Part 1 - AToL Decapoda

Carideorum Catalogus: 

The Recent Species of the Dendrobranchiate, Stenopodidean, 

Procarididean and Caridean Shrimps (Crustacea: Decapoda) 

S. De Grave & C.H.J.M. Fransen* 

De Grave, S. & C.H.J.M. Fransen. Carideorum catalogus: the recent species of the dendrobranchiate, 

stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). 

Zool. Med. Leiden 85(9): 30.ix.2011: 195-589 figs 1-59.— ISSN 0024-0672, ISBN 978-90-6519-200-4. 

S. De Grave, Oxford University Museum of Natural History, Parks Road, Oxford, OX1 3PW, United 

Kingdom (sammy.degrave@oum.ox.ac.uk). 

C.H.J.M. Fransen, Department of Marine Zoology, Netherlands Centre for Biodiversity Naturalis, P.O. 

Box 9517, 2300 RA Leiden, The Netherlands (charles.fransen@ncbnaturalis.nl). 

* The order of authorship is purely alphabetical, with both authors having equally contributed to this 

work. 

Key words: Crustacea; Decapoda; Dendrobranchiata; Stenopodidea; Procarididea; Caridea; checklist. 

A checklist of recent species of dendrobranchiate, stenopodidean, procarididean and caridean 

shrimps including synonyms and type localities. Also listed are unavailable names, larval names, 

nomina dubia and nomina nuda. A complete list of references to original descriptions of taxa listed is 

provided. 

Contents 

Introduction ............................................................................................................................................................ 196 

The higher classification of shrimp .................................................................................................. 196 

Fossil taxa ........................................................................................................................................................ 197 

How many shrimp species are there? ............................................................................................ 197 

Structure of the list ............................................................................................................................................. 197 

Notes on type locality ....................................................................................................................................... 199 

Notes on dating of some papers ................................................................................................................ 200 

Notes on author names .................................................................................................................................... 201 

Notes on authorship of taxa ......................................................................................................................... 202 

Notes on some papers of nomenclatorial significance ................................................................. 203 

Notes on the spelling of some names ..................................................................................................... 204 

Notes on the gender of generic names and gender agreement in species names ........ 204 

Acknowledgements ........................................................................................................................................... 205 

Checklist ................................................................................................................................................................... 206 

Suborder Dendrobranchiata ................................................................................................................ 206 

Suborder Pleocyemata ............................................................................................................................. 248 

Infraorder Procarididea .......................................................................................................................... 248 

Infraorder Stenopodidea ........................................................................................................................ 249 

Infraorder Caridea ..................................................................................................................................... 254 

Unavailable names ..................................................................................................................................... 471 

Penaeoidea Larvata .................................................................................................................................... 471 

Penaeoidea Nomina dubia .................................................................................................................... 471

196 De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 

Penaeoidea Nomina nuda ..................................................................................................................... 472 

Sergestoidea Larvata ................................................................................................................................. 472 

Sergestoidea Nomina dubia ................................................................................................................. 472 

Sergestoidea Nomina nuda .................................................................................................................. 473 

Caridea Larvata ............................................................................................................................................ 473 

Caridea Nomina dubia ............................................................................................................................ 475 

Caridea Nomina nuda ............................................................................................................................. 476 

References ................................................................................................................................................................ 478 

Systematic index to currently valid families and genera ............................................................ 585 

Introduction 

The higher classification of shrimp 

Over the last decade or so, much has been written on the classification of Decapoda, 

fuelled by a surge in molecular phylogenetic studies, as well as close scrutiny of internal 

and external morphological characteristics. As discussed by Fransen & De Grave 

(2009), such studies on shrimps are still somewhat ”thin on the ground”, at least compared 

to the more extensive work done on the Brachyura and Anomura. At a higher 

level in decapod classification it has long been recognised that three distinct lineages of 

shrimps can be distinguished: Dendrobranchiata, Stenopodidea and Caridea, a system 

which has not been seriously challenged by recent studies. 

The internal classification of Dendrobranchiata and Stenopodidea alike has been 

stable for some time, with the only major addition being the family Macromaxillocarididae 

Alvarez, Iliffe & Villalobos (2006) to the Stenopodidea in recent years. 

A different picture has emerged for Caridea very recently with Bracken et al. (2009) 

and Chan et al. (2010), both drawing attention to the non-monophyletic status of certain 

superfamilies and families. Further, we are aware of work currently in progress (some 

by the authors of this compilation) corroborating the hypothesis that the current classification 

of Caridea is unnatural, lines of study which will lead to the resurrection of 

certain family names as well as further refinement to other families. As one of our objectives 

for the current effort was to link this compilation of species level information 

with the earlier work by Chace (1992) for families and Holthuis (1993a) for genera, we 

have elected to largely follow the classification outlined by De Grave et al. (2009) which 

builds upon this earlier work. As such, it was deemed advisable to include the recently 

resurrected family Acanthephyridae Spence Bate, 1888 in the superfamily Oplophoroidea, 

rather than in this catalogue to create a new superfamily, which would perhaps 

be more congruent with the results in Chan et al. (2010). 

Although we follow herein the classification scheme of De Grave et al. (2009), two 

recent changes have been implemented. The clarification of the status of Galatheacaris 

abyssalis Vereshchaka, 1997a, as the megalopal stage of Eugonatonotus chacei Chan & Yu, 

1991a, by De Grave et al. (2010) resulted in the removal of the family Galatheacarididae 

and superfamily Galatheacaridoidea in the current listing. Bracken et al. (2010) clarified 

the status of the family Procarididae, resulting in the recognition of a fourth group of 

shrimp, Infraorder Procarididea.

De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 197 

Fossil taxa 

The current listing is restricted to extant genera, species and subspecies only. For 

readers interested in how the current classification of extant taxa compares with those 

of fossil taxa, we refer to De Grave et al. (2009) who employed a suprageneric classification 

of Decapoda across both fossil and extant taxa, as well as Schweitzer et al. (2010) 

who list all species (and their classification) of fossil species. Schweitzer et al. (2010) list 

97 fossil species of Dendrobranchiata in 31 genera, two fossil species in Stenopodidae 

(each in its own genus) and 52 species of fossil Caridea in 33 genera. Their placement of 

the genus Udora Münster, 1839, within the superfamily Procaridoidea (herein considered 

at infraordinal level) is interesting, as this group is currently only known from six 

species, restricted to anchialine habitats. If the five known species of Udora are indeed 

correctly placed, the fossil record of the groups extends back to the Middle Jurassic, an 

interesting observation as the first extant species was only discovered in 1971, drawing 

parallels with the discovery of coelacanths, one of the better known living fossils. 

How many shrimp species are there? 

Fransen & De Grave (2009) working from a preliminary version of the present 

catalogue estimated the species/subspecies richness of shrimps as follows: Dendrobranchiata 

(505), Stenopodidea (58) and Caridea (circa 3108), with those figures slightly 

adjusted in De Grave et al. (2009). Since then numerous descriptions have appeared, 

synonymies corrected and species placed in synonymy. The current tally stands as follows: 

Dendrobranchiata (68 genera, 533 species), Procarididea (2 genera, 6 species), 

Stenopodidea (12 genera, 71 species) and Caridea (389 genera, 3438 species). Although 

the figures listed in De Grave et al. (2009) for the other decapod taxa have clearly been 

adjusted by now, the Caridea remain the second most species-rich group within the 

Decapoda, with approximately half as many species as Brachyura, the dominant taxon 

within Decapoda. 

On a more inclusive level, the Caridea are dominated by Palaemonidae (981 species), 

followed by Alpheidae (663), Atyidae (469), Hippolytidae (338) and Crangonidae 

(219). Under the traditional classification employed here, Palaemonidae comprises two 

subfamilies, with Pontoniinae being considerably richer in species (602) than Palaemoninae 

(379). In contrast to those species-rich families, are those 12 families which are 

monogeneric, some only consisting of a single species, e.g. Physetocarididae. 

On a generic level, the most speciose genera are Caridina (Atyidae, 290 species), Alpheus 

(Alpheidae, 286), Macrobrachium (Palaemoninae, 243), Synalpheus (Alpheidae, 159) 

and Periclimenes (Pontoniinae, 152). As doubt has been cast on the monophyletic status 

of these mega-genera in recent years, this ranking may substantially alter when phylogenetic 

studies progress. 

Structure of the list 

As discussed above, we recognise herein four major groups of shrimp: Infraorder 

Dendrobranchiata and suborders Procarididea, Stenopodidea and Caridea. For each 

of those higher categories we list the currently recognized superfamilies and families,


following De Grave et al. (2009) with the changes noted above. As the focus of the current 

compilation is on genera and species, for the sake of brevity we have not provided 

the historical information for superfamilies and families, nor have we listed their synonyms. 

These can be found in Holthuis (1993a) for Stenopodidae and Caridea (including 

Procarididae) and Pérez Farfante & Kensley (1997) for Dendrobranchiata. For each genus, 

we provide the reference to the original description, as well as on type species (and 

the way in which they were designated) as well as their gender. In addition, we list all 

their known synonyms. For species and subspecies, we list their original generic designation, 

in original orthography for both genus and species, as well as all known synonyms. 

For each, we provide the page number on which the species description starts, as 

well as a reference to all figures and/or plates on which the species is illustrated, as well 

as their type locality (see below). Lectotype and neotype designations are included, as 

well as additional comments when required. Additional comments concern dating issues, 

as well as figures and/or further descriptions by the same author (often again as 

sp. nov.) in a later work. For instance, Parapasiphaë gilesii was illustrated in Wood-Mason 

(1892), thus validating the name, but without any accompanying textual description 

which appeared later in Wood-Mason & Alcock (1893). Such cases have been annotated 

to provide clarity for future researchers. 

All species names are given in their original orthography. This also includes usage of 

the Latin ligatures, æ and œ when present in a species name, such as in Palæmon longicornis 

Olivier, 1811, or Œdipus gramineus Dana, 1852a. Although we are aware that the 

ICZN Code does not differentiate this orthography, we felt this to be more appropriate in 

view of the long history of taxonomy. For instance we list Pelias Niloticus Roux, 1833 (now 

Caridina nilotica) and Pal.[æmon] Swainsonii White, 1847a (a synonym of Macrobrachium 

acanthurus), as written in their original descriptions. The square brackets in the latter case 

signifying an addition on our part, to make the generic designation clearer. 

As a general rule the latest taxonomic or systematic treatment for each species were 

followed, listing species as valid or as a synonym accordingly. This does not take into 

account the uncertainties regarding the true status of some taxa, which in reality may 

be valid species instead of a synonym and vice versa, but does reflect our current understanding 

of species richness. One notable exception is that the sergestid genera 

erected by Judkins & Kensley (2008) were not recognised in a more recent revision by 

Vereshchaka (2009), who instead recognised several species groups in Sergestes sensu 

lato. As these species groups largely correspond to the new genera proposed by Judkins 

& Kensley (2008), we have elected to continue to treat them as genera, necessitating 

some generic rearrangements to new species proposed in Vereshchaka (2009). 

Nomina dubia and nomina nuda have been listed as synonyms under currently 

valid names if such information was available, if not they have been listed as a separate 

category. We were aided herein by the Holthuis card catalogue, who listed many such 

names even under appropriate family headings. As Holthuis’s logic was not always 

transparent and other names surfaced or were assigned differently by other workers, 

we have not followed his family level assignations, but have retained them under three 

categories: Penaeoidea, Sergestoidea and Caridea. 

The scientific literature abounds with misspellings of species names due to printing 

and typographical errors. Listing all of these would enlarge the current volume manyfold. 

Thus a minor error, such as Palaemon pandaliformes instead of Palaemon pandali-


formis (Stimpson, 1871) in Tavares (1993) we do not list. However, if the magnitude of 

the error is such that it could possibly be interpreted as a different name we have included 

those, to avoid further confusion. For instance, Burukovsky (1974) used Artisteus 

viritli for Aristeus virilis Spence Bate, 1881, or Acanthephyra parvirostris in Coutière 

(1911a), an erroneous spelling of A. brevirostris Spence Bate, 1888. 

During the early years of ocean exploration, numerous larval forms were described, 

for instance by Spence Bate (1888) and Ortmann (1893). Although on occasion these 

have been linked to adult forms, as is the case for Embryocaris stylicauda Ortmann, 1893, 

which is a larval stage of Stenopus hispidus (Olivier, 1811), many remain unassigned to 

adult forms at present. Those which have not been linked are herein listed under the 

headings of Penaeoidea, Sergestoidea and Caridea Larvata. 

The present catalogue includes the binomial names for shrimps from Linnaeus 

(1758) onwards, up to 1st June 2011. We have not considered pre-Linnean names, even 

though sometimes these have been mentioned in later literature, for instance in the 

works by Holthuis (1991), Tavares (1993) and others. The cut-off date for inclusion of 

names was set at 1st June 2011, although we are acutely aware that a number of names 

are currently in press and some of these will have appeared in the scientific literature 

before the present work sees the light of day. 

Notes on type locality 

For all species we have listed their type locality in one of two ways. On the whole 

we have done this by reference to the original type series, but restricted to holotype 

locality if a holotype is indicated, as is the norm in recent papers. For instance, for Periclimenes 

cannaphilus Komai & J.N. Kim, 2010, we only list the locality of Kasuga 2 Seamount 

where the holotype originated from and not the locality of the 15 paratypes from 

Nikko and Daikoku Seamounts. If no holotype was indicated in the original publication, 

then we have included all syntypic localities in the listed type locality, resulting in 

upwards of 10 stations being listed for some species from the early expeditions (e.g. 

Spence Bate, 1888). If a later nomenclatorial act selected a lectotype from the syntypic 

series, we have only listed the locality details for that particular specimen. If a neotype 

was selected, then the information for the neotype only features, and not the original 

type series. For example, Cai et al. (2006) selected neotypes for the six Japanese species 

of Atyidae described by Stimpson (1860a), the original type series being lost in the great 

Chicago fire of 1871. Thus the type locality information is only given for the neotypes, 

and the reader is referred to the discussion in Cai et al. (2006) or the original publication 

by Stimpson (1860a) for locality information for the original (now lost) syntypes. For 

the majority of taxa we have elected to adhere to the original language of the description 

in our listing of type locality, as well as the original orthography and names of localities. 

For example, Stimpson (1860a) describes Alpheus pachychirus and other species 

from ”ad insulam Loo Choo”, currently known as the Ryukyu Islands in southern Japan. 

We accept the potential criticism that this may make this part of the current listing 

somewhat difficult to work with for non-taxonomists, but in our opinion it accurately 

reflects the rich tapestry of species descriptions, and correctly emphasizes the historical 

heritage and multinational aspect of alpha-level taxonomy. By necessity, we had to accept 

our own linguistic limitations and have had (by other taxonomists who are more


proficient in such matters) localities originally listed in Russian, Chinese and Vietnamese 

translated. These are simply listed in English in the present compilation. 

In many earlier publications no type localities are listed, bar a reference to a certain 

expedition station or a vague collection locality but with a specific station included. For 

instance, Rathbun (1902a) described 50 taxa from the coast of Alaska southwards to San 

Diego, based on samples which had accumulated for many years in the Smithsonian as 

a result of the Albatross expeditions. The majority of these have vague localities for the 

syntypic series in the actual publication, but do include a specific reference to one or 

more Albatross stations. By reference to various published lists, we have added such 

information between brackets in the herein listed type locality information. These may 

need to be corrected once the material and original labels are re-examined as part of 

future revisions. Several sources were consulted for this, notably Anon. (1914) for Investigator 

stations, Murray & Hjort (1912) for the Michael Sars expedition, as well as Sewell 

(1935) for species in Calman (1939) and others. For some taxa, such information could 

not be directly obtained from published station lists or the original description, but 

later authors have tracked down such information. For example, Crosnier & Forest 

(1973) provided accurate locality information for Gennadas talismani Bouvier, 1906a, and 

Metapenaeus perlarum Nobili, 1905a. In such instances, we have always included a reference 

as to where the information came from. On a few occasions, the actual type description 

nor any later paper discussed the exact type locality details. For instance 

Macpherson (1984) when describing Nematocarcinus gladius listed three stations without 

further details with the material being deposited in the South African Museum 

(now IZIKO). In those instances we made direct enquiries with the museums involved 

and provide here the information on the specimen labels and their catalogues. 

Notes on dating of some papers 

A very important aspect of nomenclature is the accurate dating of nomenclatorial 

acts, something which is not always understood by colleagues in other biological disciplines. 

For an extreme example of this issue we refer to Low & Guinot (2010) who discuss 

a case of 48 hour precedence. Dating issues have been often reported for older 

references and are most prominent in early works, such as Guérin-Méneville (1829- 

1838) and Leach (1815-1875) to name but two. In those cases where dating issues have 

been reported in the literature, discrepancies were noted in various draft versions of the 

current catalogue or have been brought to our attention by our colleagues, we have 

endeavoured to decide upon the correct date of publication. For this we have relied on 

a variety of sources, such as Rathbun (1897) for Herbst (1791-1796), Gordon (1959) for 

Bell (1844-1853), d’Erasmo (1949) for O.G. Costa & A. Costa (1838-1871), Holthuis (1947a) 

for Sowerby (1804-1806) and Holthuis (1961a) for Guérin-Méneville (1829-1838). In addition, 

we followed Harrison & Smith (2008) for dating of all of Leach’s publications. 

For dating of individual issues of certain journals with a known history of confusion, 

we followed Duncan (1937) and Dickinson (2005) for Proceedings of the Zoological Society 

in its numerous guises as well as their Transactions, Evenhuis (2003) for the Annals and 

Magazine of Natural History and others. 

Contrary to popular belief, such dating issues are not just restricted to the earlier 

taxonomic works and are almost as ubiquitous in more recent decades, although usually


much easier to solve. For example, the description of Plesionika williamsi Forest, 1964 is 

in the volume of the Bulletin du Muséum national d’Histoire naturelle (2ème série) for 1963, 

but this particular issue was printed on the 19th June 1964 (R. Cléva, pers. comm.). A 

peculiar case are the two pontoniine genera Exoclimenella Bruce, 1995, and Periclimenella 

Bruce, 1995. As already pointed out by Holthuis (1996) the descriptions of these 

two genera in Bruce (1995) is valid under the current ICZN Code and thus predates the 

descriptions (as gen. nov.) in Ďuriš & Bruce (1995). However, this assessment was 

based on a publication date of 30th November 1994, as indicated on the frontispiece. 

Per Koeltz (Koeltz Scientific Books) has confirmed that the actual publication date of 

Bruce (1995) was 16th March 1995, which curiously does not affect the seniority of 

those descriptions as the Ďuriš & Bruce (1995) article appeared in the May/June issue 

of Journal of Natural History, which according to Evenhuis (2003) was published on 31st 

May 1995. 

In those cases where we identified a discrepancy between the date usually associated 

with a publication and the real publication date determining priority, we have 

annotated the references following the conventions used by Schweitzer et al. (2010). 

Some of the early descriptions appear in multi-year works, such as Guérin-Méneville 

(1829-1838) and H. Milne Edwards (1836-1844). For these, we follow the quotation style 

in De Grave et al. (2009). For example, the original description of Xiphocaris elongata 

appears in Guérin-Méneville (1855-1856), as Hippolyte elongatus with 1855 as the publication 

date of the actual nomenclatorial act. This is thus listed as Hippolyte elongatus 

Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856] to distinguish the nomenclatorial 

act from the bibliographical reference. 

Notes on author names 

The numerous works by Charles Spence Bate (1819-1889) have usually been cited in 

modern literature under the name of ”Bate, C.S.”, although invariably Charles himself 

used the name ”Spence Bate, C.”. It is not clear when Charles started using ”Spence 

Bate” as a surname, nor the exact reasons why, nor indeed if he just adopted this name 

in publications and continued to use ”Bate” in his private life. According to the Oxford 

National Dictionary of Biography his mother was Harriet Spence (1788-1879) and we 

postulate here that he added Spence to his surname to distinguish himself from his father 

Charles Bate (1789-1872) who was also a dentist in Cornwall. We have checked all 

his carcinological papers, as well as most of his dentistry ones and find that indeed he 

invariably signed off in all his papers as ”C. Spence Bate”. This is also the surname 

by which his contemporaries refer to him in their own publications (e.g. Semper, 1868; 

Miers, 1884a; A. Milne-Edwards, 1891, etc.), although sometimes abbreviated to ”Sp. 

Bate”. In the two editorial notes by John Murray in Spence Bate, 1888 it is stated: ”…illustrate 

the text of Mr. Spence Bate’s report…” as well as: ”The Report on the Crustacea 

Macrura, by C. Spence Bate, Esq….”. Within the text of Spence Bate (1888) itself the 

author always (with one exception) refers to his own, older work as ”Spence Bate”, often 

shortened in the synonymy listings as ”Sp. B”. The exception being when he refers 

to ”Bate & Westwood (1863, A History of British Sessile-eyed Crustacea)”. We interpret that 

here as a lapsus, as the title page of that work also identifies the authors as C. Spence 

Bate and J.O. Westwood.


The present case mirrors the consistent usage of H. Milne Edwards in publications 

by that author, whose surname was Edwards, but who added Milne to distinguish himself 

from his brother William Edwards (see Forest, 1996). As we can see no reason why 

the name Charles Spence Bate should be treated different to Henri Milne Edwards, we 

here accept the arguments put forward by Dubois (2008) in that authors are signatures 

and not persons, and thus have elected to use the name C. Spence Bate. 

For the father and son team of Henri Milne Edwards and Alphonse Milne-Edwards, 

we herein follow Forest (1996) in the spelling of their surnames, and have added, as is 

customary in carcinology, their initial to all citations. Issues surrounding the use of 

Guérin and Guérin-Méneville were discussed in Ng et al. (2008) and we follow here 

their lead in attributing names after 1836 onwards to Guérin-Méneville and before 1836 

to Guérin. 

Ng et al. (2008) also provide a more in depth discussion on the spelling of author 

names and how these should be cited in the references. As we herein adhere to the same 

rules, we refer the reader to that treatise. 

In those instances where different taxa have been described by different authors 

who share the same surname, we generally have adopted the same rules as outlined in 

Ng et al. (2008). In brief, we do not add the authors initials to the person having described 

significantly more taxa, but add those to the author having described fewer. 

For instance, for taxa described by Peter K.L. Ng we do not add P.K.L. in front of the 

surname, whilst we do so for Ngan Kee Ng (thus N.K. Ng). As in Ng et al. (2008) we 

do not include initials if the authors were not contemporaries, the intervening time lag 

making this sufficiently obvious. For instance, J.Y. Johnson (1863, 1868) and D.S. Johnson 

(1961, 1962, 1967, 1973). A special case is the citation of first names for workers of 

Chinese descent or nationality, where fashion has dictated how names are used rather 

than bibliographical accuracy. For instance, the Chinese freshwater carcinologist Guo 

Zhao Liang has published under the names Guo Zhaoliang (abbreviated to Guo, Z.), 

Guo Zhao-Liang (Guo, Z.-L.) and Guo Zhao Liang (Guo, Z.L.). As it is evident that in 

all three cases this is the same worker, we have elected not to include his initials in the 

citations, others, more purists perhaps, may disagree. In the case of the team of A.H. 

”Hank” and D.M. ”Dora” Banner, who described numerous alpheid taxa, often jointly, 

they stated several times in their larger publications (e.g. Banner & Banner, 1978; Banner 

& Banner, 1981a) that they list their own papers in strict chronological order, without 

reference to seniority of authorship. We have herein interpreted this as a desire to 

reflect a joint effort with order of authorship, perhaps somewhat arbitrarily decided 

upon and a reflection of equal share in the description. As such we have elected to not 

add initials to citations in the checklist, although order of authorship in the references 

is as given in the actual papers to facilitate bibliographical searches. All this makes for 

less cumbersome citations, however in other cases, we had no option but to include 

initials. 

Notes on authorship of taxa 

As extensively discussed by Ng et al. (2008) attributing the correct author to a certain 

taxon can be confusing, particularly (but not exclusively) in the earlier literature. 

Often in the earlier literature another carcinologists’ name is cited behind the species


name in the work of a different author. Such reference is usually to an unpublished 

manuscript, a personal letter or even a museum label and many examples abound in, 

for instance, H. Milne Edwards (1834-1840). An interesting example is the name Lysmata 

nilita as published in Hope (1851) as a nomen nudum, with Risso MS featuring 

behind the name. Clearly Hope must have had access to unpublished manuscripts by 

Risso, as is evident from Monod (1931). Fittingly, when Dohrn & Holthuis (1950) described 

the second species of Lysmata from the Mediterranean, they adopted the name 

Lysmata nilita. 

As in Ng et al. (2008) we herein strictly adhere to Article 50 of the ICZN and attribute 

authorship only to those who are directly responsible for the name and for satisfying the 

criteria of availability. For example, we thus list Caridina cavalerieioides Liu & Liang in Liang 

(2004), but in contrast in the case of Stenopus devaneyi we use Goy (1984) rather than 

Goy & Randall (1984), as was used in Goy & Randall (1986). Observant readers will thus 

notice a discrepancy between the current list and Holthuis (1993a) for certain names attributed 

to Wood-Mason & Alcock in the The Annals and Magazine of Natural History. This 

series of papers was discussed in Ng (1998) and we herein follow that interpretation. 

Notes on some papers of nomenclatorial significance 

Confusion exists in the literature regarding the type description of the rare alpheid 

Betaeus jousseaumei (now Amphibetaeus jousseaumei), which Holthuis (1993a) attributed 

to Coutière (1897a) but therein dated as 1896. Coutière himself in his magnificent opus 

on Alpheidae (Coutière, 1899a) refers instead to Coutière (1896). We have not been able 

to accurately date either of those publications and herein follow Coutière (1899a) in 

considering Coutière (1896) as the type description, as well as following him in considering 

the publication year of Coutière (1897a) as 1897 and not 1896 as indicated in 

Holthuis (1993a), as indeed do Banner & Banner (1981a) and Anker & Jeng (2006). 

Guérin-Méneville authored the Crustacea section in Ramon de la Sagra’s work 

”Histoire Physique, Politique et Naturelle de l’Île de Cuba”, which also almost contemporaneously 

appeared in a Spanish language version ”Historia fisica politica y natural de la Isla 

de Cuba”. Although nothing is known about the actual publication dates for the Crustacea 

section, it is known that for the Mollusca the French text is the original version, 

which appeared in livraisons and was afterwards translated into Spanish (G. Rosenberg, 

pers. comm.). Curiously for Caridea, the French edition has several more shrimp 

names in the text, which do not feature in the Spanish version. Perhaps this can be explained 

by the fact that these additional species did not originate from Cuba, and thus 

perhaps were deleted from a Spanish translation, targeted at a more local audience. It 

is also known that the dates appearing on the frontispiece of the Mollusca part are incorrect. 

For the French edition it is the date the text was concluded, even though the 

work appeared in livraisons over several years, whilst for the Spanish version it is an 

intermediate date (G. Rosenberg, pers. comm.). In the absence of any firm evidence regarding 

the exact publication date of the crustacean parts, both text and Plates, we have 

no other course of action available than to follow the dates on the frontispiece, even 

though future research will almost certainly prove these to be wrong, this task is now 

taken up by P. Clark. Thus, for the species mentioned in both the French and Spanish 

editions, for instance Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]


we employ 1855 as their description date, corresponding to the date on the frontispiece 

of Tomo VIII (Atlas de Zoologia), with the corresponding text in Tomo VII (Crustáceos, 

Aragnides é Insectos) with 1856 on the frontispiece. For the additional taxa listed in the 

French edition, for example Alpheus Rouxii Guérin-Méneville, 1857, we use 1857 as the 

date, corresponding to what is written on the frontispiece of that volume. 

Notes on the spelling of some names 

In general, we have listed the species in their original spelling, unless a justified 

emendation needed to be made, for example to comply with Art. 31.2 in relation to 

gender agreement between generic and species names. 

When revising the genus Pandalus, Komai (1999) proposed that the species usually 

quoted as Pandalus propinquus in the majority of the taxonomic and ecological 

literature, should revert to its original spelling in the new combination of Atlantopandalus 

propinqvus, as used (Pandalus propinqvus) in the type description by G.O. Sars 

(1870). Under Art. 32.1 we herein follow Komai (1999) and continue to list the species 

as A. propinqvus. 

A similar case exists for the commercially important species, Plesionika narval 

(Fabricius, 1787), which was originally described by Fabricius (1787) under the name 

Astacus narual. Under Art. 33.3.1 we herein maintain the species as P. narval, and do 

not revert back to the original spelling. 

The species Lebbeus microceros (Krøyer, 1841) has been variable listed as L. microceros 

and L. microceras in older literature. Krøyer (1841) originally described the species under 

the name Hippolyte microceras (Krøyer, 1841, p. 578), but a note has been inserted in 

the journal after the table of content, in which the name is corrected to Hippolyte microceros. 

Under Art. 32.5.1.1 the original spelling thus has to be treated as an inadvertent 

error and corrected to L. microceros. 

Acanthephyra eximia Smith, 1884 has often been listed as A. eximea Smith, 1884. Both 

names were used by Smith (1884). Smith himself used A. eximia in his publication of 

1886 (pp. 189, 190, 192). In between no other author had used both names in an article 

and selected a correct spelling as First Reviser, thus here Art. 24.2.4 holds and Smith 

himself becomes the First Reviser. 

Notes on the gender of generic names and gender agreement in species names 

As already discussed by Ng et al. (2009) there has been confusion about the gender 

of generic names, as well as the correct ending of species names, if adjectival in nature. 

If the original genus description stated gender, we have simply followed this. Care 

needs to be taken when assuming gender though. For instance, Eumanningia Crosnier, 

2000, may sound feminine, but as clearly stated by Crosnier (2000), the name has to be 

treated as masculine. In the majority of older descriptions the gender was not stated, in 

all those cases we follow Holthuis (1983) and Pérez Farfante & Kensley (1997) for the 

gender of the genus, with one exception. The gender of the genus Petalidium Spence 

Bate, 1881, was treated as masculine by Pérez Farfante & Kensley (1997). We herein treat 

it as neuter, in recognition of the name of the type species, Petalidium foliaceum, as well 

as the other species assigned to this genus.


Confusion exists about the correct ending of some species names, as it is not always 

clear from the type description if a species name has to be treated as a noun or an adjective, 

especially in older descriptions when derivations were not commonly included. 

For some, the adjectival nature of the name is abundantly clear, and the necessary 

changes have been introduced in the present listing. For others, our own limited knowledge 

of the classical languages has prevented us from forming a definite conclusion 

on this matter. In those few cases, we have been fortunate that Lipke Holthuis, a master 

of classical names, used these names before and we have simply followed suit. For 

instance, even though the genus Halocaridinides is masculine, Holthuis (1982) used H. 

trigonophthalma, effectively treating it as a noun, instead of the masculine form trigonophthalmus. 

Acknowledgements 

Over the long gestation period of this catalogue, numerous people provided assistance 

with tracking down references, translated relevant papers, advised on nomenclatorial 

issues, spelling of author names and dating issues, provided us with copies of 

their own work and checked our listing in their ”special” families. We are immensely 

grateful to them all. We list them here in alphabetical order: A. Anker, C.W. Ashelby, 

A.J. Bruce, R.N. Burukovsky, Y. Cai, T.-Y. Chan, P. Clark, P.C. Dworschak, C. d’Udekem 

d’Acoz, J.W. Goy, L.Y.D. Goulding, R. Gulledge, K.-I. Hayashi, M.E. Hendrickx, S. 

Ivanenko, W. Klotz, T. Komai, R. Lemaitre, X. Li, M.E.Y. Low, D. Mann, I. Marin, T. 

Naruse, P.K.L. Ng, K. Pocklington, Nguyen Thanh Son, K. Reed, J. Poupin, S.H. Tan, 

A.L. Vereshchaka, M.K. Wicksten and K. Wouters, whilst offering our apologies to 

those who assisted along the way and are not listed. We are also grateful to the librarians 

of the Koninklijk Belgisch Instituut voor Natuurwetenschappen (KBIN-IRSNB), 

Netherlands Centre for Biodiversity Naturalis, University College Cork, the Hope Library 

(OUMNH) and the Radcliffe Science Library, Oxford who provided references 

and expedited the necessary inter-library loans. For the many photographs used we 

thank A. Anker, A.J. Bruce, T.-Y. Chan, N. Ciccarese, L. Harris, O. Helker, J. Hoover, T.M. 

Iliffe, T. Komai, G.A. Lovrich, C. Lukhaup, K. Neely, T. Sakihara, E. del Solar, J. Tomas, 

and C. d’Udekem d’Acoz. We are especially grateful to A. Anker, C. d’Udekem d’Acoz, 

P.K.L. Ng and P.C. Dworschak, who went through a near complete, final version of the 

catalogue and corrected numerous errors. Any remaining errors are the authors responsibility. 

Lastly, we must pay homage to L.B. Holthuis (1921-2008) who kept meticulous 

records of species names of shrimps and erroneous spellings, many never reported 

again in the scientific literature. Without access to his files, this catalogue would have 

been far less complete. Although sadly Lipke is no longer with us and able to see the 

finished product, we are grateful for his discussions on a preliminary version, during 

which he recommended the inclusion of type localities. To his memory, we dedicate this 

catalogue.


Checklist 

Suborder Dendrobranchiata Spence Bate, 1888 

Superfamily PeNAeoIDeA Rafinesque, 1815 

Family ARISTeIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Aristaeomorpha Wood-Mason in Wood-Mason & Alcock, 1891a 

= Aristæomorpha Wood-Mason in Wood-Mason & Alcock, 1891a (type species Aristeus rostridentatus 

Spence Bate, 1881, a junior subjective synonym of Penaeus foliacea Risso, 1827, by original designation, 

gender feminine) 

Aristaeomorpha foliacea (Risso, 1827) (Fig. 1) 

= Penaeus foliacea Risso, 1827: 69; Plate 2, fig. 6. [l’Europe Méridionale, environs de Nice et des Alpes 

Maritimes, grandes profondeurs] 

= Aristeus rostridentatus Spence Bate, 1881: 189. [near the Fiji Islands, 300 fms; according to Spence 

Bate, 1888: Challenger stn 173, 19°09'35"S 179°41'50"E] 

= Aristaeomorpha Giglioliana Wood-Mason, 1892: Plate 2, fig. 2. [Type locality not indicated] 

= Aristaeomorpha mediterranea Adensamer, 1898: 627; unnumbered text figure. [Pola stns 128, Nordküste 

von Afrika, 725 m; 192, Südkuste von Kreta (36°33'N 28°59'E, 1242 m); 204, Meer von Kandia 

(36°25'N 24°2'E, 808 m)] 

= Penaeus Meridionalis Hope, 1851: 19. [Nizza; nomen nudum] 

= Aristeus japonicus Yokoya, 1933: 3; Fig. 1. [Sôyô-Maru stns 383, E of Owase, Mie-ken, Japan, 353 m; 

399, near Owase, Mie-ken, 61 m] 

Aristaeomorpha woodmasoni Calman, 1925: 8. [Off Port Blair in the Andaman Sea, 271 fms] 

Aristaeopsis Wood-Mason in Wood-Mason & Alcock, 1891a 

= Aristæopsis Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus edwardsianus 

Johnson, 1868, by original designation, gender feminine) 

Aristaeopsis edwardsiana (Johnson, 1868) 

= Penaeus edwardsianus Johnson, 1868: 897. [off Madeira, Northeast Atlantic Ocean] 

= Aristeus coralinus Spence Bate, 1888: xxxii; fig. X. [Talisman expedition, exact locality not indicated] 

= Aristeus splendens Richard, 1900: 89. [Talisman expedition stn 1442, Azores; Talisman expedition stn 

1883, off Morocco; Talisman Expedition, off Cape Spartel, according to Davie, 2002: 119] 

Aristeus Duvernoy, 1840 

= Aristeus Duvernoy, 1840 (type species Penaeus antennatus Risso, 1816, by original designation, 

gender masculine) 

Aristeus alcocki Ramadan, 1938: 40; Figs 1, 2a, 3a. [John Murray Expedition, stns 34, Gulf of Aden, 1022 

m (13°05'36''N 46°24'42''E); 35, Gulf of Aden, 450-550 m (13°14'24''N 46°14'12''E to 13°13'24''N 

46°10'00''E); 176, Gulf of Aden, 650-730 m (12°04'06''N 50°38'36''E); 177, Gulf of Aden, 270-730 m 

(12°01'54''N 50°39'12''E); 193, Gulf of Aden, 1051 m (13°06'12''N 46°24'30''E to 13°03'00''N 46°21'42''E)] 

Aristeus antennatus (Risso, 1816) 

= Penaeus Antennatus Risso, 1816: 96; Plate 2, fig. 6. [Nice] 

= Sicyonia duvernoii Risso, 1844: 95. [Nice] [nomen nudum] 

= Penaeus antemarius Costes, 1890: 558. [nomen nudum] 

Aristeus antillensis A. Milne-Edwards & Bouvier, 1909: 201; Plate 1, figs 8-13. [Nevis Island, Antilles] 

Aristeus mabahissae Ramadan, 1938: 43; Figs 2b, 3b, 4a-c. [John Murray Expeditions, stns 143 (5°15'48''N 

73°22'48''E to 5°13'42''N 73°23'36''E), vicinity of the Maldives, 795 m; 145 (4°48'42"S 73°16'24"E), vicinity 

of the Maldives, 510 m] 

Aristeus pallidicauda Komai, 1993: 23; Figs 1-4. [Off Miyako, Japan, 600-700 m] 

Aristeus semidentatus Spence Bate, 1881: 189. [south of the Philippine Islands] 

? = Aristeus occidentalis Faxon, 1893: 215. [Albatross stns 3403 (off Galapagos Islands, 0°58'30"S 

89°17'00"W, 384 fms); 3410 (off Galapagos Islands, 0°19'00"N 90°34'00"W, 331 fms)]


Fig. 1. Aristaeomorpha foliacea (Risso, 1827). Photo by Tin-Yam Chan. 

Aristeus varidens Holthuis, 1952a: 71; Figs 17-18. [7°16'S 12°02'E, 53 M.W. Ambrizette, 440 m; 5°39'S 

11°25'E, 47 M.WbyS. Cabinda, 480 m; 10°45'S 13°10'E, 35 M.W. Cap Morro, 340 m; 10°45'S 13°17'E, 

40 M.W. Cap Morro, 400-500 m; 11°53'S 13°20'E, 28 M. WbyN. Egito, 500 m; 6°25'S 11°29'E, 50 M 

WSW. Moita Seca, ± 430 m] 

Aristeus virilis (Spence Bate, 1881) 

= Hemipenaeus virilis Spence Bate, 1881: 187. [near the Philippine Islands, 255 fms; according to 

Spence Bate, 1888: Challenger stn 200, near the Philippine Islands, 6°47'N 122°28'E, 250 fms] 

= Aristeus tomentosus Spence Bate, 1881: 189. [south of the Philippine Islands] 

= Aristeus viritli Burukovsky, 1974: 48. [Erroneous spelling] 

Austropenaeus Pérez Farfante & Kensley, 1997 

= Austropenaeus Pérez Farfante & Kensley, 1997 (type species Plesiopenaeus nitidus Barnard, 1947, by 

original designation, gender masculine) 

Austropenaeus nitidus (Barnard, 1947) 

= Plesiopenaeus nitidus Barnard, 1947: 383. [Off Cape Point, South Africa, 475-630 fms] 

= Aristeus crosnieri Burukovsky, 1975: 779; Figs 1-4. [SE Atlantic, 26°11'S 06°02'E, 1150 m] 

Hemipenaeus Spence Bate, 1881 

= Hemipenaeus Spence Bate, 1881 (type species Hemipenaeus spinidorsalis Spence Bate, 1881, designated 

by Faxon, 1895, gender masculine) 

Hemipenaeus carpenteri Wood-Mason & Alcock, 1891b 

= Hemipenæus Carpenteri Wood-Mason & Alcock, 1891b: 189. [Investigator stn 97 (Bay of Bengal, 

18°26'N 85°2'E), 1310 fms] 

= Hemipenaeus triton Faxon, 1893: 215. [Albatross stns 3360 (off Panama, 6°17'00"N 82°05'00"W, 1672 

fms); 3374 (off Panama, 2°35'00"N 83°53'00"W, 1823 fms); 3381 (4°56'00"N 80°52'30"W, 1772 fms)] 

Hemipenaeus spinidorsalis Spence Bate, 1881: 186. [South Atlantic, near the Island of Tristan d’Acunha, 

1900 fms; corresponding to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 35°41'S 

20°55'W, 1900 fms]


Hepomadus Spence Bate, 1881 

= Hepomadus Spence Bate, 1881 (type species Hepomadus glacialis Spence Bate, 1881, designated by 

Fowler, 1912, gender masculine) 

Hepomadus glacialis Spence Bate, 1881: 190. [mid South Atlantic, 1875 fms; corrected in Spence Bate, 1888 

to Challenger stn 237, near Yokohama, 34°37'N 140°32'E, 1875 fms] 

Hepomadus inermis Spence Bate, 1881: 190. [middle of South Pacific, 2550 fms; according to Spence Bate, 

1888 Challenger stn 289, South Pacific, 39°41'S 131°23'W, 2550 fms] 

Hepomadus tener Smith, 1884: 409; Plate 9, figs 7-8. [Albatross stn 2099, NW Atlantic, off eastern USA, 

37°12'20"N 69°30'W, 2949 fms] 

Parahepomadus Crosnier, 1978 

= Parahepomadus Crosnier, 1978 (type species Parahepomadus vaubani Crosnier, 1978, by original designation 

and monotypy, gender masculine) 

Parahepomadus vaubani Crosnier, 1978: 48; Figs 20-22. [Vauban CH 113, 22°18'S 43°59.7'E, 990-1010 m] 

Plesiopenaeus Spence Bate, 1881 

= Plesiopenaeus Spence Bate, 1881 (type species Aristeus armatus Spence Bate, 1881, designated by 

Faxon, 1895, gender masculine) 

Plesiopenaeus armatus (Spence Bate, 1881) 

= Aristeus armatus Spence Bate, 1881: 188. [among the islands of the Australasian archipelago, in the 

North Pacific, and South Atlantic, 1900-2050 fms; according to Spence Bate, 1888: Challenger stns 

133, South Atlantic Ocean, near Tristan da Cunha, 35°41'S 20°55'W, 1900 fms; 184, near Torres Strait, 

12°8'S 145°10'E, 1400 fms; 213, near the Philippines, 5°47'N 124°1'E, 2050 fms; 237, off Japan, 34°37'N 

140°32'E, 1875 fms; 246, Mid-Pacific, 36°10'N 178°00'E, 2050 fms; 276, the Low Archipelago, 13°28'S 

149°30'W, 2350 fms; 323, east of Buenos Ayres, 35°39'S 50°47'W, 1900 fms] 

Plesiopenaeus coruscans (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Aristæeus coruscans Wood-Mason in Wood-Mason & Alcock, 1891a: 280; Fig. 6. [Investigator stn 112 

(Bay of Bengal, 13°47'30"N 92°36'E), 561 fms] 

Pseudaristeus Crosnier, 1978 

= Pseudaristeus Crosnier, 1978 (type species Aristæus crassipes Wood-Mason in Wood-Mason & 

Alcock, 1891a, by original designation, gender masculine) 

Pseudaristeus crassipes (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Aristæus crassipes Wood-Mason in Wood-Mason & Alcock, 1891a: 281; Fig. 7. [Investigator stn 116, 

Andaman Sea, 11°25'05"N 92°47'06"E, 405 fms; lectotype designation by Pérez Farfante, 1987] 

Pseudaristeus gracilis (Spence Bate, 1888) 

= Hemipenaeus gracilis Spence Bate, 1888: 302; Plate 44, fig. 2. [Challenger stn 207, off Tablas Island, 

Philippines, 12°21'N 122°15'E, 700 fms] 

Pseudaristeus kathleenae Pérez Farfante, 1987: 314; Figs 1-3, 4C, 5-9. [Albatross stn 5657, Teluk Bone, 

Sulawesi (Celebes), Indonesia, 3°19'40"S 120°36'30"E, 900 m] 

Pseudaristeus protensus Pérez Farfante, 1987: 327; Figs 4D, 9, 14. [Investigator stn 370, W of Everal Gujarat, 

India (Arabian Sea), 19°51'30"N 69°07'30"E, 1569 m] 

Pseudaristeus sibogae (De Man, 1911a) 

= Hemipenaeus Sibogae De Man, 1911a: 6, 25. [Siboga stn 52, Savu Sea, Indonesia, 9°3.4'S 119°56.7'E, 

1000 m] 

Pseudaristeus speciosus (Spence Bate, 1881) 

= Hemipenaeus speciosus Spence Bate, 1881: 186. [the Atlantic off the coast of South America, 2650 

fms; according to Spence Bate, 1888: Challenger stn 325, east of Buenos Ayres, 36°44'S 46°16'W, 2650 

fms]


Family BeNTHeSICyMIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Altelatipes Crosnier & Vereshchaka, 2008 

= Altelatipes Crosnier & Vereshchaka, 2008 (type species Altelatipes falkenhaugae Crosnier & Vereshchaka, 

2008, by original designation and monotypy, gender masculine) 

Altelatipes brevirostris (Kikuchi & Nemoto, 1991) 

= Benthesicymus brevirostris Kikuchi & Nemoto, 1991: 75; figs 10-11. [23°08.4'N 150°04.8'E, 0-1300 m] 

Altelatipes carinatus (Smith, 1884) 

= Benthesicymus carinatus Smith, 1884: 396; Plate 10, figs 6-7. [Albatross stn 2094, off Long Island, NW 

Atlantic, 39°44'30"N 71°4'W, 1022 fms] 

= Benthesicymus expansus Kensley, 1977: 22; Figs 4-5. [Meiring Naude stn 107 (South Africa, off Natal, 

28°37.8'S 32°38.4'E, 1200-1000 m)] 

Altelatipes falkenhaugae Crosnier & Vereshchaka, 2008: 402; Figs 1-5, 6A-C, 7A-C. [ride médio-atlantique, 

53°04'N 37°17'W, 1478-665 m] 

Bentheogennema Burkenroad, 1936a 

= Bentheogennema Burkenroad, 1936a (type species Gennadas intermedius Spence Bate, 1888, by original 

designation, gender feminine) 

Bentheogennema borealis (Rathbun, 1902a) 

= Gennadas borealis Rathbun, 1902a: 887. [Albatross stn 3783, off Copper Island, Kamchatka, 1567 

fms] 

= Gennadas calmani Kemp, 1909: 724; Plate 74, figs 5-11; Plate 95, figs 4-5. [Challenger stns 232, S of 

Japan, 35°11'N 139°28'E, 345 fms; 236, S of Japan, 34°7'N 138°E, 565 fms; 237, near Yokohama, 

34°37'N 140°32'E, 1875 fms] 

Bentheogennema burkenroadi Krygier & Wasmer, 1975: 737; Figs 1-6. [Northeastern Pacific, 51°26'N 

138°28'W] 

Bentheogennema intermedia (Spence Bate, 1888) 

= Gennadas intermedius Spence Bate, 1888: 343; Plate 58, fig. 3. [Challenger stn 106, 1°47'N 24°26'W, 

1850 fms, off Sierra Leone; between Bermuda and Azores, surface; 137, 35°59'S, 1°34'E, surface] 

= Gennadas Alicei Bouvier, 1906a: 748. [Type locality not indicated; according to Bouvier, 1908a, the 

syntypic series came from Princesse-Alice stns 1306, a 300 milles à l’ouest de S.-Miguel, 4275 m; 1715, 

Canaries, 0-1000 m; 1749, entre Madère et les Canaries, 0-2500 m; 1768, Canaries, 0-3000; 1794, entre 

les Canaries et les Açores, 0-3000 m; 1797, entre les Canaries et les Açores, 0-3000 m; 2016, sud-ouest 

du cap Spartel, 0-1800 m; 2022, a l’est du banc Seine, 0-4000 m; 2113, Mer Sargasses, 0-1500 m; 2138, 

mer des Sargasses (région orientale), 0-2500 m; 2153, au sud des Açores, 0-2000 m; 2168, au sud de 

S.-Miguel, 0-2000 m; 2168, au sud de S.-Miguel, 0-2000 m; 2212, a l’ouest de Flores, 0-1200 m; 2244, 

au sud de S.-Miguel, 0-3000 m] 

Bentheogennema pasithea (De Man, 1907a) 

= Gennadas Pasithea De Man, 1907a: 146. [Siboga stn 230, 3°58'S 128°20'E, 0-2000 m] 

= Gennadas praecox Kemp, 1910: 176; Plate 13, figs 1-4. [Investigator stn 320, off C. Comorin, 7°23'N 

75°44'E, 1053 fms] 

= Gennadas caini Tirmizi, 1960: 368; Figs 40g, 48g, 84. [John Murray Expedition, stn 171 (9°07'06''N 

55°27'06''E to 9°08'48''N 55°31'48''E), central part of Arabian Sea, 3840-3872 m] 

Bentheogennema stephenseni Burkenroad, 1940: 37. [Dana stn 3624 I (SE of New Caledonia, 28°17.6'N 

177°01'E, 5000 m wire out)] 

Benthesicymus Spence Bate, 1881 

= Benthesicymus Spence Bate, 1881 (type species Benthesicymus crenatus Spence Bate, 1881, designated 

by Spence Bate, 1888, gender masculine) 

= Bentheocetes Smith, 1884 (type species Benthesicymus bartletti Smith, 1882; by monotypy; gender 

masculine) 

Benthesicymus altus Spence Bate, 1881: 191. [between Australia and Japan, 350-1400 fms]


Benthesicymus armatus MacGilchrist, 1905: 235. [Investigator stn 287 (Arabian Sea, 21°8'30''N 65°47'E), 

1506 fms] 

Benthesicymus bartletti Smith, 1882: 82; Plate 14, figs 1-7. [Blake stn 343, 39°45'40''N 70°55'W, 732 fms] 

= Benthesicymus pleocanthus Spence Bate, 1888: 334; Fig. 48; Plate 57, fig. 2. [Challenger stns 23, off 

Sombrero Island, 18°24'N 63°28'W, 450 fms; 205, Philippine Islands, 16°42'N 119°22'E, 1050 fms; 250, 

North Pacific Ocean, 37°49'N 166°47'W, 3050 fms] 

Benthesicymus brasiliensis Spence Bate, 1881: 191. [Atlantic and Pacific Oceans; according to Spence Bate, 

1888: Challenger stns 323, east of Buenos Ayres, 35°39'S 50°47'W, 1900 fms; 168, off New Zealand, 

40°29'S 177°43'E, 1100 fms; 173, off Matuku, Fiji Islands, 19°9'35"S 179°41'50"E, 315 fms; 181, between 

Australia and the Solomon Islands, 13°50'S 151°49'E, 2440 fms; 184, near Torres Strait, 12°8'S 

145°10'E, 1400 fms; 285, South Pacific, 32°36'S 137°43'W, 2375 fms] 

= Benthesicymus moratus Smith, 1886a: 90. [Albatross stns 2042 (39°33'00"N 68°26'45"W, 1555 fms); 

2174 (38°15'00"N 72°03'00"W, 1594 fms); 2222 (39°03'15"N 70°50'45"W, 1537 fms); 2575 (41°07'00"N 

65°26'30"W, 1710 fms)] 

Benthesicymus cereus Burkenroad, 1936a: 30; Figs 6, 11-12, 19, 24, 28, 35, 42, 47. [Pawnee stn 54, 21°15'40"N 

71°17'06"W, 900-945 fms] 

Benthesicymus crenatus Spence Bate, 1881: 190. [mid Pacific, 2600 fms; according to Spence Bate, 1888: 

Challenger stns 272, north of the Low Archipelago, 3°48'S 152°56'W, 2600 fms; 276, near the Low 

Archipelago, 13°28'S 149°30'W, 2350 fms] 

Benthesicymus investigatoris Alcock & Anderson, 1899: 282. [Investigator stns 222 (Andaman Sea, 13°27'N 

93°14'30''E), 400-200 fms (200-405 fms?); 228 (Andaman Sea, 13°7'N 94°44'15''E), 640 fms; 234 (Andaman 

Sea, 13°15'30''N 93°26'E), 498 fms; 235 (Andaman Sea, 14°13'N 93°40'E), 370-419 fms] 

Benthesicymus iridescens Spence Bate, 1881: 191. [near the island of Tristan d’Acunha, 1900 fms; according 

to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 35°41'S 20°55'W, 1900 fms] 

= Benthesicymus mollis Spence Bate, 1888: 339; Plate 58, fig. 2. [Challenger stn 133, near Tristan da 

Cunha, 35°41'S 20°55'W, 1900 fms] 

= Benthesicymus longipes Bouvier, 1906a: 747. [Princesse-Alice stn 1150, au sud-ouest des îles du Cap- 

Vert, 3890 m; according to Bouvier, 1908a] 

Fig. 2. Benthesicymus tanneri Faxon, 1893. Photo by E. del Solar.


Benthesicymus laciniatus Rathbun, 1906: 906; Fig. 59; Plate 19, fig. 3. [Albatross stn 4018, vicinity of Kauai 

Island, Hawaii, 804-724 fms] 

= Benthesicymus Hjorti Sund, 1920: 30; Fig. 48; Plate 11, fig. 4. [Michael Sars stn 35 (27°27'N 14°52'W), 

S of Canary Islands, 2603 m] 

= Gennadas pectinatus Schmitt, 1921: 25; Fig. 12; Plate 11, fig. 1. [Albatross stn 4390, off Santa Catalina 

Island, California, 1350-2182 m] 

Benthesicymus seymouri Tirmizi, 1960: 328; Figs 14-23. [John Murray Expedition, stn 118, 4°05'54"S 

41°10'12"E, 1789 m] 

Benthesicymus strabus Burkenroad, 1936a: 45; Figs 10, 23, 27, 34, 41, 46, 49. [Challenger stn 285, 32°36'S 

137°43'W] 

Benthesicymus tanneri Faxon, 1893: 215. [Albatross stns 3358 (off Panama, 6°30'00"N 81°44'00"W, 555 fms); 

3362 (off Panama, 5°56'00"N 85°10'30"W, 1175 fms); 3363 (off Panama, 5°42'00"N 85°50'00"W, 978 

fms); 3364 (off Panama, 5°30'00"N 86°08'30"W, 902 fms); 3365 (off Panama, 5°31'00"N 86°31'00"W, 

1010 fms); 3366 (off Panama, 5°30'00"N 86°45'00"W, 1067 fms); 3376 (off Panama, 3°09'00"N 

82°08'00"W, 1132 fms); 3377 (3°56'00"N 81°40'15"W, 764 fms); 3380 (off Panama, 4°03'00"N 

81°31'00"W, 899 fms); 3384 (off Panama, 7°31'30"N 79°14'00"W, 458 fms); 3393 (off Panama, 

07°15'00"N 79°36'00"W, 1020 fms); 3400 (off Galapagos Islands, 0°36'00"S 86°46'00"W, 1322 fms); 

3403 (off Galapagos Islands, 0°58'30"S 89°17'00"W, 383 fms); 3407 (off Galapagos Islands, 0°04'00"S 

90°24'30"W, 885 fms); 3410 (off Galapagos Islands, 0°19'00"N 90°24'00"W, 331 fms); 3411 (off Galapagos 

Islands, 0°54'00"N 91°09'00"W, 1189 fms); 3418 (off Mexico, 16°33'00"N 99°52'30"W, 660 fms); 

3424 (off Mexico, 21°15'00"N 106°23'00"W, 676 fms); 3425 (off Mexico, 21°19'00"N 106°24'00"W, 680 

fms); 3435 (Gulf of California, 26°48'00"N 110°45'20"W, 859 fms); 3436 (27°03'40"N 110°53'40"W, 905 

fms)] (Fig. 2) 

Benthesicymus tirmiziae Crosnier, 1978: 16; Figs 7a-b, 8a-b, 11a. [Vauban stn CH 132, Madagascar, 13°43.8'S 

47°29.0'E, 1950-2150 m] 

Benthesicymus howensis Dall, 2001 

= Benthesicymus urinator howensis Dall, 2001: 428; Figs 13A-D. [Lord Howe Rise, Tasman Sea, 28°44'S 

161°54'E, 1325 m] 

Benthesicymus urinator Burkenroad, 1936a: 29; Figs 4, 5, 8-9, 17-18, 22, 32, 33, 39-40, 45. [Challenger stn 184, 

near Torres Strait, Australia, 12°8'S 145°10'E, 1400 fms] 

Benthonectes Smith, 1885a 

= Benthonectes Smith, 1885a (type species Benthonectes filipes Smith, 1885a, by monotypy, gender 

masculine) 

= Hapalopoda Filhol, 1885a (type species Hapalopoda investigator Filhol, 1885a, by monotypy, gender 

feminine) 

Benthonectes filipes Smith, 1885a: 509. [Albatross stns 2181, 39°29'00''N 71°46'00''W, 603 fms; 2206, 

39°35'00''N 71°24'30''W, 1043 fms; 2210, 39°37'45''N 71°18'45''W, 991 fms; 2235, 39°12'00''N 

72°03'30''W, 707 fms] 

= Hapalopoda investigator Filhol, 1885a: 229; Fig. 2. [Crevette prise à 1900 mètres de profondeur, à 

bord du Talisman (33°9'N 11°58'W (of Paris) (= 9°38'W of Greenwich))] 

Gennadas Spence Bate, 1881 

= Gennadas Spence Bate, 1881 (type species Gennadas parvus Spence Bate, 1881, by monotypy, gender 

masculine) 

= Amalopenaeus Smith, 1882 (type species Amalopenaeus elegans Smith, 1882, by monotypy, gender 

masculine) 

= Pasiphodes Filhol, 1885b (type species, Pasiphodes purpureus Filhol, 1885b, by monotypy, gender 

masculine) 

Gennadas barbari Vereshchaka, 1990: 131; Fig. 1. [stn 1925, 25°39'S 85°27'W] 

Gennadas bouvieri Kemp, 1909: 726; Plate 74, figs 1-4; Plate 75, figs 6-7. [Challenger stns 206, W of Manila, 

17°54'N 117°14'E, 2100 fms; 220, N of New Guinea, 0°42'S 147°E, 1100 fms]


= Gennadas alcocki Kemp, 1910: 174 (?partim, males only); Plate 13, figs 5-6. [Investigator stns 111, Bay 

of Bengal, 12°50'N 90°52'E, 1644 fms; 103, Bay of Bengal, 15°14'N 81°9'E, 1260 fms; 108, off C. Comorin, 

7°4'N 76°34'15"E, 1043 fms; 309, near the Andaman Islands, 10°9'N 93°2'15"E, 765 fms; see 

Crosnier, 1978 for discussion of this problem] 

Gennadas brevirostris Bouvier, 1905a: 748. [provient des parages de Sainte-Lucie où elle fut trouvée par le 

Blake sur des fonds de 221 brasses] 

= Gennadas similis Stephensen, 1923: 6, 12; Fig. 1. [Atlantic, 39°35'N 9°45'W, 1150 m] 

= Gennadas chiasmifera Stephensen, 1923: 6, 13; Figs 2-3. [Atlantic, 39°35'N 9°45'W, 1150 m] 

Gennadas capensis Calman, 1925: 5; Plate 1, Figs 1-2. [SS Pickle stn 87, off Cape, South Africa, 1014 fms] 

Gennadas crassus Tirmizi, 1960: 346, 371; Figs 40h, 85. [John Murray Expedition, stn 121 (5°39'00''N 

39°38'30''E to 5°40'30''N 39°43'00''E), Zanzibar area, bottom at 925 m, net apparently not on bottom] 

Gennadas elegans (Smith, 1882) 

= Amalopenaeus elegans Smith, 1882: 87; Plate 14, figs 8-14; Plate 15,figs 1-5. [Blake stns 323, 33°19'0"N 

76°12'30"W, 457 fms; 324, 33°27'20"N 75°53'30"W, 1386 fms; 325, 33°35'20"N 76°0'0"W, 647 fms; 328, 

34°28'25"N 75°22'50"W, 1632 fms; 330, 31°41'0"N 74°35'0"W, 1047 fms; 343, 39°45'40"N 70°55'0"W, 

732 fms; 893, 39°52'20"N 70°58'0"W, 372 fms; 935, 39°45'0"N 69°44'45"W, 770 fms; 952, 39°55'0"N 

70°28'0"W, 388 fms] 

Gennadas gilchristi Calman, 1925: 6; Plate 1, figs 3-4. [SS Pickle stns 77 (755 fms), 84 (500 fms), 87 (1014 

fms), 357 (900 fms), off Cape, South Africa] 

Gennadas incertus (Balss, 1927) 

= Amalopenaeus incertus Balss, 1927: 265; figs 24-29. [Valdivia stns 232, 3°26'S 58°34'E, 0-1500 m; 235, 

4°34'S 53°42'E, 0-2000 m; 237, 4°45'S 48°58'E, 0-2000 m] 

= Amalopenaeus Gardineri Balss, 1927: 267; Fig. 31. [Valdivia stn 236, 4°38'S 51°16'E, 0-2000 m; 4 miles 

N.W. of Desroches Atoll, 750-0 fms] 

Gennadas kempi Stebbing, 1914a: 283; Plate 27. [Scotia stn 468, Cape Point, South Africa, 39°48'S 2°33'E, 

2772 fms] 

Gennadas parvus Spence Bate, 1881: 192. [off Japan, 2425 fms] 

Gennadas propinquus Rathbun, 1906: 907; Fig. 61a-b. [Between Erben Bank and Kaiwi Channel, Hawai’ian 

Islands] 

= Gennadas clavicarpus De Man, 1907a: 144 (partim). [Siboga stns 128, 4°27'N 125°25.7'E, 0-700 m; 141, 

1°0.4'S 127°25.3'E, 0-1500 m; 230, 3°58'S 128°20'E, 0-2000 m; see Burkenroad, 1936a] 

= Gennadas scutatus indicus Kemp, 1913a: 62. [Investigator stns 108, off C. Comorin, 7°4'N 76°34'15"E, 

1043 fms; 109, off C. Comorin, 7°1'N 78°21'E, 738 fms] 

Gennadas scutatus Bouvier, 1906a: 748. [l’Hirondelle stn 156, entre les Açores et Terre-Neuve, surface according 

to Bouvier, 1908a] 

= Gennadas scutatus Bouvier, 1906b: 690. [nomen nudum] 

Gennadas sordidus Kemp, 1910: 177; Plate 14, figs 1-3. [Investigator stns 193, N of the Laccadive Islands, 

15°11'N 72°28'45"E, 931 fms; 194, off the Laccadive Islands, 13°47'N 72°3'45"E, 891 fms; 198, NE of 

Ceylon, 8°55'N 81°17'30"E, 764 fms] 

Gennadas talismani Bouvier, 1906a: 749. [Talisman stn 105, 16°38'N 18°24'W, 3200 m, according to Crosnier 

& Forest, 1973] 

Gennadas tinayrei Bouvier, 1906c: 10; Figs 2-4, 14. [Mer des Sargasses, Açores, Cap Spartel (Princesse-Alice 

stn 2264, 37°30'N 22°39'W, 0-3000 m, according to Davie, 2002: 127)] 

Gennadas valens (Smith, 1884) 

= Amalopenaeus valens Smith, 1884: 402; Plate 10, fig. 2. [Albatross stn 2003, 37°16'30"N 74°20'36"W, 

640 fms] 

= Pasiphodes purpureus Filhol, 1885b: Plate 3. [Type locality not indicated; Holthuis, 1980a refers to 

A. Milne-Edwards’ original colour sketch of the species in the Paris Museum, probably from Talisman 

stn 38, off Cap Ghir, Morocco, 2210 m] 

= Gennadas bidentata Stephensen, 1923: 6, 14; Figs 4, 5. [Atlantic, 57°47'N 11°33'W, 1985 m]


Family PeNAeIDAe Rafinesque, 1815 

Alcockpenaeopsis Sakai & Shinomiya, 2011 

= Alcockpenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis hungerfordi Alcock, 1905, by 

original designation, gender feminine) 

Alcockpenaeopsis hungerfordii (Alcock, 1905) 

= Parapenaeopsis hungerfordii Alcock, 1905: 522, 530. [Hongkong] 

Arafurapenaeopsis Sakai & Shinomiya, 2011 

= Arafurapenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis arafurica Racek & Dall, 

1965, by original designation, gender feminine) 

Arafurapenaeopsis arafurica (Racek & Dall, 1965) 

= Parapenaeopsis arafurica Racek & Dall, 1965: 102; Fig. 16; Plate 8, figs 6-7; Plate 13, fig. 5. [off the Fly 

River, Papua New Guinea] 

Artemesia Spence Bate, 1888 

= Artemesia Spence Bate, 1888 (type species Artemesia longinaris Bate, 1888, by monotypy, gender 

feminine) 

Artemesia longinaris Spence Bate, 1888: 281; Plate 90. [Challenger stn 321, off Montevideo, Uruguay, 

35°02'S 55°15'W, 13 fms] 

= Artemesia brevinaris Nobili, 1901a: 1; Figs 1-2. [Mar del Plata, Argentina] 

Atypopenaeus Alcock, 1905 

= Atypopenaeus Alcock, 1905 (type species Penaeus compressipes Henderson, 1893, by original designation, 


= Miyadiella Kubo, 1949a (type species Miyadella pedunculata Kubo, 1949a, by original designation, 


Atypopenaeus bicornis Racek & Dall, 1965: 85; Fig. 14; Plate 7, figs 5-6; Plate 12, fig. 6. [Off Fly River mouth, 

Papua New Guinea] 

Atypopenaeus compressipes (Henderson, 1893) 

= Penaeus compressipes Henderson, 1893: 450. [Gulf of Martaban, India] 

= Parapenaeopsis brevirostris Kubo, 1936a: 55; Plate 61. [Inland Sea of Japan] 

Atypopenaeus dearmatus De Man, 1907a: 135. [Siboga stns 302, 10°17.9'S 123°28.7'E, 216 m; 306, 8°27'S 

122°54.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

= Miyadiella ornata Holthuis, 1955a: 76; Figs 1-2. [Off NE Java, 7°33'S 114°36'E, 200 m] 

Atypopenaeus formosus Dall, 1957: 199; Fig. 21A-H. [5 km E off Sandgate, Moreton Bay, Queensland, 

Australia, 2-3 fms] 

Atypopenaeus stenodactylus (Stimpson, 1860a) 

= Penaeus stenodactylus Stimpson, 1860a: 43. [Portu ”Hong Kong”] 

= Penaeus podophthalmus Stimpson, 1860a: 43. [Portu ”Hong Kong”] 

= Miyadella pedunculata Kubo, 1949a: 264; Figs 7N, 23G, H, 58O, 74C, I, 79E, 104, 105. [Osaka-wan, 

Japan] 

Batepenaeopsis Sakai & Shinomiya, 2011 

= Batepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus tenellus Spence Bate, 1888, by original 


Batepenaeopsis tenella (Spence Bate, 1888) 

= Penaeus tenellus Spence Bate, 1888: 270. [Challenger stn 235, Bay of Kobe, Japan, 34°39'N 135°14'E, 

8 fms] 

= Penaeus crucifer Ortmann, 1890: 451; Plate 56, fig. 5. [Maizuru, Japan] 

Batepenaeopsis venusta (De Man, 1907a) 

= Parapeneopsis venusta De Man, 1907a: 134. [Siboga expedition, station off Pulu Jedan, E coast of Aru 

Islands, shallow water]


Farfantepenaeus Burukovsky, 1997 

= Penaeus (Farfantepenaeus) Burukovsky, 1997 (type species Penaeus [as Pénée] brasiliensis Latreille, 

1817, by original designation, gender masculine) 

= Penaeus (Farfantepenaeus) Burukovsky, 1972 [no type species indicated] 

Farfantepenaeus aztecus (Ives, 1891) 

= Penaeus Brasiliensis var. Aztecus Ives, 1891: 190. [Veracruz, Mexico; lectotype designated by Pérez 

Farfante, 1969] 

Farfantepenaeus brasiliensis (Latreille, 1817) 

= Penaeus [as Pénée] brasiliensis Latreille, 1817: 156. [côtes du Brésil] 

Farfantepenaeus brevirostris (Kingsley, 1878a) 

= Penaeus brevirostris Kingsley, 1878a: 98. [Estero at Realijo, W. coast of Nicaragua] 

Farfantepenaeus californiensis (Holmes, 1900) 

= Penaeus californiensis Holmes, 1900: 218. [Santa Monica, California; neotype designation by Burkenroad, 

1938] 

Farfantepenaeus duorarum (Burkenroad, 1939) 

= Penaeus duorarum Burkenroad, 1939: 31; Figs 18-19, 23, 25-27. [Atlantis stn 2813, off Alabama, Gulf 

of Mexico, 19 fms] 

Farfantepenaeus notialis (Pérez Farfante, 1967) 

= Penaeus duorarum notialis Pérez Farfante, 1967: 94; Figs 4a-d. [Oregon stn 5664, off Las Piedras, Gulf 

of Venezuela, 11°44'N 70°22'W, 26 fms] 

= Penaeus duorarum var. cameronensis Rossignol, 1962 [unavailable name under Art. 15.2] 

Farfantepenaeus paulensis (Pérez Farfante, 1967) 

= Penaeus (Melicertus) paulensis Pérez Farfante, 1967: 84; Figs 1a-d. [Santos, Sāo Paulo, Brazil] 

Farfantepenaeus subtilis (Pérez Farfante, 1967) 

= Penaeus aztecus subtilis Pérez Farfante, 1967: 89; Figs 2a-b, 3a-c. [Oregon stn 5685, off Gallinas Point, 

Departemento de la Guajira, Colombia, 12°29'N 71°54'W, 95 fms] 

Fenneropenaeus Pérez Farfante, 1969 

= Fenneropenaeus Pérez Farfante, 1969 (type species Penaeus indicus H. Milne Edwards, 1837 [in H. 

Milne Edwards, 1834-1840], by original designation, gender masculine) 

Fenneropenaeus chinensis (Osbeck, 1765) 

= Cancer chinensis Osbeck, 1765: 151. [Bocca Tiger, auf Chinesisch Pho-hao, der Tigermund, oder Phomunn, 

die Tigersöfnung. Fischer kamen mit verschiedenen Fisharten zu uns, .... und chinesischen 

Krabben: Cancer chinensis] 

= Penaeus orientalis Kishinouye, 1917: 79. [Chosen, Kantoshu and Chintao] 

Fenneropenaeus indicus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

? = Palæmon longicornis Olivier, 1811: 662. [Type locality not indicated] 

= Penaeus indicus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 415. [les côtes de Coromandel] 

= Penaeus indicus longirostris De Man, 1892a: 511; Plate 29, fig. 53. [Celebes, Makassar] 

Fenneropenaeus konkani Chanda & Bhattacharya, 2003: 229; Fig. 1. [Mirkarwada Fish landing centre, Ratangiri, 

Maharstra, West Coast of India] 

Fenneropenaeus merguiensis (De Man, 1888a) 

= Penaeus merguiensis De Man, 1888a: 287; Plate 18, fig. 8; Plate 19, fig. 1. [Burma, Mergui Archipelago] 

Fenneropenaeus penicillatus (Alcock, 1905) 

= Penaeus indicus penicillatus Alcock, 1905: 515, 525. [Off the Orissa coast; at Bombay; Karáchi; 

Gangetic Delta; Mergui] 

Fenneropenaeus silasi (Muthu & Motoh, 1979a) 

=Penaeus silasi Muthu & Motoh, 1979a: 64; Figs 1-2. [Tawau, Sabah, 36 m] 

Funchalia Johnson, 1868 

= Funchalia Johnson, 1868 (type species Funchalia woodwardi Johnson, 1868, by monotypy, gender 

feminine)


= Grimaldiella Bouvier, 1905b (type species Grimaldiella richardi Bouvier, 1905b, by monotypy, gender 

feminine) 

= Hemipenaeopsis Bouvier, 1905b (type species Hemipenaeopsis villosus Bouvier, 1905b, by monotypy, 


Funchalia danae Burkenroad, 1940: 36. [Dana stn 4017 VII (Canary Islands, 29°13'N 14°12'W, 1000 m wire 

out)] 

Funchalia sagamiensis Fujino, 1975: 200; Figs 1-3. [Amadai-ba, Sagami Bay, central Japan, upper water 

layer] 

Funchalia taaningi Burkenroad, 1940: 36. [Dana stn 3920 III (N of Seychelles, 1°06'N 62°25'E, 3000 m wire 

out)] 

Funchalia villosa (Bouvier, 1905b) 

= Hemipenaeopsis villosus Bouvier, 1905b: 981 (pro parte). [capturée par la Princesse-Alice entre les îles 

Canaries et les Açores] 

= Funchalia vanhöffeni Lenz & Strunck, 1914: 306; Plate 17, figs 20-21. [nördlich von Tristan da Cunha] 

Funchalia woodwardi Johnson, 1868: 895. [Off Madeira, NE Atlantic Ocean] 

= Peneus Genianus Risso, 1841 [nomen nudum] 

= Sycionia geny Risso, 1844: 95. [Nice] [nomen nudum] 

= Sicyonia Genyana Hope, 1851: 19. [nomen nudum] 

= Grimaldiella Richardi Bouvier, 1905b: 981, 982. [capturée par la Princesse-Alice dans se dernière 

campagne, au large de l’île Saint-Michel, en plein groupe des Açores] 

= Peneus genyanus Monod, 1931: 107. [nomen nudum] 

Ganjampenaeopsis Sakai & Shinomiya, 2011 

= Ganjampenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis uncta Alcock, 1905, by 


Ganjampenaeopsis uncta (Alcock, 1905) 

= Parapenaeopsis uncta Alcock, 1905: 522, 528. [Ganjam coast, 10-11 fms] 

= Parapenaeopsis probata Hall, 1961: 96; Plate 19, figs 11-13. [South of Bedok, 24 fms; South of Singapore, 

24 and 45 fms] 

Heteropenaeus De Man, 1896 

= Heteropenaeus De Man, 1896 (type species Heteropenaeus longimanus De Man, 1896, by monotypy, 


Heteropenaeus longimanus De Man, 1896: 111. [Java-See] 

Holthuispenaeopsis Sakai & Shinomiya, 2011 

= Holthuispenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis atlantica Balss, 1914a, by 


Holthuispenaeopsis atlantica (Balss, 1914a) 

= Parapenaeopsis atlantica Balss, 1914a: 593. [Victoria, Kamerun] 

= Trachypenaeus constrictus var. africana Balss, 1916: 17; Figs 3-4. [Goldküste: Kap Coast Castle, 13 m; 

Addah, 11 m; Acra, 9 m. Togo, Lome, 13 m. Dahomey, Whyda, Cabinda, Landana, 7 m. Französisch- 

Kongo: Nyanga-Fluß, 13 m; Loango, 10 m. Angola: Ambriz, 13 m; Kinsembo, 13 m] 

Kishinouyepenaeopsis Sakai & Shinomiya, 2011 

= Kishinouyepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus cornutus Kishinouye, 1900, by 


Kishinouyepenaeopsis amicus (V.C. Nguyên, 1971) 

= Parapenaeopsis amicus V.C. Nguyên, 1971: 46; Fig. 1. [West Tonkin Gulf] 

= Parapenaeopsis sinica Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South 

China Sea]


Kishinouyepenaeopsis cornuta (Kishinouye, 1900) 

= Penaeus cornutus Kishinouye, 1900: 23; unnumbered text fig. [Bay of Ariake, Kushu] 

Kishinouyepenaeopsis incisa (Liu & Wang, 1986) 

= Parapenaeopsis incisa Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South 

China Sea] 

Kishinouyepenaeopsis maxillipedo (Alcock, 1905) 

= Parapenaeopsis maxillipedo Alcock, 1905: 522, 527. [Bombay, Madras, Arakan coast, India] 

Litopenaeus Pérez Farfante, 1969 

= Litopenaeus Pérez Farfante, 1969 (type species Penaeus vannamei Boone, 1931, by original designation, 


Litopenaeus occidentalis (Streets, 1871a) 

= Penaeus occidentalis Streets, 1871a: 243. [Isthmus of Panama] 

Litopenaeus schmitti (Burkenroad, 1936b) 

= Penaeus schmitti Burkenroad, 1936b: 315; Figs 1-3. [Kingston Bay, Jamaica] 

Litopenaeus setiferus (Linnaeus, 1767) 

= Cancer setiferus Linnaeus, 1767: 1054. [Indiis] 

= Pæneus orbignyanus Latreille, 1817: 155. [côtes maritimes du département de la Vendée] 

= P.[enaeus] fluviatilis Say, 1818: 236. [North America] 

= Penaeus gracilirostris Thallwitz, 1892: 3; Plate 1, fig. 5. [Nord-Celébes] 

Litopenaeus stylirostris (Stimpson, 1871) 

= Penaeus stylirostris Stimpson, 1871: 134. [Panama] 

Litopenaeus vannamei (Boone, 1931) 

= Penaeus vannamei Boone, 1931: 173; Fig. 16. [Golfo de Panama] 

Macropetasma Stebbing, 1914b 

= Macropetasma Stebbing, 1914b (type species Parapenaeus africanus Balss, 1913a, by monotypy, gender 

neuter) 

Macropetasma africana (Balss, 1913a) 

= Parapenaeus africanus Balss, 1913a: 105. [Swakopmund, Namibia] 

Marsupenaeus Tirmizi, 1971a 

= Marsupenaeus Tirmizi, 1971a (type species Penaeus canaliculatus var. japonicus Spence Bate, 1888, by 

monotypy, gender masculine) 

Marsupenaeus japonicus (Spence Bate, 1888) 

= Penaeus canaliculatus japonicus Spence Bate, 1888: 245; Plates 31; 32, fig. 4; 37, fig. 2. [Japan] 

= Penaeus pulchricaudatus Stebbing, 1914b: 14; Plate 67. [Great Fish Point Lighthouse, Cape province, 

South Africa] 

Megokris Pérez Farfante & Kensley, 1997 

= Megokris Pérez Farfante & Kensley, 1997 (type species Penaeus granulosus Haswell, 1879, by original 

designation, gender masculine) 

Megokris akademik Shinomiya & Sakai, 2006a: 1220; Figs 1-2. [R.V. ”Akademik” stn PG-13-Ku, Persian 

Gulf, 29°13.587'N 49°53.955'E, 41 m] 

Megokris ghamrawyi Shinomiya & Sakai, 2006a: 1223; Figs 3-6. [R.V. ”Akademik” stn PG-13-Ku, Persian 

Gulf, 29°13.587'N 49°53.955'E, 41 m] 

Megokris gonospinifer (Racek & Dall, 1965) 

= Trachypenaeus gonospinifer Racek & Dall, 1965: 89; Fig. 15; Plate 7, figs 7-8; Plate 12, fig. 7. [S. of Port 

Romilly, Papua New Guinea, 7°55'S 144°48'E, 7 fms] 

Megokris granulosus (Haswell, 1879) 

= Penaeus granulosus Haswell, 1879: 41. [Darney Island; lectotype designation by Griffin, 1970] 

= Trachypeneus salaco De Man, 1907a: 135. [Siboga stns 205, Lohio bay, Buton Strait, Indonesia, 22 m; 

258, Tual Anchorage, Kai Islands, 22 m] 

= Trachypeneus furcilla Hall, 1961: 102; Plate 20, figs 16-17. [Outer Shoal, Malaysia, 6 fms]


Megokris halli Shinomiya & Sakai, 2006b: 1255; Figs 1-2. [Singapore Fisheries Research stn B 146, Permanent 

Buoy 10, Angler Buoy, 9 m] 

Megokris manihine Shinomiya & Sakai, 2006b: 1259; Figs 3-4. [Singapore Fisheries Research stn c5/15, 

5°12'N 114°59'E, 37 m] 

Megokris motohburiorum Shinomiya & Sakai, 2006b: 1261; Fig. 5. [Iloilo, Philippines] 

Megokris pescadoreensis (Schmitt, 1931a) 

= Trachypeneus pescadoreensis Schmitt, 1931a: 265; Plate 32, figs 2-4. [Pescadores Islands, Formosa] 

Megokris sedili (Hall, 1961) 

= Trachypeneus sedili Hall, 1961: 100; Plate 20, figs 18-19. [Malaysia, S of Bedok, 24 fms] 

Melicertus Rafinesque, 1814 

= Melicertus Rafinesque, 1814 (type species Melicertus tigrinus Rafinesque, 1814, by monotypy, gender 

masculine) 

Melicertus canaliculatus (Olivier, 1811) 

= Palæmon canaliculatus Olivier, 1811: 660. [Indonesia, N Sumatra, Pulu Weh; neotype designated 

by Pérez Farfante, 1976] 

Melicertus hathor (Burkenroad, 1959) 

= Penaeus latisulcatus hathor Burkenroad, 1959: 80; Figs 7, 8. [Mersa Thlemel, Gulf of Suez] 

Melicertus kerathurus (Forskål, 1775) 

= Cancer kerathurus Forskål, 1775: 95. [Smirnæ & Alexandriæ] 

= Palæmon sulcatus Olivier, 1811: 661. [Elle se trouve aux environs de Smyrne & d’Alexandrie] 

= Alpheus Trisulcatus Leach, 1814 [in Leach, 1813-1814]: 431. [Anglesea] 

= Melicertus tigrinus Rafinesque, 1814: 22. [Sicily] 

= Alpheus caramote Risso, 1816: 90. [Nice] 

= Peneus mars Risso, 1816: 47; Plate 2, fig. 5. [Nice, se tient à une grande profondeur] 

= Alpheus punctulatus Risso, 1822: 247. [mer de Nice, régions sablonneuses] 

= Peneus cristatus Risso, 1827: 67. [environs de Nice et des Alpes Maritimes; grandes profondeurs] 

= Penaeus fasciatus Hope, 1851: 19. [nomen nudum] 

Melicertus latisulcatus (Kishinouye, 1896) 

= Penaeus latisulcatus Kishinouye, 1896: 372. [Kagoshima, Tokyo Bay, Japan] 

Melicertus longistylus (Kubo, 1943) 

= Penaeus longistylus Kubo, 1943: 200. [off South-East coast of Hainan Island] 

= Penaeus jejunus Hall, 1956: 75; Plate 10, figs 9-10. [probably from the neighbourhood of Singapore] 

= Penaeus caesius Dall, 1957: 143; Fig. 2A-G. [12 miles NNE of Bowen, Queensland, Australia, 19-25 fms] 

Melicertus marginatus (Randall, 1840) (Fig. 3) 

= P.[enœus] marginatus Randall, 1840: 146. [Sandwich Islands] 

= Penaeus teraoi Kubo, 1949a: 288; Figs 7F, 20F, 24N-P, 49K, 58G, 73D, J, 77S, 112. [Miya, Aiti Prefecture, 

Japan] 

Melicertus plebejus (Hess, 1865) 

= Penaeus plebejus Hess, 1865: 168; Plate 7, fig. 19. [Sydney] 

= Penaeus canaliculatus var. australiensis Spence Bate, 1888: 248; Plate 32, fig. 3. [Port Jackson, Australia, 

2-10 fms] 

= Penaeus maccullochi Schmitt, 1926a: 359, 370; Plate 66, figs 1-3. [21 km NE of North Reef, Queensland, 

Australia] 

Melicertus similis Chanda & Bhattacharya, 2002: 495; Figs 1-6. [Port Blair, Andaman Is., Bay of Bengal] 

Metapenaeopsis Bouvier, 1905b 

= Metapenaeopsis Bouvier, 1905b (type species Metapenaeopsis pubescens Bouvier, 1905b, by monotypy, 


= Archipenaeopsis Bouvier, 1905b (type species Archipenaeopsis vesitus Bouvier, 1905b, by monotypy, 


= Leptopenaeus Kishinouye, 1929 (type species Penaeus philippii Spence Bate, 1888, designated by 

Pérez Farfante & Kensley, 1997, gender masculine)


Fig. 3. Melicertus marginatus (Randall, 1840). Photo by Tin-Yam Chan. 

= Ceratopenaeus Kishinouye, 1929 (type species Parapenaeus mogiensis Rathbun, 1902b, designated by 

Pérez Farfante & Kensley, 1997, gender masculine) 

= Erythropenaeus Kishinouye, 1929 (type species Parapenaeus akayebi Rathbun, 1902b, designated by 

Pérez Farfante & Kensley, 1997, gender masculine) 

Metapenaeopsis acclivis (Rathbun, 1902b) 

= Parapenaeus acclivis Rathbun, 1902b: 41; Figs 12-14. [Mogi, Japan] 

Metapenaeopsis aegyptia Galil & Golani, 1990: 229; Figs 1a-b, 2a-c, 3a. [off Palmahim, Israel, 31°56'N 

34°35'E, 50 m] 

= Metapenaeopsis judaensis Por, 1989: 211. [nomen nudum] 

Metapenaeopsis andamanensis (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Metapenaeus philippinensis var. andamanensis Wood-Mason in Wood-Mason & Alcock, 1891a: 271. 

[N of Port Blair, Andaman Sea, 112-244 fms] 

Metapenaeopsis angusta Crosnier, 1987a: 441; Figs 16b, 18b, 19c-d. [MUSORSTOM I, stn 5, 14°01.5'N 

120°23.5'E, 215-200 m] 

Metapenaeopsis assimilis (De Man, 1920a) 

= Penaeopsis assimilis De Man, 1920a: 105. [Off Pulu Weh, N Sumatra, Indonesia; lectotype designation 

by Crosnier, 1991] 

= Metapenaeopsis raceki Starobogatov, 1972: 405, 409; Plate 10, fig. 127a-b. [nomen novum for Metapenaeopsis 

distincta sensu Racek & Dall, 1965 nec De Man, 1907a] 

Metapenaeopsis barbata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penaeus barbatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 3. [Japan] 

= Parapenaeus akayebi Rathbun, 1902b: 39. [Mogi, Japan] 

Metapenaeopsis batei (Miers, 1884b) 

= Penæus batei Miers, 1884b: 296; Plate 32, fig. D. [Albany Island, 3-4 fms] 

Metapenaeopsis beebei (Burkenroad, 1938) 

= Penaeopsis (Metapenaeopsis) beebei Burkenroad, 1938: 74; Figs 18-19. [Templeton Crocker Expedition 

stn 136, dredge 30, Arena Bank, Gulf Of California, 23°27'N 109°24'W, 35 fms]


Metapenaeopsis ceylonica Starobogatov, 1972: 406, 408 (key); Plate 11, fig. 159a-b; nomen novum for M. 

hilarulus sensu De Bruin, 1965 nec De Man, 1911a. [Ceylon, W coast, W of Mannar, 6 fms] 

Metapenaeopsis commensalis Borradaile, 1899: 1001, Plate 63, fig. 1-1b. [Ellice Islands, Rotuma, Fiji] 

= Penaeopsis Borradailei De Man, 1911a: 73. [Siboga stn 131, anchorage off Beo, Karakelong Island, 

Talaud Archipelago, Indonesia; lectotype designation by Crosnier, 1991] 

Metapenaeopsis coniger (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Metapenaeus coniger Wood-Mason in Wood-Mason & Alcock, 1891a: 272. [Investigator stns 119 (Bay 

of Bengal, 12°20'N 85°8'E), 95 fms; off Mahánaddi Delta, 68 fms; 96 (Bay of Bengal, 18°30'N 84°46'E), 

98-102 fms] 

Metapenaeopsis costata Crosnier, 1991: 275; Figs 83, 84. [MUSORSTOM 3, stn CP 142, Philippines, 11°47'N 

123°01.5'E, 26 m] 

Metapenaeopsis crassissima Racek & Dall, 1965: 26; Fig. 2C; Plate 2, fig. 1; Plate 4, figs 5-6; Plate 9, fig. 3. 

[Shark Bay, Western Australia] 

Metapenaeopsis dalei (Rathbun, 1902b) 

= Parapenaeus dalei Rathbun, 1902b: 40; Figs 9-11. [Mogi, Japan] 

? = Metapenaeopsis incomptus Kubo, 1949a: 424; Figs 8D, 46A, 76L, R, 80N, 148F. [East China Sea] 

Metapenaeopsis difficilis Crosnier, 1991: 255; Figs 65-67. [MUSORSTOM 3, stn CP 134, Philippines, 

12°01.1'N 121°57.3'E, 92-95 m] 

Metapenaeopsis distincta (De Man, 1907a) 

= Metapeneus distinctus De Man, 1907a: 132. [Siboga stn 184, Anchorage off kampong Kelang, south 

coast of Manipa-island, 36 m; female specimen illustrated by De Man, 1913; designated as lectotype 

by Crosnier, 1991] 

Metapenaeopsis dura Kubo, 1949a 

= Metapenaeopsis durus Kubo, 1949a: 421; Figs 8A, 18G-L, 22O, 46D, 64C-C', 76E, J, 80L, 148C, 149. 

[Tokyo fish market, Miya, Aichi Prefecture, Japan] 

Metapenaeopsis erythraea Crosnier, 1987a: 443; Figs 16c, 18c, 19e-f. [Valdivia stn 122, Mer Rouge, 21°22'N 

39°04'E, 383-363 m] 

Metapenaeopsis evermanni (Rathbun, 1906) 

= Metapenaeus evermanni Rathbun, 1906: 904; Fig. 58. [Albatross stn 3849, south coast of Molokai Island, 

Hawaii Islands, 79-134 m] 

Metapenaeopsis faouzii (Ramadan, 1938) 

= Penaeopsis (Metapenaeus) faouzii Ramadan, 1938: 72; Fig. 14d. [John Murray Expedition stn 161, 

Maldives, 5°04'48"N 72°50'30"E, 46 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis fusca R.J.G. Manning, 1988: 91; Fig. 1A-D. [Singleton, south Western Australia (32°27'S 

115°44'E)] 

Metapenaeopsis gaillardi Crosnier, 1991: 200; Figs 25-27. [Lagon stn 314, Iles des Pins, Nouvelle-Calédonie, 

27 m] 

Metapenaeopsis gallensis (Pearson, 1905) 

= Parapenaeus gallensis Pearson, 1905: 72; Plate 1, figs 3, 3a-b. [S of Galle, Ceylon] 

Metapenaeopsis gerardoi Pérez Farfante, 1971a: 20; Figs 11-12, 13C. [Oregon stn 5440, off Mayagüez, Puerto 

Rico, 18°08.5'N 67°23'W, 22 m] 

Metapenaeopsis goodei (Smith, 1885b) 

= Parapenaeus Goodei Smith, 1885b: 176. [Bermuda; Bay of Panama] 

= Archipenæopsis vestitus Bouvier, 1905a: 747. [capturé par le Blake dans la mer des Antilles, à la faible 

profondeur de 37 brasses (Blake stn 11, NW of Dry Tortugas Islands, 24°43'N 83°25'W, 37 fms, see A. 

Milne-Edwards & Bouvier, 1909 and Pérez Farfante, 1971a)] 

= Parapenaeopsis Rathbuni Bouvier, 1905a: 748. [dans la mer des Antilles, à 17 brasses de profondeur 

(26°16'25''N 82°50'10''W, west of Florida; see Pérez Farfante, 1971a)] 

Metapenaeopsis hilarula (De Man, 1911a) 

= Penaeopsis sp. (provisional name hilarulus) De Man, 1911a: 70 (partim). [Siboga stn 240, Banda-anchorage, 

Indonesia, 9-36 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis hobbsi Pérez Farfante, 1971a: 24; Figs 13D, 14-17. [Pillsbury stn 737, NW of Cabo Cordera, 

Venezuela, 10°44'N 66°07'W to 10°45'N 66°08'W, 60-73 m]


Metapenaeopsis incisa Crosnier, 1991: 277; Figs 85-87. [BENTHEDI stn 101D, Iles Glorieuses, 11°25.7'N 

47°19.5'E, 26 m] 

Metapenaeopsis ivanovi Crosnier, 1994a: 340; Figs 1-3. [au large du Kenya] 

Metapenaeopsis kishinouyei (Rathbun, 1902c) 

= Parapenaeus kishinouyei Rathbun, 1902c: 288; Plate 12, figs 13-15. [Tagus Cove, reef N of Tagus Hill, 

Albemarle Island] 

Metapenaeopsis kuboi Ivanov & Hassan, 1976a: 1302; Figs 4, 5d, 6f. [Western Indian ocean, 23°26'S 33°31'E, 

410 m] 

Metapenaeopsis kyushuensis (Yokoya, 1933) 

= Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sôyô-Maru stns 296, E of Tanegasima Island, Japan, 

219 m; 317, coast of Miyazaki-ken, Japan, 97 m; 428, W of Amakusa Islands, Japan, 119 m 

Metapenaeopsis lamellata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penaeus lamellatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, Figs 4-5. [Japan; lectotype 

designated by Yamaguchi & Baba, 1993] 

Metapenaeopsis lata Kubo, 1949a 

= Metapenaeopsis latus Kubo, 1949a: 434; Figs 8G, 46G, 76N, T, 81B, 148H, 150. [Off Nagashima, Mie 

Prefecture, Japan, circa 350 m] 

? = Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sôyô-Maru stns 296, east of Tanegasima Isl., 219 

m; 317, coast of Miyazaki-ken, 97 m; 428, west of Amakusa Isls., 119 m] 

Metapenaeopsis laubieri Crosnier, 1991: 238; Figs 51-53. [Albatross 1907-1910, stn 5160, 2.75 mi SW de l’île 

Tinakta Island, Malaisie, 5°12'40"N 119°55'10"E, 22 m] 

Metapenaeopsis lindae R.J.G. Manning, 1988: 97; Fig. 2A-D. [Bell Buoy, NW Rottnest Island (31°59'S 

115°34'E), south Western Australia] 

Metapenaeopsis liui Crosnier, 1987a: 434; Figs 3d, 10b-d, 13d-f, 14a, 15. [MUSORSTOM I, stn 40, Philippines, 

13°57.4'N 120°27.8'E, 287-265 m] 

Metapenaeopsis mannarensis De Bruin, 1965: 88; Fig. 2a-d. [Ceylon, off Mannar, 6 fms] 

Metapenaeopsis manningi Crosnier, 1994a: 344; Figs 4-6. [Anton Bruun, Cruise 9, stn 499, Somalie, 10°03'N 

51°15'E, 31-39 m] 

Metapenaeopsis marquesas Crosnier, 1991: 244; Figs 57-59. [Nat. Geogr. Marquesas Exped. stn UP II/Haul 

5, côte ouest de Ua Pou, Iles Marquises, Polynésie, 48-51 m] 

Metapenaeopsis martinella Pérez Farfante, 1971a: 16; Figs 9-10, 13B. [Oregon stn 4252, off Acaraú, Ceará, 

Brazil, 2°10'S 39°52'W, 137 m] 

Metapenaeopsis menoui Crosnier, 1991: 180; Figs 6c-d, 8-9. [Reves 2 stn 19, Seychelles, 5°54.5'S 56°19.4'E, 

30-35 m] 

Metapenaeopsis miersi (Holthuis, 1952a) 

= Penaeopsis miersi Holthuis, 1952a: 80; Fig. 19. [nomen novum for Penæus velutinus sensu Miers, 

1881a nec Dana, 1852a and nomen novum for Metapenæopsis pubescens Bouvier, 1905b nec Penæus 

pubescens Stimpson, 1871] 

= Metapenæopsis pubescens Bouvier, 1905b: 981; nec Penæus pubescens Stimpson, 1871. [îles du Cap 

Vert] 

Metapenaeopsis mineri Burkenroad, 1934a: 25; Figs 8-10. [Conception Bay, Lower California, USA] 

Metapenaeopsis mogiensis complanata Crosnier, 1991: 219; Figs 37o-r, 38d. [Soela Cruise 682, stn 128, côte 

nord-ouest de Australie, 19°07.9'S 119°06.9'E, 78 m] 

Metapenaeopsis mogiensis consobrina (Nobili, 1904) 

= Metapenaeus consobrinus Nobili, 1904: 230. [Djibouti; lectotype designation by Crosnier, 1991] 

= Metapenaeus perlarum Nobili, 1905a: 158. [côtes d’Arabie, dragages entre 25°10'N 55°10'E – 24°55'N 

54°40'E, 16-24 m, according to Crosnier, 1991] 

= Metapenaeopsis bruini Starobogatov, 1972: 405, 408; Plate 11, fig. 158. [Sri Lanka, E of Mullaitivu 

lighthouse, off Mannar, 6 fms; nomen novum for M. mogiensis sensu De Bruin, 1965 nec Rathbun, 

1902b] 

Metapenaeopsis mogiensis intermedia Crosnier, 1991: 222; Figs 37s-w, 38e. [MUSORSTOM 3, stn CP 142, 

Philippines, 11°47'N 123°02'E, 26-27 m]


Metapenaeopsis mogiensis mogiensis (Rathbun, 1902b) 

= Parapenaeus mogiensis Rathbun, 1902b: 39; Figs 6-8. [Mogi, Japan; lectotype designation by Crosnier, 

1991] 

Metapenaeopsis novaeguineae (Haswell, 1879) 

= Penaeus Novae-Guineae Haswell, 1879: 43. [Chevert Expedition, Katow, Papua New Guinea] 

Metapenaeopsis palmensis (Haswell, 1879) 

= Penaeus Palmensis Haswell, 1879: 43. [Palm Island, Queensland, Australia] 

= Metapenaeopsis barbeensis Hall, 1962: 32; Figs 118-118f. [35 miles ESE of St. Barbe (= Pejantan), 

0°07'S 107°46'E, 20 fms (37 m)] 

Metapenaeopsis parahilarula Crosnier, 1991: 232; Figs 45e, 47. [MUSORSTOM 3, stn CP 121, Philippines, 

12°08.3'N 121°17.3'E, 73-84 m] 

Metapenaeopsis parapalmensis Crosnier, 1994b: 313; Figs 37-38. [Albatross stn 5159, archipel de Sulu, 

groupe des Tawitawi, île Tinakta, 5°11'50"N 119°54'E, 18-22 m] 

Metapenaeopsis persica Crosnier, 1991: 210; Figs 32, 33a-d. [Dammam, district d’Hasa, Arabie Saoudite, 

Golfe Persique, 26°25'N 50°06'E] 

Metapenaeopsis philippii (Spence Bate, 1881) 

= Penaeus Philippii Spence Bate, 1881: 181. [Challenger stn 201, 7°3'N 121°48'E, 150 m; lectotype designation 

by Crosnier, 1987a] 

= Penaeus philippinensis Spence Bate, 1888: 261; Plate 35, figs 2-2", 3-3". [Challenger stn 201, 7°3'N 

121°48'E, 150 m; lectotype designation by Crosnier, 1987a] 

Metapenaeopsis propinqua Crosnier, 1991: 186; Figs 12-13. [”Patate bayonnaise”, lagon est, Nouvelle-Calédonie, 

20 m] 

Metapenaeopsis provocatoria longirostris Crosnier, 1987a: 435, Figs 1d, 7g-j, 8m-o. [Soela Cruise 184 stn 08, 

Australie, 19°20.2'S 115°44.1'E, 306-308 m] 

Metapenaeopsis provocatoria provocatoria Racek & Dall, 1965: 48; Fig. 10A-D. [Challenger stn 31, NNE of 

Cape Moreton, Queensland, Australia, 80-90 fms] 

Metapenaeopsis provocatoria owstoni Shinomiya & Sakai, 2000: 123; Figs 7, 8a-f, 9a-m, 11, 12. [Okinose, 

Sagami Bay, Japan, ca. 35°N 139°35'E] 

Metapenaeopsis proxima Crosnier, 1991: 258; Figs 68-70, 78a. [REVES 2 stn 12, Seychelles, 5°13.5'S 56°08.5'E, 

60 m] 

Metapenaeopsis quadrilobata Crosnier, 1991: 199; Figs 23-24. [Nosy Be (Pointe Lokobe), Madagascar, 10 m] 

Metapenaeopsis quinquedentata (De Man, 1907a) 

= Metapeneus quenquedentatus De Man, 1907a: 133. [Siboga stn 109, off Tongkil Island, Sulu Archipelago, 

Philippines, 13 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis richeri Crosnier, 1991: 280; Figs 88-89. [CORAIL 2, stn DW 147, îlles Chesterfield, 

19°36.87'S 158°13.52'E, 25 m] 

Metapenaeopsis rosea Racek & Dall, 1965: 29; Figs 2D, 3; Plate 1, fig. 4; Plate 4, Figs 7, 8; Plate 9, fig. 4. 

[Mackay, Queensland, Australia, 4 fms] 

Metapenaeopsis scotti Champion, 1973: 195; Figs 3A, 4A-D. [South Africa, Natal, off Durban, 160 fms] 

Metapenaeopsis sibogae (De Man, 1907a) 

= Metapeneus sibogae De Man, 1907a: 131. [Siboga stns 306, 8°27'S 122°54.5'E, 247 m; 312, Saleh-bay, 

north coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

Metapenaeopsis sinica Liu & Zhong, 1988: 3, 224, 270; Fig. 136. [Western Guangdong, South China Sea, 64 m] 

= Metapenaeopsis sinensis Liu & Zhong, 1988: 24, 216 (key). [nomen nudum] 

Metapenaeopsis sinuosa Dall, 1957 

= Metapenaeopsis sinuosus Dall, 1957: 176; Fig. 14A-F. [Off shore island (Kennedy Sound), Cumberland 

Group, NE coast of Australia, 16 m] 

Metapenaeopsis smithi (Schmitt, 1924a) 

= Penaeopsis smithi Schmitt, 1924a: 62; Figs 1b-c, 2a-c. [Caracas Baai, Curaçao] 

Metapenaeopsis spatulata Crosnier, 1991: 224; Figs 40-41. [MUSORSTOM 3, stn CP 142, Philippines, 

11°47'N 123°02'E, 26-27 m] 

Metapenaeopsis spiridonovi Crosnier, 1991: 268; Figs 77a-f, 78d. [REVES 2 stn 33, Seychelles, 4°25.9'S 

54°39.0'E, 45-60 m]


Metapenaeopsis stokmani Burukovsky, 1990: 189; Fig. 1A. [25°39'S 85°24'W, 160-192 m] 

Metapenaeopsis stridulans (Alcock, 1905) 

= Metapenaeus stridulans Alcock, 1905: 518, 526. [from Orissa to Palk Strait, 20-35 fms; Gulf of Martaban, 

20 fms; various places in the Andamans, 20 fms] 

? = Metapenaeopsis tchekunovae Starobogatov, 1972: 402, 413; Plate 9, fig. 107. [Arabian Sea off Pakistan, 

off Indus delta, 23°48.5' - 23°47.7'N 67°17.5' - 67°18.7'E] 

Metapenaeopsis tarawensis Racek & Dall, 1965: 46; Fig. 9A-D. [Tarawa Group, Gilbert Islands] 

Metapenaeopsis tenella Liu & Zhong, 1988: 242; Fig. 146. [East of Hainan Island, South China Sea, 270 m] 

Metapenaeopsis toloensis Hall, 1962 

= Metapenaeopsis tolœnsis Hall, 1962: 33; Figs 119-119d. [Central part of South China Sea, 6°13'N 

107°49'E, 40 fms; 5°51'N 107°53'E, 38 fms] 

Metapenaeopsis vaillanti (Nobili, 1904) 

= Metapenaeus vaillanti Nobili, 1904: 229. [Suez; Mer Rouge] 

? = Metapenaeopsis philippii var. Attaqa Al-Kholy & El-Hawary, 1970: 361. [Gulf of Suez till El-Adabiah, 

5-8 fms] 

Metapenaeopsis velutina (Dana, 1852a) 

= Penaeus velutinus Dana, 1852a: 27. [Albatross 1902, stn 3853, côte sud de l’île Molokai, Hawaii, 210- 

245 m; neotype designation by Crosnier, 1991] 

= Metapenaeopsis insona Racek & Dall, 1965: 41; Fig. 6A-B; Plate 2, fig. 3; Plate 10, fig. 2. [7 miles off 

Long Reef, New South Wales, Australia, 73 m] 

= Metapenaeopsis caliper Liu & Zhong, 1988: 238, 269; Fig. 145. [S of mouth of Pearl River, South 

China Sea, 138 m] 

Metapenaeopsis wellsi Racek, 1967: 251; Plates 12-13. [Exmouth Gulf, Western Australia] 

Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a 

= Mangalura Miers, 1878 (type species Penæus dobsoni Miers, 1878, by monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in Opinion 864 in 1969: ruled under 

the plenary powers not to be given precedence over Metapenaeus Wood-Mason, 1891) 

= Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus affinis H. Milne 

Edwards, 1837 [in Milne Edwards, 1834-1840], by original designation, gender masculine) 

Metapenaeus affinis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus affinis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 416. [côte de Malabar] 

= Penaeus mutatis Lanchester, 1901: 572; Plate 34, fig. 6. [Type locality not indicated; lectotype designation 

by Miquel, 1982] 

= Metapenaeus necopinans Hall, 1956: 83; Plate 11, fig. 15; Plate 12, fig. 16. [Jurong, Singapore] 

= Metapenaeus ivanovi Hassan, 1978: 385; Figs 1-2. [Arabian Gulf, 29°145'N 49°30'E, 25 m] 

Metapenaeus alcocki M.J. George & Rao, 1968: 146; Fig. 1. [Gulf of Kutch, Northwest coast of India, 

22°48'N 70°03'E, 3-12 m] 

Metapenaeus anchistus (De Man, 1920a) 

= Penaeopsis intermedia anchista De Man, 1920a: 5; Plate 1, Figs 3-3d. [Ternate, Indonesia; lectotype 

designation by Miquel, 1982] 

Metapenaeus arabicus Hassan, 1978: 387; Figs 3-5. [Arabian Gulf, 29°145'N 49°30'E, 25 m] 

Metapenaeus bennettae Racek & Dall, 1965: 74. [Lake Budgewoi, Tuggerah Lakes, New South Wales, Australia] 

Metapenaeus brevicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus brevicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 417. [côtes de l’Inde] 

= Penaeus avirostris Dana, 1852a: 27. [Singapore] 

Metapenaeus conjunctus Racek & Dall, 1965: 64; Fig. 12; Plate 5, figs 8-9; Plate 11, fig. 1. [Shallow brackish 

water at mouth of Tuaran river, Sabah, N Borneo, Malaysia, 4-5 fms] 

Metapenaeus dalli Racek, 1957: 4; Plate 1, figs 1-3; Plate 2, figs 1-2; Plate 3, fig. 3. [Peel Inlet, Murray River, 

West Australia] 

Metapenaeus demani demani (Roux, 1921) 

= Penaeopsis demani Roux, 1921: 599; Plate 16, figs 4-12. [Varen and Lorentz Rivers, New Guinea]


Metapenaeus demani stephani Miquel, 1982: 74; Figs 24, 25a, c, 26a-e. [Port Moresby, Golfe de Papoua, 

Nouvelle-Guinée] 

Metapenaeus dobsoni (Miers, 1878) 

= Penæus dobsoni Miers, 1878: 302; Plate 17, fig. 2. [Mangalur (Mangalore), west coast of India] [as 

Mangalura dobsoni on page 303] 

= Metapenaeus dobsoni choprai Holthuis, 1980b: 22. [nomen nudum] 

Metapenaeus eboracensis Dall, 1957: 193; Fig. 19a-g. [Mouth of Norman River, Gulf of Carpentaria, 

Queensland, Australia, 2-4 m] 

Metapenaeus elegans De Man, 1907a: 130. [Siboga stn 121, Manado, Cèlébes, Indonesia, 55 m] 

= Metapenaeus singaporensis Hall, 1956: 84; Plate 12, figs 17-19. [Jurong, Singapore] 

Metapenaeus endeavouri (Schmitt, 1926a) 

= Penaeopsis endeavouri Schmitt, 1926a: 329; Plate 59, figs 1-3; Plate 68, fig. 4. [FIS Endeavour expedition, 

S coast of Queensland, Australia] 

Metapenaeus ensis (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penoeus Ensis De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 2. [Japan; lectotype designation 

by Miquel, 1982] 

= Penaeus Mastersii Haswell, 1879: 42. [80 km WNW of Darwin, Northern Territory, Australia] 

= Penaeus incisipes Spence Bate, 1888: 257; Plate 34, fig. 2. [Challenger stns 190, Arafura Sea, south of 

Papua, 8°56'S 136°5'E, 49 fms; 203, off Panay, Philippines, 11°6'N 123°9'E, 20 fms] 

= Metapenaeus ensis var. baramensis Hall, 1962: 23. [30 miles NNE of Tanjong Baram, 4°58'N 113°58'E, 

77 m] 

= Metapenaeus philippinensis Motoh & Muthu, 1979: 1351; Figs 1-2. [Himamaylan, Philippines, 10- 

15 m] 

Metapenaeus insolitus Racek & Dall, 1965: 69; Fig. 13A-C; Plate 6, figs 3, 4; Plate 11, fig. 4. [Chambers Bay, 

Northern Territory, Australia, 30 m] 

Metapenaeus intermedius (Kishinouye, 1900) 

= Penaeus intermedius Kishinouye, 1900: 21, unnumbered textfig. [province of Tosa] 

Metapenaeus joyneri joyneri (Miers, 1880) 

= Penaeus Joyneri Miers, 1880: 458; Plate 15, figs 8-10. [Yokohama, Japan] 

= Penaeus pallidus Kishinouye, 1897: 275. [Tokyo Bay, Ise area, Seto Inland Sea; Kagoshima Bay] 

Metapenaeus joyneri formosus Lee & Yu, 1977: 101; Fig. 71. [Donggang, Pingtong County, SW Taiwan] 

Metapenaeus krishnatrii Silas & Muthu, 1976: 645; Plate 1; fig. 1. [Corbyn’ Cove, Port Blair, Andaman Islands] 

Metapenaeus kutchensis P.C. George, M.J. George & Rao, 1963: 284; Fig. 1. [Gulf of Kutch, N.W. India, 

3-12 m] 

Metapenaeus lysianassa (De Man, 1888a) 

= Penaeus lysianassa De Man, 1888a: 290. [Mergui Archilepago, Burma] 

= Metapenaeus lysianassa var. malaccænsis Hall, 1962: 24. [Kuala Sanglang, 1 m] 

Metapenaeus macleayi (Haswell, 1879) 

= Penæus macleayi Haswell, 1879: 40. [Port Jackson, New South Wales, Australia] 

= Penaeus haswelli Phillips, 1925: 3; Plate 2, fig. 1. [shallow waters of New South Wales] 

Metapenaeus monoceros (Fabricius, 1798) 

= Penaeus monoceros Fabricius, 1798: 409. [Type locality restricted by Holthuis & Gottlieb, 1958 to 

Tranquebar, Madras Province, India] 

? = Penaeopsis spinulicauda Stebbing, 1914b: 17; Plate 68. [Durban Bay] 

? = Metapenaeus cognatus Nobili, 1904: 229. [Djibouti, Mer Rouge] 

? = Metapenaeus Deschampsi Nobili, 1903a: 452; Plate 2, fig. 1. [Pondichéry, Foce dell’Arian Koupur; 

Mahè] 

Metapenaeus motohi Shinomiya & Sakai, 2009: 1068; Fig. 1. [Tigbauan Panay I., Philippines] 

Metapenaeus moyebi (Kishinouye, 1896) 

= Penaeus moyebi Kishinouye, 1896: 373. [Japan; Makassar, Celebes, Indonesia] 

= Metapenaeus burkenroadi Kubo, 1954: 92; Fig. 1A-D. [Japan]


Metapenaeus papuensis Racek & Dall, 1965: 66; Plate 3, fig. 3; Plate 6, figs 1-2; Plate 11, fig. 2. [Mouth of 

Panaroa river, Gulf of Papua, Papua New Guinea] 

= Metapenaeus bengalensis Tirmizi, 1971b: 242; Figs 1-3. [Bay of Bengal] 

Metapenaeus stebbingi Nobili, 1904 

= Metapenaeus Stebbingi Nobili, 1904: 229. [Mer Rouge; Suez; full description in Nobili, 1906a] 

Metapenaeus suluensis Racek & Dall, 1965: 61; Fig. 11A-B; Plate 5, figs 6-7; Plate 10, fig. 8. [Philippines, 

Sulu Sea, 20 fms] 

Metapenaeus tenuipes Kubo, 1949a: 348; Figs 7R, T, 22B, 31G, H, 47G, 62B-B', 74N-M, S-T, 81H, K, 125G, 

127, 129F, 130. [S coast of Borneo, Indonesia] 

= Metapenaeus spinulatus Kubo, 1949a: 355; Figs 7T, 47G, 74M, S, 81K, 129F, 130. [Bangkok, Thailand] 

Mierspenaeopsis Sakai & Shinomiya, 2011 

= Mierspenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus sculptilis Heller, 1862a, by original 


Mierspenaeopsis cultrirostris (Alcock, 1906) 

= Parapenaeopsis sculptilis var. cultrirostris Alcock, 1906: 39; Plate 7, fig. 23. [India: Orissa and Ganjam; 

Sunderbunds] 

Mierspenaeopsis hardwickii (Miers, 1878) 

= Penæus hardwickii Miers, 1878: 300; Plate 17, figs 1-1a. [Indian Seas?] 

Mierspenaeopsis indica (Muthu, 1972) 

= Parapenaeopsis indica Muthu, 1972: 174; figs 1-6. [Kakinada Bay, Kakinada, east coast of India, 

16°56.5'N 82°16.5'E, 1-4 m] 

Mierspenaeopsis sculptilis (Heller, 1862a) 

= Penaeus sculptilis Heller, 1862a: 528. [Java] 

Parapenaeopsis Alcock, 1901 

= Parapeneopsis Alcock, 1901 (type species Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne 

Edwards, 1834-1840], by original designation, gender feminine; name emended under the plenary 

powers of the ICZN to Parapenaeopsis and placed in that emended spelling on the Official List of 

Generic Names in Zoology in Opinion 864 in 1969) 

Parapenaeopsis acclivirostris Alcock, 1905: 522, 530. [Persian Gulf; in Palk Strait; at Madras; off the Vizagapatam 

and Ganjam coasts] 

Parapenaeopsis aroaensis Hall, 1962 

= Parapenæopsis aroaensis Hall, 1962: 27; Figs 109-109a. [20 mls NNE of Aroa Islands, 3°20'N 100°39'E, 

51 m] 

Parapenaeopsis balli Burkenroad, 1934a: 64; Fig. 17. [Acajutla, El Salvador, 13°36'N 89°50'W] 

Parapenaeopsis gracillima Nobili, 1903b: 4; Fig. 1. [Buntal, Borneo] 

Parapenaeopsis longirostris Chandra & Bhattacharya, 2004: 23; Fig. 1. [Ongaria Ghat, Baleshwar, Orissa] 

Parapenaeopsis nana Alcock, 1905: 522, 529. [Off Ganjam and Orissa coasts up to 68 fms; at Madras] 

Parapenaeopsis stylifera (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [les environs de 

Bombay] 

= Parapeneopsis stylifera var. coromandelica Alcock, 1906: 37; Plate 7, figs 20, 20a-c. [Madras, Coromandel 

Coast; lectotype designation by Ravindranath, 1989] 

= Parapenaeopsis stylifera var. cochinensis M.J. George, 1974: 421; Fig. 1b, d. [unavailable name under 

Art. 15.2] 

Parapenaeus Smith, 1885b 

= Parapenæus Smith, 1885b (type species Penaeus longirostris Lucas, 1846, by original designation, 


= Neopenæopsis Bouvier, 1905a (type species Neopenæopsis paradoxus Bouvier, 1905a, by original designation 

and monotypy, gender feminine)


Parapenaeus americanus Rathbun, 1901: 102; Plate 2. [Mayaguez Harbour, 220-225 fms] 

Parapenaeus australiensis Dall, 1957: 179; Fig. 15A-F. [Off Newcastle, New South Wales, Australia] 

= Parapenaeus australiensis forma nodosus Crosnier, 1986a: 313, 339. [types not indicated; unavailable 

under Art. 15.2] 

Parapenaeus cayrei Crosnier, 2005: 262; Figs 3, 4A-C. [BORDAU 2, stn CP 1541, Îles Tonga, 21°15'S 

175°14'W, 319-333 m] 

Parapenaeus fissuroides erythraeus Crosnier, 1986a: 329; Figs 5i, 6k, m, 8c-d. [Pola stn 117, Mer Rouge, 

20°16'N 37°33'E, 638 m] 

Parapenaeus fissuroides fissuroides Crosnier, 1986a: 321; Figs 5e-f, 6d-f, 7c, 8a. [Hong Kong, 19°22.5'N 

114°07.5'E, 700-795 m] 

Parapenaeus fissuroides indicus Crosnier, 1986a: 325; Figs 5g, h, 6g-j, l, 7d, f-g, 8b. [Vauban stn CH 80, 

Madagascar, 25°02.7'S 47°05.8'E, 65-70 m] 

Parapenaeus fissurus (Spence Bate, 1881) 

= Penaeus fissurus Spence Bate, 1881: 180. [south of New Guinea, 50 fms] 

= Penaeus fissurus Spence Bate, 1888: 263; Plate 36, fig. 1. [Challenger stns 190, 8°56'S 136°5'E, 49 fms; 

204A, 12°43'S 122°9'E, off Tablas Island, 100 fms; 204B, 12°46'S 122°10'E, off Tablas Island, 115 fms; 

209, 10°14'N 123°54'E, off Zebu, 95-100 fms] 

Parapenaeus investigatoris Alcock & Anderson, 1899: 279. [Investigator stns 233 (Andaman Sea, 13°17'15''N 

93°10'25''E), 185 fms; 235 (Andaman Sea, 14°13'N 93°40'E), 370-419 fms; 166 (Bay of Bengal, 

13°34'55''N 80°32'12''E), 133 fms] 

Parapenaeus kensleyi Crosnier, 2005: 258; Figs 1-2. [HALIPRO I, stn CP 853, Nouvelle-Calédonie, 21°45'S 

166°37'E, 241-250 m] 

Parapenaeus lanceolatus Kubo, 1949a: 405; Figs 4C, 7D', 21C, 28G-I, 61C, C', 75C-I, 78I, 143C, G, 144. [Heta, 

Shizuoka Prefecture and Kumano-nada, off Owase, Mie Prefecture, Japan, 300-350 m] 

Parapenaeus longipes Alcock, 1905: 520, 525. [off Malabar coast (Mangalore), 21-26 fms; off Orissa and 

Ganjan coasts, 20-68 fms; off Vizagapatam coast, 7,5-23 fms; S coast of Ceylon, 20 fms] 

= Parapenaeus longipes forma denticulatus Crosnier, 1986a: 311, 312, 347. [Indonésie et Philippines, 

exact type locality not indicated, see Crosnier, 1986a; unavailable under Art. 15.2] 

Parapenaeus longirostris (Lucas, 1846) 

= Peneus Cocco Prestandrea, 1833: 6. [Mari di Messina; suppressed under the plenary powers for the 

purposes of the Principle of Priority but not those of the Principle of Homonymy in Opinion 611 in 

1961] 

= Penaeus longirostris Lucas, 1846: 46; Plate 4, fig. 6. [Algerie, Alger, aux environs du cap Matifou] 

= Penæus bocagei Johnson, 1863: 255. [mouth of the Tagus] 

= Penaeus lividus Filhol, 1885b: Plate 1. [Type locality not indicated, 500-600 m] 

Parapenaeus murrayi Ramadan, 1938: 74; Fig. 15a-c. [John Murray Expedition stn 105 (°36'12''N 

39°13'12''E), Zanzibar area, 238-293 m] 

Parapenaeus perezfarfantae Crosnier, 1986a: 339; Figs 12a-b, 13a-b, 14a. [Albatross expedition stn 5183, 

Philippines, 10°32'48"N 122°26'E, 176 m] 

Parapenaeus politus (Smith, 1881) 

= Penaeus politus Smith, 1881: 444. [Fish Hawk stn 878 (39°55'N 70°54'W), 142 fms, off the eastern end 

of Long Island, known as the Block Island] 

= Neopenæopsis paradoxus Bouvier, 1905a: 747. [dans la mer des Antilles par 84 et 91 brasses de profondeur] 

Parapenaeus ruberoculatus Hall, 1962: 31; Figs 115-115b. [30 Miles NNE of Tanjong Baram, 5°04'N 113°48'E, 

183 m] 

Parapenaeus sextuberculatus Kubo, 1949a: 403 (partim); Figs 7F', 21B, 28D-F, 61A-A', 75B, H, 78H, 143B, F 

(nec Fig. 47R = P. fissuroides fissuroides Crosnier, 1986a). [Heta and Maisaka, Shizuoka Prefecture, 

Japan, circa 330 m] 

Pelagopenaeus Pérez Farfante & Kensley, 1997 

= Pelagopenaeus Pérez Farfante & Kensley, 1997 (type species Penaeus balboae Faxon, 1893, designated 

by Pérez Farfante & Kensley, 1997, gender masculine)


Pelagopenaeus balboae (Faxon, 1893) 

= Penaeus balboae Faxon, 1893: 211. [Albatross stn 3371 (off Isla del Coco, Costa Rica, 5°26'20"N 

86°55'00"W), 770 fms] 

= Penaeus meridionalis Lenz & Strunck, 1914: 298, Plate 15, figs 1-15. [Deutsche Südpolar-Expedition, 

30°21'S 14°2'W, 10 m] 

Penaeopsis Spence Bate, 1881 

= Penaeopsis Spence Bate, 1881 (type species Penaeopsis serratus Spence Bate, 1881, designated by 

Bouvier, 1905b, gender feminine) 

Penaeopsis balssi Ivanov & Hassan, 1976b: 1; Figs 1-2. [off central Mozambique, 25°26'S (not 23°26'S as 

stated), 33°31'E, 410 m] 

Penaeopsis challengeri De Man, 1911a: 76. [nomen novum for Penaeus serratus Spence Bate, 1881] 

= Penaeus serratus Spence Bate, 1881: 182. [Challenger stn 173, off Matuku, Fiji Islands, 19°09'35"S 

179°41'50"E, 315 fms, lectotype designation by Pérez Farfante, 1980a] 

Penaeopsis eduardoi Pérez Farfante, 1977a: 172; Figs 1-4. [Albatross stn 5116, Balayan Bay, Luzon Island, 

Philippines, 13°41'00"N 120°47'05"E, 200 fms] 

Penaeopsis jerryi Pérez Farfante, 1979: 209; Figs 1-3. [John Murray Expedition stn 16, off Berbera, Somalia, 

Gulf of Aden, 10°29'48"N 45°01'48"E, 186 m] 

Penaeopsis mclaughlinae Crosnier, 2006: 334; Figs 2-5. [CHALCAL 2 stn CC 1, Nouvelle-Calédonie, 

24°54.96'S 168°21.91'E, 500 m] 

Penaeopsis rectacuta (Spence Bate, 1881) 

= Penaeus rectacutus Spence Bate, 1881: 180. [among the Philippine Islands, about 100 fms; according 

to Pérez Farfante, 1980a: Challenger stn 209, between Bohol and Cebu, Philippines, 10°14'N 123°54'E, 

95 fms] 

Penaeopsis serrata Spence Bate, 1881 

= Penaeopsis serratus Spence Bate, 1881: 183. [Gulf of Mexico; according to Pérez Farfante, 1980a: 

Blake stn 275, off Barbados, Gulf of Mexico, 12°58'33"N 59°36'45"W, 218 fms; name placed on the 

Official List of Specific Names in Zoology in Opinion 864 in 1969] 

= Parapenaeus megalops Smith, 1885b: 172. [Albatross stns 2125, S of Curaçao, 11°43'N 69°09'30"W, 208 

fms; 2143, Golfo de Urabá, 9°30'45"N 76°25'30"W, 155 fms] 

= Artemesia talismani Bouvier, 1905b: 982. [le Talisman captura en quantité assez grande au large des 

côtes du Maroc et du Sahara; Talisman stn 72, off Guerguerat, Western Sahara, 25°41'N 15°56'W (of 

Greenwich) 18°16'W (of Paris) on label accompanying specimen), 410 m, see Pérez Farfante, 1980a] 

= Penaeopsis serratus var. antillensis A. Milne-Edwards & Bouvier, 1909: 226; Plate 3, fig. 10; Plate 4, 

fig. 5. [Blake stn 148, off St Kitts, 208 fms] 

Penaeus Fabricius, 1798 

= Penaeus Fabricius, 1798 (type species Penaeus monodon Fabricius, 1798, designated by Latreille, 

1810, gender masculine) 

Penaeus esculentus Haswell, 1879: 38. [Port Jackson, New South Wales, Australia] 

Penaeus monodon Fabricius, 1798: 408. [Bay of Batavia [=Jakarta], Indonesia; neotype designation by 

Holthuis, 1949a] 

= Penaeus carinatus Dana, 1852a: 27. [Singapore] 

= Penaeus tahitensis Heller, 1862a: 528. [Taiti] 

= Penaeus semisulcatus exsulcatus Hilgendorf, 1879: 843. [Quelimane, Mozambique] 

= Penaeus coeruleus Stebbing, 1905: 77; Plates 21-21bis. [Nahoon River on E coast near East London, 

South Africa] 

? = Penaeus durbani Stebbing, 1917a: 442; Plate 22. [Durban Bay] 

= Penaeus bubulus Kubo, 1949a: 296; Figs 1G, 7B, 15U-B', 24K-M, 37, 49C, 53, 58D, 67N-Q, 73F, L, 77P, 

113, 114. [Hatazawa, Chiba Prefecture: Miya; Aichi Prefecture; Tainan, Formosa; Palau; Batavia; 

Menado, Celebes] 

Penaeus semisulcatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 1. [Japan] 

? = Penaeus Manilensis de Procé, 1822: 134. [Manille]


= Penaeus ashiaka Kishinouye, 1900: 14; Plate 3; Plate 7, fig. 4. [southern half of our coast] 

= Penaeus semisulcatus paucidentatus Parisi, 1919: 65; Plate 5, fig. 5. [Misaki, Baia di Sagami] 

= Penaeus monodon manillensis Villaluz & Arriola, 1938: 39; Plate 3, fig. 1. [Manila Bay, Philippines] 

Protrachypene Burkenroad, 1934a 

= Protrachypene Burkenroad, 1934a (type species Protrachypene precipua Burkenroad, 1934a, by 

monotypy, gender feminine) 

Protrachypene precipua Burkenroad, 1934a: 43. [Bella Vista Beach, Panama City, Panama] 

Rimapenaeus Pérez Farfante & Kensley, 1997 

= Rimapenaeus Pérez Farfante & Kensley, 1997 (type species Trachypenaeus faoe Obarrio, 1954, by 


Rimapenaeus byrdi (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) byrdi Burkenroad, 1934a: 51; Fig. 13. [Panama City Market, Panama] 

Rimapenaeus constrictus (Stimpson, 1871) 

= Penaeus constrictus Stimpson, 1871: 135. [Beaufort, N.C., 4 fms; Charleston Harbour] 

= Penaeopsis agassizii Bouvier, 1905a: 748. [provient de Sombrero] 

Rimapenaeus faoe (Obarrio, 1954) 

= Trachypenaeus faoe Obarrio, 1954: 3. [Golfo de Panamá, en el Pacífico] 

Rimapenaeus fuscina (Pérez Farfante, 1971b) 

= Trachypenaeus fuscina Pérez Farfante, 1971b: 637; Figs 1-2, 3A, 4A-F, 5A, 6. [Off Cocodrillo Chiapas, 

Pacific coast of México, 22 m] 

Rimapenaeus pacificus (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) similis pacificus Burkenroad, 1934a: 50. [Pearl Islands, Gulf of Panama, 

Panama, 8°29'40"N 78°52'30"W, 19-24 fms] 

Rimapenaeus similis (Smith, 1885b) 

= Parapenaeus constrictus similis Smith, 1885b: 175. [Albatross stn 2121, Gulf of Paria, 10°37'40"N 

61°42'40"W, 31 fms] 

Tanypenaeus Pérez Farfante, 1972 

= Tanypenaeus Pérez Farfante, 1972 (type species Tanypenaeus caribaeus Pérez Farfante, 1972, by original 


Tanypenaeus caribaeus Pérez Farfante, 1972: 187. [Pillsbury stn 788, off Punta Faro, Colombia, 11°08'N 

74°47'W, 155-157 m] 

Trachypenaeopsis Burkenroad, 1934a 

= Trachypeneopsis Burkenroad, 1934a (type species Metapenaeus mobilispinis Rathbun, 1915, by original 


= Trachypenaeopsis [emendment of Trachypeneopsis Burkenroad, 1934a, under the plenary powers of 

the ICZN in Opinion 684 in 1969] 

Trachypenaeopsis mobilispinis (Rathbun, 1915) 

= Metapenaeus mobilispinis Rathbun, 1915: 117. [Cave Round Bay, Saba, Netherlands Antilles, 7 m] 

Trachypenaeopsis minicoyensis Thomas, 1972: 117; Fig. 1. [western lagoon of Minicoy island, Laccadive 

Archipelago, 3 m] 

Trachypenaeopsis richtersii (Miers, 1884a) 

= Penæus richtersii Miers, 1884a: 564; Plate 52, fig. A. [Cerf Island, 10 fms] 

Trachypenaeus Alcock, 1901 

= Trachypeneus Alcock, 1901 (type species Penaeus anchoralis Spence Bate, 1881, by original designation, 


= Trachypenaeus [emendment of Trachypeneus Alcock, 1901, under the plenary powers of the ICZN in 

Opinion 864 in 1969]


Trachypenaeus anchoralis (Spence Bate, 1881) 

= Penaeus anchoralis Spence Bate, 1881: 181. [south of New Guinea, 28 fms] 

= Penaeus anchoralis Spence Bate, 1888: 258; Plate 35, fig. 1. [Challenger stns 188, 9°59'S 139°42'E, 

Arafura Sea, south of Papua, 28 fms; 190, 8°56'S 136°5'E, Arafura Sea, south of Papua, 49 fms; off 

Yokohama, 5-20 fms; between stns 190-191, near the Arrou Islands in the Arafura Sea] 

Trachysalambria Burkenroad, 1934a 

= Trachypeneus (Trachysalambria) Burkenroad, 1934a (type species Penaeus curvirostris Stimpson, 

1860a, by original designation, gender feminine) 

Trachysalambria albicoma (Hayashi & Toriyama, 1980) 

= Trachypenaeus albicomus Hayashi & Toriyama, 1980: 69; Figs 1-2. [Japan, Tosa Bay] 

Trachysalambria aspera (Alcock, 1905) 

= Trachypenaeus asper Alcock, 1905: 523, 531. [fully described and illustrated in Alcock, 1906; type 

locality, Ganjam coast, 20-35 fms, according to Alcock, 1906] 

Trachysalambria brevisuturae (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) brevisuturae Burkenroad, 1934a: 55; Fig. 14. [Acajutla, El Salvador] 

Trachysalambria curvirostris (Stimpson, 1860a) 

= Penaeus curvirostris Stimpson, 1860a: 44. [Tosa Bay, Kochi, Japan, 33°30'N 133°30'E, 35 m; neotype 

designation by Sakaji & Hayashi, 2003] 

= Metapenaeus palaestinensis Steinitz, 1932: 161; Figs 1-3. [aus der Bucht von Haifa] 

Trachysalambria fulva (Dall, 1957) 

= Trachypenaeus fulvus Dall, 1957: 206; Fig. 23A-G. [Townsville, Queensland, Australia] 

= Trachypeneus unicus Hall, 1961: 102. [S of Singapore, Malaysia, 35 fms] 

Trachysalambria longipes (Paul’son, 1875) 

= Penaeus longipes Paul’son, 1875: 125; Plate 19, figs 1-1a. [Red Sea] 

Trachysalambria malaiana (Balss, 1933) 

= Trachypenaeus curvirostris malaiana Balss, 1933: 234. [Gier 16 Exp. 2, Südl. Sumatra, 6°42'S 103°E; 

Gier 2 Exp. 3, Bai von Batavia; Gier 12 Exp. 4, Javasee vor Borneo, 3°42'S 114°30'E; Gier 4, vor Toeban 

(Nordjava); Gier 9 Exp. 20, nördl. Sumatra, 1°20'S 104°43'E; Gier 17 Exp. 4, westlich Borneo, 1°36'S 

109°46'E; Gier 6 Exp. 21, Bangkastrasse, 2°3'S 105°48,5'E; Gier 4 Exp. 20, Javasee, 6°51'S 112°56'E; 

Gier 4 Exp. 11, vor Pekalongan (Nordjava)] 

Trachysalambria nansei Sakaji & Hayashi, 2003: 162; Figs 7-9. [Japan, Tosa Bay, off Kochi, 33°30'N, 133°30'E, 

75 m] 

Trachysalambria starobogatovi (Ivanov & Hassan, 1976a) 

= Trachypenaeus starobogatovi Ivanov & Hassan, 1976a: 1300; Figs 2, 3a. [Western Indian Ocean, 

19°03'S 36°29'E, 25 m] 

Trachysalambria villaluzi (Muthu & Motoh, 1979b) 

= Trachypenaeus villaluzi Muthu & Motoh, 1979b: 57; Figs 1-2. [Tigbauan, Philippines, 7 m] 

Xiphopenaeus Smith, 1869 

= Xiphopeneus Smith, 1869 (type species Xiphopeneus hartii Smith, 1869, by monotypy, gender masculine) 

= Xiphopenaeus [emendment of Xiphopeneus Smith, 1869, under the plenary powers of the ICZN in 

Opinion 864 in 1969] 

Xiphopenaeus kroyeri (Heller, 1862b) 

= Penaeus Kroyeri Heller, 1862b: 425; Plate 2, fig. 51. [Rio Janeiro] 

= Xiphopeneus Hartii Smith, 1869: 27, 40; Plate 1, fig. 1. [Caravelas, Estado da Bahia, Brazil] 

= Xiphopeneus Riveti Bouvier, 1907: 113; Fig. 1. [achetée sur le marché de Païta]


Family SICyoNIIDAe ortmann, 1898 

Sicyonia H. Milne edwards, 1830 

= Sicyonia H. Milne Edwards, 1830 (type species Sicyonia sculpta H. Milne Edwards, 1830, designated 

by Desmarest, 1858, gender feminine) 

= Ruvulus De Natale, 1850 (type species Sicyonia sculpta H. Milne Edwards, 1830, by monotypy, 


= Synhimantites Boeck, 1864 (type species Synhimantites typicus Boeck, 1864, by monotypy, gender 

masculine) 

= Eusicyonia Stebbing, 1914b (substitute name for Sicyonia H. Milne Edwards, 1830, gender feminine) 

Sicyonia abathophila Crosnier, 2003: 321; Fig. 98. [Albatross, Philippines Expedition, 1908-1909, Camp 

Overton, Iligan Bay, Mindanao, Philippines, 8°15,24'N 124°07,18'E, 2-4 m] 

Sicyonia adunca Crosnier, 2003: 265; Figs 42-44, 107F. [BATHUS 2, stn DW 749, au large de la Nouvelle- 

Calédonie, 22°33,4'S 166°26'E, 233-258 m] 

Sicyonia affinis Faxon, 1893: 209. [Albatross stns 3367, off Isla del Coco, Costa Rica, 5°31'30"N 86°52'30"W, 

183 m; 3369, off Isla del Coco, Costa Rica, 5°32'45"N 86°55'20"W, 95 m; 3378, W of Isla de Malpelo, 

Colombia, 3°58'20"N 81°36'00"W, 205 m; 3379, W of Isla de Malpelo, Colombia, 3°59'40"N 81°35'00"W, 

95 m] 

Sicyonia aliaffinis (Burkenroad, 1934a) 

= Eusicyonia aliaffinis Burkenroad, 1934a: 92; Fig. 24. [Pawnee, Pacific coast of southern Mexico, 

14°48'40"N 92°54'40"W, 19-30 fms] 

Sicyonia altirostrum Crosnier, 2003: 297; Figs 73-75, 109G. [MUSORSTOM 6, stn CP 464, Îles Loyauté, 

21°02,3'S 167°31,6'E, 430 m] 

Sicyonia australiensis Hanamura & Wadley, 1998: 701; Figs 1-4. [Southern Surveyor, cruise SS6/96, stn 183, 

SE coast of Australia, 37°18.7'S, 150°16.7'E, 123 m] 

Sicyonia benthophila De Man, 1907a: 143. [Siboga stn 253, near Kai Islands, Indonesia, 304 m] 

Sicyonia bispinosa (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte bispinosus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 9. [Japan] 

Sicyonia brevirostris Stimpson, 1871: 132. [S Florida coast] 

Sicyonia burkenroadi Cobb, 1971: 104; Figs 1-2. [Oregon stn 1083, Gulf of Mexico off Port Isabel, Texas, 

26°13'N 96°45'W, 42 m] 

Sicyonia carinata (Brünnich, 1768) 

= Cancer carinatus Brünnich, 1768: 102. [Mari Adriatico, Dalmatinis Skilla, in finu maris juxta Salonas] 

= Cancer pulchellus Herbst, 1796 [in Herbst, 1791-1796]: 175; Plate 43, fig. 3. [das Veterland ist mir 

unbekannt] 

= Sicyonia sculpta H. Milne Edwards, 1830: 340; Plate 9, figs 1-8. [Baie de Naples] 

= Sicyonia foresti Rossignol, 1962: 145; 2 unnumbered Figs [Ile du Prince, 27 m] 

Sicyonia curvirostris Balss, 1913b: 235. [Fukuura, Sagami Bay, Japon, 150 m] 

Sicyonia dejouanneti Crosnier, 2003: 293; Figs 70-72, 108G, H, 109A-F. [MUSORSTOM 4, stn CP 194, Nouvelle-Calédonie, 

18°52,8'S 163°21,7'E, 545 m] 

Sicyonia disdorsalis (Burkenroad, 1934a) 

= Eusicyonia disdorsalis Burkenroad, 1934a: 96; Figs 25, 36. [Pawnee, Pearl Islands (Archipelago de las 

Perlas), Gulf of Panama, 8°29'40"N 78°52'30"W, 19-24 fms; Golfo de Panamá; W coast of Central 

America] 

Sicyonia disedwardsi (Burkenroad, 1934a) 

= Eusicyonia disedwardi Burkenroad, 1934a: 86; Figs 23, 29, 34. [Pawnee, Conception Bay, Lower California] 

Sicyonia disparri (Burkenroad, 1934a) 

= Eusicyonia disparri Burkenroad, 1934a: 83; Fig. 27. [Pawnee, Bahia San Luis Gonzaga, Lower California, 

Mexico]


Sicyonia dorsalis Kingsley, 1878a: 97. [Fort Jefferson, Florida] 

Sicyonia fallax De Man, 1907a: 141. [Siboga stn 105, Sulu Sea, Indonesia, 6°08'N 121°19'E, 275 m] 

Sicyonia furcata Miers, 1878: 310; Plate 17, fig. 4. [Sulu Archipelago, Philippines] 

= Sicyonia formosa Chan & Yu, 1985: 99; Fig. 1; Plates 5A, B, 6E-F. [Taiwan, 100 m] 

Sicyonia galeata Holthuis, 1952a: 84. [Mbizi Expedition stns A.S.182, Angola, Luanda, 13 miles WSW of 

Cape Ledo, 9°47'S 13°11'E, 35 m; A.S.170, Congo, 10 miles W of Pointe Noire, 4°48'S 11°41'E, 49 m] 

Sicyonia inflexa (Kubo, 1949a) 

= Eusicyonia inflexa Kubo, 1949a: 458; Figs 8O, 48D, 77C, I, 79G, 159, 160A-F. [Off Atsumi, Aichi 

Prefecture, Japan, ca. 300 m; and Kumanonada, off Owase, Mie Prefecture, Japan, ca. 400 m] 

Sicyonia ingentis (Burkenroad, 1938) (Fig. 4) 

= Eusicyonia ingentis Burkenroad, 1938: 88; Figs 31-34. [Zaca stn 127D-1, off E coast of Cedros Island 

(Isla Cedros), 28°05'N 115°09'W, Baja California, Mexico] 

Sicyonia japonica Balss, 1914b 

= Sicyonia lancifer var. japonica Balss, 1914b: 16; Fig. 9. [Dzushi, 50-100 m; Misaki, 200-300 m; Fukuura; 

Yagoshima, 150 m; zwischen Ito und Hatsushima, 150 m] 

Sicyonia komai Crosnier, 2003: 327; Figs 103-104. [Thailande, côte ouest, Ao Tang Knen, Phuket, zone intertidale] 

Sicyonia laevigata Stimpson, 1871: 131. [Charleston] 

= Sicyonia sculpta var. americana De Man, 1907b: 450. [Off Bahia] 

Sicyonia laevis Spence Bate, 1881: 173. [north of New Guinea, 150 fms; fully described and illustrated in 

Spence Bate, 1888; type locality according to Spence Bate, 1888: Challenger stn 219, north of New 

Guinea, 1°54'S 146°39'40"E, 150 fms] 

= Eusicyonia nebulosa Kubo, 1949a: 454; Figs 8N, 48C, 77B, H, 79K, 156G, 157. [Off Heta, Shizuoka 

Prefecture, Japan, ca. 350 m] 

Sicyonia lancifer (Olivier, 1811) 

= Palæmon lancifer Olivier, 1811: 664. [CORINDON stn 295, Indonesia, Makassar, Strait, 1°26,5'S, 

117°02,1'E, 51-54 m; neotype selection by Crosnier, 2003; emendation to S. lancifera by Holthuis, 

1980b is an unjustified emendation] 

= Hippolyte cristatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 10. [Japan; lectotype designation 

by Yamaguchi & Baba, 1993] 

Fig. 4. Sicyonia ingentis (Burkenroad, 1938). Photo by Sammy De Grave.


Sicyonia longicauda Rathbun, 1906: 908; Plate 20, fig. 6 (partim, nec specimens of stn 4002 = S. truncata 

Kubo, 1949; see Crosnier, 2003). [Albatross stn 3865, Pailolo Channel, Hawaii Islands, 21°09'20"N 

156°35'10"W, 468-517 m] 

Sicyonia longicornis Crosnier, 2003: 276; Figs 55-56. [KARUBAR, stn CP 20, îles Kai, Indonésie, 5°15'S 

132°59'E, 769-809 m] 

Sicyonia martini Pérez Farfante & Boothe, 1981: 424; Figs 1-4. [Pillsbury, stn 556, SW of Punta Ana Maria, 

Golfo de Panama, 7°50'30"N 78°49'00"W, 58 m] 

Sicyonia metavitulans Crosnier, 2003: 304; Figs 79-81. [Port Jackson, côte est de l’Australie] 

Sicyonia mixta Burkenroad, 1946: 3; Figs 1-4. [St. Joseph (probably San José, Lower California), Swedish 

Eugenie Expedition #818] 

Sicyonia nasica Burukovsky, 1990: 193, 211; Figs 2A(1-6), 6. [25°04'S 97°28'W, 267-280 m] 

Sicyonia ocellata Stimpson, 1860a: 43. [Portu ”Hong Kong”; in mari Sinensi quoque, lat. bor. 24°; ad prof. 

20 org] 

= Sicyonia ommanneyi Hall, 1961: 110; Plate 21, figs 26-27. [Malaysia, S of Singapore, 45 fms] 

Sicyonia olgae Pérez Farfante, 1980b: 775; Figs 1-3. [Oregon stn 2277, Off Parimaribo, Suriname, 06°37'N 

55°36'W, 35 m] 

Sicyonia parafallax Crosnier, 1995: 193; Figs 3-4. [Mer Rouge, détroit de Bab el Mandeb, 12°43,7'N 

43°15,0'E, 228 m] 

Sicyonia parajaponica Crosnier, 2003: 226; Figs 10-12; 106E-F. [MUSORSTOM 1, stn 7, Philippines, dans 

l’ouest de Luçon, 14°01,0'N, 120°20,0'E, 185-200 m] 

Sicyonia parri (Burkenroad, 1934a) 

= Eusicyonia parri Burkenroad, 1934a: 80; Fig. 22. [Crooked Island, Bahamas] 

Sicyonia parvula (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte parvulus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 6. [Japan] 

Sicyonia penicillata Lockington, 1878a: 164. [Bolinas Bay, Lower California, 14 fms; Angeles Bay, Gulf of 

California] 

Sicyonia picta Faxon, 1893: 210. [Albatross stns 3387 (off Golfo de Panamá, 7°40'00"N 79°17'50"W, 127 fms); 

3355 (off Punta Mariato, Panamá, 7°12'20"N 80°55'00"W, 182 fms] 

Sicyonia rectirostris De Man, 1907a: 141. [Siboga stn 139, Sanana bay, E coast of Sula Besi, Indonesia, 22 m] 

Sicyonia robusta Crosnier, 2003: 323; Figs 99-102, 110C, D. [CORAIL 2, stn DW 166, Îles Chesterfield, 

19°41,49'S 158°25,24'E, 56 m] 

Sicyonia rocroi Crosnier, 2003: 280; Figs 60-62. [Mutu One, Hatutaa, Îles Marquises, 7°51,7'S 140°30,6'W, 

416-460 m] 

Sicyonia rotunda Crosnier, 2003: 290; Figs 68-69, 108E-F. [SMIB 5, stn DW 84, Nouvelle-Calédonie, 

22°20,8'S 168°43,1'E, 290 m] 

Sicyonia stimpsoni Bouvier, 1905a: 748. [Blake stn 273, off Barbados, 13°03'05"N 59°36'18"W, 103 fms; lectotype 

designation by Holthuis, 1959a] 

Sicyonia taiwanensis Crosnier, 2003: 282; Figs 63-64, 108B. [au large de Tai-Chi, Taiwan] 

Sicyonia trispinosa De Man, 1907a: 142. [Siboga stn 37, Pulau Sailus ketjil, Paternoster Islands, Indonesia, 

27 m] 

Sicyonia truncata (Kubo, 1949a) 

= Eusicyonia truncata Kubo, 1949a: 456; Figs 8M, 48F, 77A, G, 79J, 156E, 158. [Kumanonada, off 

Owase, Mie Prefecture, Japan, 350 m] 

Sicyonia typica (Boeck, 1864) 

= Synhimantites typicus Boeck, 1864: 189. [Norge] 

= Palæmon carinatus Olivier, 1811: 667; nec Cancer carinatus Brünnich, 1768. [il se trouve sur les côtes 

de la Nouvelle-Hollande] 

= Sicyonia Edwardsii Miers, 1881a: 367. [nomen novum for Palæmon carinatus Olivier, 1811] 

Sicyonia vitulans (Kubo, 1949a) 

= Eusicyonia vitulans Kubo, 1949a: 448; Figs 8Q, 48G, 77F, L, 79I, 154, 156F. [Kumanonada, off Owase, 

Mie Prefecture, Japan, ca. 350 m] 

Sicyonia wheeleri Gurney, 1943: 1; Figs 1-13. [Bermuda; description of adult in Burkenroad, 1946 as Sicyonia 

wheeleri sp. nov.]


Family SoleNoCeRIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Cryptopenaeus de Freitas, 1979 

= Cryptopenaeus de Freitas, 1979 (type species Cryptopenaeus catherinae de Freitas, 1979, by original 


= Crassipenaeus Liu & Zhong, 1983 (type species Crassipenaeus sinensis Liu & Zhong, 1983, by original 

designation and monotypy, gender masculine) 

Cryptopenaeus brevirostris Hayashi in Baba, Hayashi & Toriyama, 1986: 39; Plate 1; fig. 17. [Bungo Strait, 

Japan, 360-380 m] 

Cryptopenaeus catherinae de Freitas, 1979: 125; Fig. 1A-I. [Off Cape Santa Maria, southern Mozambique, 

26°06'S 33°08'E, 350 m] 

Cryptopenaeus clevai Crosnier, 1985: 26; Figs 1a-b, 2a-e, 3a. [CORINDON IV stn IV 1 (03°28.3'N 128°24.8'E 

– 03°30.0'N 128°23.0'E), 400-300 m] 

Cryptopenaeus crosnieri Pérez Farfante & Kensley, 1985: 281; Figs 1-2. [R/V Kapala stn 78-05-07, NE of 

North Solitary Island, New South Wales, Australia, 29°47-49'S 153°41'E, 234 m] 

Cryptopenaeus sinensis (Liu & Zhong, 1983) 

= Crassipenaeus sinensis Liu & Zhong, 1983: 171; Fig. 1. [North-Eastern Guangdong near Shantou 

(Swatow), 261 m] 

Gordonella Tirmizi, 1960 

= Gordonella Tirmizi, 1960 (type species Gordonella polyarthra Tirmizi, 1960, by monotypy, gender 

feminine) 

Gordonella kensleyi Crosnier, 1988a: 597; Figs 2e, 12b, 15h-i, 16g. [BIOCAL stn CP 23, Nouvelle-Calédonie, 

22°46'N 166°20'E, 2040 m] 

Gordonella paravillosa Crosnier, 1988a: 589; Figs 2d, 3c, 12a, 13-14, 15a-e, 16a-f. [Cidaris I stn 18-1, côte est 

de l’Australie, 17°19.58'S 147°47.61'E, 1147-1132 m] 

Gordonella villosa (Alcock & Anderson, 1894) 

= Haliporus villosus Alcock & Anderson, 1894: 146. [Investigator stns 121 Laccadive Sea, 14°35'15"N, 

1140 fms; 127, Laccadive Sea, off the Island of Minnikoy, 1200 fms] 

= Gordonella polyarthra Tirmizi, 1960: 373, Figs 86-96. [John Murray Expedition stn 135, southern 

Arabian Sea, 4°47'42"N 72°35'36"E, 2727 m] 

Hadropenaeus Pérez Farfante, 1977b 

= Hadropenaeus Pérez Farfante, 1977b (type species Hymenopenaeus modestus Smith, 1885b, by original 


Hadropenaeus affinis (Bouvier, 1906d) 

= Haliporus affinis Bouvier, 1906d: 4. [capturés par le Talisman aux îles du Cap Vert, sur des fonds de 

100 et de 410 mètres; un spécimen provient des Barbades où il fut pris par le Blake à 82 brasses de 

profondeur." = Talisman stn 116, off Cape Verde Islands, 16°53'N 25°10'W, 410-460 m; uncited Talisman 

stn, off Cape Verde Islands, ”100 m”; Blake stn 273, off Barbados, 13°03'05"N 59°36'18"W, 188 m 

according to Pérez Farfante, 1977b] 

Hadropenaeus lucasii (Spence Bate, 1881) 

= Solenocera Lucasii Spence Bate, 1881: 185. [south of New Guinea, about 130 fms; according to 

Spence Bate, 1888 (as Philonicus lucasii): Challenger stn 192, off the Ki Islands, south of Papua, 

5°49'15"S 132°14'15"E, 140 fms] 

= Haliporus malhaensis Borradaile, 1910: 258; Plate 16, fig. 2. [off Saya de Malha, Indian Ocean, 145 

fms] 

Hadropenaeus modestus (Smith, 1885b) 

= Hymenopenaeus modestus Smith, 1885b: 183. [Fish Hawk stn 1047, off Bethany Beach, Delaware, 

38°31'N 73°21'W, 156 fms] 

Hadropenaeus spinicaudatus Liu & Zhong, 1983: 174; Fig. 2. [Eastern Guangdong, off shore (19°59.9'N 

113°24.0'E), 140 m]


Haliporoides Stebbing, 1914b 

= Haliporoides Stebbing, 1914b (type species Haliporoides triarthrus Stebbing, 1914b, by monotypy, 


= Parahaliporus Kubo, 1949a (type species Haliporus sibogae De Man, 1907a, by original designation, 


Haliporoides cristatus Kensley, Tranter & Griffin, 1987: 265; Figs 1, 2, 5G-L. [NE of Point Danger, Queensland, 

Australia] 

Haliporoides diomedeae (Faxon, 1893) 

= Peneopsis diomedeae Faxon, 1893: 212. [Albatross stns 3384 (off Golfo de Panamá, 7°31'30"N 

79°14'00"W, 458 fms); 3395 (SW of Golfo de Panamá, 7°30'36"N 78°39'00"W, 730 fms); 3394 (SE of 

Golfo de Panamá, 7°21'N 79°35'W, 511 fms); 3393 (off Punta Mala, Panama, 7°15'N 79°36'W, 1020 

fms); 3353 (off Punta Mariato, Panama, 7°06'15"N 80°34'00"W, 695 fms); 3358 (S of Península de 

Azuero, 6°30'N 81°44'W, 555 fms)] 

Haliporoides sibogae australiensis Kensley, Tranter & Griffin, 1987: 269; Figs 3, 4, 5A-F. [E of Broken Bay, 

New South Wales, Australia] 

Haliporoides sibogae madagascariensis Crosnier, 1978: 102; Figs 36b, 37a-e, 38a-c, 39b, 40b, 41b, 42b. [Vauban 

CH 5, Madagascar, 12°44.8'S 48°10.6'E, 570-563 m] 

Haliporoides sibogae sibogae (De Man, 1907a) 

= Haliporus sibogae De Man, 1907a: 138. [Siboga stns 38, 7°35.4'S 117°28.6'E, 521 m; 74, 5°3.5'S 119°0'E, 

450 m; 212, 5°54.5'S 120°19.2'E, 462 m; 256, 5°26.6'S 132°32.5'E, 397 m] 

Haliporoides triarthrus triarthrus Stebbing, 1914b 

= Haliporoides triarthrus Stebbing, 1914b: 21. [East London NW. 1/2N., 18 miles, (Cape Colony), 250- 

300 fms] 

Haliporoides triarthrus vniroi (Crosnier, 1978) 

= Hymenopenaeus triarthrus vniroi Crosnier, 1978: 103; Figs 36c, 38e. [Van Gogh stn 264, Mozambique, 

25°28'S 33°31'E, 410 m] 

Haliporus Spence Bate, 1881 

= Haliporus Spence Bate, 1881 (type species Haliporus curvirostris Spence Bate, 1881, designated by 


Haliporus curvirostris Spence Bate, 1881: 185. [mid Pacific, 2375 fms] 

Haliporus taprobanensis Alcock & Anderson, 1899: 280. [Investigator stn 219, Gulf of Manaar and Cape 

Comorin, southern India, 550 fms)] 

= Hymenopenaeus kannemeyeri Kensley, 1977: 16, 27; Fig. 7. [Meiring Naude stn 38, off South Africa, 

28°21.9'S 32°34.6'E, 775-825 m] 

Haliporus thetis Faxon, 1893: 214. [Albatross stn 3413 (off the Galapagos Islands, 2°34'00"N 92°06'00"W, 

1360 fms)] 

Hymenopenaeus Smith, 1882 

= Hymenopenaeus Smith, 1882 (type species Hymenopenaeus debilis Smith, 1882, by monotypy, gender 

masculine) 

Hymenopenaeus aphoticus Burkenroad, 1936a: 112; Figs 62, 65-67. [Pawnee stn 54, Turks Island Passage, 

21°15'40"N 71°17'06"W, 900-945 fms] 

Hymenopenaeus chacei Crosnier & Forest, 1969: 545; Figs 1, 2a-b. [Ombango stn 406, au large du plateau 

continental du Gabon, 8°35'S 12°51'E, 545-555 m] 

Hymenopenaeus debilis Smith, 1882: 91; Plate 15, figs 6-11; Plate 16, figs 1-3. [Blake stns 317, SE of Savannah 

Beach, Georgia, 31°57'00"N 78°18'35"W, 333 fms; 323, SE of Cape Fear, North Carolina, 33°19'00"N 

76°12'30"W, 457 fms; 326, E of Cape Fear, North Carolina, 33°42'15"N 76°00'50"W, 464 fms] 

= Hymenopenaeus debilis var. africanus Bouvier, 1908a: 83. [Type locality not indicated; according to 

Crosnier & Forest, 1973: Talisman stn 21, 33°46'N 9°02'W, 1319 m] 

Hymenopenaeus doris (Faxon, 1893) 

= Haliporus doris Faxon, 1893: 214. [Albatross stns 3414 (off Cabo Velas, Costa Rica, 10°14'N 96°28'W, 

2232 fms); 3415 (S of Punta Maldonado, Guerrero, Mexico, 14°46'N 98°40'W, 1879 fms)]


Hymenopenaeus equalis (Spence Bate, 1888) 

= Haliporus equalis Spence Bate, 1888: 285; Plate 41, fig. 1. [Challenger stn 200, 6°47'N 122°28'E, between 

the Philippine Islands and Borneo, 250 fms] 

Hymenopenaeus fallax Crosnier & Dall, 2004: 10; Figs 6-8. [Albatross stn 4106, Kaiwi Channel, 335-350 fms] 

Hymenopenaeus fattahi Ramadan, 1938: 60; Fig. 8. [John Murray Expedition stn 54 (21°58'00''N 62°19'42''E 

to 21°58'36''N 62°21'24''E), South Arabian coast, 1046 m] 

Hymenopenaeus furici Crosnier, 1978: 127; Figs 39f, 40f, 42f, 43d, 46f-h. [Vauban CH 133, Madagascar, 

13°02'S 48°02'E, 1000-1525 m] 

Hymenopenaeus halli Bruce, 1966a: 216; Figs 1-2. [Cape St. Mary Cruise 1/64, stn 26, South China Sea, 

19°22.5'N 114°07.5'E to 19°22.0'N 114°11.0'E, 400-435 fms] 

Hymenopenaeus laevis (Spence Bate, 1881) 

= Haliporus laevis Spence Bate, 1881: 185. [mid Atlantic, 2500 fms] 

= Hymenopenaeus microps Smith, 1884: 413; Plate 10, fig. 1. [Albatross stns 2076, E of Georges Bank, Massachusets, 

41°13'00"N 60°00'50"W, 906 fms; 2037, off New Jersey, 38°50'00"N 69°23'30"W, 1731 fms] 

= Haliporus androgynus Bouvier, 1906e: 253. [Talisman stns 101, entre Dakar et la Praya, 16°38'N 

20°44'W, 3200 m; 102, entre Dakar et la Praya, 15°48'N 20°23'W, 3655 m] 

Hymenopenaeus methalli Crosnier & Dall, 2004: 17; Figs 11-13, 14B, C. [MUSORSTOM 5, stn 384, 19°42.40'S 

158°50.80'E, 756-772 m] 

Hymenopenaeus neptunus (Spence Bate, 1881) 

= Haliporus neptunus Spence Bate, 1881: 185. [among the Celebes Islands, about 600 fms; according 

to Spence Bate, 1888: Challenger stns 191, off the Arrou Islands, 5°41'S 134°4'30"E, 800 fms; 196, near 

the Philippines, 0°48'30"S 126°58'30"E, 825 fms] 

Hymenopenaeus nereus (Faxon, 1893) 

= Haliporus nereus Faxon, 1893: 213. [Albatross stns 3366 (S of Cabo Blanco, Costa Rica, 5°30'N 

86°45'W, 1067 fms); 3353 (S of Morro de Puercos, Panama, 7°06'15"N 80°34'00"W, 695 fms); 3382 (off 

Pen de Azuero, Panama, 6°21'N 80°41'W, 1793 fms); 3413 (NW of Galapagos Islands, Ecuador, 

2°34'N 92°06'W, 1360 fms); 3398 (NW of Punta Galera, Ecuador, 1°07'N 80°21'W, 1573 fms); 3399 

(NW of Punta Galera, Ecuador, 1°07'N 81°04'W, 1740 fms); 3400 (E of Galapagos Islands, Ecuador, 

00°36'N 86°46'W, 1322 fms); 3407 (Galapagos Islands, Ecuador, 00°04'00"S 90°24'30"W, 885 fms)] 

Hymenopenaeus obliquirostris (Spence Bate, 1881) 

= Haliporus obliquirostris Spence Bate, 1881: 186. [off Kermadec Island; according to Spence Bate, 

1888: Challenger stn 170, off the Kermadec Islands, 29°55'S 178°14'W, 520 fms] 

Hymenopenaeus propinquus (De Man, 1907a) 

= Haliporus propinquus De Man, 1907a: 140. [Siboga stns 80, Pulu Kaniungan Ketjil, Indonesia, 11 m; 

178, 2°40'S 128°37.5'E, 835 m; 316, 7°19.4'S 116°49.5'E, 538 m] 

Hymenopenaeus sewelli Ramadan, 1938: 58; Figs 7a-e. [John Murray Expedition stn 156 (4°44'30''N 

72°46'00''E to 4°41'12''N 72°42'48''E), Maldive area, 1828 m] 

Hymenopenaeus tuerkayi Crosnier, 1995: 189; Figs 1-2. [Golfe d’Aden central, 12°56,7'N 47°47,0'E-12°55,9'N 

47°47,4'E, 2276-2282 m] 

Mesopenaeus Pérez Farfante, 1977b 

= Mesopenaeus Pérez Farfante, 1977b (type species Parartemesia tropicalis Bouvier, 1905a, by original 


Mesopenaeus brucei Crosnier, 1986b: 20; Figs 1a-f, 2a-b. [Soela Cruise 184 stn 55, Australie, 17°41.2'S 

118°42.5'E, 354-360 m] 

Mesopenaeus mariae Pérez Farfante & Ivanov, 1982: 303; Figs 1-7. [Professor Mesystsev haul 135, SE of Saya 

de Malha Bank, Indian Ocean, 11°27'36"S 61°37'36"E, 158-160 m] 

Mesopenaeus tropicalis (Bouvier, 1905a) 

= Parartemesia tropicalis Bouvier, 1905a: 748. [de la mer des Antilles où elle a été prise par le Blake par 

des profondeurs de 80 à 175 brasses] 

= Solenocera weymouthi Lindner & Anderson, 1941: 181; Fig. 1a-e. [Pelican stn 137, 29°28'N 87°30'W, 

46 fms]


Pleoticus Spence Bate, 1888 

= Philonicus Spence Bate, 1888 (type species Philonicus mülleri Spence Bate, 1888, designated by 


= Pleoticus Spence Bate, 1888 (nomen novum for Philonicus Spence Bate, 1888, a junior homonym of 

Philonicus Loew, 1849 (Diptera)) 

= Parartemesia Bouvier, 1905a (type species Parartemesia carinata Bouvier, 1905a, designated by Fowler, 

1912, gender feminine) 

= Faxonia Bouvier, 1905b (type species Penaeopsis [sic] ocularis Faxon, 1895, designated by Fowler, 


Pleoticus muelleri (Spence Bate, 1888) 

= Philonicus mülleri Spence Bate, 1888: 275; Plate 39, figs 1-2. [Challenger stn 321, off Montevideo, 

Uruguay, 35°02'N 55°15'W, 13 fms] 

= Parartemesia carinata Bouvier, 1905a: 748. [capturée par le Hassler..” ”..au large de Montevideo, par 

7 et 44 brasses de profondeur.”; off mouth Río de la Plata, 35°42'S 56°20'W, 44 fms (80 m); see Pérez 

Farfante, 1977b] 

Pleoticus robustus (Smith, 1885b) 

= Hymenopenaeus robustus Smith, 1885b: 180. [Albatross stn 2125, S of Curaçao, 11°43'00"N 69°09'30"W, 

208 fms] 

= Peneopsis ocularis Faxon, 1895: 187. [I have examined with some care a specimen in this Museum 

[Museum of Comparative Zoology, Harvard] from the ”Blake” collection, labelled ”Peneopsis ocularis” 

by A. Milne-Edwards] 

Pleoticus steindachneri (Balss, 1914c) 

= Haliporus Steindachneri Balss, 1914c: 135. [Pola stns 9, 23°21'N 37°37'E, 791 m; 20, 23°20'N 36°20'E, 

780 m; 47, 23°41'N 38°9'E, 610 m; 59, 25°43'N 36°10'E, 780 m; 61, 24°35'N 36°51'E, 828 m; 66, 26°8'N 

35°27'E, 1168 m; 107, 20°27'N 38°18'51"E, 748 m; 109, 21°19'N 37°39'E, 890 m; 110, 21°7'N 37°28'E, 

635 m; 114, 19°38'N 37°55'E, 535 m; 117, 20°16'N 37°33'E, 638 m; 120, 19°21'N 38°29'E, 560 m; 121, 

18°51'N 39°5'E, 690 m; 143, 17°7'N 39°55'E, 212 m; 156, 22°51'N 38°2'E, 712 m; 170, 27°2'N 35°17'E, 

690 m; 178, 26°19'N 34°24'E, 720 m; full description and list of syntypes in Balss, 1915] 

Solenocera lucas, 1849 

= Solenocera Lucas, 1849 (type species Solenocera Philippii Lucas, 1849, by monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in Opinion 611 in 1961) 

= Parasolenocera Wood-Mason in Wood-Mason & Alcock, 1891a (type species Parasolenocera annectens 

Wood-Mason in Wood-Mason & Alcock, 1891a, by monotypy, gender feminine) 

= Transolenocera Burkenroad, 1934b (type species Solenocera (Parasolenocera) maldivensis Borradaile, 

1910, by monotypy, gender feminine) 

Solenocera acuminata Pérez Farfante & Bullis, 1973: 8; Figs 1c, 2, 4-6, 18, 19. [Oregon stn 2022, N of Rivière 

Organabo, French Guiana, 07°15'N 53°35'W, 210 m] 

Solenocera africana Stebbing, 1917b: 32; Plate 93A. [Sebastian Bluff NW. 3/4W., 8 miles, South Africa, 34 

fms] 

? = Solenocera membranacea capensis Heegaard, 1966: 32; Figs 15-21, 23. [Discovery stns 100B, 33°20'S- 

33°46'S 15°08'E – 15°18'E, 5-0 m; 102, 25°29'30"S 18°33'40"E, 50-0 m; 260, 33°06'30"S 17°45'15"E, 100-0 

m; 277, 1°44'00"S 8°38'00"E, 63 m; 89, 34°05'15"S 16°00'45"E, 50-0 m; 99A, 33°20'00"S 17°17'00"E, 5-0 

m; 99E, 33°11'00"S 17°26'00"E, 5-0 m; 276, 5°54'00"S 11°19'00"E, 150 m] 

Solenocera agassizii Faxon, 1893: 211. [Albatross stns 3389 (off Panama, 7°16'45"N 79°56'30"W, 210 fms); 

3391 (off Panama, 7°33'40"N 79°43'20"W, 153 fms) 

Solenocera alfonso Pérez Farfante, 1981: 631; Figs 1-5. [off Capitancillo Island, W of Leyte, Philippines] 

= Solenocera alfonso forma inermis Crosnier, 1989: 56; Figs 3e-f, 5f-h, 6a, e. [many locations in the 

Philippines, 175-200 m; unavailable under Art. 15.2] 

Solenocera algoensis Barnard, 1947 

= Solenocera algoense Barnard, 1947: 383. [E portion of Algoa Bay, South Africa, 50 fms] 

= Solenocera ramadani Ivanov & Hassan, 1976c: 242; Figs 1-2. [Van Gogh stn 269, off E Africa, 29°31'S 

34°42'E, 285-305 m]


Solenocera alticarinata Kubo, 1949a: 227; Figs 8W, 45E, 72P, V, 80F, 93, 94A-C, 100. [Tainan, Formosa] 

Solenocera annectens (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Parasolenocera annectens Wood-Mason in Wood-Mason & Alcock, 1891a: 276. [Investigator stn 116 

(Andaman Sea, 11°25'5"N 92°47'6"E), 405 fms] 

Solenocera atlantidis Burkenroad, 1939: 10; Figs 5-10. [Atlantic stn 2813, off Mobile Bay, Alabama, 29°45'N 

88°11'W, 19 fms] 

Solenocera australiana Pérez Farfante & Grey, 1980: 422; Figs 1-7. [Apache cruise 21, haul 3, N of Groote 

Eylandt, Gulf of Carpentaria, Northen Territory, Australia, 13°34'S 136°30'E, 22 m] 

Solenocera barunajaya Crosnier, 1994c: 355; Figs 1a-c, 2a, 3a-c. [KARUBAR stn CP 83, Iles Tanimbar Island, 

Indonésie, 9°23'S 131°00'E, 285-297 m] 

Solenocera bedokensis Hall, 1962: 13; Figs 78-78c. [Off Bedok Village, Singapore, 17 fms] 

Solenocera bifurcata Dall, 1999: 574; Fig. 14A-F. [Off Cape Moreton, SE Queensland, Australia] 

Solenocera burukovskyi Timofeev, 1993: 37; Fig. 1. [Gulf of Aden, 12°19'4N 44°21'7E, 470-475 m] 

Solenocera choprai Nataraj, 1945: 91, Figs 1-4. [Arabian Sea, 17°27'N 71°41'E, 56-58 fms] 

Solenocera comata Stebbing, 1915 

= Solenocera comatum Stebbing, 1915: 67; Plates 13-14. [Off East London, E coast of South Africa] 

= Solenocera novae-zealandiae Borradaile, 1916: 79; Fig. 1. [7 miles E of North Cape, off New Zealand, 

70 fms] 

= Solenocera brevipes Kubo, 1949a: 246; Figs 1S, 8X, 20N, 27F-H, 45D, 66I-J, 72Q, W, 80A, 98H-J, 99, 

100. [Komanonada, off Owase, Mie Prefecture, Japan, about 300 m] 

Solenocera crassicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

? = Penaeus planicornis Fabricius, 1798: 409. [in Oceano Indico] 

= Penaeus crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [côtes de l’Inde] 

= Solenocera sinensis Yü, 1937: 112; Figs 1-5. [Amoy, Tinghai] 

= Solenocera indicus Nataraj, 1945: 94; Figs 5-8. [Cocanada Bay, Bay of Bengal] 

= Solenocera subnuda Kubo, 1949a: 255; Figs 8T, 20O, 27I-J, 45C, 66C, D, 72M, S, 80D, 100, 102D-G, 

103. [S coast of Borneo, Indonesia] 

= Solenocera kuboi Hall, 1956: 69; Plate 9, Figs 2-3. [Off Tanjong Stapa, Malaysia, 6-7 fms] 

Solenocera faxoni De Man, 1907a: 136. [Siboga stn 254, 5°40'S 132°26'E, 310 m] 

Solenocera florea Burkenroad, 1938: 64; Text-Figs 6-7. [Pearl Island, Gulf of Panama, 8°29'40"N 78°52'30"W, 

19-24 fms] 

Solenocera geijskesi Holthuis, 1959a: 56; Figs 4-5. [Surinam, 20 miles off the coast, between mouths of 

Nickerie and Coppename Rivers, 27 m] 

Solenocera gurjanovae Starobogatov, 1972: 365; Plate 3, fig. 13. [Gulf of Tonkin, Vietnam, 50 m] 

Solenocera halli Starobogatov, 1972: 412; Plate 2, fig. 7. [Andaman Sea, off India, 6°05.8' – 6°02'N 98°59' – 

99°00.5'E] 

Solenocera hextii Wood-Mason & Alcock, 1891b: 188. [Investigator stns 81 (24 miles SE of Gapalpur, Ganjam 

Coast, 89-93 fms); 96 (Bay of Bengal, 18°30'N 84°46'E, 98-102 fms), 89-102 fms; off Chittagon, 65 

fms; off the Mahánadi Delta, 68 fms; off the Godávari Delta, 70 fms] 

Solenocera koelbeli De Man, 1911a: 45, 48, 50, 51. [Japan] 

= Solenocera depressa Kubo, 1949a: 237; Figs 8R, 27T-V, 45B, 66E-F, 72O, U, 80G, 96, 98A-D, 100. 

[Komanonada, off Owase, Mie Prefecture, Japan] 

= Solenocera vietnamensis Starobogatov, 1972: 363, 384; Plate 2, fig. 6a-c. [Pelamida, stn 39, Tonkin 

Gulf, Vietnam] 

Solenocera maldivensis Borradaile, 1910 

= Solenocera (Parasolenocera) maldivensis Borradaile, 1910: 258; Plate 16, fig. 1. [Funadu Velu, Miladumadulu 

Atoll, Maldive Islands] 

Solenocera mascarensis Burukovsky, 1993a: 23; Fig. 1.3-1.7. [Sala-de-Malya Bank, 11°11'1''S 62°37'E, 2020- 

2000 m] 

Solenocera melantho De Man, 1907a: 137. [Siboga stns 302, 10°17.9'S 123°28.7'E, 216 m; 306, 8°27'S 

122°54.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

= Solenocera prominentis Kubo, 1949a: 231; Figs 8V, 14C, 16C, 20M, 27Q-S, 45A, 66G, H, 72F, L, 80E, 

94D-G, 95, 100. [Komanonada, off Owase, Mie Prefecture, Japan, about 350 m]


Solenocera membranacea (Risso, 1816) 

= Peneus membranaceus Risso, 1816: 98. [Président Théodore Tissier stn J.376, France, Ligurian Sea between 

Corsica and Nice, 42°56.5'N 9°37.2'E, 310-415 m; neotype designation by Holthuis, 1961b] 

= Peneus siphonoceros Philippi, 1840: 190; Plate 4, fig. 3. [Neapel] 

= Solenocera philippii Lucas, 1849: 300. [baie d’Alger] 

= Penaeus distinctus De Haan, 1849 [in De Haan, 1833-1850]: 194. [Japan; erroneous locality, see 

Yamaguchi & Baba, 1993] 

Solenocera moosai Crosnier, 1985: 37; Figs 5a, 6a, 7c,d, h, i. [CORINDON II stn 273 (01°56.0'S 119°16.0'E), 

120-200 m] 

Solenocera mutator Burkenroad, 1938: 61; Text-Figs 2-5. [Templeton Crocker Expedition stn 150, from Gorda 

Banks, tip of Lower California, 23°01'30"N 109°30"W, 40-100 fms] 

Solenocera necopina Burkenroad, 1939: 7; Figs 1-4. [Atlantis stn 2377, off Mobile Bay, Alabama, 29°16'N 

87°54'W, 125 fms] 

Solenocera pectinata (Spence Bate, 1888) 

= Philonicus pectinatus Spence Bate, 1888: 279; Plate 38. [Challenger stn 188, Arafura Sea, south of 

Papua, 9°59'S 139°42'E, 28 fms] 

= Philonicus cervicalis Zehntner, 1894: 210; Plate 9, fig. 26. [Amboine] 

Solenocera pectinulata Kubo, 1949a: 251; Figs 8S, 27A-B, 66 K-L, 72N-T, 83B, 101, 102A-C. [Komanonada, 

off Owase, Mie Prefecture, Japan, about 350 m] 

? = Solenocera utinomii Kubo, 1951: 263; Fig. 4. [Off Kii Peninsula, Japan, ca. 150 m] 

Solenocera phuongi Starobogatov, 1972: 366; Plate 3, fig. 12a-b. [Pelamida stn 9, Tonkin Gulf, 113 m] 

Solenocera rathbuni Ramadan, 1938: 57 (partim, nec Fig. 6 = S. algoensis Barnard, 1947). [Albatross stn 3987, 

Kauai Island, Hawai’i, 50-55 fms; lectotype designation by Crosnier, 1978] (Fig. 5) 

Solenocera spinajugo Hall, 1961: 81; Plate 17, figs 1-3. [N of the Malacca Strait, 120 miles NW of Penang, 

41 fms] 

Solenocera vioscai Burkenroad, 1934b: 65; Figs 1-4. [About 5 miles off Pass à la L’outre, Louisiana, USA, 15 

fms] 

Solenocera waltairensis M.J. George & Muthu, 1970: 292; Figs 1-4. [Waltair, east coast of India, 17°43'N 

83°23'E, 20 m] 

Solenocera zarenkovi Starobogatov, 1972: 367, Plate 3, fig. 14a-b. [Orlik, stn 18, Tonkin Gulf] 

Fig. 5. Solenocera rathbuni Ramadan, 1938. Photo by Tin-Yam Chan.


Superfamily SeRGeSToIDeA Dana, 1852a 

Family luCIFeRIDAe De Haan, 1849 [in De Haan, 1833-1850] 

Lucifer J.V. Thompson, 1829 (Fig. 6) 

= Lucifer J.V. Thompson, 1829 (type species Leucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 

1834-1840], by subsequent indication under Article 68b in H. Milne Edwards, 1834-1840, 


= Leucifer H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] [incorrect subsequent spelling 

of Lucifer] 

Lucifer chacei Bowman, 1967: 266; Figs 1-4. [Eniwetok Atoll, Marshall Islands, lagoon 4.8 km W of Parry 

Island, 3 m] 

Lucifer faxoni Borradaile, 1915a: 228, 230. [off Chesapeake Bay, U.S.A.; lectotype designation by Holthuis, 

1959a] 

= Lucifer affinis Borradaile, 1915a: 228, 230. [Floridastrom, J.N., 56, N.E. of Bermuda, roughly 35°N 

60°W; lectotype designation by Holthuis, 1959a] 

Lucifer hanseni Nobili, 1905a 

= Lucifer Hanseni Nobili, 1905a: 395. [Djibouti] 

= Lucifer inermis Borradaile, 1915a: 229. [Melbourne Harbour, Australia] 

Lucifer intermedius Hansen, 1919: 57; Plate 4, figs 8a-b; Plate 5, figs 1a-g. [Siboga stns 7, 7°55'.5S 114°26'E, 

15 m and more; 35, 8°0'.3S 116°59'E, 1310 m, surface; 37, Sailus ketjil, Paternoster-islands, 27 m and 

less, surface; 66, bank between islands of Bahuluwang and Tambolungan, south of Saleyer, 8-10 m; 

75, 4°57'.4S 119°2'.8E, from 11 m to surface; 81, Pulu Sebangkatan, Borneo-bank, 34 m; 106, Anchorage 

of Kapul-island, Sulu-archipelago, 13 m; 109, Anchorage off Pulu Tongkil, Sulu-archipelago, 13 

m; 117a, 1°15'N 123°37'E; ?132, about 5°56'.7N 126°25'E; 136, Ternate anchorage, 23 m - surface; 138, 

Anchorage on the east coast of Kajoa-island, 66 m - surface; 141, 1°0'.4S 127°25'.3E, from 1500 m; 

144, Anchorage North of Salomakice-(Damar)island, 45 m; 157, 0°32'.9S 130°14'.6E; 177a, 2°30'S 

129°28'E; 189a, 2°22'S 126°46'E] 

Lucifer orientalis Hansen, 1919: 55; Plate 4, figs 7a-g. [Siboga stns 37, Sailus Ketjil, Paternoster-islands, 27 

m and less; 66, bank between islands of Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81, 

Pulu Sebangkatan, Borneo-bank, 34 m; 104, Sulu-harbour, Sulu-island, 14 m; 106, anchorage off 

Fig. 6. Lucifer spec. Photo by Arthur Anker.


Kapul-island, Sulu-archipelago, 13 m; 133, anchorage off Lirung, Salibabu-island, depth up to 36 m; 

136, Ternate anchorage, 23 m - surface; 138, anchorage on the E coast of Kajoa-island, 66 m - surface; 

140, Bay of Batjan, 13 m; 141, 1°0'.4S 127°25'.3E, 1500 m - surface; 143, 1°4'.5S, 127°52'.6E, 1000 m - 

surface; 148, 0°17'.6S 129°14'.5E, 1000 m - surface; 157, 0°32'.9S 130°14'.6E, 45 m; 177a, 2°30'S 

129°28'E; 189a, 2°22'S 126°46'E; 194-197, 1°53'.5-1°45'.3S 126°39-127°8'.3E; 203, 3°32'.5S 124°15'.5E, 

1500 m - surface; 220, anchorage off Pasir Pandjang, W coast of Binongka, 278 m - surface; 225, N. 

279°E, from Southpoint of South-Lucipara-island, 894 m; 243.4°30'.2S 129°25'E, 1000 m - surface; 

245.4°16'.5S 130°15'.8E, surface; 276, 6°47'.5S 128°40'.5 E, 750 m - surface; near stn 300, between 

Timor and Lomblen] 

Lucifer penicillifer Hansen, 1919: 59; Plate 5, figs 2a-k. [Siboga stns 7, 7°55'.5S 114°26'E, 25 m and more; 16, 

Bay of Kankamaräan, S coast of Kangeang, 22 m; 35, 8°0'.3S 116°59'E, surface; 37, Sailus ketjil, Paternoster-islands, 

27 m and less; 40, anchorage off Pulu Kawassang, Paternoster-islands, 12 m; 66, bank 

between Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81, Pulu Sebangkatan, Borneo-bank, 

34 m; 93, Pulu Sanguisiapo, Tawi-Tawi-islands, Sulu-archipelago, 12 m; 96, SE side of Pearl-bank, 

Sulu-archipelago; 98-99, 6°9'-6°7'.5N 120°21'-120°26'E; 99, 6°7'.5N 120°26'E, 16-23 m - surface; 105, 

6°8'N 121°19'E, 275 m; 106, anchorage off Kapul-island, Sulu-archipelago, 13 m; 107, 6°1'.5N. 

121°28'E; 109, anchorage off Pulu Tongkil, Sulu-archipelago, 13 m; 112, 3°1'N 122°2'E; 117 a , 1°15'N 

123°37'E; 125, anchorage off Sawan, Siau-island, 27 m; 128, 4°27'N 125°25'.7E, from 700 m to surface; 

136, Ternate anchorage, 23 m; 138, anchorage on the E coast of Kajoa-island, 66 m; 140, Bay of Batjan, 

13 m; 141, 1°0'.4S 127°25'.3E, from 1500 m to surface; 143, 1°4'.5S 127°52'.6E, from 1000 m to 

surface; 144, anchorage north of Salomakiëe- (Damar-) island, 45 m; 146, 0°36'S 128°32'.7E, 512 m; 

157, 0°32'.9S 130°14'.6E, 45 m; 165, anchorage on NE side of Daram-island, E coast of Misool, 49 m; 

168, anchorage N of Sabuda-island, 63 m; 174, Waru-bay, N coast of Ceram, 18 m; 177 a , 2°30'S 

129°28'E; 184, anchorage off Kampong Kelang, S coast of Manipa-island, 36 m; 185, 3°20' 127°22'.9E, 

from 1536 m to surface; 189 a , 2°22'S 126°46'E; 194, 1°53' 5S 126°39'E; 194-197, 1°53'.5-1°4S'.3S 126°39'- 

127°8'.3E; 203, 3°32'5S 124°15'.5E, from 1500 m to surface; 206, Buton-strait, surface; 220, anchorage 

off Pasir Pandjang, W coast of Binongka, 278 m; 225, 5700 m N 279°E from Southpoint of South- 

Lucipara-island, 894; 243, 4°30'.2S 129°25'E, from 1000 m to surface; 245, 4°16'.5S 130°15'.8E; near stn 

300, between Timor and Lomblen; 315, anchorage E of Sailus Besar, Paternoster-islands, surface] 

Lucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= Leucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [Type locality not 

indicated] 

= Leucifer Reynaudii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [trouvé dans 

l’Ocean indien] 

= Lucifer acestra Dana, 1852b: 671. [Pacific Ocean, 6°30'S 177°E, near Sherson’s Island] 

= Lucifer acicularis Dana, 1852b: 674. [Harbour of Rio de Janeiro] 

= Lucifer Zybrantsii Philippi, 1857: 323. [im Atlantischen Ocean unter 25°N.B. und 22°50'W.L. von 

Greenwich] 

= Leucifer uracanthus Steindachner, 1861: 365. [E mari atlantico] 

= Lucifer bonitensis Borradaile, 1915a: 228, 230. [toutes les mers] 

= Lucifer clausi Borradaile, 1915a: 228, 230. [Messina] 

= Lucifer batei Borradaile, 1915a: 228, 230. [Plankton-Expedition, Floridastrom, J.N. 62, Bermuda; 

lectotype designation by Holthuis, 1959a] 

Family SeRGeSTIDAe Dana, 1852a 

Acetes H. Milne edwards, 1830 

= Acetes H. Milne Edwards, 1830 (type species Acetes indicus H. Milne Edwards, 1830, by original 


Acetes americanus americanus Ortmann, 1893: 39; Plate 2, fig. 2. [Plankton-Expedition der Humbolt-Stiftung, 

Mündung des Tocantins JN 239 (0.6°S 48.1°W, 0 m), 240 (1.6°S 49.2°W, 0-35 m), 241 (1.6°S 

49.2°W, 0-35 m), 243 (0.7°S 48.2°W, 0-13 m), Pl. 106 (0.7°S 48.2°W, 12 m) and 107 (1.6°S 49.2°W, 35 m)]


= Acetes brasiliensis Hansen, 1919: 45; Figs 1-7. [Brazil] 

= Acetes americanus limonensis Burkenroad, 1934a: 99; Figs 37-38. [Mouth of the Sweetwater River, 

Limon Bay, Canal Zone, Panama] 

Acetes americanus carolinae Hansen, 1933 

= Acetes carolinae Hansen, 1933: 31; Figs 1-8. [near the Sea Buoy off Beaufort Inlet, North Carolina] 

Acetes binghami Burkenroad, 1934a: 101; Figs 39-40. [Bella Vista Beach, Panama City] 

Acetes chinensis Hansen, 1919: 41; Plate 4, fig. 3a-b. [33°10'N 129°18'E, 40 fms; Formosa Channel, 25 fms] 

Acetes erythraeus Nobili, 1905a: 393; Fig. 1. [Mer Rouge et Périm; Djibouti, dans la cavité d’une Beroë; 

Abdelkader, près de Massaouah, et Massaouah] 

Acetes indicus H. Milne Edwards, 1830: 351; Plate 11, figs 1-9. [Le Gange] 

= Acetes spiniger Hansen, 1919: 35, 43; Plate 4, fig. 5. [Surabaya, Java, Indonesia; 10 miles off Rangoon; 

Indo-Chinese Sea] 

Acetes intermedius Omori, 1975: 40; Figs 14, 16, 17. [Off Tungkiang, Taiwan] 

Acetes japonicus Kishinouye, 1905: 167; 2 unnumbered Figs [the Bay of Ariake, the Bay of Kojima, off 

Nagasu in Buzen, Japan; off Mokpho, Korea] 

= Acetes dispar Hansen, 1919: 39; Plate 3, figs 5a-f; Plate 4, fig. 1a. [Cheribon, Java, Indonesia; Lem 

Ngob, Gulf of Siam] 

= Acetes cochinensis Rao, 1970: 298; Figs 1-10. [Inshore sea of Cochin, southwest coast of India, 3.5- 

11 m] 

Acetes johni Nataraj, 1949: 139; Fig. 1. [Travancore coast, India] 

Acetes marinus Omori, 1975: 49; Figs 8, 21, 25. [Rio Tocantins, near its junction with Rio Para, Brazil] 

Acetes natalensis Barnard, 1955: 2, 43. [Durban Bay, South Africa] 

Acetes paraguayensis Hansen, 1919: 46; Figs 8-14. [Lagoon at Rio Paraguay near its junction with Rio 

Parana, Paraguay, outlet of Riacho del Oro in Rio de la Plata, Paraguay] 

Acetes serrulatus (Krøyer, 1859) 

= Sergestes serrulatus Krøyer, 1859: 268; Plate 4, fig. 12. [nordligste Kattegat] 

= Acetes insularis Kemp, 1917a: 54; Figs 1f-g, 2c, 3c, 4c, 5b, e, 7c. [Mouth of Rajang R., Sarawak, Borneo] 

Acetes sibogae sibogae Hansen, 1919: 38; Plate 3, figs 4a-h. [Siboga stns 47, Bay of Bima, near south fort, 55 

m; 323, Sangkapura roads, Bawean Island, 12 m] 

= Acetes sibogalis Achuthankutty & M.J. George, 1973: 139; Figs 1-20. [Aroor region of the Cochin 

backwaters] 

= Acetes orientalis Achuthankutty & Ayyappan Nair, 1976: 233; Figs 1-19. [near fish jetty in Mandovi 

estuary, Arabian Sea, Goa, India] 

Acetes sibogae australis Colefax, 1940 

= Acetes australis Colefax, 1940: 341, 345; Figs 1-19. [Australia: Homebush Bay, Port Jackson (Sydney 

Harbour); Folly Point, Middle Harbour, Port Jackson; Clarence River, N coast of New South Wales; 

Tuggerah Lake, 50 miles N of Port Jackson] 

Acetes vulgaris Hansen, 1919: 35; Plate 3, figs 2a-r. [Siboga stns 4, anchorage off Djangkar (Java), 7°42'S 

114°12.6'E, 9 m; 47, Bay of Bima, near south fort, 55 m; 311, Sapeh Bay, E coast of Sumbawa, 0-36 m] 

Allosergestes Judkins & Kensley, 2008 

= Allosergestes Judkins & Kensley, 2008 (type species Sergestes sargassi Ortmann, 1893, by original 


Allosergestes index (Burkenroad, 1940) 

= Sergestes (Sergestes) index Burkenroad, 1940: 41. [Dana stn 3630 II (New Zealand, Auckland, 34°24'S 

178°42.5'E), 3000 m wire out)] 

Allosergestes nudus (Illig, 1914) 

= Sergestes nudus Illig, 1914: 366; Figs 26-28. [Atlantischen Ozean, 32°S 8°W, 1000 m] 

Allosergestes oleseni (Vereshchaka, 2009) comb. nov. 

= Sergestes oleseni Vereshchaka, 2009: 78; Figs 35, 38. [Dana stn 3602-2 (south western Pacific Ocean, 

between Fiji and New Caledonia, 20°00'S 174°29'E, 600 m wire out)]


Allosergestes pectinatus (Sund, 1920) 

= Sergestes pectinatus Sund, 1920: 24; Figs 42-43. [Michael Sars stns 23 (35°32'N 7°7'W, 1215 m); 34 

(28°52'N 14°16'W, 2170 m); 52 (31°24'N 34°47'W); 45 (28°42'N 20°0'W); 49 (29°6'N 25°2'W); 51 

(31°20'N 35°7'W, 3886 m); 53 (34°59'N 33°1'W, 2615-2865 m); 56 (36°53'N 29°17'W, 3239 m); 62 

(36°52'N 39°55'W); 64 (34°44'N 47°52'W); 67 (40°17'N 50°39'W)] 

Allosergestes pestafer (Burkenroad, 1937) 

= Sergestes pestafer Burkenroad, 1937: 318; Figs 1-3. [Templeton Crocker Expedition stn 165 T-3, 145 miles 

N of Clarion Island, Revillagigido Islands, Lower California, E Pacific, 20°36'N 115°07'W, 500 m] 

Allosergestes sargassi (Ortmann, 1893) 

= Sergestes sargassi Ortmann, 1893: 34; Plate 3, fig. 1. [Plankton-Expedition der Humbolt-Stiftung, 

Floridastrom JN 50 (39.4°N 57.8°W, 0-200 m); Sargasso-See JN 94 (31.5°N 45.6°W, 0-400 m), 99 

(31.7°N 43.6°W, 0-400 m), 102 (31.7°N 42.7°W, 0-400 m), 110 (30.3°N 37.9°W, 0-400 m), 113 (29.8°N 

36.8°W, 0-400 m), 117 (28.3°N 34.3°W, 0-400 m), 127 (24.6°N 31.0°W, 0-400 m), 262 (23.7°S 36.0°W, 0 

m), 263 (25.6°S 34.9°W, 0-400 m), Pl. 58 (25.1°N 31.5°W, 200 m); Nördl. Aequatorialstrom JN 135 

(18.9°N 26.4°W, 0-400 m), 260 (20.4°N 37.8°W, 0-400 m), Pl. 64 (16.1°N 23.1°W, 200 m); Südl. Aequatorialstrom 

Pl. 76 (0.1°S 15.2°W, 200 m)] 

Allosergestes verpus (Burkenroad, 1940) 

= Sergestes (Sergestes) verpus Burkenroad, 1940: 40. [Dana stn 3739 IX (Celebes Sea, 3°20'N 123°50'E), 

2000 m wire out); according to Vereshchaka, 2009, the holotype label indicates that the specimen 

was collected at Dana stn 3844-6 (Indo-West Pacific, 12°05'S 96°45'E, 600 m wire out)] 

Allosergestes vinogradovi (Vereshchaka, 2009) comb. nov. 

= Sergestes vinogradovi Vereshchaka, 2009: 90; Figs 35, 44. [Dana stn 3902-2, Indo-West Pacific, 6°05'N 

95°30'E, (600 m wire out)] 

Deosergestes Judkins & Kensley, 2008 

= Deosergestes Judkins & Kensley, 2008 (type species Sergestes curvatus Crosnier & Forest, 1973, by 


Deosergestes coalitus (Burkenroad, 1940) 

= Sergestes (Sergestes) coalitus Burkenroad, 1940: 39. [Dana stn 3737 I (Mindanao Sea, 7°23'N 121°29'E, 

1000 m wire out)] 

= Sergestes (Sergestes) erectus Burkenroad, 1940: 38. [Dana stn 3579 I (Rarotonga, 20°56'S 160°03'W), 


Deosergestes corniculum (Krøyer, 1855) 

= Sergestes corniculum Krøyer, 1855: 30. [fra Atlanterhavets tropiske Deel, 4.5°N 21.5°W; according to 

Krøyer, 1859] 

= Sergestes laciniatus Krøyer, 1859: 274, 282; Plate 5, fig. 15. [fra Kattegattet] 

? = Pasiphae rubroguttata Filhol, 1885b: Plate 5. [Type locality not indicated] 

= Sergestes longirostris Spence Bate, 1888: 415; Plate 75, fig. 3. [Mid Atlantic] 

= Sergestes (Sergestes) curvatus Crosnier & Forest, 1973: 315; Figs 105i-k, 107c-d, f-g. [au large de 

l’Afrique du Sud, 35°42'S 24°40'E, 500 m] 

Deosergestes disjunctus (Burkenroad, 1940) 

= Sergestes (Sergestes) disjunctus Burkenroad, 1940: 38. [Dana stn 3630 IV (New Zealand, Auckland, 

34°24'S 178°42.5'E), 1000 m wire out)] 

Deosergestes henseni (Ortmann, 1893) 

= Sergia henseni Ortmann, 1893: 38; Plate 3, fig. 3. (partim, see Crosnier & Forest, 1973). [Plankton- 

Expedition der Humboldt-Stiftung, Nördl. Aequatorialstrom JN 146 (12.3°N 22.3°W); Vertikalnetz 

0-400 m; Guineastrom JN 158 (7.5°N 21.3°W), circa 4000 m] 

Deosergestes paraseminudus (Crosnier & Forest, 1973) 

= Sergestes (Sergestes) paraseminudus Crosnier & Forest, 1973: 313; Figs 105d, 106c-d, f. [Ombango stn 

309-GS 20 bis, 1°55'S 8°30'E, 0-350 m] 

Deosergestes pediformis (Crosnier & Forest, 1973) 

= Sergestes (Sergestes) pediformis Crosnier & Forest, 1973: 313; Figs 105e-h, 107a-b, e. [Ombango stn 

302-GS 19, 4°47'S 10°42'E, 0-725 m]


Deosergestes rubroguttatus (Wood-Mason in Wood-Mason & Alcock, 1891c) 

= Sergestes rubroguttatus Wood-Mason in Wood-Mason & Alcock, 1891c: 354; Fig. 10. [Investigator 

stns 107 (Laccadive Sea, 8°23'N 75°47'E), 738 fms; 109 (S of C. Comorin, 7°41'N 78°21'E), 738 fms; 110 

(Bay of Bengal, 9°34'N 85°43'15''E), 1997 fms; 117 (Bay of Bengal, 11°58'N 88°52'17''E), 1748 fms] 

Deosergestes seminudus (Hansen, 1919) 

= Sergestes seminudus Hansen, 1919: 18; Plate 1, figs 7a-c; Plate 2, figs 1a-f. [Siboga stns 66, Bank between 

islands of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 144, Anchorage N 

of Salomakiëe (Damar-) Island, 45 m; 148, 0°17.6' S 129°14.5'E, 0-1000 m; 157, 0°32.9'S 130°14.6'E, 45 

m; 167, Anchorage N of Sabuda Island, 63 m; 177a, 2°30'S 129°28'E; 185, 3°20'S 127°22.9'E, Manipa 

Strait, 0-1536 m; 189a, 2°22'S 126°46'E; 194-197, 1°55' – 1°45.3'S 126°39'- 127°8.3'E; 203, 3°32.5'S 

124°15.5'E 0-1500 m; 223, 5°44.7'S 126°27.3'E, surface; 230, 3°58'S 128°20'E 0-2000 m; 276, 6°47.5'S 

128°40.5'E, 0-750 m] 

= Sergestes nipponensis Yokoya, 1933: 13; text-fig. 4. [Sôyô-Maru stns 21, Japan, SE of Siwoya-zaki, 209 

m; 25, Japan, E of Siwoya-zaki, 525 m; 253, Japan, Sagami Bay, 452 m] 

Eusergestes Judkins & Kensley, 2008 

= Eusergestes Judkins & Kensley, 2008 (type species Sergestes arcticus Krøyer, 1855, by original designation, 


Eusergestes antarcticus (Vereshchaka, 2009) comb. nov. 

= Sergestes antarcticus Vereshchaka, 2009: 57; Figs 25, 26. [Dana stn 3975-2 (South Atlantic, Cape 

Town, 35°42'S 18°37'E, 2500 m wire out)] 

Eusergestes arcticus (Krøyer, 1855) 

= Nika Sinuolata Risso, 1816: 87. [Nice; status of name discussed by Holthuis, 1977a] 

= Sergestes arcticus Krøyer, 1855: 27. [fra Grönland, according to Krøyer, 1859] 

= Sergestes Rinkii Krøyer, 1855: 33. [Atlanterhav på 58-59° n. Br., according to Krøyer, 1859] 

= Sergestes Meyeri Metzger, 1875: 302; Plate 6, fig. 7. [Kiel] 

= Sergestes magnificus Chun, 1888: 33; Plate 4, figs 4-5. [Ischia, 800 m] 

Eusergestes similis (Hansen, 1903) 

= Sergestes similis Hansen, 1903: 60; Plate 11, fig. 6. [Challenger stn 232, off Japan, 35°11'N 139°28'E, 

345 fms] 

? = Sergestes nasidentatus Spence Bate, 1888: 398; Plate 72, fig. 2. [Pacific Ocean between Valparaiso 

and Juan Fernandez, 0-200 fms; Holthuis (pers. comm.)] 

? = Sergestes laeviventralis Spence Bate, 1888: 425; Plate 67, fig. 3. [north of New Guinea] 

? = Sergestes longicaudatus Stimpson, 1860a: 46. [Oceano Pacifico, lat. Bor. 40°, long occ. 155°] 

? = Sergestes affinis Hansen, 1919: 7. [nomen nudum] 

Neosergestes Judkins & Kensley, 2008 

= Neosergestes Judkins & Kensley, 2008 (type species Sergestes edwardsii Krøyer, 1855, by original 


Neosergestes brevispinatus (Judkins, 1978) 

= Sergestes brevispinatus Judkins, 1978: 13; Figs 2d, 5d-f,h,k, 6-8, 216. [Eastern tropical Pacific, 11°02'S 

81°41'W, ca. 200 m at night] 

Neosergestes consobrinus (Milne, 1968) 

= Sergestes consobrinus Milne, 1968: 26; Figs 5-9. [Brown Bear cruise 199, haul 227, 33°44'N 124°53'W, 

120 m] 

Neosergestes edwardsii (Krøyer, 1855) 

= Sergestes Edwardsii Krøyer, 1855: 28. [nærheden af Linien, 3°S to 10°N, according to Krøyer, 1859] 

= Sergestes oculatus Krøyer, 1855: 28. [tropiske Atlanterhav, 4.5°N 21.5°W, according to Krøyer, 1859] 

= Sergestes brachyorrhos Krøyer, 1859: 272, 281; Plate 3, Figs 5a-f. [i Atlanterhavet, efter Angivelse paa 

omtrent 30°N 33°W] 

= Sergestes intermedius Spence Bate, 1888: 383. [Off Luzon, China Sea] 

= Sergestes ventridentatus Spence Bate, 1888: 431. [N of the Sandwich Islands]


Neosergestes orientalis (Hansen, 1919) 

= Sergestes orientalis Hansen, 1919: 22; Plate 2, fig. 2a-q. [Siboga stns 37, Sailus Ketjil, Paternoster Islands, 

0-27 m; 40, Anchorage off Pulu Kawassang, Paternoster Islands, 12 m; 66, Bank between islands 

of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 96, SE side of Pearl bank, 

Sulu Archipelago, 0-15 m; 106, anchorage off Kapul Island, Sulu Archipelago, 13 m; 118, 1°38'N 

124°28.2'E, 0-900 m; 125, anchorage off Sawan, Siau Island, 27 m; 128, 4°27'N 125°25.7'E, 0-700 m; 

131-133, about 5°56.7'N 126°25'E; 141, 1°0.4'S 127°25.3'E, 0-1500 m; 143, 1°4.5'S 127°52.6'E, 0-1000 m; 

144, Anchorage N of Salomakiëe (Damar-) Island, 45 m; 146, 0°36'S 128°32.7'E, surface; 148, 0°17.6' 

S 129°14.5'E, 0-1000 m; 157, 0°32.9'S 130°14.6'E, 45 m; 177a, 2°30'S 129°28'E; 168, anchorage N of 

Sabuda Island, surface; 172, Gisser, anchorage between this island and Ceram Island, 18 m; 177a, 

2°30'S 129°28'E, surface; 185, 3°20'S 127°22.9'E, Manipa Strait, 0-1536 m; 189a, 2°22'S 126°46'E; 

194-197, 1°55'-1°45.3'S 126°39'-127°8.3'E; 194-197, 1°53.5'-1°45.3'S 126°39'-127°8.3'E; 203, 3°32.5'S 

124°15.5'E, 0-1500 m; 206 or 207, Buton Strait, surface; 220, anchorage off Pasir Pandjang, W coast of 

Binongka, surface; 225, N. 279°E from Southpoint of South Lucipara Island, 894 m; 230, 3°58'S 

128°20'E, 0-2000 m; 245, 4°16.5'S 130°15.8'E, surface] 

= Sergestes geminus Judkins, 1978: 25; Figs 2a-c, 7, 16f-j, 17, 18, 21a. [Eastern Tropical Pacific, 4°48'N 

83°38'W, ca. 200 m at night] 

= Sergestes gibbilobatus Judkins, 1978: 27; Figs 2g, 4c, 7, 19a-h, 20, 21a. [Equatorial Pacific, 0°0'S 

165°42'W, 2550 m wire out, at night] 

Neosergestes semissis (Burkenroad, 1940) 

= Sergestes (Sergestes) semissis Burkenroad, 1940: 42. [Dana stn 3905 I (Bay of Bengal, 4°44'N 88°05.5'E), 

1000 m wire out] 

Neosergestes tantillus (Burkenroad, 1940) 

= Sergestes (Sergestes) tantillus Burkenroad, 1940: 42. [Dana stn 3556 II (Gulf of Panama, 2°52'N 

87°36'W), 2500 m wire out)] 

Parasergestes Judkins & Kensley, 2008 

= Parasergestes Judkins & Kensley, 2008 (type species Sergestes armatus Krøyer, 1855, by original 


Parasergestes armatus (Krøyer, 1855) 

= Sergestes armatus Krøyer, 1855: 31. [7°37'n.Br. og N 22.5° v. Lgd., according to Krøyer, 1859] 

= Sergestes incertus Hansen, 1896: 962. [34°50'S 4°30'W and 40°4'S 53°20'W, surface] 

= Sergestes extensus Hanamura, 1983: 64; Figs 7-8. [Kaiyo Maru stn MT 9-B, Seamount 350, off Baja 

California, 23-05.4N 124-56.9W, 0-1236 m] 

Parasergestes cylindricus (Vereshchaka, 2009) comb. nov. 

= Sergestes cylindricus Vereshchaka, 2009: 96; Figs 47, 48. [Dana stn 3639-1, Southeast Pacific, 39°19'S 

179°18'E (300 m wire out)] 

Parasergestes diapontius (Spence Bate, 1881) 

= Sergestes diapontius Spence Bate, 1881: 194. [Atlantic] 

= Sergestes penerinki Spence Bate, 1888: 418; Plate 76, fig. 3. [North Atlantic Ocean] 

= Sergestes fermerinkii Spence Bate, 1888: 419; Plate 76, fig. 4. [Pacific Ocean, 24°S 148°W] 

= Sergestes semiarmis Spence Bate, 1888: 423; Plate 67, fig. 1. [Challenger stn 354, Mid North Atlantic 

Ocean, 32°41'N 36°6'W, surface] 

Parasergestes halia (Faxon, 1893) 

= Sergestes halia Faxon, 1893: 217. [Albatross stn 3388 (off Panama, 7°06'N 79°48'W, surface to 400 fms)] 

Parasergestes sirenkoi (Vereshchaka, 2009) comb. nov. 

= Sergestes sirenkoi Vereshchaka, 2009: 103; Figs 51, 52. [Dana stn 3593-8, Southwest Pacific, 17°27'S 

179°33'E (300 m wire out)] 

Parasergestes stimulator (Burkenroad, 1940) 

= Sergestes (Sergestes) stimulator Burkenroad, 1940: 41. [Dana stn 3656 VIII (off Sydney, Australia, 

33°26'S 157°02'E), 300 m wire out)] 

Parasergestes vigilax (Stimpson, 1860a)


= Sergestes vigilax Stimpson, 1860a: 45. [Oceano Atlantico prope insulas ”Azores”] 

= Sergestes macrophthalmus Stimpson, 1860a: 46. [Oceano Pacifico, lat. bor. 27,5°, long. orient. 138,5°; 

etiam lat. bor. 35°, long. occ. 155°] 

= Sergestes parvidens Spence Bate, 1888: 409; Plate 74, fig. 3. [the tropical part of the Atlantic; Pacific 

Ocean, north of the Sandwich Islands; off Sydney and Wellington, Australia] 

? = Sergestes spiniventralis Spence Bate, 1888: 426; Plate 67, fig. 5. [North Pacific Ocean] 

? = Sergia Clausi König, 1895: 10; Plate 1, figs 1-7. [Östliches Mittelmeer, 36°12'N 28°54'E, Oberflächenfang] 

Peisos Burkenroad, 1945 

= Peisos Burkenroad, 1945 (type species Peisos petrunkevitchi Burkenroad, 1945, by original designation, 


Peisos petrunkevitchi Burkenroad, 1945: 554; Plates 1-2. [Montevideo, 5-6 fms] 

Petalidium Spence Bate, 1881 

= Petalidium Spence Bate, 1881 (type species Petalidium foliaceum Spence Bate, 1881, by monotypy, 

gender neuter) 

Petalidium foliaceum Spence Bate, 1881: 194. [taken in South Indian Ocean at a depth of about 2100 fms; 

according to Spence Bate, 1888: Challenger stn 159, South of Australia, 47°25'S 130°22'E, 2150 fms] 

Petalidium obesum (Krøyer, 1855) 

= Sergestes obesus Krøyer, 1855: 31. [4.5°N Br. i Atlanterhavet, according to Krøyer, 1959] 

= Sergestes sanguineus Chun, 1889: 538; Plate 3, fig. 1. [Canarischen Inseln] 

Petalidium suspiriosum Burkenroad, 1937: 325; Text-Figs 1-6. [Templeton Crocker Expedition stn 165, 145 

miles N of Clarion Island, Revillagigido Islands, Lower California, E Pacific, 500 fms] 

Sergestes H. Milne edwards, 1830 

= Sergestes H. Milne Edwards, 1830 (type species Sergestes atlanticus H. Milne Edwards, 1830, by 


= Acheles Cocco, 1832 (type species Acheles arachnipodus Cocco, 1832, by monotypy, gender feminine) 

Sergestes atlanticus H. Milne Edwards, 1830: 349; Plate 10, figs 1-9. [l’Océan atlantique, à une grande 

distance des côtes] 

= Sergestes Frisii Krøyer, 1855: 26. [Atlanterhav, 13°N 27.5°W and 20°N 36°W, according to Krøyer, 

1859] 

= Sergestes ancylops Krøyer, 1855: 32. [Atlanterhav 13°N 27.5'W, according to Krøyer, 1859] 

? = Sergestes pacificus Stimpson, 1860a: 45. [Oceano Pacifico, lat. bor. 27,5°, long. orient. 138°] 

= Sergestes ovatoculus Spence Bate, 1888: 408; Plate 74, fig. 2. [N Atlantic Ocean] 

Sergestes cornutus Krøyer, 1855: 29. [Atlanterhavet omtrent 4.5°N for Linien; indtil 10° nord for Linien og 

undtil 8° syd for Linien, according to Krøyer, 1859] 

= Sergestes longispinus Spence Bate, 1888: 417; Plate 76, fig. 2. [Challenger stn 106, Mid Atlantic Ocean, 

1°47'N 24°26'W, within 40 fms of the surface] 

Sergestes formosensis Yokoya & Shibata, 1965: 2; Fig. 2. [Philippine Sea, 21°42'N 123°05'E, 100 m] 

Sergestes grandipes Yokoya & Shibata, 1965: 1; Fig. 1. [Philippine Sea, 13°57'N 125°29'E, 100 m] 

Sergestes hamifer Alcock & Anderson, 1894: 148. [Investigator stn 126, Laccadive Sea, 8°49'0"N 73°18'45"E, 

1370 fms] 

Sergestes latirostris Yokoya & Shibata, 1965: 3; Fig. 3. [Philippine Sea, 13°03'N 125°50'E, 40 m] 

Sergia Stimpson, 1860a 

= Sergia Stimpson, 1860a (type species Sergia remipes Stimpson, 1860a, by monotypy, gender feminine; 

the type species was determined to be a late mastigopus stage by Burkenroad, 1945, who 

identified it with the group of species in what was then the subgenus Sergia) 

Sergia bigemmea (Burkenroad, 1940) 

= Sergestes (Sergia) bigemmeus Burkenroad, 1940: 49. [Dana stn 3570 VI (NE of Tahiti, 3°09'N 

126°09.3'E), 2150 m]


Sergia bisulcata (Wood-Mason & Alcock, 1891b) 

= Sergestes bisulcatus Wood-Mason & Alcock, 1891b: 190. [Investigator stns 100 (Bay of Bengal, 

16°55'41"N, 83°21'18"E), 840 fms; 105 (Arabian Sea, 15°02'N, 72°34'E), 740 fms] 

Sergia burukovskii Vereshchaka, 2000: 121; Figs 31-33. [Dana stn 3980-1, South Atlantic, 23°26'S, 03°56'E, 

1000 m wire out] 

Sergia challengeri (Hansen, 1903) 

= Sergestes challengeri Hansen, 1903: 61; Plate 12, fig. 2a-n. [Challenger stn 173, Western Pacific, off 

Matuku, Fiji Islands, 19°9'35"S, 179°41'50"E, 315 fms] 

Sergia crosnieri Vereshchaka, 2000: 196; Figs 85-87. [Dana stn 3809-4, off Indonesian islands, 6°22'S, 

105°12'E, 50 m wire out] 

Sergia erythraeensis Iwasaki & Couwelaar, 2001: 92; Figs 1-3. [Central Red Sea, 21°23.89'N 38°03.62'E, 150 

m] 

Sergia extenuata (Burkenroad, 1940) 

= Sergestes (Sergia) extenuates Burkenroad, 1940: 45. [Dana stn 3999 II (Saint Helena, S Atlantic, 3°45'S 

10°00'E, 1000 m wire out); see discussion in Vereshchaka, 2000] 

Sergia filicta (Burkenroad, 1940) 

= Sergestes (Sergia) filictum Burkenroad, 1940: 52. [Dana stn 3549 IV (Gulf of Panama, 7°16'N 79°30'W), 

wire out 600 m] 

Sergia foresti Kensley & Judkins, 2008: 151; Fig. 1. [Albatross stn 5241, Pujada Bay, Mindanao, 6°50'45"N 

126°14'38"E, 393 m] 

Sergia fulgens (Hansen, 1919) 

= Sergestes fulgens Hansen, 1919: 17; Plate 1, fig. 6a-g. [Siboga stn 312, Indonesia, Saleh Bay, Sumbawa, 

08°19'S 117°41'E, 174 m] 

Sergia gardineri (Kemp, 1913a) 

= Sergestes gardineri Kemp, 1913a: 55; Plate 7, figs 2-5. [S by E of Farquhar, 10°27'S, 51°17'E; NE of 

Madagascar, between Providence and Alphonse Islands, 8°16'S, 51°26'E; 5 miles off Desroches Atoll] 

Sergia grandis (Sund, 1920) 

= Sergestes grandis Sund, 1920: 16; Figs 22-26. [Michael Sars stns 34, (28°52'N 14°16'W, 2170 m); 49 

(29°6'N, 25°2'W); 51 (31°20'N 35°7'W, 3886 m); 52 (31°24'N, 34°47'W] 

Sergia hansjacobi Vereshchaka, 1994: 91; Figs 23-24. [Dana stn 1198-2, 17°43'N, 64°56'W] 

= Sergia hanseni Vereshchaka, 1994: 91. [nomen nudum] 

Sergia inequalis (Burkenroad, 1940) 

= Sergestes (Sergia) inequalis Burkenroad, 1940: 51. [Dana stn 3768 (N of NW New Guinea, 1°20'S 

138°42'E), 2500 m wire out)] 

Sergia inoa (Faxon, 1893) 

= Sergestes inous Faxon, 1893: 216. [Albatross stn 3380 (East Pacific, off Malpelo Island, Colombia, 

04°03'N, 81°31'W, 899 fms)] 

Sergia japonica (Spence Bate, 1881) 

= Sergestes japonicus Spence Bate, 1881: 194. [south of Japan; according to Spence Bate, 1888: Challenger 

stn 232, western Pacific off southern coast of Japan, 35°11'N, 139°28'E, 345 fms] 

= Sergestes mollis Smith, 1884: 419. [Albatross stns 2002, 37°20'42"N 74°17'36"W, 641 fms; 2018, 

37°12'22"N 74°17'36"W, 788 fms; 2040, 38°35'13"N 58°16'00"W, 2226 fms; 2045, 40°04'20"N 68°43'50"W, 

373 fms; 2051, 39°41'00"N 69°20'20"W, 1106 fms; 2083, 40°26'40"N 67°05'15"W, 959 fms; 2093, 

39°42'50"N 71°01'20"W, 1000 fms; 2094, 39°44'30"N 71°04'00"W, 1022 fms; 2097, 37°56'20"N 

70°57'30"W, 1917 fms; 2099, 27°12'20"N 69°39'00"W, 2949 fms; 2100, 39°22'00"N 68°34'30"W, 1628 

fms; 2101, 39°18'30"N 68°24'00"W, 1686 fms; 2103, 38°47'20"N 72°37'00"W, 1091 fms; 2104, 38°48'00"N 

72°40'30"W, 991 fms; 2105, 37°50'00"N 73°03'50"W, 1395 fms; 2110, 35°12'10"N 74°57'15"W, 516 fms; 

2116, 35°45'23"N 74°31'25"W, 888 fms] 

= Sergestes profundus Spence Bate, 1888: 428 (partim). [Challenger stn 300, southeastern Pacific west 

of Valparaiso, Chile, 33°42'S, 78°18'W, 1375 fms; lectotype designation by Hansen, 1903] 

Sergia jeppeseni Vereshchaka, 2000: 179; Figs 67, 74-75; Plate 5F. [Dana stn 3943-1, Western Indian Ocean 

off Mombasa, 5°30'S, 40°40'E, 500 m wire out]


Sergia kensleyi Vereshchaka, 2000: 110; Figs 17, 24-25; Plate 4D. [Dana stn 3970-1, Western Indian Ocean 

off Mozambique, 34°09'S, 27°38'E] 

Sergia laminata (Burkenroad, 1940) 

= Sergestes (Sergia) laminatus Burkenroad, 1940: 53. [Dana stn 3933 I (N of Madagascar, 11°18'S 

50°03'E), 4000 m wire out] 

= Sergestes (Sergia) guineensis Crosnier & Forest, 1973: 343; Fig. 118. [Ombango stn 394-8, 5°52'S 

10°00'E, 0-2500 m] 

Sergia lucens (Hansen, 1922) 

= Sergestes lucens Hansen, 1922: 12, 38. [Suruga Bay; see discussion in Vereshchaka, 2000] 

Sergia manningorum Froglia & Gramitto, 2000: 72; Figs 1-3. [Atlantis II, cruise 60 stn RH2281, off Congo, 

11°23'S 10°55'E, 190-200 m] 

Sergia maxima (Burkenroad, 1940) 

= Sergestes (Sergia) maximus Burkenroad, 1940: 47. [Dana stn 3933 I (N of Madagascar, 11°18'S 

50°03'E, 4000 m wire out)] 

Sergia oksanae Vereshchaka, 2000: 182: Figs 73, 76-77; Plate 5E. [Dana stn 3736-3, Mindanao Sea, 9°17'N, 

123°58'E] 

Sergia phorca (Faxon, 1893) 

= Sergestes phorcus Faxon, 1893: 217. [Albatross stns 3382 (off Panama, 6°21'N 80°41'W, 1793 fms); 

3388 (off Panama, 7°06'N 79°48'W, 1168 fms); 3401 (off Galapagos Islands, 0°50'S 88°58'30"W, 395 

fms); 3386 (off Panama, 7°33'12"N 79°17'15"W, 242 fms); 3437 (Gulf of California, 27°39'40"N 

111°00'30"W, 628 fms)] 

Sergia plumea (Illig, 1927) 

= Sergestes plumeus Illig, 1927: 295; Figs 30-32. [Valdivia stn 268, Indischer Ozean, südlich von Ras 

Hafun, 9°6'N, 53°41'E, im Vertikalnetzfang von 1500 m Tiefe] 

Sergia potens (Burkenroad, 1940) 

= Sergestes (Sergia) potens Burkenroad, 1940: 48. [Dana stn 3975 VII (Agulhas Bank, South Africa, 

35°42'S 18°37'E, 600 m wire out)] 

Sergia prehensilis (Spence Bate, 1881) 

= Sergestes prehensilis Spence Bate, 1881: 193. [off Japan, 500 fms; according to Spence Bate, 1888: 

Challenger stn 236, off Japan, 34°58'N, 139°29'E, 775 fms] 

= Sergestes gloriosus Stebbing, 1905: 84; Plates 22-23. [off Sandy Point, 800 fms] 

= Sergestes fujiyamaensis Nakazawa, 1932: 32. [Type locality not indicated] 

Sergia profunda (Spence Bate, 1888) 

= Sergestes profundus Spence Bate, 1888: 428. [Challenger stn 137, 35°59'S 1°34'E, 2550 fms; 300, W of 

Valparaiso, 33°42'S 78°18'W, 1375 fms] 

Sergia regalis (Gordon, 1939) 

= Sergestes regalis Gordon, 1939: 498; Figs 1-4. [Discovery stn 81, South Atlantic, 32°45'S, 8°47'W] 

= Sergestes (Sergia) creber Burkenroad, 1940: 44. [Dana stn 3766 XVIII (N of NW new Guinea, 1°13'S 

138°42'E), 2900 m wire out] 

Sergia remipes Stimpson, 1860a: 46. [Oceano Pacifico, lat bor. 27½° log. orient. 138½°] 

Sergia robusta (Smith, 1882) 

= Sergestes robustus Smith, 1882: 97; Plate 16, fig. 5-8b. [Blake stn 328, 34°28'25''N 75°22'50''W, 1632 

fms; Albatross stns 893 (37°17'N 73°21'W), 372 fms; 952 (34°28'50"N 75°22'50"W), 388 fms] 

= Sergestes dissimilis Spence Bate, 1888: 437. [St. Vincent, Cape Verde Island, at the surface] 

= Sergestes mediterraneus Hansen, 1896: 954. [Ragusa and Lesina, Adriatic Sea] 

Sergia scintillans (Burkenroad, 1940) 

= Sergestes (Sergia) scintillans Burkenroad, 1940: 43. [Dana stn 3622 I (SE of New Caledonia, 25°54'S 

172°36.9'E, 300 m wire out)] 

Sergia splendens (Sund, 1920) 

= Sergestes splendens Sund, 1920: 14; Figs 16-18. [Michael Sars stns 29 (35°10'N 7°55'W) 440 m wire 

out; 42 (28°2'N 14°17'W), 300 m wire out; 45 (28°42'N 20°0'W), 100, 200, 300, 2000, & 3000 m wire 

out; 49 (29°6'N 25°2'W), 3000 m wire out; 51 (31°20'N 35°7'W), 200, 300 & 1000 m wire out; 52 

(31°24'N 34°47'W), 100 m wire out; 53 (34°59'N 33°1'W), 100, 300, 600 & 2600 m wire out; 56


(36°53'N 29°17'W), 200 & 300 m wire out; 62 (36°52'N 39°55'W), 2000 m wire out; 64 (34°44'N 

47°52'W), 100 & 2000 m wire out; 66 (39°30'N 49°42'W), 1000 m wire out; 67 (40°17'N 50°39'W), 

1200 & 2000 m wire out)] 

= Sergestes Richardi Hansen, 1920: 482. [nomen novum for Sergestes splendens Sund, 1920, considered 

a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1920] 

= Sergestes crassus Hansen, 1922: 98; Plate 5, fig. 4. [nomen novum for Sergestes splendens Sund, 1920, 

considered a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1922] 

Sergia stellata (Burkenroad, 1940) 

= Sergestes (Sergia) stellatus Burkenroad, 1940: 44. [Dana stn 3908 I (SE of Sri Lanka, 4°28'N 82°13'E, 


Sergia talismani (Barnard, 1947) 

= Sergestes splendens Hansen, 1919: 18. [nomen nudum] 

= Sergestes splendens Hansen, 1920: 480; nec Sund, 1920. [Talisman Expedition, stn 113, canal de Saint- 

Vincent, 16°52'N 27°30 - 27°31'W, 550-760 m] 

= Sergestes talismani Barnard, 1947: 384. [nomen novum for Sergestes splendens Hansen, 1919 nec 

Sund, 1920] 

Sergia tenuiremis (Krøyer, 1855) 

= Sergestes tenuiremis Krøyer, 1855: 30. [2°N, 21°W, according to Krøyer, 1859] 

= Sergestes Kröyeri Spence Bate, 1881: 193. [off Kermadec Island, about 500 fms; according to Spence 

Bate, 1888: Challenger stn 170, off the Kermadec Islands, 29°55'S 178°14'W, 520 fms] 

= Sergestes junceus Spence Bate, 1888: 416; Plate 76, fig. 1. [South Pacifc Ocean] 

= Sergestes longicollis Spence Bate, 1888: 421; Fig. 1. [Challenger near stn 131, South Atlantic Ocean, 

29°35'S 28°9'W; stn 295, South Pacific Ocean, 38°7'S 94°4'W, 1500 fms] 

= Sergestes tropicus Sund, 1920: 18; Figs 27, 28, 30-33. [Michael Sars stns 29 (35°10'N 7°55'W); 45 

(28°42'N 20°0'W); 49 (29°6'N 25°2'W); 51 (31°20'N 35°7'W, 3886 m); 52 (31°24'N 34°47'W); 53 (34°59'N 

33°1'W, 2615-2865 m); 56 (36°53'N 29°17'W, 3239 m); 64 (34°44'N 47°52'W)] 

Sergia umitakae Hashizume & Omori, 1995: 72; Figs 1-4. [South of Sri Lanka] 

Fig. 7. Sicyonella aff. maldivensis Borradaile, 1910. Photo by Tin-Yam Chan.


Sergia vityazi Vereshchaka, 2000: 157; Figs 51, 57-58. [Dana stn 3601-1, southwestern Pacific, 18°21'S, 

178°21'E] 

Sergia wolffi Vereshchaka, 1994: 88; Figs 19-21. [Dana stn 1217-1, 18°50'N, 79°07'W] 

Sicyonella Borradaile, 1910 

= Aphareus Paul’son, 1875 (type species Aphareus inermis Paul’son, 1875, by monotypy, gender masculine; 

invalid junior homonym of Aphareus Cuvier & Valenciennes, 1830 (Pisces)) 

= Sicyonella Borradaile, 1910 (type species Sicyonella maldivensis Borradaile, 1910, by monotypy, gender 

feminine) 

= Aphareocaris Calman, 1913 (nomen novum for Aphareus Paul’son, 1875, gender feminine) 

Sicyonella antennata Hansen, 1919: 30; Plate 2, fig. 5a-c; Plate 3, fig. 1a-f. [Siboga stn 258, Tual anchorage, 

Kei Islands, 22 m] 

Sicyonella inermis (Paul’son, 1875) 

= Aphareus inermis Paul’son, 1875: 117; Plate 18, fig. 3. [Red Sea] 

= Aphareocaris elegans Calman, 1913: 219; Plate 16, figs 1-16. [Thursday Island, Torres Straits] 

Sicyonella maldivensis Borradaile, 1910: 259; Plate 16, figs 3-3a. [North Male Atoll, Maldives; lectotype 

designation by Fukuoka, Tamaki & Kikuchi, 2005] (Fig. 7) 

Suborder PleoCyeMATA Burkenroad, 1963 

Infraorder PRoCARIDIDeA Felgenhauer & Abele, 1983 

Family PRoCARIDIDAe Chace & Manning, 1972 

Procaris Chace & Manning, 1972 

= Procaris Chace & Manning, 1972 (type species Procaris ascensionis Chace & Manning, 1972, by 


Procaris ascensionis Chace & Manning, 1972: 6; Figs 4-9. [coral pool back of Shelly Beach, Ascension Island] 

Procaris chacei C.W.J. Hart & Manning, 1986: 408; Figs 1-26. [Bermuda, Hamilton Parish, Green Bay Cave 

(North Shore passage), 32°19'N 64°44'W] 

Procaris hawaiana Holthuis, 1973a: 12; Figs 4-6. [near Nukuele Point, Cape Kinau, Maui, Hawaiian Islands, 

in lava pool] (Fig. 8) 

Procaris mexicana von Sternberg & Schotte, 2004: 514-522; Figs 1-3. [México, Cueva Quebrada, Chankanaab 

Park, Cozumel, Quintana Roo] 

Procaris noelensis Bruce & Davie, 2006: 23-33; Figs 1-3. [Runaway Cave, c. 1.5 km south from the North 

East Point of Christmas Island] 

Fig. 8. Procaris hawaiana Holthuis, 1973. Photo by Troy Sakihara.


Vetericaris Kensley & Williams, 1986 

= Vetericaris Kensley & Williams, 1986 (type species Vetericaris chaceorum Kensley & Williams, 1986, 

by monotypy, gender feminine) 

Vetericaris chaceorum Kensley & Williams, 1986: 419, Figs 2-7. [Lua o Palahemo, 150 m north of shoreline 

at Ka Lae (South Point), Hawaii Island, 18°55'N 155°42'W, 33 m] 

Infraorder STeNoPoDIDeA Spence Bate, 1888 

Family MACROMAXILLOCARIDIDAE Alvarez, Iliffe & Villalobos, 2006 

Macromaxillocaris Alvarez, Iliffe & Villalobos, 2006 

= Macromaxillocaris Alvarez, Iliffe & Villalobos, 2006 (type species Macromaxillocaris bahamaensis 

Alvarez, Iliffe & Villalobos, 2006, by original designation and monotypy, gender feminine) 

Macromaxillocaris bahamaensis Alvarez, Iliffe & Villalobos, 2006: 370; Figs 2-7. [Oven Rock Cave, Great 

Guana Cay, Exuma Cays, Bahamas] (Fig. 9) 

Fig. 9. Macromaxillocaris bahamaensis Alvarez, Iliffe & Villalobos, 2006. Photo by T.M. Iliffe. 

Family SPoNGIColIDAe Schram, 1986 

Engystenopus Alcock & Anderson, 1894 

= Engystenopus Alcock & Anderson, 1894 (type species Engystenopus palmipes Alcock & Anderson, 

1894, by monotypy, gender masculine) 

Engystenopus palmipes Alcock & Anderson, 1894: 149; Plate 9, fig. 1. [Investigator stn 172, Bay of Bengal, 

off Trincomallee, 200-350 fms] 

Globospongicola Komai & Saito, 2006 

= Globospongicola Komai & Saito, 2006 (type species Globospongicola nudibranchus Komai & Saito, 


Globospongicola nudibranchus Komai & Saito, 2006: 268; Figs 1-6. [Kai Islands, Banda Sea, Indonesia, 

6°05'S 132°44'E, 268-210 m] 

Globospongicola spinulatus Komai & Saito, 2006: 276; Figs 7-12. [Vanuatu, 17°50.35'S 168°39.33'E, 437-504 m]


Microprosthema Stimpson, 1860a 

= Microprosthema Stimpson, 1860a (type species Microprosthema valida Stimpson, 1860a, by monotypy, 

gender neuter) 

= Stenopusculus Richters, 1880 (type species Stenopusculus crassimanus Richters, 1880, designated by 

Holthuis, 1955, a junior subjective synonym of Microprosthema valida Stimpson, 1860a, gender masculine) 

Microprosthema emmiltum Goy, 1987: 717; Figs 1-4. [1°16'36''S 90°29'42''W, off Black Beach, Isla Santa Maria, 

Galapagos Islands; rocky shores] 

Microprosthema fujitai Saito & Okuno, 2011: 84; Figs 1-7. [Maeda-misaki, Okinawa Island, Ryukyu Islands, 

3 m] 

Microprosthema granatense Criales, 1997: 538; Figs 1-4. [Granate Bay, Caribbean coast of Colombia, 

11°18'N 74°9'W, 23 m] 

Microprosthema inornatum Manning & Chace, 1990: 26; Figs 12-13. [off North Point, Ascension Island, 18 

m] 

Microprosthema jareckii Martin, 2002: 109; Figs 1-5. [Monkey Point, Guana Island, British Virgin Islands, 

10 m] 

Microprosthema looensis Goy & Felder, 1988: 1286; Figs, 5-7. [near study finger number 3, Looe Key, Florida, 

USA, 4-5 m] 

Microprosthema lubricum Saito & Okuno, 2011: 93; Figs 8-11. [Agenashiku Islet, the Kerama Islands, the 

Ryukyu Islands, 3m] 

Microprosthema manningi Goy & Felder, 1988: 1277; Figs 1-4. [17°29'N 88°10'W, Stann Creek District, Carrie 

Bow Cay, Belize, near shore in dead conch shell] 

Microprosthema plumicorne (Richters, 1880) (Fig. 10) 

= Stenopusculus plumicornis Richters, 1880: 167; Plate 18, figs 16-26. [Fouquets, Mauritius] 

Microprosthema scabricaudatum (Richters, 1880) 

= Stenopusculus scabricaudatus Richters, 1880: 168; Plate 18, figs 30-32. [Fouquets, Mauritius] 

Microprosthema semilaeve (von Martens, 1872) 

= Stenopus semilaevis von Martens, 1872: 144. [Cuba] 

= Stenopusculus spinosus Pocock, 1890: 523. [Fernando de Noronha] 

Fig. 10. Microprosthema plumicorne (Richters, 1880). Photo by Arthur Anker.


Microprosthema validum Stimpson, 1860a 

= Microprosthema valida Stimpson, 1860a: 45. [in sinu insulae Ousima, sublittoralis, in locis lapillosis 

algosisque] 

= Stenopusculus crassimanus Richters, 1880: 168; Plate 18, figs 27-29. [Fouquets, Mauritius] 

= Stenopus robustus Borradaile, 1910: 260; Plate 16, fig. 4. [Salomon Atoll, Chagos Archipelago] 

Paraspongicola de Saint laurent & Cleva, 1981 

= Paraspongicola de Saint Laurent & Cleva, 1981 (type species Paraspongicola pusilla de Saint Laurent 

& Cleva, 1981, by original designation and monotypy, gender masculine) 

Paraspongicola acantholepis Komai, 2011: 305; Figs 12-15. [R/V Tansei-maru, KT07-31, stn L3-200, Isu Islands, 

N of Toshima Island, 34°34.04'N 139°18.37'E to 34°33.56'N 139°17.81'E, 187-261 m] 

Paraspongicola inflatus de Saint Laurent & Cleva, 1981 

= Spongicola inflata de Saint Laurent & Cleva, 1981: 179; Fig. 13. [14°02.2'N 120°17.7'E, 193-184 m] 

Paraspongicola pusillus de Saint Laurent & Cleva, 1981 

= Paraspongicola pusilla de Saint Laurent & Cleva, 1981: 181; Figs 14-15. [13°56.3'N 120°16.2'E, 150- 

159 m] 

Spongicola De Haan, 1844 [in De Haan, 1833-1850] 

= Spongicola De Haan, 1844 [in De Haan, 1833-1850] (type species Spongicola venusta De Haan, 1844 

[in De Haan, 1833-1850], by monotypy, gender masculine) 

Spongicola andamanicus Alcock, 1901 

= Spongicola andamanica Alcock, 1901: 148; Plate 2, fig. 2. [Andaman Sea, 170 and 238-290 fms] 

= Spongicola henshawi Rathbun, 1906: 901; Plate 24, fig. 8. [south coast of Molokai Island, Hawaii, 

169-182 fms] 

= Spongicola henshawi spinigera de Saint Laurent & Cleva, 1981: 174; Figs 10b, 11c, g-h. [8°47'N 

123°31.2'E, 333 m] 

= Spongicola holthuisi de Saint Laurent & Cleva, 1981: 177; Fig. 12. [13°59.8'N 120°18.6'E, 192-188 m] 

Spongicola cubanicus Ortiz, Gómez & Lalana R., 1994 

= Spongicola cubanica Ortiz, Gómez & Lalana R., 1994: 189; Figs 2-7. [Bahía Honda, costa norte de la 

Provincia de Pinar del Rio, 300 m, asociado a la esponja silícea Dactylocalyx pumiceus] 

Spongicola depressus Saito & Komai, 2008: 103; Figs 8-10. [Loyalty Islands, depth unknown] 

Spongicola goyi Saito & Komai, 2008: 107; Figs 11-15. [22°03.60'S 167°27.00'E, 850 m] 

Spongicola japonicus Kubo, 1942a 

= Spongicola japonica Kubo, 1942a: 90; Figs 1-2. [Kumanonada, off Mie Prefecture, about 300 m, commensal 

with Euplectella marshalii] 

Spongicola levigatus Hayashi & Ogawa, 1987 

= Spongicola levigata Hayashi & Ogawa, 1987: 367; Figs 1-4. [East China Sea, 30°44.7'N 127°48.3'E, 

about 200 m] 

Spongicola parvispinus Zarenkov, 1990 

= Spongicola parvispina Zarenkov, 1990: 218. [Sala-y-Gomez ridges, southeastern Pacific, 470-485 m] 

Spongicola robustus Saito & Komai, 2008: 118; Figs 20-24. [Mauritius, 10°29.7'S 61°12.4'E, 115-110 m] 

Spongicola venustus De Haan, 1844 [in De Haan, 1833-1850] 

= Spongicola venusta De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 9 (1844); 194, Plate P (1849). 

[Japan; lectotype designation by Yamaguchi & Baba, 1993] 

Spongicoloides Hansen, 1908 

= Spongicoloides Hansen, 1908 (type species Spongicoloides profundus Hansen, 1908, by monotypy, 


Spongicoloides evolutus (Bouvier, 1905b) 

= Spongicola evoluta Bouvier, 1905b: 983. [au large des côtes du Sahara, 882 m] 

Spongicoloides galapagensis Goy, 1980: 760; Figs 1-4. [Galapagos Islands, 00°29' 89°54'30''W, 717 m] 

Spongicoloides hawaiiensis Baba, 1983: 477; Figs 1-2. [French Frigate Shoals, north-western Hawaiian Islands, 

640 m, associated with glass sponge Euplectella sp.]


Spongicoloides iheyaensis Saito, Tsuchida & Yamamoto, 2006: 224; Figs 3-8. [the North Knoll of the Iheya 

Ridge, Ryukyu Islands, 27°47.17'N 26°53.91'E, 988 m] 

Spongicoloides inermis (Bouvier, 1905a) 

= Richardina inermis Bouvier, 1905a: 749. [dans les parages de Sainte-Lucie, 22-423 brasses] 

Spongicoloides koehleri (Caullery, 1896) 

= Spongicola Kœhleri Caullery, 1896: 382; Plate 16, figs 1-9. [Golfe de Gascogne, 45°57'N 6°21'E, 1410 

m] 

Spongicoloides novaezelandiae Baba, 1979: 311; Figs 1-2. [Chatham Rise, 44°44.0'S 175°42.0'E, 990-1110 m] 

Spongicoloides profundus Hansen, 1908: 45; Plate 3, figs 5a-k; Plate 4, figs 1a-l. [78 miles off the southwest 

coast of Iceland, 60°37'N 27°52'W, 1480 m; lectotype designation by Goy, 1982] 

Spongicoloides tabachnicki Burukovsky, 2009: 498; Figs 1-3. [Atlantic Ocean, Florida Straits, 23°54'N 

81°27'W, 1003-1336 m] 

Spongiocaris Bruce & Baba, 1973 

= Spongiocaris Bruce & Baba, 1973 (type species Spongiocaris semiteres Bruce & Baba, 1973, by original 


Spongiocaris hexactinellicola Berggren, 1993: 784; Figs 1-5. [24°30'N 74°28'W, Tartar Bank, Cat Island, Bahamas, 

60-610 m; in the internal cavity of Euplectella jovis] 

Spongiocaris goyi Ortiz, Lalana & Varela, 2007: 26; Figs 1-2. [sur de la Bahia de Cienfuegos, 297-351 m, en 

la esponja Pheronema annae ?] 

Spongiocaris semiteres Bruce & Baba, 1973: 155; Figs 1-6. [13 miles S.E. of Durban, South Africa, 230 fms, 

from hexactinellid sponge] 

Spongiocaris yaldwyni Bruce & Baba, 1973: 163; Figs 7-10. [15 miles N, 50°E of Plate Island in the Bay of 

Plenty, northeast of North Island, New Zealand, 320-340 fms, from Regadrella okinoseana] 

Family STeNoPoDIDAe Claus, 1872 

Juxtastenopus Goy, 2010 

= Juxtastenopus Goy, 2010 (type species Engystenopus spinulatus Holthuis, 1946a, by original designation 


Juxtastenopus spinulatus (Holthuis, 1946a) 

= Engystenopus spinulatus Holthuis, 1946a: 45; Plate 4, figs a-b. [Lobetobi Strait, E. of Flores, 8°27'S 

122°54.5'E, 247 m] 

Odontozona Holthuis, 1946a 

= Odontozona Holthuis, 1946a (type species Stenopus ensiferus Dana, 1852a, by original designation, 


Odontozona addaia Pretus, 1990: 343; Figs 1-6. [40°01'57''N 4°11'46''E, littoral marine cave near Port 

Addaia Bay, NE coast of Minorca, Balearic Islands, Spain] 

Odontozona anaphorae Manning & Chace, 1990: 29; Figs 14-15. [off North Point, Ascension Island, 10 m] 

Odontozona crinoidicola Saito & Fujita, 2009: 124; Figs 1-7. [Mizugama, Okinawa islands, Ryukyu Islands, 

3.2 m, associated with Phanogenia gracilis] 

Odontozona edwardsi (Bouvier, 1908b) 

= Richardina Edwardsi Bouvier, 1908b: 888. [au large des côtes marocaines et soudanaises] 

Odontozona ensifera (Dana, 1852a) 

= Stenopus ensiferus Dana, 1852a: 27. [archipelago Viti] 

Odontozona fasciata Okuno, 2003: 167; Figs 1-6. [Shimoji-shima Island, Miyako Group, Ryukyu Islands, 

24°49.6'N 125°08.2'E, 25 m] 

Odontozona foresti Hendrickx, 2002: 406; Figs 1-3. [southern Gulf of California, 25°43.5'N 109°53.7'W, 

1240-1270 m] 

Odontozona libertae Gore, 1981: 153; Figs 4-5. [Elbow Reef, off Key Largo, Monroe County, Florida] 

Odontozona minoica Dounas & Koukouras, 1989: 341; Figs 1-4. [35°29'10''N 24°17'20''E, off north-western 

coast of Crete, 330 m]


Odontozona rubra Wicksten, 1982: 130; Figs 1-2. [Isla Blanca, off Guaymas, Sonora, Mexico, approx. 

27°52'N 110°52'W, 6-9 m] 

Odontozona sculpticaudata Holthuis, 1946a: 37; Plate 2, fig. F; Plate 4, fig. C. [Sape Strait, east of Soembawa, 

70 m] 

Odontozona spinosissima Kensley, 1981: 66; Figs 4-5. [31°59'S 29°22'E, 150-200 m] 

Odontozona spongicola (Alcock & Anderson, 1899) 

= ?Richardina spongicola Alcock & Anderson, 1899: 291. [Andaman Sea, 498 fms, from hexactinellid 

sponge] 

Odontozona striata Goy, 1981: 843; Figs 1-5. [Gulf of Mexico, West of Cabo San Antonio, Cuba] 

Richardina A. Milne-edwards, 1881a 

= Richardina A. Milne-Edwards, 1881a (type species Richardina spinicincta A. Milne-Edwards, 1881a, 

by monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in 

Opinion 712 in 1964) 

Richardina fredericii Lo Bianco, 1903: 250; Plate 8, figs 27-28. [a circa 9 chilometri da Punta Campanella, 

direzione S.E., 1100 m; a circa 6.5 chilometri dai Galli di Positano, direzione E., 950 m; a circa 9 

chilometri dai Galli di Positano, direzione O.S.O., 1100 m; a circa 10 chilometri da Punta Carena, 

direzione E.N.E., 100 m] 

Richardina ohtsukai Saito & Komatsu, 2009: 31; Figs 1-6. [Kerama Islands, N of Nagan-nu Island, 

26°23.15'N 127°30.09'E, 730-728 m] 

Richardina parvioculata Saito & Komatsu, 2009: 39; Figs 7-9. [off Tosa Bay, western Japan, 32°56.45'N 

133°30.52'E, 517 m] 

Richardina spinicincta A. Milne-Edwards, 1881a: 933. [44°48'30''N 7°0'30''E, Golfe de Gascogne, 5100 m] 

Stenopus latreille, 1819 

= Byzenus Rafinesque, 1814 (type species Byzenus scaber Rafinesque, 1814, by monotypy, an invalid 

senior synonym of Stenopus spinosus Risso, 1827, gender masculine; name of genus and of its type 

species suppressed for the purposes of the Principle of Priority but not for those of the Principle of 

Homonymy, placed on the Official Index of Rejected and Invalid Generic (resp. Specific) Names in 

Zoology in Opinion 522 in 1958) 

= Stenopus Latreille, 1819 (type species Palæmon hispidus Olivier, 1811, by monotypy, gender masculine; 

name placed on the Official list of Generic Names in Zoology in Opinion 522 in 1958) 

= Embryocaris Ortmann, 1893 (type species, Embryocaris stylicauda Ortmann, 1893, by monotypy, a 

junior subjective synonym of Palæmon hispidus Olivier, 1811, gender feminine) 

Stenopus chrysexanthus Goy, 1992: 80; Figs 1-4. [Heron Island, Capricorn Group, Queensland, Australia, 

ca. 23°25'S 151°55'E, south west reef flat, algal crest] 

Stenopus cyanoscelis Goy, 1984: 116. [Broadhurst Reef, Off Townsville, Queenland] 

Stenopus devaneyi Goy, 1984: 117. [French Polynesia, Marquesas Islands, Nuka Hiva Island, outer portion 

of Taiohae Baie, W side of islet, Sentinelle de l’Est, 6.2 m, according to Goy & Randall, 1986] 

Stenopus earlei Goy, 1984: 117. [off Makua, O’ahu Island, Hawaiian Islands, 39.6 m, according to Goy & 

Randall, 1986] 

Stenopus goyi Saito, Okuno & Chan, 2009: 111; Figs 1-5. [Donggang fishing port, Pingtung County, Taiwan, 

commercial trawler] 

Stenopus hispidus (Olivier, 1811) (Fig. 11) 

= Squilla Groenlandica Seba, 1759: 54; Plate 21, figs 6-7. [Type locality not indicated; see Davie, 2002 

for status of the older synonyms] 

= Astacus muricatus Olivier, 1791: 346. [mer du Groënland] 

= Cancer (Astacus) longipes Herbst, 1793 [in Herbst, 1791-1796]: 90; Plate 31, fig. 2. [Grönland] 

= Penæus borealis Latreille, 1802: 250. [mers du Nord] 

= Palæmon hispidus Olivier, 1811: 666. [Type locality not indicated] 

= Palæmon ? asper Latreille, 1818: 3; Plate 293, fig. 3. [Type locality not indicated] 

= Embryocaris stylicauda Ortmann, 1893: 85; Plate 6, fig. 4. [Plankton-Expedition der Humboldt-Stiftung, 

südl. Aequatorialstrom JN 235 (0.1°S 44.2°W, 0-400 m)]


Fig. 11. Stenopus hispidus (Olivier, 1811). Photo by Arthur Anker. 

Stenopus pyrsonotus Goy & Devaney, 1980: 781; Figs 1-6. [off Pokai Bay, Oahu Island, Hawaii, 22.9 m] 

Stenopus scutellatus Rankin, 1898: 242; Plate 24, fig. 3. [Silver Cay, New Providence] 

Stenopus spinosus Risso, 1827: 66; Plate 3, fig. 8. [golfe de Nice, régions profondes] 

= Byzenus scaber Rafinesque, 1814: 23. [Sicile; name suppressed for the purposes of the Principle of 

Priority but not for those of the Principle of Homonymy, placed on the Official Index of Rejected 

and Invalid Specific Names in Zoology in Opinion 522 in 1958] 

Stenopus tenuirostris De Man, 1888b: 567; Plate 22a, fig. 5. [Amboina] 

Stenopus zanzibaricus Bruce, 1976a: 90; Figs 1-5. [5°46.6'S 39°23.5'E, Mwenba Island, off N.E. Zanzibar 

Island, 0.5 m] 

Infraorder CARIDeA Dana, 1852b 

Superfamily PASIPHAeoIDeA Dana, 1852a 

Family PASIPHAeIDAe Dana, 1852a 

Alainopasiphaea Hayashi, 1999 

= Alainopasiphaea Hayashi, 1999 (type species Pasiphaea nudipedia Burukovsky, 1993b, by original 

designation and monotypy, gender feminine) 

Alainopasiphaea australis (Hanamura, 1989) 

= Pasiphaea australis Hanamura, 1989: 59; Figs 5-7. [Maria Island area, off east coast of Tasmania, 

southern Australia, 42°42.0'S 148°24.0'E, 345-350 m] 

Alainopasiphaea nudipeda (Burukovsky, 1993b) 

= Pasiphaea nudipeda Burukovsky, 1993b: 35; Figs 1(8-13). [25°28'-25°34'5"S 35°08'-35°00'E, 535-490 m]


Eupasiphae Wood-Mason in Wood-Mason & Alcock, 1893 

= Eupasiphaë Wood-Mason in Wood-Mason & Alcock, 1893 (type species Parapasiphaë (Eupasiphaë) 

Gilesii Wood-Mason, 1892, designated by Holthuis, 1955b, gender feminine) 

Eupasiphae gilesii (Wood-Mason, 1892) 

= Parapasiphaë Gilesii Wood-Mason, 1892: Plate 3, fig. 8; accompanying text in Wood-Mason in 

Wood-Mason & Alcock, 1893: 166 (as Parapasiphaë (Eupasiiphaë) Gilesii). [off Cinque Island, Andaman 

Sea, 650 fms] 

= Eupasiphae rhinocerata Burukovsky, 1977: 473; Figs A-C. [off Morocco, near Meteor Shoal, 30°00'N 

28°43'W, 1400-2060 m] 

Eupasiphae latirostris (Wood-Mason & Alcock, 1891b) 

= Parapasiphaë latirostris Wood-Mason & Alcock, 1891b: 196. [Investigator stn 105 (Arabian Sea, 

15°2'N 72°34'E), 740 fms] 

Eupasiphae paucidentata Crosnier, 1988b: 789; Figs 2c, 3-4, 5b-c. [Madagascar, N.O., 21°26.5S 43°11'E, 810- 

1020 m] 

Eupasiphae serrata (Rathbun, 1902a) 

= Parapasiphæ serrata Rathbun, 1902a: 904. [off Cortez Bank, California, 984 fms] 

= Parapasiphae Grimaldii Coutière, 1911a: 157. [Princesse-Alice stn 3036, off Spain and Portugal, 

36°06'40"N, 10°18'W, 0-4740 m, according to Crosnier, 1988b] 

Glyphus Filhol, 1884 

= Glyphus Filhol, 1884 (type species Glyphus marsupialis Filhol, 1884, by monotypy, gender masculine) 

= Sympasiphaea Alcock, 1901 (type species Sympasiphaea annectens Alcock, 1901, by monotypy, gender 

feminine) 

Glyphus marsupialis Filhol, 1884: 231. [Talisman stn 75 (25°39'N 16°02'W)] 

= Sympasiphaea annectens Alcock, 1901: 63. [Arabian Sea, south-west of Cape Comorin, 487 fms] 

= Sympasiphaea imperialis Terao, 1922: 110; unnumbered figure. [purchased at the market of Uchiura, 

Prov. Idzu] 

Leptochela Stimpson, 1860a 

= Leptochela Stimpson, 1860a (type species Leptochela gracilis Stimpson, 1860a, designated by Kemp, 


= Proboloura Chace, 1976 (type species Leptochela carinata Ortmann, 1893, by original designation 

and monotypy, gender feminine) 

Leptochela (Leptochela) aculeocaudata Paul’son, 1875 

= Lep.[tochela] aculeocaudata Paul’son, 1875: 100; Plate 16, figs 1-1s. [Red Sea] 

Leptochela (Leptochela) bermudensis Gurney, 1939a 

= Leptochela bermudensis Gurney, 1939a: 427; Figs 1-10. [about 7 miles off the south shore, between 

1000 m and surface] 

? = Leptochela carinata Ortmann, 1893: 41 (partim, not Plate 4, fig. 1). [Plankton-Expedition der Humboldt-Stiftung, 

Küstenbank vor der Tocantins-Mündung, JN 236 (0.1°S 45.2°W), 50-100 m] 

Leptochela (Leptochela) chacei Hayashi, 1995: 85; Figs 2-4. [Viet-Nam, Cai Dua] 

Leptochela (Leptochela) crosnieri Hayashi, 1995: 89; Figs 5-7. [New Caledonia, st. Balabio, 20°03.6'S 

164°07.7'E, 13 m] 

Leptochela (Leptochela) gracilis Stimpson, 1860a 

= Leptochela gracilis Stimpson, 1860a: 42. [Sinu ”Kagosima”, in profundis] 

= Leptochela pellucida Boone, 1935: 105; Plates 26-27. [Near Equator, to the South Brother’s Island, 

south entrance of Durian Straits, Dutch East Indies, 29'N by E 104°47'E, 14 fms] 

Leptochela (Leptochela) hawaiiensis Chace, 1976: 15; Figs 11-13. [Hawaii, south coast of Oahu, Diamond 

Head Light, 62°S 3.9'E] 

Leptochela (Leptochela) irrobusta Chace, 1976: 19; Figs 14-18. [Bikini Atoll, northeast end of lagoon at 

Bowditch anchorage, surface light at night]


Leptochela (Leptochela) japonica Hayashi & Miyake, 1969 

= Leptochela japonica Hayashi & Miyake, 1969: 1; Figs 1-2. [Chijiwa Bay, northwestern Kyushu, 65-66 m] 

Leptochela (Leptochela) papulata Chace, 1976: 26; Figs 22-24. [east of Cape Lookout, North Carolina, 

34°35'30"N 75°45'30"W, 59 m] 

Leptochela (Leptochela) pugnax De Man, 1916 

= Leptochela pugnax De Man, 1916: 148. [Bay of Bima, near south fort, 55 m; Tual-anchorage, Kei-islands, 

22 m; Elat, west coast of Great-Kei-island, 27 m] 

Leptochela (Leptochela) robusta Stimpson, 1860a 

= Leptochela robusta Stimpson, 1860a: 43. [Mari sinensi, prof. 20 org; prope insulam ”Loo Choo” 

quoque] 

Leptochela (Leptochela) serratorbita Spence Bate, 1888 (Fig. 12) 

= Leptochela serratorbita Spence Bate, 1888: 859; Plate 139, fig. 1. [Saint Thomas, Virgin Islands, shallow 

water] 

Leptochela (Leptochela) sydniensis Dakin & Colefax, 1940 

= Leptochela sydniensis Dakin & Colefax, 1940: 153; Figs 245-246. [New South Wales] 

? = Leptochela hainanensis Yu, 1936: 87; Figs 1-3. [Hai-kiu-sche] 

Leptochela (Proboloura) carinata Ortmann, 1893 

= Leptochela carinata Ortmann, 1893: 41 (partim); Plate 4, fig. 1. [Plankton-Expedition der Humboldt- 

Stiftung, Küstenbank vor der Tocantins-Mündung, JN 236 (0.1°S 45.2°W), 50-100 m] 

Leptochela (Proboloura) soelae Hanamura, 1987: 21; Figs 6-8. [north-west Australian shelf, 19°05.0'S 

118°53.8'E, 82 m] 

Parapasiphae Smith, 1884 

= Parapasiphaë Smith, 1884 (type species Parapasiphaë sulcatifrons Smith, 1884, designated by Fowler, 

1912, gender feminine; name placed in its corrected spelling on the Official List of Generic Names 

in Zoology in Opinion 470 in 1957) 

= Orphania Spence Bate, 1888 (type species Orphania tenuimana Spence Bate, 1888, a junior subjective 

synonym of Parapasiphaë sulcatifrons Smith, 1884, gender feminine) 

= Dantecia Caullery, 1896 (type species Dantecia Caudani Caullery, 1896, a subjective synonym of 

Parapasiphaë sulcatifrons Smith, 1884, by monotypy, gender feminine) 

Parapasiphae compta Smith, 1884 

= Parapasiphaë compta Smith, 1884: 389. [Albatross stn 2039, 38°19'26"N 68°20'20"W, 2369 fms] 

Fig. 12. Leptochela (Leptochela) serratorbita Spence Bate, 1888. Photo by Arthur Anker.


Parapasiphae cristata Smith, 1884 

= Parapasiphaë cristata Smith, 1884: 388; Plate 5, fig. 3. [Albatross stn 2100, 39°22'N 68°34'30"W, 1628 

fms] 

= Parapasiphaë macrodactyla Chace, 1939: 33. [S.S.E. of Bahia de Guantanamo, Oriente Province, 

Cuba, 19°40'N 75°03'W, 1920 fms] 

Parapasiphae kensleyi Wasmer, 2005: 165; Figs 1-3. [31°N 159°W, 4200 m] 

Parapasiphae sulcatifrons Smith, 1884 

= Parapasiphaë sulcatifrons Smith, 1884: 384; Plate 5, fig. 4; Plate 6, figs 1-7. [Albatross stns 2002, 

37°02'42''N 74°17'36''W, 641 fms; 2034, 39°27'10''N 69°56'20''W, 1346 fms; 2037, 38°53'00''N 69°23'30''W, 

1731 fms; 2072, 41°53'00''N 65°35'00''W, 858 fms; 2074, 41°43'00''N 65°21'50''W, 1309 fms; 2094, 

39°44'30''N 71°04'00''W, 1022 fms; 2097, 37°56'20''N 70°57'30''W, 1917 fms; 2099, 37°12'20''N 

69°39'00''W, 2949 fms; 2105, 37°50'00''N 73°03'50''W, 1395 fms; 2110, 35°12'10''N 74°57'15''W, 516 fms] 

= Orphania tenuimana Spence Bate, 1888: 872; Plate 141, fig. 4. [Challenger stn 45, 38°34'N 72°10'W, 

south-east of New York, 1240 fms] 

= Dantecia Caudani Caullery, 1896: 372, Plate 14, figs 1-11. [Golfe de Gascogne, 46°28'N 7°W, 1710 m] 

= Pasiphaea metriomma Dohrn, 1908: 9. [Valdivia Expedition stn 26 (ca. 1000 mls E of Madeira, 31°59'N 

5°5'W, 1800 m)] 

Pasiphaea Savigny, 1816 

= Pasiphaea Savigny, 1816 (type species A.[lpheus] sivado Risso, 1816, by monotypy, gender feminine) 

= Pasiphaë (Phye) Wood-Mason, 1892 (type species Pasiphaë (Phye) Parapasiphaë Alcocki Wood-Mason 

& Alcock, 1891b, by monotypy, gender feminine) 

Pasiphaea acutifrons Spence Bate, 1888 

= Pasiphæa acutifrons Spence Bate, 1888: 871; Plate 141, fig. 3. [Challenger stns 311, 52°45'30''S 

73°46'0''W, off Port Churruca, Patagonia, 245 fms; 236, 34°58'N 139°29'E, south of Japan, 77 fms] 

= Pasiphæa forceps A. Milne-Edwards, 1891: 51; Plate 6, figs 2-2e. [détroit de Magellan, au sud-est de 

Port-Famine, 326 m] 

Pasiphaea affinis Rathbun, 1902a 

= Pasiphæa affinis Rathbun, 1902a: 905. [near Cortez Bank, California, 984 fms] 

Pasiphaea alcocki Wood-Mason & Alcock, 1891b 

= Parapasiphaë Alcocki Wood-Mason & Alcock, 1891b: 196. [Bay of Bengal, 16°11'15"N 82°30'30"W, 

922 fms] 

Pasiphaea americana Faxon, 1893 

= Pasiphaeia (sic) cristata americana Faxon, 1893: 208. [Gulf of Panama, 7°32'36"N 79°16'W, 523 m; 

lectotype designation by Hayashi, 2004] 

Pasiphaea amplidens Spence Bate, 1888 

= Pasiphæa amplidens Spence Bate, 1888: 870; Plate 141, fig. 2. [Challenger stn 34°58'N 139°29'E, 775 

fms] 

= Pasiphaea vereschhaka Burukovsky, 1993b: 33; Fig. 1(1-7). [12°22'S 53°02'E, 1045-1050 m] 

Pasiphaea arabica Timofeev, 1997: 142; Fig. 1. [Arabian Sea, 16°32'2''N 64°25'5''E, collecting horizon 110- 

100 m, bottom at 3600 m] 

Pasiphaea balssi Burukovsky & Romensky, 1987: 52; Figs 1, 2(9), 4. [South Atlantic, 41°55'S, 0°00'E, 910 m] 

Pasiphaea barnardi Yaldwyn, 1971: 86 [Cook Strait, 41°42.5'S 175°9'E, ca. 500 fms] 

Pasiphaea berentsae Kensley, Tranter & Griffin, 1987: 294; Figs 7-9. [New South Wales, northeast of Norah 

Head, 33°11'S 152°24'E to 33°09'S 152°25'E, 720 m] 

Pasiphaea burukovskyi Wasmer, 1993: 79; Figs 22-24. [40°22'S 168°25'E to 40°26'S 168°30'E, 600-750 m] 

Pasiphaea chacei Yaldwyn, 1962: 18; Figs 1-17. [33°36'11"N 118°32'13"W – 33°30'52"N 118°22'48"W, 475 

fms] 

Pasiphaea corteziana Rathbun, 1902a 

= Pasiphæa corteziana Rathbun, 1902a: 905. [near Cortez Bank, California, 776 fms] 

Pasiphaea cristata Spence Bate, 1888 

= Pasiphæa cristata Spence Bate, 1888: 865; Plate 140; Plate 141, fig. 1. [Challenger stn 173, 19°9'35"S 

179°41'50"E, off Matuka, Fiji Islands, 315 fms]


Pasiphaea crosnieri Hayashi, 2004: 334; Figs 7-9. [Philippines, 13°40.7'N 120°30.0'E, 685-757 m] 

Pasiphaea debitusae Hayashi, 1999: 281; Figs 8-10. [Indonesia, 05°15'S 133°01'E, 605-576 m] 

Pasiphaea diaphana Burukovsky & Romensky, 1980: 1097. [South-East Atlantic, 31°48'S 02°18'E, 1100 m] 

Pasiphaea dofleini Schmitt, 1932a: 333; Fig. 1. [Punta Arenas, now Magelhanes, Chile] 

Pasiphaea ecarina Crosnier, 1969: 537; Figs 19, 23-36. [Congo, 5°08'S 11°22'E, 795-805 m] 

Pasiphaea emarginata Rathbun, 1902a 

= Pasiphæa emarginata Rathbun, 1902a: 905. [Gulf of California, 857 fms] 

Pasiphaea faxoni Rathbun, 1904: 22; nomen novum for Pasiphaea acutifrons sensu Faxon, 1895 nec Spence 

Bate, 1888 [Albatross stns 3402 (0°57'30"S 89°03'30"W), 421 fms; 3403 (0°58'30" 89°17'00"W) 384 fms 

and 3406 (0°16'00"S 90°21'30"W), 551 fms] 

Pasiphaea flagellata Rathbun, 1906 

= Pasiphæa flagellata Rathbun, 1906: 928; Fig. 78; Plate 23, fig. 2. [Kaiwi Channel, Hawaii, 411-442 

fms] 

Pasiphaea fragilis Hayashi, 1999: 285; Fig. 11. [Loyalty Islands, 20°23.0'N 166°20.1'E, 780 m] 

Pasiphaea gelasinus Hayashi & Yaldwyn, 1998: 512; Figs 1-3. [South Indian Ocean, Seamounts off Saint 

Paul and Amsterdam Islands, 37°37.8'S, 77°51.8'E, 730-905 m] 

Pasiphaea gracilis Hayashi, 1999: 290; Figs 15-17. [New Caledonia, 23°08'S 166°51'E, 850 m] 

Pasiphaea grandicula Burukovsky, 1976: 17; Fig. 1. [South Atlantic, 26°11'S, 06°02'8"E, 1150 m] 

Pasiphaea hoplocerca Chace, 1940: 124; Figs 4-5. [Bermuda Oceanographic Expedition Nets 713, 753 and 

778 (32°12'N 64°36'W), 700 fms] 

Pasiphaea japonica Omori, 1976: 250; Figs 1-2. [Off Iwase, Toyama Bay, 0-300 m] 

Pasiphaea kapala Kensley, Tranter & Griffin, 1987: 297; Figs 10-12. [New South Wales, northeast of Norah 

Head, 33°08'S 152°27'E to 33°10'S 152°24'E, 580 m] 

Pasiphaea kaiwiensis Rathbun, 1906 

= Pasiphæa kaiwiensis Rathbun, 1906: 927; Fig. 76; Plate 23, fig. 4. [Kaiwi Channel, 343-337 fms] 

Pasiphaea korzuni Burukovsky, 1995: 121; Figs 1-5. [Aden Bay, near Ras-Fartak Island, depth unknown] 

Pasiphaea laevis Hayashi, 1999: 287; Figs 12-14. [Indonesia, Moluccas, 06°08'S 132°45'E, 390-502 m] 

Pasiphaea ledoyeri Hayashi, 2006a: 215; Figs 6-8. [South Indian Ocean, 46°33.8'S 41°23.7'E, 660-911 m, in 

stomach of Dissostichus eleginoides] 

Pasiphaea levicarinata Hanamura, 1994: 167; Figs 1-2. [north-western Australia, 18°30'S 117°23'E, 530 m] 

Pasiphaea liocerca Chace, 1940: 122; Figs 2-3. [Bermuda Oceanographic Expeditions Net 1160 (32°12'N 

64°36'W), 800 fms] 

Pasiphaea longitaenia Kensley, Tranter & Griffin, 1987: 301; Figs 13-14. [New South Wales, east of Long 

Reef Point, 33°43'S 151°56'E to 33°39'S 151°58'E, 626-637 m] 

Pasiphaea magna Faxon, 1893: 209. [Albatross stn 3384 (off Panama, 7°31'30"N, 79°15'00"W), 458 fms] 

Pasiphaea major Hayashi, 2006a: 219; Figs 9-11. [Norfolk Ridge, 25°09'S 168°53'E, 1173-1340 m] 

Pasiphaea marisrubri Iwasaki, 1989: 178; Figs 1-3. [Central Red Sea, 21°14.80'N 37°15.40'E – 21°18.20'N 

37°12.00'E, 0-220 m] 

Pasiphaea mclaughlinae Hayashi, 2006b: 342; Figs 1-3. [SE coast of Taiwan, 22°13.8'N 121°01.8'E, 1222-1226 m] 

Pasiphaea meiringnaudei Kensley, 1977: 34-37; Figs 11-14. [Meiring Naude stn 107 (South Africa, off Natal 

coast, 28°37.8'S 32°38.4'E, 1200-1000 m)] 

Pasiphaea merriami Schmitt, 1931b 

= Pasiphæa merriami Schmitt, 1931b: 391. [south of the Dry Tortugas, 253-283 fms] 

= Pasiphaea nishiei Iwasaki, 1990: 190; Figs 1-2. [off Jamaica, 17°34.3'N, 76°03.3'W, 1490-1504 m] 

Pasiphaea multidentata Esmark, 1866: 259. [Bjørumsfjorden omtrent 1/8 Miil fra Namsos] 

= P.[asiphaë] norvegica M. Sars, 1866a: 260. [nomen nudum] 

= Pasiphaë norvegica M. Sars, 1866b: 314. [Christianiafjorden; fully illustrated in M. Sars, 1868] 

= Pasiphaë (Phye) sicula Riggio, 1896 [in Riggio, 1895-1896]: 41; Plate 1 (1865, no legend), Figs 2a-b. 

[proviene dal mare di Augusta] 

Pasiphaea natalensis Burukovsky & Romensky, 1982: 1797; Figs 1-6. [south-western part of the Indian 

Ocean, 33°48.5'S 44°30.8'E, 900-910 m] 

Pasiphaea notosivado Yaldwyn, 1971: 86. [Cook Strait, 41°32'S 174°56'E, 50-100 fms] 

Pasiphaea orientalis Schmitt, 1931a: 267; Plate 32, figs 1, 5. [Ryukyusho, Takao, Formosa]


Pasiphaea oshoroae Komai & Amaoka, 1993: 367; Figs 1-3. [Off Aleuten Islands, 49°59.9'N 176°55.4'W, 

0-400 m] 

Pasiphaea pacifica Rathbun, 1902a 

= Pasiphæa pacifica Rathbun, 1902a: 905. [off Point Sur, California, 328 fms] 

Pasiphaea philippinensis Hayashi, 1999: 277; Figs 5-7. [Philippines, 13°39'N 120°43'E, 520-550 m] 

Pasiphaea planidorsalis Hayashi, 2004: 348; Figs 13-14. [New Caledonia, 19°42.4'S 158°50.8'E, 772-756 m] 

Pasiphaea poeyi Chace, 1939: 31. [Bahia de Cochinos, Santa Clara Province, Cuba, 22°07'N 81°08'W, 220- 

275 fms] 

Pasiphaea princeps Smith, 1884 

= Pasiphaë princeps Smith, 1884: 381; Plate 5, fig. 2. [Albatross stn 2095, 39°29'N 70°58'40"W, 1451 fms] 

Pasiphaea propinqua De Man, 1916: 147. [06°11'N 120°37.5'E, off the Sulu Islands, 450 m] 

Pasiphaea pseudacantha Hayashi, 2004: 353; Figs 16-18. [Kai Islands, Indonesia, 06°08'S 132°45'E, 390-502 m] 

Pasiphaea rathbunae (Stebbing, 1914a) 

= Phye rathbunæ Stebbing, 1914a: 295; Plate 31. [Scotia stn 450, 48°00'S 9°50'W, 1332 fms] 

Pasiphaea romenskyi Burukovsky, 1995: 123; Figs 6-11. [Ecuador Seamount, 00°23'S, 56°02'E, 200-235 m] 

Pasiphaea scotiae (Stebbing, 1914a) 

= Phye scotiæ Stebbing, 1914a: 294; Plate 30. [Scotia stns 417, 71°22'S 16°34'W, 1410 fms; 422, 68°32'S 

12°49'W, from surface to 600 fms] 

= Pasiphaea longispina Lenz & Strunck, 1914: 315; Plate 19, figs 1-11. [65°15'S 80°19'E, also in der Nähe 

des Eisrandes, im Magen eines Pinguins, Aptenodytes forsteri, gefunden] 

Pasiphaea semispinosa Holthuis, 1951a: 9; Fig. 1. [off Angola, 7°55'S 12°38'E, 235-460 m] 

Pasiphaea sinensis Hayashi & Miyake, 1971: 39; Fig. 1. [East China Sea, 29°27.2'N 128°16.2'E – 29°33.3'N 

128°23.5'E, 1065-1075 m] 

Pasiphaea sirenkoi Burukovsky, 1987: 37; Figs 1-5. [SE Indian Ocean, 11°27.5'N 109°56.2'E, 600 m] 

Pasiphaea sivado (Risso, 1816) 

= A.[lpheus] sivado Risso, 1816: 93; Plate 3, fig. 4. [sur la plage de Nice] 

= P.[asiphaea] Savignyi H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 426. [Patrie inconnue] 

= Pasiphæa brevirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: Plate 54bis, fig. 2. 

[Type locality not indicated] 

? = Pasiphæa distincta Guérin-Méneville, 1844 [in Guérin-Méneville, 1829-1844]: 16, Plate 22, fig. 3. 

(as Pasiphæa sivado). [la Méditerranée] 

= Pasiphaea neapolitana Hope, 1851: 44. [Type locality not indicated] 

Pasiphaea tarda Krøyer, 1845: 453. [Type locality not indicated] 

= Pasiphæa principalis Sund, 1913: 6; Figs 5-7, 9a-f. [Sognefjord; Øxsund (E. of the Lofoten isles, about 

67.5°N) 

Pasiphaea telacantha Hayashi, 2004: 363; Figs 22-24. [New Caledonia, 23°08'S 166°51'E, 850 m] 

Pasiphaea timofeevi Burukovsky, 1993b: 35; Fig. 1(14-19). [Aden Bay, Socotra Island, 12°20'N 53°09'W, 515- 

480 m] 

Pasiphaea truncata Rathbun, 1906 

= Pasiphæa truncata Rathbun, 1906: 928; Fig. 7; Plate 23, fig. 5. [vicinity of Modu Manu, 293-800 fms] 

Pasiphaea unispinosa Wood-Mason, 1892 

= Pasiphaë unispinosa Wood-Mason, 1892: Plate 3, fig. 7; accompanying text as Wood-Mason in 

Wood-Mason & Alcock, 1893: 163. [7 miles SE by S of Ross Island, Andaman Sea, 265 fms] 

Pasiphaea westindica Tchesunov, 1984: 993; Fig. 1. [Atlantic Ocean, 12°48'N 62°07'W, 1500 m] 

Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893 

= Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893 (type species Psathyrocaris fragilis 

Wood-Mason in Wood-Mason & Alcock, 1893, by monotypy, gender feminine) 

Psathyrocaris fragilis Wood-Mason in Wood-Mason & Alcock, 1893: 171; Plates 10-11. [15°56'50"N 

81°30'30"E, 240 fms] 

= Psathyrocaris infirma var. atlantica Caullery, 1896: 374; Plate 14, figs 12-15. [Golfe de Gascogne, 

44°39'N, 2°10'W, 800 m]


Psathyrocaris hawaiiensis Rathbun, 1906: 928; Fig. 79. [vicinity of Modu Manu, 876 fms] 

Psathyrocaris infirma Alcock & Anderson, 1894: 159. [Investigator stn 116, Andaman Sea, 11°25'5''N 

92°47'6''E, 405 fms] 

Psathyrocaris platyophthalmus Alcock & Anderson, 1894: 158. [Investigator stn 124, Laccadive Sea, 

10°47'45"N 72°40'20"E, 705 fms] 

Psathyrocaris plumosa Alcock & Anderson, 1894: 159. [Investigator stn 128, off Ceylon, 6°58'N 77°26'50''E, 

902 fms] 

Superfamily oPloPHoRoIDeA Dana, 1852a 

Family ACANTHePHyRIDAe Spence Bate, 1888 

Acanthephyra A. Milne-edwards, 1881b (Fig. 13) 

= Ephyra Roux, 1831 (an invalid junior homonym of Ephyra Peron & Leseur, 1810 (Coelenterata), 

type species A.[lpheus] Pelagicus Risso, 1816, designated by Kingsley, 1880, gender feminine; name 

placed on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in 

1955) 

= Miersia Kingsley, 1880 (nomen novum for Ephyra Roux, 1831, gender feminine; name suppressed 

for the purposes of the Principle of Priority but not for those of the Principle of Homonymy, placed 

on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in 1955) 

= Acanthephyra A. Milne-Edwards, 1881b (type species Acanthephyra armata A. Milne-Edwards, 

1881b, by original designation, gender feminine; name placed on the Official List of Generic Names 

in Zoology in Opinion 359 in 1955) 

= Bentheocaris Spence Bate, 1888 (type species Bentheocaris stylorostratis Spence Bate, 1888, designated 

by Holthuis, 1955b, gender feminine) 

= Acanthephyropsis Riggio, 1895 [in Riggio, 1895-1896] (type species Acanthephyra pulchra A. Milne- 

Edwards, 1890, a junior subjective synonym of Acanthephyra eximia Smith, 1884, gender feminine) 

= Hoplocaricyphus Coutière, 1907a (type species Hoplocaricyphus similis Coutière, 1907a, probably a 

subjective synonym of A.[lpheus] Pelagicus Risso, 1816, gender masculine) 

Fig. 13. Acanthephyra spec. Photo by Tin-Yam Chan.


Acanthephyra acanthitelsonis Spence Bate, 1888: 745; Plate 125, fig. 3. [Challenger stns 106, 1°47'N 24°26'W, 

Atlantic, south-west of Sierra Leone, 1850 fms; 107, 1°22'N, 26°36'W, Atlantic, south-west of Sierra 

Leone, 1500 fms] 

Acanthephyra acutifrons Spence Bate, 1888: 749 (partim); Plate 126, fig. 3. [off Kepulauan Aru, Indonesia, 

5°41'S 134°04'30"E, 1463 m; type locality restricted by Kemp, 1906a] 

Acanthephyra armata A. Milne-Edwards, 1881b: 12. [près de Sainte-Lucie, 422 brasses] 

Acanthephyra brevicarinata Hanamura, 1984: 65; Figs 1-2. [13°28.7'N, 119°53.9'W, 1617-0 m] 

Acanthephyra brevirostris Smith, 1885a: 504. [Albatross stns 5448, 37°12'20"N 69°39'00"W, 2949 fms; 5449, 

39°22'00"N, 68°34'30"W, 1686 fms; 5673, 37°50'00"N, 73°03'50"W, 1395 fms] 

= Hymenodora duplex Spence Bate, 1888: 843; Plate 136, fig. 3. [Challenger stn 147, 46°16'S 48°27'E, off 

Marion Island, 1600 fms] 

Acanthephyra carinata Spence Bate, 1888: 748; Plate 126, fig. 2. [Challenger stn 310, 51°27'30"S 74°03'00"W, 

Sarmiento channel, Patagonia, 400 fms] 

= Acanthephyra approxima Spence Bate, 1888: 755; Plate 126, fig. 8. [Challenger stn 310, 51°27'30"S 

74°03'00"W, Sarmiento Channel, Patagonia, 400 fms] 

Acanthephyra chacei Krygier & Forss, 1981: 96; Figs 1-2. [44°45.2'N 127°44.0'W, 0-2600 m] 

Acanthephyra cucullata Faxon, 1893: 206. [Albatross stn 3381 (North Pacific Ocean, Colombia, 4°56'00"N, 

80°52'30"W), 1772 fms] 

Acanthephyra curtirostris Wood-Mason & Alcock, 1891b: 195. [Investigator stns 100 (11°12'47"N 74°25'30"E), 

840 fms; 104 (16°55'41"N 83°21'18"E), 1000 fms] 

= Acanthephyra acutifrons Spence Bate, 1888: 749 (partim) (nec Plate 126, fig. 3) 

Acanthephyra eximia Smith, 1884: 376. [Albatross stn 2111, 35°09'50"N 74°57'40"W, 938 fms] 

= Acanthephyra angusta Spence Bate, 1888: 737; Plate 124, fig. 6. [Challenger stn 194, 4°34'00"S 

129°57'30"E, off Banda Island, 200 fms] 

= Acanthephyra edwardsii Spence Bate, 1888: 747; Plate 126, fig. 1. [Challenger stn 126, 10°46'S 36°08'W, 

south of Pernambuco, 770 fms] 

= Acanthephyra brachytelsonis Spence Bate, 1888: 753; Plate 126, fig. 7. [Challenger stns 170, 29°55'S 

178°14'W, off the Kermadec Islands, 520 fms; 170A, 29°45'S 178°11'W, north of the Kermadec Islands, 

630 fms; 171, 28°33'S 177°50'W, north of the Kermadec Islands, 600 fms; 194, 4°34'N 

129°57'30''E, off Banda Island, 200 fms; 214, 4°33'N 127°6'E, south of the Philippine Islands, 500 fms; 

232, 35°11'N 139°28'E, Japan, 345 fms; 236, 34°58'N, 139°29'E, south of Japan, 775 fms; 318, 42°32'S 

56°29'W, north of the Falkland Islands, 2040 fms] 

= Acanthephyra pulchra A. Milne-Edwards, 1890: 163. [au large de Monaco, 1650 m] 

= Acanthephyra frontieri Crosnier, 1987b: 699; Fig. 1. [Madagascar, 13°02'S 48°02'E, 1000-1525 m] 

Acanthephyra faxoni Calman, 1939 

= Acanthephyra Faxoni Calman, 1939: 191; Fig. 1. [John Murray Expedition stn 184, Gulf of Aden 

(14°36'06''N 51°00'18''E - 14°38'42''N 50°57'42''E), 1270 m] 

Acanthephyra fimbriata Alcock & Anderson, 1894 

= Acanthephyra armata var. fimbriata Alcock & Anderson, 1894: 156. [Bay of Bengal [off Madras, 

12°50'N 81°30'E], 475 fms; off the Malabar coast [off Goa, 15°29'N 72°41'E], 559 fms; illustrated in 

Wood-Mason, 1892: Plate 3, fig. 1. (as Acanthephyra armata var.)] 

Acanthephyra indica Balss, 1925a: 264; Figs 34-35. [Valdivia stn 215, 7°01'N 85°56'E, 0-2500 m, zwischen 

Sumatra und Ceylon] 

Acanthephyra kingsleyi Spence Bate, 1888: 751; Plate 126, fig. 4. [Challenger stn 104, 2°25'N 20°1'W, Atlantic, 

south-west of Sierra Leone, 2500 fms] 

= Acanthephyra sexspinosa Kemp, 1939: 575. [Central and South Atlantic from 17°N to 18°S] 

Acanthephyra media Spence Bate, 1888: 736; Plate 124, fig. 5. [Challenger stn 207, 12°21'N 122°15'E, off 

Tablas Island, 700 fms] 

= Acanthephyra media var. obliquirostris De Man, 1916: 150. [0°17.6'S 129°14.5'E, Halmaheira Sea, 1855 m] 

Acanthephyra pelagica (Risso, 1816) 

= A.[lpheus] Pelagicus Risso, 1816: 91; Plate 2, fig. 7. [la mer de Nice; neotype listing by Fransen, 

Holthuis & Adema, 1997 not valid] 

= Ephyra Haeckelii von Martens, 1868: 52; Plate 1, Figs 7a-b. [Messina]


= Acanthephyra Agassizzi mediterranea Riggio, 1900: 20. [Mare di Messina] 

= Acanthephyra rectirostris Riggio, 1900: 20. [Mare di Messina] 

= Acanthephyra purpurea var. multispina Coutière, 1905a: 10. [Princesse-Alice stns 1797 (32°18'N 

23°58'W), 0-2000 m; 1851 (46°15'N 7°09'W), 0-3000 m; 1991 (42°53'N 8°22'E), 0-2000 m; 1583 (47°36'N 

7°38'W, 1490 m) and 1639 (36°17'N 28°53'W, 0-3000 m)] 

= Acanthephyra parva multidens Coutière, 1905a: 15; Fig. 5. [Princesse-Alice stns 1856 (36°46'N 

26°41'W), 0-3250 m; 1834 (37°28'N 25°53'30"W), 0-1000 m] 

Acanthephyra prionota Foxton, 1971: 35; Figs 1-2. [East African coast, 05°39'S 46°22'E, 1900-1850-(0) m] 

Acanthephyra purpurea A. Milne-Edwards, 1881a: 933. [au large des îles Berlingues par 2.590 mètres] 

= Miersia Agassizii Smith, 1882: 67; Plate 11, Figs 5-7; Plate 12, Figs 1-4. [Blake stns 305, 41°23'15''N 

65°51'25''W, 810 fms; 323, 33°19'0''N 76°12'30''W, 457 fms; 330, 31°41'0''N 74°35'0''W, 1047 fms] 

= Acanthephyra abyssorum Filhol, 1885b: Plate 7. [Type locality not indicated, 4000 m] 

= Acanthephyra parva paucidens Coutière, 1905a: 16; Fig. 5(3). [Princesse-Alice stn 1856 (36°17'N, 

28°53'W), 0-3250 m] 

= Hoplocaricyphus similis Coutière, 1907a: 7; Fig. 1. [28°04'N, 16°49'30"W, 0-1000 m] 

Acanthephyra quadrispinosa Kemp, 1939: 576. [Indo-Pacific from the E. African coast to 163°W, and from 

25°N to 42°S; South Atlantic from 32°S to 40°S] 

= Acanthephyra batei Stebbing, 1905: 107; Plate 24B; nec Faxon, 1895. [Cape Point Lighthouse, S 83°E, 

35.5 miles, 360 fms] 

Acanthephyra rostrata (Spence Bate, 1888) 

= Hymenodora rostrata Spence Bate, 1888: 846; Plate 136, fig. 4. [Challenger stns 184, 12°8'S 145°10'E, 

near Torres Strait, 1400 fms; 205, 16°42'N 119°22'S, Philippine Islands, 1050 fms; 245, 36°23'N 

174°31'E, North Pacific, 2775 fms] 

Acanthephyra sanguinea Wood-Mason in Wood-Mason & Alcock, 1892: 358. [Laccadive Sea, 9°53'34''N 

75°16.5'E, 1091 fms; Laccadive Sea, 8°23'N 75°47'E, 738 fms; Bay of Bengal, 12°58'N 88°52'17''E, 

1748 fms] 

= Acanthephyra kempii Balss, 1914a: 595. [70°01'N 85°56'E, 0-2500 m] 

Acanthephyra sibogae De Man, 1916 

= Acanthephyra (Meningodora) sibogae De Man, 1916: 149. [5°26'S 121°18'E, entrance of Gulf of Boni, 

1944 m] 

Acanthephyra sica Spence Bate, 1888: 739; Plate 125, fig. 1. [Challenger stns 168, 40°28'S 177°43'E, off 

New Zealand, 100 fms; 40, 34°51'N 68°30'W, north-west of Bermuda, 2675 fms; 159, 47°25'S 

130°22'E, south of Australia, 2150 fms; 169, 37°34'S 179°22'E, near New Zealand, 700 fms; 170, 

29°55'S 178°14'W, off the Kermadec Islands, 520 fms; 170A, 29°45'S 178°11'W, near the Kermadec 

Islands, 630 fms; 181, 13°50'S 151°49'E, between Australia and the Solomon Islands, 2440 fms; 194, 

4°34'0''S 129°57'30''E, off Banda Island, 200 fms; 230, 26°29'N 137°57'E, south of Japan, 2425 fms; 

235, 34°7'N 138°0'E, south of Japan, 565 fms; 318, 42°32'S 56°29'W, north of the Falkland Islands, 

2040 fms] 

Acanthephyra smithi Kemp, 1939: 577. [Indo-Pacific from the East African coast to mid-Pacific in 131°W; 

at the western end of its range it extends to 14°S and at the eastern end from 20°N to 24°S] 

Acanthephyra stylorostratis (Spence Bate, 1888) 

= Bentheocaris stylorostratis Spence Bate, 1888: 726; Fig. 72; Plate 123, fig. 4. [Challenger stn 13, 21°38'N 

44°39'W, Mid North Atlantic, 1900 fms] 

? = Bentheocaris exuens Spence Bate, 1888: 724; Plate 123, fig. 3. [Challenger stn 285, 32°36'S 137°43'W, 

South Pacific Ocean, 2357 fms] 

Acanthephyra tenuipes (Spence Bate, 1888) 

= Tropiocaris tenuipes Spence Bate, 1888: 836; Plate 136, fig. 2. [Challenger stn 184, 12°8'S 145°10'E, 1400 

fms] 

= Acanthephyra gracilipes Chace, 1940: 149; Figs 26-27. [Bermuda Oceanographic Expedition Net 689 

or 734 (approx. 32°12'N 64°36'W), 1000 fms] 

Acanthephyra trispinosa Kemp, 1939: 577. [west coast of Central America from 7°N to 4°S, extending 

westwards to 116°W]


Ephyrina Smith, 1885a 

= Ephyrina Smith, 1885a (type species Ephyrina Benedicti Smith, 1885a, by monotypy, gender feminine) 

= Calymarina Spence Bate, 1888 (nomen nudum, gender feminine) 

= Tropirinus Spence Bate, 1888 (nomen nudum, gender masculine) 

= Tropiocaris Spence Bate, 1888 (type species Tropiocaris planipes Spence Bate, 1888, a junior subjective 

synonym of Ephyrina Benedicti Smith, 1885a, gender feminine) 

Ephyrina benedicti Smith, 1885a 

= Ephyrina Benedicti Smith, 1885a: 506. [Albatross stn 2083, 40°26'40''N 67°5'15''W, 959 fms] 

= Tropiocaris planipes Spence Bate, 1888: 835; Plate 136, fig. 1. [Challenger stn 230, 26°29'N 137°57'E, 

2425 fms] 

Ephyrina bifida Stephensen, 1923: 58; Fig. 18. [45°37'N 7°03'W, 4300 m of water] 

Ephyrina childressi Chace, 1986: 34; Fig. 17. [Halmahera Sea, Indonesia, 0°33'42"S 128°52'06"E, 950-1200 m] 

Ephyrina figueirai figueirai Crosnier & Forest, 1973 

= Ephyrina figueirai Crosnier & Forest, 1973: 73; Figs 20b, 21g-h, 22c-d, 23. [Bay of Biscay, 47°12'30"N 

6°48'W, 2120-2350 m] 

Ephyrina figueirai spinicauda Lin & Chan, 2001: 184; Figs 1, 2a, d-e, 3. [N.E. Taiwan, I-Lan County, fishing 

port at Tai-Shi, about 600 m] 

Ephyrina hoskynii Wood-Mason & Alcock, 1891b 

= Ephyrina Hoskynii Wood-Mason & Alcock, 1891b: 194. [Investigator stn 105 (Arabian Sea, 15°2'N 

72°34'E), 740 fms] 

Ephyrina ombango Crosnier & Forest, 1973: 68; Figs 20a, 21a-f, 22a-b. [off São Tomé, Gulf of Guinea, 

0°30'N 6°30'E, 0-1000 m in total depth of 2900 m] 

Heterogenys Chace, 1986 

= Heterogenys Chace, 1986 (type species Acanthephyra microphthalma Smith, 1885a, by original designation 


Heterogenys microphthalma (Smith, 1885a) 

= Acanthephyra microphthalma Smith, 1885a: 502. [Albatross stn 2224, 36°16'30"N 68°21'00"W, 2574 fms] 

= Acanthephyra longidens Spence Bate, 1888: 735; Plate 124, fig. 4. [Challenger stns 198, 2°55'N 124°53'E, 

near the Philippine Islands, 2150 fms; 285, 32°36'S 137°43'W, South Pacific Ocean, 2375 fms] 

Heterogenys monnioti Crosnier, 1987b: 704; Fig. 3. [29°50.9'S 48°35.5'E, 3668-3800 m] 

Hymenodora G.o. Sars, 1877 

= Hymenodora G.O. Sars, 1877 (type species Pasiphaë glacialis Buchholz, 1874, by monotypy, gender 

feminine) 

Hymenodora acanthitelsonis Wasmer, 1972: 87; Figs 1-8. [45°18.0'N 125°43.2'W to 45°17.2'N 125°48.3'W, 

0-2560 m] 

Hymenodora frontalis Rathbun, 1902a: 904. [west of Unalaska, 322 fms] 

Hymenodora glacialis (Buchholz, 1874) 

= Pasiphaë glacialis Buchholz, 1874: 279; Plate 1, Figs 2-2g. [an der Oberfläche des Meeres, also in der 

Nähe des 74.° nördl. Br. in beträchtlicher Entfernung von der Grenze des Packeises] 

= Hymenodora glauca Spence Bate, 1888: 847; Plate 137, fig. 1. [Challenger stns 159, 47°25' 130°22'E, 

south of Australia, 2150 fms; 215, 4°19'N 130°15'E, south of the Philippines, 2550 fms] 

= Hymenodora mollicutis Spence Bate, 1888: 848; Plate 137, fig. 2. [Challenger stns 104, 2°25'N 20°1'W, 

Atlantic, 2500 fms; 87, 25°49'N 20°55'W, near the Canary Islands, 1675 fms; 133, 35°41'S 20°55'W, 

near Tristan da Cunha, 1900 fms; 156, 62°26'S 95°44'S, 1975 fms; 157, 53°55'S 108°35'S (sic), 1950 fms; 

318, 42°32'S 56°29'W, 2040 fms] 

Hymenodora gracilis Smith, 1886a: 680; Plate 12, fig. 6. [Albatross stns 2036, 38°52'40''N 69°24'40''W, 1735 

fms; 2083, 40°26'40''N 67°05'15''W, 959 fms; 2095, 39°29'00''N 70°58'40''W, 1342 fms; 2099, 37°12'20''N 

69°36'00''W, 2949 fms; 2100, 39°22'00''N 68°34'430''W, 1628 fms; 2101, 39°18'30''N 68°24'00''W, 1686 


70°10'30''W, 1122 fms]


Kemphyra Chace, 1986 

= Kemphyra Chace, 1986 (type species Notostomus corallinus A. Milne-Edwards, 1883, by original 


Kemphyra corallina (A. Milne-Edwards, 1883) 

= Notostomus corallinus A. Milne-Edwards, 1883: Plate 32. [Travailleur dragage 20 (41°30'N 11°40'W), 

2080 m] 

= Acanthephyra valdiviae Balss, 1914a: 595. [30°6'S 87°50'E, 0-2070 m] 

Meningodora Smith, 1882 

= Meningodora Smith, 1882 (type species Meningodora mollis Smith, 1882, by monotypy, gender feminine) 

Meningodora compsa (Chace, 1940) 

= Notostomus compsus Chace, 1940: 156; Figs 31-32. [Bermuda Oceanographic Expedition Net 748 

(32°12'N 64°36'W), 900 fms] 

Meningodora longisulca Kikuchi, 1985: 191; Figs 1-14. [23°08.1'N 149°33.8'E, 0-975 m] 

Meningodora marptocheles (Chace, 1940) 

= Notostomus marptocheles Chace, 1940: 158; Figs 33-34. [N.E Providence Channel, Bahamas, 25°29'N 

77°18'W, 875 fms of wire] 

Meningodora miccyla (Chace, 1940) 

= Notostomus miccylus Chace, 1940: 161; Figs 35-36. [north of Little Inagua Island, Bahamas, 21°44'N 

72°43'W, 1167 fms of wire] 

Meningodora mollis Smith, 1882: 74; Plate 11, Figs 8-9; Plate 12, Figs 5-9. [Blake stn 328, 34°28'25"N 

75°22'50"W, 1632 fms] 

= Notostomus fragilis Faxon, 1893: 207. [Albatross stn 3371 (North Pacific Ocean, Costa Rica, 5°26'20"N, 

86°55'00"W), 770 fms] 

Meningodora vesca (Smith, 1886a) 

= Notostomus viscus Smith, 1886b: 189. [nomen nudum] 

= Notostomus vescus Smith, 1886a: 676; Plate 12, fig. 5. [Albatross stn 2099, 37°12'20"N 69°39'00"W, 

2949 fms] 

= Acanthephyra brevirostris Spence Bate, 1888: 751; Plate 126, Figs 5-6; nec Smith, 1885a. [Challenger 

stn 107, 1°22'N 26°36'W, Atlantic, south-west of Sierra Leone, 1500 fms] 

= Acanthephyra batei Faxon, 1895: 167. [nomen novum for A. brevirostris Spence Bate, 1888] 

= Acanthephyra parvirostris Coutière, 1911a: 157. [erroneous spelling of A. brevirostris] 

Notostomus A. Milne-edwards, 1881b 

= Notostomus A. Milne-Edwards, 1881b (type species Notostomus gibbosus A. Milne-Edwards, 1881b, 

by monotypy, gender masculine) 

Notostomus auriculatus Barnard, 1950: 670; Fig. 124h-i. [off Cape Point, 800 fms] 

Notostomus crosnieri Macpherson, 1984: 54; Figs 6a, 7a, 8a-b. [02°33.5'S, 05°43.2'E, 4088 m] 

Notostomus distirus Chace, 1940: 166; Figs 39-40. [Bermuda Oceanographic Expedition Net 1281 (32°12'N 

64°36'W), 1000 fms] 

Notostomus elegans A. Milne-Edwards, 1881b: 8. [la mer des Antilles, 24°36'N 84°05'W, 955 brasses] 

= Notostomus patentissimus Spence Bate, 1888: 826; Plate 133; Plate 134, Figs 1-2. [Challenger stn 198, 

2°55'N 124°53'E, south of the Philippine Islands, 2150 fms] 

= Notostomus longirostris Spence Bate, 1888: 833; Plate 135; fig. 4. [Challenger stn 195, 4°21'S 129°7'E, 

off Banda Island, 1425 fms] 

= Notostomus westergreni Faxon, 1893: 208. [Albatross stn 3399 (North Pacific Ocean, Ecuador, 1°07'N, 

81°04'W), 1740 fms] 

= Notostomus atlanticus Lenz & Strunck, 1914: 330. [20°41'N 31°53'W, 3000 m, Westich von den Kapverden 

und Kanarischen Inseln] 

Notostomus gibbosus A. Milne-Edwards, 1881b: 7. [Grenade, 626 brasses] 

= Notostomus perlatus Spence Bate, 1888: 831; Plate 135, fig. 2. [Challenger stn 198, 2°55'N 124°53'E, 

near the Philippine Islands, 2150 fms]


= Notostomus brevirostris Spence Bate, 1888: 832; Plate 135, fig. 3. [Challenger stn 120, 8°37'S 34°28'W, 

off Pernambuco, 675 fms] 

Notostomus japonicus Spence Bate, 1888: 830; Plate 135, fig. 1. [Challenger stn 235, 34°7'N 138°0'E, south of 

Japan, 565 fms] 

Notostomus murrayi Spence Bate, 1888: 829; Plate 134, fig. 3. [Challenger stn 133, 35°41'S 20°55'W, near 

Tristan da Cunha, 1900 fms] 

Notostomus robustus Smith, 1884: 377; Plate 7, fig. 2. [Albatross stns 2042, 39°33'N 68°26'45"W, 1555 fms; 

2074, 41°43'N 65°21'50"W, 1309 fms] 

= Notostomus beebei Boone, 1930a: 39; Fig. 6. [five miles south of Nonsuch Island, Bermuda, 32°16'N 

64°36'W, 900 fms] 

Notostomus sparsidenticulatus Wasmer, 1986: 44; Fig. 9. [44°40'S 145°26'W, 2050-0 m] 

Family oPloPHoRIDAe Dana, 1852a 

Janicella Chace, 1986 

= Janicella Chace, 1986 (type species Oplophorus spinicauda A. Milne-Edwards, 1883, by original designation 


Janicella spinicauda (A. Milne-Edwards, 1883) 

= Oplophorus spinicauda A. Milne-Edwards, 1883: Plate 30. [off Casablanca, Morocco, 34°13'30"N 

7°43'W, 636 m] 

= Oplophorus foliaceus Rathbun, 1906: 922; Fig. 72; Plate 20, fig. 8. [Kaiwi Channel, 337-442 fms] 

= Acanthephyra anomala Boone, 1927: 104; Fig. 21. [north of Glover Reef, 484 fms] 

Oplophorus H. Milne edwards, 1837 [in H. Milne edwards, 1834-1840] 

= Oplophorus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species O.[plophorus] 

typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy, gender masculine) 

Oplophorus gracilirostris A. Milne-Edwards, 1881b: 6. [la Dominique, 118 brasses] (Fig. 14) 

= Oplophorus longirostris Spence Bate, 1888: 765; Plate 127, fig. 2. [Challenger stn 174c, 19°07'50"S 

178°19'35"E, off Kandavu, Fiji Islands, 610 fms] 

Fig. 14. Oplophorus gracilirostris A. Milne-Edwards, 1881. Photo by Tin-Yam Chan.


= Hoplophorus Smithii Wood-Mason & Alcock, 1891b: 194 [Investigator stns 62 (16°44'30"N, 88°19-32'E), 

1439 fms; 103 (15°15'N, 81°09'E), 1260 fms] 

= Oplophorus okitsuensis Yokoya, 1922: 302. [Okitsu, Suruga Bay] 

Oplophorus novaezeelandiae (De Man, 1931) 

= Hoplophorus (Oplophorus) novæ-zeelandiæ De Man, 1931: 369; Figs 1-20. [off Kaikoura, on the east 

coast of the South Island of New Zealand, 2 fms] 

Oplophorus spinosus (Brullé, 1839) 

= Palæmon spinosus Brullé, 1839: 18; vignette on page 3 (as Palémon épineux). [îles Canaries] 

= Hoplophorus Grimaldii Coutière, 1905a: 1; Fig. 1. [full description in Coutière, 1905a; 32°18'N 

23°58'W, 0-2000 m] 

Oplophorus typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= O.[plophorus] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 424. [Nouvelle- 

Guinée] 

= Oplophorus brevirostris Spence Bate, 1888: 766; Plate 127, fig. 3. [Challenger stn 207, 12°21'N 122°15'E, 

off Tablas Island, 700 fms] 

Systellaspis Spence Bate, 1888 

= Systellaspis Spence Bate, 1888 (type species Systellaspis lanceocaudata Spence Bate, 1888, by original 


= Hoplopasiphaea Yokoya & Shibata, 1965 (type species Hoplopasiphaea philippinensis Yokoya & Shibata, 


Systellaspis braueri braueri (Balss, 1914a) 

= Acanthephyra Braueri Balss, 1914a: 594. [0°26'N 6°32'W, 0-4000 m; 0°56'N 4°34'W, 0-4000 m] 

? = Systellaspis echinurus Coutière, 1911a: 158. [au large des côtes d’Espagne et du Portugal entre les 

parallèles 36 et 45, de part et d'autre du 11 e degré W] 

= Systellaspis densispina Stephensen, 1923: 57; Fig. 17. [46°30'N 7°00'W, > 4000 m] 

Systellaspis braueri paucispinosa Crosnier, 1987c: 953; Fig. 3a-b. [31°59'08"N, 158°04'04"E, entre 1685 m de 

profondeur et la surface] 

Systellaspis cristata (Faxon, 1893) 

= Acanthephyra cristata Faxon, 1893: 206. [Albatross stns 3361 (North Pacific Ocean, Panama, 

06°10'00''N 083°06'00''W), 1471 fms; 3381 (North Pacific Ocean, Colombia, 4°56'00"N, 80°52'30"W), 

1772 fms] 

Systellaspis curvispina Crosnier, 1987b: 711; Figs 6a-b, 8a-b. [Madagascar, 13°22'S, 47°17'38"E, 2000-0 m] 

Systellaspis debilis (A. Milne-Edwards, 1881b) 

= Acanthephyra debilis A. Milne-Edwards, 1881b: 13. [trouvée à une profondeur de 500 brasses dans 

le canal de Bahama] 

= Miersia gracilis Smith, 1882: 70, Plate 1, figs 4-4d. [Blake stn 328, 34°28'25"N 75°22'50"W, 1632 fms] 

= Acanthephyra debilis var. Europæa A. Milne-Edwards, 1883: Plate 33, fig. 2. [Type locality not indicated] 

= Systellaspis Bouvieri Coutière, 1905a: 8; Fig. 3. [Princesse-Alice stn 1856, 36°46'N 26°41'W, 0-3250 m] 

= Systellaspis debilis var. indica De Man, 1916: 151. [1°10.5'S 130°09'E, Halmaheira Sea, 798 m] 

= Hoplopasiphaea philippinensis Yokoya & Shibata, 1965: 4; Figs 4, 5. [13°17'N 125°50'E, 100 m] 

Systellaspis eltanini Wasmer, 1986: 52; Figs 13-14. [South Atlantic, 34°12'S 16°35'E, 1550-0 m] 

Systellaspis guillei Crosnier, 1987b: 718; Fig. 11. [Madagascar, 13°04'S 47°56'E, 1500-0 m] 

Systellaspis intermedia Crosnier, 1987c: 948; Figs 1b, 2, 3d. [Ouest-Pacifique, au nord des îles Marquises, 

0°-140°E, chalutage pélagique] 

Systellaspis lanceocaudata Spence Bate, 1888: 758; Plate 124, fig. 7. [Challenger stn 232, 35°11'N 139°28'E, off 

Japan, 345 fms] 

Systellaspis pellucida (Filhol, 1884) 

= Acanthephyra pellucida Filhol, 1884: 199. [26°20'N 14°53'W, 782 m, lectotype designation by Crosnier 

& Forest, 1973] 

= Acanthephyra affinis Faxon, 1896: 162; Plate 2, figs 1-3. [Blake stn 258 (Off Granada, 12°03'15"N, 

61°46'25"W), 159 fms]


Superfamily ATyoIDeA De Haan, 1849 [in De Haan, 1833-1850] 

Family ATyIDAe De Haan, 1849 [in De Haan, 1833-1850] 

Antecaridina edmondson, 1954 

= Mesocaris Edmondson, 1935a (type species Mesocaris lauensis Edmondson, 1935a, by monotypy, 

gender feminine; invalid junior homonym of Mesocaris Ortmann, 1893 (Crustacea Larvata)) 

= Antecaridina Edmondson, 1954 (nomen novum for Mesocaris Edmondson, 1935a, gender feminine) 

Antecaridina lauensis (Edmondson, 1935a) 

= Mesocaris lauensis Edmondson, 1935a: 13; Fig. 4. [cave containing brackish water, Island of Namuka, 

Lau islands, Fiji] 

Archaeatya Villalobos F., 1960a 

= Archaeatya Villalobos F., 1960a (type species Archaeatya chacei Villalobos F., 1960a, by original designation 


Archaeatya chacei Villalobos F., 1960a: 332; Figs 1-25. [Isla de Cocos, 87°30'W 5°30'N] 

Atya leach, 1816a 

= Atys Leach, 1816b (type species Atys scaber Leach, 1816b, by monotypy, gender masculine; name 

placed on the Official List of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 

1957) 

= Atya Leach, 1816a (type species Atys scaber Leach, 1816b, gender feminine; name placed on the 

Official List of Generic Names in Zoology in Opinion 470 in 1957) 

= Evatya Smith, 1871 (type species Evatya crassa Smith, 1871, by monotypy, gender feminine) 

Atya abelei Felgenhauer & Martin, 1983: 333; Figs 1-4. [Panama, El Valle, Cocle Province, unnamed tributary 

of Rio Anton, 600 m elevation] 

Atya africana Bouvier, 1904a: 138. [Samkitta, dans la rivière Ogooué] 

Atya brachyrhinus H.H.Jr. Hobbs & C.W.J. Hart, 1982: 35; Figs 1c, 12g, 15. [Cole’s Cave, Barbados, West 

Indies, 13°10'40"N 59°34'3"W] 

Atya crassa (Smith, 1871) 

= Evatya crassa Smith, 1871: 95. [Fresh water streams, Polvon, and the ”Rio Fulva, two and a half 

miles northwest of Realejo”; both localities in the Occidental Department of Nicaragua] 

Atya dressleri Abele, 1975a: 51; Figs 1-2. [Panamá, Pacific drainage, Veraguas Province, Río Santa María 

drainage, headwaters of Río San Juan, about 15 km above Calobre, 566 m elevation] 

Atya gabonensis Giebel, 1875: 52. [Gabon] 

= Euatya sculptilis Koelbel, 1884: 317; Plate 2, fig. 8; Plate 3, figs 1-8. [Orinoco] 

= Atya sculptata Ortmann, 1890: 465. [Afrika, vielleicht aus West Afrika] 

Atya innocous (Herbst, 1792) 

= Cancer (Astacus) Innocous Herbst, 1792 [in Herbst, 1791-1796]: 62; Plate 28, fig. 3. [das vaderland is 

unbekannt; considered to be ”Oceano Americano ad Martinicam”, see H.H.Jr. Hobbs & C.W.J. Hart, 

1982] 

= Astacus 988 Gronovius, 1764: 231; Plate 17, fig. 6. [in Oceano Americano ad Martinicam; included 

in a work rejected for nomenclatorial purposes in Opinion 261 in 1955] 

= Astacus Nasoscopus Meuschen, 1778: 86. [nomen novum for Astacus 988 Gronovius, 1764; included 

in a work rejected for nomenclatorial purposes in Opinion 260 in 1954] 

= A.[tya] occidentalis Newport, 1847: 159. [West India Islands] 

= Atya robusta A. Milne-Edwards, 1864: 148; Plate 3, fig. 1. [Nouvelle-Calédonie] 

= Atya tenella Smith, 1871: 94, 95. [fresh water streams, Polvon, Occidental Department, Nicaragua] 

Atya intermedia Bouvier, 1904a: 137. [Afrique occidentale, île Saint-Thomas] 

Atya lanipes Holthuis, 1963a: 61; Figs 1-2. [St. Thomas, Virgin Islands] 

Atya limnetes Holthuis, 1986a: 438; Figs 1-3. [Rio Anchicaya, Departemento del Valle del Cauca] 

Atya margaritacea A. Milne-Edwards, 1864 

= Atya Margaritacea A. Milne-Edwards, 1864: 148; Plate 3, fig. 2-2c. [Nouvelle-Calédonie (evidently 

erroneous, most certainly originates from East American waters, see Holthuis, 1966)]


= Atya rivalis Smith, 1871: 94. [fresh water streams Polvon, Occidental Department, Nicaragua] 

Atya ortmannioides Villalobos F., 1956: 459; Figs 1-6. [Río de las Truchas, La Mira, 52.5 km S.SE de Arteaga, 

Michoacán, 2.5 km de la costa, en la vertiente del Pacifico] 

Atya scabra (Leach, 1816b) 

= Atys scaber Leach, 1816b: 345. [Misantla, Estado de Veracruz, Mexico, 19°56'N 96°50'W; neotype 

designation by H.H.Jr. Hobbs & C.W.J. Hart, 1982] 

= Atya mexicana Wiegmann, 1836: 145. [Misantla] 

? = A.[tya] sulcatipes Newport, 1847: 159; Plate 8, fig. 1. [in fresh water, San Nicolao, Cape Verd Islands] 

= Atya punctata Kingsley, 1878a: 91, 92. [Hayti] 

= Atya margaritacea var. claviger Aurivillius, 1898a: 14; Plate 3, Figs 5-8. [Kamerun, Etome, in Bächen] 

= Atya margaritaria clavipes Holthuis, 1966: 234. [lapsus for Atya margaritacea var. claviger Aurivillius, 

1898a] 

Atyaephyra de Brito Capello, 1867 

= Symethus Rafinesque, 1814 (type species Symethus fluviatilis Rafinesque, 1814, an invalid senior 

subjective synonym of Hippolyte Desmarestii Millet, 1831, gender masculine; name suppressed for 

the purposes of the Principle of Priority but not for those of the Principle of Homonymy and placed 


= Acilius Rafinesque, 1815 (nomen novum for Symethus Rafinesque, 1814, gender masculine; name 

suppressed for the purposes of the Principle of Priority for those of the Principle of Homonymy and 

placed on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 522 in 

1958 and Opinion 619 in 1961) 

= Symaethus Agassiz, 1846 [in Agassiz, 1842-1846] (invalid emendation of Symethus Rafinesque, 

1814, gender masculine) 

= Atyaephyra de Brito Capello, 1867 (type species Atyaephyra Rosiana de Brito Capello, 1867, by 

monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

522 in 1958) 

Atyaephyra desmarestii (Millet, 1831) 

= Hippolyte Desmarestii Millet, 1831: 56; Plate 1, figs 1-1b. [les eaux de la Mayenne, de la Sarthe, du 

Loir, du Thouet et du Layon] 

= Symethus fluviatilis Rafinesque, 1814: 23. [dans les ruisseaux et les mares; name suppressed under 

the plenary powers for the purposes of the Principle of Priority but not for those of the Principle of 

Homonymy in Opinion 522 in 1958] 

= Atyaëphyra Desmaresti var. occidentalis Bouvier, 1913a: 72. [le Nord de l’Afrique depuis la Tunisie, 

tout le Sud de l’Europe depuis la Macédonie] 

= Atyaephyra rosiana de Brito Capello, 1867: 6; Plate 1, figs 1-1e. [São Barnabe River, Algarve, Portugal; 

neotype selection by Anastiadou, Kitsos & Koukouras, 2008] 

Atyaephyra orientalis Bouvier, 1913a 

= Atyaëphyra Desmaresti var. orientalis Bouvier, 1913a: 72. [Syrie] 

= Atyaephyra desmarestii mesopotamica Al-Adhub, 1987: 1; Fig. 1. [Shatt Al-Arab river] 

Atyaephyra stankoi Karaman, 1972 

= Atyaephyra desmarestii stankoi Karaman, 1972: 82; Figs 3, 6, 9-10. [Mazedonien, Vardar Flussystem] 

Atydina Cai, 2010a 

= Atydina Cai, 2010a (type species Caridina atyoides Nobili, 1900a, by original designation, gender 

feminine) 

Atydina atyoides (Nobili, 1900a) 

= Caridina atyoides Nobili, 1900a: 478. [Si Oban; Sioban, Pulau Sipura, Kepulauan Mentawei, off west 

coast of Sumatra, Indonesia, according to Cai, 2010a] 

Atyella Calman, 1906a 

= Atyella Calman, 1906a (type species Atyella brevirostris Calman, 1906a, by original designation, gen-


der feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Atyella brevirostris Calman, 1906a: 201; Plate 14, figs 57-64. [Mbete; near mouth of Lofu; Kala; rock-pool 

at Kasakalawe (all Lake Tanganyika)] 

Atyella longirostris Calman, 1906a: 202; Plate 14, figs 65-72. [Mbete; Kala (both Lake Tanganyika)] 

Atyoida Randall, 1840 

= Atyoida Randall, 1840 (type species A.[tyoida] bisulcata Randall, 1840, by monotypy, gender feminine) 

= Ortmannia Rathbun, 1901 (type species Ortmannia henshawi Rathbun, 1901, a junior subjective 

synonym of A.[tyoida] bisulcata Randall, 1840, gender feminine) 

= Pseudatya Roux, 1928a (type species Pseudatya beauforti Roux, 1928a, a junior subjective synonym 

of A.[tya] pilipes Newport, 1847, by monotypy, gender feminine) 

= Vanderbiltia Boone, 1935 (type species Vanderbiltia rosamondae Boone, 1935, a junior subjective synonym 

of A.[tya] pilipes Newport, 1847, by original designation and monotypy, gender feminine) 

Atyoida bisulcata Randall, 1840 

= A.[tyoida] bisulcata Randall, 1840: 140; Plate 5, fig. 5. [Sandwich Islands] 

= Ortmannia henshawi Rathbun, 1901: 120. [Kaiwiki, Hilo, Hawaii, 1800ft altitude, 3 miles from the sea] 

Atyoida pilipes (Newport, 1847) 

= A.[tya] pilipes Newport, 1847: 160. [Apia, Upoln, New Zealand] 

= Atyoida tahitensis Stimpson, 1860a: 28. [in aquis dulcibus insulae ’Tahiti’] 

= Caridina acuminata Stimpson, 1860a: 29. [stream of Naka-kaigan coast, 60-70 m above sealevel, 

Chichi-jiam Island, Ogasawar Islands; neotype designation by Cai et al., 2006] 

= Caridina brevirostris Stimpson, 1860a: 29. [River Aritsu, Kume-jima Island; neotype designation by 

Cai et al., 2006] 

= Atya brevirostris De Man, 1892a: 360; Plate 21, figs 21-21d. [Fluss Wukur bei Sikka; Fluss Lella bei 

Sikka; Fluss bei Mbawa unterhalbs des Wasserfalls (all Flores); Fluss Koinino bei Kupang (Timor)] 

= Pseudatya beauforti Roux, 1928a: 209; Figs 1-9. [Batjan] 

= Vanderbiltia rosamondae Boone, 1935: 160; Plates 41-42. [Venus Point Reef, Tahiti, Society Islands, in 

coral] 

Fig. 15. Atyopsis moluccensis (De Haan, 1849). Photo by Tin-Yam Chan.


= Vanderbiltia mirabilis Holthuis, 1953a: 114. [manuscript name published as a synonym] 

Atyoida serrata (Spence Bate, 1888) 

= Atya serrata Spence Bate, 1888: 699; Plate 119, fig. 2. [Valley of San Antonia, San Iago, Cape Verde 

Islands from a fresh-water stream; probably erroneous, see Chace, 1983a] 

= Atya serrata mutation Alluaudi Bouvier, 1904b: 448. [un torrent de la montagne d’Ambre, à Madagascar; 

l’île Bourbon; dans les ravines des montagnes de Salasie et d’Helbour; l’île Maurice] 

Atyopsis Chace, 1983a 

= Atyopsis Chace, 1983a (type species A.[tya] spinipes Newport, 1847, by original designation, gender 

feminine) 

Atyopsis moluccensis (De Haan, 1849 [in De Haan, 1833-1850]) (Fig. 15) 

= Atya moluccensis De Haan, 1849 [in De Haan, 1833-1850]: 186; Plate O. [Moluccas, Indonesia; lectotype 

designation by Holthuis, 1993b] 

= Atya armata A. Milne-Edwards, 1864: 149; Plate 3, figs 3-3a. [Batavia (p. 145, 152), not les îles Philippines 

(p. 149), see Chace, 1983a] 

= Atya gustavi Ortmann, 1890: 467; Plate 36, fig. 9a-c. [Sumatra, Indrapura-Fluss] 

= Atya lineolata De Man, 1892a: 357 (footnote). [manuscript name attributed to Kulh, cited as synonym 

of Atya moluccensis] 

Atyopsis spinipes (Newport, 1847) 

= A.[tya] spinipes Newport, 1847: 159. [Philippine Islands] 

= Atya dentirostris Thallwitz, 1891: 101. [Nord-Celebes; fully described in Thallwitz, 1892] 

? = Atya brevirostris var. De Mani Nobili, 1900a: 475, fig. 1. [Fiume Sereinu, Isole Mentawei] 

Australatya Chace, 1983a 

= Australatya Chace, 1983a (type species Atya striolata McCulloch & McNeill, 1923, by original designation 


Australatya striolata (McCulloch & McNeill, 1923) 

= Atya striolata McCulloch & McNeill, 1923: 55; Plate 9, figs 3-4. [Norton’s Basin, Nepean River, New 

South Wales] 

Caridella Calman, 1906a 

= Caridella Calman, 1906a (type species Caridella cunningtoni Calman, 1906a, by original designation, 

gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

Caridella cunningtoni Calman, 1906a: 199; Plate 13, figs 45-52. [Kala, on rocks, shallow water; Kirando, 8 

fms; Utinta, 15 fms (all Lake Tanganyika)] 

Caridella minuta Calman, 1906a: 200; Plate 13, figs 53-56. [near mouth of Lofu, on rocks, shallow water; 

Karema, tow-netting, surface (both Lake Tanganyika)] 

Caridella paski Calman, 1928: 739; Figs 1-3. [Kigoma, east side of Lake Tanganyika] 

Caridina H. Milne edwards, 1837 [in H. Milne edwards, 1834-1840] 

= Caridina H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species Caridina typus H. 

Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy and by indication under 

Article 68c of the ICZN, gender feminine; name placed on the Official List of Generic Names in 


Caridina ablepsia Guo, Jiang & Zhang, 1992: 4; Figs 1-9. [cave river, Wangcun Village, 109°56'E 28°46'N, 

Yongshun County, Hunan Province] 

Caridina acuta Liang, Chen & W.-X. Li, 2005: 530; Figs 15-29. [caves of Libo County (25°41'N 107°83'E), 

Maolan Nature Sanctuary of Guizhou] 

Caridina acutirostris Schenkel, 1902: 496; Plate 8, figs 3a-c. [Süden des Posso-See] 

Caridina africana Kingsley, 1883: 127; Plate 1, figs 3-3a. [Zulu Mission, South Africa] 

= Caridina africana forme typica Bouvier, 1925: 213; Fig. 470. [pays des Zoulous]


= Caridina wyckii var. paucipara Weber, 1897: 168. [River Umhlasine, Natal; lectotype designation by 

Richard & Clark, 2010a] 

Caridina alba J. Li & S. Li, 2010: 19; Figs 2-5. [Tenglongdong cave, Lichuan city, Enshi Autonomous Prefecture, 

Hubei province, China] 

Caridina alphonsi Bouvier, 1919: 330. [localité inconnue; austro-malaise according to Bouvier, 1925] 

Caridina amnicolizambezi Richard & Clark, 2009: 45; Figs 22-23. [Angola, Rio Lucoge, branch of the Chicapu 

River, 73°36'S 20°31'E] 

Caridina amoyensis Liang & Yan, 1977: 219; Figs 1-4. [Xiamen, suburabm rivulet, Fujian Province] 

Caridina angulata Bouvier, 1905c: 84; Fig. 8. [rivière Ranofotsy, près de Fianarantsoa] 

Caridina angustipes Guo & Liang, 2003: 45; Figs 1A-C. [Wentang Village, Xinhua County, Hunan Province, 

400 m] 

Caridina anislaq Cai, Choy & Ng, 2009: 82; Fig. 11. [Spring Anislaq, outside Cave Valencia, Valencia] 

Caridina annandalei Kemp, 1918a: 96; Plate 25, figs 6-15. [Inlé Lake] 

Caridina apodosis Cai & N.K. Ng, 1999: 1627; Figs 14-17. [mountain stream near Taitong village, New 

Territory, Hong Kong] 

Caridina appendiculata Jalihal & Shenoy, 1998: 128. [River Bari, Flores; lectotype designation by Cai & Ng, 

2007] 

Caridina aruensis Roux, 1911: 82. [ruisseau Matora, Soungi Manoumbai, Arou; ruisseau Panoua Bori, 

Soungi Manoumbai, Arou] 

Caridina babaulti babaulti Bouvier, 1918 

= Caridina Babaulti Bouvier, 1918: 388; Figs 4-6. [Mukhi; Majghaon; Rajadhar] 

Caridina babaulti basrensis Al-Adhub & Hamzah, 1987: 225; Fig. 1. [small channels connected to Shatt al- 

Arab River, Iraq] 

Caridina babaultioides babaultioides Yü, 1938 

= Caridina babaultioides Yü, 1938: 301; Figs 13-14. [small mountain stream, at an altitude of about 910 

m, at Shui-tuan-pa near Yen-tsing city, Yunnan Province] 

Caridina babaultioides angustifolia Cai & Yuan, 1996 

= Caridina angustifolia Cai & Yuan, 1996: 385; Figs 14-15. [Wafang stream, near Jinsha village] 

Caridina babaultioides emeica Liang, 2004: 218; Fig. 105. [Emeishan, Sichuan] 

Caridina babaultioides phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province] 

= Caridina phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province] 

Caridina bakoensis Ng, 1995a: 192; Figs 8-10. [Middle stretch of Sungai Serait] 

Caridina bamaensis Liang & Yan, 1983a: 252; Figs 1-10. [Bama County, Guangxi, Southern China] 

Caridina baojingensis Guo, He & Bai, 1992: 609; Figs 1-9. [China, Hunan Province, Baojing County, Qingshui 

village] 

Caridina batuan Cai, Choy & Ng, 2009: 80; Fig. 10. [Cave Castigio, Batuan] 

Caridina belazoniensis Richard & Clark, 2009: 33; Figs 14-15. [Kenya, Mbelazoni, Lower Ahti River, Sabaki 

Estuary] 

Caridina boholensis Cai, Choy & Ng, 2009: 75; Figs 7-8. [Quila Cave, Nueva Vida Norte, Batuan] 

Caridina brachydactyla De Man, 1908a 

= Caridina nilotica var. brachydactyla De Man, 1908a: 269; Plate 20, figs 8a-c. [Indonesia, by Reo, Flores; 

lectotype designation by Richard & Clark, 2010a] 

= Caridina nilotica var. natalensis De Man, 1908a: 262; Plate 20, figs 3-3b [South Africa, Umgeni River, 

Durban, Natal; lectotype designation by Richard & Clark, 2010a] 

= Caridina nilotica var. brevidactyla Roux, 1920: 320. [Aru-Inseln: Bach Matora, am Sungi Manumbai; 

Bach Panua Bori am Sungi Manumbai; Bach bei Papakula, Kobroor; Wokamar, Wokam; Samang; 

Seltutti, Kobroor; Dobo, Wammer; Pobdietur, Terangan; Udjir] 

Caridina breviata N.K. Ng & Cai, 2000: 171; Figs 4-6. [Zhapu village, Yangjiang County, Guangdong Province, 

southern China] 

Caridina brevicarpalis De Man, 1892a: 397; Plate 24, figs 30-30d. [Celebes, aus einem Flusse bei Palopo, 

Luwu] 

Caridina brevispina Liang & Yan, 1986a: 203; Fig. 5. [Tongren County, Guizhou Province]


Caridina bruneiana Choy, 1992: 49; Figs 1-4. [Negara Brunei Darussalam, on the upper reaches of Temburong 

River at Batang Duri, 04°36'05''N 115°06'45''E, altitude approx. 33 m] 

Caridina buehleri Roux, 1934a 

= Caridina bühleri Roux, 1934a: 219; Figs 1-5. [Bimoun, côte occidentale Nouvelle-Irlande] 

Caridina buergersi Karge, von Rintelen & Klotz, 2010: 146; Figs 5-6. [Papua New Guinea, former 

”Deutsch-Neuguinea”, near Mäanderberg] 

Caridina buhi Cai & Shokita, 2006a: 253; Figs 3-4. [Binahugan River, Buhi, Camarines Sur, Luzon, Philippines] 

Caridina bunyonyiensis Richard & Clark, 2005: 715; Figs 5-6. [Uganda, Lake Bunyonyi] 

Caridina burmensis Cai & Ng, 2000: 941; Fig. 7. [Myitkyina, Myanmar] 

Caridina caerulea von Rintelen & Cai, 2009: 428; Figs 51-53. [Lake Poso, west shore, Cape Bancea, 

01°59.023'S 120°35.108'E, on rocks in shallow water] 

Caridina calmani Bouvier, 1919 

= Caridina Calmani Bouvier, 1919: 334. [Ambatoubavara, Madagascar] 

= Caridina Bouvieri Roux, 1929: 312; Figs 7-12. [Manjakatompo, altit. 1940 m, ruisseau descendant de 

l’Ankaratra] 

Caridina camaro Cai, Choy & Ng, 2009: 82; Fig. 12. [Cave Camaro, Batuan] 

Caridina cantonensis Yü, 1938: 290; Figs 7-8. [Qing’ao vilage, Nan’ao County, Guangdong Province, China; 

neotype designation by Cai & N.K. Ng, 1999] 

= Caridina mutata Cai & N. K. Ng, 1999: 1624; Figs 12-13. [mountain stream near Fangcheng Town, 

Fangcheng County, Guangxi Province, China] 

Caridina caobangensis S.-Q. Li & Liang, 2002: 711; Figs 3-4. [PacBo Village, District Ha Quang, Cao Bang 

province, Vietnam] 

Caridina carli Roux, 1931a: 38; Figs 4-11. [Anaimalais, environs de Valparai, dans la rivière Naduar et ses 

affluents] 

Caridina cavaleriei cavaleriei Bouvier, 1919 

= Caridina Cavalerii Bouvier, 1919: 332. [Gan-chouen-fou (Kouy-Tchéou)] 

Caridina cavaleriei industana Roux, 1931a: 35; Figs 1-3. [Aliyar Riv., près de Malayandi Pattanam, 6 milles 

au Sud de Pollachi] 

Caridina cavalerieioides Liu & Liang in Liang, 2004: 207; Fig. 100. [Malin Village, near Guiyang City, 

Guizhou Province, China] 

Caridina caverna Liang, Chen & W.-X. Li, 2005: 529; Figs 1-14. [caves of Libo County (25°41'N 107°83'E), 

Maolan Nature Sanctuary of Guizhou] 

Caridina cavernicola Liang & Zhou, 1993: 232; Figs 2(1)-2(2). [Lenggu Cave, Duan County, Guangxi Province] 

Caridina cebuensis Cai & Shokita, 2006a: 250; Figs 1-2. [spring water to Sagay River, Cebu Island, Philippines] 

Caridina celebensis De Man, 1892a 

= Caridina serratirostris var. celebensis De Man, 1892a: 385; Plate 23; figs 28f-h. [Celebes, Fluss bei 

Palopo, Luwu] 

= Caridina serratirostris koterai Kamita, 1951: 75; Figs A-J. [Shimoko, Iwami province, SE Hunshu, 

Japan] 

Caridina celestinoi Blanco, 1939a: 392; Plate 3, Figs 8-10. [mountain stream, Helosig, Leyte] 

Caridina chauhani Chopra & Tiwari, 1949 

= Caridina nilotica var. chauhani Chopra & Tiwari, 1949: 219; Figs 2-3 [tanks at Sale bhata, Chandanbhati, 

Bolangir, Salepali and Titilagarh; Ang River at Salebhata] 

= Caridina williamsoni Jalihal, Shenoy & Sankolli, 1984: 1; Figs 1-3. [Sadhankeri tank, Dharwar] 

Caridina chishuiensis Cai & Yuan, 1996: 379; Figs 8-10. [stream near Binan village] 

= Caridina euryphylla Cai & Yuan, 1996: 382; Figs 11-13. [Qintan village, near Dongping village] 

Caridina clavipes Guo & Liang, 2003: 47; Figs 1D-F. [Wentang Village, Xinhua County, Hunan Province] 

Caridina clinata Cai, X.Q. Nguyên & Ng, 1999: 531; Figs 1-2. [ditch at Cuc Phuong National Park, Ninh 

Binh Province, Northern Vietnam]


Caridina cognata De Man, 1915a: 397; Plate 28, figs 3-3g, 4-4b [in einem kleinen Bache von süssem Wasser 

in der Küsten-gegend zwischen der Humboldt-Bai und dem unteren Laufe des Tami-Flusses; in 

Bächlein zwischen den kleinen Flüssen Tjahé and Jasa; aus dem Tjahé-Flusse; aus dem kleinen 

Tjano-Flusse bei Njao; Zoutbron] 

Caridina confusa Choy & Marshall, 1997: 27; Figs 1a, 2i-j, 3b-c, 4a-e. [Thiaki Creek at Seamark Road crossing, 

17°23.5'S 145°32.5'E] 

Caridina congoensis Richard & Clark, 2009: 48; Figs 24-25. [Congo, Bakou, in river, in total darkness, 30 

m] 

Caridina cornuta Liang & Yan, 1986a: 200; Fig. 3. [Zhijin County, Guizhou Province] 

Caridina crassipes Liang, 1993: 23; Fig. 2. [Xianshui Cave, Pangshi village, Hunan] 

Caridina crurispinata Gurney, 1984: 574; Figs 4-7. [Dark zone, la Grotte d’Antsatrobonko, Ankarana Massif, 

40 km north of Ambilobé, 60 km south of Diego Suarez, northern province of Diego Suarez] 

Caridina cucphuongensis Đăng, 1980 

= Caridina serrata cucphuongensis Đăng, 1980: 404; Fig. 230. [streams in Cucphuong area, Ninh Binh 

Province, Northern Vietnam] 

Caridina curta Liang & Cai, 2000: 179; Fig. 2. [Longquan spring in Huanglong (Yellow Dragon) Temple, 

Jianshui County, Yunnan] 

Caridina demani Roux, 1911: 94. [Tawarin] 

Caridina demenica Cai & Li, 1997: 315; Fig. 1. [Demen Cave, near Yongkang village, Libo County of 

Guizhou Province, southern China] 

Caridina dennerli von Rintelen & Cai, 2009: 355; Figs 4-6. [Lake Matano, east shore, just at entrance to 

outlet bay, 02°31.54'S 121°27.00'E] 

Caridina dentifrons N.K. Ng & Cai, 2000: 167; Figs 1-3. [Baijin village, Huishui County, Guizhou Province, 

southern China] 

Caridina devaneyi Choy, 1991: 348; Figs 2-4. [stream at 8-mile Point near Suva, Viti Levu, Fiji Islands] 

Caridina dianchiensis Liang & Yan, 1985a: 196; Fig. 1. [Songhuaba Reservoir- Panlong River, Kunming, 

Jiangchuan County, Yunnan province] 

Caridina disjuncta Cai & Liang, 1999: 77; Fig. 4. [Shilin (Stone forest) village, Lunan County, central Yunnan] 

Caridina disparidentata Liang, Yan & Wang, 1984: 8; Figs 1-12. [mountain stream, Qujing County, Yunnan 

Province] 

= Caridina heterodentata Liang & Yan, 1985a: 196. [unjustified emendation of Caridina disparidentata 

Liang, Yan & Wang, 1984; see Cai & Ng, 2001a] 

Caridina ebuneus Richard & Clark, 2009: 39; Figs 18-19. [Ivory Coast, vicinity Abengourou] 

Caridina edulis Bouvier, 1904a: 135. [Anantsahalankely] 

Caridina elisabethae Karge, von Rintelen & Klotz, 2010: 142; Figs 3-4. [Papua New Guinea, Morobe, Herzog 

Mts., Bundun, 700-800 m, 06°51.598'S 146°37.07'E] 

Caridina elliptica Cai & Yuan, 1996: 376; Figs 5-7. [stream near Panlong town] 

Caridina elongapoda Liang & Yan, 1977 

= Caridina nilotica elongapoda Liang & Yan, 1977: 220; Figs 5-8. [Xingzai, Zhangpu County, Fujian 

Province] 

Caridina endehensis De Man, 1892a 

= Caridina brevicarpalis var. endehensis De Man, 1892a: 399; Plate 24, fig. 30e. [aus dem Flusse Ba bei 

Endeh; Fluss bei Mbawa (both Flores)] 

Caridina ensifera Schenkel, 1902: 490; Plate 8, figs 1-11e. [Lake Poso, Celebes (Sulawesi), Indonesia; lectotype 

designation by Cai & Wowor, 2007] 

Caridina evae Richard & Clark, 2009: 28; Figs 11-13. [Nigeria, Chubra division] 

Caridina excavata Kemp, 1913b: 306; Plate 20, figs 32-35; Plate 21, figs 36-37. [backwater of the Rowta 

River, Brahmaputra drainage system] 

Caridina excavatoides Johnson, 1961: 127; Figs 3-11. [stream running between rubber plantations and ricefields, 

about nine miles from Alor Star, Kedah, on the Pokok Sena Road] 

Caridina fasciata Hung, Chan & Yu, 1993: 489; Figs 1E, 6. [Ping Tong County, Heng Chun]


Caridina fecunda Roux, 1911: 95. [Lac Jamour] 

Caridina feixiana Cai & Liang, 1999: 74; Figs 1-2. [stream in Feixia cave, Geju County, Yunnan, China] 

Caridina fernandoi Arudpragasam & Costa, 1962: 8; Fig. 1. [shallow streamlets at Warakapola, and Seelangama, 

and also from shallow sheltered areas of the Maha Oya at Mawanella and the Deduru Oya 

and the Magura Oya at Kurunegala, Sri Lanka] 

Caridina fijiana Choy, 1983: 147; Fig. 1. [Fiji, Viti Levu, Nadarivatu, Nukunuku Creek, 17°35'40''S 

177°57'25''E, 640 m] 

Caridina flavilineata Đăng, 1975: 70; Fig. 5. [Nam Ha Province] 

Caridina formosae Hung, Chan & Yu, 1993: 487; Figs 1D, 5. [Keelung, Pa Chih Men] 

Caridina fossarum Heller, 1862b 

= C.[aridina] fossarum Heller, 1862b: 411. [Schiraz] 

? = Caridina syriaca Bouvier, 1904a: 132. [Syrie] 

Caridina gabonensis Roux, 1927a: 239; Figs 1-7. [Lambaréné, Gabon] 

Caridina ghanensis Richard & Clark, 2009: 35; Figs 16-17. [Republic of Ghana, Pond Vume] 

Caridina glaubrechti von Rintelen & Cai, 2009: 359; Figs 7-9. [Lake Towuti, west shore, south of Cape 

Timbalo, 02°42.91'S 121°26.78'E] 

Caridina glossopoda Liang, Guo & Gao, 1993: 41; Fig. 1. [Ala, Fenghuang County, Hunan, 27°95'N 

109°65'E] 

Caridina gordonae Richard & Clark, 2005: 717; Figs 7-8. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina gortio Cai & Anker, 2004: 247; Figs 8, 9a-f. [Can Gortio Cave, 11°59'00''N 124°53'00''E, altitude ca 

200 m, Matalud, West Samar, Philippines] 

Caridina gracilipes De Man, 1892a 

= Caridina Wyckii var. gracilipes De Man, 1892a: 387; Plate 24, figs 29e-k [aus dem Flusse von Maros; 

Makassar; aus einem kleinen mit dem Meere nicht in Verbindung stehenden Bache zu Balangnipa; 

Palima, aus dem Tjenrana; Pampanua, aus dem Tjenrana (all Celebes); Saleyer, aus dem Flusse 

Bonéa] 

= Caridina nilotica var. bengalensis De Man, 1908a: 265; Plate 20, figs 6-6b. [Port Canning an Dhappa, 

Calcutta] 

Caridina gracilirostris De Man, 1892a: 399; Plate 25, fig. 31-31d. [river near Maros, Sulawesi (Celebes), 

Indonesia; lectotype designation by Cai & Ng, 2007] 

= Caridina pseudogracilirostris Thomas, V.K. Pillai & N.N. Pillai, 1976: 871; Fig. 1. [Cochin backwater] 

Caridina gracillima Lanchester, 1901: 560; Plate 34, fig. 1. [Malay Peninsula; lectotype designation by Cai 

& Ng, 2007] 

Caridina grandirostris Stimpson, 1860a: 28 [upper stream of Tima River, about 4 km from river mouth, 

Okinawa Island, Ruykyu Islands; neotype designation by Cai et al., 2006] 

Caridina guangxiensis Liang & Zhou, 1993: 234; Figs 3(1)-3(2). [Dawangshan cave, Guilin City, Guangxi 

Province] 

Caridina gueryi Marquet, Keith & Kalfatak, 2009: 160; Figs 1-3. [unnamed river, Santo Island (Vanuatu), 

15.29590 S 167.16 E] 

Caridina guiyangensis Liang, 2002a: 119; Fig. 2. [China, Guizhou Province, Guiyang County] 

Caridina gurneyi Jalihal, Shenoy & Sankolli, 1984: 29; Figs 12-14. [Malaprabha river, near old bridge, 

Khanapur (Belgaum District)] 

Caridina hainanensis Liang & Yan, 1983b: 211; Fig. 1. [Wencang County, Hainan Island, China] 

Caridina hanshanensis Tan, 1990: 278; Figs 1-9. [Hanshan County, 31°45'N 118°7'E, Anhui Province] 

Caridina harmandi Bouvier, 1906f 

= Caridina Harmandi Bouvier, 1906f: 483. [Japon] 

Caridina hodgarti Kemp, 1913b: 309; Plate 20, Figs 29-31; Plate 21, figs 38-39. [Kobo, Abor county] 

Caridina holthuisi von Rintelen & Cai, 2009: 364; Figs 10-12. [Lake Matano, south shore, canal between 

island and mainland, 02°28.46'S 121°15.83'E] 

Caridina hongyanensis Cai & Yuan, 1996: 373; Figs 2-4. [Yantanggou stream near Hongyan village] 

Caridina hova Nobili, 1905c: 499. [Fort Dauphin] 

Caridina huananensis Liang, 2004: 147; Fig. 69. [Guangdong]


Caridina hubeiensis Liang & S.-Q. Li, 1993: 213; Figs 1-9. [Badong, Hubei Province] 

Caridina hunanensis Liang, Guo & Gao, 1993: 44; Fig. 3. [Zhonglian, Lengshuijiang City, Hunan, 27°7'N 

111°4'E] 

Caridina imitatrix Holthuis, 1970: 100; Fig. 3. [Rivière Bleue near the bridge near Ouénarou (Mont des 

Sources); Marsh of Rivière Blanche, Ouénarou region; Side branch of Rivière Blanche, near the forest 

road leading from Ouénarou forestry station to the forestry log cabin on the western slope of Mt. 

Pouédihi, Ouénarou region; same branch, near the log cabin; Thir River near the confluence with a 

side branch, granite area near St. Louis, S.E. of Nouméa; Rivière des Lacs, which empties in the Lac 

en Huit, near highway to Nouméa] 

Caridina indistincta indistincta Calman, 1926 

= Caridina indistincta Calman, 1926: 244; Fig. 3. [St. George District, Queensland] 

Caridina indistincta sobrina Riek, 1953: 119; Fig. 9. [Rocky Creek, Fraser Island, Queensland] 

Caridina isaloensis isaloensis Coutière, 1899b 

= Caridina typus var. Isaloensis Coutière, 1899b: 383. [Fleuve Onilahy, côte Ouest] 

= Caridina madagascariensis Bouvier, 1904a: 134. [rivière Fandramanona] 

= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar] 

Caridina isaloensis grandidieri Bouvier, 1904a 

= Caridina Grandidieri Bouvier, 1904a: 133. [rivière Fandiamanana, Madagascar] 

Caridina jalihali Mariappan & Richard, 2006: 47; Figs 22-24. [River Arani at Periapalayam; lake at Thenneri; 

ponds at Maduranmangalam, Manimangalam, Redipalayam, RoshanNagar and Singaperumal 

Koli] 

= Caridina gurneyi lonavalensis Kadrekar & Sankolli, 1987: 60. [nomen nudum] 

Caridina jeani Cai, 2010b: 80; Figs 1-2; nomen novum for Caridina typus var. brevirostris Roux, 1911. [Elat, 

Kei Islands, Moluccas, Indonesia; lectotype designation by Cai, 2010b] 

Caridina jiangxiensis Liang & Zheng, 1985 

= Caridina pingi jiangxiensis Liang & Zheng, 1985: 319; Fig. 1(1b-7b). [Shixi stream, Fengxin County, 

Jiangxi Province] 

Caridina johnsoni Cai, Ng & Choy, 2007: 301; Figs 15-16. [Singapore, Lower Peirce Reservoir, north arm] 

Caridina kaombeflutilis Richard & Clark, 2010a: 331; Figs 14-15. [Malawi, Kaombe River] 

Caridina kempi Jalihal, Shenoy & Sankolli, 1984: 13; Figs 6-8. [Hosayellapur tank (Hirekeri), Dharwar] 

Caridina kilimae Hilgendorf, 1898: 35. [Marangu am Kilimandscharo] 

Caridina kunmingensis Z.-Z. Wang & Liang, 2001: 303; Fig. 1. [Kunming, Yunnan Province] 

Caridina kunnathurensis Richard & Chandran, 1994: 250; Figs 4, 5A. [Kunnathur pond, Madras, India] 

Caridina laevis Heller, 1862b 

= C.[aridina] laevis Heller, 1862b: 411. [Java] 

Caridina lamiana Holthuis, 1965a: 39; Fig. 14. [Ambodivoangy near Maroantsetra, Antongil Bay, NE 

Madagascar; Forest between 20 and 30 km NW of Ambahoabé, Simianona River, N of Soanierana, 

east coast of Madagascar] 

= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar] 

Caridina lanceifrons Yu, 1936: 89; Figs 4-7. [near the light house at Hai-kiu-sche in the salt water] 

Caridina lanceolata Woltereck, 1937: 224; Fig. 1.7a-c; Table 1. [Lake Mantano, Mengonuwai, Sulawesi; 

neotype designated by Cai, Wowor & Choy, 2009] 

Caridina lanzana Holthuis, 1980c: 2; Figs 1-3. [small well named Bog Der, 08°35'54''N 48°46'30''E, Nogal 

Valley, northern Somalia] 

Caridina laoagensis Blanco, 1939a: 390; Plate 2. [?Laoag River, Laoag, Ilocos Norte Province, Luzon] 

Caridina leclerci Cai & Ng, 2009: 1108; Fig. 9. [Pange, cave at Tallasa, Maros, Sulawesi Selatan] 

Caridina leucosticta Stimpson, 1860a: 28. [138°56.43'E 34°41.83'E, Inouzawa River, Shimoda City, Izu Peninsula; 

neotype designation by Cai et al., 2006] 

Caridina leytensis Blanco, 1939a: 391; Plate 3, Figs 1-7. [Helosig, Leyte] 

Caridina liangi Jiang, Guo & Zhang, 2002: 220; Fig. 1. [small stream at an elevation of 450 m near Yongping 

Village, Yuanling County] 

Caridina liaoi Cai, Choy & Ng, 2009: 72; Figs 5-6. [Bilar River, Bilar] 

Caridina lima Liang, Guo & Gao, 1993: 43; Fig. 2. [Buermen, Yongshun County, Hunan, 29°N 109°8'E]


Caridina linduensis Roux, 1904: 541; Plate 9, figs 1-4. [Lac Lindu, région centrale de Célèbes (part. Occid.), 

altitude 980 m] 

Caridina lineorostris Richard & Clark, 2009: 51; Figs 26-27. [Gabon, Keri, forest stream, 20 km North of 

Lambarene] 

Caridina lingkonae Woltereck, 1937 

= Caridina Lingkonae Woltereck, 1937: 218; Fig. 1.1; Table 1. [Lake Towuti, west shore, at entrance to 

outlet bay, Cape Larona, 02°48.43'S 121°24.75'E; neotype designated by von Rintelen & Cai, 2009] 

Caridina lipalmaria Richard & Clark, 2010b: 639; Figs 1-2. [Bestiboka River Basin, Beharena River, Antaniditra, 

48°02'55''E 18° 25'43''S, 13730 asl] 

Caridina liui Liang & Yan, 1986a: 202; Fig. 4. [Tongzi County, Guizhou Province] 

Caridina lobocensis Cai, Choy & Ng, 2009: 68; Figs 2-4. [tributuary of Loboc River, Loboc] 

Caridina loehae Woltereck, 1937 

= Caridina Loéhae Woltereck, 1937: 222; Fig. 1.5a-d; Table 1. [Lake Towuti, about 3 km south of Timampu, 

estuary of Sungei Batuopa, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina longa Liang & Yan, 1985a: 200; Fig. 3. [spring water, Huanglongsi, Jianshui, Yunnan Province] 

Caridina longiacuta Guo & Wang, 2005: 14; Fig. 1. [near Yuanliping village, Yizhang County, Hunan Province 

(ca. 25°25'N 112°57'E)] 

Caridina longicarpus Roux, 1926a 

= Caridina weberi var. longicarpus Roux, 1926a: 212; Figs 37-39. [Nouvelle-Calédonie, au-dessus 

d’Oubatche, pentes du Mt. Ignambi, env. 600 m d’altitude] 

Caridina longidigita Cai & Wowor, 2007: 317; Figs 4-5. [west coast of Lake Poso at Taipa area, Kab. Poso, 

Sulawesi Tengah] 

Caridina longifrons Cai & Ng, 2007: 1599; Fig. 6. [Kabupaten Maros, stream above Bantimurung waterfall, 

Sulawesi, Indonesia] 

Caridina longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= C.[aridina] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363. [la rivière 

de la Macta, près d’Oran [probably erroneous]] 

= Caridina nilotica var. meridionalis Roux, 1926a: 207. [Pemboa; Haute Tiouaka; Koné; Coula-Boréaré; 

La Foa; Coindé, Ruisseua Katiramona, entre Nouméa et Païta (all Nouvelle-Calédonie)] 

= Caridina acuticaudata Đăng, 1975: 70; Fig. 4. [Boi River, Hoa Binh Province] 

Caridina lovoensis Roth-Woltereck, 1955: 197; Figs 1-2. [Lovo B., 20 km von Thysville (Bas-Congo), 

Kalkhöhlen] 

Caridina lufengensis Cai & Duan, 1998: 330; Figs 1-3. [pond near Sizhi Town, Lufeng County, Yunnan, 

China, 25.07°N, 102.10°E, elevation 1800 m] 

Caridina lumilympha Richard & Clark, 2010a: 328; Figs 12-13. [Kenya, Lumi River] 

Caridina macrodentata Cai & Shokita, 2006b: 2156; Figs 13-14. [123°51.84'E 24°23.65'N, fast flowing water, 

about 200 m from sea, Omija River, Iriomote Island, Ryukyu Islands] 

Caridina macrophora Kemp, 1918b 

= Caridina nilotica var. macrophora Kemp, 1918b: 277; Fig. 9. [Tale Sap, Peninsular Siam] 

= Caridina subnilotica Đăng, 1975: 69; Fig. 3. [ponds in Hanoi] 

Caridina maculata L. Wang, Liang & F. Li, 2008: 49; Figs 1-2. [Yingde County, Guangdong Province, 

N23°54.5' E113°13.6'] 

Caridina mahalona Cai, Wowor & Choy, 2009: 29; Figs 8-9. [Lake Mahalona, at southwest coast] 

Caridina malayensis Cai, Ng & Choy, 2007: 305; Fig. 17. [stream at Nee Soon Swamp, Singapore] 

Caridina malawensis Richard & Clark, 2009: 69; Figs 38-39. [Malawi, rocky shore of Lake Malawi] 

Caridina masapi Woltereck, 1937 

= Caridina Masapi Woltereck, 1937: 223; Fig. 1.6a-h; Table 1. [southern shore of Lake Masapi, South 

Sulawesi, 2°50.84'S 121°21.09'E; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina mathiassi Silas & Jayachandran, 2010: 1; Figs 1-5, 6J, 7. [Mahendragiri Estate, part of southern 

Western Ghats, Kanyakumari District, Tamil Nadu, India, 8° 27'32.2''N 77°23'32.5''E] 

Caridina mauritii Bouvier, 1912a 

= Caridina Mauritii Bouvier, 1912a: 298. [Maurice] 

Caridina mccullochi Roux, 1926b: 249. [Pallal, Horton River, near Bingara]


Caridina medifolia Cai & Yuan, 1996: 388; Figs 16-18. [Xiaojin village near Daqun town] 

Caridina mengae Liang, 1993: 22; Fig. 1. [Xianshui Cave, Panshi village, Songtao County (28.19°N 

109.20°E), Guizhou Province] 

Caridina mengaeoides Guo & Suzuki, 1996: 97; Figs 1-2. [Yuanling County, Hunan Province, China, ca. 

28°18'N 110°20'E, altitude ca. 750 m] 

Caridina menghaiensis Cai & Dai, 1999: 217; Figs 5-6. [reservoir in Menghai County] 

Caridina meridionalis L. Wang, Liang & F. Li, 2008: 56; Figs 7-8. [Heyuan County, Guangdong Province] 

Caridina mertoni Roux, 1911: 84. [Elat, île de Grand-Kei; entre Elat et Ohinangan, île de Grand-Kei; Enraalan, 

île de Grand-Kei; Warka, île de Grand-Kei] 

Caridina mesofluminis Richard & Clark, 2009: 66; Figs 36-37. [Cameroon, Mess stream] 

Caridina mindanao Cai & Shokita, 2006a: 259; Figs 7-9. [Lake Mainit, Mindanao, Philippines] 

Caridina minidentata Cai & Anker, 2004: 250; Figs 10-11. [Can Gortio Cave, 11°59'00''N 124°53'00''E, altitude 

ca 200 m, Matalud, West Samar] 

Caridina minnanica Liang, 2002a: 121; Fig. 3. [China, Fujian Province, Yunxiao County] 

Caridina modiglianii Nobili, 1900a 

= Caridina Modiglianii Nobili, 1900a: 477. [Kifa-juc] 

Caridina moeri Roth-Woltereck, 1984: 102; Figs 1 (1a-b, 2, 3 (1a-e). [Mwerusee] 

Caridina mongziensis Liang, Yan & Z.-Z. Wang, 1987: 133; Figs 1-9. [Mongzi County, 23°20'N 103°25'E, 

Yunnan Province] 

Caridina multidentata Stimpson, 1860a: 29. [stream in Tenno-ura, Chichi-jima Island, Ogasawara Islands; 

neotype designation by Cai et al., 2006] 

= Caridina japonica De Man, 1892b: 261; Plate 9, Figs 7-7a. [Kagar Hayagana, Japan] 

= Caridina Voeltzkowi Lenz, 1910a: 569. [Alaotra-See (Madagaskar)] 

= Caridina japonica sikokuensis Kubo, 1938a: 91; Fig. 20. [Ryûgadô, Kôti Prefecture] 

Caridina nanaoensis Cai & N. K. Ng, 1999: 1617; Figs 6a, f, 8. [Xiaoliao stream near Xishan Town, Nan’ao 

County, Guangdong Province, China] 

Caridina natalensis Bouvier, 1925 

= Caridina africana forme natalensis Bouvier, 1925: 214. [South Africa, Umholti River, Verulum, Natal 

(according to Richard & Clark, 2009)] 

Caridina natarajani Tiwari & R.S. Pillai, 1968: 163; Fig. 1. [Trivandrum, Kerala] 

Caridina neglecta Cai & Ng, 2007: 1595; Figs 4-5. [Sungai Batang, 13 km on road from Palopo to Wotu, 

Sulawesi, Indonesia] 

Caridina nguyeni S.-Q. Li & Liang, 2002: 709; Figs 1-2. [PacBo Village, District Ha Quang, Cao Bang 

Province,Vietnam] 

Caridina nilotica (Roux, 1833) 

= Pelias Niloticus Roux, 1833: 73; Plate 7, fig. 1. [les eaux du Nil] 

= Caridina nilotica var. typica Bouvier, 1925: 146; Fig. 308. [Caire] 

Caridina norvestica Holthuis, 1965a: 35; Fig. 12. [River north of Majunga; near Majunga, small brook at 

km 530 of the highway to Tananarive; Mahajambe lake, Madagascar] 

Caridina novaecaledoniae Roux, 1926a 

= Caridina novae-caledoniae Roux, 1926a: 214; Figs 40-46. [Tchalabel; Vallée du Diahot, chemin du Col 

Poraris, 150 m; versant Est du Col Poraris, 100 m; Pemboa; Haute Tiouaka (all Nouvelle-Calédonie)] 

Caridina nudirostris Choy, 1984: 288; Fig. 1. [above Vuwa Falls, Wainisavulevu Creek, Nadrau Plateau, 

Central Viti Levu, Fiji, 17°50'30''S 178°01'30''E, altitude 800 m; below Vuwa Falls, Wainisavulevu 

Creek, 17°48'10''S 178°05'30''E, altitude 160 m; above Monasavu Falls, Nanuku Creek, Nadrau Plateau, 

Central Viti Levu, Fiji; 17°44'30''S 178°02'30''E, altitude 700 m] 

Caridina okiamnis Richard & Clark, 2009: 54; Figs 28-29. [Cameroon, Okia stream] 

Caridina okinawa Cai & Shokita, 2006b: 2154; Fig. 12. [Ingue Cave at Yomitan Village, Okinawa Island, 

Ryukyu Islands, Japan] 

Caridina oligospina Liang, Guo & Tang, 1999: 72; Fig. 3. [China, Hunan Province, Fengfang County, Muli 

village] 

Caridina opaensis Roux, 1904: 547; Plate 9, figs 8-10. [Lac Opa. Bras S.-E. de l’île, region centrale, alt. 30 m]


Caridina palawanensis Cai & Shokita, 2006a: 256; Figs 5, 6A-B. [Panitian River, upper stream, ca. 30 km 

from river mouth, Palawan Island, Philippines] 

Caridina panikkari Jalihal, Shenoy & Sankolli, 1984: 9; Figs 4-5. [Hosayellapur tank (Hirekeri), Dharwar] 

Caridina papuana Nobili, 1905d 

= Caridina Weberi var. papuana Nobili, 1905d: 481; Plate 12, fig. 1-1b. [Nova Guinea Tedesca] 

Caridina paracornuta Cai & Yuan, 1996: 391; Figs 19-21. [Chuangfengao stream, Hongyan village] 

Caridina pareparensis De Man, 1892a: 379; Plate 22, Figs 25-25b. [river near Pare Pare, Sulawesi, Indonesia; 

lectotype designated by Cai & Ng, 2009] 

Caridina parvidentata Roux, 1904 

= Caridina pareparensis var. parvidentata Roux, 1904: 545; Plate 9, figs 5-7. [Malawa Quelle, Celebes (= 

Sulawesi); lectotype designation by Cai & Ng, 2009] 

Caridina parvirostris De Man, 1892a: 375; Plate 22, fig. 24. [Flores, Fluss bei Bombang] 

Caridina parvocula Gurney, 1984: 569; Figs 1-3. [Dark zone, la Grotte d’Antsatrobonko, Ankarana Massif, 

40 km north of Ambilobé and 60 km south of Diego Suarez, northern province of Diego Suares] 

Caridina parvula von Rintelen & Cai, 2009: 392; Figs 27-29. [Lake Towuti, southwest shore, Cape Sioloya, 

02°50.7'S 121°26.32'E] 

Caridina paucidentata L.-Q. Wang & Liang, 2005: 748; Figs 1-15. [Mongzi County, Yunnan Province] 

Caridina pedicultrata Guo & Choy, 1994: 123; Figs 1-2. [stream near Huaihua City, at Xuefeng Mountain, 

Hunan, c. 27°40'N 110°11'E] 

Caridina peninsularis Kemp, 1918b 

= Caridina brachydactyla peninsularis Kemp, 1918b: 279; Fig. 10. [Botanical Garden, Penang, Malaysia; 

lectotype designation by Cai, Ng & Choy, 2007] 

Caridina petiti Roux, 1929 

= Caridina Petiti Roux, 1929: 108; Figs 13-18. [Ambila, lagunes orientales] 

Caridina pingi Yü, 1938: 294; Figs 9-10. [Amoy, Fukien] 

Caridina pingioides Yü, 1938: 298; Figs 11-12. [locality being unkown to the writer] 

Caridina plicata Liang, 2004: 282; Fig. 138. [Xita, Jiangxi] 

Caridina prashadi Tiwari & R.S. Pillai, 1971: 87; Figs 3-4. [Aberdeen (Port Blair), South Andaman, from a 

freshwater stream] 

= Caridina sakishimensis Fujino & Shokita, 1975: 99; Fig. 5. [Yonaguni I.: Arakawabana River; Spring 

of Tendabanata; Tabaru River; Ishigaki I.: Miyara River; Yonehara River; Sakutara River; Fukido 

River; Miyako I.: Izaga Cave; Bora Rover; Kume I.: Shirase River] 

Caridina pristis Roux, 1931b: 63; Figs 1-3. [alentours de Perdenyia; prés de Kandy, Ceylan] 

= Caridina pristis cruszi de Silva, 1982: 131; Fig. 3. [stream in Sinharaja forest] 

Caridina profundicola von Rintelen & Cai, 2009: 396; Figs 30-32. [Lake Towuti, Loeha Island, southwest 

shore, 02°45.58'S 121°31.14'E, on boulders in deeper water] 

Caridina propinqua De Man, 1908b: 227; Plate 19, Figs 6-6f. [Dhappa, near Calcutta] 

= Caridina blancoi Chace, 1997: 6; Fig. 2. [Philippines, near mouth of Tyabas River, Luzon, 13°54'N 

121°36'E] 

Caridina pseudodenticulata Hung, Chan & Yu, 1993: 498; Figs 9D, 13. [Yun Lin County, Ku Keng] 

Caridina pseudonilotica Richard & Clark, 2005: 722; Figs 9-10. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina pseudoserrata Đăng & Ðỗ, 2007: 1; Figs 1-2. [Bang River, Cao Bang Province] 

Caridina qingyuanensis Guo & He, 2007: 47; Figs 1-19. [Baishikeng Village, Qingyuan City, Guangdong- 

Province (ca. 23°35'N 113°10'E)] 

Caridina rajadhari Bouvier, 1918 

= Caridina Rajadhari Bouvier, 1918: 386; Figs 1-3. [Rajadhar, dans l’état de Kawarda, massif montagneux 

situé dans les provinces centrales, entre Jubbulpoor et Nagpoor; Majghaon, non loin de Rajadhar; 

Mukhi, même région] 

Caridina rangoona Cai & Ng, 2000: 939; Fig. 6. [17°09.97'N 96°99.20'E, Win Paw Hta River, near border 

between Pegu (Bago) and Yangon, Yangon State, Myanmar] 

Caridina rapaensis Edmondson, 1935b: 12; Fig. 5a-h. [freshwater stream, Rapa] 

Caridina richtersi Thallwitz, 1892


= Caridina serrata Richters, 1880: 163; Plate 17, Figs 24-27; nec Stimpson, 1860a. [Botan. Garten Pamplemousses; 

Black river] 

= Caridina richtersii Thallwitz, 1892: 27. [nomen novum for Caridina serrata Richters, 1880] 

= Caridina apiocheles Bouvier, 1904a: 134. [peut-être des Seychelles] 

= Caridina apiocheles mutation Edwardsi Bouvier, 1904a: 134. [peut-être des Seychelles] 

= Caridina Richtersi forma typica Bouvier, 1925: 203. [Type locality not indicated] 

Caridina roubaudi Bouvier, 1925 

= Caridina africana forme Roubaudi Bouvier, 1925: 217; Fig. 477. [Congo, Brazzaville; lectotype designated 

by Richard & Clark, 2009] 

Caridina rouxi De Man, 1915a: 387; Plate 27, figs 1-11. [in einem kleinen, schnell fliessenden Flusse im 

Bougainville-Gebirge, 500 m] 

Caridina rubella Fujino & Shokita, 1975: 102; Fig. 6. [Miyako I.: Izaga Cave, Morikaga Cave; Nikadori, 

Hirara City, in well] 

Caridina rubropunctata Đăng & Ðỗ, 2007: 3; Figs 3-4. [streams in Van Lang, Dong Hy District, Thai Nguyen 

Province] 

Caridina samar Cai & Anker, 2004: 240; Fig. 4-5. [Sulpan Cave, 12°02'00''N 124°55'30''E, altitude 100 m, 

Matalud, West Samar, Philippines] 

Caridina sarasinorum Schenkel, 1902: 492; Plate 8, Figs 2a-e. [Lake Poso, central Sulawesi, Indonesia; lectotype 

designation by Cai & Wowor, 2007] 

Caridina schenkeli von Rintelen & Cai, 2009: 442; Figs 60-62. [small stream, west of Lake Poso, 02°2.613'S 

120°37.311'E, on macrophytes] 

Caridina semiblepsia Guo, Choy & Gui, 1996: 66; Figs 1-4. [Dongpaoshan Cave (c. 28°44'N 109°39'E), Hunan 

Province, China] 

Caridina serrata Stimpson, 1860a: 29. [hill above Bekhers, Hong Kong island, Hong Kong; neotype designation 

by Cai & N. K. Ng, 1999] 

Caridina serratirostris De Man, 1892a: 382; Plate 23, Figs 28-28e. [Bangkalanvir River, Saleyer; lectotype 

designation by Richard & Clark, 2010a] 

Caridina shenoyi Jalihal & Sankolli in Jalihal, Shenoy & Sankolli, 1984: 21; Figs 9-11. [Malaprabha river– 

near old bridge, Khanapur (Belgaum District), India] 

Caridina shilinica Liang & Cai, 2000: 177; Fig. 1. [Shilin (Stone forest) Tour Resort in Lunan County Yunnan 

Province, southern China] 

Caridina similis Bouvier, 1904a: 135. [Mahé, Seychelles; lectotype designation by Marquet & Keith, 2008] 

= Caridina brevirostris forma typica Bouvier, 1912b: 916. [Seychelles] 

= Caridina brevirostris forma Gardineri Bouvier, 1912b: 916. [Seychelles] 

Caridina simoni Bouvier, 1904a 

= Caridina Simoni Bouvier, 1904a: 131. [Kandee, l’île de Ceylan] 

= Caridina costai de Silva, 1982: 126; Fig. 2. [three streams in Sinharaja forest, Sri Lanka] 

Caridina sodenensis Richard & Clark, 2009: 42; Figs 20-21. [Cameroon, Lake Soden, West Cameroons] 

Caridina solearipes Guo & De Grave, 1997: 127; Figs 2-3. [small stream near Dabaozi, Jingzhou County, 

Hunan province, circa 26°41'N 09°25'E, altitude between 400-900 m] 

Caridina songtaoensis Liang, 2004: 270; Fig. 32. [Songtao, Guizhou] 

Caridina spathulirostris Richters, 1880: 163; Plate 17, fig. 28. [Botanischer Garten Pamplemousses] 

Caridina spelunca Choy, 1996: 103; Fig. 1. [Old Napier Downs Cave, Western Australia, Australia, 17°14'S 

124°39'E] 

Caridina sphyrapoda Liang & Zhou, 1993: 236; Figs 4(1)-4(2). [Longdong River in Laibing County, 

Guangxi, southern China] 

Caridina spinalifrons Guo & De Grave, 1997: 124; Fig. 1. [stream near Sangzhi city, Hunan Province, circa 

29°10'N 110°16'E, 650 m altitude] 

Caridina spinata Woltereck, 1937: 221; Fig. 1.3; Table 1. [Lake Towuti, estuary of Sungei Batuopa, about 2 

km south of Timampu, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina spinipoda Liang, Hong & Yang, 1990: 161; Figs 1-9. [Mianzhu County, 31°20'N 104°11'E, Sichuan 

Province]


Caridina spinosipes Liang, Guo & Tang, 1999: 71; Fig. 2. [China, Hunan Province, Fengfang County, Muli 

village] 

Caridina spinula Choy & Marshall, 1997: 31; Figs 1c, 2g-h, 4f-q. [east flowing first order tributary of 

Leo Creek, near crossing of Leo Creek Mine Road, 13°44.6'S 143°21.5'E, Nesbit River catchment, 

McIlwraith Range, Cape York] 

Caridina spongicola Zitzler & Cai, 2006: 271; Figs 1-3. [Indonesia, Sulawesi Selatan, Lake Towuti, west 

shore, outlet bay, west of Cape Tokaluku, 02°47.261'S 121°23.17'E] 

Caridina steineri Cai, 2005: 314; Figs 2-3. [Lakata Zafera, western Madagascar, 19°45.075'S 45°11.436'E, 

stream pool at far end of cave] 

Caridina striata von Rintelen & Cai, 2009: 410; Figs 39-41. [Indonesia, Sulawesi Selatan, Lake Towuti, 

north shore, 02°38.56'S 121°27.82'E, on rocks] 

Caridina subventralis Richard & Clark, 2005: 725; Figs 11-12. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina sulawesi Cai & Ng, 2009: 1103; Figs 6-8. [Sungai Beru, Kappang, Maros, Sulawesi Selatan] 

Caridina sumatianica Cai & Yuan, 1996: 395; Figs 22-23. [Sumatian stream, near Hongyan village] 

Caridina sumatrensis De Man, 1892a 

= Caridina Weberi var. sumatrensis De Man, 1892a: 375; Plate 22, fig. 23g. [flüssen des unteren Battaklandes 

bei Deli, an der Ostküste von Sumatra] 

Caridina sundanella Holthuis, 1978a: 32; Figs 11-12. [Waikamburu Brook, 4 km N. of Waimangura, West 

Sumba, about 250 m above sea level] 

Caridina susuruflabra Richard & Clark, 2009: 60; Figs 32-33. [South Africa, Eoshowe, Zululand] 

Caridina temasek Choy & Ng, 1991: 266; Figs 2-5. [freshwater stream near Sime road, MacRitchie catchment 

area, Singapore, ca. 01°20'14''N 103°48'47''E] 

Caridina tenuirostris Woltereck, 1937: 224; Fig. 1.8; Table 1. [Lake Towuti, southwest shore, west of Cape 

Tetetu, 02°54.13'S 121°23.78'E; neotype designation by von Rintelen & Cai, 2009] 

= Caridina Towutensis Woltereck, 1937: 220; Fig. 1.2; Table 1. [Towuti-Süd] 

Caridina thambipillai Johnson, 1961: 138; Figs 25-35. [Sungei Putat near the pumping station, Malacca] 

Caridina thermophila Riek, 1953: 120; Fig. 10. [Muttaburra, western Queensland] 

Caridina thomasi von Rintelen, Karge & Klotz, 2008: 2247; Figs 2-3. [Indonesia, Central Sulawesi, Banggai 

islands, Peleng, west peninsula, east of Alani, river with lake-like extension, 01°28.315'S 122°52.473'E] 

Caridina timorensis De Man, 1893: 300; Plate 8, fig. 6. [freshwater lake of Nefko on the island of Timor] 

Caridina togoensis Hilgendorf, 1893a: 156. [Togo, Bismarksburg; lectotype designation by Richard & 

Clark, 2009] 

= Caridina togoënsis var. stuhlmanni Hilgendorf, 1898: 35. [Undussuma und Bach bei Undussuma] 

= Caridina togoensis var. Decorsei Bouvier, 1904a: 131. [Congo français, rivière prés de Mpoko; région 

Chari-Tchad, Krébédje (Fort-Sibut) et Binguétou, rivière Gribingui; Bangoran] 

= Caridina togoensis var. breviatus Lenz, 1910b: 131. [Urwaldbache N.W. von Beni] 

= Caridina africana forme ægyptiaca Bouvier, 1925: 214. [Caire] 

= Caridina togoensis var. Kasaiensis De Man, 1925: 5; Figs 2 n1 -2 n3 . [Kamaiembe; Kondué] 

= Caridina togoensis var. Kwamouthensis De Man, 1925: 9; Fig. 2 o1 -2 012 . [Kwamouth au confluent du 

fleuve Congo et de la rivière Kasaï] 

= Caridina togoensis var. Schoutedeni De Man, 1925: 20; Figs 2 q1 -2 q21 . [les rivières du bassin du Chiloango; 

Buto-Polo dans la rivière Mapanga] 

Caridina tonkinensis Bouvier, 1919: 331. [Tonkin] 

Caridina trifasciata Yam & Cai, 2003: 277; Figs 1, 3-5. [Tsak Yue Wu, New Territories, Hong Kong] 

Caridina troglodytes Holthuis, 1978b: 214; Figs 3-4. [Danmin Cave near Konogusgus, New Ireland, in 

subterranean stagnant pool, 0.3-0.6 m] 

Caridina troglophila Holthuis, 1965a: 37; Fig. 13. [Ambovonomby cave, Namoroka, NW Madagascar] 

Caridina tumida L. Wang, Liang & F. Li, 2008: 54; Figs 5-6. [Yingde County, Guangdong Province, 

N23°54.4' E113°15.1'] 

Caridina typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= C.[aridina] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363; Plate 25bis, Figs 

4-5. [Type locality unknown, probably l’île Maurice according to Bouvier, 1925]


= Caridina exilirostris Stimpson, 1860a: 29. [Okuma River, Okinawa Island, Rykyu islands; neotype 

designation by Cai, Ng, Shokita & Satake, 2006] 

= Caridina siamensis Giebel, 1863: 329. [Siam] 

? = Caridina Spencebatei De Man, 1892a: 371. [?Cap Verdischen Inseln] 

= Caridina typus forme typica Bouvier, 1925: 250. [l’île Maurice] 

= Caridina typus forme caledonica Bouvier, 1925: 253; Figs 296-297. [Nouvelle-Calédonie] 

Caridina uminensis Đăng & Ðỗ, 2007: 6; Figs 5-6. [U Minh Thuong wetland, Ca Mau Province] 

Caridina umtatensis Richard & Clark, 2009: 63; Figs 34-35. [South Africa, Kraal Dam, Umtata] 

Caridina unca Gurney, 1984: 579; Figs 8-11. [Dark zone, Grotte d’Antsatrobonko, Andarana Massif, northern 

province of Diego Suarez] 

Caridina valencia Cai, Choy & Ng, 2009: 78; Fig. 9. [spring Anislaq, outside Valencia] 

Caridina venusta L. Wang, Liang & F. Li, 2008: 52; Figs 3-4. [Yingde County, Guagndon Province, 

N23°54.3' E113°15.8'] 

Caridina vietriensis Đăng & Ðỗ, 2007: 9; Figs 7-8. [confluence of rivers in Viet Tri City, Phu Tho Province] 

Caridina villadolidi Blanco, 1939a: 389; Plate 1. [Laoag River, Laoag, Ilocos Norte Province, Luzon] 

= Caridina typus var. longirostris De Man, 1892a: 370; Plate 22, fig. 22f-i; nec C.[aridina] longirostris H. 

Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]. [Flores, as einem Flusse bei Reo; auch auf 

Celebes und Saleyer] 

Caridina vitiensis Borradaile, 1899: 1003; Plate 63, figs 3-3a. [Suva, Tamavua River, Viti Levu, Fiji] 

= Caridina vitiensis var. canacorum Roux, 1926a: 199; Figs 32-36. [Nouvelle-Calédonie, Canala] 

Caridina weberi De Man, 1892a 

= Caridina Weberi De Man, 1892a: 371; Plate 22, figs 23-23g. [Kotting; Fluss bei Mbawa, oberhalb des 

Wasserfalls; Fluss bei Bombang (all Flores); Fluss bei Palopo, Luwu; Wasserfall bei Bantimurong 

unweit Maros; Fluss bei Pare-pare (all Celebes); Saleyer, Fluss Bangkalan] 

= Caridina weberi var. typica Bouvier, 1925: 243; Figs 562-566. [Flores, Kotting, environs de Mbawa, 

de Bombang; Java; Kifa-juc, Ile Engano] 

? = Caridina weberi var. keiensis Roux, 1911: 85. [Elat, île de Grand-Kei; ruisseau entre Elat et Ohinangan; 

Warka, île de Grand-Kei; ruisseau entre Elat et Ohilim, île de Grand-Kei] 

Caridina williamsi Cai & Ng, 2000: 933; Figs 4-5. [17°48.11'N 96°09.20'E, Balar stream, 12 km Yangon to 

Mandalay highway, Htauk Kyant, Yangon State, Myanmar] 

Caridina woltereckae Cai, Wowor & Choy, 2009: 19; Fig. 3. [Cape Larona, near Sungai Larona outlet, Lae 

Towuti, Sulawesi, Indonesia] 

Caridina wumingensis Cai & N.K. Ng, 1999: 1632; Figs 18-19. [cave near Wuming, Guangxi Province, 

China] 

Caridina wyckii (Hickson, 1888) 

= Atya Wyckii Hickson, 1888: 358; Plates 13-14. [Lake Tondano, Minahassa, North Celebes] 

= Caridina nilotica var. minahassae De Man, 1902: 895. [Minahassa, Celebes] 

Caridina xiangnanensis X.-Y. Liu, Guo & Yu, 2006: 44; Fig. 1. [near Lingxiu Village, Rucheng County, Hunan 

Province (ca. 25°33'N 113°40'E)] 

Caridina xiphias Bouvier, 1925 

= Caridina nilotica var. xiphias Bouvier, 1925: 149; Figs 310-312. [district d’Ambatondrazaka, Madagascar] 

= Caridina nilotica var. stylirostris Bouvier, 1925: 148; Fig. 309. [Madagascar] 

= Caridina gladiifera Roux, 1929: 306; Figs 1-6. [Périnet, forêt, Madagascar] 

Caridina yilong Cai & Liang, 1999: 76; Fig. 3. [Yilong Lake, Shiping County, Yunnan] 

Caridina yulinica Cai & N.K. Ng, 1999: 1620; Figs 9-11. [Niuwo cave near Kuiyang Town, Yulin County, 

Guangxi Province, China] 

Caridina yunnanensis Yü, 1938: 304; Figs 14-16. [Mountain stream at Hsün-tien, Yunnan Province] 

= Caridina impensa Cai & Ng, 2001a: 223; Figs 8-9. [Xilong Tang (Pond), Chengjiang County, Yunnan] 

Caridina zebra Short, 1993: 62; Figs 1-3. [O’Leary Creek, 17°50.7'S 145°37.7'E, altitude ca. 750 m] 

Caridina zeylanica Arudpragasam & Costa, 1962: 17; Fig. 4. [Nawala, suburb of Colombo] 

= Caridina nilotica var. veliensis R.S. Pillai, 1964: 44. [Veli Lake, Trivandrum]


Caridina zhejiangensis Liang & Zheng, 1985 

= Caridina pingi zhejiangensis Liang & Zheng, 1985: 321; Fig. 1(1c-7c). [Baisha Village, Jiande County, 

Zhejiang Province, China] 

Caridina zhongshanica Liang, 2004: 182; Fig. 88. [Zhongshan, Guangdong] 

Caridinides Calman, 1926 

= Caridinides Calman, 1926 (type species Caridinides wilkinsi Calman, 1926, by monotypy, gender 

masculine) 

Caridinides wilkinsi Calman, 1926: 242; Figs 1-2. [Olive River, Temple Bay, east coast of Cape York Peninsula] 

Caridinopsis Bouvier, 1912c 

= Caridinopsis Bouvier, 1912c (type species Caridinopsis Chevalieri Bouvier, 1912c, by monotypy, 


1957) 

Caridinopsis chevalieri Bouvier, 1912c 

= Caridinopsis Chevalieri Bouvier, 1912c: 300; Figs 1-4. [région du Haut Niger, à Sampouyara, bassin 

de la Makowa] 

= Caridinopsis brevinaris Holthuis, 1956a: 56. [Garrigues Cave near Sougouéta, French Guinea] 

Dugastella Bouvier, 1912d 

= Dugastella Bouvier, 1912d (type species Dugastella marocana Bouvier, 1912d, by monotypy, gender 

feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Dugastella marocana Bouvier, 1912d: 993. [la source de Settat] 

Dugastella valentina (Ferrer Galdiano, 1924) 

= Atyaephira valentina Ferrer Galdiano, 1924: 210; Figs 1, 3. [Laguna de Almenara (Castellón) y Albufera 

de Valencia] 

= Dugastella marocana var. hispanica Balss, 1925b: 206; Figs 1-4. [Silla, bei Valencia, Acequia communera, 

1.5 km v. Albufera] 

Edoneus Holthuis, 1978b 

= Edoneus Holthuis, 1978b (type species Edoneus atheatus Holthuis, 1978b, by original designation 


Edoneus atheatus Holthuis, 1978b: 220; Figs 5-6. [the Philippines, Luzon, Quirino province, Aglipay Municipality, 

Barrio Palasian, cave near Sitio Disiluad; see Balete & Holthuis, 1992] 

Edoneus erwini Cai & Husana, 2009: 54; Figs 2-3. [Bantakay Cave, station 2, Luzon, the Philippines] 

Edoneus marulas Cai & Husana, 2009: 60; Figs 6-7. [at middle of Marulas Cave, Luzon, the Philippines] 

Edoneus sketi Cai & Husana, 2009: 57; Figs 4-5. [small pool in Bantakay cave, Luzon, the Philippines] 

Gallocaris Sket & Zakšek, 2009 

= Gallocaris Sket & Zakšek, 2009 (type species Troglocaris Schmidti inermis Fage, 1937, by original 


Gallocaris inermis (Fage, 1937) 

= Troglocaris Schmidti inermis Fage, 1937: 215; Figs 1-6. [grotte de Cambous, près Saint-Hippolyte-du- 

Fort, département du Gard] 

Halocaridina Holthuis, 1963b 

= Halocaridina Holthuis, 1963b (type species Halocaridina rubra Holthuis, 1963b, by monotypy, gender 

feminine) 

Halocaridina palahemo Kensley & Williams, 1986: 429; Figs 9-11. [Hawaii Island, Ka Lae, Lua o Palahemo, 

lava tube pool, 18°55'N 155°42'W] 

Halocaridina rubra Holthuis, 1963b: 262; Fig. 1. [pool in the base of Lohena Rock, between the deserted 

villages of Wai-o-ahu-kini and Kaili-kii, W. of Ka Lae or South Point, Kau district, island of Hawaii]


Halocaridinides Fujino & Shokita, 1975 

= Halocaridinides Fujino & Shokita, 1975 (type species Halocaridina (Halocaridinides) trigonophthalma 

Fujino & Shokita, 1975, by monotypy, gender masculine) 

= Palauatya C.W.J. Hart, 1980 (type species Palauatya dasyomma C.W.J. Hart, 1980, by original designation 


Halocaridinides fowleri Gordon in Gordon & Monod, 1968 

= Parisia (?) fowleri Gordon in Gordon & Monod, 1968: 514; Figs 30-31. [subterranean lake, Kufile, 

Zanzibar] 

Halocaridinides trigonophthalma (Fujino & Shokita, 1975) 

= Halocaridina (Halocaridinides) trigonophthalma Fujino & Shokita, 1975: 106; Figs 7-8. [Okinawa Island: 

Kaneshi, Nakijin, in well, 10 m; Aja, Naha City, in well] 

= Palauatya dasyomma C.W.J. Hart, 1980: 481; Figs 1-31. [anchialine lake on Anguar Island, Palau, 

Caroline Islands, 6°54'49''N 134°08'12''E] 

Jolivetya Cals, 1986 

= Jolivetya Cals, 1986 (type species Jolivetya foresti Cals, 1986, by monotypy, gender feminine) 

Jolivetya foresti Cals, 1986: 387; Fig. 1. [Nouvelle-Bretagne, la grotte de Pogo Ngim, lieu d’accès au réseau 

souterrain d’Arais, dans un karst à dolines, à quelques dizaines de kilomètres du littoral de la Nouvelle-Bretagne] 

Jonga C.W.J. Hart, 1961 

= Jonga C.W.J. Hart, 1961 (type species Ortmannia Serrei Bouvier, 1909a, by original designation and 


Jonga serrei (Bouvier, 1909a) 

= Ortmannia Serrei Bouvier, 1909a: 332. [Cuba] 

Lancaris Cai & Bahir, 2005 

= Lancaris Cai & Bahir, 2005 (type species Caridina singhalensis Ortmann, 1894, by original designation, 


Lancaris kumariae (de Silva, 1990) 

= Caridina kumariae de Silva, 1990: 9; Fig. 1. [small stream near railway station at Rozella, Sri Lanka, 

6°56'-6°57'N 80°33'-80°35'E] 

Lancaris singhalensis (Ortmann, 1894) 

= Caridina singhalensis Ortmann, 1894: 11; Plate 1, fig. 2. [Ceylon, Newera Elya] 

Limnocaridella Bouvier, 1913b 

= Limnocaridella Bouvier, 1913b (type species Limnocaridina Alberti Lenz, 1910b, by monotypy, gender 

feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

Limnocaridella alberti (Lenz, 1910b) 

= Limnocaridina Alberti Lenz, 1910b: 132; Plate 3, Figs 6-9. [Albert-See] 

Limnocaridina Calman, 1899a 

= Limnocaridina Calman, 1899a (type species Limnocaridina tanganyikæ Calman, 1899a, by monotypy, 


1957) 

Limnocaridina iridinae Roth-Woltereck, 1958: 188; Figs 1-5. [Tanganjika-See bei Mpulungu (Nordrhodesien, 

Abercorn District), 3 m, aus dem Kiemenraum einer Iridina spekei] 

Limnocaridina latipes Calman, 1906a: 196; Plate 12, figs 23-29. [Mbete, shallow water amongst rocks; near 

mouth of Lofu, on rocks, shallow water; Kalambo, tow-netting; Tembwi, on rocks, shallow water 

(all Lake Tanganyika)] 

Limnocaridina parvula Calman, 1906a: 193; Plate 11, figs 9-14. [Kasawa, tow-netting; Kalambo, tow-netting; 

Karema tow-netting, surface (all lake Tangaynika)]


Limnocaridina retiarius Calman, 1906a: 192; Plate 11, figs 2-8. [Lake Tanganyika, Mbete, taken in shrimpnet, 

shore wading] 

Limnocaridina similis Calman, 1906a: 195; Plate 12, figs 15-22. [Lake Tanganyika, Kalambo, tow-netting; 

Rusisi River, close to Tanganyika] 

Limnocaridina socius Calman, 1906a: 196; Plate 12, figs 30-37. [Niamkolo Harbour, 3 fms; Utinta, 10 fms; 

Kirando, about 8 fms (all Lake Tangyanika)] 

Limnocaridina spinipes Calman, 1906a: 197; Plate 13, figs 38-44. [Niamkolo Harbour, 3 fms; Kirando, 8 

fms; Utinta, about 15 fms (all Lake Tanganyika)] 

Limnocaridina tanganyikae Calman, 1899a 

= Limnocaridina tanganyikæ Calman, 1899a: 704; Plate 39, figs 1-2, 4-9a; Plate 40, figs 10-19. [Lake 

Tanganyika, shallow water] 

Mancicaris liang, Guo & Tang, 1999 

= Mancicaris Liang, Guo & Tang, 1999 (type species Mancicaris sinensis Liang, Guo & Tang, 1999, by 

original designation and monotypy, gender feminine) 

Mancicaris sinensis Liang, Guo & Tang, 1999: 70; Fig. 1. [Tianxinpu village, Tianxin district, Lanshan 

County, Hunan Province, China] 

Marosina Cai & Ng, 2005 

= Marosina Cai & Ng, 2005 (type species Marosina brevirostris Cai & Ng, 2005, by original designation, 


Marosina brevirostris Cai & Ng, 2005: 131; Fig. 1. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi 

Selatan, Indonesia] 

Marosina longirostris Cai & Ng, 2005: 133; Figs 2-4. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi 

Selatan] 

Micratya Bouvier, 1913b 

= Calmania Bouvier, 1909b (type species Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 

1855-1856], by monotypy, gender feminine; an invalid junior homonym of Calmania Laurie, 1906 

(Crustacea Brachyura) and Calmania Nobili, 1907 (a junior subjective synonym of Brachycarpus 

Spence Bate, 1888); name placed on the Official Index of Rejected and Invalid Generic Names in 


= Micratya Bouvier, 1913b (type species Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 

1855-1856], by monotypy, gender feminine; name placed on the Official List of Generic Names in 


= Balssiola Strand, 1922 (nomen novum for Calmania Bouvier, 1909b, gender feminine; name placed 


Micratya poeyi (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) 

= Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xviii; Plate 2, Figs 7, 7a-b. 

[Cuba] 

Neocaridina Kubo, 1938a 

= Neocaridina Kubo, 1938a (type species Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833- 

1850], by original designation, gender feminine) 

Neocaridina anhuiensis (Liang, Zhu & Xiong, 1984) 

= Caridina denticulata anhuiensis Liang, Zhu & Xiong, 1984: 251; Figs 1-7. [Taiping County, Anhui 

Province] 

Neocaridina bamana Liang, 2004: 112; Fig. 55. [Bama, Guangxi] 

Neocaridina brevidactyla Liang, Chen & W.-X. Li, 2005: 532; Figs 3-43. [caves of Libo County (25°41'N 

107°83'E), Maolan Nature Sanctuary of Guizhou] 

Neocaridina curvifrons (Liang, 1979) 

= Caridina curvifrons Liang, 1979: 118; Figs 1-10. [Baisha, Zhejiang Province]


Neocaridina denticulata davidi (Bouvier, 1904a) 

= Caridina Davidi Bouvier, 1904a: 133. [Inkiafou, Chensi méridonale] 

Neocaridina denticulata denticulata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 8. [Japan] 

= Caridina denticulata sinensis Kemp, 1918b: 287; Fig. 11c-d. [creeks and irrigation channels at the 

edge of the Tai Hu lake in Kiangsu province] 

Neocaridina denticulata moganica Liang, 2004: 82; Fig. 41. [Mogashan, Zhejiang] 

Neocaridina euspinosa Cai, 1996: 150; Figs 15-16. [Yanshan mountain near Cuilin City (25°20'N 110°18'E), 

Guangxi Zhuang Autonomous Region, China] 

Neocaridina fukiensis (Liang & Yan, 1977) 

= Caridina spinosa fukiensis Liang & Yan, 1977: 222; Figs 9-11. [Gutian, Shnaghang County (rivulet); 

Xinquan, Liancheng County (pool); Luokou and Ansha, Yongan County (pool) (all Fujian Province)] 

Neocaridina gracilipoda Liang, 2004: 88; Fig. 44. [Junlian, Sichuan] 

Neocaridina heteropoda heteropoda Liang, 2002b 

= Neocaridina heteropoda Liang, 2002b: 167; Fig. 1a-h. [Baisha Village, Jiande County, Zhejiang Province, 

China] 

Neocaridina heteropoda koreana Kubo, 1938a 

= Neocaridina denticulata koreana Kubo, 1938a: 81; Figs 5c, 7g-m, 7g’-m’, 12. [Huzan, Korea] 

Neocaridina heteropoda luoyangensis Cai, 1996 

= Neocaridina denticulata luoyangensis Cai, 1996: 139, Figs 7-8. [from a mountain stream near Luoyan 

City (34°40'N 112°20'E), Henan Province, China] 

Neocaridina hofendopoda (Shen, 1948) 

= Caridina hofendopoda Shen, 1948: 122; Plate 13, Figs f-l. [Kweilin] 

Neocaridina homospina Liang, 2002b 

= Neocaridina euspinosa homospina Liang, 2002b: 169; Fig. 2. [China, Hunan Province, Shaodong County] 

Neocaridina iriomotensis Naruse, Shokita & Cai, 2006: 26; Figs 1, 2. [upper reaches of the Nishifunatsuki, 

Nakama River, Iriomote Island] 

Neocaridina ishigakiensis (Fujino & Shokita, 1975) 

= Caridina denticulata ishigakiensis Fujino & Shokita, 1975: 95; Figs 3-4. [Yonehara River; Miyara 

River; Sakutara River, Arakawa River (all Ishigaki Island)] 

Neocaridina ketagalan Shih & Cai, 2007: 687; Figs 5-6, 7D-E. [Sijhih, Taipei Co., Taiwan] 

Neocaridina keunbaei (H.S. Kim, 1976) 

= Caridina denticulata keunbaei H.S. Kim, 1976: 155; Figs 1-3. [upper stream of Cheonji fall, Jeju Island] 

Neocaridina linfenensis Cai, 1996 

= Neocaridina denticulata linfenensis Cai, 1996: 137; Figs 5-6. [near Linfen City, 36°5'N 111°30'E, Shanxi 

Province, China] 

Neocaridina longipoda (Cai, 1995a) 

= Caridina longipoda Cai, 1995a: 166; Figs 1-15. [Jinbianxi stream, Zhangjiajie National Forest Park, 

Hunan Province] 

Neocaridina palmata bosensis Cai, 1996: 145; Figs 11-12. [Longlin County (25°15'N 105°22'E), in Guangxi 

Zuang Autonomous Region, China] 

Neocaridina palmata luodianica Liang, 2004: 110; Fig. 54. [Luodian, Guizhou] 

Neocaridina palmata meridionalis Liang, 2004: 108; Fig. 53. [Luoding, Guangdong] 

Neocaridina palmata palmata (Shen, 1948) 

= Caridina palmata Shen, 1948: 120; Plate 12. [Sha-ping-pa Chungking] 

= Caridina elongata Shen, 1948: 121; Plate 13, figs a-e. [Kweilin] 

= Caridina denticulata vietnamensis Đăng, 1967: 157; Fig. 4. [Cao Bang province; Lang Son province] 

Neocaridina saccam Shih & Cai, 2007: 682; Figs 2-4, 7A-B. [mountain stream at Longci, Tainan Co., 

Taiwan] 

Neocaridina spinosa (Liang, 1964) 

= Caridina spinosa Liang, 1964: 187; Figs 1-12. [mountain stream, Lantang, Tzechin Hsien, Kwangtung]


Neocaridina xiapuensis Zheng, 2002: 19; Fig. 1. [Xiapu County, 26°55'N 120°E, Fujian Province, China] 

Neocaridina zhangjiajiensis Cai, 1996: 147; Figs 13-14. [Zhangjiajie National Forest Park in Dayong City, 

Hunan Province of China] 

Neocaridina zhoushanensis Cai, 1996 

= Neocaridina denticulata zhoushanensis Cai, 1996: 142; Figs 9-10. [Dinhai County in Zhoushan Island 

(30°02'N 122°08'E), Zhejiang Province, China] 

Palaemonias Hay, 1902 

= Palæmonias Hay, 1902 (type species Palæmonias ganteri Hay, 1902, by monotypy, gender masculine; 


Palaemonias alabamae Smalley, 1961: 127; Fig. 1. [Shelta Cave, Huntsville, Madison County, Alabama] 

Palaemonias ganteri Hay, 1902 

= Palæmonias ganteri Hay, 1902: 179. [Roaring River passage, Mammoth Cave] 

Paracaridina liang, Guo & Tang, 1999 

= Paracaridina Liang, Guo & Tang, 1999 (type species Caridina longispina Guo, He, Xu & Gui, 1992, 

by original designation and monotypy, gender feminine) 

Paracaridina chenxiensis Guo & De Grave, 2004: 203; Figs 2-3. [small stream at an elevation of 450 m near 

Houmachong village, Chenxi County, ca. 27°47'N 110°16'E] 

Paracaridina guizhouensis (Liang & Yan, 1986a) 

= Caridina guizhouensis Liang & Yan, 1986a: 199; Fig. 2. [Maopo, Yuping County, Guizhou] 

Paracaridina leptocarpa (Liang & Zheng, 1988) 

= Caridina leptocarpa Liang & Zheng, 1988: 15; Figs 1-9. [Fuzhou, 26°N 119°E, Min River, Fujian] 

Paracaridina longispina (Guo, He, Xu & Gui, 1992) 

= Caridina longispina Guo, He, Xu & Gui, 1992: 717; Figs 1-9. [Yuelu Hills, Changsha, China, 112°56'E 

28°10'N] 

Paracaridina zijinica Liang, 2002b: 171; Fig. 3. [Zijin, Guangdong, southern China] 

Paratya Miers, 1882 

= Paratya Miers, 1882 (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850], by 

monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

470 in 1957 and in Direction 85 in 1957) 

= Xiphocaridina Bouvier, 1909b (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833- 

1850], designated by Holthuis, 1955b, gender feminine; name placed on the Official Index of Rejected 

and Invalid Names in Zoology in Opinion 470 in 1957) 

= Xiphatyoida Roux, 1915 (established without included nominal species; type species Paratya 

(Xiphatyoida) typa Roux, 1926a, designated by Roux, 1926a, gender feminine) 

Paratya annamensis Balss, 1924: 45. [Annam, Phuc Son] 

Paratya australiensis Kemp, 1917b: 303; Fig. 5. [Seven Hills, near Sydney; neotype designation by W.D. 

Williams & Smith, 1979] 

= Paratya australiensis arrostra Riek, 1953: 114, Fig. 2. [Terrors Creek, Dayboro, Queensland] 

= Paratya atacta Riek, 1953: 114; Fig. 3. [Upper Nerang River, south Queensland] 

= Paratya atacta adynata Riek, 1953: 115; Fig. 4. [small creek in upper reaches of Middle Harbour, 

Sydney, N.S.W.] 

= Paratya tasmaniensis Riek, 1953: 115; Fig. 5. [small stream at Kingston, Tasmania] 

Paratya boninensis Satake & Cai, 2005: 306; Figs 1-2. [Oku-ohotaki river, a tributary of the west inlet 

stream of Shigure Dam, 210-220 m above sealevel, Chichijima Island, Ogasawara Islands, Japan] 

Paratya borealis Volk, 1938: 123; Figs 1-4. [Uluncha River, southern part of the Peter the Great Basin, Japan 

Sea] 

Paratya bouvieri Roux, 1926a 

= Paratya (Xiphatyoida) bouvieri Roux, 1926a: 189; Figs 1-5. [Nouvelle-Calédonie, Rivière du Mt. Panié] 

= Paratya (Xiphatyoida) bouvieri ngoiensis Roux, 1926a: 190; Figs 6-8. [Nouvelle-Calédonie, Vallée de 

Ngoï, 200 m alt.]


Parataya caledonica Roux, 1926a 

= Parataya (Xiphatyoida) caledonica Roux, 1926a: 192; Figs 9-16. [Nouvelle-Calédonie, Lac en 8] 

= Paratya caledonica var. magna Roux, 1926a: 194; Figs 17-19. [Nouvelle-Calédonie, Lac en 8] 

Paratya compressa (De Haan, 1844 [in De Haan, 1833-1850]) 

= Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 7 (1844); 186, Plate O (1849). 

[Japan; lectotype designation by Yamaguchi & Baba, 1993] 

Paratya improvisa Kemp, 1917b 

= Paratya compressa improvisa Kemp, 1917b: 299; Figs 2-3. [Lake Haruna, near Ikao, altitude about 

3000ft] 

Paratya curvirostris (Heller, 1862a) 

= Caridina curvirostris Heller, 1862a: 525. [Aukland] 

= Palæmon (Leander) fluviatilis Thomson, 1879: 231. [Waikato River; Taieri River; lagoons in Taieri 

plain] 

Paratya howensis Roux, 1926b 

= Paratya (Xiphatyoida) howensis Roux, 1926b: 244. [Big Creek, Lord Howe Island] 

Paratya intermedia Roux, 1926a 

= Paratya caledonica var. intermedia Roux, 1926a: 195; Figs 20-22. [Nouvelle-Calédonie, Plaine des 

Lacs, 200 m alt.] 

Paratya martensi Roux, 1925: 146. [Insel Adenare bei Flores] 

Paratya norfolkensis Kemp, 1917b 

= Paratya australiensis norfolkensis Kemp, 1917b: 305; Fig. 6. [west side of Norfolk Island] 

Paratya typa Roux, 1926a 

= Paratya (Xiphatyoida) typa Roux, 1926a: 196; Figs 23-31. [Nouvelle-Calédonie, La Madeleine, Plaine 

des Lacs, 200 m] 

Parisia Holthuis, 1956a 

= Parisia Holthuis, 1956a (type species Caridina microphthalma Fage, 1946, by original designation, 


Parisia dentata Gurney, 1984: 584; Figs 12-14. [Dark zone, Grotte d'Andafiabe, Ankarana Massif, 40 km 

north of Ambilobé and 60 km south of Diego Suarez, northern province of Diego Suarez] 

Parisia deharvengi Cai & Ng, 2009: 1110; Fig. 10. [Gua (cave) Tanette, Kappang, Maros, Sulawesi] 

Parisia edentata Holthuis, 1956a: 55. [southern part of Antsingly Mts., near Bekopaka, Mahilaka Province, 

W Madagascar] 

Parisia gracilis W.D. Williams, 1964: 94; Figs 1, 2A-H. [caves 16 miles south of of Katherine, Northern 

Territories (approx. 14°30'S 132°E)] 

Parisia holthuisi Cai, 2010c: 174; Figs 1-2. [Tigibi, Tari subdistrict, Southern Highlands, Territory of Papua 

Australian New Guinea (= Papua New Guinea), 1600 m alt.] 

Parisia macrophora Cai & Anker, 2004: 253; Figs 12-14. [Nalubog Cave, Quezon Province, Luzon, Philippines] 

Parisia macrophthalma Holthuis, 1956a: 55. [Grotte des Fanihy, Ankarana Mts., N of Ambilobé, N.-W. 

Madagascar] 

Parisia microphthalma (Fage, 1946) 

= Caridina microphthalma Fage, 1946: 324; Figs 1-2. [grotte des Fanihys, Ankarana] 

Parisia unguis W.D. Williams, 1964: 99; Figs 2L-S, 3. [caves 16 miles south of of Katherine, Northern Territories 

(approx. 14°30'S 132°E)] 

Potimirim Holthuis, 1954 

= Potimirim Holthuis, 1954 (type species Caridina mexicana de Saussure, 1857a, by original designation, 


Potimirim americana (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) 

= Caridina americana Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, Figs 13- 

13a. [la Isla de Cuba]


Potimirim glabra (Kingsley, 1878a) 

= Atyoida glabra Kingsley, 1878a: 93. [Polvon and Corcuera, west coast of Nicaragua] 

= Potimirim brasiliana Villalobos F., 1960b: 275; Figs 1-5. [Río Ariró, Angra dos Reis, Edo. De Río de 

Janeiro, Brasil] 

Potimirim mexicana (de Saussure, 1857a) 

= Caridina mexicana de Saussure, 1857a: 505. [Vera-Cruz] 

Potimirim potimirim (Müller, 1881) 

= Atyoida Potimirim Müller, 1881: 117; Figs 1, 8-15. [Itajahy] 

Puteonator Gurney, 1987 

= Puteonator Gurney, 1987 (type species Puteonator iraqiensis Gurney, 1987, by original designation 


Puteonator iraqiensis Gurney, 1987: 162; Figs 1-4. [southern Iraq, Samawa, anchialine habitat at 160 m] 

Pycneus Holthuis, 1986b 

= Pycneus Holthuis, 1986b (type species Pycneus morsitans Holthuis, 1986b, by original designation 


Pycneus morsitans Holthuis, 1986b: 105; Figs 1-2. [cave at Munjingerra, Gibson Desert, Western Australia, 

at about 22°30'S 124°10'E] 

Pycnisia Bruce, 1992a 

= Pycnisia Bruce, 1992a (type species Pycnisia raptor Bruce, 1992a, by original designation and monotypy, 


Pycnisia bunyip Suzuki & Davie, 2003: 447; Figs 1-4. [Forbes Inferno Cave, Riversleigh, Lawn Hill National 

Park, northwestern Queensland, in shallow subterranean pools] 

Pycnisia raptor Bruce, 1992a: 554; Figs 1-6. [Cave 8MD26, Katherine Region, Northern Territory, 13°55'S 

132°28'E, 33.5 m] 

Sinodina liang & Cai, 1999 

= Sinodina Liang & Cai, 1999 (type species Caridina gregoriana Kemp, 1923, by original designation, 


Sinodina acutipoda (Liang, 1989) 

= Caridina acutipoda Liang, 1989: 282; Figs 1-10. [Lugu Lake, 27°43'N 100°46'E] 

Sinodina angulata Liang, 2002a: 118; Fig. 1. [Longtan stream, Jiangchuan County, Yunnan Province, China] 

Sinodina banna (Cai & Dai, 1999) 

= Caridina banna Cai & Dai, 1999: 214; Figs 3-4. [Runjinghong, Hinghong County] 

= Sinodina bannica Liang, 2004: iv. [Erroneous spelling] 

Sinodina bispinosa (Liang, 1990) 

= Caridina bispinosa Liang, 1990: 221; Fig. 3. [Yunnan Province, Chuxiong (Jiulongdian)] 

Sinodina dianica Liang & Cai, 1999: 583; Figs 3-4. [Malipo village, Wenshan County, south Yunnan] 

Sinodina gregoriana (Kemp, 1923) 

= Caridina gregoriana Kemp, 1923: 437; Figs 1-2. [Tali-fu (Erhai Lake), near Dali city, Yunnan; lectotype 

designation by Liang & Cai, 1999] 

Sinodina heterodactyla (Liang & Yan, 1985a) 

= Caridina heterodactyla Liang & Yan, 1985a: 198; Fig. 2. [rivulet, Shizong, Yunnan Province] 

Sinodina leptopropoda Liang, 1990 

= Caridina leptopropoda Liang, 1990 (sensu Yü, 1938 (partim)): 220; Fig. 2. [several localities in Yunnan] 

Sinodina lijiang Liang & Cai, 1999: 587; Fig. 7. [Lijiang County, Yunnan Province] 

Sinodina wangtai Liang & Cai, 1999: 585; Figs 5-6. [Wangtai village, Lijing County] 

Sinodina yongshengica Chen & Liang, 2002: 240; Fig. 1. [Taoyuan, Yongsheng County, Yunnan Province, 

26°75'N 100°70'E, 1100 m]


Sinodina yui (Liang & Yan, 1985a) 

= Caridina yui Liang & Yan, 1985a: 200; Fig. 4. [Shuanglongyin, Qiubei County, Yunnan Province] 

Stygiocaris Holthuis, 1960 

= Stygiocaris Holthuis, 1960 (type species Stygiocaris lancifera Holthuis, 1960, by original designation, 


Stygiocaris lancifera Holthuis, 1960: 48; Figs 1-2. [Kuddumurra Well, Yardie Creek Station, North-West 

Cape Peninsula, W. Australia] 

Stygiocaris stylifera Holthuis, 1960: 54; Fig. 3. [Kuddumurra Well, Yardie Creek Station, North-West Cape 

Peninsula, W. Australia] 

Syncaris Holmes, 1900 

= Syncaris Holmes, 1900 (type species Miersia pacifica Holmes, 1895, by original designation, gender 

feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Syncaris pacifica (Holmes, 1895) 

= Miersia pacifica Holmes, 1895: 577; Plate 21, figs 27-28. [Sonoma County] 

Syncaris pasadenae (Kingsley, 1897) 

= Caradina pasadenæ Kingsley, 1897: 98; Plate 3, figs 1-7. [streams about Pasadena, California] 

= Syncaris Trewi Holmes, 1900: 213; Plate 4, fig. 63. [small stream near San Gabriel, Los Angeles 

County, California] 

Troglocaris Dormitzer, 1853 

= Troglocaris Dormitzer, 1853 (type species Troglocaris Schmidtii Dormitzer, 1853 (a junior subjective 

synonym of Palaemon anophthalmus Kollar, 1848), by monotypy, gender feminine; name placed on 

the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

= Troglocaridella Babić, 1922 (type species Troglocaridella hercegovinensis Babić, 1922, by monotypy, 


= Xiphocaridinella Sadowsky, 1930 (type species Xiphocaridinella kutaissiana Sadowsky, 1930, by 


= Spelaeocaris Matjašič, 1956 (type species Spelaeocaris pretneri Matjašič, 1956, by monotypy, gender 

feminine) 

Troglocaris (Troglocaridella) hercegovinensis (Babić, 1922) 

= Troglocaridella hercegovinensis Babić, 1922: 303; Fig. 5. [Höhle Vjeternica bei Zavala (Hercegovina)] 

= Troglocaris hercegovinensis Absolon, 1916: 609. [nomen nudum] 

Troglocaris (Troglocaris) anophthalmus anophthalmus (Kollar, 1848) 

= Palaemon anophthalmus Kollar, 1848: 137. [aus den unterirdischen Gewässern von Krain] 

= Troglocaris Schmidtii Dormitzer, 1853: 87. [Grotte von Kumpole in Unterkrain] 

= Troglocaris Schmidti Schmidti forma typica Fage, 1937: 226. [Type locality not indicated] 

Troglocaris (Troglocaris) anophthalmus intermedia Babić, 1922 

= Troglocaris schmidti var. intermedia Babić, 1922: 300; Figs 1-4. [Mikašinović’s Höhle (in der Nähe von 

Gornje Dubrave), ungefähr 6 km von Ogulin] 

Troglocaris (Troglocaris) bosnica Sket & Zakšek, 2009: 799; Figs 7-8. [Bosnia and Herzegovina, Bosanka 

Krajina, Lušci polje, Suvaja cave] 

Troglocaris (Troglocaris) planinensis Birstein, 1948 

= Troglocaris schmidti planinensis Birstein, 1948: 5; Fig. 2. [Planina Cave, 9 km from Postoina Cave, 

Rieka River, Yugoslavia] 

Troglocaris (Spelaeocaris) kapelana Sket & Zakšek, 2009: 806; Fig. 11Ska. [Croatia, Jezerane, Obajdini, Obajdinova 

pećina cave] 

Troglocaris (Spelaeocaris) neglecta Sket & Zakšek, 2009: 808; Figs 11Sne, 12. [Croatia, Novigrad, Karišnica 

cave near Karin] 

Troglocaris (Spelaeocaris) prasence Sket & Zakšek, 2009: 804; Figs 9-10, 11Spra. [Bosnia and Herzegovina, 

Fatničko polje, Obod cave]


Troglocaris (Spelaeocaris) pretneri (Matjašič, 1956) 

= Spelaeocaris pretneri Matjašič, 1956: 66; Figs 1-2. [einer Höhle aus der südöstlichen Herzegowina (in 

der Nähe von Bileće)] 

Troglocaris (Xiphocaridinella) ablaskiri Birstein, 1939 

= Troglocaris schmidti ablaskiri Birstein, 1939: 961; Figs 1-2. [Abkhasia, cave near Achkshe-Tyz-Gua] 

Troglocaris (Xiphocaridinella) fagei Birstein, 1939 

= Troglocaris schmidti fagei Birstein, 1939: 964; Figs 3-5. [Abkhasia, cave near Psyrtzkha] 

Troglocaris (Xiphocaridinella) jusbaschjani Birstein, 1948 

= Troglocaris schmidti jusbaschjani Birstein, 1948: 4; Fig. 1. [Russian Federation, Krasnodarskiy Krai, 

Sochi area, cave at Matzesta] 

Troglocaris (Xiphocaridinella) kutaissiana (Sadowsky, 1930) 

= Xiphocaridinella kutaissiana Sadowsky, 1930: 95. [unterirdische Gewässer in Karsthöhlen bei Kutais 

(West-Georgien)] 

Troglocaris (Xiphocaridinella) osterloffi Juzbaš’jan , 1940: 73. [Shakuran] 

Typhlatya Creaser, 1936 

= Typhlatya Creaser, 1936 (type species Typhlatya pearsei Creaser, 1936, by monotypy, gender feminine) 

Typhlatya arfeae Jaume & Bréhier, 2005: 390; Figs 3-18. [Résurgence de Font Estramar, Salses-le-château, 

Pyrénées-orientales, France] 

Typhlatya campecheae H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 6; Figs 3-4. [Grutas de Xtacumbilxunam, 

Bolonchenticul, Campeche, Mexico] 

Typhlatya consobrina Botoşăneanu & Holthuis, 1970: 123; Figs 1-2. [Cueva del Agua, Sierra de Cubitas, at 

the foot of Cerro Tuabaquei, very near to ”Finca la Entrada” (NE from the town Camagüey, prov. 

Camagüey, in a subterranean lake] 

Typhlatya dzilamensis Alvarez, Iliffe & Villalobos, 2005: 83; Figs 2-3. [Cenote Buya Uno, Dzilam de Bravo, 

Yucatan, Mexico] 

Typhlatya elenae Juarrero, 1994: 58; Figs 1-10. [Cueva Perico Sanchez, 5 km al N de Jaguey Grande, Provincia 

de Matanzas] 

Typhlatya galapagensis Monod & Cals, 1970: 70; Figs 1-46, 50, 53, 55, 58, 60, 63-64. [Ile de Santa Cruz, 

versant S-E, eau courante à peine sodique, dans une crevasse profonde de 15-20 m, à 50 m d’altitude 

et à 2 km de la côte] 

Typhlatya garciadebrasi Juarrero de Varona & Ortiz, 2000: 46; Figs 1-3. [Cueva Chicarrones, Bolondrón, 

Procincia de Matanzas] 

Typhlatya garciai Chace, 1942a: 99; Plate 29. [Potrero del Molino Cave, Las Cuatrocientas, Banes, Oriente 

Province, Cuba] 

Typhlatya iliffei C.W.J. Hart & Manning, 1981: 447; Figs 29-55. [Tucker’s Town cave, Bermuda] 

Typhlatya kakuki Alvarez, Iliffe & Villalobos, 2005: 86; Figs 4-5. [Shrimp Hole, Salinas Point, Acklins Island, 

Bahamas] 

Typhlatya miravetensis Sanz & Platvoet, 1995: 80; Figs 1-4. [Cave ”Ullal de la Rambla de Miravet” between 

the towns of Cabanes and Orpesa, province of Castellón, eastern Spain] 

Typhlatya mitchelli H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 2; Figs 1-2. [Cenote Kabahchen, Maní, Yucatán, 

Mexico] 

Typhlatya monae Chace, 1954: 318; Fig. 1. [Mona Island, Puerto Rico, from well 30ft deep at ”El Molino”, 

about 1 mile southeast of NYA camp at Sardinera] 

Typhlatya pearsei Creaser, 1936: 128; Figs 31-41. [Balam Canche Cave, 4.8 km east 0.8 km south of Chichen 

Itza, Yucatan] 

Typhlatya rogersi Chace & Manning, 1972: 14; Figs 10-11. [Marl Pool, back of Shelly Beach, Ascension Island] 

Typhlatya taina Estrada & Gómez, 1987: 3; Figs 2-5. [Cueva de la India, 2 km al E de Puerto Escondido, 

Santa Cruz del Norte, La Habana, Cuba] 

Typhlatya utilaensis Alvarez, Iliffe & Villalobos, 2005: 89; Figs 6-7. [George Gaberel’s Cave, Utila, Bay Islands, 

Honduras]


Typhlocaridina liang & yan, 1981 

= Typhlocaridina Liang & Yan, 1981 (type species Typhlocaridina lanceifrons Liang & Yan, 1981, by 

original designation and monotypy, gender feminine) 

Typhlocaridina lanceifrons Liang & Yan, 1981: 32; Figs 1-19. [Daji Cave, Wuming County, Guangxi] 

Typhlocaridina liui Liang & Zhou, 1993: 232; Figs 1(1-3). [Paobingyan Cave, Lingui County, Guangxi 

Province] 

Typhlocaridina semityphlata Cai, 1995b: 157; Fig. 1. [cave in Guilin, Guangxi Autonomous Region, South 

China] 

Typhlopatsa Holthuis, 1956a 

= Typhlopatsa Holthuis, 1956a (type species Typhlopatsa pauliani Holthuis, 1956a, by original designation 


Typhlopatsa pauliani Holthuis, 1956a: 53. [Mitoho Cave, N.-E. corner of Tsimanampetsotsa Lake, Mahafaly 

Province, S.-W. Madagascar] 

Superfamily BReSIlIoIDeA Calman, 1896 

Family AGoSToCARIDIDAe C.W.J. Hart & Manning, 1986 

Agostocaris C.W.J. Hart & Manning, 1986 

= Agostocaris C.W.J. Hart & Manning, 1986 (type species Agostocaris williamsi C.W.J. Hart & Manning, 


Agostocaris acklinsensis Alvarez, Villalobos & Iliffe, 2004: 370; Figs 2-4. [Jumby Hole Cave, Snug Corner, 

Acklins Island, Bahamas] 

Agostocaris bozanici Kensley, 1988: 688; Figs 1-3 [Xcan-ha Cenote (Cenote Roja), Cozumel Island, Quintana 

Roo, Mexico, 80-100 feet] 

Agostocaris williamsi C.W.J. Hart & Manning, 1986: 412; Figs 27-47. [Bahamas, 24°27'N, 75°30'W, Big 

Fountain Blue Hole] (Fig. 16) 

Fig. 16. Agostocaris williamsi C.W.J. Hart & Manning, 1986. Photo by Thomas M. Iliffe.


Family ALVINOCARIDIDAE Christoffersen, 1986 

Alvinocaridinides Komai & Chan, 2010a 

= Alvinocaridinides Komai & Chan, 2010a (type species Alvinocaridinides formosa Komai & Chan, 

2010a, by original designation and monotypy, gender feminine) 

Alvinocaridinides formosa Komai & Chan, 2010a: 26; Figs 1C, 7-10. [Gueishandao, Yilan County, Taiwan, 

24°51.231'N 121°59.204'E, 252-275 m] 

Alvinocaris Williams & Chace, 1982 

= Alvinocaris Williams & Chace, 1982 (type species Alvinocaris lusca Williams & Chace, 1982, by 


Alvinocaris alexander Ahyong, 2009: 777; Figs 1-3. [Rumble V Seamount, 36°08.27-07.96'S 178°11.74- 

11.70'E, 485-415 m] 

Alvinocaris brevitelsonis Kikuchi & Hashimoto, 2000: 136; Figs 1-3. [”Depression C” of the Minami-Ensei 

Knoll, 28°23.35'N 127°38.38'E, 705 m] 

Alvinocaris chelys Komai & Chan, 2010a: 16; Figs 1A-B, 2-6. [Gueishandao, Yilan County, Taiwan, 

24°49.682'N 122°0.254'E, 300-276 m] (Fig. 17) 

Alvinocaris dissimilis Komai & Segonzac, 2005a: 1158; Figs 25-26, 29. [Depression C, Minami-Ensei Knoll, 

28°23.35'N 127°38.38'E, 705 m] 

Alvinocaris komaii Zelnio & Hourdez, 2009: 55; Figs 1-6. [Kilo Moana, 20°9'S 76°12'E, 2620 m, Eastern Lau 

Spreading Center, Lau Basin, southwest Pacific] 

Alvinocaris longirostris Kikuchi & Ohta, 1995: 772-778, Figs 1-7. [Iheya Ridge, Clam Site, Okinawa Trough, 

27°32.70'N 126°58.20'E, 1360 m] 

Alvinocaris lusca Williams & Chace, 1982: 137; Figs 1-7. [Galapagos Rift Rose Garden area, 0°48.25'N 

86°13.48'W, maximum of 2450 m] 

Alvinocaris markensis Williams, 1988: 264; Figs 1-2, 7l. [Mid-Atlantic Rift Valley about 70 km south of 

Kane Fracture Zone, 23°22.09'N 44°57.12'W, 3437 m] 

Fig. 17. Alvinocaris chelys Komai & Chan, 2010. Photo by Tin-Yam Chan.


Alvinocaris methanophila Komai, Shank & Van Dover, 2005: 29; Figs 1-4. [ODP site 996, Blake Ridge Diapir, 

32°29.623'N 76°11.467'W, 2155 m] 

Alvinocaris muricola Williams, 1988: 268; Figs 3-4, 7e-k. [West Florida Escarpment, 26°01'N 84°54.61'W, 

3277 m] 

Alvinocaris niwa Webber, 2004: 5; Figs 1-4. [Rumble V, 36°8.63-8.57'S 178°11.77-11.50'E, 877-655 m] 

Alvinocaris stactophila Williams, 1988: 272; Figs 5-6, 7a-d. [north central Gulf of Mexico about 129 km S of 

Louisiana, 27°46.94'N 91°30.34'W, 534 m] 

Alvinocaris williamsi Shank & Martin, 2003: 159; Figs 1-3. [Menez Gwen hydrothermal field, North Atlantic 

Ocean, 37°50.5'N 31°31.3'W, 850 m] 

Chorocaris Martin & Hessler, 1990 

= Chorocaris Martin & Hessler, 1990 (type species Chorocaris vandoverae Martin & Hessler, 1990, by 


Chorocaris chacei (Williams & Rona, 1986) 

= Rimicaris chacei Williams & Rona, 1986: 455; Figs 8-10. [TAG Hydrothermal Field, Mid-Atlantic 

Ridge, 26°08.3'N 44°49.6'W, 3620-3650 m] 

Chorocaris paulexa Martin & Shank, 2005: 186; Figs 1-8. [Homer Vent (347OC black smoker), 17°37.220'S 

113°15.123'W, 2595 m, southern East Pacific Rise] 

Chorocaris vandoverae Martin & Hessler, 1990: 2; Figs 1-2, 3a-e. [Alice springs vent field, Mariana Back- 

Arc Basin, 18°12.599'N 144°42.431'E, 3640 m] 

Mirocaris Vereshchaka, 1997b 

= Mirocaris Vereshchaka, 1997b (type species Mirocaris keldyshi Vereshchaka, 1997b (junior subjective 

synonym of Chorocaris fortunata Martin & Christiansen, 1995), by monotypy, gender feminine) 

Mirocaris fortunata (Martin & Christiansen, 1995) 

= Chorocaris fortunata Martin & Christiansen, 1995: 221; Figs 1-3. [Vent site 3, Lucky Strike hydrothermal 

vent, Azores, 37°17.6'N 32°16.5'W, 1624 m] 

= Mirocaris keldyshi Vereshchaka, 1997b: 431; Figs 1-5. [TAG location, 26°09'N 44°50'W, 3650 m] 

Mirocaris indica Komai, Martin, Zala, Tsuchida & Hashimoto, 2006: 110; Figs 1-5. [Central Indian Ridge, 

Kairei Field, 25°19.2'S 70°02.4'E, 2422 m] 

Nautilocaris Komai & Segonzac, 2004 

= Nautilocaris Komai & Segonzac, 2004 (type species Nautilocaris saintlaurentae Komai & Segonzac, 


Nautilocaris saintlaurentae Komai & Segonzac, 2004: 1179-1188; Figs 2-6. [North Fiji Basin, White Lady 

site, 16°59.50'S 173°55.47'E, 2000 m] 

Opaepele Williams & Dobbs, 1995 

= Opaepele Williams & Dobbs, 1995 (type species Opaepele loihi Williams & Dobbs, 1995, by monotypy, 


Opaepele loihi Williams & Dobbs, 1995: 229; Figs 1-3. [Loihi Seamount, Hawaii, 18°55'N 155°16'W, 980 m] 

Opaepele susannae Komai, Giere & Segonzac, 2007: 240; Figs 2-7. [Lilliput, southern Mid-Atlantic Ridge, 

09°32.845'S 13°12.546'W, 1500 m, mussel field with diffuse vent fluids] 

Opaepele vavilovi Lunina & Vereshchaka, 2010: 70; Figs 1-2. [Mid-Atlantic Ridge, Broken Spur vent site, 

stn 4797] 

Rimicaris Williams & Rona, 1986 

= Rimicaris Williams & Rona, 1986 (type species Rimicaris exoculata Williams & Rona, 1986, by original 


= Iorania Vereshchaka, 1996 (type species Iorania concordia Vereshchaka, 1996 (junior subjective synonym 

of Rimicaris exoculata Williams & Rona, 1986), by monotypy, gender feminine)


Rimicaris exoculata Williams & Rona, 1986: 448; Figs 5-7. [TAG Hydrothermal Field, Mid-Atlantic Ridge, 

26°08.3'N 44°49.6'W, 3620-3650 m] 

= Iorania concordia Vereshchaka, 1996: 954; Figs 1-5. [TAG location, 26°09N 44°50W, point D, 3650 m] 

= Rimicaris aurantiaca Martin, Signorovitch & Patel, 1997: 400; Figs 1-5. [Snake Pit hydrothermal vent 

field, Mid-Atlantic Ridge, 23°22.1'N 44°57.0'W] 

Rimicaris kairei Watabe & Hashimoto, 2002: 1168; Figs 1-4. [The Central Indian ridge, Indian Ocean, the 

Kairei Field, 25°19.16'S 70°02.40'E, 2454 m] 

Shinkaicaris Komai & Segonzac, 2005a 

= Shinkaicaris Komai & Segonzac, 2005a (type species Alvinocaris leurokolos Kikuchi & Hashimoto, 


Shinkaicaris leurokolos (Kikuchi & Hashimoto, 2000) 

= Alvinocaris leurokolos Kikuchi & Hashimoto, 2000: 141; Figs 4-7. [”Depression C” of the Minami- 

Ensei Knoll, 28°23.35'N 127°38.38'E, 705 m] 

Family BReSIlIIDAe Calman, 1896 

Bresilia Calman, 1896 

= Bresilia Calman, 1896 (type species Bresilia atlantica Calman, 1896, by monotypy, gender feminine; 


Bresilia antipodarum Bruce, 1990a: 855; Figs 6-10. [Tasman Sea, off Cape Freycinet, Tasmania, 42°2.20'S 

148°38.70'E, 800 m] 

Bresilia atlantica Calman, 1896: 7; Plate 1; Plate 2, figs 15-18. [south west of Ireland, 750 fms] 

Bresilia briankensleyi Bruce, 2005a: 176; Figs 1-3. [Egyptian Red Sea, 22°85.2'N 36°45.9'E to 22°34'N 

36°46.2'E, 750-753 m] 

Bresilia corsicana Forest & Cals, 1977: 551; Figs 1-20. [canal de Corse, 545 milles dans le 285 du cap Trattoja, 

sud-ouest de l’île Capraia, 450 m] 

Bresilia gibbosa Komai & Yamada, 2010: 42; Figs 1-5. [Apo-gama Cave, Onna Village, Okinawa Island, 

Ryukyu Islands, 30 m] 

Bresilia plumifera Bruce, 1990b: 1; Figs 1-4. [Taupo Sea Mount, Western Tasman Sea, 33°14.21'S 56°10.68'E, 

133 m] 

Bresilia rufioculus Komai & Yamada, 2011: 72; Figs 1-6. [Ohoba Cave, Ie Island, Okinawa Islands] (Fig. 18) 

Fig. 18. Bresilia rufioculus Komai & Yamada, 2011. Photo by Tomoyuki Komai.


Bresilia saldanhai Calado, Chevaldonné & dos Santos, 2004: 191; Figs 1-4. [underwater cave at Ponta garajau, 

south of Madeira island, eastern Atlantic, 32.63°N 16.85°W, 15 m] 

Encantada Wicksten, 1989a 

= Encantada Wicksten, 1989a (type species Encantada spinoculata Wicksten, 1989a, by monotypy, gender 

feminine) 

Encantada spinoculata Wicksten, 1989a: 667; Figs 1-2. [Beagle Island, east of Jervis Island, Galapagos, 

0°26'S 90°40'W, 55-92 m] 

Family DISCIADIDAe Rathbun, 1902c 

Discias Rathbun, 1902c (Fig. 19) 

= Anisocaris Ortmann, 1893 (type species Anisocaris dromedarius Ortmann, 1893 (probably an invalid 

senior subjective synonym of Discias atlanticus Gurney, 1939b), by monotypy, gender feminine; 

name suppressed for the purposes of the Principle of Priority, but not for those of the Principle of 

Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology in 


= Discias Rathbun, 1902c (type species Discias serrifer Rathbun, 1902c, by monotypy, gender masculine) 

Discias atlanticus Gurney, 1939b: 388; Figs 1-13. [in plankton at night in the shallow waters of the Reach 

close to the Bermuda Biological Station] 

? = Anisocaris dromedarius Ortmann, 1893: 74; Plate 4, figs 3-3z. [Plankton Expedition, Nördl. 

Aequa torialstrom, JN 148 (10.2°N 22.2°W, 0-400 m); Südl. Aequatorialstrom, JN 218 (3.8°S 32.6°W, 

0-400 m)] 

Discias brownae Kensley, 1983: 8; Figs 6-9. [Green Point, Port Jackson, New South Wales, 33°50'S 151°19'E, 

9.8 m] 

Fig. 19. Discias spec. Photo by Leslie Harris.


Discias exul Kemp, 1920: 138; Figs 1-3. [Port Blair, Andaman Islands, on the reef at the N end of Ross Island, 

low water] 

= Discias mvitae Bruce, 1976b: 119; Figs 1-5. [Fort Jesus, Mombasa, 4°04.0'S 39°42.18'E, 1 m] 

Discias musicus Holthuis, 1981: 787; Figs 1-2. [Lagoon west of Saipan, Marianas Islands, immediately 

shoreward of barrier reef flat] 

= Discias musicus Holthuis, 1953b: 52. [nomen nudum] 

Discias pascuensis Fransen, 1987: 501; Figs 1-3. [Tahai, west coast of Easter Island, Chile, 39 m] 

Discias serratirostris Lebour, 1949a: 1107; Figs 1-2. [about three miles off Castle Roads, Bermuda, near 

bottom, at about 100ft or more] 

Discias serrifer Rathbun, 1902c: 290; Figs 1-4. [Tagus Cove, Albemarle Island, on the reef north of Tagus 

Hill] 

Discias vernbergi Boothe & Heard, 1987: 506; Figs 1-3. [65 nautical miles W of Egmont Key, Florida, 

27°37'N 83°58'W, 55 m] 

Kirnasia Burukovsky, 1988 

= Kirnasia Burukovsky, 1988 (type species Kirnasia nesisi Burukovsky, 1988, by original designation, 


Kirnasia nesisi Burukovsky, 1988: 457; Fig. 1. [42°39'N, 28°45'W, 850-1200 m] 

Kirnasia siedlecki Burukovsky, 1988: 458; Fig. 2. [36°18'N, 19°50'W, 874 m] 

Lucaya Chace, 1939 

= Lucaya Chace, 1939 (type species Lucaya bigelowi Chace, 1939, by monotypy, gender feminine) 

Lucaya bigelowi Chace, 1939: 34. [east of Great Abaco Island, Bahamas, 26°12'N, 76°26'W, 2610 fms] 

Tridiscias Kensley, 1983 

= Tridiscias Kensley, 1983 (type species Tridiscias transkei Kensley, 1983, by monotypy, gender masculine) 

Tridiscias transkei Kensley, 1983: 18; Figs 15-17. [off Transkei, South Africa, 31°59'S 29°22.5'E, 150-200 m] 

Family PSeuDoCHelIDAe De Grave & Moosa, 2004 

Pseudocheles Chace & Brown, 1978 

= Pseudocheles Chace & Brown, 1978 (type species Pseudocheles enigma Chace & Brown, 1978, by 


Pseudocheles chacei Kensley, 1983: 22; Figs 18-22. [Looe Key, Florida, 6 m] 

Pseudocheles enigma Chace & Brown, 1978: 757; Figs 1-4. [west side of lagoon, Lizard Island, Queensland, 

Australia, 1-15 m] 

Pseudocheles neutra De Grave & Moosa, 2004: 1; Figs 1-4. [Kaledupa Reef, Kaledupa, Tukangbesi Archipelago, 

south-eastern Sulawesi, Indonesia, 10 m] 

Superfamily NeMAToCARCINoIDeA Smith, 1884 

Family euGoNAToNoTIDAe Chace, 1937a 

Eugonatonotus Schmitt, 1926a 

= Gonatonotus A. Milne-Edwards, 1881b (type species Gonatonotus crassus A. Milne-Edwards, 1881b, 

by monotypy, gender masculine; invalid junior synonym of Gonatonotus Adams & White, 1847 

(Crustacea Brachyura); name placed on the Official Index of Rejected and Invalid Generic Names in 


= Eugonatonotus Schmitt, 1926a (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, type species 

therefore Gonatonotus crassus A. Milne-Edwards, 1881b, gender masculine; name placed on the 

Official List of Generic Names in Zoology in Opinion 470 in 1957)


Fig. 20. Eugonatonotus crassus (A. Milne-Edwards, 1881). Photo Pillsbury Expedition sta 936b. 

= Gomphonotus Chace, 1936 (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, gender masculine; 

name placed on the Official Index of Rejected and Invalid Generic Names in Zoology in 


= Galatheacaris Vereshchaka, 1997a (type species Galatheacaris abyssalis Vereshchaka, 1997a, by original 


Eugonatonotus crassus (A. Milne-Edwards, 1881b) (Fig. 20) 

= Gonatonotus crassus A. Milne-Edwards, 1881b: 10. [Grenade par 262 brasses de profondeur] 

= Gonatonotus crassus var. longirostris A. Milne-Edwards, 1883: Planche 34. [Off Sand Key] 

Eugonatonotus chacei Chan & Yu, 1991a: 144; Fig. 1; Plate 1A. [Ta-Shi, I-Lan County, Taiwan] 

= Galatheacaris abyssalis Vereshchaka, 1997a: 365; Figs 1-9. [Celebes Sea, 01°50'N 119°20'E, 4940-4970 m] 

Family NeMAToCARCINIDAe Smith, 1884 

Lenzicarcinus Burukovsky, 2005 

= Lenzicarcinus Burukovsky, 2005 (type species Lenzicarcinus struncki Burukovsky, 2005, by original 


Lenzicarcinus struncki Burukovsky, 2005: 572; Figs 1-2. [Atlantic Ocean, south of Ascension Island, 12°11'S 

6°W, 2000 m] 

Lipkius yaldwyn, 1960 

= Lipkius Yaldwyn, 1960 (type species Lipkius holthuisi Yaldwyn, 1960, by original designation and 


Lipkius holthuisi Yaldwyn, 1960: 16; Fig. 1. [41°39'30''S 175°17'E, between 50 and 200 fms] 

Nematocarcinus A. Milne-edwards, 1881b (Fig. 21) 

= Nematocarcinus A. Milne-Edwards, 1881b (type species Nematocarcinus cursor A. Milne-Edwards, 

1881b, by monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology 

in Opinion 470 in 1957) 

= Eumiersia Smith, 1882 (type species Eumiersia ensifera Smith, 1882, by monotypy, gender feminine) 

= Stochasmus Spence Bate, 1888 (type species Stochasmus exilis Spence Bate, 1888, by monotypy, 


Nematocarcinus africanus Crosnier & Forest, 1973: 101; Figs 29a, 30a-e, 31a-b. [9°05'N 15°10'W, 310-380 m]


Fig. 21. Nematocarcinus spec. Photo by Tin-Yam Chan. 

Nematocarcinus agassizii Faxon, 1893: 204. [Albatross stn 3358 (06°30'N 081°44'W, 555 fms); lectotype designation 

by Burukovsky, 2001] 

Nematocarcinus batei Burukovsky, 2000a: 284; Figs 1b-c. [Challenger stn 237, 34°37'N 140°32'E, near Yogama, 

Japan, 3470 m] 

Nematocarcinus bituberculatus Chace, 1986: 69; Fig. 37. [west of Halmahera, Indonesia, 0°12'15"N 

127°29'30"E, 527 m] 

Nematocarcinus chacei Burukovsky, 2002a: 5; Fig. 1. [Indian Ocean, Gulf of Aden, 11°31'40''N 42°46'40''E, 

335 m] 

Nematocarcinus challengeri Burukovsky, 2006a: 896; Fig. 1. [Challenger stn 250, Pacific Ocean, 37°49'N 

166°47'W, 5477 m] 

Nematocarcinus combensis Burukovsky, 2000b: 1038; Fig. 2. [south-west part of Indian Ocean, Comb Bank, 

12°15'S 177°28'W, 800-810 m] 

Nematocarcinus crosnieri Burukovsky, 2000c: 666; Fig. 3. [New Caledonia, 23°19'S 167°59'E 850-920 m] 

Nematocarcinus cursor A. Milne-Edwards, 1881b: 14. [la Mer des Antilles, 500 brasses] 

Nematocarcinus ensifer (Smith, 1882) 

= Eumiersia ensifera Smith, 1882: 77; Plate 13, Figs 1-9. [Blake stns 305, 41°33'15''N 65°51'25''W, 810 


73°10'30''W, 1186 fms; 340, 39°25'30''N 70°58'40''W, 1394 fms; 341, 39°38'20''N 70°56'0''W, 1241 

fms] 

Nematocarcinus evansi Burukovsky, 2000d: 1291; Fig. 2. [south-west Australia, 20°16.3'S 113°13.5'E, 913- 

916 m] 

Nematocarcinus exilis (Spence Bate, 1888) 

= Stochasmus exilis Spence Bate, 1888: 823; Plate 132, fig. 14. [Challenger stn 84, 30°38'N 18°5'W, off the 

Canary Islands] 

Nematocarcinus faxoni Burukovsky, 2001: 1429; Figs 1-2. [Eastern Pacific, 16°33'N 99°52'W, 1208 m]


Nematocarcinus gladius Macpherson, 1984: 67; Figs 13-14, 15a-b. [34°37'S 17°03'E, 1580-1620 fms; exact 

locality not indicated in publication, information from IZIKO collection label] 

Nematocarcinus gracilipes Filhol, 1884: 232; Fig. 1. [pêché à 850 mètres de profondeur; no further indication 

of type locality] 

Nematocarcinus gracilis Spence Bate, 1888: 815; Plate 132, fig. 8. [Challenger stn 174c, 19°7'50"S 178°19'35"E, 

off Kandavu, Fiji Islands, 610 fms; lectotype designation by Burukovsky, 2007a] 

Nematocarcinus hanamuri Burukovsky, 2000d: 1290; Fig. 1. [south-west Australia, 20°16.3'S, 113°13.5'E, 

913-916 m] 

Nematocarcinus hiatus Spence Bate, 1888: 821; Plate 132, fig. 12. [Challenger stn 169, 37°34'S 179°22'E, off 

New Zealand, 700 fms] 

Nematocarcinus kaiensis Burukovsky, 2000b: 1040; Fig. 3. [Indonesia, Selan-Timur Archipelago, 05°15'S 

133°59'E, 769-800 m] 

Nematocarcinus lanceopes Spence Bate, 1888: 804; Plate 131. [Challenger stn 152, 60°52'S 80°20'E, Antarctic 

Sea, 1260 fms] 

= Acanthephyra antarctica Bage, 1938: 6; Plate 4, fig. 1. [63°13'S 101°42'E, 870 fms] 

Nematocarcinus longirostris Spence Bate, 1888: 806; Plate 132, fig. 2. [Challenger stn 237, 34°37'N 140°32'E, 

near Yokohama, Japan, 1875 fms] 

Nematocarcinus machaerophorus Burukovsky, 2003: 116; Fig. 33. [Pacific Ocean, Eiao Island, 07°52'S 

140°39'W, 1000 m] 

= Nematocarcinus machaerophorus Burukovsky, 2004: 1181; Figs a-e. [Pacific Ocean, Eiao Island, 

07°52'S 140°39'W, 1000 m] 

Nematocarcinus manningi Burukovsky, 2003: 118; Fig. 34. [North Pacific Ocean, 35°06'45''N 139°19'45''E, 

1138 m] 

Nematocarcinus novaezealandicus Burukovsky, 2006b: 441; Fig. 1. [Chatham Rise, 42°51.7'-42°49.5'S 

175°23.3'-175°46.1'W, 1057 m] 

Nematocarcinus nudirostris Burukovsky, 1991a: 41; Fig. 1(5-8). [9°33'5"-9°35'S 59°52'7"-59°58'8"E, 830-430 m] 

= Nematocarcinus rectirostris Burukovsky, 1991a: 39; Fig. 1(1-4). [25°28'-25°29'S 35°14'-35°11'E, 715- 

700 m] 

Nematocarcinus ovalis Komai & Segonzac, 2005b: 355; Figs 8-10. [East Pacific Rise, 12°45.50'N 103°55.80'W, 

2558-2619 m (c. 120 m above bottom)] 

Nematocarcinus paucidentatus Spence Bate, 1888: 816; Plate 132, fig. 9. [Challenger stn 174c, off Kandavu, 

Fiji Islands, 19°7'50''S 178°19'35'', 610 fms] 

= Nematocarcinus alisae Burukovsky, 2000e: 903; Fig. 4. [New Caledonia, 21°05'S 165°50'E, 960-1100 m] 

Nematocarcinus parvus Burukovsky, 2000f: 1163; Fig. 7. [Madagascar, 12°26'-12°27'S 42°08'5''-48°13'E, 600- 

705 m] 

Nematocarcinus poupini Burukovsky, 2007b: 1; Fig. 1. [Austral Islands, East of Rapa Island, 27°38.2'S 

144°14.4'W, 840-1200 m] 

Nematocarcinus productus Spence Bate, 1888: 810; Plate 132, fig. 5. [Challenger stns 205, 16°42'N 119°22'E, 

off Luzon, Philippine Islands, 1050 fms; 195, 4°21'S 129°7'E, off Banda Island, 1425 fms; 237, 34°37'N 

140°32'E, near Yokohama, Japan, 1875 fms; 176, 18°30'S 173°52'E, off the New Hebrides, 1450 fms] 

= Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fig. 6. [Challenger stn 218, 2°33'S 

144°4'E, off New Guinea, 1070 fms ] 

= Nematocarcinus intermedius Spence Bate, 1888: 821; Plate 132, fig. 13. [Challenger stn 218, 2°33'S 

144°4'E, off New Guinea, 1070 fms] 

Nematocarcinus proximatus Spence Bate, 1888: 808; Plate 132, fig. 3. [Challenger stn 146, 33°42'S 78°18'W, 

west of Valparaiso, 1375 fms; lectotype designation by Burukovsky, 2003] 

Nematocarcinus pseudocursor Burukovsky, 1990: 194; Fig. 2B. [25°07'S 99°26'W, 780-790 m] 

Nematocarcinus pseudogracilis Burukovsky, 2007a: 807; Figs 5-7. [Pacific Ocean, Austral Islands, Tubuai, 

23°12.3'S 149°34.4'W, 750-1000 m] 

Nematocarcinus richeri Burukovsky, 2000f: 1155; Figs 1-3. [Indonesia, Kai Islands, 05°15'S 133°01'E, 605- 

576 m] 

Nematocarcinus romenskyi Burukovsky, 2000a: 288; Fig. 5. [Atlantic Ocean, Discovery Bank, 42°3'8''S 

00°03'W, 1030-1050 m]


Nematocarcinus rotundus Crosnier & Forest, 1973: 103; Figs 29c, 30f-i, 31e-f. [28°43'N 87°14'30''W, 960 m] 

Nematocarcinus safari Burukovsky, 2000c: 662; Fig. 1. [south-west part of Indian Ocean, Madagascar sink, 

30°42'9''S 48°20'6''E, 4297 m] 

Nematocarcinus serratus Spence Bate, 1888: 819; Plate 132, fig. 11. [Challenger stn 169, 37°34'S 179°22'E, off 

New Zealand, 700 fms] 

Nematocarcinus sigmoideus Macpherson, 1984: 63; Figs 10-12 [area del Cabo entre 840 y 2200 m; Banco 

Valdivia, entre 754 y 940 m; costas de Namibia (28°19'S 14°18,6'E)] 

Nematocarcinus subtegulisfactus Burukovsky, 2000c: 664; Fig. 2. [Philippines, 12°05'N 122°14'E, 1404 m] 

Nematocarcinus subtilis Burukovsky, 2000b: 1042; Fig. 4. [Philippines, 11°58'N 121°06'E, 448-466 m] 

Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fig. 6. [Challenger stns 235, 34°07'N 

138°0'E, south of Japan, 565 fms; 218, 2°33'S 144°04'E, near the Admiralty Islands, 1070 fms; 232, 

35°11'N 139°28'E, Japan, 345 fms] 

= Nematocarcinus parvidentatus Spence Bate, 1888: 814; Plate 132, fig. 7. [Challenger stn 237, 34°37'N 

140°32'E, near Yokohama, Japan, 1875 fms] 

= Nematocarcinus serratirostris Burukovsky, 1991a: 41; Fig. 1(9-11). [24°58'-25°05'S 35°40'-35°34'E, 

1010-910 m] 

Nematocarcinus tenuirostris Spence Bate, 1888: 817; Plate 132, fig. 10. [Challenger stns 174c, 19°7'50"S 

178°19'35"E, off Kandavu, Fiji Islands, 610 fms; 214, 4°33'N 127°6'E, south of the Philippine Islands, 

500 fms] 

= Nematocarcinus tenuirostris var. sibogae De Man, 1917: 279. [10°48'.6S 123°23'.1E, 918 m] 

Nematocarcinus tuerkayi Burukovsky, 2005: 573; Fig. 3. [Indian Ocean, Gulf of Aden, 12°52.5'N 45°53.3'E, 

1185 m] 

Nematocarcinus undulatipes Spence Bate, 1888: 801; Plate 130. [Challenger stn 200, 6°47'N 122°28'E, off 

Sibago, Philippine Islands, 250 fms; lectotype designation by Burukovsky, 2002a] 

Nematocarcinus webberi Burukovsky, 2006b: 444; Fig. 2. [Off New Zealand, 42°41.7'S 174°28'E, 1723-1549 m] 

Nematocarcinus yaldwyni Burukovsky, 2006b: 445; Fig. 3. [NE of Ritchie Bane, New Zealand, 39°47.9'S 

178°17.9'E, 832-788 m] 

Nigmatullinus Burukovsky, 1991a 

= Nigmatullinus Burukovsky, 1991a (type species Nematocarcinus acanthitelsonis Pequegnat, 1970, by 

original designation and monotypy, gender masculine) 

Nigmatullinus acanthitelsonis (Pequegnat, 1970) 

= Nematocarcinus acanthitelsonis Pequegnat, 1970: 69; Figs 4-3, 4-4. [Southwestern Gulf of Mexico, 

23°36'N 93°57'W, 2045 fms] 

Segonzackomaius Burukovsky, 2011 

= Segonzackomaius Burukovsky, 2011 (type species Nematocarcinus burukovskyi Komai & Segonzac, 


Nematocarcinus burukovskyi Komai & Segonzac, 2005b: 346; Figs 2-7. [South East Pacific Rise, Garrett Segment, 

Wormwood site, 17°34.91'S 113°14.68'W, 2595 m] 

Nematocarcinus altus Spence Bate, 1888: 809; Plate 132, fig. 4. [Challenger stn 198, 2°55'N 124°53'E, south 

of the Philippine Islands, 2150 fms] 

Family RHyNCHoCINeTIDAe ortmann, 1890 

Cinetorhynchus Holthuis, 1995 

= Rhynchocinetes (Cinetorhynchus) Holthuis, 1995 (type species Rhynchocinetes rigens Gordon, 1936, 

by original designation, gender masculine) 

Cinetorhynchus brucei Okuno, 2009a: 940; Figs 1-3. [Imazuni, Kume-jima Island, Rykyu Islands, Japan, 

26°21.8'N 126°49.6'E, 18 m] 

Cinetorhynchus concolor (Okuno, 1994a) 

= Rhynchocinetes concolor Okuno, 1994a: 66; Figs 1-2, 3A, 4A-D. [26°11.2'N 127°16.8'E, Hizushi-hama, 

Aka-jima Islet, Kerama Group, Ryukyu Islands, 3 m]


Fig. 22. Cinetorhynchus manningi Okuno, 1996. Photo by Arthur Anker. 

Cinetorhynchus erythrostictus Okuno, 1997a: 36; Plate 1A-B; Figs 2A, 3, 4A-C. [Ryukyu Islands, Aka Harbor, 

Aka-jima Islet, Kerama Group, 26°11.2'N 127°17.1'E, 3 m] 

Cinetorhynchus fasciatus Okuno & Tachikawa, 1997: 16; Figs 1-4. [Tenno-ura beach, Chichi-jima Island, 

Ogasawara Islands, Japan, 5 m] 

Cinetorhynchus hawaiiensis Okuno & Hoover, 1998: 33; Figs 1-3, 4A-B, 5A. Hawaiian Islands, 19°39.0'N 

156°00.0'W, Kailua Harbor, Hawai’i, 8 m] 

Cinetorhynchus hendersoni (Kemp, 1925) 

= Rhynchocinetes hendersoni Kemp, 1925: 265; Figs 3-7. [Pamban and Kilakarai, Gulf of Manaar] 

= Rhynchocinetes intermedius Edmondson, 1952: 72; Fig. 3. [of the southwest coast of Oahu, at about 

16 fms] 

= Rhynchocinetes marshallensis Edmondson, 1952: 75; Figs 4-6. [Ebenina, Eniwetok, Marshall Islands] 

Cinetorhynchus hiatti (Holthuis & Hayashi, 1967) 

= Rhynchocinetes hiatti Holthuis & Hayashi, 1967: 162; Figs 1-2. [Formosa, Kosho Bay, southern extremity 

of Formosa, 3 or 4 m] 

= Rhynchocinetes hiatti Holthuis, 1953b: 54. [nomen nudum] 

Cinetorhynchus manningi Okuno, 1996a: 725; Figs 1-2. [Caribbean Sea, Virgin Islands, Eagle Shoal, 10.5 

m] (Fig. 22) 

Cinetorhynchus reticulatus Okuno, 1997a: 49; Plate 1G-H; Figs 10-11, 12A-C. [Loyalty Islands, Banya Islet, 

Uvea Island, 20°35.8'S 166°16.7'E, 27 m] 

Cinetorhynchus rigens (Gordon, 1936) 

= Rhynchocinetes rigens Gordon, 1936: 76; Figs 1-4, 5e. [Potinha Bay, Madeira] 

Cinetorhynchus striatus (Nomura & Hayashi, 1992) 

= Rhynchocinetes striatus Nomura & Hayashi, 1992: 199; Figs 1-4. [Kadena Port, Okinawa Island, 1-10 m] 

Rhynchocinetes H. Milne edwards, 1837 

= Rhynchocinetes H. Milne Edwards, 1837 (type species Rhynchocinetes typus H. Milne Edwards, 

1837, by monotypy, gender masculine) 

Rhynchocinetes albatrossae Chace, 1997: 28; Figs 15-16. [Surigao Strait, east of Leyte, 10°27'30"N 125°18'E, 

123 m] 

Rhynchocinetes australis Hale, 1941: 270; Fig. 8. [Edithburgh, South Australia]


Rhynchocinetes balssi Gordon, 1936: 85; Fig. 7a-b; nomen novum for Rhynchocinetes typus sensu Balss, 

1922 nec H. Milne Edwards, 1837. [Masatierra] 

Rhynchocinetes brucei Okuno, 1994b: 29; Figs 1-3, 4a-b; Plate 1. [Long Ke Wan, Hong Kong, 8 m] 

Rhynchocinetes conspiciocellus Okuno & Takeda, 1992: 64; Figs 1-3, 4A-D; Plate 1A-B. [Sokodo Beach, 

Hachijo-jima Island, Izu Islands, Japan, 33°07'N 139°49'E, 1 m] 

Rhynchocinetes durbanensis Gordon, 1936: 83; Figs 5b-b, 7c-d; nomen novum for Rhynchocinetes typus 

sensu Stebbing, 1917c nec H. Milne Edwards, 1837. [Durban] 

Rhynchocinetes enigma Okuno, 1997b: 13; Figs 1-3. [34°17'S 132°42'E, Great Australian Bight, approx. 15 

km west-south-west off Pearson Islands, 140-160 m] 

Rhynchocinetes holthuisi Okuno, 1997c: 43; Figs 1-3, 4a-e. [Aqaba, Gulf of Aqaba, Jordan] 

Rhynchocinetes ikatere Yaldwyn, 1971: 87. [off Mayor Island, Bay of Plenty, 37°15'S 176°12'E, 80-120 fms] 

Rhynchocinetes kuiteri Tiefenbacher, 1983: 121; Figs 1-3. [before Portsea at the mouth of Port Phillip Bay, 

Victoria] 

Rhynchocinetes rathbunae Okuno, 1996b: 309; Figs 1-3. [Hawaiian Islands, O’ahu, Waimea Bay, 21°38.0'N 

158°4.0'W] 

Rhynchocinetes serratus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= H.[ippolyte] serratus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 377. [la baie de 

Jarvis] 

= Rhynchocinetes rugulosus Stimpson, 1860a: 36. [in portu Jacksoniensi Australiae] 

Rhynchocinetes typus H. Milne Edwards, 1837: 168; Plate 4c. [Type locality not indicated] 

Rhynchocinetes uritai Kubo, 1942b: 30; Figs 1-3. [Kominato, Tiba Prefecture; lectotype designation by 

Okuno & Takeda, 1992] 

Family XIPHoCARIDIDAe ortmann, 1895 

Xiphocaris von Martens, 1872 

= Xiphocaris von Martens, 1872 (type species Hippolyte elongatus Guérin-Méneville, 1855 [in Guérin- 

Méneville, 1855-1856], by monotypy, gender feminine); name placed on the Official List of Generic 

Names in Zoology in Opinion 470 in 1957) 

Fig. 23. Xiphocaris elongata (Guérin-Méneville, 1855). Photo by Chris Lukhaup.


Xiphocaris elongata (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) (Fig. 23) 

= Hippolyte elongatus Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xx; Plate 2, Figs 16- 

16a. [costas de Cuba] 

= Oplophorus americanus de Saussure, 1858: 56; Fig. 31. [embouchure des rivières de l’Ile de Haïti] 

= Xiphocaris gladiator Pocock, 1889: 18; Plate 2, fig. 6. [Laiou] 

= Xiphocaris gladiator var. intermedia Pocock, 1889: 19; Plate 2, fig. 7. [Laiou] 

= Xiphocaris brevirostris Pocock, 1889: 20; Plate 2, figs 5-5a. [Dominica] 

= Xiphocaris elongata typica Ortmann, 1895: 400. [Type locality not indicated] 

Xiphocaris gomezi Juarrero de Varona, 1993: 2; Figs 2-4. [Agua Revé, Sierra Maestra, provincia Santiago 

de Cuba, Cuba, altura aproximada de 1000 m snm] 

Superfamily PSAlIDoPoDoIDeA Wood-Mason [in Wood-Mason & Alcock, 1892] 

Family PSAlIDoPoDIDAe Wood-Mason [in Wood-Mason & Alcock, 1892] 

Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892] 

= Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892] (type species Psalidopus Huxleyi Wood- 

Mason [in Wood-Mason & Alcock, 1892] designated by Holthuis, 1955b, gender masculine; name 

placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Psalidopus barbouri Chace, 1939: 36. [Nicholas Channel, south of Cay Sal Bank, 23°21'N 79°58'W, 300-315 

fms] 

Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892] (Fig. 24) 

= Psalidopus Huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]: 273; Plate 14, figs 1-2, 7. [7.5 

miles E of N Cinque Island, Andaman Sea, 490 fms] 

= Psalidopus spiniventris Wood-Mason [in Wood-Mason & Alcock, 1892]: 274; Plate 14, figs 3-6a, 8; 

Plate 15. [8 miles SE of Cinque Island, Andaman Sea, 500 fms] 

= Psalidopus japonensis Kubo, 1952: 91; Figs 1-2; Plate 5. [off Daiō-zaki, Kii Peninsula, ca. 530 m] 

Psalidopus tosaensis Toriyama & Horikawa, 1993: 1; Figs 1-3, 5. [off Muroto Cape, Kochi Pref., Japan, 

32°13'12''N 134°01'36''E, 2765-2881 m] 

Fig. 24. Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]. Photo by Tin-Yam Chan.


Superfamily STyloDACTyloIDeA Spence Bate, 1888 

Family STyloDACTylIDAe Spence Bate, 1888 

Bathystylodactylus Hanamura & Takeda, 1996 

= Bathystylodactylus Hanamura & Takeda, 1996 (type species Stylodactylus bathyalis Cleva, 1994, by 


Bathystylodactylus bathyalis (Cleva, 1994) 

= Stylodactylus bathyalis Cleva, 1994: 56; Fig. 2. [Bligh Canyon, Coral Sea, 12°23'S 146°08'E. 3515- 

3502 m] 

Bathystylodactylus echinus Wicksten & Martin, 2004: 377; Figs 1-5. [basin off Magdalena Bay, Baja California, 

Mexico, 24°35'N 113°25'W, 3563-3621 m] 

Bathystylodactylus inflatus Hanamura & Takeda, 1996: 930; Figs 1-3. [off east coast of Taiwan, 23°42'03''N 

123°45'08''E, 3436-3452 m] 

Neostylodactylus Hayashi & Miyake, 1968a 

= Neostylodactylus Hayashi & Miyake, 1968a (type species Stylodactylus amarynthis De Man, 1902, by 


Neostylodactylus affinis Hayashi & Miyake, 1968a: 605; Fig. 7. [Korea Strait, 33°34.9'N 128°25.2'E, 120 m] 

Neostylodactylus amarynthis (De Man, 1902) 

= Stylodactylus amarynthis De Man, 1902: 897; Plate 27, figs 64-64b. [Ternate] 

Neostylodactylus investigatoris (Kemp, 1925) 

= Stylodactylus investigatoris Kemp, 1925: 260; Figs 1-2. [2 miles off Great Torres Island, Mergui 40 fms] 

Neostylodactylis litoralis Okuno & Tachikawa, 2000: 39; Figs 1-4. [Takinoura, Ani-jima Island, Ogasawara 

Islands, 5 m] 

Neostylodactylus sibogae (De Man, 1918a) 

= Stylodactylus Sibogae De Man, 1918a: 159. [Siboga Expedition stn 95, 5°43'.5N 119°40'E, Sulu Sea, 

522 m] 

Parastylodactylus Figueira, 1971 

= Parastylodactylus Figueira, 1971 (type species Stylodactylus bimaxillaris Spence Bate, 1888, by original 


Parastylodactylus bimaxillaris (Spence Bate, 1888) 

= Stylodactylus bimaxillaris Spence Bate, 1888: 855; Plate 138, fig. 3. [Challenger stn 219, 1°54'0"S 

146°39'40"E, off the Admiralty Islands, 150 fms] 

Parastylodactylus hayashii (Komai, 1997) 

= Neostylodactylus hayashii Komai, 1997a: 125 (in part); Figs 1-3. [R/V Tansei-maru, KT95-5 cruise, stn 

TB18-2, Okinoyama, off Sunosaki, Boso Peninsula, 34°59'N 139°39'E, 105-113 m] 

Parastylodactylus longidactylus Cleva, 1990: 125; Figs 12d, 15a-b, 16a-d. [Philippines, 13°55.2'N 120°30.5'E, 

320-318 m] 

Parastylodactylus moluccensis Cleva, 1997: 398; Fig. 3. [Indonésie, 05°17'06''S 132°51'19''E, 315-348 m] 

Parastylodactylus richeri Cleva, 1990: 127; Figs 15c, 16e-h. [Nouvelle-Calédonie, 23°45'S 167°12'E, 380 

m] 

Parastylodactylus semblatae Cleva, 1990: 122; Fig. 12c, 14, 18c. [Nouvelle-Calédonie, 22°16.5'S 167°16.5'E, 

475-500 m] 

Parastylodactylus tranterae Cleva, 1990: 119; Figs 11b, 12b, 13. [Nouvelle-Calédonie, 22°11.3'S 167°15.0'E, 

495-550 m] 

Stylodactyloides Cleva, 1990 

= Stylodactyloides Cleva, 1990 (type species Stylodactyloides crosnieri Cleva, 1990, by monotypy, gender 

masculine) 

Stylodactyloides crosnieri Cleva, 1990: 129; Figs 17, 18d-e. [Iles Chesterfield, 19°53.20'S 158°39.50'E, 370- 

400 m]

Part 1 - AToL Decapoda

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