 192paratypes MIGT BS.D.12.1.† bothrum, Dentalium (Dentalium) Woodring, 1973:482, 483, pl. 75, figs 14, 23. Middle Miocene,upper part <strong>of</strong> Gatun Formation. USGS locality8413, upper edge <strong>of</strong> Chilas village, Río Indios,Colón Province, Panama. Holotype USNMPaleobiology 646730.boucheti, Antalis Scarabino, 1995: 234, 235, figs 42,45h, 45j, 73g. <strong>Recent</strong>. Loyalty Isl<strong>and</strong>s, RV Alis,MUSORSTOM 6 stn DW 428, 20°24’S,166°13’E, 420 m. Holotype MNHN; paratypesMNHN (12; MUSORSTOM 6: 1 from stnCC175; 1 from stn DW 397; 1 from stn DW 406,2 from stn DW 411; 3 from stn DW 428; 2 fromDW 446; 1 from stn DW 451; 1 from stn 481),AMS C201723 (1; stn DW 428), NMNZM268969 (1; stn DW 428), USNM 890859 (1; stnDW 397). Antalis boucheti Scarabino, 1995. Indo-Pacific: Philippines to New Caledonia. 360-842 m.boucheti, Cadulus (Gadila) Nicklès, 1979: 64, 65,fig. 23. <strong>Recent</strong>. SE <strong>of</strong> Lu<strong>and</strong>a, Angola, WALDA stnCY 15, 10°30.3’S, 11°55’E, 1756 m. HolotypeMNHN. Gadila boucheti (Nicklès, 1979). Knownfrom type locality only.† bouei, Dentalium Deshayes, 1825: 355, 356, pl. 18,fig. 8. Tertiary. Baden, near Vienna, Austria. Juniorsynonym: Dentalium intermedium Hörnes, 1856(fide Pilsbry & Sharp 1898: 200). Antalis bouei(Deshayes, 1825) (fide Pavia 1991: 114). Dentaliumbonei [Arduini 1895: 195] <strong>and</strong> Dentalium borcei[Michelotti, 1861: 136] are incorrect subsequentspellings.† bouryi, Siphonodentalium (Dischides) Cossmann,1888: 13, pl. 1, figs 6, 7. Eocene. Parnes <strong>and</strong>Montainville, France. Dischides bouryi (Cossmann,1888) (fide Pilsbry & Sharp 1898: 235; fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 86).† boussaci, Dentalium Emerson, 1954: 183.Oligocene, Priabonian. Faudon, Allons <strong>and</strong>Martinets, France. Replacement name for Dentaliummartini Boussac, 1911, a junior homonym <strong>of</strong>Dentalium martini Whitfield, 1882 (Polychaeta).brazieri, Cadulus Finlay, 1927. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.braziliensis, Cadulus (Platyschides) Henderson, 1920:124, 125, pl. 19, fig. 16. <strong>Recent</strong>. Off Rio deJaneiro, USBF stn 2762, 23°08’S, 41°34’W, 59 fms[108 m]. Holotype <strong>and</strong> paratype USNM 96113.Gadila braziliensis (Henderson, 1920). WesternAtlantic: Brazil. 23-197 m.† brazosensis, Cadulus (Gadila) Hodgkinson, 1974:11, 12, text-figs 8p, q; pl. 1, figs 7-10, pl. 2, fig. 7.Middle Eocene, Claibornian Stage, Cook MountainFormation, Wheelock Member. Little Brazos River,Brazos County, Texas, USA. Holotype USNM Paleobiology180431; paratypes USNM Paleobiology180448, PRI 29222, ANSP 31500 (all from Scott’slocality MS-2-N), USNM Paleobiology 180432-180434 (locality MS-2A-S).† breve, Dentalium Deshayes, 1861: 201, 202, pl. 1,figs 7, 8. Eocene. Paris Basin (Châlons-sur-Vesles,Jonchery), France. Fustiaria brevis (Deshayes, 1861)(fide Newton 1891: 284). Remark: a valid <strong>species</strong> <strong>of</strong>Dentalium (Entaliopsis) (fide Le Renard 1995: 175;Le Renard & Pacaud 1995: 85).brevicornu, Dentalium (Compressidens) Sharp &Pilsbry in Pilsbry & Sharp, 1897: 125, pl. 22,figs 53-55. <strong>Recent</strong>. Near Galapagos Isl<strong>and</strong>s, USFCstn 2808, 0°36’30”S, 89°19’W, 634 fms [1159 m].Syntypes USNM 122809 (1), 600415 (1); LACM1757 (1, as paratype), SBMNH 35013 (1). Compressidensbrevicornu (Sharp & Pilsbry in Pilsbry &Sharp, 1897). Eastern Pacific, Galapagos. 1159-1819 m.† brevifissum, Dentalium Deshayes, 1825: 366, pl. 17,figs 13, 14. [Fossil]. Angers, Touraine, France.Dentalium brevissimum [Anton, 1838: 25] is anincorrect subsequent spelling (fide Pilsbry & Sharp1898: 200).brevifissum, Dentalium – Galeotti 1837: 150. Tertiary.Jette, Belgium. Junior homonym <strong>of</strong> Dentalium brevifissumDeshayes, 1825. Remarks: Pilsbry & Sharp(1898: 201) stated that Dentalium butini [sic] Nystis a “new name” for Galeotti’s <strong>species</strong>. However,Nyst did not cite Galeotti’s work, <strong>and</strong> D. burtiniNyst in d’Omalius-d’Halloy, 1853 is a nomennudum. We prefer not to rename this <strong>species</strong> untilits status is clarified.brevifissum, Dentalium Brugnone, 1876: 20, 21,fig. 30. Tertiary. Monte Ficarazzi, Italy. Juniorhomonym <strong>of</strong> Dentalium brevifissum Deshayes,1825. Replacement name: Dentalium brugnonein. nom.† breviforme, Dentalium Mayer-Eymar, 1904: 312[description]; 1903: 283 [name only]. Tertiary.Siout Mountain, Egypt.† brevis, Gadus Deshayes, 1861: 219, 220, pl. 3,figs 25-28. Eocene. Paris Basin (Aizy <strong>and</strong> Hérouval),France. Remark: a valid <strong>species</strong> <strong>of</strong> Dischides(fide Le Renard 1995: 175; Le Renard & Pacaud1995: 86).brevissimum, see Dentalium brevifissum Deshayes,1825.† brongniarti, Dentalium Deshayes, 1861: 212, pl. 2,figs 20, 21. Eocene. Paris Basin (Montmirel,Parnes, Damery, Brasles, Mouch, Vaudancourt,Fercourt <strong>and</strong> Saint-Félix), France. Remark: a valid<strong>species</strong> <strong>of</strong> Dentalium (Entaliopsis) (fide Le Renard1995: 175; Le Renard & Pacaud 1995: 85).browni, Dentalium Hisinger, 1837. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† brugnonei, Dentalium n. nom. Tertiary. MonteFicarazzi, Italy. Replacement name for Dentaliumbrevifissum Brugnone, 1876 non Dentalium brevifissumDeshayes, 1825.570ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)† bruneti, Dentalium n. nom. Upper Miocene,Entrerriense Formation. Baliza Punta Flecha,Puerto Madryn, Chubut Province, Argentina.Holotype MPEF-PI-080. Replacement name forDentalium asperum Brunet, 1995 non Dentaliumasperum Michelotti, 1847.† bruscasensis, Cadulus (Gadila) Weisbord, 1964: 130,131, pl. 18, figs 15, 16. Pliocene, Playa Gr<strong>and</strong>eFormation. Station W-26, Quebrada Las Bruscas,Venezuela. Holotype PRI 26897.brycei, Gadila Lamprell & Healy, 1998: 155, 156,figs 155H, 156G, 164, 166A-C. <strong>Recent</strong>. NNW <strong>of</strong>Dampier, Western Australia, 19°34’S, 116°08’E,110 m. Holotype WAM 237/94; paratypes WAM64-93 (7; NNW <strong>of</strong> Dampier, 19°34’S, 116°08’E,110 m), WAM 238/94 (10; Exmouth Gulf[Western Australia]), AMS C311959 (1; MonkeyMia, Shark Bay [Western Australia], 25°48’S,113°43’E), AMS C173593 (1; 19°00.8’S,118°01.3’E, 133 km NNW <strong>of</strong> Port Hedl<strong>and</strong>,112 m), AMS C173754 (5; 220 km NW <strong>of</strong>Roebuck Bay, 17°34’S, 120°22’E, 188 m), MV69582 (2; 19°05’S, 117°26’E, North West Shelf,between Port Hedl<strong>and</strong> <strong>and</strong> Dampier, 122 m).Gadila brycei Lamprell & Healy, 1998. IndianOcean: Western Australia. 40-330 m.buccinulum, Dentalium Gould, 1859: 166. <strong>Recent</strong>.Kagosima [Japan]. Lectotype USNM 24160 (designatedby Johnson, 1964: 48; as “holotype”); paralectotypesMCZ 39073 (1), MCZ 169062 (1).Junior synonym: Dentalium motidukii Otuka, 1935(fide Habe 1964: 25). Dentalium buccinulumGould, 1859. Indo-Pacific: Japan, China. 0-50 m.bulbosum, Dentalium Bronn, 1831. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† burdigalinum, Dentalium Mayer, 1864: 357, pl. 14,fig. 4. Miocene. Near Bordeaux, Dax <strong>and</strong> Saucats,France. Dentalium (Laevidentalium) burdigalinumMayer, 1864 (fide Cossmann & Peyrot 1917: 171,pl. 1, figs 53, 58, 59).† buricum, Dentalium (Fissidentalium) Olsson, 1942:233, 234 [81, 82], pl. 6, figs 3-6, 9. Pleistocene.Tuffaceous shales, west side <strong>of</strong> Burica Peninsula,Costa Rica. Holotype PRI 4071; paratypes PRI4072 (4) (fide Brann & Kent 1960: 317).† burnsii, Cadulus floridanus var. Dall, 1892: 446.Miocene. Alum Bluff, Appalachicola River, Florida,USA. Syntypes USNM Paleobiology 112758 (6).Synonym <strong>of</strong> Cadulus floridanus Dall, 1892.burtini, Dentalium Nyst in d’Omalius-d’Halloy, 1853:382. Tertiary. Belgium. Nomen nudum. Remarks:Pilsbry & Sharp (1898: 201) stated that this <strong>species</strong>was a “new name for Dentalium brevifissumGaleotti, 1837 not <strong>of</strong> Deshayes, 1825”. However,Nyst did not mention Galeotti’s <strong>species</strong>.Furthermore, we have not found any subsequentvalidation <strong>of</strong> Nyst’s name. Bellini (1909: 235) listedthis name as one <strong>of</strong> 12 <strong>species</strong> <strong>of</strong> “Dentalium “ thatwere not scaphopods (i.e. Polychaeta), but Bellinidid not cite any references for this transfer <strong>of</strong> Nyst’snomen nudum to the Polychaeta. Dentalium butini[Pilsbry & Sharp, 1898: 201] is an incorrect subsequentspelling.burtonae, Dentalium (Dentalium) Lamprell & Healy,1998: 33, 34, figs 20F, 21F, 28. <strong>Recent</strong>. RoebuckBay, Broome, Western Australia; 18°03’S, 122°17’E.Holotype AMS C173327; paratypes AMS C174897(1), AMS C307828 (1; Lady Musgrave Isl<strong>and</strong>,Bunker Group, Queensl<strong>and</strong>, dredged in lagoon),AMS C174526 (11; 17°58’S, 122°14’E, BroomeTownship, Western Australia), AMS C173297 (16;18°06’S, 122°20’E, Roebuck Bay, Broome, WesternAustralia, 1-3 m). Dentalium burtonae Lamprell &Healy, 1998. Indo-Pacific: Australia. 0-32 m.bushi, Siphonodentalium (Pulsellum) Henderson,1920: 94, pl. 16, fig. 6. <strong>Recent</strong>. Off Barnegat Bay,New Jersey [USA], Albatross USBF stn 2710,40°06’N, 68°01’30”W, 984 fms [1794 m].Holotype USNM 76723; paratypes USNM 76723(4), MCZ 186822. Pulsellum bushi (Henderson,1920). Western Atlantic: New Engl<strong>and</strong> Banks toCape Hatteras (North Carolina). 582-1796 m.bushii, Cadulus “carolinensis var.?” Dall, 1889: 430.<strong>Recent</strong>. Barbados, 100 fms [183 m]. LectotypeMCZ 7745 (designated by Henderson 1920: 126);paralectotypes USNM 95376 (as “holotype” inTurner 1955: 318, in error). Gadila bushii (Dall,1889) (fide Pilsbry & Sharp 1898: 153). WesternAtlanic: Caribbean. 64-360 m.butini, see burtini.† byorituense, Dentalium (Fissidentalium) Nomura,1935: 90, 91 [38, 39], pl. 6 [1], figs 1a-1c. Tertiary.Hakusyaton, Wangwa, Taikwa, Zyo-tusyowan,Kokwan, Sankwako, Sinpo, <strong>and</strong> Bosiho, Taiwan.† caduloide, Dentalium Dall, 1892: 442. Miocene. StMary’s River, Maryl<strong>and</strong>, USA. Syntypes USNMPaleobiology 112745 (numerous). Remark:Schuchert et al. (1905: 219) cited USNM 112745as the “holotype” but did not specify which syntypewas to be the holotype, so this does not constitute avalid lectotype designation.caesura, Dentalium (Pseudantalis) Colman, 1958: 145,fig. 11. <strong>Recent</strong>. Off Woollongong, New SouthWales, Australia, 100 fms [183 m]. Holotype AMSC62230; paratype AMS C62229. Fustiaria caesura(Colman, 1958) (fide Scarabino 1995: 279;Lamprell & Healy 1998: 102). Indo-Pacific: Philippinesto eastern Australia <strong>and</strong> New Caledonia.60-399 m.† calabrum, Dentalium O. G. Costa, 1851: 35, pl. 3,fig. 4. Type locality not specified [Italy?]. Typematerial not located. Remark: although not explicitlystated in the original description, this is not considereda <strong>Recent</strong> <strong>species</strong>, following the opinion <strong>of</strong>Pilsbry & Sharp (1898: 201) listing calabrum in the<strong>fossil</strong> section, as “? Pliocene, Calabria”.calabrum, Dentalium (Antalis) Crema, 1910: 68-70,text-figs 1-4 [ex Monterosato MS]. Pliocene-ZOOSYSTEMA • 2004 • 26 (4)571


Steiner G. & Kabat A. R.Pleistocene. Valle dei Molini near Monteleone <strong>and</strong>Vallone delli Manci near S. Vincenzo la Costa,Italy. Junior synonym <strong>of</strong> Antalis agilis (G. O. Sars,1872) (fide CLEMAM entry No. 24572). Juniorhomonym <strong>of</strong> Dentalium calabrum O. G. Costa,1851. The synonymy with Antalis agilis makes itunnecessary to introduce a replacement name.† calafium, Dentalium (Laevidentalium) Vokes, 1939:105, pl. 16, figs 30, 31. Eocene, DomengueFormation. UCMP locality A-1027, Valdes Ranch,Silver Creek, Priest Valley Quadrangle, California,USA. Lectotype UCMP 15750 (designated bySquires 1984: 16); paralectotype UCMP 15751(UCMP locality A-1003, Pine Canyon, MountDiablo, California, USA).calamus, Dentalium Dall, 1889: 421. <strong>Recent</strong>. TurtleHarbor, Florida, 4 fms [7 m]. Lectotype USNM68117 (designated by Henderson 1920: 72); paralectotypesUSNM 887456. Graptacme calamus(Dall, 1889) (fide Pilsbry & Sharp 1897: 97).Western Atlantic: Cape Lookout (North Carolina)to Brazil <strong>and</strong> Caribbean. 0-124 m.caledonicum, Dentalium Scarabino, 1995: 209,figs 17, 28a, 71c. <strong>Recent</strong>. Southern New Caledonia,RV Coriolis, CHALCAL 2, stn DW74, 24°40’S,168°38’E, 650 m. Holotype MNHN <strong>and</strong> paratypes(1 from stn DW 73; 1 from stn DW 75).Dentalium caledonicum Scarabino, 1995. Indo-Pacific: New Caledonia. 573-650 m.† californicum, Dentalium Stanton, 1895: 62, 63,pl. 13, fig. 3. Cretaceous. Upper part <strong>of</strong> theKnoxville Beds, Shelton’s ranch, 5 miles north <strong>of</strong>Paskenta, California, USA. Type material USNMPaleobiology 23063. Remark: Schuchert et al.(1905: 219) cited USNM 23063 as “holotype <strong>and</strong>paratypes” but did not specify which syntype was tobe the holotype, so this does not constitute a validlectotype designation.californicus, Cadulus (Gadila) Pilsbry & Sharp, 1898:180, 181, pl. 34, figs 5-8. <strong>Recent</strong>. Off San Diego,California [Albatross stn 2923, 32°40’30”N,117°31’30”W, 822 fms (1500 m)]. LectotypeUSNM 107698 (designated by Shimek 1989: 235);paralectotypes USNM 203136, SBMNH 35038,AMNH 148391 (4). Cadulus californicus Pilsbry& Sharp, 1898. Eastern Pacific: Oregon (USA) toPanama. 400-2322 m. Remark: probably referableto Polyschides (fide Emerson in Turgeon 1988: 51);further research is required.† callioglyptum, Dentalium Pilsbry & Sharp, 1898a:468, pl. 10, figs 10-12, pl. 11, fig. 21. Oligocene.Dominican Republic. Syntypes ANSP 2713 (fidePilsbry 1922: 399) <strong>and</strong> ANSP 79569.callipeplum, Dentalium Dall, 1889: 419, 420, pl. 27,fig. 12-B. <strong>Recent</strong>. USBF [Albatross] stn 2400,28°41’N, 86°07’W, between the Missisippi Riverdelta <strong>and</strong> Cedar Keys [<strong>of</strong>f Florida], 169 fms[308 m]. Lectotype USNM 94064 (designated byHenderson 1920: 74); paralectotypes USNM887457 (2), MCZ 7672, USNM 95361 (1; Blakestn 220), USNM 95362 (1; Samana Bay,St Domingo), MCZ 7673 (1; Blake stn 128), MCZ7674 (1; Blake stn 143; erroneously stated to be the“holotype” in Turner, 1955: 318), MCZ 7675(1; Blake stn 262), MCZ 186549 (3; Blake stn143), not found: USFC stn 2314 <strong>and</strong> “Coast <strong>of</strong>Honduras”. Calliodentalium callipeplum (Dall,1889) (fide Scarabino 1995: 273). WesternAtlantic, Carribean: South Carolina to Colombia.14-4000 m.callithrix, Dentalium Dall, 1889: 427, pl. 27, fig. 10.<strong>Recent</strong>. Yucatan Strait, 640 fms [1170 m].Lectotype USNM 95348 (designated by Henderson1920: 57, 58); paralectotypes USNM 93426(4; USFC stn 2678), USNM 93758 (2; USFCstn 2383), USNM 95349 (2; Blake stn 20), USNM95350 (2; Blake stn 236), USNM 95354 (1; Blakestn 248; now the holotype <strong>of</strong> Entalina quadrataHenderson, 1920), MCZ 7730 (1; Blake stn 20),MCZ 7731 (2; Blake stn 41), MCZ 7732 (1; Blakestn 162), MCZ 7733 (1; Blake stn 221), MCZ7734 (1; Blake stn 236; erroneously stated to be the“holotype” in Turner 1955: 318). Pertusiconchacallithrix (Dall, 1889) (fide Chistikov 1982a: 678).Western Atlantic <strong>and</strong> Eastern Atlantic: fromCarribean to West Africa. 12-3500 m.† caloosaense, Dentalium Dall, 1892: 441, pl. 23,fig. 24. Pliocene. Caloosahatchie River <strong>and</strong> ShellCreek, Florida, USA. Syntypes USNM Paleobiology112741 (4); possible syntypes USNMPaleobiology 112740 (6). Remark: Schuchert et al.(1905: 219) cited USNM 112741 as the “holotype”but did not specify which syntype was to be theholotype, so this does not constitute a valid lectotypedesignation.† calvertensis, Prodentalium Palmer, 1975: 378-380,pl. 52, figs 1-11. Middle Jurassic, Callovian.Calvert, Buckinghamshire, Engl<strong>and</strong>, UnitedKingdom. Holotype BMNH GG 13330; paratypesBMNH GG 13331-13339.† camaronesia, Dentalium sulcosum var. Ihering, 1907:224. Tertiary. Camarones, Patagonia, Argentina.Dentalium sulcosum camaronesium Ihering, 1907(fide Ihering 1914: 67).campylus, Cadulus Melvill, 1906: 80, pl. 8, fig. 32.<strong>Recent</strong>. Gulf <strong>of</strong> Oman, 156 fms [285 m]. LectotypeBMNH 1906.10.23.76 (designated by Ludbrook1954: 113, as “holotype”; the figured specimen, fideTrew 1987: 29); paralectotypes BMNH 1955.158.599 (3), NMW. Cadulus campylus Melvill,1906. Indian Ocean: Africa to Maldive Isl<strong>and</strong>s.183-285 m.canaliculatum, Dentalium “Klipstein” Pilsbry & Sharp,1898: 201. Error or emendation for Dentaliumcannaliculatum Klipstein, 1843.cancellatum, Dentalium Sowerby, 1860: 101, pl. 224,fig. 36. <strong>Recent</strong>. China. Syntypes BMNH 1996087(2). Coccodentalium cancellatum (Sowerby, 1860)572ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)n. comb. Indo-Pacific: China. 4 m. Remark: <strong>of</strong>the two syntypes, only the figured specimen correspondswith the <strong>species</strong> description; the secondspecimen is Paradentalium intercalatum (Gould,1859) (fide V. Scarabino, in litt. 20.II.2002).c<strong>and</strong>elatum, Dentalium (Episiphon) Kira, 1959: 105,pl. 40, fig. 5. <strong>Recent</strong>. Tosa Bay, Shikoku, Japan,200 m. Type material not located. Nomen nudum inHabe (1953: 296), <strong>and</strong> in Kira (1954: 80), fideBieler & Petit (1990: 141). Episiphon c<strong>and</strong>elatum(Kira, 1959) (fide Scarabino 1995: 289). Indo-Pacific: Japan. 100-200 m.c<strong>and</strong>idum, Dentalium Jeffreys, 1877: 153. <strong>Recent</strong>.Type locality not specified. Syntypes BMNH85.11.58.1239 (Northern <strong>and</strong> eastern Atlantic <strong>and</strong>Bay <strong>of</strong> Biscay, 410-1750 fms [748-3194 m], figuredsyntype), BMNH 85.11.58.1240 (west <strong>of</strong> Irel<strong>and</strong>,664 -1476 fms [1212-2694 m]), USNM 169428(Valorous stn 5, 66°59’N, 55°27’W, 410 fms [DavisStrait, Greenl<strong>and</strong>, North Atlantic, 748 m]), USNM169429 (Valorous stn 8, 62°06’N, 55°56’W,1350 fms [Davis Strait, Greenl<strong>and</strong>, North Atlantic,2464 m]), USNM 169430 (Valorous stn 6,64°05’N, 56°47’W, 1100 fms [Davis Strait,Greenl<strong>and</strong>, North Atlantic, 2008 m]), USNM169431 (Porcupine 1869 stn 28, 56°44’N,12°52’W, 1215 fms [Hebrides, Scotl<strong>and</strong>, NorthAtlantic, 2217 m]), USNM 169432 (Porcupine1869 stn 23, 56°07’N, 14°19’W, 664 fms [Irel<strong>and</strong>,North Atlantic, 1212 m]), USNM 169433(Porcupine 1869 stn 21, 55°40’N, 12°46’W,1476 fms [Irel<strong>and</strong>, North Atlantic, 2694 m]),USNM 169434 (Porcupine 1869 stn 23-A,56°07’N, 14°18’W, 420 fms [Irel<strong>and</strong>, NorthAtlantic, 767 m]), USNM 169435 (Porcupine 1869stn 31, 56°15’N, 11°25’W, 1360 fms [Irel<strong>and</strong>,North Atlantic, 2482 m]), USNM 169436,(Porcupine 1869 stn 38, 47°39’N, 11°33’W,2090 fms [Irel<strong>and</strong>, North Atlantic, 3814 m]),USNM 169438 (Off Cape Carnveira, Portugal,Porcupine 1870 stn 16, 39°55’N, 9°56’W, 994 fms[1814 m]). Junior synonyms: Dentalium solidumVerrill, 1884 (fide Pilsbry & Sharp 1897: 72);Dentalium caudani Locard, 1896; Dentaliumcaudani var. parfaiti Locard, 1898 (B. Métivier,n. syn.); Dentalium (Fissidentalium) c<strong>and</strong>idum var.meridionale Pilsbry & Sharp, 1897; Dentalium(Fissidentalium) meridionale var. jamaicenseHenderson, 1920 (fide Scarabino 1994: 307).Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877) (fidePilsbry & Sharp 1897: 72). Western Atlantic:Greenl<strong>and</strong> to Brazil; Eastern Atlantic: Greenl<strong>and</strong> toSouth Africa. 403-3814 m.† canna, Dentalium White, 1874: 23; 1875: 156,pl. 12, fig. 6. “Carboniferous” [Permian]. Near SaltLake, New Mexico, USA; near Relief Spring,Arizona, USA. Syntypes USNM Paleobiology8473. Prodentalium canna (White, 1874) (fideYancey 1978: 308, 309, pl. 6, figs 7-11). Juniorsynonyms: Dentalium mexicanum Girty, 1909 (fideYancey 1978: 308); Plagioglypta monolineataBranson, 1930 (fide Yancey 1973: 1126; 1978:308, 309).† cannaliculatum, Dentalium Klipstein, 1843: 206,pl. 14, figs 28a, 28b. Triassic (St Cassian?). Austria/northern Italy. Remark: Dentalium “canaliculatum”Pilsbry & Sharp, 1898: 201, error or emendation.capense, Dentalium (Compressidens) Tomlin, 1931:340. <strong>Recent</strong>. South Africa, <strong>of</strong>f Cape Point, 900 fms[1620 m]. Holotype SAFM A6191. Junior synonym<strong>of</strong> Rhomboxiphus tricarinatus (Boissevain,1906) (fide Scarabino 1995: 316).capillosum, Dentalium Jeffreys, 1877: 153, 154.<strong>Recent</strong>. Type locality not designated. SyntypesBMNH 1885.11.5.1336 (Porcupine 1869, 862 fms[Bay <strong>of</strong> Biscay, 1573 m]), BMNH 1877.11.28.55(Valorous stn 12, 56°11’N, 37°41’W, 1450 fms[Greenl<strong>and</strong>, 2646 m]), BMNH 1887.2.9.4-6 (<strong>of</strong>fthe Azores, Challenger stn 73, 38°30’N, 31°14’W,1000 fms [1825 m]), USNM 169417 (2) (Porcupine1869 stn 47, 59°34’N, 7°18’W, 542 fms[Hebrides, North Atlantic, 989 m]), USNM169419 (Valorous stn 12, 56°11’N, 37°41’W,1450 fms [Greenl<strong>and</strong>, 2646 m]), USNM 169420(Valorous stn 13, 56°01’N, 34°42’W, 690 fms[Greenl<strong>and</strong>, 1259 m]), USNM 169421 (Valorousstn 16, 55°10’N, 25°58’W, 1785 fms [Greenl<strong>and</strong>,3258 m]), USNM 169423 (2) (<strong>of</strong>f Cape Mondego,Portugal, Porcupine 1870, 220 fms [402 m]),USNM 169424 (<strong>of</strong>f Cape Carnveira, Portugal,Porcupine 1870, 1095 fms [1998 m]), USNM169425 (<strong>of</strong>f Cape Carnveira, Portugal, Porcupine1870, 984 fms [1796 m]). Junior synonyms:Dentalium ergasticum Fischer, 1883, Dentaliumscamnatum Locard, 1897, Dentalium milneedwardsiLocard, 1898 (B. Métivier, n. syn.; see Appendix 2).Fissidentalium capillosum (Jeffreys, 1877) (fidePilsbry & Sharp 1897: 77). Eastern Atlantic: Greenl<strong>and</strong>to Western Africa <strong>and</strong> Azores. 100-4088 m.caprinum, Dentalium Bosc, 1802: 191. <strong>Recent</strong>. IndianOcean (“mer des Indes”) [citing Martini 1769: I,t. 1, f. 4 B]. Type material not located. Junior synonym<strong>of</strong> Dentalium aprinum Linnaeus, 1767 (fidePilsbry & Sharp 1897: 3). Remarks: Pilsbry &Sharp (1897: 3) attributed this name to Anton(1838: 25); in fact, Anton cited the same figurefrom Martini, which indicates that Anton usedBosc’s name without attribution. It is tempting toassume that Bosc used caprinum in error for aprinum.† caprottii, Cadulus (Sulcogadila) Moroni & Ruggieri,1981: 30, figs 1-3. Lower Pleistocene. Acqua deiCorsari, Palermo, Italy. Holotype Museo Geologico,Univ. Palermo, O.C.R. 3131.carduus, Dentalium Dall, 1889: 423, pl. 27, fig. 3.<strong>Recent</strong>. Off St Lucia, Lesser Antilles, Blake stn 220,13°50’N, 61°03’W, 116 fms [211 m]. LectotypeMCZ 7692 (designated by Henderson 1920: 33);paralectotypes USNM 95321 (Blake stn 246),ZOOSYSTEMA • 2004 • 26 (4)573


Steiner G. & Kabat A. R.USNM 95322 (USBF stn 2655), MCZ 7691(Blake stn 246). Junior synonym: Dentalium floridenseHenderson, 1920 (fide Scarabino 1994: 307).Fissidentalium carduus (Dall, 1889) (fide Emersonin Turgeon 1988: 50; 1998: 200). Western Atlantic,Caribbean. 180-600 m. Remarks: Emerson (1952a: 2)assigned carduus to Coccodentalium. Emerson inTurgeon (1988: 50; 1998: 54) listed Fissidentaliumfloridense as a distinct <strong>species</strong>.carinatum, Dentalium O. G. Costa 1851. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.carlessi, Gadila Lamprell & Healy, 1998: 154, 155,figs 155a, 156a, 161. <strong>Recent</strong>. East <strong>of</strong> Cape Melville,northern Queensl<strong>and</strong>, 14°22’S, 145°52’E, 2710 m.Holotype QM MO40084. Gadila carlessi Lamprell& Healy, 1998. Known from the type locality only.carneum, Dentalium (Episiphon) Boissevain, 1906: 48,pl. 6, figs 42-45. <strong>Recent</strong>. Indonesia, Flores Sea,Siboga stn 45, 07°24’S, 118°15.2’E, 794 m.Lectotype ZMA 3.06.052 (designated by Scarabino1995: 287); paralectotypes ZMA 3.06.053 (14),ZMA 3.06.054 (1; Timor Sea, Siboga stn 300,10°48.6’S, 123°23.1’E, 918 m), ZMA 3.06.055(2; Ambon, Siboga stn 181, 36-54 m). Junior synonym<strong>of</strong> Episiphon virgula (Hedley, 1903) (fideScarabino 1995: 287).† carolinense, Dentalium Conrad, 1862a: 288; 1863:570. Miocene. North Carolina, USA.carolinensis, Cadulus Bush in Verrill, 1885b: 587;Bush 1885: 471, pl. 45, fig. 19. <strong>Recent</strong>. Off CapeHatteras, USBF stn 2114, 35°20’N, 75°20’W,14 fms [26 m]. Syntypes USNM 35811 (figured),MCZ 186818 (4). Polyschides carolinensis (Bush inVerrill, 1885) (fide Pilsbry & Sharp 1898: 152).Western Atlantic, Caribbean. 5-159 m.† caronensis, Cadulus Mansfield, 1925: 61, pl. 7,fig. 6. Miocene. About 1 mile S <strong>of</strong> Brasso, CaroniCounty, Montserrat Ward, Trinidad. Figured syntypeUSNM Paleobiology 352692 (1).† cartagenense, Dentalium Pilsbry & Brown, 1917: 37,38, pl, 5, fig. 10. Oligocene. Near Cartagena,Colombia. Holotype ANSP 3058.† caseyi, Dentalium n. nom. Oligocene. Red Bluff,Mississippi, USA. Doubtful syntype USNMPaleobiology 481673 (1) (fide MacNeil & Dockery1984: 247). Replacement name for Dentalium polygonumCasey, 1903 non Dentalium polygonumReuss, 1844.† casselensis, Gadila R. Janssen, 1989: 89, pl. 2,fig. 18. Upper Oligocene, Chattian. Glimmerode,near Hessisch-Lichtenau, Niederhessen, Germany.Holotype SMF 308322; paratypes SMF 308323/2<strong>and</strong> Collection Gürs.castellaccii, see castellacei.† castellacei, Dentalium Scalia, 1914: 2, pl. 1 [7],fig. 5. Triassic. Castellace, Italy. Type material inIstituto di Geologia della R. Università Catania.Dentalium castellaccii [Diener, 1926: 228] is anincorrect subsequent spelling.† castellanensis, Dentalium d’Orbigny, 1850 (volume 2):320. Eocene, Suessonian. Le Vit, near Castellanne(Basses-Alpes) [Alpes-de-Haute-Provence] <strong>and</strong>Bidart (Basses-Pyrénées) [Pyrénées-Atlantiques],France. Syntypes MNHN (fide Boussac 1911b:256, pl. 16, figs 7, 8). Boussac (1911) erroneouslyused Dentalium haeringense Dreger, 1892 as thevalid name for this <strong>species</strong>.catenulatum, Dentalium Chenu, 1843: pl. 4, fig. 11;1850: 2. Piedmont (Miocene/Pliocene; Italy).Junior synonym <strong>of</strong> Dentalium <strong>fossil</strong>e Gmelin, 1791(fide Pilsbry & Sharp 1898: 201).catharus, Cadulus (Platyschides) Henderson, 1920:112, 113, pl. 18, fig. 5. <strong>Recent</strong>. Off EnglishHarbor, Antigua, State Univ. Iowa Exp. 1898stn 15, 120 fms [219 m]. Holotype USNM 596596.Cadulus catharus Henderson, 1920. WesternAtlantic, Caribbean. 100-219 m. Remark: althoughthe subgenus Platyschides was put in synonymy withGadila by Scarabino (1995: 196, 355), this <strong>species</strong>with its simple apical orifice <strong>and</strong> the maximumdiameter in the middle <strong>of</strong> the shell belongs toCadulus.† catulloi, Dentalium Vinassa de Regny, 1896: 204,205, pl. 5, fig. 2a, b. Miocene. Italy. Remark: as anomen nudum in Vinassa de Regny (1895: 262).caudani, Dentalium Locard, 1896a: 213; 1896b: 171,pl. 6, fig. 2. <strong>Recent</strong>. Bay <strong>of</strong> Biscay, Caudan stn 15,3°02’W [5°22’W, Paris meridian], 44°02’N,1300 m. Holotype (preserved specimen withoutshell) in Faculté des Sciences, Lyon (fide B. Métivierpers. comm.). Junior synonym <strong>of</strong> Fissidentaliumc<strong>and</strong>idum (Jeffreys, 1877) (B. Métivier, n. syn.).caymanica, Costentalina Chistikov, 1982b: 1314, pl. 1figs 5, 6, pl. 3 figs 1-11. <strong>Recent</strong>. Cayman Trench[Caribbean Sea], Akademik Kurtschatow stn 1267 T,19°38.7’N, 76°37.1’W, 6740-6780 m. HolotypeIOM; paratypes IOM (7 from 1267 T; 1 from1267 D, the type locality; 1 from stn 1259 D,19°05.6’N, 80°30.1’W, 5930 m; 5 from stn 1266,19°45.4’N, 76°43.8’W, 6300-5900 m). Costentalinacaymanica Chistikov, 1982. Western Atlantic,Caribbean: Cayman Trench. 5900-6780 m.† cazadorianum, Dentalium Wilckens, 1907: 121,122 [25, 26], pl. 4, fig. 9. Upper Cretaceous.Cazador <strong>and</strong> Cancha Carrera, Patagonia, Argentina.ceciliae, Compressidentalium Scarabino, 1995: 263,figs 69, 70k, 71i. <strong>Recent</strong>. New Caledonia, RVVauban MUSORSTOM 4, stn DW 220, 22°58’S,167°38’E, 505-550 m. Holotype MNHN; paratypesMNHN (7 from MUSORSTOM 4 stn DW220; 4 from BIOCAL stn DW 51), AMS C201725(1; BIOCAL stn DW 54), NMNZ M268958(1; MUSORSTOM 4 stn DW 220), USNM 890864(1; BIOCAL stn DW 51). Compressidentaliumceciliae Scarabino, 1995. Indo-Pacific: NewCaledonia. 505-700 m.cedrosense, Dentalium (Rhabdus) Hertlein & Strong,1951: 69, 70, pl. 11, fig. 9. <strong>Recent</strong>. <strong>of</strong>f the east574ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)coast <strong>of</strong> Cedros Isl<strong>and</strong>, Lower California, Mexico,Templeton Crocker stn 126-D-12, 28°20’00”N115°10’30”W, 45 fms [82 m]. Holotype CAS.Junior synonym <strong>of</strong> Episiphon innumerabile (Pilsbry& Sharp, 1897) (fide Keen 1971: 888).† centenniale, Dentalium Marwick, 1942: 279, pl. 24,fig. 26. Eocene, Bortonian. Hampden Beach, NE <strong>of</strong>Otago, New Zeal<strong>and</strong>. Holotype NZGS TM4802(fide Keyes 1972: 95). Remarks: based on theOnekakara (Hampden Beach) paralectotypes <strong>of</strong>Dentalium mantelli Zittel, 1864, which Marwickconcluded represented two <strong>species</strong> (fide Dawson1991: 23; 1994: 118). Maxwell (1992: 185)suggested that this <strong>species</strong> “may also belong inFissidentalium”.ceras, Dentalium Watson, 1879: 510, 511 [Watson1886: 3, pl. 1, fig. 4]. <strong>Recent</strong>. Mid Pacific, East <strong>of</strong>Japan, Challenger stn 246, 36°10’N, 178°00’E,2050 fms [3690 m]. Syntypes BMNH 1887.2.9.10-11 (the specimen from stn 299, 33°31’S,74°43’W, 2160 fms, west <strong>of</strong> Valparaiso, Chile wasnot found in BMNH). Fissidentalium ceras(Watson, 1879) (fide Ludbrook 1954: 93). Indo-Pacific: India to Japan; Eastern Pacific: Chile.3690-3747 m. Remark: Scarabino (1995: 250)considered megathyris a junior synonym <strong>of</strong>Fissidentalium ceras (Watson, 1879). However, thisproposed synonymy requires a comparative study <strong>of</strong>all taxa included in this <strong>species</strong> <strong>group</strong>.ceratum, Dentalium Dall, 1881: 38. <strong>Recent</strong>. Off LaHavana, Cuba, Blake stn 2, 23°14’N, 82°25’W,805 fms [1472 m]. Lectotype USNM 95326 (designatedby Henderson 1920: 50); paralectotypesUSNM 887458, MCZ 7697, MCZ 7706 (WestFlorida, 50 fms [91 m]). Junior synonym: Dentalium(Antalis) ceratum tenax Henderson, 1920(n. syn.). Antalis cerata (Dall, 1881) (fide Pilsbry &Sharp 1897: 57). Western Atlantic, Caribbean:Florida to Rio de la Plata. 32-1960 m.cerinum, Dentalium (Rhabdus) Pilsbry, 1905: 117, 118,pl. 5, figs 40, 41. <strong>Recent</strong>. Shimizu, Suruga, Japan.Holotype ANSP 88305. Junior synonym: Dentaliumtracheatum Boissevain, 1906 (fide Scarabino1995: 297). Omniglypta cerina (Pilsbry, 1905) (fideHabe 1953: 296). Indo-Pacific: Indonesia to Japan<strong>and</strong> northeastern Australia. 800-3925 m.cestum, Dentalium (Dentalium) texasianum Henderson1920: 29, pl. 2, figs 1, 4, 8. <strong>Recent</strong>. Pass Cabello,Texas. Holotype USNM 159503. Synonym <strong>of</strong>Dentalium texasianum Philippi, 1848, a junior synonym<strong>of</strong> Paradentalium americanum (Chenu,1843) (fide Scarabino 1975: 183).cheverti, Dentalium (Dentalium) Sharp & Pilsbry inPilsbry & Sharp, 1897: 9. <strong>Recent</strong>. Evans Bay, CapeYork, North Australia, 6 fms [11 m]. Lectotype (<strong>of</strong>septemcostatum Brazier) AMS A96 (designated byPonder & Stanbury 1972: 53) <strong>and</strong> 3 paralectotypes.Replacement name for Dentalium septemcostatumBrazier, 1877, a junior homonym <strong>of</strong> Dentaliumseptemcostatum Abich, 1859. Dentalium chevertiSharp & Pilsbry in Pilsbry & Sharp, 1897. Indo-Pacific: eastern Australia. 4-738 m.† chilense, Dentalium d’Orbigny in Grange, 1847:pl. 4 (Paléontologie pl. 1), figs 37, 38. No localitygiven. Remarks: no description <strong>of</strong> this <strong>species</strong> waspublished in the text; the <strong>species</strong> attributed tod’Orbigny in Grange are all “name <strong>and</strong> figure” only(fide Sherborn & Woodward 1901a: 391). Pilsbry& Sharp (1898: 220) stated that this <strong>species</strong> wasfrom the Cretaceous <strong>of</strong> Chile. Feruglio (1949a:304; 1949b: 249) stated that this <strong>species</strong> was fromthe Senonian <strong>of</strong> Quiriquina (Chile).chinensis, Striodentalium Qi & Ma, 1989: 119, fig. 11.<strong>Recent</strong>. East China Sea, 52-173 m. Type depositorynot specified. Striodentalium chinensis Qi & Ma,1989. Indo-Pacific: China Sea. 52-173 m.† chipolanum, Dentalium (Antalis) Gardner, 1947:625, 626, pl. 57, figs 18-18a. Oligocene. AlumBluff Group, Chipola Formation. USGS locality3419, McClell<strong>and</strong> Farm, 1 mile below Bailey’sFerry, Calhoun County, Florida, USA. HolotypeUSNM Paleobiology 329137.† chubutensis, Dentalium (Laevidentalium) Sabattini,1979: 21, pl. 1, figs 1, 2. Upper Carboniferous,Tepuel Group. County Toba, Colán-Conhué,Sierra de Languineo, Chubut Province, Argentina.Holotype MLP 13164; paratype MLP 13165.chuni, Cadulus (Cadulus) Jaeckel, 1932: 309, fig. 6.<strong>Recent</strong>. Somalia, Africa, Valdivia stn 256, 1°49’N,45°29.5’E, 1134 m. Lectotype ZMB 75374a (designatedby Kilias 1995: 173); paralectotypes ZMB75374b (stn 256), ZMB 75375 (stn 245). Caduluschuni Jaeckel, 1932. Indo-Pacific: Africa toPhilippines. 205-1134 m. Remark: the figuredspecimen from Valdivia stn 245, ZMB 75375, wasso degraded that Kilias chose a specimen from stn256 as the lectotype.chuni, Dentalium (Fissidentalium) Plate, 1908a: 341-343, pl. 30, figs 1-9. <strong>Recent</strong>. East African Coast,Valdivia stn 251, 1°40.6’S, 41°47.1’E, 693 m.Lectotype ZMB 63824 (designated by Kilias 1995:172); paralectotypes ZMB 61100 (stn 256), ZMB61102 (stn 254), ZMB 61103 (stn 191), ZMB63821 (stn 264), ZMB 63822 (stn 256), ZMB63823 (stn 254). Junior synonym <strong>of</strong> Fissidentaliumshopl<strong>and</strong>i (Jousseaume, 1894) (fide Scarabino 1995:248). Remark: V. Scarabino (in litt. 5.XI.2001)suggested that the status <strong>of</strong> D. chuni vis-à-visFissidentalium transversostriatum (Boissevain, 1906)requires further research.† cidaris, Dentalium Geinitz, 1850: 144, 145.Cretaceous. Niederwarthla <strong>and</strong> Strehlen, Germany;Bohemia, Czech Republic. Name for Dentaliumstriatum “J. Sowerby, 1814” sensu Geinitz 1839.Junior synonym: Dentalium reussianum Ryckholt,1851 (fide Pilsbry & Sharp 1898: 220).cinctum, Dentalium de Koninck, 1843. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.ZOOSYSTEMA • 2004 • 26 (4)575


Steiner G. & Kabat A. R.† cinctum, Dentalium Goldfuss, 1841: 3, pl. 166, fig. 7.Jurassic. Derneberg, near Hannover, Germany.Objective junior synonym: Dentalium muensteriPilsbry & Sharp, 1898. Remark: Pilsbry & Sharp(1898: 224) erroneously dated this name to 1844, <strong>and</strong>therefore concluded that it was a junior homonym <strong>of</strong>Dentalium cinctum de Koninck, 1843. Thus, Pilsbry& Sharp proposed Dentalium muensteri as a replacementname for what is, in fact, the senior homonym.cinerascens, Dentalium Anton, 1838: 25. <strong>Recent</strong>.Adriatic Sea [in error]. Tentative (senior!) synonym<strong>of</strong> Antalis occidentalis (Stimpson, 1851) (fidePilsbry & Sharp 1897: 48). Remark: pursuant toICZN (1999) article 23.9 we propose to preserveStimpson’s name, occidentalis, for this <strong>species</strong>. Thename cinerascens has not been used whereas thename occidentalis has consistently been used for this<strong>species</strong> after 1851.cingulatum, Dentalium Hoeninghaus, 1831: 155.Tertiary. Nomen nudum.cingulatus, Dentalites Schlotheim, 1820. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† circinatum, Dentalium G. B. Sowerby, 1823:pl. 138, fig. 5. No locality given. Remarks: Pilsbry& Sharp (1898: 201) stated that this <strong>species</strong> wasfrom the Eocene, Paris Basin, France; Newton(1891: 284, 285) recorded this <strong>species</strong> from theMiddle Eocene <strong>of</strong> Engl<strong>and</strong>. Type <strong>species</strong> <strong>of</strong> thegenus Fustiaria Stoliczka, 1868, by subsequent designation(Pilsbry & Sharp 1897: 127). Newton(1891: 285, 286) misidentified Dentalium subeburneumd’Orbigny, 1850, as Fustiaria circinata(fide Pilsbry & Sharp 1898: 216).circumcinctum, Dentalium Watson, 1879: 513, 514[Watson 1886: 7, pl. 1, fig. 7]. <strong>Recent</strong>. Type localitynot specified. Syntypes BMNH 1887.2.9.24-25(Challenger stn II, Setubal [Portugal], 38°10’N,9°14’W, 470 fms [858 m]), BMNH 1887.2.9.26(Challenger stn 23, Sombrero Isl<strong>and</strong>, St Thomas,450 fms [821 m]), BMNH 1887.2.9.30 (figured specimen;Challenger stn 122, Pernambuco [Brazil],9°05’S, 34°50’W, 350 fms [639 m]). Antalis circumcincta(Watson, 1879) (fide Thomas 1975: 293). WesternAtlantic: Caribbean <strong>and</strong> Brazil; Eastern Atlantic:North Africa <strong>and</strong> Gulf <strong>of</strong> Guinea. 238-1935 m.Remark: Watson (1886: 7) subsequently recorded2 specimens from Challenger stn 56 (<strong>of</strong>f Bermuda,32°8’45”N, 64°59’35”W, 1075 fms [1962 m]),which are in the type collection (BMNH 1887.2.9.27-28), but these specimens are not syntypes.citrina, Dentalium vulgare var. Monterosato, 1884: 32.Nomen nudum without type locality or description.† claibornense, Dentalium (Antalis) thalloides var.Palmer, 1937: 14, 15, pl. 1, figs 1, 2, 5. Eocene,Claiborne Formation. Locality 104, Gosport s<strong>and</strong>,Mississippi, USA. Syntypes PRI 2626, PRI 2627.† clarae, Cadulus Maury, 1910: 151, pl. 8, fig. 7.Oligocene. Oak Grove, Santa Rosa County, Florida,USA. Syntype USNM Paleobiology 639096 (1).clathratum, Dentalium von Martens, 1881: 66.<strong>Recent</strong>. Off Moreton Bay [Queensl<strong>and</strong>], Australia,550 fms [1006 m]. Holotype ZMB 33122.Compressidentalium clathratum (von Martens,1881) (fide Scarabino 1995: 263). Indo-Pacific:Australia. 23-1606 m. Remark: Lamprell & Healy(1998) listed Dentalium sedecimcostatum Boissevain,1906 as a junior synonym <strong>of</strong> C. clathratum, anopinion not shared by Scarabino (1995). However,V. Scarabino (in litt. 5.XI.2001) stated that he nowagrees with this synonymy.clausum, Dentalium Turton, 1819. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.clava, Dentalium Lamarck, 1818. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.clavatum, Dentalium Gould, 1859: 166. <strong>Recent</strong>.Hong Kong [China]. Lectotype USNM 24245(designated by Johnson 1964: 239, as “holotype”);paralectotypes MCZ 169089 (1), MCZ 216584 (1).Gadila clavata (Gould, 1859) (fide Scarabino 1995:364). Indo-Pacific: South Africa, Red Sea, China.10-175 m.clavus, Dentalium Cooke, 1885: 275. <strong>Recent</strong>. Gulf <strong>of</strong>Suez. Lectotype (<strong>of</strong> D. mac<strong>and</strong>rewi Boissevain,1906 designated by Scarabino 1995: 204) not inZMA (fide R. Moolenbeek & A. N. Van der Bijl inlitt. 23.VI.2000) or in UMZC; paralectotypesUMZC I.100,745. Junior synonyms: Dentaliumaratorum Cooke, 1885; Dentalium mac<strong>and</strong>rewiBoissevain, 1906 (fide Scarabino 1995: 204; Singer2003: 11, 14). Dentalium clavus Cooke, 1885 (fideSinger 2003: 13, 14). Indian Ocean: Red Sea,Africa. 14-60 m. Remarks: Boissevain (1906: 25)replaced this name, together with <strong>and</strong> Dentaliumlineolatum, with Dentalium mac<strong>and</strong>rewi. Scarabino(1995: 204) placed D. clavus in synonymy withD. reevei Fischer, 1871.coarctatum, Dentalium Brocchi, 1814. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.Remarks: Dentalium coarctatum – Lamarck 1818:346 was subsequently described as D. ventricosumBronn, 1828; D. coarctatum – O. G. Costa 1851:38, 41, 42, was subsequently described as Cadulusmedius Pilsbry & Sharp, 1898.coarti, Dentalium Dautzenberg, 1912: 81, pl. 3,figs 44, 45. <strong>Recent</strong>. Type locality not specified.Syntype MNHN (Wharf <strong>of</strong> Tamara [Guinea]);other syntypes not located, described from the followinglocalities: between Conakry <strong>and</strong> Boulbine,Guinea; southwest <strong>of</strong> Conakry, Guinea; west <strong>of</strong>Crawford Bank, Guinea; near Île Roume, Archipelde Los, Guinea; Gr<strong>and</strong> Bassam, 15 m, Côte d’Ivoire;<strong>of</strong>f Cotonou, 20-25 m, Benin; Libreville Bay,Gabon. Junior synonym <strong>of</strong> Dentalium congoensisPlate, 1908 (fide Nicklès 1979: 44).cobbi, Gadila Lamprell & Healy, 1998: 155,figs 155G, 156D, 163. <strong>Recent</strong>. Off Cairns, northernQueensl<strong>and</strong>, Australia, 16°37.7’S, 146°20.4’E,685-658 m. Holotype AMS C174638; paratypes576ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)AMS C173427 (1), AMS C8017 (2; DarnleyIsl<strong>and</strong>, Torres Strait, Queensl<strong>and</strong>), AMS C173685(6; NE <strong>of</strong> Lady Musgrave Isl<strong>and</strong>, Queensl<strong>and</strong>,23°52.5-51.9’S, 152°42.7-41.7’E, 296 m), AMSC173438 (1; E <strong>of</strong> Lady Musgrave Isl<strong>and</strong>,Queensl<strong>and</strong>, 23°52.5’-23°51.9’S, 152°42.7-152°41.7’E, 296 m), AMS C173655 (1; <strong>of</strong>f Cairns,Queensl<strong>and</strong>, 17°09.53’S, 146°42.36’E, 253 m),AMS C174639 (12; E <strong>of</strong> Lady Musgrave Isl<strong>and</strong>,Queensl<strong>and</strong>, 23°52.5’-23°51.9’S, 152°42.7-152°41.7’E, 296 m), QM MO40083 (14; <strong>of</strong>fCairns, northern Queensl<strong>and</strong>, 17°22’S, 146°48’E,303 m). Gadila cobbi Lamprell & Healy, 1998.Indo-Pacific: northern <strong>and</strong> eastern Australia. 150-1161 m.cocconii, Dentalium (Fustiaria) Sharp & Pilsbry inPilsbry & Sharp, 1897: 87. Replacement name forDentalium dispar Cocconi, 1874, a junior homonym<strong>of</strong> Dentalium dispar Sowerby, 1860. Objectivejunior synonym <strong>of</strong> Fustiaria rubescens var. exdisparSacco, 1897 (fide Pilsbry & Sharp 1898: 214).cocentum, Dentalium Hoeninghaus, 1831: 155.Tertiary. Tabbiano, Italy. Nomen nudum.† coelatulum, Dentalium Baily in Salter, 1857: 87,pl. 2, figs 8a, 8b. Cretaceous. Upper Greens<strong>and</strong>,Aberdeenshire, Scotl<strong>and</strong>, United Kingdom.† coislinense, Dentalium (Entalis) Cossmann, 1902:107 [157], pl. 10 [15], fig. 17. Eocene. Coislin <strong>and</strong>Campbon, northwestern France.colamni, see colmani.colligens, Dentalium sexangulum var. Sacco, 1897: 94.Pliocene. Italy. Nomen nudum; published in synonymy<strong>of</strong> Dentalium sexangulum var. acutangularisCocconi, 1873.collinsae, Dentalium (Dentalium) Lamprell & Healy,1998: 31, figs 20C, 21C, 24, 25C, D. <strong>Recent</strong>. Inchannel, 0.8 km south <strong>of</strong> Green Isl<strong>and</strong>, OysterHarbour via Albany, Western Australia. HolotypeWAM 230/94; paratypes WAM N3126 (10),WAM N3129 (1; in channel W <strong>of</strong> Green Isl<strong>and</strong>,Oyster Harbour via Albany, Western Australia,10 m). Dentalium collinsae Lamprell & Healy,1998. Indian Ocean: western Australia. 10 m.colliverae, Cadulus Lamprell & Healy, 1998: 163, figs167C, 168F, 171. <strong>Recent</strong>. Off Cairns, northernQueensl<strong>and</strong>, Australia, 17°22’S, 146°48’E, 296-303 m. Holotype QM MO17654. Cadulus colliveraeLamprell & Healy, 1998. Known from typelocality only.colmani, Dentalium Palmer, 1974b: 124. <strong>Recent</strong>.35 miles east <strong>of</strong> Sydney, 1463 m. Holotype (<strong>of</strong>applanatum Colman) AMS C22652; paratypesAMS C170645 (24; 35 miles E <strong>of</strong> Sydney,1463 m). Replacement name for Dentalium(Bathoxiphus) applanatum Colman, 1958, a juniorhomonym <strong>of</strong> Dentalium applanatum Torley, 1908.Rhomboxiphus colmani (Palmer, 1974) (fideScarabino 1995: 317). Indo-Pacific: Australia toNew Caledonia. 241-3658 m. Remarks: Chistikov(1983: 183) synonymized this <strong>species</strong> withBathoxiphus soyomaruae Okutani, 1964; Scarabino(1995: 314, 317) rejected Chistikov’s synonymy.Bathoxiphus colamni [Okutani, 1975: 77] is anincorrect subsequent spelling.† colobus, Cadulus Pilsbry & Sharp, 1898a: 474,pl. 11, figs 17-20. Oligocene. Dominican Republic.Type material ANSP 2886 (fide Pilsbry 1922: 401).colonense, Dentalium (Dentalium) gouldii Henderson,1920: 31, pl. 3, fig. 6. <strong>Recent</strong>. Albatross stn 2145,near Colon, Panama, 9°27’N, 79°54’W, 25 fms[46 m]. Holotype USNM 94085. Synonym <strong>of</strong>Dentalium gouldii Dall, 1889 (fide Scarabino inRios 1975: 183).colubridens, Cadulus Watson, 1879: 523, 524[Watson 1886: 18, pl. 3, fig. 1]. <strong>Recent</strong>. North EastPoint <strong>of</strong> New Zeal<strong>and</strong>, Challenger stn 169, 37°34’S,179°22’E, 700 fms [1276 m]. Holotype BMNH1887.2.9.71. Siphonodentalium colubridens(Watson, 1879) (fide Scarabino 1995: 329). Indo-Pacific: Africa to New Caledonia, Japan, NewZeal<strong>and</strong>. 630-2800 m.columbiana, Antalis Clessin 1896: 43, pl. 10, fig. 4.<strong>Recent</strong>. British Columbia [Canada]. Type materialnot located in the Naturmuseum Stuttgart. Juniorsynonym <strong>of</strong> Antalis pretiosum (Sowerby, 1860)(fide Pilsbry & Sharp 1897: 45).complexum, Dentalium, Dall, 1895: 686, 687, pl. 26,fig. 3. <strong>Recent</strong>. Off Honolulu, Hawaii, Albatross stn3472, 21°12’N, 157°49’W, 295-298 fms [538-544 m]. Syntypes USNM 107023, USNM 107022(stn 3476, 21°09’N, 157°53’W, 298 fms [544 m]);BPBM. Fissidentalium complexum (Dall, 1895)(fide Pilsbry & Sharp 1897: 76). Indo-Pacific:Hawaii. 468-544 m.compressiusculum, Dentalium Boissevain, 1906: 33,34, pl. 6, fig. 12. <strong>Recent</strong>. B<strong>and</strong>a Sea, Sibogastn 241, 04°24.3’S, 129°49.3’E, 1570 m. HolotypeZMA 3.06.044. Compressidentalium compressiusculum(Boissevain, 1906) (fide Habe & Kosuge1964: 4). Indo-Pacific: Madagascar to Indonesia<strong>and</strong> New Caledonia. 540-2570 m.compressum, Dentalium d’Orbigny, 1850. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.compressum, Dentalium Watson, 1879: 516 [Watson1886: 9, 10, pl. 1, fig. 9]. <strong>Recent</strong>. North <strong>of</strong> CulebraIsl<strong>and</strong> [Virgin Is.], West Indies, Challenger stn 24,18°38’30”N, 65°05’30”W, 390 fms [712 m].Holotype BMNH 1887.2.9.35. Junior homonym<strong>of</strong> Dentalium compressum d’Orbigny, 1850.Replacement name: Dentalium (Compressidens) pressumSharp & Pilsbry in Pilsbry & Sharp, 1897: 124.Compressidens pressum (Sharp & Pilsbry in Pilsbry& Sharp, 1897).compressum, Dentalium Meyer, 1883: 258, 259, fig. 4.Oligocene. Joachimsthal <strong>and</strong> Hermsdorf, Germany.Junior homonym <strong>of</strong> Dentalium compressum d’Orbigny,1850 (Polychaeta) <strong>and</strong> Dentalium compressumZOOSYSTEMA • 2004 • 26 (4)577


Steiner G. & Kabat A. R.Watson, 1879. Replacement name: Dentalium ottoiSharp & Pilsbry in Pilsbry & Sharp, 1897: 119;1898: 211.compressum, Dentalium Martin, 1885: 189, 190,pl. 10, fig. 190; 1887: 189, 190, pl. 10, fig. 190.Tertiary. Ngembak, Java, Indonesia. SyntypesRGM 7291 (6) (fide Van den Hoek Ostende et al.2002: 112). Junior homonym <strong>of</strong> Dentalium compressumd’Orbigny, 1850 (Polychaeta); Dentaliumcompressum Watson, 1879; Dentalium compressumMeyer, 1883. Martin (1919: 69) transferred his<strong>species</strong> to Entalina. The replacement name, Cadulusmartini Finlay, 1927, is itself a junior homonym <strong>of</strong>C. martini Whitfield, 1882. Replacement name:Dentalium javense Palmer, 1974.compressum, Dentalium Sowerby, 1889: 569, 570,pl. 28, fig. 18. <strong>Recent</strong>. Hong Kong. Junior homonym<strong>of</strong> Dentalium compressum d’Orbigny, 1850;D. compressum Watson, 1879; D. compressumMeyer, 1883; D. compressum Martin, 1885.Replacement name: Dentalium (Fissidentalium)hungerfordi Pilsbry & Sharp, 1897: 84.Compressidentalium hungerfordi (Pilsbry & Sharp,1897) (fide Habe 1963: 260).comprimatum, Dentalium (Compressidens) Plate,1908a: 349, 350, pl. 30, figs 26-34. <strong>Recent</strong>.Zanzibar Channel, Valdivia stn 245, 5°27.9’S,39°18.8’E, 463 m. Lectotype ZMB 61101a (designatedby Kilias 1995: 173); paralectotypes ZMB61101b (2). Compressidens comprimatum (Plate,1908) (fide Scarabino 1995: 370). Indian Ocean:East Africa. 238-463 m.concinnum, Dentalium von Martens, 1878: 134.<strong>Recent</strong>. West Africa, Gazelle Exp., 10°6.9’N,17°16’W, 360 fms [657 m]. Lectotype ZMB29310a (designated by Kilias 1995: 173); paralectotypeZMB 29310b. Fissidentalium concinnum (vonMartens, 1878) (fide Pilsbry & Sharp 1898: 249).Eastern Atlantic: Africa. 270-677 m. Remark: BothPilsbry & Sharp (1898: 249) <strong>and</strong> Kilias (1995: 173)erroneously gave 150 fms as the original depth.concretum, Dentalium Colman, 1958: 141, fig. 2.<strong>Recent</strong>. 27.5 miles E <strong>of</strong> Sydney, 300 fms [548 m].Holotype AMS C62231; paratypes AMS C170638(11). Striodentalium concretum (Colman, 1958)(fide Scarabino 1995: 241). Pacific Ocean: SEAustralia. 6-1000 m.† confusum, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 220. Cretaceous: Turonian. Visé,Belgium. Replacement name for Dentaliumalternans Ryckholt, 1851, a junior homonym <strong>of</strong>Dentalium alternans Chenu, 1843, <strong>and</strong> D. alternansMüller, 1851.congoensis, Dentalium Plate, 1908a: 349, pl. 30,figs 24, 25. <strong>Recent</strong>. Mouth <strong>of</strong> Congo River,Valdivia stn 71, 6°18.7’S, 12°2.1’E, 44 m. HolotypeZMB 61085. Junior synonym: Dentaliumcoarti Dautzenberg, 1912, <strong>and</strong> Dentalium coarti var.septemcostatum Dautzenberg, 1912 (fide Nicklès1979: 44). Dentalium congoensis Plate, 1908.Eastern Atlantic: Africa. 4-150 m. Remark: Plategave the latitude as “N” instead <strong>of</strong> “S”.congruens, Cadulus curtus var. Watson, 1879: 527[Watson 1886: 21, pl. 3, fig. 7a]. <strong>Recent</strong>. CulebraIsl<strong>and</strong> [Virgin Is.], St Thomas, Challenger stn 24,390 fms [712 m]. Syntype BMNH 1887.2.9.83.Cadulus congruens Watson, 1879 (fide Henderson1920: 142). Known from the type locality only.Remark: Watson described this variety from 2 specimens,but the lot in BMNH is labelled “holotype”<strong>and</strong> contains a single specimen; the second specimenhas not been found.conicum, Dentalium Hutton, 1873b: 1. Tertiary,Wanganui (Pliocene) <strong>and</strong> Ahuriri (Miocene)Formations. Shakespeare Cliff <strong>and</strong> Broken River,New Zeal<strong>and</strong>. Lectotype NZGS TM4810 (designatedby Keyes 1972: 96). Dentalium zel<strong>and</strong>icumSowerby, 1866 (fide Keyes 1972: 96); Fissidentaliumopacum (Sowerby, 1829) (fide Suter 1913:819).conicum, Dentalium Seguenza, 1876: 266. Pliocene.Italy (various localities). Junior homonym <strong>of</strong>Dentalium conicum Hutton, 1873. Remark: we prefernot to rename this junior homonym because <strong>of</strong>the insufficient description <strong>and</strong> the lack <strong>of</strong> typematerial we consider it a nomen dubium.† conradi, Cadulus (Gadila) Pilsbry & Harbison,1933: 117, pl. 4, figs 18, 19. Miocene. Shepherd’sPit, near Shiloh, New Jersey, USA. Holotype ANSP12596; paratypes ANSP 19554 (57).conradi, Dentalium Dall, 1909: 136. Miocene. Astoria,Oregon, USA. An unnecessary replacement namefor Teredo substriatum Conrad, 1849, a secondaryhomonym <strong>of</strong> Dentalium substriatum Deshayes,1825. Remarks: Conrad’s name was alreadyrenamed Dentalium pseudonyma Pilsbry & Sharp,1898; Dall’s replacement name is a junior objectivesynonym. Since Conrad’s name encompassed whatDall regarded as two <strong>species</strong>, Dall restricted thename substriatum Conrad, by describing the“smooth” form as a separate <strong>species</strong>, D. petricola.Emerson (1958: 93) concluded that petricola was avalid <strong>species</strong> <strong>of</strong> Dentalium (Rhabdus).conspicuum, Dentalium Melvill, 1897: 21, pl. 7,fig. 28. <strong>Recent</strong>. Karachi [Pakistan]. SyntypesBMNH 1897.7.30.80-81. Junior synonym <strong>of</strong>Tesseracme quadrapicalis (Hanley in Sowerby,1860) (fide Scarabino 1995: 222).† constrictum, Dentalium Newton & Harris, 1894:64, text-fig. Eocene, London Clay. Fareham <strong>and</strong>Portsmouth, Engl<strong>and</strong>, Great Britain. Junior synonyms:Entaliopsis annulata Newton & Harris,1894; Dentalium entaliopsis Pilsbry & Sharp, 1898(fide Jeffery & Tracey 1997: 90). Antalis constricta(Newton & Harris, 1894) (fide Jeffery & Tracey1997: 90, pl. 15, figs 3-6).cookei, Dentalium (Dentalium) Sharp & Pilsbry inPilsbry & Sharp, 1897: 29. <strong>Recent</strong>. Gulf <strong>of</strong> Suez.578ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Holotype (<strong>of</strong> Dentalium acus Cooke, 1885) UMZCI.100,735. Replacement name for Dentalium acusCooke, 1885, a junior homonym <strong>of</strong> Dentalium acusEichwald, 1856. Dentalium cookei Sharp & Pilsbryin Pilsbry & Sharp, 1897. Indian Ocean, Red Sea.75-148 m.† coonensis, Cadulus Stephenson, 1941: 256, pl. 46,fig. 23. Cretaceous, Ripley Formation. Coon Creek,McNairy County, Tennessee, USA. HolotypeUSNM Paleobiology 32824; 12 paratypes USNMPaleobiology 76784. Remarks: Stephenson (1941:256) also recorded this taxon from the NavarroGroup (Cretaceous) <strong>of</strong> Texas. This <strong>species</strong> wasmisidentified as Cadulus obnutus Conrad, 1869 byWade (1926).† cooperi, Dentalium Gabb, 1864: 139, pl. 21,fig. 100. Cretaceous. San Diego; NE <strong>of</strong> Martinez;near Fort Téjon; Curry’s, S <strong>of</strong> Mount Diablo; <strong>and</strong>Siskiyou Mountains, California, USA. Remark:Anderson & Hanna (1925: 143) concluded that “itis entirely probable that the type came from” RoseCanyon, near San Diego.† corallinum, Dentalium d’Orbigny, 1850 (volume 2):12. Jurassic. La Rochelle, France.cordiseturata, Entalina Habe, 1953: 399 [ex KurodaMS]. Nomen nudum.coridisecturatum, Dentalium (Entalina) Shikama,1964: 35, caption to fig. 62(6) [ex Kuroda MS].Nomen nudum. <strong>Recent</strong>. Okezoko, <strong>of</strong>f CapeAshizuri, southwestern Shikoku, Japan. Remark: asa nomen nudum in Goto & Poppe (1996: 1025).corneum, Dentalium Linnaeus, 1767. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.cornicula, Dentalium d’Orbigny, 1852. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.corniculum, Dentalium O. G. Costa, 1851. Not ascaphopod. See Other taxa described as<strong>Scaphopoda</strong>.† cornu, Dentalium de Koninck, 1877: 315, 316,pl. 23, fig. 4; 1898: 249, pl. 23, fig. 4. Carboniferous.Karúa, New South Wales, Australia.cornubovis, Dentalium E. A. Smith, 1906: 249, 250.<strong>Recent</strong>. Indian Ocean, 1154 fms [2106 m].Lectotype BMNH 1906.10.12.142-144 (designatedby Scarabino 1995: 249, “the largest (63 mm) <strong>of</strong>the 3 syntypes in BMNH”); paralectotypes ZSIM3735/1. Fissidentalium cornubovis (E. A. Smith,1906) (fide Scarabino 1995: 249). Indo-Pacific:Africa to New Caledonia. 2089-4350 m. Remark:the manuscript list <strong>of</strong> Investigator Exp. types inBMNH gives the holotype <strong>and</strong> 3 paratypes in ZSIM3735/1, <strong>and</strong> 3 paratypes BMNH 1906.10.12.142-144.cornucopiae, Entalina Boissevain, 1906: 63, 64, pl. 6,fig. 89, text-fig. 30. <strong>Recent</strong>. Indonesia, Siboga stn52, 09°03.4’S, 119°56.7’E, 959 m. Holotype ZMA3.06.086. Megaentalina cornucopiae (Boissevain,1906) (fide Habe 1963: 272). Indo-Pacific:Indonesia to Japan. 200-3250 m.† corpulentus, Cadulus Meyer, 1886: 66, pl. 3, fig. 5.Eocene. Red Bluff, Mississippi, USA. SyntypeUSNM Paleobiology 644579. Cadulus (Polyschides)corpulentus Meyer, 1886 (fide MacNeil & Dockery1984: 249).corrugatum, Dentalium Hupé in Gay, 1854: 276, pl. 2,fig. 8. Tertiary. Topocalma, Colchagua Province,Chile. Junior synonym <strong>of</strong> Dentalium subgiganteumd’Orbigny, 1852 (fide Pilsbry & Sharp 1898: 217).corrugatum, Dentalium Carpenter, 1857: 189. <strong>Recent</strong>.Mazatlan [Mexico]. Holotype BMNH 1857.6.4.881 (fide Keen 1968: 403, text-fig. 24). Juniorhomonym <strong>of</strong> Dentalium corrugatum Hupé in Gay,1854. Replacement name: Cadulus (Gadila) panamensisSharp & Pilsbry in Pilsbry & Sharp, 1898:191, which is a junior synonym <strong>of</strong> Gadila perpusilla(Sowerby in Broderip & Sowerby, 1832) (fideEmerson 1971: 78).coruscum, Dentalium (Laevidentalium) Pilsbry, 1905:117, pl. 5, figs 42, 43. <strong>Recent</strong>. Heda, Izu, Japan,167 fms [305 m]. Holotype ANSP 88320.Laevidentalium coruscum (Pilsbry, 1905) (fideHabe 1963: 268). Indo-Pacific: Indonesia to NewCaledonia <strong>and</strong> Japan. 100-1890 m.† cossmannianum, Dentalium Pilsbry & Sharp, 1898a:467, 468, pl. 10, fig. 1, pl. 11, figs 10, 11.Oligocene. Dominican Republic. Holotype ANSP2709 (fide Pilsbry 1922: 399); paratypes ANSP79475.† costae, Dentalium Pilsbry & Sharp, 1898: 202.Pliocene. Italy. Name for Dentalium lacteum sensuO. G. Costa 1851 non Deshayes, 1825. Remark:Pilsbry & Sharp erroneously attributed this name to“Deshayes, 1898”.† costaricense, Dentalium Pilsbry, 1911: 165, 166,fig. 3. Pliocene? Costa Rica. Holotype ANSP 3204;paratypes ANSP 78955 (2). Remark: as a nomennudum in Pilsbry & Sharp (1898: 254).† costatior, Dentalium (Entalis) miopseudoentalis “?”var. Sacco, 1897: 106, pl. 9, figs 12, 13 [as a nomennudum in Sacco 1896: 97]. Miocene. Turin hills,Italy. Syntypes MIGT BS.106.03.006-BS.106.03.007 (fide Ferrero Mortara et al. 1984: 306).† costatum, Dentalium J. Sowerby, 1814: 162, pl. 70,fig. 8. Pliocene. Holywell Craig, Engl<strong>and</strong>, UnitedKingdom. Remarks: Dentalium costatum – Nyst(1845: 344, pl. 25, fig. 2; 1881: 121, pl. 7, fig. 15)was subsequently described as D. dollfusi Koenen,1883; D. costatum – J. de C. Sowerby in Dixon(1850: 162, pl. 70, fig. 8) was subsequentlydescribed as D. angustum Deshayes, 1861 (fidePilsbry & Sharp 1898: 198).† costulatior, Dentalium (Antale “?”) taurocostatum var.Sacco, 1897: 101, pl. 8, fig. 55 [as a nomen nudumin Sacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.02.047 (fide FerreroMortara et al. 1984: 304).† costulatior, Dentalium michelotti var. Sacco, 1897:96, pl. 7, figs 93, 94 [as a nomen nudum in SaccoZOOSYSTEMA • 2004 • 26 (4)579


Steiner G. & Kabat A. R.1896: 97]. Miocene. Turin hills, Italy. SyntypeMIGT BS.106.01.042 (fide Ferrero Mortara et al.1984: 303).† costulatissima, Dentalium michelotti var. Sacco,1897: 96, pl. 7, figs 95, 96 [as a nomen nudum inSacco 1896: 97]. Pliocene. Castenuova d’Asti <strong>and</strong>Viale, Montafia, Italy. Syntypes MIGT BS.106.01.043, BS.106.01.044 (fide Ferrero Mortara et al.1984: 303).† crassulum, Dentalium Stoliczka, 1868: 444, pl. 27,fig. 21. Cretaceous. Trichinopoly <strong>group</strong>. South <strong>of</strong>Serdamungalum, India.crassum, Dentalium Salis Marschlins, 1793: 358 [citingGualtieri 1742: pl. 10, fig H; Martini 1769: pl. 1,fig. 4]. <strong>Recent</strong>. Tarento [Italy]. Junior synonym <strong>of</strong>Fissidentalium rectum (Gmelin, 1791) (as SalisMarschlins cited the same figures as Gmelin).crassum, Dentalium Deshayes, 1825. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† crebrestriata, Dentalium sexangulum var. Coppi,1881: 85 [original spelling crebre striata]. Pliocene.Bagalo <strong>and</strong> Niciola, Italy.crenatocinctum, Coleolus Hall, 1879. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>. Remark:considered to be a Dentalium by Whitfield (1882).† cretaceum, Dentalium Conrad, 1852: 228, App.pl. 1, fig. 1. Cretaceous. Safed, Syria. Type materialpresumed lost (not found by Beecher 1900). Juniorsynonym: Dentalium syriacum Fraas, 1867 (fidePilsbry & Sharp 1898: 221; Blanckenhorn 1927:123).† cretaustralium, Laevidentalium Stilwell, 1999: 223,224, figs 2, 3H-K. Cretaceous, Upper Albian?Coober Pedy, S[outh]. Australia. Holotype AMSF31169-1; paratypes AMS F31169-2, AMSF31169-2, JCU 11614.† cristatus, Entalis Mansuy, 1914: 52, pl. 7, fig. 17.Tertiary. Phnom Nien, near Sisophon, Cambodia.Remark: Hayasaka (1925: 24) stated that: “Thiscannot be an Entalis but is a Dentalium.”crocea, Dentalium dentale var. Dautzenberg, 1884: 302[ex Monterosato MS; nomen nudum in Monterosato1878: 76]. “Coste de Barbaria” [= North Africa].<strong>Recent</strong>. Mediterranean Sea: Gulf <strong>of</strong> Gabes, Tunisia.Synonym <strong>of</strong> Antalis dentalis (Linnaeus, 1758).Remark: Dautzenberg appears to be the first authorto have validated Monterosato’s 1878 nomen nudum.crocinum, Dentalium Dall, 1907: 169. <strong>Recent</strong>. Gulf <strong>of</strong>Tokio, Japan, USFC stn 5094, 35°04.42’N,139°38.20’E, 88 fms [162 m]. Holotype USNM110508. Calliodentalium crocinum (Dall, 1907)(fide Habe 1964: 31). Indo-Pacific: Indonesia toJapan <strong>and</strong> Australia. 27-1200 m.crosnieri, Dentalium Scarabino, 1995: 211, 212,figs 18, 28b. <strong>Recent</strong>. New Caledonia, LoyaltyIsl<strong>and</strong>s, RV Alis, MUSORSTOM 6, stn DW428,20°24’S, 166°13’E, 420 m. Holotype MNHN <strong>and</strong>7 paratypes (3 from stn MUSORSTOM 6, stn DW428; 1 from BIOGEOCAL, stn DW 292; 2 fromstn MUSORSTOM 6, stn DW 459; 1 from stnMUSORSTOM 6, stn DW 487). Dentalium crosnieriScarabino, 1995. Indo-Pacific: NewCaledonia. 420-500m.crosnieri, Fustiaria (Laevidentalium) Nicklès, 1979:60, figs 15, pl. 1, fig. III. <strong>Recent</strong>. Vridi, Côted’Ivoire, 20 m. Holotype <strong>and</strong> 4 paratypes MNHN.Fustiaria crosnieri Nicklès, 1979. Eastern Atlantic:Africa, Côte d’Ivoire. 19-20 m.† crox, Dentalium “?” Keyserling, 1847: 228, pl. 1,figs 8, 8a. Devonian. Inä [Lake Baikal region],Siberia, Russia.† crux, Dentalium (Antale) Boettger, 1907: 210.Miocene. Valea Semini, Valea Casilor, ParauUngurului, <strong>and</strong> Parau Lui, Kostej, Romania.Lectotype SMF XIII.2a (selected by Zilch 1934:279, pl. 22, fig. 29). Dentalium (Antalis) cruxBoettger, 1907 (fide Zilch 1934: 279). Remark:Hörnes (1856) misidentified this <strong>species</strong> asDentalium pseudoentalis Lamarck, 1818.cubcanaliculatum, see subcanaliculatum.† cucumis, Cadulus Koenen, 1892: 990, 991, pl. 59,figs 21, 22. Lower Oligocene. Atzendorf <strong>and</strong>Unseburg, Germany.cucurbitus, Cadulus Dall, 1881: 35. <strong>Recent</strong>. Off BahiaHonda, Cuba, Blake stn 19, 23°03’N, 83°10.30’W,310 fms [567 m]. Holotype MCZ 7750. Caduluscucurbitus Dall, 1881. Western Atlantic, Caribbean.566 m.curtum, Dentalium Sowerby, 1860: 100, pl. 225,fig. 62. <strong>Recent</strong>. Type locality not designated. Typematerial not located. Dentalium curtum Sowerby,1860. Western Atlantic, Caribbean.curtus, Cadulus Watson, 1879: 527 [Watson 1886:21, pl. 3, fig. 7]. <strong>Recent</strong>. Challenger stn 24, <strong>of</strong>fCulebra Isl<strong>and</strong> [Virgin Is.], St Thomas, 18°38.30’N, 65°05.30’W, 390 fms [712 m]. SyntypesBMNH 1994042 (5). Cadulus curtus Watson,1879. Western Atlantic, Caribbean. 300-712 m.curvotracheatum, Dentalium (Plagioglypta) Plate,1908a: 358, pl. 30, fig. 47. <strong>Recent</strong>. ZanzibarChannel, Valdivia stn 245, 5°27.9’S, 39°18.8’E,463 m. Holotype ZMB 61086. Junior synonym <strong>of</strong>Calliodentalium semitracheatum (Boissevain,1906) (fide Scarabino 1995: 273).† curvum, Siphodentalium [sic] Gardner, 1878: 63,pl. 3, figs 45-47. Cretaceous, Gaultian. Folkestone<strong>and</strong> Sevenoaks (Atherfield beds), Engl<strong>and</strong>, UnitedKingdom. Entalina curva (Gardner, 1878) (fidePilsbry & Sharp 1898: 234).† curvus, Cadulus (Gadila) Hodgkinson, 1974: 12,15, text-fig. 8z-aa; pl. 1, figs 12-14. Middle Eocene,Claibornian Stage, Cook Mountain Formation,Wheelock Member. Little Brazos Bluff, BrazosRiver Valley, Brazos County, Texas, USA.Holotype USNM Paleobiology 180437; paratypesUSNM Paleobiology 180436, PRI 29223, ANSP31501 (all from Little Brazos Bluff), USNM180438 (Stone City Bluff).580ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)cuspidatus, Cadulus (Polyschides) Nicklès, 1979: 73,fig. 30. <strong>Recent</strong>. Bay <strong>of</strong> Saloum, Senegal, 50 m.Holotype <strong>and</strong> 36 paratypes MNHN. Polyschidescuspidatus (Nicklès, 1979). Eastern Atlantic: Africa,Senegal. 50 m.cyathoides, Cadulus (Cadulus) Jaeckel, 1932: 308,fig. 5. <strong>Recent</strong>. Padang Isl<strong>and</strong>s, Sumatra, Valdiviastn 191, 0°39.2’S, 98°52.3’E, 750 m. LectotypeZMB 75361a (designated by Kilias 1995: 174);paralectotypes ZMB 76361b-75362, ZMB 75363(stn 211, Nicobar Isl<strong>and</strong>s, 7°48.8’N, 93°7.6’E,805 m). Cadulus cyathoides Jaeckel, 1932. Indo-Pacific: Africa to Philippines, Red Sea(?). 10-2050 m.† cyathus, Creseis Crist<strong>of</strong>ori & Jan, 1832: 1. Pliocene.Italy. Cadulus cyathus (Crist<strong>of</strong>ori & Jan, 1832) (fidePilsbry & Sharp 1898: 157, 158, pl. 32, figs 36-39).Remark: Di Geronimo & La Perna (1997: 417)concluded that Cadulus attenuatus Monterosato,1875 was a valid <strong>species</strong> <strong>and</strong> not a junior synonym<strong>of</strong> C. cyathus, as stated by previous authors.cylindraceum, Dentalium O. G. Costa, 1851. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.cylindratus, Cadulus Jeffreys, 1877: 158. <strong>Recent</strong>. Typelocality not specified. Syntypes USNM 169479 (1;Valorous stn 12, 56°11’N, 37°41’W, 1450 fms[2646 m]), USNM 169480 (1; Porcupine 1869stn 31, 56°15’N, 11°25’W, 1360 fms [2482 m]),USNM 169481 (1; Porcupine 1869 stn 30, 54°24’N,11°49’W, 1380 fms [2519 m]), USNM 169482(1; Porcupine 1869 stn 20, 55°11’N, 11°31’W,1443 fms [2633 m]), BMNH 85.11.5.1284-1285(Porcupine, no locality). Cadulus cylindratusJeffreys, 1877. Eastern Atlantic: Norwegian Basin toBay <strong>of</strong> Biscay <strong>and</strong> Azores; Western Atlantic:Massachusetts (USA). 1200-3000 m.cylindrica, Hensonella Elliott, 1960. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.cylindricum, Dentalium J. Sowerby, 1814. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.Remarks: Dentalium cylindricum – Roemer1836 is Laevidentalium elongatum (Goldfuss, 1841)(fide Engeser & Riedel 1992: 37); Dentalium cylindricum– Fischer de Waldheim 1843 is Dentaliumgladiolus Eichwald, 1846 (fide Pilsbry & Sharp1898: 223).cylindricum, Dentalium – Gardner 1878: 61, pl. 3,figs 21-25 non J. Sowerby, 1814. Cretaceous.Blackdown, Engl<strong>and</strong>, United Kingdom. Remarks:Gardner (1878: 61) recognized that Sowerby’s<strong>species</strong> was actually a Ditrupa (Polychaeta), butGardner then used Sowerby’s name for a scaphopod:“I have here transferred the name to the Blackdown<strong>fossil</strong>s, as it has been for a long time almost universallyapplied to them.” Replacement name: Dentaliumsubplanum Pilsbry & Sharp, 1898.† cylindricus, Cadulus R. Janssen, 1989: 88, pl. 2,fig. 16. Middle Oligocene, Rupelian, Boom ClayFormation. Kruibeke, East Fl<strong>and</strong>ers, Belgium.Holotype RGM 221002a; paratype RGM221002b.† cyrtoceratoides, Entalis de Koninck, 1883: 216,pl. 49, figs 13-15. Carboniferous. Visé, Belgium.Dentalium cyrtoceratoides (de Koninck, 1883) (fidePilsbry & Sharp 1898: 230).† cyrtoceroides, Dentalium Broili, 1907: 71, pl. 6,fig. 3. Triassic, Carnian. Seiser Alp, Tirol, Austria.dacostianum, Dentalium Chenu, 1843: pl. 6, fig. 33;1850: 3. <strong>Recent</strong>. No information on habitat <strong>and</strong>type material. Dentalium dacostianum Chenu,1843. Remark: Pilsbry & Sharp (1897: 61) consideredthis a doubtful <strong>species</strong>, although they listed itas valid.dalli, Cadulus (Polyschides) Pilsbry & Sharp, 1898:155, 156, pl. 30, figs 19-23. <strong>Recent</strong>. West coast <strong>of</strong>Patagonia [Chile] USFC stn 2783, 51°02’30”S,74°08’30”W, 122 fms [223 m], <strong>and</strong> USFC stn2784, 48°41’S, 74°24’W, 194 fms [354 m].Syntypes USNM 602253 (2, stn 2784), USNM122746 (1, stn 2784; in error as USNM 123736 inoriginal description), SBMNH 35039 (1, stn 2783),AMNH 148395 (3). Siphonodentalium dalli(Pilsbry & Sharp, 1898) (fide Steiner & Linse 2000:21-25). Southern Atlantic, Antarctica. 93-695 m.Remark: Although Pilsbry & Sharp (1898: 156)specified two type lots, they also mentioned“Magellan Strait, 369 fms [673 m]” which is notpart <strong>of</strong> the type series.dalli, Dentalium (Rhabdus) Pilsbry & Sharp, 1897:114, 115, pl. 21, fig. 46. <strong>Recent</strong>. Santa BarbaraChannel, north <strong>of</strong> Santa Rosa Isl<strong>and</strong>, USFC stn3200, 34°15’N, 120°14’30”W, 265 fms [485 m].Syntypes USNM 107696, USNM 599443, LACM1756 (1, as paratype), SBMNH 35040. Rhabdusdalli (Pilsbry & Sharp, 1897). Eastern Pacific:California to Chile. 360-1205 m. Remark: Shimek(1998: 83) synonymized this <strong>species</strong> with Rhabdusrectius. However, it remains to be determinedwhether the specimens reported from the type localitysouthwards to Chile are correctly referred to this<strong>species</strong>, which was described from the northeasternPacific.† danai, Dentalium Meyer, 1885: 462; 1886: 64,pl. 3, figs 2, 2a. Eocene. Jackson, Mississippi, USA.Syntype USNM Paleobiology 638703 (as “holotype”in Palmer & Brann 1965: 371). Fustiaria(Laevidentalium) danai (Meyer, 1885) (fide Palmer& Brann 1965: 371).† danvillense, Dentalium (Antalis) Palmer, 1947: 210-212, pl. 26, figs 23-27. Eocene, Jackson Formation.Louisiana, USA. Holotype PRI 4472; paratypesPRI 4473-4476.dartevellei, Cadulus (Dischides) Nicklès, 1979: 72,fig. 28. <strong>Recent</strong>. Malembe Beach (Zaire) [Congo].Holotype MRAC 792481. Dischides dartevellei(Nicklès, 1979). Known from the type locality only.† davasense, Dentalium (Entalis) Oppenheim, 1919:55, pl. 2, figs 4, 4b. Miocene. Kaleh-Davas, Turkey.ZOOSYSTEMA • 2004 • 26 (4)581


Steiner G. & Kabat A. R.decemcostata, Dentalium (Antale) novemcostatum var.Sacco, 1896: 97. Nomen nudum. Remark: subsequentlydescribed as Dentalium (Antale) novemcostatumvar. decemcostulata Sacco, 1897.† decemcostata, Dentalium (Entalis) taurostriata var.Sacco, 1897: 109, pl. 9, fig. 56 [as a nomen nudumin Sacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.03.031 (fide FerreroMortara et al. 1984: 308).decemcostatum, Dentalium antverpiense “forma”Goddeeris, 1978: 72. Miocene. Belgium. Nomennudum. Remark: this is also an unavailable infrasubspecificname (ICZN 1999: articles 45.5 <strong>and</strong>45.6.3).decemcostatum, Dentalium Brazier, 1877: 55. <strong>Recent</strong>.Katow, New Guinea, 8 fms [15 m]. Lectotype AMSA90 (designated by Lamprell & Healy 1998: 34);paralectotype AMS C308152. Dentalium decemcostatumBrazier, 1877. Indo-Pacific: New Guinea<strong>and</strong> Australia. 2-271 m.† decemcostatum, Dentalium Quaas, 1902: 233, 234,pl. 25, figs 19-21. Cretaceous. Overwegischichten,Ammonitenberge, Libya. Junior homonym <strong>of</strong>Dentalium decemcostatum Brazier, 1877. Remark:we prefer not to rename this junior homonym untilits <strong>species</strong> status <strong>and</strong> systematics are better documented.† decemcostulata, Dentalium (Antale) novemcostatumvar. Sacco, 1897: 103, pl. 8, fig. 65. Pliocene. Villavernia,Bordighera <strong>and</strong> Astigiana, Italy. SyntypeMIGT BS.106.02.057 (fide Ferrero Mortara et al.1984: 305).† decoratum, Dentalium Goldfuss, 1841: 3, pl. 166,fig. 9 [ex Münster MS]. Triassic. St Cassian, Tirol,Austria/Italy. Remark: also cited in Wissmann1841: 91, pl. 9, fig. 7.decurtata, Dentalium vulgare var. Monterosato, 1884:32. Nomen nudum. <strong>Recent</strong>. Mediterranean.† decussatum, Dentalium J. Sowerby, 1814: 161,pl. 70, fig. 5. Cretaceous. Sussex, Engl<strong>and</strong>, UnitedKingdom. Junior synonyms: Dentalium ellipticum J.Sowerby, 1814 (fide Gardner 1878: 58, FirstReviser according to ICZN 1999: article 24);Dentalium nutans Kner, 1852 (fide Pilsbry & Sharp1898: 221); Dentalium bicostale Ryckholt, 1851(tentatively, fide Ryckholt 1851).deforgesi, Dentalium Scarabino, 1995: 213, 214,figs 20, 28d, 71a, b. <strong>Recent</strong>. South <strong>of</strong> NewCaledonia, RV Coriolis, CHALCAL 2, stn DW 73,24°40’S, 168°38’E, 573 m. Holotype MNHN <strong>and</strong>6 paratypes (2 from CHALCAL 2, stn DW 73;4 from CHALCAL 2, stn 1146). Dentalium deforgesiScarabino, 1995. Indo-Pacific, New Caledonia.185-575 m.deforme, Dentalium Lamarck, 1818. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† defrancii, Dentalium Deshayes, 1861: 211, pl. 2,figs 14-16. Eocene. Paris Basin (Le Guépelle,Ermenonville <strong>and</strong> Montagny), France. Remark: avalid <strong>species</strong> <strong>of</strong> Dentalium (Entaliopsis) (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 85).† degrangei, Dentalium (Antale) Cossmann & Peyrot,1917: 163-165, pl. 1, figs 19-22. Miocene, Helvetian.Orthez (Le Paren), Sallespisse <strong>and</strong> Saint-Étienned’Orthe,France.† degrangei, Gadila (Loxoporus) Cossmann & Peyrot,1917: 184, pl. 2, figs 14, 15. Miocene, Helvetian.Saint-Étienne-d’Orthe, France.† dehmi, Dentalium Kuhn, 1935: 127, pl. 8, fig. 7a-c.Jurassic, Dogger. Forth, Bavaria, Germany.† dejtarense, Dentalium densitextum var. Báldi, 1973:337, pl. 50, fig. 9. Upper Oligocene. Dejtár,Hungary.† dekagonum, Dentalium Koenen, 1892: 984-986,pl. 59, fig. 2. Lower Oligocene. Lattorf <strong>and</strong>Mühlingen, Germany.delesserti, Entalis Chenu, 1859: 374, fig. 2804.Unjustified emendation <strong>of</strong> Dentalium delessertianumChenu, 1843 (fide Pilsbry & Sharp 1897:82). Remarks: Dentalium delesserti sensu de Stefani(1884: 236) from the Tertiary <strong>of</strong> Italy is Dentaliumrectum var. monterosatoi Pilsbry & Sharp, 1898.Pilsbry & Sharp (1898: 214) established the varietymonterosatoi as a replacement name for Dentaliumphilippi Monterosato, 1872 non Chenu, 1843.delessertianum, Dentalium Chenu, 1843: pl. 6, fig. 10;1850: 3. Type locality not designated (“Fossilede ?”). Type material not located. Junior synonym<strong>of</strong> Fissidentalium rectum (Gmelin, 1791), Pliocene<strong>of</strong> Italy (fide Pilsbry & Sharp 1898: 213; fideCaprotti 1979: 223). Remark: oldest available namefor Dentalium striatum Lamarck, 1818 non Born,1790 (fide Bellini 1909: 228).delicatulus, Cadulus (Gadila) Suter, 1913: 823; 1915:pl. 32, fig. 7. <strong>Recent</strong>. Milford Sound [NewZeal<strong>and</strong>], 100-120 fms [183-219 m]. HolotypeNZGS TM1220; paratype NZGS TM1221 (fideBoreham 1959: 75). Cadulus delicatulus Suter,1913. New Zeal<strong>and</strong>. 20-602 m. Remark: Scarabino(1995: 333) tentatively referred this <strong>species</strong> toSiphonodentalium.† delli, Cadulus Marwick, 1965: 47, pl. 10, figs 17,18. Late Pliocene, Nukumaruan. NZGS locality GS2818, railway tunnel, Waihua River, New Zeal<strong>and</strong>.Holotype NZGS TM3772 (locality GS 2818);paratype NZGS TM3771 (NZGS localityGS1409). Striocadulus delli (Marwick, 1965) (fideBeu & Maxwell 1990: 424).† delphinense, Dentalium Fontannes, 1880: 10; 1882:227-229, pl. 12, figs 3-5. Pliocene. Fay-d’Albon(Drôme), France. Remark: Fontannes (1882: 228)recorded this <strong>species</strong> from a number <strong>of</strong> additionallocalities in southeastern France.demersum, Dentalium Pilsbry, 1927: 142. UpperCretaceous, Ripley Formation. Dave Weeks propertyon Coon Creek, McNairy County, Tennessee, USA.Type material USNM Paleobiology 32821. Replacementname for Dentalium inornatum Wade,582ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)1926, a junior homonym <strong>of</strong> Dentalium inornatumMcCoy in Griffith, 1844. Junior synonym <strong>of</strong>Dentalium leve Stephenson, 1923 (fide Stephenson1941: 253, 254).denacta, Dentalia Perry, 1811: caption to pl. 52.<strong>Recent</strong>. “Eastern Ocean”. Nomen dubium. Noinformation on type material.† denotatum, Dentalium (Antalis) Ludbrook, 1956:3, 4, pl. 1, figs 7-9. Pliocene. Abbatoirs Bore <strong>and</strong>Dry Creek S<strong>and</strong>s, South Australia. Holotype SAMF 15140; 14 paratypes SAM. Remark: Ludbrook(1941: 101) misidentified this <strong>species</strong> as Dentaliumbifrons Tate, 1887.densatum, Dentalium Conrad, 1865: 212, pl. 20,fig. 15. Eocene. Type locality not given in originaldescription. Junior synonym <strong>of</strong> Dentalium mississippienseConrad, 1848 (fide Pilsbry & Sharp 1898:209).denseliratum, Antalis Pilsbry & Sharp, 1898: 250 [exCarpenter MS]. <strong>Recent</strong>. Eastern Pacific: San Pedro,California, USA. Holotype USNM 19463. Juniorsynonym <strong>of</strong> Antalis pretiosa (Sowerby, 1860) (fidePilsbry & Sharp 1897: 45; 1898: 250). Remarks:Pilsbry & Sharp were the first to publish this manuscriptname: “There is also a museum name afloat‘Antalis denseliratum Cpr.’ for young indianorumwith a distinct slit on the convex side. One such inU.S. Nat. Mus. from San Pedro (No. 19.463), is19 mill. long, 2.9 wide at aperture”. This nameappears to have been overlooked in the subsequentliterature. Pursuant to ICZN (1999) article 11.6,this name is considered to have been published insynonymy <strong>of</strong> Dentalium indianorum Carpenter,1864, hence not an available name.† densitexta, Dentalium haeringense var. Noszky,1940: 51, 73. Lower Oligocene, Rupelian.Budapest district, Hungary. Remark: Báldi (1973:337) treated this taxon as a full <strong>species</strong>.† densmuris, Dentalium Mayer, 1858: 79, 80, pl. 4,fig. 3 [original spelling: dens muris]. Miocene. Saint-Jean-de-Marsacq, France. Dentalium densmurisMayer, 1858 (fide Pilsbry & Sharp 1898: 202).dentale, Dentalium Conrad, 1845: 78, pl. 44, fig. 9.<strong>Recent</strong> <strong>and</strong> Tertiary. North Atlantic. Junior synonym<strong>of</strong> Dentalium attenuatum Say, 1824 (fidePilsbry & Sharp 1898: 199). Remark: Conrad(1845), in the plate caption, stated that: “Dentaliumdentale Conr. not Lin.; p. 78. It is D. attenuatumSay”.† dentalina, Ditrupa Guppy, 1873: 87, 88, pl. 1,fig. 11 (description); 1874: 444, 445, pl. 16, fig. 11(name <strong>and</strong> figure only); 1875: 42 (description).Miocene. Jamaica. Syntypes USNM Paleobiology115607 (figured specimen), USNM Paleobiology646435 (2). Described as a serpulid polychaete butin reality a scaphopod. Junior synonyms: Cadulus(Gadila) spiniformis Gardner, 1947; Cadulus hendersoniWoodring, 1925 (fide Woodring 1973:487). Gadilopsis dentalina (Guppy, 1873) (fideWoodring 1925: 206, 207, as Cadulus (Gadilopsis)dentalinus).dentalis, Dentalium Linnaeus, 1758: 785 [citing Rumphius1705: t. 41, f. 6]. <strong>Recent</strong>. Mediterranean.Syntypes LS 612, ZMUU 924, ZMUU 1011a-b.Junior synonyms: Dentalium linnaeanum Locard,1887 (145, footnote [incorrectly proposed replacementname to avoid tautonymy]); Dentalium simileBiondi Giunti, 1859 (fide Pilsbry & Sharp 1897:53). Antalis dentalis (Linnaeus, 1758) (fide Pilsbry& Sharp 1897: 53). Eastern Atlantic, Mediterranean.0-300 m.† dentaloideum, Orthoceras Phillips, 1836: 239, pl. 21,fig. 12. Carboniferous. Boll<strong>and</strong>, Yorkshire,Engl<strong>and</strong>. Dentalium? dentaloideum (Phillips, 1836)(fide Pilsbry & Sharp 1898: 230). Remark: Pilsbry& Sharp (1898: 230) stated that this was “a doubtfulDentalium. L.-G. de Koninck considers itequally likely to be a Cyrtoceras [Cephalopoda]”.† denticulatum, Dentalium Deshayes, 1850: 36,pl. 61, figs 13, 15, 16. Fossil. Locality not specified.Junior synonym: Gadus parisiensis Deshayes, 1861(fide Pilsbry & Sharp 1898: 238). Polyschides denticulatum(Deshayes, 1850) (fide Pilsbry & Sharp1898: 238). Remark: Pilsbry & Sharp (1898: 238)erroneously dated this name to “1864” <strong>and</strong> theytherefore listed it as a junior synonym <strong>of</strong> Polyschidesparisiensis (Deshayes, 1861).† denticulustigris, Cadulus Maury, 1917: 324 [160],pl. 52 [26], fig. 7 [original spelling: denticulus-tigris].Miocene. Rio Gurabo at Los Quemados, DominicanRepublic. Cadulus denticulustigris Maury, 1917.† depressicollis, Cadulus Pilsbry & Sharp, 1898a: 473,474, pl. 11, figs 25-27. Oligocene. DominicanRepublic. Type material ANSP 2884 (fide Pilsbry1922: 401).† depressum, Laevidentalium Pavia, 1991: 126-128,pl. 6, figs 1-5. Upper Miocene. Borelli, near Turin,Italy. Holotype <strong>and</strong> 12paratypes MIGT BS.D.08.1.† depressus, Cadulus Meyer, 1884: 111, text-fig. Eocene.Claiborne, Alabama, USA. Syntype USNMPaleobiology 638695.† dertornensis, Dentalium (Entalis) recta var. Sacco,1897: 110, pl. 10, figs 4, 5 [as a nomen nudum inSacco 1896: 97]. Miocene. San Agata, Italy. SyntypesMIGT BS.106.03.035, BS.106.03.036 (fideFerrero Mortara et al. 1984: 308).desaintlaurentae, Gadila Scarabino, 1995: 361, 362,figs 166, 169f. <strong>Recent</strong>. Philippines, RV CoriolisMUSORSTOM 2, stn DR33, 13°32’N, 121°07’E,130-137 m. Holotype MNHN; paratypes MNHN(8), AMS C201734, USNM 890855. Gadiladesaintlaurentae Scarabino, 1995. Indo-Pacific:Philippines to New Caledonia. 137-250 m.† deshayesi, Dentalium Risso, 1826b: 400. Tertiary. LaTrinité <strong>and</strong> Saint-Jean, France. Type material presumedlost (fide Arnaud 1978: 114). Remark:Risso’s manuscript figure <strong>of</strong> this <strong>species</strong> was publishedby Arnaud (1978: pl. 10, fig. 169).ZOOSYSTEMA • 2004 • 26 (4)583


Steiner G. & Kabat A. R.deshayesianum, Dentalium Galeotti, 1837. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.deshayesii, Dentalium Cocconi, 1874: 644, 645, pl. 6,fig. 17 [ex Guidotti MS]. Pliocene. Rivalta,Piacenza, Italy. Junior homonym <strong>of</strong> Dentaliumdeshayesi Risso, 1826. Replacement name:Dentalium guidotti Sacco, 1897.desmoulinsi, Dentalium (s.s.) Le Renard, 1994: 39.Replacement name for Dentalium affine Deshayes,1861, a junior homonym <strong>of</strong> Dentalium affineBiondi, 1859. An objective junior synonym <strong>of</strong>Dentalium xiphias Pilsbry & Sharp, 1898.diarrhox, Dentalium Watson, 1879: 511, 512[Watson 1886: 4, pl. 1, fig. 5]. <strong>Recent</strong>. 400 milesNE <strong>of</strong> New Zeal<strong>and</strong>, Challenger stn 169, 37°34’S,179°22’E, 700 fms [1260 m]. Syntype BMNH1887.2.9.13. Antalis diarrhox (Watson, 1879) (fideScarabino 1995: 236). Indo-Pacific: New Zeal<strong>and</strong>.401-1260 m. Remark: this <strong>species</strong> was describedfrom four specimens but only one is in BMNH.Scarabino (1995: 236) cited BMNH 1887.2.9.65in error.† diazicum, Dentalium Maury, 1920: 46, 47, pl. 7,fig. 1. Tertiary. Maury Station 185, near JuanaDiaz, Puerto Rico.dichelum, Siphodentalium Watson, 1879: 521[Watson 1886: 15, pl. 2, fig. 7]. <strong>Recent</strong>. LevukaIsl<strong>and</strong>, Fiji Isl<strong>and</strong>s, 12 fms [22 m]. Syntype BMNH1887.2.9.65. Dischides dichelus (Watson, 1879)(fide Pilsbry & Sharp 1898: 145; as Cadulus(Dischides) dichelus; fide Scarabino 1995: 340).Indo-Pacific: from Red Sea <strong>and</strong> Africa to FijiIsl<strong>and</strong>s. 22-1295 m. Remark: the two fragmentslisted in the original description were not located.didymum, Dentalium Watson, 1879: 517 [Watson1886: 10, pl. 1, fig. 11]. <strong>Recent</strong>. Off CulebraIsl<strong>and</strong>, Challenger stn 24, 18°38’30”N, 65°5’30”W,390 fms [712 m]. Syntypes BMNH 1887.2.9.43-44, BMNH without reg. No. (Challenger Acc.no.2284, 2 possible syntypes). Junior synonym:Dentalium (Episiphon) johnsoni Emerson, 1952 (fideScarabino 1975: 186). Episiphon didymum (Watson,1879) (fide Scarabino 1994: 308). WesternAtlantic: Caribbean to Brazil. 2-740 m.difforme, Dentalium J. de C. Sowerby in Dixon, 1850.Not a scaphopod. See Other taxa described as<strong>Scaphopoda</strong>.diffusum, Dentalium Chenu, 1843: pl. 6, figs 11, 12;1850: 3. “Fossil” [type locality not specified]. Juniorsynonym <strong>of</strong> Graptacme secta (Deshayes, 1825) (fidePilsbry & Sharp 1897: 96).† dijki, Dentalium Martin, 1885: 186, 187, pl. 10,figs 184, 185; 1887: 186, 187, pl. 10, figs 184, 185.Miocene. Fatu Lulih, Fialarang, Timor; Grissee <strong>and</strong>Ngembak, Java, Indonesia. Syntype RGM 7243(fide Van den Hoek Ostende et al. 2002: 111, whonoted that the description encompassed 15 specimens;evidently the remainder could not be located).† dilatatum, Dentalium Philippi, 1887a: 105, pl. 12,fig. 13; 1887b: 99, pl. 12, fig. 13. Cretaceous.Tumbez, Chile.† dilatatum, Pulsellum Cossmann, 1888: 11, pl. 1,fig. 21. Eocene. Near Soissons, France. Siphonodentaliumdilatatum (Cossmann, 1888) (fide Pilsbry& Sharp 1898: 234). Remark: a valid <strong>species</strong> <strong>of</strong>Pulsellum (fide Le Renard 1995: 175; Le Renard &Pacaud 1995: 86).† dingdenensis, Cadulus (Gadila) Goddeeris, 1977:160, 161, fig. 4. Miocene. Dingden-Königsmühle,Germany. Holotype KBINW 5863.† diopon, Dentalium (Antalis) Gardner, 1947: 626,pl. 57, figs 23-23a. Oligocene. Alum Bluff Group,Shoal River Formation. USGS locality 3856, 6 milesNNW <strong>of</strong> Mossyhead, Walton County, Florida,USA. Two syntypes USNM Paleobiology 498398.† diploconus, Cadulus Seguenza, 1876: 266; 1879:276. Pliocene. Messina, Italy. Remark: this <strong>species</strong>was erroneously attributed to “Seguenza, 1880” byPilsbry & Sharp (1898: 236).dipsycha, Dentalium (Dentalium) Pilsbry & Sharp,1897: 33, 34, pl. 4, figs 57-60. <strong>Recent</strong>. Type localityunknown. Holotype ANSP 45468. Junior synonym<strong>of</strong> Tesseracme quadrapicalis (Hanley inSowerby, 1860) (fide Scarabino 1995: 222).discrepans, Dentalium Risso, 1826a: 125. Fossil. NearNice, France. Nomen nudum.dispar, Dentalium Sowerby, 1860: 103, pl. 224,fig. 37. <strong>Recent</strong>. Singapore <strong>and</strong> Samar Isl<strong>and</strong>,Philippines. Syntypes BMNH 1993007 (SamarIsl<strong>and</strong>, 4 fms [7 m]), BMNH 1993008 (Singapore,6 fms [11 m]). Tesseracme dispar (Sowerby, 1860)(fide Habe & Kosuge 1964: 5). Indo-Pacific. 0-54 m.dispar, Dentalium Cocconi, 1874: 650, pl. 6, figs 20-22. Miocene. Castell’Arquato <strong>and</strong> Majatico, Italy.Junior homonym <strong>of</strong> Dentalium dispar Sowerby,1860. Replacement name: Dentalium (Pseudantalis)rubescens var. exdispar Sacco, 1897. Remark:Dentalium cocconii Sharp & Pilsbry, 1897 is anobjective junior synonym <strong>of</strong> Dentalium rubescensvar. exdispar, as it was also introduced as a replacementname later in the same year (fide Pilsbry &Sharp 1898: 214).disparile, Dentalium d’Orbigny, 1853: 202, pl. 25,figs 14-17. <strong>Recent</strong>. Martinique. Syntypes BMNH1854.10.4.465. Antalis disparile (d’Orbigny, 1853)(fide Pilsbry & Sharp 1897: 56). Western Atlantic,Caribbean to Santa Catarina, Brazil. 5-103 m.† dissimile, Dentalium Guppy, 1866: 292, pl. 17,fig. 4. Miocene (“Oligocene”), Bowden Formation.Bowden, Jamaica. Holotype BMNH (Palaeontology)64082. Tesseracme dissimilis (Guppy, 1866)(fide Woodring 1973: 483). Remark: Dentaliumponderosum Gabb, 1873 was considered a variety <strong>of</strong>Tesseracme dissimilis by Pilsbry & Sharp (1898a:470).divae, Cadulus Vélain, 1877: 128, 129, pl. 5, figs 1, 2.<strong>Recent</strong>. Isl<strong>and</strong> <strong>of</strong> St Paul [South Indian Ocean],584ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)38°S, 75°E, 90 m. Holotype MNHN. Gadila divae(Vélain, 1877) (fide Pilsbry & Sharp 1898: 188).Known from type locality only.† divisiense, Dentalium Gardner, 1878: 60, pl. 3,fig. 15. Cretaceous, Upper Greens<strong>and</strong>. Devizes,United Kingdom.† divulgatum, Dentalium (Dentalium) Jung, 1969:313, 314, pl. 13, figs 1, 2. Miocene, SpringvaleFormation. Melajo River area, Trinidad, West Indies.Holotype NMB G-12718; paratypes USNM Paleobiology(5; without catalog number). Remark: alsorecorded from numerous other stations in Trinidad.† dodecacostatum, Antalis Brunet, 1995: 54, pl. 5,fig. 9. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-083.† dolichus, Cadulus (Gadilopsis) Woodring, 1973:486, pl. 70, fig. 7. Late Eocene, Bohio(?) Formation.USGS locality 18839, east side <strong>of</strong> PalenquillaPoint, SW <strong>of</strong> Corozo Isl<strong>and</strong>, Canal Zone, Panama.Holotype USNM Paleobiology 646689.† dollfusi, Dentalium Koenen, 1883: 326, 327.Miocene. Germany <strong>and</strong> Belgium. No informationon type material. Name introduced for “Dentaliumcostatum Nyst, 1880”, a junior homonym, or a misapplication,<strong>of</strong> Dentalium costatum J. Sowerby,1814. Remarks: Pilsbry & Sharp (1898: 203) questionedthe validity <strong>of</strong> this <strong>species</strong>: “Koenen statesthat this is not the costatum <strong>of</strong> Sowerby; but it isdoubtful whether he had the real costatum <strong>of</strong> Nyst.The <strong>species</strong> dollfusi rests therefore upon Koenen’sdescription only.dollfusi, Dentalium Cossmann, 1899a: 137. <strong>Recent</strong>.Roussillon [Gulf <strong>of</strong> Lion], Mediterranean Sea.Replacement name for Dentalium alternansBucquoy, Dautzenberg & Dollfus, 1882, a juniorhomonym <strong>of</strong> Dentalium alternans Chenu, 1843. Anobjective junior synonym <strong>of</strong> Dentalium inaequicostatumDautzenberg, 1891 (fide Cossmann1900: 186). A junior homonym <strong>of</strong> Dentaliumdollfusi Koenen, 1883. Antalis inaequicostata(Dautzenberg, 1891).dominguense, Cadulus d’Orbigny, 1853: 201, pl. 25,figs 7-9. <strong>Recent</strong>. Saint-Domingue [DominicanRepublic]; Martinique; Saint-Thomas; Cuba.Syntypes BMNH 1854.10.4.466 (Saint-Domingue).Gadila dominguensis (d’Orbigny, 1853) (fidePilsbry & Sharp 1898: 191). Western Atlantic:Caribbean to Brazil. 33-110 m. Remark: doubtfulrecords from the West African coast by Nicklés(1955: 100; 1979: 64).dorsicostata, Entalina Lamprell & Healy, 1998: 138,139, figs 141B, 142C, D, 144. <strong>Recent</strong>. Off Cairns,northern Queensl<strong>and</strong>, Australia, 17°50.67’S,147°18.2’E, 703 m. Holotype QM MO17729;paratypes QM MO40069 (1), QM MO17756 (1;northern Queensl<strong>and</strong>, 17°33’S, 146°55’E), AMSC169774 (6; 80 km SW <strong>of</strong> Cape Martin, SouthAustralia, 37°50’-37°55’S, 139°15’-139°25’E,1555 m), AMS C169795 (30; 48 km south <strong>of</strong> CapeNelson, Victoria, 2000 m), AMS C169989 (3; <strong>of</strong>fWoollongong, New South Wales, 34°27-26’S,151°27’E, 1200 m), AMS C169796 (1; east <strong>of</strong>S<strong>and</strong>y Cape, Queensl<strong>and</strong>, 24°43.5-8’S, 153°33.4-33.3’E, 604 m). Entalina dorsicostata Lamprell &Healy, 1998. Indo-Pacific, Australia. 153-2000 m.doumenci, Gadila Scarabino, 1995: 360, 361, figs 165,169a. <strong>Recent</strong>. N Mozambique Channel, Banc duGeyser, BENTHEDI stn DS11, 12°16,3’S,46°42,2’E, 2300-2450 m. Holotype MNHN <strong>and</strong> 3paratypes (1 from stn DS103; 2 from BENTHEDIstn DS40). Gadila doumenci Scarabino, 1995.Indo-Pacific: Madagascar to New Caledonia. 100-2700 m.duartei, Wemersoniella Scarabino, 1986a: 5, figs 6-11,17-21. <strong>Recent</strong>. Cuenca Argentina, Vema stn V-17-80, 43°58’S, 52°09’W, 5781 m. Holotype AMNH202770; paratypes AMNH 202771, 202772(stn V-17-81, 5332 m), MCZ 293962, MHNM14760, 14761, MNHN. Wemersoniella duarteiScarabino, 1986. Western Atlantic: Puerto Rico toBrazil. 5332-5781 m.† dufresnii, Dentalium Deshayes, 1825: 361, 362,pl. 17, fig. 18. Pliocene. Marcigny, Bourgogne,France.† dujardini, Dentalium (Antale) Peyrot, 1938: 10, 11.Miocene, Helvetian. Sainte-Catherine-de-Fierbois,Pont-Levoy, Manthelan, <strong>and</strong> Louans, LigerianBasin, Touraine, France. Type material stated to bein author’s collection. Remark: Dujardin (1837)erroneously identified this <strong>species</strong> as Dentaliumpseudoantalis Lamarck, 1818.† dumasi, Dentalium Cossmann & Peyrot, 1917: 162,163, pl. 1, figs 14, 15, pl. 2, fig. 18. Miocene,Tortonian <strong>and</strong> Helvetian. Saubrigues, Saint-Je<strong>and</strong>e-Marsacq<strong>and</strong> Parleboscq (La Guir<strong>and</strong>e), France.Tentative junior synonym <strong>of</strong> Dentalium michelottiHörnes, 1856 (fide Pavia 1991: 111).† dumasi, Fustiaria “(?)” Cossmann, 1919: 141 [89],pl. 3, fig. 49. Eocene. Le Bois-Gouët, France.dumblei, Dentalium minutistriatum var. Kennedy,1895 : 97 [spelled dumbli], 114, 123, 126, 128, 130[ex Harris MS]. Eocene. Texas. Nomen nudum; notmade available by subsequent citation by Dumble(1920 : 171) <strong>and</strong> Plummer (in Sellards et al. 1933 :693) (fide Palmer 1937: 21; 1947: 215).dunkeri, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 221, 222. Cretaceous. Near Cassel,Germany. Replacement name for Dentalium rugosumDunker, 1848 non Eichwald “1846” [sic,1857]. Remark: an unnecessary replacement name,since Pilsbry & Sharp (1898: 222) erroneouslydated Eichwald’s name to 1846, not 1857.duodecemcostatum, see duodecimcostatum Brazier, 1877.duodecenaria, Dentalium Conrad, 1862b: 570 (citingEmmons 1858: 274, fig. 188). Miocene. NorthCarolina. Junior synonym <strong>of</strong> Dentalium attenuatumSay, 1824 (fide Pilsbry & Sharp 1898: 199).ZOOSYSTEMA • 2004 • 26 (4)585


Steiner G. & Kabat A. R.Remark: the specimen figured by Emmons (1858)was labeled as “Dentalium attenuatum Say,” butConrad considered this specimen to be a distinct,new <strong>species</strong>.† duodecimcostata, Dentalium (Antale) novemcostatumvar. Sacco, 1897: 103, 104, pl. 8, fig. 69 [as a nomennudum in Sacco 1896: 97]. Pliocene. Villavernia<strong>and</strong> Zinola, Italy. Syntype MIGT BS.106.02.061(fide Ferrero Mortara et al. 1984: 305).† duodecimcostatum, Dentalium d’Archiac, 1850: 443.Oligocene? Biarritz, France.duodecimcostatum, Dentalium Brazier, 1877: 56.<strong>Recent</strong>. Darnley Isl<strong>and</strong>, Torres Straits [northernAustralia], 30 fms [55 m]. Holotype AMS C170764.Junior homonym <strong>of</strong> Dentalium duodecimcostatumd’Archiac, 1850. Replacement name: Dentaliumhealyi n. nom. Dentalium duodecemcostatum[Lamprell & Healy, 1998: 61] is an incorrect subsequentspelling.† duodecimplicatum, Dentalium Repelin, 1907: 16,pl. 1, figs 12, 13. Upper Cretaceous, Saumatrian,Lower Campanian. Plan d’Aups, France.† duplex, Dentalium Defrance, 1819a: 71. Eocene.Parnes, Seine-et-Oise [Oise], France. Junior synonym:Dentalium bicarinatum Deshayes, 1825 (fideDeshayes 1864: 203). Remark: a valid <strong>species</strong> <strong>of</strong>Lobantale (fide Le Renard 1995: 175; Le Renard &Pacaud 1995: 85).duplicatum, Dentalium Blainville, 1825. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† durhami, Cadulus (Platyschides) Emerson, 1957:989, 990, pl. 126, fig. 3. Oligocene, Lincoln stage,Lincoln formation. UCMP loc. A-8723, GraysHarbor County, Washington, USA. HolotypeUCMP 37582; paratype UCMP 37583.eboracense, Siphodentalium Watson, 1879: 523[Watson 1886: 17, pl. 2, fig. 10]. <strong>Recent</strong>. TorresStrait, Cape York, Australia, 3-11 fms [5.5-20 m].Lectotype BMNH 1887.2.9.69/1 (designated byLamprell & Healy 1998: 144); paralectotypesBMNH 1887.2.9.69/2-4 (3). Pulsellum eboracense(Watson, 1879) (fide Scarabino 1995: 322). Indo-Pacific: Australia. 5-321 m.eboreum, Dentalium Conrad, 1846b: 27. <strong>Recent</strong>.Tampa Bay [west coast <strong>of</strong> Florida]. LectotypeANSP 35530 (designated by Pilsbry & Sharp 1897:89); paralectotype ANSP 320995. Junior synonyms:Dentalium leptum Bush [in Verrill], 1885b;Dentalium matara Dall, 1889 (fide Henderson1920: 66); Dentalium amaliense Henderson, 1920(fide Scarabino 1975: 185). Graptacme eborea(Conrad, 1846) (fide Pilsbry & Sharp 1897: 89).Western Atlantic: Caribbean to Brazil. 0-195 m.Remarks: original type locality was “southern coast<strong>of</strong> Florida” <strong>and</strong> the restriction to Tampa by Pilsbry& Sharp is pursuant to ICZN (1999) article 76.2.Since the Latin stem “ebur” is not used as a nameproper but in its adjective form, the ending needs tobe adjusted to the female gender <strong>of</strong> the genus name.eburneum, Dentalium Linnaeus, 1767: 1264 [no referencesgiven]. <strong>Recent</strong>. India. Type material notlocated. Junior synonyms: Dentalium annulareSowerby, 1829; Dentalium indicum Chenu, 1843(fide Pilsbry & Sharp 1897: 116); Dentalium philippinarumSowerby, 1860 (fide Scarabino 1995: 291).Laevidentalium eburneum (Linnaeus, 1767) (fideHabe & Kosuge 1964: 7). Indo-Pacific: India toPhilippines. 44-200 m. Remark: the <strong>species</strong> misidentifiedas Dentalium eburneum by G. B. Sowerby(1825: pl. 138, fig. 6) is Dentalium subeburneumd’Orbigny, 1850.ecostatum, Dentalium Kirk, 1880. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.edenensis, Fissidentalium Lamprell & Healy, 1998: 87,figs 82E, 83E, 89. <strong>Recent</strong>. Tasman Sea, east <strong>of</strong>Eden, New South Wales, Australia, 36°57.95’S,150°22’E, 616-1050 m. Holotype AMS C303583;paratypes AMS C174458 (2), AMS C174170 (3),AMS C174457 (3). Fissidentalium edenensisLamprell & Healy, 1998. Pacific Ocean: easternAustralia. 322-1050 m.edoense, Dentalium Tokunaga, 1906. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† elassum, Dentalium (Episiphon) innumerabile subsp.Woodring, 1973: 485, pl. 75, fig. 26. MiddleMiocene, Gatun Formation. USGS locality 5845,Quebrancha Hills, overlooking Gatun Lake, CanalZone, Panama. Holotype USNM Paleobiology646738.† elaticosta, Dentalium (Entalis) recta var. Sacco, 1897:110, pl. 10, fig. 6 [as a nomen nudum in Sacco1896: 97]. Miocene. San Agata, Italy. SyntypeMIGT BS.106.03.037 (fide Ferrero Mortara et al.1984: 308).elegans, Costentalina Chistikov, 1982b: 1310, 1311,pl. 1, figs 1, 2, pl. 2, figs 1-9. <strong>Recent</strong>. Indian Ocean,Vitjaz stn 6744-5 T, 12°46.7’S, 88°54.4’E, 5100-5200 m. Holotype ZIN; paratypes IOM (1 fromstn 5193, 32°48.9’S, 103°58.2’E, 5300 m; 1 fromstn 6742-11, 13°01.2’S, 91°15.5’E, 5600-5800 m;2 from stn 6742-15, 13°01.2’S, 91°15.5’E,5280 m). Costentalina elegans Chistikov, 1982.Indo-Pacific: Australia. 5100-5800 m.† elegantissimus, Cadulus Pilsbry & Sharp, 1898a:473, pl. 11, figs 28-30. Oligocene. DominicanRepublic. Syntypes ANSP 2885 (fide Pilsbry 1922:401), ANSP 79571.elegantissimus, Striocadulus (Sagamicadulus) Sakurai &Shimazu, 1963: 250, 251, text-fig. 1. <strong>Recent</strong>. OffHayama, Sagami Bay, 30-70 fms [55-128 m].Holotype <strong>and</strong> paratype NSMT. Sagamicaduluselegantissimus (Sakurai & Shimazu, 1963) (fideScarabino 1995: 334). Indo-Pacific: Japan. 55-128 m.elegantulum, Dentalium (Antalis) Penna-Neme, 1974:108, figs 1, 3, 8. <strong>Recent</strong>. Akaroa stn 183,10°41’02”S, 36°20’25”W, 100 m. HolotypeMZUSP 18699; paratypes MZUSP 18700,MZUSP 18702, MNRJ 3646. Junior synonym <strong>of</strong>586ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Antalis infracta (Odhner, 1931) (fide Scarabino1975: 184).elenae, Gadila Scarabino, 1995: 359, 360, figs 164,169d. <strong>Recent</strong>. New Caledonia, RV Vauban MU-SORSTOM 4 stn DW 156, 18°54’S, 163°19’E,525 m. Holotype MNHN; paratypes MNHN (7; 2from MUSORSTOM 4 stn DW 156; 2 from BIO-CAL stn DW 08; 1 from MUSORSTOM 4 stn DW160; 2 from MUSORSTOM 6 stn DW 410), AMSC201733 (1; MUSORSTOM 4 stn DW 160),NMNZ M268956 (1; MUSORSTOM 4 stn DW160). Gadila elenae Scarabino, 1995. Indo-Pacific:New Caledonia. 435-668 m.elephantinum, Dentalium Linnaeus, 1758: 785 [citing:Buonanni 1684: I. t. 8; Lister 1685-1692: 4.s. 2. f. 3; Rumphius 1705: t. 41. f. I; Petiver 1713:t. 16. f. 33, 1702-1711: t. 13. f. 9; Gualtieri 1742:t. 10. f. I; d’Argenville 1742: t. 7. f. H]. <strong>Recent</strong>.Indian Ocean [Ambon, Indonesia]. SyntypesZMUU 805a-b, ZMUU 1008a-b, LS 610. Juniorsynonyms: Dentalium arcuatum Gmelin, 1791;?Dentalium recurvum Deshayes, 1825; Dentaliumstriatum Born, 1778 (partim) (fide Pilsbry & Sharp1897: 2); Dentalium viridis Perry, 1811 (fide Habe1964: 6). Dentalium elephantinum Linnaeus,1758. Indo-Pacific: Red Sea to Australia. 0-40 m.Remark: placed on the Official List <strong>of</strong> Specific Namesin Zoology as the type <strong>species</strong> <strong>of</strong> Dentalium byICZN Direction 73 (1957).† elephantinus, Dentalites Schlotheim, 1820: 95.Mesozoic [?]. Sternberg <strong>and</strong> Ludwigslust (Mecklenberg),Germany. Remark: although Schlotheim(1820) compared this <strong>fossil</strong> <strong>species</strong> with the <strong>Recent</strong>Dentalium elephantinum Linnaeus, 1758, these two<strong>species</strong> are not synonyms.elephas, Cadulus (Platyschides) Henderson, 1920: 107,108, pl. 17, fig. 10. <strong>Recent</strong>. Off St. Vincent [LesserAntilles], West Indies, Blake stn 230, 13°13’20”N,61°18’45”W, 464 fms [846 m]. Holotype MCZ7741. Gadila elephas (Henderson, 1920). WesternAtlantic, Caribbean. 640-1200 m.elizabethae, Fissidentalium Lamprell & Healy, 1998:91, 92, figs 92A, 93A, 94. <strong>Recent</strong>. North <strong>of</strong>Newcastle, New South Wales, 32°49.3’S,152°49.1’E, 1075 m. Holotype AMS C174634;paratypes AMS C152019 (6). Fissidentalium elizabethaeLamprell & Healy, 1998. Known from typelocality only. Remark: V. Scarabino (in litt.5.XI.2001) suggested that the status <strong>of</strong> F. elizabethaeLamprell & Healy, 1998 vis-à-visCompressidentalium subcurvatum (E.A. Smith,1906) requires further research.elliptica, Dentalium undulatum var. Kittl, 1894: 244[263]. Triassic, St Cassian Formation. Tirol,Austria/Italy. Junior homonym <strong>of</strong> Dentalium ellipticumJ. Sowerby, 1814. Replacement name:Dentalium kittli Cossmann, 1900.ellipticum, Dentalium J. Sowerby, 1814: 161, 162,pl. 70, figs 6, 7. Cretaceous. Folkstone, Kent,Engl<strong>and</strong>, United Kingdom. Junior synonym <strong>of</strong>Dentalium decussatum J. Sowerby, 1814 (fidePilsbry & Sharp, 1898: 221). Remark: the <strong>species</strong>misidentified as Dentalium ellipticum by Reuss(1845: 41, pl. 11, fig. 20) was subsequentlydescribed as Dentalium michauxianum Ryckholt,1851 (fide Pilsbry & Sharp 1898: 224).ellipticum, Dentalium (Laevidentalium) Koenen, 1892:986, pl. 59, fig. 16. Lower Oligocene. Lattorf,Germany. Junior homonym <strong>of</strong> Dentalium ellipticumJ. Sowerby, 1814. Replacement name:Laevidentalium pseudellipticum n. nom.elongata, Cadulus monterosatoi var. Locard, 1898: 142.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Cadulus monterosatoi Locard, 1897.elongata, Dentalium ergasticum var. Locard, 1898: 106.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium ergasticum, a junior synonym <strong>of</strong>Fissidentalium capillosum (Jeffreys, 1877).elongata, Dentalium vulgare var. Monterosato, 1884:32. <strong>Recent</strong>. Mediterranean. Nomen nudum.† elongatum, Dentalium Goldfuss, 1841: 2, pl. 166,fig. 5 [ex Münster MS]. Cretaceous. Banz,Bamberg, Germany. Laevidentalium elongatum(Goldfuss, 1841) (fide Engeser & Riedel 1992: 37).Remark: Richardson (1906: 575) tentatively suggestedthat Dentalium filicauda Quenstedt, 1852might be a junior synonym <strong>of</strong> this <strong>species</strong>; Brauns(1865: 113; 1869: 191) listed these two taxa as synonyms.elongatus, Cadulus (Platyschides?) Henderson, 1920:122, 123, pl. 19, fig. 15. <strong>Recent</strong>. Off the mouth <strong>of</strong>Mississippi River, USBF stn 2378, 29°14.30’N,88°09.30’W, 68 fms [124 m]. Holotype USNM323596. Polyschides elongatus (Henderson, 1920)(fide Emerson in Turgeon 1998: 55). Known fromtype locality only.elpis, Dentalium Winckworth, 1927: 168, 169, fig. 1,pl. 14, figs 6, 7. <strong>Recent</strong>. Mannar Isl<strong>and</strong>, Ceylon [SriLanka], 3 fms [5 m]. Lectotype BMNH 1952.3.21.13(designated by Ludbrook 1954: 110); paralectotypesBMNH 1952.3.21.14, NMW. Graptacmeelpis (Winckworth, 1927) (fide Scarabino 1995:246). Indian Ocean: Maldives, Sri Lanka. 14-238 m.† emersoni, Coccodentalium Squires, 1988: 8, 9, figs 7, 8.Eocene, Juncal Formation. CSUN locality 979,northern Lockwood Valley, Ventura County, California,USA. Holotype LACM Invertebrate Paleontology7690.† emersoni, Entalina Maxwell, 1992: 187, pl. 29,figs h, i, l, o. Eocene. NZGS localities GS9480,GS9508, Waihao River, near McCulloch’s Bridge,South Canterbury, New Zeal<strong>and</strong>. Holotype NZGSTM7230; paratypes, NZGS (3).emersoni, Fustiaria (Fustiaria) Caprotti, 1979:240, 241, pl. 11, figs 2-5. Pliocene, Piacenzian.ZOOSYSTEMA • 2004 • 26 (4)587


Steiner G. & Kabat A. R.Castell’Arquato, Italy. Holotype MSNM I 5526.Junior synonym <strong>of</strong> Gadilina jani (Hörnes, 1856)(fide Pavia 1991: 128).engischistum, Dentalium Barnard, 1963b: 351, 352,fig. 30f. <strong>Recent</strong>. Cape Natal, Durban, South Africa,62 fms [112 m]. Syntypes SAFM A5463 (6),BMNH 19642510 (6). Fustiaria engischista(Barnard, 1963) (fide Scarabino 1995: 278). IndianOcean: South Africa. 85-460 m. Remark: Scarabino(1995: 278) stated that SAFM A5463 was the holotype,but there are six syntypes in that lot <strong>and</strong> hedid not designate any one <strong>of</strong> them as a lectotype.† enneagonum, Dentalium Martin, 1885: 187, 188,pl. 10, fig. 186; 1887: 187, 188, pl. 10, fig. 186.Pliocene? Samarang, Bohrlock Blakan Kebon, Java,Indonesia. Syntypes RGM 7244 (2) (fide Van denHoek Ostende et al. 2002: 111).ensiculus, Dentalium Jeffreys, 1877: 153. <strong>Recent</strong>.Type locality not specified. Syntypes BMNH1885.11.5.1344-46 (Porcupine, no locality),BMNH 1877.11.28.56 (Valorous, 1450 fms[2624 m]) [fide Warén 1980: 54; not located inBMNH], USNM 176126 (Valorous stn 12,56°11’N, 37°41’W, 1450 fms [Greenl<strong>and</strong>,2646 m]), USNM 176127 (Porcupine 1869 stn 19a[stn 19: 54°53’N, 10°56’W, Irel<strong>and</strong>, 1360 fms,2482 m]), USNM 176128 (Porcupine 1869 stn 42,49°12’N, 12°52’W, 862 fms [Greenl<strong>and</strong>, 1573 m]),USNM 176129 (Porcupine 1870 stn 17, 39°42’N,9°43’W, 1095 fms [Portugal, 1998 m]), USNM176130 (Porcupine 1870 stn 16, 39°55’N, 9°56’W,994 fms [Portugal, 1814 m]), USNM 176131(Valorous stn 16, 55°10’N, 15°58’W, 1785 fms[Greenl<strong>and</strong>, 3258 m]). Junior synonym: Dentaliumsigsbeanum Dall, 1881 (fide Dall 1889: 428).Bathoxiphus ensiculus (Jeffreys, 1877) (fide Pilsbry& Sharp 1897: 121). Western Atlantic: Greenl<strong>and</strong>to Caribbean; Eastern Atlantic: Greenl<strong>and</strong> toNamibia. 350-4680 m.ensiforme, Dentalium Chenu, 1842: pl. 6, fig. 18;1850: 3. <strong>Recent</strong>. Antilles. Type material not located.Laevidentalium(?) ensiforme (Chenu, 1842)(fide Pilsbry & Sharp 1897: 101).ensis, Dentalium H<strong>and</strong>mann 1888: 26 [ex Rolle MS].Nomen nudum. Neogene. Vöslau, Austria. Remark:although H<strong>and</strong>mann attributed this name to“Rolle,” we were unable to find this in the publications<strong>of</strong> Friedrich Rolle that were available to us.entaliopsis, Dentalium Pilsbry & Sharp, 1898: 203.Eocene, London Clay. Portsmouth, Southampton<strong>and</strong> Primrose Hill, Engl<strong>and</strong>, Great Britain.Replacement name for Entaliopsis annulata Newton& Harris, 1894, a junior secondary homonym <strong>of</strong>Dentalium annulatum Gmelin, 1791; D. annulatumCrist<strong>of</strong>ori & Jan, 1832; D. annulatum S<strong>and</strong>berger,1842; D. annulatum Meyer, 1886; Entalis annulatumTate, 1887. Junior synonym <strong>of</strong> Antalis constricta(Newton & Harris, 1894) (fide Jeffery & Tracey1997: 90, 91).entalis, Dentalium Linnaeus, 1758: 785, 786 [citing:Linnaeus 1746: 1327; Buonanni 1684: I. t. 9; Lister1685-1692: 4. s. 2. f. 3; Rumphius 1705: t. 41. f. 5;Gualtieri 1742: t. 10. f. F; d’Argenville 1742: t. 7. f.K]. <strong>Recent</strong>. Atlantic Ocean. Syntypes ZMUU1009, ZMUU 1010, LS 613. Junior synonym:Dentalium (Antalis) entale stimpsoni Henderson,1920 (replacement name for Dentalium striolatumStimpson, 1851a) (fide Pilsbry & Sharp 1897: 43).Antalis entalis (Linnaeus, 1758) (fide G. O. Sars1878: 100, 101; Pilsbry & Sharp 1897: 42). EasternAtlantic: Spitzbergen to Morocco; WesternAtlantic: Arctic Ocean to New Engl<strong>and</strong>. 5-1400 m.Remarks: placed on the Official List <strong>of</strong> SpecificNames in Zoology as the type <strong>species</strong> <strong>of</strong> Antalis byICZN Opinion 361 (1955). Dentalium entanis[Nyst & Westendorp 1839: 412] is an incorrectsubsequent spelling.entalis, Dentalium – J. Sowerby 1814: 159, 160,pl. 70, fig. 3, non Linnaeus, 1758. Tertiary.Hordwell Cliffs, Engl<strong>and</strong>, United Kingdom.Dentalium entaloides Fleming, 1825 (fide Pilsbry &Sharp 1898: 203). Unnecessary replacement name:Dentalium acuminatum Brown, 1849.† entaloides, Dentalium Fleming, 1825: 240. Pliocene.Hordwell Cliffs <strong>and</strong> Stubbington, Engl<strong>and</strong>, UnitedKingdom. Founded on Dentalium entalis sensuJ. Sowerby 1814 non Linnaeus, 1758. Objectivejunior synonym: Dentalium acuminatum Brown, 1849.entaloides, Dentalium Eudes-Deslongchamps, 1842:128, 129, pl. 7, figs 36-38. Jurassic. Moutiers,Bayeaux, France. Junior homonym <strong>of</strong> Dentaliumentaloides Fleming, 1825. Next available name:Dentalium oolithicum Piette, 1856 (fide Pilsbry &Sharp 1898: 225). Remarks: Brauns (1869: 191)concluded that this <strong>species</strong> was a junior synonym <strong>of</strong>Dentalium elongatum Goldfuss, 1841. The <strong>species</strong>Cossmann (1885) misidentified as D. entaloidesEudes-Deslongchamps, 1842 was later described asFustiaria (Laevidentalium) rigauxi Fischer & Vadet,1985.entanis, see entalis Linnaeus, 1758.† epetrion, Cadulus (Platyschides) Woodring, 1973:487, pl. 75, figs 1, 2. Middle Miocene, GatunFormation. USGS locality 22018, Payardi Isl<strong>and</strong>,Panama. Holotype USNM Paleobiology 646741;paratype USNM Paleobiology 646732.† equisetum, Dentalium (Dentalium) Olsson, 1964:203, 204, pl. 38, figs 8-8c. Pliocene. EsmeraldasFormation. Quebrada Camarones, Ecuador.Holotype USNM Paleobiology 645438; paratypesUSNM Paleobiology 645439 (1), 645440 (1),645441 (1).erectum, Dentalium Sowerby, 1860: 99, pl. 225, fig.55. <strong>Recent</strong>. Sydney Harbor, Australia. LectotypeBMNH 1993130/1 (designated by Lamprell &Healy 1998: 109); paralectotypes BMNH1993130/2-3 (2). Junior synonym: Dentalium pluteumColman, 1958 (fide Lamprell & Healy 1998:588ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)109). Laevidentalium erectum (Sowerby, 1860)(fide Pilsbry & Sharp 1897: 111). Indo-Pacific:Australia. 11-2569 m.erectum, Dentalium “Verkrz.” Paetel, 1888: 593.“Jam.” [= Jamaica]. Nomen nudum. Remark: Pilsbry& Sharp (1898: 253) stated that this <strong>species</strong> “isunknown to us”.ergasticum, Dentalium Fischer, 1883: 275, 276.<strong>Recent</strong>. Atlantic Ocean, 1900 m. SyntypesMNHN; MCZ 133729 (1; Talisman, <strong>of</strong>f Fayal,Azores, 38°33’25”N, 30°28’54”W). Junior synonym<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877)(B. Métivier, n. syn.).† erleneae, Cadulus (Gadila) Hodgkinson, 1974: 15,16, text-fig. 8a, b; pl. 1, figs 1, 2, pl. 2, fig. 5.Middle Eocene, Claibornian Stage, Cook MountainFormation, Wheelock Member. Little Brazos River,Brazos County, Texas, USA. Holotype USNMPaleobiology 180425; paratypes USNM Paleobiology180426, 180446; PRI 29224; ANSP 31502.erosum, Fissidentalium Shimek & Moreno, 1996: 77-81, figs 2, 4, 5. <strong>Recent</strong>. Station P3 (Moss L<strong>and</strong>ingMarine Laboratory Designation), Pioneer Canyon,<strong>of</strong>f Central California, 37°03’N, 123°26’W, 3090-3300 m. Holotype LACM 2755; paratypes LACM2756, USNM 880041, BMNH 1994054. Fissidentaliumerosum Shimek & Moreno, 1996. EasternPacific: California. 3090-3300 m.“esinense”, Dentalium – Kittl 1899: 4. Error forDentalium lombardicum Kittl, 1899. Remark: Kittl(1899: 191) corrected this error in the description<strong>of</strong> Dentalium lombardicum but erred again spellingit “longobardicum”. Kittl used the correct originalspelling on pages 197 <strong>and</strong> 213.† esmeraldum, Dentalium (Fissidentalium) Olsson,1942: 232, 233 [80, 81], pl. 6, figs 1, 2, 10.Pliocene, Esmeraldas Formation. Punta Gorda,Esmeraldas province, Ecuador. Holotype PRI 4069;paratypes PRI 4070 (2) (fide Brann & Kent 1960:318).† etalense, Dentalium Terquem & Piette, 1865: 67,pl. 2, fig. 43. Jurassic, Liassic. Saint-Menge, France.Laevidentalium etalense (Terquem & Piette, 1865)(fide Engeser & Riedel 1992: 37). Remark:Woodward (1893: 352) listed Dentalium portlockiTate, 1870 as a junior synonym <strong>of</strong> this <strong>species</strong>, butwithout any explanation.eualdes, Dentalium Barnard, 1963a: 444, 445. <strong>Recent</strong>.South Africa, 33°36’S, 16°15’E, 1520-1570 fms[2774-2865 m]. Syntypes SAFM A9736 (2),BMNH 1964241. Fissidentalium eualdes (Barnard,1963) (fide Scarabino 1995: 255). Indian Ocean,South Africa. 2780-2880 m.† eugenii, Dentalium Dall, 1892: 438. Eocene. PrairieCreek beds, Wilcox County, Alabama, USA.Lectotype USNM Paleobiology 112631 (designatedby Palmer & Brann 1965: 367, as “holotype”).euloides, Cadulus Melvill & St<strong>and</strong>en, 1901: 459,pl. 24, fig. 24. <strong>Recent</strong>. Gulf <strong>of</strong> Oman, Karachi,24°49’N, 58°56’E, 345 fms [630 m]. LectotypeBMNH 1901.12.9.12 (designated by Ludbrook1954: 115); paralectotypes ANSP 164737 (1);ZMA 3.01.011 (2 possible paralectotypes). Caduluseuloides Melvill & St<strong>and</strong>en, 1901. Indian Ocean:Red Sea to India <strong>and</strong> Maldives. 183-630 m.eupatrides, Dentalium Melvill & St<strong>and</strong>en, 1907: 142,pl. 1, fig. 12. <strong>Recent</strong>. Antarctica, East <strong>of</strong> WeddellSea, N. <strong>of</strong> Queen Maud L<strong>and</strong>, 71°22’S, 16°34’W,1410 fms [2573 m]. Syntypes RSM 1921.143.584(figured syntype <strong>and</strong> 32 syntypes), NMW1955.158.590-91 (5), BMNH 1908.6.30.3 (1).Dentalium eupatrides Melvill & St<strong>and</strong>en, 1907.Antarctica, South Pacific. 2538-2700 m. Remarks:C. P. Palmer in Oliver (1984) assigned this <strong>species</strong>to Graptacme; V. Scarabino (in litt. 5.XI.2001) suggestedthat this <strong>species</strong> is referable to Fissidentalium.The type locality was “accidentally omitted” fromthe original description <strong>and</strong> was subsequently providedby Melvill & St<strong>and</strong>en (1907: 358).euzkadii, Annulipulsellum Scarabino, 1986b: 4, 5,figs 8-20. <strong>Recent</strong>. North Atlantic, R/V KnorrCruise 25, stn 307, 12°34.4’-12°40.8’N, 58°59.3’-59°09.2’W, 3835-3862 m. Holotype MCZ293952; paratypes MCZ 293954, MCZ 293953(Atlantis II cruise 31, stn 155, 00°03.0’S, 27°48.0’W,3730-3783 m), MCZ 293955 (Chain cruise 106,stn 326, 50°04.9’N, 14°23.8’W, 3859 m), MCZ293956 (Knorr cruise 25, stn 306, 09°31.1’N,56°20.6’W, 3392-3429 m), MNHN, MHNM14770-14772, AMNH 202773 (1; fide Boyko &Sage 1986: 29). Annulipulsellum euzkadiiScarabino, 1986. Eastern <strong>and</strong> Central Atlantic:Icel<strong>and</strong> to <strong>of</strong>f Brazil. 1407-3862 m.† ewekoroense, Dentalium (Antalis) Adegoke, 1977:64, pl. 6, figs 10, 11. Paleocene. Nigeria. HolotypeUIMG 182; paratype USNM Paleobiology174767.exaratum, Dentalium Risso, 1826a: 125. Fossil. NearNice, France. Nomen nudum.† exaratum, Dentalium Yokoyama, 1927b: 198,pl. 51, fig. 14. Pliocene. Upper Chikbets Beds(branch stream <strong>of</strong> the Haboro) <strong>and</strong> Lower ChikbetsBeds (Rorenai), Haboro, Teshio, Japan. Syntype,Geological Survey <strong>of</strong> Japan (fide Hanzawa et al.1961: 209).exasperatum, Dentalium Sowerby, 1903: 225, pl. 5,fig. 12. <strong>Recent</strong>. South Africa, Umphloti Rivermouth, 27 fms [49 m]. Syntypes BMNH1903.7.27.57-59. Junior synonym: Dentaliumplatei Jaeckel, 1932 (fide Barnard 1963a: 347).Fissidentalium exasperatum (Sowerby, 1903) (fideScarabino 1995: 255). Eastern Atlantic, IndianOcean: South Africa. 49-180 m.† exdispar, Dentalium (Pseudantalis) rubescens var.Sacco, 1897: 112. Miocene. Castell’Arquato <strong>and</strong>Majatico, Italy. Replacement name for Dentaliumdispar Mayer, 1874 non Sowerby, 1860. Remark:Dentalium cocconii Sharp & Pilsbry, 1897 is anZOOSYSTEMA • 2004 • 26 (4)589


Steiner G. & Kabat A. R.objective junior synonym as it was also introducedas a replacement name later in the same year (fidePilsbry & Sharp 1898: 214).† exiguum, Dentalium Koenen, 1892: 980, 981,pl. 59, figs 9-11. Lower Oligocene. Lattorf,Germany.exiguus, Cadulus Watson, 1879: 528, 529 [Watson1886: 23, pl. 3, fig. 10]. <strong>Recent</strong>. Off CulebraIsl<strong>and</strong>, Challenger stn 24, 18°38’30”N, 65°5’30”W,390 fms [712 m]. Syntypes BMNH 1887.2.9.94 (2).Cadulus exiguus Watson, 1879. Known from typelocality only.† exile, Dentalium Tommasi, 1901: 57, pl. 1, fig. 10.Middle Triassic, Ladinian (fide Diener 1926: 229).Canal di Pézo, Valle del Dezzo, Italy.† exlamarcki, Dentalium (Entalis) Sacco, 1897: 106.Miocene. Bordeaux <strong>and</strong> Dax, France. Replacementname for Dentalium lamarcki Mayer, 1864, a juniorhomonym <strong>of</strong> Dentalium lamarcki Chenu, 1843.exmouthensis, Dentalium (Dentalium) Lamprell &Healy, 1998: 19, 20, figs 9B, 10B, 12, 13D-F.<strong>Recent</strong>. Near lighthouse, North West Cape,Exmouth Gulf, northern Western Australia, in littorals<strong>and</strong>. Holotype QM MO38994; paratypesQM MO38985 (1), AMS C172174 (1), WAM282-93 (2). Dentalium exmouthensis Lamprell &Healy, 1998. Indian Ocean: Western Australia.0-82 m.† expolita, Entalina Della Bella & Tabanelli, 1996:14-16, text-figs A, B. Lower Pleistocene, Santerniano.Samoggia, near Monte Mario, Romagna,Italy. Holotype <strong>and</strong> paratypes, Laboratorio diMalacologia dell’Istituto di Zoologia dell’Universitàdi Bologna, No. 11631.exuberans, Dentalium Locard, 1897b: 10 [Locard,1898: 110, pl. 6, figs 15-20]. <strong>Recent</strong>. Senegal,“Sahara” [Morocco], Azores between Pico <strong>and</strong> SaintGeorges, 1258-3650 m. Syntypes MNHN (1 fromAzores; 3 from Azores, Talisman 1883 dr. 126,4255 m; 2 from Talisman 1883 dr. 102, 3655 m).Junior synonym <strong>of</strong> Fissidentalium paucicostatum(Watson, 1879) (B. Métivier, n. syn.).exvitreum, Siphonodentalium Sacco, 1897: 115, footnote.Replacement name for Dentalium vitreumM. Sars, 1851 [1831 in error] non Dentaliumvitreum Gmelin, 1791. Junior synonym <strong>of</strong>Siphonodentalium lobatum (Sowerby, 1860) (fidePilsbry & Sharp 1898: 253). Remark: Sacco consideredS. vitreum (M. Sars, 1851) <strong>and</strong> S. lobatum(Sowerby, 1860) to be different <strong>species</strong>.† ezoense, Dentalium cooperi var. Nagao, 1938: 140,141, pl. 15 [2], figs 12-14. Cretaceous. Abesinaigawa,Tezio province, Japan; Umpei-sawa, Uryugawa,Rutaka-gun, Sakhalin [Russia]. Syntype,Dept. Geol., Hokkaido Univ. No. 8248 (fideHanzawa et al. 1961: 209).fabrizioi, Bathycadulus, Scarabino, 1995: 354, 355,figs 159, 160 m, n. <strong>Recent</strong>. South Africa, MeiringNaude stn SM 109, 28°41’S, 32°37’E, 1300 m.Holotype SAFM A36258; paratypes MHNM14751 (1), SAM (1), MNHN (11; MD 32 Réunionstn DS109), NM (1; MD 32 Réunion stn DS109),USNM 890862 (1; stn DS109). Bathycadulusfabrizioi Scarabino, 1995. Indo-Pacific: SouthAfrica to New Caledonia. 1050-1915 m.falcatum, Dentalium Conrad, 1869. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.falkl<strong>and</strong>icum, Siphonodentalium (Pulsellum) Dell,1964: 131, fig. 4, No. 13. <strong>Recent</strong>. Falkl<strong>and</strong> Isl<strong>and</strong>s,Scoresby stn WS 210, 50°17’S, 60°06’W , 161 m.Holotype BMNH 1962878/1; paratypes BMNH1962878/2-6, BMNH 1962879/6 (stn 211,50°17’S, 60°06’W, 174 m), BMNH 1962879/4(stn 212, 49°22’S, 60°10’W, 242-249 m), BMNH1962881/6 (stn 214, 48°25’S, 60°40’W, 208-219m), BMNH 1962882/1 (stn 229, 50°35’S,57°20’W, 210-271 m). Pulsellum falkl<strong>and</strong>icum(Dell, 1964). Western Atlantic: Uruguay toMagellanic Region, Falkl<strong>and</strong> Isl<strong>and</strong>s. 81-1274 m.fasciatum, Dentalium Gmelin, 1791: 3738 [citingMartini 1769: I. t. I. f. 3. B]. <strong>Recent</strong>. Sicily [MediterraneanSea]. Type material not located. Juniorsynonym <strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fidePilsbry & Sharp 1897: 42).fausta, Polyschides Kuroda, Habe & Oyama, 1971:496 [in Japanese], 313, 314 [in English], pl. 65,figs 18, 19. <strong>Recent</strong>. Sagami Bay, Amadaiba-Kannontsukadashi-Maruyamadashi, Japan, 86 m(as a nomen nudum in Habe 1963: 276). Holotype<strong>and</strong> paratype in Imperial Household BiologicalLaboratory (Japan). Polyschides fausta Kuroda,Habe & Oyama, 1971 (fide Scarabino 1995: 339).Indo-Pacific: Japan. 50-100 m.† feruglioi, Laevidentalium Brunet, 1995: 55, pl. 5,fig. 11. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-060.festivum, Dentalium Sowerby, 1914: 8. <strong>Recent</strong>. NewCaledonia. Holotype BMNH 1914.4.2.1. Pictodentaliumfestivum (Sowerby, 1914) (fide Habe1964: 15, as Dentalium (Pictodentalium) formosum).Indo-Pacific: New Caledonia. 20-62 m. Remark:Dentalium festivum was synonymized with P. formosum(Adams & Reeve, 1850) by Habe (1963: 255)<strong>and</strong> Lamprell & Healy (1998: 72); however,Scarabino (1995: 269) treated this as a valid <strong>species</strong>.† fibula, Gadila Boettger, 1907: 212. Miocene. ValeaSemini, Kostej, Banat, Romania. Holotype SMFXIII.6a (fide Zilch 1934: 278, pl. 22, fig. 24).Cadulus (Gadila) fibula (Boettger, 1907) (fide Zilch1934: 278).† filicauda, Dentalium Quenstedt, 1852: 443, pl. 35,fig. 18; 1867: 530. Jurassic, Liassic. Near Berlin,Germany. Remarks: Richardson (1906: 575) tentativelysuggested that this <strong>species</strong> might be a juniorsynonym <strong>of</strong> Dentalium elongatum Goldfuss, 1841;Brauns (1865: 113; 1869: 191) listed these two taxaas synonyms.590ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)filosa, Entalis de Koninck, 1883: 219, pl. 49, figs 23,24. Carboniferous. Visé, Belgium. Junior secondaryhomonym <strong>of</strong> Dentalium filosum Broderip &Sowerby, 1830. Replacement name: Dentaliumorthoceras Pilsbry & Sharp, 1898.filosum, Dentalium Broderip & Sowerby, 1830: 48.<strong>Recent</strong>. Tenasserim [Burma/Myanmar], on shore.Syntypes (3) not located. Dentalium filosumBroderip & Sowerby, 1830. Reported from typelocality only.filum, Dentalium Sowerby, 1860: 98, pl. 225, fig. 45.<strong>Recent</strong>. Gibraltar; “Philippines”. Type material notlocated. Junior synonyms: Dentalium gracileJeffreys, 1870 (fide Jeffreys 1883: 661), Dentaliumfuniculus Brugnone, 1878 (fide Pilsbry & Sharp1898: 204, 253), Dentalium (Antalis) laeve Hilgard& Hopkins, 1878 (fide Pilsbry & Sharp 1898: 207).Episiphon filum (Sowerby, 1860) (fide Pilsbry &Sharp 1897: 118). Eastern Atlantic, Mediterranean.20-4784 m.filum, Dentalium (Laevidentalium) Marwick, 1931:158, pl. 18, figs 342, 343. Miocene, Altonian-Clifdenian, Ihungia Series. Waihora River,Waingaromia, Gisborne, New Zeal<strong>and</strong>. HolotypeNZGS TM4813; paratypes NZGS TM4814 (1),NZGS (5) (fide Keyes 1972: 97). Junior homonym<strong>of</strong> Dentalium filum Sowerby, 1860. Replacementname: Dentalium waihoraensis Emerson, 1954.Laevidentalium waihoraense (Emerson, 1954) (fideMaxwell 1988: 716).fisheri, Dentalium (Dentalium) Pilsbry & Sharp, 1897:36, 37, pl. 5, figs 61-65 [ex Stearns, 1894: 157,nomen nudum). <strong>Recent</strong>. Los Animas Bay, BajaCalifornia, Mexico. Holotype USNM 46204. Juniorsynonym <strong>of</strong> Tesseracme quadrangularis (Sowerbyin Broderip & Sowerby, 1832) (fide Emerson 1956: 2).† fissura, Dentalium Lamarck, 1818: 346. Eocene.Grignon, France. Junior synonym: Dentaliumacuminatum Deshayes, 1825 (fide Deshayes 1864:213). Fustiaria fissura (Lamarck, 1818) (fideNewton & Harris 1894: 65). Remarks: a valid<strong>species</strong> <strong>of</strong> Pseudantalis (fide Le Renard 1995: 175;Le Renard & Pacaud 1995: 85). For Dentalium fissuraauct. non Lamarck, 1818 reported from theUpper Oligocene, Janssen (1978a) introducedDentalium (Antalis) pseud<strong>of</strong>issura.fistula, Dentalium Sowerby, 1860: 99, pl. 225, fig. 62.<strong>Recent</strong>. Cochin, southwest India (label <strong>of</strong> BMNHtype). Syntype (possible holotype) BMNH1907.12.30.317. Episiphon fistula (Sowerby, 1860)(fide Pilsbry & Sharp 1897: 118). Indian Ocean:India. Remark: V. Scarabino (in litt. 20.II.2002)suggested that this <strong>species</strong> is not referable toEpisiphon, but its generic position is uncertain.flavum, Dentalium (Antalis) ceratum Henderson,1920: 51, 52, pl. 7, fig. 1. <strong>Recent</strong>. Off Key West,Eolis stn 333, 110 fms [201 m]. Holotype USNM314391; paratypes ANSP 132113 (2), USNM314400 (Eolis stn 325, 95 fms [173 m], <strong>of</strong>f S<strong>and</strong>Key, Florida), AMNH 148298 (1, stn 325),USNM 314391a-314405, USNM 323016.Sub<strong>species</strong> <strong>of</strong> Antalis cerata (Henderson, 1920).Western Atlantic: Florida Key region. 173-201 m.flavum, Dentalium Scarabino, 1995: 212, 213, figs 19,28c, e. <strong>Recent</strong>. South Pacific, Coral Sea, BancNova, RV Coriolis, MUSORSTOM 5, stn DW301, 22°07’S, 159°25’E, 478-610 m. HolotypeMNHN; paratypes MNHN (10; 1 from stn 301;1 from stn DW 306; 1 from stn LAGON 444;2 from stn DW 72; 1 from stn DW 38; 1 from stnDW 01; 1 from stn DW 05; 1 from stn DW 478;1 from stn DW 479), AMS C201722 (1; stnDW 479); NMNZ M268960 (1; stn DW 479).Junior homonym <strong>of</strong> Dentalium flavum Henderson,1920. Replacement name: Dentalium scarabinoin. nom.† flechensis, Antalis Brunet, 1995: 53, 54, pl. 5, fig. 8.Upper Miocene, Entrerriense Formation. BalizaPunta Flecha, Puerto Madryn, Chubut Province,Argentina. Holotype MPEF-PI-081.flindersi, Dentalium (Paradentalium) Cotton &Ludbrook, 1938: 210, pl. 12, fig. 4. <strong>Recent</strong>. StVincent Gulf, South Australia, 22 fms [40 m].Holotype SAM D 13338. Paradentalium flindersi(Cotton & Ludbrook, 1938). Indo-Pacific: West<strong>and</strong> South Australia. 18-276 m.† floratum, Dentalium Zimmermann, 1847: 243 [exPhilippi MS]. Upper Miocene. Schwarzen Thon,Saxony, Germany. Fissidentalium floratum(Zimmermann, 1847) (fide Janssen 1989: 84).florenciae, Bathycadulus Palmer, 1986: 396 [exScarabino MS]. Nomen nudum.florenciae, Cadulus Scarabino, 1995: 351, figs 156,160h. <strong>Recent</strong>. Réunion, Indian Ocean, MD32,stn DS139, 20°47’S, 55°38’E, 1575-1600 m.Holotype MNHN <strong>and</strong> 3 paratypes (from stnDR 104). Cadulus florenciae Scarabino, 1995.Known from Réunion only.florenciae, Dentalium Moraes Rego, 1936. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† floridanus, Cadulus Dall, 1892: 446, pl. 23, fig. 26.Miocene. Alum Bluff, Appalachicola River, Florida,USA. Syntypes USNM Paleobiology 112757(numerous). Remark: Schuchert et al. (1905: 112)cited USNM 112757 as the “holotype” but did notspecify which syntype was to be the holotype, sothis does not constitute a valid lectotype designation.floridense, Dentalium (Fissidentalium) Henderson,1920: 64, 65, pl. 10, figs 1, 2, 6, 7. <strong>Recent</strong>. OffS<strong>and</strong> Key, Florida, Eolis stn 301, 95 fms [173 m].Holotype USNM 314457. Junior synonym <strong>of</strong>Fissidentalium carduus (Dall, 1889) (fide Scarabino1994: 307; generic placement fide Emerson inTurgeon 1988: 50, <strong>and</strong> 1998: 54, 200, where helisted Fissidentalium floridense as a distinct <strong>species</strong>).† fodinense, Laevidentalium (s.l.) Maxwell, 1988: 76,pl. 13, figs f-h. Late Miocene, Stillwater Mudstone.ZOOSYSTEMA • 2004 • 26 (4)591


Steiner G. & Kabat A. R.NZGS localities GS3341 <strong>and</strong> GS11556, Karoro,near Greymouth, Westl<strong>and</strong>, New Zeal<strong>and</strong>.Holotype NZGS TM6613; paratypes NZGS (61).formosum, Dentalium Adams & Reeve, 1850: 71,pl. 5, fig. 1a, b. <strong>Recent</strong>. Sulu Archipelago, Philippines,outside coral reef near city <strong>of</strong> Sulu [= Jolo],16-20 fms [29-37 m]. Lectotype BMNH 1951.2.14.1(designated by Lamprell & Healy 1998: 72); paralectotypesBMNH 1951.2.14.2-3 (2). Synonyms:Fissidentalium (Pictodentalium) formosum harrisoniHabe, 1970, Dentalium (Pictodentalium) formosumhirasei Kira, 1959 (fide Scarabino 1995: 267).Pictodentalium formosum (Adams & Reeve, 1850)(fide Habe 1964: 15). Indo-Pacific: Australia toChina Sea <strong>and</strong> New Caledonia. 5-145 m. Remark:P. festivum (Sowerby, 1914) was considered a juniorsynonym by Habe (1963: 255) <strong>and</strong> Lamprell &Healy (1998: 72) but not by Scarabino (1995: 269).† <strong>fossil</strong>e, Dentalium Gmelin, 1791: 3738 [citingSchröter 1784b: 4 t. 3 f. 7]. [Fossil]. Type localitynot specified. Junior synonym: Dentalium catenulatumChenu, 1843 (fide Pilsbry & Sharp 1898: 201).Remark: the <strong>species</strong> misidentified as D. <strong>fossil</strong>e byPhilippi (1847: 80) was later described as D. kickxiiNyst, 1843 (fide Pilsbry & Sharp 1898: 207).foweyensis, Cadulus (Platyschides) Henderson, 1920:114, 115, pl. 18, figs 7, 13. <strong>Recent</strong>. Off TriumphReef, Florida, Eolis stn 350, 90 fms [164 m].Holotype USNM 314721. Polyschides foweyensis(Henderson, 1920) n. comb. Western Atlantic,Caribbean. 10-164 m.fragile, Pharetrium König, 1825. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.fragile, Pulsellum Scarabino, 1995: 320, 321, figs 122,125a. <strong>Recent</strong>. Réunion, Indian Ocean, MD32 stnDS151, 20°51’S, 56°03’E, 3300-3240 m. HolotypeMNHN; paratypes MNHN (14); NM (1).Pulsellum fragile Scarabino, 1995. Indian Ocean:Réunion. 3240-3300 m.† fragilis, Dentalium Meek & Hayden, 1856: 69.Cretaceous. Yellowstone River, “150 miles abovemouth”, Montana, USA. Syntypes USNM Paleobiology296 (fide Schuchert et al. 1905: 219),ANSP 79585. Dentalium fragile Meek & Hayden,1856 (fide Pilsbry & Sharp 1898: 222).francisense, Dentalium Verco, 1911a: 207, 208, pl. 26,figs 1, 1a. <strong>Recent</strong>. Petrel Bay, St Francis Isl<strong>and</strong>,South Australia, 15-20 fms [27-37 m]. HolotypeSAM D 13724; paratypes SAM D 16004 (1), AMSC334 (2; Gulf <strong>of</strong> St Vincent, South Australia, 27-40 m), USNM 251989 (Gulf <strong>of</strong> St Vincent, SouthAustralia). Paradentalium francisense (Verco,1911) (fide Cotton & Godfrey 1933: 139). Indo-Pacific: South Australia. 27-550 m. Remark:Lamprell & Healy (1998) did not recognize thegenus Paradentalium <strong>and</strong>, thus, listed francisenseunder Dentalium.franklinae, Fissidentalium Lamprell & Healy, 1998:93, figs 91F, G, 92C, 93C, 96. <strong>Recent</strong>. Off Cairns,northern Queensl<strong>and</strong>, Australia, 17°51’S,147°18’E, 703 m. Holotype QM MO40030;paratypes QM MO17771 (3), QM MO18003(1; <strong>of</strong>f Cairns, 17°49’S, 148°40’E, northernQueensl<strong>and</strong>, 990 m), QM MO18004 (3; <strong>of</strong>fCairns, northern Queensl<strong>and</strong>, 672 m), AMSC201735 (2; <strong>of</strong>f Cairns, Queensl<strong>and</strong>, 17°36.98’S,146°57.43’E, 672-744 m), AMS C201735 (2, preserved).Fissidentalium franklinae Lamprell &Healy, 1998. Indo-Pacific: eastern Australia. 366-1460 m.fraseri, Cadulus (Gadila) Nicklès, 1955: 98, fig. 1.<strong>Recent</strong>. Ghana, Atlantide stn 84, 5°37’N, 0°35’E,150-175 m. Holotype + 3 fragments not located inZMUC. Gadila fraseri Nicklès, 1955. Known fromthe type locality only.fredericae, Prodentalium Engeser & Riedel, 1992: 41.Carboniferous. Iowa, USA. Objective junior synonym<strong>of</strong> Prodentalium obsoletum (Hall, 1858).Remarks: the name was introduced as a replacementname for Dentalium sublaeve Hall in Miller, 1877, ajunior homonym <strong>of</strong> Dentalium dentalis var. sublaevisCocconi, 1874, which itself is an unnecessaryreplacement name for Dentalium obsoletum Hall,1858. Dentalium obsoletum Schlotheim, 1832 is anomen nudum <strong>and</strong> Hall’s name, therefore, not ajunior homonym.† fritschi, Dentalium Koenen, 1892: 988, 989, pl. 59,fig. 13. Lower Oligocene. Lattorf, Calbe, Atzendorf,Unseburg <strong>and</strong> Br<strong>and</strong>horst, Germany.† frontierensis, Cadulus Sidwell, 1932: 316, pl. 48,fig. 13. Cretaceous, Colorado Group, FrontierFormation. 3 miles NE <strong>of</strong> Medicine Bow, ComoRidge, Wyoming, USA. Holotype University <strong>of</strong>Iowa, Paleontological Museum, No. 8-629.funiculus, Dentalium Brugnone, 1878: 44, pl. 1, fig. 5.Pliocene. Ficarazzi, Italy. Junior synonym <strong>of</strong>Dentalium filum Sowerby, 1860 (fide Pilsbry &Sharp 1898: 204, 253).† furcata, Plagioglypta “?” Waterhouse, 1980: 197,198, figs 4-11. Permian, Productus Creek Group,Mangarewa Formation. NZGS localities GS 6071,6072, New Zeal<strong>and</strong>. Holotype NZGS TM5686;paratypes NZGS TM5678-TM5685.† fusiforme, Dentalium Link, 1807: 23. [Fossil].Sternberg, Germany.fusiformis, Cadulus (Gadila) Pilsbry & Sharp, 1898:193, 194, pl. 35, fig. 14 [ex Cooper MS]. <strong>Recent</strong>.San Pedro, California, 4 fms [7 m]. HolotypeUSNM 133809; paratypes USNM 602247,AMNH 148398 (1). Junior synonym <strong>of</strong> Gadilaaberrans (Whiteaves, 1887) (fide Shimek 1989:234).fusticulus, Dentalium Brugnone, 1876: 21, fig. 31.Tertiary. Ficarazzi, Italy. Junior synonym <strong>of</strong> Antalisagilis (G. O. Sars, 1872) (fide Monterosato 1880:64).† gabbi, Cadulus Pilsbry & Sharp, 1898: 236.“Cretaceous” [Eocene]. NE <strong>of</strong> Martinez; Alizos592ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Creek, near Fort Téjon; <strong>and</strong> Tuscan Springs,California, USA. Lectotype ANSP 79582 (designatedby Emerson 1957: 988, 989, pl. 26, figs 1-5);paralectotypes ANSP 79583. Replacement namefor Dentalium (Ditrupa “?”) pusillum Gabb, 1864, ajunior homonym <strong>of</strong> Dentalium pusillum Philippi,1836. Remark: Emerson (1957: 988, 989) determinedthat Gabb’s geological age, “Cretaceous” wasincorrect <strong>and</strong> that this <strong>species</strong> is actually from theEocene.† gabbi, Dentalium Pilsbry & Sharp, 1898a: 470, 471,pl. 10, figs 6, 7, 13, pl. 11, figs 1, 2. Oligocene.Dominican Republic. Type material ANSP 2711(fide Pilsbry 1922: 400). Replacement name forDentalium affine Gabb, 1873, a junior homonym <strong>of</strong>Dentalium affine Deshayes, 1861, <strong>and</strong> D. affineBiondi, 1859.† gadula, Cadulus (Gadila) gadus var. Sacco, 1897:118, pl. 10, figs 88, 89 [as a nomen nudum, Gadusgadulus Doderlein, 1864: 98, <strong>and</strong> Sacco, 1896: 98].Lower Pliocene <strong>of</strong> Zinola, Italy. Lectotype MIGTBS.106.12.010 (selected by Pavia 1991: 138) (figuredby Ferrero Mortara 1984: 311, pl. 56, fig. 3).Junior synonym: Cadulus (Gadila) razzoreiCaprotti, 1979 (fide Pavia 1991: 137, 138). Gadilagadulus (Sacco, 1897) (fide Pavia 1991: 137, 138,pl. 9, figs 12-17).gadulus, Gadus Doderlein, 1864: 98 [16]. Miocene.Monte Gibio, Italy. Nomen nudum. This <strong>species</strong>name was subsequently validated as Cadulus(Gadila) gadus var. gadula Sacco, 1897.gadulus, see gadus.gadus, Dentalium Montagu, 1803: 496, pl. 14, fig. 7.<strong>Recent</strong> (?). English Channel. Possible syntypesBMNH 1950.11.24.1-3. Junior synonym: Dentaliumcoarctatum Lamarck, 1818 (<strong>fossil</strong>; non D. coarctatumBrocchi, 1814) (fide Pilsbry & Sharp 1898:240). Gadila gadus (Montagu, 1803) (fide Pilsbry &Sharp 1898: 186). Eastern <strong>and</strong> Western Atlantic:Africa to Caribbean; (doubtful record from the RedSea by Cooke 1885: 275). 18-280 m. Remarks:Pilsbry & Sharp (1898: 243) quote Hanley sayingthat Dentalium minutu[m] Linnaeus, 1758 “[...] isnot identifiable but [...] that it may be Cadulusgadus”. Rang (1828: 309-312, pl. 18, figs 3-6; 1829:498, pl. 19, fig. E) transferred this <strong>species</strong> to Cleodora(Creseis) [Gastropoda], but this may be based uponmisidentified specimens. Dentalium gadulus[Arduini, 1895: 195] is an incorrect subsequentspelling. Sacco (1897: 118) <strong>and</strong> Pavia (1991: 137,138) determined that Arduini’s material was notconspecific with Montagu’s <strong>species</strong>, but instead isreferable to Gadila gadulus Sacco, 1897.galatheae, Siphonodentalium Knudsen, 1964: 125,126, figs 1, 2. <strong>Recent</strong>. Sunda Trench, Galathea stn465, 10°20’S, 109°55’E, 7000-6900 m. HolotypeZMUC. Striopulsellum galatheae (Knudsen, 1964)(fide Scarabino 1995: 324). Known from type localityonly.gallardoi, Dentalium Rovereto, 1921: 30. Tertiary.Argentina. Junior synonym <strong>of</strong> Dentalium octocostellatumPilsbry & Sharp, 1898 (fide Feruglio 1949b:203, footnote).† gallensteini, Dentalium Gugenberger, 1934: 43, 44,pl. 1, figs 10a, 10b [nomen nudum in Gugenberger1933a: 102; 1933b: 184]. Triassic. North <strong>of</strong>Launsdorf, Austria. Prodentalium gallensteini(Gugenberger, 1934) (fide Stiller 2001: 630).† ganense, Laevidentalium Cossmann in O’Gorman &Cossmann, 1923: 28, pl. 3, figs 5, 6. Eocene. Gan,S <strong>of</strong> Pau, France.gardineri, Dentalium Melvill, 1909: 120, pl. 5, fig. 9.<strong>Recent</strong>. Amirantes Isl<strong>and</strong>, c. 900 miles East <strong>of</strong>fZanzibar, South <strong>of</strong> Seychelles, Percy Sladen TrustExp. stn 20, 160-209 fms [292-381 m]. HolotypeBMNH 1910.3.17.12. Antalis gardineri (Melvill,1909) (fide Ludbrook 1954: 93). Indo-Pacific:Madagascar to New Caledonia. 270-705 m.gardneri, Dentalium Sharp & Pilsbry in Pilsbry & Sharp,1898: 222. Cretaceous, Gaultian. Folkestone,Engl<strong>and</strong>, United Kingdom. Replacement name forDentalium acuminatum Gardner, 1878, a junior secondaryhomonym <strong>of</strong> Dentalium acuminatumDeshayes, 1825, <strong>and</strong> itself a junior homonym <strong>of</strong>Dentalium gardneri (Holzapfel, 1888); replacementname: Dentalium infortunatum Pilsbry & Sharp, 1898.† gardneri, Entalina Sharp & Pilsbry in Pilsbry &Sharp, 1898: 234. Cretaceous, Gaultian. Folkestone,Engl<strong>and</strong>, United Kingdom. Replacementname for Siphodentalium affine Gardner, 1878, ajunior homonym <strong>of</strong> Siphonodentalium affineM. Sars, 1865.† gardneri, Entalis Holzapfel, 1888: 177, pl. 20,fig. 10. Cretaceous. Near Vaals, Netherl<strong>and</strong>s.Fustiaria gardneri (Holzapfel, 1888) (fide Pilsbry &Sharp 1898: 255; Fustiaria here as a subgenus,hence the homonymy!).garrardi, Dentalium (Lentigodentalium) Lamprell &Healy, 1998: 71, figs 68C, 69D, E, 72. <strong>Recent</strong>.Lady Musgrave Isl<strong>and</strong>, Capricorn-Bunker Group,E <strong>of</strong> Gladstone, Queensl<strong>and</strong>, Australia. HolotypeAMS C169770; paratypes AMS C174887 (30),MV F67452 (2; Lady Musgrave Isl<strong>and</strong>, Queensl<strong>and</strong>).Dentalium garrardi Lamprell & Healy,1998. Indo-Pacific: eastern Australia; Queensl<strong>and</strong>.10-37 m.† gaultinus, Cadulus Gardner, 1878: 63, pl. 3, fig. 48.Cretaceous, Gaultian. Folkestone, Engl<strong>and</strong>, UnitedKingdom.gaussianum, Dentalium majorinum var. Plate, 1908b:5, 6, figs 1-4. <strong>Recent</strong>. Antarctica, Deutsche SüdpolarExp. stn 56, 66°02’S, 89°38’E, 385 <strong>and</strong> 350 m.Lectotype ZMB 59730a (designated by Kilias 1995:175); paralectotype ZMB 59730b. Synonym <strong>of</strong>Dentalium majorinum Mabille & Rochebrune,1889.gayi, Dentalium Philippi, 1887a: 107, pl. 12, fig. 19;1887b: 101, pl. 12, fig. 19. Tertiary. Matanzas <strong>and</strong>ZOOSYSTEMA • 2004 • 26 (4)593


Steiner G. & Kabat A. R.Carauma, Chile. Junior synonym <strong>of</strong> Dentaliumsulcosum J. de C. Sowerby, 1846 (fide Ortmann1902: 157, 159).gazellae, Dentalium (Episiphon) Plate, 1908a: 356,pl. 30, figs 40, 41. <strong>Recent</strong>. Northwest Australia,Gazelle Exp., 9 fms [16 m]. Holotype ZMB 33195.Episiphon(?) gazellae (Plate, 1908) (fide Scarabino1995: 289). Known from type locality only.† geinitzi, Dentalium (Fustiaria) Koenen, 1892: 989,990, pl. 59, fig. 12. Lower Oligocene. Lattorf,Germany. Fustiaria geinitzi (Koenen, 1892).Remarks: name for Dentalium fissura sensu Koenennon Dentalium fissura Lamarck, 1818. Junior secondaryhomonym <strong>of</strong> Dentalium geinitzi (Böhm,1885). We have no information <strong>of</strong> Koenen’s namebeing used since 1899 <strong>and</strong>, therefore, do notrename this homonym until future research provesit a valid <strong>species</strong>.† geinitzi, Fustiaria Böhm, 1885: 34, 35, pl. 1, fig. 7.Cretaceous. Vaals [Vael], Netherl<strong>and</strong>s. Dentaliumgeinitzi (Böhm, 1885) (fide Pilsbry & Sharp 1898:254).† geinitzianum, Dentalium Ryckholt, 1851: 70.Cretaceous: Turonian. Tournay <strong>and</strong> Montigniessur-Roc,Belgium. Remark: name for Dentaliummedium sensu Geinitz 1840 non Dentaliummedium J. Sowerby, 1814.† geminatum, Dentalium Goldfuss, 1841: 4, pl. 166,fig. 13. Oligocene. Bünde, Westphalia, Germany.Syntypes GPIB 1072. Remarks: Janssen (1978a: 14,15) originally considered this name to be a “nomenoblitum” <strong>and</strong> thus not a senior synonym <strong>of</strong>Dentalium kickxi Nyst, 1843. Janssen (1978b: 138,139) concluded from the Goldfuss types that it wasa valid <strong>species</strong> <strong>and</strong> a senior synonym <strong>of</strong> D. kickxii.Most recently, Janssen (1987: 174-176) determinedthat Antalis geminatum was found in the LateOligocene <strong>and</strong> A. kickxii was found in the MiddleOligocene; the two <strong>species</strong> are not synonyms.gemmiparum, Dentalium Melvill, 1909: 120, pl. 5,fig. 10. <strong>Recent</strong>. Chagos Archipelago, Diego GarciaLagoon, Indian Ocean. Holotype BMNH 1910.3.17.13. Coccodentalium gemmiparum (Melvill,1909) (fide Scarabino 1995: 264). Indo-Pacific:Diego Garcia to Philippines <strong>and</strong> New Caledonia.240-2250 m.georgiense, Dentalium (Antalis) occidentale Henderson,1920: 43. <strong>Recent</strong>. Georges Bank [Massachusetts],Albatross stn 2079, 41°13’N, 66°19’50”W, 75 fms[137 m]. Syntypes (<strong>of</strong> Dentalium occidentalis sulcatumVerrill, 1884) USNM 38398 (1; USBF 2079,75 fms [137 m]), USNM 35093 (4; USBF 2077,1255 fms [2290 m]). Replacement name forDentalium occidentalis sulcatum Verrill, 1884, a juniorhomonym <strong>of</strong> Dentalium sulcatum Lamarck,1818. Synonym <strong>of</strong> Antalis occidentalis (Stimpson,1851) (fide Pilsbry & Sharp 1897: 48, 49).† germanicum, Dentalium Chenu, 1843: pl. 5,fig. 15a, b (name <strong>and</strong> figure only). Type locality notspecified. Remarks: Chenu (1852: 7), in the text,instead used Dentalium striatum Lamarck, 1818 forthe specimen figured as “D. germanicum”. However,Lamarck’s name is a junior homonym <strong>of</strong> D. striatumBorn, 1778 <strong>and</strong> a junior synonym <strong>of</strong>Fissidentalium rectum (Gmelin, 1791). Pilsbry &Sharp (1898: 205) suggested that this was “evidentlya European Tertiary <strong>species</strong>”.† gibba, Cadulus ovulum var. Seguenza, 1879: 276.Pliocene. Reggia, Calabria, Italy. Senior homonym<strong>of</strong> Cadulus gibbus Jeffreys, 1883.gibbosus, Cadulus (Polyschides) Verco, 1911a: 213,pl. 26, fig. 6. <strong>Recent</strong>. Off Cape Jaffa, SouthAustralia, 300 fms [548 m]. Holotype SAM D13729. Polyschides gibbosus (Verco, 1911) (fideLamprell & Healy 1998: 168). Indo-Pacific,Australia. 17-2780 m. Remark: the three paratypesmentioned in the original description were notlocated.gibbus, Cadulus Jeffreys, 1883: 666, pl. 49, fig. 10.<strong>Recent</strong>. Type locality not specified. SyntypesBMNH 1885.11.5.1287 (1; Porcupine stn 13,40°16’N, 9°37’W, 217 fms [396 m]), USNM175858 (1; Bay <strong>of</strong> Biscay, Travailleur). Cadulusgibbus Jeffreys, 1883. Junior homonym <strong>of</strong> Cadulusovulum var. gibba Seguenza, 1879. Eastern Atlantic:British Isl<strong>and</strong>s, Bay <strong>of</strong> Biscay. 396 m. Remark: theauthors intend to submit a petition to the InternationalCommission <strong>of</strong> Zoological Nomenclature,pursuant to ICZN (1999) article 23.9.3, to preserveJeffreys’ name.giganteum, Dentalium Phillips, 1829. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.giganteum, Dentalium “?” Zenker, 1836: 232. Tertiary.Flohberge (Mühlthal), near Jena, Germany. Typesnot located. Junior homonym <strong>of</strong> Dentalium giganteumPhillips, 1829. Remark: the lack <strong>of</strong> type material<strong>and</strong> an illustration in combination with theshort description renders this a dubious <strong>species</strong> forwhich we prefer not to introduce a replacementname.giganteum, Dentalium Chenu, 1842: pl.1, fig. 3; 1850:4. “Fossil” [type locality not specified]. Types notlocated. Junior homonym <strong>of</strong> Dentalium giganteumPhillips, 1829. Replacement name: Dentalium rexPilsbry & Sharp, 1898.giganteum, Dentalium J. de C. Sowerby, 1846: 263,pl. 2, fig. 1. Tertiary. Navidad, Chile. Types notlocated. Junior homonym <strong>of</strong> Dentalium giganteumPhillips, 1829 <strong>and</strong> D. giganteum Chenu, 1842.Replacement name: Dentalium subgiganteumd’Orbigny, 1852.gigas, Dentalium Jeffreys, 1870b: 432. Nomen nudum.† girtyi, Plagioglypta Knight, 1940: 315, pl. 6, fig. 2.Late Permian, Whitehorse Group, Rush SpringsFormation, Dozier S<strong>and</strong>stone Member. Dolomitebeds at base <strong>of</strong> Dozier, Localities 7a (CollingsworthCounty) <strong>and</strong> 8a (6 miles S <strong>of</strong> Memphis, at summit<strong>of</strong> Hogback Butte, Hall County), Texas, USA.594ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Holotype YPM 15336; paratype YPM 50071 (fideWhite et al. 1999: 43).† glabellum, Dentalium Blake, 1907: 101, 102, pl. 9,fig. 12 [nomen nudum in Bean 1839: 62]. Jurassic.Cornbrash, Scarborough, Engl<strong>and</strong>, UnitedKingdom. Holotype BMNH G.16996.† glabratum, Antale Stoliczka, 1868: 445, pl. 27,figs 24, 25. Cretaceous. Ootatoor <strong>group</strong>. Odium,India.glabrum, Dentalium Montagu, 1803. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.glabrum, Dentalium Geinitz, 1842: 74, pl. 18, fig. 28;1843: 11, pl. 1, fig. 27. Upper Cretaceous. Tisa[Tyssa], Bohemia, Czech Republic. Junior homonym<strong>of</strong> Dentalium glabrum Montagu, 1803.Replacement name: Dentalium ovosectum Sharp &Pilsbry in Pilsbry & Sharp, 1898.glabrum, Dentalium Fuenzalida, 1942: 412, 423, 425.Tertiary. Brunswick Peninsula, Patagonia, Chile.Nomen nudum. Remark: Feruglio (1949b: 160)cited this name, but did not make it available.† gladiolus, Dentalium Eichwald, 1846: 447. Jurassic.Goliowo, Dorogomilow, Mniowniki <strong>and</strong> Khoroschowo,near Moscow; Mount Worobyew; Jelatmaon the Oka River, near Antonowo-Poustosch,Russia. Junior synonyms: Dentalium subancepsTrautschold, 1861 (fide Eichwald 1868: 799;Pilsbry & Sharp 1898: 223). Remarks: Eichwald’soriginal description was cursory; the locality informationherein is from the subsequent re-description(Eichwald 1868: 799, 800). This <strong>species</strong> wasmisidentified as Dentalium cylindricum by G. Fischer(1843).glans, Cadulus Scarabino, 1995: 350, 351, figs 155,160g. <strong>Recent</strong>. New Caledonia, Loyalty Basin, RVCoriolis, BIOGEOCAL, stn KG 227, 21°33’S,166°24’E, 500 m. Holotype MNHN <strong>and</strong> 7 paratypes(2 from stn KG 219; 5 from Vauban stn 40).Cadulus glans Scarabino, 1995. Indo-Pacific: NewCaledonia. 250-570 m.glaucarena, Dentalium (Antalis) Dell, 1953: 48,figs 16, 18, 21. <strong>Recent</strong>. Chatham Rise, Discovery IIstn 2733, 45°48’S, 178°58’W, 200-300 fms [365-548 m]. Holotype NMNZ M5678; paratypesNMNZ M5679 (5). Antalis glaucarena (Dell,1953) (fide Scarabino 1995: 236). Indo-Pacific,New Zeal<strong>and</strong>. 365-602 m.† glaucoterrarum, Dentalium Maury, 1917: 323 [159],pl. 52 [26], fig. 4. Miocene. Rio Gurabo at LosQuemados, Dominican Republic.† gnizum, Dentalium de Gregorio, 1890: 173, pl. 17,figs 42, 43. Eocene. Claiborne, Alabama. Remark:“Probably a young Cadulus. Unrecognizable” (fidePilsbry & Sharp 1898: 205).g<strong>of</strong>asi, Laevidentalium Scarabino, 1995: 294, 295,figs 99, 102a. <strong>Recent</strong>. Philippines, RV Coriolis,MUSORSTOM 2, stn CP 70, 14°01’N, 120°17’E,191 m. Holotype MNHN; paratypes MNHN (8;1 from stn CP 70; 2 from stn CP 68; 1 from stn CP83; 1 from stn CP 97; 3 from stn CP 102), AMSC201728 (1; stn CP 96), USNM 890869 (1; stnCP 20). Laevidentalium g<strong>of</strong>asi Scarabino, 1995.Indo-Pacific: Philippines to New Caledonia. 195-320 m.g<strong>of</strong>toni, Dentalium (Dentalium) Lamprell & Healy,1998: 45, 46, figs 37C, 38E, 41. <strong>Recent</strong>. MiddleBanks, Moreton Bay, Queensl<strong>and</strong>, Australia, 31 m.Holotype QM MO38982; paratypes AMSC172168 (2), MV 68195 (2), WAM 283-93 (2),QM MO38996 (2; Redl<strong>and</strong> Bay, south Queensl<strong>and</strong>),QM MO38983 (11; Dunwich, StradbrokeIsl<strong>and</strong>, Moreton Bay, south Queensl<strong>and</strong>), AMSC169791 (21; Shoal Point, Mackay, northQueensl<strong>and</strong>). Dentalium g<strong>of</strong>toni Lamprell &Healy, 1998. Indo-Pacific: Queensl<strong>and</strong>, Australia.2-183 m.† golovtschenki, Dentalium Makarenko, 1970: 111,112, text-fig. 11, pl. 22, figs 6-9. Paleocene.Luzanovka, northern Ukraine. Holotype AkademiiaNauka Ukraine (Kiev), No. 17/50.† gonatodes, Dentalium Martin, 1885: 192, pl. 10,fig. 194; 1887: 192, pl. 10, fig. 194. Pliocene/Quaternary. Samarang <strong>and</strong> Batavia (Jakarta), Java,Indonesia. Syntypes RGM 7280 (1), RGM 7281(2), RGM 7278 (3), RGM 7279 (1) (fide Van denHoek Ostende et al. 2002: 111). Dentalium(Laevidentalium) gonatodes Martin, 1885 (fideMartin 1919: 118).gordonis, Cadulus Yokoyama, 1920. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.goreanum, Dentalium Clessin, 1896. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.gouldii, Dentalium Dall, 1889: 424, 425, pl. 26, fig. 4.<strong>Recent</strong>. 12 miles E from Frying Pan Shoals, SouthCarolina, 12 fms [22 m]. Lectotype USNM 82997(designated by Henderson 1920: 30); paralectotypesUSNM 94085 (USFC stn 2145; now theholotype <strong>of</strong> Dentalium gouldii colonense Henderson,1920), MCZ 7707 (Blake stn 65). Synonyms:Dentalium sexangulare Hilgard & Hopkins, 1878non Lamarck, 1818 (fide Pilsbry & Sharp 1897:21), Dentalium gouldii colonense Henderson, 1920,Dentalium gouldii portoricense Henderson, 1920.Dentalium gouldii Dall, 1889. Western Atlantic:South Carolina (USA) <strong>and</strong> Bermuda to Brazil(doubtful record from the coast <strong>of</strong> West Africa byThomas 1975: 292; Nicklés 1979: 48). 10-76 m.gracile, Dentalium Moore, 1866: 86, pl. 5, fig. 23.Jurassic, Middle Liassic. Camerton, Engl<strong>and</strong>,United Kingdom. Holotype in Bath Museum(Engl<strong>and</strong>) (fide Richardson 1906: 576, pl. 45,fig. 19). Junior homonym <strong>of</strong> Dentalium gracilisHall & Meek, 1855. Remark: a junior synonym <strong>of</strong>Dentalium elongatum Goldfuss, 1841 (fideWoodward 1893: 352; Richardson 1906: 575).gracile, Dentalium Jeffreys, 1870a: 74. <strong>Recent</strong>.Mediterranean, 310 fms [566 m]; Aegean Sea, 100-250 fms [183-456 m]; Vigo, Spain. Syntypes:ZOOSYSTEMA • 2004 • 26 (4)595


Steiner G. & Kabat A. R.USNM 175979 (E <strong>of</strong> Malta, 310 fms [566 m]),USNM 175977 (Aegean Sea, 100-250 fms [183-456 m]). Junior homonym <strong>of</strong> Dentalium gracilisHall & Meek, 1855 <strong>and</strong> Dentalium gracile Moore,1866. Junior synonym <strong>of</strong> Episiphon filum(Sowerby, 1860) (fide Jeffreys 1883: 661).gracile, Dentalium Philippi, 1887a: 107, pl. 12, fig. 15;1887b: 101, pl. 12, fig. 15. Tertiary. Navidad,Chile. Junior homonym <strong>of</strong> Dentalium gracilis Hall& Meek, 1855, Dentalium gracile Moore, 1866 <strong>and</strong>Dentalium gracile Jeffreys, 1870. Replacementname: Dentalium navidadense Pilsbry & Sharp,1898: 210. Remark: Dentalium philippii Cossmann,1907; an unnecessary replacement name.gracile, Dentalium Grönwall & Harder, 1907: 36,pl. 1, fig. 19. Paleocene. Jutl<strong>and</strong>, Denmark. Juniorhomonym <strong>of</strong> Dentalium gracilis Hall & Meek,1854, Dentalium gracile Moore, 1866, Dentaliumgracilis Jeffreys, 1870, Dentalium gracilis Philippi,1887; replacement name: Dentalium subgracileEmerson, 1954.† gracilicostatum, Dentalium (Fissidentalium)Singleton, 1943: 275-276, pl. 12, fig. 6, pl. 13,fig. 9. Lower Eocene. Pebble Point Beds, CoastalCliffs, 2.5 miles southeast <strong>of</strong> Princetown, Victoria,Australia. Holotype NMV P127989 (formerlyMelbourne University Geology Department 1871).Fissidentalium gracilicostatum (Singleton, 1943)(fide Darragh 1997: 92, 93).† gracilina, Cadulus (Gadila) Sacco, 1897: 117, pl. 10,figs 86, 87 [as a nomen nudum in Sacco 1896: 98].Miocene. Turin hills <strong>and</strong> San Agata, Italy.Lectotype MIGT BS.106.12.009 (selected by Pavia1991: 144); paralectotype BS.106.12.008 (fideFerrero Mortara et al. 1984: 311, pl. 56, fig. 2).Gadila gracilina (Sacco, 1897) (fide Pavia 1991:144, 145, pl. 8, figs 1-4).gracilis, Cadulus Jeffreys, 1877: 157. <strong>Recent</strong>. Valorousstn 13, 56°01’N, 34°42’W, 690 fms [NorthAtlantic, 1259 m]. Holotype USNM 175853.Cadulus gracilis Jeffreys, 1877. Eastern Atlantic:Greenl<strong>and</strong> to Portugal. 453-1998 m. Remarks:Jeffreys referred to a single specimen from Valorousstn 13. However, Warén (1980: 52) stated that thetype locality was not specified <strong>and</strong> listed additionalsyntypes: BMNH 1885.11.5.1300-1301 (Porcupine1870 stns 16-17a [<strong>of</strong>f Portugal]; stn 16: 39°55’N,9°56’W, 994 fms [1723 m]; stn 17: 39°42’N,9°43’W, 600-1095 fms [1095-1998 m]; stn 17a:39°39’N, 9°39’W, 740 fms [1351 m]), USNM175854 (Valorous stn 16, 55°10’N, 25°58’W, 1785fms [Greenl<strong>and</strong>, 3258 m]), USNM 175855(Porcupine 1870 stn 17, 39°42’N, 9°43’W, 600-1095 fms [1095-1998 m]), USNM 175856(Porcupine 1870 stn 19a, Portugal). The lot BMNH1885.11.5.1300 is labeled “figd. syntyp”. Warén (inlitt. 10.II.2000) explained this discrepancy: “Jeffreyslisted material from the Lightning <strong>and</strong> Porcupineexpeditions for most new molluscs in the ‘New <strong>and</strong>peculiar...’ series. He also had specimens <strong>of</strong> Cadulusgracilis, as listed later in the report on these expeditions.I therefore considered it an omission from hisside, not to mention these”.† gracilis, Dentalium Hall & Meek, 1855: 393, pl. 3,figs 11a-11c. Cretaceous. Sage Creek, “Nebraska”(Montana <strong>and</strong> Wyoming?), USA. Dentalium gracileHall & Meek, 1855 (fide Pilsbry & Sharp 1898: 197).† gracilis, Episiphon Garvie, 1996: 45, 46, pl. 9, figs 7,8. Eocene, Lower Claibornian, Reklaw Formation,Marquez Member. Locality 4, Bluff on RidgeCreek, Texas, USA. Holotype PRI 30310; paratypesPRI 30311, PRI 30312.† gracillima, Dentalium (Coccodentalium) radula var.Sacco, 1897: 111, pl. 10, figs 16-18 [as a nomennudum in Sacco 1896: 98]. Miocene. San Agata,Italy. Syntypes MIGT BS.106.04.010, BS.106.04.011 (fide Ferrero Mortara et al. 1984: 308).gradile, Paradentalium Chistikov, 1979a: 109, 111,fig. 1. <strong>Recent</strong>. Vietnam, Orlik stn 203, 20°44’N,112°13’E, 60 m. Holotype ZIN; paratype MMSU.Paradentalium gradile Chistikov, 1979. Indo-Pacific: Vietnam. 60-106 m.grahamense, Dentalium majorinum var. Odhner, 1931:4. <strong>Recent</strong>. Antarctica, Graham L<strong>and</strong>, 65°19’S,56°48’W, 400 m. Syntypes SMNH 4193.Synonym <strong>of</strong> Dentalium majorinum Mabille &Rochebrune, 1889.grahami, Dentalium (Dentalium) Lamprell & Healy,1998: 23-25, figs 9E, 10E, 16, 17D-F. <strong>Recent</strong>. OffClarence River, New South Wales, Australia,29°25’S, 153°22’E, 37-55 m. Holotype AMSC174892; paratypes AMS C154599 (1), AMSC173469 (1; 29°39.8’S, 153°26.4’E, south <strong>of</strong>Yamba, New South Wales, 55 m), AMS C154602(55; <strong>of</strong>f Solitary Isl<strong>and</strong>, New South Wales, 29°50-53’S, 153°24-23’E, 54-58 m), AMS C169957(1; Keppel Bay, southern Queensl<strong>and</strong>, 54 m), AMSC169958 (2; Keppel Bay, southern Queensl<strong>and</strong>,54 m). Dentalium grahami Lamprell & Healy,1998. Indo-Pacific: eastern Australia. 28-80 m.† granadanum, Dentalium Anderson, 1929: 144,pl. 13, fig. 3. Miocene. CAS locality 267, LasPerdices Group, 1 mile W <strong>of</strong> Puerto Colombia,northern Colombia. Holotype CAS 4638.† gr<strong>and</strong>aevum, Dentalium Winchell, 1863: 18.Carboniferous. Burlington, Iowa, USA.† gr<strong>and</strong>e, Dentalium Deshayes, 1825: 365, pl. 17,figs 1-3. Eocene. Grignon, La Chapelle near Senlis<strong>and</strong> Valmondois, Paris Basin, France. Remarks: avalid <strong>species</strong> <strong>of</strong> Fissidentalium (fide Le Renard 1995:175; Le Renard & Pacaud 1995: 85). This namehas been inappropriately used by Nyst (1836: 103)for a <strong>species</strong> subsequently described as Dentaliumacutum Hébert, 1849 (fide Pilsbry & Sharp 1898:197; Dentalium nystii d’Orbigny, 1852 is a juniorsynonym), <strong>and</strong> by Abich (1858: 561 [25]) for a<strong>species</strong> tentatively synonymized with Dentaliumtrauscholdi Koenen, 1868 (fide Koenen 1868: 160).596ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)gr<strong>and</strong>is, Cadulus Verrill, 1884: 219, pl. 44, fig. 20.<strong>Recent</strong>. 230 miles E <strong>of</strong> Martha’s Vineyard[Massachusetts], USBF stn 2076, 41°13’N,66°00’50”W, 906 fms [1651 m]. Syntypes USNM34735 (16, USBF stn 2076), USNM 35184 (USBF2084, 1290 fms [2354 m]), USNM 38116 (USBF2103, 1091 fms [1991 m]); MCZ 186804 (2); furtherlocations in original description: USBF 2052,1098 fms [2004 m]; USBF 2111, 938 fms[1712 m]; USBF 2115, 843 fms [1538 m]; <strong>and</strong>USBF 2043, 1467 fms [2677 m]). Polyschides gr<strong>and</strong>is(Verrill, 1884) (fide Pilsbry & Sharp 1898: 154).Western Atlantic. 1538-2677 m.† gr<strong>and</strong>is, Eodentalium Medina & del Valle, 1985: 3-5(Spanish), 3, 4 (English), pl. 1, figs a-f, i-k, p,q. Upper Cretaceous, Sobral Formation. MarambioIsl<strong>and</strong>, James Ross Group Isl<strong>and</strong>s, Antarctica.Holotype CIRGEO (Centro de Investigaciones enRecursos Geológicos) PI 247 (now housed inMuseo Argentino de Ciencias Naturales, BuenosAires, fide G. Pastorino in litt. 17.I.2001).† granosa, Entalis Martin, 1885: 193, 194, pl. 10,fig. 196; 1887: 193, 194, pl. 10, fig. 196. Miocene.Ngembak, Java, Indonesia. Syntype RGM 7275(fide Van den Hoek Ostende et al. 2002: 111).Remarks: Van den Hoek Ostende et al. (2002: 111)listed this <strong>species</strong> in Dentalium, following two publishedchecklists. If this transfer from Antalis (thesenior synonym <strong>of</strong> Entalis, is accepted, thenDentalium granosum (Martin 1885) is a junior secondaryhomonym <strong>of</strong> D. granosum Eichwald, 1857,<strong>and</strong> will need to be renamed. We have not renamedthis taxon, as further study is merited.† granosum, Dentalium Eichwald, 1857: 584. Silurian[“Grauwackenkalksteine”]. Pulkowa [Poulkova],Russia. Remarks: figured by Eichwald 1860: 1061,pl. 40, fig. 7. Koken (1897: 214) remarked that theshape <strong>of</strong> the shell <strong>of</strong> this <strong>species</strong> was completelydivergent from that <strong>of</strong> Dentalium (“die Structurdieser Röhren ist ganz abweichend von der einesDentalium”), <strong>and</strong> questioned whether this <strong>species</strong>was referable to the <strong>Scaphopoda</strong>.† granvillensis, Dentalium Herrick, 1888: 92.Paleozoic (“Subcarboniferous”). Granville, LickingCounty, Ohio, USA.greenlawi, Cadulus (Platyschides) Henderson, 1920:123, 124, pl. 18, fig. 11. <strong>Recent</strong>. Off Fowey Light,Florida, Eolis stn 380, 130 fms [237 m]. HolotypeUSNM 314767. Gadila greenlawi (Henderson,1920). Known from type locality only.† groenl<strong>and</strong>icum, Dentalium Ravn, 1918: 354, 355,pl. 8, figs 3, 4. Cretaceous. Ata <strong>and</strong> Niakornat,Nugsuak-Halvö, western Greenl<strong>and</strong>.† groenl<strong>and</strong>icus, Cadulus Ravn, 1918: 356, pl. 8,fig. 8. Cretaceous. Alianaitsunguak, Nugsuak-Halvö, western Greenl<strong>and</strong>.† grossheimi, Dentalium Merklin, 1961: 156, 157,text-fig. 1. Middle Eocene. Northern Caucasus,Goryachiy Kluych, near Krasnodar, Russia.gruveli, Dentalium (Laevidentalium) Dautzenberg,1910: 109, 110, pl. 3, figs 21-23. <strong>Recent</strong>. Baie deCansado [Mauretania] (1908); Baie de Cansado[Mauretania], 6-8 m (1909); 5 milles E de la Pointedu Repos [Mauretania], 9-10 m; Baie de Rufisque[Senegal], 18-20 m; W[est] du Cap Rouge [Ghana].Syntypes MNHN (2 lots from Baie de Cansado<strong>and</strong> 1 from Cap Rouge). Fustiaria(?) gruveli(Dautzenberg, 1910) (fide Nicklès 1979: 59).Eastern Atlantic: Africa. 6-20 m.† gucuoensis, Dentalium Yao, 1990: 31, 32, pl. 1,figs 1-3, 5-8. Lower Cretaceous, Gucuo II Formation.Gucuo village, Nyalam, southern Xizang[Tibet], China. Holotype (XG886-19) <strong>and</strong>paratypes, Institute <strong>of</strong> Geology, Chinese Academy<strong>of</strong> Geological Sciences, Beijing.† guidottii, Dentalium Sacco, 1897: 95 [as a nomennudum in Sacco 1896: 97]. Pliocene. Rivalta,Piacenza, Italy. Replacement name for Dentaliumdeshayesii Cocconi, 1874, a junior homonym <strong>of</strong>Dentalium deshayesii Risso, 1826.guillei, Antalis Scarabino, 1995: 235, figs 43, 45k.<strong>Recent</strong>. Réunion, Indian Ocean, MD32, stn DC136, 20°46’S, 55°36’E, 915-922 m. HolotypeMNHN; paratypes MNHN (11; 4 from stn 136;7 from stn DC 46), NM (1; stn DC 56). Antalisguillei Scarabino, 1995. Indian Ocean: Réunion.150-915 m.† guineense, Dentalium (Laevidentalium) Adegoke,1977: 62, 63, pl. 5, figs 15-18. Paleocene, EwekoroFormation. Nigeria. Holotype UIMG 176;paratypes UIMG 177; USNM Paleobiology174763, 174764, PRI 29764-PRI 29769.† guizhouensis, Plagioglypta Stiller, 2001: 623-626,figs 4, 5.5-5.7. Upper Anisian, Middle Triassic.Bangtoupo, NNE <strong>of</strong> Qingyan, Guizhou Province,southwestern China. Holotype GPIM B6D-1.S2.F35-1; paratype GPIM B6D-1.S2.F35-2.habutae, Dentalium Kuroda & Kikuchi, 1933: 8, pl.1, figs 12, 13, text-figs 3, 4. <strong>Recent</strong>. Toyama Bay,Japan, 150-350 m. Types in FGK. Junior synonym:Dentalium tessellatum Scarabino, 1995 (V. Scarabino,n. syn.). Entalinopsis habutae (Kuroda &Kikuchi, 1933) (fide Habe 1957: 13). Indo-Pacific:Japan. 93-350 m.† haeringense, Dentalium Dreger, 1892: 12, pl. 1,figs 2a, 2b. Pliocene? Häring, near Kirchbichl,Tirol, Austria. Junior synonym <strong>of</strong> Coccodentaliumtrautscholdi Koenen, 1868 (fide Merklin &Goncharova 1967: 118).halius, Cadulus (Cadulus) Henderson, 1920: 149,pl. 20, fig. 12. <strong>Recent</strong>. Off English Harbor,Antigua, 120 fms [219 m]. Holotype USNM596599; paratypes USNM 314936, AMNH148332 (1). Junior synonym <strong>of</strong> Cadulus podagrinusHenderson, 1920 (n. syn., by First Reviser actionherein). Known from type locality only.hamatum, Dentalium? Forbes, 1846. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.ZOOSYSTEMA • 2004 • 26 (4)597


Steiner G. & Kabat A. R.“hamatus”, Dentalium – Conrad 1870b. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.hancocki, Dentalium (Tesseracme) Emerson, 1956: 3-6,fig. 1. <strong>Recent</strong>. One-half mile SE <strong>of</strong> Hughes Point,Santa Maria Bay, Baja California, Mexico,24°44’31”N, 112°14’59”W, 11 fms [20 m].Holotype LACM 1758; paratypes LACM 2887(2; <strong>of</strong>f Punta Penasco, Sonora, 11 fms [20 m]),LACM 2888 (1; 2.25 miles SE <strong>of</strong> Cape Pulmo, BajaCalifornia, Mexico, 20 fms [37 m]), LACM 2889(1; Agua Verde Bay, Baja California, Mexico,20 fms [37 m]). Tesseracme hancocki (Emerson,1956). Eastern Pacific: Mexico. 9-36 m.† hanguense, Dentalium Cox, 1931: 203, 204, pl. 21,figs 12, 13. Tertiary. Hangu Shales, Samana Range,India. Holotype Geological Survey India 14703;paratype Geological Survey India 14702.hannai, Dentalium Baker, 1925: 84, 85, pl. 10, figs 4,5. <strong>Recent</strong>. Baja California Norte; Pacific coast, IslasLos Coronados, c. 1 mi S <strong>of</strong> South CoronadoIsl<strong>and</strong>, [10-18 fms, 18-33 m]. Holotype CAS066343; paratypes CAS 060763, SDNHM 1788,SDNHM 1789, SDNHM 1811 (fide Hertz 1994:11), UCM 23659 (fide Wu & Br<strong>and</strong>auer 1982:40). Junior synonym <strong>of</strong> Graptacme semistriata(Turton, 1819) (fide Keen 1971: 886, as Dentalium(Graptacme) semipolitum).† hannonicum, Dentalium Briart & Cornet, 1889: 81.Upper Eocene. Mons, Belgium.harrisoni, Fissidentalium (Pictodentalium) formosumHabe, 1970: 95, figs 1, 2. <strong>Recent</strong>. South China Sea.Holotype NSMT Mo 37303; paratype NSMT Mo37304. Synonym <strong>of</strong> Pictodentalium formosum(Adams & Reeve, 1850) (fide Scarabino 1995: 267).hatterasensis, Cadulus (Gadila) agassizii var. Sharp &Pilsbry in Pilsbry & Sharp, 1898: 169, 170, pl. 33,figs 50-54. <strong>Recent</strong>. Off Cape Hatteras [NorthCarolina], 35°19’30”N, 75°14’12”W, 293 fms[535 m]. Syntypes ANSP 35578. Synonym <strong>of</strong>Gadila agassizii (Dall, 1881) (n. syn.).† haytensis, Dentalium Gabb, 1873: 244. Miocene.Dominican Republic. Lectotype ANSP 2715(selected by Woodring 1925: 203 as “holotype”);paralectotype ANSP 79219. Laevidentaliumhaytense (Gabb, 1873) (fide Woodring 1925: 202,203).healyi, Dentalium n. nom. <strong>Recent</strong>. Darnley Isl<strong>and</strong>,Torres Straits [northern Australia], 30 fms [55 m].Holotype AMS C170764. Replacement name forDentalium duodecimcostatum Brazier, 1877 nonDentalium duodecimcostatum d’Archiac, 1850.Dentalium healyi Steiner & Kabat, 2004. Indo-Pacific: northern Australia. 9-500 m.hedleyi, Dentalium (Dentalium) Lamprell & Healy,1998: 23, figs 9D, 10D, 15. <strong>Recent</strong>. Swain Reefs,Queensl<strong>and</strong>, Australia, dredged <strong>of</strong>f s<strong>and</strong> bottom.Holotype AMS C171343; paratypes AMSC151517 (15 full specimens, 84 broken pieces, SE<strong>of</strong> Swain Reefs, Queensl<strong>and</strong>, 22°31.4’S, 152°32.6’E,100 m). Dentalium hedleyi Lamprell & Healy,1998. Indo-Pacific: Queensl<strong>and</strong>, Australia. 1-100 m.† helveticum, Dentalium sexangulum “mut.”Cossmann & Peyrot, 1917: 160, 161, pl. 1, figs 1-3. Miocene, Helvetian. Salles (Largileyre), France.† helveticum, Siphonodentalium (Dischides “?”)Cossmann & Peyrot, 1917: 186, pl. 2, figs 11-13.Miocene, Helvetian <strong>and</strong> Tortonian. Orthez (LeParen), Sallespisse, Salles (Largileyre) <strong>and</strong>Saubrigues, France.† hemigymnum, Dentalium (Antale) degrangei var.Cossmann & Peyrot, 1917: 165, pl. 1, figs 25, 26.Miocene, Helvetian. Salies-de-Béarn, La Sime <strong>and</strong>Orthez (Houssé), France.hemileuron, Dentalium Verco, 1911a: 208, pl. 26,fig. 2. <strong>Recent</strong>. Off Cape Jaffa, South Australia,300 fms [549 m]. Holotype SAM D 13727;paratypes USNM 251985 [BMNH 1913.4.30.6-9,possible paratypes]. Junior synonym: Dentaliumtignum Colman, 1958 (fide Lamprell & Healy1998: 29). Paradentalium hemileuron (Verco,1911) (fide Cotton & Godfrey 1933: 139). Indo-Pacific: South Australia. 0-1463 m.hendersoni, Cadulus (Gadilopsis) Woodring, 1925:207, pl. 28, fig. 4. Miocene. Bowden, Jamaica.Holotype USNM Paleobiology 352724. Junior synonym<strong>of</strong> Gadilopsis dentalina (Guppy, 1867) (fideWoodring 1973: 487). Remark: Woodring (1973:487) concluded that this “should be treated as aform <strong>of</strong> C. dentalinus. Whether it is a sub<strong>species</strong> oran infrasubspecific form is indeterminable”.hepburni, Cadulus Dall, 1897: 12, 13, pl. 1, fig. 13.<strong>Recent</strong>. Near Victoria, Vancouver Isl<strong>and</strong>, BritishColumbia, 60 fms [110 m]. Lectotype USNM107612 (designated by Shimek 1989: 234). Juniorsynonym <strong>of</strong> Gadila aberrans (Whiteaves, 1887) (fideShimek 1898: 234). Remark: the lectotype designationby Shimek was probably unnecessary because hestated that the USNM 107612 is “1 specimen, aparatype, the figured type”; furthermore, there is noindication in Dall’s original description that he hadmore than one specimen. If there were no other syntypes,then this specimen is the holotype.† heptagonum, Dentalium Boettger, 1882: 140, pl. 12,figs 5a, 5b. Oligocene. Bawang River, ResidenzDjokjakarta, Java, Indonesia.† herculeum, Dentalium de Koninck, 1863: 8, pl. 4,figs 10-12. Carboniferous. Salt Range, India.Remarks: Pilsbry & Sharp (1898: 231) listed Entalisherculea Waagen, “1887” as a junior synonym, butWaagen (1890: 181) is merely a new generic combinationfor de Koninck’s <strong>species</strong>. Plagioglypta herculea(de Koninck, 1863) (fide Waterhouse 1980:198).† herritschi, Dentalium Gugenberger, 1934: 44, pl. 1,fig. 10 [nomen nudum in Gugenberger 1933a: 102;1933b: 184]. Triassic. North <strong>of</strong> Launsdorf, Austria.Prodentalium herritschi (Gugenberger, 1934) (fideStiller 2001: 630).598ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)heteroschides, Cadulus (Polyschides) Niino & Habe(MS) in Habe & Kosuge, 1964: 11. <strong>Recent</strong>. OffPhuket, Malaya Archipelago. 50-80 m. Nomennudum.† hexagonale, Dentalium Richardson, 1906: 581,pl. 45, fig. 1. Jurassic, Sinemurian-Pliensbachian.Gasworks, Gloucester, Engl<strong>and</strong>, United Kingdom.Remark: Engeser & Riedel (1992: 37) stated that“the features described <strong>and</strong> figured by Richardson(1906) do not coincide with the diagnosis <strong>of</strong> anyknown scaphopod genus”.hexagonum, Dentalium Gould, 1859: 166. <strong>Recent</strong>.Hong Kong [China]. Lectotype USNM 2053(designated by Johnson 1964: 88, as “holotype”);paralectotype MCZ 169184. Junior synonyms:Dentalium sexcostatum Sowerby, 1860; Dentaliumminus Boissevain, 1906 (fide Scarabino 1995: 218).Paradentalium hexagonum (Gould, 1859) (fideKira 1954: 80). Indo-Pacific: China to Australia.16-296 m. Remarks: the synonymization <strong>of</strong>Dentalium sexcostatum <strong>and</strong> Dentalium minus withthis <strong>species</strong> by Scarabino (1995) was not followedby Lamprell & Healy (1998). Both publicationsrejected the synonymization with Dentalium octangulatumDonovan, 1804 by Nomura (1938)hexagonum, Dentalium Van Ingen, 1905: 363, pl. A,fig. b; pl. B., figs 1, 2. Late Cretaceous, Senonian.Near Kerak, Jordan Valley, Syria. Syntype YPM35573 (fide White 1998: 4). Replacement name:Dentalium vaningeni n. nom.hexangulare, Dentalium Risso, 1826a: 125. Fossil.Near Nice, France. Nomen nudum.† hexapleuron, Dentalium Kaunhowen, 1897: 13,pl. 1, fig. 2. Cretaceous, Maestrichtian. Kunraed,Germany.hexaschistus, Cadulus (Polyschides) Boissevain, 1906:67, 68, pl. 6, fig. 53, text-figs 32, 32a. <strong>Recent</strong>.Ceram Sea, Siboga stn 178, 02°40’S, 128°37.5’E,835 m. Holotype ZMA 3.06.090. Siphonodentaliumhexaschistum (Boissevain, 1906) (fideScarabino 1995: 330). Indo-Pacific: Red Sea toNew Caledonia. 528-1740 m. Remark: the designation<strong>of</strong> ZMA 3.06.178 as the lectotype by Scarabino(1995: 330) is unnecessary, since Boissevain indicatedthat the <strong>species</strong> was described from a singlespecimen.hige, Pulsellum Habe, 1963: 273, 275, text-figs 47, 48.<strong>Recent</strong>. Zushi, Kanagawa Pref. Honshu, 350 m.Holotype <strong>and</strong> paratype in NSMT. Pulsellum higeHabe, 1963. Indo-Pacific: Japan. 5-350 m.Remarks: name for Siphonodentalium (Pulsellum)ozawai sensu Hirase (1931: 140) non Yokoyama,1926. Otuka (1935: 880) <strong>and</strong> Nomura & Hatai(1940: 74) reported this <strong>species</strong> from the <strong>Recent</strong>fauna <strong>of</strong> Japan.hillae, Dentalium (Dentalium) Lamprell & Healy,1998: 59, figs 53C, 54C, 57. <strong>Recent</strong>. SW <strong>of</strong> ScottReef, 410 km N <strong>of</strong> Broome, northern WesternAustralia, 14°39’S, 121°29’E, 500 m. HolotypeAMS C169776. Dentalium hillae Lamprell &Healy, 1998. Known from type locality only.hintoni, Dischides Lamprell & Healy, 1998: 167, 168,figs 174B, 175A, 177. <strong>Recent</strong>. 3 km NW <strong>of</strong> westside <strong>of</strong> Gillett Cay, Swain Reef, Queensl<strong>and</strong>,Australia, 80 m. Holotype AMS C174894;paratypes AMS C173611 (28). Dischides hintoniLamprell & Healy, 1998. Known from type localityonly. Remark: V. Scarabino (in litt. 5.XI.2001) suggestedthat the status <strong>of</strong> D. hintoni Lamprell &Healy, 1998 vis-à-vis Cadulus aratus Hedley, 1899requires further research.hirasei, Dentalium (Pictodentalium) formosum subsp.Kira, 1959: 105, pl. 40, fig. 11. <strong>Recent</strong>. SouthernJapan, 20 fms [37 m]. Nomen nudum in Kira (1954:80) (fide Bieler & Petit 1990: 141). Junior synonym<strong>of</strong> Pictodentalium formosum (Adams & Reeve,1850) (fide Scarabino 1995: 267).† hoernesi, Gadila Boettger, 1902: 182. Tertiary.Baden, near Vienna, Austria. Holotype SMFXIII.7a (fide Zilch 1934: 278, 279, pl. 22, fig. 25).Cadulus (Gadila) hoernesi (Boettger, 1902) (fideZilch 1934: 278, 279).honoluluensis, Cadulus Watson, 1879: 89, 90 [Watson1886: 17, pl. 2, fig. 10]. <strong>Recent</strong>. Reefs <strong>of</strong>fHonolulu, Hawaii, 40 fms [72 m]. HolotypeBMNH 1887.2.9.70. Gadila honoluluensis(Watson, 1879) (fide Pilsbry & Sharp 1898: 185).Known from type locality only.horikoshii, Fissidentalium Okutani, 1982: 1-3; figs 1-5. <strong>Recent</strong>. East <strong>of</strong> Kinkazan [Japan], Soyo-maru stnR3, 38°21.7’N, 143°25.6’E, 2930-3020 m.Holotype NSMT Mo.59950; paratypes NSMTMo.59951-52. Fissidentalium horikoshii Okutani,1982. Indo-Pacific: Japan to eastern Australia.1615-3500 m.hosoi, Dentalium Habe, 1963: 263, text-figs 4-6.<strong>Recent</strong>. Off Tosa Bay, Shikoku, “deep bottom”.Holotype <strong>and</strong> paratype in NSMT. Striodentaliumhosoi (Habe, 1963) (fide Habe 1964: 23). Indo-Pacific: Japan. 200-760 m.houbricki, Laevidentalium Scarabino, 1995: 295, 296,figs 100, 102d. <strong>Recent</strong>. New Caledonia, secteur dePoindimie, LAGON stn 835, 20°47’S, 165°17’E,135-150 m. Holotype MNHN; paratypes MNHN(7; 2 from stn 835; 5 from stn 858), AMS C201729(1; stn 858), NMNZ M268955 (1; stn 858),USNM 890861 (1; stn 858). Laevidentaliumhoubricki Scarabino, 1995. Indo-Pacific: Indonesiato New Caledonia. 135-220 m. Remark: V.Scarabino (in litt. 5.XI.2001) suggested that the status<strong>of</strong> L. arnoldi Lamprell & Healy, 1998 vis-à-visL. houbricki Scarabino, 1995 requires furtherresearch.houthalenense, Dentalium Goddeeris, 1978: 70, 72.Miocene. Belgium. Nomen nudum.† houthalenensis, Cadulus (Gadila) benoisti var.Goddeeris, 1977: 155-157, fig. 1. Miocene.Houthalen S<strong>and</strong>s, Wijshagen, Belgium. HolotypeZOOSYSTEMA • 2004 • 26 (4)599


Steiner G. & Kabat A. R.KBINW 5596. Gadila benoisti houthalenensis(Goddeeris, 1977) (fide Janssen 1984: 117, 118,pl. 42, figs 18a, 18b). Remark: Goddeeris (1978:70) listed Cadulus benoisti houthalensis <strong>and</strong>Dentalium houthalense MS as separate taxa.howardensis, Clavulites Girty, 1904. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.howchini, Dentalium (Paradentalium) Cotton &Ludbrook, 1938: 224, pl. 12, fig. 6. LowerPliocene. Abattoirs Bore, South Australia, Australia.Holotype SAM. Junior synonym <strong>of</strong> Dentalium(Dentalium) latesulcatum Tate, 1899 (fideLudbrook 1956: 1, 2).hungarica, Gadila Szöts, 1953 [ex Hantken MS]: 81[Hungarian], 203 [French]. Nomen nudum.Remark: under Cadulus (Gadila) pseudohungaricusn. sp., Szöts refered to “La G. hungarica Hantk. incoll. est assez abondante [...]”.hungerfordi, Dentalium (Fissidentalium) Pilsbry &Sharp, 1897: 84, pl. 6, fig. 83. <strong>Recent</strong>. Hong Kong.Lectotype BMNH 1881.2.1.1 (designated byScarabino 1995: 258, as “holotype”); paralectotypeBMNH 1881.2.1.2. Replacement name forDentalium compressum Sowerby, 1888, a juniorhomonym <strong>of</strong> Dentalium compressum Watson, 1879,Dentalium compressum Meyer, 1884, <strong>and</strong> Dentaliumcompressum d’Orbigny, 1850. Compressidentaliumhungerfordi (Pilsbry & Sharp, 1897)(fide Habe 1963: 260). Indo-Pacific: Indonesia toJapan. 50-1570 m.† hurupiensis, Cadulus Dell, 1952: 83, 84, fig. 18.Miocene, Tongaporutuan. Hurupi Beds, southernWairarapa, Palliser Bay, New Zeal<strong>and</strong>. HolotypeNMNZ M.5590. Cadulus (s.l.) hurupiensis Dell,1952 (fide Beu & Maxwell 1990: 424).huttoni, Dentalium Kirk, 1880: 306. <strong>Recent</strong>. Threespecimens “from the stomach <strong>of</strong> a trumpeter (Latrishecateia)”. Type locality not specified [althoughpresumably New Zeal<strong>and</strong>]. Lectotype NMNZM261 (designated by Dell 1957: 570); paralectotypesNMNZ M131453. Junior synonym <strong>of</strong>Antalis nana (Hutton, 1873) (fide Dell 1957: 570).Remark: Dell (1957: 570) stated the type materialwas obtained from a fish “probably caught nearWellington”.huttoni, Dentalium Bather, 1905: 540, figs 4-7.Carboniferous. New Zeal<strong>and</strong>. Junior homonym <strong>of</strong>Dentalium huttoni Kirk, 1880 (<strong>Recent</strong>). Replacementname: Dentalium batheri Finlay, 1927.hyalinum, Dentalium Philippi, 1846: 55. <strong>Recent</strong>.Mazatlan [Mexico]. Junior synonym <strong>of</strong> Graptacmesemistriata (Turton, 1819) (fide Pilsbry & Sharp1897: 91).hyalinum, Siphodentalium Brugnone, 1876: 21,fig. 32. Plio-Pleistocene. Monte Ficarazzi, Italy.Holotype UPMN (fide R. La Perna in litt.). Juniorsynonym <strong>of</strong> Polyschides olivi (Scacchi, 1835) (fidePilsbry & Sharp 1898: 171). Remarks: Brugnone(1878: 44) transferred this <strong>species</strong> to “Helonix” [sic,Helonyx]. Bellini (1909: 233) listed this as a variety<strong>of</strong> Siphonodentalium (Gadila) gadus.hyperhemileuron, Dentalium Verco, 1911b: 217, 218,pl. 26, figs 3, 3a. <strong>Recent</strong>. King George Sound, WAustralia, 12-35 fms [22-64 m]. Lectotype SAM D13726 (designated by Zeidler & Macphail 1978:383, as “holotype”); paralectotypes SAM D 16000(4) (incorrectly labelled co-types, fide Zeidler &Macphail 1978: 383), USNM 251993 (KingGeorge Sound, Western Australia, 22-26 m).Dentalium hyperhemileuron Verco, 1911. Indo-Pacific: Australia. 22-600 m. Remark: the otherparatypes (or paralectotypes) listed by Verco(200 from King George Sound, Western Australia,12-14 fms; 60 from King George Sound, 22-28 fms; 4 from King George Sound, 35 fms; 6 fromGeographe Bay, Western Australia, 15 fms; 4 fromGeographe Bay, 22 fms; 20 from <strong>of</strong>f Fremantle,Western Australia, 10-12 fms) were not located ornot associated with the catalog numbers listed.† ibergense, Dentalium Römer, 1855: 36, pl. 7, fig. 7.Devonian. Ibergs, near Grund, Harz Mountains,Germany.† ignotum, Dentalium Korobkov, 1955: 210, 211,pl. 3, figs 8-11. Middle Eocene. NorthernCaucasus, Russia.† iheringi, Dentalium Brunet, 1995: 48, pl. 5, fig. 1.Upper Miocene, Entrerriense Formation. BalizaPunta Flecha, Puerto Madryn, Chubut Province,Argentina. Holotype MPEF-PI-074.† ihungia, Dentalium Marwick, 1931: 158, pl. 18,figs 344, 345. Miocene, Clifdenian, Ihungia Series.Muddy Creek, Gisborne, New Zeal<strong>and</strong>. HolotypeNZGS TM4803; paratypes NZGS TM4804 (1),NZGS G 1237 (2) (fide Keyes 1972: 95).Dentalium ihungia Marwick, 1931 (fide Beu &Maxwell 1990: 424).† illinoiensis, Dentalium Worthen, 1883: 325. Carboniferous.Chester Limestone, Chester, Illinois, USA.Holotype Illinois State Collection No. 2488.imparcostata, Dentalium (Entalis “?”) Nechaev, 1897:124, pl. 10, figs 3, 4. Eocene. Near Saratov, Russia.Junior synonym <strong>of</strong> Dentalium rugiferum Koenen,1885 (fide Makarenko 1970: 113).imperforatum, Dentalium G. Adams, 1798. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.inaequale, Dentalium Bronn, 1831: 84, 85. Miocene.Italy. Junior synonym <strong>of</strong> Dentalium sexangulumGmelin, 1791 (fide Bernasconi 1996: 26). Juniorsynonym: Dentalum orsum Bonelli in Sismonda,1847 (fide Pilsbry & Sharp 1898: 205).inaequale, Dentalium Ryckholt, 1851: 67, 68, pl. 2,figs 41, 42. Carboniferous. Visé, Belgium. Juniorhomonym <strong>of</strong> Dentalium inaequale Bronn, 1831.Junior synonym <strong>of</strong> Plagioglypta ingens (de Koninck,1843) (fide Pilsbry & Sharp 1898: 231).† inaequicostata, Dentalium Seguenza, 1879: 117,pl. 11, figs 48, 48a. Miocene. Reggio, Calabria, Italy.600ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)inaequicostatum, Dentalium Dautzenberg, 1891:53 (1895: 370). <strong>Recent</strong>. Roussillon, France,Mediterrannean. Syntypes (<strong>of</strong> Dentalium alternans)MNHN. Replacement name for Dentalium alternansBucquoy, Dautzenberg & Dollfus, 1882, ajunior homonym <strong>of</strong> Dentalium alternans Chenu,1843. Objective junior synonym: Dentalium dollfusiCossmann, 1899, which is a junior homonym <strong>of</strong>Dentalium dollfusi Koenen, 1883. Junior homonym<strong>of</strong> Dentalium inaequicostatum Seguenza, 1879.Antalis inaequicostata (Dautzenberg, 1891) (genericplacement fide Pilsbry & Sharp 1897: 52).Eastern Atlantic, Mediterranean. 5-120 m.Remarks: although there is no valid junior synonymavailable, we prefer not to introduce yet anotherreplacement name for this <strong>species</strong> in order to limitthe nomenclatural damage suffered by this taxon.Antalis inaequicostata (Dautzenberg, 1891) is incurrent use for this Mediterranean <strong>species</strong>, whereasSeguenza’s name is not. Moreover, it remains to bedetermined whether Antalis inaequicostata isconspecific with the allied Atlantic Antalis novemcostata(Lamarck, 1818). Should this be the case,Lamarck’s junior synonym becomes an availablename for inaequicostata. Even if these <strong>species</strong> turnout to be distinct, the name inaequicostataDautzenberg can be preserved pursuant to ICZN(1999) article 23.9.1, which allows for homonymsthat have been widely used in the literature to bepreserved if the senior name has not been used since1899. Antale ininaequicostata [Ferrera Mortara etal., 1984: 305] is an incorrect subsequent spelling.inaequisculpta, Entalina Ludbrook, 1954: 111, 112,fig. 16. <strong>Recent</strong>. Gulf <strong>of</strong> Aden, John Murray Exp.stn 185, 13°48’06”N, 49°16’48”E-13°48’36”N,49°16’24”E, 2000 m. Holotype BMNH1952.3.25.5; paratypes BMNH 1952.3.25.6-BMNH 1952.3.25.19. Entalina inaequisculptaLudbrook, 1954. Indo-Pacific: Red Sea to Celebes.835-2198 m.incertula, Dentalium (Fustiaria) Sacco, 1897: 113,pl. 10, fig. 34 [as a nomen nudum in Sacco 1896: 98].Miocene. Montegibio, Niciola <strong>and</strong> Fossetta, Italy.Replacement name for Dentalium intermediumCoppi, 1876, a junior homonym <strong>of</strong> Dentalium intermediumHupé, 1854. Junior synonym <strong>of</strong> Gadilinajani (Hörnes, 1856) (fide Pavia 1991: 128, 129).† incertum, Dentalium Deshayes, 1825: 362, 363,pl. 17, fig. 17. Tertiary? Bordeaux <strong>and</strong> Maulette,near Houdan, Paris Basin, France. Referable toLaevidentalium (Le Renard 1995: 175; Le Renard& Pacaud 1995: 85), possibly a junior synonym <strong>of</strong>Dentalium nitens J. Sowerby, 1814 (fide Deshayes1864: 202).† incisissimum, Dentalium Meyer & Aldrich, 1886:40, pl. 2, fig. 1. Eocene <strong>and</strong> Oligocene. Wautubbee,Mississippi, USA. Lectotype USNM Paleobiology638724 (designated by Palmer & Brann 1965: 367as “holotype”).† incisum, Dentalium Chenu, 1843: pl. 6, figs 8, 9;1850: 4. Pliocene? Italy.incisus, Cadulus Bush, 1885: 471, pl. 45, fig. 20.<strong>Recent</strong>. Off Cape Hatteras, Albatross stn 2272, 15fms [27 m]. Lectotype USNM 44860 (designatedby Johnson 1989: 43); paralectotypes YPM 15865(2). Junior synonym <strong>of</strong> Polyschides tetraschistus(Watson, 1879) (fide Scarabino 1980: 12). Remark:Johnson (1989: 43) stated that there were “twoparatypes” but Bush only referred to two specimenstotal, so the second specimen in YPM 15865 musthave been added at a later date.incolor, Dentalium (Dentalium) aprinum Boissevain,1906: 10, pl. 4, figs 3-6. <strong>Recent</strong>. Indonesia,Anchorage <strong>of</strong>f Lirung, Salibabu Isl<strong>and</strong>, Siboga stn133, up to 36 m. Lectotype ZMA 3.06.005 (designatedby Scarabino 1995: 198); paralectotypesZMA 3.06.006 (3 in original description; 4 fideR. Moolenbeek & A. N. Van der Bijl in litt.23.VI.2000), ZMA 3.06.007 (1; anchorage nearSeget, west-entrance Selee (Galewo) strait, Sibogastn 163, 29 m). Junior synonym <strong>of</strong> Dentaliumaprinum Linnaeus, 1767 (fide Scarabino 1995:198).incrassatum, Dentalium J. Sowerby, 1814. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.incurvum, Dentalium Renier, 1804. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.indianorum, Dentalium pretiosum var. Carpenter,1864: 612, 648. <strong>Recent</strong>. Monterey (to) CatalinaIsl<strong>and</strong>, California [USA], 20 fms [37 m]. SyntypesUSNM 14790 (1), USNM 601274 (4), MCZ29203 (7). Junior synonym: Antalis denseliratumPilsbry & Sharp, 1898 [ex Carpenter MS].Synonym <strong>of</strong> Antalis pretiosa (Sowerby, 1860) (fidePilsbry & Sharp 1897: 45). Remark: Pilsbry &Sharp (1898: 250) published Carpenter’s manuscriptname, Antalis denseliratum, in synonymy withA. indianorum.indianum, Dentalium Girty, 1915: 135. Carboniferous,Pennsylvanian. Locality 2006, WewokaFormation, Wewoka quadrangle, Oklahoma, USA.Syntype USNM Paleobiology 121077 (labelled as“lectotype”). Junior synonym <strong>of</strong> Prodentalium semicostatum(Girty, 1911) (fide Yancey 1978: 308, byFirst Reviser action). Remark: syntype illustrated byGirty (1915: 146, pl. 25, figs 12, 12a).indica, Costentalina Chistikov, 1982b: 1318, 1319,pl. 1, figs 12-14, pl. 4, figs 4-7. <strong>Recent</strong>. IndianOcean, Vitjaz stn 4922 D, 6°54.2’N, 83°00.7’E,3980 m. Holotype IOM; paratypes IOM (2 fromstn 4922; 1 from stn 4721, 16°48.3’N, 62°20.3’E,3860 m; 2 from stn 4925, 11°42.5’N, 82°49.3’E,3428 m; 1 from stn 4934, 16°57’N, 86°59.2’E,2596 m; 1 from stn 5292, 5°07.3’N, 91°43.5’E,3860 m; 1 from stn 5304, 8°08.2’N, 95°21.7’E,2480 m; 1 from Akademik Kurtschatov stn 25-8,4°32.2’S, 63°18.3’E, 4440 m). Costentalina indicaZOOSYSTEMA • 2004 • 26 (4)601


Steiner G. & Kabat A. R.Chistikov, 1982. Indian Ocean: Red Sea toMadagascar <strong>and</strong> India. 2480-5285 m.indica, Wemersoniella Scarabino, 1995: 325, 326,figs 126, 129a. <strong>Recent</strong>. NE <strong>of</strong> MozambiqueChannel (Madagascar), RV Suroit, BENTHEDIstn CH87, 11°44’S, 47°35’E, 3716 m. HolotypeMNHN. Wemersoniella indica Scarabino, 1995.Known from the type locality only.indicum, Dentalium Chenu, 1843: pl. 3, fig. 11; 1850:4. <strong>Recent</strong>. Indian Ocean. Type material not located.Junior synonym <strong>of</strong> Laevidentalium eburneum(Linnaeus, 1767) (fide Pilsbry & Sharp 1897: 116).indicum, Dentalium (Antalis) entalis var. Boissevain,1906: 44, pl. 6, fig. 15. <strong>Recent</strong>. Indonesia, Sibogastn 159, 00°59.1’S, 129°48.8’E, 411 m. HolotypeZMA 3.06.051. Junior homonym <strong>of</strong> Dentaliumindicum Chenu, 1843. Replacement name: Antalisboissevainae Palmer, 1974; a junior synonym <strong>of</strong>Antalis tibana (Nomura, 1940) (fide Habe 1961:105). ). Remark: Scarabino (1995: 231) unnecessarilydesignated the single specimen as lectotype.indistinctum, Dentalium Fleming, 1825. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.inexpectatus, Bathoxiphus Scarabino, 1995: 315, 316,figs 117, 121c. <strong>Recent</strong>. New Caledonia, N NorfolkRidge, RV Jean Charcot, BIOCAL, stn DW46,22°53’S, 167°17’E, 570-610 m. Holotype MNHN;paratypes MNHN (1 from LAGON stn 830;10 from stn DW44), AMS C201732, NMNZM268952, USNM 890866. Bathoxiphus inexpectatusScarabino, 1995. Indo-Pacific: New Caledonia.410-570 m.† infans, Cadulus (Gadila) Tate, 1899: 266, 267,pl. 8, fig. 11. Miocene. Muddy Creek, Victoria,Australia. Holotype SAM T 1614. Cadulus (Gadila)infans Tate, 1899 (fide Ludbrook 1959: 148, pl. 1,fig. 6).infimus, Compressidens Scarabino, 1995: 368, 369,figs 173, 174d. <strong>Recent</strong>. Réunion, MD32stn DS109, 20°52’S, 55°06’E, 1050-1240 m.Holotype <strong>and</strong> 8 paratypes MNHN. Compressidensinfimus Scarabino, 1995. Indian Ocean. 1050-1240 m.inflata, Cadulus olivii var. Locard, 1898: 134, pl. 7,figs 13-15. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Type material not located.Synonym <strong>of</strong> Polyschides olivi (Scacchi, 1835).inflata, Cadulus propinquus var. Locard, 1898: 139.<strong>Recent</strong>. Bay <strong>of</strong> Biscay, Travailleur 1880 stn 2,1019 m; North <strong>of</strong> Spain, Travailleur 1880 stn 9,1190 m. Type material not located. Synonym <strong>of</strong>Cadulus propinquus G. O. Sars, 1878.inflexum, Dentalium Sowerby, 1903: 224, pl. 5,fig. 11. <strong>Recent</strong>. Tugela River Mouth (Natal), bearingnorth west by west, distant 3.5 miles, 14 fms[26 m]. Syntypes BMNH 1903.7.27.60, NMW.Antalis inflexa (Sowerby, 1903) (fide Scarabino1995: 236). Eastern Atlantic, Indian Ocean: SouthAfrica. 25-44 m. Remarks: Sowerby gave the measurements<strong>of</strong> the single specimen in BMNH labelled“holotype”, but the original description did notspecify a holotype. Barnard (1963b: 351) listed“cotypes” in SAFM A5470, which were not mentionedby Giles & Gosliner (1983).† infortunatum, Dentalium Pilsbry & Sharp, 1898:255. Cretaceous, Gaultian. Folkestone, Engl<strong>and</strong>,United Kingdom. Replacement name forDentalium gardneri Sharp & Pilsbry in Pilsbry &Sharp, 1898, a junior homonym <strong>of</strong> Dentalium gardneri(Holzapfel, 1888).infractum, Dentalium Odhner, 1931: 1-3, pl. 1, figs 1-3, 8, 10, 12, pl. 2, 15-17, 22, 23. <strong>Recent</strong>. Coast <strong>of</strong>Uruguay, Swedish Antarctic Exp., stn 1, 33°00’S,51°10’W, 80 m. Syntypes SMNH 1554 (fideS<strong>and</strong>berg & Warén 1993: 125). Junior synonym:Dentalium (Antalis) elegantulum Penna-Neme,1974 (fide Scarabino 1975: 184). Antalis infracta(Odhner, 1931) (fide Penna-Neme 1974: 110;Scarabino 1975: 184). Western Atlantic: Brazil toArgentina. 20-190 m.† infraeocaenicanum, Pulsellum Cossmann, 1907a:204, 205, pl. 9. Paleocene, Thanetian. Noailles,near Paris, France. Remark: a valid <strong>species</strong> <strong>of</strong>Compressidens (fide Le Renard 1995: 175; LeRenard & Pacaud 1995: 86).† infraoligocaenicum, Compressidens Welle, 1997: 95,96, pl. 13, figs 17-19. Lower Oligocene, Ratheim-Schichten. Schacht 8, Bergwerksgesellschaft SophiaJacoba, near Erkelenz (north <strong>of</strong> Aachen), Germany.Holotype GPIM D.8-9-1.199; paratypes GPIM (5).infundibulum, Dentalium entalis var. Jeffreys, 1869:197. <strong>Recent</strong>. Loch Fyne [Scotl<strong>and</strong>]. Type materialnot located. Synonym <strong>of</strong> Antalis entalis (Linnaeus,1758).† infundibulum, Pulsellum Cossmann & Peyrot, 1917:181, pl. 1, figs 60-62, pl. 2, fig. 16. Miocene,Helvetian <strong>and</strong> Burdigalian. Salles (Le Minoy), Saint-Étienne-d’Orthe, <strong>and</strong> Saint-Paul-lès-Dax, France.† ingens, Dentalium de Koninck, 1843: 317, pl. 22,figs 2a-2c. Carboniferous. Visé, Belgium. Juniorsynonym: Dentalium inaequale Ryckholt, 1851 (fidePilsbry & Sharp 1898: 231). Plagioglypta ingens (deKoninck, 1843) (fide Waterhouse 1990: 198, 199).ingrata, Spadentalina Scarabino, 1995: 310-312,figs 111e-g, 113, 115f. <strong>Recent</strong>. Loyalty Isl<strong>and</strong>s,Pacific, RV Alis MUSORSTOM 6, stn DW444,20°54’S, 167°18’E, 300 m. Holotype MNHN;paratypes MNHN (1 from DW444; 1 fromDW253; 2 from DW282; 1 from DW334; 1 fromDW344; 3 from DW446; 1 from Vauban 34; 3from Vauban 40), AMS C201731 (1; DW282),NMNZ M268951 (1; Vauban 40), USNM 890863(1; DW280). Spadentalina ingrata Scarabino,1995. Indo-Pacific: Indonesia to New Caledonia.182-440 m.ininaequicostata, see inaequicostatum.innumerabile, Dentalium (Episiphon) Pilsbry & Sharp,1897: 119, pl. 18, figs 6-8. <strong>Recent</strong>. Type locality602ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)not specified. Localities <strong>of</strong> syntypes: ANSP 35564(Panama Bay, USFC stn 2799, 8°44’N, 79°09’W,29.5 fms [54 m]), USNM 96983 (Panama Bay,Albatross stn 2803, 8°27’N, 79°35’W, 26 fms[47 m]), USNM 97024 (Off Lower California nearCerros Isl<strong>and</strong>, Albatross stn 2838, 28°12’N,115°09’W, 44 fms [80 m]), USNM 107697 <strong>and</strong>USNM 203131 (Panama Bay, Albatross stn 2799,8°44’N, 79°09’W, 54 m), USNM 122770 (PanamaBay, Albatross stn 2803, 8°27’N, 79°35’W, 26 fms[47 m]), USNM 122804 (Panama Bay, Albatrossstn 2805, 7°56’N, 79°41’30W, 51 fms [93 m]),USNM 122811 (<strong>of</strong>f Lower California near SantaMargarita Isl<strong>and</strong>, Albatross stn 2831, 24°32’N,111°59’W, 12 fms [22 m]), USNM 203132 (OffGuaymas, Albatross stn 3037, 27°45’N, 110°45’W,20 fms [37 m]), SBMNH 35012 (Panama Bay,USFC stn 2799, 8°44’N, 79°09’W, 54 m). Juniorsynonym: Dentalium (Rhabdus) cedrosense Hertlein& Strong, 1951 (fide Keen 1971: 888). Episiphoninnumerabile (Pilsbry & Sharp, 1897). EasternPacific: California to Panama. 22-93 m.† inopinatum, Dentalium Mayer-Eymar, 1887: 52,104, pl. 4, fig. 7. Eocene, Bartonian, Auversian.Schimberg, near Thun, Switzerl<strong>and</strong>.inornatum, Dentalium McCoy in Griffith, 1844. Nota scaphopod. See Other taxa described as <strong>Scaphopoda</strong>.inornatum, Dentalium Wade, 1926: 100, 101, pl. 33,figs 2, 3. Upper Cretaceous, Ripley Formation.Dave Weeks property on Coon Creek, McNairyCounty, Tennessee, USA. Type material USNMPaleobiology 32821. Junior homonym <strong>of</strong>Dentalium inornatum McCoy in Griffith, 1844.Replacement name: Dentalium demersum Pilsbry,1927. Junior synonym <strong>of</strong> Dentalium leveStephenson, 1923 (fide Stephenson 1941: 253, 254).insolitum, Dentalium E. A. Smith, 1894: 168, 169,pl. 4, fig. 17, 17a. <strong>Recent</strong>. Bay <strong>of</strong> Bengal [India], 597fms [1090 m]. Lectotype BMNH 1894.9.11.9 (designatedby Ludbrook 1954: 109); paralectotypesBMNH 1952.3.25.83-93 (not located), ZSI 5412.Junior synonym: Dentalium stapes Boissevain, 1906(fide Habe 1964: 32). Gadilina insolita (E. A.Smith, 1894) (fide Plate 1908a: 353). Indo-Pacific:Red Sea to New Caledonia <strong>and</strong> Japan. 625-1328 m.† insuranum, Dentalium Emerson, 1954: 183, 184.Oligocene, Priabonian. Locality 813, Saädang,Celebes, Indonesia. Replacement name for Dentaliummartini Dollfus, 1915 non Whitfield, 1882.† interbinarium, Dentalium sexangulum “mut.”Cossmann & Peyrot, 1917: 161, pl. 1, figs 4-7.Miocene. Saubrigues, France.intercalatum, Dentalium Gould, 1859: 166. <strong>Recent</strong>.China Seas. Lectotype USNM 24183 (designatedby Johnson 1964: 93, as “holotype”); paralectotypeMCZ 169197 [ex NYSM 148). Junior synonym:Dentalium bednalli Pilsbry & Sharp, 1897 (fideVerco 1904; fide Lamprell & Healy 1998: 59).Paradentalium intercalatum (Gould, 1859) (fideHabe & Kosuge 1964: 1). Indo-Pacific: China toAustralia. 0-668 m.intercalatum, Dentalium Wade, 1926: 100, pl. 33,figs 18, 19. Upper Cretaceous, Ripley Formation.Dave Weeks property on Coon Creek, McNairyCounty, Tennessee, USA. Type material USNMPaleobiology 32829. Junior homonym <strong>of</strong>Dentalium intercalatum Gould, 1859. Replacementname: Dentalium wadei Pilsbry, 1927.intercosicillata, see intercosticillata.intercostatum, Dentalium Boissevain, 1906: 14, pl. 6,fig. 4, text-fig. 11. <strong>Recent</strong>. Indonesia, Ceram Sea,Siboga stn 178, 02°40’S, 128°37’E, 835 m.Lectotype ZMA 3.06.010 (designated by Scarabino1995: 306 as “holotype”); paralectotypes ZMA3.06.011 (2; stn 314, 694 m; lost, fide R. Moolenbeek& A. N. Van der Bijl in litt. 23.VI.2000).Junior synonyms: Dentalium siberutense Plate,1908a (fide Scarabino 1995: 305); Dentalium nivosumKuroda & Kikuchi, 1933 (fide Habe 1964: 40);Dentalium tugaruense Nomura & Hatai, 1940 (fideHabe 1964: 40). Entalinopsis intercostata (Boissevain,1906) (fide Habe 1963: 272). Indo-Pacific:Indonesia to Japan. 50-1500 m. Remark: Lamprell& Healy (1998: 48) considered Dentalium intercostatumto be a junior synonym <strong>of</strong> Dentalium thetidisHedley, 1903 (currently in the genusStriodentalium), thus rejecting Habe’s assignment <strong>of</strong>this <strong>species</strong> to Entalinopsis.† intercosticillata, Dentalium michelotti var. Sacco,1897: 97, pl. 7, figs 87, 88 [as a nomen nudum inSacco 1896: 97]. Pliocene. Italy (various localities).Syntypes MIGT BS.106.01.036, BS.106.01.037(fide Ferrero Mortara et al. 1984: 302). Dentaliummichelotti var. intercosicillata [Pilsbry & Sharp,1898: 209] is an incorrect subsequent spelling.intermedia, Dentalium caudani var. Locard, 1898: 105.<strong>Recent</strong>. Type locality not specified [North Atlantic].Syntype MNHN. Synonym <strong>of</strong> Dentalium caudani,a junior synonym <strong>of</strong> Fissidentalium c<strong>and</strong>idum(Jeffreys, 1877).† intermedium, Dentalium Hupé, 1854: 276, pl. 2,fig. 9. Tertiary. Topocalma, Colchagua Province,Chile.intermedium, Dentalium R. Hörnes, 1875: 364, 365[32, 33], pl. 10, figs 16, 17. Miocene. Ottnang,northern Austria. Syntypes GBA 1875/01/0043 (3).Junior homonym <strong>of</strong> Dentalium intermedium Hupé,1854; junior synonym <strong>of</strong> Antalis bouei (Deshayes,1825) (fide Pilsbry & Sharp 1898: 200).intermedium, Dentalium Coppi, 1876: 203. Miocene.Montegibio, Niciola <strong>and</strong> Fossetta, Italy. Juniorhomonym <strong>of</strong> Dentalium intermedium Hupé, 1854;replacement name: Dentalium (Fustiaria) incertulaSacco, 1897, a junior synonym <strong>of</strong> Gadilina jani(Hörnes, 1856) (fide Pavia 1991: 128, 129).† interruptum, Dentalium Gmelin, 1791: 3739 [citingSchröter 1784a: 530 n. 10]. Miocene. Piedmont,ZOOSYSTEMA • 2004 • 26 (4)603


Steiner G. & Kabat A. R.Italy. Antalis interrupta (Gmelin, 1791) (fide Pavia1991: 115-120, pl. 2, figs 1-4, table 1; text-figs 2-5).interstriatum, Dentalium Sowerby, 1860: 102, pl. 223,fig. 7. <strong>Recent</strong>. Isl<strong>and</strong> <strong>of</strong> Bohol, Philippines.Syntypes BMNH 1950.11.2.42-44 (3) (incorrectlylabelled “holotype” <strong>and</strong> “paratypes”). Junior synonym<strong>of</strong> Dentalium aprinum Linnaeus, 1767 (fideHabe 1964: 6, 7).† intertrinarium, Dentalium sexangulum var.Cossmann & Peyrot, 1917: 162, pl. 1, figs 8-11.Miocene, Helvetian. Salies-de-Béarn, France.intesi, Dentalium (Antalis) Nicklès, 1979: 49, 50, textfig.7, pl. 1, fig I. <strong>Recent</strong>. Gonzagueville, IvoryCoast, ORSTOM 1966-67 stn 17, 35 m. HolotypeMNHN; paratypes (1 from ORSTOM 1966-67stn 18, Vridi, 20 m; 1 from ORSTOM 1966-67stn 19, Jacqueville, 30 m) not located in MNHN,not in SAFM (fide Giles & Gosliner 1983). Antalisintesi (Nicklès, 1979). Eastern Atlantic: Africa,Ivory Coast, Ghana. 20-35 m.intestiniforme, Dentalium Hanley, 1860. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† intumescens, Gadus Koenen, 1885: 72, 73, pl. 3,fig. 16. Paleocene. Denmark. Remark: a valid<strong>species</strong> <strong>of</strong> Cadulus (fide Makarenko 1970: 115).invalidum, Dentalium Emerson, 1954: 184. <strong>Recent</strong>.Jamaica. Holotype (<strong>of</strong> regulare Henderson) USNM107887. Replacement name for Dentalium(Dentalium) laqueatum regulare Henderson, 1920non Ahlburg, 1906, non E. A. Smith, 1903.Dentalium invalidum Emerson, 1954. WesternAtlantic, Caribbean. 78-255 m.inversum, Dentalium Deshayes, 1825: 370, 371 [50,51 in reprint], pl. 16, fig. 21. <strong>Recent</strong>. Type localitynot specified. Syntypes MNHN. Graptacme inversa(Deshayes, 1825) (fide Pilsbry & Sharp 1897: 95).Eastern Atlantic: Africa, Senegal (Nicklés 1979:55), ? Gulf <strong>of</strong> California (Keen 1958: 238, 239;“accidently omitted” in Keen 1971: 886, fide Keen& Coan 1975: 48). No data on bathymetric distributionavailable.inversum, Dentalium (Fustiaria) Cossmann, 1915: 7[ex Rutot MS]. Paleocene, Montian. Puits Coppée,Belgium. Junior homonym <strong>of</strong> Dentalium inversumDeshayes, 1825. Tentative junior synonym <strong>of</strong>Fustiaria montense (Briart & Cornet, 1887)(n. syn.). Remark: Cossmann (1915: 7) indicatesthat the only difference between the single specimen<strong>of</strong> inversum <strong>and</strong> montense is the position <strong>of</strong> theslit. We, therefore, prefer to place inversumCossmann in synonymy with Fustiaria montenseinstead <strong>of</strong> introducing a replacement name.iota, Cadulus (Gadila) Henderson, 1920: 134, 135, pl.19, fig. 2. <strong>Recent</strong>. Mayaguez Harbor, Puerto Rico,USBF stn 6062, 25 fms [46 m]. Holotype USNM161581. Gadila iota (Henderson, 1920). WesternAtlantic, Caribbean. 46-146 m.† iowaensis, Coleolus “?” James, 1890: 354, 355.Upper Ordovician, Cincinnatian. Near Graf,Dubuque County, Iowa, USA. Lectotype USNMPaleobiology 34628 (designated by Bretsky &Bermingham 1970: 911, pl. 131, fig. 2); numerousparalectotypes USNM Paleobiology 34628.Remark: transferred to the <strong>Scaphopoda</strong> by Bretsky& Bermingham (1970), as a valid <strong>species</strong> <strong>of</strong>Plagioglypta.irregulare, Dentalium Seguenza, 1879: 275, pl. 16,figs 33, 33a. Pliocene, Astian stage. Gallina, Reggio,Calabria, Italy. Junior homonym <strong>of</strong> Dentaliumirregularis Hutton, 1873; replacement name:Dentalium subirregulare Pilsbry & Sharp, 1898.irregularis, Dentalium Risso, 1826a: 125; 1826b: 400.Fossil. La Trinité, France. Nomen nudum.† irregularis, Dentalium Hutton, 1873b: 1. UpperMiocene, Pareora Formation. Kanieri <strong>and</strong> Awamoa,New Zeal<strong>and</strong>. Lectotype NZGS TM4805 (designatedby Suter 1914: 32). Dentalium irregulareHutton, 1873 (fide Beu & Maxwell 1990: 424).Remark: Fleming (1955: 1059) <strong>and</strong> Keyes (1972:95) determined that Suter had erroneously synonymizedD. irregulare with D. mantelli Zittel,1864.isaotakii, Siphonodentalium Habe, 1953: 299. <strong>Recent</strong>.Tokyo Bay, Honshu, Japan. Holotype NSMT54817; paratypes NSMT 54818, AMNH 148421(6). Siphonodentalium isaotakii Habe, 1953. Indo-Pacific: Japan.ivanovi, Dentalium Palmer, 1974b: 125. Carboniferous.Near Moscow, Russia. Replacement namefor Dentalium striatum Ivanov, 1926 non Born,1780. Remark: as Ivanov’s taxon was not validlydescribed, there was no need to rename it.ivanovi, Epirhabdoides Steiner, 1999: 153-155, figs 2-6. <strong>Recent</strong>. Vitjaz stn 24-3578, 38°35’N, 142°53’E,Japan Trench, 1641 m. Holotype MMSU Lb-23 ;paratypes MMSU Lb-24 (2), Lb-25 (4), MNHN(2), BMNH 1996379 (2). Epirhabdoides ivanoviSteiner, 1999. Known from the type locality only.† ixhuatlanensis, Dentalium Alencaster-Ibarra, 1950:223, 224, fig. 2. Miocene. Kilometer 22 on railroadSE <strong>of</strong> Ixhuatlán, Vera Cruz, Mexico. HolotypeColección Paleontológica, Gerencia de Exploración,Petróleros Mexicanos, PM 2003.† jacksonense, Dentalium (Antalis) mississippiense var.Palmer, 1947: 212, 213, pl. 26, figs 20-22. Eocene,Jackson Formation. Moodys Branch marl <strong>and</strong>Lower Yazoo clay, Mississippi, USA. Holotype PRI4477; paratypes PRI 4478, PRI 4479.† jacksonensis, Cadulus Meyer, 1885: 462; 1886: 65,pl. 3, figs 8-8b. Eocene. Jackson, Mississippi, USA.Syntype USNM Paleobiology 638704. Remark:Pilsbry & Harbison (1933: 117) suggested thatCadulus newtonensis Meyer & Aldrich, 1886 was ajunior synonym <strong>of</strong> this <strong>species</strong>; however, Palmer &Brann (1965: 362) listed both taxa as valid.jaeckeli, Siphonodentalium Scarabino, 1995: 332,figs 134, 135a. <strong>Recent</strong>. West Indian Ocean, Madagascar,West Banc du Leven, BENTHEDI stn604ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)DS03, 12°35.5’S, 47°38.2’E, 1150-1100 m.Holotype <strong>and</strong> 1 paratype MNHN. Siphonodentaliumjaeckeli Scarabino, 1995. Known fromthe type locality only.† jaegeri, Suevidontis Engeser, Riedel & B<strong>and</strong>el, 1993:94, 95, pl. 3, figs 4-10. Late Middle Jurassic,Middle Callovian. Plettenberg, near Dotternhausen,c. 40 km south <strong>of</strong> Stuttgart, Germany.Holotype GPIMH 2834; paratypes GPIMH 2835-2839.jaffaensis, Dentalium (Fissidentalium) Cotton &Ludbrook, 1938: 221, 222, pl. 12, fig. 5. <strong>Recent</strong>.Off Cape Jaffa, South Australia, 90 fms [165 m].Holotype SAM D 13337. Laevidentalium jaffaensis(Cotton & Ludbrook, 1938) (fide Lamprell &Healy 1998: 113). Indian <strong>and</strong> Southern Oceans:S <strong>and</strong> SW Australia. 75-300 m.jamaicense, Dentalium (Fissidentalium) meridionale var.Henderson, 1920: 62, pl. 9 fig. 4. <strong>Recent</strong>. NearJamaica, USBF stn 2140, 966 fms [1763 m].Holotype USNM 103519. Junior synonym <strong>of</strong>Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877) (fideScarabino 1994: 307).jamiesoni, Episiphon Lamprell & Healy, 1998: 131,132, figs 135A, B, 136A, B, 137. <strong>Recent</strong>. OffCairns, northern Queensl<strong>and</strong>, Australia,17°50.67’S, 147°18.2’E, 703 m. Holotype QMMO45835; paratypes QM MO17730 (6), QMMO17950 (2), AMS C173422 (26; <strong>of</strong>f Cairns,Queensl<strong>and</strong>, 17°09.42-36’S, 146°42.24’-42.00’E,668 m), AMS C169798 (2; NE <strong>of</strong> Sydney, NewSouth Wales, 33°33’S, 152°05’E, 750 m), AMSC173306 (8; E <strong>of</strong> S<strong>and</strong>y Cape, Queensl<strong>and</strong>,24°43.5-8’S, 153°33.4-33.3’E, 604 m). Episiphonjamiesoni Lamprell & Healy, 1998. Indo-Pacific:northern <strong>and</strong> eastern Australia. 192-770 m.Remark: V. Scarabino (in litt. 5.XI.2001) suggestedthat the status <strong>of</strong> E. jamiesoni Lamprell & Healy,1998 vis-à-vis Gadilina insolita (E.A. Smith, 1894)requires further research.† jani, Dentalium Hörnes, 1856: 657, 658, pl. 50,fig. 37. Miocene. Baden, Wiener Becken, Austria.Lectotype NMW 1846.37.540 (designated by Pavia1991: 129); paralectotypes NMW (11). Junior synonyms:Dentalium intermedium Coppi, 1876,Fustiaria emersoni Caprotti, 1979 (fide Pavia 1991:128). Gadilina jani (Hörnes, 1856) (fide Pavia1991: 128-130).japonicum, Dentalium Dunker, 1877: 68. <strong>Recent</strong>.Japan. Holotype ZMB 101995. Junior synonym <strong>of</strong>Dentalium octangulatum Donovan, 1804 (fideHabe 1964: 8).japonicum, Siphonodentalium Habe, 1960: 294.<strong>Recent</strong>. Tomioka, Amakusa Isl<strong>and</strong>, Kyushu, Japan.Holotype NSMT 54819; paratypes NSMT 54820,AMNH 148416 (3). Siphonodentalium japonicumHabe, 1960. Indo-Pacific: Japan. 30-300 m.Remark: this name with “sp.nov.” also appears inHabe & Kikuchi (1960: 30) as nomen nudum.javanum, Dentalium Sowerby, 1860: 102, pl. 233,fig. 12. <strong>Recent</strong>. Malacca [Java], 22 m. LectotypeBMNH 1993006/1 (designated by Lamprell &Healy 1998: 42); paralectotypes BMNH1993006/2-3 (2). Dentalium javanum Sowerby,1860. Indo-Pacific: India to Australia <strong>and</strong> NewGuinea. 4-150 m.† javense, Dentalium Palmer, 1974b: 124. Tertiary.Ngembak, Java, Indonesia. Syntypes RGM 7291(6) (fide Van den Hoek Ostende et al. 2002: 112).Replacement name for Cadulus martini Finlay,1927 non Dentalium martini Whitfield, 1882.Finlay’s name is itself a replacement name forDentalium compressum Martin, 1885 non d’Orbigny,1850. Palmer (1974b: 124) claimed thatFinlay’s <strong>species</strong> was actually referable to Dentalium,so it was a junior secondary homonym <strong>of</strong>Dentalium martini Whitfield, 1882.jeanae, Dentalium (Dentalium) Lamprell & Healy,1998: 46, figs 37D, 38F, 42. <strong>Recent</strong>. 37 km E <strong>of</strong>Lady Musgrave Isl<strong>and</strong>, Queensl<strong>and</strong>, Australia,23°44’S, 152°49’E, 348-357 m. Holotype AMSC174889; paratype AMS C305472. Dentaliumjeanae Lamprell & Healy, 1998. Indo-Pacific:Queensl<strong>and</strong>, Australia. 348-357 m.jeffereysi, see jeffreysi Gardner, 1878.† jeffreysi, Dentalium Gardner, 1878: 61, 62, pl. 3,figs 26-33. Cretaceous, Gaultian. Folkestone,Cambridge <strong>and</strong> Atherfield, Engl<strong>and</strong>, UnitedKingdom. Dentalium jeffereysi [Yao 1990: 31, 32] isan incorrect subsequent spelling.jeffreysi, Helonyx Monterosato, 1875: 20. <strong>Recent</strong>.Mediterranean <strong>and</strong> North Atlantic, 91-456 m.Type material not located. Junior synonym:Helonyx subfusiformis var. abyssicola Monterosato,1875 (fide Pilsbry & Sharp 1898: 163, by FirstReviser action, according to ICZN 1999: article24). Cadulus jeffreysi (Monterosato, 1875) (fidePilsbry & Sharp 1898: 163). Eastern Atlantic:Norway to St Helena; Mediterranean. 90-2200 m.Remark: this <strong>species</strong> was misidentified as Cadulussubfusiformis M. Sars by Jeffreys (1869: 196).jelli, Dentalium (Dentalium) Lamprell & Healy, 1998:25, 26, figs 9F, 10F, 18, 19. <strong>Recent</strong>. Off Cairns,northern Queensl<strong>and</strong>, Australia, 17°50.67’S,147°18.2’E, 703 m. Holotype QM MO45837;paratypes QM MO17766 (9), AMS C171341 (2),MV F69579 (7; 26°10’S, 153°13’E, southernQueensl<strong>and</strong>, Australia). Dentalium jelli Lamprell &Healy, 1998. Indo-Pacific: Queensl<strong>and</strong>, Australia.703 m (depth range not given in original description).johnsoni, Dentalium (Episiphon) Emerson, 1952a: 5, 6,pl. 1, fig. 2. <strong>Recent</strong>. Off Puerto Rico, Johnson-Smithsonian Deep-Sea Exp. stn 25, 18°32’15”N,66°22’10”W, 240-300 fms [438-548 m]. HolotypeUSNM 603543; paratypes USNM 429714,AMNH 148289 (2). Junior synonym <strong>of</strong> Episiphondidymum (Watson, 1879) (fide Scarabino 1975:186).ZOOSYSTEMA • 2004 • 26 (4)605


Steiner G. & Kabat A. R.† jonkeri, Dentalium Martin, 1885: 188, pl. 10,fig. 187; 1887: 188, pl. 10, fig. 187. Miocene/Pliocene. Tambak Batu, Ngembak <strong>and</strong> Batavia(Jakarta), Java, Indonesia; Kassi Marinu, Fialarang,Timor. Syntypes RGM 7259 (2), RGM 7258 (1),RGM 7260 (1) (fide Van den Hoek Ostende et al.2002: 111)† juliana, Dentalium sulcosum var. Ihering, 1907: 224.Tertiary. San Julián, Canada de los Artilleros,Patagonia, Argentina. Dentalium sulcosum julianumIhering, 1907 (fide Ihering 1914: 67). Type materialnot located.† junghuhni, Dentalium Martin, 1880: 87, 88, pl. 12,fig.11. Miocene. Locality R, Tji-Karang, E <strong>of</strong>Tijieri, Kendeg district, Sukapura Regency, Java,Indonesia. Holotype RGM 7252 (fide Van denHoek Ostende et al. 2002: 111). Remark: Dentaliumjunghuhni – Boettger 1882 <strong>and</strong> Dentaliumjunghuhni – Noetling 1901 were subsequentlydescribed as Dentalium nanggulanense <strong>and</strong>Dentalium noetlingi, respectively, by Martin (1915).jungii, Dentalium d’Orbigny, 1852. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† juvenis, Cadulus Meyer, 1886: 66, pl. 3, fig. 4.Eocene, Jackson stage. Moody’s Branch Marl,Jackson, Mississippi, USA. Lectotype USNMPaleobiology 638814 (designated by Palmer &Brann 1965: 362 as “holotype”). Cadulus (Gadila)juvenis Meyer, 1886 (fide Hodgkinson 1974: 16,17).† kahirahense, Dentalium Cossmann, 1901 [“1900”]:188, 189. Middle Eocene. Ouadi Abou Choucha,near Hélouan, Egypt. Junior synonym: Dentaliumpraecursor Mayer-Eymar, 1904 (fide Oppenheim1906: 214, 215).† kahleri, Dentalium Gugenberger, 1934: 44, 45,pl. 1, fig. 16. Triassic. N <strong>of</strong> Launsdorf, Austria.Remark: nomen nudum in Gugenberger 1933a:102; 1933b: 184.kaiyomaruae, Polyschides Okutani, 1975: 77, 78, pl. 3,fig. 5. <strong>Recent</strong>. North Pacific: Kaiyo-Maru stn 57,29°49’-28°48’N, 147°09’E, 6200 m. HolotypeNSMT Mo.69591 (fide Anonymous 1996: 82, pl.5, fig. 9). Polyschides kaiyomaruae Okutani, 1975.Known from the type locality only.kanakorum, Striodentalium Scarabino, 1995: 241,figs 48, 52c. <strong>Recent</strong>. New Caledonia, Coral Sea,RV Coriolis CHALCAL 2, stn DW74, 24°40’S,168°38’E, 650 m. Holotype MNHN <strong>and</strong> paratypes(3; 1 each from DW72, DW73, <strong>and</strong> DW197).Striodentalium kanakorum Scarabino, 1995. Indo-Pacific, New Caledonia. 527-650 m.† kansasense, Dentalium (Paleodentalium) Gentile,1974: 1213-1215, text-fig. 1a-1e. Carboniferous,Pennsylvanian, Missourian Series. 2.5 miles S <strong>of</strong>Stanley, Wy<strong>and</strong>otte Limestone, Johnson County,Kansas, USA. Holotype University <strong>of</strong> MissouriKansas City MS 1; paratypes University <strong>of</strong> MissouriKansas City MS 2, MS 3, MS 4.† kapuaensis, Polyschides Maxwell, 1992: 188, pl. 9,figs e, f. Eocene. NZGS locality GS11200 WaihaoRiver, near McCulloch’s Bridge, South Canterbury,New Zeal<strong>and</strong>. Holotype NZGS TM7233; paratypeNZGS (1).† karreri, Dentalium Hörnes, 1875: 365 [33], pl. 10,fig. 18. Miocene. Ottnang, northern Austria.Syntypes GBA 1875/01/0045 (2). Remark: Sacco(1897: 114) <strong>and</strong> Pilsbry & Sharp (1898: 234) listedthis name, without bibliographic reference, as a juniorsynonym <strong>of</strong> Entalina tetragona (Brocchi, 1814).katchekense, Dentalium Fischer-Piette & Nicklès,1946: 70, 71, pl. 1, fig. 4, text-figs 23-25. <strong>Recent</strong>.Katchek, Îles Tristao, Guinea. Holotype <strong>and</strong>paratypes MNHN. Dentalium katchekense Fischer-Piette & Nicklès, 1946. Eastern Atlantic: westernAfrica. 16-150 m.kathwayae, Dentalium (Dentalium) Lamprell &Healy, 1998: 38, 39, figs 29D, 30D, 34. <strong>Recent</strong>.26 km NE <strong>of</strong> North Reef, Queensl<strong>and</strong>, Australia,23°08.4’S, 152°12.3’E, 115 m. Holotype AMSC169943. Dentalium kathwayae Lamprell &Healy, 1998. Known from the type locality only.katowense, Dentalium Brazier, 1877: 56. <strong>Recent</strong>.Katow, New Guinea, 8 fms [15 m]. Holotype AMSC170758. Paradentalium katowense (Brazier,1877) (fide Scarabino 1995: 222). Indo-Pacific:New Guinea, Australia. 0-100 m.kawamurai, Fissidentalium Kuroda & Habe in Habe,1961: 106, pl. 47, fig. 14. As a nomen nudum inHabe (1953: 293). <strong>Recent</strong>. Ashizurimisaki, KochiPref., Shikoku, Japan, 100-200 m. Types inNSMT. Fissidentalium kawamurai Kuroda &Habe in Habe, 1961. Indo-Pacific: Japan. 100-400 m.† kentuckyensis, Rhytiodentalium Pojeta & Runnegar,1979: 532-535, pl. 1, figs 11-17, pl. 2, figs 1-13.Upper Middle Ordovician, Shermanian, LexingtonLimestone. USGS locality 4073-CO, along ShryockFerry Road, 1.5 miles SW <strong>of</strong> Grier Creek Churchnear Milner, Woodford County, Kentucky, USA.Holotype USNM Paleobiology 254392; paratypesUSNM Paleobiology 254393 (22), 254394-254400(various USGS localities near Lexington, Kentucky).Remark: Engeser & Riedel (1996: 123) <strong>and</strong>Yochelson & Holl<strong>and</strong> (2004: 98) express doubtsabout the scaphopod nature <strong>of</strong> this <strong>fossil</strong> <strong>species</strong>.kermadecae, Chistikovia Scarabino, 1995: 327, 328,figs 128, 129d, e. <strong>Recent</strong>. New Zeal<strong>and</strong>, KermadecTrench, Galathea stn 664, 36°34’S, 178°57’W,4510-4570 m. Holotype ZMUC; paratypesZMUC (4), MNHN (1; Galathea stn 665).Chistikovia kermadecae Scarabino, 1995. NewZeal<strong>and</strong>. 2470-4570 m.kessneri, Dentalium (Dentalium) Lamprell & Healy,1998: 31, figs 20D, 21D, 26. <strong>Recent</strong>. NE <strong>of</strong> RatIsl<strong>and</strong>, Easter Group, Abrolhos, Western Australia;28°42’S, 113°47.3’E, 201 m. Holotype WAM112/93. Dentalium kessneri Lamprell & Healy,1998. Known from the type locality only.606ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)kiaochowwanense, Dentalium (Episiphon) Tchang &Tsi, 1950: 6, 7, pl. 1, figs 1-6. <strong>Recent</strong>. KiaochowBay [Shantung, China], stn 358, 25 m. HolotypeASP (IOS?) 3542; paratypes ASP 3976 (3; stn 382,16 m), ASP 3293 (6; stn 336, 10 m), ASP 2227 (1;stn 248, 5 m), ASP 3261 (3; stn 331, 10 m), ASP4218 (stn 411, 18 m), ASP 2228 (9; stn 244, 5 m).Synonym: Episiphon kiaochowwanense var. rectum(Tchang & Tsi, 1950). Episiphon kiaochowwanense(Tchang & Tsi, 1950) (fide Steiner 1999: 153, 160,161). Indo-Pacific: China. 5-25 m.† kickxii, Dentalium Nyst, 1843: 454. Tertiary(Oligocene). Belgium. Based on Dentalium acuticostasensu Nyst 1835 non Deshayes, 1825. Antaliskickxii (Nyst, 1843) (fide Bonci et al. 2000: 207,pl. 1, fig. 2). Remarks: Janssen (1978a: 14, 15) originallyconsidered Dentalium geminatum to be a seniorsynonym <strong>of</strong> Dentalium kickxi Nyst, 1843; buthe considered Goldfuss’ name to be a “nomen oblitum”.Janssen (1978b: 138, 139) subsequentlyobtained the Goldfuss type material <strong>and</strong> concludedthat it was a valid <strong>species</strong> <strong>and</strong> a senior synonym <strong>of</strong>D. kickxii. Most recently, Janssen (1987: 174-176)determined that Antalis geminatum was found inthe Late Oligocene <strong>and</strong> A. kickxii was found in theMiddle Oligocene; hence the two <strong>species</strong> are notsynonyms. Dentalium kicksii sensu Tenison-Woods(1876) <strong>and</strong> sensu Etheridge (1878) are misidentifications<strong>of</strong> Dentalium mantelli Zittel, 1864 (fideTate 1887: 191; Pilsbry & Sharp 1898: 208).kikuchii, Siphonodentalium Kuroda & Habe, 1952:36. <strong>Recent</strong>. Toyama Bay, Honshu, Japan Sea.Types in NSMT. Compressidens kikuchii (Kuroda& Habe, 1952) (fide Habe 1964: 30). Indo-Pacific:Japan. 100-500 m. Remark: this <strong>species</strong> wasmisidentified as Dentalium (Compressidens) stearnsiPilsbry & Sharp, 1898 by Kuroda & Kikuchi(1933: 9).† kittli, Dentalium Cossmann, 1900a: 143. Triassic,St Cassian Formation. Tirol, Austria/Italy. Replacementname for Dentalium undulatum var. ellipticaKittl, 1894 non Dentalium ellipticum J. Sowerby,1814.† klipsteini, Dentalium Kittl, 1891: 172 [7], pl. 1,fig. 3. Triassic, St Cassian Formation. Tirol,Austria/Italy. Holotype NHM 1899/7/1.Prodentalium klipsteini (Kittl, 1891) (fide Stiller2001: 630). Remark: introduced for Dentalium“quadrangulatum Klipstein” (a manuscript name).knudseni, Wemersoniella Scarabino, 1995: 326, 327,figs 127, 129b, c. <strong>Recent</strong>. Kermadec Trench,Galathea stn 664, 36°34’S, 178°57’E, 4510-4570 m. Holotype ZMUC; paratypes ZMUC (2;1 each from stn 601 <strong>and</strong> stn 662), MNHN (1;stn 664), NMNZ M268545 (2; Tangaroastn P934). Wemersoniella knudseni Scarabino,1995. New Zeal<strong>and</strong>. 4105-4630 m.koeningianum, Dentalium Risso, 1826a: 125. Fossil.Near Nice, France. Nomen nudum.† komiensis, Laevidentalium “?” Kulikov, 1967: 116,pl. 1, fig. 1. Permian. Vem River, Soviet Union[Komi Republic, Russia]. Holotype GeologicalInstitute (St Petersbourg), 8758/3.komooksense, Dentalium Meek, 1876: 364, pl. 3, fig. 6.Cretaceous. Nanaimo, Vancouver Isl<strong>and</strong>, BritishColumbia, Canada. Holotype USNM Paleobiology4270. An objective junior synonym <strong>of</strong> Dentaliumnanaimoense Meek, 1858 (fide Pilsbry & Sharp1898: 225, “The description <strong>of</strong> both <strong>of</strong> these <strong>species</strong>is from the same specimen”).† korobkovi, Cadulus (Gadila) Merklin, 1961: 157,text-fig. 2. Middle Eocene. Northern Caucasus,Goryachiy Kluych, near Krasnodar, Russia.† korytnicense, Dentalium (Antalis) Baluk, 1972: 557-559, pl. 2, figs 4-6, text-figs 6, 7. Miocene, LowerTortonian (Badenian). Korytnica, 24 km SSW <strong>of</strong>Kielce, Holy Cross Mountains, Pol<strong>and</strong>. Holotype<strong>and</strong> paratypes, Baluk collection.† kubraense, Dentalium Palmer, 1974b: 124.Cretaceous, Barremian. River Kubre, near VolgaRiver, Russia. Replacement name for Dentaliumbarremicus Sasanova, 1958 non Dentalium barremicumCossmann, 1916.kurogenge, Pulsellum Habe & Kosuge, 1964: 9.<strong>Recent</strong>. Off Choshi, Chiba Pref., Honshu, Japan,about 350 m. Holotype <strong>and</strong> paratype NSMT.Pulsellum kurogenge Habe & Kosuge, 1964. Indo-Pacific: Japan. 200-500 m.labeyriei, Cadulus Scarabino, 1995: 353, figs 158,160j. <strong>Recent</strong>. Philippines, RV Jean Charcot,ESTASE 2 stn DW1, 14°05’N, 120°01’E, 2200 m.Holotype MNHN; paratypes MNHN (2 fromDW1; 7 from CP2), USNM 890872 (1; CP 2).Cadulus labeyriei Scarabino, 1995. Indo-Pacific:Philippines. 2050-2200 m.labiatum, Dentalium Turton, 1819: 38. <strong>Recent</strong>.Torbay [Engl<strong>and</strong>]. Syntypes USNM 174971 (6).Junior synonym <strong>of</strong> Antalis vulgaris (Da Costa,1778) (fide Pilsbry & Sharp 1897: 41).labiatum, Dentalium Brown, 1827: caption to pl. 1,fig. 4. <strong>Recent</strong>. Eastern Atlantic: Lough Strangford,Irel<strong>and</strong>. Junior homonym <strong>of</strong> Dentalium labiatumTurton, 1819. Remarks: it is not clear whetherBrown referred to Turton’s <strong>species</strong>. Further researchis required to support the suspected synonymy withAntalis vulgaris (Da Costa, 1778).lacteolum, Dentalium (Laevidentalium) Tate, 1899:264. Eocene. Muddy Creek <strong>and</strong> Gelibr<strong>and</strong> River,Victoria, Australia. Holotype <strong>and</strong> 6 paratypes SAMT 253. Junior synonym <strong>of</strong> Dentalium (Laevidentalium)acriculum (Tate, 1887) (fide Ludbrook1959: 144). Remark: this <strong>species</strong> was misidentifiedas Dentalium lacteum by Tate (1887: 193).lacteum, Dentalium Deshayes, 1825: 362 [42 inreprint], pl. 16, fig. 27. <strong>Recent</strong>. Indian Ocean,India. Lectotype MNHN (designated by Scarabino1995: 242); paralectotypes MNHN (1); MHNG1155/35 (“Seychelles”; don. M. Desh.). GraptacmeZOOSYSTEMA • 2004 • 26 (4)607


Steiner G. & Kabat A. R.lactea (Deshayes, 1825) (fide Scarabino 1995: 242).Indo-Pacific: Seychelles to New Caledonia <strong>and</strong>China. 32-42 m. Remark: the <strong>species</strong> from thePliocene <strong>of</strong> Italy O. G. Costa (1851: 37, pl. 3,fig. 7) misidentified as Dentalium lacteum was laternamed Dentalium costae Pilsbry & Sharp, 1898.† ladinum, Dentalium Dall, 1916: 522. Oligocene.USGS station 6774, at Rock Isl<strong>and</strong>, Suwanee River,near White Springs, Hamilton County, Florida,USA. Syntypes USNM Paleobiology 166746 (3).laeve, Dentalium Turton, 1819: 256. <strong>Recent</strong>. Torbay[Engl<strong>and</strong>]. Syntypes USNM 174969 (6). Juniorsynonym <strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fidePilsbry & Sharp 1897: 42).laeve, Dentalium Brazier, 1877. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.laeve, Dentalium (Antalis) “n. sp.?” Hilgard &Hopkins, 1878: 48, pl. 3, fig. 6. Quaternary. LakeBorgne, Louisiana, USA. Junior homonym <strong>of</strong>Dentalium laeve Turton, 1819; junior secondaryhomonym <strong>of</strong> Dentalium laeve (Schlotheim, 1820).Remark: Pilsbry & Sharp (1898: 207) stated thatthis <strong>species</strong> was described from “young shells <strong>and</strong>fragments, probably referable to D. filum Sowerby<strong>of</strong> the <strong>Recent</strong> <strong>and</strong> Pliocene faunas”.† laevigatum, Dentalium Eichwald, 1830: 199.Pliocene. Zukowce, Volhynia, Ukraine.laevigatum, Dentalium Rayneval, Hecke & Ponzi,1854: 19, 20. Pliocene. Monte Mario, Rome, Italy.Junior homonym <strong>of</strong> Dentalium laevigatum Eichwald,1830. Junior synonym <strong>of</strong> Cadulus (Dischides)politus (S. Wood, 1842) (fide Pilsbry & Sharp 1897:144). Remark: this name was inadvertently reusedfor a new <strong>species</strong> by Ponzi (1876: 947, 956, pl. 3,figs 1a, 1b), even though Ponzi was the junior(third) author <strong>of</strong> the 1854 publication.laevigatum, Dentalium Ponzi, 1876: 947, 956, pl. 3,figs 1a, 1b. Pliocene. Monte Vaticano, Rome, Italy.Junior homonym <strong>of</strong> Dentalium laevigatumEichwald, 1830. Tentative junior synonym <strong>of</strong>Antalis vulgaris vitrea (Gmelin, 1791) (fide Sacco1897: 101; Pavia 1991: 122, 123, pl. 3, fig. 8).Remarks: although Ponzi was the junior (third)author <strong>of</strong> Dentalium laevigatum Rayneval, Hecke &Ponzi, 1854, he apparently overlooked that descriptionwhen he reused this name for a distinct <strong>species</strong>from a nearby locality. Sacco (1897: 101) listed thisas a potential junior synonym <strong>of</strong> Dentalium (Antale)vitreum Gmelin, 1791 now considered a sub<strong>species</strong><strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fide Pavia1991). CLEMAM lists it in synonymy withDischides politus (S. Wood, 1842).† laevis, Dentalites Schlotheim, 1820: 93, 94. MiddleTriassic. Thangelstadt <strong>and</strong> Holzendorf (nearWeimar), Germany; near Copenhagen, Denmark.Entalis laevis (Schlotheim, 1820) (fide Diener 1926:230).laevis, Dentalium Hutton, 1873b: 2. Miocene, PareoraFormation. Kanieri; Chatham Isl<strong>and</strong>s; Motanau,New Zeal<strong>and</strong>. Lectotype NZGS TM4812 (fideKeyes 1972: 97). Junior homonym <strong>of</strong> Dentalites laevisSchlotheim 1820; replacement name: Dentaliumpareorensis Pilsbry & Sharp, 1898. Antalis pareorensis(Pilsbry & Sharp, 1898) (fide Maxwell 1988:76). Remark: Keyes did not indicate the source <strong>of</strong>this lectotype designation.† laguncula, Gadila Darragh, 1997: 94, 95, figs 8i, 8lp.Paleocene, Pebble Point Formation. LocalityPL3001, Otway Basin, Victoria, Australia. HolotypeNMV P301881; paratypes NMV P301882,P301883.† lamarcki, Dentalium Mayer, 1864: 357, 358, pl. 14,fig. 5. Miocene. Bordeaux <strong>and</strong> Dax, France. Juniorhomonym <strong>of</strong> Dentalium lamarckii Chenu, 1843.Replacement name: Dentalium (Entalis) exlamarckiSacco, 1897.lamarckii, Dentalium Chenu, 1843: pl. 6, fig. 15;1852: 5. <strong>Recent</strong>. “Les mers de la Chine” [ChinaSea?]. Possible syntype MHNG 1155/36 (labelled“Zanzibar”, fide Y. Finet in litt. 16.X.1996). Juniorsynonym <strong>of</strong> Antalis longitrorsum (Reeve, 1843)(fide Pilsbry & Sharp 1897: 111).† lamberti, Dentalium Cossmann, 1897: 268, pl. 2,fig. 32. Cretaceous, Turonian. Gizaucourt, Marne;Novy-Chevrières, Ardennes; <strong>and</strong> Berzieux, Marne,France.† lanceolata, Theca Morris, 1845: 289, 290, pl. 18,fig. 8. Permian. Illawara, New South Wales,Australia. Remark: Waterhouse (1980: 199) suggestedthat this <strong>species</strong> “is possibly similar” toPlagioglypta.† l<strong>and</strong>inense, Dentalium Vincent, 1876: 158, 159,pl. 9, figs 12a-12c. Lower Eocene. Gelinden <strong>and</strong>Maret, Belgium.† laneensis, Dentalium (“?” Fissidentalium) Hickman,1969: 74, pl. 9, figs 1-6. Oligocene, EugeneFormation. Near Eugene, Oregon, USA (numerouslocalities). Holotype UO 27332; paratypes UO27333-UO 27339.langfordi, Laevidentalium Habe, 1963: 268, pl. 38,fig. 22. <strong>Recent</strong>. Itoman, Okinawa Isl<strong>and</strong>, RyukyuIsl<strong>and</strong>s, Japan, 30-50 fms [55-91 m]. Holotype <strong>and</strong>paratype in NSMT. Fustiaria langfordi (Habe,1963) (fide Scarabino 1995: 280). Indo-Pacific:Indonesia to New Caledonia. 50-380 m.laqueatum, Dentalium Verrill, 1885a: 431, 432,pl. 44, fig. 18. <strong>Recent</strong>. Off Cape Hatteras, NorthCarolina, Albatross stn 2268, 35°10’40”N,75°06’10”W, 68 fms [124 m]. Holotype USNM44671. Dentalium laqueatum Verrill, 1885.Western Atlantic, Caribbean: from Cape Hatteras(USA) to Guyana. 18-900 m.lardum, Dentalium Barnard, 1963a: 445. <strong>Recent</strong>.South Africa, 33°44’S, 16°30’E, 1500 fms [2738 m].Syntypes SAFM A9768 (3; fide Giles & Gosliner1983: 40), BMNH 1964242. Compressidentaliumlardum (Barnard, 1963) (fide Scarabino 1995:264). Indian Ocean: South Africa. 2519-3212 m.608ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)largicrescens, Dentalium (Laevidentalium) Tate, 1899:264, pl. 8, figs 10-10a. Cheltenhamian (UpperMiocene [or Early Pliocene]). Beaumaris, Victoria,Australia, S<strong>and</strong>ringham S<strong>and</strong>s, Black RockMember. Lectotype SAM D 18934 (ex Tate Coll.T1611) (designated by Ludbrook 1959: 145); paralectotypesSAM D 18935 (5), SAM D 18936 (1;Pliocene, Muddy Creek). Laevidentalium largicrescens(Tate, 1899) (fide Lamprell & Healy 1998:115). Indo-Pacific: eastern Australia. 284-3058 m.laseroni, Dentalium (Laevidentalium) Colman, 1958:143, fig. 7. <strong>Recent</strong>. Broken Bay, New South Wales,Australia. Holotype AMS C62221; paratypes AMSC62220 (4). Junior synonym <strong>of</strong> Laevidentaliumlubricatum (Sowerby, 1860) (fide Lamprell &Healy 1998: 118).laterischismum, Fissidentalium Shikama & Habe,1963: 249, text-figs 1, 2. <strong>Recent</strong>. Off Monbetsu,Hokkaido, Ochotsk Sea. Holotype in GIY.Fissidentalium laterischismum Shikama & Habe,1963. Ochotsk Sea, Japan. 100-200 m.latesuclatum, see latesulcatum.† latesulcatum, Dentalium (Fissidentalium) Tate, 1899:262, pl. 8, fig. 9. “Miocene” (Pliocene). GrangeBurn, near Hamilton, Victoria, Australia. Holotype<strong>and</strong> 6 paratypes SAM T 1610A. Junior synonym:Dentalium (Paradentalium) howchini Cotton &Ludbrook, 1938 (fide Ludbrook 1959: 142).Dentalium (Dentalium) latesulcatum Tate, 1899 (fideLudbrook 1959: 142, pl. 1, fig. 1). Remarks: theheader to the <strong>species</strong> description is erroneouslyspelled as “latesuclatum” but the name is correctlyspelled as latesulcatum in the Index <strong>and</strong> plate caption.Dentalium (Fissidentalium) latesuclatum [Tate,1899: 262] is an incorrect subsequent spelling.† laticornuata, Compressidens Darragh, 1997: 94,fig. 8a-c, f. Paleocene, Pebble Point Formation.Locality PL3003, Otway Basin, Victoria, Australia.Holotype NMV P301880; paratypes NMVP301878, P303879, WAM 94.403a.† laticostata, Dentalium (Entalis) badensis var. Sacco,1897: 108, pl. 9, fig. 22 [as a nomen nudum inSacco 1896: 97]. Pliocene. Turin Hills, Italy. Juniorhomonym <strong>of</strong> Dentalium laticostatum Reuss, 1844.Type material not mentioned by Ferrero Mortara etal. (1984: 307).† laticostatum, Dentalium Reuss, 1844: 201.Cretaceous. Luschitz, Bohemia, Czech Republic.laubieri, Siphonodentalium Bouchet & Warén, 1979:219, 220, figs 8, 35, 36. <strong>Recent</strong>. Norwegian Sea,NORBI stn CP17, 73°30.7’N, 13°39.6’W,2502 m. Holotype <strong>and</strong> paratypes MNHN. Siphonodentaliumlaubieri Bouchet & Warén, 1979.Eastern Atlantic: Norway. 2212-2502 m.laugieri, Dentalium Jousseaume, 1894: 103. <strong>Recent</strong>.Aden, Suez. Syntypes (10) MNHN. Junior synonym<strong>of</strong> Dentalium reevei Fischer, 1871 (fideScarabino 1995: 204). Remark: Ludbrook (1954:98) argued against the synonymy.leai, Dentalium Meyer, 1885: 462; 1886: pl. 1, figs 2,2a. Eocene. Claiborne, Alabama, USA. LectotypeUSNM Paleobiology 638702 (designated byPalmer & Brann 1965: 366 as “holotype”). Juniorsynonym <strong>of</strong> Dentalium arciformis Conrad, 1846(fide Pilsbry & Sharp 1898: 199).lebruni, Dentalium Mabille & Rochebrune inRochebrune & Mabille, 1889: 99. <strong>Recent</strong>. SantaCruz [Argentinia]. Type material not located.Laevidentalium(?) lebruni (Mabille & Rochebrunein Rochebrune & Mabille, 1889) (fide Pilsbry &Sharp 1897: 102). Western Atlantic: Argentina;Eastern Pacific: Chile, Magellan Region. Remarks:due to its inadequate description <strong>and</strong> that the typematerial could not be found, the generic assignment<strong>of</strong> this <strong>species</strong> remains uncertain, as is the tentativesynonymy by Dell (1964: 124) with Dentaliummajorinum or Rhabdus perceptus.† lebuense, Dentalium Philippi, 1887a: 106, 107,pl. 12, fig. 18; 1887b: 101, pl. 12, fig. 18. Tertiary.Lebu <strong>and</strong> Llancahue, Chile.leloeuffi, Cadulus (Dischides) Nicklès, 1979: 71, 72,fig. 26. <strong>Recent</strong>. W <strong>of</strong> Gr<strong>and</strong> Bassam, Ivory Coast,ORSTOM 1966-1967, 19-20 m. Holotype <strong>and</strong>paratypes MNHN. Dischides leloeuffi (Nicklès,1979). Eastern Atlantic: Africa, Ivory Coast. 19-20 m.† lentum, Dentalium Yokoyama, 1923: 54, 55, pl. 6,figs 18-20. Tertiary. Takinai, near Tanabe Bay, Kii,Japan.† leonense, Dentalium (Dentalium “?”) disparile subsp.Mansfield, 1930: 139, pl. 21, fig. 8. Miocene.USGS station 3422, Jackson Bluff, OcklockneeRiver, Leon County, Florida, USA. HolotypeUSNM Paleobiology 370542.† leoniae, Dentalium Meunier, 1878: 122. Eocene.Jaignes, Seine-et-Marne, France. Dentalium leoninae[Simroth, 1894: 375] is an incorrect subsequentspelling.leoninae, see leoniae.leploskeles, see leptosceles.† leptoceras, Dentalium Ryckholt, 1862: vol. 3, pl. 35,fig. 20 (name <strong>and</strong> figure only).leptoconcha, Costentalina Chistikov, 1982b: 1319,1320, pl. 1, figs 15, 16, pl. 5, figs 1-9. <strong>Recent</strong>.Eastern Pacific, Akademik Kurtschatow stn 271 T,17°42’S, 78°59.2’W, 3080-2710 m. HolotypeIOM; paratypes (2 from Akademik Kurtschatow stn271; 1 from Dimitry Mendeleev stn 525-3,5°13.6’N, 107°31.7’W, 3830 m; 1 from DimitryMendeleev stn 522-2, 5°10.3’N, 121°29.5’W,4426 m). Costentalina leptoconcha Chistikov,1982. Eastern Pacific: Peru. 2710-4426 m.† leptodoma, Cadulus (Gadila) Pilsbry & Olsson,1941: 49, pl. 10, fig. 11. Pliocene, CanoaFormation. Punta Blanca, western Ecuador.Holotype ANSP 13720.leptosceles, Dentalium Watson, 1879: 513 [Watson1886: 7, pl. 1, fig. 6 as leptoskeles]. <strong>Recent</strong>. South <strong>of</strong>ZOOSYSTEMA • 2004 • 26 (4)609


Steiner G. & Kabat A. R.Australia, Challenger stn 160, 42°42’S, 134°10’E,2600 fms [4758 m]. Lectotype BMNH1887.2.9.21 (designated by Ludbrook 1954: 104);paralectotypes BMNH 1887.2.9.22 (1), BMNH1889.11.11.162 (1). Junior synonym: Dentaliumbanale Boissevain, 1906 (fide Scarabino 1995: 292).Laevidentalium leptosceles (Watson, 1879) (fidePilsbry & Sharp 1897: 110). Indo-Pacific: Africa toAustralia <strong>and</strong> New Caledonia. 918-5300 m.Remarks: Lamprell & Healy (1998: 112, 114, 115)treated Laevidentalium banale as a separate <strong>species</strong>.Dentalium leploskeles [Pilsbry & Sharp, 1897: 110]is an incorrect subsequent spelling.leptum, Dentalium Bush in Verrill, 1885b: 586, 587[also in Bush 1885: 470, 471, pl. 45, figs 18, 18a].<strong>Recent</strong>. Off Cape Hatteras, USBF, stn 2112, 14-15 fms [26-27 m]. Holotype USNM 41562. Juniorsynonym <strong>of</strong> Graptacme eborea (Conrad, 1846) (fideHenderson 1920: 66).† leroyi, Fustiaria (Fustiaria) Hodgkinson, 1974: 23,24, text-fig. 8l, m; pl. 3, fig. 12, pl. 8, figs 1,2. Middle Eocene, Claibornian Stage, CookMountain Formation, Wheelock Member. LittleBrazos Bluff, Brazos River Valley, Brazos County,Texas, USA. Holotype USNM Paleobiology180460; paratypes USNM Paleobiology 180478,PRI 29229, ANSP 31507 (all from Scott’s localityMS-2-N, Little Brazos River).lessoni, Dentalium Deshayes, 1825: 357 [37 inreprint], pl. 2, fig. 13. <strong>Recent</strong>. New Guinea. LectotypeMNHN (designated by Scarabino 1995: 215);paralectotypes MNHN (87). Dentalium lessoniDeshayes, 1825. Indo-Pacific: New Guinea. 904-1141 m. Remark: it remains to be determined howthis deep-water <strong>species</strong> could have reached Deshayesbefore 1825.letsonae, Dentalium (Dentalium) Sharp & Pilsbry inPilsbry & Sharp, 1897: 4, 5, pl. 1, fig. 13, pl. 5,figs 66-68. <strong>Recent</strong>. Isl<strong>and</strong> <strong>of</strong> Bohol, Philippines.Holotype ANSP 35446. Dentalium letsonae Sharp& Pilsbry in Pilsbry & Sharp, 1897. Known fromthe type locality only.leucoryx, Dentalium Boissevain, 1906: 20, 21, pl. 6,fig. 23, text-fig. 15. <strong>Recent</strong>. Indonesia, Sibogastn 285, 08°39.1’S, 127°04.4’E, 34 m. LectotypeZMA 3.06.016 (designated by Scarabino 1995:206); paralectotypes ZMA 3.06.017 (2). Juniorsynonym: Dentalium sinuosum Boissevain 1906(fide Scarabino 1995: 206, by First Reviser action).Dentalium leucoryx Boissevain, 1906. Indo-Pacific:Indonesia to Chinese Sea. 34-157 m.† leve, Dentalium Stephenson, 1923: 351, 352, pl. 88,figs 1-3. Upper Cretaceous. USGS Locality 6815,0.5 miles W <strong>of</strong> Union Springs, Alabama, USA.Holotype USNM Paleobiology 31837; paratypesUSNM Paleobiology 31838 <strong>and</strong> 31993 (USGSLocality 5348, Snow Hill, Greene County, NorthCarolina). Junior synonym: Dentalium demersumPilsbry, 1927 (fide Stephenson 1941: 253, 254)(replacement name for Dentalium inornatum Wade,1926).levii, Fissidentalium Scarabino, 1995: 252, 253,figs 60, 70c. <strong>Recent</strong>. New Caledonia, RV Vauban,MUSORSTOM 4, stn DW160, 18°42’S,163°13’E, 668 m. Holotype MNHN; paratypesMNHN (2 from DW160; 1 from DW 341; 1 fromDW358; 1 from DW33; 1 from DW21; 1 fromDW55; 1 from Vauban 1976); AMS C201724(1; DW39); NMNZ M268957 (1; DW396).Fissidentalium levii Scarabino, 1995. Indo-Pacific:New Caledonia. 200-1400 m.† liassicum, Dentalium Moore, 1866: 86, pl. 5, fig. 24.Jurassic, Middle Liassic. Camerton, Engl<strong>and</strong>,United Kingdom. Holotype Bath Museum(Engl<strong>and</strong>) (fide Richardson 1906: 581, 582, pl. 45,fig. 6). Prodentalium “?” liassicum (Moore, 1866)(fide Engeser & Riedel 1992: 38).† ligeriense, Dentalium Peyrot, 1938: 8, 9, pl. 1,figs 46, 47. Miocene, Helvetian. Mirebeau(Moulin-Pochard), Ligerian Basin, Touraine,France. Type material stated to be in author’s collection.† lighti, Dentalium (Laevidentalium “?”) Vokes, 1939:105, 106, pl. 16, fig. 26. Eocene, Arroyo HondoFormation. UCMP locality 1817, N <strong>of</strong> Coalinga,Coalinga Quadrangle, California, USA. HolotypeUCMP 15752.ligusticus, Siphonodentalium (Loxoporus) Razzore,1896: 256, pl. 3, figs 3a, 3b. Pliocene. Liguria,Italy. Junior synonym <strong>of</strong> Gadila ventricosa (Bronn,1828) (fide Pilsbry & Sharp 1898: 240).lima, Fissidentalium Kuroda & Habe in Habe, 1963:260, pl. 37, fig. 15. As a nomen nudum in Habe(1953: 293). <strong>Recent</strong>. Okinoshima, KochiPrefecture, Shikoku, Japan, 40 m. Types in NSMT.Junior synonym <strong>of</strong> Fissidentalium pr<strong>of</strong>undorum(E.A. Smith, 1894) (fide Habe 1964: 13).† limatuloides, Dentalium Kuhn, 1935: 126, 127,pl. 8, fig. 6a-c. Jurassic, Dogger. Forth, Bavaria,Germany. Baltodentalium limatuloides (Kuhn,1935) (fide Engeser & Riedel 1992: 50).† limatulum, Dentalium Tate, 1870a: 402, pl. 26,fig. 1. Jurassic, Lower Liassic. Gloucestershire,Engl<strong>and</strong>, United Kingdom. Holotype GeologicalSociety <strong>of</strong> London 4441 (fide Richardson 1906:582). Baltodentalium limatulum (Tate, 1870) (fideEngeser & Riedel 1992: 38, 50).† limatum, Dentalium (Laevidentalium) Stanton,1901: 28, 29, pl. 6, fig. 9. Cretaceous. 10 miles E <strong>of</strong>Lago Pueyrrydón, Santa Cruz, Patagonia,Argentina.lineare, Dentalium Renier, 1804: xii. Nomen nudum.Remark: Renier’s publication was placed on theOfficial Index <strong>of</strong> Rejected <strong>and</strong> Invalid Works inZoological Nomenclature (ICZN 1954, Opinion316), so this name is not available.† lineatum, Dentalium Gueranger, 1853: 33.Cretaceous, Greens<strong>and</strong> stage. Le Mans, France.610ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)lineatum, Dentalium Moore, 1870: 256. Cretaceous.Wollumbilla, Queensl<strong>and</strong>, Australia. Junior homonym<strong>of</strong> Dentalium lineatum Gueranger, 1853.Replacement <strong>names</strong>: Dentalium wollumbillaensisEtheridge in Jack & Etheridge, 1892, Dentaliummoorei Pilsbry & Sharp, 1898.lineolatum, Dentalium Cooke, 1885: 274. <strong>Recent</strong>. Gulf<strong>of</strong> Suez. Syntypes UMZC I.100,750 (7). Junior synonym<strong>of</strong> Dentalium reevei Fischer, 1871 (fideScarabino 1995: 204). Remark: Boissevain (1904:25) substituted Cook’s name with Dentalium mac<strong>and</strong>rewi,which became a junior synonym <strong>of</strong> Dentaliumclavus Cooke, 1885 by a lectotype designation byScarabino (1995: 204); see Singer (2003) for details.linnaeanum, Dentalium Locard, 1887: 145. Juniorsynonym <strong>of</strong> Antalis dentalis (Linnaeus, 1758).Remark: this name was proposed in a footnote as asuggestion to avoid allegedly tautonymous nomenclaturebut is an invalid name (ICZN 1999: articles18 <strong>and</strong> 23.3.7).† linnei, Dentalium Foresti, 1895: 246. Pliocene.Ponticello in val di Saveno, near Bologna, Italy.liodon, Dentalium (Laevidentalium) Pilsbry & Sharp,1897: 107, 108, pl. 21, figs 37-39. <strong>Recent</strong>.St Martin, West Indies. Holotype ANSP 35554.Fustiaria liodon (Pilsbry & Sharp, 1897) (fideScarabino 1975: 185). Western Atlantic,Caribbean: Florida to Brazil. 8-366 m.liratum, Dentalium Carpenter, 1857: 188. <strong>Recent</strong>.Mazatlan [Mexico]. Lectotype BMNH 1857.6.4.879/1 (designated by Keen 1968: 402, text-fig. 23);paralectotypes BMNH 1857.6.4.879/2-3 (2).Junior synonym <strong>of</strong> Graptacme semistriata (Turton,1819) (fide Pilsbry & Sharp 1897: 91).lirulatum, Dentalium Mörch, 1861: 177. <strong>Recent</strong>. Gulf<strong>of</strong> Nicoya, west coast <strong>of</strong> Costa Rica. HolotypeZMUC (fide Keen 1966: 18, fig. 20). Junior synonym<strong>of</strong> Graptacme semistriata (Turton, 1819)(fide Pilsbry & Sharp 1897: 91).lobatum, Dentalium Sowerby, 1860: 100, pl. 3,fig. 44. <strong>Recent</strong>. Type locality not designated.Syntypes BMNH 1951.2.14.4-5 (labelled“Spitzbergen”). Junior synonym <strong>of</strong> Dentalium vitreumM. Sars, 1851, which is a junior homonym <strong>of</strong>Dentalium vitreum Gmelin, 1791; validated as substitutename by Pilsbry & Sharp (1897: 136, 137).Junior synonym: S. exvitreum Sacco, 1897 (fidePilsbry & Sharp 1898: 253). Siphonodentaliumlobatum (Sowerby, 1860) (fide M. Sars 1859: 52;fide Pilsbry & Sharp 1897: 136). Eastern Atlantic:British Isl<strong>and</strong>s to Spitzbergen. 38(?)-3100 m.† lobion, Cadulus (Polyschides) Gardner, 1947: 628,pl. 57, fig. 21. Oligocene. Alum Bluff Group,Chipola Formation. USGS locality 2213, 1 milebelow Bailey’s Ferry, Chipola River, CalhounCounty, Florida, USA. Holotype USNMPaleobiology 498411.lochi, Dentalium (Dentalium) Lamprell & Healy,1998: 37, figs 29B, 30B, 32. <strong>Recent</strong>. Arafura Sea,Northern Territory, Australia, 9°17.5’S, 132°20’E,168 m. Holotype AMS C174628; paratypes AMSC84257 (270), WAM 153/93 (2; 18°25’S,118°22’E, 210 km N <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 201 m). Dentalium lochi Lamprell &Healy, 1998. Indo-Pacific: northern <strong>and</strong> westernAustralia. 124-201 m.l<strong>of</strong>olense, see l<strong>of</strong>otense.l<strong>of</strong>otense, Siphonodentalium M. Sars, 1865a: 297-300,pl. 6, figs 29-33. <strong>Recent</strong>. L<strong>of</strong>oten Isl<strong>and</strong>s, Nidaros[Trondheimfjord], Flekkefjord [Norway], 30-120 fms [55-219 m]. Syntypes SMNH 4624(Norway, L<strong>of</strong>oten, 100 fms [183 m]). Pulselluml<strong>of</strong>otense (M. Sars, 1865) (fide Fischer 1885: 894).Eastern Atlantic: Norway to Spain, Mediterranean.26-3500 m. Pulsellum l<strong>of</strong>olense [Cossmann 1888:11], Pulsellum lophotense [Coppi 1881: 86; Sacco1896: 98; 1897: 114] <strong>and</strong> Pulsellum lophotensis areincorrect subsequent spellings.† lombardicum, Dentalium Kittl, 1899: 4, pl. 1, fig. 1.Triassic? Esino <strong>and</strong> Canova, on Monte Croce, Italy.Holotype NHM 1889/D/2608. Remarks: as “longobardicum”[sic] (Kittl, 1899: 191); as “esinense” [inerror] (Kittl, 1899: 4).† longifissum, Dentalium Harmer, 1923: 823, pl. 63,figs 40, 41. Pliocene, Coralline Crag. Sutton,Gedgrave, Engl<strong>and</strong>.longilobatus, Cadulus (Polyschides) Boissevain, 1906:68, 69, pl. 6, figs 55, 56, text-fig. 34. <strong>Recent</strong>.Indonesia, Siboga stn 133, <strong>of</strong>f Lirung, SalibabuIsl<strong>and</strong>, up to 36 m. Syntypes ZMA 3.06.094-095(3). Siphonodentalium longilobatum (Boissevain,1906) (fide Scarabino 1995: 333). Known from thetype locality only.longirostrum, see longitrorsum.† longistriatum, Fissidentalium “?” Yoo, 1988: 248,figs 141-143. Early Carboniferous, Late Tournaisian,Dangarfield Formation. Locality A-3,2.6 km SW <strong>of</strong> Gundy, Upper Hunter, New SouthWales, Australia. Holotype AMS F.61992; paratypeAMS F.61993 (1).longitrorsum, Dentalium Reeve, 1843: pl. 130, fig. 6[1843: 197]. <strong>Recent</strong>. Type locality not specified[Zanzibar, Africa, in 1843]. Holotype BMNH1952.3.24.1. Junior synonym: Dentalium lamarckiiChenu, 1843 (fide Pilsbry & Sharp 1897: 111).Antalis longitrorsa (Reeve, 1843) (fide Scarabino1995: 227). Indo-Pacific: Red Sea <strong>and</strong> easternAfrica to China <strong>and</strong> Australia. 46-155 m. Dentaliumlongirostrum [Paetel 1873: 78] is an incorrect subsequentspelling (fide Pilsbry & Sharp 1897: 111).longobardicum, see lombardicum.longum, Dentalium (Episiphon) Sharp & Pilsbry inPilsbry & Sharp, 1897: 120, pl. 18, figs 1-3.<strong>Recent</strong>. Habitat unknown. Holotype ANSP 71080.Episiphon longum (Sharp & Pilsbry in Pilsbry &Sharp, 1897). Remark: possible junior synonym <strong>of</strong>Dentalium attenuatum Sowerby, 1860 non Say,1824 (fide Pilsbry & Sharp 1897: 120).ZOOSYSTEMA • 2004 • 26 (4)611


Steiner G. & Kabat A. R.lophotense, see l<strong>of</strong>otense.lophotensis, see l<strong>of</strong>otense.lubricatum, Dentalium Sowerby, 1860: 97, pl. 225,fig. 56. <strong>Recent</strong>. Off Port Jackson Heads, NewSouth Wales [Australia], 45 fms [82 m]. LectotypeBMNH 1957.10.10.2 (designated by Colman1958: 143); paralectotypes BMNH 1957.10.10.3-4(2). Junior synonym: Dentalium laseroni Colman,1958 (fide Lamprell & Healy 1998: 118; whoinvalidly selected a specimen from the lot BMNH1957.10.10.2-4 as the lectotype <strong>of</strong> lubricatum).Laevidentalium lubricatum (Sowerby, 1860) (fidePilsbry & Sharp 1897: 110). Indo-Pacific: Australia.82-1125 m.luchuanum, Dentalium Dall, 1926: 63. <strong>Recent</strong>. Nago,Okinawa, Japan, 15 fms [27 m]. Holotype USNM333692. Junior synonym <strong>of</strong> Dentalium aciculumGould, 1859 (fide Habe 1964: 24).† lucidum, Dentalium Deshayes, 1861: 214, pl. 1,figs 18-20. Eocene. Paris Basin (Cuise-la-Motte,Aisy, Mercin, Laversine, Coeuvres, Cuisy-en-Almont, Laon, Héronval, Noailles <strong>and</strong> Abbecourt),France. Fustiaria lucida (Deshayes, 1861) (fideNewton & Harris 1894: 65). Remarks: a valid<strong>species</strong> <strong>of</strong> Pseudantalis (fide Le Renard 1995: 175;Le Renard & Pacaud 1995: 85). This <strong>species</strong> wasmisidentified as Dentalium incertum Deshayes,1825 by d’Orbigny (1850).ludbrookae, Cadulus (Gadila) Cotton & Godfrey,1940: 340, 341, fig. 363. <strong>Recent</strong>. Northwest <strong>of</strong> CapeBorda, Kangaroo Isl<strong>and</strong>, South Australia, 62 fms[113 m]. Holotype SAM D 13760. Gadila ludbrookae(Cotton & Godfrey, 1940) (fide Lamprell &Healy 1998: 154). South Australia. 80-155 m.ludbrooki, Dentalium (Gadilina) Caprotti, 1962. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.ludbrooki, Striocadulus Scarabino, 1995: 336, 337,figs 138, 139c. <strong>Recent</strong>. Aden, John Murray Exp.stn 176, 12°05’N, 50°38’E, 655-732 m. HolotypeBMNH 1952.3.25.342; paratypes BMNH1952.3.25.343, BMNH 1952.3.25.138, 139 (2; stn188, 13°46’N, 47°50’E, 528 m). Striocadulus ludbrookaeScarabino, 1995. Indian Ocean, Red Sea.528-732 m. Remarks: this <strong>species</strong> was misidentifiedas Cadulus (Polyschides) hexaschistus Boissevain,1906 by Ludbrook (1954: 118). As Nelly HooperLudbrook was a woman, the gender <strong>of</strong> the name ishere emended to its female form.lunulus, Cadulus Dall, 1881: 35. <strong>Recent</strong>. Off MorroLight, Havana, Cuba, Blake stn 2, 23°14’N,82°25’W, 805 fms [1469 m]. Holotype MCZ7747. Cadulus lunulus Dall, 1881. WesternAtlantic, Caribbean. 128-1469 m.lupinum, Dentalium Röding, 1798: 69. [Citing Martini,1769: pl. 1, fig. 5; Gmelin, 1791: Dentalium striatulum].Junior objective synonym <strong>of</strong> Dentalium striatulumGmelin, 1791, which is a junior subjectivesynonym (partim) <strong>of</strong> Dentalium aprinum Linnaeus,1758 (fide Pilsbry & Sharp 1897: 3).† lusanensis, Dentalium Makarenko, 1970: 112, text.fig. 12, pl. 22, figs 11-13. Paleocene. Luzanovka,northern Ukraine. Holotype Akademiia NaukaUkraine (Kiev), No. 18/27.mac<strong>and</strong>rewi, Dentalium Boissevain, 1906: 25-27, pl. 5,figs 25-34 (ex Cooke, 1885: 274). <strong>Recent</strong>. Gulf <strong>of</strong>Suez, Red Sea. Lectotype (designated by Scarabino1995: 204) not in ZMA (fide R. Moolenbeek &A. N. Van der Bijl in litt. 23.VI.2000) or in UMZC.Substitute name for Dentalium aratorum, Dentaliumclavus, <strong>and</strong> Dentalium lineolatum, all Cooke, 1885(fide Boissevain 1906: 25); junior synonym <strong>of</strong>Dentalium clavus Cooke, 1885 (fide Singer 2003:11, 14).† macilentum, Dentalium Pilsbry, 1911: 166, 167,figs 1-23. Miocene. Bowden, Jamaica. SyntypeANSP 654. Remark: as a nomen nudum in Pilsbry& Sharp (1898: 254).macleani, Cadulus (Platyschides) Emerson, 1978: 122,figs 1-3. <strong>Recent</strong>. Muertos Bay, Baja California,Mexico, LACM stn 66-22, 24°55’N, 109°46’W,18-55 m. Holotype LACM 1886; paratypes LACM1887 (2); AMNH 213440 (1). Cadulus macleaniEmerson, 1978. Eastern Pacific: Mexico. 18-55 m.Remark: the subgenus Platyschides was put in synonymywith Gadila by Scarabino (1995: 196, 355).However, the maximum diameter <strong>of</strong> this <strong>species</strong> islocated at the middle <strong>of</strong> the shell, which is typicalfor Cadulus.† maculatellata, Dentalium (Antale “?”) dentale var.Sacco, 1897: 105, pl. 8, figs 79, 80 [as a nomennudum in Sacco 1896: 97]. Pliocene. Bussana, Italy.Syntypes MIGT BS.106.02.070, BS.106.02.071(fide Ferrero Mortara et al. 1984: 306).magellanicum, Dentalium (Dentalium) majorinumPilsbry & Sharp, 1897: 28, pl. 12, figs 95-97.<strong>Recent</strong>. USFC stn 2777 <strong>and</strong> 2780, Strait <strong>of</strong> Magellan,77.5-369 fms [141-673 m]. Syntypes USNM87651, USNM 601272. Synonym <strong>of</strong> Dentaliummajorinum Mabille & Rochebrune in Rochebrune& Mabille, 1889.magnificum, Dentalium E. A. Smith, 1896: 371, 372.<strong>Recent</strong>. Tricomalee, Sri Lanka, 8°40’N, 81°27’E,637-800 fms [1163-1460 m]. Syntypes ZSI M299/1; BMNH 1895.12.13.1 (1), NMW (1).Possible junior synonym: Dentalium noetlingiMartin, 1915 (see Noetling 1901: 250; Martin1915: 180). Fissidentalium magnificum (E. A.Smith, 1896) (fide Pilsbry & Sharp 1897: 78).Indo-Pacific: Madagascar to Philippines <strong>and</strong> NewCaledonia. 520-2600 m.“magnistriatum”, Dentalium “Desh.” – Anton 1838:25. Apparently an incorrect subsequent spelling forDentalium multistriatum Deshayes, 1825 (fidePilsbry & Sharp 1898: 208).† magnocostata, Dentalium sexangulum var. Sacco,1897: 94, pl. 7, fig. 62 [as a nomen nudum in Sacco1896: 97]. Pliocene. Astigiana, Italy. Type materialnot listed by Ferrero Mortara, et al. (1984: 301).612ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)† magnum, Dentalium Briart & Cornet, 1889: 81, 82,pl. 24, fig. 6. Upper Eocene. Mons, Belgium.† magnumcostatum, Dentalium Brunet, 1995: 50, 52,pl. 5, fig. 5. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-078.magnus, Cadulus (Polyschides) Boissevain, 1906: 68,pl. 6, fig. 54, text-fig. 33. <strong>Recent</strong>. Celebes Sea,Siboga stn 88, 00°34.6’S, 119°08.5’E, 1301 m.Lectotype ZMA 3.06.092 (designated by Scarabino1995: 330); paralectotypes ZMA 3.06.091 (1),ZMA 3.06.093 (1; Flores Sea, Siboga stn 45,07°24’S, 118°15.2’E, 794 m), ZMA 3.06.107(1; Timor Sea, Siboga stn 300, 10°48.6’S,123°23.1’E, 918 m). Siphonodentalium magnum(Boissevain, 1906) (fide Scarabino 1995: 330).Indo-Pacific: Indonesia to Japan <strong>and</strong> NewCaledonia. 300-2350 m. Remarks: Siphonodentaliumsumma (Okutani, 1964) is a possible juniorsynonym <strong>of</strong> this <strong>species</strong> (Habe 1964: 51, 52).Habe (1964: 51) referred to Siboga stn 45 (FloresSea, 7°24’S, 118°15.2’E, 794 m) as the type locality,but did not designate a lectotype pursuant toICZN (1999) article 74a.† magnus, Gadus Mayer-Eymar, 1887: 53, pl. 4,fig. 8. Eocene, Bartonian, Auversian. Schimberg,near Thun, Switzerl<strong>and</strong>.majestica, Entalina Kira, 1959: 105, pl. 40, fig. 3.<strong>Recent</strong>. Tosa Bay, Japan, 30-50 fms [55-91 m].Type material not located. Nomen nudum in Habe(1953: 299) <strong>and</strong> in Kira (1954: 80) (fide Bieler &Petit 1990: 141). Junior synonym <strong>of</strong> Entalina mirifica(E. A. Smith, 1895) (fide Scarabino 1995: 300).major, Cadulus (Gadila) panamensis var. Pilsbry &Sharp, 1898: 192, pl. 36, figs 28-30. <strong>Recent</strong>. LowerCalifornia, Albatross stn 2830, 66 fms [120 m].Lectotype USNM 96570 (designated by Emerson1971: 79); paralectotype USNM 602249. Synonym<strong>of</strong> Gadila perpusilla (Sowerby in Broderip &Sowerby, 1832) (fide Emerson 1971: 78).major, Dentalium ergasticum var. Locard, 1896: 170,pl. 6, fig. 1. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Type material not located.Synonym <strong>of</strong> Dentalium ergasticum, a junior synonym<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877).† major, Dentalium Gardner, 1877: 556, pl. 16, fig. 2.Cretaceous. Grey Chalk, Folkestone, Dover,Engl<strong>and</strong>. Dentalium majus [Gardner 1878] is anincorrect subsequent spelling.major, Dentalium striolatum var. Locard, 1898: 120.<strong>Recent</strong>. “Talisman, between 2400 <strong>and</strong> 2638 m”.Type material not located. Junior synonym <strong>of</strong>Antalis entalis (Linnaeus, 1758) (fide Pilsbry &Sharp 1897: 43).major, Pseudantalis rubescens var. Monterosato, 1884:33. Nomen nudum without type locality or description.major, Siphodentalium [sic] l<strong>of</strong>otense var. Monterosato,1875: 20. <strong>Recent</strong>. Palermo, 210 m. Nomen nudum.majorinum, Dentalium Mabille & Rochebrune inRochebrune & Mabille, 1889: 100, pl. 4, fig. 10.<strong>Recent</strong>. Baie Orange [Chile]. Syntypes MNHN (2).Dentalium majorinum Mabille & Rochebrune inRochebrune & Mabille, 1889. Antarctica: Ross Sea;Western Atlantic: Magellanic Region. 133-1097 m.majus, see major Gardner, 1877.majus, Dentalium J. de C. Sowerby, 1846: 263, pl. 2,fig. 3. Tertiary. Hufo Isl<strong>and</strong>, Chile. Dentalium sulcosumJ. de C. Sowerby, 1846 (fide Ortmann 1902:157, by First Reviser action, according to ICZN1999: article 24).makiyamai, Dentalium (Episiphon) (subrectum var.?)Kuroda & Kikuchi, 1933: 11, 12, pl. 1, fig. 8.<strong>Recent</strong>. Toyama Bay, Japan, 60-200 m. Types inFGK. Replacement name for Dentalium (E.) filumMakiyama, 1929 non Sowerby, 1860, which is aninappropriate use <strong>of</strong> Episiphon virgula (Hedley,1903) (fide Scarabino 1995: 287).malayanum, Dentalium Boissevain, 1906: 32, 33,text-fig. 19, pl. 5, figs 4, 5. <strong>Recent</strong>. Timor Sea,Siboga stn 300, 10°48.6’S, 123°23.1’E, 918 m.Lectotype ZMA 3.06.033 (designated by Scarabino1995: 251); paralectotypes ZMA 3.06.030(2; anchorage <strong>of</strong>f Lirung, Salibabu Isl<strong>and</strong> up to 36m, Siboga stn 133), ZMA 3.06.031 (1; north <strong>of</strong>K<strong>of</strong>ian Isl<strong>and</strong>, Siboga stn 159, 0°59.1’S,129°48.8’E, 411 m), ZMA 3.06.032 (1). Fissidentaliummalayanum (Boissevain, 1906) (fide Habe& Kosuge 1964: 3). Indo-Pacific: Madagascar toIndonesia <strong>and</strong> New Caledonia. 36-2760 m.maltzani, Dentalium Clessin, 1896: 41, 42, pl. 11,fig. 5 [Dunker in Paetel 1888: 593, as nomennudum]. <strong>Recent</strong>. Gorée, Senegal, West Africa.Types in SMF. Junior synonym <strong>of</strong> Fustiariarubescens (Deshayes, 1825) (fide Pilsbry & Sharp1897: 107). Dentalium malzani [Pilsbry & Sharp,1897: 107] is an incorrect subsequent spelling.malzani, see maltzani.† mancorens, Dentalium Olsson, 1930: 71, 72, pl. 12,figs 12, 13. Eocene, Talara Formation. QuebradaBarranca, near Catalinas, Peru. Holotype PRI24296; paratype PRI 24925 (fide Brann & Kent1960: 318-319).mannarense, Dentalium Winckworth, 1927: 167, 168,text-fig. <strong>and</strong> pl. 14, figs 1, 2. <strong>Recent</strong>. MannarIsl<strong>and</strong>, Sri Lanka, 3 fms [5 m]. Syntypes BMNH1952.3.21.6-12, NMW. Dentalium mannarenseWinckworth, 1927. Indian Ocean: Sri Lanka,India. 5-6 m.† mantelli, Dentalium Zittel, 1864: 45, pl. 13, fig. 7.Eocene. The Cliffs, near Nelson, New Zeal<strong>and</strong>.Dentalium mantelli Zittel, 1864 (fide Beu &Maxwell 1990: 424). Remarks: Maxwell (1992:186) noted that this <strong>species</strong> was described from twolocalities, <strong>and</strong> selected the specimen figured byMantell, from “The Cliffs, Nelson” as the lectotype(the other syntype was from Awatere Valley,Onekakara, New Zeal<strong>and</strong>, <strong>and</strong> is a paralectotypeZOOSYSTEMA • 2004 • 26 (4)613


Steiner G. & Kabat A. R.<strong>of</strong> Dentalium centenniale Marwick, 1942). Flügel(1959: 839) noted that the type material <strong>of</strong> this<strong>species</strong> could not be found.† maoria, Fustiaria Maxwell, 1992: 186, pl. 30, figs k-m.Eocene. NZGS locality GS9508, Waihao River,near McCulloch’s Bridge, South Canterbury, NewZeal<strong>and</strong>. Holotype NZGS TM7228, paratypesNZGS (2).marchadi, Cadulus (Gadila) Nicklès, 1979: 64, fig. 22.<strong>Recent</strong>. Senegal. Holotype MNHN. Gadila marchadi(Nicklès, 1979). Eastern Atlantic: WestAfrica. No bathymetric information available.marchadi, Dentalium (Graptacme) Nicklès, 1979: 55,text-fig. 10, pl. 1, fig. II. <strong>Recent</strong>. Dakar, Senegal.Holotype MNHN. Graptacme marchadi (Nicklès,1979). Eastern Atlantic: West Africa.† margarita, Cadulus (Polyschides) Palmer, 1947: 217,218, pl. 26, figs 8-11. Eocene, Jackson Formation.Moodys Branch marl, Mississippi, USA. HolotypePRI 4483; paratype PRI 4484.marshae, Laevidentalium Lamprell & Healy, 1998:122, 123, figs 121F, 125F, 128. <strong>Recent</strong>. 78 milesNNE <strong>of</strong> Port Hedl<strong>and</strong>, Western Australia,19°04.1’-04.7’S, 118°47.8’-47.2’E, 82 m. HolotypeAMS C174622; paratypes AMS C174546 (3),AMS C174542 (4; N <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 19°03.4-6’S, 119°03.3-6’E, 81 m), AMSC308353 (1; 18°03’S, 122°17’E), AMS C173298(1; 94 miles NNE <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 18°48’S, 119°00.8’E, 92-94 m). Laevidentaliummarshae Lamprell & Healy, 1998. Indo-Pacific: western Australia. 81-94 m.martensi, Dentalium Boissevain, 1906 [December]: 34,text-fig. 20, pl. 4, fig. 19, pl. 5, figs 1-3. <strong>Recent</strong>.Macassar Strait, Siboga stn 88, 00°34.6’N,119°08.5’E, 1301 m. Lectotype ZMA 3.06.045(designated by Ludbrook 1954: 101; <strong>and</strong> independentlyby Scarabino 1995: 261); paralectotypes ZMA3.06.046 (4 + fragments), ZMA 3.06.047(1; stn 223, 5°44.7’S, 126°27.3’E, 4391 m). Juniorsynonym <strong>of</strong> Compressidentalium subcurvatum(E. A. Smith, 1906) [October] (fide Scarabino1995: 261).† marthae, Dentalium Cossmann & Pissarro, 1902:179, pl. 31, fig. 45. Eocene. Hauteville, Cotentin,France.martini, Cadulus Finlay, 1927: 521. Tertiary.Ngembak, Java, Indonesia. Replacement name forDentalium compressum Martin, 1885 non d’Orbigny,1850 (Polychaeta) nec Watson, 1879 necMeyer, 1883 nec Sowerby, 1889. Remarks: Palmer(1974b: 124) claimed that Finlay’s <strong>species</strong> was actuallyreferable to Dentalium, so it was a junior secondaryhomonym <strong>of</strong> Dentalium martini Whitfield,1882; replacement name: Dentalium javensePalmer, 1974.martini, Cadulus Scarabino, 1995: 349, 350, figs 154,160f. <strong>Recent</strong>. New Caledonia, Bassin des Loyauté,RV Coriolis BIOGEOCAL, stn CP232, 21°34’S,166°27’E, 760-790 m. Holotype MNHN <strong>and</strong>paratypes (3 from CP232; 1 from CP75; 2 fromDW106). Junior homonym <strong>of</strong> Cadulus martiniFinlay, 1927. Replacement name Cadulus scarabinoin. nom.martini, Dentalium Whitfield, 1882. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.martini, Dentalium Boussac, 1911b: 256, pl. 16,figs 4, 5, 49-51, 53, 54. Oligocene, Priabonian.Faudon, Allons <strong>and</strong> Martinets, France. Juniorhomonym <strong>of</strong> Dentalium martini Whitfield, 1882.Replacement name: Dentalium boussaci Emerson,1954. Remark: this <strong>species</strong> was misidentified asDentalium striatum J. Sowerby, 1814 by Renevier(1890).martini, Dentalium (Fustiaria) Dollfus, 1915: 993,994. Oligocene, Priabonian Formation. Locality813, Saädang, Celebes, Indonesia. Junior homonym<strong>of</strong> Dentalium martini Whitfield, 1882.Replacement name: Dentalium insuranum Emerson,1954.martyi, Laevidentalium Lamprell & Healy, 1998: 123-125, figs 130A, 131C, D, 132, 134E, F. <strong>Recent</strong>.Gulf <strong>of</strong> Carpentaria, Queensl<strong>and</strong>, Australia,10°58.5’S, 140°21’E, 54 m. Holotype QMMO39004; paratypes QM MO39006 (1; Gulf <strong>of</strong>Carpentaria, Queensl<strong>and</strong>, 13°1.5’S, 136°43’E,22 m), QM MO39009 (1; Gulf <strong>of</strong> Carpentaria,Queensl<strong>and</strong>, 11°4.1’S, 142°52.4E, 21 m), AMSC171342 (6; 98 km W <strong>of</strong> Cape Wessel, ArafuraSea, Northern Territory, 10°48.4’S, 135°52.0’E,44 m). Laevidentalium martyi Lamprell & Healy,1998. Indo-Pacific: northern Australia. 15-296 m.marukawai, Dentalium (Antalis) Otuka, 1933: 159,160, fig. 1. <strong>Recent</strong>. Off Koshikijima on the westcoast <strong>of</strong> Kyushu, Japan, Soyo-maru stn 425, 200 m.Type material not located. Antalis marukawai(Otuka, 1933) (fide Habe 1963: 264). Indo-Pacific,Japan. 100-400 m. Remark: the type materialdeposited in the “Earthquake Research Institute,Tokyo Imp. Univ.” is now presumed lost, sincethey were not listed by Ichikawa (1983: 463, 464).marwicki, Dentalium Mestayer, 1926: 583, pl. 101,figs 11, 12. Pliocene, Castlecliffian. New Zeal<strong>and</strong>.Holotype NMNZ M1394. Junior synonym <strong>of</strong>Antalis nana (Hutton, 1873) (fide Dell 1957: 570).† masoni, Gadilina Maxwell, 1992: 187, pl. 30, figs g,h. Eocene. NZGS localities GS1986, GS11200,Waihao River, near McCulloch’s Bridge, SouthCanterbury, New Zeal<strong>and</strong>. Holotype NZGSTM7229; numerous paratypes NZGS.† matanzasense, Dentalium Ihering, 1907: 224.Tertiary. Matanzas, Chile. Dentalium matanzasenseIhering, 1907 (fide Ihering 1914: 66). Remark: this<strong>species</strong> was misidentified as Dentalium sulcosumJ. de C. Sowerby, 1846 by Philippi (1887).matara, Dentalium Dall, 1889: 420. <strong>Recent</strong>. Off CapeLookout, USFC stn 2611, 34°15’N, 76°11.30’W,31 fms [57 m]. Lectotype USNM 95363 (designat-614ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)ed by Henderson 1920: 67); paralectotypes USNM92823 (2; USFC stn 2608), USNM 94095(1; USFC 2401), USNM 95364 (1; USFCstn 2409), USNM 95365 (7; Samana Bay, DominicanRepublic). Junior synonym <strong>of</strong> Graptacmeeborea (Conrad, 1846) (fide Scarabino 1974: 185).† mauryae, Dentalium Penna-Neme & da CostaBarros Muniz, 1976: 524, 525, fig. 2. Paleocene,Maria Farinha Formation. Pernambuco, Brazil.Holotype <strong>and</strong> paratypes, Departamento deGeologia, Centro de Tecnologia, UniversidadeFederal de Pernambuco, Brasil.† mawsoni, Dentalium (Fissidentalium) Ludbrook,1956: 2, 3, pl. 1, figs 5, 6. Miocene. River MurrayCliffs, South Australia. Holotype SAM F 15139;paratypes SAM (8), paratypes BMNH (3; LowerBeds, Muddy Creek, South Australia).† mayeri, Dentalium Gümbel, 1861: 745. Oligocene.Lohergraben, Bavaria, Germany.† mayor, Siphondentalium (Gadila) gadus var. Foresti,1895: 258. Pliocene. Italy.mayori, Cadulus (Gadila) Henderson, 1920: 133, 134,pl. 19, fig. 3. <strong>Recent</strong>. East <strong>of</strong> Tortugas, Eolis stn 33,16 fms [29 m]. Holotype USNM 314783;paratypes USNM 314784 (64), USNM 157809(32; USFC 2404), USNM 323953 (14; USFC2404), USNM 314785 (46; Eolis stn 42), USNM314786 (153; Eolis stn 43), USNM 314787 (1;Eolis stn 73), USNM 314788 (42; Eolis stn 48),USNM 314789 (1; Eolis stn 49), USNM 314790(5; Eolis stn 68), USNM 314791 (6; Eolis stn 69),USNM 314792 (2; Eolis stn 87), USNM 314793(1; Eolis stn 117), USNM 314794 (6; Eolisstn 140), USNM 314795 (1; Eolis stn 311), USNM314796 (1; Eolis stn 128), USNM 314797 (2; Eolisstn 147), USNM 314798 (2; Eolis stn 152), USNM314799 (2; Eolis stn 156), USNM 314800 (1; Eolisstn 167), USNM 314801 (1; Eolis stn 172), USNM314802 (3; Eolis stn 179), USNM 314803 (2; Eolisstn 182), USNM 314804 (1; Eolis stn 186), USNM314805 (1; Eolis stn 306), USNM 314806 (1; Eolisstn 354), USNM 314807 (3; Eolis stn 356), USNM314808 (2; Eolis stn 360), USNM 314809 (3; Eolisstn 364), USNM 314810 (16; Eolis stn 368),USNM 314811 (13; Eolis stn 370), USNM 314812(2; Eolis stn 196), USNM 314813 (39; Eolisstn 32), USNM 314814 (2; Eolis stn 100), USNM314815 (2; Eolis stn 300), USNM 314816 (1; Eolisstn 350), MCZ 48411 (6; Eolis stn 43), ANSP44914 (Eolis stn 43), AMNH 148339 (7). Gadilamayori (Henderson, 1920). Western Atlantic,Caribbean. 29-182 m.† mcfarl<strong>and</strong>ae, Cadulus (Cadulus) Emerson, 1957:986, 987, pl. 126, fig. 4. Eocene, Vacaville shale,Capay formation. UCMP loc. A-3365, Ualtis CreekSW <strong>of</strong> Dunns Peak, Vaca Valley, Solano County,California, USA. Holotype UCMP 37590;paratypes UCMP 37591-UCMP 37594.mechelinii, see michelinii.† mediaviense, Dentalium Harris, 1896: 187 [73],pl. 17 [7], figs 1, 1a. Eocene, Midway Stage.0.5 miles N <strong>of</strong> Ripley, Mississippi; Black Bluff,Graveyard Hill, Dale’s Branch <strong>and</strong> Allenton,Alabama, USA. Syntypes PRI 82 [lost], PRI 2349(fide Brann & Kent 1960: 319).mediocarinata, Entalina Boissevain, 1906: 63, textfig.29, pl. 6, figs 70-72, 87, 88. <strong>Recent</strong>. Off KaiIsl<strong>and</strong>s [Indonesia], Siboga stn 256, 5°26.6’S,132°32.5’E, 387 m. Lectotype ZMA 3.06.084 (designatedby Habe: 1964: 42); paralectotypes ZMA3.06.085 (2; Bali Sea, Siboga stn 5, 07°46’S,114°30.5’E, 330 m). Junior synonym: Megaentalinateramachii Kuroda & Habe in Habe, 1963 (fideHabe 1964: 42). Megaentalina mediocarinata(Boissevain, 1906) (fide Kuroda & Habe in Habe1963: 273 as Megaentalina teramachii). Indo-Pacific: eastern Africa to Japan. 200-1035 m.Remark: Scarabino (1995: 366) invalidly selectedone <strong>of</strong> the specimens from Siboga stn 5 as the lectotype.mediopacificensis, Dentalium Rehder & Ladd, 1973:48, 49, pl. 3, figs 1, 2. <strong>Recent</strong>. Agassiz Guyot, Mid-Pacific Mountains, stn 4, 17°58.5’N, 178°14.2’E,865-884 fms [1579-1613 m]. Holotype USNM703273; paratype USNM 703274 (1). Dentaliummediopacificensis Rehder & Ladd, 1973. Knownfrom the type locality only. Remark: V. Scarabino(in litt. 5.XI.2001) suggested that this <strong>species</strong> isreferable to Compressidentalium.† medium, Dentalium J. Sowerby, 1814: 181, pl. 79,fig. 5. Cretaceous. Blackdown, Engl<strong>and</strong>, UnitedKingdom. Remark: Geinitz (1850: 74, pl. 18,figs 25, 26) erroneously used this name for the laterdescribed Dentalium geinitzianum Ryckholt, 1851 .† medius, Cadulus Pilsbry & Sharp, 1898: 237 [exDeshayes MS]. Tertiary. Naples, Italy. Based onDentalium coarctatum sensu O. G. Costa 1851 nonLamarck 1818.† meekianum, Dentalium Geinitz, 1866: 13, pl. 1,fig. 20. Carboniferous. Locality 60, Nebraska City,Missouri River, Nebraska, USA. Plagioglyptameekiana (Geinitz, 1866) (fide Yancey 1978: 310).† meganosensis, Cadulus Palmer, 1923: 303, pl. 54,fig. 5. Middle Eocene. UCMP Locality 3573, nearVacaville, California, USA. Holotype UCMP30601.megathyris, Dentalium Dall, 1890: 293, pl. 9, fig. 1.<strong>Recent</strong>. Localities <strong>of</strong> syntypes: USNM 87558 (OffChiloe Isl<strong>and</strong> [Chile], Albatross stn 2789, 42°35’S,75°28’W, 1342 fms [2449 m]), USNM 95851,594262, ANSP 72361, <strong>and</strong> SBMNH 35017 (nearGalapagos Isl<strong>and</strong>s, USFC stn 2807, 0°24’S,89°06’W, 812 fms [1480 m]), USNM 110665 (<strong>of</strong>fsouthwest Chili [Chile], Albatross stn 2788,45°35’S, 75°55’W, 1050 fms [1916 m]), LACM1816 (1485 m, Islas Galápagos). Fissidentaliummegathyris (Dall, 1890) (fide Pilsbry & Sharp 1897:67). Eastern Pacific: California to Chile, Galapagos.ZOOSYSTEMA • 2004 • 26 (4)615


Steiner G. & Kabat A. R.1480-4300 m. Remark: Scarabino (1995: 250)considered megathyris a junior synonym <strong>of</strong> Fissidentaliumceras Watson, 1879. This proposed synonymyrequires a comparative study <strong>of</strong> all taxaincluded in this <strong>species</strong> <strong>group</strong>.† menthifonta, Fustiaria (Episiphon) Dockery inMacNeil & Dockery, 1984: 248, pl. 71, figs 1-3.Lower Oligocene, Mint Spring Formation.Mississippi Geological Survey locality 90, IchusaCreek, Smith County, Mississippi, USA. HolotypeUSNM Paleobiology 376681.meridionale, Dentalium (Fissidentalium) c<strong>and</strong>idum var.Pilsbry & Sharp, 1897: 73, pl. 15, figs 32-34. <strong>Recent</strong>.SE <strong>of</strong> Rio de Janeiro, Brazil, Albatross stn 2763,24°17’S, 42°48.30’W, 671 fms [1225 m]. HolotypeUSNM 887471; paratypes USNM 87538, 87557.Synonym <strong>of</strong> Fissidentalium c<strong>and</strong>idum (Jeffreys,1877) (fide Scarabino 1994: 307).metivieri, Fissidentalium Scarabino, 1995: 254, 255,figs 61, 62a, 70d, e. <strong>Recent</strong>. Madagascar, MozambiqueChannel, 22°24’S, 43°03’E, 600 m.Holotype MNHN; paratypes MNHN (1 from stn106; 1 from stn 65; 1 from stn 76; 1 from stn 84;2 from stn 88; 1 from stn 91; 2 from stn 126; 1from stn CH49), NM (1; stn 91), USNM 890876(1; stn 104), BMNH 1994041 (2; stn CH102).Fissidentalium metivieri Scarabino, 1995. Indo-Pacific: Madagascar. 310-1020 m.† mexicanum, Dentalium Girty, 1909: 95, pl. 11,fig. 10. Carboniferous, San Andreas Formation.USGS localities 3738a (Caballos Mountain) <strong>and</strong>3743 (Fra Cristobal), New Mexico, USA. Typematerial USNM Paleobiology 120807 (CaballosMountain).mexicanum, Dentalium (Tesseracme) Perrilliat, 1975:57, 58, pl. 10, figs 3-5. Middle Miocene. Head <strong>of</strong>stream in Arroyo Tomás Martínez, about 3 km NE<strong>of</strong> Santa Rosa, <strong>and</strong> approximately 28 km NW <strong>of</strong>Santa Lucrecia, by the railroad track, state <strong>of</strong>Veracruz, Mexico. Holotype USNM Paleobiology350085; paratypes UNAM 2520. Junior homonym<strong>of</strong> Dentalium mexicanum Girty, 1909. Replacementname: Tesseracme perrilliatae n. nom.† meyeri, Entalis Gardner, 1878: 62, pl. 3, fig. 40.Cretaceous. Blackdown beds, Engl<strong>and</strong>, UnitedKingdom.† meyeri, Siphonodentalium (Siphonodentalium)Cossmann, 1888: 12, 13, pl. 1, figs 4, 5. Eocene.Chaussy <strong>and</strong> Houdan, France. Cadulus meyeri(Cossmann, 1888) (fide Pilsbry & Sharp 1898:237). Remark: a valid <strong>species</strong> <strong>of</strong> Polyschides (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 86).miamiensis, Cadulus (Platyschides) Henderson, 1920:129, 130, pl. 19, fig. 18. <strong>Recent</strong>. Off Fowey Light,Florida, Eolis stn 340, 209 fms [381 m]. HolotypeUSNM 314772; paratypes USNM 314772a (24),USNM 314769 (2; Eolis stn 303), USNM 314770(1; Eolis stn 304), USNM 314771 (1; Eolisstn 306), USNM 314773 (16; Eolis stn 347),USNM 314774 (15; Eolis stn 348), USNM 314775(1; Eolis stn 371), USNM 314776 <strong>and</strong> USNM108176a (2; Eolis stn 353), USNM 314777 (10;Eolis stn 377), USNM 314778 (6; Eolis stn 378),USNM 314779 (1; Eolis stn 379), USNM 314780(3; Eolis stn 381), USNM 314781 (9), USNM330522 (2; USBF stn 2644), MCZ 48413 (2; Eolisstn 348), AMNH 148351 (2), ANSP 44912 (Eolisstn 348). Polyschides miamiensis (Henderson,1920) (fide Emerson in Turgeon 1988: 51).Western Atlantic, Caribbean. 155-381 m.† michauxianum, Dentalium Ryckholt, 1851: 72, 73,pl. 2, figs 47, 48. Cretaceous, Turonian. Liège,Belgium. Remark: based on Dentalium ellipticumsensu Reuss 1845 non J. Sowerby, 1814.† michelinii, Dentalium Rouault, 1850: 473, pl. 15,figs 6a-6c. Eocene. Bos d’Arros, near Pau, France.Dentalium mechelinii [Pilsbry & Sharp, 1898: 208]is an incorrect subsequent spelling.michelottii, Dentalium Hörnes, 1856: 654, figs 84-86.Miocene. Baden, Vienna Basin, Austria. LectotypeNHM 1846.37.544 (designated by Pavia 1991:111) + 11 paralectotypes. Tentative junior synonym:Dentalium dumasi Cossmann & Peyrot, 1938 (fidePavia 1991: 111). Dentalium michelottii Hörnes,1856. Indian Ocean (0°59’S, 129°48.8’E). 411 m.Remark: this Miocene <strong>species</strong> was reported fromthe <strong>Recent</strong> fauna <strong>of</strong> the Indian Ocean by Boissevain(1906: 16).micra, Entalinopsis Scarabino, 1995: 306, 307,figs 109, 111 c, d, 115c. <strong>Recent</strong>. South NewCaledonia, RV Vauban stn 40, 22°30’S, 166°24’E,250-350 m. Holotype <strong>and</strong> 6 paratypes MNHN.Entalinopsis micra Scarabino, 1995. Indo-Pacific:New Caledonia. 250-350 m.microceras, Siphonodentalium Meyer, 1883: 259 [exBoettger MS]. Oligocene. Joachimsthal, Germany.Nomen nudum.† microreticulata, Antalis “?” Welle, 1997: 93, 94,pl. 13, figs 10, 11. Upper Oligocene. GrafenbergerS<strong>and</strong>. Schacht 8, Bergwerksgesellschaft SophiaJacoba, near Erkelenz (N <strong>of</strong> Aachen), Germany.Holotype GPIM D.8-9-1.192 ; paratypes GPIM (6).† microstria, Dentalium Heilprin, 1881: 375, pl. 20,fig. 3 [original spelling micro-stria]. Eocene. CaveBranch <strong>and</strong> Wood’s Bluff, Clarke County,Alabama, USA. Holotype Geological Survey <strong>of</strong>Alabama, No. 20 (fide Palmer & Brann 1965: 367).milneedwardsi, Dentalium Locard, 1897b: 10, 11[Locard 1898: 113, pl. 6, figs 21-24]. <strong>Recent</strong>.Talisman stn 78, 23°55’N, 19°35’W, “Sur les côtesoccidentales d’Afrique, au large du Soudan”[Morocco], 1435 m. Syntypes MNHN. Junior synonym<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877)(n. syn. B. Métivier, in litt. 3.IV.2000). Describedfrom the type locality only.† minimum, Dentalium Buckman in Murchison,1845: 101 [ex Strickl<strong>and</strong> MS]. Jurassic. Cheltenham,Engl<strong>and</strong>, Great Britain. Neotype Museum <strong>of</strong>616ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Practical Geology, London, No. 5799 (designatedby Richardson 1906: 584) (now in BMNH?). This isalso the holotype <strong>of</strong> Dentalium portlocki Tate, 1870.Junior objective synonym: Dentalium portlocki Tate,1870. Laevidentalium minimum (Buckman inMurchison, 1845) (fide Engeser & Riedel 1992: 38).minimum, Dentalium Etheridge & Bell in Bell, 1898:156, 157. Pliocene. St Erth, Cornwall, Engl<strong>and</strong>,United Kingdom. Junior homonym <strong>of</strong> Dentaliumminimum Buckman in Murchison, 1845. Remark:Pilsbry & Sharp (1898: 310) stated that: “Thename is preoccupied; but as this is probably not avalid <strong>species</strong>, it need not be renamed”.minimum, Siphonodentalium Plate, 1908b: 4, 5, fig. 5.<strong>Recent</strong>. Antarctica, Gauss Winterstation, 66°02’S,89°38’W, 3423 m. Lectotype ZMB 59728a (designatedby Kilias 1995:175); paralectotype ZMB59728b. Striopulsellum minimum (Plate, 1908)(fide Scarabino 1995: 323). Circumantarctic; Indo-Pacific: New Caledonia. 3423-6179 m.† minoensis, Siphonodentalium (Pulsellum) Itoigawa,1960: 274, pl. 3 figs 2, 3. Akeyo Miocene. Stn S11-1, Kujiri facies, near Mizunami, Honshu, Japan.Holotype ESN 20031; paratype ESN 20032.† minor, Cadulus gracilina var. Báldi, Horvath &Makk, 1974: 27. Oligocene, Kiscellian. Budafok,S <strong>of</strong> Budapest, Hungary.minor, Cadulus monterosatoi var. Locard, 1898: 142.<strong>Recent</strong>. Type locality not specified [North Atlantic].Type material not located. Synonym <strong>of</strong> Cadulusmonterosatoi Locard, 1897a.minor, Cadulus olivi var. Locard, 1898: 134, pl. 7,figs 10-12. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Type material not located.Synonym <strong>of</strong> Polyschides olivi (Scacchi, 1835).minor, Cadulus tumidosus var. Locard, 1898: 143[nomen nudum in Jeffreys 1883: 665]. <strong>Recent</strong>. Typelocality not specified [North Atlantic]. Type materialnot located. Synonym <strong>of</strong> Cadulus tumidosusJeffreys, 1877.minor, Dentalium agile var. Locard, 1898: 117, 118.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Antalis agilis (G. O. Sars ex M. Sars, 1872).minor, Dentalium caudani var. Locard, 1898: 104.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Syntype MNHN. Synonym <strong>of</strong> Dentaliumcaudani, a junior synonym <strong>of</strong> Fissidentaliumc<strong>and</strong>idum (Jeffreys, 1877).minor, Dentalium ergasticum var. Locard, 1898: 106.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium ergasticum, a junior synonym <strong>of</strong>Fissidentalium capillosum (Jeffreys, 1877).minor, Dentalium exuberans var. Locard, 1898: 112.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium exuberans, a junior synonym <strong>of</strong> Fissidentaliumpaucicostatum (Jeffreys, 1877).minor, Dentalium panomitanum var. Locard, 1898:123. <strong>Recent</strong>. Type locality not specified. Typematerial not located. Synonym <strong>of</strong> Antalis panorma(Chenu, 1843).minor, Dentalium scamnatum var. Locard, 1898: 110.<strong>Recent</strong>. Type locality not specified [North Atlantic].Syntype MNHN. Synonym <strong>of</strong> Dentalium scamnatum,a junior synonym <strong>of</strong> Fissidentalium capillosum(Jeffreys, 1877).minor, Dentalium semivestitum var. Locard, 1898: 108,pl. 6, figs 28, 29. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Type material not located.Synonym <strong>of</strong> Fissidentalium semivestitum (Fischerin Locard, 1898).minor, Dentalium Stephenson, 1952: 143, pl. 34,figs 4, 5. Upper Cretaceous, Cenomanian, WoodbineFormation. Tarrant County Locality 38,Lewisville member, near railroad, 0.9 mile W <strong>of</strong>Dallas County border, Tarrant County, Texas,USA. Holotype USNM Paleobiology 105610;paratypes USNM Paleobiology 105611-105612(Tarrant County localities 41 <strong>and</strong> 46). Junior homonym<strong>of</strong> Dentalium caudani var. minor Locard,1898 (<strong>and</strong> seven other minor varieties in the sameLocard paper). Replacement name: Dentaliumminorinum n. nom.minor, Pseudantalis rubescens var. Monterosato, 1884:33. Nomen nudum without type locality.minor, Siphodentalium [sic] l<strong>of</strong>otense var. Monterosato,1878: 76. <strong>Recent</strong>. Mediterranean <strong>and</strong> Adriatic Seas.Nomen nudum.† minorinum, Dentalium n. nom. Upper Cretaceous,Cenomanian, Woodbine Formation. TarrantCounty Locality 38, Lewisville member, near railroad,0.9 mile W <strong>of</strong> Dallas County border, TarrantCounty, Texas, USA. Holotype USNM Paleobiology105610; paratypes USNM Paleobiology105611-105612 (localities 41 <strong>and</strong> 46). Replacementname for Dentalium minor Stephenson,1952 non Dentalium caudani var. minor Locard,1898 (<strong>and</strong> seven other minor varieties in the sameLocard paper). Dentalium minorinum Steiner &Kabat, 2004.minus, Dentalium Boissevain, 1906: 14, pl. 6, fig. 3,text-fig. 10. <strong>Recent</strong>. Java Sea, Siboga stn 319,6°16.5’S, 114°37’E, 82 m. Lectotype ZMA3.06.008 (designated by Scarabino 1995: 218);paralectotype ZMA 3.06.009 (1). Junior synonym<strong>of</strong> Paradentalium hexagonum (Gould, 1859) (fideScarabino 1995: 218).minusculus, Cadulus Dall, 1889: 432. <strong>Recent</strong>. OffCape Hatteras, USA, USBF stn 2595, 35°08’N,75°05’30”W, 63 fms [115 m]. Lectotype USNM93122 (designated by Henderson 1920: 134); paralectotypeUSNM 887459 (6). Cadulus minusculusDall, 1889. Known from the type locality only.minutalis, Gadila Scarabino, 1995: 362, 363, figs 167,169g. <strong>Recent</strong>. Réunion, MD32 stn DS151,20°51’S, 56°03’E, 3300-3240 m. Holotype <strong>and</strong>ZOOSYSTEMA • 2004 • 26 (4)617


Steiner G. & Kabat A. R.8 paratypes MNHN. Gadila minutalis Scarabino,1995. Indo-Pacific: Réunion Isl<strong>and</strong>. 3240-3300 m.† minutiannulatum, Dentalium (Antalis) Hodgkinson,1974: 20, 21, text-fig. 7f-i; pl. 4, figs 2-7, pl. 6,figs 1-3, pl. 7, fig. 2. Middle Eocene, ClaibornianStage, Cook Mountain Formation, WheelockMember. Little Brazos River Valley, Brazos County,Texas, USA. Holotype USNM Paleobiology180467; paratypes USNM Paleobiology 180466,180468-180471, 180474-180476, PRI 29228;ANSP 31506.minutissimum, Dentalium (Episiphon) Ludbrook,1954: 108, fig. 11. <strong>Recent</strong>. Maldives Is., JohnMurray Exp. stn 147, 04°53’N, 72°54’E, 27 m.Holotype BMNH 1952.3.25.123; paratypesBMNH 1952.3.25.113-122 (9). Antalis(?) minutissima(Ludbrook, 1954) (fide V. Scarabino, in litt.20.II.2002, tentatively as Episiphon in Scarabino1995: 289). Known from the type locality only.minutissimus, Cadulus (Gadila) Niino & Habe inHabe & Kosuge, 1964: 11. Nomen nudum. <strong>Recent</strong>.Off Puket [Thail<strong>and</strong>], 50-80 m.† minutistriatum, Dentalium Gabb, 1860: 386, pl. 67,fig. 46. Eocene, Cook Mountain Formation.Wheelock, Robertson County, Texas, USA.Holotype ANSP 13264 (“broken” fide Hodgkinson1974: 22). Dentalium (Antalis) minutistriatumGabb, 1860 (fide Hodgkinson 1974: 21, 22).minutu[m], Dentalium Linnaeus, 1758: 786 [citingPlancus 1739: t. 2., f. 2]; emended to minutum byLinnaeus (1767: 1264). <strong>Recent</strong>. Mediterranean.Syntype LS 615. Possible junior synonym: Gadilagadus (Montagu, 1803) (fide Pilsbry & Sharp 1898:243, who listed Dentalium minutum under spurious<strong>Scaphopoda</strong>, “[...] is not identifiable, but Hanleysurmises that it may be Cadulus gadus”).minutus, Cadulus H. Adams, 1872: 10, pl. 3, fig. 9.<strong>Recent</strong>. Red Sea. Syntypes BMNH 1872.4.9.6 (4),UMZC I.100,755. Dischides minutus (H. Adams,1872) (fide Scarabino 1995: 340). Indian Ocean:Red Sea <strong>and</strong> Gulf <strong>of</strong> Aden. Bathymetric rangeunknown; type specimens “probably washedashore” (Scarabino 1995: 340).minutus, Dentalites Schlotheim, 1813: 91. Cretaceous.Insel Moen, Denmark. Nomen nudum.† miocaenicum, Pulsellum Boettger, 1902: 182; 1907:211. Miocene. Kostej <strong>and</strong> Lapugy, Banat, Romania.Lectotype SMF XIII.3a (designated by Zilch 1934:279, pl. 22, fig. 27). Episiphon miocaenicum(Boettger, 1902) (fide Baluk 1972: 560, 561).† miocenicum, Dentalium Michelotti, 1847: 144, 145,pl. 16, fig. 12. Miocene. Tortona, Italy. Entalismiocaenica (Michelotti, 1847) (fide Cossmann &Peyrot 1917: 176-178); Dentalium (Entalis)miocenica Michelotti, 1847 (fide Sacco 1897: 108).miopseudodentalis, see miopseudoentalis.† miopseudoentalis, Dentalium (Entalis) Sacco, 1897:106, pl. 9, figs 7-11 [as a nomen nudum in Sacco1896: 97]. Miocene. Turin hills, Italy. SyntypesMIGT BS.106.03.003-BS.106.03.005. Entalis miopseudodentalis[Ferrero Mortara et al. 1984: 306] isan incorrect subsequent spelling.mirificum, Dentalium E. A. Smith, 1895: 9, pl. 2,fig. 1. <strong>Recent</strong>. Ceylon [Sri Lanka], <strong>of</strong>f Trincomalee,Investigator stn 172, 200-350 fms [365-639 m].Lectotype BMNH 1895.7.2.26 (designated byLamprell & Healy 1998: 137 as “holotype”. Their“paratype” could not be located.); paralectotype ZSIM194/1 (1). Junior synonyms: Entalina quadrangularisBoissevain, 1906 (fide Scarabino 1995: 300);E. majestica Kira, 1959 (fide Habe 1964: 39; Scarabino1995: 147). Entalina mirifica (E. A. Smith,1895) (fide Pilsbry & Sharp 1897: 134). Indo-Pacific: from Africa to New Caledonia. 55-2050 m.Remarks: two <strong>of</strong> the four cited type specimens couldnot be located; Scarabino’s (1995: 300) record <strong>of</strong>four syntypes in the BMNH cannot be confirmed.† misatoensis, Dentalium Araki, 1958: 164, 165,pl. 18, figs 5a, 5b. Miocene. Kaisekizan Formation,near Funayama, Misato-mura, Age-gun, MiePrefecture, Japan. Holotype Geol. Dept., MieUniv. (fide Hanzawa et al. 1961: 209).† mississippiensis, Dentalium Conrad, 1848a: 282;1848b: 112, 113, pl. 11, fig. 1. “Eocene” [Oligocene].Vicksburg, Mississippi, USA. Holotype <strong>and</strong>paratype ANSP 30659 (fide MacNeil & Dockery1984: 245). Tentative senior synonym: Dentaliumvirginianum Chenu, 1843 (fide Pilsbry & Sharp1898: 209). Junior synonym: Dentalium densatumConrad, 1865. Dentalium mississippiense Conrad,1848 (fide Pilsbry & Sharp 1898: 209).† missouriense, Dentalium Swallow, 1863: 99.Carboniferous. Archimedes Limestone, Chester,Illinois [USA] <strong>and</strong> St Mary’s, Missouri [USA].modicellum, Dentalium Kurtz, 1860: 6. SouthCarolina. Nomen nudum.† molengraaffi, Dentalium (s.s.) Martin, 1915: 179,pl. 7, figs 178, 179. Eocene. Kali Puru, Nanggulan,Java, Indonesia. Syntypes RGM 7265 (1), RGM7264 (2), RGM 7268 (3), RGM 7261 (1), RGM7267 (3), RGM 7259 (3), RGM 7266 (1), RGM7263 (1) (fide Van den Hoek Ostende et al. 2002:111, 112).monodonta, Gadila Scarabino, 1995: 363, figs 168,169i. <strong>Recent</strong>. Mozambique Channel, West [<strong>of</strong>]Glorieuses Is., BENTHEDI stn DS10, 11°28.5’S,47°17.7’E, 440 m. Holotype MNHN; paratypesMNHN (1 each from DS120 <strong>and</strong> DS122). Gadilamonodonta Scarabino, 1995. Indo-Pacific:Madagascar to New Caledonia. 390-625 m.monolineata, Plagioglypta Branson, 1930: 58, 59,pl. 15, fig. 7. Late Pennsylvanian to early Triassic.Top Limestone, Phosphoria Formation, WindRiver <strong>and</strong> Owl Creek Mountains, Wyoming, USA.Junior synonym <strong>of</strong> Prodentalium canna (White,1874) (fide Yancey 1973: 1126; 1978: 308, 309).† montense, Dentalium Briart & Cornet, 1889: 80,pl. 24, figs 12a, 12b. Upper Eocene. Mons, Belgium.618ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Tentative junior synonym: Fustiaria inversa(Cossmann, 1915) (n. syn.). Fustiaria montense(Briart & Cornet, 1889). Remark: Cossmann(1915: 6, 7) noted that the only difference betweenDentalium (Fustiaria) inversum <strong>and</strong> Dentaliummontense is the position <strong>of</strong> the apical slit.monterosatoi, Cadulus Locard, 1897a: 4. <strong>Recent</strong>. West<strong>of</strong> Cape Finistere [Cabo Finisterra], Spain,Travailleur 1881 dr. 1, 2018 m. Type material notlocated. Synonym: Cadulus monterosatoi var. ventricosaLocard, 1898. Cadulus monterosatoi Locard,1897. Eastern Atlantic: Bay <strong>of</strong> Biscay to Portugal.896-3307 m.† monterosatoi, Dentalium rectum var. Pilsbry &Sharp, 1898: 214. Tertiary. Sicily, Italy. Replacementname for Dentalium philippi Monterosato,1872 non Chenu, 1843.moorei, Dentalium Pilsbry & Sharp, 1898: 224.Cretaceous. Wollumbilla, Queensl<strong>and</strong>, Australia.Replacement name for Dentalium lineatum Moore,1870 non Gueranger, 1853. Objective junior synonym<strong>of</strong> Dentalium wollumbillaensis Etheridge inJack & Etheridge, 1892. Remark: Pilsbry & Sharp(1898) inadvertently overlooked the earlier replacementname.† moreanum, Dentalium d’Orbigny, 1845: 454,pl. 38, fig. 10. Jurassic. Kaminka, Russia; Vieil-Saint-Rémy, Ardennes, France. Remark: this <strong>species</strong>was misidentified as Dentalium cylindricum J.Sowerby, 1814 by Fischer (1845).moreauanum, Dentalium Bronn, 1848: 415; 1849:359. Unjustified emendation <strong>of</strong> Dentalium moreanumd’Orbigny, 1845.† morganianum, Dentalium (Laevidentalium)Wilckens, 1922: 24, 25, pl. 5, figs 8, 9. Cretaceous,Upper Senonian. NZGS Localities 5 <strong>and</strong> 13 (AmuriBluff) <strong>and</strong> NZGS locality 589 (Selwyn Rapid beds,Selwyn River, Malvern Hills), New Zeal<strong>and</strong>.mosae, Dentalium – Bronn 1837: 706, pl. 32, fig. 18,non Pyrgopolon mosae Montfort, 1808 (Polychaeta).Cretaceous. Petersberges near Maastricht (Netherl<strong>and</strong>s);Ciply, near Mons (Belgium); Köpingemolla,Schoonen (Sweden?). Remark: Hisinger (1840:Supp., p. 8) suggested that Bronn’s specimens couldbe equivalent to Dentalium browni Hisinger, 1837(Polychaeta); but Goldfuss (1841: 3, pl. 166,fig. 10) used mosae as a valid <strong>species</strong> <strong>of</strong> Dentalium.mosae, Pyrgopolon Montfort, 1808. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† moseleyensis, Cadulus (Gadila) Hodgkinson, 1974:17, text-fig. 8g, h; pl. 1, figs 3, 4, pl. 2, fig. 3.Middle Eocene, Claibornian Stage, Cook MountainFormation, Wheelock Member. Stone City Bluff,Brazos River Valley, Burleson County, Texas, USA.Holotype USNM Paleobiology 180428; paratypesUSNM Paleobiology 180427, 180444, PRI 29221,ANSP 31499.motidukii, Dentalium (Antalis) Otuka, 1935: 879,880, pl. 54, fig. 89. <strong>Recent</strong>. Hiradoko, Japan.Holotype GIY 2400. Junior synonym <strong>of</strong> Dentaliumbuccinulum Gould, 1859 (fide Habe 1964: 25).† mucronatus, Cadulus Tate, 1887: 193, 194, pl. 20,fig. 10. Lower Miocene. Muddy Creek, Victoria,Australia. Holotype <strong>and</strong> 14 paratypes SAM T229A. Cadulus (Gadila) mucronatus Tate, 1887(fide Ludbrook 1959: 147, pl. 1, fig. 8).† muellerianum, Dentalium Pilsbry & Sharp, 1898:224. Cretaceous. Königsthore, Germany. Replacementname for Dentalium alternans Müller, 1849non Chenu, 1843.muensteri, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 224. Jurassic. Derneberg, nearHannover, Germany. Replacement name forDentalium cinctum “Goldfuss, 1844” [sic!, shouldbe 1841] non de Koninck, 1843. Remark: Pilsbry &Sharp (1898: 224) erroneously dated this name to1844, <strong>and</strong> therefore concluded that it was a juniorhomonym <strong>of</strong> Dentalium cinctum de Koninck, 1843.Thus, Pilsbry & Sharp proposed Dentalium muensterias a replacement name for what is, in fact, thesenior homonym.† multannulatum, Dentalium Aldrich, 1895: 55 [3],pl. 2 [1], fig. 3. Eocene. Gregg’s L<strong>and</strong>ing, Alabama,USA. Lectotype USNM Paleobiology 638920 (designatedby Palmer & Brann 1965: 369 as “holotype”).† multicanaliculatum, Dentalium Gümbel, 1861: 572.Cretaceous. Siegsdorf, Bavaria, Germany.† multiclathratum, Dentalium Martelli, 1905: 351,pl. 11, fig. 39. Triassic, Ladinian. Wengen,Montenegro.† multicostatum, Dentalium Favre, 1869: 101, pl. 12,fig. 8. Cretaceous. Nagorzany, Lviv [Lemberg],Ukraine. Holotype NHM 1862/0005/0130.Remark: this <strong>species</strong> was misidentified as Dentaliumdecussatum Sowerby, 1814 by Alth (1850).multicostata, Dentalium arctoides var. Gugenberger,1934: 43, pl. 1, fig. 15 [nomen nudum inGugenberger 1933a: 102; 1933b: 184]. Triassic.N <strong>of</strong> Launsdorf, Austria. Junior homonym <strong>of</strong>Dentalium multicostatum Favre, 1869; replacementname: Prodentalium angelae Engeser & Riedel,1992.multistriatum, Dentalium Deshayes, 1825: 358, pl. 18,fig. 11. <strong>Recent</strong>. “[presumably from India]” [label <strong>of</strong>type lot: Île de France = Mauritius]. LectotypeMNHN (designated by Scarabino 1995: 201);paralectotypes MNHN (13). Junior synonym <strong>of</strong>Dentalium variabile Deshayes, 1825 (fideScarabino 1995: 200, by First Reviser action,according to ICZN 1999: article 24).† multistrictum, Laevidentalium Finlay & Marwick,1937: 93, pl. 13, fig. 3. Paleocene. Wangaloa, NewZeal<strong>and</strong>. Holotype NZGS TM4811; paratypesNZGS (2) (fide Keyes 1972: 96). Antalis multistricta(Finlay & Marwick, 1937) (fide Maxwell 1988: 76).† mutabile, Dentalium Hörnes, 1856: 654, pl. 50,fig. 32 [ex Doderlein MS]. Miocene, Badenian.ZOOSYSTEMA • 2004 • 26 (4)619


Steiner G. & Kabat A. R.Steinabrunn, Wiener Becken, Austria. LectotypeNMW 1855.XLV.271 (selected by Pavia 1991:124); paralectotypes NMW 1855.XLV.271a-d (4).Fissidentalium mutabile (Hörnes, 1856) (fide Pavia1991: 124-126).nagoense, Dentalium Dall, 1927: 1. <strong>Recent</strong>. Nago,Okinawa, Ryukyu Isl<strong>and</strong>s, 15 fms [27 m]. HolotypeUSNM 333718. Junior synonym <strong>of</strong> Fustiarianipponica (Yokoyama, 1922) (fide Habe 1963:266).† nanaimoensis, Dentalium Meek, 1858: 44, 45.Cretaceous. Nanaimo, Vancouver Isl<strong>and</strong>, BritishColumbia, Canada. Holotype USNM Paleobiology4270. Junior synonym: Dentalium komooksenseMeek, 1876 (fide Pilsbry & Sharp 1898: 225).Dentalium nanaimoense Meek, 1858 (fide Pilsbry &Sharp 1898: 225).† nanggulanense, Dentalium (s.s.) Martin, 1915: 179,180, pl. 7, figs 180-182. Eocene. Sumatra, Indonesia.Syntypes RGM 7250 (17), RGM 7245 (3),RGM 7247 (2), RGM 7248 (5), RGM 47182 (5),RGM 7246 (2), RGM 7251 (1), RGM 7249 (4)(fide Van den Hoek Ostende et al. 2002: 112).Remark: this <strong>species</strong> was misidentified as Dentaliumjunghuhni by Boettger (1882).nanum, Dentalium Hutton, 1873b: 73, pl. 8, fig. 78.Castlecliffian, Pliocene. New Zeal<strong>and</strong>. LectotypeNZGS TM.4807 (designated by Keyes 1972: 96).Junior synonyms: Dentalium huttoni Kirk, 1880;Dentalium marwicki Mestayer, 1926 (fide Dell1957: 570). Antalis nana (Hutton, 1873) (fide Dell1957: 570). New Zeal<strong>and</strong>; Antarctica. 30-320 m.nanus, Cadulus (Gadila) iota Clench & Aguayo, 1939:197. <strong>Recent</strong>. Gibara, Oriente Province, Cuba,40 fms [73 m]. Holotype MCZ 89222; 1 paratypeMuseo Poey, Habana (Havana), Cuba. Synonym <strong>of</strong>Gadila iota Henderson, 1920.natalense, Dentalium Barnard, 1963b: 350, fig. 30e.<strong>Recent</strong>. South Africa, <strong>of</strong>f Cape Natal, Durban,85 fms [153 m]. Syntypes BMNH 1964257 (6),SAFM A9364 (49) (fide Giles & Gosliner 1983:41). Paradentalium natalense (Barnard, 1963) (fideScarabino 1995: 219). Indian Ocean: South Africato Madagascar. 50-670 m.† navarroi, Dentalium Stephenson, 1941: 253, pl. 46,figs 16, 17. Cretaceous, Navarro Group, KempClay. USGS locality 762, Nacatoch s<strong>and</strong> nearChatfield, Navarro County, Texas, USA. HolotypeUSNM Paleobiology 76767, paratypes USNMPaleobiology 76768 (1); 76769 (1).† navicanum, Dentalium Ryckholt, 1851: pl. 10,fig. 2; 1854: 198. Carboniferous. Visé, Belgium.† navidadense, Dentalium Pilsbry & Sharp, 1898:210. Tertiary. Navidad, Chile. Replacement namefor Dentalium gracile Philippi, 1887 non Dentaliumgracilis Hall & Meek, 1855 nec Dentalium gracileMoore, 1866 nec Dentalium gracile Jeffreys, 1870.nebulosum, Dentalium Gmelin, 1791: 3738. [No referencesgiven]. <strong>Recent</strong>. Sicily [Mediterranean]. Juniorsynonym <strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fidePilsbry & Sharp 1897: 42).nedallisoni, Cadulus (Polyschides) Emerson, 1978:120-122, figs 4-6. <strong>Recent</strong>. Corinto, Nicaragua,Zaca Exp. stn 200-D-19, 12°28’03”N, 87°12’39”W,22-24 m. Holotype AMNH 160349; paratypesAMNH 183875 (30), LACM 2117 (2). Polyschidesnedallisoni (Emerson, 1978). Eastern Pacific:Nicaragua. 22-24 m.† neglectum, Pulsellum Cossmann, 1888: 11, pl. 1,figs 2, 3. Eocene. Le Guépelle, Marines <strong>and</strong>Chaussy, France. Siphonodentalium neglectum(Cossmann, 1888) (fide Pilsbry & Sharp 1898:235). Remark: a valid <strong>species</strong> <strong>of</strong> Dischides (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 86).neohexagonum, Dentalium (Dentalium) Sharp &Pilsbry in Pilsbry & Sharp, 1897: 19, 20, pl. 11,figs 74-86. <strong>Recent</strong>. Santa Barbara; San Pedro Bay;San Diego [all California]. Syntypes ANSP 35459(4). Dentalium neohexagonum Sharp & Pilsbry inPilsbry & Sharp, 1897. Eastern Pacific: California.0-145 m.† neornatum, Dentalium Hayasaka, 1925: 108, 109[24, 25], pl. 8 [1], figs 18-18b. Permian.Kinshozan, Akasaka-Machi, Mino province, Japan.Holotype Instn Geol. Paleont., Tohoku Univ. (fideHanzawa et al. 1961: 209).newcombei, Cadulus (Gadila) tolmiei var. Pilsbry &Sharp, 1898: 182, pl. 34, figs 1, 2. <strong>Recent</strong>. Typelocality not specified. Type material not located.Synonym <strong>of</strong> Gadila tolmiei (Dall, 1897).† newtonensis, Cadulus Meyer & Aldrich, 1886: 41,pl. 2, figs 3, 3a, 3b. Eocene. Newton, NewtonCounty, Mississippi, USA. Syntype USNMPaleobiology 638727. Remarks: this name was tentativelyproposed for “two depressed fragments” <strong>of</strong>Cadulus: “If, however, the form should prove to bea new <strong>species</strong> we propose the name Cadulus newtonensisfor it”. Pilsbry & Harbison (1933: 117) suggestedthat it is a junior synonym <strong>of</strong> Cadulusjacksonensis Meyer, 1885, but Palmer & Brann(1965: 362) listed both <strong>species</strong> as valid.† niceforoi, Dentalium Royo y Gomez, 1945: 460:pl. 70, fig. 1. Lower Cretaceous, Barremian. TowerVilleta Formation, San Gil, Departmento deSant<strong>and</strong>er, Colombia.† nicense, Dentalium Bellardi, 1852: 229, pl. 15,figs 9, 10 [as niceense, a nomen nudum in Bellardi1850: 681]. Oligocene. Palarca <strong>and</strong> Mortola, Italy.Dentalium nitense [Gümbel, 1861: 604] is an incorrectsubsequent spelling (fide Pilsbry & Sharp 1898:210).nicklesi, Cadulus Dell, 1964: 129, 130, pl. 7, fig. 5.<strong>Recent</strong>. Cape Lopez, Angola, Discovery stn 279, 58-67 m. Holotype BMNH 1962883/1; paratypesBMNH 1963300-303 (1 <strong>of</strong> them from stn 274,St Paul de Lo<strong>and</strong>a, Angola, 65 m). Gadila nicklesi(Dell, 1964) (fide Nicklès 1979: 69). EasternAtlantic: West Africa. 29-200 m. Remark: Nicklès620ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)(1979) listed this <strong>species</strong> as C. (Platyschides) nicklesi,<strong>and</strong> the subgenus Platyschides was put in synonymywith Gadila by Scarabino (1995: 196, 355).nicolasi, Annulipulsellum Palmer, 1986: 396 [exScarabino MS]. Nomen nudum.nielseni, Graptacme Lamprell & Healy, 1998: 98, 99,figs 99B, 100A, 102A-D, 103. <strong>Recent</strong>. Off Cairns,northern Queensl<strong>and</strong>, Australia, 11°22’S, 144°35’E,2016 m. Holotype QM MO40055; paratypes QMMO40057 (8), QM MO40056 (1; northernQueensl<strong>and</strong>, 14°27’S, 146°51’E, 1855 m), QMMO40058 (3; northern Queensl<strong>and</strong>, 11°23’S,144°36’E, 1999 m), AMS C201736 (2; northernQueensl<strong>and</strong>, 11°23’S, 144°36’E, 1999 m).Graptacme nielseni Lamprell & Healy, 1998. Indo-Pacific: northern Australia. 1855-2016 m.† nigeriense, Dentalium (Laevidentalium) Adegoke,1977: 63, 64, pl. 7, figs 5-10. Paleocene. Nigeria.Holotype UIMG 179; paratypes UIMG 180,UIMG 181, USNM Paleobiology 174765, 174766.nigr<strong>of</strong>asciatum, Dentalium Eichwald, 1830. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.nigrum, Dentalium Lamarck, 1818. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.nipponicum, Dentalium (Fustiaria) Yokoyama, 1922:119, pl. 6, fig. 7. Pleistocene. Shito, Honshu, Japan.Lectotype UMT CM.21050 (designated by Taki &Oyama 1954: caption to pl. 26, fig. 7, as “holotype”;see Oyama [1973: 3] for an explanation <strong>of</strong>Taki & Oyama’s [1954] lectotype designations).Junior synonyms: Dentalium nagoense Dall, 1927;Dentalium numatai Hirase, 1931 (fide Habe 1963:266, 267). Fustiaria nipponica (Yokoyama, 1922)(fide Habe 1963: 266). Indo-Pacific: Indonesia toJapan <strong>and</strong> New Caledonia. 5-200 m.† nipponicum, Siphonodentalium Makiyama, 1927:59, pl. 2, figs 17-19. Lower Pliocene. Hônohasi,Saigo-mura, near Kakegawa, Japan. Holotype FGKJC.200224 (fide Hanzawa et al. 1961: 210).† nitens, Dentalium J. Sowerby, 1814: 159, pl. 70,figs 1, 2. Lower Eocene. Highgate, London Clay,Engl<strong>and</strong>, United Kingdom. Fustiaria nitens(J. Sowerby, 1814) (fide Newton 1891: 285, 286).Possible junior synonym: Dentalium incertumDeshayes, 1825 (fide Deshayes 1861: 202).Remarks: this name was incorrectly applied byEudes-Deslongchamps (1842: 129, 130) to the<strong>species</strong> subsequently described as Dentalium normianumd’Orbigny, 1850, from the Jurassic <strong>of</strong> France,<strong>and</strong> by Dixon (1850) to the <strong>species</strong> subsequentlydescribed as Dentalium lucidum Deshayes, 1861(fide Pilsbry & Sharp 1898: 208).nitense, see nicense.nitentior, Cadulus Arnold, 1903: 187, pl. 8, fig. 13 [exCarpenter MS]. <strong>Recent</strong>. California coast;Pleistocene <strong>of</strong> San Pedro, San Diego; Pliocene <strong>of</strong>San Diego well. Lectotype CAS (designated byPalmer 1958: 119); paralectotypes CAS (San Pedro,California), USNM 23729. Junior synonym <strong>of</strong>Gadila aberrans (Whiteaves, 1887) (fide Shimek1989: 235). Remarks: Pilsbry (1904: 108) expresseddoubts as to the scaphopod nature <strong>of</strong> nitentior:“The figure <strong>and</strong> description given fix the namenitentior on what seems to be the tube <strong>of</strong> a serpulidannelid”. Shimek (1989: 235) designated USNM23729 as lectotype <strong>of</strong> C. aberrans; this is invalidbecause it is not a syntype <strong>of</strong> that <strong>species</strong>. Palmer(1958: 119) stated that the figured specimen (whichArnold said was “now in the collection <strong>of</strong> DelosArnold”) was the holotype <strong>of</strong> this <strong>species</strong>. SinceArnold reported this <strong>species</strong> from several localities,Palmer’s action constitutes a lectotype designation;the CAS type specimens are not labeled as to whichspecimen is the figured specimen, so it remains tobe determined which specimen, if any, is the lectotype.However, Palmer (1958: 119) erroneouslystated that USNM 23729 (30 fms, Catalina Isl<strong>and</strong>)“has no status in a type category” since this specimenis from a locality included by Arnold in hisdescription, it was a syntype <strong>and</strong> is now a paralectotype.† nitidum, Dentalium Deshayes, 1861: 203, pl. 1,figs 29, 30. Eocene. Paris Basin (Châlons-sur-Vesles, Aizy, Mercin, Laversine, <strong>and</strong> Cuise-la-Motte),France. Remark: a valid <strong>species</strong> <strong>of</strong> Laevidentalium(fide Le Renard 1995: 175; Le Renard & Pacaud1995: 85).nitidus, Cadulus (Platyschides) Henderson, 1920: 129,pl. 19, fig. 9. <strong>Recent</strong>. Mayaguez Harbor, PuertoRico, USBF stn 6062, 25 fms [46 m]. HolotypeUSNM 887474; paratypes USNM 161582,AMNH 148353 (3). Polyschides nitidus (Henderson,1920) n. comb. Known from the type localityonly.nivosum, Dentalium Kuroda & Kikuchi, 1933: 7, 8,pl. 1, figs 9-11, text-figs 1, 2. <strong>Recent</strong>. Toyama Bay,Japan, 80-150 m. Type material not located. Juniorsynonym <strong>of</strong> Entalinopsis intercostata (Boissevain,1906) (fide Habe 1963: 272).† nobile, Dentalium Mayer, 1863: 97, 98. Tertiary.Hoering, Germany [“Couches tongriennes deHoering”]. Remark: type specimen(s) stated to bein the “Musée de mines de Munich [Germany]”.nodosus, see nodulosus.† nodulosus, Dentalites Schlotheim, 1820: 94, 95.Cretaceous. Insel Moen, Denmark. Dentalitesnodosus [Krueger, 1823: 312] is an incorrect subsequentspelling.noe, Dentalium Sismonda, 1847: 24 [ex Bonelli MS].Nomen nudum. Pliocene. Piedmont, Italy.† noe, Dentalium sexangulum var. Sacco, 1897: 94,pl. 7, figs 57-61 [as a nomen nudum in Sacco 1896:97]. Pliocene. Italy (various localities). Syntypes MIGTBS.106.01.010, BS.106.01.011, BS.106.01.012(fide Ferrero Mortara et al. 1984: 301).† noetlingi, Dentalium Martin, 1915: 180. Miocene.Singu <strong>and</strong> Kama, Burma. Remark: Martin (1915)ZOOSYSTEMA • 2004 • 26 (4)621


Steiner G. & Kabat A. R.rejected Noetling’s claim that his Burma specimenswere conspecific with Dentalium junghuhnidescribed from Java, Indonesia. Noetling (1901:250) claimed that his Burma specimens <strong>of</strong>“junghuhni” were “absolutely identical with”Dentalium magnificum E. A. Smith, 1896, a <strong>Recent</strong><strong>species</strong> from the Indian Ocean. Martin’s subsequentconclusion that the Burma specimens were notreferable to Dentalium junghuhni means thatDentalium magnificum is no longer a junior synonym,<strong>and</strong> indeed may now be a senior synonym <strong>of</strong>Dentalium noetlingi.† normanianum, Dentalium d’Orbigny, 1850 (volume2): 46. Jurassic. Villerville, France. Remark:this <strong>species</strong> was misidentified as Dentalium nitens byEudes-Deslongchamp (1842).† notabile, Dentalium Eichwald, 1857: 583. Silurian[“Grauwackenkalksteine”]. Pulkowa, Russia.Remark: figured by Eichwald, 1860: 1061, pl. 40,fig. 9; 1868: 800, 801, pl. 28, fig. 1.novaeholl<strong>and</strong>iae, Dentalium Chenu, 1843: pl. 6,fig. 14; 1852: 5. <strong>Recent</strong>. Australia. Type materialnot located. Graptacme? novaeholl<strong>and</strong>iae (Chenu,1843) (fide Pilsbry & Sharp 1897: 93). No informationon geographic or bathymetric distributionavailable.† novaki, Dentalium Koenen, 1892: 978-980, pl. 59,figs 7, 8. Oligocene. Morigny, France. Remark:Merklin & Goncharova (1967: 118) listed novaki asa junior synonym <strong>of</strong> acutum Hébert, but Janssen(1978b: 199) treated it as a valid <strong>species</strong> <strong>of</strong> Fissidentalium.† novaresei, Dentalium (Plagioglypta) de Stefani, 1917:42, 43, pl. 4, fig. 6. Carboniferous. Elba, Italy.† novemcinctum, Dentalium “?” Sacco, 1897: 96, 97,pl. 7, fig. 97 [as a nomen nudum in Sacco 1896: 97].Oligocene. Sassello, Italy. Holotype MIGTBS.106.01.045 (fide Ferrero Mortara et al. 1984:303, pl. 56, fig. 1). Remark: Bonci et al. (2000:206) concluded that this was a valid <strong>species</strong>, referableto Dentalium (Dentalium).novemcostatum, Dentalium Lamarck, 1818: 344.<strong>Recent</strong>. Around La Rochelle, France [Atlantic].Possible syntypes MHNG 1081/54 (5; fide Y.Finet, in litt. 16.X.1996). Antalis novemcostata(Lamarck, 1818) (fide Pilsbry & Sharp 1897: 51).Eastern Atlantic: Europe. 20-300 m.noviluna, see novilunatus.novilunatus, Cadulus (Platyschides) Kira, 1959: 104,pl. 40, fig. 2 [nomen nudum in Habe 1953: 298,<strong>and</strong> in Kira 1954: 80, pl. 40, fig. 2; see Bieler &Petit 1990: 141]. <strong>Recent</strong>. Shikoku, Japan. Typematerial not located. Junior synonym <strong>of</strong> Gadila virginalis(Boissevain, 1906) (fide Habe 1963: 279).Remarks: although Habe (1963: 279, 1964: 49,1977: 342) <strong>and</strong> Habe & Kosuge (1964: 12) includedG. novilunata in the synonymy <strong>of</strong> virginalis,Scarabino (1995: 358) stated that he “... prefer[s] tokeep them separate pending further study”. Gadilanoviluna [Habe 1977: 342] is an incorrect subsequentspelling.† novum, Dentalium Chenu, 1843: pl. 6, fig. 23;1850: 5. Type locality not specified (Tertiary, fidePilsbry & Sharp 1898: 210).† nudum, Dentalium Zekeli, 1852: 118, pl. 24, figs 1,2. Jurassic. Edelbachgraben, Gosautal, Austria.numatai, Dentalium (Fustiaria) Hirase, 1931: 139,pl. 3, fig. 11. <strong>Recent</strong>. Osumi <strong>and</strong> Oshima [Japan]. Typematerial not located. Junior synonym <strong>of</strong> Fustiarianipponica (Yokoyama, 1922) (fide Habe 1963: 267).† numerosa, Plagioglypta Yoo, 1988: 248, figs 148-151.Early Carboniferous, Late Tournaisian, DangarfieldFormation. Locality A-3, 2.6 km SW <strong>of</strong> Gundy,Upper Hunter, New South Wales, Australia. HolotypeAMS F.61996; paratypes AMS F.61997 (20).numerosum, Dentalium Dall, 1895: 686. <strong>Recent</strong>. Fromthe Galapagos to California. Syntypes LACM 1755(near Isla Cedros, Pacific coast Baja California,México, 28°12’N, 115°09’W, 80 m), SBMNH35016, USNM 87559, USNM 203102, USNM600413 (all from Baja California Norte, near IslaCedros, Albatross stn 2823, 24°18’N, 110°22’W,26 fms [48 m]). Junior synonym <strong>of</strong> Dentalium oerstediiMörch, 1861 (fide Pilsbry & Sharp 1897: 25).† nunomae, Dentalium Takeda, 1953: 62, 63, pl. 4,fig. 8, pl. 5, fig. 12. Oligocene, Poronai Formation.Hokkaido University Locality T268, southernSakhalin Isl<strong>and</strong>, Russia. Holotype HokkaidoUniversity No. 11090; paratype HokkaidoUniversity No. 11089 (Locality T211).† nutans, Cadulus Böhm, 1891: 70, pl. 4, fig. 26.Cretaceous. Höpfling, near Siegsdorf, southernBavaria, Germany.nutans, Dentalium Kner, 1850: 23, 24, pl. 4, figs 10,10a. Cretaceous. Nagórzany, Ukraine. Junior synonym<strong>of</strong> Dentalium decussatum J. Sowerby, 1814(fide Pilsbry & Sharp 1898: 221).nysti, Dentalium d’Orbigny, 1852 (volume 3): 18.Tertiary, Falunian. Klein-Spauwen, Hoesselt, Lethen,Belgium. Junior synonym <strong>of</strong> Dentalium acutumHébert, 1849 (fide Pilsbry & Sharp 1898: 197).Remark: this <strong>species</strong> was misidentified as Dentaliumgr<strong>and</strong>e Deshayes, 1825 by Nyst (1843).nysti, Dentalium Van den Binkhorst, 1861: 61, pl. 6,fig. 2a-2c. Cretaceous. Limbourg, Belgium. Juniorhomonym <strong>of</strong> Dentalium nysti d’Orbigny, 1852.Replacement name: Dentalium binkhorsti Pilsbry &Sharp, 1898.obesus, Cadulus Watson, 1879: 527, 528 [Watson1886: 22, pl. 3, fig. 8]. <strong>Recent</strong>. Culebra Isl<strong>and</strong>[Lesser Antilles], Challenger stn 24, 18°38’30”N,65°05’30”W, 390 fms [712 m]. Syntypes BMNH1887.2.9.84-87 (labelled “in advanced stage <strong>of</strong>decay; Palmer 1971”). Cadulus obesus Watson,1879. Western Atlantic, Caribbean. 365-712 m.† obliquatus, Cadulus Koenen, 1892: 990, pl. 59, figs19, 20. Lower Oligocene. Lattorf, Calbe, Atzendorf,<strong>and</strong> Unseburg, Germany.622ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)† oblongum, Dentalium Richardson, 1906: 585,pl. 45, fig. 3. Jurassic, Pliensbachian. Railwaycutting,Dixton West, near Gotherington, nearCheltenham, Engl<strong>and</strong>, United Kingdom. Remark:Engeser & Riedel (1992: 38) stated that “This<strong>species</strong> cannot be attributed to any known scaphopodgenus <strong>and</strong> therefore a new genus should beproposed”.obnutus, see obrutus.† obrutus, Gadus Conrad, 1870a: 227. Emendationfor Gadus “obnutus” Conrad, 1869b. Cretaceous.Haddonfield, New Jersey, USA. Syntype ANSP19502. Cadulus obrutus (Conrad, 1870) (fidePilsbry & Sharp 1898: 238).obscurum, Dentalium gouldii var. Dall, 1889: 425,pl. 27, fig. 4. <strong>Recent</strong>. Off Barbados, Lesser Antilles,Blake stn 299, 13°05’N, 59°39’40”W, 140 fms[256 m]. Lectotype USNM 95351 (designated byHenderson 1920: 32); paralectotypes USNM887460, MCZ 7695, MCZ 7708 (erroneouslystated to be the holotype by Turner 1955: 319).Dentalium obscurum Dall, 1889 (fide Henderson1920: 32). Western Atlantic: Caribbean <strong>and</strong> Brazil.146-1540 m.† obsoletum, Dentalium Hall, 1858: 724, pl. 29,figs 16, 17. Carboniferous. Iowa, USA. SyntypeFM UC.10722 (fide Forney & Nitecki 1976: 11).Objective junior synonyms: Dentalium sublaeveHall, 1877; Prodentalium fredericae Engeser &Riedel, 1992. Junior synonym: Dentalium acutisulcatumGurley, 1883 (fide Yancey 1978: 307).Prodentalium obsoletum (Hall, 1858) (fide Engeser& Riedel 1992: 41). Remarks: this is a valid name<strong>and</strong> not a junior secondary homonym <strong>of</strong> Dentalitesobsoletum Schlotheim, 1832, which is a nomennudum. The replacement <strong>names</strong>, Dentalium sublaeveHall in Miller, 1877 (being itself a juniorhomonym <strong>of</strong> Dentalium sublaeve Cocconi, 1874)<strong>and</strong> Prodentalium fredericae Engeser & Riedel, 1992are, therefore, unnecessary.obsoletum, Dentalium Doderlein, 1864: 97 [15].Nomen nudum. Miocene. Monte Gibio (Modena),Santo Agata (Tortona) <strong>and</strong> Vigoleno (Piacenza),Italy. Junior homonym <strong>of</strong> Dentalium obsoletumHall, 1858. See Dentalium dentalis var. sublaevisCocconi, 1874.obsoletus, Dentalites Schlotheim, 1813: 100; 1832: 67.Tonna, Sachsenberg, Germany. Nomen nudum.obtusum, Dentalium Qi & Ma, 1989: 114, fig. 3.<strong>Recent</strong>. Zhejiang Province, China. Type in IOAS.Dentalium obtusum Qi & Ma, 1989. Indo-Pacific:China Sea. 0-5 m.occidentale, Dentalium Stimpson, 1851b: 28. <strong>Recent</strong>.Deep water on the coast <strong>of</strong> Maine, <strong>and</strong> in MassachusettsBay [Western Atlantic]. Type material presumablylost (fide Johnson 1976: 66). Juniorsynonyms: Dentalium abyssorum M. Sars, 1859 (fidePilsbry & Sharp 1897: 47); Dentalium cinerascensAnton, 1838 (fide Pilsbry & Sharp 1897: 48, as tentativesenior(!) synonym); Dentalium aenigmaticumJordan, 1895 (fide Friele & Grieg 1901: 49).Antalis occidentalis (Stimpson, 1851) (fide Pilsbry& Sharp 1897: 47). North Atlantic. 36-1823 m.Remarks: Emerson in Turgeon (1998: 54) listedthis <strong>species</strong> in the synonymy <strong>of</strong> Antalis entalis(Linnaeus, 1758); further study <strong>of</strong> the taxonomy <strong>of</strong>these <strong>species</strong> is needed. This <strong>species</strong> was misidentifiedas Dentalium dentale by Gould (1841).occidentale, Siphonodentalium (Pulsellum) Henderson,1920: 93, pl. 16, fig. 4. <strong>Recent</strong>. Off Martha’sVineyard [Massachusetts], USBF stn 871,40°05’39”N, 70°23’52”W, 86 fms [157 m]. HolotypeUSNM 38702; paratype MCZ 186823 (2).Pulsellum occidentale (Henderson, 1920). WesternAtlantic: Massachusetts, USA. 157-1794 m.occiduus, Cadulus Verco, 1911b: 218, pl. 26, fig. 7.<strong>Recent</strong>. Geographe Bay, <strong>of</strong>f Bunbury, WestAustralia, 15 fms [27 m]. Holotype SAM D 13759.Cadulus occiduus Verco, 1911. Indo-Pacific: westernAustralia. 27-2900 m. Remark: the 7 paratypescould not be located.octangulatum, Dentalium Donovan, 1803: pl. 162.<strong>Recent</strong>. Japan. Neotype BMNH 1952.2.23.1 (designatedby Ludbrook 1954: 97). Junior synonyms:Dentalium octogonum Lamarck, 1818; Dentaliumstriatulum Turton, 1819 (fide Pilsbry & Sharp1897: 17); Dentalium yokohamense Watson, 1879;Dentalium japonicum Dunker, 1877 (fide Habe1964: 8). Dentalium octangulatum Donovan,1803. Indo-Pacific: Red Sea to Japan <strong>and</strong> NewGuinea. 5-1380 m. Remarks: the original typelocality was “near Lelant, Cornwall, Engl<strong>and</strong>”.However, the type material was mislocalized, <strong>and</strong>Ludbrook’s designation <strong>of</strong> a neotype from Japanfixes the type locality as Japan (ICZN 1999: article76.3).† octevillense, Dentalium Cossmann, 1899b: 329,pl. 2, fig. 8. Cretaceous, Cenomanian. Octeville,Seine-Inférieure [Seine-Maritime], France.octocostatum, Dentalium Fraas, 1867. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.Remark: Blanckenhorn (1927: 123, 124, pl. 1,figs 2-8) considered this to be a valid <strong>species</strong> <strong>of</strong>Dentalium, but he did not discuss the contraryconclusion <strong>of</strong> Pilsbry & Sharp (1898: 245).octocostatum, Dentalium Ihering, 1897: 266, 267,pl. 4, fig. 16. Tertiary, Santa Cruz Formation.Yegua Quemada, Patagonia, Argentina. Juniorhomonym <strong>of</strong> Dentalium octocostatum Fraas, 1867[Polychaeta]. Replacement name: Dentalium octocostellatumPilsbry & Sharp, 1898.† octocostellatum, Dentalium Pilsbry & Sharp, 1898:211. Tertiary, Santa Cruz Formation. YeguaQuemada, Patagonia, Argentina. Replacementname for Dentalium octocostatum Ihering, 1897 nonFraas, 1867 (Polychaeta). Junior synonym:Dentalium gallardoi Rovereto, 1921 (fide Feruglio1949b: 203, footnote).ZOOSYSTEMA • 2004 • 26 (4)623


Steiner G. & Kabat A. R.† octogonalis, Dentalium (Antale “?”) taurocostatumvar. Sacco, 1897: 102, pl. 8, fig. 57 [as a nomennudum in Sacco 1896: 97]. Miocene. Turin hills,Italy. Syntype MIGT BS.106.02.049 (fide FerreroMortara et al. 1984: 305).octogonum, Dentalium Lamarck, 1818: 344. <strong>Recent</strong>.Sea <strong>of</strong> China. Syntypes MNHN (2; fide Scarabino1995: 200), possible syntypes MHNG 1081/53(4; fide Y. Finet, in litt. 16.X.1996). Junior synonym<strong>of</strong> Dentalium octangulatum Donovan, 1803(fide Pilsbry & Sharp 1897: 17).octogonum, Dentalium Rayneval, Hecke & Ponzi1854: 13, 19. Pliocene. Monte Mario, Rome, Italy.Junior homonym <strong>of</strong> Dentalium octogonum Lamarck,1818. Remark: the very short general description<strong>and</strong> the lack <strong>of</strong> illustrations <strong>and</strong> types renders this adubious <strong>species</strong> for which we prefer not to introducea replacement name.octopleuron, Dentalium Verco, 1911a: 206. <strong>Recent</strong>.St Vincent Gulf, South Australia, 15-22 fms [27-40 m]. Holotype SAM D 13725. Paradentalium octopleuron(Verco, 1911) (fide Cotton & Godfrey1933: 139). Pacific Ocean: South Australia. 12-1210 m.Remark: this <strong>species</strong> was described from a lot <strong>of</strong>88 specimens; the 87 paratypes could not be located.octostriatum, Dentalium antverpiense “forma”Goddeeris, 1978: 70, 72. Miocene. Belgium.Nomen nudum. Remark: this is also an unavailableinfrasubspecific name (ICZN 1999: articles 45.5<strong>and</strong> 45.6.3).oerstedii, Dentalium Mörch, 1861: 177. <strong>Recent</strong>. Gulf<strong>of</strong> Nicoya, west coast <strong>of</strong> Costa Rica, 30 fms [55 m].Syntypes ZMUC (3; fide Keen 1966: 18, fig. 21).Junior synonym: Dentalium numerosum Dall, 1895(fide Pilsbry & Sharp 1897: 25). Dentalium oerstediiMörch, 1861. Eastern Pacific: California toEcuador, Galapagos Isl<strong>and</strong>s. 4-1480 m.okudai, Siphonodentalium Habe, 1953: 299, figs 759,760. <strong>Recent</strong>. Akkeshi Bay, Hokkaido, Japan, 20 m.Holotype NSMT 54867; paratypes NSMT 54868,AMNH 148420 (1). Siphonodentalium okudaiHabe, 1953. Known from the type locality only.† oleacinum, Dentalium agile var. Dall, 1892: 441.Pliocene. Caloosahatchie River <strong>and</strong> Shell Creek,Florida, USA. Syntypes USNM Paleobiology112739 (3).† oligocenica, Gadila Vergneau-Saubade, 1968: 208,pl. 1, fig. 23. Oligocene, Aquitanian. Gaas,Sarcignan, France.olivi, Dentalium Scacchi, 1835: 12, pl. 2, fig. 6.Pleistocene. Gravina, Italy. Holotype UPMN (fideR. La Perna pers. comm.). Tentative junior synonym:Siphodentalium hyalinum Brugnone, 1876(fide Pilsbry & Sharp 1898: 171), non Dentaliumhyalinum Philippi, 1846. Polyschides olivi (Scacchi,1835) (fide Steiner & Dreyer 2003: 346, 349).Eastern Atlantic, Mediterranean. 500-2242 m.Remark: Jeffreys (1877: 157, 158) listed this <strong>species</strong>in the genus Gadila.onyx, Hamulus Morton, 1834. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.† oolithicum, Dentalium Piette, 1856: 598, pl. 15,figs 28, 29. Jurassic. Rumigny, France. Junior synonym:Dentalium entaloides Eudes-Deslongchamps,1842 non Fleming, 1825 (fide Pilsbry & Sharp1898: 225).† opaculum, Dentalium Casey, 1903: 266, 267.Oligocene. Vicksburg, Mississippi, USA.opacum, Dentalium Sowerby, 1829: 198, 199. <strong>Recent</strong>.“From South Sea ships, supposedly from NewZeal<strong>and</strong>” (G. Humphrey’s collection, Pilsbry &Sharp 1897: 70). Type material not located. Fissidentaliumopacum (Sowerby, 1829) (fide Pilsbry &Sharp 1897: 70). New Zeal<strong>and</strong>. Remarks: Finlay(1928: 262) considered this <strong>species</strong> indeterminate.Dell (1957: 564) treated it as nomen dubium due tothe absence <strong>of</strong> the type material <strong>and</strong> pointed out apossible synonymy with Fissidentalium zel<strong>and</strong>icum(Sowerby, 1860).† opalina, Dentalium filicauda var. Quenstedt, 1856:328, pl. 44, fig. 16. Jurassic, Liassic. Near Berlin,Germany. Remark: Richardson (1906: 575, 576)tentatively suggested that this name might be ajunior synonym <strong>of</strong> Dentalium elongatum Goldfuss,1841.† operculatus, Dentalites Krueger, 1823: 312 [exBohadsch MS]. Near Gosathal, Hallstadt,Germany.ophiodon, Dentalium Dall, 1881: 38. <strong>Recent</strong>. OffBahia Honda, Cuba, Blake stn 20, 23°02’30”N,83°11’W, 220 fms [402 m]. Lectotype USNM95344 (designated by Henderson 1920: 85); paralectotypesUSNM 95345 (1; Blake stn 21), MCZ7727 (1; Blake stn 19), MCZ 7728 (1; Blakestn 20), MCZ 7729 (1; Blake stn 21; erroneouslystated to be the holotype by Turner 1955: 319).Compressidens ophiodon (Dall, 1881) (fide Pilsbry& Sharp 1897: 126). Western Atlantic, Caribbean.128-1280 m.opiparium, Quasidentalium Shimansky, 1974. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.opportuna, Gadila Kuroda & Habe in Habe, 1961:105, pl. 47, fig. 2 [nomen nudum in Habe 1953:298]. <strong>Recent</strong>. Off Zushi City, Sagami Bay,Honshu, Japan, 100-200 m. Holotype in NSMT.Gadila opportuna Kuroda & Habe in Habe, 1961.Indo-Pacific: Japan. 100-780 m.† ornatum, Dentalium de Koninck, 1843: 318, 319,pl. 22, figs 3a-3c. Carboniferous. Visé, Belgium.† ornatus, Cadulus (Dischides) Baluk, 1972: 563, 564,pl. 5, figs 9-13. Miocene, Lower Tortonian(Badenian). Korytnica, 24 km SSW <strong>of</strong> Kielce, HolyCross Mountains, Pol<strong>and</strong>. Holotype <strong>and</strong> paratypes,Baluk collection.orsum, Dentalium Bonelli in Sismonda, 1842: 25.Tertiary. Piedmont, Italy. Junior synonym <strong>of</strong>Dentalium inaequale Bronn, 1831 (fide Sismonda624ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)1847: 24; Pilsbry & Sharp 1898: 205), itself a juniorsynonym <strong>of</strong> Dentalium sexangulum Gmelin,1791 (fide Bernasconi 1996: 26).† orthezense, Dentalium Tournouër, 1873: 466 [40],pl. 6, fig. 12. Eocene. Orthez, Adour Basin, southwesternFrance.† orthoceras, Dentalium Pilsbry & Sharp, 1898: 225.Carboniferous. Visé, Belgium. Replacement namefor Entalis filosa de Koninck, 1883, a junior secondaryhomonym <strong>of</strong> Dentalium filosum Broderip &Sowerby, 1830.orthrum, Dentalium entalis var. Watson, 1879: 512,513. <strong>Recent</strong>. Type locality not specified. SyntypesBMNH 1887.2.9.18-20 (Setubal, Challenger stn II,470 fms [858 m]; Fayal, Azores, Challenger stn 75,450 fms [821 m]; Prince Edward Isl<strong>and</strong>, Challengerstn 145, 150 fms [274 m]). Synonym <strong>of</strong> Antalisentalis (Linnaeus, 1758).† ortus, Fissidentalium Palmer, 1974b: 124. Miocene.Locality R, Java, Indonesia. Syntypes RGM 7272(5) (fide Van den Hoek Ostende et al. 2002: 112,pl. 3). Replacement name for Dentalium tenuistriatumMartin, 1880 non Rouault, 1850.oryx, Dentalium Boissevain, 1906: 20, pl. 6, fig. 24, textfig.14. <strong>Recent</strong>. Sunda Isl<strong>and</strong>s, Siboga stn 302,10°27.9’S, 123°28.7’E, 216 m. Lectotype ZMA3.06.014 (designated by Scarabino 1995: 205); paralectotypesZMA 3.06.015 (4). Junior synonym:Dentalium paucicontortum Boissevain, 1906 (fideScarabino 1995: 205, by First Reviser action, accordingto ICZN 1999: article 24). Dentalium oryx Boissevain,1906. Indo-Pacific: Africa to Philippines. 216-965 m.osceola, Spirodentalium Walcott, 1890. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† otamaringaensis, Dentalium Marwick, 1926: 326,327, pl. 74, fig. 15. Miocene. Mouth <strong>of</strong> WaiauStream E to Otamaringa Pa, Waitara SurveyDistrict, New Zeal<strong>and</strong>. Holotype NZGS TM4808;paratypes NZGS (7) (fide Keyes 1972: 96).Dentalium otamaringaense Marwick, 1926 (fide Beu& Maxwell 1990: 424).† otatumei, Dentalium Nagao, 1938: 141, 142, pl. 15[2], figs 11-11b. Cretaceous. Ponporokabetu, nearYubari coal mines, Isikari province, Japan.Holotype Dept. Geol., Hokkaido Univ., No. 8235(fide Hanzawa et al. 1961: 209).† ottoi, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1897: 119; 1898: 211. Oligocene.Joachimsthal <strong>and</strong> Hermsdorf, Germany. Replacementname for Dentalium compressum Meyer, 1883non d’Orbigny, 1850.† ouachitensis, Cadulus (Cadulus) Palmer, 1937: 21,pl. 2, figs 18, 19. Eocene, Lower ClaiborneFormation. PRI Locality 741, Lapiniere L<strong>and</strong>ing,east bank <strong>of</strong> Ouachita River, Louisiana <strong>and</strong>Columbus, Louisiana, USA. Syntypes PRI 2644,PRI 2645.ovale, Dentalium “Malm.” – Pilsbry & Sharp 1898:225. Nomen nudum? Remarks: we have not foundthe original description <strong>of</strong> this <strong>species</strong>; Pilsbry &Sharp (1898: 225) wrote: “Description not seen byus. Jurassic, Dogger: Germany”, <strong>and</strong> R. Janssen (inlitt.) stated that the putative senior homonymappears to be a manuscript name that was neverdescribed.† ovale, Dentalium Cooke, 1928: 9, pl. 2, fig. 5.Oligocene. Alazan Clay, Rio Buena Vista, west <strong>of</strong>Alazan, Vera Cruz, Mexico. Type material USNMPaleobiology 352708.ovalis, Cadulus (Dischides) Boissevain, 1906: 66, textfig.31, pl. 6, fig. 52. <strong>Recent</strong>. Madura Bay <strong>and</strong> otherlocalities in the southern part <strong>of</strong> Molo-strait[Indonesia], Siboga stn 51, 69-91 m. SyntypesZMA 3.06.088-089 (2). Dischides ovalis (Boissevain,1906). Indo-Pacific: Africa to Indonesia. 69-101 m.† ovatum, Dentalium Hu, 1992: 659, figs 9, 14-17,22. Pleistocene. Limestone <strong>of</strong> Henchun Peninsula,Taiwan. Holotype NMT 1639d; paratypes NMT1639, 1639a-c. Remark: this <strong>species</strong> probablybelongs to the genus Compressidentalium.† ovosectum, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 226. Upper Cretaceous. Tisa [Tyssa],Bohemia, Czech Republic. Replacement name forDentalium glabrum Geinitz, 1839 non Montagu,1803.ovulum, Dentalium Philippi, 1844: 208, 209, pl. 27,fig. 21. Pleistocene near Crotone, Calabria, Italy.Type material not located. Cadulus ovulum(Philippi, 1844) (fide Philippi 1844, who also establishedthe monotypic genus Cadulus for this type<strong>species</strong>). Eastern Atlantic, Mediterranean. 982-2643 m. Remark: this may be an <strong>fossil</strong> <strong>species</strong>, as DiGeronimo & La Perna (1997: 417, 418) claimedthat reports <strong>of</strong> <strong>Recent</strong> specimens in theMediterranean <strong>and</strong> in the Atlantic are based onreworked <strong>fossil</strong>s or misidentified specimens <strong>of</strong>Cadulus gibbus Jeffreys, 1883.† ozawai, Siphonodentalium (Pulsellum) Yokoyama,1926: 289, pl. 34, fig. 11. Pliocene, SawanéFormation. Sado Isl<strong>and</strong>, Japan. Holotype GIUT(fide Hanzawa et al. 1961: 210).pachypleurum, Dentalium (Gadilina) Boissevain,1906: 51, text-fig. 26, pl. 5, figs 21, 22. <strong>Recent</strong>.Locality <strong>of</strong> syntypes: ZMA 3.06.063 (2; B<strong>and</strong>a Sea,Siboga stn 208, 5°39’S, 122°12’E, 1886 m), ZMA3.06.064 (1; Arafura Sea, Siboga stn 271, 5°46.7’S,134°0’E, 1788 m). Gadilina pachypleura(Boissevain, 1906) (fide Scarabino 1995: 285).Indo-Pacific: Indonesia. 1788-1886 m.pacifica, Costentalina Chistikov, 1982b: 1311, 1314,pl. 1, figs 3, 4, pl. 2, figs 10, 14. <strong>Recent</strong>. AleutianTrench, Vitjaz stn 6136 T, 53°25’N, 163°23’W,4180-4186 m. Holotype IOM; paratypes IOM (1from stn 5159, 7°36’N, 162°01’W, 4916 m; 3 fromstn 6097, 57°00’N, 148°18.5’W, 4740 m; 1 from6107-1, 57°37.8’N, 143°12’W, 3800 m). Costentalinapacifica Chistikov, 1982. Indo-Pacific:ZOOSYSTEMA • 2004 • 26 (4)625


Steiner G. & Kabat A. R.Micronesia; Tasman Sea; Eastern Pacific: Alaska.3253-4916 m.pacificum, Dentalium Hutton, 1873a: 5, 6. <strong>Recent</strong>.New Zeal<strong>and</strong>. Holotype NMNZ M263 (fideMarshall 1996: 43); paratype ZMB 24023 (fideKilias 1995: 175). Junior synonym <strong>of</strong> Fissidentaliumzel<strong>and</strong>icum (Sowerby, 1860) (fide Suter1913: 819).† pagellai, Dentalium Negri, 1934: 172, 173, pl. 14,figs 15-17. Paleogene. Gef Gef el-Abiad (Zella-Marada) <strong>and</strong> Haifàt (Zella-Marada), Libya.pallida, Pseudantalis rubescens var. Monterosato, 1884:33. Nomen nudum without type locality.† palmerae, Cadulus (Gadila) Hodgkinson, 1974: 17,text-fig. 8t, u; pl. 1, figs 15, 16, pl. 2, fig. 8. MiddleEocene, Claibornian Stage, Cook MountainFormation, Wheelock Member. Stone City Bluff,Brazos River Valley, Burleson County, Texas, USA.Holotype USNM Paleobiology 180439; paratypesUSNM Paleobiology 180440, 180449, PRI 29225,ANSP 31503.† palmerae, Dentalium (Antalis) Garvie, 1996: 46,pl. 9, fig. 2. Eocene, Lower Claibornian, ReklawFormation, Marquez Member. Locality 4, Bluff onRidge Creek, Texas, USA. Holotype PRI 30313.panamense, Dentalium megathyris var. Dall, 1908: 358.<strong>Recent</strong>. Gulf <strong>of</strong> Panama. Syntypes LACM 1836(Isla Malpelo, Colombia, 3241 m), USNM 122975(16; Albatross stn 3381, 4°56’N, 80°52’30”W, 1772fms [3234 m]), USNM 122977 (7) <strong>and</strong> 594263 (1)(Albatross stn 3361, 6°10’N, 83°06’W, 1471 fms[2685 m]), USNM 122982 (1; Albatross stn 3360,6°17’N, 82°05’W, 1672 fms [3051 m]), SBMNH35019 (1; Albatross stn 3361, 6°10’N, 83°06’W,1471 fms [2685 m]). Synonym <strong>of</strong> Fissidentaliummegathyris (Dall, 1889).panamensis, Cadulus (Gadila) Sharp & Pilsbry inPilsbry & Sharp, 1898: 191, 192, pl. 36, figs 23-25.<strong>Recent</strong>. Panama Bay, 51 fms [93 m]. Holotype (<strong>of</strong>Dentalium corrugatum) BMNH 1857.6.4.881 (fideKeen 1968: 403, text-fig. 24). Replacement namefor Dentalium corrugatum Carpenter, 1857 non corrugatumHupé in Gay, 1854; junior synonym <strong>of</strong>Gadila perpusilla (Sowerby in Broderip &Sowerby, 1832) (fide Emerson 1971: 78). Remark:Emerson (1971: 78) erroneously selected a “lectotype”for Cadulus panamensis Pilsbry & Sharp,1898; this action is incorrect under ICZN (1999)article 72.7, since panamensis was a replacementname for corrugatum <strong>and</strong> thus both taxa have thesame type material. For that reason, the “paralectotypes”<strong>of</strong> panamensis (e.g., Scott et al. 1990: 18;Boyko & Sage 1996: 30, 31) have no nomenclaturalstatus.p<strong>and</strong>ionis, Cadulus Verrill & Smith in Verrill, 1880:392. <strong>Recent</strong>. Off Martha’s Vineyard [Massachusetts],USBF stn 876, 120 fms [219 m].Lectotype USNM 38644 (designated by Johnson1989: 56 as holotype); paralectotypes MCZ 186811(3; USFC stn 877, 126 fms [230 m]), MCZ186812 (1; USFC 873, 100 fms [183 m]), MCZ186813 (2; USFC 871, 115 fms [210 m]), MCZ186814 (USFC stn 870, 155 fms [283 m]). Gadilap<strong>and</strong>ionis (Verrill & Smith in Verrill, 1880) (fidePilsbry & Sharp 1898: 171). Western Atlantic:New Jersey to Florida (USA). 46-704 m. Remark:Emerson in Turgeon (1998: 55) placed this <strong>species</strong>in the genus Polyschides.panormitanum, Dentalium “Chenu” – Jeffreys 1883:657. Unjustified emendation <strong>of</strong> Dentalium panormumChenu. Antalis panorma (Chenu, 1843) (fidePilsbry & Sharp 1897: 54).panormum, Dentalium Chenu, 1843: pl. 6, fig. 13;1852: 6. <strong>Recent</strong>. “Fossil”, Sicily [Italy]. Type materialnot located. Junior synonyms: Dentalium pseudoentalisO. G. Costa, 1851; Dentalium arguticostaBrugnone, 1878 (fide Pilsbry & Sharp 1897: 54,Bellini 1909: 226). Antalis panorma (Chenu, 1843)(fide Pilsbry & Sharp 1897: 54). Eastern Atlantic,Mediterranean: Europe <strong>and</strong> North Africa. 54-350 m.† parallelum, Dentalium Zinndorf, 1928: 38, pl. 1,fig. 8 [ex Boettger MS]. Oligocene (Rupelton).Offenbach, Germany. Rhabdus parallelus (Zinndorf,1928) (fide Janssen 1978b: 140, 141).† parannulatum, Dentalium n. nom. Eocene.Claiborne, Alabama, USA. Syntype USNMPaleobiology 638812. Replacement name forDentalium annulatum Meyer, 1886 non Dentaliumannulatum Gmelin, 1791.† pardelensis, Dentalium Brunet, 1995: 49, pl. 5,fig. 2. Upper Miocene, Entrerriense Formation.Punta Pardela, Valdés Peninsula, Chubut Province,Argentina. Holotype MPEF-PI-075.† pareorensis, Dentalium Pilsbry & Sharp, 1898: 211.Miocene, Pareora Formation. Kanieri; ChathamIsl<strong>and</strong>s; Motanau, New Zeal<strong>and</strong>. Replacementname for Dentalium laevis Hutton, 1873 nonSchlotheim, 1820. Lectotype NZGS TM4812 (designatedby Keyes 1972: 97). Antalis pareorensis(Pilsbry & Sharp, 1898) (fide Beu & Maxwell 1990:424).parfaiti, Dentalium caudani var. Locard, 1898: 104 [exFischer MS]. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Syntype MNHN. Synonym<strong>of</strong> Dentalium caudani, a junior synonym <strong>of</strong>Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877).† parianus, Cadulus Guppy in Guppy & Dall, 1896:325, pl. 30, fig. 7. Oligocene. Pointapier, Trinidad.Syntypes USNM Paleobiology 107155 (12).Remark: Schuchert et al. (1905: 112) cited USNM107155 as the “holotype” but did not specify whichsyntype was to be the holotype, so this does notconstitute a valid lectotype designation.† parisiensis, Dentalium d’Orbigny, 1850 (volume 2):372. Eocene. Chaumont, Parnes, Mouchy-le-Châtel, Senlis (Oise), France. Replacement namefor Dentalium semistriatum Deshayes, 1825 non626ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Turton, 1819. Dentalium parisiense d’Orbigny,1850 (fide Deshayes 1864: 210, pl. 2, figs 17-19;Pilsbry & Sharp 1898: 211). Remark: a valid <strong>species</strong><strong>of</strong> Dentalium (Entaliopsis) (fide Le Renard 1995:175; Le Renard & Pacaud 1995: 85).parisiensis, Gadus Deshayes, 1861: 218, 219, pl. 3,figs 18-20. Eocene. Paris Basin (numerous localities),France. Junior synonym <strong>of</strong> Polyschides denticulatum(Deshayes, 1850) (fide Pilsbry & Sharp 1898:238). Remark: Pilsbry & Sharp (1898: 238) erroneouslylisted Dentalium denticulatum Deshayes,“1864” as a junior synonym <strong>of</strong> this <strong>species</strong>; in fact,D. denticulatum was described in 1850, not 1864,<strong>and</strong> the synonymy is herein reversed. Used as a valid<strong>species</strong> <strong>of</strong> Siphonodentalium by Le Renard (1995:175) <strong>and</strong> Le Renard & Pacaud (1995: 86), whowere unaware that it was a junior synonym.† parkinsoni, Dentalium Quenstedt, 1852: 443,pl. 35, fig. 18; 1867: 531. Jurassic. Germany.Remarks: Brauns (1865: 137; 1869: 191) statedthat this <strong>species</strong> was a junior synonym <strong>of</strong> Dentaliumentaloides Eudes-Deslongchamps, 1842 (= Dentaliumoolithicum Piette, 1856). However, that <strong>species</strong>name is a junior homonym <strong>of</strong> Dentalium entaloidesFleming, 1825. Pilsbry & Sharp (1898: 226)instead treated Quenstedt’s <strong>species</strong> as valid.† parvula, Fustiaria Stoliczka, 1868: 445, pl. 27,fig. 22. Cretaceous, Valudayur <strong>group</strong>. Pondicherry,India.† parvulina, Cadulus tumidosus var. Sacco, 1897: 116,pl. 10, figs 74-76 [as a nomen nudum in Sacco 1896:98]. Miocene. San Agata, Piedmont, Italy. SyntypesMIGT BS.106.10.010-BS.106.10.012 (fide FerreroMortara et al. 1984: 310, pl. 55, figs 13, 14).Cadulus (Cadulus) parvulinus Sacco, 1897 (fideMoroni & Ruggeri 1980: 47, figs 1, 2). Remark:Moroni & Ruggeri (1980: 47, by First Reviseraction) listed Cadulus tumidosus var. perinflataSacco, 1897 as a junior synonym <strong>of</strong> this <strong>species</strong>.† parvulum, Dentalium Philippi, 1887a: 107, 108,pl. 12, fig. 16; 1887b: 102, pl. 12, fig. 16. Tertiary.Navidad <strong>and</strong> Lebu, Chile. Objective junior synonym:Dentalium philippianum Pilsbry & Sharp,1898. Remark: Pilsbry & Sharp (1898: 212)considered this <strong>species</strong> as congeneric with Fustiariaparvula Stoliczka, 1868 <strong>and</strong>, therefore, introducedthe replacement name. Although the two <strong>species</strong>belong to Dentalium <strong>and</strong> Fustiaria respectively,Philippi’s name cannot be restored according toICZN (1999).parvulum, Dentalium Richardson, 1906: 585-587,pl. 45, figs 9, 12 [ex J. Buckman MS]. Jurassic,Pliensbachian. Leckhampton Station clay-pit,Cheltenham, Engl<strong>and</strong>, United Kingdom. Juniorhomonym <strong>of</strong> Dentalium parvulum Philippi, 1887.Replacement name: Dentalium subparvulumEmerson, 1954.† parvum, Dentalium Mayer, 1864: 358, 359, pl. 14,fig. 6. Miocene. Menou, near Léognan, France.parvus, Cadulus (Platyschides) Henderson, 1920: 113,114, pl. 18, figs 2, 4. <strong>Recent</strong>. Off Barbados, Blake,100 fms [183 m]. Holotype USNM 314713;paratype MCZ 48414 (1). Cadulus parvusHenderson, 1920. Western Atlantic: Caribbean,Brazil. 60-219 m. Remarks: Henderson (1920: 113,114) recognized that the original description <strong>of</strong>Cadulus amiantus Dall, 1889 encompassed two<strong>species</strong>; therefore, he described parvus for thosespecimens <strong>of</strong> “amiantus” that were not conspecificwith amiantus s.s. Emerson in Turgeon (1998: 55)placed this <strong>species</strong> in the genus Polyschides, althoughthe position <strong>of</strong> the maximum diameter in the middle<strong>of</strong> the shell suggests it to belong to Cadulus.passeriniamum, see passerinianum.† passerinianum, Dentalium Cocconi, 1874: 646, 647,pl. 6, figs 18, 19. Miocene. San Agata, Italy.Remarks: Dentalium striatissimum Doderlein, 1862(nomen nudum) listed in synonymy. Dentaliumpasseriniamum [Sacco, 1897: 96] is an incorrectsubsequent spelling.† patagonicum, Dischides Brunet, 1995: 55, pl. 5,fig. 12. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-082.† patriciae, Laevidentalium Engeser & Riedel, 1992:41. Middle Devonian. Grube Rothe Erde, nearGiessen, Germany. Replacement name for Dentaliumrobustum Maurer, 1885 non Brazier, 1877.† patulum, Dentalium Potiez & Michaud, 1838: 545.Cretaceous, Green S<strong>and</strong>. Varenne, Meuse, northeasternFrance.paucicontortum, Dentalium Boissevain, 1906: 28, 29,pl. 6, figs 25-28. <strong>Recent</strong>. Sulu Archipelago [Philippines],Siboga stn 95, 05°43.5’N, 119°40’E,522 m. Lectotype ZMA 3.06.022 (designated byScarabino 1995: 205); paralectotypes ZMA3.06.023 (34; fide R. Moolenbeek & A. N. Van derBijl, in litt. 23.VI.2000), ZMA 3.06.024 (1;stn 105, 6°08’N, 121°19’E, 275 m). Junior synonym<strong>of</strong> Dentalium oryx Boissevain, 1906 (fideScarabino 1995: 205, by First Reviser action,according to ICZN 1999: article 24).† paucicostata, Dentalium (Entalis) badensis var. Sacco,1897: 108, pl. 9, figs 24-28 [as a nomen nudum inSacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.03.013 (fide FerreroMortara et al. 1984: 307).paucicostatum, Dentalium capillosum var. Watson,1879: 509. <strong>Recent</strong>. Setubal [Portugal], Challengerstn II, 38°10’N, 9°14’W, 470 fms [858 m].Holotype BMNH 1887.2.9.1. Junior synonym:Dentalium exuberans Locard, 1897 (B. Métivier,n. syn.). Fissidentalium paucicostatum (Watson,1879) (fide Pilsbry & Sharp 1897: 78). EasternAtlantic: Europe to northern Africa. 403-3650 m.† paucicostulata, Dentalium (Antale “?”) dentale var.Sacco, 1897: 105, pl. 8, fig. 78 [as a nomen nudumin Sacco 1896: 97]. Pliocene. Italy (various localities).ZOOSYSTEMA • 2004 • 26 (4)627


Steiner G. & Kabat A. R.Syntype MIGT BS.106.02.069 (fide FerreroMortara et al. 1984: 306).† paucistriata, Pulsellum (Entalina) tetragona var.Sacco, 1897: 114, pl. 10, figs 54, 55 [as a nomennudum in Sacco 1896: 97]. Miocene/Pliocene. Italy(various localities). Syntypes MIGT BS.106.08.008, BS.106.08.009 (fide Ferrero Mortara et al.1984: 310).† paulini, Dentalium Maury, 1925: 403, 404, pl. 24,fig. 1. Cretaceous. Estacao Agronomica (CamadaNo. 7), Pará, Brasil.† pauperculum, Dentalium Meek & Hayden, 1860:178. Cretaceous. Lance Formation, Fox HillsS<strong>and</strong>stone, Moreau River, South Dakota, USA.Holotype USNM Paleobiology 311 (figured byStanton 1920: 34, pl. 6, fig. 6).peitaihoensis, Dentalium King & Ping, 1935: 103-105, text-figs 1, 2. <strong>Recent</strong>. Peitaiho, beach[Beidaihe], China. Type material <strong>and</strong> type depositorynot specified. Dentalium? peitaihoensis King& Ping, 1935. Indo-Pacific: China. Known fromthe type locality only. Remarks: generic placementuncertain. Figures 1a <strong>and</strong> 1b resemble fragments orjuveniles <strong>of</strong> Omniglypta cerina (Pilsbry, 1905) orEpisiphon subtorquatum (Fischer, 1871); figure 1cdepicts a Caecum shell. Moreover, the presence <strong>of</strong> athick orange to brownish periostracum suggests thatsome <strong>of</strong> the types do not belong to the <strong>Scaphopoda</strong>.The authors erroneously classified their new <strong>species</strong>in the “Order Scutibranchiata” which is actually anordinal-level name in the Gastropoda.pelamidae, Polyschides Chistikov, 1979a: 113, 114,fig. 7. <strong>Recent</strong>. Tonkin, Vietnam, Pelamide stn 46,56 m. Holotype ZIN. Polyschides pelamidaeChistikov, 1979. Indo-Pacific: Indonesia to China.31-150 m. Polyschides pelamide [Scarabino 1995:338] is an incorrect subsequent spelling.pelamide, see pelamidae.† pellati, Dentalium Loriol in Loriol & Pellat, 1874:405, pl. 10, figs 36, 37. Upper Jurassic, Portl<strong>and</strong>ian.Ningle, Alpreck, Boulogne-sur-Mer,northern France. Remark: Laevidentalium pellati(Loriol in Loriol & Pellat, 1874), fide Gerasimov(1969: 52).pelliceri, Dentalium (Episiphon) sowerbyi Henderson,1920: 80, 81, pl. 13, figs 7-9. <strong>Recent</strong>. BahiaHonda, Cuba, Tomas Barrera stn 208, 1-12 fms [2-22 m]. Holotype USNM 314565; paratypesUSNM 314566-314573, ANSP 119157, AMNH148322 (35; stn 203, Cabanas Harbor, Cuba, 3-12 fms [5-22 m]), AMNH 148323 (6). Synonym<strong>of</strong> Episiphon sowerbyi (Guilding, 1834).† pellucens, Dentalium Deshayes, 1861: 214, 215,pl. 1, figs 21-23. Eocene. Paris Basin (Damery,Montmirel, Parnes <strong>and</strong> Grignon), France. Fustiariapellucens (Deshayes, 1861) (fide Le Renard 1995:175; Le Renard & Pacaud 1995: 85).pellucidum, Dentalium Gmelin, 1791. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† pentagonale, Dentalium Pilsbry & Sharp, 1898: 226.Triassic. Bavaria, Germany. Replacement name forDentalium quinquangulare Gümbel, 1861 nonForbes, 1844.pentagonum, Siphonodentalium M. Sars, 1865a: 307-312, pl. 7, figs 45-51. <strong>Recent</strong>. Drøbak, near Oslo;Langesund <strong>and</strong> Flekkefjord, 50-120 fms [91-219 m]; L<strong>of</strong>oten Isl<strong>and</strong>s, 50-300 fms [91-548 m][Norway]. Type material not located. Junior synonym<strong>of</strong> Entalina tetragona (Brocchi, 1814) (fideM. Sars 1870: 195, as S. quinquangulare). Remark:in his description, Sars referred to the specimenmisidentified as Dentalium abyssorum juv. (M. Sars1865b: 43, figs 107-109).† peracuta, Dentalium sexangulum var. Sacco, 1897:94, pl. 7, fig. 63 [as a nomen nudum in Sacco 1896:97]. Pliocene. Carrú, Italy. Syntype MIGTBS.106.01.013 (fide Ferrero Mortara et al. 1984:301).† peranulatus, Cadulus Guppy, 1912: 4. Tertiary.Corosal, Trinidad.† perarmatum, Dentalium Ryckholt, 1851: 67, pl. 2,figs 39, 40. Carboniferous. Visé, Belgium. Pilsbry& Sharp (1898: 232) questioned the placement <strong>of</strong>this <strong>species</strong> in the <strong>Scaphopoda</strong>, because the posteriorspines are unknown in this class.perceptum, Dentalium Mabille & Rochebrune inRochebrune & Mabille, 1889: 99-100. <strong>Recent</strong>.Type locality not specified. Syntypes MNHN (2;labelled “Beagle Channel” [Patagonia]). Rhabdusperceptus (Mabille & Rochebrune in Rochebrune& Mabille, 1889) (fide Pilsbry & Sharp 1897: 115).Western Atlantic: Argentina; Eastern Pacific: Chile.278-933 m.† perfragile, Dentalium (Pulsellum) Koenen, 1892:986, 987, pl. 59, figs 17, 18. Lower Oligocene.Lattorf, Westeregeln <strong>and</strong> Helmstadt, Germany.† perigonicum, Dentalium Doderlein, 1864: 97 [15].Miocene. Monte Gibio, Modena, Italy.perinflata, Cadulus tumidosus var. Sacco, 1897: 116,pl. 10, fig. 77 [as a nomen nudum in Sacco 1896:98]. Miocene. San Agata, Piedmont, Italy. SyntypeMIGT BS.106.10.013 (fide Ferrero Mortara et al.1984: 310). Junior synonym <strong>of</strong> Cadulus (Cadulus)parvulinus Sacco, 1897 (fide Moroni & Ruggeri1980: 47, figs 1, 2, as First Reviser action).† perinii, Dentalium Malaroda, 1951: 190, 191, pl. 5,fig. 23, pl. 7, fig. 19. Eocene. Monteccio, Costozza,Colli Berici, near Vicenza, Italy. Holotype Museodell’Istituto Geologia dell’Università di Padova.perinvolutum, Dentalium (Fissidentalium) Ludbrook,1954: 101, 102, fig. 7. <strong>Recent</strong>. Gulf <strong>of</strong> Aden, JohnMurray Exp. stn 185, 13°48’N, 49°16’E, 2000 m.Holotype BMNH 1952.3.25.65. Antalis perinvoluta(Ludbrook, 1954) (fide Scarabino 1995: 233).Indo-Pacific: Red Sea to Indonesia <strong>and</strong> Australia.918-2000 m.† perlaevis, Dentalium (Antale) bouei var. Sacco, 1897:99, pl. 8, fig. 18 [as a nomen nudum in Sacco 1896:628ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)97]. Miocene. Turin hills, Italy. Lectotype MIGTBS.106.02.012 (designated by Bonci et al. 2000:228, 229); paralectotypes MIGT BS.106.02.005-BS.106.02.007, MIGT BS.106.02.007/01 (34),MIGT BS.106.02.007/02 (36), MIGT BS.106.02.014/01 (14), MIGT BS.106.02.012/01 (25) (fideBonci et al. 2000: 228).perlongum, Dentalium Dall, 1881: 36. <strong>Recent</strong>. North<strong>of</strong> Yucatan Bank, Blake stn 33, 24°01’N, 88°58’W,1568 fms [2868 m]. Lectotype MCZ 7752 (designatedby Turner 1955: 319 as “holotype”); paralectotypesMCZ 7660 (1), MCZ 7661 (1; Blakestn 41), MCZ 7663 (1; Blake stn 46), MCZ 7664(3; Yucatan Strait, 640 fms [1168 m]). Graptacmeperlonga (Dall, 1881) (fide Scarabino 1994: 307).Western Atlantic, Caribbean: Cape Hatteras (USA)to Brazil. 200-4850 m. Remark: Henderson (1920:caption to pl. 9, fig. 1) stated “selected type,99 mm”, however, none <strong>of</strong> the then-syntype specimensis over 80 mm in length, hence Henderson’sstatement does not constitute a valid lectotypeselection.† perplexa, Gadila Boettger, 1907: 211, 212.Miocene. Valea Semini <strong>and</strong> Valea Casilor, Kostej,Banat, Romania. Lectotype SMF XIII.5a (designatedby Zilch 1934: 278, pl. 22, fig. 23).perpusillum, Dentalium Sowerby in Broderip &Sowerby, 1832: 29. <strong>Recent</strong>. Puerto Solango,Colombia [Eastern Pacific]. Lectotype BMNH1950.10.10.1-5 (best preserved specimen, lenght =7.2 mm, <strong>of</strong> 5 shells glued to a wooden block) (designatedby Emerson 1971) <strong>and</strong> 4 paralectotypes.Junior synonyms: Dentalium corrugatum Carpenter,1857 (fide Pilsbry & Sharp 1898: 192); Cadulus(Gadila) panamensis Sharp & Pilsbry in Pilsbry &Sharp, 1898 (fide Emerson 1971: 78). Gadila perpusilla(Sowerby in Broderip & Sowerby, 1832)(fide Pilsbry & Sharp 1898: 190). Eastern Pacific:Baja California to Panama, Galapagos Isl<strong>and</strong>s,?Ecuador. 18-120 m.† perrilliatae, Tesseracme n. nom. Middle Miocene.Stns 9995 <strong>and</strong> 23737, Santa Rosa, Veracruz,Mexico. Holotype USNM Paleobiology 350085;paratypes UNAM 2520. Replacement name forDentalium (Tesseracme) mexicanum Perrilliat, 1975non Dentalium mexicanum Girty, 1909.† perseveratum, Dentalium Brunet, 1995: 49, 50,pl. 5, fig. 4. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-077.† perstriolata, Dentalium (Antale) vulgare var. Sacco,1897: 98, pl. 8, figs 1-5 [as a nomen nudum in Sacco1896: 97]. Pliocene. Bordighera, Astigiana <strong>and</strong>Vezza d’Alba, Italy. Syntypes MIGT BS.106.02.001, BS.106.02.002 (fide Ferrero Mortara et al.1984: 303).pertracheatum, Dentalium (Plagioglypta) Plate,1908a: 357, 358, pl. 30. figs 45, 46. <strong>Recent</strong>. NiasNorth Channel (15 sm [= nautical miles] South <strong>of</strong>Bangkam), Valdivia stn 203, 1°47.1’N, 96°58.7’E,660 m. Lectotype ZMB 61099a (designated byKilias 1995: 175); paralectotypes ZMB 61099b (1),ZMB 63839 (1; between Cocos Isl<strong>and</strong> <strong>and</strong>Sumatra, Valdivia stn 185, 3°41.3’S, 100°59.5’E,614 m). Plagioglypta pertracheata (Plate, 1908)(fide Scarabino 1995: 236). Indo-Pacific:Madagascar to New Caledonia. 460-1740 m.peruvianum, Dentalium Dall, 1908: 358. <strong>Recent</strong>. OffPeru, Albatross stn 4656, 6°55’S, 83°34’W,2222 fms [4055 m]. Holotype USNM 110667;paratype USNM 110666 (Albatross stn 4649, fideKabat 1996: 17). Fissidentalium peruvianum (Dall,1908) n. comb. (V. Scarabino in litt. 6.III.2000).Eastern Pacific: Peru. 40-4085 m.peruvianus, Cadulus (Gadila) Dall, 1908: 361. <strong>Recent</strong>.Off Point Aguja, Peru, Albatross stn 4654, 1036 fms[1885 m]. Lectotype USNM 110671 (designatedby Keen 1971: 965 as “holotype” in caption to figure21); paralectotypes USNM 122806 (Albatrossstn 2807, fide Kabat 1996: 17), USNM 602250;MCZ 27972. Gadila peruviana (Dall, 1908).Eastern Pacific: Peru. 1484-1894 m.† petricola, Dentalium Dall, 1909: 136. Miocene.Astoria <strong>and</strong> Coos Bay, Oregon; Clallam Bay,Washington, USA. Holotype USNM Paleobiology3481. Rhabdus petricola (Dall, 1909) (fide Emerson1958: 93). Remark: because Dall considered Dentaliumsubstriatum (Conrad, 1849) to encompasstwo <strong>species</strong>, he proposed this name for the smoothform, <strong>and</strong> Dentalium conradi Dall [= Dentaliumpseudonyma Pilsbry & Sharp, 1898] for the striatedform; see Emerson (1958) <strong>and</strong> Moore (1963: 50,51) for further discussion.† peyreirense, Dentalium (Antale) Cossmann &Peyrot, 1917: 166, 167, pl. 1, figs 32, 33. Neogene,Helvetian <strong>and</strong> Tortonian. Peyrère, Saint-Étienned’Orthe<strong>and</strong> Saint-Jean-de-Marsacq, France.phaneum, Dentalium Dall, 1895: 686, pl. 26, fig. 1.<strong>Recent</strong>. Hawaii, Albatross stn 3476, 21°09’N,157°53’E, 298 fms [545 m]; Albatross stn 3475,21°08’N, 157°43’E, 351 fms [641 m]. SyntypesUSNM 107025 (3; stn 3476), USNM 107026 (11;stn 3475). Antalis phanea (Dall, 1895) (fideScarabino 1995: 233). Indo-Pacific: New Caledoniato French Polynesia <strong>and</strong> Hawaii. 545-860 m.Remark: V. Scarabino (in litt. 5.XI.2001) statedthat the type material comprises two <strong>species</strong>, thesecond potentially new.† phenax, Cadulus Pilsbry & Sharp, 1898a: 472, 473,pl. 11, figs 23, 24. Oligocene. Domincan Republic.Type material ANSP 2883 (fide Pilsbry 1922: 400).philcolmani, Tesseracme Lamprell & Healy, 1998: 77,figs 76B, 77C, D, 79, 81A-D. <strong>Recent</strong>. 13 km NE <strong>of</strong>Redcliff Point, Bernier Isl<strong>and</strong>, Australia, 24°54’S,113°17.75’E, 22.5 m. Holotype WAM 233/94;paratypes WAM 118-93 (3), AMS C173310 (1;18.5 km NNE <strong>of</strong> Port Hedl<strong>and</strong>, Western Australia,19°30.31’S, 118°52.49’E, 40 m), AMS C174539ZOOSYSTEMA • 2004 • 26 (4)629


Steiner G. & Kabat A. R.(1; 50 miles NNE <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 19°29.2-5’S, 118°52.5-3’E , 40-41 m),AMS C174543 (1; N <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 19°29.9-4’S, 118°52.0-3’E, 37 m), AMSC174548 (2; N <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 19°30.5-31.0’S, 118°49.5’E, 39-41 m),AMS C174547 (1; N <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 19°30.5-31.0’S, 118°49.5’E, 39-41 m),AMS C174645 (2; N <strong>of</strong> Port Hedl<strong>and</strong>, 19°29.6-9’S, 118°51.7-0’E, 40-41 m). Tesseracme philcolmaniLamprell & Healy, 1998. Indo-Pacific:Western Australia. 40-124 m.† philippi, Dentalium Chenu, 1843: pl. 6, fig. 17;1850: 6. Miocene or Pliocene. Italy.philippi, Dentalium Monterosato, 1872: 27. Tertiary.Sicily, Italy. Junior homonym <strong>of</strong> Dentalium philippiChenu, 1843. Replacement name: Dentalium rectumvar. monterosatoi Pilsbry & Sharp, 1898.Remark: this <strong>species</strong> was misidentified as Dentaliumstriatum Lamarck, 1818 by Philippi (1844).philippii, Dentalium Cossmann, 1907c: 202. Tertiary.Navidad, Chile. Unnecessary replacement name forDentalium gracile Philippi, 1887 non Moore, 1866non Hall & Meek, 1855. Junior objective synonym<strong>of</strong> Dentalium navidadense Pilsbry & Sharp, 1898.Junior homonym <strong>of</strong> Dentalium philippi Chenu,1843 <strong>and</strong> Monterosato, 1872.philippianum, Dentalium Pilsbry & Sharp, 1898: 212.Tertiary. Navidad <strong>and</strong> Lebu, Chile. Objective juniorsynonym <strong>of</strong> Dentalium parvulum Philippi,1887. Remark: Pilsbry & Sharp (1898: 212) consideredDentalium parvulum Philippi, 1887 as congenericwith Fustiaria parvula Stoliczka, 1868 <strong>and</strong>,therefore, introduced D. philippianum as a replacementname. Although the two <strong>species</strong> belong toDentalium <strong>and</strong> Fustiaria respectively, Philippi’sname cannot be restored according to ICZN (1999).philippinarum, Dentalium Sowerby, 1860: 98, 99,pl. 225, fig. 54. <strong>Recent</strong>. Isl<strong>and</strong> <strong>of</strong> Samar,Philippines. Type material not located. Junior synonym<strong>of</strong> Laevidentalium eburneum (Linnaeus,1767) (fide Scarabino 1995: 291).† phoenicea, Cadulus Gardner, 1933: 199, 200, pl. 20,fig. 2. Eocene, Midway Group, Kincaid Formation.USGS Station 5282, clay from bluff at Webberville,Travis County, Texas, USA. Holotype USNMPaleobiology 370996.pichoni, Episiphon Lamprell & Healy, 1998: 134, figs135E, 136G, 140. <strong>Recent</strong>. Off Cairns, northernQueensl<strong>and</strong>, Australia, 17°22’S, 146°48’E, 303 m.Holotype QM MO40064; paratypes QMMO40068 (44), QM MO40065 (32), QMMO40066 (180), AMS C172490 (1; SE <strong>of</strong> SwainsReefs, Queensl<strong>and</strong>, 22°26.27-20.2’S, 153°17.13-17.6’E, 187 m), AMS C172479 (1; c. 160 km N <strong>of</strong>Croker Isl<strong>and</strong>, Northern Territory, 9°30’S,132°34’E). Episiphon pichoni Lamprell & Healy,1998. Indo-Pacific: northern <strong>and</strong> eastern Australia.18-351 m. Remark: V. Scarabino (in litt. 5.XI.2001)suggested that the status <strong>of</strong> E. pichoni Lamprell &Healy, 1998 vis-à-vis Gadilina insolita (E.A. Smith,1894) requires further research.picteti, Dentalium (Dentalium) Pilsbry & Sharp, 1897:22, 23, pl. 11, fig. 87 [ex Deshayes MS]. <strong>Recent</strong>.West Indies “?”. Syntype MHNG 1155/15. Juniorsynonym <strong>of</strong> Paradentalium americanum (Chenu,1843) (fide Henderson 1920: 28; fide Scarabino1975: 183).† pictile, Dentalium (Laevidentalium) Tate, 1899:263, pl. 8, fig. 8. Oligocene. Table Cape, Tasmania,Australia. Holotype <strong>and</strong> paratype SAM T 1608.Laevidentalium pictile Tate, 1899 (fide Ludbrook1959: 146, pl. 2, fig. 4).† pilsbryi, Cadulus (Platyschides) Woodring, 1925:209, 210, pl. 28, fig. 10. Miocene. Bowden,Jamaica. Holotype USNM Paleobiology 352726.pilsbryi, Dentalium (Antalis) Rehder, 1942: 69.<strong>Recent</strong>. Brazil. Holotype (<strong>of</strong> pseudohexagonumHenderson) USNM 225817. Replacement namefor Dentalium pseudohexagonum Henderson, 1920non Arnold, 1903. Antalis pilsbryi (Rehder, 1942).Western Atlantic, Caribbean: Florida to Brazil.0-15 m.pistis, Dentalium (Graptacme) Winckworth, 1940: 43,fig. 6. <strong>Recent</strong>. Madras [Chennai, southern India].Holotype BMNH 1940.7.22.7; paratypes(?) MAM.Paradentalium pistis (Winckworth, 1940) (fideScarabino 1995: 222). Indian Ocean: India. Nobathymetric information available.† planatum, Dentalium Bronn, 1831: 84. Miocene.Castell’Arquato, between Parma <strong>and</strong> Piacenza,Italy.† planicostata, Dentalium (Entalis) badensis var. Sacco,1897: 108, pl. 9, figs 29, 30 [as a nomen nudum inSacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.03.014 (fide FerreroMortara et al. 1984: 307).† planicostatum, Dentalium Hébert, 1855: 374,pl. 29, fig. 11. Cretaceous. Meudon, France.planum, Dentalium J. Sowerby, 1814. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.platamodes, Siphodentalium Watson, 1879: 519, 520[Watson 1886: 13, pl. 2, fig. 4]. <strong>Recent</strong>. WestIndies: Culebra Isl<strong>and</strong>, Challenger stn 24,18°38’30”N, 65°05’30”W, 390 fms [712 m].Syntypes BMNH 1887.2.9.61-62 (3). Junior synonym:Entalina quadrata Henderson, 1920 (fideEmerson 1952a: 7, 8; Scarabino 1975: 181).Entalina platamodes (Watson, 1879) (fide Pilsbry& Sharp 1897: 133). Western Atlantic, Caribbean:Florida to Brazil. 200-1400 m.platei, Cadulus (Gadila) Jaeckel, 1932: 311, fig. 9.<strong>Recent</strong>. Cape Agulhas, South Africa, Valdivia stn109, 35°19’S, 20°12’E, 126 m. Holotype ZMB75376. Cadulus platei Jaeckel, 1932. Known fromthe type locality only.platei, Dentalium (Dentalium) Jaeckel, 1932: 303,304, text-figs 1, 2. <strong>Recent</strong>. Southwest Africa,630ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Valdivia stn 104, 35°16’S, 22°26,7’E, 155 m.Holotype ZMB 75386; paratypes ZMB 78431,ZMB 75387 (1; South Africa, Valdivia stn 84,16°26.5’S, 11°41.5’E, no depth given; fide Kilias1995: 176). Junior synonym <strong>of</strong> Fissidentaliumexasperatum (Sowerby, 1903) (fide Barnard 1963a:347).platensis, Cadulus (Cadulus) Henderson, 1920: 147,148, pl. 20, fig. 1. <strong>Recent</strong>. Off Rio de la Plata,USBF stn 2764, 36°42’S, 56°23’W, 11.5 fms[21 m; an obvious error as the coordinates indicatea depth <strong>of</strong> around 2000 m; V. Scarabino pers.comm.]. Holotype USNM 887476; paratypesUSNM 95447, USNM 108172, USNM 108279,USNM 330845, AMNH 148340 (1; <strong>of</strong>fFern<strong>and</strong>ina, Florida, USBF 2668, 294 fms [537 m];fide Boyko & Sage 1996: 31). Cadulus platensisHenderson, 1920. Western Atlantic: Georgia(USA) to Brazil. 538-1857 m.platyceras, Dentalium (Compressidens) Sharp & Pilsbryin Pilsbry & Sharp, 1897: 126, 127, pl. 22, figs 58-60. <strong>Recent</strong>. Port Stephens, Salam<strong>and</strong>er Bay, NewSouth Wales, Australia, 7-15 m [type locality pursuantto ICZN 1999: article 76.2]. LectotypeANSP 35565 (designated by Lamprell & Healy,1998: 147); paralectotypes ANSP 35565 (17), AMSC55085 (7; Salam<strong>and</strong>er Bay, Port Stephens, NewSouth Wales, 24 m), AMS C11721 (7; Salam<strong>and</strong>erBay, Port Stephens, New South Wales, 15-20 m).Compressidens platyceras (Sharp & Pilsbry inPilsbry & Sharp, 1897) (fide Scarabino 1995: 370).Pacific Ocean: southeastern Australia. 7-2900 m.platypleurum, Dentalium (Fissidentalium) Tomlin,1931: 339, 340. <strong>Recent</strong>. Off Itongazi River, Natal[South Africa], 25 fms [46 m]. Holotype SAFMA3631. Fissidentalium platypleurum (Tomlin,1931). Known from the type locality only.platystoma, Cadulus (Gadila) Pilsbry & Sharp, 1898:180, pl. 35, figs 17, 18. <strong>Recent</strong>. Off Manta,Ecuador, USFC stn 2792, 0°37’S, 81°00’W,401 fms [733 m]. Holotype USNM 107699.Gadila platystoma (Pilsbry & Sharp, 1898). Knownfrom the type locality only.† playagr<strong>and</strong>ensis, Cadulus (Gadila) Weisbord, 1964:131, 132, pl. 18, figs 17, 18. Pliocene, Playa Gr<strong>and</strong>eFormation. Stn W-24, Quebrada Las Pailas,Venezuela. Holotype PRI 26898.† pleiocenum, Dentalium Tuomey & Holmes, 1856:105, 106, pl. 25, fig. 2. Pliocene. Pee Dee, SouthCarolina, USA.† pliocenica, Dentalium (Entalis) badensis var. Sacco,1897: 107, 108, pl. 9, fig. 21 [as a nomen nudum inSacco 1896: 97]. Pliocene. Vezza d’Alba, Italy.Syntype MIGT BS.106.03.011 (fide FerreroMortara et al. 1984: 307).† pliocenica, Dentalium (Entalis) recta var. Sacco,1897: 110, pl. 10, fig. 3. Pliocene. Piacentino <strong>and</strong>Rio Torsero, Liguria, Italy. Syntype MIGTBS.106.03.034 (fide Ferrero Mortara et al. 1984:308). Remark: if both <strong>of</strong> Sacco’s (1897) “pliocenica”taxa prove valid <strong>species</strong> <strong>of</strong> Fissidentalium (as suggestedby the current placement <strong>of</strong> the <strong>species</strong> theywere described as varieties <strong>of</strong>) this one will have tobe renamed as a junior homonym.† pliocenica, Dentalium (Entalina) mediocarinatasubsp. Koperberg, 1931: 33. Pliocene. Locality 6,“Nono Fatoe Fekoe (Toi Oesapi Soka),” Timor.Junior homonym <strong>of</strong> Dentalium (Entalis) badensisvar. pliocenica Sacco, 1897 <strong>and</strong> D. (Entalis) rectavar. pliocenica Sacco, 1897. Remark: if both <strong>of</strong>Sacco’s (1897) “pliocenica” taxa prove to be valid<strong>species</strong>, Koperberg’s <strong>species</strong> will have to be renamedas since it is a junior homonym.pluricostatum, Dentalium Boissevain, 1906: 30, pl. 6,figs 6, 7. <strong>Recent</strong>. Indonesia, Buton Strait, Sibogastn 204, 04°20’S, 122°58’E, 75-94 m. LectotypeZMA 3.06.027 (designated by Scarabino 1995:207); paralectotypes ZMA 3.06.025 (1; BougainvilleStrait, stn 153, 0°3.8’N, 130°24.3’E,141 m), ZMA 3.06.026 (3, fide R. Moolenbeek &A. N. Van der Bijl, in litt. 23.VI.2000; probably2 specimens lost). Dentalium pluricostatumBoissevain, 1906. Indo-Pacific: East Africa toPhilippines. 9-240 m.plurifissuratum, Schizodentalium Sowerby, 1894: 158,159, pl. 12, fig. 24. <strong>Recent</strong>. Hong Kong (probablyerroneous, fide Pilsbry & Sharp 1897: 83, 84, <strong>and</strong>fide Scarabino 1995: 257). Lectotype BMNH1895.4.29.179 (designated by Sowerby 1903: 231;as “the type”); paralectotypes BMNH 1895.4.29.180, 181 (2). Schizodentalium plurifissuratumSowerby, 1894. South Africa. 80-564 m. Remark:Sowerby (1894: 159) was uncertain about the origin<strong>of</strong> the specimens. The subsequently reportedmaterial from South Africa renders Hong Kong anunlikely type locality.pluteum, Dentalium (Laevidentalium) Colman, 1958:143, 144, fig. 8. <strong>Recent</strong>. Off Woollongong, NewSouth Wales, Australia, 100 fms [183 m]. Holotype<strong>and</strong> 4 paratypes AMS C18217. Junior synonym <strong>of</strong>Laevidentalium erectum (Sowerby, 1860) (fideLamprell & Healy 1998: 109).poculum, Cadulus Dall, 1889: 429. <strong>Recent</strong>. OffSt Vincent, West Indies, Blake stn 226, 13°09’05”N,61°16’20”W, 424 fms [774 m; 646 fms in error inoriginal description]. Lectotype USNM 95374(designated by Henderson 1920: 109); paralectotypesUSNM 887461 (1), MCZ 7743 (1; Cape SanAntonio, 640 fms [1168 m]). Polyschides poculum(Dall, 1889) n. comb. Western Atlantic, Caribbean.774-1168 m. Remark: Turner (1955: 319)erroneously stated that MCZ 7743 was the“holotype”.podagrinus, Cadulus (Cadulus) Henderson, 1920: 148,149, pl. 20, fig. 5. <strong>Recent</strong>. Off English Harbor,Antigua, 120 fms [219 m]. Holotype USNM596597; paratypes USNM 314935. Junior synonym:Cadulus halius Henderson, 1920 (n. syn., byZOOSYSTEMA • 2004 • 26 (4)631


Steiner G. & Kabat A. R.First Reviser action, according to ICZN 1999: article24). Cadulus podagrinus Henderson, 1920.Known from the type locality only.politum, Dentalium Linnaeus, 1767: 1264 [citingGualtieri 1742: t. 10. f. F]. <strong>Recent</strong>(?). Indian Ocean<strong>and</strong> Sicily. Syntypes LS 614 (2), ZMUU 1012.Fustiaria polita (Linnaeus, 1767) (fide Pilsbry &Sharp 1897: 128). Indian Ocean. Remarks:Deshayes (1861: 216) considered Linnaeus’ specimenslikely to be <strong>fossil</strong>. Pilsbry & Sharp (1897:128) identified as Dentalium (Fustiaria) politum<strong>Recent</strong> specimens which Deshayes (1825: 368) <strong>and</strong>Sowerby (1860: 99) had reported as Laevidentaliumeburneum (Linnaeus, 1767).politum, Dentalium Blainville, 1819: 70, nonLinnaeus, 1767. Junior synonym <strong>of</strong> Antalis vulgaris(Da Costa, 1778) (fide Pilsbry & Sharp 1897: 42).politus, Ditrupa S. Wood, 1842: 459, pl. 5, fig. 14.Coralline Crag (Pliocene), Sutton, Engl<strong>and</strong>, UnitedKingdom. Type material not located. Junior synonym:Dentalium bifissum S. Wood, 1848 (fidePilsbry & Sharp 1897: 144). Dischides politus(S. Wood, 1842) (fide Pilsbry & Sharp 1897: 144).Eastern Atlantic, Mediterranean. 9-324 m.politus, Gadus Benoist, 1873: 268, 269. Miocene.Giraudeau, Lagus <strong>and</strong> Pont-Pourquey, France.Junior secondary homonym <strong>of</strong> Cadulus politus(S. Wood, 1842). Replacement name: Siphonodentalium(Dischides) subpolitum Cossmann &Peyrot, 1917.polycostatum, Striodentalium Qi & Ma, 1989: 119,120, fig. 12. <strong>Recent</strong>. East China Sea, 184 m. Typesin IOS(?). Striodentalium polycostatum Qi & Ma,1989. Indo-Pacific: China Sea. 184 m.† polyedrum, Dentalium Seguenza, 1879: 275.Pliocene, Astian stage. Gallina, Reggio, Calabria,Italy.† polygonum, Dentalium Reuss, 1844: 201, 202.Cretaceous. Wollenitz <strong>and</strong> Priesen a/d Eger,Bohemia, Czech Republic.polygonum, Dentalium Casey, 1903: 267. Oligocene.Red Bluff, Mississippi, USA. Doubtful syntypeUSNM Paleobiology 481673 (1) (fide MacNeil &Dockery 1984: 247). Junior homonym <strong>of</strong>Dentalium polygonum Reuss, 1844. Replacementname: Dentalium casey: n. nom. Remarks: there is ah<strong>and</strong>written label in the type lot, with no author,stating that the “specimen associated with this numberis not the specimen described 9/8/64 [8 Sept.1964]”. MacNeil & Dockery (1984: 247, pl. 67,figs 10-13) redescribed this <strong>fossil</strong> <strong>and</strong> treated it as avalid <strong>species</strong> <strong>of</strong> Dentalium.† polypleurum, Dentalium Seifert, 1959: 25, 26, pl. 1,figs 5, 6. Oligocene. Hoerstgen, Kapellen, Rossenray,Sternberger Gestein, Germany. Holotype KielGeologisches Institut No. 738; paratype KielGeologisches Institut No. 739.† poncensis, Cadulus Maury, 1920: 47, pl. 7, fig. 13.Tertiary. Maury stn 283, near Ponce, Puerto Rico.ponderi, Fissidentalium Lamprell & Healy, 1998: 83,84, figs 82A, 83A, 84. <strong>Recent</strong>. Off Newcastle, NewSouth Wales, Australia, 32°59’S, 152°33.5’E,444 m. Holotype AMS C174624; paratypes AMSC173468 (1), AMS C307826 (2; NE <strong>of</strong> Woollongong,New South Wales, 34°18-24’S, 151°26-21’E,457-484 m), SAM D 18888 (2; 50 km E <strong>of</strong> GaboIsl<strong>and</strong>, Victoria, Australia, 37°37’S, 150°17’E, 436m). Fissidentalium ponderi Lamprell & Healy,1998. Pacific Ocean, Tasman Sea: southernAustralia. 82-770 m.† ponderosum, Dentalium Gabb, 1873: 244, 245.Tertiary. Dominican Republic. Type materialANSP 2708 (fide Pilsbry 1922: 400). Tesseracmedissimilis var. ponderosum (Gabb, 1873) (fide Pilsbry& Sharp 1898a: 470, pl. 10, figs 1-3, pl. 11, figs 15,16).porcatum, Dentalium Gould, 1859: 166. <strong>Recent</strong>.Hong Kong Harbor [China]. Lectotype MCZ169304 (designated by Johnson 1964: 130 as holotype);paralectotypes USNM 24142 (2). Antalisporcata (Gould, 1859) (fide Scarabino 1995: 228).Indo-Pacific: Madagascar to Hong Kong. 75-90 m.Remark: the lectotype is missing, with a note in thetray saying “Specimen missing 8-15-77”.† porterensis, Dentalium Weaver, 1912: 79, pl. 13,fig. 113. Lower Miocene, Blakeley Formation.U.W. [University <strong>of</strong> Washington] Loc. 90, W <strong>of</strong>Porter, on bluff on north side <strong>of</strong> Chehalis River,Grays Harbor County, Washington, USA.Holotype CAS 7517 (fide Weaver 1942: 268).Remarks: the type locality was originally indicatedas from Chehalis County, <strong>and</strong> corrected by Weaver(1942) to Grays Harbor County. The holotype <strong>of</strong>this <strong>species</strong> was formerly WSM 81 (“U.W. 81”),but many <strong>of</strong> the Weaver types were subsequentlytransferred to the CAS (R. Eng in litt. 29.VI.2001).† portlocki, Dentalium Tate, 1870b: 17, pl. 1, fig. 15.Replacement name for Dentalium tenue Portlock,1843 non Goldfuss, 1841. Jurassic, Liassic.Aghanloo, Ballymaglin <strong>and</strong> Ballycarton; Magilligan,Craig <strong>and</strong> Gortmore, Irel<strong>and</strong>. Holotype Museum <strong>of</strong>Practical Geology, London, No. 5799 (fideRichardson 1906: 584) (now in BMNH?). Richardsonalso designated this specimen as the neotype <strong>of</strong>Laevidentalium minimum (Buckman in Murchison,1845). Junior objective synonym <strong>of</strong> Laevidentaliumminimum (Buckman in Murchison, 1845).Remark: Woodward (1893: 352) listed this name asa junior synonym <strong>of</strong> Dentalium etalense Terquem &Piette, 1865, but without any explanation.portoricense, Dentalium (Dentalium) gouldii Henderson,1920: 30, 31, pl. 2, fig. 5. <strong>Recent</strong>. MayaguezHarbor, Puerto Rico, USBF stn 6062, 25-30 fms[46-55 m]. Holotype USNM 161568. Sub<strong>species</strong><strong>of</strong> Dentalium gouldii Dall, 1889.portoricensis, Cadulus (Platyschides) Henderson, 1920:115, 116, pl. 18, fig. 6. <strong>Recent</strong>. Mayaguez Harbor,Puerto Rico, USBF stn 6062, 25 fms [46 m].632ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Holotype USNM 314712. Polyschides portoricensis(Henderson, 1920) n. comb. Western Atlantic,Caribbean. 46-86 m.† potiezi, Dentalium (s.s.) Le Renard, 1994: 39.Eocene. Grignon, France. For Dentalium sulcatumLamarck, 1818 non Schumacher, 1817. Dentaliumpotiezi Le Renard, 1994 (fide Le Renard 1995: 175;Le Renard & Pacaud 1995: 85).potteri, Dentalium (Dentalium) Lamprell & Healy,1998: 53, 54, figs 44E, 45E, 50A-C, 51. <strong>Recent</strong>.Off Cairns, northern Queensl<strong>and</strong>, 17°50.67’S,147°18.2’E, 703 m. Holotype QM MO17888;paratypes QM MO17732 (5), QM MO17766(1; 17°51’S, 147°18’E, north Queensl<strong>and</strong>, 703 m),AMS C201739 (1; 17°51’S, 147°18’E, northQueensl<strong>and</strong>, 703 m). Dentalium potteri Lamprell& Healy, 1998. Indo-Pacific: eastern Australia.410-703 m.† praecursor, Dentalium (Compressidens) Pilsbry &Sharp, 1898a: 472, pl. 11, figs 12-14. Oligocene.Dominican Republic. Type material ANSP 2717(fide Pilsbry 1922: 400).praecursor, Dentalium Mayer-Eymar, 1904: 312, 313[description; 1903: 283 name only]. Tertiary.Hélouan <strong>and</strong> Wadi el Fih, Egypt. Junior homonym<strong>of</strong> Dentalium praecursor Pilsbry & Sharp, 1898;junior synonym <strong>of</strong> Dentalium kahirahenseCossmann, 1901 (fide Oppenheim 1906: 214,215).† praetenuis, Cadulus Stephenson, 1952: 143, pl. 34,figs 7-9. Upper Cretaceous, Cenomanian,Woodbine Formation. Grayson County Locality154, gullies south <strong>of</strong> the old Sherman road,2.8 miles E <strong>of</strong> Whitesboro, Grayson County, Texas,USA. Holotype USNM Paleobiology 105616;paratypes USNM Paleobiology 105616-105618.pressum, Dentalium (Compressidens) Sharp & Pilsbry inPilsbry & Sharp, 1897: 124, 125, pl. 22, figs 50-52,pl. 7, fig. 11. <strong>Recent</strong>. Culebra Isl<strong>and</strong>, West Indies,Challenger stn 24, 18°38’30”N, 65°05’30”W,390 fms [712 m]. Holotype (<strong>of</strong> compressum Watson)BMNH 1887.2.9.35. Replacement name forDentalium compressum Watson, 1879 non d’Orbigny,1850. Compressidens pressum (Sharp & Pilsbryin Pilsbry & Sharp, 1897). Western Atlantic: BlakePlateau (<strong>of</strong>f the USA coast between Cape Hatteras<strong>and</strong> Florida) to Brazil. 170-1430 m.pretiosum, Dentalium Sowerby, 1860: 97, pl. 225,fig. 57 [ex Nuttall MS]. <strong>Recent</strong>. California.Syntypes BMNH 1950.11.8.4-7. Junior synonyms:Dentalium indianorum Carpenter, 1864; Antaliscolumbiana Clessin, 1896; A. denseliratum Pilsbry& Sharp, 1898 (fide Pilsbry & Sharp 1897: 45,1898: 250). Antalis pretiosa (Sowerby, 1860) (fidePilsbry & Sharp 1897: 44). Eastern Pacific: Canadato California. 2-152 m.† primarium, Dentalium Hall, 1858: 666, pl. 23,fig. 16. Carboniferous. Warsaw Group, HancockCounty, Illinois, USA.prionotum, Siphonodentalium Watson, 1879: 522, 523[Watson 1886: 16, pl. 2, fig. 9]. <strong>Recent</strong>. RaineIsl<strong>and</strong>, Cape York, Queensl<strong>and</strong>, Australia,Challenger stn 185, 11°35’S, 143°03’E, 155 fms[283 m]. Lectotype BMNH 1887.52.9.67 (designatedby Ludbrook 1954: 116); paralectotypesBMNH 1887.52.9.68 (3). Dischides prionotus(Watson, 1879) (fide Pilsbry & Sharp 1898: 146).Indo-Pacific: Red Sea, Madagascar to Australia. 8-284 m. Remark: Lamprell & Healy (1998: 167)independently selected the same lectotype.† priscum, Dentalium Goldfuss, 1841: 2, pl. 166,fig. 3 [ex Münster MS]. Carboniferous. Tournay,Belgium. Remark: as nomen nudum in S<strong>and</strong>berger(1842: 399).† prisma, Dentalium Dall, 1892: 442, pl. 15, fig. 5.Pliocene. Caloosahatchie River, Florida, USA.Syntypes USNM Paleobiology 112742 (3 + fragments).Remark: Schuchert et al. (1905: 220) citedUSNM 112742 as the “holotype” but did not specifywhich syntype was to be the holotype, so thisdoes not constitute a valid lectotype designation.† prismaticum, Dentalium Seguenza, 1879: 117,pl. 11, figs 49, 49a. Miocene. Reggio, Calabria, Italy.pr<strong>of</strong>undorum, Dentalium E. A. Smith, 1894: 167,168, pl. 4, fig. 18. <strong>Recent</strong>. Colombo, Sri Lanka,India, Investigator stn 167, 6°32’N, 79°37’E,675 fms [1234 m]. Lectotype ZSI M60/1 (fide labelin BMNH) or ZSI 4120/7 (fide list at BMNH)(designated by Lamprell & Healy 1998: 93 as holotype);paralectotypes BMNH 1894.9.11.11. Juniorsynonym: Fissidentalium lima Kuroda & Habe inHabe, 1963 (fide Habe 1964: 13). Fissidentaliumpr<strong>of</strong>undorum (E. A. Smith, 1894) (fide Pilsbry &Sharp 1897: 79). Indo-Pacific: Madagascar to Japan<strong>and</strong> New Caledonia. 40-3560 m.† proliferum, Dentalium Chenu, 1843: pl. 4, fig. 5;1850: 6. Pliocene/Miocene. Italy.promontorii, Cadulus Barnard, 1963b: 353, fig. 30h-l(nomen nudum in Barnard 1963a: 446). <strong>Recent</strong>. OffCape Point, South Africa, 700 fms [1278 m].Syntypes SAFM A7460 (4; fide Giles & Gosliner1983: 41), NMW. Siphonodentalium promontorii(Barnard, 1963) (fide Scarabino 1995: 331). SouthAfrica. 850-1280 m.propinquus, Cadulus G. O. Sars, 1878: 106, 107,pl. 20, fig. 15a, b, pl. I, fig. 5. <strong>Recent</strong>. Finmark <strong>and</strong>west coast <strong>of</strong> Norway, 100-450 fms [183-821 m].Syntypes ZMO 26071-72. Cadulus propinquusG. O. Sars, 1878. Eastern Atlantic: Europe <strong>and</strong>North Africa. 60-2300 m.† prosperus, Cadulus (Gadilopsis) Marwick, 1931: 159,pl. 18, fig. 346. Miocene, Tongaporutuan, Ihungia<strong>and</strong> Tutamoe Series. Waikohu, New Zeal<strong>and</strong>.Holotype NZGS TM4816 (fide Keyes 1972: 97).Striocadulus prosperus (Marwick, 1931) (fide Beu &Maxwell 1990: 424).† prosseri, Plagioglypta Morningstar, 1922: 245, 246,pl. 16, figs 3, 4. Carboniferous, Lower Pennsylvanian.ZOOSYSTEMA • 2004 • 26 (4)633


Steiner G. & Kabat A. R.Lower Mercer Limestone (Perry County, MuskingumCounty <strong>and</strong> Licking County); McArthurLimestone (Jackson County <strong>and</strong> Vinton County),Ohio, USA. Type material USNM Paleobiology340010 (1) (Bald Knob, 2 miles southeast <strong>of</strong>Newark, Licking County, Ohio).† proteiforme, Dentalium Cossmann, 1910: 81, 82,pl. 5, figs 26-28. Pliocene. Kariakal [Karikal], India.Three syntypes stated to be in the Bonnet collection.† proterum, Dentalium (Dentalium) armillatum subsp.Woodring, 1973: 482, pl. 71, figs 8, 17. LateOligocene, Caimito Formation. USGS locality18841, Barro Colorado Isl<strong>and</strong>, Canal Zone,Panama. Holotype USNM Paleobiology 646697.Synonym <strong>of</strong> Dentalium armillatum Toula, 1911.† protuberans, Pipadentalium Yoo, 1988: 249, 250,figs 152-154. Early Carboniferous, Late Tournaisian,Dangarfield Formation. Locality A-3,2.6 km SW <strong>of</strong> Gundy, Upper Hunter, New SouthWales, Australia. Holotype AMS F.61998; paratypeAMS F.61999 (1).providensis, Cadulus (Platyschides) Henderson, 1920:122, pl. 18, fig. 14. <strong>Recent</strong>. Off Old ProvidenceIsl<strong>and</strong>, Bahamas, USBF stn 2150, 13°34’45”N,81°21’10”W, 382 fms [696 m]. Holotype USNM94055; paratypes USNM 108176, AMNH 148352(1; USBF stn 2668, 294 fms [537 m]; fide Boyko &Sage 1996: 31). Gadila providensis (Henderson,1920). Western Atlantic, Caribbean. 536-696 m.Remark: the subgenus Platyschides was put in synonymywith Gadila by Scarabino (1995: 196, 355).† prudens, Dentalium Brunet, 1995: 49, pl. 5, fig. 3.Upper Miocene, Entrerriense Formation. BalizaPunta Flecha, Puerto Madryn, Chubut Province,Argentina. Holotype MPEF-PI-076.† pseudacutum, Fissidentalium novaki subsp. R. Janssen,1989: 81, 82, pl. 1, figs 1-12. Middle Oligocene,Rupelian. Welschberg, near Waldböckelheim,Mainzer Becken, Germany. Holotype SMF308300; paratypes SMF 308301/6, SMF308302/7, SMF 308303/25, SMF 308304/100,RGM 229723/10.† pseudaprina, Dentalium (Antale) novemcostatum var.Sacco, 1897: 102, pl. 8, figs 59, 60 [as a nomennudum in Sacco 1896: 97]. Pliocene. Italy (variouslocalities). Syntypes MIGT BS.106.02.051,BS.106.02.052 (fide Ferrero Mortara et al. 1984:305).† pseudellipticum, Laevidentalium n. nom. LowerOligocene. Lattorf, Germany. Replacement namefor Dentalium (Laevidentalium) ellipticum Koenen,1892 non Dentalium ellipticum J. Sowerby, 1814.† pseudoantalis, Dentalium Lamarck, 1818: 345 [originalspelling pseudo-antalis]. Tertiary. Grignon,France. Junior synonym: Dentalium pseudoentalisDefrance, 1819 (fide Deshayes 1825: pl. 1, figs 4-6). Remarks: a valid <strong>species</strong> <strong>of</strong> Dentalium(Entaliopsis) (fide Le Renard 1995: 175; Le Renard& Pacaud 1995: 85). Dentalium pseudoentalis[Defrance, 1819: 72] is an incorrect subsequentspelling.† pseudobouei, Dentalium (Entalis) badensis var. Sacco,1897: 108, pl. 9, fig. 23 [as a nomen nudum inSacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.03.012 (fide FerreroMortara et al. 1984: 307).† pseudocircinata, Fustiaria Gougerot, 1968: 235,236, pl. 1, fig. 2 [original spelling pseudo-circinata].Eocene, Upper Lutetian, Paris Basin. Montmirail,Seine-et-Marne, France. Fustiaria pseudocircinataGougerot, 1968 (fide Le Renard 1995: 175; LeRenard & Pacaud 1995: 85).pseudoentalis, see pseudoantalis.pseudoentalis, Dentalium O. G. Costa, 1851: 17, 18,pl. 1, fig. 2. <strong>Recent</strong>. “Golfo Tarantino” [Italy].Type material not located. Junior synonym <strong>of</strong>Antalis panorma (Chenu, 1843) (fide Pilsbry &Sharp 1897: 54; as “pseudo-antalis” non Lamarck,1818; misspelling corrected by Pilsbry & Sharp,1898: 250).† pseud<strong>of</strong>issura, Dentalium (Antalis) R. Janssen, 1978a:14-16, pl. 1, figs 1, 2, pl. 4, fig. 1; 1978b: 139, 140,pl. 9, fig. 3. Upper Oligocene. Kasseler Meeress<strong>and</strong>,Tagebau Höllkopf, near Glimmerode, Niederhessen,Germany. Holotype SMF 245901 ; paratypesSMF 245902, SMF 245903. For Dentaliumfissura auct. (reported from the Upper Oligocene)non Lamarck, 1818.pseudohexagonum, Dentalium Arnold, 1903: 186, 187,pl. 8, figs 12, 12a [ex Dall MS]. Pliocene (San PedroSeries) <strong>and</strong> Pleistocene. Deadman Isl<strong>and</strong>; San Pedro;San Diego, California. Junior synonym <strong>of</strong> Dentaliumneohexagonum Sharp & Pilsbry in Pilsbry &Sharp, 1897. Remark: Arnold described this taxon asa new <strong>species</strong>, even though he listed D. neohexagonumas a synonym; thus, Arnold’s name was publishedin synonymy <strong>and</strong> is, thus, not available.pseudohexagonum, Dentalium (Antalis) Henderson,1920: 46, 47, pl. 6, figs 1-3 [ex Ihering MS].[<strong>Recent</strong>]. Brazil. Holotype USNM 225817. Juniorhomonym <strong>of</strong> Dentalium pseudohexagonum Arnold,1903; replacement name: Dentalium (Antalis) pilsbryiRehder, 1941: 96. Antalis pilsbryi Rehder,1942 (fide Scarabino 1975: 183).pseudohexagonum “Deshayes, 1825”, see pseudosexagonum.† pseudohungaricus, Cadulus (Gadila) Szöts, 1953: 81[Hungarian], 203 [French], pl. 7, figs 53, 54.Eocene. Gánti-szölök, Hosszúharasztos <strong>and</strong> Újfeltárás,Hungary. Lectotype Hungarian GeologicalInstitute E.180 (designated by Boda 1964: 67 as“holotype”). Remark: this <strong>species</strong> was describedfrom three localities in Hungary. Boda (1964: 67),by stating that the specimen from Hosszúharasztoswas the holotype, selected that specimen as the lectotype<strong>and</strong> restricted the type locality.pseudolivi, Cadulus (Gadila) Boissevain, 1906: 73, 74,pl. 6, fig. 67, text-fig. 36. <strong>Recent</strong>. B<strong>and</strong>a Sea, Siboga634ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)stn 211, 05°40.7’S, 120°45.5’E, 1158 m. HolotypeZMA 3.06.101. Gadila pseudolivae (Boissevain,1906) (fide Scarabino 1995: 364). Known fromtype locality only. Remark: holotype incorrectlycited as a syntype by Scarabino (1995: 364) (fideR. Moolenbeek & A. N. Van der Bijl, in litt.23.VI.2000).† pseudomutabile, Lentigodentalium R. Janssen, 1989:85, 86, pl. 1, figs 10-13. Miocene, Hemmoorian/Reinbekian. Twistringen, Niedersachsen, Germany.Holotype SMF 308312; paratypes SMF 308313,SMF 308314, SMF 308315/50; RGM 227725.† pseudonyma, Dentalium Pilsbry & Sharp, 1898:213. Miocene. Astoria, Oregon, USA. Type specimenlost. Moore (1963: 50) selected the illustrationin Conrad (1849: pl. 20, fig. 7a) as the lectotype <strong>of</strong>this <strong>species</strong>. Replacement name for Teredo substriataConrad, 1849 non Dentalium substriatumDeshayes, 1825. Remark: Dentalium conradi Dall,1909, an unnecessary replacement name, is a juniorsynonym; see Emerson (1958) for further discussion.† pseudorakhiense, Dentalium (Laevidentalium)Eames, 1952: 6, pl. 1, figs 3a, 3b. Eocene. RakhiaNala section, Upper Chocolate Clays, near RakhiGaj, Pakistan. Holotype BMNH G.68014.pseudosexagonum, Dentalium Deshayes, 1825: 358[38], pl. 16, figs 14-16. <strong>Recent</strong>. Type locality notspecified. Lectotype <strong>and</strong> 5 paralectotypes MNHN(designated by Lamprell & Healy 1998: 63).Paradentalium pseudosexagonum (Deshayes,1825). Indo-Pacific: Red Sea to China Sea <strong>and</strong>Australia. 3-499 m. Dentalium (Paradentalium)pseudohexagonum “Deshayes, 1825” [Habe &Kosuge, 1964: 1] is an incorrect subsequent spelling.† pukaea, Fissidentalium Maxwell, 1992: 185, pl. 30,figs a-d. Eocene. NZGS localities GS9508 <strong>and</strong> GS11200, Waihao River, near McCulloch’s Bridge,South Canterbury, New Zeal<strong>and</strong>. Holotype NZGSTM7225; numerous paratypes, NZGS.pulcherrimus, Cadulus (Gadila) Boissevain, 1906: 74,75, pl. 6, figs 58, 59, text-fig. 38. <strong>Recent</strong>. FloresSea, Siboga stn 314, 07°36’S, 117°30.8’E, 694 m.Lectotype ZMA 3.06.104 (designated by Scarabino1995: 334); paralectotype ZMA 3.06.105 (5).Striocadulus pulcherrimus (Boissevain, 1906) (fideScarabino 1995: 334). Indo-Pacific: Indonesia.530-694 m.† pulchrum, Dentalium Wilckens, 1911: 30, 31, pl. 1,fig. 38. Tertiary. Seymour Isl<strong>and</strong>, Antarctica. HolotypeSMNH 2968 (fide Stilwell & Zinsmeister1992: 173). Dentalium (Dentalium) pulchrumWilckens, 1911 (fide Stilwell & Zinsmeister 1992:173, 174).punctatostriatum, Dentalium Gümbel, 1861: 274.Triassic. Lodensee, Germany. Nomen nudum.pusillum, Dentalium Philippi, 1836. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.pusillum, Dentalium (Ditrupa “?”) Gabb, 1864: 139,pl. 21, fig. 99. Cretaceous. NE <strong>of</strong> Martinez; AlizosCreek, near Fort Téjon; <strong>and</strong> Tuscan Springs,California, USA. Lectotype ANSP 79582 (designatedby Emerson 1957: 988, 989, pl. 26, figs 1, 5)(from Martinez); paralectotype ANSP 79583.Junior homonym <strong>of</strong> Dentalium pusillum Philippi,1836. Replacement name: Cadulus gabbi Pilsbry &Sharp, 1898. Remark: Gabb (1869: 230) transferredthis <strong>species</strong> to “Gadus”.pusillum, Siphodentalium Watson, 1879: 520 [Watson1886: 14, pl. 2, fig. 6]. <strong>Recent</strong>. Palma, Canaries,Challenger stn 85, 28°42’N, 18°06’W, 1125 fms[2025 m]. Syntypes BMNH 1887.2.9.64 (2).Pulsellum pusillum (Watson, 1879) (fide Pilsbry &Sharp 1897: 140). Known from the type localityonly.pygmaeus, Dentalium Defrance, 1819. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† pyrum, Dentalium Pilsbry & Sharp, 1898a: 472,pl. 11, figs 6, 7. Oligocene. Dominican Republic.Type material ANSP 2714 (fide Pilsbry 1922: 400).† qingyanicum, Prodentalium Stiller, 2001: 626-630,figs 5.8-5.12, 6. Upper Anisian, Middle Triassic.Bangtoupo, NNE <strong>of</strong> Qingyan, Guizhou Province,southwestern China. Holotype GPIM B6D-1.S3.Fbt-1; paratypes GPIM B6D-1.S3.F30-1,GPIM B6D-1.S3.F30-2, GPIM B6D-1.S3.Fbt-2.quadrangulare, Dentalium Sowerby in Broderip &Sowerby, 1832: 29. <strong>Recent</strong>. Xipixapi [WestColombia], 7-11 fms [13-20 m]. Syntypes BMNH1950.11.8.10-14. Junior synonym: Dentalium fisheriPilsbry & Sharp, 1897 (fide Emerson 1956: 2).Tesseracme quadrangularis (Sowerby in Broderip& Sowerby, 1832) (fide Emerson 1956: 2). EasternPacific: Baja California to Ecuador. 0-37 m.quadrangulare, Dentalium Deshayes, 1835: 132,pl. 26, fig. 14. Mesozoic. Morea, Peloponnesus,Greece. Junior homonym <strong>of</strong> Dentalium quadrangulareBroderip & Sowerby, 1832. Replacementname: Dentalium subquadrangulare Emerson, 1954.quadrangularis, Entalina Boissevain, 1906: 62, 63,text-fig. 28, pl. 6, figs 73-75, 85, 86. <strong>Recent</strong>. OffKai-Isl<strong>and</strong>s [Indonesia], Siboga stn 256, 5°26.6’S,132°32.5’E, 397 m. Lectotype ZMA 3.06.081 (designatedby Habe 1964: 39; <strong>and</strong> unnecessarily byScarabino, 1995: 300); paralectotypes ZMA3.06.082 (1; Celebes Sea, Siboga stn 88, 0°34.6’N,119°08.5’E, 1301 m), ZMA 3.06.083 (HalmaheraSea, Siboga stn 151, 0°12.6’S, 129°48’E, 845 m).Junior synonym <strong>of</strong> Entalina mirifica (E.A. Smith,1895) (fide Scarabino 1995: 300). Remarks: Habe(1964: 39) gave the wrong type locality (Siboga stn88) for the lectotype specimen (fide R. Moolenbeek& A. N. Van der Bijl, in litt. 23.VI.2000). Entalinaquadriangularis [Habe 1963: 271] is an incorrectsubsequent spelling.quadrangulatum, Dentalium “Klipstein.” Manuscriptname; see Dentalium klipsteini Kittl, 1891.† quadrangulatum, Dentalium Hu & Lee, 1991: 121,figs 18, 19, 29-31. Pliocene. Henchun Peninsula,ZOOSYSTEMA • 2004 • 26 (4)635


Steiner G. & Kabat A. R.Maan Mountain, Taiwan. Holotype NMT 296;paratypes NMT 296a-d. Remark: this <strong>species</strong> probablybelongs to the genus Entalina.† quadrangulum, Dentalium Assmann, 1937: 98,pl. 18, fig. 22. Triassic. Gorasdze, Upper Silesia,Pol<strong>and</strong>.quadrapicale, Dentalium Hanley in Sowerby, 1860:103, pl. 225, fig. 61. <strong>Recent</strong>. Cochin, Malabar,Southwest India. Lectotype BMNH 1907.10.28.147 (designated by Ludbrook 1954: 103 as holotype);paralectotypes BMNH 1907.10.28.148-149 (2). Junior synonyms: Dentalium dipsychaPilsbry & Sharp, 1897; Dentalium conspicuumMelvill, 1897 (fide Scarabino 1995: 222).Tesseracme quadrapicalis (Hanley in Sowerby,1860) (fide Plate 1908a: 350). Indo-Pacific: Africato northern Australia. 6-743 m. Remark: Habe &Kosuge (1964: 5) synonymized T. tetrapleura(Boissevain, 1906) with this <strong>species</strong>.quadrata, Entalina Henderson, 1920: 88, pl. 15,figs 2, 3, 6, 10. <strong>Recent</strong>. Off Grenada [LesserAntilles], Blake stn 248, 12°02’35”N, 61°47’15”W,159 fms [290 m]. Holotype USNM 95354 (formerlya syntype <strong>of</strong> Dentalium callithrix Dall, 1889).Junior secondary homonym <strong>of</strong> Entalina quadrata(Martin, 1885). Junior synonym <strong>of</strong> Entalina platamodes(Watson, 1879) (fide Emerson 1952a: 7, 8;Scarabino 1975: 181).† quadratum, Dentalium Martin, 1885: 190, pl. 10,fig. 191; 1887: 190, pl. 10, fig. 191. Miocene.Grissee, Java, Indonesia. Syntypes RGM 7290 (5)(fide Van den Hoek Ostende et al. 2002: 112).Junior secondary homonym: Entalina quadrataHenderson, 1920. Entalina quadrata (Martin,1885) (fide Martin 1919: 69).quadriangularis, see quadrangularis.quadricostatum, Dentalium Brazier, 1877: 58. <strong>Recent</strong>.Princess Charlotte Bay, Queensl<strong>and</strong>, Australia,13 fms [24 m]. Lectotype AMS C170763 (ex AMS94ii) (designated by Lamprell & Healy 1998: 78);paralectotypes AMS C308638 (11), AMS C170762(11; Katow, New Guinea, 8 fms [15m]), AMSC170761 (1; York Isl<strong>and</strong>, Torres Strait, 13 fms[24m]). Eudentalium quadricostatum (Brazier,1877) (fide Cotton & Godfrey 1933: 140). Indo-Pacific: northern Australia <strong>and</strong> New Guinea. 9-124 m.quadridentatum, Siphonodentalium Dall, 1881: 36.<strong>Recent</strong>. West coast <strong>of</strong> Florida, 30 fms [55 m].Holotype MCZ 7739. Junior synonym <strong>of</strong>Polyschides tetraschistus (Watson, 1879) (fideScarabino 1980: 11). Remark: Emerson in Turgeon(1998: 55) recognized quadridentatum as a valid<strong>species</strong> <strong>of</strong> Polyschides without justifying the rejection<strong>of</strong> Scarabino’s synonymy.quadrifissatus, Cadulus (Polyschides) Pilsbry & Sharp,1898: 150, 151, pl. 29, figs 10-13 [ex CarpenterMS]. <strong>Recent</strong>. San Pedro; San Diego, California[32°45’N, 117°10’W], 10 fms [18 m]. SyntypesUSNM 19462 (1), ANSP 35583 (1). Polyschidesquadrifissatus (Pilsbry & Sharp, 1898). EasternPacific: California. 18 m. Remark: this <strong>species</strong> wasplaced in Siphonodentalium by Keen (1971: 890)<strong>and</strong> Emerson in Turgeon (1998: 55).† quadriturritus, Cadulus Meyer, 1886: 65, pl. 3,figs 7, 7a. Eocene. Red Bluff, Mississippi, USA.Syntype USNM Paleobiology 644578 (as “holotype”in MacNeil & Dockery 1984: 248, 249).† quatuordecimcostata, Dentalium (Antale “?”) dentalevar. Sacco, 1897: 105, pl. 8, figs 76, 77 [as a nomennudum in Sacco 1896: 97]. Pliocene. Italy (variouslocalities). Syntypes MIGT BS.106.02.067,BS.106.02.068 (fide Ferrero Mortara et al. 1984: 306).† quenstedti, Dentalium Blake, 1875: 226 (citingQuenstedt 1857: 797, pl. 98, fig. 20). Jurassic.Söflingen, Germany. Remark: Blake (1875) basedthis <strong>species</strong> upon a figure: “Quenstedt does not givea name to the Dentalium figured by him, whichdoes not appear to differ from those found in theKimmeridge Clay generally throughout Engl<strong>and</strong>.I therefore propose the above name in accordancewith a now accepted custom”.† quindeciesstriatum, Dentalium Eichwald, 1853: 137;pl. 3, fig. 19. Pliocene. Zukowce, Volhynia,Ukraine. Replacement name for Dentalium striatumEichwald, 1830 non Born, 1778. Antalis quindeciesstriata(Eichwald, 1853) (fide Harzhauser 2002).quinquangulare, Dentalium Forbes, 1844: 135, 188.<strong>Recent</strong>. Aegean Sea, 80-230 fms [46-420 m].Syntypes BMNH 1948.2.11.6-8 [3 syntypes in badcondition], BMNH 1948.4.8.28 (1). Junior synonym<strong>of</strong> Entalina tetragona (Brocchi, 1814) (fideMonterosato 1872: 27).quinquangulare, Dentalium Gümbel, 1861: 397, 409.Triassic. Bavaria, Germany. Junior homonym <strong>of</strong>Dentalium quinquangulare Forbes, 1844. Replacementname: Dentalium pentagonale Pilsbry &Sharp, 1898.† quitus, Cadulus (Polyschides) Pilsbry & Olsson,1941: 48, 49, pl. 10, figs 9, 10. Pliocene, CanoaFormation. Punta Blanca, western Ecuador.Holotype ANSP 13719 ; paratype ANSP 13718.radicula, Dentalium Lamarck, 1818. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† radiolineata, Dentalium Clark, 1918: 192, pl. 22,fig. 12. Oligocene, San Ramon Formation. NearMount Diablo, California, USA. Holotype UCMP11227.† radula, Dentalium Gmelin, 1791: 3738, 3739 [citingSchröter 1784a: 530 n. 9]. Miocene. Piedmont,Italy. Junior synonym: Dentalium asperumMichelotti, 1847 (fide Sacco 1897: 111). Coccodentaliumradula (Gmelin, 1791) (fide Sacco 1896: 98).† radularis, Dentalites Schlotheim, 1820: 95.Miocene. Piedmont, Italy. Remark: it is not apparentwhether this name is an emendation <strong>of</strong>Dentalium radula Gmelin.† rakhiense, Dentalium (Laevidentalium) Eames,1952: 5, 6, pl. 1, figs 2a, 2b. Eocene. Rakhi Nala636ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)section, Upper Chocolate Clays, near Rakhi Gaj,Pakistan. Holotype BMNH G.68007.† raricostata, Dentalium (Antale) <strong>fossil</strong>e var. Sacco,1897: 100, pl. 8, figs 32-41 [as a nomen nudum inSacco 1896: 97]. Miocene/Pliocene. Italy (variouslocalities). Syntypes MIGT BS.106.02.028-BS.106.02.034 (fide Ferrero Mortara et al. 1984:304, pl. 56, fig. 5).† raricostata, Dentalium (Coccodentalium) radula var.Sacco, 1897: 111, pl. 10, figs 19-21 [as a nomennudum in Sacco 1896: 98]. Miocene. San Agata,Italy. Syntypes MIGT BS.106.04.012, BS.106.04.014 (fide Ferrero Mortara et al. 1984: 308).† rarinodosum, Dentalium haeringense var. Báldi,Horvath & Makk, 1974: 27, pl. 2, fig. 11.Paleocene, Kiscellian. Budafok, S <strong>of</strong> Budapest,Hungary.rastridens, Cadulus Watson, 1879: 525 [Watson 1886:19, pl. 3, fig. 3]. <strong>Recent</strong>. Culebra Isl<strong>and</strong>, Challengerstn 24, 18°38’30”N, 65°05’30”W, 390 fms[712 m]. Syntypes BMNH 1887.2.9.73-75 (completelydisintegrated), BMNH 1994043 (11). Gadilarastridens (Watson, 1879) (fide Pilsbry & Sharp1898: 174). Western Atlantic, Caribbean. 500-712 m.† raymondi, Prodentalium Young, 1942: 120, 121,pl. 20, figs 3-6, 8, 12. Pennsylvanian, MagdalenaGroup. Near Talpa, Sangre de Cristo mountains,northern New Mexico, USA. Holotype MCZInvertebrate Paleontology 27938.razzorei, Cadulus (Gadila) Caprotti, 1979: 248, 249,pl. 15, figs 1-7. Lower Pliocene. Rio Torsero,Liguria, Italy. Holotype MSNM I 5569. Remark:Caprotti (1979: 248) proposed this name for“Gadila gadus var. gadula” Sacco, 1897 [ex DoderleinMS], which Caprotti did not consider to be avalid name. However, Ruggieri (1980: 95, 96)demonstrated that Sacco’s usage constitutes a validdescription; thus, C. (G.) razzorei is an junior synonym<strong>of</strong> gadula Sacco, 1897 (see also Pavia 1991:137, 138).† readi, Dentalium Emerson, 1954: 184. Triassic,Carnian. Seiser Alp, Tirol, Austria. Replacementname for Dentalium simile Read in Broili, 1907 nonDentalium simile Wissmann, 1841 nec Giunti,1859. Laevidentalium readi (Emerson, 1954) (fideStiller 2001: 622).rebecaense, see rebeccaense.rebeccaense, Dentalium (Dentalium) Henderson, 1920:31, 32, pl. 3, fig. 2. <strong>Recent</strong>. Between Tortugas <strong>and</strong>Rebecca Shoals, Florida, Eolis stn 33, 16 fms[29 m]. Holotype USNM 314293; paratypesUSNM 314294 (25), AMNH 148314 (1). Juniorsynonym <strong>of</strong> Paradentalium americanum (Chenu,1843) (fide Scarabino 1975: 183). Dentalium rebecaense[Scarabino in Rios 1975: 183; 1994: 305] isan incorrect subsequent spelling.recta, Cadulus strangulatus var. Locard, 1898: 136.<strong>Recent</strong>. Type locality not specified. Type materialnot located. Synonym <strong>of</strong> Gadila strangulata(Locard, 1897).rectius, Dentalium Carpenter, 1864: 648. <strong>Recent</strong>. PugetSound [Washington, USA]. Lectotype USNM 5283(designated by Palmer 1958: 118, 119, pl. 17, figs 1,2, as “holotype”); paralectotype MCZ 178535 (1).Junior synonym: Rhabdus watsoni Pilsbry & Sharp1897 (fide Shimek 1998: 83). Rhabdus rectius(Carpenter, 1864) (fide Pilsbry & Sharp 1897: 113).Eastern Pacific: Alaska to California. 91-1794 m.Remarks: although Carpenter’s (1864: 648) descriptionwas rather cursory, the name must be consideredavailable notwithst<strong>and</strong>ing the claim by Pilsbry &Sharp (1897: 114) that this <strong>species</strong> was not describedby Carpenter until the following year (Carpenter1865: 59). Shimek (1998: 83) listing <strong>of</strong> Rhabdus dalliPilsbry & Sharp, 1897 in synonymy with this <strong>species</strong>remains to be confirmed.† rectiusculum, Dentalium Eichwald, 1857: 583[nomen nudum in Eichwald 1846: 425]. Carboniferous[“Bergkalke”]. Kasakendörfer, Russia.Remark: figured by Eichwald (1860: 1062, 1063,pl. 40, fig. 12); locality given as Kasatschy-datschy,Ural, near Serpoukhow, Russia.rectum, Dentalium (Episiphon) kiaochowwanense var.Tchang & Tsi, 1950: 7, 8, pl. 1, figs 7, 8. <strong>Recent</strong>.Kiaochow Bay [China], stn 331, 10 m. HolotypeASP 3261; 6 paratypes ASP 3261, ASP 2897 (13;stn 314, 5 m). Synonym <strong>of</strong> Episiphon kiaochowwanense(Tchang & Tsi, 1950).† rectum, Dentalium Gmelin, 1791: 3738 [citingGualtieri 1742: t. 10. f. H; Martini, 1769: I. t. I.f. 4A; Schröter, 1784b, 4 t. 3. f. 5]. Type localitynot specified. Type material not located. Junior synonyms:Dentalium striatum Link, 1807 (based uponthe same figure from Schröter, 1784b); Dentaliumsulcatum Schumacher, 1817) (based upon the samefigure from Martini, 1769); Dentalium crassum SalisMarschlins, 1793 (cited the same figures asGmelin); Fissidentalium delessertianum (Chenu,1843) (fide Pilsbry & Sharp 1898: 213; Caprotti1979: 223). Remarks: Pilsbry & Sharp (1897: 81)synonymized Dentalium elephantinum “Linnaeus”Deshayes, 1825, partim with Dentalium rectum <strong>and</strong>used Deshayes’ locality information, “India”, toindicate a <strong>Recent</strong> occurrence, which they later correctedby citing Fischer (Pilsbry & Sharp: 1898:252). Fischer (1883) reported this <strong>species</strong> as livingin the Eastern Atlantic, which was questioned byCaprotti (1979: 223, 224). Thus, Fissidentaliumrectum is considered to be a valid <strong>fossil</strong> <strong>species</strong> butnot a <strong>Recent</strong> one.recurvum, Dentalium Deshayes, 1825: 30. <strong>Recent</strong>.Junior synonym <strong>of</strong> ?Dentalium elephantinumLinnaeus, 1758 (fide Pilsbry & Sharp 1897: 2; withthe epithet “ by error?”, probably applying toDeshayes new <strong>species</strong>).reesi, Cadulus (Gadila) Ludbrook, 1954: 115, fig. 20.<strong>Recent</strong>. Zanzibar area, John Murray Exp. stn 105B,ZOOSYSTEMA • 2004 • 26 (4)637


Steiner G. & Kabat A. R.5°34’24”-37’S, 39°14’06”-36”E, 238-293 m.Holotype BMNH 1952.3.25.150; paratypesBMNH 1952.3.25.390-BMNH 1952.3.25.394,BMNH 1952.3.25.385-BMNH 1952.3.25.388 (3;stn 176, Gulf <strong>of</strong> Aden, 12°04’06”N, 50°38’3”E,655-732 m), BMNH 1952.3.25.389 (1; stn 179B,Gulf <strong>of</strong> Aden, 12°02’06”N, 50°40’1”E, 275 m),BMNH 1952.3.25.395-BMNH 1952.3.25.401(7; stn 191, Gulf <strong>of</strong> Aden, 13°46’3”N, 47°48’5”E,274 m). Junior synonym <strong>of</strong> Gadila boissevainae(Jaeckel, 1932) (fide Scarabino 1995: 358).reevei, Dentalium Fischer, 1871: 212 [ex DeshayesMS]. <strong>Recent</strong>. Suez, Red Sea. Lectotype MNHN(designated by Scarabino 1995: 204; the remainingsyntypes were not located by Scarabino). Juniorsynonyms: Dentalium lineolatum Cooke, 1885;Dentalium laugieri Jousseaume, 1894 (fide Ludbrook1954: 98; Singer 2003: 14). Dentaliumreevei Fischer, 1871. Red Sea. 15-120 m. Remark:Singer (2003) restricted the distribution <strong>of</strong> D. reeveito the Red Sea.† regium, Dentalium Assmann, 1937: 97, 98, pl. 18,figs 19-21. Triassic. Gorasdze <strong>and</strong> Greater Strehlitz,Upper Silesia, Pol<strong>and</strong>. Laevidentalium regium(Assmann, 1937) (fide Stiller 2001: 622).regulare, Dentalium E. A. Smith, 1903: 393, pl. 15,fig. 2. <strong>Recent</strong>. Port Shepstone, Natal, South Africa.Syntypes BMNH 1903.9.9.24, BMNH 1903.7.26.29-32. Dentalium regulare E. A. Smith, 1903.Indian Ocean: South Africa. 10-22 m.regulare, Dentalium Ahlburg, 1906: 86, 87, pl. 3,fig. 2. Triassic. Granietz, Upper Silesia, Pol<strong>and</strong>.Junior homonym <strong>of</strong> Dentalium regulare E. A.Smith, 1903; replacement name: Dentalium subregularePalmer, 1974.regulare, Dentalium (Dentalium) laqueatum var.Henderson 1920: 26, pl. 1, fig. 8. <strong>Recent</strong>. Jamaica,73-140 fms [78-256 m]. Holotype USNM 107887.Junior homonym <strong>of</strong> Dentalium regulare E. A.Smith, 1903 <strong>and</strong> Ahlburg, 1906. Replacementname: Dentalium invalidum Emerson, 1954.regularis, Cadulus (Gadila) Henderson, 1920: 137,pl. 19, fig. 14. <strong>Recent</strong>. Off Cape Canaveral, Florida,USBF stn 2660, 28°40’10”N, 78°46’W, 504 fms[919 m]. Holotype USNM 887475; paratypesUSNM 108175, USNM 108278, USNM 330675,AMNH 148341 (21; stn 2668, 294 fms [537 m],<strong>of</strong>f Fern<strong>and</strong>ina, Florida; fide Boyko & Sage 1996:31). Gadila regularis (Henderson, 1920). WesternAtlantic, Caribbean. 536-919 m.† rei, Fissidentalium “?” Maxwell, 1992: 186, pl. 30,figs e, f. Eocene. NZGS localities GS1162,GS3274, GS11200, GS4872, Waihao River, nearMcCulloch’s Bridge, South Canterbury, NewZeal<strong>and</strong>. Holotype NZGS TM7227; numerousparatypes NZGS.† reticulatum, Dentalium Tournouër in de Boullé,1873: 466 [40], pl. 6, fig. 13. Eocene, Auversian.Peyrehorade, Adour Basin, southwestern France.Remark: the original description was reprinted byBoussac (1911a: 33).reussianum, Dentalium Ryckholt, 1851: 70, 71.Cretaceous: Turonian. Visé, Belgium. Junior synonym<strong>of</strong> Dentalium cidaris Geinitz, 1850 (fidePilsbry & Sharp 1898: 220).† rex, Dentalium Pilsbry & Sharp, 1898: 214. Replacementname for Dentalium giganteum Chenu,1842 non Phillips, 1829.rhabdotum, Dentalium Pilsbry, 1905: 116, 117, pl. 5,figs 45-47. <strong>Recent</strong>. Heda, Izu, Japan. Syntypes ANSP88319 (2), ANSP A4085 (1). Striodentalium rhabdotum(Pilsbry, 1905) (fide Okutani 1966: 12). Indo-Pacific: China Sea <strong>and</strong> Japan to New Caledonia.61-1350 m. Remark: Scarabino (1995: 240) statedthat ANSP 88319 was the “holotype” but there aretwo syntypes in that lot, so Scarabino’s statementdoes not constitute a valid lectotype designation.† rhenanus, Dentalium R. Janssen, 1978b: 141, pl. 9,figs 4, 5. Oligocene, Chattian. Schacht Diergardt,Rumeln, near Moers, Niederrhein-Gebeit, Germany.Holotype SMF 250385a.† rhodani, Dentalium Pictet & Roux, 1849: 150, 151,pl. 27, figs 13a-13e. Cretaceous. Rhône <strong>and</strong>Saxonet, France.“rhotomagense”, Dentalium – d’Orbigny 1850(volume 2): 156. Cretaceous, Cenomanian. Rouen,France. Incorrect original spelling <strong>of</strong> Dentaliumrothomagense. Remark: Rothomagus was the Latinname <strong>of</strong> Rouen.† richardsoni, Dentalium Cossmann, 1907b: 172,173. Jurassic, Pliensbachian. Railway-cutting,Dixton West, near Gotherington, near Cheltenham,Engl<strong>and</strong>, United Kingdom. Replacementname for Dentalium acutum Richardson, 1906 nonHébert, 1849. Objective junior synonym: Dentaliumacutoides Emerson, 1954. Laevidentaliumrichardsoni (Cossmann, 1907) (fide Engeser &Riedel 1992: 37).richardsoni, Dentalium Emerson, 1954: 184. Jurassic,Sinemurian. Honeybourne, near Evesham, Engl<strong>and</strong>,United Kingdom. Replacement name forDentalium subquadratum Richardson, 1906 nonMeek, 1860, itself a junior homonym <strong>of</strong> Dentaliumrichardsoni Cossmann, 1907; replacement name:Dentalium sabrinae Engeser & Riedel, 1992.† rigauxi, Fustiaria (Laevidentalium) Fischer & Vadet,1985: 5, pl. 1, figs 8a, 8b. Middle Jurassic,Bathonian. Hydrequent, Pas de Calais, France.Holotype MHNB 115. Remark: this <strong>species</strong> wasmisidentified as Dentalium entaloides Eudes-Deslongchamps, 1842 by Cossmann (1885).† rimosum, Dentalium Böse, 1906: 55, 56, pl. 3,fig. 1. Upper Miocene, Division Rio Coatzacoalcos.Kilometres 123-124 del Ferrocarril de Tehuantepec,Mexico. Holotype UNAM 16374 (fideAlencaster Ibarra 1950: 222, fig. 1).rioense, Dentalium (Dentalium) texasianumHenderson, 1920: 29, pl. 2, figs 2, 3. <strong>Recent</strong>. East638ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)<strong>of</strong> Rio de Janeiro, USBF stn 2762, 23°08’S,41°34’W, 59 fms [108 m]. Holotype USNM96114. Junior synonym <strong>of</strong> Paradentalium americanum(Chenu, 1843) (fide Scarabino 1975: 183).† ripleyanum, Dentalium Gabb, 1860: 393, pl. 69,fig. 48. Cretaceous, Ripley Group. Eufala,Alabama, USA.† robusta, Plagioglypta Detre, 1993: 164, pl. 1. UpperPermian. Bükk Mountains, NE <strong>of</strong> Nagyvisnó,Hungary.robustum, Dentalium Brazier, 1877: 56. <strong>Recent</strong>.Katow, New Guinea, 8 fms [15 m]. Lectotype AMSC170765 (designated by Ponder & Stanbury 1972:53); paralectotypes AMS C170766 (3). Dentaliumrobustum Brazier, 1877. Indo-Pacific: New Guinea,northern Australia. 13-3150 m.robustum, Dentalium Maurer, 1885: 233, 234, pl. 10,fig. 1. Middle Devonian. Grube Rothe Erde, nearGiessen, Germany. Junior homonym <strong>of</strong> Dentaliumrobustum Brazier, 1877. Replacement name:Laevidentalium patriciae Engeser & Riedel, 1992.rosea, Dentalium dentale var. Dautzenberg, 1884: 302[ex Monterosato MS]. <strong>Recent</strong>. Mediterranean Sea:Gulf <strong>of</strong> Gabes, Tunisia. Type material not located.Synonym <strong>of</strong> Antalis dentalis (Linnaeus, 1767).Remark: Dautzenberg appears to be the first authorto have validated Monterosato’s (1878) nomennudum.rosea, Dentalium vulgaris var. Bucquoy, Dautzenberg& Dollfus, 1882: 561. <strong>Recent</strong>. Beach <strong>of</strong> Saint-Briac, Côtes d’Amor, France. Synonym <strong>of</strong> Antalisvulgaris (Da Costa, 1778).rossati, Dentalium (Antalis) Caprotti, 1966: 194-196,fig. 11. <strong>Recent</strong>. Shiqmona Bay, Haifa, Israel.Holotype <strong>and</strong> 9 paratypes in MSNM. Antalisrossati (Caprotti, 1966). Mediterranean. 3-25 m.rossoi, Cadulus (Cadulus) Nicklès, 1979: 62, textfig.18, pl. 1, figs IV, V. <strong>Recent</strong>. Off Cape Vert,Senegal, Atlantide stn 10-1-56A, 150-250 m.Holotype <strong>and</strong> paratypes MNHN. Cadulus rossoiNicklès, 1979. Eastern Atlantic, Africa. 150-250 m.† rothomagense, Dentalium d’Orbigny, 1852 (Index):59. Cretaceous, Cenomanian. Rouen, France.Emendation <strong>of</strong> Dentalium rhotomagense d’Orbigny,1850. Remark: Rothomagus was the Latin name <strong>of</strong>Rouen.† rotundatior, Dentalium inaequale var. Sacco, 1897:96, pl. 7, figs 74, 75 [as a nomen nudum in Sacco1896: 97]. Miocene. Italy (various localities).Syntypes MIGT BS.106.01.024, BS.106.01.025(fide Ferrero Mortara et al. 1984: 302).† rotundosimplex, Dentalium michelotti var. Sacco,1897: 96, pl. 7, figs 91, 92 [as a nomen nudum inSacco 1896: 97]. Pliocene. Italy (various localities).Syntypes MIGT BS.106.01.040, BS.106.01.041(fide Ferrero Mortara et al. 1984: 303).† rotundulina, Dentalium michelotti var. Sacco, 1897:96, pl. 7, figs 89, 90 [as a nomen nudum in Sacco1896: 97]. Pliocene. Italy (various localities).Syntypes MIGT BS.106.01.038, BS.106.01.039(fide Ferrero Mortara et al. 1984: 302).rowei, Dentalium (Dentalium) Lamprell & Healy,1998: 38, figs 29C, 30C, 33. <strong>Recent</strong>. 3.2 km NE <strong>of</strong>Cape Bossut, northern Western Australia, 7-10 m.Holotype WAM 231/94; paratypes WAM 53-93 (1), WAM 67-93 (2; 24°54’S, 113°17.75’E,13 km NE <strong>of</strong> Redcliff Point, Bernier Isl<strong>and</strong>,Western Australia, 24.7 m), WAM 66-93(2; 24°52’S, 113°17.5’E, ENE <strong>of</strong> Redcliff Point,Bernier Isl<strong>and</strong>, Western Australia), AMS C201738(12; Exmouth Gulf, northern Western Australia).Dentalium rowei Lamprell & Healy, 1998. IndianOcean: western Australia. 7-196 m.rubescens, Dentalium Deshayes, 1825: 363 [43 inreprint], pl. 16, figs 23-25. <strong>Recent</strong>. Mediterranean Sea.Syntypes MNHN. Junior synonyms: Dentalium splendensO. G. Costa, 1829; Pseudantalis tenuifissumMonterosato, 1875; Dentalium maltzani Clessin,1896 (fide Pilsbry & Sharp 1897: 106); Dentaliumsiculum Pilsbry & Sharp, 1897 [ex Deshayes MS](n. syn.). Fustiaria rubescens (Deshayes, 1825) (fideEmerson 1952c: 204; 1962: 471). Eastern Atlantic,Mediterranean: Europe, North Africa. 4-618 m.Remarks: Scarabino (1995: 282) questioned the record<strong>of</strong> this <strong>species</strong> from the Indian Ocean by Ludbrook(1954: 105, 106). Dentalium rufescens [Weinkauff,1868: 420] is an incorrect subsequent spelling.rudis, ?Dentalium Gabb, 1873. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.rudmani, Cadulus Lamprell & Healy, 1998: 164,figs 167E, 168H, 173. <strong>Recent</strong>. Off Cairns, northernQueensl<strong>and</strong>, Australia, 17°09.42’S, 146°42.24’E, 668 m. Holotype AMS C174626; paratypeAMS C173407 (1). Cadulus rudmani Lamprell &Healy, 1998. Known from the type locality only.rudoi, Paradentalium Scarabino, 1995: 219, 220,figs 27, 28j, l. <strong>Recent</strong>. Northeastern Madagascar,RV Vauban 12°39’S, 48°17’E, 240 m. HolotypeMNHN; paratypes MNHN (6 from the type locality;3 from BENTHEDI stn DS72, 12°71’S,45°02’E, 300-350 m), NM (1). Paradentaliumrudoi Scarabino, 1995. Indian Ocean: Madagascar.240-350 m.rufescens, see rubescens.† rugiferum, Dentalium Koenen, 1885: 71, 72, pl. 3,fig. 18. Paleocene. Near Copenhagen, Denmark.Junior synonym: Dentalium imparcostata Nechaev,1897 (fide Makarenko 1970: 113). Remark: a valid<strong>species</strong> <strong>of</strong> Dentalium (fide Makarenko 1970: 113).ruginosa, Dentalium dentalis var. “Monterosato” fideStork 1934: 5. No further information located;potentially a nomen nudum.† rugosum, Dentalium Dunker, 1849: 16, 17.Cretaceous. Near Cassel, Germany. Unnecessaryreplacement name: Dentalium dunkeri Sharp &Pilsbry in Pilsbry & Sharp, 1898: 221, 222.rugosum, Dentalium Müller, 1851. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.ZOOSYSTEMA • 2004 • 26 (4)639


Steiner G. & Kabat A. R.rugosum, Dentalium Eichwald, 1857: 584 [1860:1063, 1064, pl. 40, fig. 8]. Carboniferous[“Berglehme”]. Near Dorf Sloboda, Tula [Toula],Russia. Junior homonym <strong>of</strong> Dentalium rugosumDunker, 1849 <strong>and</strong> D. rugosum Müller, 1851.Remarks: Eichwald (1860: 1063) erroneously datedthis name to Eichwald (1846: 425), which is actuallya citation for Dentalium rectiusculum Eichwald;Pilsbry & Sharp (1898: 232) repeated this error. Iffuture studies confirm the validity <strong>of</strong> this <strong>species</strong> itneeds to be renamed.rugosum, Dentalium Spillman, 1860: 389. Nomennudum. Cretaceous (Green S<strong>and</strong>). Tombigbee,Mississippi, USA.rugosum, Entalium Defrance, 1819b. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† runnegari, Scissuradentalium Yoo, 1988: 249,figs 144-147. Early Carboniferous, Late Tournaisian,Dangarfield Formation. Locality A-3,2.6 km SW <strong>of</strong> Gundy, Upper Hunter, New SouthWales, Australia. Holotype AMS F.61994 ;paratypes AMS F.61995 (8).† rupeliensis, Gadila R. Janssen, 1989: 88, 89, pl. 2,fig.17. Middle Oligocene, Rupelian, Boom ClayFormation. Herselt (Tongrube 400 m N <strong>of</strong> Kirchevon Ramsel), Antwerpen Province, Belgium.Holotype RGM 221152a; paratypes RGM221152b; RGM 220907/1; SMF.rushii, Cadulus (Gadila) Pilsbry & Sharp, 1898: 168,pl. 27, figs 94-97. <strong>Recent</strong>. Off Cape Hatteras[North Carolina], 35°19’30”N, 75°17’12”W, 293fms [535 m]. Syntypes ANSP 35579 (1), ANSP333440 (1). Junior synonym: Polyschides rushii arne(Henderson , 1920). Polyschides rushii (Pilsbry &Sharp, 1898) (fide Emerson in Turgeon 1998: 55).Western Atlantic: Canada to North Carolina (USA).210-1934 m.† rutteni, Dentalium Martin, 1917: 262, pl. 4, fig. 91.Miocene. Kembang Sokkoh, Java, Indonesia. SyntypesRGM 7256 (3), RGM 7257 (14), RGM 47284(2) (fide Van den Hoek Ostende et al. 2002: 112).† sabrinae, Dentalium Engeser & Riedel, 1992: 38,39, 41. Jurassic, Sinemurian. Honeybourne, nearEvesham, Engl<strong>and</strong>, United Kingdom. Replacementname for Dentalium richardsoni Emerson, 1954 nonCossmann, 1907. Remark: Engeser & Riedel(1992: 39) stated that “The generic assignment isunclear. It probably belongs to a new genus”.† saccoi, Cadulus taurovulum subsp. Pavia, 1991: 133-137, pl. 9, figs 1-5, 7, 8; table 2; text-fig. 6. UpperMiocene. Borelli, near Turin, Italy. Holotype <strong>and</strong>218 paratypes MIGT B-S.D.02.1. Remark: Sacco(1897: 115) misidentified this <strong>species</strong> as Cadulusovulus Philippi.† sacheri, Dentalium Alth, 1850: 226, pl. 12, fig. 2.Cretaceous. Lviv [Lemberg], Ukraine. HolotypeNHM 1862/0005/0251.sagamiensis, Gadila Kuroda, Habe & Oyama, 1971:495 [in Japanese], 312 [in English], pl. 65, figs 14-17 (as a nomen nudum in Habe 1953: 298; 1963:276; Okutani 1964: 79). <strong>Recent</strong>. Sagami Bay,Japan, 400-740 m. Holotype <strong>and</strong> 3 paratypes inImperial Household Biological Laboratory (Japan).Gadila sagamiensis Kuroda, Habe & Oyama, 1971.Indo-Pacific: Japan. 400-780 m.sagei, Striocadulus Scarabino, 1995: 335, 336,figs 137, 139 d, e, 172c, d. <strong>Recent</strong>. Philippines, RVCoriolis MUSORSTOM 2 stn CP24, 13°37’S,120°42’E, 640-647 m. Holotype <strong>and</strong> 4 paratypesMNHN. Striocadulus sagei Scarabino, 1995. Indo-Pacific: Indonesia, Philippines. 550-800 m.† sagittarii, Dentalium Pilsbry & Johnson, 1917: 185.Oligocene. Dominican Republic. Holotype ANSP652. Remark: figured by Pilsbry (1922: 400, pl. 18,fig. 12).sakuraii, Platyschides Kuroda & Habe in Habe, 1961:105, pl. 47, fig. 3. <strong>Recent</strong>. Off Choshi, Chiba Pref.,Honshu, Japan, 100-200 m. Holotype <strong>and</strong> paratypein NSMT. Polyschides sakuraii (Kuroda & Habe inHabe, 1961) (fide Habe 1963: 278). Indo-Pacific:Japan. 100-1400 m. Platyschides sokuraii [Okutani,1966: 14] is an incorrect subsequent spelling.† salicensis, Cadulus Seguenza, 1879: 276. Pliocene.Reggia, Calabria, Italy.salishorum, Pulsellum Marshall, 1980: 149-152, figs 1-6.<strong>Recent</strong>. East Sound, Orcas Isl<strong>and</strong>, San Juan Isl<strong>and</strong>s,Puget Sound, Washington, 48°36’N, 122°51’W,18-22 m [lower limit probably in error as the holotypeis from 27 m]. Holotype USNM 782263;paratypes USNM 782264 (6), ANSP 352474 (4),AMNH 198610 (6), LACM 1929 (6), NMC 86068(4), WSM 35663 (4). Pulsellum salishorumMarshall, 1980. Eastern Pacific, Canada, USA. 3-91 m.salpinx, Dentalium (Fissidentalium) Tomlin, 1931:338, 339, text-fig. <strong>Recent</strong>. Off Cape Point, SouthAfrica, 700-800 fms [1278-1460 m]. LectotypeSAFM A5459 (designated by Giles & Gosliner1983: 40); paralectotypes NMW (9). Fissidentaliumsalpinx (Tomlin, 1931). South Africa.1260-1462 m.† samanicum, Dentalium Berry, 1926: 19, figs 3, 4.Eocene, Saman shale. Negritos, Peru.† s<strong>and</strong>bergeri, Dentalium Bosquet, 1859: 20, 21, pl. 2,figs 7a-7d. Tertiary (Eocene?). Limburg, Netherl<strong>and</strong>s.Remark: Deshayes (1861: 215) recognizedthat Dentalium fissura sensu Nyst (1836: 180) nonLamarck, 1818 was equivalent to Dentalium s<strong>and</strong>bergeri.† sangiorgii, Dentalium Emerson, 1954: 184.Pliocene. Ponticella di Sàvena, near Bologna, Italy.Replacement name for Antale dentale var. alternansSangiorgi, 1926 [ex Berti MS] non Dentalium alternansChenu, 1843 nec Müller, 1851 nec Ryckholt,1851 nec Bucquoy, Dautzenberg & Dollfus, 1882.Antalis sangiorgii (Emerson, 1954) (fide Caprotti1979: 237).† santarosanum, Dentalium Maury, 1910: 151, pl. 8,fig. 6. Oligocene. Oak Grove, Santa Rosa County,640ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Florida, USA. Syntype USNM Paleobiology639095 (1).† saturni, Dentalium Hoeninghaus, 1836a: 1, pl. 1[nomen nudum in Hoeninghaus 1836b: 12]. “Übergangskalk”,Gerolstein, Eifel Mountain, Germany[Devonian]. Remarks: this <strong>species</strong> has been attributedto Goldfuss, 1841 by most authors, whoconcluded that Goldfuss had validated a manuscriptname <strong>of</strong> Hoeninghaus (1836b: 12). However,Langer (1967: 137) has shown that the <strong>species</strong> wasfirst validated by Hoeninghaus (1836a), in a lithographedpublication, which included an illustration<strong>of</strong> the holotype. It remains to be determined whetherGoldfuss’ concept <strong>of</strong> this <strong>species</strong> is equivalent tothat <strong>of</strong> the original description in Hoeninghaus(1836a). Yochelson & Holl<strong>and</strong> (2004) concludedthat the supposed “holotype” <strong>of</strong> Goldfuss’ namewas not the type specimen, <strong>and</strong> that the description<strong>of</strong> this <strong>species</strong> was too imprecise to allow determination.We do not follow their conclusion that this isprobably not a scaphopod until the problem withthe type is solved.† saucatsense, Dentalium Benoist, 1873: 423, 460[239, 276]. Miocene, Helvetian. Sime, France.Remark: this <strong>species</strong> was misidentified as Dentaliumpseudoantalis Lamarck, 1818 by Benoist (1873: 268[84]); see Cossmann & Peyrot (1917: 165) forfurther discussion.sauridens, Cadulus Watson, 1879: 525, 526 [Watson1886: 19, pl. 3, fig. 4]. <strong>Recent</strong>. West Indies,Culebra Isl<strong>and</strong>, Challenger stn 24, 18°38’30”N,65°05’30”W, 390 fms [712 m]. Holotype BMNH1887.2.9.76. Gadila sauridens (Watson, 1879)(fide Pilsbry & Sharp 1898: 173). Western Atlantic,Caribbean. 650-712 m. Remark: BMNH 1994044is also labelled “holotype”, obviously in error as thedescription referred to a single specimen.scamnatum, Dentalium Locard, 1897b: 10 [ex FischerMS] [Locard, 1898: 109, 110, pl. 7, figs 1-7].<strong>Recent</strong>. “Côtes occidentales d’Afrique, cap Ghir, lesAçores, mer des Sargasses entre 1235 et 2087mètres”. Syntypes MNHN. Junior synonym <strong>of</strong>Fissidentalium capillosum (Jeffreys, 1877)(B. Métivier, n. syn.).scarabinoi, Cadulus n. nom. <strong>Recent</strong>. New Caledonia,Bassin des Loyauté, RV Coriolis BIOGEOCAL stnCP232, 21°34’S, 166°27’E, 760-790 m. HolotypeMNHN; paratypes MNHN (3 from CP232; 1from CP75; 2 from DW106). Replacement namefor Cadulus martini Scarabino, 1995 non Cadulusmartini Finlay, 1927. Cadulus scarabinoi Steiner &Kabat, 2004. Indo-Pacific: Réunion Isl<strong>and</strong> to NewCaledonia. 625-3500 m.scarabinoi, Dentalium n. nom. <strong>Recent</strong>. ChesterfieldIsl<strong>and</strong>s, Mer de Corail, Banc Nova, RV Coriolis,MUSORSTOM 5. stn DW 301, 22°07’S, 159°25’E,478-610 m. Holotype MNHN; paratypes MNHN(10; 1 from stn 301; 1 from stn DW 306; 1 fromstn LAGON 444; 2 from stn DW 72; 1 fromstn DW 38; 1 from stn DW 01; 1 from stn DW 05;1 from stn DW 478; 1 from stn DW 479), AMSC201722 (1; stn DW 479), NMNZ M268960(1; stn DW 479). Replacement name for Dentaliumflavum Scarabino, 1995 non Dentalium flavumHenderson, 1920. Dentalium scarabinoi Steiner &Kabat, 2004. Indo-Pacific: Chesterfield Isl<strong>and</strong>s(Coral Sea); New Caledonia. 300-610 m.† schafferi, Dentalium Gugenberger, 1934: 45, pl. 1,fig. 9 [nomen nudum in Gugenberger 1933a: 102;1933b: 184]. Triassic. N <strong>of</strong> Launsdorf, Austria.† schencki, Dentalium (Rhabdus) Moore, 1963: 51,pl. 31, fig. 3. Miocene, Astoria Formation. SpencerCreek, Oregon, USA. Holotype USNM Paleobiology563335.† schlosseri, Dentalium Traub, 1938: 65, pl. 5, fig. 8.Cretaceous/Tertiary. Rupertiwinkel, N <strong>of</strong> Salzburg,Austria.† schumoi, Dentalium Pilsbry, 1911: 167, 168, fig. 5[nomen nudum in Pilsbry & Sharp 1898: 254].Miocene. Bowden, Jamaica. Syntype ANSP 653.scoticum, Dentalium Kirkby, 1880: 563, 589 [exYoung MS]. Nomen nudum. Carboniferous. NearSt Andrews, Scotl<strong>and</strong>, United Kingdom.† sectiforme, Dentalium (Graptacme) Tate, 1899: 262,263, pl. 8, figs 6-6a. Miocene. Muddy Creek,Victoria, Australia. Holotype <strong>and</strong> 5 paratypes SAMT 1615A. Dentalium (Antalis) sectiforme Tate, 1899(fide Ludbrook 1959: 143, pl. 2, fig. 5).sectum, Dentalium Deshayes, 1825: 367, 368 [47, 48],pl. 18, figs 12-14. <strong>Recent</strong>. “Habitat in mari Indico”[Indian Ocean]. Syntypes MNHN. Junior synonym:Dentalium diffusum Chenu, 1850 (fidePilsbry & Sharp 1897: 96). Graptacme secta(Deshayes, 1825) (fide Pilsbry & Sharp 1897: 96).Geographical range unknown. Remark: the citationby Keen (1971: 886) from the “Gulf <strong>of</strong> California”was in error (fide Keen & Coan 1975: 49 citing W.K. Emerson in litt.).sedecimcostatum, Dentalium Boissevain, 1906: 33,pl. 6, figs 8-11. <strong>Recent</strong>. B<strong>and</strong>a Sea, Indonesia,Siboga stn 52, 09°03.4’S, 119°56.7’E, 959 m.Lectotype ZMA 3.06.035 (designated by Scarabino1995: 260); paralectotypes ZMA 3.06.034 (1;stn 45, 07°24’S, 118°15.2’E, 794 m), ZMA3.06.036 (8), ZMA 3.06.037 (1; stn 88, 00°34.6’N,119°8.5’E, 1301 m), ZMA 3.06.038 (1; stn 156,00°29.2’S, 130°5.3’E, 469 m), ZMA 3.06.039(1; stn 178, 02°40’S, 128°37.5’E, 835 m), ZMA3.06.040 (1; stn 223, 05°44.7’S, 126°27.3’E,4391 m), ZMA 3.06.041 (1; stn 241, 04°24.3’S,129°49.3’E, 1570 m), ZMA 3.06.042 (8, fideR. Moolenbeek & A. N. Van der Bijl in litt.23.VI.2000; stn 300, 10°48.6’S, 123°23.1’E,918 m), ZMA 3.06.043 (1; stn 314, 07°36’S,117°30.8’E, 694 m). Compressidentalium sedecimcostatum(Boissevain, 1906) (fide Habe & Kosuge1964: 4). Indo-Pacific: East Africa to NewCaledonia <strong>and</strong> China Sea. 250-4391 m. Remark:ZOOSYSTEMA • 2004 • 26 (4)641


Steiner G. & Kabat A. R.Lamprell & Healy (1998: 87) listed this <strong>species</strong> as ajunior synonym <strong>of</strong> C. clathratum (von Martens,1881) (as Fissidentalium), an opinion rejected byScarabino (1995).† seguenzianus, Cadulus (Gadila) Moroni & Ruggieri,1980: 49, 50, text-figs 3a, 3b, 4. Upper Miocene.Near Bonfornello, Palermo, Sicily, Italy. Holotype<strong>and</strong> paratype Museo di Geologia della Università diPalermo.† semarangense, Dentalium Altena, 1938: 211.Miocene. Ngembak <strong>and</strong> Grissee, Java, Indonesia.Syntypes RGM 7239 (1), RGM 7240 (1), RGM7238 (1), RGM 7241 (1) (fide Van den HoekOstende et al. 2002: 112). Replacement name forDentalium subrectum Martin, 1885 non Jeffreys,1883. Junior synonym <strong>of</strong> Dentalium teschiKoperberg, 1931. Remark: see under Dentaliumsubrectum Martin, 1885.† semialternans, Dentalium Chenu, 1843: pl. 4, fig. 7;1850: 7. Type locality not specified (presumablyTertiary, fide Pilsbry & Sharp 1898: 215).† semiaratum, Dentalium Chapman & Crespin, 1928:105, 106, pl. 6, fig. 28. Tertiary. Victoria, Australia.† semiclausum, Dentalium Nyst, 1835: 36, pl. 5,fig. 53. Tertiary. Near Antwerp, Belgium.† semicostatum, Dentalium Girty, 1911: 135.Carboniferous, Pennsylvanian. USGS Locality2006, Wewoka Formation, Wewoka quadrangle,Oklahoma, USA. Junior synonym: Dentalium indianumGirty, 1911 (fide Yancey 1978: 308, by FirstReviser action). Prodentalium semicostatum (Girty,1911) (fide Yancey 1978: 308). Remark: Yancey(1978) erroneously dated this <strong>species</strong> to itsredescription in Girty (1915), not to its originaldescription in 1911.† seminuda, Dentalium (Antale) raricostatum “race”Cossmann & Peyrot, 1917: 168, pl. 1, figs 29-31.Miocene, Tortonian. Saint-Jean-de-Marsacq,France.† seminudum, Dentalium Deshayes, 1861: 200, 201,pl. 3, figs 11-14. Eocene. Paris Basin (Jeures <strong>and</strong>Ormoy), France; Kaufingen, Drausfeldt <strong>and</strong> nearCassel, Germany.semipolitum, Dentalium Broderip & Sowerby, 1829:369. <strong>Recent</strong>. Type locality not specified [EasternPacific, California]. Syntypes BMNH 1950.11.8.1-3.Junior synonym <strong>of</strong> Graptacme semistriata (Turton,1819) (fide Pilsbry & Sharp 1897: 91). Remarks:Emerson in Turgeon (1998: 54) listed Graptacmesemipolita as a distinct <strong>species</strong>; further study <strong>of</strong> thetaxonomy <strong>of</strong> these <strong>species</strong> is needed.semistriatum, Dentalium Turton, 1819: 39, pl. 18,fig. 68. <strong>Recent</strong>. Near the Pigeon-House, DublinBay, Irel<strong>and</strong> [in error]. Syntype USNM 175006 (1).Junior synonyms: Dentalium semipolitum Broderip& Sowerby, 1829; Dentalium semistriolatumGuilding, 1834; Dentalium translucidum Chenu,1843; Dentalium hyalinum Philippi, 1846;Dentalium liratum Carpenter, 1857b; Dentaliumlirulatum Mörch, 1861 (fide Pilsbry & Sharp 1897:90, 91); Dentalium sericatum Dall, 1881 (fideScarabino 1994: 307); Dentalium hannai Baker,1925 (fide Keen 1971: 886). Graptacme semistriata(Turton, 1819) (fide Pilsbry & Sharp 1897: 90).Western Atlantic, Caribbean: Bermuda to Brazil;Eastern Pacific: California. 3-164 m. Remarks:Warén (1983: 169) erroneously stated that the typematerial was lost. Since Jeffreys could not determinethe origin <strong>of</strong> this <strong>species</strong>, Warén also stated that“I regard the name as a nomen dubium”. The allegedoccurrence <strong>of</strong> this <strong>species</strong> in both the Caribbean<strong>and</strong> the Eastern Pacific may represent a case <strong>of</strong> sibling<strong>species</strong>. The second specimen mentioned in theoriginal description was not located.semistriatum, Dentalium Deshayes, 1825: 367, pl. 17,figs 15, 16. Eocene. Parnes, Chaumont, Mouchy<strong>and</strong> Senlis, Paris Basin, France. Junior homonym <strong>of</strong>Dentalium semistriatum Turton, 1819. Replacementname: Dentalium parisiensis d’Orbigny, 1850.semistriatus, Cadulus Jeffreys, 1880: 317. <strong>Recent</strong>. Bay<strong>of</strong> Biscay. Nomen nudum.semistriolatum, Dentalium Guilding, 1834: 34, pl. 3,figs 1-5. <strong>Recent</strong>. Caribbean. Type material notlocated. Junior synonym <strong>of</strong> Graptacme semistriata(Turton, 1819) (fide Pilsbry & Sharp 1897: 90).Remarks: Emerson in Turgeon (1998: 54) listedGraptacme semistriolata as a distinct <strong>species</strong>; furtherstudy <strong>of</strong> the taxonomy <strong>of</strong> these <strong>species</strong> is needed.semitracheatum, Dentalium (Plagioglypta) Boissevain,1906: 56, pl. 4, figs 20, 21. <strong>Recent</strong>. Timor Sea,Siboga stn 284, 08°43.1’S, 127°16.7’E, 828 m.Lectotype ZMA 3.06.072 (designated by Scarabino1995: 273); paralectotypes ZMA 3.06.070 (7, fideR. Moolenbeek & A. N. Van der Bijl in litt.23.VI.2000; Flores Sea, Siboga stn 45, 7°24’S,118°15.2’E, 794 m), ZMA 3.06.071 (1; not mentionedin the text but illustrated on plate 4, figs 20,21; “St. 248” in caption was error for 284, fideR. Moolenbeek & A. N. Van der Bijl in litt.23.VI.2000), ZMA 3.06.073 (2; Timor Sea, Sibogastn 300, 10°48.6’S, 123°23.1’E, 918 m), ZMA3.06.074 (1 [2 in error in original description];Flores Sea, Siboga stn 314, 7°36’S, 117°30.8’E,694 m). Junior synonym: Dentalium curvotracheatumPlate, 1908 (fide Scarabino 1995: 273).Calliodentalium semitracheatum (Boissevain,1906) (fide Scarabino 1995: 273). Indo-Pacific:Madagascar to Philippines. 450-918 m.semivestitum, Dentalium Locard, 1897b: 9, 10 [exFischer MS] [1898: 107, pl. 6, figs 25-29]. <strong>Recent</strong>.“Les Tropiques et le Sahara entre 830 et 1113 mètres”.Syntypes MNHN (Talisman stn 85, “Sahara”,830 m). Fissidentalium semivestitum (Locard,1897) (fide Pilsbry & Sharp 1897: 75). EasternAtlantic: Europe, Africa. 800-1113 m.senegalense, Dentalium Dautzenberg, 1891: 53, 54,pl. 3, fig. 8. <strong>Recent</strong>. Dakar, Senegal, Mélita Exp.stn 99, 5 m. Type material not located. Antalis642ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)senegalensis (Dautzenberg, 1891) (fide Pilsbry &Sharp 1897: 55). Eastern Atlantic: Senegal toGuinea. 5-41 m.senegalensis, Cadulus Locard, 1897a: 3, 4. <strong>Recent</strong>.Senegal. Holotype MNHN. Gadila senegalensis(Locard, 1897) (fide Pilsbry & Sharp 1898: 176).Known from the type locality only. Remark: typelocality subsequently cited as Talisman 1883stn 101, 3200 m, by Locard (1898: 133).septangulare, Dentalium Fleming, 1825. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† septemcostata, Dentalium (Antale “?”) taurocostatumvar. Sacco, 1897: 102, pl. 8, fig. 58 [as a nomennudum in Sacco 1896: 97]. Miocene. Turin hills,Italy. Syntype MIGT BS.106.02.050 (fide FerreroMortara et al. 1984: 305).septemcostatum, Dentalium Abich in Trautschold,1859. Not a scaphopod. See Other taxa described as<strong>Scaphopoda</strong>.septemcostatum, Dentalium Brazier, 1877: 57. <strong>Recent</strong>.Evan’s Bay, Cape York, Queensl<strong>and</strong>, Australia,6 fms [11 m]. Lectotype AMS A96 (designated byPonder & Stanbury 1972: 53); paralectotypes AMS(3). Junior homonym <strong>of</strong> Dentalium septemcostatumAbich, 1859. Replacement name: Dentaliumcheverti Sharp & Pilsbry in Pilsbry & Sharp, 1897.septemcostatum, Dentalium coarti var. Dautzenberg,1912: 81. <strong>Recent</strong>. Wharf de Tamara, Guinea. Typematerial MNHN. Junior homonym <strong>of</strong> Dentaliumseptemcostatum Abich, 1859. Junior synonym <strong>of</strong>Dentalium congoensis (Plate, 1908a) (fide Nicklès1979: 44).septentrionalis, Antalis Kuroda & Habe in Habe, 1963:262, 263, pl. 38, fig. 34, text-figs 15-17 [nomennudum in Habe 1953: 294, as “Dentalium(Dentalium) septentrionale”, <strong>and</strong> in Taki & Oyama1954: 30, pl. 7, figs 19-21, pl. 26. fig. 6]. <strong>Recent</strong>.Off Hachinoe, Aomori Pref., Honshu, Japan, 30 m.Lectotype NSMT (designated by Oyama 1973: 72,pl. 20, figs 7, 11, 14); paralectotype NSMT. Juniorsynonym <strong>of</strong> Antalis weinkauffi (Dunker, 1877)(fide Habe 1964: 20).† seriaeforme, Siphonodentalium Ravn, 1939: 48, 49,pl. 1, figs 18a, 18b. Paleocene. Vestre Gasvaerk VI,near Copenhagen, Denmark.sericatum, Dentalium Dall, 1881: 37, 38. <strong>Recent</strong>. OffYucatan Bank, Blake (no stn given), 640 fms[1168 m]. Holotype USNM 203191. Known fromthe type locality only. Junior synonym <strong>of</strong> Graptacmesemistriata (Turton, 1819) (fide Scarabino1994: 307).serratum, Dentalium Pictet & Roux, 1849. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.serratum, Dentalium Martin, 1885: 189, pl. 10,fig. 189; 1887: 189, 190, pl. 10, fig. 189. Miocene.Grissee, Java, Indonesia. Holotype RGM 7270 (fideVan den Hoek Ostende et al. 2002: 112). Juniorhomonym <strong>of</strong> Dentalium serratum Pictet & Roux,1849 (Polychaeta); replacement name: Dentaliumsubserratum Palmer, 1974.serrulatum, Dentalium E. A. Smith, 1906: 249.<strong>Recent</strong>. Andaman Isl<strong>and</strong>, Bay <strong>of</strong> Bengal, 60 fms[110 m]. Syntypes ZSI M877/1 (fide list inBMNH); BMNH 1906.10.12.4. Fissidentaliumserrulatum (E. A. Smith, 1906) (fide Scarabino1995: 250). Indo-Pacific: Andaman Isl<strong>and</strong>s toPhilippines. 96-182 m.sewelli, Dentalium (Episiphon) Ludbrook, 1954: 107,fig. 10. <strong>Recent</strong>. Gulf <strong>of</strong> Oman, John Murray Exp.stn 75, 25°09’-10’N, 56°47’30”E, 201 m. HolotypeBMNH 1952.3.25.125; paratypes BMNH1952.3.25.73 (1; stn MBII(c), Hadramut Coast,17°33’30”N, 56°01’30”E, 29 m), BMNH1952.3.25.236-BMNH 1952.3.25.236.242 (5;stn 188, Gulf <strong>of</strong> Aden, 13°43’-46’N, 47°50’-56’E,528 m), BMNH 1952.3.25.243 (1; stn 103,Zanzibar Area, 05°39’30”S, 39°11’30”E, 101 m),BMNH 1952.3.25.244-BMNH 1952.3.25.244.250 (6). Junior synonym <strong>of</strong> Episiphon subtorquatum(Fischer, 1871) (fide Scarabino 1995: 286).sexangulare, Dentalium Lamarck, 1818: 344. Italy.Remark: the original description tentatively listedthis name as a synonym <strong>of</strong> Dentalium sexangulumGmelin, 1791.sexangulare, Dentalium “n. sp.?” Hilgard & Hopkins,1878: 48, pl. 3, fig. 7. Quaternary. Lake Borgne,Louisiana, USA. Junior homonym <strong>of</strong> Dentaliumsexangulare Lamarck, 1818. Oldest available name:Dentalium gouldii Dall, 1889 (fide Pilsbry & Sharp1897: 21).† sexangulum, Dentalium Gmelin, 1791: 3739 [citingSchröter 1784a: 531 n. 10]. Pliocene. Laretto, Italy.Junior synonyms: Dentalium sexangulare Lamarck,1818 (tentatively in the original description);Dentalium inaequale Bronn, 1831 (fide Bernasconi1996: 26, as synonym <strong>of</strong> D. orsum); Dentaliumorsum Bonelli in Sismonda, 1842 (fide Sismonda1847: 24; Pilsbry & Sharp 1898: 205); Dentaliumnoe Sismonda, 1847 (tentatively, fide Pilsbry &Sharp 1898: 215); Dentalium sexangulare var. acutangularis,<strong>and</strong> var. subrecta Cocconi, 1873 (fidePilsbry & Sharp 1898: 215).sexcarinatum, Dentalium Goldfuss, 1841. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.sexcostatum, Dentalium Sowerby, 1860: 103, pl. 223,fig. 11. <strong>Recent</strong>. Type locality not specified.Holotype BMNH 1996084. Junior synonym <strong>of</strong>Paradentalium hexagonum (Gould, 1859) (fideScarabino 1995: 218). Remark: The synonymization<strong>of</strong> Paradentalium hexagonum with this <strong>species</strong>by Scarabino (1995) was not followed by Lamprell& Healy (1998: 58).† sexdecimcostata, Dentalium (Antale “?”) dentale var.Sacco, 1897: 105, pl. 8, figs 74, 75 [as a nomennudum in Sacco 1896: 97]. Pliocene. Italy (variouslocalities). Syntypes MIGT BS.106.02.065,ZOOSYSTEMA • 2004 • 26 (4)643


Steiner G. & Kabat A. R.BS.106.02.066 (fide Ferrero Mortara et al. 1984:306).sexradiatum, Dentalium Goldfuss – Pilsbry & Sharp1898: 244. Not a scaphopod. See Other taxadescribed as <strong>Scaphopoda</strong>.shirleyae, Fissidentalium Lamprell & Healy, 1998: 84,figs 82B, 83B, 85, 86A, B. <strong>Recent</strong>. SW <strong>of</strong>Imperieuse Reef, Rowley Shoals, 250 km NNW <strong>of</strong>Western Australia, 18°06’S, 118°10’E, 356 m.Holotype WAM 234/94; paratypes AMS C201740(2, ex WAM 4340.83), QM MO52532 (1, exWAM 4340.83), WAM 4304.83 (38), WAM 119-93 (1; 220 km NW <strong>of</strong> Port Hedl<strong>and</strong>, WesternAustralia, 18°44’S, 117°20’E, 326 m). Fissidentaliumshirleyae Lamprell & Healy, 1998. Indo-Pacific: Western Australia. 326-356 m.shopl<strong>and</strong>i, Dentalium Jousseaume, 1894: 102. <strong>Recent</strong>.50 miles from Aden, 670 fms [1223 m]. LectotypeMNHN (designated by Ludbrook 1954: 100, as“holotype”). Junior synonyms: Dentalium transversostriatumBoissevain, 1906; Dentalium chuni Plate,1908a (fide Scarabino 1995: 248). Fissidentaliumshopl<strong>and</strong>i (Jousseaume, 1894) (fide Plate 1908a:341, as Dentalium (Fissidentalium) chuni). Indo-Pacific: Red Sea to New Caledonia. 528-1350 m.siberutense, Dentalium Plate, 1908a: 348, pl. 30,figs 17-20. <strong>Recent</strong>. Siberut Isl<strong>and</strong>s, Indonesia,Valdivia stn 191, 00°39’S, 98°52’E, 750 m.Holotype ZMB 61093 (fide Kilias 1995: 176).Junior synonym <strong>of</strong> Entalinopsis intercostata(Boissevain, 1906) (fide Scarabino 1995: 305).siberutensis, Cadulus (Cadulus) Jaeckel, 1932: 309,310, fig. 7. <strong>Recent</strong>. Siberut Isl<strong>and</strong>s, Indonesia,Valdivia stn 191, 00°39.2’S, 98°52.3’E, 750 m.Lectotype ZMB 75368a (designated by Kilias 1995:176); paralectotypes ZMB 75368b (2), ZMB75369 (48). Cadulus siberutensis Jaeckel, 1932.Known from the type locality only.sibogae, Dentalium (Fissidentalium) Boissevain, 1906:39, pl. 4, figs 17, 18, text-fig. 22. <strong>Recent</strong>.Indonesia, Siboga stn 159, 00°59.1’S, 129°48.8’E,411 m. Holotype ZMA 3.06.048. Compressidentaliumsibogae (Boissevain, 1906) (fideScarabino 1995: 264). Indo-Pacific: Indonesia,Japan. 200-411 m.† siciliensis, Dentalium sexangulare var. Brugnone,1880: 132. Pliocene. Caltanisetta <strong>and</strong> Altavilla,Italy. Junior synonym <strong>of</strong> Dentalium sexangulumGmelin, 1791 (fide Bellini 1909: 221).siculum, Dentalium (Laevidentalium) Pilsbry & Sharp,1897: 107, pl. 19, fig. 6 [ex Deshayes MS]. <strong>Recent</strong>.Adriatic Sea <strong>and</strong> Ionian Isl<strong>and</strong>s. Type material notlocated. Replacement name for Dentalium politum“Linnaeus” sensu O. G. Costa 1850: 23 (non politumLinnaeus, 1767). Junior synonym <strong>of</strong> Fustariarubescens (Deshayes, 1825) (n. syn.).sigsbeanum, Dentalium Dall, 1881: 38. <strong>Recent</strong>.Yucatan Strait, 640 fms [1168 m]. Lectotype MCZ7737 (designated by Turner 1955: 319 as “holotype”);paralectotype USNM 95347 (1). Junior synonym<strong>of</strong> Bathoxiphus ensiculus (Jeffreys, 1877) (fideDall 1889: 428).† simile, Dentalium Wissmann, 1841: 91, pl. 9, fig. 8.Triassic. St Cassian, Tirol, Austria/Italy. Remark:Dentalium simile – Laube 1870 non Wissmann,1841 is a misidentified Orthoceras (fide Broili 1907:22).simile, Dentalium Biondi, 1859: 120, pl. 1, fig. 6.<strong>Recent</strong>. Sicily, Italy. Type material not located.Junior homonym <strong>of</strong> Dentalium simile Wissmann,1851. Junior synonym <strong>of</strong> Antalis dentalis Linnaeus,1758 (fide Pilsbry & Sharp 1897: 53).simile, Dentalium Read in Broili, 1907: 71, 72, pl. 6,fig. 4. Triassic, Carnian. Seiser Alp, Tirol, Austria.Non Dentalium simile Wissmann, 1841 nec Biondi,1859; replacement name: Dentalium readiEmerson, 1954.simillimus, Cadulus Watson, 1879: 526, 527 [Watson1886: 20, pl. 3, fig. 6]. <strong>Recent</strong>. Raine (sic) Isl<strong>and</strong>,Cape York, Queensl<strong>and</strong>, Australia, Challenger stn185, 11°35’S, 144°03’E, 155 fms [284 m].Lectotype BMNH 1887.2.9.80/1 (designated byLamprell & Healy 1998: 161); paralectotypesBMNH 1887.2.9.80/2 (2; Challenger stn 185).Cadulus simillimus Watson, 1879. Indo-Pacific:Australia to New Caledonia. 5-1646 m. Remark:Lamprell & Healy (1995: 161, 162) explained thatthe second syntype lot, BMNH 1887.2.9.81(Challenger stn 187), was erroneously labeled ascontaining the figured specimen, <strong>and</strong> is insteadCadulus vincentianus Cotton & Godfrey, 1940.† simplex, Dentalium Michelotti, 1861: 136, pl. 12,figs 12, 13. Lower Miocene. Dego, Mioglia, Pareto,Italy. Laevidentalium simplex (Michelotti, 1861)(fide Bonci et al. 2000: 207, 208, pl. 1, fig. 3).simplex, Dentalium (Compressidens) Pilsbry & Sharp,1897: 125, 126, pl. 27, figs 88, 89. <strong>Recent</strong>. OffTillamook Harbor, Oregon, USBF 3346, 786 fms[1434 m]. Syntypes USNM 107700, USNM600414. Junior homonym <strong>of</strong> Dentalium simplexMichelotti 1861. Replacement name: Compressidensstearnsii (Pilsbry & Sharp, 1898).† simplicior, Dentalium (Entalis) taurostriata var.Sacco, 1897: 109, pl. 9, figs 53, 54 [as a nomennudum in Sacco 1896: 97]. Miocene. Turin hills,Italy. Syntypes MIGT BS.106.03.028, BS.106.03.029 (fide Ferrero Mortara et al. 1984: 308).simpsoni, Cadulus (Platyschides) Henderson, 1920:127, 128, pl. 19, fig. 17. <strong>Recent</strong>. MayaguezRoadstead, Puerto Rico, USBF stn 6062, 25 fms[46 m]. Holotype USNM 161580; paratypesUSNM 314677, USNM 314932, AMNH 148357(1). Cadulus simpsoni Henderson, 1920. WesternAtlantic, Caribbean. 46-575 m. Remark: the subgenusPlatyschides was put in synonymy with Gadilaby Scarabino (1995: 196, 355). However, this<strong>species</strong> has its maximum diameter not in the anteriorthird <strong>of</strong> the shell as it is characteristic for Gadila,644ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)but in the middle <strong>of</strong> the shell as it is typical forCadulus.† simrothi, Cadulus Pilsbry, 1911: 168, 169, fig. 4.Miocene. Bowden, Jamaica. Syntype ANSP 650. Asa nomen nudum in Pilsbry & Sharp 1898: 238.singaporensis, Cadulus (Gadila) Sharp & Pilsbry inPilsbry & Sharp, 1898: 195, pl. 36, figs 30, 31.<strong>Recent</strong>. Singapore. Syntypes ANSP 35584 (4),ZMA 2.98.001 (2). Gadila singaporensis (Sharp &Pilsbry in Pilsbry & Sharp, 1898). Known from thetype locality only.sinuosum, Dentalium Boissevain, 1906: 28, pl. 6,fig. 22. <strong>Recent</strong>. Indonesia, Timor Sea, Siboga stn294, 10°12’S, 124°27’E, 73 m. Lectotype ZMA3.06.021 (designated by Scarabino 1995: 206);paralectotypes ZMA 3.06.018 (1, second specimenlost, fide R. Moolenbeek & A. N. Van der Bijl inlitt. 23.VI.2000; Kw<strong>and</strong>ang-bay-entrance, Sibogastn 114, 0°58.5’N, 122°55’E, 75 m), ZMA3.06.019 (1, lost fide R. Moolenbeek & A. N. V<strong>and</strong>er Bijl in litt. 23.VI.2000; between isl<strong>and</strong>sWowoni <strong>and</strong> Buton, Siboga stn 204, 4°20’S,122°58’E, 75-95 m), ZMA 3.06.020 (5 in originaldescription; 6 fide R. Moolenbeek & A. N. Van derBijl in litt. 23.VI.2000). Junior synonym <strong>of</strong>Dentalium leucoryx Boissevain, 1906 (fideScarabino 1995: 206).† sobrantensis, Dentalium radiolineata var. Clark,1918: 192. Oligocene. Near Mount Diablo,California, USA. Holotype UCMP 11261.s<strong>of</strong>iae, Cadulus Scarabino, 1995: 352, figs 157, 160i,l. <strong>Recent</strong>. Réunion, MD32 stn DS151, 20°51’S,56°03’E, 3300-3240 m. Holotype MNHN;paratypes MNHN (3 from DS151; 1 from DS11;2 from CH87; 1 from CH90), NM (1; DS149).Cadulus s<strong>of</strong>iae Scarabino, 1995. Indo-Pacific:Madagascar to New Caledonia. 1600-3716 m.† sokkohense, Dentalium Martin, 1917: 262, pl. 4,fig. 92. Miocene. Kembang Sokkoh, Java,Indonesia. Holotype RGM 7271 (fide Van denHoek Ostende et al. 2002: 111).sokuraii, see sakuraii.† solidissimum, Dentalium Pilsbry & Brown, 1917:37, pl. 5, fig. 8. Oligocene. Near Cartagena,Colombia. Holotype ANSP 3057. Remark theauthors suggested that this <strong>species</strong> “is probably aFissidentalium”.† solidum, Dentalium Hutton, 1873b: 2. Miocene,Pareora Formation. Kanieri; Sherry River; Waikari,New Zeal<strong>and</strong>. Lectotype NZGS TM.4809 (designatedby Keyes 1972: 96). Fissidentalium solidum(Hutton, 1873) (fide Beu & Maxwell 1990: 424).solidum, Dentalium Verrill, 1884: 215, 216, pl. 44,figs 16, 17. <strong>Recent</strong>. Off Georges Banks, USBF stn2083, 959 fms [1750 m]. Lectotype USNM 34687(designated by Henderson 1920: 63); paralectotypes:USNM 34687 (USBF stn 2083, 959 fms[1750 m]), USNM 34904 (USBF stn 2077, 1255fms [2290 m]), USNM 34688 <strong>and</strong> USNM 34911(USBF stn 2084, 1290 fms [2354 m]), USNM35636 (USBF stn 2103, 1091 fms [1991 m]),USNM 35635 (USBF stn 2111, 938 fms [1712m]), USNM 35645 (USBF stn 2115, 843 fms[1538 m]), AMNH 148370 (3; USBF stn 2084,1290 fms [2354 m]). Junior homonym <strong>of</strong>Dentalium solidum Hutton, 1873. Replacementname: Dentalium (Fissidentalium) meridionale verrilliHenderson, 1920. Junior synonym <strong>of</strong>Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877) (fidePilsbry & Sharp 1897: 72).† soliticum, Dentalium Piette, 1855: 1122. Jurassic.Rumigny, France.sominium, Laevidentalium Okutani, 1964: 75, 76,text-fig. 3. <strong>Recent</strong>. Sagami Bay, Japan, 35°03.6’N,139°22.4’E, 1320-1400 m. Holotype NSMTMo.69572; paratypes NSMT Mo.69573 (47).Episiphon sominium (Okutani, 1964) (fide Steiner1999: 155). Indo-Pacific: Japan. 620-1400 m.Remarks: Okutani (1964: 75) listed material fromthe following Soyo-maru stations: 1 specimen fromstn T29, Sagami Bay, 35°04.8’N, 139°30.2’E, 620-780 m; 6 specimens from stn T29, Sagami Bay,35°05.5’N, 139°29.3’E, 710-770 m; 23 specimensfrom stn T28, Sagami Bay, 35°03.6’N, 139°22.4’E,1400 m; 24 specimens from stn T28, Sagami Bay,35°01.0’N, 139°19.6’E, 1385 m; 9 Specimens fromstn T28’, Sagami Bay, 35°03.5’N, 139°24.5’E,1190-1220 m; 26 specimens from stn 169, SagamiBay, 35°04.9’N, 139°33.2’E, 780 m; 1 specimenfrom stn 9, Sea <strong>of</strong> Kashima-Nada, 36°19.2’N,141°15.6’E, 1030 m; about 20 specimens from stn17, Sea <strong>of</strong> Kashima-Nada, 36°19.7’N, 141°09.0’E,870 m. None <strong>of</strong> these match in number <strong>of</strong> specimenswith the located paratype lot which is labelled“Sagami Bay” <strong>and</strong> may represent a partial combinationfrom these stations.† sorbii, Dentalium King, 1850: 218. Permian.Connigsborough, near Doncaster, Irel<strong>and</strong>.Holotype Sheffield City Museum H.27.11.80.777(fide Riley 1974: 36). Junior synonym: Dentaliumspeyeri Geinitz, 1852 (fide Pilsbry & Sharp 1898:233).† soriense, Dentalium Eames, 1952: 5, pl. 1, figs 1a,1b. Eocene. Zinda Pir section, Ghazij Shales, N <strong>of</strong>Dera Ghazi Khan, Pakistan. Holotype BMNHG.68000.† sougraignense, Dentalium Cossmann, 1904: 637,pl. 3, figs 12, 13. Cretaceous, Santonian. Sougraigne,Aude, France.sowerbyi, Dentalium Guilding, 1834: 35, pl. 3, fig. 7.<strong>Recent</strong>. Caribbean. Type material not located.Synonym: Episiphon sowerbyi pelliceri (Henderson,1920). Episiphon sowerbyi (Guilding, 1834) (fidePilsbry & Sharp 1897: 117). Western Atlantic,Caribbean: Cape Hatteras (North Carolina, USA)to Brazil. 7-329 m.sowerbyi, Dentalium Chenu, 1843: pl. 6, fig. 2; 1850:7. Type locality not specified. Non DentaliumZOOSYSTEMA • 2004 • 26 (4)645


Steiner G. & Kabat A. R.sowerbyi Guilding, 1834. Tentative junior synonym<strong>of</strong> Antalis entalis (Linnaeus, 1758) (fide Pilsbry &Sharp 1898: 216).sowerbyi, Dentalium Michelotti, 1847. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.soyomaruae, Bathoxiphus Okutani, 1964: 77, 78, textfig.4. <strong>Recent</strong>. 56 miles <strong>of</strong>f Aogashima Isl<strong>and</strong>,Japan, 32°20’N, 140°55.5’E, 3150-3350 m.Holotype NSMT Mo.9574 (fide Anonymous 1996:82) <strong>and</strong> 2 paratypes. Bathoxiphus soyomaruaeOkutani, 1964. Indo-Pacific: Madagascar to Japan<strong>and</strong> New Caledonia. 975-5750 m. Remark:Chistikov (1983: 183) synonymized Rhomboxiphuscolmani (Palmer, 1974) with B. soyomaruae, a decisionrejected by Scarabino (1995: 314, 317).† spaethi, Progadilina Engeser & Riedel, 1992: 47, 48,pl. 1, fig. 4, text-figs 3, 4. Jurassic, Middle Liassic,Pliensbachian. Hoisdorf, Schleswig-Holstein,Germany. Holotype GPIMH 2516/3; paratypeGPIMH 2516/4.† speciosum, Dentalium Gümbel, 1861: 668. Eocene.Bavaria, Germany.spectabilis, Cadulus Verrill, 1885a: 432, 433, pl. 44,fig. 19. <strong>Recent</strong>. 180 miles <strong>of</strong>f Ocean City [Maryl<strong>and</strong>],USBF stn 2221, 39°05’30”N, 70°44’30”W,2780 m. Lectotype USNM 37935 (designated byHenderson 1920: 107; USNM “40498” error forUSNM 37935); paralectotypes USNM 38116(USBF stn 2043, 1467 fms [2677 m]), USNM38691, USNM 73051, MCZ 186801 (1), MCZ186802 (USFC 2174, 1594 fms [2909 m]).Polyschides spectabilis (Verrill, 1885) (fide Pilsbry& Sharp 1898: 153). Western <strong>and</strong> Eastern Atlantic:Caribbean to West Africa. 62-3389 m.speyeri, Dentalium Geinitz, 1852: 198. Permian.Rückingen, Wetterau, Germany. Junior synonym<strong>of</strong> Dentalium sorbii King, 1850 (fide Pilsbry &Sharp 1898: 233).† spiniforme, Dentalium Martin, 1885: 191, 192,pl. 10, fig. 193; 1887: 191, 192, pl. 10, fig. 193.Miocene. Ngembak, Java, Indonesia. SyntypesRGM 7277 (9) (fide Van den Hoek Ostende et al.2002: 112).† spiniformis, Cadulus (Gadila) Gardner, 1947: 630,pl. 57, fig. 19. Oligocene. Alum Bluff Group, ShoalRiver Formation. USGS locality 5079, 0.5 milesbelow Shell Bluff, Shoal River, Walton County,Florida, USA. Syntype USNM Paleobiology351645 (1).spinulosum, Dentalium “Miller, MSS” Pilsbry &Sharp, 1898. Not a scaphopod. See Other taxadescribed as <strong>Scaphopoda</strong>.spirale, Dentalium Risso, 1826b. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.† spitiense, Dentalium Gümbel, 1866: 360, pl. 1,fig. 7. Triassic. Near Spiti, Hamachal Pradesh,northwestern India [formerly Tibet].splendens, Dentalium O. G. Costa, 1829: cxxv, cxxvi.<strong>Recent</strong>. Golfo di Taranto [Italy]. Type material notlocated. Junior synonym <strong>of</strong> Fustiaria rubescens(Deshayes, 1825) (fide Pilsbry & Sharp 1897: 106).splendens, Dischides Raines, 2002: 37, 38, fig. 47.<strong>Recent</strong>. Tahai, Easter Isl<strong>and</strong>, 50-80 m. HolotypeLACM 2949; paratypes LACM 2950, ANSP410697 (2), AMS (2), Bailey-Matthews ShellMuseum (Florida) 1383 (2). Dischides splendensRaines, 2002. Pacific Ocean, Easter Isl<strong>and</strong>, 50-80 m.splendidum, Dentalium Sowerby in Broderip &Sowerby, 1832: 29. <strong>Recent</strong>. Xipixapi, West Colombia,10-16 fms [18-29 m]. Syntypes BMNH1950.11.8.8-9. Graptacme splendida (Sowerby inBroderip & Sowerby, 1832) (fide Pilsbry & Sharp1897: 96). Eastern Pacific: Colombia. 10-110 m.Remark: Keen (1971: 888) placed this <strong>species</strong> intoFustiaria (Laevidentalium).spretus, Cadulus (Gadilia [sic]) Tate & May, 1900:102. <strong>Recent</strong>. Port Esperance, Tasmania, 24 fms[44 m]. Lectotype TMH E1202/8543 (specimenon left h<strong>and</strong> side <strong>of</strong> board; ex TM1095) (designatedby Lamprell & Healy 1998: 156); paralectotypesTMH E1202/8543 (specimen on right h<strong>and</strong> side <strong>of</strong>board), SAM D 15848 (21), SAM D 16002 (5).Gadila spreta (Tate & May, 1900) (fide Cotton &Godfrey 1933: 148). Indo-Pacific: Tasmania,Australia, New Zeal<strong>and</strong>. 9-1770 m. Remark: SAMD 16009, potential type material, consists <strong>of</strong> “fivedry shells with no locality or collection data”(Zeidler & Macphail 1978: 384).stanisici, Bathoxiphus Lamprell & Healy, 1998: 142,figs 145G, 149, 150A. <strong>Recent</strong>. Off Cairns, northernQueensl<strong>and</strong>, Australia, 16°43.06’S, 146°37.18’E, 1477 m. Holotype AMS C173420; paratypeAMS C173421 (1). Bathoxiphus stanisici Lamprell& Healy, 1998. Known from the type locality only.Remark: V. Scarabino (in litt. 5.XI.2001) suggestedthat the status <strong>of</strong> B. stanisici Lamprell & Healy,1998 vis-à-vis Solenoxiphus striatulus Chistikov,1983 requires further research.stapes, Dentalium (Gadilina) Boissevain, 1906: 50, 51,pl. 5, figs 16-20, pl. 6, figs 79, 81, 83. <strong>Recent</strong>.B<strong>and</strong>a Sea, Indonesia, Siboga stn 212, 05°54.5’S,120°19.2’E, 462 m. Syntypes ZMA 3.06.061-062(2). Junior synonym <strong>of</strong> Gadilina insolita(E. A. Smith, 1894) (fide Habe 1964: 32).stearnsii, Dentalium Pilsbry & Sharp, 1898: 253,pl. 27, figs 88, 89. <strong>Recent</strong>. Off Tillamook Harbor,Oregon, 45°30’N, 125°42’W, 786 fms [1434 m].Syntypes USNM 107700, 600414. Replacementname for Dentalium (Compressidens) simplex Pilsbry& Sharp, 1897 non Michelotti, 1861. Compressidensstearnsii (Pilsbry & Sharp, 1898). Indo-Pacific:Japan; Eastern Pacific: Oregon. 1434 m.steineri, Bathoxiphus Lamprell & Healy, 1998: 142,figs 141E, 145F, 148, 166E, F. <strong>Recent</strong>. NE [<strong>of</strong>]Thursday Isl<strong>and</strong> [Australia], 10°51’S, 145°49’E,1379 m. Holotype QM MO40078; paratypes QMMO40079 (3), QM MO40080 (2; northern646ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Queensl<strong>and</strong>, 10°01’S, 145°00’E, 1777 m), QMMO40081 (12; northern Queensl<strong>and</strong>, 17°46’S,147°49’E, 1224 m), AMS C175551 (2; <strong>of</strong>f Cairns,northern Queensl<strong>and</strong>, 16°43.1-42.1’S, 146°37.3-36.4’S 1468-1477 m). Bathoxiphus steineriLamprell & Healy, 1998. Indo-Pacific:Queensl<strong>and</strong>, Australia. 950-1777 m. Remark:V. Scarabino (in litt. 5.XI.2001) suggested that thestatus <strong>of</strong> B. steineri Lamprell & Healy, 1998 vis-àvisSolenoxiphus striatulus Chistikov, 1983 requiresfurther research.stellata, Entalinopsis Scarabino, 1995: 307, 308,figs 110, 111a, b, 115b. <strong>Recent</strong>. Philippines,MUSORSTOM 2, stn CP18, 14°00’N, 120°18’E,188-195 m. Holotype MNHN; paratypes MNHN(5 from stn CP55; 1 from stn DR94), AMSC201730 (1; stn CP55), USNM 890871 (1;stn CP55). Entalinopsis stellata Scarabino, 1995.Indo-Pacific: Indonesia, Philippines, New Caledonia.195-900 m.stenoschizum, Dentalium (Fustiaria) Pilsbry & Sharp,1897: 128, 129, pl. 19, figs 10-15. <strong>Recent</strong>. WestIndies. Syntypes ANSP 71081 (1), ANSP 320994(1). Replacement name for Dentalium translucidumsensu Sowerby (1860, 1872) non Deshayes, 1825.Fustiaria stenoschiza (Pilsbry & Sharp, 1897).Western Atlantic, Caribbean: Florida to Brazil, 40-400 m. Remarks: Scarabino (1995: 277) determinedthat the specimens identified as Dentaliumstenoschizum by Boissevain (1906: 59) were actuallyreferable to Fustiaria nipponica. Since Lamprell &Healy (1998: 100, 102) cited to Boissevain (1906)in their use <strong>of</strong> Fustiaria stenoschiza, their Australianrecords should also be attributed to F. nipponica,thus restricting the distribution <strong>of</strong> Fustiariastenoschiza to the western Atlantic <strong>and</strong> theCaribbean.† stentor, Dentalium Anderson & Hanna, 1925: 145,pl. 13, fig. 17. Eocene, Tejon Group. CAS locality792, west side <strong>of</strong> Tecuya Creek, Kern County,California, USA. Type material CAS 819.stimpsoni, Dentalium (Antalis) entale Henderson,1920: 35-38, pl. 4, figs 2-4. <strong>Recent</strong>. Near GottsIsl<strong>and</strong>, <strong>of</strong>f Mount Desert, Maine, Eolis, 20 fms[37 m]. Neotype USNM 314938 (designated byHenderson 1920). Replacement name forDentalium striolatum Stimpson 1851 non Dentaliumstriolatum Risso, 1826. Junior synonym <strong>of</strong>Antalis entalis (Linnaeus, 1758) (fide Pilsbry &Sharp 1897: 43). Remarks: according to Johnson(1976: 66), Stimpson’s type material was destroyedin the great Chicago fire. Henderson, when replacingStimpson’s name, chose a recently collectedspecimen from the USNM collection as the “type”.This specimen is the neotype, although Hendersondid not use that term.† stonecityensis, Cadulus (Gadila) Hodgkinson, 1974:18, text-fig. 8r, s; pl. 1, fig. 17. Middle Eocene,Claibornian Stage, Cook Mountain Formation,Wheelock Member. Stone City Bluff, Brazos RiverValley, Burleson County, Texas, USA. HolotypeUSNM Paleobiology 180441; paratypes PRI29226, ANSP 31504.† stramineum, Dentalium Gabb, 1864: 139, 140,pl. 21, fig. 100. Cretaceous. NE <strong>of</strong> Martinez; SanDiego; <strong>and</strong> Curry’s, S <strong>of</strong> Mount Diablo, California,USA.strangulatum, Dentalium Deshayes, 1825. Not ascaphopod. See Other taxa described as <strong>Scaphopoda</strong>.strangulatus, Cadulus Locard, 1897a: 4. <strong>Recent</strong>. Bay<strong>of</strong> Biscay, Travailleur stn 44 [depth <strong>of</strong> 667 m providedin Locard 1898: 137]. Type material notlocated. Gadila strangulata (Locard, 1897) (fidePilsbry & Sharp 1898: 176). Eastern Atlantic,Mediterranean. 555-2018 m.“strangulosum”, Dentalium “Deshayes” – Gümbel1861. Not a scaphopod. See Other taxa described as<strong>Scaphopoda</strong>.† strenuum, Dentalium Casey, 1903: 266. Oligocene.Vicksburg, Mississippi, USA. Syntype ANSP 998.† striatellulata, Dentalium (Fustiaria) jani var. Sacco,1897: 113, pl. 10, figs 28-33 [as a nomen nudum inSacco 1896: 98]. Miocene/Pliocene. Italy (variouslocalities). Syntypes MIGT BS.106.06.005-BS.106.06.008 (fide Ferrero Mortara et al. 1984:309).striatinum, Siphonodentalium (Siphonodentalium)Henderson, 1920: 90, 91, pl. 16, figs 3, 8. <strong>Recent</strong>.Off Fern<strong>and</strong>ina, Florida, Albatross stn 2668,294 fms [537 m]. Holotype <strong>and</strong> 9 paratypesUSNM 108166. Striopulsellum striatinum(Henderson, 1920) (fide Scarabino 1995: 323).Known from the type locality only. Remark:Henderson (1920: 90, 91) referred to a type specimenfrom a lot consisting <strong>of</strong> 10 specimens; however,the holotype has not yet been segregated fromthe paratypes.striatissimum, Dentalium Doderlein, 1864: 97 [15].Nomen nudum. Miocene. Monte Gibio (Modena)<strong>and</strong> Santo Agata (Tortona), Italy. Remark: Cocconi(1874: 647) mentioned this manuscript name in hisdescription <strong>of</strong> Dentalium passerinianum.† striatissimum, Dentalium Razzore, 1896: 248 [exDoderlein MS]. Pliocene. Borzoli, Zinola <strong>and</strong> RioTorsero, Italy. Lectotype MIGT BS.106.01.030(selected by Pavia 1991: 113); paralectotypeBS.106.01.031 (fide Ferrero Mortara et al. 1984:302). Remarks: Dentalium striatissimum Vinassa deRegny, 1897 is a junior homonym. This name hasbeen attributed to Sacco, 1897 (e.g., Pavia 1991:112-114) but Razzore had already validatedDoderlein’s manuscript name. Dentalium(Dentalium) passerinanum subsp. striatissimumRazzore, 1896 (fide Pavia 1991: 112-114, pl. 1,figs 4, 6, 8-10).striatissimum, Dentalium Vinassa de Regny, 1897: 163[ex Meneghini MS]. Tertiary. Roncà, Italy. JuniorZOOSYSTEMA • 2004 • 26 (4)647


Steiner G. & Kabat A. R.homonym <strong>of</strong> Dentalium striatissimum Razzore,1896.striatissimum, Dentalium Fuenzalida, 1942: 414, 423.Tertiary. Patagonia, Chile. Nomen nudum. Remarks:Feruglio (1949b: 160) cited this name, but did notvalidate it. If this name ever was validated, then itwould be a junior homonym <strong>of</strong> Dentalium striatissimumRazzore, 1896 <strong>and</strong> Vinassa de Regny, 1897.† striatulinum, Antale “?” Cossmann in O’Gorman &Cossmann, 1923: 28, pl. 3, fig. 4. Eocene. Gan,S <strong>of</strong> Pau, southwestern France.striatulum, Dentalium Gmelin, 1791: 3738. [citingLister 1685-1692: t. 547. f.I.b; Martini 1769: I. t. I.f. 5. b]. <strong>Recent</strong>. Sicily [in error]. Type material notlocated. Junior synonym <strong>of</strong> Dentalium aprinumLinnaeus, 1767 (fide Pilsbry & Sharp 1897: 3).Objective junior synonym: Dentalium lupinumRöding, 1798.striatulum, Dentalium Turton, 1819: 352, pl. 16,figs 5, 6. <strong>Recent</strong>. Type locality not known. Noinformation on type material. Junior synonym <strong>of</strong>Dentalium octangulatum Donovan, 1803 (fidePilsbry & Sharp 1897: 17).striatulus, Solenoxiphus Chistikov, 1983: 187, 189, pl.3, figs 2-13, l pl. 5, figs 10, 11. <strong>Recent</strong>. North FijiBasin, Vitjaz stn 5944-2D, 14°21.1’S, 170°38.2’W,2380 m. Holotype IOM; paratypes IOM (3 fromstn 59442D -2D; 1 from stn 6818-2, 5°05.8’N,130°58.8’E, 1530-1680 m). Solenoxiphus striatulusChistikov, 1983. Indo-Pacific: South Philippine Seato Fiji Isl<strong>and</strong>s <strong>and</strong> New Caledonia. 1500-3700 m.striatum, Dentalium Born, 1778: 449, 450 [citingGualtieri 1742: pl. 10, fig. 6; Hill 1752, pl. 7;Martini 1769: pl. 5, fig. B]; 1780: 431. <strong>Recent</strong>.Type locality not specified. Syntypes NHM. Juniorsynonym <strong>of</strong> Dentalium elephantinum Linnaeus,1758 (partim) <strong>and</strong> Dentalium aprinum Linnaeus,1767 (partim) (fide Pilsbry & Sharp 1897: 2, 3).striatum, Dentalium Link, 1807: 23 [citing Schröter1784b: 4 t. 3 f. 5]. Tertiary. Sternberg, Germany.Junior homonym <strong>of</strong> Dentalium striatum Born,1778. Objective junior synonym <strong>of</strong> Dentalium rectumGmelin, 1791; both taxa are based upon thesame illustration.striatum, Dentalium J. Sowerby, 1814: 160, pl. 70,fig. 4. Upper Eocene. Barton, Hampshire, Engl<strong>and</strong>,United Kingdom. Junior homonym <strong>of</strong> Dentaliumstriatum Born, 1778. Replacement name:Dentalium acuticosta Deshayes, 1825; Dentaliumbartonense Palmer, 1974 is an unnecessary replacementname. Remarks: Deshayes (1861: 206) attributedthis name to “Br<strong>and</strong>er, 1775, Foss. hant., pl. 1,fig. 10”. Pilsbry & Sharp (1898: 198) noted thatthe 1766 edition <strong>of</strong> Br<strong>and</strong>er used Dentalium elephantinumfor this <strong>species</strong>, <strong>and</strong> wondered whether“Deshayes may quote from another edition whichwe have not seen”. We have been unable to trackany reference to a “1775” edition <strong>of</strong> Br<strong>and</strong>er <strong>and</strong>conclude that Deshayes’ attribution was in error.striatum, Dentalium Lamarck, 1818: 344. Tertiary.Siena, Italy. Junior homonym <strong>of</strong> Dentalium striatumBorn, 1780, D. striatum J. Sowerby, 1814.Remark: Bellini (1909: 228) concluded thatDentalium delessertianum Chenu, 1843 was the nextavailable name for Dentalium striatum Lamarck,1818, which however is a junior synonym <strong>of</strong>Fissidentalium rectum (Gmelin, 1791) (fide Pilsbry& Sharp 1898: 213; fide Caprotti 1979: 223).striatum, Dentalium Brown, 1827: caption to pl. 1,fig. 8. <strong>Recent</strong>. Eastern Atlantic: Holy Isl<strong>and</strong>, coast<strong>of</strong> Northumberl<strong>and</strong>, Engl<strong>and</strong>, United Kingdom.Junior homonym <strong>of</strong> Dentalium striatum Born,1780, D. striatum J. Sowerby, 1814, <strong>and</strong> D. striatumLamarck, 1818.striatum, Dentalium Eichwald, 1830: 199. Pliocene.Zukowce, Volhynia, Ukraine. Junior homonym <strong>of</strong>Dentalium striatum Born, 1780, D. striatumJ. Sowerby, 1814, D. striatum Lamarck, 1818, <strong>and</strong>D. striatum Brown, 1827. Replacement name:Dentalium quindeciesstriatum Eichwald, 1853.striatum, Dentalium sensu Philippi 1844: 208 nonBorn, 1778. Tertiary. Sicily, Italy. Replacementname: Dentalium philippi Monterosato, 1872, ajunior homonym <strong>of</strong> Dentalium philippi Chenu,1843; renamed Dentalium rectum var. monterosatoiPilsbry & Sharp, 1898: 214.striatum, Dentalium Ivanov, 1926: 174. Nomennudum. Carboniferous. Near Moscow, Russia. NonDentalium striatum Born, 1780 (<strong>Recent</strong>). Replacementname: Dentalium ivanovi Palmer, 1974.Remark: as Ivanov’s taxon was not validlydescribed, there was no need to rename it.striatus, Cadulus (Gadila) Pilsbry & Sharp, 1898 [exDall MS]: 179, 180, pl. 35, figs 9-13. <strong>Recent</strong>. Gulf<strong>of</strong> Panama, USFC stn 3354, 7°09’45”N, 80°50’W,322 fms [588 m]. Syntypes USNM 122992 (1),USNM 602252 (15), SBMNH 35023 (2), AMNH148405 (3). Gadila striata (Pilsbry & Sharp,1898). Eastern Pacific: Panama. 588-1206 m.striatus, Dentalites Schlotheim, 1813: 91. S<strong>and</strong>stein,Mecklenberg, Germany. Nomen nudum.† striatus, Entalis Romanovskii, 1890: 112, pl. 16,fig. 4. Upper Paleozoic. Pasha-Ata <strong>and</strong> Namangan,eastern Uzbekistan [“Turkestan”]. Prodentaliumstriatus (Romanovskii, 1890) (fide Likharev 1967:12, 13, pl. 1, figs 1-3).strigatum, Dentalium Gould, 1859: 166. <strong>Recent</strong>. FalseBay, Cape <strong>of</strong> Good Hope. Lectotype USNM 159(designated by Johnson 1964: 153); paralectotypesMCZ 169382 (2). Junior synonym: Dentaliumagulhasense Plate, 1908 (fide Barnard 1974: 742).Dentalium strigatum Gould, 1859. Indian Ocean:South Africa, Madagascar, Réunion. 33-317 m.Remark: Tomlin (1931: 337) considered Dentaliumbelcheri Sowerby, 1860 to be a junior synonym <strong>of</strong>Dentalium strigatum.† striolatissima, Dentalium sexangulum var. Sacco,1897: 94, pl. 7, figs 55, 56 [as a nomen nudum in648ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Sacco 1896: 97]. Pliocene. Italy (various localities).Syntypes MIGT BS.106.01.008, BS.106.01.009(fide Ferrero Mortara et al. 1984: 301).striolatum, Dentalium Risso, 1826b: 398. <strong>Recent</strong>.Mediterranean. Syntypes MNHN (8). Junior synonym<strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fidePilsbry & Sharp 1897: 42). Remark: one <strong>of</strong> the syntypesis actually a specimen <strong>of</strong> Antalis inaequicostata(Dautzenberg, 1891) (fide Arnaud 1978: 135).striolatum, Dentalium Stimpson, 1851a: 114. <strong>Recent</strong>.At the mouth <strong>of</strong> the Bay <strong>of</strong> Fundy, 10-60 fms [18-110 m]. Type material lost in the Chicago fire(Johnson: 1976: 66). Neotype USNM 314938(designated by Henderson 1920: 35, pl. 4, figs 2-4).Near Gotts Isl<strong>and</strong>, <strong>of</strong>f Mount Desert, Maine, Eolis,20 fms [37 m]). Junior homonym <strong>of</strong> Dentaliumstriolatum Risso, 1826. Replacement name:Dentalium (Antalis) entale stimpsoni Henderson,1920, a junior synonym <strong>of</strong> Antalis entalis(Linnaeus, 1758) (fide Pilsbry & Sharp 1897: 43).† stromeri, Dentalium (Fustiaria) Oppenheim, 1906:215, pl. 25, figs 6, 6a. Miocene. Fajûn, west <strong>of</strong> KasrKerûn, Libya.stumkatae, Dentalium (Lentigodentalium) Lamprell &Healy, 1998: 70, 71, figs 68B, 69C, 71. <strong>Recent</strong>.32 km <strong>of</strong>f Point Charles, NW <strong>of</strong> Darwin, 12°10’S,130°22’E, 27-37 m. Holotype AMS C174641;paratypes AMS C174642 (2), AMS C169964(2; 96 km NE <strong>of</strong> Port Hedl<strong>and</strong>, Western Australia,19°30.9-28.2’S, 118°49.2-55.4’E, 36-37 m).Dentalium stumkatae Lamprell & Healy, 1998.Indo-Pacific: northern Australia. 37-154 m.subagile, Dentalium (Antalis) agile Henderson, 1920:38, 39, pl. 4, fig. 8. <strong>Recent</strong>. USBF stn 2513[between Halifax <strong>and</strong> Le Havre Bank], 43°34’N,63°56’30”W, 134 fms [245 m]. Holotype USNM887470; paratypes USNM 50188, USNM 50189,USNM 50191, USNM 50193, USNM 52302,USNM 314325, AMNH 148306 (1), ANSP119158. Sub<strong>species</strong> <strong>of</strong> Antalis agilis (G. O. Sars exM. Sars, 1872). Remarks: Abbott (1974: 385) listedsubagile as a valid <strong>species</strong>. This opinion is not followedherein because the observed high variability<strong>of</strong> Antalis agilis across the northern Atlantic encompassesthe form <strong>of</strong> subagile.subanceps, Dentalium Trautschold, 1861: 350-352,pl. 8, figs 16, 17. Jurassic. Mniowniki, Galiowa <strong>and</strong>Dorogomil<strong>of</strong>, near Moscow, Russia. Junior synonym<strong>of</strong> Dentalium gladiolus Eichwald, 1846 (fideEichwald 1868: 799; Pilsbry & Sharp 1898: 223).† subannulata, Plagioglypta Easton, 1962: 101, pl. 13,figs 20, 21. Carboniferous. USGS locality 13399,Alaska Bench limestone, Heath, Fergus County,Montana, USA. Holotype <strong>and</strong> 9 paratypes (in oneblock) USNM Paleobiology 118878.† subarcuatum, Dentalium Conrad, 1853: 276, pl. 24,fig. 13. Cretaceous. New Egypt, New Jersey, USA.† subcanaliculatum, Dentalium S<strong>and</strong>berger &S<strong>and</strong>berger, 1856: 240, pl. 26, fig. 19, 19a.Carboniferous. Villmar (“Stringocephalenkalk”),Germany. Remarks: Nomen nudum in S<strong>and</strong>berger(1842: 399). Dentalium cubcanaliculatum“G. S<strong>and</strong>berger, 1842” [de Koninck, 1843: 319] isan incorrect subsequent spelling.“subcarinatum”, Dentalium “Münster in Goldfuss” –Ryckholt 1851. Not a scaphopod. See Other taxadescribed as <strong>Scaphopoda</strong>.subcentralis, Costentalina tuscarorae Chistikov, 1982b:1317, 1318, pl. 1, figs 9-11, pl. 4, fig. 3. <strong>Recent</strong>.Vitjaz stn 4199-T [4191 in error], 35°06.8’N,137°53’W, 5056-5072 m. Holotype IOM;paratypes IOM (2 from stn 4199-T; 1 from stn4074-T, 40°19’N, 175°45.3’W, 6065 m; 1 fromstn 4239-T, 24°50.3’N, 144°05.4’W, 5120 m;2 from stn 4281-T, 20°01’N, 121°59.6’W,4370 m; 1 from stn 4289-T, 20°00.3’N,130°01.2’W, 4984-4902 m; 4 from stn 5065-T,16°23’N, 146°36’W, 5363-5570 m; 1 from stn5996-2-T, 10°58.2’N, 153°21.5’W, 5060-5080 m;2 from stn 6298-24-T, 22°44’S, 160°54’W,4860 m; 1 from stn 6704, 18°36.8’N, 128°07.8’E).Sub<strong>species</strong> <strong>of</strong> Costentalina tuscarorae Chistikov,1982. Indo-Pacific <strong>and</strong> Eastern Pacific: fromPhilippines to <strong>of</strong>f California <strong>and</strong> Mexico. 4370-6065 m.subcoarctata, Ditrupa (Gabb, 1860) – Dall 1892: 444.Unjustified emendation <strong>of</strong> Dentalium subcoarcuataGabb, 1860.† subcoarcuata, Ditrupa Gabb, 1860: 386, pl. 67,fig. 47. Eocene. Wheelock, Texas, USA. HolotypeANSP 13263 (“possibly lost” fide Hodgkinson1974: 20). Unjustified emendation: Cadulus subcoarctatus(Gabb, 1860) (Dall 1892: 444). Cadulus(Polyschides) subcoarcuatus (Gabb, 1860) (fideHodgkinson 1974: 19). Remark: Aldrich (1895: 4)<strong>and</strong> Pilsbry & Sharp (1898: 235) listed Cadulusabruptus Meyer & Aldrich, 1886 as a probable juniorsynonym <strong>of</strong> Cadulus subcoarcuatus.subcolubridens, Cadulus (Gadila) Ludbrook, 1954:115, 116, fig. 21. <strong>Recent</strong>. Gulf <strong>of</strong> Aden, 2000 m,John Murray Exp. stn 185, 13°48’N, 49°16’E.Holotype BMNH 1952.3.25.32; paratypes BMNH1952.3.25.33-BMNH 1952.3.25.44 (12). Gadilasubcolubridens (Ludbrook, 1954). Indo-Pacific:Red Sea to New Zeal<strong>and</strong>. 1158-2000 m.† subcompressum, Dentalium Meyer, 1885: 462; 1886:64, pl. 3, figs 3, 3a. Eocene. Vicksburg <strong>and</strong> Jackson,Mississippi, USA. Syntype USNM Paleobiology638701 (as “holotype” in Palmer & Brann 1965:371). Fustiaria (Laevidentalium) subcompressa(Meyer, 1885) (fide Palmer & Brann 1965: 371).subcostulata, Dentalium agile var. Locard, 1898: 118.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Antalis agilis (G. O. Sars ex M. Sars, 1872).subcurvatum, Dentalium E. A. Smith, 1906[October]: 251. <strong>Recent</strong>. SW <strong>of</strong> Cape Comorin,[Maldives], India, Investigator stn 275, 731-771 fmsZOOSYSTEMA • 2004 • 26 (4)649


Steiner G. & Kabat A. R.[1334-1407 m]. The holotype in BMNH as mentionedby Scarabino (1995: 261) could not belocated. Junior synonym: Dentalium martensiBoissevain, 1906 [December] (fide Scarabino 1995:261). Compressidentalium subcurvatum (E. A.Smith, 1906) (fide Scarabino 1995: 261). Indo-Pacific: Madagascar to New Caledonia. 142-1411 m.subcylindrica, Cadulus tumidosus var. Locard, 1898:143. <strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Cadulus tumidosus Jeffreys, 1877.† subcylindricum, Dentalium Philippi, 1887a: 105,pl. 12, fig 14; 1887b: 99, pl. 12, fig. 14.Cretaceous. Algarrobo <strong>and</strong> S. Vicente, Chile.† subeburneum, Dentalium d’Orbigny, 1850(volume 2): 372. Eocene. Chaumont, Halaincourt,Acy-en-Mulcien (Oise), France. Replacement namefor Dentalium eburneum sensu G. B. Sowerby 1825non Linnaeus, 1767. Junior synonym: Fustiaria circinataNewton, 1891 (fide Pilsbry & Sharp 1898:216). Remarks: Deshayes (1864: 216) listed thisname as a junior synonym <strong>of</strong> Dentalium eburneumLinnaeus, 1767; Newton & Harris (1894: 65, 66),Le Renard (1995: 175), <strong>and</strong> Le Renard & Pacaud(1995: 85) considered it as a valid <strong>species</strong> <strong>of</strong>Fustiaria.subentalis, Dentalium d’Orbigny, 1850 (volume 2):320. Eocene, Suessonian. Cuise-Lamotte, Varinfroy(Oise), France. Tentative junior synonym <strong>of</strong> Antalisentalis (Linnaeus, 1758) (fide Pilsbry & Sharp 1898:216).† subfissura, Entalis Tate, 1887: 191, pl. 20, figs 4a,4b. Eocene. River Murray Cliffs, near Morgan;Nine-Mile Camp, near Nor’West Bend; MuddyCreek, Corio Bay; Schnapper Point, Victoria;Blanche Point, Aldinga Bay, Australia. Holotype<strong>and</strong> 15 paratypes SAM T 249A. Dentalium(Laevidentalium) subfissura (Tate, 1887) (fideLudbrook 1959: 146, 147, pl. 2, fig. 3).subfusiforme, Siphonodentalium M. Sars, 1865a: 301,pls 6, 7, figs 36-44. <strong>Recent</strong>. Drøbach, Langesund<strong>and</strong> Flekkefjord, 50-120 fms; Bergen, 50-60 fms;L<strong>of</strong>oten, 100-300 fms [west coast <strong>of</strong> Norway, 91-548 m]. Syntypes SMNH 4626 (Norway, L<strong>of</strong>oten,360-540 m). Cadulus subfusiformis (M. Sars, 1865)(fide Jeffreys 1867: 251). Eastern Atlantic:Northern Europe to Northern Africa. 73-1185 m.† subgiganteum, Dentalium d’Orbigny, 1852 (volume3): 94. Miocene, “Falunian”. Navidad, Chile.Replacement name for Dentalium giganteum J. deC. Sowerby, 1846 non Phillips, 1829. Junior synonym:Dentalium corrugatum Hupé in Gay, 1854(fide Pilsbry & Sharp 1898: 217).† subgracile, Dentalium Emerson, 1954: 185.Paleocene. Jutl<strong>and</strong>, Denmark. Replacement namefor Dentalium gracile Grönwall & Harder, 1907non Dentalium gracile Hall & Meek, 1854, necMoore, 1866, nec Jeffreys, 1870, nec Philippi, 1887.† subirregulare, Dentalium Pilsbry & Sharp, 1898:217. Pliocene, Astian stage. Gallina, Reggio,Calabria, Italy. Replacement name for Dentaliumirregulare Seguenza, 1879 non Dentalium irregularisRisso, 1826 nec Hutton, 1873.† subjuvenis, Dentalium (Entalis) taurostriata var.Sacco, 1897: 109, pl. 9, fig. 55 [as a nomen nudumin Sacco 1896: 97]. Miocene. Turin hills, Italy.Syntype MIGT BS.106.03.030 (fide FerreroMortara et al. 1984: 308).sublaeve, Dentalium Hall in Miller, 1877: 244.Carboniferous. Iowa, USA. Objective junior synonym<strong>of</strong> Prodentalium obsoletum (Hall, 1858).Remarks: introduced as a replacement name forDentalium obsoletum Hall, 1858 non Schlotheim,1832. However, Schlotheim’s name is a nomennudum. Dentalium sublaeve is also a junior homonym<strong>of</strong> Dentalium dentalis var. sublaevis Cocconi, 1874.sublaevigata, Dentalium semivestitum var. Locard,1898: 108. <strong>Recent</strong>. Type locality not specified[North Atlantic]. Type material not located.Synonym <strong>of</strong> Fissidentalium semivestitum (Fischerin Locard, 1898).† sublaevis, Dentalium dentalis var. Cocconi, 1874:648. Miocene. Vigoleno, Italy. Said by Cocconi tobe the same as Dentalium obsoletum Doderlein,1862 [nomen nudum].sublaevis, Dentalium sexangulum var. Coppi, 1881: 85.Pliocene. Savignano <strong>and</strong> Zappolino, Italy. Nomennudum.† sublineatum, Dentalium Stephenson, 1952: 142,143, pl. 34, fig. 3. Upper Cretaceous, Cenomanian,Woodbine Formation. Lamar County Locality 201,near old Slate Shoals, Red River, 8 miles E <strong>of</strong>Arthur City, Lamar County, Texas, USA. HolotypeUSNM Paleobiology 105607; paratypes USNMPaleobiology 105608-105609.subornata, Dentalium semivestitum var. Locard, 1898:109. <strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Fissidentalium semivestitum (Fischer in Locard,1898).† subovatum, Dentalium Richardson, 1906: 587, 588,pl. 45, fig. 7. Jurassic, “Lower Lias”. Redcar,Yorkshire, Engl<strong>and</strong>, United Kingdom. For Dentaliumcompressum sensu Terquem 1885 non d’Orbigny,1850. Progadilina subovata (Richardson,1906) (fide Engeser & Riedel 1992: 38).† subparvulum, Dentalium Emerson, 1954: 185.Jurassic, Pliensbachian. Leckhampton Station claypit,Cheltenham, Engl<strong>and</strong>, United Kingdom.Replacement name for Dentalium parvulumRichardson, 1906 non Philippi, 1887. Laevidentaliumsubparvulum (Emerson, 1954) (fideEngeser & Riedel 1992: 38).† subplanum, Dentalium Pilsbry & Sharp, 1898: 228.Cretaceous. Blackdown, Engl<strong>and</strong>, United Kingdom.Replacement name for Dentalium cylindricumsensu Gardner 1878 non J. Sowerby, 1814.650ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)† subpolitum, Siphonodentalium (Dischides) Cossmann& Peyrot, 1917: 185, 186, pl. 2, figs 8-10. Miocene.Giraudeau, Lagus <strong>and</strong> Pont-Pourquey,France. Replacement name for Gadus politus Benoist,1873 non Cadulus politus (S. Wood, 1842).† subprismaticum, Dentalium (Dentalium) Baluk,1972: 450, 451, pl. 6, figs 2-7, text-fig. 2. Miocene,Lower Tortonian (Badenian). Korytnica, 24 kmSSW <strong>of</strong> Kielce, Holy Cross Mountains, Pol<strong>and</strong>.Holotype <strong>and</strong> paratypes, Baluk collection.† subquadrangulare, Dentalium Emerson, 1954: 185.Mesozoic. Morea, Peloponnesus, Greece. Replacementname for Dentalium quadrangulare Deshayes1835 non Sowerby in Broderip & Sowerby, 1832.† subquadratum, Dentalium Meek, 1860: 311.Jurassic. North Platte River, at Red Buttes,42°50’N, 106°40’W, Wyoming, USA. SyntypesUSNM Paleobiology 677 (slab with numerousspecimens).subquadratum, Dentalium Richardson, 1906: 588,589, pl. 45, fig. 8. Jurassic, Sinemurian. Honeybourne,near Evesham, Engl<strong>and</strong>, United Kingdom.Junior homonym <strong>of</strong> Dentalium subquadratumMeek, 1860. The replacement name, Dentaliumrichardsoni Emerson, 1954, is itself a junior homonym<strong>of</strong> Dentalium richardsoni Cossmann, 1907.Replacement name: Dentalium sabrinae Engeser &Riedel, 1992.subrecta, Dentalium agile var. Locard, 1898: 117.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Antalis agilis (G. O. Sars ex M. Sars, 1872).subrecta, Dentalium scamnatum var. Locard, 1898:110. <strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Dentalium scamnatum, a junior synonym <strong>of</strong>Fissidentalium capillosum (Jeffreys, 1877).† subrecta, Dentalium sexangulare var. Cocconi, 1874:645. Pliocene. Castell’Arquato, Italy.subrectum, Dentalium Jeffreys, 1883: 661. <strong>Recent</strong>.Philippines. Holotype USNM 176124. Juniorhomonym <strong>of</strong> Dentalium sexangulare var. subrectaCocconi, 1874. Episiphon virgula (Hedley, 1903)(fide Scarabino 1995: 287). Remarks: since Habe(1964: 29) claimed that E. subrectum was a juniorhomonym <strong>of</strong> the <strong>fossil</strong> taxon Dentalium sexangulumvar. subrecta Cocconi, 1873, Hedley’s (1903) nameis the oldest available one for this <strong>species</strong>, <strong>and</strong> notEpisiphon subrectum var. makiyamai (Kuroda &Kikuchi, 1933). V. Scarabino (in litt. 5.XI.2001),based on examination <strong>of</strong> the type material, suggestedthat the status <strong>of</strong> D. subrectum vis-à-visLaevidentalium eburneum (Linnaeus, 1767) requiresfurther research.subrectum, Dentalium Martin, 1885: 185, pl. 10,figs 180, 181; 1887: 185, pl. 10, figs 180, 181.Miocene. Ngembak <strong>and</strong> Grissee, Java, Indonesia.Syntypes RGM 7239 (1), RGM 7240 (1), RGM7238 (1), RGM 7241 (1) (fide Van den HoekOstende et al. 2002: 112). Junior homonym <strong>of</strong>Dentalium sexangulum var. subrecta Cocconi, 1873.Dentalium teschi Koperberg, 1931. Remark: thisname introduced as sub<strong>species</strong> by Koperberg(1931) has priority over the replacement nameDentalium semarangense Altena, 1938.† subregulare, Dentalium Palmer, 1974: 124. Triassic.Granietz, Upper Silesia, Pol<strong>and</strong>. Replacement namefor Dentalium regulare Ahlburg, 1906 nonE. A. Smith, 1903.† subserratum, Dentalium Palmer, 1974: 125.Miocene. Grissee, Java, Indonesia. Holotype RGM7270 (fide Van den Hoek Ostende et al. 2002:112). Replacement name for Dentalium serratumMartin, 1885 non Pictet & Roux, 1849(Polychaeta).† subsexangulare, Dentalium d’Orbigny, 1852 (volume3): 94. Miocene, Falunian. Piemont, Dertona,Italy. Replacement name for Dentalium sexangularesensu Deshayes 1825 non Lamarck, 1818. Tentativejunior synonym: Dentalium noe Sismonda, 1847(fide Pilsbry & Sharp 1898: 215). Remarks:d’Orbigny (1852) erroneously credited sexangulareto Gmelin, 1791 instead <strong>of</strong> Lamarck, 1818; Gmelin(1791) is author <strong>of</strong> Dentalium sexangulum.Dentalium subsexangulatum [d’Orbigny, 1852(Index): 59] is an incorrect subsequent spelling.subsexangulatum, see subsexangulare.substriata, Teredo Conrad, 1849: 728, pl. 20, figs 7,7a, 7b. Miocene. Astoria, Oregon, USA. Types notlocated. A junior secondary homonym <strong>of</strong>Dentalium substriatum Deshayes, 1825. Replacement<strong>names</strong>: Dentalium pseudonyma Pilsbry &Sharp, 1898; Dentalium conradi Dall; 1909Dentalium petricola Dall, 1909. Remark: this<strong>species</strong> was independently renamed twice; seeEmerson (1958) for further discussion.† substriatum, Dentalium Deshayes, 1825: 366, 367,pl. 18, figs 1, 2. Eocene. Parnes, Chaumont <strong>and</strong>l’Orme, Paris Basin, France. Types not located. ForDentalium fissura sensu J. Sowerby 1823 nonLamarck, 1818. Remarks: this <strong>species</strong> was misidentifiedas Dentalium acuticosta Deshayes, 1825 byDixon (1850: 96) (fide Deshayes 1864: 208).A valid <strong>species</strong> <strong>of</strong> Antalis (fide Le Renard 1995: 175;Le Renard & Pacaud 1995: 85).subterfissum, Dentalium Jeffreys, 1877: 154, 155.<strong>Recent</strong>. Localities <strong>of</strong> syntypes: USNM 175017(Porcupine 1869 stn 19a), USNM 175018(Porcupine 1869 stn 16, 54°19’N, 11°50’W,816 fms [1489 m]), USNM 175019 (Valorousstn 12, 56°11’N, 37°41’W, 1450 fms [Greenl<strong>and</strong>,2646 m]), USNM 175020 (Porcupine 1869 stn 21,55°40’N, 12°46’W, 1476 fms [2694 m]); BMNH85.11.5.1393-1394 (Porcupine, without locality).Heteroschismoides subterfissus (Jeffreys, 1877) (fideHenderson 1920: 58, as Dentalium (Heteroschisma)subterfissum). Western Atlantic: Caribbean toBrazil; Eastern Atlantic: Greenl<strong>and</strong> to West Africa.ZOOSYSTEMA • 2004 • 26 (4)651


Steiner G. & Kabat A. R.940-3109 m. Remark: also described fromChallenger Exp. (station number not given),37°26’N, 45°14’W, 1000 fms [1825 m], but thistype lot was neither located in BMNH nor mentionedby Warén (1980).subterlineatum, Dentalium Tomlin, 1931: 337.<strong>Recent</strong>. Off Cape Point [South Africa], 900 fms[1642 m]. Lectotype SAFM 6192 (designated byGiles & Gosliner 1983: 41); paralectotype NMW(1). Entalina subterlineata (Tomlin, 1931) (fideChistikov 1982c: 1497). South Africa. 690-1645 m.subtilis, Cadulus (Gadila) Plate, 1908a: 360, pl. 30,fig. 48. <strong>Recent</strong>. Near Dar-es-Salaam, Tanzania,Valdivia stn 244, 5°55.8’S, 39°1.2’E, 50 m.Lectotype ZMB 61108a (designated by Kilias 1995:176); paralectotypes ZMB 61108b (1), ZMB63842 (4). Gadila subtilis (Plate, 1908) (fideJaeckel 1932: 310). Known from the type localityonly. Remark: Jaeckel (1932: 310) remarked uponcertain similarities with Ditrupa (Polychaeta), <strong>and</strong>Kilias (1995: 176) parenthetically remarked thatthis taxon is probably Ditrupa. Plate’s illustrationindeed resembles a worm tube, <strong>and</strong> the taxon mayhave to be removed from the <strong>Scaphopoda</strong> afterexamination <strong>of</strong> the types.subtorquatum, Dentalium Fischer, 1871: 212, 218,pl. 11, fig. 1. <strong>Recent</strong>. Suez, Red Sea. Lectotype <strong>and</strong>4 paralectotypes MNHN (designated by Scarabino1995: 286). Junior synonyms: Dentalium anulosumBrazier, 1877; Dentalium tornatum Watson, 1879;Dentalium (Episiphon) sewelli Ludbrook, 1954 (fideScarabino 1995: 286). Episiphon subtorquatum(Fischer, 1871) (fide Habe & Kosuge 1964: 8).Indo-Pacific: South Africa to New Caledonia <strong>and</strong>Fiji; Eastern Atlantic: South Africa. 17-650 m.Remark: Lamprell & Healy (1998: 104) listed anulosumas a valid <strong>species</strong> <strong>of</strong> Omniglypta.† subtrigonale, Dentalium Richardson, 1906: 589,pl. 45, fig. 2. Jurassic, Pliensbachian. Folly-Laneclay pit, Cheltenham, Engl<strong>and</strong>, United Kingdom.Progadilina subtrigonalis (Richardson, 1906) (fideEngeser & Riedel 1992: 39).subula, Cadulus (Gadila) Henderson, 1920: 135, 136,pl. 19, fig. 5. <strong>Recent</strong>. Off Bahia Honda, Cuba, USBFstn 2352, 22°35’N, 84°23’W, 463 fms [845 m].Holotype USNM 323113. Gadila subula (Henderson,1920). Known from the type locality only.subulatum, Dentalium Deshayes, 1825. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† sueroi, Calstevenus Sabattini, 1979: 23-25, pl. 1,figs 4-9. Lower Permian, Tepuel Group. Subida delPaisano, Sierra de Languineo, Chubut Province,Argentina. Holotype MLP 13169; paratypes MLP13167, 13168, 13170, 13171.† sulcatulum, Dentalium Koenen, 1892: 987, 988,pl. 59, figs 14, 15. Lower Oligocene. Lattorf,Germany.sulcatum, Dentalium Schumacher, 1817: 263 [citingMartini 1769: I t. I f. 4A]. Type locality not specified.Types not located. Junior synonym <strong>of</strong>Fissidentalium rectum (Gmelin, 1791); both taxa arebased upon the same illustrations from Martini1769.sulcatum, Dentalium Lamarck, 1818: 343. Eocene.Grignon, France. Junior homonym <strong>of</strong> Dentaliumsulcatum Schumacher, 1817; replacement name:Dentalium potiezi Le Renard, 1994.sulcatum, Dentalium (Antalis) occidentale var. Verrill,1884: 217. <strong>Recent</strong>. Georges Bank [Massachusetts].Syntypes USNM 38398 (1; USBF 2079, Albatrossstn 2079, 41°13’N, 66°19’50”W, 75 fms [137 m]),USNM 35093 (4; USBF 2077, 1255 fms [2290 m]).Junior homonym <strong>of</strong> Dentalium sulcatumSchumacher, 1817 <strong>and</strong> Dentalium sulcatumLamarck, 1818. Replacement name: Dentalium(Antalis) occidentale georgiense Henderson, 1920: 43.Synonym <strong>of</strong> Antalis occidentalis (Stimpson, 1851).† sulcosum, Dentalium J. de C. Sowerby, 1846: 263,pl. 2, fig. 2. Tertiary. Navidad, Chile. Junior synonyms:Dentalium majus J. de C. Sowerby, 1846;D. gayi Philippi, 1887 (fide Ortmann 1902: 157,159; by First Reviser action, according to ICZN1999: article 24).sumatrense, Dentalium (Compressidens) Plate, 1908a:350 (illustrated in Jaeckel 1932: 304, text-fig. 3).<strong>Recent</strong>. Sumatra, Valdivia stn 186, 3°22.1’S,101°11.5’E, 903 m. Holotype ZMB 61081 (fideKilias 1995: 176). Compressidentalium sumatrense(Plate, 1908) (fide Scarabino 1995: 264). Indo-Pacific: Indonesia. 750-903 m.summa, Polyschides Okutani, 1964: 79, 80 [ex KurodaMS] (nomen nudum in Habe 1953: 298; 1963:278). <strong>Recent</strong>. Sagami Bay, Japan, 700-780 m; <strong>of</strong>fKushikino, Japan, 730-760 m. Types in NSMT(?).Siphonodentalium summa (Okutani, 1964) (fideHabe 1964: 51). Indo-Pacific: Japan. 700-780 m.Remark: Siphonodentalium magnum (Boissevain,1906) may be a senior synonym <strong>of</strong> this <strong>species</strong>(Habe 1964: 51).† sundkrogensis, Dentalium (Dentalium) Schnetler,2001: 43, 44, pl. 1, figs 10, 11. Sel<strong>and</strong>ian,Paleocene. Sundkrogen, Copenhagen, Denmark.Holotype MGUH 24839; paratype MGUH24838. Remark: Schnetler determined that somespecimens identified in the literature as Dentaliumrugiferum von Koenen 1885 were actually a distinct,new <strong>species</strong>.† sunieri, Dentalium (Antalis) Koperberg, 1931: 31,32, pl. 1, fig. 4. Neogene/Quaternary. Locality 5,“Toi Oesapi Soka aan de [on the] Noi Lioe(gemengd met fossielen von Nono Fatoe Fekoe),”[mixed with <strong>fossil</strong>s from Nono Fatoe Fekoe] Timor.† superbus, Dentalium Brunet, 1995: 52, 53, pl. 5,fig. 6. Upper Miocene, Entrerriense Formation.Baliza Punta Flecha, Puerto Madryn, ChubutProvince, Argentina. Holotype MPEF-PI-079.suteri, Dentalium Emerson, 1954: 185. <strong>Recent</strong>. PortPegasus, Stewart Isl<strong>and</strong>, New Zeal<strong>and</strong>, 33 m.652ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Holotype ( <strong>of</strong> Dentalium arenarium Suter) NZIGNTM1219 (fide Boreham 1959: 75); paratypes SAMD 16001 (fide Zeidler & Macphail 1978: 83), ZMB61911 (fide Kilias 1995: 172). Replacement namefor Dentalium arenarium Suter, 1913 non Römer,1855. Antalis suteri (Emerson, 1954) (fide Dell1957: 573). Pacific Ocean: Australia, New Zeal<strong>and</strong>.33-374 m.sutherl<strong>and</strong>i, Polyschides Lamprell & Healy, 1998: 169,170, figs 174D, 175F, 179. <strong>Recent</strong>. Off Cairns,northern Queensl<strong>and</strong>, Australia, 17°22’S,146°48’E, 296-303 m. Holotype QM MO53677;paratypes QM MO45834 (1), QM MO40085(11), AMS C172620 (1; E <strong>of</strong> North West Isl<strong>and</strong>,Queensl<strong>and</strong>, 23°15.2’S, 152°24.1’E, 284 m), AMSC173955 (4; mouth <strong>of</strong> Caiman Creek, BerkeleyBay, Port Essington, Cobourg Peninsula, NorthernTerritory [Australia]). Polyschides sutherl<strong>and</strong>iLamprell & Healy, 1998. Indo-Pacific: northern<strong>and</strong> eastern Australia. 243-1764 m.† suzukii, Dentalium (Fustiaria) Yokoyama, 1929: 14,15, pl. 7, figs 6, 6a. Pliocene. Tonohama, nearYasuda, Aki County, Tosa Prefecture, Japan.Holotype Geol. Survey <strong>of</strong> Japan (fide Hanzawa et al.1961: 209).† syltense, Fissidentalium floratum subsp. R. Janssen,1989: 84, 85, pl. 1, figs 7-9. Upper Miocene,Syltian. Morsumkliff, Sylt, Schleswig-Holstein,Germany. Holotype SMF 308309; paratypes SMF308310, SMF 308311/20, RGM 220270/32.† sylvaerupis, Serpulorbis Harris, 1899: 73, pl. 10,fig. 1. Lower Eocene. Woods Bluff, TombigbeeRiver, Clark County, Alabama, USA. Holotype PRI268 (fide Brann & Kent 1960: 793). Remark: transferredfrom the Polychaeta to the <strong>Scaphopoda</strong>:Dentaliidae by Palmer & Brann (1965: 369).syriacum, Dentalium Fraas, 1867: 95 [attributed to“Conrad”]. Cretaceous. Syria. Junior synonym <strong>of</strong>Dentalium cretaceum Conrad, 1852 (fide Pilsbry &Sharp 1898: 221; Blanckenhorn 1927: 123).† taeniolatum, Dentalium S<strong>and</strong>berger & S<strong>and</strong>berger,1856: 241, pl. 26, fig. 21, 21a. Carboniferous.Nassau, Germany.taiwanum, Dentalium aprinum Kuroda, 1941: 149,pl. 14, fig. 45. <strong>Recent</strong>. Takao, Taiwan. Types inNSMT. Junior synonym <strong>of</strong> Dentalium aprinumLinnaeus, 1767 (fide Scarabino 1995: 198).takakoae, Graptacme Tsuchida & Tachi, 1999: 9-11,figs 1-12 [ex Kira MS]. <strong>Recent</strong>. Ise Bay, MiePrefecture, central Japan. Holotype NSMT Mo71400; paratype NSMT Mo 71401. Graptacmetakakoae Tsuchida & Tachi, 1999. Known fromthe type locality only. Remark: this name was firstused in the “Yume-Hamaguri” which was a h<strong>and</strong>writtenJapanese malacological journal. Because itwas h<strong>and</strong>-written, it was not a valid publication(ICZN 1999: article 9.1). In 1994, severalunnamed members <strong>of</strong> the Tokai Shell Clubcompiled various articles from “Yume-Hamaguri”<strong>and</strong> other sources, <strong>and</strong> republished them in a typesetformat (Anonymous 1994: 10, pl. 3, fig. 7 [exKira MS]). Although properly printed, this 1994book was anonymously published, <strong>and</strong> hence is nota valid publication (ICZN 1999: articles 14, 50.1).taphrium, Dentalium Dall, 1889: 422. <strong>Recent</strong>.Between Mississippi Delta <strong>and</strong> Cedar Keys, USBFstn 2405, 28°45’N, 85°02’W, 30 fms [55 m].Lectotype USNM 95320 (designated by Henderson1920: 53); paralectotypes USNM 92891, USNM92819 (2; USFC stn 2598), USNM 93119(2; USFC stn 2608), USNM 93120 (14; USFC stn2612), MCZ 7690 (8; USFC stn 2612), USFC,Havana, 182 fms [332 m] not located. Antalistaphria (Dall, 1889) (fide Pilsbry & Sharp 1897:58). Western Atlantic, Caribbean: Cape Hatteras(North Carolina, USA) to Brazil. 15-692 m.tarentinum, Dentalium Lamarck, 1818: 345. <strong>Recent</strong>.Gulf <strong>of</strong> Tarento [Italy]. Possible syntypes MHNG1081/57 (7; fide Y. Finet in litt. 16.X.1996). Juniorsynonym <strong>of</strong> Antalis vulgaris (Da Costa, 1778) (fidePilsbry & Sharp 1897: 41). Remark: Palmer(1987b) argued that this taxon was a distinct <strong>species</strong><strong>and</strong> not a junior synonym <strong>of</strong> Antalis vulgaris.tasmaniensis, Dentalium Tenison-Woods, 1877: 140.<strong>Recent</strong>. Northwest coast <strong>of</strong> Tasmania, Australia.Type material apparently lost (fide Lamprell &Healy 1998: 172, citing E. Turner, TMH, pers.comm.). Cotton & Godfrey (1940: 327) gave measurements<strong>of</strong> the purported holotype, but their figure348 depicts a sub<strong>fossil</strong> specimen from SouthAustralia. Lamprell & Healy (1998: 172) concludedthat this was an unrecognizable <strong>species</strong>.† tatei, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 218. Upper Eocene. Glauconitics<strong>and</strong>s, Adelaide bore, Kent Town, South Australia,Australia. Holotype <strong>and</strong> 6 paratypes SAM T 252A.Replacement name for Dentalium triquetrum Tate,1887 non Brocchi, 1814. Dentalium (Gadilina) tateiSharp & Pilsbry in Pilsbry & Sharp, 1898 (fideLudbrook 1959: 144, pl. 1, fig. 5).† tauraspera, Dentalium (Antale) bouei var. Sacco,1897: 99, pl. 8, figs 14-17 [as a nomen nudum inSacco 1896: 97]. Miocene. Stazzano, Italy.Lectotype MIGT BS.106.02.011 (selected by Pavia,1991: 148); paralectotypes MIGT BS.106.02.008-BS.106.02.010 (fide Ferrero Mortara et al. 1984:303). Fissidentalium tauroasperum (Sacco, 1897)(fide Pavia 1991: 148-150, pl. 5, figs 1, 3, 6-8; table5). Remark: Pavia (1991) emended the <strong>species</strong>name from “tauraspera” to “tauroasperum”.† tauroasperum, Fissidentalium Pavia, 1991: 148-150,pl. 5, figs 1, 3, 6-8. Emendation <strong>of</strong> Dentalium(Antale) bouei var. tauraspera Sacco, 1897.† taurocostatum, Dentalium (Antale “?”) Sacco, 1897:101, pl. 8, figs 50-54 [as a nomen nudum in Sacco1896: 97]. Miocene. Turin hills, Italy. LectotypeMIGT BS.106.02.043 (selected by Pavia 1991:121); paralectotypes MIGT BS.106.02.043,ZOOSYSTEMA • 2004 • 26 (4)653


Steiner G. & Kabat A. R.BS.106.02.044, BS.106.02.046 (fide FerreroMortara et al. 1984: 304, pl. 56, figs 7, 8). Antalistaurocostata (Sacco, 1897) (fide Pavia 1991: 120-123, pl. 3, figs 1-7).† taurogracilis, Dentalium (Antale) bouei var. Sacco,1897: 99, pl. 8, figs 19-21 [as a nomen nudum inSacco 1896: 97]. Miocene. Turin hills, Italy.Syntypes MIGT BS.106.02.013, BS.106.02.014(fide Ferrero Mortara et al. 1984: 303).† taurogracilis, Dentalium (Gadilina) triquetra var.Sacco, 1897: 114, pl. 10, figs 44-46 [as a nomennudum in Sacco 1896: 98]. Miocene. Turin hills,Italy. Syntypes MIGT BS.106.07.013-BS.106.07.010 (fide Ferrero Mortara et al. 1984: 309).Fustiaria taurogracilis (Sacco, 1897) (fide Báldi1973: 339, pl. 51, fig. 4).† taurominima, Cadulus (Loxoporus) Sacco, 1897:116, pl. 10, fig. 78 [as a nomen nudum in Sacco1896: 98]. Miocene. Turin hills, Italy. SyntypeMIGT BS.106.11.001 (fide Ferrero Mortara et al.1984: 310).† taurostriata, Dentalium (Entalis) Sacco, 1897: 109,pl. 9, figs 47-52 [as a nomen nudum in Sacco 1896:97]. Miocene. Turin hills <strong>and</strong> Baldissero, Italy.Syntypes MIGT BS.106.03.026, BS.106.03.027(fide Ferrero Mortara et al. 1984: 307, pl. 56,fig. 4).† taurotumidosus, Cadulus (Cadulus) Sacco, 1897:115, pl. 10, figs 68-73 [as a nomen nudum in Sacco1896: 98]. Miocene. Turin hills, Italy. SyntypesMIGT BS.106.10.008, BS.106.10.009 (fide FerreroMortara et al. 1984: 310, pl. 55, figs 11, 12).† taurovulus, Cadulus (Cadulus) Sacco, 1897: 115,pl. 10, figs 64-67 [as a nomen nudum in Sacco1896: 98]. Miocene. Turin hills, Italy. LectotypeMIGT BS.106.10.006 (selected by Pavia 1991:134); paralectotypes MIGT BS.106.10.005,BS.106.10.007 (fide Ferrero Mortara et al. 1984:310, pl. 55, figs 7, 8).teliger, Cadulus Finlay, 1926: 444, pl. 19, figs 53, 54.Tertiary <strong>of</strong> New Zeal<strong>and</strong>. <strong>Recent</strong> <strong>of</strong>f Auckl<strong>and</strong>Isl<strong>and</strong>, 95 fms [New Zeal<strong>and</strong>, 173 m]. HolotypeAIM; paratypes ANSP 232249, DMNH 22370(fide Mikkelsen & Bradford 1997: 44). Cadulusteliger Finlay, 1926. Pacific Ocean: New Zeal<strong>and</strong>.9-365 m.tenax, Dentalium (Antalis) ceratum Henderson, 1920:52, pl. 7, fig. 3. <strong>Recent</strong>. Barbados, Blake stn 272, 76fms [139 m]. Holotype USNM 887472. Synonym<strong>of</strong> Antalis cerata (Dall, 1881) (n. syn.). WesternAtlantic, Caribbean: Barbados. 60-182 m.† tenue, Dentalium Goldfuss, 1841: 2, pl. 166, fig. 6[ex Münster MS]. Jurassic. Pappenheim, Eichstadt,Bavaria, Germany.tenue, Dentalium Portlock, 1843: 118. Jurassic,Liassic. Aghanloo, Ballymaglin <strong>and</strong> Ballycarton;Magilligan, Craig <strong>and</strong> Gortmore, Irel<strong>and</strong>. HolotypeMuseum <strong>of</strong> Practical Geology, London, No. 5799(fide Richardson 1906: 584) (now in BMNH?).Junior homonym <strong>of</strong> Dentalium tenue Goldfuss,1841. Replacement name: Dentalium portlockiTate, 1870, a junior objective synonym <strong>of</strong> Laevidentaliumminimum (Buckman in Murchison,1845). Remark: Tate (1870b: 17) suggested thatDentalium <strong>and</strong>leri Oppel, 1856 “is probably a synonym”<strong>of</strong> this <strong>species</strong>, “but, assuming that the two<strong>names</strong> refer to the same <strong>species</strong>, the imperfect diagnosis<strong>of</strong> Oppel justifies me in discarding his name”.† tenue, Gadila Ozaki, 1956: 2, 3, pl. 1, fig. 7.Miocene. Nisinotani, Nobori, Hane Town, AkiCounty, Kôti Prefecture, Japan. Holotype NSMT4375.† tenuicincta, Fustiaria Gougerot, 1968: 236. Eocene,Upper Bartonian. Le Ruel, near Marines, Val-d’Oise,France. Remark: a valid <strong>species</strong> <strong>of</strong> Plagioglypta (fideLe Renard 1995: 175; Le Renard & Pacaud 1995:85).† tenuicinctum, Dentalium Koenen, 1892: 983, 984,pl. 59, figs 5, 6. Lower Oligocene. Lattorf <strong>and</strong>Calbe, Germany.† tenuicostatum, Dentalium Böhm, 1891: 69, pl. 2,fig. 34. Cretaceous. Gerhardtstreiter Graben, nearSiegsdorf, southern Bavaria, Germany.tenuicostatum, Fissidentalium Qi & Ma, 1989: 114,115, figs 5a, b. <strong>Recent</strong>. South China Sea, 61-117 m. Type depository not indicated (?IOS).Fissidentalium tenuicostatum Qi & Ma, 1989.Indo-Pacific: China Sea. 61-117 m.tenuifissum, Pseudantalis Monterosato, 1875: 20.<strong>Recent</strong>. Mediterranean. Type material not located.Junior synonym <strong>of</strong> Fustiaria rubescens (Deshayes,1825) (fide Pilsbry & Sharp 1897: 106).† tenuis, Dentalium Hutton, 1873b: 1. Eocene,Otatara Group. Whangape Lake, Waikato, NewZeal<strong>and</strong>; Raglan, New Zeal<strong>and</strong>. Lectotype NZGSTM4806 (designated by Keyes 1972: 95).tenuis, Dentalium novemcostatum var. Monterosato,1878: 76 (non tenuis Hutton, 1873, <strong>fossil</strong>). <strong>Recent</strong>.Nomen nudum.† tenuis, Helonyx Seguenza, 1879: 118, pl. 11, fig. 50.Miocene. Reggio, Calabria, Italy. Cadulus tenuis(Seguenza, 1879) (fide Pilsbry & Sharp 1898: 239).† tenuissimum, Dentalium de Koninck, 1876: 117,118, pl. 4, fig. 3; 1898: 93, 94, pl. 4i fig. 3.Devonian? Yaas, New South Wales, Australia.† tenuistriata, Cyrtoconella Patrulius, 1996: 18, pl. 5,figs 8, 9. Jurassic, Bathonian/Lower Callovian.Crisul Repede, Vadu Crisului, Apuseni Mountains,northwestern Romania. Holotype IGG P-17196;paratypes IGG P-17197 (19). Remark: Patruliustentatively referred his new <strong>species</strong> to the<strong>Scaphopoda</strong>.† tenuistriatum, Dentalium Rouault, 1850: 473,pl. 15, fig. 5. Eocene. Bos d’Arros, near Pau, France.tenuistriatum, Dentalium Martin, 1880: 88, pl. 12,fig.12. Miocene. Locality R, Tji-Karang, E <strong>of</strong>Tijieri, Kendeg district, Sukapura Regency, Java,Indonesia. Syntypes RGM 7272 (5) (fide Van den654ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Hoek Ostende et al. 2002: 112, pl. 3). Junior homonym<strong>of</strong> Dentalium tenuistriatum Rouault, 1850;replacement name: Fissidentalium ortus Palmer,1974.tenuistriatum, Dentalium Tavera, 1942: 600. Tertiary.Arauco, Chile. Nomen nudum. Remarks: Feruglio(1949b: 239) cited this name, but did not validateit. We could not determine if this name was evervalidated; if so, it would be a junior homonym <strong>of</strong>Dentalium tenuistriatum Rouault, 1850 <strong>and</strong>Martin, 1880.tenuistriatum, Dentalium klipsteini “n.f.” Zardini,1978: 56, pl. 39, figs 19, 22, 23, pl. 40, fig. 13.Triassic, St. Cassian Formation. Cortina d’Ampezzo,Italy. Remarks: this taxon is an infrasubspecificname, pursuant to ICZN (1999) article 46.6.3,<strong>and</strong> therefore is not an available name pursuant toICZN (1999) article 46.5. Stiller (2001: 630) erroneouslyused this name as a valid sub<strong>species</strong> <strong>of</strong>Prodentalium klipsteini (Kittl, 1891).† tenuisulcatum, Dentalium (Fustiaria) Hölzl, 1962:135, 136, pl. 8, fig. 7. Oligocene, Chattian. Isaruferam Kalvarienberg (Tölz), Bavaria, Germany.Holotype Bayerisches Staatsammlung fürPaläontologie und Historische Geologie 2160(Sammlung Hölzl).teramachii, Entalina Habe, 1953: 299. Nomen nudum.Remark: validated by Kuroda & Habe in Habe1963: 273, pl. 38, figs 12, 13.teramachii, Entalina Kuroda & Habe in Habe, 1963:273, pl. 38, figs 12, 13. As a nomen nudum in Habe(1953: 299). <strong>Recent</strong>. Tosa Bay, Shikoku, Japan,200 m. Holotype <strong>and</strong> paratype NSMT. Junior synonym<strong>of</strong> Megaentalina mediocarinata (Boissevain,1906) (fide Habe 1964: 42).teres, Siphodentalium Jeffreys, 1883: 661, pl. 49, fig. 5.<strong>Recent</strong>. Type locality not specified. SyntypesBMNH 85.11.5.1318-1319 (Porcupine 1870 stn16-17a [<strong>of</strong>f Portugal]; stn 16: 39°55’N, 9°56’W,994 fms [1723 m]; stn 17: 39°42’N, 9°43’W, 600-1095 fms [1095-1998 m]; stn 17a: 39°39’N,9°39’W, 740 fms [1351 m]). Pulsellum teres(Jeffreys, 1883) n. comb. Eastern Atlantic: Europe.1095-2400 m.† terquemi, Dentalium Richardson, 1906: 589, 590[ex Tate MS]. Jurassic, Hettangian. Aiglemont,France. Remark: Engeser & Riedel (1992: 39) statedthat this <strong>species</strong> was a nomen dubium.tersus, Cadulus (Cadulus) Henderson, 1920: 149, 150,pl. 20, fig. 4. <strong>Recent</strong>. Off Barbados, State Univ.Iowa Exp. stn 51, 33 fms [60 m]. Holotype USNM596600. Cadulus tersus Henderson, 1920. WesternAtlantic, Caribbean. 60-137 m.† teschi, Dentalium subrectum subsp. Koperberg,1931: 25, 26. Neogene/Quaternary. Locality 1(“Clypeasterblok Noil Boenoe”); locality 2(“Tusschen [between] Niki-Niki en [<strong>and</strong>] HorstPené”); locality 4 (“Omgeving van Niki-Niki(gekocht van [purchased from (Mr.?)] Venema)”);locality 6 (“Nono Fatoe Fekoe (gemengd met[mixed with (shells from)] Toi Oesapi Soka)”),Timor. Junior synonym: Dentalium semarangenseAltena, 1938. Dentalium teschi Koperberg, 1931.Remark: see under Dentalium subrectum Martin,1885.tessellatum, Dentalium Scarabino, 1995: 214, 215, figs21a-e, 22. <strong>Recent</strong>. Philippines, RV CoriolisMUSORSTOM 3, stn DR140, 11°43’N,122°34’E, 93-99 m. Holotype <strong>and</strong> 3 paratypesMNHN. Junior synonym <strong>of</strong> Entalinopsis habutae(Kuroda & Kikuchi, 1933) (V. Scarabino, n. syn.).tesseragonum, Dentalium Sowerby in Broderip &Sowerby, 1832: 29, pl. 224, figs 21, 22. <strong>Recent</strong>.Gulf <strong>of</strong> Nicoya, Costa Rica; Puerto Portrero [CostaRica]; Xipixapi, Colombia. Syntypes BMNH1950.11.5.15-18. Tesseracme tesseragona (Sowerbyin Broderip & Sowerby, 1832) (fide Pilsbry &Sharp 1897: 34). Eastern Pacific: Colombia, CostaRica. 16-23 m.tetragonum, Dentalium (Dentalium) Brocchi, 1814:627, 628, pl. 15, fig. 26. Pliocene. Northern Italy.Holotype <strong>and</strong> paratype MSNM 505 (fide RossiRonchetti 1955: 342, 343, fig. 185). Junior synonyms:Dentalium quinquangulare Forbes, 1844 (fideMonterosato 1872: 27); Siphonodentalium pentagonumM. Sars, 1865 (fide M. Sars 1870: 195, placingS. pentagonum in synonymy with quinquangulare);Dentalium karreri Hörnes, 1875 (fide Pilsbry &Sharp 1898: 234). Entalina tetragona (Brocchi,1814) (fide Monterosato 1872: 27). Eastern Atlantic:Europe; Mediterranean. 68-2664 m. Remark: Pavia(1991: 133) suggested that Entalina viallii Caprotti,1962 might also be a junior synonym.tetrapleurum, Dentalium Boissevain, 1906: 41, 42,pl. 6, fig. 37. <strong>Recent</strong>. Anchorage <strong>of</strong> Djankar, Java,Siboga stn 4, 7°42’S, 114°12.6’E, 9 m. LectotypeZMA 3.06.049 (designated by Scarabino 1995:224); paralectotype ZMA 3.06.050 (1; Ruma-KudaBay, Roma Isl<strong>and</strong>, Siboga stn 279, 36 m).Tesseracme tetrapleura (Boissevain, 1906) (fideHabe & Kosuge 1964: 5, as synonym <strong>of</strong>T. quadrapicale; this synonymy was rejected byScarabino 1995: 224). Indo-Pacific: Indonesia toNew Caledonia. 9-70 m.tetraschistum, Siphodentalium Watson, 1879: 521[Watson 1886: 15, pl. 2, fig. 8]. <strong>Recent</strong>. Anchorage<strong>of</strong>f Fern<strong>and</strong>o Noronha Isl<strong>and</strong> [Brazil], Challengerstn 113a, 3°57’S, 32°24’30”W, 7-25 fms [13-46 m]. Holotype BMNH 1887.2.9.66. Junior synonyms:Siphonodentalium quadridentatum Dall,1881; Cadulus incisus Bush, 1885; Cadulus(Polyschides) quadridentatus acompsus Henderson,1920 (fide Scarabino 1980: 11). Polyschidestetraschistus (Watson, 1879) (fide Pilsbry & Sharp1898: 148). Western Atlantic, Caribbean: NorthCarolina (USA) to Argentina. 0-120 m.tetrodon, Cadulus (Polyschides) Pilsbry & Sharp, 1898:151, 152, pl. 29, figs 14-18. <strong>Recent</strong>. 5 miles <strong>of</strong>fZOOSYSTEMA • 2004 • 26 (4)655


Steiner G. & Kabat A. R.Cape Florida, 8 fms [15 m]. Syntypes ANSP 71070(5), USNM 84953, AMNH 148360. Polyschidestetrodon (Pilsbry & Sharp, 1898). Western Atlantic,Florida. 5-155 m. Remarks: Scarabino (1994: 310)listed this name as a junior synonym <strong>of</strong> P. tetraschistus(Watson, 1879), rejecting his previous conclusions(Scarabino 1980: 12) that this was a valid<strong>species</strong>. Since Scarabino (1994) did not explain whyhe changed his mind, <strong>and</strong> because the descriptions<strong>and</strong> comparison <strong>of</strong> tetrodon <strong>and</strong> tetraschistus givenby Pilsbry & Sharp (1898) clearly indicate two distinct<strong>species</strong>, the synonymy with tetraschistus byScarabino (1994) is here rejected. Boyko & Sage(1996: 31) listed the AMNH specimen as a paralectotype,although Henderson’s (1920: 101) citationto ANSP 71070 as the “type” referred to a lotcontaining 5 specimens, without designating alectotype.texasianum, Dentalium Philippi, 1849: 144. <strong>Recent</strong>.Galveston, Texas. Type material not located. Juniorsynonym <strong>of</strong> Paradentalium americanum (Chenu,1843) (fide Scarabino 1975: 183).† thalloides, Dentalium Conrad, 1833: 34, pl. 15,fig. 10. Eocene. Claiborne, Alabama, USA.Lectotype ANSP 65242 (designated by Palmer1937: 14); paralectotype ANSP 65243 (6). Juniorsynonyms: Dentalium alternatum Lea, 1833;Dentalium asgum de Gregorio, 1890; Dentaliumasgum var. tirpum de Gregorio, 1890; Dentaliumbimixtum de Gregorio, 1890 (fide Pilsbry & Sharp1898: 218). Dentalium thaloides [de Gregorio,1890: 171] is an incorrect subsequent spelling.† thallus, Dentalium Conrad, 1834: 142. Miocene.Suffolk, Virginia, USA. Cadulus thallus (Conrad,1834) (fide Dall 1892: 445 <strong>and</strong> Pilsbry & Sharp1898: 239).thaloides, see thalloides.thetidis, Dentalium Hedley, 1903: 327, fig. 61.<strong>Recent</strong>. 5-8 miles <strong>of</strong>f Port Kembla, New SouthWales, Australia, 63-75 fms [115-137 m].Lectotype AMS C16212 (designated by Colman1958: 141, 142 as “holotype”); paralectotypes AMSC170641 (2; <strong>of</strong>f Cape Three Points, 41-50 fms [75-91 m]). Striodentalium thetidis (Hedley, 1903)(fide Scarabino 1995: 239). Indo-Pacific: Australia,Philippines. 80-242 m. Remark: Lamprell & Healy(1998: 48) listed Entalinopsis intercostata(Boissevain, 1906) as a junior synonym <strong>of</strong> this<strong>species</strong>.thielei, Cadulus Plate, 1908b: 3, 4, text-figs 6-11.<strong>Recent</strong>. Deutsche Südpolar Exp. stn 56, 380-385 m. Lectotype ZMB 59726a (designated byKilias 1995: 177); paralectotype ZMB 59726b (4).Cadulus thielei Plate, 1908. East Antarctica. 362-385 m.thomassini, Pulsellum Scarabino, 1995: 321-322,figs 123, 125b. <strong>Recent</strong>. Madagascar, NortheastBanc du Geyser, BENTHEDI stn DR11,12°16.3’S, 46°42.2’E, 2450-2300 m. HolotypeMNHN; paratypes MNHN (6 from DR11; 2 fromDR88, 11°46’S, 47°34’E, 3700 m; 2 from DS106,20°28’S, 55°05’E, 1710-1730 m; 2 from DS151,20°51’S, 56°03’E, 3240-3300 m). Pulsellumthomassini Scarabino, 1995. Indo-Pacific:Madagascar to New Caledonia. 1730-3716 m.tibanum, Dentalium (Antalis) Nomura, 1940: 101,102, pl. 1, fig. 11. <strong>Recent</strong>. E <strong>of</strong> Inubo-zaki, Japan,Husa-Maru stn 15, 40 m. Syntypes SHM 17981(10). Junior synonym: Antalis boissevainae Palmer,1974, a replacement name for Dentalium entalis var.indicum Boissevain, 1906 non Dentalium indicumChenu, 1843 (fide Habe 1964: 19; in contrast toHabe 1963: 262, where he did not synonymize thetwo <strong>species</strong>). Antalis tibana (Nomura, 1940) (fideHabe 1961: 105). Indo-Pacific: Indonesia to Japan<strong>and</strong> Vanuatu. 0-200 m.† tigillum, Dentalium Yokoyama, 1931: 194, pl. 11,figs 5, 6. Neogene. Unetomari, Obirashibets,Teshio, Japan; Morai, near Atsuta, Ishikari, Japan.Syntype GIUT (fide Hanzawa et al. 1961: 209).tignum, Dentalium (Dentalium) Colman, 1958: 141,fig. 1. <strong>Recent</strong>. 35 miles E <strong>of</strong> Sydney, New SouthWales, 800 fms [1460 m]. Holotype AMS C24485;paratype AMS C170643 (27.5 miles E <strong>of</strong> Sydney,New South Wales, 550 m; fide Lamprell & Healy1998: 29). Junior synonym <strong>of</strong> Paradentaliumhemileuron (Verco, 1911) (fide Lamprell & Healy1998: 29).tirpum, Dentalium asgum var. de Gregorio, 1890: 172.Eocene. Claiborne, Alabama. Type material lost(fide Palmer & Brann 1965: 369). Junior synonym<strong>of</strong> Dentalium thalloides Conrad, 1833 (fide Pilsbry& Sharp 1898: 218).tiwhana, Dentalium Dell, 1953. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.tolmiei, Cadulus Dall, 1897: 13, pl. 1, fig. 8. <strong>Recent</strong>.Near Victoria, Vancouver Isl<strong>and</strong> [British Columbia,Canada], 60 fms [110 m]. Holotype not located;figured paratype USNM 107613 (Shimek [1989:235] referred to USNM 107613 as the “figuredparatype”, but is more likely to be the holotype,since Dall [1897] gave the measurements <strong>of</strong> a singleshell; it is improbable that only a paratype wouldhave been figured in the original). Gadila tolmiei(Dall, 1897) (fide Pilsbry & Sharp 1898: 181).Eastern Pacific: Canada to California. 80-350 m.Remark: Emerson in Turgeon (1998: 55) placedthis <strong>species</strong> in the genus Polyschides.tomlini, Dentalium Melvill, 1918: 155, 156, pl. 5,fig. 31. <strong>Recent</strong>. Karachi [Pakistan]. SyntypesBMNH 1921.1.28.36-40, NMW 1955.158.592-93 (fide Trew, 1987: 68). Dentalium tomliniMelvill, 1918. Indian Ocean, India. No bathymetricdata available.tonosum, see torosum.† tornatissimum, Dentalium (Episiphon) Tate, 1899:265, 266, pl. 8, figs 7-7a. “Miocene” (Pliocene).Gippsl<strong>and</strong> Lakes, Australia. Holotype <strong>and</strong> paratype656ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)SAM T 1609. Fustiaria tornatissima (Tate, 1899)(fide Ludbrook 1959: 143, 144, pl. 2, figs 6, 7).tornatum, Dentalium Watson, 1879: 518, 519[Watson 1886: 13, pl. 2, fig. 3]. <strong>Recent</strong>. Levuka,Fiji, Challenger stn 173, 174, 12 fms [22 m].Syntypes BMNH 1887.2.9.57-60 (6). Junior synonym<strong>of</strong> Episiphon subtorquatum (Fischer, 1871)(fide Scarabino 1995: 286).torosum, Dentalium Zenker, 1836: 231, 232. Triassic.Rauhthal, near Jena, Germany. Junior synonym <strong>of</strong>Dentalium torquatus (Schlotheim, 1820) (fidePilsbry & Sharp 1898: 228). Dentalium tonosum[Geinitz, 1837: caption to pl. 1, fig. 2] is an incorrectsubsequent spelling.† torquatus, Dentalites Schlotheim, 1820: 94. Triassic,Wellenkalk. Farrenstadt, near Querfurth, Germany.Entalis torquata (Schlotheim, 1820) (fide Diener1926: 230).tosaensis, Antalis Habe, 1963: 264, pl. 38, fig. 2.<strong>Recent</strong>. Off Tosa Bay, Shikoku, Japan, 200 m.Holotype <strong>and</strong> paratype in NSMT. Striodentaliumtosaensis (Habe, 1963) (fide Habe 1964: 23). Indo-Pacific: Japan. 200-620 m.† totomiensis, Dentalium (Antalis) Makiyama, 1931:44, 45, pl. 1, figs 2-4. Pliocene, Hosoya Beds.Localities 522 <strong>and</strong> 523, Totomi, Ugari District,Japan. Holotype FGK (fide Hanzawa et al. 1961:209).toyamaense, Dentalium Kuroda & Kikuchi, 1933: 11,pl. 1, figs 5, 6. <strong>Recent</strong>. Toyama Bay, Honshu,Japan, 200-400 m. Syntypes in FGK. Rhabdustoyamaense (Kuroda & Kikuchi, 1933) (fide Steiner1999: 154). Indo-Pacific: Japan. 200-1400 m.Remarks: the original description did not specify aholotype. Habe (1963: 269) referred to the “type<strong>and</strong> paratype specimens”, but did not indicatewhich specimen was the “type”, so this did notconstitute a valid lectotype designation.trachea, Dentalium Montagu, 1803. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.tracheatum, Dentalium (Plagioglypta) Boissevain, 1906:56, 57, pl. 4, fig. 22. <strong>Recent</strong>. Indonesia, B<strong>and</strong>a Sea,Siboga stn 208, 05°39’S, 122°12’E, 1886 m.Lectotype ZMA 3.06.076 (designated by Scarabino1995: 297); paralectotypes ZMA 3.06.075(3; Flores Sea, Siboga stn 45, 7°24’S, 118°15.2’E,794 m), ZMA 3.06.077 (2), ZMA 3.06.078(11; Arafura Sea, Siboga stn 271, 5°46.7’S,134°00’E, 1788 m), ZMA 3.06.079 (4; Timor Sea,Siboga stn 300, 10°47.6’S, 123°23.1’E, 918 m),ZMA 3.06.080 (1, second specimen probably lost;Flores Sea, Siboga stn 314, 7°36’S, 117°30.8’E,694 m). Junior synonym <strong>of</strong> Omniglypta cerina(Pilsbry, 1905) (fide Habe 1964: 37; fide Scarabino1995: 297).transiens, Dentalium (Antalis) kickxi var. Steininger,1963b: 36, 37, pl. 12, figs 1a, b [nomen nudum inSteininger 1963a: 34]. Burdigal, Lower Miocene.Dornergraben bei Fels am Wagram, Niederösterreich[Lower Austria, Austria]. Holotype PIUV1663. Junior homonym <strong>of</strong> Dentalium michelottiivar. transiens Boettger, 1907. Junior synonym <strong>of</strong>Antalis kickxii (Nyst, 1843).† transiens, Dentalium michelotti var. Boettger, 1907:209. Miocene. Parau Ungurului, Valea Semini, <strong>and</strong>Valea Casilor, Kostej, Romania. Lectotype SMFXIII.1a (designated by Zilch 1934: 279, pl. 22,fig. 28). Junior homonym: Dentalium (Antalis)kickxii var. transiens Steininger, 1963. Dentaliummichelottii transiens Boettger, 1907 (fide Zilch1934: 279).transitorius, Cadulus (Cadulus) Henderson, 1920:143, 144, pl. 19, figs 6, 7. <strong>Recent</strong>. Little BahamaBank, USBF stn 2654, 27°57’30”N, 77°27’30”W,660 fms [1204 m]. Holotype USNM 330589;paratypes USNM 95375, USNM 108170, USNM314821, USNM 312822, USNM 314823, USNM323779, USNM 323813, USNM 330676, AMNH148334 (8; USBF stn 2668, <strong>of</strong>f Fern<strong>and</strong>ina,Florida, 294 fms [537 m], fide Boyko & Sage 1996:31), MCZ 7744. Cadulus transitorius Henderson,1920. Western Atlantic, Caribbean: Bahamas toBrazil. 148-1349 m.translucidum, Dentalium Deshayes, 1825: 362 [42 inreprint], pl. 16, fig. 26. <strong>Recent</strong>. Type locality notspecified. Type material not located. Nomen dubium.Laevidentalium(?) translucidum (Deshayes,1825) (fide Pilsbry & Sharp 1897: 99), but not incurrent use. Remark: generic assignment uncertain.translucidum, Dentalium Chenu, 1843: pl. 3, fig. 12[1852: 8]. <strong>Recent</strong>. Type locality not specified. Typematerial not located. Junior homonym <strong>of</strong>Dentalium translucidum Deshayes, 1825. Juniorsynonym <strong>of</strong> Graptacme semistriata (Turton, 1819)(fide Pilsbry & Sharp 1897: 90).† transsilvanicum, Siphonodentalium Boettger, 1897:55, 56. Miocene. Kostej <strong>and</strong> Ober-Lapugy, Banat,Romania. Lectotype SMF XIII.4a (designated byZilch 1934: 279, pl. 22, fig. 26). Siphonodentaliumtranssylvanicum Boettger, 1897 (fide Zilch 1934:279).† transversatum, Dentalium Link, 1807: 23. [Fossil].Sternberg, Germany.transversostriatum, Dentalium Boissevain, 1906: 32,pl. 4, fig. 23. <strong>Recent</strong>. Indian Ocean, Off Saleyer,Siboga stn 212, 5°54.5’S, 120°19.2’E, 462 m.Syntypes ZMA 3.06.028-029 (3). Junior synonym<strong>of</strong> Fissidentalium shopl<strong>and</strong>i (Jousseaume, 1894)(fide Scarabino 1995: 248). Remark: V. Scarabino(in litt. 5.XI.2001) suggested that the status <strong>of</strong>F. transversostriatum vis-à-vis F. shopl<strong>and</strong>i requiresfurther research.† trautscholdi, Dentalium Koenen, 1868: 160.Oligocene. Aral Sea. Junior synonym: Dentaliumhaeringense Dreger, 1892 (fide Merklin & Goncharova1967: 118). Dentalium (Coccodentalium)trautscholdi Koenen, 1868 (fide Merklin &Goncharova 1967: 118).ZOOSYSTEMA • 2004 • 26 (4)657


Steiner G. & Kabat A. R.† tredecimcostatum, Dentalium (Antale) novemcostatumvar. Sacco, 1897: 104, pl. 8, fig. 70 [as a nomennudum in Sacco 1896: 97]. Pliocene. San Agata,Italy. Type material not mentioned by FerreroMortara et al. (1984: 305).tricarinatum, Dentalium (Bathoxiphus) Boissevain,1906: 48, 49, text-fig. 23, pl. 6, figs 40, 41. <strong>Recent</strong>.Ceram Sea, Siboga stn 178, 2°40’S, 128°37.5’E,835 m. Lectotype ZMA 3.06.060 (designated byHabe 1964: 33, 34, <strong>and</strong> unnecessarily by Scarabino1995: 316); paralectotypes ZMA 3.06.056 (1),ZMA 3.06.057 (1; B<strong>and</strong>a Sea, Siboga stn 208,5°39’S, 122°12’E, 1886 m), ZMA 3.06.058 (1;B<strong>and</strong>a Sea, Siboga stn 211, 5°40.7’S, 120°45.5’E,1158 m), ZMA 3.06.059 (3; Timor Sea, Siboga stn300, 10°48.6’S, 123°23.1’E, 918 m). Junior synonym:Dentalium (Compressidens) capense Tomlin,1931 (fide Scarabino 1995: 316). Rhomboxiphustricarinatus (Boissevain, 1906) (fide Chistikov1983: 185). Indo-Pacific: Africa to Japan <strong>and</strong>Tasman Sea. 495-4830 m.tricostatum, Dentalium Goldfuss, 1844. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.tridentata, Pertusiconcha Chistikov, 1982a: 678-681,pl. 4, figs 2-13, pl. 5, figs 10, 11. <strong>Recent</strong>. TasmanSea, Mendeleev stn 1244, 31°42.6’S, 159°00.2’E,1640 m. Holotype IOM. Pertusiconcha tridentataChistikov, 1982. Indo-Pacific: Madagascar to NewZeal<strong>and</strong>. 825-2470 m.† triedra, Serpula Quenstedt, 1856: 200, 329, pl. 25,fig. 55. Jurassic. Dörlbach, Germany. Remarks:Quenstedt (1856: 329) suggested that this <strong>species</strong>might be referrable to Dentalium; Richardson (1906:590-592) suggested that this <strong>species</strong> might be a seniorsynonym <strong>of</strong> Dentalium trigonalis Moore, 1866.trigonale, Dentalium Martin, 1885: 191, pl. 10,fig. 192; 1887: 191, pl. 10, fig. 192. Miocene.Ngembak, Java, Indonesia. Holotype RGM 7276(fide Van den Hoek Ostende et al. 2002: 112).Junior homonym <strong>of</strong> Dentalium trigonalis Moore,1866; replacement name: Gadilina trigonaloidesPalmer, 1974.† trigonalis, Dentalium “?” Moore, 1866: 86, pl. 5,fig. 22. Jurassic, Middle Lias (Pliensbachian).Camerton, Engl<strong>and</strong>, United Kingdom. HolotypeGeology Museum Bath (Engl<strong>and</strong>), M973 (fidePalmer, 1987a: 30). Progadilina trigonalis (Moore,1866) (fide Emerson 1987: 30). Remark:Richardson (1906: 590-592) suggested that Serpulatriedra Quenstedt, 1856 might be a senior synonym<strong>of</strong> Dentalium trigonalis.† trigonaloides, Gadilina Palmer, 1974b: 124.Miocene. Ngembak, Java, Indonesia. HolotypeRGM 7276 (fide Van den Hoek Ostende et al.2002: 112). Replacement name for Dentalium trigonaleMartin, 1885 non Dentalium trigonalis Moore,1866.trigonum, Dentalium Hoeninghaus, 1831: 155. Nomennudum. Tertiary. Tabbiano [Italy].† triquetrum, Dentalium Brocchi, 1814: 628.Pliocene. Italy. Lectotype MSNM 506 (selected byRossi Ronchetti 1955: 340, 341, fig. 184). Gadilinatriquetra (Brocchi, 1814) (fide Pavia 1991: 130,131, pl. 7, figs 7, 8).triquetrum, Dentalium “(?)” Tate, 1887: 193, pl. 20,fig. 3. Upper Eocene. Glauconitic s<strong>and</strong>s, Adelaidebore, Kent Town, South Australia, Australia.Holotype <strong>and</strong> 6 paratypes SAM T 252A. Juniorhomonym <strong>of</strong> Dentalium triquetrum Brocchi, 1814.Replacement name: Dentalium tatei Sharp &Pilsbry in Pilsbry & Sharp, 1898.truncatum, Dentalium (Gadilina) Boissevain, 1906:51, text-fig. 27, pl. 6, fig. 33. <strong>Recent</strong>. Localities <strong>of</strong>syntypes: ZMA 3.06.065 (1; Celebes Sea, Siboga stn90, 01°17.5’N, 118°53’E, 281 m), ZMA 3.06.066-067 (2; Arafura Sea, Siboga stn 256, 5°26.6’S,132°32.5’E, 397 m). Episiphon(?) truncatum(Boissevain, 1906) (fide Scarabino 1995: 289).Indo-Pacific: Indonesia. 281-397 m.† tryoni, Dentalium Pilsbry & Sharp, 1898a: 468,469, pl. 10, figs 5, 9, pl. 11, fig. 22. Oligocene.Dominican Republic. Syntypes ANSP 2710 (fidePilsbry 1922: 399), ANSP 79570.tubiforme, Dentalium Boissevain, 1906: 19, 20, pl. 6,fig. 5. <strong>Recent</strong>. Indonesia, Siboga stn 212, 05°56’S,120°19’E, 462 m. Lectotype ZMA 3.06.013 (designatedby Scarabino 1995: 310); paralectotype ZMA3.06.012 (1; stn 159, 0°59.1’S, 129°19.2’E, 411m). Spadentalina tubiformis (Boissevain, 1906)(fide Habe 1963: 264). Indo-Pacific: Indonesia toJapan. 200-702 m.tubulatum, Dentalium (Antalis) Henderson, 1920: 56,pl. 8, fig. 5. <strong>Recent</strong>. Bahia Honda, Cuba, Blake stn20, 23°02’30”N, 83°11’W, 220 fms [402 m].Syntypes USNM 95367 (2). Antalis tubulata(Henderson, 1920). Western Atlantic, Caribbean.31-536 m.tugaruense, Dentalium Nomura & Hatai, 1940: 73,pl. 3, fig. 4. <strong>Recent</strong>. Kyûruko-sima Isl<strong>and</strong>s, AomoriPrefecture, Northeast Honsyû, Japan, stn 18,40°36’N, 139°45’45”E, 103 m. Holotype SHM10997. Junior synonym <strong>of</strong> Entalinopsis intercostata(Boissevain, 1906) (fide Habe 1963: 272).tumida, see tumidula.tumidosus, Cadulus Jeffreys, 1877: 156, 157. <strong>Recent</strong>.Type locality not specified. Syntypes BMNH1885.11.5.1288-90 (3; Porcupine, no locality),BMNH 1877.11.28.58 (3) <strong>and</strong> USNM 169497 (1)(Valorous, stn 12, 56°11’N, 37°41’W, 1450 fms[2646 m]), USNM 169498 (1; Porcupine 1869 stn39), USNM 169500 (18; Porcupine , 1870 stn 22,26, 31, 34), USNM 169501 (15; Porcupine 1870stn 16, 54°19’N, 11°50’W, 816 fms [1489 m]),USNM 169502 (many; Porcupine 1870 stn 17a,39°39’N, 9°39’W, 740 fms [1351 m]), USNM169503 (many; Porcupine 1870 stn 17 39°42’N,9°43’W, 600-1095 fms [1095-1998 m]). Cadulustumidosus Jeffreys, 1877. Western Atlantic:658ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Caribbean to Brazil; Eastern Atlantic: Europe,Africa. 295-2853 m. Remarks: also described fromJosephine Exp. (110-550 fms [201-1004 m]) <strong>and</strong><strong>fossil</strong> outcrops at “Messina [Sicily] (Seguenza)”;these specimens were not located. Jeffreys (1883:665) mentions a var. minor (nomen nudum) fromPorcupine 1870 stns 17a, 22, 24, <strong>and</strong> 31-34, thetypes <strong>of</strong> which have not been segregated from those<strong>of</strong> the full <strong>species</strong>.tumidula, Cadulus jeffreysi var. Jeffreys, 1883: 665.<strong>Recent</strong>. Travailleur Exp. 1880 [Bay <strong>of</strong> Biscay].Syntypes USNM 175843 (21). Synonym <strong>of</strong>Cadulus jeffreysi (Monterosato, 1875). Cadulus jeffreysivar. tumida “Jeffreys” [Locard, 1898: 141] isan incorrect subsequent spelling.† tumidum, Dentalium W. D. Smith, 1913: 292,pl. 19, fig. 2. “Pleistocene or <strong>Recent</strong> shales.”Locality 1054, San Rafael, Agusan River, Mindanao,Philippines.† turgidus, Cadulus Meyer, 1886: 65, pl. 1, fig. 10.Eocene. Matthews’ L<strong>and</strong>ing, Alabama, USA.Syntype USNM Paleobiology 638813 (as “holotype”in Palmer & Brann 1965: 364).turnerae, Wemersoniella Scarabino, 1986a: 3, 4, figs 1-5, 12-16. <strong>Recent</strong>. Northwest Atlantic, Chain 50 stn80, 34°49.8’N, 66°34’W, 4970 m. Holotype MCZ293959; paratypes MCZ 293957, 293958, 293959,293960, 293961, MHNM 14757, 14758, 14759,MNHN. Wemersoniella turnerae Scarabino, 1986.North Atlantic: Puerto Rican Trench to Europe.4240-4970 m. Remarks: MCZ paratype lots not yetdeposited. Scarabino (1986a: 7) listed the localities<strong>of</strong> all the lots <strong>of</strong> this <strong>species</strong>, but did not correlatethe stations with the museum catalog numbers.† turoniense, Dentalium Woods, 1896: 96, pl. 4,figs 16, 17. Cretaceous. Chalk Rock <strong>of</strong> Winchester,Cuckhamsley <strong>and</strong> Luton, Engl<strong>and</strong>, UnitedKingdom.† turritum, Dentalium Lea, 1833: 35, pl. 1, fig. 3.Eocene. Claiborne, Alabama, USA. HolotypeANSP 5015 (fide Palmer & Brann 1965: 365).Polyschides turritus (Lea, 1833) (fide Pilsbry & Sharp1898: 239).tuscarorae, Costentalina Chistikov, 1982b: 1316, 1317,pl. 1, figs 7, 8, pl. 3, figs 12-18, pl. 4, figs 1, 2.<strong>Recent</strong>. Off Japan, Vitjaz stn 3575-T, 38°02.1’N,146°33.1’E, 5475 m. Holotype IOM; paratypesIOM (6 from stn 3575-T; 1 from stn 3198-T,39°02.9’N, 151°50.6’E, 5817-5807 m; 1 from stn3232-D, 33°18’N, 149°45.7’E, 6070 m; 1 from3240-D, 37°42.5’N, 156°23.6’E, 5790 m; 1 fromstn 5620-T, 44°48’N, 156°33’E, 5045-5005 m;1 from stn 5623-T, 45°26’N, 154°59’E, 5045-4995m; 8 from 5624-T, 45°26’N 154°12’E, 5200 m;1 from stn 7499-T, 37°58’N, 144°12’E, 6499-6640 m). Costentalina tuscarorae Chistikov, 1982.Indo-Pacific: Philippines to Japan. 2050-6640 m.† tutongense, Dentalium (Dentalium) javanum subsp.Beets, 1984: 26, pl. 1, figs 20-23. Late Miocene.Preangerian. Near Sekurau, northern Kutai,Kalimantan Timur [East Borneo], Indonesia.Holotype RGM 315069; paratypes RGM 315068,315070, 315071.† twistringense, Fissidentalium R. Janssen, 1989: 82,84, pl. 1, figs 3-6. Miocene, Reinbekian. Twistringen,Niedersachsen, Germany. Holotype SMF 308305 ;paratypes SMF 308306, SMF 308307, SMF308309/200; RGM 227724.typa, Throopella Greger, 1933. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.tytthum, Siphodentalium Watson, 1879: 520 [Watson1886: 14, pl. 2, fig. 5]. <strong>Recent</strong>. North <strong>of</strong> CulebraIsl<strong>and</strong>, West Indies, Challenger stn 24, 18°38’30”N,65°05’30”W, 390 fms [712 m]. Syntypes BMNH1887.2.9.63 (4). Siphonodentalium(?) tytthumWatson, 1879. Known from the type locality only.† uchauxiense, Dentalium Cossmann, 1897: 267, 268,pl. 2, figs 33, 34. Cretaceous, Turonian. Uchaux,Vaucluse, France.undatum, Dentalium Defrance, 1819. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.† undecimcostata, Dentalium (Antale) novemcostatumvar. Sacco, 1897: 103, pl. 8, figs 66-68 [as a nomennudum in Sacco 1896: 97]. Pliocene. San Agata <strong>and</strong>Astigiana, Italy. Syntypes MIGT BS.106.02.058-BS.106.02.60 (fide Ferrero Mortara et al. 1984:305).† undiferum, Dentalium Koenen, 1885: 72, pl. 3,fig. 17. Paleocene. Near Copenhagen, Denmark.† undulatum, Dentalium Goldfuss, 1841: 3, pl. 166,fig. 8 [ex Münster MS]. Triassic. St Cassian, Tirol,Austria/Italy. Plagioglypta undulata (Goldfuss,1841) (fide Stiller 2001: 624). Remark: also cited inWissmann, 1841: 91, pl. 9, fig. 6.† uscarianum, Dentalium (Fissidentalium) Olsson,1922: 338, 339 [166, 167], pl. 15, fig. 1. Miocene,Uscari Stage. Costa Rica <strong>and</strong> Panama (various localities).Holotype PRI 21115 (Rio Coches, CostaRica) (fide Brann & Kent 1960: 322).usitatum, Dentalium E. A. Smith, 1894: 168, pl. 4,figs 16, 16a. <strong>Recent</strong>. Off Colombo, Ceylon [SriLanka], 6°32’N, 79°37’E, 675 fms [1232 m].Syntypes ZSI 5410 (fide list in BMNH), BMNH1894.9.11.8 (1; Bay <strong>of</strong> Bengal, 597 fms [1090 m]).Antalis usitata (E. A. Smith, 1894) (fide Ludbrook1954: 91). Indo-Pacific: Red Sea to New Caledonia.56-1542 m.† vacavillensis, Dentalium Palmer, 1923: 302, 303.Middle Eocene. UCMP Locality 3573, nearVacaville, California, USA. Holotype UCMP30593; paratypes UCMP 30594-30595.vagina, Dentalium Jeffreys, 1877: 155. <strong>Recent</strong>.Greenl<strong>and</strong>, Valorous stn 16, 55°10’N, 25°58’W,1785 fms [3215 m]. Holotype USNM 174993.Junior synonym <strong>of</strong> Antalis agilis (G. O. Sars, 1872)(fide Pilsbry & Sharp 1897: 46). Remark: Jeffreysalso referred to “perhaps two imperfect specimens<strong>of</strong> a Dentalium from Station 12, 1450 fathoms,ZOOSYSTEMA • 2004 • 26 (4)659


Steiner G. & Kabat A. R.[that] may belong to the same <strong>species</strong>”. However,this is too tentative to consider those two specimens,now catalogued as USNM 174992 (Valorousstn 12, 56°11’N, 37°41’W, 1450 fms [Greenl<strong>and</strong>,2646 m]), to constitute type material.vagina, Fustiaria Scarabino, 1995: 281, figs 71h, 87,88i. <strong>Recent</strong>. New Caledonia, Chesterfield Isl<strong>and</strong>s,MUSORSTOM 5, stn DW340, 19°49’S, 158°41’E,675-680 m. Holotype MNHN; paratypes MNHN(1 from DW341; 1 from DW357; 2 from DW46;2 from DW49; 1 from CP290; 1 from DW468),AMS C201726 (1), NMNZ M268953 (1; DW49),USNM 890868. Fustiaria vagina Scarabino, 1995.Indo-Pacific: New Caledonia. 290-820 m.† valangiense, Dentalium Pictet & Campiche, 1864:723, pl. 98, figs 16-18. Cretaceous, Neocomian,Valanginian. Villers-le-Lac, near Locle, Sainte-Croix, Switzerl<strong>and</strong>.valdiviae, Cadulus (Gadila) Jaeckel, 1932: 312,fig. 11. <strong>Recent</strong>. Off Somalia, Valdivia stn 251,01°41’S, 41°47’E, 693 m. Lectotype ZMB 75370a(designated by Kilias 1995: 177); paralectotypesZMB 75370b (1), ZMB 75371 (1; stn 256,01°49’S, 45°29.5’E, 1134 m). Cadulus valdiviaeJaeckel, 1932. Indian Ocean, Africa. 693-1134 m.valdiviae, Dentalium (Antalis) Plate, 1908a: 355, 356,pl. 30, figs 38, 39. <strong>Recent</strong>. North Atlantic, S <strong>of</strong>[Wyville] Thomson Rise, Valdivia stn 11,58°36.6’N, 11°33’W, 1750 m. Lectotype ZMB61079 (designated by Kilias 1995: 177). Antalisvaldiviae (Plate, 1908). Known from the typelocality only. Remarks: in addition to the singlewell preserved specimen, Plate mentioned twoincomplete specimens <strong>and</strong> a fragment that were notlisted by Kilias (1995: 177). These specimenswould constitute paralectotypes, since Plate did notspecify a holotype. The description, the illustration,<strong>and</strong> the locality fit well with Fissidentalium c<strong>and</strong>idum(Jeffreys, 1877), but we have not yet seen thetypes.vallicolens, Dentalium Raymond, 1904: 123, 124.<strong>Recent</strong>. Type locality not specified; syntypes fromSanta Monica Bay [California], <strong>of</strong>f Redondo,Vincente (or Redondo) submerged valley, 145 fms[265 m]. Syntypes CAS 064381 (1), CAS 064383(1), CAS 064385 (1), CAS 066519 (4); the syntypesfrom the following localities could not belocated: stn 14, <strong>of</strong>f Point Fermin, California,100 fms [183 m]; stn 70, La Jolla submerged valley,California, 54-117 fms [99-214 m]; stn 79, <strong>of</strong>f SanDiego, California, 64 fms [117 m]. Dentalium vallicolensRaymond, 1904. Eastern Pacific:California. 154-410 m.† vaningeni, Dentalium n. nom. Late Cretaceous,Senonian. Near Kerak, Jordan Valley, Jordan.Syntype YPM 35573 (fide White 1998: 4).Replacement name for Dentalium hexagonum VanIngen, 1905 non Dentalium hexagonum Gould,1859.variabile, Dentalium Deshayes, 1825: 367 [32, 33 inreprint], pl. 16, ig fig. 30. <strong>Recent</strong>. India (?).Lectotype <strong>and</strong> 163 paralectotypes MNHN (designatedby Scarabino 1995: 201). Junior synonyms:Dentalium multistriatum Deshayes, 1825 (fideScarabino 1995: 200, by First Reviser action,according to ICZN 1999: article 24); Dentaliumbelcheri Sowerby, 1860 (fide Habe 1963: 258).Dentalium variabile Deshayes, 1825. Indo-Pacific:Indonesia to Japan. 10-75 m.varians, Dentalium (Antalis) ceratum Scarabino, 1973:196, 197, pl. 1, figs 4, 4a-d. <strong>Recent</strong>. 24°20’S,44°40’W, 130 m. Holotype MHNM 2702;paratypes MHNM 2703-2706. Sub<strong>species</strong> <strong>of</strong>Antalis cerata (Dall, 1881). Western Atlantic:Brazil to Uruguay. 110-175 m.† varicostata, Dentalium Dockery in MacNeil &Dockery, 1984: 246, 247, pl. 69, figs 9, 11-16,pl. 70, figs 1-6. Lower Oligocene, Mint SpringFormation. Mississippi Geological Survey 99,Mississippi, USA. Holotype USNM Paleobiology376680.† vaughani, Cadulus Dall, 1912: 6. Pleistocene. USGSstn 5850, Canal Zone, Panama. Syntypes USNMPaleobiology 214361 (numerous).† vaughani, Dentalium Stephenson, 1941: 252, 253,pl. 46, fig. 15. Cretaceous, Navarro Group. USGSlocality 16170, Neyl<strong>and</strong>ville marl, on the CorsicanaRoad, 2.5 miles N <strong>of</strong> Corbet, Navarro County,Texas, USA. Holotype USNM Paleobiology 76765;17 paratypes USNM Paleobiology 76766.vayssierei, Dentalium Fenaux, 1942. Not a scaphopod.See Other taxa described as <strong>Scaphopoda</strong>.vemae, Costentalina Scarabino, 1986b: 2, 3, figs 1-7.<strong>Recent</strong>. Vema stn 81 (V-17-81), 44°35’S, 44°19’W[Argentine Basin], 5332 m. Holotype AMNH202762; paratype AMNH 202763 (1; stn 85 [V-17-85], 47°35’S, 43°21’W, 5495 m). Costentalinavemae Scarabino, 1986. Western Atlantic:Argentine Basin. 5332-5495 m.† venezuelana, Fustiaria (Laevidentalium “?”)Weisbord, 1964: 127, 128, pl. 17, fig. 7, pl. 18,figs 7, 8. Pliocene, Playa Gr<strong>and</strong>e Formation.Weisbord Station W-23, north flank <strong>of</strong> PuntaGorda anticline,Venezuela. Holotype PRI 26888.ventricosa, Cadulus monterosatoi var. Locard, 1898:142, pl. 7, figs 19-21. <strong>Recent</strong>. Type locality notspecified [North Atlantic]. Type material not located.Synonym <strong>of</strong> Cadulus monterosatoi Locard,1897.ventricosa, Cadulus tumidosus var. Locard, 1898: 143.<strong>Recent</strong>. Type locality not specified [NorthAtlantic]. Type material not located. Synonym <strong>of</strong>Cadulus tumidosus Jeffreys, 1877.† ventricosum, Dentalium Bronn, 1828: 539. Miocene.Castell’Arquato, Piacenza, Italy. Junior synonym:Siphonodentalium (Loxoporus) ligusticus Razzore,1896 (fide Pilsbry & Sharp 1898: 240). Remarks:Doderlein (1864: 98 [16]) listed this as “Gadus ven-660ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)tricosus nob.” but this was merely a new generic combination,not a new <strong>species</strong>. Gadila ventricosa (Bronn,1828) (fide Pavia 1991: 139-144, pl. 8, figs 9-14;table 4; text-fig. 7). This <strong>species</strong> was misidentified asDentalium coarctatum by Lamarck, 1818.ventricosus, Gadus Benoist, 1873: 269. Miocene.Giradueau, France. Junior secondary homonym <strong>of</strong>Cadulus ventricosus (Bronn, 1828). Replacementname: Gadila benoisti Cossmann & Peyrot, 1917.† venustum, Dentalium Meek & Worthen, 1861: 145.Carboniferous. Waterloo, Monroe County, Illinois,USA.verconis, Dentalium (Fissidentalium) Cotton &Ludbrook, 1938: 221, 222, pl. 12, fig. 1. <strong>Recent</strong>.Off Beachport, South Australia, 200 fms [366 m].Holotype SAM D 13341. Fissidentalium verconis(Cotton & Ludbrook, 1938). Pacific Ocean: SouthAustralia. 150-1210 m.vernedei, Dentalium Hanley in Sowerby, 1860: 101,pl. 223, fig. 3. <strong>Recent</strong>. Japan. Holotype UMZCI.100,760. Pictodentalium vernedei (Hanley inSowerby, 1860) (fide Habe 1964: 16). Indo-Pacific:China, Japan. 20-128 m.verrilli, Cadulus (Gadila) Henderson, 1920: 136, 137,pl. 19, fig. 13. <strong>Recent</strong>. Off Martha’s Vineyard[Massachusetts], USBF stn 871, 40°02’54”N,70°23’40”W, 115 fms [210 m]. Holotype USNM38693; paratypes USNM 38693 (7), USNM 38694(1; <strong>of</strong>f Martha’s Vineyard, USFB stn 873, 100 fms[183 m]), USNM 61144 (5), MCZ 186819 (2;USFC 871, 115 fms [210 m]), AMNH 148343 (1).Gadila verrilli (Henderson, 1920). WesternAtlantic. 182-210 m.verrilli, Dentalium (Fissidentalium) meridionaleHenderson, 1920: 62, 63, pl. 9, fig. 3. <strong>Recent</strong>. OffGeorge Banks, USBF stn 2083, 959 fms [1750 m].Lectotype (<strong>of</strong> solidum Verrill) USNM 34687 (designatedby Henderson 1920: 63); paralectotypesUSNM 34688, USNM 34904, USNM 34911,USNM 35635, USNM 35636, USNM 35645,AMNH 148370 (3). Replacement name forDentalium solidum Verrill, 1884 non Dentaliumsolidum Hutton, 1873. Junior synonym <strong>of</strong> Fissidentaliumc<strong>and</strong>idum (Jeffreys, 1877) (fide Pilsbry& Sharp 1897: 72).verrilli, Siphonodentalium (Siphonodentalium)Henderson, 1920: 91, 92, pl. 16, figs 2, 7. <strong>Recent</strong>.Off Nantucket [Massachusetts], Albatross USBF stn2710, 40°06’N, 68°01’30”W, 984 fms [1796 m].Holotype USNM 314844; paratype MCZ 186821(1). Pulsellum verrilli (Henderson, 1920) (fideEmerson in Turgeon 1988: 51). Western Atlantic:Massachusetts (USA). 700-1796 m.† verrucosum, Dentalium Eichwald, 1857: 584 [illustratedby Eichwald 1860: 1063, pl. 40, fig. 6].Carboniferous [“Bergkalke”]. Artinsk, Urals,Russia.† viallii, Entalina Caprotti, 1962: 97, 98, pl. 16,figs 1-3. Lower Pliocene. Castell’Arquato, Piacenza,Italy. Holotype MSNM. Remark: Pavia (1991:133) suggested that Entalina viallii might be a juniorsynonym <strong>of</strong> Entalina tetragona (Brocchi, 1815).vicdani, Fissidentalium Kosuge, 1981: 114, 115,pl. 39, figs 5-7. <strong>Recent</strong>. Bohol, Philippines. HolotypeIMT 81-36. Fissidentalium vicdani Kosuge,1981. Indo-Pacific: Philippines. 448-466 m.† vicksburgensis, Cadulus Meyer, 1885: 463; 1886: 65,pl. 3, fig. 6. Oligocene. Vicksburg, Mississippi,USA. Syntype USNM Paleobiology 644575.vietnamicus, Polyschides Chistikov, 1979a: 113, fig. 6.<strong>Recent</strong>. Tonking, Vietnam, VD-12 stn 1, 8 m.Holotype ZIN. Polyschides vietnamicus Chistikov,1979. Indo-Pacific: Indonesia, China, Vietnam,Philippines, Borneo. 8-13 m.† vinassai, Dentalium Cossmann, 1912: 215.Oligocene. Replacement name for Dentalium ancepsVinassa de Regny, 1898 [ex Meneghini MS] nonSowerby, 1837.† vincense, Dentalium (Antalis) Palmer, 1947: 209,210, pl. 26, figs 29-31. Eocene, Jackson Formation.Arkansas, USA. Holotype PRI 4468; paratypes PRI4469, PRI 4470.vincentianus, Cadulus (Gadila) Cotton & Godfrey,1940: 338, 339, fig. 360. <strong>Recent</strong>. Holdfast Bay, StVincent Gulf, South Australia. Holotype SAM D13730. Cadulus vincentianus Cotton & Godfrey,1940. Pacific Ocean: eastern <strong>and</strong> southernAustralia; Antarctica. 4-2780 m.viperidens, Cadulus Melvill & St<strong>and</strong>en, 1896: 314,pl. 11, fig. 79. <strong>Recent</strong>. Lifu <strong>and</strong> Uvea, LoyaltyIsl<strong>and</strong>s, New Caledonia, beach. Syntypes MM EE3797 (figured syntype), NMW 1955.158.594-595(6; fide Trew 1987: 70, 71). Dischides viperidens(Melvill & St<strong>and</strong>en, 1896) (fide Scarabino 1995:342). Known from the type locality only.virginalis, Cadulus (Gadila) Boissevain, 1906: 72, 73,text-fig. 35, pl. 6, figs 60-64. <strong>Recent</strong>. Indonesia,Savu Sea, Siboga stn 52, 09°03.4’S, 119°56.7’E,959 m. Lectotype ZMA 3.06.096 (designated byHabe 1964: 49; <strong>and</strong> unnecessarily by Scarabino1995: 356); paralectotypes ZMA 3.06.097 (4),ZMA 3.06.098 (3 in original description; 4 fideR. Moolenbeek & A. N. Van der Bijl in litt.23.VI.2000; B<strong>and</strong>a Sea, Siboga stn 208, 5°39’S,122°12’E, 1886 m), ZMA 3.06.099 (1 in originaldescription; 2 fide R. Moolenbeek & A. N. Van derBijl in litt. 23.VI.2000; Arafura Sea, Siboga stn 271,5°46.7’S, 134°0’E, 1788 m), ZMA 3.06.100 (2;Timor Sea, Siboga stn 284, 8°43.1’S, 127°16.7’E,828 m). Junior synonym: G. novilunata Kira, 1955(fide Habe 1963: 279). Gadila virginalis(Boissevain, 1906) (fide Scarabino 1995: 356).Indo-Pacific: Indonesia to Japan <strong>and</strong> NewCaledonia. 200-3010 m. Remark: Habe (1963,1964, 1977) included G. novilunata Kira, 1959 inthe synonymy <strong>of</strong> virginalis, but Scarabino (1995:358) stated that he “[...] prefer[s] to keep them separatepending further study”.ZOOSYSTEMA • 2004 • 26 (4)661


Steiner G. & Kabat A. R.virginianum, Dentalium Chenu, 1843: pl. 4, figs 8-8b(name <strong>and</strong> figure only). Type locality not specified.Remarks: Chenu (1852: 7), in the text, instead usedDentalium striatum Lamarck, 1818 for the specimenfigured as “D. virginianum”. However,Lamarck’s name is a junior homonym <strong>of</strong> D. striatumBorn, 1778. Pilsbry & Sharp (1898: 209) suggestedthat D. virginianum “may possibly be”Dentalium mississippiense Conrad, 1848.virginieae, Episiphon Scarabino, 1995: 288, 289,figs 93, 95a. <strong>Recent</strong>. New Caledonia, LoyaltyIsl<strong>and</strong>s, RV Alis MUSORSTOM 6, stn DW399,20°42’S, 167°00’E, 282 m. Holotype MNHN;paratypes MNHN (9), AMS C201727, NMNZM268954. Episiphon virginieae Scarabino, 1995.Known from the type locality only.virgula, Dentalium (Episiphon) Hedley, 1903: 328,fig. 62. <strong>Recent</strong>. 5.8 miles <strong>of</strong>f Port Kembla, NewSouth Wales, Australia, 63-75 fms [115-137 m].Lectotype AMS C16217 (designated by Colman1958: 144, as “holotype”); paralectotypes AMSC170639, AMS C16215 (<strong>of</strong>f Cape Three Points,New South Wales, 41-50 fms [75-91 m]), AMSC16218 (<strong>of</strong>f Wata Mooli, New South Wales, 54-59 fms [99-108 m]), AMS C62225 (<strong>of</strong>f BotanyBay, New South Wales, 50-52 fms [91-95 m]).Oldest available name for Episiphon subrectum(Jeffreys, 1883) non Dentalium sexangulum subrectaCocconi, 1873. Junior synonyms: Dentalium(Episiphon) carneum Boissevain, 1906; Dentalium(Episiphon) makiyamai Kuroda & Kikuchi, 1933(fide Scarabino 1995: 287); Dentalium yamakawaiYokoyama, 1927 (fide Habe 1964: 28, as E. makiyamai).Episiphon virgula (Hedley, 1903) (fideScarabino 1995: 287). Indo-Pacific: Africa toAustralia. 29-918 m.viridis, Dentalia Perry, 1811: pl. 52, fig. 3. <strong>Recent</strong>.South Seas. Type material not located. Junior synonym<strong>of</strong> Dentalium elephantinum Linnaeus, 1758(fide Habe 1964: 6).† vitreum, Dentalium Gmelin, 1791: 3739 [citingSchröter 1784a: 531 n. 11]. Miocene. Piedmont,Italy. Antalis vulgaris subsp. vitrea (Gmelin, 1791)(fide Pavia 1991: 122, 123, pl. 3, fig. 8); Dentaliumlaevigatum Ponzi, 1876 (tentatively, fide Sacco1897: 101).vitreum, Dentalium M. Sars, 1851: 178. <strong>Recent</strong>.Øksfjord, Norway, 100 fms [183 m]. SyntypesZMO 27067. Junior homonym <strong>of</strong> Dentalium vitreumGmelin, 1791 (Miocene <strong>of</strong> Northern Italy).Sacco (1897: footnote on p. 115) renamed Sars’taxon as Dentalium exvitreum. Now known by thefirst available name from its synonymy: Siphonodentaliumlobatum (Sowerby, 1860) (fide Pilsbry &Sharp 1897: 136). North Atlantic, Europe. 38-3100 m.† volvulus, Cadulus (Gadila?) Gardner, 1947: 629,630, pl. 57, fig. 22. Oligocene. Alum Bluff Group,Chipola Formation. USGS locality 2213, 1 milebelow Bailey’s Ferry, Chipola River, CalhounCounty, Florida, USA. Syntype USNMPaleobiology 112754 (1).† vredenburgi, Dentalium Cossmann & Pissarro,1909: 82, 83, pl. 3, fig. 23. Eocene, UppermostRanikot Series. Jhirak, India.vulgare, Dentalium Da Costa, 1778: 24, 25, pl. 2,fig. 10. <strong>Recent</strong>. Southern Engl<strong>and</strong>, Scilly Isl<strong>and</strong>s,Cornwall, Devonshire, Hampshire. Possible syntypesMHNG 1081/57 (7). Junior synonyms:Dentalium fasciatum Gmelin, 1791; ?Dentaliumnebulosum Gmelin, 1791; Dentalium tarentinumLamarck, 1818; Dentalium labiatum Turton, 1819;Dentalium laeve Turton, 1819; Dentalium striolatumRisso, 1826; Dentalium affine Biondi Giunti,1859; Dentalium vulgaris var. rosea Bucquoy,Dautzenberg & Dollfus, 1882 (fide Pilsbry & Sharp1897: 41, 42). Antalis vulgaris (Da Costa, 1778)(fide Stoliczka 1868: 438). Eastern Atlantic,Mediterranean: Europe, North Africa. 5-1000 m.† vulpeculum, Dentalium Nardo, 1847: 101 [exChiereghini MS]. Fossil. “Quarnero” [Kvarner],northwestern Croatia.vulpidens, Cadulus Watson, 1879: 524, 525 [Watson1886: 18, pl. 3, fig. 2]. <strong>Recent</strong>. Culebra Isl<strong>and</strong>,Challenger stn 24, 18°38’30”N, 65°05’30”W,390 fms [712 m]. Holotype BMNH 1887.2.9.72.Gadila vulpidens (Watson, 1879) (fide Pilsbry &Sharp 1898: 172). Western Atlantic, Caribbean.650-712 m.† vulpinum, Dentalium Nardo, 1847: 101 [exChiereghini MS]. Fossil. “Quarnero” [Kvarner],northwestern Croatia.† waccamawense, Dentalium (Antalis?) Gardner, 1948:179, pl. 24, fig. 1. Pliocene, Waccamaw Formation.Robinsons L<strong>and</strong>ing, on the Cape Fear River,Bladen County, North Carolina, USA. HolotypeUSNM Paleobiology 325479.† wadei, Dentalium Pilsbry, 1927: 142. UpperCretaceous, Ripley Formation. Dave Weeks propertyon Coon Creek, McNairy County, Tennessee,USA. Type material USNM Paleobiology 32829.Replacement name for Dentalium intercalatumWade, 1926 non Gould, 1859.† waihoraensis, Dentalium Emerson, 1954: 184.Miocene, Altonian-Clifdenian, Ihungia Series.Waihora River, Waingaromia, Gisborne, NewZeal<strong>and</strong>. Holotype NZGS TM4813; paratypesNZGS TM4814 (1), NZGS (5) (fide Keyes 1972:97). Replacement name for Dentalium (Laevidentalium)filum Marwick, 1931 non Sowerby,1860. Laevidentalium waihoraense (Emerson, 1954)(fide Maxwell 1988: 716).† waisiuense, Dentalium (Dentalium) Beets, 1943:307, 308, pl. 29, figs 113-116. Upper Oligocene.Waisiu, Buton Isl<strong>and</strong>, Indonesia.† walciodorensis, Entalis de Koninck, 1883: 215, 216,pl. 49, figs 16, 17. Carboniferous. Waulsort,Pauquys <strong>and</strong> Dréhance, Belgium. Dentalium wal-662ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)ciodorense (de Koninck, 1883) (fide Pilsbry & Sharp1898: 233).† wangwaensis, Cadulus Nomura, 1935: 93 [41], pl. 6[1], fig. 3. Pliocene. Nomura Station 14, Byoritubeds, southeast <strong>of</strong> Zyo-tusyowan, Taiwan; NomuraStation 18, Wangwa, Taiwan.† wanneri, Dentalium n. nom. Cretaceous. N <strong>of</strong> OasisDachel, Libya. Replacement name for Dentaliumbicarinatum Wanner, 1902 non Dentalium bicarinatumDeshayes, 1825.† watanabei, Dentalium (Fissidentalium) Kanno,1958: 200, 201, pl. 5, figs 7-9. Upper Oligocene toMiocene. Kanno Locality 207, riverside exposurenear a fall, Nenokami, Hikokubo, Yoshida-machi,Japan (holotype); Kanno Locality 203, riversideexposure <strong>of</strong> the Shinoha-zawa, Obashira, Chichibucity,Japan (paratype). Holotype GIUT No. 6191;paratype GIUT No. 6192.waterhousae, Fissidentalium Lamprell & Healy, 1998:94, 97, figs 92E, 93F, 98. <strong>Recent</strong>. Elizabeth Reef,Tasman Sea [<strong>of</strong>f New South Wales], 29°53.82’S,159°01.65’E, 420 m. Holotype AMS C309881;paratypes AMS C311734 (1), AMS C174647 (1).Fissidentalium waterhousae Lamprell & Healy,1998. Known from the type locality only.watsoni, Cadulus Dall, 1881: 34, 35. <strong>Recent</strong>. OffCape San Antonio, Cuba, 413 fms [754 m].Lectotype MCZ 7742 (designated by Henderson1920: 120, 121); paralectotype USNM 95380(1; Yucatan Strait, 640 fms [1168 m]). Gadila watsoni(Dall, 1881) (fide Pilsbry & Sharp 1898: 167).Western Atlantic, Caribbean: Cape Hatteras (NorthCarolina, USA) to Cuba. 370-1800 m.watsoni, Dentalium (Rhabdus) Pilsbry & Sharp,1897: 113, pl. 21, fig. 44. <strong>Recent</strong>. Localities <strong>of</strong>syntypes: USNM 107702 (<strong>of</strong>f San Diego,California, USFC stn 2923, 32°40’30”N,117°31’30”W, 822 fms [1500 m]), USNM107706 (<strong>of</strong>f Tillamook Bay, Oregon, USFC stn3346, 45°30’N, 125°52’W, 786 fms [1430 m]).Junior synonym <strong>of</strong> Rhabdus rectius Carpenter,1864 (fide Shimek 1998: 83).† waynensis, Cadulus Mansfield, 1940: 202, pl. 27,fig. 33. Oligocene. USGS station 14346, LowerChickasawhay Marl, Taylor Mill Creek, 1.5 milesN <strong>of</strong> Waynesboro, Wayne County, Mississippi,USA. Holotype USNM Paleobiology 498491.† weinbrechti, Polyschides R. Janssen, 1989: 89, 90,pl. 2, figs 19-21. Upper Miocene, Langenfeldian,Gramian <strong>and</strong> Syltian. Gram, Jyll<strong>and</strong>, Denmark.Holotype SMF 308324; paratypes SMF 308325,SMF 308326, SMF 308327/12, SMF 255698/11,RGM 229727/12, SMF 308328/5, RGM229728/5 (latter two lots from Flensburg-Weiche,Schleswig-Holstein, Germany).† weinheimensis, Antalis R. Janssen, 1989: 86-88,pl. 2, figs 14, 15. Middle Oligocene, Rupelian.Weinheim (Trift) near Alzey, Mainzer Becken,Germany. Holotype SMF 308316; paratypes SMF308317/4, SMF 308318/5, SMF 308319/4, SMF308320/3, SMF 308321/12, RGM 229726/2.weinkauffi, Dentalium Dunker, 1877: 68. <strong>Recent</strong>.Japan. Holotype ZMB 101996 (fide Kilias 1995:177). Junior synonym: Antalis septentrionalisKuroda & Habe in Habe, 1963 (fide Habe 1964:20). Antalis weinkauffi (Dunker, 1877) (fide Hirase1931: 135). Indo-Pacific: Madagascar to China <strong>and</strong>New Caledonia. 30-500 m.† weitschati, Baltodentalium Engeser & Riedel, 1992:50, 51, pl. 2, figs 3, 4, text-figs 6, 7. Jurassic,Middle Liassic, Pliensbachian. Hoisdorf, Schleswig-Holstein, Germany. Holotype GPIMH 2516/5.weldianum, Dentalium Tenison-Woods, 1877: 140.<strong>Recent</strong>. North coast <strong>of</strong> Tasmania. Holotype TMHE1200/8541. Lamprell & Healy (1998: 172)concluded that this was an unrecognizable <strong>species</strong>.wellsi, Dentalium (Dentalium) Lamprell & Healy,1998: 52, 53, figs 44D, 45D, 49. <strong>Recent</strong>. 133 kmNNW <strong>of</strong> Dampier, Western Australia, 19°28.9’-29.0’S, 116°29.4’-29.0’E, 110 m. Holotype AMSC149930; paratypes AMS C172573 (2), AMSC172572 (2; 19°00.8’S, 118°01.3’E, 157 kmNNW <strong>of</strong> Port Hedl<strong>and</strong>, Western Australia, 112 m),WAM 142-93 (4; 18°47’S, 117°58’E, 185 km NW<strong>of</strong> Port Hedl<strong>and</strong>, north Western Australia, 154 m).Dentalium wellsi Lamprell & Healy, 1998. Indo-Pacific: Western Australia. 110-168 m.wellsiana, Pulsellum (Compressidens) Kraeuter, 1972:26-28, fig. 3a-f. <strong>Recent</strong>. 1 mile W <strong>of</strong> Georgetown,Gr<strong>and</strong> Cayman Isl<strong>and</strong>s, outer slope <strong>of</strong> beach, 31-38m. Holotype ANSP 320993; paratypes ANSP300672 (5). Compressidens wellsiana (Kraeuter,1972). Known from the type locality only.† whiteavesi, Dentalium (Entalis) Anderson & Hanna,1935: 27, 28, pl. 6, fig. 5. Cretaceous. BetweenRosario <strong>and</strong> Santa Catarina L<strong>and</strong>ing, BajaCalifornia, Mexico. Holotype CAS-IP 4252.Remark: this <strong>species</strong> was misidentified as Entaliscooperi (Gabb, 1864) by Whiteaves (1879: 134;1903: 372).whitneyae, Gadila Lamprell & Healy, 1998: 152, 153,figs 155C, 156F, 159. <strong>Recent</strong>. Buchanan Isl<strong>and</strong>,southern side <strong>of</strong> Melville Isl<strong>and</strong>, near Darwin,Northern Territory, Australia, 11°49’S, 130°39’E.Holotype AMS C174636; paratypes AMSC173790 (21), AMS C173913 (19; Bowen,Queensl<strong>and</strong>), AMS C173924 (5; Facing Isl<strong>and</strong>,Port Curtis, Queensl<strong>and</strong>), AMS C173795 (1; beachbelow Crocodile research Station, Maningrida,Arnhem L<strong>and</strong>, Northern Territory), AMSC174637 (1; 32 km <strong>of</strong>f Point Charles, Darwin,Northern Territory, 12°10’S, 130°22’E, 27-37 m),AMS C173791 (1; Port Darwin, NorthernTerritory). Gadila whitneyae Lamprell & Healy,1998. Indo-Pacific: northern Australia. 0-238 m.† wilckensi, Laevidentalium Medina & del Valle,1985: 5 (Spanish), 5, 6 (English), pl. 1, figs g, h,l-o. Upper Cretaceous, López de BertodanoZOOSYSTEMA • 2004 • 26 (4)663


Steiner G. & Kabat A. R.Formation. Marambio Isl<strong>and</strong>, James Ross GroupIsl<strong>and</strong>s, Antarctica. Holotype CIRGEO (Centro deInvestigaciones en Recursos Geológicos) PI 242(now housed in Museo Argentino de CienciasNaturales, Buenos Aires, fide G. Pastorino in litt.17.I.2001).wilsoni, Dentalium Fraas, 1867. Not a scaphopod. SeeOther taxa described as <strong>Scaphopoda</strong>.† wollumbillaensis, Dentalium Etheridge in Jack &Etheridge, 1892: 483. Cretaceous. Wollumbilla,Queensl<strong>and</strong>, Australia. Neotype PIRS M2430 (designatedby Stilwell 1999: 220). Replacement namefor Dentalium lineatum Moore, 1870, a juniorhomonym <strong>of</strong> Dentalium lineatum Gueranger, 1853.Objective junior synonym: Dentalium mooreiPilsbry & Sharp, 1898. Fustiaria wollumbillaensis(Etheridge in Jack & Etheridge, 1892) (fideLudbrook 1966: 188; fide Stilwell 1999: 220).woodhousae, Cadulus Lamprell & Healy, 1998: 163,164, figs 167D, 168G, 172. <strong>Recent</strong>. Arafura Sea,64.5 km N <strong>of</strong> Croker Isl<strong>and</strong>, Northern Territory,10°17’S, 132°38’E, 65 m. Holotype AMSC174625; paratypes AMS C173679 (7). Caduluswoodhousae Lamprell & Healy, 1998. Indo-Pacific:northern Australia. 65-1380 m.woolacottae, Dentalium (Antalis) Colman, 1958: 142,fig. 4. <strong>Recent</strong>. Cronulla, New South Wales,Australia. Holotype AMS C21230; paratypes AMSC170642 (ex C21230) (1), AMS C32706 (2;Middle Harbour, Sydney, New South Wales,Australia). Dentalium woolacottae Colman, 1958.Indo-Pacific: eastern Australia. 9-40 m.† wymensis, Laevidentalium“?” Kulikov, 1967: 116,pl. 1, fig. 2. Permian. Vem River, Komi Republic,Russia. Holotype Geological Institute (St Petersbourg),8758/3.† xiphias, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 219. Eocene. Paris Basin (Parnes),France. Replacement name for Dentalium affineDeshayes, 1861 non Biondi, 1859. Objective juniorsynonym: Dentalium desmoulinsi Le Renard, 1994.Remark: a valid <strong>species</strong> <strong>of</strong> Dentalium (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 85,as “D. desmoulinsi”).yamakawai, Dentalium Yokoyama, 1927: 427, pl. 48,fig. 6. Pleistocene. Shinagawa, Tokyo [Japan].Lectotype UMT CM.23906 (designated by Taki &Oyama 1954: caption to pl. 45, as “holotype”).Junior synonym <strong>of</strong> Episiphon virgula (Hedley,1903) (fide Habe 1964: 28, as E. makiyamai).Remarks: reported from the <strong>Recent</strong> fauna <strong>of</strong> Japanby Nomura & Niino (1940: 63) <strong>and</strong> Nomura &Hatai (1940: 73, 74). Oyama (1973: 3) explained thelectotype designations <strong>of</strong> Taki & Oyama (1954).† yasilum, Dentalium Olsson, 1930: 72, pl. 12,figs 14-20. Eocene, Talara Formation. Yasila, Peru.Holotype PRI 24299; paratypes PRI 24297, PRI24298, PRI 24300-24303 (fide Brann & Kent1960: 323).† yatalensis, Cadulus (Dischides) Ludbrook, 1956: 4, 5,pl. 1, figs 3, 4. Pliocene. Weymouth’s Bore, DryCreek S<strong>and</strong>s, South Australia. Holotype SAM F15142; paratypes SAM (28). Remark: this <strong>species</strong>was misidentified as Cadulus mucronatus Tate, 1887by Ludbrook (1941).yateensis, Dischides Scarabino, 1995: 343, figs 146,147g. <strong>Recent</strong>. Yaté, New Caledonia, LAGON stn619, 22°03’S, 166°54’E, 27-42 m. Holotype <strong>and</strong>paratype MNHN. Dischides yateensis Scarabino,1995. Indo-Pacific: Philippines <strong>and</strong> New Caledonia.0-42 m.yokohamense, Dentalium Watson, 1879: 517 [Watson1886: 11, pl. 2, fig. 1]. <strong>Recent</strong>. Yokohama, Japan,Challenger stn 233, 34°39’N, 135°14’E, 8-14 fms[15-26 m]. Syntypes BMNH 1887.2.9.45-47 (3).Junior synonym <strong>of</strong> Dentalium octangulatumDonovan, 1803 (fide Habe 1964: 8).yokoyamai, Dentalium Makiyama, 1931: 44, pl. 1,fig. 1. <strong>Recent</strong>. Koshiba, Yokosuka City, Honshu,Pleistocene <strong>of</strong> Japan. Holotype KIU JC.310001(fide Hanzawa et al. 1961: 210). Fissidentaliumyokoyamai (Makiyama, 1931) (fide Kira 1959:106). Indo-Pacific: northern Australia to China Sea.56-504 m. Remark: this <strong>species</strong> was misidentified asDentalium complexum Dall, 1895 by Yokoyama(1920).† yotsukurensis, Dentalium Hirayama, 1955: 109-110,pl. 4, figs 26, 28. Upper Oligocene, Uchigo Group.Hirayama Locality A15, 1 km N <strong>of</strong> YoksukuraFishing Port, Fukushima Prefecture, Japan.Holotype GIUT 10233; paratypes GIUT 10208,10210 (Hirayama localities A28 <strong>and</strong> A45; fideHanzawa et al. 1961: 210).zanzibarense, Dentalium Plate, 1908a: 348, pl. 30,figs 35, 36. <strong>Recent</strong>. Zanzibar Channel, Valdiviastn 245, 05°28’S, 39°18’E, 463 m. Holotype ZMB61092 (fide Kilias 1995: 177). Compressidentaliumzanzibarense (Plate, 1908) (fide Scarabino 1995:262). Indo-Pacific: Africa to Madagascar. 463-1850 m.† zecaninus, Cadulus Laws, 1939: 503, pl. 63, fig. 18.Miocene. Pakaurangi Point, Kaipara Harbour, NewZeal<strong>and</strong>. Holotype NZGS TM1374 (fide Keyes1972: 98). Gadila zecanina (Laws, 1939) (fideMaxwell 1992: 187).zeidleri, Laevidentalium Lamprell & Healy, 1998:114, figs 112E, 113F, 119, 120. <strong>Recent</strong>. 29 km SE<strong>of</strong> Scam<strong>and</strong>er, Tasmania, Australia, 41°33’S,148°36’E, 122 m. Holotype SAM D 18903.Laevidentalium zeidleri Lamprell & Healy, 1998.Known from the type locality only.zel<strong>and</strong>icum, Dentalium Sowerby, 1860: 101, pl. 223,fig. 13. <strong>Recent</strong>. New Zeal<strong>and</strong>. Type material notlocated in BMNH. Junior synonym: Dentaliumpacificum Hutton, 1873 (fide Suter 1913: 819).Fissidentalium zel<strong>and</strong>icum (Sowerby, 1860) (fideSuter 1913: 819). Indo-Pacific: New Zeal<strong>and</strong>;Antarctica. 45-548 m. Remark: Dentalium opacum664ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Sowerby, 1829 may be a senior synonym for this<strong>species</strong> (fide Dell 1957: 564).† zephyrinum, Dentalium Casey, 1903: 267.Oligocene. Red Bluff, Mississippi, USA. LectotypeUSNM Paleobiology 481672 (designated byMacNeil & Dockery 1984: 247, as “holotype”).† zingulai, Cadulus (Gadila) Hodgkinson, 1974: 18, textfig.8v, w; pl. 1, fig. 11, pl. 2, fig. 6a, b. Middle Eocene,Claibornian Stage, Cook Mountain Formation,Wheelock Member. Little Brazos Bluff, Brazos RiverValley, Brazos County, Texas, USA. Holotype USNMPaleobiology 180435; paratypes USNM Paleobiology180447, PRI 29227, ANSP 31505.zonatum, Dentalium “Orb.” Paetel, 1888: 594. “Jam.”[= Jamaica]. Nomen nudum. Remark: Pilsbry &Sharp (1898: 253) stated that this <strong>species</strong> “isunknown to us”.zonatus, Cadulus (Gadila) Boissevain, 1906: 74, pl. 6,fig. 57, text-fig. 37. <strong>Recent</strong>. B<strong>and</strong>a Sea, Indonesia,Siboga stn 214, 06°30’S, 121°55’E, 2796 m.Lectotype ZMA 3.06.102 (designated by Scarabino1995: 358) ; paralectotype ZMA 3.06.103(1; Arafura Sea, Siboga stn 254, 05°40’S, 132°26’E,310 m). Gadila zonata (Boissevain, 1906) (fideScarabino 1995: 358). Indo-Pacific: Indonesia toPhilippines. 195-2796 m.OTHER TAXA DESCRIBEDAS SCAPHOPODAThe list <strong>of</strong> non-scaphopod <strong>species</strong> erroneouslydescribed or assigned to this class, as compiled byPilsbry & Sharp (1898) is reproduced here, supplementedby <strong>names</strong> subsequently removed fromthe <strong>Recent</strong> <strong>Scaphopoda</strong>. We have included thetype <strong>species</strong> <strong>of</strong> the six genera – Entalium, Falcula,Hamulus, Pharetrium, Pyrgopolon <strong>and</strong> Spirodentaliumthat Pilsbry & Sharp (1898: 244-247)excluded from the <strong>Scaphopoda</strong>. However, wehave not included any <strong>species</strong> that were subsequentlydescribed in those six genera, since theydo not enter into homonymy with valid taxa <strong>of</strong>the <strong>Scaphopoda</strong>. The entry form is that <strong>of</strong> the list<strong>of</strong> genus-<strong>group</strong> <strong>and</strong> <strong>Recent</strong> <strong>species</strong>-<strong>group</strong> <strong>names</strong>.Detailed information on type localities, typematerial deposits <strong>and</strong> synonymy is only providedfor the <strong>names</strong> removed from <strong>Scaphopoda</strong> subsequentto Pilsbry & Sharp (1898).abbreviatum, Dentalium Deshayes, 1825: 352,pl. xviii, figs 21, 22 (“1864”: 199, pl. 3, figs 5-7).Eocene <strong>of</strong> Paris. “Serpula” heptagona Sowerby (fidePilsbry & Sharp 1898: 241) (Polychaeta).aciculatum, Dentalium Hall, 1860: 107. MiddleDevonian, Marcellus Shale <strong>and</strong> Portage Groups.Manilus, New York, USA. Coleolus aciculatus, transferredto Coleolus (phylum uncertain) by Hall(1879: 190), but it may as likely be a scaphopod(fide Pilsbry & Sharp 1898: 240).acus, Dentalium Eichwald, 1857: 584. Silurian[“Grauwacke”]. Putilowa [Poulkovo], near StPetersbourg, Russia; near Erras, Estonia.Criconarida (Cephalopoda: Mollusca) (fide Kiselev2001). Remarks: figured by Eichwald, 1860: 1062,pl. 40, fig. 10. Koken (1897: 214) remarked thatEichwald’s illustration showed an unusual irregularityin its curvature (“eine Unregelmässigkeit in derBiegung”), <strong>and</strong> questioned whether this <strong>species</strong> wasreferable to the <strong>Scaphopoda</strong>. Kiselev (2001) concludedthat this <strong>species</strong> was actually from theOrdovician, <strong>and</strong> belongs to the Cephalopoda.annulata, Dentalium McClell<strong>and</strong>, 1834: 629, pl. 35,figs 1-4. Ponar River, Kamaon, northern India.Remark: the biological nature <strong>of</strong> this taxon, whichappears to be a trace <strong>fossil</strong>, cannot be determined.annulatum, Dentalium Jay, 1850: 96 [ex Mighels MS]non Dentalium annulatum Gmelin, 1791. “Maine”.Nomen nudum. Remarks: Pilsbry & Sharp (1898:240) suggested that this taxon might be a Caecum(Gastropoda). We know <strong>of</strong> no basis for suggestingthat a nomen nudum could be referable to a knowngenus.antiquum, Dentalium Goldfuss, 1841: 2, pl. 166,fig. 2. Devonian? Eifel Mountain, Germany.Coleolus antiquum (Goldfuss, 1841) (fide Yochelson2002). Yochelson (2002) concluded that this<strong>species</strong> was not a scaphopod, <strong>and</strong> suggested that itbe referred to Coleolus Hall, a calcareous “worm”tube.arietinum, Dentalium Müller, 1776: 236. Ditrupa arietina(Müller) (Polychaeta) (fide Pilsbry & Sharp1898: 242).beachportensis, Tesseracme Cotton & Ludbrook, 1938:220, pl. 12, fig. 2. Off Beachport, South Australia,201 m. Holotype SAM D 13339. Pennatulidrhachis (Cnidaria) (fide Lamprell & Healy 1998:172).belcheri, Prodentalium Nassichuk & Hodgkinson,1976: 1153-1156, pl. 1, figs 1-9. Early Permian,Assistance Formation. GSC Locality 246406, LyallRiver, Grinnell Peninsula, Devon Isl<strong>and</strong>, ArcticOcean, Canada. Holotype GSC 1377; numerousparatypes GSC 13776, 44794-44797 (Polychaeta?).Remark: Yancey (1978: 306) stated that this <strong>species</strong>“is not a <strong>species</strong> <strong>of</strong> Prodentalium, <strong>and</strong> probably nota scaphopod” <strong>and</strong> concluded that “it is probablethat it is not a molluscan <strong>species</strong>, but a calcareousworm tube”.billingsi, Salterella Safford, 1869: 289. Ordovician <strong>of</strong>Tennessee. Transferred to the <strong>Scaphopoda</strong> byZOOSYSTEMA • 2004 • 26 (4)665


Steiner G. & Kabat A. R.Fisher (1958: 144-146). Yochelson (1968) concludedthat this taxon was not scaphopod <strong>and</strong> was probablyreferable to the Hyolitha.brazieri, Cadulus Finlay, 1927: 521. Princess CharlotteBay, North East Australia, 23 m. AMS. Replacementname for Dentalium laeve Brazier, 1877 (fideEmerson 1954: 186). Both Finlay (1927: 521) <strong>and</strong>Bretnall (1921: 156) independently renamedBrazier’s laeve, <strong>and</strong> both independently used thesame replacement name, but transferred the <strong>species</strong>into different genera. Finlay named it Cadulus brazieri;while Bretnall had already renamed it Ditrypa[sic] brazieri. Thus, Finlay’s replacement name isboth an objective junior synonym <strong>and</strong> a secondaryhomonym <strong>of</strong> Bretnall’s name. Ditrupa gracillimaGrube, 1878 (Polychaeta, Serpulida) (fide ten Hove &Smith 1990: 107; fide Lamprell & Healy 1998: 172).browni, Dentalium Hisinger, 1837: 21, pl. iv, fig. 9.Köpingemölla, Scania [Sweden]. Types inSMNH(?). Polychaeta (fide Pilsbry & Sharp 1898:246).bulbosum, Dentalium Bronn, 1831: 85 [nomen nudumin Bronn 1828: 538]. Tertiary <strong>of</strong> Italy. Polychaeta(fide Pilsbry & Sharp 1898: 242.carinatum, Dentalium O. G. Costa, 1851: 24. Mare diTaranto [Mediterranean Sea]. Polychaeta (fidePilsbry & Sharp 1898: 242).cinctum, Dentalium de Koninck, 1843: 318, pl. 23,fig. 3. Carboniferous, Belgium. Orthoceras subcentrale(de Koninck, 1843) (Cephalopoda) (fidePilsbry & Sharp 1898: 241).cingulatus, Dentalites Schlotheim, 1820: 94.Polychaeta (fide Pilsbry & Sharp 1898: 242).clausum, Dentalium Turton, 1819: 39. Calves Isl<strong>and</strong>,Irel<strong>and</strong>. Quill <strong>of</strong> a seabird’s wing feather(Vertebrata) (fide Pilsbry & Sharp 1898: 247; citingForbes & Hanley, <strong>and</strong> Jeffreys).clava, Dentalium Lamarck, 1818: 346. Fossil fromCypli, near Mons [Belgium]. Polychaeta (fidePilsbry & Sharp 1898: 246).coarctatum, Dentalium Brocchi, 1814: 628. Ditrupa(Polychaeta) (fide Pilsbry & Sharp 1898: 242).compressum, Dentalium d’Orbigny, 1850: 233.Jurassic. Chalon-sur-Saône, France. Polychaeta.Remark: Richardson (1906: 592, 593) concludedthat this <strong>species</strong> “is not a Dentalium at all, but theshell <strong>of</strong> a tubicolous annelid”.corneum, Dentalium Linnaeus, 1767: 1263 [no referencesgiven]. “O. Africano”. Ditrupa arietina(Polychaeta) (fide Pilsbry & Sharp 1898: 242).“cornicula, Dentalium” – d’Orbigny 1852: index,p. 59. Indexing error for Dentalina cornicula(Foraminifera) (fide Pilsbry & Sharp 1898: 247).corniculum, Dentalium O. G. Costa, 1851: 55, pl. 4,figs 2a-A. Caecidae (Gastropoda) (fide Pilsbry &Sharp 1898: 241).crassum, Dentalium Deshayes, 1825: 373, pl. 18,fig. 20. Fossil near Mons [Belgium]. Polychaeta(fide Pilsbry & Sharp 1898: 246).crenatocinctum, Coleolus Hall, 1879: 188, 189, pl. 23,figs 1-3, pl. 23A, figs 3, 4. Upper HelderbergFormation, Jamesville, Ontario County, New York.Phylum uncertain. Considered a Dentalium byWhitfield (fide Pilsbry & Sharp 1898: 241).cylindraceum, Dentalium O. G. Costa, 1851: 39, pl. 3,fig. 10. Pliocene near Caramanico [Italy]. Ditrupa(Polychaeta) (fide Pilsbry & Sharp 1898: 242).cylindrica, Hensonella Elliott, 1960: 229, 230, pl. 8,fig. 1. Lower Cretaceous, Surdash, Sulemania Liwa,Iraq. Syntypes BMNH (Palaeontology) No. Z.902.Remarks: this <strong>species</strong> was also recorded from theLower Cretaceous <strong>of</strong> Iraqi Kurdistan <strong>and</strong> at Kirkuk,as well as northern Iran. Elliot (1960) removed the<strong>species</strong> from the dasyclad alga <strong>and</strong> tentatively placedit in the <strong>Scaphopoda</strong>. The microstructure <strong>of</strong> thetube does not resemble that <strong>of</strong> scaphopod shells,therefore it is unlikely to belong to this taxon.cylindricum, Dentalium J. Sowerby, 1814: 179, 180,pl. 79, fig. 2. Eocene. Emsworth, Engl<strong>and</strong>. Ditrupa(Polychaeta) (fide Pilsbry & Sharp 1898: 242).Remark: Gardner (1878: 57, 61) noted that thetype locality was given in error as “Exmouth” <strong>and</strong>hence this <strong>species</strong> was mistakenly assumed to beCretaceous, <strong>and</strong> not Eocene.deforme, Dentalium Lamarck, 1818: 344. Fossil,Sarthe, France. Polychaeta (fide Pilsbry & Sharp1898: 243).deshayesianum, Dentalium Galeotti, 1837: 62, 150,pl. 4, fig. 7. Tertiary. Belgium (numerous localities).Polychaeta (fide Pilsbry & Sharp 1898: 242).Remark: Galeotti (1837: 192), under “Corrections”,listed his new <strong>species</strong> as a junior synonym <strong>of</strong>Dentalium incrassatum J. Sowerby, 1814 (Polychaeta).difforme, Dentalium J. de C. Sowerby in Dixon, 1850:348, pl. xxix, fig. 10. Sussex, Engl<strong>and</strong>. Polychaeta(fide Pilsbry & Sharp 1898: 243).duplicatum, Dentalium Blainville, 1825: 628.Polychaeta (fide Pilsbry & Sharp 1898: 245, 246).Remark: Pilsbry & Sharp (1898: 245) explainedthat Blainville, in using “Entale” Defrance[= Entalium Defrance, 1819], changed the name <strong>of</strong>the type <strong>species</strong>, “for some occult reason”, fromEntalium rugosum to Dentalium duplicatum.ecostatum, Dentalium Kirk, 1880: 306, 307.Waikanae, New Zeal<strong>and</strong>. Although Dell (1957:566) <strong>and</strong> Powell (1976: 121) considered this to be ascaphopod, Marshall (1996: 32) concluded that itwas the shell <strong>of</strong> a planktonic gastropod, Creseis acicula(Rang, 1829) (Cavoliniidae).edoense, Dentalium Tokunaga, 1906: 34, 35, pl. 2,fig. 17. Shinagawa <strong>and</strong> Oji, Tokyo [Pleistocene <strong>of</strong>Japan]. Ditrupa arietina (Polychaeta) (fide Habe1963: 12).falcatum, Dentalium Conrad, 1869: 44, pl. 1, figs 12,16. Cretaceous. Crosswicks, New Jersey [USA].Polychaeta (fide Pilsbry & Sharp 1898: 243).florenciae, Dentalium Moraes Rego, 1936: 43-46, textfigure.Corumbataí Formation (Triassic?), Guareí,666ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)southern Sao Paulo province, Brasil. Holotype(Paleontology Section, Departamento Nacional daProdução Mineral (DNPM), Rio de Janeiro,Brasil). Remark: Wehmuth Ragonha & da SilvaSantos (1987: 2) determined that this <strong>fossil</strong> wasactually an external impression <strong>of</strong> a dorsal fin-spine<strong>of</strong> Hybodus, a <strong>fossil</strong> shark (elasmobranch).fragile, Pharetrium König, 1825: 4, pl. 7, fig. 80.Cretaceous. Mount San Petri (Italy?). Polychaeta(fide Pilsbry & Sharp 1898: 245, 246).giganteum, Dentalium Phillips, 1829: 164, pl. 14,fig. 8; 1835: 136, pl. 14, fig. 8. Jurassic, Liassic.Marlstone, Yorkshire, Engl<strong>and</strong>. Holotype YorkshireMuseum 1998/718. Ditrupa-like serpulid polychaete(fide Palmer 2001: 253-255). Remark: this<strong>species</strong> was originally described as Dentalia giganteum.glabrum, Dentalium Montagu, 1803: 497. Caecumglabrum (Montagu) (Gastropoda) (fide Pilsbry &Sharp 1898: 241).gordonis, Cadulus Yokoyama, 1920: 104, pl. 6, figs 25,26. Miyata <strong>and</strong> Naganuma Zones [Pliocene <strong>of</strong>Japan]. Lectotype UMT CM.20329 (designated byTaki & Oyama 1954: caption to pl. 7, as “holotype”);paralectotype UMT CM.20328. Ditrupaarietina (Polychaeta) (fide Habe 1963: 12). Remark:Oyama (1973: 3) explained the lectotype designations<strong>of</strong> Taki & Oyama (1954).goreanum, Dentalium Clessin, 1896: 42, pl. 10, figs 9,10. Gorée [Senegal]. Polychaeta (fide Pilsbry &Sharp 1898: 243).hamatum, Dentalium? Forbes, 1846: 138, pl. xv, fig. 8.Trichinopoli [Tiruchchirappalli, Cretaceous <strong>of</strong>southern India]. Polychaeta (fide Pilsbry & Sharp1898: 243).“hamatus”, Dentalium – Conrad 1870b: 77. Error forDentalium falcatus Conrad, 1869 (Polychaeta) (fidePilsbry & Sharp 1898: 244, 245).howardensis, Clavulites Girty, 1904: 732, 733, pl. 47,figs 4-11. Burlingame Shale, Howard, Kansas, USA(Carboniferous). Type material USNM Paleontology35134. Tentatively placed in the serpulidpolychaetes by Yochelson (1971).imperforatum, Dentalium G. Adams, 1798: 635,pl. 14, fig. 8 [ex E. Jacob MS]. S<strong>and</strong>wich, Engl<strong>and</strong>,United Kingdom. This taxon appears to be a juvenileCaecum (Gastropoda) or possibly even a microscopicProtista. It is likely that the usage <strong>of</strong> thisname by Montagu (1803: 496) <strong>and</strong> by Turton(1819: 39) for a <strong>species</strong> <strong>of</strong> Caecum (fide Pilsbry &Sharp 1898: 241) were merely reiterations <strong>of</strong>G. Adams’ name, <strong>and</strong> not new <strong>species</strong>.incrassatum, Dentalium J. Sowerby, 1814: 180, pl. 79,figs 3, 4. Polychaeta (fide Pilsbry & Sharp 1898:243).incurvum, Dentalium Renier, 1804: xii, xx. Polychaeta(fide Pilsbry & Sharp 1898: 243). Remark: Renier’spublication was placed on the Official Index <strong>of</strong>Rejected <strong>and</strong> Invalid Works in Zoological Nomenclature(ICZN 1954, Opinion 316), so this name isnot available.indistinctum, Dentalium Fleming, 1825: 241, pl. ix,fig. 2. West Lothian [Scotl<strong>and</strong>], Carboniferous.Polychaeta (fide Pilsbry & Sharp 1898: 243).inornatum, Dentalium McCoy in Griffith, 1844: 47,pl. 5, fig. 30. Carboniferous. Irel<strong>and</strong>. Possible typematerial National Museum <strong>of</strong> Irel<strong>and</strong>, NMINGF19087, NMING F4983 <strong>and</strong> NMING 19806 (fideHoll<strong>and</strong> & Yochelson 2000). “Ortoceras” (Cephalopoda)(fide Holl<strong>and</strong> & Yochelson 2000). Remark:Holl<strong>and</strong> & Yochelson (2000) stated that “we suggestthat [this taxon] be confined to use with the type materialin the National Museum <strong>of</strong> Irel<strong>and</strong> noted above”.intestiniforme, Dentalium Hanley, 1860: 89 [49] [exLinnaeus MS]. Published in synonymy <strong>of</strong> Vermetusgigas. Thylacodes polyphragma (Sassi, 1827)(Gastropoda: Vermetidae) (fide Mörch 1862: 66;Pilsbry & Sharp 1898: 241).jungii, “Dentalium” – d’Orbigny 1852: index, p. 59.Lapsus for Fusus jungii (fide Pilsbry & Sharp 1898:241).laeve, Dentalium Brazier, 1877: 59. Darnley Isl<strong>and</strong>,Torres Strait [northern Australia]. Lectotype AMSC7525 (designated by Ponder & Stanbury 1972:53) <strong>and</strong> paralectotypes (from Darnley Isl<strong>and</strong> <strong>and</strong>elsewhere). Junior homonym <strong>of</strong> Dentalium laeve(Schlotheim, 1820). Replacement name: Ditrypa[sic] brazieri Bretnall, 1921 (= Ditrupa gracillimaGrube, 1878 (Polychaeta, Serpulida) (fide ten Hove& Smith 1990: 107; fide Lamprell & Healy 1998:172). Remark: Finlay’s (1927: 521) replacementname for Dentalium laeve Brazier, 1877, Cadulusbrazieri, is both an objective junior synonym <strong>and</strong> asecondary homonym <strong>of</strong> Bretnall’s name.ludbrooki, Dentalium (Gadilina) Caprotti, 1962: 96,97, pl. 16, figs 4-6. Pliocene. Castell’Arquato,Piacenza, Italy. Holotype MSNM. Cuvierina jagtiJanssen, 1995 (Gastropoda) (fide Janssen 1999).Remarks: Janssen (1999) concluded that Caprotti’s<strong>species</strong> was actually the shell <strong>of</strong> a planktonic pteropod(Gastropoda, Euthecosomata). Janssen alsonoted that Caprotti’s type material was not in theMuseo Civico di Storia Naturale (Milano), contraryto the original description.martini, Dentalium Whitfield, 1882: 203. UpperHelderberg, near Dublin, Ohio [USA]. Coleolus crenatocinctumHall, 1879 (phylum uncertain) (fideYochelson & Goodison 1999: 634-640).mosae, Pyrgopolon Montfort, 1808: 394-396, text-fig.Cretaceous. Maastricht, Netherl<strong>and</strong>s. Polychaeta(fide Pilsbry & Sharp 1898: 245, 246).nigr<strong>of</strong>asciatum, Dentalium Eichwald, 1830: 199.Pliocene, Zukowce, Volhynia [Ukraine]. Polychaeta(fide Pilsbry & Sharp 1898: 243).nigrum, Dentalium Lamarck, 1818: 345, 346.Polychaeta (fide Pilsbry & Sharp 1898: 243).octocostatum, Dentalium Fraas, 1867a: 239, pl. 4,fig. 13; 1867b: 95, pl. 1, fig. 13. Cretaceous. Marsaba,ZOOSYSTEMA • 2004 • 26 (4)667


Steiner G. & Kabat A. R.Palestine. Polychaeta (fide Pilsbry & Sharp 1898:245). Remark: Blanckenhorn (1927: 123, 124, pl.1, figs 2-8) considered this to be a valid <strong>species</strong> <strong>of</strong>Dentalium, but he did not discuss the contraryconclusion <strong>of</strong> Pilsbry & Sharp.onyx, Hamulus Morton, 1834: 73, 74, pl. 2, fig. 8,pl. 16, fig. 5. Cretaceous. Lynch’s Creek, SouthCarolina <strong>and</strong> Erie, Alabama, USA. Polychaeta (fidePilsbry & Sharp 1898: 245).opiparium, Quasidentalium Shimansky, 1974: 135,figs 2a-2c. Lower Carboniferous, Donets Basin[Russia]. Coleolus (fide Yochelson 1999: 63-65).osceola, Spirodentalium Walcott, 1890: 271, 272,pl. 20, fig. 12. Upper Cambrian, Potsdam Terrane.Osceola Mills, Wisconsin, USA. Lectotype USNMPaleobiology 23853a (selected by Yochelson 1987:66, 67, fig. 1.4); two paralectotypes USNMPaleobiology 23853b, 23853c. Remark: Yochelson(1987) concluded that this taxon was actually anopen-coiled gastropod, which he referred to theEuomphaloidea.pellucidum, Dentalium Gmelin, 1791: 3738 [citingSchröter 1783-1786: 529]. Oceano septentrionali[northern ocean]. Polychaeta (fide Pilsbry & Sharp1898: 243, pl. 6, fig. 17).planum, Dentalium J. Sowerby, 1814: 179, pl. 79,fig. 1. Bognor, Engl<strong>and</strong>. Polychaeta (fide Pilsbry &Sharp 1898: 243).pusillum, Dentalium Philippi, 1836: 245. Panormi,Catania [Sicily]. “Has been referred to Dischidespolitus, but it may be a Ditrupa” (Pilsbry & Sharp1898: 243).pygmaeus, Dentalium Defrance, 1819a: 71. Caecum(Gastropoda) (fide Pilsbry & Sharp 1898: 241).radicula, Dentalium Lamarck, 1818: 345. Eocene.Grignon, France. Polychaeta (fide Pilsbry & Sharp1898: 243).rudis, ?Dentalium Gabb, 1873: 244. San Domingo[Dominican Republic]. Polychaeta (fide Pilsbry &Sharp 1898a: 474, pl. 10, figs 4, 8; Pilsbry & Sharp1898: 243).rugosum, Dentalium Müller, 1851: 6, pl. 3, fig. 2.Cretaceous. Lusberg, Germany. Gastrochaena(Bivalvia) (fide Holzapfel 1888: 179). Remarks:Pilsbry & Sharp (1898: 227) initially suggested thatthis <strong>species</strong> might belong to the Serpulidae[Polychaeta] or Teredinidae [Bivalvia]; in theiraddendum, Pilsbry & Sharp (1898: 255) thennoted that Holzapfel (1888) had already placed thistaxon in the Bivalvia.rugosum, Entalium Defrance, 1819b: 518. Cretaceous.Saint-Pierre, near Maastricht, Netherl<strong>and</strong>s.Polychaeta (fide Pilsbry & Sharp 1898: 245).septangulare, Dentalium Fleming, 1825: 240, pl. 9,fig. 1. Belfast [Irel<strong>and</strong>]. Probably “Serpula” heptagonaSowerby (Polychaeta) (fide Pilsbry & Sharp1898: 243).septemcostatum, Dentalium Abich in Trautschold,1859: 314, 315, pl. 6, fig. 5. Eocene <strong>and</strong> Oligocene<strong>of</strong> Armenia. Probably “Serpula” heptagona Sowerby(Polychaeta) (fide Pilsbry & Sharp 1898: 244).serratum, Dentalium Pictet & Roux, 1849: 150, pl. 27,fig. 12a, b. Cretaceous, Bouches-du-Rhône, France.Probably belongs to Hamulus (Polychaeta) (fidePilsbry & Sharp 1898: 244).sexcarinatum, Dentalium Goldfuss, 1841: 4, pl. 166,fig. 12. Cretaceous. Friedl<strong>and</strong>, Silesia (Pomerania?),Germany; St Petri, Maastricht, Netherl<strong>and</strong>s.Polychaeta (fide Pilsbry & Sharp 1898: 244).sexradiatum, Dentalium Goldfuss – Pilsbry & Sharp1898: 244. “Craie super. Maestricht”. Nomennudum. May be an incorrect subsequent spelling <strong>of</strong>sexcarinatum Goldfuss, 1844 (fide Pilsbry & Sharp1898: 244). Remark: Pilsbry & Sharp (1898: 244)state that they “have seen no work in which this<strong>species</strong> is described”.sowerbyi, Dentalium Michelotti, 1847: 145. Miocene,Italy. Junior homonym <strong>of</strong> Dentalium sowerbyiGuilding, 1834. Polychaeta (fide Pilsbry & Sharp1898: 244).spinulosum, Dentalium Miller, MSS – Pilsbry & Sharp(1898: 241) cited this name in the synonymy <strong>of</strong>.Hamites spinulosus J. Sowerby, 1818 (Cephalopoda).spirale, Dentalium Risso, 1826b: 125; 1826b: 401.(Fossil). France. Type material presumed lost (fideArnaud 1978: 135). Polychaeta. Remark: Arnaud(1978: 135) noted that Risso’s unpublished illustration<strong>of</strong> the type specimen showed spiral ornamentationthat is unknown in the <strong>Scaphopoda</strong>.strangulatum, Dentalium Deshayes, 1825: 372, 373,pl. 16, fig. 28. Australian Sea?, <strong>fossil</strong> in Engl<strong>and</strong>,France, <strong>and</strong> Italy. Polychaeta (fide Pilsbry & Sharp1898: 244). Dentalium strangulosum “Deshayes”[Gümbel, 1861: 604] is an incorrect subsequentspelling.strangulosum, see strangulatum.“subcarinatum”, Dentalium “Münster in Goldfuss” –Ryckholt 1851. May be an incorrect subsequentspelling <strong>of</strong> Dentalium sexcarinatum Münster inGoldfuss, 1844 (fide Pilsbry & Sharp 1898: 244).subtilis, Cadulus (Gadila) Plate, 1908a: 360, pl. 30,fig. 48. <strong>Recent</strong>. Near Dar-es-Salaam, Tanzania,Valdivia stn 244, 5°55.8’S, 39°1.2’E, 50 m. LectotypeZMB 61108a (designated by Kilias 1995:176); paralectotypes ZMB 61108b (1), ZMB63842 (4). Remarks: Jaeckel (1932: 310) remarkedupon certain similarities with Ditrupa (Polychaeta),<strong>and</strong> Kilias (1995: 176) parenthetically remarkedthat this taxon is probably Ditrupa. Plate’s illustrationindeed resembles a worm tube, <strong>and</strong> the taxonmay have to be removed from the <strong>Scaphopoda</strong> afterexamination <strong>of</strong> the types.subulatum, Dentalium Deshayes, 1825: 373, pl. 16,fig. 29. Type locality not specified. Polychaeta (fidePilsbry & Sharp 1898: 244).tiwhana, Dentalium Dell, 1953: 48, figs 17, 22.Chatham Rise, New Zeal<strong>and</strong>, 361 m. Serpula crenata668ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)(Ehlers, 1908) (Polychaeta, Serpulida) (fide tenHove & Smith 1990: 107; fide Dell 1995: 21).trachea, Dentalium Montagu, 1803: 497, pl. 14,fig. 10. Caecum (Gastropoda) (fide Pilsbry & Sharp1898: 241).tricostatum, Dentalium Goldfuss, 1841: 3, 4, pl. 166,fig. 11a, b. Cretaceous. Essen, Ruhr Valley,Westphalia [Germany]. Polychaeta (fide Pilsbry &Sharp 1898: 246).typa, Throopella Greger, 1933: 373, 374, pl. 21. UpperDevonian, Cow Creek stage. Snyder Creek, centralMissouri, USA. Emerson (1962: 480) <strong>and</strong>Starobogatov (1974: 8) excluded this taxon fromthe <strong>Scaphopoda</strong>. Starobogatov (1974) placed it inthe toxeumorphorid Xenoconchia.undatum, Dentalium Defrance, 1819a: 72. Saint-Clément, near Angers [France]. Ditrupa (Polychaeta)(fide Pilsbry & Sharp 1898: 244).vayssierei, Dentalium Fenaux, 1942: 3, text-fig. 5.<strong>Recent</strong>, 80 m, <strong>of</strong>f Carro, France (MediterraneanSea). Two syntypes (presumably in MOM).Polychaeta (fide V. Scarabino, in litt. 5.XI.2001).wilsoni, Dentalium Fraas, 1867a: 239, pl. 4, fig. 12;1867b: 95, pl. 1, fig. 12. Cretaceous. Marsaba,Palestine. Polychaeta (fide Pilsbry & Sharp 1898:246). Blanckenhorn (1927: 123, 124, pl. 1, figs 2-8) listed this as a junior synonym <strong>of</strong> Dentalium octocostatumFraas, 1867, which he treated as a valid<strong>species</strong> <strong>of</strong> Dentalium, but he did not discuss thecontrary conclusion <strong>of</strong> Pilsbry & Sharp.AcknowledgementsThe first author is most indebted to P. Bouchet,B. Métivier (MNHN), S. G<strong>of</strong>as (University <strong>of</strong>Malaga), J. Taylor, K. Way, J. Pickering (BMNH)<strong>and</strong> D. Ivanov (MMSU) for making it possible towork at their institutions <strong>and</strong> for their helpwith the collections. I owe special thanks toV. Scarabino (Paris) for providing a wealth <strong>of</strong>information <strong>and</strong> help.The second author thanks K. J. Boss (MCZ) <strong>and</strong>M. G. Harasewych (USNM) for allowing him touse their departmental libraries, T. R. Waller <strong>and</strong>J. Thompson (USNM Paleobiology) for allowinghim to examine the <strong>fossil</strong> type material in theircollections, V. Héros (MNHN) for her help withthe Dautzenberg type localities, <strong>and</strong> R. I. Johnson(MCZ) for his fine hospitality <strong>and</strong> access to hispersonal library.The following colleagues provided us with valuableinformation on type material <strong>and</strong> synonyms, or sentus copies <strong>of</strong> obscure publications: A. Baldinger(MCZ), A. Beu (NZGS), R. Bieler (FMNH),E. Brunstad (Bergen, Norway), P. Callomon(Japan), K. Chinzei (FGK), G. Coan (CAS),A. Dhondt (KBINW), R. Eng (WSM), Y. Finet(MHNG), D. Geiger (SBMNH), B. Goddeeris(KBINW), D. Graf (ANSP), L. Groves (LACM),R. Janssen (SMF), E. Kools (CAS), S. Kosuge(IMT), K. B. Kvalsvik (ZMO), R. La Perna (Univ.Catania, Italy), A. Matsukuma (KyushuUniversity), R. Moolenbeek <strong>and</strong> A. N. Van derBijl (ZMA), G. Pastorino (Museo Argentino deCiencias Naturales), R. E. Petit (North MyrtleBeach, South Carolina), M. Recevik (MCZ),G. Rosenberg (ANSP), H. Saito (NSMT),K. Sindemark (SMNH), J. A. Sneli (TrondheimBiological Station, Norway), J. Stilwell (JCU),R. Symonds (UMZC), S. Takeuchi (SHM),F. Wesselingh (RMNH), <strong>and</strong> R. White (YPM).We most appreciate the reviews provided byPhilippe Bouchet <strong>and</strong> Victor Scarabino, whichenhanced the accuracy <strong>and</strong> presentation <strong>of</strong> thiscatalog. Helmut Zibrowius (Centre d’Océanologiede Marseille) provided helpful commentson various <strong>species</strong> that were originally describedas scaphopods but subsequently determined to benon-molluscan. 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P. 1839-1853. — Traité élémentaire deconchyliologie avec les applications de cette science à lagéologie. Explication des planches. Victor Masson,Paris, 80 p. + 132 pls Notes: p. 1-8 (1839); p. 9-48(1850); p. 49-80 (1853).DESHAYES G. P. 1861 [30 July] [in 1861-1864]. —Description des animaux sans vèrtebres découverts dansle Bassin de Paris pour servir de supplément à ladescription des coquilles <strong>fossil</strong>es des environs de Pariscomprenant une revue générale de toutes les espècesactuellement connues. Vol. 2. J. B. Baillière et fils,Paris, 968 p., pls 1-62.DE STEFANI C. 1884. — Escursione scientifica nellaCalabria (1877-78), Jejo, Montalto e Capo Vaticano.Studio geologico. Atti della Reale Accademia deiLincei (Anno CCLXXX 1882-83), Serie Terza,Memorie della Classe di scienze fisiche, matematiche enaturali 18: 290 p.DE STEFANI C. 1917 [15 December]. — Fossili carboniferidell’Isola d’Elba. 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Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Akademie der wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse 1933 (11): 101-102.GUGENBERGER O. 1933b. — Die Cardita-Schichtenvon Launsdorf in Mittelkärnten und ihre Fauna. II.Gastropoda. Akademie der wissenschaften in Wien,Mathematisch-Naturwissenschaftliche Klasse,Sitzungsberichte, Abteilung I, Mineralogie, Biologie,Erdkunde 142 (3-4): 157-184, pl. 1.GUGENBERGER O. 1934. — Die Cardita-Schichtenvon Launsdorf in Mittelkärnten und ihre Fauna.III. Scaphopoden; IV. Cephalopoden. Akademie derwissenschaften in Wien, Mathematisch-NaturwissenschaftlicheKlasse, Sitzungsberichte, Abteilung I,Mineralogie, Biologie, Erdkunde 143 (1-2): 41-59.GUILDING L. 1834. — Observations on Naticina <strong>and</strong>Dentalium, two genera <strong>of</strong> molluscous animals.Transactions <strong>of</strong> the Linnean Society <strong>of</strong> London 17 (1):29-35, pl. 3.GÜMBEL C. W. 1861. — Geognostische Beschreibungdes Königreichs Bayern, Abtheilung I, GeognostischeBeschreibung des bayerischen Alpengebirges und seinesVorl<strong>and</strong>es. Justus Perthes, Gotha, xx + 950 p.,42 pls.GÜMBEL C. W. 1866 [“1865”]. — Über das Vorkommenvon unteren Triasschichten in Hochasien.Sitzungsberichte der königlichen bayerisches Akademieder Wissenschaften zu München 1865 (II)(4): 348-366, 1 pl.GUPPY R. J. L. 1866 [1 August]. — On the TertiaryMollusca <strong>of</strong> Jamaica. 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Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)HODGKINSON K. A. 1974 [10 July]. — Stone City <strong>and</strong>Cook Mountain (Middle Eocene) scaphopods fromsouthwest Texas. University <strong>of</strong> Kansas,Paleontological Contributions 70: 1-25, pls 1-8.HOENINGHAUS F. W. 1831. — Versuch einer geognostischenEintheilung seiner Versteinerung-Sammlung (Dritter Theil. Tertiär-Gebirge). Jahrbuchfür Mineralogie, Geognosie, Geologie und Petrefaktenkunde1831 (2): 132-170.HOENINGHAUS F. W. 1836a. — Dentalium saturni.Krefeld: private publication, 1 p., 1 pl. [see Langer1967 for discussion <strong>of</strong> publication].HOENINGHAUS F. W. 1836b. — [Abbildung einesDentalium]. Mittheilungen aus den Verh<strong>and</strong>lungender Gesellschaft naturforschender Freunde zu Berlin1836 (1): 12.HOERNES R. 1875. — Die Fauna des Schliers vonOttnang. Jahrbuch der kaiserlich-königlichen geologischenReichstanstalt, Wien 25 (4): 333-400 [1-68],pls 10-15.HOLLAND C. H. & YOCHELSON E. L. 2000. — Dentaliuminornatum McCoy, 1844, is not a scaphopod.Irish Journal <strong>of</strong> Earth Sciences 18: 123-125.HOLZAPFEL E. 1887-1888. — Die Mollusken derAachener Kreide. Palaeontographica 34 (1): 29-72,pls 4-5 [July 1887]; 34 (2-4): 73-180, pls 6-21[March 1888].HÖLZL O. 1962. — Die Molluskenfauna der oberbayerischenmarinen Oligozänmolasse zwischen Isarund Inn und ihre stratigraphische Auswertung.Geologica Bavarica 50: 1-275, 12 pls.HÖRNES M. 1856. — Die Fossilen Mollusken desTertiär-Beckens von Wien unter der Mitwirkungvon Paul Partsch. Abh<strong>and</strong>lungen der Kaiserlich-Königlichen Geologischen Reichsanstalt, Wien, vol. 3,I. B<strong>and</strong>: Univalven: 736 p.HOVE H. A. TEN & SMITH R. 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Renier commonly attributed tothe year 1804. Opinions <strong>and</strong> Declarations renderedby the International Commission on ZoologicalNomenclature 9 (5): 91-106.INTERNATIONAL COMMISSION ON ZOOLOGICALNOMENCLATURE 1955. — Opinion 361: Validationunder the plenary powers <strong>of</strong> the generic nameAntalis Adams (H.) & Adams (A.) [1854] (Class<strong>Scaphopoda</strong>). Opinions <strong>and</strong> Declarations rendered bythe International Commission on Zoological Nomenclature11 (11): 161-172.INTERNATIONAL COMMISSION ON ZOOLOGICALNOMENCLATURE 1957. — Direction 73: Additionto the Official List <strong>of</strong> Specific Names in Zoology.Opinions <strong>and</strong> Declarations rendered by theInternational Commission on Zoological Nomenclature1 (E) (E 12): 193-214.ITOIGAWA J. 1960 [December]. — Paleoecologicalstudies <strong>of</strong> the Miocene Mizunami Group, centralJapan. Journal <strong>of</strong> Earth Sciences, Nagoya University8(2): 246-300, pls 1-6.IVANOV A. 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Steiner G. & Kabat A. R.Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition 1898-1899 21 (2): 301-315.JAMES J. F. 1890 [June]. — On the Maquoketa shales,<strong>and</strong> their correlation with the Cincinnati <strong>group</strong> <strong>of</strong>southwestern Ohio. The American Geologist 5 (6):335-356.JANSSEN A. W. 1984. — Mollusken uit het Mioceen vanWinterswijk-Miste. Een inventarisatie, met beschrijvingenen afbeeldingen van alle aangetr<strong>of</strong>fen soorten.Koninklijke Nederl<strong>and</strong>se NatuurhistorischeVereniging, Leiden, 451 p., 82 pls.JANSSEN A. W. 1999 [22 December]. — Notes on thesystematics, morphology <strong>and</strong> biostratigraphy <strong>of</strong> <strong>fossil</strong>holoplanktonic Mollusca, 5. Cuvierina jagtiJanssen, 1995: a junior synonym <strong>of</strong> Dentalium(Gadilina) ludbrooki Caprotti, 1962. 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Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)MONTEROSATO T. A. DI 1884. — Nomenclatura genericae specifica di alcune Conchiglie Mediterranee.Stabilimento Tipografico Virzi, Palermo, 152 p.MONTFORT P. DENYS DE 1808 [in 1808-1810]. —Conchyliologie systématique, et classification méthodiquedes coquilles... Tome Premier. F. Schoell,Paris, lxxxviii + 410 p.MOORE C. 1866. — On the Middle <strong>and</strong> Upper Lias<strong>of</strong> the South West <strong>of</strong> Engl<strong>and</strong>. Proceedings <strong>of</strong> theSomersetshire Archaeological <strong>and</strong> Natural HistorySociety 13: 119-230, 7 pls.MOORE C. 1870 [2 May]. — Australian Mesozoicgeology <strong>and</strong> palaeontology. Quarterly Journal <strong>of</strong>the Geological Society <strong>of</strong> London 26 (2): 226-261,pls 10-18.MOORE E. J. 1963. — Miocene marine mollusks fromthe Astoria Formation in Oregon. United StatesGeological Survey Pr<strong>of</strong>essional Paper 419: iv + 109 p.,33 pls.MORAES REGO L. F. DE 1936 [31 March]. — Contribuiçãoao estudo das camadas superiores da seriePassa Dous. Annaes da Academia Brasileira deSciencias 8 (1): 41-54.MÖRCH O. A. L. 1861. — Beiträge zur MolluskenfaunaCentral-Amerika’s. MalakozoologischeBlätter 7: 170-213.MÖRCH O. A. L. 1862 [June]. — Review <strong>of</strong> theVermetidae (Part III). Proceedings <strong>of</strong> the ZoologicalSociety <strong>of</strong> London 1862 (1): 54-83.MORNINGSTAR H. 1922. — Pottsville fauna <strong>of</strong> Ohio.Geological Survey <strong>of</strong> Ohio Bulletin (ser. 4) 25: 1-312,pls 1-16.MORONI M. A. & RUGGIERI G. 1980 [20 September].— Due Siphodentaliidae [sic] del Saheliano delPalermitano (Mollusca, <strong>Scaphopoda</strong>). Il NaturalistaSiciliano (ser. 4) 4 (1-2): 45-51.MORONI M. A. & RUGGIERI G. 1981 [March]. —Cadulus (Sulcogadila n. subgen.) caprottii n. sp.,nuovo scafopode del Siciliano (Pleistocene Inf.) diPalermo. Bolletino Malacologica 17 (1-2): 27-31.MORRIS J. 1845. — Fossil fauna: Mollusca, inSTRZELECKI P. E. DE (ed.), Physical Description <strong>of</strong>New South Wales <strong>and</strong> Van Diemen’s L<strong>and</strong>, Accompaniedby a Geological Map, Sections <strong>and</strong> Diagrams,<strong>and</strong> Figures <strong>of</strong> the Organic Remains. Longman,Brown, Green & Longmans, London: 270-291,pls 10-19.MORTON S. G. 1834. — Synopsis <strong>of</strong> the OrganicRemains <strong>of</strong> the Cretaceous Group <strong>of</strong> the United States.Key & Biddle, Philadelphia, 88 + 8 + [xii] p.,19 pls.MÜLLER J. 1851 [in 1847-1859]. — Monographie derPetrefacten der Aachener Kreideformation. ZweiteAbtheilung. Henry & Cohen, Bonn, 88 p., pls 3-6.MÜLLER O. F. 1776. — Zoologiae Danicae prodromus,seu animalium Daniae et Norvegiae indigenarumcharacters, nomina, et synonyma imprimis popularium.Hallageriis, Havniae [Copenhagen], xxxii +282 p.MURCHISON R. I. 1845. — Outline <strong>of</strong> the Geology <strong>of</strong>the neighbourhood <strong>of</strong> Cheltenham (new ed.). Murray,London, 109 p., 14 pls.NAGAO T. 1928. — Palaeogene <strong>fossil</strong>s <strong>of</strong> the Isl<strong>and</strong> <strong>of</strong>Kyûshû, Japan. Part II. The Science Reports <strong>of</strong> theTôhoku Imperial University, Sendai, Second Series(Geology) 12 (1): 1-140, pls 1-23.NAGAO T. 1938 [March]. — Some molluscan <strong>fossil</strong>sfrom the Cretaceous deposits <strong>of</strong> Hokkaidô <strong>and</strong>Japanese Saghalien. Journal <strong>of</strong> the Faculty <strong>of</strong> Science,Hokkaido Imperial University, Series IV, Geology <strong>and</strong>Mineralogy 4 (1-2): 117-142, pls 14-16 [1-3].NARDO G. D. 1847. — Sinonimia moderna delle specieregistrate nell’ opera intitolata: Descrizione de’ crostacei,de’ testacei e de’ pesci che abitano le Lagune etGolfo Veneto, rappresentati in figure a chiaroscuro eda colori, dall’Abate Stefano Chiereghini, Ven.Clodiense. Antonelli, Venezia, xi + 128 [= 64] p.NASSICHUK W. W. & HODGKINSON K. A. 1976[29 December]. — Scaphopods from the PermianAssistance Formation, Canadian Arctic Archipelago.Journal <strong>of</strong> Paleontology 50 (6): 1150-1156,pl. 1.NECHAEV A. V. 1897. — Fauna Eotzenovykhotlozhenii na Volghye mezhdu Saratovuim iTzaritzuinuim / Die Fauna der Eocänablagerungenan der Wolga unterhalb Saratow. Trudy, ObshchestvoEstestvoispuitatelei pri Imperatorskom KazanskomUniversitetye [Transactions <strong>of</strong> the Society <strong>of</strong>Naturalists <strong>of</strong> the Imperial Kazan University] 32 (1):1-247 + 10 pls (in Russian).NEGRI L. 1934 [December]. — Gasteropodi,Scafopodi e Cefalopodi paleogenici della Sirtica edel Fezzan orientale. Missione Scientifica dellaReale Accademia d’Italia a Cufra (1931-IX), Roma3: 131-181, pl. 14.NEWTON R. B. 1891. — Systematic List <strong>of</strong> the FrederickE. Edwards Collection <strong>of</strong> British Oligocene <strong>and</strong> EoceneMollusca in the British Museum (Natural History),with References to the Type-Specimens from SimilarHorizons Contained in other Collections Belonging tothe Geological Department <strong>of</strong> the Museum. BritishMuseum (Natural History), London, xxviii + 365 p.NEWTON R. B. 1902 [July]. — List <strong>of</strong> Thomas Say’stypes <strong>of</strong> Maryl<strong>and</strong> (U.S.) Tertiary Mollusca in theBritish Museum. Geological Magazine (n.s.) (decadeIV) 9 (7): 303-305.NEWTON R. B. & HARRIS G. F. 1894 [March]. — Arevision <strong>of</strong> the British Eocene <strong>Scaphopoda</strong>, withdescriptions <strong>of</strong> some new <strong>species</strong>. Proceedings <strong>of</strong> theMalacological Society <strong>of</strong> London 1 (2): 63-69, pl. 6.NICKLÈS M. 1955 [25 November]. — Scaphopodes etLamellibranches récoltés dans l’Ouest Africain.Atlantide Report. Scientific Results <strong>of</strong> the DanishExpedition to the Coasts <strong>of</strong> Tropical West Africa1945-1946 3: 93-237.NICKLÈS M. 1979 [31 March]. — Scaphopodes del’Ouest-Africain (Mollusca, <strong>Scaphopoda</strong>). Bulletindu Muséum national d’Histoire naturelle, Section A,ZOOSYSTEMA • 2004 • 26 (4)693


Steiner G. & Kabat A. R.Zoologie, Biologie et Écologie animales (sér. 4) 1 (1):41-77, 1 pl.NOETLING F. 1901. — Fauna <strong>of</strong> the Miocene Beds <strong>of</strong>Burma. Memoirs <strong>of</strong> the Geological Survey <strong>of</strong> India,Palaeontologia Indica (n.s.) 1 (3): 1-378, pls 1-25.NOMURA S. 1935. — <strong>Catalog</strong>ue <strong>of</strong> the Tertiary <strong>and</strong>Quaternary Mollusca from the isl<strong>and</strong> <strong>of</strong> Taiwan(Formosa) in the Institute <strong>of</strong> Geology <strong>and</strong>Palaeontology, Tôhoku Imperial University,Sendai, Japan. Part 2, <strong>Scaphopoda</strong> <strong>and</strong> Gastropoda.Science Reports <strong>of</strong> the Tôhoku Imperial University,Sendai, Japan, Second Series (Geology) 18 (2): 53-228 [1-176], pls 6-10 [1-5].NOMURA S. 1938 [December]. — Variation <strong>of</strong> ribs inDentalium octangulatum Donovan. Venus 8 (3-4):155-158.NOMURA S. 1940 [March]. — Mollusca dredged bythe Husa-Maru from the Pacific coast <strong>of</strong> TibaPrefecture, Japan. Records <strong>of</strong> the OceanographicWorks in Japan 12 (1): 81-116, 1 pl.NOMURA S. & HATAI K. 1940 [July]. — The marinefaune <strong>of</strong> Kyûroku-sima <strong>and</strong> its vicinity, northeastHonsyû, Japan. Saito Ho-on Kai Museum ResearchBulletin 19: 57-115, pls 3-4.NOMURA S. & NIINO H. 1940 [March]. — Molluscadredged from <strong>of</strong>f the coast <strong>of</strong> Hukui Prefecture inthe Japan Sea. Records <strong>of</strong> Oceanographic Works inJapan 12 (1): 23-79, 1 pl.NOSZKY J. 1936 [29 October]. — Az egri felsöCattien molluszkafaunája / Die Molluskenfaunades oberen Cattiens von Eger, in Ungarn. AnnalesHistorico-Naturales Musei Nationalis Hungarici,Budapest, Pars mineralogica, geologica, palaeontologica30: 53-115, pls 5-6 (in Hungarian withGerman summary).NOSZKY J. 1940. — A kiscelli agyag Molluszka-faunája.II. Rész Loricata, Gastropoda, <strong>Scaphopoda</strong> / DieMolluskenfauna des Kisceller Tones (Rupelien) ausder Umgebung von Budapestn II. Teil Loricata,Gastropoda und <strong>Scaphopoda</strong>. Annales Historico-Naturales Musei Nationalis Hungarici, ParsMineralogica, Geologica et Palaeontologica 33: 1-80,pls 1-2 (in Hungarian with German summary).NYST P. H. 1835. — Recherches sur les coquilles <strong>fossil</strong>esde la province d’Anvers. Perichon, Bruxelles, iii +36 p., 5 pls [not seen].NYST P. H. 1836. — Recherches sur les coquilles <strong>fossil</strong>esde Kleyn-Spauwen et Housselt (Province duLimbourg). Messager des Sciences et des Arts de laBelgique, ou Nouvelles Archives historiques, littéraireset scientifiques (G<strong>and</strong>) 4: 139-180, pls 1-4.NYST P. H. 1843. — [Sur les coquilles <strong>fossil</strong>es de laprovince d’Anvers]. Bulletin de la Société géologiquede France 14: 451-456.NYST P. H. 1845 [“1843”]. — Description descoquilles et des polypiers <strong>fossil</strong>es des terrains tertiairesde la Belgique. Mémoire couronné de l’Académieroyale des Sciences et Belles-Lettres de Bruxelles 17:1-697, pls 1-15.NYST P. H. 1883. — Conchyliologie des terrainstertiaires de la Belgique. Première Partie. TerrainPliocène Scaldisien. Annales du Musée royald’Histoire naturelle de Belgique, Série paléontologique3: lv + 263 p., 28 pls.NYST P. H. & WESTENDORP G. D. 1839. —Nouvelles recherches sur les coquilles <strong>fossil</strong>es de laprovince d’Anvers. Bulletins de l’Académie royale desSciences et Belles-Lettres de Bruxelles 6 (2) 393-414,3 pls.ODHNER N. HJ. 1931. — Die Scaphopoden. FurtherZoological Results <strong>of</strong> the Swedish Antarctic Expedition1901-1903 2 (5): 1-8, pls 1-2.O’GORMAN G. & COSSMANN M. 1923. — Le gisementCuisien de Gan (Basses-Pyrénées). Privately published,Pau, xxvii + 188 p., 14 pls [“Description desmollusques” by M. Cossmann: 1-188, pls 1-11].OKUTANI T. 1964 [July]. — Report on the archibenthal<strong>and</strong> abyssal scaphopod Mollusca mainly collectedfrom Sagami Bay <strong>and</strong> adjacent waters by theR.V. Soyo-Maru during the years 1955-1963, withsupplementary notes for the previous report onLamellibranchiata. Venus 23 (2): 72-90.OKUTANI T. 1966 [May]. — Archibenthal <strong>and</strong> abyssalMollusca collected by the R.V. Soyo-Maru fromJapanese waters during 1964. Bulletin <strong>of</strong> the TokaiRegional Fisheries Research Laboratory 46: 1-32,pls 1-2.OKUTANI T. 1975 [August]. — Deep-sea bivalves <strong>and</strong>scaphopods collected from deeper than 2,000 m inthe northwestern Pacific by the R/V Soyo-Maru <strong>and</strong>the R/V Kaiyo-Maru during the years 1969-1974.Bulletin <strong>of</strong> the Tokai Regional Fisheries ResearchLaboratory 82: 57-87, pls 1-3.OKUTANI T. 1982 [April]. — A remarkable tusk shell,Fissidentalium horikoshii n. sp., trawled from thedeep sea shelf <strong>of</strong>f Sanriku, Pacific coast <strong>of</strong> NortheastHonshu, Japan. Venus 41: 1-4.OLIVER P. G. 1984. — <strong>Scaphopoda</strong>. National Museum<strong>of</strong> Wales, H<strong>and</strong>lists <strong>of</strong> the Molluscan Collections in theDepartment <strong>of</strong> Zoology. Series 1: The Melvill-TomlinCollection 26: ii + 8 + vi + v + ii p.OLSSON A. A. 1922 [21 April]. — The Miocene <strong>of</strong>northern Costa Rica; with notes on its generalstratigraphic relations. Part I. Bulletins <strong>of</strong> AmericanPaleontology 9 (39):174-482 [1-310], pls 4-35 [1-32].OLSSON A. A. 1930 [25 December]. — Contributionsto the Tertiary paleontology <strong>of</strong> northern Peru.Part 3, Eocene Mollusca. Bulletins <strong>of</strong> AmericanPaleontology 17 (62): 1-96, pls 1-12.OLSSON A. A. 1942 [25 December]. — Tertiary <strong>and</strong>Quaternary <strong>fossil</strong>s from the Burica Peninsula <strong>of</strong>Panama <strong>and</strong> Costa Rica. Bulletins <strong>of</strong> AmericanPaleontology 27 (106): 153-258 [1-106], pls 14-25[1-12].OLSSON A. A. 1964 [28 October]. — NeogeneMollusks from Northwestern Ecuador. PaleontologicalResearch Institution, Ithaca (NY), 256 p.,38 pls.694ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)OMALIUS-D’HALLOY J. J. D’ 1853. — Géologie de laBelgique. Encyclopédie populaire. A. Jamar,Bruxelles: 279-400. Remark: see Dupont (1876:272, 289) for date <strong>of</strong> publication.OPPEL A. 1856-58. — Die Juraformation Engl<strong>and</strong>s,Frankreich und des sudwestlichen Deutschl<strong>and</strong>s. Nachihren einzelnen Gliedern eingetheilt und verglichen.Ebner & Seubert, Stuttgart, 857 p. Dates <strong>of</strong> publication:1-438 (1856); 439-694 + map (1857);695-857 + table (1858).OPPENHEIM P. 1906. — Zur Kenntnis alttertiärerFaunen in Ägypten. 2. Lieferung: Der Bivalvenzweiter Teil, Gastropoda und Cephalopoda.Palaeontographica 30 (III) (7): 165-348, pls 18-27.OPPENHEIM P. 1919 [“1918”]. — Das Neogen inKleinasien. I. Teil. Zeitschrift der deutschen GeologischenGesellschaft 70 (1-4): 1-210, pls 1-12.ORBIGNY A. C. D. D’ 1843 [in 1842-1843]. —Paléontologie française. Description zoologique etgéologique de tous les animaux mollusques et rayonnés<strong>fossil</strong>es de France. Terraines cretacées, Tome II[Gastropoda]. V. Masson, Paris, 456 p., pls 149-236. Remark: pages 1-80 were published in1842; pages 81-456 in 1843 (fide Sherborn 1899:223).ORBIGNY A. C. D. D’ 1845. — Terrain Secondaire.Système Jurassique (étage Oxfordien). Mollusques,in MURCHISON R. I., VERNEUIL É. DE & KEYSERLINGA. DE (eds), Géologie de la Russie d’Europe et desmontagnes de l’Oural, Volume II, Troisième Partie,Paléontologie. John Murray, London; Bertr<strong>and</strong>,Paris: 419-488, pls 28-42.ORBIGNY A. C. D. D’ 1850-1852. — Prodrome depaléontologie stratigraphique universelle des animauxmollusques et rayonnés faisant suite au cours élémentairede paléontologie et de géologie stratigraphiques.Victor Masson, Paris 1: lx + 394 p. (1850); 2:428 p. (1850); 3: 196 p. + 190 p. [Index] (1852).Remark: The new <strong>species</strong> in this work are erroneouslydated as “d’Orb., 1847” in the text.ORBIGNY A. C. D. D’ 1853 [in 1842-1853]. — Mollusques,in SAGRA R. DE LA (ed.), Histoire physique,politique et naturelle de l’Île de Cuba. Vol. 2.Bertr<strong>and</strong>, Paris, 380 p., 28 pls. Remark: refer toAguayo (1943) for dates <strong>of</strong> publication.ORTMANN A. E. 1902 [19 April]. — TertiaryInvertebrates. Reports <strong>of</strong> the Princeton UniversityExpeditions to Patagonia 1896-1899, IV, PaleontologyI (Part II): 45-332, pls 11-39.OTUKA Y. 1933 [December]. — Description <strong>of</strong> a newDentalium from Southern Japan. Venus 4 (3): 159-161. Remark: the name <strong>of</strong> this author was transliteratedin the original publication as “Otuka” butsome modern authorities have transliterated thisname as “Otsuka”.OTUKA Y. 1935 [December]. — The Oti Graben inSouthern Noto Peninsula (Part 3). Bulletin <strong>of</strong> theEarthquake Research Institute 13 (4): 846-909,pls 53-57.OYAMA K. 1973 [30 November]. — Revision <strong>of</strong>Matajiro Yokoyama’s type Mollusca from theTertiary <strong>and</strong> Quaternary <strong>of</strong> the Kanto Area.Palaeontological Society <strong>of</strong> Japan, Special Papers 17:148 p., 57 pls.OZAKI H. 1956 [March]. — Some new <strong>and</strong> interestingmolluscs from Miocene Nobori Formation inthe eastern Kôti Prefecture, Japan. Bulletin <strong>of</strong> theNational Science Museum, Tokyo 3 (1): 1-6, 1 pl.PAETEL F. 1873. — <strong>Catalog</strong> der Conchylien-Sammlungvon F. Paetel, nebst uebersicht des angew<strong>and</strong>tenSystems. Berlin, iv + 172 p.PAETEL F. 1887-1888. — <strong>Catalog</strong> der Conchylien-Sammlung, mit Hinzufügung der bis jetzt publicirtenrecenten Arten, sowie der ermittelten Synonyma.Fourth Edition. Vol. 1. Berlin, 639 p. Remark:pages 1-480 (parts 1-6) were published in 1887;pages 481-639 (parts 7-8) were published in 1888.PALMER C. P. 1974a [1 October]. — A supraspecificclassification <strong>of</strong> the scaphopod Mollusca. TheVeliger 17: 115-123.PALMER C. P. 1974b [1 October]. — Rectification <strong>of</strong>nomenclature in the molluscan class <strong>Scaphopoda</strong>.The Veliger 17: 124-125.PALMER C. P. 1975 [17 May]. — A new Jurassicscaphopod from the Oxford Clay <strong>of</strong> Buckinghamshire.Palaeontology 18 (2): 377-383, pl. 52.PALMER C. P. 1986 [December]. — <strong>Catalog</strong>ue <strong>of</strong>genera in the class <strong>Scaphopoda</strong>. The Conchologists’Newsletter 99: 395-397.PALMER C. P. 1987a [September]. — Confessions <strong>of</strong> ataxonomic sinner-mea culpa. The ConchologistsNewletter 102: 30-31.PALMER C. P. 1987b [December]. — The case forAntalis tarentinum Lamarck, 1818. The Conchologists’Newsletter 103: 54-56.PALMER C. P. 2001 [May]. — Dentalium giganteumPhillips: a serpulid worm tube. Proceedings <strong>of</strong> theYorkshire Geological Society 53 (3): 253-255.PALMER C. P. & STEINER G. 1998. — <strong>Scaphopoda</strong>,Introduction, in BEESLEY P. L., ROSS G. J. P. &WELLS A. (eds), Mollusca: The Southern Synthesis.Fauna <strong>of</strong> Australia. Vol. 5A. CSIRO Publishing,Melbourne: 431-438.PALMER D. B. K. 1923 [3 November]. — A faunafrom the Middle Eocene shales near Vacaville,California. University <strong>of</strong> California Publications,Bulletin <strong>of</strong> the Department <strong>of</strong> Geological Sciences14 (8): 289-318, pls 52-57.PALMER K. VAN W. 1937 [20-25 December]. — TheClaibornian <strong>Scaphopoda</strong>, Gastropoda <strong>and</strong>dibranchiate Cephalopoda <strong>of</strong> the southern UnitedStates. Bulletins <strong>of</strong> American Paleontology 7 (32):1-730, pls 1-90.PALMER K. VAN W. 1947 [6 February]. — The Mollusca<strong>of</strong> the Jackson Eocene <strong>of</strong> the Mississippi Embayment(Sabine River to the Alabama River). Part II,Univalves <strong>and</strong> Index. Bulletins <strong>of</strong> AmericanPaleontology 30 (117) (Section 2): 207-564, pls 26-65.ZOOSYSTEMA • 2004 • 26 (4)695


Steiner G. & Kabat A. R.PALMER K. VAN W. 1958 [8 December]. — Typespecimens <strong>of</strong> marine Mollusca described byP. P. Carpenter from the West Coast (San Diego toBritish Columbia). Memoirs <strong>of</strong> the Geological Society<strong>of</strong> America 76: viii + 376 p., pls 1-35.PALMER K. VAN W. & BRANN D. C. 1965 [25 June].— <strong>Catalog</strong>ue <strong>of</strong> the Paleocene <strong>and</strong> Eocene Mollusca<strong>of</strong> the southern <strong>and</strong> eastern United States. Part I.Pelecypoda, Amphineura, Pteropoda, <strong>Scaphopoda</strong>,<strong>and</strong> Cephalopoda. Bulletins <strong>of</strong> American Paleontology48 (218): 1-466, 3 pls.PATRULIUS D. 1996. — Heteromorphic ammonites<strong>and</strong> other parkinsoniids <strong>of</strong> the Bathonian-LowerCallovian <strong>of</strong> Vadu Crisului (Apuseni Mountains-Romania). Memoriile Institutului Geologic alRomaniei 36: 13-19.PAVIA G. 1991 [27 February]. — I molluschi delMessiniano di Borelli (Torino). 2. <strong>Scaphopoda</strong>.Bolletino del Museo Regionale di Scienze Naturali(Torino) 9 (1): 105-172, 9 pls.PENNA-NEME L. 1974 [28 June]. — Consideraçõessobre os <strong>Scaphopoda</strong> (Mollusca) da costa brasilieira,com descrição de uma nova espécie. Papéis Avulsosde Zoologia, São Paulo 28 (6): 105-126.PENNA-NEME L. & DA COSTA BARROS MUNIZ G.1976 [30 September]. — Um novo Dentalium(Mollusca – <strong>Scaphopoda</strong>) da Formacâo MariaFarinha, Paleoceno de Pernambuco. Anais daAcademia Brasiliera de Ciências 48(3): 523-525.PERRILLIAT M. DEL C. 1974. — Monografia de losMoluscos del Mioceno medio de Santa Rosa,Veracruz, México. Parte IV (Escafópodos:Dentaliidae, Siphonodentaliidae). PalaeontologiaMexicana 37: 51-64, pls 9-14.PERRY G. 1811. — Conchology, or the Natural History<strong>of</strong> Shells: Containing a New Arrangement <strong>of</strong> theGenera <strong>and</strong> Species, Illustrated by ColouredEngravings, Executed from the Natural Specimens <strong>and</strong>Including the Latest Discoveries. Miller, London,4p., 62 pls.PETIT R. E. & BOSS K. J. 1989 [30 August]. — SearlesV. Wood’s “Monograph <strong>of</strong> the Crag Mollusca”(1848-1882), with notes on the generic nameAstartella. Occasional Papers on Mollusks 5 (68):145-153.PETIVER J. 1702-1711. — Gazophylacium Naturae etArtis. London.PETIVER J. 1713. — Aquatilium Animalium Amboinae,&c. Icones & nomina. Containing near 400 figures...<strong>of</strong> aquatick crustaceous <strong>and</strong> testaceous Animals; ...found about Amboina, <strong>and</strong> the neighboring Indianshores, with their Latin, English, Dutch, <strong>and</strong> native<strong>names</strong>. J. Bateman, London, 4 p., 20 pl.PEYROT A. 1938. — Les mollusques testacés univalvesdes dépots helvétiens du Bassin Ligérien. <strong>Catalog</strong>uecritique, descriptif et illustré. Actes de la Société linnéennede Bordeaux 89 (Supplément): 361 p., 5 pls.PHILIPPI R. A. 1836. — Enumeratio MolluscorumSiciliae, cum viventium tum in tellure tertiaria <strong>fossil</strong>ium,quae in itinere suo observavit. Vol. I. SimonisSchroppii, Berlin, xiv + 268 p., pls 1-12.PHILIPPI R. A. 1844. — Enumeratio MolluscorumSiciliae, cum viventium tum in tellure tertiaria <strong>fossil</strong>ium,quae in itinere suo observavit. Continens addendaet emendenda, nec non comparationem faunae recentisSiciliae cum faunis aliarum terrarum et cum faunaperiodi tertiariae. Vol. II. E. Anton, Halle, iv +303 p., pls 13-28.PHILIPPI R. A. 1846 [April]. — Diagnosen neuerConchylien-Arten. Zeitschrift für Malakozoologie3: 49-55.PHILIPPI R. 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Proceedings <strong>of</strong>the Malacological Society <strong>of</strong> London 22 (3): 153-156.WINCKWORTH R. 1940 [15 July]. — New <strong>species</strong> <strong>of</strong>shells from Madras. Proceedings <strong>of</strong> the MalacologicalSociety <strong>of</strong> London 24 (2): 41-43.WISSMANN H. L. 1841. — Beiträge zur Geognosieund Petrefakten-Kunde des südöstlichen Tirol’svorzüglich der Schichten von St Cassian. Beiträgezur Petrefacten-Kunde 4: 1-152, 16 pls.WOOD S. V. 1842 [August]. — A catalogue <strong>of</strong> shellsfrom the Crag. Annals <strong>and</strong> Magazine <strong>of</strong> NaturalHistory (series 1) 9 (60): 455-462, pl. 5.WOOD S. V. 1848. — A monograph <strong>of</strong> the CragMollusca. Palaeontographical Society <strong>of</strong> London,v-xii + 209 p., pls 1-21 [refer to Petit & Boss (1989)for dates <strong>and</strong> collation].WOODRING W. P. 1925 [20 May]. — Contributionsto the geology <strong>and</strong> palaeontology <strong>of</strong> the West-Indies: Miocene mollusks from Bowden, Jamaica.Part I. Pelecypods <strong>and</strong> Scaphopods. CarnegieInstitution <strong>of</strong> Washington Publication 366: viii +222 p., 28 pls.WOODRING W. P. 1973. — Geology <strong>and</strong> paleontology<strong>of</strong> Canal Zone <strong>and</strong> adjoining parts <strong>of</strong> Panama.Description <strong>of</strong> Tertiary Mollusca (additions toGastropods, Scaphopods, Pelycypods: Nuculidae toMalleidae). United States Geological SurveyPr<strong>of</strong>essional Paper 306-E: 453-539, pls 67-82.706ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)WOODS H. 1896 [1 February]. — The Mollusca <strong>of</strong>the Chalk Rock. Quarterly Journal <strong>of</strong> the GeologicalSociety <strong>of</strong> London 52 (1): 68-98, pls 2-4.WOODWARD H. B. 1893. — Memoirs <strong>of</strong> the GeologicalSurvey <strong>of</strong> the United Kingdom. The Jurassic Rocks <strong>of</strong>Britain. Volume III. The Lias <strong>of</strong> Engl<strong>and</strong> <strong>and</strong> Wales(Yorkshire excepted). HMSO, London, xii + 399 p.WORTHEN A. H. 1883. — Descriptions <strong>of</strong> some new<strong>species</strong> <strong>of</strong> <strong>fossil</strong> shells from the UpperCarboniferous Limestones <strong>and</strong> Coal Measures <strong>of</strong>Illinois. Geological Survey <strong>of</strong> Illinois (Geology <strong>and</strong>Palaeontology) 8: 323-338.WRIGHT C. W. & CLEEVELY R. J. 1985 [2 April]. —Authorship <strong>and</strong> dates <strong>of</strong> the Sowerby’s MineralConchology <strong>of</strong> Great Britain. Bulletin <strong>of</strong> ZoologicalNomenclature 42 (1): 64-71.WU S.-K. & BRANDAUER N. E. 1982 [1 October]. —Type specimens <strong>of</strong> <strong>Recent</strong> Mollusca in theUniversity <strong>of</strong> Colorado Museum. Natural HistoryInventory <strong>of</strong> Colorado 7: 1-47.YANCEY T. E. 1973 [November]. — A new genus <strong>of</strong>Permian siphonodentalid scaphopods, <strong>and</strong> its bearingon the origin <strong>of</strong> Siphonodentaliidae. Journal <strong>of</strong>Paleontology 47 (6): 1062-1064.YANCEY T. E. 1978 [26 October]. — Brachiopods <strong>and</strong>molluscs <strong>of</strong> the Lower Permian Arcturus Group,Nevada <strong>and</strong> Utah. Part 1: brachiopods, scaphopods,rostroconchs, <strong>and</strong> bivalves. Bulletins <strong>of</strong> AmericanPaleontology 74 (303): 257-367, pls 1-10.YAO P. 1990 [June]. — Discovery <strong>of</strong> Dentalium(Dentaliidae, <strong>Scaphopoda</strong>) from Lower Cretaceous<strong>of</strong> Gucuo village, Nyalam, southern Xizang (Tibet).Geoscience, Journal <strong>of</strong> Graduate School, ChinaUniversity <strong>of</strong> Geosciences 4 (2): 30-32, pl. 1.YOCHELSON E. L. 1968. — On the nature <strong>of</strong> Polylopia.United States Geological Survey Pr<strong>of</strong>essional Paper593-F: 1-7, pl. 1.YOCHELSON E. L. 1971 [3 February]. — The littleknownPennsylvanian Clavulites reinterpreted as a“worm”. Journal <strong>of</strong> Paleontology 45 (1): 126-129.YOCHELSON E. L. 1987 [14 January]. — Redescription<strong>of</strong> Spirodentalium Walcott (Gastropoda: LateCambrian) from Wisconsin. Journal <strong>of</strong> Paleontology61 (1): 66-69.YOCHELSON E. L. 1999 [25 January]. — Rejection <strong>of</strong>Carboniferous Quasidentalium Shimansky, 1974,from the phylum Mollusca. Journal <strong>of</strong> Paleontology73 (1): 63-65.YOCHELSON E. L. 2002. — Restudy <strong>and</strong> reassignment <strong>of</strong>Dentalium antiquum Goldfuss, 1841 (Middle Devonian).Paläontologische Zeitschrift 76 (2): 297-304.YOCHELSON E. L. & GOODISON R. 1999 [12 July]. —Devonian Dentalium martini Whitfield, 1882, isnot a mollusk but a worm. Journal <strong>of</strong> Paleontology73 (4): 634-640.YOCHELSON E. L. & HOLLAND, C. H. 2004 [April].— Dentalium saturni Goldfuss, 1841 (Eifelian:Mollusca): complex issues from a simple <strong>fossil</strong>.Paläontologische Zeitschrift 78: 97-102.YOCHELSON E. L. & SAUNDERS B. W. 1967. — A bibliographicindex <strong>of</strong> North American Late PaleozoicHyolitha, Amphineura, <strong>Scaphopoda</strong>, <strong>and</strong>Gastropoda. United States Geological Survey Bulletin1210: 271 p.YOKOYAMA M. 1920 [22 March]. — Fossils from theMiura Peninsula <strong>and</strong> its immediate north. Journal<strong>of</strong> the College <strong>of</strong> Science, Imperial University <strong>of</strong> Tokyo39 (6): 1-198, pls 1-20.YOKOYAMA M. 1922 [7 July]. — Fossils from theUpper Musashino <strong>of</strong> Kazusa <strong>and</strong> Shimosa. Journal<strong>of</strong> the College <strong>of</strong> Science, Imperial University <strong>of</strong> Tokyo44 (1): viii + 1-200, pls 1-17.YOKOYAMA M. 1923. — Tertiary <strong>fossil</strong>s from Kii.Japanese Journal <strong>of</strong> Geology <strong>and</strong> Geography 2 (3):47-58, pls 6-7.YOKOYAMA M. 1926 [18 September]. — Fossil shellsfrom Sado. Journal <strong>of</strong> the Faculty <strong>of</strong> Science,Imperial University <strong>of</strong> Tokyo. Section II: Geology,Mineralogy, Geography, Seismology 1 (8): 249-312,pls 32-37.YOKOYAMA M. 1927a. — Mollusca from the UpperMusashino <strong>of</strong> Toyko <strong>and</strong> its suburbs. Journal <strong>of</strong> theFaculty <strong>of</strong> Science, Imperial University <strong>of</strong> Tokyo.Section II: Geology, Mineralogy, Geography,Seismology 1 (10): 391-437, pls 46-50.YOKOYAMA M. 1927b [28 July]. — Tertiary shellsfrom the coal-field <strong>of</strong> Haboro, Teshio. Journal <strong>of</strong>the Faculty <strong>of</strong> Science, Imperial University Tokyo,Section II: Geology, Mineralogy, Geography, Seismology2: 191-204, pls 51-52YOKOYAMA M. 1929. — Pliocene shells fromTonohama, Tosa. Report, Imperial Geological Survey<strong>of</strong> Japan 104: 9-17, pls 7-8.YOKOYAMA M. 1931 [30 June]. — Neogene shellsfrom Karafto <strong>and</strong> the Hokkaido. Journal <strong>of</strong> theFaculty <strong>of</strong> Science, Imperial University Tokyo, SectionII: Geology, Mineralogy, Geography, Seismology 3 (4):197-204, pl. 11.YOO E. K. 1988 [16 December]. — Early CarboniferousMollusca from Gundy, Upper Hunter,New South Wales. Records <strong>of</strong> the Australian Museum40 (3-4): 233-264.YOUNG J. A. JR 1942 [9 January]. — Pennsylvanian<strong>Scaphopoda</strong> <strong>and</strong> Cephalopoda from New Mexico.Journal <strong>of</strong> Paleontology 16 (1): 120-125, pl. 20.ZARDINI R. 1978 [November]. — Fossili Cassiani(Trias Medio-Superiore). Atlante dei gasteropodidella formazione di S. Cassiano raccolta nellaregione dolomitica attorno a Cortina d’Ampezzo.Edizioni Ghedina, Cortina d’Ampezzo, 58 p.,41 pls.ZEIDLER W. & MACPHAIL M. K. 1978 [12 June]. —Mollusc type-specimens in the South AustralianMuseum; I: Cephalopoda <strong>and</strong> <strong>Scaphopoda</strong>.Records <strong>of</strong> the South Australian Museum 17 (26):381-385.ZEKELI L. F. 1852. — Die Gastropoden der Gosaugebilde.Abh<strong>and</strong>lungen der kaiserlich-königlichenZOOSYSTEMA • 2004 • 26 (4)707


Steiner G. & Kabat A. R.geologischen Reichstanstalt in Wien, 2. Abtheilung1(2): 1-124, pls 1-24.ZENKER J. C. 1836. — Historisch-topographischesTaschenbuch von Jena und seiner Umgebung besondersin naturwissenschaftlicher u. medicinischerBeziehung. F. Frommann, Jena, x + 338 p.ZILCH A. 1934 [21 December]. — Zur Fauna desMittel-Miocäns von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger’sBearbeitungen. Senckenbergiana 16 (4-6): 193-302,22 pls.ZIMMERMANN K. G. 1847. — Ueber Tertiärpetrefacteaus dem Sachsenwalde. Amtlicher Bericht über die24. Versammlung Deutscher Naturforscher und Aerztein Kiel im September 1846: 240-244.ZINNDORF J. 1928. — Die Versteinerungen aus denTertiär-Ablagerungen von Offenbach a. M. I. Teil.Die Conchylien des Rupeltones (Septarientones).Bericht über die Tätigkeit des Offenbacher Vereins fürNaturkunde 66-68: 1-65, pls 1-6.ZITTEL K. A. 1864. — Fossile Mollusken undEchinodermen aus Neu-Seel<strong>and</strong>. Novara-Expedition,Geologischer Theil, I., B<strong>and</strong> 2. Abtheilung,Paläontologie von Neu-Seel<strong>and</strong>. Beiträge zurKenntniss der Fossilen Flora und Fauna der ProvinzenAuckl<strong>and</strong> und Nelson: 15-68, pls 6-15.Submitted on 19 October 2001;accepted on 15 October 2004.708ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)APPENDIX 1Rectification <strong>of</strong> nomenclature. Renaming <strong>of</strong> homonyms.In the course <strong>of</strong> preparing this catalog, we identifieda significant number <strong>of</strong> junior homonyms.Since the majority <strong>of</strong> scaphopod <strong>species</strong> wereoriginally described in the genus Dentalium, it isnot surprising that the same <strong>species</strong> <strong>names</strong>hould be used twice, or even more <strong>of</strong>ten, forthis genus. Pilsbry & Sharp (1897-1898)renamed numerous homonyms, except for severalthat they considered to be nomina dubia orotherwise invalid (e.g., “The name is preoccupied;but as this is probably not a valid <strong>species</strong> itneed not be renamed”, Pilsbry & Sharp 1898:310).Subsequently, Emerson (1954) renamed 12 juniorhomonyms, <strong>and</strong> Palmer (1974b) renamed11 junior homonyms. These authors tartly notedthat nearly all <strong>of</strong> these homonyms “could havebeen avoided” if the authors had taken the timeto consult Pilsbry & Sharp (1897-1898) beforedescribing their new <strong>species</strong>.We have uncovered an additional 29 juniorhomonyms, 13 <strong>of</strong> which are listed below <strong>and</strong> areherein renamed. Full bibliographic data for each<strong>species</strong> is given in the main catalog. There are 11taxa that we have not renamed, although ICZN(1999) article 60.1 requires replacement <strong>names</strong>for junior homonyms that do not have an “available<strong>and</strong> potentially valid synonym”. If either thesenior or the junior homonym is a dubious<strong>species</strong> due to the lack <strong>of</strong> type material, localitydata, or proper descriptions, or if a junior synonymis available, we avoided introducing new<strong>names</strong> for probably invalid taxa. For each <strong>of</strong> thesejunior homonyms we justified our decisions inthe main catalog: Dentalium attenuatumSowerby, 1860 non Say, 1824; Dentalium ovaleCooke, 1928 non “Malm.”; Dentalium dentalisvar. aequicostata Foresti, 1895 non Dentaliumaequicostatum Koenen, 1892; Dentalium angulatumVinassa de Regny, 1897 non Buckman inMurchison, 1845; Dentalium annulatumCrist<strong>of</strong>ori & Jan, 1832 non Gmelin, 1791; Dentaliumbrevifissum Galeotti, 1837 non Deshayes,1825; Dentalium calabrum Crema, 1910 nonO. G. Costa, 1851; Dentalium conicum Seguenza,1876 non Hutton, 1873; Dentalium decemcostatumQuaas, 1902 non Brazier, 1877; Dentalium(Fustiaria) geinitzi Koenen, 1892 non Böhm,1885; Dentalium(?) giganteum Zenker, 1836 nonPhillips, 1829; Dentalium inaequicostatumDautzenberg, 1891 non Seguenza, 1879;Dentalium (Fustiaria) inversum Cossmann, 1915non Deshayes, 1825; Dentalium octogonumRayneval, Hecke & Ponzi, 1854 non Lamarck,1818; Dentalium (Entalina) mediocarinatapliocenica Koperberg, 1931 non Sacco, 1897;Dentalium rugosum Eichwald, 1857 non Dunker,1849.Furthermore, this list <strong>of</strong> renamed homonymsdoes not include the numerous subspecific-rank<strong>names</strong> <strong>of</strong> Sacco (1897) <strong>and</strong> Locard, who repeatedlyused the same name below the <strong>species</strong> level.Should a subsequent researcher recognize one <strong>of</strong>Sacco’s or Locard’s junior homonyms as beingvalid, it can then be renamed.Further, to the best <strong>of</strong> our knowledge, only one<strong>of</strong> these junior homonyms, Antalis inaequicostata(Dautzenberg, 1891) can be preserved pursuantto ICZN (1999) article 23.9.1, which allows forhomonyms that have been widely used in the literatureto be preserved if the senior name has notbeen used since 1899.Where possible, we contacted the authors <strong>of</strong> the<strong>species</strong> taxa that are junior homonyms, i.e.Rodolfo Brunet, Maria del Carmen Perrilliat <strong>and</strong>Victor Scarabino, to ascertain whether theywould prefer to rename their own homonyms.Drs Perrilliat <strong>and</strong> Scarabino kindly consented toour renaming their homonyms; no reply wasreceived from Mr Brunet, despite severalinquiries.Dentalium (Graptacme) acuticostatum Plate, 1908anon Dentalium acuticostata Deshayes, 1825. Hereinrenamed Graptacme acutistriata Steiner & Kabat,2004.ZOOSYSTEMA • 2004 • 26 (4)709


Steiner G. & Kabat A. R.Dentalium annulatum Meyer, 1886 non Dentaliumannulatum Gmelin, 1791. Herein renamedDentalium parannulatum Steiner & Kabat, 2004.Dentalium asperum Brunet, 1995 non Dentalium asperumMichelotti, 1847. Herein renamed Dentaliumbruneti Steiner & Kabat, 2004.Dentalium bicarinatum Wanner, 1902 non Dentaliumbicarinatum Deshayes, 1825. Herein renamedDentalium wanneri Steiner & Kabat, 2004.Dentalium brevifissum Brugnone, 1876 non Dentaliumbrevifissum Deshayes, 1825. Herein renamedDentalium brugnonei Steiner & Kabat, 2004.Dentalium duodecimcostatum Brazier, 1877 nonDentalium duodecimcostatum d’Archiac, 1850.Herein renamed Dentalium healyi Steiner & Kabat,2004.Dentalium (Laevidentalium) ellipticum Koenen, 1892non Dentalium ellipticum J. Sowerby, 1814. Hereinrenamed Laevidentalium pseudellipticum Steiner &Kabat, 2004.Dentalium flavum Scarabino, 1995 non Dentaliumflavum Henderson, 1920. Herein renamed Dentaliumscarabinoi Steiner & Kabat, 2004.Dentalium hexagonum Van Ingen, 1905 nonDentalium hexagonum Gould, 1859. Hereinrenamed Dentalium vaningeni Steiner & Kabat,2004.Cadulus martini Scarabino, 1995 non Cadulus martiniFinlay, 1927. Herein renamed Cadulus scarabinoiSteiner & Kabat, 2004.Dentalium (Tesseracme) mexicanum Perrilliat, 1975non Dentalium mexicanum Girty, 1909. Hereinrenamed Dentalium perrilliatae Steiner & Kabat,2004.Dentalium minor Stephenson, 1952 non Dentaliumcaudani var. minor Locard, 1898. Herein renamedDentalium minorinum Steiner & Kabat, 2004.Dentalium polygonum Casey, 1903 non Dentaliumpolygonum Reuss, 1844. Herein renamed Dentaliumcaseyi Steiner & Kabat, 2004.APPENDIX 2Arguments for the taxonomic decisions.NEW GENERIC COMBINATIONSDentalium americanum Chenu, 1843 has previouslybeen placed in Dentalium (e.g., Pilsbry &Sharp 1897: 22; Henderson 1920: 27; Scarabino1975: 183). This <strong>species</strong> shows six primary ribscharacteristic for the genus Paradentalium,whereas Dentalium has eight to 12 primary ribs.Therefore, we have transferred this <strong>species</strong> toParadentalium.Dentalium peruvianum Dall, 1908 has the followingcharacters diagnostic <strong>of</strong> Fissidentalium:large (90 mm), almost straight shell with slightlydepressed anterior aperture; a high number <strong>of</strong>primary ribs (about 26); a long <strong>and</strong> wide slit onthe ventral side <strong>of</strong> the apex. In agreement with V.Scarabino (in litt. 6.III.2000), we have transferredthis <strong>species</strong> to Fissidentalium.Dentalium cancellatum Sowerby, 1860 shows sixto eight primary ribs (intercalating towards theanterior aperture) intersected by raised transversestriae giving the shell a cancellated appearancetypical for Coccodentalium. Similar patterns <strong>of</strong>sculpture are also present in some <strong>species</strong> <strong>of</strong>Fissidentalium <strong>and</strong> Compressidentalium, both <strong>of</strong>which have more than 12 primary ribs. We,therefore, transferred this <strong>species</strong> to Coccodentalium.Cadulus (Platyschides) foweyensis Henderson,1920 has four apical lobes, two lateral <strong>and</strong> oneeach dorsal <strong>and</strong> ventral, separated by distinctnotches. This is typical for the genus Polyschides.We, therefore, transferred this <strong>species</strong> toPolyschides.Cadulus (Platyschides) nitidus Henderson, 1920has four apical lobes, two lateral <strong>and</strong> one eachdorsal <strong>and</strong> ventral, separated by distinct notches.This is typical for the genus Polyschides. We,therefore, have transferred this <strong>species</strong> toPolyschides.Cadulus poculum Dall, 1889 has been transferredto the genus Polyschides because <strong>of</strong> the position <strong>of</strong>the maximum diameter being in the anteriorthird <strong>of</strong> the shell <strong>and</strong> the lateral projections at theapex indicating four apical lobes. Henderson710ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)(1920: 108, 109) listed this <strong>species</strong> in his subgenusPlatyschides that was synonymized withGadila by Scarabino (1995). Although the placementinto Gadila is possible due to the inadequatelypreserved apical features, the stronglyoblique anterior aperture resembles that <strong>of</strong> severalPolyschides <strong>species</strong>.Cadulus (Platyschides) portoricensis Henderson,1920 has four apical lobes typical for the genusPolyschides. We, therefore, have transferred this<strong>species</strong> to Polyschides.Siphodentalium teres Jeffreys, 1883 has beenremoved from Siphonodentalium because it hasneither more than four apical lobes nor an anteriorconstriction, which is diagnostic <strong>of</strong>Siphonodentalium. Because <strong>of</strong> the long <strong>and</strong> slightlytapering shell <strong>and</strong> the radula morphology(polygonal, unicuspidate rhachis; lateral toothwith three cusps, the posterior two being closer toeach other <strong>and</strong> separated from the anterior cuspby a nodulous ridge), we have transferred this<strong>species</strong> to Pulsellum. The two lateral lobes atthe shell apex, however, are unique among the<strong>Scaphopoda</strong>.NEW SYNONYMIESThe following taxa have previously been recognizedas a valid <strong>species</strong>, or as infrasubspecific taxa,<strong>of</strong> Fissidentalium: Dentalium ergasticum Fischer,1883 (Pilsbry & Sharp 1897: 74, 75) <strong>and</strong> itsforms D. e. var. albida Locard, 1898, D. e. var.elongata Locard, 1898, D. e. var. major Locard,1896, D. e. var. minor Locard, 1898; D. milneedwardsiLocard, 1897; D. scamnatum Locard,1897 (Pilsbry & Sharp 1897: 79) <strong>and</strong> its formsD. s. var. alba Locard, 1898, D. s. var. minorLocard, 1898, D. s. var. subrecta Locard, 1898.However, these taxa fall within the range <strong>of</strong> variation<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synoymy <strong>of</strong> D. ergasticum Fischer, 1883,D. milneedwardsi Locard, 1897, <strong>and</strong> D. scamnatumLocard, 1897 with Fissidentalium capillosum(Jeffreys, 1877) is, therefore, credited toB. Métivier. Note that this synonymy changesthe name <strong>of</strong> the type <strong>species</strong> <strong>of</strong> the genusFissidentalium from Fissidentalium ergasticum toFissidentalium capillosum.Dentalium caudani Locard, 1896 <strong>and</strong> its formsD. c. var. parfaiti Locard, 1898, D. c. var. intermediaLocard, 1898, <strong>and</strong> D. c. var. minor Locard,1898 have previously been recognized as a valid<strong>species</strong>, or as infrasubspecific taxa, <strong>of</strong> Laevidentalium(Pilsbry & Sharp 1897: 104).However, these taxa fall within the range <strong>of</strong> variation<strong>of</strong> Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synonymy <strong>of</strong> D. caudani Fischer, 1883 withF. c<strong>and</strong>idum (Jeffreys, 1877) is, therefore, creditedto B. Métivier.Dentalium exuberans Locard, 1897 <strong>and</strong> its formD. e. var. minor Locard, 1898 have previouslybeen recognized as a valid <strong>species</strong>, or as infrasubspecifictaxon, <strong>of</strong> Fissidentalium (Pilsbry & Sharp1897: 78, 79). However, shell shape <strong>and</strong> sculpture<strong>of</strong> these taxa are identical with the previouslydescribed F. paucicostatum (Watson, 1879), asBernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synonymy <strong>of</strong> D. exuberans Locard, 1897with F. paucicostatum (Watson, 1879) is, therefore,credited to B. Métivier.Dentalium (Fustiaria) inversum Cossmann, 1915is placed in tentative synonymy with Fustiariamontense (Briart & Cornet, 1889) because it differsonly in the dorsal position <strong>of</strong> the apical slit.Cossmann (1915: 7) remarked that the type specimenmay be only an inverted specimen <strong>of</strong>Fustiaria montense. Dorsal displacement <strong>of</strong> the slitinfrequently occurs in other <strong>species</strong> <strong>of</strong> the genus(G. S. pers. obs.) <strong>and</strong> is, thus, not considered sufficientto separate a <strong>species</strong>, especially since thedescription was based on a single specimen only.Dentalium tessellatum Scarabino, 1995 matchesthe description <strong>and</strong> illustration <strong>of</strong> Dentaliumhabutae Kuroda & Kikuchi, 1933 in all details <strong>of</strong>shell shape <strong>and</strong> sculpture (V. Scarabino pers.comm., <strong>and</strong> in litt. 14.I.1997). Dentaliumhabutae is considered a valid <strong>species</strong> <strong>of</strong>ZOOSYSTEMA • 2004 • 26 (4)711


Steiner G. & Kabat A. R.Entalinopsis (Habe, 1957: 13). Therefore, wecredit the synonymization <strong>of</strong> D. tessellatumScarabino, 1995 with Entalinopsis habutae(Kuroda & Kikuchi, 1933) to V. Scarabino.Since this <strong>species</strong> is known from its shell only, itssystematic position remains tentative.Cadulus (Cadulus) halius Henderson, 1920(p. 149) falls within the range <strong>of</strong> variation <strong>of</strong>Cadulus (Cadulus) podagrinus Henderson, 1920(p. 148, 149). Both taxa were described from thesame sample. Examination <strong>of</strong> the type materialshowed that the shells are identical except for thevarying length between the posterior aperture<strong>and</strong> the location <strong>of</strong> the maximum diameter.Because these differences are insignificant, wehave synonymized Cadulus halius Henderson,1920 with C. podagrinus Henderson, 1920.Dentalium (Antalis) ceratum tenax Henderson,1920 falls within the range <strong>of</strong> variation <strong>of</strong> thenominate <strong>species</strong>, Antalis cerata (Dall, 1881), <strong>and</strong>is, therefore, placed in synonymy with the nominate<strong>species</strong>.Cadulus (Platyschides) rushii arne Henderson,1920 falls within the range <strong>of</strong> variation <strong>of</strong> thenominate <strong>species</strong>, Polyschides rushii (Pilsbry &Sharp, 1898), <strong>and</strong> is, therefore, placed in synonymywith the nominate <strong>species</strong>.Cadulus (Gadila) agassizii var. hatterasensis Sharp& Pilsbry in Pilsbry & Sharp, 1898 falls withinthe range <strong>of</strong> variation <strong>of</strong> the nominate <strong>species</strong>,Gadila agassizii (Dall, 1881), <strong>and</strong> is, therefore,placed in synonymy with the nominate <strong>species</strong>.Dentalium bartonense Palmer, 1974 is an objectivejunior synonym <strong>of</strong> Dentalium acuticostataDeshayes, 1825. Both <strong>names</strong> are replacement<strong>names</strong> for Dentalium striatum J. Sowerby, 1814non Born, 1778. Palmer (1974b: 125) proposedDentalium bartonense as a replacement name,apparently being unaware that Deshayes hadalready renamed this <strong>species</strong>.APPENDIX 3Generic catalog <strong>of</strong> <strong>Recent</strong> <strong>species</strong>.The compilation <strong>of</strong> valid <strong>Recent</strong> <strong>species</strong> <strong>names</strong> istaken from Steiner & Kabat (2001) <strong>and</strong> modifiedaccording to the nomenclatural <strong>and</strong> taxonomicacts in this catalog. It alphabetically lists the<strong>species</strong> <strong>names</strong> for each genus, the authority foreach taxon, <strong>and</strong> the number <strong>of</strong> valid <strong>Recent</strong><strong>species</strong> for each genus in brackets.Annulipulsellum Scarabino, 1986 [1]euzkadii Scarabino, 1986Antalis H. & A. Adams, 1854 [41]aculeata (Sowerby, 1860)agilis (G. O. Sars, 1872)antillaris (d’Orbigny, 1853)bartletti (Henderson, 1920)berryi (Smith & Gordon, 1948)boucheti Scarabino, 1995cerata (Dall, 1881)circumcincta (Watson, 1879)dentalis (Linnaeus, 1758)diarrhox (Watson, 1879)disparile (d’Orbigny, 1853)entalis (Linnaeus, 1758)gardineri (Melvill, 1909)glaucarena (Dell, 1953)guillei Scarabino, 1995inaequicostata (Dautzenberg, 1891)inflexa (Sowerby, 1903)infracta (Odhner, 1931)intesi (Nicklès, 1979)longitrorsa (Reeve, 1843)marukawai (Otuka, 1933)minutissima (Ludbrook, 1954)nana (Hutton, 1873)novemcostata (Lamarck, 1818)occidentalis (Stimpson, 1851)panorma (Chenu, 1843)perinvoluta (Ludbrook, 1954)phanea (Dall, 1895)pilsbryi (Rehder, 1942)porcata (Gould, 1859)712ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)pretiosa (Sowerby, 1860)rossati (Caprotti, 1966)senegalensis (Dautzenberg, 1891)suteri (Emerson, 1954)taphria (Dall, 1889)tibana (Nomura, 1940)tubulata (Henderson, 1920)usitata (E. A. Smith, 1894)valdiviae (Plate, 1908)vulgaris (Da Costa, 1778)weinkauffi (Dunker, 1877)Anulidentalium Chistikov, 1975 [1]bambusa Chistikov, 1975Bathoxiphus Pilsbry & Sharp, 1897 [5]ensiculus (Jeffreys, 1877)inexpectatus Scarabino, 1995soyomaruae Okutani, 1964stanisici Lamprell & Healy, 1998steineri Lamprell & Healy, 1998Bathycadulus Scarabino, 1995 [1]fabrizioi Scarabino, 1995Cadulus Philippi, 1844 [54]aequatorialis Jaeckel, 1932amphora Jeffreys, 1883ampullaceus Watson, 1879aratus Hedley, 1899artatus Locard, 1897atlanticus Henderson, 1920attenuatus Monterosato, 1875californicus Pilsbry & Sharp, 1898campylus Melvill, 1906catharus Henderson, 1920chuni Jaeckel, 1932colliverae Lamprell & Healy, 1998congruens Watson, 1879cucurbitus Dall, 1881curtus Watson, 1879cyathoides Jaeckel, 1932cylindratus Jeffreys, 1877delicatulus Suter, 1913euloides Melvill & St<strong>and</strong>en, 1901exiguus Watson, 1879florenciae Scarabino, 1995gibbus Jeffreys, 1883glans Scarabino, 1995gracilis Jeffreys, 1877jeffreysi (Monterosato, 1875)labeyriei Scarabino, 1995lunulus Dall, 1881macleani Emerson, 1978minusculus Dall, 1889monterosatoi Locard, 1897obesus Watson, 1879occiduus Verco, 1911ovulum (Philippi, 1844)parvus Henderson, 1920platei Jaeckel, 1932platensis Henderson, 1920podagrinus Henderson, 1920propinquus G. O. Sars, 1878rossoi Nicklès, 1979rudmani Lamprell & Healy, 1998scarabinoi Steiner & Kabat, 2004siberutensis Jaeckel, 1932simillimus Watson, 1879simpsoni Henderson, 1920s<strong>of</strong>iae Scarabino, 1995subfusiformis (M. Sars, 1865)teliger Finlay, 1926tersus Henderson, 1920thielei Plate, 1908transitorius Henderson, 1920tumidosus Jeffreys, 1877valdiviae Jaeckel, 1932vincentianus Cotton & Godfrey, 1940woodhousae Lamprell & Healy, 1998Calliodentalium Habe, 1964 [4]balanoides (Plate, 1908)callipeplum (Dall, 1889)crocinum (Dall, 1907)semitracheatum (Boissevain, 1906)Chistikovia Scarabino, 1995 [1]kermadecae Scarabino, 1995Coccodentalium Sacco, 1896 [2]cancellatum (Sowerby, 1860)gemmiparum (Melvill, 1909)Compressidens Pilsbry & Sharp, 1897 [9]brevicornu (Sharp & Pilsbry in Pilsbry & Sharp, 1897)comprimatum (Plate, 1908)infimus Scarabino, 1995kikuchii (Kuroda & Habe, 1952)ophiodon (Dall, 1881)platyceras (Sharp & Pilsbry in Pilsbry & Sharp, 1897)pressum (Sharp & Pilsbry in Pilsbry & Sharp, 1897)stearnsii (Pilsbry & Sharp, 1898)wellsiana (Kraeuter, 1972)Compressidentalium Habe, 1963 [10]ceciliae Scarabino, 1995clathratum (von Martens, 1881)compressiusculum (Boissevain, 1906)hungerfordi (Pilsbry & Sharp, 1897)lardum (Barnard, 1963)sedecimcostatum (Boissevain, 1906)sibogae (Boissevain, 1906)subcurvatum (E. A. Smith, 1906)sumatrense (Plate, 1908)zanzibarense (Plate, 1908)Costentalina Chistikov, 1982 [7]caymanica Chistikov, 1982ZOOSYSTEMA • 2004 • 26 (4)713


Steiner G. & Kabat A. R.elegans Chistikov, 1982indica Chistikov, 1982leptoconcha Chistikov, 1982pacifica Chistikov, 1982tuscarorae Chistikov, 1982vemae Scarabino, 1986Dentalium Linnaeus, 1758 [68]aciculum Gould, 1859adenense Ludbrook, 1954agassizi Pilsbry & Sharp, 1897aprinum Linnaeus, 1767austini Lamprell & Healy, 1998bisexangulatum Sowerby, 1860buccinulum Gould, 1859burtonae Lamprell & Healy, 1998caledonicum Scarabino, 1995cheverti Sharp & Pilsbry in Pilsbry & Sharp, 1897clavus Cooke, 1885collinsae Lamprell & Healy, 1998congoensis Plate, 1908cookei Sharp & Pilsbry in Pilsbry & Sharp, 1897crosnieri Scarabino, 1995curtum Sowerby, 1860dacostianum Chenu, 1843decemcostatum Brazier, 1877deforgesi Scarabino, 1995elephantinum Linnaeus, 1758eupatrides Melvill & St<strong>and</strong>en, 1907exmouthensis Lamprell & Healy, 1998filosum Broderip & Sowerby, 1830garrardi Lamprell & Healy, 1998g<strong>of</strong>toni Lamprell & Healy, 1998gouldii Dall, 1889grahami Lamprell & Healy, 1998healyi Steiner & Kabat, 2004hedleyi Lamprell & Healy, 1998hillae Lamprell & Healy, 1998hyperhemileuron Verco, 1911invalidum Emerson, 1954javanum Sowerby, 1860jeanae Lamprell & Healy, 1998jelli Lamprell & Healy, 1998katchekense Fischer-Piette & Nicklès, 1946kathwayae Lamprell & Healy, 1998kessneri Lamprell & Healy, 1998laqueatum Verrill, 1885lessoni Deshayes, 1825letsonae Sharp & Pilsbry in Pilsbry & Sharp, 1897leucoryx Boissevain, 1906lochi Lamprell & Healy, 1998majorinum Mabille & Rochebrune in Rochebrune &Mabille, 1889mannarense Winckworth, 1927mediopacificensis Rehder & Ladd, 1973michelottii Hörnes, 1856neohexagonum Sharp & Pilsbry in Pilsbry & Sharp, 1897obscurum Dall, 1889obtusum Qi & Ma, 1989octangulatum Donovan, 1803oerstedii Mörch, 1861oryx Boissevain, 1906peitaihoensis King & Ping, 1935pluricostatum Boissevain, 1906potteri Lamprell & Healy, 1998reevei Fischer, 1871regulare E. A. Smith, 1903robustum Brazier, 1877rowei Lamprell & Healy, 1998scarabinoi Steiner & Kabat, 2004strigatum Gould, 1859stumkatae Lamprell & Healy, 1998tomlini Melvill, 1918vallicolens Raymond, 1904variabile Deshayes, 1825wellsi Lamprell & Healy, 1998woolacottae Colman, 1958Dischides Jeffreys, 1867 [13]atlantideus (Nicklès, 1955)belcheri Pilsbry & Sharp, 1898dartevellei (Nicklès, 1979)dichelus (Watson, 1879)hintoni Lamprell & Healy, 1998leloeuffi (Nicklès, 1979)minutus (H. Adams, 1872)ovalis (Boissevain, 1906)politus (S. Wood, 1842)prionotus (Watson, 1879)splendens Raines, 2002viperidens (Melvill & St<strong>and</strong>en, 1896)yateensis Scarabino, 1995Entalina Monterosato, 1872 [7]adenensis Ludbrook, 1954dorsicostata Lamprell & Healy, 1998inaequisculpta Ludbrook, 1954mirifica (E. A. Smith, 1895)platamodes (Watson, 1879)subterlineata (Tomlin, 1931)tetragona (Brocchi, 1814)Entalinopsis Habe, 1957 [4]habutae (Kuraoda & Kikuchi, 1933)intercostata (Boissevain, 1906)micra Scarabino, 1995stellata Scarabino, 1995Epirhabdoides Steiner, 1999 [1]ivanovi Steiner, 1999Episiphon Pilsbry & Sharp, 1897 [17]bordaensis (Cotton & Ludbrook, 1938)c<strong>and</strong>elatum (Kira, 1959)didymum (Watson, 1879)filum (Sowerby, 1860)fistula (Sowerby, 1860)gazellae (Plate, 1908)714ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)innumerabile (Pilsbry & Sharp, 1897)jamiesoni Lamprell & Healy, 1998kiaochowwanense (Tchang & Tsi, 1950)longum (Sharp & Pilsbry in Pilsbry & Sharp, 1897)pichoni Lamprell & Healy, 1998sominium (Okutani, 1964)sowerbyi (Guilding, 1834)subtorquatum (Fischer, 1871)truncatum (Boissevain, 1906)virginieae Scarabino, 1995virgula (Hedley, 1903)Eudentalium Cotton & Godfrey, 1933 [1]quadricostatum (Brazier, 1877)Fissidentalium Fischer, 1885 [41]actiniophorum Shimek, 1997aegeum (Watson, 1879)amphialum (Watson, 1879)c<strong>and</strong>idum (Jeffreys, 1877)capillosum (Jeffreys, 1877)carduus (Dall, 1889)ceras (Watson, 1879)complexum (Dall, 1895)concinnum (von Martens, 1878)cornubovis (E. A. Smith, 1906)edenensis Lamprell & Healy, 1998elizabethae Lamprell & Healy, 1998erosum Shimek & Moreno, 1996eualdes (Barnard, 1963)exasperatum (Sowerby, 1903)franklinae Lamprell & Healy, 1998horikoshii Okutani, 1982kawamurai Kuroda & Habe in Habe, 1961laterischismum Shikama & Habe, 1963levii Scarabino, 1995magnificum (E. A. Smith, 1896)malayanum (Boissevain, 1906)megathyris (Dall, 1890)metivieri Scarabino, 1995opacum (Sowerby, 1829)paucicostatum (Watson, 1879)peruvianum (Dall, 1908) n. comb.platypleurum (Tomlin, 1931)ponderi Lamprell & Healy, 1998pr<strong>of</strong>undorum (E. A. Smith, 1894)salpinx (Tomlin, 1931)semivestitum (Locard, 1897)serrulatum (E. A. Smith, 1906)shirleyae Lamprell & Healy, 1998shopl<strong>and</strong>i (Jousseaume, 1894)tenuicostatum Qi & Ma, 1989verconis (Cotton & Ludbrook, 1938)vicdani Kosuge, 1981waterhousae Lamprell & Healy, 1998yokoyamai (Makiyama, 1931)zel<strong>and</strong>icum (Sowerby, 1860)Fustiaria Stoliczka, 1868 [11]caesura (Colman, 1958)crosnieri Nicklès, 1979engischista (Barnard, 1963)gruveli (Dautzenberg, 1910)langfordi (Habe, 1963)liodon (Pilsbry & Sharp, 1897)nipponica (Yokoyama, 1922)polita (Linnaeus, 1767)rubescens (Deshayes, 1825)stenoschiza (Pilsbry & Sharp, 1897)vagina Scarabino, 1995Gadila Gray, 1847 [63]aberrans (Whiteaves, 1887)abruptoinflata (Boissevain, 1906)acuminata (Tate, 1887)acus (Dall, 1889)aequalis (Dall, 1881)agassizii (Dall, 1881)amianta (Dall, 1889)anguidens (Melvill & St<strong>and</strong>en, 1898)angustior (Verco, 1911)arctus (Henderson, 1920)austinclarki (Emerson, 1951)boissevainae (Jaeckel, 1932)bordaensis (Cotton & Godfrey, 1940)boucheti (Nicklès, 1979)braziliensis (Henderson, 1920)brycei Lamprell & Healy, 1998bushii (Dall, 1889)carlessi Lamprell & Healy, 1998clavata (Gould, 1859)cobbi Lamprell & Healy, 1998desaintlaurentae Scarabino, 1995divae (Vélain, 1877)dominguense (d’Orbigny, 1853)doumenci Scarabino, 1995elenae Scarabino, 1995elephas (Henderson, 1920)fraseri Nicklès, 1955gadus (Montagu, 1803)greenlawi (Henderson, 1920)honoluluensis (Watson, 1879)iota (Henderson, 1920)ludbrookae (Cotton & Godfrey, 1940)marchadi (Nicklès, 1979)mayori (Henderson, 1920)minutalis Scarabino, 1995monodonta Scarabino, 1995nicklesi (Dell, 1964)opportuna Kuroda & Habe in Habe, 1961p<strong>and</strong>ionis (Verrill & Smith in Verrill, 1880)perpusilla (Sowerby in Broderip & Sowerby, 1832)peruviana (Dall, 1908)platystoma (Pilsbry & Sharp, 1898)providensis (Henderson, 1920)pseudolivae (Boissevain, 1906)rastridens (Watson, 1879)regularis (Henderson, 1920)sagamiensis Kuroda, Habe & Oyama, 1971ZOOSYSTEMA • 2004 • 26 (4)715


Steiner G. & Kabat A. R.sauridens (Watson, 1879)senegalensis (Locard, 1897)singaporensis (Sharp & Pilsbry in Pilsbry & Sharp, 1898)spreta (Tate & May, 1900)strangulata (Locard, 1897)striata (Pilsbry & Sharp, 1898)subcolubridens (Ludbrook, 1954)subtilis (Plate, 1908)subula (Henderson, 1920)tolmiei (Dall, 1897)verrilli (Henderson, 1920)virginalis (Boissevain, 1906)vulpidens (Watson, 1879)watsoni (Dall, 1881)whitneyae Lamprell & Healy, 1998zonata (Boissevain, 1906)Gadilina Foresti, 1895 [2]insolita (E. A. Smith, 1894)pachypleura (Boissevain, 1906)Graptacme Pilsbry & Sharp, 1897 [16]acutissima (Watson, 1879)acutistriata Steiner & Kabat, 2004africana (Sowerby, 1903)calamus (Dall, 1889)eborea (Conrad, 1846)elpis (Winckworth, 1927)inversa (Deshayes, 1825)lactea (Deshayes, 1825)marchadi (Nicklès, 1979)nielseni Lamprell & Healy, 1998novaeholl<strong>and</strong>iae (Chenu, 1843)perlonga (Dall, 1881)secta (Deshayes, 1825)semistriata (Turton, 1819)splendida (Sowerby in Broderip & Sowerby, 1832)takakoae Tsuchida & Tachi, 1999Heteroschismoides Ludbrook, 1960 [1]subterfissus (Jeffreys, 1877)Laevidentalium Cossmann, 1888 [18]ambiguum (Chenu, 1843)arnoldi Lamprell & Healy, 1998banale (Boissevain, 1906)bisinuatum (André, 1896)coruscum (Pilsbry, 1905)eburneum (Linnaeus, 1767)ensiforme (Chenu, 1842)erectum (Sowerby, 1860)g<strong>of</strong>asi Scarabino, 1995houbricki Scarabino, 1995jaffaensis (Cotton & Ludbrook, 1938)largicrescens (Tate, 1899)lebruni (Mabille & Rochebrune in Rochebrune &Mabille, 1889)leptosceles (Watson, 1879)lubricatum (Sowerby, 1860)marshae Lamprell & Healy, 1998martyi Lamprell & Healy, 1998zeidleri Lamprell & Healy, 1998Megaentalina Habe, 1963 [2]cornucopiae (Boissevain, 1906)mediocarinata (Boissevain, 1906)Omniglypta Kuroda & Habe in Habe, 1953 [1]cerina (Pilsbry, 1905)Paradentalium Cotton & Godfrey, 1933 [14]americanum (Chenu, 1843) n. comb.angustistriatum Chistikov, 1979flindersi (Cotton & Ludbrook, 1938)francisense (Verco, 1911)gradile Chistikov, 1979hemileuron (Verco, 1911)hexagonum (Gould, 1859)intercalatum (Gould, 1859)katowense (Brazier, 1877)natalense (Barnard, 1963)octopleuron (Verco, 1911)pistis (Winckworth, 1940)pseudosexagonum (Deshayes, 1825)rudoi Scarabino, 1995Pertusiconcha Chistikov, 1982 [2]callithrix (Dall, 1889)tridentata Chistikov, 1982Pictodentalium Habe, 1963 [3]festivum (Sowerby, 1914)formosum (A. Adams & Reeve, 1850)vernedei (Hanley in Sowerby, 1860)Plagioglypta Pilsbry in Pilsbry & Sharp, 1898 [1]pertracheata (Plate, 1908)Polyschides Pilsbry & Sharp, 1898 [25]<strong>and</strong>ersoni Lamprell & Healy, 1998arnaudi Scarabino, 1995carolinensis (Bush in Verrill, 1885)cuspidatus (Nicklès, 1979)elongatus (Henderson, 1920)fausta Kuroda, Habe & Oyama, 1971foweyensis (Henderson, 1920) n. comb.gibbosus (Verco, 1911)gr<strong>and</strong>is (Verrill, 1884)kaiyomaruae Okutani, 1975miamiensis (Henderson, 1920)nedallisoni (Emerson, 1978)nitidus (Henderson, 1920) n. comb.olivi (Scacchi, 1835)pelamidae Chistikov, 1979poculum (Dall, 1889) n. comb.portoricensis (Henderson, 1920) n. comb.quadrifissatus (Pilsbry & Sharp, 1898)rushii (Pilsbry & Sharp, 1898)716ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)sakuraii (Kuroda & Habe in Habe, 1961)spectabilis (Verrill, 1885)sutherl<strong>and</strong>i Lamprell & Healy, 1998tetraschistus (Watson, 1879)tetrodon (Pilsbry & Sharp, 1898)vietnamicus Chistikov, 1979Pulsellum Stoliczka, 1868 [15]affine (M. Sars, 1865)beecheyi Lamprell & Healy, 1998bushi (Henderson, 1920)eboracense (Watson, 1879)falkl<strong>and</strong>icum (Dell, 1964)fragile Scarabino, 1995hige Habe, 1963kurogenge Habe & Kosuge, 1964l<strong>of</strong>otense (M. Sars, 1865)occidentale (Henderson, 1920)pusillum (Watson, 1879)salishorum Marshall, 1980teres (Jeffreys, 1883)thomassini Scarabino, 1995verrilli (Henderson, 1920)Rhabdus Pilsbry & Sharp, 1897 [5]aequatorius (Pilsbry & Sharp, 1897)dalli (Pilsbry & Sharp, 1897)perceptus (Mabille & Rochebrune in Rochebrune &Mabille, 1889)rectius (Carpenter, 1864)toyamaense (Kuroda & Kikuchi, 1933)Rhomboxiphus Chistikov, 1983 [2]colmani (Palmer, 1974)tricarinatus (Boissevain, 1906)Sagamicadulus Sakurai & Shimazu, 1963 [1]elegantissimus Sakurai & Shimazu, 1963Schizodentalium Sowerby, 1894 [1]plurifissuratum Sowerby, 1894Siphonodentalium M. Sars, 1859 [16]australasiae Boissevain, 1906booceras (Tomlin, 1926)colubridens (Watson, 1879)dalli (Pilsbry & Sharp, 1898)hexaschistum (Boissevain, 1906)isaotakii Habe, 1953jaeckeli Scarabino, 1995japonicum Habe, 1960laubieri Bouchet & Warén, 1979lobatum (Sowerby, 1860)longilobatum (Boissevain, 1906)magnum (Boissevain, 1906)okudai Habe, 1953promontorii (Barnard, 1963)summa (Okutani, 1964)tytthum Watson, 1879Solenoxiphus Chistikov, 1983 [1]striatulus Chistikov, 1983Spadentalina Habe, 1963: 264 [2]ingrata Scarabino, 1995tubiformis (Boissevain, 1906)Striocadulus Emerson, 1962 [4]albicomatus (Dall, 1890)ludbrookae Scarabino, 1995pulcherrimus (Boissevain, 1906)sagei Scarabino, 1995Striodentalium Habe, 1964 [8]chinensis Qi & Ma, 1989concretum (Colman, 1958)hosoi (Habe, 1963)kanakorum Scarabino, 1995polycostatum Qi & Ma, 1989rhabdotum (Pilsbry, 1905)thetidis (Hedley, 1903)tosaensis (Habe, 1963)Striopulsellum Scarabino, 1995 [3]galatheae (Knudsen, 1964)minimum (Plate, 1908)striatinum (Henderson, 1920)Tesseracme Pilsbry & Sharp, 1898 [7]dispar (Sowerby, 1860)hancocki (Emerson, 1956)philcolmani Lamprell & Healy, 1998quadrangularis (Sowerby in Broderip & Sowerby,1832)quadrapicalis (Hanley in Sowerby, 1860)tesseragona (Sowerby in Broderip & Sowerby, 1832)tetrapleura (Boissevain, 1906)Wemersoniella Scarabino, 1986 [4]duartei Scarabino, 1986indica Scarabino, 1995knudseni Scarabino, 1995turnerae Scarabino, 1986ZOOSYSTEMA • 2004 • 26 (4)717


Steiner G. & Kabat A. R.APPENDIX 4Geological catalog <strong>of</strong> <strong>fossil</strong> <strong>species</strong>.The <strong>fossil</strong> <strong>species</strong> that are currently treated as valid– i.e. neither junior synonyms nor juniorhomonyms – are indexed by geological period(Paleozoic <strong>and</strong> Mesozoic) <strong>and</strong> epoch (Cenozoic).However, it must be emphasized that some <strong>of</strong> thisinformation is based on the older literature, whichmay have been based on erroneous or outdatedstratigraphic determinations. A question mark (“?”)before a <strong>species</strong> name indicates a doubtful placement;at the end are those elusive <strong>species</strong> describedonly as “<strong>fossil</strong>” but without further stratigraphicdetails. Many <strong>of</strong> the Pleistocene <strong>and</strong> some <strong>of</strong> thePliocene <strong>species</strong> are also known from the <strong>Recent</strong>, asindicated in the main catalog. Generic placementsare as in the original description, except for thoserelatively few <strong>fossil</strong> <strong>species</strong> that were subsequentlytransferred to other genera. The sequence <strong>of</strong> geologicaleras, periods, <strong>and</strong> epochs, <strong>and</strong> the assignments<strong>of</strong> stages <strong>and</strong> other subdivisions to epochs, istaken from Harl<strong>and</strong> et al. (1990).It must be emphasized that the majority <strong>of</strong> <strong>fossil</strong><strong>species</strong> listed herein have not been analyzed in anysubstantive manner since the time <strong>of</strong> Pilsbry &Sharp (1897-1898). Hence, their inclusion in thisAppendix is merely a compilation <strong>of</strong> nomenclaturallyavailable <strong>names</strong> arranged by stratigraphictime periods. Thus, the numbers <strong>of</strong> <strong>species</strong> <strong>names</strong>for each time period should not be taken as anactual measure <strong>of</strong> paleontological diversity, whichwill require a modern determination <strong>of</strong> the phylogeneticvalidity <strong>of</strong> each described <strong>species</strong>.PALEOZOICCambrianNoneOrdovicianiowaensis, Plagioglypta (James, 1890)kentuckyensis, Rhytiodentalium Pojeta & Runnegar,1979Siluriangranosum, Dentalium Eichwald, 1857notabile, Dentalium Eichwald, 1857Devonianapplanatum, Dentalium Torley, 1908crox, Dentalium Keyserling, 1847ibergense, Dentalium Römer, 1855patriciae, Laevidentalium Engeser & Riedel, 1992?saturni, Dentalium Hoeninghaus, 1836?tenuissimum, Dentalium de Koninck, 1876Carboniferousacumen, Dentalium (de Koninck, 1883)acutisulcatum, Dentalium Gurley, 1883annuliferum, Dentalium Pilsbry & Sharp, 1898annulostriatum, Dentalium Meek & Worthen, 1871barquense, Dentalium Winchell, 1862batheri, Dentalium Finlay, 1927chubutensis, Dentalium (Laevidentalium) Sabattini,1979cornu, Dentalium de Koninck, 1877cyrtoceratoides, Dentalium (de Koninck, 1883)dentaloideum, Dentalium (Phillips, 1836)gr<strong>and</strong>aevum, Dentalium Winchell, 1863herculea, Plagioglypta (de Koninck, 1863)illinoiensis, Dentalium Worthen, 1883ingens, Plagioglypta (de Koninck, 1843)kansasense, Dentalium (Paleodentalium) Gentile,1974longistriatum, Fissidentalium “?” Yoo, 1988meekiana, Plagioglypta (Geinitz, 1866)mexicanum, Dentalium Girty, 1909missouriense, Dentalium Swallow, 1863navicanum, Dentalium Ryckholt, 1851novaresei, Dentalium (Plagioglypta) de Stefani, 1917numerosa, Plagioglypta Yoo, 1988obsoletum, Prodentalium (Hall, 1858)ornatum, Dentalium de Koninck, 1843orthoceras, Dentalium Pilsbry & Sharp, 1898perarmatum, Dentalium Ryckholt, 1851primarium, Dentalium Hall, 1858priscum, Dentalium Goldfuss, 1841prosseri, Plagioglypta Morningstar, 1922protuberans, Pipadentalium Yoo, 1988raymondi, Prodentalium Young, 1942rectiusculum, Dentalium Eichwald, 1857runnegari, Scissuradentalium Yoo, 1988semicostatum, Prodentalium (Girty, 1911)subannulata, Plagioglypta Easton, 1962subcanaliculatum, Dentalium S<strong>and</strong>berger &S<strong>and</strong>berger, 1856taeniolatum, Dentalium S<strong>and</strong>berger & S<strong>and</strong>berger,1856venustum, Dentalium Meek & Worthen, 1861verrucosum, Dentalium Eichwald, 1857walciodorense, Dentalium (de Koninck, 1883)718ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)Permianakasakensis, Dentalium Hayasaka, 1925arcturus, Calstevenus Yancey, 1973canna, Prodentalium (White, 1874)furcata, Plagioglypta “?” Waterhouse, 1980girtyi, Plagioglypta Knight, 1940komiensis, Laevidentalium “?” Kulikov, 1967lanceolata, Theca Morris, 1845neornatum, Dentalium Hayasaka, 1925robusta, Plagioglypta Detre, 1993sorbii, Dentalium King, 1850sueroi, Calstevenus Sabattini, 1979wymensis, Laevidentalium “?” Kulikov, 1967“Paleozoic” (period not specified)arenarium, Dentalium Römer, 1855granvillensis, Dentalium Herrick, 1888 (“Subcarboniferous”)striatus, Prodentalium (Romanovskii, 1890)MESOZOICTriassicangelae, Prodentalium Engeser & Riedel, 1992anisicum, Dentalium Schnetzer, 1934arctoides, Dentalium Gugenberger, 1934arctum, Dentalium Pichler, 1857bangtoupoensis, Laevidentalium Stiller, 2001boreale, Dentalium Böhm, 1904cannaliculatum, Dentalium Klipstein, 1843castellacei, Dentalium Scalia, 1914cyrtoceroides, Dentalium Broili, 1907decoratum, Dentalium Goldfuss, 1841exile, Dentalium Tommasi, 1901gallensteini, Dentalium Gugenberger, 1934guizhouensis, Plagioglypta Stiller, 2001herritschi, Dentalium Gugenberger, 1934kahleri, Dentalium Gugenberger, 1934kittli, Dentalium Cossmann, 1900klipsteini, Dentalium Kittl, 1891laevis, Entalis (Schlotheim, 1820)?lombardicum, Dentalium Kittl, 1899multiclathratum, Dentalium Martelli, 1905pentagonale, Dentalium Pilsbry & Sharp, 1898qingyanicum, Prodentalium Stiller, 2001quadrangulum, Dentalium Assmann, 1937readi, Dentalium Emerson, 1954regium, Dentalium Assmann, 1937schafferi, Dentalium Gugenberger, 1934simile, Dentalium Wissmann, 1841spitiense, Dentalium Gümbel, 1866subregulare, Dentalium Palmer, 1974torquata, Entalis (Schlotheim, 1820)undulatum, Dentalium Goldfuss, 1841Jurassicamalthei, Laevidentalium (Engel, 1891)<strong>and</strong>leri, Dentalium Oppel, 1856angulatum, Dentalium Buckman in Murchison, 1845b<strong>and</strong>eli, Prodentalium Engeser & Riedel, 1992boonei, Dentalium Cossmann, 1924calvertensis, Prodentalium Palmer, 1975cinctum, Dentalium Goldfuss, 1841corallinum, Dentalium d’Orbigny, 1850dehmi, Dentalium Kuhn, 1935etalense, Laevidentalium (Terquem & Piette, 1865)filicauda, Dentalium Quenstedt, 1852glabellum, Dentalium Blake, 1907gladiolus, Dentalium Eichwald, 1846hexagonale, Dentalium Richardson, 1906jaegeri, Suevidontis Engeser, Riedel & B<strong>and</strong>el, 1993liassicum, Prodentalium (Moore, 1866)limatuloides, Baltodentalium (Kuhn, 1935)limatulum, Baltodentalium (Tate, 1870)minimum, Laevidentalium (Buckman in Murchison,1845)moreanum, Dentalium d’Orbigny, 1845normanianum, Dentalium d’Orbigny, 1850nudum, Dentalium Zekeli, 1852oblongum, Dentalium Richardson, 1906oolithicum, Dentalium Piette, 1856opalina, Dentalium filicauda var. Quenstedt, 1856ovale, Dentalium “Malm.” Pilsbry & Sharp, 1898parkinsoni, Dentalium Quenstedt, 1852pellati, Laevidentalium (Loriol in Loriol & Pellat,1874)portlocki, Dentalium Tate, 1870quenstedti, Dentalium Blake, 1875richardsoni, Laevidentalium (Cossmann, 1907)rigauxi, Fustiaria (Laevidentalium) Fischer & Vadet,1985sabrinae, Dentalium Engeser & Riedel, 1992soliticum, Dentalium Piette, 1855spaethi, Progadilina Engeser & Riedel, 1992subovata, Progadilina (Richardson, 1906)subparvulum, Laevidentalium (Emerson, 1954)subquadratum, Dentalium Meek, 1860subtrigonalis, Progadilina (Richardson, 1906)tenue, Dentalium Goldfuss, 1841tenuistriata, Cyrtoconella Patrulius, 1996terquemi, Dentalium Richardson, 1906triedra, Serpula Quenstedt, 1856trigonalis, Progadilina “?” (Moore, 1866)weitschati, Baltodentalium Engeser & Riedel, 1992Cretaceousalatum, Dentalium Gardner, 1878alineatum, Dentalium Stephenson, 1952angulare, Dentalium Kaunhowen, 1897aquensis, Cadulus (Holzapfel, 1888)arcotinum, Dentalium Forbes, 1846barremicum, Dentalium Cossmann, 1916binkhorsti, Dentalium Pilsbry & Sharp, 1898californicum, Dentalium Stanton, 1895cazadorianum, Dentalium Wilckens, 1907chilense, Dentalium d’Orbigny in Grange, 1847ZOOSYSTEMA • 2004 • 26 (4)719


Steiner G. & Kabat A. R.cidaris, Dentalium Geinitz, 1850coelatulum, Dentalium Baily in Salter, 1857confusum, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898coonensis, Cadulus Stephenson, 1941cooperi, Dentalium Gabb, 1864crassulum, Dentalium Stoliczka, 1868cretaceum, Dentalium Conrad, 1852cretaustralium, Laevidentalium Stilwell, 1999curva, Entalina (Gardner, 1878)decemcostatum, Dentalium Quaas, 1902decussatum, Dentalium J. Sowerby, 1814dilatatum, Dentalium Philippi, 1887divisiense, Dentalium Gardner, 1878duodecimplicatum, Dentalium Repelin, 1907elongatum, Laevidentalium (Goldfuss, 1841)ezoense, Dentalium cooperi var. Nagao, 1938fragilis, Dentalium Meek & Hayden, 1856frontierensis, Cadulus Sidwell, 1932gardneri, Entalina Sharp & Pilsbry in Pilsbry & Sharp,1898gardneri, Fustiaria (Holzapfel, 1888)gaultinus, Cadulus Gardner, 1878geinitzi, Dentalium (Böhm, 1885)geinitzianum, Dentalium Ryckholt, 1851glabratum, Antale Stoliczka, 1868gracilis, Dentalium Hall & Meek, 1855gr<strong>and</strong>is, Eodentalium Medina & del Valle, 1985groenl<strong>and</strong>icum, Dentalium Ravn, 1918groenl<strong>and</strong>icus, Cadulus Ravn, 1918gucuoensis, Dentalium Yao, 1990hexapleuron, Dentalium Kaunhowen, 1897infortunatum, Dentalium Pilsbry & Sharp, 1898jeffreysi, Dentalium Gardner, 1878kubraense, Dentalium Palmer, 1974lamberti, Dentalium Cossmann, 1897laticostatum, Dentalium Reuss, 1844leve, Dentalium Stephenson, 1923limatum, Dentalium (Laevidentalium) Stanton, 1901lineatum, Dentalium Gueranger, 1853major, Dentalium Gardner, 1877medium, Dentalium J. Sowerby, 1814meyeri, Entalis Gardner, 1878michauxianum, Dentalium Ryckholt, 1851minorinum, Dentalium Steiner & Kabat, 2004 n. nom.morganianum, Dentalium (Laevidentalium) Wilckens,1922muellerianum, Dentalium Pilsbry & Sharp, 1898multicanaliculatum, Dentalium Gümbel, 1861multicostatum, Dentalium Favre, 1869navarroi, Dentalium Stephenson, 1941niceforoi, Dentalium Royo y Gomez, 1945nodulosus, Dentalites Schlotheim, 1820nutans, Cadulus Böhm, 1891obrutus, Cadulus (Conrad, 1870)octevillense, Dentalium Cossmann, 1899otatumei, Dentalium Nagao, 1938ovosectum, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898parvula, Fustiaria Stoliczka, 1868patulum, Dentalium Potiez & Michaud, 1838paulini, Dentalium Maury, 1925pauperculum, Dentalium Meek & Hayden, 1860planicostatum, Dentalium Hébert, 1855polygonum, Dentalium Reuss, 1844praetenuis, Cadulus Stephenson, 1952rhodani, Dentalium Pictet & Roux, 1849rothomagense, Dentalium d’Orbigny, 1850ripleyanum, Dentalium Gabb, 1860rugosum, Dentalium Dunker, 1849sacheri, Dentalium Alth, 1850schlosseri, Dentalium Traub, 1938sougraignense, Dentalium Cossmann, 1904stramineum, Dentalium Gabb, 1864subarcuatum, Dentalium Conrad, 1853subcylindricum, Dentalium Philippi, 1887sublineatum, Dentalium Stephenson, 1952subplanum, Dentalium Pilsbry & Sharp, 1898tenuicostatum, Dentalium Böhm, 1891turoniense, Dentalium Woods, 1896uchauxiense, Dentalium Cossmann, 1897valangiense, Dentalium Pictet & Campiche, 1864vaningeni, Dentalium Steiner & Kabat, 2004 n. nom.vaughani, Dentalium Stephenson, 1941wadei, Dentalium Pilsbry, 1927wanneri, Dentalium Steiner & Kabat, 2004 n. nom.whiteavesi, Dentalium (Entalis) Anderson & Hanna,1935wilckensi, Laevidentalium Medina & del Valle, 1985wollumbillaensis, Dentalium Etheridge in Jack &Etheridge, 1892“Mesozoic” (period not specified)?elephantinus, Dentalites Schlotheim, 1820subquadrangulare, Dentalium Emerson, 1954CENOZOICPaleogene [Paleocene to Oligocene, epoch notspecified]pagellai, Dentalium Negri, 1934Paleoceneashiyaensis, Dentalium Nagao, 1928ewekoroense, Dentalium (Antalis) Adegoke, 1977golovtschenki, Dentalium Makarenko, 1970guineense, Dentalium (Laevidentalium) Adegoke,1977infraeocaenicanum, Compressidens (Cossmann, 1907)intumescens, Cadulus (Koenen, 1885)laguncula, Gadila Darragh, 1997laticornuata, Compressidens Darragh, 1997lusanensis, Dentalium Makarenko, 1970mauryae, Dentalium Penna-Neme & da Costa BarrosMuniz, 1976720ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)montense, Fustiaria (Briart & Cornet, 1889) [asD. inversum Cossmann, 1915 non Deshayes]multistricta, Antalis (Finlay & Marwick, 1937)nigeriense, Dentalium (Laevidentalium) Adegoke, 1977rarinodosum, Dentalium haeringense var. Báldi,Hovarth & Makk, 1974rugiferum, Dentalium Koenen, 1885seriaeforme, Siphonodentalium Ravn, 1939subgracile, Dentalium Emerson, 1954sundkrogensis, Dentalium (Dentalium) Schnetler, 2001undiferum, Dentalium Koenen, 1885Eoceneabsconditum, Dentalium Deshayes, 1861acicula, Fustiaria (Episiphon) Hodgkinson, 1974acre, Dentalium Sharp & Pilsbry in Pilsbry & Sharp,1898acuticostata, Dentalium Deshayes, 1825aequale, Dentalium Deshayes, 1861aldrichi, Cadulus Gardner, 1933anceps, Dentalium J. Sowerby in Wetherell, 1837<strong>and</strong>rewsi, Dentalium Cuvillier, 1933angustum, Dentalium Deshayes, 1861annulopunctatum, Dentalium (Fustiaria) Cossmann &Pissarro, 1902apicostatum, Dentalium (Antalis) Vokes, 1939arabicum, Dentalium Cuvillier, 1933archiaci, Dentalium Tournouër in de Boullé, 1873arciformis, Dentalium Conrad, 1846armoricense, Siphonodentalium Cossmann, 1902arnoensis, Polyschides Maxwell, 1992atavus, Cadulus (Gadila) Finlay & Marwick, 1937bellulus, Cadulus Clark, 1895bilabiatus, Dischides (Deshayes, 1861)bironense, Dentalium Douvillé, 1929bisissura, Cadulus “(?)” Garvie, 1996bitubatum, Dentalium Meyer, 1886bl<strong>and</strong>um, Dentalium de Gregorio, 1890boggsi, Dentalium Berry, 1926bouryi, Siphonodentalium (Dischides) Cossmann, 1888brazosensis, Cadulus (Gadila) Hodgkinson, 1974breve, Dentalium Deshayes, 1861brevis, Dischides (Deshayes, 1861)brongniarti, Dentalium Deshayes, 1861calafium, Dentalium (Laevidentalium) Vokes, 1939castellanensis, Dentalium d’Orbigny, 1850centenniale, Dentalium Marwick, 1942circinatum, Fustiaria (G. B. Sowerby, 1823)claibornense, Dentalium (Antalis) thalloides var. Palmer,1937coislinense, Dentalium (Entalis) Cossmann, 1902constricta, Antalis (Newton & Harris, 1894)corpulentus, Cadulus Meyer, 1886curvus, Cadulus (Gadila) Hodgkinson, 1974danai, Fustiaria (Laevidentalium) (Meyer, 1885)danvillense, Dentalium (Antalis) Palmer, 1947defrancii, Dentalium Deshayes, 1861depressus, Cadulus Meyer, 1884dilatatum, Pulsellum Cossmann, 1888dolichus, Cadulus (Gadilopsis) Woodring, 1973dumasi, Fustiaria “(?)” Cossmann, 1919duplex, Lobantale (Defrance, 1819)emersoni, Coccodentalium Squires, 1988emersoni, Entalina Maxwell, 1992erleneae, Cadulus (Gadila) Hodgkinson, 1974eugenii, Dentalium Dall, 1892gabbi, Cadulus Pilsbry & Sharp, 1898ganense, Laevidentalium Cossmann in O’Gorman &Cossmann, 1923gnizum, Dentalium de Gregorio, 1890gracilicostatum, Dentalium (Fissidentalium) Singleton,1943gracilis, Episiphon Garvie, 1996gr<strong>and</strong>e, Dentalium Deshayes, 1825grossheimi, Dentalium Merklin, 1961hannonicum, Dentalium Briart & Cornet, 1889ignotum, Dentalium Korobkov, 1955incisissimum, Dentalium Meyer & Aldrich, 1886inopinatum, Dentalium Mayer-Eymar, 1887jacksonense, Dentalium (Antalis) mississippiense var.Palmer, 1947jacksonensis, Cadulus Meyer, 1885juvenis, Cadulus Meyer, 1886kahirahense, Dentalium Cossmann, 1901kapuaensis, Polyschides Maxwell, 1992korobkovi, Cadulus (Gadila) Merklin, 1961l<strong>and</strong>inense, Dentalium Vincent, 1876leoniae, Dentalium Meunier, 1878leroyi, Fustiaria (Fustiaria) Hodgkinson, 1974lighti, Dentalium (Laevidentalium “?”) Vokes, 1939lucidum, Pseudantalis (Deshayes, 1861)magnum, Dentalium Briart & Cornet, 1889magnus, Gadus Mayer-Eymar, 1887mancorens, Dentalium Olsson, 1930mantelli, Dentalium Zittel, 1864maoria, Fustiaria Maxwell, 1992margarita, Cadulus (Polyschides) Palmer, 1947marthae, Dentalium Cossmann & Pissarro, 1902masoni, Gadilina Maxwell, 1992mcfarl<strong>and</strong>ae, Cadulus (Cadulus) Emerson, 1957mediaviense, Dentalium Harris, 1896meganosensis, Cadulus Palmer, 1923meyeri, Polyschides (Cossmann, 1888)michelinii, Dentalium Rouault, 1850microstria, Dentalium Heilprin, 1881minutiannulatum, Dentalium (Antalis) Hodgkinson,1974minutistriatum, Dentalium Gabb, 1860molengraaffi, Dentalium (s.s.) Martin, 1915montense, Dentalium Briart & Cornet, 1889moseleyensis, Cadulus (Gadila) Hodgkinson, 1974multannulatum, Dentalium Aldrich, 1895nanggulanense, Dentalium (s.s.) Martin, 1915neglectum, Dischides (Cossmann, 1888)newtonensis, Cadulus Meyer & Aldrich, 1886nitens, Fustiaria (J. Sowerby, 1814)nitidum, Laevidentalium (Deshayes, 1861)ZOOSYSTEMA • 2004 • 26 (4)721


Steiner G. & Kabat A. R.orthezense, Dentalium Tournouër, 1873ouachitensis, Cadulus (Cadulus) Palmer, 1937palmerae, Cadulus (Gadila) Hodgkinson, 1974palmerae, Dentalium (Antalis) Garvie, 1996parannulatum, Dentalium Steiner & Kabat, 2004n. nom.parisiensis, Dentalium d’Orbigny, 1850pellucens, Fustiaria (Deshayes, 1861)perinii, Dentalium Malaroda, 1951phoenicea, Cadulus Gardner, 1933potiezi, Dentalium (s.s.) Le Renard, 1994pseudocircinata, Fustiaria Gougerot, 1968pseudohungaricus, Cadulus (Gadila) Szöts, 1953pseudorakhiense, Dentalium (Laevidentalium) Eames,1952pukaea, Fissidentalium Maxwell, 1992quadriturritus, Cadulus Meyer, 1886rakhiense, Dentalium (Laevidentalium) Eames, 1952rei, Fissidentalium “?” Maxwell, 1992reticulatum, Dentalium Tournouër in de Boullé, 1873samanicum, Dentalium Berry, 1926?s<strong>and</strong>bergeri, Dentalium Bosquet, 1859seminudum, Dentalium Deshayes, 1861soriense, Dentalium Eames, 1952speciosum, Dentalium Gümbel, 1861stentor, Dentalium Anderson & Hanna, 1925stonecityensis, Cadulus (Gadila) Hodgkinson, 1974striatulinum, Antale “?” Cossmann in O’Gorman &Cossmann, 1923subcoarcuatus, Cadulus (Polyschides) (Gabb, 1860)subcompressa, Fustiaria (Laevidentalium) (Meyer,1885)subeburneum, Fustiaria (d’Orbigny, 1850)subfissura, Dentalium (Laevidentalium) (Tate, 1887)substriatum, Dentalium Deshayes, 1825sylvaerupis, Serpulorbis Harris, 1899tatei, Dentalium Sharp & Pilsbry in Pilsbry & Sharp,1898tenuicincta, Plagioglypta (Gougerot, 1968)tenuis, Dentalium Hutton, 1873tenuistriatum, Dentalium Rouault, 1850thalloides, Dentalium Conrad, 1833turgidus, Cadulus Meyer, 1886turritum, Dentalium Lea, 1833vacavillensis, Dentalium Palmer, 1923vincense, Dentalium (Antalis) Palmer, 1947vredenburgi, Dentalium Cossmann & Pissarro, 1909xiphias, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898yasilum, Dentalium Olsson, 1930zingulai, Cadulus (Gadila) Hodgkinson, 1974Oligoceneabruptus, Cadulus Meyer & Aldrich, 1886acuta, Antalis (Hébert, 1849)aequicostatum, Dentalium Koenen, 1892alazanum, Dentalium (Dentalium) Cooke, 1928asphaltodes, Dentalium (Dentalium) Beets, 1943boussaci, Dentalium Emerson, 1954callioglyptum, Dentalium Pilsbry & Sharp, 1898cartagenense, Dentalium Pilsbry & Brown, 1917caseyi, Dentalium Steiner & Kabat, 2004 n. nom.casselensis, Gadila R. Janssen, 1989chipolanum, Dentalium (Antalis) Gardner, 1947clarae, Cadulus Maury, 1910colobus, Cadulus Pilsbry & Sharp, 1898cossmannianum, Dentalium Pilsbry & Sharp, 1898cucumis, Cadulus Koenen, 1892cylindricus, Cadulus R. Janssen, 1989dejtarense, Dentalium densitextum var. Báldi, 1973dekagonum, Dentalium Koenen, 1892densitexta, Dentalium haeringense var. Noszky, 1940depressicollis, Cadulus Pilsbry & Sharp, 1898diopon, Dentalium (Antalis) Gardner, 1947duodecimcostatum, Dentalium d’Archiac, 1850durhami, Cadulus (Platyschides) Emerson, 1957elegantissimus, Cadulus Pilsbry & Sharp, 1898exiguum, Dentalium Koenen, 1892fritschi, Dentalium Koenen, 1892gabbi, Dentalium Pilsbry & Sharp, 1898geinitzi, Dentalium (Fustiaria) Koenen, 1892geminatum, Antalis (Goldfuss, 1841)heptagonum, Dentalium Boettger, 1882infraoligocaenicum, Compressidens Welle, 1997insuranum, Dentalium Emerson, 1954kickxii, Antalis (Nyst, 1843)ladinum, Dentalium Dall, 1916laneensis, Dentalium (“?” Fissidentalium) Hickman,1969mayeri, Dentalium Gümbel, 1861menthifonta, Fustiaria (Episiphon) Dockery in MacNeil& Dockery, 1984microreticulata, Antalis “?” Welle, 1997minor, Cadulus gracilina var. Báldi, Hovarth & Makk,1974mississippiensis, Dentalium Conrad, 1848nicense, Dentalium Bellardi, 1852novaki, Dentalium Koenen, 1892novemcinctum, Dentalium “?” Sacco, 1897nunomae, Dentalium Takeda, 1953obliquatus, Cadulus Koenen, 1892oligocenica, Gadila Vergneau-Saubade, 1968opaculum, Dentalium Casey, 1903ottoi, Dentalium Sharp & Pilsbry in Pilsbry & Sharp,1897ovale, Dentalium Cooke, 1928parallelus, Rhabdus (Zinndorf, 1928)parianus, Cadulus Guppy in Guppy & Dall, 1896perfragile, Dentalium (Pulsellum)Koenen, 1892phenax, Cadulus Pilsbry & Sharp, 1898pictile, Dentalium (Laevidentalium) Tate, 1899polypleurum, Dentalium Seifert, 1959praecursor, Dentalium (Compressidens) Pilsbry &Sharp, 1898proterum, Dentalium (Dentalium) armillatumWoodring, 1973pseudacutum, Fissidentalium novaki R. Janssen, 1989722ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)pseudellipticum, Laevidentalium Steiner & Kabat, 2004n. nom.pseud<strong>of</strong>issura, Dentalium (Antalis) R. Janssen, 1978pyrum, Dentalium Pilsbry & Sharp, 1898radiolineata, Dentalium Clark, 1918rhenanus, Dentalium R. Janssen, 1978rupeliensis, Gadila R. Janssen, 1989sagittarii, Dentalium Pilsbry & Johnson, 1917santarosanum, Dentalium Maury, 1910sobrantensis, Dentalium radiolineata var. Clark, 1918solidissimum, Dentalium Pilsbry & Brown, 1917spiniformis, Cadulus (Gadila) Gardner, 1947strenuum, Dentalium Casey, 1903sulcatulum, Dentalium Koenen, 1892tenuicinctum, Dentalium Koenen, 1892tenuisulcatum, Dentalium (Fustiaria) Hölzl, 1962trautscholdi, Dentalium Koenen, 1868tryoni, Dentalium Pilsbry & Sharp, 1898varicostata, Dentalium Dockery in MacNeil &Dockery, 1984vicksburgensis, Cadulus Meyer, 1885vinassai, Dentalium Cossmann, 1912volvulus, Cadulus (Gadila?) Gardner, 1947waisiuense, Dentalium (Dentalium) Beets, 1943watanabei, Dentalium (Fissidentalium) Kanno, 1958waynensis, Cadulus Mansfield, 1940weinheimensis, Antalis R. Janssen, 1989yotsukurensis, Dentalium Hirayama, 1955zephyrinum, Dentalium Casey, 1903Neogene [Miocene to Pleistocene, unspecified]bifrons, Dentalium “(?)” Tate, 1887tigillum, Dentalium Yokoyama, 1931Mioceneacriculum, Dentalium (Laevidentalium) (Tate, 1887)addicotti, Cadulus (Platyschides “?”) Emerson, 1957angsananum, Dentalium (s.s.) Martin, 1921angusticostatum, Dentalium (Antalis) Baluk, 1972annectens, Cadulus (Platyschides) Woodring, 1925annulatus, Polyschides (Pilsbry, 1911)anomalocostata, Dentalium (Entalis) taurostriata “?”var. Sacco, 1897anomalum, Dentalium (Entalina) Cossmann & Peyrot,1917aratum, Dentalium Tate, 1887armillatum, Dentalium Toula, 1911arrosus, Cadulus (Platyschides) Woodring, 1925atava, Dentalium (Antale “?”) taurocostatum var. Sacco,1897attenuatum, Dentalium Say, 1824aturensis, Pseudantalis Cossmann & Peyrot, 1917australe, Laevidentalium (Sharp & Pilsbry in Pilsbry &Sharp, 1898)badense, Fissidentalium (Hörnes, 1856)bakoniense, Dentalium Palmer, 1974basteroti, Entalis Cossmann & Peyrot, 1917benoisti, Gadila Cossmann & Peyrot, 1917blountense, Cadulus Mansfield, 1935bocasensis, Dentalium (Dentalium) Olsson, 1922boettgeri, Dentalium Noetling, 1901borealis, Dentalium (Entalis) badense var. Kautsky, 1925borelliana, Gadilina Pavia, 1991bothrum, Dentalium (Dentalium) Woodring, 1973bruneti, Dentalium Steiner & Kabat, 2004 n. nom.burdigalinum, Dentalium Mayer, 1864burnsii, Cadulus floridanus var. Dall, 1892caduloide, Dentalium Dall, 1892carolinense, Dentalium Conrad, 1862caronensis, Cadulus Mansfield, 1925catulloi, Dentalium Vinassa de Regny, 1896conradi, Cadulus (Gadila) Pilsbry & Harbison, 1933costatior, Dentalium (Entalis) miopseudoentalis “?” var.Sacco, 1897costulatior, Dentalium (Antale “?”) taurocostatum var.Sacco, 1897costulatior, Dentalium michelotti var. Sacco, 1897davasense, Dentalium (Entalis) Oppenheim, 1919decemcostata, Dentalium (Entalis) taurostriata var.Sacco, 1897degrangei, Dentalium (Antale) Cossmann & Peyrot,1917degrangei, Gadila (Loxoporus) Cossmann & Peyrot,1917densmuris, Dentalium Mayer, 1858dentalina, Gadilopsis (Guppy, 1873)denticulustigris, Cadulus Maury, 1917depressum, Laevidentalium Pavia, 1991dertornensis, Dentalium (Entalis) recta var. Sacco, 1897dijki, Dentalium Martin, 1885dingdenensis, Cadulus (Gadila) Goddeeris, 1977dissimilis, Tesseracme (Guppy, 1866)divulgatum, Dentalium (Dentalium) Jung, 1969dodecacostatum, Antalis Brunet, 1995dollfusi, Dentalium Koenen, 1883dujardini, Dentalium (Antale) Peyrot, 1938dumasi, Dentalium Cossmann & Peyrot, 1917elassum, Dentalium (Episiphon) innumerabile Woodring,1973elaticosta, Dentalium (Entalis) recta var. Sacco, 1897epetrion, Cadulus (Platyschides) Woodring, 1973exdispar, Dentalium (Pseudantalis) rubescens var. Sacco,1897exlamarcki, Dentalium (Entalis) Sacco, 1897feruglioi, Laevidentalium Brunet, 1995fibula, Gadila Boettger, 1907flechensis, Antalis Brunet, 1995floratum, Fissidentalium (Zimmermann, 1847)floridanus, Cadulus Dall, 1892fodinense, Laevidentalium (s.l.) Maxwell, 1988glaucoterrarum, Dentalium Maury, 1917gracilina, Cadulus (Gadila) Sacco, 1897gracillima, Dentalium (Coccodentalium) radula var.Sacco, 1897granadanum, Dentalium Anderson, 1929granosa, Entalis Martin, 1885ZOOSYSTEMA • 2004 • 26 (4)723


Steiner G. & Kabat A. R.helveticum, Dentalium sexangulum “mut.” Cossmann& Peyrot, 1917helveticum, Siphonodentalium (Dischides “?”)Cossmann & Peyrot, 1917hemigymnum, Dentalium (Antale) degrangei var.Cossmann & Peyrot, 1917houthalenensis, Cadulus (Gadila) benoisti var.Goddeeris, 1977hurupiensis, Cadulus Dell, 1952iheringi, Dentalium Brunet, 1995ihungia, Dentalium Marwick, 1931inaequicostata, Dentalium Seguenza, 1879infans, Cadulus (Gadila) Tate, 1899infundibulum, Pulsellum Cossmann & Peyrot, 1917interbinarium, Dentalium sexangulum “mut.”Cossmann & Peyrot, 1917interrupta, Antalis (Gmelin, 1791)intertrinarium, Dentalium sexangulum var. Cossmann& Peyrot, 1917irregularis, Dentalium Hutton, 1873ixhuatlanensis, Dentalium Alencaster-Ibarra, 1950jani, Dentalium Hörnes, 1856jonkeri, Dentalium Martin, 1885junghuhni, Dentalium Martin, 1880karreri, Dentalium R. Hörnes, 1875korytnicense, Dentalium (Antalis) Baluk, 1972leonense, Dentalium (Dentalium “?”) disparileMansfield, 1930ligeriense, Dentalium Peyrot, 1938macilentum, Dentalium Pilsbry, 1911magnumcostatum, Dentalium Brunet, 1995mawsoni, Dentalium (Fissidentalium) Ludbrook, 1956minoensis, Siphonodentalium (Pulsellum) Itoigawa,1960miocaenicum, Pulsellum Boettger, 1902miocenicum, Dentalium Michelotti, 1847miopseudoentalis, Dentalium (Entalis) Sacco, 1897misatoensis, Dentalium Araki, 1958mucronatus, Cadulus Tate, 1887mutabile, Fissidentalium (Hörnes, 1856)noetlingi, Dentalium Martin, 1915octogonalis, Dentalium (Antale “?”) taurocostatum var.Sacco, 1897ornatus, Cadulus (Dischides) Baluk, 1972ortus, Fissidentalium Palmer, 1974otamaringaensis, Dentalium Marwick, 1926pardelensis, Dentalium Brunet, 1995pareorensis, Antalis (Pilsbry & Sharp, 1898)parvulina, Cadulus tumidosus var. Sacco, 1897parvum, Dentalium Mayer, 1864passerinianum, Dentalium Cocconi, 1874patagonicum, Dischides Brunet, 1995paucicostata, Dentalium (Entalis) badensis var. Sacco,1897paucistriata, Pulsellum (Entalina) tetragona var. Sacco,1897perigonicum, Dentalium Doderlein, 1864perlaevis, Dentalium (Antale) bouei var. Sacco, 1897perplexa, Gadila Boettger, 1907perrilliatae, Tesseracme Steiner & Kabat, 2004 n. nom.perseveratum, Dentalium Brunet, 1995petricola, Rhabdus (Dall, 1909)peyreirense, Dentalium (Antale) Cossmann & Peyrot,1917?philippi, Dentalium Chenu, 1843pilsbryi, Cadulus (Platyschides) Woodring, 1925planatum, Dentalium Bronn, 1831planicostata, Dentalium (Entalis) badensis var. Sacco,1897porterensis, Dentalium Weaver, 1912prismaticum, Dentalium Seguenza, 1879?proliferum, Dentalium Chenu, 1843prosperus, Striocadulus (Marwick, 1931)prudens, Dentalium Brunet, 1995pseudobouei, Dentalium (Entalis) badensis var. Sacco,1897pseudomutabile, Lentigodentalium R. Janssen, 1989pseudonyma, Dentalium Pilsbry & Sharp, 1898quadratum, Dentalium Martin, 1885radula, Coccodentalium (Gmelin, 1791)radularis, Dentalites Schlotheim, 1820raricostata, Dentalium (Antale) <strong>fossil</strong>e var. Sacco, 1897raricostata, Dentalium (Coccodentalium) radula var.Sacco, 1897rimosum, Dentalium Böse, 1906rotundatior, Dentalium inaequale var. Sacco, 1897rutteni, Dentalium Martin, 1917saccoi, Cadulus taurovulum Pavia, 1991saucatsense, Dentalium Benoist, 1873schencki, Dentalium (Rhabdus) Moore, 1963schumoi, Dentalium Pilsbry, 1911sectiforme, Dentalium (Graptacme) Tate, 1899seguenzianus, Cadulus (Gadila) Moroni & Ruggieri,1980semarangense, Dentalium Altena, 1938seminuda, Dentalium (Antale) raricostatum “race”Cossmann & Peyrot, 1917septemcostata, Dentalium (Antale “?”) taurocostatumvar. Sacco, 1897simplex, Laevidentalium (Michelotti, 1861)simplicior, Dentalium (Entalis) taurostriata var. Sacco,1897simrothi, Cadulus Pilsbry, 1911sokkohense, Dentalium Martin, 1917solidum, Fissidentalium (Hutton, 1873)spiniforme, Dentalium Martin, 1885striatellulata, Dentalium (Fustiaria) jani var. Sacco,1897stromeri, Dentalium (Fustiaria) Oppenheim, 1906subgiganteum, Dentalium d’Orbigny, 1852subjuvenis, Dentalium (Entalis) taurostriata var. Sacco,1897sublaevis, Dentalium dentalis var. Cocconi, 1874subpolitum, Siphonodentalium (Dischides) Cossmann &Peyrot, 1917subprismaticum, Dentalium (Dentalium) Baluk, 1972subserratum, Dentalium Palmer, 1974subsexangulare, Dentalium d’Orbigny, 1852724ZOOSYSTEMA • 2004 • 26 (4)


Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)superbus, Dentalium Brunet, 1995syltense, Fissidentalium floratum R. Janssen, 1989taurasperum, Fissidentalium (Sacco, 1897)taurocostata, Antalis (Sacco, 1897)taurogracilis, Dentalium (Antale) bouei var. Sacco,1897taurogracilis, Fustiaria (Sacco, 1897)taurominima, Cadulus (Loxoporus) Sacco, 1897taurostriata, Dentalium (Entalis) Sacco, 1897taurotumidosus, Cadulus (Cadulus) Sacco, 1897taurovulus, Cadulus (Cadulus) Sacco, 1897tenue, Gadila Ozaki, 1956tenuis, Cadulus (Seguenza, 1879)thallus, Cadulus (Conrad, 1834)transiens, Dentalium michelotti var. Boettger, 1907transsilvanicum, Siphonodentalium Boettger, 1897trigonaloides, Gadilina Palmer, 1974tutongense, Dentalium (Dentalium) javanum Beets,1984twistringense, Fissidentalium R. Janssen, 1989uscarianum, Dentalium (Fissidentalium) Olsson, 1922ventricosa, Gadila (Bronn, 1828)vitreum, Dentalium Gmelin, 1791waihoraense, Laevidentalium (Emerson, 1954)weinbrechti, Polyschides R. Janssen, 1989zecaninus, Gadila (Laws, 1939)Plioceneacuminatus, Gadila (Tate, 1887)acutangularis, Dentalium sexangulare var. Cocconi,1874adelaidense, Siphonodentalium (Pulsellum) Ludbrook,1956arcticus, Siphonodentalium (Dall, 1920)astensis, Dentalium (Antale “?”) dentale var. Sacco,1897astensis, Dentalium (Fustiaria) polita var. Sacco, 1897bruscasensis, Cadulus (Gadila) Weisbord, 1964calabrum, Dentalium O. G. Costa, 1851caloosaense, Dentalium Dall, 1892costae, Dentalium Pilsbry & Sharp, 1898?costaricense, Dentalium Pilsbry, 1911costatum, Dentalium J. Sowerby, 1814costulatissima, Dentalium michelotti var. Sacco, 1897crebrestriata, Dentalium sexangulum var. Coppi, 1881crux, Dentalium (Antale) Boettger, 1907cyathus, Cadulus (Crist<strong>of</strong>ori & Jan, 1832)decemcostulata, Dentalium (Antale) novemcostatum var.Sacco, 1897delli, Striocadulus (Marwick, 1965)delphinense, Dentalium Fontannes, 1880denotatum, Dentalium (Antalis) Ludbrook, 1956diploconus, Cadulus Seguenza, 1876dufresnii, Dentalium Deshayes, 1825duodecimcostata, Dentalium (Antale) novemcostatumvar. Sacco, 1897?enneagonum, Dentalium Martin, 1885entaloides, Dentalium Fleming, 1825equisetum, Dentalium (Dentalium) Olsson, 1964esmeraldum, Dentalium (Fissidentalium) Olsson, 1942exaratum, Dentalium Yokoyama, 1927gadula, Cadulus (Gadila) gadus var. Sacco, 1897gibba, Cadulus ovulum var. Seguenza, 1879gonatodes, Dentalium Martin, 1885guidottii, Dentalium Sacco, 1897?incisum, Dentalium Chenu, 1843intercosticillata, Dentalium michelotti var. Sacco, 1897laevigatum, Dentalium Eichwald, 1830latesulcatum, Dentalium (Fissidentalium) Tate, 1899laticostata, Dentalium (Entalis) badensis var. Sacco,1897leptodoma, Cadulus (Gadila) Pilsbry & Olsson, 1941linnei, Dentalium Foresti, 1895longifissum, Dentalium Harmer, 1923maculatellata, Dentalium (Antale “?”) dentale var.Sacco, 1897magnocostata, Dentalium sexangulum var. Sacco, 1897mayor, Siphondentalium (Gadila) gadus var. Foresti,1895nana, Antalis (Hutton, 1873)nipponicum, Siphonodentalium Makiyama, 1927noe, Dentalium sexangulum var. Sacco, 1897oleacinum, Dentalium agile var. Dall, 1892ozawai, Siphonodentalium (Pulsellum) Yokoyama,1926paucicostulata, Dentalium (Antale “?”) dentale var.Sacco, 1897peracuta, Dentalium sexangulum var. Sacco, 1897perstriolata, Dentalium (Antale) vulgare var. Sacco,1897playagr<strong>and</strong>ensis, Cadulus (Gadila) Weisbord, 1964pleiocenum, Dentalium Tuomey & Holmes, 1856pliocenica, Dentalium (Entalis) badensis var. Sacco,1897pliocenica, Dentalium (Entalis) recta var. Sacco, 1897pliocenica, Dentalium (Entalina) mediocarinataKoperberg, 1931politus, Dischides (S. Wood, 1842)polyedrum, Dentalium Seguenza, 1879prisma, Dentalium Dall, 1892proteiforme, Dentalium Cossmann, 1910pseudaprina, Dentalium (Antale) novemcostatum var.Sacco, 1897quadrangulatum, Dentalium Hu & Lee, 1991quatuordecimcostata, Dentalium (Antale “?”) dentalevar. Sacco, 1897quindeciesstriatum, Dentalium Eichwald, 1853quitus, Cadulus (Polyschides) Pilsbry & Olsson, 1941rotundosimplex, Dentalium michelotti var. Sacco,1897rotundulina, Dentalium michelotti var. Sacco, 1897salicensis, Cadulus Seguenza, 1879sangiorgi, Antalis (Emerson, 1954)sexangulum, Dentalium Gmelin, 1791sexdecimcostata, Dentalium (Antale “?”) dentale var.Sacco, 1897striatissimum, Dentalium Razzore, 1896ZOOSYSTEMA • 2004 • 26 (4)725


Steiner G. & Kabat A. R.striolatissima, Dentalium sexangulum var. Sacco, 1897subirregulare, Dentalium Pilsbry & Sharp, 1898subrecta, Dentalium sexangulare var. Cocconi, 1874suzukii, Dentalium (Fustiaria) Yokoyama, 1929tetragona, Entalina (Brocchi, 1814)tornatissimum, Dentalium (Episiphon) Tate, 1899totomiensis, Dentalium (Antalis) Makiyama, 1931tredecimcostatum, Dentalium (Antale) novemcostatumvar. Sacco, 1897triquetra, Gadilina (Brocchi, 1814)undecimcostata, Dentalium (Antale) novemcostatum var.Sacco, 1897venezuelana, Fustiaria (Laevidentalium “?”) Weisbord,1964viallii, Entalina Caprotti, 1962waccamawense, Dentalium (Antalis?) Gardner, 1948wangwaensis, Cadulus Nomura, 1935yatalensis, Cadulus (Dischides) Ludbrook, 1956Pleistocene <strong>and</strong> Quaternaryburicum, Dentalium (Fissidentalium)Olsson, 1942caprottii, Cadulus (Sulcogadila) Moroni & Ruggieri,1981expolita, Entalina Della Bella & Tabanelli, 1996nipponicum, Dentalium (Fustiaria) Yokoyama, 1922olivi, Polyschides (Scacchi, 1835)ovatum, Dentalium Hu, 1992tumidum, Dentalium W. D. Smith, 1913vaughani, Cadulus Dall, 1912“Tertiary” (epoch not specified)antarctotubulus, Cadulus (Gadila) Stilwell &Zinsmeister, 1992araucanum, Dentalium Philippi, 1887bouei, Dentalium Deshayes, 1825breviforme, Dentalium Mayer-Eymar, 1904brugnonei, Dentalium Steiner & Kabat, 2004 n. nom.byorituense, Dentalium (Fissidentalium) Nomura, 1935camaronesia, Dentalium sulcosum var. Ihering, 1907cristatus, Entalis Mansuy, 1914deshayesi, Dentalium Risso, 1826diazicum, Dentalium Maury, 1920fissura, Pseudantalis (Lamarck, 1818)hanguense, Dentalium Cox, 1931haytense, Laevidentalium (Gabb, 1873)hoernesi, Cadulus (Gadila) Boettger, 1902incertum, Laevidentalium (Deshayes, 1825)intermedium, Dentalium Hupé, 1854javense, Dentalium Palmer, 1974juliana, Dentalium sulcosum var. Ihering, 1907lebuense, Dentalium Philippi, 1887lentum, Dentalium Yokoyama, 1923lobion, Cadulus (Polyschides) Gardner, 1947matanzasense, Dentalium Ihering, 1907medius, Cadulus Pilsbry & Sharp, 1898monterosatoi, Dentalium rectum var. Pilsbry & Sharp,1898navidadense, Dentalium Pilsbry & Sharp, 1898nobile, Dentalium Mayer, 1863?novum, Dentalium Chenu, 1843octocostellatum, Dentalium Pilsbry & Sharp, 1898parvulum, Dentalium Philippi, 1887peranulatus, Cadulus Guppy, 1912poncensis, Cadulus Maury, 1920ponderosum, Dentalium Gabb, 1873pseudoantalis, Dentalium Lamarck, 1818pulchrum, Dentalium Wilckens, 1911?semialternans, Dentalium Chenu, 1843semiaratum, Dentalium Chapman & Crespin, 1928semiclausum, Dentalium Nyst, 1835sulcosum, Dentalium J. de C. Sowerby, 1846sunieri, Dentalium (Antalis) Koperberg, 1931teschi, Dentalium subrectum Koperberg, 1931“FOSSIL” [WITHOUT FURTHER SPECIFICATION]annulatum, Dentalium Gmelin, 1791?annulatus, Dentalites Krueger, 1823bifissuratum, Dentalium Deshayes, 1850brevifissum, Dentalium Deshayes, 1825denticulatum, Dentalium Deshayes, 1850<strong>fossil</strong>e, Dentalium Gmelin, 1791fusiforme, Dentalium Link, 1807leptoceras, Dentalium Ryckholt, 1862operculatus, Dentalites Krueger, 1823rectum, Fissidentalium (Gmelin, 1791)rex, Dentalium Pilsbry & Sharp, 1898transversatum, Dentalium Link, 1807vulpeculum, Dentalium Nardo, 1847vulpinum, Dentalium Nardo, 1847726ZOOSYSTEMA • 2004 • 26 (4)

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