Part 4 – LITERATURE CITED (complete) - Gastrotricha Global ...

Phylum Gastrotricha 

Marine; Freshwater, or both 

For the SYNOPSIS OF THE DESCRIBED GASTROTRICHA OF THE WORLD © July 2007 

Compiled by William D. Hummon, Department of Biological Sciences, Ohio University, Athens, Ohio 45701 

hummon@ohio.edu 001-740-592-5703 h 001-740-593-2993 w 001-740-593-0300 fax 

Server at: www.132.235.243.28 or www.hummon-nas.biosci.ohiou.edu 

Rev. F 24 October 2008 

♪ Note: Meeting abstracts have been omitted if the results have been subsequently published by the same author(s). 

Yellow, papers I do not have; Blue, papers not in the Todaro, Gastrotricha Portal Bibliography; Gray, Missing data. 

Papers listed in magenta are listed as ‘In Press’. 

A 

Allen, E.J. 1915. Polychaeta of Plymouth and the South Devon Coast, including a list of the Archiannelida. Journal 

of the Marine Biological Association of the UK, 4: 595. 

Amato, A.J. & M.J. Weiss. 1982. Development flexibility in the cuticular pattern of a cell-constant organism, 

Lepidodermella squamata (Gastrotricha). Transactions of the American Microscopical Society, 101: 229-240. 

Anderson, M.T. & C.E. Robbins. 1980. A new species of gastrotrich from Illinois. Transactions of the American 

Microscopical Society, 99: 225-227. 

Angelier, E. 1953. Le peuplement des sables submerges d'eau douce. Annes Biologie, 29: 467-486. 

Annandale, N. 1907. The fauna of brackish water pond at Port Canning, Lower Bengal I. Records of the Indian 

Museum, 1: 35-43. 

Ansari, Z.A., C. Rodrigues, A. Chatterji, & A.H. Parulekar. 1982. Distribution of meiobenthos and macrobenthos 

at the mouth of some rivers of the east coast of India. Indian Journal of Marine Science, 11: 341-343. 

Applewhite, P.B. & H.J. Morowitz. 1966. The micrometazoa as model systems for studying the physiology of 

memory. Yale Journal of Biology and Medicine, 39: 90-105. 

Armonies, W. & K. Reise. 2000. Faunal diversity across a sandy shore. Marine Ecology, Progress Series, 196: 

49-57. 

Austin, W.C. 1985. Gastrotricha, Volume 1, pp. 158-159. In: An Annotated Checklist of Marine Invertebrates 

in the Cold Temperate Northeast Pacific, 3 volumes. Khoyatan Marine Laboratory, Cowichan Bay, BC, 

Canada. 

Ax, P. 1970. Das Vertielungsprinzip des subterranen Psammon am Uebergang Meer-Süsswasser. Mikrofauna 

Meeresbodens 1: 1-51. 

Ax, P. 1993. Turbanella lutheri (Gastrotricha, Macrodasyoidea) im Brackwasser der Färöer. Microfauna Marina, 8: 

139-144. 

Ax, P. & P. Schmidt. 1973. Interstitielle Fauna von Galapagos. I. Einführung. Mikrofauna des Meeresbodens, 

20: 1-38. 

B 

Balavoine, G. & A. Adoutte. 1998. One or three Cambrian radiations? Science, 280: 397-398. 

Balsamo, M. 1977. Prime ricerche sui Gastrotrichi dulciacquicoli italiani. Atti della società toscana Scienze 

naturale, Memorie (ser. B), 84: 87-150. 

Balsamo, M. 1980a. Secondo contributo alla conoscenza dei Gastrotrichi delle acque dulci italiane. Atti della 

società toscana Scienze naturale, Memorie (ser. B), 87: 97-126. 

Balsamo, M. 1980b. Spectral sensitivity in a fresh-water gastrotrich (Lepidodermella squammatum Dujardin). 

Experientia, 36: 830-831. 

Balsamo, M. 1981 (for 1980). Gastrotrichi della Toscana: il Lago di Sibolla. Bolletino del Museo Civico di Storia 

Naturale Verona, 7: 547-571. 

Balsamo, M. 1982. Three new gastrotrichs from a Tuscan-Emilian Apennine lake. Bolletino di Zoologia, 49: 287- 

295. 

Balsamo, M. 1983. Gastrotrichi (Gastrotricha). Guide per il Riconoscimento delle Specie Animali delle Acque 

Interne Italiane. Consiglio Nazionale della Richerche AQ/1/199, 20: 1-92. 

Balsamo, M. 1990. Gastrotrichs from Lakes Bolsena, Chiusi and Montepulciano (central Italy), with the 

description of four new species. Bollettino di Zoologia, 57: 165-178. 

1

Balsamo, M. 1992. Hermaphroditism and parthenogenesis in lower Bilateria: Gnathostomulida and Gastrotricha, 

pp. 309-327. In: R. Dallai, ed. Sex: Origin and Evolution. Mucchi, Modena. 

Balsamo, M., R. Bertolani, P. Tongiorgi, L. Tosi, & R. Zunarelli Vandini. 1987. Ricerche faunistiche al lago 

Calamone (RE). Inf. Bot. Ital., 19: 89-92. 

Balsamo, M., M. Ferraguti, L. Guidi, M.A. Todaro & P. Tongiorgi. 2002. Reproductive system and spermatozoa 

of Paraturbanella teissieri (Gastrotricha, Macrodasyida): Implications for sperm transfer modality in 

Turbanellidae. Zoomorphology, 121: 235-241. 

Balsamo, M. & E. Fregni. 1995. Gastrotrichs from interstitial fresh water, with a description of four new species. 

Hydrobiologia, 302: 163-175. 

Balsamo, M., E. Fregni, & M. Ferraguti. 1999. Gastrotricha, pp. 171-191. In: K.G. Adiyodi & R. Adiyodi, eds. 

Reproductive Biology of Invertebrates. Vol. IX, Part A (B. G. M. Jamieson, ed.) Progress in male gamete 

ultrastructure and phylogeny. Oxford & IBH Publishing, New Delhi. 

Balsamo, M., E Fregni & P. Tongiorgi. 1994. Marine and Freshwater Gastrotricha from the Island of Montecristo 

(Tuscan Archipelago, Italy), with the description of new species. Bollettino di Zoologia, 61: 217-227. 

Balsamo, M., E. Fregni & P. Tongiorgi. 1995. Marine Gastrotricha from the coasts of Sardinia (Italy). 

Bollettino di Zoologia, 62: 273-286. 

Balsamo, M., E. Fregni & P. Tongiorgi. 1996. Marine Gastrotricha from Sicily with the description of a new 

species of Chaetonotus. Italian Journal of Zoology, 63: 173-183. 

Balsamo, M., L. Guidi, L. Pierboni, R. Marotta, M.A. Todaro & M. Ferraguti. 2007. Living without mitochondria: 

Spermatazoa and spermatogenesis in two species of Urodasys (Gastrotricha, Macrodasyida) from dysoxic 

sediments. Invertebrate Biology, 126: 1-9. 

Balsamo, M., J.-L. d’Hondt, J. Kisielewski & L. Pierboni. 2008. Global diversity of gastrotrichs (Gastrotricha) in 

fresh waters. Hydrobiologia, 595: 85-91. 

Balsamo, M., W.D. Hummon, M.A. Todaro & P. Tongiorgi. 1997. Italian marine Gastrotricha: IV. Four new 

species Chaetonotida. Italian Journal of Zoology, 64: 83-89. 

Balsamo, M. & J. Kisielewski. 1986. Chaetonotus fluviatalis, a new freshwater gastrotrich. Bollettino di Zoologia, 

53: 111-114. 

Balsamo, M. & G.C. Manicardi. 1995. Nuclear DNA content in Gastrotricha. Experientia, 51: 356-359. 

Balsamo, M. & M.A.D. Todaro. 1987. Aspidiophorus polystictos, a new marine species (Gastrotricha, 

Chaetonotida) and its life cycle. Bollettino di Zoologia, 54: 147-153. 

Balsamo, M. & M.A.D. Todaro. 1988. Life history traits of two chaetonotids (Gastrotricha) under different 

experimental conditions. Invertebrate Reproduction and Development, 14: 161-176. 

Balsamo, M. & M.A. Todaro. 1995 (for 1993). Gastrotrichi del Trentino: le Viotte del Monte Bondone. Studi 

Trentini di Scienze Naturali, Acta Biologica, 70: 9-22. 

Balsamo, M. & M.A. Todaro. 2002. Gastrotricha, pp. 45-61. In: S.D. Rundle, A.L. Robertson, & J.M. Schmid- 

Araya, eds. Freshwater Meiofauna: Biology and Ecology. Backhuys Publishers, Leiden. 

Balsamo, M., M.A. Todaro & P. Tongiorgi. 1992. Marine gastrotrichs from the Tuscan Archipelago (Tyrrhenian 

Sea): Chaetonotida, with description of three new species. Bollettino di Zoologia, 59: 487-498. 

Balsamo, M., M.A. Todaro, & P. Tongiorgi. 2004. Gastrotricha, pp. 127-135. In: C.M.Yule & H.S.Yong, eds. The 

freshwater fauna of Malaysia and Singapore. Academy of Science Malaysia, Kuala Lampur. 

Balsamo, M. & P. Tongiorgi. 1995. Gastrotricha, pp. 1-11. In: A. Minelli, S. Ruffo e S. La Posta, editors. Checklist 

delle specie della fauna italiana, Volume 7. Calderini, Bologna. 

Balsamo, M., A. Wirz, S. Cassanelli, M.A. Todaro & P. Tongiorgi. 2001. An updated phylogeny of Gastrotricha 

based on 18S rRNA gene. Proceedings of the 11th International Meiofauna Conference. Boston, USA 14-20 

July 2001. (Abstract) 

Banchetti, R. & N. Ricci. 1998. The behavior of Heterolepidoderma sp. (Gastrotricha). Zoological Science, 15: 

131- 137. 

Barnes, R.D. 1985. Current perspectives on lower invertebrates, pp. xx-xx. In: S.Conway Morris, J.D. George, 

R.Gibson & H.M.Platt, eds. The Origins and Relationships of Lower Invertebrates. Oxford University Press, 

New York. 

Baumann, F. 1910. Beiträge zur Biologie der Stockhornseen. Revue Suisse de Zoologie, 18: 647-728. 

Baumeister, W. 1936. Planktonkunde für Jedermann. Franckhsche Verlagshandlungen, Stuttgart. 62 pp. 

Beatty, J.A. & R.E. Blackwelder. 1974. Names of invertebrate phyla. Systematic Zoology, 23: 545-547. 

Beauchamp, P.M. 1911. Analyse de recents travaux sur les Rotiferes et Gastrotriches. Annales de Biologie 

Lacustre, 4: 399-412. 

2

Beauchamp, P.M. 1928. Coup d'oeil sur les recerches recents relatives aux Rotiferes. Bull. Biol. France et 

Belgique, 62: 51-125. 

Beauchamp, P.M. 1929. Le developpement des Gastrotriches. (Note prelimaire). Bulletin de la Société Zoologique 

de France, 54: 549-558. 

Beauchamp, P.M. 1932. Scientific results of the Cambridge Expedition to the East African Lakes. Journal of the 

Linnean Society of London (Zoology), 38: 241-248. 

Beauchamp, P.M. 1934. Sur La morphologie et l'ethologie des Neogossea (Gastrotriches). Bulletin de la Société 

Zoologique de France, 58: 331-342. 

Beauchamp, P. de. 1965. Classe des Gastrotriches (Gastrotricha Metschnikoff, 1864), pp 1381-1406. In: P. Grassé, 

Traité de Zoologie (Anatomie, Systématique, Biologie), Tome 4, Fasicule 3. Masson et Cie, Paris. 

Behning, A.L. 1924. Zur Erforschung der am Fussboden der Wolga leben der Oranismen. Monogr. Biol. Wolga 

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Beklemischev, V.N. 1963. On the relationship of the Turbellaria to other groups of the animal kingdom, pp. xx-xx. 

In: E.C. Dougherty, ed. The Lower Metazoa. University of California Press, Berkeley. 

Bennett, L.W. 1975. Partial trophic analysis of a freshwater Gastrotrich. Bulletin of the Association of 

Southeastern Biologists, 22: 41-42. 

Bennett, L.W. 1978. Lipid globules in fresh-water Gastrotricha. Transactions of the American Microscopical 

Society, 97: 390-392. 

Bennett, L.W. 1979. Experimental analysis of the trophic ecology of Lepidodermella squammata (Gastrotricha: 

Chaetonotida) in mixed culture. Transactions of the American Microscopical Society, 98: 254-260. 

Bennett, L.W. 1979. Distributional ecology of selected marine meiobenthic metazoa. Transactions of the American 

Microscopical Society, 98: 150. (Abstract) 

Bernhard, J.M., K.R. Buck, M.A. Farmer & S.S. Bowser. 2000. The Santa Barbara Basin is a symbiosis oasis. 

Nature, 403: 77-80. 

Bertolani, R. & M. Balsamo. 1989. Tardigradi e Gastrotrichi del Trentino: il Lago di Tovel. Studi Trentini di Scienze 

Naturali, 65: 83-93. 

Bilio, M. 1963. Die Zonierung der aquatischen Bodenfauna in den Küstensalzwiesen Schleswig-Holsteins. 

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Bilio, M. 1966. Die aquatische Bodenfauna von Salzwiesen der Nord- und Ostsee. II. Ökologische Faunenanalyse: 

Hydrozoa, Nematodes, Rotatoria, Gastrotricha, Nemertini, Polychaeta, Oligochaeta, Halacaridae, Ostracoda, 

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Birge, E.A. & C. Juday. 1911. The inland lakes of Wisconsin. Wisconsin Geological and Natural History Survey 

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Blackwelder, 

Blake, C.H. 1933. Nomenclatorial notes on Gastrotricha. Science, 77: 606. 

Blanc, H. 1929. Vie latente en milieu potamique: Exposé rétrospectif – Faits nouveau observes avec des cultures 

de vases desséchées detant de 1908 et 1910, origaires de la Gambie et du Congo. Mémoires de la Société 

Vaudoise des Sciences Naturelle, 20: 185-205. 

Blinn, D.W. & J. Green. 1986. A pump sampler study of microdistribution in Walker Lake, Arizona, U.S.A.: A 

scenescent crater lake. Freshwater Biology, 16: 175-185. 

Boaden, P.J.S. 1960. Three new gastrotrichs from the Swedish west coast. Cahiers de Biologie Marine, 1: 379- 

406. 

Boaden, P.J.S. 1962. Colonization of a graded sand by an interstitial fauna. Cahiers de Biologie Marine, 3: 245- 

248. 

Boaden, P.J.S. 1963a. Marine Gastrotricha from the interstitial fauna of some North Wales beaches. Proceedings 

of the Zoological Society of London, 140: 485-502. 

Boaden, P.J.S. 1963b. The interstitial fauna of some North Walse beaches. Journal of the Marine Biological 

Association of the UK, 43: 79-96. 

Boaden, P.J.S. 1963c. Tetranchyroderma coeliopodium n.sp., a marine interstitial gastrotrich from Ireland. Annals 

and Magazine of Natural History (series 13), 6: 375-379. 

Boaden, P.J.S. 1965a. Two new interstitial Gastrotricha of the family Thaumastodermatidae. Pubblicazioni della 

Stazione Zoologica di Napoli, 34: 219-225. 

Boaden, P.J.S. 1964. Grazing in the interstitial habitat, pp. xxx-xxx. In: D.J. Crisp, ed. Grazing in Terrestrial and 

Marine Environments. Blackwell, Oxford. 

Boaden, P.J.S. 1965b. Interstitial fauna from Porto Paone. Pubblicazioni della Stazione Zoologica di Napoli, 34: 

235-239. 

3

Boaden, P.J.S. 1966. Interstitial fauna from Northern Ireland. Veröffentlichen des Instituts für Meeresforschung in 

Bremerhaven, Sonderband 2: 125-130. 

Boaden, P.J.S. 1968. Water movement – A dominant factor in interstitial ecology. Sarsia, 34: 125-136. 

Boaden, P.J.S. 1971. Daily migration patterns in an intertidal meiobenthic community. Thalassia Jugoslavica, 7: 

1-12. 

Boaden, P.J.S. 1974. Three new thiobiotic Gastrotricha. Cahiers de Biology Marine, 15: 367-378. 

Boaden, P.J.S. 1975. Anaerobiosis, meiofauna and early metazoan evolution. Zoologica Scripta, 4: 21-24. 

Boaden, P.J.S. 1976. Soft meiofauna of sand from the Delta region of the Rhine, Meuse and Scheldt. Netherlands 

Journal of Sea Research, 10: 461-471.o 

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meiofauna). Mikrofauna Meeresboden, 61: 45-63. 

Boaden, P.J.S. 1979 

Boaden, P.J.S. 1980. Meiofaunal thiobios and “the Arenicola negation”: Case not proven. Marine Biology, 58: 

25-29. 

Boaden, P.J.S. 1985. Why is a gastrotrich? pp. 248-260. In: S. Conway-Morris, J.D. George, R. Gibson & H.M. 

Platt, eds. The Origins and Relationships of Lower Invertebrates. Oxford University Press, New York. 

Boaden, P.J.S. & E.A.G. Elhag. 1984. Meiobenthos and the oxygen budget of an intertidal sand beach. 


Boaden, P.J.S. & D.G. Erwin. 1971. Turbanella hyalina versus Protodriloides symbioticus: A study in interstitial 

ecology. Vie et Milieu, Supplément Nº 22: 479-492. 

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Jugoslavica, 7: 1-12. 

Boldyreva, N.V. 1930. Die Überwinterung von Wasserorganismen im Eise. Russ. Hidrobiol. Zhur., 9: 45-84. 

Bovee, E.C. & D.L. Cordell. 1971. Feeding on gastrotrichs by the heliozoon Actinophrys sol. Transactions of the 

American Microscopical Society, 90: 365-369. 

Brandenburg, J. 1962. Elektronenmikroskopische Untersuchung des Terminalapparates von Chaetonotus sp. 

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144. 

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Brinck, Dahl & Wieser, 1955 

Brunson, R.B. 1947. Gastrotricha of North America. II. Four new species of Ichthydium from Michigan. Papers of 

the Michigan Academy of Science, Arts, and Letters, 33: 59-62. 

Brunson, R.B. 1948. Chaetonotus tachyneusticus, a new species of gastrotrich from Michigan. Transactions of the 


Brunson, R.B. 1949. The life history and ecology of two North American gastrotrichs. Transactions of the American 


Brunson, R.B. 1950. An Introduction to the taxonomy of the Gastrotricha with a study of eighteen species from 

Michigan. Transactions of the American Microscopical Society, 69: 325-352. 

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York. 

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C 

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5

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of Macrodasyida. Meiofauna Marina, 16: 117-174. 

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species of Chaetonotida. Bollettino di Zoologia, 59: 499-516. 

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USA. Ohio Journal of Science, 80 Suppl.: 92. (Absract) 

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abundance in sandbars of acid mine polluted, reclaimed and unpolluted streams in southeastern Ohio, pp. 

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Reproductive Biology of Invertebrates. Vol. I. Oogenesis, Oviposition, and Oosorption. Wiley, London. 

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32: 115A. (Absract) 

Hummon, W.D. & M. R. Hummon. 1995. Marine Gastrotricha of the eastern Mediterranean (EMED) and Red Seas 

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Hummon, W.D., M.R. Hummon & J.W. Fleeger. 1976. Meiofauna-macrofauna interactions: I. Sand beach 

meiofauna affected by maturing Limulus eggs. Chesapeake Science, 17: 297-299. 

Hummon, W.D., M.R. Hummon & H.M. Mostafa. 1994. Marine Gastrotricha of Mediterranean Egypt. American 

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Hummon W.D. & E. Roidou. 1995. Marine Gastrotricha of Greece: A preliminary report. Biologia Gallo- 

Hellenica, 22: 279-289. 

Hummon, W.D. & M.A. Todaro. 2007. A new species of Xenotrichulidae (Gastrotricha) from southern and 

southeastern USA. Cahiers de Biologie Marine, 48: 297-302. 

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in the northwestern Adriatic Sea. Marine Pollution Bulletin, 21: 241-243. 

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Hummon, W.D., M.A. Todaro & M.R. Hummon. 1990. Marine Gastrotricha of the central Mediterranean Sea. 

American Zoologist, 31: 20A. (Absract) 

Hummon W.D., M.A. Todaro & P. Tongiorgi. 1993. Italian marine Gastrotricha: II. One new genus and ten 

new species of Macrodasyida. Bollettino di Zoologia, 60: 109-127. 

Hummon W.D., M.A. Todaro, M. Balsamo & P. Tongiorgi. 1996. Italian marine Gastrotricha: III. Four new 

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