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Carideorum Catalogus: 

The Recent Species of the Dendrobranchiate, Stenopodidean, 

Procarididean and Caridean Shrimps (Crustacea: Decapoda) 

S. De Grave & C.H.J.M. Fransen* 

De Grave, S. & C.H.J.M. Fransen. Carideorum catalogus: the recent species of the dendrobranchiate, 

stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). 

Zool. Med. Leiden 85(9): 30.ix.2011: 195-589 figs 1-59.— ISSN 0024-0672, ISBN 978-90-6519-200-4. 

S. De Grave, Oxford University Museum of Natural History, Parks Road, Oxford, OX1 3PW, United 

Kingdom (sammy.degrave@oum.ox.ac.uk). 

C.H.J.M. Fransen, Department of Marine Zoology, Netherlands Centre for Biodiversity Naturalis, P.O. 

Box 9517, 2300 RA Leiden, The Netherlands (charles.fransen@ncbnaturalis.nl). 

* The order of authorship is purely alphabetical, with both authors having equally contributed to this 

work. 

Key words: Crustacea; Decapoda; Dendrobranchiata; Stenopodidea; Procarididea; Caridea; checklist. 

A checklist of recent species of dendrobranchiate, stenopodidean, procarididean and caridean 

shrimps including synonyms and type localities. Also listed are unavailable names, larval names, 

nomina dubia and nomina nuda. A complete list of references to original descriptions of taxa listed is 

provided. 

Contents 

Introduction ............................................................................................................................................................ 196 

The higher classification of shrimp .................................................................................................. 196 

Fossil taxa ........................................................................................................................................................ 197 

How many shrimp species are there? ............................................................................................ 197 

Structure of the list ............................................................................................................................................. 197 

Notes on type locality ....................................................................................................................................... 199 

Notes on dating of some papers ................................................................................................................ 200 

Notes on author names .................................................................................................................................... 201 

Notes on authorship of taxa ......................................................................................................................... 202 

Notes on some papers of nomenclatorial significance ................................................................. 203 

Notes on the spelling of some names ..................................................................................................... 204 

Notes on the gender of generic names and gender agreement in species names ........ 204 

Acknowledgements ........................................................................................................................................... 205 

Checklist ................................................................................................................................................................... 206 

Suborder Dendrobranchiata ................................................................................................................ 206 

Suborder Pleocyemata ............................................................................................................................. 248 

Infraorder Procarididea .......................................................................................................................... 248 

Infraorder Stenopodidea ........................................................................................................................ 249 

Infraorder Caridea ..................................................................................................................................... 254 

Unavailable names ..................................................................................................................................... 471 

Penaeoidea Larvata .................................................................................................................................... 471 

Penaeoidea Nomina dubia .................................................................................................................... 471

196 De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 

Penaeoidea Nomina nuda ..................................................................................................................... 472 

Sergestoidea Larvata ................................................................................................................................. 472 

Sergestoidea Nomina dubia ................................................................................................................. 472 

Sergestoidea Nomina nuda .................................................................................................................. 473 

Caridea Larvata ............................................................................................................................................ 473 

Caridea Nomina dubia ............................................................................................................................ 475 

Caridea Nomina nuda ............................................................................................................................. 476 

References ................................................................................................................................................................ 478 

Systematic index to currently valid families and genera ............................................................ 585 

Introduction 

The higher classification of shrimp 

Over the last decade or so, much has been written on the classification of Decapoda, 

fuelled by a surge in molecular phylogenetic studies, as well as close scrutiny of internal 

and external morphological characteristics. As discussed by Fransen & De Grave 

(2009), such studies on shrimps are still somewhat ”thin on the ground”, at least compared 

to the more extensive work done on the Brachyura and Anomura. At a higher 

level in decapod classification it has long been recognised that three distinct lineages of 

shrimps can be distinguished: Dendrobranchiata, Stenopodidea and Caridea, a system 

which has not been seriously challenged by recent studies. 

The internal classification of Dendrobranchiata and Stenopodidea alike has been 

stable for some time, with the only major addition being the family Macromaxillocarididae 

Alvarez, Iliffe & Villalobos (2006) to the Stenopodidea in recent years. 

A different picture has emerged for Caridea very recently with Bracken et al. (2009) 

and Chan et al. (2010), both drawing attention to the non-monophyletic status of certain 

superfamilies and families. Further, we are aware of work currently in progress (some 

by the authors of this compilation) corroborating the hypothesis that the current classification 

of Caridea is unnatural, lines of study which will lead to the resurrection of 

certain family names as well as further refinement to other families. As one of our objectives 

for the current effort was to link this compilation of species level information 

with the earlier work by Chace (1992) for families and Holthuis (1993a) for genera, we 

have elected to largely follow the classification outlined by De Grave et al. (2009) which 

builds upon this earlier work. As such, it was deemed advisable to include the recently 

resurrected family Acanthephyridae Spence Bate, 1888 in the superfamily Oplophoroidea, 

rather than in this catalogue to create a new superfamily, which would perhaps 

be more congruent with the results in Chan et al. (2010). 

Although we follow herein the classification scheme of De Grave et al. (2009), two 

recent changes have been implemented. The clarification of the status of Galatheacaris 

abyssalis Vereshchaka, 1997a, as the megalopal stage of Eugonatonotus chacei Chan & Yu, 

1991a, by De Grave et al. (2010) resulted in the removal of the family Galatheacarididae 

and superfamily Galatheacaridoidea in the current listing. Bracken et al. (2010) clarified 

the status of the family Procarididae, resulting in the recognition of a fourth group of 

shrimp, Infraorder Procarididea.

De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 197 

Fossil taxa 

The current listing is restricted to extant genera, species and subspecies only. For 

readers interested in how the current classification of extant taxa compares with those 

of fossil taxa, we refer to De Grave et al. (2009) who employed a suprageneric classification 

of Decapoda across both fossil and extant taxa, as well as Schweitzer et al. (2010) 

who list all species (and their classification) of fossil species. Schweitzer et al. (2010) list 

97 fossil species of Dendrobranchiata in 31 genera, two fossil species in Stenopodidae 

(each in its own genus) and 52 species of fossil Caridea in 33 genera. Their placement of 

the genus Udora Münster, 1839, within the superfamily Procaridoidea (herein considered 

at infraordinal level) is interesting, as this group is currently only known from six 

species, restricted to anchialine habitats. If the five known species of Udora are indeed 

correctly placed, the fossil record of the groups extends back to the Middle Jurassic, an 

interesting observation as the first extant species was only discovered in 1971, drawing 

parallels with the discovery of coelacanths, one of the better known living fossils. 

How many shrimp species are there? 

Fransen & De Grave (2009) working from a preliminary version of the present 

catalogue estimated the species/subspecies richness of shrimps as follows: Dendrobranchiata 

(505), Stenopodidea (58) and Caridea (circa 3108), with those figures slightly 

adjusted in De Grave et al. (2009). Since then numerous descriptions have appeared, 

synonymies corrected and species placed in synonymy. The current tally stands as follows: 

Dendrobranchiata (68 genera, 533 species), Procarididea (2 genera, 6 species), 

Stenopodidea (12 genera, 71 species) and Caridea (389 genera, 3438 species). Although 

the figures listed in De Grave et al. (2009) for the other decapod taxa have clearly been 

adjusted by now, the Caridea remain the second most species-rich group within the 

Decapoda, with approximately half as many species as Brachyura, the dominant taxon 

within Decapoda. 

On a more inclusive level, the Caridea are dominated by Palaemonidae (981 species), 

followed by Alpheidae (663), Atyidae (469), Hippolytidae (338) and Crangonidae 

(219). Under the traditional classification employed here, Palaemonidae comprises two 

subfamilies, with Pontoniinae being considerably richer in species (602) than Palaemoninae 

(379). In contrast to those species-rich families, are those 12 families which are 

monogeneric, some only consisting of a single species, e.g. Physetocarididae. 

On a generic level, the most speciose genera are Caridina (Atyidae, 290 species), Alpheus 

(Alpheidae, 286), Macrobrachium (Palaemoninae, 243), Synalpheus (Alpheidae, 159) 

and Periclimenes (Pontoniinae, 152). As doubt has been cast on the monophyletic status 

of these mega-genera in recent years, this ranking may substantially alter when phylogenetic 

studies progress. 

Structure of the list 

As discussed above, we recognise herein four major groups of shrimp: Infraorder 

Dendrobranchiata and suborders Procarididea, Stenopodidea and Caridea. For each 

of those higher categories we list the currently recognized superfamilies and families,


following De Grave et al. (2009) with the changes noted above. As the focus of the current 

compilation is on genera and species, for the sake of brevity we have not provided 

the historical information for superfamilies and families, nor have we listed their synonyms. 

These can be found in Holthuis (1993a) for Stenopodidae and Caridea (including 

Procarididae) and Pérez Farfante & Kensley (1997) for Dendrobranchiata. For each genus, 

we provide the reference to the original description, as well as on type species (and 

the way in which they were designated) as well as their gender. In addition, we list all 

their known synonyms. For species and subspecies, we list their original generic designation, 

in original orthography for both genus and species, as well as all known synonyms. 

For each, we provide the page number on which the species description starts, as 

well as a reference to all figures and/or plates on which the species is illustrated, as well 

as their type locality (see below). Lectotype and neotype designations are included, as 

well as additional comments when required. Additional comments concern dating issues, 

as well as figures and/or further descriptions by the same author (often again as 

sp. nov.) in a later work. For instance, Parapasiphaë gilesii was illustrated in Wood-Mason 

(1892), thus validating the name, but without any accompanying textual description 

which appeared later in Wood-Mason & Alcock (1893). Such cases have been annotated 

to provide clarity for future researchers. 

All species names are given in their original orthography. This also includes usage of 

the Latin ligatures, æ and œ when present in a species name, such as in Palæmon longicornis 

Olivier, 1811, or Œdipus gramineus Dana, 1852a. Although we are aware that the 

ICZN Code does not differentiate this orthography, we felt this to be more appropriate in 

view of the long history of taxonomy. For instance we list Pelias Niloticus Roux, 1833 (now 

Caridina nilotica) and Pal.[æmon] Swainsonii White, 1847a (a synonym of Macrobrachium 

acanthurus), as written in their original descriptions. The square brackets in the latter case 

signifying an addition on our part, to make the generic designation clearer. 

As a general rule the latest taxonomic or systematic treatment for each species were 

followed, listing species as valid or as a synonym accordingly. This does not take into 

account the uncertainties regarding the true status of some taxa, which in reality may 

be valid species instead of a synonym and vice versa, but does reflect our current understanding 

of species richness. One notable exception is that the sergestid genera 

erected by Judkins & Kensley (2008) were not recognised in a more recent revision by 

Vereshchaka (2009), who instead recognised several species groups in Sergestes sensu 

lato. As these species groups largely correspond to the new genera proposed by Judkins 

& Kensley (2008), we have elected to continue to treat them as genera, necessitating 

some generic rearrangements to new species proposed in Vereshchaka (2009). 

Nomina dubia and nomina nuda have been listed as synonyms under currently 

valid names if such information was available, if not they have been listed as a separate 

category. We were aided herein by the Holthuis card catalogue, who listed many such 

names even under appropriate family headings. As Holthuis’s logic was not always 

transparent and other names surfaced or were assigned differently by other workers, 

we have not followed his family level assignations, but have retained them under three 

categories: Penaeoidea, Sergestoidea and Caridea. 

The scientific literature abounds with misspellings of species names due to printing 

and typographical errors. Listing all of these would enlarge the current volume manyfold. 

Thus a minor error, such as Palaemon pandaliformes instead of Palaemon pandali-


formis (Stimpson, 1871) in Tavares (1993) we do not list. However, if the magnitude of 

the error is such that it could possibly be interpreted as a different name we have included 

those, to avoid further confusion. For instance, Burukovsky (1974) used Artisteus 

viritli for Aristeus virilis Spence Bate, 1881, or Acanthephyra parvirostris in Coutière 

(1911a), an erroneous spelling of A. brevirostris Spence Bate, 1888. 

During the early years of ocean exploration, numerous larval forms were described, 

for instance by Spence Bate (1888) and Ortmann (1893). Although on occasion these 

have been linked to adult forms, as is the case for Embryocaris stylicauda Ortmann, 1893, 

which is a larval stage of Stenopus hispidus (Olivier, 1811), many remain unassigned to 

adult forms at present. Those which have not been linked are herein listed under the 

headings of Penaeoidea, Sergestoidea and Caridea Larvata. 

The present catalogue includes the binomial names for shrimps from Linnaeus 

(1758) onwards, up to 1st June 2011. We have not considered pre-Linnean names, even 

though sometimes these have been mentioned in later literature, for instance in the 

works by Holthuis (1991), Tavares (1993) and others. The cut-off date for inclusion of 

names was set at 1st June 2011, although we are acutely aware that a number of names 

are currently in press and some of these will have appeared in the scientific literature 

before the present work sees the light of day. 

Notes on type locality 

For all species we have listed their type locality in one of two ways. On the whole 

we have done this by reference to the original type series, but restricted to holotype 

locality if a holotype is indicated, as is the norm in recent papers. For instance, for Periclimenes 

cannaphilus Komai & J.N. Kim, 2010, we only list the locality of Kasuga 2 Seamount 

where the holotype originated from and not the locality of the 15 paratypes from 

Nikko and Daikoku Seamounts. If no holotype was indicated in the original publication, 

then we have included all syntypic localities in the listed type locality, resulting in 

upwards of 10 stations being listed for some species from the early expeditions (e.g. 

Spence Bate, 1888). If a later nomenclatorial act selected a lectotype from the syntypic 

series, we have only listed the locality details for that particular specimen. If a neotype 

was selected, then the information for the neotype only features, and not the original 

type series. For example, Cai et al. (2006) selected neotypes for the six Japanese species 

of Atyidae described by Stimpson (1860a), the original type series being lost in the great 

Chicago fire of 1871. Thus the type locality information is only given for the neotypes, 

and the reader is referred to the discussion in Cai et al. (2006) or the original publication 

by Stimpson (1860a) for locality information for the original (now lost) syntypes. For 

the majority of taxa we have elected to adhere to the original language of the description 

in our listing of type locality, as well as the original orthography and names of localities. 

For example, Stimpson (1860a) describes Alpheus pachychirus and other species 

from ”ad insulam Loo Choo”, currently known as the Ryukyu Islands in southern Japan. 

We accept the potential criticism that this may make this part of the current listing 

somewhat difficult to work with for non-taxonomists, but in our opinion it accurately 

reflects the rich tapestry of species descriptions, and correctly emphasizes the historical 

heritage and multinational aspect of alpha-level taxonomy. By necessity, we had to accept 

our own linguistic limitations and have had (by other taxonomists who are more


proficient in such matters) localities originally listed in Russian, Chinese and Vietnamese 

translated. These are simply listed in English in the present compilation. 

In many earlier publications no type localities are listed, bar a reference to a certain 

expedition station or a vague collection locality but with a specific station included. For 

instance, Rathbun (1902a) described 50 taxa from the coast of Alaska southwards to San 

Diego, based on samples which had accumulated for many years in the Smithsonian as 

a result of the Albatross expeditions. The majority of these have vague localities for the 

syntypic series in the actual publication, but do include a specific reference to one or 

more Albatross stations. By reference to various published lists, we have added such 

information between brackets in the herein listed type locality information. These may 

need to be corrected once the material and original labels are re-examined as part of 

future revisions. Several sources were consulted for this, notably Anon. (1914) for Investigator 

stations, Murray & Hjort (1912) for the Michael Sars expedition, as well as Sewell 

(1935) for species in Calman (1939) and others. For some taxa, such information could 

not be directly obtained from published station lists or the original description, but 

later authors have tracked down such information. For example, Crosnier & Forest 

(1973) provided accurate locality information for Gennadas talismani Bouvier, 1906a, and 

Metapenaeus perlarum Nobili, 1905a. In such instances, we have always included a reference 

as to where the information came from. On a few occasions, the actual type description 

nor any later paper discussed the exact type locality details. For instance 

Macpherson (1984) when describing Nematocarcinus gladius listed three stations without 

further details with the material being deposited in the South African Museum 

(now IZIKO). In those instances we made direct enquiries with the museums involved 

and provide here the information on the specimen labels and their catalogues. 

Notes on dating of some papers 

A very important aspect of nomenclature is the accurate dating of nomenclatorial 

acts, something which is not always understood by colleagues in other biological disciplines. 

For an extreme example of this issue we refer to Low & Guinot (2010) who discuss 

a case of 48 hour precedence. Dating issues have been often reported for older 

references and are most prominent in early works, such as Guérin-Méneville (1829- 

1838) and Leach (1815-1875) to name but two. In those cases where dating issues have 

been reported in the literature, discrepancies were noted in various draft versions of the 

current catalogue or have been brought to our attention by our colleagues, we have 

endeavoured to decide upon the correct date of publication. For this we have relied on 

a variety of sources, such as Rathbun (1897) for Herbst (1791-1796), Gordon (1959) for 

Bell (1844-1853), d’Erasmo (1949) for O.G. Costa & A. Costa (1838-1871), Holthuis (1947a) 

for Sowerby (1804-1806) and Holthuis (1961a) for Guérin-Méneville (1829-1838). In addition, 

we followed Harrison & Smith (2008) for dating of all of Leach’s publications. 

For dating of individual issues of certain journals with a known history of confusion, 

we followed Duncan (1937) and Dickinson (2005) for Proceedings of the Zoological Society 

in its numerous guises as well as their Transactions, Evenhuis (2003) for the Annals and 

Magazine of Natural History and others. 

Contrary to popular belief, such dating issues are not just restricted to the earlier 

taxonomic works and are almost as ubiquitous in more recent decades, although usually


much easier to solve. For example, the description of Plesionika williamsi Forest, 1964 is 

in the volume of the Bulletin du Muséum national d’Histoire naturelle (2ème série) for 1963, 

but this particular issue was printed on the 19th June 1964 (R. Cléva, pers. comm.). A 

peculiar case are the two pontoniine genera Exoclimenella Bruce, 1995, and Periclimenella 

Bruce, 1995. As already pointed out by Holthuis (1996) the descriptions of these 

two genera in Bruce (1995) is valid under the current ICZN Code and thus predates the 

descriptions (as gen. nov.) in Ďuriš & Bruce (1995). However, this assessment was 

based on a publication date of 30th November 1994, as indicated on the frontispiece. 

Per Koeltz (Koeltz Scientific Books) has confirmed that the actual publication date of 

Bruce (1995) was 16th March 1995, which curiously does not affect the seniority of 

those descriptions as the Ďuriš & Bruce (1995) article appeared in the May/June issue 

of Journal of Natural History, which according to Evenhuis (2003) was published on 31st 

May 1995. 

In those cases where we identified a discrepancy between the date usually associated 

with a publication and the real publication date determining priority, we have 

annotated the references following the conventions used by Schweitzer et al. (2010). 

Some of the early descriptions appear in multi-year works, such as Guérin-Méneville 

(1829-1838) and H. Milne Edwards (1836-1844). For these, we follow the quotation style 

in De Grave et al. (2009). For example, the original description of Xiphocaris elongata 

appears in Guérin-Méneville (1855-1856), as Hippolyte elongatus with 1855 as the publication 

date of the actual nomenclatorial act. This is thus listed as Hippolyte elongatus 

Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856] to distinguish the nomenclatorial 

act from the bibliographical reference. 

Notes on author names 

The numerous works by Charles Spence Bate (1819-1889) have usually been cited in 

modern literature under the name of ”Bate, C.S.”, although invariably Charles himself 

used the name ”Spence Bate, C.”. It is not clear when Charles started using ”Spence 

Bate” as a surname, nor the exact reasons why, nor indeed if he just adopted this name 

in publications and continued to use ”Bate” in his private life. According to the Oxford 

National Dictionary of Biography his mother was Harriet Spence (1788-1879) and we 

postulate here that he added Spence to his surname to distinguish himself from his father 

Charles Bate (1789-1872) who was also a dentist in Cornwall. We have checked all 

his carcinological papers, as well as most of his dentistry ones and find that indeed he 

invariably signed off in all his papers as ”C. Spence Bate”. This is also the surname 

by which his contemporaries refer to him in their own publications (e.g. Semper, 1868; 

Miers, 1884a; A. Milne-Edwards, 1891, etc.), although sometimes abbreviated to ”Sp. 

Bate”. In the two editorial notes by John Murray in Spence Bate, 1888 it is stated: ”…illustrate 

the text of Mr. Spence Bate’s report…” as well as: ”The Report on the Crustacea 

Macrura, by C. Spence Bate, Esq….”. Within the text of Spence Bate (1888) itself the 

author always (with one exception) refers to his own, older work as ”Spence Bate”, often 

shortened in the synonymy listings as ”Sp. B”. The exception being when he refers 

to ”Bate & Westwood (1863, A History of British Sessile-eyed Crustacea)”. We interpret that 

here as a lapsus, as the title page of that work also identifies the authors as C. Spence 

Bate and J.O. Westwood.


The present case mirrors the consistent usage of H. Milne Edwards in publications 

by that author, whose surname was Edwards, but who added Milne to distinguish himself 

from his brother William Edwards (see Forest, 1996). As we can see no reason why 

the name Charles Spence Bate should be treated different to Henri Milne Edwards, we 

here accept the arguments put forward by Dubois (2008) in that authors are signatures 

and not persons, and thus have elected to use the name C. Spence Bate. 

For the father and son team of Henri Milne Edwards and Alphonse Milne-Edwards, 

we herein follow Forest (1996) in the spelling of their surnames, and have added, as is 

customary in carcinology, their initial to all citations. Issues surrounding the use of 

Guérin and Guérin-Méneville were discussed in Ng et al. (2008) and we follow here 

their lead in attributing names after 1836 onwards to Guérin-Méneville and before 1836 

to Guérin. 

Ng et al. (2008) also provide a more in depth discussion on the spelling of author 

names and how these should be cited in the references. As we herein adhere to the same 

rules, we refer the reader to that treatise. 

In those instances where different taxa have been described by different authors 

who share the same surname, we generally have adopted the same rules as outlined in 

Ng et al. (2008). In brief, we do not add the authors initials to the person having described 

significantly more taxa, but add those to the author having described fewer. 

For instance, for taxa described by Peter K.L. Ng we do not add P.K.L. in front of the 

surname, whilst we do so for Ngan Kee Ng (thus N.K. Ng). As in Ng et al. (2008) we 

do not include initials if the authors were not contemporaries, the intervening time lag 

making this sufficiently obvious. For instance, J.Y. Johnson (1863, 1868) and D.S. Johnson 

(1961, 1962, 1967, 1973). A special case is the citation of first names for workers of 

Chinese descent or nationality, where fashion has dictated how names are used rather 

than bibliographical accuracy. For instance, the Chinese freshwater carcinologist Guo 

Zhao Liang has published under the names Guo Zhaoliang (abbreviated to Guo, Z.), 

Guo Zhao-Liang (Guo, Z.-L.) and Guo Zhao Liang (Guo, Z.L.). As it is evident that in 

all three cases this is the same worker, we have elected not to include his initials in the 

citations, others, more purists perhaps, may disagree. In the case of the team of A.H. 

”Hank” and D.M. ”Dora” Banner, who described numerous alpheid taxa, often jointly, 

they stated several times in their larger publications (e.g. Banner & Banner, 1978; Banner 

& Banner, 1981a) that they list their own papers in strict chronological order, without 

reference to seniority of authorship. We have herein interpreted this as a desire to 

reflect a joint effort with order of authorship, perhaps somewhat arbitrarily decided 

upon and a reflection of equal share in the description. As such we have elected to not 

add initials to citations in the checklist, although order of authorship in the references 

is as given in the actual papers to facilitate bibliographical searches. All this makes for 

less cumbersome citations, however in other cases, we had no option but to include 

initials. 

Notes on authorship of taxa 

As extensively discussed by Ng et al. (2008) attributing the correct author to a certain 

taxon can be confusing, particularly (but not exclusively) in the earlier literature. 

Often in the earlier literature another carcinologists’ name is cited behind the species


name in the work of a different author. Such reference is usually to an unpublished 

manuscript, a personal letter or even a museum label and many examples abound in, 

for instance, H. Milne Edwards (1834-1840). An interesting example is the name Lysmata 

nilita as published in Hope (1851) as a nomen nudum, with Risso MS featuring 

behind the name. Clearly Hope must have had access to unpublished manuscripts by 

Risso, as is evident from Monod (1931). Fittingly, when Dohrn & Holthuis (1950) described 

the second species of Lysmata from the Mediterranean, they adopted the name 

Lysmata nilita. 

As in Ng et al. (2008) we herein strictly adhere to Article 50 of the ICZN and attribute 

authorship only to those who are directly responsible for the name and for satisfying the 

criteria of availability. For example, we thus list Caridina cavalerieioides Liu & Liang in Liang 

(2004), but in contrast in the case of Stenopus devaneyi we use Goy (1984) rather than 

Goy & Randall (1984), as was used in Goy & Randall (1986). Observant readers will thus 

notice a discrepancy between the current list and Holthuis (1993a) for certain names attributed 

to Wood-Mason & Alcock in the The Annals and Magazine of Natural History. This 

series of papers was discussed in Ng (1998) and we herein follow that interpretation. 

Notes on some papers of nomenclatorial significance 

Confusion exists in the literature regarding the type description of the rare alpheid 

Betaeus jousseaumei (now Amphibetaeus jousseaumei), which Holthuis (1993a) attributed 

to Coutière (1897a) but therein dated as 1896. Coutière himself in his magnificent opus 

on Alpheidae (Coutière, 1899a) refers instead to Coutière (1896). We have not been able 

to accurately date either of those publications and herein follow Coutière (1899a) in 

considering Coutière (1896) as the type description, as well as following him in considering 

the publication year of Coutière (1897a) as 1897 and not 1896 as indicated in 

Holthuis (1993a), as indeed do Banner & Banner (1981a) and Anker & Jeng (2006). 

Guérin-Méneville authored the Crustacea section in Ramon de la Sagra’s work 

”Histoire Physique, Politique et Naturelle de l’Île de Cuba”, which also almost contemporaneously 

appeared in a Spanish language version ”Historia fisica politica y natural de la Isla 

de Cuba”. Although nothing is known about the actual publication dates for the Crustacea 

section, it is known that for the Mollusca the French text is the original version, 

which appeared in livraisons and was afterwards translated into Spanish (G. Rosenberg, 

pers. comm.). Curiously for Caridea, the French edition has several more shrimp 

names in the text, which do not feature in the Spanish version. Perhaps this can be explained 

by the fact that these additional species did not originate from Cuba, and thus 

perhaps were deleted from a Spanish translation, targeted at a more local audience. It 

is also known that the dates appearing on the frontispiece of the Mollusca part are incorrect. 

For the French edition it is the date the text was concluded, even though the 

work appeared in livraisons over several years, whilst for the Spanish version it is an 

intermediate date (G. Rosenberg, pers. comm.). In the absence of any firm evidence regarding 

the exact publication date of the crustacean parts, both text and Plates, we have 

no other course of action available than to follow the dates on the frontispiece, even 

though future research will almost certainly prove these to be wrong, this task is now 

taken up by P. Clark. Thus, for the species mentioned in both the French and Spanish 

editions, for instance Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]


we employ 1855 as their description date, corresponding to the date on the frontispiece 

of Tomo VIII (Atlas de Zoologia), with the corresponding text in Tomo VII (Crustáceos, 

Aragnides é Insectos) with 1856 on the frontispiece. For the additional taxa listed in the 

French edition, for example Alpheus Rouxii Guérin-Méneville, 1857, we use 1857 as the 

date, corresponding to what is written on the frontispiece of that volume. 

Notes on the spelling of some names 

In general, we have listed the species in their original spelling, unless a justified 

emendation needed to be made, for example to comply with Art. 31.2 in relation to 

gender agreement between generic and species names. 

When revising the genus Pandalus, Komai (1999) proposed that the species usually 

quoted as Pandalus propinquus in the majority of the taxonomic and ecological 

literature, should revert to its original spelling in the new combination of Atlantopandalus 

propinqvus, as used (Pandalus propinqvus) in the type description by G.O. Sars 

(1870). Under Art. 32.1 we herein follow Komai (1999) and continue to list the species 

as A. propinqvus. 

A similar case exists for the commercially important species, Plesionika narval 

(Fabricius, 1787), which was originally described by Fabricius (1787) under the name 

Astacus narual. Under Art. 33.3.1 we herein maintain the species as P. narval, and do 

not revert back to the original spelling. 

The species Lebbeus microceros (Krøyer, 1841) has been variable listed as L. microceros 

and L. microceras in older literature. Krøyer (1841) originally described the species under 

the name Hippolyte microceras (Krøyer, 1841, p. 578), but a note has been inserted in 

the journal after the table of content, in which the name is corrected to Hippolyte microceros. 

Under Art. 32.5.1.1 the original spelling thus has to be treated as an inadvertent 

error and corrected to L. microceros. 

Acanthephyra eximia Smith, 1884 has often been listed as A. eximea Smith, 1884. Both 

names were used by Smith (1884). Smith himself used A. eximia in his publication of 

1886 (pp. 189, 190, 192). In between no other author had used both names in an article 

and selected a correct spelling as First Reviser, thus here Art. 24.2.4 holds and Smith 

himself becomes the First Reviser. 

Notes on the gender of generic names and gender agreement in species names 

As already discussed by Ng et al. (2009) there has been confusion about the gender 

of generic names, as well as the correct ending of species names, if adjectival in nature. 

If the original genus description stated gender, we have simply followed this. Care 

needs to be taken when assuming gender though. For instance, Eumanningia Crosnier, 

2000, may sound feminine, but as clearly stated by Crosnier (2000), the name has to be 

treated as masculine. In the majority of older descriptions the gender was not stated, in 

all those cases we follow Holthuis (1983) and Pérez Farfante & Kensley (1997) for the 

gender of the genus, with one exception. The gender of the genus Petalidium Spence 

Bate, 1881, was treated as masculine by Pérez Farfante & Kensley (1997). We herein treat 

it as neuter, in recognition of the name of the type species, Petalidium foliaceum, as well 

as the other species assigned to this genus.


Confusion exists about the correct ending of some species names, as it is not always 

clear from the type description if a species name has to be treated as a noun or an adjective, 

especially in older descriptions when derivations were not commonly included. 

For some, the adjectival nature of the name is abundantly clear, and the necessary 

changes have been introduced in the present listing. For others, our own limited knowledge 

of the classical languages has prevented us from forming a definite conclusion 

on this matter. In those few cases, we have been fortunate that Lipke Holthuis, a master 

of classical names, used these names before and we have simply followed suit. For 

instance, even though the genus Halocaridinides is masculine, Holthuis (1982) used H. 

trigonophthalma, effectively treating it as a noun, instead of the masculine form trigonophthalmus. 

Acknowledgements 

Over the long gestation period of this catalogue, numerous people provided assistance 

with tracking down references, translated relevant papers, advised on nomenclatorial 

issues, spelling of author names and dating issues, provided us with copies of 

their own work and checked our listing in their ”special” families. We are immensely 

grateful to them all. We list them here in alphabetical order: A. Anker, C.W. Ashelby, 

A.J. Bruce, R.N. Burukovsky, Y. Cai, T.-Y. Chan, P. Clark, P.C. Dworschak, C. d’Udekem 

d’Acoz, J.W. Goy, L.Y.D. Goulding, R. Gulledge, K.-I. Hayashi, M.E. Hendrickx, S. 

Ivanenko, W. Klotz, T. Komai, R. Lemaitre, X. Li, M.E.Y. Low, D. Mann, I. Marin, T. 

Naruse, P.K.L. Ng, K. Pocklington, Nguyen Thanh Son, K. Reed, J. Poupin, S.H. Tan, 

A.L. Vereshchaka, M.K. Wicksten and K. Wouters, whilst offering our apologies to 

those who assisted along the way and are not listed. We are also grateful to the librarians 

of the Koninklijk Belgisch Instituut voor Natuurwetenschappen (KBIN-IRSNB), 

Netherlands Centre for Biodiversity Naturalis, University College Cork, the Hope Library 

(OUMNH) and the Radcliffe Science Library, Oxford who provided references 

and expedited the necessary inter-library loans. For the many photographs used we 

thank A. Anker, A.J. Bruce, T.-Y. Chan, N. Ciccarese, L. Harris, O. Helker, J. Hoover, T.M. 

Iliffe, T. Komai, G.A. Lovrich, C. Lukhaup, K. Neely, T. Sakihara, E. del Solar, J. Tomas, 

and C. d’Udekem d’Acoz. We are especially grateful to A. Anker, C. d’Udekem d’Acoz, 

P.K.L. Ng and P.C. Dworschak, who went through a near complete, final version of the 

catalogue and corrected numerous errors. Any remaining errors are the authors responsibility. 

Lastly, we must pay homage to L.B. Holthuis (1921-2008) who kept meticulous 

records of species names of shrimps and erroneous spellings, many never reported 

again in the scientific literature. Without access to his files, this catalogue would have 

been far less complete. Although sadly Lipke is no longer with us and able to see the 

finished product, we are grateful for his discussions on a preliminary version, during 

which he recommended the inclusion of type localities. To his memory, we dedicate this 

catalogue.


Checklist 

Suborder Dendrobranchiata Spence Bate, 1888 

Superfamily PeNAeoIDeA Rafinesque, 1815 

Family ARISTeIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Aristaeomorpha Wood-Mason in Wood-Mason & Alcock, 1891a 

= Aristæomorpha Wood-Mason in Wood-Mason & Alcock, 1891a (type species Aristeus rostridentatus 

Spence Bate, 1881, a junior subjective synonym of Penaeus foliacea Risso, 1827, by original designation, 

gender feminine) 

Aristaeomorpha foliacea (Risso, 1827) (Fig. 1) 

= Penaeus foliacea Risso, 1827: 69; Plate 2, fig. 6. [l’Europe Méridionale, environs de Nice et des Alpes 

Maritimes, grandes profondeurs] 

= Aristeus rostridentatus Spence Bate, 1881: 189. [near the Fiji Islands, 300 fms; according to Spence 

Bate, 1888: Challenger stn 173, 19°09'35"S 179°41'50"E] 

= Aristaeomorpha Giglioliana Wood-Mason, 1892: Plate 2, fig. 2. [Type locality not indicated] 

= Aristaeomorpha mediterranea Adensamer, 1898: 627; unnumbered text figure. [Pola stns 128, Nordküste 

von Afrika, 725 m; 192, Südkuste von Kreta (36°33'N 28°59'E, 1242 m); 204, Meer von Kandia 

(36°25'N 24°2'E, 808 m)] 

= Penaeus Meridionalis Hope, 1851: 19. [Nizza; nomen nudum] 

= Aristeus japonicus Yokoya, 1933: 3; Fig. 1. [Sôyô-Maru stns 383, E of Owase, Mie-ken, Japan, 353 m; 

399, near Owase, Mie-ken, 61 m] 

Aristaeomorpha woodmasoni Calman, 1925: 8. [Off Port Blair in the Andaman Sea, 271 fms] 

Aristaeopsis Wood-Mason in Wood-Mason & Alcock, 1891a 

= Aristæopsis Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus edwardsianus 

Johnson, 1868, by original designation, gender feminine) 

Aristaeopsis edwardsiana (Johnson, 1868) 

= Penaeus edwardsianus Johnson, 1868: 897. [off Madeira, Northeast Atlantic Ocean] 

= Aristeus coralinus Spence Bate, 1888: xxxii; fig. X. [Talisman expedition, exact locality not indicated] 

= Aristeus splendens Richard, 1900: 89. [Talisman expedition stn 1442, Azores; Talisman expedition stn 

1883, off Morocco; Talisman Expedition, off Cape Spartel, according to Davie, 2002: 119] 

Aristeus Duvernoy, 1840 

= Aristeus Duvernoy, 1840 (type species Penaeus antennatus Risso, 1816, by original designation, 

gender masculine) 

Aristeus alcocki Ramadan, 1938: 40; Figs 1, 2a, 3a. [John Murray Expedition, stns 34, Gulf of Aden, 1022 

m (13°05'36''N 46°24'42''E); 35, Gulf of Aden, 450-550 m (13°14'24''N 46°14'12''E to 13°13'24''N 

46°10'00''E); 176, Gulf of Aden, 650-730 m (12°04'06''N 50°38'36''E); 177, Gulf of Aden, 270-730 m 

(12°01'54''N 50°39'12''E); 193, Gulf of Aden, 1051 m (13°06'12''N 46°24'30''E to 13°03'00''N 46°21'42''E)] 

Aristeus antennatus (Risso, 1816) 

= Penaeus Antennatus Risso, 1816: 96; Plate 2, fig. 6. [Nice] 

= Sicyonia duvernoii Risso, 1844: 95. [Nice] [nomen nudum] 

= Penaeus antemarius Costes, 1890: 558. [nomen nudum] 

Aristeus antillensis A. Milne-Edwards & Bouvier, 1909: 201; Plate 1, figs 8-13. [Nevis Island, Antilles] 

Aristeus mabahissae Ramadan, 1938: 43; Figs 2b, 3b, 4a-c. [John Murray Expeditions, stns 143 (5°15'48''N 

73°22'48''E to 5°13'42''N 73°23'36''E), vicinity of the Maldives, 795 m; 145 (4°48'42"S 73°16'24"E), vicinity 

of the Maldives, 510 m] 

Aristeus pallidicauda Komai, 1993: 23; Figs 1-4. [Off Miyako, Japan, 600-700 m] 

Aristeus semidentatus Spence Bate, 1881: 189. [south of the Philippine Islands] 

? = Aristeus occidentalis Faxon, 1893: 215. [Albatross stns 3403 (off Galapagos Islands, 0°58'30"S 

89°17'00"W, 384 fms); 3410 (off Galapagos Islands, 0°19'00"N 90°34'00"W, 331 fms)]


Fig. 1. Aristaeomorpha foliacea (Risso, 1827). Photo by Tin-Yam Chan. 

Aristeus varidens Holthuis, 1952a: 71; Figs 17-18. [7°16'S 12°02'E, 53 M.W. Ambrizette, 440 m; 5°39'S 

11°25'E, 47 M.WbyS. Cabinda, 480 m; 10°45'S 13°10'E, 35 M.W. Cap Morro, 340 m; 10°45'S 13°17'E, 

40 M.W. Cap Morro, 400-500 m; 11°53'S 13°20'E, 28 M. WbyN. Egito, 500 m; 6°25'S 11°29'E, 50 M 

WSW. Moita Seca, ± 430 m] 

Aristeus virilis (Spence Bate, 1881) 

= Hemipenaeus virilis Spence Bate, 1881: 187. [near the Philippine Islands, 255 fms; according to 

Spence Bate, 1888: Challenger stn 200, near the Philippine Islands, 6°47'N 122°28'E, 250 fms] 

= Aristeus tomentosus Spence Bate, 1881: 189. [south of the Philippine Islands] 

= Aristeus viritli Burukovsky, 1974: 48. [Erroneous spelling] 

Austropenaeus Pérez Farfante & Kensley, 1997 

= Austropenaeus Pérez Farfante & Kensley, 1997 (type species Plesiopenaeus nitidus Barnard, 1947, by 

original designation, gender masculine) 

Austropenaeus nitidus (Barnard, 1947) 

= Plesiopenaeus nitidus Barnard, 1947: 383. [Off Cape Point, South Africa, 475-630 fms] 

= Aristeus crosnieri Burukovsky, 1975: 779; Figs 1-4. [SE Atlantic, 26°11'S 06°02'E, 1150 m] 

Hemipenaeus Spence Bate, 1881 

= Hemipenaeus Spence Bate, 1881 (type species Hemipenaeus spinidorsalis Spence Bate, 1881, designated 

by Faxon, 1895, gender masculine) 

Hemipenaeus carpenteri Wood-Mason & Alcock, 1891b 

= Hemipenæus Carpenteri Wood-Mason & Alcock, 1891b: 189. [Investigator stn 97 (Bay of Bengal, 

18°26'N 85°2'E), 1310 fms] 

= Hemipenaeus triton Faxon, 1893: 215. [Albatross stns 3360 (off Panama, 6°17'00"N 82°05'00"W, 1672 

fms); 3374 (off Panama, 2°35'00"N 83°53'00"W, 1823 fms); 3381 (4°56'00"N 80°52'30"W, 1772 fms)] 

Hemipenaeus spinidorsalis Spence Bate, 1881: 186. [South Atlantic, near the Island of Tristan d’Acunha, 

1900 fms; corresponding to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 35°41'S 

20°55'W, 1900 fms]


Hepomadus Spence Bate, 1881 

= Hepomadus Spence Bate, 1881 (type species Hepomadus glacialis Spence Bate, 1881, designated by 

Fowler, 1912, gender masculine) 

Hepomadus glacialis Spence Bate, 1881: 190. [mid South Atlantic, 1875 fms; corrected in Spence Bate, 1888 

to Challenger stn 237, near Yokohama, 34°37'N 140°32'E, 1875 fms] 

Hepomadus inermis Spence Bate, 1881: 190. [middle of South Pacific, 2550 fms; according to Spence Bate, 

1888 Challenger stn 289, South Pacific, 39°41'S 131°23'W, 2550 fms] 

Hepomadus tener Smith, 1884: 409; Plate 9, figs 7-8. [Albatross stn 2099, NW Atlantic, off eastern USA, 

37°12'20"N 69°30'W, 2949 fms] 

Parahepomadus Crosnier, 1978 

= Parahepomadus Crosnier, 1978 (type species Parahepomadus vaubani Crosnier, 1978, by original designation 

and monotypy, gender masculine) 

Parahepomadus vaubani Crosnier, 1978: 48; Figs 20-22. [Vauban CH 113, 22°18'S 43°59.7'E, 990-1010 m] 

Plesiopenaeus Spence Bate, 1881 

= Plesiopenaeus Spence Bate, 1881 (type species Aristeus armatus Spence Bate, 1881, designated by 

Faxon, 1895, gender masculine) 

Plesiopenaeus armatus (Spence Bate, 1881) 

= Aristeus armatus Spence Bate, 1881: 188. [among the islands of the Australasian archipelago, in the 

North Pacific, and South Atlantic, 1900-2050 fms; according to Spence Bate, 1888: Challenger stns 

133, South Atlantic Ocean, near Tristan da Cunha, 35°41'S 20°55'W, 1900 fms; 184, near Torres Strait, 

12°8'S 145°10'E, 1400 fms; 213, near the Philippines, 5°47'N 124°1'E, 2050 fms; 237, off Japan, 34°37'N 

140°32'E, 1875 fms; 246, Mid-Pacific, 36°10'N 178°00'E, 2050 fms; 276, the Low Archipelago, 13°28'S 

149°30'W, 2350 fms; 323, east of Buenos Ayres, 35°39'S 50°47'W, 1900 fms] 

Plesiopenaeus coruscans (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Aristæeus coruscans Wood-Mason in Wood-Mason & Alcock, 1891a: 280; Fig. 6. [Investigator stn 112 

(Bay of Bengal, 13°47'30"N 92°36'E), 561 fms] 

Pseudaristeus Crosnier, 1978 

= Pseudaristeus Crosnier, 1978 (type species Aristæus crassipes Wood-Mason in Wood-Mason & 

Alcock, 1891a, by original designation, gender masculine) 

Pseudaristeus crassipes (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Aristæus crassipes Wood-Mason in Wood-Mason & Alcock, 1891a: 281; Fig. 7. [Investigator stn 116, 

Andaman Sea, 11°25'05"N 92°47'06"E, 405 fms; lectotype designation by Pérez Farfante, 1987] 

Pseudaristeus gracilis (Spence Bate, 1888) 

= Hemipenaeus gracilis Spence Bate, 1888: 302; Plate 44, fig. 2. [Challenger stn 207, off Tablas Island, 

Philippines, 12°21'N 122°15'E, 700 fms] 

Pseudaristeus kathleenae Pérez Farfante, 1987: 314; Figs 1-3, 4C, 5-9. [Albatross stn 5657, Teluk Bone, 

Sulawesi (Celebes), Indonesia, 3°19'40"S 120°36'30"E, 900 m] 

Pseudaristeus protensus Pérez Farfante, 1987: 327; Figs 4D, 9, 14. [Investigator stn 370, W of Everal Gujarat, 

India (Arabian Sea), 19°51'30"N 69°07'30"E, 1569 m] 

Pseudaristeus sibogae (De Man, 1911a) 

= Hemipenaeus Sibogae De Man, 1911a: 6, 25. [Siboga stn 52, Savu Sea, Indonesia, 9°3.4'S 119°56.7'E, 

1000 m] 

Pseudaristeus speciosus (Spence Bate, 1881) 

= Hemipenaeus speciosus Spence Bate, 1881: 186. [the Atlantic off the coast of South America, 2650 

fms; according to Spence Bate, 1888: Challenger stn 325, east of Buenos Ayres, 36°44'S 46°16'W, 2650 

fms]


Family BeNTHeSICyMIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Altelatipes Crosnier & Vereshchaka, 2008 

= Altelatipes Crosnier & Vereshchaka, 2008 (type species Altelatipes falkenhaugae Crosnier & Vereshchaka, 

2008, by original designation and monotypy, gender masculine) 

Altelatipes brevirostris (Kikuchi & Nemoto, 1991) 

= Benthesicymus brevirostris Kikuchi & Nemoto, 1991: 75; figs 10-11. [23°08.4'N 150°04.8'E, 0-1300 m] 

Altelatipes carinatus (Smith, 1884) 

= Benthesicymus carinatus Smith, 1884: 396; Plate 10, figs 6-7. [Albatross stn 2094, off Long Island, NW 

Atlantic, 39°44'30"N 71°4'W, 1022 fms] 

= Benthesicymus expansus Kensley, 1977: 22; Figs 4-5. [Meiring Naude stn 107 (South Africa, off Natal, 

28°37.8'S 32°38.4'E, 1200-1000 m)] 

Altelatipes falkenhaugae Crosnier & Vereshchaka, 2008: 402; Figs 1-5, 6A-C, 7A-C. [ride médio-atlantique, 

53°04'N 37°17'W, 1478-665 m] 

Bentheogennema Burkenroad, 1936a 

= Bentheogennema Burkenroad, 1936a (type species Gennadas intermedius Spence Bate, 1888, by original 

designation, gender feminine) 

Bentheogennema borealis (Rathbun, 1902a) 

= Gennadas borealis Rathbun, 1902a: 887. [Albatross stn 3783, off Copper Island, Kamchatka, 1567 

fms] 

= Gennadas calmani Kemp, 1909: 724; Plate 74, figs 5-11; Plate 95, figs 4-5. [Challenger stns 232, S of 

Japan, 35°11'N 139°28'E, 345 fms; 236, S of Japan, 34°7'N 138°E, 565 fms; 237, near Yokohama, 

34°37'N 140°32'E, 1875 fms] 

Bentheogennema burkenroadi Krygier & Wasmer, 1975: 737; Figs 1-6. [Northeastern Pacific, 51°26'N 

138°28'W] 

Bentheogennema intermedia (Spence Bate, 1888) 

= Gennadas intermedius Spence Bate, 1888: 343; Plate 58, fig. 3. [Challenger stn 106, 1°47'N 24°26'W, 

1850 fms, off Sierra Leone; between Bermuda and Azores, surface; 137, 35°59'S, 1°34'E, surface] 

= Gennadas Alicei Bouvier, 1906a: 748. [Type locality not indicated; according to Bouvier, 1908a, the 

syntypic series came from Princesse-Alice stns 1306, a 300 milles à l’ouest de S.-Miguel, 4275 m; 1715, 

Canaries, 0-1000 m; 1749, entre Madère et les Canaries, 0-2500 m; 1768, Canaries, 0-3000; 1794, entre 

les Canaries et les Açores, 0-3000 m; 1797, entre les Canaries et les Açores, 0-3000 m; 2016, sud-ouest 

du cap Spartel, 0-1800 m; 2022, a l’est du banc Seine, 0-4000 m; 2113, Mer Sargasses, 0-1500 m; 2138, 

mer des Sargasses (région orientale), 0-2500 m; 2153, au sud des Açores, 0-2000 m; 2168, au sud de 

S.-Miguel, 0-2000 m; 2168, au sud de S.-Miguel, 0-2000 m; 2212, a l’ouest de Flores, 0-1200 m; 2244, 

au sud de S.-Miguel, 0-3000 m] 

Bentheogennema pasithea (De Man, 1907a) 

= Gennadas Pasithea De Man, 1907a: 146. [Siboga stn 230, 3°58'S 128°20'E, 0-2000 m] 

= Gennadas praecox Kemp, 1910: 176; Plate 13, figs 1-4. [Investigator stn 320, off C. Comorin, 7°23'N 

75°44'E, 1053 fms] 

= Gennadas caini Tirmizi, 1960: 368; Figs 40g, 48g, 84. [John Murray Expedition, stn 171 (9°07'06''N 

55°27'06''E to 9°08'48''N 55°31'48''E), central part of Arabian Sea, 3840-3872 m] 

Bentheogennema stephenseni Burkenroad, 1940: 37. [Dana stn 3624 I (SE of New Caledonia, 28°17.6'N 

177°01'E, 5000 m wire out)] 

Benthesicymus Spence Bate, 1881 

= Benthesicymus Spence Bate, 1881 (type species Benthesicymus crenatus Spence Bate, 1881, designated 

by Spence Bate, 1888, gender masculine) 

= Bentheocetes Smith, 1884 (type species Benthesicymus bartletti Smith, 1882; by monotypy; gender 

masculine) 

Benthesicymus altus Spence Bate, 1881: 191. [between Australia and Japan, 350-1400 fms]


Benthesicymus armatus MacGilchrist, 1905: 235. [Investigator stn 287 (Arabian Sea, 21°8'30''N 65°47'E), 

1506 fms] 

Benthesicymus bartletti Smith, 1882: 82; Plate 14, figs 1-7. [Blake stn 343, 39°45'40''N 70°55'W, 732 fms] 

= Benthesicymus pleocanthus Spence Bate, 1888: 334; Fig. 48; Plate 57, fig. 2. [Challenger stns 23, off 

Sombrero Island, 18°24'N 63°28'W, 450 fms; 205, Philippine Islands, 16°42'N 119°22'E, 1050 fms; 250, 

North Pacific Ocean, 37°49'N 166°47'W, 3050 fms] 

Benthesicymus brasiliensis Spence Bate, 1881: 191. [Atlantic and Pacific Oceans; according to Spence Bate, 

1888: Challenger stns 323, east of Buenos Ayres, 35°39'S 50°47'W, 1900 fms; 168, off New Zealand, 

40°29'S 177°43'E, 1100 fms; 173, off Matuku, Fiji Islands, 19°9'35"S 179°41'50"E, 315 fms; 181, between 

Australia and the Solomon Islands, 13°50'S 151°49'E, 2440 fms; 184, near Torres Strait, 12°8'S 

145°10'E, 1400 fms; 285, South Pacific, 32°36'S 137°43'W, 2375 fms] 

= Benthesicymus moratus Smith, 1886a: 90. [Albatross stns 2042 (39°33'00"N 68°26'45"W, 1555 fms); 

2174 (38°15'00"N 72°03'00"W, 1594 fms); 2222 (39°03'15"N 70°50'45"W, 1537 fms); 2575 (41°07'00"N 

65°26'30"W, 1710 fms)] 

Benthesicymus cereus Burkenroad, 1936a: 30; Figs 6, 11-12, 19, 24, 28, 35, 42, 47. [Pawnee stn 54, 21°15'40"N 

71°17'06"W, 900-945 fms] 

Benthesicymus crenatus Spence Bate, 1881: 190. [mid Pacific, 2600 fms; according to Spence Bate, 1888: 

Challenger stns 272, north of the Low Archipelago, 3°48'S 152°56'W, 2600 fms; 276, near the Low 

Archipelago, 13°28'S 149°30'W, 2350 fms] 

Benthesicymus investigatoris Alcock & Anderson, 1899: 282. [Investigator stns 222 (Andaman Sea, 13°27'N 

93°14'30''E), 400-200 fms (200-405 fms?); 228 (Andaman Sea, 13°7'N 94°44'15''E), 640 fms; 234 (Andaman 

Sea, 13°15'30''N 93°26'E), 498 fms; 235 (Andaman Sea, 14°13'N 93°40'E), 370-419 fms] 

Benthesicymus iridescens Spence Bate, 1881: 191. [near the island of Tristan d’Acunha, 1900 fms; according 

to Spence Bate, 1888: Challenger stn 133, near Tristan da Cunha, 35°41'S 20°55'W, 1900 fms] 

= Benthesicymus mollis Spence Bate, 1888: 339; Plate 58, fig. 2. [Challenger stn 133, near Tristan da 

Cunha, 35°41'S 20°55'W, 1900 fms] 

= Benthesicymus longipes Bouvier, 1906a: 747. [Princesse-Alice stn 1150, au sud-ouest des îles du Cap- 

Vert, 3890 m; according to Bouvier, 1908a] 

Fig. 2. Benthesicymus tanneri Faxon, 1893. Photo by E. del Solar.


Benthesicymus laciniatus Rathbun, 1906: 906; Fig. 59; Plate 19, fig. 3. [Albatross stn 4018, vicinity of Kauai 

Island, Hawaii, 804-724 fms] 

= Benthesicymus Hjorti Sund, 1920: 30; Fig. 48; Plate 11, fig. 4. [Michael Sars stn 35 (27°27'N 14°52'W), 

S of Canary Islands, 2603 m] 

= Gennadas pectinatus Schmitt, 1921: 25; Fig. 12; Plate 11, fig. 1. [Albatross stn 4390, off Santa Catalina 

Island, California, 1350-2182 m] 

Benthesicymus seymouri Tirmizi, 1960: 328; Figs 14-23. [John Murray Expedition, stn 118, 4°05'54"S 

41°10'12"E, 1789 m] 

Benthesicymus strabus Burkenroad, 1936a: 45; Figs 10, 23, 27, 34, 41, 46, 49. [Challenger stn 285, 32°36'S 

137°43'W] 

Benthesicymus tanneri Faxon, 1893: 215. [Albatross stns 3358 (off Panama, 6°30'00"N 81°44'00"W, 555 fms); 

3362 (off Panama, 5°56'00"N 85°10'30"W, 1175 fms); 3363 (off Panama, 5°42'00"N 85°50'00"W, 978 

fms); 3364 (off Panama, 5°30'00"N 86°08'30"W, 902 fms); 3365 (off Panama, 5°31'00"N 86°31'00"W, 

1010 fms); 3366 (off Panama, 5°30'00"N 86°45'00"W, 1067 fms); 3376 (off Panama, 3°09'00"N 

82°08'00"W, 1132 fms); 3377 (3°56'00"N 81°40'15"W, 764 fms); 3380 (off Panama, 4°03'00"N 

81°31'00"W, 899 fms); 3384 (off Panama, 7°31'30"N 79°14'00"W, 458 fms); 3393 (off Panama, 

07°15'00"N 79°36'00"W, 1020 fms); 3400 (off Galapagos Islands, 0°36'00"S 86°46'00"W, 1322 fms); 

3403 (off Galapagos Islands, 0°58'30"S 89°17'00"W, 383 fms); 3407 (off Galapagos Islands, 0°04'00"S 

90°24'30"W, 885 fms); 3410 (off Galapagos Islands, 0°19'00"N 90°24'00"W, 331 fms); 3411 (off Galapagos 

Islands, 0°54'00"N 91°09'00"W, 1189 fms); 3418 (off Mexico, 16°33'00"N 99°52'30"W, 660 fms); 

3424 (off Mexico, 21°15'00"N 106°23'00"W, 676 fms); 3425 (off Mexico, 21°19'00"N 106°24'00"W, 680 

fms); 3435 (Gulf of California, 26°48'00"N 110°45'20"W, 859 fms); 3436 (27°03'40"N 110°53'40"W, 905 

fms)] (Fig. 2) 

Benthesicymus tirmiziae Crosnier, 1978: 16; Figs 7a-b, 8a-b, 11a. [Vauban stn CH 132, Madagascar, 13°43.8'S 

47°29.0'E, 1950-2150 m] 

Benthesicymus howensis Dall, 2001 

= Benthesicymus urinator howensis Dall, 2001: 428; Figs 13A-D. [Lord Howe Rise, Tasman Sea, 28°44'S 

161°54'E, 1325 m] 

Benthesicymus urinator Burkenroad, 1936a: 29; Figs 4, 5, 8-9, 17-18, 22, 32, 33, 39-40, 45. [Challenger stn 184, 

near Torres Strait, Australia, 12°8'S 145°10'E, 1400 fms] 

Benthonectes Smith, 1885a 

= Benthonectes Smith, 1885a (type species Benthonectes filipes Smith, 1885a, by monotypy, gender 


= Hapalopoda Filhol, 1885a (type species Hapalopoda investigator Filhol, 1885a, by monotypy, gender 

feminine) 

Benthonectes filipes Smith, 1885a: 509. [Albatross stns 2181, 39°29'00''N 71°46'00''W, 603 fms; 2206, 

39°35'00''N 71°24'30''W, 1043 fms; 2210, 39°37'45''N 71°18'45''W, 991 fms; 2235, 39°12'00''N 

72°03'30''W, 707 fms] 

= Hapalopoda investigator Filhol, 1885a: 229; Fig. 2. [Crevette prise à 1900 mètres de profondeur, à 

bord du Talisman (33°9'N 11°58'W (of Paris) (= 9°38'W of Greenwich))] 

Gennadas Spence Bate, 1881 

= Gennadas Spence Bate, 1881 (type species Gennadas parvus Spence Bate, 1881, by monotypy, gender 


= Amalopenaeus Smith, 1882 (type species Amalopenaeus elegans Smith, 1882, by monotypy, gender 


= Pasiphodes Filhol, 1885b (type species, Pasiphodes purpureus Filhol, 1885b, by monotypy, gender 


Gennadas barbari Vereshchaka, 1990: 131; Fig. 1. [stn 1925, 25°39'S 85°27'W] 

Gennadas bouvieri Kemp, 1909: 726; Plate 74, figs 1-4; Plate 75, figs 6-7. [Challenger stns 206, W of Manila, 

17°54'N 117°14'E, 2100 fms; 220, N of New Guinea, 0°42'S 147°E, 1100 fms]


= Gennadas alcocki Kemp, 1910: 174 (?partim, males only); Plate 13, figs 5-6. [Investigator stns 111, Bay 

of Bengal, 12°50'N 90°52'E, 1644 fms; 103, Bay of Bengal, 15°14'N 81°9'E, 1260 fms; 108, off C. Comorin, 

7°4'N 76°34'15"E, 1043 fms; 309, near the Andaman Islands, 10°9'N 93°2'15"E, 765 fms; see 

Crosnier, 1978 for discussion of this problem] 

Gennadas brevirostris Bouvier, 1905a: 748. [provient des parages de Sainte-Lucie où elle fut trouvée par le 

Blake sur des fonds de 221 brasses] 

= Gennadas similis Stephensen, 1923: 6, 12; Fig. 1. [Atlantic, 39°35'N 9°45'W, 1150 m] 

= Gennadas chiasmifera Stephensen, 1923: 6, 13; Figs 2-3. [Atlantic, 39°35'N 9°45'W, 1150 m] 

Gennadas capensis Calman, 1925: 5; Plate 1, Figs 1-2. [SS Pickle stn 87, off Cape, South Africa, 1014 fms] 

Gennadas crassus Tirmizi, 1960: 346, 371; Figs 40h, 85. [John Murray Expedition, stn 121 (5°39'00''N 

39°38'30''E to 5°40'30''N 39°43'00''E), Zanzibar area, bottom at 925 m, net apparently not on bottom] 

Gennadas elegans (Smith, 1882) 

= Amalopenaeus elegans Smith, 1882: 87; Plate 14, figs 8-14; Plate 15,figs 1-5. [Blake stns 323, 33°19'0"N 

76°12'30"W, 457 fms; 324, 33°27'20"N 75°53'30"W, 1386 fms; 325, 33°35'20"N 76°0'0"W, 647 fms; 328, 

34°28'25"N 75°22'50"W, 1632 fms; 330, 31°41'0"N 74°35'0"W, 1047 fms; 343, 39°45'40"N 70°55'0"W, 

732 fms; 893, 39°52'20"N 70°58'0"W, 372 fms; 935, 39°45'0"N 69°44'45"W, 770 fms; 952, 39°55'0"N 

70°28'0"W, 388 fms] 

Gennadas gilchristi Calman, 1925: 6; Plate 1, figs 3-4. [SS Pickle stns 77 (755 fms), 84 (500 fms), 87 (1014 

fms), 357 (900 fms), off Cape, South Africa] 

Gennadas incertus (Balss, 1927) 

= Amalopenaeus incertus Balss, 1927: 265; figs 24-29. [Valdivia stns 232, 3°26'S 58°34'E, 0-1500 m; 235, 

4°34'S 53°42'E, 0-2000 m; 237, 4°45'S 48°58'E, 0-2000 m] 

= Amalopenaeus Gardineri Balss, 1927: 267; Fig. 31. [Valdivia stn 236, 4°38'S 51°16'E, 0-2000 m; 4 miles 

N.W. of Desroches Atoll, 750-0 fms] 

Gennadas kempi Stebbing, 1914a: 283; Plate 27. [Scotia stn 468, Cape Point, South Africa, 39°48'S 2°33'E, 

2772 fms] 

Gennadas parvus Spence Bate, 1881: 192. [off Japan, 2425 fms] 

Gennadas propinquus Rathbun, 1906: 907; Fig. 61a-b. [Between Erben Bank and Kaiwi Channel, Hawai’ian 

Islands] 

= Gennadas clavicarpus De Man, 1907a: 144 (partim). [Siboga stns 128, 4°27'N 125°25.7'E, 0-700 m; 141, 

1°0.4'S 127°25.3'E, 0-1500 m; 230, 3°58'S 128°20'E, 0-2000 m; see Burkenroad, 1936a] 

= Gennadas scutatus indicus Kemp, 1913a: 62. [Investigator stns 108, off C. Comorin, 7°4'N 76°34'15"E, 

1043 fms; 109, off C. Comorin, 7°1'N 78°21'E, 738 fms] 

Gennadas scutatus Bouvier, 1906a: 748. [l’Hirondelle stn 156, entre les Açores et Terre-Neuve, surface according 

to Bouvier, 1908a] 

= Gennadas scutatus Bouvier, 1906b: 690. [nomen nudum] 

Gennadas sordidus Kemp, 1910: 177; Plate 14, figs 1-3. [Investigator stns 193, N of the Laccadive Islands, 

15°11'N 72°28'45"E, 931 fms; 194, off the Laccadive Islands, 13°47'N 72°3'45"E, 891 fms; 198, NE of 

Ceylon, 8°55'N 81°17'30"E, 764 fms] 

Gennadas talismani Bouvier, 1906a: 749. [Talisman stn 105, 16°38'N 18°24'W, 3200 m, according to Crosnier 

& Forest, 1973] 

Gennadas tinayrei Bouvier, 1906c: 10; Figs 2-4, 14. [Mer des Sargasses, Açores, Cap Spartel (Princesse-Alice 

stn 2264, 37°30'N 22°39'W, 0-3000 m, according to Davie, 2002: 127)] 

Gennadas valens (Smith, 1884) 

= Amalopenaeus valens Smith, 1884: 402; Plate 10, fig. 2. [Albatross stn 2003, 37°16'30"N 74°20'36"W, 

640 fms] 

= Pasiphodes purpureus Filhol, 1885b: Plate 3. [Type locality not indicated; Holthuis, 1980a refers to 

A. Milne-Edwards’ original colour sketch of the species in the Paris Museum, probably from Talisman 

stn 38, off Cap Ghir, Morocco, 2210 m] 

= Gennadas bidentata Stephensen, 1923: 6, 14; Figs 4, 5. [Atlantic, 57°47'N 11°33'W, 1985 m]


Family PeNAeIDAe Rafinesque, 1815 

Alcockpenaeopsis Sakai & Shinomiya, 2011 

= Alcockpenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis hungerfordi Alcock, 1905, by 

original designation, gender feminine) 

Alcockpenaeopsis hungerfordii (Alcock, 1905) 

= Parapenaeopsis hungerfordii Alcock, 1905: 522, 530. [Hongkong] 

Arafurapenaeopsis Sakai & Shinomiya, 2011 

= Arafurapenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis arafurica Racek & Dall, 

1965, by original designation, gender feminine) 

Arafurapenaeopsis arafurica (Racek & Dall, 1965) 

= Parapenaeopsis arafurica Racek & Dall, 1965: 102; Fig. 16; Plate 8, figs 6-7; Plate 13, fig. 5. [off the Fly 

River, Papua New Guinea] 

Artemesia Spence Bate, 1888 

= Artemesia Spence Bate, 1888 (type species Artemesia longinaris Bate, 1888, by monotypy, gender 

feminine) 

Artemesia longinaris Spence Bate, 1888: 281; Plate 90. [Challenger stn 321, off Montevideo, Uruguay, 

35°02'S 55°15'W, 13 fms] 

= Artemesia brevinaris Nobili, 1901a: 1; Figs 1-2. [Mar del Plata, Argentina] 

Atypopenaeus Alcock, 1905 

= Atypopenaeus Alcock, 1905 (type species Penaeus compressipes Henderson, 1893, by original designation, 


= Miyadiella Kubo, 1949a (type species Miyadella pedunculata Kubo, 1949a, by original designation, 


Atypopenaeus bicornis Racek & Dall, 1965: 85; Fig. 14; Plate 7, figs 5-6; Plate 12, fig. 6. [Off Fly River mouth, 

Papua New Guinea] 

Atypopenaeus compressipes (Henderson, 1893) 

= Penaeus compressipes Henderson, 1893: 450. [Gulf of Martaban, India] 

= Parapenaeopsis brevirostris Kubo, 1936a: 55; Plate 61. [Inland Sea of Japan] 

Atypopenaeus dearmatus De Man, 1907a: 135. [Siboga stns 302, 10°17.9'S 123°28.7'E, 216 m; 306, 8°27'S 

122°54.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

= Miyadiella ornata Holthuis, 1955a: 76; Figs 1-2. [Off NE Java, 7°33'S 114°36'E, 200 m] 

Atypopenaeus formosus Dall, 1957: 199; Fig. 21A-H. [5 km E off Sandgate, Moreton Bay, Queensland, 

Australia, 2-3 fms] 

Atypopenaeus stenodactylus (Stimpson, 1860a) 

= Penaeus stenodactylus Stimpson, 1860a: 43. [Portu ”Hong Kong”] 

= Penaeus podophthalmus Stimpson, 1860a: 43. [Portu ”Hong Kong”] 

= Miyadella pedunculata Kubo, 1949a: 264; Figs 7N, 23G, H, 58O, 74C, I, 79E, 104, 105. [Osaka-wan, 

Japan] 

Batepenaeopsis Sakai & Shinomiya, 2011 

= Batepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus tenellus Spence Bate, 1888, by original 


Batepenaeopsis tenella (Spence Bate, 1888) 

= Penaeus tenellus Spence Bate, 1888: 270. [Challenger stn 235, Bay of Kobe, Japan, 34°39'N 135°14'E, 

8 fms] 

= Penaeus crucifer Ortmann, 1890: 451; Plate 56, fig. 5. [Maizuru, Japan] 

Batepenaeopsis venusta (De Man, 1907a) 

= Parapeneopsis venusta De Man, 1907a: 134. [Siboga expedition, station off Pulu Jedan, E coast of Aru 

Islands, shallow water]


Farfantepenaeus Burukovsky, 1997 

= Penaeus (Farfantepenaeus) Burukovsky, 1997 (type species Penaeus [as Pénée] brasiliensis Latreille, 

1817, by original designation, gender masculine) 

= Penaeus (Farfantepenaeus) Burukovsky, 1972 [no type species indicated] 

Farfantepenaeus aztecus (Ives, 1891) 

= Penaeus Brasiliensis var. Aztecus Ives, 1891: 190. [Veracruz, Mexico; lectotype designated by Pérez 

Farfante, 1969] 

Farfantepenaeus brasiliensis (Latreille, 1817) 

= Penaeus [as Pénée] brasiliensis Latreille, 1817: 156. [côtes du Brésil] 

Farfantepenaeus brevirostris (Kingsley, 1878a) 

= Penaeus brevirostris Kingsley, 1878a: 98. [Estero at Realijo, W. coast of Nicaragua] 

Farfantepenaeus californiensis (Holmes, 1900) 

= Penaeus californiensis Holmes, 1900: 218. [Santa Monica, California; neotype designation by Burkenroad, 

1938] 

Farfantepenaeus duorarum (Burkenroad, 1939) 

= Penaeus duorarum Burkenroad, 1939: 31; Figs 18-19, 23, 25-27. [Atlantis stn 2813, off Alabama, Gulf 

of Mexico, 19 fms] 

Farfantepenaeus notialis (Pérez Farfante, 1967) 

= Penaeus duorarum notialis Pérez Farfante, 1967: 94; Figs 4a-d. [Oregon stn 5664, off Las Piedras, Gulf 

of Venezuela, 11°44'N 70°22'W, 26 fms] 

= Penaeus duorarum var. cameronensis Rossignol, 1962 [unavailable name under Art. 15.2] 

Farfantepenaeus paulensis (Pérez Farfante, 1967) 

= Penaeus (Melicertus) paulensis Pérez Farfante, 1967: 84; Figs 1a-d. [Santos, Sāo Paulo, Brazil] 

Farfantepenaeus subtilis (Pérez Farfante, 1967) 

= Penaeus aztecus subtilis Pérez Farfante, 1967: 89; Figs 2a-b, 3a-c. [Oregon stn 5685, off Gallinas Point, 

Departemento de la Guajira, Colombia, 12°29'N 71°54'W, 95 fms] 

Fenneropenaeus Pérez Farfante, 1969 

= Fenneropenaeus Pérez Farfante, 1969 (type species Penaeus indicus H. Milne Edwards, 1837 [in H. 

Milne Edwards, 1834-1840], by original designation, gender masculine) 

Fenneropenaeus chinensis (Osbeck, 1765) 

= Cancer chinensis Osbeck, 1765: 151. [Bocca Tiger, auf Chinesisch Pho-hao, der Tigermund, oder Phomunn, 

die Tigersöfnung. Fischer kamen mit verschiedenen Fisharten zu uns, .... und chinesischen 

Krabben: Cancer chinensis] 

= Penaeus orientalis Kishinouye, 1917: 79. [Chosen, Kantoshu and Chintao] 

Fenneropenaeus indicus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

? = Palæmon longicornis Olivier, 1811: 662. [Type locality not indicated] 

= Penaeus indicus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 415. [les côtes de Coromandel] 

= Penaeus indicus longirostris De Man, 1892a: 511; Plate 29, fig. 53. [Celebes, Makassar] 

Fenneropenaeus konkani Chanda & Bhattacharya, 2003: 229; Fig. 1. [Mirkarwada Fish landing centre, Ratangiri, 

Maharstra, West Coast of India] 

Fenneropenaeus merguiensis (De Man, 1888a) 

= Penaeus merguiensis De Man, 1888a: 287; Plate 18, fig. 8; Plate 19, fig. 1. [Burma, Mergui Archipelago] 

Fenneropenaeus penicillatus (Alcock, 1905) 

= Penaeus indicus penicillatus Alcock, 1905: 515, 525. [Off the Orissa coast; at Bombay; Karáchi; 

Gangetic Delta; Mergui] 

Fenneropenaeus silasi (Muthu & Motoh, 1979a) 

=Penaeus silasi Muthu & Motoh, 1979a: 64; Figs 1-2. [Tawau, Sabah, 36 m] 

Funchalia Johnson, 1868 

= Funchalia Johnson, 1868 (type species Funchalia woodwardi Johnson, 1868, by monotypy, gender 

feminine)


= Grimaldiella Bouvier, 1905b (type species Grimaldiella richardi Bouvier, 1905b, by monotypy, gender 

feminine) 

= Hemipenaeopsis Bouvier, 1905b (type species Hemipenaeopsis villosus Bouvier, 1905b, by monotypy, 


Funchalia danae Burkenroad, 1940: 36. [Dana stn 4017 VII (Canary Islands, 29°13'N 14°12'W, 1000 m wire 

out)] 

Funchalia sagamiensis Fujino, 1975: 200; Figs 1-3. [Amadai-ba, Sagami Bay, central Japan, upper water 

layer] 

Funchalia taaningi Burkenroad, 1940: 36. [Dana stn 3920 III (N of Seychelles, 1°06'N 62°25'E, 3000 m wire 

out)] 

Funchalia villosa (Bouvier, 1905b) 

= Hemipenaeopsis villosus Bouvier, 1905b: 981 (pro parte). [capturée par la Princesse-Alice entre les îles 

Canaries et les Açores] 

= Funchalia vanhöffeni Lenz & Strunck, 1914: 306; Plate 17, figs 20-21. [nördlich von Tristan da Cunha] 

Funchalia woodwardi Johnson, 1868: 895. [Off Madeira, NE Atlantic Ocean] 

= Peneus Genianus Risso, 1841 [nomen nudum] 

= Sycionia geny Risso, 1844: 95. [Nice] [nomen nudum] 

= Sicyonia Genyana Hope, 1851: 19. [nomen nudum] 

= Grimaldiella Richardi Bouvier, 1905b: 981, 982. [capturée par la Princesse-Alice dans se dernière 

campagne, au large de l’île Saint-Michel, en plein groupe des Açores] 

= Peneus genyanus Monod, 1931: 107. [nomen nudum] 

Ganjampenaeopsis Sakai & Shinomiya, 2011 

= Ganjampenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis uncta Alcock, 1905, by 


Ganjampenaeopsis uncta (Alcock, 1905) 

= Parapenaeopsis uncta Alcock, 1905: 522, 528. [Ganjam coast, 10-11 fms] 

= Parapenaeopsis probata Hall, 1961: 96; Plate 19, figs 11-13. [South of Bedok, 24 fms; South of Singapore, 

24 and 45 fms] 

Heteropenaeus De Man, 1896 

= Heteropenaeus De Man, 1896 (type species Heteropenaeus longimanus De Man, 1896, by monotypy, 


Heteropenaeus longimanus De Man, 1896: 111. [Java-See] 

Holthuispenaeopsis Sakai & Shinomiya, 2011 

= Holthuispenaeopsis Sakai & Shinomiya, 2011 (type species Parapenaeopsis atlantica Balss, 1914a, by 


Holthuispenaeopsis atlantica (Balss, 1914a) 

= Parapenaeopsis atlantica Balss, 1914a: 593. [Victoria, Kamerun] 

= Trachypenaeus constrictus var. africana Balss, 1916: 17; Figs 3-4. [Goldküste: Kap Coast Castle, 13 m; 

Addah, 11 m; Acra, 9 m. Togo, Lome, 13 m. Dahomey, Whyda, Cabinda, Landana, 7 m. Französisch- 

Kongo: Nyanga-Fluß, 13 m; Loango, 10 m. Angola: Ambriz, 13 m; Kinsembo, 13 m] 

Kishinouyepenaeopsis Sakai & Shinomiya, 2011 

= Kishinouyepenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus cornutus Kishinouye, 1900, by 


Kishinouyepenaeopsis amicus (V.C. Nguyên, 1971) 

= Parapenaeopsis amicus V.C. Nguyên, 1971: 46; Fig. 1. [West Tonkin Gulf] 

= Parapenaeopsis sinica Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South 

China Sea]


Kishinouyepenaeopsis cornuta (Kishinouye, 1900) 

= Penaeus cornutus Kishinouye, 1900: 23; unnumbered text fig. [Bay of Ariake, Kushu] 

Kishinouyepenaeopsis incisa (Liu & Wang, 1986) 

= Parapenaeopsis incisa Liu & Wang, 1986: 214, 215. [Wailuo, Xuwen County, Guangdong, South 

China Sea] 

Kishinouyepenaeopsis maxillipedo (Alcock, 1905) 

= Parapenaeopsis maxillipedo Alcock, 1905: 522, 527. [Bombay, Madras, Arakan coast, India] 

Litopenaeus Pérez Farfante, 1969 

= Litopenaeus Pérez Farfante, 1969 (type species Penaeus vannamei Boone, 1931, by original designation, 


Litopenaeus occidentalis (Streets, 1871a) 

= Penaeus occidentalis Streets, 1871a: 243. [Isthmus of Panama] 

Litopenaeus schmitti (Burkenroad, 1936b) 

= Penaeus schmitti Burkenroad, 1936b: 315; Figs 1-3. [Kingston Bay, Jamaica] 

Litopenaeus setiferus (Linnaeus, 1767) 

= Cancer setiferus Linnaeus, 1767: 1054. [Indiis] 

= Pæneus orbignyanus Latreille, 1817: 155. [côtes maritimes du département de la Vendée] 

= P.[enaeus] fluviatilis Say, 1818: 236. [North America] 

= Penaeus gracilirostris Thallwitz, 1892: 3; Plate 1, fig. 5. [Nord-Celébes] 

Litopenaeus stylirostris (Stimpson, 1871) 

= Penaeus stylirostris Stimpson, 1871: 134. [Panama] 

Litopenaeus vannamei (Boone, 1931) 

= Penaeus vannamei Boone, 1931: 173; Fig. 16. [Golfo de Panama] 

Macropetasma Stebbing, 1914b 

= Macropetasma Stebbing, 1914b (type species Parapenaeus africanus Balss, 1913a, by monotypy, gender 

neuter) 

Macropetasma africana (Balss, 1913a) 

= Parapenaeus africanus Balss, 1913a: 105. [Swakopmund, Namibia] 

Marsupenaeus Tirmizi, 1971a 

= Marsupenaeus Tirmizi, 1971a (type species Penaeus canaliculatus var. japonicus Spence Bate, 1888, by 

monotypy, gender masculine) 

Marsupenaeus japonicus (Spence Bate, 1888) 

= Penaeus canaliculatus japonicus Spence Bate, 1888: 245; Plates 31; 32, fig. 4; 37, fig. 2. [Japan] 

= Penaeus pulchricaudatus Stebbing, 1914b: 14; Plate 67. [Great Fish Point Lighthouse, Cape province, 

South Africa] 

Megokris Pérez Farfante & Kensley, 1997 

= Megokris Pérez Farfante & Kensley, 1997 (type species Penaeus granulosus Haswell, 1879, by original 

designation, gender masculine) 

Megokris akademik Shinomiya & Sakai, 2006a: 1220; Figs 1-2. [R.V. ”Akademik” stn PG-13-Ku, Persian 

Gulf, 29°13.587'N 49°53.955'E, 41 m] 

Megokris ghamrawyi Shinomiya & Sakai, 2006a: 1223; Figs 3-6. [R.V. ”Akademik” stn PG-13-Ku, Persian 

Gulf, 29°13.587'N 49°53.955'E, 41 m] 

Megokris gonospinifer (Racek & Dall, 1965) 

= Trachypenaeus gonospinifer Racek & Dall, 1965: 89; Fig. 15; Plate 7, figs 7-8; Plate 12, fig. 7. [S. of Port 

Romilly, Papua New Guinea, 7°55'S 144°48'E, 7 fms] 

Megokris granulosus (Haswell, 1879) 

= Penaeus granulosus Haswell, 1879: 41. [Darney Island; lectotype designation by Griffin, 1970] 

= Trachypeneus salaco De Man, 1907a: 135. [Siboga stns 205, Lohio bay, Buton Strait, Indonesia, 22 m; 

258, Tual Anchorage, Kai Islands, 22 m] 

= Trachypeneus furcilla Hall, 1961: 102; Plate 20, figs 16-17. [Outer Shoal, Malaysia, 6 fms]


Megokris halli Shinomiya & Sakai, 2006b: 1255; Figs 1-2. [Singapore Fisheries Research stn B 146, Permanent 

Buoy 10, Angler Buoy, 9 m] 

Megokris manihine Shinomiya & Sakai, 2006b: 1259; Figs 3-4. [Singapore Fisheries Research stn c5/15, 

5°12'N 114°59'E, 37 m] 

Megokris motohburiorum Shinomiya & Sakai, 2006b: 1261; Fig. 5. [Iloilo, Philippines] 

Megokris pescadoreensis (Schmitt, 1931a) 

= Trachypeneus pescadoreensis Schmitt, 1931a: 265; Plate 32, figs 2-4. [Pescadores Islands, Formosa] 

Megokris sedili (Hall, 1961) 

= Trachypeneus sedili Hall, 1961: 100; Plate 20, figs 18-19. [Malaysia, S of Bedok, 24 fms] 

Melicertus Rafinesque, 1814 

= Melicertus Rafinesque, 1814 (type species Melicertus tigrinus Rafinesque, 1814, by monotypy, gender 


Melicertus canaliculatus (Olivier, 1811) 

= Palæmon canaliculatus Olivier, 1811: 660. [Indonesia, N Sumatra, Pulu Weh; neotype designated 

by Pérez Farfante, 1976] 

Melicertus hathor (Burkenroad, 1959) 

= Penaeus latisulcatus hathor Burkenroad, 1959: 80; Figs 7, 8. [Mersa Thlemel, Gulf of Suez] 

Melicertus kerathurus (Forskål, 1775) 

= Cancer kerathurus Forskål, 1775: 95. [Smirnæ & Alexandriæ] 

= Palæmon sulcatus Olivier, 1811: 661. [Elle se trouve aux environs de Smyrne & d’Alexandrie] 

= Alpheus Trisulcatus Leach, 1814 [in Leach, 1813-1814]: 431. [Anglesea] 

= Melicertus tigrinus Rafinesque, 1814: 22. [Sicily] 

= Alpheus caramote Risso, 1816: 90. [Nice] 

= Peneus mars Risso, 1816: 47; Plate 2, fig. 5. [Nice, se tient à une grande profondeur] 

= Alpheus punctulatus Risso, 1822: 247. [mer de Nice, régions sablonneuses] 

= Peneus cristatus Risso, 1827: 67. [environs de Nice et des Alpes Maritimes; grandes profondeurs] 

= Penaeus fasciatus Hope, 1851: 19. [nomen nudum] 

Melicertus latisulcatus (Kishinouye, 1896) 

= Penaeus latisulcatus Kishinouye, 1896: 372. [Kagoshima, Tokyo Bay, Japan] 

Melicertus longistylus (Kubo, 1943) 

= Penaeus longistylus Kubo, 1943: 200. [off South-East coast of Hainan Island] 

= Penaeus jejunus Hall, 1956: 75; Plate 10, figs 9-10. [probably from the neighbourhood of Singapore] 

= Penaeus caesius Dall, 1957: 143; Fig. 2A-G. [12 miles NNE of Bowen, Queensland, Australia, 19-25 fms] 

Melicertus marginatus (Randall, 1840) (Fig. 3) 

= P.[enœus] marginatus Randall, 1840: 146. [Sandwich Islands] 

= Penaeus teraoi Kubo, 1949a: 288; Figs 7F, 20F, 24N-P, 49K, 58G, 73D, J, 77S, 112. [Miya, Aiti Prefecture, 

Japan] 

Melicertus plebejus (Hess, 1865) 

= Penaeus plebejus Hess, 1865: 168; Plate 7, fig. 19. [Sydney] 

= Penaeus canaliculatus var. australiensis Spence Bate, 1888: 248; Plate 32, fig. 3. [Port Jackson, Australia, 

2-10 fms] 

= Penaeus maccullochi Schmitt, 1926a: 359, 370; Plate 66, figs 1-3. [21 km NE of North Reef, Queensland, 

Australia] 

Melicertus similis Chanda & Bhattacharya, 2002: 495; Figs 1-6. [Port Blair, Andaman Is., Bay of Bengal] 

Metapenaeopsis Bouvier, 1905b 

= Metapenaeopsis Bouvier, 1905b (type species Metapenaeopsis pubescens Bouvier, 1905b, by monotypy, 


= Archipenaeopsis Bouvier, 1905b (type species Archipenaeopsis vesitus Bouvier, 1905b, by monotypy, 


= Leptopenaeus Kishinouye, 1929 (type species Penaeus philippii Spence Bate, 1888, designated by 

Pérez Farfante & Kensley, 1997, gender masculine)


Fig. 3. Melicertus marginatus (Randall, 1840). Photo by Tin-Yam Chan. 

= Ceratopenaeus Kishinouye, 1929 (type species Parapenaeus mogiensis Rathbun, 1902b, designated by 

Pérez Farfante & Kensley, 1997, gender masculine) 

= Erythropenaeus Kishinouye, 1929 (type species Parapenaeus akayebi Rathbun, 1902b, designated by 

Pérez Farfante & Kensley, 1997, gender masculine) 

Metapenaeopsis acclivis (Rathbun, 1902b) 

= Parapenaeus acclivis Rathbun, 1902b: 41; Figs 12-14. [Mogi, Japan] 

Metapenaeopsis aegyptia Galil & Golani, 1990: 229; Figs 1a-b, 2a-c, 3a. [off Palmahim, Israel, 31°56'N 

34°35'E, 50 m] 

= Metapenaeopsis judaensis Por, 1989: 211. [nomen nudum] 

Metapenaeopsis andamanensis (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Metapenaeus philippinensis var. andamanensis Wood-Mason in Wood-Mason & Alcock, 1891a: 271. 

[N of Port Blair, Andaman Sea, 112-244 fms] 

Metapenaeopsis angusta Crosnier, 1987a: 441; Figs 16b, 18b, 19c-d. [MUSORSTOM I, stn 5, 14°01.5'N 

120°23.5'E, 215-200 m] 

Metapenaeopsis assimilis (De Man, 1920a) 

= Penaeopsis assimilis De Man, 1920a: 105. [Off Pulu Weh, N Sumatra, Indonesia; lectotype designation 

by Crosnier, 1991] 

= Metapenaeopsis raceki Starobogatov, 1972: 405, 409; Plate 10, fig. 127a-b. [nomen novum for Metapenaeopsis 

distincta sensu Racek & Dall, 1965 nec De Man, 1907a] 

Metapenaeopsis barbata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penaeus barbatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 3. [Japan] 

= Parapenaeus akayebi Rathbun, 1902b: 39. [Mogi, Japan] 

Metapenaeopsis batei (Miers, 1884b) 

= Penæus batei Miers, 1884b: 296; Plate 32, fig. D. [Albany Island, 3-4 fms] 

Metapenaeopsis beebei (Burkenroad, 1938) 

= Penaeopsis (Metapenaeopsis) beebei Burkenroad, 1938: 74; Figs 18-19. [Templeton Crocker Expedition 

stn 136, dredge 30, Arena Bank, Gulf Of California, 23°27'N 109°24'W, 35 fms]


Metapenaeopsis ceylonica Starobogatov, 1972: 406, 408 (key); Plate 11, fig. 159a-b; nomen novum for M. 

hilarulus sensu De Bruin, 1965 nec De Man, 1911a. [Ceylon, W coast, W of Mannar, 6 fms] 

Metapenaeopsis commensalis Borradaile, 1899: 1001, Plate 63, fig. 1-1b. [Ellice Islands, Rotuma, Fiji] 

= Penaeopsis Borradailei De Man, 1911a: 73. [Siboga stn 131, anchorage off Beo, Karakelong Island, 

Talaud Archipelago, Indonesia; lectotype designation by Crosnier, 1991] 

Metapenaeopsis coniger (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Metapenaeus coniger Wood-Mason in Wood-Mason & Alcock, 1891a: 272. [Investigator stns 119 (Bay 

of Bengal, 12°20'N 85°8'E), 95 fms; off Mahánaddi Delta, 68 fms; 96 (Bay of Bengal, 18°30'N 84°46'E), 

98-102 fms] 

Metapenaeopsis costata Crosnier, 1991: 275; Figs 83, 84. [MUSORSTOM 3, stn CP 142, Philippines, 11°47'N 

123°01.5'E, 26 m] 

Metapenaeopsis crassissima Racek & Dall, 1965: 26; Fig. 2C; Plate 2, fig. 1; Plate 4, figs 5-6; Plate 9, fig. 3. 

[Shark Bay, Western Australia] 

Metapenaeopsis dalei (Rathbun, 1902b) 

= Parapenaeus dalei Rathbun, 1902b: 40; Figs 9-11. [Mogi, Japan] 

? = Metapenaeopsis incomptus Kubo, 1949a: 424; Figs 8D, 46A, 76L, R, 80N, 148F. [East China Sea] 

Metapenaeopsis difficilis Crosnier, 1991: 255; Figs 65-67. [MUSORSTOM 3, stn CP 134, Philippines, 

12°01.1'N 121°57.3'E, 92-95 m] 

Metapenaeopsis distincta (De Man, 1907a) 

= Metapeneus distinctus De Man, 1907a: 132. [Siboga stn 184, Anchorage off kampong Kelang, south 

coast of Manipa-island, 36 m; female specimen illustrated by De Man, 1913; designated as lectotype 

by Crosnier, 1991] 

Metapenaeopsis dura Kubo, 1949a 

= Metapenaeopsis durus Kubo, 1949a: 421; Figs 8A, 18G-L, 22O, 46D, 64C-C', 76E, J, 80L, 148C, 149. 

[Tokyo fish market, Miya, Aichi Prefecture, Japan] 

Metapenaeopsis erythraea Crosnier, 1987a: 443; Figs 16c, 18c, 19e-f. [Valdivia stn 122, Mer Rouge, 21°22'N 

39°04'E, 383-363 m] 

Metapenaeopsis evermanni (Rathbun, 1906) 

= Metapenaeus evermanni Rathbun, 1906: 904; Fig. 58. [Albatross stn 3849, south coast of Molokai Island, 

Hawaii Islands, 79-134 m] 

Metapenaeopsis faouzii (Ramadan, 1938) 

= Penaeopsis (Metapenaeus) faouzii Ramadan, 1938: 72; Fig. 14d. [John Murray Expedition stn 161, 

Maldives, 5°04'48"N 72°50'30"E, 46 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis fusca R.J.G. Manning, 1988: 91; Fig. 1A-D. [Singleton, south Western Australia (32°27'S 

115°44'E)] 

Metapenaeopsis gaillardi Crosnier, 1991: 200; Figs 25-27. [Lagon stn 314, Iles des Pins, Nouvelle-Calédonie, 

27 m] 

Metapenaeopsis gallensis (Pearson, 1905) 

= Parapenaeus gallensis Pearson, 1905: 72; Plate 1, figs 3, 3a-b. [S of Galle, Ceylon] 

Metapenaeopsis gerardoi Pérez Farfante, 1971a: 20; Figs 11-12, 13C. [Oregon stn 5440, off Mayagüez, Puerto 

Rico, 18°08.5'N 67°23'W, 22 m] 

Metapenaeopsis goodei (Smith, 1885b) 

= Parapenaeus Goodei Smith, 1885b: 176. [Bermuda; Bay of Panama] 

= Archipenæopsis vestitus Bouvier, 1905a: 747. [capturé par le Blake dans la mer des Antilles, à la faible 

profondeur de 37 brasses (Blake stn 11, NW of Dry Tortugas Islands, 24°43'N 83°25'W, 37 fms, see A. 

Milne-Edwards & Bouvier, 1909 and Pérez Farfante, 1971a)] 

= Parapenaeopsis Rathbuni Bouvier, 1905a: 748. [dans la mer des Antilles, à 17 brasses de profondeur 

(26°16'25''N 82°50'10''W, west of Florida; see Pérez Farfante, 1971a)] 

Metapenaeopsis hilarula (De Man, 1911a) 

= Penaeopsis sp. (provisional name hilarulus) De Man, 1911a: 70 (partim). [Siboga stn 240, Banda-anchorage, 

Indonesia, 9-36 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis hobbsi Pérez Farfante, 1971a: 24; Figs 13D, 14-17. [Pillsbury stn 737, NW of Cabo Cordera, 

Venezuela, 10°44'N 66°07'W to 10°45'N 66°08'W, 60-73 m]


Metapenaeopsis incisa Crosnier, 1991: 277; Figs 85-87. [BENTHEDI stn 101D, Iles Glorieuses, 11°25.7'N 

47°19.5'E, 26 m] 

Metapenaeopsis ivanovi Crosnier, 1994a: 340; Figs 1-3. [au large du Kenya] 

Metapenaeopsis kishinouyei (Rathbun, 1902c) 

= Parapenaeus kishinouyei Rathbun, 1902c: 288; Plate 12, figs 13-15. [Tagus Cove, reef N of Tagus Hill, 

Albemarle Island] 

Metapenaeopsis kuboi Ivanov & Hassan, 1976a: 1302; Figs 4, 5d, 6f. [Western Indian ocean, 23°26'S 33°31'E, 

410 m] 

Metapenaeopsis kyushuensis (Yokoya, 1933) 

= Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sôyô-Maru stns 296, E of Tanegasima Island, Japan, 

219 m; 317, coast of Miyazaki-ken, Japan, 97 m; 428, W of Amakusa Islands, Japan, 119 m 

Metapenaeopsis lamellata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penaeus lamellatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, Figs 4-5. [Japan; lectotype 

designated by Yamaguchi & Baba, 1993] 

Metapenaeopsis lata Kubo, 1949a 

= Metapenaeopsis latus Kubo, 1949a: 434; Figs 8G, 46G, 76N, T, 81B, 148H, 150. [Off Nagashima, Mie 

Prefecture, Japan, circa 350 m] 

? = Leptopenaeus Kyushuensis Yokoya, 1933: 5; Fig. 2. [Sôyô-Maru stns 296, east of Tanegasima Isl., 219 

m; 317, coast of Miyazaki-ken, 97 m; 428, west of Amakusa Isls., 119 m] 

Metapenaeopsis laubieri Crosnier, 1991: 238; Figs 51-53. [Albatross 1907-1910, stn 5160, 2.75 mi SW de l’île 

Tinakta Island, Malaisie, 5°12'40"N 119°55'10"E, 22 m] 

Metapenaeopsis lindae R.J.G. Manning, 1988: 97; Fig. 2A-D. [Bell Buoy, NW Rottnest Island (31°59'S 

115°34'E), south Western Australia] 

Metapenaeopsis liui Crosnier, 1987a: 434; Figs 3d, 10b-d, 13d-f, 14a, 15. [MUSORSTOM I, stn 40, Philippines, 

13°57.4'N 120°27.8'E, 287-265 m] 

Metapenaeopsis mannarensis De Bruin, 1965: 88; Fig. 2a-d. [Ceylon, off Mannar, 6 fms] 

Metapenaeopsis manningi Crosnier, 1994a: 344; Figs 4-6. [Anton Bruun, Cruise 9, stn 499, Somalie, 10°03'N 

51°15'E, 31-39 m] 

Metapenaeopsis marquesas Crosnier, 1991: 244; Figs 57-59. [Nat. Geogr. Marquesas Exped. stn UP II/Haul 

5, côte ouest de Ua Pou, Iles Marquises, Polynésie, 48-51 m] 

Metapenaeopsis martinella Pérez Farfante, 1971a: 16; Figs 9-10, 13B. [Oregon stn 4252, off Acaraú, Ceará, 

Brazil, 2°10'S 39°52'W, 137 m] 

Metapenaeopsis menoui Crosnier, 1991: 180; Figs 6c-d, 8-9. [Reves 2 stn 19, Seychelles, 5°54.5'S 56°19.4'E, 

30-35 m] 

Metapenaeopsis miersi (Holthuis, 1952a) 

= Penaeopsis miersi Holthuis, 1952a: 80; Fig. 19. [nomen novum for Penæus velutinus sensu Miers, 

1881a nec Dana, 1852a and nomen novum for Metapenæopsis pubescens Bouvier, 1905b nec Penæus 

pubescens Stimpson, 1871] 

= Metapenæopsis pubescens Bouvier, 1905b: 981; nec Penæus pubescens Stimpson, 1871. [îles du Cap 

Vert] 

Metapenaeopsis mineri Burkenroad, 1934a: 25; Figs 8-10. [Conception Bay, Lower California, USA] 

Metapenaeopsis mogiensis complanata Crosnier, 1991: 219; Figs 37o-r, 38d. [Soela Cruise 682, stn 128, côte 

nord-ouest de Australie, 19°07.9'S 119°06.9'E, 78 m] 

Metapenaeopsis mogiensis consobrina (Nobili, 1904) 

= Metapenaeus consobrinus Nobili, 1904: 230. [Djibouti; lectotype designation by Crosnier, 1991] 

= Metapenaeus perlarum Nobili, 1905a: 158. [côtes d’Arabie, dragages entre 25°10'N 55°10'E – 24°55'N 

54°40'E, 16-24 m, according to Crosnier, 1991] 

= Metapenaeopsis bruini Starobogatov, 1972: 405, 408; Plate 11, fig. 158. [Sri Lanka, E of Mullaitivu 

lighthouse, off Mannar, 6 fms; nomen novum for M. mogiensis sensu De Bruin, 1965 nec Rathbun, 

1902b] 

Metapenaeopsis mogiensis intermedia Crosnier, 1991: 222; Figs 37s-w, 38e. [MUSORSTOM 3, stn CP 142, 

Philippines, 11°47'N 123°02'E, 26-27 m]


Metapenaeopsis mogiensis mogiensis (Rathbun, 1902b) 

= Parapenaeus mogiensis Rathbun, 1902b: 39; Figs 6-8. [Mogi, Japan; lectotype designation by Crosnier, 

1991] 

Metapenaeopsis novaeguineae (Haswell, 1879) 

= Penaeus Novae-Guineae Haswell, 1879: 43. [Chevert Expedition, Katow, Papua New Guinea] 

Metapenaeopsis palmensis (Haswell, 1879) 

= Penaeus Palmensis Haswell, 1879: 43. [Palm Island, Queensland, Australia] 

= Metapenaeopsis barbeensis Hall, 1962: 32; Figs 118-118f. [35 miles ESE of St. Barbe (= Pejantan), 

0°07'S 107°46'E, 20 fms (37 m)] 

Metapenaeopsis parahilarula Crosnier, 1991: 232; Figs 45e, 47. [MUSORSTOM 3, stn CP 121, Philippines, 

12°08.3'N 121°17.3'E, 73-84 m] 

Metapenaeopsis parapalmensis Crosnier, 1994b: 313; Figs 37-38. [Albatross stn 5159, archipel de Sulu, 

groupe des Tawitawi, île Tinakta, 5°11'50"N 119°54'E, 18-22 m] 

Metapenaeopsis persica Crosnier, 1991: 210; Figs 32, 33a-d. [Dammam, district d’Hasa, Arabie Saoudite, 

Golfe Persique, 26°25'N 50°06'E] 

Metapenaeopsis philippii (Spence Bate, 1881) 

= Penaeus Philippii Spence Bate, 1881: 181. [Challenger stn 201, 7°3'N 121°48'E, 150 m; lectotype designation 

by Crosnier, 1987a] 

= Penaeus philippinensis Spence Bate, 1888: 261; Plate 35, figs 2-2", 3-3". [Challenger stn 201, 7°3'N 

121°48'E, 150 m; lectotype designation by Crosnier, 1987a] 

Metapenaeopsis propinqua Crosnier, 1991: 186; Figs 12-13. [”Patate bayonnaise”, lagon est, Nouvelle-Calédonie, 

20 m] 

Metapenaeopsis provocatoria longirostris Crosnier, 1987a: 435, Figs 1d, 7g-j, 8m-o. [Soela Cruise 184 stn 08, 

Australie, 19°20.2'S 115°44.1'E, 306-308 m] 

Metapenaeopsis provocatoria provocatoria Racek & Dall, 1965: 48; Fig. 10A-D. [Challenger stn 31, NNE of 

Cape Moreton, Queensland, Australia, 80-90 fms] 

Metapenaeopsis provocatoria owstoni Shinomiya & Sakai, 2000: 123; Figs 7, 8a-f, 9a-m, 11, 12. [Okinose, 

Sagami Bay, Japan, ca. 35°N 139°35'E] 

Metapenaeopsis proxima Crosnier, 1991: 258; Figs 68-70, 78a. [REVES 2 stn 12, Seychelles, 5°13.5'S 56°08.5'E, 

60 m] 

Metapenaeopsis quadrilobata Crosnier, 1991: 199; Figs 23-24. [Nosy Be (Pointe Lokobe), Madagascar, 10 m] 

Metapenaeopsis quinquedentata (De Man, 1907a) 

= Metapeneus quenquedentatus De Man, 1907a: 133. [Siboga stn 109, off Tongkil Island, Sulu Archipelago, 

Philippines, 13 m; lectotype designation by Crosnier, 1991] 

Metapenaeopsis richeri Crosnier, 1991: 280; Figs 88-89. [CORAIL 2, stn DW 147, îlles Chesterfield, 

19°36.87'S 158°13.52'E, 25 m] 

Metapenaeopsis rosea Racek & Dall, 1965: 29; Figs 2D, 3; Plate 1, fig. 4; Plate 4, Figs 7, 8; Plate 9, fig. 4. 

[Mackay, Queensland, Australia, 4 fms] 

Metapenaeopsis scotti Champion, 1973: 195; Figs 3A, 4A-D. [South Africa, Natal, off Durban, 160 fms] 

Metapenaeopsis sibogae (De Man, 1907a) 

= Metapeneus sibogae De Man, 1907a: 131. [Siboga stns 306, 8°27'S 122°54.5'E, 247 m; 312, Saleh-bay, 

north coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

Metapenaeopsis sinica Liu & Zhong, 1988: 3, 224, 270; Fig. 136. [Western Guangdong, South China Sea, 64 m] 

= Metapenaeopsis sinensis Liu & Zhong, 1988: 24, 216 (key). [nomen nudum] 

Metapenaeopsis sinuosa Dall, 1957 

= Metapenaeopsis sinuosus Dall, 1957: 176; Fig. 14A-F. [Off shore island (Kennedy Sound), Cumberland 

Group, NE coast of Australia, 16 m] 

Metapenaeopsis smithi (Schmitt, 1924a) 

= Penaeopsis smithi Schmitt, 1924a: 62; Figs 1b-c, 2a-c. [Caracas Baai, Curaçao] 

Metapenaeopsis spatulata Crosnier, 1991: 224; Figs 40-41. [MUSORSTOM 3, stn CP 142, Philippines, 

11°47'N 123°02'E, 26-27 m] 

Metapenaeopsis spiridonovi Crosnier, 1991: 268; Figs 77a-f, 78d. [REVES 2 stn 33, Seychelles, 4°25.9'S 

54°39.0'E, 45-60 m]


Metapenaeopsis stokmani Burukovsky, 1990: 189; Fig. 1A. [25°39'S 85°24'W, 160-192 m] 

Metapenaeopsis stridulans (Alcock, 1905) 

= Metapenaeus stridulans Alcock, 1905: 518, 526. [from Orissa to Palk Strait, 20-35 fms; Gulf of Martaban, 

20 fms; various places in the Andamans, 20 fms] 

? = Metapenaeopsis tchekunovae Starobogatov, 1972: 402, 413; Plate 9, fig. 107. [Arabian Sea off Pakistan, 

off Indus delta, 23°48.5' - 23°47.7'N 67°17.5' - 67°18.7'E] 

Metapenaeopsis tarawensis Racek & Dall, 1965: 46; Fig. 9A-D. [Tarawa Group, Gilbert Islands] 

Metapenaeopsis tenella Liu & Zhong, 1988: 242; Fig. 146. [East of Hainan Island, South China Sea, 270 m] 

Metapenaeopsis toloensis Hall, 1962 

= Metapenaeopsis tolœnsis Hall, 1962: 33; Figs 119-119d. [Central part of South China Sea, 6°13'N 

107°49'E, 40 fms; 5°51'N 107°53'E, 38 fms] 

Metapenaeopsis vaillanti (Nobili, 1904) 

= Metapenaeus vaillanti Nobili, 1904: 229. [Suez; Mer Rouge] 

? = Metapenaeopsis philippii var. Attaqa Al-Kholy & El-Hawary, 1970: 361. [Gulf of Suez till El-Adabiah, 

5-8 fms] 

Metapenaeopsis velutina (Dana, 1852a) 

= Penaeus velutinus Dana, 1852a: 27. [Albatross 1902, stn 3853, côte sud de l’île Molokai, Hawaii, 210- 

245 m; neotype designation by Crosnier, 1991] 

= Metapenaeopsis insona Racek & Dall, 1965: 41; Fig. 6A-B; Plate 2, fig. 3; Plate 10, fig. 2. [7 miles off 

Long Reef, New South Wales, Australia, 73 m] 

= Metapenaeopsis caliper Liu & Zhong, 1988: 238, 269; Fig. 145. [S of mouth of Pearl River, South 

China Sea, 138 m] 

Metapenaeopsis wellsi Racek, 1967: 251; Plates 12-13. [Exmouth Gulf, Western Australia] 

Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a 

= Mangalura Miers, 1878 (type species Penæus dobsoni Miers, 1878, by monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in Opinion 864 in 1969: ruled under 

the plenary powers not to be given precedence over Metapenaeus Wood-Mason, 1891) 

= Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891a (type species Penaeus affinis H. Milne 

Edwards, 1837 [in Milne Edwards, 1834-1840], by original designation, gender masculine) 

Metapenaeus affinis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus affinis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 416. [côte de Malabar] 

= Penaeus mutatis Lanchester, 1901: 572; Plate 34, fig. 6. [Type locality not indicated; lectotype designation 

by Miquel, 1982] 

= Metapenaeus necopinans Hall, 1956: 83; Plate 11, fig. 15; Plate 12, fig. 16. [Jurong, Singapore] 

= Metapenaeus ivanovi Hassan, 1978: 385; Figs 1-2. [Arabian Gulf, 29°145'N 49°30'E, 25 m] 

Metapenaeus alcocki M.J. George & Rao, 1968: 146; Fig. 1. [Gulf of Kutch, Northwest coast of India, 

22°48'N 70°03'E, 3-12 m] 

Metapenaeus anchistus (De Man, 1920a) 

= Penaeopsis intermedia anchista De Man, 1920a: 5; Plate 1, Figs 3-3d. [Ternate, Indonesia; lectotype 

designation by Miquel, 1982] 

Metapenaeus arabicus Hassan, 1978: 387; Figs 3-5. [Arabian Gulf, 29°145'N 49°30'E, 25 m] 

Metapenaeus bennettae Racek & Dall, 1965: 74. [Lake Budgewoi, Tuggerah Lakes, New South Wales, Australia] 

Metapenaeus brevicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus brevicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 417. [côtes de l’Inde] 

= Penaeus avirostris Dana, 1852a: 27. [Singapore] 

Metapenaeus conjunctus Racek & Dall, 1965: 64; Fig. 12; Plate 5, figs 8-9; Plate 11, fig. 1. [Shallow brackish 

water at mouth of Tuaran river, Sabah, N Borneo, Malaysia, 4-5 fms] 

Metapenaeus dalli Racek, 1957: 4; Plate 1, figs 1-3; Plate 2, figs 1-2; Plate 3, fig. 3. [Peel Inlet, Murray River, 

West Australia] 

Metapenaeus demani demani (Roux, 1921) 

= Penaeopsis demani Roux, 1921: 599; Plate 16, figs 4-12. [Varen and Lorentz Rivers, New Guinea]


Metapenaeus demani stephani Miquel, 1982: 74; Figs 24, 25a, c, 26a-e. [Port Moresby, Golfe de Papoua, 

Nouvelle-Guinée] 

Metapenaeus dobsoni (Miers, 1878) 

= Penæus dobsoni Miers, 1878: 302; Plate 17, fig. 2. [Mangalur (Mangalore), west coast of India] [as 

Mangalura dobsoni on page 303] 

= Metapenaeus dobsoni choprai Holthuis, 1980b: 22. [nomen nudum] 

Metapenaeus eboracensis Dall, 1957: 193; Fig. 19a-g. [Mouth of Norman River, Gulf of Carpentaria, 

Queensland, Australia, 2-4 m] 

Metapenaeus elegans De Man, 1907a: 130. [Siboga stn 121, Manado, Cèlébes, Indonesia, 55 m] 

= Metapenaeus singaporensis Hall, 1956: 84; Plate 12, figs 17-19. [Jurong, Singapore] 

Metapenaeus endeavouri (Schmitt, 1926a) 

= Penaeopsis endeavouri Schmitt, 1926a: 329; Plate 59, figs 1-3; Plate 68, fig. 4. [FIS Endeavour expedition, 

S coast of Queensland, Australia] 

Metapenaeus ensis (De Haan, 1844 [in De Haan, 1833-1850]) 

= Penoeus Ensis De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 2. [Japan; lectotype designation 

by Miquel, 1982] 

= Penaeus Mastersii Haswell, 1879: 42. [80 km WNW of Darwin, Northern Territory, Australia] 

= Penaeus incisipes Spence Bate, 1888: 257; Plate 34, fig. 2. [Challenger stns 190, Arafura Sea, south of 

Papua, 8°56'S 136°5'E, 49 fms; 203, off Panay, Philippines, 11°6'N 123°9'E, 20 fms] 

= Metapenaeus ensis var. baramensis Hall, 1962: 23. [30 miles NNE of Tanjong Baram, 4°58'N 113°58'E, 

77 m] 

= Metapenaeus philippinensis Motoh & Muthu, 1979: 1351; Figs 1-2. [Himamaylan, Philippines, 10- 

15 m] 

Metapenaeus insolitus Racek & Dall, 1965: 69; Fig. 13A-C; Plate 6, figs 3, 4; Plate 11, fig. 4. [Chambers Bay, 

Northern Territory, Australia, 30 m] 

Metapenaeus intermedius (Kishinouye, 1900) 

= Penaeus intermedius Kishinouye, 1900: 21, unnumbered textfig. [province of Tosa] 

Metapenaeus joyneri joyneri (Miers, 1880) 

= Penaeus Joyneri Miers, 1880: 458; Plate 15, figs 8-10. [Yokohama, Japan] 

= Penaeus pallidus Kishinouye, 1897: 275. [Tokyo Bay, Ise area, Seto Inland Sea; Kagoshima Bay] 

Metapenaeus joyneri formosus Lee & Yu, 1977: 101; Fig. 71. [Donggang, Pingtong County, SW Taiwan] 

Metapenaeus krishnatrii Silas & Muthu, 1976: 645; Plate 1; fig. 1. [Corbyn’ Cove, Port Blair, Andaman Islands] 

Metapenaeus kutchensis P.C. George, M.J. George & Rao, 1963: 284; Fig. 1. [Gulf of Kutch, N.W. India, 

3-12 m] 

Metapenaeus lysianassa (De Man, 1888a) 

= Penaeus lysianassa De Man, 1888a: 290. [Mergui Archilepago, Burma] 

= Metapenaeus lysianassa var. malaccænsis Hall, 1962: 24. [Kuala Sanglang, 1 m] 

Metapenaeus macleayi (Haswell, 1879) 

= Penæus macleayi Haswell, 1879: 40. [Port Jackson, New South Wales, Australia] 

= Penaeus haswelli Phillips, 1925: 3; Plate 2, fig. 1. [shallow waters of New South Wales] 

Metapenaeus monoceros (Fabricius, 1798) 

= Penaeus monoceros Fabricius, 1798: 409. [Type locality restricted by Holthuis & Gottlieb, 1958 to 

Tranquebar, Madras Province, India] 

? = Penaeopsis spinulicauda Stebbing, 1914b: 17; Plate 68. [Durban Bay] 

? = Metapenaeus cognatus Nobili, 1904: 229. [Djibouti, Mer Rouge] 

? = Metapenaeus Deschampsi Nobili, 1903a: 452; Plate 2, fig. 1. [Pondichéry, Foce dell’Arian Koupur; 

Mahè] 

Metapenaeus motohi Shinomiya & Sakai, 2009: 1068; Fig. 1. [Tigbauan Panay I., Philippines] 

Metapenaeus moyebi (Kishinouye, 1896) 

= Penaeus moyebi Kishinouye, 1896: 373. [Japan; Makassar, Celebes, Indonesia] 

= Metapenaeus burkenroadi Kubo, 1954: 92; Fig. 1A-D. [Japan]


Metapenaeus papuensis Racek & Dall, 1965: 66; Plate 3, fig. 3; Plate 6, figs 1-2; Plate 11, fig. 2. [Mouth of 

Panaroa river, Gulf of Papua, Papua New Guinea] 

= Metapenaeus bengalensis Tirmizi, 1971b: 242; Figs 1-3. [Bay of Bengal] 

Metapenaeus stebbingi Nobili, 1904 

= Metapenaeus Stebbingi Nobili, 1904: 229. [Mer Rouge; Suez; full description in Nobili, 1906a] 

Metapenaeus suluensis Racek & Dall, 1965: 61; Fig. 11A-B; Plate 5, figs 6-7; Plate 10, fig. 8. [Philippines, 

Sulu Sea, 20 fms] 

Metapenaeus tenuipes Kubo, 1949a: 348; Figs 7R, T, 22B, 31G, H, 47G, 62B-B', 74N-M, S-T, 81H, K, 125G, 

127, 129F, 130. [S coast of Borneo, Indonesia] 

= Metapenaeus spinulatus Kubo, 1949a: 355; Figs 7T, 47G, 74M, S, 81K, 129F, 130. [Bangkok, Thailand] 

Mierspenaeopsis Sakai & Shinomiya, 2011 

= Mierspenaeopsis Sakai & Shinomiya, 2011 (type species Penaeus sculptilis Heller, 1862a, by original 


Mierspenaeopsis cultrirostris (Alcock, 1906) 

= Parapenaeopsis sculptilis var. cultrirostris Alcock, 1906: 39; Plate 7, fig. 23. [India: Orissa and Ganjam; 

Sunderbunds] 

Mierspenaeopsis hardwickii (Miers, 1878) 

= Penæus hardwickii Miers, 1878: 300; Plate 17, figs 1-1a. [Indian Seas?] 

Mierspenaeopsis indica (Muthu, 1972) 

= Parapenaeopsis indica Muthu, 1972: 174; figs 1-6. [Kakinada Bay, Kakinada, east coast of India, 

16°56.5'N 82°16.5'E, 1-4 m] 

Mierspenaeopsis sculptilis (Heller, 1862a) 

= Penaeus sculptilis Heller, 1862a: 528. [Java] 

Parapenaeopsis Alcock, 1901 

= Parapeneopsis Alcock, 1901 (type species Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne 

Edwards, 1834-1840], by original designation, gender feminine; name emended under the plenary 

powers of the ICZN to Parapenaeopsis and placed in that emended spelling on the Official List of 

Generic Names in Zoology in Opinion 864 in 1969) 

Parapenaeopsis acclivirostris Alcock, 1905: 522, 530. [Persian Gulf; in Palk Strait; at Madras; off the Vizagapatam 

and Ganjam coasts] 

Parapenaeopsis aroaensis Hall, 1962 

= Parapenæopsis aroaensis Hall, 1962: 27; Figs 109-109a. [20 mls NNE of Aroa Islands, 3°20'N 100°39'E, 

51 m] 

Parapenaeopsis balli Burkenroad, 1934a: 64; Fig. 17. [Acajutla, El Salvador, 13°36'N 89°50'W] 

Parapenaeopsis gracillima Nobili, 1903b: 4; Fig. 1. [Buntal, Borneo] 

Parapenaeopsis longirostris Chandra & Bhattacharya, 2004: 23; Fig. 1. [Ongaria Ghat, Baleshwar, Orissa] 

Parapenaeopsis nana Alcock, 1905: 522, 529. [Off Ganjam and Orissa coasts up to 68 fms; at Madras] 

Parapenaeopsis stylifera (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Penaeus styliferus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [les environs de 

Bombay] 

= Parapeneopsis stylifera var. coromandelica Alcock, 1906: 37; Plate 7, figs 20, 20a-c. [Madras, Coromandel 

Coast; lectotype designation by Ravindranath, 1989] 

= Parapenaeopsis stylifera var. cochinensis M.J. George, 1974: 421; Fig. 1b, d. [unavailable name under 

Art. 15.2] 

Parapenaeus Smith, 1885b 

= Parapenæus Smith, 1885b (type species Penaeus longirostris Lucas, 1846, by original designation, 


= Neopenæopsis Bouvier, 1905a (type species Neopenæopsis paradoxus Bouvier, 1905a, by original designation 

and monotypy, gender feminine)


Parapenaeus americanus Rathbun, 1901: 102; Plate 2. [Mayaguez Harbour, 220-225 fms] 

Parapenaeus australiensis Dall, 1957: 179; Fig. 15A-F. [Off Newcastle, New South Wales, Australia] 

= Parapenaeus australiensis forma nodosus Crosnier, 1986a: 313, 339. [types not indicated; unavailable 

under Art. 15.2] 

Parapenaeus cayrei Crosnier, 2005: 262; Figs 3, 4A-C. [BORDAU 2, stn CP 1541, Îles Tonga, 21°15'S 

175°14'W, 319-333 m] 

Parapenaeus fissuroides erythraeus Crosnier, 1986a: 329; Figs 5i, 6k, m, 8c-d. [Pola stn 117, Mer Rouge, 

20°16'N 37°33'E, 638 m] 

Parapenaeus fissuroides fissuroides Crosnier, 1986a: 321; Figs 5e-f, 6d-f, 7c, 8a. [Hong Kong, 19°22.5'N 

114°07.5'E, 700-795 m] 

Parapenaeus fissuroides indicus Crosnier, 1986a: 325; Figs 5g, h, 6g-j, l, 7d, f-g, 8b. [Vauban stn CH 80, 

Madagascar, 25°02.7'S 47°05.8'E, 65-70 m] 

Parapenaeus fissurus (Spence Bate, 1881) 

= Penaeus fissurus Spence Bate, 1881: 180. [south of New Guinea, 50 fms] 

= Penaeus fissurus Spence Bate, 1888: 263; Plate 36, fig. 1. [Challenger stns 190, 8°56'S 136°5'E, 49 fms; 

204A, 12°43'S 122°9'E, off Tablas Island, 100 fms; 204B, 12°46'S 122°10'E, off Tablas Island, 115 fms; 

209, 10°14'N 123°54'E, off Zebu, 95-100 fms] 

Parapenaeus investigatoris Alcock & Anderson, 1899: 279. [Investigator stns 233 (Andaman Sea, 13°17'15''N 

93°10'25''E), 185 fms; 235 (Andaman Sea, 14°13'N 93°40'E), 370-419 fms; 166 (Bay of Bengal, 

13°34'55''N 80°32'12''E), 133 fms] 

Parapenaeus kensleyi Crosnier, 2005: 258; Figs 1-2. [HALIPRO I, stn CP 853, Nouvelle-Calédonie, 21°45'S 

166°37'E, 241-250 m] 

Parapenaeus lanceolatus Kubo, 1949a: 405; Figs 4C, 7D', 21C, 28G-I, 61C, C', 75C-I, 78I, 143C, G, 144. [Heta, 

Shizuoka Prefecture and Kumano-nada, off Owase, Mie Prefecture, Japan, 300-350 m] 

Parapenaeus longipes Alcock, 1905: 520, 525. [off Malabar coast (Mangalore), 21-26 fms; off Orissa and 

Ganjan coasts, 20-68 fms; off Vizagapatam coast, 7,5-23 fms; S coast of Ceylon, 20 fms] 

= Parapenaeus longipes forma denticulatus Crosnier, 1986a: 311, 312, 347. [Indonésie et Philippines, 

exact type locality not indicated, see Crosnier, 1986a; unavailable under Art. 15.2] 

Parapenaeus longirostris (Lucas, 1846) 

= Peneus Cocco Prestandrea, 1833: 6. [Mari di Messina; suppressed under the plenary powers for the 

purposes of the Principle of Priority but not those of the Principle of Homonymy in Opinion 611 in 

1961] 

= Penaeus longirostris Lucas, 1846: 46; Plate 4, fig. 6. [Algerie, Alger, aux environs du cap Matifou] 

= Penæus bocagei Johnson, 1863: 255. [mouth of the Tagus] 

= Penaeus lividus Filhol, 1885b: Plate 1. [Type locality not indicated, 500-600 m] 

Parapenaeus murrayi Ramadan, 1938: 74; Fig. 15a-c. [John Murray Expedition stn 105 (°36'12''N 

39°13'12''E), Zanzibar area, 238-293 m] 

Parapenaeus perezfarfantae Crosnier, 1986a: 339; Figs 12a-b, 13a-b, 14a. [Albatross expedition stn 5183, 

Philippines, 10°32'48"N 122°26'E, 176 m] 

Parapenaeus politus (Smith, 1881) 

= Penaeus politus Smith, 1881: 444. [Fish Hawk stn 878 (39°55'N 70°54'W), 142 fms, off the eastern end 

of Long Island, known as the Block Island] 

= Neopenæopsis paradoxus Bouvier, 1905a: 747. [dans la mer des Antilles par 84 et 91 brasses de profondeur] 

Parapenaeus ruberoculatus Hall, 1962: 31; Figs 115-115b. [30 Miles NNE of Tanjong Baram, 5°04'N 113°48'E, 

183 m] 

Parapenaeus sextuberculatus Kubo, 1949a: 403 (partim); Figs 7F', 21B, 28D-F, 61A-A', 75B, H, 78H, 143B, F 

(nec Fig. 47R = P. fissuroides fissuroides Crosnier, 1986a). [Heta and Maisaka, Shizuoka Prefecture, 

Japan, circa 330 m] 

Pelagopenaeus Pérez Farfante & Kensley, 1997 

= Pelagopenaeus Pérez Farfante & Kensley, 1997 (type species Penaeus balboae Faxon, 1893, designated 

by Pérez Farfante & Kensley, 1997, gender masculine)


Pelagopenaeus balboae (Faxon, 1893) 

= Penaeus balboae Faxon, 1893: 211. [Albatross stn 3371 (off Isla del Coco, Costa Rica, 5°26'20"N 

86°55'00"W), 770 fms] 

= Penaeus meridionalis Lenz & Strunck, 1914: 298, Plate 15, figs 1-15. [Deutsche Südpolar-Expedition, 

30°21'S 14°2'W, 10 m] 

Penaeopsis Spence Bate, 1881 

= Penaeopsis Spence Bate, 1881 (type species Penaeopsis serratus Spence Bate, 1881, designated by 

Bouvier, 1905b, gender feminine) 

Penaeopsis balssi Ivanov & Hassan, 1976b: 1; Figs 1-2. [off central Mozambique, 25°26'S (not 23°26'S as 

stated), 33°31'E, 410 m] 

Penaeopsis challengeri De Man, 1911a: 76. [nomen novum for Penaeus serratus Spence Bate, 1881] 

= Penaeus serratus Spence Bate, 1881: 182. [Challenger stn 173, off Matuku, Fiji Islands, 19°09'35"S 

179°41'50"E, 315 fms, lectotype designation by Pérez Farfante, 1980a] 

Penaeopsis eduardoi Pérez Farfante, 1977a: 172; Figs 1-4. [Albatross stn 5116, Balayan Bay, Luzon Island, 

Philippines, 13°41'00"N 120°47'05"E, 200 fms] 

Penaeopsis jerryi Pérez Farfante, 1979: 209; Figs 1-3. [John Murray Expedition stn 16, off Berbera, Somalia, 

Gulf of Aden, 10°29'48"N 45°01'48"E, 186 m] 

Penaeopsis mclaughlinae Crosnier, 2006: 334; Figs 2-5. [CHALCAL 2 stn CC 1, Nouvelle-Calédonie, 

24°54.96'S 168°21.91'E, 500 m] 

Penaeopsis rectacuta (Spence Bate, 1881) 

= Penaeus rectacutus Spence Bate, 1881: 180. [among the Philippine Islands, about 100 fms; according 

to Pérez Farfante, 1980a: Challenger stn 209, between Bohol and Cebu, Philippines, 10°14'N 123°54'E, 

95 fms] 

Penaeopsis serrata Spence Bate, 1881 

= Penaeopsis serratus Spence Bate, 1881: 183. [Gulf of Mexico; according to Pérez Farfante, 1980a: 

Blake stn 275, off Barbados, Gulf of Mexico, 12°58'33"N 59°36'45"W, 218 fms; name placed on the 

Official List of Specific Names in Zoology in Opinion 864 in 1969] 

= Parapenaeus megalops Smith, 1885b: 172. [Albatross stns 2125, S of Curaçao, 11°43'N 69°09'30"W, 208 

fms; 2143, Golfo de Urabá, 9°30'45"N 76°25'30"W, 155 fms] 

= Artemesia talismani Bouvier, 1905b: 982. [le Talisman captura en quantité assez grande au large des 

côtes du Maroc et du Sahara; Talisman stn 72, off Guerguerat, Western Sahara, 25°41'N 15°56'W (of 

Greenwich) 18°16'W (of Paris) on label accompanying specimen), 410 m, see Pérez Farfante, 1980a] 

= Penaeopsis serratus var. antillensis A. Milne-Edwards & Bouvier, 1909: 226; Plate 3, fig. 10; Plate 4, 

fig. 5. [Blake stn 148, off St Kitts, 208 fms] 

Penaeus Fabricius, 1798 

= Penaeus Fabricius, 1798 (type species Penaeus monodon Fabricius, 1798, designated by Latreille, 

1810, gender masculine) 

Penaeus esculentus Haswell, 1879: 38. [Port Jackson, New South Wales, Australia] 

Penaeus monodon Fabricius, 1798: 408. [Bay of Batavia [=Jakarta], Indonesia; neotype designation by 

Holthuis, 1949a] 

= Penaeus carinatus Dana, 1852a: 27. [Singapore] 

= Penaeus tahitensis Heller, 1862a: 528. [Taiti] 

= Penaeus semisulcatus exsulcatus Hilgendorf, 1879: 843. [Quelimane, Mozambique] 

= Penaeus coeruleus Stebbing, 1905: 77; Plates 21-21bis. [Nahoon River on E coast near East London, 

South Africa] 

? = Penaeus durbani Stebbing, 1917a: 442; Plate 22. [Durban Bay] 

= Penaeus bubulus Kubo, 1949a: 296; Figs 1G, 7B, 15U-B', 24K-M, 37, 49C, 53, 58D, 67N-Q, 73F, L, 77P, 

113, 114. [Hatazawa, Chiba Prefecture: Miya; Aichi Prefecture; Tainan, Formosa; Palau; Batavia; 

Menado, Celebes] 

Penaeus semisulcatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 1. [Japan] 

? = Penaeus Manilensis de Procé, 1822: 134. [Manille]


= Penaeus ashiaka Kishinouye, 1900: 14; Plate 3; Plate 7, fig. 4. [southern half of our coast] 

= Penaeus semisulcatus paucidentatus Parisi, 1919: 65; Plate 5, fig. 5. [Misaki, Baia di Sagami] 

= Penaeus monodon manillensis Villaluz & Arriola, 1938: 39; Plate 3, fig. 1. [Manila Bay, Philippines] 

Protrachypene Burkenroad, 1934a 

= Protrachypene Burkenroad, 1934a (type species Protrachypene precipua Burkenroad, 1934a, by 

monotypy, gender feminine) 

Protrachypene precipua Burkenroad, 1934a: 43. [Bella Vista Beach, Panama City, Panama] 

Rimapenaeus Pérez Farfante & Kensley, 1997 

= Rimapenaeus Pérez Farfante & Kensley, 1997 (type species Trachypenaeus faoe Obarrio, 1954, by 


Rimapenaeus byrdi (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) byrdi Burkenroad, 1934a: 51; Fig. 13. [Panama City Market, Panama] 

Rimapenaeus constrictus (Stimpson, 1871) 

= Penaeus constrictus Stimpson, 1871: 135. [Beaufort, N.C., 4 fms; Charleston Harbour] 

= Penaeopsis agassizii Bouvier, 1905a: 748. [provient de Sombrero] 

Rimapenaeus faoe (Obarrio, 1954) 

= Trachypenaeus faoe Obarrio, 1954: 3. [Golfo de Panamá, en el Pacífico] 

Rimapenaeus fuscina (Pérez Farfante, 1971b) 

= Trachypenaeus fuscina Pérez Farfante, 1971b: 637; Figs 1-2, 3A, 4A-F, 5A, 6. [Off Cocodrillo Chiapas, 

Pacific coast of México, 22 m] 

Rimapenaeus pacificus (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) similis pacificus Burkenroad, 1934a: 50. [Pearl Islands, Gulf of Panama, 

Panama, 8°29'40"N 78°52'30"W, 19-24 fms] 

Rimapenaeus similis (Smith, 1885b) 

= Parapenaeus constrictus similis Smith, 1885b: 175. [Albatross stn 2121, Gulf of Paria, 10°37'40"N 

61°42'40"W, 31 fms] 

Tanypenaeus Pérez Farfante, 1972 

= Tanypenaeus Pérez Farfante, 1972 (type species Tanypenaeus caribaeus Pérez Farfante, 1972, by original 


Tanypenaeus caribaeus Pérez Farfante, 1972: 187. [Pillsbury stn 788, off Punta Faro, Colombia, 11°08'N 

74°47'W, 155-157 m] 

Trachypenaeopsis Burkenroad, 1934a 

= Trachypeneopsis Burkenroad, 1934a (type species Metapenaeus mobilispinis Rathbun, 1915, by original 


= Trachypenaeopsis [emendment of Trachypeneopsis Burkenroad, 1934a, under the plenary powers of 

the ICZN in Opinion 684 in 1969] 

Trachypenaeopsis mobilispinis (Rathbun, 1915) 

= Metapenaeus mobilispinis Rathbun, 1915: 117. [Cave Round Bay, Saba, Netherlands Antilles, 7 m] 

Trachypenaeopsis minicoyensis Thomas, 1972: 117; Fig. 1. [western lagoon of Minicoy island, Laccadive 

Archipelago, 3 m] 

Trachypenaeopsis richtersii (Miers, 1884a) 

= Penæus richtersii Miers, 1884a: 564; Plate 52, fig. A. [Cerf Island, 10 fms] 

Trachypenaeus Alcock, 1901 

= Trachypeneus Alcock, 1901 (type species Penaeus anchoralis Spence Bate, 1881, by original designation, 


= Trachypenaeus [emendment of Trachypeneus Alcock, 1901, under the plenary powers of the ICZN in 

Opinion 864 in 1969]


Trachypenaeus anchoralis (Spence Bate, 1881) 

= Penaeus anchoralis Spence Bate, 1881: 181. [south of New Guinea, 28 fms] 

= Penaeus anchoralis Spence Bate, 1888: 258; Plate 35, fig. 1. [Challenger stns 188, 9°59'S 139°42'E, 

Arafura Sea, south of Papua, 28 fms; 190, 8°56'S 136°5'E, Arafura Sea, south of Papua, 49 fms; off 

Yokohama, 5-20 fms; between stns 190-191, near the Arrou Islands in the Arafura Sea] 

Trachysalambria Burkenroad, 1934a 

= Trachypeneus (Trachysalambria) Burkenroad, 1934a (type species Penaeus curvirostris Stimpson, 

1860a, by original designation, gender feminine) 

Trachysalambria albicoma (Hayashi & Toriyama, 1980) 

= Trachypenaeus albicomus Hayashi & Toriyama, 1980: 69; Figs 1-2. [Japan, Tosa Bay] 

Trachysalambria aspera (Alcock, 1905) 

= Trachypenaeus asper Alcock, 1905: 523, 531. [fully described and illustrated in Alcock, 1906; type 

locality, Ganjam coast, 20-35 fms, according to Alcock, 1906] 

Trachysalambria brevisuturae (Burkenroad, 1934a) 

= Trachypeneus (Trachysalambria) brevisuturae Burkenroad, 1934a: 55; Fig. 14. [Acajutla, El Salvador] 

Trachysalambria curvirostris (Stimpson, 1860a) 

= Penaeus curvirostris Stimpson, 1860a: 44. [Tosa Bay, Kochi, Japan, 33°30'N 133°30'E, 35 m; neotype 

designation by Sakaji & Hayashi, 2003] 

= Metapenaeus palaestinensis Steinitz, 1932: 161; Figs 1-3. [aus der Bucht von Haifa] 

Trachysalambria fulva (Dall, 1957) 

= Trachypenaeus fulvus Dall, 1957: 206; Fig. 23A-G. [Townsville, Queensland, Australia] 

= Trachypeneus unicus Hall, 1961: 102. [S of Singapore, Malaysia, 35 fms] 

Trachysalambria longipes (Paul’son, 1875) 

= Penaeus longipes Paul’son, 1875: 125; Plate 19, figs 1-1a. [Red Sea] 

Trachysalambria malaiana (Balss, 1933) 

= Trachypenaeus curvirostris malaiana Balss, 1933: 234. [Gier 16 Exp. 2, Südl. Sumatra, 6°42'S 103°E; 

Gier 2 Exp. 3, Bai von Batavia; Gier 12 Exp. 4, Javasee vor Borneo, 3°42'S 114°30'E; Gier 4, vor Toeban 

(Nordjava); Gier 9 Exp. 20, nördl. Sumatra, 1°20'S 104°43'E; Gier 17 Exp. 4, westlich Borneo, 1°36'S 

109°46'E; Gier 6 Exp. 21, Bangkastrasse, 2°3'S 105°48,5'E; Gier 4 Exp. 20, Javasee, 6°51'S 112°56'E; 

Gier 4 Exp. 11, vor Pekalongan (Nordjava)] 

Trachysalambria nansei Sakaji & Hayashi, 2003: 162; Figs 7-9. [Japan, Tosa Bay, off Kochi, 33°30'N, 133°30'E, 

75 m] 

Trachysalambria starobogatovi (Ivanov & Hassan, 1976a) 

= Trachypenaeus starobogatovi Ivanov & Hassan, 1976a: 1300; Figs 2, 3a. [Western Indian Ocean, 

19°03'S 36°29'E, 25 m] 

Trachysalambria villaluzi (Muthu & Motoh, 1979b) 

= Trachypenaeus villaluzi Muthu & Motoh, 1979b: 57; Figs 1-2. [Tigbauan, Philippines, 7 m] 

Xiphopenaeus Smith, 1869 

= Xiphopeneus Smith, 1869 (type species Xiphopeneus hartii Smith, 1869, by monotypy, gender masculine) 

= Xiphopenaeus [emendment of Xiphopeneus Smith, 1869, under the plenary powers of the ICZN in 

Opinion 864 in 1969] 

Xiphopenaeus kroyeri (Heller, 1862b) 

= Penaeus Kroyeri Heller, 1862b: 425; Plate 2, fig. 51. [Rio Janeiro] 

= Xiphopeneus Hartii Smith, 1869: 27, 40; Plate 1, fig. 1. [Caravelas, Estado da Bahia, Brazil] 

= Xiphopeneus Riveti Bouvier, 1907: 113; Fig. 1. [achetée sur le marché de Païta]


Family SICyoNIIDAe ortmann, 1898 

Sicyonia H. Milne edwards, 1830 

= Sicyonia H. Milne Edwards, 1830 (type species Sicyonia sculpta H. Milne Edwards, 1830, designated 

by Desmarest, 1858, gender feminine) 

= Ruvulus De Natale, 1850 (type species Sicyonia sculpta H. Milne Edwards, 1830, by monotypy, 


= Synhimantites Boeck, 1864 (type species Synhimantites typicus Boeck, 1864, by monotypy, gender 


= Eusicyonia Stebbing, 1914b (substitute name for Sicyonia H. Milne Edwards, 1830, gender feminine) 

Sicyonia abathophila Crosnier, 2003: 321; Fig. 98. [Albatross, Philippines Expedition, 1908-1909, Camp 

Overton, Iligan Bay, Mindanao, Philippines, 8°15,24'N 124°07,18'E, 2-4 m] 

Sicyonia adunca Crosnier, 2003: 265; Figs 42-44, 107F. [BATHUS 2, stn DW 749, au large de la Nouvelle- 

Calédonie, 22°33,4'S 166°26'E, 233-258 m] 

Sicyonia affinis Faxon, 1893: 209. [Albatross stns 3367, off Isla del Coco, Costa Rica, 5°31'30"N 86°52'30"W, 

183 m; 3369, off Isla del Coco, Costa Rica, 5°32'45"N 86°55'20"W, 95 m; 3378, W of Isla de Malpelo, 

Colombia, 3°58'20"N 81°36'00"W, 205 m; 3379, W of Isla de Malpelo, Colombia, 3°59'40"N 81°35'00"W, 

95 m] 

Sicyonia aliaffinis (Burkenroad, 1934a) 

= Eusicyonia aliaffinis Burkenroad, 1934a: 92; Fig. 24. [Pawnee, Pacific coast of southern Mexico, 

14°48'40"N 92°54'40"W, 19-30 fms] 

Sicyonia altirostrum Crosnier, 2003: 297; Figs 73-75, 109G. [MUSORSTOM 6, stn CP 464, Îles Loyauté, 

21°02,3'S 167°31,6'E, 430 m] 

Sicyonia australiensis Hanamura & Wadley, 1998: 701; Figs 1-4. [Southern Surveyor, cruise SS6/96, stn 183, 

SE coast of Australia, 37°18.7'S, 150°16.7'E, 123 m] 

Sicyonia benthophila De Man, 1907a: 143. [Siboga stn 253, near Kai Islands, Indonesia, 304 m] 

Sicyonia bispinosa (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte bispinosus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 9. [Japan] 

Sicyonia brevirostris Stimpson, 1871: 132. [S Florida coast] 

Sicyonia burkenroadi Cobb, 1971: 104; Figs 1-2. [Oregon stn 1083, Gulf of Mexico off Port Isabel, Texas, 

26°13'N 96°45'W, 42 m] 

Sicyonia carinata (Brünnich, 1768) 

= Cancer carinatus Brünnich, 1768: 102. [Mari Adriatico, Dalmatinis Skilla, in finu maris juxta Salonas] 

= Cancer pulchellus Herbst, 1796 [in Herbst, 1791-1796]: 175; Plate 43, fig. 3. [das Veterland ist mir 

unbekannt] 

= Sicyonia sculpta H. Milne Edwards, 1830: 340; Plate 9, figs 1-8. [Baie de Naples] 

= Sicyonia foresti Rossignol, 1962: 145; 2 unnumbered Figs [Ile du Prince, 27 m] 

Sicyonia curvirostris Balss, 1913b: 235. [Fukuura, Sagami Bay, Japon, 150 m] 

Sicyonia dejouanneti Crosnier, 2003: 293; Figs 70-72, 108G, H, 109A-F. [MUSORSTOM 4, stn CP 194, Nouvelle-Calédonie, 

18°52,8'S 163°21,7'E, 545 m] 

Sicyonia disdorsalis (Burkenroad, 1934a) 

= Eusicyonia disdorsalis Burkenroad, 1934a: 96; Figs 25, 36. [Pawnee, Pearl Islands (Archipelago de las 

Perlas), Gulf of Panama, 8°29'40"N 78°52'30"W, 19-24 fms; Golfo de Panamá; W coast of Central 

America] 

Sicyonia disedwardsi (Burkenroad, 1934a) 

= Eusicyonia disedwardi Burkenroad, 1934a: 86; Figs 23, 29, 34. [Pawnee, Conception Bay, Lower California] 

Sicyonia disparri (Burkenroad, 1934a) 

= Eusicyonia disparri Burkenroad, 1934a: 83; Fig. 27. [Pawnee, Bahia San Luis Gonzaga, Lower California, 

Mexico]


Sicyonia dorsalis Kingsley, 1878a: 97. [Fort Jefferson, Florida] 

Sicyonia fallax De Man, 1907a: 141. [Siboga stn 105, Sulu Sea, Indonesia, 6°08'N 121°19'E, 275 m] 

Sicyonia furcata Miers, 1878: 310; Plate 17, fig. 4. [Sulu Archipelago, Philippines] 

= Sicyonia formosa Chan & Yu, 1985: 99; Fig. 1; Plates 5A, B, 6E-F. [Taiwan, 100 m] 

Sicyonia galeata Holthuis, 1952a: 84. [Mbizi Expedition stns A.S.182, Angola, Luanda, 13 miles WSW of 

Cape Ledo, 9°47'S 13°11'E, 35 m; A.S.170, Congo, 10 miles W of Pointe Noire, 4°48'S 11°41'E, 49 m] 

Sicyonia inflexa (Kubo, 1949a) 

= Eusicyonia inflexa Kubo, 1949a: 458; Figs 8O, 48D, 77C, I, 79G, 159, 160A-F. [Off Atsumi, Aichi 

Prefecture, Japan, ca. 300 m; and Kumanonada, off Owase, Mie Prefecture, Japan, ca. 400 m] 

Sicyonia ingentis (Burkenroad, 1938) (Fig. 4) 

= Eusicyonia ingentis Burkenroad, 1938: 88; Figs 31-34. [Zaca stn 127D-1, off E coast of Cedros Island 

(Isla Cedros), 28°05'N 115°09'W, Baja California, Mexico] 

Sicyonia japonica Balss, 1914b 

= Sicyonia lancifer var. japonica Balss, 1914b: 16; Fig. 9. [Dzushi, 50-100 m; Misaki, 200-300 m; Fukuura; 

Yagoshima, 150 m; zwischen Ito und Hatsushima, 150 m] 

Sicyonia komai Crosnier, 2003: 327; Figs 103-104. [Thailande, côte ouest, Ao Tang Knen, Phuket, zone intertidale] 

Sicyonia laevigata Stimpson, 1871: 131. [Charleston] 

= Sicyonia sculpta var. americana De Man, 1907b: 450. [Off Bahia] 

Sicyonia laevis Spence Bate, 1881: 173. [north of New Guinea, 150 fms; fully described and illustrated in 

Spence Bate, 1888; type locality according to Spence Bate, 1888: Challenger stn 219, north of New 

Guinea, 1°54'S 146°39'40"E, 150 fms] 

= Eusicyonia nebulosa Kubo, 1949a: 454; Figs 8N, 48C, 77B, H, 79K, 156G, 157. [Off Heta, Shizuoka 

Prefecture, Japan, ca. 350 m] 

Sicyonia lancifer (Olivier, 1811) 

= Palæmon lancifer Olivier, 1811: 664. [CORINDON stn 295, Indonesia, Makassar, Strait, 1°26,5'S, 

117°02,1'E, 51-54 m; neotype selection by Crosnier, 2003; emendation to S. lancifera by Holthuis, 

1980b is an unjustified emendation] 

= Hippolyte cristatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 10. [Japan; lectotype designation 

by Yamaguchi & Baba, 1993] 

Fig. 4. Sicyonia ingentis (Burkenroad, 1938). Photo by Sammy De Grave.


Sicyonia longicauda Rathbun, 1906: 908; Plate 20, fig. 6 (partim, nec specimens of stn 4002 = S. truncata 

Kubo, 1949; see Crosnier, 2003). [Albatross stn 3865, Pailolo Channel, Hawaii Islands, 21°09'20"N 

156°35'10"W, 468-517 m] 

Sicyonia longicornis Crosnier, 2003: 276; Figs 55-56. [KARUBAR, stn CP 20, îles Kai, Indonésie, 5°15'S 

132°59'E, 769-809 m] 

Sicyonia martini Pérez Farfante & Boothe, 1981: 424; Figs 1-4. [Pillsbury, stn 556, SW of Punta Ana Maria, 

Golfo de Panama, 7°50'30"N 78°49'00"W, 58 m] 

Sicyonia metavitulans Crosnier, 2003: 304; Figs 79-81. [Port Jackson, côte est de l’Australie] 

Sicyonia mixta Burkenroad, 1946: 3; Figs 1-4. [St. Joseph (probably San José, Lower California), Swedish 

Eugenie Expedition #818] 

Sicyonia nasica Burukovsky, 1990: 193, 211; Figs 2A(1-6), 6. [25°04'S 97°28'W, 267-280 m] 

Sicyonia ocellata Stimpson, 1860a: 43. [Portu ”Hong Kong”; in mari Sinensi quoque, lat. bor. 24°; ad prof. 

20 org] 

= Sicyonia ommanneyi Hall, 1961: 110; Plate 21, figs 26-27. [Malaysia, S of Singapore, 45 fms] 

Sicyonia olgae Pérez Farfante, 1980b: 775; Figs 1-3. [Oregon stn 2277, Off Parimaribo, Suriname, 06°37'N 

55°36'W, 35 m] 

Sicyonia parafallax Crosnier, 1995: 193; Figs 3-4. [Mer Rouge, détroit de Bab el Mandeb, 12°43,7'N 

43°15,0'E, 228 m] 

Sicyonia parajaponica Crosnier, 2003: 226; Figs 10-12; 106E-F. [MUSORSTOM 1, stn 7, Philippines, dans 

l’ouest de Luçon, 14°01,0'N, 120°20,0'E, 185-200 m] 

Sicyonia parri (Burkenroad, 1934a) 

= Eusicyonia parri Burkenroad, 1934a: 80; Fig. 22. [Crooked Island, Bahamas] 

Sicyonia parvula (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte parvulus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 6. [Japan] 

Sicyonia penicillata Lockington, 1878a: 164. [Bolinas Bay, Lower California, 14 fms; Angeles Bay, Gulf of 

California] 

Sicyonia picta Faxon, 1893: 210. [Albatross stns 3387 (off Golfo de Panamá, 7°40'00"N 79°17'50"W, 127 fms); 

3355 (off Punta Mariato, Panamá, 7°12'20"N 80°55'00"W, 182 fms] 

Sicyonia rectirostris De Man, 1907a: 141. [Siboga stn 139, Sanana bay, E coast of Sula Besi, Indonesia, 22 m] 

Sicyonia robusta Crosnier, 2003: 323; Figs 99-102, 110C, D. [CORAIL 2, stn DW 166, Îles Chesterfield, 

19°41,49'S 158°25,24'E, 56 m] 

Sicyonia rocroi Crosnier, 2003: 280; Figs 60-62. [Mutu One, Hatutaa, Îles Marquises, 7°51,7'S 140°30,6'W, 

416-460 m] 

Sicyonia rotunda Crosnier, 2003: 290; Figs 68-69, 108E-F. [SMIB 5, stn DW 84, Nouvelle-Calédonie, 

22°20,8'S 168°43,1'E, 290 m] 

Sicyonia stimpsoni Bouvier, 1905a: 748. [Blake stn 273, off Barbados, 13°03'05"N 59°36'18"W, 103 fms; lectotype 

designation by Holthuis, 1959a] 

Sicyonia taiwanensis Crosnier, 2003: 282; Figs 63-64, 108B. [au large de Tai-Chi, Taiwan] 

Sicyonia trispinosa De Man, 1907a: 142. [Siboga stn 37, Pulau Sailus ketjil, Paternoster Islands, Indonesia, 

27 m] 

Sicyonia truncata (Kubo, 1949a) 

= Eusicyonia truncata Kubo, 1949a: 456; Figs 8M, 48F, 77A, G, 79J, 156E, 158. [Kumanonada, off 

Owase, Mie Prefecture, Japan, 350 m] 

Sicyonia typica (Boeck, 1864) 

= Synhimantites typicus Boeck, 1864: 189. [Norge] 

= Palæmon carinatus Olivier, 1811: 667; nec Cancer carinatus Brünnich, 1768. [il se trouve sur les côtes 

de la Nouvelle-Hollande] 

= Sicyonia Edwardsii Miers, 1881a: 367. [nomen novum for Palæmon carinatus Olivier, 1811] 

Sicyonia vitulans (Kubo, 1949a) 

= Eusicyonia vitulans Kubo, 1949a: 448; Figs 8Q, 48G, 77F, L, 79I, 154, 156F. [Kumanonada, off Owase, 

Mie Prefecture, Japan, ca. 350 m] 

Sicyonia wheeleri Gurney, 1943: 1; Figs 1-13. [Bermuda; description of adult in Burkenroad, 1946 as Sicyonia 

wheeleri sp. nov.]


Family SoleNoCeRIDAe Wood-Mason in Wood-Mason & Alcock, 1891a 

Cryptopenaeus de Freitas, 1979 

= Cryptopenaeus de Freitas, 1979 (type species Cryptopenaeus catherinae de Freitas, 1979, by original 


= Crassipenaeus Liu & Zhong, 1983 (type species Crassipenaeus sinensis Liu & Zhong, 1983, by original 

designation and monotypy, gender masculine) 

Cryptopenaeus brevirostris Hayashi in Baba, Hayashi & Toriyama, 1986: 39; Plate 1; fig. 17. [Bungo Strait, 

Japan, 360-380 m] 

Cryptopenaeus catherinae de Freitas, 1979: 125; Fig. 1A-I. [Off Cape Santa Maria, southern Mozambique, 

26°06'S 33°08'E, 350 m] 

Cryptopenaeus clevai Crosnier, 1985: 26; Figs 1a-b, 2a-e, 3a. [CORINDON IV stn IV 1 (03°28.3'N 128°24.8'E 

– 03°30.0'N 128°23.0'E), 400-300 m] 

Cryptopenaeus crosnieri Pérez Farfante & Kensley, 1985: 281; Figs 1-2. [R/V Kapala stn 78-05-07, NE of 

North Solitary Island, New South Wales, Australia, 29°47-49'S 153°41'E, 234 m] 

Cryptopenaeus sinensis (Liu & Zhong, 1983) 

= Crassipenaeus sinensis Liu & Zhong, 1983: 171; Fig. 1. [North-Eastern Guangdong near Shantou 

(Swatow), 261 m] 

Gordonella Tirmizi, 1960 

= Gordonella Tirmizi, 1960 (type species Gordonella polyarthra Tirmizi, 1960, by monotypy, gender 

feminine) 

Gordonella kensleyi Crosnier, 1988a: 597; Figs 2e, 12b, 15h-i, 16g. [BIOCAL stn CP 23, Nouvelle-Calédonie, 

22°46'N 166°20'E, 2040 m] 

Gordonella paravillosa Crosnier, 1988a: 589; Figs 2d, 3c, 12a, 13-14, 15a-e, 16a-f. [Cidaris I stn 18-1, côte est 

de l’Australie, 17°19.58'S 147°47.61'E, 1147-1132 m] 

Gordonella villosa (Alcock & Anderson, 1894) 

= Haliporus villosus Alcock & Anderson, 1894: 146. [Investigator stns 121 Laccadive Sea, 14°35'15"N, 

1140 fms; 127, Laccadive Sea, off the Island of Minnikoy, 1200 fms] 

= Gordonella polyarthra Tirmizi, 1960: 373, Figs 86-96. [John Murray Expedition stn 135, southern 

Arabian Sea, 4°47'42"N 72°35'36"E, 2727 m] 

Hadropenaeus Pérez Farfante, 1977b 

= Hadropenaeus Pérez Farfante, 1977b (type species Hymenopenaeus modestus Smith, 1885b, by original 


Hadropenaeus affinis (Bouvier, 1906d) 

= Haliporus affinis Bouvier, 1906d: 4. [capturés par le Talisman aux îles du Cap Vert, sur des fonds de 

100 et de 410 mètres; un spécimen provient des Barbades où il fut pris par le Blake à 82 brasses de 

profondeur." = Talisman stn 116, off Cape Verde Islands, 16°53'N 25°10'W, 410-460 m; uncited Talisman 

stn, off Cape Verde Islands, ”100 m”; Blake stn 273, off Barbados, 13°03'05"N 59°36'18"W, 188 m 

according to Pérez Farfante, 1977b] 

Hadropenaeus lucasii (Spence Bate, 1881) 

= Solenocera Lucasii Spence Bate, 1881: 185. [south of New Guinea, about 130 fms; according to 

Spence Bate, 1888 (as Philonicus lucasii): Challenger stn 192, off the Ki Islands, south of Papua, 

5°49'15"S 132°14'15"E, 140 fms] 

= Haliporus malhaensis Borradaile, 1910: 258; Plate 16, fig. 2. [off Saya de Malha, Indian Ocean, 145 

fms] 

Hadropenaeus modestus (Smith, 1885b) 

= Hymenopenaeus modestus Smith, 1885b: 183. [Fish Hawk stn 1047, off Bethany Beach, Delaware, 

38°31'N 73°21'W, 156 fms] 

Hadropenaeus spinicaudatus Liu & Zhong, 1983: 174; Fig. 2. [Eastern Guangdong, off shore (19°59.9'N 

113°24.0'E), 140 m]


Haliporoides Stebbing, 1914b 

= Haliporoides Stebbing, 1914b (type species Haliporoides triarthrus Stebbing, 1914b, by monotypy, 


= Parahaliporus Kubo, 1949a (type species Haliporus sibogae De Man, 1907a, by original designation, 


Haliporoides cristatus Kensley, Tranter & Griffin, 1987: 265; Figs 1, 2, 5G-L. [NE of Point Danger, Queensland, 

Australia] 

Haliporoides diomedeae (Faxon, 1893) 

= Peneopsis diomedeae Faxon, 1893: 212. [Albatross stns 3384 (off Golfo de Panamá, 7°31'30"N 

79°14'00"W, 458 fms); 3395 (SW of Golfo de Panamá, 7°30'36"N 78°39'00"W, 730 fms); 3394 (SE of 

Golfo de Panamá, 7°21'N 79°35'W, 511 fms); 3393 (off Punta Mala, Panama, 7°15'N 79°36'W, 1020 

fms); 3353 (off Punta Mariato, Panama, 7°06'15"N 80°34'00"W, 695 fms); 3358 (S of Península de 

Azuero, 6°30'N 81°44'W, 555 fms)] 

Haliporoides sibogae australiensis Kensley, Tranter & Griffin, 1987: 269; Figs 3, 4, 5A-F. [E of Broken Bay, 

New South Wales, Australia] 

Haliporoides sibogae madagascariensis Crosnier, 1978: 102; Figs 36b, 37a-e, 38a-c, 39b, 40b, 41b, 42b. [Vauban 

CH 5, Madagascar, 12°44.8'S 48°10.6'E, 570-563 m] 

Haliporoides sibogae sibogae (De Man, 1907a) 

= Haliporus sibogae De Man, 1907a: 138. [Siboga stns 38, 7°35.4'S 117°28.6'E, 521 m; 74, 5°3.5'S 119°0'E, 

450 m; 212, 5°54.5'S 120°19.2'E, 462 m; 256, 5°26.6'S 132°32.5'E, 397 m] 

Haliporoides triarthrus triarthrus Stebbing, 1914b 

= Haliporoides triarthrus Stebbing, 1914b: 21. [East London NW. 1/2N., 18 miles, (Cape Colony), 250- 

300 fms] 

Haliporoides triarthrus vniroi (Crosnier, 1978) 

= Hymenopenaeus triarthrus vniroi Crosnier, 1978: 103; Figs 36c, 38e. [Van Gogh stn 264, Mozambique, 

25°28'S 33°31'E, 410 m] 

Haliporus Spence Bate, 1881 

= Haliporus Spence Bate, 1881 (type species Haliporus curvirostris Spence Bate, 1881, designated by 


Haliporus curvirostris Spence Bate, 1881: 185. [mid Pacific, 2375 fms] 

Haliporus taprobanensis Alcock & Anderson, 1899: 280. [Investigator stn 219, Gulf of Manaar and Cape 

Comorin, southern India, 550 fms)] 

= Hymenopenaeus kannemeyeri Kensley, 1977: 16, 27; Fig. 7. [Meiring Naude stn 38, off South Africa, 

28°21.9'S 32°34.6'E, 775-825 m] 

Haliporus thetis Faxon, 1893: 214. [Albatross stn 3413 (off the Galapagos Islands, 2°34'00"N 92°06'00"W, 

1360 fms)] 

Hymenopenaeus Smith, 1882 

= Hymenopenaeus Smith, 1882 (type species Hymenopenaeus debilis Smith, 1882, by monotypy, gender 


Hymenopenaeus aphoticus Burkenroad, 1936a: 112; Figs 62, 65-67. [Pawnee stn 54, Turks Island Passage, 

21°15'40"N 71°17'06"W, 900-945 fms] 

Hymenopenaeus chacei Crosnier & Forest, 1969: 545; Figs 1, 2a-b. [Ombango stn 406, au large du plateau 

continental du Gabon, 8°35'S 12°51'E, 545-555 m] 

Hymenopenaeus debilis Smith, 1882: 91; Plate 15, figs 6-11; Plate 16, figs 1-3. [Blake stns 317, SE of Savannah 

Beach, Georgia, 31°57'00"N 78°18'35"W, 333 fms; 323, SE of Cape Fear, North Carolina, 33°19'00"N 

76°12'30"W, 457 fms; 326, E of Cape Fear, North Carolina, 33°42'15"N 76°00'50"W, 464 fms] 

= Hymenopenaeus debilis var. africanus Bouvier, 1908a: 83. [Type locality not indicated; according to 

Crosnier & Forest, 1973: Talisman stn 21, 33°46'N 9°02'W, 1319 m] 

Hymenopenaeus doris (Faxon, 1893) 

= Haliporus doris Faxon, 1893: 214. [Albatross stns 3414 (off Cabo Velas, Costa Rica, 10°14'N 96°28'W, 

2232 fms); 3415 (S of Punta Maldonado, Guerrero, Mexico, 14°46'N 98°40'W, 1879 fms)]


Hymenopenaeus equalis (Spence Bate, 1888) 

= Haliporus equalis Spence Bate, 1888: 285; Plate 41, fig. 1. [Challenger stn 200, 6°47'N 122°28'E, between 

the Philippine Islands and Borneo, 250 fms] 

Hymenopenaeus fallax Crosnier & Dall, 2004: 10; Figs 6-8. [Albatross stn 4106, Kaiwi Channel, 335-350 fms] 

Hymenopenaeus fattahi Ramadan, 1938: 60; Fig. 8. [John Murray Expedition stn 54 (21°58'00''N 62°19'42''E 

to 21°58'36''N 62°21'24''E), South Arabian coast, 1046 m] 

Hymenopenaeus furici Crosnier, 1978: 127; Figs 39f, 40f, 42f, 43d, 46f-h. [Vauban CH 133, Madagascar, 

13°02'S 48°02'E, 1000-1525 m] 

Hymenopenaeus halli Bruce, 1966a: 216; Figs 1-2. [Cape St. Mary Cruise 1/64, stn 26, South China Sea, 

19°22.5'N 114°07.5'E to 19°22.0'N 114°11.0'E, 400-435 fms] 

Hymenopenaeus laevis (Spence Bate, 1881) 

= Haliporus laevis Spence Bate, 1881: 185. [mid Atlantic, 2500 fms] 

= Hymenopenaeus microps Smith, 1884: 413; Plate 10, fig. 1. [Albatross stns 2076, E of Georges Bank, Massachusets, 

41°13'00"N 60°00'50"W, 906 fms; 2037, off New Jersey, 38°50'00"N 69°23'30"W, 1731 fms] 

= Haliporus androgynus Bouvier, 1906e: 253. [Talisman stns 101, entre Dakar et la Praya, 16°38'N 

20°44'W, 3200 m; 102, entre Dakar et la Praya, 15°48'N 20°23'W, 3655 m] 

Hymenopenaeus methalli Crosnier & Dall, 2004: 17; Figs 11-13, 14B, C. [MUSORSTOM 5, stn 384, 19°42.40'S 

158°50.80'E, 756-772 m] 

Hymenopenaeus neptunus (Spence Bate, 1881) 

= Haliporus neptunus Spence Bate, 1881: 185. [among the Celebes Islands, about 600 fms; according 

to Spence Bate, 1888: Challenger stns 191, off the Arrou Islands, 5°41'S 134°4'30"E, 800 fms; 196, near 

the Philippines, 0°48'30"S 126°58'30"E, 825 fms] 

Hymenopenaeus nereus (Faxon, 1893) 

= Haliporus nereus Faxon, 1893: 213. [Albatross stns 3366 (S of Cabo Blanco, Costa Rica, 5°30'N 

86°45'W, 1067 fms); 3353 (S of Morro de Puercos, Panama, 7°06'15"N 80°34'00"W, 695 fms); 3382 (off 

Pen de Azuero, Panama, 6°21'N 80°41'W, 1793 fms); 3413 (NW of Galapagos Islands, Ecuador, 

2°34'N 92°06'W, 1360 fms); 3398 (NW of Punta Galera, Ecuador, 1°07'N 80°21'W, 1573 fms); 3399 

(NW of Punta Galera, Ecuador, 1°07'N 81°04'W, 1740 fms); 3400 (E of Galapagos Islands, Ecuador, 

00°36'N 86°46'W, 1322 fms); 3407 (Galapagos Islands, Ecuador, 00°04'00"S 90°24'30"W, 885 fms)] 

Hymenopenaeus obliquirostris (Spence Bate, 1881) 

= Haliporus obliquirostris Spence Bate, 1881: 186. [off Kermadec Island; according to Spence Bate, 

1888: Challenger stn 170, off the Kermadec Islands, 29°55'S 178°14'W, 520 fms] 

Hymenopenaeus propinquus (De Man, 1907a) 

= Haliporus propinquus De Man, 1907a: 140. [Siboga stns 80, Pulu Kaniungan Ketjil, Indonesia, 11 m; 

178, 2°40'S 128°37.5'E, 835 m; 316, 7°19.4'S 116°49.5'E, 538 m] 

Hymenopenaeus sewelli Ramadan, 1938: 58; Figs 7a-e. [John Murray Expedition stn 156 (4°44'30''N 

72°46'00''E to 4°41'12''N 72°42'48''E), Maldive area, 1828 m] 

Hymenopenaeus tuerkayi Crosnier, 1995: 189; Figs 1-2. [Golfe d’Aden central, 12°56,7'N 47°47,0'E-12°55,9'N 

47°47,4'E, 2276-2282 m] 

Mesopenaeus Pérez Farfante, 1977b 

= Mesopenaeus Pérez Farfante, 1977b (type species Parartemesia tropicalis Bouvier, 1905a, by original 


Mesopenaeus brucei Crosnier, 1986b: 20; Figs 1a-f, 2a-b. [Soela Cruise 184 stn 55, Australie, 17°41.2'S 

118°42.5'E, 354-360 m] 

Mesopenaeus mariae Pérez Farfante & Ivanov, 1982: 303; Figs 1-7. [Professor Mesystsev haul 135, SE of Saya 

de Malha Bank, Indian Ocean, 11°27'36"S 61°37'36"E, 158-160 m] 

Mesopenaeus tropicalis (Bouvier, 1905a) 

= Parartemesia tropicalis Bouvier, 1905a: 748. [de la mer des Antilles où elle a été prise par le Blake par 

des profondeurs de 80 à 175 brasses] 

= Solenocera weymouthi Lindner & Anderson, 1941: 181; Fig. 1a-e. [Pelican stn 137, 29°28'N 87°30'W, 

46 fms]


Pleoticus Spence Bate, 1888 

= Philonicus Spence Bate, 1888 (type species Philonicus mülleri Spence Bate, 1888, designated by 


= Pleoticus Spence Bate, 1888 (nomen novum for Philonicus Spence Bate, 1888, a junior homonym of 

Philonicus Loew, 1849 (Diptera)) 

= Parartemesia Bouvier, 1905a (type species Parartemesia carinata Bouvier, 1905a, designated by Fowler, 

1912, gender feminine) 

= Faxonia Bouvier, 1905b (type species Penaeopsis [sic] ocularis Faxon, 1895, designated by Fowler, 


Pleoticus muelleri (Spence Bate, 1888) 

= Philonicus mülleri Spence Bate, 1888: 275; Plate 39, figs 1-2. [Challenger stn 321, off Montevideo, 

Uruguay, 35°02'N 55°15'W, 13 fms] 

= Parartemesia carinata Bouvier, 1905a: 748. [capturée par le Hassler..” ”..au large de Montevideo, par 

7 et 44 brasses de profondeur.”; off mouth Río de la Plata, 35°42'S 56°20'W, 44 fms (80 m); see Pérez 

Farfante, 1977b] 

Pleoticus robustus (Smith, 1885b) 

= Hymenopenaeus robustus Smith, 1885b: 180. [Albatross stn 2125, S of Curaçao, 11°43'00"N 69°09'30"W, 

208 fms] 

= Peneopsis ocularis Faxon, 1895: 187. [I have examined with some care a specimen in this Museum 

[Museum of Comparative Zoology, Harvard] from the ”Blake” collection, labelled ”Peneopsis ocularis” 

by A. Milne-Edwards] 

Pleoticus steindachneri (Balss, 1914c) 

= Haliporus Steindachneri Balss, 1914c: 135. [Pola stns 9, 23°21'N 37°37'E, 791 m; 20, 23°20'N 36°20'E, 

780 m; 47, 23°41'N 38°9'E, 610 m; 59, 25°43'N 36°10'E, 780 m; 61, 24°35'N 36°51'E, 828 m; 66, 26°8'N 

35°27'E, 1168 m; 107, 20°27'N 38°18'51"E, 748 m; 109, 21°19'N 37°39'E, 890 m; 110, 21°7'N 37°28'E, 

635 m; 114, 19°38'N 37°55'E, 535 m; 117, 20°16'N 37°33'E, 638 m; 120, 19°21'N 38°29'E, 560 m; 121, 

18°51'N 39°5'E, 690 m; 143, 17°7'N 39°55'E, 212 m; 156, 22°51'N 38°2'E, 712 m; 170, 27°2'N 35°17'E, 

690 m; 178, 26°19'N 34°24'E, 720 m; full description and list of syntypes in Balss, 1915] 

Solenocera lucas, 1849 

= Solenocera Lucas, 1849 (type species Solenocera Philippii Lucas, 1849, by monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in Opinion 611 in 1961) 

= Parasolenocera Wood-Mason in Wood-Mason & Alcock, 1891a (type species Parasolenocera annectens 

Wood-Mason in Wood-Mason & Alcock, 1891a, by monotypy, gender feminine) 

= Transolenocera Burkenroad, 1934b (type species Solenocera (Parasolenocera) maldivensis Borradaile, 

1910, by monotypy, gender feminine) 

Solenocera acuminata Pérez Farfante & Bullis, 1973: 8; Figs 1c, 2, 4-6, 18, 19. [Oregon stn 2022, N of Rivière 

Organabo, French Guiana, 07°15'N 53°35'W, 210 m] 

Solenocera africana Stebbing, 1917b: 32; Plate 93A. [Sebastian Bluff NW. 3/4W., 8 miles, South Africa, 34 

fms] 

? = Solenocera membranacea capensis Heegaard, 1966: 32; Figs 15-21, 23. [Discovery stns 100B, 33°20'S- 

33°46'S 15°08'E – 15°18'E, 5-0 m; 102, 25°29'30"S 18°33'40"E, 50-0 m; 260, 33°06'30"S 17°45'15"E, 100-0 

m; 277, 1°44'00"S 8°38'00"E, 63 m; 89, 34°05'15"S 16°00'45"E, 50-0 m; 99A, 33°20'00"S 17°17'00"E, 5-0 

m; 99E, 33°11'00"S 17°26'00"E, 5-0 m; 276, 5°54'00"S 11°19'00"E, 150 m] 

Solenocera agassizii Faxon, 1893: 211. [Albatross stns 3389 (off Panama, 7°16'45"N 79°56'30"W, 210 fms); 

3391 (off Panama, 7°33'40"N 79°43'20"W, 153 fms) 

Solenocera alfonso Pérez Farfante, 1981: 631; Figs 1-5. [off Capitancillo Island, W of Leyte, Philippines] 

= Solenocera alfonso forma inermis Crosnier, 1989: 56; Figs 3e-f, 5f-h, 6a, e. [many locations in the 

Philippines, 175-200 m; unavailable under Art. 15.2] 

Solenocera algoensis Barnard, 1947 

= Solenocera algoense Barnard, 1947: 383. [E portion of Algoa Bay, South Africa, 50 fms] 

= Solenocera ramadani Ivanov & Hassan, 1976c: 242; Figs 1-2. [Van Gogh stn 269, off E Africa, 29°31'S 

34°42'E, 285-305 m]


Solenocera alticarinata Kubo, 1949a: 227; Figs 8W, 45E, 72P, V, 80F, 93, 94A-C, 100. [Tainan, Formosa] 

Solenocera annectens (Wood-Mason in Wood-Mason & Alcock, 1891a) 

= Parasolenocera annectens Wood-Mason in Wood-Mason & Alcock, 1891a: 276. [Investigator stn 116 

(Andaman Sea, 11°25'5"N 92°47'6"E), 405 fms] 

Solenocera atlantidis Burkenroad, 1939: 10; Figs 5-10. [Atlantic stn 2813, off Mobile Bay, Alabama, 29°45'N 

88°11'W, 19 fms] 

Solenocera australiana Pérez Farfante & Grey, 1980: 422; Figs 1-7. [Apache cruise 21, haul 3, N of Groote 

Eylandt, Gulf of Carpentaria, Northen Territory, Australia, 13°34'S 136°30'E, 22 m] 

Solenocera barunajaya Crosnier, 1994c: 355; Figs 1a-c, 2a, 3a-c. [KARUBAR stn CP 83, Iles Tanimbar Island, 

Indonésie, 9°23'S 131°00'E, 285-297 m] 

Solenocera bedokensis Hall, 1962: 13; Figs 78-78c. [Off Bedok Village, Singapore, 17 fms] 

Solenocera bifurcata Dall, 1999: 574; Fig. 14A-F. [Off Cape Moreton, SE Queensland, Australia] 

Solenocera burukovskyi Timofeev, 1993: 37; Fig. 1. [Gulf of Aden, 12°19'4N 44°21'7E, 470-475 m] 

Solenocera choprai Nataraj, 1945: 91, Figs 1-4. [Arabian Sea, 17°27'N 71°41'E, 56-58 fms] 

Solenocera comata Stebbing, 1915 

= Solenocera comatum Stebbing, 1915: 67; Plates 13-14. [Off East London, E coast of South Africa] 

= Solenocera novae-zealandiae Borradaile, 1916: 79; Fig. 1. [7 miles E of North Cape, off New Zealand, 

70 fms] 

= Solenocera brevipes Kubo, 1949a: 246; Figs 1S, 8X, 20N, 27F-H, 45D, 66I-J, 72Q, W, 80A, 98H-J, 99, 

100. [Komanonada, off Owase, Mie Prefecture, Japan, about 300 m] 

Solenocera crassicornis (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

? = Penaeus planicornis Fabricius, 1798: 409. [in Oceano Indico] 

= Penaeus crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 418. [côtes de l’Inde] 

= Solenocera sinensis Yü, 1937: 112; Figs 1-5. [Amoy, Tinghai] 

= Solenocera indicus Nataraj, 1945: 94; Figs 5-8. [Cocanada Bay, Bay of Bengal] 

= Solenocera subnuda Kubo, 1949a: 255; Figs 8T, 20O, 27I-J, 45C, 66C, D, 72M, S, 80D, 100, 102D-G, 

103. [S coast of Borneo, Indonesia] 

= Solenocera kuboi Hall, 1956: 69; Plate 9, Figs 2-3. [Off Tanjong Stapa, Malaysia, 6-7 fms] 

Solenocera faxoni De Man, 1907a: 136. [Siboga stn 254, 5°40'S 132°26'E, 310 m] 

Solenocera florea Burkenroad, 1938: 64; Text-Figs 6-7. [Pearl Island, Gulf of Panama, 8°29'40"N 78°52'30"W, 

19-24 fms] 

Solenocera geijskesi Holthuis, 1959a: 56; Figs 4-5. [Surinam, 20 miles off the coast, between mouths of 

Nickerie and Coppename Rivers, 27 m] 

Solenocera gurjanovae Starobogatov, 1972: 365; Plate 3, fig. 13. [Gulf of Tonkin, Vietnam, 50 m] 

Solenocera halli Starobogatov, 1972: 412; Plate 2, fig. 7. [Andaman Sea, off India, 6°05.8' – 6°02'N 98°59' – 

99°00.5'E] 

Solenocera hextii Wood-Mason & Alcock, 1891b: 188. [Investigator stns 81 (24 miles SE of Gapalpur, Ganjam 

Coast, 89-93 fms); 96 (Bay of Bengal, 18°30'N 84°46'E, 98-102 fms), 89-102 fms; off Chittagon, 65 

fms; off the Mahánadi Delta, 68 fms; off the Godávari Delta, 70 fms] 

Solenocera koelbeli De Man, 1911a: 45, 48, 50, 51. [Japan] 

= Solenocera depressa Kubo, 1949a: 237; Figs 8R, 27T-V, 45B, 66E-F, 72O, U, 80G, 96, 98A-D, 100. 

[Komanonada, off Owase, Mie Prefecture, Japan] 

= Solenocera vietnamensis Starobogatov, 1972: 363, 384; Plate 2, fig. 6a-c. [Pelamida, stn 39, Tonkin 

Gulf, Vietnam] 

Solenocera maldivensis Borradaile, 1910 

= Solenocera (Parasolenocera) maldivensis Borradaile, 1910: 258; Plate 16, fig. 1. [Funadu Velu, Miladumadulu 

Atoll, Maldive Islands] 

Solenocera mascarensis Burukovsky, 1993a: 23; Fig. 1.3-1.7. [Sala-de-Malya Bank, 11°11'1''S 62°37'E, 2020- 

2000 m] 

Solenocera melantho De Man, 1907a: 137. [Siboga stns 302, 10°17.9'S 123°28.7'E, 216 m; 306, 8°27'S 

122°54.5'E, 247 m; 312, Saleh bay, N coast of Sumbawa, Indonesia, 8°19'S 117°41'E, 274 m] 

= Solenocera prominentis Kubo, 1949a: 231; Figs 8V, 14C, 16C, 20M, 27Q-S, 45A, 66G, H, 72F, L, 80E, 

94D-G, 95, 100. [Komanonada, off Owase, Mie Prefecture, Japan, about 350 m]


Solenocera membranacea (Risso, 1816) 

= Peneus membranaceus Risso, 1816: 98. [Président Théodore Tissier stn J.376, France, Ligurian Sea between 

Corsica and Nice, 42°56.5'N 9°37.2'E, 310-415 m; neotype designation by Holthuis, 1961b] 

= Peneus siphonoceros Philippi, 1840: 190; Plate 4, fig. 3. [Neapel] 

= Solenocera philippii Lucas, 1849: 300. [baie d’Alger] 

= Penaeus distinctus De Haan, 1849 [in De Haan, 1833-1850]: 194. [Japan; erroneous locality, see 

Yamaguchi & Baba, 1993] 

Solenocera moosai Crosnier, 1985: 37; Figs 5a, 6a, 7c,d, h, i. [CORINDON II stn 273 (01°56.0'S 119°16.0'E), 

120-200 m] 

Solenocera mutator Burkenroad, 1938: 61; Text-Figs 2-5. [Templeton Crocker Expedition stn 150, from Gorda 

Banks, tip of Lower California, 23°01'30"N 109°30"W, 40-100 fms] 

Solenocera necopina Burkenroad, 1939: 7; Figs 1-4. [Atlantis stn 2377, off Mobile Bay, Alabama, 29°16'N 

87°54'W, 125 fms] 

Solenocera pectinata (Spence Bate, 1888) 

= Philonicus pectinatus Spence Bate, 1888: 279; Plate 38. [Challenger stn 188, Arafura Sea, south of 

Papua, 9°59'S 139°42'E, 28 fms] 

= Philonicus cervicalis Zehntner, 1894: 210; Plate 9, fig. 26. [Amboine] 

Solenocera pectinulata Kubo, 1949a: 251; Figs 8S, 27A-B, 66 K-L, 72N-T, 83B, 101, 102A-C. [Komanonada, 

off Owase, Mie Prefecture, Japan, about 350 m] 

? = Solenocera utinomii Kubo, 1951: 263; Fig. 4. [Off Kii Peninsula, Japan, ca. 150 m] 

Solenocera phuongi Starobogatov, 1972: 366; Plate 3, fig. 12a-b. [Pelamida stn 9, Tonkin Gulf, 113 m] 

Solenocera rathbuni Ramadan, 1938: 57 (partim, nec Fig. 6 = S. algoensis Barnard, 1947). [Albatross stn 3987, 

Kauai Island, Hawai’i, 50-55 fms; lectotype designation by Crosnier, 1978] (Fig. 5) 

Solenocera spinajugo Hall, 1961: 81; Plate 17, figs 1-3. [N of the Malacca Strait, 120 miles NW of Penang, 

41 fms] 

Solenocera vioscai Burkenroad, 1934b: 65; Figs 1-4. [About 5 miles off Pass à la L’outre, Louisiana, USA, 15 

fms] 

Solenocera waltairensis M.J. George & Muthu, 1970: 292; Figs 1-4. [Waltair, east coast of India, 17°43'N 

83°23'E, 20 m] 

Solenocera zarenkovi Starobogatov, 1972: 367, Plate 3, fig. 14a-b. [Orlik, stn 18, Tonkin Gulf] 

Fig. 5. Solenocera rathbuni Ramadan, 1938. Photo by Tin-Yam Chan.


Superfamily SeRGeSToIDeA Dana, 1852a 

Family luCIFeRIDAe De Haan, 1849 [in De Haan, 1833-1850] 

Lucifer J.V. Thompson, 1829 (Fig. 6) 

= Lucifer J.V. Thompson, 1829 (type species Leucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 

1834-1840], by subsequent indication under Article 68b in H. Milne Edwards, 1834-1840, 


= Leucifer H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] [incorrect subsequent spelling 

of Lucifer] 

Lucifer chacei Bowman, 1967: 266; Figs 1-4. [Eniwetok Atoll, Marshall Islands, lagoon 4.8 km W of Parry 

Island, 3 m] 

Lucifer faxoni Borradaile, 1915a: 228, 230. [off Chesapeake Bay, U.S.A.; lectotype designation by Holthuis, 

1959a] 

= Lucifer affinis Borradaile, 1915a: 228, 230. [Floridastrom, J.N., 56, N.E. of Bermuda, roughly 35°N 

60°W; lectotype designation by Holthuis, 1959a] 

Lucifer hanseni Nobili, 1905a 

= Lucifer Hanseni Nobili, 1905a: 395. [Djibouti] 

= Lucifer inermis Borradaile, 1915a: 229. [Melbourne Harbour, Australia] 

Lucifer intermedius Hansen, 1919: 57; Plate 4, figs 8a-b; Plate 5, figs 1a-g. [Siboga stns 7, 7°55'.5S 114°26'E, 

15 m and more; 35, 8°0'.3S 116°59'E, 1310 m, surface; 37, Sailus ketjil, Paternoster-islands, 27 m and 

less, surface; 66, bank between islands of Bahuluwang and Tambolungan, south of Saleyer, 8-10 m; 

75, 4°57'.4S 119°2'.8E, from 11 m to surface; 81, Pulu Sebangkatan, Borneo-bank, 34 m; 106, Anchorage 

of Kapul-island, Sulu-archipelago, 13 m; 109, Anchorage off Pulu Tongkil, Sulu-archipelago, 13 

m; 117a, 1°15'N 123°37'E; ?132, about 5°56'.7N 126°25'E; 136, Ternate anchorage, 23 m - surface; 138, 

Anchorage on the east coast of Kajoa-island, 66 m - surface; 141, 1°0'.4S 127°25'.3E, from 1500 m; 

144, Anchorage North of Salomakice-(Damar)island, 45 m; 157, 0°32'.9S 130°14'.6E; 177a, 2°30'S 

129°28'E; 189a, 2°22'S 126°46'E] 

Lucifer orientalis Hansen, 1919: 55; Plate 4, figs 7a-g. [Siboga stns 37, Sailus Ketjil, Paternoster-islands, 27 

m and less; 66, bank between islands of Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81, 

Pulu Sebangkatan, Borneo-bank, 34 m; 104, Sulu-harbour, Sulu-island, 14 m; 106, anchorage off 

Fig. 6. Lucifer spec. Photo by Arthur Anker.


Kapul-island, Sulu-archipelago, 13 m; 133, anchorage off Lirung, Salibabu-island, depth up to 36 m; 

136, Ternate anchorage, 23 m - surface; 138, anchorage on the E coast of Kajoa-island, 66 m - surface; 

140, Bay of Batjan, 13 m; 141, 1°0'.4S 127°25'.3E, 1500 m - surface; 143, 1°4'.5S, 127°52'.6E, 1000 m - 

surface; 148, 0°17'.6S 129°14'.5E, 1000 m - surface; 157, 0°32'.9S 130°14'.6E, 45 m; 177a, 2°30'S 

129°28'E; 189a, 2°22'S 126°46'E; 194-197, 1°53'.5-1°45'.3S 126°39-127°8'.3E; 203, 3°32'.5S 124°15'.5E, 

1500 m - surface; 220, anchorage off Pasir Pandjang, W coast of Binongka, 278 m - surface; 225, N. 

279°E, from Southpoint of South-Lucipara-island, 894 m; 243.4°30'.2S 129°25'E, 1000 m - surface; 

245.4°16'.5S 130°15'.8E, surface; 276, 6°47'.5S 128°40'.5 E, 750 m - surface; near stn 300, between 

Timor and Lomblen] 

Lucifer penicillifer Hansen, 1919: 59; Plate 5, figs 2a-k. [Siboga stns 7, 7°55'.5S 114°26'E, 25 m and more; 16, 

Bay of Kankamaräan, S coast of Kangeang, 22 m; 35, 8°0'.3S 116°59'E, surface; 37, Sailus ketjil, Paternoster-islands, 

27 m and less; 40, anchorage off Pulu Kawassang, Paternoster-islands, 12 m; 66, bank 

between Bahuluwang and Tambolungan, S of Saleyer, 8-10 m; 81, Pulu Sebangkatan, Borneo-bank, 

34 m; 93, Pulu Sanguisiapo, Tawi-Tawi-islands, Sulu-archipelago, 12 m; 96, SE side of Pearl-bank, 

Sulu-archipelago; 98-99, 6°9'-6°7'.5N 120°21'-120°26'E; 99, 6°7'.5N 120°26'E, 16-23 m - surface; 105, 

6°8'N 121°19'E, 275 m; 106, anchorage off Kapul-island, Sulu-archipelago, 13 m; 107, 6°1'.5N. 

121°28'E; 109, anchorage off Pulu Tongkil, Sulu-archipelago, 13 m; 112, 3°1'N 122°2'E; 117 a , 1°15'N 

123°37'E; 125, anchorage off Sawan, Siau-island, 27 m; 128, 4°27'N 125°25'.7E, from 700 m to surface; 

136, Ternate anchorage, 23 m; 138, anchorage on the E coast of Kajoa-island, 66 m; 140, Bay of Batjan, 

13 m; 141, 1°0'.4S 127°25'.3E, from 1500 m to surface; 143, 1°4'.5S 127°52'.6E, from 1000 m to 

surface; 144, anchorage north of Salomakiëe- (Damar-) island, 45 m; 146, 0°36'S 128°32'.7E, 512 m; 

157, 0°32'.9S 130°14'.6E, 45 m; 165, anchorage on NE side of Daram-island, E coast of Misool, 49 m; 

168, anchorage N of Sabuda-island, 63 m; 174, Waru-bay, N coast of Ceram, 18 m; 177 a , 2°30'S 

129°28'E; 184, anchorage off Kampong Kelang, S coast of Manipa-island, 36 m; 185, 3°20' 127°22'.9E, 

from 1536 m to surface; 189 a , 2°22'S 126°46'E; 194, 1°53' 5S 126°39'E; 194-197, 1°53'.5-1°4S'.3S 126°39'- 

127°8'.3E; 203, 3°32'5S 124°15'.5E, from 1500 m to surface; 206, Buton-strait, surface; 220, anchorage 

off Pasir Pandjang, W coast of Binongka, 278 m; 225, 5700 m N 279°E from Southpoint of South- 

Lucipara-island, 894; 243, 4°30'.2S 129°25'E, from 1000 m to surface; 245, 4°16'.5S 130°15'.8E; near stn 

300, between Timor and Lomblen; 315, anchorage E of Sailus Besar, Paternoster-islands, surface] 

Lucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= Leucifer typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [Type locality not 

indicated] 

= Leucifer Reynaudii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 469. [trouvé dans 

l’Ocean indien] 

= Lucifer acestra Dana, 1852b: 671. [Pacific Ocean, 6°30'S 177°E, near Sherson’s Island] 

= Lucifer acicularis Dana, 1852b: 674. [Harbour of Rio de Janeiro] 

= Lucifer Zybrantsii Philippi, 1857: 323. [im Atlantischen Ocean unter 25°N.B. und 22°50'W.L. von 

Greenwich] 

= Leucifer uracanthus Steindachner, 1861: 365. [E mari atlantico] 

= Lucifer bonitensis Borradaile, 1915a: 228, 230. [toutes les mers] 

= Lucifer clausi Borradaile, 1915a: 228, 230. [Messina] 

= Lucifer batei Borradaile, 1915a: 228, 230. [Plankton-Expedition, Floridastrom, J.N. 62, Bermuda; 

lectotype designation by Holthuis, 1959a] 

Family SeRGeSTIDAe Dana, 1852a 

Acetes H. Milne edwards, 1830 

= Acetes H. Milne Edwards, 1830 (type species Acetes indicus H. Milne Edwards, 1830, by original 


Acetes americanus americanus Ortmann, 1893: 39; Plate 2, fig. 2. [Plankton-Expedition der Humbolt-Stiftung, 

Mündung des Tocantins JN 239 (0.6°S 48.1°W, 0 m), 240 (1.6°S 49.2°W, 0-35 m), 241 (1.6°S 

49.2°W, 0-35 m), 243 (0.7°S 48.2°W, 0-13 m), Pl. 106 (0.7°S 48.2°W, 12 m) and 107 (1.6°S 49.2°W, 35 m)]


= Acetes brasiliensis Hansen, 1919: 45; Figs 1-7. [Brazil] 

= Acetes americanus limonensis Burkenroad, 1934a: 99; Figs 37-38. [Mouth of the Sweetwater River, 

Limon Bay, Canal Zone, Panama] 

Acetes americanus carolinae Hansen, 1933 

= Acetes carolinae Hansen, 1933: 31; Figs 1-8. [near the Sea Buoy off Beaufort Inlet, North Carolina] 

Acetes binghami Burkenroad, 1934a: 101; Figs 39-40. [Bella Vista Beach, Panama City] 

Acetes chinensis Hansen, 1919: 41; Plate 4, fig. 3a-b. [33°10'N 129°18'E, 40 fms; Formosa Channel, 25 fms] 

Acetes erythraeus Nobili, 1905a: 393; Fig. 1. [Mer Rouge et Périm; Djibouti, dans la cavité d’une Beroë; 

Abdelkader, près de Massaouah, et Massaouah] 

Acetes indicus H. Milne Edwards, 1830: 351; Plate 11, figs 1-9. [Le Gange] 

= Acetes spiniger Hansen, 1919: 35, 43; Plate 4, fig. 5. [Surabaya, Java, Indonesia; 10 miles off Rangoon; 

Indo-Chinese Sea] 

Acetes intermedius Omori, 1975: 40; Figs 14, 16, 17. [Off Tungkiang, Taiwan] 

Acetes japonicus Kishinouye, 1905: 167; 2 unnumbered Figs [the Bay of Ariake, the Bay of Kojima, off 

Nagasu in Buzen, Japan; off Mokpho, Korea] 

= Acetes dispar Hansen, 1919: 39; Plate 3, figs 5a-f; Plate 4, fig. 1a. [Cheribon, Java, Indonesia; Lem 

Ngob, Gulf of Siam] 

= Acetes cochinensis Rao, 1970: 298; Figs 1-10. [Inshore sea of Cochin, southwest coast of India, 3.5- 

11 m] 

Acetes johni Nataraj, 1949: 139; Fig. 1. [Travancore coast, India] 

Acetes marinus Omori, 1975: 49; Figs 8, 21, 25. [Rio Tocantins, near its junction with Rio Para, Brazil] 

Acetes natalensis Barnard, 1955: 2, 43. [Durban Bay, South Africa] 

Acetes paraguayensis Hansen, 1919: 46; Figs 8-14. [Lagoon at Rio Paraguay near its junction with Rio 

Parana, Paraguay, outlet of Riacho del Oro in Rio de la Plata, Paraguay] 

Acetes serrulatus (Krøyer, 1859) 

= Sergestes serrulatus Krøyer, 1859: 268; Plate 4, fig. 12. [nordligste Kattegat] 

= Acetes insularis Kemp, 1917a: 54; Figs 1f-g, 2c, 3c, 4c, 5b, e, 7c. [Mouth of Rajang R., Sarawak, Borneo] 

Acetes sibogae sibogae Hansen, 1919: 38; Plate 3, figs 4a-h. [Siboga stns 47, Bay of Bima, near south fort, 55 

m; 323, Sangkapura roads, Bawean Island, 12 m] 

= Acetes sibogalis Achuthankutty & M.J. George, 1973: 139; Figs 1-20. [Aroor region of the Cochin 

backwaters] 

= Acetes orientalis Achuthankutty & Ayyappan Nair, 1976: 233; Figs 1-19. [near fish jetty in Mandovi 

estuary, Arabian Sea, Goa, India] 

Acetes sibogae australis Colefax, 1940 

= Acetes australis Colefax, 1940: 341, 345; Figs 1-19. [Australia: Homebush Bay, Port Jackson (Sydney 

Harbour); Folly Point, Middle Harbour, Port Jackson; Clarence River, N coast of New South Wales; 

Tuggerah Lake, 50 miles N of Port Jackson] 

Acetes vulgaris Hansen, 1919: 35; Plate 3, figs 2a-r. [Siboga stns 4, anchorage off Djangkar (Java), 7°42'S 

114°12.6'E, 9 m; 47, Bay of Bima, near south fort, 55 m; 311, Sapeh Bay, E coast of Sumbawa, 0-36 m] 

Allosergestes Judkins & Kensley, 2008 

= Allosergestes Judkins & Kensley, 2008 (type species Sergestes sargassi Ortmann, 1893, by original 


Allosergestes index (Burkenroad, 1940) 

= Sergestes (Sergestes) index Burkenroad, 1940: 41. [Dana stn 3630 II (New Zealand, Auckland, 34°24'S 

178°42.5'E), 3000 m wire out)] 

Allosergestes nudus (Illig, 1914) 

= Sergestes nudus Illig, 1914: 366; Figs 26-28. [Atlantischen Ozean, 32°S 8°W, 1000 m] 

Allosergestes oleseni (Vereshchaka, 2009) comb. nov. 

= Sergestes oleseni Vereshchaka, 2009: 78; Figs 35, 38. [Dana stn 3602-2 (south western Pacific Ocean, 

between Fiji and New Caledonia, 20°00'S 174°29'E, 600 m wire out)]


Allosergestes pectinatus (Sund, 1920) 

= Sergestes pectinatus Sund, 1920: 24; Figs 42-43. [Michael Sars stns 23 (35°32'N 7°7'W, 1215 m); 34 

(28°52'N 14°16'W, 2170 m); 52 (31°24'N 34°47'W); 45 (28°42'N 20°0'W); 49 (29°6'N 25°2'W); 51 

(31°20'N 35°7'W, 3886 m); 53 (34°59'N 33°1'W, 2615-2865 m); 56 (36°53'N 29°17'W, 3239 m); 62 

(36°52'N 39°55'W); 64 (34°44'N 47°52'W); 67 (40°17'N 50°39'W)] 

Allosergestes pestafer (Burkenroad, 1937) 

= Sergestes pestafer Burkenroad, 1937: 318; Figs 1-3. [Templeton Crocker Expedition stn 165 T-3, 145 miles 

N of Clarion Island, Revillagigido Islands, Lower California, E Pacific, 20°36'N 115°07'W, 500 m] 

Allosergestes sargassi (Ortmann, 1893) 

= Sergestes sargassi Ortmann, 1893: 34; Plate 3, fig. 1. [Plankton-Expedition der Humbolt-Stiftung, 

Floridastrom JN 50 (39.4°N 57.8°W, 0-200 m); Sargasso-See JN 94 (31.5°N 45.6°W, 0-400 m), 99 

(31.7°N 43.6°W, 0-400 m), 102 (31.7°N 42.7°W, 0-400 m), 110 (30.3°N 37.9°W, 0-400 m), 113 (29.8°N 

36.8°W, 0-400 m), 117 (28.3°N 34.3°W, 0-400 m), 127 (24.6°N 31.0°W, 0-400 m), 262 (23.7°S 36.0°W, 0 

m), 263 (25.6°S 34.9°W, 0-400 m), Pl. 58 (25.1°N 31.5°W, 200 m); Nördl. Aequatorialstrom JN 135 

(18.9°N 26.4°W, 0-400 m), 260 (20.4°N 37.8°W, 0-400 m), Pl. 64 (16.1°N 23.1°W, 200 m); Südl. Aequatorialstrom 

Pl. 76 (0.1°S 15.2°W, 200 m)] 

Allosergestes verpus (Burkenroad, 1940) 

= Sergestes (Sergestes) verpus Burkenroad, 1940: 40. [Dana stn 3739 IX (Celebes Sea, 3°20'N 123°50'E), 

2000 m wire out); according to Vereshchaka, 2009, the holotype label indicates that the specimen 

was collected at Dana stn 3844-6 (Indo-West Pacific, 12°05'S 96°45'E, 600 m wire out)] 

Allosergestes vinogradovi (Vereshchaka, 2009) comb. nov. 

= Sergestes vinogradovi Vereshchaka, 2009: 90; Figs 35, 44. [Dana stn 3902-2, Indo-West Pacific, 6°05'N 

95°30'E, (600 m wire out)] 

Deosergestes Judkins & Kensley, 2008 

= Deosergestes Judkins & Kensley, 2008 (type species Sergestes curvatus Crosnier & Forest, 1973, by 


Deosergestes coalitus (Burkenroad, 1940) 

= Sergestes (Sergestes) coalitus Burkenroad, 1940: 39. [Dana stn 3737 I (Mindanao Sea, 7°23'N 121°29'E, 

1000 m wire out)] 

= Sergestes (Sergestes) erectus Burkenroad, 1940: 38. [Dana stn 3579 I (Rarotonga, 20°56'S 160°03'W), 


Deosergestes corniculum (Krøyer, 1855) 

= Sergestes corniculum Krøyer, 1855: 30. [fra Atlanterhavets tropiske Deel, 4.5°N 21.5°W; according to 

Krøyer, 1859] 

= Sergestes laciniatus Krøyer, 1859: 274, 282; Plate 5, fig. 15. [fra Kattegattet] 

? = Pasiphae rubroguttata Filhol, 1885b: Plate 5. [Type locality not indicated] 

= Sergestes longirostris Spence Bate, 1888: 415; Plate 75, fig. 3. [Mid Atlantic] 

= Sergestes (Sergestes) curvatus Crosnier & Forest, 1973: 315; Figs 105i-k, 107c-d, f-g. [au large de 

l’Afrique du Sud, 35°42'S 24°40'E, 500 m] 

Deosergestes disjunctus (Burkenroad, 1940) 

= Sergestes (Sergestes) disjunctus Burkenroad, 1940: 38. [Dana stn 3630 IV (New Zealand, Auckland, 

34°24'S 178°42.5'E), 1000 m wire out)] 

Deosergestes henseni (Ortmann, 1893) 

= Sergia henseni Ortmann, 1893: 38; Plate 3, fig. 3. (partim, see Crosnier & Forest, 1973). [Plankton- 

Expedition der Humboldt-Stiftung, Nördl. Aequatorialstrom JN 146 (12.3°N 22.3°W); Vertikalnetz 

0-400 m; Guineastrom JN 158 (7.5°N 21.3°W), circa 4000 m] 

Deosergestes paraseminudus (Crosnier & Forest, 1973) 

= Sergestes (Sergestes) paraseminudus Crosnier & Forest, 1973: 313; Figs 105d, 106c-d, f. [Ombango stn 

309-GS 20 bis, 1°55'S 8°30'E, 0-350 m] 

Deosergestes pediformis (Crosnier & Forest, 1973) 

= Sergestes (Sergestes) pediformis Crosnier & Forest, 1973: 313; Figs 105e-h, 107a-b, e. [Ombango stn 

302-GS 19, 4°47'S 10°42'E, 0-725 m]


Deosergestes rubroguttatus (Wood-Mason in Wood-Mason & Alcock, 1891c) 

= Sergestes rubroguttatus Wood-Mason in Wood-Mason & Alcock, 1891c: 354; Fig. 10. [Investigator 

stns 107 (Laccadive Sea, 8°23'N 75°47'E), 738 fms; 109 (S of C. Comorin, 7°41'N 78°21'E), 738 fms; 110 

(Bay of Bengal, 9°34'N 85°43'15''E), 1997 fms; 117 (Bay of Bengal, 11°58'N 88°52'17''E), 1748 fms] 

Deosergestes seminudus (Hansen, 1919) 

= Sergestes seminudus Hansen, 1919: 18; Plate 1, figs 7a-c; Plate 2, figs 1a-f. [Siboga stns 66, Bank between 

islands of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 144, Anchorage N 

of Salomakiëe (Damar-) Island, 45 m; 148, 0°17.6' S 129°14.5'E, 0-1000 m; 157, 0°32.9'S 130°14.6'E, 45 

m; 167, Anchorage N of Sabuda Island, 63 m; 177a, 2°30'S 129°28'E; 185, 3°20'S 127°22.9'E, Manipa 

Strait, 0-1536 m; 189a, 2°22'S 126°46'E; 194-197, 1°55' – 1°45.3'S 126°39'- 127°8.3'E; 203, 3°32.5'S 

124°15.5'E 0-1500 m; 223, 5°44.7'S 126°27.3'E, surface; 230, 3°58'S 128°20'E 0-2000 m; 276, 6°47.5'S 

128°40.5'E, 0-750 m] 

= Sergestes nipponensis Yokoya, 1933: 13; text-fig. 4. [Sôyô-Maru stns 21, Japan, SE of Siwoya-zaki, 209 

m; 25, Japan, E of Siwoya-zaki, 525 m; 253, Japan, Sagami Bay, 452 m] 

Eusergestes Judkins & Kensley, 2008 

= Eusergestes Judkins & Kensley, 2008 (type species Sergestes arcticus Krøyer, 1855, by original designation, 


Eusergestes antarcticus (Vereshchaka, 2009) comb. nov. 

= Sergestes antarcticus Vereshchaka, 2009: 57; Figs 25, 26. [Dana stn 3975-2 (South Atlantic, Cape 

Town, 35°42'S 18°37'E, 2500 m wire out)] 

Eusergestes arcticus (Krøyer, 1855) 

= Nika Sinuolata Risso, 1816: 87. [Nice; status of name discussed by Holthuis, 1977a] 

= Sergestes arcticus Krøyer, 1855: 27. [fra Grönland, according to Krøyer, 1859] 

= Sergestes Rinkii Krøyer, 1855: 33. [Atlanterhav på 58-59° n. Br., according to Krøyer, 1859] 

= Sergestes Meyeri Metzger, 1875: 302; Plate 6, fig. 7. [Kiel] 

= Sergestes magnificus Chun, 1888: 33; Plate 4, figs 4-5. [Ischia, 800 m] 

Eusergestes similis (Hansen, 1903) 

= Sergestes similis Hansen, 1903: 60; Plate 11, fig. 6. [Challenger stn 232, off Japan, 35°11'N 139°28'E, 

345 fms] 

? = Sergestes nasidentatus Spence Bate, 1888: 398; Plate 72, fig. 2. [Pacific Ocean between Valparaiso 

and Juan Fernandez, 0-200 fms; Holthuis (pers. comm.)] 

? = Sergestes laeviventralis Spence Bate, 1888: 425; Plate 67, fig. 3. [north of New Guinea] 

? = Sergestes longicaudatus Stimpson, 1860a: 46. [Oceano Pacifico, lat. Bor. 40°, long occ. 155°] 

? = Sergestes affinis Hansen, 1919: 7. [nomen nudum] 

Neosergestes Judkins & Kensley, 2008 

= Neosergestes Judkins & Kensley, 2008 (type species Sergestes edwardsii Krøyer, 1855, by original 


Neosergestes brevispinatus (Judkins, 1978) 

= Sergestes brevispinatus Judkins, 1978: 13; Figs 2d, 5d-f,h,k, 6-8, 216. [Eastern tropical Pacific, 11°02'S 

81°41'W, ca. 200 m at night] 

Neosergestes consobrinus (Milne, 1968) 

= Sergestes consobrinus Milne, 1968: 26; Figs 5-9. [Brown Bear cruise 199, haul 227, 33°44'N 124°53'W, 

120 m] 

Neosergestes edwardsii (Krøyer, 1855) 

= Sergestes Edwardsii Krøyer, 1855: 28. [nærheden af Linien, 3°S to 10°N, according to Krøyer, 1859] 

= Sergestes oculatus Krøyer, 1855: 28. [tropiske Atlanterhav, 4.5°N 21.5°W, according to Krøyer, 1859] 

= Sergestes brachyorrhos Krøyer, 1859: 272, 281; Plate 3, Figs 5a-f. [i Atlanterhavet, efter Angivelse paa 

omtrent 30°N 33°W] 

= Sergestes intermedius Spence Bate, 1888: 383. [Off Luzon, China Sea] 

= Sergestes ventridentatus Spence Bate, 1888: 431. [N of the Sandwich Islands]


Neosergestes orientalis (Hansen, 1919) 

= Sergestes orientalis Hansen, 1919: 22; Plate 2, fig. 2a-q. [Siboga stns 37, Sailus Ketjil, Paternoster Islands, 

0-27 m; 40, Anchorage off Pulu Kawassang, Paternoster Islands, 12 m; 66, Bank between islands 

of Bahuluwang and Tambolungan, S of Salayer, Indonesia, 8-10 m; 96, SE side of Pearl bank, 

Sulu Archipelago, 0-15 m; 106, anchorage off Kapul Island, Sulu Archipelago, 13 m; 118, 1°38'N 

124°28.2'E, 0-900 m; 125, anchorage off Sawan, Siau Island, 27 m; 128, 4°27'N 125°25.7'E, 0-700 m; 

131-133, about 5°56.7'N 126°25'E; 141, 1°0.4'S 127°25.3'E, 0-1500 m; 143, 1°4.5'S 127°52.6'E, 0-1000 m; 

144, Anchorage N of Salomakiëe (Damar-) Island, 45 m; 146, 0°36'S 128°32.7'E, surface; 148, 0°17.6' 

S 129°14.5'E, 0-1000 m; 157, 0°32.9'S 130°14.6'E, 45 m; 177a, 2°30'S 129°28'E; 168, anchorage N of 

Sabuda Island, surface; 172, Gisser, anchorage between this island and Ceram Island, 18 m; 177a, 

2°30'S 129°28'E, surface; 185, 3°20'S 127°22.9'E, Manipa Strait, 0-1536 m; 189a, 2°22'S 126°46'E; 

194-197, 1°55'-1°45.3'S 126°39'-127°8.3'E; 194-197, 1°53.5'-1°45.3'S 126°39'-127°8.3'E; 203, 3°32.5'S 

124°15.5'E, 0-1500 m; 206 or 207, Buton Strait, surface; 220, anchorage off Pasir Pandjang, W coast of 

Binongka, surface; 225, N. 279°E from Southpoint of South Lucipara Island, 894 m; 230, 3°58'S 

128°20'E, 0-2000 m; 245, 4°16.5'S 130°15.8'E, surface] 

= Sergestes geminus Judkins, 1978: 25; Figs 2a-c, 7, 16f-j, 17, 18, 21a. [Eastern Tropical Pacific, 4°48'N 

83°38'W, ca. 200 m at night] 

= Sergestes gibbilobatus Judkins, 1978: 27; Figs 2g, 4c, 7, 19a-h, 20, 21a. [Equatorial Pacific, 0°0'S 

165°42'W, 2550 m wire out, at night] 

Neosergestes semissis (Burkenroad, 1940) 

= Sergestes (Sergestes) semissis Burkenroad, 1940: 42. [Dana stn 3905 I (Bay of Bengal, 4°44'N 88°05.5'E), 

1000 m wire out] 

Neosergestes tantillus (Burkenroad, 1940) 

= Sergestes (Sergestes) tantillus Burkenroad, 1940: 42. [Dana stn 3556 II (Gulf of Panama, 2°52'N 

87°36'W), 2500 m wire out)] 

Parasergestes Judkins & Kensley, 2008 

= Parasergestes Judkins & Kensley, 2008 (type species Sergestes armatus Krøyer, 1855, by original 


Parasergestes armatus (Krøyer, 1855) 

= Sergestes armatus Krøyer, 1855: 31. [7°37'n.Br. og N 22.5° v. Lgd., according to Krøyer, 1859] 

= Sergestes incertus Hansen, 1896: 962. [34°50'S 4°30'W and 40°4'S 53°20'W, surface] 

= Sergestes extensus Hanamura, 1983: 64; Figs 7-8. [Kaiyo Maru stn MT 9-B, Seamount 350, off Baja 

California, 23-05.4N 124-56.9W, 0-1236 m] 

Parasergestes cylindricus (Vereshchaka, 2009) comb. nov. 

= Sergestes cylindricus Vereshchaka, 2009: 96; Figs 47, 48. [Dana stn 3639-1, Southeast Pacific, 39°19'S 

179°18'E (300 m wire out)] 

Parasergestes diapontius (Spence Bate, 1881) 

= Sergestes diapontius Spence Bate, 1881: 194. [Atlantic] 

= Sergestes penerinki Spence Bate, 1888: 418; Plate 76, fig. 3. [North Atlantic Ocean] 

= Sergestes fermerinkii Spence Bate, 1888: 419; Plate 76, fig. 4. [Pacific Ocean, 24°S 148°W] 

= Sergestes semiarmis Spence Bate, 1888: 423; Plate 67, fig. 1. [Challenger stn 354, Mid North Atlantic 

Ocean, 32°41'N 36°6'W, surface] 

Parasergestes halia (Faxon, 1893) 

= Sergestes halia Faxon, 1893: 217. [Albatross stn 3388 (off Panama, 7°06'N 79°48'W, surface to 400 fms)] 

Parasergestes sirenkoi (Vereshchaka, 2009) comb. nov. 

= Sergestes sirenkoi Vereshchaka, 2009: 103; Figs 51, 52. [Dana stn 3593-8, Southwest Pacific, 17°27'S 

179°33'E (300 m wire out)] 

Parasergestes stimulator (Burkenroad, 1940) 

= Sergestes (Sergestes) stimulator Burkenroad, 1940: 41. [Dana stn 3656 VIII (off Sydney, Australia, 

33°26'S 157°02'E), 300 m wire out)] 

Parasergestes vigilax (Stimpson, 1860a)


= Sergestes vigilax Stimpson, 1860a: 45. [Oceano Atlantico prope insulas ”Azores”] 

= Sergestes macrophthalmus Stimpson, 1860a: 46. [Oceano Pacifico, lat. bor. 27,5°, long. orient. 138,5°; 

etiam lat. bor. 35°, long. occ. 155°] 

= Sergestes parvidens Spence Bate, 1888: 409; Plate 74, fig. 3. [the tropical part of the Atlantic; Pacific 

Ocean, north of the Sandwich Islands; off Sydney and Wellington, Australia] 

? = Sergestes spiniventralis Spence Bate, 1888: 426; Plate 67, fig. 5. [North Pacific Ocean] 

? = Sergia Clausi König, 1895: 10; Plate 1, figs 1-7. [Östliches Mittelmeer, 36°12'N 28°54'E, Oberflächenfang] 

Peisos Burkenroad, 1945 

= Peisos Burkenroad, 1945 (type species Peisos petrunkevitchi Burkenroad, 1945, by original designation, 


Peisos petrunkevitchi Burkenroad, 1945: 554; Plates 1-2. [Montevideo, 5-6 fms] 

Petalidium Spence Bate, 1881 

= Petalidium Spence Bate, 1881 (type species Petalidium foliaceum Spence Bate, 1881, by monotypy, 

gender neuter) 

Petalidium foliaceum Spence Bate, 1881: 194. [taken in South Indian Ocean at a depth of about 2100 fms; 

according to Spence Bate, 1888: Challenger stn 159, South of Australia, 47°25'S 130°22'E, 2150 fms] 

Petalidium obesum (Krøyer, 1855) 

= Sergestes obesus Krøyer, 1855: 31. [4.5°N Br. i Atlanterhavet, according to Krøyer, 1959] 

= Sergestes sanguineus Chun, 1889: 538; Plate 3, fig. 1. [Canarischen Inseln] 

Petalidium suspiriosum Burkenroad, 1937: 325; Text-Figs 1-6. [Templeton Crocker Expedition stn 165, 145 

miles N of Clarion Island, Revillagigido Islands, Lower California, E Pacific, 500 fms] 

Sergestes H. Milne edwards, 1830 

= Sergestes H. Milne Edwards, 1830 (type species Sergestes atlanticus H. Milne Edwards, 1830, by 


= Acheles Cocco, 1832 (type species Acheles arachnipodus Cocco, 1832, by monotypy, gender feminine) 

Sergestes atlanticus H. Milne Edwards, 1830: 349; Plate 10, figs 1-9. [l’Océan atlantique, à une grande 

distance des côtes] 

= Sergestes Frisii Krøyer, 1855: 26. [Atlanterhav, 13°N 27.5°W and 20°N 36°W, according to Krøyer, 

1859] 

= Sergestes ancylops Krøyer, 1855: 32. [Atlanterhav 13°N 27.5'W, according to Krøyer, 1859] 

? = Sergestes pacificus Stimpson, 1860a: 45. [Oceano Pacifico, lat. bor. 27,5°, long. orient. 138°] 

= Sergestes ovatoculus Spence Bate, 1888: 408; Plate 74, fig. 2. [N Atlantic Ocean] 

Sergestes cornutus Krøyer, 1855: 29. [Atlanterhavet omtrent 4.5°N for Linien; indtil 10° nord for Linien og 

undtil 8° syd for Linien, according to Krøyer, 1859] 

= Sergestes longispinus Spence Bate, 1888: 417; Plate 76, fig. 2. [Challenger stn 106, Mid Atlantic Ocean, 

1°47'N 24°26'W, within 40 fms of the surface] 

Sergestes formosensis Yokoya & Shibata, 1965: 2; Fig. 2. [Philippine Sea, 21°42'N 123°05'E, 100 m] 

Sergestes grandipes Yokoya & Shibata, 1965: 1; Fig. 1. [Philippine Sea, 13°57'N 125°29'E, 100 m] 

Sergestes hamifer Alcock & Anderson, 1894: 148. [Investigator stn 126, Laccadive Sea, 8°49'0"N 73°18'45"E, 

1370 fms] 

Sergestes latirostris Yokoya & Shibata, 1965: 3; Fig. 3. [Philippine Sea, 13°03'N 125°50'E, 40 m] 

Sergia Stimpson, 1860a 

= Sergia Stimpson, 1860a (type species Sergia remipes Stimpson, 1860a, by monotypy, gender feminine; 

the type species was determined to be a late mastigopus stage by Burkenroad, 1945, who 

identified it with the group of species in what was then the subgenus Sergia) 

Sergia bigemmea (Burkenroad, 1940) 

= Sergestes (Sergia) bigemmeus Burkenroad, 1940: 49. [Dana stn 3570 VI (NE of Tahiti, 3°09'N 

126°09.3'E), 2150 m]


Sergia bisulcata (Wood-Mason & Alcock, 1891b) 

= Sergestes bisulcatus Wood-Mason & Alcock, 1891b: 190. [Investigator stns 100 (Bay of Bengal, 

16°55'41"N, 83°21'18"E), 840 fms; 105 (Arabian Sea, 15°02'N, 72°34'E), 740 fms] 

Sergia burukovskii Vereshchaka, 2000: 121; Figs 31-33. [Dana stn 3980-1, South Atlantic, 23°26'S, 03°56'E, 

1000 m wire out] 

Sergia challengeri (Hansen, 1903) 

= Sergestes challengeri Hansen, 1903: 61; Plate 12, fig. 2a-n. [Challenger stn 173, Western Pacific, off 

Matuku, Fiji Islands, 19°9'35"S, 179°41'50"E, 315 fms] 

Sergia crosnieri Vereshchaka, 2000: 196; Figs 85-87. [Dana stn 3809-4, off Indonesian islands, 6°22'S, 

105°12'E, 50 m wire out] 

Sergia erythraeensis Iwasaki & Couwelaar, 2001: 92; Figs 1-3. [Central Red Sea, 21°23.89'N 38°03.62'E, 150 

m] 

Sergia extenuata (Burkenroad, 1940) 

= Sergestes (Sergia) extenuates Burkenroad, 1940: 45. [Dana stn 3999 II (Saint Helena, S Atlantic, 3°45'S 

10°00'E, 1000 m wire out); see discussion in Vereshchaka, 2000] 

Sergia filicta (Burkenroad, 1940) 

= Sergestes (Sergia) filictum Burkenroad, 1940: 52. [Dana stn 3549 IV (Gulf of Panama, 7°16'N 79°30'W), 

wire out 600 m] 

Sergia foresti Kensley & Judkins, 2008: 151; Fig. 1. [Albatross stn 5241, Pujada Bay, Mindanao, 6°50'45"N 

126°14'38"E, 393 m] 

Sergia fulgens (Hansen, 1919) 

= Sergestes fulgens Hansen, 1919: 17; Plate 1, fig. 6a-g. [Siboga stn 312, Indonesia, Saleh Bay, Sumbawa, 

08°19'S 117°41'E, 174 m] 

Sergia gardineri (Kemp, 1913a) 

= Sergestes gardineri Kemp, 1913a: 55; Plate 7, figs 2-5. [S by E of Farquhar, 10°27'S, 51°17'E; NE of 

Madagascar, between Providence and Alphonse Islands, 8°16'S, 51°26'E; 5 miles off Desroches Atoll] 

Sergia grandis (Sund, 1920) 

= Sergestes grandis Sund, 1920: 16; Figs 22-26. [Michael Sars stns 34, (28°52'N 14°16'W, 2170 m); 49 

(29°6'N, 25°2'W); 51 (31°20'N 35°7'W, 3886 m); 52 (31°24'N, 34°47'W] 

Sergia hansjacobi Vereshchaka, 1994: 91; Figs 23-24. [Dana stn 1198-2, 17°43'N, 64°56'W] 

= Sergia hanseni Vereshchaka, 1994: 91. [nomen nudum] 

Sergia inequalis (Burkenroad, 1940) 

= Sergestes (Sergia) inequalis Burkenroad, 1940: 51. [Dana stn 3768 (N of NW New Guinea, 1°20'S 

138°42'E), 2500 m wire out)] 

Sergia inoa (Faxon, 1893) 

= Sergestes inous Faxon, 1893: 216. [Albatross stn 3380 (East Pacific, off Malpelo Island, Colombia, 

04°03'N, 81°31'W, 899 fms)] 

Sergia japonica (Spence Bate, 1881) 

= Sergestes japonicus Spence Bate, 1881: 194. [south of Japan; according to Spence Bate, 1888: Challenger 

stn 232, western Pacific off southern coast of Japan, 35°11'N, 139°28'E, 345 fms] 

= Sergestes mollis Smith, 1884: 419. [Albatross stns 2002, 37°20'42"N 74°17'36"W, 641 fms; 2018, 

37°12'22"N 74°17'36"W, 788 fms; 2040, 38°35'13"N 58°16'00"W, 2226 fms; 2045, 40°04'20"N 68°43'50"W, 

373 fms; 2051, 39°41'00"N 69°20'20"W, 1106 fms; 2083, 40°26'40"N 67°05'15"W, 959 fms; 2093, 

39°42'50"N 71°01'20"W, 1000 fms; 2094, 39°44'30"N 71°04'00"W, 1022 fms; 2097, 37°56'20"N 

70°57'30"W, 1917 fms; 2099, 27°12'20"N 69°39'00"W, 2949 fms; 2100, 39°22'00"N 68°34'30"W, 1628 

fms; 2101, 39°18'30"N 68°24'00"W, 1686 fms; 2103, 38°47'20"N 72°37'00"W, 1091 fms; 2104, 38°48'00"N 

72°40'30"W, 991 fms; 2105, 37°50'00"N 73°03'50"W, 1395 fms; 2110, 35°12'10"N 74°57'15"W, 516 fms; 

2116, 35°45'23"N 74°31'25"W, 888 fms] 

= Sergestes profundus Spence Bate, 1888: 428 (partim). [Challenger stn 300, southeastern Pacific west 

of Valparaiso, Chile, 33°42'S, 78°18'W, 1375 fms; lectotype designation by Hansen, 1903] 

Sergia jeppeseni Vereshchaka, 2000: 179; Figs 67, 74-75; Plate 5F. [Dana stn 3943-1, Western Indian Ocean 

off Mombasa, 5°30'S, 40°40'E, 500 m wire out]


Sergia kensleyi Vereshchaka, 2000: 110; Figs 17, 24-25; Plate 4D. [Dana stn 3970-1, Western Indian Ocean 

off Mozambique, 34°09'S, 27°38'E] 

Sergia laminata (Burkenroad, 1940) 

= Sergestes (Sergia) laminatus Burkenroad, 1940: 53. [Dana stn 3933 I (N of Madagascar, 11°18'S 

50°03'E), 4000 m wire out] 

= Sergestes (Sergia) guineensis Crosnier & Forest, 1973: 343; Fig. 118. [Ombango stn 394-8, 5°52'S 

10°00'E, 0-2500 m] 

Sergia lucens (Hansen, 1922) 

= Sergestes lucens Hansen, 1922: 12, 38. [Suruga Bay; see discussion in Vereshchaka, 2000] 

Sergia manningorum Froglia & Gramitto, 2000: 72; Figs 1-3. [Atlantis II, cruise 60 stn RH2281, off Congo, 

11°23'S 10°55'E, 190-200 m] 

Sergia maxima (Burkenroad, 1940) 

= Sergestes (Sergia) maximus Burkenroad, 1940: 47. [Dana stn 3933 I (N of Madagascar, 11°18'S 

50°03'E, 4000 m wire out)] 

Sergia oksanae Vereshchaka, 2000: 182: Figs 73, 76-77; Plate 5E. [Dana stn 3736-3, Mindanao Sea, 9°17'N, 

123°58'E] 

Sergia phorca (Faxon, 1893) 

= Sergestes phorcus Faxon, 1893: 217. [Albatross stns 3382 (off Panama, 6°21'N 80°41'W, 1793 fms); 

3388 (off Panama, 7°06'N 79°48'W, 1168 fms); 3401 (off Galapagos Islands, 0°50'S 88°58'30"W, 395 

fms); 3386 (off Panama, 7°33'12"N 79°17'15"W, 242 fms); 3437 (Gulf of California, 27°39'40"N 

111°00'30"W, 628 fms)] 

Sergia plumea (Illig, 1927) 

= Sergestes plumeus Illig, 1927: 295; Figs 30-32. [Valdivia stn 268, Indischer Ozean, südlich von Ras 

Hafun, 9°6'N, 53°41'E, im Vertikalnetzfang von 1500 m Tiefe] 

Sergia potens (Burkenroad, 1940) 

= Sergestes (Sergia) potens Burkenroad, 1940: 48. [Dana stn 3975 VII (Agulhas Bank, South Africa, 

35°42'S 18°37'E, 600 m wire out)] 

Sergia prehensilis (Spence Bate, 1881) 

= Sergestes prehensilis Spence Bate, 1881: 193. [off Japan, 500 fms; according to Spence Bate, 1888: 

Challenger stn 236, off Japan, 34°58'N, 139°29'E, 775 fms] 

= Sergestes gloriosus Stebbing, 1905: 84; Plates 22-23. [off Sandy Point, 800 fms] 

= Sergestes fujiyamaensis Nakazawa, 1932: 32. [Type locality not indicated] 

Sergia profunda (Spence Bate, 1888) 

= Sergestes profundus Spence Bate, 1888: 428. [Challenger stn 137, 35°59'S 1°34'E, 2550 fms; 300, W of 

Valparaiso, 33°42'S 78°18'W, 1375 fms] 

Sergia regalis (Gordon, 1939) 

= Sergestes regalis Gordon, 1939: 498; Figs 1-4. [Discovery stn 81, South Atlantic, 32°45'S, 8°47'W] 

= Sergestes (Sergia) creber Burkenroad, 1940: 44. [Dana stn 3766 XVIII (N of NW new Guinea, 1°13'S 

138°42'E), 2900 m wire out] 

Sergia remipes Stimpson, 1860a: 46. [Oceano Pacifico, lat bor. 27½° log. orient. 138½°] 

Sergia robusta (Smith, 1882) 

= Sergestes robustus Smith, 1882: 97; Plate 16, fig. 5-8b. [Blake stn 328, 34°28'25''N 75°22'50''W, 1632 

fms; Albatross stns 893 (37°17'N 73°21'W), 372 fms; 952 (34°28'50"N 75°22'50"W), 388 fms] 

= Sergestes dissimilis Spence Bate, 1888: 437. [St. Vincent, Cape Verde Island, at the surface] 

= Sergestes mediterraneus Hansen, 1896: 954. [Ragusa and Lesina, Adriatic Sea] 

Sergia scintillans (Burkenroad, 1940) 

= Sergestes (Sergia) scintillans Burkenroad, 1940: 43. [Dana stn 3622 I (SE of New Caledonia, 25°54'S 

172°36.9'E, 300 m wire out)] 

Sergia splendens (Sund, 1920) 

= Sergestes splendens Sund, 1920: 14; Figs 16-18. [Michael Sars stns 29 (35°10'N 7°55'W) 440 m wire 

out; 42 (28°2'N 14°17'W), 300 m wire out; 45 (28°42'N 20°0'W), 100, 200, 300, 2000, & 3000 m wire 

out; 49 (29°6'N 25°2'W), 3000 m wire out; 51 (31°20'N 35°7'W), 200, 300 & 1000 m wire out; 52 

(31°24'N 34°47'W), 100 m wire out; 53 (34°59'N 33°1'W), 100, 300, 600 & 2600 m wire out; 56


(36°53'N 29°17'W), 200 & 300 m wire out; 62 (36°52'N 39°55'W), 2000 m wire out; 64 (34°44'N 

47°52'W), 100 & 2000 m wire out; 66 (39°30'N 49°42'W), 1000 m wire out; 67 (40°17'N 50°39'W), 

1200 & 2000 m wire out)] 

= Sergestes Richardi Hansen, 1920: 482. [nomen novum for Sergestes splendens Sund, 1920, considered 

a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1920] 

= Sergestes crassus Hansen, 1922: 98; Plate 5, fig. 4. [nomen novum for Sergestes splendens Sund, 1920, 

considered a junior homonym of Sergestes splendens Hansen, 1919 by Hansen, 1922] 

Sergia stellata (Burkenroad, 1940) 

= Sergestes (Sergia) stellatus Burkenroad, 1940: 44. [Dana stn 3908 I (SE of Sri Lanka, 4°28'N 82°13'E, 


Sergia talismani (Barnard, 1947) 

= Sergestes splendens Hansen, 1919: 18. [nomen nudum] 

= Sergestes splendens Hansen, 1920: 480; nec Sund, 1920. [Talisman Expedition, stn 113, canal de Saint- 

Vincent, 16°52'N 27°30 - 27°31'W, 550-760 m] 

= Sergestes talismani Barnard, 1947: 384. [nomen novum for Sergestes splendens Hansen, 1919 nec 

Sund, 1920] 

Sergia tenuiremis (Krøyer, 1855) 

= Sergestes tenuiremis Krøyer, 1855: 30. [2°N, 21°W, according to Krøyer, 1859] 

= Sergestes Kröyeri Spence Bate, 1881: 193. [off Kermadec Island, about 500 fms; according to Spence 

Bate, 1888: Challenger stn 170, off the Kermadec Islands, 29°55'S 178°14'W, 520 fms] 

= Sergestes junceus Spence Bate, 1888: 416; Plate 76, fig. 1. [South Pacifc Ocean] 

= Sergestes longicollis Spence Bate, 1888: 421; Fig. 1. [Challenger near stn 131, South Atlantic Ocean, 

29°35'S 28°9'W; stn 295, South Pacific Ocean, 38°7'S 94°4'W, 1500 fms] 

= Sergestes tropicus Sund, 1920: 18; Figs 27, 28, 30-33. [Michael Sars stns 29 (35°10'N 7°55'W); 45 

(28°42'N 20°0'W); 49 (29°6'N 25°2'W); 51 (31°20'N 35°7'W, 3886 m); 52 (31°24'N 34°47'W); 53 (34°59'N 

33°1'W, 2615-2865 m); 56 (36°53'N 29°17'W, 3239 m); 64 (34°44'N 47°52'W)] 

Sergia umitakae Hashizume & Omori, 1995: 72; Figs 1-4. [South of Sri Lanka] 

Fig. 7. Sicyonella aff. maldivensis Borradaile, 1910. Photo by Tin-Yam Chan.


Sergia vityazi Vereshchaka, 2000: 157; Figs 51, 57-58. [Dana stn 3601-1, southwestern Pacific, 18°21'S, 

178°21'E] 

Sergia wolffi Vereshchaka, 1994: 88; Figs 19-21. [Dana stn 1217-1, 18°50'N, 79°07'W] 

Sicyonella Borradaile, 1910 

= Aphareus Paul’son, 1875 (type species Aphareus inermis Paul’son, 1875, by monotypy, gender masculine; 

invalid junior homonym of Aphareus Cuvier & Valenciennes, 1830 (Pisces)) 

= Sicyonella Borradaile, 1910 (type species Sicyonella maldivensis Borradaile, 1910, by monotypy, gender 

feminine) 

= Aphareocaris Calman, 1913 (nomen novum for Aphareus Paul’son, 1875, gender feminine) 

Sicyonella antennata Hansen, 1919: 30; Plate 2, fig. 5a-c; Plate 3, fig. 1a-f. [Siboga stn 258, Tual anchorage, 

Kei Islands, 22 m] 

Sicyonella inermis (Paul’son, 1875) 

= Aphareus inermis Paul’son, 1875: 117; Plate 18, fig. 3. [Red Sea] 

= Aphareocaris elegans Calman, 1913: 219; Plate 16, figs 1-16. [Thursday Island, Torres Straits] 

Sicyonella maldivensis Borradaile, 1910: 259; Plate 16, figs 3-3a. [North Male Atoll, Maldives; lectotype 

designation by Fukuoka, Tamaki & Kikuchi, 2005] (Fig. 7) 

Suborder PleoCyeMATA Burkenroad, 1963 

Infraorder PRoCARIDIDeA Felgenhauer & Abele, 1983 

Family PRoCARIDIDAe Chace & Manning, 1972 

Procaris Chace & Manning, 1972 

= Procaris Chace & Manning, 1972 (type species Procaris ascensionis Chace & Manning, 1972, by 


Procaris ascensionis Chace & Manning, 1972: 6; Figs 4-9. [coral pool back of Shelly Beach, Ascension Island] 

Procaris chacei C.W.J. Hart & Manning, 1986: 408; Figs 1-26. [Bermuda, Hamilton Parish, Green Bay Cave 

(North Shore passage), 32°19'N 64°44'W] 

Procaris hawaiana Holthuis, 1973a: 12; Figs 4-6. [near Nukuele Point, Cape Kinau, Maui, Hawaiian Islands, 

in lava pool] (Fig. 8) 

Procaris mexicana von Sternberg & Schotte, 2004: 514-522; Figs 1-3. [México, Cueva Quebrada, Chankanaab 

Park, Cozumel, Quintana Roo] 

Procaris noelensis Bruce & Davie, 2006: 23-33; Figs 1-3. [Runaway Cave, c. 1.5 km south from the North 

East Point of Christmas Island] 

Fig. 8. Procaris hawaiana Holthuis, 1973. Photo by Troy Sakihara.


Vetericaris Kensley & Williams, 1986 

= Vetericaris Kensley & Williams, 1986 (type species Vetericaris chaceorum Kensley & Williams, 1986, 

by monotypy, gender feminine) 

Vetericaris chaceorum Kensley & Williams, 1986: 419, Figs 2-7. [Lua o Palahemo, 150 m north of shoreline 

at Ka Lae (South Point), Hawaii Island, 18°55'N 155°42'W, 33 m] 

Infraorder STeNoPoDIDeA Spence Bate, 1888 

Family MACROMAXILLOCARIDIDAE Alvarez, Iliffe & Villalobos, 2006 

Macromaxillocaris Alvarez, Iliffe & Villalobos, 2006 

= Macromaxillocaris Alvarez, Iliffe & Villalobos, 2006 (type species Macromaxillocaris bahamaensis 

Alvarez, Iliffe & Villalobos, 2006, by original designation and monotypy, gender feminine) 

Macromaxillocaris bahamaensis Alvarez, Iliffe & Villalobos, 2006: 370; Figs 2-7. [Oven Rock Cave, Great 

Guana Cay, Exuma Cays, Bahamas] (Fig. 9) 

Fig. 9. Macromaxillocaris bahamaensis Alvarez, Iliffe & Villalobos, 2006. Photo by T.M. Iliffe. 

Family SPoNGIColIDAe Schram, 1986 

Engystenopus Alcock & Anderson, 1894 

= Engystenopus Alcock & Anderson, 1894 (type species Engystenopus palmipes Alcock & Anderson, 

1894, by monotypy, gender masculine) 

Engystenopus palmipes Alcock & Anderson, 1894: 149; Plate 9, fig. 1. [Investigator stn 172, Bay of Bengal, 

off Trincomallee, 200-350 fms] 

Globospongicola Komai & Saito, 2006 

= Globospongicola Komai & Saito, 2006 (type species Globospongicola nudibranchus Komai & Saito, 


Globospongicola nudibranchus Komai & Saito, 2006: 268; Figs 1-6. [Kai Islands, Banda Sea, Indonesia, 

6°05'S 132°44'E, 268-210 m] 

Globospongicola spinulatus Komai & Saito, 2006: 276; Figs 7-12. [Vanuatu, 17°50.35'S 168°39.33'E, 437-504 m]


Microprosthema Stimpson, 1860a 

= Microprosthema Stimpson, 1860a (type species Microprosthema valida Stimpson, 1860a, by monotypy, 


= Stenopusculus Richters, 1880 (type species Stenopusculus crassimanus Richters, 1880, designated by 

Holthuis, 1955, a junior subjective synonym of Microprosthema valida Stimpson, 1860a, gender masculine) 

Microprosthema emmiltum Goy, 1987: 717; Figs 1-4. [1°16'36''S 90°29'42''W, off Black Beach, Isla Santa Maria, 

Galapagos Islands; rocky shores] 

Microprosthema fujitai Saito & Okuno, 2011: 84; Figs 1-7. [Maeda-misaki, Okinawa Island, Ryukyu Islands, 

3 m] 

Microprosthema granatense Criales, 1997: 538; Figs 1-4. [Granate Bay, Caribbean coast of Colombia, 

11°18'N 74°9'W, 23 m] 

Microprosthema inornatum Manning & Chace, 1990: 26; Figs 12-13. [off North Point, Ascension Island, 18 

m] 

Microprosthema jareckii Martin, 2002: 109; Figs 1-5. [Monkey Point, Guana Island, British Virgin Islands, 

10 m] 

Microprosthema looensis Goy & Felder, 1988: 1286; Figs, 5-7. [near study finger number 3, Looe Key, Florida, 

USA, 4-5 m] 

Microprosthema lubricum Saito & Okuno, 2011: 93; Figs 8-11. [Agenashiku Islet, the Kerama Islands, the 

Ryukyu Islands, 3m] 

Microprosthema manningi Goy & Felder, 1988: 1277; Figs 1-4. [17°29'N 88°10'W, Stann Creek District, Carrie 

Bow Cay, Belize, near shore in dead conch shell] 

Microprosthema plumicorne (Richters, 1880) (Fig. 10) 

= Stenopusculus plumicornis Richters, 1880: 167; Plate 18, figs 16-26. [Fouquets, Mauritius] 

Microprosthema scabricaudatum (Richters, 1880) 

= Stenopusculus scabricaudatus Richters, 1880: 168; Plate 18, figs 30-32. [Fouquets, Mauritius] 

Microprosthema semilaeve (von Martens, 1872) 

= Stenopus semilaevis von Martens, 1872: 144. [Cuba] 

= Stenopusculus spinosus Pocock, 1890: 523. [Fernando de Noronha] 

Fig. 10. Microprosthema plumicorne (Richters, 1880). Photo by Arthur Anker.


Microprosthema validum Stimpson, 1860a 

= Microprosthema valida Stimpson, 1860a: 45. [in sinu insulae Ousima, sublittoralis, in locis lapillosis 

algosisque] 

= Stenopusculus crassimanus Richters, 1880: 168; Plate 18, figs 27-29. [Fouquets, Mauritius] 

= Stenopus robustus Borradaile, 1910: 260; Plate 16, fig. 4. [Salomon Atoll, Chagos Archipelago] 

Paraspongicola de Saint laurent & Cleva, 1981 

= Paraspongicola de Saint Laurent & Cleva, 1981 (type species Paraspongicola pusilla de Saint Laurent 

& Cleva, 1981, by original designation and monotypy, gender masculine) 

Paraspongicola acantholepis Komai, 2011: 305; Figs 12-15. [R/V Tansei-maru, KT07-31, stn L3-200, Isu Islands, 

N of Toshima Island, 34°34.04'N 139°18.37'E to 34°33.56'N 139°17.81'E, 187-261 m] 

Paraspongicola inflatus de Saint Laurent & Cleva, 1981 

= Spongicola inflata de Saint Laurent & Cleva, 1981: 179; Fig. 13. [14°02.2'N 120°17.7'E, 193-184 m] 

Paraspongicola pusillus de Saint Laurent & Cleva, 1981 

= Paraspongicola pusilla de Saint Laurent & Cleva, 1981: 181; Figs 14-15. [13°56.3'N 120°16.2'E, 150- 

159 m] 

Spongicola De Haan, 1844 [in De Haan, 1833-1850] 

= Spongicola De Haan, 1844 [in De Haan, 1833-1850] (type species Spongicola venusta De Haan, 1844 

[in De Haan, 1833-1850], by monotypy, gender masculine) 

Spongicola andamanicus Alcock, 1901 

= Spongicola andamanica Alcock, 1901: 148; Plate 2, fig. 2. [Andaman Sea, 170 and 238-290 fms] 

= Spongicola henshawi Rathbun, 1906: 901; Plate 24, fig. 8. [south coast of Molokai Island, Hawaii, 

169-182 fms] 

= Spongicola henshawi spinigera de Saint Laurent & Cleva, 1981: 174; Figs 10b, 11c, g-h. [8°47'N 

123°31.2'E, 333 m] 

= Spongicola holthuisi de Saint Laurent & Cleva, 1981: 177; Fig. 12. [13°59.8'N 120°18.6'E, 192-188 m] 

Spongicola cubanicus Ortiz, Gómez & Lalana R., 1994 

= Spongicola cubanica Ortiz, Gómez & Lalana R., 1994: 189; Figs 2-7. [Bahía Honda, costa norte de la 

Provincia de Pinar del Rio, 300 m, asociado a la esponja silícea Dactylocalyx pumiceus] 

Spongicola depressus Saito & Komai, 2008: 103; Figs 8-10. [Loyalty Islands, depth unknown] 

Spongicola goyi Saito & Komai, 2008: 107; Figs 11-15. [22°03.60'S 167°27.00'E, 850 m] 

Spongicola japonicus Kubo, 1942a 

= Spongicola japonica Kubo, 1942a: 90; Figs 1-2. [Kumanonada, off Mie Prefecture, about 300 m, commensal 

with Euplectella marshalii] 

Spongicola levigatus Hayashi & Ogawa, 1987 

= Spongicola levigata Hayashi & Ogawa, 1987: 367; Figs 1-4. [East China Sea, 30°44.7'N 127°48.3'E, 

about 200 m] 

Spongicola parvispinus Zarenkov, 1990 

= Spongicola parvispina Zarenkov, 1990: 218. [Sala-y-Gomez ridges, southeastern Pacific, 470-485 m] 

Spongicola robustus Saito & Komai, 2008: 118; Figs 20-24. [Mauritius, 10°29.7'S 61°12.4'E, 115-110 m] 

Spongicola venustus De Haan, 1844 [in De Haan, 1833-1850] 

= Spongicola venusta De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 9 (1844); 194, Plate P (1849). 

[Japan; lectotype designation by Yamaguchi & Baba, 1993] 

Spongicoloides Hansen, 1908 

= Spongicoloides Hansen, 1908 (type species Spongicoloides profundus Hansen, 1908, by monotypy, 


Spongicoloides evolutus (Bouvier, 1905b) 

= Spongicola evoluta Bouvier, 1905b: 983. [au large des côtes du Sahara, 882 m] 

Spongicoloides galapagensis Goy, 1980: 760; Figs 1-4. [Galapagos Islands, 00°29' 89°54'30''W, 717 m] 

Spongicoloides hawaiiensis Baba, 1983: 477; Figs 1-2. [French Frigate Shoals, north-western Hawaiian Islands, 

640 m, associated with glass sponge Euplectella sp.]


Spongicoloides iheyaensis Saito, Tsuchida & Yamamoto, 2006: 224; Figs 3-8. [the North Knoll of the Iheya 

Ridge, Ryukyu Islands, 27°47.17'N 26°53.91'E, 988 m] 

Spongicoloides inermis (Bouvier, 1905a) 

= Richardina inermis Bouvier, 1905a: 749. [dans les parages de Sainte-Lucie, 22-423 brasses] 

Spongicoloides koehleri (Caullery, 1896) 

= Spongicola Kœhleri Caullery, 1896: 382; Plate 16, figs 1-9. [Golfe de Gascogne, 45°57'N 6°21'E, 1410 

m] 

Spongicoloides novaezelandiae Baba, 1979: 311; Figs 1-2. [Chatham Rise, 44°44.0'S 175°42.0'E, 990-1110 m] 

Spongicoloides profundus Hansen, 1908: 45; Plate 3, figs 5a-k; Plate 4, figs 1a-l. [78 miles off the southwest 

coast of Iceland, 60°37'N 27°52'W, 1480 m; lectotype designation by Goy, 1982] 

Spongicoloides tabachnicki Burukovsky, 2009: 498; Figs 1-3. [Atlantic Ocean, Florida Straits, 23°54'N 

81°27'W, 1003-1336 m] 

Spongiocaris Bruce & Baba, 1973 

= Spongiocaris Bruce & Baba, 1973 (type species Spongiocaris semiteres Bruce & Baba, 1973, by original 


Spongiocaris hexactinellicola Berggren, 1993: 784; Figs 1-5. [24°30'N 74°28'W, Tartar Bank, Cat Island, Bahamas, 

60-610 m; in the internal cavity of Euplectella jovis] 

Spongiocaris goyi Ortiz, Lalana & Varela, 2007: 26; Figs 1-2. [sur de la Bahia de Cienfuegos, 297-351 m, en 

la esponja Pheronema annae ?] 

Spongiocaris semiteres Bruce & Baba, 1973: 155; Figs 1-6. [13 miles S.E. of Durban, South Africa, 230 fms, 

from hexactinellid sponge] 

Spongiocaris yaldwyni Bruce & Baba, 1973: 163; Figs 7-10. [15 miles N, 50°E of Plate Island in the Bay of 

Plenty, northeast of North Island, New Zealand, 320-340 fms, from Regadrella okinoseana] 

Family STeNoPoDIDAe Claus, 1872 

Juxtastenopus Goy, 2010 

= Juxtastenopus Goy, 2010 (type species Engystenopus spinulatus Holthuis, 1946a, by original designation 


Juxtastenopus spinulatus (Holthuis, 1946a) 

= Engystenopus spinulatus Holthuis, 1946a: 45; Plate 4, figs a-b. [Lobetobi Strait, E. of Flores, 8°27'S 

122°54.5'E, 247 m] 

Odontozona Holthuis, 1946a 

= Odontozona Holthuis, 1946a (type species Stenopus ensiferus Dana, 1852a, by original designation, 


Odontozona addaia Pretus, 1990: 343; Figs 1-6. [40°01'57''N 4°11'46''E, littoral marine cave near Port 

Addaia Bay, NE coast of Minorca, Balearic Islands, Spain] 

Odontozona anaphorae Manning & Chace, 1990: 29; Figs 14-15. [off North Point, Ascension Island, 10 m] 

Odontozona crinoidicola Saito & Fujita, 2009: 124; Figs 1-7. [Mizugama, Okinawa islands, Ryukyu Islands, 

3.2 m, associated with Phanogenia gracilis] 

Odontozona edwardsi (Bouvier, 1908b) 

= Richardina Edwardsi Bouvier, 1908b: 888. [au large des côtes marocaines et soudanaises] 

Odontozona ensifera (Dana, 1852a) 

= Stenopus ensiferus Dana, 1852a: 27. [archipelago Viti] 

Odontozona fasciata Okuno, 2003: 167; Figs 1-6. [Shimoji-shima Island, Miyako Group, Ryukyu Islands, 

24°49.6'N 125°08.2'E, 25 m] 

Odontozona foresti Hendrickx, 2002: 406; Figs 1-3. [southern Gulf of California, 25°43.5'N 109°53.7'W, 

1240-1270 m] 

Odontozona libertae Gore, 1981: 153; Figs 4-5. [Elbow Reef, off Key Largo, Monroe County, Florida] 

Odontozona minoica Dounas & Koukouras, 1989: 341; Figs 1-4. [35°29'10''N 24°17'20''E, off north-western 

coast of Crete, 330 m]


Odontozona rubra Wicksten, 1982: 130; Figs 1-2. [Isla Blanca, off Guaymas, Sonora, Mexico, approx. 

27°52'N 110°52'W, 6-9 m] 

Odontozona sculpticaudata Holthuis, 1946a: 37; Plate 2, fig. F; Plate 4, fig. C. [Sape Strait, east of Soembawa, 

70 m] 

Odontozona spinosissima Kensley, 1981: 66; Figs 4-5. [31°59'S 29°22'E, 150-200 m] 

Odontozona spongicola (Alcock & Anderson, 1899) 

= ?Richardina spongicola Alcock & Anderson, 1899: 291. [Andaman Sea, 498 fms, from hexactinellid 

sponge] 

Odontozona striata Goy, 1981: 843; Figs 1-5. [Gulf of Mexico, West of Cabo San Antonio, Cuba] 

Richardina A. Milne-edwards, 1881a 

= Richardina A. Milne-Edwards, 1881a (type species Richardina spinicincta A. Milne-Edwards, 1881a, 

by monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in 

Opinion 712 in 1964) 

Richardina fredericii Lo Bianco, 1903: 250; Plate 8, figs 27-28. [a circa 9 chilometri da Punta Campanella, 

direzione S.E., 1100 m; a circa 6.5 chilometri dai Galli di Positano, direzione E., 950 m; a circa 9 

chilometri dai Galli di Positano, direzione O.S.O., 1100 m; a circa 10 chilometri da Punta Carena, 

direzione E.N.E., 100 m] 

Richardina ohtsukai Saito & Komatsu, 2009: 31; Figs 1-6. [Kerama Islands, N of Nagan-nu Island, 

26°23.15'N 127°30.09'E, 730-728 m] 

Richardina parvioculata Saito & Komatsu, 2009: 39; Figs 7-9. [off Tosa Bay, western Japan, 32°56.45'N 

133°30.52'E, 517 m] 

Richardina spinicincta A. Milne-Edwards, 1881a: 933. [44°48'30''N 7°0'30''E, Golfe de Gascogne, 5100 m] 

Stenopus latreille, 1819 

= Byzenus Rafinesque, 1814 (type species Byzenus scaber Rafinesque, 1814, by monotypy, an invalid 

senior synonym of Stenopus spinosus Risso, 1827, gender masculine; name of genus and of its type 

species suppressed for the purposes of the Principle of Priority but not for those of the Principle of 

Homonymy, placed on the Official Index of Rejected and Invalid Generic (resp. Specific) Names in 

Zoology in Opinion 522 in 1958) 

= Stenopus Latreille, 1819 (type species Palæmon hispidus Olivier, 1811, by monotypy, gender masculine; 

name placed on the Official list of Generic Names in Zoology in Opinion 522 in 1958) 

= Embryocaris Ortmann, 1893 (type species, Embryocaris stylicauda Ortmann, 1893, by monotypy, a 

junior subjective synonym of Palæmon hispidus Olivier, 1811, gender feminine) 

Stenopus chrysexanthus Goy, 1992: 80; Figs 1-4. [Heron Island, Capricorn Group, Queensland, Australia, 

ca. 23°25'S 151°55'E, south west reef flat, algal crest] 

Stenopus cyanoscelis Goy, 1984: 116. [Broadhurst Reef, Off Townsville, Queenland] 

Stenopus devaneyi Goy, 1984: 117. [French Polynesia, Marquesas Islands, Nuka Hiva Island, outer portion 

of Taiohae Baie, W side of islet, Sentinelle de l’Est, 6.2 m, according to Goy & Randall, 1986] 

Stenopus earlei Goy, 1984: 117. [off Makua, O’ahu Island, Hawaiian Islands, 39.6 m, according to Goy & 

Randall, 1986] 

Stenopus goyi Saito, Okuno & Chan, 2009: 111; Figs 1-5. [Donggang fishing port, Pingtung County, Taiwan, 

commercial trawler] 

Stenopus hispidus (Olivier, 1811) (Fig. 11) 

= Squilla Groenlandica Seba, 1759: 54; Plate 21, figs 6-7. [Type locality not indicated; see Davie, 2002 

for status of the older synonyms] 

= Astacus muricatus Olivier, 1791: 346. [mer du Groënland] 

= Cancer (Astacus) longipes Herbst, 1793 [in Herbst, 1791-1796]: 90; Plate 31, fig. 2. [Grönland] 

= Penæus borealis Latreille, 1802: 250. [mers du Nord] 

= Palæmon hispidus Olivier, 1811: 666. [Type locality not indicated] 

= Palæmon ? asper Latreille, 1818: 3; Plate 293, fig. 3. [Type locality not indicated] 

= Embryocaris stylicauda Ortmann, 1893: 85; Plate 6, fig. 4. [Plankton-Expedition der Humboldt-Stiftung, 

südl. Aequatorialstrom JN 235 (0.1°S 44.2°W, 0-400 m)]


Fig. 11. Stenopus hispidus (Olivier, 1811). Photo by Arthur Anker. 

Stenopus pyrsonotus Goy & Devaney, 1980: 781; Figs 1-6. [off Pokai Bay, Oahu Island, Hawaii, 22.9 m] 

Stenopus scutellatus Rankin, 1898: 242; Plate 24, fig. 3. [Silver Cay, New Providence] 

Stenopus spinosus Risso, 1827: 66; Plate 3, fig. 8. [golfe de Nice, régions profondes] 

= Byzenus scaber Rafinesque, 1814: 23. [Sicile; name suppressed for the purposes of the Principle of 

Priority but not for those of the Principle of Homonymy, placed on the Official Index of Rejected 

and Invalid Specific Names in Zoology in Opinion 522 in 1958] 

Stenopus tenuirostris De Man, 1888b: 567; Plate 22a, fig. 5. [Amboina] 

Stenopus zanzibaricus Bruce, 1976a: 90; Figs 1-5. [5°46.6'S 39°23.5'E, Mwenba Island, off N.E. Zanzibar 

Island, 0.5 m] 

Infraorder CARIDeA Dana, 1852b 

Superfamily PASIPHAeoIDeA Dana, 1852a 

Family PASIPHAeIDAe Dana, 1852a 

Alainopasiphaea Hayashi, 1999 

= Alainopasiphaea Hayashi, 1999 (type species Pasiphaea nudipedia Burukovsky, 1993b, by original 

designation and monotypy, gender feminine) 

Alainopasiphaea australis (Hanamura, 1989) 

= Pasiphaea australis Hanamura, 1989: 59; Figs 5-7. [Maria Island area, off east coast of Tasmania, 

southern Australia, 42°42.0'S 148°24.0'E, 345-350 m] 

Alainopasiphaea nudipeda (Burukovsky, 1993b) 

= Pasiphaea nudipeda Burukovsky, 1993b: 35; Figs 1(8-13). [25°28'-25°34'5"S 35°08'-35°00'E, 535-490 m]


Eupasiphae Wood-Mason in Wood-Mason & Alcock, 1893 

= Eupasiphaë Wood-Mason in Wood-Mason & Alcock, 1893 (type species Parapasiphaë (Eupasiphaë) 

Gilesii Wood-Mason, 1892, designated by Holthuis, 1955b, gender feminine) 

Eupasiphae gilesii (Wood-Mason, 1892) 

= Parapasiphaë Gilesii Wood-Mason, 1892: Plate 3, fig. 8; accompanying text in Wood-Mason in 

Wood-Mason & Alcock, 1893: 166 (as Parapasiphaë (Eupasiiphaë) Gilesii). [off Cinque Island, Andaman 

Sea, 650 fms] 

= Eupasiphae rhinocerata Burukovsky, 1977: 473; Figs A-C. [off Morocco, near Meteor Shoal, 30°00'N 

28°43'W, 1400-2060 m] 

Eupasiphae latirostris (Wood-Mason & Alcock, 1891b) 

= Parapasiphaë latirostris Wood-Mason & Alcock, 1891b: 196. [Investigator stn 105 (Arabian Sea, 

15°2'N 72°34'E), 740 fms] 

Eupasiphae paucidentata Crosnier, 1988b: 789; Figs 2c, 3-4, 5b-c. [Madagascar, N.O., 21°26.5S 43°11'E, 810- 

1020 m] 

Eupasiphae serrata (Rathbun, 1902a) 

= Parapasiphæ serrata Rathbun, 1902a: 904. [off Cortez Bank, California, 984 fms] 

= Parapasiphae Grimaldii Coutière, 1911a: 157. [Princesse-Alice stn 3036, off Spain and Portugal, 

36°06'40"N, 10°18'W, 0-4740 m, according to Crosnier, 1988b] 

Glyphus Filhol, 1884 

= Glyphus Filhol, 1884 (type species Glyphus marsupialis Filhol, 1884, by monotypy, gender masculine) 

= Sympasiphaea Alcock, 1901 (type species Sympasiphaea annectens Alcock, 1901, by monotypy, gender 

feminine) 

Glyphus marsupialis Filhol, 1884: 231. [Talisman stn 75 (25°39'N 16°02'W)] 

= Sympasiphaea annectens Alcock, 1901: 63. [Arabian Sea, south-west of Cape Comorin, 487 fms] 

= Sympasiphaea imperialis Terao, 1922: 110; unnumbered figure. [purchased at the market of Uchiura, 

Prov. Idzu] 

Leptochela Stimpson, 1860a 

= Leptochela Stimpson, 1860a (type species Leptochela gracilis Stimpson, 1860a, designated by Kemp, 


= Proboloura Chace, 1976 (type species Leptochela carinata Ortmann, 1893, by original designation 

and monotypy, gender feminine) 

Leptochela (Leptochela) aculeocaudata Paul’son, 1875 

= Lep.[tochela] aculeocaudata Paul’son, 1875: 100; Plate 16, figs 1-1s. [Red Sea] 

Leptochela (Leptochela) bermudensis Gurney, 1939a 

= Leptochela bermudensis Gurney, 1939a: 427; Figs 1-10. [about 7 miles off the south shore, between 

1000 m and surface] 

? = Leptochela carinata Ortmann, 1893: 41 (partim, not Plate 4, fig. 1). [Plankton-Expedition der Humboldt-Stiftung, 

Küstenbank vor der Tocantins-Mündung, JN 236 (0.1°S 45.2°W), 50-100 m] 

Leptochela (Leptochela) chacei Hayashi, 1995: 85; Figs 2-4. [Viet-Nam, Cai Dua] 

Leptochela (Leptochela) crosnieri Hayashi, 1995: 89; Figs 5-7. [New Caledonia, st. Balabio, 20°03.6'S 

164°07.7'E, 13 m] 

Leptochela (Leptochela) gracilis Stimpson, 1860a 

= Leptochela gracilis Stimpson, 1860a: 42. [Sinu ”Kagosima”, in profundis] 

= Leptochela pellucida Boone, 1935: 105; Plates 26-27. [Near Equator, to the South Brother’s Island, 

south entrance of Durian Straits, Dutch East Indies, 29'N by E 104°47'E, 14 fms] 

Leptochela (Leptochela) hawaiiensis Chace, 1976: 15; Figs 11-13. [Hawaii, south coast of Oahu, Diamond 

Head Light, 62°S 3.9'E] 

Leptochela (Leptochela) irrobusta Chace, 1976: 19; Figs 14-18. [Bikini Atoll, northeast end of lagoon at 

Bowditch anchorage, surface light at night]


Leptochela (Leptochela) japonica Hayashi & Miyake, 1969 

= Leptochela japonica Hayashi & Miyake, 1969: 1; Figs 1-2. [Chijiwa Bay, northwestern Kyushu, 65-66 m] 

Leptochela (Leptochela) papulata Chace, 1976: 26; Figs 22-24. [east of Cape Lookout, North Carolina, 

34°35'30"N 75°45'30"W, 59 m] 

Leptochela (Leptochela) pugnax De Man, 1916 

= Leptochela pugnax De Man, 1916: 148. [Bay of Bima, near south fort, 55 m; Tual-anchorage, Kei-islands, 

22 m; Elat, west coast of Great-Kei-island, 27 m] 

Leptochela (Leptochela) robusta Stimpson, 1860a 

= Leptochela robusta Stimpson, 1860a: 43. [Mari sinensi, prof. 20 org; prope insulam ”Loo Choo” 

quoque] 

Leptochela (Leptochela) serratorbita Spence Bate, 1888 (Fig. 12) 

= Leptochela serratorbita Spence Bate, 1888: 859; Plate 139, fig. 1. [Saint Thomas, Virgin Islands, shallow 

water] 

Leptochela (Leptochela) sydniensis Dakin & Colefax, 1940 

= Leptochela sydniensis Dakin & Colefax, 1940: 153; Figs 245-246. [New South Wales] 

? = Leptochela hainanensis Yu, 1936: 87; Figs 1-3. [Hai-kiu-sche] 

Leptochela (Proboloura) carinata Ortmann, 1893 

= Leptochela carinata Ortmann, 1893: 41 (partim); Plate 4, fig. 1. [Plankton-Expedition der Humboldt- 

Stiftung, Küstenbank vor der Tocantins-Mündung, JN 236 (0.1°S 45.2°W), 50-100 m] 

Leptochela (Proboloura) soelae Hanamura, 1987: 21; Figs 6-8. [north-west Australian shelf, 19°05.0'S 

118°53.8'E, 82 m] 

Parapasiphae Smith, 1884 

= Parapasiphaë Smith, 1884 (type species Parapasiphaë sulcatifrons Smith, 1884, designated by Fowler, 

1912, gender feminine; name placed in its corrected spelling on the Official List of Generic Names 

in Zoology in Opinion 470 in 1957) 

= Orphania Spence Bate, 1888 (type species Orphania tenuimana Spence Bate, 1888, a junior subjective 

synonym of Parapasiphaë sulcatifrons Smith, 1884, gender feminine) 

= Dantecia Caullery, 1896 (type species Dantecia Caudani Caullery, 1896, a subjective synonym of 

Parapasiphaë sulcatifrons Smith, 1884, by monotypy, gender feminine) 

Parapasiphae compta Smith, 1884 

= Parapasiphaë compta Smith, 1884: 389. [Albatross stn 2039, 38°19'26"N 68°20'20"W, 2369 fms] 

Fig. 12. Leptochela (Leptochela) serratorbita Spence Bate, 1888. Photo by Arthur Anker.


Parapasiphae cristata Smith, 1884 

= Parapasiphaë cristata Smith, 1884: 388; Plate 5, fig. 3. [Albatross stn 2100, 39°22'N 68°34'30"W, 1628 

fms] 

= Parapasiphaë macrodactyla Chace, 1939: 33. [S.S.E. of Bahia de Guantanamo, Oriente Province, 

Cuba, 19°40'N 75°03'W, 1920 fms] 

Parapasiphae kensleyi Wasmer, 2005: 165; Figs 1-3. [31°N 159°W, 4200 m] 

Parapasiphae sulcatifrons Smith, 1884 

= Parapasiphaë sulcatifrons Smith, 1884: 384; Plate 5, fig. 4; Plate 6, figs 1-7. [Albatross stns 2002, 

37°02'42''N 74°17'36''W, 641 fms; 2034, 39°27'10''N 69°56'20''W, 1346 fms; 2037, 38°53'00''N 69°23'30''W, 

1731 fms; 2072, 41°53'00''N 65°35'00''W, 858 fms; 2074, 41°43'00''N 65°21'50''W, 1309 fms; 2094, 

39°44'30''N 71°04'00''W, 1022 fms; 2097, 37°56'20''N 70°57'30''W, 1917 fms; 2099, 37°12'20''N 

69°39'00''W, 2949 fms; 2105, 37°50'00''N 73°03'50''W, 1395 fms; 2110, 35°12'10''N 74°57'15''W, 516 fms] 

= Orphania tenuimana Spence Bate, 1888: 872; Plate 141, fig. 4. [Challenger stn 45, 38°34'N 72°10'W, 

south-east of New York, 1240 fms] 

= Dantecia Caudani Caullery, 1896: 372, Plate 14, figs 1-11. [Golfe de Gascogne, 46°28'N 7°W, 1710 m] 

= Pasiphaea metriomma Dohrn, 1908: 9. [Valdivia Expedition stn 26 (ca. 1000 mls E of Madeira, 31°59'N 

5°5'W, 1800 m)] 

Pasiphaea Savigny, 1816 

= Pasiphaea Savigny, 1816 (type species A.[lpheus] sivado Risso, 1816, by monotypy, gender feminine) 

= Pasiphaë (Phye) Wood-Mason, 1892 (type species Pasiphaë (Phye) Parapasiphaë Alcocki Wood-Mason 

& Alcock, 1891b, by monotypy, gender feminine) 

Pasiphaea acutifrons Spence Bate, 1888 

= Pasiphæa acutifrons Spence Bate, 1888: 871; Plate 141, fig. 3. [Challenger stns 311, 52°45'30''S 

73°46'0''W, off Port Churruca, Patagonia, 245 fms; 236, 34°58'N 139°29'E, south of Japan, 77 fms] 

= Pasiphæa forceps A. Milne-Edwards, 1891: 51; Plate 6, figs 2-2e. [détroit de Magellan, au sud-est de 

Port-Famine, 326 m] 

Pasiphaea affinis Rathbun, 1902a 

= Pasiphæa affinis Rathbun, 1902a: 905. [near Cortez Bank, California, 984 fms] 

Pasiphaea alcocki Wood-Mason & Alcock, 1891b 

= Parapasiphaë Alcocki Wood-Mason & Alcock, 1891b: 196. [Bay of Bengal, 16°11'15"N 82°30'30"W, 

922 fms] 

Pasiphaea americana Faxon, 1893 

= Pasiphaeia (sic) cristata americana Faxon, 1893: 208. [Gulf of Panama, 7°32'36"N 79°16'W, 523 m; 

lectotype designation by Hayashi, 2004] 

Pasiphaea amplidens Spence Bate, 1888 

= Pasiphæa amplidens Spence Bate, 1888: 870; Plate 141, fig. 2. [Challenger stn 34°58'N 139°29'E, 775 

fms] 

= Pasiphaea vereschhaka Burukovsky, 1993b: 33; Fig. 1(1-7). [12°22'S 53°02'E, 1045-1050 m] 

Pasiphaea arabica Timofeev, 1997: 142; Fig. 1. [Arabian Sea, 16°32'2''N 64°25'5''E, collecting horizon 110- 

100 m, bottom at 3600 m] 

Pasiphaea balssi Burukovsky & Romensky, 1987: 52; Figs 1, 2(9), 4. [South Atlantic, 41°55'S, 0°00'E, 910 m] 

Pasiphaea barnardi Yaldwyn, 1971: 86 [Cook Strait, 41°42.5'S 175°9'E, ca. 500 fms] 

Pasiphaea berentsae Kensley, Tranter & Griffin, 1987: 294; Figs 7-9. [New South Wales, northeast of Norah 

Head, 33°11'S 152°24'E to 33°09'S 152°25'E, 720 m] 

Pasiphaea burukovskyi Wasmer, 1993: 79; Figs 22-24. [40°22'S 168°25'E to 40°26'S 168°30'E, 600-750 m] 

Pasiphaea chacei Yaldwyn, 1962: 18; Figs 1-17. [33°36'11"N 118°32'13"W – 33°30'52"N 118°22'48"W, 475 

fms] 

Pasiphaea corteziana Rathbun, 1902a 

= Pasiphæa corteziana Rathbun, 1902a: 905. [near Cortez Bank, California, 776 fms] 

Pasiphaea cristata Spence Bate, 1888 

= Pasiphæa cristata Spence Bate, 1888: 865; Plate 140; Plate 141, fig. 1. [Challenger stn 173, 19°9'35"S 

179°41'50"E, off Matuka, Fiji Islands, 315 fms]


Pasiphaea crosnieri Hayashi, 2004: 334; Figs 7-9. [Philippines, 13°40.7'N 120°30.0'E, 685-757 m] 

Pasiphaea debitusae Hayashi, 1999: 281; Figs 8-10. [Indonesia, 05°15'S 133°01'E, 605-576 m] 

Pasiphaea diaphana Burukovsky & Romensky, 1980: 1097. [South-East Atlantic, 31°48'S 02°18'E, 1100 m] 

Pasiphaea dofleini Schmitt, 1932a: 333; Fig. 1. [Punta Arenas, now Magelhanes, Chile] 

Pasiphaea ecarina Crosnier, 1969: 537; Figs 19, 23-36. [Congo, 5°08'S 11°22'E, 795-805 m] 

Pasiphaea emarginata Rathbun, 1902a 

= Pasiphæa emarginata Rathbun, 1902a: 905. [Gulf of California, 857 fms] 

Pasiphaea faxoni Rathbun, 1904: 22; nomen novum for Pasiphaea acutifrons sensu Faxon, 1895 nec Spence 

Bate, 1888 [Albatross stns 3402 (0°57'30"S 89°03'30"W), 421 fms; 3403 (0°58'30" 89°17'00"W) 384 fms 

and 3406 (0°16'00"S 90°21'30"W), 551 fms] 

Pasiphaea flagellata Rathbun, 1906 

= Pasiphæa flagellata Rathbun, 1906: 928; Fig. 78; Plate 23, fig. 2. [Kaiwi Channel, Hawaii, 411-442 

fms] 

Pasiphaea fragilis Hayashi, 1999: 285; Fig. 11. [Loyalty Islands, 20°23.0'N 166°20.1'E, 780 m] 

Pasiphaea gelasinus Hayashi & Yaldwyn, 1998: 512; Figs 1-3. [South Indian Ocean, Seamounts off Saint 

Paul and Amsterdam Islands, 37°37.8'S, 77°51.8'E, 730-905 m] 

Pasiphaea gracilis Hayashi, 1999: 290; Figs 15-17. [New Caledonia, 23°08'S 166°51'E, 850 m] 

Pasiphaea grandicula Burukovsky, 1976: 17; Fig. 1. [South Atlantic, 26°11'S, 06°02'8"E, 1150 m] 

Pasiphaea hoplocerca Chace, 1940: 124; Figs 4-5. [Bermuda Oceanographic Expedition Nets 713, 753 and 

778 (32°12'N 64°36'W), 700 fms] 

Pasiphaea japonica Omori, 1976: 250; Figs 1-2. [Off Iwase, Toyama Bay, 0-300 m] 

Pasiphaea kapala Kensley, Tranter & Griffin, 1987: 297; Figs 10-12. [New South Wales, northeast of Norah 

Head, 33°08'S 152°27'E to 33°10'S 152°24'E, 580 m] 

Pasiphaea kaiwiensis Rathbun, 1906 

= Pasiphæa kaiwiensis Rathbun, 1906: 927; Fig. 76; Plate 23, fig. 4. [Kaiwi Channel, 343-337 fms] 

Pasiphaea korzuni Burukovsky, 1995: 121; Figs 1-5. [Aden Bay, near Ras-Fartak Island, depth unknown] 

Pasiphaea laevis Hayashi, 1999: 287; Figs 12-14. [Indonesia, Moluccas, 06°08'S 132°45'E, 390-502 m] 

Pasiphaea ledoyeri Hayashi, 2006a: 215; Figs 6-8. [South Indian Ocean, 46°33.8'S 41°23.7'E, 660-911 m, in 

stomach of Dissostichus eleginoides] 

Pasiphaea levicarinata Hanamura, 1994: 167; Figs 1-2. [north-western Australia, 18°30'S 117°23'E, 530 m] 

Pasiphaea liocerca Chace, 1940: 122; Figs 2-3. [Bermuda Oceanographic Expeditions Net 1160 (32°12'N 

64°36'W), 800 fms] 

Pasiphaea longitaenia Kensley, Tranter & Griffin, 1987: 301; Figs 13-14. [New South Wales, east of Long 

Reef Point, 33°43'S 151°56'E to 33°39'S 151°58'E, 626-637 m] 

Pasiphaea magna Faxon, 1893: 209. [Albatross stn 3384 (off Panama, 7°31'30"N, 79°15'00"W), 458 fms] 

Pasiphaea major Hayashi, 2006a: 219; Figs 9-11. [Norfolk Ridge, 25°09'S 168°53'E, 1173-1340 m] 

Pasiphaea marisrubri Iwasaki, 1989: 178; Figs 1-3. [Central Red Sea, 21°14.80'N 37°15.40'E – 21°18.20'N 

37°12.00'E, 0-220 m] 

Pasiphaea mclaughlinae Hayashi, 2006b: 342; Figs 1-3. [SE coast of Taiwan, 22°13.8'N 121°01.8'E, 1222-1226 m] 

Pasiphaea meiringnaudei Kensley, 1977: 34-37; Figs 11-14. [Meiring Naude stn 107 (South Africa, off Natal 

coast, 28°37.8'S 32°38.4'E, 1200-1000 m)] 

Pasiphaea merriami Schmitt, 1931b 

= Pasiphæa merriami Schmitt, 1931b: 391. [south of the Dry Tortugas, 253-283 fms] 

= Pasiphaea nishiei Iwasaki, 1990: 190; Figs 1-2. [off Jamaica, 17°34.3'N, 76°03.3'W, 1490-1504 m] 

Pasiphaea multidentata Esmark, 1866: 259. [Bjørumsfjorden omtrent 1/8 Miil fra Namsos] 

= P.[asiphaë] norvegica M. Sars, 1866a: 260. [nomen nudum] 

= Pasiphaë norvegica M. Sars, 1866b: 314. [Christianiafjorden; fully illustrated in M. Sars, 1868] 

= Pasiphaë (Phye) sicula Riggio, 1896 [in Riggio, 1895-1896]: 41; Plate 1 (1865, no legend), Figs 2a-b. 

[proviene dal mare di Augusta] 

Pasiphaea natalensis Burukovsky & Romensky, 1982: 1797; Figs 1-6. [south-western part of the Indian 

Ocean, 33°48.5'S 44°30.8'E, 900-910 m] 

Pasiphaea notosivado Yaldwyn, 1971: 86. [Cook Strait, 41°32'S 174°56'E, 50-100 fms] 

Pasiphaea orientalis Schmitt, 1931a: 267; Plate 32, figs 1, 5. [Ryukyusho, Takao, Formosa]


Pasiphaea oshoroae Komai & Amaoka, 1993: 367; Figs 1-3. [Off Aleuten Islands, 49°59.9'N 176°55.4'W, 

0-400 m] 

Pasiphaea pacifica Rathbun, 1902a 

= Pasiphæa pacifica Rathbun, 1902a: 905. [off Point Sur, California, 328 fms] 

Pasiphaea philippinensis Hayashi, 1999: 277; Figs 5-7. [Philippines, 13°39'N 120°43'E, 520-550 m] 

Pasiphaea planidorsalis Hayashi, 2004: 348; Figs 13-14. [New Caledonia, 19°42.4'S 158°50.8'E, 772-756 m] 

Pasiphaea poeyi Chace, 1939: 31. [Bahia de Cochinos, Santa Clara Province, Cuba, 22°07'N 81°08'W, 220- 

275 fms] 

Pasiphaea princeps Smith, 1884 

= Pasiphaë princeps Smith, 1884: 381; Plate 5, fig. 2. [Albatross stn 2095, 39°29'N 70°58'40"W, 1451 fms] 

Pasiphaea propinqua De Man, 1916: 147. [06°11'N 120°37.5'E, off the Sulu Islands, 450 m] 

Pasiphaea pseudacantha Hayashi, 2004: 353; Figs 16-18. [Kai Islands, Indonesia, 06°08'S 132°45'E, 390-502 m] 

Pasiphaea rathbunae (Stebbing, 1914a) 

= Phye rathbunæ Stebbing, 1914a: 295; Plate 31. [Scotia stn 450, 48°00'S 9°50'W, 1332 fms] 

Pasiphaea romenskyi Burukovsky, 1995: 123; Figs 6-11. [Ecuador Seamount, 00°23'S, 56°02'E, 200-235 m] 

Pasiphaea scotiae (Stebbing, 1914a) 

= Phye scotiæ Stebbing, 1914a: 294; Plate 30. [Scotia stns 417, 71°22'S 16°34'W, 1410 fms; 422, 68°32'S 

12°49'W, from surface to 600 fms] 

= Pasiphaea longispina Lenz & Strunck, 1914: 315; Plate 19, figs 1-11. [65°15'S 80°19'E, also in der Nähe 

des Eisrandes, im Magen eines Pinguins, Aptenodytes forsteri, gefunden] 

Pasiphaea semispinosa Holthuis, 1951a: 9; Fig. 1. [off Angola, 7°55'S 12°38'E, 235-460 m] 

Pasiphaea sinensis Hayashi & Miyake, 1971: 39; Fig. 1. [East China Sea, 29°27.2'N 128°16.2'E – 29°33.3'N 

128°23.5'E, 1065-1075 m] 

Pasiphaea sirenkoi Burukovsky, 1987: 37; Figs 1-5. [SE Indian Ocean, 11°27.5'N 109°56.2'E, 600 m] 

Pasiphaea sivado (Risso, 1816) 

= A.[lpheus] sivado Risso, 1816: 93; Plate 3, fig. 4. [sur la plage de Nice] 

= P.[asiphaea] Savignyi H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 426. [Patrie inconnue] 

= Pasiphæa brevirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: Plate 54bis, fig. 2. 

[Type locality not indicated] 

? = Pasiphæa distincta Guérin-Méneville, 1844 [in Guérin-Méneville, 1829-1844]: 16, Plate 22, fig. 3. 

(as Pasiphæa sivado). [la Méditerranée] 

= Pasiphaea neapolitana Hope, 1851: 44. [Type locality not indicated] 

Pasiphaea tarda Krøyer, 1845: 453. [Type locality not indicated] 

= Pasiphæa principalis Sund, 1913: 6; Figs 5-7, 9a-f. [Sognefjord; Øxsund (E. of the Lofoten isles, about 

67.5°N) 

Pasiphaea telacantha Hayashi, 2004: 363; Figs 22-24. [New Caledonia, 23°08'S 166°51'E, 850 m] 

Pasiphaea timofeevi Burukovsky, 1993b: 35; Fig. 1(14-19). [Aden Bay, Socotra Island, 12°20'N 53°09'W, 515- 

480 m] 

Pasiphaea truncata Rathbun, 1906 

= Pasiphæa truncata Rathbun, 1906: 928; Fig. 7; Plate 23, fig. 5. [vicinity of Modu Manu, 293-800 fms] 

Pasiphaea unispinosa Wood-Mason, 1892 

= Pasiphaë unispinosa Wood-Mason, 1892: Plate 3, fig. 7; accompanying text as Wood-Mason in 

Wood-Mason & Alcock, 1893: 163. [7 miles SE by S of Ross Island, Andaman Sea, 265 fms] 

Pasiphaea westindica Tchesunov, 1984: 993; Fig. 1. [Atlantic Ocean, 12°48'N 62°07'W, 1500 m] 

Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893 

= Psathyrocaris Wood-Mason in Wood-Mason & Alcock, 1893 (type species Psathyrocaris fragilis 

Wood-Mason in Wood-Mason & Alcock, 1893, by monotypy, gender feminine) 

Psathyrocaris fragilis Wood-Mason in Wood-Mason & Alcock, 1893: 171; Plates 10-11. [15°56'50"N 

81°30'30"E, 240 fms] 

= Psathyrocaris infirma var. atlantica Caullery, 1896: 374; Plate 14, figs 12-15. [Golfe de Gascogne, 

44°39'N, 2°10'W, 800 m]


Psathyrocaris hawaiiensis Rathbun, 1906: 928; Fig. 79. [vicinity of Modu Manu, 876 fms] 

Psathyrocaris infirma Alcock & Anderson, 1894: 159. [Investigator stn 116, Andaman Sea, 11°25'5''N 

92°47'6''E, 405 fms] 

Psathyrocaris platyophthalmus Alcock & Anderson, 1894: 158. [Investigator stn 124, Laccadive Sea, 

10°47'45"N 72°40'20"E, 705 fms] 

Psathyrocaris plumosa Alcock & Anderson, 1894: 159. [Investigator stn 128, off Ceylon, 6°58'N 77°26'50''E, 

902 fms] 

Superfamily oPloPHoRoIDeA Dana, 1852a 

Family ACANTHePHyRIDAe Spence Bate, 1888 

Acanthephyra A. Milne-edwards, 1881b (Fig. 13) 

= Ephyra Roux, 1831 (an invalid junior homonym of Ephyra Peron & Leseur, 1810 (Coelenterata), 

type species A.[lpheus] Pelagicus Risso, 1816, designated by Kingsley, 1880, gender feminine; name 

placed on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in 

1955) 

= Miersia Kingsley, 1880 (nomen novum for Ephyra Roux, 1831, gender feminine; name suppressed 

for the purposes of the Principle of Priority but not for those of the Principle of Homonymy, placed 

on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 359 in 1955) 

= Acanthephyra A. Milne-Edwards, 1881b (type species Acanthephyra armata A. Milne-Edwards, 

1881b, by original designation, gender feminine; name placed on the Official List of Generic Names 


= Bentheocaris Spence Bate, 1888 (type species Bentheocaris stylorostratis Spence Bate, 1888, designated 

by Holthuis, 1955b, gender feminine) 

= Acanthephyropsis Riggio, 1895 [in Riggio, 1895-1896] (type species Acanthephyra pulchra A. Milne- 

Edwards, 1890, a junior subjective synonym of Acanthephyra eximia Smith, 1884, gender feminine) 

= Hoplocaricyphus Coutière, 1907a (type species Hoplocaricyphus similis Coutière, 1907a, probably a 

subjective synonym of A.[lpheus] Pelagicus Risso, 1816, gender masculine) 

Fig. 13. Acanthephyra spec. Photo by Tin-Yam Chan.


Acanthephyra acanthitelsonis Spence Bate, 1888: 745; Plate 125, fig. 3. [Challenger stns 106, 1°47'N 24°26'W, 

Atlantic, south-west of Sierra Leone, 1850 fms; 107, 1°22'N, 26°36'W, Atlantic, south-west of Sierra 

Leone, 1500 fms] 

Acanthephyra acutifrons Spence Bate, 1888: 749 (partim); Plate 126, fig. 3. [off Kepulauan Aru, Indonesia, 

5°41'S 134°04'30"E, 1463 m; type locality restricted by Kemp, 1906a] 

Acanthephyra armata A. Milne-Edwards, 1881b: 12. [près de Sainte-Lucie, 422 brasses] 

Acanthephyra brevicarinata Hanamura, 1984: 65; Figs 1-2. [13°28.7'N, 119°53.9'W, 1617-0 m] 

Acanthephyra brevirostris Smith, 1885a: 504. [Albatross stns 5448, 37°12'20"N 69°39'00"W, 2949 fms; 5449, 

39°22'00"N, 68°34'30"W, 1686 fms; 5673, 37°50'00"N, 73°03'50"W, 1395 fms] 

= Hymenodora duplex Spence Bate, 1888: 843; Plate 136, fig. 3. [Challenger stn 147, 46°16'S 48°27'E, off 

Marion Island, 1600 fms] 

Acanthephyra carinata Spence Bate, 1888: 748; Plate 126, fig. 2. [Challenger stn 310, 51°27'30"S 74°03'00"W, 

Sarmiento channel, Patagonia, 400 fms] 

= Acanthephyra approxima Spence Bate, 1888: 755; Plate 126, fig. 8. [Challenger stn 310, 51°27'30"S 

74°03'00"W, Sarmiento Channel, Patagonia, 400 fms] 

Acanthephyra chacei Krygier & Forss, 1981: 96; Figs 1-2. [44°45.2'N 127°44.0'W, 0-2600 m] 

Acanthephyra cucullata Faxon, 1893: 206. [Albatross stn 3381 (North Pacific Ocean, Colombia, 4°56'00"N, 

80°52'30"W), 1772 fms] 

Acanthephyra curtirostris Wood-Mason & Alcock, 1891b: 195. [Investigator stns 100 (11°12'47"N 74°25'30"E), 

840 fms; 104 (16°55'41"N 83°21'18"E), 1000 fms] 

= Acanthephyra acutifrons Spence Bate, 1888: 749 (partim) (nec Plate 126, fig. 3) 

Acanthephyra eximia Smith, 1884: 376. [Albatross stn 2111, 35°09'50"N 74°57'40"W, 938 fms] 

= Acanthephyra angusta Spence Bate, 1888: 737; Plate 124, fig. 6. [Challenger stn 194, 4°34'00"S 

129°57'30"E, off Banda Island, 200 fms] 

= Acanthephyra edwardsii Spence Bate, 1888: 747; Plate 126, fig. 1. [Challenger stn 126, 10°46'S 36°08'W, 

south of Pernambuco, 770 fms] 

= Acanthephyra brachytelsonis Spence Bate, 1888: 753; Plate 126, fig. 7. [Challenger stns 170, 29°55'S 

178°14'W, off the Kermadec Islands, 520 fms; 170A, 29°45'S 178°11'W, north of the Kermadec Islands, 

630 fms; 171, 28°33'S 177°50'W, north of the Kermadec Islands, 600 fms; 194, 4°34'N 

129°57'30''E, off Banda Island, 200 fms; 214, 4°33'N 127°6'E, south of the Philippine Islands, 500 fms; 

232, 35°11'N 139°28'E, Japan, 345 fms; 236, 34°58'N, 139°29'E, south of Japan, 775 fms; 318, 42°32'S 

56°29'W, north of the Falkland Islands, 2040 fms] 

= Acanthephyra pulchra A. Milne-Edwards, 1890: 163. [au large de Monaco, 1650 m] 

= Acanthephyra frontieri Crosnier, 1987b: 699; Fig. 1. [Madagascar, 13°02'S 48°02'E, 1000-1525 m] 

Acanthephyra faxoni Calman, 1939 

= Acanthephyra Faxoni Calman, 1939: 191; Fig. 1. [John Murray Expedition stn 184, Gulf of Aden 

(14°36'06''N 51°00'18''E - 14°38'42''N 50°57'42''E), 1270 m] 

Acanthephyra fimbriata Alcock & Anderson, 1894 

= Acanthephyra armata var. fimbriata Alcock & Anderson, 1894: 156. [Bay of Bengal [off Madras, 

12°50'N 81°30'E], 475 fms; off the Malabar coast [off Goa, 15°29'N 72°41'E], 559 fms; illustrated in 

Wood-Mason, 1892: Plate 3, fig. 1. (as Acanthephyra armata var.)] 

Acanthephyra indica Balss, 1925a: 264; Figs 34-35. [Valdivia stn 215, 7°01'N 85°56'E, 0-2500 m, zwischen 

Sumatra und Ceylon] 

Acanthephyra kingsleyi Spence Bate, 1888: 751; Plate 126, fig. 4. [Challenger stn 104, 2°25'N 20°1'W, Atlantic, 

south-west of Sierra Leone, 2500 fms] 

= Acanthephyra sexspinosa Kemp, 1939: 575. [Central and South Atlantic from 17°N to 18°S] 

Acanthephyra media Spence Bate, 1888: 736; Plate 124, fig. 5. [Challenger stn 207, 12°21'N 122°15'E, off 

Tablas Island, 700 fms] 

= Acanthephyra media var. obliquirostris De Man, 1916: 150. [0°17.6'S 129°14.5'E, Halmaheira Sea, 1855 m] 

Acanthephyra pelagica (Risso, 1816) 

= A.[lpheus] Pelagicus Risso, 1816: 91; Plate 2, fig. 7. [la mer de Nice; neotype listing by Fransen, 

Holthuis & Adema, 1997 not valid] 

= Ephyra Haeckelii von Martens, 1868: 52; Plate 1, Figs 7a-b. [Messina]


= Acanthephyra Agassizzi mediterranea Riggio, 1900: 20. [Mare di Messina] 

= Acanthephyra rectirostris Riggio, 1900: 20. [Mare di Messina] 

= Acanthephyra purpurea var. multispina Coutière, 1905a: 10. [Princesse-Alice stns 1797 (32°18'N 

23°58'W), 0-2000 m; 1851 (46°15'N 7°09'W), 0-3000 m; 1991 (42°53'N 8°22'E), 0-2000 m; 1583 (47°36'N 

7°38'W, 1490 m) and 1639 (36°17'N 28°53'W, 0-3000 m)] 

= Acanthephyra parva multidens Coutière, 1905a: 15; Fig. 5. [Princesse-Alice stns 1856 (36°46'N 

26°41'W), 0-3250 m; 1834 (37°28'N 25°53'30"W), 0-1000 m] 

Acanthephyra prionota Foxton, 1971: 35; Figs 1-2. [East African coast, 05°39'S 46°22'E, 1900-1850-(0) m] 

Acanthephyra purpurea A. Milne-Edwards, 1881a: 933. [au large des îles Berlingues par 2.590 mètres] 

= Miersia Agassizii Smith, 1882: 67; Plate 11, Figs 5-7; Plate 12, Figs 1-4. [Blake stns 305, 41°23'15''N 

65°51'25''W, 810 fms; 323, 33°19'0''N 76°12'30''W, 457 fms; 330, 31°41'0''N 74°35'0''W, 1047 fms] 

= Acanthephyra abyssorum Filhol, 1885b: Plate 7. [Type locality not indicated, 4000 m] 

= Acanthephyra parva paucidens Coutière, 1905a: 16; Fig. 5(3). [Princesse-Alice stn 1856 (36°17'N, 

28°53'W), 0-3250 m] 

= Hoplocaricyphus similis Coutière, 1907a: 7; Fig. 1. [28°04'N, 16°49'30"W, 0-1000 m] 

Acanthephyra quadrispinosa Kemp, 1939: 576. [Indo-Pacific from the E. African coast to 163°W, and from 

25°N to 42°S; South Atlantic from 32°S to 40°S] 

= Acanthephyra batei Stebbing, 1905: 107; Plate 24B; nec Faxon, 1895. [Cape Point Lighthouse, S 83°E, 

35.5 miles, 360 fms] 

Acanthephyra rostrata (Spence Bate, 1888) 

= Hymenodora rostrata Spence Bate, 1888: 846; Plate 136, fig. 4. [Challenger stns 184, 12°8'S 145°10'E, 

near Torres Strait, 1400 fms; 205, 16°42'N 119°22'S, Philippine Islands, 1050 fms; 245, 36°23'N 

174°31'E, North Pacific, 2775 fms] 

Acanthephyra sanguinea Wood-Mason in Wood-Mason & Alcock, 1892: 358. [Laccadive Sea, 9°53'34''N 

75°16.5'E, 1091 fms; Laccadive Sea, 8°23'N 75°47'E, 738 fms; Bay of Bengal, 12°58'N 88°52'17''E, 

1748 fms] 

= Acanthephyra kempii Balss, 1914a: 595. [70°01'N 85°56'E, 0-2500 m] 

Acanthephyra sibogae De Man, 1916 

= Acanthephyra (Meningodora) sibogae De Man, 1916: 149. [5°26'S 121°18'E, entrance of Gulf of Boni, 

1944 m] 

Acanthephyra sica Spence Bate, 1888: 739; Plate 125, fig. 1. [Challenger stns 168, 40°28'S 177°43'E, off 

New Zealand, 100 fms; 40, 34°51'N 68°30'W, north-west of Bermuda, 2675 fms; 159, 47°25'S 

130°22'E, south of Australia, 2150 fms; 169, 37°34'S 179°22'E, near New Zealand, 700 fms; 170, 

29°55'S 178°14'W, off the Kermadec Islands, 520 fms; 170A, 29°45'S 178°11'W, near the Kermadec 

Islands, 630 fms; 181, 13°50'S 151°49'E, between Australia and the Solomon Islands, 2440 fms; 194, 

4°34'0''S 129°57'30''E, off Banda Island, 200 fms; 230, 26°29'N 137°57'E, south of Japan, 2425 fms; 

235, 34°7'N 138°0'E, south of Japan, 565 fms; 318, 42°32'S 56°29'W, north of the Falkland Islands, 

2040 fms] 

Acanthephyra smithi Kemp, 1939: 577. [Indo-Pacific from the East African coast to mid-Pacific in 131°W; 

at the western end of its range it extends to 14°S and at the eastern end from 20°N to 24°S] 

Acanthephyra stylorostratis (Spence Bate, 1888) 

= Bentheocaris stylorostratis Spence Bate, 1888: 726; Fig. 72; Plate 123, fig. 4. [Challenger stn 13, 21°38'N 

44°39'W, Mid North Atlantic, 1900 fms] 

? = Bentheocaris exuens Spence Bate, 1888: 724; Plate 123, fig. 3. [Challenger stn 285, 32°36'S 137°43'W, 

South Pacific Ocean, 2357 fms] 

Acanthephyra tenuipes (Spence Bate, 1888) 

= Tropiocaris tenuipes Spence Bate, 1888: 836; Plate 136, fig. 2. [Challenger stn 184, 12°8'S 145°10'E, 1400 

fms] 

= Acanthephyra gracilipes Chace, 1940: 149; Figs 26-27. [Bermuda Oceanographic Expedition Net 689 

or 734 (approx. 32°12'N 64°36'W), 1000 fms] 

Acanthephyra trispinosa Kemp, 1939: 577. [west coast of Central America from 7°N to 4°S, extending 

westwards to 116°W]


Ephyrina Smith, 1885a 

= Ephyrina Smith, 1885a (type species Ephyrina Benedicti Smith, 1885a, by monotypy, gender feminine) 

= Calymarina Spence Bate, 1888 (nomen nudum, gender feminine) 

= Tropirinus Spence Bate, 1888 (nomen nudum, gender masculine) 

= Tropiocaris Spence Bate, 1888 (type species Tropiocaris planipes Spence Bate, 1888, a junior subjective 

synonym of Ephyrina Benedicti Smith, 1885a, gender feminine) 

Ephyrina benedicti Smith, 1885a 

= Ephyrina Benedicti Smith, 1885a: 506. [Albatross stn 2083, 40°26'40''N 67°5'15''W, 959 fms] 

= Tropiocaris planipes Spence Bate, 1888: 835; Plate 136, fig. 1. [Challenger stn 230, 26°29'N 137°57'E, 

2425 fms] 

Ephyrina bifida Stephensen, 1923: 58; Fig. 18. [45°37'N 7°03'W, 4300 m of water] 

Ephyrina childressi Chace, 1986: 34; Fig. 17. [Halmahera Sea, Indonesia, 0°33'42"S 128°52'06"E, 950-1200 m] 

Ephyrina figueirai figueirai Crosnier & Forest, 1973 

= Ephyrina figueirai Crosnier & Forest, 1973: 73; Figs 20b, 21g-h, 22c-d, 23. [Bay of Biscay, 47°12'30"N 

6°48'W, 2120-2350 m] 

Ephyrina figueirai spinicauda Lin & Chan, 2001: 184; Figs 1, 2a, d-e, 3. [N.E. Taiwan, I-Lan County, fishing 

port at Tai-Shi, about 600 m] 

Ephyrina hoskynii Wood-Mason & Alcock, 1891b 

= Ephyrina Hoskynii Wood-Mason & Alcock, 1891b: 194. [Investigator stn 105 (Arabian Sea, 15°2'N 

72°34'E), 740 fms] 

Ephyrina ombango Crosnier & Forest, 1973: 68; Figs 20a, 21a-f, 22a-b. [off São Tomé, Gulf of Guinea, 

0°30'N 6°30'E, 0-1000 m in total depth of 2900 m] 

Heterogenys Chace, 1986 

= Heterogenys Chace, 1986 (type species Acanthephyra microphthalma Smith, 1885a, by original designation 


Heterogenys microphthalma (Smith, 1885a) 

= Acanthephyra microphthalma Smith, 1885a: 502. [Albatross stn 2224, 36°16'30"N 68°21'00"W, 2574 fms] 

= Acanthephyra longidens Spence Bate, 1888: 735; Plate 124, fig. 4. [Challenger stns 198, 2°55'N 124°53'E, 

near the Philippine Islands, 2150 fms; 285, 32°36'S 137°43'W, South Pacific Ocean, 2375 fms] 

Heterogenys monnioti Crosnier, 1987b: 704; Fig. 3. [29°50.9'S 48°35.5'E, 3668-3800 m] 

Hymenodora G.o. Sars, 1877 

= Hymenodora G.O. Sars, 1877 (type species Pasiphaë glacialis Buchholz, 1874, by monotypy, gender 

feminine) 

Hymenodora acanthitelsonis Wasmer, 1972: 87; Figs 1-8. [45°18.0'N 125°43.2'W to 45°17.2'N 125°48.3'W, 

0-2560 m] 

Hymenodora frontalis Rathbun, 1902a: 904. [west of Unalaska, 322 fms] 

Hymenodora glacialis (Buchholz, 1874) 

= Pasiphaë glacialis Buchholz, 1874: 279; Plate 1, Figs 2-2g. [an der Oberfläche des Meeres, also in der 

Nähe des 74.° nördl. Br. in beträchtlicher Entfernung von der Grenze des Packeises] 

= Hymenodora glauca Spence Bate, 1888: 847; Plate 137, fig. 1. [Challenger stns 159, 47°25' 130°22'E, 

south of Australia, 2150 fms; 215, 4°19'N 130°15'E, south of the Philippines, 2550 fms] 

= Hymenodora mollicutis Spence Bate, 1888: 848; Plate 137, fig. 2. [Challenger stns 104, 2°25'N 20°1'W, 

Atlantic, 2500 fms; 87, 25°49'N 20°55'W, near the Canary Islands, 1675 fms; 133, 35°41'S 20°55'W, 

near Tristan da Cunha, 1900 fms; 156, 62°26'S 95°44'S, 1975 fms; 157, 53°55'S 108°35'S (sic), 1950 fms; 

318, 42°32'S 56°29'W, 2040 fms] 

Hymenodora gracilis Smith, 1886a: 680; Plate 12, fig. 6. [Albatross stns 2036, 38°52'40''N 69°24'40''W, 1735 

fms; 2083, 40°26'40''N 67°05'15''W, 959 fms; 2095, 39°29'00''N 70°58'40''W, 1342 fms; 2099, 37°12'20''N 

69°36'00''W, 2949 fms; 2100, 39°22'00''N 68°34'430''W, 1628 fms; 2101, 39°18'30''N 68°24'00''W, 1686 


70°10'30''W, 1122 fms]


Kemphyra Chace, 1986 

= Kemphyra Chace, 1986 (type species Notostomus corallinus A. Milne-Edwards, 1883, by original 


Kemphyra corallina (A. Milne-Edwards, 1883) 

= Notostomus corallinus A. Milne-Edwards, 1883: Plate 32. [Travailleur dragage 20 (41°30'N 11°40'W), 

2080 m] 

= Acanthephyra valdiviae Balss, 1914a: 595. [30°6'S 87°50'E, 0-2070 m] 

Meningodora Smith, 1882 

= Meningodora Smith, 1882 (type species Meningodora mollis Smith, 1882, by monotypy, gender feminine) 

Meningodora compsa (Chace, 1940) 

= Notostomus compsus Chace, 1940: 156; Figs 31-32. [Bermuda Oceanographic Expedition Net 748 

(32°12'N 64°36'W), 900 fms] 

Meningodora longisulca Kikuchi, 1985: 191; Figs 1-14. [23°08.1'N 149°33.8'E, 0-975 m] 

Meningodora marptocheles (Chace, 1940) 

= Notostomus marptocheles Chace, 1940: 158; Figs 33-34. [N.E Providence Channel, Bahamas, 25°29'N 

77°18'W, 875 fms of wire] 

Meningodora miccyla (Chace, 1940) 

= Notostomus miccylus Chace, 1940: 161; Figs 35-36. [north of Little Inagua Island, Bahamas, 21°44'N 

72°43'W, 1167 fms of wire] 

Meningodora mollis Smith, 1882: 74; Plate 11, Figs 8-9; Plate 12, Figs 5-9. [Blake stn 328, 34°28'25"N 

75°22'50"W, 1632 fms] 

= Notostomus fragilis Faxon, 1893: 207. [Albatross stn 3371 (North Pacific Ocean, Costa Rica, 5°26'20"N, 

86°55'00"W), 770 fms] 

Meningodora vesca (Smith, 1886a) 

= Notostomus viscus Smith, 1886b: 189. [nomen nudum] 

= Notostomus vescus Smith, 1886a: 676; Plate 12, fig. 5. [Albatross stn 2099, 37°12'20"N 69°39'00"W, 

2949 fms] 

= Acanthephyra brevirostris Spence Bate, 1888: 751; Plate 126, Figs 5-6; nec Smith, 1885a. [Challenger 

stn 107, 1°22'N 26°36'W, Atlantic, south-west of Sierra Leone, 1500 fms] 

= Acanthephyra batei Faxon, 1895: 167. [nomen novum for A. brevirostris Spence Bate, 1888] 

= Acanthephyra parvirostris Coutière, 1911a: 157. [erroneous spelling of A. brevirostris] 

Notostomus A. Milne-edwards, 1881b 

= Notostomus A. Milne-Edwards, 1881b (type species Notostomus gibbosus A. Milne-Edwards, 1881b, 

by monotypy, gender masculine) 

Notostomus auriculatus Barnard, 1950: 670; Fig. 124h-i. [off Cape Point, 800 fms] 

Notostomus crosnieri Macpherson, 1984: 54; Figs 6a, 7a, 8a-b. [02°33.5'S, 05°43.2'E, 4088 m] 

Notostomus distirus Chace, 1940: 166; Figs 39-40. [Bermuda Oceanographic Expedition Net 1281 (32°12'N 

64°36'W), 1000 fms] 

Notostomus elegans A. Milne-Edwards, 1881b: 8. [la mer des Antilles, 24°36'N 84°05'W, 955 brasses] 

= Notostomus patentissimus Spence Bate, 1888: 826; Plate 133; Plate 134, Figs 1-2. [Challenger stn 198, 

2°55'N 124°53'E, south of the Philippine Islands, 2150 fms] 

= Notostomus longirostris Spence Bate, 1888: 833; Plate 135; fig. 4. [Challenger stn 195, 4°21'S 129°7'E, 

off Banda Island, 1425 fms] 

= Notostomus westergreni Faxon, 1893: 208. [Albatross stn 3399 (North Pacific Ocean, Ecuador, 1°07'N, 

81°04'W), 1740 fms] 

= Notostomus atlanticus Lenz & Strunck, 1914: 330. [20°41'N 31°53'W, 3000 m, Westich von den Kapverden 

und Kanarischen Inseln] 

Notostomus gibbosus A. Milne-Edwards, 1881b: 7. [Grenade, 626 brasses] 

= Notostomus perlatus Spence Bate, 1888: 831; Plate 135, fig. 2. [Challenger stn 198, 2°55'N 124°53'E, 

near the Philippine Islands, 2150 fms]


= Notostomus brevirostris Spence Bate, 1888: 832; Plate 135, fig. 3. [Challenger stn 120, 8°37'S 34°28'W, 

off Pernambuco, 675 fms] 

Notostomus japonicus Spence Bate, 1888: 830; Plate 135, fig. 1. [Challenger stn 235, 34°7'N 138°0'E, south of 

Japan, 565 fms] 

Notostomus murrayi Spence Bate, 1888: 829; Plate 134, fig. 3. [Challenger stn 133, 35°41'S 20°55'W, near 

Tristan da Cunha, 1900 fms] 

Notostomus robustus Smith, 1884: 377; Plate 7, fig. 2. [Albatross stns 2042, 39°33'N 68°26'45"W, 1555 fms; 

2074, 41°43'N 65°21'50"W, 1309 fms] 

= Notostomus beebei Boone, 1930a: 39; Fig. 6. [five miles south of Nonsuch Island, Bermuda, 32°16'N 

64°36'W, 900 fms] 

Notostomus sparsidenticulatus Wasmer, 1986: 44; Fig. 9. [44°40'S 145°26'W, 2050-0 m] 

Family oPloPHoRIDAe Dana, 1852a 

Janicella Chace, 1986 

= Janicella Chace, 1986 (type species Oplophorus spinicauda A. Milne-Edwards, 1883, by original designation 


Janicella spinicauda (A. Milne-Edwards, 1883) 

= Oplophorus spinicauda A. Milne-Edwards, 1883: Plate 30. [off Casablanca, Morocco, 34°13'30"N 

7°43'W, 636 m] 

= Oplophorus foliaceus Rathbun, 1906: 922; Fig. 72; Plate 20, fig. 8. [Kaiwi Channel, 337-442 fms] 

= Acanthephyra anomala Boone, 1927: 104; Fig. 21. [north of Glover Reef, 484 fms] 

Oplophorus H. Milne edwards, 1837 [in H. Milne edwards, 1834-1840] 

= Oplophorus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species O.[plophorus] 

typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy, gender masculine) 

Oplophorus gracilirostris A. Milne-Edwards, 1881b: 6. [la Dominique, 118 brasses] (Fig. 14) 

= Oplophorus longirostris Spence Bate, 1888: 765; Plate 127, fig. 2. [Challenger stn 174c, 19°07'50"S 

178°19'35"E, off Kandavu, Fiji Islands, 610 fms] 

Fig. 14. Oplophorus gracilirostris A. Milne-Edwards, 1881. Photo by Tin-Yam Chan.


= Hoplophorus Smithii Wood-Mason & Alcock, 1891b: 194 [Investigator stns 62 (16°44'30"N, 88°19-32'E), 

1439 fms; 103 (15°15'N, 81°09'E), 1260 fms] 

= Oplophorus okitsuensis Yokoya, 1922: 302. [Okitsu, Suruga Bay] 

Oplophorus novaezeelandiae (De Man, 1931) 

= Hoplophorus (Oplophorus) novæ-zeelandiæ De Man, 1931: 369; Figs 1-20. [off Kaikoura, on the east 

coast of the South Island of New Zealand, 2 fms] 

Oplophorus spinosus (Brullé, 1839) 

= Palæmon spinosus Brullé, 1839: 18; vignette on page 3 (as Palémon épineux). [îles Canaries] 

= Hoplophorus Grimaldii Coutière, 1905a: 1; Fig. 1. [full description in Coutière, 1905a; 32°18'N 

23°58'W, 0-2000 m] 

Oplophorus typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= O.[plophorus] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 424. [Nouvelle- 

Guinée] 

= Oplophorus brevirostris Spence Bate, 1888: 766; Plate 127, fig. 3. [Challenger stn 207, 12°21'N 122°15'E, 

off Tablas Island, 700 fms] 

Systellaspis Spence Bate, 1888 

= Systellaspis Spence Bate, 1888 (type species Systellaspis lanceocaudata Spence Bate, 1888, by original 


= Hoplopasiphaea Yokoya & Shibata, 1965 (type species Hoplopasiphaea philippinensis Yokoya & Shibata, 


Systellaspis braueri braueri (Balss, 1914a) 

= Acanthephyra Braueri Balss, 1914a: 594. [0°26'N 6°32'W, 0-4000 m; 0°56'N 4°34'W, 0-4000 m] 

? = Systellaspis echinurus Coutière, 1911a: 158. [au large des côtes d’Espagne et du Portugal entre les 

parallèles 36 et 45, de part et d'autre du 11 e degré W] 

= Systellaspis densispina Stephensen, 1923: 57; Fig. 17. [46°30'N 7°00'W, > 4000 m] 

Systellaspis braueri paucispinosa Crosnier, 1987c: 953; Fig. 3a-b. [31°59'08"N, 158°04'04"E, entre 1685 m de 

profondeur et la surface] 

Systellaspis cristata (Faxon, 1893) 

= Acanthephyra cristata Faxon, 1893: 206. [Albatross stns 3361 (North Pacific Ocean, Panama, 

06°10'00''N 083°06'00''W), 1471 fms; 3381 (North Pacific Ocean, Colombia, 4°56'00"N, 80°52'30"W), 

1772 fms] 

Systellaspis curvispina Crosnier, 1987b: 711; Figs 6a-b, 8a-b. [Madagascar, 13°22'S, 47°17'38"E, 2000-0 m] 

Systellaspis debilis (A. Milne-Edwards, 1881b) 

= Acanthephyra debilis A. Milne-Edwards, 1881b: 13. [trouvée à une profondeur de 500 brasses dans 

le canal de Bahama] 

= Miersia gracilis Smith, 1882: 70, Plate 1, figs 4-4d. [Blake stn 328, 34°28'25"N 75°22'50"W, 1632 fms] 

= Acanthephyra debilis var. Europæa A. Milne-Edwards, 1883: Plate 33, fig. 2. [Type locality not indicated] 

= Systellaspis Bouvieri Coutière, 1905a: 8; Fig. 3. [Princesse-Alice stn 1856, 36°46'N 26°41'W, 0-3250 m] 

= Systellaspis debilis var. indica De Man, 1916: 151. [1°10.5'S 130°09'E, Halmaheira Sea, 798 m] 

= Hoplopasiphaea philippinensis Yokoya & Shibata, 1965: 4; Figs 4, 5. [13°17'N 125°50'E, 100 m] 

Systellaspis eltanini Wasmer, 1986: 52; Figs 13-14. [South Atlantic, 34°12'S 16°35'E, 1550-0 m] 

Systellaspis guillei Crosnier, 1987b: 718; Fig. 11. [Madagascar, 13°04'S 47°56'E, 1500-0 m] 

Systellaspis intermedia Crosnier, 1987c: 948; Figs 1b, 2, 3d. [Ouest-Pacifique, au nord des îles Marquises, 

0°-140°E, chalutage pélagique] 

Systellaspis lanceocaudata Spence Bate, 1888: 758; Plate 124, fig. 7. [Challenger stn 232, 35°11'N 139°28'E, off 

Japan, 345 fms] 

Systellaspis pellucida (Filhol, 1884) 

= Acanthephyra pellucida Filhol, 1884: 199. [26°20'N 14°53'W, 782 m, lectotype designation by Crosnier 

& Forest, 1973] 

= Acanthephyra affinis Faxon, 1896: 162; Plate 2, figs 1-3. [Blake stn 258 (Off Granada, 12°03'15"N, 

61°46'25"W), 159 fms]


Superfamily ATyoIDeA De Haan, 1849 [in De Haan, 1833-1850] 

Family ATyIDAe De Haan, 1849 [in De Haan, 1833-1850] 

Antecaridina edmondson, 1954 

= Mesocaris Edmondson, 1935a (type species Mesocaris lauensis Edmondson, 1935a, by monotypy, 

gender feminine; invalid junior homonym of Mesocaris Ortmann, 1893 (Crustacea Larvata)) 

= Antecaridina Edmondson, 1954 (nomen novum for Mesocaris Edmondson, 1935a, gender feminine) 

Antecaridina lauensis (Edmondson, 1935a) 

= Mesocaris lauensis Edmondson, 1935a: 13; Fig. 4. [cave containing brackish water, Island of Namuka, 

Lau islands, Fiji] 

Archaeatya Villalobos F., 1960a 

= Archaeatya Villalobos F., 1960a (type species Archaeatya chacei Villalobos F., 1960a, by original designation 


Archaeatya chacei Villalobos F., 1960a: 332; Figs 1-25. [Isla de Cocos, 87°30'W 5°30'N] 

Atya leach, 1816a 

= Atys Leach, 1816b (type species Atys scaber Leach, 1816b, by monotypy, gender masculine; name 

placed on the Official List of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 

1957) 

= Atya Leach, 1816a (type species Atys scaber Leach, 1816b, gender feminine; name placed on the 

Official List of Generic Names in Zoology in Opinion 470 in 1957) 

= Evatya Smith, 1871 (type species Evatya crassa Smith, 1871, by monotypy, gender feminine) 

Atya abelei Felgenhauer & Martin, 1983: 333; Figs 1-4. [Panama, El Valle, Cocle Province, unnamed tributary 

of Rio Anton, 600 m elevation] 

Atya africana Bouvier, 1904a: 138. [Samkitta, dans la rivière Ogooué] 

Atya brachyrhinus H.H.Jr. Hobbs & C.W.J. Hart, 1982: 35; Figs 1c, 12g, 15. [Cole’s Cave, Barbados, West 

Indies, 13°10'40"N 59°34'3"W] 

Atya crassa (Smith, 1871) 

= Evatya crassa Smith, 1871: 95. [Fresh water streams, Polvon, and the ”Rio Fulva, two and a half 

miles northwest of Realejo”; both localities in the Occidental Department of Nicaragua] 

Atya dressleri Abele, 1975a: 51; Figs 1-2. [Panamá, Pacific drainage, Veraguas Province, Río Santa María 

drainage, headwaters of Río San Juan, about 15 km above Calobre, 566 m elevation] 

Atya gabonensis Giebel, 1875: 52. [Gabon] 

= Euatya sculptilis Koelbel, 1884: 317; Plate 2, fig. 8; Plate 3, figs 1-8. [Orinoco] 

= Atya sculptata Ortmann, 1890: 465. [Afrika, vielleicht aus West Afrika] 

Atya innocous (Herbst, 1792) 

= Cancer (Astacus) Innocous Herbst, 1792 [in Herbst, 1791-1796]: 62; Plate 28, fig. 3. [das vaderland is 

unbekannt; considered to be ”Oceano Americano ad Martinicam”, see H.H.Jr. Hobbs & C.W.J. Hart, 

1982] 

= Astacus 988 Gronovius, 1764: 231; Plate 17, fig. 6. [in Oceano Americano ad Martinicam; included 

in a work rejected for nomenclatorial purposes in Opinion 261 in 1955] 

= Astacus Nasoscopus Meuschen, 1778: 86. [nomen novum for Astacus 988 Gronovius, 1764; included 

in a work rejected for nomenclatorial purposes in Opinion 260 in 1954] 

= A.[tya] occidentalis Newport, 1847: 159. [West India Islands] 

= Atya robusta A. Milne-Edwards, 1864: 148; Plate 3, fig. 1. [Nouvelle-Calédonie] 

= Atya tenella Smith, 1871: 94, 95. [fresh water streams, Polvon, Occidental Department, Nicaragua] 

Atya intermedia Bouvier, 1904a: 137. [Afrique occidentale, île Saint-Thomas] 

Atya lanipes Holthuis, 1963a: 61; Figs 1-2. [St. Thomas, Virgin Islands] 

Atya limnetes Holthuis, 1986a: 438; Figs 1-3. [Rio Anchicaya, Departemento del Valle del Cauca] 

Atya margaritacea A. Milne-Edwards, 1864 

= Atya Margaritacea A. Milne-Edwards, 1864: 148; Plate 3, fig. 2-2c. [Nouvelle-Calédonie (evidently 

erroneous, most certainly originates from East American waters, see Holthuis, 1966)]


= Atya rivalis Smith, 1871: 94. [fresh water streams Polvon, Occidental Department, Nicaragua] 

Atya ortmannioides Villalobos F., 1956: 459; Figs 1-6. [Río de las Truchas, La Mira, 52.5 km S.SE de Arteaga, 

Michoacán, 2.5 km de la costa, en la vertiente del Pacifico] 

Atya scabra (Leach, 1816b) 

= Atys scaber Leach, 1816b: 345. [Misantla, Estado de Veracruz, Mexico, 19°56'N 96°50'W; neotype 

designation by H.H.Jr. Hobbs & C.W.J. Hart, 1982] 

= Atya mexicana Wiegmann, 1836: 145. [Misantla] 

? = A.[tya] sulcatipes Newport, 1847: 159; Plate 8, fig. 1. [in fresh water, San Nicolao, Cape Verd Islands] 

= Atya punctata Kingsley, 1878a: 91, 92. [Hayti] 

= Atya margaritacea var. claviger Aurivillius, 1898a: 14; Plate 3, Figs 5-8. [Kamerun, Etome, in Bächen] 

= Atya margaritaria clavipes Holthuis, 1966: 234. [lapsus for Atya margaritacea var. claviger Aurivillius, 

1898a] 

Atyaephyra de Brito Capello, 1867 

= Symethus Rafinesque, 1814 (type species Symethus fluviatilis Rafinesque, 1814, an invalid senior 

subjective synonym of Hippolyte Desmarestii Millet, 1831, gender masculine; name suppressed for 

the purposes of the Principle of Priority but not for those of the Principle of Homonymy and placed 


= Acilius Rafinesque, 1815 (nomen novum for Symethus Rafinesque, 1814, gender masculine; name 

suppressed for the purposes of the Principle of Priority for those of the Principle of Homonymy and 


1958 and Opinion 619 in 1961) 

= Symaethus Agassiz, 1846 [in Agassiz, 1842-1846] (invalid emendation of Symethus Rafinesque, 


= Atyaephyra de Brito Capello, 1867 (type species Atyaephyra Rosiana de Brito Capello, 1867, by 

monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

522 in 1958) 

Atyaephyra desmarestii (Millet, 1831) 

= Hippolyte Desmarestii Millet, 1831: 56; Plate 1, figs 1-1b. [les eaux de la Mayenne, de la Sarthe, du 

Loir, du Thouet et du Layon] 

= Symethus fluviatilis Rafinesque, 1814: 23. [dans les ruisseaux et les mares; name suppressed under 

the plenary powers for the purposes of the Principle of Priority but not for those of the Principle of 

Homonymy in Opinion 522 in 1958] 

= Atyaëphyra Desmaresti var. occidentalis Bouvier, 1913a: 72. [le Nord de l’Afrique depuis la Tunisie, 

tout le Sud de l’Europe depuis la Macédonie] 

= Atyaephyra rosiana de Brito Capello, 1867: 6; Plate 1, figs 1-1e. [São Barnabe River, Algarve, Portugal; 

neotype selection by Anastiadou, Kitsos & Koukouras, 2008] 

Atyaephyra orientalis Bouvier, 1913a 

= Atyaëphyra Desmaresti var. orientalis Bouvier, 1913a: 72. [Syrie] 

= Atyaephyra desmarestii mesopotamica Al-Adhub, 1987: 1; Fig. 1. [Shatt Al-Arab river] 

Atyaephyra stankoi Karaman, 1972 

= Atyaephyra desmarestii stankoi Karaman, 1972: 82; Figs 3, 6, 9-10. [Mazedonien, Vardar Flussystem] 

Atydina Cai, 2010a 

= Atydina Cai, 2010a (type species Caridina atyoides Nobili, 1900a, by original designation, gender 

feminine) 

Atydina atyoides (Nobili, 1900a) 

= Caridina atyoides Nobili, 1900a: 478. [Si Oban; Sioban, Pulau Sipura, Kepulauan Mentawei, off west 

coast of Sumatra, Indonesia, according to Cai, 2010a] 

Atyella Calman, 1906a 

= Atyella Calman, 1906a (type species Atyella brevirostris Calman, 1906a, by original designation, gen-


der feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Atyella brevirostris Calman, 1906a: 201; Plate 14, figs 57-64. [Mbete; near mouth of Lofu; Kala; rock-pool 

at Kasakalawe (all Lake Tanganyika)] 

Atyella longirostris Calman, 1906a: 202; Plate 14, figs 65-72. [Mbete; Kala (both Lake Tanganyika)] 

Atyoida Randall, 1840 

= Atyoida Randall, 1840 (type species A.[tyoida] bisulcata Randall, 1840, by monotypy, gender feminine) 

= Ortmannia Rathbun, 1901 (type species Ortmannia henshawi Rathbun, 1901, a junior subjective 

synonym of A.[tyoida] bisulcata Randall, 1840, gender feminine) 

= Pseudatya Roux, 1928a (type species Pseudatya beauforti Roux, 1928a, a junior subjective synonym 

of A.[tya] pilipes Newport, 1847, by monotypy, gender feminine) 

= Vanderbiltia Boone, 1935 (type species Vanderbiltia rosamondae Boone, 1935, a junior subjective synonym 

of A.[tya] pilipes Newport, 1847, by original designation and monotypy, gender feminine) 

Atyoida bisulcata Randall, 1840 

= A.[tyoida] bisulcata Randall, 1840: 140; Plate 5, fig. 5. [Sandwich Islands] 

= Ortmannia henshawi Rathbun, 1901: 120. [Kaiwiki, Hilo, Hawaii, 1800ft altitude, 3 miles from the sea] 

Atyoida pilipes (Newport, 1847) 

= A.[tya] pilipes Newport, 1847: 160. [Apia, Upoln, New Zealand] 

= Atyoida tahitensis Stimpson, 1860a: 28. [in aquis dulcibus insulae ’Tahiti’] 

= Caridina acuminata Stimpson, 1860a: 29. [stream of Naka-kaigan coast, 60-70 m above sealevel, 

Chichi-jiam Island, Ogasawar Islands; neotype designation by Cai et al., 2006] 

= Caridina brevirostris Stimpson, 1860a: 29. [River Aritsu, Kume-jima Island; neotype designation by 

Cai et al., 2006] 

= Atya brevirostris De Man, 1892a: 360; Plate 21, figs 21-21d. [Fluss Wukur bei Sikka; Fluss Lella bei 

Sikka; Fluss bei Mbawa unterhalbs des Wasserfalls (all Flores); Fluss Koinino bei Kupang (Timor)] 

= Pseudatya beauforti Roux, 1928a: 209; Figs 1-9. [Batjan] 

= Vanderbiltia rosamondae Boone, 1935: 160; Plates 41-42. [Venus Point Reef, Tahiti, Society Islands, in 

coral] 

Fig. 15. Atyopsis moluccensis (De Haan, 1849). Photo by Tin-Yam Chan.


= Vanderbiltia mirabilis Holthuis, 1953a: 114. [manuscript name published as a synonym] 

Atyoida serrata (Spence Bate, 1888) 

= Atya serrata Spence Bate, 1888: 699; Plate 119, fig. 2. [Valley of San Antonia, San Iago, Cape Verde 

Islands from a fresh-water stream; probably erroneous, see Chace, 1983a] 

= Atya serrata mutation Alluaudi Bouvier, 1904b: 448. [un torrent de la montagne d’Ambre, à Madagascar; 

l’île Bourbon; dans les ravines des montagnes de Salasie et d’Helbour; l’île Maurice] 

Atyopsis Chace, 1983a 

= Atyopsis Chace, 1983a (type species A.[tya] spinipes Newport, 1847, by original designation, gender 

feminine) 

Atyopsis moluccensis (De Haan, 1849 [in De Haan, 1833-1850]) (Fig. 15) 

= Atya moluccensis De Haan, 1849 [in De Haan, 1833-1850]: 186; Plate O. [Moluccas, Indonesia; lectotype 

designation by Holthuis, 1993b] 

= Atya armata A. Milne-Edwards, 1864: 149; Plate 3, figs 3-3a. [Batavia (p. 145, 152), not les îles Philippines 

(p. 149), see Chace, 1983a] 

= Atya gustavi Ortmann, 1890: 467; Plate 36, fig. 9a-c. [Sumatra, Indrapura-Fluss] 

= Atya lineolata De Man, 1892a: 357 (footnote). [manuscript name attributed to Kulh, cited as synonym 

of Atya moluccensis] 

Atyopsis spinipes (Newport, 1847) 

= A.[tya] spinipes Newport, 1847: 159. [Philippine Islands] 

= Atya dentirostris Thallwitz, 1891: 101. [Nord-Celebes; fully described in Thallwitz, 1892] 

? = Atya brevirostris var. De Mani Nobili, 1900a: 475, fig. 1. [Fiume Sereinu, Isole Mentawei] 

Australatya Chace, 1983a 

= Australatya Chace, 1983a (type species Atya striolata McCulloch & McNeill, 1923, by original designation 


Australatya striolata (McCulloch & McNeill, 1923) 

= Atya striolata McCulloch & McNeill, 1923: 55; Plate 9, figs 3-4. [Norton’s Basin, Nepean River, New 

South Wales] 

Caridella Calman, 1906a 

= Caridella Calman, 1906a (type species Caridella cunningtoni Calman, 1906a, by original designation, 

gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

Caridella cunningtoni Calman, 1906a: 199; Plate 13, figs 45-52. [Kala, on rocks, shallow water; Kirando, 8 

fms; Utinta, 15 fms (all Lake Tanganyika)] 

Caridella minuta Calman, 1906a: 200; Plate 13, figs 53-56. [near mouth of Lofu, on rocks, shallow water; 

Karema, tow-netting, surface (both Lake Tanganyika)] 

Caridella paski Calman, 1928: 739; Figs 1-3. [Kigoma, east side of Lake Tanganyika] 

Caridina H. Milne edwards, 1837 [in H. Milne edwards, 1834-1840] 

= Caridina H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] (type species Caridina typus H. 

Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by monotypy and by indication under 

Article 68c of the ICZN, gender feminine; name placed on the Official List of Generic Names in 


Caridina ablepsia Guo, Jiang & Zhang, 1992: 4; Figs 1-9. [cave river, Wangcun Village, 109°56'E 28°46'N, 

Yongshun County, Hunan Province] 

Caridina acuta Liang, Chen & W.-X. Li, 2005: 530; Figs 15-29. [caves of Libo County (25°41'N 107°83'E), 

Maolan Nature Sanctuary of Guizhou] 

Caridina acutirostris Schenkel, 1902: 496; Plate 8, figs 3a-c. [Süden des Posso-See] 

Caridina africana Kingsley, 1883: 127; Plate 1, figs 3-3a. [Zulu Mission, South Africa] 

= Caridina africana forme typica Bouvier, 1925: 213; Fig. 470. [pays des Zoulous]


= Caridina wyckii var. paucipara Weber, 1897: 168. [River Umhlasine, Natal; lectotype designation by 

Richard & Clark, 2010a] 

Caridina alba J. Li & S. Li, 2010: 19; Figs 2-5. [Tenglongdong cave, Lichuan city, Enshi Autonomous Prefecture, 

Hubei province, China] 

Caridina alphonsi Bouvier, 1919: 330. [localité inconnue; austro-malaise according to Bouvier, 1925] 

Caridina amnicolizambezi Richard & Clark, 2009: 45; Figs 22-23. [Angola, Rio Lucoge, branch of the Chicapu 

River, 73°36'S 20°31'E] 

Caridina amoyensis Liang & Yan, 1977: 219; Figs 1-4. [Xiamen, suburabm rivulet, Fujian Province] 

Caridina angulata Bouvier, 1905c: 84; Fig. 8. [rivière Ranofotsy, près de Fianarantsoa] 

Caridina angustipes Guo & Liang, 2003: 45; Figs 1A-C. [Wentang Village, Xinhua County, Hunan Province, 

400 m] 

Caridina anislaq Cai, Choy & Ng, 2009: 82; Fig. 11. [Spring Anislaq, outside Cave Valencia, Valencia] 

Caridina annandalei Kemp, 1918a: 96; Plate 25, figs 6-15. [Inlé Lake] 

Caridina apodosis Cai & N.K. Ng, 1999: 1627; Figs 14-17. [mountain stream near Taitong village, New 

Territory, Hong Kong] 

Caridina appendiculata Jalihal & Shenoy, 1998: 128. [River Bari, Flores; lectotype designation by Cai & Ng, 

2007] 

Caridina aruensis Roux, 1911: 82. [ruisseau Matora, Soungi Manoumbai, Arou; ruisseau Panoua Bori, 

Soungi Manoumbai, Arou] 

Caridina babaulti babaulti Bouvier, 1918 

= Caridina Babaulti Bouvier, 1918: 388; Figs 4-6. [Mukhi; Majghaon; Rajadhar] 

Caridina babaulti basrensis Al-Adhub & Hamzah, 1987: 225; Fig. 1. [small channels connected to Shatt al- 

Arab River, Iraq] 

Caridina babaultioides babaultioides Yü, 1938 

= Caridina babaultioides Yü, 1938: 301; Figs 13-14. [small mountain stream, at an altitude of about 910 

m, at Shui-tuan-pa near Yen-tsing city, Yunnan Province] 

Caridina babaultioides angustifolia Cai & Yuan, 1996 

= Caridina angustifolia Cai & Yuan, 1996: 385; Figs 14-15. [Wafang stream, near Jinsha village] 

Caridina babaultioides emeica Liang, 2004: 218; Fig. 105. [Emeishan, Sichuan] 

Caridina babaultioides phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province] 

= Caridina phyllopoda Huang, 1984: 1; Figs 1-8. [Kwan Shien, Sichuan Province] 

Caridina bakoensis Ng, 1995a: 192; Figs 8-10. [Middle stretch of Sungai Serait] 

Caridina bamaensis Liang & Yan, 1983a: 252; Figs 1-10. [Bama County, Guangxi, Southern China] 

Caridina baojingensis Guo, He & Bai, 1992: 609; Figs 1-9. [China, Hunan Province, Baojing County, Qingshui 

village] 

Caridina batuan Cai, Choy & Ng, 2009: 80; Fig. 10. [Cave Castigio, Batuan] 

Caridina belazoniensis Richard & Clark, 2009: 33; Figs 14-15. [Kenya, Mbelazoni, Lower Ahti River, Sabaki 

Estuary] 

Caridina boholensis Cai, Choy & Ng, 2009: 75; Figs 7-8. [Quila Cave, Nueva Vida Norte, Batuan] 

Caridina brachydactyla De Man, 1908a 

= Caridina nilotica var. brachydactyla De Man, 1908a: 269; Plate 20, figs 8a-c. [Indonesia, by Reo, Flores; 

lectotype designation by Richard & Clark, 2010a] 

= Caridina nilotica var. natalensis De Man, 1908a: 262; Plate 20, figs 3-3b [South Africa, Umgeni River, 

Durban, Natal; lectotype designation by Richard & Clark, 2010a] 

= Caridina nilotica var. brevidactyla Roux, 1920: 320. [Aru-Inseln: Bach Matora, am Sungi Manumbai; 

Bach Panua Bori am Sungi Manumbai; Bach bei Papakula, Kobroor; Wokamar, Wokam; Samang; 

Seltutti, Kobroor; Dobo, Wammer; Pobdietur, Terangan; Udjir] 

Caridina breviata N.K. Ng & Cai, 2000: 171; Figs 4-6. [Zhapu village, Yangjiang County, Guangdong Province, 

southern China] 

Caridina brevicarpalis De Man, 1892a: 397; Plate 24, figs 30-30d. [Celebes, aus einem Flusse bei Palopo, 

Luwu] 

Caridina brevispina Liang & Yan, 1986a: 203; Fig. 5. [Tongren County, Guizhou Province]


Caridina bruneiana Choy, 1992: 49; Figs 1-4. [Negara Brunei Darussalam, on the upper reaches of Temburong 

River at Batang Duri, 04°36'05''N 115°06'45''E, altitude approx. 33 m] 

Caridina buehleri Roux, 1934a 

= Caridina bühleri Roux, 1934a: 219; Figs 1-5. [Bimoun, côte occidentale Nouvelle-Irlande] 

Caridina buergersi Karge, von Rintelen & Klotz, 2010: 146; Figs 5-6. [Papua New Guinea, former 

”Deutsch-Neuguinea”, near Mäanderberg] 

Caridina buhi Cai & Shokita, 2006a: 253; Figs 3-4. [Binahugan River, Buhi, Camarines Sur, Luzon, Philippines] 

Caridina bunyonyiensis Richard & Clark, 2005: 715; Figs 5-6. [Uganda, Lake Bunyonyi] 

Caridina burmensis Cai & Ng, 2000: 941; Fig. 7. [Myitkyina, Myanmar] 

Caridina caerulea von Rintelen & Cai, 2009: 428; Figs 51-53. [Lake Poso, west shore, Cape Bancea, 

01°59.023'S 120°35.108'E, on rocks in shallow water] 

Caridina calmani Bouvier, 1919 

= Caridina Calmani Bouvier, 1919: 334. [Ambatoubavara, Madagascar] 

= Caridina Bouvieri Roux, 1929: 312; Figs 7-12. [Manjakatompo, altit. 1940 m, ruisseau descendant de 

l’Ankaratra] 

Caridina camaro Cai, Choy & Ng, 2009: 82; Fig. 12. [Cave Camaro, Batuan] 

Caridina cantonensis Yü, 1938: 290; Figs 7-8. [Qing’ao vilage, Nan’ao County, Guangdong Province, China; 

neotype designation by Cai & N.K. Ng, 1999] 

= Caridina mutata Cai & N. K. Ng, 1999: 1624; Figs 12-13. [mountain stream near Fangcheng Town, 

Fangcheng County, Guangxi Province, China] 

Caridina caobangensis S.-Q. Li & Liang, 2002: 711; Figs 3-4. [PacBo Village, District Ha Quang, Cao Bang 

province, Vietnam] 

Caridina carli Roux, 1931a: 38; Figs 4-11. [Anaimalais, environs de Valparai, dans la rivière Naduar et ses 

affluents] 

Caridina cavaleriei cavaleriei Bouvier, 1919 

= Caridina Cavalerii Bouvier, 1919: 332. [Gan-chouen-fou (Kouy-Tchéou)] 

Caridina cavaleriei industana Roux, 1931a: 35; Figs 1-3. [Aliyar Riv., près de Malayandi Pattanam, 6 milles 

au Sud de Pollachi] 

Caridina cavalerieioides Liu & Liang in Liang, 2004: 207; Fig. 100. [Malin Village, near Guiyang City, 

Guizhou Province, China] 

Caridina caverna Liang, Chen & W.-X. Li, 2005: 529; Figs 1-14. [caves of Libo County (25°41'N 107°83'E), 

Maolan Nature Sanctuary of Guizhou] 

Caridina cavernicola Liang & Zhou, 1993: 232; Figs 2(1)-2(2). [Lenggu Cave, Duan County, Guangxi Province] 

Caridina cebuensis Cai & Shokita, 2006a: 250; Figs 1-2. [spring water to Sagay River, Cebu Island, Philippines] 

Caridina celebensis De Man, 1892a 

= Caridina serratirostris var. celebensis De Man, 1892a: 385; Plate 23; figs 28f-h. [Celebes, Fluss bei 

Palopo, Luwu] 

= Caridina serratirostris koterai Kamita, 1951: 75; Figs A-J. [Shimoko, Iwami province, SE Hunshu, 

Japan] 

Caridina celestinoi Blanco, 1939a: 392; Plate 3, Figs 8-10. [mountain stream, Helosig, Leyte] 

Caridina chauhani Chopra & Tiwari, 1949 

= Caridina nilotica var. chauhani Chopra & Tiwari, 1949: 219; Figs 2-3 [tanks at Sale bhata, Chandanbhati, 

Bolangir, Salepali and Titilagarh; Ang River at Salebhata] 

= Caridina williamsoni Jalihal, Shenoy & Sankolli, 1984: 1; Figs 1-3. [Sadhankeri tank, Dharwar] 

Caridina chishuiensis Cai & Yuan, 1996: 379; Figs 8-10. [stream near Binan village] 

= Caridina euryphylla Cai & Yuan, 1996: 382; Figs 11-13. [Qintan village, near Dongping village] 

Caridina clavipes Guo & Liang, 2003: 47; Figs 1D-F. [Wentang Village, Xinhua County, Hunan Province] 

Caridina clinata Cai, X.Q. Nguyên & Ng, 1999: 531; Figs 1-2. [ditch at Cuc Phuong National Park, Ninh 

Binh Province, Northern Vietnam]


Caridina cognata De Man, 1915a: 397; Plate 28, figs 3-3g, 4-4b [in einem kleinen Bache von süssem Wasser 

in der Küsten-gegend zwischen der Humboldt-Bai und dem unteren Laufe des Tami-Flusses; in 

Bächlein zwischen den kleinen Flüssen Tjahé and Jasa; aus dem Tjahé-Flusse; aus dem kleinen 

Tjano-Flusse bei Njao; Zoutbron] 

Caridina confusa Choy & Marshall, 1997: 27; Figs 1a, 2i-j, 3b-c, 4a-e. [Thiaki Creek at Seamark Road crossing, 

17°23.5'S 145°32.5'E] 

Caridina congoensis Richard & Clark, 2009: 48; Figs 24-25. [Congo, Bakou, in river, in total darkness, 30 

m] 

Caridina cornuta Liang & Yan, 1986a: 200; Fig. 3. [Zhijin County, Guizhou Province] 

Caridina crassipes Liang, 1993: 23; Fig. 2. [Xianshui Cave, Pangshi village, Hunan] 

Caridina crurispinata Gurney, 1984: 574; Figs 4-7. [Dark zone, la Grotte d’Antsatrobonko, Ankarana Massif, 

40 km north of Ambilobé, 60 km south of Diego Suarez, northern province of Diego Suarez] 

Caridina cucphuongensis Đăng, 1980 

= Caridina serrata cucphuongensis Đăng, 1980: 404; Fig. 230. [streams in Cucphuong area, Ninh Binh 

Province, Northern Vietnam] 

Caridina curta Liang & Cai, 2000: 179; Fig. 2. [Longquan spring in Huanglong (Yellow Dragon) Temple, 

Jianshui County, Yunnan] 

Caridina demani Roux, 1911: 94. [Tawarin] 

Caridina demenica Cai & Li, 1997: 315; Fig. 1. [Demen Cave, near Yongkang village, Libo County of 

Guizhou Province, southern China] 

Caridina dennerli von Rintelen & Cai, 2009: 355; Figs 4-6. [Lake Matano, east shore, just at entrance to 

outlet bay, 02°31.54'S 121°27.00'E] 

Caridina dentifrons N.K. Ng & Cai, 2000: 167; Figs 1-3. [Baijin village, Huishui County, Guizhou Province, 

southern China] 

Caridina devaneyi Choy, 1991: 348; Figs 2-4. [stream at 8-mile Point near Suva, Viti Levu, Fiji Islands] 

Caridina dianchiensis Liang & Yan, 1985a: 196; Fig. 1. [Songhuaba Reservoir- Panlong River, Kunming, 

Jiangchuan County, Yunnan province] 

Caridina disjuncta Cai & Liang, 1999: 77; Fig. 4. [Shilin (Stone forest) village, Lunan County, central Yunnan] 

Caridina disparidentata Liang, Yan & Wang, 1984: 8; Figs 1-12. [mountain stream, Qujing County, Yunnan 

Province] 

= Caridina heterodentata Liang & Yan, 1985a: 196. [unjustified emendation of Caridina disparidentata 

Liang, Yan & Wang, 1984; see Cai & Ng, 2001a] 

Caridina ebuneus Richard & Clark, 2009: 39; Figs 18-19. [Ivory Coast, vicinity Abengourou] 

Caridina edulis Bouvier, 1904a: 135. [Anantsahalankely] 

Caridina elisabethae Karge, von Rintelen & Klotz, 2010: 142; Figs 3-4. [Papua New Guinea, Morobe, Herzog 

Mts., Bundun, 700-800 m, 06°51.598'S 146°37.07'E] 

Caridina elliptica Cai & Yuan, 1996: 376; Figs 5-7. [stream near Panlong town] 

Caridina elongapoda Liang & Yan, 1977 

= Caridina nilotica elongapoda Liang & Yan, 1977: 220; Figs 5-8. [Xingzai, Zhangpu County, Fujian 

Province] 

Caridina endehensis De Man, 1892a 

= Caridina brevicarpalis var. endehensis De Man, 1892a: 399; Plate 24, fig. 30e. [aus dem Flusse Ba bei 

Endeh; Fluss bei Mbawa (both Flores)] 

Caridina ensifera Schenkel, 1902: 490; Plate 8, figs 1-11e. [Lake Poso, Celebes (Sulawesi), Indonesia; lectotype 

designation by Cai & Wowor, 2007] 

Caridina evae Richard & Clark, 2009: 28; Figs 11-13. [Nigeria, Chubra division] 

Caridina excavata Kemp, 1913b: 306; Plate 20, figs 32-35; Plate 21, figs 36-37. [backwater of the Rowta 

River, Brahmaputra drainage system] 

Caridina excavatoides Johnson, 1961: 127; Figs 3-11. [stream running between rubber plantations and ricefields, 

about nine miles from Alor Star, Kedah, on the Pokok Sena Road] 

Caridina fasciata Hung, Chan & Yu, 1993: 489; Figs 1E, 6. [Ping Tong County, Heng Chun]


Caridina fecunda Roux, 1911: 95. [Lac Jamour] 

Caridina feixiana Cai & Liang, 1999: 74; Figs 1-2. [stream in Feixia cave, Geju County, Yunnan, China] 

Caridina fernandoi Arudpragasam & Costa, 1962: 8; Fig. 1. [shallow streamlets at Warakapola, and Seelangama, 

and also from shallow sheltered areas of the Maha Oya at Mawanella and the Deduru Oya 

and the Magura Oya at Kurunegala, Sri Lanka] 

Caridina fijiana Choy, 1983: 147; Fig. 1. [Fiji, Viti Levu, Nadarivatu, Nukunuku Creek, 17°35'40''S 

177°57'25''E, 640 m] 

Caridina flavilineata Đăng, 1975: 70; Fig. 5. [Nam Ha Province] 

Caridina formosae Hung, Chan & Yu, 1993: 487; Figs 1D, 5. [Keelung, Pa Chih Men] 

Caridina fossarum Heller, 1862b 

= C.[aridina] fossarum Heller, 1862b: 411. [Schiraz] 

? = Caridina syriaca Bouvier, 1904a: 132. [Syrie] 

Caridina gabonensis Roux, 1927a: 239; Figs 1-7. [Lambaréné, Gabon] 

Caridina ghanensis Richard & Clark, 2009: 35; Figs 16-17. [Republic of Ghana, Pond Vume] 

Caridina glaubrechti von Rintelen & Cai, 2009: 359; Figs 7-9. [Lake Towuti, west shore, south of Cape 

Timbalo, 02°42.91'S 121°26.78'E] 

Caridina glossopoda Liang, Guo & Gao, 1993: 41; Fig. 1. [Ala, Fenghuang County, Hunan, 27°95'N 

109°65'E] 

Caridina gordonae Richard & Clark, 2005: 717; Figs 7-8. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina gortio Cai & Anker, 2004: 247; Figs 8, 9a-f. [Can Gortio Cave, 11°59'00''N 124°53'00''E, altitude ca 

200 m, Matalud, West Samar, Philippines] 

Caridina gracilipes De Man, 1892a 

= Caridina Wyckii var. gracilipes De Man, 1892a: 387; Plate 24, figs 29e-k [aus dem Flusse von Maros; 

Makassar; aus einem kleinen mit dem Meere nicht in Verbindung stehenden Bache zu Balangnipa; 

Palima, aus dem Tjenrana; Pampanua, aus dem Tjenrana (all Celebes); Saleyer, aus dem Flusse 

Bonéa] 

= Caridina nilotica var. bengalensis De Man, 1908a: 265; Plate 20, figs 6-6b. [Port Canning an Dhappa, 

Calcutta] 

Caridina gracilirostris De Man, 1892a: 399; Plate 25, fig. 31-31d. [river near Maros, Sulawesi (Celebes), 

Indonesia; lectotype designation by Cai & Ng, 2007] 

= Caridina pseudogracilirostris Thomas, V.K. Pillai & N.N. Pillai, 1976: 871; Fig. 1. [Cochin backwater] 

Caridina gracillima Lanchester, 1901: 560; Plate 34, fig. 1. [Malay Peninsula; lectotype designation by Cai 

& Ng, 2007] 

Caridina grandirostris Stimpson, 1860a: 28 [upper stream of Tima River, about 4 km from river mouth, 

Okinawa Island, Ruykyu Islands; neotype designation by Cai et al., 2006] 

Caridina guangxiensis Liang & Zhou, 1993: 234; Figs 3(1)-3(2). [Dawangshan cave, Guilin City, Guangxi 

Province] 

Caridina gueryi Marquet, Keith & Kalfatak, 2009: 160; Figs 1-3. [unnamed river, Santo Island (Vanuatu), 

15.29590 S 167.16 E] 

Caridina guiyangensis Liang, 2002a: 119; Fig. 2. [China, Guizhou Province, Guiyang County] 

Caridina gurneyi Jalihal, Shenoy & Sankolli, 1984: 29; Figs 12-14. [Malaprabha river, near old bridge, 

Khanapur (Belgaum District)] 

Caridina hainanensis Liang & Yan, 1983b: 211; Fig. 1. [Wencang County, Hainan Island, China] 

Caridina hanshanensis Tan, 1990: 278; Figs 1-9. [Hanshan County, 31°45'N 118°7'E, Anhui Province] 

Caridina harmandi Bouvier, 1906f 

= Caridina Harmandi Bouvier, 1906f: 483. [Japon] 

Caridina hodgarti Kemp, 1913b: 309; Plate 20, Figs 29-31; Plate 21, figs 38-39. [Kobo, Abor county] 

Caridina holthuisi von Rintelen & Cai, 2009: 364; Figs 10-12. [Lake Matano, south shore, canal between 

island and mainland, 02°28.46'S 121°15.83'E] 

Caridina hongyanensis Cai & Yuan, 1996: 373; Figs 2-4. [Yantanggou stream near Hongyan village] 

Caridina hova Nobili, 1905c: 499. [Fort Dauphin] 

Caridina huananensis Liang, 2004: 147; Fig. 69. [Guangdong]


Caridina hubeiensis Liang & S.-Q. Li, 1993: 213; Figs 1-9. [Badong, Hubei Province] 

Caridina hunanensis Liang, Guo & Gao, 1993: 44; Fig. 3. [Zhonglian, Lengshuijiang City, Hunan, 27°7'N 

111°4'E] 

Caridina imitatrix Holthuis, 1970: 100; Fig. 3. [Rivière Bleue near the bridge near Ouénarou (Mont des 

Sources); Marsh of Rivière Blanche, Ouénarou region; Side branch of Rivière Blanche, near the forest 

road leading from Ouénarou forestry station to the forestry log cabin on the western slope of Mt. 

Pouédihi, Ouénarou region; same branch, near the log cabin; Thir River near the confluence with a 

side branch, granite area near St. Louis, S.E. of Nouméa; Rivière des Lacs, which empties in the Lac 

en Huit, near highway to Nouméa] 

Caridina indistincta indistincta Calman, 1926 

= Caridina indistincta Calman, 1926: 244; Fig. 3. [St. George District, Queensland] 

Caridina indistincta sobrina Riek, 1953: 119; Fig. 9. [Rocky Creek, Fraser Island, Queensland] 

Caridina isaloensis isaloensis Coutière, 1899b 

= Caridina typus var. Isaloensis Coutière, 1899b: 383. [Fleuve Onilahy, côte Ouest] 

= Caridina madagascariensis Bouvier, 1904a: 134. [rivière Fandramanona] 

= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar] 

Caridina isaloensis grandidieri Bouvier, 1904a 

= Caridina Grandidieri Bouvier, 1904a: 133. [rivière Fandiamanana, Madagascar] 

Caridina jalihali Mariappan & Richard, 2006: 47; Figs 22-24. [River Arani at Periapalayam; lake at Thenneri; 

ponds at Maduranmangalam, Manimangalam, Redipalayam, RoshanNagar and Singaperumal 

Koli] 

= Caridina gurneyi lonavalensis Kadrekar & Sankolli, 1987: 60. [nomen nudum] 

Caridina jeani Cai, 2010b: 80; Figs 1-2; nomen novum for Caridina typus var. brevirostris Roux, 1911. [Elat, 

Kei Islands, Moluccas, Indonesia; lectotype designation by Cai, 2010b] 

Caridina jiangxiensis Liang & Zheng, 1985 

= Caridina pingi jiangxiensis Liang & Zheng, 1985: 319; Fig. 1(1b-7b). [Shixi stream, Fengxin County, 

Jiangxi Province] 

Caridina johnsoni Cai, Ng & Choy, 2007: 301; Figs 15-16. [Singapore, Lower Peirce Reservoir, north arm] 

Caridina kaombeflutilis Richard & Clark, 2010a: 331; Figs 14-15. [Malawi, Kaombe River] 

Caridina kempi Jalihal, Shenoy & Sankolli, 1984: 13; Figs 6-8. [Hosayellapur tank (Hirekeri), Dharwar] 

Caridina kilimae Hilgendorf, 1898: 35. [Marangu am Kilimandscharo] 

Caridina kunmingensis Z.-Z. Wang & Liang, 2001: 303; Fig. 1. [Kunming, Yunnan Province] 

Caridina kunnathurensis Richard & Chandran, 1994: 250; Figs 4, 5A. [Kunnathur pond, Madras, India] 

Caridina laevis Heller, 1862b 

= C.[aridina] laevis Heller, 1862b: 411. [Java] 

Caridina lamiana Holthuis, 1965a: 39; Fig. 14. [Ambodivoangy near Maroantsetra, Antongil Bay, NE 

Madagascar; Forest between 20 and 30 km NW of Ambahoabé, Simianona River, N of Soanierana, 

east coast of Madagascar] 

= Caridina brevirostris var. brevipes Bouvier, 1925: 230 (partim). [baie Amposindova, Madagascar] 

Caridina lanceifrons Yu, 1936: 89; Figs 4-7. [near the light house at Hai-kiu-sche in the salt water] 

Caridina lanceolata Woltereck, 1937: 224; Fig. 1.7a-c; Table 1. [Lake Mantano, Mengonuwai, Sulawesi; 

neotype designated by Cai, Wowor & Choy, 2009] 

Caridina lanzana Holthuis, 1980c: 2; Figs 1-3. [small well named Bog Der, 08°35'54''N 48°46'30''E, Nogal 

Valley, northern Somalia] 

Caridina laoagensis Blanco, 1939a: 390; Plate 2. [?Laoag River, Laoag, Ilocos Norte Province, Luzon] 

Caridina leclerci Cai & Ng, 2009: 1108; Fig. 9. [Pange, cave at Tallasa, Maros, Sulawesi Selatan] 

Caridina leucosticta Stimpson, 1860a: 28. [138°56.43'E 34°41.83'E, Inouzawa River, Shimoda City, Izu Peninsula; 

neotype designation by Cai et al., 2006] 

Caridina leytensis Blanco, 1939a: 391; Plate 3, Figs 1-7. [Helosig, Leyte] 

Caridina liangi Jiang, Guo & Zhang, 2002: 220; Fig. 1. [small stream at an elevation of 450 m near Yongping 

Village, Yuanling County] 

Caridina liaoi Cai, Choy & Ng, 2009: 72; Figs 5-6. [Bilar River, Bilar] 

Caridina lima Liang, Guo & Gao, 1993: 43; Fig. 2. [Buermen, Yongshun County, Hunan, 29°N 109°8'E]


Caridina linduensis Roux, 1904: 541; Plate 9, figs 1-4. [Lac Lindu, région centrale de Célèbes (part. Occid.), 

altitude 980 m] 

Caridina lineorostris Richard & Clark, 2009: 51; Figs 26-27. [Gabon, Keri, forest stream, 20 km North of 

Lambarene] 

Caridina lingkonae Woltereck, 1937 

= Caridina Lingkonae Woltereck, 1937: 218; Fig. 1.1; Table 1. [Lake Towuti, west shore, at entrance to 

outlet bay, Cape Larona, 02°48.43'S 121°24.75'E; neotype designated by von Rintelen & Cai, 2009] 

Caridina lipalmaria Richard & Clark, 2010b: 639; Figs 1-2. [Bestiboka River Basin, Beharena River, Antaniditra, 

48°02'55''E 18° 25'43''S, 13730 asl] 

Caridina liui Liang & Yan, 1986a: 202; Fig. 4. [Tongzi County, Guizhou Province] 

Caridina lobocensis Cai, Choy & Ng, 2009: 68; Figs 2-4. [tributuary of Loboc River, Loboc] 

Caridina loehae Woltereck, 1937 

= Caridina Loéhae Woltereck, 1937: 222; Fig. 1.5a-d; Table 1. [Lake Towuti, about 3 km south of Timampu, 

estuary of Sungei Batuopa, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina longa Liang & Yan, 1985a: 200; Fig. 3. [spring water, Huanglongsi, Jianshui, Yunnan Province] 

Caridina longiacuta Guo & Wang, 2005: 14; Fig. 1. [near Yuanliping village, Yizhang County, Hunan Province 

(ca. 25°25'N 112°57'E)] 

Caridina longicarpus Roux, 1926a 

= Caridina weberi var. longicarpus Roux, 1926a: 212; Figs 37-39. [Nouvelle-Calédonie, au-dessus 

d’Oubatche, pentes du Mt. Ignambi, env. 600 m d’altitude] 

Caridina longidigita Cai & Wowor, 2007: 317; Figs 4-5. [west coast of Lake Poso at Taipa area, Kab. Poso, 

Sulawesi Tengah] 

Caridina longifrons Cai & Ng, 2007: 1599; Fig. 6. [Kabupaten Maros, stream above Bantimurung waterfall, 

Sulawesi, Indonesia] 

Caridina longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= C.[aridina] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363. [la rivière 

de la Macta, près d’Oran [probably erroneous]] 

= Caridina nilotica var. meridionalis Roux, 1926a: 207. [Pemboa; Haute Tiouaka; Koné; Coula-Boréaré; 

La Foa; Coindé, Ruisseua Katiramona, entre Nouméa et Païta (all Nouvelle-Calédonie)] 

= Caridina acuticaudata Đăng, 1975: 70; Fig. 4. [Boi River, Hoa Binh Province] 

Caridina lovoensis Roth-Woltereck, 1955: 197; Figs 1-2. [Lovo B., 20 km von Thysville (Bas-Congo), 

Kalkhöhlen] 

Caridina lufengensis Cai & Duan, 1998: 330; Figs 1-3. [pond near Sizhi Town, Lufeng County, Yunnan, 

China, 25.07°N, 102.10°E, elevation 1800 m] 

Caridina lumilympha Richard & Clark, 2010a: 328; Figs 12-13. [Kenya, Lumi River] 

Caridina macrodentata Cai & Shokita, 2006b: 2156; Figs 13-14. [123°51.84'E 24°23.65'N, fast flowing water, 

about 200 m from sea, Omija River, Iriomote Island, Ryukyu Islands] 

Caridina macrophora Kemp, 1918b 

= Caridina nilotica var. macrophora Kemp, 1918b: 277; Fig. 9. [Tale Sap, Peninsular Siam] 

= Caridina subnilotica Đăng, 1975: 69; Fig. 3. [ponds in Hanoi] 

Caridina maculata L. Wang, Liang & F. Li, 2008: 49; Figs 1-2. [Yingde County, Guangdong Province, 

N23°54.5' E113°13.6'] 

Caridina mahalona Cai, Wowor & Choy, 2009: 29; Figs 8-9. [Lake Mahalona, at southwest coast] 

Caridina malayensis Cai, Ng & Choy, 2007: 305; Fig. 17. [stream at Nee Soon Swamp, Singapore] 

Caridina malawensis Richard & Clark, 2009: 69; Figs 38-39. [Malawi, rocky shore of Lake Malawi] 

Caridina masapi Woltereck, 1937 

= Caridina Masapi Woltereck, 1937: 223; Fig. 1.6a-h; Table 1. [southern shore of Lake Masapi, South 

Sulawesi, 2°50.84'S 121°21.09'E; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina mathiassi Silas & Jayachandran, 2010: 1; Figs 1-5, 6J, 7. [Mahendragiri Estate, part of southern 

Western Ghats, Kanyakumari District, Tamil Nadu, India, 8° 27'32.2''N 77°23'32.5''E] 

Caridina mauritii Bouvier, 1912a 

= Caridina Mauritii Bouvier, 1912a: 298. [Maurice] 

Caridina mccullochi Roux, 1926b: 249. [Pallal, Horton River, near Bingara]


Caridina medifolia Cai & Yuan, 1996: 388; Figs 16-18. [Xiaojin village near Daqun town] 

Caridina mengae Liang, 1993: 22; Fig. 1. [Xianshui Cave, Panshi village, Songtao County (28.19°N 

109.20°E), Guizhou Province] 

Caridina mengaeoides Guo & Suzuki, 1996: 97; Figs 1-2. [Yuanling County, Hunan Province, China, ca. 

28°18'N 110°20'E, altitude ca. 750 m] 

Caridina menghaiensis Cai & Dai, 1999: 217; Figs 5-6. [reservoir in Menghai County] 

Caridina meridionalis L. Wang, Liang & F. Li, 2008: 56; Figs 7-8. [Heyuan County, Guangdong Province] 

Caridina mertoni Roux, 1911: 84. [Elat, île de Grand-Kei; entre Elat et Ohinangan, île de Grand-Kei; Enraalan, 

île de Grand-Kei; Warka, île de Grand-Kei] 

Caridina mesofluminis Richard & Clark, 2009: 66; Figs 36-37. [Cameroon, Mess stream] 

Caridina mindanao Cai & Shokita, 2006a: 259; Figs 7-9. [Lake Mainit, Mindanao, Philippines] 

Caridina minidentata Cai & Anker, 2004: 250; Figs 10-11. [Can Gortio Cave, 11°59'00''N 124°53'00''E, altitude 

ca 200 m, Matalud, West Samar] 

Caridina minnanica Liang, 2002a: 121; Fig. 3. [China, Fujian Province, Yunxiao County] 

Caridina modiglianii Nobili, 1900a 

= Caridina Modiglianii Nobili, 1900a: 477. [Kifa-juc] 

Caridina moeri Roth-Woltereck, 1984: 102; Figs 1 (1a-b, 2, 3 (1a-e). [Mwerusee] 

Caridina mongziensis Liang, Yan & Z.-Z. Wang, 1987: 133; Figs 1-9. [Mongzi County, 23°20'N 103°25'E, 

Yunnan Province] 

Caridina multidentata Stimpson, 1860a: 29. [stream in Tenno-ura, Chichi-jima Island, Ogasawara Islands; 

neotype designation by Cai et al., 2006] 

= Caridina japonica De Man, 1892b: 261; Plate 9, Figs 7-7a. [Kagar Hayagana, Japan] 

= Caridina Voeltzkowi Lenz, 1910a: 569. [Alaotra-See (Madagaskar)] 

= Caridina japonica sikokuensis Kubo, 1938a: 91; Fig. 20. [Ryûgadô, Kôti Prefecture] 

Caridina nanaoensis Cai & N. K. Ng, 1999: 1617; Figs 6a, f, 8. [Xiaoliao stream near Xishan Town, Nan’ao 

County, Guangdong Province, China] 

Caridina natalensis Bouvier, 1925 

= Caridina africana forme natalensis Bouvier, 1925: 214. [South Africa, Umholti River, Verulum, Natal 

(according to Richard & Clark, 2009)] 

Caridina natarajani Tiwari & R.S. Pillai, 1968: 163; Fig. 1. [Trivandrum, Kerala] 

Caridina neglecta Cai & Ng, 2007: 1595; Figs 4-5. [Sungai Batang, 13 km on road from Palopo to Wotu, 

Sulawesi, Indonesia] 

Caridina nguyeni S.-Q. Li & Liang, 2002: 709; Figs 1-2. [PacBo Village, District Ha Quang, Cao Bang 

Province,Vietnam] 

Caridina nilotica (Roux, 1833) 

= Pelias Niloticus Roux, 1833: 73; Plate 7, fig. 1. [les eaux du Nil] 

= Caridina nilotica var. typica Bouvier, 1925: 146; Fig. 308. [Caire] 

Caridina norvestica Holthuis, 1965a: 35; Fig. 12. [River north of Majunga; near Majunga, small brook at 

km 530 of the highway to Tananarive; Mahajambe lake, Madagascar] 

Caridina novaecaledoniae Roux, 1926a 

= Caridina novae-caledoniae Roux, 1926a: 214; Figs 40-46. [Tchalabel; Vallée du Diahot, chemin du Col 

Poraris, 150 m; versant Est du Col Poraris, 100 m; Pemboa; Haute Tiouaka (all Nouvelle-Calédonie)] 

Caridina nudirostris Choy, 1984: 288; Fig. 1. [above Vuwa Falls, Wainisavulevu Creek, Nadrau Plateau, 

Central Viti Levu, Fiji, 17°50'30''S 178°01'30''E, altitude 800 m; below Vuwa Falls, Wainisavulevu 

Creek, 17°48'10''S 178°05'30''E, altitude 160 m; above Monasavu Falls, Nanuku Creek, Nadrau Plateau, 

Central Viti Levu, Fiji; 17°44'30''S 178°02'30''E, altitude 700 m] 

Caridina okiamnis Richard & Clark, 2009: 54; Figs 28-29. [Cameroon, Okia stream] 

Caridina okinawa Cai & Shokita, 2006b: 2154; Fig. 12. [Ingue Cave at Yomitan Village, Okinawa Island, 

Ryukyu Islands, Japan] 

Caridina oligospina Liang, Guo & Tang, 1999: 72; Fig. 3. [China, Hunan Province, Fengfang County, Muli 

village] 

Caridina opaensis Roux, 1904: 547; Plate 9, figs 8-10. [Lac Opa. Bras S.-E. de l’île, region centrale, alt. 30 m]


Caridina palawanensis Cai & Shokita, 2006a: 256; Figs 5, 6A-B. [Panitian River, upper stream, ca. 30 km 

from river mouth, Palawan Island, Philippines] 

Caridina panikkari Jalihal, Shenoy & Sankolli, 1984: 9; Figs 4-5. [Hosayellapur tank (Hirekeri), Dharwar] 

Caridina papuana Nobili, 1905d 

= Caridina Weberi var. papuana Nobili, 1905d: 481; Plate 12, fig. 1-1b. [Nova Guinea Tedesca] 

Caridina paracornuta Cai & Yuan, 1996: 391; Figs 19-21. [Chuangfengao stream, Hongyan village] 

Caridina pareparensis De Man, 1892a: 379; Plate 22, Figs 25-25b. [river near Pare Pare, Sulawesi, Indonesia; 

lectotype designated by Cai & Ng, 2009] 

Caridina parvidentata Roux, 1904 

= Caridina pareparensis var. parvidentata Roux, 1904: 545; Plate 9, figs 5-7. [Malawa Quelle, Celebes (= 

Sulawesi); lectotype designation by Cai & Ng, 2009] 

Caridina parvirostris De Man, 1892a: 375; Plate 22, fig. 24. [Flores, Fluss bei Bombang] 

Caridina parvocula Gurney, 1984: 569; Figs 1-3. [Dark zone, la Grotte d’Antsatrobonko, Ankarana Massif, 

40 km north of Ambilobé and 60 km south of Diego Suarez, northern province of Diego Suares] 

Caridina parvula von Rintelen & Cai, 2009: 392; Figs 27-29. [Lake Towuti, southwest shore, Cape Sioloya, 

02°50.7'S 121°26.32'E] 

Caridina paucidentata L.-Q. Wang & Liang, 2005: 748; Figs 1-15. [Mongzi County, Yunnan Province] 

Caridina pedicultrata Guo & Choy, 1994: 123; Figs 1-2. [stream near Huaihua City, at Xuefeng Mountain, 

Hunan, c. 27°40'N 110°11'E] 

Caridina peninsularis Kemp, 1918b 

= Caridina brachydactyla peninsularis Kemp, 1918b: 279; Fig. 10. [Botanical Garden, Penang, Malaysia; 

lectotype designation by Cai, Ng & Choy, 2007] 

Caridina petiti Roux, 1929 

= Caridina Petiti Roux, 1929: 108; Figs 13-18. [Ambila, lagunes orientales] 

Caridina pingi Yü, 1938: 294; Figs 9-10. [Amoy, Fukien] 

Caridina pingioides Yü, 1938: 298; Figs 11-12. [locality being unkown to the writer] 

Caridina plicata Liang, 2004: 282; Fig. 138. [Xita, Jiangxi] 

Caridina prashadi Tiwari & R.S. Pillai, 1971: 87; Figs 3-4. [Aberdeen (Port Blair), South Andaman, from a 

freshwater stream] 

= Caridina sakishimensis Fujino & Shokita, 1975: 99; Fig. 5. [Yonaguni I.: Arakawabana River; Spring 

of Tendabanata; Tabaru River; Ishigaki I.: Miyara River; Yonehara River; Sakutara River; Fukido 

River; Miyako I.: Izaga Cave; Bora Rover; Kume I.: Shirase River] 

Caridina pristis Roux, 1931b: 63; Figs 1-3. [alentours de Perdenyia; prés de Kandy, Ceylan] 

= Caridina pristis cruszi de Silva, 1982: 131; Fig. 3. [stream in Sinharaja forest] 

Caridina profundicola von Rintelen & Cai, 2009: 396; Figs 30-32. [Lake Towuti, Loeha Island, southwest 

shore, 02°45.58'S 121°31.14'E, on boulders in deeper water] 

Caridina propinqua De Man, 1908b: 227; Plate 19, Figs 6-6f. [Dhappa, near Calcutta] 

= Caridina blancoi Chace, 1997: 6; Fig. 2. [Philippines, near mouth of Tyabas River, Luzon, 13°54'N 

121°36'E] 

Caridina pseudodenticulata Hung, Chan & Yu, 1993: 498; Figs 9D, 13. [Yun Lin County, Ku Keng] 

Caridina pseudonilotica Richard & Clark, 2005: 722; Figs 9-10. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina pseudoserrata Đăng & Ðỗ, 2007: 1; Figs 1-2. [Bang River, Cao Bang Province] 

Caridina qingyuanensis Guo & He, 2007: 47; Figs 1-19. [Baishikeng Village, Qingyuan City, Guangdong- 

Province (ca. 23°35'N 113°10'E)] 

Caridina rajadhari Bouvier, 1918 

= Caridina Rajadhari Bouvier, 1918: 386; Figs 1-3. [Rajadhar, dans l’état de Kawarda, massif montagneux 

situé dans les provinces centrales, entre Jubbulpoor et Nagpoor; Majghaon, non loin de Rajadhar; 

Mukhi, même région] 

Caridina rangoona Cai & Ng, 2000: 939; Fig. 6. [17°09.97'N 96°99.20'E, Win Paw Hta River, near border 

between Pegu (Bago) and Yangon, Yangon State, Myanmar] 

Caridina rapaensis Edmondson, 1935b: 12; Fig. 5a-h. [freshwater stream, Rapa] 

Caridina richtersi Thallwitz, 1892


= Caridina serrata Richters, 1880: 163; Plate 17, Figs 24-27; nec Stimpson, 1860a. [Botan. Garten Pamplemousses; 

Black river] 

= Caridina richtersii Thallwitz, 1892: 27. [nomen novum for Caridina serrata Richters, 1880] 

= Caridina apiocheles Bouvier, 1904a: 134. [peut-être des Seychelles] 

= Caridina apiocheles mutation Edwardsi Bouvier, 1904a: 134. [peut-être des Seychelles] 

= Caridina Richtersi forma typica Bouvier, 1925: 203. [Type locality not indicated] 

Caridina roubaudi Bouvier, 1925 

= Caridina africana forme Roubaudi Bouvier, 1925: 217; Fig. 477. [Congo, Brazzaville; lectotype designated 

by Richard & Clark, 2009] 

Caridina rouxi De Man, 1915a: 387; Plate 27, figs 1-11. [in einem kleinen, schnell fliessenden Flusse im 

Bougainville-Gebirge, 500 m] 

Caridina rubella Fujino & Shokita, 1975: 102; Fig. 6. [Miyako I.: Izaga Cave, Morikaga Cave; Nikadori, 

Hirara City, in well] 

Caridina rubropunctata Đăng & Ðỗ, 2007: 3; Figs 3-4. [streams in Van Lang, Dong Hy District, Thai Nguyen 

Province] 

Caridina samar Cai & Anker, 2004: 240; Fig. 4-5. [Sulpan Cave, 12°02'00''N 124°55'30''E, altitude 100 m, 

Matalud, West Samar, Philippines] 

Caridina sarasinorum Schenkel, 1902: 492; Plate 8, Figs 2a-e. [Lake Poso, central Sulawesi, Indonesia; lectotype 

designation by Cai & Wowor, 2007] 

Caridina schenkeli von Rintelen & Cai, 2009: 442; Figs 60-62. [small stream, west of Lake Poso, 02°2.613'S 

120°37.311'E, on macrophytes] 

Caridina semiblepsia Guo, Choy & Gui, 1996: 66; Figs 1-4. [Dongpaoshan Cave (c. 28°44'N 109°39'E), Hunan 

Province, China] 

Caridina serrata Stimpson, 1860a: 29. [hill above Bekhers, Hong Kong island, Hong Kong; neotype designation 

by Cai & N. K. Ng, 1999] 

Caridina serratirostris De Man, 1892a: 382; Plate 23, Figs 28-28e. [Bangkalanvir River, Saleyer; lectotype 

designation by Richard & Clark, 2010a] 

Caridina shenoyi Jalihal & Sankolli in Jalihal, Shenoy & Sankolli, 1984: 21; Figs 9-11. [Malaprabha river– 

near old bridge, Khanapur (Belgaum District), India] 

Caridina shilinica Liang & Cai, 2000: 177; Fig. 1. [Shilin (Stone forest) Tour Resort in Lunan County Yunnan 

Province, southern China] 

Caridina similis Bouvier, 1904a: 135. [Mahé, Seychelles; lectotype designation by Marquet & Keith, 2008] 

= Caridina brevirostris forma typica Bouvier, 1912b: 916. [Seychelles] 

= Caridina brevirostris forma Gardineri Bouvier, 1912b: 916. [Seychelles] 

Caridina simoni Bouvier, 1904a 

= Caridina Simoni Bouvier, 1904a: 131. [Kandee, l’île de Ceylan] 

= Caridina costai de Silva, 1982: 126; Fig. 2. [three streams in Sinharaja forest, Sri Lanka] 

Caridina sodenensis Richard & Clark, 2009: 42; Figs 20-21. [Cameroon, Lake Soden, West Cameroons] 

Caridina solearipes Guo & De Grave, 1997: 127; Figs 2-3. [small stream near Dabaozi, Jingzhou County, 

Hunan province, circa 26°41'N 09°25'E, altitude between 400-900 m] 

Caridina songtaoensis Liang, 2004: 270; Fig. 32. [Songtao, Guizhou] 

Caridina spathulirostris Richters, 1880: 163; Plate 17, fig. 28. [Botanischer Garten Pamplemousses] 

Caridina spelunca Choy, 1996: 103; Fig. 1. [Old Napier Downs Cave, Western Australia, Australia, 17°14'S 

124°39'E] 

Caridina sphyrapoda Liang & Zhou, 1993: 236; Figs 4(1)-4(2). [Longdong River in Laibing County, 

Guangxi, southern China] 

Caridina spinalifrons Guo & De Grave, 1997: 124; Fig. 1. [stream near Sangzhi city, Hunan Province, circa 

29°10'N 110°16'E, 650 m altitude] 

Caridina spinata Woltereck, 1937: 221; Fig. 1.3; Table 1. [Lake Towuti, estuary of Sungei Batuopa, about 2 

km south of Timampu, Sulawesi, Indonesia; neotype designated by Cai, Wowor & Choy, 2009] 

Caridina spinipoda Liang, Hong & Yang, 1990: 161; Figs 1-9. [Mianzhu County, 31°20'N 104°11'E, Sichuan 

Province]


Caridina spinosipes Liang, Guo & Tang, 1999: 71; Fig. 2. [China, Hunan Province, Fengfang County, Muli 

village] 

Caridina spinula Choy & Marshall, 1997: 31; Figs 1c, 2g-h, 4f-q. [east flowing first order tributary of 

Leo Creek, near crossing of Leo Creek Mine Road, 13°44.6'S 143°21.5'E, Nesbit River catchment, 

McIlwraith Range, Cape York] 

Caridina spongicola Zitzler & Cai, 2006: 271; Figs 1-3. [Indonesia, Sulawesi Selatan, Lake Towuti, west 

shore, outlet bay, west of Cape Tokaluku, 02°47.261'S 121°23.17'E] 

Caridina steineri Cai, 2005: 314; Figs 2-3. [Lakata Zafera, western Madagascar, 19°45.075'S 45°11.436'E, 

stream pool at far end of cave] 

Caridina striata von Rintelen & Cai, 2009: 410; Figs 39-41. [Indonesia, Sulawesi Selatan, Lake Towuti, 

north shore, 02°38.56'S 121°27.82'E, on rocks] 

Caridina subventralis Richard & Clark, 2005: 725; Figs 11-12. [Uganda, Bufundi, Lake Bunyonyi] 

Caridina sulawesi Cai & Ng, 2009: 1103; Figs 6-8. [Sungai Beru, Kappang, Maros, Sulawesi Selatan] 

Caridina sumatianica Cai & Yuan, 1996: 395; Figs 22-23. [Sumatian stream, near Hongyan village] 

Caridina sumatrensis De Man, 1892a 

= Caridina Weberi var. sumatrensis De Man, 1892a: 375; Plate 22, fig. 23g. [flüssen des unteren Battaklandes 

bei Deli, an der Ostküste von Sumatra] 

Caridina sundanella Holthuis, 1978a: 32; Figs 11-12. [Waikamburu Brook, 4 km N. of Waimangura, West 

Sumba, about 250 m above sea level] 

Caridina susuruflabra Richard & Clark, 2009: 60; Figs 32-33. [South Africa, Eoshowe, Zululand] 

Caridina temasek Choy & Ng, 1991: 266; Figs 2-5. [freshwater stream near Sime road, MacRitchie catchment 

area, Singapore, ca. 01°20'14''N 103°48'47''E] 

Caridina tenuirostris Woltereck, 1937: 224; Fig. 1.8; Table 1. [Lake Towuti, southwest shore, west of Cape 

Tetetu, 02°54.13'S 121°23.78'E; neotype designation by von Rintelen & Cai, 2009] 

= Caridina Towutensis Woltereck, 1937: 220; Fig. 1.2; Table 1. [Towuti-Süd] 

Caridina thambipillai Johnson, 1961: 138; Figs 25-35. [Sungei Putat near the pumping station, Malacca] 

Caridina thermophila Riek, 1953: 120; Fig. 10. [Muttaburra, western Queensland] 

Caridina thomasi von Rintelen, Karge & Klotz, 2008: 2247; Figs 2-3. [Indonesia, Central Sulawesi, Banggai 

islands, Peleng, west peninsula, east of Alani, river with lake-like extension, 01°28.315'S 122°52.473'E] 

Caridina timorensis De Man, 1893: 300; Plate 8, fig. 6. [freshwater lake of Nefko on the island of Timor] 

Caridina togoensis Hilgendorf, 1893a: 156. [Togo, Bismarksburg; lectotype designation by Richard & 

Clark, 2009] 

= Caridina togoënsis var. stuhlmanni Hilgendorf, 1898: 35. [Undussuma und Bach bei Undussuma] 

= Caridina togoensis var. Decorsei Bouvier, 1904a: 131. [Congo français, rivière prés de Mpoko; région 

Chari-Tchad, Krébédje (Fort-Sibut) et Binguétou, rivière Gribingui; Bangoran] 

= Caridina togoensis var. breviatus Lenz, 1910b: 131. [Urwaldbache N.W. von Beni] 

= Caridina africana forme ægyptiaca Bouvier, 1925: 214. [Caire] 

= Caridina togoensis var. Kasaiensis De Man, 1925: 5; Figs 2 n1 -2 n3 . [Kamaiembe; Kondué] 

= Caridina togoensis var. Kwamouthensis De Man, 1925: 9; Fig. 2 o1 -2 012 . [Kwamouth au confluent du 

fleuve Congo et de la rivière Kasaï] 

= Caridina togoensis var. Schoutedeni De Man, 1925: 20; Figs 2 q1 -2 q21 . [les rivières du bassin du Chiloango; 

Buto-Polo dans la rivière Mapanga] 

Caridina tonkinensis Bouvier, 1919: 331. [Tonkin] 

Caridina trifasciata Yam & Cai, 2003: 277; Figs 1, 3-5. [Tsak Yue Wu, New Territories, Hong Kong] 

Caridina troglodytes Holthuis, 1978b: 214; Figs 3-4. [Danmin Cave near Konogusgus, New Ireland, in 

subterranean stagnant pool, 0.3-0.6 m] 

Caridina troglophila Holthuis, 1965a: 37; Fig. 13. [Ambovonomby cave, Namoroka, NW Madagascar] 

Caridina tumida L. Wang, Liang & F. Li, 2008: 54; Figs 5-6. [Yingde County, Guangdong Province, 

N23°54.4' E113°15.1'] 

Caridina typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= C.[aridina] typus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 363; Plate 25bis, Figs 

4-5. [Type locality unknown, probably l’île Maurice according to Bouvier, 1925]


= Caridina exilirostris Stimpson, 1860a: 29. [Okuma River, Okinawa Island, Rykyu islands; neotype 

designation by Cai, Ng, Shokita & Satake, 2006] 

= Caridina siamensis Giebel, 1863: 329. [Siam] 

? = Caridina Spencebatei De Man, 1892a: 371. [?Cap Verdischen Inseln] 

= Caridina typus forme typica Bouvier, 1925: 250. [l’île Maurice] 

= Caridina typus forme caledonica Bouvier, 1925: 253; Figs 296-297. [Nouvelle-Calédonie] 

Caridina uminensis Đăng & Ðỗ, 2007: 6; Figs 5-6. [U Minh Thuong wetland, Ca Mau Province] 

Caridina umtatensis Richard & Clark, 2009: 63; Figs 34-35. [South Africa, Kraal Dam, Umtata] 

Caridina unca Gurney, 1984: 579; Figs 8-11. [Dark zone, Grotte d’Antsatrobonko, Andarana Massif, northern 

province of Diego Suarez] 

Caridina valencia Cai, Choy & Ng, 2009: 78; Fig. 9. [spring Anislaq, outside Valencia] 

Caridina venusta L. Wang, Liang & F. Li, 2008: 52; Figs 3-4. [Yingde County, Guagndon Province, 

N23°54.3' E113°15.8'] 

Caridina vietriensis Đăng & Ðỗ, 2007: 9; Figs 7-8. [confluence of rivers in Viet Tri City, Phu Tho Province] 

Caridina villadolidi Blanco, 1939a: 389; Plate 1. [Laoag River, Laoag, Ilocos Norte Province, Luzon] 

= Caridina typus var. longirostris De Man, 1892a: 370; Plate 22, fig. 22f-i; nec C.[aridina] longirostris H. 

Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]. [Flores, as einem Flusse bei Reo; auch auf 

Celebes und Saleyer] 

Caridina vitiensis Borradaile, 1899: 1003; Plate 63, figs 3-3a. [Suva, Tamavua River, Viti Levu, Fiji] 

= Caridina vitiensis var. canacorum Roux, 1926a: 199; Figs 32-36. [Nouvelle-Calédonie, Canala] 

Caridina weberi De Man, 1892a 

= Caridina Weberi De Man, 1892a: 371; Plate 22, figs 23-23g. [Kotting; Fluss bei Mbawa, oberhalb des 

Wasserfalls; Fluss bei Bombang (all Flores); Fluss bei Palopo, Luwu; Wasserfall bei Bantimurong 

unweit Maros; Fluss bei Pare-pare (all Celebes); Saleyer, Fluss Bangkalan] 

= Caridina weberi var. typica Bouvier, 1925: 243; Figs 562-566. [Flores, Kotting, environs de Mbawa, 

de Bombang; Java; Kifa-juc, Ile Engano] 

? = Caridina weberi var. keiensis Roux, 1911: 85. [Elat, île de Grand-Kei; ruisseau entre Elat et Ohinangan; 

Warka, île de Grand-Kei; ruisseau entre Elat et Ohilim, île de Grand-Kei] 

Caridina williamsi Cai & Ng, 2000: 933; Figs 4-5. [17°48.11'N 96°09.20'E, Balar stream, 12 km Yangon to 

Mandalay highway, Htauk Kyant, Yangon State, Myanmar] 

Caridina woltereckae Cai, Wowor & Choy, 2009: 19; Fig. 3. [Cape Larona, near Sungai Larona outlet, Lae 

Towuti, Sulawesi, Indonesia] 

Caridina wumingensis Cai & N.K. Ng, 1999: 1632; Figs 18-19. [cave near Wuming, Guangxi Province, 

China] 

Caridina wyckii (Hickson, 1888) 

= Atya Wyckii Hickson, 1888: 358; Plates 13-14. [Lake Tondano, Minahassa, North Celebes] 

= Caridina nilotica var. minahassae De Man, 1902: 895. [Minahassa, Celebes] 

Caridina xiangnanensis X.-Y. Liu, Guo & Yu, 2006: 44; Fig. 1. [near Lingxiu Village, Rucheng County, Hunan 

Province (ca. 25°33'N 113°40'E)] 

Caridina xiphias Bouvier, 1925 

= Caridina nilotica var. xiphias Bouvier, 1925: 149; Figs 310-312. [district d’Ambatondrazaka, Madagascar] 

= Caridina nilotica var. stylirostris Bouvier, 1925: 148; Fig. 309. [Madagascar] 

= Caridina gladiifera Roux, 1929: 306; Figs 1-6. [Périnet, forêt, Madagascar] 

Caridina yilong Cai & Liang, 1999: 76; Fig. 3. [Yilong Lake, Shiping County, Yunnan] 

Caridina yulinica Cai & N.K. Ng, 1999: 1620; Figs 9-11. [Niuwo cave near Kuiyang Town, Yulin County, 

Guangxi Province, China] 

Caridina yunnanensis Yü, 1938: 304; Figs 14-16. [Mountain stream at Hsün-tien, Yunnan Province] 

= Caridina impensa Cai & Ng, 2001a: 223; Figs 8-9. [Xilong Tang (Pond), Chengjiang County, Yunnan] 

Caridina zebra Short, 1993: 62; Figs 1-3. [O’Leary Creek, 17°50.7'S 145°37.7'E, altitude ca. 750 m] 

Caridina zeylanica Arudpragasam & Costa, 1962: 17; Fig. 4. [Nawala, suburb of Colombo] 

= Caridina nilotica var. veliensis R.S. Pillai, 1964: 44. [Veli Lake, Trivandrum]


Caridina zhejiangensis Liang & Zheng, 1985 

= Caridina pingi zhejiangensis Liang & Zheng, 1985: 321; Fig. 1(1c-7c). [Baisha Village, Jiande County, 

Zhejiang Province, China] 

Caridina zhongshanica Liang, 2004: 182; Fig. 88. [Zhongshan, Guangdong] 

Caridinides Calman, 1926 

= Caridinides Calman, 1926 (type species Caridinides wilkinsi Calman, 1926, by monotypy, gender 


Caridinides wilkinsi Calman, 1926: 242; Figs 1-2. [Olive River, Temple Bay, east coast of Cape York Peninsula] 

Caridinopsis Bouvier, 1912c 

= Caridinopsis Bouvier, 1912c (type species Caridinopsis Chevalieri Bouvier, 1912c, by monotypy, 


1957) 

Caridinopsis chevalieri Bouvier, 1912c 

= Caridinopsis Chevalieri Bouvier, 1912c: 300; Figs 1-4. [région du Haut Niger, à Sampouyara, bassin 

de la Makowa] 

= Caridinopsis brevinaris Holthuis, 1956a: 56. [Garrigues Cave near Sougouéta, French Guinea] 

Dugastella Bouvier, 1912d 

= Dugastella Bouvier, 1912d (type species Dugastella marocana Bouvier, 1912d, by monotypy, gender 

feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Dugastella marocana Bouvier, 1912d: 993. [la source de Settat] 

Dugastella valentina (Ferrer Galdiano, 1924) 

= Atyaephira valentina Ferrer Galdiano, 1924: 210; Figs 1, 3. [Laguna de Almenara (Castellón) y Albufera 

de Valencia] 

= Dugastella marocana var. hispanica Balss, 1925b: 206; Figs 1-4. [Silla, bei Valencia, Acequia communera, 

1.5 km v. Albufera] 

Edoneus Holthuis, 1978b 

= Edoneus Holthuis, 1978b (type species Edoneus atheatus Holthuis, 1978b, by original designation 


Edoneus atheatus Holthuis, 1978b: 220; Figs 5-6. [the Philippines, Luzon, Quirino province, Aglipay Municipality, 

Barrio Palasian, cave near Sitio Disiluad; see Balete & Holthuis, 1992] 

Edoneus erwini Cai & Husana, 2009: 54; Figs 2-3. [Bantakay Cave, station 2, Luzon, the Philippines] 

Edoneus marulas Cai & Husana, 2009: 60; Figs 6-7. [at middle of Marulas Cave, Luzon, the Philippines] 

Edoneus sketi Cai & Husana, 2009: 57; Figs 4-5. [small pool in Bantakay cave, Luzon, the Philippines] 

Gallocaris Sket & Zakšek, 2009 

= Gallocaris Sket & Zakšek, 2009 (type species Troglocaris Schmidti inermis Fage, 1937, by original 


Gallocaris inermis (Fage, 1937) 

= Troglocaris Schmidti inermis Fage, 1937: 215; Figs 1-6. [grotte de Cambous, près Saint-Hippolyte-du- 

Fort, département du Gard] 

Halocaridina Holthuis, 1963b 

= Halocaridina Holthuis, 1963b (type species Halocaridina rubra Holthuis, 1963b, by monotypy, gender 

feminine) 

Halocaridina palahemo Kensley & Williams, 1986: 429; Figs 9-11. [Hawaii Island, Ka Lae, Lua o Palahemo, 

lava tube pool, 18°55'N 155°42'W] 

Halocaridina rubra Holthuis, 1963b: 262; Fig. 1. [pool in the base of Lohena Rock, between the deserted 

villages of Wai-o-ahu-kini and Kaili-kii, W. of Ka Lae or South Point, Kau district, island of Hawaii]


Halocaridinides Fujino & Shokita, 1975 

= Halocaridinides Fujino & Shokita, 1975 (type species Halocaridina (Halocaridinides) trigonophthalma 

Fujino & Shokita, 1975, by monotypy, gender masculine) 

= Palauatya C.W.J. Hart, 1980 (type species Palauatya dasyomma C.W.J. Hart, 1980, by original designation 


Halocaridinides fowleri Gordon in Gordon & Monod, 1968 

= Parisia (?) fowleri Gordon in Gordon & Monod, 1968: 514; Figs 30-31. [subterranean lake, Kufile, 

Zanzibar] 

Halocaridinides trigonophthalma (Fujino & Shokita, 1975) 

= Halocaridina (Halocaridinides) trigonophthalma Fujino & Shokita, 1975: 106; Figs 7-8. [Okinawa Island: 

Kaneshi, Nakijin, in well, 10 m; Aja, Naha City, in well] 

= Palauatya dasyomma C.W.J. Hart, 1980: 481; Figs 1-31. [anchialine lake on Anguar Island, Palau, 

Caroline Islands, 6°54'49''N 134°08'12''E] 

Jolivetya Cals, 1986 

= Jolivetya Cals, 1986 (type species Jolivetya foresti Cals, 1986, by monotypy, gender feminine) 

Jolivetya foresti Cals, 1986: 387; Fig. 1. [Nouvelle-Bretagne, la grotte de Pogo Ngim, lieu d’accès au réseau 

souterrain d’Arais, dans un karst à dolines, à quelques dizaines de kilomètres du littoral de la Nouvelle-Bretagne] 

Jonga C.W.J. Hart, 1961 

= Jonga C.W.J. Hart, 1961 (type species Ortmannia Serrei Bouvier, 1909a, by original designation and 


Jonga serrei (Bouvier, 1909a) 

= Ortmannia Serrei Bouvier, 1909a: 332. [Cuba] 

Lancaris Cai & Bahir, 2005 

= Lancaris Cai & Bahir, 2005 (type species Caridina singhalensis Ortmann, 1894, by original designation, 


Lancaris kumariae (de Silva, 1990) 

= Caridina kumariae de Silva, 1990: 9; Fig. 1. [small stream near railway station at Rozella, Sri Lanka, 

6°56'-6°57'N 80°33'-80°35'E] 

Lancaris singhalensis (Ortmann, 1894) 

= Caridina singhalensis Ortmann, 1894: 11; Plate 1, fig. 2. [Ceylon, Newera Elya] 

Limnocaridella Bouvier, 1913b 

= Limnocaridella Bouvier, 1913b (type species Limnocaridina Alberti Lenz, 1910b, by monotypy, gender 

feminine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

Limnocaridella alberti (Lenz, 1910b) 

= Limnocaridina Alberti Lenz, 1910b: 132; Plate 3, Figs 6-9. [Albert-See] 

Limnocaridina Calman, 1899a 

= Limnocaridina Calman, 1899a (type species Limnocaridina tanganyikæ Calman, 1899a, by monotypy, 


1957) 

Limnocaridina iridinae Roth-Woltereck, 1958: 188; Figs 1-5. [Tanganjika-See bei Mpulungu (Nordrhodesien, 

Abercorn District), 3 m, aus dem Kiemenraum einer Iridina spekei] 

Limnocaridina latipes Calman, 1906a: 196; Plate 12, figs 23-29. [Mbete, shallow water amongst rocks; near 

mouth of Lofu, on rocks, shallow water; Kalambo, tow-netting; Tembwi, on rocks, shallow water 

(all Lake Tanganyika)] 

Limnocaridina parvula Calman, 1906a: 193; Plate 11, figs 9-14. [Kasawa, tow-netting; Kalambo, tow-netting; 

Karema tow-netting, surface (all lake Tangaynika)]


Limnocaridina retiarius Calman, 1906a: 192; Plate 11, figs 2-8. [Lake Tanganyika, Mbete, taken in shrimpnet, 

shore wading] 

Limnocaridina similis Calman, 1906a: 195; Plate 12, figs 15-22. [Lake Tanganyika, Kalambo, tow-netting; 

Rusisi River, close to Tanganyika] 

Limnocaridina socius Calman, 1906a: 196; Plate 12, figs 30-37. [Niamkolo Harbour, 3 fms; Utinta, 10 fms; 

Kirando, about 8 fms (all Lake Tangyanika)] 

Limnocaridina spinipes Calman, 1906a: 197; Plate 13, figs 38-44. [Niamkolo Harbour, 3 fms; Kirando, 8 

fms; Utinta, about 15 fms (all Lake Tanganyika)] 

Limnocaridina tanganyikae Calman, 1899a 

= Limnocaridina tanganyikæ Calman, 1899a: 704; Plate 39, figs 1-2, 4-9a; Plate 40, figs 10-19. [Lake 

Tanganyika, shallow water] 

Mancicaris liang, Guo & Tang, 1999 

= Mancicaris Liang, Guo & Tang, 1999 (type species Mancicaris sinensis Liang, Guo & Tang, 1999, by 

original designation and monotypy, gender feminine) 

Mancicaris sinensis Liang, Guo & Tang, 1999: 70; Fig. 1. [Tianxinpu village, Tianxin district, Lanshan 

County, Hunan Province, China] 

Marosina Cai & Ng, 2005 

= Marosina Cai & Ng, 2005 (type species Marosina brevirostris Cai & Ng, 2005, by original designation, 


Marosina brevirostris Cai & Ng, 2005: 131; Fig. 1. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi 

Selatan, Indonesia] 

Marosina longirostris Cai & Ng, 2005: 133; Figs 2-4. [Gua Salukkan, Kallang, Kappang, Maros, Sulawesi 

Selatan] 

Micratya Bouvier, 1913b 

= Calmania Bouvier, 1909b (type species Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 

1855-1856], by monotypy, gender feminine; an invalid junior homonym of Calmania Laurie, 1906 

(Crustacea Brachyura) and Calmania Nobili, 1907 (a junior subjective synonym of Brachycarpus 

Spence Bate, 1888); name placed on the Official Index of Rejected and Invalid Generic Names in 


= Micratya Bouvier, 1913b (type species Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 

1855-1856], by monotypy, gender feminine; name placed on the Official List of Generic Names in 


= Balssiola Strand, 1922 (nomen novum for Calmania Bouvier, 1909b, gender feminine; name placed 


Micratya poeyi (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) 

= Atya Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xviii; Plate 2, Figs 7, 7a-b. 

[Cuba] 

Neocaridina Kubo, 1938a 

= Neocaridina Kubo, 1938a (type species Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833- 

1850], by original designation, gender feminine) 

Neocaridina anhuiensis (Liang, Zhu & Xiong, 1984) 

= Caridina denticulata anhuiensis Liang, Zhu & Xiong, 1984: 251; Figs 1-7. [Taiping County, Anhui 

Province] 

Neocaridina bamana Liang, 2004: 112; Fig. 55. [Bama, Guangxi] 

Neocaridina brevidactyla Liang, Chen & W.-X. Li, 2005: 532; Figs 3-43. [caves of Libo County (25°41'N 

107°83'E), Maolan Nature Sanctuary of Guizhou] 

Neocaridina curvifrons (Liang, 1979) 

= Caridina curvifrons Liang, 1979: 118; Figs 1-10. [Baisha, Zhejiang Province]


Neocaridina denticulata davidi (Bouvier, 1904a) 

= Caridina Davidi Bouvier, 1904a: 133. [Inkiafou, Chensi méridonale] 

Neocaridina denticulata denticulata (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte denticulatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 8. [Japan] 

= Caridina denticulata sinensis Kemp, 1918b: 287; Fig. 11c-d. [creeks and irrigation channels at the 

edge of the Tai Hu lake in Kiangsu province] 

Neocaridina denticulata moganica Liang, 2004: 82; Fig. 41. [Mogashan, Zhejiang] 

Neocaridina euspinosa Cai, 1996: 150; Figs 15-16. [Yanshan mountain near Cuilin City (25°20'N 110°18'E), 

Guangxi Zhuang Autonomous Region, China] 

Neocaridina fukiensis (Liang & Yan, 1977) 

= Caridina spinosa fukiensis Liang & Yan, 1977: 222; Figs 9-11. [Gutian, Shnaghang County (rivulet); 

Xinquan, Liancheng County (pool); Luokou and Ansha, Yongan County (pool) (all Fujian Province)] 

Neocaridina gracilipoda Liang, 2004: 88; Fig. 44. [Junlian, Sichuan] 

Neocaridina heteropoda heteropoda Liang, 2002b 

= Neocaridina heteropoda Liang, 2002b: 167; Fig. 1a-h. [Baisha Village, Jiande County, Zhejiang Province, 

China] 

Neocaridina heteropoda koreana Kubo, 1938a 

= Neocaridina denticulata koreana Kubo, 1938a: 81; Figs 5c, 7g-m, 7g’-m’, 12. [Huzan, Korea] 

Neocaridina heteropoda luoyangensis Cai, 1996 

= Neocaridina denticulata luoyangensis Cai, 1996: 139, Figs 7-8. [from a mountain stream near Luoyan 

City (34°40'N 112°20'E), Henan Province, China] 

Neocaridina hofendopoda (Shen, 1948) 

= Caridina hofendopoda Shen, 1948: 122; Plate 13, Figs f-l. [Kweilin] 

Neocaridina homospina Liang, 2002b 

= Neocaridina euspinosa homospina Liang, 2002b: 169; Fig. 2. [China, Hunan Province, Shaodong County] 

Neocaridina iriomotensis Naruse, Shokita & Cai, 2006: 26; Figs 1, 2. [upper reaches of the Nishifunatsuki, 

Nakama River, Iriomote Island] 

Neocaridina ishigakiensis (Fujino & Shokita, 1975) 

= Caridina denticulata ishigakiensis Fujino & Shokita, 1975: 95; Figs 3-4. [Yonehara River; Miyara 

River; Sakutara River, Arakawa River (all Ishigaki Island)] 

Neocaridina ketagalan Shih & Cai, 2007: 687; Figs 5-6, 7D-E. [Sijhih, Taipei Co., Taiwan] 

Neocaridina keunbaei (H.S. Kim, 1976) 

= Caridina denticulata keunbaei H.S. Kim, 1976: 155; Figs 1-3. [upper stream of Cheonji fall, Jeju Island] 

Neocaridina linfenensis Cai, 1996 

= Neocaridina denticulata linfenensis Cai, 1996: 137; Figs 5-6. [near Linfen City, 36°5'N 111°30'E, Shanxi 

Province, China] 

Neocaridina longipoda (Cai, 1995a) 

= Caridina longipoda Cai, 1995a: 166; Figs 1-15. [Jinbianxi stream, Zhangjiajie National Forest Park, 

Hunan Province] 

Neocaridina palmata bosensis Cai, 1996: 145; Figs 11-12. [Longlin County (25°15'N 105°22'E), in Guangxi 

Zuang Autonomous Region, China] 

Neocaridina palmata luodianica Liang, 2004: 110; Fig. 54. [Luodian, Guizhou] 

Neocaridina palmata meridionalis Liang, 2004: 108; Fig. 53. [Luoding, Guangdong] 

Neocaridina palmata palmata (Shen, 1948) 

= Caridina palmata Shen, 1948: 120; Plate 12. [Sha-ping-pa Chungking] 

= Caridina elongata Shen, 1948: 121; Plate 13, figs a-e. [Kweilin] 

= Caridina denticulata vietnamensis Đăng, 1967: 157; Fig. 4. [Cao Bang province; Lang Son province] 

Neocaridina saccam Shih & Cai, 2007: 682; Figs 2-4, 7A-B. [mountain stream at Longci, Tainan Co., 

Taiwan] 

Neocaridina spinosa (Liang, 1964) 

= Caridina spinosa Liang, 1964: 187; Figs 1-12. [mountain stream, Lantang, Tzechin Hsien, Kwangtung]


Neocaridina xiapuensis Zheng, 2002: 19; Fig. 1. [Xiapu County, 26°55'N 120°E, Fujian Province, China] 

Neocaridina zhangjiajiensis Cai, 1996: 147; Figs 13-14. [Zhangjiajie National Forest Park in Dayong City, 

Hunan Province of China] 

Neocaridina zhoushanensis Cai, 1996 

= Neocaridina denticulata zhoushanensis Cai, 1996: 142; Figs 9-10. [Dinhai County in Zhoushan Island 

(30°02'N 122°08'E), Zhejiang Province, China] 

Palaemonias Hay, 1902 

= Palæmonias Hay, 1902 (type species Palæmonias ganteri Hay, 1902, by monotypy, gender masculine; 


Palaemonias alabamae Smalley, 1961: 127; Fig. 1. [Shelta Cave, Huntsville, Madison County, Alabama] 

Palaemonias ganteri Hay, 1902 

= Palæmonias ganteri Hay, 1902: 179. [Roaring River passage, Mammoth Cave] 

Paracaridina liang, Guo & Tang, 1999 

= Paracaridina Liang, Guo & Tang, 1999 (type species Caridina longispina Guo, He, Xu & Gui, 1992, 

by original designation and monotypy, gender feminine) 

Paracaridina chenxiensis Guo & De Grave, 2004: 203; Figs 2-3. [small stream at an elevation of 450 m near 

Houmachong village, Chenxi County, ca. 27°47'N 110°16'E] 

Paracaridina guizhouensis (Liang & Yan, 1986a) 

= Caridina guizhouensis Liang & Yan, 1986a: 199; Fig. 2. [Maopo, Yuping County, Guizhou] 

Paracaridina leptocarpa (Liang & Zheng, 1988) 

= Caridina leptocarpa Liang & Zheng, 1988: 15; Figs 1-9. [Fuzhou, 26°N 119°E, Min River, Fujian] 

Paracaridina longispina (Guo, He, Xu & Gui, 1992) 

= Caridina longispina Guo, He, Xu & Gui, 1992: 717; Figs 1-9. [Yuelu Hills, Changsha, China, 112°56'E 

28°10'N] 

Paracaridina zijinica Liang, 2002b: 171; Fig. 3. [Zijin, Guangdong, southern China] 

Paratya Miers, 1882 

= Paratya Miers, 1882 (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850], by 

monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

470 in 1957 and in Direction 85 in 1957) 

= Xiphocaridina Bouvier, 1909b (type species Ephyra compressa De Haan, 1844 [in De Haan, 1833- 

1850], designated by Holthuis, 1955b, gender feminine; name placed on the Official Index of Rejected 

and Invalid Names in Zoology in Opinion 470 in 1957) 

= Xiphatyoida Roux, 1915 (established without included nominal species; type species Paratya 

(Xiphatyoida) typa Roux, 1926a, designated by Roux, 1926a, gender feminine) 

Paratya annamensis Balss, 1924: 45. [Annam, Phuc Son] 

Paratya australiensis Kemp, 1917b: 303; Fig. 5. [Seven Hills, near Sydney; neotype designation by W.D. 

Williams & Smith, 1979] 

= Paratya australiensis arrostra Riek, 1953: 114, Fig. 2. [Terrors Creek, Dayboro, Queensland] 

= Paratya atacta Riek, 1953: 114; Fig. 3. [Upper Nerang River, south Queensland] 

= Paratya atacta adynata Riek, 1953: 115; Fig. 4. [small creek in upper reaches of Middle Harbour, 

Sydney, N.S.W.] 

= Paratya tasmaniensis Riek, 1953: 115; Fig. 5. [small stream at Kingston, Tasmania] 

Paratya boninensis Satake & Cai, 2005: 306; Figs 1-2. [Oku-ohotaki river, a tributary of the west inlet 

stream of Shigure Dam, 210-220 m above sealevel, Chichijima Island, Ogasawara Islands, Japan] 

Paratya borealis Volk, 1938: 123; Figs 1-4. [Uluncha River, southern part of the Peter the Great Basin, Japan 

Sea] 

Paratya bouvieri Roux, 1926a 

= Paratya (Xiphatyoida) bouvieri Roux, 1926a: 189; Figs 1-5. [Nouvelle-Calédonie, Rivière du Mt. Panié] 

= Paratya (Xiphatyoida) bouvieri ngoiensis Roux, 1926a: 190; Figs 6-8. [Nouvelle-Calédonie, Vallée de 

Ngoï, 200 m alt.]


Parataya caledonica Roux, 1926a 

= Parataya (Xiphatyoida) caledonica Roux, 1926a: 192; Figs 9-16. [Nouvelle-Calédonie, Lac en 8] 

= Paratya caledonica var. magna Roux, 1926a: 194; Figs 17-19. [Nouvelle-Calédonie, Lac en 8] 

Paratya compressa (De Haan, 1844 [in De Haan, 1833-1850]) 

= Ephyra compressa De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 7 (1844); 186, Plate O (1849). 


Paratya improvisa Kemp, 1917b 

= Paratya compressa improvisa Kemp, 1917b: 299; Figs 2-3. [Lake Haruna, near Ikao, altitude about 

3000ft] 

Paratya curvirostris (Heller, 1862a) 

= Caridina curvirostris Heller, 1862a: 525. [Aukland] 

= Palæmon (Leander) fluviatilis Thomson, 1879: 231. [Waikato River; Taieri River; lagoons in Taieri 

plain] 

Paratya howensis Roux, 1926b 

= Paratya (Xiphatyoida) howensis Roux, 1926b: 244. [Big Creek, Lord Howe Island] 

Paratya intermedia Roux, 1926a 

= Paratya caledonica var. intermedia Roux, 1926a: 195; Figs 20-22. [Nouvelle-Calédonie, Plaine des 

Lacs, 200 m alt.] 

Paratya martensi Roux, 1925: 146. [Insel Adenare bei Flores] 

Paratya norfolkensis Kemp, 1917b 

= Paratya australiensis norfolkensis Kemp, 1917b: 305; Fig. 6. [west side of Norfolk Island] 

Paratya typa Roux, 1926a 

= Paratya (Xiphatyoida) typa Roux, 1926a: 196; Figs 23-31. [Nouvelle-Calédonie, La Madeleine, Plaine 

des Lacs, 200 m] 

Parisia Holthuis, 1956a 

= Parisia Holthuis, 1956a (type species Caridina microphthalma Fage, 1946, by original designation, 


Parisia dentata Gurney, 1984: 584; Figs 12-14. [Dark zone, Grotte d'Andafiabe, Ankarana Massif, 40 km 

north of Ambilobé and 60 km south of Diego Suarez, northern province of Diego Suarez] 

Parisia deharvengi Cai & Ng, 2009: 1110; Fig. 10. [Gua (cave) Tanette, Kappang, Maros, Sulawesi] 

Parisia edentata Holthuis, 1956a: 55. [southern part of Antsingly Mts., near Bekopaka, Mahilaka Province, 

W Madagascar] 

Parisia gracilis W.D. Williams, 1964: 94; Figs 1, 2A-H. [caves 16 miles south of of Katherine, Northern 

Territories (approx. 14°30'S 132°E)] 

Parisia holthuisi Cai, 2010c: 174; Figs 1-2. [Tigibi, Tari subdistrict, Southern Highlands, Territory of Papua 

Australian New Guinea (= Papua New Guinea), 1600 m alt.] 

Parisia macrophora Cai & Anker, 2004: 253; Figs 12-14. [Nalubog Cave, Quezon Province, Luzon, Philippines] 

Parisia macrophthalma Holthuis, 1956a: 55. [Grotte des Fanihy, Ankarana Mts., N of Ambilobé, N.-W. 

Madagascar] 

Parisia microphthalma (Fage, 1946) 

= Caridina microphthalma Fage, 1946: 324; Figs 1-2. [grotte des Fanihys, Ankarana] 

Parisia unguis W.D. Williams, 1964: 99; Figs 2L-S, 3. [caves 16 miles south of of Katherine, Northern Territories 

(approx. 14°30'S 132°E)] 

Potimirim Holthuis, 1954 

= Potimirim Holthuis, 1954 (type species Caridina mexicana de Saussure, 1857a, by original designation, 


Potimirim americana (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) 

= Caridina americana Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, Figs 13- 

13a. [la Isla de Cuba]


Potimirim glabra (Kingsley, 1878a) 

= Atyoida glabra Kingsley, 1878a: 93. [Polvon and Corcuera, west coast of Nicaragua] 

= Potimirim brasiliana Villalobos F., 1960b: 275; Figs 1-5. [Río Ariró, Angra dos Reis, Edo. De Río de 

Janeiro, Brasil] 

Potimirim mexicana (de Saussure, 1857a) 

= Caridina mexicana de Saussure, 1857a: 505. [Vera-Cruz] 

Potimirim potimirim (Müller, 1881) 

= Atyoida Potimirim Müller, 1881: 117; Figs 1, 8-15. [Itajahy] 

Puteonator Gurney, 1987 

= Puteonator Gurney, 1987 (type species Puteonator iraqiensis Gurney, 1987, by original designation 


Puteonator iraqiensis Gurney, 1987: 162; Figs 1-4. [southern Iraq, Samawa, anchialine habitat at 160 m] 

Pycneus Holthuis, 1986b 

= Pycneus Holthuis, 1986b (type species Pycneus morsitans Holthuis, 1986b, by original designation 


Pycneus morsitans Holthuis, 1986b: 105; Figs 1-2. [cave at Munjingerra, Gibson Desert, Western Australia, 

at about 22°30'S 124°10'E] 

Pycnisia Bruce, 1992a 

= Pycnisia Bruce, 1992a (type species Pycnisia raptor Bruce, 1992a, by original designation and monotypy, 


Pycnisia bunyip Suzuki & Davie, 2003: 447; Figs 1-4. [Forbes Inferno Cave, Riversleigh, Lawn Hill National 

Park, northwestern Queensland, in shallow subterranean pools] 

Pycnisia raptor Bruce, 1992a: 554; Figs 1-6. [Cave 8MD26, Katherine Region, Northern Territory, 13°55'S 

132°28'E, 33.5 m] 

Sinodina liang & Cai, 1999 

= Sinodina Liang & Cai, 1999 (type species Caridina gregoriana Kemp, 1923, by original designation, 


Sinodina acutipoda (Liang, 1989) 

= Caridina acutipoda Liang, 1989: 282; Figs 1-10. [Lugu Lake, 27°43'N 100°46'E] 

Sinodina angulata Liang, 2002a: 118; Fig. 1. [Longtan stream, Jiangchuan County, Yunnan Province, China] 

Sinodina banna (Cai & Dai, 1999) 

= Caridina banna Cai & Dai, 1999: 214; Figs 3-4. [Runjinghong, Hinghong County] 

= Sinodina bannica Liang, 2004: iv. [Erroneous spelling] 

Sinodina bispinosa (Liang, 1990) 

= Caridina bispinosa Liang, 1990: 221; Fig. 3. [Yunnan Province, Chuxiong (Jiulongdian)] 

Sinodina dianica Liang & Cai, 1999: 583; Figs 3-4. [Malipo village, Wenshan County, south Yunnan] 

Sinodina gregoriana (Kemp, 1923) 

= Caridina gregoriana Kemp, 1923: 437; Figs 1-2. [Tali-fu (Erhai Lake), near Dali city, Yunnan; lectotype 

designation by Liang & Cai, 1999] 

Sinodina heterodactyla (Liang & Yan, 1985a) 

= Caridina heterodactyla Liang & Yan, 1985a: 198; Fig. 2. [rivulet, Shizong, Yunnan Province] 

Sinodina leptopropoda Liang, 1990 

= Caridina leptopropoda Liang, 1990 (sensu Yü, 1938 (partim)): 220; Fig. 2. [several localities in Yunnan] 

Sinodina lijiang Liang & Cai, 1999: 587; Fig. 7. [Lijiang County, Yunnan Province] 

Sinodina wangtai Liang & Cai, 1999: 585; Figs 5-6. [Wangtai village, Lijing County] 

Sinodina yongshengica Chen & Liang, 2002: 240; Fig. 1. [Taoyuan, Yongsheng County, Yunnan Province, 

26°75'N 100°70'E, 1100 m]


Sinodina yui (Liang & Yan, 1985a) 

= Caridina yui Liang & Yan, 1985a: 200; Fig. 4. [Shuanglongyin, Qiubei County, Yunnan Province] 

Stygiocaris Holthuis, 1960 

= Stygiocaris Holthuis, 1960 (type species Stygiocaris lancifera Holthuis, 1960, by original designation, 


Stygiocaris lancifera Holthuis, 1960: 48; Figs 1-2. [Kuddumurra Well, Yardie Creek Station, North-West 

Cape Peninsula, W. Australia] 

Stygiocaris stylifera Holthuis, 1960: 54; Fig. 3. [Kuddumurra Well, Yardie Creek Station, North-West Cape 

Peninsula, W. Australia] 

Syncaris Holmes, 1900 

= Syncaris Holmes, 1900 (type species Miersia pacifica Holmes, 1895, by original designation, gender 


Syncaris pacifica (Holmes, 1895) 

= Miersia pacifica Holmes, 1895: 577; Plate 21, figs 27-28. [Sonoma County] 

Syncaris pasadenae (Kingsley, 1897) 

= Caradina pasadenæ Kingsley, 1897: 98; Plate 3, figs 1-7. [streams about Pasadena, California] 

= Syncaris Trewi Holmes, 1900: 213; Plate 4, fig. 63. [small stream near San Gabriel, Los Angeles 

County, California] 

Troglocaris Dormitzer, 1853 

= Troglocaris Dormitzer, 1853 (type species Troglocaris Schmidtii Dormitzer, 1853 (a junior subjective 

synonym of Palaemon anophthalmus Kollar, 1848), by monotypy, gender feminine; name placed on 

the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

= Troglocaridella Babić, 1922 (type species Troglocaridella hercegovinensis Babić, 1922, by monotypy, 


= Xiphocaridinella Sadowsky, 1930 (type species Xiphocaridinella kutaissiana Sadowsky, 1930, by 


= Spelaeocaris Matjašič, 1956 (type species Spelaeocaris pretneri Matjašič, 1956, by monotypy, gender 

feminine) 

Troglocaris (Troglocaridella) hercegovinensis (Babić, 1922) 

= Troglocaridella hercegovinensis Babić, 1922: 303; Fig. 5. [Höhle Vjeternica bei Zavala (Hercegovina)] 

= Troglocaris hercegovinensis Absolon, 1916: 609. [nomen nudum] 

Troglocaris (Troglocaris) anophthalmus anophthalmus (Kollar, 1848) 

= Palaemon anophthalmus Kollar, 1848: 137. [aus den unterirdischen Gewässern von Krain] 

= Troglocaris Schmidtii Dormitzer, 1853: 87. [Grotte von Kumpole in Unterkrain] 

= Troglocaris Schmidti Schmidti forma typica Fage, 1937: 226. [Type locality not indicated] 

Troglocaris (Troglocaris) anophthalmus intermedia Babić, 1922 

= Troglocaris schmidti var. intermedia Babić, 1922: 300; Figs 1-4. [Mikašinović’s Höhle (in der Nähe von 

Gornje Dubrave), ungefähr 6 km von Ogulin] 

Troglocaris (Troglocaris) bosnica Sket & Zakšek, 2009: 799; Figs 7-8. [Bosnia and Herzegovina, Bosanka 

Krajina, Lušci polje, Suvaja cave] 

Troglocaris (Troglocaris) planinensis Birstein, 1948 

= Troglocaris schmidti planinensis Birstein, 1948: 5; Fig. 2. [Planina Cave, 9 km from Postoina Cave, 

Rieka River, Yugoslavia] 

Troglocaris (Spelaeocaris) kapelana Sket & Zakšek, 2009: 806; Fig. 11Ska. [Croatia, Jezerane, Obajdini, Obajdinova 

pećina cave] 

Troglocaris (Spelaeocaris) neglecta Sket & Zakšek, 2009: 808; Figs 11Sne, 12. [Croatia, Novigrad, Karišnica 

cave near Karin] 

Troglocaris (Spelaeocaris) prasence Sket & Zakšek, 2009: 804; Figs 9-10, 11Spra. [Bosnia and Herzegovina, 

Fatničko polje, Obod cave]


Troglocaris (Spelaeocaris) pretneri (Matjašič, 1956) 

= Spelaeocaris pretneri Matjašič, 1956: 66; Figs 1-2. [einer Höhle aus der südöstlichen Herzegowina (in 

der Nähe von Bileće)] 

Troglocaris (Xiphocaridinella) ablaskiri Birstein, 1939 

= Troglocaris schmidti ablaskiri Birstein, 1939: 961; Figs 1-2. [Abkhasia, cave near Achkshe-Tyz-Gua] 

Troglocaris (Xiphocaridinella) fagei Birstein, 1939 

= Troglocaris schmidti fagei Birstein, 1939: 964; Figs 3-5. [Abkhasia, cave near Psyrtzkha] 

Troglocaris (Xiphocaridinella) jusbaschjani Birstein, 1948 

= Troglocaris schmidti jusbaschjani Birstein, 1948: 4; Fig. 1. [Russian Federation, Krasnodarskiy Krai, 

Sochi area, cave at Matzesta] 

Troglocaris (Xiphocaridinella) kutaissiana (Sadowsky, 1930) 

= Xiphocaridinella kutaissiana Sadowsky, 1930: 95. [unterirdische Gewässer in Karsthöhlen bei Kutais 

(West-Georgien)] 

Troglocaris (Xiphocaridinella) osterloffi Juzbaš’jan , 1940: 73. [Shakuran] 

Typhlatya Creaser, 1936 

= Typhlatya Creaser, 1936 (type species Typhlatya pearsei Creaser, 1936, by monotypy, gender feminine) 

Typhlatya arfeae Jaume & Bréhier, 2005: 390; Figs 3-18. [Résurgence de Font Estramar, Salses-le-château, 

Pyrénées-orientales, France] 

Typhlatya campecheae H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 6; Figs 3-4. [Grutas de Xtacumbilxunam, 

Bolonchenticul, Campeche, Mexico] 

Typhlatya consobrina Botoşăneanu & Holthuis, 1970: 123; Figs 1-2. [Cueva del Agua, Sierra de Cubitas, at 

the foot of Cerro Tuabaquei, very near to ”Finca la Entrada” (NE from the town Camagüey, prov. 

Camagüey, in a subterranean lake] 

Typhlatya dzilamensis Alvarez, Iliffe & Villalobos, 2005: 83; Figs 2-3. [Cenote Buya Uno, Dzilam de Bravo, 

Yucatan, Mexico] 

Typhlatya elenae Juarrero, 1994: 58; Figs 1-10. [Cueva Perico Sanchez, 5 km al N de Jaguey Grande, Provincia 

de Matanzas] 

Typhlatya galapagensis Monod & Cals, 1970: 70; Figs 1-46, 50, 53, 55, 58, 60, 63-64. [Ile de Santa Cruz, 

versant S-E, eau courante à peine sodique, dans une crevasse profonde de 15-20 m, à 50 m d’altitude 

et à 2 km de la côte] 

Typhlatya garciadebrasi Juarrero de Varona & Ortiz, 2000: 46; Figs 1-3. [Cueva Chicarrones, Bolondrón, 

Procincia de Matanzas] 

Typhlatya garciai Chace, 1942a: 99; Plate 29. [Potrero del Molino Cave, Las Cuatrocientas, Banes, Oriente 

Province, Cuba] 

Typhlatya iliffei C.W.J. Hart & Manning, 1981: 447; Figs 29-55. [Tucker’s Town cave, Bermuda] 

Typhlatya kakuki Alvarez, Iliffe & Villalobos, 2005: 86; Figs 4-5. [Shrimp Hole, Salinas Point, Acklins Island, 

Bahamas] 

Typhlatya miravetensis Sanz & Platvoet, 1995: 80; Figs 1-4. [Cave ”Ullal de la Rambla de Miravet” between 

the towns of Cabanes and Orpesa, province of Castellón, eastern Spain] 

Typhlatya mitchelli H.H.III Hobbs & H.H.Jr. Hobbs, 1976: 2; Figs 1-2. [Cenote Kabahchen, Maní, Yucatán, 

Mexico] 

Typhlatya monae Chace, 1954: 318; Fig. 1. [Mona Island, Puerto Rico, from well 30ft deep at ”El Molino”, 

about 1 mile southeast of NYA camp at Sardinera] 

Typhlatya pearsei Creaser, 1936: 128; Figs 31-41. [Balam Canche Cave, 4.8 km east 0.8 km south of Chichen 

Itza, Yucatan] 

Typhlatya rogersi Chace & Manning, 1972: 14; Figs 10-11. [Marl Pool, back of Shelly Beach, Ascension Island] 

Typhlatya taina Estrada & Gómez, 1987: 3; Figs 2-5. [Cueva de la India, 2 km al E de Puerto Escondido, 

Santa Cruz del Norte, La Habana, Cuba] 

Typhlatya utilaensis Alvarez, Iliffe & Villalobos, 2005: 89; Figs 6-7. [George Gaberel’s Cave, Utila, Bay Islands, 

Honduras]


Typhlocaridina liang & yan, 1981 

= Typhlocaridina Liang & Yan, 1981 (type species Typhlocaridina lanceifrons Liang & Yan, 1981, by 


Typhlocaridina lanceifrons Liang & Yan, 1981: 32; Figs 1-19. [Daji Cave, Wuming County, Guangxi] 

Typhlocaridina liui Liang & Zhou, 1993: 232; Figs 1(1-3). [Paobingyan Cave, Lingui County, Guangxi 

Province] 

Typhlocaridina semityphlata Cai, 1995b: 157; Fig. 1. [cave in Guilin, Guangxi Autonomous Region, South 

China] 

Typhlopatsa Holthuis, 1956a 

= Typhlopatsa Holthuis, 1956a (type species Typhlopatsa pauliani Holthuis, 1956a, by original designation 


Typhlopatsa pauliani Holthuis, 1956a: 53. [Mitoho Cave, N.-E. corner of Tsimanampetsotsa Lake, Mahafaly 

Province, S.-W. Madagascar] 

Superfamily BReSIlIoIDeA Calman, 1896 

Family AGoSToCARIDIDAe C.W.J. Hart & Manning, 1986 

Agostocaris C.W.J. Hart & Manning, 1986 

= Agostocaris C.W.J. Hart & Manning, 1986 (type species Agostocaris williamsi C.W.J. Hart & Manning, 


Agostocaris acklinsensis Alvarez, Villalobos & Iliffe, 2004: 370; Figs 2-4. [Jumby Hole Cave, Snug Corner, 

Acklins Island, Bahamas] 

Agostocaris bozanici Kensley, 1988: 688; Figs 1-3 [Xcan-ha Cenote (Cenote Roja), Cozumel Island, Quintana 

Roo, Mexico, 80-100 feet] 

Agostocaris williamsi C.W.J. Hart & Manning, 1986: 412; Figs 27-47. [Bahamas, 24°27'N, 75°30'W, Big 

Fountain Blue Hole] (Fig. 16) 

Fig. 16. Agostocaris williamsi C.W.J. Hart & Manning, 1986. Photo by Thomas M. Iliffe.


Family ALVINOCARIDIDAE Christoffersen, 1986 

Alvinocaridinides Komai & Chan, 2010a 

= Alvinocaridinides Komai & Chan, 2010a (type species Alvinocaridinides formosa Komai & Chan, 

2010a, by original designation and monotypy, gender feminine) 

Alvinocaridinides formosa Komai & Chan, 2010a: 26; Figs 1C, 7-10. [Gueishandao, Yilan County, Taiwan, 

24°51.231'N 121°59.204'E, 252-275 m] 

Alvinocaris Williams & Chace, 1982 

= Alvinocaris Williams & Chace, 1982 (type species Alvinocaris lusca Williams & Chace, 1982, by 


Alvinocaris alexander Ahyong, 2009: 777; Figs 1-3. [Rumble V Seamount, 36°08.27-07.96'S 178°11.74- 

11.70'E, 485-415 m] 

Alvinocaris brevitelsonis Kikuchi & Hashimoto, 2000: 136; Figs 1-3. [”Depression C” of the Minami-Ensei 

Knoll, 28°23.35'N 127°38.38'E, 705 m] 

Alvinocaris chelys Komai & Chan, 2010a: 16; Figs 1A-B, 2-6. [Gueishandao, Yilan County, Taiwan, 

24°49.682'N 122°0.254'E, 300-276 m] (Fig. 17) 

Alvinocaris dissimilis Komai & Segonzac, 2005a: 1158; Figs 25-26, 29. [Depression C, Minami-Ensei Knoll, 

28°23.35'N 127°38.38'E, 705 m] 

Alvinocaris komaii Zelnio & Hourdez, 2009: 55; Figs 1-6. [Kilo Moana, 20°9'S 76°12'E, 2620 m, Eastern Lau 

Spreading Center, Lau Basin, southwest Pacific] 

Alvinocaris longirostris Kikuchi & Ohta, 1995: 772-778, Figs 1-7. [Iheya Ridge, Clam Site, Okinawa Trough, 

27°32.70'N 126°58.20'E, 1360 m] 

Alvinocaris lusca Williams & Chace, 1982: 137; Figs 1-7. [Galapagos Rift Rose Garden area, 0°48.25'N 

86°13.48'W, maximum of 2450 m] 

Alvinocaris markensis Williams, 1988: 264; Figs 1-2, 7l. [Mid-Atlantic Rift Valley about 70 km south of 

Kane Fracture Zone, 23°22.09'N 44°57.12'W, 3437 m] 

Fig. 17. Alvinocaris chelys Komai & Chan, 2010. Photo by Tin-Yam Chan.


Alvinocaris methanophila Komai, Shank & Van Dover, 2005: 29; Figs 1-4. [ODP site 996, Blake Ridge Diapir, 

32°29.623'N 76°11.467'W, 2155 m] 

Alvinocaris muricola Williams, 1988: 268; Figs 3-4, 7e-k. [West Florida Escarpment, 26°01'N 84°54.61'W, 

3277 m] 

Alvinocaris niwa Webber, 2004: 5; Figs 1-4. [Rumble V, 36°8.63-8.57'S 178°11.77-11.50'E, 877-655 m] 

Alvinocaris stactophila Williams, 1988: 272; Figs 5-6, 7a-d. [north central Gulf of Mexico about 129 km S of 

Louisiana, 27°46.94'N 91°30.34'W, 534 m] 

Alvinocaris williamsi Shank & Martin, 2003: 159; Figs 1-3. [Menez Gwen hydrothermal field, North Atlantic 

Ocean, 37°50.5'N 31°31.3'W, 850 m] 

Chorocaris Martin & Hessler, 1990 

= Chorocaris Martin & Hessler, 1990 (type species Chorocaris vandoverae Martin & Hessler, 1990, by 


Chorocaris chacei (Williams & Rona, 1986) 

= Rimicaris chacei Williams & Rona, 1986: 455; Figs 8-10. [TAG Hydrothermal Field, Mid-Atlantic 

Ridge, 26°08.3'N 44°49.6'W, 3620-3650 m] 

Chorocaris paulexa Martin & Shank, 2005: 186; Figs 1-8. [Homer Vent (347OC black smoker), 17°37.220'S 

113°15.123'W, 2595 m, southern East Pacific Rise] 

Chorocaris vandoverae Martin & Hessler, 1990: 2; Figs 1-2, 3a-e. [Alice springs vent field, Mariana Back- 

Arc Basin, 18°12.599'N 144°42.431'E, 3640 m] 

Mirocaris Vereshchaka, 1997b 

= Mirocaris Vereshchaka, 1997b (type species Mirocaris keldyshi Vereshchaka, 1997b (junior subjective 

synonym of Chorocaris fortunata Martin & Christiansen, 1995), by monotypy, gender feminine) 

Mirocaris fortunata (Martin & Christiansen, 1995) 

= Chorocaris fortunata Martin & Christiansen, 1995: 221; Figs 1-3. [Vent site 3, Lucky Strike hydrothermal 

vent, Azores, 37°17.6'N 32°16.5'W, 1624 m] 

= Mirocaris keldyshi Vereshchaka, 1997b: 431; Figs 1-5. [TAG location, 26°09'N 44°50'W, 3650 m] 

Mirocaris indica Komai, Martin, Zala, Tsuchida & Hashimoto, 2006: 110; Figs 1-5. [Central Indian Ridge, 

Kairei Field, 25°19.2'S 70°02.4'E, 2422 m] 

Nautilocaris Komai & Segonzac, 2004 

= Nautilocaris Komai & Segonzac, 2004 (type species Nautilocaris saintlaurentae Komai & Segonzac, 


Nautilocaris saintlaurentae Komai & Segonzac, 2004: 1179-1188; Figs 2-6. [North Fiji Basin, White Lady 

site, 16°59.50'S 173°55.47'E, 2000 m] 

Opaepele Williams & Dobbs, 1995 

= Opaepele Williams & Dobbs, 1995 (type species Opaepele loihi Williams & Dobbs, 1995, by monotypy, 


Opaepele loihi Williams & Dobbs, 1995: 229; Figs 1-3. [Loihi Seamount, Hawaii, 18°55'N 155°16'W, 980 m] 

Opaepele susannae Komai, Giere & Segonzac, 2007: 240; Figs 2-7. [Lilliput, southern Mid-Atlantic Ridge, 

09°32.845'S 13°12.546'W, 1500 m, mussel field with diffuse vent fluids] 

Opaepele vavilovi Lunina & Vereshchaka, 2010: 70; Figs 1-2. [Mid-Atlantic Ridge, Broken Spur vent site, 

stn 4797] 

Rimicaris Williams & Rona, 1986 

= Rimicaris Williams & Rona, 1986 (type species Rimicaris exoculata Williams & Rona, 1986, by original 


= Iorania Vereshchaka, 1996 (type species Iorania concordia Vereshchaka, 1996 (junior subjective synonym 

of Rimicaris exoculata Williams & Rona, 1986), by monotypy, gender feminine)


Rimicaris exoculata Williams & Rona, 1986: 448; Figs 5-7. [TAG Hydrothermal Field, Mid-Atlantic Ridge, 

26°08.3'N 44°49.6'W, 3620-3650 m] 

= Iorania concordia Vereshchaka, 1996: 954; Figs 1-5. [TAG location, 26°09N 44°50W, point D, 3650 m] 

= Rimicaris aurantiaca Martin, Signorovitch & Patel, 1997: 400; Figs 1-5. [Snake Pit hydrothermal vent 

field, Mid-Atlantic Ridge, 23°22.1'N 44°57.0'W] 

Rimicaris kairei Watabe & Hashimoto, 2002: 1168; Figs 1-4. [The Central Indian ridge, Indian Ocean, the 

Kairei Field, 25°19.16'S 70°02.40'E, 2454 m] 

Shinkaicaris Komai & Segonzac, 2005a 

= Shinkaicaris Komai & Segonzac, 2005a (type species Alvinocaris leurokolos Kikuchi & Hashimoto, 


Shinkaicaris leurokolos (Kikuchi & Hashimoto, 2000) 

= Alvinocaris leurokolos Kikuchi & Hashimoto, 2000: 141; Figs 4-7. [”Depression C” of the Minami- 

Ensei Knoll, 28°23.35'N 127°38.38'E, 705 m] 

Family BReSIlIIDAe Calman, 1896 

Bresilia Calman, 1896 

= Bresilia Calman, 1896 (type species Bresilia atlantica Calman, 1896, by monotypy, gender feminine; 


Bresilia antipodarum Bruce, 1990a: 855; Figs 6-10. [Tasman Sea, off Cape Freycinet, Tasmania, 42°2.20'S 

148°38.70'E, 800 m] 

Bresilia atlantica Calman, 1896: 7; Plate 1; Plate 2, figs 15-18. [south west of Ireland, 750 fms] 

Bresilia briankensleyi Bruce, 2005a: 176; Figs 1-3. [Egyptian Red Sea, 22°85.2'N 36°45.9'E to 22°34'N 

36°46.2'E, 750-753 m] 

Bresilia corsicana Forest & Cals, 1977: 551; Figs 1-20. [canal de Corse, 545 milles dans le 285 du cap Trattoja, 

sud-ouest de l’île Capraia, 450 m] 

Bresilia gibbosa Komai & Yamada, 2010: 42; Figs 1-5. [Apo-gama Cave, Onna Village, Okinawa Island, 

Ryukyu Islands, 30 m] 

Bresilia plumifera Bruce, 1990b: 1; Figs 1-4. [Taupo Sea Mount, Western Tasman Sea, 33°14.21'S 56°10.68'E, 

133 m] 

Bresilia rufioculus Komai & Yamada, 2011: 72; Figs 1-6. [Ohoba Cave, Ie Island, Okinawa Islands] (Fig. 18) 

Fig. 18. Bresilia rufioculus Komai & Yamada, 2011. Photo by Tomoyuki Komai.


Bresilia saldanhai Calado, Chevaldonné & dos Santos, 2004: 191; Figs 1-4. [underwater cave at Ponta garajau, 

south of Madeira island, eastern Atlantic, 32.63°N 16.85°W, 15 m] 

Encantada Wicksten, 1989a 

= Encantada Wicksten, 1989a (type species Encantada spinoculata Wicksten, 1989a, by monotypy, gender 

feminine) 

Encantada spinoculata Wicksten, 1989a: 667; Figs 1-2. [Beagle Island, east of Jervis Island, Galapagos, 

0°26'S 90°40'W, 55-92 m] 

Family DISCIADIDAe Rathbun, 1902c 

Discias Rathbun, 1902c (Fig. 19) 

= Anisocaris Ortmann, 1893 (type species Anisocaris dromedarius Ortmann, 1893 (probably an invalid 

senior subjective synonym of Discias atlanticus Gurney, 1939b), by monotypy, gender feminine; 

name suppressed for the purposes of the Principle of Priority, but not for those of the Principle of 

Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology in 


= Discias Rathbun, 1902c (type species Discias serrifer Rathbun, 1902c, by monotypy, gender masculine) 

Discias atlanticus Gurney, 1939b: 388; Figs 1-13. [in plankton at night in the shallow waters of the Reach 

close to the Bermuda Biological Station] 

? = Anisocaris dromedarius Ortmann, 1893: 74; Plate 4, figs 3-3z. [Plankton Expedition, Nördl. 

Aequa torialstrom, JN 148 (10.2°N 22.2°W, 0-400 m); Südl. Aequatorialstrom, JN 218 (3.8°S 32.6°W, 

0-400 m)] 

Discias brownae Kensley, 1983: 8; Figs 6-9. [Green Point, Port Jackson, New South Wales, 33°50'S 151°19'E, 

9.8 m] 

Fig. 19. Discias spec. Photo by Leslie Harris.


Discias exul Kemp, 1920: 138; Figs 1-3. [Port Blair, Andaman Islands, on the reef at the N end of Ross Island, 

low water] 

= Discias mvitae Bruce, 1976b: 119; Figs 1-5. [Fort Jesus, Mombasa, 4°04.0'S 39°42.18'E, 1 m] 

Discias musicus Holthuis, 1981: 787; Figs 1-2. [Lagoon west of Saipan, Marianas Islands, immediately 

shoreward of barrier reef flat] 

= Discias musicus Holthuis, 1953b: 52. [nomen nudum] 

Discias pascuensis Fransen, 1987: 501; Figs 1-3. [Tahai, west coast of Easter Island, Chile, 39 m] 

Discias serratirostris Lebour, 1949a: 1107; Figs 1-2. [about three miles off Castle Roads, Bermuda, near 

bottom, at about 100ft or more] 

Discias serrifer Rathbun, 1902c: 290; Figs 1-4. [Tagus Cove, Albemarle Island, on the reef north of Tagus 

Hill] 

Discias vernbergi Boothe & Heard, 1987: 506; Figs 1-3. [65 nautical miles W of Egmont Key, Florida, 

27°37'N 83°58'W, 55 m] 

Kirnasia Burukovsky, 1988 

= Kirnasia Burukovsky, 1988 (type species Kirnasia nesisi Burukovsky, 1988, by original designation, 


Kirnasia nesisi Burukovsky, 1988: 457; Fig. 1. [42°39'N, 28°45'W, 850-1200 m] 

Kirnasia siedlecki Burukovsky, 1988: 458; Fig. 2. [36°18'N, 19°50'W, 874 m] 

Lucaya Chace, 1939 

= Lucaya Chace, 1939 (type species Lucaya bigelowi Chace, 1939, by monotypy, gender feminine) 

Lucaya bigelowi Chace, 1939: 34. [east of Great Abaco Island, Bahamas, 26°12'N, 76°26'W, 2610 fms] 

Tridiscias Kensley, 1983 

= Tridiscias Kensley, 1983 (type species Tridiscias transkei Kensley, 1983, by monotypy, gender masculine) 

Tridiscias transkei Kensley, 1983: 18; Figs 15-17. [off Transkei, South Africa, 31°59'S 29°22.5'E, 150-200 m] 

Family PSeuDoCHelIDAe De Grave & Moosa, 2004 

Pseudocheles Chace & Brown, 1978 

= Pseudocheles Chace & Brown, 1978 (type species Pseudocheles enigma Chace & Brown, 1978, by 


Pseudocheles chacei Kensley, 1983: 22; Figs 18-22. [Looe Key, Florida, 6 m] 

Pseudocheles enigma Chace & Brown, 1978: 757; Figs 1-4. [west side of lagoon, Lizard Island, Queensland, 

Australia, 1-15 m] 

Pseudocheles neutra De Grave & Moosa, 2004: 1; Figs 1-4. [Kaledupa Reef, Kaledupa, Tukangbesi Archipelago, 

south-eastern Sulawesi, Indonesia, 10 m] 

Superfamily NeMAToCARCINoIDeA Smith, 1884 

Family euGoNAToNoTIDAe Chace, 1937a 

Eugonatonotus Schmitt, 1926a 

= Gonatonotus A. Milne-Edwards, 1881b (type species Gonatonotus crassus A. Milne-Edwards, 1881b, 

by monotypy, gender masculine; invalid junior synonym of Gonatonotus Adams & White, 1847 

(Crustacea Brachyura); name placed on the Official Index of Rejected and Invalid Generic Names in 


= Eugonatonotus Schmitt, 1926a (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, type species 

therefore Gonatonotus crassus A. Milne-Edwards, 1881b, gender masculine; name placed on the 

Official List of Generic Names in Zoology in Opinion 470 in 1957)


Fig. 20. Eugonatonotus crassus (A. Milne-Edwards, 1881). Photo Pillsbury Expedition sta 936b. 

= Gomphonotus Chace, 1936 (nomen novum for Gonatonotus A. Milne-Edwards, 1881b, gender masculine; 

name placed on the Official Index of Rejected and Invalid Generic Names in Zoology in 


= Galatheacaris Vereshchaka, 1997a (type species Galatheacaris abyssalis Vereshchaka, 1997a, by original 


Eugonatonotus crassus (A. Milne-Edwards, 1881b) (Fig. 20) 

= Gonatonotus crassus A. Milne-Edwards, 1881b: 10. [Grenade par 262 brasses de profondeur] 

= Gonatonotus crassus var. longirostris A. Milne-Edwards, 1883: Planche 34. [Off Sand Key] 

Eugonatonotus chacei Chan & Yu, 1991a: 144; Fig. 1; Plate 1A. [Ta-Shi, I-Lan County, Taiwan] 

= Galatheacaris abyssalis Vereshchaka, 1997a: 365; Figs 1-9. [Celebes Sea, 01°50'N 119°20'E, 4940-4970 m] 

Family NeMAToCARCINIDAe Smith, 1884 

Lenzicarcinus Burukovsky, 2005 

= Lenzicarcinus Burukovsky, 2005 (type species Lenzicarcinus struncki Burukovsky, 2005, by original 


Lenzicarcinus struncki Burukovsky, 2005: 572; Figs 1-2. [Atlantic Ocean, south of Ascension Island, 12°11'S 

6°W, 2000 m] 

Lipkius yaldwyn, 1960 

= Lipkius Yaldwyn, 1960 (type species Lipkius holthuisi Yaldwyn, 1960, by original designation and 


Lipkius holthuisi Yaldwyn, 1960: 16; Fig. 1. [41°39'30''S 175°17'E, between 50 and 200 fms] 

Nematocarcinus A. Milne-edwards, 1881b (Fig. 21) 

= Nematocarcinus A. Milne-Edwards, 1881b (type species Nematocarcinus cursor A. Milne-Edwards, 

1881b, by monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology 

in Opinion 470 in 1957) 

= Eumiersia Smith, 1882 (type species Eumiersia ensifera Smith, 1882, by monotypy, gender feminine) 

= Stochasmus Spence Bate, 1888 (type species Stochasmus exilis Spence Bate, 1888, by monotypy, 


Nematocarcinus africanus Crosnier & Forest, 1973: 101; Figs 29a, 30a-e, 31a-b. [9°05'N 15°10'W, 310-380 m]


Fig. 21. Nematocarcinus spec. Photo by Tin-Yam Chan. 

Nematocarcinus agassizii Faxon, 1893: 204. [Albatross stn 3358 (06°30'N 081°44'W, 555 fms); lectotype designation 

by Burukovsky, 2001] 

Nematocarcinus batei Burukovsky, 2000a: 284; Figs 1b-c. [Challenger stn 237, 34°37'N 140°32'E, near Yogama, 

Japan, 3470 m] 

Nematocarcinus bituberculatus Chace, 1986: 69; Fig. 37. [west of Halmahera, Indonesia, 0°12'15"N 

127°29'30"E, 527 m] 

Nematocarcinus chacei Burukovsky, 2002a: 5; Fig. 1. [Indian Ocean, Gulf of Aden, 11°31'40''N 42°46'40''E, 

335 m] 

Nematocarcinus challengeri Burukovsky, 2006a: 896; Fig. 1. [Challenger stn 250, Pacific Ocean, 37°49'N 

166°47'W, 5477 m] 

Nematocarcinus combensis Burukovsky, 2000b: 1038; Fig. 2. [south-west part of Indian Ocean, Comb Bank, 

12°15'S 177°28'W, 800-810 m] 

Nematocarcinus crosnieri Burukovsky, 2000c: 666; Fig. 3. [New Caledonia, 23°19'S 167°59'E 850-920 m] 

Nematocarcinus cursor A. Milne-Edwards, 1881b: 14. [la Mer des Antilles, 500 brasses] 

Nematocarcinus ensifer (Smith, 1882) 

= Eumiersia ensifera Smith, 1882: 77; Plate 13, Figs 1-9. [Blake stns 305, 41°33'15''N 65°51'25''W, 810 


73°10'30''W, 1186 fms; 340, 39°25'30''N 70°58'40''W, 1394 fms; 341, 39°38'20''N 70°56'0''W, 1241 

fms] 

Nematocarcinus evansi Burukovsky, 2000d: 1291; Fig. 2. [south-west Australia, 20°16.3'S 113°13.5'E, 913- 

916 m] 

Nematocarcinus exilis (Spence Bate, 1888) 

= Stochasmus exilis Spence Bate, 1888: 823; Plate 132, fig. 14. [Challenger stn 84, 30°38'N 18°5'W, off the 

Canary Islands] 

Nematocarcinus faxoni Burukovsky, 2001: 1429; Figs 1-2. [Eastern Pacific, 16°33'N 99°52'W, 1208 m]


Nematocarcinus gladius Macpherson, 1984: 67; Figs 13-14, 15a-b. [34°37'S 17°03'E, 1580-1620 fms; exact 

locality not indicated in publication, information from IZIKO collection label] 

Nematocarcinus gracilipes Filhol, 1884: 232; Fig. 1. [pêché à 850 mètres de profondeur; no further indication 

of type locality] 

Nematocarcinus gracilis Spence Bate, 1888: 815; Plate 132, fig. 8. [Challenger stn 174c, 19°7'50"S 178°19'35"E, 

off Kandavu, Fiji Islands, 610 fms; lectotype designation by Burukovsky, 2007a] 

Nematocarcinus hanamuri Burukovsky, 2000d: 1290; Fig. 1. [south-west Australia, 20°16.3'S, 113°13.5'E, 

913-916 m] 

Nematocarcinus hiatus Spence Bate, 1888: 821; Plate 132, fig. 12. [Challenger stn 169, 37°34'S 179°22'E, off 

New Zealand, 700 fms] 

Nematocarcinus kaiensis Burukovsky, 2000b: 1040; Fig. 3. [Indonesia, Selan-Timur Archipelago, 05°15'S 

133°59'E, 769-800 m] 

Nematocarcinus lanceopes Spence Bate, 1888: 804; Plate 131. [Challenger stn 152, 60°52'S 80°20'E, Antarctic 

Sea, 1260 fms] 

= Acanthephyra antarctica Bage, 1938: 6; Plate 4, fig. 1. [63°13'S 101°42'E, 870 fms] 

Nematocarcinus longirostris Spence Bate, 1888: 806; Plate 132, fig. 2. [Challenger stn 237, 34°37'N 140°32'E, 

near Yokohama, Japan, 1875 fms] 

Nematocarcinus machaerophorus Burukovsky, 2003: 116; Fig. 33. [Pacific Ocean, Eiao Island, 07°52'S 

140°39'W, 1000 m] 

= Nematocarcinus machaerophorus Burukovsky, 2004: 1181; Figs a-e. [Pacific Ocean, Eiao Island, 

07°52'S 140°39'W, 1000 m] 

Nematocarcinus manningi Burukovsky, 2003: 118; Fig. 34. [North Pacific Ocean, 35°06'45''N 139°19'45''E, 

1138 m] 

Nematocarcinus novaezealandicus Burukovsky, 2006b: 441; Fig. 1. [Chatham Rise, 42°51.7'-42°49.5'S 

175°23.3'-175°46.1'W, 1057 m] 

Nematocarcinus nudirostris Burukovsky, 1991a: 41; Fig. 1(5-8). [9°33'5"-9°35'S 59°52'7"-59°58'8"E, 830-430 m] 

= Nematocarcinus rectirostris Burukovsky, 1991a: 39; Fig. 1(1-4). [25°28'-25°29'S 35°14'-35°11'E, 715- 

700 m] 

Nematocarcinus ovalis Komai & Segonzac, 2005b: 355; Figs 8-10. [East Pacific Rise, 12°45.50'N 103°55.80'W, 

2558-2619 m (c. 120 m above bottom)] 

Nematocarcinus paucidentatus Spence Bate, 1888: 816; Plate 132, fig. 9. [Challenger stn 174c, off Kandavu, 

Fiji Islands, 19°7'50''S 178°19'35'', 610 fms] 

= Nematocarcinus alisae Burukovsky, 2000e: 903; Fig. 4. [New Caledonia, 21°05'S 165°50'E, 960-1100 m] 

Nematocarcinus parvus Burukovsky, 2000f: 1163; Fig. 7. [Madagascar, 12°26'-12°27'S 42°08'5''-48°13'E, 600- 

705 m] 

Nematocarcinus poupini Burukovsky, 2007b: 1; Fig. 1. [Austral Islands, East of Rapa Island, 27°38.2'S 

144°14.4'W, 840-1200 m] 

Nematocarcinus productus Spence Bate, 1888: 810; Plate 132, fig. 5. [Challenger stns 205, 16°42'N 119°22'E, 

off Luzon, Philippine Islands, 1050 fms; 195, 4°21'S 129°7'E, off Banda Island, 1425 fms; 237, 34°37'N 

140°32'E, near Yokohama, Japan, 1875 fms; 176, 18°30'S 173°52'E, off the New Hebrides, 1450 fms] 

= Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fig. 6. [Challenger stn 218, 2°33'S 

144°4'E, off New Guinea, 1070 fms ] 

= Nematocarcinus intermedius Spence Bate, 1888: 821; Plate 132, fig. 13. [Challenger stn 218, 2°33'S 

144°4'E, off New Guinea, 1070 fms] 

Nematocarcinus proximatus Spence Bate, 1888: 808; Plate 132, fig. 3. [Challenger stn 146, 33°42'S 78°18'W, 

west of Valparaiso, 1375 fms; lectotype designation by Burukovsky, 2003] 

Nematocarcinus pseudocursor Burukovsky, 1990: 194; Fig. 2B. [25°07'S 99°26'W, 780-790 m] 

Nematocarcinus pseudogracilis Burukovsky, 2007a: 807; Figs 5-7. [Pacific Ocean, Austral Islands, Tubuai, 

23°12.3'S 149°34.4'W, 750-1000 m] 

Nematocarcinus richeri Burukovsky, 2000f: 1155; Figs 1-3. [Indonesia, Kai Islands, 05°15'S 133°01'E, 605- 

576 m] 

Nematocarcinus romenskyi Burukovsky, 2000a: 288; Fig. 5. [Atlantic Ocean, Discovery Bank, 42°3'8''S 

00°03'W, 1030-1050 m]


Nematocarcinus rotundus Crosnier & Forest, 1973: 103; Figs 29c, 30f-i, 31e-f. [28°43'N 87°14'30''W, 960 m] 

Nematocarcinus safari Burukovsky, 2000c: 662; Fig. 1. [south-west part of Indian Ocean, Madagascar sink, 

30°42'9''S 48°20'6''E, 4297 m] 

Nematocarcinus serratus Spence Bate, 1888: 819; Plate 132, fig. 11. [Challenger stn 169, 37°34'S 179°22'E, off 

New Zealand, 700 fms] 

Nematocarcinus sigmoideus Macpherson, 1984: 63; Figs 10-12 [area del Cabo entre 840 y 2200 m; Banco 

Valdivia, entre 754 y 940 m; costas de Namibia (28°19'S 14°18,6'E)] 

Nematocarcinus subtegulisfactus Burukovsky, 2000c: 664; Fig. 2. [Philippines, 12°05'N 122°14'E, 1404 m] 

Nematocarcinus subtilis Burukovsky, 2000b: 1042; Fig. 4. [Philippines, 11°58'N 121°06'E, 448-466 m] 

Nematocarcinus tenuipes Spence Bate, 1888: 812 (partim); Plate 132, fig. 6. [Challenger stns 235, 34°07'N 

138°0'E, south of Japan, 565 fms; 218, 2°33'S 144°04'E, near the Admiralty Islands, 1070 fms; 232, 

35°11'N 139°28'E, Japan, 345 fms] 

= Nematocarcinus parvidentatus Spence Bate, 1888: 814; Plate 132, fig. 7. [Challenger stn 237, 34°37'N 

140°32'E, near Yokohama, Japan, 1875 fms] 

= Nematocarcinus serratirostris Burukovsky, 1991a: 41; Fig. 1(9-11). [24°58'-25°05'S 35°40'-35°34'E, 

1010-910 m] 

Nematocarcinus tenuirostris Spence Bate, 1888: 817; Plate 132, fig. 10. [Challenger stns 174c, 19°7'50"S 

178°19'35"E, off Kandavu, Fiji Islands, 610 fms; 214, 4°33'N 127°6'E, south of the Philippine Islands, 

500 fms] 

= Nematocarcinus tenuirostris var. sibogae De Man, 1917: 279. [10°48'.6S 123°23'.1E, 918 m] 

Nematocarcinus tuerkayi Burukovsky, 2005: 573; Fig. 3. [Indian Ocean, Gulf of Aden, 12°52.5'N 45°53.3'E, 

1185 m] 

Nematocarcinus undulatipes Spence Bate, 1888: 801; Plate 130. [Challenger stn 200, 6°47'N 122°28'E, off 

Sibago, Philippine Islands, 250 fms; lectotype designation by Burukovsky, 2002a] 

Nematocarcinus webberi Burukovsky, 2006b: 444; Fig. 2. [Off New Zealand, 42°41.7'S 174°28'E, 1723-1549 m] 

Nematocarcinus yaldwyni Burukovsky, 2006b: 445; Fig. 3. [NE of Ritchie Bane, New Zealand, 39°47.9'S 

178°17.9'E, 832-788 m] 

Nigmatullinus Burukovsky, 1991a 

= Nigmatullinus Burukovsky, 1991a (type species Nematocarcinus acanthitelsonis Pequegnat, 1970, by 

original designation and monotypy, gender masculine) 

Nigmatullinus acanthitelsonis (Pequegnat, 1970) 

= Nematocarcinus acanthitelsonis Pequegnat, 1970: 69; Figs 4-3, 4-4. [Southwestern Gulf of Mexico, 

23°36'N 93°57'W, 2045 fms] 

Segonzackomaius Burukovsky, 2011 

= Segonzackomaius Burukovsky, 2011 (type species Nematocarcinus burukovskyi Komai & Segonzac, 


Nematocarcinus burukovskyi Komai & Segonzac, 2005b: 346; Figs 2-7. [South East Pacific Rise, Garrett Segment, 

Wormwood site, 17°34.91'S 113°14.68'W, 2595 m] 

Nematocarcinus altus Spence Bate, 1888: 809; Plate 132, fig. 4. [Challenger stn 198, 2°55'N 124°53'E, south 

of the Philippine Islands, 2150 fms] 

Family RHyNCHoCINeTIDAe ortmann, 1890 

Cinetorhynchus Holthuis, 1995 

= Rhynchocinetes (Cinetorhynchus) Holthuis, 1995 (type species Rhynchocinetes rigens Gordon, 1936, 

by original designation, gender masculine) 

Cinetorhynchus brucei Okuno, 2009a: 940; Figs 1-3. [Imazuni, Kume-jima Island, Rykyu Islands, Japan, 

26°21.8'N 126°49.6'E, 18 m] 

Cinetorhynchus concolor (Okuno, 1994a) 

= Rhynchocinetes concolor Okuno, 1994a: 66; Figs 1-2, 3A, 4A-D. [26°11.2'N 127°16.8'E, Hizushi-hama, 

Aka-jima Islet, Kerama Group, Ryukyu Islands, 3 m]


Fig. 22. Cinetorhynchus manningi Okuno, 1996. Photo by Arthur Anker. 

Cinetorhynchus erythrostictus Okuno, 1997a: 36; Plate 1A-B; Figs 2A, 3, 4A-C. [Ryukyu Islands, Aka Harbor, 

Aka-jima Islet, Kerama Group, 26°11.2'N 127°17.1'E, 3 m] 

Cinetorhynchus fasciatus Okuno & Tachikawa, 1997: 16; Figs 1-4. [Tenno-ura beach, Chichi-jima Island, 

Ogasawara Islands, Japan, 5 m] 

Cinetorhynchus hawaiiensis Okuno & Hoover, 1998: 33; Figs 1-3, 4A-B, 5A. Hawaiian Islands, 19°39.0'N 

156°00.0'W, Kailua Harbor, Hawai’i, 8 m] 

Cinetorhynchus hendersoni (Kemp, 1925) 

= Rhynchocinetes hendersoni Kemp, 1925: 265; Figs 3-7. [Pamban and Kilakarai, Gulf of Manaar] 

= Rhynchocinetes intermedius Edmondson, 1952: 72; Fig. 3. [of the southwest coast of Oahu, at about 

16 fms] 

= Rhynchocinetes marshallensis Edmondson, 1952: 75; Figs 4-6. [Ebenina, Eniwetok, Marshall Islands] 

Cinetorhynchus hiatti (Holthuis & Hayashi, 1967) 

= Rhynchocinetes hiatti Holthuis & Hayashi, 1967: 162; Figs 1-2. [Formosa, Kosho Bay, southern extremity 

of Formosa, 3 or 4 m] 

= Rhynchocinetes hiatti Holthuis, 1953b: 54. [nomen nudum] 

Cinetorhynchus manningi Okuno, 1996a: 725; Figs 1-2. [Caribbean Sea, Virgin Islands, Eagle Shoal, 10.5 

m] (Fig. 22) 

Cinetorhynchus reticulatus Okuno, 1997a: 49; Plate 1G-H; Figs 10-11, 12A-C. [Loyalty Islands, Banya Islet, 

Uvea Island, 20°35.8'S 166°16.7'E, 27 m] 

Cinetorhynchus rigens (Gordon, 1936) 

= Rhynchocinetes rigens Gordon, 1936: 76; Figs 1-4, 5e. [Potinha Bay, Madeira] 

Cinetorhynchus striatus (Nomura & Hayashi, 1992) 

= Rhynchocinetes striatus Nomura & Hayashi, 1992: 199; Figs 1-4. [Kadena Port, Okinawa Island, 1-10 m] 

Rhynchocinetes H. Milne edwards, 1837 

= Rhynchocinetes H. Milne Edwards, 1837 (type species Rhynchocinetes typus H. Milne Edwards, 


Rhynchocinetes albatrossae Chace, 1997: 28; Figs 15-16. [Surigao Strait, east of Leyte, 10°27'30"N 125°18'E, 

123 m] 

Rhynchocinetes australis Hale, 1941: 270; Fig. 8. [Edithburgh, South Australia]


Rhynchocinetes balssi Gordon, 1936: 85; Fig. 7a-b; nomen novum for Rhynchocinetes typus sensu Balss, 

1922 nec H. Milne Edwards, 1837. [Masatierra] 

Rhynchocinetes brucei Okuno, 1994b: 29; Figs 1-3, 4a-b; Plate 1. [Long Ke Wan, Hong Kong, 8 m] 

Rhynchocinetes conspiciocellus Okuno & Takeda, 1992: 64; Figs 1-3, 4A-D; Plate 1A-B. [Sokodo Beach, 

Hachijo-jima Island, Izu Islands, Japan, 33°07'N 139°49'E, 1 m] 

Rhynchocinetes durbanensis Gordon, 1936: 83; Figs 5b-b, 7c-d; nomen novum for Rhynchocinetes typus 

sensu Stebbing, 1917c nec H. Milne Edwards, 1837. [Durban] 

Rhynchocinetes enigma Okuno, 1997b: 13; Figs 1-3. [34°17'S 132°42'E, Great Australian Bight, approx. 15 

km west-south-west off Pearson Islands, 140-160 m] 

Rhynchocinetes holthuisi Okuno, 1997c: 43; Figs 1-3, 4a-e. [Aqaba, Gulf of Aqaba, Jordan] 

Rhynchocinetes ikatere Yaldwyn, 1971: 87. [off Mayor Island, Bay of Plenty, 37°15'S 176°12'E, 80-120 fms] 

Rhynchocinetes kuiteri Tiefenbacher, 1983: 121; Figs 1-3. [before Portsea at the mouth of Port Phillip Bay, 

Victoria] 

Rhynchocinetes rathbunae Okuno, 1996b: 309; Figs 1-3. [Hawaiian Islands, O’ahu, Waimea Bay, 21°38.0'N 

158°4.0'W] 

Rhynchocinetes serratus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= H.[ippolyte] serratus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 377. [la baie de 

Jarvis] 

= Rhynchocinetes rugulosus Stimpson, 1860a: 36. [in portu Jacksoniensi Australiae] 

Rhynchocinetes typus H. Milne Edwards, 1837: 168; Plate 4c. [Type locality not indicated] 

Rhynchocinetes uritai Kubo, 1942b: 30; Figs 1-3. [Kominato, Tiba Prefecture; lectotype designation by 

Okuno & Takeda, 1992] 

Family XIPHoCARIDIDAe ortmann, 1895 

Xiphocaris von Martens, 1872 

= Xiphocaris von Martens, 1872 (type species Hippolyte elongatus Guérin-Méneville, 1855 [in Guérin- 

Méneville, 1855-1856], by monotypy, gender feminine); name placed on the Official List of Generic 

Names in Zoology in Opinion 470 in 1957) 

Fig. 23. Xiphocaris elongata (Guérin-Méneville, 1855). Photo by Chris Lukhaup.


Xiphocaris elongata (Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]) (Fig. 23) 

= Hippolyte elongatus Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xx; Plate 2, Figs 16- 

16a. [costas de Cuba] 

= Oplophorus americanus de Saussure, 1858: 56; Fig. 31. [embouchure des rivières de l’Ile de Haïti] 

= Xiphocaris gladiator Pocock, 1889: 18; Plate 2, fig. 6. [Laiou] 

= Xiphocaris gladiator var. intermedia Pocock, 1889: 19; Plate 2, fig. 7. [Laiou] 

= Xiphocaris brevirostris Pocock, 1889: 20; Plate 2, figs 5-5a. [Dominica] 

= Xiphocaris elongata typica Ortmann, 1895: 400. [Type locality not indicated] 

Xiphocaris gomezi Juarrero de Varona, 1993: 2; Figs 2-4. [Agua Revé, Sierra Maestra, provincia Santiago 

de Cuba, Cuba, altura aproximada de 1000 m snm] 

Superfamily PSAlIDoPoDoIDeA Wood-Mason [in Wood-Mason & Alcock, 1892] 

Family PSAlIDoPoDIDAe Wood-Mason [in Wood-Mason & Alcock, 1892] 

Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892] 

= Psalidopus Wood-Mason [in Wood-Mason & Alcock, 1892] (type species Psalidopus Huxleyi Wood- 

Mason [in Wood-Mason & Alcock, 1892] designated by Holthuis, 1955b, gender masculine; name 

placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Psalidopus barbouri Chace, 1939: 36. [Nicholas Channel, south of Cay Sal Bank, 23°21'N 79°58'W, 300-315 

fms] 

Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892] (Fig. 24) 

= Psalidopus Huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]: 273; Plate 14, figs 1-2, 7. [7.5 

miles E of N Cinque Island, Andaman Sea, 490 fms] 

= Psalidopus spiniventris Wood-Mason [in Wood-Mason & Alcock, 1892]: 274; Plate 14, figs 3-6a, 8; 

Plate 15. [8 miles SE of Cinque Island, Andaman Sea, 500 fms] 

= Psalidopus japonensis Kubo, 1952: 91; Figs 1-2; Plate 5. [off Daiō-zaki, Kii Peninsula, ca. 530 m] 

Psalidopus tosaensis Toriyama & Horikawa, 1993: 1; Figs 1-3, 5. [off Muroto Cape, Kochi Pref., Japan, 

32°13'12''N 134°01'36''E, 2765-2881 m] 

Fig. 24. Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]. Photo by Tin-Yam Chan.


Superfamily STyloDACTyloIDeA Spence Bate, 1888 

Family STyloDACTylIDAe Spence Bate, 1888 

Bathystylodactylus Hanamura & Takeda, 1996 

= Bathystylodactylus Hanamura & Takeda, 1996 (type species Stylodactylus bathyalis Cleva, 1994, by 


Bathystylodactylus bathyalis (Cleva, 1994) 

= Stylodactylus bathyalis Cleva, 1994: 56; Fig. 2. [Bligh Canyon, Coral Sea, 12°23'S 146°08'E. 3515- 

3502 m] 

Bathystylodactylus echinus Wicksten & Martin, 2004: 377; Figs 1-5. [basin off Magdalena Bay, Baja California, 

Mexico, 24°35'N 113°25'W, 3563-3621 m] 

Bathystylodactylus inflatus Hanamura & Takeda, 1996: 930; Figs 1-3. [off east coast of Taiwan, 23°42'03''N 

123°45'08''E, 3436-3452 m] 

Neostylodactylus Hayashi & Miyake, 1968a 

= Neostylodactylus Hayashi & Miyake, 1968a (type species Stylodactylus amarynthis De Man, 1902, by 


Neostylodactylus affinis Hayashi & Miyake, 1968a: 605; Fig. 7. [Korea Strait, 33°34.9'N 128°25.2'E, 120 m] 

Neostylodactylus amarynthis (De Man, 1902) 

= Stylodactylus amarynthis De Man, 1902: 897; Plate 27, figs 64-64b. [Ternate] 

Neostylodactylus investigatoris (Kemp, 1925) 

= Stylodactylus investigatoris Kemp, 1925: 260; Figs 1-2. [2 miles off Great Torres Island, Mergui 40 fms] 

Neostylodactylis litoralis Okuno & Tachikawa, 2000: 39; Figs 1-4. [Takinoura, Ani-jima Island, Ogasawara 

Islands, 5 m] 

Neostylodactylus sibogae (De Man, 1918a) 

= Stylodactylus Sibogae De Man, 1918a: 159. [Siboga Expedition stn 95, 5°43'.5N 119°40'E, Sulu Sea, 

522 m] 

Parastylodactylus Figueira, 1971 

= Parastylodactylus Figueira, 1971 (type species Stylodactylus bimaxillaris Spence Bate, 1888, by original 


Parastylodactylus bimaxillaris (Spence Bate, 1888) 

= Stylodactylus bimaxillaris Spence Bate, 1888: 855; Plate 138, fig. 3. [Challenger stn 219, 1°54'0"S 

146°39'40"E, off the Admiralty Islands, 150 fms] 

Parastylodactylus hayashii (Komai, 1997) 

= Neostylodactylus hayashii Komai, 1997a: 125 (in part); Figs 1-3. [R/V Tansei-maru, KT95-5 cruise, stn 

TB18-2, Okinoyama, off Sunosaki, Boso Peninsula, 34°59'N 139°39'E, 105-113 m] 

Parastylodactylus longidactylus Cleva, 1990: 125; Figs 12d, 15a-b, 16a-d. [Philippines, 13°55.2'N 120°30.5'E, 

320-318 m] 

Parastylodactylus moluccensis Cleva, 1997: 398; Fig. 3. [Indonésie, 05°17'06''S 132°51'19''E, 315-348 m] 

Parastylodactylus richeri Cleva, 1990: 127; Figs 15c, 16e-h. [Nouvelle-Calédonie, 23°45'S 167°12'E, 380 

m] 

Parastylodactylus semblatae Cleva, 1990: 122; Fig. 12c, 14, 18c. [Nouvelle-Calédonie, 22°16.5'S 167°16.5'E, 

475-500 m] 

Parastylodactylus tranterae Cleva, 1990: 119; Figs 11b, 12b, 13. [Nouvelle-Calédonie, 22°11.3'S 167°15.0'E, 

495-550 m] 

Stylodactyloides Cleva, 1990 

= Stylodactyloides Cleva, 1990 (type species Stylodactyloides crosnieri Cleva, 1990, by monotypy, gender 


Stylodactyloides crosnieri Cleva, 1990: 129; Figs 17, 18d-e. [Iles Chesterfield, 19°53.20'S 158°39.50'E, 370- 

400 m]


Stylodactylus A. Milne-edwards, 1881b 

= Stylodactylus A. Milne-Edwards, 1881b (type species Stylodactylus serratus A. Milne-Edwards, 

1881b, by monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology 


Stylodactylus brevidactylus Cleva, 1990: 106; Fig. 8a-g. [Philippines, 13°40.5'N 120°53.7'E, 230-204 m] 

Stylodactylus brucei Cleva, 1994: 54; Fig. 1A, C-F. [south-west Pacific, Wallis Island, 13°37'S 179°56'E, 820- 

840 m] 

Stylodactylus discissipes Spence Bate, 1888: 851; Plate 138, fig. 1. [Challenger stn 171, 28°33'S 177°50'W, 

north of the Kermadec Islands, 600 fms] 

= Stylodactylus orientalis Spence Bate, 1888: 854; Plate 138, fig. 2. [Challenger stn 171, 28°33'S 177°50'W, 

north of the Kermadec Islands, 600 fms] 

Stylodactylus gracilis Cleva, 2008b: 68. [nomen novum for Stylodactylus gracilis Cleva, 2008a] 

= Stylodactylus gracilis Cleva, 2008a: 32; Figs 1, 3A. [Philippine Islands, Panglao Island, 8°49.3'N 

123°1.9'E, 2149-2217 m; not validly published name according to Art. 16.4.2] 

Stylodactylus kauaiensis Figueira, 1971: 3. [Kauai Island, Hawaii, 230-253 fms] 

Stylodactylus laurentae Cleva, 1990: 96; Figs 6, 19. [Nouvelle-Calédonie, 24°55.48'S 168°21.29'E, 500 m] 

Stylodactylus libratus Chace, 1983b: 12; Fig. 5. [Indonesia, Selat Butung, Celebes, 5°29'06"S 122°36'06"E, 

377 m] 

Stylodactylus licinus Chace, 1983b: 14; Fig. 6. [Philippines, Palawan Passage, 10°57'45"N 118°38'15"E, 686 m] 

Stylodactylus macropus Chace, 1983b: 16; Fig. 7. [Philippines, north of Samar, 12°44'42"N 124°59'50"E, 

700 m] 

Stylodactylus major Hayashi & Miyake, 1968a: 590; Figs 2-3. [east China Sea, 30°53.2'N 127°26.0'E – 

30°44.0'N 127°29.0'E, 122-124 m] 

Stylodactylus multidentatus multidentatus Kubo, 1942b (Fig. 25) 

= Stylodactylus multidentatus Kubo, 1942b: 34; Figs 4-5. [Kumanonada, off Mie Prefecture, about 300 m] 

= Stylodactylus brevidactylus Cleva, 1990: 106; Fig. 8a-g. [Philippines, 13°40.5'N 120°53.7'E, 230-204 m] 

Fig. 25. Stylodactylus multidentatus multidentatus Kubo, 1942. Photo by Tin-Yam Chan.


Stylodactylus multidentatus robustus Cleva, 1990: 105; Fig. 8n-s. [Madagascar, 22°20.5'S 43°06.1'E, 400 m] 

Stylodactylus profundus Cleva, 1990: 85; Fig. 2. [Nouvelle-Calédonie, 24°19'S 167°08'E, 1530-1480 m] 

Stylodactylus pubescens Burukovsky, 1990: 198; Fig. 3B. [25°09'S 96°18'W, 545-800 m] 

Stylodactylus rectirostris A. Milne-Edwards, 1883: Plate 35. [off Dominica, 15°18'12''N 61°26'32''W, 542 

fms] 

Stylodactylus serratus A. Milne-Edwards, 1881b: 11. [près de la Dominique, 524 brasses] 

Stylodactylus stebbingi Hayashi & Miyake, 1968a: 595; Fig. 4. [Off Buffalo River, NW ½ W, 19 miles, East 

London, Cape Colony, 300 fms] 

Stylodactylus tokarensis Zarenkov, 1968a: 58; Figs 2-3. [Vitjaz stn 3768, Tokara Strait, East China Sea, 820 

m] 

Superfamily CAMPyloNoToIDeA Sollaud, 1913 

Family BATHyPAlAeMoNellIDAe De Saint laurent, 1985 

Bathypalaemonella Balss, 1914a 

= Bathypalaemonella Balss, 1914a (type species Bathypalaemonella zimmeri Balss, 1914a, by monotypy, 


Bathypalaemonella adenensis Cleva, 2001: 770; Fig. 6. [Golfe d’Aden, 11°56.5'N 43°34.5'E, 1600-1400 m] 

Bathypalaemonella delsolari Wicksten & Méndez, 1983: 225; Figs 1-4. [SW of Lobos de Tierra, Peru, 6°31'S 

81°01'W, 712-714 m] 

Bathypalaemonella hayashii Komai, 1995a: 40; Figs 1-4. [southeast of Amami-oshima Island, 28°02.5'N 

129°34.5'E / 28°02.8'N 129°34.9'E, 815-789 m] 

Bathypalaemonella humilis Bruce, 1966b: 277; Figs 1-3. [15°55.7'N 109°28.5'E to 15°57.9'N 109°31.5'E, 172- 

142 fms] 

Bathypalaemonella pandaloides (Rathbun, 1906) 

= Palaemon pandaloides Rathbun, 1906: 924; Fig. 73; Plate 22, fig. 4. [vicinity of Kauai Island, 528 fms] 

Bathypalaemonella serratipalma Pequegnat, 1970: 77; Figs 4.5-4.6. [Southwestern Gulf of Mexico, 21°44'N 

96°46'W, 1774 m] (Fig. 26) 

Bathypalaemonella texana Pequegnat, 1970: 81; Figs 4.7-4.8. [Northwestern Gulf of Mexico, 27°01.6'N 

94°42'W, 1463 m] 

Bathypalaemonella zimmeri Balss, 1914a: 598. [6°18'N 49°32'E, 1079 m] 

Fig. 26. Bathypalaemonella serratipalma Pequegnat, 1970. Photo Gilliss Expedition stn G-1111.


Bathypalaemonetes Cleva, 2001 

= Bathypalaemonetes Cleva, 2001 (type species Bathypalaemonella brevirostris Bruce, 1986a, by original 


Bathypalaemonetes brevirostris (Bruce, 1986a) 

= Bathypalaemonella brevirostris Bruce, 1986a: 252; Figs 1-5. [13°52.1'S 123°00.4'E, 306-308 m] 

Bathypalaemonetes chani Cleva, 2004: 508; Figs 4-5, 6h. [22°0.98'N 120°6.73'E, 690-700 m] 

Bathypalaemonetes pilosipes (Bruce, 1986a) 

= Bathypalaemonella pilosipes Bruce, 1986a: 257; Figs 6-10. [13°33.8'S 122°53.4'E, 390-394 m] 

Family CAMPyloNoTIDAe Sollaud, 1913 

Campylonotus Spence Bate, 1888 

= Campylonotus Spence Bate, 1888 (type species Campylonotus semistriatus Spence Bate, 1888, designated 

by Holthuis, 1955b, gender masculine; name placed on the Official List of Generic Names in 


= Anchistiella A. Milne-Edwards, 1891 (type species Anchistiella Hyadesi A. Milne-Edwards, 1891 

(junior subjective synonym of Campylonotus vagans Spence Bate, 1888), designated by Holthuis, 

1955b, gender feminine) 

Campylonotus arntzianus Thatje, 2003: 243; Figs 2-4. [Saunders Island, South Sandwich Islands, Scotia 

Sea, Antarctica, 57°40.31'S 26°27.81'W, 475-589 m] 

Campylonotus capensis Spence Bate, 1888: 773; Plate 128, fig. 3. [Challenger stns 145, 46°43'S 38°4'30''E, off 

Marion Island, 140 fms; 122, 9°5'S 34°50'W, off Pernambuco, 350 ms] 

Campylonotus rathbunae Schmitt, 1926a 

= Campylonotus rathbunæ Schmitt, 1926a: 373; Plate 67, figs 1-5. [South of Eucla, Great Australian 

Bight, 129°28'E, 250-450 fms] 

Fig. 27. Campylonotus vagans Spence Bate, 1888. Photo by Karen Neely.


Campylonotus semistriatus Spence Bate, 1888: 768; Plate 128, figs 1-2. [Challenger stns 309, 50°56'S 74°15'W, 

Puerto Bueno, Patagonia, 40 fms; 305A, 47°48'30''S 74°47'0''W, Messier Channel, 125 fms; 306A, 

48°27'S 74°30'W, Messier Channel, 345 fms; 307, 49°24'30''S 74°23'30''W, off Port Grappler, 140 fms; 

308, 50°8'30''S 74°41'0''W, off Tom Bay, 175 fms; 310, 51°27'30''S 74°3'0''W, Sarmiento Channel, 400 

fms; 311, 52°45'30''S 73°46'0''W, off Port Churruca, 245 fms] 

Campylonotus vagans Spence Bate, 1888: 775; Plate 122, fig. 3. [Challenger stn 308, 50°8'30''S 74°41'0''W, off 

Tom Bay, Patagonia, 175 fms] (Fig. 27) 

= Anchistiella Hyadesi A. Milne-Edwards, 1891: 38; Plate 4, figs 1-1n. [à l’entrée est du canal de Franklin, 

entre Herschel et Wollaston, 51 m; dans la baie Orange, 26-28 m; dans la baie du Naturaliste, 

35 m; dans le détroit de Magellan, 18 m; à l’île de Wollaston, 95 m] 

= Anchistiella Hahni A. Milne-Edwards, 1891: 41; Plate 4, figs 2-2d. [à l’île Wollaston dans la baie 

Gretton, à Lajaifwaya, 30 m; aux Murray narrows, 200 m; à 10 milles au sud-est de la baie Seogatt, 

816 m] 

= Anchistiella Seneuilli A. Milne-Edwards, 1891: 42; Plate 3, figs 2-2d. [dans le détroit de Magellan, 

en face de Punta Arenas, 143 m] 

Superfamily PALAEMONOIDEA Rafinesque, 1815 

Family ANCHISTIoIDIDAe Borradaile, 1915b 

Anchistioides Paul’son, 1875 

= Anchistioides Paul’son, 1875 (type species Anchistioides compressus Paul’son, 1875, by monotypy, 

gender masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

= Palaemonopsis Borradaile, 1900 (type species Palaemonopsis willeyi Borradaile, 1900, by monotypy, 


= Amphipalaemon Nobili, 1901a (nomen novum for Palaemonopsis Borradaile, 1900, gender masculine) 

Anchistioides antiguensis (Schmitt, 1924b) (Fig. 28) 

= Periclimenes antiguensis Schmitt, 1924b: 84. [English Harbour, Antigua, electric light] 

= Periclimenes barbadensis Schmitt, 1924b: Plates 3-4. [English Harbour, Antigua, electric light] 

Anchistioides compressus Paul’son, 1875 

= Anch.[istioides] compressus Paul’son, 1875: 115; Plate 19, figs 5-5n. [Red Sea] 

= Amphipalaemon Seurati Nobili, 1906b: 259. [Tearia, Tuamotu Islands according to Bruce, 1967a] 

Fig. 28. Anchistioides antiguensis (Schmitt, 1924). Photo by Arthur Anker.


Anchistioides willeyi (Borradaile, 1900) 

= Palaemonopsis willeyi Borradaile, 1900: 410; Plate 36, figs 7-7b; Plate 37, figs 7c-e. [Ralun, New 

Britain] 

= Amphipalaemon cooperi Borradaile, 1915b: 209. [S. Nilandu Atoll, Maldive Islands] 

= Amphipalaemon gardineri Borradaile, 1915b: 209. [N. Malé Atoll, Maldive Islands] 

= Amphipalaemon australiensis Balss, 1921a: 11; Figs 3-6. [Cape Jaubert, 45 meilen WSW, 66 Fuss Tiefe] 

Family DeSMoCARIDIDAe Borradaile, 1915b 

Desmocaris Sollaud, 1911a 

= Desmocaris Sollaud, 1911a (type species Palaemonetes trispinosus Aurivillius, 1898a, by monotypy, 


1957) 

Desmocaris trispinosa (Aurivillius, 1898a) (Fig. 29) 

= Palaemonetes trispinosus Aurivillius, 1898a: 29; Plate 4, figs 1-2. [Kamerun, Kitta, in einem Bache] 

Desmocaris bislineata Powell, 1977: 667; Fig. 5. [River Ethiope at Adagbrassa-Amukpe, 5°51'N 5°45.5'E] 

Family euRyRHyNCHIDAe Holthuis, 1950a 

Euryrhynchina Powell, 1976 

= Euryrhynchina Powell, 1976 (type species Euryrhynchina edingtonae Powell, 1976, by monotypy, 


Euryrhynchina edingtonae Powell, 1976: 895; Figs 5-7. [swamp forest along Ogba River, 1 km south of 

Ogba village and approx. 8 km southwest of Benin City centre, 6°16.5'N 5°35'E] 

Euryrhynchoides Powell, 1976 

= Euryrhynchoides Powell, 1976 (type species Euryrhynchoides holthuisi Powell, 1976, by monotypy, 


Euryrhynchoides holthuisi Powell, 1976: 886; Figs 1-4. [Sierra Leone: River Taja (= River Jong) at Njala, 

8°06'N 12°04'W] 

Euryrhynchus Miers, 1877 

= Euryrhynchus Miers, 1877 (type species Euryrhynchus wrzesniowskii Miers, 1877, by monotypy, gender 

masculine; placed on the Official List of Generic Names in Zoology in Opinion 518 in 1958) 

= Euryrhynchella Balss, 1955 (nomen novum for Euryrhynchus Miers, 1877, gender feminine) 

Euryrhynchus amazoniensis Tiefenbacher, 1978: 183; Fig. 2a-b. [A15-1, Rio Tupana, am 2. Vermessungspunkt] 

Euryrhynchus burchelli Calman, 1907: 297; Figs 2-8. [Pará] 

Euryrhynchus pemoni Pereira, 1985: 619; Fig. 5. [Creek in La Gran Sabana road, between Sn. Rafael Town 

and Kama Fall, Venezuela, 6°15'N 60°1'W] 

Euryrhynchus tomasi De Grave, 2007a: 194; Figs 1-7. [Crique Timothy, near the N2 road from Règina to St 

Georges de l’Oyapock, 20 km NW of St Georges de l’Oyapock, French Guyana, approx. 4°00'N 

51°52'W] (Fig. 30) 

Euryrhynchus wrzesniowskii Miers, 1877: 662; Plate 67, figs 2-2b. [Cayenne] 

Family GNATHoPHyllIDAe Dana, 1852a 

Gnathophylleptum d’udekem d’Acoz, 2001 

= Gnathophylleptum d’Udekem d’Acoz, 2001 (type species Gnathophylleptum tellei d’Udekem d’Acoz, 

2001, by monotypy, gender neuter) 

Gnathophylleptum tellei d’Udekem d’Acoz, 2001: 114; Figs 1-7. [Canary Islands, Gran Canaria, Sardina del 

Norte, Caulerpa racemosa meadow, 15 m]


Fig. 29. Desmocaris trispinosa (Aurivillius, 1898). Photo by Chris Lukhaup. 

Fig. 30. Euryrhynchus tomasi De Grave, 2007. Photo by J. Tomas & O. Helker.


Gnathophylloides Schmitt, 1933 

= Gnathophylloides Schmitt, 1933 (type species Gnathophylloides mineri Schmitt, 1933, by monotypy, 


Gnathophylloides mineri Schmitt, 1933: 7; Fig. 3. [Ballena Point, Ensenada, Puerto Rico] 

Gnathophylloides robustus Bruce, 1973a: 17; Figs 1-7. [off Point Moore, Geraldton, Western Australia, 

28°47.0'S 114°34.2'E, 3 m] 

Gnathophyllum latreille, 1819 

= Gnatophyllum Latreille, 1819 (type species A.[lpheus] elegans Risso, 1816, designated by H. Milne 

Edwards, 1837 [in H. Milne Edwards, 1836-1844], gender neuter; name emended under the plenary 

power of the ICZN to Gnathophyllum and placed in that emended spelling on the Official List of 


= Drimo Risso, 1827 (type species A.[lpheus] elegans Risso, 1816, gender masculine; name placed on 

the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 1957) 

Gnathophyllum americanum Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xx; Plate 2, fig. 14. 

[Cuba] 

= Gnathophyllum fasciolatum Stimpson, 1860a: 28. [Rose Bay, Port Jackson, NSW; neotype designation 

by Ahyong, 2003] 

= Gnathophyllum zebra Richters, 1880: 161; Plate 17, Figs 18-20. [Fouquets] 

= Gnathophyllum pallidum Ortmann, 1890: 537. [Tahiti] 

= Gnathophyllum tridens Nobili, 1906b: 259. [Rikitea] 

= Gnathophyllum minuscularium Armstrong, 1940: 9; Fig. 4C-K. [in the Reach, St. George Island, Bermuda] 

Gnathophyllum ascensione Manning & Chace, 1990: 11; Figs 5-6. [Northeast Bay, Ascension Island] 

Gnathophyllum circellum Manning, 1963: 54; Figs 3-4. [300 yards southwest of Alligator Reef Light, Monroe 

Co., Florida, rock ledge at 15-20 feet] 

Gnathophyllum elegans (Risso, 1816) 

= Peneus quadricolor Rafinesque, 1814: 22. [Type locality not indicated; suppressed under the plenary 

power for the purposes of the Principle of Priority but not for those of the Principle of Homonymy 

in Opinion 522 in 1958] 

= A.[lpheus] elegans Risso, 1816: 92; Plate 2, fig. 4. [environs de Nice, dans les profondeurs rocailleuses] 

= Gnathophyllum elegans var. brevirostris Gourret, 1887a: 1034. [l’estaque jusqu’à la Corbière; près de 

Somaty; Mourepiano; Roucas blanc] 

Gnathophyllum modestum Hay, 1917: 72. [about 20 miles S.W. of Beaufort, N.C., in about 15 fms] 

Gnathophyllum panamense Faxon, 1893: 198. [Panama] (Fig. 31) 

Gnathophyllum precipuum Titgen, 1989: 203; Figs 1-3. [Oahu, Mahaka, 12 m] 

Gnathophyllum splendens Chace & Fuller, 1971: 493; Figs 1-5. [Puerto Yabucoa, one-half mile east of Playa 

de Guayanés, Municipio de Yabucoa, Puerto Rico] 

Gnathophyllum taylori Ahyong, 2003: 237; Figs 1-2. [S side of Long Bay, Malabar, New South Wales, 

33°58'S 151°15'E, 4 m, under Centrostephanus rogersi] 

Levicaris Bruce, 1973a 

= Levicaris Bruce, 1973a (type species Coralliocaris mammilatus Edmondson, 1931, by monotypy, gender 

feminine) 

Levicaris mammillata (Edmondson, 1931) 

= Coralliocaris mammillata Edmondson, 1931: 5; Fig. 2; Plate 1. [Waikiki Reef, Oahu, associated with 

echinoid Heterocentrotus mamillatus] 

Pycnocaris Bruce, 1972a 

= Pycnocaris Bruce, 1972a (type species Pycnocaris chagoae Bruce, 1972a, by monotypy, gender feminine) 

Pycnocaris chagoae Bruce, 1972a: 54; Figs 1-7. [East Point, Diego Garcia, Chagos Archipelago] (Fig. 32)


Fig. 31. Gnathophyllum panamense Faxon, 1893. Photo by Arthur Anker. 

Fig. 32. Pycnocaris chagoae Bruce, 1972. Photo by Tin-Yam Chan.


Family HyMeNoCeRIDAe ortmann, 1890 

Hymenocera latreille, 1819 

= Hymenocera Latreille, 1819 (type species Hymenocera picta Dana, 1852b, designated under the plenary 

power of the ICZN, gender feminine; name placed on the Official List of Generic Names in 


= Nematophyllum Bleeker, 1856 (type species Hymenocera picta Dana, 1852b, designated by Holthuis, 

1955b, gender neuter; invalid junior hononym of Nematophyllum H. Milne Edwards & Haime, 

1850 (Coelenterata); name placed on the Official List of Rejected and Invalid Names in Zoology in 


Hymenocera picta Dana, 1852b: 593 (1852); Plate 39, figs 3a-c (1855). [coral reefs of Raraka, Paumotu Islands] 

(Fig. 33) 

= H.[ymenocera] elegans Heller, 1861: 25. [Zu Tor zwischen Korallen] 

= H.[ymenocera] elegans Heller, 1862c: 264; Plate 3, figs 9-14. [Tor] 

= Hymenocera latreillii Sharp, 1893: 119 [Agalega, Seychelles] 

Phyllognathia Borradaile, 1915b 

= Phyllognathia Borradaile, 1915b (type species Hymenocera (?) ceratophthalma Balss, 1913b, by monotypy, 

gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

470 in 1957) 

Phyllognathia ceratophthalma (Balss, 1913b) (Fig. 34) 

= Hymenocera (?) ceratophthalma Balss, 1913b: 236. [Satsuma] 

Phyllognathia simplex Fujino, 1973a: 90; Figs 1-3. [near Shuragane, Sagami Bay, central Japan, 40-50 m deep] 

Fig. 33. Hymenocera picta Dana, 1852. Photo by Arthur Anker.


Fig. 34. Phyllognathia ceratophthalma (Balss, 1913). Photo by Tin-Yam Chan. 

Family KAKADuCARIDIDAe Bruce, 1993a 

Calathaemon Bruce & Short, 1993 

= Calathaemon Bruce & Short, 1993 (type species Palaemonetes holthuisi Strenth, 1976, by monotypy, 


Calathaemon holthuisi (Strenth, 1976) 

= Palaemonetes holthuisi Strenth, 1976: 3, Fig. 1. [subterranean waters of Ezell’s Cave in the city of San 

Marcos, Hays County, Texas] 

Kakaducaris Bruce, 1993a 

= Kakaducaris Bruce, 1993a (type species Kakaducaris glabra Bruce, 1993a, by monotypy, gender feminine) 

Kakaducaris glabra Bruce, 1993a: 28; Figs 1-11, 12A. [Lightening Dreaming Creek, Nourlangi, Kakadu 

National Park, Arnhem Land, Northern Territory, 12°55.3'S 132°55.8'E, 90 m alt.] (Fig. 35) 

Leptopalaemon Bruce & Short, 1993 

= Leptopalaemon Bruce & Short, 1993 (type species Leptopalaemon gagadjui Bruce & Short, 1993, by 


Leptopalaemon gagadjui Bruce & Short, 1993: 75; Figs 1-6. [Barramundi Creek, Kakadu National Park, 

Northern Territory, 13°20'S 132°27'E] 

Family PALAEMONIDAE Rafinesque, 1815 

Subfamily PALAEMONINAE Rafinesque, 1815 

Arachnochium Wowor & Ng, 2010a 

= Arachnochium Wowor & Ng, 2010a (type species Palaemon mirabilis Kemp, 1917c, by original designation, 


Arachnochium kulsiense (Jayachandran, Lal Mohan & Raji, 2007) 

= Macrobrachium kulsiense Jayachandran, Lal Mohan & Raji, 2007: 43; Figs 1-3. [Kulsi River] 

Arachnochium mirabilis (Kemp, 1917c) 

= Palaemon mirabilis Kemp, 1917c: 227; Plate 10. [Pazudaung and Dala Creeks, Rangoon]


Fig. 35. Kakaducaris glabra Bruce, 1993. Photo by A.J. Bruce. 

Brachycarpus Spence Bate, 1888 

= Brachycarpus Spence Bate, 1888 (type species Brachycarpus savignyi Spence Bate, 1888 (junior subjective 

synonym of Brachycarpus biunguiculatus Lucas, 1846), by original designation, gender masculine; 


= Retrocaris Ortmann, 1893 (type species Retrocaris spinosa (junior subjective synonym of Palæmon 

biunguiculatus Lucas, 1846), designated by Holthuis, 1955b, gender feminine) 

= Calmania Nobili, 1907 (type species Palæmon biunguiculatus Lucas, 1846, by original designation 

and monotypy, gender feminine; invalid junior homonym of Calmania Laurie, 1906 (Crustacea 

Brachyura)) 

Brachycarpus biunguiculatus (Lucas, 1846) 

= Palæmon biunguiculatus Lucas, 1846: 45; Plate 4, figs 4-4a. [Oran et Bône] 

= Brachycarpus savignyi Spence Bate, 1888: 795; Plate 124, fig. 4. [Bermuda] 

= Brachycarpus neapolitanus Cano, 1890: 38; Plate 4, fig. 1. [Golfo di Napoli] 

= Retrocaris spinosa Ortmann, 1893: 84; Plate 6, fig. 2. [Plankton-Expedition der Humboldt-Stiftung, 

Sargasso-See JN 62 (33.2°N 63.8°W, 0-400 m) and Pl. 34 (32.1°N 63.4°W, 200 m); Südl. Aequatorialstrom 

JN 249 (5.6°S 44.0°W, 0 m)] 

= Brachycarpus advena Nobili, 1905b: 395. [Mer Rouge] 

= Palæmonella rathbunensis Borradaile, 1917: 358. [Hawaiian Is.] 

Brachycarpus crosnieri Bruce, 1998a: 158; Figs 1-4. [New Caledonia, Loyalty Islands, Uvea, Passe de la 

Meurthe, 6-10 m] 

Brachycarpus holthuisi Fausto Filho, 1966: 123; Figs 1-11. [Acaraú, Ceará] 

Coutierella Sollaud, 1914 

= Coutierella Sollaud, 1914 (type species Coutierella tonkinensis Sollaud, 1914, by monotypy, gender 

feminine) 

Coutierella tonkinensis guangdongensis (Liu, Liang & Yan, 1990a) 

= Palaemonetes tonkinensis guangdongensis Liu, Liang & Yan, 1990a: 250; Fig. 41. [Guangzhou]


Coutierella tonkinensis tonkinensis Sollaud, 1914 

= Coutierella tonkinensis Sollaud, 1914: 318; Figs 2A, 3B, 4. [delta du fleuve Rouge, Tonkin] 

Creaseria Holthuis, 1950a 

= Creaseria Holthuis, 1950a (type species Palæmon morleyi Creaser, 1936, by original designation, 


Creaseria morleyi (Creaser, 1936) 

= Palæmon morleyi Creaser, 1936: 126; Figs 25-30. [San Isidro Cave, Salar Colony, Merida, Yucatan] 

Cryphiops Dana, 1852a 

= Cryphiops Dana, 1852a (type species Cryphiops spinuloso-manus Dana, 1852a (junior subjective synonym 

of Cancer caementarius Molina, 1782), by monotypy, gender masculine) 

= Bithynis Philippi, 1860 (type species Bithynis longimana Philippi, 1860 (junior subjective synonym 

of Cancer caementarius Molina, 1782), by monotypy, gender feminine) 

= Bithynops Holthuis, 1973b (type species Bythinops luscus Holthuis, 1973b, by original designation 


Cryphiops (Bithynops) brasiliensis (Gomes Corrêa, 1973) 

= Cryphiops brasiliensis Gomes Corrêa, 1973: 169; Figs 1-21. [riacho da Granja do Ipê, em Brasília, 

distrito Federal] 

Cryphiops (Bithynops) luscus (Holthuis, 1973b) 

= Bythinops luscus Holthuis, 1973b: 136; Figs 1-2. [Grutas de l’Arco, near San Raphael de l’Arco, La 

Trinitaria, about 16°10'N 92°01'W, Chiapas State, Mexico, altitude 1470 m] 

Cryphiops (Bithynops) perspicax (Holthuis, 1977b) 

= Bythinops perspicax Holthuis, 1977b: 182; Figs 3-4. [Cenote la Cueva, Lagunas de Montebello, La 

Trinitaria, Chiapas, altitude 1480 m] 

Cryphiops (Bithynops) sbordonii Baldari, Mejía-Ortiz & López-Mejía, 2010: 48; Figs 2-4. [Cueva Chamburro, 

Las Margaritas, Chiapas, Mexico, 16°25'57''N 91°56'40''W] 

Cryphiops (Bithynops) villalobosi Villalobos Hiriart, Nates Rodriguez & Cantú Diaz Barriga, 1989: 166; 

Figs 1-5, 7b-d, 8c. [arroyo La Laja, km. 140 carretera Palenque-Ocosingo, a 5 km al NO del poblado 

de Ocosingo. Mpio. de Ocosingo, Chiapas] 

Cryphiops (Cryphiops) caementarius (Molina, 1782) (Fig. 36) 

= Cancer cæmentarius Molina, 1782: 208. [Chili] 

= P.[alémon] Gaudichaudii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 400. [Chili] 

= Cryphiops spinuloso-manus Dana, 1852a: 26. [in fluminibus Chilensibus mari remotis] 

= Bithynis longimana Philippi, 1860: 164. [im Flusse la Ligua, Chile] 

= Macrobrachium africanum Spence Bate, 1868a: 366; Plate 31, fig. 3. [Tambo river; according to Semper, 

1868 on the west coast of South America, near Islay in Peru] 

Exopalaemon Holthuis, 1950a 

= Exopalaemon Holthuis, 1950a (type species P.[alemon] styliferus H. Milne Edwards, 1840, by original 


Exopalaemon annandalei (Kemp, 1917c) 

= Leander annandalei Kemp, 1917c: 211; Fig. 1-4. [China, Whangpoo River, between Shanghai and 

Woosung, 5.5-7.5 m] 

= Leander annandalei var. stylirostris Yu, 1930a: 460; Fig. 4C-P. [Tangku] 

Exopalaemon carinicauda (Holthuis, 1950a) 

= Palaemon (Exopalaemon) carinicauda Holthuis, 1950a: 48; Fig. 9. [nomen novum for Leander longirostris 

var. carinatus Ortmann, 1890] 

= Leander longirostris var. carinatus Ortmann, 1890: 521. [China] 

Exopalaemon guangdongensis Guo, Wang & Zhang, 2005: 840; Figs 1-2. [Zhujiang river, near Tangjiawan, 

Zhuhai City, circa 22°36'N 113°36'E] 

Exopalaemon hainanensis Liang, 2000: 278; Figs 11-19. [fishery market, Haikou, Hainan Island] 

Exopalaemon mani (Sollaud, 1914)


Fig. 36. Cryphiops (Cryphiops) caementarius (Molina, 1782). Photo by Arthur Anker. 

= Leander Mani Sollaud, 1914: 315; Figs 1, 2B. [delta du fleuve Rouge, Tonkin] 

Exopalaemon modestus (Heller, 1862a) 

= L.[eander] modestus Heller, 1862a: 527. [Schanghai] 

= Leander modestus Heller, 1865: 111; Plate 10, fig. 6. [Shanghai] 

= Leander czerniavskyi Bražnikov, 1907: 176. [Amurskiy Liman] 

= Leander modestus var. sibirica Czerniavsky, 1878; 24. [nomen nudum] 

= Leander macrogenitus Yu, 1930b: 559; Figs 1A-C. [Hangchow] 

= Leander czerniavsky lacustris Buldovsky, 1933: 50; Plate 2, figs 21, 23. [Lake Khanka, Soviet Far East] 

Exopalaemon orientis (Holthuis, 1950a) 

= Palaemon (Exopalaemon) orientis Holthuis, 1950a: 49. [nomen novum for Leander longirostris var. 

japonicus Ortmann, 1890] 

= Leander longirostris var. japonicus Ortmann, 1890: 519; Plate 37, figs 14, 14z. [Japan, Tokiobai] 

Exopalaemon styliferus (H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840]) 

= P.[alemon] styliferus H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840]: 638. [nomen novum 

for P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 394 nec 

P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 392] 

= P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 394. [l’embouchure 

du Gange] 

Exopalaemon vietnamicus Nguyên, 1992: 25; Figs 4, 12 (part). [Can Giò, 15 km west of Vu Tà] 

Exopalaemon xinjiangensis Liang, 2000: 277; Figs 1-10. [Talimu River, Xinjiang, northwest China] 

Leander Desmarest, 1849 

= Leander Desmarest, 1849 (type species Leander erraticus Desmarest, 1849 (junior subjective synonym 

of P.[alaemon] tenuicornis Say, 1818), by original designation and monotypy, gender masculine; 


= Cryptoleander Gurney & Lebour, 1941 (type species P.[alaemon] tenuicornis Say, 1818, by monotypy, 


Leander kempi Holthuis, 1950a: 31; Fig. 3. [Siboga stn 121, Menado anchorage, shore exploration, 55 m; 

Snellius Expedition, Beo, Talaud Islands]


Leander manningi Bruce, 2002a: 71; Figs 1-4. [BP Oil Refinery Jetty, Kwinana, Cockburn Sound, Western 

Australia] 

Leander paulensis Ortmann, 1897: 192; Plate 1, fig. 14. [canal entre o continente e a ilha de São Sebastião, 

Estado de São Paulo] 

Leander plumosus Bruce, 1994a: 39; Figs 1-5, 6A-B; Plate 2. [Ari Atoll, Maldive Islands] 

Leander tenuicornis (Say, 1818) 

= P.[alæmon] tenuicornis Say, 1818: 249. [Banks of Newfoundland] 

= Astacus locusta Fabricius, 1781: 513; nec Astacus locusta Pennant, 1777. [Type locality not indicated] 

? = Penaeus punctatissimus Bosc, 1802: 109; Plate 14, fig. 3. [dans sa traversée d’Europe en Amérique, 

sur les fucus nageans] 

? = Penaeus adspersus Tilesius, 1819: 4; Plate 21, fig. 1. [suppressed under the Plenary Power for the 

Purpose of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 564 

in 1959] 

= P.[alemon] natator H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 393. [dans l’océan 

Indien, sur du fucus nageans] 

= Palaemon latirostris De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 12 (1844); 170, Plate P 

(1849). [Japan; lectotype designation by Yamaguchi & Baba, 1993] 

= Leander erraticus Desmarest, 1849: 92; unnumbered figure. [Océan Atlantique, à cent ou cent cinquante 

lieues des côtes de la Guadeloupe] 

= P.[alaemon] torensis Paul’son, 1875: 116; Plate 17, fig. 3. [Red Sea] 

= P.[alaemon] torensis var. A Paul’son, 1875: 116; Plate 17, fig. 4. [Red Sea] 

= P.[alaemon] torensis var. B Paul’son, 1875: 116; Plate 17, fig. 5. [Red Sea] 

? = Palaemon mauritii Spamer & Bogan, 1992: 163. [nomen nudum] 

? = Palaemon mexicanus Spamer & Bogan, 1992: 163. [nomen nudum] 

Leandrites Holthuis, 1950a 

= Leandrites Holthuis, 1950a (type species Leander celebensis De Man, 1881 by original designation, 


Leandrites celebensis (De Man, 1881) 

= Leander celebensis De Man, 1881: 141. [on the road of Macassar (Celebes)] 

= Palaemonetes hornelli Kemp, 1925: 318; Figs 14-15. [Silavathuri Lagoon, Tuticorin, S. India] 

= Leander wieneckii Holthuis, 1950a: 36. [nomen nudum, manuscript name for Leander celebensis De 

Man, 1881] 

Leandrites deschampsi (Nobili, 1903c) 

= Leander Deschampsi Nobili, 1903c: 8. [Singapore] 

Leandrites indicus Holthuis, 1950a: 37; Fig. 5. [Off Makassar] 

Leandrites stenopus Holthuis, 1950a: 40; Fig. 6. [Madoera Strait, 7°25'S 113°16'E, 56 m] 

Leptocarpus Holthuis, 1950a 

= Leptocarpus Holthuis, 1950a (type species Leander fluminicola Kemp, 1917c, by original designation, 


Leptocarpus fluminicola (Kemp, 1917c) 

= Leander fluminicola Kemp, 1917c: 223; Plate 9, fig. 2. [River Ganges, Mirzapur, United Provinces] 

Leptocarpus kempi Jayachandran, 1992: 129; Fig. 2. [Cochin backwaters at Panangad] 

Leptocarpus potamiscus (Kemp, 1917c) 

= Leander potamiscus Kemp, 1917c: 225; Fig. 7. [Patani River, below the town of Patani, Siamese Malay 

States] 

Macrobrachium Spence Bate, 1868a 

= Macrobrachium Spence Bate, 1868a (type species Macrobrachium americanum Spence Bate, 1868a, 

designated by Fowler, 1912, gender neuter; name placed on the Official List of Generic Names in 

Zoology in Opinion 564 in 1959)


= Eupalaemon Ortmann, 1891 (type species Palaemon (s.s.) dolichodactylus Hilgendorf, 1879 (junior 

subjective synonym of Palaemon scabriculus Heller, 1862a), designated by Holthuis, 1955b, gender 


= Parapalaemon Ortmann, 1891 (type species Palaemon (s.s.) dolichodactylus Hilgendorf, 1879 (junior 

subjective synonym of Palaemon scabriculus Heller, 1862a, designated by Holthuis, 1955b, gender 


= Macroterocheir Stebbing, 1908 (type species Palaemon (s.s.) lepidactylus Hilgendorf, 1879, by monotypy, 


Macrobrachium acanthochirus Villalobos F., 1967: 168; Plates 1-2. [Río Valdeflores, Valdeflores de Tonameca, 

Pochutla, Estado de Oaxaca, Mexico] 

Macrobrachium acanthurus (Wiegmann, 1836) 

= P.[alaemon] acanthurus Wiegmann, 1836: 150. [Küste Brasiliens] 

= P.[alemon] forceps H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 397. [Rio-Janeiro] 

= Pal.[æmon] Swainsonii White, 1847a: 78. [nomen nudum] 

= Palæmon mexicanus de Saussure, 1857a: 504. [Cuba et Mexique] 

= Macrobrachium longidigitum Spence Bate, 1868a: 365; Plate 31, fig. 2. [Type locality not indicated] 

= Palæmon dasydactylus Streets, 1871b: 225; Plate 2, Figs 3-3a. [tide-water of the Coatzacoalcos River, 

Isthmus of Tehuantepec] 

= Palæmon sexdentatus Streets, 1871b: 226; Plate 2, Figs 4-4a. [tide-water of the Coatzacoalcos River, 

Isthmus of Tehuantepec] 

? = Palaemon Potieté Müller, 1892: 184. [nomen nudum] 

Macrobrachium acherontium Holthuis, 1977b: 188; Figs 6-7. [Grutas del Cocona, 3 km NE of Teapa, Tabasco, 

altitude 60 m above sea level] 

= Macrobrachium coconaensis Guzman, Cabrera & Kensler, 1977: 208. [nomen nudum] 

Macrobrachium aemulum (Nobili, 1906b) 

= Palaemon (Parapalaemon) aemulus Nobili, 1906b: 258. [Gatavake, 180 mètres d’altitude] 

Macrobrachium agwi Klotz, 2008: 49; Figs 1-3. [India, West Bengal, Alipurduar District, Barobisha, approx. 

800 km north of Calcutta] 

Macrobrachium ahkowi Chong & Khoo, 1987a: 562. [nomen novum for Macrobrachium johnsoni Chong & 

Khoo, 1987b] 

= Macrobrachium johnsoni Chong & Khoo, 1987b: 360; Figs 1-3; nec Macrobrachium johnsoni Ravindranath, 

1979. [first landing, Gunong Palai waterfall stream, Johore, ca. 200 m altitude] 

Macrobrachium altifrons altifrons (Henderson, 1893) 

= Palæmon altifrons Henderson, 1893: 444; Plate 40, Figs 4-6. [Delhi; River Jumna; Lahore] 

Macrobrachium altifrons ranjhai Tiwari, 1964a 

= Macrobrachium altifrons ranjhai Tiwari, 1964a: 237; Figs 2A-F, 5-6. [Kabul River at Nowshera, Peshawar 

District (West Pakistan)] 

Macrobrachium amazonicum (Heller, 1862b) 

= P.[alaemon] amazonicus Heller, 1862b: 418; Plate 2, fig. 45. [im Amazonenstrome] 

= Palæmon ensiculus Smith, 1869: 26; Plate 1, figs 2-2c. [Pará, Brazil] 

= Palaemon Dieperinkii De Man, 1879: 167. [Suriname] 

Macrobrachium americanum Spence Bate, 1868a: 363; Plate 30. [Lake of Amatitlan, Guatemala] 

Macrobrachium amplimanus Cai & Dai, 1999: 231; Figs 13-15. [forest stream near Mengla County] 

Macrobrachium andamanicum (Tiwari, 1952) 

= Palaemon andamanicum Tiwari, 1952: 30. [Andaman Islands] 

Macrobrachium aracamuni Rodríguez, 1982: 379; Fig. 2. [Cerro Aracamuni, Territorio Federal Amazonas, 

680 m above sea level] 

Macrobrachium asperulum (von Martens, 1868) 

= Palaemon asperulus von Martens, 1868: 43; Plate 1, fig. 5. [Shanghai, aus dem Fischmarkt] 

= Palaemon asperulus var. brevirostris Yu, 1931a: 287; Fig. 4. [Ichang, Changsha, Soochow] 

= Macrobrachium pinguis Dai, 1984: 245; Figs 1-5. [Longhai County, Fujian Province] 

= Macrobrachium anhuiense Tan, 1991: 286; Figs 1-4. [Wuwei County, (31°20'N 117°54'E), Anhui 

Province]


Macrobrachium assamense assamense (Tiwari, 1958) 

= Palaemon assamensis assamensis Tiwari, 1958: 298. [Someswari River, near Siju, Garo Hills, Assam] 

Macrobrachium assamense peninsulare (Tiwari, 1958) 

= Palaemon assamensis peninsularis Tiwari, 1958: 298. [Nerbudda River at Khetgaon, 22.50°N 81.20°E, 

Mandla District, Madhya Pradesh] 

Macrobrachium atabapense Pereira S., 1986: 202; Figs 4, 5, 6A. [Atabapo River, Sta. Cruz, Territorio Federal 

Amazonas, Venezuela, 3°20'N 67°29'W] 

Macrobrachium auratum Short, 2004: 71; Figs 27-28, 36C. [Flame Tree Creek between Airlie Beach and 

Shute Harbour, 20°16'S 148°45'E] 

Macrobrachium australe (Guérin-Méneville, 1838 [in Guérin-Méneville, 1829-1838]) 

= Palaemon australis Guérin-Méneville, 1838 [in Guérin-Méneville, 1829-1838]: 37. [île de Taïti] 

? = Palaemon Danae Heller, 1865: 120; Plate 11, fig. 3. [Sidney; considered erroneous by Short, 2004] 

= Palaemon dispar von Martens, 1868: 41. [Insel Adenare, umweit Flores] 

= Palaemon alphonsianus Hoffman, 1874: 33; Plate 9, figs 63-65. [l’île de la Réunion] 

= Palaemon parvus Hoffman, 1874: 35; Plate 7, fig. 59. [l’île de Nossy-Faly] 

= Palaemon Malliardi Richters, 1880: 166; Plate 18, figs 1-3. [Creole river, Black river] 

= P.[alæmon] (Eupalæmon) ustulatus Nobili, 1899: 241. [Rigo] 

= Leander lepidus De Man, 1915a: 410; Plate 28, figs 6-6d. [Mündung des kleinen Flusses zu Oinaké, 

einem Dorfe an der Küste östlich von der Humboldt-Bai] 

Macrobrachium australiense Holthuis, 1950a: 174. [nomen novum for Palaemon australis Ortmann, 1891] 

= Palaemon australis Ortmann, 1891: 709; nec Palaemon australis Guérin-Méneville, 1838. [Gayndah, 

Rockhampton (both Queensland); Peak Downs, Ost-Australien] 

= Macrobrachium adscitum adscitum Riek, 1951: 363; Fig. 3. [Queensland, Accomodation Creek, 

Oakey] 

= Macrobrachium adscitum subsp. Riek, 1951: 363. [Angus River, Strathalbyn, south of Adelaide, 

South Australia] 

= Macrobrachium atactum atactum Riek, 1951: 364; Fig. 5. [Conondale, Mary River, Queensland] 

= Macrobrachium atactum ischnomorphum Riek, 1951: 364; Fig. 6. [Elimbah Creek, Elimbah, Queensland] 

= Macrobrachium atactum sobrinum Riek, 1951: 364; Fig. 7. [Muttaburra, Queensland] 

= Macrobrachium australiense crassum Riek, 1951: 366; Fig. 10. [Cairns, Queensland] 

= Macrobrachium australiense cristatum Riek, 1951: 366; Fig. 9. [Pallal, Horton River, near Bingara, 

New South Wales] 

= Macrobrachium australiense eupharum Riek, 1951: 365; Fig. 8. [Burdekin River, Macrossan, Queensland] 

Macrobrachium banjarae (Tiwari, 1958) 

= Palaemon banjarae Tiwari, 1958: 299. [Banjar River, off Aonrai Forest Village, Baihar Tehsil, Dist. 

Balaghat, M.P.] 

Macrobrachium bariense (De Man, 1892a) 

= Palaemon (Macrobrachium) bariensis De Man, 1892a: 496; Plate 29, figs 50-50d. [Flores, aus dem 

Flusse bei Bari] 

Macrobrachium birai Lobão, Melo & Fernandes, 1986: 50. [Rio Branco, afluente do Itapitangui, Cananéia, 

SP, 24°54'44''S 47°5'30''W; fully described in Melo, Lobão & Fernandes, 1988] 

Macrobrachium birmanicum Schenkel, 1902 

= Palaemon spinipes var. birmanicus Schenkel, 1902: 503; Plate 9, figs 8-8c. [Mandalay and Bhamd; 

lectotype designated by Cai & Ng, 2002] 

Macrobrachium bombajense Almelkar & Sankolli, 2006: 187; Figs 1-2. [Maharasthra State Fisheries Department 

fish farm Aarey, Goregaon Mumbai, Maharasthra, India] 

= Macrobrachium bombayensis Indulkar & Shirgur, 1995: 40. [nomen nudum] 

Macrobrachium borellii (Nobili, 1896) 

= Palæmon Borellii Nobili, 1896: 2. [San Lorenzo (Jujuy) e età diversa della provincia di San Luis] 

Macrobrachium brasiliense (Heller, 1862b)


= P.[alaemon] brasiliensis Heller, 1862b: 419; Plate 2, fig. 46. [in einem Bache zu Camaroes, Brasilien] 

= Palaemon appuni var. aequatorialis Ortmann, 1891: 731; Plate 47, fig. 6. [Ecuador] 

Macrobrachium brevicarpum Tan & Dong, 1996: 287; Figs 1-4. [Qimen County (29°52'N 117°43'E), Anhui 

Province] 

Macrobrachium bullatum Fincham, 1987: 351; Fig. 1. [Magela Creek, Northern Territory, Australia] 

Macrobrachium cacharense (Tiwari, 1952) 

= Palaemon hendersoni cacharensis Tiwari, 1952: 32. [Cachar and Silchar districts, Assam] 

Macrobrachium caledonicum (Roux, 1926a) 

= Palaemon (Macrobrachium) caledonicus Roux, 1926a: 224; Figs 52-54. [Bondé; Tao; Canala; Koné et 

environs; Bopope; Coula-Boréaré; Ni; La Foa; Coindé (all Nouvelle-Calédonie)] 

Macrobrachium callirrhoe (De Man, 1898) 

= Palaemon (Macrobrachium) callirrhoë De Man, 1898: 152; Plate 8. [Mandai river at Nanga Raoen; 

Ketoengau river] 

Macrobrachium canarae (Tiwari, 1958) 

= Palaemon canarae Tiwari, 1958: 298. [Sitanadi River, near Ghats, South Kanara, Madras] 

Macrobrachium carcinus (Linnaeus, 1758) 

= Cancer Carcinus Linnaeus, 1758: 631. [Americæ fluvis] 

= Cancer (Astacus) Jamaicensis Herbst, 1792 [in Herbst, 1791-1796]: 57; Plate 27, fig. 2. [Jamaica, in 

Flussen] 

= Palaemon brachydactylus Wiegmann, 1836: 148. [Ostküste Mexiko] 

= P.[alemon] punctatus Randall, 1840: 146. [?East Indies and West Indies] 

= Palemon brevicarpus De Haan, 1849 [in De Haan, 1833-1850]: 172. [Japan; lectotype designation by 

Yamaguchi & Baba, 1993; specimens considered to be from Suriname, see Yamaguchi & Baba, 1993] 

= Palæmon aztecus de Saussure, 1857a: 504. [Vera-Cruz] 

? = Palæmon Montezumæ de Saussure, 1857a: 504. [Vera-Cruz] 

= P.[alaemon] laminatus von Martens, 1869: 24. [Caracas] 

= Palaemon ornatus Torralbas, 1917: 616; Figs 56, 57; nec Olivier, 1811. [corrientes de agua dulce, 

Cuba] 

= Periclimenes portoricensis Schmitt, 1933: 3; Fig. 2. [Porto Rico] 

Macrobrachium catonium H.H.III Hobbs & H.H.Jr. Hobbs, 1995: 50; Fig. 1. [Lake in Actun Chapat, Cayo 

District, Belize] 

Macrobrachium cavernicola (Kemp, 1924) 

= Palaemon cavernicola Kemp, 1924: 42; Plate 3, figs 1-4. [Siju Cave, Garo Hills, Assam] 

Macrobrachium chevalieri (Roux, 1935a) 

= Palaemon chevalieri (Macrobrachium) Roux, 1935a: 193; Figs 1-2. [Iles du Cap-vert, Saint Antonio 

Paul] 

Macrobrachium clymene (De Man, 1902) 

= Palaemon (Macrobrachium) clymene De Man, 1902: 794; Plate 25, figs 50-50h. [aus dem Baramflusse, 

Borneo] 

Macrobrachium cocoense Abele & W. Kim, 1984: 951; Figs 1-2. [Isla del Coco, Costa Rica, stream on east 

side of Wafer Bay] 

Macrobrachium cortezi Rodríguez, 1982: 383; Fig. 3. [Tobogán, 30 km S of Puerto Ayacucho, Río Orinoco] 

Macrobrachium cowlesi Holthuis, 1950a: 257. [Manila water supply, Luzon, Philippines; described and 

illustrated by Cowles, 1914 as Palaemon sp.] 

Macrobrachium crebrum Abele & W. Kim, 1989: 6; Fig. 2. [Republic of Panama, Panama Canal, Miraflores 

Third Locks Lake] 

Macrobrachium crenulatum Holthuis, 1950b: 95. [Pejebobo River. E. Panama] 

Macrobrachium dalatense Nguyên, 2003: 454; Figs 1-3. [Mountain stream near the village of Krean at about 

1250 m altitude, 11°52'N 108°29'E, south of Dalat] 

Macrobrachium dayanum (Henderson, 1893) 

= Palaemon Dayanus Henderson, 1893: 443; Plate 40, figs 7-13. [Orissa; Jubbulpore; Calcutta; Beerbhoom; 

Debroo; Delhi; Roorkee; Hurdwar; Loodiana; River Jumna; Lahore]


Macrobrachium denticulatum Ostrovski, Da Fonseca & Da Silva-Ferreira, 1996: 361; Figs 4-5. [Brazil, border 

between the states of Alagoas and Sergipe, São Francisco river, 9°37'30''S 37°46'W] 

Macrobrachium depressimanum Pereira S., 1993 

= Macrobrachium depressimanus Pereira S., 1993: 339; Figs 1, 2A. [Perú, Amazonas Department, Santiago 

River, La Poza, 4°01'S 77°47'W] 

Macrobrachium dienbienphuense Đăng & B.Y. Nguyên, 1972: 4; Fig. 3. [Nam Rom River, Dien Bien Phu, Lai 

Chau Province] 

= Macrobrachium longidigitum Dai, 1984: 248; Figs 18-22; nec Spence Bate, 1868a. [Ganlanba, Lancang 

River, Yunnan Province, China] 

= Macrobrachium eriocheirum Dai, 1984: 247; Figs 13-17. [Jingsan, Xishuangbanna Dai Aut. Pref., Yunnan 

Province] 

Macrobrachium dierythrum Pereira S., 1986: 204; Figs 7-8, 12C. [Aguaro River, Paso Garzerito, Edo. Guarico, 

Venezuela; 8°10'N 66°25'W] 

Macrobrachium digitus Abele & W. Kim, 1989: 8; Figs 3-4. [Republic of Panama, Panama Canal, Miraflores 

Locks] 

Macrobrachium digueti (Bouvier, 1895) 

= Palemon Digueti Bouvier, 1895: 159; Fig. 2. [rivière Mulege et canaux d’irrigation issus de cette 

rivière] 

Macrobrachium dolatum Cai, Naiyanetr & Ng, 2004: 598; Figs 6-7. [Trang, South Thailand] 

Macrobrachium dolichodactylus (Hilgendorf, 1879) 

= Palaemon (s.s.) dolichodactylus Hilgendorf, 1879: 840; Plate 4, fig. 18. [Tette] 

Macrobrachium duri Wowor & Ng, 2010b: 279; Figs 1-2. [Banten, Banten Province, Java] 

Macrobrachium dux (Lenz, 1910b) 

= Palaemon (Eupalaemon) dux Lenz, 1910b: 129; Plate 3, figs 2-5. [Avakubi am Ituri] 

= Palaemon (Eupalaemon) Lenzii De Man, 1911b: 225. [Congo River, probably near Boma] 

= Palaemon (Eupalaemon) dux var. congoensis De Man, 1912a: 416. [la rivière Kole, affluent du Lohali 

(Aruwimi), nord-est du Congo Belge] 

= Palæmon (Eupalæmon) dux var. congoensis De Man, 1912b: 229; Plate 4, figs 6-6a. [la rivière Kole, 

affluent du Lohali (Aruwimi), nord-est du Congo Belge] 

= Palaemon (Eupalaemon) dux var. tenuicarpus De Man, 1925: 47; Fig. 12k, g, l, h-j . [Kikada, Congo 

Belge] 

Macrobrachium edentatum Liang & Yan, 1986b: 107; Figs 1-4. [Junlian County, Sichuan Province] 

Macrobrachium elatum Jayachandran, 1987: 370; Fig. 2. [Ashtamudi estuary at Perumon, Quilon District, 

Kerala] 

Macrobrachium elegantum Pan, Hou & S. Li, 2010: 86; Figs 2-4. [unnamed cave in Xiaorui Village, Ludong 

Town, Jingxi County (23.1°N 106.4°E), Guangxi Zhuang Autonomous Region, China] 

Macrobrachium empulipke Wowor, 2010: 717; Figs 1-2. [sg. Cibogo, Sukabumi Regency, W. Java] 

Macrobrachium equidens (Dana, 1852a) 

= Palæmon equidens Dana, 1852a: 26. [in mari prope portum ”Singapore”] 

= Palaemon sundaicus var. bataviana De Man, 1897: 784. [Fluss zu Batavia] 

= P.[alæmon] (Eupalæmon) sundaicus var. brachydactyla Nobili, 1899: 238. [Amboina] 

= P.[alæmon] sundaicus var. De Mani Nobili, 1899: 239. [Atjeh; Amboina] 

= P.[alæmon] (Eupalæmon) acanthosoma Nobili, 1899: 242. [Katau] 

= Palaemon (Eupalaemon) sundaicus var. baramensis De Man, 1902: 770. [aus dem Baramflusse] 

= Palaemon (Eupalaemon) nasutus Nobili, 1903c: 9; unnumbered fig. [Singapore] 

= Palaemon delagoae Stebbing, 1915: 74; Plate 16. [mouth of river flowing into Delagoa Bay] 

= Urocaridella borradailei Stebbing, 1923: 8; Plate 14. [Umhlotzu River, Natal] 

Macrobrachium esculentum (Thallwitz, 1891) 

= Palaemon esculentus Thallwitz, 1891: 98. [Nord-Celebes; fully described in Thallwitz, 1892] 

= Palaemon dulcis Thallwitz, 1891: 99. [Nord-Celebes; fully described in Thallwitz, 1892] 

Macrobrachium faustinum (de Saussure, 1857a) 

= Palæmon Faustinus de Saussure, 1857a: 505. [Haiti] 

= Palaemon cubanus Sharp, 1893: 123. [Cuba; nomen nudum]


= P.[alemon] spinimanus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 399 (partim); nec 

Latreille, 1818. [les Antilles et les côtes du Brésil] 

Macrobrachium felicinum Holthuis, 1949b: 183. [Catumbela near Benguella, Angola] 

Macrobrachium ferreirai Kensley & Walker, 1982: 4; Figs 5-6, 12b. [Igarapé near Castanhal, Aripuaná] 

Macrobrachium feunteuni Keith & Vigneux, 2002: 130; Photo 25-33. [Nuku Hiva, îles Marquises, Polynésie 

Française, rivière Hakaui] 

Macrobrachium foai (Coutière, 1902a) 

= P.[alaemon] (Eupalæmon) Foai Coutière, 1902a: 517. [Haut Congo] 

Macrobrachium forcipatum Ng, 1995b: 251; Figs 2-4. [Tasik Temengor, south of Banding, Sungai Halong] 

Macrobrachium formosense Spence Bate, 1868a: 364; Plate 31, fig. 1. [River Tamsuy, Island of Formosa] 

= Palemon longipes De Haan, 1849 [in De Haan, 1833-1850]: 171; nec Olivier, 1811. [Japan; lectotype 

designation by Yamaguchi & Baba, 1993] 

= Palæmon similis Yu, 1931a: 281; Fig. 2. [Amoy] 

? = Palaemon riukiuensis Kubo, 1940a: 21; Figs 12-13; Plate 1B. [Okinawa-zima (Riu-Kiu)] 

Macrobrachium fukienense Liang & Yan, 1980: 30; Figs 1-7. [Fujian Province: Xiamen (Bantou, Dongfu), 

Tong’an (Dingxi), Anxi, Haicang (Jiaomei), Yongchun, Zhangpu (Shiliu)] 

Macrobrachium gallus Holthuis, 1952b: 67; Fig. 1. [Rio Peripa, Ecuador] 

Macrobrachium gangeticum Spence Bate, 1868a: 365. [Ganges River near Dufferin Bridge at Rajghat, Banaras 

India; neotype selection by Tiwari & Holthuis, 1996] 

= Palaemon choprai Tiwari, 1949a: 333; Figs 1-2. [Ganges River near Dufferin Bridge at Rajghat, Banaras 

India; lectotype selection by Tiwari & Holthuis, 1996] 

Macrobrachium glabrum Holthuis, 1995: 139; Figs 1-2. [Kotomay, sous-préfecture Mitsinjo, prov. Majunga, 

N.W. Madagascar] 

Macrobrachium gracilirostre (Miers, 1875) 

= Palaemon gracilirostris Miers, 1875: 343. [Samoa Islands, Upolu] 

= Palaemon (Parapalaemon) modestus De Man, 1892a: 469; Plate 27, figs 43-43d; nec Heller, 1862a. 

[Flores, Fluss bei Wukur unweit Sikka] 

= Palaemon (Parapalaemon) modestus brevimanus Roux, 1934a: 228; Figs 9- 10. [Bimun, Nouvelle- 

Irlande] 

= Macrobrachium sophronicum Holthuis, 1950a: 198; Fig. 40. [nomen novum name for Palaemon (Parapalaemon) 

modestus De Man, 1892a] 

Macrobrachium grandimanus (Randall, 1840) 

= P.[alemon] grandimanus Randall, 1840: 142. [Sandwich Islands] 

= P.[alemon] gracilimanus Randall, 1840: 143. [Sandwich Islands] 

= Palæmon acutirostris Dana, 1852a: 26. [insulis Hawaiensibus] 

Macrobrachium gua Chong, 1989: 32; Figs 1-2. [stream at resurgence from Gomantong Hill, ca. 5°33'N 

118°06'E, Sabah, Borneo] 

Macrobrachium guangxiense Liang & Yan, 1981: 33; Figs 20-21. [Longzhou County, Guangxi Province] 

Macrobrachium gurudeve Jayachandran & Raji, 2005: 1180; Figs 1-2, 6. [Bhavani River (at Thavalam), an 

east flowing mountain river, Palakkad District of Kerala State] 

Macrobrachium hainanense (Parisi, 1919) 

= Palaemon (Parapalaemon) hainanense Parisi, 1919: 87; Plate 2, fig. 1; Plate 6, figs 1, 7. [Fiume Ken- 

Kong, Isla di Hainan] 

Macrobrachium hancocki Holthuis, 1950b: 96. [Esparta, Rio Barranca, Costa Rica] 

Macrobrachium handschini (Roux, 1933) 

= Palaemon (Macrobrachium) handschini Roux, 1933: 345. [Katherine River, Northern Territory, Australia; 

lectotype designation by Bruce, 1992b] 

= Macrobrachium glypticum Riek, 1951: 363; Fig. 4. [Coen, north Queensland] 

Macrobrachium hendersodayanum (Tiwari, 1952) 

= Palaemon henderso-dayanus Tiwari, 1952: 29. [Western Ghats, from Satara District up to Mysore State] 

Macrobrachium hendersoni (De Man, 1906) 

= Palæmon (Parapalæmon ?) Hendersoni De Man, 1906: 405. [Darjeeling, Bengal, at a height of 2500 

feet]


= Palaemon yunnanensis Yü, 1936: 308; Figs 3, 4. [Mann-Tchi-Pan] 

Macrobrachium heterochirus (Wiegmann, 1836) 

= Palaemon heterochirus Wiegmann, 1836: 149. [Ostküste Mexiko] 

= Palaemon Appuni von Martens, 1869: 31; Plate 2, fig. 5. [Porto Cabello in Venezuela] 

Macrobrachium heterorhynchos Guo & He, 2008: 12; Figs 1-2, 7A-B. [Xijiang river near Jiangmen City, ca. 

22°33'N 113°08'E] 

Macrobrachium hildebrandti (Hilgendorf, 1893b) 

= Bithynis? hildebrandti Hilgendorf, 1893b: 244. [Central-Madagaskar] 

Macrobrachium hirsutimanus (Tiwari, 1952) 

= Palaemon hirsutimanus Tiwari, 1952: 31. [forest stream 95 km north from Nan Town, Nan province, 

about 15-20 km north from Ban Pon, Nam Gae, north of Ban Sala, near Laos, North Thailand; neotype 

selection by Cai, Naiyanetr & Ng, 2004] 

Macrobrachium hirtimanus (Olivier, 1811) 

= Palæmon hirtimanus Olivier, 1811: 663. [la mer des Indes] 

Macrobrachium hobbsi Villalobos Hiriart & Nates Rodriguez, 1990: 7; Fig. 3. [México, Chiapas, Río El 

Naranjo, aprox. a 8 km al NO de Pijijiapan, carretera Tonalá-Pijijipan] 

Macrobrachium holthuisi Genofre & Lobão, 1978: 273; Fig. 1. [Guaecá river of São Sebastião, State of São 

Paulo, Brazil] 

Macrobrachium horstii (De Man, 1892a) 

= Palaemon (Parapalaemon) Horstii De Man, 1892a: 460; Plate 27, figs 39-39c. [Celebes, aus dem Flusse 

bei Palopo, Luwu] 

= Palaemon (Parapal.) horsti brevidigitus Roux, 1930: 358. [Bali] 

Macrobrachium idae (Heller, 1862b) 

= P.[alaemon] Idae Heller, 1862b: 416; Plate 2, figs 40-41. [Borneo; lectotype selection by Cai & Ng, 

2001b] 

= Palaemon (Eupalaemon) ritsemae De Man, 1897: 774; Plate 37, figs 70-70c. [Atjeh] 

= Palæmon (Eupalæmon) Idæ var. subinermis Nobili, 1899: 237. [Innawi, Fiume San Guiseppe] 

= Palæmon (Eupalæmon) Mariæ Coutière, 1900a: 1266. [Rivière Ivaloina, près Tamatave] 

= Palaemon (Eupalaemon) robustus De Man, 1902: 771; Plate 24, fig. 48. [Tabelo, Halmahera; lectotype 

selection by Cai & Ng, 2001b] 

= Macrobrachium palawanensis Johnson, 1962: 307; Fig. 1. [Palawan] 

Macrobrachium idella idella (Hilgendorf, 1898) 

= Palaemon (Eupalaemon) idae var. idella Hilgendorf, 1898: 29; Fig. A. [Ungúu, Teich bei Matomondo; 

Usaramo] 

= Palæmon (Eupalæmon) multidens Coutière, 1900a: 1266. [Riv. Kotofotsy, Bras de l’Onilahy] 

Macrobrachium idella georgii Jayachandran & Joseph, 1985a: 130; Fig. 1. [Pallickal, Pamba, Manimala and 

Meenachil rivers] 

Macrobrachium iheringi (Ortmann, 1897) 

= Palaemon iheringi Ortmann, 1897: 211; Plate 1, figs 7-8. [Alto da Serra and Rio Tieté, Estado de São 

Paulo] 

Macrobrachium inca Holthuis, 1950b: 93. [Rio Mochè, near Salavery, N. Peru] 

Macrobrachium indicum Jayachandran & Joseph, 1986: 217; Figs 1-4. [Vellayani Lake, 8°24'09"-8°6'30"N 

76°59'08"-76°59'47"E] 

Macrobrachium inflatum Liang & Yan, 1985b: 254; Fig. 3. [Kunshan County, 31°20'N 120°58'E, Jiangsu 

Province] 

Macrobrachium inpa Kensley & Walker, 1982: 6; Figs 7-9, 12c. [Igarapé da Cachoeira] 

Macrobrachium insulare (Parisi, 1919) 

= Palaemon (Parapalaemon) insularis Parisi, 1919: 85; Fig. 7; Plate 3, figs 2-3; Plate 6, fig. 12. [Formosa] 

Macrobrachium jacobsoni Holthuis, 1950a: 227; Fig. 47. [Lugu, Sinabang Bay, Simalur, off the west coast of 

Sumatra] 

Macrobrachium japonicum (De Haan, 1849 [in De Haan, 1833-1850]) 

= Palaemon japonicus De Haan, 1849 [in De Haan, 1833-1850]: 172. [Japan; lectotype designation by 

Yamaguchi & Baba, 1993]


= Palaemon boninensis Stimpson, 1860a: 41. [insulis ”Bonin”, in rivulis montanis] 

Macrobrachium jaroense (Cowles, 1914) 

= Palaemon jaroensis Cowles, 1914: 385; Plate 3, figs 8-8k. [Hibucawan River, near Jaro, Leyte] 

Macrobrachium jayasreei Jayachandran & Raji, 2005: 1186; Figs 3-4. [Kabbini River (at Mananthavadi), an 

east flowing mountain river, Wynad District of Kerala State] 

Macrobrachium jelskii (Miers, 1877) 

= Palæmon jelskii Miers, 1877: 661; Plate 67, fig. 1-1b. [Guiana, Oyapock] 

Macrobrachium jiangxiense Liang & Yan, 1985b: 256; Fig. 4. [Boyang County, 29°N 116°40'E, Jiangxi 

Provinces] 

Macrobrachium johnsoni Ravindranath, 1979: 184; Figs 1-2. [Guntur fish market] 

Macrobrachium joppae Holthuis, 1950a: 233; Fig. 48. [Nias] 

Macrobrachium kelianense Wowor & Short, 2007: 82; Figs 3-4. [Sg. Kelian confluence with Sg. Ketang, 

Makaham River basin, E. Kalimantan] 

Macrobrachium kempi (Tiwari, 1949b) 

= Palaemon kempi Tiwari, 1949b: 330. [small stream between Chittagong and Sultan Bagu Bastan] 

Macrobrachium kistnense (Tiwari, 1952) 

= Palaemon kistnensis Tiwari, 1952: 28. [Vindhyan Satpuran range of mountains, beginning from the 

Tanda Falls in the Mirzapur District, U.P.; Western Ghats; Ceylon, Aravalli Hills] 

Macrobrachium kiukianense (Yu, 1931a) 

= Palæmon kiukianensis Yu, 1931a: 279; Fig. 1. [Kiukiang] 

Macrobrachium koombooloomba Short, 2004: 61; Figs 23-24, 36A. [Koombooloomba area, creek on Koombooloomba 

road near Koombooloomba Dam turnoff, 7°50'S 145°34'E] 

Macrobrachium kulkarnii Almelkar & Sankolli, 2006: 190; Figs 3-4. [fish farm of M/s Tata Hydro-Electric 

Company, Walvan Dam, Lonavala (Pune District), Maharasthra State, India] 

Macrobrachium kunjuramani Jayachandran & Raji, 2005: 1188; Fig. 5. [Kabbini River (at Chundal, Kalpetta, 

Vythiri, Thalapuzha, Noolpuzha, Kenipuzha), east flowing mountain river, Wynad District of 

Kerala State] 

Macrobrachium lamarrei lamarrei (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= P.[alemon] Lamarrei H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 397. [côtes du Bengale] 

Macrobrachium lamarrei lamarroides (Tiwari, 1952) 

= Palaemon lamarrei lamarroides Tiwari, 1952: 28. [Logtak Lake, Manipur, Assam] 

Macrobrachium lanatum Cai & Ng, 2002: 72; Figs 13-16. [southern Myanmar] 

Macrobrachium lanceifrons (Dana, 1852a) 

= Palæmon lanceifrons Dana, 1852a: 26. [insula ”Luzon”, archiepelagi Phillipinsis, prope portum 

”Manila”] 

= Palaemon lanceifrons var. montalbanensis Cowles, 1914: 371; Plate 2, figs 6-6i. [city of Manila at Montalban] 

Macrobrachium lanchesteri (De Man, 1911c) 

= Pal.[aemon] (Eupalaemon) Lanchesteri De Man, 1911c: 264. [nomen novum for Palæmon paucidens 

Lanchester, 1901] 

= Palæmon paucidens Lanchester, 1901: 568; Plate 33, fig. 4; nec De Haan, 1844, nec Hilgendorf, 1893a. 

[Singora] 

Macrobrachium lar (Fabricius, 1798) 

= Palaemon Lar Fabricius, 1798: 402. [in India] 

= Palaemon longimanus Fabricius, 1798: 402. [in India orientali] 

= Palæmon ornatus Olivier, 1811: 660. [Nouvelle-Hollande] 

= Palæmon tridens White, 1847a: 78. [nomen nudum] 

= P.[alaemon] vagus Heller, 1862b: 417; Plate 2, figs 42-43. [Amboina] 

= Palaemon spectabilis Heller, 1862a: 527. [Taiti] 

= Palaemon ruber Hess, 1865: 165; Plate 7, fig. 20. [Viti-Inseln] 

= Palaemon spectabilis Heller, 1865: 113; Plate 10, fig. 8. [Taiti]


= Palaemon longimanus Hoffman, 1874: 34; Plate 9, figs 68-69; nec Fabricius, 1798. [l’île de la Réunion] 

= Palaemon mayottensis Hoffman, 1874: 32; Plate 9, figs 61-62. [l’île de Mayotte; l’île de Nossy-Faly] 

= Palaemon reunionnensis Hoffman, 1874: 33; Plate 9, figs 66-67. [l’île de la Réunion] 

= Palaemon madagascariensis Hoffman, 1874: 35; Plate 7, fig. 58. [l’île de Nossy-Faly] 

= Leander dionyx Nobili, 1905d: 482; Plate 12, fig. 2. [Stephansort] 

= Cancer teatae Curtiss, 1938: 162. [Vaitapiha River, Tautira township, Tahiti] 

? = Macrobrachium ornatus Jayachandran & Raji, 2004: 41; Figs 1-2. [small rivulet of Muvattupuzha 

river at Pothanikkad, Kerala] 

Macrobrachium latidactylus (Thallwitz, 1891) 

= Palaemon latidactylus Thallwitz, 1891: 97. [Nord-Celebes; fully described in Thallwitz, 1892] 

= Palaemon (Eupalaemon) endehensis De Man, 1892a: 465; Plate 27, figs 42-42h. [Endeh, aus dem Nanga 

Ba; aus dem Flusse bei Reo; aus dem Flusse bei Bari; Mbawa, dicht am Meere; Sungei Nargi bei 

Konga (all localities from Flores)] 

= Palaemon (Macrobrachium) lampropus De Man, 1892a: 493; Plate 29, figs 49-49c. [aus einem Flusse 

bei Palopo; aus dem Wasserfalle Bantimurong unweit Maros; aus einem Flusse bei Pare-Pare (all 

from Celebes); Timor, aus dem Flusse Koinino bei Kupang] 

Macrobrachium latimanus (von Martens, 1868) 

= Palaemon latimanus von Martens, 1868: 44. [Philippinen, bei Loquilocun auf der Insel Samar] 

= Palaemon euryrhynchus Ortmann, 1891: 738; Plate 47, fig. 12. [Fidji-Levu; Fidji-Ins.] 

= Palœmon (Macrobrachium) singalangensis Nobili, 1900a: 487. [Aier Mantcior] 

Macrobrachium lepidactyloides (De Man, 1892a) 

= Palaemon (Macrobrachium) lepidactyloides De Man, 1892a: 497; Plate 29, figs 51-51c. [Flores, aus dem 

Flusse bei Mbawa, oberhalb des Wasserfalls] 

Macrobrachium lepidactylus (Hilgendorf, 1879) 

= Palaemon (s.s.) lepidactylus Hilgendorf, 1879: 838; Plate 4, figs 14-16. [Quellimane; Tette] 

= Palæmon (Macrobrachium) Hilgendorfi Coutière, 1899b: 382. [Madagascar, Côte Est, region des 

grandes forêts] 

Macrobrachium leptodactylus (De Man, 1892a) 

= Palaemon pilimanus var. leptodactylus De Man, 1892a: 476; Plate 28, fig. 44i-l. [Buitenzorg, Java, Indonesia; 

lectotype selection by Cai, Naiyanetr & Ng, 2004] 

Macrobrachium leucodactylus Wowor & Choy, 2001: 274; Figs 4-6. [tributary of Sg. Temburong-Machang, 

Temburong District, altitude 580 m] 

Macrobrachium lingyunense J. Li, Cai & Clarke, 2006: 277; Figs 1-3. [Shadong (Sand Cave), Guancang Village, 

Sicheng Town, Lingyun County, Guangxi Province, southern China, 106°23'-106°55'33''E 

24°05'-24°37'N] 

Macrobrachium lopopodus Wowor & Choy, 2001: 270; Figs 2-3. [Ulu Sg. Sawatan, Kimanis, Papar District, 

Sabah, Malaysia] 

Macrobrachium lorentzi (Roux, 1921) 

= Palaemon (Parapalaemon) lorentzi Roux, 1921: 596; Plate 16, figs 1-3. [Kllof-Biwak; Alkmaar; Went- 

Geb; Sabang (all Irian Jaya)] 

Macrobrachium lucifugum Holthuis, 1974 

= Macrobrachium faustinum lucifugum Holthuis, 1974: 233; Figs 2-3. [Cueva del Agua de Yara, on the 

territory of the ”bario” Yara, E. of Baracoa, Oriente Province, Cuba] 

= Macrobrachium crybelum Chace, 1975: 30; Figs 1-4. [Cave at Ciudad del Caribe, 18°58'N 70°23'W, 

Santo Domingo, D.N., Dominican Republic] 

Macrobrachium lujae (De Man, 1912a) 

= Palaemon (Eupalaemon) Lujæ De Man, 1912a: 415. [Kondué, dans le district de Kasaï, sur la rivière 

Sankuru, dans les parages de Lusambo] 

= Palæmon (Eupalæmon) Lujæ De Man, 1912b: 215; Plate 2, figs 3-3b; Plate 3, figs 3-3e. [Kondué, dans 

le district de Kasaï, sur la rivière Sankuru, dans les parages de Lusambo] 

Macrobrachium macrobrachion (Herklots, 1851)


= Palemon macrobrachion Herklots, 1851: 15. [Boutry] [species name spelled mabrobrachion on page 15 

and macrobrachion on page 25; former treated as an inadvertent spelling mistake by Holthuis, 1951a] 

= Palæmon africanus Kingsley, 1883: 107. [West coast of Africa] 

Macrobrachium maculatum Liang & Yan, 1980: 31; Figs 8-14. [Fujian Province: Yong’an (Jiulongxi), 

Liancheng (Xianshuitang)] 

Macrobrachium madhusoodani Unnikrishnan, P.M. Pillai & Jayachandran, 2011: 123; Fig. 1. [Ithikkara River, 

8°56'42''N 76°56'20''E, Kerala, South India] 

= Macrobrachium madhusoodani Unnikrishnan, P.M. Pillai & Jayachandran, 2010: 1116; Figs 1-2. [upper 

reaches of the Ithikkara River, 8°56'42''N 76°56'20''E, Kerala, South India; unavailable under Art. 16.4.2] 

Macrobrachium malayanum (Roux, 1935b) 

= Palæmon (Macrobrachium) pilimanus malayanus Roux, 1935b: 32. [Lasah, Plus Valley, East Perak, 

Malay Peninsula] 

= Macrobrachium geron Holthuis, 1950a: 258; Fig. 52. [Banka, E. of Sumatra] 

Macrobrachium malcolmsonii kotreeanum Johnson, 1973: 279. [Kotree, Indus River, Pakistan] 

Macrobrachium malcolmsonii malcolmsonii (H. Milne Edwards, 1844) 

= Palemon Malcolmsonii H. Milne Edwards, 1844: 8; Plate 3. [eaux douces près de Naguepore, à une 

distance d’environ 200 lieues de la mer] 

Macrobrachium mammillodactylus (Thallwitz, 1892) 

= Palaemon idea var. mammillodactylus Thallwitz, 1892: 15. [Nord-Celébes; Luzon] 

= Palaemon (Eupalaemon) Wolterstorffi Nobili, 1900b: 1. [Sœrabaia-Giava] 

= Palaemon philippinensis Cowles, 1914: 340; Plate 2, figs 2-2m. [San Juan River; Pasig River in the city 

of Manila] 

? = Palæmon talavaræ Blanco, 1939b: 168; Plate 2. [Lake Sampaloc, San Pablo, Laguna province, Luzon] 

Macrobrachium manipurense (Tiwari, 1952) 

= Palaemon manipurensis Tiwari, 1952: 30. [Manipur, Assam] 

Macrobrachium manningi Pereira & Lasso, 2007: 134; Figs 1-3. [Cucurital River, affluent of Caroni River, 

Orinoco Basin, Cainama National Park, Bolivar State, Venezuela, 6°00'N 62°47'W, 380 m a.s.l.] 

Macrobrachium meridionale Liang & Yan, 1983b 

= Macrobrachium meridionalis Liang & Yan, 1983b: 213; Fig. 2. [Jongjiang River, Chengmai County, 

Hainan Island, China] 

Macrobrachium michoacanus Villalobos Hiriart & Nates Rodriguez, 1990: 2; Fig. 2. [México, Michoacán, 

Río Mexcalhuacán, aprox. 40 km al NO de Playa Azul, carretera Azul-Caleta de Campos] 

Macrobrachium microps Holthuis, 1978b: 210; Figs 1-2. [Danmin Cave, near Konogusgus, New Ireland] 

Macrobrachium mieni Đăng, 1975: 68; Fig. 2. [streams in Kim Boi, Hoa Binh Province] 

Macrobrachium minutum (Roux, 1917) 

= Palaemon minutus Roux, 1917: 599; Plate 27, Figs 1-3. [Lac Sentani] 

Macrobrachium miyakoense Komai & Fujita, 2005: 14; Figs 1-6. [anchialine cave located at southern coast 

of Miyako Island, Ryukyu Islands] 

Macrobrachium moorei (Calman, 1899a) 

= Palaemon moorei Calman, 1899a: 709; Plate 40, figs 20-24. [Lake Tanganyika, 50ft] 

Macrobrachium naso (Kemp, 1918a) 

= Palaemon naso Kemp, 1918a: 91; Plate 25, figs 1-5. [Sizon, Tang-do cirle of Yawnghwe State in the 

river that runs south from the Inlé Lake] 

Macrobrachium nattereri (Heller, 1862b) 

= P.[alaemon] Nattereri Heller, 1862b: 414; Plate 2, figs 36-37. [Brasilien, im Rio Negro] 

Macrobrachium natulorum Holthuis, 1984a: 164; Figs 2-3. [Jawej River near Tigi Lake] 

Macrobrachium neglectum (De Man, 1905) 

= Palaemon (Eupalaemon) neglectus De Man, 1905: 201; Plate 15, fig. 6. [lower Batak landen, near Deli, 

North Sumatra; lectotype designation by Wowor & Ng, 2010b] 

Macrobrachium nepalense Kamita, 1974: 10; Fig. 4; Plate 2, fig. 2. [Tarahara Farm (Birat.) off the Breeding 

and Distribution Centre] 

Macrobrachium niloticum (Roux, 1833) 

= Palæmon Niloticus Roux, 1833: 73; Plate 7, fig. 2. [les eaux de Nil]


Macrobrachium niphanae Shokita & Takeda, 1989: 148: Figs 1-2; Plate 1. [Nang Rong waterfall stream, 

Thailand] 

Macrobrachium nipponense (De Haan, 1849 [in De Haan, 1833-1850]) 

= Palaemon nipponense De Haan, 1849 [in De Haan, 1833-1850]: 171. [Japan; lectotype designation by 

Yamaguchi & Baba, 1993] 

= Palaemon asper Stimpson, 1860a: 41; nec Latreille, 1818. [in fluvii et rivulis Sinenses prope urbem 

”Canton”] 

= Palaemon sinensis Heller, 1862a: 528. [Schanghai] 

= Palaemon sinensis Heller, 1865: 119; Plate 10, fig. 11. [Shanghai] 

= Macrobrachium obtusifrons Dai, 1984: 246; Figs 6-12. [Guanting Reservoir, Miyun County, Beijing] 

= Macrobrachium meishanense Tan & Lu, 1992: 169; Figs 1-4. [rivulet from Meishan Reservoir, Jinzhai 

County, west Anhui Province] 

Macrobrachium nobilii (Henderson & Matthai, 1910) 

= Palaemon nobilii Henderson & Matthai, 1910: 295; Plate 17, figs 6a-e. [Walajabad, Chingleput District] 

Macrobrachium novaehollandiae (De Man, 1908c) 

= Pal. (Eupal.) novæ-hollandiæ De Man, 1908c: 370; Plate 16. [neighbourhood of Sydney] 

Macrobrachium occidentale Holthuis, 1950b: 95. [Rio de los Esclavos, just S of Cuilapa, Guatemala] 

Macrobrachium oenone (De Man, 1902) 

= Palaemon (Macrobrachium) oenone De Man, 1902: 784; Plate 25, figs 49-49c. [Kau and Saluta, Halmahera] 

= Palaemon (Macrobrachium) oenone papuana Roux, 1927b: 324; Fig. 2. [Pionierbovak, Fl. Mamberamo, 

Nouvelle-Guinée] 

Macrobrachium ohione (Smith, 1874) 

= Palæmon Ohionis Smith, 1874: 640. [Ohio River at Cannelton, Indiana] 

= P.[alaemon] sallei Kingsley, 1883: 108. [Mississippi] 

Macrobrachium olfersii (Wiegmann, 1836) 

= Palaemon Olfersii Wiegmann, 1836: 150. [Küste Brasiliens] 

= P.[alemon] spinimanus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] nec Latreille, 

1818: 399 (partim). [les Antilles et les côtes du Brésil] 

= Palæmon consobrinus de Saussure, 1857a: 504. [Vera-Cruz] 

= Palaemon Desausuri Heller, 1862b: 420; Plate 2, fig. 47. [Neu-Granada] 

= Palaemon Potiporanga Müller, 1880: 152. [Itajahy] 

Macrobrachium oxyphilus Ng, 1992: 442; Figs 1A, C, E, G; 2A, C; 3A, C. [39 km road marker, Sungai Besar 

to Tanjong Malim trunk road, north Selangor peat swamp forest, Peninsular Malaysia] 

Macrobrachium panamense Rathbun, 1912a 

= Macrobrachium acanthurus panamense Rathbun, 1912a: 1. [Rio Calabre, Republic of Panama] 

Macrobrachium patheinense Phone & Suzuki, 2004: 524; Figs 1-2. [Mayan Creek, Thayet Kone village, 

Pathein City, Ayeyawaddy Division] 

Macrobrachium patsa (Coutière, 1899b) 

= Palæmon (Parapalæmon) Patsa Coutière, 1899b: 382. [Madagascar, Rivière Mahanara (Côte Est); 

Bras de l’Onilahy (Côte Ouest)] 

Macrobrachium pectinatum Pereira S., 1986: 200; Figs 2, 3, 6B. [Atabapo River, Sta. Cruz, Territorio Federal 

Amazonas, Venezuela, 3°20'N 67°29'W] 

Macrobrachium peguense (Tiwari, 1952) 

= Palaemon peguensis Tiwari, 1952: 27. [Burma] 

Macrobrachium pentazona He, Gao & Guo, 2009: 39; Figs 1-3. [Beijiang River, near Qingyuan City (ca. 

23°42'N 113°01'E)] 

Macrobrachium petersii (Hilgendorf, 1879) 

= Palaemon (s.s.) Petersii Hilgendorf, 1879: 841; Plate 44, fig. 19. [Tette] 

Macrobrachium petiti (Roux, 1934b) 

= Palaemon (Macrobrachium) Petiti Roux, 1934b: 537; Figs 1-2. [Vatomandry]


Macrobrachium petronioi Melo, Lobão & Fernandes, 1986: 51. [Rio Branco, afluente do Itapitangui, 

Cananéia, SP, 24°54'44''S 47°58'30''W; fully described in Melo, Lobão & Fernandes, 1988] 

Macrobrachium pilimanus (De Man, 1879) 

= Palaemon pilimanus De Man, 1879: 181. [Moearalboeh, West Sumatra, Indonesia; lectotype selection 

by Cai, Naiyanetr & Ng, 2004] 

= Palaemon (Macrobrachium) pygmaeus Roux, 1928b: 222; Figs 1-4. [Telaga Kastobo, Bawean] 

Macrobrachium pilosum Cai & Dai, 1999: 226: Figs 10-12. [mountain stream near Mengban village, Mengla 

County] 

Macrobrachium placidulum (De Man, 1892a) 

= Palaemon (Macrobrachium) placidulus De Man, 1892a: 489; Plate 28, figs 48-48g. [Fluss bei Mbawa, 

oberhalb des Wasserfalls; aus dem Flusse Dona bei Endeh; Fluss Ba bei Endeh; Fluss Lella bei Sikka, 

aus dem Flusse Wukur bei Sikka (all from Celebes); Saleyer, aus dem Flusse Bangkalan; Celebes, 

Fluss bei Palopo; Timor, aus dem Flusse Koinino bei Kupang] 

? = Palæmon spinimanus Latreille, 1818: 5; Plate 319, fig. 1. [Type locality not indicated] 

Macrobrachium placidum (De Man, 1892a) 

= Palaemon (Macrobrachium) placidus De Man, 1892a: 483; Plate 28, figs 46-46c. [Sumatra, Fluss bei 

Kaju-tanam] 

Macrobrachium platycheles Ou & Yeo, 1995: 300; Figs 1-3. [stream leading from Nee Soon swamp forest, 

150 m upstream from pipeline, Singapore] 

Macrobrachium platyrostris (Tiwari, 1952) 

= Palaemon hendersoni platyrostris Tiwari, 1952: 32. [Type locality not indicated] 

Macrobrachium poeti Holthuis, 1984b: 143; Fig. 1. [Luwang Jurangjero, grid 668115, ca 100 m below entrance] 

Macrobrachium potiuna (Müller, 1880) 

= Palaemon Potiuna Müller, 1880: 152. [aus dem Itajahy; fully described and illustrated in Müller, 1892] 

Macrobrachium praecox (Roux, 1928c) 

= Palaemon (Eupalaemon) praecox Roux, 1928c: 43. [El Mene; El Pozon (both Prov. Falcon, Vénézuela); 

Santander, Colombie] 

Macrobrachium pumilum Pereira S., 1986: 208; Figs 11, 12B. [Aguaro River, Cachimbo pass, Edo. Guarico, 

Venezuela; 8°10'N 66°35'W] 

Macrobrachium purpureamanus Wowor, 1999: 34; Figs 1A-B, 2A, C, E-F, 3A, C, E, 4. [Sungai Nibong B, 

nearby Kampung Baru, Kundur Island, Riau Province, Indonesia, 0°46'46.8''N 103°28'05.4''E] 

Macrobrachium quelchi (De Man, 1900) 

= Palæmon (Macrobrachium) Quelchi De Man, 1900: 57; Plate 6, figs 1-8. [Upper Mazaruni River, 

2500ft; Mt. Roirama range, 3500ft] 

Macrobrachium raridens (Hilgendorf, 1893c) 

= Palaemon (Eupalaemon?) paucidens Hilgendorf, 1893a: 155; nec De Haan, 1844 [in De Haan, 1833- 

1850]. [Adeli bei Bismarckburg, Togoland] 

= Palaemon (Eu.[palaemon]) raridens Hilgendorf, 1893c: 181. [nomen novum for Palaemon (Eupalaemon?) 

paucidens Hilgendorf, 1893a] 

Macrobrachium rathbunae Holthuis, 1950b: 94. [Hog Creek Valley, San José Island, Archipielago de las 

Perlas, Gulf of Panama] 

Macrobrachium reyesi Pereira S., 1986: 198; Figs 1, 6C. [Quebrada Corral de Piedra, El Limón, Maracay, 

Edo. Aragua, Venezuela, 10°15'N 67°35'W] 

Macrobrachium rhodochir Ng, 1995a: 182; Fig. 1. [small stream parallel to Tajur Waterfall, Sungaj Tajur] 

Macrobrachium rodriguezi Pereira S., 1986: 206; Figs 10, 12A. [Caris River, El Tigre, Edo. Anzoategui, 

Venezuela, 8°45'N 64°50'W] 

Macrobrachium rogersi (Tiwari, 1952) 

= Palaemon rogersi Tiwari, 1952: 31. [Arakan and Pegu Yomas, Burma] 

Macrobrachium rosenbergii (De Man, 1879) 

= Palaemon Rosenbergii De Man, 1879: 167. [Batavia (Jakarta), Java, Indonesia; neotype designated by 

Wowor & Ng, 2008; see Anon., 2010] 

= Palaemon d’Acqueti Sunier, 1925: cxvii. [Batavia, Java, Indonesia; lectotype designation by Wowor 

& Ng, 2007]


= Palaemon whitei Sharp, 1893: 122. [Bombay] 

= Macrobrachium rosenbergii schenkeli Johnson, 1973: 274, 277. [Tavoy, Burma] 

Macrobrachium rostratum X. Wang, 1997: 460; Fig. 2. [Jishou, Hunan Province] 

Macrobrachium rude (Heller, 1862a) 

= Palaemon rudis Heller, 1862a: 527. [Ceylon] 

= Palaemon rudis Heller, 1865: 114. [Ceylon] 

= Palaemon (s.s.) Mossambicus Hilgendorf, 1879: 839; Plate 4, fig. 17. [Quellimane; Moçambique] 

= Palaemon (Eupalaemon) Alcocki Nobili, 1903a: 9; Plate 2, fig. 5. [Pondichéry] 

Macrobrachium sabanus Ng, 1994: 76; Figs 4-5. [waterfall stream, off Safoda Oli Palm Plantation, between 

5 and 10 km marks] 

Macrobrachium saigonense Nguyên, 2006a: 236; Figs 1, 2a-b, 3a-j, 4-6. [near Hoa An Bridge, on the Dong 

Nai river, Bien Hoa province, about 30 km northwest of Ho-Chi-Minh City, 10°55'N 106°50'E] 

Macrobrachium sankolli Jalihal & Shenoy in Jalihal, Shenoy & Sankolli, 1988: 11; Figs 3-4. [Malaprabha 

River, near old bridge, Khanapur] 

Macrobrachium santanderensis Garcia-Perez & Villamizar, 2009: 62; Figs 2-4. [70 km on an east to west 

road connecting the cities of Bucaramanga and Barrancabermeja (Department of Santander, Central 

East of Colombia, South America), 900 m above sea level, 07°06'15''N 73°24'44''W] 

Macrobrachium sbordonii Mejía-Ortíz, Baldari & López-Mejía, 2008: 50; Figs 2-4. [terminal lake in the 

Sistema de la Lucha, Chipas, Mexico, 17°03'40''N 93°53'23''W] 

Macrobrachium scabriculum (Heller, 1862a) 

= Palaemon scabriculus Heller, 1862a: 527. [Ceylon] 

= Palaemon scabriculus Heller, 1865: 117; Plate 10, fig. 9. [Ceylon] 

= P.[alaemon] dubius Henderson & Matthai, 1910: 300; Plate 18, figs 9a-d. [Walajabad, Saidapet and 

other localities in the Chingleput District] 

Macrobrachium scorteccii Maccagno, 1961: 336; Plates 17-18. [Càl Galloàn, Somalia] 

Macrobrachium shaoi Cai & Jeng, 2001: 276; Figs 1-2. [stream of tributary of Shuangchi river in Taipei 

County, northern Taiwan] 

Macrobrachium shokitai Fujino & Baba, 1973: 101; Figs 1-4. [River head, Urauchi River, Iriomote Island, 

Ryukyu Islands] 

Macrobrachium sintangense (De Man, 1898) 

= Palaemon (Eupalaemon) sintangensis De Man, 1898: 138; Plate 6. [Sintang] 

= Palaemon (Eupalaemon) elegans De Man, 1892a: 440; Plate 26, figs 36-36d; nec Rathke, 1837. [Buitenzorg; 

Sinagar (Java)] 

Macrobrachium sirindhorn Naiyanetr, 2001: 610; Figs 1-2; Plate 1. [Pong Nam Dung Waterfall, Mae Soon, 

Amphoe Fang, Chiang Mai Province, northern Thailand] 

Macrobrachium siwalikense (Tiwari, 1952) 

= Palaemon siwalikensis Tiwari, 1952: 28. [base of Simla Hills] 

Macrobrachium sollaudii (De Man, 1912a) 

= Palaemon (Eupalaemon) Sollaudii De Man, 1912a: 413. [dans la rivière Ottenge, près de Banzyville, 

dans le Congo Belge] 

= Palæmon (Eupalæmon) Sollaudii De Man, 1912b: 205; Plate 1, figs 2-2i. [dans la rivière Ottenge, près 

de Banzyville, dans le Congo Belge] 

Macrobrachium spinipes (Schenkel, 1902) 

= Palaemon spinipes Schenkel, 1902: 501; Plate 9, figs 7-7b; nec Desmarest, 1817. [Kema, Minahasa; see 

Ng & Wowor, 2011] 

= Macrobrachium wallacei Wowor & Ng, 2008 (nomen novum for Macrobrachium rosenbergii sensu 

Wowor & Ng, 2007): 291. [Laloki River, ca. 30 miles from Port Moresby, Papua New Guinea] 

Macrobrachium spinosum Cai & Ng, 2001b: 676; Figs 8-9. [Sungai Dadago, Halmahera] 

Macrobrachium srilankense Costa, 1979: 60; Fig. 6; Plate 1D. [Negombo lagoon; Kelni river at Kelaniya; 

Panadura] 

Macrobrachium striatum N.N. Pillai, 1991 

= Macrobrachium striatus N.N. Pillai, 1991: 249; Figs 1-3. [Cochin backwater around Thevara area, 

Kerala State, South India]


Macrobrachium sulcatus (Henderson & Matthai, 1910) 

= Palaemon sulcatus Henderson & Matthai, 1910: 289; Plate 16, fig. 4a-g. [Cochin] 

Macrobrachium sulcicarpale Holthuis, 1950a: 220; Fig. 45. [Bangkalan River, Salajar, SE of Celebes] 

Macrobrachium sundaicum (Heller, 1862b) 

= P.[alaemon] sundaicus Heller, 1862b: 415; Plate 2, figs 38-39. [Java, Indonesia; lectotype designated 

by Wowor & Ng, 2010b] 

= P.[alaemon] javanicus Heller, 1862b: 421; Plate 2, fig. 48. [Java, Indonesia; lectotype designation by 

Wowor & Ng, 2010b] 

= Palaemon (Parapalaemon) Trompii De Man, 1898: 144; Plate 7. [Ketungau river, Sintang Regency, W. 

Kalimantan; lectotype designation by Wowor & Ng, 2010b] 

= Palaemon (Parapalaemon) thienemanni Roux, 1932: 565, 570; Figs a-b. [Sg. Musi near Muara Klingi, 

S. Sumatra; lectotype designation by Wowor & Ng, 2010b] 

= Palæmon (Parap.) trompi armatus Roux, 1936: 30. [Gunong Pulai, Johore] 

Macrobrachium suongae Nguyên, 2003: 460; Figs 4-5. [Stream near An Phu village, 10 km south of Pleiku 

city (altitude about 880 m), 13°57'N 108°02'E, centre of Vietnam] 

Macrobrachium superbum (Heller, 1862a) 

= Palaemon superbus Heller, 1862a: 528. [Schanghai] 

= Palaemon superbus Heller, 1865: 118; Plate 10, fig. 10. [Schanghai] 

Macrobrachium surinamicum Holthuis, 1948: 1112. [Plantation ”Geyersvlijt” near Paramaribo, Surinam] 

Macrobrachium tenellum (Smith, 1871) 

= Palæmon tenellus Smith, 1871: 98. [Polvon, Occidental Department of Nicaragua] 

= Palaemon longipes Lockington, 1878a: 161. [Mulege River, West Coast of Gulf of California] 

Macrobrachium tenuirostrum X. Wang, 1997: 459; Fig. 1. [Tongren, Guizhou Province] 

Macrobrachium thai Cai, Naiyanetr & Ng, 2004: 605; Figs 8-9. [Nong Khai, north-est Thailand] 

Macrobrachium therezieni Holthuis, 1965b: 281; Fig. 1. [Fénérive district, Tamatave province, eastern Madagascar] 

Macrobrachium thuylami Nguyên, 2006b: 1; Figs 1-3. [local market in Duc Lap, a district in the highlands 

of South Vietnam, at 12°26'N 107°39'E (about 300 km northeast of Ho Chi Minh City)] 

Macrobrachium thysi Powell, 1980: 318; Figs 1-3. [Côte d’Ivoire, Banco forêt] 

Macrobrachium tiwarii Jalihal, Shenoy & Sankolli, 1988: 27; Figs 8-9. [Malaprabha River, near old bridge, 

Khanapur] 

Macrobrachium tolmerum Riek, 1951: 362; Fig. 1. [Black River, Macrossan, Queensland, Australia; considered 

erroneous by Short, 2004] 

Macrobrachium totonacum Mejía, Alvarez & Hartnoll, 2003: 79; Figs 2-3. [spring of the San Antonio River, 

Oaxaca, Mexico,18°28'8"N 96°38'6"W, 90 m altitude] 

Macrobrachium transandicum Holthuis, 1950b: 94. [Rio Telembi, near San Lorenzo, S.W. Colombia] 

Macrobrachium tratense Cai, Naiyanetr & Ng, 2004: 595; Figs 3-4, 5A. [Khlong Fuai, road 3271, 12°23'44.8''N 

102°39'34.5''E, Trat Province] 

Macrobrachium trichodactylum Liang, Liu & Chen in Li, Liu, Liang & Chen, 2007: 136; Fig. 54. [Qiong-Hai 

city, Hainan] 

Macrobrachium tuxtlaense Villalobos & Alvarez, 1999: 746; Figs 1-3. [small stream 1 km S of Coyame, 

18°26'N 95°7'W, tributary of lake Catemaco, Municipio de Catemaco, Veracruz, México] 

Macrobrachium unikarnatakae Jalihal, Shenoy & Sankolli, 1988: 21; Figs 6-7. [Hosayellapur tank (Hirekeri), 

Dharwad] 

Macrobrachium urayang Wowor & Short, 2007: 78; Figs 1-2. [Sg. Enggeng I’ut, a tributary of Sg. Bahau, 

Kayan Basin, East Kalimantan] 

Macrobrachium veliense Jayachandran & Joseph, 1985b: 185; Figs 1-2. [Veli Lake, 08°30'-08°32'N 76°53'- 

76°55'E] 

Macrobrachium venustum (Parisi, 1919) 

= Palaemon (Eupalaemon) venustus Parisi, 1919: 82 (as P. vernustus); Plate 4, fig. 1; Plate 6, figs 5-13 (as 

P. venustus). [Isla di Hainan] 

Macrobrachium vicconi Román, Ortega & Mejía, 2000: 186-192; Figs 1-12. [Río Perlas, Municipio de 

Ocosingo, state of Chiapas, México, 16°47'78″N, 91°29′29″W]


Macrobrachium vietnamiense Đăng in Đăng & B.Y. Nguyên, 1972: 2; Fig. 2. [Ky Phu stream, Bac Thai 

Province] 

Macrobrachium villalobosi H.H.Jr. Hobbs, 1973a: 77; Fig. 3. [Cueva del Naciemento del Río San Antonio, 

10 km SSW Acatlán, Oaxaca, México] 

Macrobrachium villosimanus (Tiwari, 1949b) 

= Palaemon villosimanus Tiwari, 1949b: 329. [Calcutta, Pulta Waterworks] 

Macrobrachium vollenhoveni (Herklots, 1857) 

= Palaemon Vollenhovenii Herklots, 1857: 96. [côte de Guinée] 

= P.[alaemon] jamaicensis var. africanus Bouvier, 1895: 160. [les rivières de l’Afrique occidentale] 

= Palæmon (Macrobrachium) jamaicensis var. angolensis De Man, 1904: 314; Plate 19, figs 39-45, 47; 

Plate 20, Figs 46, 48-53. [river at Catumbella, near Benguella, Angola] 

= Palæmon (Macrobrachium) jamaicensis var. Herklotsii De Man, 1912b: 239. [Mayumba, Bas-Congo] 

Macrobrachium walvanense Almelkar, Jalihal & Sankolli, 1999: 328; Figs 1-19. [fish rearing ponds of 

Messrs. Tata Hydro-Electric Company’s fish farm, Walvan Dam, Lonavala (Pune district), Maharashtra 

State, India, 540 m above mean sea level] 

Macrobrachium wannanense Dai & Tan, 1993: 301; Figs 1-4. [Shitai County, 30°11'N 117°27'E, Anhui Province] 

Macrobrachium weberi (De Man, 1892a) 

= Palaemon (Eupalaemon) Weberi De Man, 1892a: 421; Plate 25, figs 33-33h. [See von Tempe; See von 

Sidenreng; aus dem Flusse Tjenara bei Pampanua (all from Celebes)] 

Macrobrachium yui Holthuis, 1950a: 211. [nomen novum for Palaemon brevicarpus var. heterochirus Yü, 1936] 

= Palaemon brevicarpus var. heterochirus Yü, 1936: 305; Figs 1-2; nec Palaemon heterochirus Wiegmann, 

1836. [ning-Erh] 

= Macrobrachium yeti Đăng, 1975: 67; Fig. 1. [streams in Son La Province] 

Macrobrachuim zariquieyi Holthuis, 1949b: 178; Figs 1-2. [Fernando Poo, Rio Consul, 1 km from its mouth] 

Nematopalaemon Holthuis, 1950a 

= Nematopalaemon Holthuis, 1950a (type species Leander tenuipes Henderson, 1893, by original designation, 


Nematopalaemon colombiensis (Squires & Mora L., 1971) 

= Palaemon (Nematopalaemon) colombiensis Squires & Mora L., 1971: 102; Fig. 1. [off Pizarro, Colombia, 

5°09'N 77°28'W, 9.1 m] 

Nematopalaemon hastatus (Aurivillius, 1898a) 

= Palaemon (Leander) hastatus Aurivillius, 1898a: 27; Plate 4, figs 3-6. [Kamerun, im Meere bei Beticka 

ba Mallale] 

Nematopalaemon karnafuliensis (Ali Azam Khan, Fincham & Mahmood, 1980) 

= Palaemon (Nematopalaemon) karnafuliensis Ali Azam Khan, Fincham & Mahmood, 1980: 85; Figs 1-2. 

[Karnafuli Estuary, Chittagong, Bangladesh] 

Nematopalaemon schmitti (Holthuis, 1950b) 

= Palaemon schmitti Holthuis, 1950b: 97. [mouth of the Surinam River, near Resolutie, Dutch Guiana] 

Nematopalaemon tenuipes (Henderson, 1893) 

= Leander tenuipes Henderson, 1893: 440; Plate 40, Figs 14-15. [Bombay; Gulf of Martaban; Madras] 

= Palæmon luzonensis Blanco, 1939c: 201; Plate 1. [Aparri, Cagayan Province, Luzon] 

Neopalaemon H.H.Jr. Hobbs, 1973b 

= Neopalaemon H.H.Jr. Hobbs, 1973b (type species Neopalaemon nahuatlus H.H.Jr. Hobbs, 1973b, by 


Neopalaemon nahuatlus H.H.Jr. Hobbs, 1973b: 26, Figs 1-2. [Cueva del Guano, 10 km NE Valle Nacional, 

Oaxaca, México] 

Palaemon Weber, 1795 

= Palaemon Weber, 1795 (type species Palaemon adspersus Rathke, 1837, by direction under the plenary 

power of the ICZN, gender masculine; name placed on the Official List of Generic Names in 



= Palaemon Fabricius, 1798 (type species Cancer Squilla Linnaeus, 1758, designated by Latreille, 1810, 

gender masculine; name placed on the Official Index of Rejected and Invalid Names in Zoology in 


= Palaeander Holthuis, 1950a (type species Palaemon elegans Rathke, 1837, by original designation, 


Palaemon adspersus Rathke, 1837 

= Palaemon adspersus Rathke, 1837: 368; Plate 4, fig. 4. [an den vom schwarzen Meere bespülten 

Küsten der Krym] 

= Cancer Squilla Linnaeus, 1758: 632. [Mare Balthico, Oceano Europæo] 

= Palaemon communis Anslijn, 1826: 262. [nomen nudum] 

= Palaemon Fabricii Rathke, 1843: 6. [Christiansund] 

= Palaemon rectirostris Zaddach, 1844: 1. [maris baltici in littore Gedanensi] 

= Palaemon Leachii Bell, 1851 [in Bell, 1844-1853]: 307; unnumbered text figure. [Poole Harbour] 

? = Palaemon imbellis Fischer, 1872: 422. [Bassin d’Arcachon, dans les Zostera (1-2 brasses)] 

= P.[alaemon] rectirostris var. octodentata Neumann, 1878: 37. [Palam de Mallorka] 

= Leander rectirostris var. transitans Czerniavsky, 1884: 41. [sinus Sevastopol, portu Streletzkaja ad 

littor.] 

? = Leander Brandti Czerniavsky, 1884: 43; Plate 3, fig. 8. [Sevastopol] 

= Palaemon foliirostris Phan, 1971: 35; Figs 1-5. [Caspian Sea] 

Palaemon affinis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= P.[alemon] affinis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 391. [Nouvelle-Zélande] 

= P.[alemon] Quoianus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 393. [Nouvelle- 

Zélande] 

Palaemon capensis (De Man in Weber, 1897) 

= Leander capensis De Man in Weber, 1897: 174; Plate 15, figs 3-3g. [Cap-Colonie, im Knysna-Fluss, in 

süssem Wasser] 

Palaemon concinnus Dana, 1852a 

= Palæmon concinnus Dana, 1852a: 26. [archipelago ”Viti”] 

= Palæmon exilimanus Dana, 1852a: 26. [archipelago ”Viti”] 

= Leander longicarpus Stimpson, 1860a: 40. [Portu ”Hong Kong” Sinensi] 

= Palæmon lagdaoensis Blanco, 1939b: 167; Plate 1. [Cagayan River] 

Palaemon curvirostris Nguyên, 1992: 34; Figs 7-8, 12 (part). [small fishery centre near Thanh An, a village 

20 km north-west of Vung Tàu ( = Cap St. Jacques)] 

Palaemon debilis Dana, 1852a 

= Palæmon debilis Dana, 1852a: 26. [insulas Hawaiensibus] 

= Palæmon debilis var. α Dana, 1852a: 26. [insulas Hawaiensibus] 

= Palæmon debilis var. β, attenuatus Dana, 1852a: 26. [insulas Hawaiensibus] 

= Leander gardineri Borradaile, 1901: 98. [fresh-water kuli in Ekasdu, Miladummadulu Atoll] 

= Leander beauforti Roux, 1923: 18; Figs 1-2. [Céram: Kairatoe, eau saumâtre] 

= Palaemonetes pacificus Gurney, 1939c: 145; Plates 5-6. [brackish canal at Charanka in the island of 

Saipan, Marianne Islands] 

Palaemon dolospinus Walker & Poore, 2003 

= Palaemon dolospina Walker & Poore, 2003: 251; Figs 1E, 2F-J, 3C, 6-7. [Margate Beach, North West 

Bay, Tasmania, in Zostera muelleri and Heterozostera tasmanica on sand, 0.2-1 m] 

Palaemon elegans Rathke, 1837 

= Palaemon elegans Rathke, 1837: 370; Plate 4, fig. 5. [bei Sudagh und am Cap Porthenion] 

= Palæmon minans Norman, 1861: 279; Plate 14, figs 1-2. [Guernsey] 

= Pal.[aemonella] gracilis Paul’son, 1875: 117; Plate 17, figs 6-6f. [Red Sea] 

= Leander squilla var. prototypa Czerniavsky, 1884: 50. [sinus Sevatopol, in portu Streletzk, ad littor.] 

= Leander squilla var. brevidigitata Czerniavsky, 1884: 51. [promontorium Santa, prope Theodosiam]


= Leander squilla var. intermedia De Man, 1915b: 127; Plate 10, figs 10-1v. [Oosterschelde, near Yerseke; 

Goesche Sas (Sluis van Goes), Oosterschelde; off Kats, Oosterschelde; Oosterschelde; Plymouth; 

Brixham, Tor Bay; Le Portel, Straits of Calais; Straits of Calais] 

Palaemon floridanus Chace, 1942b: 80; Plate 23. [tide pool or sinkhole in back of beach at south end of 

Captiva Island, Florida] 

Palaemon gladiator Holthuis, 1950b: 96. [Academy Bay, Indefatigable Island, Galápagos Archipelago] 

Palaemon gracilis (Smith, 1871) 

= Leander gracilis Smith, 1871: 97. [”Estero at Realejo”, west coast of Nicaragua] 

Palaemon gravieri (Yu, 1930b) 

= Leander Gravieri Yu, 1930b: 555, 564; Fig. 3. [Tientsin, Tangkou] 

Palaemon guangdongensis Liu, Liang & Yan, 1990a: 234; Fig. 27. [Guanghai, Taishan County, Guangdong] 

Palaemon hancocki Holthuis, 1950b: 97. [Guayas River, Ecuador] 

Palaemon intermedius (Stimpson, 1860a) 

= Leander intermedius Stimpson, 1860a: 41. [in portu Jacksoniensi Australiae, prof. 2 org] 

Palaemon khori De Grave & Al-Maslamani, 2006: 38; Figs 1-4. [creek on the northern side of Al-Khor, east 

coast of Qatar, 51°33'30''E 25°41'30''N] 

Palaemon litoreus (McCulloch, 1909) 

= Leander litoreus McCulloch, 1909: 308; Fig. 16. [rock-pools on the coast near Sydney] 

Palaemon longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= P.[alemon] longirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 392. [l’embouchure 

de la Garonne, près de Bordeaux] 

= Astacus albescens Pennant, 1812: 25. [coasts of Kent] 

= Palaemon Edwardsii Heller, 1863a: 265. [Mittelmeer, Corsika] 

= Palaemon longipes Fischer, 1872: 421. [nomen nudum] 

= Leander edwardsi var. brevidigitata forma similis Czerniavsky, 1884: 54. [sinus Sevastopol; promontorium 

Santa, prope Theodosiam] 

= Leander edwardsi var. brevidigitata forma simplicior Czerniavsky, 1884: 55. [sinus Sevastopol; promontorium 

Santa] 

= Leander edwardsi forma intermedia Czerniavsky, 1884: 55. [littus Tauriae] 

= Leander edwardsi var. prototypa Czerniavsky, 1884: 55. [sinus Sevastopol] 

= Leander longirostris var. robusta De Man, 1925b: 4. [mouth of the river Adour; Hollandsch Diep] 

= Palaemon garciacidi Zariquiey Álvarez, 1968: 167; Figs 69c, 70. [río Gaudalquivir, Ilegando hasta 

Sevilla; río Francolí, en Tarragona; río Fluviá, en el Golfo de Rosas; Portugal] 

Palaemon macrodactylus Rathbun, 1902b 

= Palæmon macrodactylus Rathbun, 1902b: 52; Fig. 24. [Aomori, Rikuoku] 

= Leander serrifer var. longidactylus Yu, 1930b: 570 (partim); Fig. 4B'-C' (nec text figures). [Yangmatoa, 

Peitaiho, Tangkou, Chefoo] 

Palaemon maculatus (Thallwitz, 1892) 

= Leander maculatus Thallwitz, 1892: 19; Plate 1, fig. 4. [Ogowé, West-Afrika] 

Palaemon miyadii (Kubo, 1938b) 

= Leander miyadii Kubo, 1938b: 538; Fig. 1A, C-F, M-P. [Liao-yang] 

Palaemon northropi (Rankin, 1898) 

= Leander northropi Rankin, 1898: 245; Plate 30, fig. 4. [Nassau, New Providence] 

= Palæmon brachylabis Rathbun, 1900: 154; Plate 8, fig. 10. [Rio Parahyba do Norte, on mangroves] 

Palaemon ogasawaraensis Kato & Takeda, 1981: 101; Figs 2-5. [stream mouth, a stream flowing into Buta 

beach (Minami-fukurosawa), Chichi-jimi Island, Ogasawara Islands] 

Palaemon ortmanni Rathbun, 1902b 

= Palæmon ortmanni Rathbun, 1902b: 53. [nomen novum for Leander longipes Ortmann, 1890] 

= Leander longipes Ortmann, 1890: 519; Plate 37, fig. 13; nec Olivier, 1811. [Japan, Sagamibai] 

Palaemon pacificus (Stimpson, 1860a) 

= Leander pacificus Stimpson, 1860a: 40. [ad insulas ”Hong Kong” et ”Hawaii” etiam in portu ”Simoda”] 

? = Leander okiensis Kamita, 1950: 216; Fig. 2. [innermost coast of Suwa Bay]


Palaemon paivai Fausto Filho, 1967 

= Palaemon (Palaemon) paivai Fausto Filho, 1967: 19; Figs 1-12. [Praia de Meireles, Fortaleza, Ceará, 

Brasil] 

Palaemon pandaliformis (Stimpson, 1871) 

= Leander pandaliformis Stimpson, 1871: 130. [found by Prof. T. Gill in fresh-water streams near the 

sea, either in Barbados or Trinidad, he is uncertain which] 

? = Leander Potitinga Müller, 1892: 181. [nomen nudum] 

= Palæmonetes cubensis Hay, 1903: 433; Fig. 3. [Palacio, Cuba] 

Palaemon paucidens De Haan, 1844 [in De Haan, 1833-1850] 

= Palemon paucidens De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 11 (1844); 170 (1849). [Japan; 

lectotype designation by Yamaguchi & Baba, 1993] 

Palaemon peringueyi (Stebbing, 1915) 

= Leander peringueyi Stebbing, 1915: 75; Plate 17. [33°49'S 25°56'E] 

= Leander gilchristi Stebbing, 1915: 76; Plate 18. [East London wood] 

Palaemon peruanus Holthuis, 1950b: 97. [La Palisada, near Tumbes, N. Peru] 

Palaemon powelli Ashelby & De Grave, 2009: 33; Figs 3-8. [Oguck, ½ km from entrance to Bonny River, 

Niger Delta, Nigeria, 04°39'40''N 07°09'20''W] 

Palaemon ritteri Holmes, 1895 

= Palæmon Ritteri Holmes, 1895: 579; Plate 21, figs 29-35. [San Diego] 

Palaemon semmelinkii (De Man, 1881) 

= Leander semmelinkii De Man, 1881: 137. [on the road of Makassar, Celebes] 

Palaemon serenus Heller, 1862a 

= Leander serenus Heller, 1862a: 527. [Sidney; fully described in Heller, 1865] 

Palaemon serratus (Pennant, 1777) 

= Astacus Serratus Pennant, 1777: 15; Plate 16, fig. 28. [Type locality not indicated] 

= M.[elicerta] Triliana Risso, 1816: 111; Plate 3, fig. 6. [environs de Nice, dans les moyennes profondeurs] 

= Cancer captivus Nardo, 1847: 6. [golfo tra gli Asprei] 

= Leander Latreillianus forma gigantea Czerniavsky, 1884: 44. [nomen novum for Melicerta Triliana 

Risso, 1816] 

= Leander Latreillianus forma typica Czerniavsky, 1884: 45. [sinus Sevastopol, portu Streletzkaja ad 

litt.; sinus Novorossijsk, 6-10 m] 

= Leander Latreillianus forma transitans Czerniavsky, 1884: 46. [sinus Suchum, 1-1.5 m] 

= Leander Latreillianus var. intermedia Czerniavsky, 1884: 47. [Sevastopol] 

= Leander Latreillianus var. aberrans Czerniavsky, 1884: 47. [lacus salinus prope Sevastopol] 

= Palaemon rostratus Gimenez, 1922: 141. [nomen nudum] 

= Palemon Oratelli Monod, 1931: 133. [nomen nudum] 

Palaemon serrifer (Stimpson, 1860a) 

= Leander serrifer Stimpson, 1860a: 41. [Portu ”Hong Kong” et sinibus insulae ”Ousima”, littoralis] 

= Leander Fagei Yu, 1930b: 555, 561; Fig. 2. [Péninsule de Shantong] 

= Leander serrifer var. longidactylus Yu, 1930b: 570 (partim); text figures (nec Fig. 4B’-C’). [Yangmatoa, 

Peitaiho, Tangkou, Chefoo] 

Palaemon sewelli (Kemp, 1925) 

= Leander sewelli Kemp, 1925: 299; Figs 9-10. [off Betim Point, opposite Nova Goa, Portuguese India] 

Palaemon tenuidactylus Liu, Liang & Yan, 1990a: 238; Fig. 31; Plate 1, Figs 2-4. [Changjiang (Yangtze) 

River estuary] 

Palaemon vicinus Ashelby, 2009: 826; Figs 1-40. [Boa Vista, W coast, NW coast of Ilhéu de Sal rei, 16°10'N 

22°58'E, intertidal (rockpool) and shallow sublittoral] 

Palaemon xiphias Risso, 1816 

= P.[alemon] Xiphias Risso, 1816: 102. [environs de Nice, dans les zostères du rivage] 

? = P.[alemon] Trisetaceus Risso, 1816: 103. [environs de Nice, dans les fucus, loin du rivage] 

= Palaemon crenulatus Risso, 1827: 60. [environs de Nice, région des algues] 

= Palaemon Sogiontii Monod, 1931: 133. [nomen nudum]


Palaemon yamashitai Fujino & Miyake, 1970a 

= Palaemon (Palaemon) yamashitai Fujino & Miyake, 1970a: 244; Figs 1-2. [33°14.3'N 122°16.9'E, 26 m] 

Palaemonetes Heller, 1869 

= Palaemonetes Heller, 1869 (type species Palaemon Varians Leach, 1813 [in Leach, 1813-1814], by 

monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology in 


= Palaemonopsis Stimpson, 1871 (type species Palaemonopsis carolinus Stimpson, 1871 (junior subjective 

synonym of P.[alæmon] vulgaris Say, 1818), designated by Holthuis, 1955b, gender feminine) 

= Allocaris Sollaud, 1911b (type species Allocaris sinensis Sollaud, 1911b, by monotypy, gender feminine) 

= Alaocaris Holthuis, 1949c (type species Palaemonetes antrorum Benedict, 1896, by monotypy, gender 

feminine) 

Palaemonetes africanus Balss, 1916: 27; Fig. 9. [Nigeria, Old Calabar] 

Palaemonetes antennarius (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= P.[alemon] antennarius H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 391. [la mer 

Adriatique] 

= Palaemon lacustris von Martens, 1857: 183; Plate 10, Figs 1-9. [Albanersee] 

= P.[elias] migratorius Heller, 1862b: 409; Plate 2, fig. 35. [aus dem Adriatischen Meere, aus dem 

Gardasee und aus ägypten] 

= Palaemonetes varians var. thermajophilus Garbini, 1879: 187. [nomen nudum] 

= Palaemon varians var. termaiophilus Garbini, 1881: 108. [nomen dubium] 

= Palaemonetes varians var. macrogenitor Boas, 1889: 800; Figs (partim) 2 (N1-5), 3 (N1-5), 4 (N1-5); 

Plate 23, figs 2, 4, 6. [im südlichen Europa, in den Mittelmeerländern] 

Palaemonetes antrorum Benedict, 1896 

= Palæmonetes antrorum Benedict, 1896: 615. [artesian well, San Marcos, Texas] 

Palaemonetes argentinus Nobili, 1901a: 3. [Buenos-Ayres; Tigre, Rio de la Plata] 

= Leander brasiliensis Ortmann, 1890: 524; Plate 37, fig. 16. [Rio Grande do Sul] 

Palaemonetes atrinubes Bray, 1976: 76; Figs 23-42. [Canning Bridge, lower Swan River] 

Palaemonetes australis Dakin, 1915: 572; Plate 1. [Upper Swan Bridge; neotype designation by Bray, 1976] 

Palaemonetes camranhi Nguyên, 1997: 85; Fig. 1. [Ba Ngoi, Cam Ranh, a coastal district 50 km south of 

Nha Trang, 11°54'N 190°9'E] 

Palaemonetes carteri Gordon, 1935: 324; Fig. 12. [creek near Camp 1, Upper Cuyuni] 

Palaemonetes cummingi Chace, 1954 

= Palaemonetes (Palaemonetes) cummingi Chace, 1954: 319; Fig. 2. [Squirrel Chimney, Alachua County, 

Florida] 

Palaemonetes hiltoni Schmitt, 1921: 36; Plate 12, fig. 5. [San Pedro, California] 

Palaemonetes hobbsi Strenth, 1994: 292; Fig. 1. [Nacimiento del Río Mante, 8 km southwest of Ciudad 

Mante, Tamaulipas, México] 

Palaemonetes intermedius Holthuis, 1949c 

= Palaemonetes (Palaemonetes) intermedius Holthuis, 1949c: 94; Fig. 2j-l. [Box Iron Bay, Chincoteague 

Bay, Virginia; see Holthuis, 1952b] 

Palaemonetes ivonicus Holthuis, 1950b: 98. [Ivon at the Beni River, N. Bolivia] 

Palaemonetes kadiakensis Rathbun, 1902a 

= Palæmonetes kadiakensis Rathbun, 1902a: 903. [Kadiak Island, Alaska, under stones at low water] 

Palaemonetes karukera Carvacho, 1979 

= Palaemonetes (Palaemonetes) karukera Carvacho, 1979: 449; Figs 2-3. [Rivière Lézarde, 300 m de 

l’embouchure] 

Palaemonetes lindsayi Villalobos Figueroa & H.H.Jr. Hobbs, 1974 

= Palaemonetes (Palaemonetes) lindsayi Villalobos Figueroa & H.H.Jr. Hobbs, 1974: 9; Figs 4-7, 8e. 

[springs and canals in the watershed of Laguna de la Media Luna, 4.8 miles south of Río Verde (on 

the orad to Pedro Montoya) and 2.5 miles to the west on a road to the Mina El Refugio, San Luis 

Potosí, Mexico]


Palaemonetes mercedae Pereira S., 1986 

= Palaemonetes (Palaemonetes) mercedae Pereira S., 1986: 209; Fig. 13. [Atabapo River at Chamuchina, 

Territorio Federal Amazonas, Venezuela, 3°20'N 67°29'W] 

Palaemonetes mesogenitor Sollaud, 1912 

= Palæmonetes varians mesogenitor Sollaud, 1912: 1270. [Gabès, dans l’oued qui arrose cet oasis] 

= Palaemonetes punicus Sollaud, 1923: 589; Figs 23-24. [Tunisie] 

Palaemonetes mesopotamicus Pesta, 1913 

= Palaemonetes mesopotamicus Pesta, 1913: 23; Figs 6-7, 9. [Hsitsche (= Heseke) am Khabur] 

Palaemonetes mexicanus Strenth, 1976: 7; Fig. 3. [small river 15 road miles (24 km) west of Ciudad Valles, 

San Luis Potosí, Mexico] 

Palaemonetes octaviae Chace, 1972 

= Palaemonetes (Palaemonetes) octaviae Chace, 1972: 22; Figs 3-4. [Guadeloupe, Pointe-à-Pitre, between 

Îlet à Monroux and Îlet Rat, sandy mudflats] 

Palaemonetes paludosus (Gibbes, 1850) 

= Hippolyte caroliniana Gibbes, 1848: 16. [nomen nudum] 

= Hippolyte paludosa Gibbes, 1850: 197. [in fresh water ponds, St. Andrew’s Parish, South Carolina] 

= Palaemonetes exilipes Stimpson, 1871: 130. [freshwater at Somerville, South Carolina] 

Palaemonetes pugio Holthuis, 1949c 

= Palaemonetes (Palaemonetes) pugio Holthuis, 1949c: 95; Fig. 2m-o. [Lagoon near Cove Point Light, 

Chesapeake Bay; see Holthuis, 1952b] 

Palaemonetes schmitti Holthuis, 1950b: 99. [Upper Chamber, east side, Miraflores Locks, Panama Canal] 

Palaemonetes sinensis (Sollaud, 1911b) 

= Allocaris sinensis Sollaud, 1911b: 50; Figs 1-2. [prés de Pèkin] 

= Palaemonetes chankensis Buldovsky, 1933: 43; Plate 1; Plate 2, figs 12-16, 18-19. [First river (Jerick) to 

the south of the Village of Astrachanka, lake Khanka, Far East] 

= Palaemonetes venephicus Birstein & Vinogradov, 1934: 45; Fig. 3. [in der Nähe von Daubiche (Bassin 

des Flusses Ussuri)] 

Palaemonetes suttkusi Smalley, 1964: 229; Fig. 1. [Rio Grande drainage, 24 miles north of Monclova, 

Coahuila Province, Mexico] 

Palaemonetes texanus Strenth, 1976: 5; Fig. 2. [San Marcos River, within the city limits of San Marcos, 

Hays County, Texas] 

Palaemonetes turcorum Holthuis, 1961c: 14; Figs 6-8. [Sakarya River near the bridge of the highway from Ankara 

to Eskişehir, about halfway between Sivrihisar and Haymana, about 95 km S.W. of Ankara, Turkey] 

Palaemonetes varians (Leach, 1813 [in Leach, 1813-1814]) 

= Palaemon Varians Leach, 1813 [in Leach, 1813-1814]: 401. [Devonshire and Glamorgan coasts] 

= Palaemon variabilis Bouchard-Chanteraux, 1829: 15. [les fossés d’eau saumâtre de Capécure] 

= Palaemonetes varians var. microgenitor Boas, 1889: 800; Figs (partim) 2 (A1-5), 3 (A1-5), 4 (A1-5); 

Plate 23, figs 1, 3, 5. [nördliches Europa] 

= Palaemonetes varians occidentalis Sollaud, 1923: 547; Figs 12-13; Plate 18, figs 5-7. [eaux saumâtres le 

long des côtes occidentales de l’Europe et de l’Afrique] 

Palaemonetes vulgaris (Say, 1818) 

= P.[alæmon] vulgaris Say, 1818: 248. [bays and estuaries] 

= Palaemonetes carolinus Stimpson, 1871: 129. [harbor of Charleston, South Carolina; Great Egg Harbor, 

New Jersey; harbor of Beaufort, North Carolina] 

Palaemonetes zariquieyi Sollaud, 1938 

= Palæmonetes Zariquieyi Sollaud, 1939: 638; Fig. 1c-d. [l’Albufera (lac) de Valence; les lagunes de la 

Almenara, les canaux d’Amposta (delta de l’Ebre), une petit étang littoral au cap Salou (près de 

Tarragone)] 

Pseudopalaemon Sollaud, 1911c 

= Pseudopalaemon Sollaud, 1911c (type species Pseudopalaemon Bouvieri Sollaud, 1911c, by monotypy, 


1957)


Pseudopalaemon amazonensis Ramos-Porto, 1979: 693. [arredores de Manaus, Amazonas] 

Pseudopalaemon bouvieri Sollaud, 1911c 

= Pseudopalæmon Bouvieri Sollaud, 1911c: 12; Figs 1, 2a. [environs de Montévidéo] 

= Pseudopalaemon Iheringi Sollaud, 1911d: 285; Fig. 1. [Arroyo del Bellaco, Brésil] 

Pseudopalaemon chryseus Kensley & Walker, 1982: 16; Figs 18-19. [Rio Tarumazinho] 

Pseudopalaemon funchiae García-Dávila & Magalhães, 2004: 677; Figs 28-38. [Loreto, Província de Requena: 

distrito Jenaro Herrera, Jenaro Herrera, cocha Carahuaite, afluente da cocha Supay, bacia do 

rio Ucayali] 

Pseudopalaemon gouldingi Kensley & Walker, 1982: 18; Figs 20-21. [Rio Negro, Ilha de Cumuru, Lago da 

Ilha] 

Pseudopalaemon iquitoensis García-Dávila & Magalhães, 2004: 679; Figs 39-50. [Província de Maynas, 

distrito de Iquitos, Iquitos, quebrada Paujil, km 22 da Estrada Iquitos-Nauta, afluente do rio 

Itaya] 

Pseudopalaemon nigramnis Kensley & Walker, 1982: 20; Figs 22-23. [Rio Marania, Rio Negro basin] 

Rhopalaemon Ashelby & De Grave, 2010 

= Rhopalaemon Ashelby & De Grave, 2010 (type species Leander belindae Kemp, 1925, by original 


Rhopalaemon belindae (Kemp, 1925) 

= Leander belindae Kemp, 1925: 308; Figs 12-13. [Kilakarai, Gulf of Manaar] 

Tenuipedium Wowor & Ng, 2010a 

= Tenuipedium Wowor & Ng, 2010a (type species Macrobrachium palaemonoides Holthuis, 1950a, by 

original designation and monotypy, gender neuter) 

Tenuipedium palaemonoides (Holthuis, 1950a) 

= Macrobrachium palaemonoides Holthuis, 1950a: 136; Fig. 31. [Laut Tawar, Laulo Lake, N. Simalur, off 

west coast of Sumatra] 

Troglindicus Sankolli & Shenoy, 1979 

= Troglindicus Sankolli & Shenoy, 1979 (type species Troglindicus phreaticus Sankolli & Shenoy, 1979, 


Troglindicus phreaticus Sankolli & Shenoy, 1979: 86; Figs 1-2. [northernmost old freshwater well (13 th 

century?) in the fort area of Ratnagiri, west coast of India] 

Troglocubanus Holthuis, 1949c 

= Troglocubanus Holthuis, 1949c (type species Palæmonetes eigenmanni Hay, 1903, by original designation, 


Troglocubanus calcis (Rathbun, 1912b) 

= Palaemonetes calcis Rathbun, 1912b: 451; Plate 1, figs 1-5. [Cuba, pool in a cave between Madruga 

and Aguacate] 

Troglocubanus eigenmanni (Hay, 1903) 

= Palæmonetes eigenmanni Hay, 1903: 431; Fig. 2. [cavern at Ashton, Cuba] 

Troglocubanus gibarensis (Chace, 1943) 

= Palaemonetes gibarensis Chace, 1943: 28; Plate 7. [a well, 29 yards deep, entering an underground 

stream, Aguada del Montañes, en el Jobal, Barrio de Cupeysillo, Termino de Gibara, Oriente province, 

Cuba] 

Troglocubanus inermis (Chace, 1943) 

= Palaemonetes inermis Chace, 1943: 26; Plate 6. [pool in a cave situated on the calzada from Madruga 

to Aguacate, about one kilometer from the junction with the calzada from Madruga to Matanzas, 

Havana Province, Cuba] 

Troglocubanus jamaicensis Holthuis, 1963a: 67; Fig. 3. [stream in limestone cave near Lucky Hill Cooperative 

farm, near Goshen, Jamaica]


Troglomexicanus Villalobos, Alvarez & Iliffe, 1999 

= Troglomexicanus Villalobos, Alvarez & Iliffe, 1999 (type species Troglocubanus perezfarfantae Villalobos 

Figueroa, 1971, by original designation, gender masculine) 

Troglomexicanus huastecae Villalobos, Alvarez & Iliffe, 1999: 117; Figs 4-5. [Manantial de San Rafael de los 

Castros, 22°46'N 99°03'W, Municipio de Ciudad Mante, San Luis Potosí] 

Troglomexicanus perezfarfanteae (Villalobos Figueroa, 1971) 

= Troglocubanus perezfarfanteae Villalobos Figueroa, 1971: 1; Figs 1-23, 24 (partim). [Sótano de Tinaja, 

11.7 km E. NE. Valles, San Luis Potosí, México] 

Troglomexicanus tamaulipasensis Villalobos, Alvarez & Iliffe, 1999: 113; Figs 2-3. [Cueva del Nacimiento 

del Río Frío, 23°01'N 99°04'W, Municipio de Gómez Farís, Tamaulipas] 

Urocaridella Borradaile, 1915b 

= Urocaridella Borradaile, 1915b (type species Urocaridella gracilis Borradaile, 1915b (invalid senior 

synonym of Urocaridella urocaridella (Holthuis, 1950a), by monotypy, gender feminine) 

Urocaridella antonbruunii (Bruce, 1967b) 

= Periclimenes antonbruunii Bruce, 1967b: 45; Figs 19-22. [Pamanzi Island reef, Dzaoudzi, Ile Mayotte, 

Archipel des Comores] 

= Leandrites cyrtorhynchus Fujino & Miyake, 1969a: 143; Figs 1-3. [Tô-shima Island, Tanabe Bay, Wakayama 

Prefecture] 

= Leandrites longipes Liu, Liang & Yan, 1990b: 127; Fig. 24. [Sanya, Hainan Isl., China; additional 

figure in Liu, Liang & Yan, 1990a] 

Urocaridella pulchella Yokes & Galil, 2006: 749; Figs 2A-B, 3. [Turkey, Güvercin Ada, Kaş, 12 m] 

Urocaridella urocaridella (Holthuis, 1950a) 

= Leander urocaridella Holthuis, 1950a: 28. [nomen novum for Urocaridella gracilis Borradaile, 1915b] 

= Urocaridella gracilis Borradaile, 1915b: 210. [Maldive Islands] 

Urocaridella vestigialis Chace & Bruce, 1993: 45; Fig. 17. [Indonesia, Selat Butung, Celebes, 4°31'40"S 

122°49'42"E, 68 m] 

Subfamily PoNToNIINAe Kingsley, 1879 

Allopontonia Bruce, 1972b 

= Allopontonia Bruce, 1972b (type species Allopontonia iani Bruce, 1972b, by original designation and 

monotypy, gender feminine; a junior synonym of Anchistia Brockii De Man, 1888b) 

Allopontonia brockii (De Man, 1888b) 

= Anchistia Brockii De Man, 1888b: 548; Plate 22a, figs 3-3d. [Amboina] 

? = Periclimenes (Periclimenes) signatus Kemp, 1925: 322; Figs 16-17. [Andamans] 

= Allopontonia iani Bruce, 1972b: 7; Figs 1-4. [Zanzibar Harbour, 6°09.5'S 39°10.2'E] 

= Periclimenes priodactylus Bruce, 1992c: 53; Figs 1, 7-11. [Lizard island, Queensland, 14°40'S 145°28'E, 

northern entrance to Blue Lagoon, 3.5 m] 

Allopontonia alastairi Bruce, 2010a: 33; Fig. 1. [North-western Australia, Imperieuse L23 transect, 

18°25'31''S 120°0555''E to 18°26'12''S 120°06'30''E, 105-103 m] 

Altopontonia Bruce, 1990c 

= Altopontonia Bruce, 1990c (type species Altopontonia disparostris Bruce, 1990c, by original designation 


Altopontonia disparostris Bruce, 1990c: 192; Figs 25-33, 39k. [New Caledonia, 23°03'S 167°19'E, 503 m] 

Amphipontonia Bruce, 1991a 

= Amphipontonia Bruce, 1991a (type species Amphipontonia kanak Bruce, 1991a, by original designation 


Amphipontonia kanak Bruce, 1991a: 382; Figs 58-63. [Loyalty Islands]


Anapontonia Bruce, 1966c 

= Anapontonia Bruce, 1966c (type species Anapontonia denticauda Bruce, 1966c, by monotypy, gender 

feminine) 

= Anapontonia Bruce, 1967b (type species Anapontonia denticauda Bruce, 1967b, by monotypy, gender 

feminine) 

Anapontonia denticauda Bruce, 1966c: 596. [Pange Reef, Zanzibar; fully described in Bruce, 1967b] 

Anchiopontonia Bruce, 1992d 

= Anchiopontonia Bruce, 1992d (type species Pontonia hurii Holthuis, 1981, by original designation 


Anchiopontonia hurii (Holthuis, 1981) 

= Pontonia hurii Holthuis, 1981: 796; Fig. 4. [Arno Atoll, Marshall Islands, from mantle cavity of rock 

clam, Spondylus spec.] 

= Pontonia hurii Holthuis, 1953b: 57. [nomen nudum] 

Anchistus Borradaile, 1898 

= Anchistus Borradaile, 1898 (type species Harpilius Miersi De Man, 1888a, by original designation, 

gender masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 

in 1957) 

= Tridacnocaris Nobili, 1899 (nomen novum for Anchistus Borradaile, 1898, gender feminine) 

= Marygrande Pesta, 1911 (type species Marygrande mirabilis Pesta, 1911, by monotypy, gender 

feminine) 

= Ensiger Borradaile, 1915b (type species Anchistia aurantiaca Dana, 1852a (junior subjective synonym 

of Cancer custos Forskål, 1775), being the first species subsequently placed in this subgenus, 

which was described without any nominal species, first placed in Ensiger by Borradaile, 1917) 

Anchistus australis Bruce, 1977a 

= Anchistus australis forma typica Bruce, 1977a: 56; Figs 7-9. [Capre Cay, Swain’s Reefs; unavailable 


= Anchistus australis forma dendricauda Bruce, 1977a: 62; Fig. 10. [West Cay, Diamond Islets; unavailable 


Anchistus custoides Bruce, 1977a: 50; Figs 4-6. [north-west end, Gillett Cay, Great Barrier Reef, 21°43'S 

152°25'E, in bivalve mollusk Atrina vexillum according to Springthorpe & Lowry, 1994] 

Anchistus custos (Forskål, 1775) 

= Cancer custos Forskål, 1775: 94. [Lohajæ, intra Pinna nigras] 

= Pontonia inflata H. Milne Edwards, 1840 [in H. Milne Edwards, 1834-1840]: 633. [latinization of the 

French vernacular name Pontonie enflée H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 

360. [Ceylan et sur les côtes de Vanicoso] 

= Anchistia aurantiaca Dana, 1852a: 25. [archipelago Viti] 

= Harpilius inermis Miers, 1884b: 291; Plate 32, fig. B. [Port Molle, in the interior of a shell of a species 

of Pinna] 

= Pontonia pinnae Ortmann, 1894: 16; Plate 1, Figs 3-3n; nec Lockington, 1878a. [Dar-es-Salaam, 

Chokirbank, in den Schalen einer Pinna-Art] 

= Marygrande mirabilis Pesta, 1911: 573; Figs 1-5 (partim). [Samoa] 

= Pontonia spinax Dawydoff, 1952: 136. [nomen nudum] 

Anchistus demani Kemp, 1922: 256; Figs 86-88. [Aberdeen, Port Blair, Andamans, low tide] 

Anchistus gravieri Kemp, 1922: 252; Figs 82-84. [Vanikoro, Santa Cruz Islands, Polynesia] 

Anchistus miersi (De Man, 1888a) 

= Harpilius Miersi De Man, 1888a: 274; Plate 17, figs 6-10. [Elphinstone Island] 

= Marygrande mirabilis Pesta, 1911: 571, Figs 1-5 (partim). [Samoa] 

Anchistus pectinis Kemp, 1925: 327; Figs 19-20. [Octavia Bay, Nancowry Harbor, Nicobars] 

= Anchistus misakiensis Yokoya, 1936: 136; Fig. 5. [vicinity of the Misaki Marine Biological Station, 

inside of a bivalve, Amusium japonicum]


Ancylomenes okuno & Bruce, 2010 

= Ancylomenes Okuno & Bruce, 2010 (type species Periclimenes venustus Bruce, 1989a, by original 


Ancylomenes adularans (Bruce, 2003a) 

= Periclimenes adularans Bruce, 2003a: 116; Fig. 2. [Cape Flattery Service Jetty, 2 m] 

Ancylomenes aesopius (Spence Bate, 1863) 

= Anchistia æsopia Spence Bate, 1863: 502; Plate 41, Figs 5-5z. [Gulf of St. Vincent] 

Ancylomenes amirantei (Bruce, 2007a) 

= Periclimenes amirantei Bruce, 2007a: 52; Figs 1-4. [Amirante Islands, Seychelle Islands, station E.15, 

70 m] 

Ancylomenes aqabai (Bruce, 2008a) 

= Periclimenes aqabai Bruce, 2008a: 27; Figs 1-4. [Aqaba, Jordan, 35-45 m] 

Ancylomenes grandidens (Bruce, 2005b) 

= Periclimenes grandidens Bruce, 2005b: 350; Figs 8-10, 22C. [Loloata Island, 16 m] 

Ancylomenes holthuisi (Bruce, 1969a) 

= Periclimenes holthuisi Bruce, 1969a: 258. [Lung Ha Wan, N.T., Hong Kong, 22°18.5'N 114°18.2'E, 

2 fms] (Fig. 37) 

Ancylomenes kobayashii (Okuno & Nomura, 2002) 

= Periclimenes kobayashii Okuno & Nomura, 2002: 84; Figs 1-5. [off Akazawa, Ito, Izu Peninsula, Honshu, 

Japan, 34°51.2'N 139°05.5'E, 32 m] 

Ancylomenes kuboi Bruce, 2010b: 170; Figs 1-2. [Northwest Shelf, Western Australia, 19°05.3'S 118°53.8'E, 

80 m] 

Ancylomenes longicarpus (Bruce & Svoboda, 1983) 

= Periclimenes longicarpus Bruce & Svoboda, 1983: 13; Figs 4-8. [Aqaba, Jordan, 15 m] 

Ancylomenes lucasi Chace, 1937b 

= Periclimenes (Ancylocaris) lucasi Chace, 1937b: 133; Fig. 8. [San Lucas Bay, 22°53'N 109°54'W, 3-9 fms] 

Fig. 37. Ancylomenes holthuisi (Bruce, 1969). Photo by Charles H.J.M. Fransen.


Ancylomenes luteomaculatus Okuno & Bruce, 2010: 88; Figs 1-5, 6A-B. [off Gima Harbor, Kume-jima Island, 

Ruykyu Islands, Japan, 26°19.7'N 126°45.6'E, 32 m, associated with Pachycerianthus sp.] 

Ancylomenes magnificus (Bruce, 1979a) 

= Periclimenes magnificus Bruce, 1979a: 195; Figs 1-5; Plate 1A-C. [Wistari Reef, Heron Island, Capricorn 

Islands, Queensland, Australia, 85-95 feet] 

Ancylomenes okunoi Bruce, 2010b: 172; Fig. 3. [19°05.0'S 118°50.5'E, Northwest Shelf, Western Australia, 

83 m] 

Ancylomenes pedersoni (Chace, 1958) 

= Periclimenes (Periclimenes) pedersoni Chace, 1958: 125; Figs 1-17. [Lyford Cay, New Providence Island, 

Bahamas] 

= Periclimenes (Periclimenes) anthophilus Holthuis & Eibl-Eibesfeldt, 1964: 185; Figs 1-4. [Whalebone 

Bay, Bermuda, near the shore, 2-3 m, on sea anemones; considered valid species by Okuno & Bruce, 

2010] 

Ancylomenes sarasvati (Okuno, 2002) 

= Periclimenes sarasvati Okuno, 2002: 212; Figs 1-4, 6A-B. [off Hateno-hama, Kume-jima Island, 

26°25.9'N 127°46.5'E, 26 m] 

Ancylomenes speciosus (Okuno, 2004) 

= Periclimenes speciosus Okuno, 2004: 866; Figs 1-5, 6A-B. [Hira-ne, Hasama, Tateyama, Boso Peninsula, 

Honshu, Japan, 34°58.6'N 139°47.1'E, 18 m] 

Ancylomenes tenuirostris (Bruce, 1991b) 

= Periclimenes tenuirostris Bruce, 1991b: 247; Figs 13-16. [New Caledonia, 22°35.1'S 166°59.5'E, Grand 

Récif Sud, 82 m] 

Ancylomenes tosaensis (Kubo, 1951) 

= Periclimenes (Ancylocaris) tosaensis Kubo, 1951: 268; Figs 7-8. [off Usa, Takaoka-gun, Kôchi Prefecture] 

Ancylomenes venustus (Bruce, 1989a) 

= Periclimenes venustus Bruce, 1989a: 178. [Nailon] 

= Periclimenes venustus Bruce, 1990d: 230; Figs 1-6. [Table Head, Port Essington, 3 m] 

Anisomenaeus Bruce, 2010c 

= Anisomenaeus Bruce, 2010c (type species Periclimenaeus spinimanus Bruce, 1969b, gender masculine) 

Anisomenaeus spinimanus (Bruce, 1969b) 

= Periclimenaeus spinimanus Bruce, 1969b: 165. [North-western Indian Ocean, off Ras Asir, 11°37'N 

51°27'E – 11°38'N 51°27'E, 37-40 fms] 

Apopontonia Bruce, 1976c 

= Apopontonia Bruce, 1976c (type species Apopontonia falcirostris Bruce, 1976c, by original designation 


Apopontonia falcirostris Bruce, 1976c: 303; Figs 1-5. [NW coast of Madagascar, 12°44.5'S 48°25.2'E, 73 m] 

Araiopontonia Fujino & Miyake, 1970b 

= Araiopontonia Fujino & Miyake, 1970b (type species Araiopontonia odontorhyncha Fujino & Miyake, 

1970b, by original designation and monotypy, gender feminine) 

Araiopontonia odontorhyncha Fujino & Miyake, 1970b: 2; Figs 1-4. [Koniya, Amami O Shima Island, Ryukyu 

Islands, Japan] 

Ascidonia Fransen, 2002 

= Ascidonia Fransen, 2002 (type species P.[ontonia] flavomaculata Heller, 1864, by original designation, 


Ascidonia californiensis (Rathbun, 1902a) 

= Pontonia californiensis Rathbun, 1902a: 902. [Off Santa Cruz Island, California, 30 fms]


Ascidonia flavomaculata (Heller, 1864) 

= P.[ontonia] flavomaculata Heller, 1864: 51. [aus dem Adriatischen Meere, in der Mantelhöhle von 

Phallusia mamillata] 

= Alciope heterochelus Rafinesque, 1814: 24. [Sicile; suppressed under the plenary power or the purposes 

of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 522 in 1958] 

= Pontonia Phallusiae Marion in de Folin & Périer, 1879: 226. [nomen nudum; les eaux de Marseille] 

= Pontonia Diazona Joliet, 1882: 118 (as Pontonia Diazonæ on page 120). [Menton] 

= Pontonia elegans Sarato, 1887: 2. [les eaux de Saint-Jean, près de Nice] [nomen nudum] 

Ascidonia miserabilis (Holthuis, 1951b) 

= Pontonia miserabilis Holthuis, 1951b: 148; Plate 47, figs d-i. [off Vieques Island, Porto Rico, Culebritas 

light house, NE 3/8 E 7¼ miles, 16 fms] 

= Pontonia grayi Rathbun, 1901: 122 (partim; nec fig. 25). [between San Antonio Bridge and San 

Geronimo, San Juan, Puerto Rico] 

Ascidonia pusilla (Holthuis, 1951b) 

= Pontonia pusilla Holthuis, 1951b: 142; Plate 45. [Ecuador, Salango Island, 3 fms] 

= Pontonia spighti Fujino, 1972: 293; Figs 1-3. [Playas del Coco, Pacific coast of Costa Rica, shallow 

water of sublittoral zone] 

Ascidonia quasipusilla (Chace, 1972) 

= Pontonia quasipusilla Chace, 1972: 41; Fig. 10. [Charlotte Point, English Harbour, Antigua Island] 

Balssia Kemp, 1922 

= Balssia Kemp, 1922 (type species Amphipalaemon Gasti Balss, 1921b, by monotypy, gender feminine; 


Balssia antipodarum Bruce, 2004a: 367; Figs 1-3. [North Norfolk Ridge, north of Norfolk Island, 28°54.39'S 

167°41.05'E, 111-115 m] 

Balssia gasti (Balss, 1921b) 

= Amphipalaemon Gasti Balss, 1921b: 524; Figs 1-8. [Golf von Neapel, von Corallium rubrum] 

? = Brachycarpus Antonini Magri, 1926 [lapsus for Brachycarpus Audouini but considered a synonym 

as here listed by L.B. Holthuis (pers. comm.)] 

Balssia noeli Bruce, 1998b: 604; Figs 1-4. [Mediterranean Sea, vicinity of Banyuls] 

Blepharocaris Mitsuhashi & Chan, 2007 

= Blepharocaris Mitsuhashi & Chan, 2007 (type species Blepharocaris panglao Mitsuhashi & Chan, 

2007, by original designation and monotypy, gender feminine) 

Blepharocaris panglao Mitsuhashi & Chan, 2007: 2; Figs 1-4. [9°37.6'N 123°47.3'E, Bingag, Panglao Island, 

Philippines, 20 m] 

Brucecaris Marin & Chan, 2006 

= Brucecaris Marin & Chan, 2006 (type species Periclimenes tenuis Bruce, 1969a, by original designation 


Brucecaris tenuis (Bruce, 1969a) 

= Periclimenes tenuis Bruce, 1969a: 272. [Chukwani, Zanzibar Island, 6°15.1'S 39°12.7'E, 1ft] 

Bruceonia Fransen, 2002 

= Bruceonia Fransen, 2002 (type species Pontonia ardeae Bruce, 1981a, by original designation and 


Bruceonia ardeae (Bruce, 1981a) 

= Pontonia ardeae Bruce, 1981a: 113; Figs 1-8. [Heron Island, Wistari Reef, 23°27.5'S 151°55.0'E, 60-70 

feet] 

Cainonia Bruce, 2005b 

= Cainonia Bruce, 2005b (type species Pontonia medipacifica Edmondson, 1935a, by original designation 



Cainonia medipacifica (Edmondson, 1935a) 

= Pontonia medipacifica Edmondson, 1935a: 6; Fig. 2. [Midway Island, in shallow water] 

Carinopontonia Bruce, 1988a 

= Carinopontonia Bruce, 1988a (type species Carinopontonia paucipes Bruce, 1988a, by original designation 


Carinopontonia paucipes Bruce, 1988a: 1264; Figs 1-3. [19°04.4'S 118°47.55'E, 83 m] 

Chacella Bruce, 1986b 

= Chacella Bruce, 1986b (type species Dasycaris kerstitchi Wicksten, 1983a, by original designation 


Chacella kerstitchi (Wicksten, 1983a) 

= Dasycaris kerstitchi Wicksten, 1983a: 16; Fig. 2. [Punta Doble, San Carlos, Sonora, Mexico, 27°56'N 

111°02'W, 30 m] 

Chernocaris Johnson, 1967 

= Chernocaris Johnson, 1967 (type species Chernocaris placunae Johnson, 1967, by monotypy, gender 

feminine) 

Chernocaris placunae Johnson, 1967: 500; Figs 1-12. [Telok Paku, Singapore, in Placuna sella taken at low 

tide] 

Climeniperaeus Bruce, 1995 

= Climeniperaeus Bruce, 1995 (type species Periclimenaeus truncoides Chace & Bruce, 1993, by original 


= Climeniperaeus Bruce, 1996 (type species Periclimenaeus truncoides Chace & Bruce, 1993, by original 


Climeniperaeus orbitospinatus (Bruce, 1969b) 

= Periclimenaeus orbitospinatus Bruce, 1969b: 160. [Gulf of Carpentaria, northern Australia, 10-15 

fms] 

= Apopontonia tridentata Bruce, 1988a: 1270; Figs 4-7. [19°41.9'S 17°57.15'E, 54 m] 

Climeniperaeus truncatus (Rathbun, 1906) 

= Coralliocaris truncata Rathbun, 1906: 920; Fig. 70; Plate 24, fig. 2. [South coast of Molokai, 23-24 fms] 

Climeniperaeus truncoideus (Chace & Bruce, 1993) 

= Periclimenaeus truncoideus Chace & Bruce, 1993: 93. [2.3 miles [3.7 km] N, 63°W from north point 

of Kai Besar, Kepulauan Kai, Indonesia, 5°36'5'S 132°55.2'E, 90 m] 

Colemonia Bruce, 2005b 

= Colemonia Bruce, 2005b (type species Colemonia litodactylus Bruce, 2005b, by original designation 


Colemonia litodactylus Bruce, 2005b: 335; Figs 1-4, 22A. [Milne Bay, 12 m] 

Conchodytes Peters, 1852 

= Conchodytes Peters, 1852 (type species Conchodytes tridacnae Peters, 1852, designated by Hilgendorf, 

1879, gender masculine; name placed on the Official List of Generic Names in Zoology in 


Conchodytes biunguiculatus (Paul’son, 1875) 

= P.[ontonia] biunguiculata Paul'son, 1875: 111; Plate 15, fig. 1-1n. [Red Sea] 

= Conchodytes kempi Bruce, 1989a: 183; Fig. 3B-E. [Moalboal, in Isognomon isognomon] 

Conchodytes maculatus Bruce, 1989b: 182; Figs 1-6. [Pao Yuan No. 1, 16°34.5'S 121°27.0'E, 40 m] 

Conchodytes meleagrinae Peters, 1852: 594. [Südostküste Africas, Ibo, im 12°S Br., zwischen den Mantellapen 

von Melaegrina margaritifera] 

Conchodytes monodactylus Holthuis, 1952c: 200; Figs 96-98. [Takao, Southern Taiwan (in bivalve mollusk, 

Pinna); Lesser Sunda Island]


Conchodytes nipponensis (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hymenocera niponensis De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 8 (1844); 180 (1849, as 

Pontonia nipponensis). [Japan; incorrect original spelling corrected in Opinion 383 in 1956] 

Conchodytes philippinensis Bruce, 1996: 212; Figs 6, 29b. [Philippines, few meters] 

Conchodytes pteriae Fransen, 1994: 97; Figs 38-58. [N of d’Arros Island, 5°24'S 53°19'E, 45-55 m, in small 

Pteria loveni (Dunker, 1872)] 

Conchodytes tridacnae Peters, 1852: 594. [Südostküste Africas, bei Ibo, im 12° S Br., zwischen den Mantellappen 

von Tridacna squamosa] 

Coralliocaris Stimpson, 1860a 

= Oedipus Dana, 1852a (type species Oedipus superbus Dana, 1852a, designated by Kingsley, 1880, 

gender masculine; invalid junior homonym of Oedipus Berthold, 1827 (Orthoptera), Oedipus Tschudi, 

1838 (Amphibia) and Oedipus Lesson, 180 (Mammalia); name placed on the Official Index of Rejected 

and Invalid Generic Names in Zoology in Opinion 470 in 1957) 

= Coralliocaris Stimpson, 1860a (nomen novum for Oedipus Dana, 1852a, gender feminine; name 


Coralliocaris brevirostris Borradaile, 1898: 386. [Ellice Islands] 

Coralliocaris graminea (Dana, 1852a) 

= Œdipus gramineus Dana, 1852a: 25. [archipelago Viti] 

= Coralliocaris inaequalis Ortmann, 1890: 510; Plate 36, fig. 21. [Japan, Kagoshima and Samoa] 

Coralliocaris labyrintha Mitsuhashi & Takeda, 2008: 16; Figs 1E-F, 6E-F, 11-14. [Urasoko Bay, Ishigaki-jima 

Island, Rykyu Islands, coral reef, 2 m, from Acropora sp.] 

Coralliocaris macrophthalma (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= P.[ontonia] macrophthalma H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 359. [les 

mers d’Asie] 

Coralliocaris nudirostris (Heller, 1861) 

= O.[edipus] nudirostris Heller, 1861: 27. [im rothen Meere] 

= O.[edipus] nudirostris Heller, 1862c: 279; Plate 3, fig. 25. [im rothen Meere] 

= Coralliocaris tahitoei Boone, 1935: 180; Fig. 12; Plate 49. [Venus Point reef, Tahiti, Society Islands] 

Coralliocaris sandyi Mitsuhashi & Takeda, 2008: 11; Figs 1C-D, 6C-D, 8-10. [off Kuro-shima Island, Rykyu 

Islands, 8 m, from Acropora sp.] 

Fig. 38. Coralliocaris viridis Bruce, 1974. Photo by Arthur Anker.


Coralliocaris superba (Dana, 1852a) 

= Œdipus superbus Dana, 1852a: 25. [insula Tongatabu] 

= Oed.[ipus] dentirostris Paul’son, 1875: 112; Plate 14, Figs 7-7d. [Red Sea] 

Coralliocaris taiwanensis Fujino & Miyake, 1972: 92; Figs 1-3. [Herngchuen, Shiangtiau Bay, southern 

Taiwan, 2-5 m] 

= Coralliocaris pavonae Bruce, 1972c: 77; Figs 8-11. [fringing reef at Singatoka, Viti Levu, Fiji] 

Coralliocaris tridens Mitsuhashi, Fujino & Takeda, 2001: 944; Figs 1-3. [Kabira Bay, 24°26'N 124°8'E, Ishigaki 

Island, Ryukyu Islands, southwestern Japan, from tabular coral (Acropora sp)] 

Coralliocaris venusta Kemp, 1922: 274; Figs 100-101. [N.E. Tholayiram Paar, Gulf of Manaar] 

Coralliocaris viridis Bruce, 1974a: 222; Fig. 1A-B. [seaward reefs of Mombasa Island, 4°04.5'S 39°40.5'E] 

(Fig. 38) 

Coutierea Nobili, 1901b 

= Coutièrea Nobili, 1901b (type species Coralliocaris Agassizi Coutière, 1901, by monotypy, gender 

feminine; name placed in the corrected spelling (Coutierea) on the Official List of Generic Names in 


Coutierea agassizi (Coutière, 1901) 

= Coralliocaris Agassizi Coutière, 1901: 115, unnumbered text fig. [Barbades, 94 brasses] 

Crinotonia Marin, 2006 

= Crinotonia Marin, 2006 (type species Crinotonia anastasiae Marin, 2006, by original designation, 


Crinotonia anastasiae Marin, 2006: 322; Figs 1-6. [South China Sea, Vietnam, Nha Thrang Bay, Mun Island, 

10-12 m] 

Crinotonia attenuatus (Bruce, 1971a) 

= Periclimenes attenuatus Bruce, 1971a: 533; Figs 1-5. [Waterhouse Cove, Burukuk, Duke of York Island, 

4°7.3'S 152°27.3'E, 1-2 m] 

Ctenopontonia Bruce, 1979b 

= Ctenopontonia Bruce, 1979b (type species Ctenopontonia cyphastreophila Bruce, 1979b, by original 


Ctenopontonia cyphastreophila Bruce, 1979b: 425; Figs 1-6. [Enewetak Atoll, Marshall Islands, 5-15 m] 

Cuapetes Clark, 1919 

= Cuapetes Clark, 1919 (nomen novum for Falciger Borradaile, 1915b, gender masculine) 

= Falciger Borradaile, 1915b (type species Periclimenes (Falciger) nilandensis Borradaile, 1915b, designated 

by Holthuis, 1955b, gender masculine) 

= Kemponia Bruce, 2004b (type species Anchistia grandis Stimpson, 1860a, by original designation, 


Cuapetes agag (Kemp, 1922) 

= Periclimenes (Ancylocaris) agag Kemp, 1922: 197; Figs 47-50; Plate 7, fig. 9. [Ross Channel, Port Blair, 

Andamans, 4-8 fms] 

Cuapetes akiensis (Kubo, 1936b) 

= Periclimenes (Ancylocaris) akiensis Kubo, 1936b: 47; Plate 14. [Simokamogari-mura, Province 

Aki] 

Cuapetes americanus (Kingsley, 1878a) 

= Anchistia americana Kingsley, 1878a: 96. [Key West, Florida] 

= Periclimenes (Ancylocaris) bermudensis Lebour, 1949a: 1115; Fig. 6. [Mangrove Lake, Bermuda] 

= Periclimenes (Ancylocaris) rhizophorae Lebour, 1949b: 605. [nomen novum for Periclimenes (Ancylocaris) 

bermudensis Lebour, 1949a nec Periclimenes (Periclimenaeus) bermudensis Armstrong, 1940] 

Cuapetes amymone (De Man, 1902) 

= Periclimenes amymone De Man, 1902: 829; Plate 25, Figs 53-53g. [Ternate]


Cuapetes anacanthus (Bruce, 1988b) 

= Periclimenes anacanthus Bruce, 1988b: 105; Figs 1-5. [Polka Point, Dunwich North Stradbroke Island, 

Moreton Bay, Queensland, 27°29'S 153°24'E, 0.5-1.0 m below LWS] 

Cuapetes andamanensis (Kemp, 1922) 

= Periclimenes (Ancylocaris) andamanensis Kemp, 1922: 204; 54-57. [Ross Channel, Port Blair, Andamans, 

4-8 fms] 

Cuapetes calmani (Tattersall, 1921) 

= Periclimenes calmani Tattersall, 1921: 385; Plate 27, fig. 11; Plate 28, figs 14-15. [Sudan coast] 

Cuapetes darwiniensis (Bruce, 1987a) 

= Periclimenes darwiniensis Bruce, 1987a: 29; Figs 1-5. [Weed Reef, Darwin Harbour, 12°31.6'S 

130°47.3'E, LWS] 

Cuapetes demani (Kemp, 1915) 

= Periclimenes demani Kemp, 1915: 279; Fig. 27; Plate 13, fig. 10. [Ennur backwater and Adar river 

(both neighbourhood of Madras); Chilka Lake] 

Cuapetes edwardsii (Paul’son, 1875) 

= Anch.[istia] Edwardsii Paul’son, 1875: 114; Plate 17, fig. 2-2b. [Red Sea] 

Cuapetes elegans (Paul’son, 1875) 

= Anch.[istia] elegans Paul’son, 1875: 113; Plate 17, fig. 1-1h. [Red Sea] 

= Periclimenes (Falciger) dubius Borradaile, 1915b: 211. [Minikoi] 

Cuapetes ensifrons (Dana, 1852a) 

= Anchistia ensifrons Dana, 1852a: 25. [in freto ”Balabac”] 

Cuapetes grandis (Stimpson, 1860a) 

= Anchistia grandis Stimpson, 1860a: 39. [ad insulam ”Ousima”] 

= Periclimenes vitiensis Borradaile, 1898: 383. [Fiji] 

Cuapetes johnsoni (Bruce, 1987b) 

= Periclimenes johnsoni Bruce, 1987b: 115; Figs 1-5. [Pasir Laba, Singapore, 1°21'N 103°38'E] 

Cuapetes kororensis (Bruce, 1977b) 

= Periclimenes kororensis Bruce, 1977b: 33; Figs 1-4. [Koror, Palau Islands] 

Cuapetes lacertae (Bruce, 1992c) 

= Periclimenes lacertae Bruce, 1992c: 46; Figs 1, 2-6. [off Chinaman’s Ridge, Mrs Watson’s Bay, Lizard 

Island, 22-33 m] 

Cuapetes longirostris (Borradaile, 1915b) 

= Palæmonella longirostris Borradaile, 1915b: 210. [Fardiffolu Atoll, Maldive Islands] 

= Periclimenes (Falciger) affinis Borradaile, 1915b: 211. [Salomon Island] 

= Periclimenes (Ancylocaris) proximus Kemp, 1922: 201; Figs 51-53. [Ross Channel, Port Blair, Andamans, 

4-8 fms] 

Cuapetes nilandensis (Borradaile, 1915b) 

= Periclimenes (Falciger) nilandensis Borradaile, 1915b: 211. [S. Nilandu Atoll, Maldive Islands] 

Cuapetes paulsoni (Bruce, 2003a) 

= Periclimenes paulsoni Bruce, 2003a: 118; Fig. 3A-P. [Cape Flattery, inner wharf pile, 7 m] 

Cuapetes platycheles (Holthuis, 1952c) 

= Periclimenes (Harpilius) platycheles Holthuis, 1952c: 85; Fig. 33. [off Fau Island, west coast of Gebe 

Island (31 m); off Atiationin, west coast of New Guinea (to 57 m)] 

Cuapetes rapanui (Fransen, 1987) 

= Periclimenes rapanui Fransen, 1987: 519; Figs 13-15. [Tahai, W coast of Easter Island, Chile, 39 m] 

Cuapetes seychellensis (Borradaile, 1915b) 

= Periclimenes (Falciger) seychellensis Borradaile, 1915b: 212. [Praslin, Seychelles] 

Cuapetes suvadivensis (Borradaile, 1915b) 

= Periclimenes (Falciger) suvadivensis Borradaile, 1915b: 212. [Suvadiva Atoll, Maldive Islands] 

Cuapetes tenuipes (Borradaile, 1898) 

= Periclimenes tenuipes Borradaile, 1898: 384. [New Britain] 

= Periclimenes (Falciger) kolumadulensis Borradaile, 1915b: 213. [Kolumadulu Atoll, Maldive Islands]


= Periclimenes borradailei Rathbun, 1904: 34. [nomen novum for Periclimenes tenuipes Borradaile, 

1898] 

Cuapetes ungujaensis (Bruce, 1969a) 

= Periclimenes ungujaensis Bruce, 1969a: 275. [Unguja Ukuu, Zanzibar Island, 6°18.8'S 39°21.1'E, 

0.5 ft] 

Dactylonia Fransen, 2002 

= Dactylonia Fransen, 2002 (type species Pontonia ascidicola Borradaile, 1898, by original designation, 


Dactylonia anachoreta (Kemp, 1922) 

= Pontonia anachoreta Kemp, 1922: 264; Figs 93-95. [Off Madras coast, 20 fms] 

Dactylonia ascidicola (Borradaile, 1898) 

= Pontonia ascidicola Borradaile, 1898: 389. [New Britain] 

Dactylonia borradalei Bruce, 2005b: 375. [Neokumbi Reef, New Caledonia, 20-40 m] 

Dactylonia carinicula Bruce, 2006a: 4; Figs 2-5. [Rhyi di Hamri, E of Hawlaf, N coast, 12°40.429'N 

54°11.731'E, 7-9 m] 

Dactylonia franseni Bruce, 2003b: 299; Figs 1-4. [Nyali, Old Port, Mombasa, Kenya, 2 m, in Ascidia sp.] 

Dactylonia holthuisi Fransen, 2002: 295; Figs 194-201; Plate 13A. [Ambon, S coast, Seri Bay, 03°45'S 

128°09'E, 25 m, in colony of Plurella spec.] 

Dactylonia monnioti (Bruce, 1990c) 

= Pontonia monnioti Bruce, 1990c: 183; Figs 21-24, 38e-h, 39i-j. [Chesterfield Islands, 24°46.6'S 

159°40.3'E, 285 m] 

Dactylonia okai (Kemp, 1922) 

= Pontonia okai Kemp, 1922: 261; Figs 89-92. [off Cape Negrais, Burma, 15°25'N 93°45'E, 40-49 fms] 

Dasella lebour, 1945 

= Dasia Lebour, 1938 (type species Dasia herdmaniæ Lebour, 1938, by monotypy, gender feminine; 

invalid junior homonym of Dasia Gray, 1839 (Reptilia)) 

= Dasella Lebour, 1945 (nomen novum for Dasia Lebour, 1938, gender feminine) 

Dasella ansoni Bruce, 1983a: 22; Figs 1-5. [Arafura Sea, 10°58'S 132°10'E, 27 m; see Bruce & Coombes, 

1995] 

Dasella brucei Berggren, 1990: 558. [Heron Island, Queensland, Australia, 15 m] 

Dasella herdmaniae (Lebour, 1938) 

= Dasia herdmaniæ Lebour, 1938: 650; Plate 1. [Tuticorin, Gulf of Manaar, Madras] 

Dasycaris Kemp, 1922 

= Dasycaris Kemp, 1922 (type species Dasycaris symbiotes Kemp, 1922, by monotypy, gender feminine; 


Dasycaris ceratops Holthuis, 1952c: 176; Figs 87-88. [Borneo Bank, 2°25'S 117°43'E, 34-0 m] 

Dasycaris doederleini (Balss, 1924) 

= Dasygius doederleini Balss, 1924: 49; Fig. 2. [Dzushi, Sagamibai, 130 m] 

Dasycaris symbiotes Kemp, 1922: 240; Figs 76-77; Plate 9. [2.5 miles ESE of Santapilli Lt., near Vizagapatam, 

Madras Coast, 15-17 fms and 3 miles ESE of Kabusa Island, Mergui, 12°44'30''N 97°55'30''E, 35 

fms] 

Dasycaris zanzibarica Bruce, 1973b: 247; Figs 1-5. [Changu Island reefs, off west coast of Zanzibar, 6°06.2'S 

39°08.9'E, 8 m] 

Diapontonia Bruce, 1986c 

= Diapontonia Bruce, 1986c (type species Diapontonia maranulus Bruce, 1986c, by original designation 


Diapontonia maranulus Bruce, 1986c: 126; Figs 1-5. [off Wood Cay, West End, Grand Bahama Island, 

26°42.55'N 79°01.72'W, 244-309 m]


Epipontonia Bruce, 1977c 

= Epipontonia Bruce, 1977c (type species Epipontonia spongicola Bruce, 1977c, by original designation 


Epipontonia anceps Bruce, 1983b: 19; Figs 1-10. [Heron Island, Queensland, 23°28.0'S 151°59.2'E, 18 m] 

Epipontonia hainanensis Li, 1999: 357; Figs 1-3. [Yezhu Island, Yalong Bay, Hainan Island, 6-9 m, from 

dead corals] 

Epipontonia spongicola Bruce, 1977c: 308; Figs 1-5. [Wasini Channel, Kenya, 4°39.4'S 39°22.2'E, 6 fms] 

Epipontonia tahitiensis Bruce, 2004c: 280; Figs 1-4. [French Polynesia, Tahiti] 

Eupontonia Bruce, 1971b 

= Eupontonia Bruce, 1971b (type species Eupontonia noctalbata Bruce, 1971b, by original designation 


Eupontonia noctalbata Bruce, 1971b: 227; Figs 1-5. [Anse Etoile, Mahé, Seychelle Islands, 04°35'12"S 

55°27'48"E, coral reef flats, 0.3 m] 

Eupontonia oahu Bruce, 2010d: 406; Figs 1-5. [Kahe Point, Oahu, Hawai’ian Islands] 

Exoclimenella Bruce, 1995 

= Exoclimenella Bruce, 1995 (type species Periclimenes Petitthouarsi var. denticulata Nobili, 1906b, by 


= Exoclimenella Ďuriš & Bruce, 1995 (type species Periclimenes Petitthouarsi var. denticulata Nobili, 

1906b, by original designation, gender feminine) 

Exoclimenella denticulata (Nobili, 1906b) 

= Periclimenes Petitthouarsi var. denticulata Nobili, 1906b: 257. [Gatavake] 

Exoclimenella maldivensis Ďuriš & Bruce, 1995: 622; Figs 1-5. [Genego Islet, North Nilandu Atoll, Maldive 

Islands, outer reef slope, 20 m] 

Exoclimenella sibogae (Holthuis, 1952c) 

= Periclimenes (Harpilius) sibogae Holthuis, 1952c: 73; Figs 28-29. [Banda anchorage, 9-36 m] 

Exoclimenella sudanensis Ďuriš & Bruce, 1995: 631; Figs 6-8. [coral reef near Port Sudan, Sudan, Red Sea, 

in coral head of Stylophora pistillata] 

Exopontonia Bruce, 1988c 

= Exopontonia Bruce, 1988c (type species Exopontonia malleatrix Bruce, 1988c, by original designation 


Exopontonia malleatrix Bruce, 1988c: 123; Figs 1-5. [Ashmore Reef, Timor Sea, 12°16'S 123°02'E, intertidal] 

Fennera Holthuis, 1951b 

= Fennera Holthuis, 1951b (type species Fennera chacei Holthuis, 1951b, by original designation and 


Fennera chacei Holthuis, 1951b: 171; Plate 54. [Secas Islands, Bay of South Island, shallow water] 

Fennera holthuisi Marin, 2011: 34; Figs 5-7, 8d-f. [French Polynesia, Moorea, outer reef slope, 17°29’02”S 

149°52’09’W, 10 m, in Pocillopora] 

Hamiger Borradaile, 1916 

= Hamiger Borradaile, 1916 (type species Periclimenes (Hamiger) novae-zealandiae Borradaile, 1916, by 


Hamiger novaezealandiae (Borradaile, 1916) 

= Periclimenes (Hamiger) novae-zealandiae Borradaile, 1916: 87; Fig. 4. [seven miles E. of North Cape, 

New Zealand, 128 m] 

Hamodactyloides Fujino, 1973b 

= Hamodactyloides Fujino, 1973b (type species Hamodactylus incompletus Holthuis, 1958, by original 

designation, gender masculine)


Hamodactyloides incompletus (Holthuis, 1958) 

= Hamodactylus incompletus Holthuis, 1958: 11; Fig. 4. [Sharam a Sheikh, Sinai Peninsula] 

= Hamodactyloides ishigakiensis Fujino, 1973b: 174; Figs 1-3. [Kabira Bay, Ishigaki-jima Island, Ryukyu 

Islands, 1 m] 

Hamodactylus Holthuis, 1952c 

= Hamodactylus Holthuis, 1952c (type species Hamodactylus boschmai Holthuis, 1952c, by original 


Hamodactylus aqabai Bruce & Svoboda, 1983: 26; Figs 10-14. [Aqaba, Jordan] 

Hamodactylus boschmai Holthuis, 1952c: 209; Figs 102-104. [Ternate, off Halmahera, and Djedan, Kepulauan 

Aru, Indonesia, 2-13 m] 

Hamodactylus noumeae Bruce, 1970a: 539; Fig. 2. [between Isle aux Canards and Islôt Maître, near 

Noumea, New Caledonia, 25 m] 

Hamopontonia Bruce, 1970b 

= Hamopontonia Bruce, 1970b (type species Hamopontonia corallicola Bruce, 1970b, by original designation 


Hamopontonia corallicola Bruce, 1970b: 41; Figs 1-4. [Kat O Chau, Mirs Bay, New Territories, Hong Kong, 

22°32.1'N 114°17.95'E] 

Hamopontonia essingtoni Bruce, 1986d: 158; Figs 1C, 11-14, 15D-G. [Coral Bay, Port Essington, 11°11.05'S 

132°03.4'E, 6 m] 

Harpiliopsis Borradaile, 1917 

= Harpiliopsis Borradaile, 1917 (type species Palæmon Beaupressi Audouin, 1826, by original designation, 


Harpiliopsis beaupresii (Audouin, 1826) 

= Palæmon beaupresii Audouin, 1826: 91 (accompanying figures in Savigny, 1826). [Egypt] 

= Pontonia (Harpilius) dentata Richters, 1880: 165; Plate 17, Figs 36-38. [Fouquets] 

Harpiliopsis depressa (Stimpson, 1860a) 

? = Anchistia gracilis Dana, 1852a: 25. [in mari Suluensi] 

= Harpilius depressus Stimpson, 1860a: 38. [ad insulam ”Hawaii”, inter madreporas] 

= Pelias notatus Heller, 1862a: 526. [Nicobaren] 

= Anchistia notata Heller, 1865: 109; Plate 10, fig. 3. [Nicobaren] 

= Periclimenes pusillus Rathbun, 1906: 921; Fig. 71; Plate 24, fig. 7. [south coast of Oahu, surface] 

Harpiliopsis spinigera (Ortmann, 1890) 

= Anchistia spinigera Ortmann, 1890: 511; Plate 36, fig. 23-23a. [Samoa] 

= Harpilius depressus var. gracilis Kemp, 1922: 234; Fig. 71. [Andamans] 

Harpilius Dana, 1852a 

= Harpilius Dana, 1852a (type species Harpilius lutescens Dana, 1852a, by monotypy, gender masculine; 


Harpilius bayeri (Holthuis, 1981) 

= Periclimenes bayeri Holthuis, 1981: 792; Fig. 3. [Ine village, Arno Atoll, Marshall Islands, outer edge 

of sea reef, on coral, from head of Pocillopora spec. according to K. Reed (pers. comm.)] 

= Periclimenes (Harpilius) bayeri Holthuis, 1953b: 56. [nomen nudum] 

Harpilius consobrinus De Man, 1902: 836; Plate 26, fig. 54-54c. [Ternate] 

Harpilius lutescens Dana, 1852a 

= Harpilius lutescens Dana, 1852a: 25. [insula Tongatabu] 

= Periclimenes (Ancylocaris) amamiensis Kubo, 1940b: 44; Figs 11, 12. [Ôsima, Riu-Kiu Islands] 

Harpilius spinifer Bruce, 2006b: 272; Figs 1-5. [Ilôt Maitre, New Caledonia, 4 m]


Holthuisaeus Anker & De Grave, 2010 

= Holthuisaeus Anker & De Grave, 2010 (type species Periclimenes (Periclimenaeus) bermudensis Armstrong, 

1940, by original designation and monotypy, gender masculine) 

Holthuisaeus bermudensis (Armstrong, 1940) 

= Periclimenes (Periclimenaeus) bermudensis Armstrong, 1940: 4; Figs 2, 3A-F. [The Reach, St. George 

Island, Bermuda] 

Ischnopontonia Bruce, 1966c 

= Ischnopontonia Bruce, 1966c (type species Philarius lophos Barnard, 1962, by original designation 


Ischnopontonia lophos (Barnard, 1962) 

= Philarius lophos Barnard, 1962: 242; Fig. 2. [Inhaca Island, Delagoa Bay, U.W.] 

Isopontonia Bruce, 1982a 

= Isopontonia Bruce, 1982a (type species Isopontonia platycheles Bruce, 1982a, by original designation 


Isopontonia platycheles Bruce, 1982a: 55; Figs 1-4. [North Cay, Bennet Island, Chesterfield Islands, 19°48.0'S 

158°17.0'E, 15 m] 

Izucaris okuno, 1999a 

= Izucaris Okuno, 1999a (type species Izucaris masudai Okuno, 1999a, by original designation and 


Izucaris crosnieri Li, 2008: 207; Figs 1-4, 5A. [Marquesas Islands, Eiao, 7°59.2'S 140°44.2'W, 100 m] 

Izucaris masudai Okuno, 1999a: 399; Figs 1-6. [34°52.7'N 39°08.2'E, Izu Oceanic Park, Izu Peninsula, Honshu, 

Japan, 27 m] 

Jocaste Holthuis, 1952c 

= Jocaste Holthuis, 1952c (type species Coralliocaris lucina Nobili, 1901c, by monotypy, gender feminine) 

Jocaste japonica (Ortmann, 1890) 

= Coralliocaris superba var. japonica Ortmann, 1890: 50; Plate 36, fig. 22. [Japan, Kagoshima] 

? = Cavicheles kempi Holthuis, 1952c: 205; Figs 99-101. [Ternate, 4 m] 

Jocaste lucina (Nobili, 1901c) 

= C.[oralliocaris] lucina Nobili, 1901c: 5. [Eritrea] 

? = Coralliocaris lamellirostris Stimpson, 1860a: 38. [ad insulam ”Loo Choo”, inter corallia ad prof 2 org] 

Jocaste platysoma Fransen, 1994: 111; Figs 65-84; Plate 2C. [St. Joseph Atoll, S rim, 5°27'S 53°21'E, 10 m, on 

tabular Acropora] 

Laomenes Clark, 1919 

= Corniger Borradaile, 1915b (type species Periclimenes (Corniger) ceratophthalmus Borradaile, 1915b, 

designated by Borradaile, 1917, gender masculine; invalid junior homonym of Corniger Agassiz, 

1831 (Pisces) and Corniger Boehm, 1879 (Pycnogonida)) 

= Laomenes Clark, 1919 (nomen novum for Corniger Borradaile, 1915b) 

= Parapontonia Bruce, 1968a (type species Parapontonia nudirostris Bruce, 1968a, by original designation 


Laomenes amboinensis (De Man, 1888b) 

= Anchistia amboinensis De Man, 1888b: 546; Plate 22a, figs 2-2b. [Amboina] 

Laomenes ceratophthalmus (Borradaile, 1915b) 

= Periclimenes (Corniger) ceratophthalmus Borradaile, 1915b: 211. [Malé Atoll, Maldive Islands, on 

crinoid] 

Laomenes clarki Marin, 2009a: 13; Figs 13-19. [Vietnam, Nhatrang Bay, Nok Island, 15 m, on crinoid Zygometra 

cf. punctata]


Laomenes cornutus (Borradaile, 1915b) 

= Periclimenes (Corniger) cornutus Borradaile, 1915b: 211. [Malé Atoll, Maldive Islands, on crinoid] 

Laomenes holthuisi Marin & Okuno, 2010: 462; Figs 1-6. [Pacific Ocean, Japan, Izu Islands, Hachijo Island, 

Aozumi, 15 m] 

Laomenes jackhintoni (Bruce, 2006c) 

= Periclimenes jackhintoni Bruce, 2006c: 23; Figs 1-5. [Nukualofa, Tongatapu, Tonga, 1.5 m] 

Laomenes nudirostris (Bruce, 1968a) 

= Parapontonia nudirostris Bruce, 1968a: 1149; Figs 1-5. [Tiaré Bay, Noumea, New Caledonia, 22°10'S 

166°15'E] 

Laomenes pardus Marin, 2009a: 19; Figs 15-18. [Vietnam, Nhatrang Bay, Mun Island, 5 m, on crinoid 

Comantheria briareus] 

Laomenes tigris Marin, 2009a: 25; Figs 20-24. [Mun Island, Nhatrang Bay, Vietnam, 15-20 m, on crinoid 

Stephanometra sp.] 

Leptomenaeus Bruce, 2007b 

= Leptomenaeus Bruce, 2007b (nomen novum for Leptomenes Bruce, 2006d, gender masculine) 

= Leptomenes Bruce, 2006d (type species Periclimenes dolichosternum Okuno & Mitsuhashi, 2003, by 

original designation and monotypy, gender masculine; invalid junior homonym of Leptomenes Soika, 

1939 (Insecta)) 

Leptomenaeus dolichosternum (Okuno & Mitsuhashi, 2003) 

= Periclimenes dolichosternum Okuno & Mitsuhashi, 2003: 487-496; Figs 1-5. [Ou-shima Harbor, 

Kume Island, Ryukyu Islands, 26°20.1'N 126°49.2'E, 1.5 m] 

Leptomenaeus nhatrangensis Ďuriš & Horká, 2008: 46; Figs 1-5. [Hon Chong Beach, N of Nhatrang Bay 

(South China Sea), Vietnam, 12°17'45.8''N 109°14'05.7''E, 1 m] 

Lipkebe Chace, 1969 

= Lipkebe Chace, 1969 (type species Lipkebe holthuisi Chace, 1969, by original designation and monotypy, 


Lipkebe holthuisi Chace, 1969: 263; Figs 8-9. [Gulf of Mexico, west-northwest of Dry Tortugas, Florida, 

25°13'N 83°55'W, 119 m] 

Lipkemenes Bruce & okuno, 2010 

= Lipkemenes Bruce & Okuno, 2010 (type species Periclimenes (Periclimenes) lanipes Kemp, 1922, by 


Periclimenes lanipes Kemp, 1922 

= Periclimenes (Periclimenes) lanipes Kemp, 1922: 156; Plate 4, fig. 4. [Mergui Archipelago, 12°48'N 

98°16'10"E, 24 fms] 

Manipontonia Bruce, okuno & li, 2005 

= Manipontonia Bruce, Okuno & Li, 2005 (type species Urocaris psamathe De Man, 1902, by original 


Manipontonia persiana Marin, 2010: 64; Figs 3-4. [Indian Ocean, Persian Gulf, 28°49.661'N 49°48.506'E, 51 

m depth] 

Manipontonia psamathe (De Man, 1902) 

= Urocaris psamathe De Man, 1902: 816; Plate 25, fig. 51. [Ternate] 

Margitonia Bruce, 2007c 

= Margitonia Bruce, 2007c (type species Periclimenes insolitus Bruce, 1974b, by original designation 


Margitonia insolitus (Bruce, 1974b) 

= Periclimenes insolitus Bruce, 1974b: 293; Figs 1-8. [Waikiki Beach, Oahu, Hawaii, 21°15.9'N 

157°50.5'W]


Mesopontonia Bruce, 1967b 

= Mesopontonia Bruce, 1967b (type species Mesopontonia gorgoniophila Bruce, 1967b, by original designation 


Mesopontonia brevicarpus Li & Bruce, 2006: 652; Figs 7-8. [La Réunion, 20°51.5'S 55°36.87'E, 270 m] 

Mesopontonia brucei Burukovsky, 1991b: 38; Figs 1.8-1.18. [33°16'S 43°53'E, 415-460 m] 

Mesopontonia gorgoniophila Bruce, 1967b: 13; Figs 5-9. [21°47.7'N 116°28.5'E to 21°43.3'N 116°28.0'E, 64-72 fms] 

Mesopontonia gracilicarpus Bruce, 1990c: 202; Figs 34-37, 39l-m. [New Caledonia, 22°56'N 167°14'E, 398- 

410 m] 

Mesopontonia monodactylus Bruce, 1991a: 392; Figs 65-69. [Loyalty Islands, 20°35.0'S 166°54.0'E, Uvéa, 460 

m] 

Mesopontonia verrucimanus Bruce, 1996: 216; Figs 8a-n, 29c. [Indonesia, Tanimbar Islands, 7°59'S 133°02'E, 

184-186 m] 

Metapontonia Bruce, 1967b 

= Metapontonia Bruce, 1967b (type species Metapontonia fungiacola Bruce, 1967b, by original designation 


Metapontonia fungiacola Bruce, 1967b: 24: Figs 10-12. [Pamanzi Reef, Mayotte Island, Comore Archipelago] 

Miopontonia Bruce, 1985a 

= Miopontonia Bruce, 1985a (type species Miopontonia yongei Bruce, 1985a by original designation 


Miopontonia yongei Bruce, 1985a: 168; Figs 1-5. [19°04.3'S 118°15.5'E, 80 m] 

Neoanchistus Bruce, 1975a 

= Neoanchistus Bruce, 1975a (type species Neoanchistus cardiodytes Bruce, 1975a, by original designation 


Neoanchistus cardiodytes Bruce, 1975a: 151; Figs 1-6. [Nosy Bé, Madagascar] 

Neoanchistus nasalis Holthuis, 1986c: 264; Figs 1-2. [artificial reef near Raysut Harbour, Dhofar, southern 

Oman, 16°57'45''N 54°00'E, in Chlamys townsendi] 

Neoclimenes Mitsuhashi, li & Chan, 2010 

= Neoclimenes Mitsuhashi, Li & Chan, 2010 (type species Neoclimenes holthuisi Mitsuhashi, Li & 

Chan, 2010, by original designation, gender masculine) 

Neoclimenes holthuisi Mitsuhashi, Li & Chan, 2010: 518; Figs 1-6. [21°40.2'N 117°43.4'E, 513-509 m] 

Neopericlimenes Heard, Spotte & Bubucis, 1993 

= Neopericlimenes Heard, Spotte & Bubucis, 1993 (type species Neopericlimenes thornei Heard, Spotte 

& Bubucis, 1993, by original designation and monotypy, gender masculine) 

Neopericlimenes thornei Heard, Spotte & Bubucis, 1993: 795; Figs 1-4. [fringing reef off north shore of Pine 

Cay, Turks and Caicos Islands, British West Indies, 40 m, approx. 21°53'N 72°06'W] 

Neopontonides Holthuis, 1951b 

= Neopontonides Holthuis, 1951b (type species Periclimenes beaufortensis Borradaile, 1920, by original 


Neopontonides beaufortensis (Borradaile, 1920) 

= Periclimenes beaufortensis Borradaile, 1920: 132. [Beaufort, North Carolina] 

Neopontonides brucei Fransen & de Almeida, 2009: 838; Figs 1-4. [Boquirão beach, Ubatuba, NE São Paulo, 

Brazil, 32 m, mud] 

Neopontonides chacei Heard, 1986: 472; Figs 1A, 2-3, 4B-D. [reef just south of Marigot Bay, St. Lucia Island, 

West Indies, 4-6 m] 

Neopontonides dentiger Holthuis, 1951b: 193; Plate 61. [Ecuador, off Cape San Francisco, 2 fms] 

Neopontonides henryvonprahli Ramos, 1995: 231; Figs 1-3. [Punta Alta, Bahía Málaga, simbionte de Muricea 

robusta, 5 m]


Nippontonia Bruce & Bauer, 1997 

= Nippontonia Bruce & Bauer, 1997 (type species Nippontonia minirostris Bruce & Bauer, 1997, by 


Nippontonia minirostris Bruce & Bauer, 1997: 1711; Figs 1-8. [Nishihama, Aka-jima, Kerama group, Ryukyu 

Islands, Japan, 15 m] 

Notopontonia Bruce, 1991c 

= Notopontonia Bruce, 1991c (type species Notopontonia platycheles Bruce, 1991c, by original designation 


Notopontonia platycheles Bruce, 1991c: 608; Figs 1-6, 14E. [northwest of Robe, South Australia, 36°53'S 

139°53'E, 64 m] 

Odontonia Fransen, 2002 

= Odontonia Fransen, 2002 (type species Pontonia katoi Kubo, 1940b, by original designation, gender 

feminine) 

Odontonia compacta (Bruce, 1996) 

= Pontonia compacta Bruce, 1996: 242; Figs 15-18. [New Caledonia, 22°36.3'S 167°14.2'E, 10-60 m] 

Odontonia katoi (Kubo, 1940b) 

= Pontonia katoi Kubo, 1940b: 55; Figs 21-23. [Off Simoda, Siduoka Prefecture] 

Odontonia maldivensis Fransen, 2006a: 718; Figs 2-5. [Maldives, S Malé Atoll, ocean side North reef, 

04°07.54'N 73°30.55'E, 10 m, in Polycarpa cryptocarpa] 

Odontonia rufopunctata Fransen, 2002: 352; Figs 229-234; Plate 17. [SW Sulawesi, Spermonde Archipelago, 

Kudingareng Keke, 20 m, in ascidian] 

Odontonia seychellensis Fransen, 2002: 361; Figs 235-240; Plate 18. [Seychelles, Bird Island, off N coast, 

3°42'S 55°12'E, 20 m, in Ascidiidae] 

Odontonia sibogae (Bruce, 1972d) 

= Pontonia sibogae Bruce, 1972d: 182; Fig. 1. [Curtis Channel, Port Curtis, Queensland, Australia, 23 

fms] 

Odontonia simplicipes (Bruce, 1996) 

= Pontonia simplicipes Bruce, 1996: 247; Figs 19-20, 29i. [Chesterfield Islands, 19°15.15'S 158°47.73'E, 71 m] 

Onycocaridella Bruce, 1981b 

= Onycocaridella Bruce, 1981b (type species Onycocaridella prima Bruce, 1981b, by original designation, 


Onycocaridella monodoa (Fujino & Miyake, 1969b) 

= Onycocaridella monodoa Fujino & Miyake, 1969b: 405; Figs 1-5. [Kasari-zaki, Amami-oshima Island, 

Ryukyu Islands, Japan, 1 m] 

Onycocaridella prima Bruce, 1981b: 243; Figs 1-6. [Wistari Reef, Heron Island, Capricorn Islands, Queensland, 

23°25.7'S 151°55.0'E, 12 m] 

Onycocaridella stenolepis (Holthuis, 1952c) 

= Onycocaris stenolepis Holthuis, 1952c: 148; Figs 66-68. [southeast side of Pearl Bank, Sulu Archipelago, 

15 m] 

Onycocaridites Bruce, 1987c 

= Onycocaridites Bruce, 1987c (type species Onycocaridites anomodactylus Bruce, 1987c, by original 


Onycocaridites anomodactylus Bruce, 1987c: 772; Figs 1-4. [Arafura Sea, 10°40'S 133°50'E, 60 m] 

Onycocaris Nobili, 1904 

= Onycocaris Nobili, 1904 (type species Coralliocaris (Onycocaris) aualitica Nobili, 1904, designated by 

Holthuis, 1952c, gender feminine) 

Onycocaris amakusensis Fujino & Miyake, 1969b: 413; Figs 6, 8a-c, 9a-c. [Tsuji-shima Islet, Amakusa Island, 

in sponge collected at low tide level]


Onycocaris aualitica (Nobili, 1904) 

= Coralliocaris (Onycocaris) aualitica Nobili, 1904: 233. [Djibouti] 

Onycocaris balssi Bruce, 2011b: 478; Figs 1-5. [Table Head, Port Essington, Northern Territory, Australia, 

11°13.5'S 132°10.5'E, 3 m, in unidentified sponge] 

Onycocaris bocki Bruce, 1992e: 335; Figs 4-6. [Tuvalu, Niue, leeward reef] 

Onycocaris callyspongiae Fujino & Miyake, 1969b: 422; Figs 10-12. [Tomioka, Amakusa Island, in sponges] 

Onycocaris fujinoi Bruce, 2011b: 487. [Terasaki, Yoron-jima, Ryukyu Islands, Japan] 

Onycocaris furculata Bruce, 1979c: 324; Figs 1-4. [La Saline, La Réunion, approximately 21°20'S 55°00'E, 20 m] 

Onycocaris longirostris Bruce, 1980a: 15; Figs 6-10. [Ilot Maître, Nouméa, New Caledonia, 20 m] 

Onycocaris nieli Bruce, 2011a: 320; Figs 1-5. [Heron Island, Capricorn Islands, Queensland, reef flat] 

Onycocaris oligodentata Fujino & Miyake, 1969b: 415; Figs 7, 8d-f, 9d-f. [Tomioka, Amakusa Island, in 

sponge fished up entangling with gill net, 35 m] 

Onycocaris profunda Bruce, 1985b: 241; Figs 8-11. [13°33.0'N 122°10.1'E, 81-84 m] 

Onycocaris quadratophthalma (Balss, 1921a) 

= Pontonia quadratophthalma Balss, 1921a: 15; Fig. 7. [Cape Jaubert] 

Onycocaris seychellensis Bruce, 1971c: 208; Figs 1-6. [Anse Etoile, Mahé, Seychelle Islands] 

Onycocaris spinosa Fujino & Miyake, 1969b: 429; Figs 13-15. [Terasaki, Yoron-jima Island, Ryukyu Islands, 

in sponge, 1 m] 

Onycocaris stradbrokei Bruce, 1998c: 390; Figs 2-3. [Myora Reef, North Stradbroke Island, Queensland, 

27°29'S 153°25'E, 9 m] 

Onycocaris temiri Marin, 2005: 113; Figs 1-8. [Mung Island, 16 m, inside boring sponge Kallipilidion] 

Onycocaris trullata Bruce, 1978a: 269; Figs 36-41. [Tany-Kely, Madagascar, 13°28'S 48°12'E, 28 m] 

Onycocaris zanzibarica Bruce, 1971d: 293; Figs 1-2. [channel between Chumbe Island and mainland of 

Zanzibar Island, 6°16.0'S 39°12.6'E, 10 fms] 

Onycomenes Bruce, 2009a 

= Onycomenes Bruce, 2009a (type species Onycocaridella antokha Marin, 2007a, by original designation 


Onycomenes antokha (Marin, 2007a) 

= Onycocaridella antokha Marin, 2007a: 221; Figs 5-9. [South China Sea, Vietnam, Nhatrang Bay, Nok 

Island, 10 m] 

Orthopontonia Bruce, 1982b 

= Orthopontonia Bruce, 1982b (type species Periclimenaeus ornatus Bruce, 1970c, by original designation 


Orthopontonia ornata (Bruce, 1970c) 

= Periclimenaeus ornatus Bruce, 1970c: 313. [Heron Island, Capricorn Group, Great Barrier Reef, Australia, 

littoral] 

Palaemonella Dana, 1852a 

= Palaemonella Dana, 1852a (type species Palaemonella tenuipes Dana, 1852a, designated by Kingsley, 

1880, gender feminine; name placed on the Official List of Generic Names in Zoology in Opinion 

470 in 1957) 

Palaemonella aliska Marin, 2008a: 376; Figs 1-5, 9a. [South China Sea, Vietnam, Nhatrang Bay, Tre Island, 

Tre Bay, mangrove littoral] 

Palaemonella asymmetrica Holthuis, 1951b: 19; Plate 5. [Sullivan Bay, James Island] 

Palaemonella atlantica Holthuis, 1951a: 152; Fig. 31. [São Pedro Bay, São Vicente, Cape Verde Islands, 

16°50'N 25°04'W] 

Palaemonella burnsi Holthuis, 1973a: 24; Figs 8-9. [Cape Kinau Peninsula, Maui, Hawaiian Islands; small 

lava pool near coast of Keoneoio (= La Perouse) Bay, at the extreme east end of the Cape] 

Palaemonella crosnieri Bruce, 1978a: 210; Figs 2-4. [Iles Glorieuses, 11°28.1'S 27°[sic]21.1'E, 20 m] 

Palaemonella dijonesae Bruce, 2010e: 151; Fig. 1. [Rocky head, Enderby Island, Dampier Archipelago, 

20°40'14''S 116°27'96''E, 12.5 m]


Palaemonella disalvoi Fransen, 1987: 511; Figs 7-12. [Tahai, west coast of Easter Islands, 35 m] 

Palaemonella dolichodactylus Bruce, 1991b: 232; Figs, 6f-l, 7. [22°14.5'S 167°02.0'E, Lagon Est, 65-70 m] 

Palaemonella foresti Bruce, 2002b: 279; Figs 1-4. [BP Oil Refinery Jetty, Cockburn Sound, Western Australia, 

32°13.9'S 115°45.14'E, 7 m] 

Palaemonella hachijo Okuno, 1999b: 739; Figs 1-3. [33°03.5'N 139°47.9'E, southern Japan, Izu Islands, 

Hachijo-jima Island, Occho-gahama, submarine cave, 20 m] 

Palaemonella holmesi (Nobili, 1907) 

= Periclimenes Holmesi Nobili, 1907: 5. [nomen novum for Anchista tenuipes Holmes, 1900] 

= Anchista tenuipes Holmes, 1900: 216. [Santa Catalina Island] 

Palaemonella komaii Li & Bruce, 2006: 657; Figs 9-11. [Tonga, Vava’u group, 18°37'S 174°03'W, 327-360 

m] 

Palaemonella lata Kemp, 1922: 127; Figs 3-6. [Aberdeen, Port Blair, Andamans, rock pool at low tide] 

Palaemonella longidactylus Hayashi, 2009: 97; Figs 5-7. [Beppu Bay, Ooita Prefecture] 

Palaemonella maziwi Bruce, 2002c: 15; Fig. 1. [Maziwi Island, Pangani, Tanganyika, 30.6'S 39°04.06'E, 2 m] 

Palaemonella meteorae Bruce, 2008b: 64; Figs 1-2. [Red Sea, off Port Sudan, 19°33.3'N 37°16.4'E - 19°33.9'N 

37°16.2'E, 519-544 m] 

Palaemonella pottsi (Borradaile, 1915b) 

= Periclimenes (Falciger) pottsi Borradaile, 1915b: 212. [Torres Straits, on Comanthus] 

Palaemonella pusilla Bruce, 1975b: 169; Figs 1-5. [Kisiti Island, near Wasin, Kenya, 4°43.3'S 39°22.15'E, on 

sheltered coral reef at LWS] 

Palaemonella rotumana (Borradaile, 1898) 

= Periclimenes rotumanus Borradaile, 1898: 383. [Rotuma] 

= Palaemonella vestigialis Kemp, 1922: 123; Figs 1-2; Plate 3, fig. 2. [Port Blair, Andamans] 

Palaemonella spinulata Yokoya, 1936: 135; Fig. 4. [Sagami Bay, off Misaki to Yokosuka, Miura Peninsula, 

35°10.53'N 139°34.56'E to 39°10.53'N 139°34.53'E, 73-62 m; neotype designation by Hayashi, 2009] 

Palaemonella tenuipes Dana, 1852a 

= Palæmonella tenuipes Dana, 1852a: 25. [in mari Suluensi] 

= Palæmonella tridentata Borradaile, 1899: 1007; Plate 64, Figs 8-8c. [Funafuti] 

= Palæmonella elegans Borradaile, 1915b: 210. [Salomon Island] 

Paraclimenaeus Bruce, 1988d 

= Paraclimenaeus Bruce, 1988d (type species Periclimenaeus fimbriatus Borradaile, 1915b, by original 


Paraclimenaeus dentimanus (Mitsuhashi & Chan, 2008) 

= Apopontonia dentimanus Mitsuhashi & Chan, 2008: 32; Figs 3-7. [North of Doljo, Panglao Island, 

9°35.9'N 123°44.5'E, 24 m] 

Paraclimenaeus fimbriatus (Borradaile, 1915b) 

= Periclimenæus fimbriatus Borradaile, 1915b: 213. [Makalu Atoll, Maldive Islands; holotype designation 

by Bruce, 1988d invalid] 

Paraclimenaeus seticauda (Bruce, 2008c) 

= Apopontonia seticauda Bruce, 2008c: 3; Figs 2-5, 14A. [18°27'37''S 120°08'41''E, 80 m] 

Paraclimenaeus spinicauda (Bruce, 1969b) 

= Periclimenaeus spinicauda Bruce, 1969b: 164. [South China Sea, 20°57.5'N 115°55.0'E – 20°57.5'N 

115°58.6'E, 35-36 fms] 

= Apopontonia dubia Bruce, 1981c: 225; Figs 1-3. [Shag Rock, E of North Stradbroke Island, Queensland, 

Australia, 27°25'S 153°32'E, 20 m] 

Paraclimenes Bruce, 1995 

= Paraclimenes Bruce, 1995 (type species Periclimenes franklini Bruce, 1990e, by original designation, 


Paraclimenes franklini (Bruce, 1990e) 

= Periclimenes franklini Bruce, 1990e: 55; Figs 1-5. [17°21.7'S 146°48.52'E, 296-302 m]


Paraclimenes gorgonicola (Bruce, 1969a) 

= Periclimenes gorgonicola Bruce, 1969a: 257. [South China Sea, 21°47.7'N 116°28.5'E to 21°43.3'N 

116°28.0'E, 60-70 fms] 

Paraclimenes setirostris (Bruce, 1991a) 

= Periclimenes setirostris Bruce, 1991a: 364; Figs 45-49. [Chesterfield Islands, 25°32.8'S 159°46.1'E, 300 m] 

Paranchistus Holthuis, 1952c 

= Paranchistus Holthuis, 1952c (type species Anchistus biunguiculatus Borradaile, 1898 (junior subjective 

synonym of Pontonia armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840], by 


Paranchistus armatus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Pontonia armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 359. [près de côtes de 

la Nouvelle-Irlande] 

= Anchistus biunguiculatus Borradaile, 1898: 387. [British New Guinea] 

= Anchistus oshimai Kubo, 1949b: 26; Figs 1-2. [Helen Atoll] 

Paranchistus liui Li, Bruce & Manning, 2004: 535; Figs 21-23. [21°N 108°E, 30 m] 

Paranchistus nobilii Holthuis, 1952c: 100; Figs 41-42. [Arzana Island, Rakas Zakoum Bank, Persian Gulf, 

out of Spondylus gaederopus] 

Paranchistus ornatus Holthuis, 1952c: 97; Figs 39-40. [Mozambique] 

Paranchistus pycnodontae Bruce, 1978b: 233; Figs 1-5; Plate 39. [Heron Island, Capricorn Group, Queensland, 

3 m] 

= Paranchistus serenei Bruce, 1983c: 890; Figs 7H-I, 9. [Seleman Bay, Seram Island] 

Paranchistus spondylis Suzuki, 1971: 15; Figs 8-9. [Shiraiso, near the Manazuru M.B.L.] 

Paratypton Balss, 1914d 

= Paratypton Balss, 1914d (type species Paratypton siebenrocki Balss, 1914d, by monotypy, gender 

masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Paratypton siebenrocki Balss, 1914d: 84; Fig. 1. [Senafir, Koseir, Mersa Scheikh; Jaluit, Samoa] 

Patonia Mitsuhashi & Chan, 2006 

= Patonia Mitsuhashi & Chan, 2006 (type species Patonia mclaughlinae Mitsuhashi & Chan, 2006, by 


Patonia mclaughlinae Mitsuhashi & Chan, 2006: 390; Figs 1-6. [Taiwan northeastern coast, 24°34.71'N 

122°4.02'E, 209-280 m] 

Periclimenaeus Borradaile, 1915b 

= Periclimenæus Borradaile, 1915b (type species Periclimenæus robustus Borradaile, 1915b, designated 

by Borradaile, 1917, gender masculine; name placed on the Official List of Generic Names in Zoology 


Periclimenaeus arabicus (Calman, 1939) 

= Periclimenes (Periclimenæus) arabicus Calman, 1939: 210; Fig. 4. [Khalij al Masirah, eastern Oman, 

19°22.6'N 57°53.0'E, 13.5 m, from surface of sponge] 

= Periclimenaeus ohshimai Miyake & Fujino, 1967: 275; Fig. 1. [Takamatsu, Amakusa Islands, Kyushu] 

Periclimenaeus ardeae Bruce, 1970c: 310. [Heron Island, Capricorn Group, Great Barrier Reef, Queensland, 

Australia, littoral] 

Periclimenaeus arthrodactylus Holthuis, 1952c: 122; Figs 51-53. [Sailus-ketjil, Paternoster Islands, up to 18 

m] 

Periclimenaeus ascidiarum Holthuis, 1951b: 80; Plate 22, figs g-l; Plate 23. [Bird Key Reef, Tortugas] 

Periclimenaeus atlanticus (Rathbun, 1901) 

= Coralliocaris atlantica Rathbun, 1901: 122; Fig. 26. [Off St. Thomas 20-23 fms] 

Periclimenaeus aurae dos Santos, Calado & Araújo, 2008: 156; Figs 1-4, 5C, K. [Garajau, south of Madeira 

Island, eastern Atlantic, 32.36°N 16.85°W, 6-7 m]


Periclimenaeus bidentatus Bruce, 1970c: 305. [Heron Island, Capricorn Group, Great Barrier Reef, Queensland, 

Australia, littoral] 

Periclimenaeus bouvieri (Nobili, 1904) 

= Typton Bouvieri Nobili, 1904: 233. [Djibouti] 

Periclimenaeus bredini Chace, 1972: 26; Fig. 5. [Isla Mujeres] 

Periclimenaeus brucei Cardoso & Young, 2007: 302; Figs 20-25. [Rocas Atoll, outer reef, 20 m, in Ircinia sp.] 

Periclimenaeus caraibicus Holthuis, 1951b: 110; Plate 32, figs h-j; Plate 34. [British West Indies, Tobago, 

Buccoo Reef] 

Periclimenaeus chacei Abele, 1971: 38; Figs 1-2. [Gulf of Mexico, off the West coast of Florida, 28°31'N 

84°16'W, 26 m] 

Periclimenaeus colodactylus Bruce, 1996: 222; Figs 9-10. [New Caledonia, Lagoon, Uatio Islet, 20-25 m] 

Periclimenaeus crassipes (Calman, 1939) 

= Periclimenes (Ancylocaris) crassipes Calman, 1939: 211; Fig. 5. [Ghubbat Sawquirah, southeastern 

Oman, 18°025.5'N 57°025'E, 38 m; animals pulled off calcareous sponges or removed from the debris 

in the jar containing them] 

Periclimenaeus creefi Bruce, 2010f: 29; Figs 7-11. [Heron island reef, north-eastern side, 23°25.990'S 

151°55.601'E, 10 m, dead coral heads] 

Periclimenaeus crosnieri Cardoso & Young, 2007: 309; Figs 26-31. [Rocas Atoll, outer reef, 15-20 m, in 

sponge] 

Periclimenaeus devaneyi Bruce, 2010g: 380; Figs 1-5. [Kahe Point, Oahu, Hawai’ian Islands] 

Periclimenaeus diplosomatis Bruce, 1980b: 39; Figs 1-6. [Heron Island, Capricorn Islands, Queensland, 

Australia, 23°26.9'S 151°55'E, low water] 

Periclimenaeus djiboutensis Bruce, 1970c: 307. [Jibouti, French Somaliland] 

Periclimenaeus fawatu Bruce, 2006e: 33; Figs 1-6. [off Fungu Fawatu, W coast of Unguja, Zanzibar, 33-36.5 m] 

Periclimenaeus garthi Bruce, 1976d: 443; Figs 2-4. [Dunidu Island, Male Atoll] 

Periclimenaeus gorgonidarum (Balss, 1913b) 

= Periclimenes gorgonidarum Balss, 1913b: 236. [full description in Balss, 1914b, type locality therein 

as Sagamibai, bei Miski, an Gorgoniden, 20-30 m Tiefe] 

Periclimenaeus hancocki Holthuis, 1951b: 97; Plate 29. [Piñas Bay, Panama, 32 fms] 

Periclimenaeus hebedactylus Bruce, 1970c: 308. [Makunduchi, Zanzibar, 50 fms] 

Periclimenaeus hecate (Nobili, 1904) 

= Coralliocaris hecate Nobili, 1904: 232. [Djibouti] 

Periclimenaeus heronensis Bruce, 2010f: 21; Figs 1-5. [Heron Island reef, north-eastern side, 23°29.990'S 

151°55.601'E, 12 m, dead Acropora] 

Periclimenaeus holthuisi Bruce, 1969b: 159. [Banda, Moluccas, Indonesia, 9 fms] 

Periclimenaeus jeancharcoti Bruce, 1991a: 371; Figs 50-55. [New Caledonia, 21°31'S 166°21'E, 375-450 m] 

Periclimenaeus kottae Bruce, 2005c: 325; Figs 1-3. [Australia, Western Australia, Ashmore Reef, outer 

slope, 6-16 m] 

Periclimenaeus leptodactylus Fujino & Miyake, 1968: 90; Figs 3-5. [Kasari-cho, Amami-oshima Island] 

Periclimenaeus lobiferus Bruce, 1978a: 260; Figs 30-35. [Mozambique Channel, 15°21.7'S 46°12.6'E, 80-85 m] 

Periclimenaeus manihinei Bruce, 1976e: 138; Figs 29-30. [Baie St. Anne, Praslin] 

Periclimenaeus matherae Bruce, 2005c: 331; Figs 4-8. [Australia, Western Australia, Ashmore Reef, West 

Laggon entrance, 2°013.26'S 122°059.28'E, 4-7 m] 

Periclimenaeus maxillulidens (Schmitt, 1936) 

= Periclimenes maxillulidens Schmitt, 1936: 371; Plate 13. [Bonaire, in the Lac (entrance), about 4ft] 

Periclimenaeus minutus Holthuis, 1952c: 134; Figs 57-59. [Banda anchorage, 18-36 m] 

Periclimenaeus mortenseni Bruce, 1993b: 829; Figs 1-2. [Toeal, Kei-Doelah Islands, 2 m] 

Periclimenaeus myora Bruce, 1998c: 394; Figs 4-5. [Myora Reef, North Stradbroke Island, Queensland, 

27°29'S 153°25'E, 3 m] 

Periclimenaeus nobilii Bruce, 1975c: 1577; Figs 13F, 14. [Red Sea] 

Periclimenaeus nielbrucei Bruce, 2006f: 2; Figs 1-6. [Wreck Island, Capricorn Islands, Queensland] 

Periclimenaeus nufu Ďuriš, Horká & Hoc, 2009: 454; Figs 1-5. [Bai Tré site, Lon Island, 12°36'25''N 

109°19'58''E, Van Phong Bay, Vietnam, South China Sea, 5 m, coral rubble]


Periclimenaeus orbitocarinatus Fransen, 2006a: 732; Figs 13-15. [Loyalty Islands, 20°22.25'S 166°10.00'E, 21 

m, in Lissoclinum verrilli] 

Periclimenaeus orontes Bruce, 1986d: 151; Figs 1B, 6-10. [Orontes Reef, Port Essington, 11°03.6'S 132°05.0'E, 

3 m] 

Periclimenaeus pachydentatus Bruce, 1969b: 162. [Great Barrier Reef, Australia, 14°12'N 142°48'E, 19 fms] 

Periclimenaeus pachyspinosus Marin, 2007a: 230-235; Figs 12-15. [South China Sea, Vietnam, Nhatrang 

Bay, Nok Island, 15-20 m] 

Periclimenaeus pacificus Holthuis, 1951b: 85; Plate 25. [Piñas Bay, Panama, 32 fms] 

Periclimenaeus palauensis Miyake & Fujino, 1968: 417; Fig. 5. [Ngadarák Reef] 

Periclimenaeus pearsei (Schmitt, 1932b) 

= Coralliocaris pearsei Schmitt, 1932b: 123; Fig. 1. [in the vicinity of the Tortugas Marine Laboratory 

of the Carnegie Institution, from Spongia officinalis] 

Periclimenaeus pectinidactylus Ďuriš, Horká & Sandford, 2009: 32; Figs 1-4, 5B-E. [off Southwater Caye, 

Belizean Barrier Reef, SE off Dangriga, Belize, Caribbean Sea, 16.4 m, from an unidentified sponge 

(Smenospongia sp.?)] 

Periclimenaeus perlatus (Boone, 1930b) 

= Coralliocaris perlatus Boone, 1930b: 45; Fig. 8. [Gonave Bay, Haiti] 

Periclimenaeus quadridentatus (Rathbun, 1906) 

= Coralliocaris quadridentata Rathbun, 1906: 920; Fig. 69; Plate 24, fig. 1. [Auau Channel, 28-43 fms] 

= Coralliocaris rathbuni Borradaile, 1917: 385. [nomen novum for Coralliocaris quadridentata Rathbun, 

1906] 

Periclimenaeus rastrifer Bruce, 1980a: 27; Figs 12, 13a-b. [Ilot Maître, Nouméa, 20 m] 

Periclimenaeus rhodope (Nobili, 1904) 

= Coralliocaris (Onycocaris) rhodope Nobili, 1904: 233. [Djibouti] 

Periclimenaeus robustus Borradaile, 1915b 

= Periclimenæus robustus Borradaile, 1915b: 213. [Amirante Islands] 

Periclimenaeus serrula Bruce & Coombes, 1995: 111; Figs 3-5. [Orontes Reef, 11°4.5'S 132°4.8'E, 11-12 m] 

Periclimenaeus schmitti Holthuis, 1951b: 90; Plate 27. [Tortugas, Florida] (Fig. 39) 

Periclimenaeus solitus Bruce & Coombes, 1995: 115; Figs 6-7. [Orontes Reef, 11°03.6'S 132°05.0'E, 3 m] 

Fig. 39. Periclimenaeus schmitti Holthuis, 1951. Photo by Arthur Anker.


Periclimenaeus spinosus Holthuis, 1951b: 113; Plate 35. [Costa Rica, near South Viradores Islands, Port 

Culebra, shallow water] 

Periclimenaeus spongicola Holthuis, 1952c: 137; Figs 60-62. [Java Sea, 4°41'S 113°2'E, 28-32 m, in sponge] 

Periclimenaeus storchi Bruce, 1989a: 181; Fig. 5. [Cuaming Island, N of Bohol Island, Visayan Sea] 

Periclimenaeus stylirostris Bruce, 1969b: 167. [South China Sea, 20°34.0'N 113°30.5'E – 20°30.3'N 113°29.0'E, 

49-50 fms] 

Periclimenaeus tchesunovi Ďuriš, 1990a: 615; Figs 1-2. [Genego Islet, North Nilandu Atoll, Maldive Islands, 

20 m] 

Periclimenaeus tridentatus (Miers, 1884b) 

= Coralliocaris ? tridentata Miers, 1884b: 294; Plate 32, fig. C. [Thursday Island, 4-6 fms] 

Periclimenaeus trispinosus Bruce, 1969b: 169. [Mkokotoni, N. end of Zanzibar Island, East Africa, 14-15 fms] 

Periclimenaeus tuamotae Bruce, 1969b: 170. [Mururoa Island, Tuamotu Archipelago, Polynesia] 

Periclimenaeus uropodialis Barnard, 1958: 18; Fig. 6. [Delagoa Bay] 

Periclimenaeus usitatus Bruce, 1969b: 172. [Indian Ocean, 7°46'48"S 39°42'36"E, 11 fms] 

Periclimenaeus wilsoni (Hay, 1917) 

= Coralliocaris wilsoni Hay, 1917: 71. [20 miles off Beaufort, N. C., in 10 to 20 fms] 

Periclimenaeus wolffi Bruce, 1993b: 833; Figs 4-5. [23°20'N 118°30'E, southern Tai-Wan Hai-Hsia, 31 m] 

Periclimenaeus zanzibaricus Bruce, 1969b: 174. [Uroa, east coast of Zanzibar Island, East Africa, littoral] 

Periclimenaeus zarenkovi Ďuriš, 1990a: 620; Figs 3-4. [Genego Islet, North Nilandu Atoll, Maldive Islands; 

0.7 m depth] 

Periclimenella Bruce, 1995 

= Periclimenella Bruce, 1995 (type species Palæmon Petitthouarsii Audouin, 1826, by original designation, 


= Periclimenella Ďuriš & Bruce, 1995 (type species Palæmon Petitthouarsii Audouin, 1826, by original 


Periclimenella petitthouarsii (Audouin, 1826) 

= Palæmon Petitthouarsii Audouin, 1826: 91 (accompanying figures in Savigny, 1826). [Egypt] 

= A.[nchistia] inaequimana Heller, 1861: 28. [im rothen Meere; fully described in Heller, 1862c] 

Periclimenella spinifera (De Man, 1902) 

= Periclimenes pettithouarsii var. spinifera De Man, 1902: 824. [Ternate; Pulo Edam; Bai von Batavia; 

Amboina; Tahiti] 

Periclimenes o.G. Costa, 1844a 

= Pelias Roux, 1831 (type species A.[lpheus] amethystea Risso, 1827, designated by Holthuis, 1955b, 

gender masculine; a junior invalid homonym of Pelias Merrem, 1820 (Reptilia)) 

= Periclimenes O.G. Costa, 1844a (type species Periclimenes insignis O.G. Costa, 1844a, by monotypy, 

gender masculine; name placed on the Official List of Generic Names in Zoology in Opinion 

470 in 1957) 

= Anchistia Dana, 1852a (type species Anchistia gracilis Dana, 1852a, designated by Kingsley, 1880, 


= Dennisia Norman, 1861 (type species Dennisia sagittifera Norman, 1861, by monotypy, gender 

feminine) 

= Ancylocaris Schenkel, 1902 (type species Ancylocaris brevicarpalis Schenkel, 1902, by monotypy, 


= Cristiger Borradaile, 1915b (type species Periclimenes (Cristiger) commensalis Borradaile, 1915b, designated 


Periclimenes acanthimerus Bruce, 2006g: 704; Figs 1-6. [New Caledonia, 21°52.3'S 166°47.9'E, 174 m] 

Periclimenes aegylios Grippa & d’Udekem d’Acoz, 1996 

= Periclimenes sagittifer aegylios Grippa & d’Udekem d’Acoz, 1996: 406; Fig. 1, 2b; Plate 1b. [western 

Italy, Giglio Island, Punta della Secca, 42°21'N 10°54'E, 10 m, associated with Anemonia viridis]


Periclimenes affinis (Zehntner, 1894) 

= Palæmonella affinis Zehntner, 1894: 208. [Amboine, sur Actinometra] 

= Periclimenes (Falciger) brocketti Borradaile, 1915b: 212. [Malé Atoll, Maldive Islands, on brown 

crinoid; lectotype designation by Bruce, 1978c] 

Periclimenes albatrossae Chace & Bruce, 1993: 100; Fig. 20. [Philippines, South China Sea off western Luzon, 

16°33'52"N 119°52'54"E, 315 m] 

Periclimenes albolineatus Bruce & Coombes, 1997: 305; Figs 1-2. [Lee Point, Darwin Harbour, 12°19.0'S 

130°52.5'E, ELWST] 

Periclimenes alcocki Kemp, 1922 

= Periclimenes (Periclimenes) alcocki Kemp, 1922: 154; Figs 21-24. [Laccadive Sea, 9°34'57"N 75°36'30"E, 

406 fms] 

Periclimenes aleator Bruce, 1991a: 315; Figs 10-14. [Loyalty Islands, 20°53.0'S 167°17.0'E, 570-610 m] 

Periclimenes alexanderi Li, 2008: 219; Figs 7-10. [Marquesas Islands, Nuku Hiva, 8°45.1'S 140°15.1'W, 108- 

112 m] 

Periclimenes amethysteus (Risso, 1827) 

= A.[lpheus] amethystea Risso, 1827: 77; Plate 4, fig. 16. [golfe de Nice, rochers peu profonds] 

= Periclimenes insignis O.G. Costa, 1844a: 291. [Napoli] 

Periclimenes andresi Macpherson, 1988: 52; Figs 1-4. [17°15'S 11°27'E, 185 m] 

Periclimenes antipathophilus Spotte, Heard & Bubucis, 1994: 214; Figs 1-6. [Pine Cay, Turks and Caicos 

Islands, near 21°53'N 72°05'W, 37-40 m, from the antipatharian Antipathes gracilis] 

Periclimenes batei (Borradaile, 1917) 

= Palæmonella batei Borradaile, 1917: 358. [Philippine Is.] 

Periclimenes boucheti Li, Mitsuhashi & Chan, 2008: 386; Fig. 1C, 2-4, 5A-B, 16B-D. [8°42.4'N 123°13.4'E, 

123 m] 

Periclimenes bowmani Chace, 1972: 32; Figs 6-7. [Saint Lucia Island, reef south of Marigot Harbour, 2-3 m] 

Periclimenes brevicarpalis (Schenkel, 1902) 

= Ancylocaris brevicarpalis Schenkel, 1902: 563; Plate 13, figs 21a-m. [Makassar] 

= Palæmonella amboinensis Zehntner, 1894: 206; Plate 9, figs 27-27a. [Amboine] 

= Palaemonella aberrans Nobili, 1904: 234. [Djibouti] 

= Harpilius latirostris Lenz, 1905: 380; Plate 47, figs 14-14b. [Zanzibar, Kokotoni, Bawi] 

= Periclimenes potina Nobili, 1905a: 159. [Côtes d’Arabie, sur une algue brune flottante] 

= Periclimenes hermitensis Rathbun, 1914: 655; Plate 1, figs 1-3. [Hermite, under rock] 

Periclimenes brevinaris Nobili, 1906c: 42; Plate 3, fig. 7-7a. [nomen novum for Periclimenes Borradailei Nobili, 

1905a nec Rathbun, 1904] 

Periclimenes brevirostris Bruce, 1991a: 322; Figs 15-20. [22°05.8'S 167°10.3'E, Isle of Pines, 500-550 m] 

Periclimenes brucei Ďuriš, 1990b: 1; Figs 1-2. [Indian Ocean, the Maldive Islands, North Nilandu Atoll, 

Genego Island, 52 m] 

Periclimenes burrup Bruce, 2007d: 113; Figs 5-7. [Burrup Peninsula, 20°31.59'S 116°52.09'E, 11.0 m] 

Periclimenes calcaratus Chace & Bruce, 1993: 104; Fig. 21. [Philippines, Albay Gulf, east of southern Luzon, 

13°12'N 123°49'18"E, 267 m] 

Periclimenes canalinsulae Bruce & Coombes, 1997: 309; Figs 3-4. [Channel Island, 12°33.4'S 130°52.3'E, 

ELWS] 

Periclimenes cannaphilus Komai, Nemoto & Tsuchida, 2010: 800; Figs 2-6. [Kasuga 2 Seamount, 21°36.574'N 

142°38.175'E, 392 m] 

Periclimenes carinidactylus Bruce, 1969a: 254. [Bottle and Glass Rocks, Port Jackson, Sydney Harbour, 

Australia, 20 feet] 

Periclimenes chacei Li, Bruce & Manning, 2004: 542; Figs 27-28. [Xincun, Hainan Island, 1-3 m, with living 

corals] 

Periclimenes colemani Bruce, 1975d: 488; Figs 1-7. [Wistari Reef, Heron Island, Capricorn Group, Queensland, 

Australia, 12 m] 

Periclimenes colesi De Grave & Anker, 2009: 141; Figs 1-26. [Black Coral Wall dive site, Utila, Bay Islands, 

Honduras, 16°05.027'N 86°54.944'W, 15 m, inside osculum of Callyspongia vaginalis]


Periclimenes commensalis Borradaile, 1915b 

= Periclimenes (Cristiger) commensalis Borradaile, 1915b: 211. [Torres Strait, on Comanthus annulatus] 

Periclimenes compressus Borradaile, 1915b 

= Periclimenes (Falciger) compressus Borradaile, 1915b: 212. [Saya de Malha] 

Periclimenes coriolis Bruce, 1985b: 234; Figs 4-7. [14°01.0'N 120°17.1'E, 186-184 m] 

Periclimenes crinoidalis Chace, 1969: 251; Figs 1-2. [Jan Thiel Beach, Curaçao, Netherlands Antilles, 38 m, 

associated with Nemaster grandis] 

Periclimenes cristimanus Bruce, 1965: 487; Figs 1-2. [1°12.7'N 103°43.65'E, Pulau Sudong, near Pulau Salu, 

Singapore] 

Periclimenes crosnieri Li & Bruce, 2006: 681; Figs 16-18. [Indonesia, Tanimbar Islands 7°54'S 132°47'E, 302- 

305 m] 

Periclimenes curvirostris Kubo, 1940b 

= Periclimenes (Periclimenes) curvirostris Kubo, 1940b: 35; Figs 3-5. [Kumano-nada, off Mie Prefecture, 

about 170 fms] 

Periclimenes dardanicola Bruce & Okuno, 2006: 368; Figs 1-4. [Japan. Kuryo, Numazu, Izu Peninsula, Suruga 

Bay, Honshu, 35°01.2'N 138°49.9'E, 21 m] 

Periclimenes delagoae Barnard, 1958: 14; Fig. 4-B. [Delagoa Bay, in coral] 

Periclimenes dentidactylus Bruce, 1984: 7; Figs 1-6. [0°31.4'N 117°50.1'E, 592-595 m] 

Periclimenes difficilis Bruce, 1976e: 111; Figs 15-17. [Baie St. Anne, Praslin] 

Periclimenes digitalis Kemp, 1922 

= Periclimenes (Ancylocaris) digitalis Kemp, 1922: 224; Figs 65; Plate 8, fig. 12. [off Viper Island, Port 

Blair, Andamans, 3-5 fms] 

Periclimenes diversipes Kemp, 1922 

= Periclimenes (Ancylocaris) diversipes Kemp, 1922: 179; Figs 36-39. [Kilakarai, Gulf of Manaar] 

Periclimenes eleftherioui Koukouras & Türkay, 1996: 136; Figs 1-3. [Aegean Sea, between Aspronisi and 

Thera, 36°22'20''N 25°21'20''E, 40-73 m depth] 

Periclimenes exederens Bruce, 1969a: 255. [South China Sea, 20°36.0'N 113°54.2'E – 20°38.8'N 113°57.8'E, 

47-48 fms] 

Periclimenes fenneri Bruce, 2005d: 5; Figs 2-4, 8C. [West Norfolk Ridge, 34°37.20'S 158°59.03'E, 521-539 m] 

Periclimenes finlayi Chace, 1972: 35; Fig. 8. [Off Marigot Bay, St. Lucia, Windward Islands, 165 m, mollusk 

traps] 

Periclimenes forcipulatus Bruce, 1991a: 330; Figs 21-25. [Loyalty Islands, 20°35.0'S 166°54'E, 460 m] 

Periclimenes foresti Bruce, 1981d: 201; Figs 10-11, 17c. [14°00.0'N 120°18.0'E to 14°01.7'N 120°20.2'E, 11 

miles N. of Ambil island, 189-209 m] 

Periclimenes forgesi Li & Bruce, 2006: 686; Figs 19-20. [New Caledonia, 23°02.90'S 166°57.97'E, 500-514 m] 

Periclimenes foveolatus Bruce, 1981d: 196; Figs 6-9, 17a-b, 18b, e. [14°01.0'N 120°15.8'E to 13°59.2'N 

120°18.8'E, 10 miles N. of Ambil Island, 191-188 m] 

Periclimenes fujinoi Bruce, 1990c: 161; Figs 8-11, 39a-b. [Chesterfield Islands, 22°06.9'S 159°24.6'E, 487-610 m] 

Periclimenes gonioporae Bruce, 1989c: 149; Figs 1-3, 4A. [Ras Iwatine, Mombasa, 4°01.15'S 39°43.78'E] 

Periclimenes granulatus Holthuis, 1950c 

= Periclimenes (Periclimenes) granulatus Holthuis, 1950c: 4; Fig. 1, Plate 1. [le travers de Bou-Haroun 

(Départ. D’Alger), environ 100 m] 

Periclimenes granulimanus Bruce, 1978a: 237; Figs 16-19. [Tany-Kely, NW coast near Nosy Bé, on grey 

sand with foraminifera] 

Periclimenes granuloides Hayashi in Baba, Hayashi & Toriyama, 1986: 103; Fig. 18; unnumbered plate. 

[Tosa Bay, Japan, 130 m] 

Periclimenes guarapari De Grave, 2008: 15; Figs 1-5. [Ilha Escalvada, in front of Guarapri City, Espirito 

Santo, Brazil, approx. 20°42'S 04°24'W, 17 m, from Muricea flammea] 

Periclimenes harringtoni Lebour, 1949a 

= Periclimenes (Periclimenes) harringtoni Lebour, 1949a: 1110; Fig. 3. [Harrington Sound] 

Periclimenes hertwigi Balss, 1913b: 235. [Sagamibai, 120 m] 

= Periclimenes (Ancylocaris) gracilirostris Kubo, 1940b: 41; Figs 8-10. [Kumano-nada, off Mie Prefecture, 

about 170 fms]


Periclimenes hongkongensis Bruce, 1969a: 259. [Rocky Harbour, Hong Kong, 22°20.0'N 114°21'E, 14 

fms] 

Periclimenes hydrothermophilus Hayashi & Ohtomi, 2001: 161; Figs 1-4. [Kagoshima Bay, 90-100 m, associated 

with Lamellibranchia satsuma (Polychaeta)] 

Periclimenes imperator Bruce, 1967b: 53; Figs 23-25. [Tumbatu Island, west coast of Zanzibar Island, on 

nudibranch, 3 ft below low water spring tide level] 

Periclimenes incertus Borradaile, 1915b 

= Periclimenes (Cristiger) incertus Borradaile, 1915b: 210. [Maldive Islands] 

= Periclimenes (Periclimenes) impar Kemp, 1922: 147; Figs 16, 17; Plate 3, fig. 1. [Port Blair, Andamans, 

5 fms] 

Periclimenes infraspinis (Rathbun, 1902a) 

= Urocaris infraspinis Rathbun, 1902a: 903. [Gulf of California in Concepcion Bay, Lower California] 

Periclimenes ingressicolumbi Berggren & Svane, 1989: 432; Figs 1-5. [San Salvador Island, 24°02.91'N 

74°32.7'W, 1900ft] 

Periclimenes inornatus Kemp, 1922 

= Periclimenes (Ancylocaris) inornatus Kemp, 1922: 191; Figs 43-46. [Port Blair, Andamans] 

= Periclimenes paraornatus Bruce, 1979a: 207. [nomen nudum] 

Periclimenes investigatoris Kemp, 1922 

= Periclimenes (Periclimenes) investigatoris Kemp, 1922: 160; Fig. 26-27; Plate 5, fig. 6. [Persian Gulf, 

29°20'N 48°47'E, 13 fms] 

Periclimenes involens Bruce, 1996: 234; Figs 13, 28h. [Philippines, off Mindoro, 12°31'N 120°39'E, 92-97 m] 

Periclimenes iridescens Lebour, 1949a 

= Periclimenes (Periclimenes) iridescens Lebour, 1949a: 1112; Figs 4-5. [tow-net about 10 miles outside 

Castle Roads over about 8-100 fms] 

Periclimenes ischiospinosus Bruce, 1991b: 240; Figs 3b, 9-12. [New Caledonia, 21°44'S 166°32'E, 50 m] 

Periclimenes josephi Li, 2008: 224; Figs 11-14. [Austral Islands, Neilson Reef, 27°03.4'S 146°04'W, 130- 

140 m] 

Periclimenes jugalis Holthuis, 1952c 

= Periclimenes (Harpilius) jugalis Holthuis, 1952c: 67; Fig. 26. [anchorage off Djedan Island, east coast 

of the Aru Islands, 13 m] 

Periclimenes kallisto Bruce, 2008d: 89; Figs 1-6. [Keeper Reef, Queensland, 18°44'S 147°16'E, 38 m] 

Periclimenes kempi Bruce, 1969a: 260. [Hurghada, Red Sea, Egypt, 27°14'N 38°50'E, 0.5 fms] 

Periclimenes kornii (Lo Bianco, 1903) 

= Anchistia Kornii Lo Bianco, 1903: 250; Plate 7, fig. 13. [a circa 12.5 chilometri da Punta Carena, direzone 

S.E., 1100 m; a circa 8.5 chilometri da Punta Tragera, direzione O.S.O., 1000 m; a circa 10 

chilometri da Punta Carena, direzione E.N.E., 1000 m] 

Periclimenes laccadivensis (Alcock & Anderson, 1894) 

= Palæmonella laccadivensis Alcock & Anderson, 1894: 157. [Investigator stn 124, Laccadive Sea, 

10°47'45''N 72°40'20''E, 705 fms] 

Periclimenes laevimanus Ďuriš, 2010: 107; Figs 1-6. [Vietnam, Nhatrang Bay, 12°10'06''N 109°17'45''E, 14 m, 

on hydroid cf. Lytocarpia sp.] 

Periclimenes latipollex Kemp, 1922 

= Periclimenes (Periclimenes) latipollex Kemp, 1922: 150; Fig. 18; Plate 4, fig. 3. [Mergui Archipelago, 

12°15'20"N 97°10'10"E, 62 fms] 

Periclimenes lepidus Bruce, 1978a: 244; Figs 20-24. [NW coast of Madagascar near Nosy Bé, 40 m] 

Periclimenes leptodactylus Bruce, 1991a: 338; Figs 26-30. [Loyalty Islands, 20°37.8'S 167°02.7'E, Lifou/Uvéa, 

825-370 m] 

Periclimenes leptopus Kemp, 1922 

= Periclimenes (Ancylocaris) leptopus Kemp, 1922: 173; Figs 31-33. [Brigade Creek, Port Blair, Andamans, 

2-5 fms] 

Periclimenes leptunguis Li, Mitsuhashi & Chan, 2008: 393; Figs 6-8, 17B. [8°49.9'N 123°34.9'E, 437-443 m] 

Periclimenes longimanus (Dana, 1852a) 

= Anchistia longimana Dana, 1852a: 25. [Type locality not indicated]


Periclimenes longipes (Stimpson, 1860a) 

= Urocaris longipes Stimpson, 1860a: 39. [insulam ”Ousima”, prof 20 org] 

Periclimenes loyautensis Li & Bruce, 2006: 700; Fig. 24 [Loyalty Islands, 21°02.30'S 167°31.60'E, 430 m] 

Periclimenes macrophthalmus Fujino & Miyake, 1970a 

= Periclimenes (Harpilius) macrophthalmus Fujino & Miyake, 1970a: 250; Figs 3-5. [32°36.7'N 127°42.8'E, 

145 m] 

Periclimenes madreporae Bruce, 1969a: 262. [Erskine Island, Capricorn Group, Great Barrier Reef, Queensland, 

Australia, 3-6 fms] 

Periclimenes magnus Holthuis, 1951b 

= Periclimenes (Harpilius) magnus Holthuis, 1951b: 52; Plate 15. [Gulf of Mexico, off Aransas, Texas, 

27°40'N 96°34'W, 27.5 fms] 

Periclimenes mahei Bruce, 1969a: 263. [North West Bay, Mahé, Seychelles Islands, 4°36'15"S 55°26'01"E, 

1-2 fms] 

Periclimenes maldivensis Bruce, 1969a: 264. [Suvadiva Atoll, Maldive Islands] 

Periclimenes manihine Bruce, 2006h: 46; Figs 1-4. [off Ras Ngomeni, Kenya, 3°01'S 40°22.5'E to 2°59.5'N 

40°22.6'E, 245-256 m] 

Periclimenes mclaughlinae Fransen, 2006b: 380; Figs 1-6. [Philippines, Cebu Strait, W of Bohol, NW side of 

Cabilao Island, La Estrella Dive Resort, 9°53.20'N 123°45.53'E, 5 m, on Phyllacanthus imperialis] 

Periclimenes mclellandi Heard & Spotte, 1997: 39; Figs 1-5. [forereef slope west of Pine Cay, Turks and 

Caicos Islands, British West Indies, on gorgonian Pseudopterogorgia americana, 29 m] 

Periclimenes meyeri Chace, 1969: 255; Figs 3-4. [Jan Thiel Beach, Curaçao, Netherlands Antilles, 24 m, associated 

with Nemaster?] 

Periclimenes milleri Bruce, 1986e: 637; Figs 1-5. [24°02.75'N 74°32.53'W, 527 m] 

Periclimenes murcielagensis Vargas, 2000: 17; Figs 1-5. [Isla San Pedrito, Archipiélago de las Islas Murciélago, 

Guanacaste, Costa Rica, 25 m] 

Periclimenes nevillei Bruce, 2010h: 45; Figs 1-6. [Espiritu Santo, Vanuatu, 8 m] 

Periclimenes ngi Li, Mitsuhashi & Chan, 2008: 397; Figs 9-11, 17C-D. [9°27.4'N 123°49.4'E, 273-302 m] 

Periclimenes nomadophila Berggren, 1994a: VII.3; Figs 1-5. [Inhaca Island, Mocambique] 

= Periclimenes nomadophila Berggren, 1994b: 783; Figs 1-5. [Inhaca Island, Mocambique] 

Periclimenes novaffinis Bruce & Coombes, 1997: 315; Fig. 6. [off East Point, Darwin Harbour, 12°25.92'S 

130°48.72'E, 8 m] 

Periclimenes obscurus Kemp, 1922 

= Periclimenes (Periclimenes) obscurus Kemp, 1922: 144; Figs 14-15. [Springhaven, Madras Harbor] 

Periclimenes ordinarius Bruce, 1991a: 344; Figs 31-35. [New Caledonia, 18°04.0'S 163°27.5'E, 260 m] 

Periclimenes ornatellus Bruce, 1979d: 219; Figs 4-6; Plate 1C-E. [Enewetak Atoll] 

Periclimenes ornatus Bruce, 1969a: 266. [Lung Ha Wan, N.T., Hong Kong, 22°18.5'N 114°18.2'E, 2 fms] 

Periclimenes paivai Chace, 1969: 259; Figs 5-7. [Cananéia, Estado de São Paulo, Brazil] 

Periclimenes pandionis Holthuis, 1951b 

= Periclimenes (Periclimenes) pandionis Holthuis, 1951b: 41; Plate 11. [Gulf Stream off Key West, 

Florida, 24°21'55"N 81°58'25"W, 98 fms] 

Periclimenes panglaonis Li, Mitsuhashi & Chan, 2008: 402; Figs 12-13, 14A-D, 17E. [9°38.8'N 123°45.9'E, 

418-477 m] 

Periclimenes paralcocki Li & Bruce, 2006: 707; Fig. 27. [Tuvalu, Banc Bayonnaise, 11°54'S 179°32'W, 600 m] 

Periclimenes paraleator Li & Bruce, 2006: 711; Fig. 28. [New Caledonia, 18°56.3'S 163°12.9'E, 520 m] 

Periclimenes paraparvus Bruce, 1969a: 267. [South China Sea, 20°28.2'N 112°46.5'E to 20°23.3'N 112°52.2'E, 

46-48 fms] 

Periclimenes parvispinatus Bruce, 1990c: 154; Figs 3-6, 39d. [New Caledonia, S.W. Récif Jouan, 200 m] 

Periclimenes parvus Borradaile, 1898: 384. [New Britain] 

Periclimenes patae Heard & Spotte, 1991: 40; Figs 1-4. [fringing reef northwest of Pine Cay, Turks and 

Caicos Islands, British West Indies, near 21°53'N 072°05'W, on gorgonian, 3-4 m] 

Periclimenes pauper Holthuis, 1951b 

= Periclimenes (Harpilius) pauper Holthuis, 1951b: 50; Plate 14. [Venezuela, Isla Cubagua, rocky 

shore]


Periclimenes pectiniferus Holthuis, 1952c 

= Periclimenes (Periclimenes) pectiniferus Holthuis, 1952c: 48; Figs 15-16. [Kabala dua Island, Borneo 

Bank, 22 m] 

Periclimenes pectinipes Bruce, 1991a: 351; Figs 36-40, 75. [New Caledonia, 23°41.2'S 168°00.5'E, Norfolk 

Ridge, 280 m] 

Periclimenes perlucidus Bruce, 1969a: 268. [South China Sea, 16°06.5'N 114°41.5'E to 16°05.8'N 114°38.2'E, 

43-44 fms] 

Periclimenes perryae Chace, 1942b 

= Periclimenes (Periclimenes) perryae Chace, 1942b: 82; Plate 24. [off Sanibel Island, Florida, about 5.5 

fms, on Astrophyton muricatum] 

Periclimenes perturbans Bruce, 1978a: 253; Figs 25-26. [NW coast of Madagascar, near Nosy Bé, 40 m] 

Periclimenes pholeter Holthuis, 1973a: 30; Figs 10-11; Plate 1, fig. 1. [Ras Muhammad Crack, Ras Muhammad, 

southern tip of Sinai Peninsula, in pool in crack in coral rock about 150 m from the sea] 

Periclimenes platydactylus Li, 2008: 231; Figs 16-19. [Marquesas Islands, Nuku Hiva, 8°45.1'S 140°15.1'W, 

108-112 m] 

Periclimenes platyrhynchus Bruce, 1991a: 358; Figs 41-44. [New Caledonia, 19°04.0'S 163°27.0'E, 260 m] 

Periclimenes polynesiensis Li, 2008: 236; Figs 20-23. [Marquesas Islands, Nuku Hiva, 8°45.1'S 140°15.1'W, 

108-112 m] 

Periclimenes poriphilus Bruce, 2010i: 53; Figs 2-6. [Heron Island reef, north-eastern side, 23°25.990'S 

151°55.601'E, 10 m] 

Periclimenes poupini Bruce, 1989d: 852; Figs 1-5, 6A. [Tubuai, 23°19'S 142°22'W, 430-520 m] 

Periclimenes pseudalcocki Li & Bruce, 2006: 716; Fig. 30. [Indonesia, Kai Islands, 5°17'S 132°50'E, 315-349 m] 

Periclimenes rathbunae Schmitt, 1924a: 70; Figs 5-6. [Spanish Port] 

Periclimenes rectirostris Bruce, 1981d: 204; Figs 12-15. [13°53.1'N 120°08.9'E to 13°53.3'N 120°10.7'E, 3 

miles NE. of town of Lubang, 134-129 m] 

Periclimenes rex Kemp, 1922 

= Periclimenes (Periclimenes) rex Kemp, 1922: 158; Fig. 25; Plate 5, fig. 5. [Ross Channel, Port Blair, 

Andamans, 8 fms] 

Periclimenes richeri Bruce, 1990c: 181; Figs 20, 39f. [New Caledonia, 24°54.5'S 168°23.3'E, 527 m] 

Periclimenes ruber Bruce, 1982c: 197; Figs 3-5, 8f. [Bribie Passage, Pumicestone Channel, Queensland, 

Australia, in 8 m depth, from crinoid host Zygometra microdiscus] 

Periclimenes sagittifer (Norman, 1861) 

= Dennisia sagittifer Norman, 1861: 278; Plate 13, figs 8-13. [St. Catherine’s Bay, Jersey, 4 fms] 

Periclimenes sandybrucei Mitsuhashi & Chan, 2009: 920; Figs 1-4. [24°30.55'N 122°5.78'E, 399-397 m] 

Periclimenes sandyi De Grave, 2009: 830; Figs 1-4. [Crawl Cay, Bocas del Toro Archipelago, Caribbean 

Panama, 09°15.028'N 82°07.897'W, 10 m, from unidentified sponge] 

Periclimenes sarkanae Bruce, 2007e: 61; Figs 1-5. [Fisherman Island, near mouth of Brisbane River, Moreton 

Bay, 27°22'S 153°10'E, 0.2-0.5 m] 

Periclimenes scriptus (Risso, 1822) 

= A.[lpheus] scriptus Risso, 1822: 247. [mer de Nice] 

? = Periclimenes elegans Gourret, 1884: 15; nec Anch.[istia] elegans Paul’son, 1875. [nomen nudum] 

= Urocaris de Mani Balss, 1916: 29; Fig. 10. [Französisch-Kongo, Setté Cama, salzwasser] 

Periclimenes sibogae Holthuis, 1952c 

= Periclimenes (Harpilius) sibogae Holthuis, 1952c: 73; Figs 28-29. [Banda anchorage, 9-36 m] 

Periclimenes sinensis Bruce, 1969a: 270. [Hong Kong] 

= Periclimenes (Periclimenes) setoensis Fujino & Miyake, 1969a: 149; Figs 4, 5. [Shisô-jima Island, Tanabe 

Bay, Wakayama Prefecture, 5 m] 

Periclimenes soror Nobili, 1904: 232. [Djibouti] 

? = Periclimenes parasiticus Borradaile, 1898: 384. [British New Guinea] 

= Periclimenes (Cristiger) frater Borradaile, 1915b: 210. [Seychelles] 

= Periclimenes bicolor Edmondson, 1935a: 10; Fig. 3. [Kaneohe Bay, Oahu, on Linckia multiflora in shallow 

water]


Periclimenes tangeroa Bruce, 2005d: 12; Figs 5-7, 8D. [South Norfolk Ridge, 34°09.14'S 171°27.95'E, 242-254 m] 

Periclimenes tenellus (Smith, 1882) 

= Anchistia tenella Smith, 1882: 55; Plate 9, figs 1-1b. [Blake stn 316, 32°07'N 78°37'05"W, 229 fms] 

Periclimenes terangeri Bruce, 1998c: 387; Fig. 1. [Polka Point, North Stradbroke Island, Queensland, 

27°30'S 153°23'E, 0.5 m] 

Periclimenes thermohydrophilus Hayashi & Ohtomi, 2001: 161; Figs 1-4. [Kagoshima Bay, 90-100 m] 

Periclimenes toloensis Bruce, 1969a: 273. [Ap Chau, Tolo Channel, Hong Kong, 5-15 fms] 

Periclimenes tonga Bruce, 1988e: 23; Figs 1-5. [Nuapapa Island (southside), Vava’u Group, Tonga, 5-50 ft] 

Periclimenes uniunguiculatus Bruce, 1990c: 167; Figs 12-15, 39e. [New Caledonia, 23°06'S 167°47'E, 540- 

600 m] 

Periclimenes vanellus Fransen, 2010: 242; Figs 1-6. [Indonesia, off Halmahera mainland, Teluk Dodinga, 

Karang Galiasa Besar E, 0°50'45.6''N 127°35'7.4''E, on Ianthella basta, 15 m] 

Periclimenes vaubani Bruce, 1990c: 174; Figs 16-9, 38a-d, 39g. [New Caledonia, 23°38'S 167°42'S, 470 m] 

Periclimenes veleronis Holthuis, 1951b 

= Periclimenes (Harpilius) veleronis Holthuis, 1951b: 67; Plate 20, figs a-h. [off beach at La Libertad, 4 fms] 

Periclimenes vicinus Li, 2008: 242; Figs 24-27. [Austral Islands, Tubuai, 23°21.4'S 149°34.2'W, 600-1200 m] 

Periclimenes watamuae Bruce, 1976f: 16; Figs 5-6. [Watamu Park, Kenya, 3°22.0'S 40°00.5'E, 2 m, from alcyonarian; 

information from NCB-Naturalis collection label] 

Periclimenes wirtzi d’Udekem d’Acoz, 1996a: 133; Figs 1-7. [Madeira, small seamount in front of the bay 

of Machico, from Antipathes bushes at 30-40 m] 

Periclimenes yaldwyni Holthuis, 1959b 

= Periclimenes (Harpilius) yaldwyni Holthuis, 1959b: 197. [nomen novum for Periclimenes batei 


= Brachycarpus audouini Spence Bate, 1888: 798; Plate 129, fig. 5. [Challenger stn 167A, 41°4'S 174°19'E, 

off New Zealand, 10 fms] 

= Periclimenes batei Holthuis, 1950a: 22. [nec Palæmonella batei Borradaile, 1917] 

Periclimenes yucatanicus (Ives, 1891) 

= Palæmonella Yucatanica Ives, 1891: 183; Plate 5, fig.8. [off Progreso, 20ft] 

Periclimenes zanzibaricus Bruce, 1967b: 62; Figs 26-29. [Fawatu Reef, off west coast of Zanzibar Island, on 

echinoid at low water spring tide level on sand flats] 

Periclimenes zevinae Ďuriš, 1990b: 4; Figs 3-4. [Indian Ocean, the Maldive Islands, North Nilandu Atoll, 

Genego Island, 50 m] 

Periclimenoides Bruce, 1990f 

= Periclimenoides Bruce, 1990f (type species Periclimenaeus odontodactylus Fujino & Miyake, 1968, by 


Periclimenoides odontodactylus (Fujino & Miyake, 1968) 

= Periclimenaeus odontodactylus Fujino & Miyake, 1968: 85; Figs 1-2. [Ushibuka, Amakusa Island, 

west of Kyushu, Japan] 

Periclimenoides socotrae Bruce, 2006a: 22; Figs 6-9. [12°40.264'N 53°27.204'E, SW of Qualansiyah, NWcoast, 

5-7 m] 

Philarius Holthuis, 1952c 

= Philarius Holthuis, 1952c (type species Harpilius Gerlachei Nobili, 1905a, by original designation, 


Philarius albimaculatus Marin & Anker, 2011: 22; Figs 16-18, 19E-F. [Madagascar, Nosy-Bé W side of Sakatia, 

13.2989°S 48.1472°E, 3-8 m] 

Philarius gerlachei (Nobili, 1905a) 

= Harpilius Gerlachei Nobili, 1905a: 160. [Golfe Persique] 

Philarius imperialis (Kubo, 1940c) 

= Harpilius imperialis Kubo, 1940c: 1; Figs 1-3. [Nankin-Hama, Haha-zima, Bonin Islands] 

Philarius lifuensis (Borradaile, 1898) 

= Periclimenes lifuensis Borradaile, 1898: 384. [Loyalty Islands]


Philarius minor Marin & Anker, 2011: 13; Figs 13-15, 19C-D. [Australia, Great Barrier Reef off S Queensland, 

Heron Island, 23.47262°S 151.95988°E, 14-16 m] 

Philarius polynesicus Marin & Anker, 2011: 6; Figs 4-8, 9A-E. [French Polynesia, Society Islands, Moorea, 

S of Vaiare Pass, 17.5303°S 149.7621°W, 5-6 m] 

Philarius rufus Marin & Anker, 2011: 13; Figs 10-14, 19A-B. [Australia, Great Barrier Reef off S Queensland, 

Heron Island, 23.45322°S 151.90045°E, 24-26 m] 

Phycomenes Bruce, 2008e 

= Phycomenes Bruce, 2008e (type species Phycomenes zostericola Bruce, 2008e, by original designation 


Phycomenes cobourgi (Bruce & Coombes, 1995) 

= Periclimenes cobourgi Bruce & Coombes, 1995: 125; Figs 10-11. [Table Head, 3 m, on gorgonian 

Rumphella aggregata] 

Phycomenes indicus (Kemp, 1915) 

= Urocaris indica Kemp, 1915: 275; Fig. 26; Plate 13, fig. 9. [Chilka Lake; Ennur and Adyar River near 

Madras; fringing coral-reef at Kilakarai, northern end of Gulf of Manaar] 

Phycomenes siankaanensis (Martínez-Mayén & Román-Contreras, 2006) 

= Periclimenes siankaanensis Martínez-Mayén & Román-Contreras, 2006: 33; Figs 1-3. [Cayo Culebras, 

Bahía de la Ascensión, Quintana Roo, México, 1 m] 

Phycomenes sulcatus (Ďuriš, Horká & Marin, 2008) 

= Periclimenes sulcatus Ďuriš, Horká & Marin, 2008: 36; Figs 1-6. [S of Nhatrang Bay, Vietnam, 

12°09'42.0''N 109°12'13.1''E, 0.1-0.2 m] 

Phycomenes zostericola Bruce, 2008e: 221; Figs 1-7. [Loder’s Creek, Labrador, Queensland] 

Fig. 40. Platypontonia hyotis Hipeau-Jacquotte, 1971. Photo by Arthur Anker.


Pinnotheronia Marin & Paulay, 2010 

= Pinnotheronia Marin & Paulay, 2010 (type species Pinnotheronia rumphiusi Marin & Paulay, 2010, by 


Pinnotheronia rumphiusi Marin & Paulay, 2010: 38; Figs 1-6. [Pacific Ocean, Palau, Koror Island, Iwayama 

Bay, in mantle cavity of Periglypta crispata, 0-10 m] 

Platycaris Holthuis, 1952c 

= Platycaris Holthuis, 1952c (type species Platycaris latirostris Holthuis, 1952c, by monotypy, gender 

feminine) 

Platycaris latirostris Holthuis, 1952c: 173; Figs 85-86. [Ende, Flores] 

Platypontonia Bruce, 1968b 

= Platypontonia Bruce, 1968b (type species Pontonia ? brevirostris Miers, 1884a, by original designation 


Platypontonia brevirostris (Miers, 1884a) 

= Pontonia? brevirostris Miers, 1884a: 562; Plate 51, figs Bb-b'''. [Seychelles, 12 fms, in ”clamp shells” 

(bivalves?)] 

Platypontonia hyotis Hipeau-Jacquotte, 1971: 126; Figs 1-7. [Madagascar, S of Tuléar, outer slope of Banc 

de Belosa, in Pycnodonta hyotis; according to Fransen, Holthuis & Adema, 1997] (Fig. 40) 

= Platypontonia pterostreae Suzuki, 1971: 5; Figs 3-4; Plate 3. [Hatsu-shima, Sagami Bay] 

Plesiomenaeus Bruce, 2009b 

= Plesiomenaeus Bruce, 2009b (type species Plesiomenaeus poorei Bruce, 2009b, by original designation 


Plesiomenaeus poorei Bruce, 2009b: 26; Figs 1-7. [Chukwani, Unguja, Zanzibar, 6°13'60''S 30°13'00''E, at 

low water spring tide] 

Plesiopontonia Bruce, 1985b 

= Plesiopontonia Bruce, 1985b (type species Plesiopontonia monodi Bruce, 1985b, by original designation 


Plesiopontonia monodi Bruce, 1985b: 250; Figs 13-17. [13°49.6'N 120°51'E, 299-320 m] 

Pliopontonia Bruce, 1973c 

= Pliopontonia Bruce, 1973c (type species Pliopontonia furtiva Bruce, 1973c, by original designation 


Pliopontonia furtiva Bruce, 1973c: 99; Figs 1-5; Plate 1. [Ras Iwatine, Mombasa, Kenya, 4°00.55'S 39°44.17'E, 

1 m] 

Pliopontonia harazakii Okuno, 2009b: 106; Figs 1-4, 5A-B, 6. [Onoaida, Yaku-shima Island, Osumi Islands, 

Japan, 30°14.1'N 130°33.2'E, 3 m] 

Pontonia latreille, 1829 

= Alciope Rafinesque, 1814 (type species Alciope heterochelus Rafinesque, 1814, gender feminine; an 

invalid senior subjective synonym of P.[ontonia] flavomaculata Heller, 1864; name suppressed for the 

purposes of the Principle of Priority, but not for those of the Principle of Homonymy, placed on the 

Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 522 in 1958) 

= Pontonia Latreille, 1829 (type species Palaemon pinnophylax Otto, 1821, gender feminine; designated 

under the plenary power of the ICZN and placed on the Official List of Generic Names in Zoology 


Pontonia chimaera Holthuis, 1951b: 125; Plate 39. [Panama, Archipielago de las Perlas, north side of Pedro 

Gonzalez Island, west of Cocal] 

Pontonia domestica Gibbes, 1850: 196. [coast of South Carolina] 

= Pontonia occidentalis Gibbes, 1848: 16. [South Carolina; nomen nudum]


Fig. 41. Pontonia pinnae Lockington, 1878. Photo by Arthur Anker. 

Pontonia longispina Holthuis, 1951b: 128; Plate 40. [Lower California, Mexico, Angel de la Guardia Island, 

Puerto Refugio, shore] 

Pontonia manningi Fransen, 2000: 101; Figs 1-3, 4a-f. [Caribbean Sea, 25 miles N of Margarita Island, 36.5 m] 

Pontonia margarita Verrill, 1869: 245. [Bay of Panama, in Pearl Oyster (Margaritophora fimbriata)] 

= Coralliocaris Camerani Nobili, 1901b: 3. [Isola Flamenco] 

Pontonia mexicana Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, fig. 12. [Mexico; 

see Holthuis, 1951b] 

= Pontonia grayi Rathbun, 1901: 122 (partim); Fig. 5. [between San Antonio Bridge and San Geronimo, 

San Juan, Porto Rico] 

Pontonia panamica Marin & Anker, 2008: 503; Figs 1-7. [Panama, Playa Venao, 10 km west of Panama City, 

lower rocky intertidal, in ascidian, Ascidia cf. interrupta] 

Pontonia pilosa Fransen, 2002: 147; Figs 91-99; Plate 4. [East Atlantic, Cape Verde Islands, Sao Tiago Island, 

Tarrafal, from Pseudochama radians] 

Pontonia pinnae Lockington, 1878a: 163. [interior of the shell of the common Pinna, Gulf of California, 

Angelas Bay, Mulege Bay, San José Island] (Fig. 41) 

Pontonia pinnophylax (Otto, 1821) 

= Palaemon pinnophylax Otto, 1821: 12. [Neapoli in Pinna nobili marem; fully described in Otto, 1828 

as Alpheus pinnophylax] 

= A.[lpheus] Tyrhenus Risso, 1816: 94; Plate 2, fig. 2. [environs de Nice, dans les valves du jambonneau 

marin] 

= Pontonia parasitica Roux, 1831: 26. [sur les cotes de Morée, dans l’interieur de la Pinna Nobilis] 

= Pontonia custos Guérin, 1832: 36; Plate 37, fig. 1. [Sapience dans la Pinne marine (Pinna nobilis)] 

= Pontonia heterochelis Guérin, 1832: 37. [see Holthuis, 1947b] 

Pontonia simplex Holthuis, 1951b: 135; Plate 42. [Jalisco, Mexico, Tenacatita Bay, lagoon]


Pontonides Borradaile, 1917 

= Pontonides Borradaile, 1917 (type species Pontonia maldivensis Borradaile, 1915b, by monotypy, gender 

masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Pontonides ankeri Marin, 2007b: 3; Figs 1-3, 10B, 11F-G. [Vietnam, Nhatrang Bay, Nok Island, 15 m] 

Pontonides asperulatus Bruce, 2005b: 359; Figs 13-16, 22E. [Horseshoe Reef, Loloata Island, 16 m] 

Pontonides loloata Bruce, 2005b: 367; Figs 17-21, 22F. [Loloata Island] 

Pontonides maldivensis (Borradaile, 1915b) 

= Pontonia maldivensis Borradaile, 1915b: 213. [Fadiffolu Atoll, Maldive Islands] 

Pontonides sibogae Bruce, 2005b: 377. [Sape Strait, Indonesia, 23°30'S 119°4.6'E, 70 m] 

Pontonides tatianae Marin, 2007b: 13; Fig. 8. [Vietnam, Nhatrang Bay, Dun Island, 40 m] 

Pontonides unciger Calman, 1939: 213; Figs 6-7. [Red Sea, 13°31'00''N 42°31'00''E, 55 m] 

Pontoniopsides Bruce, 2005e 

= Pontoniopsides Bruce, 2005e (type species Pontoniopsis paulae Gore, 1981, by original designation 


Pontoniopsides paulae (Gore, 1981) 

= Pontoniopsis paulae Gore, 1981: 139; Fig. 1. [Carysfort Reef, off Key Largo, Monroe County, Florida, 

25°10.30'N 80°12.82'W, 62.5 m] 

Pontoniopsis Borradaile, 1915b 

= Pontoniopsis Borradaile, 1915b (type species Pontoniopsis comanthi Borradaile, 1915b, by monotypy, 


1959) 

Pontoniopsis comanthi Borradaile, 1915b: 213. [Torres Straits, on Comanthus] 

Poripontonia Fransen, 2003 

= Poripontonia Fransen, 2003 (type species Poripontonia dux Fransen, 2003, by original designation 


Poripontonia dux Fransen, 2003: 130; Figs 1-6. [Tanjung Benoa, Loloan Benoa, 08°45'46''S 115°14'01''E, to 25 m] 

Poripontonia cornuta Marin, 2007a: 235; Figs 16-18. [South China Sea, Vietnam, Nhatrang Bay, Nok Island, 

10 m] 

Propontonia Bruce, 1969c 

= Propontonia Bruce, 1969c (type species Propontonia pellucida Bruce, 1969c, by original designation 


Propontonia pellucida Bruce, 1969c: 142; Figs 1-5. [Remire Island, Amirante Islands, Seychelles, 05°04'S 

53°22'E, 1.5 m] 

Pseudoclimenes Bruce, 2008c 

= Pseudoclimenes Bruce, 2008c (type species Pseudoclimenes holthuisi Bruce, 2008c, by original designation 


Pseudoclimenes holthuisi Bruce, 2008c: 17; Figs 10-13. [Jurien Bay, 29°48'25''S 114°25'52''E to 29°48'53''S 

114°25'55''E, 113-114 m] 

Pseudocoutierea Holthuis, 1951b 

= Pseudocoutièrea Holthuis, 1951b (type species Pseudocoutièrea elegans Holthuis, 1951b, by original 


Pseudocoutierea antillensis Chace, 1972: 43; Fig. 11. [Saba Bank, 17°28'N 63°13'W, dredged in 13 m] 

Pseudocoutierea conchae Criales, 1981: 174; Figs 6-9. [Bahía Concha, Colombia, 11°18'N 74°10'W, 15 m] 

Pseudocoutierea dotae De Grave, 2007b: 30; Figs 1-5. [Hospital Point, Cayo Solarte, Bocas Province, Panama, 

09°20.016'N 082°13.133'W, 15 m, on Stichopathes lutkeni] 

Pseudocoutierea edentata Criales, 1981: 168; Figs 2-5. [Bahía Concha, Colombia, 11°18'N 74°10'W, 18 m]


Pseudocoutièrea elegans Holthuis, 1951b: 182; Plate 57. [southern California, Santa Catalina Island, ½ mile 

E of Long Point, 45-50 fms] 

Pseudocoutierea wirtzi d’Udekem d’Acoz, 2000: 70; Figs 1-12. [Cape Verde Islands, São Tiago Island, 20-30 

m, on Leptogorgia gaini] 

Pseudopontonia Bruce, 1992d 

= Pseudopontonia Bruce, 1992d (type species Pontonia minuta Baker, 1907, by original designation 


Pseudopontonia minuta (Baker, 1907) 

= Pontonia minuta Baker, 1907: 189; Plate 24, Figs 9-12. [unidentified locality in South Australia; see 

Bruce, 1992d] 

Pseudopontonides Heard, 1986 

= Pseudopontonides Heard, 1986 (type species Neopontonides principes Criales, 1980, by original designation 


Pseudopontonides plumosus Snijders & Fransen, 2010: 7; Figs 1-6. [south coast, Otrobanda, harbour, Superior 

Producer, Curaçao, Lesser Antilles, 12°05'21.5''N 068°56'35.5''W, on Antipathes spec., depth 29 

m] 

Pseudopontonides principis (Criales, 1980) 

= Neopontonides principis Criales, 1980: 75; Figs 25-29. [Awa di Oostpunt, Curaçao, 18 m] 

Pseudoveleronia Marin, 2008b 

= Pseudoveleronia Marin, 2008b (type species Veleronia laevifrons Holthuis, 1951b, by original designation 


Pseudoveleronia laevifrons (Holthuis, 1951b) 

= Veleronia laevifrons Holthuis, 1951b: 199; Plate 63, figs f-m. [Galapagos Islands, Ecuador, Gardner 

Bay, Hood Island, 4 fms] 

Rapipontonia Marin, 2007c 

= Rapipontonia Marin, 2007c (type species Rapipontonia paragalene Marin, 2007c, by original designation, 


Rapipontonia galene (Holthuis, 1952c) 

= Periclimenes (Harpilius) galene Holthuis, 1952c: 62; Fig. 24. [Amboina, pier, 0-2 m; Islet near Menado] 

Rapipontonia hydra Marin, 2009b: 10; Figs 5-7. [Pacific Ocean, Vietnam, Nhatrang Bay, on pile under pier 

of MMA, depth 2 m, on hydroid Lytocarpia sp.] 

Rapipontonia paragalene Marin, 2007c: 777; Figs 1-6. [Vietnam, South China Sea, Nhatrang Bay, on pile 

under pier of Marine Military Academy, 2 m, on hydroid Lytocarpia sp.] 

Rapipontonia platalea (Holthuis, 1951a) 

= Periclimenes (Harpilius) platalea Holthuis, 1951a: 157; Fig. 32. [off French Guinea, 9°23'N 15°07'W, 

30-34 m] 

Rostronia Fransen, 2002 

= Rostronia Fransen, 2002 (type species Pontonia stylirostris Holthuis, 1952c, by original designation 


Rostronia stylirostris (Holthuis, 1952c) 

= Pontonia stylirostris Holthuis, 1952c: 169; Figs 82-84. [between Misool and New Guinea, 1°42.5'S 

130°47.5'W, 32 m] 

Sandimenes li, 2009 

= Sandimenes Li, 2009 (type species Periclimenes hirsutus Bruce, 1971e, by original designation and 


Sandimenes hirsutus (Bruce, 1971e) 

= Periclimenes hirsutus Bruce, 1971e: 91; Figs 1-6. [Nukulau Island, Laucala Bay Suva, Viti Levu, Fiji]


Sandyella Marin, 2009c 

= Sandyella Marin, 2009c (type species Chacella tricornuta Hendrickx, 1990a, by original designation, 


Sandyella bicornuta (Li & Poupin, 2009) 

= Chacella bicornuta Li & Poupin, 2009: 581; Figs 1-4. [Clipperton, 10°19.22'N 109°13.38'W, 55 m, 

outer reef] 

Sandyella mclaughlinae (Li, 2006a) 

= Chacella mclaughlinae Li, 2006a: 360; Figs 1-4. [Clipperton Island, 10°19.22'N 109°13.38'W, 55 m] 

Sandyella sexicornuta (Li & Poupin, 2009) 

= Chacella sexicornuta Li & Poupin, 2009: 587; Figs 9-12. [Clipperton, 10°18.03'N 109°13.77'W, 54 m, 

outer reef] 

Sandyella tricornuta (Hendrickx, 1990a) 

= Chacella tricornuta Hendrickx, 1990a: 326; Figs 1-3. [Mexico, Nayarit, Islas Tres Marias, east coast 

of Isla Maria Madre, 21°38'N 106°32'W, 30-40 m] 

Sandyella quadricornuta (Li & Poupin, 2009) 

= Chacella quadricornuta Li & Poupin, 2009: 584; Figs 5-8. [Clipperton, 10°19.22'N 109°13.38'W, 55 m, 

outer reef] 

Stegopontonia Nobili, 1906b 

= Stegopontonia Nobili, 1906b (type species Stegopontonia commensalis Nobili, 1906b, by monotypy, gender 


Stegopontonia commensalis Nobili, 1906b: 258. [lagon de Hao, commensal de Echinothrix turcarum] 

Tectopontonia Bruce, 1973d 

= Tectopontonia Bruce, 1973d (type species Tectopontonia maziwiae Bruce, 1973d, by original designation 


Tectopontonia maziwiae Bruce, 1973d: 172; Figs 1-4. [Maziwi Island, off Pangani, Tanganyika, 5°30.0'S 

39°04.1'E] 

Thaumastocaris Kemp, 1922 

= Thaumastocaris Kemp, 1922 (type species Thaumastocaris streptopus Kemp, 1922, by original designation 


Thaumastocaris streptopus Kemp, 1922: 244; Figs 78-80. [Noumea, New Caledonia] 

Tuleariocaris Hipeau-Jacquotte, 1965 

= Tuleariocaris Hipeau-Jacquotte, 1965 (type species Tuleariocaris holthuisi Hipeau-Jacquotte, 1965, by 


Tuleariocaris holthuisi Hipeau-Jacquotte, 1965: 248; Plates 1-5. [Tuléar, Madagascar, on echinoids Echinometra 

and Stomopneustes; see Chace & Bruce, 1993] 

Tuleariocaris neglecta Chace, 1969: 266; Figs 10-11. [Bellairs Research Institute of McGill University, St. 

James, Barbados, on Diadema antillarum] 

Tuleariocaris sarec Berggren, 1994a: VIII.19; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west of 

the northern half of Inhaca Island, approx. 26°S 33°E, Moçambique] 

Tuleariocaris sarec Berggren, 1994b: 794; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west of the 

northern half of Inhaca Island, approx. 26°S 33°E, Moçambique] 

= Tuleariocaris sarec Berggren, 1994b: 794; Figs 10-14. [coral reef of Ilha des Portugueses, 500 m west 

of the northern half of Inhaca Island, approx. 26°S 33°E, Moçambique] 

Tuleariocaris zanzibarica Bruce, 1967b: 33; Figs 13-18. [Mtoni, Zanzibar, from an echinoid collected at 

L.W.S.] 

Typton o.G. Costa, 1844a 

= Typton O.G. Costa, 1844a (type species Typton spongicola O.G. Costa, 1844a, by monotypy, gender 

masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957)


= Pontonella Heller, 1856 (type species Pontonella glabra Heller, 1856 (junior subjective synonym of 

Typton spongicola O.G. Costa, 1844a), by monotypy, gender feminine) 

Typton ascensionis Manning & Chace, 1990: 10; Fig. 4. [McArthur Point, Ascension Island] 

Typton australis Bruce, 1973e: 254; Figs 1-4. [Chinaman’s Reef, between Opal and Rudder Reefs, Great 

Barrier Reef, Queensland, Australia, 10ft] 

Typton bawii Bruce, 1972e: 243; Figs 1-5. [South of Bawi Island, off west coast of Zanzibar, 6°9.7'S 39°8.3'E, 

18-25 m] 

Typton capricorniae Bruce, 2000: 94; Figs 6-8. [Heron Island, Capricorn Islands, Queensland] 

Typton carneus Holthuis, 1951b: 162; Plate 51. [Tortugas, Florida] 

Typton crosslandi Bruce, 1978d: 294; Figs 1-3. [Off Onslow Island, near Charles (= Floreana, or Santa Maria 

Island), Galapagos Archipelago, 4 fms] 

Typton distinctus Chace, 1972: 49; Figs 13-14. [Los Arroyos, Provincia de Pinar del Rio, in sponge] 

Typton gnathophylloides Holthuis, 1951b: 159; Plate 50. [Tortugas, about 13 miles S of No. 2 Red Buoy, up 

to 45 fms] 

Typton hephaestus Holthuis, 1951b: 159; Plate 49, figs o-p. [southern part of the Gulf of California, 24°12'N 

109°55'W, 9.5 fms] 

Typton holthuisi De Grave, 2010: 210; Figs 1-2. (Ascension Island, Comfortless Cove, 07°54.615'S 

014°24.142'W, 2 m) 

Typton manningi Bruce, 2000: 87; Figs 1-5. [Abbot Point, Queensland, Australia, 19°53'S 148°05'E] 

Typton prionurus Holthuis, 1951b: 165; Plate 52. [Tortugas, Florida, in channel between Middle Ground 

and White Shoal, 10 fms] 

Typton serratus Holthuis, 1951b: 167; Plate 53. [Galapagos Islands, Ecuador, Albemarle Island, Tagus 

Cove, shallow water] 

Typton spongicola O.G. Costa, 1844a: 289. [Bay of Naples, Italy; neotype designated by Bruce, 2009c] 

= Pontonia pulsatrix Nardo, 1847: 6. [le lagune e golfo Veneto] 

= Pontonella glabra Heller, 1856: 634; Plate 9. [Meeresküste von Zara] 

= Typton spongiosus Spence Bate, 1868b: 119; Plate 11, Figs 1-1z. [Type locality not indicated] 

? = Pontonia Vagans Gourret, 1887b: 1132. [nomen nudum] 

? = Pontonia Vagans Gourret, 1888: 39. [Fonds vaseux entre l’île de Tiboulen et la calanque de Méjean, 

par 35 brasses] 

Typton tortugae McClendon, 1911 

= Typton tortugæ McClendon, 1911: 57; Plate 1, fig. 2. [Tortugas, Florida] 

Typton vulcanus Holthuis, 1951b: 157; Plate 49, Figs a-n. [south of Tortugas, Florida] 

Typton wasini Bruce, 1977d: 272; Figs 1-6. [Wasini Island Channel, Kenya, 4°39.4'S 39°22.2'E, 6 fm] 

= Typton nanus Bruce, 1987d: 49; Figs 1-5. [Australian North-West Shelf, 16°34'S 121°27'E, 40-46 m] 

Typtonoides Bruce, 2010j 

= Typtonoides Bruce, 2010j (type species Typtonoides neili Bruce, 2010j, by original designation and 


Typtonoides neili Bruce, 2010j: 69; Figs 1-6. [19°53'S 158°19'E, Long Island, Chesterfield Islands, New 

Caledonia, 15 m] 

Typtonychus Bruce, 1996 

= Typtonychus Bruce, 1995 (unavailable name; see Holthuis, 1996) 

= Typtonychus Bruce, 1996 (type species Typtonychus crassimanus Bruce, 1996, by original designation, 


Typtonychus anomalus (Bruce, 1979e) 

= Onycocaris anomala Bruce, 1979e: 69; Figs, 1-4. [between North and South Shell Islands, Port Darwin, 

Northern Australia, 3-7 fms] 

Typtonychus crassimanus Bruce, 1996: 253; Figs 22-27. [New Caledonia, Anaa Reef, 21°22.70'S 166°01.50'E, 38 m] 

Typtonychus dentatus (Fujino & Miyake, 1969c) 

= Typton dentatus Fujino & Miyake, 1969c: 80; Figs 1-2. [Ukachi, Yoron-jima Island, Ryukyu Islands, 

Japan]


Typtonychus dimorphus (Bruce, 1986f) 

= Typton dimorphus Bruce, 1986f: 278; Figs 1-4. [Ashmore Reef, Timor Sea, 12°15.0'S 123°00.0'E, outer 

slope of NW reef, 5 m] 

Unguicaris Marin & Chan, 2006 

= Unguicaris Marin & Chan, 2006 (type species Unguicaris panglaonis Marin & Chan, 2006, by original 


Unguicaris alegrias (Bruce, 1986d) 

= Periclimenes alegrias Bruce, 1986d: 143; Figs 1A, 2-5, 15A-C. [Coral Bay, Port Essington, 11°11.2'S 

132°02.8'E, 2-4 m] 

Unguicaris novaecaledoniae (Bruce, 1968a) 

= Periclimenes novaecaledoniae Bruce, 1968a: 1157; Figs 6-9. [Ilot Maître, Noumea, New Caledonia, 

22°20'S 116°25'E] 

Unguicaris panglaonis Marin & Chan, 2006: 530; Figs 6-13. [Philippines, 9°29.4'N 123°56.0'E, 6-37 m] 

Unguicaris pilipes (Bruce & Zmarzly, 1983) 

= Periclimenes pilipes Bruce & Zmarzly, 1983: 644; Figs 1-5. [southern tip of Medren Islet, Enewetak 

Atoll, Marshall Islands, 11°24'N 162°22'E, 3 m] 

Urocaris Stimpson, 1860a 

= Urocaris Stimpson, 1860a (type species Urocaris longicaudata Stimpson, 1860a, by original selection, 


Urocaris longicaudata Stimpson, 1860a: 39. [in littoribus Carolinensibus] 

Veleronia Holthuis, 1951b 

= Veleronia Holthuis, 1951b (type species Veleronia serratifrons Holthuis, 1951b, by original selection, 


Veleronia serratifrons Holthuis, 1951b: 196; Plate 62; Plate 63, figs a-e. [Ecuador, La Libertad, 4 fms] 

Veleronia sympathes (De Ridder & Holthuis, 1979) 

= Pontonides sympathes De Ridder & Holthuis, 1979: 101; Figs 1-3. [Punta Pitt, north-east coast of San 

Cristobal Island, Galapagos Archipelago, Ecuador, 8 m, on Antipathes galapagensis] 

Veleroniopsis Gore, 1981 

= Veleroniopsis Gore, 1981 (type species Veleroniopsis kimallynae Gore, 1981, by original designation 


Veleroniopsis kimallynae Gore, 1981: 147; Fig. 2. [Elbow Reef, off Key Largo, Monroe County, Florida, 

25°07.70'N 80°15.90'W, 18.3 m] 

Vir Holthuis, 1952c 

= Vir Holthuis, 1952c (type species Palaemonella orientalis Dana, 1852a, by original designation and 


Vir colemani Bruce, 2003c: 119; Figs 1-6. [Loloata Island, Papua New Guinea, 15 m] 

Vir euphyllius Marin & Anker, 2005: 118; Figs 1-5. [Vietnam, Nha Trang Bay, Tre Island, southern bay, 

near lighthouse, 15 m, on Euphyllia cf. divisa] 

= Vir pareuphyllius Marin & Anker, 2005: 123; Figs 6-8. [Vietnam, Nha Trang Bay, Tre Island, northern 

”Bay Tre”, 7 m, on Euphyllia cf. parancora] 

Vir longidactylus Marin, 2008a: 383; Figs 6-8, 9c. [South China Sea, Vietnam, Nhatrang Bay, Tre Island, Tre 

Bay, on Physogyra lichtensteini, 5 m] 

Vir orientalis (Dana, 1852a) 

= Palaemonella orientalis Dana, 1852a: 26. [in mari Suluensi] 

Vir philippinensis Bruce & Svoboda, 1984: 87; Figs 1-4. [island of Cebu, associated with Plerogyra sinuosa] 

Vir smiti Fransen & Holthuis, 2007: 101; Figs 1-27, 32-34. [Philippines, Cebu Strait, W of Bohol, N side of 

Cabilao Island, NE of Looc, 9°53.59'N 123°46.92'E, 15 m, on Physogyra lichtensteini]


Waldola Holthuis, 1951b 

= Waldola Holthuis, 1951b (type species Waldola schmitti Holthuis, 1951b, by original designation 


Waldola schmitti Holthuis, 1951b: 186; Plate 58; Plate 59, Figs a-f. [Tepic, Mexico, off Isabel Island, 10-25 

fms] 

Yemenicaris Bruce, 1997a 

= Yemenicaris Bruce, 1997a (type species Yemenicaris trullicauda Bruce, 1997a, by original designation 


Yemenicaris trullicauda Bruce, 1997a: 1215; Figs 1-4. [Khalf, Yemen, 14°35'N 49°10'E, to 9 m] 

Zenopontonia Bruce, 1975e 

= Zenopontonia Bruce, 1975e (type species Periclimenes (Periclimenes) noverca Kemp, 1922, by original 


Zenopontonia noverca (Kemp, 1922) 

= Periclimenes (Periclimenes) noverca Kemp, 1922: 162; Figs 28-30. [New Caledonia] 

Family TyPHloCARIDIDAe Annandale & Kemp, 1913 

Typhlocaris Calman, 1909 

= Typhlocaris Calman, 1909 (type species Typhlocaris galilea Calman, 1909, by monotypy, gender feminine; 


Typhlocaris ayyaloni Tsurnamal, 2008: 490; Figs 1-4. [Ayyalon Cave, Israel] 

Typhlocaris galilea Calman, 1909: 94; Plate 19. [small pond near the town of Tiberias] 

Typhlocaris lethaea Parisi, 1921: 241; Figs 1-5. [Grotta del Lete (Giok-Kebir), 10 km d Est di Bengasi, Cirenaica] 

Fig. 42. Typhlocaris salentina Caroli, 1923. Photo by Ninni Ciccarese.


Typhlocaris salentina Caroli, 1923: 265. [grotta presso Castro, in Terra d’Ortanto; full description in Caroli, 

1924; type locality corrected therein to grotte la Zinzulusa, costa della Penisola Salentina copreso 

tra il Capo d’Otranto e il Capo di Leuca] (Fig. 42) 

Superfamily AlPHeoIDeA Rafinesque, 1815 

Family AlPHeIDAe Rafinesque, 1815 

Acanthanas Anker, Poddoubtchenko & Jeng, 2006 

= Acanthanas Anker, Poddoubtchenko & Jeng, 2006 (type species Acanthanas pusillus Anker, Poddoubtchenko 

& Jeng, 2006, by original designation and monotypy, gender masculine) 

Acanthanas pusillus Anker, Poddoubtchenko & Jeng, 2006: 342; Figs 1-5. [north of Doljo, Panglao Island, 

Philippines, 9°35.9'N 123°44.5'E, 24 m] 

Alpheopsis Coutière, 1897b 

= Alpheopsis Coutière, 1897b (type species Betaeus trispinosus Stimpson, 1860a, designated by 

Holthuis, 1955b, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Alpheopsis aequalis Coutière, 1897b 

= Alpheopsis æqualis Coutière, 1897b: 382. [mer Rouge; océan Indien] 

= Alpheopsis equalis var. truncatus Coutière, 1903: 89; Figs 37-38. [Goifufahendu Atoll] 

= Alpheopsis equalis var. truncatus Coutière, 1905b: 869; Fig. 139. [Goifufahendu Atoll] 

= Alpheopsis consobrinus De Man, 1910a: 305. [8°25.2'S 127°18.4'E, anchorage between Nusa Besi and 

the NE-point of Timor, 27-54 m] 

Alpheopsis africana Holthuis, 1952a 

= Alpheopsis africanus Holthuis, 1952a: 45; Fig. 11 (as A. africana). [8°30'S 13°E, 20 m W. Pointa do 

Dandé, 150 m] 

Alpheopsis allanhancocki Wicksten, 1992: 51; Figs 1-2. [Isla Candelero, Sonora, Gulf of California, Mexico, 

27°56'N 111°03'W, 10 m] 

Alpheopsis azorica Anker, d’Udekem d’Acoz & Poddoubtchenko, 2005: 98; Figs 1-3. [Monte da Guia near 

Horta, Faial Island, Azores, in large marine cave, 35 m] 

Alpheopsis biunguiculata Banner, 1953 

= Alpheopsis biunguiculatus Banner, 1953: 18; Fig. 5. [Halape, Kau Coast, Hawaii, from Pocillopora 

meandrina] 

Alpheopsis chalciope De Man, 1910a 

= Alpheopsis Chalciope De Man, 1910a: 306. [0°7.2'N 130°25.5'E, 83-59 m] 

Alpheopsis chilensis Coutière, 1897b: 382. [Chili] 

Alpheopsis cortesiana Wicksten & Hendrickx, 1986: 196, Fig. 1. [Off Río Fuerte, Sinaloa, Mexico, 25°48'N 

109°34'W, 90 m, mud] 

Alpheopsis diabolus Banner, 1956: 325 (as A. diabilus); Fig. 3. (as A. diabolus). [Saipan, Mariana Islands; 

name corrected to A. diabolus by Banner & Banner, 1964 under First Reviser Principle] 

Alpheopsis equidactylus (Lockington, 1877a) 

= Alpheus equidactylus Lockington, 1877a: 35. [Monterey] 

Alpheopsis garricki Yaldwyn, 1971: 87. [off Rangitoto Island, Cook Strait, 40°44'S 174°0'E, 40-50 fms] 

Alpheopsis harperi Wicksten, 1984a: 97; Fig. 1. [off Freeport, Texas, 28°43'41''N 95°14'10''W, 21 m] 

Alpheopsis idiocarpus Coutière, 1908: 194. [Providence, 50-78 brasses] 

Alpheopsis labis Chace, 1972: 55; Fig. 15. [Antigua, Charlotte Point, English Harbour] 

Alpheopsis shearmii (Alcock & Anderson, 1899) 

= Alpheus Shearmii Alcock & Anderson, 1899: 283. [Investigator stn 232 (Laccadive Sea, 7°17'30''N 

76°54'30''E), 430 fms] 

Alpheopsis tetrarthri Banner, 1956: 328; Fig. 4. [north side of passage into Saipan harbor, 40-45ft] 

Alpheopsis trigona (Rathbun, 1901) 

= Jousseaumea trigona Rathbun, 1901: 111; Fig. 21. [off Vieques, 6 fms]


Fig. 43. Alpheopsis yaldwyni Banner & Banner, 1973. Photo by Arthur Anker. 

Alpheopsis trispinosa (Stimpson, 1860a) 

= Betaeus trispinosus Stimpson, 1860a: 32. [Green Cape, New South Wales, 30 fms; neotype selection 

by Banner & Banner, 1973a] 

Alpheopsis undicola Banner & Banner, 1973a: 340; Fig. 15. [Opal Reef, outermost margin of Great Barrier 

Reef, 5ft] 

Alpheopsis vietnamensis Tiwari, 1964b: 314. [Anchorage of Itu Aba; fully described in Tiwari, 1965, as 

Alpheopsis vietnami] 

Alpheopsis yaldwyni Banner & Banner, 1973a: 344; Fig. 17. [One Tree Island, Capricorn Group] (Fig. 43) 

Alpheus Fabricius, 1798 

= Crangon Weber, 1795 (type species Astacus Malabaricus Fabricius, 1775, by monotypy, gender feminine; 

name suppressed under the plenary power of the ICZN for both the Principle of Priority and 

that of Hononymy, and placed on the Official Index of Rejected and Invalid Names in Zoology in 


= Alpheus Fabricius, 1798 (type species Alpheus avarus Fabricius, 1798, designated by Latreille, 1810, 

gender masculine; junior homonym of Alpheus Weber, 1795 (Crustacea Brachyura); name conserved 

by the suppression for the Principle of Priority and that of Homonymy under the plenary powers 

of the ICZN of the generic name Alpheus Weber, 1795; name placed on the Official List of Generic 


= Cryptophthalmus Rafinesque, 1814 (type species Cryptophtalmus ruber Rafinesque, 1814 (junior subjective 

synonym of Cancer glaber Olivi, 1792), by monotypy, gender masculine) 

= Autonomæa Risso, 1816 (type species A.[utonomæa] Olivii Risso, 1816 (junior subjective synonym of 

Cancer glaber Olivi, 1792), by monotypy, gender feminine) 

= Asphalius Roux, 1831 (type species Palæmon brevirostris Olivier, 1811, by monotypy, gender masculine) 

= Dienècia Westwood, 1835a (type species Hippólyte rùbra Hailstone, 1835a (junior objective synonym 

of Hippólyte macrochèles Hailstone, 1835b), by monotypy, gender feminine) 

= Nauplius Risso, 1844 (type species N.[ika] Variegata Risso, 1816 (invalid senior subjective synonym 

of Alpheus dentipes Guérin, 1832), designated by Holthuis, 1977, gender masculine) 

= Phleusa Nardo, 1847 (type species Phleusa cynea Nardo, 1847 (junior subjective synonym of Cancer 

glaber Olivi, 1792), gender feminine)


= Halopsyché de Saussure, 1857b (type species Halopsyché lutaria de Saussure, 1857b, a junior subjective 

synonym of A.[lpheus] heterochælis Say, 1818, by monotypy, gender feminine) 

= Alpheoides Paul’son, 1875 (type species A.[lpheus] insignis Heller, 1861, designated by Holthuis, 

1955b, gender masculine) 

= Paralpheus Spence Bate, 1888 (type species Palæmon diversimanus Olivier, 1811, by monotypy, gender 


= Thunor Armstrong, 1949 (type species Crangon rathbunae Schmitt, 1924b (junior subjective synonym 

of Alpheus Simus Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856), by monotypy, gender 


Alpheus acutocarinatus De Man, 1909a: 104. [7°25'S 113°16'E, Madura Strait, 56 m; 8°44'.5S 116°2'.5E, Bay 

of Labuan Tring, west coast of Lombok, 18-27 m; 0°58'.5N 122°42'.5E, west of Kwandang-bay-entrance 

72 m; Sapeh-bay, east coast of Sumbawa, up to 36 m] 

Alpheus acutofemoratus Dana, 1852a 

= Alpheus acuto-femoratus Dana, 1852a: 22. [in freto ”Balabac”] 

= Alpheus parabrevipes Coutière, 1898a: 151; Figs 2-2c. [Samoa] 

Alpheus adamastor Coutière, 1908 

= Alpheus Adamastor Coutière, 1908: 209. [Salomon, Chagos] 

Alpheus aequus W. Kim & Abele, 1988: 55; Fig. 23. [Playa Blancas, Costa Rica] 

Alpheus agilis Anker, Hurt & Knowlton, 2009: 11; Figs 4, 5F. [São Tomé, 200 m west of Lagoa Azul, rocky 

intertidal] 

Alpheus agrogon Ramos, 1997: 395; Figs 1-3. [La Ventana, Gorgona Island, 2°56'10''N 78°12'05''W] 

Alpheus albatrossae (Banner, 1953) 

= Crangon albatrossae Banner, 1953: 60; Fig. 18. [Penguin Bank, west of Molokai, 27 fm] 

Alpheus alcyone De Man, 1902: 870; Plate 27, fig. 61. [Ternate] 

= Alpheus aculeipes Coutière, 1905b: 892; Plate 79, figs 31-31g. [South Nilandu Atoll; Hulule Male 

Atoll; North Male Atoll; Milandumadulu Atoll, Naifaro Reef; Djibouti; Fernando-Veloso] 

Alpheus alpheopsides Coutière, 1905b: 901; Plate 83; figs 40-40d. [Hulule Male Atoll] 

Alpheus amblyonyx Chace, 1972: 59; Fig. 16. [Dominica, north end of Woodbridge Bay] 

Alpheus amirantei Coutière, 1908 

= Alpheus Amirantei Coutière, 1908: 205. [Amirante, 25-80 brasses] 

Alpheus anchistus De Man, 1920a: 108. [off Ambon, 54 m] 

Alpheus angulosus McClure, 2002: 368. [nomen novum for Alpheus angulatus McClure, 1995; nec Alpheus 

strenuus var. angulatus Coutière, 1905b] 

= Alpheus angulatus McClure, 1995: 85; Figs 1-2; nec Alpheus strenuus var. angulatus Coutière, 1905b. 

[South Padre Island, Texas, Laguna Madre just north of Brazos-Santiago Pass] 

Alpheus angustilineatus Nomura & Anker, 2005: 111; Figs 3-4, 13C-D. [Fuki, Kuro-shima, Yaeyama Islands, 

southern Ryukyu Archipelago, Japan, 3 m] 

Alpheus antepaenultimus W. Kim & Abele, 1988: 103; Fig. 44. [Amador, Panama] 

Alpheus architectus De Man, 1897: 726; Plate 34, figs 60-60f. [Atjeh] 

? = Alpheus bullatus Barnard, 1955: 45; fig. 22. [Delagoa Bay; St. Lucia Bay] 

Alpheus arenensis (Chace, 1937b) 

= Crangon arenensis Chace, 1937b: 119; Fig. 4. [off Arena Bank, 2.5 fms, in coral] 

Alpheus arenicolus Banner & Banner, 1983: 14; Fig. 1. [Madagascar] 

Alpheus arethusa De Man, 1909a 

= Alpheus Arethusa De Man, 1909a: 100. [Haingsisi, Samau island, Timor, 36 m] 

Alpheus armatus Rathbun, 1901: 108; Fig. 20. [Ponce] 

Alpheus armillatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= A.[lpheus] armillatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 354. [les Antilles] 

= Alpheus lancirostris Rankin, 1900: 541; Plate 17, fig. 5. [Bailey’s Bay, at low tide] 

= Crangon verrilli Schmitt, 1924b: 77; Plate 2, figs 7-10. [Barbados] 

Alpheus arnoa Banner, 1957: 199; Fig. 5. [lagoon reef, Arno Atoll, Marshall Islands, 4-60 ft] 

Alpheus astrinx Banner & Banner, 1982a: 35; Fig. 5. [Garden Island, near Perth, Western Australia, 100 

yards offshore]


Alpheus australiensis Banner & Banner, 1982a: 256; Fig. 79. [Caloundra, Queensland] 

Alpheus australosulcatus Banner & Banner, 1982a: 83; Fig. 21. [North Head, Flinders, Victoria] 

Alpheus baccheti Anker, 2010a: 50; Figs 1-5. [French Polynesia, Tuamotu Islands, Makemo, lagoon, reef 

flat, around 0.5 m at low tide] 

Alpheus bahamensis Rankin, 1898 

= Alpheus hippothoë var. bahamensis Rankin, 1898: 247; Plate 30, fig. 5. [New Providence and Nassau, 

N. P.] 

Alpheus balaenodigitus Banner & Banner, 1982a: 223; Fig. 70. [Port Walcott, Western Australia, 20°39'S 

117°10'E, 8 fms] 

Alpheus bannerorum Bruce, 1987e: 61; Figs 1-6; Plate 1. [Dudley Point, Darwin, Northern Territory] 

Alpheus barbadensis (Schmitt, 1924b) 

= Crangon barbadensis Schmitt, 1924b: 70; Plate 2, figs 1-3. [Off the Castel, E side Barbados, 1-4 fms] 

Alpheus barbatus Coutière, 1897c: 235. [Djibouti] (Fig. 44) 

Alpheus batesi Banner & Banner, 1964: 94; nomen novum for Alpheus leviusculus sensu Spence Bate, 1888; 

nec Dana, 1852b. [Challenger stn 203, 11°6'N 123°9'E, off Panay, Philippine Islands, 20 fms] 

Alpheus belli Coutière, 1898a 

= Alpheus Belli Coutière, 1898a: 149; Figs 1-1a. [Fernando-Noronha] 

Alpheus bellimanus Lockington, 1877a: 34. [San Diego, among kelp] 

Alpheus bellulus Miya & Miyake, 1969: 308; Figs 1-2. [Ezura, Tanabe Bay, Wakayama Prefecture, burrowing 

on gravely sand, 5 m] 

Alpheus bicostatus De Man, 1908d: 102. [Sialus Ketjil, Paternoster-islands, 27 m and less; 6°7'.5N 120°26'E, 

anchorage of Nort-Ubian, 16-23 m; Buton-strait, between floating seaweed] 

Alpheus bidens (Olivier, 1811) 

= Palæmon bidens Olivier, 1811: 663. [côtes de la Nouvelle-Hollande] 

= Alpheus tridentatus Zehntner, 1894: 204; Plate 8, fig. 24. [Amboine] 

= Alpheus dissodontonotus Stebbing, 1915: 83; Plate 22. [33°50'S 25°46'E, 20 fms] 

Alpheus bisincisus De Haan, 1849 [in De Haan, 1833-1850] 

= Alpheus bis-incisus De Haan, 1849 [in De Haan, 1833-1850]: Plate 45, fig. 3. (1844); 179 (1849, as 

Alpheus avarus). [Japan] 

Fig. 44. Alpheus barbatus Coutière, 1897. Photo by Arthur Anker.


= Alpheus bis-incisus var. Malensis Coutière, 1905b: 910; Plate 86, figs 48-48d. [South Male Atoll; Suvadiva 

Atoll; South Nilandu Atoll; Mulaku Atoll] 

= Alpheus bis-incisus var. stylirostris Coutière, 1905b: 911; Plate 86, figs 49-49a. [Mulaku Atoll] 

? = Alpheus bisincisus var. variabilis De Man, 1909a: 109. [Makassar, up to 32 m] 

Alpheus blachei Crosnier & Forest, 1965a: 358; Fig. 3. [lieudit Pointe Indienne, situé dans les environs de 

Pointe-Noire, Congo] 

Alpheus bouvieri A. Milne-Edwards, 1878 

= Alpheus Bouvieri A. Milne-Edwards, 1878: 231. [toutes les îles du Cap-Vert] 

Alpheus brachymerus (Banner, 1953) 

= Crangon brachymerus Banner, 1953: 58; Fig. 17. [between Maui and Lanai Islands, 21-28 fms] 

Alpheus bradypus Coutière, 1905b: 891; Plate 78, figs 30-30a’; Plate 79, figs 30b-30e. [Minikoi] 

Alpheus brevicristatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 1. (1844); 177 (1849, as Alpheus 

malabaricus Fabricius) [Japan; lectotype designation by Yamaguchi & Baba, 1993] 

= Alpheus kingsleyi Miers, 1879: 54. [35°7'N 136°55'E, 3 fms] 

Alpheus brevipes Stimpson, 1860a: 30. [ad insulas Hawaiensis, inter ramos madreporarum] 

Alpheus brevirostris (Olivier, 1811) 

= Palæmon brevirostris Olivier, 1811: 664. [côtes de la Nouvelle-Hollande] 

= Alpheus brevirostris var. angustodigitus De Man, 1911d: 385. [Balikpapan, east coast of Borneo] 

= A.[lpheus] dispar Randall, 1840: 141. [Manilla] 

Alpheus brucei Banner & Banner, 1982b: 359; Fig. 1. [Heron Island reef flat, 0.5 m from Porites andrewsi] 

Alpheus bucephaloides Nobili, 1905e: 238. [Golfe Persique, Banc au N.E. d’île Arzana et pêche côtières 

dans les polypiers; lectotype selection by Banner & Banner, 1981a] 

Alpheus bucephalus Coutière, 1905b: 890; Plate 78, figs 29-29f. [Hulule Male Atoll; Minikoi; Cap; Fernando-Veloso; 

Mahé; Djibouti] 

= Alpheus consobrinus De Man, 1908d: 101 [Lumu-Lum-shoal, Borneo Bank, reef; South-east side of 

Pearl-bank, Sulu-archipelago, 15 m; East side of Pajunga-island, Kwandang-bay, reef exploration; 

Anchorage off Sawan, Siau-island, 27 m; Saleyer-anchorage and surroundings, up to 36 m; 10°52'.4S 

123°1'.1E, Boeka- or Cyrus-bay, South coast of Rotti-island, up to 36 m; Anchorage east of Sailus 

Besar, Paternoster-islands, up to 36 m] 

Alpheus buchanorum Banner & Banner, 1983: 19; Fig. 2. [Round Island, off Victoria, Mahé, Seychelles] 

Alpheus bunburius Banner & Banner, 1982a: 213; Fig. 66. [Bunbury, Western Australia] 

Alpheus californiensis Holmes, 1900: 186; Plate 2, fig. 42; Plate 3, figs 43-44. [San Pedro, California, dredged 

in the harbour] 

Alpheus canaliculatus Banner & Banner, 1968: 141; Fig. 1. [20°05'N 115°11'E, 137 fms] 

Alpheus candei Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856] 

= Alpheus Candei Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, figs 9-9a. 

[costas de la Isla de Cuba] 

Alpheus chacei Carvacho, 1979: 455; Figs 4-6. [entre les canaux Belle Plaine et Perrin, prés de la mangrove] 

= Alpheus maxilliplanus Christoffersen, 1979: 318; Figs 11-13. [Balneário de Atalaia, Sergipe, Brazil] 

Alpheus chamorro Banner, 1956: 349; Fig. 14. [Saipan] 

Alpheus chilensis Lenz, 1902 

= Alpheus bouvieri var. chilensis Lenz, 1902: 732. [Calbuco] 

Alpheus chiragricus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= A.[lpheus] chiragricus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 354. [les mers 

d’Asie] 

Alpheus christofferseni Anker, Hurt & Knowlton, 2007a: 3; Figs 1-6, 11a-c, 12b-c. [Brazil, Atol das Rocas, 

Ilha do Cemitério] 

Alpheus clamator Lockington, 1877b: 43. [Santa Barbara Island] 

= Alpheus transverso-dactylus Kingsley, 1878b: 196. [Santa Barbara and San Diego, California; Bermudas] 

? = Alpheus barbara Lockington, 1878b: 471. [Santa Barbara, California] 

Alpheus clypeatus Coutière, 1905b: 897; Plate 81, figs 36-36’; Plate 82, figs 36a-36g. [Minikoi; Hulule Male 

Atoll]


Alpheus coetivensis Coutière, 1908 

= Alpheus Coetivensis Coutière, 1908: 210. [Coetivy; Salomon] 

= Crangon edmondsoni Banner, 1953: 78; Fig. 26. [Kawela Bay, Oahu] 

Alpheus collumianus Stimpson, 1860a: 30. [ad insulas ”Bonin”, inter corallia vientia ad prof. 1 org] 

= Alpheus Seurati Coutière, 1905b: 881; Plate 75, figs 20-20e. [l’île Marutea, archipel Gambier] 

= Alpheus Malhaensis Coutière, 1908: 205. [Saya de Malha, 29 brasses] 

= Alpheus collumianus probabilis Banner, 1956: 338; Fig. 10. [about 1.4 miles offshore, north of Puntan 

Muchot, Saipan, 8-9ft] 

= Alpheus collumianus medius Banner, 1956: 340; Fig. 11. [Saipan] 

= Alpheus collumianus inermis Banner, 1956: 342; Fig. 12. [about 1 mile off Muchot Point, near entrance 

to Saipan harbor, 22ft] 

Alpheus colombiensis Wicksten, 1988: 1; Fig. 1. [Bahía Malaga, Pacific coast of Colombia, 4°0'N 77°15'W, 

under rocks] 

= Alpheus hamus W. Kim & Abele, 1988: 91; Fig. 38. [Golfito, Costa Rica] 

Alpheus compressus Banner & Banner, 1981b: 227; Fig. 3. [13°59.8'N 120°18.6'E, 188-192 m] 

Alpheus confusus Carvacho, 1989: 59; Fig. 1. [Isla San José] 

Alpheus coutierei De Man, 1909a 

= Alpheus Coutierei De Man, 1909a: 107. [South-east side of Pearl-bank, Sulu-archipelago, 15 m; 

8°25'.2S 127°18'.4E, Anchorage between Nusa-Besi and the N.E.-point of Timor, 27-54 m] 

Alpheus crinitus Dana, 1852a: 21. [in freto ”Balabac”] 

Alpheus cristatus Coutière, 1897d: 303. [Thursday-Island] 

Alpheus cristulifrons Rathbun, 1900: 152. [Brazil, Fernando de Noronha; lectotype selection by Anker, 

Hurt & Knowlton, 2008a] 

Alpheus crockeri (Armstrong, 1941) 

= Crangon crockeri Armstrong, 1941: 8; Figs 2-3. [eastern reef, Mataatu Harbor, Savaii, shallow water] 

= Crangon tuthilli Banner, 1953: 63; Fig. 19. [off southwest Oahu, 40-350 ft] 

Alpheus cyanoteles Yeo & Ng, 1996: 46; Figs 4-7. [Sungei Tementang, Kota Tinggi, Johore, 1°52'0.81''N 

103°55'49.2''E] 

Alpheus cylindricus Kingsley, 1878b: 196. [Pearl Island, Bay of Panama; neotype designation by Anker, 

Hurt, Jara & Knowlton, 2008 invalid, as holotype extant] 

Alpheus cythereus Banner & Banner, 1966a: 123; Fig. 45. [Patong Beach, Phuket, Thailand, 2 m] 

Alpheus dasycheles Coutière, 1908: 211. [Seychelles, 37 brasses] 

Alpheus davaoensis Chace, 1988: 21; Fig. 2. [Philippines, Davao Gulf, Mindanao, 7°04'48''N 125°39'38''E, 

51 m] 

Alpheus dentipes Guérin, 1832: 39; Plate 27, fig. 3. [Sapience; cap Ténare; golfe de Gènes] 

= N.[ika] Variegata Risso, 1816: 86. [le golfe de Beaulieu, dans les algues profondes; name placed on 

the Official Index of Rejected and Invalid Names in Zoology in Opinion 606 in 1961] 

= Alpheus streptochirus Stimpson, 1860a: 30. [ad insulas ”Cape de Verdes”, inter nulliporas ad prof. 

20 org] 

= Crypt.[ophthalmus] Costa O.G. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 2; Plate 7, fig. 2. 

[vive nel Mediterraneo egualmente che nello Ionio] 

= Alpheus similis Czerniavsky, 1884: 30. [Portu Suchum, circa 2 m] 

= Alpheus cristidigitus Spence Bate, 1888: 546; Plate 97, fig. 3. [Challenger stn 93C, 16°57'15''N 25°1'W, 

St. Vincent, Cape Verde Islands, 52 fms] 

= Alpheus Gabrieli Gourret, 1887a: 1033. [pointe rouge de Montredon] 

Alpheus deuteropus Hilgendorf, 1879: 834; Plate 4, figs 8-10. [Zanzibar] 

Alpheus diadema Dana, 1852a: 23. [ad insulam ”Maui” Hawaiensem] 

= A.[lpheus] insignis Heller, 1861: 24. [im rothen Meere; fully described in Heller, 1862c] 

Alpheus digitalis De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 4 (1844); 178 (1849). [Japan] 

= Alpheus distinguendus De Man, 1909b: 155; Plate 7, Figs 9-14. [Japan] 

Alpheus distinctus W. Kim & Abele, 1988: 95; Fig. 40. [Punta Paitilla, Panama] 

Alpheus djeddensis Coutière, 1897e: 202. [Djeddah] 

Alpheus djiboutensis De Man, 1909b: 160; Plate 8, Figs 17-24. [Djibouti]


Alpheus dolerus Banner, 1956: 362; Fig. 21. [Saipan] 

Alpheus edamensis De Man, 1888b 

= Alpheus Hippothoë var. edamensis De Man, 1888b: 518. [Pulo Edam; Amboina] 

= Alpheus acanthomerus Ortmann, 1890: 474; Plate 36, fig. 12. [Tahiti] 

Alpheus edwardsii (Audouin, 1826) 

= Athanas Edwarsii Audouin, 1826: 91 (lapsus for Athanas Edwardsii, justified emendation in Audouin, 

1827; accompanying figures in Savigny, 1826: Plate 10, fig. 1). [Suez; neotype selection by 

Banner & Banner, 1973b] 

= Alpheus Audouini Coutière, 1905b: 911; Plate 87, Figs 52a-52a’’. [Miladumadulu Atoll, Goidu Atoll, 

Hulule Male Atoll, Reef Naifaro, Suvadiva Atoll, Minokoi] 

Alpheus ehlersii De Man, 1909c: 663; Plate 70. [Edam, Bay of Batavia] 

Alpheus estuariensis Christoffersen, 1984: 191; Figs 1-2. [Rio Potengi estuary, Natal, State of Rio Grande 

do Norte, Brazil] 

Alpheus euchirus Dana, 1852a: 21. [in freto ”Balabac”] 

Alpheus euphrosyne De Man, 1897: 745; Plate 36, figs 64-64d. [Java-See] 

= Alpheus eurydactylus De Man, 1920a: 109. [Java] 

Alpheus exilis W. Kim & Abele, 1988: 105; Fig. 45. [Sullivan Bay, James Island, Galapagos Islands] 

Alpheus explorator Boone, 1935: 139; Fig. 11; Plate 36. [Flores Strait, near Larantuka village, Flores Island, 

Dutch East Indies, 140 fms] 

Alpheus eulimene De Man, 1909a 

= Alpheus Eulimene De Man, 1909a: 101. [0°7'.2N 130°25'.5E, 83 m] 

Alpheus facetus De Man, 1908d: 100. [Anchorage off Pulu Jedan, east coast of Aru-islands (Pearl-banks), 

13 m] 

Alpheus fagei Crosnier & Forest, 1965b: 603; Fig. 1. [São Tomé, 0°20'N 6°47'E, 10 m, algues calcaires] 

Alpheus fasciatus Lockington, 1878b: 478. [Port Escondido, Gulf of California] 

Alpheus fasqueli Anker, 2001: 59; Figs 1c-d, 2b, 3f-j, 4e-g, 5i-n, 6l-n. [imported from Sri Lanka] 

Alpheus felgenhaueri W. Kim & Abele, 1988: 40; Fig. 16. [outer Miramar Bay on Cape Haro, north of Guaymas, 

Sonora, Mexico] 

Alpheus fenneri Bruce, 1994b: Figs 1-2, 5A. [Manado, Sulawesi, Indonesia, off Nusantara diving Centre, 6 m] 

Alpheus firmus W. Kim & Abele, 1988: 93; Fig. 39. [Miraflores Locks, Panama] 

Alpheus floridanus Kingsley, 1878b: 193. [Fort Jefferson, Florida] 

= Alpheus floridanus var. africana Balss, 1916: 21; Fig. 5. [Elfenbein-Küste, Wappu, 40 m] 

= Alpheus platycheirus Boone, 1927: 131; Figs 29-30. [Siguanea Bay, Isle of Pines, 12 fms] 

Alpheus foresti Banner & Banner, 1981b: 229; Fig. 4. [14°02.7'N 120°20.3'E, 191-200 m] 

Alpheus formosus Gibbes, 1850: 196. [USA, Florida, Big Pine Key, Bahia Honda State Park, shallow subtidal 

(less than 1.5 m), under rocks; neotype designation by Anker, Hurt & Knowlton, 2008b] 

= Alpheus Poeyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, fig. 10. [Cuba] 

Alpheus frontalis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= A.[lpheus] frontalis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 356. [les côtes de la 

Nouvelle-Hollande] 

= Alpheus latifrons A. Milne-Edwards, 1873: 87. [Upolu] 

= Betaeus utricola Richters, 1880: 164; Plate 17, Figs 34-35. [Insel Mauritius und Seychellen] 

Alpheus funafutensis Borradaile, 1899: 1013; Plate 65, fig. 10. [Funafuti] 

= Alpheus acanthomerus var. inermis Lanchester, 1901: 564. [Kelantan] 

Alpheus fujitai Nomura & Anker, 2005: 115; Figs 5-6, 13E-F. [Mizugama, Kadena, Okinawa-jima, Okinawa 

Islands, central Ryukyu Archipelago, Japan, submarine cave, 5-8 m] 

Alpheus fushima Nomura, 2009: 115; Figs 1-3. [Nishinohama, Kuro-shima Island, Yaeyama Islands, 

southern Ryukyu Archipelago, Japan, 5 m, internal canals of large colony of an unidentified 

sponge] 

Alpheus galapagensis Sivertsen, 1933 

= Alpheus strenuus var. galapagensis Sivertsen, 1933: 3; Plate 1, figs 1-5. [Floreana, Post Office Bay, 

Ranvik]


= Alpheus canalis W. Kim & Abele, 1988: 72; Fig. 30. [North Bay, San Francisco Island, Baja California] 

? = Alpheus cryptodentatus Christoffersen & Ramos, 1988: 61; Figs 1- 2. [Isla Playa Blanca, Ensenada 

de Utría, Chocó, Pacific coast of Colombia, 6°02'N 77°20'W] 

Alpheus georgei Banner & Banner, 1982a: 200; Fig. 61. [40 miles W of Cape Jaubert, 23 fms, from sponge] 

Alpheus glaber (Olivi, 1792) 

= Cancer glaber Olivi, 1792: 51; Plate 3, fig. 4. [la Golfo e delle Lagune di Venezia] 

= Cryptophthalmus ruber Rafinesque, 1814: 23. [Sicile] 

= A.[utonomæa] Olivii Risso, 1816: 166. [golfe de Nice] 

= Phleusa cynea Nardo, 1847: 6. [le lagune e golfo Veneto] 

Alpheus gracilipes Stimpson, 1860a: 31. [French Polynesia, Tahiti; neotype selection by Nomura & Anker, 

2005] 

Alpheus gracilis Heller, 1861 

= A.[lpheus] gracilis Heller, 1861: 27. [im rothen Meere] 

= Alpheus gracilis var. Alluaudi Coutière, 1905b: 882. [Mahé] 

= Alpheus gracilis var. luciparensis De Man, 1911d: 337. [5700 m N, 279° E from South Point of South- 

Lucipara-island, reef] 

= Crangon gracilis var. simplex Banner, 1953: 75; Fig. 25. [Waikiki Reef, Oahu] 

Alpheus grahami Abele, 1975b: 72; Fig. 29 A-C, E-J. [southeastern side of Malpelo Island, Colombia, 10 

m] 

Alpheus haanii Ortmann, 1890 

= Alpheus Haanii Ortmann, 1890: 472. [nomen novum for Alpheus minor De Haan, 1844 [in De Haan, 

1833-1850]] 

= Alpheus minor De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 6 (1844); 180 (1849). [Japan] 

Alpheus hailstonei Coutière, 1905b 

= Alpheus Hailstonei Coutière, 1905b: 879; Plate 74, figs 18-18c. [North Male Atoll; Mahlos Atoll; 

Hulule Male Atoll] 

= Alpheus Hailstonei var. laetabilis De Man, 1908d: 98. [8°23'.5S 119°4'6E, Sapeh-Strait, 70 m; 7°0'S 

120°34'5E, 400-120 m; 0°7'2N 130°25'5E, 83-59 m; Banda-Anchorage, 9-36 m; 8°25'2S 127°18'4E, 

anchorage between Nusa Besi and the N.E.-point of Timor, 27-54 m; 8°39'.1S 127°4'.4E, anchorage 

south of Timor, 34 m; Haingsis, Samau-island, up to 36 m; 8°30'S 119°7'.5E, Sapeh-Strait, 73 m] 

= Alpheus Hailstonei var. assimulans De Man, 1908d: 99. [5°36'.5S 132°55'.2E, 2.3 miles N, 63°W from 

the North Point of Nuhu Jaan, Kei-islands, 90 m; 8°25'2S 127°18'.4E, anchorage between Nusa Besi 

and the N.E.-point of Timor, 27-54 m; Mid-channel in Solor-strait off Kampong Menanga, 113 m] 

= Crangon hailstonei var. paucispinata Banner, 1953: 51; Fig. 6. [off Kauai, 68-90 fms] 

Alpheus halesii Kirk, 1887: 194; Plate 6d. [East Coast of Wellington Province] 

Alpheus hebes W. Kim & Abele, 1988: 62; Fig. 26. [James Island, Galapagos Islands] 

? = Alpheus bouvieri var. Bastardi Coutière, 1898b: 133 (partim); Fig. 1a. [specimen from Panama only] 

Alpheus heeia Banner & Banner, 1975a: 433; Fig. 5. [Kaneohe Bay, Oahu, Hawaii, 21°26'55''N 157°47'48''W] 

Alpheus heronicus Banner & Banner, 1982a: 220; Fig. 69. [Heron Island, Capricorn Group, Queensland, 

probably intertidal] 

Alpheus heterocarpus (Yu, 1935) 

= Crangon heterocarpus Yu, 1935: 63; Figs 3-4. [Peitaijo] 

Alpheus heterochaelis Say, 1818 

= A.[lpheus] heterochælis Say, 1818: 243. [Fort Saint George Inlet, 8 miles south of Amelia Island, Duval 

County, Florida, intertidal; neotype selection by McClure, 1995] 

= Halopsyché lutaria de Saussure, 1857b: 100. [les côtes de Cuba] 

= Alpheus andronyx Christoffersen, 1998: 358. [nomen nudum] 

Alpheus heurteli Coutière, 1897d 

= Alpheus Heurteli Coutière, 1897d: 305. [baie de Fernando-Velosa] 

Alpheus hippothoe De Man, 1888a 

= Alpheus Hippothoë De Man, 1888a: 268; Plate 17, Figs 1-5. [Sullivan Island and King Island Bay]


Alpheus holthuisi Ribeiro, 1964: 1; Figs 1-11. [St. Antão, Bahía de Porto-Novo, 12 m; São Vicente, Baía de 

Porto Grande, 3-5-11 m] 

Alpheus homochirus (Yu, 1935) 

= Grangon [sic] homochirus Yu, 1935: 61; Figs 1-2. [Weihaiwei] 

Alpheus hoonsooi W. Kim & Abele, 1988: 18; Fig. 7. [Academy Bay, Indefatigable Island, Galapagos Islands] 

Alpheus hoplocheles Coutière, 1897e: 197. [China, Amoy; lectotype designation by Fransen, Holthuis & 

Adema, 1997] 

Alpheus hortensis Wicksten & McClure, 2003: 26; Figs 1-3. [Stetson Bank, 28°10'N 94°17'W, 21-24 m] 

Alpheus hululensis Coutière, 1905b 

= Alpheus Bouvieri var. Hululensis Coutière, 1905b: 908; Plate 85, figs 46-46b. [Hulule Male Atoll] 

Alpheus hutchingsae Banner & Banner, 1982a: 191; Fig. 58. [Lizard Island, Queensland] 

Alpheus hyeyoungae W. Kim & Abele, 1988: 75; Fig. 31. [northeast beach of Isla Jaltimba, off Rincon de los 

Guayabitas, Nayarit, Mexico] 

Alpheus hyphalus Chace, 1988: 31; Figs 4-5. [Philippines, Verde Island Passage, 13°38'45''N 121°01'12''E, 

296 m] 

Alpheus idiocheles Coutière, 1905b: 883; Plate 75, Figs 21-21c. [Minikoi] 

= Alpheus baculifer Coutière, 1908: 16. [Ile du Coin, Chagos] 

Alpheus immaculatus Knowlton & Keller, 1983: 354; Figs 1-4. [East Discovery Bay, Jamaica, near the central 

harbor buoy at approx. 10 m, from the anemone Bartholomea annulata] 

Alpheus inca Wicksten & Méndez G., 1981: 137; Figs 1-16. [Bahía de los Mejilones, Chile, 22°8'S 70.5'W] 

Alpheus inopinatus Holthuis & Gottlieb, 1958: 42; Figs 8-9. [Herzliya, littoral, Israel] 

Alpheus intrinsecus Spence Bate, 1888: 557; Plate 100, fig. 1. [Challenger (no stn), off Bahia, 7-20 fms] 

Alpheus japonicus Miers, 1879: 53. [34°6'N 136°15'E, 11 fms; 35°7'N 136°55'E, 3 fms; holotype designation 

by Banner & Banner, 1984 not valid] 

= Alpheus longimanus Spence Bate, 1888: 551; Plate 98, fig. 4. [Challenger (no stn), off Yokoska, Japan, 

5-20 fms; Challenger stns 233, (34°39'0"N 135°14'0"E) off Kobé, Japan, 8 fms; 233a (34°38'0"N 

135°1'0"E), off Kobé, Japan, 50 fms] 

Alpheus javieri Anker, Hurt & Knowlton, 2009: 7; Figs 2, 5C-D. [Panama, Pacific Coast, Río Mar, rocky 

intertidal, under rocks and from crevices in rocky reef platform] 

Alpheus kagoshimanus Hayashi & Nagata, 2000: 1111; Figs 1-5. [Off Tarumizu, Kagoshima Bay, 223- 

225 m] 

Alpheus kuroshimensis Nomura & Anker, 2005: 119; Figs 7-8, 13G-H. [Cyan, Kuro-shima, Yaeyama Islands, 


Alpheus labis Banner & Banner, 1982a: 127; Fig. 35. [Albany Passage area, Torres Straits, Queensland] 

Alpheus lacertosus W. Kim & Abele, 1988: 68; Fig. 28. [Algarrobo, CA, 7 km south of Valparaiso, Valparaiso 

Province, Chile] 

Alpheus ladronis Banner, 1956: 360; Fig. 20. [Saipan] 

Alpheus lanceloti Coutière, 1905b 

= Alpheus Lanceloti Coutière, 1905b: 900; Plate 83, fig. 39-39e. [North Male Atoll; Hulule Male Atoll; 

Goidu Atoll] 

Alpheus lanceostylus Banner, 1959: 136; Fig. 3. [Pearl and Hermes Reef, Hawaiian Archipelago, 27°N 

146°W] 

Alpheus latus W. Kim & Abele, 1988: 88; Fig. 37. [west side of Panama Canal, Panama, in mangroves] 

Alpheus lentiginosus Anker & Nizinski, 2011: 50; Figs 1-3. [North Atlantic Ocean, Gulf of Mexico, 188.8 

km south of Louisiana, 28°11'41"N 89°47'59"W, depth 434-438 m, base of Lophelia pertusa] 

Alpheus lepidus De Man, 1908d: 106. [Madura-bay and other localities in the southern part of Molo-strait, 

54-90 m; 0°58'.5N 122°55'E, Kwandang-bay-entrance, 75 m] 

Alpheus leptocheles Banner & Banner, 1975b: 261; Fig. 1. [mouth of Sepik River, New Guinea, between 

Cape Girgir and Kaup, 1.5-10 fms] 

Alpheus leptochiroides De Man, 1909a: 110. [5°36'5S 132°55'.2E, 2.3 miles N, 63°W from the north point of 

Nuhu Jaan, Kei-islands, 90 m]


Alpheus leptochirus Coutière, 1905b: 914; Plate 87, Figs 54-54e. [North Male Atoll, Mahlosmadulu Atoll] 

Alpheus leviusculus Dana, 1852b 

= Alpheus Edwardsii var. leviusculus Dana, 1852b: 543; Plate 34, Figs 3a-f. [Wakes Island, north Pacific] 

= Alpheus Bouvieri var. Bastardi Coutière, 1898b: 133 (partim); Fig. 1a. [côte Ouest, Nossi-Lava, baie 

de Bombétock (Madagascar); Mascate; Djibouti; nec specimen from Panama (? = Alpheus hebes W. 

Kim & Abele, 1988)] 

Alpheus lobidens De Haan, 1849 [in De Haan, 1833-1850] 

= Alpheus lobidens De Haan, 1849 [in De Haan, 1833-1850]: 179. [Japan] 

= Alpheus crassimanus Heller, 1862a: 526. [Nicobaren] 

= Alpheus crassimanus Heller, 1865: 107; Plate 10, fig. 2. [Nicobaren] 

= Alpheus lobidens polynesica Banner & Banner, 1975a: 429; Fig. 3A-H, J-L. [outer side of Heeia Fish 

Pond, Kaneohe Bay, Oahu, Ahwaii, 21°26'23''N 157°48'29''W] 

Alpheus longecarinatus Hilgendorf, 1879: 833; Plate 4, Figs 3-7. [Zanzibar] 

Alpheus longichaelis Carvacho, 1979: 459; Figs 7-8. [Grand-Cul-de-Sac] 

Alpheus longiforceps Hayashi & Nagata, 2002: 79; Figs 3-6. [Japan, Kii Strait, Wakanoura Bay, Wakayama 

Prefecture, 35 m] 

Alpheus longinquus W. Kim & Abele, 1988: 65; Fig. 27. [Braithwaite Bay, Socoro Island, Mexico] 

Alpheus lottini Guérin, 1829 [in Guérin-Méneville, 1829-1838] 

= Alpheus Lottini Guérin, 1829 [in Guérin-Méneville, 1829-1838]: Plate 3, fig. 3 (1829); 38 (1838). 

[Nouvelle-Irlande] 

= Cancer (Astacus) sublucanus Forskål, 1775: 94. [Djiddæ; suppressed under the plenary powers for 

the purposes of the Principle of Priority but not those of the Principle of Homonymy in Opinion 

1367 in 1987] 

= A.[lpheus] ventrosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 352. [les côtes de 

l’Ile-de-France] 

= A.[lpheus] lævis Randall, 1840: 141. [Sandwich Islands] 

= Alpheus Thetis White, 1847a: 75. [nomen nudum] 

= Alpheus Rouxii Guérin-Méneville, 1857: 50. [Bombay] 

= Crangon latipes Banner, 1953: 82; Fig. 27. [off Waikiki Reef, Oahu, 20 ft] 

Alpheus lutosus Anker & De Grave, 2009: 812; Figs 1-18. [Kuwait, Shatt-Al-Arab delta, Bubiyan Island, 

Ras Al-Barshah and area bordering Khor Al-Subbiya, approx. 29°36'N 48°188'E, muddy intertidal, 

in burrow with gobies] 

Alpheus macellarius Chace, 1988: 35; Figs 6-7. [vicinity of Cebu City, Cebu, Philippines] 

Alpheus macrocheles (Hailstone, 1835b) 

= Hippólyte rùbra Hailstone, 1835a: 272; Fig. 31. [off Hastings] 

= Hippólyte rùbra Westwood, 1835b: 274. [off Hastings] 

= Hippólyte macrochèles Hailstone, 1835b: 395. [nomen novum for Hippólyte rùbra Hailstone, 1835a 

nec Cryptophthalmus ruber Rafinesque, 1814] 

= A.[lpheus] Edwardsii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 352. [la Méditerranée] 

= Alphaeus latimanus Hope, 1851: 43. [Type locality not indicated] 

= Alpheus affinis Guise, 1854: 280; Figs 1-6. [Herm, Channel Islands, tide pool at low water] 

= Alpheus Milnei Guérin-Méneville, 1857: 49. [nomen novum for A.[lpheus] Edwardsii H. Milne Edwards, 

1837 [in H. Milne Edwards, 1834-1840 [nec Athanas Edwarsii Audouin, 1826] 

= Alpheus platyrhynchus Heller, 1862b: 400; Plate 1, Figs 21-24. [im adriatischen Meere, Lissa und 

Lesina] 

= Alpheus megacheles Norman, 1868: 175. [lapsus for Hippólyte macrochèles Hailstone, 1835b] 

= Nika platyura Fischer, 1872: 419. [Cap Breton (Landes), 28 brasses] 

Alpheus macroskeles Alcock & Anderson, 1899: 153. [Investigator stn 129, Bay of Bengal, off Godávari 

Delta, 270 fms; 162, off Pulicat, 13°51'12''N 80°28'12''E, 145-250 fms, the Swatch, 193 fms; Andaman 

Sea, 193 fms] 

Alpheus maindroni Coutière, 1898b 

= Alpheus Maindroni Coutière, 1898b: 133; Figs 2-2’. [Mascate; Dijbouti]


Alpheus malabaricus (Fabricius, 1775) 

= Astacus Malabaricus Fabricius, 1775: 415. [ad littora Malabarica] 

= Alpheus forceps White, 1847a: 75. [nomen nudum] 

= Alpheus dolichodactylus Ortmann, 1890: 473; Plate 36, fig. 11. [Japan, Tokiobai] 

= Alpheus macrodactylus Ortmann, 1890: 473; Plate 36, fig. 10, 10l. [Sydney] 

= Alpheus dolichodactylus var. leptopus De Man, 1910a: 289. [7°15'S 115°15'E, 289 m; Bay of Labuan 

Tring, west coast of Lombok, 8°44'.5S 116°2'.5E, 18-27 m; Saleyer-anchorage and surroundings, 18- 

45 m; Sapeh-bay, East coast of Sumbawa, up to 36 m] 

= Alpheus malabaricus mackayi Banner, 1959: 149; Fig. 12. [Wailupe fish pond, Oahu, Hawaii] 

= Alpheus malabaricus songkla Banner & Banner, 1966a: 147; Fig. 56. [Lake Songkla, 1 m] 

= Alpheus malabaricus trefzae Banner & Banner, 1982a: 207; Fig. 64. [Brammo Bay, Dunk Island, near 

Tully, northern Queensland, 0 tide level] 

Alpheus malleator Dana, 1852a: 23. [in portu Rio Janeiro?] 

= Alpheus pugilator A. Milne-Edwards, 1878: 229. [toutes les îles du Cap-Vert] 

? = Alpheus tuberculosus Osorio, 1892: 201. [Iogo-Iogo] 

= Alpheus malleator var. edentatus Zimmer, 1913: 387; Figs G-M. [Barbados (Riff)] 

Alpheus malleodigitus (Spence Bate, 1888) 

= Betæus malleodigitus Spence Bate, 1888: 565; Plate 101, fig. 5. [Levuka, Fiji Islands, reefs] 

= Alpheus obesomanus var. japonicus Ortmann, 1890: 478. [Japan, Tokiobai] 

= Alpheus Danae Coutière, 1905b: 887; Plate 72, figs 26-26d. [Hulule Male Atoll] 

= Alpheus phrygianus Coutière, 1905b: 886; Plate 72, figs 25-25e. [Hulule Male Atoll; Suvadiva Atoll; 

Madagascar; l’île Tague] 

= Alpheus persicus Nobili, 1905e: 238. [Golfe Persique] 

= Alpheus malleodigitus var. gracilicarpus De Man, 1909a: 99. [Bank between islands of Bahuluwang 

and Tambolungan, south of Saleyer, 8-10 m; Pulu Sanguisiapo, Tawi-Tawi-islands, Sulu archipelago, 

12 m; 6°7'.5N 120°26'E, anchorage off North-Ubian, 16-23 m; 8°25'.2S 127°18'.4E, anchorage between 

Nusa-Besi and the N.E. point of Timor, 27-54 m] 

Alpheus martini W. Kim & Abele, 1988: 98; Fig. 41. [Venado Beach, Panama] 

Alpheus mazatlanicus Wicksten, 1983a: 46; Figs 7-8. [Laguna Caimaero, Sinaloa, México] 

Alpheus microrhynchus De Man, 1897 

= Alpheus microrhynchus De Man, 1897: 754; Plate 36, Figs 65-65b (as Alpheus sp.). [Pontianak, Westküste 

von Borneo] 

Alpheus microscaphis (Banner, 1959) 

= Thunor microscaphis Banner, 1959: 151; Fig. 13. [Likiep Atoll, Marshall Islands] 

Alpheus microstylus (Spence Bate, 1888) 

= Betæus microstylus Spence Bate, 1888: 566; Plate 101, fig. 6. [Albany Island, Cape York] 

= Alpheus microstylus var.? De Man, 1911d: 345. [Anchorage off Pulu Kawassang, Paternoster-islands, 

coralreef] 

Alpheus miersi Coutière, 1898c 

= Alpheus rapax var. Miersi Coutière, 1898c: 166; Figs 1-1b. [Port-Molle; Ceylan; Japon, 32°49'N 

128°54'E] 

? = Alpheus gracilipes var. serratus Coutière, 1898a: 166. [Type locality not indicated] 

Alpheus migrans Lewinsohn & Holthuis, 1978: 76; Figs 1-2. [Mediterranean, off the north coast of Sinai 

Peninsula, 31°26'N 34°11'E, 36.6 m] 

Alpheus millsae Anker, Hurt & Knowlton, 2007b: 50; Figs 3-4, 5C, 6C, 7G-H, 8C. [Panama, Panama City, 

off Casco Viejo, near extreme low tide mark] 

Alpheus mitis Dana, 1852a: 22. [in freto ”Balabac”] 

Alpheus moretensis Banner & Banner, 1982a: 177; Fig. 54. [27°10'S 153°21'E, 8 fms] 

Alpheus naos Anker, Hurt & Knowlton, 2007a: 12; Figs 7-9, 11d-e. [Panama, Pacific coast, Amador causeway 

near Panama City, Punta Culebra, rocky shore, under large mud-covered rocks on muddy 

sand, in burrow of Listriolobus sp., extreme low tide] 

Alpheus nipa Banner & Banner, 1985: 21; Fig. 2. [4°20'N 98°54'E, Straits of Malacca, roughly 60 km from 

shore in the vicinity of Medan, Sumatera [sic], from floating frond of nipa palm]


Alpheus nobili Banner & Banner, 1966b: 172; nomen novum for Alpheus sp. Nobili, 1906c. [bancs d’huîtres 

perlières entre 25°10'-24°55'N 55°10'-54°40'E] 

Alpheus nonalter Kensley, 1969: 172; Fig. 15. [29°34'S 31°39'E, 118 m] 

Alpheus normanni Kingsley, 1878a: 93. [nomen novum for Alpheus affinis Kingsley, 1878b] 

= Alpheus affinis Kingsley, 1878b: 195; nec Guise, 1854. [Panama] 

Alpheus notabilis Stebbing, 1915: 80; Plates 20- 21. [Delagoa Bay] 

Alpheus novaezealandiae Miers, 1876a 

= Alpheus novæ-zealandiæ Miers, 1876a: 224. [New Zealand; illustrated in Miers, 1876b; lectotype 

selection by Yaldwyn, 1956] 

Alpheus nuttingi (Schmitt, 1924b) 

= Crangon nuttingi Schmitt, 1924b: 78; Plate 2, Figs 4-6. [Pelican Island, shallow] 

Alpheus oahuensis (Banner, 1953) 

= Crangon oahuensis Banner, 1953: 64; Fig. 20. [off Waikiki Reef, Oahu, 20ft] 

Alpheus obesomanus Dana, 1852a 

= Alpheus obeso-manus Dana, 1852a: 21. [in archipelago ”Viti”] 

= Alpheus Lutini Coutière, 1905b: 885; Plate 76, Figs 24-24e. [Hulule Male Atoll; Samoa; l’île Tague] 

Alpheus ovaliceps Coutière, 1905b: 888; Plate 77, figs 27-27e. [Minikoi] 

Alpheus pachychirus Stimpson, 1860a: 30. [ad insulam ”Loo Choo”] 

Alpheus pacificus Dana, 1852a: 21. [Kalama Park, about 18 miles southeast of Lahaina, Maui, 3ft; neotype 

selection by Banner, 1953, as Crangon pacifica] 

= Alpheus gracilidigitus Miers, 1884b: 287. [Fiji Islands; Toyota; Sandwich Islands] 

Alpheus packardii Kingsley, 1880: 417. [Key West] 

= Alpheus bermudensis Spence Bate, 1888: 547; Plate 98, fig. 3. [Bermuda, St. Thomas, West Indies] 

= Alpheus beanii Verrill, 1922: 81; Fig. 7; Plate 22, fig. 1; Plate 32, figs 1a-u. [Challenger Bank, 24 fms] 

Alpheus paludicola Kemp, 1915: 303; Fig. 33; Plate 13; figs 11-13. [Outer Channel, Lake Cilka, Orissa, India; 

lectotype selection by Yeo & Ng, 1996] 

Alpheus panamensis Kingsley, 1878b: 192. [Acajutla, Central America; Panama] 

Alpheus papillosus Banner & Banner, 1982a: 260; Fig. 80. [Sandgate, Moreton Bay, Queensland] 

Alpheus paracrinitus Miers, 1881a: 365; Plate 16, fig. 6. [Goree Bay, Senegambia, 9-15 fms] 

= Alpheus ascensionis Ortmann, 1893: 45. [Plankton-Expedition der Humboldt-Stiftung, Ascension, 

JN 200 (7.9°S 14.4°W) 20 m] 

= Alpheus paracrinitus var. Bengalensis Coutière, 1905b: 901; Plate 82, figs 37-37f. [Minikoi] 

= Crangon togatus Armstrong, 1940: 2; Fig. 1. [Piedra Prieta Reef, Barahone, St. Domingo; Grazbury’s 

Island, Bermuda] 

Alpheus paradentipes Coutière, 1905b: 880; Plate 74, figs 17-17d. [Miladumadulu Atoll; North Male Atoll] 

Alpheus paraformosus Anker, Hurt & Knowlton, 2008b: 12; Figs 5-6, 7f-h. [Bocas del Toro, San Cristóbal, 

Punta Coco, shallow subtidal (1-1.5 m), from rock and coral rubble crevices] 

Alpheus paralcyone Coutière, 1905b: 895; Plate 80, figs 34-34e; Plate 81, figs 34f-h. [South Nilandu Atoll; 

Miladumadulu Atoll; North Male Atoll; Mahlos Atoll] 

= Alpheus Providencei Coutière, 1908: 208. [Providence, 50-78 brasses] 

= Crangon laysani Edmondson, 1925: 17; Fig. 3. [Laysan Island, shallow water] 

Alpheus paralpheopsides Coutière, 1905b: 902; Plate 83, figs 41-41e. [North Male Atoll, Suvadiva Atoll] 

Alpheus parasocialis Banner & Banner, 1982a: 72; Fig. 17. [Palm Beach, Rockingham, Western Australia, 

from jetty piles] 

Alpheus pareuchirus pareuchirus Coutière, 1905b 

= Alpheus pareuchirus Coutière, 1905b: 906; Plate 84, figs 43-43d. [North Male Atoll] 

= Alpheus pareuchirus var. Leucothea De Man, 1909a: 105. [7°25'S 113°16'E, Madura-Strait, 56 m; 

8°23'.5S 119°4'.6E, Sapeh-Strait, 70 m; Anchorage off Dongala, Palos-bay, Celebes, 36 m; South-east 

side of Pearl-bank, Sulu-archipelago, 15 m] 

Alpheus pareuchirus imitatrix De Man, 1909a 

= Alpheus pareuchirus var. imitatrix De Man, 1909a: 106. [0°3'8N 130°24'3E, 141 m; Between Loslos 

and Broken-islands, west-coast of Salawatti, 18 m; 1°42'.5S 130°47'.5E, 32 m; Mid-channel in Solorstrait 

of Kampog Menanga, 113 m]


Alpheus parvimaculatus Nomura & Anker, 2005: 122; Figs 9-10, 14A-C. [Fuki, Kuro-shima, Yaeyama Islands, 


Alpheus parvirostris Dana, 1852a 

= Alpheus parvi-rostris Dana, 1852a: 22. [in freto ”Balabac”] 

= Alpheus lineifer Miers, 1875: 343. [Samoa Islands] 

= Alpheus euchiroides Nobili, 1906b: 257. [Marutea] 

= Alpheus braschi Boone, 1935: 131; Fig. 10; Plate 34. [Pago Pago, Samoa] 

Alpheus parvus De Man, 1909a: 102. [Anchorage off Lirung, Salibabu island, 36 m] 

Alpheus peasei (Armstrong, 1940) 

= Crangon peasei Armstrong, 1940: 1. [Castle Harbor Reefs, Bermuda] 

Alpheus percyi Coutière, 1908 

= Alpheus Percyi Coutière, 1908: 211. [Cargados Carajos, 30 brasses] 

Alpheus perezi Coutière, 1908 

= Alpheus Perezi Coutière, 1908: 212. [Golfe Persique] 

Alpheus perplexus Banner, 1956: 347; Fig. 13. [1.2 miles offshore, immediately south of uncovering reef SE 

of Mañgaha Islet, 24ft] 

Alpheus philoctetes De Man, 1909a 

= Alpheus Philoctetes De Man, 1909a: 103. [Banda, 9-36 m] 

Alpheus platydactylus Coutière, 1897d: 306. [Travailleur stn 52, au large de Madère (32°30' 31°N 18°51'E 

(of Paris)), 100 m; lectotype designation by Forest, 1965] 

Alpheus platyunguiculatus (Banner, 1953) 

= Crangon platyunguiculata Banner, 1953: 130; Fig. 47. [Waialua Bay, Oahu, 6ft] 

Alpheus polystictus Knowlton & Keller, 1985: 894; Fig. 1. [Morrocoy, Venezuela, 1-10 m] 

Alpheus polyxo De Man, 1909a 

= Alpheus Polyxo De Man, 1909a: 108. [Banda-anchorage, 9-45 m] 

Alpheus pontederiae de Rochebrune, 1883 

= Alpheus Pontederiæ de Rochebrune, 1883: 174. [Marigots de Leybar, Thiank, Dakar-Bango] 

= Crangon langi Schmitt, 1926b: 20; Fig. 63. [Banana] 

? = Alpheus heterochaelis var. orientalis Vilela, 1949: 53; Fig. 3. [canal de Mantambua, porto de Passõ; 

rio Baboc, Canchungo; Ilha de Rubane; foz de rio Grande (all Guiné Portuguesa)] 

Alpheus pouang Christoffersen, 1979: 324; Figs 14-15. [23°39'S 43°37'W, off the State of São Paulo, 120- 

121 m] 

Alpheus praedator De Man, 1908d: 103. [Ambon, reef-exploration] 

Alpheus proseuchirus De Man, 1908d: 111. [0°58'.5N 122°55'E, Kwandang-bay-entrance, 75 m; 0°58'.5N 

122°42'.5E, West of Kwandang-bay-entrance, 72 m] 

Alpheus pseudopugnax (Banner, 1953) 

= Crangon pseudopugnax Banner, 1953: 122; Fig. 44. [Kalama Park, southeast Maalea Bay, Maui, 8ft] 

Alpheus puapeba Christoffersen, 1979: 328; Figs 16-18. [35°05'S 52°33'W, off the province of Buenos Aires, 

115 m] 

Alpheus pubescens De Man, 1908d: 109. [Anchorage off Pulu Sarassa, Postillon-islands, up to 30 m; 

Haingsisi, Samau-island, Timor, 36 m; Makassar, up to 32 m; 3°27'S 117°36'E, Borneo-bank, 59 m; 

between Loslos and Broken-islands, west-coast of Salawatti, 18 m; Anchorage off Pulu Jedan, East 

coast of Aru-islands, 13 m] 

Alpheus pugnax Dana, 1852a: 23. [ad insulam ”Maui” Hawaiensem] 

Alpheus pustulosus Banner & Banner, 1968: 143; Fig. 2. [21°N 114°E, 30-40 fms] 

Alpheus quasirapacida Chace, 1988: 50; Fig. 12. [Off Tawitawi, Sulu Archipelago, Philppines, 5°11'50''N 

119°54'E, 18 m] 

Alpheus randalli Banner & Banner, 1980: 401; Figs 1-3. [Nuka Hiva, Marquesas Islands, northwest side 

Sentinelle de l’Est, 18 m commensal with Amblyeleotris sp.] 

Alpheus rapacida De Man, 1908d: 105. [Ruma-Kuda-bay, Roma-island, 36 m; Anchorage east of Dangar 

Besar, Saleh-bay, up to 36 m] 

Alpheus rapax Fabricius, 1798: 405. [in India orientali]


Alpheus rectus W. Kim & Abele, 1988: 16; Fig. 6. [Bahia Honda, Panama] 

Alpheus ribeiroae Anker & Dworschak, 2004: 48; Figs 1-27. [tidal flat at Moia-Moia, NE coast of Ilha do 

São Tiago, Cabo Verde] 

Alpheus richardsoni Yaldwyn, 1971: 88. [off Mariott Id., Waikare River Inlet, Bay of Islands, 1-3 fms] 

Alpheus rudolphi Almeida & Anker, 2011: 1; Figs 1-22. [REVIZEE Program, Score Nordeste III stn 171, off 

Alagoas, Brazil, 9°55.11'S 35°32.73'W, 49 m] 

Alpheus romensky Burukovsky, 1990: 197; Fig. 3A.1-6. [25°34'S 89°04'W, 540-560 m] 

Alpheus roquensis Knowlton & Keller, 1985: 897; Fig. 1. [Los Roques, Venezuela, 3-16 m] 

Alpheus roseodigitalis Nomura & Anker, 2005: 126; Figs 10-11, 14D-F. [Aragusuku-jima, Yaeyama Islands, 

southern Ryukyu Archipelago, Japan, intertidal] 

Alpheus rostratus W. Kim & Abele, 1988: 51; Fig. 21. [Darwin Bay, Tower Island, Galapagos Islands] 

Alpheus rugimanus A. Milne-Edwards, 1878: 230. [les îles du Cap-Vert] 

Alpheus samoa Banner & Banner, 1966b: 174. [”Lion’s Head”, Tutuila, American Samoa] 

Alpheus savuensis De Man, 1908d: 110. [Anchorage off Seba, Savu] 

Alpheus saxidomus Holthuis, 1980d: 47; Figs 1-2. [Playa Tamarindo, Nicoya Peninsula, Pacific coast of 

Costa Rica, about 10°19'N 85°53'W, boring in basaltic rock in the surf zone] 

Alpheus schmitti Chace, 1972: 70; Figs 21-22. [Antigua Island, Falmouth Harbour, beach north of Black’s 

Point, 2-3 ft] 

Alpheus scopulus W. Kim & Abele, 1988: 77; Fig. 32. [Whorehouse reef tidepools, Panama] 

Alpheus serenei Tiwari, 1964b: 314. [St. Cauda, 3-4 m; fully described in Tiwari, 1965] 

Alpheus sibogae De Man, 1908d: 107. [8°23'.5S 119°4'.6E, Sapeh-strait, 70 m; Madura-bay and other localities 

on the southern part of Moho-strait, 54-90 m; Mid-channel in Solor-strait off Kampong Menanga, 

113 m] 

Alpheus simus Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856] 

= Alpheus Simus Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xix; Plate 2, fig. 11. [Cuba] 

= Crangon rathbunae Schmitt, 1924b: 74; Plate 1; nec Alpheus Rathbuni Coutière, 1900b. [Needham’s 

Point, Barbados] 

Alpheus sizou Banner & Banner, 1967 

= Alpheus amirantei sizou Banner & Banner, 1967: 265. [north side of Bora Bora, outer reef platform, 1ft] 

Alpheus socialis Heller, 1862a 

= Alpheus socialis Heller, 1862a: 526. [Aukland] 

= Alpheus socialis Heller, 1865: 106; Plate 10, fig. 1. [Auckland; Sidney] 

= Alpheus Doto White, 1847a: 75 [nomen nudum] 

Alpheus soelae Banner & Banner, 1986: 21; Fig. 1. [18°41'S 120°07'E, 430 m] 

Alpheus soror Bruce, 1999: 454; Figs 1A, 2-3. [off Trincomalee, Sri Lanka] 

Alpheus spatulatus Banner & Banner, 1968: 146; Fig. 3. [19°17'N 112°81'E, 52 fms] 

Alpheus splendidus Coutière, 1897c: 236. [Djibouti] 

= Alpheus pomatoceros Banner & Banner, 1966a: 93; Fig. 32. [Lam Chongklan, Koh Samui, Thailand, 

outer reef front, 2-3 m] 

Alpheus spongiarum Coutière, 1897c: 236. [Djibouti] 

= Alpheus paraculeipes Coutière, 1905b: 894; Plate 79, figs 32-32b'; Plate 80, figs 32c-32g. [South Nilandu 

Atoll; Mahlosmadulu Atoll; North et South Male Atoll] 

Alpheus stanleyi Coutière, 1908 

= Alpheus Stanleyi Coutière, 1908: 207. [Amirante, 30 brasses] 

= Alpheus Stanleyi var. dearmatus De Man, 1910a: 287. [Wunoh-bay, N.W. coast of Waigeu-island, 

reef; Banda-anchorage, 9-36 m] 

= Alpheus cloudi Banner, 1956: 352; Fig. 16. [about 0.6 mile off Muchot Point, between reef clusters at 

south side of Saipan harbor, 5-15ft] 

Alpheus stantoni Banner & Banner, 1986: 24; Fig. 2. [Heron Island, Great Barrier Reef, dug out of coral 

rubble, 13 m] 

Alpheus staphylinus Coutière, 1908: 204. [Salomon Island, Chagos] 

Alpheus stephensoni Banner & Smalley, 1969: 43; Fig. 2. [Moreton Bay, Queensland, 4-5 fms]


Alpheus strenuus strenuus Dana, 1852a: 21. [insula Tongabatu] 

= Alpheus Alope White, 1847a: 74. [nomen nudum] 

= Alpheus Rhode White, 1847a: 74. [nomen nudum] 

= Alpheus Doris White, 1847a: 75. [nomen nudum] 

= Alpheus strenuus var. angulatus Coutière, 1905b: 914. [Hulule Male Atoll] 

Alpheus strenuus cremnus Banner & Banner, 1982a: 229; Fig. 72. [Minnie Waters, near Grafton, New South 

Wales, intertidal rock platform] 

Alpheus styliceps Coutière, 1905b: 889; Plate 78, figs 28-28d. [Naifaro Reef] 

Alpheus sudara Banner & Banner, 1966a: 153; Fig. 59. [Phuket, Thailand, outer reef edge, 2 m] 

Alpheus sulcatus Kingsley, 1878b: 193. [Bay of Panama; Zorritas, Peru] 

= Alpheus macrochirus Richters, 1880: 164; Plate 17, figs 31-33. [Fouquets] 

= Alpheus luciæ Barnard, 1947: 389. [St. Lucia Bay, Zululand] 

Alpheus suluensis Chace, 1988: 57; Figs 15-16. [near Siasi, Sulu Archipelago, 5°41'40''N 120°47'10''E, 38 m] 

Alpheus supachai Banner & Banner, 1966a: 89; Fig. 30. [Koh Samui, west side of Gulf of Thailand, 3 m] 

Alpheus superciliaris Coutière, 1905b: 896; Plate 81, figs 35-35g. [Reef Naifaro] 

Alpheus talismani Coutière, 1898d 

= Alpheus Talismani Coutière, 1898d: 31; Figs 3-4. [îles du Cap-Vert, 410-450 m] 

Alpheus tasmanicus Banner & Banner, 1982a: 215; Fig. 67. [Tasmania] 

Alpheus tenuicarpus De Man, 1908d: 104. [0°58'.5N 122°55'E, Kwandang-bay-entrance, 75 m; Sapeh-bay, 

East coast of Sumbawa up to 36 m] 

Alpheus tenuipes De Man, 1910a: 288. [4°20'S 122°58'E, between islands of Wowoni and Buton, northern 

entrance of Buton-Strait, 75-94 m] 

Alpheus tenuis W. Kim & Abele, 1988: 79; Fig. 33. [Farfan Point, Panama] 

Alpheus thomasi Hendrix & Gore, 1973: 415; Figs 1-3. [Virginia Beach, Virginia Key, Miami, Florida, 1 m] 

Alpheus tirmiziae Kazmi, 1974: 170; Figs 1-4. [Bay of Bengal, 20°05'-20°30'N 91°50'-90°50'E] 

Alpheus tricolor Anker, 2001: 58; Figs 1a-b, 2a, 3a-e, 4a-d, 5a-h, 6a-k. [Bali, Indonesia] 

Alpheus triphopus Nobili, 1906b 

= Alpheus aculeipes var. triphopus Nobili, 1906b: 257. [Marokau] 

Alpheus tungii Banner & Banner, 1966b: 160. [south coast of Tongatabu, Tonga, outer edge of fringing 

reef near surge channel] 

Alpheus umbo W. Kim & Abele, 1988: 84; Fig. 35. [Isabel Island, Sinaloa, Mexico] 

Alpheus utriensis Ramos & von Prahl, 1989: 477; Figs 1-2. [Utria Sound] 

Alpheus vanderbilti Boone, 1930c: 163; Plate 58, fig. 5. [south of Sand Key, Key West, Florida, 20 fms] 

Alpheus villosus (Olivier, 1811) 

= Palæmon villosus Olivier, 1811: 664. [la mer des Indes] 

= Palæmon diverfimanus Olivier, 1811: 663. [côtes de la Nouvelle-Hollande] 

Alpheus villus W. Kim & Abele, 1988: 82; Fig. 34. [spit north of mill site, Angeles Bay, Baja California, 

Mexico] 

Alpheus viridari (Armstrong, 1949) 

= Crangon viridari Armstrong, 1949: 8; Fig. 2. [inside North end of El Cayo, Dominican Republic] 

Alpheus waltervadi Kensley, 1969: 175; Fig. 16. [Walter’s Shoal, 33°13'S 43°51'E, 38-46 m] 

Alpheus websteri Kingsley, 1880: 416. [Key West] 

= Alpheus Ridleyi Pocock, 1890: 518. [Fernando de Noronha] 

= Alpheus nigro-spinatus Rankin, 1898: 249; Plate 30, fig. 6. [New Providence] 

Alpheus wickstenae Christoffersen & Ramos, 1987: 333; Figs 1-3. [Rampa de los Suecos, Bahía Málaga, 

Pacific coast of Colombia, 3°59'N 77°20'W] 

Alpheus williamsi Bruce, 1994b: 19; Figs 3, 5C. [north of Charles Point, Northern Territory, Australia, 

12°17.18'S 30°40.06'N, 18-24 m] 

Alpheus xishaensis Liu & Lan, 1980: 102; Fig. 13. [Xisha Island, coral reef] 

Alpheus xanthocarpus Anker, Hurt & Knowlton, 2008a: 556; Figs 6, 7E-F. [São Tomé, Ilha Santana, 15-20 m] 

Alpheus zimmermani Anker, 2007a: 242; Figs 1-3. [Caribbean Sea, British Virgin Islands, Guana island, 

near end of Long Point, 1.5-11 m]


Alpheus zulfaquiri Kazmi, 1982: 137; Figs 1-2. [Manora Island, coastal waters of Karachi, northern Arabian 

Sea] 

Amphibetaeus Coutière, 1897b 

= Amphibetæus Coutière, 1897b (type species Betæus Jousseaumei Coutière, 1896, by monotypy, gender 


Amphibetaeus jousseaumei (Coutière, 1896) 

= Betæus Jousseaumei Coutière, 1896: 313; Figs 1-12. [Perim; Tadjourah] 

= Amphibetæus Jousseaumei Coutière, 1897b: 384. [nomen novum for Betæus Jousseaumei Coutière, 1896] 

Arete Stimpson, 1860a 

= Arete Stimpson, 1860a (type species Arete dorsalis Stimpson, 1860a, by monotypy, gender feminine; 


Arete acanthocarpus (Miya & Miyake, 1968a) 

= Athanas acanthocarpus Miya & Miyake, 1968a: 157; Figs 10D, H, J, 11C, E, 12E, 13. [Kamiyama-jima, 

near Naha City, Okinawa-jima, Ryukyu Islands] 

Arete dorsalis Stimpson, 1860a: 32. [Cape D’Aguilar, Hong Kong Island, 1 m; neotype designation by 

Bruce, 1990g, as Athanas dorsalis] 

? = Athanas mascarenicus Richters, 1880: 164. [Insel Mauritius und Seychellen] 

= Arete dorsalis var. Pacificus Coutière, 1903: 85; Figs 31-34. [Hong-Kong (?); Nouvelle Calédonie; 

Samoa; Amérique Centrale; Amboine (??)] 

= Arete Maruteensis Coutière, 1905b: 864. [Marutea] 

= Arete maruteensis var. salibabuensis De Man, 1910a: 313. [Anchorage off Lirung, Salibabu-island, up 

to 36 m] 

Arete amboinensis De Man, 1910b: 25; Figs 1-6. [Insel Amboina] 

Arete indicus Coutière, 1903 

= Arete dorsalis var. Indicus Coutière, 1903: 84; Figs 25-29. [Djibouti; Hulule Male Atoll; Mascareignes 

(?)] 

= Arete Indicus Coutière, 1905b: 863; Figs 134-135. [Hulule Male Atoll] 

= Arete Iphianassa De Man, 1910a: 312. [Anchorage off Sawan, Siau-island, reef] 

= Arete intermedius Yu, 1931b: 513; Fig. 1. [?Amoy] 

? = Athanas kominatoensis Kubo, 1942c: 82; Figs 1-2. [Kominato, Prov. Awa (Bôsyû) 

Aretopsis De Man, 1910a 

= Aretopsis De Man, 1910a (type species Aretopsis amabilis De Man, 1910a, by monotypy, gender 

feminine) 

Aretopsis amabilis De Man, 1910a: 311. [Pulu Kaniungan ketjil, reef] 

= Aretopsis aegyptiaca Ramadan, 1936: 16; Plate 1, figs 9-10; Plate 2, figs 9-17. [Ghardaga, Red Sea] 

Aretopsis manazuruensis Suzuki, 1971: 19; Figs 10-12. [Shikkake, near the Manazuru M.B.L., intertidal] 

Athanas leach, 1814 [in leach, 1813-1814] 

= Athanas Leach, 1814 [in Leach, 1813-1814] (type species Palaemon nitescens Leach, 1813 [in Leach, 

1813-1814], by monotypy, gender masculine; name placed on the Official List of Generic Names in 


Athanas ahyongi Anker & Komai, 2010: 46; Figs 1-6. [Madagascar, Nosy-Bé, west of Hellville, 13°41.56'S 

48°24.65'E, 0.5-1 m] 

Athanas amazone Holthuis, 1951a: 111; Fig. 23. [off Nigeria, 4°01'N 7°12'E, 52 m] 

Athanas anatidactylus Anker & Marin, 2007: 163; Figs 1-2, 3a. [Vietnam, Nhatrang Bay, Tre Island, 7 m, on 

unidentified crinoid] 

Athanas areteformis Coutière, 1903: 79; Figs 17-18. [Naifaro Reef; Hulule Male Atoll] 

= Athanas Naifaroensis Coutière, 1903: 77; Figs 14-16. [Naifaro Reef; Hulule Male Atoll] 

= Athanas Naifaroensis Coutière, 1905b: 859; Fig. 131. [Naifaro Reef; Hulule Male Atoll]


= Athanas aretiformis Coutière, 1905b: 860; Fig. 132. [Naifaro Reef; Fadifolu Atoll; Hulule Male Atoll] 

? = Athanas crosslandi Tattersall, 1921: 372; Plate 27, figs 13-17. [Khor Dongonab, Engineer Island, 

20°50'N, reef flat] 

= Athanas erythraeus Ramadan, 1936: 13; Plate 1, fig. 1. [Ghardaga, Red Sea] 

= Athanas dubius Banner, 1956: 322; Fig. 2. [Saipan] 

Athanas daviei Anker, 2011a: 5; Figs 3-4, 8E-F. [Australia, Queensland, Great Barrier Reef, Lizard Island, 

Watson’s Bay, 14°39'50.4''S 145°27'46.8''E, 1 m] 

Athanas dentirostris Anker, Jeng & Chan, 2001: 1049; Figs 1-3. [Cua Luc, N Vietnam, together with Upogebia 

aff. takaoensis] 

Athanas dimorphus Ortmann, 1894: 12; Plate 1, fig. 1-1z. [Dar-es-Salaam, Upanga-Riff] 

? = A.[lpheus] monoceros Heller, 1862c: 274. [im rothen Meere] 

? = Athanas transitans var. longispina Czerniavsky, 1884: 25. [nomen novum for A.[lpheus] monoceros 

Heller, 1862c] 

= Athanas solenomerus Coutière, 1897b: 381. [mer Rouge] 

= Athanas leptocheles Coutière, 1897b: 381. [mer Rouge] 

= Athanas dispar Coutière, 1897c: 233. [mer Rouge] 

= Athanas setoensis Kubo, 1951: 265; Figs 5-6. [vicinity of Seto Marine Biological Laboratory of Kyoto 

University] 

= Athanas dimorphus seedang Banner & Banner, 1966a: 28; Fig. 4. [Koh Krarard, Trad., southwest side 

of island, 1.5 m] 

Athanas djiboutensis Coutière, 1897c 

= Athanas Djiboutensis Coutière, 1897c: 234. [Djibouti] 

= Athanas sulcatipes Borradaile, 1899: 1011; Plate 65, figs 9-9i. [Funafuti] 

Athanas gracilipes Banner & Banner, 1978: 234; Fig. 3a-g. [12°02'N 112°49'E, 365-385 m] 

Athanas gracilis Boone, 1935: 122; Plate 32. [coral at Teviatoa reef, Raiatea Island, Society Islands] 

Athanas granti Coutière, 1908 

= Athanas Granti Coutière, 1908: 192. [South Adelaïde Coast] 

Athanas grimaldii Coutière, 1911b 

= Athanas Grimaldii Coutière, 1911b: 1; Figs 1-6. [Belle Ile, 155 m; Iles du Cap Vert, 16 m + 91 m] 

Athanas hasswelli Coutière, 1908 

= Athanas Hasswelli Coutière, 1908: 192. [South Adelaïde Coast; emendation to A. haswelli by Banner 

& Banner, 1973 is an unjustified emendation] 

Athanas hongkongensis Bruce, 1990f: 624; Figs 7-8. [Long Ke Wan, low water] 

Athanas iranicus Anker, Naderloo & Marin, 2010: 54; Figs 1-4. [Iran, Hormozgan, Qeshm Island north of 

Strait of Hormuz, northern coast, 2 km east of desalination center, 26°56'N 55°47'E, intertidal] 

Athanas ivoiriensis Anker & Ahyong, 2007: 23; Figs 3-4. [Côte d’Ivoire, Grand Bassam, 20 m] 

Athanas japonicus Kubo, 1936c: 43; Plate 8. [Mitaziri, Yamaguti Prefecture] 

= Athanas lamellifer Kubo, 1940d: 102; Figs 3-5. [Kominato (Matuga-hana), Tiba Prefecture, near low 

water mark] 

Athanas jedanensis De Man, 1910a: 313. [Anchorage off Pulau Jedan, east coast of Aru-islands (Pearlbanks)] 

Athanas locincertus Banner & Banner, 1973a: 311; Fig. 5. [Panchoran Buoy] 

Athanas marshallensis Chace, 1955: 17; Fig. 8. [Eniwetok Atoll, Bogombogo Island, intertidal] 

? = Athanas esakii Kubo, 1940e: 93; Fig. 13. [Kusaie, Caroline Islands] 

Athanas minikoensis Coutière, 1903: 76; Figs 9-11. [Minikoi Atoll] 

= Athanas Minikoensis Coutière, 1905b: 858; Fig. 130. [Minikoi Atoll] 

Athanas naga Banner & Banner, 1966a: 26; Fig. 3. [Gulf of Thailand, 10°12'48''N 103°32'30''E, 27-30 m] 

Athanas nitescens (Leach, 1813 [in Leach, 1813-1814]) 

= Palaemon nitescens Leach, 1813 [in Leach, 1813-1814]: 401. [southern coast of Devonshire] 

= P.[alemon] Lævirhincus Risso, 1816: 108. [environs de Nice, sur les bas fonds] 

= Cancer listellus Nardo, 1847: 6. [Trovato nel golfo, tra gli Asprei] 

= Alpheus ? vittatus Nardo, 1847: 6. [Trovato nel golfo, tra gli Asprei]


= Arete Diolectiana Heller, 1862b: 404; Plate 1, Figs 28-33. [Pirano] 

= Athanas veloculus Spence Bate, 1888: 529; Plate 96, fig. 1. [Cape Verde Islands] 

= Athanas transitans var. pontica Czerniavsky, 1884: 25. [Black Sea] 

= Athanas nitescens forma rotundicauda Czerniavsky, 1884: 24; Plate 2, figs 5A-D. [promontorium 

Ivan-baba (prope Theodosia, Strembitzki)] 

= Athanas alpheoides Czerniavsky, 1884: 26. [Portus Suchum, zona littor., 1-2 m] 

Athanas nouvelae Holthuis, 1951a: 104; Fig. 22. [off São Pedro Bay, São Vicente, Cape Verde Islands, 

22 m] 

Athanas orientalis Pearson, 1905: 88; Plate 2, figs 10-10b. [Cheval Paar, 6.5-13 fms; across Muttuvaratu 

Paar, 6-9 fms; south of Adam’s bridge, Gulf of Manaar, 4-40 fms; northern end of Periya Paar, 12- 

14 fms] 

Athanas ornithorhynchus Banner & Banner, 1973a: 319; Fig. 8. [Chambers Bay, Van Diemen Gulf, Northern 

Territory] 

Athanas ohsimai Yokoya, 1936: 129; Fig. 1. [tide pools at Aburatubo] 

Athanas parvus De Man, 1910a: 315. [8°39.1'S 127°4.4'E, anchorage south coast of Timor, 34 m] 

= Athanas Sibogae De Man, 1910a: 314. [Anchorage off Pulu Jedan, east coast of Aru-islands (pearlbanks), 

13 m] 

Athanas phyllocheles Banner & Banner, 1983: 152; Figs 13-14. [off Réunion, 450 m] 

Athanas polymorphus Kemp, 1915: 295; Figs 31-32. [outer channel off Satpara and Barhampur Island, 

Chilka Lake, 6-10 ft] 

Athanas rhothionastes Banner & Banner, 1960a: 142; Fig. 3. [Canton Island, Phoenix group] 

Athanas shawnsmithi Anker, 2011a: 2; Figs 1-2, 8A-D. [Australia, Queensland, Great Barrier Reef, Lizard 

Island, about 50 m east of Lumus Beach (in front of marine station), 14°40'92.6''S 145°26'95.2''E, less 

than 0.5 m] 

Athanas squillophilus Hayashi, 2002: 396; Figs 1-3. [Ishikari Bay, off Takashima Port, Otaru, Hokkaido, 10 

m, associated with Oratosquilla oratoria] 

Athanas stebbingii De Man, 1920a 

= Athanas Stebbingii De Man, 1920a: 106. [Sangkapoera-roads, Bawean-island, 12 m] 

Athanas sydneyensis Anker & Ahyong, 2007: 18; Figs 1-2. [mid-stream between Juno Head and Hungry 

Beach, Hawkesbry River, New South Wales, Australia, 33°34'S 151°16'E, 10 m] 

Athanas tenuipes De Man, 1910a: 316. [0°58.5'N 122°42.5'E, west of Kwandang-by-entrance, 72 m] 

Athanopsis Coutière, 1897f 

= Athanopsis Coutière, 1897f (type species Athanopsis platyrhynchus Coutière, 1897f, by monotypy, 


Athanopsis australis Banner & Banner, 1982a: 9; Fig. 1. [Beaumaris, Port Phillip Bay, Victoria, 8 m] 

Athanopsis brevirostris Banner & Banner, 1981a: 45; Fig. 5. [Cundabilu Island, Dahlak Archipelago, southern 

Red Sea, 0.3 m] 

Athanopsis dentipes Miya, 1980: 118; Figs 1-3. [in front of the Aquaculture Research Laboratory, Nagasaki 

Prefectural Institute of Fisheries, Nomo Bay, Nomozaki, Nagasaki Peninsula, LWS, in burrow of 

Thalassema ?mucosa] 

Athanopsis platyrhynchus Coutière, 1897f: 301. [Djibouti] 

Athanopsis rubricinctuta Berggren, 1991: 166; Figs 1-6. [Inhaca Island, Moçambique] 

Athanopsis saurus Anker, 2011a: 9; Figs 5-7, 8G-H. [Australia, Queensland, Great Barrier Reef, Lizard Island, 

Casuarina Beach near marine station, 14°40'31.4''S 145°26'39.5''E, 1 m] 

Automate De Man, 1888b 

= Arethusa De Man, 1887 (nomen nudum) 

= Automate De Man, 1888b (type species Automate dolichognatha De Man, 1888b, by monotypy, gender 


Automate anacanthopus De Man, 1910a: 317. [Kwadang-bay entrance, 0°58'.5N 122°55'E, 75 m; Sananabay, 

East coast of Sula Besi, 22 m]


Automate branchialis Holthuis & Gottlieb, 1958: 34; Figs 5-6. [off Gaza] 

Automate dolichognatha De Man, 1888b: 529; Plate 22, fig. 5-5i. [Insel Noordwachter] 

= Automate Gardineri Coutière, 1902b: 337. [Minikoi Atoll; Madhu Atoll; Goidu Atoll; Hulule Atoll] 

= Automate kingsleyi Hay, 1917: 72. [Beaufort, North Carolina] 

= Automate haightæ Boone, 1931: 184; Fig. 22. [north shore of Tabogilla Island, Pearl Islands, Bay of 

Panama] 

= Automate johnsoni Chace, 1955: 13; Fig. 7. [Bikini Atoll] 

Automate evermanni Rathbun, 1901: 112; Fig. 22. [off Aguadilla, 137 fms] 

Automate hayashii Anker & Komai, 2004: 1897; Figs 1-4. [tidal flat at Kamiiso, Hakodate Bay, southern 

Hokkaido, Japan, intertidal] 

Automate rectifrons Chace, 1972: 75; Fig. 24. [Antigua Island, north side of Bird Island, Nonsuch Bay, 1-4 ft] 

Automate rugosa Coutière, 1902b: 341. [8°44N 79°09W, baie de Panama, 30 fms] 

Automate salomoni Coutière, 1908 

= Automate Salomoni Coutière, 1908: 192. [Chagos, Salomon Island] 

Automate talismani Coutière, 1902b: 340. [Puerto-Grande, Açores, 20 m] 

Bannereus Bruce, 1988f 

= Bannereus Bruce, 1988f (type species Bannereus anomalus Bruce, 1988f, by original designation and 


Bannereus anomalus Bruce, 1988f: 140; Figs 1-6. [Marian Plateau, Coral Sea, 22°35.3'S 153°46.7'E to 

22°36.3'S 153°50.0'E, 345-350 m] 

Batella Holthuis, 1955b 

= Cheirothrix Spence Bate, 1888 (type species Cheirothrix parvimanus Spence Bate, 1888, by monotypy, 

gender feminine; invalid junior homonym of Cheirothrix Pictet & Humbert, 1866 (Pisces)) 

= Batella Holthuis, 1955b (nomen novum for Cheirothrix Spence Bate, 1888, gender feminine) 

Batella leptocarpus Chace, 1988: 65; Fig. 17. [Philippines, western Mindanao Sea, 8°47'15''N 123°35'00''E, 

296 m] 

Batella parvimanus (Spence Bate, 1888) 

= Cheirothrix parvimanus Spence Bate, 1888: 533; Plate 96, fig. 2-2b. [Challenger stn 186, 10°30'S 

142°18'E, off Cape York, 8 fms] 

= Batella bifurcata Miya & Miyake, 1968b: 116; Figs 2-4. [near Dajo Group, Nagasaki Prefecture, Japan, 

32°14.0'N 127°50.4'E, 156 m] 

Batella praecipua De Grave, 2004a: 504; Figs 1-3. [New Caledonia, 22°53'8"S 167°13'9"E, 425-440 m, dans 

une éponge avec des sténopodides] 

Bermudacaris Anker & Iliffe, 2000 

= Bermudacaris Anker & Iliffe, 2000 (type species Bermudacaris harti Anker & Iliffe, 2000, by original 


Bermudacaris australiensis Anker & Komai, 2004: 1903; Figs 5-7. [north-west shelf, Western Australia, 

19°54.0'S 117°52.2'E, 38 m] 

Bermudacaris britayevi Anker, Poddoubtchenko & Marin, 2006: 1677; Figs 1-4. Vietnam, Nha Trang Bay, 

Tre Island, Dam Bay, intertidal] 

Bermudacaris harti Anker & Iliffe, 2000: 762; Figs 1-4. [Bermuda Island, Christie’s Cave] 

Betaeopsis yaldwyn, 1971 

= Betaeopsis Yaldwyn, 1971 (type species Betæus æquimanus Dana, 1852a, by original designation, 


= Hamalpheus Bruce & Iliffe, 1991 (type species Hamalpheus acanthops Bruce & Iliffe, 1991, by original 


Betaeopsis acanthops (Bruce & Iliffe, 1991) 

= Hamalpheus acanthops Bruce & Iliffe, 1991: 584; Figs 1-5. [Tosua-Tolesua lave tube cave, near Lotofaga 

village, south coast of Upolu Island, Western Samoa]


Betaeopsis aequimanus (Dana, 1852a) 

= Betæus æquimanus Dana, 1852a: 23. [in portu ”Bay of Islands”, ad insulas ”Black Rocks”, Novi- 

Zealandiæ] 

Betaeopsis indica (De Man, 1910a) 

= Betaeus indicus De Man, 1910a: 309. [Anchorage off Labuan Pandan, Lombok, 18 m] 

Betaeus Dana, 1852a 

= Betaeus Dana, 1852a (type species Betæus truncatus Dana, 1852a, designated by Fowler, 1912, gender 

masculine; name placed on the Official List of Generic Names in Zoology in Opinion 470 in 

1957) 

Betaeus australis Stimpson, 1860a: 31. [Long Reef, Collaroy, New South Wales, intertidal rock platform; 

neotype selection by Banner & Banner, 1973a] 

Betaeus emarginatus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) (Fig. 45) 

= A.[lpheus] emarginatus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 357. [Type locality 

not indicated] 

= Alpheus lævigatus Guérin-Méneville, 1838 [in Guérin-Méneville, 1829-1838]: 38. [l’île de Timor] 

= Betæus scabro-digitus Dana, 1852a: 23. [juxta urbem ”Valparaiso” Chilensem] 

Betaeus ensenadensis Glassell, 1938: 416; Plate 28. [Estero de la Punta Banda, Ensenada, Baja California, 

Mexico, low tide] 

Betaeus gelasinifer Nomura & Komai, 2000: 45; Figs 1-5. [Kominato, Boso Peninsula, intertidal] 

Betaeus gracilis J.F.L. Hart, 1964: 453; Figs 50-51, 56, 65-67, 77-78. [Laguna Beach] 

Betaeus granulimanus Yokoya, 1927: 173; Plate 7, figs 17-22. [Bay of Kagoshima] 

= Betaeus yokoyai Kubo, 1936b: 50; Plate 15. [near low tide mark at Kominato] 

= Betaeus murayamai Yokoya, 1936: 132; Fig. 2. [vicinity of the Misaki Marine Biological Station] 

Betaeus harfordi (Kingsley, 1878b) 

= Alpheus harfordi Kingsley, 1878b: 198. [Santa Barbara and Catalina Island, California, under the 

mouth of Haliotis rufescens] 

= Betæus equimanus Lockington, 1877b: 43. [Catalina Island, under the mantle of Haliotis rufescens] 

= Alpheus lævigatus Nicolet, 1849: 215. [costas de Chile] 

= Alpheus æqualis Kingsley, 1878b: 199. [nomen novum for Betæus equimanus Lockington, 1877b] 

Betaeus harrimani Rathbun, 1904 

= Betæus harrimani Rathbun, 1904: 108; Fig. 49. [Sitka] 

Betaeus jucundus Barnard, 1947 

= Betæus jucundus Barnard, 1947: 388. [Keurbooms River estuary, Plettenberg Bay] 

Betaeus levifrons Vinogradov, 1950: 198; Plate 10, fig. 32. [Zolotoi Rog Bay, near Vladivostok region, Russian 

Far East] 

Betaeus lilianae Boschi, 1966: 84; Figs 1-3. [Quequén, provincia de Buenos Aires, entre algas en la playa] 

Betaeus longidactylus Lockington, 1877a: 35. [San Diego, sandy mud flat, between tides] 

Betaeus macginitieae J.F.L. Hart, 1964: 451; Figs 48-49, 55, 62-64, 75-76. [California, Corona del Mar, off 

Ladder Rock, 4 fms] 

Betaeus pingi Yu, 1930a 

= Betaeus Pingi Yu, 1930a: 454; Figs 1-3. [Chefoo?] 

Betaeus setosus J.F.L. Hart, 1964: 455; Figs 52-53, 57, 68-72, 79-80; Plate 2. [British Colombia, Vancouver 

Island, west coast, Clayoquot Sound] 

Betaeus truncatus Dana, 1852a 

= Betæus truncatus Dana, 1852a: 23. [in mari prope insulam ”Hermite”] 

= Alpheus Sinuosus Guérin-Méneville, 1857: 51. [Callao, au Chili] 

Bruceopsis Anker, 2010b 

= Bruceopsis Anker, 2010b (type species Bruceopsis projectus Anker, 2010b, by original designation, 


Bruceopsis projectus Anker, 2010b: 392; Figs 1A-B, 2-5. [Mariana Islands, Guam, Agat Bay north of Alutom 

Islet, under rocks, 3-7 m]


Fig. 45. Betaeus emarginatus (H. Milne Edwards, 1837). Photo by Arthur Anker. 

Bruceopsis guamensis Anker, 2010b: 399; Figs 1C, 6-8. [Marian Islands, Guam, Agat Bay north of Alutom 

Islet, under rocks, 3-7 m] 

Coronalpheus Wicksten, 1999 

= Coronalpheus Wicksten, 1999 (type species Coronalpheus natator Wicksten, 1999, by original designation 


Coronalpheus natator Wicksten, 1999: 106; Figs 1-4. [Devil’s Crown, Isla Onslow, north of Isla Floreana, 15 m] 

Coutieralpheus Anker & Felder, 2005 

= Coutieralpheus Anker & Felder, 2005 (type species Coutieralpheus setirostris Anker & Felder, 2005, 

by original designation and monotypy, gender masculine) 

Coutieralpheus setirostris Anker & Felder, 2005: 42; Figs 1-6. [south margin of Fort Pierce Inlet channel, 

beside US Highway A1A South Causeway, Indian River Lagoon, St. Lucie County, Florida, 27°27.7'N 

80°18.7'W, intertidal sand flat] 

Deioneus Dworschak, Anker & Abed-Navandi, 2000 

= Deioneus Dworschak, Anker & Abed-Navandi, 2000 (type species Deioneus sandizelli Dworschak, 

Anker & Abed-Navandi, 2000, by original designation and monotypy, gender masculine) 

Deioneus sandizelli Dworschak, Anker & Abed-Navandi, 2000: 304; Figs 1-26. [Cape Verde Islands, Ilha 

do São Tiago, NE-coast, tidal flat at Moia-Moia, with Corallianassa intesi] 

Fenneralpheus Felder & Manning, 1986 

= Fenneralpheus Felder & Manning, 1986 (type species Fenneralpheus chacei Felder & Manning, 1986, 


Fenneralpheus chacei Felder & Manning, 1986: 498; Figs 1-3. [Florida, St. Lucie County, Indian River lagoon, 

Fort Pierce Inlet, 27°27.7'N 80°18.7'W] 

Harperalpheus Felder & Anker, 2007 

= Harperalpheus Felder & Anker, 2007 (type species Harperalpheus pequegnatae Felder & Anker, 2007, 


Harperalpheus pequegnatae Felder & Anker, 2007: 457; Figs 1-4. [northwestern Gulf of Mexico, southeast 

of Galveston Island, Texas, USA, near 29°18.5'N 94°40.0'W, about 5 km offshore, 11.5-12 m]


Jengalpheops Anker & Dworschak, 2007a 

= Jengalpheops Anker & Dworschak, 2007a (type species Jengalpheops rufus Anker & Dworschak, 

2007a, by original designation and monotypy, gender masculine) 

Jengalpheops rufus Anker & Dworschak, 2007a: 291; Figs 1-7. [Philippines, Panglao Island, Sungcolan 

Bay, 9°38.3'N 123°49.6'E, tidal flat, from Glypturus armatus mound] 

Leptalpheus Williams, 1965 

= Leptalpheus Williams, 1965 (type species Leptalpheus forceps Williams, 1965, by original designation 


Leptalpheus axianassae Dworschak & Coelho, 1999: 477; Figs 1-30. [Praia do Araçá, São Sebastião, São 

Paulo, Brazil] 

Leptalpheus denticulatus Anker & Marin, 2009: 92; Figs 1-6, 7a-b, 8. [Vietnam, Nhatrang Bay, Dam Bay, 

intertidal sand flat, from burrow of Glypturus cf. armatus] 

Leptalpheus dworschaki Anker & Marin, 2009: 100; Figs 9-12. [Philippines, Panglao Island, Sungcolan 

Bay, 9°38.3'N 123°49.6'E, intertidal flat, from burrow of Glypturus sp.] 

Leptalpheus felderi Anker, Vera Caripe & Lira, 2006: 688; Figs 1-5, 6A-B. [Venezuela, Estado Nueva Esparta, 

Isla Margarita, southern shore near Guamache, 0.5-1 m, from burrow of Upogebia omissa] 

Leptalpheus forceps Williams, 1965: 194; Figs 1-2. [Gallants Point, Newport River, Carteret County, 

North Carolina] 

Leptalpheus mexicanus Ríos & Carvacho, 1983: 306; Figs 1-3. [estuary of Mulegé River, Baja California 

Sur, Mexico, 26°54'N 111°59'W] 

Leptalpheus pacificus Banner & Banner, 1975a: 424; Fig. 1. [reef in Kaneohe Bay, Oahu, 21°27'00''N 

157°47'16''W, low intertidal] 

Leptalpheus pierrenoeli Anker, 2008: 783; Figs 1-2, 3A-B. [Panama, Caribbean coast, Isla Grande, southern 

shore, village, near Cabañas Super Jackson, from burrow, 0.5-1.0 m] 

Fig. 46. Leslibetaeus coibita Anker, Poddoubtchenko & Wehrtmann, 2006. Photo by Arthur Anker.


Leptathanas De Grave & Anker, 2008 

= Leptathanas De Grave & Anker, 2008 (type species Leptathanas powelli De Grave & Anker, 2008, 


Leptathanas powelli De Grave & Anker, 2008: 45; Figs 1-5. [Nigeria, Niger Delta, Eagle Island, Bonny 

River system, from burrows of Upogebia furcata] 

Leslibetaeus Anker, Poddoubtchenko & Wehrtmann, 2006 

= Leslibetaeus Anker, Poddoubtchenko & Wehrtmann, 2006 (type species Leslibetaeus coibita Anker, 

Poddoubtchenko & Wehrtmann, 2006, by original designation and monotypy, gender masculine) 

Leslibetaeus caribbeaus Anker, 2011b: 64; Figs 1-2. [Trinidad & Tobago, Tobago Island, Sandy Bay, 1 m] 

Leslibetaeus coibita Anker, Poddoubtchenko & Wehrtmann, 2006: 30; Figs 1-5. [Isla Coibita, on side facing 

Isla Coiba, north side of STRI beach house, first small island connected by causeway, 7°38'80''N 

81°42'159''W, Isla Coiba National Park, Pacific Coast of Panama] (Fig. 46) 

Metabetaeus Borradaile, 1899 

= Metabetaeus Borradaile, 1899 (type species Betaeus minutus Whitelegge, 1897, by monotypy, gender 


Metabetaeus lohena Banner & Banner, 1960b: 299; Fig. 1. [Lohena pool, a few hundred yards from the 

shore between the deserted villages of Wai-o-ahu-kini and Ka’ili-k’I, immediately west of Lae 

(South Point), Hawaii] 

Metabetaeus mcphersonae Anker, 2010c: 40; Figs 1-7. [French Polynesia, Society Islands, Moorea, in front 

of Hilton hotel, 149.84247 W 17.47564 S, 13-17 m] 

Metabetaeus minutus (Whitelegge, 1897) 

= Betaeus minutus Whitelegge, 1897: 147; Plate 7, figs 4-4b. [Funafuti, under stones and in sponges 

in the mangrove swamp] 

Metalpheus Coutière, 1908 

= Metalpheus Coutière, 1908 (type species Alpheus rostratipes Pocock, 1890, designated by Shelford, 


Metalpheus hawaiiensis (Edmondson, 1925) 

= Crangon hawaiiensis Edmondson, 1925: 14; Fig. 2. [Lisiansky Island, shallow water] 

Metalpheus paragracilis (Coutière, 1897d) 

= Alpheus paragracilis Coutière, 1897d: 304. [Ile Tague] 

? = Alpheus aglaopheniae Borradaile, 1900: 417; Plate 38, figs 13a-f. [Engineer Group, British New 

Guinea] 

Metalpheus rostratipes (Pocock, 1890) 

= Alpheus rostratipes Pocock, 1890: 522. [Fernando de Noronha] 

= Alpheus sp.?, Metalpheus n. gen. Coutière, 1908: 213. 

= Crangon hawaiiensis clippertoni Schmitt, 1939: 11. [Clipperton Island, on rocks to south of landing 

place, on north-eastern shore of atoll, shore collecting] 

= Crangon nanus Banner, 1953: 90; Figs 30-31; nec Krøyer, 1843. [Hanauma Bay, Oahu, 10ft] 

= Alpheus huikau Banner, 1959: 139; Fig. 5. [nomen novum for Crangon nanus Banner, 1953] 

Mohocaris Holthuis, 1973c 

= Mohocaris Holthuis, 1973c (type species Mohocaris bayeri Holthuis, 1973c, by original designation 


Mohocaris bayeri Holthuis, 1973c: 490; Figs 1-2. [Off French Guiana, 6°58'N 53°10'W to 7°01'N 53°15'W, 

127-131 m] 

Nennalpheus Banner & Banner, 1981b 

= Nennalpheus Banner & Banner, 1981b (type species Alpheopsis Sibogae De Man, 1910a, by original 



Nennalpheus inarticulatus Banner & Banner, 1981b: 221; Fig. 1. [13°59.2'N 120°20.3'E, 208-222 m] 

Nennalpheus sibogae (De Man, 1910a) 

= Alpheopsis Sibogae De Man, 1910a: 307. [8°23.5'S 119°4.6'E, Sapeh-strait, 70 m] 

Notalpheus Méndez G. & Wicksten, 1982 

= Notalpheus Méndez G. & Wicksten, 1982 (type species Notalpheus imarpe Méndez G. & Wicksten, 


Notalpheus imarpe Méndez G. & Wicksten, 1982: 709; Figs 1-2. [off Paita, Peru, 5°03'S 81°20'W, 143 m] 

Orygmalpheus De Grave & Anker, 2000 

= Orygmalpheus De Grave & Anker, 2000 (type species Orygmalpheus polites De Grave & Anker, 2000, 


Orygmalpheus polites De Grave & Anker, 2000: 250; Figs 1-4. [Liang Island Lagoon, about 4°10'30"S 

144°52'47"E, Hansa Bay, Madang Province, Papua New Guinea, 10 m] 

Parabetaeus Coutière, 1897b 

= Parabetæus Coutière, 1897b (type species Parabetæus Culliereti Coutière, 1897b, by monotypy, gender 


= Neoalpheopsis Banner, 1953 (type species Neoalpheopsis hiatti Banner, 1953, by original designation, 


Parabetaeus culliereti Coutière, 1897b 

= Parabetaeus Culliereti Coutière, 1897b: 383. [Papeete] 

Parabetaeus euryone (De Man, 1910a) 

= Alpheopsis? Euryone De Man, 1910a: 308. [anchorage off Kawio- and Kamboling-islands, Karkarolong-group, 

reef] 

= Neoalpheopsis hiatti Banner, 1953: 21; Fig. 6. [Hanauma Bay, Oahu, 20ft] 

Parabetaeus hummelincki (Schmitt, 1936) 

= Alpheopsis hummelincki Schmitt, 1936: 364; Plate 11, figs 1a-h. [Kralendijk (de Hoop) Island, Bonaire, 

about 3ft] 

Pomagnathus Chace, 1937b 

= Pomagnathus Chace, 1937b (type species Pomagnathus corallinus Chace, 1937b, by original designation 


Pomagnathus corallinus Chace, 1937b: 124; Fig. 5. [off Arena Bank, 2.5 fms in coral] 

Potamalpheops Powell, 1979 

= Potamalpheops Powell, 1979 (type species Potamalpheops pylorus Powell, 1979, by original designation, 


Potamalpheops amnicus Yeo & Ng, 1997: 165; Figs 1-2. [1°51'46.5''N 104°00'31.0''E, Sungei Selangi, Kota 

Tinggi, Johore] 

Potamalpheops darwiniensis Bruce, 1993c: 698; Figs 1-3. [Hudson’s Creek, Darwin Harbour, Northern Territory, 

Australia, 12°28.75'S 130°55.67'E, intertidal] 

Potamalpheops galle Anker, 2005: 32; Fig. 1. [SW Sri Lanka, Lelkada near Dodangoda, Galle District, Gib 

River basin, 6 km from the sea, altitude 5 m, 06°08'N 80°10'E, 0.5-1 m] 

Potamalpheops hanleyi Bruce, 1991d: 629; Figs 1-4. [Creek H, East Arm, Darwin Harbor, Northern Territory, 

12°32.6'S 130°56.6'E, low water spring tide] 

Potamalpheops haugi (Coutière, 1906) 

= Alpheopsis Haugi Coutière, 1906: 377, 378; Figs 1-2. [Ngômô, petit lac d’eau douce des bords de 

l’Ogoué, Gabon] 

Potamalpheops johnsoni Anker, 2003a: 290; Figs 5-7, 20c-d. [Sungei Buloh mangrove forest, Singapore] 

Potamalpheops miyai Yeo & Ng, 1997: 175; Figs 3-4. [Sungei Lagoi, bridge leading to Kampung Lagoi, 

downstream of reservoir outlet, Pulau Bintan, Indonesia]


Potamalpheops monodi (Sollaud, 1932) 

= Alpheopsis monodi Sollaud, 1932: 377; Figs 1-2. [Cameroun] 

Potamalpheops palawanensis Cai & Anker, 2004: 259; Figs 15-16. [St Paul Cave subterranean river, near 

cave entrance, Palawan] 

Potamalpheops pininsulae Bruce & Iliffe, 1992: 231; Figs 1-53. [Grotte de la Troisième (Paradise Cave), Ile 

de Pins, New Caledonia, 0-6 m] 

Potamalpheops pylorus Powell, 1979: 120; Figs 1, 3. [Nigeria, mangrove creek at southwest corner of College 

of Science and Technology, Port Harcourt, approx. 4°47'15''N 6°58'45''E] 

Potamalpheops stygicola (H.H.Jr. Hobbs, 1973a) 

= Alpheopsis stygicola H.H.Jr. Hobbs, 1973a: 73; Figs 1-2. [Cueva del Nacimineto del Río San Antonio, 

10 km SSW Acatlán, Oaxaca, México] 

Potamalpheops tigger Yeo & Ng, 1997: 182; Figs 5-6. [Sungi Buloh mangrove, Singapore] 

Prionalpheus Banner & Banner, 1960c 

= Prionalpheus Banner & Banner, 1960c (type species Prionalpheus triarticulatus Banner & Banner, 

1960c, by original designation and monotypy, gender masculine) 

Prionalpheus brachytomeus Banner & Banner, 1971: 267. [outer edge of fringing reef at Korolevu, Vitilevu, 

Fiji] 

Prionalpheus fissipes Coutière, 1908 

= Alpheopsis fissipes Coutière, 1908: 193. [Providence, 50-78 brasses] 

Prionalpheus gomezi Martínez-Iglesias & Carvacho, 1991: 85; Figs 1-12. [récif Juan García, Golfe de Batabanó, 

SW de Cuba, 10 m] 

Prionalpheus mortoni Bruce, 1990f: 647; Fig. 18. [Tolo Channel, 5 m; full description as Prionalpheus sp. aff. 

triarticulatus Bruce, 1990f: 628; Figs 10-12] 

Prionalpheus nayaritae Alvarez, Camacho & Villalobos, 1996: 719; Fig. 4. [Punta de Mita, Nayarit, 20°46'N 

105°31'W] 

Prionalpheus sulu Banner & Banner, 1971: 268; Fig. 2. [eastern end of Big Santa Cruz Island, Zamboanga, 

Mindanao, 10ft] 

Prionalpheus triarticulatus Banner & Banner, 1960c: 293; Fig. 1. [Korolevu, Vitilevu, Fiji, reef flat on fringing 

reef] 

Pseudalpheopsis Anker, 2007b 

= Pseudalpheopsis Anker, 2007b (type species Pseudalpheopsis guana Anker, 2007b, by original designation 


Pseudalpheopsis guana Anker, 2007b: 430; Figs 1-8. [Caribbean Sea, British Virgin Islands, Guana Island, 

North Bay, North Beach, 9 m] 

Pseudathanas Bruce, 1983d 

= Pseudathanas Bruce, 1983d (type species Pseudathanas darwiniensis Bruce, 1983d, by original designation 


Pseudathanas darwiniensis Bruce, 1983d: 464; Figs 1-5. [Dudley Point Reef, East Point, Darwin, Australia, 

12°25.0'S 30°49.1'E, LWS] 

Pterocaris Heller, 1862b 

= Pterocaris Heller, 1862b (type species Pterocaris typica Heller, 1862b, by monotypy, gender feminine) 

Pterocaris typica Heller, 1862b: 398; Plate 1, figs 7-18. [Amboina] 

Racilius Paul’son, 1875 

= Racilius Paul’son, 1875 (type species R.[acilius] compressus Paul’son, 1875, by monotypy, gender 


Racilius compressus Paul’son, 1875 

= R.[acilius] compressus Paul’son, 1875: 107; Plate 14, fig. 2. [Red Sea]


Richalpheus Anker & Jeng, 2006 

= Richalpheus Anker & Jeng, 2006 (type species Richalpheus palmeri Anker & Jeng, 2006, by original 


Richalpheus dahabensis Anker & Dworschak, 2007b: 2333; Figs 1-3. [Egypt, Red Sea, Dahab, Laguna, 13 m, 

from mound of Glypturus sp.] 

Richalpheus palmeri Anker & Jeng, 2006: 381; Figs 1-6. [Philippines, Panglao Island, Pontod Islet lagoon, 

9°33.1'N 123°44.0'E, 3-4 m] 

Rugathanas Anker & Jeng, 2007 

= Rugathanas Anker & Jeng, 2007 (type species Athanas verrucosus Banner & Banner, 1960a, by original 


Rugathanas borradailei (Coutière, 1903) 

= Arete Borradailei Coutière, 1903: 80; Figs 19-24. [Hulule Male Atoll] 

= Arete Borradailei Coutière, 1905b: 861; Fig. 133. [Hulule Male Atoll] 

= Arete ghardaqensis Ramadan, 1936: 15; Plate 1, figs 2-8. [Ghardaga, Red Sea] 

= Athanas polynesia Banner & Banner, 1966b: 152; Fig. 7. [Alofau, Tuttuila, 3-10ft] 

Rugathanas verrucosus (Banner & Banner, 1960a) 

= Athanas verrucosus Banner & Banner, 1960a: 147; Fig. 4. [ocean (windward) reef, Parry Island, Eniwetok] 

Salmoneus Holthuis, 1955b 

= Jousseaumea Coutière, 1897b (type species Jousseaumea serratidigitus Coutière, 1897b, designated by 

Holthuis, 1955b, gender feminine; invalid junior homonym of Jousseaumia Sacco, 1894 (Mollusca); 

name placed on the Official Index of Rejected and Invalid Names in Zoology in Opinion 673 in 

1963) 

= Salmoneus Holthuis, 1955b (nomen novum for Jousseaumea Coutière, 1897b, gender masculine; 


Salmoneus alpheophilus Anker & Marin, 2006: 310; Figs 12-13. [South China Sea, Vietnam, Nhatrang Bay, 

Tre Island, Dam Bay, 12°10'20''N 109°15'30''E, low tide, in burrow of goby-associated Alpheus sp.] 

Salmoneus armatus Anker, 2010d: Figs 3-4, 14a. [Panama, Caribbean coast, Isla Grande, southern shore, 

100 m east of western-most point, under rock on muddy sand, 0.5 m depth] 

Salmoneus arubae (Schmitt, 1936) 

= Jousseaumea arubae Schmitt, 1936: 366; Plate 12, Figs 2a-g. [Aruba, Punta Braboe (Oranjestad), 

Schorrenfläche, about 1 ft] 

Salmoneus auroculatus Anker & Marin, 2006: 300; Figs 4-6, 7a. [South China Sea, Vietnam, Nhatrang Bay, 

Tre Island, 6-8 m] 

Salmoneus babai Miyake & Miya, 1966: 133; Fig. 1. [Suno coral reef, Kasari, Amami-oshima] 

Salmoneus brevirostris (Edmondson, 1930) 

= Jousseaumea brevirostris Edmondson, 1930: 7; Fig. 3. [Waikiki reef, Oahu, in shallow water] 

Salmoneus brucei Komai, 2009: 870; Figs 1-5. [sand flat in Uehara Port, Iriomote Island, Yaeyama Islands, 

southern Rykyu Islands, intertidal burrow of Lepidophthalmus tridentatus] 

Salmoneus bruni Banner & Banner, 1966a: 42; Fig. 11. [Gulf of Thailand, 28 m] 

Salmoneus caboverdensis Dworschak, Anker & Abed-Navandi, 2000: 309; Figs 27-41. [Cape Verde Islands, 

Ilha do São Tiago, NE-coast, tidal flat at Moia-Moia, with Neocallichirus pachydactylus] 

Salmoneus camaroncito Anker, 2010d: 184; Figs 5-6, 14b. [Panama, Caribbean coast, Isla Grande, western 

point, under rock on coarse sand mixed with rubble, 0.5 m depth] 

Salmoneus carvachoi Anker, 2007c: 27; Fig. 3. [Guadeloupe, estuary of Rivière Lézarde, dredge between 

estuary and canal] 

Salmoneus cavicolus Felder & Manning, 1986: 503; Figs 4-6. [Florida, St. Lucie County, Indian River lagoon, 

south side of Fort Pierce Inlet, 27°27.7'N 80°18.7'W] 

Salmoneus colinorum De Grave, 2004b: 41; Figs 1-4. [Ngerbeched mangrove, SW of Malakal Harbour, 

Palau, 07°19.703'N 134°28.611'E]


Salmoneus cristatus (Coutière, 1897c) 

= Jousseaumea cristata Coutière, 1897c: 234. [Djibouti] 

Salmoneus degravei Anker, 2010d: 188; Figs 7-9, 14c. [Panama, Caribbean coast, Isla Grande, southern 

shore, about 100 m east of western point and Hotel Isla Grande, from burrow of unknown host, 

0.5-1 m depth] (Fig. 47) 

Salmoneus erasimorum Dworschak, Anker & Abed-Navandi, 2000: 313; Figs 42-56. [Adria, Kvarner, 

Opatja, ”Hotel Beach”, ca. 1.5 m, with Callianassa tyrrhena] 

Salmoneus falcidactylus Anker & Marin, 2006: 304; Figs 8-9. [South China Sea, Vietnam, Nhatrang Bay, 

about 2 km SW off Mung Island, 40 m] 

Salmoneus gracilipes Miya, 1972: 38; Plate 3. [Amakusa Islands, Shino-shima Island, Tomioka, low tide mark] 

Salmoneus hilarulus (De Man, 1910a) 

= Jousseaumea hilarula De Man, 1910a: 304. [1°42.5'S 130°47.5'E, 32 m] 

Salmoneus hispaniolensis Anker, 2010d: 178; Figs 1-2. [Dominican Republic, Boca Chica, sandy lagoon 

with artificial barrier in front of hotel, 1 m] 

Salmoneus jarli (Holthuis, 1951a) 

= Jousseaumea jarli Holthuis, 1951a: 94; Fig. 20. [off Nigeria, 4°12'N 7°05'E, 19 m] 

Salmoneus kekovae Grippa, 2004: 45; Figs 1-2. [Kekova, southwest coast of Turkey] 

Salmoneus komaii Anker, 2011c: 73; Figs 5-7. [Mariana Islands, Guam, west coast of Guam, south of Orote 

Point, Barracuda Rock, less than 10 m] 

Salmoneus latirostris (Coutière, 1897b) 

= Jousseaumea latirostris Coutière, 1897b: 382. [Mer Rouge] 

Salmoneus mauiensis (Edmondson, 1930) 

= Jousseaumea mauiensis Edmondson, 1930: 5; Fig. 2. [island of Maui, shallow water] 

Salmoneus nhatrangensis Anker & Marin, 2006: 296; Figs 1-3. [South China Sea, Nhatrang Bay, Tre Island, 

8-10 m] 

Salmoneus ortmanni (Rankin, 1898) 

= Athanas ortmanni Rankin, 1898: 251; Plate 30, fig. 7. [near Nassau, New Providence, along shore] 

Salmoneus paulayi Anker, 2011c: 68; Figs 1-4. [Mariana Islands, Guam, west coast of Guam, north of Apra 

Harbor, between Piti Channel and Cabras, reef flat, 0.5-1.5 m] 

Salmoneus poupini Anker, 2011c: 78; Figs 8-11. [French Polynesia, Society Islands, Moorea, Moorea lagoon, 

public beach between Temae point and Sofitel, 0.5-2 m] 

Salmoneus pusillus Anker & Marin, 2006: 307; Figs 10-11. [South China Sea, Vietnam, Nhatrang Bay, 

Pyramides, 14-16 m] 

Fig. 47. Salmoneus degravei Anker, 2010. Photo by Arthur Anker.


Salmoneus rocas Anker, 2007c: 29; Fig. 4. [Brazil, Atol das rocas, east of Laguna Interna, in calcareous algae, 

1 m] 

Salmoneus rostratus Barnard, 1962: 240; Fig. 1. [Nosi Bé, U. W.; lectotype designation by De Grave & 

Wilkins, 1997] 

Salmoneus serratidigitus (Coutière, 1897b) 

= Jousseaumea serratidigitus Coutière, 1897b: 382. [Mer Rouge] 

Salmoneus seticheles Anker, 2003b: 102; Figs 1-5. [mouth of Ludmilla Creek, Darwin, 12°24.8'S 130°51.7'E, 

LWS] 

Salmoneus setosus Manning & Chace, 1990: 17; Fig. 9. [South West Bay, Ascension Island] 

Salmoneus sibogae De Man, 1910a 

= Jousseaumea Sibogae De Man, 1910a: 303. [5700 M. N. 279°E from south point of South-Luciparaisland, 

reef-exploration] 

Salmoneus singaporensis Anker, 2003a: 284; Figs 1-2, 3a-b. [Tanjong Penuru, Jurong, Singapore, mudflats] 

Salmoneus sketi Fransen, 1991a: 171; Figs 1-29. [underwater cave, off the N coast of the small island Laernaka, 

Kornati Archipelago, Middle Dalmatia, Adriatic] 

Salmoneus tafaongae Banner & Banner, 1966b: 155. [3 miles east of Apia, Upolu, Western Samoa, 1ft below 

low water] 

Salmoneus teres Manning & Chace, 1990: 20; Fig. 10. [South West Bay, Ascension Island] 

Salmoneus tricristatus Banner, 1959 

= Salmoneus tricristata Banner, 1959: 131; Fig. 1. [Tomil Harbor, Yap island, Caroline Archipelago] 

Salmoneus wehrtmanni Anker, 2010d: 193; Figs 10-11, 12a, 13b, 14d. [Panama, Caribbean coast, Isla 

Grande, southern shore, under rocks and rubble on sand, near seagrass, 0.5-1 m depth] 

Stenalpheops Miya, 1997 

= Stenalpheops Miya, 1997 (type species Stenalpheops anacanthus Miya, 1997, by original designation 


= Cavipelta Hayashi, 1998 (type species Cavipelta yamashitai Hayashi, 1998 (junior subjective synonym 

of Stenalpheops anacanthus Miya, 1997), by original designation and monotypy, gender feminine) 

= Chelomalpheus W. Kim, 1998 (type species Chelomalpheus koreanus W. Kim, 1998 (junior subjective 

synonym of of Stenalpheops anacanthus Miya, 1997), by original designation and monotypy, gender 


Stenalpheops anacanthus Miya, 1997: 149; Figs 1A, 2-5. [Seto Inland Sea, Hiroshima Prefecture, Miyajima 

Island, intertidal zone, found in a burrow of Upogebia (Upogebia) major] 

= Chelomalpheus koreanus W. Kim, 1998: 141; Figs 1-3. [Daecheon, the Yellow Sea, Korea, 36°19'05''N 

126°30'25''E, from pool dug in mud-sand beach] 

= Cavipelta yamashitai Hayashi, 1998: 231; Figs 1-4. [Seto Inland Sea, tidal flat off Miyajima, Horishima 

Prefecture, living in tube of Upogebia major] 

Stenalpheops crangonus (Anker, Jeng & Chan, 2001) 

= Chelomalpheus crangonus Anker, Jeng & Chan, 2001: 1053; Figs 4-8. [Chung-Hua County, W Taiwan, 

mudflats, collected together with Upogebia edulis] 

Synalpheus Spence Bate, 1888 

= Homaralphæus Spence Bate, 1876 (nomen nudum) 

= Homaralpheus Spence Bate, 1888 (type species A.[lpheus] minus Say, 1818, designated by Holthuis, 

1955b, gender masculine) 

= Synalpheus Spence Bate, 1888 (type species Synalpheus falcatus Spence Bate, 1888 (junior subjective 

synonym of Alpheus Comatularum Haswell, 1882), by monotypy, gender masculine) 

= Alpheinus Borradaile, 1900 (type species Alpheinus tridens Borradaile, 1900 (junior subjective synonym 

of Alpheus Stimpsonii De Man, 1888b), gender masculine) 

= Zuzalpheus Ríos & Duffy, 2007 (type species Zuzalpheus kensleyi Ríos & Duffy, 2007, by original 



Synalpheus albatrossi Coutière, 1909: 89; Fig. 54. [Laysan Island, 10-19 fms] 

Synalpheus agelas Pequegnat & Heard, 1979: 110; Figs 1-4. [27°52'N 93°48'W, West Flower Garden Bank, 

Gulf of Mexico, 25 m] 

Synalpheus amabilis De Man, 1910a: 295. [Banda, 9-36 m] 

Synalpheus anasimus Chace, 1972: 82; Figs 25-28. [Bahía de l’Ascensión, Punta Solimán to 300 m southwest, 

5ft] 

Synalpheus anceps Banner, 1956: 334; Fig. 8. [Saipan] 

Synalpheus ancistrorhynchus De Man, 1909a: 124. [Anchorage off Pulu Jedan, east cost of Aru-islands 

(pearl-banks), 13 m] 

Synalpheus androsi Coutière, 1909: 82; Fig. 50. [Andros Island, Bahamas] 

Synalpheus antenor De Man, 1910a 

= Synalpheus Antenor De Man, 1910a: 293. [0°7'.2N 130°25'.5E, 83-59 m; 1°42'.5S 130°47'.5E, 32 m; 

Banda-anchorage, 9-36 m] 

Synalpheus apioceros apioceros Coutière, 1909 

= Synalpheus apioceros Coutière, 1909: 27; Fig. 9. [Marco, Florida] 

Synalpheus apioceros desterroensis Coutière, 1909: 31; Fig. 13. [Desterro] 

Synalpheus apioceros leiopes Coutière, 1909: 30; Fig. 12. [Venezuela] 

Synalpheus apioceros mayaguensis Coutière, 1909: 30; Fig. 11. [Mayaguez, Porto Rico, on coral reef] 

Synalpheus arostris Wicksten, 1989b: 78; Fig. 1. [Morro de Los Agujeros, Bahía de Malaga, Colombia, 

3°55'N 77°20'W, rocky intertidal zone] 

Synalpheus bannerorum Abele, 1975b: 79; Fig. 32. [southeastern side of Malpelo Island, Colombia, 10 m] 

Synalpheus barahonensis Armstrong, 1949: 20; Fig. 7. [behind Piedra Prieta Reef, Dominican Republic, 

from Agaricia agaricites] 

Synalpheus belizensis Anker & Tóth, 2008: 19; Figs 12-13, 14I. [Belize, Carrie Bow Cay, approx. 12 m, in 

cryptic sponge] 

Synalpheus bispinosus De Man, 1910a: 302. [Bay of Pidjot, Lombok, 9-22 m; Banda-anchorage, 9-36 m] 

Synalpheus bituberculatus De Man, 1910a: 294. [Haingsisi, Samau Island, Timor, 36 m; Lumu-lumu-shoal, 

Borneo-bank, reef, 1°42'.5S 130°47'.5E, 32 m; Anchorage off Pulu Jedan, east coast of Aru-islands 

(Pearl-banks), 13 m; 8°25'.2S 127°18'.4E, anchorage between Nusa Besi and the N.E. point of Timor, 

27-54 m; Anchorage East of Sailus Besar, Paternoster islands, up to 36 m] 

Synalpheus biunguiculatus (Stimpson, 1860a) 

= Alpheus biunguiculatus Stimpson, 1860a: 31. [ad insulas Hawaiensis, inter madreporas] 

Synalpheus bocas Anker & Tóth, 2008: 17; Figs 10-11, 14E-H. [Panama, Caribbean coast, Bocas del Toro, 

STRI Bay, in Xestospongia rosariensis, 5-15 m] 

Synalpheus bousfieldi Chace, 1972: 86; Figs 29-30. [Virgin Gorda, Colquhoun Reef at entrance to North 

Sound] 

Synalpheus bradleyi Verrill, 1922: 108; Plate 33, fig. 1. [Pearl Islands, Bay of Panama] 

Synalpheus brevicarpus (Herrick, 1891) 

= Alpheus saulcyi var. brevicarpus Herrick, 1891: 384 (?partim); Plate 4, figs 1-3; Plate. 21, figs 1-4, 8, 9; 

Plate 22, figs 1, 2, 4-10, 12-16; Plate 23, figs 1-8; Plate 24, figs 1, 3. [Bahamas, assumed to be Nassau, 

New Providence] 

= Synalpheus brevicarpus guerini Coutière, 1909: 52; Fig. 30. [Key West] 

Synalpheus brevidactylus Anker & Tóth, 2008: 14; Figs 8-9, 14C-D. [Panama, Caribbean coast, Isla Grande, 

between village and west point, in cryptic sponge, 1-3 m] 

Synalpheus brevifrons Chace, 1972: 89; Figs 31-32. [Dominica, north end of Woodbridge Bay] 

Synalpheus brevispinis Coutière, 1909 

= Synalpheus townsendi brevispinis Coutière, 1909: 34; Fig. 16. [Lower California] 

Synalpheus brooksi Coutière, 1909: 69; Fig. 41. [Sugar Loaf Key, Florida] 

= Synalpheus brooksi strepsiceros Coutière, 1909: 73; Fig. 42. [St. Thomas] 

= Synalpheus brooksi eleutheræ Coutière, 1909: 73; Fig. 43. [The Current, Eleuthera Island, Bahamas] 

Synalpheus carinatus (De Man, 1888b) 

= Alpheus carinatus De Man, 1888b: 508; Plate 22, fig. 2-2c. [Amboina]


= Synalpheus carinatus var. binongcensis De Man, 1909a: 111. [Anchorage off Pasir Pandjang, west 

coast of Binongka, 278 m] 

= Synalpheus carinatus var. ubianensis De Man, 1909a: 111. [6°7'5.N 120°26'E, anchorage off North- 

Ubian, 16-23 m; Anchorage off Pasir Pandjang, west coast of Binongka, reef] 

Synalpheus carpenteri MacDonald & Duffy, 2006: 2; Figs 1-7. [Long Reef, Belize, from canals of Agelas 

clathrodes] 

Synalpheus chacei Duffy, 1998: 389; Figs 1-5. [Carrie Bow Cay, Belize, from Agelas dispar, 3 m] 

Synalpheus charon (Heller, 1861) 

= A.[lpheus] charon Heller, 1861: 25. [im rothen Meere; fully described in Heller, 1862c] 

= Alpheus prolificus Spence Bate, 1888: 556; Plate 99, fig. 4. [off Honolulu, Sandwich Islands, 18 fms] 

= Synalpheus Helleri De Man, 1911d: 246. [Nicobar Islands] 

= Synalpheus charon obscurus Banner, 1956: 329; Fig. 5. [reef flat on the SE side of Unai Obyan, Saipan] 

Synalpheus comatularum (Haswell, 1882) 

= Alpheus Comatularum Haswell, 1882: 762. [Albany Passage, near Cape York, Queensland, few fms] 

= Synalpheus falcatus Spence Bate, 1888: 574; Plate 103. [Challenger stn 186, 10°30'S 142°18'E, between 

Cape York and the Arrou Islands, 8 fms] 

Synalpheus corallinus MacDonald, Hultgren & Duffy, 2009: 16; Figs 5-9; Plates 2D, 3A. [Jamaica, Dairy 

Bull Reef, 18°28.083'N 77°23.289'W, from canals of Hyattella intestinalis] 

Synalpheus coutierei Banner, 1953: 36. [nomen novum for Synalpheus biunguiculatus sensu Coutière, 1898e 

nec Stimpson, 1860a] 

? = Synalpheus biunguiculatus var. exilipes Coutière, 1905b: 874; Plate 71, figs 10-10a’’. [Haddumati 

Atoll; North Male Atoll; South Nilandu Atoll] 

Synalpheus cretoculatus Banner & Banner, 1979: 30; Fig. 2. [10°37'N 97°34'E, Andaman Sea, off southern 

tip of Burma, 75-80 m] 

Synalpheus crosnieri Banner & Banner, 1983: 92; Fig. 11. [northwest coast of Madagascar, 12°44'S 48°25.2'E, 

73 m] 

Synalpheus curacaoensis Schmitt, 1924a 

= Synalpheus curaçaoensis Schmitt, 1924a: 66; Fig. 3. [Spanish Bay, between stones in the surf] 

Synalpheus dardeaui (Ríos & Duffy, 2007) 

= Zuzalpheus dardeaui Ríos & Duffy, 2007: 18; Figs 3-6; Plate 2. [Twin Cays, Belize, from Lissodendoryx 

colombiensis, 1 m] 

Synalpheus demani Borradaile, 1900: 416. [Lifu, Loyalty Islands] 

= Alpheus triunguiculatus De Man, 1888b: 504; Plate 22, fig. 1-1e; nec Paul’son, 1875. [Amboina] 

= Synalpheus Brockii Nobili, 1901c: 2. [Amboina] 

Synalpheus digueti Coutière, 1909: 48; Fig. 28a-a’’, c-c’, k’-m. [Lower California] 

? = Alpheus læviusculus Lockington, 1878b: 474. [Port Escondido, Mulege Bay and other points on the 

Californian shore of the Gulf of California] 

= Synalpheus digueti ecuadorensis Coutière, 1909: 50; Fig. 28aa, cc. [St.Helena, Ecuador] 

Synalpheus disparodigitus Armstrong, 1949: 17; Fig. 6. [quiet water behind Piedra Pietra Reef, Dominican 

Republic] 

Synalpheus dominicensis Armstrong, 1949: 23; Fig. 8. [quit water inside Piedra Prieta Reef] 

Synalpheus dorae Bruce, 1988g: 843; Figs 1-6. [19°05.0'S 118°57.8'E, 82 m] 

Synalpheus duffyi Anker & Tóth, 2008: 8; Figs 4-5, 14A, 15A-B. [Panama, Caribbean coast, Isla Grande, 

southern shore facing La Guaira, between village and west point, in cryptic sponge, 1-1.5 m] 

Synalpheus echinus Banner & Banner, 1975c: 374; Fig. 27. [Trigg Island, near Perth, Western Australia] 

Synalpheus elizabethae (Ríos & Duffy, 2007) 

= Zuzalpheus elizabethae Ríos & Duffy, 2007: 23; Figs 7-10; Plate 2. [The Pinnacles (Sand Bores), SW of 

Carrie Bow Cay, Belize, in Hymeniacidion caerulae, 2 m] 

Synalpheus filidigitus Armstrong, 1949: 15; Fig. 5. [halfway between the Piedra Prieta Reef of Barahona 

Harbour and the shore, Dominican Republic] 

Synalpheus fossor (Paul’son, 1875) 

= Alph.[eus] fossor Paul’son, 1875: 103; Plate 13, figs 5-5g. [Red Sea]


= Synalpheus trionyx Coutière, 1908: 196. [Saya de Malha, 26 + 29 fms] 

= Synalpheus Bakeri Coutière, 1908: 199. [South-Adelaïde] 

= Synalpheus fossor var. propinqua De Man, 1909a: 121. [South-east side of Pearl-bank, Sulu-archipelago, 

15 m; 1°42'.5S 130°47'.5E, 32 m; Anchorage off Pulu Jedan, East coast of Aru-islands (Pearlbanks), 

13 m; 8°25'2.S 127°18'.4E, anchorage between Nusa Besi and the N.E. point of Timor, 27-54 

m; Anchorage East of Sailus Besar, Paternoster-islands, up to 36 m] 

= Synalpheus Bakeri var. Stormi De Man, 1911d: 253. [Balikpapan, east coast of Borneo; Atjeh] 

Synalpheus fritzmuelleri Coutière, 1909 

= Synalpheus fritzmülleri Coutière, 1909: 35; Fig. 18. [Marco, Florida] 

= Synalpheus fritzmülleri elongatus Coutière, 1909: 37; Fig. 19. [South Carolina, Mouth of Bull Creek] 

= Synalpheus fritzmuelleri var. caribæa Verrill, 1922: 97; Fig. 8; Plate 39, figs 3a-c. [Dominica Island] 

= Synalpheus fritzmuelleri var. carolinensis Verrill, 1922: 99; Plate 22, fig. 6; Plate 39, figs 1-1d. [Fort 

Macon, North Carolina] 

Synalpheus gambarelloides (Nardo, 1847) 

= Alpheus gambarelloides Nardo, 1847: 6. [trovato di rado nella valle Gambarelli vicino Chioggia] 

= Cancer glaber var. Olivi, 1792: 51; Plate 3, fig. 5; nec Cancer glaber Olivi, 1792. [la Golfo e delle Lagune 

di Venezia] 

= Alphaeus spincicerus Hope, 1851: 43. [mari neapolitano] 

= Alpheus laevimanus Heller, 1862b: 401; Plate 1, figs 25-27. [Sicilien, sowie von Pirano aus dem adriatischen 

Meere] 

Synalpheus goodei Coutière, 1909: 58; Fig. 33. [Bermuda] 

Synalpheus gracilirostris De Man, 1910a: 291. [8°25.2'S 127°18.4'E, anchorage between Nusa Besi and the 

N.E. point of Timor, 27-54 m] 

Synalpheus haddoni Coutière, 1900c 

= Synalpheus laevimanus var. Haddoni Coutière, 1900c: 411, unnumbered figures. [Saibai Channel, 

Torres Strait] 

Synalpheus harpagatrus Banner & Banner, 1975c: 311; Fig. 9. [5 miles west of North Beach, Perth, Western 

Australia, 50ft] 

Synalpheus hastilicrassus Coutière, 1905b: 875; Plate 72, figs 12-12d. [South Nilandu Atoll; North Male 

Atoll; Felidu Atoll; Kendikolu Reef; Miladumadulu Atoll] 

? = Synalpheus acanthitelsonis Coutière, 1905b: 875; Plate 72, figs 13-13c. [Hulule Male Atoll; Fadifolu 

Atoll; Addu Atoll; Mulaku Atoll; North male Atoll] 

= Synalpheus hastilicrassus var.? De Man, 1911d (partim): 264. [mid-channel in Solor-Strait off Kampong 

Menanga, 113 m; 8°30'S 119°7'.5E, Sapeh Strait, 73 m] 

= Synalpheus hastilicrassus var. acanthitelsoniformis De Man, 1920a: 108. [east side of Pajunga island, 

Kwandang-bay, on the reef] 

Synalpheus heardi Dardeau, 1984: 47; Figs 23-26. [Florida Middle ground, 28°36'N 84°16'W, 30-34 m] 

Synalpheus hemphilli Coutière, 1909: 39; Fig. 20. [West coast of Florida, 21-28 fms] (Fig. 48) 

= Synalpheus hemphilli longicornis Coutière, 1909: 39; Fig. 39. [West coast of Florida] 

Synalpheus herdmaniae Lebour, 1938 

= Synalpheus herdmaniæ Lebour, 1938: 651; Plate 2, figs 21-34. [Tuticorin, Gulf of Manaar, 5-12 fms] 

Synalpheus heroni Coutière, 1909: 42; Fig. 24. [Djibouti, Héron] 

Synalpheus herricki Coutière, 1909: 74; Fig. 44. [Anclote, Florida; lectotype selection by Dardeau, 1984] 

= Synalpheus herricki angustipes Coutière, 1909: 76; Fig. 45. [Anclote Section, Florida, 12.5 fms] 

= Synalpheus herricki dimidiatus Coutière, 1909: 77; Fig. 46. [Anclote Section, Florida, 12.5 fms] 

= Synalpheus tanneri Coutière, 1909: 78; Fig. 47. [Gulf of Mexico, 29°15'30''N 85°29'30''W, 27 fms] 

Synalpheus hilarulus De Man, 1910a: 290. [Wunoh-bay, N.W. coast of Waigeu-island, reef] 

Synalpheus hoetjesi Hultgren, MacDonald & Duffy, 2010: 231; Figs 3-9; Plate 4B-D. [Caracas Baai, 

Curaçao, 12°04'11.64''N 68°51'43.59''W, from the canals of Hyatella intestinalis] 

Synalpheus idios (Ríos & Duffy, 2007) 

= Zuzalpheus idios Ríos & Duffy, 2007: 36; Figs 13-17; Plate 3. [Carrie Bow Cay, in Hymeniacidion amphilecta, 

6 m]


Fig. 48. Synalpheus hemphilli Coutière, 1909. Photo by Arthur Anker. 

Synalpheus iphinoe De Man, 1909a 

= Synalpheus Iphinoë De Man, 1909a: 116. [8°23'.5S 119°4'.6E, Sapeh-strait, 70 m; Madura-bay and 

other localities in the southern part of Molo-strait, 54-90 m; Banda-anchorage, 9-36 m; Rumah-kuda-bay, 

Roma-island, 36 m] 

Synalpheus iocasta De Man, 1909a 

= Synalpheus Iocasta De Man, 1909a: 119. [Makassar and surroundings, up to 32 m; 0°58'.5N 

122°42'.5E, west of Kwadang-bay-entrance, 72 m; Anchorage north of Salomakiëe (Damar) island, 

45 m; 1°42'.5S 130°47'.5E, 32 m; 4°20'S 122°58'E, between islands of Wowoni and Buton, 

northern entrance of Buton-strait, 75-94 m; Banda-anchorage, 9-36 m; Anchorage off Pulu Jedan, 

east coast of Aru-islands (Pearl-banks), 13 m; 5°28'.2S 134°53'.9E, 57 m; 8°25'.2S 127°18'.4E, anchorage 

between Nusa Besi and the N.E. point of Timor, 27-54 m; 8°39'.1 127°4'.4E, anchorage 

south coast of Timor, 34 m; Mid-channel in Solor-strait off Kampong Menanga, 113 m; 8°30'S 

119°7'.5E, 73 m] 

Synalpheus irie MacDonald, Hultgren & Duffy, 2009: 25; Figs 11-16; Plate 3C-D. [fore-reef (near M1 channel 

marker), 18°28.083'N 77°23.289'W, from canals of Auletta cf. sycinularia] 

Synalpheus jedanensis De Man, 1909a: 117. [Anchorage off Pulu Jedan, east coast of Aru-islands (Pearlbanks), 

13 m] 

Synalpheus kensleyi (Ríos & Duffy, 2007) 

= Zuzalpheus kensleyi Ríos & Duffy, 2007: 41; Figs 18-22; Plate 3. [The Pinnacles (Sand Bores), SW of 

Carrie Bow Cay, Belize, 6 m] 

Synalpheus kuadramanus Hultgren, MacDonald & Duffy, 2010: 240; Figs 10-13; Plate 5A. [Piscadera Baai 

east, 12°07'15.96''N 68°58'11.64''W, in Xestospongia subtriangularis] 

Synalpheus kusaiensis Kubo, 1940e: 87; Fig. 10. [Kusaie, Malem, Caroline Islands] 

Synalpheus lani Hermoso & Alvarez, 2005: 522; Figs 1-2. [Punta de Mita, Nayarit, México] 

Synalpheus laticeps Coutière, 1905b: 874; Plate 72, figs 11-11d. [North Male Atoll] 

Synalpheus lockingtoni Coutière, 1909: 21; Fig. 1; nomen novum for Alpheus leviusculus Lockington, 1878b 

nec Alpheus edwardsi var. leviusculus Dana, 1852b. [Port Escondido, Mulege Bay, and other points on 

the Californian shore of the Gulf of California; Albatross stn 4421, eastern point of San Nicolas Island, 

N. 26° W. 3.8 miles, 229-298 fms]


Synalpheus longicarpus (Herrick, 1891) 

= Alpheus saulcyi var. longicarpus Herrick, 1891: 383; Plate 21, figs 5-7; Plate 22, figs 3, 11, ? 14, 17; 

Plate 24, ? figs 2, 4-9 (partim). [Bahamas, assumed to be Nassau, New Providence] 

= Synalpheus longicarpus approxima Coutière, 1909: 57; Fig. 32. [Gulf of Mexico, 26 fms] 

Synalpheus lophodactylus Coutière, 1908: 197. [Lagon de Diego] 

Synalpheus macromanus Edmondson, 1925: 9; Fig. 1. [Lisiansky Island] 

Synalpheus mcclendoni Coutière, 1910: 487; Fig. 3. [Dry Tortugas, Florida] 

Synalpheus merospiniger Coutière, 1908: 195. [Amirante, 20-44 brasses] 

Synalpheus mexicanus Coutière, 1909 

= Synalpheus townsendi mexicanus Coutière, 1909: 34; Fig. 17. [southern part of Gulf of California, 9.5 fms] 

Synalpheus microneptunus Hultgren, MacDonald & Duffy, 2011: 8; Figs 3-6. [Cement Factory, 13°17'21.84''N 

59°39'27.72''W] 

Synalpheus minus (Say, 1818) 

= A.[lpheus minus] Say, 1818: 245. [coasts of the southern states, and of East Florida] 

= Synalpheus morinus Coutière, 1898f: 189. [nomen nudum, lapsus for Synalpheus minus (Say, 1818)] 

= Synalpheus minus bahiensis Coutière, 1909: 45; Fig. 26. [Brazil, Plataforma, Bahia] 

= Synalpheus minus antillensis Coutière, 1909: 46; Fig. 27. [St. Thomas, 20-23 fms] 

= Synalpheus minus var. somersi Verrill, 1922: 108; Plate 33, figs 4-4a; Plate 34, figs 1-1u; Plate 36, figs 

1-1e, 2. [Bermuda, on the coral reefs] 

Synalpheus modestus De Man, 1909a: 115. [Banda, reef-exploration] 

Synalpheus mortenseni Banner & Banner, 1985: 47; Fig. 5. [Kei Islands, Ambon, 13-18 m] 

Synalpheus mulegensis Ríos, 1992: 9; Fig. 6. [Bahía Concepción, 22 m] 

Synalpheus mushaensis Coutière, 1908 

= Synalpheus Mushaensis Coutière, 1908: 202. [Iles Musha] 

Synalpheus neomeris (De Man, 1897) 

= Alpheus neomeris De Man, 1897: 734; Plate 35, fig. 61. [Atjeh] 

= Synalpheus Gravieri Coutière, 1905b: 870; Plate 70, figs 2-2c. [North Male Atoll] 

= Synalpheus Bougainvillei Coutière, 1905b: 871 [lapsus for Synalpheus Gravieri Coutière, 1905b] 

= Synalpheus miscellaneus De Man, 1909a: 118. [8°25'.2S 127°18'.4E, anchorage between Nusa Besi 

and the N.E. point of Timor, 27-54 m] 

Synalpheus neptunus neptunus (Dana, 1852a) 

= Alpheus neptunus Dana, 1852a: 22. [Tictabon Island, near Zamboanga, Philippines, Sulu Sea, scattered 

coral heads, 4-20 ft; neotype selection by Banner & Banner, 1972a] 

Synalpheus neptunus germanus Banner & Banner, 1975c: 321; Fig. 12. [Cottesloe, Western Australia] 

Synalpheus nilandensis Coutière, 1905b 

= Synalpheus Nilandensis Coutière, 1905b: 871; Plate 70, figs 4-4d. [North Male Atoll; Suvadiva Atoll; 

South Male Atoll; South Nilandu Atoll] 

= Synalpheus Nilandensis var. oxyceros Coutière, 1905b: 871; Plate 70, Figs 5-5a. [South Nilandu Atoll] 

= Synalpheus Nilandensis var. bandaensis De Man, 1909a: 121. [8°23'.5S 119°4'6E, Sapeh-strait, 70 m; 

Banda, 9-36 m] 

Synalpheus nobilii Coutière, 1909: 40; Fig. 22. [St. Helena, Ecuador] 

Synalpheus obtusifrons Chace, 1972: 99; Figs 37-38. [Bahía de la Ascensión, just behind center of Arrecife 

Nicchehabin, 1-5 ft] 

Synalpheus occidentalis Coutière, 1909 

= Synalpheus goodei occidentalis Coutière, 1909: 60; Fig. 34. [Bermudas] 

Synalpheus odontophorus De Man, 1909a: 113. [very near Siboga stn 65, 7°0'S 120°34'.5E, 120-400 m; 5°36'.5S 

132°55'.2E, 2.3 miles N, 63°W from the north point of Nuhu Jaan, Kei-islands, 90 m; 9°0'.3S 

126°24'.5E, 112 m] 

Synalpheus orapilosus Hultgren, MacDonald & Duffy, 2010: 246; Figs 14-17, Plate 5D-E. [Caracas Baai, 

Curaçao, 12°04'11.64''N 68°51'43.56''W, in an unidentified white web-like sponge embedded in Madracis 

sp. rubble] 

Synalpheus osburni Schmitt, 1933: 1; Fig. 1. [inside Cayo Maria Langa, near Guayanilla Harbor, Porto 

Rico]


Synalpheus otiosus Coutière, 1908: 195. [Coevity] 

Synalpheus pachymeris Coutière, 1905b 

= Synalpheus biunguiculatus var. pachymeris Coutière, 1905b: 873; Plate 71, figs 9-9a’. [South Nilandu 

Atoll] 

= Synalpheus pachymeris var. Cargadosi Coutière, 1921: 417; Plate 61, fig. 10. [Cargados Carajos] 

Synalpheus pandionis Coutière, 1909: 67; Fig. 39. [St. Thomas, 20-23 fms] 

= Synalpheus pandionis extentus Coutière, 1909: 69; Fig. 40. [St. Thomas, 20-23 fms] 

= Synalpheus grampusi Coutière, 1909: 62; Fig. 36. [Gulf of Mexico, 26°33'N 83°10'W, 28 fms] 

Synalpheus paradoxus Banner & Banner, 1981a: 68; Fig. 10. [Negus Salomon, 15°37'N 40°43'E, 15 fms] 

Synalpheus paralaticeps Banner & Banner, 1982a: 299; Fig. 92. [Rudder Reef, off Port Douglas, Queensland, 

outer reef flat, 200 yards from edge] 

Synalpheus paraneomeris Coutière, 1905b: 872; Plate 71, figs 7-7e. [Goidu Atoll, Minikoi] 

= Synalpheus paraneomeris Coutière, 1899a: 456, 466, 486, 501. [nomen nudum] 

= Synalpheus paraneomeris oxyceros Coutière, 1909: 9. [nomen nudum] 

= Synalpheus paraneomeris prolatus Coutière, 1909: 9. [nomen nudum; see discussion in Banner, 1953] 

= Synalpheus paraneomeris var. halmaherensis De Man, 1909a: 122. [Wunoh-bay, NW coast of Waigeuisland] 

= Synalpheus paraneomeris var. praedabunda De Man, 1909a: 123. [Anchorage off Labuan Pandan, 

Lombok, 18 m; Sailus Keyjil, Paternoster-islands, 27 m; Anchorage off Kawio-and Kamboling-island, 

Karkarolong-group, reef; Anchorage off Kilsuin, west-coast of Kur-island, reef-exploration] 

= Synalpheus paraneomeris var. prolatus De Man, 1911d: 241. [Wunoh-bay, NW coast of Waigeu-island, 

reef; Anchorage off the south point of Kabaëna-island, reef] 

= Synalpheus Sluiteri De Man, 1920a: 107. [Bay of Batavia] 

= Synalpheus paraneomeris var. praslini Coutière, 1921: 415; Plate 61, fig. 6. [Seychelles, Praslin Reef] 

= Synalpheus paraneomeris var. seychellensis Coutière, 1921: 415; Plate 61, fig. 7. [Seychelles] 

Synalpheus paraneptunus Coutière, 1909: 86; Fig. 52 (partim). [Panama, Caribbean coast, off Golfo de 

Morrosquillo, 09°30'N 76°20'W, 77 m] 

Synalpheus parfaiti (Coutière, 1898f) 

= Synalpheus laevimanus var. Parfaiti Coutière, 1898f: 191; Figs 3-3a. [Annobon] 

Synalpheus paulsoni paulsoni Nobili, 1906c 

= Synalpheus Paulsoni Nobili, 1906c: 28; Fig. 1. [Banc au NE de l’île Arzana] 

Synalpheus paulsoni kurracheensis Coutière, 1908 

= Synalpheus Paulsoni Kurracheensis Coutière, 1908: 203. [Kurrachee] 

Synalpheus paulsoni liminaris Coutière, 1908 

= Synalpheus Paulsoni liminaris Coutière, 1908: 201. [Djibouti; Golfe Persique] 

Synalpheus paulsoni rameswarensis Coutière, 1908 

= Synalpheus Paulsoni Rameswarensis Coutière, 1908: 201. [Rameswaran] 

Synalpheus paulsonoides Coutière, 1909 

= Synalpheus paulsonoïdes Coutière, 1909: 24; Fig. 6. [island of San José, Lower California] 

Synalpheus pectiniger Coutière, 1907b: 611. [Albatross stn 2413 (26°N 082°57'30''W), golfe de Mexico, 

banc de la Floride, 24 brasses; Curaçao; full description in Coutière, 1909] 

= Alpheus Praecox Herrick, 1888: 34. [nomen oblitum; see De Grave & Anker, 2010] 

Synalpheus pescadorensis Coutière, 1905b 

= Synalpheus Pescadorensis Coutière, 1905b: 877; Plate 73, figs 15-15e. [Miladumadulu Atoll; South 

Nilandu Atoll; North Male Atoll, Pescadores] 

Synalpheus peruvianus Rathbun, 1910 

= Synalpheus townsendi peruvianus Rathbun, 1910: 563; Plate 53, fig. 4. [oysterbeds of Matapalo, near 

Capon] 

Synalpheus plumosetosus MacDonald, Hultgren & Duffy, 2009: 36; Figs 17-21. [Jamaica, Dairy Bull Reef, 

18°28.083'N 77°23.289'W, from canals of Auletta cf. sycinularia] 

Synalpheus pococki Coutière, 1898c 

= Synalpheus neomeris var. Pococki Coutière, 1898c: 167; Figs 2-2a’. [Holothuria Bank, NW Australie; 

Macclesfield Bank, Arafura See]


Synalpheus quadriarticulatus Banner & Banner, 1975c: 297; Fig. 5. [between Hammond and Waivea Islands, 

Torres Straits, 3 m] 

Synalpheus quadrispinosus De Man, 1910a: 298. [8°23'.5S 119°4'.6E, Sapeh-Strait, 70 m; 1°42'.5S 130°47'.5E, 

32 m; Anchorage off Pulu jedan, East coast of Aru-islands (Pearl-banks), 13 m; 8°30'S 119°7'.5E, 73 

m; Anchorage East of Sailus Besar, Paternoster-islands, up to 36 m] 

= Synalpheus quadridens De Man, 1910a: 299. [8°25'.2S 127°18'.4E, anchorage between Nusa besi and 

the N.E. point of Timor, 27-54 m] 

Synalpheus quinquedens Tattersall, 1921: 376; Plate 28, figs 1-5. [Suez, among coral; Khor Dongonab, 

among coral on reef; Mersa Ar-rakiya, 20°15'N, among coral in 1ft; Suakin Harbour, 19°8'N, washed 

from sponges] 

Synalpheus rathbunae Coutière, 1909: 84; Fig. 51. [St. Thomas, 20-30 fms] 

Synalpheus readi Banner & Banner, 1972b: 137; Fig. 1. [Malakal Harbor, Urukthapel, Palau Island, 5 m] 

Synalpheus recessus Abele & W. Kim, 1989: 15; Figs 7- 8. [Republic of Panama, Panama Canal, Miraflores 

Locks] 

Synalpheus redactocarpus Banner, 1953: 29; Fig. 8. [Keaoi Island, Halape, Kau Coast, Hawaii, intertidal] 

Synalpheus regalis Duffy, 1996: 564; Figs 1-5. [outer reef ridge at Carrie Bow Cay, Belize, 16°48'N 88°05'W, 

15 m, from Xestospongia cf. subtriangularis] 

Synalpheus riosi Anker & Tóth, 2008: 11; Figs 6-7. [Dominica, Prince Rupert Bay, 1.5 m] 

Synalpheus ruetzleri MacDonald & Duffy, 2006: 8; Figs 8-13. [Sand Bores, Belize, from canals of Hymeniacidon 

cf. caerulea] 

Synalpheus sanctithomae Coutière, 1909: 61; Fig. 35. [St. Thomas, 20-30 fms] 

Synalpheus sanjosei Coutière, 1909 

= Synalpheus apioceros sanjosei Coutière, 1909: 29; Fig. 10. [San José, Lower California] 

Synalpheus sanlucasi Coutière, 1909: 41; Fig. 23. [Cape St. Lucas, Lower California] 

Synalpheus scaphoceris Coutière, 1910 

= Synalpheus townsendi scaphoceris Coutière, 1910: 486; Fig. 2. [Dry Tortugas, Florida] 

Synalpheus sciro Banner & Banner, 1975c: 304; Fig. 7. [27°40'S 113°20'E, northwest of Bluff Point, Western 

Australia, 7.5 fms] 

Synalpheus senegambiensis Coutière, 1908 

= Synalpheus Paulsoni Senegambiensis Coutière, 1908: 202. [Cap Vert] 

Synalpheus septemspinosus De Man, 1910a: 297. [Anchorage east of Sailus Besar, Paternoster-islands, up 

to 36 m] 

Synalpheus sladeni Coutière, 1908 

= Synalpheus Sladeni Coutière, 1908: 198. [Cargados Carajos, 30 brasses] 

Synalpheus somalia Banner & Banner, 1979: 31; Fig. 3. [11°11'N 51°14'E, off Somalia, 47-49 m] 

Synalpheus spinifrons (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= A.[lpheus] spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 355. [côtes de 

Chili] 

= Synalpheus latastei Coutière, 1909: 25; Fig. 7. [Chile; ?New Holland] 

Synalpheus spiniger (Stimpson, 1860a) 

= Alpheus spiniger Stimpson, 1860a: 31. [ad insulas ”Amakirrima” prope ”Loo Choo”] 

Synalpheus spongicola Banner & Banner, 1981a: 78; Fig. 12. [Marsa Murach, from inside sponge] 

Synalpheus stimpsonii (De Man, 1888b) 

= Alpheus Stimpsonii De Man, 1888b: 513; Plate 22, fig. 3a-b. [Amboina] 

= Alpheus amboinæ Zehntner, 1894: 202; Plate 8, figs 23-23b. [Amboine] 

= Alpheinus tridens Borradaile, 1900: 415; Plate 38, Figs 12a-g. [Sandal Bay, Lifu, Loyalty Islands] 

= Alpheus stimpsoni Var.? Schenkel, 1902: 567; Plate 13, figs 22a-c. [Kema] 

= Synalpheus Stimpsoni var. Maldivensis Coutière, 1905b: 878; Plate 73, figs 16-16d. [Hulule Male 

Atoll] 

= Synalpheus consobrinus De Man, 1909a: Moearas-reef, inner side, east coast of Borneo, up to 54 m; 

Saleyer-anchorage and surroundings, reef-exploration; 8°25'.2S 127°18'.4E, anchorage between 

Nusa Besi and the NE point of Timor, 27-54 m; Mid-channel in Solor-strait off Kampong Menanga, 

113 m]


= Synalpheus brucei Potts, 1915: 76; Figs 1-3; Plate 1, fig. 2. [Murray Island] 

= Synalpheus striatus Kubo, 1938c: 89; Figs 1-2. [Ôsima Island, Wakayam Prefecture] 

Synalpheus streptodactylus Coutière, 1905b 

= Synalpheus neomeris var. streptodactylus Coutière, 1905b: 870; Plate 70, fig. 1’. [Haddumati Atoll; 

Suvadiva Atoll; South Nilandu Atoll] 

= Synalpheus streptodactyloides De Man, 1909a: 114. [8°25'.2S 127°18'.4E, anchorage between Nusa 

Besi and the NE point of Timor, 27-54 m] 

= Synalpheus metaneomeris Coutière, 1921: 414; Plate 60, fig. 4. [Amirante Bank, 20-80 fms; Saya de 

Malha, 47 and 90 fms; Cargados carajos, 28-33 fms; Providence, 58 fms] 

= Synalpheus streptodactylus hadrungus Banner & Banner, 1966b: 158. [nomen novum for S. metaneomeris 

streptodactylus Coutière, 1921] 

Synalpheus stylopleuron Hermoso Salazar & Hendrickx, 2006: 1109; Figs 3-4. [off Lobos Island, 23°13'49"N 

106°27'43"W, Sinaloa, Mexico, in sponge Mycale parishii, 3-4 m] 

Synalpheus superus Abele & W. Kim, 1989: 18; Figs 9-10. [Republic of Panama, Panama Canal, Miraflores 

Locks] 

Synalpheus tenuispina Coutière, 1909 

= Synalpheus latastei tenuispina Coutière, 1909: 26; Fig. 8. [Desterro, Brazil] 

Synalpheus thai Banner & Banner, 1966a: 61; Fig. 19. [off Koh Samet, Rayong, Thailand, 2 m] 

Synalpheus thele MacDonald, Hultgren & Duffy, 2009: 43; Figs 22-27; Plate 5C-D. [Jamaica, Columbus 

Park, Discovery Bay, 18°27.955'N 77°24.843'W, from canals of Agelas cf. clathrodes] 

Synalpheus theano De Man, 1910a 

= Synalpheus Theano De Man, 1910a: 296. [1°42.5'S 130°47.5'E, 32 m] 

Synalpheus tijou Banner & Banner, 1982a: 296; Fig. 91. [Tijou Reef, Lizard Island, with a crinoid] 

Synalpheus townsendi Coutière, 1909: 32; Fig. 14. [Gulf of Mexico (29°14'00''N 085°29'15''W, 25 fms)] 

= Synalpheus townsendi productus Coutière, 1909: 33; Fig. 15. [Gulf of Mexico (28°46'00''N 084°49'00''W, 

26 fms)] 

Synalpheus triacanthus De Man, 1910a: 301. [9°0.3'S 126°24.5'E, 112 m, living in Solenocaulon] 

Synalpheus tricuspidatus (Heller, 1861) 

= A.[lpheus] tricuspidatus Heller, 1861: 24. [im rothen Meere] 

= A.[lpheus] tricuspidatus Heller, 1862c: 267; Plate 3, fig. 15. [im rothen Meere] 

Synalpheus tridentulatus (Dana, 1852a) 

= Alpheus tridentulatus Dana, 1852a: 22. [in portu ”Rio Janeiro”?] 

Synalpheus trispinosus De Man, 1910a: 300. [Madura-bay and other localities in the southern part of 

Molo-strait, 54-90 m] 

Synalpheus triunguiculatus (Paul’son, 1875) 

= Alph.[eus] triunguiculatus Paul’son, 1875: 103; Plate 14, figs 1-1g. [Red Sea] 

= Synalpheus physocheles Coutière, 1908: 200. [Djibouti] 

Synalpheus tropidodactylus Banner & Banner, 1975c: 286; Fig. 2a-k, o. [west of Geraldton, 28°14'S 113°14'E, 

60 fms] 

Synalpheus tumidomanus tumidomanus (Paul’son, 1875) 

= Alph.[eus] tumido-manus Paul’son, 1875: 101; Plate 13, figs 2-2i. [Red Sea] 

= Alph.[eus] tumido-manus Var. Alph.[eus] gracili-manus Paul’son, 1875: 102; Plate 13, figs 3-3c. [Red 

Sea] 

= Synalpheus Hululensis Coutière, 1908: 202. [Maldives] 

= Synalpheus Mac-Cullochi Coutière, 1908: 203. [Côte S.W. d’Australie] 

= Synalpheus tumidomanus var. exilimanus Coutière, 1909: 10. [lapsus for Alph.[eus] tumido-manus 

Var. Alph.[eus] gracili-manus Paul’son, 1875] 

= Synalpheus Theophane De Man, 1910a: 292. [Lumu-lumu-shoal, Borneo-bank, reef; Anchorage 

north of Salomakiee-(Damar)-island, reef; 8°25'.2S 127°18'.4E, anchorage between Nusa-Besi and 

the N.E. point of Timor] 

= Synalpheus anisocheir Stebbing, 1915: 86; Plate 87. [Gordon’s Bay, False Bay] 

= Synalpheus japonicus Yokoya, 1936: 133; Fig. 3. [vicinity of the Misaki Marine Biological Station]


Synalpheus tumidomanus africanus Crosnier & Forest, 1965b 

= Synalpheus hululensis africanus Crosnier & Forest, 1965b: 607; Fig. 2. [São Tomé, Morro Peixe, 2-6 

m] 

Synalpheus tumidomanus congoensis Crosnier & Forest, 1965b 

= Synalpheus hululensis congoensis Crosnier & Forest, 1965b: 608; Fig. 3. [Djéno, près de Pointe- 

Noire (Congo)] 

Synalpheus tuthilli Banner, 1959: 133; Fig. 2. [Tomil Harbor, Yap Island, Caroline Archipelago] 

Synalpheus ul (Ríos & Duffy, 2007) 

= Zuzalpheus ul Ríos & Duffy, 2007: 63; Figs 27-30; Plate 5. [Sandbores, Belize, in Hymeniacidion 

caerulea, 2 m] 

Synalpheus wickstenae Hermoso Salazar & Hendrickx, 2006: 1101; Figs 1-2. [off Altata (24°35'N 

107°55'W), Sinaloa, Mexico, in sponges, 10 m] 

Synalpheus williamsi Ríos & Duffy, 1999: 542; Figs 1-6. [Carrie Bow Cay, Belize, from Hymeniacidon 

caerulea, 18 m] 

Synalpheus yano (Ríos & Duffy, 2007) 

= Zuzalpheus yano Ríos & Duffy, 2007: 69; Figs 31-34; Plate 5. [Twin Cays, Belize, in Lissodendoryx 

colombiensis, 1-2 m] 

Thuylamea Nguyên, 2001 

= Thuylamea Nguyên, 2001 (type species Thuylamea camelus Nguyên, 2001, by original designation 


Thuylamea camelus Nguyên, 2001: 218; Figs 1-22. [Donghoa, a coastal community of the Can Gio district, 

21 km northwest of Vung Tau (Cap St Jacques), Vietnam, 10°20'N 106°52'E, in an estuarine 

area formed by the confluence of two large rivers, 5-9 m] 

Triacanthoneus Anker, 2010e 

= Triacanthoneus Anker, 2010e (type species Triacanthoneus toro Anker, 2010e, by original designation, 


Triacanthoneus alacranes Anker, 2010e: 57; Figs 7-9, 10G. [Mexico, Yucatán, Arrecife Alacranes, La Anegada, 

22°27'37''N 89°36'40.6''W, 0.5 m] 

Triacanthoneus pacificus Anker, 2010e: 54; Figs 5-6, 10D-F. [Panama, Pacific coast, Playa Venao, intertidal] 

Triacanthoneus toro Anker, 2010e: 49; Figs 1-4, 10A-C. [Panama, Caribbean coast, Bocas del Toro, Isla 

Colón, Punta Caracol, 1.5 m] 

Vexillipar Chace, 1988 

= Vexillipar Chace, 1988 (type species Vexillipar repandum Chace, 1988, by original designation and 

monotypy, gender neuter) 

Vexillipar repandum Chace, 1988: 91; Figs 23-25. [Off Murcielago Bay, Mindanao, Philippines, 8°47'15''N 

123°35'00''E, 296 m] 

Yagerocaris Kensley, 1988 

= Yagerocaris Kensley, 1988 (type species Yagerocaris cozumel Kensley, 1988, by original designation 


Yagerocaris cozumel Kensley, 1988: 693; Figs 4-6. [Areolito Cenote, Cozumel, Quintana Roo, Mexico, 40 ft] 

Family BARBouRIIDAe Christoffersen, 1987 

Barbouria Rathbun, 1912b 

= Barbouria Rathbun, 1912b (type species Barbouria poeyi Rathbun, 1912b (junior subjective synonym 

of Hippolyte cubensis von Martens, 1872), by original designation and monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in Opinion 470 in 1957)


Barbouria cubensis (von Martens, 1872) 

= Hippolyte Cubensis von Martens, 1872: 136; Plate 5, fig. 14. [Cuba] 

= Barbouria poeyi Rathbun, 1912b: 455; Plates 2-5. [Cave near seashore, between Morro Castle and 

Cojimar] 

Barbouria yanezi Mejía, Zarza & López, 2008: 663; Figs 1-2. [Cenote Tres Potrillos, Cozumel Island, Mexico, 

20°27'05''N 86°59'15''W, 26 m] 

Janicea Manning & C.W.J. Hart, 1984 

= Janicea Manning & C.W.J. Hart, 1984 (type species Barbouria antiguensis Chace, 1972, by original 


Janicea antiguensis (Chace, 1972) 

= Barbouria antiguensis Chace, 1972: 107; Figs 40-41. [Antigua Island, English Harbour] 

Parhippolyte Borradaile, 1900 

= Parhippolyte Borradaile, 1900 (type species Parhippolyte uveae Borradaile, 1900, by monotypy, gender 

feminine) 

= Somersiella C.W.J. Hart & Manning, 1981 (type species Somersiella sterreri C.W.J. Hart & Manning, 


= Koror J. Clark, 1989 (type species Koror misticius J. Clark, 1989, by original designation and monotypy, 


Parhippolyte cavernicola Wicksten, 1996a: 201; Figs 1-4. [San Diego Reef, west of San Diego Island (north 

of San Jose Island), Baja California Sur, Mexico, 25°20'N 110°40'W, 6 m, in sea cave] 

Parhippolyte misticia (J. Clark, 1989) (Fig. 49) 

= Koror misticius J. Clark, 1989: 446; Figs 1-4. [South Point Cave, Koror, Ngermeuangel, Palau, 

7°18'32"N 134°30'05"E] 

Parhippolyte rukuensis Burukovsky, 2007c: 3; Fig. 2. [Japan, Ryukyu Islands, Ie-jima, W of Okinawa, in 

sea-connected cave, 30 m] 

Fig. 49. Parhippolyte misticia (J. Clark, 1989). Photo by John Hoover.


Parhippolyte sterreri (C.W.J. Hart & Manning, 1981) 

= Somersiella sterreri C.W.J. Hart & Manning, 1981: 442; Figs 1-28. [Tucker’s Town Cave, Tucker’s 

Town, Bermuda] 

Parhippolyte uveae Borradaile, 1900: 414; Plate 38, figs 11a-g. [Uvea, Loyalty Islands] 

Family HIPPolyTIDAe Spence Bate, 1888 

Alcyonohippolyte Marin, Chan & okuno, 2011 

= Alcyonohippolyte Marin, Chan & Okuno, 2011 (type species Alcyonohippolyte dossena Marin, Chan 

& Okuno, 2011, by original designation, gender feminine) 

Alcyonohippolyte commensalis (Kemp, 1925) 

= Hippolyte commensalis Kemp, 1925: 331; Figs 21-22. [Coral reef off Reed Point, Nancowry Island, 

Nicobars, on a compound actinian] 

Alcyonohippolyte dossena Marin, Chan & Okuno, 2011: 34; Figs 1-4, 16A. [Izu Islands, Hachijo-jima Island, 

Yaene fishing port, 5 m] 

Alcyonohippolyte maculata Marin, Chan & Okuno, 2011: 46; Figs 12-13, 14D-F, 15B-H, 16C. [Taiwan, Keelung, 

Badouzih, 5-6 m] 

Alope White, 1847b 

= Alope White, 1847b (type species Alope palpalis White, 1847b (junior subjective synonym of Hippolyte 

spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]), by monotypy, gender 


= Hetairocaris De Man, 1890 (type species Hetairocaris orientalis De Man, 1890, by monotypy, gender 

feminine) 

Alope orientalis (De Man, 1890) 

= Hetairocaris orientalis De Man, 1890: 122; Plate 6, fig. 16-16c. [Ponapé] 

= Hippolyte ponapensis Ortmann, 1890: 502; Plate 36, figs 20-20d. [Karolinen, Ponapé] 

= Alope australis Baker, 1904: 154; Plate 30, figs 1-7. [Smith’s Bay, Kangaroo Island, shallow water] 

Alope spinifrons (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Hippolyte spinifrons H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 377. [les côtes de la 

Nouvelle-Zélande] 

= Alope palpalis White, 1847b: 124. [New Zealand] 

Bathyhippolyte Hayashi & Miyake, 1970 

= Bathyhippolyte Hayashi & Miyake, 1970 (type species Bathyhippolyte yaldwyni Hayashi & Miyake, 


Bathyhippolyte yaldwyni Hayashi & Miyake, 1970: 42; Figs 1-16. [Chatham Rise, New Zealand, 44°44'S 

175°42'E, 995-1110 m] 

Birulia Bražnikov, 1903 

= Birulia Bražnikov, 1903 (type species Birulia sachalensis Bražnikov, 1903, by monotypy, gender 


= Paraspirontocaris Yokoya, 1930 (type species Paraspirontocaris kishinouyei Yokoya, 1930, by monotypy, 


Birulia kishinouyei (Yokoya, 1930) 

= Paraspirontocaris kishinouyei Yokoya, 1930: 536; Plate 16, figs 1-6. [on the line between Kanita and 

Yokohama, off Ôshima Isl.; off Tairadate] 

Birulia sachalinensis Bražnikov, 1903: 44. [southern Sakhalin Island, Russian Far East] 

Bythocarides Sokolov, 2002 

= Bythocarides Sokolov, 2002 (type species Bythocarides menshutkinae Sokolov, 2002, by original designation 


Bythocarides menshutkinae Sokolov, 2002: 139; Figs 1-3. [Laptev Sea, 81°10.8'N 141°47.1'E, 945 m]


Bythocaris G.o. Sars, 1870 

= Bythocaris G.O. Sars, 1870 (type species Bythocaris simplicirostris G.O. Sars, 1870, by monotypy; 


Bythocaris akidopleura Fransen, 1993: 588; Figs 41-62. [Azores, West of Formigas, 37°17'N 25°14'W, 2070- 

2120 m] 

Bythocaris biruli Kobjakova, 1964 

= Bythocaris leucopis biruli Kobjakova, 1964: 326; Fig. 1. [Arctic Ocean, 81°18'N 9°36'E, 1300 m] 

Bythocaris cosmetops Holthuis, 1951a: 135; Fig. 30. [off Sierra Leone, 7°29'N 13°38'W, 74-78 m] 

Bythocaris cryonesus Bowman & Manning, 1972: 189; Fig. 1. [Arctic Ocean, 81°33.9'N 157°12.5'W, 3803 

m] 

Bythocaris curvirostris Kobjakova, 1957: 363; Figs 1-2. [Arctic Basin, eastern sector, 3255 m] 

Bythocaris elegans Bryazgin, 1982: 603; Figs 1-7. [Arctic Basin, western flank of Franz-Victoria Trough, 

81°51'N 36°20'E, 420 m] 

Bythocaris floridensis Abele & Martin, 1989: 29; Fig. 1. [Florida, east of Jacksonville, 31°09'N 79°33'30"W, 

644 m] 

Bythocaris gorei Abele & Martin, 1989: 38; Fig. 2. [Florida, East of St. Augustine, 29°41'N 79°55'W, 682 m] 

Bythocaris gracilis Smith, 1885a: 497. [Albatross stns 2116, 35°5'23''N 74°31'25''W, 888 fms; 2206, 

39°35'00''N 71°24'30''W, 1043 fms] 

Bythocaris grumanti Burukovsky, 1966: 538; Fig. 2. [south end of Spitzbergen, Strait between island’s 

edge and Nadejda, 50 m] 

Bythocaris irene Retowsky, 1946: 298; Fig. 1. [IS ”G.Sedov” stn 35/71, Kara Sea, 520 m according to 

Sokolov, 2000] 

Bythocaris kobjakovae Sokolov, 2000: 461; Figs 37-39. [Laptev Sea; 1368 m] 

Bythocaris leucopis G.O. Sars, 1879: 427. [71.59°N 11.40°W, 1110 fms] 

Bythocaris miserabilis Abele & Martin, 1989: 41; Fig. 3. [Southeast Florida, 27°11'N 79°30'W, 677 m] 

Bythocaris nana Smith, 1885a: 499. [Albatross stns 865, 40°05'00''N 70°23'00''W, 65 fms; 872, 40°05'39''N 

70°23'52''W, 86 fms; 874, 40°00'00''N 70°57'00''W, 85 fms; 878, 39°55'00''N 70°54'15''W, 143 fms (all 

off Martha’s Vineyard)] 

Bythocaris payeri (Heller, 1875a) 

= Hippolyte Payeri Heller, 1875a: 609. [full description in Heller, 1875b: 26; Plate 1, figs 1-4; type locality 

therein as Nordpol, 182 m; corrected in Fransen, 1993 to S of Franz Josef Land, 79°0.4'N 

62°29.7'E, 182 m] 

Bythocaris simplicirostris G.O. Sars, 1870: 149. [Lofoten, 250 orgyarum] 

= Hippolyte Panschii Buchholz, 1874: 277; Plate 1, fig. 1. [Nordshannon-Insel, 30 faden] 

= Bythocaris spinipleura Squires, 1990: 158; Figs 82-83. [off Bonavista Bay, Newfoundland, 48°49'N 

51°30'W, 309 m, from cod stomachs] 

Calliasmata Holthuis, 1973a 

= Calliasmata Holthuis, 1973a (type species Calliasmata pholidota Holthuis, 1973a, by original designation 


Calliasmata nohochi Escobar-Briones, Camacho & Alcocer, 1997: 733; Figs 2-4. [Crack House Cave, Nohoch 

Nah Chich system, Quintana Roo, Mexico] 

Calliasmata pholidota Holthuis, 1973a: 37; Figs 12-13; Plate 1, fig. 2; Plate 2, fig. 1. [Ras Muhammad 

Crack, near Ras Muhammad, southern tip of Sinai Peninsula, in pool in narrow crack in elevated 

coral rock, about 150 m from the sea] 

Calliasmata rimolii Chace, 1975: 37; Figs 5-7. [cave 4 km from town of Estero Hondo (19°51'N 71°11'W), 

Provincia de Puerto Plate, northern Dominican Republic] 

Caridion Goës, 1864 

= Doryphorus Norman, 1861 (type species Hippolyte Gordoni Spence Bate, 1858, by monotypy, gender 

masculine; invalid junior homonym of Doryphorus Cuvier, 1829 (Reptilia); name placed on the Official 

Index of Rejected and Invalid Generic Names in Zoology in Opinion 470 in 1957)


= Caridion Goës, 1864 (nomen novum for Doryphorus Norman, 1861, gender neuter; name placed on 

the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Caridion gordoni (Spence Bate, 1858) 

= Hippolyte Gordoni Spence Bate, 1858: iv, 52; unnumbered figures. [Moray Firth] 

Caridion steveni Lebour, 1930: 185; Plate 1; Plate 4, fig. 2. [rocky pools under stones at low water and between 

tide-marks in bays around Plymouth, Wembury Bay, Rum Bay, Cawsand Bay, Drake’s Island, 

below Mount Edgcumbe] 

Chorismus Spence Bate, 1888 

= Chorismus Spence Bate, 1888 (type species Chorismus tuberculatus Spence Bate, 1888, by monotypy, 


1957) 

Chorismus antarcticus (Pfeffer, 1887) 

= Hippolyte antarctica Pfeffer, 1887: 51; Plate 1, figs 22-27. [Süd-Georgien, 7-9 Faden] 

= Hippolyte Romanchei A. Milne-Edwards, 1891: 45; Plate 5, figs 1-1d. [Punta Arenas, 25 m] 

Chorismus tuberculatus Spence Bate, 1888: 617; Plate 110, fig. 2. [Challenger stn 145A, 46°41'S 38°10'E, off 


Cryptocheles G.o. Sars, 1870 

= Cryptocheles G.O. Sars, 1870 (type species Cryptocheles pygmaea G.O. Sars, 1870, by original designation 

and monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology 


Cryptocheles pygmaea G.O. Sars, 1870: 150. [Lofoten, 120-800 orgyarum] 

= Cryptocheles abyssicola M. Sars, 1869: 262, 275. [nomen nudum] 

Eualus Thallwitz, 1892 

= Eualus Thallwitz, 1892 (type species Eualus obses Thallwitz, 1892 (junior subjective synonym of 

Hippolyte Gaimardii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840), gender masculine; 


= Helia Thallwitz, 1892 (type species Hippolyte Fabricii Krøyer, 1841, by monotypy, gender feminine; 

invalid junior homonym of Helia Hübner (Lepidoptera)) 

? = Vianellia Nardo, 1847 (type species Vianellia dorsioculata Nardo, 1847, by monotypy, gender feminine; 

name suppressed for the purposes of the Principle of Priority but not for those of the Principle 

of Homonomy under the plenary power of the ICZN, and placed on the Official Index of Rejected 

and Invalid Generic Names in Zoology, in Opinion 671, in 1963) 

= Lysippe Kinahan, 1858a (type species Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875], by 

monotypy, gender masculine; generic name suppressed for the purposes of both the Principle of 

Priority and the Principle of Homonomy under the plenary power of the ICZN, and placed on the 

Official Index of Rejected and Invalid Generic Names in Zoology, in Opinion 671, in 1963) 

= Spirontocarella Bražnikov, 1907 (type species Hippolyte macilenta Krøyer, 1841, by monotypy, gender 

feminine) 

= Thoralus Holthuis, 1947a (type species Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875], by 

original designation, gender masculine; name placed on the Official List of Generic Names in Zoology 


Eualus avinus (Rathbun, 1899) 

= Spirontocaris avina Rathbun, 1899: 557. [North of Unalaska Island, 54°00'45"N 166°53'50"W, 351 fms] 

Eualus barbatus (Rathbun, 1899) 

= Spirontocaris barbata Rathbun, 1899: 556. [56°18'N 160°38'W, 86 fms] 

Eualus berkeleyorum Butler, 1971: 1616; Figs 1-2. [off Gabriola Island, Strait of Georgia, 49°09.0'N 

123°32.6'W, 384 m] 

Eualus biunguis (Rathbun, 1902a) 

= Spirontocaris biunguis Rathbun, 1902a: 899. [off Cape St. James, Queen Charlotte Islands, British 

Columbia, 876 fms]


Eualus bulychevae Kobjakova, 1955: 238; Figs 4-5. [South Kurile Straits, Kuril Islands, max depth 54 m] 

Eualus butleri Jensen, 2004: 463; Figs 1-2. [Gastineau Channel, Juneau, Alaska, 58°17'N 134°24'W, 30 m, 

with sponges] 

Eualus cteniferus (Barnard, 1950) 

= Spirontocaris ctenifera Barnard, 1950: 696; Fig. 129c-k. [Algoa Bay; Durban] 

Eualus cranchii (Leach, 1817 [in Leach, 1815-1875]) 

? = P.[alemon] Microramphos Risso, 1816: 104. [dans les rochers du rivage, environs de Nice] 

= Hippolyte Cranchii Leach, 1817 [in Leach, 1815-1875]: Plate 38, Figs 17-21. [southern point of the 

Saltstone, in the Kingsbridge Estuary] 

? = Hippolyte Crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 375. [St Malo, 

Brittany, France] 

= Hippolyte mutila Krøyer, 1841: 573. [Norges Vestkyst] 

= Hippolyte Bunseni Neumann, 1878: 36. [Palma de Mallorka] 

? = Hippolyte Lovenii Rathke, 1843: 13. [Molde] 

? = Vianellia dorsioculata Nardo, 1847: 8. [Lagune e Golfo Veneto] 

? = Hippolyte Yarrellii W. Thompson, 1853: 112; Plate 6, figs 2-2b. [Weymouth Bay, 5-7 fms] 

Eualus dozei (A. Milne-Edwards, 1891) 

= Hippolyte Dozei A. Milne-Edwards, 1891: 46; Plate 5, figs 3-3c. [à l’île Grevy, 65 m] 

Eualus drachi Noël, 1978: 23; Figs 1-2. [Banyuls-sur-mer, Méditerranée, 95 m] 

Eualus fabricii (Krøyer, 1841) 

= Hippolyte Fabricii Krøyer, 1841: 571. [Grønland] 

Eaulus gaimardii belcheri (Bell, 1855) 

= Hippolyte belcheri Bell, 1855: 402; Plate 34, fig. 1. [between Beechey Island and Northumberland 

Sound] 

Eualus gaimardii gaimardii (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

? = Cancer nautilor Herbst, 1796 [in Herbst, 1791-1796]: 175; Plate 43, fig. 4. 

= Hippolyte Gaimardii H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les mers 

d’Islande] 

= Hippolyte gibba Krøyer, 1841: 572. [Spitsbergen og Grønland] 

= Hippolyte recurvirostris Rathke, 1843: 12. [Christiansund] 

= Hippolyte lentiginosa Rathke, 1843: 14. [bei Molde und Christiansund] 

= Hippolyte Retzii Rathke, 1843: 16. [Meerbusen von Drontheim] 

= H.[ippolyte] gracilis Liljeborg, 1851: 83. [Tromsö] 

= Hippolyte pandaliformis Bell, 1851 [in Bell, 1844-1853]: 294; unnumbered vignette. [Loch Fyne, 

about 20 fms] 

= Eualus obses Thallwitz, 1892: 23. [Grönland] 

= Spirontocaris recurvirostris Molander, 1914: 1; Plate 1, fig. 1. [Vaigattet, Greenland, 315 m] 

Eualus gracilipes Crosnier & Forest, 1973: 163; Fig. 50. [San Tiago, Cape Verde Islands, 275-150 m] 

Eualus gracilirostris (Stimpson, 1860a) 

= Hippolyte gracilirostris Stimpson, 1860a: 34. [portu ”Hakodadi”, in regione laminarium] 

Eualus horii Komai & Hayashi, 2002: 382; Figs 1-5. [Off Hayatsuki-gawa Rivermouth, Toyama Bay, 80 m] 

Eualus kikuchii Miyake & Hayashi, 1967: 261; Figs 6-7. [Tomioka Bay, Amakusa Island] 

Eualus kinzeri Tiefenbacher, 1990: 117; Fig. 1. [Weddell Sea, 71°06.2'S 12°53.8'W, 771 m] 

Eualus kuratai Miyake & Hayashi, 1967: 253; Fig. 3. [between Rebun I. and Rishiri I., Hokkaido, 100- 

150 m] 

Eualus lebourae Holthuis, 1951a: 124; Fig. 26. [off French Guinea, 10°49'N 16°39'W, 42 m] 

Eualus leptognathus (Stimpson, 1860a) 

= Hippolyte leptognatha Stimpson, 1860a: 34. [sinu ”Hakodadi”, vulgaris in fundis algoso-arenoso, 

prof. 2-6 org.] 

= Spirontocaris japonica Yokoya, 1930: 533; Fig. 3. [Between Yuno-Shima isl. and Asamushi, 5-6 fms] 

= Spirontocaris fabricii var. minuta Urita, 1942: 25; Fig. 6. [Otomari, 2-3 fms] [nec Spirontocaris minuta 

Yokoya, 1930]


Eualus lindbergi Kobjakova, 1955: 240; Fig. 6. [near Levenorn, off Se Sakhalin, 110 m] 

Eualus lineatus Wicksten & Butler, 1983: 3; Figs 1-2. [1.5 mi. southwest of Gull Island, off Santa Cruz Island, 

California, 33°56'00"N 119°50'55"W, 89 m] 

Eualus macilentus (Krøyer, 1841) 

= Hippolyte macilenta Krøyer, 1841: 574 [Grønland] 

= Spirontocaris stoneyi Rathbun, 1902a: 899. [Bering Sea, 62°15'N 167°48'W, 20.5 fms] 

Eualus macrophthalmus (Rathbun, 1902a) 

= Spirontocaris macrophthalma Rathbun, 1902a: 900. [off Tawhit Head, Washington, 178 fms] 

Eualus middendorffi Bražnikov, 1907: 165; Fig. 23. [Sea of Okhotsk, off South Sakhalin Island] 

Eualus occultus (Lebour, 1936) 

= Spirontocaris occulta Lebour, 1936: 96; Plate 1; Plate 2, fig. 2. [Plymouth] 

? = Hippolyte Crassicornis H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 375. [la rade de 

Saint-Malo] 

? = Hippolyte Lovenii Rathke, 1843: 13. [Molde] 

? = Vianellia dorsiocullata Nardo, 1847: 8. [Lagune e Golfo Veneto] 

? = Hippolyte Yarrellii W. Thompson, 1853: 112; Plate 4, fig. 2. [Weymouth Bay] 

Eualus pax (Stebbing, 1915) 

= Spirontocaris pax Stebbing, 1915: 91; Plate 24. [34°11'N 18°31'E, 20 fms; off Buffels Bay (False Bay), 

30 fms] 

Eualus pectiniformis Hanamura, 2008: 88; Figs 1-3. [38°08.3'S 149°31.9'E, 150-189 m] 

Eualus pusiolus (Krøyer, 1841) 

= Hippolyte pusiola Krøyer, 1841: 576. [Norges Vestkyst] 

= Hippolyte subula Rathke, 1843: 9. [bei Molde] 

= Hippolyte vittata Rathke, 1843: 10. [Christiansund] 

= Hippolyte Barleei Spence Bate, 1852: 357; Plate 5B(1). [off the Haaf, Shetlands] 

= H.[ippolyte] Korenii Danielssen, 1859: 6. [Vadsøe paa Leerbund, 60 fms] 

= Hippolyte Andrewsii Kinahan, 1860a: 48. [Dalkey Sound] 

Eualus ratmanovi Makarov, 1941: 125. [Bering Sea, stns 40, 51 (1932); 10, 16, 18 (1933)] 

Eualus sinensis (Yu, 1931b) 

= Spirontocaris sinensis Yu, 1931b: 514; Fig. 2. [Chefoo] 

Eualus sollaudi (Zariquiey Cenarro, 1936) 

= Thor Sollaudi Zariquiey Cenarro, 1936: 242; Figs 17-21. [Cadaqués, Arenys] 

Eualus spathulirostris (Yokoya, 1933) 

= Spirontocaris spathulirostris Yokoya, 1933: 28; Fig. 10. [Sôyô-Maru stn 70, near middle between Siriya-zaki 

and Todo-saki, 285 m; 474, north-west of Is. Tusima, 219 m; 652, Tugaru Strait, 110 m] 

Eualus suckleyi (Stimpson, 1864) 

= Hippolyte Suckleyi Stimpson, 1864: 154. [Puget Sound, circumlittoral zone] 

Eualus subtilis Carvacho & Olson, 1984: 61; Figs 1-2. [Punta Banda, extremo sur de Bahía de Todos Santos, 

Baja California, 30 m] 

Eualus townsendi (Rathbun, 1902a) 

= Spirontocaris townsendi Rathbun, 1902a: 897. [Washington Sound, Straits of Fuca, 48 fms] 

Eumanningia Crosnier, 2000 

= Eumanningia Crosnier, 2000 (type species Eumanningia pliarthron Crosnier, 2000, by original designation 


Eumanningia pliarthron Crosnier, 2000: 110; Figs 1-4. [South-West Pacific, Wallis Island, 14°11.0'S 

176°17.3'W, 890-915 m] 

Exhippolysmata Stebbing, 1915 

= Exhippolysmata Stebbing, 1915 (type species Hippolysmata ensirostris Kemp, 1914, designated by 

Holthuis, 1955b, gender feminine) 

Exhippolysmata ensirostris ensirostris (Kemp, 1914) 

= Hippolysmata ensirostris Kemp, 1914: 113, 118; Plate 7, figs 1-4. [Colombo]


Exhippolysmata ensirostris punctata (Kemp, 1914) 

= Hippolysmata ensirostris var. punctata Kemp, 1914: 120; Plate 7, figs 5-7. [Sandheads, Ganges delta; 

Green Isl., Amherst, Tenasserim; Thongwa, Burma] 

Exhippolysmata hastatoides (Balss, 1914a) 

= Mimocaris hastatoides Balss, 1914a: 596. [Victoria, Cameroon, Reuse (Geringe tiefe)] 

Exhippolysmata oplophoroides (Holthuis, 1948) 

= Hippolysmata (Exhippolysmata) oplophoroides Holthuis, 1948: 1106; Figs 2-3. [Mouth of Surinam 

River, near Resolutie] 

Exhippolysmata tugelae Stebbing, 1915: 94; Plate 89. [Off South Head, Tugela River, 12 fms; Cape Henderson, 

NW 2.5 miles, 26 fms] 

Gelastocaris Kemp, 1914 

= Gelastocaris Kemp, 1914 (type species Latreutes Paronae Nobili, 1905f, by monotypy, gender feminine; 


Gelastocaris paronae (Nobili, 1905f) 

= Latreutes Paronae Nobili, 1905f: 2. [isola di Zanzibar] 

Gelastreutes Bruce, 1990h 

= Gelastreutes Bruce, 1990h (type species Gelastreutes crosnieri Bruce, 1990h, by original designation 


Gelastreutes crosnieri Bruce, 1990h: 139; Figs 1-4. [New Caledonia, 19°08'30"S 163°29'30"E, 65-120 m] 

Heptacarpus Holmes, 1900 

= Heptacarpus Holmes, 1900 (type species Hippolyte palpator Owen, 1839, by original designation, 


1957) 

Heptacarpus acuticarinatus Komai & Ivanov, 2008: 9; Figs 5-9. [Sagami Bay, 35°07.858'N 139°33.698'E, 100- 

101 m] 

Heptacarpus brachydactylus (Rathbun, 1902a) 

= Spirontocaris brachydactyla Rathbun, 1902a: 898. [off Santa Cruz Island, 266 fms] 

Heptacarpus brevirostris (Dana, 1852a) 

= Hippolyte brevirostris Dana, 1852a: 24. [in freto ”de Fuca”, juxta portum ”Dungeness”] 

Heptacarpus camtschaticus (Stimpson, 1860a) 

= Hippolyte camtschatica Stimpson, 1860a: 33. [Type locality not indicated] 

Heptacarpus carinatus Holmes, 1900: 202; Plate 3, fig. 60. [Monterey Bay, shallow water] 

Heptacarpus commensalis Hayashi, 1979: 14; Figs 1-2. [just north of Seto Marine Biological Laboratory, 

Shirahama, from Acropora sp.] 

Heptacarpus decorus (Rathbun, 1902a) 

= Spirontocaris decora Rathbun, 1902a: 896. [off Santa Cruz Island, California, 150 fms] 

Heptacarpus flexus (Rathbun, 1902a) 

= Spirontocaris flexa Rathbun, 1902a: 896. [north of Bird Island, Shumagins, Alaska, 21 fms] 

Heptacarpus franciscanus (Schmitt, 1921) 

= Spirontocaris franciscana Schmitt, 1921: 60; Plate 12, figs 8-9. [San Francisco Bay] 

Heptacarpus fuscimaculatus Wicksten, 1986: 47; Figs 1-2. [Big Fisherman’s Cove, Santa Catalina Island, 

California, 33°27'N 118°28'W, among low-growing algae on floating dock] 

Heptacarpus futilirostris (Spence Bate, 1888) 

= Nauticaris futilirostris Spence Bate, 1888: 606; Plate 109, fig. 1. [Challenger stn 233A, 34°38'N 

135°01'E, off Japan, 50 fms] 

Heptacarpus geniculatus (Stimpson, 1860a) 

= Hippolyte geniculata Stimpson, 1860a: 34. [Sinu ”Hakodadi”, inter lapides ad prof 2 org.] 

= Spirontocaris alcimede De Man, 1906: 404. [Inland sea of Japan] 

Heptacarpus grebnitzkii (Rathbun, 1902b) 

= Spirontocaris grebnitzkii Rathbun, 1902b: 44; Fig. 18. [Mororan, Hokkaido]


Heptacarpus herdmani (Walker, 1898) 

= Spirontocaris herdmani Walker, 1898: 277; Plate 16, fig. 2. [Puget Sound] 

Heptacarpus igarashii Hayashi & Chiba, 1989: 71; Figs 1-3. [Toni Bay, Kamaishi City, Iwate Prefecture] 

Heptacarpus jordani (Rathbun, 1902b) 

= Spirontocaris jordani Rathbun, 1902b: 44; Fig. 17. [Hakodata, Hokkaido] 

Heptacarpus kincaidi (Rathbun, 1902a) 

= Spirontocaris kincaidi Rathbun, 1902a: 899. [off Santa Cruz, California, 21 fms] 

Heptacarpus longirostris (Kobjakova, 1936a) 

= Eualus geniculata var. longirostris Kobjakova, 1936a: 211; Fig. 38. [Peter the Great Bay, Sea of Japan; 

fully described in Kobjakova, 1937] 

Heptacarpus maxillipes (Rathbun, 1902a) 

= Spirontocaris maxillipes Rathbun, 1902a: 898. [Off Segouam, Aleutians, 283 fms] 

Heptacarpus minutus (Yokoya, 1930) 

= Spirontocaris minuta Yokoya, 1930: 531; Fig. 2. [Off Arito, 19 fms] 

Heptacarpus moseri (Rathbun, 1902a) 

= Spirontocaris moseri Rathbun, 1902a: 897. [off Segouam, Aleutians, 283 fms] 

Heptacarpus palpator (Owen, 1839) 

= Hippolyte palpator Owen, 1839: 89; Plate 28, fig. 3. [Monteray, California] 

= Hippolyte? Hemphillii Lockington, 1877a: 35. [San Diego] 

Heptacarpus paludicola Holmes, 1900: 201; Plate 3, figs 56-57. [Humboldt Bay; Shelter Cove; Bodega Bay] 

Heptacarpus pandaloides (Stimpson, 1860a) 

= Hippolyte pandaloides Stimpson, 1860a: 34. [Sinu ”Hakodadi”, inter lapides ad prof 2 org.] 

= Spirontocaris propugnatrix De Man, 1906: 404. [Inland Sea of Japan, 6 fms] 

Heptacarpus pugettensis Jensen, 1983: 314; Figs 1-3. [Alki Point, Seattle, Washington, 47°34'N 122°25'W, 

low intertidal, under rock] 

Heptacarpus rectirostris (Stimpson, 1860a) 

= Hippolyte rectirostris Stimpson, 1860a: 33. [Portu ”Hakodadi” Japoniæ borealis, in locis profundis 

maris] 

Heptacarpus sitchensis (Brandt, 1851) 

= Hippolyte sitchensis Brandt, 1851: 116; Plate 5, figs 18-18c. [Insel Sitcha] 

= Hippolyte picta Stimpson, 1871: 125. [Monterey, Cal.] 

= Heptacarpus littoralis Butler, 1980: 220; unnumbered figure; Fig. 17. [Bunsby Islands, 2-9 m] 

Heptacarpus stimpsoni Holthuis, 1947a: 13. [nomen novum for Hippolyte cristata Stimpson, 1860a] 

= Hippolyte cristata Stimpson, 1860a: 33, nec De Haan, 1844 [in De Haan, 1833-1850]. [Portu ”San 

Francisco” Californiae, fundo arenoso prof. 5-10 org.] 

Heptacarpus stylus (Stimpson, 1864) 

= Hippolyte stylus Stimpson, 1864: 154. [Straits of De Fuca] 

= Hippolyte esquimaltiana Spence Bate, 1864: 666. [Esquimalt Harbor] 

Heptacarpus taylori (Stimpson, 1857) 

= Hippolyte taylori Stimpson, 1857: 500. [Monterey] 

Heptacarpus tenuissimus Holmes, 1900: 203. [Monterey] 

= Hippolyte gracilis Stimpson, 1864: 155; nec Liljeborg, 1851. [Puget Sound, deep water] 

= Hippolyte amabilis Lenz, 1901: 432; Plate 32, figs 2-3. [Bare Island] 

Heptacarpus tridens (Rathbun, 1902a) 

= Spirontocaris tridens Rathbun, 1902a: 896. [Admiralty Inlet, Puget Sound, 40 fms] 

Heptacarpus yaldwyni Wicksten, 1984b: 241; Figs 1-3. [off Salina Cruz, Mexico, 14°47'N 96°19'W – 

14°50.5'N 96°13'W, 1052-1145 m] 

Hippolyte leach, 1814 [in leach, 1813-1814] 

= Hippolyte Leach, 1814 [in Leach, 1813-1814] (type species Hippolyte Varians Leach, 1814 [in Leach, 




= Nectoceras Rafinesque, 1817 (type species Nectoceras pelagica Rafinesque, 1817, by monotypy, gender 

neuter, a junior subjective synonym of Astacus coerulescens Fabricius, 1775; name placed on the 

Official Index of Rejected and Invalid Names in Zoology in Opinion 522 in 1958) 

= Virbius Stimpson, 1860a (type species Hippolyte acuminatus Dana, 1852a (junior subjective synonym 

of Astacus coerulescens Fabricius, 1775), designated by Kingsley, 1880, gender masculine) 

= Bellidia Gosse, 1877 (type species Bellidia Huntii Gosse, 1877, by monotypy, gender feminine) 

Hippolyte acuta (Stimpson, 1860a) 

= Virbius acutus Stimpson, 1860a: 35. [ad insulam ”Loo Choo”, littoralis in rupibus algosis] 

Hippolyte australiensis (Stimpson, 1860a) 

= Virbius australiensis Stimpson, 1860a: 35. [in portu Jacksoniensi Australiae, inter algas ad prof. org. 2] 

= Caradina cincinnuli Spence Bate, 1863: 500; Plate 40, figs 3-3z. [Gulf St. Vincent, 4.5 fms] 

Hippolyte bifidirostris (Miers, 1876a) 

= Virbius bifidirostris Miers, 1876a: 224. [New Zealand; illustrated in Miers, 1876b] 

Hippolyte californiensis Holmes, 1895: 576; Plate 20, figs 21-26. [Bodega Bay] 

Hippolyte caradina Holthuis, 1947a: 14. [nomen novum for Caradina tenuirostris Spence Bate, 1863 nec 

Hippolyte tenuirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]] 

= Caradina tenuirostris Spence Bate, 1863: 501; Plate 40, figs 4-4k. [Gulf St. Vincent, 4.5 fms] 

Hippolyte catagrapha d’Udekem d’Acoz, 2007: 185; Figs 1-4. [South Africa, False Bay, 6-8 m, from the crinoid 

Tropiometra carinata] 

Hippolyte clarki Chace, 1951: 37; Fig. 1f-p. [Friday Harbor, Washington, in eel grass] 

Hippolyte coerulescens (Fabricius, 1775) 

= Astacus coerulescens Fabricius, 1775: 414. [Pelago inter Tropicos frequens] 

= Palaemon pelasgicus Bosc, 1802: 105; Plate 14, fig. 2. [la haute mer, sur fucus nageans] 

= Nectoceras pelagica Rafinesque, 1817: 41. [Atlantic Ocean, in the gulph stream on the Fucus natans] 

= Hippolyte tenuirostris H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 374. [en haute 

mer, près de Açores] 

= Hippolyte acuminatus Dana, 1852a: 24. [in mari Atlantico cum Sargasso lat. bor. 36°07' - 4°07', long. 

occid. 20°43' - 71°36'] 

= Hippolyte bidentatus Spence Bate, 1888: 591; Plate 105, figs 1-2. [Atlantic Ocean, on gulf-weed; between 

32°07'-35°29'N and 50°53'-52°32'W] 

= Hippolyte Martiali A. Milne-Edwards, 1891: 47; Plate 6, figs 1-1g. [canal du Beagle, en vue de 

Loupataya, 198 m] 

Hippolyte edmondsoni Hayashi, 1981a: 185; Figs 1-3. [Waimanalo, Oahu] 

Hippolyte garciarasoi d’Udekem d’Acoz, 1996b: 16; Figs 5-6, 8-10. [Banyuls, between algae on rocks at the 

laboratory Arago, 42°29'N 03°08'E, 0-1 m] 

Hippolyte holthuisi Zariquiey Álvarez, 1953: 104; Figs 3-4, 6, 8. [Cadaqués, junto al Cabo de Creus, 22-60 m] 

Hippolyte inermis Leach, 1816b: 347. [Cornwall (United Kingdom), Plymouth, 50°13'N 04°10'W; neotype 

selection by d’Udekem d’Acoz, 1996b] 

? = Carida viridis Rafinesque, 1814: 24. [Sicile; nomen nudum] 

= P.[alemon] Olivieri Risso, 1816: 107. [dans les fucus et les conferves, environs de Nice] 

= P.[alemon] Margaritaceus Risso, 1816: 108. [dans les endroits pierreux, environs de Nice] 

= Alpheus viridis Otto, 1821: 12. [in mari circa Nicaeam; fully described in Otto, 1828] 

= A.[lpheus] elongatus Risso, 1827: 77. [golfe de Nice, dans les fucus] 

= Hippolytus Brullei Guérin, 1832: 41; Plate 27, fig. 2. [Grèce] 

= Hippolyte virescens H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: Plate 53, figs 3-3b 

(as Hippolyte viridis). [Type locality not indicated] 

= Hippolyte mauritanicus Lucas, 1846: 42; Plate 4, fig. 3. [rade de Mers-el-Kebir; Lager, Stora, Bône] 

= Hippolyte Whitei W. Thompson, 1853: 110; Plate 6, figs 1-1a. [Weymouth Bay, 4-6 fms] 

= Hippolyte Whitei var. ensis W. Thompson, 1853: 111; Plate 6, fig. 1b. [Weymouth Bay] 

= Hippolyte Whitei var. falcatus W. Thompson, 1853: 112; Plate 6, fig. 1c. [Weymouth Bay] 

= Hippolyte Mitchelli W. Thompson, 1853: 114; Plate 6, figs 4-4a. [Weymouth Bay] 

= Hippolyte producta Norman, 1861: 275; Plate 13, fig. 5. [Lamlash Bay] 

Hippolyte jarvinensis Hayashi, 1981a: 190; Figs 4-5. [Jarvis Islands, the Line Islands]


Hippolyte kraussiana (Stimpson, 1860a) 

= Virbius Kraussianus Stimpson, 1860a: 36. [in sinu ”Simons Bay” prope promontorium Bonae Spei] 

= Virbius capensis Lenz & Strunck, 1914: 319; Plate 20, figs 1-4. [Simonsbai] 

Hippolyte lagarderei d’Udekem d’Acoz, 1995: 494; Fig. 1. [Maroc, côte Atlantique, Témara, 33°55'N 

06°55'W] 

Hippolyte leptocerus (Heller, 1863a) 

= Virbius leptocerus Heller, 1863a: 289; Plate 10, Figs 5-6. [Hafen von Genua] 

= V.[irbius] gracilis Heller, 1862b: 399; nec H.[ippolyte] gracilis Liljeborg, 1851. [aus dem adriatischen 

Meere] 

= Virbius gracilis varietas intermedia Czerniavsky, 1868: 68; Plate 5, figs 2-7. [sinu Jaltensi] 

= Virbius gracilis var. longirostris Czerniavsky, 1868: 68; Plate 5, figs 8-25. [Black Sea] 

= Virbius gracilis forma typica Czerniavsky, 1884: 14. [Black Sea] 

= Virbius gracilis var. articulirostris Czerniavsky, 1884: 15; Plate 1, fig.1a. [Black Sea] 

= Virbius gracilis var. brevirostris Czerniavsky, 1884: 15; Plate 1, figs 1p-s. [Black Sea] 

= Virbius Brullei var. elongata Czerniavsky, 1884: 18. [sinus Jaltensis, zona littoralis; sinus Sevastopol] 

= Virbius Brullei forma fortior Czerniavsky, 1884: 19; Plate 2, figs 3a-n. [sinus Jaltensis, zona littoralis, 

1-1.5 m; sinus Suchum, zona littoralis, 1-1.5 m] 

= Virbius tenuirostris Czerniavsky, 1884: 20; Plate 2, figs 4a-g. [sinus Suchum, zona littoralis, 1-1.5 m] 

= Virbius rectifrons Czerniavsky, 1884: 21; Plate 1, fig. 2. [sinus Suchum, zona littoralis, 1-1.5 m] 

= Hippolyte longirostris armoricana Sollaud in Bourdon, 1965: 6, 39. [côtes bretonnes] 

Hippolyte leptometrae Ledoyer, 1969: 343; Plate 2. [côtes de Provence, 110 m] 

Hippolyte longiallex d’Udekem d’Acoz, 2007: 193; Figs 7-9. [West Africa, Gulf of Guinea, Principe Island, 

Pedra da Galé, 35 m, from Muriceops tuberculata] 

Hippolyte multicolorata Yaldwyn, 1971: 90. [Island Bay, Wellington, from intertidal algae] 

Hippolyte nicholsoni Chace, 1972: 113; Figs 46-47. [Milford Bay, between Pigeon Point and Crown Point, 

Tobago, 9-12 m] 

Hippolyte niezabitowskii d’Udekem d’Acoz, 1996b: 58; Figs 25-28, 29a. [Ionian coast of Greece, Drepanos, 

western part, 39°30'N 20°15'E, small seagrasses on non muddy sand] 

Hippolyte obliquimanus Dana, 1852a: 24. [in portu Rio Janeiro] 

= Hippolyte exilirostratus Dana, 1852a: 24. [in portu ”Rio Janeiro”] 

= Virbius gracilis var. brasiliensis Czerniavsky, 1884: 14. [Type locality not indicated] 

= Hippolyte curaçaoensis Schmitt, 1924a: 68; Fig. 4. [West Punt, Curaçao] 

Hippolyte palliola Kensley, 1970: 183; Figs 1-2. [4 or 5 miles south of the Kunene River mouth, South West 

Africa, in brown algae growing on intertidal rocks] 

Hippolyte pleuracantha (Stimpson, 1871) 

= Virbius pleuracanthus Stimpson, 1871: 127. [harbor of Norfolk, Va.; at Somers’ Point, in Great Egg 

Harbor, N. J.] 

Hippolyte prideauxiana Leach, 1817 [in Leach, 1815-1875] 

= Hippolyte Prideauxiana Leach, 1817 [in Leach, 1815-1875]: Plate 38, figs 1, 3-5. [Near Bantham, on 

the southern coast of Devon] 

= Hippolyte Moorii Leach, 1817 [in Leach, 1815-1875]: Plate 38, fig. 2. [Plymouth Sound] 

= Bellidia Huntii Gosse, 1877: 313; Plate 10. [off the Shag Rock, northern end of Torbay, 6 fms] 

Hippolyte proteus (Paul’son, 1875) 

= V.[irbius] Proteus Paul’son, 1875: 109, Plate 16, figs 2-5, Plate 18, figs 1-1k. [Red Sea] 

Hippolyte sapphica d’Udekem d’Acoz, 1993: 56; Figs 1, 5, 7, 8-9. [20 m à l’est du port de Skala Kallonis, 

herbiers de Zostéracees, 0.5-1 m] 

= Hippolyte sapphica forma A d’Udekem d’Acoz, 1996b: 79; Figs 38-39. [Gulf of Kallonis, 20 m at East 

of Skala Kallonis harbour, 39°12'N 26°15'E, 0.5-1 m] 

= Hippolyte sapphica forma B d’Udekem d’Acoz, 1996b: 84; Figs 40-43. [Gulf of Amvrakikos] 

Hippolyte varians Leach, 1814 [in Leach, 1813-1814] 

= Hippolyte Varians Leach, 1814 [in Leach, 1813-1814]: 432. [rocky shores of Devon] 

= Hippolyte smaragdina Krøyer, 1841: 570. [Norges Vestkyst og Kattegattet] 

= Hippolyte fascigera Gosse, 1853: 153. [Weymouth Bay, a few miles from land]


= Hippolyte Grayana W. Thompson, 1853: 113; Plate 6, figs 3-3a. [Weymouth Bay, 4 fms] 

= Caradina tenuis Spence Bate, 1866: 28; Plate 2, fig. 1. [Plymouth] 

Hippolyte ventricosa H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840] 

= Hippolyte ventricosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 371. [mers d’Asie; 

India according to d’Udekem d’Acoz, 1999] 

= H.[ippolyte] orientalis Heller, 1862c: 277. [im rothen Meere] 

? = Virbius Mossambicus Hilgendorf, 1879: 836; Plate 4, fig. 1. [Zambeze] 

Hippolyte williamsi Schmitt, 1924c: 163; Fig. 40. [Eden, rock pools] 

= Hippolyte mexicana Chace, 1937b: 127; Fig. 6. [Santa Inez Bay, 26°50'45''N 111°54'20''W, 1 fm] 

Hippolyte zostericola (Smith, 1873) 

= Virbius zostericola Smith, 1873: 550; Plate 3, fig. 11. [among eel-grass about Vineyard Sound] 

= Hippolyte pleuracantha bermudensis Gurney, 1936: 27; Plate 1, figs 4-12; Plate 2, figs 13-21. [Zosterabed 

in Walsingham Bay, on the west of Castle Harbour; in Sargassum growing on rocks at Tobacco 

Bay on the north shore] 

Latreutes Stimpson, 1860a 

= Cyclorhynchus De Haan, 1849 [in De Haan, 1833-1850] (type species Hippolyte planirostris De Haan, 

1844 [in De Haan, 1833-1850], by monotypy, gender neuter; invalid junior homonym of Cyclorhynchus 

Kaup, 1829 (Aves), Cyclorhynchus Sundevall, 1836 (Aves) and Cyclorhynchus Macquart, 1841 

(Diptera)) 

= Latreutes Stimpson, 1860a (type species Hippolyte ensiferus H. Milne Edwards, 1837 [in H. Milne 

Edwards, 1834-1840], designated by Kingsley, 1880, gender masculine; name placed on the Official 

List of Generic Names in Zoology in Opinion 470 in 1957) 

= Rhynchocyclus Stimpson, 1860a (nomen novum for Cyclorhynchus De Haan, 1849 [in De Haan, 1833- 

1850], gender masculine; invalid junior homonym of Rhynchocyclus Cabanis & Heine, 1859 (Aves)) 

= Concordia Kingsley, 1880 (type species Concordia gibberosus Kingsley, 1880 (junior subjective synonym 

of Rhynchocyclus parvulus Stimpson, 1871), by monotypy, gender feminine) 

= Platybema Spence Bate, 1888 (nomen novum for Cyclorhynchus De Haan, 1849 [in De Haan, 1833- 

1850] and Rhynchocyclus Stimpson, 1860a, gender neuter) 

Latreutes acicularis Ortmann, 1890: 506; Plate 37, fig. 6. [Japan, Kadsiyama] 

Latreutes anoplonyx Kemp, 1914: 104; Plate 4, figs 3-5. [Bombay] 

Latreutes antiborealis Holthuis, 1952d: 62; Fig. 14. [Golgo de Ancud, Canal San Antonio, inner part, 

41°44'10"S 73°15'15"W, 15 m] 

Latreutes compressus (Stimpson, 1860a) 

= Rhynchocyclus compressus Stimpson, 1860a: 28. [in portu ”Jackson” Australiae, f. algoso p. 2 org.] 

= Caradina truncifrons Spence Bate, 1863: 499; Plate 40, figs 2-2u. [Gulf St. Vincent, 4.5 fms] 

Latreutes foliirostris Kobjakova, 1935: 91; Fig. 4. [Bucht Peters des Großen] 

Latreutes fucorum (Fabricius, 1798) 

= Palaemon fucorum Fabricius, 1798: 404. [in Oceani Fuco natante] 

= Hippolyte ensiferus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 374. [haute mer, 

près de Açores] 

Latreutes inermis Chace, 1972: 122; Figs 51-52. [Reef just south of Marigot Harbour, Saint Lucia Island, 4-6 m] 

Latreutes laminirostris Ortmann, 1890: 506; Plate 37, fig. 5. [Japan, Tanagava] 

Latreutes mucronatus (Stimpson, 1860a) 

= Rhynchocyclus mucronatus Stimpson, 1860a: 27. [in freto ”Ly-I-moon” prope Hong Kong, f. conchoso 

p. 25 org] 

= Latreutes Gravieri Nobili, 1904: 231. [Djibouti] 

= Latreutes mucronatus var. multidens Nobili, 1905b: 394. [Mer Rouge, Java] 

= Latreutes natalensis Lenz & Strunck, 1914: 320; Plate 21, figs 1-11. [Port Natal] 

Latreutes parvulus (Stimpson, 1871) 

= Rhynchocyclus parvulus Stimpson, 1866: 48. [nomen nudum] 

= Rhynchocyclus parvulus Stimpson, 1871: 124. [St. Joseph I., Texas] 

= Concordia gibberosa Kingsley, 1880: 414. [Fort Macon (Beaufort Inlet), North Carolina, U.S.A.]


Latreutes phycologus Nobili, 1905a: 159; unnumbered figure. [Côtes d’Arabie, sur une algue brune flottante] 

Latreutes planirostris (De Haan, 1844 [in De Haan, 1833-1850]) 

= Hippolyte planirostris De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 7. [Japan] 

= Latreutes dorsalis Stimpson, 1860a: 27. [in sinu ”Hakodadi” Japoniae, prof. 8 org.] 

Latreutes planus Spence Bate, 1888: 584; Plate 89, fig. 5. [Off Sibago Island, Samboangan, Philippine Islands] 

Latreutes porcinus Kemp, 1916a: 397; Fig. 3; Plate 36, fig. 3. [jetty on Ross Island, Port Blair, Andamans, 

among weeds] 

Latreutes pristis (Nobili, 1899) 

= Platybema pristis Nobili, 1899: 233. [Beagle Bay, sulle secche fra Beagle e Volverine Passage] 

Latreutes pymoeus Nobili, 1904: 231. [Djibouti; original spelling maintained under Art. 32.1] 

Latreutes unidentatus Spence Bate, 1888: 586; Plate 89, fig. 6. [Off Sibago, Samboangan, Philippine Islands] 

Lebbeus White, 1847a 

= Lebbeus White, 1847a (type species Lebbeus orthorhynchus White, 1847a (junior subjective synonym 

of Alpheus Polaris Sabine, 1824), by monotypy, gender masculine; name conserved under the plenary 

power of the ICZN, and placed on the Official List of Generic Names in Zoology in Opinion 

671 in 1963) 

= Hetairus Spence Bate, 1888 (type species Alpheus Polaris Sabine, 1824, designated under the plenary 

powers of the ICZN, gender masculine; name placed on the Official List of Rejected and Invalid 


= Birulaecaris Dons, 1915 (type species Hippolyte mysis Birulya, 1898 (junior subjective synonym of 

Alpheus Polaris Sabine, 1824), gender feminine) 

Lebbeus acudactylus Jensen, 2006: 34; Figs 4D, 7-9. [Aleutian Islands, Alaska, 51.86°N 174.92°W, 155 m] 

Lebbeus africanus Fransen, 1997a: 231; Figs 1-29. [Mauritania, off Banc d’Arguin, 19°43'N 17°30'W, 1500 

m] 

Lebbeus antarcticus (Hale, 1941) 

= Spirontocaris antarcticus Hale, 1941: 267; Figs 5-6. [Off Adelie Coast, Wilkes Land, Antarctica, 

66°21'S 138°28'E, 640 m] 

Lebbeus balssi Hayashi, 1992: 112; Figs 1-3. [East China Sea, 33°59.4'N 128°48.0'E, 102 m] 

Lebbeus bidentatus Zarenkov, 1976: 13; Fig. 5. [R.V. Akademik Kurchatov stn 276, off Peru, 1680 m] 

Lebbeus brandti (Bražnikov, 1907) 

= Hetairus brandti Bražnikov, 1907: 157; Fig. 20. [near Cape Terpeniya, East coast of Sakhaln Island, 

Sea of Okhotsk] 

Lebbeus brevicornis Komai, 2011: 291; Figs 5-8. [R/V Rinkai-maru, Sagami Bay, off Misaki, 351-338 m] 

Lebbeus brevirostris Chang, Komai & Chan, 2010: 727; Figs 1-3, 10A-B. [24°52.33'N 122°03.10'E, 600 m] 

Lebbeus carinatus Zarenkov, 1976: 9; Fig. 2. [R.V. Akademik Kurchatov stn 290, off Peru, 1850 m] 

Lebbeus catalepsis Jensen, 1987: 89; Figs 1-3. [Strait of Juan de Fuca between Sekiu and Neah Bay, Washington, 

48°19'N 124°28'W, low intertidal] 

Lebbeus clarehannah McCallum & Poore, 2010: 127; Figs 1C, 2-3. [Western Australia, off Bald Island, 

35°12.49'S 118°39.04'E - 35°12.14'S 118°40.08'E, 408-431 m] 

Lebbeus comanthi Hayashi & Okuno, 1997: 49; Figs 1-2, 3(2), 4(3). [Izu Ocean Park, Sagami Bay, 15 m, from 

Oxycomanthus japonicus] 

Lebbeus compressus Holthuis, 1947a: 9, 40. [nomen novum for Spirontocaris gibberosa Yokoya, 1933 sensu 

Balss, 1914b] 

= Spirontocaris gibberosa Yokoya, 1933: 25; Fig. 8. [Sôyô-Maru stn 26, north-east of Siwoya-zaki, 232 m] 

Lebbeus cristagalli McCallum & Poore, 2010: 131; Figs 1A-B, 4. [Western Australia, north-west Australia, 

near Ashmore Reef, 12°28.53'S 123°25.04'E – 12°29.58'S 123°25.00'E, 397-405 m] 

Lebbeus cristatus Ahyong, 2010: 341; Figs 1-2, 3A. [36°54.76'S 167°32.10'E, Challenger Plateau, New Zealand, 

1231-1226 m]


Lebbeus curvirostris Zarenkov, 1976: 12; Fig. 4. [R.V. Akademic Kurchatov stn 290, off Peru, 1680-1860 m] 

Lebbeus elegans Komai, Hayashi & Kohtsuka, 2004: 103; Figs 1-3. [off Togi, Hakui District, Ishikawa Prefecture, 

west coast of Noto Peninsula, Sea of Japan, 400 m] 

Lebbeus eludus Jensen, 2006: 24; Figs 1-3, 4A-B. [Burrows Channel, Anacortes, Washington, 48.29.4°N 

122.41.7°W, 14 m] 

Lebbeus fasciatus (Kobjakova, 1936a) 

= Hetairus fasciata Kobjakova, 1936a: 222; Fig. 17. [Okhotsk Sea; fully described in Kobjakova, 1937] 

= Hetairus zebra Makarov, 1935: 319; Fig. 1; nec Leim, 1921. [Bering-Insel, Bucht Nikolskaja, sandiges 

Ufer am Ende der Bucht; Bering-Insel, Fazies der Steine und Felsen im Litoral nahe dem Dorf Nokolskaja; 

Bering-Meer, 59°53'N 170°26'E, 32 m; Awatsch-Golf, nahe der Bucht Betschewinskaja, 30 m] 

= Hetairus zebra Makarov, 1941: 123. [Karaginsky Island, Bering Island, Mednyi Island, Avanchisky 

Gulf (near Becheviskaya Bay), Bering Sea stn 85] 

= Spirontocaris makarofi Urita, 1942: 18. [nomen novum for Hetairus zebra Makarov, 1935] 

Lebbeus formosus Chang, Komai & Chan, 2010: 732; Figs 4-5, 10C. [22°06.3345'N 121°08.2237'E, 1260-1275 

m] 

Lebbeus grandimanus (Bražnikov, 1907) 

= Hetairus grandimana Bražnikov, 1907: 152; Fig. 18. [coasts of Kamtchatka, Sea of Okhotsk; Sea of 

Japan, 19-118 m] 

Lebbeus groenlandicus (Fabricius, 1775) 

= Astacus Groenlandicus Fabricius, 1775: 416. [in mari groenlandico] 

= Cancer aculeatus O. Fabricius, 1780: 239. [Naularnak] 

= Hippolite armata Owen, 1839: 88; Plate 27, fig. 2. [shores of Kamtschatka] 

= Hippolite cornuta Owen, 1839: 89; Plate 28, fig. 2. [Type locality not indicated] 

Lebbeus heterochaelus (Kobjakova, 1936a) 

= Hetairus heterochaela Kobjakova, 1936a: 222; Figs 18-19. [Okhotsk Sea, 165 m; fully described in 

Kobjakova, 1937] 

Lebbeus indicus Holthuis, 1947a: 40; Figs 1-3. [Bali Sea, 7°28.2'S 115°24.6'E, 1018 m] 

Lebbeus kuboi Hayashi, 1992: 123; Figs 6-8. [central Sea of Japan, off Namerikawa, Toyama Prefecture, 200 

m] 

Lebbeus laevirostris Crosnier, 1999a: 454; Figs 1-3. [Indonésie (détroit de Makassar), 02°37.6'S 118°10.9'E, 

798-779 m] 

Lebbeus lagunae (Schmitt, 1921) 

= Spirontocaris lagunae Schmitt, 1921: 57; Fig. 35; Plate 12, Figs 10-11. [Laguna Beach, California, 12-15 

fms] 

Lebbeus laurentae Wicksten, 2010: 196; Figs 1-4; nomen novum for Lebbeus carinatus de Saint Laurent, 1984 

nec Zarenkov, 1976. [Northeastern Pacific, 12°49'N 103°57'W, 2630 m] 

Lebbeus longidactylus (Kobjakova, 1936a) 

= Hetairus longidactyla Kobjakova, 1936a: 222; Figs 12-13. [Okhotsk Sea, 440-504 m; fully described in 


Lebbeus longipes (Kobjakova, 1936a) 

= Hetairus longipes Kobjakova, 1936a: 222; Fig. 16. [Peter the Great Bay and Tatar Strait, Sea of Japan; 


Lebbeus manus Komai & Collins, 2009: 29; Figs 1-3. [Manus Basin, 03°79'S 146°09'E, 1575 m] 

Lebbeus microceros (Krøyer, 1841) 

= Hippolyte microceras Krøyer, 1841: 578. [Grønland; microceras is an incorrect original spelling under 

Art. 33.3.1 and corrected to microceros] 

= Spirontocaris zebra Leim, 1921: 133; Plates 2-3. [Passamquoddy bay, 30 m; Joe’s Point, St. Croix 

River; Head Harbour, Campobello Island, low tide and 10 m (all New Brunswick); St. Mary’s Bay 

(Nova Scotia)] 

Lebbeus miyakei Hayashi, 1992: 127; Figs 10-11. [Sea of Genkainada, Orono-shima Island, Fukuoka Prefecture, 

30-40 m] 

Lebbeus mundus Jensen, 2006: 30; Figs 4C, 5-6. [Willis Point, Vancouver Island, British Columbia, Canada, 

48.34.601°N 123.29.319°W, 9-12 m]


Lebbeus nudirostris Komai & Takeda, 2004: 78; Figs 1A, 2-3. [west of Taibusa-misaki, Boso Peninsula, 

35°04.68'N 139°45.32'E, 250 m, trap for scampi] 

Lebbeus polaris (Sabine, 1824) 

= Alpheus polaris Sabine, 1824: ccxxxviii; Plate 2, figs 5-8. [on the coast of Melville Island, 50 fms] 

= Hippolite borealis Ross, 1835: lxxxiv; Plate B, fig. 3. [off Elizabeth Harbour, 80 fms] 

= Lebbeus orthorhynchus White, 1847a: 76. [Type locality not indicated] 

= Hippolyte St. Pauli Brandt, 1851: 118; Plate 5, fig. 19. [Beeringschen Meere bei Pauls-Insel] 

= Hippolyte cultellata Norman, 1867: 200. [the Minch] 

= Hippolyte incerta Buchholz, 1874: 272. [Ostgrönland] 

= Hippolyte Amazo Pfeffer, 1886: 46; Plate 1, figs 7a-b. [Cumberland-Sund, Baffins Land, 66°35'40''N 

67°19'15''W] 

= Hetarius debilis Spence Bate, 1888: 615; Plate 109, fig. 4. [Challenger stn 49, 43°3'N 63°39'W, 85 fms] 

= Hetairus tenuis Spence Bate, 1888: 613; Plate 109, fig. 3. [Challenger stn 49, 43°3'N 63°39'W, south of 

Halifax, Nova Scotia, 85 fms] 

= Hippolyte projecta Spence Bate, 1888: 594; Plate 105, fig. 3. [Challenger stn 49, 43°3'N 63°39'W, south 

of Halifax, Nova Scotia, 85 fms] 

= Hippolyte mysis Birulya, 1898: 184; Plate 1, figs 1-15. [White Sea, SW coast of Kandalak Bay, Kovda] 

Lebbeus polyacanthus Komai, Hayashi & Kohtsuka, 2004: 109; Figs 4-6. [off Togi, Hakui District, west 

coast of Noto Peninsula, 400 m] 

Lebbeus profundus (Rathbun, 1906) 

= Spirontocaris profunda Rathbun, 1906: 914; Plate 24, fig. 10. [vicinity of Modu Manu, 762-1000 fms] 

Lebbeus rubrodentatus Bruce, 2010k: 75; Figs 1-2. [9°35'S 129°28'E, Timor Sea, NW Australia, 360-396 m] 

Lebbeus saldanhae (Barnard, 1947) 

= Spirontocaris saldanhae Barnard, 1947: 385. [off Saldanha Bay, 145 fms] 

Lebbeus schrencki (Bražnikov, 1907) 

= Hetairus schrenki Bražnikov, 1907: 161; Fig. 21. [East coast of Sakhalin Island, Sea of Okhotsk, 43-100 m] 

Lebbeus scrippsi Wicksten & Méndez G., 1982: 106; Plates 1-2. [Off Arica, Chile, 18°40.5'S 70°36.0'W to 

18°32.2'S 70°29.8'W, 768-968 m] 

Lebbeus similior Komai & Komatsu, 2009: 540; Figs 10-12. [off Soh-ma, Fukushima Prefecture, 37°47.6'N 

142°37.1'E to 37°47.4'N 142°37.2'E, 1196-1196 m] 

Lebbeus speciosus (Urita, 1942) 

= Spirontocaris makarofi speciosa Urita, 1942: 19; Fig. 4. [Otomari, 2-3 fms] 

= Lebbeus possjeticus Kobjakova, 1967: 235; Fig. 4. [Minosok Bay, Possjet Gulf, Sea of Japan, Russian 

Far East] 

Lebbeus spinirostris (Kobjakova, 1936a) 

= Hetairus spinirostris Kobjakova, 1936a: 222; Fig. 10. [Okhotsk Sea; fully described in Kobjakova, 

1937] 

Lebbeus splendidus Wicksten & Méndez G., 1982: 110; Plates 3-5. [Southwest of Lobos de Tierra, Peru, 

6°31'S 81°01'W, 712-744 m] 

Lebbeus spongiaris Komai, 2001a: 57; Figs 1-4. [W of Izu-Oshima Island, Izu Islands, 34°44.538'N 

139°19.723'E, 257-263 m] 

Lebbeus tosaensis Hanamura & Abe, 2003: 17; Figs 1-5. [33°17'34''8N 134°13'07''E, 344 m] 

Lebbeus unalaskensis (Rathbun, 1902a) 

= Spirontocaris unalaskensis Rathbun, 1902a: 895. [north of Unalaska, 350 fms] 

= Hetairus brevipes Kobjakova, 1936a: 222; Figs 9a-c. [Okhotsk Sea; fully described in Kobjakova, 

1937] 

= Hetairus unalaskensis var. japonica Kobjakova, 1936a: 222; Fig. 14. [Tatar Straits, Sea of Japan; fully 

described in Kobjakova, 1937] 

= Hetairus unalaskensis var. ochotensis Kobjakova, 1936a: 222; Fig. 15. [Okhotsk Sea; fully described in 


Lebbeus unguiculatus Chang, Komai & Chan, 2010: 737; Figs 7-9, 10D. [22°09.556'N 121°07.174'E, 1257- 

1262 m]


Lebbeus ushakovi (Kobjakova, 1936a) 

= Hetairus ushakovi Kobjakova, 1936a: 222; Fig. 11. [Okhotsk Sea, 165 m; fully described in Kobjakova, 

1937] 

Lebbeus vicinus vicinus (Rathbun, 1902a) 

= Spirontocaris vicina Rathbun, 1902a: 895. [north of Unalaska Island, 309 fms] 

Lebbeus vicinus montereyensis Wicksten & Méndez G., 1982: 114; Plate 6. [West of Punta Banda, Baja California, 

Mexico, 31°18'N 117°36'W, 2068-2086 m] 

Lebbeus vinogradowi Zarenkov, 1960: 346; Fig. 3. [Sea of Okhotsk, 56°57.5'N 145°57'E, 204 m] 

Lebbeus washingtonianus (Rathbun, 1902a) 

= Spirontocaris washingtoniana Rathbun, 1902a: 895. [off Sea Lion Rock, Washington, 685 fms] 

Lebbeus wera Ahyong, 2009: 786; Figs 5-7. [Brothers Caldera, 34°52.93'S 179°04.13'E, 1208-1209 m] 

Lebbeus yaldwyni Kensley, Tranter & Griffin, 1987: 304; Figs 15-17; frontispiece (B). [New South Wales, 

east of Sydney, 33°43'S 151°51 – 53'E, 450 m] 

Leontocaris Stebbing, 1905 

= Leontocaris Stebbing, 1905 (type species Leontocaris paulsoni Stebbing, 1905, by monotypy, gender 


= Problemacaris Stebbing, 1921a (type species Problemacaris spinetum Stebbing, 1921a (junior subjective 

synonym of Leontocaris paulsoni Stebbing, 1905), by monotypy, gender feminine) 

Leontocaris alexander Poore, 2009: 958; Figs 2-4. [Seamount Hill U, 147°10.8'S 44°19.6'E, 1200-1300 m] 

Leontocaris amplectipes Bruce, 1990i: 121; Figs 1-5. [Victoria, South of Point Hicks, 38°21.9'S 149°20.0'E, 

1000 m] 

Leontocaris bulga Taylor & Poore, 1998: 59; Figs 2-4. [Tasmania, 94.5 km SSE of Southeast Cape, ”V” seamount, 

44°24.0'S 147°09.0'E, 1400-1650 m] 

Leontocaris lar Kemp, 1906b: 299. [outside the Porcupine Bank, 53°07'N 14°50'W, 500 fms] 

? = Problemacaris boschmai Gordon, 1964: 337; Figs 3-6, 9B. [48°03'N 9°04'W, 500-0 m] 

Leontocaris pacifica Zarenkov, 1976 

= Leontocaris pacificus Zarenkov, 1976: 10; Fig. 3. [off Chile, 680-700 m] 

Leontocaris paulsoni Stebbing, 1905: 99; Plate 26. [25 miles off Lion’s Head, N. 67°E, 131-136 fms] 

? = Problemacaris spinetum Stebbing, 1921a: 626. [about 300 fms, Table Mountain E. by S. 40 miles 

according to Stebbing, 1924] 

Leontocaris vanderlandi Fransen, 2001: 57; Figs 1-26. [Seychelles, N of Platte Island atoll, 05°48'S 55°22'E, 

600 m] 

Leontocaris yarramundi Taylor & Poore, 1998: 64; Figs 5-7. [Tasmania, 82.8 km SSE of Southeast Cape, ”U” 

seamount, 44°19.2'S 147°07.2'E, 1083-1448 m] 

Ligur Sarato, 1885 

= Lybia Risso, 1844 (type species P.[alaemon] Ensiferus Risso, 1816, by monotypy, gender feminine; 

invalid junior homonym of Lybia H. Milne Edwards, 1834 (Crustacea Brachyura)) 

= Ligur Sarato, 1885 (type species Ligur Edwardsii Sarato, 1885 (junior subjective synonym of 

P.[alaemon] Ensiferus Risso, 1816), by monotypy, gender masculine; name placed on the Official List 

of Generic Names in Zoology in Opinion 470 in 1957) 

Ligur ensiferus (Risso, 1816) 

= P.[alaemon] Ensiferus Risso, 1816: 106. [dans la région de coraux, environs de Nice] 

= Hippolyte Carneus Roux, 1831: 28. [nomen nudum] 

= Hippolytus Incarnatus Hope, 1851: 18. [nomen nudum] 

= Palemon Vedianti Monod, 1931: 133. [nomen nudum] 

= Ligur Edwardsii Sarato, 1885: 2. [mers de Saint-Jean et d’Eza] 

Lysmata Risso, 1816 

= Aglaope Rafinesque, 1814 (type species Aglaope striata Rafinesque, 1814 (invalid senior subjective 

synonym of M.[elicerta] Seti Caudata Risso, 1816), by monotypy, gender feminine; invalid junior


homonym of Aglaope Latreille, 1809 (Lepidoptera); name placed on the Official Index of Rejected 

and Invalid Names in Zoology in Opinion 522 in 1958) 

= Niphea Rafinesque, 1815 (nomen novum for Aglaope Rafinesque, 1814, gender feminine; name suppressed 

for the purposes of the Principle of Priority but not for those of the Principle of Homonymy 

under the plenary powers of the ICZN, placed on the Official Index of Rejected and Invalid Names 


= Melicerta Risso, 1816 (type species M.[elicerta] Seti Caudata Risso, 1816, designated by H. Milne 

Edwards, 1837 [in H. Milne Edwards, 1836-1844], gender feminine; invalid junior homonym of 

Melicerta Schrank, 1803 (Rotifera) and Melicerta Peron & Leseur, 1810 (Colenterata); name placed on 

the Official Index of Rejected and Invalid Names in Zoology in Opinion 522 in 1958) 

= Lysmata Risso, 1816 (nomen novum for Melicerta Risso, 1816, gender feminine; name placed on the 


? = Opithiocheirus Leach, 1830a (type species Opithiocheirus Chrysophthalmus Leach, 1830a, by monotypy, 


? = Usterocheirus Leach, 1830a (type species Usterocheirus Macropocoilium Leach, 1830a, designated 

by Holthuis, 1993a, gender masculine) 

= Arno Roux, 1831 (nomen novum for Aglaope Rafinesque, 1814, gender feminine) 

= Eretmocaris Spence Bate, 1888 (type species Eretmocaris stylorostris Spence Bate, 1888, designated 

by Holthuis, 1955b, gender feminine) 

= Hippolysmata Stimpson, 1860a (type species Hippolysmata vitatta Stimpson, 1860a, gender feminine; 


Lysmata amboinensis (De Man, 1888b) 

= Hippolysmata vittata var. amboinensis De Man, 1888b: 495. [Amboina] 

Lysmata anchisteus Chace, 1972: 125; Figs 53-54. [Grenada, Point Saline, rocks at southwest end of first 

beach on lee coast] 

Lysmata ankeri Rhyne & Lin, 2006: 179; Figs 7-9; Plate 1C. [Pompano Beach, Florida, southeast of inlet, 4 m] 

Lysmata argentopunctata Wicksten, 2000a: 3; Figs 1-3. [Morro Colorado, Sonora, 28°20'N 111°18'W, 5-10 m] 

Lysmata bahia Rhyne & Lin, 2006: 191; Figs 16-18; Plate 1F. [Bahia, Salvador, Brazil, seawall at harbor, 

purchased from a pet store] 

Lysmata boggessi Rhyne & Lin, 2006: 183; Figs 10-12; Plate 1D. [northwest of Hernando Beach, Florida, 

seagrass bed, 4 m] 

Lysmata californica (Stimpson, 1866) 

= Hippolysmata californica Stimpson, 1866: 48. [San Diego, Cal.] 

= Hippolyte lineata Lockington, 1877a: 35. [San Diego, between tides] 

Lysmata chica Wicksten, 2000a: 8; Figs 5-7. [Bahía Cartago, Isla Isabela (Albemarle Island), Galapagos, 

0°34'N 90°58'W, shore] 

Lysmata debelius Bruce, 1983e: 115; Figs 1-9. [Polillo Island, east of Luzon, Philippines, 28 m] 

Lysmata dispar Hayashi, 2007: 85; Figs 1-3. [Goodwyn I., 20°32.000'S 116°32.420'E, intertidal] 

Lysmata galapagensis Schmitt, 1924c: 165; Fig. 41. [Northeast of Eden, 7 fms] 

Lysmata grabhami (Gordon, 1935) 

= Hippolysmata grabhami Gordon, 1935: 319; Figs 10, 11a-b. [Funchal, Madeira] 

Lysmata gracilirostris Wicksten, 2000b: 208; Figs 1-3. [Golfo de Papagayo, Costa Rica, 10°42.4'N 85°55'W, 

8-86 m] 

Lysmata guamensis Anker & Cox, 2011: 205; Figs 1-3. [Mariana Archipelago, Guam, Glass Breakwater 

near mouth of Apra Harbor, 3-6 m] 

Lysmata hochi Baeza & Anker, 2008: 149; Figs 1-2. [Panama, Caribbean coast, Bocas del Toro, Isla Colon, 

Playa Bluff, 0.5-1 m] 

Lysmata holthuisi Anker, Baeza & De Grave, 2009: 684; Figs 1-5. [Taboga I., intertidal, low tide, under 

rocks and coral rubble] 

Lysmata intermedia (Kingsley, 1878a) 

= Hippolysmata intermedia Kingsley, 1878a: 90. [Fort Jefferson; Tortugas, Florida] 

Lysmata kempi Chace, 1997: 53. [nomen novum for Hippolysmata dentata Kemp, 1914 nec Palaemon dentatus 

De Haan, 1844]


= Hippolysmata dentata Kemp, 1914: 117; Plate 6, fig. 5. [off M. of Irrawaddy R, 15°20'N 94°55'E, 20 

fms] 

Lysmata kuekenthali (De Man, 1902) 

= Hippolyte kükenthali De Man, 1902: 850; Plate 26, fig. 56; nomen novum for Merhippolyte orientalis 

sensu De Man, 1892a nec Spence Bate, 1888. [Flores; Ternate] 

= Hippolysmata marleyi Stebbing, 1919: 120; Plate 18. [Isezela, Natal] 

Lysmata lipkei Okuno & Fiedler, 2010: 599; Figs 1-4. [Ubara-jima Islet, Katsuura, Boso Peninsula, Honshu, 

Japan, 35°07.7'N 140°16.7'E, 15 m] 

Lysmata moorei (Rathbun, 1901) 

= Hippolysmata moorei Rathbun, 1901: 115; Fig. 23. [Playa de Ponce] 

Lysmata morelandi (Yaldwyn, 1971) 

= Hippolysmata (Hippolysmata) morelandi Yaldwyn, 1971: 90. [Bay of Islands, North Auckland, subtidal 

algal beds on rocky substrate to a depth of about 20 ft] 

Lysmata multiscissa (Nobili, 1904) 

= Hippolysmata multiscissa Nobili, 1904: 232. [Djibouti] 

Lysmata nayaritensis Wicksten, 2000a: 14; Figs 9-11. [Playa Mantac, about 5 miles SE of San Blas, Nayarit, 

Mexico, 21°33'N 105°19'W] 

Lysmata nilita Dohrn & Holthuis, 1950: 339; Fig. 1; Plate 9. [Western half of Bay of Naples] 

= Lysmata Nilita Hope, 1851: 17. [nomen nudum] 

Lysmata olavoi Fransen, 1991b: 63; Figs 1-34. [Azores, Pico, Ponto da Ilha, 38°25'00"N 27°59'10"W, 135 

m] 

Lysmata pederseni Rhyne & Lin, 2006: 187; Figs 13-15, Plate 1E. [southeast of Sombrero Reef, Marathon, 

Florida, in tube sponge, 15 m] 

Lysmata philippinensis Chace, 1997: 53, 75; Figs 19-20. [Albay Gulf, east of southern Luzon, 13°12'N 

123°49'18"E, 267 m] 

Lysmata porteri (Rathbun, 1907) 

= Hippolysmata porteri Rathbun, 1907: 49; Plate 3, fig. 4. [Bay of Valparaiso] 

Lysmata rafa Rhyne & Anker, 2007: 292; Figs 2-4. [Florida, Key west Lakes, west of Key West, under 

ledge, about 1 m] 

Lysmata rathbunae Chace, 1970: 59; Figs 1-3. [Off Boynton Beach, Florida, 26°31'N 80°01'W, 55-64 m] 

Lysmata rauli Laubenheimer & Rhyne, 2010: 299; Figs 1-3. [Salvador, Bahia, Brazil, under rocky ledge in 

tide pool, 2 m depth] 

Lysmata seticaudata (Risso, 1816) 

= M.[elicerta] Seti Caudata Risso, 1816: 110; Plate 2, fig. 1. [environs de Nice, dans les eaux profondes] 

= Aglaope striata Rafinesque, 1814: 24. [Name suppressed under the plenary powers for the purposes 

of the Principle of Priority but not for those of the Principle of Homonymy in Opinion 522 in 

1958] 

= P.[alemon] Cognetii Risso, 1816: 106. [environs de Nice, dans la région des madrépores] 

= Lysmata aberrans Czerniavsky, 1884: 63; Plate 3, Figs 7A-K. [Portus Suchum, 1-1.5 m] 

= Miersia clavigera Chun, 1888: 34; Plate 4, fig. 6. [Type locality not indicated, presumed to be around 

Gulf of Naples, 0-600 m] 

Lysmata splendida Burukovsky, 2000g: 223; Figs 1-3. [Maldives, Ari-Atoll, inner reef Maayafushi, in 

caverns on reef roof wall, 6-35 m] 

Lysmata stenolepis Crosnier & Forest, 1973: 177; Figs 55, 56a-c. [Off Sao Tiago, 275-150 m] 

Lysmata ternatensis De Man, 1902 

= Palaemon dentatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 13 (1844); 176 (1849, as 

Lysmata seticaudata Risso). [Japan] [nec Palaemon dentatus Roemer, 1841] 

= Lysmata seticaudata var. ternatensis De Man, 1902: 846. [Ternate] 

= Hippolysmata acicula Rathbun, 1906: 912; Plate 24, fig. 6. [vicinity of Kauai Island, 7-148 fms] 

= Lysmata affinis Borradaile, 1915b: 209. [Minikoi, Peros Banhos, Salomon, Seychelles] 

Lysmata trisetacea (Heller, 1861) 

= H.[ippolyte] trisetacea Heller, 1861: 29. [im rothen Meere] 

= Lysmata pusilla Heller, 1862c: 287; Plate 3, fig. 26. [im rothen Meere]


= Lysmata chiltoni Kemp, 1914: 110; Plate 6, figs 1-4. [Meyer Island, Kermadec group] 

= Hippolysmata paucidens Rathbun, 1906: 913; Plate 24, fig. 4. [Waikiki beach] 

Lysmata udoi Baeza, Bolaños, Hernandez & López, 2009: 62; Figs 1-3. [Caribbean Coast, Cubagua Island, 

Venezuela, from dens of the ”sapo bocón” toadfish Amphichthys criptocentrus, 2-4 m] 

Lysmata uncicornis Holthuis & Maurin, 1952: 198; Figs 1-2. [port de Casablanca, 4-5 m] 

Lysmata vittata (Stimpson, 1860a) 

= Hippolysmata vittata Stimpson, 1860a: 26. [in portu ”Hong Kong”] 

= Nauticaris unirecedens Spence Bate, 1888: 608; Plate 110, fig. 1. [Hong Kong] 

= Hippolysmata vittata var. subtilis Thallwitz, 1892: 22. [Cebú] 

= Hippolysmata durbanensis Stebbing, 1921b: 20; Plate 5. [cast up on the beach in Durban Bay] 

Lysmata wurdemanni (Gibbes, 1850) 

= Hippolyte Wurdemanni Gibbes, 1850: 197. [Key West Lakes, west of Key West, Florida, 1 m; neotype 

selection by Rhyne & Lin, 2006] 

Lysmata zacae Armstrong, 1941: 10; Fig. 4. [Mataatu Harbor, Savaii, 8ft] 

Lysmatella Borradaile, 1915b 

= Lysmatella Borradaile, 1915b (type species Lysmatella prima Borradaile, 1915b, by monotypy, gender 

feminine) 

Lysmatella prima Borradaile, 1915b: 209. [Maldive Islands] 

Merguia Kemp, 1914 

= Merguia Kemp, 1914 (type species Hippolyte oligodon De Man, 1888a, by monotypy, gender feminine; 


Merguia oligodon (De Man, 1888a) 

= Hippolyte oligodon De Man, 1888a: 277; Plate 18, figs 1-6. [Elphinstone Island] 

Merguia rhizophorae (Rathbun, 1900) 

= Hippolysmata rhizophoræ Rathbun, 1900: 153; Plate 8, fig. 9. [Rio Parahyba do Norte, on mangroves] 

Merhippolyte Spence Bate, 1888 

= Merhippolyte Spence Bate, 1888 (type species Merhippolyte agulhasensis Spence Bate, 1888, by original 

designation, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Merhippolyte agulhasensis Spence Bate, 1888: 619; Plate 110, fig. 4. [Challenger stn 142, 35°04'S 18°37'E, 

south of the Cape of Good Hope, 150 fms] 

Merhippolyte americana Holthuis, 1961d: 1; Fig. 1. [Yucatan Channel, 20°59'30"N 86°23'45"W, 130 fms] 

Merhippolyte ancistrota Crosnier & Forest, 1973: 167; Figs 52-53. [îles du Cap Vert, 15°34.5'N 23°11.5'W, 

185 m] 

Merhippolyte calmani Kemp & Sewell, 1912: 20; Plate 1, figs 1-4. [off the S.W. coast of India, 9°14'10"N 

75°45'E, 237 fms] 

Merhippolyte chacei Kensley, Tranter & Griffin, 1987: 309; Figs 18-19. [New South Wales, 33°43-37'S 151°55' 

– 152°02'E, 686 m] 

Merhippolyte kauaiensis (Rathbun, 1906) 

= Spirontocaris kauaiensis Rathbun, 1906: 913; Plate 24, fig. 5. [vicinity of Kauai Island, 55-362 fms] 

Merhippolyte peroni Ledoyer, 1990: 48; Fig. 3A. [Sud de l’île Saint-Paul] 

Mimocaris Nobili, 1903b 

= Mimocaris Nobili, 1903b (type species Mimocaris heterocarpoides Nobili, 1903b, by monotypy, gender 


Mimocaris heterocarpoides Nobili, 1903b: 6; Fig. 2. [Pulu Burong] 

Nauticaris Spence Bate, 1888 

= Nauticaris Spence Bate, 1888 (type species Nauticaris marionis Spence Bate, 1888, designated by


Fig. 50. Nauticaris magellanica (A. Milne-Edwards, 1891). Photo by Arthur Anker. 

Calman, 1906b, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Nauticaris brucei Stebbing, 1914a: 292; Plate 24. [Scotia stn 461, Gough Island, 100 fms] 

Nauticaris magellanica (A. Milne-Edwards, 1891) (Fig. 50) 

= Hippolyte magellanicus A. Milne-Edwards, 1891: 46; Plate 5, figs 2-2i. [dans la baie Orange, 17 m; au 

nord-ouest de l’île Grévy (cap Hall), 65 m] 

= Hippolyte consobrinus A. Milne-Edwards, 1891: 47; Plate 5, fig. 4. [à terre dans la baie Orange] 

= Nauticaris marionis var. chilensis Doflein & Balss, 1912: 30. [Port Stanley, Falkland-Inseln; Magalhaens-Straße] 

Nauticaris marionis Spence Bate, 1888: 603; Plate 108. [Challenger stns 144A, 46°48'0''S 37°49'30''E, off 

Marion Island, 69 fms; 145, 46°43'0''S 38°4'30''E, off Prince Edward Island, 140 fms; 315, 51°40'S 

57°50'W, off the Falkland Islands, 12 fms] 

= Hippolyte stewarti Thomson, 1889: 259; Plate 13, fig. 1. [Paterson Inlet, Stewart Island] 

= Merhippolyte australis Hodgson, 1902: 233; Plate 29 (as Hippolyte australis). [Auckland Island, 10 

fms] 

Paralatreutes Kemp, 1925 

= Paralatreutes Kemp, 1925 (type species Paralatreutes bicornis Kemp, 1925, by original designation 

and monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology in 


Paralatreutes bicornis Kemp, 1925: 334; Figs 23-24. [Ross Channel, Port Blair, Andamans, 3-4 fms] 

Paralebbeus Bruce & Chace, 1986 

= Paralebbeus Bruce & Chace, 1986 (type species Paralebbeus zotheculatus Bruce & Chace, 1986, by 


Paralebbeus zotheculatus Bruce & Chace, 1986: 238; Figs 1-6. [17°30.1'S 118°28.9'E, 505-506 m] 

Paralebbeus zygius Chace, 1997: 82; Figs 25-28. [Selat Butung, Sulawesi (Celebes), Indonesia, 5°35'00"S 

122°20'00''E, 1023 m]


Phycocaris Kemp, 1916a 

= Phycocaris Kemp, 1916a (type species Phycocaris simulans Kemp, 1916a, by original designation 

and monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Phycocaris simulans Kemp, 1916a: 392; Fig. 2; Plate 36, fig. 2. [near Ross Island, Port Blair, Andaman Islands, 

2-4 fms] 

Saron Thallwitz, 1891 

= Saron Thallwitz, 1891 (type species Hippolyte gibberosus H. Milne Edwards, 1837 [in H. Milne Edwards, 

1834-1840], by original designation and monotypy, gender masculine; name placed on the 


Saron inermis Hayashi in Debelius, 1983: 117 (partim, see Hayashi, 1989). [Indonesien] 

Saron marmoratus (Olivier, 1811) 

= Palæmon marmoratus Olivier, 1811: 665. [Nouvelle-Hollande] 

= Hippolyte gibberosus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les côtes de 

la Nouvelle-Hollande] 

= Hippolyte Leachii Guérin-Méneville, 1838 [in Guérin-Méneville, 1829-1838]: 37. [côtes de l’île de 

Strong ou Oualan] 

= H.[ippolyte] Hemprichii Heller, 1861: 29. [im rothen Meere] 

= Hyppolite Kraussii Bianconi, 1869: 209; Plate 1, fig. 2a. [Mosambico] 

= Nauticaris grandirostris Pearson, 1905: 79; Plate 1, figs 6-6c. [entrance to Galle Harbour, 4.5-7 fms] 

Saron neglectus De Man, 1902: 854; Plate 26, figs 58-58b. [Ternate; Pulo Edam in der Bai von Batavia; 

Kagoshima, Japan] 

Saron rectirostris Hayashi, 1984: 116. [Indonesia] 

Spirontocaris Spence Bate, 1888 

= Sowerbyus Hoek, 1887 (type species Sowerbyus spinus Hoek, 1887, probably a later combination of 

Cancer Spinus Sowerby, 1805 [in Sowerby, 1804-1806], by monotypy, gender masculine; name place 


= Spirontocaris Spence Bate, 1888 (type species Cancer Spinus Sowerby, 1805 [in Sowerby, 1804-1806], 

by monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Spirontocaris arcuata Rathbun, 1902a: 893. [Washington Sound, Straits of Fuca, 48 fms] 

Spirontocaris arcuatoides Kobjakova, 1962: 244; Fig. 2. [Ostrov Zelenii (Green Island), 14 m; Shitokan Island, 

10-4 m] 

Spirontocaris brashnikowi Kobjakova, 1936a: 221. [Tatar Strait and Peter the Great Bay, Sea of Japan; nomen 

novum for Spirontocaris dalli sensu Bražnikov, 1907 nec Rathbun, 1902a; fully described in 


= Spirontocaris brashnikovi Makarov, 1941: 121. [Avachinsk Bay; nomen nudum] 

Spirontocaris brevidigitata Kobjakova, 1935: 88; Fig. 3. [von der Bucht Peters des Großen nach Norden bis 

zur Bucht Nelma, 75-1380 m] 

Spirontocaris dalli Rathbun, 1902a: 894. [Coal Harbor, Unga Island, Alaska, 9-8 fms] 

Spirontocaris gurjanovae Kobjakova, 1955: 238; Fig. 3. [4th Kuril Straits, Kurile Islands, 100 m] 

Spirontocaris holmesi Holthuis, 1947a: 8. [nomen novum for Spirontocaris bispinosus Holmes, 1900 nec 

Hippolyte bispinosa De Haan, 1844 [in De Haan, 1833-1850]] 

= Spirontocaris bispinosus Holmes, 1900: 207. [Puget Sound] 

Spirontocaris intermedia Kobjakova, 1936a 

= Spirontocaris spina intermedia Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova, 

1937] 

= Spirontocaris spina intermedia Makarov, 1941: 117; Figs 1-2. [Bering Sea and Chukchi Sea, 30-100 m] 

Spirontocaris lamellicornis (Dana, 1852a) 

= Hippolyte lamellicornis Dana, 1852a: 24. [in freta ”de Fuca” Oregoniæ, juxta portum ”Dungeness”]


Spirontocaris lilljeborgii (Danielssen, 1859) 

? = Astacus histrio Fabricius, 1775: 415. [Groenlandia] 

= H.[ippolyte] Lilljeborgii Danielssen, 1859: 5. [Slotholem (Lofoten) paa sandig Leerbund, 40 fms] 

= Hippolyte securifrons Norman, 1862: 151. [off the Shetland Isles; full description in Norman, 1863] 

Spirontocaris microdentata Kobjakova, 1962 

= Spirontocaria [sic] microdentatus Kobjakova, 1962: 245; Fig. 4. [Ushishir Island, about 18 m] 

Spirontocaris murdochi Rathbun, 1902a: 893. [off Robben Island, east coast of Sghalin, Okhostk Sea] 

Spirontocaris ochotensis (Brandt, 1851) 

= Hippolyte ochotensis Brandt, 1851: 120; Plate 5, Figs 17-17b. [Type locality not indicated] 

= Spirontocaris mororani Rathbun, 1902b: 43; Fig. 16. [Mororan, Hokkaido] 

= Spirontocaris makarovi Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova, 1937] 

= Spirontocaris makarovi spatula Kobjakova, 1936a: 221. [Okhotsk Sea; fully described in Kobjakova, 

1937] 

= Spirontocaris onagawaensis Yokoya, 1939: 268; Fig. 5. [Takasiro, 7.5 m; Nonohama, 4.6 m] 

Spirontocaris pectinifera (Stimpson, 1860a) 

= Hippolyte pectinifera Stimpson, 1860a: 35. [sinu ”Hakodadi” f. conchoso org 8.] 

= Spirontocaris crassirostris Kubo, 1951: 274; Figs 11-12. [off Heta, Izu peninsula, ca. 300 m] 

Spirontocaris phippsii (Krøyer, 1841) 

= Hippolyte Phippsii Krøyer, 1841: 575. [Spitsbergen, Norges Vestkyst (og Grønland ?)] 

= Hippolyte turgida Krøyer, 1841: 575. [Spitsbergen, Norges Vestkyst, Grønland] 

= Hippolyte vibrans Stimpson, 1871: 125. [Massachusetts Bay] 

Spirontocaris prionota (Stimpson, 1864) 

= Hippolyte prionota Stimpson, 1864: 153. [Hale’s Passage, 10 fms; off Lummi Island, 8-12 fms, near 

San Juan Island, 2-4 fms (all Puget Sound)] 

= Spirontocaris macrodonta J.F.L. Hart, 1930: 102; Plate 1. [Gonzales Point, tide pool; False Narrows, 

near the water’s edge; Departure Bay in about 10 fms (all Vancouver Island)] 

Spirontocaris sica Rathbun, 1902a: 894. [Santa Barbara Channel, 265 fms] 

Spirontocaris snyderi Rathbun, 1902a: 894. [Monterey Bay, California] 

Spirontocaris spinus (Sowerby, 1805 [in Sowerby, 1804-1806]) 

? = Astacus histrio Fabricius, 1775: 415. [Groenlandia] 

= Cancer spinus Sowerby, 1805 [in Sowerby, 1804-1806]: 47; Plate 23. [among oysters on the Scottish 

coast] 

= Hippolyte Sowerbæi Leach, 1817 [in Leach, 1815-1875]: Plate 39, figs 1-10. [Newhaven, in the Frith 

of Forth] 

= Spirontocaris spina laevidens Kobjakova, 1936a: 221. [Tatar Strait and Peter the Great Bay, Sea of Japan; 


Spirontocaris truncata Rathbun, 1902a: 894. [Heceta Bank, Oregon, 50 fms] 

Spirontocaris urupensis Kobjakova, 1962: 243; Fig. 1. [Urup Island, 5-7 m] 

Thinora Bruce, 1998d 

= Thinora Bruce, 1998d (nomen novum for Thorina Bruce, 1997b, gender feminine) 

= Thorina Bruce, 1997b (type species Thor maldivensis Borradaile, 1915b, by original designation 

and monotypy, gender feminine; invalid junior homonym of Thorina Stephensen, 1944 (Amphipoda)) 

Thinora maldivensis (Borradaile, 1915b) 

= Thor maldivensis Borradaile, 1915b: 208. [Minikoi, Maldives; Salomon] 

Thor Kingsley, 1878a 

= Thor Kingsley, 1878a (type species Thor floridanus Kingsley, 1878a, by monotypy, gender masculine; 


= Paschocaris Nobili, 1905b (type species H.[ippolyte] paschalis Heller, 1862c, by original designation 



Thor algicola Wicksten, 1987: 27; Figs 1-3. [Bahía Bacochibampo, Guaymas, Sonora, Mexico, 27°57'N 

111°02'W, 5 m, among Sargassum] 

Thor amboinensis (De Man, 1888b) 

= Hippolyte amboinensis De Man, 1888b: 535. [Amboina] 

= Thor discosomatis Kemp, 1916a: 388; Fig. 1; Plate 36, fig. 1. [Port Blair, Andamans] 

Thor cocoensis Wicksten & Vargas, 2001: 139; Figs 1-4. [Bajo Alcyone, Isla del Coco, Costa Rica (Pacific), 

35 m] 

Thor cordelli Wicksten, 1996b: 295; Fig. 1. [Sulphur Bay, Clarion Island, Mexico, shallow] 

Thor dobkini Chace, 1972: 133; Fig. 57. [Florida, Punta Rassa, 2 m] 

Thor floridanus Kingsley, 1878a: 95. [Key West, Florida] 

Thor intermedius Holthuis, 1947a: 51; Figs 4-6. [Sissie near Misool, shore and reef] 

Thor manningi Chace, 1972: 137; Figs 59-60. [English Harbour, Antigua, from bottom of yacht anchored 

for several months] 

Thor marguitae Bruce, 1978e: 159; Figs 1-6. [reef flat, Heron Island, Capricorn Islands, Queensland, Australia, 

23°25.25'S 151°55.20'E] 

Thor paschalis (Heller, 1862c) 

= H.[ippolyte] paschalis Heller, 1862c: 276; Plate 3, fig. 24. [im rothen Meere] 

Thor spinipes Bruce, 1983f: 2; Figs 1-6. [Burford Island, Cobourg Peninsula, Northern Territory, Australia, 

11°29.3'S 131°57.5'E, muddy reef flat pools at LWS tide] 

Thor spinosus Boone, 1935: 192; Plate 52. [Tmukus Roads, Bali, Dutch East Indies, in coral] 

Thorella Bruce, 1982d 

= Thorella Bruce, 1982d (type species Thorella cobourgi Bruce, 1982d, by original designation and 


Thorella cobourgi Bruce, 1982d: 452; Figs 1-5. [Black Point, Port Essington, Cobourg Peninsula, Northern 

Territory, 11°09.0'S 132°08.2'E, 1-2 m, in Sargassum] 

Tozeuma Stimpson, 1860a 

= Tozeuma Stimpson, 1860a (type species Tozeuma lanceolatum Stimpson, 1860a, by monotypy, gender 

neuter) 

= Angasia Spence Bate, 1863 (type species Angasia pavonina Spence Bate, 1863, by monotypy, gender 

feminine) 

Tozeuma armatum Paul’son, 1875 

= Toz.[euma] armata Paul’son, 1875: 99; Plate 15, fig. 2. [Red Sea] 

= Angasia Stimpsonii Henderson, 1893: 437; Plate 40, figs 18-20. [Gulf of Martaban] 

Tozeuma carolinense Kingsley, 1878a (Fig. 51) 

= Tozeuma carolinensis Kingsley, 1878a: 90. [Fort Macon, North Carolina] 

Fig. 51. Tozeuma carolinense Kingsley, 1878. Photo by Arthur Anker.


Tozeuma cornutum A. Milne-Edwards, 1881b: 16. [près de Barbades, 40 brasses] 

Tozeuma elongata (Baker, 1904) 

= Angasia elongata Baker, 1904: 147; Plate 27, figs 1-4. [south Australian coast, 15 fms; Port Victor] 

Tozeuma erythraeum Nobili, 1904 

= Tozeuma erythræum Nobili, 1904: 231. [Mer Rouge] 

Tozeuma kimberi (Baker, 1904) 

= Angasia kimberi Baker, 1904: 149; Plate 27, fig. 5. [Port Willunga, 4 fms] 

Tozeuma lanceolatum Stimpson, 1860a: 27. [in portu ”Hong Kong”, in fundo limoso prof. sex. org. sat 

vulgaris] 

Tozeuma novaezealandiae Borradaile, 1916 

= Tozeuma novae-zealandiae Borradaile, 1916: 86; Fig. 3. [7 miles E of North Cape, New Zealand, 128 

m] 

Tozeuma pavoninum (Spence Bate, 1863) 

= Angasia pavonina Spence Bate, 1863: 498; Plate 90, figs 1-1z. [St. Vincent’s Gulf, 4.5 fms] 

= Angasia robusta Baker, 1904: 150; Plate 28, figs 1-8. [St.Vincent Gulf, 10-12 fms] 

Tozeuma serratum A. Milne-Edwards, 1881b: 16. [près de Barbades, 56 brasses] 

Tozeuma tomentosum (Baker, 1904) 

= Angasia tomentosa Baker, 1904: 152; Plate 29, figs 1-4. [south Australian coast, about 20 fms] 

Trachycaris Calman, 1906b 

= Trachycaris Calman, 1906b (type species Platybema rugosus Spence Bate, 1888, by original designation 

and monotypy, gender feminine; name placed on the Official List of Generic names in Zoology 


Trachycaris restricta (A. Milne-Edwards, 1878) 

= Hippolyte restrictus A. Milne-Edwards, 1878: 231. [îles du Cap-Vert] 

Trachycaris rugosa (Spence Bate, 1888) 

= Platybema rugosus Spence Bate, 1888: 579; Plate 104, fig. 2. [Challenger stn 24, 18°38'30''N 65°5'30''W, 

off Culebra island, West Indies, 390 fms] 

Family oGyRIDIDAe Holthuis, 1955b 

Ogyrides Stebbing, 1914b 

= Ogyris Stimpson, 1860a (type species Ogyris orientalis Stimpson, 1860a, by monotypy, invalid junior 

homonym of Ogyris Westwood, 1851 (Lepidoptera), gender feminine; name placed on the Official 

Index of Rejected and Invalid Names in Zoology in Opinion 470 in 1957) 

= Ogyrides Stebbing, 1914b (nomen novum for Ogyris Stimpson, 1860a, gender masculine; name 


Ogyrides alphaerostris (Kingsley, 1880) 

= Ogyris alphærostris Kingsley, 1880: 420; Plate 14, fig. 7. [Northampton County, Virginia, Eastern 

shore, Atlantic side] 

= Ogyris occidentalis Ortmann, 1893: 46; Plate 3, fig. 4. [Plankton-Expedition der Humbolt-Stiftung: 

Tocantins-Mündung JN 239 (0.6°S 48.1°W, 0 m); Pl. 106 (0.7°S 48.2°W, 12 m)] 

= Ogyrides yaquiensis Armstrong, 1949: 3; Fig. 1. [Rio Yaqui del Sul, Dominican Republic, 1-3 ft] 

= Ogyrides limicola Williams, 1955: 57; Fig. 1. [mouth of Far Creek at Ebgelhard, Hyde Count, North 

Carolina, 1-8 ft] 

Ogyrides delli Yaldwyn, 1971: 89. [Lowry Bay, Wellington Harbour, 4 fms] 

Ogyrides hayi Williams, 1981: 145. [off Bogue Bank, west of Fort Macon, North Carolina, ca. 3.5 m] (Fig. 52) 

Ogyrides mjoebergi (Balss, 1921a) 

= Ogyris Mjöbergi Balss, 1921a: 7; Figs 1-2 [Cape Jaubert, 45 meilen WSW, 54 fuss] 

Ogyrides orientalis (Stimpson, 1860a) 

= Ogyris orientalis Stimpson, 1860a: 36. [mari Sinensi, et in sinu ”Kagosima”, 5-25 org] 

Ogyrides rarispina Holthuis, 1951a: 119; Fig. 25. [anchorage of Monrovia, Liberia, 11 m]


Fig. 52. Ogyrides hayi Williams, 1981. Photo by Arthur Anker. 

Ogyrides saldanhae Barnard, 1947 

= Ogyrides saldanhæ Barnard, 1947: 387. [Saldanha Bay, 10 fms] 

Ogyrides sibogae (De Man, 1910a) 

= Ogyris Sibogae De Man, 1910a: 318. [6°4.1'N 120°44'E, 535 m] 

Ogyrides striaticauda Kemp, 1915: 284; Figs 28-30. [Chilka Lake] 

Ogyrides tarazonai Wicksten & Mendez G., 1988: 622; Fig. 1. [Ventanilla, Peru, 11°50'S, shallow water, 50 

m from tide line] 

Superfamily PRoCeSSoIDeA ortmann, 1896a 

Family PRoCeSSIDAe ortmann, 1896a 

Ambidexter Manning & Chace, 1971 

= Ambidexter Manning & Chace, 1971 (type species Ambidexter symmetricus Manning & Chace, 1971, 


Ambidexter panamensis Abele, 1972: 373; Figs 4-5. [Panama, Canal Zone, Pacific coast, Ft. Amador, Naos 

Island, from burrows on sand-mud beach adjacent to Smithsonian Tropical Marine Laboratory] 

Ambidexter swifti Abele, 1972: 366; Figs 1-3. [Panama, Panama Province, Pacific coast, Punta Paitilla, 0.5 

m] 

Ambidexter symmetricus Manning & Chace, 1971: 3-7; Figs 1-2. [Florida, Dade County, Miami, Biscayne 

Bay, Matheson Hammock Wading Beach, push nets on grass flats] 

Clytomanningus Chace, 1997 

= Clytomanningus Chace, 1997 (type species Processa molaris Chace, 1955, by original designation, 


Clytomanningus coutierei (Nobili, 1904) 

= Processa Coutierei Nobili, 1904: 234. [Djibouti] 

Clytomanningus molaris (Chace, 1955) 

= Processa molaris Chace, 1955: 11; Fig. 5. [Rongelap Atoll, Burok Island, intertidal coral] 

Hayashidonus Chace, 1997 

= Hayashidonus Chace, 1997 (type species Nika japonica De Haan, 1844 [in De Haan, 1833-1850], by 

original designation and monotypy, gender masculine)


Hayashidonus japonicus (De Haan, 1844 [in De Haan, 1833-1850]) 

= Nika japonica De Haan, 1844 [in De Haan, 1833-1850]: Plate 46, fig. 6 (1844); 184, Plate N (1849). 


Nikoides Paul’son, 1875 

= Nikoides Paul’son, 1875 (type species Nik.[oides] Danae Paul’son, 1875, by monotypy, gender masculine; 


Nikoides boraboraensis Burukovsky, 2002b: 1389; unnumbered figure. [French Polynesia, Bora-Bora Atoll, 

near Motutagu, 1-1.5 m, on coral boulder] 

Nikoides danae Paul’son, 1875 

= Nik.[oides] Danae Paul’son, 1875: 98; Plate 14, Figs 5-5d. 

Nikoides gurneyi Hayashi, 1975a: 58; Figs 3-4. [Kikambala, Kenya, weedy pools inner reef flat at LWS] 

Nikoides longicarpus Noël, 1986: 264; Figs 1-8. [Philippines, 13°59'N 120°10'E, 164-150 m] 

Nikoides maldivensis Borradaile, 1915b: 209. [Amirante island, 25-28 ft; see Chace, 1997] 

= Processa Jacobsoni De Man, 1921: 95. [Sinabang, island of Simalur, near the west coast of Atjeh, 

Sumatra] 

Nikoides multispinatus Hayashi, 1981b: 53; Figs 1-3. [Heron Island, Great Barrier Reef, Australia, 35ft] 

Nikoides plantei Burukovsky, 2007c: 1; Fig. 1. [Madagascar, Antseranana, 47 m] 

Nikoides schmitti Manning & Chace, 1971: 8; Figs 3-5. [Florida, Monroe County, Tortugas, near black 

buoy, 1.25 km south of Garden Key, 26 m] 

Nikoides sibogae De Man, 1918a 

= Nikoides Sibogae De Man, 1918a: 160. [Siboga Expedition stns 71, Makassar and surroundings, up 

to 32 m; 274, 5°28'.2S 134°53'.9E, east coast of Aru-islands, 57 m; 282, 8°25'.2S 127°18'.4E, anchorage 

between Nusa Besi and the N.E.-point of Timor, 27-54 m] 

Nikoides steinii (Edmondson, 1935a) 

= Processa steinii Edmondson, 1935a: 3; Fig. 1. [shoal water reef of Maui, in a coral head] 

= Nikoides nanus Chace, 1955: 8; Fig. 4. [Eniwetok Atoll, Runit island, intertidal] 

Processa leach, 1815 [in leach, 1815-1875] 

= Thalassalpes Bosc, 1813 (type species N.[ika] Edulis Risso, 1816, designated by Holthuis, 1955b, 

gender masculine; name suppressed for the purposes of the Principle of Priority, but not those of 

the Principle of Homonymy, placed on the Official Index of Rejected and Invalid Generic Names in 


= Processa Leach, 1815 [in Leach, 1815-1875] (type species Processa canaliculata Leach, 1815 [in Leach, 



= Nika Risso, 1816 (type species N.[ika] Edulis Risso, 1816, designated by H. Milne Edwards, 1837 [in 

H. Milne Edwards, 1836-1844], gender feminine) 

= Velocina Gistel, 1848 (nomen novum for Processa Leach, 1815 [in Leach, 1815-1875], gender feminine) 

? = Chiereghina Nardo, 1869 (type species Cancer pellucidus Nardo, 1847 (probably a junior subjective 

synonym of N.[ika] Edulis Risso, 1816), gender feminine) 

= Hectarthropus Spence Bate, 1888 (type species Hectarthropus expansus Spence Bate, 1888, designated 


Processa acutirostris Nouvel & Holthuis, 1957: 23; Figs 81-107. [prés de Monaco] 

? = Cancer longipes Nardo, 1847: 6. [trovato di rado nel golfo] 

Processa aequimana (Paul’son, 1875) 

= N.[ika] aequimana Paul’son, 1875: 97; Plate 14, figs 6-6a. [Red Sea] 

Processa affinis Hayashi, 1975a: 85; Fig. 12. [Sanana Bay, east coast of Sanana, Soela Islands, 22 m] 

Processa australiensis Baker, 1907: 185; Plate 25, figs 2-2e. [south Australian coast] 

Processa austroafricana Barnard, 1947: 386. [Agulhas Bank] 

Processa bermudensis (Rankin, 1900) 

= Nika bermudensis Rankin, 1900: 536; Plate 17, figs 2, 2a-b. [Harrington Sound, 1fm]


Processa borboronica Holthuis, 1951a: 43; Fig. 7. [off Gold Coast, 5°37'N 0°38'E, 50 m] 

Processa brasiliensis Christoffersen, 1979: 364; Figs 34-35. [8°15'S 34°42'W, off Pernambuco, 33 m] 

Processa canaliculata Leach, 1815 [in Leach, 1815-1875]: Plate 41. [Torcross, southern coast of Devon] 

= Nika cannelata Griffith & Pidgeon, 1833: Plate 17, fig. 2. [Type locality not indicated] 

= Nika Couchii Bell, 1847 [in Bell, 1844-1853]: 278; unnumbered vignette. [coast of Cornwall] 

= Nika edulis var. britannica Czerniavsky, 1884: 59. [nomen novum for Nika Couchii Bell, 1847 [in Bell, 

1844-1853]] 

= Nika mediterranea Parisi, 1915: 65. [Nizza] 

= Processa prostatica Zariquiey Cenarro, 1941: 345; Figs 17-35. [Région Tramontana] 

Processa compacta Crosnier, 1971: 577; Fig. 3. [Congo, baie de Pointe-Noire, 10 m] 

= Processa barnardi Hayashi, 1975a: 92; Figs 15-16. [South Africa, 33°50'S 25°46'E, 20 fms] 

Processa crosnieri Noël, 1986: 274; Figs 9-12. [Ile de la Réunion, radiale B au large de St Pierre, 21°20'7[S] 

55°26'9[E], 73-77 m] 

Processa demani Hayashi, 1975a: 98; Figs 19-20. [Elat, west coast of Great Kei Islands, 27 m] 

Processa dimorpha Hayashi, 1975a: 102; Fig. 21. [Japan, Sea of Genkai, off Shingu, Fukuoka Prefecture, 

weed belts] 

Processa edulis crassipes Nouvel & Holthuis, 1957: 16; Figs 28-37. [herbier de l’île Verte, Roscoff, France, 

Finistère] 

Processa edulis edulis (Risso, 1816) 

= N.[ika] Edulis Risso, 1816: 85; Plate 3, fig. 3. [dans la région des algues, environs de Nice] 

? = Cancer pellucidus Nardo, 1847: 6. [trovato nel golfo, tra gli Asprei] 

? = Nikoides pontica Sowinsky, 1882: 226. [mer Noire] 

= Nika edulis var. mediterranea forma typica Czerniavsky, 1884: 60. 

= Processa edulis arcassonensis Nouvel & Holthuis, 1957: 18; Figs 38-52. [basin d’Arcachon] 

Processa elegantula Nouvel & Holthuis, 1957: 37; Figs 174-204. [baie de Cadaqués, 30-40 m] 

Processa famelica Manning & C.W.J. Hart, 1991: 317; Figs 1-3. [Hungry Bay, Bermuda, 32°17'N 64°46'W] 

Processa fimbriata Manning & Chace, 1971: 19; Figs 8-10. [Florida, Monroe County, Tortugas, off East Key, 

3 m] (Fig. 53) 

Processa foresti Noël, 1986: 280; Fig.13. [Philippines, 13°59'N 120°18'E, 187-205 m] 

Processa gracilis Baker, 1907: 187; Plate 25, Figs 3-3c. [South Australia coast] 

Processa guyanae Holthuis, 1959a: 115, Figs 18-19. [NW of the Coppename River, 6°54'N 56°14'W, 49 m] 

= Processa tenuipes Manning & Chace, 1971: 31; Figs 17-18. [Gulf of Mexico, off west coast of Florida, 

29°12'N 84°22'W, 31 m] 

Processa hawaiensis (Dana, 1852a) 

= Nika Hawaiensis Dana, 1852a: 20. [prope insulam ”Maui” Hawaiensem] 

= Processa paucirostris Edmondson, 1930: 3; Fig. 1. [Kahana Bay, Oahu, in shallow water] 

Processa hemphilli Manning & Chace, 1971: 23; Figs 11-12. [Florida, Collier County, Marco, 2-6 m] 

Processa indica Noël, 1986: 282; Figs 14-17. [Inde, côte N-E (baie du Bengale), à proximité de Visakhapatnam, 

40-60 m] 

Processa intermedia Holthuis, 1951a: 38; Figs 5-6. [off French Guinea, 9°57'N 15°22'W, 25 m] 

Processa kotiensis (Yokoya, 1933) 

= Nika kotiensis Yokoya, 1933: 30; Fig. 13. [Sôyô-Maru stn 322, west of Muroto-zaki, 110 m] 

Processa longirostris Hayashi, 1975a: 118; Figs 26-27. [South Viet Nam, Bay of Nha Trang, 5 m] 

Processa macrodactyla Holthuis, 1952a: 30; Fig. 8. [Baie de Gorréi, au S. d’Angra de Cintra, Rio de Oro] 

Processa macrognatha (Stimpson, 1860a) 

= Nica macrognatha Stimpson, 1860a: 26. [in portu ”Hong Kong” f. conchoso, p. 8 org.] 

Processa macrophthalma Nouvel & Holthuis, 1957: 27; Figs 108-133. [prés de Monaco] 

Processa moana Yaldwyn, 1971: 91. [off Cuvier Id, Bay of Plenty, 30-40 fms] 

Processa modica modica Williamson in Williamson & Rochanaburanon, 1979: 12; Figs 1, 2a-e, 3f-g, 4-5, 9d. 

[52°19'N 4°22'E, about 25 km north of The Hague, The Netherlands] 

Processa modica carolii Williamson in Williamson & Rochanaburanon, 1979: 21; Figs 2f-h, 3h-i, 6a, d, g. 

[Tunisia, Carthage-Salammbô, from off Baths of Antoninus to INSTOP, 4-5 m]


Fig. 53. Processa fimbriata Manning & Chace, 1971. Photo by Arthur Anker. 

Processa namibiensis Macpherson, 1983: 60; Figs 31-32. [norte de Namibia, 17°44'S 11°34'E, 70 m] 

Processa neglecta Hayashi, 1975a: 127; Figs 31-32. [South Viet Nam, Bay of Nha Trang, 11 m] 

Processa nouveli nouveli Al-Adhub & Williamson, 1975: 694; Figs 1a-b, 2a-b, 3. [Monaco] 

Processa nouveli holthuisi Al-Adhub & Williamson, 1975: 701. [20 km north-west of Port Erin, Isle of Man, 

90 m] 

Processa packeri Manning & Chace, 1990: 24; Fig. 11. [Ascension Island, McArthur Point, northern edge 

of South West Bay] 

Processa parva Holthuis, 1951a: 47; Fig. 8. [off Bathurst, Gambia, 18 m] 

Processa peruviana Wicksten, 1983a: 29; Figs 4-6. [Isla Manuelita, Costa Rica, 5°36'09''N 87°01'14''W, 146 m] 

Processa philippinensis Noël, 1986: 288; Fig. 18. [Philippines, 3°53'N 120°08'E, 134-129 m] 

Processa pippinae Wicksten & Méndez G., 1985: 16; Figs 1-4. [E of Isla Ángel de la Guardia, Gulf of California, 

Mexico, 29°20'N 113°11.5'W, 265-284 m] 

Processa processa (Spence Bate, 1888) 

= Nika processa Spence Bate, 1888: 527. (nec Plate 95, fig. 1) [Amboina, 15 fms] 

Processa profunda Manning & Chace, 1971: 25; Figs 13-15. [Gulf of Mexico, off west coast of Florida, 

28°36'N 85°33'30''W, 202 m] 

Processa pygmaea Burukovsky, 1990: 191; Fig. 1B. [25°04'S 97°29'W, 267-280 m] 

Processa riveroi Manning & Chace, 1971: 28; Fig. 16. [Puerto Rico, Lajas, La Parguera, east side of Maguey 

Island, on Thalassia flats] 

Processa robusta Nouvel & Holthuis, 1957: 19; Figs 53-80. [prés de Monaco, Baie de Monte-Caro, fond de 

Posidonies, 15 m] 

Processa sulcata Hayashi, 1975a: 134; Fig. 34. [Japan, Sea of Genkai, Ainoshima Island, Fukuoka Prefecture]


Processa vicina Manning & Chace, 1971: 34; Figs 19-20. [off north Carolina, 34°35'30''N 75°45'30''W, 59 m] 

Processa vossi Manning, 1992: 552; Figs 1-2. [Florida, Atlantic Ocean, Martin County, about 1 mile north 

of St. Lucie Inlet, 27°11'N 80°09.5'W, Bathtub Beach] 

Processa wheeleri Lebour, 1941: 403; Figs 1-9, 11-27. [5 miles from Bermuda, 150 m] 

Processa zostericola Hayashi, 1975a: 137; Fig. 35. [Japan, Tomioka Bay, Amakusa Islands, Kumamoto Prefecture, 

Zostera belt] 

Superfamily PANDAloIDeA Haworth, 1825 

Family PANDAlIDAe Haworth, 1825 

Anachlorocurtis Hayashi, 1975b 

= Anachlorocurtis Hayashi, 1975b (type species Anachlorocurtis commensalis Hayashi, 1975b, by original 


Anachlorocurtis commensalis Hayashi, 1975b: 175; Figs 1-3. [off Andonohana, Shionomisaki, southern Kii 

Peninsula, Japan, 8 m, associated with Antipathes sp.] 

Atlantopandalus Komai, 1999 

= Atlantopandalus Komai, 1999 (type species Pandalus propinqvus G.O. Sars, 1870, by original designation 


Atlantopandalus propinqvus (G.O. Sars, 1870) 

= Pandalus propinqvus G.O. Sars, 1870: 148. [Lofoten near Skrava, Guldbrandsøy, 360-540 m; lectotype 

designation by Komai, 1999; spelling maintained as propinqvus under Art. 32.1] 

Austropandalus Holthuis, 1952d 

= Austropandalus Holthuis, 1952d (type species Hippolyte Grayi Cunningham, 1871, by original designation 


Austropandalus grayi (Cunningham, 1871) 

= Hippolyte Grayi Cunningham, 1871: 496; Plate 59, fig. 8. [Port Otway] 

= Pandalus paucidens Miers, 1881b: 74; Plate 7, figs 6-7. [Tom Bay, on a bottom of rock, kelp, and mud; 

Trinidad Channel, 30 fms, on a sandy bottom] 

= Nothocaris spiniserratus Spence Bate, 1888: 663; Plate 132, fig. 1. [Challenger stn 304, 46°53'15''S 

75°12'0''W, Port Otway, Messier Channel, 45 fms] 

Bitias Fransen, 1990 

= Bitias Fransen, 1990 (type species Bitias stocki Fransen, 1990, by original designation and monotypy, 


Bitias brevis (Rathbun, 1906) 

= Pandalus brevis Rathbun, 1906: 916; Fig. 65; Plate 21, fig. 3. [vicinity of Niihau Island, 426-417 fms] 

Bitias stocki Fransen, 1990: 68; Figs 1-3. [Azores, S of Pico, 38°09'N 28°31'W, 1320-1350 m] 

Calipandalus Komai & Chan, 2003 

= Calipandalus Komai & Chan, 2003 (type species Calipandalus elachys Komai & Chan, 2003, by original 


Calipandalus elachys Komai & Chan, 2003: 880-889; Figs 1-4. [22°18.59'N 121°29.39'E, 267-302 m] 

Chelonika Fransen, 1997b 

= Chelonika Fransen, 1997b (type species Chelonika macrochela Fransen, 1997b, by original designation 


Chelonika macrochela Fransen, 1997b: 179, Figs 1-7. [New Caledonia, 22°21.5'S 166°14.6'E, 144-155 m] 

Chlorocurtis Kemp, 1925 

= Chlorocurtis Kemp, 1925 (type species Chlorocurtis miser Kemp, 1925 (junior subjective synonym of


Virbius (?) jactans Nobili, 1904), by original designation and monotypy, gender masculine; name 


Chlorocurtis jactans (Nobili, 1904) 

= Virbius (?) jactans Nobili, 1904: 230. [Djibouti] 

= Chlorocurtis miser Kemp, 1925: 280. [Port Blair, Andamans, low water] 

Chlorotocella Balss, 1914b 

= Chlorotocella Balss, 1914b (type species Chlorotocella gracilis Balss, 1914b by monotypy, gender feminine; 


Chlorotocella gracilis Balss, 1914b: 33; Figs 16-22. [Type locality restricted to Sagami Bay (Japan) by 

Hayashi & Miyake, 1968b] 

Chlorotocella spinicaudus (H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]) 

= Hippolyte spinicaudus H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 378. [les côtes de 

la Nouvelle Hollande] 

= Pandalus leptorhynchus Stimpson, 1860a: 38; nec Kinahan, 1858b. [Portu Jacksoniensi Australiae, ad 

littora arenosa et algosa] 

= Pandalus (Parapandalus) spinicauda var. gibber Hale, 1924: 68; Plate 4, figs 6-7. [St. Vincent Gulf] 

Chlorotocus A. Milne-edwards, 1882 

= Chlorotocus A. Milne-Edwards, 1882 (type species Chlorotocus gracilipes A. Milne-Edwards, 1882, 

by monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology in 


Chlorotocus crassicornis (A. Costa, 1871) (Fig. 54) 

= Pandalus crassicornis A. Costa, 1871: 89; Plate 2, fig. 2. [golfo di Napoli] 

= Chlorotocus gracilipes A. Milne-Edwards, 1882: 14. [Alboran Sea off Mediterranean coast of Morocco, 

35°21'30"N 4°33'05"W – 35°24'20"N 4°32'45'W, 322-432 m; see Chace, 1985] 

= Palemon chlorotocus Filhol, 1885b: Plate 1. [Type locality not indicated] 

? = Chlorotocus incertus Spence Bate, 1888: 674; Plate 116, Figs 1-2. [Challenger stn 142, 35°4'S 18°37'E, 

Agulhas Bank, off the Cape of Good Hope, 150 fms] 

Fig. 54. Chlorotocus crassicornis (A. Costa, 1871). Photo by Tin-Yam Chan.


= Chlorotocus gracilipes var. andamanensis Alcock & Anderson, 1899: 284. [Andaman Sea, east of 

North Andaman Island, 13°17'15"N 93°10'15"E, 185 fms] 

Chlorotocus novaezealandiae (Borradaile, 1916) 

= Thalassocaris novae-zealandiae Borradaile, 1916: 84; Fig. 2. [7 miles E of North Cape, New Zealand, 

128 m] 

Dichelopandalus Caullery, 1896 

= Dichelopandalus Caullery, 1896 (type species Dichelopandalus Bonnieri Caullery, 1896, by monotypy, 


1957) 

Dichelopandalus bonnieri Caullery, 1896 

= Dichelopandalus Bonnieri Caullery, 1896: 379; Plate 15, figs 7-15. [45°30'N 5°53'W, 1200 m; 45°18'N 

5°23'W, 180 m; 45°18'N 5°23'W, 300-400 m; 47°13'N 5°56'W, 110 m (all Golfe de Gascogne)] 

Dichelopandalus leptocerus (Smith, 1881) 

= Pandalus leptocerus Smith, 1881: 437. [Fish Hawk stns 870 (off Block Island, R.I., 40°02'36"N 

70°22'58"W, 155 fms); 873 (off Newport, R.I., 40°02'N 70°57'W, 100 fms); 878 (off Block Island, R.I., 

39°55'N 70°54'15"W, 142 fms)] 

= Pandalus falcipes Spence Bate, 1888: 668; Plate 115, fig. 2. [Challenger stn 49, 43°03'N 63°39'W, south 

of Halifax, Nova Scotia, 85 fms] 

Dorodotes Spence Bate, 1888 

= Dorodotes Spence Bate, 1888 (type species Dorodotes reflexus Spence Bate, 1888, designated by 

Holthuis, 1955b, gender masculine; name placed on the Official List of Generic Names in Zoology 


Dorodotes reflexus Spence Bate, 1888: 678; Plate 116, fig. 3. [Challenger stns 195, 4°21'S 129°07'E, near 

Banda Island, 1425 fms; 205, 16°42'N 119°22'E, Philippine Islands, 1050 fms] 

Heterocarpus A. Milne-edwards, 1881b 

= Heterocarpus A. Milne-Edwards, 1881b (type species Heterocarpus ensifer A. Milne-Edwards, 1881b, 

by monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology in 


= Atlantocaris Ortmann, 1893 (type species Atlantocaris gigas Ortmann, 1893 (junior subjective synonym 

of Heterocarpus ensifer A. Milne-Edwards, 1881b), designated by Holthuis, 1955b, gender feminine) 

Heterocarpus abulbus Yang, Chan & Chu, 2010: 207; Figs 1, 3A-B, 4A-D, 5A. [Taiwan, Dasi fishing port, 

Yilan County, commercial trawler] 

Heterocarpus affinis Faxon, 1893: 204. [Albatross stns 3418 (16°30'00''N 099°52'30''W), 660 fms; 3424 

(21°15'00''N 106°23'00''W), 676 fms; 3425 (21°19'00''N 106°24'00''W), 680 fms] 

Heterocarpus agassizi Allen & Butler, 1994: 431; Figs 7-9. [Mid-Pacific Ocean, Darwin Guyot, 171°36.0'E 

22°07.7'N, 1281 m] 

Heterocarpus amacula Crosnier, 1988c: 72; Fig. 4a; Plate 1b; Plate 2h. [Polynésie française, Tuamotu 

(Mururoa), 250-600 m] 

Heterocarpus calmani Crosnier, 1988c: 59; Fig. 1a-h. [Madagascar, 12°52'S 48°10.3'E, 420-428 m] 

Heterocarpus chani Li, 2006b: 362; Figs 1-4. [Philippines, 8°52.1'N 123°37.1'E, 569-597 m depth] 

Heterocarpus corona Yang, Chan & Chu, 2010: 210; Figs 2, 3D-F, 4G-H, 5C-D. [Vanuatu, 15°41.1'S 

167°00.5'E, 509-659 m] 

Heterocarpus cutressi Monterrosa, 1988: 633; Figs 1-3. [southwest of Puerto Rico, 17°51.7'N 67°15.4'W, 777 

m] 

Heterocarpus dorsalis Spence Bate, 1888: 630; Plate 112. [Challenger stn 194, 4°34'00"S 129°57'30"E, off Banda 

Island, 200 fms] 

= Heterocarpus alphonsi Spence Bate, 1888: 632; Plate 112, fig. 1. [4°33'N 127°6'E, south of the Philippines, 

500 fms; 35°11'N 139°28'E, off Japan, 345 fms] 

= Heterocarpus affinis Borradaile, 1915b: 208; nec Faxon, 1893. [Saya de Malha, 300-500 fms]


Heterocarpus ensifer A. Milne-Edwards, 1881b: 8. [près des Barbades à 218 brasses] 

= Pandalus carinatus Smith, 1882: 63; Plate 10, figs 2-2f; Plate 11, figs 1-3. [Blake stn 327, 34°0'30"N 

76°10'30"W, 178 fms] 

= Atlantocaris gigas Ortmann, 1893: 80; Plate 5, fig. 3. [Plankton-Expedition der Humboldt-Stiftung, 

Südl. Aequatorialstrom, JN 209 (5.7°S 26.5°W, 0-400 m); Pl. 88 (6.6°S 24.5°W, 200 m)] 

= Procletes (?) atlanticus Lenz & Strunck, 1914: 334; Plate 22, figs 9-4. [nähe der Inseln Ascension, 

8°43'S 11°55'W, 3000 m] 

Heterocarpus gibbosus Spence Bate, 1888: 634; Plate 112, fig. 2. [Bohol Sea, 9°31.5'N 123°41.7'E, 336 m; 

neotype selection by Li et al., 2007] 

Heterocarpus grimaldii A. Milne-Edwards & Bouvier, 1900 

= Heterocarpus Grimaldii A. Milne-Edwards & Bouvier, 1900: 58. [entre Pico et São Jorge, 1330 m] 

Heterocarpus hayashii Crosnier, 1988c: 81; Figs 4b, 5b; Plate 1, fig. d; Plate 3, figs c-e. [îles Chesterfield Islands, 

19°40.86'S 158°46.0'E, 615-600 m] (Fig. 55) 

Heterocarpus hostilis Faxon, 1893: 204. [Albatross stns 3353 (07°06'15''N 080°34'00''W), 695 fms; 3363 

(05°43'00''N 085°50'00''W), 978 fms; 3364 (05°30'00''N 086°08'30''W), 902 fms; 3371 (05°26'20''N 

086°55'00''W), 770 fms; 3380 (04°03'00''N 081°31'00''W), 899 fms; 3393 (07°15'00''N 079°36'00''W), 

1020 fms; 3395 (07°30'36''N 078°39'00''W), 730 fms] 

Heterocarpus inopinatus Tavares, 1999: 673; Fig. 1. [19°38'S 38°43'W, 960 m] 

Heterocarpus intermedius Crosnier, 1999b: 346; Fig. 1. [MUSORSTOM 5, stn CP 363(19°47'53.9874''S 

158°44'17.9874''E), Chesterfield Islands, Coral Sea, 685-700 m] 

Heterocarpus laevigatus Spence Bate, 1888 

= Heterocarpus lævigatus Spence Bate, 1888: 636; Plate 112, fig. 3. [4°34'0"S 129°57'30"E, off Banda Island, 

200 fms] 

Heterocarpus lepidus De Man, 1917: 282. [West 1000 m distant from North point of Kabia-island reef, 

Flores Sea, 500 m; 5°53'.8S 132°48'8E, Kei-islands, 560 m] 

Heterocarpus longirostris MacGilchrist, 1905: 237. [Investigator stn 310, Bay of Bengal, 960 fms (13°29'30"N, 

95°29'E)] 

= Heterocarpus facetus Zarenkov & Khodkina, 1981: 83; Fig. 1. [central part of the Marcus-Nekker 

seamount chain, 1270-1320 m] 

Fig. 55. Heterocarpus hayashii Crosnier, 1988. Photo by Tin-Yam Chan.


Heterocarpus nasicus Timofeev, 1993: 40; Fig. 2. [Gulf of Aden, 11°31'3N 52°40'1E, 490-500 m] 

Heterocarpus oryx A. Milne-Edwards, 1881b: 10. [24°36'N 84°05'W, 955 brasses] 

Heterocarpus parvispina De Man, 1917 

= Heterocarpus ensifer var. parvispina De Man, 1917: 282. [7°35'.4S, 117°28'.6E, Bali Sea, 521 m; 6°8'N 

121°19'E, north of Sulu Island, 275 m; 5°40'S 132°26'E, off the Kei-islands, 310 m] 

Heterocarpus reedi Bahamonde, 1955: 106; Figs 1-4. [entres los Puertos de Valparaiso y Quinteros, Chili, 

329 m] 

Heterocarpus sibogae De Man, 1917: 283. [7°15'S 115°15'.6E, Bali Sea, 289 m; 7°35'.4S 117°28'.6E, Bali Sea, 

521 m; 5°3'.5S 119°0'E, southern entrance of Strait of Makassar, 450 m; 0°11'S 127°25'E, North of 

Batjan, 397 m; 5°54'.5S 120°19'.2E, West of Saleyer, 462 m; 5°26'.6S 132°32'.5E, Kei-islands, 397 m; 

5°53'.8S 132°48'.8E, Kei-islands, 560 m; 7°19.'4S 116°49'.5E, Bali Sea, 538 m] 

Heterocarpus signatus Rathbun, 1906: 918; Plate 21, fig. 6. [West coast of Hawaii Island, 382-253 fms] 

Heterocarpus tenuidentatus Cleva & Crosnier, 2006: 61; Figs 1-3. [Solomon Islands, New Georgia Sound, 

8°41.5'S 157°38.2'E, 814-980 m] 

Heterocarpus tricarinatus Alcock & Anderson, 1894: 154. [Investigator stn 122, Laccadive Sea, 12°05'35"N 

71°35'50"E, 880 fms] 

= Heterocarpus tricarinatus angustus Crosnier, 1988c: 84; Plate 4, Figs g-h. [5°39'S 122°12'E, au nord de 

l’île Muna, 1886 m] 

Heterocarpus vicarius Faxon, 1893: 203. [Albatross stns 3385 (07°32'36''N 079°16'00''W), 286 fms; 3386 

(07°33'12''N 079°17'15''W), 242 fms; 3389 (07°16'45''N 079°56'30''W), 210 fms; 3396 (07°32'00''N 

078°36'30''W), 259 fms] 

Heterocarpus woodmasoni Alcock, 1901 

= Heterocarpus Wood-masoni Alcock, 1901: 108. [Andaman Sea, 265 fms] 

Miropandalus Bruce, 1983g 

= Miropandalus Bruce, 1983g (type species Miropandalus hardingi Bruce, 1983g, by original designation 


Miropandalus hardingi Bruce, 1983g: 483; Figs 1-5. [Enewetak Island, Marshall Islands, 23 m] 

Notopandalus yaldwyn, 1960 

= Notopandalus Yaldwyn, 1960 (type species Pandalus magnoculus Spence Bate, 1888, by original designation 


Notopandalus magnoculus (Spence Bate, 1888) 

= Pandalus magnoculus Spence Bate, 1888: 667; Plate 115, fig. 1. [38°50'S 169°20'E, near New Zealand, 

275 fms; 39°32'S 171°48'E, off the west coast of New Zealand, 150 fms] 

Pandalina Calman, 1899b 

= Pandalina Calman, 1899b (type species Pandalus brevirostris Rathke, 1843, by original designation 

and monotypy, gender feminine; name placed on the Official List of Generic Names in Zoology in 


Pandalina brevirostris (Rathke, 1843) 

= Pandalus brevirostris Rathke, 1843: 17. [Molde] 

= Hippolyte Thompsoni Bell, 1851 [in Bell, 1844-1853]: 290; unnumbered vignette [?Belfast] 

= Pandalus Rathkii Heller, 1863b: 441; Plate 3, fig. 31. [Lissa und Lesina in 20-30 Faden Tiefe] 

Pandalina modesta (Spence Bate, 1888) 

= Pandalus modestus Spence Bate, 1888: 670; Plate 114, fig. 4. [35°4'S 18°37'E, at the Agulhas Bank, off 

the Cape of Good Hope, 150 fms] 

Pandalina nana Burukovsky, 1990: 201; Fig. 4A. [25°03'S 97°27'W, 271 m] 

Pandalina profunda Holthuis, 1946b: 281; Fig. 1a-c. [Barents Sea; Bergen, Norway] 

Pandalina spinicauda Komai & Chan, 2010b: 626; Figs 1-2, 7A. [Mozambique Channel, MAINBAZA stn 

CP 3130, 25°53'S 33°07'E, 112-127 m]


Pandalopsis Spence Bate, 1888 

= Pandalopsis Spence Bate, 1888 (type species Pandalopsis amplus Spence Bate, 1888, by monotypy, 


1959) 

Pandalopsis aleutica Rathbun, 1902a: 901. [off Segouam, Aleutians, 283 fms] 

Pandalopsis ampla Spence Bate, 1888 

= Pandalopsis amplus Spence Bate, 1888: 671; Plate 115, fig. 3. [37°17'S 53°52'W, off Monte Video, 

600 fms] 

Pandalopsis coccinata Urita, 1941: 12; Figs 1-2. [off Tottori-mura, near Kushiro, Hokkaido, Japan] 

Pandalopsis dispar Rathbun, 1902a: 903. [Chernoffski Harbor, Unalaska, 109 fms] 

Pandalopsis gibba Komai & Takeda, 2002: 91; Figs 1-5. [Sagami Bay, 35°04.68'N 139°45.32'E, 250 m] 

Pandalopsis glabra Kobjakova, 1936b 

= Pandalopsis aleutica var. glabra Kobjakova, 1936b: 192; Fig. 4. [Ochotsk Meer, 515 and 1076 m] 

Pandalopsis japonica Balss, 1914b 

= Pandalopsis dispar var. japonica Balss, 1914b: 32. [Etschiu] 

Pandalopsis lamelligera (Brandt, 1851) 

= Pandalus lamelligerus Brandt, 1851: 124; Plate 5, figs 20-20b. [von der Chantarischen Bäreninsel; im 

Ochotskischen Meer] 

Pandalopsis longipes Komai, 1994: 545; Figs 4-5. [off Nemuro, Hokkaido] 

Pandalopsis longirostris Rathbun, 1902a: 902. [off Iliuliuk Harbor, Unalaska, 309 fms] 

Pandalopsis lucidirimicola Jensen, 1998: 82; Figs 1-3. [Point Atkinson, Burrard Inlet, British Colombia, 

Canada, 49°19'N 123°15'W, 18 m] 

Pandalopsis miyakei Hayashi in Baba, Hayashi & Toriyama, 1986: 123; Fig. 19; unnumbered plate. [Tosa 

Bay, 700 m] 

Pandalopsis multidentatus Kobjakova, 1936b: 188; Fig. 1. [Japanisches Meer von der Peter-des-Großen- 

Bucht nördlich bis zum Kap Terpenjia (Tataren-Sund), 64-500 m] 

Pandalopsis ochotensis Kobjakova, 1936b: 190; Fig. 2. [Ochotsk Meer, 500-504 m] 

Pandalopsis pacifica (Doflein, 1902) 

= Pandalus pacificus Doflein, 1902: 619; Plate 4, fig. 1. [Nemuro, Nordspitze der Insel Yesso] 

= Pandalopsis mitsukurii Rathbun, 1902b: 48; Figs 22-23. [Mororan, Hokkaido] 

Pandalopsis profundus Zarenkov, 1971: 189; Fig. 3(11-20). [south of Japan, 820 m] 

Pandalopsis punctata Kobjakova, 1936b 

= Pandalopsis punctatus Kobjakova, 1936b: 191; Fig. 3. [Ochotsk Meer, 165 m] 

Pandalopsis rubra Komai, 1994: 553; Figs 1, 9-10. [eastern Hokkaido, 42°46.0'N 145°19.5'E, 1080-1120 m] 

Pandalopsis spinosior Hanamura, Khono & Sakaji, 2000: 27; Figs 1-3. [Urup (or Etorofu) Strait, around 

400 m] 

Pandalopsis zarenkovi Ivanov & Sokolov, 2001: 160; Figs 1-3. [Bering Sea, 61°43.4'N 177°37.8'W, 362 m] 

Pandalus leach, 1814 [in leach, 1813-1814] 

= Pandalus Leach, 1814 [in Leach, 1813-1814] (type species Pandalus Montagui Leach, 1814 [in Leach, 

1813-1814], by monotypy, gender masculine; name placed on the Official List of Generic Names in 

Zoology in Opinion 104 in 1928 and in Direction 47 in 1956) 

= Dyma Krøyer, 1861 (type species Dymas typus Krøyer, 1861, by monotypy and by indication under 

Art. 68c of the ICZN) 

= Boreocaris Ortmann, 1893 (type species Boreocaris möbiusi Ortmann, 1893 (probably a junior subjective 

synonym of Pandalus Montagui Leach, 1814 [in Leach, 1815-1875]), gender feminine) 

Pandalus borealis Krøyer, 1838: 254. [Sydpröven, Julianehaab, Fiskæsset, &c] 

= Dymas typus Krøyer, 1861: 63. [Grønland] 

= Pandalus borealis v. edenticulatus Retowsky, 1946: 300; Figs 2a-d. [Arctic Ocean] 

Pandalus chani Komai, 1999: 1337; Figs 33-35, 44B. [Su-Aou, I-Lan County, northeast of Taiwan, circa 

400 m] 

Pandalus curvatus Komai, 1999: 1302; Figs 16-17. [southern Japan, Amakusa-nada, western Kyushu, 

32°08.6'N 129°41.2'E, 501 m]


Pandalus danae Stimpson, 1857 

= Pandalus danæ Stimpson, 1857: 87; Plate 21, figs 6-7. [opposite Fort Townsend, Puget Sound] 

= Pandalus franciscorum Kingsley, 1878a: 94. [San Francisco, California] 

Pandalus eous Makarov, 1935 

= Pandalus borealis var. eous Makarov, 1935: 321; Figs 2, 3b. [im Bering-Meere nahe dem Nawarin- 

Kap, 61°33'N 179°09'W, 128 m; Bering-Meer bei dem Kap Oljutorsky, 59°48'N 170°43'E, 142 m; nahe 

der Bering-Insel, 52°25'N 165°44'E, 234 m] 

Pandalus formosanus Komai, 1999: 1342; Figs 36-37, 44C. [Su-Aou, I-Lan County, NE Taiwan, about 

500 m] 

Pandalus goniurus Stimpson, 1860a: 36. [in sinu ”Avatska” Kamtschatkae, in fundo limoso prof. 10 org] 

= Pandalus dapifer Murdoch, 1884: 519. [off Point Franklin, 13.5 fms] 

Pandalus gracilis Stimpson, 1860a: 37. [Hakodate Bay, Hokkaido Island, Japan; lectotype designation by 


= Pandalus robustus Stimpson, 1860a: 37. [sinu ”Hakodadi”, in profundis] 

= Pandalus stimpsoni Thallwitz, 1892: 3. [Japan oder China] 

Pandalus gurneyi Stimpson, 1871 

= Pandalus Gurneyi Stimpson, 1871: 128. [Monterey, Cal.] 

Pandalus hypsinotus Brandt, 1851: 125. [Unalaschka] 

Pandalus ivanovi Komai & Eletskaya, 2008: 47; Figs 1-6. [off Eastern Sakhalin, Sea of Okhotsk, 46°N 

145°E, 150-200 m] 

Pandalus jordani Rathbun, 1902a: 900. [off Santa Cruz Island, California, 155 fms] 

Pandalus latirostris Rathbun, 1902b: 46; Figs 20-21. [Mororan, Hokkaido] 

= Pandalus Kessleri Czerniavsky, 1878: 23. [nomen nudum] 

Pandalus montagui Leach, 1814 [in Leach, 1813-1814] 

= Pandalus Montagui Leach, 1814 [in Leach, 1813-1814]: 432. [Zetland, Scotland; lectotype designation 

by Holthuis, 1956b] 

= Astacus maculatus Leach, 1814 [in Leach, 1813-1814]: 432. [nomen nudum] 

= Pandalus annulicornis Leach, 1815 [in Leach, 1815-1875]: Plate 40. [Zetland, Scotland; lectotype 

designation by Holthuis, 1956b] 

? = Pandalus levigatus Stimpson, 1854: 58. [Grand Manan] 

= P.[andalus] Leptorhynchus Kinahan, 1858b: 41; unnumbered figures. [Sandycove, prope Dublin] 

= P.[andalus] Leptorhynchus Kinahan, 1860b: 80; unnumbered figures. [Sandycove, prope Dublin] 

? = Boreocaris möbiusi Ortmann, 1893: 84; Plate 6, fig. 3; Plate 7, fig. 1. [Plankton-Expedition der 

Humboldt-Stiftung, Golfstrom, JN 4 (59.2°N 11.8°W, 0-400 m)] 

Pandalus nipponensis Yokoya, 1933: 16; Fig. 5. [off Tohtomi, Suruga Bay, 100 m; lectotype designation by 

Komai, 1999] 

Pandalus platyceros Brandt, 1851: 123. [Insel Unalaschka] 

= Pandalus pubesculentus Dana, 1852a: 24. [in freta ”de Fuca” Oregoniæ, juxta portum ”Dungeness”] 

Pandalus prensor Stimpson, 1860a: 37. [sinu ”Hakodadi”, fundo conchoso, prof. 8 org.] 

= Pandalus hypsinotus var. meridionalis Balss, 1914b: 29. [Nagasaki bis Nemuro und Wladiwostok] 

Pandalus stenolepis Rathbun, 1902a: 901. [Straits of Fuca, 40 fms] 

Pandalus teraoi Kubo, 1937a: 98; Figs 4-6. [off Ôyama, Prefecture Aiti] 

Pandalus tridens Rathbun, 1902a 

= Pandalus montagui tridens Rathbun, 1902a: 901. [off North Head, Akutan Island, Alaska, 72 fms] 

Pantomus A. Milne-edwards, 1883 

= Pantomus A. Milne-Edwards, 1883 (type species Pantomus parvulus A. Milne-Edwards, 1883, by 



Pantomus affinis Chace, 1937b: 116; Fig. 3a-b. [Santa Inez Bay, 26°56'30''N 111°48'30''W, 60 fms] 

Pantomus parvulus A. Milne-Edwards, 1883: Plate 26. [North part of Yucatan Bank, 23°13'N 89°16'W, 84 

brasses]


Peripandalus De Man, 1917 

= Peripandalus De Man, 1917 (type species Pandalus serratus A. Milne-Edwards, 1873, by monotypy, 


Peripandalus serratus (A. Milne-Edwards, 1873) 

= Pandalus serratus A. Milne-Edwards, 1873: 87. [Upolu] 

Plesionika Spence Bate, 1888 

= Nisea Risso, 1844 (type species Nisea formosa Risso, 1844 (junior subjective synonym of Astacus 

Narual Fabricius, 1787), gender feminine; nomen nudum, see Holthuis, 1977) 

= Plesionika Spence Bate, 1888 (type species Plesionika uniproducta Spence Bate, 1888 (junior subjective 

synonym of Acanthephyra ensis A. Milne-Edwards, 1881b), designated by Alcock, 1901, gender 


= Nothocaris Spence Bate, 1888 (type species Nothocaris rostricrescentis Spence Bate, 1888, designated 

by Fowler, 1912, gender feminine) 

= Parapandalus Borradaile, 1900 (type species Pandalus (Parapandalus) serratifrons Borradaile, 1900, 

designated by Alcock, 1901, gender masculine; name placed on the Official List of Generic Names 


= Parapandalopsis Dong, Chen & Wang, 1986 (type species Plesionika spinipes Spence Bate, 1888, by 

monotypy, gender feminine; see Holthuis, 1993a) 

Plesionika acanthonotus (Smith, 1882) 

= Pandalus acanthonotus Smith, 1882: 61; Plate 13, figs 10-11. [Blake stn 321, 32°43'25"N 77°20'30"W, 

233 fms] 

= Pandalus Parfaitii A. Milne-Edwards, 1883: Plate 21. [East Atlantic, 44°4'30"N 9°25'W (of Paris), 

119 m] 

= Pandalus geniculatus A. Milne-Edwards, 1883: Plate 25. [East Atlantic, 44°4'30"N 9°25'W (of Paris), 

119 m] 

Plesionika acinacifer Chace, 1985: 51; Figs 21-23. [Between southern Cebu and Siquijor, Philippines, 

9°27'15"N 123°31'48"E, 790 m] 

Plesionika adensameri (Balss, 1914c) 

= Parapandalus Adensameri Balss, 1914c: 134. [Rote Meer, zwischen 18 und 26° n. Br., von 800 bis 1300 

m; fully described in Balss, 1915] 

Plesionika albocristata Chan & Chuang, 2002: 611; Figs 1-2. [Taiwan, NE coast, Su-Aou fishing port, I-Lan 

County, about 350 m] 

Plesionika alcocki (Anderson, 1896) 

= Pandalus alcocki Anderson, 1896: 92. [9°34'57"N 75°36'30"E, 406 fms] 

= Plesionika adenensis Timofeev, 1993: 44; Fig. 4. [Gulf of Aden, 12°18'7N 44°28'E, 610 m] 

Plesionika alexandri (A. Milne-Edwards, 1883) 

= Heterocarpus Alexandri A. Milne-Edwards, 1883: Plate 28. [north of Havana, 23°14'00''N 82°25'00''W, 

805 fms] 

Plesionika antigai Zariquiey Álvarez, 1955: 11. [Barcelona, barcas de arrastre, más de 300 m] 

Plesionika assimilis De Man, 1917: 280. [Madura-bay and other localities in the southern part of Molostrait, 

54-90 m] 

Plesionika beebei Chace, 1937b: 114; Fig. 2. [13 miles northeast by north of San Ildefonso Island, 26°48'40''N 

111°20'30''W, 300 fms] 

Plesionika bifurca Alcock & Anderson, 1894: 155. [Investigator stn 177, Laccadive Sea, 13°47'49"N 73°07'E, 

636 fms] 

Plesionika bimaculata Chan, 2004: 295; Figs 1, 10. [Vanuatu, 20°18.87'S 169°53.12'E, 250-315 m] 

Plesionika binoculus (Spence Bate, 1888) 

= Nothocaris binoculus Spence Bate, 1888: 656; Plate 114, fig. 2. [8°56'S 136°05'E, Arafura Sea, south of 

New Guinea, 49 fms] 

Plesionika brevipes (Crosnier & Forest, 1968) 

= Parapandalus brevipes Crosnier & Forest, 1968: 1136; Fig. 5. [au large de Pointe-Noir, Congo, par 

5°00'S 11°22'E et 5°06'S 11°26'E, 190-355 m]


Plesionika brevirostris Spence Bate, 1888: 650; Plate 113, fig. 5. [6°47'N 122°28'E, between the Philippine 

Islands and Borneo, 250 fms] 

Plesionika carinata Holthuis, 1951a: 59; Fig. 12. [off Angola, 7°55'S 12°38'E, 235-460 m] 

Plesionika carinirostris Hendrickx, 1990b: 38; Figs 2-3. [off Isla San Lorenzo, 28°46'N 112°54'W, 360-380 m] 

Plesionika carsini Crosnier, 1986c: 369; Fig. 3. [Iles Tuamotu, Mururoa, 450 m] 

Plesionika chwitchii Burukovsky, 1978: 1729; Figs 1-4 [south-west Atlantic, 33°26'S 50°58'W, 130 m] 

Plesionika costelloi (Yaldwyn, 1971) 

= Parapandalus costelloi Yaldwyn, 1971: 91. [off Astrolabe Reef, Bay of Plenty, 65 fms] 

Plesionika crosnieri Chan & Yu, 1991b: 546; Figs 1, 3a. [Ta-Chi, I-Lan County, northeastern Taiwan, fish 

market] 

? = Plesionika dentirostris Tung, Wang & Li, 1988: 20; Fig. 1. [East China Sea, 400 m] 

Plesionika curvata Chan & Crosnier, 1991: 440; Figs 10b, 11d-f, 33. [Tubuai, French Polynesia, 23°40'S 

149°40'W, 200 m] 

Plesionika echinicola Chan & Crosnier, 1991: 416; Figs 1a, 2a, 3a-b, 19-20. [New Caledonia, 24°47.0'S 

168°09.4'E, 274 m] 

Plesionika edwardsii (Brandt, 1851) 

= Pandalus Guerinii Risso, 1844: 95. [nomen nudum] 

= Pandalus (Pontophilus) Edwardsii Brandt, 1851: 122; nomen novum for Pandalus narval H. Milne 

Edwards, 1841 [in H. Milne Edwards, 1836-1844] nec Astacus Narual Fabricius, 1787. [Mediterranean, 

type locality designated by Chan & Yu, 1991b] 

= Pandalus (Parapandalus) longirostris Borradaile, 1900: 413; Plate 37, fig. 10a-h. [New Britain] 

Plesionika ensis (A. Milne-Edwards, 1881b) 

= Acanthephyra ensis A. Milne-Edwards, 1881b: 14. [près de Barbades, 237 brasses] 

= Plesionika uniproducta Spence Bate, 1888: 641 (partim); Plate 113, fig. 1. [9°5'S 34°50'W, off Barra 

Grande, 350 fms] 

Plesionika erythrocyclus Chan & Crosnier, 1997: 214; Figs 16, 34-35. [Tuamotu Islands, Mururoa, French 

Polynesia, 350-600 m] 

Plesionika exigua (Rathbun, 1906) 

= Pandalus exiguus Rathbun, 1906: 916; Plate 21, fig. 2. [vicinity of Kauai, Hawaii Islands, 233-40 fms] 

Plesionika fenneri Crosnier, 1986d: 691. [nomen novum for Plesionika chacei Crosnier, 1986c] 

= Plesionika chacei Crosnier, 1986c: 363; Figs 1a-b, 2a-h; nec Hayashi in Baba, Hayashi & Toriyama, 

1986. [Iles Australes, Tubuai, 700 m] 

Plesionika fimbriata Chace, 1985: 63; Fig. 27. [Samar Sea east of Masbate, Philippines, 12°13'15"N 

124°05'03"E, 216 m] 

Plesionika flavicauda Chan & Crosnier, 1991: 438; Figs 10a, 11a-c, 29-32. [Rurutu, French Polynesia, 

22°27.8'S 151°22.9'W, 240-260 m] 

Plesionika gigliolii (Senna, 1902) 

= Pandalus Gigliolii Senna, 1902: 315; Plate 16, Figs 5-16. [39°15'37''3'''N 9°26'37''7'''E, presso il capo 

Carbonara, Sardegna, 508 m] 

= Pandalus subtilirostris Riggio, 1905 [in Riggio, 1904-1905]: 283. [mare di Messina] 

Plesionika gracilis (Zarenkov, 1971) 

= Parapandals (sic) gracilis Zarenkov, 1971: 187; Fig. 3.1-3.10. [South China Sea, off Vietnam, 38 m] 

Plesionika grahami Kensley, Tranter & Griffin, 1987: 315; Figs 20-21. [New South Wales, north-east of Port 

Jackson, 33°44'S 151°49'E to 33°42'S 151°50'E, 405 m] 

Plesionika grandis Doflein, 1902 

= Plesionika spinipes var. grandis Doflein, 1902: 618; Plate 3, figs 3-5. [Sagamibai] 

Plesionika heterocarpus (A. Costa, 1871) 

= Pandalus heterocarpus A. Costa, 1871: 89; Plate 2, fig. 3. [golfo di Napoli] 

= Pandalus sagittarius A. Milne-Edwards, 1883: Plate 23. [38°3'N 11°32'E (of Paris), 370 m] 

= Pandalus longicarpus A. Milne-Edwards, 1883: Plate 25. [44°N 9°17'E (of Paris), 156 m] 

Plesionika holthuisi Crosnier & Forest, 1968: 1141; Fig. 7b-c. [Ombango stn 394, off Pointe Noir, Congo, 

500-505 m]


Plesionika hsuehyui Chan, 2004: 300; Figs 3, 12-13. [Taiwan, Tai-Shi fishing port, NE coast, I-Lan County] 

Plesionika hypanodon Doflein, 1902: 615; Plate 3, fig. 1. [Sagamibai] 

Plesionika indica De Man, 1917 

= Plesionika longipes var. indica De Man, 1917: 279. [5°40'S 132°26'E, near the Kei-islands, 310 m] 

Plesionika intermedia Chace, 1985: 72; Fig. 33. [Tañon Straight, east of Negros, Philippines, 10°29'45"N 

123°31'15"E, 472 m] 

Plesionika izumiae Omori, 1971: 242; Figs 1-2; Plate 1. [34°54.1'N 138°24.7'E, 45 m] 

Plesionika kensleyi Chace, 1985: 77; Figs 35-36. [Mindanao Sea, off Murcielagos Bay, Mindanao Philippines, 

8°45'30"N 123°33'45"E, 309 m] 

Plesionika laevis (A. Milne-Edwards, 1883) 

= Heterocarpus laevis A. Milne-Edwards, 1883: Plate 28. [off Martinique, 14°43'48''N 61°11'25''W, 169 

fms] 

Plesionika laurentae Chan & Crosnier, 1991: 431; Figs 6, 24. [îles Chesterfield, 19°25.49'S 158°37.96'E: 215- 

217 m] 

Plesionika longicauda (Rathbun, 1901) 

= Pandalus longicauda Rathbun, 1901: 117; Fig. 24. [Gulf of Mexico, 28°42'30''N 85°29'W, 88 fms] 

Plesionika longidactylus Li & Komai, 2003: 262; Figs 2-5. [northern South China Sea, 17°30'N 109°30'E, 

106 m] 

Plesionika longipes (A. Milne-Edwards, 1881b) 

= Pandalus longipes A. Milne-Edwards, 1881b: 15. [près de Barbades, 204 brasses; lectotype designation 

by Forest & Holthuis, 1997] 

Plesionika lophotes Chace, 1985: 81; Fig. 37. [Samar Sea between southeastern Masbate and Almagro Island, 

Philippines, 11°57'27"N 124°10'42"E, 245 m] 

Plesionika macropoda Chace, 1939: 37. [Bahia de Cochinos, Santa Clara Province, Cuba, 22°07'N 81°08'W, 

195-235 fms] 

Plesionika martia (A. Milne-Edwards, 1883) 

= Pandalus martius A. Milne-Edwards, 1883: Plate 21. [Atlantic Ocean, roughly between 34° and 

44°N, 512-1290 m; see Forest & Holthuis, 1997] 

= Plesionika uniproducta Spence Bate, 1888: 641 (partim); Plate 113, fig. 1. [9°5'S 34°50'W, off Barra 

Grande, 350 fms] 

= Plesionika (Pandalus) sicherii Riggio, 1900: 20. [mare Siculo] 

Plesionika miles (A. Milne-Edwards, 1883) 

= Pandalus miles A. Milne-Edwards, 1883: Plate 18. [Dominica, 98 fms; see Forest & Holthuis, 1997] 

Plesionika minor Calman, 1939: 199; Fig. 2. [John Murray Expedition, stn 75 (25°10'48''N 56°47'30''E to 

25°09'48''N 56°47'30''E), Gulf of Oman, 210 m] 

Plesionika multispinosa (Zarenkov, 1971) 

= Parapandalus multispinosus Zarenkov, 1971: 185; Fig. 2(11-20). [off Easter Island, 80-150 m] 

Plesionika narval (Fabricius, 1787) 

= Astacus Narual Fabricius, 1787: 331. [in mari mediterraneo] 

= P.[alemon] Pristis Risso, 1816: 105. [sur les fonds rocailleux, environs de Nice] 

= Palaemon tarentinum O.G. Costa, 1844b: 9. [Mare di Taranto] 

= Nisea formosa Risso, 1844: 95. [nomen nudum] 

= Pandalus escatilis Stimpson, 1860a: 37. [prope insulam Madeirae, in profundis] 

? = Pandalus stylopus A. Milne-Edwards, 1883: Plate 19. [34°11'30''N 9°59'E (of Paris), 530 m] 

Plesionika neon Komai & Chan, 2010b: 631; Figs 3-4, 7B. [Mozambique Channel, MAINBAZA st CP 3130, 

25°53'S 33°07'E, 112-127 m] 

Plesionika nesisi (Burukovsky, 1986) 

= Heterocarpus nesisi Burukovsky, 1986: 62; Fig. 1. [East Pacific sea-mount, 13°34'N 120°33'W, 800 m] 

Plesionika ocellus (Spence Bate, 1888) 

= Nothocaris ocellus Spence Bate, 1888: 657; Plate 114, fig. 3. [7°03'N 121°48'E, near Samboangan, 

Philippine Islands, 82 fms] 

= Plesionika chacei Hayashi in Baba, Hayashi & Toriyama, 1986: 129; Fig. 20; unnumbered plate. [off 

Kagoshima Prefecture, 107-110 m]


Plesionika orientalis Chace, 1985 

? = Plesionika cottei Kotte, 1903: 622; Plates 23-26. [im Indischen Ocean, von der Suaheliküste, 630 m] 

= Plesionika martia orientalis Chace, 1985: 84; Figs 38-39. [Sulu Archipelago, between Jolo and Tawitawi 

Islands, 5°48'00"N 129°33'45"E, 490 m] 

= Plesionika semilævis Spence Bate, 1888: 644. (partim) 

Plesionika ortmanni Doflein, 1902: 616; Plate 3, fig. 2. [Sagamibai] 

Plesionika pacifica Edmondson, 1952 

= Plesionika pacificus Edmondson, 1952: 67; Fig. 1. [off the north Kona coast of Hawaii, little more 

than 100 fms, from stomach of fish, the opakapaka (Pristipomoides sp.)] 

Plesionika parvimartia Chace, 1985: 93; Figs 42-43. [Samar Sea south of Tagapula Island, 11°57'27"N 

124°10'42"E, 245 m] 

Plesionika payeni Chan & Crosnier, 1997: 221; Figs 19, 39. [Tuamoto Islands, Mururoa, French Polynesia, 

21°47.6'S 138°55.7'W, 200 m] 

Plesionika persica (Kemp, 1925) 

= Parapandalus persicus Kemp, 1925: 273; Fig. 8. [Gulf of Oman, 23°44'30"N 58°52'15"E, 261 fms] 

= Parapandalus filipes Calman, 1939: 202; Fig. 3. [John Murray Expedition, Red Sea, stns 5 (18°59'00''N 

39°13'30''E to 18°49'18''N 38°23'42''E), 500-0 m; 76B (24°13'54''N 59°03'30''E to 24°16'42''N 59°06'06''E), 

South Arabian coast, 600-2500 m] 

Plesionika philippinensis Chace, 1985: 97; Fig. 44. [Northern Surigao Strait between Cabugan Grande and 

Hibuson Islands, Philippines, 10°27'30"N 125°19'15"E, 135 m] 

Plesionika picta Chan & Crosnier, 1997: 223; Figs 20, 40. [Rurutu, Tubuai Islands, French Polynesia, 

22°27.8'S 151°23.0'W, 570 m] 

Plesionika polyacanthomerus Pequegnat, 1970: 97; Figs 4-13, 4-14. [Northeastern Gulf of Mexico, 28°51'N 

88°47.5'W, 472-289 fms] 

Plesionika poupini Chan & Crosnier, 1997: 189; Figs 1-2, 22. [Tubuai Islands, Raevavae, 23°55.0'S 

147°40.0'W, 450 m] 

Plesionika protati Chan & Crosnier, 1997: 219; Figs 16, 38. [Tahuata, Marquesas Islands, French Polynesia, 

9°54.5'S 139°07.9'W, 190 m] 

Plesionika pumila Chace, 1985: 100; Figs 45-46. [Western part of Basilan Strait, northwest of Basilan Island, 

Sulu Archipelago, 6°44'45"N 121°48'E, 46 m] 

Plesionika quasigrandis Chace, 1985: 104; Figs 47-48. [Davao Gulf about 2 miles (3.2 km) off Davao, Mindanao, 

Philippines, 7°02'N 125°38'45"E, 247 m] 

= Parapandalus yugniroi Timofeev, 1993: 42; Fig. 3. [Gulf of Aden, 14°53'2N 50°32'9E, 240-250 m] 

Plesionika reflexa Chace, 1985: 108; Fig. 49. [East of southern Luzon, northeast of San Bernardino Strait, 

Philippines, 12°56'24"N 124°25'24"E, 494 m] 

Plesionika rossignoli Crosnier & Forest, 1968: 1139; Fig. 6b. [Ombango stn 323 (off St. P. de Loanda, Angola, 

9°S 9°E)] 

Plesionika rostricrescentis (Spence Bate, 1888) 

= Nothocaris rostricrescentis Spence Bate, 1888: 653; Plate 114, fig. 1. [5°49'15"S 132°14'15"E, off the Ki 

Islands, south of New Guinea, 140 fms] 

Plesionika rubrior Chan & Crosnier, 1991: 433; Figs 7, 25-28. [Maiao, French Polynesia, 17°38.6'S 

150°39.0'W, 320 m] 

Plesionika rufomaculata Chan, 2004: 309; Figs 7, 16. [Loyalty Islands, 20°41.8'S 167°0.2'E, 282 m] 

Plesionika sanctaecatalinae Wicksten, 1983b: 138; Figs 1-3. [7.3 miles, 17° true from Long Point Light, Santa 

Catalina Island, California, 33°33'29''N 18°20'42''W – 33°30'12''N 118°17'28''W, 886 m] 

Plesionika scopifera Chan, 2004: 311; Figs 8-9, 17. [Chesterfield Islands, 19°48.6'S 158°29.1'E, 260-270 m] 

Plesionika semilaevis Spence Bate, 1888 

= Plesionika semilævis Spence Bate, 1888; Plate 113, fig. 3. [Moro Gul east of Basilan Strait, Philippines, 

6°47'N 122°28'E, 457 m; lectotype designation by Chace, 1985] 

Plesionika serratifrons (Borradaile, 1900) 

= Pandalus (Parapandalus) serratifrons Borradaile, 1900: 411; Plate 37, figs 8a-8d. [Blanche Bay, New 

Britain, 50-100 fms]


= Pandalus (Parapandalus) tenuipes Borradaile, 1900: 412; Plate 37, fig. 9; nec Smith, 1881. [Blanche 

Bay, New Britain; D’Entrecasteaux Group, British New Guinea] 

Plesionika simulatrix Chace, 1985: 125; Fig. 57. [Samar Sea, east of Masbate, Philippines, 12°13'15"N 

124°05'03"E, 216 m] 

Plesionika sindoi (Rathbun, 1906) 

= Pandalus sindoi Rathbun, 1906: 915; Plate 21, fig. 4. [Albatross stn 3998, vicinity of Kauai Island 

(21°56'25''N 159°48'35''W), 235-228 fms] 

Plesionika spinensis Chace, 1985: 128; Figs 58-59. [Western Mindanao Sea, off Murcielagos Bay, Philippines, 

8°45'30"N 123°33'45"E, 309 m] 

Plesionika spinidorsalis (Rathbun, 1906) 

= Pandalus spinidorsalis Rathbun, 1906: 917; Plate 21, fig. 5. [Albatross stn 3986 (22°04'35''N 

159°18'40''W, 55-362 fms), Vicinity of Kauai Island] 

Plesionika spinipes Spence Bate, 1888: 646; Plate 113, fig. 2. [1°54'0"S 146°39'40"E, north of New Guinea, 

150 fms] 

Plesionika suffusa Chan, 2004: 305; Figs 5, 14. [New Caledonia, 18°57.8'S 163°14'E, 425 m] 

Plesionika taiwanica Chan & Yu, 2000: 120; Figs 1-3. [Taiwan, southwestern coast, fishing port at Tungkang, 

Pingtong County, about 150 m] 

Plesionika tenuipes (Smith, 1881) 

= Pandalus tenuipes Smith, 1881: 441. [Fish Hawk stns 870 (40°02'36''N 70°22'58''W, 155 fms), 871 

(40°02'54''N 70°23'40''W, 115 fms), 873 (40°02'00''N 70°57'00''W, 100 fms), 877 (39°56'00''N 70°54'18''W, 

126 fms), 880 (39°48'30''N 70°54'00''W, 252 fms); all off Block Island, 100-252 fms] 

Plesionika trispinus Squires & Barragan, 1976: 113; Figs 1-2. [off Pizarro, Colombia, 05°10'N 77°25'W, 257- 

275 m] 

Plesionika unicarinatus (Borradaile, 1915b) 

= Heterocarpus unicarinatus Borradaile, 1915b: 208. [Providence Island, 637-665 fms] 

Plesionika unidens Spence Bate, 1888: 648; Plate 113, fig. 4. [1°54'00"S 146°39'48"E, north of New Guinea, 

150 fms] 

= Plesionika affinis Alcock & Anderson, 1899: 285. [Investigator stn 236 (Andaman Sea, 13°59'N 93°E), 

172-303 fms] 

Plesionika williamsi Forest, 1964: 620; Figs 1-4. [4°39'N 2°46'W, 300-400 m] 

? = Plesionika dentirostris Tung, Wang & Li, 1988: 20; Fig. 1. [East China Sea, 400 m] 

= Plesionika crosnieri Burukovsky, 1992: 145; Figs 1-4; nec Chan & Yu, 1991b. [25°39'S 86°51'E, 583- 

600 m] 

= Plesionika alaini Burukovsky, 1993c: 18. [nomen novum for Plesionika crosnieri Burukovsky, 1992] 

Plesionika willisi (Pequegnat, 1970) 

= Parapandalus willisi Pequegnat, 1970: 87; Figs 4.9-4.10. [Northeastern Gulf of Mexico, 29°27.6'N 

86°46.5'W, 210 fms] 

Plesionika yui Chan & Crosnier, 1991: 428; Figs 4b, 5c, e, g-h, 23. [Tong-Kong, Ping-Tong County, SW 

Taiwan] 

Procletes Spence Bate, 1888 

= Procletes Spence Bate, 1888 (type species Procletes biangulatus Spence Bate, 1888 (junior subjective 

synonym of Dorodotes levicarina Spence Bate, 1888), designated by Holthuis, 1955b, gender masculine) 

= Heterocarpoides De Man, 1917 (type species Dorodotes levicarina Spence Bate, 1888, by monotypy, 


1957) 

Procletes levicarina (Spence Bate, 1888) (Fig. 56) 

= Dorodotes levicarina Spence Bate, 1888: 680; Plate 112, fig. 5. [9°59'S 139°42'E, Arafura Sea, south of 

Papua, 28 fms] 

= Procletes biangulatus Spence Bate, 1888: 884; Plate 127, fig. 4. [Challenger stn 191, 5°41'0''S 134°4'30''E, 

off the Arrou islands, 800 fms]


? = Heterocarpus (Heterocapoides) [sic] glabrus Zarenkov, 1971: 193; Fig. 4(16-27). [South-China Sea, off 

S Vietnam, 75 m] 

Fig. 56. Procletes levicarina (Spence Bate, 1888). Photo by Tin-Yam Chan. 

Pseudopandalus Crosnier, 1997 

= Pseudopandalus Crosnier, 1997 (type species Pseudopandalus curvirostris Crosnier, 1997, by original 


Pseudopandalus curvirostris Crosnier, 1997: 172; Figs 1-3. [Nouvelle-Calédonie, 22°58.9'S 167°20.2'E, 508- 

514 m] 

Stylopandalus Coutière, 1905c 

= Stylopandalus Coutière, 1905c (type species Pandalus (Stylopandalus) Richardi Coutière, 1905c, by 


Stylopandalus richardi (Coutière, 1905a) 

= Pandalus (Stylopandalus) Richardi Coutière, 1905c: 1115. [full description in Coutière, 1905a; Princesse-Alice 

stns west of Madeira, 32°18'N 23°58'W, 2000-0 m; and Canary Islands, 27°43'N 18°28'W, 

3000-0 m; see Chace, 1985] 

= Plesionika nana Murray & Hjort, 1912: 585. [Spain to Newfoundland; nomen nudum] 

= Parapandalus zur strasseni Balss, 1914a: 597. [10°08'S 97°14'E, 0-2400 m; 3°24'S 58°38'E, 0-2000 m] 

= Pandalus (Plesionica) gracilis Borradaile, 1915b: 208; nec Stimpson, 1860a. [Western Indian Ocean, 

200 fms] 

Family THAlASSoCARIDIDAe Spence Bate, 1888 

Chlorotocoides Kemp, 1925 

= Chlorotocoides Kemp, 1925 (type species Chlorotocus spinicauda De Man, 1902, by original designation 

and monotypy, gender masculine; name placed on the Official List of Generic Names in Zoology 


Chlorotocoides spinicauda (De Man, 1902) 

= Chlorotocus spinicauda De Man, 1902: 856; Plate 26, fig. 59 [Ternate] 

= Thalassocaris maldivensis Borradaile, 1915b: 208. [Maldive Islands]


Fig. 57. Thalassocaris crinita (Dana, 1852). Photo by Arthur Anker. 

Thalassocaris Stimpson, 1860a 

= Regulus Dana, 1852a (type species Regulus lucidus Dana, 1852a, designated by Kingsley, 1880; invalid 

junior homonym of Regulus Cuvier, 1880 (Aves); name placed on the Official List of Rejected 

and Invalid Generic Names in Zoology in Opinion 470 in 1957) 

= Thalassocaris Stimpson, 1860a (nomen novum for Regulus Dana, 1852a, gender masculine; name 


Thalassocaris crinita (Dana, 1852a) (Fig. 57) 

= Regulus crinitus Dana, 1852a: 27. [in mari Suluensi] 

= Thalassocaris affinis Borradaile, 1915b: 208. [Maldives; Saya de Malha; lectotype designation by 

Gopala Menon & Williamson, 1971] 

= Thalassocaris crinita var.? De Man, 1920b: 100; Plate 10, figs 23-23c. [Saleyer-anchorage and surroundings, 

including Pulu Pasi Tanette, near the North point of Saleyer-island, up to 36 m] 

Thalassocaris lucida (Dana, 1852a) 

= Regulus lucidus Dana, 1852a: 27. [in mari Pacifico, prope insulas ”Ladrone”] 

= Thalassocaris danæ Spence Bate, 1888: 683; Plate 117, fig. 1. [Fiji Islands, at the surface] 

= Thalassocaris stimpsoni Spence Bate, 1888: 684; Plate 117, fig. 2. [Japan, near Yokohama, surface; Fiji 

Islands, surface] 

Thalassocaris obscura Gopala Menon & Williamson, 1971: 36; Figs 1c, 4, 5c, 6e [Arabian Sea, 09°00'N 

75°20'E, 200-0 m] 

Superfamily PHySeToCARIDoIDeA Chace, 1940 

Family PHySeToCARIDIDAe Chace, 1940 

Physetocaris Chace, 1940 

= Physetocaris Chace, 1940 (type species Physetocaris microphthalma Chace, 1940, by monotypy, gender 

feminine) 

Physetocaris microphthalma Chace, 1940: 196, Figs 62-63. [Bermuda Oceanographic Expedition Net 798 

(32°12'N 64°36'W), 600 fms]


Superfamily CRANGoNoIDeA Haworth, 1825 

Family CRANGoNIDAe Haworth, 1825 

Aegaeon Agassiz, 1846 [in Agassiz, 1842-1846] 

= Egeon Bosc, 1813 (type species Cancer cataphractus Olivi, 1792, by monotypy, gender masculine; 

invalid junior homonym of Egon de Montfort, 1808 (Protozoa); name placed on the Official Index of 

Rejected and Invalid Generic Names in Zoology in Opinion 434 in 1956) 

= Aegaeon Agassiz, 1846 [in Agassiz, 1842-1846] (nomen novum for Egeon Bosc, 1813, gender masculine; 

see Holthuis, 1993a) 

Aegaeon boschii (Christoffersen, 1988) 

= Pontocaris boschii Christoffersen, 1988: 49; Figs 2-3. [37°S 56°W, Argentina, province of Buenos 

Aires] 

Aegaeon cataphractus (Olivi, 1792) 

= Cancer cataphractus Olivi, 1792: 50; Plate 3, fig. 1. [la Golfo e delle Lagune di Venezia] 

= E.[geon] loricatus Risso, 1827: 58; Plate 1, fig. 3. [golfe de Nice, sur les fonds rocailleux] 

Aegaeon lacazei (Gourret, 1887a) 

= Crangon Lacazei Gourret, 1887a: 1033. [région nord-ouest du golfe de Marseille, 70-80 m] 

= Ægeon Brendani Kemp, 1906b: 299. [about 50 miles W. by N. of Inishtearaght Lighthouse, Co. 

Kerry, 320-370 fms] 

= Pontocaris habereri Doflein, 1902: 620; Fig. A; Plate 1, figs 4-5. [Sagamibai] 

Aegaeon orientalis Henderson, 1893 

= Ægeon orientalis Henderson, 1893: 446; Plate 40, figs 16-17. [Gulf of Martaban] 

= Ægeon rugulosum Borradaile, 1915b: 210. [Western Indian Ocean] 

Aegaeon rathbuni De Man, 1918b 

= Aegeon Rathbuni De Man, 1918b: 304. [Pulu Kaniungan Ketjil, 11 m] 

Argis Krøyer, 1843 

= Argis Krøyer, 1843 (type species Crangon lar Owen, 1839, by monotypy, gender feminine; name 


= Nectrocrangon Brandt, 1850 (type species Crangon lar Owen, 1839, by monotypy, gender feminine) 

= Nectocrangon Brandt, 1851 (type species Crangon lar Owen, 1839, by monotypy, gender feminine; 

name placed on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 

470 in 1957) 

Argis alaskensis (Kingsley, 1883) 

= Nectocrangon alaskensis Kingsley, 1883: 128. [Marmot Island, Kodiac Archipelago, Alaska] 

Argis californiensis (Rathbun, 1902a) 

= Nectocrangon californiensis Rathbun, 1902a: 892. [off Santa Catalina Island, California, 80 fms] 

Argis crassa (Rathbun, 1899) 

= Nectocrangon crassa Rathbun, 1899: 556. [57°4'N 170°24'W, 26 fms] 

= Nectocrangon sadoensis Yokoya, 1933: 40; Fig. 21. [Sôyô-Maru stn 600, north of Is. Sado, 168 m] 

Argis dentata (Rathbun, 1902a) 

= Nectocrangon dentata Rathbun, 1902a: 892. [off Sitkalidak Island, Alaska, 69 fms] 

= Argis argilicola Krøyer, 1843: 256. [nomen nudum] 

Argis hozawai (Yokoya, 1939) 

= Nectocrangon hozawai Yokoya, 1939: 276; Fig. 9. [about 5 miles off Ozaki, 130 fms] 

= Nectocrangon lar kobjakovi Vinogradov, 1950: 221. [Sea of Japan, southern Sea of Okhotsk and 

southern Kuril Islands] 

Argis lar (Owen, 1839) 

= Crangon lar Owen, 1839: 88; Plate 28, fig. 1. [between Bristol Bay and the Pribilof islands, 25.5 fms; 

neotype selection by Squires, 1964] 

Argis levior (Rathbun, 1902a) 

= Nectocrangon levior Rathbun, 1902a: 892. [Admiralty Inlet, Puget Sound, 40 fms]


Argis ochotensis ochotensis Komai, 1997b 

= Argis ochotensis Komai, 1997b: 149; Figs 12-15. [Okhotsk Sea, Kitami, Yamato Bank, 44°39.1'N 

144°18.3'E, 815-820 m] 

Argis ochotensis kamtschatica Sokolov, 2001: 1056; Figs 3-5, 6f-h. [Ochotsk Sea, 52°45'N 154°43'E, 245 m] 

Argis ovifer (Rathbun, 1902a) 

= Nectocrangon ovifer Rathbun, 1902a: 892. [off the Trinity Islands, Alaska, 159 fms] 

Argis toyamaensis (Yokoya, 1933) 

= Nectocrangon toyamaensis Yokoya, 1933: 39; Fig. 20. [Sôyô-Maru stn 591, Toyama Bay, 311 m] 

Crangon Fabricius, 1798 

= Crangon Fabricius, 1798 (type species Cancer Crangon Linnaeus, 1758, by absolute tautonymy, 

gender feminine; junior homonym of Crangon Weber, 1795 (invalid senior synonym of Alpheus Fabricius, 

1798); name conserved under the plenary powers of the ICZN and placed on the Official List 

of Generic Names in Zoology in Opinion 334 in 1955) 

= Crago Lamarck, 1801 (type species Cancer Crangon Linnaeus, 1758, by monotypy, gender masculine; 

name placed on the Official Index of Rejected and Invalid Generic Names in Zoology in Opinion 

334 in 1955) 

= Crangonus Rafinesque, 1815 (nomen novum for Crangon Fabricius, 1798, gender masculine; name 


1955) 

= Steiracrangon Kinahan, 1862 (type species Crangon Allmanni Kinahan, 1860c, designated by Fowler, 


Crangon affinis De Haan, 1849 [in De Haan, 1833-1850]: 183. [Bay of Kobé, Japan, 34°39'N 135°14'E, 8 fms; 

neotype designation by Hayashi & J.N. Kim, 1998] 

= Crangon consobrinus De Man, 1906: 401. [Inland Sea of Japan, deep water] 

Crangon alaskensis Lockington, 1877a 

= Crangon Alaskensis Lockington, 1877a: 34. [Mutiny Bay, Alaska] 

= Crangon alaskensis elongata Rathbun, 1902a: 888. [off Santa Barbara, California, 29 fms] 

Crangon alba Holmes, 1900: 174. [Monterey] 

Crangon allmanni Kinahan, 1860c 

= Crangon Allmanni Kinahan, 1860c: 28; unnumbered text Figs [Zonam corallinam Maris Hibernici 

juxta, ’Bray,’ comitat: ’Wicklow’] 

Crangon amurensis Bražnikov, 1907 

= Crangon septemspinosa f. amurensis Bražnikov, 1907: 68. [Amuriskiy Liman] 

= Crangon septemspinosa f. anivensis Bražnikov, 1907: 68. [Aniva Bay] 

Crangon capensis Stimpson, 1860a: 24. [in sinu ”Simon’s Bay”, promont. Bonae Spei, prof. 12 org] 

Crangon cassiope De Man, 1906: 402. [Inland Sea of Japan, deep water] 

Crangon crangon (Linnaeus, 1758) 

= Cancer Crangon Linnaeus, 1758: 632. [Mare Balthico] 

= Crangon vulgaris Fabricius, 1798: 410. [mari balthico] 

= Crangon Rubro punctatus Risso, 1816: 83. [environs de Nice, dans les endroits sabloneux] 

= Cancer schillinus Nardo, 1847: 7. [Trovato nel golfo in estate] 

= Crangon maculosus Rathke, 1837: 366. [Cap Porthenion und bei Sevastopol] 

= Crangon maculosus forma typica Czerniavsky, 1884: 71. [Sevastopol; Odessa; Theodosia] 

= Crangon maculosus forma brevirostris Czerniavsky, 1884: 72. [sinus Sevatopol et Odessa] 

= Crangon maculosus var. suchumica Czerniavsky, 1884: 72. [sin. Suchum] 

= Steiracrangon orientalis var. longicauda forma intermedia Czerniavsky, 1884: 74. [sinus Sevastopol; 

Orianda] 

= Steiracrangon orientalis var. brevicauda Czerniavsky, 1884: 75. [sinus Sevastopol; Orianda] 

= Crangon crangon typicus Doflein, 1900: 325. [Type locality not indicated] 

= Crangon crangon f. mediterranea Bražnikov, 1907: 70. [Naples] 

= Crangon crangon f. typica Bražnikov, 1907: 70. [White Sea; Norway] 

Crangon dalli Rathbun, 1902a: 889. [Bering Sea, off Cape Seniavin, Alaska, 30 fms]


Crangon franciscorum franciscorum Stimpson, 1856 

= Crangon Franciscorum Stimpson, 1856: 89. [sandy coves around the bay (San Francisco)] 

Crangon franciscorum angustimana Rathbun, 1902a: 889. [off Chuckanuts Island, Bellingham Bay, Washington, 

11 fms] 

Crangon hakodatei Rathbun, 1902b: 42; Fig. 15. [Hakodate, Hokkaido] 

Crangon handi Kuris & Carlton, 1977: 543; Figs 1, 2a, d, g. [Horseshoe Cove, Bodega Head, Sonoma 

County, 38°18'30''N 123°4'W] 

Crangon holmesi Rathbun, 1902a: 888. [Off Wilmington, California, 27 fms] 

Crangon lockingtonii Holmes, 1904 

= Crangon Lockingtonii Holmes, 1904: 312; Plate 35, figs 14-16. [Magdalen Bay, Lower California] 

Crangon nigricauda Stimpson, 1856: 89. [California, deeper water] 

Crangon nigromaculata Lockington, 1877a: 34. [San Diego, 6 fms] 

Crangon propinquus Stimpson, 1860a: 25. [Lake Saroma, Hokkaido; neotype selection by Hayashi & J.N. 

Kim, 1999] 

= Steiracrangon orientalis var. longicauda forma pacifica Czerniavsky, 1884: 74. [Sachalin (ocean. Pacific); 

colonia Dui (Itere Amurensi)] 

= Crangon vulgaris var. Shidlovskii Ostroumov, 1896: 81. [Vladivostok] 

Crangon septemspinosa Say, 1818 

= C.[rangon] septemspinosus Say, 1818: 246. [bay shores and inlets of the sea] 

Crangon uritai Hayashi & J.N. Kim, 1999: 86; Figs 13-16. [Dadaepo, Pusan, Krea, 1 m] 

Lissocrangon Kuris & Carlton, 1977 

= Lissocrangon Kuris & Carlton, 1977 (type species Crangon stylirostris Holmes, 1900, by original 


Lissocrangon stylirostris (Holmes, 1900) 

= Crangon stylirostris Holmes, 1900: 174. [Trinidad, Humboldt County, California] 

Lissosabinea Christoffersen, 1988 

= Lissosabinea Christoffersen, 1988 (type species Sabinea tridentata Pequegnat, 1970, by original designation, 


Lissosabinea armata Komai, 2006a: 44; Figs 7-9. [New Caledonia, 24°44.24'S 170°08.01'E, 770-830 m] 

Lissosabinea arthuri Taylor & Collins, 2009: 177; Figs 3-4. [Australia, Vic., 38°09.80'S 149°41.71'E - 

38°10.11'S 149°41.01'E, 260-265 m] 

Lissosabinea beresfordi Taylor & Collins, 2009: 176; Figs 1-2, 8A-B. [Australia, Southern Ocean, south of 

Tasmania, 43°59.5'S 147°32.76'E - 43°59.7'S 147°33.80'E, 370-410 m] 

Lissosabinea ecarina Komai, 2006a: 49; Figs 10-12. [Indonesia, Kai Islands, 05°30'N 132°52'E, 336-346 m] 

Lissosabinea indica (De Man, 1918b) 

= Sabinea indica De Man, 1918b: 304. [7°0'S 120°34'.5E, North of Tanah Djampeah Island, 400 m] 

Lissosabinea lynseyae Taylor & Collins, 2009: 186; Figs 15-6, 8C. [Australia, WA, off Bunbury, 33°00.5'S 

114°59.26'E - 33°00.11'S 114°34.50'E, 421-414 m] 

Lissosabinea tridentata (Pequegnat, 1970) 

= Sabinea tridentata Pequegnat, 1970: 115; Figs 4-16, 4-17. [Southeastern Gulf of Mexico, 24°58'N 

84°17'W, 214 fms] 

Lissosabinea unispinosa Komai, 2006a: 53; Figs 13-16. [New Caledonia, 18°54.8'S 163°22.2'E, 465 m] 

Mesocrangon Zarenkov, 1965 

= Mesocrangon Zarenkov, 1964 (unavailable name under Art. 13.3) 

= Mesocrangon Zarenkov, 1965 (type species Crangon intermedius Stimpson, 1860a, by original designation 


Mesocrangon intermedia (Stimpson, 1860a) 

= Crangon intermedius Stimpson, 1860a: 25. [mari Beringiano prope promontorium ”Chepoonski”, 

ad prof. 40 org]


= Crangon tenuifrons Kingsley, 1883: 128; Plate 1, fig. 10. [Marmot Island, Kodiac Archipelago, Alaska] 

= Sclerocrangon intermedius var. bidentata Balss, 1914b: 65. [Sagamibai] 

Mesocrangon munitella (Walker, 1898) 

= Crangon munitellus Walker, 1898: 275; Plate 16, fig. 1. [Puget Sound, pacific coast of North America] 

Mesocrangon volki (Birstein & Vinogradov, 1953) 

= Sclerocrangon volki Birstein & Vinogradov, 1953: 217; Fig. 1. [near Bering Island, Bering Sea, 93- 

112 m] 

Metacrangon Zarenkov, 1965 

= Metacrangon Zarenkov, 1964 (unavailable name under Art. 13.3) 

= Metacrangon Zarenkov, 1965 (type species Crangon variabilis Rathbun, 1902a, by original designation, 


Metacrangon acclivis (Rathbun, 1902a) 

= Crangon acclivis Rathbun, 1902a: 890. [off Santa Cruz Island, California, 266 fms] 

Metacrangon agassizii (Smith, 1882) 

= Cheraphilus Agassizii Smith, 1882: 32; Plate 7, figs 4-5a. [Blake stns 317, 31°57'0''N 78°18'35''W, 333 


74°48'0''W, 263 fms; off Block Island, 39°46'N 71°10'W, 500 fms] 

Metacrangon australis Komai & Taylor, 2010: 46; Figs 1-3. [Australia, Tasmania, southwestern Pacific, 

west of Macquarie Island, 54°42.42'S 158°45.12'E – 54°41.36'S 158°43.12'E, 700-900 m] 

Metacrangon bahamondei Retamal & Gorny, 2003: 85; Fig. 1. [55°44'00''S 66°14'05''W, sur de la Isla Nueva, 

Tierra del Fuego, 746 m] 

Metacrangon bellmarleyi (Stebbing, 1914b) 

= Sclerocrangon bellmarleyi Stebbing, 1914b: 29; Plate 10. [Cape Natal N by E 24 miles, Natal, 440 fms] 

Metacrangon cornuta Komai & Komatsu, 2009: 515; Figs 2-4, 19G. [off Kushi-ro, eastern Hokkaido, 

42°30.3'N 144°50.5'E to 42°30.6'N 144°2.2'E, 1535-1543 m] 

Metacrangon crosnieri Komai, 1997c: 662; Figs 1c, 5-6. [Madagascar, off Cape Ambre, 13°02'S 48°02'E, 

1000-1525 m] 

Metacrangon haona Komai & Ahyong, 2011: 78; Figs 1-3. [Pukaki Rise, Campbell Plateau, 49°21.50'S 

171°53.00'E, 353 m] 

Metacrangon hikurangi Komai & Ahyong, 2011: 83; Figs 4-6. [Hikurangi Margin, east coast of North Island, 

New Zealand, 40°02.22-02.14'S 177°48.06-47.97'E, 1171-1172 m] 

Metacrangon holthuisi Komai, 2010a: 370; Figs 1-2. [west of Sunosaki, Tateyama, Sagami Sea, 34°56.46'N 

139°33.12'E, 1039-1300 m] 

Metacrangon jacqueti (A. Milne-Edwards, 1881a) 

= Pontophilus Jacqueti A. Milne-Edwards, 1881a: 933. [golfe de Gascogne, 44°48'30''N 7°0'30''W, 

5100 m] 

Metacrangon knoxi (Yaldwyn, 1960) 

= Sclerocrangon knoxi Yaldwyn, 1960: 35; Fig. 7. [Chatham Rise, 43°40'S 179°28'E, 220 fms] 

Metacrangon laevis (Yokoya, 1933) 

= Crangon laevis Yokoya, 1933: 36; Fig. 16. [Sôyô-Maru stn 638, west of Tubaki-zaki, Aomori-ken, 

73 m] 

Metacrangon longirostris (Yokoya, 1933) 

= Crangon longirostris Yokoya, 1933: 36; Fig. 17. [Sôyô-Maru stn 381, east of Owase, Mie-ken, 329 m] 

Metacrangon miyakei J.N. Kim, 2005: 246; Figs 5-6. [Sea of Japan, off Taiza, Tango Peninsula, 110 m] 

Metacrangon monodon (Birstein & Vinogradov, 1951) 

= Sclerocrangon monodon Birstein & Vinogradov, 1951: 357; Figs 1-3. [4th Kuril Straits, Kuril Islands, 

NW Pacific, 630 m] 

Metacrangon munita (Dana, 1852a) 

= Crangon munitus Dana, 1852a: 20. [in freto Pugettensi] 

Metacrangon nipponensis (Yokoya, 1933) 

= Crangon nipponensis Yokoya, 1933: 38; Fig. 19. [Sôyô-Maru stn 22, east of Siwoya-zaki, 539 m]


Metacrangon ochotensis (Kobjakova, 1955) 

= Sclerocrangon ochotensis Kobjakova, 1955: 235; Fig. 1. [Ochotsk Sea, Kuril Island area, off Ekaterina 

Straits, 2850 m] 

Metacrangon procax (Faxon, 1893) 

= Sclerocrangon procax Faxon, 1893: 199. [Albatross stns 3380 (04°03'00''N 81°31'00''W, 899 fms); 3418 

(16°33'00''N 99°52'30''W, 660 fms); 3435 (26°48'00''N 110°45'20''W, 859 fms); 3436 (27°03'40''N 

110°53'40''W, 905 fms] 

= Crago lomae Schmitt, 1921: 100; Plate 12, figs 3-4. [Off Point Loma, California, 525-541 fms] 

Metacrangon proxima J.N. Kim, 2005: 242; Figs 1-3. [Pacific coast of central Japan, Amadai-ba, Sagami Bay, 

280 m] 

Metacrangon poorei Komai & Taylor, 2010: 49; Figs 4-5. [Australia, off southeastern Victoria, 39°53.76'S 

149°03.39'E, 1608 m] 

Metacrangon rau Komai & Ahyong, 2011: 91; Figs 7-10. [west of Northland, North Island, 35°19.99'S 

172°19.99'E, 1029-1074 m] 

Metacrangon richardsoni (Yaldwyn, 1960) 

= Sclerocrangon richardsoni Yaldwyn, 1960: 39; Fig. 8. [41°42'30''S 175°9'E, circa 550 fms] 

Metacrangon robusta (Kobjakova, 1935) 

= Sclerocrangon robustus Kobjakova, 1935: 85; Figs 1-2. [Japanischen Meer von der Bucht Peters des 

Großen nach Norden bis zu 50° n. Br., 70-1380 m; Ochotskisches Meer im nordwestlichen Teil, in 

der Umgegend von der St.-Jonas-Insel, 50-515 m] 

Metacrangon similis Komai, 1997c: 675; Figs 1h, 11-12. [Japan, off Owase, Kumano-nada, 34°05.8'N 

136°35.9'E, 369-412 m] 

Metacrangon sinensis Fujino & Miyake, 1970a: 269; Figs 10-15. [37°01.5'N 123°57.9'E, 74 m] 

Metacrangon spinidorsalis Komai & Taylor, 2010: 52; Figs 6-10. [Australia, Western Australia, off Point 

Hillier, 35°22.54'S 117°12.25'E – 35°22.54'S 117°12.25'E, 539 m] 

Metacrangon spinirostris (Rathbun, 1902a) 

= Crangon spinirostris Rathbun, 1902a: 891. [North of Unalaska, 399 fms, Albatross stn 3329 

(53°56'50''N 167°08'15''W)] 

Metacrangon spinosissima (Rathbun, 1902a) 

= Crangon spinosissima Rathbun, 1902a: 891. [Off Point Arena, California, 51 fms] 

Metacrangon teina Komai & Ahyong, 2011: 95; Figs 11-15. [Challenger Plateau, 39°32.61'S 169°42.87'E, 

636-634 m] 

Metacrangon trigonorostris Yokoya, 1933 

= Crangon trigonorostris Yokoya, 1933: 37; Fig. 18. [Sôyô-Maru stn 73, south of Siwoya-zaki, 390 m] 

Metacrangon variabilis variabilis (Rathbun, 1902a) 

= Crangon variabilis Rathbun, 1902a: 890. [Off North Head, Akutan Island, Alaska, 72 fms] 

Metacrangon variabilis asiaticus (Kobjakova, 1955) 

= Sclerocrangon variabilis asiaticus Kobjakova, 1955: 236; Fig. 2. [SE of Shikotan Island, Kuril Islands, 

136-414 m] 

Neocrangon Zarenkov, 1965 

= Neocrangon Zarenkov, 1964 (unavailable name under Art. 13.3) 

= Neocrangon Zarenkov, 1965 (type species Crangon communis Rathbun, 1899, by original designation, 


Neocrangon abyssorum (Rathbun, 1902a) 

= Crangon abyssorum Rathbun, 1902a: 890. [Bering Sea, southwest of Pribilof Islands, 1771 fms] 

Neocrangon communis (Rathbun, 1899) 

= Crangon communis Rathbun, 1899: 556. [57°4'20''N 170°52'30''W, 51 fms] 

Neocrangon geniculata (Yokoya, 1933) 

= Crangon geniculata Yokoya, 1933: 35; Fig. 15. [Sôyô-Maru stn 213, Kii Strait, 353 m] 

Neocrangon joloensis (De Man, 1929) 

= Crangon joloensis De Man, 1929: 131; Figs 9-9b. [off the Jolo Islands, 16 fms]


Neocrangon resima (Rathbun, 1902a) 

= Crangon resima Rathbun, 1902a: 889. [Off San Diego, California, 124 fms] 

= Crago zacae Chace, 1937b: 136; Fig. 9. [east of Cedros Island, 28°13'N 115°07'W, 44 fms] 

Neocrangon sagamiensis (Balss, 1913b) 

= Crangon (Crangon) sagamiense Balss, 1913b: 237. [Fukuura, Sagamibai] 

= Neocrangon orientalis Han & Li, 2009: 65; Figs 1-2. [East China Sea, 30°31.0'N 127°56.5'E, 365-395 m] 

Notocrangon Coutière, 1900d 

= Notocrangon Coutière, 1900d (type species Crangon antarcticus Pfeffer, 1887, by monotypy, gender 


Notocrangon antarcticus (Pfeffer, 1887) (Fig. 58) 

= Crangon antarcticus Pfeffer, 1887: 45; Plate 1, Figs 1-21. [Süd-Georgien] 

= Crangon (Notocrangon) antarcticus var. gracilis Borradaile, 1916: 89. [Terra Nova Expedition stns 294 

(74°25'S 166°47'W, 0-1750 m); 314 (5 miles N of Inaccesible Island, McMurdo Sound, 406-411 m); 316 

(off Glacier Tongue, about 8 miles N of Hut Point, McMurdo Sound, 348-457 m); 338 (77°13'S 

164°18'E, 379 m); 339 (77°5'S 164°17'E, 256 m); 348 (off Barne Glacier, McMurdo Sound, 366 m); 355 

(77°46'S 166°8'E, 547 m)] 

Fig. 58. Notocrangon antarcticus (Pfeffer, 1887). Photo by Cedric d'Udekem d'Acoz. 

Paracrangon Dana, 1852a 

= Paracrangon Dana, 1852a (Paracrangon echinatus Dana, 1852a, by monotypy, gender feminine; 

name placed on the Official List of Generic Names in Zoology in 470 in 1957) 

Paracrangon abei Kubo, 1937b: 3; Figs 2-3. [Kumano-nada, off Nagasima, Mie Prefecture, about 170 fms] 

Paracrangon areolata Faxon, 1893: 200. [Albatross stns 3424 (21°15'00''N 106°23'00''E), 676 fms; 3425 

(21°19'00''N 106°24'00''E), 680 fms] 

Paracrangon australis Hanamura, Wadley & Taylor, 1999: 311; Figs 1-3. [Tasmania, approx. 84 km SSE of 

South East Cape, 44°16'S 147°20'E, 987 m] 

Paracrangon echinata Dana, 1852a 

= Paracrangon echinatus Dana, 1852a: 20. [in freto Pugettensi] 

Paracrangon furcata Kubo, 1937b: 7; Figs 4-5. [Kumano-nada, off Nagasima, Mie Prefecture, about 170 

fms] 

Paracrangon okutanii Ohé & Takeda, 1986: 76; Figs 2-3. [35°04'N 139°28.4'E, 770-930 m] 

Paracrangon ostlingos Komai & J.N. Kim, 2004: 531; Figs 1-13. [Pacific coast of Japan, off Taito-saki, Boso 

Peninsula, 35°09.4'N 140°49.0'E, 311-323 m]


Parapontocaris Alcock, 1901 

= Parapontocaris Alcock, 1901 (type species Crangon bengalensis Wood-Mason in Wood-Mason & 

Alcock, 1891c, designated by Holthuis, 1955b, gender feminine) 

Parapontocaris andamanensis (Wood-Mason in Wood-Mason & Alcock, 1891c) 

= Crangon andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c: 360. [Andaman Sea, 

11°31'40''N 92°46'40''E, 188-220 fms] 

Parapontocaris aspera Chace, 1984: 30; Figs 9-11. [Albay Gulf, east of southern Luzon, Philippines, 

13°10'21''N 123°59'54'', 368 m] 

Parapontocaris bengalensis (Wood-Mason in Wood-Mason & Alcock, 1891c) 

= Crangon bengalensis Wood-Mason in Wood-Mason & Alcock, 1891c: 360. [Bay of Bengal, 15°56'50''N 

81°30.5'E, 240-276 fms] 

Parapontocaris caribbaea (Boone, 1927) 

= Aegeon caribbaeus Boone, 1927: 125; Fig. 28. [north of Glover Reef, 484 fms] 

Parapontocaris levigata Chace, 1984: 34; Figs 12-14. [Balayan Bay, southern Luzon, Philippines, 13°51'30''N 

120°50'30''E, 291 m] 

Parapontocaris vicina (Dardeau & Heard, 1983) 

= Pontocaris vicina Dardeau & Heard, 1983: 10; Figs 2g, 4-7. [Straits of Florida, 23°35'N 80°22'W, 457 m] 

Parapontophilus Christoffersen, 1988 

= Parapontophilus Christoffersen, 1988 (type species Pontophilus gracilis Smith, 1882, by original designation, 


Parapontophilus abyssi (Smith, 1884) 

= Pontophilus abyssi Smith, 1884: 363. [Albatross stns 7023/7025, 37°56'20''N 70°57'30''W, 1917 fms; 

7024, 37°40'30''N 70°37'30''W, 2221 fms] 

= Pontophilus gracilis Spence Bate, 1888: 487 (partim); Plate 87; nec Smith, 1882. [Challenger stn 133, 

35°41'S 20°55'W, near Tristan da Cunha, 1900 fms only] 

= Pontophilus challengeri Ortmann, 1893: 49. [nomen novum for Pontophilus gracilis Spence Bate, 

1888] 

= Pontophilus batei Faxon, 1893: 200. [nomen novum for Pontophilus gracilis Spence Bate, 1888] 

Parapontophilis caledonicus Komai, 2008: 313; Figs 24-25. [New Caledonia, 20°35.9'S 164°58.3'E, 400- 

500 m] 

Parapontophilus cornutus Komai, 2008: 292; Figs 10-12. [Austral Islands, Tubuai Island, 23°21.3'S 

149°33.9'W, 500-200 m] 

Parapontophilus cyrton Komai, 2008: 295; Fig. 13. [New Caledonia, 23°52'S 167°58'E, 1220-1225 m] 

Parapontophilus demani (Chace, 1984) 

= Pontophilus demani Chace, 1984: 48. [Selat Roti, between Timor and Pulau Roti, Lesser Sunda Islands, 

Indonesia, 10°39'S 123°40'E, 520 m] 

Parapontophilus difficilis Komai, 2008: 297; Fig. 6C, 14-16. [Vanuatu, 16°00.73'S 166°39.94'E, 1014-1050 m] 

Parapontophilus geminus Komai, 2008: 302; Figs 17, 20F. [Japan, Hyuga Basin, 32°2.58'N 132°30.54'E, 1644- 

1657 m] 

Parapontophilus gracilis (Smith, 1882) 

= Pontophilus gracilis Smith, 1882: 36; Plate 7, Figs 2-3a. [Blake stn 315, 32°18'20''N 78°43'W, 225 fms; 

Albatross stn 1029, off Martha’s Vineyard, 458 fms] 

Parapontophilus junceus (Spence Bate, 1888) 

= Pontophilus junceus Spence Bate, 1888: 491; Plate 88, Figs 2-4. [Challenger stn 200, 6°47'N 122°28'E, 

between the Philippines and Borneo, 250 fms] 

= Pontophilus occidentalis var. indica De Man, 1918a: 161. [Siboga Expedition stns 45, 7°24'S 118°15'.2E, 

Flores Sea, 794 m; 88, 0°34'.6N 119°8'.5E, northern part of the Strait of Makassar, 1301 m; 178, 2°40'S 

128°37'.5E, Ceram Sea, 835 m; 211, 5°40'.7S 120°45'.5E, E of Saleyer Island, 1158 m; 300, 10°48'.6S 

123°23'.1E, S of Rotti island, 918 m; 316, 7°19'.4S 116°49'.5E, Bali Sea, 538 m] 

Parapontophilus juxta Komai, 2008: 316; Figs 26-27. [Western Indian Ocean, off Réunion, 21°03.6'S 

55°09.8'E, 412-460 m]


Parapontophilus longirostris Komai, 2008: 305; Figs 18-19, 21A. [Austral Islands, Tubuai island, 23°21.3'S 

149°33.9'W] 

Parapontophilus modumanuensis (Rathbun, 1906) 

= Pontophilus modumanuensis Rathbun, 1906: 910; Fig. 63. [vicinity of Modu Manu, Hawaii Islands, 

293-800 fms] 

Parapontophilus occidentalis (Faxon, 1893) 

= Pontophilus occidentalis Faxon, 1893: 200. [02°34.0'N 92°06.0'W, 2448 m; 04°56.0'N 80°52.30'W, 3243 

m; 05°43.0'N 85°50.0'W, 1760 m; 06°10.0'N 83°06.0'W, 2648 m; 06°21.0'N 80°41.0'W, 3281 m; 

07°05.30'N 79°40.0'W, 2286 m (all Gulf of Panama); off Ecuador, 01°07.0'N 80°21.0'W, 2831 m; off 

Costa Rica, 10°14.0'N 96°28.0'W, 4018 m; off Guatemala, 14°46.0'N 98°40.0'W, 3190 m] 

Parapontophilus profundus (Spence Bate, 1888) 

= Pontophilus profundus Spence Bate, 1888: 490; Plate 88, fig. 1-1z. [Challenger stn 165, 34°50'S 155°28'E, 

off Sydney, 2600 fms] 

Parapontophilus psyllus Komai, 2008: 319; Figs 28-29. [Japan, off Nago, Okinawa Island, Ryukyu Islands, 

26°32.90'N 127°43.94'E, 396-407 m] 

Parapontophilus sibogae Komai, 2008: 322; Figs 30-31. [Indonesia, Pulu Kaniungan Ketjil, 11 m] 

Parapontophilus stenorhinus Komai, 2008: 324; Figs 32-33. [Tonga, 21°02'S 175°19'W, 555-581 m] 

Parapontophilus talismani (Crosnier & Forest, 1973) 

= Pontophilus talismani Crosnier & Forest, 1973: 245; Fig. 80a-d. [16°38'N 18°24'W, 3200 m] 

Philocheras Stebbing, 1900 

= Mesapus Rafinesque, 1814 (type species Mesapus fasciatus Rafinesque, 1814 (invalid senior subjective 

synonym of Crangon Fasciatus Risso, 1816), gender masculine; name of genus and its type species 

suppressed for the purposes of the Principle of Priority, but not those of the Principle of Homonymy; 

placed on the Official Index of Rejected and Invalid Generic (resp. Specific) Names in 


= Philocheras Stebbing, 1900 (type species Crangon nanus Krøyer, 1843, selected by Holthuis, 1955b, 


Philocheras acutirostratus (Yaldwyn, 1960) 

= Pontophilus acutirostratus Yaldwyn, 1960: 41; Fig. 9. [Chatham Rise, 43°42'S 179°55'E, 280 fms] 

Philocheras aglyptus (Crosnier, 1971) 

= Pontophilus aglyptus Crosnier, 1971: 581; Figs 4a-b, 5. [Congo, baie de Point-Noire, senne de plage] 

Philocheras angustirostris (De Man, 1918a) 

= Pontophilus angustirostris De Man, 1918a: 163. [Siboga Expedition stns 7, 7°55'.5S 114°26'E, reef of 

Batjulmani (Java); 50, Bay of Bajo, W coast of Flores, up to 40 m; 164, 1°42'.5S 130°47'.5E, between 

Misool and New Guinea, 32 m; 273, anchorage of Pulu Jedan, E coast of Aru-islands, 13 m] 

Philocheras anthonyi Taylor, 2010: 159; Figs 1-2. [Australia, Western Australia, 17°35.98'S 118°5908'E to 

17°38.56'S 119°01.26'E, 222 m] 

Philocheras australis (Thomson, 1879) 

= Crangon australis Thomson, 1879: 231; Plate 10, Fig. A1a-c. [Cook Straits, Dunedin; Stewart Island] 

Philocheras bidens (Holthuis, 1951a) 

= Pontophilus bidens Holthuis, 1951a: 165; Fig. 33. [off Portuguese Guinea, 11°54'N 17°14'W, 55-80 m] 

Philocheras bidentatus (De Haan, 1844 [in De Haan, 1833-1850]) 

= Crangon bidentatus De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 14 (1844); 183 (1849). [Japan] 

Philocheras bispinosus bispinosus (Hailstone, 1835a) 

= Pontóphilus bispinòsus Hailstone, 1835a: 271; Fig. 30. [off Hastings] 

= Pontóphilus bispinòsus Westwood, 1835c: 273. [off Hastings] 

= Crangon nanus Krøyer, 1843: 231; Plate 4, Figs 15-28. [i det sydligste Kattegat (udfor Hornbet, paa 

en halv Snees Favnes Dybbe og paa Leerbund)] 

= Pontophilus mbizi Holthuis, 1952a: 64; Fig. 16. [15°57'S 11°40'E, 5 M.SW. Pointa Albina] 

Philocheras bispinosus neglectus (G.O. Sars, 1883)


Fig. 59. Philocheras coralliophilus Komai & Kim, 2010. Photo by Arthur Anker. 

= Ceraphilus neglectus G.O. Sars, 1883: 45; Plate 1, fig. 7. [ad oras meridionales et occidentalis Norvegiæ 

in prof 2-6 orgyarum] 

Philocheras breviflagella Komai, 2001b: 419; Figs 1-3. [Ala Moana, O’ahu, Hawai’i, subtidal] 

Philocheras brucei Komai, 2004a: 665; Figs 1-4. [Cairns, northeast Queensland, 16°55'S 145°46'E, 10 m] 

Philocheras candidus (Kemp, 1916b) 

= Pontophilus candidus Kemp, 1916b: 365; Fig. 3; Plate 8, fig. 3. [Port Blair, Andamans] 

Philocheras carinicauda (Stimpson, 1860a) 

= Crangon carinicauda Stimpson, 1860a: 25. [portu Sinensi ”Hong Kong”] 

Philocheras chiltoni (Kemp, 1911) 

= Pontophilus chiltoni Kemp, 1911: 5; Plate 2, Figs 6-10. [New Zealand] 

Philocheras coralliophilus Komai & J.N. Kim, 2010: 12; Figs 1-4. [Uehara Beach, Iriomote Island, Yaeyama 

Islands, Ryukyus, subtidal] (Fig. 59) 

Philocheras dimorphus (Fujino & Miyake, 1971) 

= Pontophilus dimorphus Fujino & Miyake, 1971: 27; Figs 1-2. [Sagami Bay, Amadai-ba, Maruyamadashi, 

80 m] 

Philocheras echinulatus (M. Sars, 1862) 

= Crangon echinulatus M. Sars, 1862: 186. [Norges Vestkyst, Manger ved Bergen] 

? = Cheraphilus Pattersonii Kinahan, 1860d: 130. [at the Gobbins, full description in Kinahan, 1862] 

= Crangon serratus Norman, 1862: 151. [off the Shetland Isles] 

Philocheras fasciatus (Risso, 1816) 

= Mesapus fasciatus Rafinesque, 1814: 23. [Type locality not indicated; name suppressed for the purpose 

of the Principle of Priority, but not those of the Principle of Homonymy; placed on the Official 

Index of Rejected and Invalid Specific Names in Zoology in Opinion 522 in 1958] 

= Crangon Fasciatus Risso, 1816: 82; Plate 3, fig. 5. [environs de Nice, sure les bas fond sabloneux] 

Philocheras flindersi (Fulton & Grant, 1902) 

= Pontophilus flindersi Fulton & Grant, 1902: 67; Plate 10, fig. 3. [off Shoreham, Western Port, about 

4 fms] 

Philocheras fujinoi J.N. Kim & Hayashi, 2000: 694; Figs 4-5. [off Shantou, northern South China Sea, 80 m] 

Philocheras gaillardi (Crosnier, 1971) 

= Pontophilus gaillardi Crosnier, 1971: 588; Figs 7-8. [Congo, au large de Pointe-Noire] 

Philocheras gemmaceus J.N. Kim & Hayashi, 2000: 688; Figs 1-2, 3a-e. [off Kaminoseki, Seto Inland Sea, 

Japan, 50 m] 

Philocheras gorei (Dardeau, 1980) 

= Pontophilus gorei Dardeau, 1980: 563; Figs 1-4. [26°24'N 83°22'W, 55 m] 

Philocheras hamiltoni (Yaldwyn, 1971)


= Pontophilus hamiltoni Yaldwyn, 1971: 92. [off Portobello Marine Biological Station wharf, Otago 

Harbour, 2-4 fms] 

Philocheras hendersoni (Kemp, 1915) 

= Pontophilus hendersoni Kemp, 1915: 261; Fig. 25; Plate 13, fig. 8. [outer channel of Chilka Lake, 

1-2 ft] 

Philocheras incisus (Kemp, 1916b) 

= Pontophilus incisus Kemp, 1916b: 357; Fig. 1; Plate 8, fig. 1. [Andamans, 20 fms; Port Blair, Andamans, 

2-12 fms] 

Philocheras intermedius (Spence Bate, 1863) 

= Crangon intermedius Spence Bate, 1863: 503; Plate 41, figs 6-6l; nec Stimpson, 1860a. [St. Vincent 

Gulf, 4.5 fms] 

= Crangon batei Kingsley, 1883: 129. [nomen novum for Crangon intermedius Spence Bate, 1863] 

Philocheras japonicus (Doflein, 1902) 

= Pontophilus japonicus Doflein, 1902: 621; Fig. B; Plate 3, fig. 6. [Süden der Sagamibai] 

Philocheras kempii (De Man, 1918a) 

= Pontophilus Kempii De Man, 1918a: 165. [very near Siboga Expedition stn 65 (7°0'S 120°34'.5E), between 

the islands of Saleyer and Tanah Djampeah, 300-400 m] 

Philocheras lapillus Wicksten, 1989b: 80; Figs 2-4. [Off Gardner Bay, Hood Island (Isla Española), Galápagos 

(Archipelago de Colón), 1°22'S 90°40'W, 46-65 m] 

Philocheras lowisi (Kemp, 1916b) 

= Pontophilus lowisi Kemp, 1916b: 361; Fig. 2; Plate 8, fig. 2. [Port Blair, Andamans, 3-12 fms] 

Philocheras magnioculus Komai & Chan, 2007: 159; Figs 1-3. [between Libaong and Pamilacan, 9°31.3'N 

123°51.4'E, 145-163 m] 

Philocheras megalocheir Stebbing, 1915: 71; Plate 15. [near East London, Cove Rock, NE 2 miles, 25 fms; 

33°13'S 27°39'E, 37 fms] 

Philocheras modestus (De Man, 1918a) 

= Pontophilus modestus De Man, 1918a: 162. [Siboga Expedition stn 253, 5°48'.2S 132°13'E, W of Keiislands, 

304 m] 

Philocheras monacanthus (Holthuis, 1961c) 

= Pontophilus monacanthus Holthuis, 1961c: 26; Fig. 9. [harbour of Mersin, south-east coast of Turkey, 

0.5-1.5 m] 

Philocheras nikiforovi (Burukovsky, 1990) 

= Pontophilus nikiforovi Burukovsky, 1990: 209; Fig. 5B. [25°03'S 97°27'W, 271 m] 

Philocheras obliquus (Fulton & Grant, 1902) 

= Pontophilus obliquus Fulton & Grant, 1902: 63; Plate 10, fig. 1. [off Shoreham, Western Port, about 5 

fms] 

Philocheras opici (Crosnier, 1971) 

= Pontophilus opici Crosnier, 1971: 585; Figs 4c-d, 6. [Ile Annobon, 1°26'30''S 5°35'30''E, 50 m] 

Philocheras parasculptus Burukovsky, 1991b: 40; Figs 2.4-2.9. [33°16'S 43°53'E, 415-460 m] 

Philocheras parvirostris (Kemp, 1916b) 

= Pontophilus parvirostris Kemp, 1916b: 372; Fig. 6; Plate 8, fig. 6. [Kilakarai, Ramnad district, S. India] 

Philocheras pilosoides (Stephensen, 1927) 

= Pontophilus pilosoides Stephensen, 1927: 299; Fig. 1. [Auckland Island, Carnley Harbour, 45 fms; 

Campbell Island, Perseverance Harbour, 10-20 fms] 

Philocheras pilosus (Kemp, 1916b) 

= Pontophilus pilosus Kemp, 1916b: 367; Fig. 4; Plate 8, fig. 4. [Kilakarai, Ramnad district, S. India] 

Philocheras planoculminus Bruce, 1994c: 753; Fig. 3. [Flat Top Bank, Timor Sea, 12°35'S 129°35'E, about 

30 m] 

Philocheras plebs (Kemp, 1916b) 

= Pontophilus plebs Kemp, 1916b: 370; Fig. 5; Plate 8, fig. 5. [Port Blair, Andamans, 2 fms] 

Philocheras prionolepis (Holthuis, 1952a) 

= Pontophilus prionolepis Holthuis, 1952a: 61: Fig. 15. [6°06'S 10°36'E, 35 M. W. Moita Seca] 

Philocheras quadrispinosus (Yaldwyn, 1971)


= Pontophilus quadrispinosus Yaldwyn, 1971: 93. [off Cape Reinga, North Auckland, approx. 34°21.5'S 

172°49'E, 58 m] 

Philocheras sabsechota (Kemp, 1911) 

= Pontophilus sabsechota Kemp, 1911: 6; Plate 2, Figs 11-14. [S. Sentinel Island, Andamans] 

Philocheras sculptus (Bell, 1847 [in Bell, 1844-1853]) 

= Crangon sculptus Bell, 1847 [in Bell, 1844-1853]: 263, unnumbered text figure. [Weymouth] 

Philocheras triangulus Komai, 2006b: 31; Figs 1-4. [West Alligator Head, Kakadu National Park, Northern 

Territory, 12°9.725'S 132°13.959'E, 7.3 m] 

Philocheras trispinosus (Hailstone in Hailstone & Westwood, 1835) 

= Pontóphilus trispinòsus Hailstone in Hailstone & Westwood, 1835: 261; Fig. 25. [Hastings; see 

Holthuis, 1976b for discussion of authorship] 

Philocheras vanderbilti (Boone, 1935) 

= Pontophilus vanderbilti Boone, 1935: 200; Plate 55. [Flores Strait, near Larantuka village, Flores Island, 

140 fms] 

Philocheras vestigialis (Fujino & Miyake, 1971) 

= Pontophilus vestigialis Fujino & Miyake, 1971: 32; Figs 3-4. [Kagoshima Bay, Kagoshima Prefecture] 

Philocheras victoriensis (Fulton & Grant, 1902) 

= Pontophilus victoriensis Fulton & Grant, 1902: 65; Plate 10, fig. 2. [Altona Bay and Beaumaris, Port 

Phillip, about 5 fms] 

Philocheras wilkinsae De Grave, 2000: 49; Figs 1-4. [north-western side of Laing Island lagoon, Hansa Bay, 

Madang Province, Papua New Guinea, 4°10'30''S 144°52'47''E, 7 m] 

Philocheras wolffi (Holthuis, 1951a) 

= Pontophilus wolffi Holthuis, 1951a: 170; Fig. 34. [off French Guinea, 9°27'N 14°48'W, 50 m] 

Philocheras yaldwyni (Zarenkov, 1968b) 

= Pontophilus yaldwyni Zarenkov, 1968b: 165; Fig. 10. [southwest of New Zealand, 334 m] 

Placopsicrangon Komai & Chan, 2009 

= Placopsicrangon Komai & Chan, 2009 (type species Placopsicrangon formosa Komai & Chan, 2009, by 


Placopsicrangon formosa Komai & Chan, 2009: 256; Figs 1-6. [22°26.44'N 122°21.18'E, 4824-4807 m] 

Pontocaris Spence Bate, 1888 

= Pontocaris Spence Bate, 1888 (type species Pontocaris propensalata Spence Bate, 1888, designated by 

Holthuis, 1947b, gender feminine; name placed on the Official List of Generic Names in Zoology in 


= Pontocheras Bruce, 1988h (type species Pontocheras arafurae Bruce, 1988h, by original designation 


Pontocaris affinis affinis (Alcock, 1901) 

= Aegeon affine Alcock, 1901: 118. [off Bombay, 56-58 fms] 

Pontocaris affinis allodactylus Chan, 1996: 303; Fig. 12. [Red Sea, north of Massawa Channel] 

Pontocaris arafurae (Bruce, 1988h) 

= Pontocheras arafurae Bruce, 1988h: 213; Figs 1-6, 7A-D. [Arafura Sea, 11°04.5'S 132°04.5'E, 15-22 

m] 

Pontocaris hilarula (De Man, 1918b) 

= Aegeon propensalata var. hilarula De Man, 1918b: 301. [Bay of Bima, near south fort, 55 m] 

Pontocaris laurentae Chan, 1996: 307; Fig. 14. [Indonesia, Makassar Strait, 1°56.6'S 119°16.7'E, 134-186 m] 

Pontocaris major Chan, 1996: 304, Fig. 13. [Philippines, 14°0.3'N 120°17'E, 187-210 m] 

Pontocaris pennata Spence Bate, 1888: 499; Plate 91. [Challenger stn 190, 8°56'S 136°5'E, Arafura Sea, south 

of Papua, 49 fms] 

= Aegeon obsoletum Balss, 1914b: 70; Plate 1, fig. 3. [bei Enoshima, 80 m; Dzushi, 50-100 m; Sagamibai; 

Sagamibai, zwischen Ito und Hatsushima, 150 m] 

Pontocaris profundior Chan, 1996: 294; Fig. 8. [Red Sea, off Sudan, 21°33'N 38°21'E, 753-804 m]


Pontocaris propensalata Spence Bate, 1888: 496; Plate 86, fig. 5; Plate 90, figs 2-3. [Challenger stn 192, 

5°49'15''S 132°4'15''E, off the Ki Islands, south of Papua, 140 fms] 

= Pontocaris media Alcock & Anderson, 1899: 282. [Andamans, 55 fms] 

Pontocaris sibogae (De Man, 1918b) 

= Aegeon Sibogae De Man, 1918b: 302. [7°2'.6S 115°23'.6E, Bali Sea, south of Kangeang, 100 m] 

= Aegeon Sibogae var. intermedia De Man, 1918b: 303 [10°27'.9S 123°28'.7E, between the islands of 

Rotti and Timor, 216 m] 

Pontocaris spinifera Chan, 1996: 317; Fig. 18. [Indonesia, Saleh Bay, Sumbawa, Flora Sea, 8°19'S 117°41'E, 

274 m] 

Pontophilus leach, 1817 [in leach, 1815-1875] 

= Pontophilus Leach, 1817 [in Leach, 1815-1875] (type species Crangon spinosus Leach, 1816b, by 



= Cheraphilus Kinahan, 1862 (nomen novum for Pontophilus Leach, 1817 [in Leach, 1815-1875], gender 


Pontophilus brevirostris Smith, 1881: 435. [Fish Hawk stns 865 (40°05'00''N 70°23'00''W, 65 fms); 866 

(40°05'18''N 70°22'18''W, 65 fms); 867 (40°05'42''N 70°22'06''W, 64 fms); 870 (40°02'36''N 70°22'58''W, 

155 fms); 871 (40°02'54''N 70°23'40''W, 11 fms); 872 (40°05'39''N 70°23'52''W, 86 fms); 873 (40°02'00''N 

70°57'00''W, 100 fms); 874 (40°00'00''N 70°57'00''W, 85 fms); 877 (39°56'00''N 70°54'18''W, 126 fms); 

878 (39°55'00''N 70°54'15''W, 142.5 fms)] 

Pontophilus norvegicus (M. Sars, 1861) 

= Crangon norvegicus M. Sars, 1861: 248. [Florøen; Manger; Bundeflorden ved Christiana, 30-50 m] 

? = Hippolyte costata Leuckart, 1847: 89. [Helgoland] 

Pontophilus spinosus (Leach, 1816b) 

= Crangon spinosus Leach, 1816b: 346. [in Danmoniæ australi mari; apud Plymouth Sound] 

Prionocrangon Wood-Mason in Wood-Mason & Alcock, 1891c 

= Prionocrangon Wood-Mason in Wood-Mason & Alcock, 1891c (type species Prionocrangon ommatosteres 

Wood-Mason in Wood-Mason & Alcock, 1891c, by monotypy, gender feminine; name placed 

on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

Prionocrangon curvicaulis Yaldwyn, 1960: 46; Fig. 10. [Chatham Rise, 43°40'S 179°28'E, 220 fms] 

Prionocrangon demani J.N. Kim & Chan, 2005: 1617; Fig. 9. [Sulu Sea, 10°00.70'-10°01.62'N 120°54.72'- 

120°55.27'E, 1482-1488 m] 

Prionocrangon dofleini Balss, 1913b: 238. [Sagamibai, 350-600 m] 

Prionocrangon formosa J.N. Kim & Chan, 2005: 1615; Figs 8, 11B. [24°48.86'N 122°5.31'E, 365-447 m] 

Prionocrangon ommatosteres Wood-Mason in Wood-Mason & Alcock, 1891c: 362. [Andaman Sea, 

11°25'5"N 92°47'6"E, 405 fms] 

Prionocrangon paucispina J.N. Kim & Chan, 2005: 1620; Figs 10, 11C. [22°0.54'N 119°27.94'E, 2334-2543 m] 

Prionocrangon pectinata Faxon, 1896: 157; Plate 2, figs 4-7. [off Martinique, 565 fms] 

Pseudopontophilus Komai, 2004b 

= Pseudopontophilus Komai, 2004b (type species Pseudopontophilus serratus Komai, 2004b, by original 


Pseudopontophilus serratus Komai, 2004b: 76; Figs 1-6. [New Caledonia, Sponge Bank, Norfolk Ridge, 

24°55.1'S 168°21.6'E, 510 m] 

Rhynocrangon Zarenkov, 1965 

= Rhynocrangon Zarenkov, 1964 (unavailable name under Art. 13.3) 

= Rhynocrangon Zarenkov, 1965 (type species Crangon (Sclerocrangon) sharpi Ortmann, 1896b, by 


Rhynocrangon alata (Rathbun, 1902a)


= Sclerocrangon alata Rathbun, 1902a: 891. [Admiralty Inlet, Puget Sound, 40 fms] 

Rhynocrangon rugosa Komai & Komatsu, 2008: 184; Figs 1-5. [off Kuji, Iwate prefecture, northeastern Japan, 

40°14.98'N 142°06.64'E to 40°13.31'N 142°7.38'E, 156 m] 

Rhynocrangon sharpi (Ortmann, 1896b) 

= Crangon (Sclerocrangon) sharpi Ortmann, 1896b: 178. [Alaska, Kodiak Archipelago, Marmot Isle, 45 

fms] 

Sabinea Ross, 1835 

= Sabinea Ross, 1835 (type species Crangon septemcarinata Sabine, 1824, by monotypy; name placed 

on the Official List of Generic Names in Zoology in Opinion 470 in 1957) 

= Myto Krøyer, 1845 (type species Myto Gaimardii Krøyer, 1845 (junior subjective synonym of Crangon 

septemcarinata Sabine, 1824), by monotypy, gender masculine) 

Sabinea hystrix (A. Milne-Edwards, 1881b) 

= Paracrangon hystrix A. Milne-Edwards, 1881b: 6. [près de la Guadeloupe, 734 brasses] 

= Sabinea princeps Smith, 1882: 38; Plate 8, Figs 1-1b. [Blake stns 312, 39°50'45''N 70°11'0''W, 466 fms; 

326, 33°42'15''N 76°0'50''W, 464 fms; 337, 38°20'8''N 73°23'20''W, 740 fms; Albatross stns 892, off Block 

Island, 39°46'N 71°5'W, 487 fms; 893, 39°52'20''N 70°58'W, 372 fms] 

Sabinea sarsii Smith, 1879 

= Sabinea Sarsii Smith, 1879: 59; Plate 11, Figs 6-8. [St. George’s Banks, 60 fms; Gulf of Maine, about 

east-southeast from Cape Sable, Nova Scotia, 42°40'N 66°58'W, 112 fms; Le Have Bank, 60 fms; Lofoten 

Islands, coast of Norway] 

Sabinea septemcarinata (Sabine, 1824) 

= Crangon septemcarinata Sabine, 1824: ccxxxvi; Plate 2, figs 11-13. [west coast of Davis Strait] 

? = Myto gaimardi Krøyer, 1845: 470. [Spitsbergen] 

Sclerocrangon G.o. Sars, 1883 

= Sclerocrangon G.O. Sars, 1883 (type species Cancer Boreas Phipps, 1774, by monotypy, gender feminine; 


Sclerocrangon atrox Faxon, 1893: 199. [Albatross stns 3418 (16°33'00''N 99°52'30''W, 660 fms); 3424 

(21°15'00''N 106°23'00''W, 676 fms)] 

Sclerocrangon boreas (Phipps, 1774) 

= Cancer Boreas Phipps, 1774: 190; Plate 12, fig. 1. [stomach of a seal, caught near the coast of Spitzbergen] 

= Cancer homaroides O. Fabricius, 1780: 241. [Umitak] 

= Cancer arctica Dewhurst, 1834: 254. [Northern and Greenland Ocean] 

Sclerocrangon derjugini Kobjakova, 1936a: 224. [Okhotsk Sea, 182-664 m; fully described in Kobjakova, 

1937] 

Sclerocrangon ferox (G.O. Sars, 1877) 

= Cheraphilus ferox G.O. Sars, 1877: 239. [62°10.2N 4°59.6E, 417 fms] 

Sclerocrangon igarashii Komai & Amaoka, 1991: 27; Figs 1-4. [off Urup Island, Kurile Islands, 45°17'2N 

149°28'2E, 450 m] 

Sclerocrangon salebrosa (Owen, 1839) 

= Crangon salebrosus Owen, 1839: 88; Plate 27, fig. 1. [shores of Kamtschatka] 

= Sclerocrangon gasuyebi Yokoya, 1933: 41; Fig. 22. [Sôyô-Maru stns 602, north of Is. Sado, 205 m; 628, 

south-west of Oga, 148 m; 652, Tugaru Strait, 110 m] 

Sclerocrangon unidentata Komai & Takeda, 1989: 77, Figs 1-5. [off Miyako, Iwate Pref., northeastern Honshu, 

600-700 m] 

Sclerocrangon zenkevitchi Birstein & Vinogradov, 1953: 218; Figs 2-5. [Bering Sea, 2995-3940 m] 

Syncrangon J.N. Kim & Hayashi, 2003 

= Syncrangon J.N. Kim & Hayashi, 2003 (type species Crangon angusticauda De Haan, 1844 [in De 

Haan, 1833-1850], by original designation, gender feminine) 

Syncrangon angusticauda (De Haan, 1844 [in De Haan, 1833-1850])


= Crangon angusticauda De Haan, 1844 [in De Haan, 1833-1850]: Plate 45, fig. 15 (1844); 183 (1849). 

[Japan] 

Syncrangon dentata (Balss, 1914b) 

= Crangon (Sclerocrangon) angusticauda var. dentata Balss, 1914b: 65; Fig. 40 (as Sclerocrangon angusticauda 

dentala). [Negishi Harbour bei Yokohama, 45 m, and Dzushi, 130 m] 

Vercoia Baker, 1904 

= Vercoia Baker, 1904 (type species Vercoia gibbosa Baker, 1904, by monotypy, gender feminine) 

Vercoia gibbosa Baker, 1904: 158; Plate 31. [South Australia, 36-54 m; see Ďuriš, 1992] 

Vercoia interrupta J.N. Kim & Fujita, 2004: 2014; Figs 1-7. [Cape Maeda, Okinawa island, Ryukyu Islands, 

Japan, 8.5 m] 

Vercoia japonica Komai, 1995b: 123; Figs 1-4. [Omurodashi, off Izu-Ohshima Island, 34°31.7'N 139°23.2'E, 

138-167 m] 

Vercoia socotrana Ďuriš, 1992: 1448; Figs 7-11. [Socotra Island, Gulf of Aden, western Indian Ocean, 

12°21'08''N 54°28'00''E, 80 m] 

Family GlyPHoCRANGoNIDAe Smith, 1884 

Glyphocrangon A. Milne-edwards, 1881b 

= Thalascaris Spence Bate, 1878 (type species Glyphocrangon rimapes Spence Bate, 1888, designated by 

Holthuis, 1971, gender feminine; name suppressed for the purposes of the Principle of Priority but 

not for those of the Principle of Homonymy, placed on the Official Index of Rejected and Invalid 


= Glyphocrangon A. Milne-Edwards, 1881b (type species Glyphocrangon spinicauda A. Milne-Edwards, 

1881b, by original designation, gender feminine; name placed on the Official List of Generic 


= Rhachocaris Smith, 1882 (type species Rhachocaris Agassizii Smith, 1882 (junior subjective synonym 

of Glyphocrangon aculeatum A. Milne-Edwards, 1881b), designated by Fowler, 1912, gender feminine) 

= Plastocrangon Alcock, 1901 (type species Glyphocrangon cæcescens Wood-Mason in Wood-Mason & 

Alcock, 1891c, designated by Fowler, 1912, gender feminine) 

Glyphocrangon aculeata A. Milne-Edwards, 1881b 

= Glyphocrangon aculeatum A. Milne-Edwards, 1881b: 5. [à Saint-Vincent, 593 brasses] 

= Rhachocaris Agassizii Smith, 1882: 43; Plate 5, fig. 2; Plate 6, fig. 2. [Blake stn 326, 33°42'15''N 

76°0'50''W, 464 fms] 

Glyphocrangon acuminata Spence Bate, 1888: 522; Plate 94, Figs 2-3. [Challenger stn 175, Fiji Islands, 19°02'S 

177°10'E, 2468 m; lectotype designation by Komai, 2004c] 

Glyphocrangon africana Komai, 2010b: 84; Figs 1-4. [off Cape Point, South Africa, 2269 m] 

Glyphocrangon alata Faxon, 1893: 201. [Albatross stns 3395 (North Pacific Ocean, 07°30'36''N 078°39'00''W), 

730 fms; 3418 (North Pacific Ocean, 16°33'00''N 098°52'30''W), 660 fms; ?] 

Glyphocrangon albatrossae Komai, 2004c: 468; Figs 40-41. [Philippines, South China Sea, off W Luzon, 

15°58.15'N 119°40.20'E, 1719 m] 

Glyphocrangon alispina Chace, 1939: 39. [off Bahia Cardenas, Matanzas Province, Cuba, 23°24'N 

81°00'30''W, 370-605 fms] 

Glyphocrangon amblytes Komai, 2004c: 561; Figs 89-90. [Madagascar, 22°25.3'N 43°04.9'E, 525-550 m] 

Glyphocrangon andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c 

= Glyphocrangon investigatoris var. andamanensis Wood-Mason in Wood-Mason & Alcock, 1891c: 356; 

accompanying illustration in Wood-Mason, 1894: Plate 6, figs 2-2ba. [Andaman Sea, 11°31.40'N 

92°46'40''E, 188-220 fms] 

Glyphocrangon arduus Komai, 2007: 566; Figs 1-3. [French Polynesia, Austral Islands, off Tubuai Island, 

23°21.4'S 149°34.2'W, 600-1200 m] 

Glyphocrangon armata Komai, 2004c: 570; Figs 95-96. [New Caledonia, 23°10.41'S 166°49.16'E, 870-1000 m] 

Glyphocrangon assimilis De Man, 1918b: 294. [7°19'.4S 116°49'.5E, 538 m; lectotype designation by Komai, 

2004c]


Glyphocrangon atlantica Chace, 1939 

= Glyphocrangon (Plastocrangon) caecescens atlantica Chace, 1939: 40. [south of Santa Clara Province, 

Cuba, 20°47'N 80°24'W, 2125 fms] 

Glyphocrangon aurantiaca Holthuis, 1971: 303; Fig. 8. [Off Tobago, 11°37.3'N 60°59.4'W to 11°41'N 

61°01.3'W, 400-700 fms] 

Glyphocrangon boletifera Komai, 2004c: 558; Figs 87-88. [Gulf of Aden, 13°06.12'N 46°24.30'E, 1061-1080 

m] 

Glyphocrangon brevis Komai, 2006c: 250; Figs 1A, 2A, 3. [Madagascar, 17°40'S 43°12'E, 1075-1175 m] 

Glyphocrangon caeca Wood-Mason in Wood-Mason & Alcock, 1891c 

= Glyphocrangon cæca Wood-Mason in Wood-Mason & Alcock, 1891c; 358 accompanying illustration 

in Wood-Mason, 1894: Plate 7, figs 1-1a. [Bay of Bengal, 13°47'30''N 92°36'E, 561 fms] 

Glyphocrangon caecescens Wood-Mason in Wood-Mason & Alcock, 1891c 

= Glyphocrangon cæcescens Wood-Mason in Wood-Mason & Alcock, 1891c: 357; accompanying illustration 

in Wood-Mason, 1894: Plate 7, figs 5-5a. [Bay of Bengal, 11°58'N 88°52'17''E, 1748 fms] 

Glyphocrangon cerea Alcock & Anderson, 1894: 151. [Investigator stn 150, Laccadive Sea, off the Northern 

Maldive Atoll, 7°05'45''N 75°04'E, 719 fms] 

Glyphocrangon chacei Komai, 2004c: 534; Figs 73-74. [Philippines, between Bohol and Siquijor Islands, 

Mindanao Sea, 09°22.30'N 123°47.40'E, 719 m] 

Glyphocrangon confusa Komai, 2004c: 597; Figs 110, 112A-D. [Indonesia, Halmahera, 00°15.00'N 

127°24.35'E, 545 m] 

Glyphocrangon conodactylus Komai, 2004c: 555; Figs 85-86. [New Caledonia, Mount K, Loyalty Islands, 

24°44.55'S 170°07.00'E, 790-825 m] 

Glyphocrangon cornuta Komai, 2004c: 472; Figs 42-43. [New Caledonia, 23°56'S 166°41'E, 2660-2750 m] 

Glyphocrangon crosnieri Komai, 2004c: 431; Figs 22-23, 115. [Madagascar, 22°26'S 43°03'E, 540 m] 

Glyphocrangon demani Komai, 2006c: 253; Figs 1B, 2B, 4. [Indonesia, between Ceram and New Guinea, 

03°37.7'S 131°26.4'E, 924 m] 

Glyphocrangon dentata Barnard, 1926 

= Glyphocrangon gilesii var. dentata Barnard, 1926: 128. [25°59'S 33°31'E, 540 m] 

Glyphocrangon dimorpha Komai, 2004c: 459; Figs 37-38. [Norfolk Ridge, 23°56'S 168°28'E, 1846-1862 m] 

Glyphocrangon elephas Komai, 2005: 253; Figs 1-3. [Southern Australia, Great Australian Bight, 33°17'S 

128°32'E, 360 m] 

Glyphocrangon faxoni De Man, 1918b 

= Glyphocrangon (Plastocrangon) Faxoni De Man, 1918b: 298. [7°15'S 115°15'.6E, Bali Sea, 289 m; 

10°39'S 123°40'E, between the islands of Roti and Timor, 520 m] 

Glyphocrangon ferox Komai, 2004c: 506; Figs 59-60. [Madagascar, 13°16'S 47°33'E, 1490-1600 m] 

Glyphocrangon fimbriata Komai & Takeuchi, 1994: 458; Figs 1-3. [Sio Guyot, Mid-Pacific Mountains, 

18°16.05'N-18°15.87'N 171°20.99'E-171°22.01'E, 1300-1312 m] 

= Glyphocrangon joani Allen & Butler, 1994: 435; Figs 10-14. [Mid-Pacific Ocean, 171°15.0'E 18°14.0'N, 

1281-1651 m] 

Glyphocrangon formosana Komai, 2004c: 594; Figs 108, 109E-H. [Taiwan, off Su-Aou, I-Lan County, NE of 

Taiwan, 300-400 m] 

Glyphocrangon gilesii Wood-Mason & Alcock, 1891b 

= Glyphocrangon Gilesii Wood-Mason & Alcock, 1891b: 193; accompanying illustration in Wood- 

Mason, 1894: Plate 7, Figs 4-4a. [eight miles south-east of Cinque Island, in the Andaman Sea, 500 

fms] 

Glyphocrangon grandis Komai & Chan, 2008: 41; Figs 1-5, 12D. [3°21.32'N 124°12.26'E, 1037-1100 m] 

Glyphocrangon granulosis Spence Bate, 1888: 507; Plate 92; Plate 93, fig. 1. [Challenger stn 218, 02°33'S 

144°04'E, between New Guinea and Admiralty Islands, 1070 fms] 

Glyphocrangon haematonotus Holthuis, 1971: 315; Figs 6c, 7c. [Straits of Florida, 26°34'N 79°43'W, 270 fms] 

Glyphocrangon hakuhoae Takeda & Hanamura, 1994: 24; Figs 11-13. [Flores Sea, 05°47.3'N 119°35.4'E - 

05°46.9'N 119°34.6'E, 280 m] 

Glyphocrangon hastacauda Spence Bate, 1888: 519; Plate 93, fig. 5. [Challenger stn 232, 35°11'N 139°28'E, off 

Japan, 345 fms]


Glyphocrangon holthuisi Kensley, Tranter & Griffin, 1987: 321; Figs 22, 25C. [New South Wales, south-east 

of Cape Byron, 28°41'S 153°52'E, 153 m] 

Glyphocrangon humilis Komai, 2006c: 254; Figs 1C, 2C, 5. [Japan, Tosa Bay, 33°00'N 133°35'E, 660-700 m] 

Glyphocrangon indonesiensis Komai, 2004c: 408-412; Figs 10-11. [Indonesia, Tanimbar Islands, 08°16'S 

131°59'E, 769-809 m] 

Glyphocrangon investigatoris Wood-Mason & Alcock, 1891b: 191; accompanying illustration in Wood- 

Mason, 1894: Plate 6, Figs 3-3b. [19°35'N 92°24'E, 272 fms] 

Glyphocrangon isos Komai & Chan, 2008: 54; Figs 9-11, 15. [14°55'N 123°8'E, 922-909 m] 

Glyphocrangon kapala Komai, 2004c: 428; Figs 20-21. [Eastern Australia, East of Broken Bay, New South 

Wales, 33°35'S 152°00'E, 810 m] 

Glyphocrangon lineata Komai, 2004c: 545; Figs 79-80. [Indonesia, Kai Islands, 05°15'S 132°59'E, 769-809 m] 

Glyphocrangon longipes Komai, 2004c: 567; Figs 93-94. [Philippines, North of Palawan Passage, 10°57.45'N 

138°38.15'E, 686 m] 

Glyphocrangon longirostris (Smith, 1882) 

= Rhachocaris longirostris Smith, 1882: 51; Plate 5, fig. 1; Plate 6, fig. 1. [Blake stn 330, 31°41'N 74°35'W, 

1047 fms] 

Glyphocrangon longleyi Schmitt, 1931b: 393. [south of Tortugas, 18-220 fms] 

Glyphocrangon loricata Faxon, 1895: 140; Plate 38bis. [Albatross stns 3402 (South Pacific Ocean, 00°57'30''S 

089°03'30''W), 421 fms; and 3410 (North Pacific Ocean, 00°19'00''N 090°34'00''W), 331 fms] 

Glyphocrangon lowryi Kensley, Tranter & Griffin, 1987: 322; Figs 23, 25D. [Queensland, north-east of Danger 

Point, 28°03'S 154°04'E, 720 m] 

Glyphocrangon mabahissae Calman, 1939: 217, fig. 8. [John Murray Expedition stn 115 (05°05.18'S 

39°22.12'E), Zanzibar area, 640-658 m] 

Glyphocrangon major Komai, 2004c: 514; Figs 63-64. [Indonesia, N Makassar Strait, 0°34.6'N 119°08.5'E, 

1301 m] 

Glyphocrangon megalophthalma De Man, 1918b: 296. [4°22'.1S 118°16'.9E, Strait of Makassar, 2029 m; lectotype 

designation by Komai, 2004c] 

Glyphocrangon musorstomia Komai, 2006c: 256; Figs 1D-E, 6-7. [Wallis and Futuna Islands, Tuscarora 

Bank, 11°47.4'S 178°25.3'E, 900 m] 

Glyphocrangon neglecta Faxon, 1896: 159; Plate 1, Figs 5-6. [off Grenada, 340 fms; off Montserrat, 303 fms; 

off Grenada, 291 fms] 

Glyphocrangon nobilis A. Milne-Edwards, 1881b 

= Glyphocrangon nobile A. Milne-Edwards, 1881b: 5. [près de la Dominique, 1131 brasses; neotype 

designation by Holthuis, 1971; see discussion in Forest & Holthuis, 1997] 

Glyphocrangon novaecastellum Kensley, Tranter & Griffin, 1987: 324; Figs 24, 25E. [New South Wales, 

south-east of Newcastle, 33°08'S 152°27'E, 720 m] 

Glyphocrangon panglao Komai & Chan, 2008: 47; Figs 6-8, 13B1-B2. [8°51'N 123°10'E, 982-1040 m] 

Glyphocrangon parva Komai, 2004c: 412; Figs 12-13. [Philippines, Sulu Sea, East of Palawan, 09°13'N 

118°51.15'E, 2021 m] 

Glyphocrangon parviocullus Komai, 2006c: 259; Figs 1F, 2E, 8. [New Caledonia, 23°10'S 166°49'E, 870- 

1000 m] 

Glyphocrangon perplexa Komai, 2004c: 579; Figs 101-102. [East China Sea, 29°21.5'N 127°28.7'E, 543-600 m] 

Glyphocrangon podager Spence Bate, 1888: 516; Plate 93, fig. 2. [Challenger stn 146, 46°46'S 45°31'E, near 


Glyphocrangon priononota Wood-Mason & Alcock, 1891b: 192; accompanying illustration in Wood-Mason, 

1894: Plate 6, figs 1-1a. [Investigator stn 104 (Laccadive Sea, 11°12'47''N 74°25'30''E), 1000 fms] 

Glyphocrangon proxima Komai, 2004c: 416; Figs 14-15. [Indonesia, Kai Islands, 05°14'S 133°00'E, 688-694 

m] 

Glyphocrangon pugnax De Man, 1918b: 293. [10°39'S 123°40'E, between the islands of Roti and Timor, 520 

m; lectotype designation by Komai, 2004c] 

Glyphocrangon punctata Komai, 2004c: 475; Figs 44-45. [Philippines, South of Mindoro, 06°06.78'N 

125°58.91'E, 2820-2770 m]


Glyphocrangon regalis Spence Bate, 1888: 517; Plate 93, figs 3-4. [Challenger stn 194, 4°34'S 129°57'30"E, off 

Banda Island, 200 fms; see Komai, 2004c] 

= Glyphocrangon juxtaculeata Chace, 1984: 15; figs 3-4. [Indonesia, off southern Buru, 3°47.15'S 

126°23.40'E, 946 m] 

Glyphocrangon richeri Komai, 2004c: 495; Figs 54-55. [Wallis and Futuna Islands, SW slope of Combe 

Bank, 12°34'S 178°11'W, 1280-1300 m] 

Glyphocrangon rimapes Spence Bate, 1888: 523; Plate 94, fig. 4. [Challenger stns 300, 33°42'S 78°18'W, near 

Juan Fernandez, 1375 fms; 237, 34°37'N 140°32'E, near Yokohama, 1875 fms; 331, 37°47'S 30°20'W, 

South Atlantic, between Buenos Ayres and Tristan de Cunha, 1715 fms] 

Glyphocrangon robusta Komai, 2004c: 526; Figs 69-70. [Philippines, South of Mindoro, 11°59'N 121°13'E, 

320-337 m] 

Glyphocrangon rubricinctuta Komai, 2004c: 599; Figs 111, 112E-H. [Wallis and Futuna Islands, Bayonnaise 

Bank, 11°54.2'S 179°31.4'W, 597-600 m] 

Glyphocrangon rudis Komai, 2006c: 261; Figs 1G, 2F, 9. [Solomon Islands, 08°31.2'S 160°37.7'E, 1036- 

1138 m] 

Glyphocrangon runcinata Komai, 2004c: 420; Figs 16-17. [East China Sea, East of Kuchiminose Bank, 

29°21.5'N 127°28.7'E, 543-600 m] 

Glyphocrangon saintlaurentae Komai, 2004c: 487; Figs 50-51. [Réunion Island, 21°28.1'S 56°32.4'E, 4030 m] 

Glyphocrangon sculpta (Smith, 1882) 

= Rhacocharis sculpta Smith, 1882: 49; Plate 5, fig. 3; Plate 6, figs 3-3d. [Blake stn 339, 38°16'45''N 

73°10'30''W, 1186 fms] 

Glyphocrangon sibogae De Man, 1918b 

= Glyphocrangon Sibogae De Man, 1918b: 295. [Flores Sea, 07°24'S 118°15.2'E, 794 m; lectotype designation 

by Komai, 2004c] 

Glyphocrangon sicaria Faxon, 1893 

= Glyphocrangon sicarius Faxon, 1893: 202. [Albatross stn 3382 (North Pacific Ocean, 06°21'00''N 

080°41'00''W), 1793 fms] 

Glyphocrangon similior Komai, 2004c: 590; Figs 107, 109A-D. [Vanuatu, off Epi, 16°33'S 167°55'E, 602- 

620 m] 

Glyphocrangon smithii Wood-Mason in Wood-Mason & Alcock, 1891c 

= Glyphocrangon Smithii Wood-Mason in Wood-Mason & Alcock, 1891c: 357; accompanying illustration 

in Wood-Mason, 1894: Plate 7, figs 3-3a. [Bay of Bengal, 13°47'30''N 92°36'E, 561 fms] 

Glyphocrangon speciosa Komai, 2004c: 500; Figs 56-58. [New Caledonia, 25°04'S 168°46'E, 1093-1193 m] 

Glyphocrangon spinicauda A. Milne-Edwards, 1881b: 3. [près de St-Kitts, 250 brasses] 

Glyphocrangon spinossisima Brand & Takeda, 1996: 264; Figs 1-4. [Sulu Sea, 08°31.6'S 118°35.7'E, 1712- 

1840 m] 

Glyphocrangon spinulosa Faxon, 1893: 202. [Albatross stns 3353 (North Pacific Ocean, 07°06'15''N 

080°34'00''W), 695 fms; 3418 (North Pacific Ocean, 16°33'00''N 099°52'30''W), 660 fms; 3419 (North 

Pacific Ocean, 16°34'30''N 100°03'00''W), 672 fms; 3424 (North Pacific Ocean, 21°15'00''N 

106°23'00''W), 676 fms; 3425 (North Pacific Ocean, 21°19'00''N 080°34'00''W), 695 fms; 3435 (North 

Pacific Ocean, 26°48'00''N 110°45'20''W), 859 fms] 

Glyphocrangon stenolepis Chace, 1984: 22; Figs 5-6. [South China Sea, southeast of Hong Kong, 20°31'N 

115°49'E, 485 m] 

Glyphocrangon taludensis Hendrickx, 2010: 359; Figs 1-3. [18°26'45''N 104°16'10''W, 1858-1879 m] 

Glyphocrangon tasmanica Komai, 2004c: 574; Figs 97-98. [Lord Howe Ridge, 400 m] 

Glyphocrangon unguiculata Wood-Mason & Alcock, 1891b: 193; accompanying illustration in Wood-Mason, 

1894: Plate 7, figs 2-2a. [Investigator stn 105 (Arabian Sea, 15°2'N 72°34'E), 740 fms] 

Glyphocrangon vicaria Faxon, 1896: 159. [0°54'N 91°9'W, 1189 fms] 

Glyphocrangon wagini Burukovsky, 1990: 206; Fig. 4B (as G. wagigi), 5A. [24°56'S 88°31'W, 580-564 m]


unavailable names 

Macrobrachium divakarani Jayachandran, 2001: 91; Fig. 21. [unavailable under Art. 16.4.1] 

Macrobrachium josephi Jayachandran, 2001: 123; Fig. 32. [Veli Lake, Kulathoor rivulet (Kerala, India); 

unavailable under Art. 16.4.1] 

Allobrachium Jayachandran, 2001 [unavailable under Art. 13.3] 

Penaeoidea larvata 

Cerataspis affinis Kishinouye, 1926: 66; Fig. 2. [Between Kyushyu and the Ryukyu Archipelago, Japan] 

Cerataspis longiremis Dohrn, 1871: 360; Plates 28-29. [dessen Herkunft mir aber entfallen is] 

Cerataspis monstruosus Gray, 1828: 8; Plate 6, fig. 5. [Found in stomach of a delphin off the coast of Brazil] 

Cerataspis Petitii Guérin-Méneville, 1844 [in Guérin-Méneville, 1829-1844]: 18; Plate 22. [trouvé en pleine 

mer dans l’Océan Indien] 

Cryptopus Defrancii H. Milne Edwards, 1837 [in H. Milne Edwards, 1836-1844]: 149; Plate 54bis, fig. 4. [la 

Méditerranée] 

Euphema armata H. Milne Edwards, 1837 [in H. Milne Edwards, 1834-1840]: 421. [dans l’océan Atlantique 

austral] 

Euphema polyacantha Ortmann, 1893: 77. [Plankton-Expedition der Humbolt-Stiftung, JN 252 (9.4°S 

41.9°W, 0-400 m), Guineastrom] 

Hoplites longirostris Philippi, 1857: 329; Plate 14, figs 1a-i. [Atlantischen Ocean in 25°N.B. und 22°50'W.L.] 

Loxopis tridens Dana, 1852b: 661 (1852); Plate 44, fig. 4 (1855). [Sooloo Archipelago] 

Ophthalmeryon transitionalis Spence Bate, 1889: 6; Plate 9. [swallowed by a dolphin, and therefore been 

affected somewhat by the gastric juices of the fish’s stomach] 

Opisthocaris mülleri Ortmann, 1893: 77; Plate 4, fig. 5. [Plankton-Expedition der Humbolt-Stiftung, Floridastrom, 

JN 45 (41.6°N 56.3°W, 0-200 m) and 47 (40.4°N 57.0°W, 0-300 m); Südl. Aequatorialstrom 

JN 218 (3.8°S 32.6°W, 0-400 m), 228 (1.8°S 38.1°W, 0-600 m), 231 (1.5°S 39.2°W, 0-400 m), 232 (0.4°S 

42.4°W, 0-400 m), 235 (0.1°S 44.2°W, 0-400 m), 246 (0.4°S 46.6°W, 0-400 m)] 

Platysacus crenatus Spence Bate, 1888: 363; Plate 63. [Challenger stn 352, on the surface of the Atlantic 

Ocean, Sierra Leone, 10°55'N 17°46'W] 

Rachitia spinalis Dana, 1852b: 667 (1852); Plate 44, fig. 8 (1855). [Atlantic, off the harbour of Rio de 

Janeiro] 

Solenocera sp. larva barbata Heegaard, 1966: 89; Figs 181-192. [Great Barrier Reef stn 46, 14°32'S 145°32'E, 

inside Cooks Passage, 33 m; Great Barrier Reef Expedition, no further data; Discovery stn 276, 

5°54'00"S 11°19'00"E, 110-0 m] 

Solenocera sp. larva danae Heegaard, 1966: 53; Figs 66-108. [Dana stns 3922-II, 3°45'S 56°33'E, 600 m wire 

out; 3903-II, III, IV, 5°30'N 93°38'E, 600, 300, 100 m wire out resp.; 3860-XX, XXI, 2°57'S 99°36'E, 600, 

300 m wire out resp.] 

Solenocera sp. larva elongata Heegaard, 1966: 85; Figs 65, 170-180. [Terra Nova stn 43, 22°06'S 39°40'W, 

surface] 

Solenocera sp. larva nodulosa Heegaard, 1966: 81; Figs 154-169. [Dana stn 3921-III, 3°36'S 58°19'E 300 m 

wire out] 

Solenocera sp. larva sumatransis Heegaard, 1966: 66; Figs 109-153. [Dana stn 3903-III, 5°30'N 93°28'E, 300 

m wire out] 

Penaeoidea Nomina Dubia 

Hemipenaeus dubius Spence Bate, 1881: 187. [Among Philippine Islands, less than 20 fms] 

Pandalus tenuicornis Rankin, 1900: 544. [Type locality not indicated] 

Penaeus amazonicus Kingsley, 1883: 106. [Upper Amazon] 

Penaeus gracilis Dana, 1852a: 27. [in mari Suluensi; Burkenroad, 1934a considers this an immature Metapenaeopsis 

species] 

Penaeus pubescens Stimpson, 1871: 133. [St. Thomas] 

Penaeus telsodecacanthus Spence Bate, 1881: 182. [in the channels of the Japanese Islands, 8-10 fms]


Penaeus tenuis Dana, 1852a: 27. [in mari Atlantico prope portum ”Rio Negro” Patagoniae] 

Penaeus villosus Guérin-Méneville, 1838 [in Guérin-Méneville, 1829-1838]: 36. [Nouvelle-Irlande; quite 

probably Metapenaeus monoceros or M. mastersii, according to Burkenroad, 1934a] 

Penacus (Protosolenocera) kessleri Czerniavsky, 1878: 23. [Han’kou, Yan-Tse-Kian’, north of Nankin, China] 

Sergestes caudatus Krøyer, 1859: 270; Plate 5, fig. 14. [nordlige Kattegat] 

Penaeoidea Nomina Nuda 

Metapenaeus tweedii Morris & Bennett, 1952: 165. 

Sergestoidea larvata 

Acanthosoma brevitelsonis Spence Bate, 1888: 367; Fig. 51; Plate 64, fig. 1. [Western Pacific Ocean] 

Acanthosoma dohrni Gurney, 1924: 92; Fig. 31. [Terra Nova Expedition stn 129, off Three Kings Island, New 

Zealand; 311, Atlantic, 35°29'S 50°26'W] 

Acanthosoma dorsispinalis Spence Bate, 1888: 370: Plate 65, fig. 1. [Western Pacific Ocean] 

Acanthosoma hispida Gurney, 1924: 90; Figs 29-30. [Terra Nova Expedition stn 129, off Three Kings Island, 

New Zealand] 

Acanthosoma laevirostratis Spence Bate, 1888: 374; Plate 65, fig. 2. [North of Admiralty Islands, between 

Challenger stations 221 and 222] 

Acanthosoma longitelsonis Spence Bate, 1888: 371; Plate 44, fig. 3. [South Pacific Ocean] 

Acanthosoma macrotelsonis Spence Bate, 1888: 373; Plate 66, fig. 1. [Samboangan, Philippines] 

Acanthosoma tynitelsonis Spence Bate, 1888: 369; Plate 64, fig. 2. [Western Pacific Ocean] 

Elaphocaris crassus Spence Bate, 1888: 362; Plate 61, fig. 4. [South Atlantic, between Montevideo and 

Tristan da Cunha, 32°S 45°W] 

Elaphocaris dohrni Spence Bate, 1888: Plate 62. [South Atlantic, 32°S 45°W, between Monte Video and 

Tristan da Cunha] 

Elaphocaris hispida Gurney, 1924: 85; Figs 24-28. [Terra Nova Expedition: stns 65, Atlantic, 23°28'N 34°45'W; 

66, Atlantic, 25°35'N 34°10'W; 109, off New Zealand, 34°15'S 172°E; 120, off New Zealand, 34°26'S 

172°14'E] 

Elaphocaris ortmanni Gurney, 1924: 84; Figs 21-23. [Terra Nova Expedition stn 17, Atlantic, 26°17'N 

18°14'W] 

Elaphocaris suhmi Spence Bate, 1888: 353; Figs 49-50; Plate 61, figs 1-3. [South Atlantic, between Montevideo 

and Tristan da Cunha, 32°S 45°W] 

Erichthina demissa Dana, 1852b: 634; Plate 42, fig. 3. [Near eastern entrance of Straits of Sunda] 

Mastigopus acetiformis Spence Bate, 1888: 376. [Western Pacific Ocean] 

Mastigopus crassus Spence Bate, 1888: 381; Figs 52-53. [Taken on the voyage from Hilo to Tahiti, 5°0'N] 

Mastigopus dorsispinalis Spence Bate, 1888: 375; Plate 65, fig. 3. [Pacific Ocean, NW of the Admiralty Islands, 

1°N 146°E] 

Mastigopus spiniventralis Spence Bate, 1888: 379; Plate 67, fig. 4. [Western Pacific Ocean] 

Mastigopus suhmi Spence Bate, 1888: 378; Plate 66, fig. 2. [North Atlantic Ocean] 

Mastigopus tenuis Spence Bate, 1888: 428; Plate 65, fig. 4. [Western Pacific Ocean] 

Sceletina armata Dana, 1852b: 663. [Atlantic, 0°30'S 17°30'W] 

Sceletina laticeps Dana, 1852b: 664. [Pacific, 30 miles W of Assumption Island, one of the Ladrones] 

Sceletina orientalis Dana, 1852b: 665. [Sooloo Sea, SW of Panay] 

Sciacaris telsonis Spence Bate, 1888: 438; Plate 78, fig. 1. [N of New Guinea] 

Sergestoidea Nomina Dubia 

Acheles arachnipodus Cocco, 1832: 204; unnumbered Plate, fig. 1. [Messina, Sicily] 

Lucifer pacificus Dana, 1852b: 673 (1852); Plate 45, fig. 2. (1855). [Pacific, 15°20'S 148°W] 

Mastigopus spinosus Leuckart, 1853: 258. [Type locality not indicated] 

Phasmatocarcinus discophthalmus Tilesius, 1819: 7; Plate 21a, fig. 10. [placed on the Official Index of Rejected 

and Invalid Works in Zoological Nomenclature in Opinion 864 in 1969] 

Phasmatocarcinus glaucus Tilesius, 1819: 6; Plate 21a, fig. 9. [placed on the Official Index of Rejected and 

Invalid Works in Zoological Nomenclature in Opinion 864 in 1969]


Prionorhynchus Apus Tilesius, 1819: 6; Plate 21a, fig. 7. [placed on the Official Index of Rejected and Invalid 

Works in Zoological Nomenclature in Opinion 864 in 1969] 

Sergestes dorsispinalis Spence Bate, 1888: 394; Plate 72, fig. 1. [S of Australia; possibly a juvenile stage of 

Sergestes arcticus T. Kikuchi (in litt.)] 

Sergestes hanseni Dennell, 1955: 400. [Bermuda area] 

Sergestes laterodentatus Spence Bate, 1888: 395. [S of Australia; possibly a mastigopus stage of Sergestes 

arcticus (T. Kikuchi (in litt.))] 

Sergestes oculatus Perrier, 1886: 81. [Le Challenger et le Talisman, la faune des Sargasses] 

Sergestes praecollus Spence Bate, 1888: 423; Plate 77, fig. 2. [North Pacific Ocean] 

Sergestes sundi Dennell, 1955: 400. [Bermuda area] 

Sergestes tenuis Spence Bate, 1888: 233, Figs 43-45. [North of New Guinea, and on the voyage along the 

Marianne Islands] 

Sergestes utrinquedens Spence Bate, 1888: 433. [North Pacific Ocean] 

Sergestes Colosii Cecchini, 1933: 4; Plate 1, fig. 1. [Ammiraglio Magnaghi stns 138, a sud ouest di Aden, 12°5' 

- 12°7'N 44°47' - 44°45'E, 388 m; 150, a nord est di Porto Sudan, 20°47'N 38°22'30"E, 1200 m] 

Symphysopus hirtus Tilesius, 1819: 9; Plate 21b, fig. 19. [placed on the Official Index of Rejected and Invalid 


Sergestoidea Nomina Nuda 

Sergestes forins Bakus, 1989: 36. 

Caridea larvata 

Amphiplectus depressus Spence Bate, 1888: 623; Plate 110, fig. 3. [Off Recife, Brazil, 9°05'S, 34°50'W, 6640 

m] 

Anebocaris ancylifer Coutière, 1907c: 407; unnumbered figure. [Géorgie du Sud] 

Anebocaris quadroculus Spence Bate, 1888: 722; Plate 123, fig. 1. [Off Port Jackson, Australia, surface; off 

Sibago, Samboangan, Philippine Islands; off Basilan Strait; China Sea, off Luzon; Zebu Harbour, 

Philippine Islands] 

Anisocaris puerilis Coutière, 1907c: 410; unnumbered figure. [Géorgie du Sud] 

Astacus cylindrus Fabricius, 1793: 483. [in Oceani Indico] 

Astacus planatus Fabricius, 1798: 482. [Type locality not indicated] 

Atlantocaris longirostris Ortmann, 1893: 80; Plate 5, fig. 3. [Plankton-Expedition der Humboldt-Stiftung, 

JN 153 (7.9°N 21.4°W), im Guineastrom] 

Camptocaris maxima Ortmann, 1893: 81; Plate 5, fig. 4. [Plankton-Expedition der Humboldt-Stiftung, Sargasso-See, 

JN 60 (35.0°N 62.1°W, 0-300 m); Südl Aequatorialstrom, JN 188 (2.6°S 14.6°W, 0-400 m)] 

Caricyphus acutus Coutière, 1905a: 21; Fig. 7. [Princesse-Alice stn 1834 (37°28'N 25°53'30"W), 0-1000 m] 

Caricyphus angulatus Spence Bate, 1888: 718; Plate 121, fig. 6. [46°46'S 45°31'E, near Marion Island, surface] 

Caricyphus bigibbosus Coutière, 1905c: 1114. [fully described in Coutière, 1905a: 26; Fig. 8]. [Princesse-Alice 

stn 1851 (46°15'N 7°09'W), 0-3000 m] 

Caricyphus cornutus Spence Bate, 1888: 712; Plate 121, fig. 2. [near Samboangan, Philippine Islands] 

Caricyphus edentulus Ortmann, 1893: 74; Plate 2, fig. 3. [Plankton-Expedition der Humboldt-Stiftung, 

Sargasso-See, JN 72 (31.5°N 59.0°W, 0 m); 102 (31.7°N 42.7°W, 0-400 m); Pl.55 (28.3°N 34.3°W, 200 m, 

in table as Nördl. Aequatorialstrom und Canarienstrom)] 

Caricyphus gibberosus Spence Bate, 1888: 716; Plate 121, fig. 4. [near the Sandwich Islands, Pacific Ocean, 

surface] 

Caricyphus serramarginis Spence Bate, 1888: 714; Plate 121, fig. 3. [Kandavu, Fiji Islands, surface] 

Caricyphus targidus Spence Bate, 1888: 717; Plate 121, fig. 5. [off Luzon, China Sea, surface] 

Caulurus pelagicus Stimpson, 1860a: 42. [in Oceano Pacifico, lat. Bor. 34° long. orient. 126°] 

Copiocaris messinensis Thiele, 1905: 454; Plate 15, Figs 17-30. [Straße von Messina] 

Coronocaris brevis Ortmann, 1893: 82; Plate 5, fig. 5. [Plankton-Expedition der Humboldt-Stiftung, Südl. 

Aequatorialstrom JN 234 (0.4°S 42.4°W, 0 m); 235 (0.1°S 44.2°W, 0-400 m); 249 (5.6°S 44.0°W, 0 m); 

Brasilianische Küstenbank, Pl. 112 (0.4°S 46.6°W, 207 m); Pl. 113 (0.4°S 46.6°W, 200 m)]


Coronocaris gracilis Ortmann, 1893: 81; Plate 6, fig. 1. [Plankton-Expedition der Humboldt-Stiftung, 

Nördl. Aequatorialstrom JN 145 (13.3°N 22.7°W, 0-400 m); 146 (12.3°N 22.3°W, 0-400 m); 148 (10.2°N 

22.2°W, 0-400 m); 150 (10.2°N 22.2°W, 0-1000 m); Südl. Aequatorialstrom JN 232 (0.4°S 42.4°W, 0-400 

m); 234 (0.4°S 42.4°W, 0 m); 235 (0.1°S 44.2°W, 0-400 m); 246 (0.4°S 46.6°W, 0-400 m); Pl. 103 (0.4°S 

42.4°W, 200 m); Brasilianische Küstenbank, Pl. 112 (0.4°S 46.6°W, 207 m)] 

Coronocaris humilis Coutière, 1907a: 58; Fig. 21. [Açores, 0-1000 m] 

Diaphoropus longidorsalis Spence Bate, 1888: 688; Plate 117, fig. 4. [Cape Verde Islands] 

Diaphoropus versipellis Spence Bate, 1888: 687; Plate 107, fig. 3. [Off Cape Horn, Australia, surface] 

Eretmocaris corniger Spence Bate, 1888: 900; Plate 145, fig. 4. [Cape Verde] 

Eretmocaris dolichops Ortmann, 1893: 79; Plate 5, fig. 1. [Plankton-Expedition der Humboldt-Stiftung, bei 

Boavista, JN 141 (16.1°N 23.1°W, 0-500 m)] 

Eretmocaris longicaulis Spence Bate, 1888: 897; Plate 145, fig. 2. [17°29'N 141'31°E, south of Japan, surface] 

Eretmocaris stylorostris Spence Bate, 1888: 898; Plate 145, fig. 3. [Off Cape Verde Islands, surface.] 

Falcicaris tenuis Ortmann, 1893: 74; Plate 3, fig. 6. [Plankton-Expedition der Humboldt-Stiftung, Floridastrom 

JN 55 (37.9° N 59.1°W, 0-400 m); Südl. Aequatorialstrom JN 213 (5.7°S 26.5°W, 0-440 m) and 

218 (3.8°S 32.6°W, 0-400 m)] 

Hectarthropus compressus Spence Bate, 1888: 891; Plate 144, fig. 3. [Pacific, between Api and Cape York, 

surface] 

Hectarthropus exilis Spence Bate, 1888: 889; Plate 144, fig. 2. [off Basilan Strait, Philippine Islands, surface] 

Hectarthropus expansus Spence Bate, 1888: 893; Plate 144, figs 4-5. [off Basilan Strait, Philippine Islands, 

surface] 

Hectarthropus nikiformis Coutière, 1907a: 34; Fig. 11. [SAS Le Prince de Monaco stn 1715, Canary Islands, 

près de Ténériffe, 0-1000 m] 

Hectarthropus tenuis Spence Bate, 1888: 893; Plate 144, fig. 6. [North Atlantic, probably near the Cape 

Verde Islands, surface] 

Icotopus amplissimus Coutière, 1907a: 23; Fig. 7. [SAS Le Prince de Monaco stns 1715, Canary Islands, 

0-1000 m; 1760, Canary Islands, 0-3000 m; 2276, au voisinage des Baléares, 1-1700 m] 

Icotopus amproxima Coutière, 1907a: 27; Fig. 8. [SAS Le Prince de Monaco stn 1715, Canary Islands, 

0-1000 m] 

Icotopus arcurostris Spence Bate, 1888: 886; Plate 144, fig. 1. [off Cape Howe, Australia, surface] 

Kyptocaris oligodon Coutière, 1907a: 29; Fig. 9. [SAS Le Prince de Monaco stn 1874, Açores, 0-2000 m] 

Kyptocaris stylofrontalis Spence Bate, 1888: 690; Plate 121, fig. 1. [off Sibago, Samboangan, Philippine Islands, 

surface] 

Mesocaris recurva Ortmann, 1893: 82; Plate 5, fig. 6. [Plankton-Expedition der Humboldt-Stiftung, 

Brasilianische Küstenbank, JN 246 (0.4°S 46.6°W, 0-400 m)] 

Odontolophus serratus Spence Bate, 1888: 665. [off Malta, Mediterranean] 

Oligocaris bispinosa Ortmann, 1893: 85; Plate 7, fig. 2. [Plankton-Expedition der Humboldt-Stiftung, 

Irminger-See, JN 9 (60.2°N 22.7°W), JN 15 (60.3°N 27.0°W)] 

Oligocaris (?) brevirostris Coutière, 1907a: 32; Fig. 10. [SAS Le Prince de Monaco stn 1781, Açores, 

0-5000 m] 

Pandacaricyphus pandaliformis Coutière, 1907a: 21; Fig. 6. [SAS Le Prince de Monaco stn 1834, région de 

Açores, 0-1000 m] 

Posydon depressus Fabricius, 1798: 417. [in Oceano Indico] 

Procletes ellioti Spence Bate, 1888: 885. [off Waltair, on the coast of Coromandel] 

Retrocaris antarcticus Coutière, 1907c: 411; unnumbered figure. [Géorgie du Sud] 

Retrocaris contraria Ortmann, 1893: 83; Plate 5, fig. 7. [Plankton-Expedition der Humboldt-Stiftung, Südl. 

Aequatorialstrom JN 226 (2.4°S 36.4°W, 0 m); 235 (0.1°S 44.2°W, 0-400 m); Brasilianische Küstenbank, 

Pl. 112 (0.4°S 46.6°W, 207 m)] 

Retrocaris serrata Nobili, 1906c: 54; Plate 4, fig. 12. [Port d’Aden] 

Rhomaleocaris hamulus Spence Bate, 1888: 720: Plate 123, fig. 2. [on the passage from Api, New Hebrides, 

to Cape York, Pacific, surface]


Caridea Nomina Dubia 

Alpheus avarus Fabricius, 1798: 404. [in India orientali] 

Alpheus bispinosus Streets, 1871a: 242. [Isthmus of Panama] 

Alpheus flavescens Latreille, 1806: 53. [in Indiæ Orientalis Oceano] 

Alpheus galathea Miers, 1874: 5; Plate 4, fig. 4. [Port Essington] 

= Alpheus Galathea White, 1847a: 75. [Port Essington; nomen nudum] 

Alpheus hoplites Nobili, 1906b: 257. [îles Touamotou] 

Alpheus Jourdainii Guérin-Méneville, 1857: 51. [Type locality not indicated] 

Alpheus parvimanus Kingsley, 1878b: 195. [Panama] 

Alpheus Saulcyi Guérin-Méneville, 1855 [in Guérin-Méneville, 1855-1856]: xviii; Plate 2, fig. 8. [Cuba] 

Alpheus Savignyi Guérin-Méneville, 1857: 49. [nomen novum for Athanas nitescens Audouin, 1826 nec 

Leach, 1813 [in Leach, 1813-1814]] 

Alpheus sculptimanus Guérin-Méneville, 1857: 49. [De Janaon] 

Alpheus setimanus Cano, 1888: 180. [provenienza ?] 

Alpheus spinicaudus Lockington, 1878b: 476. [Port Escondido, Gulf of Cal.] 

Alpheus tamulus Fabricius, 1798: 405. [in Oceano Indico] 

Alpheus tenuimanus Lockington, 1878b: 473. [Port Escondido, Gulf of California] 

Alphous [sic] Gundlach Torralbas, 1917: 72; Figs 50-51. [Cuba] 

Anchistia antennata Hansen, 1925: 140. [Type locality not indicated] 

Anchistia brachiata Stimpson, 1860a: 39. [Portu ”Lloyd” ad insulas ”Bonin”] 

Anchistia Danae Stimpson, 1860a: 39. [Ins. ”Tahiti”, in corrallis] 

Astacus carinatus Meuschen, 1778: 86. [Type locality not indicated] 

Astacus ferratus Fabricius, 1793: 486. [in Oceano Norwegico] 

Astacus Gibbosus Pennant, 1812: 32. [Type locality not indicated] 

Astacus Squilla major Meuschen, 1778: 86. [Type locality not indicated] 

Astacus tettigonus Fabricius, 1775: 417. [Islandia] 

Betaeus vladivostokiensis Vinogradov, 1950: 198; Plate 10, fig. 31. [Zolotoi Bay, near Vladivostok] 

Caridina africana forma brevis Roth-Woltereck, 1984: 111. [Itimbiri] 

Caridina africana forma longa Roth-Woltereck, 1984: 111. [Mwerusee] 

Caridina africana nigerdeltae Roth-Woltereck, 1984: 117. [Nigerdelta] 

Caridina barombi Roth-Woltereck, 1984: 110. [Barombisee, Kamerun] 

Caridina powelli Roth-Woltereck, 1984: 110. [Nigerdelta] 

Caridina weberi var. celebensis Schenkel, 1902: 499. [Tabela Bach, System des Kalaena, Luwu] 

Cr.[yptophthalmus] ventricosus Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 4; Plate 7, figs 3A-B. 

[trovato nel Golfo di Taranto e sul faro di Messina] 

Crangon marginatus Fabricius, 1798: 410. [Isle de France] 

Crangon monopodium Bosc, 1802: 96; Plate 13, fig. 2. [la mer des Indes] 

Harpilius spinuliferus Miers, 1884b: 203. [from the interior of Tridacna without precise indication of locality] 

Hippolite affinis Owen, 1839: 90; Plate 27, fig. 4. [Monterey, California] 

Hippolyte gracilipes Randall, 1840: 142. [Sandwich Islands] 

Hippolyte ignobilis Kinahan, 1858c: 131. [Port Philip] 

Hippolyte Marioni Gourret, 1887a: 1033. [graviers et sables vaseux qui s’étendent depuis l’île de Riou 

jusqu’au delà de Planier, 108 m] 

Hymenodora parva Coutière, 1905c: 1114. [Princesse-Alice stns 1834 (37°28'N 25°53'30"W), 1000-0 m; 1851 

(46°15'N 7°09'W), 3000-0 m] 

Latreutes ceylonensis Pearson, 1905: 81; Plate 2, fig. 7. [10-12 miles north of Cheval Paar and 12 miles west 

of Vankali Church, 7.5-9 fms] 

Leander distans Heller, 1862a: 526. [Nicobaren] 

Leander hammondii Kingsley, 1883: 108; Plate 1, fig. 2. [Baker’s Island, North Pacific] 

Leander indicus Heller, 1862b: 413. [Java und Borneo] 

Leptalpheus petronii Ramos-Porto & de Souza, 1994: 22. [Ilha de Itamaraca, Pernambuco] 

Macrobrachium parasintangense Nguyên, 2004: 24. [Tri An Lake] 

Merhippolyte orientalis Spence Bate, 1888: 621. [5°41'0"S, 134°4'30"E, off New Guinea, 800fms] 

Odontocerus lutescens Leach, 1830b: 170. [shores of Guinea]


Opithiocheirus Chrysophthalmus Leach, 1830a: 172. [Atlantic ocean near the river Congo] 

Palaemon (Eupalaemon) cognatus Roux, 1927b: 322; Fig. 1. [Pionierbivak, Fl. Mamberamo] 

Palaemon (Parapalaemon) stresemanni Roux, 1918: 113; Figs 1-2. [Tjelukan Bawang, N.-O. Bali] 

Palaemon adriaticus O.G. Costa & A. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 7. [Golfo di 

Napoli] 

Palaemon Audouini Heller, 1861: 26. [im rothen Meere] 

Palaemon brevimanus Fabricius, 1798: 403. [in India orientali] 

Palaemon brevirostris Andrzeiowksi, 1839: 22. [Type locality not indicated] 

Palaemon coromandelianus Fabricius, 1798: 403. [in India orientali] 

Palæmon flavescens Olivier, 1811: 667. [Nouvelle-Hollande] 

Palæmon fluvialis Streets, 1871b: 227; Plate 2, figs 5-5a. [Tributary of the Coatzacoalcos River among the 

Cordilleras] 

Palaemon noctilucus Tilesius, 1819: 5; Plate 21a, fig. 2. [placed on the Official Index of Rejected and Invalid 


Palaemon parvulus O.G. Costa & A. Costa, 1871 [in O.G. Costa & A. Costa, 1838-1871]: 7. [Golfo di Napoli] 

Palaemon parvus Olivier, 1811: 666. [dans la Méditerranée] 

Palaemon rosalesi Rodríguez de la Cruz, 1965: 100; Plate 7. [Ciudad del Carmen, arroyo de agua salobre 

(cerca Av. Ultimo Paseo), Estado de Campeche] 

Palaemon tranquebaricus Fabricius, 1798: 403. [in India orientalis] 

Palaemonetes (Palaemonetes) granulosus Rodríguez de la Cruz, 1965: 89; Plate 6. [Laguna Las Palomas, 

Tampico, Tam.] 

Palaemonetes natalensis Stebbing, 1915: 78; Plate 19. [Cape Natal N. by E. 24 miles; depth 440 fms] 

Pandalus bipennus Poisson, 1947: 18. [l’ile Maurice] 

Pandalus quadridentata A. Milne-Edwards, 1883: Plate 19. [Travailleur dragage 54, 32°40'20''N 18°54'30''W, 

400 m] 

Plesionika capreensis Lo Bianco, 1903: 186. [vicinanze di Capri] 

Pontonia maculata Stimpson, 1860a: 38. [ad insulas ”Bonin”, in Tridacnis] 

Pontonia unidens Kingsley, 1880: 422; Plate 14, fig. 9. [Key West, Fla.] 

Prionorhynchus Cranchianus Leach, 1830b: 171. [Atlantic Ocean Lat. 24°13' Long. 18°51'15'', on the surface 

of the sea] 

Spirontocaris makrognathus Stebbing, 1921b: 19; Plate 4. [Durban waters] 

Synalpheus brachyceros Nobili, 1906b: 256. [Makatea] 

Typhlocaridina lingyunensis W.-X. Li & Luo, 2001: 72; Fig. 1. [cave in Shadong, Lingyun County, Guangxi] 

Usterocheirus Brachypocoilion Leach, 1830a: 174. [the Atlantic, adhering to fucus natans] 

Usterocheirus Macropocoilium Leach, 1830a: 173. [the Atlantic, adhering to fucus natans] 

Zuphanusa Smithiana Leach, 1830a: 175. [the Atlantic, adhering to fucus natans] 

Caridea Nomina Nuda 

Acanthephyra pleuracantha Coutière, 1914: 886. 

Alpheus albertei Kazmi & Kazmi, 1979: 154. 

Alpheus Amphitrite White, 1847a: 74. 

Alpheus Doris White, 1847a: 75. 

Alpheus Faxoni Coutière, 1900b: 356. 

Alpheus forceps White, 1847a: 75. 

Alpheus intermedius Stimpson, 1860b: 444. 

Alpheus isodactylus Afzal, Javed & Barkati, 1986: 339. 

Alpheus isthmalleator Hurt, Anker & Knowlton, 2009: 526. 

Alpheus manorensis Afzal, Javed & Barkati, 1986: 339. 

Alpheus miyakei Miya, 1974: 106. 

Alpheus ochrostriatus Karplus, Szlep & Tsurnamal, 1981: 5. 

Alpheus pseudoedwardsii Afzal, Javed & Barkati, 1986: 338. 

Alpheus purpurilenticularis Karplus & Ben Tuvia, 1979: 226. 

Alpheus Rathbuni Coutière, 1900b: 356.


Alpheus Rhode White, 1847a: 74. 

Alpheus rubromaculatus Karplus, Szlep & Tsurnamal, 1981: 5. 

Alpheus Triton White, 1847a: 74. 

Athanas arabicus Afzal, Javed & Barkati, 1986: 339. 

Brachycarpus dentatus Nobili, 1907: 4. 

Caricyphus acutus Coutière, 1905c: 1114. 

Caridion monctoni Citarella, 1993: 15. [off Buctouche (46°32'N – 64°32'30''W)] 

Crangon cymodocea Price, 1971: 166. 

Egeon tricuspidatum O.G. Costa, 1844c: 72. 

Gasconella parvula de Saint Laurent, 1985: 473. 

Hippolyte buruënsis Bleeker, 1856: 37. 

Hippolyte Hasseltii Bleeker, 1856: 64. 

Hippolyte lygdamis White, 1847a: 76. 

Hippolyte Macandreae Bell, 1847: 81. 

Hippolyte metis White, 1847a: 76. 

Hippolyte rubrosignata Wagner, 1885: 60. 

Macrobrachium honnaense Jayachandran, 2010: 47. 

Palæmon Abbotii White, 1847a: 78. 

Palæmon aciculatus White, 1847a: 79. 

Palaemon Affinis Hope, 1851: 17. 

Palaemon affinis O.G. Costa & A. Costa, 1840 [in O.G. Costa & A. Costa, 1838-1871]: 4. 

Palaemon Bipunctatus Hope, 1851: 17. 

Palæmon Brasiliensis White, 1847a: 79. 

Palæmon Colombicus White, 1847a: 78. 

Palæmon Creusa White, 1847a: 78. 

Palaemon cubanus Spamer & Bogan, 1992: 150. 

Palæmon Cydippe White, 1847a: 78. 

Palaemon Delaserii Hope, 1851: 17. 

Palæmon Electra White, 1847a: 78. 

Palæmon fluvialis Spence Bate, 1876: 378. 

Palæmon Glauce White, 1847a: 79. 

Palæmon Hypsa White, 1847a: 78. 

Palæmon Latreillei White, 1847a: 79. 

Palæmon Nicippe White, 1847a: 79. 

Palæmon Procles White, 1847a: 79. 

Palemon pusillum Rafinesque, 1814: 24. 

Palaemon Rathki Aurivillius, 1898b: 35. 

Palaemon recticornis Sherborn, 1933: 785. 

Palaemon seriatus Suckling, 1876: 270. 

Palaemon Siamensis von Martens, 1868: 30. 

Palaemon Splendens Hope, 1851: 17. 

Palaemon splendens O.G. Costa & A. Costa, 1840 [in O.G. Costa & A. Costa, 1838-1871]: 4. 

Palaemon walkeri O’Hara & Barmby, 2000: 43, 46. 

Palemon Inermis Roux, 1831: 15. 

Tizeuma Stimpsoni Perrier, 1886: 81.


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Hakuho-Maru during the KH-72-1 cruise.— Bulletin of the National Science Museum, Tokyo 22: 

263-281. 

Brandt, J.F., 1850. Bericht über die für die Reisebeschreibung des Herrn von Middendorf von J.F. Brandt 

bearbeiteten Krebsthiere aus den Abtheilungen der Brachyuren (Krabben), Anomuren und Makrouren 

(Krebse).— Bulletin de la Classe physico-mathématique de l’Académie Imperiale des Sciences 

de Saint-Pétersbourg 8: 234-238. 

Brandt, J.F., 1851. Krebse. In: Middendorff, A.T. von, Reise in den äussersten Norden und Osten Sibiriens 

während der Jahre 1843 und 1844 mit allerhöchster Genehmigung auf Veranstaltung der Kaiserlichen 

Akademie der Wissenschaften zu St. Petersburg ausgeführt und in Verbinding mit vielen 

Gelehrten herausgegeben, 2 (Theil 1): 77-148, Plates 5-6. St. Petersburg. 

Bray, D.M., 1976. A review of two Western Australian shrimp of the genus Palaemonetes, P. australis 

Dakin 1915 and P. atrinubes sp. nov. (Decapoda, Palaemonidae).— Records of the Western Australian 

Museum 4: 65-84. 

Bražnikov, V., 1903. Sur une nouveau genre et une nouvelle espèce de décapodes, famille Hippolytidae 

[in Russian].— Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.- Pétersbourg 

8: 44-45. 

Bražnikov, V., 1907. Matériaux pour servir à la connaissance de la faune des mers russes de l’Est rassemblés 

par le schooner ”Storož” en 1899-1902 [in Russian].— Mémoires de l’Académie Impériale 

des Sciences de St-Pétersbourg. Classe des Sciences Physiques et Mathématiques (8) 20: i-ii, 1-185, 

unumbered Plate. 

Brito Capello, F. de, 1867. Descripção de algunas especies novas ou pouco coniecidas de Crustaceos e 

Arachnidos de Portugal e possessões Portuguezas do ultramar.— Memorias da Academia Real de 

Lisboa 4: 1-17, Plates 1-2.


Bruce, A.J., 1965. Notes on Indo-Pacific Pontoniinae, X. Periclimenes cristimanus sp. nov., a new pontoniid 

shrimp from Singapore.— The Annals and Magazine of Natural History (13) 8: 487-493. 

Bruce, A.J., 1966a. Hymenopenaeus halli sp. nov., a new species of penaeid prawn from the South China 

Sea (Decapoda, Penaeidae).— Crustaceana 11: 216-224. 

Bruce, A.J., 1966b. Bathypalaemonella humilis sp.nov., a new species of shrimp from the South China Sea 

(Decapoda, Campylonotidae).— Crustaceana 11: 277-287. 

Bruce, A.J., 1966c. Notes on some Indo-Pacific Pontoniinae. XI. A re-examination of Philarius lophos Barnard, 

with the designation of a new genus, Ischnopontonia.— Bulletin of Marine Science 16: 584-598. 

Bruce, A.J., 1967a. The results of the re-examination of the type specimens of some pontoniid shrimps in 

the collection of the Muséum national d’Histoire naturelle, Paris.— Bulletin du Muséum national 

d’Histoire naturelle (2) 39: 564-572. 

Bruce, A.J., 1967b. Notes on some Indo-Pacific Pontoniinae III-IX. Descriptions of some new genera and 

species from the western Indian Ocean and the South China Sea.— Zoologische Verhandelingen 87: 

1-73. 

Bruce, A.J., 1968a. A report on some pontoniid shrimps from New Caledonia (Crustacea Decapoda Natantia).— 


Bruce, A.J., 1968b. Notes on some Indo-Pacific Pontoniinae. XII. The re-examination of the types of Pontonia? 

brevirostris Miers, 1884, with the designation of a new genus, Platypontonia (Decapoda, Natantia).— 


Bruce, A.J., 1969a. Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 

(Crustacea, Decapoda Natantia, Pontoniinae).— Zoologische Mededelingen 43: 253-278. 

Bruce, A.J., 1969b. Preliminary descriptions of ten new species of the genus Periclimenaeus Borradaile, 

1915 (Crustacea, Decapoda Natantia, Pontoniinae).— Zoologische Mededelingen 44: 159-176. 

Bruce, A.J., 1969c. Notes on some Indo-Pacific Pontoniinae. XIII. Propontonia pellucida gen. nov., sp. nov., 

a new pontoniid shrimp from the Amirante Islands.— Crustaceana 17: 141-150. 

Bruce, A.J., 1970a. Report on some commensal pontoniid shrimps (Crustacea: Palaemonidae) associated 

with an Indo-Pacific gorgonian host (Coelenterata: Gorgonacea).— Journal of Zoology, London 160: 

537-544. 

Bruce, A.J., 1970b. Notes on some Indo-Pacific Pontoniinae. XV. Hamopontonia corallicola gen. nov., sp. 

nov., a new pontoniid shrimp from Hong Kong.— Crustaceana 18: 37-48. 

Bruce, A.J., 1970c. Further preliminary descriptions of new species of the genus Periclimenaeus Borradaile, 

1915, (Crustacea, Decapoda Natantia, Pontoniinae).— Zoologische Mededelingen 44: 305- 

315. 

Bruce, A.J., 1971a. Periclimenes attenuatus sp. nov. (Crustacea Decapoda, Natantia, Pontoniinae), a new 

commensal shrimp from the Duke of York Islands.— Pacific Science 25: 533-544. 

Bruce, A.J., 1971b. Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. 

nov., a new pontoniid shrimp from Mahé, the Seychelle islands.— Crustaceana 20: 225-236. 

Bruce, A.J., 1971c. Notes on some Indo-Pacific Pontoniinae. XVI. Onycocaris seychellensis sp. nov., a new 

species of shrimp from Mahé.— Crustaceana 20: 208-218. 

Bruce, A.J., 1971d. Onycocaris zanzibarica sp. nov., a new pontoniid shrimp from East Africa.— Journal of 

Natural History 5: 293-298. 

Bruce, A.J., 1971e. On a new commensal shrimp Periclimenes hirsutus sp. nov. (Crustacea, Decapoda 

Natantia, Pontoniinae) from Fiji.— Pacific Science 25: 91-99. 

Bruce, A.J., 1972a. Pycnocaris chagoae gen. nov., sp. nov., a new shrimp from the Chagos Archipelago 

(Decapoda Natantia, Gnathophyllidae).— Crustaceana 23: 50-64. 

Bruce, A.J., 1972b. Notes on some Indo-Pacific Pontoniinae. XIX. Allopontonia iani gen nov., sp. nov., a 

new echinoid associate from Zanzibar.— Crustaceana 22: 1-12. 

Bruce, A.J., 1972c. A report on a small collection of pontoniid shrimps from Fiji, with the description of 

a new species of Coralliocaris Stimpson (Crustacea, Decapoda, Natantia, Pontoniinae).— Pacific Science 

26: 63-86. 

Bruce, A.J., 1972d. Notes on some Indo-Pacific Pontoniinae, XX. Pontonia sibogae sp. nov., a new species 

of Pontonia from eastern Australia and Indonesia (Decapoda Natantia, Palaemonidae).— Crustaceana 

23: 182-186.


Bruce, A.J., 1972e. Notes on some Indo-Pacific Pontoniinae, XXI. Typton bawii sp. nov., the first occurrence 

of the genus Typton Costa in the Indian Ocean (Decapoda Natantia, Palaemonidae).— Crustaceana 

23: 243-254. 

Bruce, A.J., 1973a. Gnathophylloides robustus sp. nov., a new commensal gnathophylloid shrimp from 

Western Australia, with the designation of a new genus Levicaris (Decapoda, Caridea).— Crustaceana 

24: 17-32. 

Bruce, A.J., 1973b. Notes on some Indo-Pacific Pontoniinae, XXIV. Dasycaris zanzibarica sp. nov. from the 

western Indian Ocean, with remarks on other species of Dasycaris Kemp, 1922 (Decapoda Natantia).— 


Bruce, A.J., 1973c. Notes on some Indo-Pacific Pontoniinae. XXII. Pliopontonia furtiva gen. nov., sp. nov., 

a new shrimp associated with a corallimorph zoantharian.— Crustaceana 24: 97-109. 

Bruce, A.J., 1973d. Notes on some Indo-Pacific Pontoniinae, XXIII. Tectopontonia maziwiae gen. nov., sp. 

nov., a new coral associate from Tangaynika (Decapoda, Palaemonidae).— Crustaceana 24: 169-180. 

Bruce, A.J., 1973e. Typton australis sp. nov., a new pontoniid shrimp from the Great Barrier Reef, Australia.— 

Records of the Australian Museum 28: 253-263. 

Bruce, A.J., 1974a. Coralliocaris viridis sp. nov., a preliminary note (Decapoda Natantia, Pontoniinae).— 


Bruce, A.J., 1974b. Periclimenes insolitus sp. nov. (Decapoda Natantia, Pontoniinae), a new commensal 

shrimp from Waikiki Beach, Oahu, Hawaii.— Crustaceana 26: 293-307. 

Bruce, A.J., 1975a. Notes on some Indo-Pacific Pontoniinae. XXVI. Neoanchistus cardiodytes gen. nov., sp. 

nov., a new mollusc-associated shrimp from Madagascar (Decapoda, Palaemonidae).— Crustaceana 

29: 149-165. 

Bruce, A.J., 1975b. Further observations on the Indo-West Pacific species of the genus Palaemonella Dana, 

1852 (Decapoda Natantia, Pontoniinae).— Crustaceana 29: 169-185. 

Bruce, A.J., 1975c. Observations upon some specimens of the genus Periclimenaeus Borradaile (Decapoda 

Natantia, Pontoniinae) originally described by G. Nobili.— Bulletin du Muséum national d’Histoire 

naturelle (3), Zoologie 258: 1557-1583. 

Bruce, A.J., 1975d. Periclimenes colemani sp. nov., a new shrimp associate of a rare sea urchin from Heron 

Island, Queensland (Decapoda Natantia, Pontoniinae).— Records of the Australian Museum 29: 

486-502. 

Bruce, A.J., 1975e. Notes on some Indo-Pacific Pontoniinae, XXV. Further observations upon Periclimenes 

noverca Kemp, 1922, with the designation of a new genus Zenopontonia, and some remarks upon 

Periclimenes parasiticus Borradaile (Decapoda Natantia, Palaemonidae).— Crustaceana 28: 275-285. 

Bruce, A.J., 1976a. Studies on Indo-West Pacific Stenopodidea, 1. Stenopus zanzibaricus sp. nov., a new 

species from East Africa.— Crustaceana 31: 90-102. 

Bruce, A.J., 1976b. Discias mvitae sp. nov., a new sponge associate from Kenya (Decapoda Natantia, Disciadidae).— 


Bruce, A.J., 1976c. Notes on some Indo-Pacific Pontoniinae, XXVII. Apopontonia falcirostris gen. nov., sp. 

nov., from Madagascar.— Crustaceana 31: 301-311. 

Bruce, A.J., 1976d. A report on a small collection of pontoniine shrimps from the Northern Indian 

Ocean.— Journal of the Marine Biological Association of India 16 [for 1974]: 437-454. 

Bruce, A.J., 1976e. A report on some pontoniid shrimps collected from the Seychelle Islands by the F.R.V. 

Manihine, 1972, with a review of the Seychelles pontoniid shrimp fauna.— Zoological Journal of the 

Linnean Society 59: 89-153. 

Bruce, A.J., 1976f. A report on a small collection of shrimps from the Kenya National Marine Parks at 

Malindi, with notes on selected species.— Zoologische Verhandelingen 145: 1-72. 

Bruce, A.J., 1977a. Pontoniine shrimps in the collections of the Australian Museum.— Records of the 


Bruce, A.J., 1977b. Periclimenes kororensis n. sp., an unusual shrimp associate of the fungiid coral, Heliofungia 

actiniformis.— Micronesica 13: 33-43. 

Bruce, A.J., 1977c. Notes on some Indo-Pacific Pontoniinae, XXIX. Epipontonia spongicola gen. nov., sp. 

nov., from Wasin Island, Kenya.— Crustaceana 32: 304-315.


Bruce, A.J., 1977d. Notes on some Indo-Pacific Pontoniinae, XXVIII. Typton wasini sp. nov., from Wasin 

Island, Kenya.— Crustaceana 32: 272-285. 

Bruce, A.J., 1978a. A report on a collection of pontoniine shrimps from Madagascar and adjacent seas.— 

Zoological Journal of the Linnean Society 62: 205-290. 

Bruce, A.J., 1978b. Paranchistus pycnodontae sp. nov., a new pontoniine shrimp associated with an ostreid 

bivalve host.— Memoirs of the Queensland Museum 18: 233-243, Plate 39. 

Bruce, A.J., 1978c. The re-examination of some pontoniine shrimp types first described by L.A. Borradaile 

(Decapoda, Palaemonidae).— Crustaceana 34: 251-268. 

Bruce, A.J., 1978d. Typton crosslandi sp. nov., a new pontoniine shrimp, from the Galapagos Islands.— 


Bruce, A.J., 1978e. Thor marguitae sp. nov., a new hippolytid shrimp from Heron Island, Australia.— 


Bruce, A.J., 1979a. Notes on some Indo-Pacific Pontoniinae, XXXI. Periclimenes magnificus sp. nov., a 

coelenterate associate from the Capricorn Islands (Decapoda, Palaemonidae).— Crustaceana Suppl. 

5: 195-208, Plate 1. 

Bruce, A.J., 1979b. Ctenopontonia cyphastreophila, a new genus and species of coral associated pontoniine 

shrimp from Eniwetok Atoll.— Bulletin of Marine Science 29: 423-435. 

Bruce, A.J., 1979c. Onycocaris furculata sp. nov., a new pontoniine shrimp from La Réunion.— Cahiers de 

l’Indo-Pacifique 1: 323-334. 

Bruce, A.J., 1979d. A report on a small collection of pontoniine shrimps from Eniwetok Atoll.— Crustaceana 

Supplement 5: 209-230. 

Bruce, A.J., 1979e. Onycocaris anomala sp. nov., a new pontoniine shrimp from the Northern Territory, 

Australia.— Records of the Australian Museum 32: 69-79. 

Bruce, A.J., 1980a. On some Pontoniine shrimps from Noumea, New Caledonia.— Cahiers de l’Indo- 

Pacifique 2: 1-39. 

Bruce, A.J., 1980b. Notes on some Indo-Pacific Pontoniinae, XXXIII. Periclimenaeus diplosomatis sp. nov., 

an ascidian associate from Heron Island, Australia.— Crustaceana 39: 39-51. 

Bruce, A.J., 1981a. Notes on some Indo-Pacific Pontoniinae, XXXVI. Pontonia ardeae sp. nov., a new bivalve 

associate from the Capricorn Islands (Decapoda, Natantia).— Crustaceana 40: 113-126. 

Bruce, A.J., 1981b. Onycocaridella prima, new genus, new species, a new pontoniine sponge-associate 

from the Capricorn Islands, Australia (Decapoda, Caridea, Pontoniinae).— Journal of Crustacean 

Biology 1: 241-250. 

Bruce, A.J., 1981c. Notes on some Indo-Pacific Pontoniinae, XXXVIII. Apopontonia dubia sp. nov., from a 

southern Queensland sponge host.— Crustaceana 41: 225-232. 

Bruce, A.J., 1981d. Results of the MUSORSTOM expeditions. I. — Philippines (18-28 March 1976). Decapod 

Crustacea: Pontoniinae.— Memoirs ORSTOM 91: 189-215. 

Bruce, A.J., 1982a. Notes on some Indo-Pacific Pontoniinae, XXXIX. Isopontonia platycheles gen. nov., sp. 

nov., from the Chesterfield Islands, New Caledonia (Decapoda, Caridea).— Crustaceana 42: 54-64. 

Bruce, A.J., 1982b. Notes on some Indo-Pacific Pontoniinae, XLI. Orthopontonia, a new genus proposed 

for Periclimenaeus ornatus Bruce.— Crustaceana 43: 163-176. 

Bruce, A.J., 1982c. The shrimps associated with Indo-West Pacific echinoderms, with the descriptions of 

a new species in the genus Periclimenes Costa, 1844 (Crustacea: Pontoniinae).— Memoirs of the 


Bruce, A.J., 1982d. Thorella cobourgi, new genus, new species, a hippolytid shrimp from the Northern 

Territory, Australia.— Journal of Crustacean Biology 2: 451-458. 

Bruce, A.J., 1983a. A second species of the pontoniine shrimp genus Dasella Lebour, D. ansoni sp. nov., 

from the Arafura Sea.— The Beagle, Records of the Northern Territory Museum of Arts and Sciences 

1: 21-29. 

Bruce, A.J., 1983b. Epipontonia anceps n. sp., a sponge-associated pontoniine shrimp from Heron Island, 

Queensland (Crustacea: Decapoda: Palaemonidae).— Records of the Australian Museum 35: 19-28. 

Bruce, A.J., 1983c. Expédition Rumphius II (1975). Crustacés parasites, commensaux, etc. (Th. Monod 

éd). IX. Crustacés Décapodes (1 ere partie: Natantia Pontoniinae).— Bulletin du Muséum national 

d’Histoire naturelle (4), section A, Zoologie, Biologie et Écologie animales 5: 871-902.


Bruce, A.J., 1983d. Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the 

Northern territory, Australia.— Journal of Crustacean Biology 3: 463-471. 

Bruce, A.J., 1983e. Lysmata debelius new species, a new hippolytid shrimp from the Philippines.— Revue 

française d’Aquariologie 9 [for 1982]: 115-120. 

Bruce, A.J., 1983f. Thor spinipes sp. nov., a new hippolytid shrimp from the Cobourg Peninsula, northern 

Australia.— The Beagle, Records of the Northern Territory Museum of Arts and Sciences 1: 1-10. 

Bruce, A.J., 1983g. Miropandalus hardingi, new genus, new species, a bizarre commensal pandalid shrimp 

from the Marshall Islands.— Journal of Crustacean Biology 3: 482-490. 

Bruce, A.J., 1984. Periclimenes dentidactylus, a new deep water pontoniine shrimp from Makassar Strait, 

Indonesia.— Marine Research in Indonesia 24: 7-17. 

Bruce, A.J., 1985a. Notes on some Indo-Pacific Pontoniinae, XLII. Miopontonia yongei gen. nov., sp. nov., 

from the Australian north west shelf (Decapoda, Caridea).— Crustaceana 48: 167-178. 

Bruce, A.J., 1985b. Decapod Crustacea: Pontoniinae (MUSORSTOM II).— Mémoires du Muséum national 

d’Histoire naturelle (A) Zoologie 133: 229-260. 

Bruce, A.J., 1986a. Two new species of Bathypalaemonella Balss (Crustacea, Decapoda, Campylonotidae) 

from the Australian Northwest shelf.— Zoologica Scripta 15: 251-264. 

Bruce, A.J., 1986b. Chacella, a new palaemonid shrimp genus proposed for Dasycaris kerstitchi Wicksten, 

1983 (Crustacea: Decapoda: Natantia).— Journal of Crustacean Biology 6: 485-490. 

Bruce, A.J., 1986c. Diapontonia maranulus, new genus, new species, a pontoniine shrimp associate of a 

deep-water echinoid.— Journal of Crustacean Biology 6: 125-133. 

Bruce, A.J., 1986d. Three new species of commensal shrimps from Port Essington, Arnhem Land, northern 

Australia (Crustacea: Decapoda: Palaemonidae).— The Beagle, Records of the Northern Territory 

Museum of Arts and Sciences 3: 143-166. 

Bruce, A.J., 1986e. Periclimenes milleri new species, a bathyal echinoid-associated pontoniine shrimp 

from the Bahamas.— Bulletin of Marine Science 39: 637-645. 

Bruce, A.J., 1986f. Notes on some Indo-Pacific Pontoniinae, XLIII. A new species of Typton from Ashmore 

Reef, Timor Sea (Decapoda, Palaemonidae).— Crustaceana 50: 278-286. 

Bruce, A.J., 1987a. Notes on some Indo-Pacific Pontoniinae, XLIV. Periclimenes darwiniensis sp. nov. from 

the Northern Territory, Australia (Decapoda, Caridea).— Crustaceana 52: 29-39. 

Bruce, A.J., 1987b. Periclimenes johnsoni sp.nov., a new species of shrimp from Singapore (Crustacea: 

Decapoda: Palaemonidae).— Indo-Malayan Zoology 4: 113-126. 

Bruce, A.J., 1987c. Onycocaridites anomodactylus, new genus, new species (Decapoda: Palaemonidae), a 

commensal shrimp from the Arafura Sea.— Journal of Crustacean Biology 7: 771-779. 

Bruce, A.J., 1987d. Typton nanus sp. nov., a new commensal shrimp (Crustacea: Decapoda: Palaemonidae) 

from the Australian north-west shelf.— The Beagle, Records of the Northern Territory Museum 

of Arts and Sciences 4: 49-56. 

Bruce, A.J., 1987e. A new species of alpheid shrimp, Alpheus bannerorum from northern Australia.— The 

Beagle, Records of the Northern Territory Museum of Arts and Sciences 4: 61-72. 

Bruce, A.J., 1988a. Two new palaemonid shrimps (Crustacea: Decapoda) from the Australian North 

West shelf.— Journal of Natural History 22: 1263-1276. 

Bruce, A.J., 1988b. A new palaemonid shrimp from the Zostera-beds of Moreton Bay, Queensland, Australia 

(Decapoda: Palaemonidae).— The Beagle, Records of the Northern Territory Museum of Arts 

and Sciences 5: 105-114. 

Bruce, A.J., 1988c. Exopontonia malleatrix, new genus, new species, a palaemonid shrimp from Ashmore 

Reef, Timor Sea.— Journal of Crustacean Biology 8: 122-130. 

Bruce, A.J., 1988d. A redescription of Periclimenaeus fimbriatus Borradaile, 1915, with the designation of 

a new genus (Crustacea: Decapoda: Palaemonidae).— Zoological Journal of the Linnean Society 94: 

219-232. 

Bruce, A.J., 1988e. Periclimenes tonga sp. nov., a commensal shrimp associated with a scyphozoan host 

from Tonga (Crustacea: Decapoda: Palaemonidae).— Micronesica 21: 23-32. 

Bruce, A.J., 1988f. Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the 

Coral Sea.— Pacific Science 42: 139-149.


Bruce, A.J., 1988g. Synalpheus dorae, a new commensal alpheid shrimp from the Australian northwest 

shelf.— Proceedings of the Biological Society of Washington 101: 843-852. 

Bruce, A.J., 1988h. A new crangonid shrimp, Pontocheras arafurae gen. et sp. nov., from the Arafura Sea.— 

Zoologica Scripta 17: 213-221. 

Bruce, A.J., 1989a. A report on some coral reef shrimps from the Philippine Islands.— Asian Marine Biology 

6: 173-192. 

Bruce, A.J., 1989b. Notes on some Indo-Pacific Pontoniinae, XLV. Conchodytes maculatus sp. nov., a new 

bivalve associate from the Australian northwest shelf.— Crustaceana 56: 182-192. 

Bruce, A.J., 1989c. Periclimenes gonioporae sp. nov. (Crustacea: Decapoda: Palaemonidae), a new coelenterate-associated 

shrimp.— The Beagle, Records of the Northern Territory Museum of Arts and 

Sciences 6: 149-156. 

Bruce, A.J., 1989d. Periclimenes poupini sp. nov., a new anemone-associated shrimp from deep-water 

traps (Crustacea, Decapoda, Palaemonidae).— Bulletin du Muséum national d’Histoire naturelle, 

4ème série, section A, Zoologie, Biologie et Écologie animales 11: 851-863. 

Bruce, A.J., 1990a. Two deep-sea shrimps new to the Australian fauna, Psathyrocaris hawaiiensis Rathbun 

(Pasiphaeidae) and Bresilia antipodarum, sp. nov. (Bresiliidae), with remarks on Encantada spinoculata 

Wicksten (Bresiliidae).— Invertebrate Taxonomy 4: 847-866. 

Bruce, A.J., 1990b. A second species of Bresilia, B. plumifera sp. nov., new to the Australian fauna (Crustacea: 

Decapoda: Bresiliidae).— The Beagle, Records of the Northern Territory Museum of Arts and 

Sciences 7: 1-8. 

Bruce, A.J., 1990c. Crustacea Decapoda: Deep-sea palaemonid shrimps from New Caledonian waters. 

In: Crosnier, A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 6.— Mémoires du Muséum 

national d’Histoire naturelle (A) Zoologie 145: 149-215. 

Bruce, A.J., 1990d. A new cnidarian-associated palaemonid shrimp from Port Essington, Cobourg Peninsula.— 

Indo-Malayan Zoology 6 [for 1989]: 229-243. 

Bruce, A.J., 1990e. Periclimenes franklini sp. nov., a new deep-sea shrimp from the Coral Sea (Crustacea: 

Decapoda: Palaemonidae).— The Beagle, Records of the Northern Territory Museum of Arts and 


Bruce, A.J., 1990f. Additions to the marine shrimp fauna of Hong Kong. In: Morton, B. (ed.), Proceedings 

of the second international marine biological workshop: The marine fauna and flora of Hong Kong 

and southern China, Hong Kong, 1986: 611-648. Hong Kong University Press, Hong Kong. 

Bruce, A.J., 1990g. Redescriptions of five Hong Kong carideans first described by William Stimpson, 

1860. In: Morton, B. (ed.), Proceedings of the second international marine biological workshop: The 

marine fauna and flora of Hong Kong and southern China, Hong Kong, 1986: 569-610. Hong Kong 

University Press, Hong Kong. 

Bruce, A.J., 1990h. Crustacea Decapoda: Gelastreutes crosnieri gen. nov., sp. nov. (Hippolytidae) from 

New Caledonia. In: Crosnier, A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 6.— Mémoires 

du Muséum national d’Histoire naturelle (A) Zoologie 145: 137-147. 

Bruce, A.J., 1990i. Leontocaris amplectipes sp. nov. (Hippolytidae), a new deep-water shrimp from southern 

Australia.— Memoirs of the Museum of Victoria 51: 121-130. 

Bruce, A.J., 1991a. Crustacea Decapoda: Further deep-sea palaemonoid shrimps from New Caledonian 

waters. In: Crosnier, A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 9.— Mémoires du Muséum 


Bruce, A.J., 1991b. Shallow-water palaemonoid shrimps from New Caledonia (Crustacea: Decapoda). 

In: Richer de Forges, B. (ed.), Le benthos des Fonds Meubles des Lagons de Nouvelle-Calédonie, 1: 

221-279. ORSTOM, Paris. 

Bruce, A.J., 1991c. Notopontonia platycheles, new genus, new species (Decapoda: Pontoniinae) from South 

Australia, with remarks on Pontonia pinnophylax (Otto), the type species of Pontonia Latreille.— 


Bruce, A.J., 1991d. The ”African” shrimp genus Potamalpheops in Australia, with the description of 

P.hanleyi, new species (Decapoda: Alpheidae).— Journal of Crustacean Biology 11: 629-638. 

Bruce, A.J., 1992a. Pycnisia raptor, a new genus and species of predatory troglobitic shrimp (Crustacea: 

Decapoda: Atyidae) from northern Australia.— Invertebrate Taxonomy 6: 553-566.


Bruce, A.J., 1992b. A re-description of Macrobrachium handschini (Roux, 1933) (Crustacea: Decapoda: Palaemonidae).— 

Hydrobiologia 231: 131-139. 

Bruce, A.J., 1992c. Two new species of Periclimenes (Crustacea: Decapoda: Palaemonidae) from Lizard 

Island, Queensland, with notes on related taxa.— Records of the Australian Museum 44: 

45-84. 

Bruce, A.J., 1992d. Designation of two new pontoniine shrimp genera (Decapoda: Palaemonidae).— 

Journal of Natural History 26: 1273-1282. 

Bruce, A.J., 1992e. Additions to the marine caridean fauna of Hong Kong, with a description of a new 

species of Onycocaris (Crustacea: Decapoda: Palaemonidae) from Tuvalu. In: Morton, B. (ed.), The 

marine fauna and flora of Hong Kong and southern China III. Proceedings of the Fourth International 

Marine Biological Workshop. The Marine Fauna and Flora of Hong Kong and southern China: 

329-343. Hong Kong University Press, Hong Kong. 

Bruce, A.J., 1993a. Kakaducaris glabra gen. nov., sp. nov., a new freshwater shrimp from the Kakadu National 

Park, Northern Territory, Australia, Crustacea: Decapoda: Palaemonidae with the designation 

of a new subfamily Kakaducaridinae.— Hydrobiologia 268: 27-44. 

Bruce, A.J., 1993b. Pontoniine shrimps from the Zoological Museum, Copenhagen.— Journal of Natural 

History 28: 829-840. 

Bruce, A.J., 1993c. Potamalpheops darwiniensis (Crustacea: Decapoda: Alpheidae), the third Indo-West 

Pacific species.— Proceedings of the Biological Society of Washington 106: 698-704. 

Bruce, A.J., 1994a. Leander plumosus sp. nov., a new palaemonine shrimp (Crustacea: Palaemonidae) 

from the Maldive Islands.— The Beagle, Records of the Museums and Art Galleries of the Northern 

Territory 11: 39-51. 

Bruce, A.J., 1994b. Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid 

shrimps of the brevirostris species group.— The Beagle, Records of the Museums and Art Galleries 

of the Northern Territory 11: 15-28. 

Bruce, A.J., 1994c. Shrimps from Flat-Top Bank, Timor Sea (Crustacea: Decapoda: Caridea).— Raffles 

Bulletin of Zoology 42: 743-756. 

Bruce, A.J., 1995. A synopsis of the Indo-West Pacific genera of the Pontoniinae (Crustacea: Decapoda: 

Pontoniinae).— Theses Zoologicae 25: 1-172 [imprint 1994, published 16st March 1995]. 

Bruce, A.J., 1996. Crustacea Decapoda: Palaemonoid shrimps from the Indo-West Pacific region mainly 

from New Caledonia. In: Crosnier, A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 15.— Mémoires 

du Muséum national d’Histoire naturelle 168: 197-267. 

Bruce, A.J., 1997a. A new pontoniine shrimp genus (Crustacea: decapoda) from the Yemen, with a note 

on other species.— Journal of Natural History 31: 1213-1222. 

Bruce, A.J., 1997b. A new genus of hippolytid shrimp (Crustacea: Decapoda: Hippolytidae) for Thor 

maldivensis Borradaile.— Memoirs of the Queensland Museum 42: 13-23. 

Bruce, A.J., 1998a. A new species of the genus Brachycarpus (Decapoda, Caridea, Palaemonidae) from 

New Caledonia.— Zoosystema 20: 157-165. 

Bruce, A.J., 1998b. A second species of the genus Balssia Kemp, 1922 (Crustacea, Decapoda, Pontoniinae).— 

Zoosystema 20: 603-611. 

Bruce, A.J., 1998c. Pontoniine shrimps from Moreton Bay, Queensland (Crustacea: Decapoda: Pontoniinae).— 

Memoirs of the Queensland Museum 42: 387-398. 

Bruce, A.J., 1998d. A new name, Thinora, proposed for the preoccupied name Thorina Bruce (Crustacea: 

Decapoda: Hippolytidae).— Memoirs of the Queensland Museum 42: 398. 

Bruce, A.J., 1999. Alpheus soror, a new snapping shrimp cryptospecies from Sri Lanka (Crustacea: Decapoda: 

Alpheidae).— Raffles Bulletin of Zoology 47: 453-463. 

Bruce, A.J., 2000. Typton manningi and T. capricorniae, new species, new pontoniine shrimps from Northern 

Queensland, with a review of the Indo-West Pacific species of Typton Costa (Decapoda: Palaemonidae).— 

Journal of Crustacean Biology 20 (Special number 2): 87-100. 

Bruce, A.J., 2002a. Leander manningi, a new palaemonine shrimp from Western Australia (Crustacea, 

Decapoda, Palaemonidae), with a review of the Indo-West Pacific species of the genus Leander E. 

Desmarest, 1849.— Records of the Western Australian Museum 21: 71-81.


Bruce, A.J., 2002b. Notes on some Indo-Pacific Pontoniinae, XLVI. Palaemonella foresti sp.nov., a new 

pontoniine shrimp from Western Australia (Decapoda, Palaemonidae), with a review of the Indo- 

West Pacific species of the genus Palaemonella Dana, 1852.— Crustaceana 75: 277-298. 

Bruce, A.J., 2002c. A new species of Palaemonella (Crustacea: Decapoda: Pontoniinae) from East Africa.— 

The Beagle, Records of the Museums and Art Galleries of the Northern Territory 18: 15-18. 

Bruce, A.J., 2003a. Periclimenes species (Crustacea: Decapoda: Pontoniinae) from far North Queensland.— 


Bruce, A.J., 2003b. A new species of Dactylonia Fransen (Crustacea: Decapoda: Pontoniinae) from East 

Africa.— Cahiers de Biologie Marine 44: 299-306. 

Bruce, A.J., 2003c. Vir colemani sp. nov., a new commensal pontoniine shrimp (Crustacea: decapoda: 

Palaemonidae) from Papua New Guinea.— The Beagle, Records of the Museums and Art Galleries 


Bruce, A.J., 2004a. Balssia antipodarum sp. nov., the first occurrence of the genus Balssia Kemp in the Indo- 

West Pacific region (Crustacea: Decapoda: Pontoniinae).— Cahiers de Biologie Marine 45: 365-372. 

Bruce, A.J., 2004b. A partial revision of the genus Periclimenes Costa, 1844 (Crustacea: Decapoda: Palaemonidae).— 

Zootaxa 582: 1-26. 

Bruce, A.J., 2004c. A new pontoniine shrimp from Tahiti, French Polynesia (Crustacea, Decapoda, Palaemonidae).— 


Bruce, A.J., 2005a. Bresiliid shrimps from the Red Sea (Crustacea: Decapoda: Caridea) with the description 

of a new species.— Proceedings of the Biological Society of Washington 118: 176-182. 

Bruce, A.J., 2005b. Pontoniine shrimps from Papua New Guinea, with designation of two new genera, 

Cainonia and Colemonia (Crustacea: Decapoda: Palaemonidae).— Memoirs of the Queensland Museum 

51: 333-383. 

Bruce, A.J., 2005c. New species of Periclimenaeus Borradaile (Crustacea: Decapoda: Pontoniinae) from 

Ashmore reef, North Western Australia.— Records of the Western Australian Museum 22: 325-342. 

Bruce, A.J., 2005d. Pontoniine shrimps from the 2003 NORFANZ Expedition, 10 May-16 June (Crustacea: 

Decapoda: Palaemonidae).— Zootaxa 981: 1-20. 

Bruce, A.J., 2005e. A new genus Pontoniopsides, for Pontoniopsis paulae Gore, 1981 (Crustacea: Decapoda: 

Pontoniinae).— Zootaxa 826: 1-4. 

Bruce, A.J., 2006a. Pontoniine shrimps (Decapoda: Palaemonidae) from the island of Socotra, with descriptions 

of new species of Dactylonia Fransen, 2002 and Periclimenoides Bruce, 1990.— Zootaxa 

1137: 1-36. 

Bruce, A.J., 2006b. A new species of the pontoniine shrimp genus Harpilius Dana, 1852, H. spinifer (Crustacea: 

Decapoda: Palaemonidae), from New Caledonia.— Cahiers de Biologie Marine 47: 271-280. 

Bruce, A.J., 2006c. Periclimenes jackhintoni sp. nov. (Crustacea: Decapoda: Palaemonidae), a new pontoniine 

shrimp and crinoid associate from Tonga.— The Beagle, Records of the Museums and Art Galleries 


Bruce, A.J., 2006d. A new genus Leptomenes, for the pontoniine shrimp Periclimenes dolichosternum Okuno 

& Mitsuhashi, 2003 (Crustacea: Decapoda: Palaemonidae).— Cahiers de Biologie Marine 47: 

223-225. 

Bruce, A.J., 2006e. Periclimenaeus fawatu spec. nov. (Crustacea: Decapoda: Pontoniinae), from Zanzibar.— 

Zoologische Mededelingen 84: 33-43. 

Bruce, A.J., 2006f. Periclimenaeus nielbrucei sp. nov. (Crustacea: Decapoda: Pontoniinae), a new sponge 

associate from the Capricorn Islands, Queensland, with notes on related Periclimenaeus species.— 

Zootaxa 1224: 1-22. 

Bruce, A.J., 2006g. An unusual new Periclimenes (Crustacea, Decapoda, Palaemonidae) from New Caledonia.— 


Bruce, A.J., 2006h. Periclimenes manihine sp. nov., a new pontoniine shrimp of the P.alcocki species group 

(Crustacea: Decapoda: Palaemonidae).— Zootaxa 1309: 45-54. 

Bruce, A.J., 2007a. Re-examination of Borradaile’s Urocaris longicaudata specimens from the 1905 J.S. 

Gardiner Collection (Crustacea: Decapoda: Pontoniinae).— Zootaxa 1644: 51-57. 

Bruce, A.J., 2007b. Leptomenaeus gen. nov., a new name for Leptomenes Bruce, 2006 (Crustacea: Decapoda: 

Pontoniinae).— Cahiers de Biologie Marine 48: 411-412.


Bruce, A.J., 2007c. A re-definition of the genus Periclimenes Costa, 1844 and the designation of a new 

genus Margitonia (Crustacea: Decapoda: Pontoniinae).— Cahiers de Biologie Marine 48: 403-406. 

Bruce, A.J., 2007d. Palaemonoid shrimps from the Dampier Archipelago (Crustacea: Decapoda), with a 

review of the Western Australian pontoniine shrimp fauna.— Records of the Western Australian 

Museum Supplement 73: 97-129. 

Bruce, A.J., 2007e. Periclimenes sarkanae sp. nov. a new pontoniine shrimp from Moreton Bay, Queensland 

(Crustacea: Decapoda: Palaemonidae).— Zootaxa 1393: 61-68. 

Bruce, A.J., 2008a. Periclimenes aqabai sp. nov., a further new species of the aesopius species group 

(Caridea: Pontoniinae) from the Red Sea.— Zootaxa 1682: 27-32. 

Bruce, A.J., 2008b. A new species of Palaemonella Dana, 1852, (Crustacea: Decapoda; Pontoniinae) from 

the Red Sea.— Zootaxa 1844: 63-68. 

Bruce, A.J., 2008c. Palaemonoid shrimps from the Australian north west shelf.— Zootaxa 1815: 1-24. 

Bruce, A.J., 2008d. A new pontoniine shrimp from the Great Barrier Reef (Crustacea. Decapoda. Palaemonidae).— 


Bruce, A.J., 2008e. Phycomenes zostericola gen. nov., sp. nov., a new pontoniine shrimp (Crustacea: Decapoda: 

Palaemonidae) from Moreton Bay, Queensland.— Memoirs of the Queensland Museum 54: 

219-232. 

Bruce, A.J., 2009a. Comments on the generic position of Typton australis Bruce, 1973, and some related 

taxa (Crustacea: Decapoda: Pontoniinae).— Zootaxa 2076: 60-62. 

Bruce, A.J., 2009b. Plesiomenaeus poorei gen. nov., sp. nov., (Crustacea: Decapoda: Pontoniinae) from Zanzibar.— 

Memoirs of Museum Victoria 66: 25-34. 

Bruce, A.J., 2009c. A re-description of Typton spongicola Costa, 1844, the type species of the genus Typton 

Costa, 1844 (Crustacea: Decapoda: Pontoniinae).— Cahiers de Biologie Marine 50: 382-394. 

Bruce, A.J., 2010a. Allopontonia alastairi sp. nov., a second species of the genus Allopontonia Bruce, 1972 

(Crustacea: Decapoda: Pontoniinae), from the Australian Northwest Shelf. In: De Grave, S. & 

C.H.J.M. Fransen (eds.), Contributions to shrimp taxonomy.— Zootaxa 2372: 33-36. 

Bruce, A.J., 2010b. Ancylomenes kuboi and A.okunoi spp. nov. (Decapoda: Pontoniinae), from the Australian 

Northwest Shelf, Vietnam and the Philippines. In: De Grave, S. & C.H.J.M. Fransen (eds.), Contributions 


Bruce, A.J., 2010c. A revision of the systematic position of Periclimenaeus spinimanus Bruce, 1969 (Crustacea: 

Decapoda: Pontoniinae) and the designation of Anisomenaeus gen. nov. In: De Grave, S. & 


Bruce, A.J., 2010d. Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: 

Decapoda: Pontoniinae) from Oahu, Hawai’ian Islands. In: De Grave, S. & C.H.J.M. Fransen 

(eds.), Contributions to shrimp taxonomy.— Zootaxa 2372: 405-414. 

Bruce, A.J., 2010e. Palaemonella dijonesae sp. nov. (Crustacea: Decapoda: Pontoniinae) from Western Australia. 

In: De Grave, S. & C.H.J.M. Fransen (eds.), Contributions to shrimp taxonomy.— Zootaxa 

2372: 151-156. 

Bruce, A.J., 2010f. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the CReefs 2009 

Heron Island expedition.— Zootaxa 2604: 20-36. 

Bruce, A.J., 2010g. Periclimenaeus devaneyi sp. nov., from Oahu, Hawai’i (Crustacea: Decapoda: Pontoniinae). 


2372: 379-388. 

Bruce, A.J., 2010h. Periclimenes nevillei sp. nov. (Crustacea: Decapoda: Pontoniinae) from Vanuatu.— 

Zootaxa 2510: 45-54. 

Bruce, A.J., 2010i. Pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the CReefs 2009 Heron 

Island Expedition, with a review of the Heron island pontoniine fauna.— Zootaxa 2541: 50-68. 

Bruce, A.J., 2010j. Typtonoides nieli gen. nov., sp. nov., a new pontoniine shrimp (Crustacea: Decapoda: 

Palaemonidae) from the Chesterfield Island.— The Beagle, Records of the Museums and Art Galleries 


Bruce, A.J., 2010k. Lebbeus rubrodentatus sp. nov. (Crustacea: Caridea: Hippolytidae) from the Australian 

North West Shelf.— The Beagle, Records of the Museums and Art Galleries of the Northern Territory 

26: 75-78.


Bruce, A.J., 2011a. Notes on some Indo-Pacific Pontoniinae, XLVIII. Onycocaris nieli sp. nov., a new pontoniine 

shrimp from Heron Island, Queensland. — Crustaceana 84: 319-330. 

Bruce, A.J., 2011b. Notes on some Indo-Pacific Pontoniinae, XLIX. Onycocaris balsii sp. nov., from northern 

Australia, with the designation of O. fujinoi sp. nov. Crustaceana 84: 477-490. 

Bruce, A.J. & K. Baba, 1973. Spongiocaris, a new genus of stenopodidean shrimp from New Zealand and 

South African waters, with a description of two new species (Decapoda Natantia, Stenopodidea).— 


Bruce, A.J. & R.T. Bauer, 1997. A new pontoniine shrimp (Crustacea: Decapoda: Palaemonidae) from the 

Ryukyu Islands, Japan.— Journal of Natural History 31: 1709-1721. 

Bruce, A.J. & F.A.Jr. Chace, 1986. Paralebbeus zotheculatus, n. gen, n sp., a new hippolytid shrimp from the 

Australian northwest shelf.— Proceedings of the Biological Society of Washington 99: 237-247. 

Bruce, A.J. & K.E. Coombes, 1995. The palaemonoid shrimp fauna (Crustacea: Decapoda: Caridea) of 

the Cobourg Peninsula, Northern Territory.— The Beagle, Records of the Museums and Art Galleries 


Bruce, A.J. & K.E. Coombes, 1997. An annotated check-list of the caridean shrimps (Crustacea: decapoda) 

of Darwin Harbour, with descriptions of three new species of Periclimenes [Palaemonidae: Pontoniinae]. 

In: Hanley, J.R., G. Caswell, D. Megirian & H.K. Larson (eds.), Proceedings of the sixth 

international marine biological workshop. The marine fauna and flora of Darwin Harbour, Northern 

Territory, Australia: 301-337. Museums and Arts Galleries of the Northern Territory and the 

Australian Marine Sciences Association, Darwin. 

Bruce, A.J. & P.J.F. Davie, 2006. A new anchialine shrimp of the genus Procaris from Christmas Island: the 

first occurrence of the Procarididae in the Indian Ocean (Crustacea: Decapoda: Caridea).— Zootaxa 

1238: 23-33. 

Bruce, A.J. & T.M. Iliffe, 1991. Hamalpheus acanthops, new genus, new species, a stygiophilic alpheid 

shrimp from a Samoan lava tube.— Journal of Crustacean Biology 11: 583-593. 

Bruce, A.J. & T.M. Iliffe, 1992. Potamalheops pininsulae sp. nov., a new stygiophilic shrimp from New 

caledonia (Crustacea: Decapoda: Alpheidae).— Stygologia 7: 231-242. 

Bruce, A.J. & J. Okuno, 2006. Periclimenes dardanicola n. sp., a new species of hermit crab associated 

shrimp (Crustacea, Decapoda, Palaemonidae) from the western Pacific.— Zoosystema 28: 367-377. 

Bruce, A.J. & J. Okuno, 2010. Designation of a new genus Lipkemenes, with supplementary description 

and range extension of its type species, L.lanipes (Kemp, 1922) (Decapoda, Palaemonidae). In: Fransen, 

C.H.J.M., S. De Grave & P.K.L. Ng (eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis 

Memorial Volume.— Crustaceana Monographs, 14: 159-171. Brill, Leiden. 

Bruce, A.J., J. Okuno & X. Li, 2005. Manipontonia gen. nov., a new pontoniine shrimp genus for Periclimenes 

psamathe (De Man) (Crustacea: Decapoda: Palaemonidae).— Zootaxa 926: 1-11. 

Bruce, A.J. & J.W. Short, 1993. Leptopalaemon gagadjui gen. nov., sp. nov., a new freshwater palaemonid 

shrimp from Arnhem land, and a re-evaluation of Palaemonetes holthuisi Strenth, with the desgnation 

of a new genus, Calathaemon.— Hydrobiologia 257: 73-94. 

Bruce, A.J. & A. Svoboda, 1983. Observations upon some pontoniine shrimps from Aqaba, Jordan.— 

Zoologische Verhandelingen 205: 1-44. 

Bruce, A.J. & A. Svoboda, 1984. A report on a small collection of coelenterate-associated pontoniine 

shrimps from Cebu, Philippines Islands.— Asian Marine Biology 1: 87-99. 

Bruce, A.J. & D.L. Zmarzly, 1983. Periclimenes pilipes, new species, a crinoid associate form Enewetak 

Atoll, Marshall Islands (Crustacea: Decapoda: Pontoniinae).— Journal of Crustacean Biology 3: 644- 

654. 

Brünnich, M.T., 1768. Ichthyologia Massiliensis, sistens piscium descriptions eorumque apud incolas 

nomina. Accedunt spolia maris Adriatica: 1-110. Hafniae et Lipsiae. 

Bruin, G.H.P. de, 1965. Penaeid prawns of Ceylon (Crustacea, Decapoda, Penaeidae).— Zoologische 

Mededelingen 41: 73-104. 

Brullé, A., 1839. Crustacés. In: Barker-Webb, P. & S. Berthelot, Histoire naturelle des Iles Canaries: 15-18, 

Planche unique. Béthune, Paris. 

Bryazgin, V.F., 1982. On two species of shrimps from the genus Bythocaris in the Arctic Basin [in Russian].— 

Zoolo gicheskii Zhurnal 61: 603-605.


Buchholz, R., 1874. Crustaceen. In: Hartlaub, G. & M. Lindeman, Die zweite deutsche Nordpolarfahrt in 

den Jahren 1869 und 1870, unter Fuhrung des Kapitan Karl Koldewey: 262-399, Plates 1-15. 

Buldovsky, A.T., 1933. New data on fana of the Decapoda of the Lake basin [in Russian].— Biological 

Bulletin of the Far Eastern Branch of the Russian Academy of Science 1-3: 43-56. 

Burkenroad, M.D., 1934a. Littoral Penaeidea chiefly from the Bingham Oceanographic Collection, with 

a revision of Penaeopsis and descriptions of two new genera and eleven new American species.— 

Bulletin of the Bingham Oceanographic Collection 4: 1-109. 

Burkenroad, M.D., 1934b. The Penaeidea of Lousiana with a discussion of their world relationships.— 

Bulletin of the American Museum of Natural History 68: 61-143. 

Burkenroad, M.D., 1936a. The Aristaeinae, Solenocerinae and pelagic Penaeinae of the Bingham Oceanographic 

collection.— Bulletin of the Bingham Oceanographic Collection 5: 1-151. 

Burkenroad, M.D., 1936b. A new species of Penaeus from the American Atlantic.— Anais da Academia 

Brasileira de Ciências 8: 315-318. 

Burkenroad, M.D., 1937. The Templeton Crocker Expedition. XII. Sergestidae (Crustacea Decapoda) 

from the Lower Californian region, with descriptions of two new species and some remarks on the 

Organs of Pesta in Sergestes.— Zoologica, New York 22: 315-329. 

Burkenroad, M.D., 1938. The Templeton Crocker Expedition. XIII. Penaeidae from the region of Lower 

California and Clarion Island, with descriptions of four new species.— Zoologica, New York 23: 

55-91. 

Burkenroad, M.D., 1939. Further observations on Penaeidae of the northern Gulf of Mexico.— Bulletin 

of the Bingham Oceanographic Collection 6 (6): 1-62. 

Burkenroad, M.D., 1940. Preliminary descriptions of twenty-one new species of pelagic Penaeidae 

(Crustacea Decapoda) from the Danish Oceanographical Expeditions.— The Annals and Magazine 

of Natural History (11) 6: 35-54. 

Burkenroad, M.D., 1945. A new sergestid shrimp (Peisos petrunkevitchii, n. gen., n. sp.), with remarks on 

its relationships.— Transactions of the Connecticut Academy of Arts and Sciences 36: 553-591, 

Plates 1-2. 

Burkenroad, M.D., 1946. Status of the name Sicyonia H.M.E., with a note on S.typica (Boeck) and descriptions 

of two new species.— Arkiv för Zoologi 37A (9): 1-10. 

Burkenroad, M.D., 1959. Decapoda Macrura I. Penaeidae. Mission Robert Ph. Dollfus en Egypte XXV.— 

Mission Robert Ph. Dollfus en Égypt, Résultats Scientifique 3: 67-92. 

Burkenroad, M.D., 1963. The evolution of the Eucarida (Crustacea, Eumalacostraca), in relation to the 

fossil record.— Tulane Studies in Geology 2: 3-17. 

Burukovsky, R.N., 1966. A new species of shrimps of the genus Bythocaris, and some problems of zoogeography 

of the genus [in Russian].— Zoologicheskii Zhurnal 45: 536-542. 

Burukovsky, R.N., 1972. Some problems of the systematics and distribution of shrimps of the genus 

Penaeus [in Russian].— Trudy AtlantNIRO, Kaliningrad 2: 3-21. 

Burukovsky, R.N., 1974. Keys for the identification of shrimps, spiny lobsters and lobsters [in Russian]: 

1-126. Pishchevaya Promyshlennost’, Moscow. 

Burukovsky, R.N., 1975. Aristeus crosnieri sp. n. (Decapoda, Aristeinae) from the south-east Atlantic [in 

Russian].— Zoologicheskii Zhurnal 54: 779-781. 

Burukovsky, R.N., 1976. A new species of shrimp Pasiphaea grandicula sp. n. (Decapoda, Crustacea) and 

a short outline of the genus species [in Russian].— Byulleten’ Moskovskogo Obshchestva Ispytatelei 

Prirody Otdel Biologicheskii 4: 17-28. 

Burukovsky, R.N., 1977. A new species of the family Pasiphaeidae (Crustacea, Decapoda) [in Russian].— 

Zoologicheskii Zhurnal 56: 473-475. 

Burukovsky, R.N., 1978. About two species of shrimps (Decapoda, Caridea) from the south-west Atlantic 

[in Russian].— Zoologicheskii Zhurnal 57: 1729-1732. 

Burukovsky, R.N., 1986. A new shrimp species from the genus Heterocarpus (Crustacea: Decapoda: Pandalidae) 

and a brief review of species of the genus [in Russian].— Byulleten’ Moskovskogo Obshchestva 

Ispytatelei Prirody, Otdel Biologicheskii 91: 62-73. 

Burukovsky, R.N., 1987. On the taxonomic status of two bathypelagic species of shrimps (Crustacea, 

Decapoda, Pasiphaeidae) [in Russian].— Zoologicheskii Zhurnal 66: 37-41.


Burukovsky, R.N., 1988. New representatives of the family Bresiliidae (Crustacea, Decapoda) from the 

north-east Atlantic [in Russian].— Zoologicheskii Zhurnal 67: 456-460. 

Burukovsky, R.N., 1990. Shrimps from the Sala-y-Gomez and Nazca Ridges [in Russian].— Trudy Instituta 

Okeanologii Akademii Nauk SSSR 124: 187-217. 

Burukovsky, R.N., 1991a. Shrimps of the family Nematocarcinidae (Decapoda, Caridea) from the western 

part of the Indian Ocean [in Russian].— Zoologicheskii Zhurnal 70: 39-46. 

Burukovsky, R.N., 1991b. New and rare species of shrimps from the south-west part of the Indian Ocean 


Burukovsky, R.N., 1992. New species of the genus Plesionika (Crustacea, Decapoda, Pandalidae) from 

underwater bight in the Pacific Ocean [in Russian].— Zoologicheskii Zhurnal 71: 145-147. 

Burukovsky, R.N., 1993a. Shrimps of the Saya de Malha Bank (the Indian Ocean) [in Russian].— Zoologicheskii 

Zhurnal 72: 20-28. 

Burukovsky, R.N., 1993b. Shrimps of the genus Pasiphaea (Crustacea, Decapoda, Pasiphaeidae) from the 

western part of the Indian Ocean [in Russian].— Byulleten’ Moskovskogo Obshchestva Ispytatelei 

Prirody, Otdel Biologicheskii 98: 33-40. 

Burukovsky, R.N., 1993c. Plesionika alaini nom. nov., a new name for Plesionika crosnieri Burukovsky, 1992 

(Crustacea Decapoda Pandalidae).— Arthropoda Selecta 2: 18. 

Burukovsky, R.N., 1995. Two new species of shrimps of the genus Pasiphaea and new records of other 

shrimps [in Russian].— Zoologicheskii Zhurnal 74: 121-126. 

Burukovsky, R.N., 1997. Selection of a type species for Farfantepenaeus Burukovsky (Crustacea: Decapoda: 

Penaeidae).— Proceedings of the Biological Society of Washington 110: 154. 

Burukovsky, R.N., 2000a. Taxonomy of shrimps from the genus Nematocarcinus (Crustacea, Decapoda, 

Nematocarcinidae). 2. Revision of N. longirostris and N. altus, correction of species diagnosis in N. 

sigmoides and N. lanceopes, and description of new species, N. batei and N. romenskyi [in Russian].— 


Burukovsky, R.N., 2000b. Taxonomy of shrimps of the genus Nematocarcinus (Decapoda, Nematocarcinidae). 

5. Redescription of Nematocarcinus nudirostris and description of N. combensis, N. kaiensis, 

and N. subtilis [in Russian].— Zoologicheskii Zhurnal 79: 1036-1044. 

Burukovsky, R.N., 2000c. Taxonomy of shrimps from the genus Nematocarcinus (Decapoda, Nematocarcinidae). 

3. Description of new species [in Russian].— Zoologicheskii Zhurnal 79: 662-668. 

Burukovsky, R.N., 2000d. Taxonomy of shrimps of the genus Nematocarcinus (Crustacea, Decapoda, 

Nematocarcinidae). 7. Description of new species, N. hanamuri and N. evansi, from southwestern 

Australian waters [in Russian].— Zoologicheskii Zhurnal 79: 1290-1293. 

Burukovsky, R.N., 2000e. Taxonomy of shrimps from the genus Nematocarcinus (Crustacea, Decapoda, 

Nematocarcinidae). 4. Description of species from tenuirostris group [in Russian].— Zoologicheskii 

Zhurnal 79: 898-906. 

Burukovsky, R.N., 2000f. Taxonomy of shrimps from the genus Nematocarcinus (Decapoda, Nematocarcinidae). 

6. Redescription of species from the groups undulatipes and gracilis with descriptions of 

two new species [in Russian].— Zoologicheskii Zhurnal 79: 1155-1167. 

Burukovsky, R.N., 2000g. Lysmata splendida sp. nov., a new species of shrimp from the Maldives (Crustacea: 

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Burukovsky, R.N., 2001. Taxonomy of Nematocarcinus (Decapoda, Nematocarcinidae). Description of 

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Holthuis, L.B., 1946b. Note on the genus Pandalina (Crustacea Decapoda), with the description of a new 

species from European waters.— Zoologische Mededeelingen 26: 281-286. 

Holthuis, L.B., 1947a. The Decapoda of the Siboga Expedition. Part IX. The Hippolytidae and Rhynchocinetidae 

collected by the Siboga and Snellius expeditions with remarks on other species.— Siboga 

Expeditie 39a8: 1-100. 

Holthuis, L.B., 1947b. Nomenclatorial notes on European macrurous Crustacea Decapoda.— Zoologische 

Mededeelingen 27: 312-322. 

Holthuis, L.B., 1948. Note on some Crustacea Decapod Natantia from Surinam.— Proceedings van de 

Koninklijke Nederlandsche Akademie van Wetenschappen 51: 1104-1113. 

Holthuis, L.B., 1949a. The identity of Penaeus monodon Fabr.— Proceedings van de Koninklijke Nederlandsche 

Akademie van Wetenschappen 52: 1051-1057. 

Holthuis, L.B., 1949b. On some species of Macrobrachium (Crustacea Decapoda) from West Africa.— Eos, 

Revista Española de Entomología 25: 175-185. 

Holthuis, L.B., 1949c. Note on the species of Palaemonetes (Crustacea Decapoda) found in the United States 

of America.— Proceedings van de Koninklijke Nederlandsche Akademie van Wetenschappen 52: 87-95. 

Holthuis, L.B., 1950a. The Decapoda of the Siboga Expedition. Part X. The Palaemonidae collected by 

the Siboga and Snellius expeditions with remarks on other species. I. Subfamily Palaemoninae.— 

Siboga Expeditie 39a9: 1-268.


Holthuis, L.B., 1950b. Preliminary descriptions of twelve new species of palaemonid prawns from 

American waters (Crustacea Decapoda).— Proceedings van de Koninklijke Nederlandsche Akademie 

van Wetenschappen 53: 93-99. 

Holthuis, L.B., 1950c. Description d’une nouvelle espèce du genre Periclimenes Costa (Crustacés Décapodes, 

Natantia) des Côtes Algériennes.— Bulletin des Travaux publiés par la Station Expérimentale 

l’Aquiculture et de Pêche de Castiglione (n.s.) 2: 3-12. 

Holthuis, L.B., 1951a. The caridean Crustacea of Tropical West Africa.— Atlantide Report. Scientific 

Results of the Danish Expedition to the Coasts of Tropical West Africa 1945-1946 2: 7-187. 

Holthuis, L.B., 1951b. A general revision of the Palaemonidae (Crustacea Decapoda Natantia) of the 

Americas. I. The subfamilies Euryrhynchidae and Pontoniinae.— Occasional Papers of the Allan 

Hancock Foundation 11: 1-332. 

Holthuis, L.B., 1952a. Crustacés Décapodes, Macrures.— Expédition Océanographique Belge dans les 

Eaux Côtières Africaines de l’Atlantique Sud (1948-1949). Résultats Scientifiques 3 (2): 1-88. 

Holthuis, L.B., 1952b. A general revision of the Palaemonidae (Crustacea Decapoda Natantia) of the 

Americas. II. The subfamily Palaemoninae.— Occasional Papers of the Allan Hancock Foundation 

12: 1-396. 

Holthuis, L.B., 1952c. The Decapoda of the Siboga Expedition. Part XI. The Palaemonidae collected by 

the Siboga and Snellius Expeditions with remarks on other species II. Subfamily Pontoniinae.— 

Siboga Expeditie 39a10 : 1-253. 

Holthuis, L.B., 1952d. The Crustacea Decapoda Macrura of Chile. Reports of the Lund University Chile 

Expedition 1948-49, vol 5.— Lunds Universitets Årsskrift (N. F.) (= Kungl. Fysiografiska Sällskapets 

Handlingar. N.F. Bd 62 (10)) 47: 1-110. 

Holthuis, L.B., 1953a. On the type specimens of Vanderbiltia rosamondae Boone (Crustacea Decapoda 

Macrura).— Zoologische Mededelingen 32: 113-118. 

Holthuis, L.B., 1953b. Enumeration of the Decapod and Stomatopod Crustacea from Pacific Coral islands.— 

Atoll Research Bulletin 24: 1-66. 

Holthuis, L.B., 1954. On a collection of decapod Crustacea from the Republic of El Salvador (central 

America).— Zoologische Verhandelingen 23: 1-43, Plates 1-2. 

Holthuis, L.B., 1955a. Note on the genus Miyadiella Kubo (Crustacea Decapoda Penaeidae) with the description 

of a new species.— Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i 

Københaven 117: 75-81. 

Holthuis, L.B., 1955b. The recent genera of the caridean and stenopodidean shrimps (Class Crustacea, 

order Decapoda, supersection Natantia) with keys for their determination.— Zoologische Verhandelingen 

26: 1-157. 

Holthuis, L.B., 1956a. An enumeration of the Crustacea Decapoda Natantia inhabiting subterranean 

waters.— Vie et Milieu 7: 43-76. 

Holthuis, L.B., 1956b. Selection of a specimen preserved in the British Museum (Natural History), London, 

to be the lectotype of the nominal species ”Pandalus montagui” [Leach], [1814], and ”Pandalus 

annulicornis” Leach, [1815] (Class Crustacea, Order Decapoda). In: Hemming, A.F., Direction 47. 

Substitution on the Official List of Generic Names in Zoology of a revised entry relating to the generic 

name Pandalus Leach, 1815 (Class Crustacea, Order Decapoda)(revision of a Ruling given in 

Opinion 104) .— Opinions and declarations rendered by the International Commission on Zoological 

Nomenclature 1 (D): 251. ICZN, London. 

Holthuis, L.B., 1958. Crustacea Decapoda from the northern Red Sea (Gulf of Aqaba and Sinai Peninsula). 

I. Macrura.— Bulletin of the Sea Fisheries Research Station, Haifa 17: 1-40. 

Holthuis, L.B., 1959a. The Crustacea Decapoda of Suriname (Dutch Guiana).— Zoologische Verhandelingen 

44: 1-296, Plates 1-16. 

Holthuis, L.B., 1959b. Results of the reexamination of the type specimens of some species belonging to 

the subfamilies Pontoninae and Palaemoninae (Crustacea Decapoda Macrura).— Zoologische Mededelingen 

36: 193-200. 

Holthuis, L.B., 1960. Two new species of atyid shrimps from subterranean waters of N.W. Australia 

(Decapoda Natantia).— Crustaceana 1: 47-57.


Holthuis, L.B., 1961a. On the dates of publication of the crustacean plates in Duperrey’s ”Voyage autour 

du Monde...sur...La Coquille.— Crustaceana 3: 168-169. 

Holthuis, L.B., 1961b. Designation of a neotype for Peneus membranaceus Risso, 1816 (Class Crustacea, 

Order Decapoda) In: China, W.E., Opinion 611. Parapenaeus S.I.Smith, 1885 (Crustacea, Decapoda); 

Validation under the Plenary Powers and Interpretation of Peneus membranaceus Risso, 1816.— Bulletin 

of Zoological Nomenclature 18: 309-310, Plate 4. 

Holthuis, L.B., 1961c. Report on a collection of Crustacea Decapoda and Stomatopoda from Turkey and 

the Balkans.— Zoologische Verhandelingen 47: 1-67, Plates 1-2. 

Holthuis, L.B., 1961d. A new species of Merhippolyte (Decapoda Natantia) from east American waters.— 


Holthuis, L.B., 1963a. Two new species of freshwater shrimp (Crustacea Decapoda) from the West 

Indies.— Proceedings van de Koninklijke Nederlandsche Akademie van Wetenschappen (C) 66: 

61-69. 

Holthuis, L.B., 1963b. On red coloured shrimps (Decapoda, Caridea) from tropical land-locked saltwater 

pools.— Zoologische Mededelingen 38: 261-279. 

Holthuis, L.B., 1965a. The Atyidae of Madagascar.— Mémoires du Muséum national d’Histoire naturelle 

(A) Zoologie 33: 1-48. 

Holthuis, L.B., 1965b. A new fresh-water prawn of the genus Macrobrachium (Crustacea, Decapoda, 

Caridea) from Madagascar.— Zoologische Mededelingen 40: 281-285. 

Holthuis, L.B., 1966. The R/V Pillsbury deep-sea biological expedition to the Gulf of Guinea, 1964-65. 11. 

The freshwater shrimps of the Island of Annobon, West Africa.— Studies in Tropical Oceanography 

4: 224-239. 

Holthuis, L.B., 1970. Études hydrobiologiques en Nouvelle-Calédonie (Mission 1965 du premier Institut 

de Zoologie de l’Université de Vienne) (suite). IX. The freshwater shrimps (Crustacea Decapoda, 

Natantia) of New Caledonia.— Cahiers ORSTOM, série Hydrobiologique 3: 87-108 [imprint 1969]. 

Holthuis, L.B., 1971. Biological results of the University of Miami deep-sea Expeditions. 75. The Atlantic 

shrimps of the deep-sea genus Glyphocrangon A. Milne-Edwards, 1881.— Bulletin of Marine Science 

21: 267-373. 

Holthuis, L.B., 1973a. Caridean shrimps found in land-locked saltwater pools at four Indo-West Pacific 

localities (Sinai Peninsula, Funafuti Atoll, Maui and Hawaii Islands), with the description of one 

new genus and four new species.— Zoologische Verhandelingen 128: 1-48, Plates 1-7. 

Holthuis, L.B., 1973b. Bithynops luscus, a new genus and species of cavernicolous shrimp from Mexico 

(Crustacea Decapoda, Palaemonidae).— Quaderno del Accademia Nazionale dei Lincei 171: 135- 

142. 

Holthuis, L.B., 1973c. Biological results of the University of Miami deep-sea Expeditions. 99. Mohocaris, 

a new genus of alpheid shrimps from the Caribbean region (Crustacea, Decapoda, Natantia).— Bulletin 

of Marine Science 23: 489-495. 

Holthuis, L.B., 1974. Subterranean Crustacea Decapoda Macrura collected by Mr. L. Botosaneanu during 

the 1973 Cuban-Roumanian Biospeleological Expedition to Cuba.— International Journal of 

Speleology 6: 231-242. 

Holthuis, L.B., 1976a. The identities of Pandalus gracilis Stimpson, 1860 and Pandalus prensor Stimpson, 

1860 (Decapoda, Pandalidae).— Crustaceana 30: 49-54. 

Holthuis, L.B., 1976b. The authorship of the names of species of Crustacea Decapoda published in 1835 

by S. Hailstone and J.O. Westwood.— Zoologische Mededelingen 49: 285-291. 

Holthuis, L.B., 1977a. The Mediterranean decapod and stomatopod Crustacea in A. Risso’s published 

works and manuscripts.— Annales du Muséum d’Histoire naturelle de Nice 5: 37-88. 

Holthuis, L.B., 1977b. Cave shrimps (Crustacea Decapoda, Natantia) from Mexico.— Quaderno del Accademia 

Nazionale dei Lincei 171: 173-195. 

Holthuis, L.B., 1978a. A collection of Decapod Crustacea from Sumba, Lesser Sunda islands, Indonesia.— 

Zoologische Verhandelingen 162: 1-55, Plate 1. 

Holthuis, L.B., 1978b. Zoological results of the British speleological expedition to Papua New Guinea 

1975. 7. Cavernicolous shrimps (Crustacea Decapoda, Natantia) from New Ireland and the Philippines.— 

Zoologische Mededelingen 53: 209-224.


Holthuis, L.B., 1980a. The identity of Hapalopoda investigator Filhol, 1885 (Decapoda, Penaeidae) and 

other shrimps collected by the 1880-1883 ”Travailleur” and ”Talsiman” expeditions.— Zoologische 

Mededelingen 55: 183-194. 

Holthuis, L.B., 1980b. FAO Species Catalogue. Vol. 1 - Shrimps and Prawns of the World. An Annotated 

Catalogue of Species of Interest to Fisheries.— FAO Fisheries Synopsis 125 (Vol. 1): i-xvii, 1-271. 

Holthuis, L.B., 1980c. Caridina lanzana, a new troglobitic shrimp from Somalia (Crustacea Decapoda).— 

Monitore Zoologico Italiano (n.s) suppl. 13: 1-10. 

Holthuis, L.B., 1980d. Alpheus saxidomus new species, a rock boring snapping shrimp from the Pacific 

coast of Costa Rica, with notes on Alpheus simus Guérin-Méneville, 1856.— Zoologische Mededelingen 

55: 47-58. 

Holthuis, L.B., 1981. Description of three new species of shrimps (Crustacea: Decapoda: Caridea) from 

Pacific Islands.— Proceedings of the Biological Society of Washington 94: 787-800. 

Holthuis, L.B., 1982. Notes on Indo-West Pacific Crustacea Decapoda I and II.— Crustaceana 42: 26-36. 

Holthuis, L.B., 1984a. Two freshwater prawns of the genus Macrobrachium (Crustacea Decapoda: Palaemonidae) 

from New Guinea.— Zoologische Mededelingen 58: 163-174. 

Holthuis, L.B., 1984b. Freshwater prawns (Crustacea Decapoda: Natantia) from subterranean waters of 

the Gunung Sewu area, central Java, Indonesia.— Zoologische Mededelingen 58: 141-148. 

Holthuis, L.B., 1986a. Fresh-water shrimps of the family Atyidae (Crustacea: Decapoda) from western 

Colombia.— Journal of Crustacean Biology 6: 438-445. 

Holthuis, L.B., 1986b. A new genus and species of subterranean shrimp from Western Australia (Crustacea: 

Decapoda: Atyidae).— Zoologische Mededelingen 60: 103-111. 

Holthuis, L.B., 1986c. Some Pontoniinae (Crustacea: Decapoda: Palaemonidae) from southern Oman.— 


Holthuis, L.B., 1991. Marcgraf’s (1648) Brazilian Crustacea.— Zoologische Verhandelingen 268: 1-123. 

Holthuis, L.B., 1993a. The recent genera of the caridean and stenopodidean shrimps (Crustacea, Decapoda) 

with an appendix on the order Amphionidacea: 1-328. Nationaal Natuurhistorisch Museum, 

Leiden. 

Holthuis, L.B., 1993b. The non-Japanese new species established by W. de Haan in the Crustacea Volume 

of Fauna Japonica (1833-1850). In: Yamagushi, T. (ed.), Ph.F. von Siebold and natural history of 

Japan, Crustacea: 599-642. Carcinological Society of Japan, Tokyo. 

Holthuis, L.B., 1995. Notes on Indo-West Pacific Crustacea Decapoda III to IX.— Zoologische Mededelingen 

69: 139-151. 

Holthuis, L.B., 1996. [Review of] H.-G. Müller, 1993. Catalogue of the Indo-Pacific pontoniine shrimps; 

and: A.J. Bruce, 1994. A synopsis of the Indo-West Pacific genera of the Pontoniinae (Crustacea: 

Decapoda: Palaemonidae).— Crustaceana 69: 806-808. 

Holthuis, L.B. & I. Eibl-Eibesfeldt, 1964. A new species of the genus Periclimenes from Bermuda (Crustacea, 

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Holthuis, L.B. & E. Gottlieb, 1958. An annotated list of the decapod Crustacea of the Mediterranean 

coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean.— Bulletin of 

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Holthuis, L.B. & K.-I. Hayashi, 1967. A new species of shrimp, Rhynchocinetes hiatti (Crustacea, Decapoda).— 


Holthuis, L.B. & C. Maurin, 1952. Note sur Lysmata uncicornis nov. spec. et sur deux autres espèces intéressantes 

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Received: 16.v.2011 

Accepted: 15.vi.2011 

Edited: L.P. van Ofwegen


Acanthanas ............................. 376 

Acanthephyra ......................... 260 

Acanthephyridae ................ 260 

Acetes ...................................... 239 

Aegaeon ................................... 454 

Agostocarididae .................. 291 

Agostocaris ............................. 291 

Alainopasiphaea ..................... 254 

Alcockpenaeopsis ................... 213 

Alcyonohippolyte ................... 414 

Allopontonia ........................... 339 

Allosergestes ........................... 240 

Alope ....................................... 414 

Alpheidae .............................. 376 

Alpheopsis ............................... 376 

Alpheus ................................... 377 

Altelatipes ............................... 209 

Altopontonia ........................... 339 

Alvinocarididae ................... 292 

Alvinocaridinides ................... 292 

Ambidexter ............................. 436 

Amphibetaeus ......................... 391 

Amphipontonia ...................... 339 

Anachlorocurtis ..................... 440 

Anapontonia ........................... 340 

Anchiopontonia ...................... 340 

Anchistioides .......................... 308 

Anchistioididae ................... 308 

Anchistus ................................ 340 

Ancylomenes .......................... 341 

Anisomenaeus ........................ 342 

Antecaridina ........................... 267 

Apopontonia ........................... 342 

Arachnochium ........................ 314 

Arafurapenaeopsis ................. 213 

Araiopontonia ........................ 342 

Archaeatya .............................. 267 

Arete ........................................ 391 

Aretopsis ................................. 391 

Argis ........................................ 454 

Aristaeomorpha ...................... 206 

Aristaeopsis ............................ 206 

Aristeidae .............................. 206 

Aristeus ................................... 206 

Ascidonia ................................ 342 

Athanas ................................... 391 

Athanopsis .............................. 391 

Atlantopandalus .................... 440 

Atya ......................................... 267 

Atyaephyra ............................. 268 

Atydina ................................... 268 

Systematic index to currently valid families and genera 

Atyella ..................................... 268 

Atyidae .................................. 267 

Atyoida .................................... 269 

Atyopsis .................................. 270 

Atypopenaeus ......................... 213 

Australatya ............................. 270 

Austropandalus ...................... 440 

Austropenaeus ........................ 207 

Automate ................................ 393 

Balssia ..................................... 343 

Bannereus ............................... 394 

Barbouria ................................ 412 

Barbouriidae ......................... 412 

Batella ...................................... 394 

Batepenaeopsis ....................... 213 

Bathyhippolyte ....................... 414 

Bathypalaemonella ................. 306 

Bathypalaemonellidae ....... 306 

Bathypalaemonetes ................ 307 

Bathystylodactylus ................ 304 

Bentheogennema .................... 209 

Benthesicymidae ................. 209 

Benthesicymus ....................... 209 

Benthonectes ........................... 211 

Bermudacaris .......................... 394 

Betaeopsis ................................ 394 

Betaeus .................................... 395 

Birulia ..................................... 414 

Bitias ....................................... 440 

Blepharocaris .......................... 343 

Brachycarpus .......................... 315 

Bresilia .................................... 294 

Bresiliidae ............................. 294 

Brucecaris ............................... 343 

Bruceonia ................................ 343 

Bruceopsis ............................... 395 

Bythocarides ........................... 414 

Bythocaris ............................... 415 

Cainonia .................................. 343 

Calathaemon ........................... 313 

Calipandalus ........................... 440 

Calliasmata ............................. 415 

Campylonotidae .................. 307 

Campylonotus ........................ 307 

Caridella .................................. 270 

Caridina .................................. 270 

Caridinides ............................. 282 

Caridinopsis ........................... 282 

Caridion .................................. 415 

Carinopontonia ...................... 344 

Chacella ................................... 344 

Chelonika ................................ 440 

Chernocaris ............................. 344 

Chlorocurtis ............................ 440 

Chlorotocella ........................... 441 

Chlorotocoides ........................ 452 

Chlorotocus ............................. 441 

Chorismus ............................... 416 

Chorocaris ............................... 293 

Cinetorhynchus ...................... 300 

Climeniperaeus ...................... 344 

Clytomanningus .................... 436 

Colemonia ............................... 344 

Conchodytes ........................... 344 

Coralliocaris ........................... 345 

Coronalpheus .......................... 396 

Coutieralpheus ....................... 396 

Coutierea ................................. 346 

Coutierella .............................. 315 

Crangon .................................. 455 

Crangonidae ......................... 454 

Creaseria ................................. 316 

Crinotonia ............................... 346 

Cryphiops ................................ 316 

Cryptocheles ........................... 416 

Cryptopenaeus ....................... 232 

Ctenopontonia ........................ 346 

Cuapetes .................................. 346 

Dactylonia .............................. 348 

Dasella ..................................... 348 

Dasycaris ................................ 348 

Deioneus ................................. 396 

Deosergestes ........................... 241 

Desmocarididae .................. 309 

Desmocaris ............................. 309 

Diapontonia ............................ 348 

Dichelopandalus .................... 442 

Disciadidae ........................... 295 

Discias ..................................... 295 

Dorodotes ................................ 442 

Dugastella ............................... 282 

Edoneus ................................... 282 

Encantada ............................... 295 

Engystenopus ......................... 249 

Ephyrina ................................. 263 

Epipontonia ............................ 349 

Eualus ..................................... 416 

Eugonatonotidae ................. 296 

Eugonatonotus ....................... 296 

Eumanningia .......................... 418 

Eupasiphae .............................. 255 

Eupontonia ............................. 349


Euryrhynchidae ................... 309 

Euryrhynchina ....................... 309 

Euryrhynchoides .................... 309 

Euryrhynchus ........................ 309 

Eusergestes ............................. 242 

Exhippolysmata ...................... 418 

Exoclimenella ......................... 349 

Exopalaemon .......................... 316 

Exopontonia ............................ 349 

Farfantepenaeus ..................... 214 

Fennera ................................... 349 

Fenneralpheus ........................ 396 

Fenneropenaeus ...................... 214 

Funchalia ................................ 214 

Gallocaris ................................ 282 

Ganjampenaeopsis ................. 215 

Gelastocaris ............................ 419 

Gelastreutes ............................ 419 

Gennadas ................................ 211 

Globospongicola ..................... 249 

Glyphocrangon ....................... 467 

Glyphocrangonidae ............ 467 

Glyphus ................................... 255 

Gnathophylleptum ................. 309 

Gnathophyllidae ................. 309 

Gnathophylloides ................... 311 

Gnathophyllum ...................... 311 

Gordonella .............................. 232 

Hadropenaeus ......................... 232 

Haliporoides ............................ 233 

Haliporus ................................ 233 

Halocaridina ........................... 282 

Halocaridinides ...................... 283 

Hamiger .................................. 349 

Hamodactyloides .................... 349 

Hamodactylus ........................ 350 

Hamopontonia ........................ 350 

Harperalpheus ........................ 396 

Harpiliopsis ............................ 350 

Harpilius ................................. 350 

Hayashidonus ......................... 436 

Hemipenaeus .......................... 207 

Hepomadus ............................. 208 

Heptacarpus ........................... 419 

Heterocarpus .......................... 442 

Heterogenys ............................ 263 

Heteropenaeus ........................ 215 

Hippolyte ................................ 420 

Hippolytidae ........................ 414 

Holthuisaeus ........................... 351 

Holthuispenaeopsis ................ 215 

Hymenocera ............................ 313 

Hymenoceridae ................... 313 

Hymenodora ........................... 263 

Hymenopenaeus ..................... 233 

Ischnopontonia ....................... 351 

Isopontonia ............................. 351 

Izucaris .................................... 351 

Janicea ..................................... 413 

Janicella ................................... 265 

Jengalpheops ........................... 397 

Jocaste ...................................... 351 

Jolivetya .................................. 283 

Jonga ........................................ 283 

Juxtastenopus ......................... 252 

Kakaducarididae ................. 314 

Kakaducaris ............................ 314 

Kemphyra ................................ 264 

Kirnasia ................................... 296 

Kishinouyepenaeopsis ........... 215 

Lancaris ................................... 283 

Laomenes ................................. 351 

Latreutes ................................. 423 

Leander .................................... 317 

Leandrites ............................... 318 

Lebbeus .................................... 424 

Lenzicarcinus ......................... 297 

Leontocaris .............................. 427 

Leptalpheus ............................. 397 

Leptathanas ............................ 398 

Leptocarpus ............................ 318 

Leptochela ............................... 255 

Leptomenaeus ......................... 352 

Leptopalaemon ....................... 314 

Leslibetaeus ............................. 398 

Levicaris .................................. 311 

Ligur ........................................ 427 

Limnocaridella ........................ 283 

Limnocaridina ........................ 283 

Lipkebe .................................... 352 

Lipkemenes ............................. 352 

Lipkius ..................................... 297 

Lissocrangon .......................... 456 

Lissosabinea ............................ 456 

Litopenaeus ............................. 216 

Lucaya ..................................... 296 

Lucifer ..................................... 238 

Luciferidae ............................ 238 

Lysmata ................................... 427 

Lysmatella ............................... 430 

Macrobrachium ...................... 318 

Macromaxillocarididae ..... 249 

Macromaxillocaris ................. 249 

Macropetasma ........................ 216 

Mancicaris .............................. 284 

Manipontonia ......................... 352 

Margitonia .............................. 352 

Marosina ................................. 284 

Marsupenaeus ........................ 216 

Megokris ................................. 216 

Melicertus ............................... 217 

Meningodora .......................... 264 

Merguia ................................... 430 

Merhippolyte .......................... 430 

Mesocrangon .......................... 456 

Mesopenaeus .......................... 234 

Mesopontonia ......................... 353 

Metabetaeus ............................ 398 

Metacrangon .......................... 456 

Metalpheus ............................. 398 

Metapenaeopsis ...................... 217 

Metapenaeus ........................... 222 

Metapontonia ......................... 353 

Micratya ................................. 284 

Microprosthema ..................... 250 

Mierspenaeopsis ..................... 224 

Mimocaris ............................... 430 

Miopontonia ........................... 353 

Mirocaris ................................. 293 

Miropandalus ......................... 444 

Mohocaris ............................... 398 

Nauticaris ............................... 430 

Nautilocaris ............................ 293 

Nematocarcinidae ............... 297 

Nematocarcinus ..................... 297 

Nematopalaemon ................... 332 

Nennalpheus ........................... 398 

Neoanchistus .......................... 353 

Neocaridina ............................ 284 

Neoclimenes ............................ 353 

Neocrangon ............................ 458 

Neopalaemon .......................... 332 

Neopericlimenes ..................... 353 

Neopontonides ........................ 353 

Neosergestes ........................... 242 

Neostylodactylus .................... 304 

Nigmatullinus ........................ 300 

Nikoides .................................. 437 

Nippontonia ........................... 354 

Notalpheus .............................. 399 

Notocrangon ........................... 459 

Notopandalus ......................... 444 

Notopontonia .......................... 354 

Notostomus ............................ 264 

Odontonia ............................... 354 

Odontozona ............................ 252 

Ogyrides ................................. 435 

Ogyrididae ............................ 435 

Onycocaridella ....................... 354


Onycocaridites ....................... 354 

Onycocaris .............................. 354 

Onycomenes ........................... 355 

Opaepele .................................. 293 

Oplophoridae ....................... 265 

Oplophorus ............................. 265 

Orthopontonia ........................ 355 

Orygmalpheus ........................ 399 

Palaemon ................................. 332 

Palaemonella ........................... 355 

Palaemonetes .......................... 336 

Palaemonias ............................ 286 

Palaemonidae ....................... 314 

Pandalidae ............................ 440 

Pandalina ................................ 444 

Pandalopsis ............................. 445 

Pandalus ................................. 445 

Pantomus ................................ 446 

Parabetaeus ............................. 399 

Paracaridina ........................... 286 

Paraclimenaeus ...................... 356 

Paraclimenes .......................... 356 

Paracrangon ........................... 459 

Parahepomadus ...................... 208 

Paralatreutes .......................... 431 

Paralebbeus ............................. 431 

Paranchistus ........................... 357 

Parapasiphae ........................... 256 

Parapenaeopsis ....................... 224 

Parapenaeus ........................... 224 

Parapontocaris ....................... 460 

Parapontophilus ..................... 460 

Parasergestes .......................... 243 

Paraspongicola ....................... 251 

Parastylodactylus .................. 304 

Paratya .................................... 286 

Paratypton .............................. 357 

Parhippolyte ........................... 413 

Parisia ..................................... 287 

Pasiphaea ................................ 257 

Pasiphaeidae ........................ 254 

Patonia .................................... 357 

Peisos ....................................... 244 

Pelagopenaeus ........................ 225 

Penaeidae .............................. 213 

Penaeopsis ............................... 226 

Penaeus ................................... 226 

Periclimenaeus ....................... 357 

Periclimenella ......................... 360 

Periclimenes ........................... 360 

Periclimenoides ...................... 366 

Peripandalus ........................... 447 

Petalidium .............................. 244 

Philarius ................................. 366 

Philocheras .............................. 461 

Phycocaris ............................... 432 

Phycomenes ............................ 367 

Phyllognathia ......................... 313 

Physetocarididae ................. 453 

Physetocaris ............................ 453 

Pinnotheronia ......................... 368 

Placopsicrangon ..................... 464 

Platycaris ................................ 368 

Platypontonia ......................... 368 

Pleoticus .................................. 235 

Plesiomenaeus ........................ 368 

Plesionika ................................ 447 

Plesiopenaeus ......................... 208 

Plesiopontonia ........................ 368 

Pliopontonia ........................... 368 

Pomagnathus .......................... 399 

Pontocaris ............................... 464 

Pontonia .................................. 368 

Pontonides .............................. 370 

Pontoniopsides ....................... 370 

Pontoniopsis ........................... 370 

Pontophilus ............................ 465 

Poripontonia ........................... 370 

Potamalpheops ....................... 399 

Potimirim ................................ 287 

Prionalpheus ........................... 400 

Prionocrangon ........................ 465 

Procarididae ......................... 248 

Procaris ................................... 248 

Processa ................................... 437 

Processidae ........................... 436 

Procletes .................................. 451 

Propontonia ............................ 370 

Protrachypene ........................ 227 

Psalidopodidae .................... 303 

Psalidopus ............................... 303 

Psathyrocaris .......................... 259 

Pseudalpheopsis ..................... 400 

Pseudaristeus ......................... 208 

Pseudathanas .......................... 400 

Pseudocheles ........................... 296 

Pseudochelidae .................... 296 

Pseudoclimenes ...................... 370 

Pseudocoutierea ...................... 370 

Pseudopalaemon ..................... 337 

Pseudopandalus ..................... 452 

Pseudopontonia ...................... 371 

Pseudopontonides .................. 371 

Pseudopontophilus ................. 465 

Pseudoveleronia ..................... 371 

Pterocaris ................................ 400 

Puteonator .............................. 288 

Pycneus ................................... 288 

Pycnisia ................................... 288 

Pycnocaris .............................. 311 

Racilius ................................... 400 

Rapipontonia .......................... 371 

Rhopalaemon .......................... 338 

Rhynchocinetes ...................... 301 

Rhynchocinetidae ............... 300 

Rhynocrangon ........................ 465 

Richalpheus ............................ 401 

Richardina .............................. 253 

Rimapenaeus .......................... 227 

Rimicaris ................................. 293 

Rostronia ................................. 371 

Rugathanas ............................. 401 

Sabinea .................................... 466 

Salmoneus ............................... 401 

Sandimenes ............................. 371 

Sandyella ................................. 372 

Saron ....................................... 432 

Sclerocrangon ......................... 466 

Segonzackomaius ................... 300 

Sergestes .................................. 244 

Sergestidae ............................ 239 

Sergia ....................................... 244 

Shinkaicaris ............................ 294 

Sicyonella ................................ 248 

Sicyonia ................................... 229 

Sicyoniidae ............................ 229 

Sinodina .................................. 288 

Solenocera ............................... 235 

Solenoceridae ....................... 232 

Spirontocaris .......................... 432 

Spongicola ............................... 251 

Spongicolidae ....................... 249 

Spongicoloides ........................ 251 

Spongiocaris ........................... 252 

Stegopontonia ......................... 372 

Stenalpheops ........................... 403 

Stenopodidae ....................... 252 

Stenopus ................................. 253 

Stygiocaris .............................. 289 

Stylodactylidae .................... 304 

Stylodactyloides ..................... 304 

Stylodactylus .......................... 305 

Stylopandalus ......................... 452 

Synalpheus ............................. 403 

Syncaris .................................. 289 

Syncrangon ............................ 466 

Systellaspis ............................. 266 

Tanypenaeus ........................... 227 

Tectopontonia ......................... 372


Tenuipedium ........................... 338 

Thalassocarididae ............... 452 

Thalassocaris .......................... 453 

Thaumastocaris ...................... 372 

Thinora .................................... 433 

Thor ......................................... 433 

Thorella ................................... 434 

Thuylamea .............................. 412 

Tozeuma .................................. 434 

Trachycaris ............................. 435 

Trachypenaeopsis ................... 227 

Trachypenaeus ........................ 227 

Trachysalambria ..................... 228 

Triacanthoneus ....................... 412 

Tridiscias ................................. 296 

Troglindicus ............................ 338 

Troglocaris .............................. 289 

Troglocubanus ........................ 338 

Troglomexicanus .................... 339 

Tuleariocaris ........................... 372 

Typhlatya ................................ 290 

Typhlocarididae .................. 375 

Typhlocaridina ....................... 291 

Typhlocaris ............................. 375 

Typhlopatsa ............................. 291 

Typton ..................................... 372 

Typtonoides ............................. 373 

Typtonychus ........................... 373 

Unguicaris .............................. 374 

Urocaridella ............................ 339 

Urocaris .................................. 374 

Veleronia .................................. 374 

Veleroniopsis ........................... 374 

Vercoia ..................................... 467 

Vetericaris ............................... 249 

Vexillipar ................................. 412 

Vir ........................................... 374 

Waldola ................................... 375 

Xiphocarididae .................... 302 

Xiphocaris ............................... 302 

Xiphopenaeus ......................... 228 

Yagerocaris .............................. 412 

Yemenicaris ............................. 375 

Zenopontonia ......................... 375

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