Checklist of the Plants of the Guiana Shield - Department of Botany ...

SMITHSONIAN INSTITUTION 

Contributions from the United States National Herbarium 

Volume 55: 1-584 

Checklist of the Plants of the Guiana Shield 

(VENEZUELA: Amazonas, Bolivar, Delta Amacuro; 

GUYANA, SURINAM, FRENCH GUIANA) 

by 

V. Funk,T. Hollowell, P. Berry, C. Kelloff, and S. N. Alexander 

Department of Botany 

National Museum of Natural History 

Washington, DC 

2007

ABSTRACT 

Funk, V., T. Hollowell, P. Berry, C. Kelloff, and S. N. Alexander. Checklist of the Plants of the Guiana 

Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions 

from the United States National Herbarium, volume 55, 584 pages. This volume contains a checklist of the 

plants of the Guiana Shield Area (northeastern South America) produced using information gathered from historical and recent 

collections as well as many recent expeditions. The Guiana Shield, as treated here, includes the countries of Guyana and 

Surinam, French Guiana (a part of France) and the Venezuelan states of Amazonas, Bolivar, and Delta Amacuro. The checklist 

contains 257 vascular plant families with 13,367 species. According to the list 6,791 of those species are from Amazonas, 6,669 

from Bolívar, 1,651 from Delta Amacuro, 7,112 from Guyana, 4,984 from Surinam, and 5,406 are from French Guiana. Of these 

species, 293 (2.2%) are estimated to be introduced and naturalized and even fewer are invasive. The small number of naturalized 

species is a good indication of the large amount of intact vegetation from this area of South America. The largest family is the 

Leguminoseae with 146 genera containing 1032 species; 41 families are represented by only one species. Over 250 scientists 

from around the world participated in this project by identifying specimens and checking species lists. This volume should be of 

use to students, taxonomists, ecologists, and conservation biologists, as well as to interested amateurs. 

KEY WORDS (not in title): South America, biodiversity, endemic species, invasive species 

DATE OF PUBLICATION: 31 July 2007 

Cover Design by Alice R. Tangerini: front, illustration by Alice R. Tangerini: new species of Paloue, habit, 

leaf base detail, ovary detail, anther and stipules (K.M. Redden 4027); back, illustration by Alice R. 

Tangerini: Nautilocalyx pemphidius L. Skog; habit, flower bud, open flower, corolla face, longitudinal 

section of flower, stamens, pistil, and nectary, fruit, seeds (Bell 324) 

Contributions from the United States National Herbarium (ISSN 0097-1618) Department of Botany, National 

Museum of Natural History, MRC-166, Smithsonian Institution, Washington, DC, 20013-7012, USA. 

POSTMASTER: Send address changes to Contributions from the U.S. National Herbarium, Department 

of Botany, National Museum of Natural History, MRC-166, P.O. Box 37012, Smithsonian Institution, 

Washington, DC, 20013-7012, USA. 

The paper used in this publication meets the minimum requirements of the American National Standard 

for Permanence of Paper for Printed Library Materials Z39.48—1984. 

The periodical, Contributions from the United States National Herbarium, was first published in 1890 by 

The United States Department of Agriculture. From 1 July 1902 forward it was published as a Bulletin of 

the United States National Museum. The series was discontinued after volume 38 (1974), and has been 

revived with volume 39 (2000) as a venue for publishing longer taxonomic papers, checklists, floras, and 

monographs, produced by the staff and associates at the U.S. National Herbarium. It is externally peer 

reviewed, and published at irregular intervals. Subscription and other correspondence should be addressed 

to CUSNH, Department of Botany, National Museum of Natural History, MRC-166, Smithsonian 

Institution, Washington, DC, 20013-7012, USA e-mail: CUSNH@si.edu. The present issue is available 

for free while supplies last and PDF files of the most recent issues (vol. 49-present) of the Contr. U.S. Natl. 

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manuscript submission or comments on previous volumes, please contact the editor, Paul M. Peterson 

(peterson@si.edu).

CONTENTS 

FOREWORD by Peter H. Raven ....................................................................................... 

INTRODUCTION by V. A. Funk & T. Hollowell ............................................................ 

Geology of the Guiana Shield .............................................................................. 

Climate ...................................................................................................................... 

Diversity ................................................................................................................. 

Conservation .......................................................................................................... 

Discussion ............................................................................................................. 

Using the Checklist ................................................................................................ 

Nomenclatorial considerations ................................................................................ 

Acknowledgements ................................................................................................ 

References .............................................................................................................. 

CHECKLIST ......................................................................................................................... 

Psilotophyta ........................................................................................................... 

Lycophyta .............................................................................................................. 

Pterophyta .............................................................................................................. 

Coniferophyta ......................................................................................................... 

Cycadophyta .......................................................................................................... 

Gnetophyta ............................................................................................................. 

Anthophyta ............................................................................................................ 

Monocotyledonea ...................................................................................... 

Dicotyledonea .......................................................................................... 

LITERATURE CITATION FOR ILLUSTRATIONS ................................................................. 

APPENDIX: Individuals who identified plants ........................................................... 

INDEX TO GENERA .......................................................................................................... 

INDEX TO FAMILIES ......................................................................................................... 

5 

7 

8 

9 

9 

10 

10 

13 

14 

14 

14 

17 

17 

17 

21 

56 

56 

57 

57 

58 

183 

562 

563 

565 

584

Henriettea multiflora Naudin (Melastomataceae)


FOREWORD 

At a time when human activities have placed enormous stress on the natural world, there is still 

much that we do not know about the Earth’s biodiversity; the great majority of species have not yet been 

described, and very few of those that have been named are known in any detail. Checklists provide us 

with a first approximation of what we know, and can provide a profile of the biodiversity in relatively 

poorly-known parts of the globe. For many groups, they may represent the most complete information 

available. Since basic taxonomic information provides essential information for studies in systematic 

and evolutionary biology, we continually seek to improve and update it. Collecting specimens and writing 

checklists, flora treatments, and monographs are all essential elements in our quest to understand the 

natural world and how it evolved. 

In northern South America, one of the richest regions of the world in terms of biodiversity, our 

knowledge has grown with the recent publication of the Checklist of the Plants of Peru, Checklist of the 

Plants of Ecuador, and the nearly-completed Checklist of the Plants of Bolivia. This current volume, 

which covers all of the plants known from the Guiana Shield region in northeastern South America, is 

another outstanding contribution to this global effort. All perrtinent information about this region will 

be brought together in a comprehensive New World Checklist of Vascular Plants and Bryophytes 

(http://mobot.mobot.org/W3T/Search/nwc.html). All of these projects contribute to our overall mission 

to understand, preserve, and protect the wealth of biodiversity that is such an outstanding feature of this 

incredible part of the world. 

Checklists have many uses: they are aids in the identification and correct naming of species, essential 

resources for biodiversity estimates and biogeographic studies, and the starting points for more detailed 

studies of an area’s biota. Reviewed by many specialists, they often represent the most advanced state of 

knowledge available in their field. They also increase national and regional pride and provide important 

public outreach and fundamental information to use in addressing the biodiversity crisis. This checklist 

of the plants of the Guiana Shield, combined with the recently published Checklist of the Terrestrial 

Vertebrates of the Guiana Shield, increases our knowledge of this fascinating region and acts as a starting 

point from which additional research may be developed. It gives insight into the species richness, 

endemicity, and floral and faunal affinities of this region. Such lists can also provide the baseline 

information for analyzing species turnover rates, migration, invasion events, and many other biological 

phenomena. 

In the face of the growing biodiversity crisis, we celebrate the collaborative nature of a project such 

as this. It is an excellent example of effective interaction, involving over 250 people from around the 

world who contributed in diverse ways to its completion. These contributors variously collected plants 

from the area, provided identifications of and information about those plants, or reviewed drafts of the 

checklist, and together made possible its successful completion. 

It is with great pleasure that I welcome this important addition to our knowledge of the biodiversity 

of Earth, and at the same time congratulate the many individuals and institutions who made it possible. 

Peter H. Raven 

President, Missouri Botanical Garden 

George Engelmann Professor of Botany, Washington University in St. Louis 

5

. 

6 


Figure 1: The Guiana Shield



(VENEZUELA: Amazonas, Bolivar, Delta Amacuro; 

GUYANA, SURINAM, FRENCH GUIANA) 

by 

V. Funk 1 , T. Hollowell 2 , P. Berry 3 , C. Kelloff 1 , and S. N. Alexander 1 

The Guiana Shield region has long held a 

fascination for tropical biologists because of its 

unique geography that includes table-top 

mountains (tepuis), tropical savannas, and broad 

expanses of rainforests. These areas are home to 

many endemic taxa, as well as to unique 

ecosystems such as the Greenheart 

(Chlorocardium) forests of Guyana and the 

montane savannas of Brazil, Venezuela, and 

Guyana. Some of the largest expanses of 

untouched Neotropical forests occur in these areas. 

The Checklist of the Plants of the Guiana 

Shield covers all vascular plants known to occur 

in the Guiana Shield region of northeastern South 

America. It is the product of the combined efforts 

of two research initiatives, the Smithsonian 

Institution’s Biological Diversity of the Guiana 

Shield Program (BDG) and the Missouri Botanical 

Garden’s Flora of the Venezuelan Guayana project 

(FVG). The database created for the Checklist of 

the Plants of the Guianas (Boggan et al., 1992, 

1997) has been extensively updated and refined, 

and most importantly the nomenclature has been 

coordinated with a database extracted from the 

Flora of the Venezuelan Guayana (Steyermark, 

Berry, and Holst, 1995—2005). 

INTRODUCTION 

V. A. Funk and T. Hollowell 

This plant checklist is a companion to the 

recently published Checklist of the Terrestrial 

Vertebrates of the Guiana Shield (Hollowell and 

Reynolds, eds. 2005). Together they represent a 

new research and conservation resource which 

highlights three critical facets of taxonomic work: 

research, collections, and expeditions. These 

volumes should be of particular use to students, 

taxonomists, ecologists, and conservation 

biologists, as well as to interested amateurs. Both 

of these checklists were produced using 

information gathered from historical and recent 

collections as well as many recent expeditions to 

northeastern South America. The expertise of 

scientists from many organizations around the 

globe has been drawn together and made available 

to the scientific and conservation communities in 

both hard copy and on the web (Hollowell et al., 

2000). Research projects such as this would not 

be possible without considerable international 

collaboration from over 100 scientists who provide 

determinations for the collections (see Appendix). 

Checklists have many uses. They are aids in 

the identification of organisms, resources for 

biodiversity estimates and biogeographic studies, 

1 Department of Botany and Biological Diversity of the Guiana Shield Program (BDG), National Museum of Natural History, 

MRC-166, P.O.Box 37012, Smithsonian Institution, Washington, DC 20013-7012 U.S.A. 

2 BDG and Office of Information Technology, National Museum of Natural History, P.O.Box 37012, Smithsonian Institution, 

Washington, DC 20013-7012 U.S.A. 

3 University of Michigan Herbarium & Dept. of Ecology and Evolutionary Biology, 3600 Varsity Drive, Ann Arbor, Michigan 

48108-2287 U.S.A. 

7

8 

nomenclators, and essential starting points for more 

detailed studies of an area’s biota. The Guiana 

Shield is a geologic formation, and as such it 

defines a section of northeastern South America 

that is ideally a natural unit rather than a political 

one (Kelloff and Funk 2004). This gives these two 

checklists (plants and terrestrial vertebrates) an 

environmental focus that is a first step toward 

thinking in terms of ecosystems, evolution, and 

systematics rather than simple lists of organisms. 

For instance, checklists can tell us how many 

organisms of a certain kind are in a particular area, 

thus giving us an idea of the species richness. 

Properly utilized, the data can serve as an indicator 

of endemicity within an area and provide 

information on the number of introduced species, 

and they can be used in a comparative manner 

between and among areas. Since this is a natural 

area, they can be used to formulate questions about 

adaptive radiation, dispersal, and evolutionary 

history. In fact, data from these checklists are 

already being used to examine endemicity levels, 

species turnover rates, and the location of areas 

most in need of additional study. The checklists 

are particularly useful for these types of 

investigations because they encompass several 

groups of organisms (plants, amphibians, reptiles, 

birds, and mammals) facilitating comparisons 

among the different groups. Conservation efforts 

are improved because the checklists provide 

standardized species nomenclature that will be 

used in both governmental and academic 

undertakings, including impact studies, reserve 

planning, ecological research, production of floral 

and faunal studies, and biogeographical analyses, 

allowing data sets from different sources to be 

compared with greater accuracy and confidence. 

Both checklists exemplify the collaborative 

nature of studies in biological diversity science, 

being reflections of the efforts of so many 

specialists essential to their production. They are 

truly international efforts, for while the volumes 

were organized by the BDG staff, the authors and 

experts who supplied identifications are based all 

over the world and in many types of organizations. 

The result affirms the common interests of all of 

the people, institutions, and nations involved in 

studying, understanding, and conserving the 

irreplaceable natural heritage of our world. 


GEOLOGY OF THE GUIANA SHIELD 

The Guiana Shield (Figure 1) is a distinct 

ancient geological formation that today is roughly 

bounded by the Atlantic Ocean to the east, the 

Orinoco River to the north and west, the Rio Negro 

(a major tributary of the Amazon River) to the 

southwest, and the Amazon River to the south 

(Gibbs and Barron 1993). The Shield includes the 

mountain systems that form the watershed 

boundary between the Amazon and Orinoco rivers. 

On the Shield’s western side, the Orinoco River 

and Río Negro are connected by the Río Casiquiare. 

The base of the Guiana Shield is composed of 

crystalline rocks of Proterozoic origin; these are 

mainly granites and gneisses formed between 3.6 

and 0.8 billion years ago (Mendoza 1977; Schubert 

and Huber 1990). Large areas of the Shield were 

overlain with sediments from 1.6 to 1 billion years 

ago (Huber 1995a), and remnants of these 

sandstones comprise the Roraima group of 

formations that are scattered across the central 

portion of the Shield. Erosion has decreased the 

size of these formations, and the remaining 

sediments extend west from the Potaro escarpment 

of the Pakaraima Mountains in central Guyana 

through parts of Venezuela and Colombia 

(Arbeláez and Callejas 1999) and south into 

northern Brazil (Leechman 1913; Gibbs and Barron 

1993). Within this area, erosion has resulted in 

numerous vertical-walled, frequently flat-topped 

mountains called “tepuis,” among them Chimantátepui 

(2,550 m), Cerro Duida (2,358 m) and Auyántepui 

(2,450 m). Pico de Neblina (3,014 m) is the 

Shield’s westernmost tepui and highest point, 

located on the southernmost segment of the border 

between Venezuela and Brazil. Mount Roraima 

(2,810 m) is located at the juncture of Guyana, 

Venezuela, and Brazil and includes the highest 

point within Guyana. The easternmost peaks or 

the shield are located in Guyana and reach 

approximately 2,000 m elevation, including Mt. 

Ayanganna (2,080 m) and Mt. Wokomung (2,000 

m). Most Venezuelan tepuis are between 2,000 

and 2,400 m and the highest point is Pico de 

Neblina in Venezuela/Brazil (over 3,000 m). 

Several of these mesa-like formations are virtually 

inaccessible by foot and are so unusual that a 

fictional scientific expedition account referred to 

one as “The Lost World” (Doyle 1912), a term 

sometimes applied to all tepuis. Notable waterfalls

of the region include Angel Falls (979 m) on 

Auyán-tepui in Venezuela and Kaieteur Falls (226 

m) on the Potaro River in Guyana’s Pakaraima 

Mountains. 

Granitic dome mountains occur in the southern 

part of the three Guianas (Guyana, Surinam, French 

Guiana), where they are known as “inselbergs,” as 

well as in Venezuela where they are known as 

“lajas” and extend in a broad arc all the way from 

northeastern Bolivar state west and south along the 

Orinoco and then further south as far as Brazil. 

Deposits of low-nutrient white sands occur inland 

of the coastal plain, in belts across the region and 

in isolated pockets. Large areas of savanna are 

found in the region, particularly the complex of 

savannas that includes the Rupununi Savanna in 

southwestern Guyana, the Gran Sabana in eastern 

Bolívar, Venezuela, and the savannas of northern 

Roraima, Brazil. In some parts of these areas there 

is limited forest growth because sands overlay a 

clay hardpan that is resistant to penetration by tree 

roots and because they flood during the heavy rainy 

season. Tertiary and Quaternary sediments 

separate the southern edge of the Guiana Shield 

from the Amazon River and the eastern edge of 

the Shield from the Atlantic Ocean. 

The boundaries of the Guiana Shield are 

difficult to define precisely. In the south between 

mountains on the southern boundary of the Shield 

and the Amazon and Negro rivers there is a broad 

band of outwash materials resulting from erosion. 

In the Venezuelan state of Delta Amacuro much of 

the area is covered by thick sediments deposited 

primarily by the Orinoco River. In the west, there 

are remnants of the Shield at least as far as 

Colombia. In the east and south the hills give way 

in some areas to lowland tropical rainforest more 

similar to that found in the Amazon basin. 

However, some remains of the Guiana Shield occur 

in the southern section, and some portion of the 

sediments are derived from outwash from the 

highlands of the Shield. For more detailed 

discussions of the geology of the area, readers 

should refer to Gibbs and Barron (1993) and Huber 

(1995a). 

CLIMATE 


As a whole, the Guiana Shield region has a 

tropical climate characterized by a relatively high 

mean annual temperature exceeding 25º C at sea 

level, an annual monthly maximum temperature 

range of less than 5º C, and an average daily 

temperature range of approximately 6º C. Because 

of the Guiana Shield’s location just north of the 

equator, its climate varies primarily according to 

elevation and effects of the trade winds that 

combine to affect rainfall patterns. The trade winds 

blow consistently from the east and northeast, off 

of the Atlantic Ocean onto northeastern South 

America. Due to orographic effects, the 

easternmost escarpments of the mountains of the 

Guiana Shield are generally localities of increased 

precipitation where these moisture-laden winds 

meet the slopes (Clarke et al. 2001). Seasonal 

oscillations of the Inter-tropical Convergence Zone 

(ITCZ) also bring variations in rainfall as the 

locations of low pressure zones near the equator 

change (Snow 1976). Varying primarily by 

latitude, one or two rainy seasons result from shifts 

in the ITCZ. The heaviest rains usually occur 

between May and August, whereas the rainy season 

running from December to January is shorter and 

less intense, with rains that do not penetrate as far 

inland. In areas where there is only one rainy 

season, the driest part of the year is January to 

March; in areas where there are two rainy seasons 

the driest months are March and October. 

However, even during most dry seasons, frequent 

storms provide adequate moisture to allow 

evergreen tropical moist forests to persist in most 

low elevation parts of the region. 

DIVERSITY 

The varied topology of the Guiana Shield 

includes sandstone tepuis, granite inselbergs, white 

sands, seasonally flooded tropical savannas, 

lowlands with numerous rivers, isolated mountain 

ranges, and coastal swamps, each supporting a 

characteristic vegetation (Huber et al. 1995; Huber 

1995b). This variety accounts for a great deal of 

the high diversity and endemicity of the Shield’s 

biota. The Shield has a flora and fauna with 

numerous endemic species, ca. 40% according to 

a recent analysis (Kelloff and Funk 2004). Some 

tepui endemic species occur as low as 300 m 

elevation, with increasing numbers by 1,500 to 

1,800 m, and fully developed communities 

occurring by 2,000 m (Funk and Berry, 2005). 

9

10 

Most parts of Brazil north of the Amazon River 

and much of eastern Colombia are poorly explored, 

and they are not included in this publication (see 

below). 

CONSERVATION 

With the exceptions of the few populated 

localities such as Puerto Ayacucho, Ciudad 

Guayana, Ciudad Bolívar, and the agricultural 

coastal areas, the environment of the Guiana Shield 

has benefited from limited access and low 

population densities, although this same isolation 

has hindered biodiversity research. Estimates vary, 

but approximately 70% (Funk & Richardson 2003) 

of the vegetation is still relatively undisturbed by 

human activities. Recently, however, the pace of 

disturbance has greatly increased; current threats 

to the environment include large-scale logging by 

Asian and local companies, large- and small-scale 

gold and diamond mining, oil prospecting, bauxite 

mining, massive hydroelectric projects, wildlife 

trade, and population-related pressures such as 

burning, grazing, agriculture, and the expansion 

of Amerindian villages. Taken together, these 

impacts have begun to take their toll, with vast 

areas previously left undisturbed are now 

vulnerable to increasing disturbance. 

The status of conservation efforts varies by 

country. Throughout the Guiana Shield, many 

areas that are designated as protected or otherwise 

restricted are often only “paper” parks because of 

a lack of infrastructure and funds to protect them. 

Over the last four decades, Venezuela has 

established seven national parks, 29 natural 

monuments, and two biosphere reserves covering 

about 142,280 km², more than 30% of its share of 

the Guiana Shield (Funk and Berry 2005). In 

Guyana, the progress of conservation efforts has 

been slower, with the only substantial protected 

area being Kaieteur National Park, its 627 km² 

comprising about 0.3% of the country’s area 

(Kelloff 2003), with additional reserves under 

consideration. Guyana’s 3,710 km² Iwokrama 

Forest (Clarke et al. 2001) is dedicated to 

sustainable use. Surinam’s protected areas system 

includes one national park and a network of 11 

reserves, totaling almost 20,000 km², over 12% of 

its total area. This includes the recently created 

16,000 km² Central Suriname Nature Reserve, an 

UNESCO World Heritage Site that joined and 


expanded three existing reserves (see http:// 

www.stinasu.com). French Guiana has no 

officially designated protected areas, but 18 

proposed sites total 6,710 km², about 7.5% of its 

area (Lindeman and Mori 1989). 

The Guiana Shield encompasses part or all of 

five countries (excluding the small part in 

Colombia) with five different governments, five 

official languages, and many more indigenous 

languages. Cooperation is sometimes hampered 

by border disputes and illegal cross-border 

transportation of gold, wildlife, and more recently, 

illegal drugs. The implementation of conservation 

practices is further complicated by many issues 

concerning the indigenous peoples of the region. 

All of these challenges will have to be overcome 

on the way to designing and maintaining a viable 

reserve system for the Guiana Shield. 

DISCUSSION 

This checklist includes 257 vascular plant 

families containing 2,072 accepted genera. We 

have recorded over 14,732 accepted subgeneric 

taxa, 1,358 of those including a subspecific or 

varietal epithet and 13,367 distinct species. 

According to our list 6,791 of those species are 

from Amazonas, 6,669 from Bolívar, 1,651 from 

Delta Amacuro, 7,112 from Guyana, 4,984 from 

Surinam, and 5,406 from French Guiana. Of these 

species, 293 (2.2%) are estimated to be introduced 

and naturalized. In the three Guianas there are 

8,933 species and there are 9,814 species in the 

Venezuelan Guayana. With 5,482 species found 

in both areas there is 41% overlap between the two 

areas. Within the checklist area 3,451 species are 

listed only for the Guianas and 4,332 are found 

only in the Venezuelan Guayana. See Table 1 for 

a summary of the numbers of taxa in each family 

and Table 2 for the families with the largest number 

of species. 

The top 15 families (in number of species) 

have a total of 7,094 species, which represents over 

half of the species in the Guiana Shield area. 

According to our list 154 families have 25 species 

or less, and 34 families have 26-50 species. 

Consequently, 73% of families have no more than 

50 species and 41 families (16%) are represented 

by only one species.

Table 1. Number of species in each plant family. 

PSILOPHYTA (1) 

Psilotaceae 1 

LYCOPHYTA (107) 

Isoëtaceae 5 

Lycopodiaceae 27 

Selaginellaceae 75 

PTEROPHYTA (754) 

Adiantaceae 71 

Aspleniaceae 36 

Blechnaceae 23 

Cyatheaceae 46 

Davalliaceae 1 

Dennstaedtiaceae 63 

Dicksoniaceae 1 

Dryopteridaceae 26 

Gleicheniaceae 18 

Grammitidaceae 62 

Hymenophyllaceae 89 

Hymenophyllopsidaceae 8 

Lomariopsidaceae 79 

Lygodiaceae 4 

Marattiaceae 7 

Marsileaceae 3 

Metaxyaceae 1 

Oleandraceae 8 

Ophioglossaceae 3 

Osmundaceae 2 

Parkeriaceae 4 

Plagiogyriaceae 1 

Polypodiaceae 59 

Pteridaceae 16 

Salviniaceae 5 

Schizaeaceae 21 

Tectariaceae 21 

Thelypteridaceae 44 

Vittariaceae 16 

Woodsiaceae 16 

CONIFEROPHYTA (10) 

Pinaceae 1 

Podocarpaceae 9 

CYCADOPHYTA (5) 

Cycadaceae 1 

Zamiaceae 4 


GNETOPHYTA (6) 

Gnetaceae 6 

ANTHOPHYTA 

MONOCOTYLEDONEA (3301) 

Agavaceae 2 

Alismataceae 13 

Araceae 226 

Arecaceae 111 

Bromeliaceae 327 

Burmanniaceae 29 

Cannaceae 2 

Commelinaceae 26 

Costaceae 16 

Cyclanthaceae 33 

Cyperaceae 413 

Dioscoreaceae 54 

Eriocaulaceae 94 

Haemodoraceae 3 

Heliconiaceae 19 

Hydrocharitaceae 2 

Iridaceae 13 

Lemnaceae 8 

Liliaceae 22 

Limnocharitaceae 3 

Marantaceae 85 

Mayacaceae 3 

Musaceae 2 

Najadaceae 3 

Orchidaceae 1020 

Poaceae 526 

Pontederiaceae 9 

Rapateaceae 83 

Ruppiaceae 1 

Smilacaceae 24 

Strelitziaceae 1 

Taccaceae 1 

Thurniaceae 3 

Triuridaceae 9 

Typhaceae 1 

Velloziaceae 2 

Xyridaceae 100 

Zingiberaceae 11 

Zosteraceae 1 

DICOTYLEDONEA (9183) 

Incertae sedis 7 

11 

Acanthaceae 115 

Aizoaceae 1 

Amaranthaceae 30 

Anacardiaceae 23 

Anisophylleaceae 1 

Annonaceae 178 

Apiaceae 4 

Apocynaceae 211 

Aquifoliaceae 76 

Araliaceae 64 

Aristolochiaceae 32 

Asclepiadaceae 132 

Balanophoraceae 2 

Balsaminaceae 1 

Basellaceae 2 

Bataceae 1 

Begoniaceae 14 

Bignoniaceae 175 

Bixaceae 4 

Bombacaceae 55 

Bonnetiaceae 28 

Boraginaceae 62 

Brassicaceae 8 


Brunelliaceae 2 

Burseraceae 81 

Buxaceae 1 

Cabombaceae 4 

Cactaceae 24 

Campanulaceae 10 

Canellaceae 1 

Capparaceae 33 

Caprifoliaceae 3 

Caricaceae 3 

Caryocaraceae 10 

Caryophyllaceae 3 

Casuarinaceae 1 

Cecropiaceae 42 

Celastraceae 22 

Ceratophyllaceae 1 

Chenopodiaceae 1 

Chloranthaceae 3 

Chrysobalanaceae 186 

Clethraceae 1 

Clusiaceae 198 

Combretaceae 48 

Compositae 303

12 

Connaraceae 30 

Convolvulaceae 107 

Crassulaceae 2 

Cucurbitaceae 56 

Cunoniaceae 13 

Cuscutaceae 5 

Cyrillaceae 2 

Dichapetalaceae 10 

Dilleniaceae 38 

Droseraceae 14 

Ebenaceae 25 

Elaeocarpaceae 50 

Eremolepidaceae 3 

Ericaceae 73 

Erythroxylaceae 37 

Euphorbiaceae 317 

Euphroniaceae 3 

Flacourtiaceae 64 

Gentianaceae 92 

Gesneriaceae 94 

Haloragaceae 2 

Hernandiaceae 6 

Hippocrateaceae 52 

Hugoniaceae 8 

Humiriaceae 30 

Hydrophyllaceae 3 

Icacinaceae 22 

Ixonanthaceae 10 

Juglandaceae 1 

Juncaceae 1 

Krameriaceae 3 

Lacistemataceae 3 

Lamiaceae 38 

Lauraceae 214 

Lecythidaceae 67 

Leguminosae 

- Caesalpinioideae 294 

- Faboideae 500 

- Mimosoideae 238 

Lentibulariaceae 55 

Lissocarpaceae 3 

Loasaceae 2 

Loganiaceae 43 

Loranthaceae 57 


Table 1. Number of species in each plant family. (contd.) 

Lythraceae 35 

Magnoliaceae 2 

Malpighiaceae 205 

Malvaceae 85 

Marcgraviaceae 17 

Melastomataceae 534 

Meliaceae 49 

Mendonciaceae 14 

Menispermaceae 48 

Menyanthaceae 1 

Molluginaceae 3 

Monimiaceae 4 

Monotaceae 1 

Moraceae 83 

Moringaceae 1 

Muntingiaceae 1 

Myoporaceae 1 

Myricaceae 1 

Myristicaceae 33 

Myrsinaceae 70 

Myrtaceae 270 

Nelumbonaceae 1 

Nyctaginaceae 50 

Nymphaeaceae 15 

Ochnaceae 165 

Olacaceae 35 

Oleaceae 4 

Onagraceae 21 

Opiliaceae 2 

Oxalidaceae 12 

Papaveraceae 1 

Passifloraceae 107 

Pedaliaceae 2 

Peridiscaceae 2 

Phytolaccaceae 12 

Picramniaceae 9 

Piperaceae 172 

Plantaginaceae 2 

Plumbaginaceae 2 

Podostemaceae 67 

Polygalaceae 76 

Polygonaceae 53 

Portulacaceae 15 

Primulaceae 1 

Proteaceae 20 

Quiinaceae 32 

Rafflesiaceae 2 

Ranunculaceae 2 

Rhabdodendraceae 1 

Rhamnaceae 26 

Rhizophoraceae 13 

Rosaceae 10 

Rubiaceae 742 

Rutaceae 81 

Sabiaceae 11 

Santalaceae 1 

Sapindaceae 158 

Sapotaceae 148 

Sarraceniaceae 10 

Scrophulariaceae 52 

Simaroubaceae 12 

Siparunaceae 10 

Solanaceae 116 

Sphenocleaceae 1 

Sterculiaceae 67 

Styracaceae 22 

Surianaceae 1 

Symplocaceae 13 

Tepuianthaceae 4 

Ternstroemiaceae 31 

Tetrameristaceae 1 

Theaceae 1 

Theophrastaceae 4 

Thymelaeaceae 15 

Tiliaceae 37 

Trigoniaceae 12 

Tropaeolaceae 1 

Turneraceae 31 

Ulmaceae 5 

Urticaceae 15 

Valerianaceae 1 

Verbenaceae 90 

Violaceae 39 

Viscaceae 60 

Vitaceae 16 

Vochysiaceae 84 

Winteraceae 1 

Zygophyllaceae 5

Table 2: Families with largest numbers of species. 


Rank Family No. of Genera No. of Species 

1 Leguminoseae 146 1032 

- Mimosoideae 31 238 

- Caesalpinioideae 41 294 

- Faboideae 74 500 

2 Orchidaceae 171 1020 

3 Ferns* 95 754 

4 Rubiaceae 97 742 

5 Melastomataceae 54 534 

6 Poaceae 116 526 

7 Cyperaceae 37 413 

8 Bromeliaceae 24 327 

9 Euphorbiaceae** 60 317 

10 Compositae 104 303 

11 Myrtaceae 18 270 

12 Araceae 24 226 

13 Lauraceae 18 214 

14 Apocynaceae*** 38 211 

15 Malpighiaceae 25 205 

* Includes many families 

** Includes the small families Putrangiivaceae, Picrodendraceae, and Phyllanthaceae 

which do not belong in a monophyletic Euphorbiaceae 

*** Does not include the Asclepidaceae (18 genera and 132 species) 

The largest family on the Shield is the 

Leguminoseae (= Fabaceae) with 1032 species. 

The phylogeny of this family is under revision but 

it is well enough understood to know that this 

family has a rich and interesting history in the 

Guiana Shield area. We know from the molecular 

data that there are a number of genera that have 

their primary radiation in the area and that there 

are many endemics. Very little is known about the 

phylogeny of the Guiana Shield members of the 

other large family, the Orchidaceae. 

In a broad scope of botanical understanding, 

the combination of such checklists looks to the 

evolutionary and ecological importance of physical 

conditions beyond those of political units and 

boundaries. This process is simply a step in the 

direction of looking at a flora at the level of the 

distinct geological unit of the Guiana Shield. In 

the future there will be additional field trips and 

further progress on the Flora of the Guianas 

(Nationaal Herbarium Nederland 2006) which will 

provide additions and corrections to this list. Also, 

future developments will hopefully include the 

13 

ability to analyze plant community composition 

on finer levels of the landscape, and to produce 

customized checklists for analysis and 

conservation with the aid of Geographic 

Information System (GIS) technologies drawing 

upon improved databases incorporating 

georeferenced historical collections from many 

herbaria. 

USING THE CHECKLIST 

For each family, accepted names are listed 

followed by the known distributions. 

Abbreviations used are Amazonas (AM), Bolívar 

(BO), and Delta Amacuro (DA); and for the three 

Guianas: Guyana (GU), Surinam (SU), and French 

Guiana (FG). 

Comments following the accepted name or 

synonym are enclosed in brackets and include 

aspects such as: exotic species, distributional 

expectations or details, and in the case of 

misidentifications an indication that the taxon is 

not found in the flora area. Some taxa are listed

14 

with comments naming Mt. Roraima, Serrania de 

la Neblina or other localities without distribution 

codes. Specimens from these areas may lack a 

country designation or may be incorrect or 

unknown, especially in historical collections. 

Alternatively, a species may be known from 

neighboring Brazil and if it is thought that it is 

likely to occur in the checklist area it is included 

as part of the flora. 

NOMENCLATURAL CONSIDERATIONS 

Family names generally follow the 

classification used by Cronquist (1981) but the list 

has been modified to reflect some recent changes 

based on molecular and morphological data and 

presented on the website of Peter Stevens (Stevens, 

2001). An occasional dilemma in combining plant 

checklists is the necessity to choose among 

classifications used by different specialists. When 

decisions had to be made we selected the most 

recent comprehensive work for the group in 

question. However, our decision does not imply 

an endorsement of one or rejection of another and 

it is hoped that an adequate reference system has 

been provided to allow the use of most systems 

and to facilitate work towards future clarification. 

Authors of plant names are standardized according 

to Brummitt and Powell (1992). 

In some cases, a species has been reported 

from the flora area either because of a broad species 

circumscription that has since been restricted (or 

as an infraspecific taxon that has since been raised 

to species status), or due to misidentifications or 

other errors. Such names are listed in the 

Synonyms and Misidentification section as “not 

in the flora area.” 

In some of the cases source data for one region 

lists subspecific taxa while data for the other region 

lists only the specific level. In those cases, both 

the specific and subspecific distributions are given 

rather than eliminating the partial information for 

the subspecific taxa. 

When the list has been reviewed by a specialist 

that persons name is listed under the taxon name. 

All other Family lists were compiled by T. 

Hollowell. 


ACKNOWLEDGMENTS 

Special thanks go to botanists at the Missouri 

Botanical Garden (MO), particularly Kay 

Yatskievych, James Miller, Ronald Liesner, Robert 

Magill, and many others who assisted in the Flora 

of the Venezuelan Guayana project and shared the 

electronic manuscript with the BDG program. 

Also, we thank several colleagues including Scott 

Mori (NY) and Jean-Jacques de Granville (CAY) 

who provided data in the development of the first 

checklist of the three Guianas. In addition, our 

work would not have been possible without the 

dedication of botanical specialists from many 

institutions who contributed to checklists, 

taxonomic treatments, did countless identifications, 

and reviewed drafts of the combined lists. Each 

family treatment lists the name of the collaborating 

specialists for that family and the Appendix lists 

all of those who helped with identifications. We 

owe a great debt to the many plant collectors and 

their field crews who over more than two centuries 

have provided specimens for the museum 

collections essential to an understanding of the 

flora of the Guiana Shield (Ek, 1990, 1991; 

Hollowell et al. 2000, 2003, 2005). Equally 

indispensable to studies of biodiversity are the 

many institutions that have financed those 

expeditions and those that house the collections, 

and their staff members that maintain and organize 

those collections. In addition to the home 

institutions we must also thank the various funding 

agencies that have made field work possible, chief 

among these is the National Geographic Society 

who funded quite a few expeditions for botanists 

and zoologists and USAID in Guyana. In Guyana 

the collecting program was hosted by the Centre 

for the Study of Biological Diversity, University 

of Guyana, and we appreciate their help. Particular 

thanks for assistance are given to Marilyn Hansel, 

Christian Feuillet, Bob DeFilipps, Susan Grose, 

Karen Redden, Matt Sewell, H. David Clarke, Julie 

Barcelona, Greg McKee, Sylvia Stone, and Mark 

Strong (all from US). Finally, we must thank Peter 

Jorgensen (MO) who provided a careful review of 

the manuscript. 

This publication was prepared under the 

auspices of the Biological Diversity of the Guiana 

Shield Program. This is number 125 in the 

Smithsonian’s Biological Diversity of the Guiana 

Shield publication series.

REFERENCES 

Arbeláez, M. V., and R. Callejas. 1999. Flórula 

de la Meseta de Arensica de la comunidad de 

Monochoa (Región de Araracuara, Medio 

Caquetá. Tropenbos - Colombia, Bogota. 

Boggan, J., V. A. Funk, C. L. Kelloff, M. Hoff, G. 

Cremers, & C. Feuillet. 1992. Checklist of 

the Plants of the Guianas. Biological 

Diversity of the Guianas Program, 

Smithsonian Institution, Washington, DC. 381 

pp. 

Boggan, J., V. Funk, C. L. Kelloff, M. Hoff, G. 

Cremers, & C. Feuillet. 1997. Checklist of 

the Plants of the Guianas, 2nd Edition. 

Biological Diversity of the Guianas Program, 

Smithsonian Institution, Washington, DC. 238 

pp. 

Brummitt, R. K., and C. E. Powell, eds. 1992. 

Authors of Plant Names. Royal Botanic 

Gardens, Kew. 

Clarke, H. D., V. A. Funk, and T. Hollowell. 2001. 

Using checklists and collections data to 

investigate plant diversity. I: A comparative 

checklist of the plant diversity of the 

Iwokrama Forest, Guyana. Sida Botanical 

Miscellany 21: 1-86. 

Cronquist, A. 1981. An Integrated System of 

Classification of Flowering Plants. Columbia 

University Press, New York. 

Doyle, A. C. 1912. The Lost World. Puffin Books, 

London 

Ek, R. C. 1990. Index of Guyana Plant Collectors. 

Flora of the Guianas, A.R.A. Görts-van Rijn 

ed., Supplementary Fascicle 1. Koeltz 

Scinetific Books, Koenigstein, Germany. 

Ek, R. C. 1991. Index of Suriname Plant 

Collectors. Flora of the Guianas, A.R.A. 

Görts-van Rijn ed., Supplementary Fascicle 

2. Koeltz Scinetific Books, Koenigstein, 

Germany. 

Funk, V. A. and P. E. Berry. 2005. Chapter 4.3 

The Guiana Shield. In Plant Conservation: A 

natural history approach. University of 

Chicago Press, pages 76-79. 

Funk, V. A. and K. Richardson. 2002. Biological 

Specimen Data in Biodiversity Studies: Use 

it or lose it. Systematic Biology 51: 303-316. 

Funk, V. A., K. S. Richardson, and S. Ferrier. 2005. 

Survey-gap analysis in expeditionary research, 


15 

where do we go from here? Biological Journal 

of the Linnean Society 85: 549-567. 

Gibbs, A. K. and C. N. Barron. 1993. The Geology 

of the Guiana Shield. Oxford University 

Press, New York. 

Hollowell, T., V. Funk, and C. Kelloff. 2000 

(onwards). Biological Diversity of the Guiana 

Shield. www.mnh.si.edu/biodiversity/bdg 

Hollowell, V. A. Funk, C. L. Kelloff & G. 

Gharbarran. 2000. Smithsonian Plant 

Collections, Guyana: 1986-1987, John J. 

Pipoly III. Smithsonian Institution, 

Washington, D.C. 

Hollowell, T., L. J. Gillespie, V. A. Funk, and C. 

L. Kelloff. 2003. Smithsonian Plant 

Collections, Guyana: 1989-1991, Lynn J. 

Gillespie. Contributions from the United States 

National Herbarium 44: 1-104. 

Hollowell, T., T. McDowell, V. A. Funk, C. L. 

Kelloff, and D. Gopaul. 2005. Smithsonian 

Plant Collections, Guyana: 1990-1991, Tim 

McDowell. Contributions from the United 

States National Herbarium 50: 1-150. 

Hollowell, T. and R. P. Reynolds, eds. 2005. 

Checklist of the Terrestrial Vertebrates of the 

Guiana Shield. Bulletin of the Biological 

Society of Washington, no. 12. 

Huber, O. 1995a. Geography and Physical 

Features. Pp 1-61 in: P.E. Berry, B.K. Holst, 

and K. Yatskievych (eds.), Flora of the 

Venezuelan Guayana. Volume 1: Introduction. 

J.A. Steyermark, Paul E. Berry, and B.K. Holst 

(general eds.). Missouri Botanical Garden, St. 

Louis. 

Huber, O. 1995b. Vegetation. Pp 97-160 in: P.E. 

Berry, B.K. Holst, and K. Yatskievych (eds.), 

Flora of the Venezuelan Guayana. Volume 1: 

Introduction. J.A. Steyermark, P.E. Berry, and 

B.K. Holst (general eds.) Missouri Botanical 

Garden, St. Louis. 

Huber, O., G. Gharbarran, and V.A. Funk. 1995. 

Preliminary vegetation map of Guyana. 

Biological diversity of the Guianas program, 

Smithsonian Institution, Washington, D.C. 

Kelloff, C. L. 2003. The use of biodiversity data 

in developing Kaieteur National Park, Guyana 

for ecotourism and conservation. 

Contributions to the Study of Biological 

Diversity 1: 1-44. University of Guyana, 

Georgetown. 

Kelloff, C. L., and V.A. Funk. 2004.

16 

Phytogeography of the Kaieteur Falls, Potaro 

Plateau, Guyana: floral distributions and 

affinities. Journal of Biogeography 31: 501- 

513. 

Leechman, A. 1913. The British Guiana 

Handbook. “The Argosy” Co., Ltd.; 

Georgetown, British Guiana and Dulau & Co., 

London. 

Lindeman, J. C., and S. A. Mori. 1989. The 

Guianas. In: D.G. Campbell and H.D. 

Hammond (eds.), Floristic Inventory of 

Tropical Countries. New York Botanical 

Garden, New York. pp. 375-391. 

Mendoza, V. 1977. Petrogenesis de rocas 

volcanicas Precambrics del noroeste del 

Escudo de Guayana Venezuela. Mem. V 

Congreso Geol. Venezolano, Caracas, Venez. 


Tomo II, Pp. 555-589. 

Nationaal Herbarium Nederland. 2006. www.bio. 

uu.nl/~herba/Guyana/Flora_of_the_Guianas 

Schubert, C., and O. Huber. 1990. The Gran 

Sabana: Panorama of a Region. Caracas: 

Lagoven Booklets. 

Snow, J. W. 1976. Climates of northern South 

America. In: W. Schwerdtfeger (ed.), Climates 

of Central and South America. Elsevier 

Scientific Publishing Company, Amsterdam. pp 

295-403. 

Stevens, P. F. 2001 (onwards). Angiosperm 

Phylogeny Website. www.mobot.org/MOBOT/ 

research/APweb/. 

Steyermark, J. A., P. E. Berry, and B. K. Holst, eds. 

1995-2005. Flora of the Venezuelan Guayana, 

9 vols. Missouri Botanical Garden, St. Louis. 

Guapira eggersiana (Heimerl) Lundell (Nyctaginaceae)

Checklist of the Plants of the Guiana Shield 17 

CHECKLIST OF THE PLANTS OF THE GUIANA SHIELD 

Psilotum nudum (L.) P. Beauv. 

PTERIDOPHYTES & ALLIES 

G. Cremers, R.J. Hickey, D.B. Lellinger, J.T. Mickel, R.C. Moran, B. Øllgaard, A.R. Smith 

Psilotum triquetrum Sw. =Psilotum nudum 

Isoëtes killipii C.V. Morton [Mt. Roraima] 

Isoëtes ovata N. Pfeiff. 

Isoëtes schinzii H.P. Fuchs [nomem nudum] 

Isoëtes triangula U. Weber 

Isoëtes vermiculata Hickey 

PSILOTOPHYTA 

PSILOTACEAE 

SYNONYMS AND MISIDENTIFICATIONS 

REFERENCES 

Kramer, K.U. 1990. Psilotaceae. In K.U. Kramer & P.S. Green, Families and Genera of Vascular Plants. I. Pteridophytes and 

Gymnosperms. New York: Springer-Verlag. 

Smith, A.R. 1995. Psilotaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 250-251. 

LYCOPHYTA 

GENERAL REFERENCES 

Berry, P.E., B.K. Holst, & K. Yatskievych. 1995. Flora of the Venezuelan Guayana, vol. 2. Pteridophytes, Spermatophytes 

Acanthaceae - Araceae. Missouri Botanical Garden, St. Louis, & Timber Press, Portland, Oregon. 

Cremers, G. 1997. Lycophyta. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide 

to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the 

New York Botanical Garden 76, pt. 1: 56-65. 

Cremers, G., C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des espèces de Phanérogames et de Ptéridophytes de la 

Guyane française d’après l’Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne. 

Cremers, G. & M. Hoff. 1990. Inventaire taxonomique des plantes de la Guyane française. I. Les Ptéridophytes. Mus. Nat. Hist. 

Nat., Inventaires de Faune et de Flore 54: 1-133. 

Kramer, K.U. 1978. The Pteridophytes of Suriname. An enumeration with keys of the ferns and fern-allies. Uitgaven 

“Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen” Utrecht no. 93. 

Kramer, K.U. & P.S. Green (ed.). 1990. The Families and Genera of Vascular Plants. I. Pteridophytes and Gymnosperms. New 

York: Springer-Verlag. 

Posthumous, O. 1928. The Ferns of Surinam and of French and British Guiana. In A. Pulle, Flora of Suriname supplement: 1- 

196. 

Tryon, R.M. & A.F. 1981. Ferns and Allied Plants. New York: Springer-Verlag. 


Isoëtes ovatum auct. non N. Pfeiff.: Vareschi (pro parte) =Isoëtes killipii 

REFERENCES 

Fuchs-Eckert, H.P. von. 1982. Zur heutigen Kenntnis von Vorkommen und Verbreitung der Südamerikanischen Isoëtes-Arten. 

Proc. Kon. Ned. Akad. Wetensch. (ser. C) 85: 205-260. 

Hickey, R.J. & A.R. Smith. 1995. Isoëtaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 

2: 189-190. 

Huperzia aqualupiana (Spring) Rothm. 

Huperzia beitelii B. qllg. [Mt. Roraima] 

ISOËTACEAE 

LYCOPODIACEAE 

AM GU SU FG 

BO GU 

GU FG 

GU FG 

AM BO GU 

AM 

BO GU 

AM BO

18 


Huperzia capillaris (Sodiro) Holub 

Huperzia dichotoma (Jacq.) Trevis. 

Huperzia funiformis (Cham. ex Spring) Trevis. 

Huperzia hippuridea (H. Christ) Holub 

Huperzia huberi B. qllg. 

Huperzia intermedia (Spring) Trevis. [Mt. Roraima] 

Huperzia killipii (Herter) B. qllg. 

Huperzia linifolia (L.) Trevis. 

Huperzia linifolia var. jenmanii (Underw. & F.E. Lloyd) B. qllg. & P.G. 

Windisch 

Huperzia linifolia var. linifolia 

Huperzia myrsinites (Lam.) Trevis. 

Huperzia recurvifolia Rolleri 

Huperzia reflexa (Lam.) Trevis. [Mt. Roraima] 

Huperzia robusta (Klotzsch) Holub 

Huperzia taxifolia (Sw.) Trevis. 

Huperzia tenuicaulis (Underw. & F.E. Lloyd) B. qllg. 

Huperzia wilsonii (Underw. & F.E. Lloyd) B. qllg. 

Lycopodiella alopecuroides (L.) Cranfill 

Lycopodiella alopecuroides var. integerrima (Spring) B. qllg. & P.G. 

Windisch 

Lycopodiella camporum B. qllg. & P.G. Windisch 

Lycopodiella caroliniana (L.) Pic. Serm. 

Lycopodiella caroliniana var. meridionalis (Underw. & F.E. Lloyd) 

B. qllg. & P.G. Windisch 

Lycopodiella caroliniana var. paradoxa (Mart.) B. qllg. & P.G. Windisch 

Lycopodiella cernua (L.) Pic. Serm. 

Lycopodiella cernua var. cernua 

Lycopodiella cernua var. curvata (Sw.) Kartesz & Gandhi 

Lycopodiella contexta (Mart.) Holub 

Lycopodiella iuliformis (Underw. & F.E. Lloyd) B. qllg. 

Lycopodiella iuliformis var. iuliformis 

Lycopodiella iuliformis var. tatei (A.C. Sm.) B. qllg. [Mt. Roraima] 

Lycopodiella riofrioi (Sodiro) B. qllg. 

Lycopodiella steyermarkii B. qllg. 

Lycopodium clavatum L. 

Lycopodium clavatum subsp. clavatum 

Lycopodium clavatum subsp. contiguum (Klotzsch) B. qllg. 

Lycopodium jussiaei Poir. 

BO 

BO DA GU SU FG 

BO GU 

AM BO 

AM BO 

BO 

AM BO FG 

AM BO DA GU SU FG 


BO GU FG 

AM BO GU 

AM BO GU 

AM BO 

AM BO 

AM BO GU SU FG 

GU 

AM BO SU 

AM BO GU SU 

GU SU 




BO 


GU SU FG 

GU? 

AM BO 

AM BO GU SU 

AM BO GU SU 

AM BO 

AM BO 

AM BO 

AM BO GU 

AM BO 

BO GU 

AM BO 


Diphasium jussiaei (Desv. ex Poir.) Rothm. =Lycopodium jussiaei 

Huperzia jenmanii (Underw. & F.E. Lloyd) Holub =Huperzia linifolia var. jenmanii 

Huperzia passerinoides (Kunth) Trevis. =Huperzia taxifolia 

Huperzia phelpsii (Vareschi) Holub =Huperzia robusta 

Huperzia underwoodiana (Maxon) Holub =Huperzia capillaris 

Lepidotis cernua (L.) P. Beauv. =Lycopodiella cernua 

Lepidotis contexta (Mart.) Rothm. =Lycopodiella contexta 

Lycopodiella duidae (A.C. Sm.) B. qllg. =Lycopodiella iuliformis var. tatei 

Lycopodium alopecuroides L. =Lycopodiella alopecuroides 

Lycopodium anomalum Hook. & Grev. (Selaginellaceae) =Selaginella porelloides 

Lycopodium aqualupianum Spring =Huperzia aqualupiana 

Lycopodium aristatum Willd. =Lycopodium clavatum 

Lycopodium barbatum H. Christ =Huperzia dichotoma 

Lycopodium capillaceum (Spring) Hieron. =Lycopodiella cernua 

Lycopodium capillare Sodiro =Huperzia capillaris 

Lycopodium carolinianum L. =Lycopodiella caroliniana 

Lycopodium carolinianum var. meridionale (Underw. & F.E. Lloyd) Nessel =Lycopodiella caroliniana var. meridionalis 

Lycopodium carolinianum var. paradoxum (Mart.) H. Christ =Lycopodiella caroliniana var. paradoxa 

Lycopodium cernuum L. =Lycopodiella cernua 

Lycopodium cernuum var. capillaceum Spring =Lycopodiella cernua 

Lycopodium chamaepeuce Herter =Huperzia dichotoma 

Lycopodium contextum Mart. =Lycopodiella contexta 

Lycopodium contiguum Klotzsch =Lycopodium clavatum subsp. contiguum 

Lycopodium convolutum Arn. (Selaginellaceae) =Selaginella convoluta


Lycopodium dichotomum Jacq. =Huperzia dichotoma 

Lycopodium duidae A.C. Sm. =Lycopodiella iuliformis var. tatei 

Lycopodium funiforme Cham. ex Spring =Huperzia funiformis 

Lycopodium gramineum Spring =Huperzia dichotoma 

Lycopodium guatemalense Maxon =Huperzia capillaris 

Lycopodium hippurideum H. Christ =Huperzia hippuridea 

Lycopodium intermedium Spring =Huperzia intermedia 

Lycopodium iuliforme Underw. & F.E. Lloyd =Lycopodiella iuliformis 

Lycopodium jenmanii Underw. & F.E. Lloyd =Huperzia linifolia var. jenmanii 

Lycopodium killipii (Herter) C.V. Morton =Huperzia killipii 

Lycopodium linifolium L. =Huperzia linifolia 

Lycopodium marginatum Humb. & Bonpl. ex Willd. (Selaginellaceae) =Selaginella marginata 

Lycopodium meridionale Underw. & F.E. Lloyd =Lycopodiella caroliniana var. meridionalis 

Lycopodium myrsinites Lam. =Huperzia myrsinites 

Lycopodium nudum L. (Psilotaceae) =Psilotum nudum 

Lycopodium paradoxum Mart. =Lycopodiella caroliniana var. paradoxa 

Lycopodium parkeri Hook. & Grev. (Selaginellaceae) =Selaginella parkeri 

Lycopodium passerinoides Kunth =Huperzia taxifolia 

Lycopodium penniforme Lam. (Selaginellaceae) =Selaginella radiata 

Lycopodium pensum Lellinger & Mickel =Lycopodiella riofrioi 

Lycopodium phelpsii Vareschi =Huperzia robusta 

Lycopodium plumosum sensu Aubl., non L. (Selaginellaceae) =Selaginella epirrhizos 

Lycopodium porelloides Lam. (Selaginellaceae) =Selaginella porelloides 

Lycopodium radiatum Aubl. (Selaginellaceae) =Selaginella radiata 

Lycopodium reflexum Lam. =Huperzia reflexa 

Lycopodium reflexum Lam. var. caesium Vareschi [nom. inval.] =Huperzia reflexa 

Lycopodium riofrioi Sodiro =Lycopodiella riofrioi 

Lycopodium robustum Klotzsch =Huperzia robusta 

Lycopodium roraimense Underw. & F.E. Lloyd =Huperzia myrsinites 

Lycopodium skutchii Maxon =Huperzia myrsinites 

Lycopodium sprucei Baker =Lycopodiella contexta 

Lycopodium tatei A.C. Sm. =Lycopodiella iuliformis var. tatei 

Lycopodium taxifolium Sw. =Huperzia taxifolia 

Lycopodium underwoodianum Maxon =Huperzia capillaris 

Lycopodium wilsonii Underw. & F.E. Lloyd =Huperzia wilsonii 

Palhinhaea capillacea (Spring) Holub =Lycopodiella cernua 

Palhinhaea cernua (L.) Vasc. & Franco =Lycopodiella cernua 

Phlegmariurus taxifolius (Sw.) A. Löve & D. Löve =Huperzia taxifolia 

Pseudolycopodiella caroliniana (L.) Holub =Lycopodiella caroliniana 

Pseudolycopodiella iuliformis (Underw. & F.E. Lloyd) Holub =Lycopodiella iuliformis 

Pseudolycopodiella meridionalis (Underw. & F.E. Lloyd) Holub =Lycopodiella caroliniana var. meridionalis 

Pseudolycopodiella paradoxa (Mart.) Holub =Lycopodiella caroliniana var. paradoxa 

Urostachys commutatus Herter =Huperzia intermedia 

Urostachys hippurideus (H. Christ) Nessel =Huperzia hippuridea 

Urostachys intermedius (Trevis.) Herter =Huperzia intermedia 

Urostachys killipii Herter =Huperzia killipii 

Urostachys linifolius (L.) Herter =Huperzia linifolia 

Urostachys myrsinites (Lam.) Herter =Huperzia myrsinites 

Urostachys reflexus (Lam.) Herter =Huperzia reflexa 

Urostachys roraimensis (Underw. & F.E. Lloyd) Nessel =Huperzia myrsinites 

Urostachys wilsonii (Underw. & F.E. Lloyd) Herter =Huperzia wilsonii 

REFERENCES 

Øllgaard, B. 1987. A revised classification of the Lycopodiaceae s.l. Opera Bot. 92: 153-178. 

Øllgaard, B. 1995. Lycopodiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 190-206. 

Øllgaard, B. & P.G. Windisch. 1987. Sinopse das Licopodiáceas do Brasil. Bradea 5(1): 1-43. 

Selaginella albolineata A.R. Sm. 

Selaginella amazonica Spring 

Selaginella anaclasta Alston ex Crabbe & Jermy 

Selaginella applanata A. Braun 

Selaginella arrecta A.R. Sm. 

SELAGINELLACEAE 

AM 

AM GU 

AM 

AM 

AM BO

20 

Selaginella arthritica Alston 

Selaginella asperula Spring 

Selaginella beitelii A.R. Sm. 

Selaginella brevifolia Baker 

Selaginella breweriana A.R. Sm. 

Selaginella breynii Spring 

Selaginella cabrerensis Hieron. 

Selaginella calceolata Jermy & J.M. Rankin 

Selaginella cardiophylla Valdespino 

Selaginella cladorrhizans A. Braun 

Selaginella coarctata Spring 

Selaginella conduplicata Spring 

Selaginella convoluta (Arn.) Spring 

Selaginella cyclophylla A.R. Sm. 

Selaginella dendricola Jenman 

Selaginella densifolia Spruce ex Hook. 

Selaginella diffusa (C. Presl) Spring 

Selaginella epirrhizos Spring 

Selaginella falcata (P. Beauv.) Spring 

Selaginella flabellata (L.) Spring 

Selaginella flagellata Spring 

Selaginella fragilis A. Braun 

Selaginella hemicardia Valdespino 

Selaginella hirsuta Alston ex Crabbe & Jermy 

Selaginella humboldtiana A. Braun 

Selaginella imbricans A.R. Sm. 

Selaginella karowtipuensis Valdespino, sp. nov. ined. 

Selaginella kochii Hieron. 

Selaginella kunzeana A. Braun 

Selaginella lechleri Hieron. 

Selaginella marahuacae A.R. Sm. 

Selaginella marginata (Humb. & Bonpl. ex Willd.) Spring 

Selaginella mazaruniense Jenman 

Selaginella microdonta A.C. Sm. 

Selaginella minima Spring 

Selaginella muscosa Spring 

Selaginella neblinae A.R. Sm. 

Selaginella novae-hollandiae (Sw.) Spring 

Selaginella pallescens (C. Presl) Spring 

Selaginella palmiformis Alston ex Crabbe & Jermy 

Selaginella parkeri (Hook. & Grev.) Spring 

Selaginella plana (Desv. ex Poir.) Hieron. [introduced] 

Selaginella porelloides (Lam.) Spring 

Selaginella porphyrospora A. Braun 

Selaginella potaroensis Jenman 

Selaginella producta Baker 

Selaginella pruskiana Valdespino 

Selaginella pubens A.R. Sm. 

Selaginella radiata (Aubl.) Spring 

Selaginella revoluta Baker 

Selaginella rhodostachya Baker 

Selaginella roraimensis Baker 

Selaginella sandwithii Alston 

Selaginella scalariformis A.C. Sm. 

Selaginella schultesii Alston ex Crabbe & Jermy 

Selaginella seemannii Baker 

Selaginella smithiorum Valdespino 

Selaginella sobolifera A.R. Sm. 

Selaginella sp. A 

Selaginella substipitata Spring 

Selaginella sulcata (Desv. ex Poir.) Spring ex Mart. 

Selaginella terezoana Bautista 

Selaginella thysanophylla A.R. Sm. 

Selaginella tuberculata Spruce ex Baker 


FG? 

AM 

AM 

AM 

AM 

BO 

BO GU FG 

AM BO 

AM BO 

BO 

DA FG 

AM GU 

AM GU SU FG 

BO GU FG? 

AM 

GU SU FG 

SU? FG? 

SU FG 

DA GU SU FG 

GU? 

SU 

FG 

AM DA GU SU FG 

AM 

AM 

BO 

BO GU 

AM 

AM 

GU 

AM 

AM 

AM 

AM 

SU FG 

AM BO 

GU 

AM BO 

FG 

AM 

AM 

GU 

FG? 

SU 

AM 

AM BO GU 

GU 

SU FG 

BO 

BO 

GU FG 



AM 

AM 

GU 

GU SU FG 


BO GU 

AM BO GU 

GU SU FG 

AM 

AM 

BO GU 

BO 

GU SU FG 

AM 

BO 

GU 

AM BO 

GU SU FG 

BO 

BO 

GU 

BO GU SU FG

Selaginella tyleri A.C. Sm. 

Selaginella umbrosa Lem. ex Hieron. 

Selaginella valdepilosa Baker 

Selaginella vernicosa Baker 

Selaginella versatilis A.R. Sm. 

Selaginella wurdackii Alston 



Selaginella affinis A. Braun =Selaginella epirrhizos 

Selaginella brachyclada Baker =Selaginella rhodostachya 

Selaginella brevispicata Hieron. ex Bautista =Selaginella rhodostachya 

Selaginella calcarata A. Braun =Selaginella conduplicata 

Selaginella caudorhiza Baker =Selaginella sulcata 

Selaginella cayennensis Baker =Selaginella producta 

Selaginella diminutifolia Jenman =Selaginella rhodostachya 

Selaginella duidae A.C. Sm. =Selaginella scalariformis 

Selaginella guianensis Spring =Selaginella falcata 

Selaginella jenmanii Baker =Selaginella porelloides 

Selaginella karsteniana A. Braun =Selaginella substipitata 

Selaginella lemairei Hieron. =Selaginella umbrosa 

Selaginella lucidinervia Spring =Selaginella parkeri 

Selaginella macroclada Baker =Selaginella parkeri 

Selaginella parkeri Spring var. pedata (Klotzsch) Baker =Selaginella parkeri 

Selaginella pedata Klotzsch =Selaginella parkeri 

Selaginella penniformis (Lam.) Hieron. =Selaginella radiata 

Selaginella platyphylla Baker =Selaginella porelloides 

Selaginella platyphylla var. laxa Jenman =Selaginella porelloides 

Selaginella poeppigiana var. guyanensis Spring =Selaginella epirrhizos 

Selaginella puberula Klotzsch =Selaginella coarctata 

Selaginella rigidula Baker =Selaginella substipitata 

Selaginella scintillata Alston =Selaginella roraimensis 

Selaginella stellata Spring =Selaginella conduplicata 

Selaginella valdepilosa subsp. tricholoma Jermy & J.M. Rankin =Selaginella rhodostachya 

Selaginella vernicosa var. oligoclada Baker =Selaginella vernicosa 

REFERENCES 

Alston, A.H.G., A.C. Jermy, & J.M. Rankin. 1981. The genus Selaginella in tropical South America. Bull. Brit. Mus. (Nat. Hist.) 

Bot. 9: 233-330. 

Smith, A.R. 1995. Selaginellaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 296-314. 

PTEROPHYTA 

AM 

AM BO 

AM BO GU 

AM BO GU 

AM 

AM BO 


Berry, P.E., B.K. Holst, & K. Yatskievych. 1995. Flora of the Venezuelan Guayana, vol. 2. Pteridophytes, Spermatophytes 

Acanthaceae - Araceae. Missouri Botanical Garden, St. Louis, & Timber Press, Portland, Oregon. 

Cremers, G. 1997. Pterophyta. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 

Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs 

of the New York Botanical Garden 76, pt. 1: 65-162. 

Cremers, G., C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des espèces de Phanérogames et de Ptéridophytes de la 

Guyane française d’après l’Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne. 

Cremers, G. & M. Hoff. 1990. Inventaire taxonomique des plantes de la Guyane française. I. Les Ptéridophytes. Mus. Nat. Hist. 

Nat., Inventaires de Faune et de Flore 54: 1-133. 

Jenman, G.S. 1909. The ferns and fern allies of the British West Indies and Guiana. Trinidad: Bull. Misc. Int. Bot. Dept. 

Kramer, K.U. 1978. The Pteridophytes of Suriname. An enumeration with keys of the ferns and fern-allies. Uitgaven 

“Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen” Utrecht no. 93. 

Kramer, K.U. & P.S. Green, eds. 1990. The Families and Genera of Vascular Plants. I. Pteridophytes and Gymnosperms. New 

York: Springer-Verlag. 

Lellinger, D.B. 1989. The ferns and fern-allies of Costa Rica, Panama, and the Chocó. Part 1: Psilotaceae through Dicksoniaceae. 

Pteridologia 2A: 1–364. 

Mickel, J.T., and J.M. Beitel. 1988. Pteridophyte flora of Oaxaca, Mexico. Memoirs of the New York Botanical Garden 46: 1– 

568. 

Pichi Sermolli, R.E.G. 1977. Tentamen Pteridophytorum genera in taxonomicum ordinem redigendi. Webbia 31: 313–512. 

Proctor, G.R. 1977. Pteridophytes. Vol. 2 in Flora of the Lesser Antilles, edited by R. A. Howard. Cambridge, Massachusetts: 

Harvard University. 

Proctor, G.R. 1985. Ferns of Jamaica. London: British Museum (Natural History).

22 


Proctor, G.R. 1989. Ferns of Puerto Rico and the Virgin Islands. Memoirs of the New York Botanical Garden 53: 1–389. 

Posthumous, O. 1928. The Ferns of Surinam and of French and British Guiana. In A. Pulle, Flora of Suriname supplement: 1- 

196. 

Smith, A.R. 1981. Pteridophytes. Vol. 2 in Flora of Chiapas, edited by D. E. Breedlove. San Francisco: California Academy of 

Sciences. 

Tryon, R.M. 1972. Endemic areas and geographic speciation in tropical American ferns. Biotropica 4: 121–131. 

Tryon, R.M., and R.G. Stolze. 1989. Pteridophyta of Peru. Part I. 1. Ophioglossaceae—12. Cyatheaceae. Fieldiana, Bot., n.s. 20: 

1–145. 

Tryon, R.M., and A.F. Tryon. 1982. Ferns and Allied Plants with Special Reference to Tropical America. New York: Springer- 

Verlag. 

Vareschi, V. 1968. Helechos. In T. Lasser, Flora de Venezuela 1: 1-1033. 

ADIANTACEAE 

Adiantopsis monticola (Gardner) T. Moore 

Adiantopsis radiata (L.) Fée 

Adiantum adiantoides (J. Sm.) C. Chr. 

Adiantum amazonicum A.R. Sm. 

Adiantum argutum Splitg. 

Adiantum asperum (Poir.) Desv. 

Adiantum cajennense Willd. ex Klotzsch 

Adiantum concinnum Humb. & Bonpl. ex Willd. 

Adiantum cordatum Maxon [specimens ident. as sp. nov. by G. Cremers] 

Adiantum decoratum Maxon & Weath. 

Adiantum deflectens Mart. 

Adiantum fructuosum Poepp. ex Spreng. 

Adiantum fuliginosum Fée 

Adiantum glaucescens Klotzsch 

Adiantum glaziovii Baker 

Adiantum humile Kunze 

Adiantum latifolium Lam. 

Adiantum leprieurii Hook. 

Adiantum lucidum (Cav.) Sw. 

Adiantum lunulatum Burm. f. 

Adiantum macrophyllum Sw. 

Adiantum nudum A.R. Sm. 

Adiantum obliquum Willd. 

Adiantum olivaceum Baker 

Adiantum oyapokense Jenman 

Adiantum paraense Hieron. 

Adiantum petiolatum Desv. 

Adiantum phyllitidis J. Sm. 

Adiantum pulverulentum L. 

Adiantum serratodentatum Humb. & Bonpl. ex Willd. 

Adiantum terminatum Kunze ex Miq. [possibly conspecific with Adiantum 

humile] 

Adiantum tetraphyllum Humb. & Bonpl. ex Willd. 

Adiantum tomentosum Klotzsch 

Adiantum tripteris K.U. Kramer 

Adiantum villosum L. 

Cassebeera pinnata Kaulf. 

Cheilanthes bonariensis (Willd.) Proctor 

Cheilanthes concolor (Langsd. & Fisch.) R.M. Tryon & A.F. Tryon 

Cheilanthes eriophora (Fée) Mett. 

Doryopteris collina (Raddi) J. Sm. 

Doryopteris conformis K.U. Kramer & R.M. Tryon 

Doryopteris cyclophylla A.R. Sm. 

Doryopteris davidsei A.R. Sm. 

Doryopteris lomariacea Kunze ex Klotzsch 

Doryopteris pedata (L.) Fée 

Doryopteris pedata var. multipartita (Fée) R.M. Tryon 

Doryopteris pedata var. palmata (Willd.) Hicken 

Doryopteris rediviva Fée 

Doryopteris sagittifolia (Raddi) J. Sm. 

GU 


GU SU FG 

AM BO 

GU SU 



AM 

FG 

FG 

BO FG 

GU SU FG 


BO GU SU FG 

FG 

AM BO SU FG 


BO GU SU FG 


BO 

BO GU SU FG 

AM BO 


GU 

FG 

AM 






AM GU SU FG 


SU 

FG? 

AM BO 

BO DA 

BO 

AM BO 

GU SU 

BO GU SU 

AM 

AM 

BO GU 

BO GU SU 

GU SU 

BO 

BO 

AM BO GU SU FG


Doryopteris varians (Raddi) J. Sm. 

Eriosorus flexuosus (Kunth) Copel. 

Eriosorus flexuosus var. flexuosus 

Eriosorus hispidulus (Kunze) Vareschi 

Eriosorus hispidulus var. hispidulus 

Eriosorus paucifolius (A.C. Sm.) Vareschi [Mt. Roraima] 

Hemionitis palmata L. 

Hemionitis rufa (L.) Sw. 

Notholaena solitaria R.M. Tryon 

Pityrogramma calomelanos (L.) Link 

Pityrogramma calomelanos var. aureoflava (Hook.) Weath. ex L.H. Bailey 

Pityrogramma calomelanos var. calomelanos 

Pityrogramma ebenea (L.) Proctor 

Pityrogramma ebenea var. ebenea [Mt. Roraima] 

Pityrogramma trifoliata (L.) R.M. Tryon 

Pterozonium brevifrons (A.C. Sm.) Lellinger 

Pterozonium cyclophyllum (Baker) Diels 

Pterozonium cyclosorum A.C. Sm. 

Pterozonium elaphoglossoides (Baker) Lellinger 

Pterozonium lineare Lellinger 

Pterozonium maguirei Lellinger 

Pterozonium paraphysatum (A.C. Sm.) Lellinger 

Pterozonium reniforme (Mart.) Fée 

Pterozonium retroflexum Mickel 

Pterozonium scopulinum Lellinger 

Pterozonium spectabile Maxon & A.C. Sm. 

Pterozonium steyermarkii Vareschi 

Pterozonium tatei A.C. Sm. 

Pterozonium terrestre Lellinger 


Adiantum dolosum Kunze (considered valid in FVG) =Adiantum asperum 

Adiantum guianense Aubl. (Dennstaedtiaceae) =Lindsaea guianensis 

Adiantum hirtum Klotzsch =Adiantum terminatum 

Adiantum lancea L. (Dennstaedtiaceae) =Lindsaea lancea 

Adiantum lunulatum Houtt. (Dryopteridaceae) =Didymochlaena truncatula 

Adiantum philippense L. =Adiantum lunulatum 

Adiantum propinquum Fée =Adiantum pulverulentum 

Adiantum radiatum L. =Adiantopsis radiata 

Adiantum sagittatum Aubl. (Dennstaedtiaceae) =Lindsaea sagittata 

Adiantum strictum Sw. (Dennstaedtiaceae) =Lindsaea stricta 

Cheilanthes glandulosa Sw. (Dennstaedtiaceae) =Paesia glandulosa 

Cheilanthes radiata (L.) J. Sm. =Adiantopsis radiata 

Cheilanthes viscosa H. Karst. [nomen nudum] (Dennstaedtiaceae) =Hypolepis viscosa 

Doryopteris concolor (Langsd. & Fisch.) Kuhn =Cheilanthes concolor 

Doryopteris palmata (Willd.) J. Sm. =Doryopteris pedata var. palmata 

Eriosorus ottonis (Kunze ex Klotzsch) Vareschi =Eriosorus hispidulus var. hispidulus 

Eriosorus paucifolius var. neblinae A.F. Tryon =Eriosorus paucifolius 

Eriosorus paucifolius var. steyermarkii A.F. Tryon =Eriosorus paucifolius 

Eriosorus schomburgkianus (Kunze ex Klotzsch) Copel =Eriosorus hispidulus var. hispidulus 

Gymnogramma cyclophylla Baker =Pterozonium cyclophyllum 

Gymnogramma diplazioides Desv. (Thelypteridaceae) =Thelypteris linkiana 

Gymnogramma elaphoglossoides Baker =Pterozonium elaphoglossoides 

Gymnogramma flexuosa Desv. =Eriosorus flexuosus 

Gymnogramma hispidula (Kunze) Klotzsch =Eriosorus hispidulus 

Gymnogramma ottonis Kunze ex Klotzsch =Eriosorus hispidulus var. hispidulus 

Gymnogramma reniformis Mart. =Pterozonium reniforme 

Gymnogramma rufa (L.) Desv. =Hemionitis rufa 

Gymnogramma schomburgkiana Kunze ex Klotzsch =Eriosorus hispidulus var. hispidulus 

Gymnopteris rufa (L.) Bernh. ex Underw. =Hemionitis rufa 

Gymnopteris semipinnatifida Fée (Lomariopsidaceae) =Bolbitis semipinnatifida 

Hemionitis cajenensis Desv. (Vittariaceae) =Antrophyum cajenense 

Hemionitis lineata Sw. (Vittariaceae) =Antrophyum lineatum 

Jamesonia hispidula Kunze =Eriosorus hispidulus 

Notholaena aurea (Poir.) Desv. =Cheilanthes bonariensis 

GU 

AM BO GU 

AM BO GU 

AM BO GU 

AM BO GU 

AM BO GU? 

GU SU FG 

BO GU SU FG 

SU 


GU 


AM BO 

AM BO 

BO 

AM BO GU 

AM BO GU 

AM BO 

AM BO GU 

AM 

AM 

AM BO GU SU 

AM BO GU 

AM 

AM BO GU 

BO 

BO 

AM 

AM BO

24 


Notholaena eriophora Fée =Cheilanthes eriophora 

Notholaena steyermarkii Vareschi =Cheilanthes eriophora 

Pellaea lomariacea (Kunze ex Klotzsch) Hook. =Doryopteris lomariacea 

Pellaea pinnata (Kaulf.) Prantl (Pteridaceae) =Cassebeera pinnata 

Pityrogramma tartarea (Cav.) Maxon =Pityrogramma ebenea 

Psilogramme paucifolia A.C. Sm. =Eriosorus paucifolius 

Pterozonium reniforme var. ulei Brause =Pterozonium reniforme 

Pterozonium schneei Vareschi =Pterozonium cyclophyllum 

Pterozonium spathulatum Vareschi =Pterozonium cyclophyllum 

Syngramma brevifrons A.C. Sm. =Pterozonium brevifrons 

Syngramma elaphoglossoides (Baker) Diels =Pterozonium elaphoglossoides 

Syngramma paraphysata A.C. Sm. =Pterozonium paraphysatum 

Syngramma versicolor Vareschi =Pterozonium brevifrons 

Syngrammatopsis elaphoglossoides (Baker) Alston =Pterozonium elaphoglossoides 

Syngrammatopsis paraphysata (A.C. Sm.) Alston =Pterozonium paraphysatum 

Trismeria trifoliata (L.) Diels =Pityrogramma trifoliata 

REFERENCES 

Lellinger, D.B. 1967. Pterozonium. In B. Maguire & collaborators, Botany of the Guayana Highland part 7. Mem. New York 

Bot. Gard. 17(1): 1-439. 

Lellinger, D.B. 1972. Polypodiaceae subfam. Adiantoideae. In B. Maguire & collaborators, Botany of the Guayana Highland 

part 9. Mem. New York Bot. Gard. 23: 1-832. 

Smith, A.R. & D.B. Lellinger. 1995. Pteridaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 

2: 250-286. 

Tryon, A.F. 1970. A monograph of the fern genus Eriosorus. Contrib. Gray Herb. 200: 54-174. 

Tryon, R.M. 1942. A revision of the genus Doryopteris. Contrib. Gray Herb. 143: 1-80. 

Asplenium abscissum Willd. 

Asplenium angustum Sw. 

Asplenium auriculatum Sw. 

Asplenium auritum Sw. 

Asplenium chimantae A.R. Sm. 

Asplenium claussenii Hieron. 

Asplenium cowanii A.R. Sm. 

Asplenium cristatum Lam. 

Asplenium cuneatum Lam. 

Asplenium cuspidatum Lam. 

Asplenium delitescens (Maxon) L.D. Gómez 

Asplenium dissectum Sw. 

Asplenium feei Kunze ex Fée 

Asplenium flabellulatum Kunze 

Asplenium formosum Willd. 

Asplenium hallii Hook. 

Asplenium harpeodes Kunze 

Asplenium hastatum Klotzsch ex Kunze 

Asplenium hostmannii Hieron. 

Asplenium juglandifolium Lam. 

Asplenium laetum Sw. 

Asplenium macilentum Kunze ex Klotzsch 

Asplenium pearcei Baker 

Asplenium pedicularifolium A. St.-Hil. 

Asplenium praemorsum Sw. 

Asplenium pteropus Kaulf. 

Asplenium pumilum Sw. 

Asplenium radicans L. 

Asplenium radicans var. cirrhatum (Rich. ex Willd.) Rosenst. 

Asplenium radicans var. radicans 

Asplenium repandulum Kunze 

Asplenium riparium Liebm. 

Asplenium rutaceum (Willd.) Mett. 

Asplenium salicifolium L. 

Asplenium serra Langsd. & Fisch. 

ASPLENIACEAE 

GU FG 


GU 


BO 

AM BO GU? 

AM 



AM 

AM FG 

BO GU 

AM GU SU FG 

AM 

AM BO GU FG 

AM 

AM BO GU 

AM 

GU SU FG 


BO GU SU FG 

SU 

AM GU SU FG 

GU SU FG 

BO GU 

AM BO GU SU 

BO 

AM BO GU FG 

AM BO GU FG 

AM BO GU 

FG 

FG 

AM GU SU FG 


AM BO GU SU

Asplenium serratum L. 

Asplenium sessilifolium Desv. 

Asplenium zamiifolium Willd. 



Asplenium alloeopteron Kunze ex Klotzsch =Asplenium radicans 

Asplenium amazonicum H. Christ =Asplenium pearcei 

Asplenium anisophyllum auct. non Kunze =Asplenium feei 

Asplenium arboreum Willd. (Woodsiaceae) =Diplazium cristatum 

Asplenium asplenioides (A.C. Sm.) Pittier et al. (Hymenophyllopsidaceae) =Hymenophyllopsis asplenioides 

Asplenium auritum var. auriculatum (Hook. f.) C.V. Morton & Lellinger =Asplenium auritum 

Asplenium auritum var. bipinnatifidum Kunze =Asplenium auritum 

Asplenium auritum var. obtusum Kunze ex Mett. =Asplenium macilentum 

Asplenium blechnoides Lag. ex Sw. (Blechnaceae) =Blechnum polypodioides 

Asplenium centripetale Baker (Woodsiaceae) =Diplazium centripetale 

Asplenium cicutarium Sw. =Asplenium cristatum 

Asplenium cirrhatum Rich. ex Willd. =Asplenium radicans var. cirrhatum 

Asplenium cyrtopteron Kunze =Asplenium radicans 

Asplenium dejectum (Baker) Pittier (Hymenophyllopsidaceae) =Hymenophyllopsis dejecta 

Asplenium dimidiatum auct. non Sw. =Asplenium zamiifolium 

Asplenium fragrans Sw. =Asplenium cuspidatum 

Asplenium grandifolium Sw. (Woodsiaceae) =Diplazium grandifolium 

Asplenium haplophyllum Domin =Asplenium pearcei 

Asplenium integerrimum Spreng. =Asplenium juglandifolium 

Asplenium kapplerianum Kunze =Asplenium juglandifolium 

Asplenium lechleri Mett. (Woodsiaceae) =Diplazium lechleri 

Asplenium marginatum L. (Woodsiaceae) =Hemidictyum marginatum 

Asplenium marinum Hook. f. =Asplenium auritum 

Asplenium perkinsii Jenman =Asplenium rutaceum 

Asplenium plantaginifolium L. (Woodsiaceae) =Diplazium plantaginifolium 

Asplenium pseudoerectum Hieron. =Asplenium pteropus 

Asplenium sanguinolentum Kunze ex Mett. =Asplenium feei 

Asplenium schomburgkianum Klotzsch =Asplenium serratum 

Asplenium semicordatum Raddi =Asplenium salicifolium 

Asplenium spruceanum Hieron. =Asplenium hallii 

Asplenium sulcatum auct. non Lam. =Asplenium auritum 

Asplenium wilsonii Baker (Woodsiaceae) =Diplazium wilsonii 

REFERENCES 

(Included in Pteridaceae in Flora of the Venezuelan Guayana) 

Morton, C.V. & D.B. Lellinger. 1966. The Polypodiaceae subfam. Asplenioideae in Venezuela. Mem. New York Bot. Gard. 15: 

1-49. 

Smith, A.R. 1995. Aspleniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 12-22. 

BLECHNACEAE 

Blechnum asplenioides Sw. 

Blechnum atropurpureum A.R. Sm. 

Blechnum auratum (Fée) R.M. Tryon & Stolze 

Blechnum binervatum (Poir.) C.V. Morton & Lellinger 

Blechnum brasiliense Desv. 

Blechnum x caudatum Cav. 

Blechnum cordatum (Desv.) Hieron. 

Blechnum divergens (Kunze) Mett. 

Blechnum fraxineum Willd. 

Blechnum gracile Kaulf. 

Blechnum lanceola Sw. 

Blechnum lehmannii Hieron. 

Blechnum lherminieri (Bory) C. Chr. 

Blechnum lherminieri subsp. lherminieri 

Blechnum lineatum (Sw.) C. Chr. 

Blechnum occidentale L. 

Blechnum polypodioides Raddi 

Blechnum proliferum Rosenst. [Possibly misident. B. violaceum in the 

Guianas] 


FG? 

AM GU SU FG 

BO GU SU FG 

AM BO 

AM 

AM 

BO GU 

BO 

AM BO 

AM GU SU FG 

GU? FG 

GU SU FG 

SU 

AM BO GU 

GU 

GU 

GU 


BO GU SU FG 

GU?

26 

Blechnum schomburgkii (Klotzsch) C. Chr. 

Blechnum serrulatum Rich. 

Blechnum stipitellatum (Sodiro) C. Chr. 

Blechnum violaceum (Fée) C. Chr. 

Salpichlaena hookeriana (Kuntze) Alston 

Salpichlaena volubilis (Kaulf.) J. Sm. 



Blechnum x antillanum Proctor =Blechnum x caudatum 

Blechnum arborescens (Klotzsch & H. Karst.) Hieron =Blechnum cordatum 

Blechnum blechnoides (Lag. ex Sw.) C. Chr. =Blechnum polypodioides 

Blechnum buchtienii Rosenst. =Blechnum auratum 

Blechnum ensiforme (Liebm.) C. Chr. =Blechnum binervatum 

Blechnum exaltatum (Fée) Duek =Blechnum divergens 

Blechnum indicum auct. non Burm. f. =Blechnum serrulatum 

Blechnum maxonii (Broadh.) C. Chr. =Blechnum lehmannii 

Blechnum ornifolium (C. Presl) Ettingsh. =Blechnum cordatum 

Blechnum unilaterale Sw. [nom. superfl.] =Blechnum polypodioides 

Blechnum volubile Kaulf. =Salpichlaena volubilis 

Blechnum volubile var. lomarioidea Baker =Salpichlaena hookeriana 

Lomaria arborescens Klotzsch & H. Karst. =Blechnum cordatum 

Lomaria aurata Fée =Blechnum auratum 

Lomaria cordata Desv. =Blechnum cordatum 

Lomaria divergens Kunze =Blechnum divergens 

Lomaria ensiformis Liebm. =Blechnum binervatum 

Lomaria exaltata Fée =Blechnum divergens 

Lomaria ornifolia C. Presl =Blechnum cordatum 

Lomaria schomburgkii Klotzsch =Blechnum schomburgkii 

Lomaria stipitellata Sodiro =Blechnum stipitellatum 

Lomaria violacea Fée =Blechnum violaceum 

Salpichlaena lomarioidea (Baker) A.R. Sm. =Salpichlaena hookeriana 

Spicanta hookeriana Kuntze =Salpichlaena hookeriana 

Stenochlaena fendleri (D.C. Eaton) Underw. (Lomariopsidaceae) =Lomariopsis fendleri 

Stenochlaena japurensis (Mart.) L. Underw. (Lomariopsidaceae) =Lomariopsis japurensis 

Stenochlaena marginata auct. non (Schrad.) C. Chr. (Lomariopsidaceae) =Lomariopsis japurensis 

Struthiopteris maxonii Broadh. =Blechnum lehmannii 

REFERENCES 

Smith, A.R. 1995. Blechnaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 23-29. 

CYATHEACEAE 

Alsophila cuspidata (Kunze) D.S. Conant 

Alsophila imrayana (Hook.) D.S. Conant 

Alsophila imrayana (Hook.) D.S. Conant var. basilaris (H. Christ) Lellinger 

Alsophila miquelii Kunze 

Cnemidaria cruciata (Desv.) Stolze 

Cnemidaria horrida (L.) C. Presl 

Cnemidaria roraimensis (Domin) R.M. Tryon 

Cnemidaria spectabilis (Kunze) R.M. Tryon 

Cnemidaria spectabilis var. spectabilis 

Cyathea akawaiorum P.J. Edwards 

Cyathea andina (H. Karst.) Domin 

Cyathea bipinnatifida (Baker) Domin 

Cyathea caracasana (Klotzsch) Domin [Mt. Roraima] 

Cyathea cyatheoides (Desv.) K.U. Kramer 

Cyathea cyclodium (Tryon) Lellinger 

Cyathea delgadii Sternb. 

Cyathea demissa (C.V. Morton) A.R. Sm. ex Lellinger 

Cyathea dissimilis (C.V. Morton) Stolze 

Cyathea divergens Kunze 

Cyathea divergens var. divergens 

Cyathea fulva (M. Martens & Galeotti) Fée 

Cyathea gibbosa (Klotzsch) Domin 

Cyathea infesta (Kunze) Domin 

Cyathea intramarginalis (P.G. Windisch) Lellinger 

AM BO GU 


BO GU 

AM GU 

AM GU SU FG 


FG 

AM BO SU FG 

SU FG 

SU? 

GU FG 

AM 

AM BO GU 



GU 


AM BO GU 

BO GU? 


AM 

BO GU 

AM BO 

BO GU 

AM BO GU SU 

AM BO GU SU 

AM BO 

GU 

FG 

BO

Cyathea kalbreyeri (Baker) Domin 

Cyathea lasiosora (Kuhn) Domin 

Cyathea lechleri Mett. 

Cyathea liesneri A.R. Sm. 

Cyathea macrocarpa (C. Presl) Domin 

Cyathea macrosora (Baker) Domin 

Cyathea macrosora var. macrosora 

Cyathea macrosora var. vaupensis (P.G. Windisch) A.R. Sm. 

Cyathea marginalis (Klotzsch) Domin 

Cyathea microdonta (Desv.) Domin 

Cyathea multiflora Sm. 

Cyathea nanna (Barrington) Lellinger 

Cyathea neblinae A.R. Sm. 

Cyathea platylepis (Hook.) Domin 

Cyathea praeceps A.R. Sm. 

Cyathea pubens Domin 

Cyathea pungens (Willd.) Domin 

Cyathea schlimii (Kuhn) Domin 

Cyathea simplex R.M. Tryon 

Cyathea sipapoensis (R.M. Tryon) Lellinger 

Cyathea steyermarkii R.M. Tryon 

Cyathea surinamensis (Miq.) Domin 

Cyathea thysanolepis (Barrington) A.R. Sm. 

Cyathea traillii (Baker) Domin 

Cyathea ulei (H. Christ) Domin 

Cyathea villosa Humb. & Bonpl. ex Willd. 

Cyathea williamsii (Maxon) Domin 



Alsophila bipinnatifida Baker =Cyathea bipinnatifida 

Alsophila blechnoides Hook. (Metaxyaceae) =Metaxya rostrata 

Alsophila caracasana Klotzsch =Cyathea caracasana 

Alsophila demissa C.V. Morton =Cyathea demissa 

Alsophila fulva M. Martens & Galeotti =Cyathea fulva 

Alsophila gibbosa Klotzsch =Cyathea gibbosa 

Alsophila gleasonii Maxon =Cyathea pungens 

Alsophila kalbreyeri Baker =Cyathea kalbreyeri 

Alsophila lasiosora Mett. ex Kuhn =Cyathea lasiosora 

Alsophila lechleri Mett. =Cyathea ulei 

Alsophila macrosora Baker =Cyathea macrosora 

Alsophila marginalis Klotzsch =Cyathea marginalis 

Alsophila microdonta (Desv.) Desv. =Cyathea microdonta 

Alsophila nigra Mart. =Cyathea lasiosora 

Alsophila oblonga Klotzsch =Cyathea pungens 

Alsophila phalerata var. infesta Kunze =Cyathea infesta 

Alsophila podophylla Baker =Cyathea kalbreyeri 

Alsophila procera (Willd.) Desv. =Cyathea pungens 

Alsophila pubescens Baker =Cyathea bipinnatifida 

Alsophila pungens (Willd.) C. Presl =Cyathea pungens 

Alsophila schlimii Kuhn =Cyathea schlimii 

Alsophila subaculeata Splitg. =Cyathea pungens 

Alsophila ulei H. Christ =Cyathea ulei 

Alsophila vestita C. Presl =Cyathea spp. 

Alsophila villosa (Humb. & Bonpl.) Desv. =Cyathea villosa 

Alsophila williamsii Maxon =Cyathea williamsii 

Cyathea arborea auct. non (L.) Sm. =Alsophila imrayana 

Cyathea aurea Klotzsch =Cyathea fulva 

Cyathea boconensis H. Karst. =Cyathea caracasana 

Cyathea calva H. Karst. =Cyathea divergens var. divergens 

Cyathea caracasana var. meridensis (H. Karst.) R.M. Tryon =Cyathea caracasana 

Cyathea cuspidata Kunze =Alsophila cuspidata 

Cyathea horrida (L.) Sm. =Cnemidaria horrida 

Cyathea imrayana Hook. =Alsophila imrayana 

Cyathea macrosora var. reginae (P.G. Windisch) A.R. Sm. =Cyathea traillii 

Cyathea meridensis H. Karst. =Cyathea caracasana 

BO 

BO FG 

AM BO 

AM 

AM GU SU FG 

AM BO GU 

AM BO 

BO 



GU 

GU 

AM BO 

AM BO GU 

AM BO 

GU 


BO 

AM GU? 

BO 

AM 


BO 

AM GU 

AM BO 

BO GU 

AM

28 


Cyathea oblonga (Klotzsch) Domin =Cyathea pungens 

Cyathea oyapoka Jenman =Alsophila cuspidata 

Cyathea petiolulata H. Karst. =Cyathea divergens var. divergens 

Cyathea procera (Willd.) Domin =Cyathea pungens 

Cyathea purpurea C.V. Morton =Cyathea lechleri 

Cyathea roraimensis (Domin) Domin =Cnemidaria roraimensis 

Cyathea spectabilis (Kunze) Domin =Cnemidaria spectabilis 

Cyathea squamipes H. Karst. =Cyathea caracasana 

Cyathea subtropica Domin =Cyathea ulei 

Cyathea surukunensis V. Marcano =Cyathea kalbreyeri 

Cyathea vestita Mart. =Cyathea delgadii 

Hemitelia andina H. Karst. =Cyathea andina 

Hemitelia boryana Mett. ex Kuhn =Cyathea andina 

Hemitelia cyatheoides Desv. =Cyathea cyatheoides 

Hemitelia dissimilis C.V. Morton =Cyathea dissimilis 

Hemitelia escuquensis H. Karst. =Cyathea andina 

Hemitelia hirsuta (Desv.) Weath. =Cyathea surinamensis 

Hemitelia horrida (L.) R. Br. =Cnemidaria horrida 

Hemitelia hostmannii Hook. =Cyathea cyatheoides 

Hemitelia macrocarpa C. Presl =Cyathea macrocarpa 

Hemitelia macrosora (Baker) Jenman =Cyathea macrosora 

Hemitelia minima C.V. Morton =Cyathea platylepis 

Hemitelia multiflora (Desv.) Spreng. =Cyathea cyatheoides 

Hemitelia multiflora (Desv.) Spreng. var. sprucei Baker =Cyathea cyatheoides 

Hemitelia parkeri Hook. =Cyathea surinamensis 

Hemitelia platylepis Hook. =Cyathea platylepis 

Hemitelia roraimensis Domin =Cnemidaria roraimensis 

Hemitelia spectabilis Kunze =Cnemidaria spectabilis 

Hemitelia stigmosa Desv. =Cyathea cyatheoides 

Hemitelia surinamensis Miq. =Cyathea surinamensis 

Nephelea imrayana (Hook.) R.M. Tryon =Alsophila imrayana 

Sphaeropteris cyatheoides (Desv.) P.G. Windisch =Cyathea cyatheoides 

Sphaeropteris dissimilis (C.V. Morton) P.G. Windisch =Cyathea dissimilis 

Sphaeropteris hirsuta (Desv.) R.M. Tryon =Cyathea surinamensis 

Sphaeropteris intramarginalis P.G. Windisch =Cyathea intramarginalis 

Sphaeropteris macrocarpa (C. Presl) R.M. Tryon =Cyathea macrocarpa 

Sphaeropteris macrosora (Baker) P.G. Windisch =Cyathea macrosora 

Sphaeropteris macrosora var. reginae P.G. Windisch =Cyathea traillii 

Sphaeropteris macrosora var. vaupensis P.G. Windisch =Cyathea macrosora var. vaupensis 

Sphaeropteris marginalis (Klotzsch) R.M. Tryon =Cyathea marginalis 

Sphaeropteris sipapoensis R.M. Tryon =Cyathea sipapoensis 

Trichipteris cyclodium R.M. Tryon =Cyathea cyclodium 

Trichipteris demissa (C.V. Morton) R.M. Tryon =Cyathea demissa 

Trichipteris demissa var. thysanolepis Barrington =Cyathea thysanolepis 

Trichipteris kalbreyeri (Baker) R.M. Tryon =Cyathea kalbreyeri 

Trichipteris lechleri (Mett.) R.M. Tryon =Cyathea ulei 

Trichipteris maguirei R.M. Tryon =Cyathea williamsii 

Trichipteris microdonta (Desv.) R.M. Tryon =Cyathea microdonta 

Trichipteris nigra (Mart.) R.M. Tryon =Cyathea lasiosora 

Trichipteris procera (Willd.) R.M. Tryon =Cyathea pungens 

Trichipteris pubescens (Baker) R.M. Tryon =Cyathea bipinnatifida 

Trichipteris schlimii (Kuhn) Barrington =Cyathea schlimii 

Trichipteris villosa (Humb. & Bonpl. ex Willd.) R.M. Tryon =Cyathea villosa 

Trichipteris williamsii (Maxon) R.M. Tryon =Cyathea williamsii 

REFERENCES 

Conant, D.C. 1983. A revision of the genus Alsophila in the Americas. Journ. Arn. Arb. 64: 333-382. 

Smith, A.R. 1995. Cyatheaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 30-43. 

Stolze, R.G. 1974. A taxonomic revision of the genus Cnemidaria. Fieldiana Bot. 37: 1-98. 

Tryon, R.M. 1970. The classification of the Cyatheaceae. Contrib. Gray Herb. 200: 3-53. 

Tryon, R.M. 1976. A revision of the genus Cyathea. Contrib. Gray Herb. 206: 19-101. 

Windisch, P.G. 1977. Synopsis of the genus Sphaeropteris, with a revision of the neotropical exindusiate species. Bot. Jahrb. 

Syst. 98(2): 176-198.

Rumohra adiantiformis (G. Forst.) Ching 


DAVALLIACEAE 


Davallia domingensis Spreng. (Dennstaedtiaceae) =Saccoloma domingense 

Davallia inaequalis Kunze (Dennstaedtiaceae) =Saccoloma inaequale 

Davallia meifolia Kunth (Dennstaedtiaceae) =Lindsaea meifolia 

Davallia parkeri Hook. (Dennstaedtiaceae) =Lindsaea parkeri 

REFERENCES 

Cremers, G., K.U. Kramer, R.C. Moran, & A.R. Smith. 1994. Dryopteridaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas 

(series B) 6: 3-65. 

DENNSTAEDTIACEAE 

Blotiella lindeniana (Hook.) R.M. Tryon 

Dennstaedtia bipinnata (Cav.) Maxon 

Dennstaedtia dissecta (Sw.) T. Moore 

Dennstaedtia globulifera (Poir.) Hieron. 

Histiopteris incisa (Thunb.) J. Sm. 

Hypolepis guianensis Klotzsch [Mt. Roraima] 

Hypolepis hostilis (Kunze) C. Presl 

Hypolepis nigrescens Hook. [Mt. Roraima] 

Hypolepis parallelogramma (Kunze) C. Presl 

Hypolepis repens (L.) C. Presl 

Hypolepis sp. 1 [Mt. Roraima] 



Hypolepis sp. A 

Hypolepis viscosa Mett. [Cheilanthes] 

Lindsaea arcuata Kunze 

Lindsaea bolivarensis V. Marcano 

Lindsaea coarctata K.U. Kramer 

Lindsaea cultriformis K.U. Kramer 

Lindsaea cyclophylla K.U. Kramer 

Lindsaea divaricata Klotzsch 

Lindsaea dubia Spreng. 

Lindsaea falciformis Hook. [Mt. Roraima] [taxon of uncertain status] 

Lindsaea guianensis (Aubl.) Dryand. 

Lindsaea guianensis subsp. guianensis 

Lindsaea hemiptera K.U. Kramer 

Lindsaea javitensis Humb. & Bonpl. ex Willd. 

Lindsaea klotzschiana Moritz ex Ettingsh. 

Lindsaea lancea (L.) Bedd. 

Lindsaea lancea var. elatior (Kunze) K.U. Kramer 

Lindsaea lancea var. falcata (Dryand.) Rosenst. 

Lindsaea lancea var. lancea 

Lindsaea lancea var. leprieurii (Hook.) K.U. Kramer 

Lindsaea lancea var. remota (Kunze) K.U. Kramer 

Lindsaea lancea var. submontana Boudrie & Cremers 

Lindsaea macrophylla Kaulf. 

Lindsaea mazarunensis Jenman [species of uncertain status, probably a hybrid] 

Lindsaea meifolia (Kunth) Mett. ex Kuhn 

Lindsaea mesarum K.U. Kramer 

Lindsaea pallida Klotzsch 

Lindsaea parkeri (Hook.) Kuhn 

Lindsaea parkeri subsp. parkeri 

Lindsaea parkeri subsp. steyermarkiana K.U. Kramer 

Lindsaea pendula Klotzsch 

Lindsaea pleioptera K.U. Kramer 

Lindsaea portoricensis Desv. 

Lindsaea pumila Klotzsch [dubious species] 

Lindsaea quadrangularis Raddi 

BO GU 

AM BO 

BO 


GU 

AM BO GU 

GU 

GU SU FG 

AM GU 

GU FG 

GU SU FG 

GU 

GU 

GU? 

AM 

AM BO 

AM 

BO 

GU 

AM BO GU 

AM BO GU 



GU 



AM 

AM BO GU 

AM BO 





AM GU SU FG 


FG 

GU 

FG 

AM 

AM BO 

BO GU SU FG 


BO GU SU FG 

AM BO 

AM BO GU SU 

AM BO GU 


GU 

GU SU FG

30 


Lindsaea quadrangularis subsp. antillensis K.U. Kramer 

Lindsaea reniformis Dryand. 

Lindsaea rigidiuscula Lindm. 

Lindsaea sagittata (Aubl.) Dryand. 

Lindsaea schomburgkii Klotzsch 

Lindsaea semilunata (C. Chr.) C. Chr. 

Lindsaea sphenomeridopsis K.U. Kramer 

Lindsaea stenomeris K.U. Kramer 

Lindsaea stricta (Sw.) Dryand. 

Lindsaea stricta var. jamesoniiformis K.U. Kramer 

Lindsaea stricta var. parvula (Fée) K.U. Kramer 

Lindsaea stricta var. stricta 

Lindsaea surinamensis Posth. 

Lindsaea taeniata K.U. Kramer 

Lindsaea tenuis Klotzsch 

Lindsaea tetraptera K.U. Kramer 

Lindsaea ulei Hieron. 

Lonchitis hirsuta L. 

Microlepia speluncae (L.) T. Moore [introduced, fide Lellinger] 

Odontosoria fumarioides (Sw.) J. Sm. 

Ormoloma imrayanum (Hook.) Maxon [cited from FG in error] 

Paesia acclivis (Kunze) Kuhn 

Paesia glandulosa (Sw.) Kuhn 

Pteridium arachnoideum (Kaulf.) Maxon 

Pteridium caudatum (L.) Maxon 

Saccoloma domingense (Spreng.) C. Chr. 

Saccoloma elegans Kaulf. 

Saccoloma elegans subsp. chartaceum Nair ex Cremers & K.U. Kramer 

Saccoloma inaequale (Kunze) Mett. 


Allosorus acclivis Kunze =Paesia acclivis 

Dennstaedtia adiantoides (Willd.) T. Moore =Dennstaedtia dissecta 

Dennstaedtia obtusifolia (Willd.) T. Moore =Dennstaedtia dissecta 

Ithycaulon inaequale (Kunze) Copel. =Saccoloma inaequale 

Lindsaea aquatica Jenman =Lindsaea stricta var. parvula 

Lindsaea coriifolia Lindm. =Lindsaea schomburgkii 

Lindsaea crenata Klotzsch =Lindsaea portoricensis 

Lindsaea elegans Fée [nom. illeg.] =Lindsaea klotzschiana 

Lindsaea falcata Dryand. =Lindsaea lancea var. falcata 

Lindsaea falcata var. elatior Kunze =Lindsaea lancea var. elatior 

Lindsaea falcata var. lancea (L.) Jenman =Lindsaea lancea var. lancea 

Lindsaea falcata var. leprieurii (Hook.) Jenman =Lindsaea lancea var. leprieurii 

Lindsaea falcata var. subrotundifolia Jenman [name of uncertain application] 

Lindsaea feei C. Chr. =Lindsaea klotzschiana 

Lindsaea filiformis Hook. =Lindsaea tenuis 

Lindsaea gracilis Klotzsch =Lindsaea stricta var. parvula 

Lindsaea guianensis var. venosa Jenman [name of uncertain application] 

Lindsaea horizontalis Hook. =Lindsaea arcuata 

Lindsaea krameriana V. Marcano =Lindsaea mesarum 

Lindsaea lancea var. longifolia K.U. Kramer =Lindsaea lancea var. elatior 

Lindsaea lancea var. semilunata C. Chr. =Lindsaea semilunata 

Lindsaea leprieurii Hook. =Lindsaea lancea var. leprieurii 

Lindsaea nervosa Lindm. [nom. illeg.] =Lindsaea rigidiuscula 

Lindsaea nitidissima Rich. ex Willd. =Lindsaea lancea 

Lindsaea parvula Fée =Lindsaea stricta var. parvula 

Lindsaea pumila var. remota Kunze =Lindsaea lancea var. remota 

Lindsaea pusilla Splitg. =Lindsaea lancea var. remota 

Lindsaea raddiana Klotzsch =Lindsaea stricta 

Lindsaea rigescens Willd. =Lindsaea stricta 

Lindsaea rufescens Kunze =Lindsaea portoricensis 

Lindsaea schomburgkii f. coriifolia (Lindm.) K.U. Kramer =Lindsaea schomburgkii 

Lindsaea schomburgkii f. densa K.U. Kramer =Lindsaea schomburgkii 

Lindsaea trapeziformis Dryand. =Lindsaea lancea 

Lindsaea trapeziformis var. erythromera Kunze =Lindsaea divaricata 

GU SU FG 

GU SU FG 

AM BO 


AM BO GU 

AM BO GU 

AM GU 

AM BO 





AM 

GU SU FG 

BO GU 

AM BO GU 

AM BO GU 

SU FG 

FG 

FG? 

AM 

BO 

BO 

GU 

AM BO SU 

AM 

BO 

GU SU FG 

GU FG 

GU FG 

AM BO GU SU FG


Lonchitis lindeniana Hook. =Blotiella lindeniana 

Lonchitis pedata L. (Pteridaceae) =Pteris podophylla 

Odontoloma parkeri (Hook.) C. Presl =Lindsaea parkeri 

Orthiopteris domingensis (Spreng.) Copel. =Saccoloma domingense 

Orthiopteris inaequalis (Kunze) Copel. =Saccoloma inaequale 

Paesia viscosa A. St.-Hil. =Paesia glandulosa 

Pteridium aquilinum var. arachnoideum (Kaulf.) Brade =Pteridium arachnoideum 

Pteridium aquilinum subsp. caudatum (L.) Bonap. =Pteridium caudatum 

Saccoloma imrayana Hook. =Ormoloma imrayanum 

Saccoloma imrayanum Hook. =Ormoloma imrayanum 

Schizoloma macrophyllum (Kaulf.) C. Presl =Lindsaea macrophylla 

Schizoloma reniforme (Dryand.) Diels =Lindsaea reniformis 

Schizoloma sagittatum (Aubl.) Diels =Lindsaea sagittata 

Stenoloma gratissima Fée =Lindsaea meifolia 

REFERENCES 

Cremers, G. & K.U. Kramer. 1991. Dennstaedtiaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 4: 20-81. 

Kramer, K.U. 1957. A revision of the genus Lindsaea in the New World. Acta Bot. Neerl. 6: 97-290. 

Smith, A.R. & K.U. Kramer. 1995. Dennstaedtiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 2: 46-71. 

Culcita coniifolia (Hook.) Maxon 

DICKSONIACEAE 


Dicksonia adiantoides Humb. & Bonpl. ex Willd. (Dennstaedtiaceae) =Dennstaedtia bipinnata 

Dicksonia bipinnata Cav. (Dennstaedtiaceae) =Dennstaedtia bipinnata 

Dicksonia coniifolia Hook. =Culcita coniifolia 

Dicksonia dissecta Sw. (Dennstaedtiaceae) =Dennstaedtia dissecta 

Dicksonia obtusifolia Willd. (Dennstaedtiaceae) =Dennstaedtia dissecta 

REFERENCES 

Maxon, W.R. 1922. The genus Culcita. J. Wash. Acad. Sci. 12: 454-460. 

Smith, A.R. 1995. Dicksoniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 72-73. 

DRYOPTERIDACEAE 

Arachniodes denticulata (Sw.) Ching [Mt. Roraima] 

Arachniodes leucostegioides (C. Chr.) Ching [Mt. Roraima] 

Arachniodes macrostegia (Hook.) R.M. Tryon & D.S. Conant [Mt. Roraima] 

Arachniodes ochropteroides (Baker) Lellinger 

Cyclodium akawaiorum A.R. Sm. [Mt. Roraima] 

Cyclodium calophyllum (C.V. Morton) A.R. Sm. 

Cyclodium guianense (Klotzsch) van der Werff ex L.D. Gómez 

Cyclodium inerme (Fée) A.R. Sm. 

Cyclodium meniscioides (Willd.) C. Presl 

Cyclodium meniscioides var. meniscioides 

Cyclodium meniscioides var. rigidissimum (C. Chr.) A.R. Sm. 

Cyclodium rheophilum A.R. Sm. 

Cyclodium varians (Fée) A.R. Sm. 

Didymochlaena truncatula (Sw.) J. Sm. 

Olfersia cervina (L.) Kunze 

Polybotrya caudata Kunze 

Polybotrya cyathifolia Fée 

Polybotrya fractiserialis (Baker) Sm. 

Polybotrya glandulosa Mett. ex Kuhn 

Polybotrya lechleriana Mett. [Mt. Roraima] 

Polybotrya osmundacea Humb. & Bonpl. ex Willd. 

Polybotrya sessilisora R.C. Moran [Brazil/Guyana border area] 

Polybotrya sorbifolia Mett. ex Kuhn [expected in the Guianas] 

Polystichum platylepis Fée 

Polystichum platyphyllum (Willd.) C. Presl 

Polystichum sp. A 

Stigmatopteris longicaudata (Liebm.) C. Chr. 

AM BO GU 

AM BO GU 

AM GU 

AM BO GU? 

AM BO GU SU 

BO GU 

BO GU? 



AM BO? DA GU SU FG 


BO? GU 

FG 

BO GU 




FG 

GU SU FG 

AM 

GU 

AM GU SU FG 

BO GU? 

AM BO 

GU? 

AM 

BO 

AM FG

32 

Stigmatopteris rotundata (Humb. & Bonpl. ex Willd.) C. Chr. 


BO GU SU FG 


Aspidium articulatum Sw. (Oleandraceae) =Oleandra articulata 

Aspidium biolleyi H. Christ (Thelypteridaceae) =Thelypteris biolleyi 

Aspidium biserratum Sw. (Oleandraceae) =Nephrolepis biserrata 

Aspidium capense Willd. (Davalliaceae) =Rumohra adiantiformis 

Aspidium coarctatum Kunze (Thelypteridaceae) =Thelypteris opposita 

Aspidium confertum Kaulf. =Cyclodium meniscioides var. meniscioides 

Aspidium crenulans Fée (Tectariaceae) =Megalastrum crenulans 

Aspidium dicksonioides Fée (Tectariaceae) =Triplophyllum dicksonioides 

Aspidium extensum Blume (Thelypteridaceae) =Thelypteris opulenta 

Aspidium funestum Kunze (Tectariaceae) =Triplophyllum funestum 

Aspidium gongylodes Schkuhr (Thelypteridaceae) =Thelypteris interrupta 

Aspidium guianense Klotzsch =Cyclodium guianense 

Aspidium hispidulum Decne. (Thelypteridaceae) =Thelypteris hispidula 

Aspidium lizarzaburui (Sodiro) C. Chr. (Tectariaceae) =Tectaria lizarzaburui 

Aspidium martinicense Spreng. (Tectariaceae) =Tectaria incisa 

Aspidium meniscioides Willd. =Cyclodium meniscioides 

Aspidium nodosum Willd. (Oleandraceae) =Oleandra articulata 

Aspidium opulentum Kaulf. (Thelypteridaceae) =Thelypteris opulenta 

Aspidium pectinatum Willd. (Oleandraceae) =Nephrolepis pectinata 

Aspidium pendulum Raddi (Oleandraceae) =Nephrolepis cordifolia 

Aspidium pendulum Splitg., non Raddi 1819 (Oleandraceae) =Oleandra pilosa 

Aspidium plantagineum (Jacq.) Griseb. (Tectariaceae) =Tectaria plantaginea 

Aspidium platyphyllum Willd. =Polystichum platyphyllum 

Aspidium plumieri C. Presl (Tectariaceae) =Tectaria trifoliata 

Aspidium rostratum Kunth (Metaxyaceae) =Metaxya rostrata 

Aspidium rotundatum Humb. & Bonpl. ex Willd. =Stigmatopteris rotundata 

Aspidium rutaceum Willd. (Aspleniaceae) =Asplenium rutaceum 

Aspidium schomburgkii Klotzsch (Tectariaceae) =Ctenitis falciculata 

Aspidium sprengelii Kaulf. (Thelypteridaceae) =Thelypteris balbisii 

Aspidium truncatulum Sw. =Didymochlaena truncatula 

Cyclodium confertum (Kaulf.) C. Presl =Cyclodium meniscioides var. meniscioides 

Cyclodium rigidissimum C. Chr. =Cyclodium meniscioides var. rigidissimum 

Dryopteris amplissima (C. Presl) Kuntze (Tectariaceae) =Lastreopsis amplissima 

Dryopteris amplissima var. subeffusa C. Chr. (Tectariaceae) =Lastreopsis amplissima 

Dryopteris angustifolia (Willd.) Urb. (Thelypteridaceae) =Thelypteris angustifolia 

Dryopteris arborea Brause (Thelypteridaceae) =Thelypteris arborea 

Dryopteris balbisii (Spreng.) Urb. (Thelypteridaceae) =Thelypteris balbisii 

Dryopteris brachypoda (Baker) C. Chr. (Thelypteridaceae) =Thelypteris brachypoda 

Dryopteris calophylla C.V. Morton =Cyclodium calophyllum 

Dryopteris carrii auct. non (Baker) C. Chr. =Stigmatopteris rotundata 

Dryopteris chrysodioides (Fée) Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris chrysodioides 

Dryopteris chrysodioides var. goyazensis Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris chrysodioides var. 

goyazensis 

Dryopteris crenulans (Fée) C. Chr. (Tectariaceae) =Megalastrum crenulans 

Dryopteris decussata (L.) Urb. (Thelypteridaceae) =Thelypteris decussata 

Dryopteris dejecta (Jenman) C. Chr. (Thelypteridaceae) =Thelypteris glandulosa 

Dryopteris demerarana (Baker) C. Chr. (Thelypteridaceae) =Thelypteris demerarana 

Dryopteris densiloba C. Chr. [nom. illeg.] (Thelypteridaceae) =Thelypteris gardneriana 

Dryopteris dentata auct. non (Forssk.) C. Chr. (Thelypteridaceae) =Thelypteris hispidula 

Dryopteris denticulata (Sw.) Kuntze =Arachniodes denticulata 

Dryopteris denticulata var. aperta C. Chr. =Arachniodes denticulata 

Dryopteris effusa (Sw.) Urb. (Tectariaceae) =Lastreopsis effusa 

Dryopteris engelii Hieron. (Thelypteridaceae) =Thelypteris rudis 

Dryopteris extensa (Blume) Kuntze (Thelypteridaceae) =Thelypteris opulenta 

Dryopteris funesta (Kunze) C. Chr. (Tectariaceae) =Triplophyllum funestum 

Dryopteris gongylodes (Schkuhr) Kuntze (Thelypteridaceae) =Thelypteris interrupta 

Dryopteris gonophora Weath. (Thelypteridaceae) =Thelypteris gonophora 

Dryopteris guianensis (Klotzsch) Posth. =Cyclodium guianense 

Dryopteris hostmannii Klotzsch (Thelypteridaceae) =Thelypteris arborescens 

Dryopteris kanukuana Weath. (Thelypteridaceae) =Thelypteris pennata 

Dryopteris killipii C. Chr. & Maxon (Tectariaceae) =Lastreopsis killipii 

Dryopteris latiuscula Maxon =Thelypteris abrupta


Dryopteris leprieurii (Hook.) Kuntze (Thelypteridaceae) =Thelypteris leprieurii 

Dryopteris leucostegioides C. Chr. =Arachniodes leucostegioides 

Dryopteris lindigii C. Chr. (Thelypteridaceae) =Thelypteris deflexa 

Dryopteris linkiana (C. Presl) Maxon (Thelypteridaceae) =Thelypteris linkiana 

Dryopteris longicaudata (Liebm.) Maxon =Stigmatopteris longicaudata 

Dryopteris macrophylla (Kunze) C. Chr. (Thelypteridaceae) =Thelypteris macrophylla 

Dryopteris macrostegia (Hook.) Kuntze =Arachniodes macrostegia 

Dryopteris martinii C. Chr. (Thelypteridaceae) [not in flora area, type apparently erroneously cited from FG] =Thelypteris 

connexa 

Dryopteris megalodus (Schkuhr) Urb. (Thelypteridaceae) =Thelypteris pennata 

Dryopteris meniscioides (Willd.) Kuntze =Cyclodium meniscioides 

Dryopteris meniscioides var. conferta (Kaulf.) C.V. Morton =Cyclodium meniscioides var. meniscioides 

Dryopteris mercurii A. Braun ex Hieron. (Thelypteridaceae) =Thelypteris balbisii 

Dryopteris nephrodioides var. biolleyi (H. Christ) C. Chr. (Thelypteridaceae) =Thelypteris biolleyi 

Dryopteris nesiotica Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris nesiotica 

Dryopteris ochropteroides (Baker) C. Chr. =Arachniodes ochropteroides 

Dryopteris oligocarpa (Humb. & Bonpl. ex Willd.) Kuntze (Thelypteridaceae) =Thelypteris oligocarpa 

Dryopteris opposita (Vahl) Urb. (Thelypteridaceae) =Thelypteris opposita 

Dryopteris poiteana (Bory) Urb. (Thelypteridaceae) =Thelypteris poiteana 

Dryopteris protensa var. dicksonioides (Fée) C. Chr. (Tectariaceae) =Triplophyllum dicksonioides 

Dryopteris protensa var. funesta (Kunze) C. Chr. (Tectariaceae) =Triplophyllum funestum 

Dryopteris pteroidea (Klotzsch) C. Chr. (Thelypteridaceae) =Thelypteris pteroidea 

Dryopteris pyramidata (Fée) Maxon (Thelypteridaceae) =Thelypteris abrupta 

Dryopteris quadrangularis (Fée) Alston (Thelypteridaceae) =Thelypteris hispidula 

Dryopteris refulgens (Klotzsch ex Mett.) C. Chr. (Tectariaceae) =Ctenitis refulgens 

Dryopteris rolandii C. Chr. (Thelypteridaceae) =Thelypteris x rolandii 

Dryopteris roraimensis Brause (Thelypteridaceae) =Thelypteris arborea 

Dryopteris rotundata (Humb. & Bonpl. ex Willd.) C. Chr. =Stigmatopteris rotundata 

Dryopteris rudis (Kunze) C. Chr. (Thelypteridaceae) =Thelypteris rudis 

Dryopteris sancti-gabrieli (Hook.) Kuntze =Cyclodium guianense 

Dryopteris serrata (Cav.) C. Chr. (Thelypteridaceae) =Thelypteris serrata 

Dryopteris sorbifolia auct. non (Jacq.) Hieron. (Thelypteridaceae) =Thelypteris arborescens 

Dryopteris subcuneata (Baker) Kuntze (Thelypteridaceae) =Thelypteris abrupta 

Dryopteris subincisa (Willd.) Urb. (Tectariaceae) =Megalastrum subincisum 

Dryopteris subobliquata (Hook.) Kuntze =Cyclodium inerme 

Dryopteris tetragona (Sw.) Urb. (Thelypteridaceae) =Thelypteris tetragona 

Dryopteris tristis (Kunze) Kuntze (Thelypteridaceae) =Thelypteris tristis 

Dryopteris varians (Fée) Kuntze =Cyclodium varians 

Nephrodium antioquianum Baker (Tectariaceae) =Tectaria antioquioana 

Nephrodium aureoviridum Jenman [dubious species; =Thelypteris tetragona, Thelypteridaceae?] 

Nephrodium brachypodum Baker (Thelypteridaceae) =Thelypteris brachypoda 

Nephrodium connexum Kuhn ex Baker (Thelypteridaceae) [not in flora area, type erroneously cited from FG] =Thelypteris 

connexa 

Nephrodium deflexum C. Presl (Thelypteridaceae) =Thelypteris deflexa 

Nephrodium dejectum Jenman (Thelypteridaceae) =Thelypteris glandulosa 

Nephrodium gardnerianum Baker (Thelypteridaceae) =Thelypteris gardneriana 

Nephrodium leprieurii Hook. (Thelypteridaceae) =Thelypteris leprieurii 

Nephrodium lizarzaburui Sodiro (Tectariaceae) =Tectaria lizarzaburui 

Nephrodium macrostegium Hook. =Arachniodes macrostegia 

Nephrodium ochropteroides Baker =Arachniodes ochropteroides 

Nephrodium quadrangulare Fée (Thelypteridaceae) =Thelypteris hispidula 

Nephrodium subcuneatum Baker (Thelypteridaceae) =Thelypteris abrupta 

Nephrodium subfuscum Baker (Thelypteridaceae) =Thelypteris germaniana 

Nephrodium varians Fée =Cyclodium varians 

Polybotrya aristeguietae Brade =Polybotrya osmundacea 

Polybotrya cervina (L.) Kaulf. =Olfersia cervina 

Polybotrya pubens auct. non Mart. =Polybotrya caudata 

Polybotrya salicifolia Lellinger =Polybotrya sorbifolia 

Polybotrya subelliptica Lellinger =Polybotrya glandulosa 

Polybotrya vareschii Brade =Polybotrya osmundacea 

Polybotrya villosula H. Christ =Polybotrya caudata 

Polystichopsis ochropteroides (Baker) C.V. Morton =Arachniodes ochropteroides 

Polystichum adiantiforme (G. Forst.) J. Sm. (Davalliaceae) =Rumohra adiantiformis 

Polystichum amplissimum C. Presl (Tectariaceae) =Lastreopsis amplissima 

Polystichum capense (Willd.) J. Sm. (Davalliaceae) =Rumohra adiantiformis

34 


Polystichum inerme Fée =Cyclodium inerme 

Stigmatopteris guianensis (Klotzsch) C. Chr. =Cyclodium guianense 

Stigmatopteris meniscioides (Willd.) K.U. Kramer =Cyclodium meniscioides 

Stigmatopteris sancti-gabrieli (Hook.) C. Chr. =Cyclodium guianense 

Stigmatopteris varians (Fée) Alston =Cyclodium varians 

REFERENCES 


(series B) 6: 3-65. 

Moran, R.C. 1986. The Neotropical fern genus Olfersia. Amer. Fern J. 76: 161-178. 

Moran, R.C. 1987. Monograph of the Neotropical fern genus Polybotrya (Dryopteridaceae). Ill. Nat. Hist. Surv. Bull. 34(1): 1- 

138. 

Moran, R.C. 1991. Monograph of the Neotropical fern genus Stigmatopteris (Dryopteridaceae). Ann. Missouri Bot. Gard. 78(4): 

857-914. 

Smith, A.R. 1986. Revision of the Neotropical fern genus Cyclodium. Am. Fern J. 76(2): 56-98. 

Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopteridaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the 

Venezuelan Guayana 2: 73-128. 

GLEICHENIACEAE 

Dicranopteris flexuosa (Schrad.) Underw. 

Dicranopteris nervosa (Kaulf.) Maxon 

Dicranopteris pectinata (Willd.) Underw. 

Dicranopteris schomburgkiana (J.W. Sturm) C.V. Morton [Mt. Roraima] 

Diplopterygium bancroftii (Hook.) A.R. Sm. 

Sticherus bifidus (Willd.) Ching 

Sticherus compactus (H. Christ) Nakai 

Sticherus hypoleucus (Sodiro) Copel. 

Sticherus longipinnatus (Hook.) Ching 

Sticherus melanoblastus (Alston) qsterg. & B. qllg. 

Sticherus nervatus J. Gonzales 

Sticherus pallescens (Mett.) Vareschi 

Sticherus penniger (Mart.) Copel. 

Sticherus remotus (Kaulf.) Chrysler 

Sticherus revolutus (Kunth) Ching [Mt. Roraima] 

Sticherus sp. A 

Sticherus tepuiensis A.R. Sm. 

Sticherus tomentosus (Cav. ex Sw.) A.R. Sm. 


Dicranopteris bifida (Willd.) Maxon =Sticherus bifidus 

Dicranopteris nitidula (Rosenst.) Suess. ex Losch =Sticherus nitidulus 

Dicranopteris remota (Kaulf.) Maxon =Sticherus remotus 

Gleichenia bancroftii Hook. =Diplopterygium bancroftii 

Gleichenia bifida (Willd.) Spreng. =Sticherus bifidus 

Gleichenia hypoleuca Sodiro =Sticherus hypoleucus 

Gleichenia linearis auct. non (Burm.) C.B. Clarke =Dicranopteris flexuosa 

Gleichenia longipinnata Hook. =Sticherus longipinnatus 

Gleichenia melanoblasta Alston 

Gleichenia nitidula Rosenst. =Sticherus nitidulus 

Gleichenia pallescens Mett. =Sticherus pallescens 

Gleichenia pectinata (Willd.) C. Presl =Dicranopteris pectinata 

Gleichenia pennigera (Mart.) T. Moore =Sticherus penniger 

Gleichenia remota (Kaulf.) Spreng. =Sticherus remotus 

Gleichenia revoluta Kunth =Sticherus revolutus 

Gleichenia tomentosa (Cav. ex Sw.) Spreng. =Sticherus tomentosus 

Hicriopteris bancroftii (Hook.) Ching =Diplopterygium bancroftii 

Mertensia bifida Willd. =Sticherus bifidus 

Mertensia flexuosa Schrad. =Dicranopteris flexuosa 

Mertensia pectinata Willd. =Dicranopteris pectinata 

Mertensia pennigera Mart. =Sticherus penniger 

Mertensia pubescens Humb. & Bonpl. ex Willd. =Sticherus bifidus 

Mertensia remota Kaulf. =Sticherus remotus 

Mertensia schomburgkiana J.W. Sturm =Dicranopteris schomburgkiana 

Mertensia tomentosa Cav. ex Sw. =Sticherus tomentosus 


GU 


AM BO 

AM BO 


GU 

GU 

GU SU 

GU 

GU 

AM 

BO 


AM BO GU 

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AM BO GU


Mertensia velata Kunze =Sticherus tomentosus 

Sticherus nitidulus (Rosenst.) Copel. [not in flora area] 

Sticherus pubescens (Humb. & Bonpl. ex Willd.) Nakai =Sticherus bifidus 

Sticherus velatus (Kunze) Copel. =Sticherus tomentosus 

REFERENCES 

Smith, A.R. 1995. Gleicheniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 128-135. 

GRAMMITIDACEAE 

Ceradenia arthrothrix L.E. Bishop & A.R. Sm. 

Ceradenia capillaris (Desv.) L.E. Bishop [Mt. Roraima] 

Ceradenia discolor (Hook.) L.E. Bishop [Mt. Roraima] 

Ceradenia fragillima (Copel.) L.E. Bishop [Mt. Roraima] 

Ceradenia jungermannioides (Klotzsch) L.E. Bishop 

Ceradenia kalbreyeri (Baker) L.E. Bishop 

Ceradenia kookenamae (Jenman) L.E. Bishop [Mt. Roraima] 

Ceradenia microcystis L.E. Bishop & A.R. Sm. 

Ceradenia pruinosa (Maxon) L.E. Bishop 

Ceradenia spixiana (Mart. ex Mett.) L.E. Bishop 

Cochlidium attenuatum A.C. Sm. 

Cochlidium connellii (Baker ex C.H. Wright) A.C. Sm. 

Cochlidium furcatum (Hook. & Grev.) C. Chr. 

Cochlidium linearifolium (Desv.) Maxon ex C. Chr. 

Cochlidium rostratum (Hook.) Maxon ex C. Chr. 

Cochlidium seminudum (Willd.) Maxon 

Cochlidium serrulatum (Sw.) L.E. Bishop 

Cochlidium tepuiense (A.C. Sm.) L.E. Bishop 

Cochlidium wurdackii L.E. Bishop 

Enterosora campbellii Baker 

Enterosora campbellii subsp. campbellii 

Enterosora parietina (Klotzsch) L.E. Bishop 

Enterosora trifurcata (L.) L.E. Bishop 

Grammitis anfractuosa (Kunze ex Klotzsch) Proctor [=Melpomene 

anfractuosa] 

Grammitis apiculata (Kunze ex Klotzsch) F. Seym. 

Grammitis asplenifolia (L.) Proctor [=Terpsichore asplenifolia] 

Grammitis blanchetii (C. Chr.) A.R. Sm. [=Micropolypodium nanum] 

Grammitis bryophila (Maxon) F. Seym. 

Grammitis caucana (Hieron.) C.V. Morton 

Grammitis cultrata (Bory ex Willd.) Proctor [=Terpsichore cultrata] 

Grammitis firma (J. Sm.) C.V. Morton [=Melpomene firma] 

Grammitis flabelliformis (Poir.) C.V. Morton [Mt. Roraima] [=Melpomene 

flabelliformis] 

Grammitis kegeliana (Kunze) Lellinger [=Terpsichore kegeliana] 

Grammitis leptopoda (C.H. Wright) Copel. [Mt. Roraima] 

Grammitis liesneri A.R. Sm. 

Grammitis limula (H. Christ) L.D. Gómez 

Grammitis major (Copel.) C.V. Morton 

Grammitis melanosticta (Kunze) F. Seym. [=Melpomene melanosticta] 

Grammitis melanotrichia (Baker) Lellinger [Mt. Roraima] [=Lellingeria 

melanotrichia] 

Grammitis mollissima (Fée) Proctor [=Terpsichore mollissima] 

Grammitis moniliformis (Lag. ex Sw.) Proctor [=Melpomene moniliformis] 

Grammitis myosuroides (Sw.) Sw. [Mt. Roraima] [=Lellingeria myosuroides] 

Grammitis paramicola L.E. Bishop [Mt. Roraima] 

Grammitis peritimundi L.E. Bishop & A.R. Sm. [Mt. Roraima] 

Grammitis phlegmaria (J. Sm.) Proctor 

Grammitis phlegmaria var. phlegmaria 

Grammitis pilosissima (M. Martens & Galeotti) C.V. Morton [=Melpomene 

pilosissima] 

Grammitis plicata A.R. Sm. 

Grammitis randallii (Maxon) Proctor 

Grammitis shaferi (Maxon) Lellinger [=Lellingeria shaferi] 

AM 

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BO 

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AM BO GU 

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AM BO GU 



GU? 

GU? FG? 


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BO 

AM BO GU FG? 

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36 


Grammitis sinuosa A.R. Sm. [=Lellingeria sinuosa] 

Grammitis sp. A 

Grammitis staheliana (Posth.) Lellinger [=Terpsichore staheliana] 

Grammitis subsessilis (Baker) C.V. Morton [=Lellingeria subsessilis] 

Grammitis suspensa (L.) Proctor [=Lellingeria suspensa] 

Grammitis taenifolia (Jenman) Proctor [=Micropolypodium taenifolium] 

Grammitis taxifolia (L.) Proctor [=Terpsichore taxifolia] 

Grammitis tegetiformis L.E. Bishop 

Grammitis tenuicula (Fée) Proctor 

Grammitis trichomanoides (Sw.) Ching [=Micropolypodium trichomanoides] 

Grammitis truncicola (Klotzsch) C.V. Morton 

Grammitis xanthotrichia (Klotzsch) Duek & Lellinger [Mt. Roraima] 

[=Terpsichore xanthotrichia] 

Grammitis xiphopteroides (Liebm.) A.R. Sm. 

Zygophlebia sectifrons (Kunze ex Mett.) L.E. Bishop 

AM BO GU SU 

BO 

BO SU FG 

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BO 

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Ctenopteris anfractuosa (Kunze ex Klotzsch) Copel. =Grammitis anfractuosa 

Ctenopteris apiculata (Kunze ex Klotzsch) Copel. =Grammitis apiculata 

Ctenopteris asplenifolia (L.) Copel. =Grammitis asplenifolia 

Ctenopteris capillaris (Desv.) Copel. =Ceradenia capillaris 

Ctenopteris cultrata (Bory ex Willd.) Copel. =Grammitis cultrata 

Ctenopteris firma J. Sm. =Grammitis firma 

Ctenopteris fragillima Copel. =Ceradenia fragillima 

Ctenopteris jubiformis (Kaulf.) J. Sm. =Grammitis suspensa 

Ctenopteris kaieteura (Jenman) Copel. =Grammitis tenuicula 

Ctenopteris kalbreyeri (Baker) Copel. =Ceradenia kalbreyeri 

Ctenopteris kegeliana (Kunze) K.U. Kramer =Grammitis kegeliana 

Ctenopteris kookenamae (Jenman) Copel. =Ceradenia kookenamae 

Ctenopteris major Copel. =Grammitis major 

Ctenopteris melanosticta (Kunze) Copel. =Grammitis melanosticta 

Ctenopteris melanotrichia (Baker) Copel. =Grammitis melanotrichia 

Ctenopteris mollissima (Fée) Copel. =Grammitis mollissima 

Ctenopteris moniliformis (Lag. ex Sw.) J. Sm. =Grammitis moniliformis 

Ctenopteris phlegmaria (J. Sm.) Copel. =Grammitis phlegmaria 

Ctenopteris pilosissima (M. Martens & Galeotti) Copel. =Grammitis pilosissima 

Ctenopteris pruinosa (Maxon) Copel. =Ceradenia pruinosa 

Ctenopteris rigens (Maxon) Copel. =Grammitis xiphopteroides 

Ctenopteris rigescens (Bory ex Willd.) J. Sm. =Grammitis flabelliformis 

Ctenopteris staheliana (Posth.) K.U. Kramer =Grammitis staheliana 

Ctenopteris subsessilis (Baker) Copel. =Grammitis subsessilis 

Ctenopteris suspensa (L.) Copel. =Grammitis suspensa 

Ctenopteris taxifolia (L.) Copel. =Grammitis taxifolia 

Ctenopteris tenuicula (Fée) Copel. =Grammitis tenuicula 

Ctenopteris trifurcata (L.) Pic. Serm. =Enterosora trifurcata 

Ctenopteris xanthotrichia (Klotzsch) Copel. =Grammitis xanthotrichia 

Enterosora integra Lellinger & C.V. Morton =Enterosora campbellii subsp. campbellii 

Glyphotaenium campbellii (Baker) Copel. =Enterosora campbellii subsp. campbellii 

Glyphotaenium integrum (Lellinger & C.V. Morton) Lellinger =Enterosora campbellii subsp. campbellii 

Glyphotaenium trifurcatum (L.) Lellinger =Enterosora trifurcata 

Grammitis campbellii (Baker) Proctor =Enterosora campbellii 

Grammitis capillaris (Desv.) Proctor =Ceradenia capillaris 

Grammitis dicranophylla (C. Chr.) Vareschi =Cochlidium furcatum 

Grammitis discolor (Hook.) C.V. Morton =Ceradenia discolor 

Grammitis flexuosa Kunth (Adiantaceae) =Eriosorus flexuosus 

Grammitis fragillima (Copel.) C.V. Morton =Ceradenia fragillima 

Grammitis furcata Hook. & Grev. =Cochlidium furcatum 

Grammitis haplophlebica (A.C. Sm.) Vareschi =Grammitis bryophila 

Grammitis jamesonii (Hook.) C.V. Morton =Grammitis myosuroides 

Grammitis jubiformis (Kaulf.) Proctor =Grammitis suspensa 

Grammitis jungermannioides (Klotzsch) Ching =Ceradenia jungermannioides 

Grammitis kaieteura (Jenman) C.V. Morton =Grammitis tenuicula 

Grammitis kalbreyeri (Baker) C.V. Morton =Ceradenia kalbreyeri 

Grammitis kirkbridei Mickel =Ceradenia pruinosa 

Grammitis leptodon (C.H. Wright) Vareschi [incorrect spelling, combination in Grammitis previously made] =Grammitis


leptopoda 

Grammitis linearifolia (Desv.) Steud. =Cochlidium linearifolium 

Grammitis linkiana C. Presl (Thelypteridaceae) =Thelypteris linkiana 

Grammitis nana Fée =Grammitis blanchetii 

Grammitis parietina (Klotzsch) Fée =Enterosora parietina 

Grammitis pruinosa (Maxon) C.V. Morton =Ceradenia pruinosa 

Grammitis revoluta Spreng. ex Willd. (Polypodiaceae) =Pleopeltis astrolepis 

Grammitis rigens (Maxon) Proctor =Grammitis xiphopteroides 

Grammitis rigescens (Bory ex Willd.) Lellinger =Grammitis flabelliformis 

Grammitis sectifrons (Kunze ex Mett.) F. Seym. =Zygophlebia sectifrons 

Grammitis serrulata (Sw.) Sw. =Cochlidium serrulatum 

Grammitis sprucei (Hook.) J. Sm. =Enterosora parietina 

Grammitis tatei Copel. =Cochlidium attenuatum 

Grammitis tepuiensis (A.C. Sm.) Vareschi =Cochlidium tepuiense 

Grammitis transiens (Lindm.) F. Seym. =Ceradenia spixiana 

Grammitis trifurcata (L.) Copel. =Enterosora trifurcata 

Grammitis yarumalensis (Hieron.) Proctor =Enterosora parietina 

Lellingeria apiculata (Kunze ex Klotzsch) A.R. Sm. & R.C. Moran =Grammitis apiculata 

Lellingeria limula (H. Christ) A.R. Sm. & R.C. Moran =Grammitis limula 

Lellingeria major (Copel.) A.R. Sm. & R.C. Moran =Grammitis major 

Lellingeria melanotrichia (Baker) A.R. Sm. & R.C. Moran =Grammitis melanotrichia 

Lellingeria myosuroides (Sw.) A.R. Sm. & R.C. Moran =Grammitis myosuroides 

Lellingeria shaferi (Maxon) A.R. Sm. & R.C. Moran =Grammitis shaferi 

Lellingeria sinuosa (A.R. Sm.) A.R. Sm. & R.C. Moran =Grammitis sinuosa 

Lellingeria subsessilis (Baker) A.R. Sm. & R.C. Moran =Grammitis subsessilis 

Lellingeria suspensa (L.) A.R. Sm. & R.C. Moran =Grammitis suspensa 

Lellingeria tenicula A.R. Sm. & R.C. Moran =Grammitis tenuicula 

Melpomene anfractuosa (Kunze ex Klotzsch) A.R. Sm. & R.C. Moran =Grammitis anfractuosa 

Melpomene firma (J. Sm.) A.R. Sm. & R.C. Moran =Grammitis firma 

Melpomene flabelliformis (Poir.) A.R. Sm. & R.C. Moran =Grammitis flabelliformis 

Melpomene melanosticta (Kunze) A.R. Sm. & R.C. Moran =Grammitis melanosticta 

Melpomene moniliformis (Lag. ex Sw.) A.R. Sm. & R.C. Moran =Grammitis moniliformis 

Melpomene pilosissima (M. Martens & Galeotti) A.R. Sm. & R.C. Moran =Grammitis pilosissima 

Melpomene xiphopteroides (Liebm.) A.R. Sm. & R.C. Moran =Grammitis xiphopteroides 

Micropolypodium caucanum (Hieron.) A.R. Sm. =Grammitis caucana 

Micropolypodium liesneri (A.R. Sm.) A.R. Sm. =Grammitis liesneri 

Micropolypodium nanum (Fée) A.R. Sm. =Grammitis blanchetii 

Micropolypodium plicatum (A.R. Sm.) A.R. Sm. =Grammitis plicata 

Micropolypodium taenifolium (Jenman) A.R. Sm. =Grammitis taenifolia 

Micropolypodium trichomanoides (Sw.) A.R. Sm. =Grammitis trichomanoides 

Micropolypodium truncicola (Klotzsch) A.R. Sm. =Grammitis truncicola 

Monogramma linearifolia Desv. =Cochlidium linearifolium 

Nanopteris caucana (Hieron.) Vareschi [nom. inval.] =Grammitis caucana 

Nanopteris limula (H. Christ) Pic. Serm. =Grammitis limula 

Nanopteris nana (Fée) Vareschi [nom. inval.] =Grammitis blanchetii 

Nanopteris taenifolia (Jenman) Vareschi [nom. inval.] =Grammitis taenifolia 

Nanopteris truncicola (Klotzsch) Vareschi =Grammitis truncicola 

Terpsichore asplenifolia (L.) A.R. Sm. =Grammitis asplenifolia 

Terpsichore cultrata (Bory ex Willd.) A.R. Sm. =Grammitis cultrata 

Terpsichore kegeliana (Kunze) A.R. Sm. =Grammitis kegeliana 

Terpsichore mollissima (Fée) A.R. Sm. =Grammitis mollissima 

Terpsichore staheliana (Posth.) A.R. Sm. =Grammitis staheliana 

Terpsichore taxifolia (L.) A.R. Sm. =Grammitis taxifolia 

Terpsichore xanthotrichia (Klotzsch) A.R. Sm. =Grammitis xanthotrichia 

Xiphopteris auyantepuiensis Vareschi =Cochlidium serrulatum 

Xiphopteris duale (Maxon) Vareschi =Cochlidium serrulatum 

Xiphopteris jamesonii Hook. =Grammitis myosuroides 

Xiphopteris limula (H. Christ) Pic. Serm. =Grammitis limula 

Xiphopteris myosuroides (Sw.) Kaulf. =Grammitis myosuroides 

Xiphopteris nana (Fée) Copel. =Grammitis blanchetii 

Xiphopteris rigescens (Bory ex Willd.) Alston =Grammitis flabelliformis 

Xiphopteris serrulata (Sw.) Kaulf. =Cochlidium serrulatum 

Xiphopteris shaferi (Maxon) K.U. Kramer =Grammitis shaferi 

Xiphopteris strictissima (Hook.) Vareschi =Grammitis myosuroides 

Xiphopteris taenifolia (Jenman) Copel. =Grammitis taenifolia

38 


Xiphopteris trichomanoides (Sw.) Copel. =Grammitis trichomanoides 

Xiphopteris truncicola (Klotzsch) Copel. =Grammitis truncicola 

REFERENCES 

Bishop, L.E. 1977. The American species of Grammitis sect. Grammitis. Amer. Fern J. 67: 101-106. 

Bishop, L.E. 1978. Revision of the genus Cochlidium. Amer. Fern J. 68: 76-94. 

Bishop, L.E. 1988. Ceradenia, a new genus of Grammitidaceae. Amer. Fern J. 78: 1-5. 

Bishop, L.E. & A.R. Smith. 1992. Revision of the fern genus Enterosora (Grammitidaceae) in the New World. Syst. Bot. 17: 

345-362. 

Bishop, L.E. & A.R. Smith. 1995. Grammitidaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 2: 135-158. 

Copeland, E.B. 1952. The American species of Xiphopteris. Amer. Fern J. 42: 41-52, 93-110. 

Smith, A.R. 1992. A review of the fern genus Micropolypodium (Grammitidaceae). Novon 2(4): 419-425. 

Smith, A.R. 1993. Terpsichore, a new genus of Grammitidaceae (Pteridophyta). Novon 3(4): 478-489. 

Smith, A.R. & R.C. Moran. 1992. Melpomene, a new genus of Grammitidaceae (Pteridophyta). Novon 2(4): 426-432. 

Smith, A.R., R.C. Moran, & L.E. Bishop. 1991. Lellingeria, a new genus of Grammitidaceae. Amer. Fern J. 81: 76-88. 

Hymenophyllum angustum Bosch 

Hymenophyllum antillense (Jenman) Jenman [Mt. Roraima] 

Hymenophyllum apiculatum Mett. ex Kuhn 

Hymenophyllum asplenioides (Sw.) Sw. 

Hymenophyllum axillare Sw. 

Hymenophyllum brevifrons Kunze 

Hymenophyllum decurrens (Jacq.) Sw. 

Hymenophyllum dependens C.V. Morton 

Hymenophyllum elegans Spreng. 

Hymenophyllum fendlerianum J.W. Sturm 

Hymenophyllum fucoides (Sw.) Sw. 

Hymenophyllum funckii Bosch 

Hymenophyllum hirsutum (L.) Sw. 

Hymenophyllum jamesonii Hook. 

Hymenophyllum kaieteurum Jenman 

Hymenophyllum karstenianum J.W. Sturm 

Hymenophyllum lanatum Fée 

Hymenophyllum lehmannii Hieron. 

Hymenophyllum lindenii Hook. 

Hymenophyllum microcarpum Desv. [Mt. Roraima] 

Hymenophyllum nanostellatum Lellinger 

Hymenophyllum plumieri Hook. & Grev. 

Hymenophyllum polyanthos (Sw.) Sw. 

Hymenophyllum roraimense C.V. Morton [Mt. Roraima] 

Hymenophyllum sericeum (Sw.) Sw. 

Hymenophyllum sp. A 

Hymenophyllum sp. B 

Hymenophyllum trichomanoides Bosch 

Hymenophyllum trichophyllum Kunth 

Hymenophyllum undulatum (Sw.) Sw. 

Hymenophyllum valvatum Hook. & Grev. [Mt. Roraima] 

Trichomanes accedens C. Presl 

Trichomanes angustifrons (Fée) Wess. Boer 

Trichomanes ankersii C. Parker ex Hook. & Grev. 

Trichomanes anomalum Maxon & C.V. Morton 

Trichomanes arbuscula Desv. 

Trichomanes bancroftii Hook. & Grev. 

Trichomanes bicorne Hook. 

Trichomanes botryoides Kaulf. 

Trichomanes caliginum Lellinger 

Trichomanes capillaceum L. 

Trichomanes cellulosum Klotzsch 

Trichomanes crinitum Sw. 

Trichomanes crispum L. 

Trichomanes cristatum Kaulf. 

HYMENOPHYLLACEAE 

AM BO 

BO 

AM BO GU SU 

BO GU 

AM BO 

AM FG 

GU SU FG 

AM BO GU 

AM BO GU 

AM BO GU SU 

AM BO GU SU 

FG? 


AM 

GU 

AM BO 

SU 

BO GU 

AM 

AM BO GU 

BO 

AM 


BO 

GU? 

BO 

BO 

AM BO GU 

AM BO GU 

AM 

BO GU 


GU SU FG 


BO GU SU? 


GU 

AM BO GU 

AM GU SU FG 

GU SU FG 

AM 

AM BO GU SU? 

AM BO GU 


AM BO GU SU? FG


Trichomanes dactylites Sodiro [Mt. Roraima] 

Trichomanes diaphanum Kunth 

Trichomanes diversifrons (Bory) Mett. ex Sadeb. 

Trichomanes egleri P.G. Windisch 

Trichomanes ekmanii Wess. Boer [Guianas records due to misidentification 

fide Lellinger] 

Trichomanes elegans Rich. 

Trichomanes fimbriatum Backh. ex T. Moore [Guianas records due to 

misidentification fide Lellinger] 

Trichomanes guidoi P.G. Windisch 

Trichomanes hostmannianum (Klotzsch) Kunze 

Trichomanes humboldtii (Bosch) Lellinger 

Trichomanes hymenophylloides Bosch [Mt. Roraima] 

Trichomanes jenmanii Lellinger 

Trichomanes kapplerianum J.W. Sturm 

Trichomanes krausii Hook. & Grev. 

Trichomanes longifolium Desv. 

Trichomanes lucens Sw. [Mt. Roraima] 

Trichomanes macilentum Bosch 

Trichomanes martiusii C. Presl 

Trichomanes membranaceum L. 

Trichomanes mougeotii Bosch 

Trichomanes osmundoides DC. ex Poir. 

Trichomanes ovale (E. Fourn.) Wess. Boer 

Trichomanes pedicellatum Desv. 

Trichomanes pellucens Kunze 

Trichomanes pinnatinervium Jenman 

Trichomanes pinnatum Hedw. 

Trichomanes plumosum Kunze 

Trichomanes polypodioides L. 

Trichomanes punctatum Poir. 

Trichomanes punctatum subsp. labiatum (Jenman) Wess. Boer 

Trichomanes pyxidiferum L. 

Trichomanes radicans Sw. 

Trichomanes radicans var. radicans 

Trichomanes resinosum R.C. Moran 

Trichomanes rigidum Sw. 

Trichomanes robustum E. Fourn. 

Trichomanes roraimense Jenman 

Trichomanes rupestre (Raddi) Bosch 

Trichomanes spruceanum Hook. 

Trichomanes sprucei Baker 

Trichomanes steyermarkii P.G. Windisch & A.R. Sm. [Mt. Roraima] 

Trichomanes tanaicum Hook. ex J.W. Sturm 

Trichomanes trollii Bergdolt 

Trichomanes tuerckheimii H. Christ 

Trichomanes vandenboschii P.G. Windisch 

Trichomanes vittaria DC. ex Poir. 


Davalliopsis elegans (Rich.) Copel. =Trichomanes elegans 

Didymoglossum krausii (Hook. & Grev.) C. Presl =Trichomanes krausii 

Didymoglossum membranaceum (L.) Vareschi =Trichomanes membranaceum 

Didymoglossum ovale E. Fourn. =Trichomanes ovale 

Didymoglossum palmarum Vareschi =Trichomanes kapplerianum 

Didymoglossum punctatum (Poir.) Desv. =Trichomanes punctatum 

Feea boryi Bosch, nom. superfl. =Trichomanes diversifrons 

Feea botryoides (Kaulf.) Bosch =Trichomanes botryoides 

Feea humboldtii Bosch =Trichomanes humboldtii 

Feea nana Bory =Trichomanes botryoides 

Feea trollii (Bergdolt) Vareschi =Trichomanes trollii 

Homoeotes heterophylla (Humb. & Bonpl. ex Willd.) C. Presl =Trichomanes humboldtii 

Hymenophyllum amoenum (Klotzsch) J.W. Sturm =Hymenophyllum fendlerianum 

Hymenophyllum ciliatum (Sw.) Sw. =Hymenophyllum hirsutum 

AM BO 



AM BO GU SU 

AM 


AM BO GU? SU? 

AM BO 


AM BO GU 

AM BO GU? 

GU 


BO GU SU FG 

GU 

AM BO GU? 

AM BO GU 

AM BO GU SU 


SU FG 

GU FG 

AM SU 

BO GU SU FG 


GU SU FG 


GU 

AM GU SU FG 



FG 



BO GU 


AM BO FG 

AM BO GU SU? 

FG 

BO GU 

AM BO GU SU 

AM BO GU 

AM BO GU 



AM BO GU SU 

AM GU SU FG

40 


Hymenophyllum ciliatum var. ornifolium Kunze =Hymenophyllum hirsutum 

Hymenophyllum dejectum Baker (Hymenophyllopsidaceae) =Hymenophyllopsis dejecta 

Hymenophyllum lineare var. antillense Jenman =Hymenophyllum antillense 

Hymenophyllum myriocarpum Hook. [records due to misidentification fide Lellinger] 

Hymenophyllum protrusum Hook. [records due to misidentification fide Lellinger] 

Hymenophyllum schomburgkii C. Presl ex J.W. Sturm =Hymenophyllum decurrens 

Hymenophyllum siliquosum H. Christ =Hymenophyllum apiculatum 

Hymenophyllum surinamense Bosch, nom. superfl. =Hymenophyllum hirsutum 

Hymenophyllum trapezoidale Liebm. [records due to misidentification fide Lellinger] 

Hymenophyllum undulatum var. fendlerianum (J.W. Sturm) Stolze =Hymenophyllum fendlerianum 

Hymenostachys diversifrons Bory =Trichomanes diversifrons 

Hymenostachys elegans (Rudge) K. Presl =Trichomanes diversifrons 

Hymenostachys osmundoides (DC. ex Poir.) C. Presl =Trichomanes osmundoides 

Lacostea ankersii (C. Parker ex Hook. & Grev.) Prantl =Trichomanes ankersii 

Lacostea tanaica (Hook. ex J.W. Sturm) Prantl =Trichomanes tanaicum 

Lecanium membranaceum (L.) C. Presl =Trichomanes membranaceum 

Mecodium apiculatum (Mett. ex Kuhn) Vareschi =Hymenophyllum apiculatum 

Mecodium asplenioides (Sw.) Copel. =Hymenophyllum asplenioides 

Mecodium axillare (Sw.) Copel. =Hymenophyllum axillare 

Mecodium fendlerianum (J.W. Sturm) Copel. =Hymenophyllum fendlerianum 

Mecodium microcarpum (Desv.) Copel. =Hymenophyllum microcarpum 

Mecodium polyanthos (Sw.) Copel. =Hymenophyllum polyanthos 

Mecodium trichomanoides (Bosch) Pic. Serm. =Hymenophyllum trichomanoides 

Mecodium undulatum (Sw.) Copel. =Hymenophyllum undulatum 

Microgonium kapplerianum (J.W. Sturm) Pic. Serm. =Trichomanes kapplerianum 

Neuromanes abruptum Bosch, nom. superfl. =Trichomanes hostmannianum 

Neuromanes kaulfussii Bosch, nom. superfl. =Trichomanes pinnatum 

Neuromanes schomburgkianum (J.W. Sturm) Bosch =Trichomanes pinnatum 

Neuromanes vittaria (DC. ex Poir.) Bosch =Trichomanes vittaria 

Neurophyllum hostmannianum Klotzsch =Trichomanes hostmannianum 

Neurophyllum pennatum (Kaulf.) K. Presl =Trichomanes pinnatum 

Neurophyllum pinnatum (Hedw.) K. Presl =Trichomanes pinnatum 

Neurophyllum rhizophyllum (Cav.) Fée =Trichomanes pinnatum 

Neurophyllum vittaria (DC. ex Poir.) Prantl =Trichomanes vittaria 

Odontomanes hostmannianum (Klotzsch) C. Presl =Trichomanes hostmannianum 

Ptilophyllum accedens (C. Presl) Prantl =Trichomanes accedens 

Ptilophyllum bancroftii (Hook. & Grev.) Prantl =Trichomanes bancroftii 

Ptilophyllum bicorne (Hook.) Prantl =Trichomanes bicorne 

Ptilophyllum botryoides (Kaulf.) Prantl =Trichomanes botryoides 

Ptilophyllum elegans (Rudge) Prantl =Trichomanes diversifrons 

Ptilophyllum hostmannianum (Klotzsch) Prantl =Trichomanes hostmannianum 

Ptilophyllum pinnatum (Kaulf.) Prantl =Trichomanes pinnatum 

Ptilophyllum schomburgkianum (J.W. Sturm) Prantl =Trichomanes pinnatum 

Ptilophyllum vittaria (DC. ex Poir.) Prantl =Trichomanes vittaria 

Ragatelus eriophorus C. Presl =Trichomanes crispum 

Selenodesmium rigidum (Sw.) Copel. =Trichomanes rigidum 

Sphaerocionium antillense (Jenman) Copel. =Hymenophyllum antillense 

Sphaerocionium crispum var. amoenum Klotzsch =Hymenophyllum fendlerianum 

Sphaerocionium dependens (C.V. Morton) Vareschi =Hymenophyllum dependens 

Sphaerocionium elegans (Spreng.) Copel. =Hymenophyllum elegans 

Sphaerocionium hirsutum (L.) C. Presl =Hymenophyllum hirsutum 

Sphaerocionium karstenianum (J.W. Sturm) Vareschi =Hymenophyllum karstenianum 

Sphaerocionium lindenii (Hook.) Vareschi =Hymenophyllum lindenii 

Sphaerocionium microcarpum (Desv.) Copel. =Hymenophyllum microcarpum 

Sphaerocionium plumieri (Hook. & Grev.) C. Presl =Hymenophyllum plumieri 

Sphaerocionium roraimense (C.V. Morton) Vareschi =Hymenophyllum roraimense 

Sphaerocionium trichophyllum (Kunth) Copel. =Hymenophyllum trichophyllum 

Sphaerocionium valvatum (Hook. & Grev.) Copel. =Hymenophyllum valvatum 

Trichomanes abruptum (Bosch) H. Christ =Trichomanes hostmannianum 

Trichomanes ankersii var. tanaicum (Hook. ex J.W. Sturm) Sadeb. =Trichomanes tanaicum 

Trichomanes asplenioides Sw. =Hymenophyllum asplenioides 

Trichomanes carolianum Vareschi =Trichomanes tanaicum 

Trichomanes ciliatum Sw. =Hymenophyllum hirsutum 

Trichomanes commutatum J.W. Sturm =Trichomanes pedicellatum


Trichomanes curranii Weath. [Guianas records due to misidentifications of T. caliginum] =Trichomanes fimbriatum 

Trichomanes digitatum Sodiro =Trichomanes dactylites 

Trichomanes elegans Rudge, non Rich. [nom. illeg.] =Trichomanes diversifrons 

Trichomanes floribundum var. vittaria (DC. ex Poir.) Splitg. =Trichomanes vittaria 

Trichomanes fruticulosum Jenman =Trichomanes punctatum subsp. labiatum 

Trichomanes fucoides Sw. =Hymenophyllum fucoides 

Trichomanes furcatum Bosch, non Kaulf. ex Spreng. =Trichomanes pedicellatum 

Trichomanes gemmatum J. Sm. ex Hook. & Baker, nom. superfl. =Trichomanes cellulosum 

Trichomanes guianense J.W. Sturm =Trichomanes ankersii 

Trichomanes heterophyllum Humb. & Bonpl. ex Willd. =Trichomanes humboldtii 

Trichomanes hirsutum L. =Hymenophyllum hirsutum 

Trichomanes hookeri auct. non C. Presl =Trichomanes kapplerianum 

Trichomanes huberi H. Christ =Trichomanes hostmannianum 

Trichomanes kunzeanum Hook. =Trichomanes radicans 

Trichomanes labiatum Jenman =Trichomanes punctatum subsp. labiatum 

Trichomanes lambertianum Hook. =Trichomanes lucens 

Trichomanes leprieurii Hook., nom. superfl. =Trichomanes elegans 

Trichomanes leptophyllum Bosch [nom. illeg.] =Trichomanes hymenophylloides 

Trichomanes myrioneuron Lindm. =Trichomanes punctatum subsp. labiatum 

Trichomanes nanum (Bory) Hook. =Trichomanes botryoides 

Trichomanes pennatum Kaulf. =Trichomanes pinnatum 

Trichomanes pilosum Raddi =Trichomanes martiusii 

Trichomanes pinnatum var. kaulfussii Domin =Trichomanes pinnatum 

Trichomanes pinnatum var. vittaria (DC. ex Poir.) Baker =Trichomanes vittaria 

Trichomanes polyanthos Sw. =Hymenophyllum polyanthos 

Trichomanes polyphlebius V. Marcano =Trichomanes arbuscula 

Trichomanes prieurii Kunze =Trichomanes elegans 

Trichomanes rhizophyllum Cav. =Trichomanes pinnatum 

Trichomanes schomburgkianum J.W. Sturm =Trichomanes pinnatum 

Trichomanes schomburgkii Bosch =Trichomanes crispum 

Trichomanes sellowianum C. Presl =Trichomanes cristatum 

Trichomanes sphenoides auct. non Kunze =Trichomanes punctatum 

Trichomanes spicisorum Desv. =Trichomanes osmundoides 

Trichomanes subsessile Splitg. =Trichomanes pedicellatum 

Trichomanes superbum Bosch =Trichomanes fimbriatum 

Trichomanes torotumanii Vareschi =Trichomanes tuerckheimii 

Trichomanes trigonum auct. non Desv. [records due to misidentification fide Lellinger] =Trichomanes fimbriatum 

Trichomanes trigonum var. fimbriatum (Backh. ex T. Moore) Proctor =Trichomanes fimbriatum 

Trichomanes undulatum Sw. =Hymenophyllum undulatum 

Trichomanes undulatum Bosch =Trichomanes vandenboschii 

Trichomanes windischianum Lellinger =Trichomanes sprucei 

Vandenboschia capillacea (L.) Copel. =Trichomanes capillaceum 

REFERENCES 

Lellinger, D.B. 1984. Hymenophyllaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 12. Mem. New 

York Bot. Gard. 38: 1-84. 

Lellinger, D.B. 1991. Notes on neotropical Hymenophyllaceae. Amer. Fern J. 81: 42-46. 

Lellinger, D.B. 1994. Trichomanes polypodioides and its allies. Amer. Fern J. 84(1): 1-4. 

Lellinger, D.B. 1994. Hymenophyllaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 3: 1-66. 

Smith, A.R. 1995. Hymenophyllaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 159- 

185. 

Hymenophyllopsis asplenioides A.C. Sm. [Mt. Roraima] 

Hymenophyllopsis ctenitoides Lellinger [Mt. Roraima] 

Hymenophyllopsis dejecta (Baker) K.I. Goebel [Mt. Roraima] 

Hymenophyllopsis hymenophylloides L.D. Gómez 

Hymenophyllopsis incognita Lellinger 

Hymenophyllopsis steyermarkii Vareschi 

Hymenophyllopsis superba Lellinger 

Hymenophyllopsis universitatis Vareschi 

HYMENOPHYLLOPSIDACEAE 

AM BO GU 

AM BO GU? 

AM BO GU 

AM BO 

BO 

AM BO 

AM 

BO

42 


REFERENCES 

Lellinger, D.B. 1984. Hymenophyllopsidaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 12. Mem. 

New York Bot. Gard. 38: 1-84. 

Lellinger, D.B. 1991. Hymenophyllopsidaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 4: 82-86. 

Lellinger, D.B. 1995. Hymenophyllopsidaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 

2: 186-188. 

Bolbitis lindigii (Mett.) Ching 

Bolbitis nicotianifolia (Sw.) Alston 

Bolbitis portoricensis (Spreng.) Hennipman 

Bolbitis semipinnatifida (Fée) Alston 

Bolbitis serratifolia (Mert. ex Kaulf.) Schott 

Elaphoglossum acrocarpum (Mart.) T. Moore 

Elaphoglossum andicola (Fée) T. Moore 

Elaphoglossum antioquianum Hieron. 

Elaphoglossum apodum (Kaulf.) Schott ex J. Sm. 

Elaphoglossum auricomum (Kunze) T. Moore 

Elaphoglossum brachyneuron (Fée) J. Sm. 

Elaphoglossum crinitum (L.) H. Christ 

Elaphoglossum crispatum Mickel 

Elaphoglossum crispatum var. beitelii Mickel 

Elaphoglossum crispatum var. crispatum 

Elaphoglossum cuspidatum (Willd.) T. Moore 

Elaphoglossum decoratum (Kunze) T. Moore 

Elaphoglossum discolor (Kuhn) C. Chr. 

Elaphoglossum dolichopus Mickel 

Elaphoglossum engelii (H. Karst.) H. Christ 

Elaphoglossum erinaceum (Fée) T. Moore 

Elaphoglossum eximium (Mett.) H. Christ 

Elaphoglossum flaccidum (Fée) T. Moore 

Elaphoglossum floccosum Mickel 

Elaphoglossum foldatsii Vareschi 

Elaphoglossum glabellum J. Sm. 

Elaphoglossum hayesii (Mett. ex Kuhn) Maxon 

Elaphoglossum herminieri (Bory & Fée ex Fée) T. Moore 

Elaphoglossum hoffmannii (Mett. ex Kuhn) H. Christ 

Elaphoglossum horridulum (Kaulf.) J. Sm. 

Elaphoglossum incubus Mickel 

Elaphoglossum laminarioides (Bory ex Fée) T. Moore 

Elaphoglossum langsdorffii (Hook. & Grev.) T. Moore 

LOMARIOPSIDACEAE 

Elaphoglossum latifolium (Sw.) J. Sm. [see also Elaphoglossum sporadolepis] 

Elaphoglossum leptophlebium (Baker) C. Chr. 

Elaphoglossum lingua (Raddi) Brack. 

Elaphoglossum lloense (Hook.) T. Moore 

Elaphoglossum longicaudatum Mickel 

Elaphoglossum luridum (Fée) H. Christ 

Elaphoglossum macrophyllum (Mett. ex Kuhn) H. Christ 

Elaphoglossum maguirei Mickel 

Elaphoglossum meridense (Klotzsch) T. Moore 

Elaphoglossum minutum (Pohl ex Fée) T. Moore [Mt. Roraima] 

Elaphoglossum moritzianum (Klotzsch) T. Moore 

Elaphoglossum nigrescens (Hook.) T. Moore ex Diels 

Elaphoglossum obovatum Mickel 

Elaphoglossum paleaceum (Hook. & Grev.) Sledge 

Elaphoglossum papillosum (Baker) H. Christ 

Elaphoglossum parvulum Mickel 

Elaphoglossum peltatum (Sw.) Urb. 

Elaphoglossum piloselloides (C. Presl) T. Moore 

Elaphoglossum plumosum (Fée) T. Moore 

Elaphoglossum polyblepharum Mickel 

Elaphoglossum praetermissum Mickel 

Elaphoglossum productum Rosenst. [Mt. Roraima] 

GU 


BO 



AM BO 

AM BO 

AM BO GU 

GU SU FG 

BO 

AM BO GU 

BO GU 

AM 

AM 

AM 

AM GU 

AM BO GU 

AM BO GU? 

AM 

GU 

AM 

BO GU 


BO 

AM BO 


BO SU 


AM BO 

AM GU 

AM 


AM BO 

GU SU FG 

GU 

GU FG 

SU FG? 

AM BO FG 


AM BO GU FG 

AM 

BO 

BO 

GU 

AM BO GU 

AM BO 

BO 

AM 

AM 

AM BO GU 

BO SU FG 


AM 

AM BO GU 

AM BO GU?


Elaphoglossum pteropodum (Sodiro) C. Chr. 

Elaphoglossum pteropus C. Chr. 

Elaphoglossum pusillum (Mett. ex Kuhn) C. Chr. 

Elaphoglossum raywaense (Jenman) Alston 

Elaphoglossum revolutum (Liebm.) T. Moore 

Elaphoglossum scandens (Bory ex Fée) T. Moore 

Elaphoglossum sporadolepis (Kunze ex Kuhn) T. Moore ex C. Chr. 

[=Elaphoglossum latifolium?] 

Elaphoglossum squarrosum (Klotzsch) T. Moore 

Elaphoglossum steyermarkii Mickel 

Elaphoglossum strictum (Raddi) T. Moore 

Elaphoglossum styriacum Mickel 

Elaphoglossum succubus Mickel 

Elaphoglossum tantalinum Mickel 

Elaphoglossum tectum (Humb. & Bonpl. ex Willd.) T. Moore 

Elaphoglossum tenuifolium (Liebm.) T. Moore 

Elaphoglossum tovarense (Mett. ex Kuhn) T. Moore ex C. Chr. [Mt. Roraima] 

Elaphoglossum vanderwerffii Mickel 

Elaphoglossum vareschianum Mickel 

Elaphoglossum wurdackii Vareschi [Mt. Roraima] 

Elaphoglossum xiphoides Mickel 

Elaphoglossum zosteriformis Mickel 

Lomagramma guianensis (Aubl.) Ching 

Lomariopsis erythrodes (Kunze) Fée 

Lomariopsis fendleri D.C. Eaton 

Lomariopsis japurensis (Mart.) J. Sm. 

Lomariopsis prieuriana Fée 


Bolbitis aliena (Sw.) auct. non Alston =Bolbitis semipinnatifida 

Bolbitis cladorrhizans (Spreng.) Ching =Bolbitis portoricensis 

Bolbitis guianensis (Aubl.) K.U. Kramer ex Vareschi =Lomagramma guianensis 

Bolbitis maguirei Maxon & C.V. Morton =Bolbitis serratifolia 

Bolbitis scopulina Maxon & C.V. Morton =Bolbitis semipinnatifida 

Elaphoglossum agostinii Vareschi =Elaphoglossum sporadolepis 

Elaphoglossum attenuatum (Kunze ex Kuhn) H. Christ =Elaphoglossum hoffmannii 

Elaphoglossum cordigerum H. Christ =Elaphoglossum eximium 

Elaphoglossum fraseri (Mett. ex Kuhn) H. Christ =Elaphoglossum pusillum 

Elaphoglossum gayanum (Fée) T. Moore =Elaphoglossum minutum 

Elaphoglossum leptophyllum (Fée) T. Moore =Elaphoglossum minutum 

Elaphoglossum lineare var. klotzschii (Moritz) Hieron. =Elaphoglossum eximium 

Elaphoglossum longifolium (C. Presl) J. Sm. =Elaphoglossum latifolium 

Elaphoglossum reichenbachii (Moritz ex Kuhn) H. Christ =Elaphoglossum eximium 

Elaphoglossum rigidum (Aubl.) Urb. =Elaphoglossum flaccidum 

Elaphoglossum schomburgkii (Fée) T. Moore =Elaphoglossum luridum 

Elaphoglossum spathulatum auct. non (Bory) T. Moore =Elaphoglossum piloselloides 

Elaphoglossum squamosum (Sw.) J. Sm. =Elaphoglossum paleaceum 

Elaphoglossum stenopteris (Klotzsch) T. Moore =Elaphoglossum brachyneuron 

Elaphoglossum williamsii Vareschi =Elaphoglossum nigrescens 

Hymenodium crinitum (L.) Fée =Elaphoglossum crinitum 

Leptochilus alienus auct. non (Sw.) C. Chr. =Bolbitis semipinnatifida 

Leptochilus guianensis (Aubl.) C. Chr. =Lomagramma guianensis 

Leptochilus nicotianifolius (Sw.) C. Chr. =Bolbitis nicotianifolia 

Leptochilus serratifolius (Mert. ex Kaulf.) C. Chr. =Bolbitis serratifolia 

Peltapteris peltata (Sw.) C.V. Morton =Elaphoglossum obovatum 

Rhipidopteris peltata (Sw.) Schott =Elaphoglossum peltatum 

REFERENCES 

Hennipman, E. 1977. A monograph of the fern genus Bolbitis. Leiden Bot. Ser. 2: 1-331. 



Lygodium flexuosum (L.) Sw. 

LYGODIACEAE 

FG? 


BO 


FG 

SU? FG? 


AM 

BO 

GU SU FG 

AM BO FG? 

BO 

AM 

AM 

FG 

AM BO 

AM 

BO 

AM BO 

BO 

BO 


FG? 

BO 


GU SU FG 

GU

44 


Lygodium microphyllum (Cav.) R. Br. [introduced from Old World tropics] 

Lygodium venustum Sw. 

Lygodium volubile Sw. 


Lygodium mexicanum C. Presl =Lygodium venustum 

Lygodium micans J.W. Sturm =Lygodium volubile 

Lygodium polymorphum auct. non (Cav.) Kunth =Lygodium venustum 

REFERENCES 

Duek, J.J. 1978. A taxonomic revision of Lygodium in America. Feddes Repert. 89: 411-423. 

Lellinger, D.B. 1969. Schizaeaceae. In B. Maguire & collaborators, Botany of the Guayana Highland - Part VIII. Mem. New 

York Bot. Gard. 18(2): 2-11. 

Smith, A.R. 1995. Schizaeaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 288-296. 

Danaea elliptica Sm. 

Danaea leprieurii Kunze 

Danaea moritziana C. Presl 

Danaea nodosa (L.) Sm. 

Danaea simplicifolia Rudge 

Danaea trifoliata Rchb. ex Kunze 

Marattia laevis Sm. 

MARATTIACEAE 


Danaea elliptica subsp. simplicifolia (Rudge) Rolleri =Danaea simplicifolia 

Danaea nigrescens Jenman =Danaea nodosa 

Marattia kaulfussii J. Sm. =Marattia laevis 

REFERENCES 

Smith, A.R. 1995. Marattiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 206-209. 

Marsilea deflexa A. Braun 

Marsilea minuta L. 

Marsilea polycarpa Hook. & Grev. 

MARSILEACEAE 

REFERENCES 

Johnson, D.M. 1986. Systematics of the New World species of Marsilea. Syst. Bot. Monogr. 11: 1-87. 

Reed, C.F. 1965. Index Marsileata et Salviniata. Suppl. Bol. Soc. Brot. II, 39: 259-302. 

Smith, A.R. 1995. Marsileaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 209-211. 

Metaxya rostrata (Kunth) C. Presl 

METAXYACEAE 


Amphidesmium blechnoides (Hook.) J. Sm. =Metaxya rostrata 

Amphidesmium rostratum (Kunth) J. Sm. =Metaxya rostrata 

REFERENCES 

Smith, A.R. 1995. Metaxyaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 211-212. 

Nephrolepis biserrata (Sw.) Schott 

Nephrolepis cordifolia (L.) C. Presl 

Nephrolepis multiflora (Roxb.) Jarrett ex C.V. Morton 

Nephrolepis pectinata (Willd.) Schott 

Nephrolepis rivularis (Vahl) Mett. ex Krug 

Oleandra articulata (Sw.) C. Presl 

Oleandra duidae A.C. Sm. 

Oleandra pilosa Hook. 

OLEANDRACEAE 

GU 



AM BO GU? SU FG 

GU SU FG 

AM BO FG 

AM SU FG 


BO GU SU FG 

BO 

GU 

GU? 





SU 

BO GU 



AM BO 

AM BO GU SU FG



Nephrolepis pendula (Raddi) J. Sm. =Nephrolepis cordifolia 

Oleandra nodosa (Willd.) C. Presl =Oleandra articulata 

REFERENCES 

Cremers, G. & K.U. Kramer. 1994. Nephrolepidaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 6: 66-72. 

Cremers, G. & K.U. Kramer. 1994. Oleandraceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 6: 73-76. 

Smith, A.R. 1995. Davalliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 43-46. 



Cheiroglossa palmata (L.) C. Presl 

Ophioglossum nudicaule L. f. 

Ophioglossum reticulatum L. 

OPHIOGLOSSACEAE 


Ophioglossum ellipticum Hook. & Grev. =Ophioglossum nudicaule 

Ophioglossum macrorhizum Kunze =Ophioglossum nudicaule 

Ophioglossum palmatum L. =Cheiroglossa palmata 

REFERENCES 

Clausen, R.T. 1938. A monograph of the Ophioglossaceae. Mem. Torrey Bot. Club 19(2): 1-177. 

Smith, A.R. 1995. Ophioglossaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 212-214. 

Osmunda cinnamomea L. 

Osmunda regalis L. 

OSMUNDACEAE 


Osmunda cervina L. (Dryopteridaceae) =Olfersia cervina 

Osmunda humilis Cav. (Schizaeaceae) =Anemia oblongifolia 

Osmunda oblongifolia Cav. (Schizaeaceae) =Anemia oblongifolia 

Osmunda peltata Sw. (Lomariopsidaceae) =Elaphoglossum obovatum 

Ceratopteris deltoidea Benedict [=Ceratopteris pteridoides?] 

Ceratopteris pteridoides (Hook.) Hieron. 

Ceratopteris richardii Brongn. 

Ceratopteris thalictroides (L.) Brongn. 

PARKERIACEAE 


Ceratopteris calomelanos (L.) Link (Pteridaceae) =Pityrogramma calomelanos 

Ceratopteris lockhartii (Hook. & Grev.) Kunze =Ceratopteris pteridoides or Ceratopteris thalictroides 

Parkeria pteridoides Hook. =Ceratopteris pteridoides 

REFERENCES 

Lloyd, R.M. 1974. Systematics of the genus Ceratopteris Brongn. II. Taxonomy. Brittonia 26: 139-160. 

Smith, A.R. 1995. Parkeriaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 215-217. 

Plagiogyria semicordata (C. Presl) H. Christ 

PLAGIOGYRIACEAE 


Lomaridium semicordatum C. Presl =Plagiogyria semicordata 

POLYPODIACEAE 

Campyloneurum amphostenon (Kunze ex Klotzsch) Fée [Mt. Roraima] 

AM BO DA GU SU 

BO GU SU FG 

GU SU FG? 

AM 

BO 

BO 

AM BO 

GU SU 


GU SU FG 

BO DA GU SU FG

46 

Campyloneurum angustifolium (Sw.) Fée 

Campyloneurum coarctatum (Kunze) Fée 

Campyloneurum fuscosquamatum Lellinger 

Campyloneurum latum T. Moore 

Campyloneurum occultum (H. Christ) L.D. Gómez 

Campyloneurum phyllitidis (L.) C. Presl 

Campyloneurum repens (Aubl.) C. Presl 

Campyloneurum wurdackii B. León 

Dicranoglossum desvauxii (Klotzsch) Proctor 

Dicranoglossum furcatum (L.) J. Sm. 

Microgramma baldwinii Brade 

Microgramma fuscopunctata (Hook.) Vareschi 

Microgramma geminata (Schrad.) R.M. Tryon & A.F. Tryon 

Microgramma lycopodioides (L.) Copel. 

Microgramma megalophylla (Desv.) de la Sota 

Microgramma persicariifolia (Schrad.) C. Presl 

Microgramma piloselloides (L.) Copel. 

Microgramma reptans (Cav.) A.R. Sm. 

Microgramma tecta (Kaulf.) Alston 

Microgramma thurnii (Baker) R.M. Tryon & Stolze 

Neurodium lanceolatum (L.) Fée 

Niphidium crassifolium (L.) Lellinger 

Pecluma camptophyllaria (Fée) M.G. Price 

Pecluma camptophyllaria var. lachnifera (Hieron.) Lellinger 

Pecluma consimilis (Mett.) M.G. Price 

Pecluma consimilis var. consimilis 

Pecluma dispersa (A.M. Evans) M.G. Price 

Pecluma eurybasis (C. Chr.) M.G. Price [Mt. Roraima] 

Pecluma hygrometrica (Splitg.) M.G. Price 

Pecluma pectinata (L.) M.G. Price 

Pecluma plumula (Humb. & Bonpl. ex Willd.) M.G. Price 

Pecluma ptilodon (Kunze) M.G. Price 

Pecluma ptilodon var. caespitosa (Jenman) Mickel & Beitel 

Pecluma ptilodon var. pilosa (A.M. Evans) Stolze 

Pecluma ptilodon var. ptilodon 

Phlebodium aureum (L.) J. Sm. 

Phlebodium decumanum (Willd.) J. Sm. 

Phlebodium pseudoaureum (Cav.) Lellinger 

Pleopeltis astrolepis (Liebm.) E. Fourn. 

Pleopeltis macrocarpa (Bory ex Willd.) Kaulf. 

Pleopeltis percussa (Cav.) Hook. & Grev. 

Pleopeltis repanda A.R. Sm. 

Polypodium adnatum Kunze ex Klotzsch 

Polypodium attenuatum Humb. & Bonpl. ex Willd. 

Polypodium aturense Maury 

Polypodium bombycinum Maxon 

Polypodium caceresii Sodiro 

Polypodium dissimile L. 

Polypodium dulce Poir. 

Polypodium flagellare H. Christ 

Polypodium fraxinifolium Jacq. 

Polypodium funckii Mett. 

Polypodium lasiopus Klotzsch 

Polypodium levigatum Cav. 

Polypodium loriceum L. 

Polypodium panorense C. Chr. 

Polypodium polypodioides (L.) Watt 

Polypodium polypodioides var. burchellii (Baker) Weath. 

Polypodium polypodioides var. polypodioides 

Polypodium sessilifolium Desv. 

Polypodium thyssanolepis A. Braun ex Klotzsch 

Polypodium triseriale Sw. 

Polypodium wagneri Mett. 

Terpsichore bradeana Labiak 



BO 

GU SU FG 

GU SU FG 

FG 



AM BO 


SU FG 

AM BO GU 


GU 


AM GU SU? 


GU FG 


AM GU SU FG 


FG? 


BO 

BO 

GU 

GU 

BO 

AM BO 

AM BO GU SU 




SU FG 


GU 




BO GU SU FG 

AM BO 


BO 



AM BO 

AM BO GU 


GU SU FG 


SU FG 

AM BO GU 

AM BO 

BO SU FG 

AM GU SU FG 

BO GU 

GU 



GU FG? 

AM BO GU 

BO 


AM BO 

BO

Terpsichore steyermarkii Lobiak 



Campyloneurum costatum (Kunze) C. Presl =Campyloneurum phyllitidis 

Campyloneurum sphenodes (Kunze) Fée [not in flora area] 

Drynaria acuminata Fée =Microgramma thurnii 

Eschatogramme desvauxii (Klotzsch) C. Chr. =Dicranoglossum desvauxii 

Eschatogramme furcata (L.) C. Chr. =Dicranoglossum furcatum 

Eschatogramme furcata auct. non (L.) C. Chr. =Dicranoglossum desvauxii 

Goniophlebium adnatum (Kunze ex Klotzsch) T. Moore =Polypodium adnatum 

Goniophlebium attenuatum (Humb. & Bonpl. ex Willd.) C. Presl =Polypodium attenuatum 

Goniophlebium fraxinifolium (Jacq.) Moore =Polypodium fraxinifolium 

Goniophlebium glaucophyllum (Kunze ex Klotzsch) Fée =Polypodium levigatum 

Goniophlebium lasiopus (Klotzsch) T. Moore =Polypodium lasiopus 

Goniophlebium loriceum (L.) J. Sm. ex Hook. =Polypodium loriceum 

Goniophlebium triseriale (Sw.) Pic. Serm. =Polypodium triseriale 

Marginaria bombycina (Maxon) Pic. Serm. =Polypodium bombycinum 

Marginaria polypodioides (L.) Tidestr. =Polypodium polypodioides 

Marginaria thyssanolepis (A. Braun ex Klotzsch) Farw. =Polypodium thyssanolepis 

Microgramma acuminata (Fée) Lellinger =Microgramma thurnii 

Microgramma ciliata (Willd.) Alston =Microgramma reptans 

Microgramma percussa (Cav.) de la Sota =Pleopeltis percussa 

Paltonium lanceolatum (L.) C. Presl =Neurodium lanceolatum 

Pessopteris crassifolia (L.) Underw. & Maxon =Niphidium crassifolium 

Phymatodes prominula Maxon =Microgramma lycopodioides 

Pleopeltis fuscopunctata (Hook.) R.M. Tryon & A.F. Tryon =Microgramma fuscopunctata 

Pleopeltis lanceolata (L.) C. Presl =Pleopeltis macrocarpa 

Pleopeltis polypodioides (L.) E.G. Andrews & Windham =Polypodium polypodioides 

Pleopeltis revoluta (Spreng. ex Willd.) A.R. Sm. =Pleopeltis astrolepis 

Pleopeltis thyssanolepis (A. Braun ex Klotzsch) E.G. Andrews & Windham =Polypodium thyssanolepis 

Polypodium abruptum Desv. (Thelypteridaceae) =Thelypteris abrupta 

Polypodium adiantiforme G. Forst. (Davalliaceae) =Rumohra adiantiformis 

Polypodium adiantoides Aubl., non Burm. (Dryopteridaceae) =Polybotrya caudata 

Polypodium amphostenon Kunze ex Klotzsch =Campyloneurum amphostenon 

Polypodium anfractuosum Kunze ex Klotzsch (Grammitidaceae) =Grammitis anfractuosa 

Polypodium angustifolium Sw. =Campyloneurum angustifolium 

Polypodium angustifolium var. amphostenon (Kunze ex Klotzsch) Baker =Campyloneurum amphostenon 

Polypodium apiculatum Kunze ex Klotzsch (Grammitidaceae) =Grammitis apiculata 

Polypodium areolatum Humb. & Bonpl. ex Willd. =Phlebodium pseudoaureum 

Polypodium aromaticum Maxon (Grammitidaceae) =Grammitis firma 

Polypodium asplenifolium L. (Grammitidaceae) =Grammitis asplenifolia 

Polypodium astrolepis Liebm. =Pleopeltis astrolepis 

Polypodium aureum L. =Phlebodium aureum 

Polypodium aureum var. areolatum (Humb. & Bonpl. ex Willd.) Baker =Phlebodium pseudoaureum 

Polypodium balbisii Spreng. (Thelypteridaceae) =Thelypteris balbisii 

Polypodium binervatum Poir. (Blechnaceae) =Blechnum binervatum 

Polypodium blanchetii C. Chr. (Grammitidaceae) =Grammitis blanchetii 

Polypodium bolivari de la Sota =Polypodium bombycinum 

Polypodium brasiliense Poir. =Polypodium triseriale 

Polypodium bryophilum Maxon (Grammitidaceae) =Grammitis bryophila 

Polypodium camptophyllarium Fée =Pecluma camptophyllaria 

Polypodium camptophyllarium var. lachniferum (Hieron.) A.M. Evans =Pecluma camptophyllaria var. lachnifera 

Polypodium capillare Desv. (Grammitidaceae) =Ceradenia capillaris 

Polypodium caucanum Hieron. (Grammitidaceae) =Grammitis caucana 

Polypodium chimantense Vareschi (Grammitidaceae) =Ceradenia discolor 

Polypodium ciliatum Willd. =Microgramma reptans 

Polypodium coarctatum Kunze =Campyloneurum coarctatum 

Polypodium confusum J. Sm. (Grammitidaceae) =Grammitis suspensa 

Polypodium connellii Baker ex C.H. Wright (Grammitidaceae) =Cochlidium connellii 

Polypodium consimile Mett. =Pecluma consimilis 

Polypodium crassifolium L. =Niphidium crassifolium 

Polypodium crispatum (J. Sm.) C.V. Morton (Grammitidaceae) [not in flora area] =Enterosora ecostata (Sodiro) L.E.Bishop 

Polypodium cultratum auct. non Willd. (Grammitidaceae) =Grammitis mollissima 

Polypodium decumanum Willd. =Phlebodium decumanum 

BO

48 


Polypodium decussatum L. (Thelypteridaceae) =Thelypteris decussata 

Polypodium demeraranum Baker (Thelypteridaceae) =Thelypteris demerarana 

Polypodium denticulatum Sw. (Dryopteridaceae) =Arachniodes denticulata 

Polypodium dicranophyllum C. Chr. (Grammitidaceae) =Cochlidium furcatum 

Polypodium discolor Hook. (Grammitidaceae) =Ceradenia discolor 

Polypodium dispersum A.M. Evans =Pecluma dispersa 

Polypodium dissectum Sw. (Dennstaedtiaceae) =Dennstaedtia dissecta 

Polypodium divergens Willd. (Tectariaceae) =Lastreopsis effusa subsp. divergens 

Polypodium duale Maxon (Grammitidaceae) =Cochlidium serrulatum 

Polypodium effusum Sw. (Tectariaceae) =Lastreopsis effusa 

Polypodium eurybasis C. Chr. =Pecluma eurybasis 

Polypodium exiguum Fée (Grammitidaceae) =Grammitis blanchetii 

Polypodium firmum Klotzsch (Grammitidaceae) =Grammitis firma 

Polypodium flabelliforme Poir. (Grammitidaceae) =Grammitis flabelliformis 

Polypodium furcatum (Hook. & Grev.) Mett. (Grammitidaceae) =Cochlidium furcatum 

Polypodium fuscopunctatum Hook. =Microgramma fuscopunctata 

Polypodium glaucophyllum Kunze ex Klotzsch =Polypodium levigatum 

Polypodium guianense Aubl. (Lomariopsidaceae) =Lomagramma guianensis 

Polypodium haplophlebicum A.C. Sm. (Grammitidaceae) =Grammitis bryophila 

Polypodium hirsutum Desv. (Cyatheaceae) =Cyathea surinamensis 

Polypodium horridum L. (Cyatheaceae) =Cnemidaria horrida 

Polypodium hostmannii Klotzsch (Thelypteridaceae) =Thelypteris arborescens 

Polypodium hygrometricum Splitg. =Pecluma hygrometrica 

Polypodium incanum var. burchelli Baker =Polypodium polypodioides var. burchellii 

Polypodium induens Maxon (Grammitidaceae) =Grammitis anfractuosa 

Polypodium jubiforme Kaulf. (Grammitidaceae) =Grammitis suspensa 

Polypodium jungermannioides Klotzsch (Grammitidaceae) =Ceradenia jungermannioides 

Polypodium kaieteurum Jenman (Grammitidaceae) =Grammitis tenuicula 

Polypodium kalbreyeri Baker (Grammitidaceae) =Ceradenia kalbreyeri 

Polypodium kegelianum Kunze (Grammitidaceae) =Grammitis kegeliana 

Polypodium kookenamae Jenman (Grammitidaceae) =Ceradenia kookenamae 

Polypodium lachniferum Hieron. =Pecluma camptophyllaria var. lachnifera 

Polypodium lanceolatum auct. non L. =Pleopeltis astrolepis 

Polypodium leptopodon C.H. Wright (Grammitidaceae) =Grammitis leptopoda 

Polypodium l’herminieri Fée (Grammitidaceae) =Grammitis taxifolia 

Polypodium limulum H. Christ (Grammitidaceae) =Grammitis limula 

Polypodium longicaudatum Liebm. (Dryopteridaceae) =Stigmatopteris longicaudata 

Polypodium lycopodioides L. =Microgramma lycopodioides 

Polypodium macrocarpum Bory ex Willd. =Pleopeltis macrocarpa 

Polypodium megalodus Schkuhr (Thelypteridaceae) =Thelypteris pennata 

Polypodium megalophyllum Desv. =Microgramma megalophylla 

Polypodium melanostictum Kunze (Grammitidaceae) =Grammitis melanosticta 

Polypodium melanotrichium Baker (Grammitidaceae) =Grammitis melanotrichia 

Polypodium microdonton Desv. (Cyatheaceae) =Cyathea microdonta 

Polypodium mollissimum Fée (Grammitidaceae) =Grammitis mollissima 

Polypodium moniliforme Lag. ex Sw. (Grammitidaceae) =Grammitis moniliformis 

Polypodium myosuroides Sw. (Grammitidaceae) =Grammitis myosuroides 

Polypodium nanum Fée (Grammitidaceae) =Grammitis blanchetii 

Polypodium nematorhizon D.C. Eaton =Microgramma fuscopunctata 

Polypodium neriifolium Schkuhr =Polypodium triseriale 

Polypodium nervosum Klotzsch (Thelypteridaceae) =Thelypteris rudis 

Polypodium oligocarpum Humb. & Bonpl. ex Willd. (Thelypteridaceae) =Thelypteris oligocarpa 

Polypodium oppositum Vahl (Thelypteridaceae) =Thelypteris opposita 

Polypodium parietinum Klotzsch (Grammitidaceae) =Enterosora parietina 

Polypodium pectinatum L. =Pecluma pectinata 

Polypodium pennatum Poir. (Thelypteridaceae) =Thelypteris pennata 

Polypodium percussum Cav. =Pleopeltis percussa 

Polypodium persicariifolium Schrad. =Microgramma persicariifolia 

Polypodium phlegmaria J. Sm. (Grammitidaceae) =Grammitis phlegmaria 

Polypodium phyllitidis L. =Campyloneurum phyllitidis 

Polypodium plantagineum Jacq. (Tectariaceae) =Tectaria plantaginea 

Polypodium plumula Humb. & Bonpl. ex Willd. =Pecluma plumula


Polypodium plumulum Humb. & Bonpl. ex Willd. =Pecluma plumula 

Polypodium pressum Brause (Grammitidaceae) =Ceradenia kalbreyeri 

Polypodium procerum Willd. (Cyatheaceae) =Cyathea pungens 

Polypodium pruinatum Baker (Grammitidaceae) =Ceradenia pruinosa 

Polypodium pruinosum Maxon (Grammitidaceae) =Ceradenia pruinosa 

Polypodium pseudoaureum Cav. =Phlebodium pseudoaureum 

Polypodium pseudofraternum A.C. Sm. =Polypodium sessilifolium 

Polypodium pteroideum Klotzsch (Thelypteridaceae) =Thelypteris pteroidea 

Polypodium pteropus Hook. (Grammitidaceae) =Grammitis subsessilis 

Polypodium ptilodon Kunze =Pecluma ptilodon 

Polypodium ptilodon var. pilosum A.M. Evans =Pecluma ptilodon var. pilosa 

Polypodium pungens Willd. (Cyatheaceae) =Cyathea pungens 

Polypodium repens Aubl. =Campyloneurum repens 

Polypodium rigens Maxon (Grammitidaceae) =Grammitis xiphopteroides 

Polypodium rigescens Bory ex Willd. (Grammitidaceae) =Grammitis flabelliformis 

Polypodium rivulare Vahl (Oleandraceae) =Nephrolepis rivularis 

Polypodium roraimense Brause (Grammitidaceae) =Ceradenia kalbreyeri 

Polypodium roraimense Baker (Thelypteridaceae) =Thelypteris roraimensis 

Polypodium rostratum Willd. (Metaxyaceae) =Metaxya rostrata 

Polypodium rude Kunze (Thelypteridaceae) =Thelypteris rudis 

Polypodium sancti-gabrieli Hook. (Dryopteridaceae) =Cyclodium guianense 

Polypodium sectifrons Kunze ex Mett. (Grammitidaceae) =Zygophlebia sectifrons 

Polypodium semicordatum Sw. (Tectariaceae) =Cyclopeltis semicordata 

Polypodium serrulatum (Sw.) Mett. (Grammitidaceae) =Cochlidium serrulatum 

Polypodium serrulatum var. strictissimum Hook. (Grammitidaceae) =Grammitis myosuroides 

Polypodium sororium Humb. & Bonpl. ex Willd. =Polypodium dulce 

Polypodium sphenodes Kunze [not in flora area] =Campyloneurum sphenodes (Kunze) Fée 

Polypodium spixianum Mart. ex Mett. (Grammitidaceae) =Ceradenia spixiana 

Polypodium sprucei Hook. (Grammitidaceae) =Enterosora parietina 

Polypodium stahelianum Posth. (Grammitidaceae) =Grammitis staheliana 

Polypodium strictissimum (Hook.) Hieron. (Grammitidaceae) =Grammitis myosuroides 

Polypodium subincisum Willd. (Tectariaceae) =Megalastrum subincisum 

Polypodium subobliquatum Hook. (Dryopteridaceae) =Cyclodium inerme 

Polypodium subsessile Baker (Grammitidaceae) =Grammitis subsessilis 

Polypodium subtetragonum Link (Thelypteridaceae) =Thelypteris tetragona 

Polypodium surinamensis Jacq. =Microgramma lycopodioides 

Polypodium suspensum L. (Grammitidaceae) =Grammitis suspensa 

Polypodium taenifolium Jenman (Grammitidaceae) =Grammitis taenifolia 

Polypodium taxifolium L. (Grammitidaceae) =Grammitis taxifolia 

Polypodium tectum Kaulf. =Microgramma tecta 

Polypodium tenuiculum Fée (Grammitidaceae) =Grammitis tenuicula 

Polypodium tenuiculum var. acrosorum Hieron. (Grammitidaceae) =Grammitis major 

Polypodium tepuiense A.C. Sm. (Grammitidaceae) =Cochlidium tepuiense 

Polypodium tetragonum Sw. (Thelypteridaceae) =Thelypteris tetragona 

Polypodium thurnii Baker =Microgramma thurnii 

Polypodium tottum Thunb. (Thelypteridaceae) =Thelypteris interrupta 

Polypodium transiens Lindm. (Grammitidaceae) =Ceradenia spixiana 

Polypodium trichomanoides Sw. (Grammitidaceae) =Grammitis trichomanoides 

Polypodium trifoliatum L. (Tectariaceae) =Tectaria trifoliata 

Polypodium trifurcatum L. (Grammitidaceae) =Enterosora trifurcata 

Polypodium triste Kunze (Thelypteridaceae) =Thelypteris tristis 

Polypodium truncicola Klotzsch (Grammitidaceae) =Grammitis truncicola 

Polypodium xanthotrichium Klotzsch (Grammitidaceae) =Grammitis xanthotrichia 

Polypodium xiphopteroides Liebm. (Grammitidaceae) =Grammitis xiphopteroides 

Polypodium yarumalense Hieron. (Grammitidaceae) =Enterosora parietina 

Taenitis desvauxii Klotzsch =Dicranoglossum desvauxii 

REFERENCES 

Lellinger, D.B. 1972. A revision of the fern genus Niphidium. Amer. Fern J. 62: 101-120. 

Lellinger, D.B. 1988. Some new species of Campyloneurum and a provisional key to the genus. Amer. Fern J. 78(1): 14-35. 

Price, M.G. 1983. Pecluma, a new tropical American fern genus. Amer. Fern J. 73(3): 109-116. 

Smith, A.R. 1995. Polypodiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 219-249.

50 


PTERIDACEAE 

Acrostichum aureum L. 

DA GU SU FG 

Acrostichum danaeifolium Langsd. & Fisch. [expected in DA] 

GU SU FG 

Pteris altissima Poir. 


Pteris biaurita L. 

GU SU FG 

Pteris deflexa Link 

GU SU FG 

Pteris denticulata Sw. 

GU 

Pteris pearcei Baker 

AM BO GU 

Pteris plumula Desv. 

FG 

Pteris podophylla Sw. 

AM 

Pteris polita Link [see also Pteris propinqua] 

SU FG 

Pteris propinqua J. Agardh [=Pteris polita?] 

AM BO SU FG 

Pteris pungens Willd. 


Pteris quadriaurita Retz. [neotropical material =P. plumula, fide D. Lellinger] BO FG 

Pteris splendens Kaulf. 

GU 

Pteris tripartita Sw. [naturalized] 

DA SU FG 

Pteris vittata L. [naturalized] 

SU 


Acrostichum acrocarpum Mart. (Lomariopsidaceae) =Elaphoglossum acrocarpum 

Acrostichum alatum Fée (Lomariopsidaceae) =Elaphoglossum pteropus 

Acrostichum andicola Fée (Lomariopsidaceae) =Elaphoglossum andicola 

Acrostichum auricomum Kunze (Lomariopsidaceae) =Elaphoglossum auricomum 

Acrostichum bonariense Willd. (Adiantaceae) =Cheilanthes bonariensis 

Acrostichum brachyneuron Fée (Lomariopsidaceae) =Elaphoglossum brachyneuron 

Acrostichum calomelanos L. (Adiantaceae) =Pityrogramma calomelanos 

Acrostichum chrysolepis Fée (Lomariopsidaceae) =Elaphoglossum plumosum 

Acrostichum citrifolium L. (Vittariaceae) =Anetium citrifolium 

Acrostichum cladorrhizans Spreng. (Lomariopsidaceae) =Bolbitis portoricensis 

Acrostichum compactum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis 

Acrostichum corpulentum Kunze ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis 

Acrostichum crinitum L. (Lomariopsidaceae) =Elaphoglossum crinitum 

Acrostichum cuspidatum Willd. (Lomariopsidaceae) =Elaphoglossum cuspidatum 

Acrostichum decoratum Kunze (Lomariopsidaceae) =Elaphoglossum decoratum 

Acrostichum discolor Kuhn (Lomariopsidaceae) =Elaphoglossum discolor 

Acrostichum ebeneum L. (Adiantaceae) =Pityrogramma ebenea 

Acrostichum elegans Vahl (Schizaeaceae) =Schizaea elegans 

Acrostichum erinaceum Fée (Lomariopsidaceae) =Elaphoglossum erinaceum 

Acrostichum eximium Mett. (Lomariopsidaceae) =Elaphoglossum eximium 

Acrostichum flaccidum Fée (Lomariopsidaceae) =Elaphoglossum flaccidum 

Acrostichum fraseri Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum pusillum 

Acrostichum gayanum (Fée) T. Moore (Lomariopsidaceae) =Elaphoglossum minutum 

Acrostichum hayesii Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum hayesii 

Acrostichum herminieri Bory ex Fée (Lomariopsidaceae) =Elaphoglossum herminieri 

Acrostichum hoffmannii Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum hoffmannii 

Acrostichum horridulum Kaulf. (Lomariopsidaceae) =Elaphoglossum horridulum 

Acrostichum japurense Mart. (Lomariopsidaceae) =Lomariopsis japurensis 

Acrostichum laminarioides Fée (Lomariopsidaceae) =Elaphoglossum laminarioides 

Acrostichum langsdorffii Hook. & Grev. (Lomariopsidaceae) =Elaphoglossum langsdorffii 

Acrostichum leptophlebium Baker (Lomariopsidaceae) =Elaphoglossum leptophlebium 

Acrostichum leptophyllum Fée (Lomariopsidaceae) =Elaphoglossum minutum 

Acrostichum luridum Fée (Lomariopsidaceae) =Elaphoglossum luridum 

Acrostichum macrophyllum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum macrophyllum 

Acrostichum meridense Klotzsch (Lomariopsidaceae) =Elaphoglossum meridense 

Acrostichum minutum Pohl ex Fée (Lomariopsidaceae) =Elaphoglossum minutum 

Acrostichum nicotianifolium Sw. (Lomariopsidaceae) =Bolbitis nicotianifolia 

Acrostichum nigrescens Hook. (Lomariopsidaceae) =Elaphoglossum nigrescens 

Acrostichum nodosum L. (Marattiaceae) =Danaea nodosa 

Acrostichum paleaceum Hook. & Grev. (Lomariopsidaceae) =Elaphoglossum paleaceum 

Acrostichum papillosum Baker (Lomariopsidaceae) =Elaphoglossum papillosum 

Acrostichum piloselloides C. Presl (Lomariopsidaceae) =Elaphoglossum piloselloides 

Acrostichum plumosum Fée (Lomariopsidaceae) =Elaphoglossum plumosum


Acrostichum polypodioides L. (Polypodiaceae) =Polypodium polypodioides 

Acrostichum portoricense Spreng. (Lomariopsidaceae) =Bolbitis portoricensis 

Acrostichum pusillum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum pusillum 

Acrostichum raywaense Jenman (Lomariopsidaceae) =Elaphoglossum raywaense 

Acrostichum reichenbachii Moritz ex Kuhn (Lomariopsidaceae) =Elaphoglossum eximium 

Acrostichum reptans Cav. (Polypodiaceae) =Microgramma reptans 

Acrostichum rufum (L.) L. (Adiantaceae) =Hemionitis rufa 

Acrostichum schomburgkii Fée (Lomariopsidaceae) =Elaphoglossum luridum 

Acrostichum serratifolium Mert. ex Kaulf. (Lomariopsidaceae) =Bolbitis serratifolia 

Acrostichum serrulatum Sw. (Grammitidaceae) =Cochlidium serrulatum 

Acrostichum sporadolepis Kunze ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis 

Acrostichum squamosum Sw. (Lomariopsidaceae) =Elaphoglossum paleaceum 

Acrostichum squarrosum Klotzsch (Lomariopsidaceae) =Elaphoglossum squarrosum 

Acrostichum stenopteris Klotzsch (Lomariopsidaceae) =Elaphoglossum brachyneuron 

Acrostichum tartareum Cav. (Adiantaceae) =Pityrogramma ebenea 

Acrostichum tectum Humb. & Bonpl. ex Willd. (Lomariopsidaceae) =Elaphoglossum tectum 

Acrostichum thalictroides L. (Parkeriaceae) =Ceratopteris thalictroides 

Acrostichum tovarense Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum tovarense 

Acrostichum trifoliatum L. (Adiantaceae) =Pityrogramma trifoliata 

Litobrochia podophylla (Sw.) C. Presl =Pteris podophylla 

Litobrochia tripartita (Sw.) C. Presl =Pteris tripartita 

Neurogramme elaphoglossoides (Baker) H. Christ (Adiantaceae) =Pterozonium elaphoglossoides 

Ormopteris pinnata (Kaulf.) Lellinger (Adiantaceae) =Cassebeera pinnata 

Pteris angustifolia Sw. (Vittariaceae) =Ananthacorus angustifolius 

Pteris arachnoidea Kaulf. (Dennstaedtiaceae) =Pteridium arachnoideum 

Pteris aurea Poir. (Adiantaceae) =Cheilanthes bonariensis 

Pteris caudata L. (Dennstaedtiaceae) =Pteridium caudatum 

Pteris concolor Langsd. & Fisch. (Adiantaceae) =Cheilanthes concolor 

Pteris incisa Thunb. (Dennstaedtiaceae) =Histiopteris incisa 

Pteris interrupta Willd. (Thelypteridaceae) =Thelypteris interrupta 

Pteris kunzeana J. Agardh =Pteris altissima 

Pteris lindeniana (Hook.) H. Christ (Dennstaedtiaceae) =Blotiella lindeniana 

Pteris lineata L. (Vittariaceae) =Vittaria lineata 

Pteris lucida Cav. (Adiantaceae) =Adiantum lucidum 

Pteris palmata Willd. (Adiantaceae) =Doryopteris pedata var. palmata 

Pteris pedata L. (Adiantaceae) =Doryopteris pedata 

Pteris petiolulata R.M. Tryon =Pteris pearcei 

Pteris sagittifolia Raddi (Adiantaceae) =Doryopteris sagittifolia 

REFERENCES 

Smith, A.R. & D.B. Lellinger. 1995. Pteridaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 

2: 250-286. 

Azolla caroliniana Willd. 

Azolla mexicana C. Presl 

Azolla microphylla Kaulf. 

Salvinia auriculata Aubl. 

Salvinia radula Baker 

SALVINIACEAE 


Salvinia laevigata Humb. & Bonpl. ex Willd. (Hydrocharitaceae) =Limnobium laevigatum 

Salvinia rotundifolia Willd. =Salvinia auriculata 


GU 

GU 


GU SU FG 

REFERENCES 

Reed, C.F. 1965a. Distribution of Salvinia and Azolla in South America and Africa in connection with studies for control by 

insects. Phytologia 12(3): 121-130. 

Reed, C.F. 1965b. Index Marsileata et Salviniata. Suppl. Bol. Soc. Brot. II, 39: 259-302. 

Smith, A.R. 1995. Azollaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 22-23. 

Smith, A.R. 1995. Salviniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 287-288. 

Svenson, H.K. 1944. The New World species of Azolla. Amer. Fern J. 34: 69-84.

52 

Actinostachys pennula (Sw.) Hook. 

Actinostachys subtrijuga (Mart.) C. Presl 

Anemia ayacuchensis Mickel 

Anemia buniifolia (Gardner) T. Moore 

Anemia collina Raddi 

Anemia ferruginea Kunth 

Anemia ferruginea var. ahenobarba (H. Christ) Mickel 

Anemia ferruginea var. ferruginea 

Anemia flexuosa (Sav.) Sw. 

Anemia hirsuta (L.) Sw. 

Anemia hirta (L.) Sw. 

Anemia mandioccana Hook. 

Anemia millefolia (Gardner) C. Presl 

Anemia oblongifolia (Cav.) Sw. 

Anemia pastinacaria Moritz ex Prantl 

Anemia phyllitidis (L.) Sw. 

Anemia porrecta Mickel 

Schizaea elegans (Vahl) Sw. 

Schizaea fluminensis Miers ex J.W. Sturm 

Schizaea incurvata Schkuhr 

Schizaea poeppigiana J.W. Sturm 

Schizaea sprucei Hook. 

Schizaea stricta Lellinger 


SCHIZAEACEAE 


Actinostachys penicillata (Humb. & Bonpl. ex Willd.) Maxon =Actinostachys pennula 

Anemia humilis (Cav.) Sw. =Anemia oblongifolia 

Anemia villosa Humb. & Bonpl. ex Willd. [not in flora area] =Anemia flexuosa 

Coptophyllum buniifolium Gardner =Anemia buniifolia 

Coptophyllum millefolium Gardner =Anemia millefolia 

Lophidium elegans (Vahl) C. Presl =Schizaea elegans 

Lophidium poeppigianum (J.W. Sturm) Underw. =Schizaea poeppigiana 

Schizaea flabellum Mart. =Schizaea elegans 

Schizaea pennula Sw. =Actinostachys pennula 

Schizaea subtrijuga Mart. =Actinostachys subtrijuga 

REFERENCES 

Lellinger, D.B. 1969. Schizaeaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 8. Mem. New York 

Bot. Gard. 18(2): 1-290. 

Smith, A.R. 1995. Schizaeaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 288-296. 

Ctenitis falciculata (Raddi) Ching 

Ctenitis paranaensis (C. Chr.) Lellinger 

Ctenitis refulgens (Klotzsch ex Mett.) C. Chr. ex Vareschi 

Cyclopeltis semicordata (Sw.) J. Sm. 

Lastreopsis amplissima (C. Presl) Tindale 

Lastreopsis effusa (Sw.) Tindale 

Lastreopsis effusa subsp. divergens (Willd.) Tindale 

Lastreopsis killipii (C. Chr. & Maxon) Tindale 

Megalastrum crenulans (Fée) A.R. Sm. & R.C. Moran 

Megalastrum subincisum (Willd.) A.R. Sm. & R.C. Moran 

Tectaria antioquioana (Baker) C. Chr. 

Tectaria heracleifolia (Willd.) Underw. 

Tectaria incisa Cav. 

Tectaria lizarzaburui (Sodiro) C. Chr. 

Tectaria plantaginea (Jacq.) Maxon 

Tectaria plantaginea var. macrocarpa (Fée) C.V. Morton 

Tectaria plantaginea var. plantaginea 

Tectaria transiens (C.V. Morton) A.R. Sm. 

Tectaria trifoliata (L.) Cav. 

TECTARIACEAE 


AM GU SU FG 

AM 

AM BO SU 

GU 

AM BO GU SU 

SU 

AM BO GU SU 


GU 

GU SU 

GU 

BO GU 

AM BO GU SU 

BO GU SU FG 

GU SU 

AM 




AM BO GU 

AM 

AM BO GU 

GU 

GU 



AM BO GU 

BO GU SU FG 

BO GU SU FG 

AM GU? 

AM BO 

BO 

AM 

GU SU 


AM 



AM BO 

GU FG 

BO DA GU SU FG

Tectaria trinitensis Maxon 

Triplophyllum acutilobum Holttum 

Triplophyllum angustifolium Holttum 

Triplophyllum dicksonioides (Fée) Holttum 

Triplophyllum funestum (Kunze) Holttum 



Bathmium macrocarpon Fée =Tectaria plantaginea var. macrocarpa 

Bathmium sinuatum Fée =Tectaria plantaginea var. macrocarpa 

Ctenitis crenulans (Fée) Ching =Megalastrum crenulans 

Ctenitis effusa (Sw.) Copel. =Lastreopsis effusa 

Ctenitis funesta (Kunze) Vareschi =Triplophyllum funestum 

Ctenitis protensa (Afzel. ex Sw.) Ching var. funesta (Kunze) Proctor =Triplophyllum funestum 

Ctenitis subincisa (Willd.) Ching =Megalastrum subincisum 

Lastreopsis effusa subsp. effusa [not in flora area] 

Phegopteris refulgens Klotzsch ex Mett. =Ctenitis refulgens 

Tectaria martinicensis (Spreng.) Copel. =Tectaria incisa 

Triplophyllum crassifolium Holttum =Triplophyllum funestum 

Triplophyllum crassifolium var. caudatum Holttum =Triplophyllum funestum 

Triplophyllum funestum var. hirsutum Holttum =Triplophyllum funestum 

REFERENCES 

(Included in Dryopteridaceae in Flora of the Venezuelan Guayana and Flora of the Guianas) 


(series B) 6: 3-65. 

Holttum, R.E. 1986. Studies in the fern genera allied to Tectaria Cav. V. Triplophyllum, a new genus of Africa and America. Kew 

Bull. 41: 237-260. 



Tindale, M.D. 1965. A monograph of the genus Lastreopsis Ching. Contr. New South Wales Nat. Herb. 3: 249-339. 

THELYPTERIDACEAE 

Thelypteris abrupta (Desv.) Proctor 

Thelypteris angustifolia (Willd.) Proctor 

Thelypteris arborea (Brause) A.R. Sm. [Mt. Roraima] 

Thelypteris arborescens (Humb. & Bonpl. ex Willd.) C.V. Morton 

Thelypteris balbisii (Spreng.) Ching 

Thelypteris binervata A.R. Sm. 

Thelypteris biolleyi (H. Christ) Proctor 

Thelypteris brachypoda (Baker) C.V. Morton [Mt. Roraima] 

Thelypteris chrysodioides (Fée) C.V. Morton 

Thelypteris chrysodioides var. goyazensis (Maxon & C.V. Morton) C.V. 

Morton 

Thelypteris decussata (L.) Proctor 

Thelypteris decussata var. decussata 

Thelypteris deflexa (C. Presl) R.M. Tryon 

Thelypteris demerarana (Baker) C.F. Reed [Mt. Roraima] 

Thelypteris funckii (Mett.) Alston [Mt. Roraima] 

Thelypteris gardneriana (Baker) C.F. Reed 

Thelypteris gemmulifera (Hieron.) A.R. Sm. 

Thelypteris germaniana (Fée) Proctor 

Thelypteris glandulosa (Desv.) Proctor 

Thelypteris glandulosa var. glandulosa 

Thelypteris gonophora (Weath.) A.R. Sm. 

Thelypteris hispidula (Decne.) C.F. Reed 

Thelypteris hispidula var. hispidula 

Thelypteris holodictya K.U. Kramer 

Thelypteris interrupta (Willd.) K. Iwats. 

Thelypteris juruensis (C. Chr.) R.M. Tryon & D.S. Conant 

Thelypteris kunthii (Desv.) C.V. Morton 

Thelypteris leprieurii (Hook.) R.M. Tryon 

Thelypteris leprieurii var. leprieurii 

Thelypteris linkiana (C. Presl) R.M. Tryon [Mt. Roraima] 

Thelypteris macrophylla (Kunze) C.V. Morton 

AM GU SU 

GU? 

GU SU FG 




DA 

AM BO GU 


BO GU 

AM 

AM 

BO GU 

BO GU 

BO 

AM 

AM 

BO GU SU FG 

BO GU SU FG 

BO GU? 

GU 

FG 

FG 

GU SU FG 

GU SU FG 

GU 



SU FG 

BO GU SU FG 

FG 

GU 



BO GU 

AM GU SU FG

54 

Thelypteris nephrodioides (Klotzsch) Proctor 

Thelypteris nesiotica (Maxon & C.V. Morton) C.V. Morton 

Thelypteris oligocarpa (Humb. & Bonpl. ex Willd.) Ching 

Thelypteris opposita (Vahl) Ching 

Thelypteris opulenta (Kaulf.) Fosberg [naturalized] 

Thelypteris pennata (Poir.) C.V. Morton 

Thelypteris peradenia A.R. Sm. [Mt. Roraima] 

Thelypteris pilosissima C.V. Morton [Mt. Roraima] 

Thelypteris poiteana (Bory) Proctor 

Thelypteris pteroidea (Klotzsch) R.M. Tryon [Mt. Roraima] 

Thelypteris x rolandii (C. Chr.) R.M. Tryon 

Thelypteris roraimensis (Baker) C.F. Reed [Mt. Roraima] 

Thelypteris rudis (Kunze) Proctor 

Thelypteris salzmannii (Fée) C.V. Morton 

Thelypteris schomburgkii A.R. Sm. 

Thelypteris serrata (Cav.) Alston 

Thelypteris tetragona (Sw.) Small 

Thelypteris tristis (Kunze) R.M. Tryon 



Amauropelta deflexa (C. Presl) A. Löve & D. Löve =Thelypteris deflexa 

Amauropelta diplazioides (Desv.) Pic. Serm. =Thelypteris linkiana 

Amauropelta oligocarpa (Humb. & Bonpl. ex Willd.) Pic. Serm. =Thelypteris oligocarpa 

Amauropelta opposita (Vahl) Pic. Serm. =Thelypteris opposita 

Amauropelta rudis (Kunze) Pic. Serm. =Thelypteris rudis 

Amphineuron opulentum (Kaulf.) Holttum =Thelypteris opulenta 

Christella hispidula (Decne.) Holttum =Thelypteris hispidula 

Cyclosorus gongylodes (Schkuhr) Link =Thelypteris interrupta 

Glaphyropteris decussata (L.) Fée =Thelypteris decussata 

Goniopteris biolleyi (H. Christ) Pic. Serm. =Thelypteris biolleyi 

Goniopteris juruensis (C. Chr.) Brade =Thelypteris juruensis 

Goniopteris megalodus (Schkuhr) C. Presl =Thelypteris pennata 

Goniopteris pennata (Poir.) Pic. Serm. =Thelypteris pennata 

Goniopteris poiteana (Bory) Ching =Thelypteris poiteana 

Goniopteris pyramidata Fée =Thelypteris abrupta 

Goniopteris subtetragona (Link) Vareschi =Thelypteris tetragona 

Goniopteris tetragona (Sw.) C. Presl =Thelypteris tetragona 

Goniopteris tristis (Kunze) Brade =Thelypteris tristis 

Lastrea adiantiformis (G. Forst.) Vareschi (Davalliaceae) =Rumohra adiantiformis 

Lastrea brachypoda (Baker) Vareschi =Thelypteris brachypoda 

Lastrea coarctata (Kunze) T. Moore =Thelypteris opposita 

Lastrea demerarana (Baker) Vareschi =Thelypteris demerarana 

Lastrea denticulata (Sw.) T. Moore (Dryopteridaceae) =Arachniodes denticulata 

Lastrea engelii (Hieron.) Vareschi [nomen nudum] =Thelypteris rudis 

Lastrea lindigii (C. Chr.) Vareschi =Thelypteris deflexa 

Lastrea linkiana (C. Presl) Copel. =Thelypteris linkiana 

Lastrea nervosa (Klotzsch) Vareschi [comb. illeg.] =Thelypteris rudis 

Lastrea oligocarpa (Humb. & Bonpl. ex Willd.) T. Moore =Thelypteris oligocarpa 

Lastrea opposita (Vahl) Gilli =Thelypteris opposita 

Lastrea pilosissima (C.V. Morton) Vareschi =Thelypteris pilosissima 

Lastrea poiteana Bory =Thelypteris poiteana 

Lastrea roraimensis (Baker) Brade =Thelypteris roraimensis 

Meniscium angustifolium Willd. =Thelypteris angustifolia 

Meniscium arborescens Humb. & Bonpl. ex Willd. =Thelypteris arborescens 

Meniscium chyrsodioides Fée =Thelypteris chrysodioides 

Meniscium cristatum Desr. (Woodsiaceae) =Diplazium cristatum 

Meniscium guyanense Fée =Thelypteris macrophylla 

Meniscium macrophyllum Kunze =Thelypteris macrophylla 

Meniscium serratum Cav. =Thelypteris serrata 

Steiropteris gardneriana (Baker) Pic. Serm. =Thelypteris gardneriana 

Steiropteris glandulosa (Desv.) Pic. Serm. =Thelypteris glandulosa 

Steiropteris leprieurii (Hook.) Pic. Serm. =Thelypteris leprieurii 

Thelypteris coarctata (Kunze) R.M. Tryon =Thelypteris opposita 

Thelypteris connexa (Kuhn ex Baker) C.F. Reed [not in flora area, type erroneously cited from FG] 

Thelypteris extensa (Blume) C.V. Morton =Thelypteris opulenta 

GU FG 

BO GU 

BO 

BO 

AM BO GU SU FG? 


BO 

BO 

GU SU FG 

AM BO 

AM 

BO GU 

AM BO GU 

GU 

GU 

DA GU SU FG 

DA GU SU FG 

AM BO DA GU SU FG


Thelypteris gongylodes (Schkuhr) Small =Thelypteris interrupta 

Thelypteris guyanensis (Fée) C.V. Morton =Thelypteris macrophylla 

Thelypteris hostmannii (Klotzsch) C.V. Morton =Thelypteris arborescens 

Thelypteris lindigii (C. Chr.) Alston =Thelypteris deflexa 

Thelypteris megalodus (Schkuhr) Proctor =Thelypteris pennata 

Thelypteris mercurii (A. Braun ex Hieron.) C.F. Reed =Thelypteris balbisii 

Thelypteris nervosa (Klotzsch) R.M. Tryon =Thelypteris rudis 

Thelypteris quadrangularis (Fée) Schelpe =Thelypteris hispidula 

Thelypteris sprengelii (Kaulf.) Proctor =Thelypteris balbisii 

Thelypteris totta (Thunb.) Schelpe =Thelypteris interrupta 

REFERENCES 

Smith, A.R. 1994. Thelypteridaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series B) 6: 77-115. 

Smith, A.R. 1995. Thelypteridaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 315-326. 

Ananthacorus angustifolius (Sw.) Underw. & Maxon 

Anetium citrifolium (L.) Splitg. 

Antrophyum cajenense (Desv.) Spreng. 

Antrophyum guayanense Hieron. 

Antrophyum jenmanii Benedict 

Antrophyum lanceolatum (L.) Kaulf. 

Antrophyum lineatum (Sw.) Kaulf. 

Antrophyum ophioglossoides Lellinger 

Hecistopteris kaieteurensis Kelloff & G.S. McKee 

Hecistopteris pumila (Spreng.) J. Sm. 

Vittaria gardneriana Fée 

Vittaria graminifolia Kaulf. 

Vittaria lineata (L.) Sm. 

Vittaria moritziana Mett. 

Vittaria remota Fée 

Vittaria stipitata Kunze 

VITTARIACEAE 


Gymnogramme pumila Spreng. =Hecistopteris pumila 

Polytaenium brasilianum (Desv.) Benedict [records due to misidentification fide Lellinger] 

Polytaenium cajenense (Desv.) Benedict =Antrophyum cajenense 

Polytaenium guayanense (Hieron.) Alston =Antrophyum guayanense 

Polytaenium lineatum (Sw.) J. Sm. =Antrophyum lineatum 

Pteridanetium citrifolium (L.) Copel. =Anetium citrifolium 

Vittaria angustifolia (Sw.) Baker =Ananthacorus angustifolius 

Vittaria costata Kunze =Ananthacorus angustifolius 

Vittaria filifolia Fée =Vittaria graminifolia 

REFERENCES 

Lellinger, D.B. 1972. Polypodiaceae subfam. Vittarioideae. In B. Maguire & colaborators, Botany of the Guayana Highland part 

9. Mem. New York Bot. Gard. 23: 1-832. 

Smith, A.R. 1995. Vittariaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 327-334. 

WOODSIACEAE 

Diplazium celtidifolium Kunze 

Diplazium centripetale (Baker) Maxon [Mt. Roraima] 

Diplazium cristatum (Desr.) Alston 

Diplazium expansum Willd. 

Diplazium gracilescens (Mett.) C. Chr. 

Diplazium grandifolium (Sw.) Sw. 

Diplazium hians Kunze ex Klotzsch 

Diplazium lechleri (Mett.) T. Moore 

Diplazium plantaginifolium (L.) Urb. 

Diplazium radicans Desv. 

Diplazium riedelianum Kunze 

Diplazium roraimense Cremers & K.U. Kramer [Mt. Roraima] 

Diplazium striatum (L.) C. Presl 





GU SU FG 

GU SU FG 

AM BO 

AM 

GU 


AM BO GU SU 

AM BO GU SU 


GU 


BO FG 

AM GU SU FG 

AM BO GU 


GU FG 

SU FG 

AM BO GU FG 

AM 

AM BO GU SU 

AM GU 

SU FG 

GU 

BO GU 

GU FG

56 

Diplazium tabalosense Hieron. 

Diplazium wilsonii (Baker) Diels 

Hemidictyum marginatum (L.) C. Presl 



Diplazium arboreum (Willd.) C. Presl =Diplazium cristatum 

Diplazium callipteris Fée =Diplazium celtidifolium 

Diplazium delitescens Maxon (Aspleniaceae) =Asplenium delitescens 

Diplazium shepherdii (Spreng.) Link =Diplazium cristatum 

REFERENCES 

(Included in Dryopteridaceae in Flora of the Venezuelan Guayana and in Flora of the Guyanas) 


(series B) 6: 3-65. 



Pinus caribaea Morelet [cultivated, naturalized] 

GYMNOSPERMS 

D.J. de Laubenfels, D.W. Stevenson 

CONIFEROPHYTA 

PINACEAE 

REFERENCES 

Stevenson, D. & T. Zanoni. 1991. Pinaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 9: 19-22. 

PODOCARPACEAE 

Podocarpus acuminatus de Laub. 

Podocarpus aracensis de Laub. & Silba 

Podocarpus brasiliensis de Laub. [expected in GU] 

Podocarpus buchholzii de Laub. [expected in GU] 

Podocarpus celatus de Laub. 

Podocarpus magnifolius J. Buchholz & N.E. Gray [expected in GU] 

Podocarpus roraimae Pilg. [Mt. Roraima] 

Podocarpus steyermarkii J. Buchholz & N.E. Gray 

Podocarpus tepuiensis J. Buchholz & N.E. Gray 

REFERENCES 

Laubenfels, D.J. de. 1982. Podocarpaceae. In Flora de Venezuela 11(2): 7-41. 

Laubenfels, D.J. de. 1985. A taxonomic revision of the genus Podocarpus. Blumea 30: 251-278. 

Laubenfels, D.J. de. 2004. Podocarpaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 297-300. 

Stevenson, D. 1991. Podocarpaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 9: 23-30. 

CYCADOPHYTA 

CYCADACEAE 

Cycas spp. [species of this Old World family are widely cultivated] 

REFERENCES 

Stevenson, D.W. 1991. Cycadaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 9: 3-7. 

Zamia amazonum D.W. Stev. 

Zamia lecointei Ducke [expected in the Guianas] 

Zamia muricata Willd. [expected in the Guianas] 

Zamia ulei Dammer [expected in the Guianas] 

ZAMIACEAE 


Zamia jirijirmensis R.E. Schult. =Zamia lecointei 

Zamia obidensis Ducke =Zamia lecointei 

Zamia ulei subsp. lecointei (Ducke) Ducke =Zamia lecointei 

AM 

BO 

AM SU FG 

AM BO 

AM BO 

AM BO 

AM BO 

AM BO 

AM BO 

AM BO GU? 

AM BO GU 

AM BO GU 

AM 

AM BO 

BO 

GU SU FG


REFERENCES 

Jones, D.L. 1993. Cycads of the World. Washington, DC: Smithsonian Institution Press. 

Sabato, S. 1990. West Indian and South American Cycads. Mem. New York Bot. Gard. 57: 173-185. 

Stevenson, D. 1991. Zamiaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 9: 7-11. 

Gnetum camporum (Markgr.) D.W. Stev. & T. Zanoni 

Gnetum leyboldii Tul. 

Gnetum nodiflorum Brongn. 

Gnetum paniculatum Spruce ex Benth. 

Gnetum schwackeanum Taub. ex Markgr. 

Gnetum urens (Aubl.) Blume 

GNETOPHYTA 

GNETACEAE 


Gnetum amazonicum Tul. =Gnetum nodiflorum 

Gnetum cruzianum Gleason =Gnetum nodiflorum 

Gnetum leyboldii var. woodsonianum Markgr. =Gnetum urens 

Gnetum melinonii Benoist =Gnetum urens 

Gnetum microstachyum Benth. ex Tul. =Gnetum paniculatum 

Gnetum oblongifolium Huber =Gnetum nodiflorum 

Gnetum paraense Huber =Gnetum leyboldii 

Gnetum thoa Brongn. [nom. illeg.] =Gnetum urens 

Gnetum urens var. camporum Markgr. =Gnetum camporum 

Thoa urens Aubl. =Gnetum urens 

REFERENCES 

Markgraf, F. 1965. New discoveries of Gnetum in tropical America. Ann. Missouri Bot. Gard. 52: 379-386. 

Stevenson, D.W. 1997. Gnetaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 


of the New York Botanical Garden 76, pt. 1: 162 

Stevenson, D.W. 1999. Gnetaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 573-576. 

Stevenson, D. & T. Zanoni. 1991. Gnetaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 9: 12-18. 

INCERTAE SEDIS 

Aubletella macoucou (Aubl.) Pierre [excluded from Sapotaceae. 

Chrysophyllum macoucou Aubl.?] 

Macahanea guianensis Aubl. [excluded from Hippocrateaceae. Salacia 

guianensis (Aubl.) Lemée?] 

Managa guianensis Aubl. [excluded from Hippocrateaceae] 

Sarcaulus melinonianum Pierre [name of unknown origin, placed by Lemée in 

Myrsinaceae] 

Senapea guianensis Aubl. [excluded from Passifloraceae] 

Tampoa guianensis Aubl. [excluded from Hippocrateaceae] 

Vouapa simira Aubl. [excluded from Leguminosae-Caesalp. Macrolobium 

simira (Aubl.) J.F. Gmelin?] 

ANTHOPHYTA 

BO GU 

AM BO GU 


AM BO GU FG 

AM GU 



Cremers, G. 1985. Espèces nouvelles de Guyane française: richesse de l’herbier de Cayenne (CAY). Proc. Kon. Ned. Akad. 

Wet. ser. C, 88(1): 15-37. 

Cremers, G., C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des espèces de Phanérogames et de Ptéridophytes de Guyane 

française d’après l’Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne. 

Cremers, G., J.J. de Granville, & M. Hoff. 1994. Les plantes “endémiques” de Guyane française. Mus. Nat. Hist. Nat., Coll. 

Patr. Nat. (Sér. Patr. Génétique) 18: 1-50. 

Cremers, G. & M. Hoff. 1994. Inventaire taxonomique des plantes de la Guyane française. IV - Les Monocotylédones. Mus. 

Nat. Hist. Nat., Coll. Patr. Nat. (Sér. Patr. Génétique) 17: 1-252. 

DeFilipps, R.A. 1992. Ornamental Garden plants of the Guianas: An Historical Perspective of Selected Garden Plants from 

Guyana, Surinam and French Guiana. 363 pp. Washington, D.C.: Smithsonian Institution. 

Fanshawe, D.B. 1952. The vegetation of British Guiana: A preliminary review. Imperial Forestry Institute paper no. 29: 1-96. 

FG 

FG 

FG 

FG 

FG 

FG 

FG

58 


Funk, V.A. & C.L. Kelloff. 1991. A Preliminary Checklist of the Flowering Plants and Ferns of Kaieteur Falls National Park 

Guyana. Unpublished manuscript. Biological Diversity of the Guianas program, Smithsonian Institution, Washington, DC. 

Gleason, H.A. 1929. Studies on the flora of northern South America - XIII. The Tate collection from Mount Roraima and 

vicinity. Bull. Torr. Bot. Club 56(8): 391-408. 

Graham, E.H. 1934. Flora of the Kartabo Region, British Guiana. Reprinted from Ann. Carn. Mus. 22(1) (1933). 

Howard, R.A. 1983. The plates of Aublet’s Histoire des Plantes de la Guiane Françoise. J. Arn. Arb. 64: 255-292. 

Humboldt, A. von, A. Bonpland, and K.S. Kunth. 1815–1825. Nova Genera et Species Plantarum. 7 volumes. Paris: Sumtibus 

Librariae Graeco-Latino-Germanicae 

Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1989. Botanical explorations in Guyana, 1985. Annex: identifications. Institute of 

Systematic Botany, Utrecht. 

Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1990. Botanical explorations in the northern part of the western Kanuku Mountains 

(Guyana). Annex: identifications. Institute of Systematic Botany, Utrecht. 

Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1992. Botanical exploration in the Wai-Wai area of southern Guyana. Annex: 

identifications. Herbarium Division, Department of Plant Ecology and Evolutionary Biology, University of Utrecht. 

Judd, W.S., C.S. Campbell, E.A. Kellogg, P.F. Stevens, and M.J. Donoghue. 2002. Plant Systematics: A Phylogenetic Approach. 

Sinauer Associates, Inc., Sunderland, MA. 

Lemée, A. 1953-1956. Flore de la Guyane Française. Lechevalier, Paris. 

Lindeman, J.C. 1953. The Vegetation of the Coastal Region of Suriname. Drukkerij en Uitg.-Mij. v/h Kemink, Utrecht. 

Maas, P.J.M. & L.Y.Th. Westra. 1990. Botanical exploration in the northern part of the Rupununi Savanna and in the Mabura Hill 

area (Guyana). Appendix: identifications. Institute of Systematic Botany, Utrecht. 

Maguire, B. & collaborators. 1948. Plant explorations in Guiana in 1944, chiefly to the Tafelberg and the Kaieteur Plateau. Bull. 

Torrey Bot. Club 75: 56-115, 189-230, 286-323, 374-438, 523-580, 633-671. 

Mennega, E.A., W.C.M. Tammens-de Rooij, & M.J. Jansen-Jacobs. 1988. Check-List of Woody Plants of Guyana. The Tropenbos 

Foundation, Ede, The Netherlands. 

Mori, S.A. & collaborators. 1987. The Lecythidaceae of a lowland neotropical forest: La Fumée Mountain, French Guiana. 

Mem. New York Bot. Gard. 44: 1-190. 

Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular 

plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2). 776 pp. 

Omawale. 1973. Guyana’s Edible Plants. University of Guyana. 

Plotkin, M.J., B.M. Boom, & M. Allison. 1991. The ethnobotany of Aublet’s Histoire des Plantes de la Guiane Françoise 

(1775). Monogr. Syst. Bot. 35: 1-108. 

Polak, A.M. 1992. Major Timber Trees of Guyana, a Field Guide. The Tropenbos Foundation, Wageningen. 

Pulle, A. 1906. An Enumeration of the Vascular Plants Known from Surinam, Together with their Distribution and Synonymy. 

E.J. Brill, Ltd., London. 

Roosmalen, M.G.M. van. 1985. Fruits of the Guianan Flora. Institute of Systematic Botany, Utrecht University, Utrecht; 

Silvicultural Department of Wageningen Agricultural University, Wageningen. 

Smith, N., S.A. Mori, A. Henderson, D.W. Stevenson, & S.V. Heald. Flowering Plants of the Neotropics. Princeton University 

Press, Princeton. 616 pp. 

Steege, H. ter, M.J. Jansen-Jacobs, & M. Polak. 1993. Checklist of plants collected around Mabura Hill, Guyana. The Tropenbos 

Foundation, Wageningen and Department of Plant Ecology and Evolutionary Biology, Utrecht University. 

Steyermark, J.A., P.E. Berry, & B.K. Holst. 1995-2005. Flora of the Venezuelan Guayana. Missouri Botanical Garden, St. 

Louis, & Timber Press, Portland, Oregon. 

Furcraea cabuya Trel. 

Furcraea foetida (L.) Haw. 

MONOCOTYLEDONEA 

AGAVACEAE 

P.E. Berry 


Cordyline micrantha Baker (Bromeliaceae) =Brocchinia micrantha 

Furcraea gigantea Vent. =Furcraea foetida 

REFERENCES 

Berry, P.E. 1995. Agavaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 374–375. 

Bogler, D.J. and B.B. Simpson. 1995. A chloroplast DNA study of the Agavaceae. Syst. Bot. 20 (2): 191-205. 

Bos, J.J. 1984. Dracaena in West Africa. 126 pp. Wageningen, The Netherlands: Agricultural University Wageningen. 

Boterenbrood, M.J.A. 1943. Amaryllidaceae. In: Pulle, A., Flora of Suriname 1 (1): 443-454. 

Chahinian, B.J. 1986. The Sansevieria trifasciata varieties. 109 pp. Reseda, California: Trans Terra Publishing. 

Fosberg, F.R. 1985. Cordyline fruticosa (L.) Chevalier (Agavaceae). Baileya 22 (4): 180-181. 

Gentry, H.S. 1982. Agaves of Continental North America. 670 pp. Tucson, Arizona: University of Arizona Press. 

Grenand, P., Moretti, C. and H. Jacquemin. 1987. Pharmacopées Traditionnelles en Guyane: Créoles, Palikur, Wayapi. 569 pp. 

BO 

GU SU FG


Paris: ORSTOM. 

Howard, R.A. 1979. Agavaceae. Flora of the Lesser Antilles 3: 485-504. 

Morgenstern, K.D. 1979. Sansevierias. 112 pp. Kempten, Germany: Illertaler Offsetdruck. 

Ostendorf, F.W. 1962. Nuttige Planten en Sierplanten in Suriname. 325 pp. Landbouwproefstation in Suriname Bulletin No. 79. 

SECAB. 1989. Agave americana. Especies Vegetales Promisorias 1: 9-37. 

ALISMATACEAE 

R.R. Haynes, L.B. Holm-Nielsen 

Echinodorus berteroi (Spreng.) Fassett 

Echinodorus bolivianus (Rusby) Holm-Niels. 

Echinodorus grisebachii Small 

Echinodorus horizontalis Rataj 

Echinodorus macrophyllus (Kunth) Micheli 

Echinodorus macrophyllus subsp. macrophyllus 

Echinodorus macrophyllus subsp. scaber (Rataj) Haynes & Holm-Niels. 

Echinodorus paniculatus Micheli 

Echinodorus reticulatus Haynes & Holm-Niels. 

Echinodorus subalatus (Mart.) Griseb. 

Echinodorus subalatus subsp. andrieuxii (Hook. & Arn.) Haynes & Holm- 

Niels. 

Echinodorus subalatus subsp. subalatus 

Echinodorus tenellus (Mart.) Buchenau 

Sagittaria guayanensis Kunth 

Sagittaria guayanensis subsp. guayanensis 

Sagittaria lancifolia L. 

Sagittaria lancifolia subsp. lancifolia 

Sagittaria rhombifolia Cham. 

Sagittaria sprucei Micheli 


Alisma boliviana Rusby =Echinodorus bolivianus 

Alisma tenellum Mart. =Echinodorus tenellus 

Alisma tenellum f. latifolia Seub. =Echinodorus bolivianus 

Echinodorus andrieuxii (Hook. & Arn.) Small =Echinodorus subalatus subsp. andrieuxii 

Echinodorus gracilis Rataj =Echinodorus grisebachii 

Echinodorus grandiflorus (Cham. & Schltdl.) Micheli [not in flora area] 

Echinodorus guianensis (Kunth) Griseb. =Sagittaria guayanensis 

Echinodorus intermedius Griseb. [=E. grisebachii, fide Lindeman] 

Echinodorus latifolius (Seub.) Rataj =Echinodorus bolivianus 

Echinodorus scaber Rataj =Echinodorus macrophyllus subsp. scaber 

Lophotocarpus guayanensis (Kunth) J.G. Sm. =Sagittaria guayanensis 

Sagittaria pugioniformis L. =Sagittaria lancifolia 

REFERENCES 

Bogin, C. 1955. Revision of the genus Sagittaria. Mem. New York Bot. Gard. 9: 179-233. 

Fassett, N.C. 1955. Echinodorus in the American tropics. Rhodora 57: 133-212. 

Haynes, R.R. & L.B. Holm-Nielsen. 1986. Notes on Echinodorus. Brittonia 38: 325-332. 

Haynes, R.R. & L.B. Holm-Nielsen. 1994. Alismataceae. Flora Neotropica Monograph 64: 1-112. 

Haynes, R.R. & L.B. Holm-Nielsen. 1995. Alismataceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 2: 377-383. 

Jonker, F.P. 1943. Alismataceae. In A. Pulle, Flora of Suriname 1(1): 472-482. 

Jonker, F.P. 1968. Alismataceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 378-379. 

Rataj, K. 1972. Revision of the genus Sagittaria. Part II. Annot. Zool. Bot. Bratislava 78: 1-61. 

Rataj, K. 1975. Revizion (sic) of the genus Echinodorus Rich. Studie CSAV (Praha) 2: 1-156. 

Alocasia macrorrhiza (L.) G. Don [cultivated, escaped] 

Anaphyllopsis americana (Engl.) A. Hay 

Anaphyllopsis pinnata A. Hay 

Anthurium acrobates Sodiro 

Anthurium apaporanum R.E. Schult. 

Anthurium atropurpureum R.E. Schult. & Maguire 

ARACEAE 

G.S. Bunting, T. Croat, D. Nicolson 

GU 

BO GU 

BO GU SU FG 

DA 

GU SU FG? 

GU 

GU SU FG? 

BO GU SU? 

SU 

GU SU 

GU 

GU SU 

AM GU 

BO GU SU FG 

BO GU SU FG 

GU SU FG 

GU SU FG 

BO GU SU FG? 

DA 

AM 

AM 

AM 

AM? 

SU 

SU FG

60 


Anthurium bakeri Hook. f. 

Anthurium bonplandii G.S. Bunting 

Anthurium bonplandii subsp. bonplandii 

Anthurium bonplandii subsp. guayanum (G.S. Bunting) Croat 

Anthurium bonplandii subsp. rionegrense G.S. Bunting 

Anthurium cataniapoense Croat 

Anthurium clavigerum Poepp. 

Anthurium corocoroense G.S. Bunting 

Anthurium cowanii Croat 

Anthurium cremersii Croat, sp. nov. ined. 

Anthurium digitatum (Jacq.) G. Don 

Anthurium eminens Schott 

Anthurium expansum Gleason 

Anthurium friedrichsthalii Schott 

Anthurium glaucophyllum Gleason 

Anthurium gracile (Rudge) Schott 

Anthurium guaiquinimae G.S. Bunting 

Anthurium guanchezii G.S. Bunting 

Anthurium guayanum G.S. Bunting 

Anthurium hookeri Kunth 

Anthurium iramirezae G.S. Bunting 

Anthurium jenmanii Engl. 

Anthurium kunthii Poepp. 

Anthurium lanjouwii A.M.E. Jonker & Jonker 

Anthurium maguirei A.D. Hawkes 

Anthurium martianum K. Koch & Kolb 

Anthurium moonenii Croat & E.G.Gonç. 

Anthurium nymphaeifolium K. Koch & Bouché 

Anthurium obtusum (Engl.) Grayum 

Anthurium pentaphyllum (Aubl.) G. Don 

Anthurium ptarianum Steyerm. 

Anthurium roraimense N.E. Br. 

Anthurium rubrinervium (Link) G. Don 

Anthurium scandens (Aubl.) Engl. 

Anthurium sinuatum Benth. ex Schott 

Anthurium subscriptum G.S. Bunting 

Anthurium tatei G.S. Bunting 

Anthurium thrinax Madison 

Anthurium triphyllum Brongn. ex Schott 

Anthurium vinillense G.S. Bunting 

Anthurium wurdackii G.S. Bunting 

Anthurium xanthoneurum G.S. Bunting 

Anthurium yutajense G.S. Bunting 

Caladium bicolor (Aiton) Vent. 

Caladium humboldtii Schott [cultivated in Guyana] 

Caladium macrotites Schott 

Caladium picturatum K. Koch & Bouché [=Caladium bicolor?] 

Caladium schomburgkii Schott 

Colocasia esculenta (L.) Schott [cultivated, naturalized] 

Cyrtosperma spruceanum (Schott) Engl. 

Dieffenbachia duidae (Steyerm.) G.S. Bunting 

Dieffenbachia elegans A.M.E. Jonker & Jonker 

Dieffenbachia humilis Poepp. 

Dieffenbachia jenmanii Veitch ex Regel 

Dieffenbachia paludicola N.E. Br. ex Gleason 

Dieffenbachia parvifolia Engl. 

Dieffenbachia seguine (Jacq.) Schott 

Dieffenbachia seguine var. lingulata (Mart. ex Schott) Engl. 

Dieffenbachia seguine var. seguine 

Dieffenbachia seguine var. venteniana (Schott) Engl. 

Dieffenbachia seguine var. viridis Engl. 

Dieffenbachia sp. nov. 1 

Dieffenbachia sp. nov. 2 

Dracontium asperum K. Koch [expected in GU] 

AM GU 


AM BO GU FG 

GU SU FG 

AM 

AM SU 


AM 

GU SU 

FG 

BO DA FG 

AM GU SU FG 


GU? 

GU 


BO 

AM 

BO 

BO GU SU FG 

AM 

BO GU SU FG 

AM GU? 

SU FG 

GU SU 

GU SU 

FG 

GU 


GU SU FG 

AM BO GU 

AM BO GU 

GU SU FG 


GU SU FG 

AM 

AM 

GU FG 

AM 

AM 

AM 

AM 

AM 


AM GU 

AM 

AM BO DA GU FG 

BO GU SU FG 


SU 

AM BO GU 

GU SU FG 

GU SU FG 

GU 

AM GU SU FG 

AM SU FG 


SU 

GU SU FG 

SU 

FG 

SU FG 

GU 

BO SU FG


Dracontium dubium Kunth 

Dracontium foecundum Hook. f. 

Dracontium polyphyllum L. 

Heteropsis flexuosa (Kunth) G.S. Bunting 

Heteropsis flexuosa var. flexuosa 

Heteropsis flexuosa var. maguirei G.S. Bunting 

Heteropsis longispathacea Engl. 

Heteropsis melinonii (Engl.) A.M.E. Jonker & Jonker 

Heteropsis oblongata (Kunth) Croat, comb. nov. ined. 

Heteropsis spruceana Schott 

Heteropsis spruceana var. robusta G.S. Bunting 

Heteropsis spruceana var. spruceana 

Heteropsis steyermarkii G.S. Bunting 

Heteropsis tenuispadix G.S. Bunting 

Homalomena picturata (Linden & André) Regel [cultivated only?] 

Jasarum steyermarkii G.S. Bunting 

Monstera adansonii Schott 

Monstera adansonii var. klotzschiana (Schott) Madison 

Monstera adansonii var. laniata (Schott) Madison 

Monstera dubia (Kunth) Engl. & K. Krause 

Monstera gracilis Engl. 

Monstera lechleriana Schott 

Monstera obliqua Miq. 

Monstera spruceana (Schott) Engl. 

Montrichardia arborescens (L.) Schott 

Montrichardia linifera (Arruda) Schott 

Philodendron acutatum Schott 

Philodendron anaadu G.S. Bunting 

Philodendron asplundii Croat 

Philodendron atabapoense G.S. Bunting 

Philodendron auyantepuiense G.S. Bunting 

Philodendron barrosoanum G.S. Bunting 

Philodendron billietiae Croat 

Philodendron bipennifolium Schott [expected in GU] 

Philodendron brevispathum Schott 

Philodendron brevispathum subsp. holmquistii (G.S. Bunting) G.S. Bunting 

Philodendron callosum K. Krause 

Philodendron callosum subsp. callosum 

Philodendron callosum subsp. ptarianum (Steyerm.) G.S. Bunting 

Philodendron camposportoanum G.M. Barroso 

Philodendron canaimae G.S. Bunting 

Philodendron cataniapoense G.S. Bunting 

Philodendron chimantae G.S. Bunting 

Philodendron conforme G.S. Bunting 

Philodendron cremersii Croat & Grayum 

Philodendron deflexum Poepp. ex Schott 

Philodendron delascioi G.S. Bunting 

Philodendron dioscoreoides Gleason 

Philodendron duckei Croat & Grayum 

Philodendron dunstervilleorum G.S. Bunting 

Philodendron dyscarpium R.E. Schult. 

Philodendron dyscarpium var. dyscarpium 

Philodendron dyscarpium var. ventuarianum G.S. Bunting 

Philodendron ecordatum Schott 

Philodendron englerianum Steyerm. 

Philodendron englerianum subsp. duidae (Steyerm.) G.S. Bunting 

Philodendron englerianum subsp. englerianum 

Philodendron exile G.S. Bunting 

Philodendron fendleri K. Krause 

Philodendron fragrantissimum (Hook.) G. Don 

Philodendron goeldii G.M. Barroso 

Philodendron grandifolium (Jacq.) Schott 

Philodendron guaiquinimae G.S. Bunting 

Philodendron guianense Croat & Grayum 

AM BO GU 

GU SU 

SU FG 


AM BO GU SU 

AM BO 

GU? 

BO GU SU FG 

GU 

AM BO GU FG 

AM BO FG 

AM BO 

AM GU? FG 


GU SU FG 

BO GU 




AM BO GU SU? FG? 

AM 

AM BO 




AM? BO DA? GU SU FG 


AM BO 

FG 

AM 

BO GU 

AM 

GU FG 

AM BO SU FG 


AM BO 


BO GU SU FG 

AM BO GU 

GU SU FG 

BO 

AM 

BO 

AM 

SU? FG 


DA 

GU SU 

GU SU FG 

BO 

AM 

AM 

AM 

GU SU FG 

AM BO GU SU 

AM 

BO 

AM SU FG 

DA 


AM SU FG 

DA GU SU FG 

BO 

GU SU FG

62 


Philodendron guttiferum Kunth 

Philodendron hederaceum (Jacq.) Schott [expected in GU] 

Philodendron holstii G.S. Bunting 

Philodendron houlletianum Engl. 

Philodendron hylaeae G.S. Bunting 

Philodendron inaequilaterum Liebm. 

Philodendron inaequilaterum subsp. anthoblastum G.S. Bunting 

Philodendron insigne Schott 

Philodendron jonkerianum Croat 

Philodendron krauseanum Steyerm. [possibly not distinct from P. platypodum 

Gleason] 

Philodendron lemae G.S. Bunting 

Philodendron liesneri G.S. Bunting 

Philodendron linnaei Kunth 

Philodendron linnaei var. linnaei 

Philodendron linnaei var. rionegrense G.S. Bunting 

Philodendron macropodum K. Krause 

Philodendron maguirei G.S. Bunting 

Philodendron marahuacae G.S. Bunting 

Philodendron maroae G.S. Bunting 

Philodendron mawarinumae G.S. Bunting 

Philodendron melinonii Brongn. ex Regel 

Philodendron multinerve G.S. Bunting 

Philodendron muricatum Willd. ex Schott 

Philodendron nebulense G.S. Bunting 

Philodendron ornatum Schott 

Philodendron panduriforme (Kunth) Kunth 

Philodendron panduriforme var. panduriforme 

Philodendron panduriforme var. reichenbachianum (Schott) Croat 

Philodendron pedatum (Hook.) Kunth 

Philodendron peperomioides G.S. Bunting 

Philodendron peraiense G.S. Bunting 

Philodendron phlebodes G.S. Bunting 

Philodendron phlebodes var. kermesinum G.S. Bunting 

Philodendron phlebodes var. phlebodes 

Philodendron pimichinense G.S. Bunting 

Philodendron placidum Schott 

Philodendron platypodum Gleason 

Philodendron quercifolium Engl. 

Philodendron remifolium R.E. Schult. 

Philodendron remifolium subsp. sabulosum (G.S. Bunting) G.S. Bunting 

Philodendron roraimae K. Krause 

Philodendron roraimae subsp. aracamuniense G.S. Bunting 

Philodendron roraimae subsp. roraimae 

Philodendron rudgeanum Schott 

Philodendron samayense G.S. Bunting 

Philodendron scandens K. Koch & Sello 

Philodendron scitulum G.S. Bunting 

Philodendron simsii Sweet 

Philodendron simulans G.S. Bunting 

Philodendron solimoesense A.C. Sm. 

Philodendron sphalerum Schott 

Philodendron splitgerberi Schott 

Philodendron spruceanum G.S. Bunting 

Philodendron squamiferum Poepp. 

Philodendron steyermarkii G.S. Bunting 

Philodendron subsessile Gleason 

Philodendron surinamense (Miq.) Engl. 

Philodendron tatei K. Krause 

Philodendron tatei subsp. melanochlorum (G.S. Bunting) G.S. Bunting 

Philodendron tatei subsp. tatei 

Philodendron triplum G.S. Bunting 

Philodendron venezuelense G.S. Bunting 

Philodendron venustum G.S. Bunting 

AM FG? 

AM BO SU FG? 

AM 

FG 


AM 

AM 

BO GU SU FG 

SU 


BO 

AM 


AM BO DA 

AM 

AM BO FG 

AM 

AM 

AM 

AM SU FG 


AM 


AM 


AM GU 

AM GU 

AM 


BO 

BO 

AM 

AM 

AM 

AM 

GU SU FG 

GU SU FG 

GU 

AM 

AM 

AM BO GU FG? 

AM 

AM BO 

DA GU SU FG 

BO 


AM 

GU FG 

AM 


GU SU FG 

SU FG 

AM 

SU FG 

AM BO 

GU 


AM BO GU 

AM BO 

AM 

BO 

AM FG 

AM


Philodendron vinaceum G.S. Bunting 

Philodendron werkhoveniae Croat 

Philodendron wittianum Engl. 

Philodendron wurdackii G.S. Bunting 

Philodendron yavitense G.S. Bunting 

Philodendron yutajense G.S. Bunting 

Pistia stratiotes L. 

Rhodospatha bolivarana G.S. Bunting 

Rhodospatha brachypoda G.S. Bunting 

Rhodospatha cardonae G.S. Bunting 

Rhodospatha latifolia Poepp. 

Rhodospatha oblongata Poepp. 

Rhodospatha venosa Gleason 

Schismatoglottis americana A.M.E. Jonker & Jonker 

Schismatoglottis bolivarana G.S. Bunting & Steyerm. 

Schismatoglottis spruceana (Schott) G.S. Bunting 

Spathiphyllum bariense G.S. Bunting 

Spathiphyllum cannifolium (Dryand. ex Sims) Schott 

Spathiphyllum cuspidatum Schott 

Spathiphyllum humboldtii Schott 

Spathiphyllum jejunum G.S. Bunting 

Spathiphyllum maguirei G.S. Bunting [=Spathiphyllum cuspidatum fide Croat] 

Spathiphyllum mawarinumae G.S. Bunting 

Spathiphyllum monachinoi G.S. Bunting 

Spathiphyllum monachinoi var. monachinoi 

Spathiphyllum monachinoi var. perangustum G.S. Bunting 

Spathiphyllum neblinae G.S. Bunting 

Spathiphyllum schomburgkii Schott 

Stenospermation ammiticum G.S. Bunting 

Stenospermation amomifolium (Poepp.) Schott 

Stenospermation maguirei A.M.E. Jonker & Jonker 

Stenospermation multiovulatum (Engl.) N.E. Br. 

Stenospermation nebulense G.S. Bunting 

Stenospermation spruceanum Schott 

Stenospermation ulei K. Krause 

Syngonium hastifolium Engl. 

Syngonium podophyllum Schott 

Syngonium podophyllum var. vellozianum (Schott) Croat 

Typhonium blumei Nicolson & Sivad. [naturalized] 

Urospatha angustiloba Engl. 

Urospatha sagittifolia (Rudge) Schott 

Urospatha sagittifolia var. sagittifolia 

Urospathella wurdackii (G.S. Bunting) G.S. Bunting 

Xanthosoma acutum E.G. Gonç. 

Xanthosoma akkermansii (G.S. Bunting) Croat 

Xanthosoma belophyllum (Willd.) Schott [cultivated?] 

Xanthosoma bolivaranum G.S. Bunting 

Xanthosoma caracu K. Koch & Bouché [cultivated?] 

Xanthosoma caulotuberculatum G.S. Bunting 

Xanthosoma conspurcatum Schott 

Xanthosoma contractum G.S. Bunting 

Xanthosoma cordatum N.E. Br. 

Xanthosoma exiguum G.S. Bunting 

Xanthosoma granvillei Croat & S.A. Thomps. 

Xanthosoma helleborifolium (Jacq.) Schott 

Xanthosoma mafaffoides G.S. Bunting 

Xanthosoma maroae G.S. Bunting 

Xanthosoma orinocense G.S. Bunting 

Xanthosoma sagittifolium (L.) Schott [widely cultivated] 

Xanthosoma striatipes (K. Koch & Bouché) Madison 

Xanthosoma striolatum Mart. ex Schott 

Xanthosoma trilobum G.S. Bunting 

Xanthosoma undipes (K. Koch & C.D. Bouché) K. Koch 

Xanthosoma violaceum Schott [widely cultivated] 

AM 

AM 

AM 

BO 

SU 

GU FG 


BO 

AM GU SU FG? 

AM 

GU FG 


AM GU SU FG 

SU FG 

AM BO 

AM 

AM 

AM BO DA GU 

BO GU SU FG? 

AM GU SU FG 

AM 

GU 

AM 

AM BO 

BO 

AM 

AM 

BO GU 

AM BO GU 

AM 

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AM 

AM GU SU 

AM BO GU 

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SU 

AM 



AM 

FG 

AM 

GU SU FG 

BO 

SU? 

BO 

AM GU SU FG 

BO 

GU FG 

AM 

GU FG 

AM BO GU SU 

AM 

AM 

AM 

DA FG 

AM BO GU FG 

GU? FG 

AM 


FG

64 



Alocasia undipes K. Koch & C.D. Bouché =Xanthosoma undipes 

Anthurium amoenum Kunth [not in flora area] 

Anthurium andersonii Schott [not in flora area] 

Anthurium angelorum G.S. Bunting =Anthurium bakeri 

Anthurium bonplandii subsp. cuatrecasii Croat =Anthurium bonplandii subsp. bonplandii 

Anthurium cordatum G. Don [not in flora area] 

Anthurium crassinervium (Jacq.) Schott [not in flora area] 

Anthurium duidae Steyerm. =Anthurium ptarianum 

Anthurium englerianum G.S. Bunting =Anthurium jenmanii 

Anthurium galeottii (Hort.) K. Koch [not in flora area] 

Anthurium huegelii Schott =Anthurium hookeri 

Anthurium idroboanum R.E. Schult. =Anthurium ptarianum 

Anthurium igarapeum G.S. Bunting =Anthurium acrobates 

Anthurium krauseanum Steyerm. =Anthurium ptarianum 

Anthurium liesneri Croat =Anthurium tatei 

Anthurium martini Schott =Anthurium sinuatum 

Anthurium monsteroides Steyerm. =Anthurium clavigerum 

Anthurium neblinae G.S. Bunting =Anthurium triphyllum 

Anthurium nigrescens Engl. [not in flora area] 

Anthurium obovatum Gleason =Anthurium clavigerum 

Anthurium panduratum Mart. ex Schott =Anthurium clavigerum 

Anthurium pentaphyllum var. digitatum (Jacq.) Madison =Anthurium digitatum 

Anthurium polyrrhizum K. Koch & Augustin =Anthurium rubrinervium 

Anthurium potarense Gleason =Anthurium expansum 

Anthurium salvinii Hemsl. [not in flora area] 

Anthurium scandens var. trinerve (Miq.) Matuda =Anthurium obtusum 

Anthurium scolopendrinum (Ham.) Kunth =Anthurium gracile 

Anthurium solitarium Schott [not in flora area] 

Anthurium stahelii A.D. Hawkes =Philodendron deflexum 

Anthurium steyermarkii G.S. Bunting =Anthurium ptarianum 

Anthurium surinamense Miq. =Philodendron surinamense 

Anthurium trinerve Miq. [nom. illeg.] =Anthurium obtusum 

Anthurium trinervium var. obtusum Engl. =Anthurium obtusum 

Arum arborescens L. =Montrichardia arborescens 

Arum bicolor Aiton =Caladium bicolor 

Arum esculentum L. =Colocasia esculenta 

Arum grandifolium Jacq. =Philodendron grandifolium 

Arum hederaceum Jacq. =Philodendron hederaceum 

Arum helleborifolium Jacq. =Xanthosoma helleborifolium 

Arum liniferum Arruda =Montrichardia linifera 

Arum sagittaefolium L. =Xanthosoma sagittifolium 

Arum sagittaefolium Rodschied =Urospatha sagittifolia 

Arum seguine Jacq. =Dieffenbachia seguine 

Arum xanthorrhizon Jacq. =Xanthosoma sagittifolium 

Caladium aculeatum G. Mey. =Montrichardia arborescens 

Caladium akkermansii G.S. Bunting =Xanthosoma akkermansii 

Caladium angustifolium Engl. =Xanthosoma striatipes 

Caladium angustilobum Engl. =Caladium macrotites 

Caladium argyrites Lem. =Caladium humboldtii 

Caladium aturense G.S. Bunting =Caladium picturatum 

Caladium belleyenei hort. Chantin ex Lem. =Caladium picturatum 

Caladium fragrantissimum Hook. =Philodendron fragrantissimum 

Caladium maculatum Lodd. =Dieffenbachia seguine 

Caladium pedatum Hook. =Philodendron pedatum 

Caladium striatipes (Kunth & Bouché) Schott =Xanthosoma striatipes 

Colocasia antiquorum Schott =Colocasia esculenta 

Cyrtosperma americanum Engl. =Anaphyllopsis americana 

Cyrtosperma wurdackii G.S. Bunting =Urospathella wurdackii 

Dieffenbachia bolivarana G.S. Bunting =Dieffenbachia duidae 

Dieffenbachia liesneri Croat =Dieffenbachia duidae 

Dieffenbachia longipistila Croat =Dieffenbachia paludicola 

Dieffenbachia maculata (Lodd.) G. Don =Dieffenbachia seguine 

Dieffenbachia picta Schott =Dieffenbachia seguine


Dieffenbachia spruceana Schott =Dieffenbachia humilis 

Dracontium bissectum Vell. (Cyclanthaceae) =Thoracocarpus bissectus 

Dracontium changuango G.S. Bunting =Dracontium dubium 

Dracontium laciniatum Vell. =Philodendron pedatum 

Dracontium pentaphyllum Aubl. =Anthurium pentaphyllum 

Dracontium pertusum L. =Monstera adansonii 

Dracontium scandens Aubl. =Anthurium scandens 

Dracontium spruceanum (Schott) G.H. Zhu =Cyrtosperma spruceanum 

Elopium surinamense (Miq.) Schott =Philodendron surinamense 

Heteropsis jenmanii Oliv. =Heteropsis flexuosa 

Heteropsis salicifolia Kunth [not in flora area] 

Maguirea spathicarpoides A.D. Hawkes =Dieffenbachia paludicola 

Monstera acreana K. Krause =Monstera dubia 

Monstera amomifolia Poepp. =Stenospermation amomifolium 

Monstera cannifolia Schott =Philodendron rudgeanum 

Monstera egregia Schott =Monstera lechleriana 

Monstera epipremnoides Engl. [not in flora area] 

Monstera expilata Schott =Monstera obliqua 

Monstera falcifolia Engl. =Monstera obliqua 

Monstera fendleri Engl. =Monstera obliqua 

Monstera henri-pittieri G.S. Bunting =Monstera lechleriana 

Monstera irritans N.W. Simmonds =Monstera dubia 

Monstera klotzschiana Schott =Monstera adansonii var. klotzschiana 

Monstera maxima Engl. & K. Krause =Monstera lechleriana 

Monstera milleriana Schott =Monstera adansonii 

Monstera pertusa (L.) de Vriese =Monstera adansonii 

Monstera sagotiana Engl. =Monstera obliqua 

Monstera steyermarkii G.S. Bunting =Monstera spruceana 

Monstera xanthospatha Madison [not in flora area] 

Montrichardia arborescens var. aculeata (G. Mey.) Engl. =Montrichardia arborescens 

Philodendron amplectens A.C. Sm. =Philodendron muricatum 

Philodendron asperatum (K. Koch) K. Koch =Philodendron ornatum 

Philodendron brevispathum var. wurdackii Croat =Philodendron brevispathum subsp. holmquistii 

Philodendron broadwayi N.E. Br. [not in flora area] =Philodendron venosum (Willd.) Croat 

Philodendron calophyllum Brongn. ex Linden & André =Philodendron insigne 

Philodendron cannifolium (Schott) Engl. =Philodendron rudgeanum 

Philodendron cuspidatum K. Koch & Bouché =Philodendron scandens 

Philodendron cyclops A.D. Hawkes =Philodendron acutatum 

Philodendron decurrens K. Krause =Philodendron linnaei 

Philodendron demerarae Gleason =Philodendron fragrantissimum 

Philodendron deviatum Schott =Philodendron hederaceum 

Philodendron duidae Steyerm. =Philodendron englerianum subsp. duidae 

Philodendron grandipes K. Krause [not in flora area] 

Philodendron guttiferum var. rudgeanum (Schott) A.M.E. Jonker & Jonker =Philodendron rudgeanum 

Philodendron haematinum R.E. Schult. =Philodendron insigne 

Philodendron hederaceum auct. non (Jacq.) Schott =Philodendron scandens 

Philodendron hoffmannii Schott =Philodendron hederaceum 

Philodendron holmquistii G.S. Bunting =Philodendron brevispathum subsp. holmquistii 

Philodendron jacquinii Schott =Philodendron hederaceum 

Philodendron jenmanii K. Krause =Philodendron muricatum 

Philodendron laciniatum (Vell.) Engl. =Philodendron pedatum 

Philodendron latilobum Schott =Philodendron panduriforme 

Philodendron levelii G.S. Bunting =Philodendron barrosoanum 

Philodendron longipetiolatum Engl. =Philodendron sphalerum 

Philodendron megalophyllum Schott =Philodendron deflexum 

Philodendron melanochlorum G.S. Bunting =Philodendron tatei subsp. melanochlorum 

Philodendron micans Klotzsch ex K. Koch =Philodendron scandens 

Philodendron milleri Croat =Philodendron barrosoanum 

Philodendron myrmecophilum Engl. =Philodendron deflexum 

Philodendron nobile W. Bull. =Philodendron linnaei 

Philodendron oxycardium Schott =Philodendron scandens 

Philodendron polypodioides A.M.E. Jonker & Jonker =Philodendron pedatum 

Philodendron prieurianum Schott =Philodendron scandens 

Philodendron ptarianum Steyerm. =Philodendron callosum subsp. ptarianum

66 


Philodendron reichenbachianum Schott =Philodendron panduriforme var. reichenbachianum 

Philodendron riedelianum Schott =Philodendron surinamense 

Philodendron rubens Schott =Philodendron ornatum 

Philodendron sabulosum G.S. Bunting =Philodendron remifolium subsp. sabulosum 

Philodendron scabrum K. Krause =Philodendron muricatum 

Philodendron smaragdinum G.S. Bunting =Philodendron muricatum 

Philodendron striatipes Kunth & Bouché =Xanthosoma striatipes 

Philodendron tessmannii K. Krause =Philodendron guttiferum 

Philodendron tobagense Engl. =Philodendron ornatum 

Philodendron tripartitum (Jacq.) Schott [not in flora area] 

Philodendron wayombense A.M.E. Jonker & Jonker =Philodendron bipennifolium 

Philonotion spruceanum Schott =Schismatoglottis spruceana 

Philonotion williamsii Steyerm. =Schismatoglottis spruceana 

Pothos cannaefolia Dryand. =Spathiphyllum cannifolium 

Pothos cannifolius Rudge =Philodendron rudgeanum 

Pothos digitata Jacq. =Anthurium digitatum 

Pothos flexuosus Kunth =Heteropsis flexuosa 

Pothos gracilis Rudge =Anthurium gracile 

Pothos panduraeformis Kunth =Philodendron panduriforme 

Pothos sagittaefolia Rudge =Urospatha sagittifolia 

Pothos scolopendrinus Ham. =Anthurium gracile 

Rhodospatha blanda var. melinonii Engl. =Heteropsis melinonii 

Rhodospatha melinonii (Engl.) Engl. & K. Krause =Heteropsis melinonii 

Rhodospatha pachysperma Madison =Rhodospatha venosa 

Rhodospatha spruceana (Schott) J.F. Macbr. =Stenospermation spruceanum 

Spathicarpa duidae Steyerm. =Dieffenbachia duidae 

Spathiphyllum candolleanum Schott =Spathiphyllum humboldtii 

Spathiphyllum caudatum Poepp. =Urospatha sagittifolia 

Spathiphyllum floribundum (Linden & André) N.E. Br. [not in flora area] 

Spathiphyllum glaziovii Engl. =Spathiphyllum humboldtii 

Spathiphyllum huberi Engl. =Spathiphyllum humboldtii 

Spathiphyllum lanceifolium (Jacq.) Schott [not in flora area] 

Spathiphyllum liesneri Croat =Spathiphyllum humboldtii 

Spathiphyllum sipapoanum G.S. Bunting =Spathiphyllum jejunum 

Stenospermation ammiticum subsp. neblinae G.S. Bunting =Stenospermation ammiticum 

Stenospermation spruceanum var. multiovulatum Engl. =Stenospermation multiovulatum 

Stenospermation steyermarkii G.S. Bunting =Stenospermation multiovulatum 

Stenospermation verecundum R.E. Schult. =Schismatoglottis spruceana 

Syngonium connatum Gleason =Philodendron hylaeae 

Syngonium ternatum Gleason =Syngonium podophyllum 

Syngonium vellozianum Schott =Syngonium podophyllum var. vellozianum 

Thaumatophyllum spruceanum Schott =Philodendron goeldii 

Tornelia laniata Schott =Monstera adansonii var. laniata 

Tornelia spruceana Schott =Monstera spruceana 

Typhonium trilobatum (L.) Schott [misidentification of T. blumei] 

Urospatha caudata (Poepp.) Schott =Urospatha sagittifolia 

Urospatha decipiens Schott =Urospatha sagittifolia 

Urospatha dubia Schott =Urospatha sagittifolia 

Urospatha hostmannii Schott =Urospatha sagittifolia 

Urospatha sagittifolia var. spruceana (Schott) Engl. =Urospatha sagittifolia 

Urospatha savannarum Steyerm. =Urospatha sagittifolia 

Urospatha spruceana Schott =Urospatha sagittifolia 

Urospatha wurdackii (G.S. Bunting) A. Hay =Urospathella wurdackii 

Xanthosoma hoffmannii var. wendlandii (Schott) Engl. [not in flora area] 

Xanthosoma jacquinii Schott =Xanthosoma undipes p.p.; X. sagittifolium p.p. 

Zantedeschia asperata K. Koch =Philodendron ornatum 

REFERENCES 

Bunting, G.S. 1960. A revision of Spathiphyllum. Mem. New York Bot. Gard. 10(3): 1-54. 

Bunting, G.S. 1995. Araceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 600-679. 

Croat, T.B. 1981. A revision of Syngonium. Ann. Missouri Bot. Gard. 68: 565-651. 

Croat, T.B. 1991. A revision of Anthurium section Pachyneurium. Ann. Missouri Bot. Gard. 78(3): 539-855. 

Croat, T.B. 1995 (1994). Taxonomic status of Neotropical Araceae. Aroideana 17: 33-60. 

Croat, T.B. 1995. Two new species of Araceae for the Guianas. Novon 5(1): 24-29.


Croat, T.B. 1997. Araceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide to 

the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the 


Croat, T.B. & M.H. Grayum. 1994. Philodendron subgenus Pteromischum from Saül, French Guiana. Novon 4(3): 211-219. 

Jonker-Verhoef, A.M.E. & F.P. Jonker. 1953. Araceae. In A. Pulle, Flora of Suriname 1(2): 1-80. 

Jonker-Verhoef, A.M.E. & F.P. Jonker. 1968. Araceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 

1(2): 380-412. 

Madison, M. 1977. A revision of Monstera. Contrib. Gray Herb. 207: 3-100. 

Madison, M. 1981. Notes on Caladium and its allies. Selbyana 5(3-4): 342-377. 

Zhu, G., J.O. Boos, and T.B. Croat. 1998. Identity and typification of Draconitum dubium (Araceae). Novon 8(1): 101-103. 

Acrocomia aculeata (Jacq.) Lodd. ex Mart. 

Asterogyne guianensis Granv. & A.J. Hend. 

Astrocaryum acaule Mart. 

Astrocaryum aculeatum G. Mey. 

Astrocaryum chambira Burret 

Astrocaryum gynacanthum Mart. 

Astrocaryum jauari Mart. 

Astrocaryum murumuru Mart. 

Astrocaryum paramaca Mart. 

Astrocaryum sciophilum (Miq.) Pulle 

Astrocaryum vulgare Mart. 

Attalea butyracea (Mutis ex L. f.) Wess. Boer 

Attalea camopiensis (Glassman) Zona 

Attalea degranvillei (Glassman) Zona 

Attalea guianensis (Glassman) Zona 

Attalea luetzelburgii (Burret) Wess. Boer 

Attalea maripa (Aubl.) Mart. 

Attalea maripensis (Glassman) Zona 

Attalea microcarpa Mart. 

Attalea racemosa Spruce 

Attalea speciosa Mart. 

Bactris acanthocarpa Mart. 

Bactris acanthocarpa var. acanthocarpa 

Bactris acanthocarpa var. intermedia A.J. Hend. 

Bactris acanthocarpoides Barb. Rodr. 

Bactris aubletiana Trail 

Bactris balanophora Spruce 

Bactris bidentula Spruce 

Bactris brongniartii Mart. 

Bactris campestris Poepp. ex Mart. 

Bactris constanciae Barb. Rodr. 

Bactris corossilla H. Karst. 

Bactris cuspidata Mart. 

Bactris elegans Barb. Rodr. 

Bactris gasipaes Kunth [cultivated, escaped] 

Bactris gastoniana Barb. Rodr. 

Bactris guineensis (L.) H.E. Moore 

Bactris hirta Mart. 

Bactris hirta var. hirta 

Bactris hirta var. jenmanii A.J. Hend. 

Bactris hirta var. pulchra (Trail) A.J. Hend. 

Bactris major Jacq. 

Bactris major var. infesta (Mart.) Drude 

Bactris major var. major 

Bactris major var. megalocarpa (Trail) A.J. Hend. 

Bactris maraja Mart. [see also Bactris brongniartii] 

Bactris maraja var. juruensis (Trail) A.J. Hend. 

Bactris x moorei Wess. Boer 

Bactris oligocarpa Barb. Rodr. & Trail ex Barb. Rodr. 

Bactris oligoclada Burret 

ARECACEAE 

M. Balick, J.-J. de Granville, A. Henderson 


FG 

AM BO GU 

AM GU SU 

AM 



AM FG 

GU SU FG 

GU SU FG 

GU SU FG 

AM BO FG 

FG 

FG 

FG 

AM 


FG 

AM GU SU FG 

AM 

GU SU 



GU FG 

GU? SU FG 

SU FG 

AM GU 

AM 


AM GU SU FG 

GU SU FG 

AM BO GU? 

SU FG 

GU SU FG 

AM BO FG 

SU FG 

AM BO 

GU SU FG 

GU SU FG 

GU 

GU 


SU 

GU SU FG 

GU 


FG 

BO 

SU FG 

BO GU

68 


Bactris pliniana Granv. & A.J. Hend. 

Bactris ptariana Steyerm. 

Bactris rhaphidacantha Wess. Boer 

Bactris setulosa H. Karst. 

Bactris simplicifrons Mart. 

Bactris sp. nov. 1 [=Bactris vesiculifera Trail, nomen nudum] 

Bactris sp. nov. 2 

Bactris sp. nov. 3 

Bactris tomentosa Mart. 

Bactris tomentosa var. tomentosa 

Bactris trailiana Barb. Rodr. 

Bactris turbinocarpa Barb. Rodr. 

Cocos nucifera L. [cultivated, naturalized] 

Copernicia tectorum (Kunth) Mart. 

Desmoncus mitis Mart. 

Desmoncus cf. mitis Mart. 

Desmoncus orthacanthos Mart. 

Desmoncus phoenicocarpus Barb. Rodr. 

Desmoncus polyacanthos Mart. 

Desmoncus polyacanthos var. polyacanthos 

Desmoncus cf. pycnacanthos Mart. 

Desmoncus cf. riparius Spruce [Desmoncus riparius = Desmoncus 

polyacanthos var. polyacanthos] 

Dictyocaryum fuscum (H. Karst.) H. Wendl. 

Dictyocaryum ptarianum (Steyerm.) H.E. Moore & Steyerm. 

Elaeis guineensis Jacq. [cultivated, naturalized] 

Elaeis oleifera (Kunth) Cortés 

Euterpe catinga Wallace 

Euterpe catinga var. catinga 

Euterpe catinga var. roraimae (Dammer) A.J. Hend. & Galeano 

Euterpe longebracteata Barb. Rodr. 

Euterpe oleracea Mart. 

Euterpe precatoria Mart. 

Euterpe precatoria var. precatoria 

Geonoma appuniana Spruce 

Geonoma aspidiifolia Spruce 

Geonoma baculifera (Poit.) Kunth 

Geonoma densa Linden & H. Wendl. ex H. Wendl. [Mt. Roraima] 

Geonoma deversa (Poit.) Kunth 

Geonoma interrupta (Ruiz & Pav.) Mart. 

Geonoma interrupta var. euspatha (Burret) A.J. Hend. 

Geonoma interrupta var. interrupta [not in flora area fide A. Henderson] 

Geonoma leptospadix Trail 

Geonoma macrostachys Mart. 

Geonoma macrostachys var. acaulis (Mart.) Skov 

Geonoma macrostachys var. poiteauana (Kunth) A.J. Hend. 

Geonoma maxima (Poit.) Kunth 

Geonoma maxima var. ambigua (Spruce) A.J. Hend. 

Geonoma maxima var. chelidonura (Spruce) A.J. Hend. 

Geonoma maxima var. maxima 

Geonoma oldemanii Granv. 

Geonoma oligoclona Trail 

Geonoma sp. nov. 1 

Geonoma sp. nov. 2 

Geonoma stricta (Poit.) Kunth 

Geonoma stricta var. piscicauda (Dammer) A.J. Hend. 

Geonoma stricta var. stricta 

Geonoma stricta var. trailii (Burrett) A.J. Hend. 

Geonoma umbraculiformis Wess. Boer 

Hyospathe elegans Mart. 

Iriartea deltoidea Ruiz & Pav. 

Iriartella setigera (Mart.) H. Wendl. 

Leopoldinia major Wallace 

Leopoldinia piassaba Wallace 

GU SU FG 

BO GU 

GU SU FG 

BO 


GU 

FG 

FG 

FG 

FG 

SU FG 

SU 


BO 

AM 

FG 


AM GU SU FG 


GU SU FG 

FG 

SU 

GU 

AM BO GU 

GU SU FG 

GU SU FG 

AM BO GU 

AM 

AM BO GU 

AM BO GU 



GU SU FG 

AM BO GU 

AM GU FG 


BO GU? 


BO GU SU FG 

BO GU SU FG 

SU 

BO GU SU FG 


AM BO 




AM 


FG 

AM 

FG 

FG 

AM GU SU FG 

SU FG 

AM GU SU FG 

FG 

GU SU FG 


AM BO 

AM BO GU 

AM 

AM


Leopoldinia pulchra Mart. 

Lepidocaryum tenue Mart. 

Lepidocaryum tenue var. gracile (Mart.) A.J. Hend. 

Manicaria saccifera Gaertn. 

Mauritia carana Wallace 

Mauritia flexuosa L. f. 

Mauritiella aculeata (Kunth) Burret 

Mauritiella armata (Mart.) Burret 

Maximbignya dahlgreniana (Bondar) Glassman [=Attalea dahlgreniana] 

Nypa fruticans Wurmb. [Asian native, invasive along tidal rivers] 

Oenocarpus bacaba Mart. 

Oenocarpus balickii F. Kahn 

Oenocarpus bataua Mart. 

Oenocarpus bataua var. bataua 

Oenocarpus bataua var. oligocarpa (Griseb. & H. Wendl.) A.J. Hend. 

Phoenix reclinata Jacq. [cultivated, naturalized] 

Prestoea tenuiramosa (Dammer) H.E. Moore 

Roystonea oleracea (Jacq.) O.F. Cook [also widely cultivated, naturalized] 

Sabal mauritiiformis (H. Karst.) Griseb. & H. Wendl. 

Socratea exorrhiza (Mart.) H. Wendl. 

Syagrus cocoides Mart. 

Syagrus inajai (Spruce) Becc. 

Syagrus orinocensis (Spruce) Burret 

Syagrus sancona H. Karst. 

Syagrus stratincola Wess. Boer 


Acrocomia lasiospatha Mart. =Acrocomia aculeata 

Acrocomia sclerocarpa Mart. =Acrocomia aculeata 

Acrocomia wallaceana (Drude) Becc. =Acrocomia aculeata 

Asterogyne spicata (H.E. Moore) Wess. Boer [not in flora area] 

Astrocaryum caudescens Barb. Rodr. =Astrocaryum aculeatum 

Astrocaryum farinosum Barb. Rodr. =Astrocaryum sciophilum 

Astrocaryum guianense Splitg. ex Mart. =Astrocaryum vulgare 

Astrocaryum gymnopus Burret =Astrocaryum gynacanthum 

Astrocaryum humile Wallace =Bactris acanthocarpa 

Astrocaryum macrocalyx Burret =Astrocaryum murumuru 

Astrocaryum minus Trail =Astrocaryum gynacanthum 

Astrocaryum munbaca Mart. =Astrocaryum gynacanthum 

Astrocaryum murumuru var. macrocalyx (Burret) A.J. Hend. [not in flora area] 

Astrocaryum plicatum Drude =Astrocaryum sciophilum 

Astrocaryum rodriguesii Trail =Astrocaryum paramaca 

Astrocaryum segregatum Drude =Astrocaryum vulgare 

Astrocaryum tucuma Mart. =Astrocaryum aculeatum 

Astrocaryum tucumoides Drude =Astrocaryum vulgare 

Attalea attaleoides (Barb. Rodr.) Wess. Boer [non Scheelea attaleoides Karst.] =Scheelea sp. 

Attalea cryptanthera Wess. Boer =Attalea maripa 

Attalea dahlgreniana (Bondar) Wess. Boer =Maximbignya dahlgreniana 

Attalea excelsa Mart. ex Spreng. [not in flora area] =Attalea phalerata 

Attalea ferruginea Burret =Attalea racemosa 

Attalea funifera Mart. [not in flora area] 

Attalea humboldtiana Spruce =Attalea butyracea 

Attalea macrocarpa (H. Karst.) Wess. Boer =Attalea butyracea 

Attalea macrolepis (Burret) Wess. Boer =Attalea butyracea 

Attalea macropetala (Burret) Wess. Boer =Attalea maripa 

Attalea monosperma Barb. Rodr. [not in flora area] =Attalea spectabilis 

Attalea polysticha (Burret) Wess. Boer =Attalea microcarpa 

Attalea pycnocarpa Wess. Boer =Attalea butyracea 

Attalea regia (Mart.) Wess. Boer =Attalea maripa 

Attalea sagotii (Trail ex Thurn) Wess. Boer =Attalea microcarpa 

Attalea spectabilis Mart. [not in flora area] 

Bactris armata Barb. Rodr. [not in flora area] =Bactris maraja var. maraja 

Bactris capinensis Huber =Bactris tomentosa var. tomentosa 

Bactris carolensis Spruce =Bactris simplicifrons 

AM 

AM BO GU 

AM BO GU 


AM 


AM GU 


SU 

GU 


AM 


GU? SU? FG 

GU SU FG 

GU SU FG 

AM BO GU 

BO DA GU 

BO 


GU 

SU FG 

AM BO 

BO 

GU SU FG

70 


Bactris chaetochlamys Burret =Bactris maraja 

Bactris chaetospatha Mart. =Bactris maraja 

Bactris concinna Mart. [not in flora area] 

Bactris cruegeriana Griseb. & H. Wendl. ex Griseb. =Bactris major var. major 

Bactris cuspidata var. tenuis (Wallace) Drude =Bactris simplicifrons 

Bactris duidae Steyerm. =Bactris corossilla 

Bactris exscapa (Barb. Rodr.) Barb. Rodr. =Bactris acanthocarpa var. acanthocarpa 

Bactris floccosa Spruce =Bactris cuspidata 

Bactris gaviona (Trail) Trail ex Drude =Bactris major var. infesta 

Bactris geonomoides Drude =Bactris hirta var. hirta 

Bactris gymnospatha Burret =Bactris maraja 

Bactris humilis (Wallace) Burret =Bactris acanthocarpa var. acanthocarpa 

Bactris integrifolia Wallace =Bactris hirta var. hirta 

Bactris kamarupa Steyerm. =Bactris maraja 

Bactris leptocarpa Trail ex Thurn =Bactris campestris 

Bactris longicuspis Burret =Bactris maraja 

Bactris luetzelburgii Burret =Bactris simplicifrons 

Bactris megalocarpa Trail ex Thurn =Bactris major var. megalocarpa 

Bactris mitis Mart. =Bactris cuspidata 

Bactris monticola Barb. Rodr. =Bactris maraja 

Bactris pallidispina Mart. =Bactris brongniartii 

Bactris pectinata Mart. =Bactris hirta var. hirta 

Bactris penicillata Barb. Rodr. =Bactris maraja var. juruensis 

Bactris rivularis Barb. Rodr. =Bactris brongniartii 

Bactris sororpanae Steyerm. =Bactris simplicifrons 

Bactris sphaerocarpa Trail [not in flora area] =Bactris tomentosa var. sphaerocarpa 

Bactris tenuis Wallace =Bactris simplicifrons 

Bactris tomentosa var. sphaerocarpa (Trail) A.J. Hend. [not in flora area] 

Bactris trichospatha Trail =Bactris maraja 

Bactris turbinata Spruce =Bactris hirta 

Bactris ulei Burret =Bactris hirta var. pulchra 

Bactris venezuelensis Steyerm. =Bactris corossilla 

Chamaedorea gracilis Willd. [name of uncertain application] 

Cocos aculeatus Jacq. =Acrocomia aculeata 

Cocos butyracea Mutis ex L. f. =Attalea butyracea 

Cocos guineensis L. =Bactris guineensis 

Cocos orinocensis Spruce =Syagrus orinocensis 

Copernicia cerifera (Arruda) Mart. [cultivated only] =Copernicia prunifera (Miller) Moore 

Corypha tectorum Kunth =Copernicia tectorum 

Dahlgrenia ptariana Steyerm. =Dictyocaryum ptarianum 

Desmoncus apureanus L.H. Bailey =Desmoncus orthacanthos 

Desmoncus duidensis Steyerm. =Desmoncus mitis 

Desmoncus guianensis Trail [apparently a nomen nudum] 

Desmoncus horridus Splitg. ex Mart. =Desmoncus orthacanthos 

Desmoncus kaieteurensis L.H. Bailey =Desmoncus phoenicocarpus 

Desmoncus macroacanthos Mart. [specimens probably = B. phoenicocarpus according to Henderson] =Desmoncus 

polyacanthos var. polyacanthos 

Desmoncus maguirei L.H. Bailey =Desmoncus polyacanthos var. polyacanthos 

Desmoncus multijugus Steyerm. =Desmoncus orthacanthos 

Desmoncus oxyacanthos Mart. =Desmoncus polyacanthos 

Desmoncus palustris Trail =Desmoncus orthacanthos 

Desmoncus parvulus L.H. Bailey =Desmoncus phoenicocarpus 

Desmoncus pumilus Trail =Desmoncus mitis 

Desmoncus riparius Spruce =Desmoncus polyacanthos var. polyacanthos 

Desmoncus velezii L.H. Bailey & H.E. Moore =Desmoncus orthacanthos 

Euterpe aurantiaca H.E. Moore =Euterpe catinga 

Euterpe erubescens H.E. Moore =Euterpe catinga var. roraimae 

Euterpe montis-duida Burret =Euterpe catinga 

Euterpe ptariana Steyerm. =Euterpe catinga var. roraimae 

Euterpe roraimae Dammer =Euterpe catinga var. roraimae 

Euterpe stenophylla Trail =Euterpe precatoria var. precatoria 

Euterpe subruminata Burret =Euterpe precatoria 

Euterpe tenuiramosa Dammer =Prestoea tenuiramosa 

Geonoma acaulis Mart. =Geonoma macrostachys var. acaulis 

Geonoma acaulis (Poit.) Burret =Geonoma macrostachys var. poiteauana


Geonoma ambigua Spruce =Geonoma maxima var. ambigua 

Geonoma arundinacea Mart. [not in flora area] 

Geonoma barthia Engel =Geonoma densa 

Geonoma bartlettii Dammer ex Burret =Geonoma deversa 

Geonoma chaunostachys Burret =Geonoma macrostachys var. poiteauana 

Geonoma chelidoneura Spruce =Geonoma maxima var. chelidonura 

Geonoma elegans Mart. [not in flora area] 

Geonoma euspatha Burret =Geonoma interrupta var. euspatha 

Geonoma fusca Wess. Boer =Geonoma aspidiifolia 

Geonoma juruana Dammer =Geonoma maxima 

Geonoma karuaiana Steyerm. =Geonoma interrupta var. euspatha 

Geonoma macrospatha Spruce =Geonoma baculifera 

Geonoma maguirei L.H. Bailey =Geonoma stricta var. stricta 

Geonoma major Burret =Geonoma deversa 

Geonoma microspatha Spruce =Geonoma deversa 

Geonoma multiflora Mart. =Geonoma maxima var. maxima 

Geonoma negrensis Spruce =Geonoma maxima var. maxima 

Geonoma paniculigera Mart. =Geonoma deversa 

Geonoma pinnatifrons Willd. [not in flora area fide A. Henderson] =Geonoma interrupta var. interrupta 

Geonoma piscicauda Dammer =Geonoma stricta var. piscicauda 

Geonoma platycaula Drude & Trail ex Drude [not in flora area] =Geonoma rubescens H. Wendl. ex Drude 

Geonoma poiteauana Kunth =Geonoma macrostachys var. poiteauana 

Geonoma pycnostachys Mart. =Geonoma stricta var. stricta 

Geonoma robusta Burret =Geonoma maxima var. ambigua 

Geonoma roraimae Dammer =Geonoma appuniana p.p.; Geonoma baculifera p.p. 

Geonoma saramaccana L.H. Bailey =Geonoma leptospadix 

Geonoma schomburgkiana Spruce =Geonoma maxima var. ambigua 

Geonoma spruceana Trail =Geonoma maxima 

Geonoma tamandua Trail [not in flora area] =Geonoma macrostachys var. macrostachys 

Geonoma triglochin Burret [not in flora area] [confused with G. umbraculiformis fide Granville] 

Guilielma speciosa Mart. =Bactris gasipaes 

Gynestum baculiferum Poit. =Geonoma baculifera 

Gynestum deversum Poit. =Geonoma deversa 

Gynestum maximum Poit. =Geonoma maxima 

Hyospathe maculata Steyerm. =Hyospathe elegans 

Iriartea exorrhiza Mart. =Socratea exorrhiza 

Iriartea fusca (H. Karst.) Drude [not in flora area] =Dictyocaryum fuscum 

Iriartea setigera Mart. =Iriartella setigera 

Iriartea ventricosa Mart. =Iriartea deltoidea 

Jessenia bataua (Mart.) Burret =Oenocarpus bataua 

Jessenia bataua subsp. bataua =Oenocarpus bataua var. bataua 

Jessenia bataua subsp. oligocarpa (Griseb. & H. Wendl.) Balick =Oenocarpus bataua var. oligocarpa 

Jessenia oligocarpa Griseb. & H. Wendl. =Oenocarpus bataua var. oligocarpa 

Lepidocaryum casiquiarense (Spruce) Drude =Lepidocaryum tenue 

Lepidocaryum gracile Mart. =Lepidocaryum tenue var. gracile 

Lepidocaryum guainiensis (Spruce) Drude =Lepidocaryum tenue var. gracile 

Manicaria atricha Burret =Manicaria saccifera 

Manicaria plukenetii Griseb. & H. Wendl. =Manicaria saccifera 

Markleya dahlgreniana Bondar =Maximbignya dahlgreniana 

Mauritia aculeata Kunth =Mauritiella aculeata 

Mauritia casiquiarensis Spruce =Lepidocaryum tenue 

Mauritia flexuosa var. venezuelana Steyerm. =Mauritia flexuosa 

Mauritia guainiensis Spruce =Lepidocaryum tenue 

Mauritia martiana Spruce =Attalea maripa 

Mauritia minor Burret =Mauritia flexuosa 

Mauritia subinermis Spruce =Mauritiella armata 

Mauritiella martiana (Spruce) Burret =Mauritiella armata 

Maximiliana elegans H. Karst. =Attalea maripa 

Maximiliana macropetala Burret =Attalea maripa 

Maximiliana maripa (Aubl.) Drude =Attalea maripa 

Maximiliana martiana H. Karst. =Attalea maripa 

Maximiliana regia Mart. =Attalea maripa 

Oenocarpus bacaba var. grandis (Burret) Wess. Boer =Oenocarpus bacaba 

Oenocarpus bacaba var. parvus Wess. Boer =Oenocarpus balickii 

Oenocarpus grandis Burret =Oenocarpus bacaba

72 


Oenocarpus oligocarpa (Griseb. & H. Wendl.) Wess. Boer =Oenocarpus bataua var. oligocarpa 

Orbignya barbosiana Burret =Attalea speciosa 

Orbignya luetzelburgii Burret =Attalea luetzelburgii 

Orbignya microcarpa (Mart.) Burret =Attalea microcarpa 

Orbignya phalerata Mart. =Attalea speciosa 

Orbignya polysticha Burret =Attalea microcarpa 

Orbignya sagotii Trail ex Thurn =Attalea microcarpa 

Orbignya speciosa (Mart.) Barb. Rodr. =Attalea speciosa 

Orbignya spectabilis (Mart.) Burret [not in flora area] =Attalea spectabilis 

Palma maripa Aubl. =Attalea maripa 

Parascheelea luetzelburgii (Burret) Dugand =Attalea luetzelburgii 

Prestoea steyermarkii H.E. Moore =Prestoea tenuiramosa 

Raphia taedigera (Mart.) Mart. [not in flora area] 

Roystonea jenmanii (Wright) Burret [cultivated] =Roystonea regia (Kunth) Cook 

Roystonea venezuelana L.H. Bailey =Roystonea oleracea 

Scheelea butyracea (Mutis ex L. f.) H. Karst. ex H. Wendl. =Attalea butyracea 

Scheelea camopiensis Glassman =Attalea camopiensis 

Scheelea degranvillei Glassman =Attalea degranvillei 

Scheelea guianensis Glassman =Attalea guianensis 

Scheelea humboldtiana (Spruce) Burret =Attalea butyracea 

Scheelea macrocarpa H. Karst. =Attalea butyracea 

Scheelea macrolepis Burret =Attalea butyracea 

Scheelea maripensis Glassman =Attalea maripensis 

Scheelea passargei Burret =Attalea butyracea 

Socratea albolineata Steyerm. =Socratea exorrhiza 

Socratea gracilis Burret =Socratea exorrhiza 

Socratea microchlamys Burret =Socratea exorrhiza 

Trithrinax mauritiaeformis H. Karst. =Sabal mauritiiformis 

Yuyba dakamana L.H. Bailey =Bactris simplicifrons 

Yuyba essequiboensis L.H. Bailey =Bactris simplicifrons 

Yuyba gleasonii L.H. Bailey =Bactris simplicifrons 

Yuyba maguirei L.H. Bailey =Bactris simplicifrons 

Yuyba stahelii L.H. Bailey =Bactris simplicifrons 

REFERENCES 

Balick, M.J. 1986. Systematics and economic botany of the Oenocarpus-Jessenia (Palmae) complex. Advances in Economic 

Botany 3: 1-140. The New York Botanical Garden, New York. 

Dahlgren, B.E. 1936. Index of American Palms. Field Museum of Natural History, Botanical Series, 14: 1-436. 

Dransfield, J., D. Johnson, & H. Synge. 1988. The Palms of the New World, A Conservation Census. IUCN: Kew, Richmond, 

Surrey. 

Glassman, S.F. 1972. A revision of B.E. Dahlgren’s Index of American Palms. Phanerogamarum Monographiae Tomus VI: 1- 

294. 

Glassman, S.F. 1977. Preliminary taxonomic studies in the palm genus Attalea H.B.K. Fieldiana Bot. 38: 31-61. 

Glassman, S.F. 1977. Preliminary studies in the palm genus Orbignya Mart. Phytologia 36: 89-115. 

Glassman, S.F. 1978. Preliminary taxonomic studies in the palm genus Maximiliana Mart. Phytologia 38: 161-172. 

Glassman, S.F. 1987. Revisions of the palm genus Syagrus Mart. and other selected genera in the Cocos alliance. Ill. Biol. 

Monogr. 56: 1-230. 

Granville, J.-J. de. 1975. Un nouveau palmier en Guyane Française: Geonoma oldemanii. Adansonia, sér. 2 14(4): 553-559. 

Granville, J.-J. de. 1986. The Palms of the Guianas. Flora of the Guianas Newsletter 3: 36-39. 

Granville, J.-J. de. 1989. La distribucion de las palmas en Guyana Francesa. Acta Amaz. 19: 115-138. 

Granville, J.-J. de. 1990. Les Palmiers de la Guyane Française. Bois et Forêts des Tropiques 220: 43-54. 

Granville, J.-J. de. 1992. The genus Bactris: taxonomy and distribution. Flora of the Guianas Newsletter 9: 6-11. 

Granville, J.-J. de. 1997. Arecaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Granville, J.-J. de. 2002. Les Palmiers de Guyane française. Acta Botanica Gallica, Bulletin de la Société Botanique de France 

149 (3): 275-228. 

Granville, J.-J. de & A. Henderson. 1988. A new species of Asterogyne (Palmae) from French Guiana. Studies on the Flora of the 

Guianas 28. Brittonia 40(1): 76-80. 

Granville, J.-J. de & A. Henderson. 1994. A new species of Bactris (Palmae) from the Amazon region. Studies on the Flora of the 

Guianas 78. Brittonia 46(2): 147-150. 

Henderson, A. 1990. Arecaceae part I. Introduction and Iriarteinae. Flora Neotropica Monogr. 53: 1-100. 

Henderson, A. 1994. The genus Attalea. Palm Journal 114: 49-53. 

Henderson, A. 1995. The Palms of the Amazon. Oxford University Press. 

Henderson, A. 1997. Arecaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 32-122.


Henderson, A, G. Galeano, & R. Bernal. 1995. Field Guide to the Palms of the Americas. Princeton University Press. 

Kahn, F. & J.-J. de Granville. 1992. Palms in Forest Ecosystems of Amazonia. Ecological Studies 95: 1-226. Springer-Verlag. 

Kahn, F. & B. Millan. 1992. Astrocaryum (Palmae) in Amazonia. A preliminary treatment. Bull. Inst. fr. études andines 21(2): 

459-531. 

Lindeman, J.C. 1984. Palmae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 651-652. 

Wessels Boer, J.G. 1965. Palmae. In J. Lanjouw, Flora of Suriname 5(1): 1-172. 

Wessels Boer, J.G. 1968. The Geonomoid Palms. N.V. Noord-Hollandsche Uitgevers Maatschappij, Amsterdam. 

Wessels Boer, J.G. 1972. Palmae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot. 

Gard. 23: 1-832. 

BROMELIACEAE 

E.J. Gouda, J. Grant, B.K. Holst, H. Luther, M. Spencer 

Aechmea angustifolia Poepp. & Endl. 

Aechmea aquilega (Salisb.) Griseb. 

Aechmea aquilega var. aquilega 

Aechmea beeriana L.B. Sm. & M.A. Spencer [=Streptocalyx poeppigii] 

Aechmea brassicoides Baker 

Aechmea brevicollis L.B. Sm. 

Aechmea bromeliifolia (Rudge) Baker 

Aechmea campanulata L.B. Sm. 

Aechmea castelnavii Baker [expected in FG, fide Gouda] 

Aechmea chantinii (CarriPre) Baker 

Aechmea contracta (Mart. ex Schult. f.) Baker 

Aechmea corymbosa (Mart. ex Schult. f.) Mez 

Aechmea egleriana L.B. Sm. 

Aechmea egleriana var. egleriana 

Aechmea egleriana var. major L.B. Sm. [expected in FG, fide Gouda] 

Aechmea lanjouwii (L.B. Sm.) L.B. Sm. 

Aechmea lingulata (L.) Baker 

Aechmea lingulata var. lingulata 

Aechmea longifolia (Rudge) L.B. Sm. & M.A. Spencer [=Streptocalyx 

longifolius] 

Aechmea melinonii Hook. 

Aechmea mertensii (G. Mey.) Schult. & Schult. f. 

Aechmea nudicaulis (L.) Griseb. 

Aechmea nudicaulis var. cuspidata Baker 

Aechmea nudicaulis var. nudicaulis 

Aechmea pallida L.B. Sm. 

Aechmea penduliflora André 

Aechmea poitaei (Baker) L.B. Sm. & M.A. Spencer [=Streptocalyx poitaei] 

Aechmea politii L.B. Sm. 

Aechmea rubiginosa Mez [expected in FG] 

Aechmea setigera Mart. ex Schult. f. 

Aechmea tillandsioides (Mart. ex Schult. f.) Baker 

Aechmea tocantina Baker 

Ananas ananassoides (Baker) L.B. Sm. 

Ananas comosus (L.) Merr. [cultivated, naturalized] 

Ananas lucidus Mill. 

Ananas nanus (L.B. Sm.) L.B. Sm. [records misident., fide Spencer] 

Ananas parguazensis L.A. Camargo & L.B. Sm. 

Araeococcus flagellifolius Harms 

Araeococcus goeldianus L.B. Sm. 

Araeococcus micranthus Brongn. 

Ayensua uaipanensis (Maguire) L.B. Sm. 

Billbergia brachysiphon L.B. Sm. [expected in FG, fide Gouda] 

Billbergia macrolepis L.B. Sm. 

Billbergia pyramidalis (Sims) Linden 

Billbergia rosea hort. ex Beer 

Billbergia violacea Beer 

Brewcaria brocchinioides (L.B. Sm.) B. Holst 

Brewcaria duidensis L.B. Sm., Steyerm. & H. Rob. 

Brewcaria hechtioides (L.B. Sm.) B. Holst 

Brewcaria hohenbergioides (L.B. Sm.) B. Holst 

AM BO GU FG 



SU FG 

GU 

AM 


GU 

AM BO SU 

AM 

AM GU 

AM 

AM FG 

FG 

AM 

SU 

DA GU SU FG 

DA GU SU FG 

BO GU SU FG 

GU SU FG 



DA GU 


GU 

AM BO GU 

FG 

AM GU FG? 

AM BO 

AM BO SU FG 


AM GU SU FG 

GU SU FG 


AM BO FG 

SU? FG? 


AM BO SU FG 


BO 

AM BO GU 

AM 

AM 

AM 

AM 

FG 

SU? FG 

GU SU FG

74 


Brewcaria marahuacae L.B. Sm., Steyerm. & H. Rob. 

Brewcaria reflexa (L.B. Sm.) B. Holst 

Brocchinia acuminata L.B. Sm. [expected in GU] 

Brocchinia cowanii L.B. Sm. 

Brocchinia delicatula L.B. Sm. [Sierra de la Neblina] 

Brocchinia gilmartiniae G.S. Varad. 

Brocchinia hechtioides Mez 

Brocchinia hitchcockii L.B. Sm. 

Brocchinia maguirei L.B. Sm. 

Brocchinia melanacra L.B. Sm. [expected in GU] 

Brocchinia micrantha (Baker) Mez 

Brocchinia paniculata Schult. f. 

Brocchinia prismatica L.B. Sm. 

Brocchinia reducta Baker 

Brocchinia steyermarkii L.B. Sm. 

Brocchinia tatei L.B. Sm. 

Brocchinia vestita L.B. Sm. 

Brocchinia wurdackiana B. Holst 

Bromelia agavifolia Brongn. ex Houllett 

Bromelia alta L.B. Sm. 

Bromelia chrysantha Jacq. [expected in GU, fide Gouda] 

Bromelia fosteriana L.B. Sm. 

Bromelia goeldiana L.B. Sm. 

Bromelia granvillei L.B. Sm. & Gouda 

Bromelia humilis Jacq. 

Bromelia morreniana (Regel) Mez [expected in FG, fide Gouda] 

Bromelia oleveiriae L.B. Sm. [this species not expected in FG, fide Gouda] 

Bromelia pinguin L. [probably cultivated, fide Grant] 

Bromelia plumieri (E. Morren) L.B. Sm. 

Bromelia rondoniana L.B. Sm. [expected in GU, fide Gouda] 

Bromelia serra Griseb. 

Bromelia tubulosa L.B. Sm. 

Catopsis berteroniana (Schult. & Schult. f.) Mez 

Catopsis sessiliflora (Ruiz & Pav.) Mez 

Connellia augustae (M.R. Schomb.) N.E. Br. [Mt. Roraima] 

Connellia caricifolia L.B. Sm. [Mt. Roraima] 

Connellia nutans L.B. Sm. [expected in GU] 

Connellia quelchii N.E. Br. [Mt. Roraima] 

Connellia varadarajanii L.B. Sm. & Steyerm. 

Disteganthus basi-lateralis Lem. 

Disteganthus lateralis (L.B. Sm.) Gouda 

Guzmania altsonii L.B. Sm. 

Guzmania brasiliensis Ule [possibly expected in GU] 

Guzmania calothyrsa Mez 

Guzmania lingulata (L.) Mez 

Guzmania melinonis Regel 

Guzmania monostachia (L.) Rusby ex Mez 

Guzmania monostachia var. monostachia 

Guzmania nubicola L.B. Sm. [Sierra de la Neblina] 

Guzmania patula Mez & Wercklé 

Guzmania plumieri (Griseb.) Mez 

Guzmania retusa L.B. Sm. 

Guzmania roezlii (E. Morren) Mez 

Guzmania sphaeroidea (André) André ex Mez 

Guzmania squarrosa (Mez & Sodiro) L.B. Sm. & Pittendr. 

Guzmania steyermarkii L.B. Sm. [expected in GU] 

Guzmania terrestris L.B. Sm. & Steyerm. 

Hohenbergia stellata Schult. f. [expected in GU, fide Gouda] 

Lindmania arachnoidea (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. 

Lindmania argentea L.B. Sm. 

Lindmania atrorosea (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. 

Lindmania aurea L.B. Sm., Steyerm. & H. Rob. 

Lindmania brachyphylla L.B. Sm. 

Lindmania candelabriformis B. Holst 

AM 

AM 

AM BO GU? 

AM 

AM 

BO 

AM BO GU 

AM 

AM 

AM BO 

BO GU 

AM 

AM 

BO GU 

BO GU 

AM BO GU 

AM 

AM 

BO 

AM BO GU 

GU SU 

SU 

SU? 

FG 

FG 

FG? 

GU SU 


FG 

AM BO FG? 



BO GU 

BO GU? 

BO 

BO GU 

BO 

FG 

SU FG 


AM 

GU 


AM GU SU FG 

AM DA GU? 

AM DA 

AM 

AM BO 

GU? SU? FG 

BO GU 

BO DA GU 

AM BO GU 

AM BO GU 

BO 

AM 

BO DA 

AM 

BO 

BO 

BO 

BO GU 

AM


Lindmania cylindrostachya L.B. Sm. 

Lindmania dendritica (L.B. Sm.) L.B. Sm. [Sierra de la Neblina] 

Lindmania gracillima (L.B. Sm.) L.B. Sm. [expected in GU] 

Lindmania guianensis (Beer) Mez 

Lindmania guianensis var. vestita (L.B. Sm.) L.B. Sm. [recognized as distinct 

in FVG, but appropriate level not ventured] 

Lindmania holstii Steyerm. & L.B. Sm. 

Lindmania huberi L.B. Sm., Steyerm. & H. Rob. 

Lindmania imitans L.B. Sm., Steyerm. & H. Rob. 

Lindmania lateralis (L.B. Sm. & Read) L.B. Sm. & H. Rob. [Sierra de la 

Neblina] 

Lindmania longipes (L.B. Sm.) L.B. Sm. 

Lindmania maguirei (L.B. Sm.) L.B. Sm. [Sierra de la Neblina] 

Lindmania marahuacae (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. 

Lindmania minor L.B. Sm. 

Lindmania navioides L.B. Sm. 

Lindmania nubigena (L.B. Sm.) L.B. Sm. [Sierra de la Neblina] 

Lindmania oliva-estevae L.B. Sm. & Steyerm. ex B. Holst 

Lindmania phelpsiae L.B. Sm. 

Lindmania piresii L.B. Sm., Steyerm. & H. Rob. [Serra Araca, Brazil] 

Lindmania riparia L.B. Sm., Steyerm. & H. Rob. 

Lindmania saxicola L.B. Sm., Steyerm. & H. Rob. 

Lindmania serrulata L.B. Sm. 

Lindmania smithiana (Steyerm. & Luteyn) L.B. Sm. 

Lindmania sp. A [Sierra de la Neblina] 

Lindmania sp. B [Sierra de la Neblina] 

Lindmania sp. C 

Lindmania stenophylla L.B. Sm. 

Lindmania steyermarkii L.B. Sm. 

Lindmania subsimplex L.B. Sm. 

Lindmania thyrsoidea L.B. Sm. 

Lindmania tillandsioides L.B. Sm. 

Lindmania wurdackii L.B. Sm. 

Mezobromelia capituligera (Griseb.) J.R. Grant [=Vriesea capituligera] 

[possibly expected in GU] 

Mezobromelia pleiosticha (Griseb.) Utley & H. Luther [=Vriesea pleiosticha] 

Navia abysmophila L.B. Sm. [Sierra de la Neblina] 

Navia affinis L.B. Sm. [Sierra de la Neblina] 

Navia albiflora L.B. Sm., Steyerm. & H. Rob. 

Navia aliciae L.B. Sm., Steyerm. & H. Rob. 

Navia aloifolia L.B. Sm. [Sierra de la Neblina] 

Navia angustifolia (Baker) Mez 

Navia arida L.B. Sm. & Steyerm. 

Navia aurea L.B. Sm. 

Navia barbellata L.B. Sm. 

Navia berryana L.B. Sm., Steyerm. & H. Rob. 

Navia brachyphylla L.B. Sm. 

Navia breweri L.B. Sm. & Steyerm. 

Navia cardonae L.B. Sm. 

Navia caricifolia L.B. Sm. 

Navia carnevalii L.B. Sm. & Steyerm. 

Navia cataractarum Sandwith 

Navia caurensis L.B. Sm. 

Navia colorata L.B. Sm. 

Navia connata L.B. Sm. & Steyerm. 

Navia crassicaulis L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina] 

Navia cretacea L.B. Sm. 

Navia crispa L.B. Sm. 

Navia cucullata L.B. Sm. 

Navia culcitaria L.B. Sm., Steyerm. & H. Rob. 

Navia duidae L.B. Sm. [expected in Guyana] 

Navia duidae var. duidae 

Navia duidae var. glabrior L.B. Sm. 

Navia emergens L.B. Sm., Steyerm. & H. Rob. 

AM BO 

AM 

BO 

AM BO GU 

GU 

BO 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

AM 

AM BO 


AM 

AM 

AM 

AM 

AM 

GU 

BO GU 

AM 

GU 

AM 

AM 

BO 

BO 

AM GU? 

AM 

GU 

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AM 

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AM 

AM 

AM 

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AM GU? 

AM 

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BO

76 


Navia filifera L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina] 

Navia geaster L.B. Sm., Steyerm. & H. Rob. 

Navia glandulifera B. Holst 

Navia glauca L.B. Sm. 

Navia gleasonii L.B. Sm. 

Navia huberiana L.B. Sm., Steyerm. & H. Rob. 

Navia igneosicola L.B. Sm., Steyerm. & H. Rob. 

Navia immersa L.B. Sm. 

Navia incrassata L.B. Sm. & Steyerm. 

Navia intermedia L.B. Sm. & Steyerm. 

Navia involucrata L.B. Sm. 

Navia jauana L.B. Sm., Steyerm. & H. Rob. 

Navia lactea L.B. Sm., Steyerm. & H. Rob. 

Navia lanigera L.B. Sm. 

Navia lasiantha L.B. Sm. & Steyerm. 

Navia latifolia L.B. Sm. 

Navia lepidota L.B. Sm. [Sierra de la Neblina] 

Navia liesneri L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina] 

Navia lindmanioides L.B. Sm. 

Navia linearis L.B. Sm., Steyerm. & H. Rob. 

Navia luzuloides L.B. Sm., Steyerm. & H. Rob. 

Navia maguirei L.B. Sm. 

Navia mima L.B. Sm. 

Navia mosaica B. Holst 

Navia navicularis L.B. Sm. & Steyerm. 

Navia nubicola L.B. Sm. 

Navia ocellata L.B. Sm. 

Navia octopoides L.B. Sm. 

Navia ovoidea L.B. Sm., Steyerm. & H. Rob. 

Navia paruana B. Holst 

Navia parvula L.B. Sm. 

Navia parvula var. expansa L.B. Sm. [Sierra de la Neblina] 

Navia parvula var. parvula [Sierra de la Neblina] 

Navia patria L.B. Sm. & Steyerm. 

Navia pauciflora L.B. Sm. 

Navia phelpsiae L.B. Sm. 

Navia piresii L.B. Sm., Steyerm. & H. Rob. 

Navia polyglomerata L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina] 

Navia pulvinata L.B. Sm. 

Navia pungens L.B. Sm. 

Navia robinsonii L.B. Sm. 

Navia rupestris (Gleason) Sandwith 

Navia sandwithii L.B. Sm. 

Navia saxicola L.B. Sm. 

Navia scirpiflora L.B. Sm., Steyerm. & H. Rob. 

Navia scopulorum L.B. Sm. 

Navia semiserrata L.B. Sm. 

Navia serrulata L.B. Sm. 

Navia sp. A 

Navia sp. B [Sierra de la Neblina] 

Navia sp. C 

Navia splendens L.B. Sm. 

Navia stenodonta L.B. Sm. 

Navia steyermarkii L.B. Sm. 

Navia subpetiolata L.B. Sm. 

Navia tentaculata B. Holst 

Navia terramarae L.B. Sm. & Steyerm. 

Navia trichodonta L.B. Sm. 

Navia umbratilis L.B. Sm. 

Navia viridis L.B. Sm. 

Navia wurdackii L.B. Sm. 

Navia xyridiflora L.B. Sm. 

Neoregelia leviana L.B. Sm. 

Neoregelia myrmecophila (Ule ex G. Karst. & Schenck) L.B. Sm. 

Neoregelia stolonifera L.B. Sm. 

AM 

BO 

BO 

AM 

AM GU 

AM 

AM 

AM 

BO 

BO 

AM 

BO 

AM 

AM 

BO 

AM 

AM 

AM 

AM 

AM 

BO 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

BO 

BO 

BO 

AM 

AM 

AM BO 

BO 

AM 

AM 

AM 

AM 

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GU 

GU 

BO GU 

SU


Pitcairnia agavifolia L.B. Sm. 

Pitcairnia armata Maury 

Pitcairnia brittoniana Mez 

Pitcairnia bulbosa L.B. Sm. 

Pitcairnia cana B. Holst 

Pitcairnia caricifolia Mart. ex Schult. f. [=Pepinia caricifolia] 

Pitcairnia ctenophylla L.B. Sm. [=Pepinia ctenophylla] [expected in GU] 

Pitcairnia echinata Hook. 

Pitcairnia echinata Hook. var. echinata 

Pitcairnia epiphytica L.B. Sm. 

Pitcairnia filispina L.B. Sm. 

Pitcairnia geyskesii L.B. Sm. [=Pepinia geyskesii] 

Pitcairnia graniticola B. Holst 

Pitcairnia heterophylla (Lindl.) Beer [expected in GU] 

Pitcairnia incarnata Baker [=Pepinia incarnata] 

Pitcairnia juncoides L.B. Sm. 

Pitcairnia kunhardtiana L.B. Sm. 

Pitcairnia leopoldii (W. Till & S. Till) B. Holst 

Pitcairnia leprieurii Baker 

Pitcairnia maguirei L.B. Sm. 

Pitcairnia maidifolia (C. Morren) Decne. ex Planch. & Linden 

Pitcairnia nematophora L.B. Sm. & Read 

Pitcairnia nuda Baker [=Pepinia nuda] 

Pitcairnia orchidifolia Mez 

Pitcairnia patentiflora L.B. Sm. [=Pepinia patentiflora] 

Pitcairnia patentiflora var. armata L.B. Sm. 

Pitcairnia patentiflora var. patentiflora 

Pitcairnia phelpsiae (L.B. Sm.) B. Holst & L.B. Sm. 

Pitcairnia pruinosa Kunth 

Pitcairnia pusilla Mez 

Pitcairnia rubiginosa (Brongn. ex E. Morren) Baker [=Pepinia rubiginosa] 

Pitcairnia rubignosa (Baker) Mez 

Pitcairnia rubignosa var. amazonica (Baker) L.B. Sm. 

Pitcairnia sastrei L.B. Sm. & Read 

Pitcairnia sp. A 

Pitcairnia sp. B [Sierra de la Neblina] 

Pitcairnia sp. C 

Pitcairnia sprucei Baker [=Pepinia sprucei] 

Pitcairnia uaupensis Baker 

Puya floccosa (Linden) E. Morren ex Mez 

Puya floccosa var. floccosa 

Puya grafii Rauh 

Racinaea jenmanii (Baker) M.A. Spencer & L.B. Sm. 

Racinaea spiculosa (Griseb.) M.A. Spencer & L.B. Sm. 

Racinaea spiculosa var. micrantha (Baker) M.A. Spencer & L.B. Sm. 

Racinaea spiculosa var. spiculosa 

Racinaea spiculosa var. stenoglossa (L.B. Sm.) M.A. Spencer & L.B. Sm. 

Racinaea tetrantha (Ruiz & Pav.) M.A. Spencer & L.B. Sm. 

Racinaea tetrantha var. aurantiaca (Griseb.) M.A. Spencer & L.B. Sm. 

Racinaea tetrantha var. caribaea (L.B. Sm.) M.A. Spencer & L.B. Sm. 

Steyerbromelia deflexa L.B. Sm. & H. Rob. 

Steyerbromelia diffusa L.B. Sm., Steyerm. & H. Rob. 

Steyerbromelia discolor L.B. Sm. & H. Rob. 

Steyerbromelia plowmanii (L.B. Sm., Steyerm. & H. Rob.) H. Rob. & D.C. 

Taylor [Sierra de la Neblina] 

Steyerbromelia ramosa (L.B. Sm.) B. Holst 

Steyerbromelia thomasii (L.B. Sm., Steyerm. & H. Rob.) B. Holst [Sierra de 

la Neblina] 

Tillandsia adpressiflora Mez 

Tillandsia anceps Lodd. 

Tillandsia balbisiana Schult. f. [possibly expected in GU] 

Tillandsia brachycaulos Schltdl. 

Tillandsia bulbosa Hook. 

Tillandsia clavigera Mez 

Tillandsia compacta Griseb. [possibly expected in GU] 

AM 

AM BO 

AM BO GU 

AM 

AM 


BO 

AM 

AM 

AM 

AM 

GU SU FG 

BO 

AM BO 

FG 

AM 

AM 

AM 

GU FG 

AM 

AM BO GU SU 

AM 

GU SU 

BO 

AM FG 

AM 

AM FG 

AM 

AM BO 

FG 

AM FG 

AM 

AM 

GU SU? FG 

AM 

AM 

AM 

AM GU FG 

AM 

BO 

BO 

AM 

AM BO GU 


AM BO GU 


AM BO GU 

AM BO GU 

BO 

AM BO GU 

AM 

AM 

AM 

AM 

AM BO 

AM 

AM BO SU FG 


BO DA 

BO 


AM 

AM BO

78 


Tillandsia compacta var. compacta 

Tillandsia complanata Benth. 

Tillandsia confinis L.B. Sm. 

Tillandsia elongata Kunth 

Tillandsia elongata var. subimbricata (Baker) L.B. Sm. [expected in GU] 

Tillandsia fasciculata Sw. 

Tillandsia fendleri Griseb. 

Tillandsia fendleri var. fendleri 

Tillandsia fendleri var. reducta (L.B. Sm.) L.B. Sm. 

Tillandsia flexuosa Sw. 

Tillandsia gardneri Lindl. 

Tillandsia geminiflora Brongn. 

Tillandsia juncea (Ruiz & Pav.) Poir. 

Tillandsia kegeliana Mez 

Tillandsia monadelpha (E. Morren) Baker 

Tillandsia paraensis Mez 

Tillandsia polystachia (L.) L. 

Tillandsia pruinosa Sw. 

Tillandsia pyramidata André 

Tillandsia pyramidata var. pyramidata 

Tillandsia recurvata (L.) L. 

Tillandsia schiedeana Steud. 

Tillandsia stricta Sol. ex Ker Gawl. 

Tillandsia stricta var. stricta 

Tillandsia tenuifolia L. 

Tillandsia tenuifolia var. tenuifolia 

Tillandsia turneri Baker 

Tillandsia turneri var. orientalis L.B. Sm. 

Tillandsia usneoides (L.) L. 

Vriesea bibeatricis Morillo 

Vriesea duidae (L.B. Sm.) Gouda 

Vriesea fibrosa L.B. Sm. 

Vriesea glutinosa Lindl. 

Vriesea heliconioides (Kunth) Hook. ex Walp. 

Vriesea incurva (Griseb.) Read [=Tillandsia incurva] 

Vriesea johnstonii (Mez) L.B. Sm. & Pittendr. [=Vriesea hygrometrica?] 

Vriesea jonghei (K. Koch) E. Morren 

Vriesea maguirei L.B. Sm. [Sierra de la Neblina] 

Vriesea melgueiroi I. Ramírez & Carnevali 

Vriesea mitoura L.B. Sm. [Sierra de la Neblina] 

Vriesea platynema Gaudich. 

Vriesea platynema var. platynema 

Vriesea procera (Mart. ex Schult. f.) Wittm. 

Vriesea procera var. procera 

Vriesea rubra (Ruiz & Pav.) Beer 

Vriesea sanguinolenta Cogn. & Marchand 

Vriesea simplex (Vell.) Beer 

Vriesea socialis L.B. Sm. 

Vriesea soderstromii L.B. Sm. 

Vriesea splendens (Brongn.) Lem. 

Vriesea splendens var. formosa Suringar ex Witte 

Vriesea splendens var. splendens 

Vriesea sulcata L.B. Sm. 

Vriesea wurdackii L.B. Sm. 

Werauhia gigantea (Mart. ex Schult. f.) J.R. Grant [non Vriesea gigantea 

Gaudich, 1846] [=Vriesea amazonica] 

Werauhia gladioliflora (H. Wendl.) J.R. Grant [=Vriesea gladioliflora] 

Werauhia hygrometrica (André) J.R. Grant [=Vriesea hygrometrica] 

Werauhia ringens (Griseb.) J.R. Grant [=Vriesea ringens] 

Werauhia viridiflora (Regel) J.R. Grant [=Vriesea viridiflora] 


Aechmea basi-lateralis (Lem.) L.B.Sm. =Disteganthus basi-lateralis 

Aechmea fernandae (E. Morren) Baker =Aechmea rubiginosa 

Aechmea humilis Mez =Aechmea mertensii 

AM BO 

AM BO GU 

AM BO 

AM BO DA 

AM BO DA 


AM BO GU 

AM BO GU 

AM BO GU 


BO DA 

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BO GU SU FG 



BO DA 

AM BO SU 

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BO 

AM BO 

AM 

BO GU SU 

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BO GU SU FG 

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AM BO GU 

AM BO GU 


AM BO 

AM BO GU 

AM 

BO 

AM GU SU FG 

AM BO GU 

GU 

FG 

AM 

AM 

AM 

AM BO DA GU 

AM BO DA 

DA GU SU FG 

DA GU SU 

AM BO DA GU 

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BO GU SU FG 

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DA GU SU FG 

GU SU FG 

BO GU 

GU 

BO GU


Aechmea kienastii E. Morren ex Mez =Aechmea tillandsioides 

Aechmea lateralis L.B. Sm. =Disteganthus lateralis 

Aechmea magdalenae auct. non (André) André ex Baker sensu L.B. Sm. =Aechmea rubiginosa 

Aechmea megalantha Harms =Aechmea melinonii 

Aechmea mucroniflora Hook. =Aechmea mertensii 

Aechmea oligosphaera Baker =Hohenbergia stellata 

Aechmea prancei L.B. Sm. [not in flora area, fide Gouda] 

Aechmea spicata Mart. ex Schult. f. =Aechmea mertensii 

Aechmea sprucei Mez =Aechmea castelnavii 

Aechmea tillandsioides var. kienastii (E. Morren ex Mez) L.B. Sm. =Aechmea tillandsioides 

Aechmea wullschlaegeliana Mez =Aechmea mertensii 

Ananas ananassoides var. nanus L.B. Sm. =Ananas nanus 

Ananas sativus Schult. f. =Ananas comosus 

Anoplophytum guianensis Beer =Lindmania guianensis 

Billbergia chantinii CarriPre =Aechmea chantinii 

Billbergia contracta Mart. ex Schult. f. =Aechmea contracta 

Billbergia corymbosa Mart. ex Schult. f. =Aechmea corymbosa 

Billbergia tillandsioides Mart. ex Schult. f. =Aechmea tillandsioides 

Bonpartea juncea Ruiz & Pav. =Tillandsia juncea 

Brocchinia bernardii L.B. Sm. =Brocchinia melanacra 

Brocchinia cordylinoides Baker [nom. illeg.] =Brocchinia micrantha 

Brocchinia cryptantha L.B. Sm. =Brocchinia hechtioides 

Brocchinia oliva-estevae Steyerm. & L.B. Sm. =Brocchinia tatei 

Brocchinia secunda L.B. Sm. =Brocchinia tatei 

Bromelia aquilega Salisb. =Aechmea aquilega 

Bromelia comosa L. =Ananas comosus 

Bromelia karatas L. =Bromelia plumieri 

Bromelia lingulata L. =Aechmea lingulata 

Bromelia longifolia Rudge =Aechmea longifolia 

Bromelia lutea G. Mey. =Aechmea nudicaulis var. cuspidata 

Bromelia mertensii G. Mey. =Aechmea mertensii 

Bromelia nudicaulis L. =Aechmea nudicaulis 

Caraguata hygrometrica André =Werauhia hygrometrica 

Caraguata lingulata (L.) Lindl. =Guzmania lingulata 

Caraguata sphaeroidea André =Guzmania sphaeroidea 

Caraguata splendens Planch. =Guzmania lingulata 

Catopsis fendleri Baker =Racinaea tetrantha var. caribaea 

Catopsis inconspicua Baker [not in flora area] =Catopsis nitida (Hook.) Griseb. 

Chevaliera fernandae (E. Morren) L.B. Sm. & W.J. Kress =Aechmea rubiginosa 

Chevaliera pallida (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea pallida 

Chevaliera rubiginosa (Mez) L.B. Sm. & W.J. Kress =Aechmea rubiginosa 

Connellia smithiana Steyerm. & Luteyn =Lindmania smithiana 

Cottendorfia arachnoidea L.B. Sm., Steyerm. & H. Rob. =Lindmania arachnoidea 

Cottendorfia argentea (L.B. Sm.) L.B. Sm. =Lindmania argentea 

Cottendorfia atrorosea L.B. Sm., Steyerm. & H. Rob. =Lindmania atrorosea 

Cottendorfia brachyphylla (L.B. Sm.) L.B. Sm. =Lindmania brachyphylla 

Cottendorfia brachyphylla var. angustior L.B. Sm., Steyerm. & H. Rob. =Lindmania brachyphylla 

Cottendorfia cylindrostachya (L.B. Sm.) L.B. Sm. =Lindmania cylindrostachya 

Cottendorfia dendritica L.B. Sm. =Lindmania dendritica 

Cottendorfia ? dyckioides L.B. Sm. =Lindmania guianensis 

Cottendorfia geniculata (L.B. Sm.) L.B. Sm. =Lindmania guianensis 

Cottendorfia ? geniculata var. minor L.B. Sm., Steyerm. & H. Rob. =Lindmania guianensis 

Cottendorfia gracillima L.B. Sm. =Lindmania gracillima 

Cottendorfia guianensis (Beer) Klotzsch ex Baker =Lindmania guianensis 

Cottendorfia ? guianensis var. vestita L.B. Sm. =Lindmania guianensis 

Cottendorfia lateralis L.B. Sm. & Read =Lindmania lateralis 

Cottendorfia longipes L.B. Sm. =Lindmania longipes 

Cottendorfia maguirei L.B. Sm. =Lindmania maguirei 

Cottendorfia marahuacae L.B. Sm., Steyerm. & H. Rob. =Lindmania marahuacae 

Cottendorfia minor (L.B. Sm.) L.B. Sm. =Lindmania minor 

Cottendorfia navioides (L.B. Sm.) L.B. Sm. =Lindmania navioides 

Cottendorfia nubigena L.B. Sm. =Lindmania nubigena 

Cottendorfia paludosa (L.B. Sm.) L.B. Sm. =Lindmania guianensis 

Cottendorfia phelpsiae (L.B. Sm.) L.B. Sm. =Lindmania phelpsiae

80 


Cottendorfia savannensis L.B. Sm. =Lindmania nubigena 

Cottendorfia serrulata (L.B. Sm.) L.B. Sm. =Lindmania serrulata 

Cottendorfia serrulata var. reducta (L.B. Sm.) L.B. Sm. =Lindmania serrulata 

Cottendorfia stenophylla (L.B. Sm.) L.B. Sm. =Lindmania stenophylla 

Cottendorfia steyermarkii (L.B. Sm.) L.B. Sm. =Lindmania steyermarkii 

Cottendorfia subsimplex (L.B. Sm.) L.B. Sm. =Lindmania subsimplex 

Cottendorfia thyrsoidea (L.B. Sm.) L.B. Sm. =Lindmania thyrsoidea 

Cottendorfia tillandsioides (L.B. Sm.) L.B. Sm. =Lindmania tillandsioides 

Cottendorfia wurdackii (L.B. Sm.) L.B. Sm. =Lindmania wurdackii 

Disteganthus bromeliifolius L.B. Sm. [nomen nudum] =Disteganthus lateralis 

Disteganthus calatheoides (L.B. Sm.) L.B. Sm. & Read =Disteganthus basi-lateralis 

Dyckia augustae (M.R. Schomb.) Baker =Connellia augustae 

Encholirium augustae M.R. Schomb. =Connellia augustae 

Gravisia aquilega (Salisb.) Mez =Aechmea aquilega 

Gravisia lanjouwii (L.B. Sm.) L.B. Sm. =Aechmea lanjouwii 

Guzmania angustifolia (Baker) Wittm. [not in flora area] 

Guzmania capituligera (Griseb.) Mez =Mezobromelia capituligera 

Guzmania cryptantha L.B. Sm. =Guzmania squarrosa 

Guzmania erythrolepis Brongn. ex Planch. [probably misident. of G. melinonis, fide E. Gouda] 

Guzmania geniculata L.B. Sm. =Guzmania sphaeroidea 

Guzmania lingulata var. minor (Mez) L.B. Sm. & Pittendr. =Guzmania lingulata 

Guzmania lingulata var. splendens (Planch.) Mez =Guzmania lingulata 

Guzmania minor Mez =Guzmania lingulata 

Guzmania pleiosticha auct. =Guzmania altsonii 

Guzmania pleiosticha (Griseb.) Mez =Mezobromelia pleiosticha 

Guzmania plumieri auct. non (Griseb.) Mez sensu L.B. Sm. in Lasser =Guzmania steyermarkii 

Guzmania splitgerberi Mez =Mezobromelia pleiosticha 

Guzmania venamensis L.B. Sm. =Guzmania sphaeroidea 

Hepetis amazonica (Baker) Mez =Pitcairnia rubignosa var. amazonica 

Hepetis brittoniana (Mez) Mez =Pitcairnia brittoniana 

Hepetis caricifolia (Mart. ex Schult. f.) Mez =Pitcairnia caricifolia 

Hepetis kegeliana (Schltdl.) Mez =Pitcairnia caricifolia 

Hepetis pauciflora (Baker) Mez =Pitcairnia caricifolia 

Hepetis pruinosa (Kunth) Mez =Pitcairnia pruinosa 

Hepetis rubiginosa (Baker) Mez =Pitcairnia rubiginosa 

Hepetis sprucei (Baker) Mez =Pitcairnia sprucei 

Hepetis uaupensis (Baker) Mez =Pitcairnia uaupensis 

Karatas plumieri E. Morren =Bromelia plumieri 

Lamprococcus brevicollis (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea brevicollis 

Lamprococcus campanulatus (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea campanulata 

Lindmania brachyphylla var. angustior (L.B. Sm., Steyerm. & H. Rob.) R.W. Read =Lindmania brachyphylla 

Lindmania dyckioides (L.B. Sm.) L.B. Sm. =Lindmania guianensis 

Lindmania geniculata L.B. Sm. =Lindmania guianensis 

Lindmania geniculata var. minor (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. =Lindmania guianensis 

Lindmania paludosa L.B. Sm. =Lindmania guianensis 

Lindmania savannensis (L.B. Sm.) L.B. Sm. =Lindmania nubigena 

Lindmania serrulata var. reducta L.B. Sm. =Lindmania serrulata 

Lindmania sessilis L.B. Sm., Steyerm. & H. Rob. =Lindmania serrulata 

Lindmania terramarae L.B. Sm., Steyerm. & H. Rob. =Lindmania marahuacae 

Macrochordion bromeliifolia (Rudge) Beer =Aechmea bromeliifolia 

Navia brocchinioides L.B. Sm. =Brewcaria brocchinioides 

Navia delascionis L.B. Sm., Steyerm. & H. Rob. =Navia berryana 

Navia diffusa L.B. Sm. non Steyerbromelia diffusa L.B. Sm., Steyerm. & H. Rob. 1986 [1987]. =Steyerbromelia plowmanii 

Navia fluviatilis L.B. Sm. =Navia rupestris 

Navia gracilis L.B. Sm. =Brewcaria reflexa 

Navia grafii Steyerm. & B. Holst =Navia phelpsiae 

Navia hechtioides L.B. Sm. =Brewcaria hechtioides 

Navia hohenbergioides L.B. Sm. =Brewcaria hohenbergioides 

Navia iosothrix L.B. Sm. & Steyerm. =Navia nubicola 

Navia lopezii L.B. Sm. (Xyridaceae) =Aratitiyopea lopezii 

Navia pedemontana L.B. Sm., Steyerm. & H. Rob. =Navia nubicola 

Navia ? platyphylla L.B. Sm. & Steyerm. =Steyerbromelia ramosa 

Navia plowmanii L.B. Sm., Steyerm. & H. Rob. =Steyerbromelia plowmanii 

Navia ramosa L.B. Sm. =Steyerbromelia ramosa 

Navia reflexa L.B. Sm. =Brewcaria reflexa


Navia thomasii L.B. Sm., Steyerm. & H. Rob. =Steyerbromelia thomasii 

Nidularium myrmecophilum Ule ex G. Karst. & Schenck =Neoregelia myrmecophila 

Pepinia agavifolia (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia agavifolia 

Pepinia armata (Maury) G.S. Varad. & Gilmartin =Pitcairnia armata 

Pepinia breweri (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia armata 

Pepinia bulbosa (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia bulbosa 

Pepinia caricifolia (Mart. ex Schult. f.) G.S. Varad. & Gilmartin =Pitcairnia caricifolia 

Pepinia ctenophylla (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia ctenophylla 

Pepinia epiphytica (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia epiphytica 

Pepinia filispina (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia filispina 

Pepinia geyskesii (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia geyskesii 

Pepinia incarnata (Baker) G.S. Varad. & Gilmartin =Pitcairnia incarnata 

Pepinia juncoides (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia juncoides 

Pepinia kunhardtiana (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia kunhardtiana 

Pepinia leopoldii W. Till & S. Till =Pitcairnia leopoldii 

Pepinia maguirei (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia maguirei 

Pepinia nuda (Baker) G.S. Varad. & Gilmartin =Pitcairnia nuda 

Pepinia patentiflora (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia patentiflora 

Pepinia pruinosa (Kunth) G.S. Varad. & Gilmartin =Pitcairnia pruinosa 

Pepinia rubiginosa (Brongn.) G.S. Varad. & Gilmartin =Pitcairnia rubiginosa 

Pepinia sprucei (Baker) G.S. Varad. & Gilmartin =Pitcairnia sprucei 

Pepinia uaupensis (Baker) G.S. Varad. & Gilmartin =Pitcairnia uaupensis 

Pepinia wurdackii (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia bulbosa 

Phytarrhiza monadelpha E. Morren =Tillandsia monadelpha 

Pitcairnia amazonica Baker =Pitcairnia rubignosa var. amazonica 

Pitcairnia breweri L.B. Sm. =Pitcairnia armata 

Pitcairnia cinerea L.B. Sm. =Pitcairnia armata 

Pitcairnia kegeliana Schltdl. =Pitcairnia caricifolia 

Pitcairnia nuda auct. non Baker: sensu L.B. Sm. & R.J. Downs =Pitcairnia patentiflora var. patentiflora 

Pitcairnia pauciflora Baker =Pitcairnia caricifolia 

Pitcairnia viridiflora Regel =Werauhia viridiflora 

Pitcairnia wurdackii L.B. Sm. =Pitcairnia bulbosa 

Platyaechmea chantinii (CarriPre) L.B. Sm. & W.J. Kress =Aechmea chantinii 

Platyaechmea contracta (Mart. ex Schult. f.) L.B. Sm. & W.J. Kress =Aechmea contracta 

Platyaechmea tillandsioides (Mart. ex Schult. f.) L.B. Sm. & W.J. Kress =Aechmea tillandsioides 

Pogospermum berteronianum (Schult. f.) Brongn. =Catopsis berteroniana 

Pothuava nudicaulis (L.) Regel =Aechmea nudicaulis 

Pothuava nudicaulis var. cuspidata (Baker) L.B. Sm. & W.J. Kress =Aechmea nudicaulis var. cuspidata 

Pourretia floccosa Linden =Puya floccosa 

Puya augustae (M.R. Schomb.) Mez =Connellia augustae 

Puya guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Puya floccosa var. floccosa 

Puya guyanensis Baker =Puya floccosa var. floccosa 

Puya heterophylla Lindl. =Pitcairnia heterophylla 

Puya maidifolia E. Morren =Pitcairnia maidifolia 

Puya phelpsiae L.B. Sm. =Pitcairnia phelpsiae 

Puya quelchii (N.E. Br.) L.B. Sm. =Connellia quelchii 

Puya roraimae Mez =Connellia quelchii 

Quesnelia lamarckii Baker (Musaceae)[cultivated only] =Musa coccinea Andrews 

Schlumbergeria roezli E. Morren =Guzmania roezlii 

Steyerbromelia neblinae B. Holst =Steyerbromelia plowmanii 

Streptocalyx angustifolius Mez =Aechmea longifolia 

Streptocalyx longifolius (Rudge) Baker =Aechmea longifolia 

Streptocalyx poeppigii Beer =Aechmea beeriana 

Streptocalyx poitaei Baker =Aechmea poitaei 

Thecophyllum splitgerberi (Mez) Pittendr. =Mezobromelia pleiosticha 

Thecophyllum squarrosum Mez & Sodiro =Guzmania squarrosa 

Tillandsia abysmophila L.B. Sm. & Steyerm. =Tillandsia confinis 

Tillandsia aloifolia Hook. =Tillandsia flexuosa 

Tillandsia amazonica Baker =Werauhia gigantea 

Tillandsia appuniana Baker =Vriesea splendens var. splendens 

Tillandsia archeri L.B. Sm. =Tillandsia turneri 

Tillandsia aurantiaca Griseb. =Racinaea tetrantha var. aurantiaca 

Tillandsia berteroniana Schult. f. =Catopsis berteroniana 

Tillandsia bromeliaefolia Rudge =Aechmea bromeliifolia 

Tillandsia capituligera Griseb. =Mezobromelia capituligera

82 


Tillandsia caracasana Baker =Tillandsia compacta var. compacta 

Tillandsia caribaea L.B. Sm. =Racinaea tetrantha var. caribaea 

Tillandsia chinchucuana Harms =Racinaea spiculosa var. micrantha 

Tillandsia compressa Bertero [not in flora area] 

Tillandsia conspersa Miq. =Tillandsia stricta 

Tillandsia crinita Willd. ex Beer =Tillandsia usneoides 

Tillandsia dactylifera E. Morren ex Baker =Vriesea incurva 

Tillandsia deppeana var. clavigera (Mez) L.B. Sm. =Tillandsia clavigera 

Tillandsia deppeana var. latifolia (Griseb.) L.B. Sm. =Tillandsia fendleri var. fendleri 

Tillandsia deppeana var. reducta (L.B. Sm.) L.B. Sm. =Tillandsia fendleri var. reducta 

Tillandsia duidae L.B. Sm. =Vriesea duidae 

Tillandsia excelsa var. latifolia Griseb. =Tillandsia fendleri var. fendleri 

Tillandsia fendleri (Baker) Mez =Racinaea tetrantha var. caribaea 

Tillandsia ferruginascens Mez [nom. inval.] =Racinaea tetrantha var. caribaea 

Tillandsia flexuosa var. vivipara André =Tillandsia flexuosa 

Tillandsia fulgens (Klotzsch ex Beer) Mez =Racinaea tetrantha var. aurantiaca 

Tillandsia gigantea Mart. ex Schult. f. =Werauhia gigantea 

Tillandsia graminifolia Baker =Tillandsia monadelpha 

Tillandsia gymnophylla Baker =Guzmania monostachia var. monostachia 

Tillandsia heliconioides Kunth =Vriesea heliconioides 

Tillandsia heliconioides auct. non Kunth: sensu Griseb. =Guzmania monostachia var. monostachia 

Tillandsia incurva Griseb. =Vriesea incurva 

Tillandsia jenmanii Baker =Racinaea jenmanii 

Tillandsia lineatifolia Mez =Tillandsia anceps 

Tillandsia lingulata L. =Guzmania lingulata 

Tillandsia micrantha Baker =Racinaea spiculosa var. micrantha 

Tillandsia multifolia Mez =Tillandsia turneri var. orientalis 

Tillandsia pachychlamys Mez =Werauhia gladioliflora 

Tillandsia paniculata ß fendleri (Griseb.) Mez =Tillandsia fendleri 

Tillandsia parkeri Baker =Racinaea spiculosa var. spiculosa 

Tillandsia parviflora auct. non Ruiz & Pav. 1802: sensu Griseb. =Racinaea tetrantha var. caribaea 

Tillandsia patens Willd. ex Schult. f. =Tillandsia flexuosa 

Tillandsia pleiosticha Griseb. =Mezobromelia pleiosticha 

Tillandsia procera Mart. ex Schult. f. =Vriesea procera 

Tillandsia pulchella Hook. =Tillandsia tenuifolia 

Tillandsia pulchella var. surinamensis Mez =Tillandsia tenuifolia 

Tillandsia pumila Griseb. =Tillandsia bulbosa 

Tillandsia punctulata Cham. & Schltdl. [not in flora area] 

Tillandsia rhodocincta Baker =Tillandsia turneri var. orientalis 

Tillandsia rhododactyla Mez =Vriesea rubra 

Tillandsia rigidula Baker =Guzmania roezlii 

Tillandsia rubra Ruiz & Pav. =Vriesea rubra 

Tillandsia rubra var. fendleri (Griseb.) Mez =Tillandsia fendleri 

Tillandsia rubra var. reducta L.B. Sm. =Tillandsia fendleri var. reducta 

Tillandsia sessiliflora Ruiz & Pav. =Catopsis sessiliflora 

Tillandsia simplex Vell. =Vriesea simplex 

Tillandsia spiculosa var. micrantha (Baker) L.B. Sm. =Racinaea spiculosa var. micrantha 

Tillandsia spiculosa var. spiculosa =Racinaea spiculosa var. spiculosa 

Tillandsia spiculosa var. stenoglossa (L.B. Sm.) Gouda =Racinaea spiculosa var. stenoglossa 

Tillandsia splendens Brongn. =Vriesea splendens 

Tillandsia stenoglossa L.B. Sm. =Racinaea spiculosa var. stenoglossa 

Tillandsia subimbricata Baker =Tillandsia elongata var. subimbricata 

Tillandsia surinamensis Miq. ex Mez =Tillandsia tenuifolia 

Tillandsia tenuifolia var. surinamensis (Mez) L.B. Sm. =Tillandsia tenuifolia 

Tillandsia tetrantha var. aurantiaca (Griseb.) L.B. Sm. =Racinaea tetrantha var. aurantiaca 

Tillandsia tetrantha var. caribaea (L.B. Sm.) Gouda =Racinaea tetrantha var. caribaea 

Tillandsia trichoides Kunth =Tillandsia usneoides 

Tillandsia usneoides var. filiformis (André) Mez =Tillandsia usneoides 

Tillandsia vestita Schltdl. & Cham. =Tillandsia schiedeana 

Tillandsia viscidula Britton =Vriesea procera 

Tillandsia xiphostachys Griseb. =Tillandsia anceps 

Tussacia fulgens Klotzsch ex Beer =Racinaea tetrantha var. aurantiaca 

Vriesea amazonica (Baker) Mez =Werauhia gigantea 

Vriesea capituligera (Griseb.) L.B. Sm. & Pittendr. =Mezobromelia capituligera 

Vriesea gigantea (Mart. ex Schult. f.) Mez =Werauhia gigantea


Vriesea gladioliflora (H. Wendl.) Antoine =Werauhia gladioliflora 

Vriesea heliconioides var. polysticha L.B. Sm. =Vriesea heliconioides 

Vriesea heterandra (André) L.B. Sm. [misident. of possibly non-Guianan material] 

Vriesea hygrometrica (André) L.B. Sm. & Pittendr. =Werauhia hygrometrica 

Vriesea pachychlamys Mez =Werauhia gladioliflora 

Vriesea pleiosticha (Griseb.) Gouda =Mezobromelia pleiosticha 

Vriesea ringens (Griseb.) Mez =Werauhia ringens 

Vriesea splitgerberi (Mez) L.B. Sm. & Pittendr. =Mezobromelia pleiosticha 

Vriesea viridiflora (Regel) Wittm. ex Mez =Werauhia viridiflora 

Wittmackia lingulata (L.) Mez =Aechmea lingulata 

Wittmackia odora Mez =Aechmea lingulata 

REFERENCES 

Grant, J.R. 1995. The resurrection of Alcantarea and Werauhia, a new genus. Tropische und subtropische Pflanzenwelt 91: 1-59. 

Gouda, E.J. 1987. Bromeliaceae, subfamily Tillandsioideae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 3: 1-112. 

Gouda, E. 1997. Bromeliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Holst, B.K. 1994. Checklist of Venezuelan Bromeliaceae with notes on species distribution by state and levels of endemism. 

Selbyana 15(1): 132-149. 

Holst, B.K. 1997. Bromeliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 548-676. 

Kramer, K.U. & A.R.A. Görts-van Rijn. 1968. Bromeliaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 413-414. 

Smith, L.B. 1957. Bromeliaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 94-148. 

Smith, L.B. 1967. Bromeliaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 15-68. 

Smith, L.B. 1971. Bromeliaceae. Flora of Venezuela 12(1): 1-361 

Smith, L.B. 1986. Revision of the Guayana Highland Bromeliaceae. Ann. Missouri Bot. Gard. 73(4): 689-721. 

Smith, L.B. & R.J. Downs. 1974. Pitcairnioideae. Flora Neotropica Monogr. 14, part 1: 1-658. 

Smith, L.B. & R.J. Downs. 1977. Tillandsioideae. Flora Neotropica Monogr. 14, part 2: 663-1492. 

Smith, L.B. & R.J. Downs. 1979. Bromelioideae. Flora Neotropica Monogr. 14, part 3: 1493-2142. 

BURMANNIACEAE 

P.J.M. Maas, H. Maas 

Apteria aphylla (Nutt.) Barnhart ex Small 

Burmannia bicolor Mart. 

Burmannia capitata (Walter ex J.F. Gmel.) Mart. 

Burmannia compacta Maas & H. Maas 

Burmannia dasyantha Mart. 

Burmannia flava Mart. 

Burmannia foliosa Gleason 

Burmannia kalbreyeri Oliv. 

Burmannia polygaloides Schltr. 

Burmannia sanariapoana Steyerm. 

Burmannia tenella Benth. 

Campylosiphon purpurascens Benth. 

Cymbocarpa saccata Sandwith 

Dictyostega orobanchoides (Hook.) Miers 

Dictyostega orobanchoides subsp. orobanchoides 

Dictyostega orobanchoides subsp. parviflora (Benth.) Snelders & Maas 

Gymnosiphon brachycephalus Snelders & Maas 

Gymnosiphon breviflorus Gleason 

Gymnosiphon capitatus (Benth.) Urb. 

Gymnosiphon cymosus (Benth.) Benth. & Hook. f. 

Gymnosiphon divaricatus (Benth.) Benth. & Hook. f. 

Gymnosiphon fimbriatus (Benth.) Urb. 

Gymnosiphon guianensis Gleason 

Gymnosiphon minutus Snelders & Maas 

Gymnosiphon recurvatus Snelders & Maas 

Gymnosiphon suaveolens (H. Karst.) Urb. 

Gymnosiphon tenellus (Benth.) Urb. 

Hexapterella gentianoides Urb. 

Hexapterella steyermarkii Maas & H. Maas 

Miersiella umbellata (Miers) Urb. 

Thismia saülensis H. Maas & Maas 




AM 

AM 

BO GU SU FG 

AM BO 

BO 

AM 

AM BO 

AM BO GU 


GU 


AM BO GU SU 


AM BO GU SU 

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AM BO GU 

GU SU 

AM BO GU 

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BO 

AM 

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84 



Apteria aphylla var. hymenanthera (Miq.) Jonker =Apteria aphylla 

Apteria gracillima Sagot [nomen nudum] =Apteria aphylla 

Apteria hymenanthera Miq. =Apteria aphylla 

Apteria orobanchoides Hook. =Dictyostega orobanchoides 

Benitzia suaveolens H. Karst. =Gymnosiphon suaveolens 

Burmannia brachyphylla Willd. ex Schult. & Schult. f. =Burmannia bicolor 

Burmannia brachystachya Miq. =Burmannia bicolor 

Burmannia bracteosa Gleason =Burmannia capitata 

Burmannia carrenoi Steyerm. =Burmannia kalbreyeri 

Burmannia foliosa subsp. marahuacensis Maguire & Steyerm. =Burmannia foliosa 

Dictyostega schomburgkii Miers =Dictyostega orobanchoides subsp. orobanchoides 

Dictyostega schomburgkii var. parviflora Benth. =Dictyostega orobanchoides subsp. parviflora 

Dictyostega umbellata Miers =Miersiella umbellata 

Gymnosiphon altsoni Gleason =Hexapterella gentianoides 

Gymnosiphon arcuatus Urb. =Gymnosiphon divaricatus 

Gymnosiphon cornutus (Benth.) Benth. & Hook. f. =Gymnosiphon divaricatus 

Miersiella aristata Sandwith =Miersiella umbellata 

Ptychomeria cymosa Benth. =Gymnosiphon cymosus 

Ptychomeria divaricata Benth. =Gymnosiphon divaricatus 

Ptychomeria fimbriata Benth. =Gymnosiphon fimbriatus 

Ptychomeria tenella Benth. =Gymnosiphon tenellus 

Vogelia capitata Walter ex J.F. Gmel. =Burmannia capitata 

REFERENCES 

Jonker, F.P. 1938. Burmanniaceae. In A. Pulle, Flora of Suriname 1(1): 176-187. 

Jonker, F.P. 1968. Burmanniaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 321-325. 

Maas, P.J.M., H. Maas-van de Kamer, J. van Benthem, H.C.M. Snelders, & T. Rübsamen. 1986. Burmanniaceae. Flora Neotropica 

Monogr. 42: 1-189. 

Maas, P.J.M. & H. Maas-van de Kamer. 1989. Burmanniaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 6: 1-43. 

Maas, P.J.M. & H. Maas-van de Kamer. 1987. A new Thismia (Burmanniaceae) from French Guiana. Brittonia 39(3): 376-378. 

Maas, P.J.M. & H. Maas-van de Kamer. 1997. Bromeliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 3: 678-688. 

Maas-van de Kamer, H. & P. Maas. 1997. Burmanniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & 

J.D. Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and 

Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 232-237. 

Canna glauca L. 

Canna indica L. [naturalized?] 

Canna angustifolia L. =Canna glauca 

Canna coccinea Mill. =Canna indica 

Canna formosa Bouché =Canna indica 

Canna heliconiifolia Bouché =Canna indica 

Canna lambertii Lindl. =Canna indica 

Canna pallida Roscoe =Canna indica 

Canna surinamensis Bouché =Canna indica 

CANNACEAE 

P.J.M. Maas, B.A. Stein 


BO GU SU FG 


REFERENCES 

Maas, P.J.M. 1979. Cannaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 412-415. 

Maas, P.J.M. 1985. Cannaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 1: 69-73. 

Maas, P. & H. Maas-van de Kamer. 1997. Cannaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. 

Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and 

Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 237. 

Miller, J.S. 1998. Cannaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 130-131.


COMMELINACEAE 

P.E. Berry, R. Faden 

Aneilema brasiliense C.B. Clarke 

Aneilema umbrosum (Vahl) Kunth 

Aneilema umbrosum subsp. ovato-oblongum (P. Beauv.) J.K. Morton 

Buforrestia candolleana C.B. Clarke 

Callisia filiformis (M. Martens & Galeotti) D.R. Hunt 

Callisia repens L. 

Commelina benghalensis L. [naturalized] 

Commelina caroliniana Walter 

Commelina diffusa Burm. f. 

Commelina erecta L. 

Commelina obliqua Vahl 

Commelina platyphylla Klotzsch ex C.B. Clarke 

Commelina rufipes Seub. 

Commelina rufipes var. glabrata (D.R. Hunt) Faden & D.R. Hunt 

Commelina rufipes var. rufipes 

Commelina schomburgkiana Klotzsch 

Dichorisandra diederichsanae Steyerm. 

Dichorisandra hexandra (Aubl.) Standl. 

Dichorisandra sp. nov. [Saül, Central French Guiana] 

Dichorisandra villosula Mart. ex Schult. & Schult. f. 

Floscopa peruviana Hassk. ex C.B. Clarke 

Gibasis geniculata (Jacq.) Rohweder 

Murdannia nudiflora (L.) Brenan 

Murdannia schomburgkiana (Kunth) Brückn. 

Tinantia umbellata (Vahl) Urb. 

Tradescantia zanonia (L.) Sw. 

Tradescantia zebrina hort. ex Bosse 

Tripogandra glandulosa (Seub.) Rohweder 

Tripogandra serrulata (Vahl) Handlos 


Aneilema geniculata (Jacq.) Woodson =Gibasis geniculata 

Aneilema ovato-oblongum P. Beauv. =Aneilema umbrosum subsp. ovato-oblongum 

Athyrocarpus persicariifolius (DC.) Hemsl. =Commelina rufipes var. glabrata 

Athyrocarpus rufipes (Seub.) Standl. =Commelina rufipes var. rufipes 

Campelia zanonia (L.) Kunth =Tradescantia zanonia 

Commelina cayennensis Rich. =Commelina diffusa 

Commelina elegans Kunth =Commelina erecta 

Commelina hexandra Aubl. =Dichorisandra hexandra 

Commelina longicaulis Jacq. =Commelina diffusa 

Commelina nudiflora L. =Murdannia nudiflora 

Commelina nudiflora auct. non L.: sensu Burm. f. =Commelina diffusa 

Commelina persicariaefolia auct. non Delile =Commelina rufipes var. glabrata 

Commelina serrulata Vahl =Tripogandra serrulata 

Commelina umbrosa Vahl =Aneilema umbrosum 

Commelina virginica L. [not in flora area] 

Commelina zanonia L. =Tradescantia zanonia 

Commelinopsis glabrata D.R. Hunt =Commelina rufipes var. glabrata 

Commelinopsis persicariifolia (DC.) Pichon =Commelina rufipes var. glabrata 

Commelinopsis persicariifolium auct. non (Delile) Pichon =Commelina rufipes var. glabrata 

Dichorisandra aubletiana Schult. & Schult. f. =Dichorisandra hexandra 

Dichorisandra incurva Roem. & Schult. [not in flora area] 

Dichorisandra leucophthalmos Hook. [not in flora area] 

Leptorhoeo filiformis (M. Martens & Galeotti) C.B. Clarke =Callisia filiformis 

Phaeosphaerion persicariifolium auct. non (Delile) C.B. Clarke =Commelina rufipes var. glabrata 

Rhoeo discolor (L’Hér.) Hance ex Walp. [cultivated only] =Tradescantia spathacea Sw. 

Tinantia erecta (Jacq.) Schltdl. [not in flora area] 

Tradescantia elongata G. Mey. [not in flora area] 

BO 

AM GU SU FG 

AM GU SU FG 

SU FG 

BO 

BO FG 

FG 

GU 


BO GU SU FG 

FG 

GU 



BO GU SU FG 

GU 

GU 


FG 

AM 

AM SU FG 

BO GU SU FG 

AM BO DA GU SU FG? 

GU 

BO GU 

AM BO GU 

GU 

AM FG 

BO DA GU SU FG

86 


Tradescantia filiformis M. Martens & Galeotti =Callisia filiformis 

Tradescantia geniculata Jacq. =Gibasis geniculata 

Tradescantia glandulosa Seub. =Tripogandra glandulosa 

Tradescantia umbellata Vahl =Tinantia umbellata 

REFERENCES 

Aristeguieta, L. 1965. Notas sobre la familia Commelinaceae en Venezuela. Bol. Acad. Cienc. Físicas, Matemáticas y Naturales 

25(69): 94-142. 

Berry, P.E. 1998. Commelinaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 353-365. 

Gracie, C. 1997. Commelinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Uittien, H. 1938. Commelinaceae. In A. Pulle, Flora of Suriname 1(1): 197-212. 

Costus amazonicus (Loes.) J.F. Macbr. 

Costus arabicus L. 

Costus claviger Benoist 

Costus comosus (Jacq.) Roscoe 

Costus comosus var. bakeri (K. Schum.) Maas 

Costus congestiflorus Rich. ex Gagnep. 

Costus curcumoides Maas 

Costus erythrothyrsus Loes. 

Costus guanaiensis Rusby 

Costus guanaiensis var. macrostrobilus (K. Schum.) Maas 

Costus lanceolatus Petersen 

Costus lanceolatus subsp. pulchriflorus (Ducke) Maas 

Costus lasius Loes. 

Costus lima K. Schum. 

Costus longebracteolatus Maas 

Costus scaber Ruiz & Pav. 

Costus spiralis (Jacq.) Roscoe 

Costus spiralis var. spiralis 

Costus spiralis var. villosus Maas 

Costus villosissimus Jacq. 

Dimerocostus strobilaceus Kuntze 

Dimerocostus strobilaceus subsp. gutierrezii (Kuntze) Maas 

Dimerocostus strobilaceus subsp. strobilaceus 

COSTACEAE 

H. Maas, P.J M. Maas 


Costus bracteatus Gleason =Costus claviger 

Costus ciliatus Miq. =Costus scaber 

Costus congestiflorus var. glabrior K. Schum. =Costus congestiflorus 

Costus cylindricus auct. non Jacq. =Costus scaber 

Costus cylindricus var. ciliatus (Miq.) Petersen =Costus scaber 

Costus discolor Roscoe =Costus arabicus 

Costus guianicus Loes. =Costus claviger 

Costus macrostrobilus K. Schum. =Costus guanaiensis var. macrostrobilus 

Costus niveus G. Mey. =Costus arabicus 

Costus pulchriflorus Ducke =Costus lanceolatus subsp. pulchriflorus 

Costus ramosus Woodson =Costus arabicus 

Costus scaberulus Rich. ex Gagnep. =Costus scaber 

Dimerocostus gutierrezii Kuntze =Dimerocostus strobilaceus subsp. gutierrezii 

GU 


GU SU FG 

GU 

GU 

DA GU SU FG 

FG 

GU SU FG 


AM BO GU 

SU FG 

SU FG 

FG 

GU 

GU 




GU SU FG 

BO GU FG 

AM DA GU SU 

DA GU SU 

AM 

REFERENCES 

Maas, H. & P.J.M. Maas. 1997. Costaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 

424-430. 

Maas, P.J.M. 1972. Costoideae (Zingiberaceae). Flora Neotropica Monogr. 8: 1-140. 

Maas, P.J.M. 1977. Costoideae (additions). Flora Neotropica Monogr. 18: 162-218. 

Maas, P.J.M. 1979. Zingiberaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 416-441. 

Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 1: 29-67.


Maas, P. & H. Maas-van de Kamer. 1997. Costaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. 



CYCLANTHACEAE 

R. Eriksson, G. Harling 

Asplundia brachyphylla Harling 

Asplundia fanshawei (Maguire) Harling 

Asplundia flavovaginata Harling 

Asplundia glandulosa (Gleason) Harling 

Asplundia gleasonii Harling 

Asplundia guianensis Harling 

Asplundia heteranthera Harling 

Asplundia luetzelburgii Harling 

Asplundia maguirei Harling 

Asplundia moritziana (Klotzsch) Harling 

Asplundia neblinae Harling 

Asplundia nilssonii Harling 

Asplundia vaupesiana Harling 

Asplundia venezuelensis Harling 

Asplundia xiphophylla Harling 

Carludovica palmata Ruiz & Pav. 

Cyclanthus bipartitus Poit. 

Dicranopygium angustissimum (Sandwith) Harling 

Dicranopygium bolivarense Harling 

Dicranopygium imeriense Harling 

Dicranopygium nanum (Gleason) Harling 

Dicranopygium omichlophilum R.E. Schult. ex Harling 

Dicranopygium pygmaeum (Gleason) Harling 

Dicranopygium pygmaeum subsp. fimbriatum (Maguire) Harling 

Dicranopygium pygmaeum subsp. pygmaeum 

Dicranopygium robustum Harling 

Dicranopygium yacu-sisa Harling 

Evodianthus funifer (Poit.) Lindm. 

Evodianthus funifer subsp. funifer 

Evodianthus funifer subsp. trailianus (Drude) Harling 

Ludovia lancifolia Brongn. 

Sphaeradenia amazonica Harling 

Sphaeradenia duidae Harling 

Sphaeradenia lemaënsis Harling 

Stelestylis stylaris (Gleason) Harling 

Stelestylis surinamensis Harling 

Thoracocarpus bissectus (Vell.) Harling 


Carludovica acuminata (Ruiz & Pav.) Harling [not in flora area] 

Carludovica angustissima Sandwith =Dicranopygium angustissimum 

Carludovica bracteosa Gleason =Thoracocarpus bissectus 

Carludovica coronata Gleason =Evodianthus funifer subsp. funifer 

Carludovica disticha Neum. =Ludovia lancifolia 

Carludovica fanshawei Maguire =Asplundia fanshawei 

Carludovica fimbriata Maguire =Dicranopygium pygmaeum subsp. fimbriatum 

Carludovica glandulosa Gleason =Asplundia glandulosa 

Carludovica kegeliana Lem. =Thoracocarpus bissectus 

Carludovica latifrons Drude [not in flora area] =Asplundia latifrons (Drude) Harling 

Carludovica moritziana Klotzsch =Asplundia moritziana 

Carludovica nana Gleason =Dicranopygium nanum 

Carludovica pygmaea Gleason =Dicranopygium pygmaeum 

Carludovica sarmentosa Sagot ex Drude =Thoracocarpus bissectus 

Carludovica stylaris Gleason =Stelestylis stylaris 

Cyclanthus bipartitus var. gracilis Drude =Cyclanthus bipartitus 

Cyclanthus cristatus Klotzsch =Cyclanthus bipartitus 

Cyclanthus plumieri Poit. =Cyclanthus bipartitus 

AM BO 

GU SU FG 

GU SU 

GU SU FG 

GU 

GU 

SU FG 

AM 

AM BO GU SU 

BO DA 

AM 

BO 

AM BO 

AM BO 

AM BO 

FG? 


GU 

AM BO 

AM BO 

AM BO GU 

AM 

GU SU FG 

SU FG 

GU SU FG 

AM 

AM BO 



GU SU FG 


AM 

AM BO 

BO 

AM BO GU 

SU FG? 

AM BO DA GU SU FG

88 

Ludovia crenifolia Drude =Ludovia lancifolia 

Ludovia funifera Poit. =Evodianthus funifer 

Sphaeradenia maguirei Harling =Sphaeradenia duidae 

Sphaeradenia rubescens Harling =Sphaeradenia duidae 


REFERENCES 

Harling, G. 1958. Monograph of the Cyclanthaceae. Acta Horti Berg. 18: 1-428. 

Harling, G. 1972. Cyclanthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot. 

Gard. 23: 1-832. 

Harling, G. & R. Eriksson. 1998. Cyclanthaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 

4: 471-486. 

Mori, S.A. 1997. Cyclanthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



CYPERACEAE 

K. Camelbeke, P. Goetghebeur, M.S. González-Elizondo, D.M. Kearns, R. Kral, A.A. Reznicek, 

D.A. Simpson, M.T. Strong, W. W. Thomas, G.C. Tucker 

Abildgaardia ovata (Burm. f.) Kral 

Ascolepis brasiliensis (Kunth) Benth. ex C.B. Clarke 

Becquerelia cymosa Brongn. 

Becquerelia cymosa subsp. cymosa 

Becquerelia cymosa subsp. merkeliana (Nees) T. Koyama 

Becquerelia tuberculata (Böck.) H. Pfeiff. 

Bisboeckelera irrigua (Nees) Kuntze 

Bisboeckelera longifolia (Rudge) Kuntze 

Bisboeckelera microcephala (Böck.) T. Koyama 

Bulbostylis aturensis (Maury) C.B. Clarke 

Bulbostylis barbata (Rottb.) C.B. Clarke 

Bulbostylis capillaris (L.) Kunth ex C.B. Clarke 

Bulbostylis capillaris var. capillaris 

Bulbostylis capillaris var. major Uittien 

Bulbostylis circinata (Nees) C.B. Clarke 

Bulbostylis conifera (Kunth) C.B. Clarke 

Bulbostylis fasciculata Uittien 

Bulbostylis fluviatilis Kral & Davidse 

Bulbostylis hispidula (Vahl) R. Haines 

Bulbostylis junciformis (Kunth) C.B. Clarke 

Bulbostylis juncoides (Vahl) Kük. ex Osten 

Bulbostylis lanata (Kunth) C.B. Clarke [nom. cons] 

Bulbostylis leucostachya (Kunth) C.B. Clarke 

Bulbostylis medusae Camelbeke, Goetgh. & O. Huber, sp. nov. ined. [B. 

sp “A” in FVG] 

Bulbostylis paradoxa (Spreng.) Lindm. 

Bulbostylis schomburgkiana (Steud.) M.T. Strong [type erroneously reported 

from Guyana] 

Bulbostylis stenocarpa Kük. 

Bulbostylis surinamensis Uittien 

Bulbostylis svensoniana Steyerm. 

Bulbostylis tenuifolia (Rudge) J.F. Macbr. 

Bulbostylis truncata (Nees) M.T. Strong [expected in FG] 

Bulbostylis vestita (Kunth) C.B. Clarke 

Bulbostylis vestita var. glabrescens Böck. 

Bulbostylis vestita var. vestita 

Calyptrocarya bicolor (H. Pfeiff.) T. Koyama 

Calyptrocarya glomerulata (Brongn.) Urb. 

Calyptrocarya luzuliformis T. Koyama 

Calyptrocarya monocephala Hochst. ex Steud. 

Calyptrocarya poeppigiana Kunth [records may be referable to another 

species] 

Carex bonplandii Kunth 

Carex jamesonii Boott 

Carex marahuacana Reznicek 

BO GU 

BO GU SU 




AM BO GU SU 

AM GU 

GU SU FG 

GU SU FG 

AM 

SU 


GU SU FG 

SU 

GU SU FG 

BO GU SU FG 

GU SU FG 

AM 



AM BO GU FG 


AM BO 

AM 


AM 

GU SU FG 

GU SU FG? 

BO DA 


AM BO GU SU 

AM BO SU 

SU 

FG 

SU FG 






AM 

BO 

GU?

Carex neblinensis Reznicek 

Carex polystachya Sw. ex Wahlenb. 

Carex porrecta Reznicek & Camelbeke 

Carex roraimensis Steyerm. 

Carex tamana Steyerm. 

Cephalocarpus confertus Gilly 

Cephalocarpus dracaenula Nees 

Cephalocarpus lineariifolius Gilly 

Cephalocarpus obovoideus T. Koyama 

Cephalocarpus rigidus Gilly ex Gleason & Killip 

Cladium costatum Steyerm. [Mt. Roraima] 

Cladium mariscus Pohl 

Cyperus aggregatus (Willd.) Endl. 

Cyperus altsonii Kük. 

Cyperus amabilis Vahl 

Cyperus articulatus L. 

Cyperus articulatus var. articulatus 

Cyperus articulatus var. nodosus (Willd.) Kük. 

Cyperus bipartitus Torr. 

Cyperus camphoratus Liebm. 

Cyperus capillifolius A. Rich. 

Cyperus chlorocephalus (C.B. Clarke) Kük. 

Cyperus comosus Poir. 

Cyperus compressus L. 

Cyperus cornelii-ostenii Kük. 

Cyperus croceus Vahl 

Cyperus cuspidatus Kunth 

Cyperus difformis L. 

Cyperus digitatus Roxb. 

Cyperus distans L. f. 

Cyperus drummondii Torr. & Hook. 

Cyperus elegans L. 

Cyperus eragrostis Lam. 

Cyperus esculentus L. 

Cyperus esculentus var. leptostachyus Böck. 

Cyperus felipponei Kük. 

Cyperus filifolius Willd. ex Kunth 

Cyperus filiformis Sw. 

Cyperus flavescens L. 

Cyperus flavicomus Michx. 

Cyperus fugax Liebm. 

Cyperus gardneri Nees 

Cyperus gayi (C.B. Clarke) Kük. 

Cyperus giganteus Vahl 

Cyperus grandisimplex C.B. Clarke 

Cyperus guianensis (C.B. Clarke) Kük. 

Cyperus haspan L. [subsp. haspan?] 

Cyperus haspan subsp. juncoides (Lam.) Kük. 

Cyperus haspan var. riparius (Nees) Kük. 

Cyperus hayesii (C.B. Clarke) Standl. 

Cyperus hermaphroditus (Jacq.) Standl. 

Cyperus hoppiifolius Uittien 

Cyperus imbricatus Retz. 

Cyperus imbricatus var. elongatus (Böck.) Kük. 

Cyperus imbricatus Retz. var. imbricatus 

Cyperus involucratus Rottb. [escaped from cultivation] 

Cyperus iria L. [introduced] 

Cyperus kappleri Hochst. ex Steud. 

Cyperus laxus Lam. 

Cyperus ligularis L. 

Cyperus luzulae (L.) Rottb. ex Retz. 

Cyperus megapotamicus Kunth 

Cyperus meridionalis Barros 

Cyperus meyenianus Kunth 


AM 

BO 

BO 

GU 

BO 

BO 

GU 

AM 

AM 

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AM BO GU 

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AM BO DA GU? SU FG 


BO 

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AM BO GU 

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GU FG 

GU SU FG 

DA GU SU FG 

BO DA GU SU 


AM BO GU 

GU 

SU FG? 

BO DA GU SU 

AM DA GU SU 

SU 

FG 

GU SU 

AM 

AM 

GU SU 

AM 

AM 

BO DA 

GU 

BO 

BO 

GU 

GU 

AM BO DA 

GU FG 

BO 

BO 

DA GU SU FG 

GU SU 


GU SU FG 

GU FG 

FG 

BO GU FG? 

AM BO SU FG 

AM BO DA GU 

GU 

AM BO DA GU 

GU 

SU 

BO GU SU FG 




SU 

FG 

AM BO GU 

BO DA

90 


Cyperus miliifolius Poepp. & Kunth 

Cyperus mutisii (Kunth) Griseb. 

Cyperus odoratus L. 

Cyperus oxylepis Nees ex Steud. 

Cyperus polystachyos Rottb. 

Cyperus procerus Rottb. 

Cyperus prolixus Kunth 

Cyperus pseudodistans Uittien 

Cyperus pumilus L. 

Cyperus reflexus Vahl 

Cyperus rotundus L. 

Cyperus schomburgkianus Nees 

Cyperus simplex Kunth 

Cyperus sphacelatus Rottb. 

Cyperus subumbellatus Kük. 

Cyperus surinamensis Rottb. [var. surinamensis?] 

Cyperus surinamensis var. lutescens Böck. 

Cyperus swartzii (A. Dietr.) Böck. & Kük. 

Cyperus tenuis Sw. 

Cyperus tenuispica Steud. [sparingly naturalized Asian species] 

Cyperus thyrsiflorus Jungh. 

Cyperus trailii C.B. Clarke 

Cyperus unicolor Böck. 

Cyperus unioloides R. Br. 

Cyperus virens Michx. 

Didymiandrum stellatum (Böck.) Gilly 

Diplacrum africanum (Benth.) C.B. Clarke 

Diplacrum capitatum (Willd.) Böck. 

Diplacrum guianense (Nees) T. Koyama 

Diplasia karatifolia Rich. 

Egleria fluctuans L.T. Eiten 

Eleocharis acutangula (Roxb.) Schult. 

Eleocharis alveolatoides S. González & Reznicek 

Eleocharis amazonica C.B. Clarke 

Eleocharis atrospiculata S. González & Reznicek 

Eleocharis ayacuchensis S. González & Reznicek 

Eleocharis capillacea Kunth 

Eleocharis debilis Kunth [f. anomala S. González & Reznicek, found only in 

Amazonas Territory in the flora area] 

Eleocharis eglerioides S. González & Reznicek 

Eleocharis elegans (Kunth) Roem. & Schult. 

Eleocharis filiculmis Kunth 

Eleocharis flavescens (Poir.) Urb. 

Eleocharis geniculata (L.) Roem. & Schult. 

Eleocharis cf. glauca Böck. 

Eleocharis interstincta (Vahl) Roem. & Schult. 

Eleocharis jelskiana Böck. 

Eleocharis liesneri S. González & Reznicek 

Eleocharis maculosa (Vahl) Roem. & Schult. 

Eleocharis minima Kunth [var. minima?] 

Eleocharis minima var. bicolor (Chapm.) Svenson 

Eleocharis minima var. minima 

Eleocharis cf. minima ‘A’ 

Eleocharis cf. minima ‘B’ 

Eleocharis cf. minima ‘C’ 

Eleocharis mitrata (Griseb.) C.B. Clarke 

Eleocharis montana (Kunth) Roem. & Schult. 

Eleocharis mutata (L.) Roem. & Schult. 

Eleocharis nana Kunth 

Eleocharis nigrescens (Nees) Steud. 

Eleocharis aff. nigrescens ‘A’ 

Eleocharis aff. nigrescens ‘B’ 

Eleocharis pachystyla (C. Wright) C.B. Clarke 

Eleocharis plicarhachis (Griseb.) Svenson [expected in Venezuelan Guayana] 


GU FG 


GU 

DA GU SU FG 

GU 


SU 

FG 

BO 


GU 



SU 


SU FG 

AM 

BO 

BO GU 

BO 

AM GU SU 

BO 

BO SU 

BO SU FG 

AM BO GU 

SU 




AM 

AM BO GU SU 

BO 

AM BO 

AM 

AM 

BO 

BO GU 

AM BO 

DA GU SU FG 


BO GU SU FG 


BO GU 



BO 

BO GU 

AM BO DA GU FG? 

AM BO GU 

BO DA 

AM 

BO 

BO 

BO GU SU FG 

BO SU 

DA GU SU FG 

BO GU SU 

BO GU 

BO 

AM BO 

BO GU SU FG 

GU SU FG


Eleocharis retroflexa (Poir.) Urb. 

Eleocharis sellowiana Kunth 

Eleocharis sp. A 

Eleocharis sp. B [Eleocharis retroflexa complex] 

Eleocharis sp. C 

Eleocharis squamigera Svenson 

Eleocharis squamigera subsp. straminea S. González & Reznicek 

Eleocharis steyermarkii S. González & Reznicek 

Eleocharis subfoliata C.B. Clarke 

Eleocharis venezuelensis S. González & Reznicek 

Everardia angusta N.E. Br. 

Everardia debilis T. Koyama & Maguire 

Everardia diffusa T. Koyama & Maguire 

Everardia disticha T. Koyama & Maguire 

Everardia flexifolia (Gilly) T. Koyama & Maguire 

Everardia lanata T. Koyama & Maguire 

Everardia longifolia Gilly ex Gleason & Killip 

Everardia maguireana T. Koyama [Sierra de la Neblina] 

Everardia montana Ridl. ex Thurn 

Everardia montana subsp. duidae (Gilly) T. Koyama & Maguire 

Everardia montana subsp. glaucifolia (Gilly) T. Koyama & Maguire 

Everardia montana subsp. guaiquinimae (Schnee) T. Koyama & Maguire 

Everardia montana subsp. montana 

Everardia montana subsp. ptariensis (Gilly) T. Koyama & Maguire 

Everardia montana subsp. velutina T. Koyama 

Everardia recurvigluma T. Koyama & Maguire [Mt. Roraima] 

Everardia surinamensis Gilly 

Everardia vareschii Maguire 

Exochogyne amazonica C.B. Clarke 

Exochogyne megalorrhyncha Tutin 

Fimbristylis aestivalis (Retz.) Vahl 

Fimbristylis annua (All.) Roem. & Schult. 

Fimbristylis aspera Böck. 

Fimbristylis autumnalis (L.) Roem. & Schult. 

Fimbristylis complanata (Retz.) Link 

Fimbristylis cymosa R. Br. 

Fimbristylis decipiens Kral 

Fimbristylis dichotoma (L.) Vahl 

Fimbristylis dipsacea (Rottb.) C.B. Clarke 

Fimbristylis ferruginea (L.) Vahl 

Fimbristylis limosa Poepp. & Kunth 

Fimbristylis littoralis Gaudich. 

Fimbristylis quinquangularis (Vahl) Kunth 

Fimbristylis spadicea (L.) Vahl 

Fimbristylis squarrosa Vahl 

Fimbristylis vahlii (Lam.) Link 

Fuirena robusta Kunth 

Fuirena simplex Vahl 

Fuirena umbellata Rottb. 

Hypolytrum amplum Poepp. & Kunth 

Hypolytrum jenmanii C.B. Clarke 

Hypolytrum jenmanii subsp. jenmanii 

Hypolytrum jenmanii subsp. serratifolium (Uittien) T. Koyama 

Hypolytrum laxum Kunth 

Hypolytrum leptocalamum M. Alves & W.W. Thomas 

Hypolytrum longifolium (Rich.) Nees 

Hypolytrum longifolium subsp. longifolium 

Hypolytrum longifolium subsp. nicaraguense (Liebm.) T. Koyama 

Hypolytrum longifolium subsp. rubescens (Huber ex C.B. Clarke) T. Koyama 

Hypolytrum longifolium subsp. sylvaticum (Poepp. & Kunth) T. Koyama 

Hypolytrum nudum C.B. Clarke 

Hypolytrum pallidiceps S.S. Hooper & T. Koyama 

Hypolytrum pulchrum (Rudge) H. Pfeiff. 

Hypolytrum schraderianum Nees 


BO GU FG 

BO 

AM 

BO 

AM BO 

AM BO 

BO 

AM BO GU SU 

AM BO 

AM BO GU 

AM GU 

AM BO 

BO GU 

AM 

BO 

BO 

AM 

AM BO GU SU 

AM BO 

AM BO GU 

AM 

AM BO GU 

BO 

AM 

BO GU? 

AM BO GU SU 

BO 


GU 

AM BO 


GU 

DA GU? SU FG 



FG 


BO 

GU SU FG 

GU 


DA GU SU FG 

DA GU SU FG 

BO 


GU SU FG 

FG 





GU SU FG 

AM BO 

AM BO GU 



DA 

AM DA GU 


BO GU 

GU 


AM BO GU FG

92 


Hypolytrum sphaerostachyum Böck. 

Hypolytrum spongiosum T. Koyama 

Hypolytrum stemonifolium T. Koyama 

Hypolytrum strictum Poepp. & Kunth 

Koyamaea neblinensis W.W. Thomas & Davidse 

Kyllinga brevifolia Rottb. 

Kyllinga nemoralis (J.R. Forst. & G. Forst.) Dandy ex Hutch. 

Kyllinga odorata Vahl 

Kyllinga pumila Michx. 

Kyllinga pungens Link 

Kyllinga vaginata Lam. 

Lagenocarpus celiae T. Koyama & Maguire 

Lagenocarpus eriopodus T. Koyama & Maguire 

Lagenocarpus glomerulatus Gilly 

Lagenocarpus guianensis Lindl. & Nees ex Nees 

Lagenocarpus guianensis subsp. guianensis 

Lagenocarpus guianensis subsp. hypochaetus T. Koyama 

Lagenocarpus pendulus T. Koyama 

Lagenocarpus rigidus (Kunth) Nees 

Lagenocarpus rigidus subsp. rigidus 

Lagenocarpus rigidus subsp. tremulus (Nees) T. Koyama & Maguire 

Lagenocarpus sabanensis Gilly 

Lagenocarpus venezuelensis Davidse 

Lagenocarpus verticillatus (Spreng.) T. Koyama & Maguire 

Lipocarpha humboldtiana Nees 

Lipocarpha mexicana Liebm. 

Lipocarpha micrantha (Vahl) G.C. Tucker 

Lipocarpha salzmanniana Steud. 

Lipocarpha schomburgkii (Friedl.) G.C. Tucker 

Machaerina ayangannensis M.T. Strong 

Mapania assimilis T. Koyama 

Mapania assimilis subsp. guianensis D.A. Simpson 

Mapania aturensis D.A. Simpson 

Mapania cuatrecasasii T. Koyama 

Mapania effusa (C.B. Clarke) T. Koyama 

Mapania imeriensis (R. Gross) T. Koyama 

Mapania insignis Sandwith 

Mapania macrophylla (Böck.) H. Pfeiff. 

Mapania maguireana T. Koyama & Steyerm. 

Mapania micrococca (T. Koyama) D.A. Simpson 

Mapania neblina D.A. Simpson 

Mapania paradoxa J. Raynal 

Mapania pedunculata D.A. Simpson 

Mapania pycnocephala (Benth.) Benth. 

Mapania pycnocephala subsp. fluviatilis (Sandwith) T. Koyama 

Mapania pycnocephala subsp. pycnocephala 

Mapania pycnostachya (Benth.) T. Koyama 

Mapania rionegrensis D.A Simpson 

Mapania steyermarkii T. Koyama 

Mapania surinamensis Uittien 

Mapania sylvatica Aubl. 

Mapania sylvatica subsp. sylvatica 

Mapania tepuiana (Steyerm.) T. Koyama 

Mapania theobromina D.A. Simpson 

Oreobolus goeppingeri Suess. 

Oxycaryum cubense (Poepp. & Kunth) Palla 

Pleurostachys guianensis Uittien 

Pleurostachys puberula Böck. 

Remirea maritima Aubl. 

Rhynchocladium steyermarkii (T. Koyama) T. Koyama 

Rhynchospora agostiniana T. Koyama 

Rhynchospora albescens (Miq.) Kük. 

Rhynchospora albida (Nees) Böck. 

Rhynchospora albomarginata Kük. 

AM 

AM GU 

GU 

AM 

AM 

BO GU SU FG 

SU FG? 



GU SU FG 

GU SU FG 

AM GU 

AM 

AM BO GU 



AM 

AM 


AM BO GU 


AM BO SU FG 

AM 

BO GU SU 

AM BO GU SU 

AM BO 


GU 

AM GU 

GU 

FG 

FG 

AM 

GU 

BO GU FG 

AM GU SU 

GU 

BO GU SU FG 

AM GU 

BO GU 

AM 

FG 

AM 


AM BO GU FG 

AM 

AM 

AM BO 

BO GU 

GU SU 

BO GU SU FG 

BO GU SU FG 

BO GU FG? 

AM 

AM 


SU 

AM BO GU FG 

SU FG 

AM BO GU 

AM 

AM GU SU FG 

AM GU 

BO GU FG

Rhynchospora amazonica Poepp. & Kunth 

Rhynchospora amazonica subsp. amazonica 

Rhynchospora amazonica subsp. guianensis (Kük.) T. Koyama 

Rhynchospora angustipaniculata M.T. Strong 

Rhynchospora arenicola Uittien 

Rhynchospora armerioides J. Presl & C. Presl 

Rhynchospora ayangannensis M.T. Strong 

Rhynchospora bakhuisensis M.T. Strong 

Rhynchospora barbata (Vahl) Kunth 

Rhynchospora biflora Böck. 

Rhynchospora bolivarana Steyerm. 

Rhynchospora brachychaeta C. Wright 

Rhynchospora brasiliensis Böck. 

Rhynchospora brevirostris Griseb. 

Rhynchospora brownii Roem. & Schult. 

Rhynchospora brownii subsp. americana Guaglianone 

Rhynchospora cajennensis Böck. 

Rhynchospora candida (Nees) Böck. 

Rhynchospora capillifolia W.W. Thomas 

Rhynchospora capitata (Kunth) Roem. & Schult. 

Rhynchospora caracasana (Kunth) Böck. 

Rhynchospora cariciformis Nees 

Rhynchospora cephalotes (L.) Vahl 

Rhynchospora chimantensis W.W. Thomas 

Rhynchospora ciliata (G. Mey) Kük. 

Rhynchospora comata (Link) Roem. & Schult. 

Rhynchospora contracta (Nees) J. Raynal 

Rhynchospora cordatachenia M.T. Strong 

Rhynchospora corymbosa (L.) Britton 

Rhynchospora curvula Griseb. 

Rhynchospora dentinux C.B. Clarke 

Rhynchospora divaricata (Ham.) M.T. Strong 

Rhynchospora donselaarii M.T. Strong 

Rhynchospora duckei R. Gross 

Rhynchospora duidae Steyerm. 

Rhynchospora eburnea Kral & W.W. Thomas 

Rhynchospora elegantula Maury 

Rhynchospora emaciata (Nees) Böck. 

Rhynchospora exaltata Kunth 

Rhynchospora exaltata var. microcephala Kük. 

Rhynchospora eximia (Nees) Böck. 

Rhynchospora fallax Uittien 

Rhynchospora fascicularis (Michx.) Vahl 

Rhynchospora fascicularis subsp. fascicularis 

Rhynchospora filiformis Vahl 

Rhynchospora gigantea Link 

Rhynchospora globosa (Kunth) Roem. & Schult. 

Rhynchospora harperi Small 

Rhynchospora hassleri C.B. Clarke 

Rhynchospora hieronymii Böck. 

Rhynchospora hieronymii subsp. hieronymii 

Rhynchospora hirsuta (Vahl) Vahl 

Rhynchospora holoschoenoides (Rich.) Herter 

Rhynchospora imeriensis (Kük.) W.W. Thomas 

Rhynchospora immensa Kük. 

Rhynchospora junciformis (Kunth) Böck. 

Rhynchospora lechleri Steud. ex Böck. 

Rhynchospora locuples C.B. Clarke 

Rhynchospora longibracteata Böck. 

Rhynchospora macrochaeta Steud. ex Böck. 

Rhynchospora maguireana T. Koyama 

Rhynchospora marisculus Nees 

Rhynchospora mexicana (Liebm.) Steud. 

Rhynchospora montana (Uittien) H. Pfeiff. 




AM BO GU FG? 

GU 


AM GU SU 

GU 

SU 


AM BO 

AM BO GU 

GU 

AM BO GU 


BO 

BO 

AM BO GU FG 


BO GU 

AM 

AM BO GU SU 

AM BO 


BO 



BO FG 

FG 


BO GU SU FG 

AM BO FG 

BO GU SU FG 

SU 

AM 

BO 

AM BO 

AM BO 

AM BO GU SU 

AM BO GU 

GU 

BO 

BO GU SU FG 

GU SU 

GU SU 


GU SU FG 


GU SU 

GU SU FG 

AM BO 

AM BO 



AM 

GU 


AM BO 

BO 

AM BO GU 

AM BO 

AM GU 

AM BO GU SU 

AM 

SU

94 


Rhynchospora nervosa (Vahl) Böck. 

Rhynchospora nitens (Vahl) A. Gray 

Rhynchospora pallida (Nees) Steud. 

Rhynchospora papillosa W.W. Thomas 

Rhynchospora pilosa (Kunth) Böck. 

Rhynchospora planifolia (T. Koyama) W.W. Thomas 

Rhynchospora polyphylla Vahl 

Rhynchospora pubera (Vahl) Böck. 

Rhynchospora pubera subsp. parvula W.W. Thomas 

Rhynchospora pubera subsp. pubera 

Rhynchospora pubisquama M.T. Strong 

Rhynchospora radicans (Schltdl. & Cham.) H. Pfeiff. 

Rhynchospora radicans subsp. microcephala (Bertero ex Spreng.) W.W. 

Thomas 

Rhynchospora reptans (Rich.) Böck. 

Rhynchospora rigidifolia (Gilly) T. Koyama 

Rhynchospora riparia (Nees) Böck. 

Rhynchospora robusta (Kunth) Böck. 

Rhynchospora roraimae Kük. [Mt. Roraima] 

Rhynchospora rugosa (Vahl) Gale 

Rhynchospora ruiziana Böck. 

Rhynchospora rupicola M.T. Strong 

Rhynchospora sanariapensis Steyerm. 

Rhynchospora saxisavannicola M.T. Strong 

Rhynchospora schomburgkiana (Böck.) T. Koyama 

Rhynchospora sp. A 

Rhynchospora sp. B 

Rhynchospora sp. C 

Rhynchospora sp. D 

Rhynchospora sp. E 

Rhynchospora splendens Lindm. 

Rhynchospora spruceana C.B. Clarke 

Rhynchospora steyermarkii T. Koyama [expected in GU] 

Rhynchospora subdicephala T. Koyama 

Rhynchospora sublanata Lindeman & Donsel. 

Rhynchospora subplumosa C.B. Clarke 

Rhynchospora tenella (Nees) Böck. 

Rhynchospora tenerrima Nees ex Spreng. 

Rhynchospora tenuis Link 

Rhynchospora tenuis subsp. austrobrasiliensis T. Koyama 

Rhynchospora tenuis subsp. tenuis 

Rhynchospora tepuiana Steyerm. 

Rhynchospora trichochaeta C.B. Clarke 

Rhynchospora trichophora Nees 

Rhynchospora triflora Vahl 

Rhynchospora trispicata (Nees) Schrad. ex Steud. 

Rhynchospora tuerckheimii C.B. Clarke ex Kük. 

Rhynchospora uniflora Böck. 

Rhynchospora unisetosa T. Koyama 

Rhynchospora velutina (Kunth) Böck. 

Rhynchospora watsonii (Britton) Davidse 

Schoenoplectus maritimus (L.) Lye 

Schoenoplectus robustus (Pursh) M.T. Strong 

Scleria acanthocarpa Böck. 

Scleria amazonica Camelbeke, M. Strong & Goetgh. 

Scleria anceps Liebm. 

Scleria bracteata Cav. 

Scleria camaratensis Core 

Scleria cerradicola T. Koyama 

Scleria cyperina Willd. ex Kunth 

Scleria cyperinoides C.B. Clarke 

Scleria distans Poir. 

Scleria eggersiana Böck. 

Scleria flagellum-nigrorum P.J. Bergius 


GU 

AM 

GU 

GU 

AM 

AM BO GU 


AM BO GU SU 


SU 

GU SU FG 

GU SU FG 

GU SU FG 

AM BO 


AM BO 

AM BO GU 


BO GU 

GU SU 

AM BO GU 

SU FG 

AM 

AM 

GU 

AM 

AM 

AM 

BO 

BO 

GU 

GU 

AM BO 

BO 

GU SU FG? 

AM FG 

GU SU 

AM BO GU SU 


BO GU SU FG 

AM BO GU 

GU 

SU FG 

AM BO GU 

AM BO GU 

AM BO GU SU? 

AM BO 

BO GU SU FG 

AM BO 

BO 

BO 

GU 

GU 

SU FG 


GU SU FG 

SU FG? 

DA GU SU? FG? 

FG 

AM 

GU SU 

AM BO 

BO 

GU 

GU 

SU FG 


AM BO 


BO GU SU 

AM BO DA GU SU FG


Scleria hirtella Sw. [records may be referable to Scleria distans Poir.] 

Scleria interrupta Rich. 

Scleria lacustris C. Wright 

Scleria latifolia Sw. 

Scleria lindleyana C.B. Clarke 

Scleria lithosperma (L.) Sw. 

Scleria longigluma Kük. 

Scleria macrogyne C.B. Clarke 

Scleria macrophylla J. Presl & C. Presl 

Scleria martii (Nees) Steud. 

Scleria melaleuca Rchb. ex Schltdl. & Cham. 

Scleria microcarpa Nees ex Kunth 

Scleria mitis P.J. Bergius 

Scleria muehlenbergii Steud. 

Scleria neogranatensis C.B. Clarke 

Scleria parallela C.B. Clarke 

Scleria pinetorum Britton 

Scleria plusiophylla Steud. 

Scleria purdiei C.B. Clarke 

Scleria ramosa C.B. Clarke 

Scleria robusta Camelbeke & Goetgh. 

Scleria scabra Willd. 

Scleria secans (L.) Urb. 

Scleria splitgerberiana Henrard ex Uittien 

Scleria staheliana Uittien 

Scleria stipitata Uittien 

Scleria stipularis Nees 

Scleria tenacissima Steud. 

Scleria tepuiensis Core 

Scleria triquetra M.T. Strong 

Scleria cf. vaginata Steud. 

Scleria vaginata Steud. 

Scleria verticillata Muhl. ex Willd. 

Scleria violacea Pilg. 

Trilepis kanukuensis Gilly 

Uncinia hamata (Sw.) Urb. 

Websteria confervoides (Poir.) S.S. Hooper 


Abildgaardia monostachyos (L.) Vahl =Abildgaardia ovata 

Ascolepis venezuelensis Schnee =Lipocarpha humboldtiana 

Becquerelia bicolor H. Pfeiff. =Calyptrocarya bicolor 

Becquerelia divaricata (Nees ex Böck.) Palla =Becquerelia cymosa subsp. merkeliana 

Becquerelia glomerulata Brongn. =Calyptrocarya glomerulata 

Becquerelia merkeliana Nees =Becquerelia cymosa subsp. merkeliana 

Bisboeckelera angustifolia (Böck.) Kuntze =Bisboeckelera longifolia 

Bolboschoenus maritimus (L.) Palla =Schoenoplectus maritimus 

Bolboschoenus robustus (Pursh) Soják =Schoenoplectus robustus 

Bulbostylis arenaria (Nees) Lindm. =Bulbostylis juncoides 

Bulbostylis argentina Palla =Bulbostylis juncoides 

Bulbostylis capillaris var. glabra Uittien =Bulbostylis truncata 

Bulbostylis capillaris var. tenuifolia (Rudge) C.B. Clarke =Bulbostylis tenuifolia 

Bulbostylis fendleri C.B. Clarke =Bulbostylis juncoides 

Bulbostylis hirta (Thunb.) Svenson [misapplied to Bulbostylis hispidula] 

Bulbostylis hirtella (Schrad. ex Schult.) Nees ex Urb. =Bulbostylis juncoides 

Bulbostylis juncoides var. lorentzii (Böck.) Kük. =Bulbostylis juncoides 

Bulbostylis langsdorffiana (Kunth) C.B. Clarke =Bulbostylis juncoides 

Bulbostylis papillosa Kük. =Bulbostylis junciformis 

Bulbostylis spadicea (Kunth) Kük. =Bulbostylis paradoxa 

Calyptrocarya fragifera Nees ex Kunth [nomen nudum] =Calyptrocarya glomerulata 

Calyptrocarya intermedia C.B. Clarke =Calyptrocarya glomerulata 

Calyptrostylis ceylanica (Nees) Nees =Rhynchospora triflora 

Calyptrostylis longirostris Nees =Rhynchospora amazonica subsp. guianensis 

Carex hamata Sw. =Uncinia hamata 

Carex hermaphrodita Jacq. =Cyperus hermaphroditus 

GU SU FG 

BO GU SU FG 

GU SU 


FG? 

BO FG 

GU 



SU FG 





SU 

AM 

SU 

GU 

BO 

AM SU 

AM BO 

AM BO GU SU 


GU SU 

SU FG 

GU SU 


AM BO 

AM GU FG 

FG 

SU 

BO 


FG 

AM GU SU FG 

AM BO GU? 

AM BO GU SU? FG

96 


Carex ovata Burm. f. =Abildgaardia ovata 

Cephalocarpus maguirei Gilly =Cephalocarpus rigidus 

Cephalocarpus steyermarkii Gilly =Cephalocarpus rigidus 

Cephaloschoenus ceylanicus Nees =Rhynchospora triflora 

Chaetospora capitata Kunth =Rhynchospora capitata 

Chaetospora globosa Kunth =Rhynchospora globosa 

Chaetospora pterocarpa Kunth =Rhynchospora barbata 

Cladium steyermarkii T. Koyama =Rhynchocladium steyermarkii 

Cryptangium stellatum Böck. =Didymiandrum stellatum 

Cyperus albomarginatus (Nees) Steud. =Cyperus flavicomus 

Cyperus amabilis var. macrostachyus (Böck.) Kük. =Cyperus amabilis 

Cyperus amabilis var. oligostachyus (Kunth) Kük. =Cyperus amabilis 

Cyperus aurantiacus Kunth =Cyperus amabilis 

Cyperus aureus Kunth =Cyperus amabilis 

Cyperus aureus var. aurantiacus (Kunth) Böck. =Cyperus amabilis 

Cyperus aureus var. oligostachyus (Kunth) Böck. =Cyperus amabilis 

Cyperus aureus x macrostachyus Böck. =Cyperus amabilis 

Cyperus brevifolius (Rottb.) Endl. ex Hassk. =Kyllinga brevifolia 

Cyperus cayennensis (Lam.) Britton =Cyperus aggregatus 

Cyperus chalaranthus J. Presl & C. Presl =Cyperus laxus 

Cyperus corymbosus Rottb. =Cyperus articulatus 

Cyperus densicaespitosus Mattf. & Kük. =Kyllinga pumila 

Cyperus diffusus Vahl =Cyperus laxus 

Cyperus dissitiflorus Nees ex Torr. =Cyperus thyrsiflorus 

Cyperus eggersii Böck. =Cyperus odoratus 

Cyperus elegans auct. non L. =Cyperus laxus 

Cyperus ferax Rich. =Cyperus odoratus 

Cyperus flavus (Vahl) Nees =Cyperus aggregatus 

Cyperus fraternus Kunth =Cyperus reflexus 

Cyperus globulosus Aubl. =Cyperus luzulae 

Cyperus globulosus auct., non Aubl. =Cyperus croceus 

Cyperus herndoniae G.C. Tucker =Cyperus camphoratus 

Cyperus hirtus Thunb. =Bulbostylis hispidula 

Cyperus huarmensis (Kunth) M.C. Johnst. =Cyperus aggregatus 

Cyperus kyllingia Endl. =Kyllinga nemoralis 

Cyperus macrocephalus Liebm. =Cyperus odoratus 

Cyperus monostachyos L. =Abildgaardia ovata 

Cyperus nodosus Willd. =Cyperus articulatus var. nodosus 

Cyperus obtusatus (J. Presl & C. Presl) Mattf. & Kük. =Kyllinga pungens 

Cyperus oligostachyus Kunth =Cyperus amabilis 

Cyperus pedunculatus (R. Br.) Kern =Remirea maritima 

Cyperus planifolius Rich. [Not in flora area] 

Cyperus rivularis Kunth =Cyperus bipartitus 

Cyperus ruderalis Uittien =Cyperus trailii 

Cyperus sesquiflorus (Torr.) Mattf. & Kük. =Kyllinga odorata 

Cyperus tolucensis Kunth =Cyperus laxus 

Cyperus uncinatus Poir. [not in flora area] 

Dichromena albida (Nees) Steud. =Rhynchospora albida 

Dichromena brevicaulis Suess. =Rhynchospora duckei 

Dichromena candida (Nees) Ridl. =Rhynchospora candida 

Dichromena caracasana Kunth =Rhynchospora caracasana 

Dichromena cariciformis (Nees) J.F. Macbr. =Rhynchospora cariciformis 

Dichromena ciliata Vahl [nom. illeg., non Dichromena ciliata Pers. (1805)] =Rhynchospora ciliata 

Dichromena ciliata var. cinnamomea Kük. =Rhynchospora ciliata 

Dichromena exaltata (Kunth) J.F. Macbr. =Rhynchospora exaltata 

Dichromena filiformis (Vahl) Kunth =Rhynchospora filiformis 

Dichromena junciformis Kunth =Rhynchospora junciformis 

Dichromena monostachya (Böck.) C.B. Clarke =Rhynchospora albescens 

Dichromena montana Uittien =Rhynchospora montana 

Dichromena nervosa Vahl =Rhynchospora nervosa 

Dichromena persooniana Nees =Rhynchospora nervosa 

Dichromena pilosa Kunth =Rhynchospora pilosa 

Dichromena pubera Vahl =Rhynchospora pubera 

Dichromena radicans Schltdl. & Cham. =Rhynchospora radicans


Dichromena repens Vahl =Rhynchospora reptans 

Dichromena rigidifolia Gilly =Rhynchospora rigidifolia 

Dichromena robusta Kunth =Rhynchospora robusta 

Dichromena squarrosa Willd. ex Link (Rapateaceae) =Duckea squarrosa 

Dichromena tenella (Nees) Steud. =Rhynchospora tenella 

Dichromena triflora (Vahl) Macbride =Rhynchospora triflora 

Dichromena velutina Kunth =Rhynchospora velutina 

Didymiandrum flexifolium Gilly =Everardia flexifolia 

Didymiandrum guaiquinimae Schnee =Didymiandrum stellatum 

Diplacrum laevissimum (Kük.) Uittien =Diplacrum guianense 

Diplacrum longifolium (Griseb.) C.B. Clarke ex T. Durand & Schinz =Diplacrum capitatum 

Diplacrum notopterum (Nees ex Steud.) C.B. Clarke =Diplacrum guianense 

Diplasia pycnostachya Benth. =Mapania pycnostachya 

Echinoschoenus beyrichii Nees =Rhynchospora exaltata 

Eleocharis atropurpurea (Retz.) J. Presl & C. Presl [not in flora area] 

Eleocharis bicolor Chapm. =Eleocharis minima var. bicolor 

Eleocharis caribaea (Rottb.) S.F. Blake =Eleocharis geniculata 

Eleocharis confervoides (Poir.) G. Tucker =Websteria confervoides 

Eleocharis elata Böck. =Eleocharis plicarhachis 

Eleocharis fistulosa (Poir.) Schult. [nom. superfl.] =Eleocharis acutangula 

Eleocharis fistulosa (Poir.) Link [invalidly published] =Eleocharis acutangula 

Eleocharis flaccida (Rchb. ex Spreng.) Urb. =Eleocharis flavescens 

Eleocharis flaccida auct. non (Rchb. ex Spreng.) Urb. sensu Svenson =Eleocharis flavescens 

Eleocharis homonyma Steud. =Eleocharis sellowiana 

Eleocharis nodulosa (Roth) Schult. =Eleocharis montana 

Eleocharis sagotii C.B. Clarke =Eleocharis jelskiana 

Eleocharis sagotii var. glochidiata C.B. Clarke =Eleocharis plicarhachis 

Eleocharis sellowiana var. homonyma (Steud.) H. Pfeiff. =Eleocharis sellowiana 

Eleocharis sulcata Nees [nomen nudum] =Eleocharis filiculmis 

Eleocharis sulcata Nees ex Böck. [nom. superfl.] =Eleocharis filiculmis 

Ephippiorhynchium triflorum (Vahl) Nees =Rhynchospora triflora 

Ephippiorhynchium trispicatum Nees =Rhynchospora trispicata 

Everardia duidae Gilly =Everardia montana subsp. duidae 

Everardia erecto-laxa T. Koyama & Maguire =Everardia angusta 

Everardia glaucifolia Gilly =Everardia montana subsp. glaucifolia 

Everardia gracilis Gilly =Everardia angusta 

Everardia guaiquinimae Schnee =Everardia montana subsp. guaiquinimae 

Everardia hitchcockii Lasser & Maguire =Everardia montana subsp. montana 

Everardia montana var. pilosa Gilly =Everardia montana subsp. montana 

Everardia ptariensis Gilly =Everardia montana subsp. ptariensis 

Everardia revoluta Gilly =Everardia montana subsp. montana 

Everardia steyermarkii Gilly =Everardia angusta 

Exochogyne steyermarkii Gilly =Exochogyne amazonica 

Fimbristylis annua var. diphylla (Retz.) Kük. =Fimbristylis dichotoma 

Fimbristylis apus (A. Gray) S. Watson =Fimbristylis vahlii 

Fimbristylis autumnalis var. complanata (Retz.) Barros =Fimbristylis complanata 

Fimbristylis baldwiniana (Schult.) Torr. =Fimbristylis annua 

Fimbristylis bufonia (Kunth) Alain =Bulbostylis tenuifolia 

Fimbristylis capillaris var. pilosa Britton =Bulbostylis juncoides 

Fimbristylis chirigota (C. Wright) Alain =Bulbostylis hispidula 

Fimbristylis comata Nees =Fimbristylis squarrosa 

Fimbristylis congesta Torr. =Fimbristylis vahlii 

Fimbristylis cymosa subsp. spathacea (Roth) T. Koyama =Fimbristylis cymosa 

Fimbristylis dichotoma f. annua (All.) Owhi =Fimbristylis annua 

Fimbristylis diphylla (Retz.) Vahl =Fimbristylis dichotoma 

Fimbristylis divaricata Ham. =Rhynchospora divaricata 

Fimbristylis frankii Steud. =Fimbristylis autumnalis 

Fimbristylis glomerata (Retz.) Urb. =Fimbristylis cymosa 

Fimbristylis hirta (Kunth) Roem. & Schult. =Fimbristylis squarrosa 

Fimbristylis hirta (Thunb.) Roem. & Schult. ex Kük. [name has been misapplied to Bulbostylis hispidula] =Fimbristylis 

squarrosa 

Fimbristylis hispidula (Vahl) Kunth =Bulbostylis hispidula 

Fimbristylis juncoides (Vahl) Alain =Bulbostylis juncoides 

Fimbristylis laxa Vahl =Fimbristylis annua

98 


Fimbristylis miliacea (L.) Vahl [Guianas collections referable to F. littoralis] =Fimbristylis littoralis 

Fimbristylis monostachyos (L.) Hassk. =Abildgaardia ovata 

Fimbristylis obtusifolia (Vahl) Kunth =Fimbristylis cymosa 

Fimbristylis ovata (Burm. f.) J. Kern =Abildgaardia ovata 

Fimbristylis papillosa (Kük.) Alain =Bulbostylis junciformis 

Fimbristylis polymorpha Böck. =Fimbristylis dichotoma 

Fimbristylis savannarum Alain =Bulbostylis juncoides 

Fimbristylis schomburgkiana Steud. =Bulbostylis schomburgkiana 

Fimbristylis spadicea f. domingensis (Pers.) Kük. =Fimbristylis spadicea 

Fimbristylis spathacea Roth =Fimbristylis cymosa 

Fimbristylis speciosa Rohde ex Spreng. =Fimbristylis spadicea 

Fimbristylis verrucosa C. Presl =Fimbristylis annua 

Fimbristylis vestita (Kunth) Hemsl. =Bulbostylis vestita 

Fuirena paniculata L. f. =Fuirena umbellata 

Fuirena tereticulmis C. Presl =Fuirena umbellata 

Fuirena verticillata Spreng. =Lagenocarpus verticillatus 

Haloschoenus albidus Nees =Rhynchospora albida 

Haloschoenus caracasanus (Kunth) Nees =Rhynchospora caracasana 

Haloschoenus contractus Nees =Rhynchospora contracta 

Haloschoenus emaciatus Nees =Rhynchospora emaciata 

Haloschoenus riparius Nees =Rhynchospora riparia 

Haloschoenus tenellus Nees =Rhynchospora tenella 

Haplostylis mexicana Liebm. =Rhynchospora mexicana 

Hemicarpha micrantha (Vahl) Pax =Lipocarpha micrantha 

Hemicarpha schomburgkii Friedl. =Lipocarpha schomburgkii 

Hoppia irrigua Nees =Bisboeckelera irrigua 

Hypolytrum fuscum Nees =Hypolytrum longifolium 

Hypolytrum imeriense R. Gross =Mapania imeriensis 

Hypolytrum irriguum Nees =Hypolytrum longifolium 

Hypolytrum macrophyllum Böck. =Mapania macrophylla 

Hypolytrum micranthum Böck. =Hypolytrum strictum 

Hypolytrum nicaraguense Liebman =Hypolytrum longifolium subsp. nicaraguense 

Hypolytrum punctatum Nees =Hypolytrum longifolium subsp. sylvaticum 

Hypolytrum pycnocephalum Benth. =Mapania pycnocephala 

Hypolytrum rubescens Huber ex C.B. Clarke =Hypolytrum longifolium subsp. rubescens 

Hypolytrum sandwithii Gilly =Mapania imeriensis 

Hypolytrum scapigerum Nees =Hypolytrum strictum 

Hypolytrum sylvaticum Poepp. & Kunth =Hypolytrum longifolium subsp. sylvaticum 

Hypolytrum tepuianum Steyerm. =Mapania tepuiana 

Hypoporum micrococcum Liebm. =Scleria verticillata 

Iria monostachya (L.) Kuntze =Abildgaardia ovata 

Isolepis albescens Miq. =Rhynchospora albescens 

Isolepis bufonia Kunth =Bulbostylis tenuifolia 

Isolepis capillaris (L.) Roem. & Schult. =Bulbostylis capillaris 

Isolepis conifera Kunth =Bulbostylis conifera 

Isolepis dipsacea (Rottb.) Roem. & Schult. =Fimbristylis dipsacea 

Isolepis hirta Kunth =Fimbristylis squarrosa 

Isolepis hirtella Schrad. ex Schult. =Bulbostylis juncoides 

Isolepis humboldtii Roem. & Schult. =Lipocarpha micrantha 

Isolepis junciformis Kunth =Bulbostylis junciformis 

Isolepis lanata Kunth =Bulbostylis lanata 

Isolepis langsdorffiana Kunth =Bulbostylis juncoides 

Isolepis leucostachya Kunth =Bulbostylis leucostachya 

Isolepis miliacea (L.) C. Presl =Fimbristylis littoralis 

Isolepis paradoxa (Spreng.) Kunth =Bulbostylis paradoxa 

Isolepis radiciflora Steud. =Bulbostylis capillaris 

Isolepis vahlii (Lam.) Kunth =Fimbristylis vahlii 

Isolepis vestita Kunth =Bulbostylis vestita 

Isolepis willdenowii Roem. & Schult. =Fimbristylis complanata 

Kyllinga cayennensis Lam. =Cyperus aggregatus 

Kyllinga filiformis Sw. =Cyperus swartzii 

Kyllinga monocephala Rottb., nom. illeg. =Kyllinga nemoralis 

Kyllinga peruviana Lam. =Kyllinga vaginata 

Kyllinga rigida Baldwin =Kyllinga vaginata


Kyllinga sesquiflora Torr. =Kyllinga odorata 

Kyllinga squarrosa Baldwin =Cyperus aggregatus 

Lagenocarpus amazonicus Davidse =Lagenocarpus venezuelensis 

Lagenocarpus amazonicus (C.B. Clarke) H. Pfeiff. =Exochogyne amazonica 

Lagenocarpus kunthii (Miq.) Uittien =Lagenocarpus verticillatus 

Lagenocarpus tremulus Nees =Lagenocarpus rigidus subsp. tremulus 

Lagenocarpus weigelti (Spreng.) Uittien =Lagenocarpus verticillatus 

Leptoschoenus prolifer Nees =Rhynchospora schomburgkiana 

Lipocarpha maculata auct. non (Michx.) Torrey 1836: sensu neotropical =Lipocarpha mexicana 

Lipocarpha sellowiana Kunth =Lipocarpha humboldtiana 

Mapania fluviatilis Sandwith =Mapania pycnocephala subsp. fluviatilis 

Mapaniopsis effusa C.B. Clarke =Mapania effusa 

Mapaniopsis micrococca T. Koyama =Mapania micrococca 

Mariscus aggregatus Willd. =Cyperus aggregatus 

Mariscus brizaeus (Vahl) C.B. Clarke =Cyperus planifolius 

Mariscus cayennensis (Lam.) Urb. =Cyperus aggregatus 

Mariscus chlorocephalus C.B. Clarke =Cyperus chlorocephalus 

Mariscus cornelii-ostenii (Kük.) T. Koyama =Cyperus cornelii-ostenii 

Mariscus dissitiflorus (Torr.) C.B. Clarke =Cyperus thyrsiflorus 

Mariscus elatus Kunth =Cyperus aggregatus 

Mariscus flavus Vahl =Cyperus aggregatus 

Mariscus flavus var. major Nees =Cyperus aggregatus 

Mariscus gayi C.B. Clarke =Cyperus gayi 

Mariscus guianensis C.B. Clarke =Cyperus guianensis 

Mariscus laevis Kunth =Cyperus aggregatus 

Mariscus ligularis (L.) Urb. =Cyperus ligularis 

Mariscus mutisii Kunth =Cyperus mutisii 

Mariscus pedunculatus (R. Br.) T. Koyama =Remirea maritima 

Mariscus swartzii A. Dietr. =Cyperus swartzii 

Mariscus tovarii Kunth =Cyperus hermaphroditus 

Mariscus umbellatus (Rottb.) Vahl =Cyperus subumbellatus 

Micropapyrus viviparoides Suess. =Rhynchospora schomburgkiana 

Mitrospora cephalophora Nees =Rhynchospora exaltata 

Mitrospora pallida Nees =Rhynchospora pallida 

Oncostylis arenaria Nees =Bulbostylis juncoides 

Oncostylis hispida Liebm. =Bulbostylis vestita 

Oncostylis junciformis var. humboldtiana (Spreng.) Nees =Bulbostylis junciformis 

Oncostylis paradoxa (Spreng.) Nees =Bulbostylis paradoxa 

Oncostylis tenuifolia (Rudge) Nees =Bulbostylis tenuifolia 

Oncostylis truncata Nees =Bulbostylis truncata 

Oncostylis vestita (Kunth) Nees =Bulbostylis vestita 

Oreobolus obtusangulus var. rubrovaginatus Kük. =Oreobolus goeppingeri 

Oreobolus obtusangulus subsp. rubrovaginatus (Kük.) T. Koyama =Oreobolus goeppingeri 

Platylepis brasiliensis Kunth =Ascolepis brasiliensis 

Pogonostylis squarrosus (Vahl) Bertol. =Fimbristylis squarrosa 

Psilocarya candida Nees =Rhynchospora candida 

Psilocarya robusta (Kunth) Nees =Rhynchospora robusta 

Psilocarya rufa Nees =Rhynchospora velutina 

Psilocarya velutina (Kunth) Nees =Rhynchospora velutina 

Pteroscleria guianensis Nees =Diplacrum guianense 

Pteroscleria longifolia Griseb. =Diplacrum capitatum 

Pycreus albomarginatus Mart. & Schrad. ex Nees =Cyperus flavicomus 

Pycreus capillifolius (A. Rich.) C.B. Clarke =Cyperus capillifolius 

Pycreus flavescens (L.) Rchb. =Cyperus flavescens 

Pycreus megapotamicus (Kunth) Nees =Cyperus megapotamicus 

Pycreus polystachyos (Rottb.) P. Beauv. =Cyperus polystachyos 

Pycreus pumilus (L.) Nees =Cyperus pumilus 

Pycreus unioloides (R. Br.) Urb. =Cyperus unioloides 

Rhynchospora amazonica var. guianensis Kük. =Rhynchospora amazonica subsp. guianensis 

Rhynchospora aristata var. immensa (Kük.) Kük. =Rhynchospora immensa 

Rhynchospora arundinacea Böck. =Rhynchospora exaltata 

Rhynchospora aurea Vahl =Rhynchospora corymbosa 

Rhynchospora baileyi Britton ex L.H. Bailey =Rhynchospora albomarginata 

Rhynchospora beyrichii (Nees) Steud. =Rhynchospora exaltata

100 


Rhynchospora borinquensis Britton =Rhynchospora marisculus 

Rhynchospora buchii Urb. =Rhynchospora mexicana 

Rhynchospora canaliculata Böck. =Rhynchospora riparia 

Rhynchospora candida var. pilosa Schnee =Rhynchospora candida 

Rhynchospora cariciformis var. trichophora (Nees) Kük. =Rhynchospora trichophora 

Rhynchospora cephalophora (Nees) Steud. =Rhynchospora exaltata 

Rhynchospora cephalotes var. angustifolia Uittien =Rhynchospora pubisquama 

Rhynchospora cephalotes var. imeriensis Kük. =Rhynchospora imeriensis 

Rhynchospora ceylanica (Nees) Kunth =Rhynchospora triflora 

Rhynchospora clarkei Rose =Rhynchospora brevirostris 

Rhynchospora cyperoides (Sw.) Mart. =Rhynchospora holoschoenoides 

Rhynchospora denticulata Huber =Rhynchospora trichochaeta 

Rhynchospora distans Vahl =Rhynchospora fascicularis 

Rhynchospora elegans Kük. =Rhynchospora eximia 

Rhynchospora exaltata var. ovalis Kük. =Rhynchospora exaltata 

Rhynchospora fascicularis var. distans (Michx.) Chapm. [not in flora area] misidentifications =R. fascicularis subsp. 

fascicularis] 

Rhynchospora filifolia A. Gray [not in flora area. Identifications =R. harperi] 

Rhynchospora flexuosa var. tenerrima (Böck.) Kük. =Rhynchospora tepuiana 

Rhynchospora flexuosa subsp. tepuiana (Steyerm.) T. Koyama =Rhynchospora tepuiana 

Rhynchospora glauca Vahl =Rhynchospora rugosa 

Rhynchospora graminea Uittien =Rhynchospora spruceana 

Rhynchospora guianensis Lindeman & Donsel. =Rhynchospora trispicata 

Rhynchospora heterocaulis C.B. Clarke =Rhynchospora barbata p.p. & R. holoschoenoides p.p. 

Rhynchospora hispidula (Vahl) Böck. [nom. illeg.] =Rhynchospora divaricata 

Rhynchospora igarapicola T. Koyama =Rhynchospora cariciformis 

Rhynchospora junciformis var. monocarpa Kük. =Rhynchospora junciformis 

Rhynchospora karuaiana Steyerm. =Rhynchospora triflora 

Rhynchospora longispicata Böck. =Rhynchospora filiformis 

Rhynchospora macrochaeta var. ruiziana (Böck.) Kük. =Rhynchospora ruiziana 

Rhynchospora mexicana var. trichochaeta (C.B. Clarke) Kük. =Rhynchospora trichochaeta 

Rhynchospora micrantha Vahl [nom. illeg.] =Rhynchospora contracta 

Rhynchospora minutiflora (Rich. ex Spreng.) Adams =Rhynchospora contracta 

Rhynchospora monostachya Böck. =Rhynchospora albescens 

Rhynchospora nervosa var. ciliata (G. Mey.) Kük. =Rhynchospora ciliata 

Rhynchospora nervosa subsp. ciliata (G. Mey.) T. Koyama =Rhynchospora ciliata 

Rhynchospora nervosa var. cinnamomea (Kük.) Kük. =Rhynchospora ciliata 

Rhynchospora oxycephala C. Wright =Rhynchospora eximia 

Rhynchospora patuligluma Lindm. =Rhynchospora pallida 

Rhynchospora perrigida Böck. =Bulbostylis paradoxa 

Rhynchospora persooniana (Nees) Griseb. =Rhynchospora nervosa 

Rhynchospora pilosa subsp. arenicola (Uittien) T. Koyama =Rhynchospora arenicola 

Rhynchospora podosperma C. Wright =Rhynchospora filiformis 

Rhynchospora pringlei C.B. Clarke =Rhynchospora brevirostris 

Rhynchospora prolifera (Nees) C.B. Clarke =Rhynchospora schomburgkiana 

Rhynchospora psilocaryoides Griseb. =Rhynchospora eximia 

Rhynchospora ptaritepuiana Steyerm. =Rhynchospora lechleri 

Rhynchospora pterocarpa (Kunth) Roem. & Schult. =Rhynchospora barbata 

Rhynchospora pungens Liebm. =Rhynchospora rugosa 

Rhynchospora repens (Vahl) Böck. =Rhynchospora reptans 

Rhynchospora rigidifolia subsp. capillacea T. Koyama =Rhynchospora capillifolia 

Rhynchospora rigidifolia subsp. planifolia T. Koyama =Rhynchospora planifolia 

Rhynchospora rufa (Nees) Böck. =Rhynchospora velutina 

Rhynchospora setacea (Bergius) Böck. =Rhynchospora fallax 

Rhynchospora sororopana Steyerm. =Rhynchospora biflora 

Rhynchospora stellata (Lam.) Griseb. [not in flora area] =Rhynchospora colorata (L.) H. Pfeiffer 

Rhynchospora stricta var. demerarensis (C.B. Clarke) Kük. =Rhynchospora cajennensis 

Rhynchospora tamana Steyerm. =Rhynchospora macrochaeta 

Rhynchospora tenella f. haplostylis Uittien =Rhynchospora tenella 

Rhynchospora tenella var. haplostylis (Uittien) Kük. =Rhynchospora tenella 

Rhynchospora tenerrima Böck. =Rhynchospora tepuiana 

Rhynchospora tenerrima var. fallax (Uittien) Kük. =Rhynchospora fallax 

Rhynchospora tenerrima subsp. fallax (Uittien) T. Koyama =Rhynchospora fallax 

Rhynchospora tenuis subsp. riparia (Nees) T. Koyama =Rhynchospora riparia


Rhynchospora tenuiseta C. Wright =Rhynchospora marisculus 

Rhynchospora triceps Steud. =Rhynchospora exaltata 

Rhynchospora trichochaeta var. venezuelensis Steyerm. =Rhynchospora subplumosa 

Rhynchospora trichodes C.B. Clarke =Rhynchospora divaricata 

Rhynchospora venezolana Kük. =Rhynchospora hirsuta 

Rhynchospora viridilutea C.B. Clarke =Rhynchospora hassleri 

Schoenus barbatus Vahl =Rhynchospora barbata 

Schoenus ciliatus G. Mey. [nom. illeg., non Dichromena ciliata Pers. (1805)] =Rhynchospora ciliata 

Schoenus comatus Link =Rhynchospora comata 

Schoenus cyperoides Sw. =Rhynchospora holoschoenoides 

Schoenus flavus Link (Rapateaceae) =Duckea flava 

Schoenus hirsutus Vahl =Rhynchospora hirsuta 

Schoenus hispidulus Vahl =Rhynchospora divaricata 

Schoenus holoschoenoides Rich. =Rhynchospora holoschoenoides 

Schoenus juncoides Vahl =Bulbostylis juncoides 

Schoenus paradoxus Spreng. =Bulbostylis paradoxa 

Schoenus rugosus Vahl =Rhynchospora rugosa 

Schoenus secans L. =Scleria secans 

Schoenus setaceus P.J. Bergius =Rhynchospora fallax 

Scirpidium nigrescens Nees =Eleocharis nigrescens 

Scirpidium sulcatum Nees [nom. superfl.] =Eleocharis filiculmis 

Scirpus acutangulus Roxb. =Eleocharis acutangula 

Scirpus aestivalis Retz. =Fimbristylis aestivalis 

Scirpus annuus All. =Fimbristylis annua 

Scirpus apus A. Gray =Fimbristylis vahlii 

Scirpus aturensis Maury =Bulbostylis aturensis 

Scirpus autumnalis L. =Fimbristylis autumnalis 

Scirpus baldwinianus Schult. =Fimbristylis annua 

Scirpus brachyphyllus Link =Bulbostylis capillaris 

Scirpus bufonis (Kunth) Poir. =Bulbostylis tenuifolia 

Scirpus capillaris L. =Bulbostylis capillaris 

Scirpus caribaeus Rottb. =Eleocharis geniculata 

Scirpus cephalotes L. =Rhynchospora cephalotes 

Scirpus chirigota C. Wright =Bulbostylis hispidula 

Scirpus complanatus Retz. =Fimbristylis complanata 

Scirpus confervoides Poir. =Websteria confervoides 

Scirpus corymbosus L. =Rhynchospora corymbosa 

Scirpus cubensis Poepp. & Kunth =Oxycaryum cubense 

Scirpus dichotomus L. =Fimbristylis dichotoma 

Scirpus diphyllus Retz. =Fimbristylis dichotoma 

Scirpus dipsaceus Rottb. =Fimbristylis dipsacea 

Scirpus domingensis Pers. =Fimbristylis spadicea 

Scirpus elegans Kunth =Eleocharis elegans 

Scirpus fistulosus Poir. =Eleocharis acutangula 

Scirpus flavescens Poir. =Eleocharis flavescens 

Scirpus geniculatus L. =Eleocharis geniculata 

Scirpus glomeratus Retz. =Fimbristylis cymosa 

Scirpus hirtus Griseb. =Bulbostylis vestita 

Scirpus hispidulus Vahl =Bulbostylis hispidula 

Scirpus humboldtiana Spreng. =Bulbostylis junciformis 

Scirpus interstinctus Vahl =Eleocharis interstincta 

Scirpus lithospermus L. =Scleria lithosperma 

Scirpus longifolius Rich. =Hypolytrum longifolium 

Scirpus lorentzii Böck. =Bulbostylis juncoides 

Scirpus luzulae L. =Cyperus luzulae 

Scirpus maculosus Vahl =Eleocharis maculosa 

Scirpus maritimus L. =Schoenoplectus maritimus 

Scirpus maritimus var. robustus (Pursh) Kük. =Schoenoplectus robustus 

Scirpus maritimus ß macrostachyus Michx. =Schoenoplectus robustus 

Scirpus micranthus Vahl =Lipocarpha micrantha 

Scirpus miliaceus L. =Fimbristylis littoralis 

Scirpus mitratus Griseb. =Eleocharis mitrata 

Scirpus montanus Kunth =Eleocharis montana 

Scirpus mucronulatus Michx. =Fimbristylis autumnalis

102 


Scirpus mutatus L. =Eleocharis mutata 

Scirpus nodulosus Roth =Eleocharis montana 

Scirpus obtusifolius Vahl =Fimbristylis cymosa 

Scirpus pachystylus C. Wright =Eleocharis pachystyla 

Scirpus plicarhachis Griseb. =Eleocharis plicarhachis 

Scirpus pulcher Rudge =Hypolytrum pulchrum 

Scirpus quinquangularis Vahl =Fimbristylis quinquangularis 

Scirpus retroflexus Poir. =Eleocharis retroflexa 

Scirpus robustus Pursh =Schoenoplectus robustus 

Scirpus schomburgkianus Böck. =Rhynchospora schomburgkiana 

Scirpus spadiceus L. =Fimbristylis spadicea 

Scirpus submersus C. Wright =Websteria confervoides 

Scirpus sulcatus Roth =Eleocharis filiculmis 

Scirpus tenuifolius Rudge =Bulbostylis tenuifolia 

Scirpus triflorus (Vahl) Poir. =Rhynchospora triflora 

Scirpus umbellatus (Rottb.) Kuntze =Fuirena umbellata 

Scirpus vahlii Lam. =Fimbristylis vahlii 

Scirpus vestitus (Kunth) Rchb. ex Böck. =Bulbostylis vestita 

Scleria arundinacea Kunth =Scleria latifolia 

Scleria bracteata f. supragynacea H. Pfeiff. =Scleria bracteata 

Scleria capitata Willd. =Diplacrum capitatum 

Scleria flagellum Sw. =Scleria secans 

Scleria foliosa C. Wright, nom. illeg. =Scleria microcarpa 

Scleria grandis Core non Labill =Scleria robusta 

Scleria hirtella auct. non Sw. =Scleria distans 

Scleria latifolia Nees =Scleria latifolia 

Scleria latifolia var. arundinacea (Kunth) H. Pfeiff. =Scleria latifolia 

Scleria liebmannii Steud. =Scleria verticillata 

Scleria micrococca (Liebm.) Steud. =Scleria verticillata 

Scleria minutiflora Rich. ex Spreng. [nom. inval.] =Rhynchospora contracta 

Scleria nutans Willd. ex Kunth =Scleria distans 

Scleria paludosa Poepp. & Kunth =Scleria macrophylla 

Scleria pterota C. Presl [nomen nudum] =Scleria melaleuca 

Scleria pterota var. melaleuca (Rchb. ex Schltdl. & Cham.) Uittien =Scleria melaleuca 

Scleria pterota var. melanocarpa Nees [nomen nudum] =Scleria melaleuca 

Scleria pyramidalis Hochst. ex Steud. [=Scleria stipularis?] =Scleria martii 

Scleria reflexa Kunth =Scleria secans 

Scleria reticularis Michx. [Not in flora area. Restricted to E. US]] =Scleria muehlenbergii 

Scleria rigida Kunth =Lagenocarpus rigidus 

Scleria setacea Poir. =Scleria muehlenbergii 

Scleria tenella Kunth =Scleria verticillata 

Scleria tuberculata Böck. =Becquerelia tuberculata 

Scleria weigeltiana Schrad. ex Böck. =Scleria secans 

Spermodon eximius Nees =Rhynchospora eximia 

Spermodon filiformis (Vahl) Nees =Rhynchospora filiformis 

Stenophyllus capillaris (L.) Britton =Bulbostylis capillaris 

Stenophyllus paradoxus (Spreng.) Standl. =Bulbostylis paradoxa 

Stenophyllus truncata (Nees) H. Pfeiff. =Bulbostylis truncata 

Stenophyllus vestitus (Kunth) Britton =Bulbostylis vestita 

Torulinium ferax (Rich.) Urb. =Cyperus odoratus 

Torulinium odoratum (L.) S.S. Hooper =Cyperus odoratus 

Trichelostylis complanata (Retz.) Nees =Fimbristylis complanata 

Trichelostylis miliacea (L.) Nees =Fimbristylis littoralis 

Trichelostylis mucronulata (Michx.) Torr. =Fimbristylis autumnalis 

Trichelostylis quinquangularis (Vahl) Nees =Fimbristylis quinquangularis 

Trichelostylis stricta Nees =Bulbostylis schomburgkiana 

Trichochaeta tenuis Steud. =Rhynchospora trichochaeta 

REFERENCES 

Core, E.L. 1965. Cyperaceae tribe Sclerieae. In B. Maguire & collaborators, Botany of the Guayana Highland Part VI. Mem. 

New York Bot. Gard. 12(3): 1-285. 

Cremers, G. & M. Hoff. 1993. Inventaire taxonomique des plantes de la Guyane française III - les Cyperaceae et les Poaceae. 

Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Sér. Patr. Génétique) 11: 1-213.


Denton, M. F. 1978. A taxonomic treatment of the Luzulae group of Cyperus. Contributions from the University of Michigan 

Herbarium 11: 197–271. 

Kearns, D.M., W.W. Thomas, G.C. Tucker, R. Kral, K. Camelbeke, D.A. Simpson, A. A. Reznicek, M.S. González-Elizondo, 

M.T. Strong, & P. Goetghebeur. 1998. Cyperaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan 

Guayana 4: 486-663. 

Koyama, T. and B. Maguire. 1965. Cyperaceae tribe Lagenocarpeae. In B. Maguire & collaborators, Botany of the Guayana 

Highland part 6. Mem. New York Bot. Gard. 12(3): 1-285. 

Koyama, T. 1967. Cyperaceae - Mapanioideae. In B. Maguire & collaborators, Botany of the Guayana Highland part 7. Mem. 


Koyama, T. 1972. Cyperaceae - Rhynchosporeae and Cladieae. In B. Maguire & collaborators, Botany of the Guayana Highland 

part 9. Mem. New York Bot. Gard. 23: 1-832. 

Kral, R. 1971. A treatment of Abildgaardia, Bulbostylis, and Fimbristylis (Cyperaceae) for North America. Sida 4: 57-227. 

Kükenthal, G. 1935–36. Cyperaceae-Scirpoideae-Cypereae Cyperus. In H. G. A. Engler, Das Pflanzenreich IV. 20(Heft 101). 

671 pp. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Cyperaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 310-314. 

Simpson, D.A. 1992. A revision of the genus Mapania. Royal Botanic Gardens, Kew. 

Strong, M.T. 2001. Novelties in Rhynchospora (Cyperaceae) from the Guianas. Novon 11: 261-273. 

Thomas, W.W. 1984. The systematics of Rhynchospora section Dichromena. Mem. New York Bot. Gard. 37: 1-116. 

Thomas, W.W. 1997. Cyperaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Tucker, G. C. 1983. The taxonomy of Cyperus (Cyperaceae) in Costa Rica and Panama. Systematic Botany Monographs 2. 83 

pp. 

Tucker, G. C. 1987. The genera of Cyperaceae in the southeastern United States. Journal of the Arnold Arboretum 68: 361–445. 

Tucker, G. C. 1994. A revision of the Mexican species of Cyperus (Cyperaceae). Systematic Botany Monographs 43. 212 pp. 

Uittien, H. 1934. Cyperaceae. In A. Pulle, Flora of Suriname 1(1): 72-149. 

Uittien, H. 1943. Cyperaceae (additions and corrections). In A. Pulle (ed.), Flora of Suriname 1(1): 492-505. 

DIOSCOREACEAE 

O. Téllez 

Dioscorea abysmophila Maguire & Steyerm. 

Dioscorea alata L. [introduced] 

Dioscorea altissima Lam. 

Dioscorea amazonum Mart. ex Griseb. 

Dioscorea aspera Humb. & Bonpl. ex Willd. 

Dioscorea atrescens R. Knuth 

Dioscorea bolivarensis Steyerm. 

Dioscorea bulbifera L. [introduced] 

Dioscorea cayenensis Lam. [introduced?] 

Dioscorea cordifolia Laness. 

Dioscorea crotalariifolia Uline 

Dioscorea cuspidata Humb. & Bonpl. ex Willd. 

Dioscorea decorticans C. Presl 

Dioscorea discolor Kunth 

Dioscorea dodecaneura Vell. 

Dioscorea effusa Griseb. 

Dioscorea esculenta (Lour.) Prain [cultivated only?] 

Dioscorea georgensis R. Knuth [=Dioscorea amazonum?] 

Dioscorea guianensis R. Knuth 

Dioscorea hastata Vell. [=Dioscorea hasslerana?] 

Dioscorea hitchcockii R. Knuth 

Dioscorea holmioidea Maury 

Dioscorea laxiflora Mart. ex Griseb. 

Dioscorea macrostachya Benth. 

Dioscorea manicorensis Clayton, sp. nov. ined. 

Dioscorea megacarpa Gleason 

Dioscorea melastomatifolia Uline ex Prain & Burkill [=Dioscorea truncata?] 

Dioscorea microura R. Knuth 

Dioscorea neblinensis Maguire & Steyerm. 

Dioscorea nitida R. Knuth 

Dioscorea oblonga Gleason 

Dioscorea pachycarpa Kunth 

AM 




BO 

AM BO 

BO 

SU FG 

AM GU SU FG 

FG 

AM GU 

AM 

BO 

BO 

GU SU FG 

AM 

FG 

GU 

GU 

FG 

GU 

AM 

AM 

GU? 

SU FG 

GU SU FG 

FG 

GU 

AM 

AM 

GU SU 

GU?

104 


Dioscorea panamensis R. Knuth 

Dioscorea pilosiuscula Bertero ex Spreng. 

Dioscorea piperifolia Humb. & Bonpl. ex Willd. 

Dioscorea pittieri R. Knuth 

Dioscorea polygonoides Humb. & Bonpl. ex Willd. 

Dioscorea pomeroonensis R. Knuth 

Dioscorea potarensis R. Knuth 

Dioscorea pubescens Poir. [insufficiently known species] 

Dioscorea quinquefoliolata R. Knuth 

Dioscorea riparia Kunth & R.H. Schomb. [=Dioscorea samydea?] 

Dioscorea sagittata Poir. [insufficiently known species] 

Dioscorea samydea Griseb. 

Dioscorea scabra Humb. & Bonpl. ex Willd. 

Dioscorea sororopana Steyerm. 

Dioscorea sp. A 

Dioscorea syringifolia Kunth & R.H. Schomb. 

Dioscorea trachyandra Griseb. 

Dioscorea trachycarpa Kunth [insufficiently known species] 

Dioscorea trichanthera Gleason 

Dioscorea trifida L. f. 

Dioscorea trifoliata Kunth 

Dioscorea truncata Miq. [=Dioscorea trifida?] 


Dioscorea affinis Kunth =Dioscorea trifida 

Dioscorea amazonum var. consanguinea (Kunth) Uline =Dioscorea amazonum 

Dioscorea bulbifera Griseb., non L. =Dioscorea pilosiuscula 

Dioscorea cuspidata Klotzsch ex Kunth =Dioscorea amazonum 

Dioscorea elegantula Kunth =Dioscorea amazonum 

Dioscorea holtii R. Knuth =Dioscorea amazonum 

Dioscorea kegeliana Griseb. =Dioscorea polygonoides 

Dioscorea lanosa Gleason =Dioscorea pilosiuscula 

Dioscorea lindmanii Uline ex R. Knuth =Dioscorea pilosiuscula 

Dioscorea megalobotrya Kunth & R.H. Schomb. ex Kunth =Dioscorea amazonum 

Dioscorea pichinchensis R. Knuth =Dioscorea cuspidata 

Dioscorea pimichinensis R. Knuth ex Sandwith =Dioscorea cuspidata 

Dioscorea schomburgkiana Kunth & R.H. Schomb. =Dioscorea pilosiuscula 

Dioscorea surinamensis G. Mey. =Dioscorea amazonum 

Dioscorea truncata Prain, non Miq. =Dioscorea megacarpa 

Dioscorea truncata R.H. Schomb., non Miq. =Dioscorea megacarpa 

Helmia consanguinea Kunth =Dioscorea amazonum 

REFERENCES 

Gleason, H.A. 1925. Studies on the flora of northern South America. V. Miscellaneous new or noteworthy species. Bull. Torrey 

Bot. Club 52(5): 181-196. 

Knuth, R. 1924. Dioscoreaceae. In A. Engler, Das Pflanzenreich IV(43): 1-387. 

Mori, S.A. 1997. Dioscoreaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Téllez, O. 1998. Dioscoreaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 686-696. 

ERIOCAULACEAE 

M. Hakki, N. Hensold 

Eriocaulon cinereum R. Br. 

Eriocaulon dimorphopetalum Moldenke 

Eriocaulon guyanense Körn. 

Eriocaulon heterodoxum Moldenke 

Eriocaulon humboldtii Kunth 

Eriocaulon jauense Moldenke 

Eriocaulon melanocephalum Kunth 

Eriocaulon spongiola Hensold 

Eriocaulon steyermarkii Moldenke 

Eriocaulon stramineum Körn. [possibly =Eriocaulon guyanense] 

BO 


BO GU 

AM 


GU 

GU 

FG 

GU 

GU SU 

FG 

GU SU 

BO 

BO 

AM 

GU FG 

GU 

GU 

AM BO GU 

AM GU SU FG 

AM 

SU 

AM BO 

BO 

BO GU FG 

GU 


AM BO 


AM 

AM BO GU SU 

FG?


Eriocaulon tenuifolium Klotzsch ex Körn. 

Leiothrix celiae Moldenke 

Leiothrix flavescens (Bong.) Ruhland 

Leiothrix flavescens var. alpina Moldenke 

Leiothrix flavescens var. flavescens 

Leiothrix mucronata var. glabra Moldenke 

Paepalanthus apacarensis Moldenke 

Paepalanthus aristatus Moldenke 

Paepalanthus auyantepuiensis Moldenke 

Paepalanthus bifidus (Schrad.) Kunth 

Paepalanthus cardonae Moldenke 

Paepalanthus chimantensis Hensold 

Paepalanthus convexus Gleason 

Paepalanthus cristatus Moldenke 

Paepalanthus cumbricola Moldenke 

Paepalanthus dichotomus Klotzsch ex Körn. 

Paepalanthus dichotomus var. dichotomus 

Paepalanthus dichotomus var. pumilus Moldenke 

Paepalanthus fasciculatus (Rottb.) Kunth 

Paepalanthus fasciculoides Hensold 

Paepalanthus formosus Moldenke 

Paepalanthus fraternus N.E. Br. 

Paepalanthus fulgidus Moldenke 

Paepalanthus gleasonii Moldenke 

Paepalanthus holstii Steyerm. 

Paepalanthus kunhardtii Moldenke 

Paepalanthus lamarckii Kunth 

Paepalanthus major (Moldenke) Hensold 

Paepalanthus oyapockensis Herzog 

Paepalanthus parvicephalus (Moldenke) Hensold 

Paepalanthus parvicephalus var. parvicephalus 

Paepalanthus parvicephalus var. wurdackii (Moldenke) Hensold 

Paepalanthus pauper Moldenke 

Paepalanthus perpusillus Kunth 

Paepalanthus phelpsiae Moldenke 

Paepalanthus polytrichoides Kunth 

Paepalanthus roraimensis Moldenke 

Paepalanthus schomburgkii Klotzsch ex Körn. 

Paepalanthus scopulorum Moldenke 

Paepalanthus sessiliflorus Körn. 

Paepalanthus sessiliflorus var. venezuelensis Moldenke 

Paepalanthus squamuliferus Moldenke 

Paepalanthus stegolepoides Moldenke 

Paepalanthus subcaulescens N.E. Br. [expected in BO] 

Paepalanthus subtilis Miq. 

Paepalanthus sulcatus Hensold 

Paepalanthus tortilis (Bong.) Körn. 

Paepalanthus turbinatus (Gleason) Hensold 

Paepalanthus venustus Moldenke 

Paepalanthus yapacanensis (Moldenke) Hensold 

Paepalanthus yapacanensis var. hirsutus (Moldenke) Hensold 

Paepalanthus yapacanensis var. yapacanensis 

Philodice hoffmannseggii Mart. 

Rondonanthus acopanensis (Moldenke) Hensold & Giul. 

Rondonanthus acopanensis var. acopanensis 

Rondonanthus acopanensis var. obtusifolius (Moldenke) Hensold & Giul. 

Rondonanthus capillaceus (Klotzsch ex Körn.) Hensold & Giul. 

Rondonanthus caulescens (Moldenke) Hensold & Giul. 

Rondonanthus duidae (Gleason) Hensold & Giul. 

Rondonanthus flabelliformis (Moldenke) Hensold & Giul. 

Rondonanthus roraimae (Oliv.) Herzog 

Syngonanthus acephalus Hensold 

Syngonanthus albopulvinatus (Moldenke) Moldenke 

Syngonanthus amapensis Moldenke [expected in the Guianas] 

AM GU 

AM 

AM BO GU 

BO 

AM BO GU 

BO 

BO 

AM 

BO 


BO 

BO 

AM GU? 

BO 

BO 

BO GU 

BO GU 

BO 


BO 

AM BO 

BO GU 

BO 

AM GU 

BO 

AM BO 


AM 


AM 

AM 

AM 

GU 

AM 

BO 

AM BO GU SU 

BO GU 

BO GU 

BO 

BO GU 

BO GU 

BO 

BO 

BO? GU 

AM BO GU SU 

AM 

AM BO GU SU 

AM 

BO 

AM 

AM 

AM 

AM BO GU SU 

BO 

BO 

BO 

AM BO GU 

BO 

AM BO 

BO 

BO GU 

AM 

AM BO 

AM

106 


Syngonanthus amazonicus Moldenke 

Syngonanthus anomalus (Körn.) Ruhland 

Syngonanthus biformis (N.E. Br.) Gleason 

Syngonanthus bisumbellatus (Steud.) Ruhland 

Syngonanthus caulescens (Poir.) Ruhland 

Syngonanthus cowanii Moldenke 

Syngonanthus duidae Moldenke 

Syngonanthus duidae var. duidae 

Syngonanthus duidae var. humilis Hensold 

Syngonanthus fenestratus Hensold 

Syngonanthus gracilis (Bong.) Ruhland [possibly =Syngonanthus davidsei 

Huft] 

Syngonanthus heteropeploides Herzog 

Syngonanthus humboldtii (Kunth) Ruhland 

Syngonanthus humboldtii var. humboldtii 

Syngonanthus humboldtii var. parvus (Moldenke) Hensold 

Syngonanthus jenmanii (Gleason) Giul. & Hensold 

Syngonanthus kegelianus (Körn.) Ruhland 

Syngonanthus leprieurii (Körn.) Ruhland 

Syngonanthus longipes Gleason 

Syngonanthus macrocephalus (Moldenke) Hensold 

Syngonanthus minutus (Moldenke) Hensold 

Syngonanthus nitens (Bong.) Ruhland 

Syngonanthus oblongus (Körn.) Ruhland 

Syngonanthus oblongus var. aequinoctialis Ruhland 

Syngonanthus ottohuberi Hensold 

Syngonanthus pakaraimensis Moldenke 

Syngonanthus pakaraimensis var. pakaraimensis 

Syngonanthus pakaraimensis var. rivularis (Moldenke) Hensold 

Syngonanthus reflexus Gleason 

Syngonanthus setifolius Hensold 

Syngonanthus simplex (Miq.) Ruhland 

Syngonanthus surinamensis Moldenke 

Syngonanthus tenuis (Kunth) Ruhland 

Syngonanthus tenuis var. bulbifer (Huber) Hensold 

Syngonanthus tenuis var. tenuis 

Syngonanthus tiricensis Moldenke 

Syngonanthus trichophyllus Moldenke 

Syngonanthus umbellatus (Lam.) Ruhland 

Syngonanthus williamsii (Moldenke) Hensold 

Syngonanthus xeranthemoides (Bong.) Ruhland 

Tonina fluviatilis Aubl. 


Carptotepala insolita Moldenke =Syngonanthus jenmanii 

Carptotepala jenmanii (Gleason) Moldenke =Syngonanthus jenmanii 

Comanthera kegeliana (Körn.) Moldenke =Syngonanthus kegelianus 

Comanthera linderi L.B. Sm. =Syngonanthus kegelianus 

Eriocaulon ? aequinoctiale Ruhland =Eriocaulon humboldtii 

Eriocaulon aquaticum Sagot ex Körn =Eriocaulon melanocephalum 

Eriocaulon atabapense Moldenke =Eriocaulon tenuifolium 

Eriocaulon bifidum Schrad. =Paepalanthus bifidus 

Eriocaulon bisumbellatum Steud. =Syngonanthus bisumbellatus 

Eriocaulon bonplandianum Steud. =Syngonanthus humboldtii 

Eriocaulon brevifolium Klotzsch ex Körn. =Eriocaulon tenuifolium 

Eriocaulon caulescens Poir. =Syngonanthus caulescens 

Eriocaulon fasciculatum Rottb. =Paepalanthus fasciculatus 

Eriocaulon fasciculatum Lam. =Paepalanthus lamarckii 

Eriocaulon flavescens Bong. =Leiothrix flavescens 

Eriocaulon gracile Bong. =Syngonanthus gracilis 

Eriocaulon klotzschii Moldenke =Eriocaulon tenuifolium 

Eriocaulon nitens Bong. =Syngonanthus nitens 

Eriocaulon sieboldianum Siebold & Zucc. ex Steud. =Eriocaulon cinereum 

Eriocaulon tenue Kunth =Syngonanthus tenuis 

AM BO 



BO GU 


AM 

AM BO 

AM BO 

BO 

BO GU 


GU? 

AM BO GU SU 

AM BO GU SU 

AM BO 

BO GU 

BO GU SU 

FG 

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AM 

BO 

AM BO 

AM BO 

AM BO 

AM 

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BO 

AM 

AM 

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AM BO 

BO 

AM BO GU 


AM 

AM BO GU 

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Eriocaulon tortile Bong. =Paepalanthus tortilis 

Eriocaulon umbellatum Kunth =Syngonanthus humboldtii 

Eriocaulon umbellatum Lam. =Syngonanthus umbellatus 

Eriocaulon xeranthemoides Bong. =Syngonanthus xeranthemoides 

Lasiolepis aquatica Böck. =Eriocaulon melanocephalum 

Leiothrix echinulata Moldenke =Syngonanthus reflexus 

Leiothrix flavescens var. chimantensis Moldenke =Leiothrix flavescens 

Leiothrix marahuacensis Moldenke =Syngonanthus duidae 

Leiothrix steyermarkii Moldenke =Syngonanthus pakaraimensis 

Leiothrix turbinata Gleason =Paepalanthus turbinatus 

Leiothrix umbratilis Moldenke =Leiothrix flavescens 

Leiothrix umbratilis var. brevipes Moldenke =Leiothrix flavescens var. alpina 

Paepalanthus albopulvinatus Moldenke =Syngonanthus albopulvinatus 

Paepalanthus anomalus Körn. =Syngonanthus anomalus 

Paepalanthus apacarensis var. humilis Moldenke =Paepalanthus dichotomus 

Paepalanthus biformis N.E. Br. =Syngonanthus biformis 

Paepalanthus brunneus Moldenke =Paepalanthus oyapockensis 

Paepalanthus bulbifer Huber =Syngonanthus tenuis var. bulbifer 

Paepalanthus capillaceus Klotzsch ex Körn. =Rondonanthus capillaceus 

Paepalanthus capillaceus f. proliferus (Gleason) Moldenke =Rondonanthus capillaceus 

Paepalanthus capillaceus var. proliferus Gleason =Rondonanthus capillaceus 

Paepalanthus caulescens (Poir.) Kunth =Syngonanthus caulescens 

Paepalanthus congestus (Kunth) Kunth =Paepalanthus fasciculatus 

Paepalanthus convexus var. major Moldenke =Paepalanthus major 

Paepalanthus convexus var. parvicephalus Moldenke =Paepalanthus parvicephalus 

Paepalanthus convexus var. strigosus Moldenke =Paepalanthus gleasonii 

Paepalanthus duidae Gleason =Rondonanthus duidae 

Paepalanthus duidae var. parvifolius Moldenke =Rondonanthus duidae 

Paepalanthus eriophyllus Körn. =Syngonanthus simplex 

Paepalanthus fertilis Körn. =Syngonanthus humboldtii 

Paepalanthus filipes Moldenke =Syngonanthus simplex 

Paepalanthus flavescens (Bong.) Körn. =Leiothrix flavescens 

Paepalanthus fraternus var. chimantensis Moldenke =Paepalanthus stegolepoides 

Paepalanthus fraternus var. marahuacensis Moldenke =Leiothrix flavescens 

Paepalanthus fraternus var. spathulatus Moldenke =Paepalanthus squamuliferus 

Paepalanthus fulgidus var. zuloagensis Moldenke =Paepalanthus major 

Paepalanthus gracilis (Bong.) Körn. =Syngonanthus gracilis 

Paepalanthus griseus Moldenke =Paepalanthus oyapockensis 

Paepalanthus guyanensis Klotzsch ex Körn. =Paepalanthus dichotomus 

Paepalanthus heteropeplus Körn. =Syngonanthus caulescens 

Paepalanthus humboldtii Kunth =Syngonanthus humboldtii 

Paepalanthus jauensis Moldenke =Rondonanthus duidae 

Paepalanthus jauensis var. caulescens Moldenke =Rondonanthus caulescens 

Paepalanthus jenmanii Gleason =Syngonanthus jenmanii 

Paepalanthus kegelianus Körn. =Syngonanthus kegelianus 

Paepalanthus killipii Moldenke =Paepalanthus tortilis 

Paepalanthus leucocyaneus Tutin =Paepalanthus oyapockensis 

Paepalanthus lilliputianus Moldenke =Paepalanthus sessiliflorus var. venezuelensis 

Paepalanthus macrocaulon var. venamensis Moldenke =Paepalanthus schomburgkii 

Paepalanthus maguirei Moldenke =Paepalanthus tortilis 

Paepalanthus meseticola Moldenke & Steyerm. =Leiothrix flavescens 

Paepalanthus minutus Moldenke =Syngonanthus minutus 

Paepalanthus moldenkeanus R. Schult. =Paepalanthus formosus 

Paepalanthus nitens (Bong.) Kunth =Syngonanthus nitens 

Paepalanthus oblongus Körn. =Syngonanthus oblongus 

Paepalanthus pauperrimus Herzog =Paepalanthus subtilis 

Paepalanthus pendulus Moldenke =Paepalanthus schomburgkii 

Paepalanthus perplexans Moldenke =Paepalanthus schomburgkii 

Paepalanthus perplexans var. steyermarkii Moldenke =Paepalanthus fulgidus 

Paepalanthus perplexans var. wurdackii Moldenke =Paepalanthus parvicephalus var. wurdackii 

Paepalanthus robustus Gleason =Paepalanthus gleasonii 

Paepalanthus roraimae Oliv. =Rondonanthus roraimae 

Paepalanthus salticola Herzog =Paepalanthus tortilis 

Paepalanthus savannarum (Moldenke) Moldenke =Paepalanthus dichotomus 

Paepalanthus savannarum var. glabrescens (Moldenke) Moldenke =Paepalanthus dichotomus

108 


Paepalanthus saxicola var. conicus Moldenke =Syngonanthus trichophyllus 

Paepalanthus scopulorum var. auyantepuiensis Moldenke =Paepalanthus scopulorum 

Paepalanthus simplex Miq. =Syngonanthus simplex 

Paepalanthus stegolepoides var. acutalis Moldenke =Paepalanthus sulcatus 

Paepalanthus steudelianus Körn. =Syngonanthus bisumbellatus 

Paepalanthus steyermarkii Moldenke =Paepalanthus dichotomus 

Paepalanthus subulatus Klotzsch =Paepalanthus subtilis 

Paepalanthus tafelbergensis Moldenke =Paepalanthus oyapockensis 

Paepalanthus tatei Moldenke =Paepalanthus tortilis 

Paepalanthus tenuis (Kunth) Kunth =Syngonanthus tenuis 

Paepalanthus umbellatus (Lam.) Kunth =Syngonanthus umbellatus 

Paepalanthus ? venetifolius Moldenke & Steyerm. =Syngonanthus pakaraimensis var. pakaraimensis 

Paepalanthus venustoides Moldenke =Paepalanthus venustus 

Paepalanthus vigiensis Moldenke =Paepalanthus polytrichoides 

Paepalanthus villipes Moldenke =Paepalanthus polytrichoides 

Paepalanthus viscosus Moldenke =Paepalanthus polytrichoides 

Paepalanthus williamsii Moldenke =Syngonanthus williamsii 

Rondonanthus micropetalus Moldenke =Paepalanthus roraimensis 

Syngonanthus acopanensis Moldenke =Rondonanthus acopanensis 

Syngonanthus akurimensis Moldenke =Syngonanthus kegelianus 

Syngonanthus allenii var. parvus Moldenke =Syngonanthus humboldtii var. parvus 

Syngonanthus baldwinii Moldenke =Syngonanthus anomalus 

Syngonanthus brevifolius Gleason =Syngonanthus anomalus 

Syngonanthus bulbifer (Huber) Ruhland =Syngonanthus tenuis var. bulbifer 

Syngonanthus caulescens var. hirsutus Moldenke =Syngonanthus caulescens 

Syngonanthus cowanii var. involucratus Moldenke =Syngonanthus cowanii 

Syngonanthus cowanii var. longipedunculatus Moldenke =Syngonanthus cowanii 

Syngonanthus cowanii var. simplex Moldenke =Syngonanthus cowanii 

Syngonanthus cowanii var. tabulatus Moldenke =Syngonanthus cowanii 

Syngonanthus densifolius var. venezuelensis Moldenke =Syngonanthus ottohuberi 

Syngonanthus densus var. pumilus Moldenke =Syngonanthus biformis 

Syngonanthus drouetii L.B. Sm. =Syngonanthus tenuis var. bulbifer 

Syngonanthus drouetii var. parviceps Moldenke =Syngonanthus tenuis 

Syngonanthus duidae var. longifolius Moldenke =Syngonanthus pakaraimensis 

Syngonanthus eriophyllus (Körn.) Ruhland =Syngonanthus simplex 

Syngonanthus fertilis (Körn.) Ruhland =Syngonanthus humboldtii 

Syngonanthus fertilis var. glandulosus (Gleason) Moldenke =Syngonanthus humboldtii 

Syngonanthus fertilis var. orinocensis (Moldenke) Moldenke =Syngonanthus humboldtii 

Syngonanthus flavipes Moldenke =Syngonanthus nitens 

Syngonanthus glandulosus Gleason =Syngonanthus caulescens 

Syngonanthus glandulosus var. epapillosus Moldenke =Syngonanthus caulescens 

Syngonanthus glandulosus f. epapillosus (Moldenke) Moldenke =Syngonanthus caulescens 

Syngonanthus guianensis Moldenke =Paepalanthus polytrichoides 

Syngonanthus heteropeplus (Körn.) Ruhland =Syngonanthus caulescens 

Syngonanthus huberi Ruhland =Syngonanthus anomalus 

Syngonanthus humboldtii var. elongatus Moldenke =Syngonanthus humboldtii 

Syngonanthus humboldtii var. glabrescens Moldenke =Syngonanthus humboldtii 

Syngonanthus humboldtii var. glandulosus Gleason =Syngonanthus humboldtii 

Syngonanthus humboldtii var. macrocephalus Moldenke =Syngonanthus macrocephalus 

Syngonanthus humboldtii var. orinocensis Moldenke =Syngonanthus humboldtii 

Syngonanthus humboldtii var. simplex Moldenke =Syngonanthus umbellatus 

Syngonanthus lanatus var. alpinus Moldenke =Syngonanthus pakaraimensis 

Syngonanthus macrocaulon Ruhland =Syngonanthus anomalus 

Syngonanthus obtusifolius Moldenke =Rondonanthus acopanensis var. obtusifolius 

Syngonanthus paraensis Ruhland =Syngonanthus anomalus 

Syngonanthus phelpsiae Moldenke =Syngonanthus duidae 

Syngonanthus phelpsiae var. elongatus Moldenke =Syngonanthus duidae 

Syngonanthus phelpsiae var. pilosus Moldenke =Rondonanthus duidae 

Syngonanthus phelpsiae var. viridis Moldenke =Rondonanthus duidae 

Syngonanthus reflexus var. longifolius Moldenke =Syngonanthus reflexus 

Syngonanthus rivularis Moldenke =Syngonanthus pakaraimensis var. rivularis 

Syngonanthus savannarum Moldenke =Paepalanthus dichotomus 

Syngonanthus savannarum var. glabrescens Moldenke =Paepalanthus dichotomus 

Syngonanthus similis var. venezuelensis Moldenke =Syngonanthus ottohuberi 

Syngonanthus simplex var. appendiculifera Ruhland =Syngonanthus biformis


Syngonanthus ? tenuis var. minor Moldenke =Syngonanthus tenuis 

Syngonanthus tricostatus Gleason =Syngonanthus xeranthemoides 

Syngonanthus umbellatus f. proliferens Moldenke =Syngonanthus umbellatus 

Syngonanthus vareschii Moldenke =Syngonanthus simplex 

Syngonanthus vaupesanus Moldenke =Syngonanthus tenuis var. bulbifer 

Syngonanthus venezuelensis Moldenke =Paepalanthus dichotomus 

Syngonanthus xeranthemoides var. alpinus Moldenke =Syngonanthus xeranthemoides 

Syngonanthus xeranthemoides var. angustifolius Moldenke =Syngonanthus xeranthemoides 

Syngonanthus xeranthemoides f. brevifolius Moldenke =Syngonanthus xeranthemoides 

Syngonanthus xeranthemoides var. tricostatus (Gleason) Moldenke =Syngonanthus xeranthemoides 

Syngonanthus yapacanensis Moldenke =Paepalanthus yapacanensis 

Syngonanthus yapacanensis var. hirsutus Moldenke =Paepalanthus yapacanensis var. hirsutus 

Wurdackia flabelliformis Moldenke =Rondonanthus flabelliformis 

REFERENCES 

Giulietti, A.M., M.C.E. Amaral, & V. Bittrich. 1995. Phylogenetic analysis of inter- and infrageneric relationships of Leiothrix 

Ruhland (Eriocaulaceae). Kew Bull. 50(1): 55-71. 

Giulietti, A.M. & N. Hensold. 1991. Nomenclatural changes and range extension in Leiothrix flavescens (Bong.) Ruhl. 

(Eriocaulaceae). Novon 1(1): 45-49. 

Giulietti, A.M. & N. Hensold. 1991. Synonymization of the genera Comanthera and Carptotepala with Syngonanthus 

(Eriocaulaceae). Ann. Missouri Bot. Gard. 78 (2): 460-464. 

Hensold, N. 1991. Revisionary studies in the Eriocaulaceae of Venezuela. Ann. Missouri Bot. Gard. 78(2): 424-440. 

Hensold, N. 1999. Eriocaulaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 1-58. 

Hensold, N. & A.M. Giulietti. 1991. Revision and redefinition of the genus Rondonanthus Herzog (Eriocaulaceae). Ann. Missouri 

Bot. Gard. 78(2): 441-459. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Eriocaulaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 330-339. 

Moldenke, H.N. 1971. A Fifth Summary of the Verbenaceae, Avicenniaceae, Stilbaceae, Dicrostylidaceae, Symphoremaceae, 

Nyctanthaceae, and Eriocaulaceae of the World. H.N. Moldenke, Wayne, NJ. 

Mori, S.A. 1997. Eriocaulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 


of the New York Botanical Garden 76, pt. 1: 266. 

Uittien, H. & A.N.J. Heyn. 1938. Eriocaulaceae. In A. Pulle, Flora of Suriname 1(1): 213-224. 

Pyrrorhiza neblinae Maguire & Wurdack 

Schiekia orinocensis (Kunth) Meisn. 

Schiekia orinocensis subsp. orinocensis 

Schiekia orinocensis subsp. silvestris Maas & Stoel 

Xiphidium caeruleum Aubl. 

HAEMODORACEAE 

P.J.M. Maas, Hiltje Maas 

AM 


AM BO GU SU 




Schiekia flavescens Maury =Schiekia orinocensis subsp. orinocensis 

Schiekia orinocensis subsp. savannarum Maguire & Wurdack =Schiekia orinocensis subsp. orinocensis 

Troschelia orinocensis (Kunth) Klotzsch & M.R. Schomb. [nomen nudum] =Schiekia orinocensis subsp. orinocensis 

Wachendorfia orinocensis Kunth =Schiekia orinocensis subsp. orinocensis 

Xiphidium floribundum Sw. =Xiphidium caeruleum 

Xiphidium fockeanum Miq. =Xiphidium caeruleum 

Xiphidium giganteum Lindl. =Xiphidium caeruleum 

REFERENCES 

Maas-van de Kamer, H. & P. Maas. 1997. Haemodoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & 


Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 270. 

Lanjouw, J. 1938. Haemodoraceae. In A. Pulle, Flora of Suriname 1(1): 255-257. 

Maas, P.J.M. & H. Maas-van de Kamer. 1993. Haemodoraceae. Flora Neotropica Monogr. 61: 1-44. 

Maas, P.J.M. & H. Maas-van de Kamer. 1994. Haemodoraceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 15: 

103-112. 

Maas, P.J.M. & H. Maas. 1999. Haemodoraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Villarroel, F. 1971. Haemodoraceae. In T. Lasser, Flora de Venezuela 3(1): 207-222.

110 


HELICONIACEAE 

L.L. Andersson, W.J. Kress, P.J.M. Maas 

Heliconia acuminata Rich. 

Heliconia acuminata subsp. acuminata 

Heliconia acuminata subsp. occidentalis L. Andersson 

Heliconia aemygdiana Burle-Marx [CAY specimen not seen] 

Heliconia bihai (L.) L. 

Heliconia chartacea Lane ex Barreiros 

Heliconia dasyantha K. Koch & Bouché 

Heliconia densiflora B. Verl. 

Heliconia densiflora subsp. angustifolia L. Andersson 

Heliconia densiflora subsp. densiflora 

Heliconia episcopalis Vell. 

Heliconia hirsuta L. f. 

Heliconia juliani Barreiros 

Heliconia lourteigiae Emygdio & E. Santos 

Heliconia marginata (Griggs) Pittier 

Heliconia x nickeriensis Maas & de Rooij 

Heliconia pendula Wawra 

Heliconia platystachys Baker 

Heliconia psittacorum L. f. 

Heliconia revoluta (Griggs) Standl. [Sierra de la Neblina] 

Heliconia richardiana Miq. 

Heliconia spathocircinata Aristeg. [orthographic variant: H. spathocircinada] 

Heliconia stricta Huber 


Bihai marginata Griggs =Heliconia marginata 

Bihai revoluta Griggs =Heliconia revoluta 

Bihai silvestris Gleason =Heliconia psittacorum 

Heliconia aurea G. Rodr. =Heliconia bihai 

Heliconia ballia Rich. =Heliconia psittacorum 

Heliconia brasiliensis var. concolor Petersen =Heliconia psittacorum 

Heliconia cannoidea Rich. =Heliconia psittacorum 

Heliconia caribaea auct. non Lam. 1783 [1785], nec Aristeg. 1961 =Heliconia bihai 

Heliconia costanensis Aristeg. =Heliconia hirsuta 

Heliconia glauca Poit. ex B. Verl. =Heliconia richardiana 

Heliconia hirsuta var. cannoidea (Rich.) Baker =Heliconia psittacorum 

Heliconia humilis Aubl., auct. non Musa humilis =Heliconia bihai 

Heliconia humilis (Aubl.) Jacq. =Heliconia psittacorum 

Heliconia marginata f. lutea Aristeg. =Heliconia marginata 

Heliconia psittacorum var. rhizomatosa Aristeg. =Heliconia psittacorum 

Heliconia psittacorum var. schomburgkiana (Klotzsch) Baker =Heliconia psittacorum 

Heliconia psittacorum var. spathacea Eichler ex Petersen =Heliconia psittacorum 

Heliconia schaeferiana G. Rodr. =Heliconia bihai 

Heliconia schomburgkiana Klotzsch =Heliconia psittacorum 

Heliconia silvestris (Gleason) L.B. Sm. =Heliconia psittacorum 

REFERENCES 

Anderson, P., S.A. Mori, & C. Gracie. 1997. Heliconiaceae. In: Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & 



Andersson, L.L. 1999. Heliconiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 5: 583-591. 

Berry, F. & W.J. Kress. 1991. Heliconia: An Identification Guide. Washington, DC: Smithsonian Institution Press. 

Maas, P.J.M. 1985. Musaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 1: 1-28. 

Maas, P.J.M. & M.J.M. de Rooij. 1979. Musaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 393-411. 

Elodea granatensis Bonpl. 

Limnobium laevigatum (Humb. & Bonpl. ex Willd.) Heine 

HYDROCHARITACEAE 




AM 

FG? 



SU FG 

BO GU SU FG 

BO GU 

BO GU SU FG 

AM SU 


AM 

BO GU SU FG 

DA GU SU 

GU SU 

GU FG 

BO 


AM 



GU SU 


DA GU SU FG



Apalanthe granatensis (Bonpl.) Planch. =Elodea granatensis 

Elodea guianensis Rich. =Elodea granatensis 

Elodea orinocensis Rich. =Elodea granatensis 

Hydromystria laevigata (Willd.) Hanzeker =Limnobium laevigatum 

Hydromystria laevigata (Humb. & Bonpl. ex Willd.) Díaz-Mir. & Philcox =Limnobium laevigatum 

Limnobium spongia subsp. laevigatum (Humb. & Bonpl. ex Willd.) Lowden =Limnobium laevigatum 

Limnobium stoloniferum (G. Mey.) Griseb. =Limnobium laevigatum 

Philotria granatensis (Bonpl.) Vict. =Elodea granatensis 

Philotria orinocensis (Rich.) Vict. =Elodea granatensis 

Udora granatensis (Bonpl.) Spreng. =Elodea granatensis 

REFERENCES: 

Cook, C.D. 1985. A revision of the genus Apalanthe. Aquatic Bot. 21: 157-164. 

Cook, C.D. & K. Urmi-König, 1983. A revision of the genus Limnobium including Hydromystria. Aquatic Bot. 17: 1-27. 

Haynes, R.R. & L.B. Holm-Nielsen. 1999. Hydrocharitaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 

Flora of the Venezuelan Guayana 5: 641-644. 

Jonker, F.P. 1943. Hydrocharitaceae. In A. Pulle, Flora of Suriname 1(1): 467-471. 

Lowden, R.M. 1992. Floral variation and taxonomy of Limnobium L.C. Richard. Rhodora 94: 111-134. 

Phillips, R.C. & E.G. Meñez. 1988. Seagrasses. Smiths. Contrib. Marine Sci. 34: 1-104. 

IRIDACEAE 

P. Goldblatt, J.E. Henrich 

Alophia drummondii (Graham) R.C. Foster 

Cipura gigas Celis, Goldblatt & Betancur 

Cipura paludosa Aubl. 

Cipura rupicola Goldblatt & Henrich 

Cypella linearis (Kunth) Baker 

Eleutherine bulbosa (Mill.) Urb. 

Neomarica caerulea (Ker Gawl.) Sprague [escaped from cultivation] 

Sisyrinchium tinctorium Kunth 

Sisyrinchium vaginatum Spreng. [expected in FG] 

Trimezia chimantensis Steyerm. 

Trimezia fosteriana Steyerm. 

Trimezia guianensis Ravenna 

Trimezia martinicensis (Jacq.) Herb. 


Iris martinicencis Jacq. =Trimezia martinicensis 

Ixia americana Aubl. =Eleutherine bulbosa 

Ixia xiphidium Loefl. (Haemodoraceae) =Xiphidium caeruleum 

Larentia linearis (Kunth) Klatt =Cypella linearis 

Moraea linearis Kunth =Cypella linearis 

Sisyrinchium alatum Hook. =Sisyrinchium vaginatum 

Sisyrinchium americanum (Aubl.) Lemée =Eleutherine bulbosa 

Sisyrinchium marchio (Vell.) Steud. =Sisyrinchium vaginatum 

Souza marchio Vell. =Sisyrinchium vaginatum 

REFERENCES: 

Boterenbrood, M.J.A. 1943. Iridaceae. In A. Pulle, Flora of Suriname 1(1): 455-460. 

Goldblatt, P. & James E. Henrich. 1999. Iridaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Goldblatt, P. & T.M. Howard. 1992. Notes on Alophia and a new species, A. veracruzana, from Vera Cruz, Mexico. Ann. Missouri 

Bot. Gard. 79(4): 901-905. 

Goldblatt, P. & N. Snow. 1991. Systematics and chromosome cytology of Eleutherine Herbert. Ann. Missouri Bot. Gard. 78(4): 

942-949. 

Mori, S.A. 1997. Iridaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide 



Juncus densiflorus Kunth 

JUNCACEAE 

C.A. Vargas 

GU 

AM 

AM 

AM 

BO GU SU FG 

AM BO 

GU SU FG 

AM 

AM BO GU SU 

BO 

BO GU 

GU 

BO GU? FG 

AM BO

112 



Juncus cyperinus Willd. ex Spreng. =Juncus densiflorus 

Juncus densiflorus var. cyperinus (Willd.) Buchenau =Juncus densiflorus 

REFERENCES 

Vargas, C.A. 1999. Juncaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 672-674. 

Lemna aequinoctialis Welw. 

Lemna valdiviana Phil. 

Spirodela intermedia W. Koch 

Spirodela punctata (G. Mey.) C.H. Thomps. 

Wolffia brasiliensis Wedd. 

Wolffiella lingulata (Hegelm.) Hegelm. 

Wolffiella neotropica Landolt 

Wolffiella welwitschii (Hegelm.) Monod 

LEMNACEAE 

L.J. Davenport, R.A. DeFilipps† 


Lemna gibba L. [not in flora area] 

Lemna minor L. [not in flora area] 

Lemna paucicostata Hegelm. =Lemna aequinoctialis 

Lemna perpusilla Torr. [not in flora area] 

Lemna trinervis sensu Small, non (D.F. Austin) Small =Lemna aequinoctialis 

Spirodela biperforata W. Koch =Spirodela intermedia 

Spirodela polyrhiza (L.) Schleid. [not in flora area] 

Wolffia welwitschii Hegelm. =Wolffiella welwitschii 

Wolffiopsis welwitschii (Hegelm.) Hartog & Plas =Wolffiella welwitschii 

REFERENCES 

Daubs, E.H. 1965. A monograph of Lemnaceae. Ill. Biol. Monogr. 34: 1-118. 

Davenport, L.J. 1999. Lemnaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 779-781. 

Hartog, C. den. & F. van der Plas. 1970. A synopsis of the Lemnaceae. Blumea 18: 355-368. 

Landolt, E. 1986-1987. The family of Lemnaceae - a monographic study. Veröff. Geobot. Inst. E.T.H. Stift. Rübel, Zürich 71: 1- 

566, 95: 1-638. 

LILIACEAE 

J.K. Boggan, R.W. Cruden, A.W. Meerow 

Alstroemeria amazonica Ducke 

Alstroemeria sp. 

Bomarea edulis (Tussac) Herb. 

Crinum erubescens L. f. ex Sol. 

Curculigo scorzonerifolia (Lam.) Baker 

Eccremis coarctata (Ruiz & Pav.) Baker 

Echeandia bolivarensis Cruden 

Hippeastrum barbatum Herbert 

Hippeastrum elegans (Spreng.) H.E. Moore 

Hippeastrum puniceum (Lam.) Kuntze 

Hippeastrum stylosum Herbert [distribution described as “Guiana”] 

Hippeastrum vanleestenii (Traub) H.E. Moore [=Hippeastrum puniceum?] 

Hymenocallis caribaea (L., emend. Ker Gawl.) Herb. [cultivated or escaped 

from cultivation] 

Hymenocallis littoralis (Jacq.) Salisb. [cultivated or escaped from cultivation] 

Hymenocallis tubiflora Salisb. 

Hypoxis decumbens L. [most, if not all, records represent Curculigo 

scorzonerifolia] 

Isidrogalvia duidae (Steyerm.) Cruden 

Isidrogalvia schomburgkiana (Oliv.) Cruden 

Nietneria corymbosa Klotzsch & M.R. Schomb. ex B.D. Jacks. [Mt. Roraima] 

Nietneria paniculata Steyerm. 

DA GU SU FG 

SU 

DA SU 

GU 

BO SU 

SU FG 

SU 


BO 

FG 

AM BO GU SU 



AM BO 

BO 

SU 

AM BO GU SU 

AM GU SU FG 

SU 

GU SU FG 

GU? SU FG 


GU? SU? FG? 

AM 

AM BO GU 

AM BO GU 

BO GU


Zephyranthes citrina Baker [type attributed to Guyana, possibly erroneously] 

Zephyranthes tubispatha (Ker Gawl.) Herb. 


Alstroemeria edulis Tussac =Bomarea edulis 

Amaryllis belladonna L. [name incorrectly applied to Hippeastrum puniceum] 

Amaryllis belladonna var. barbata (Herb.) Traub & Moldenke =Hippeastrum barbatum 

Amaryllis eguestris Aiton =Hippeastrum puniceum 

Amaryllis elegans Spreng. =Hippeastrum elegans 

Amaryllis longiflora Steud. =Hippeastrum elegans 

Amaryllis punicea Lam. =Hippeastrum puniceum 

Amaryllis solandriflora Lindl. =Hippeastrum elegans 

Amaryllis stylosa (Herb.) Sweet =Hippeastrum stylosum 

Amaryllis vanleestenii Traub =Hippeastrum vanleestenii 

Anthericum coarctatum Ruiz & Pav. =Eccremis coarctata 

Bomarea fuscata Klotzsch ex M.R. Schomb. [nomen nudum] 

Bomarea guianensis Kraenzl. =Bomarea edulis 

Crinum commelyni Jacq. =Crinum erubescens 

Crinum guianense M. Roem. ? =Crinum erubescens 

Crinum lancei Herbert ex Sweet ? =Crinum erubescens 

Crinum lindleyanum Schult. f. ex Seub. =Crinum erubescens 

Dianella dubia Kunth =Eccremis coarctata 

Echeandia spp. [recorded from Guianas in error, but possibly expected] 

Hippeastrum equestre (Aiton) Herb. =Hippeastrum puniceum 

Hippeastrum puniceum (Lam.) Urb. =Hippeastrum puniceum 

Hippeastrum puniceum (Lam.) Voss =Hippeastrum puniceum 

Hippeastrum solandriflorum (Lindl.) Herb. =Hippeastrum elegans 

Hymenocallis guianensis (Ker Gawl.) Herb. =Hymenocallis tubiflora 

Hymenocallis guianensis var. undulata (Kunth) Herb. =Hymenocallis tubiflora 

Hymenocallis moritziana Kunth =Hymenocallis tubiflora 

Hymenocallis obtusata (Griseb.) Walp. =Hymenocallis caribaea 

Hymenocallis petiolata (Willd. ex Schult.) M. Roem. =Hymenocallis tubiflora 

Hymenocallis undulata (Kunth) Herb. =Hymenocallis tubiflora 

Hypoxis scorzonerifolia Lam. =Curculigo scorzonerifolia 

Isidrogalvia guianensis Klotzsch [nomen nudum] =Isidrogalvia schomburgkiana 

Nietneria corymbosa Klotzsch & M.R. Schomb. [nomen nudum] =Nietneria corymbosa 

Pancratium guianense Ker Gawl. =Hymenocallis tubiflora 

Pancratium petiolatum Willd. ex Schult. =Hymenocallis tubiflora 

Pancratium tubiflorum (Salisb.) Schult. =Hymenocallis tubiflora 

Pancratium undulatum Kunth =Hymenocallis tubiflora 

Tofieldia duidae Steyerm. =Isidrogalvia duidae 

Tofieldia guianensis (Klotzsch) H. Schulz [nomen nudum] =Isidrogalvia schomburgkiana 

Tofieldia rupestris Gleason (Bromeliaceae) =Navia rupestris 

Tofieldia schomburgkiana Oliv. =Isidrogalvia schomburgkiana 

REFERENCES 

Baker, J.G. 1888. Handbook of the Amaryllideae. London: George Bell & Sons. 

Bloomfield, F. 1985. Miracle Plants: Aloe vera. London: Century Publishing. 

Boterenbrood, M.J.A. 1943. Amaryllidaceae. In A. Pulle, Flora of Suriname 1(1): 443-454. 

Cruden, R.W. 1991. A revision of Isidrogalvia: recognition for Ruíz & Pavón’s genus. Syst. Bot. 16: 270-282. 

Howard, R.A. 1979. Liliaceae. Flora of the Lesser Antilles 3: 454-460. 

Killip, E.P. 1936. Bomarea, a genus of showy Andean plants. Nat. Hort. Mag. 15(2): 115-128. 

Maas, P.J.M. 1968. Amaryllidaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 376. 

Martyn, E.B. 1936. Amaryllidaceae of British Guiana. Herbertia 3: 71. 

Meerow, A.W. & R.W. Cruden. 2001. Liliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Mori. S.A. 1997. Liliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. Guide to the 

vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the New 

York Botanical Garden 76, pt. 1. 

Obermeyer, A.A. 1962. A revision of the South African species of Anthericum, Chlorophytum and Trachyandra. Bothalia 7 (4): 

669-767. 

Sealy, J.R. 1954. Review of the genus Hymenocallis. Kew Bull. 1954: 201-240. 

Traub, H.P. 1958. The Amaryllis Manual. The Macmillan Company, New York. 

Traub, H.P. & H.N. Moldenke. 1949. Amaryllidaceae: Tribe Amarylleae. American Plant Life Society. 

Uphof, J.C. Th. 1942. A review of the species of Crinum. Herbertia 9: 63-84. 

GU? 

SU

114 


LIMNOCHARITACEAE 


Hydrocleys nymphoides (Humb. & Bonpl. ex Willd.) Buchenau 

Limnocharis flava (L.) Buchenau [record not verified, but expected 

in the Guianas] 

Limnocharis laforestii Duchass. ex Griseb. 


Limnocharis flava var. minor Micheli =Limnocharis laforestii 

Stratiotes nymphoides Humb. & Bonpl. ex Willd. =Hydrocleys nymphoides 

REFERENCES 

Haynes, R.R. & L.B. Holm-Nielsen. 2001. Limnocharitaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Haynes, R.R. & L.B. Holm-Nielsen. 1992. Limnocharitaceae. Flora Neotropica Monogr. 56: 1-34. 

Jonker, F.P. 1943. Butomaceae. In A. Pulle, Flora of Suriname 1(1): 483-485. 

MARANTACEAE 

L.L. Andersson, Mats Hagberg, H. Kennedy 

Calathea acuminata Steyerm. 

Calathea altissima (Poepp. & Endl.) Körn. 

Calathea cannoides (Nicolson, Steyerm. & Sivad.) H. Kenn. 

Calathea casupito (Jacq.) Schult. 

Calathea cataractarum K. Schum. 

Calathea comosa (L. f.) Lindl. 

Calathea cyclophora Baker 

Calathea cylindrica (Roscoe) K. Schum. 

Calathea densa (K. Koch) Regel 

Calathea dilabens L. Andersson & H. Kenn. 

Calathea elliptica (Roscoe) K. Schum. 

Calathea erecta L. Andersson & H. Kenn. 

Calathea fragilis Gleason 

Calathea grandis Petersen 

Calathea granvillei L. Andersson & H. Kenn. 

Calathea inocephala (Kuntze) H. Kenn. & Nicolson 

Calathea lasseriana Steyerm. 

Calathea latifolia (Willd. ex Link) Klotzsch 

Calathea legrelleana (Linden) Regel 

Calathea liesneri H. Kenn. 

Calathea lutea (Aubl.) Schult. 

Calathea maasiorum H. Kenn. 

Calathea mansonis Körn. 

Calathea micans (Mathieu) Körn. 

Calathea microcephala (Poepp. & Endl.) Körn. 

Calathea mishuyacu J.F. Macbr. 

Calathea neblinensis H. Kenn. 

Calathea ovata (Nees & Mart.) Lindl. 

Calathea panamensis Rowlee ex Standl. 

Calathea polytricha Baker 

Calathea propinqua (Poepp. & Endl.) Körn. 

Calathea splendida (Verschaff. ex Lem.) Regel 

Calathea squarrosa L. Andersson & H. Kenn. 

Calathea variegata Linden ex Körn. 

Calathea villosa Lindl. 

Calathea zingiberina Körn. 

Ctenanthe compressa (A. Dietr.) Eichler 

Hylaeanthe hexantha (Poepp. & Endl.) A.M.E. Jonker & Jonker 

Hylaeanthe polystachya (Pulle) A.M.E. Jonker & Jonker 

Hylaeanthe unilateralis (Poepp. & Endl.) A.M.E. Jonker & Jonker 

Ischnosiphon arouma (Aubl.) Körn. 

Ischnosiphon bambusaceus (Poepp. & Endl.) Körn. 

Ischnosiphon cannoideus L. Andersson 

BO DA GU SU FG? 

FG? 

BO 

AM 

AM GU SU FG 

BO 

AM BO GU 

AM GU 

SU FG? 

AM BO DA GU 

FG 

AM SU 

FG 


AM FG 

AM BO GU SU 

BO GU SU FG 

GU SU FG 

AM BO DA 

AM BO 

AM BO 

GU SU FG 

AM 


SU FG 

GU SU FG 


FG 

BO 

AM 

AM SU? FG? 

AM BO 

GU 

GU SU FG 

FG 

SU FG 

BO GU SU 



AM BO DA 

SU FG 

SU FG? 



SU? FG? 

AM

Ischnosiphon centricifolius L. Andersson 

Ischnosiphon enigmaticus L. Andersson 

Ischnosiphon foliosus Gleason 

Ischnosiphon gracilis (Rudge) Körn. 

Ischnosiphon gracilis subsp. gracilis 

Ischnosiphon gracilis subsp. sandwithii L. Andersson 

Ischnosiphon hirsutus Petersen 

Ischnosiphon leucophaeus (Poepp. & Endl.) Körn. 

Ischnosiphon leucophaeus subsp. leucophaeus 

Ischnosiphon longiflorus K. Schum. 

Ischnosiphon longiflorus subsp. angustifolius L. Andersson 

Ischnosiphon martianus Eichler ex Petersen 

Ischnosiphon obliquus (Rudge) Körn. 

Ischnosiphon petiolatus (Rudge) L. Andersson 

Ischnosiphon polyphyllus (Poepp. & Endl.) Körn. 

Ischnosiphon puberulus Loes. 

Ischnosiphon surumuensis Loes. 

Ischnosiphon ursinus L. Andersson 

Koernickanthe orbiculata (Körn.) L. Andersson 

Maranta amplifolia K. Schum. 

Maranta arundinacea L. 

Maranta friedrichsthaliana Körn. 

Maranta gibba Sm. 

Maranta humilis Aubl. 

Maranta linearis L. Andersson 

Maranta protracta Miq. 

Maranta ruiziana Körn. 

Maranta rupicola L. Andersson 

Monophyllanthe oligophylla K. Schum. 

Monotagma contractum Huber 

Monotagma juruanum Loes. 

Monotagma laxum (Poepp. & Endl.) K. Schum. 

Monotagma ovatum Hagberg 

Monotagma plurispicatum (Körn.) K. Schum. 

Monotagma rhodanthum Maguire & Wurdack 

Monotagma secundum (Petersen) K. Schum. 

Monotagma sp. nov. ined. 

Monotagma spicatum (Aubl.) J.F. Macbr. 

Monotagma tomentosum K. Schum. ex Loes. 

Monotagma ulei K. Schum. ex Loes. 

Monotagma vaginatum Hagberg 

Monotagma yapacanensis Steyerm. & G.S. Bunting 

Myrosma cannifolia L. f. 

Stromanthe jacquinii (Roem. & Schult.) H. Kenn. & Nicolson 

Stromanthe tonckat (Aubl.) Eichler 

Thalia geniculata L. 



Calathea albicans Brongn. ex K. Schum. =Calathea micans 

Calathea allouia (Aubl.) Lindl. [not in flora area] =Calathea legrelleana 

Calathea allouia var. violacea auct. non Calathea violacea Lindl. 1825 =Calathea latifolia 

Calathea altissima auct. non (Poepp. & Endl.) Körn. 1862 =Calathea inocephala 

Calathea barbillana Cufod. =Calathea inocephala 

Calathea discolor G. Mey. =Calathea lutea 

Calathea duidae Steyerm. =Calathea fragilis 

Calathea hirsuta Standl. =Calathea villosa 

Calathea klugii J.F. Macbr. =Calathea micans 

Calathea laxa Poepp. & Endl. =Monotagma laxum 

Calathea leucophaea Poepp. & Endl. =Ischnosiphon leucophaeus 

Calathea magnifica C.V. Morton & Skutch =Calathea lutea 

Calathea marantina (Willd.) K. Koch =Calathea casupito 

Calathea micans var. amabilis (Linden) Petersen =Calathea micans 

Calathea pardina Planch. & Linden =Calathea villosa 

Calathea polyphylla Poepp. & Endl. =Ischnosiphon polyphyllus 

GU SU FG 

BO GU FG 

GU 



GU 

BO GU 



AM GU SU 

AM GU SU 

SU FG 


GU SU FG 

AM 


AM 

AM SU FG 

SU 

BO GU SU FG 


FG 

BO DA GU SU 


AM BO 

BO GU SU 

AM GU SU FG 

BO GU SU FG 

SU FG 

SU FG 

SU FG 


BO GU 


AM 

AM BO GU SU 

FG 


AM BO 

FG 

AM FG 

AM 


BO 

BO GU SU FG 

BO DA GU SU FG

116 


Calathea strobilifera (Miq.) Körn. (Zingiberaceae) =Renealmia cernua 

Calathea trinitensis Britton =Calathea casupito 

Calathea villosa var. pardina (Planch. & Linden) Körn. =Calathea villosa 

Calathea vittata hort. ex Körn. =Calathea elliptica 

Ischnosiphon gracilis var. scabra Petersen =Ischnosiphon puberulus 

Ischnosiphon lasseriana Steyerm. =Ischnosiphon gracilis 

Ischnosiphon plurispicatus Körn. =Monotagma plurispicatum 

Ischnosiphon puberulus var. scaber (Petersen) L. Andersson =Ischnosiphon puberulus 

Ischnosiphon puberulus var. verruculosus (J.F. Macbr.) L. Andersson =Ischnosiphon puberulus 

Ischnosiphon secundus Petersen =Monotagma secundum 

Ischnosiphon surinamensis (Miq.) Körn. =Ischnosiphon leucophaeus 

Ischnosiphon verruculosous J.F. Macbr. =Ischnosiphon puberulus 

Ischnosiphon violaceus Pulle =Ischnosiphon petiolatus 

Maranta allouia Aubl. =Calathea legrelleana 

Maranta amabilis Linden =Calathea micans 

Maranta arouma Aubl. =Ischnosiphon arouma 

Maranta cachibou Jacq. =Calathea lutea 

Maranta casupito Jacq. =Calathea casupito 

Maranta compressa A. Dietr. =Ctenanthe compressa 

Maranta divaricata K. Schum. 1902 auct. non Roscoe 1828, A.M.E. Jonker & Jonker1957 =Maranta gibba 

Maranta divaricata Roscoe [not in flora area] 

Maranta gracilis Rudge =Ischnosiphon gracilis 

Maranta jacquinii Roem. & Schult. =Stromanthe jacquinii 

Maranta lutea Aubl. =Calathea lutea 

Maranta lutea Jacq. =Stromanthe jacquinii 

Maranta micans Mathieu =Calathea micans 

Maranta obliqua Rudge =Ischnosiphon obliquus 

Maranta orbiculata (Körn.) K. Schum. =Koernickanthe orbiculata 

Maranta spicata Aubl. =Monotagma spicatum 

Maranta strobilifera Miq. (Zingiberaceae) =Renealmia cernua 

Maranta tonckat Aubl. =Stromanthe tonckat 

Monotagma duidae Steyerm. =Monotagma tomentosum 

Monotagma exannulatum K. Schum. =Monotagma laxum 

Monotagma guianense (Körn.) K. Schum. =Monotagma plurispicatum 

Monotagma parkeri (Roscoe) K. Schum. =Monotagma spicatum 

Monotagma surinamense Pulle =Monotagma secundum 

Myrosma polystachya Pulle =Hylaeanthe polystachya 

Myrosma unilateralis (Poepp. & Endl.) K. Schum. =Hylaeanthe unilateralis 

Phrynium altissimum Poepp. & Endl. =Calathea altissima 

Phrynium densum K. Koch =Calathea densa 

Phrynium ellipticum Roscoe =Calathea elliptica 

Phrynium micans (Mathieu) Klotzsch =Calathea micans 

Phrynium ovatum Nees & Mart. =Calathea ovata 

Phrynium pusillum K. Koch =Calathea micans 

Phrynium strobiliferum (Miq.) K. Koch (Zingiberaceae) =Renealmia cernua 

Phyllodes inocelphalum Kuntze =Calathea inocephala 

Phyllodes platyphyllum Kuntze =Calathea latifolia 

Saranthe urceolata Petersen =Maranta humilis 

Stromanthe lutea Eichler =Stromanthe jacquinii 

Thalia latifolia (Willd. ex Link) Link ex Schult. =Calathea latifolia 

Thalia trichocalyx Gagnep. =Thalia geniculata 

Thalia unilateralis Poepp. & Endl. =Hylaeanthe unilateralis 

Thymocarpus cannoides Nicolson, Steyerm. & Sivad. =Calathea cannoides 

REFERENCES 

Andersson, L. 1977. The genus Ischnosiphon. Opera Bot. 43: 1-113. 

Andersson, L. 1986. Revision of Maranta subgen. Maranta. Nord. J. Bot. 6(6): 729-756. 

Andersson, L. 1997. Marantaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Andersson, L. & H. Kennedy. 1986. Four new species of Calathea from French Guiana. Nordic J. Bot. 6(4): 447-455. 

Andersson, L., H. Kennedy & M. Hagberg. Marantaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 

the Venezuelan Guayana 6: 219-248.


Hagberg, M. 1990. Two new species of Monotagma from the Venezuelan Guayana. Ann. Missouri Bot. Gard. 77(2): 421-424. 

Jonker-Verhoef, A.M.E. & F.P. Jonker. 1957. Marantaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 149-208. 

Kennedy, H. 1995. Calathea maasiorum (Marantaceae), a new species from French Guiana and Surinam. Brittonia 47(2): 156- 

159. 

Mayaca fluviatilis Aubl. 

Mayaca longipes Mart. ex Seub. 

Mayaca sellowiana Kunth 

Mayaca aubletii Michx. =Mayaca fluviatilis 

MAYACACEAE 

A. Lourteig† 


REFERENCES 

Lanjouw, J. 1938. Mayacaceae. In A. Pulle, Flora of Suriname 1(1): 249-251. 

Lourteig, A. 1952. Mayacaceae. Notul. Syst. 14: 234-248. 

Lourteig, A. 1971. Mayacaceae. In T. Lasser, Flora de Venezuela 3(1): 197-203. 

Lourteig, A. 2001. Mayacaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 260-262. 

Mori, S.A. 1997. Mayacaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Musa x paradisiaca L. [widely cultivated] 

Musa spp. [cultivated only] 

MUSACEAE 

K. Yatskievych 


Musa bihai L. (Heliconiaceae) =Heliconia bihai 

Musa humilis Aubl. (Heliconiaceae) =Heliconia psittacorum 

Musa sapientum L. =Musa ×paradisiaca 

REFERENCES 

Maas, P.J.M. 1985. Musaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (Series A) 1: 1-28. 


Mori, S.A. 1997. Musaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide 



Yatskievych, K. 2001. Musaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 731-732. 

NAJADACEAE 


Najas guadalupensis (Spreng.) Magnus 

Najas microcarpa K. Schum. [expected in GU, SU; Mt Roraima] 

Najas wrightiana A. Braun [as circumscribed by Lowden, not in Guianas] 


Najas conferta (A. Braun) A. Braun [not in flora area] 

Najas marina L. [widespread, perhaps expected] 

Najas microdon A. Braun =Najas guadalupensis 




GU SU FG 

FG? 

BO FG 

BO FG? 

REFERENCES 

Haynes, R.R. & L.B. Holm-Nielsen. 2003. Najadaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 

the Venezuelan Guayana 7: 100-101. 

Lowden, R.M. 1986. Taxonomy of the genus Najas in the Neotropics. Aquatic Bot. 24: 147-184.

118 


ORCHIDACEAE 

G. Carnevali, E. Christenson, E. Foldats, I.M. Ramírez-Morillo, G.A. Romero-González, 

C.A. Vargas, M. Werkhoven 

Acacallis cyanea Lindl. 

Acacallis fimbriata (Rchb. f.) Schltr. 

Acineta alticola C. Schweinf. 

Acineta superba (Kunth) Rchb. f. 

Ada keiliana (Rchb. f. ex Lindl.) N.H. Williams 

Aganisia pulchella Lindl. 

Aracamunia liesneri Carnevali & I. Ramírez 

Aspasia variegata Lindl. 

Aspidogyne confusa (C. Schweinf.) Garay 

Aspidogyne foliosa (Poepp. & Endl.) Garay 

Aspidogyne longicornu (Cogn.) Garay 

Aspidogyne multifoliata (C. Schweinf.) Garay 

Aspidogyne pumila (Cogn.) Garay 

Aspidogyne robusta (C. Schweinf.) Garay 

Aspidogyne steyermarkii Carnevali & Foldats 

Barbosella orbicularis Luer 

Baskervilla colombiana Garay 

Batemannia armillata Rchb. f. 

Batemannia colleyi Lindl. 

Batemannia lepida Rchb. f. 

Beloglottis costaricensis (Rchb. f.) Schltr. 

Bifrenaria longicornis Lindl. 

Bifrenaria steyermarkii (Foldats) Garay & Dunst. 

Bifrenaria venezuelana C. Schweinf. 

Bletia altilamellata Garay 

Bletia purpurea (Lam.) DC. 

Bletia stenophylla Schltr. 

Bollea hemixantha Rchb. f. 

Bollea violacea (Lindl.) Rchb. f. 

Brachionidium brevicaudatum Rolfe 

Brachionidium juliani Carnevali & I. Ramírez 

Brachionidium longicaudatum Ames & C. Schweinf. 

Brachionidium neblinense Carnevali & I. Ramírez 

Brachionidium parvum Cogn. 

Brachystele guayanensis (Lindl.) Schltr. 

Braemia vittata (Lindl.) Jenny 

Brassavola cucullata (L.) R. Br. 

Brassavola gardneri Cogn. 

Brassavola martiana Lindl. 

Brassavola nodosa (L.) Lindl. 

Brassia angusta Lindl. 

Brassia bidens Lindl. 

Brassia caudata (L.) Lindl. 

Brassia chloroleuca Barb. Rodr. 

Brassia forgetiana hort. ex C. Schweinf. 

Brassia huebneri Schltr. 

Brassia lanceana Lindl. 

Brassia lawrenceana Lindl. 

Brassia macrostachya Lindl. 

Brassia neglecta Rchb. f. 

Brassia pumila Lindl. 

Brassia villosa Lindl. 

Brassia wageneri Rchb. f. 

Bulbophyllum bracteolatum Lindl. 

Bulbophyllum dunstervillei Garay 

Bulbophyllum exaltatum Lindl. 

Bulbophyllum geraense Rchb. f. 

Bulbophyllum meridense Rchb. f. 

Bulbophyllum meristorhachis Garay & Dunst. 

Bulbophyllum oerstedii (Rchb. f.) Hemsl. 

AM 

AM GU SU 

AM BO GU? 

SU 

GU 

BO GU SU FG 

AM 


AM BO FG 

GU SU FG 

GU SU 

GU 

BO GU SU 

BO GU 

BO 

BO 

BO 

SU 


AM SU 

SU 


BO GU 

AM BO 

GU 

SU? FG 

AM BO 

AM BO GU FG? 

GU SU FG 

AM? BO GU 

AM 

AM BO GU 

AM 

AM BO 




SU? FG 


GU FG 

AM BO GU 

AM BO GU 


GU FG 

AM BO 

FG 


BO GU SU FG 

AM BO GU SU 


GU FG 

GU 

GU 

GU SU FG 

BO 

AM BO GU 

BO GU 

AM BO 

BO 

AM BO SU FG


Bulbophyllum pachyrachis (A. Rich.) Griseb. 

Bulbophyllum quadrisetum Lindl. 

Bulbophyllum roraimense Rolfe 

Bulbophyllum setigerum Lindl. 

Caluera surinamensis Dodson & Determann 

Caluera vulpina Dodson & Determann 

Campylocentrum fasciola (Lindl.) Cogn. 

Campylocentrum hondurense Ames 

Campylocentrum huebneri Mansf. 

Campylocentrum lansbergii (Rchb. f.) Schltr. 

Campylocentrum micranthum (Lindl.) Rolfe 

Campylocentrum nataliae Carnevali & I. Ramírez 

Campylocentrum pachyrrhizum (Rchb. f.) Rolfe 

Campylocentrum panamense Ames 

Campylocentrum poeppigii (Rchb. f.) Rolfe 

Catasetum barbatum (Lindl.) Lindl. 

Catasetum barbatum var. spinosum Rolfe 

Catasetum bergoldianum Foldats 

Catasetum bicallosum Cogn. 

Catasetum bicolor Klotzsch 

Catasetum callosum Lindl. 

Catasetum callosum var. carunculatum Mansf. 

Catasetum collare Cogn. 

Catasetum costatum Rchb. f. 

Catasetum cristatum Lindl. 

Catasetum deltoideum (Lindl.) Mutel 

Catasetum discolor (Lindl.) Lindl. 

Catasetum x dunstervillei G.A. Romero & Carnevali [natural hybrid of 

C. pileatum and C. discolor] 

Catasetum ferox Kraenzl. 

Catasetum gnomus Linden & Rchb. f. 

Catasetum gomezii G.A. Romero & Carnevali 

Catasetum x guianense Jenny & G.A. Romero 

Catasetum longifolium Lindl. 

Catasetum luridum (Link) Lindl. 

Catasetum macrocarpum Rich. ex Kunth 

Catasetum maroaënse G.A. Romero & C. Gómez 

Catasetum merchae G.A. Romero 

Catasetum microglossum Rolfe 

Catasetum ochraceum Lindl. 

Catasetum parguazense G.A. Romero & Carnevali 

Catasetum pileatum Rchb. f. 

Catasetum planiceps Lindl. 

Catasetum poriferum R.H. Schomb. ex Lindl. 

Catasetum rivularium Barb. Rodr. 

Catasetum x roseo-album (Hook.) Lindl. [natural hybrid of C. discolor and 

C. longifolium] 

Catasetum saccatum Lindl. 

Catasetum x tapiriceps Rchb. f. [natural hybrid of C. macrocarpum and C. 

pileatum] 

Catasetum thompsonii Dodson 

Catasetum x wendlingeri Foldats [natural hybrid of C. pileatum and C. 

planiceps] 

Catasetum yavitaënse G.A. Romero & C Gómez 

Cattleya jenmanii Rolfe 

Cattleya lawrenceana Rchb. f. 

Cattleya violacea (Kunth) Rolfe 

Caularthron bicornutum (Hook.) Raf. 

Caularthron bilamellatum (Rchb. f.) R.E. Schult. 

Caularthron bilamellatum f. indivisa (Bradf. ex Griseb.) Carnevali & I. 

Ramírez 

Chaubardia surinamensis Rchb. f. 

Chaubardiella tigrina (Garay & Dunst.) Garay 

Cheiradenia cuspidata Lindl. 

FG 

SU FG 

BO GU FG 

AM GU 

SU 

FG 


AM GU 

AM 

BO DA 


BO 

BO GU SU FG 

AM BO 

AM BO GU 


FG? 

AM 

AM GU? 

AM BO GU? 

AM GU SU? FG 

GU SU? FG 

AM BO 

AM 


GU SU FG? 


AM BO GU? 

AM GU? 

GU? 

AM 

GU SU? FG 


GU 


AM 

AM 

AM 

GU? 

BO 

AM BO GU? 


GU 

AM 


GU 

AM BO GU? FG? 

GU 

AM BO GU 

AM 

BO GU 

AM BO GU 


AM BO GU SU 

BO GU? 

BO GU SU FG 

BO GU SU FG 

BO GU SU FG

120 


Chelyorchis ampliata (Lindl.) Dressler & N.H. Williams 

Cleistes grandiflora (Aubl.) Schltr. 

Cleistes huberi Carnevali & I. Ramírez 

Cleistes lepida (Rchb. f.) Schltr. 

Cleistes paludosa Rchb. f. 

Cleistes parviflora Lindl. 

Cleistes rosea Lindl. 

Cleistes stricta (C. Schweinf.) Garay & Dunst. 

Cleistes tenuis (Rchb. f. ex Griseb.) Schltr. 

Cleistes triflora (C. Schweinf.) Carnevali & I. Ramírez [Mt. Roraima] 

Cleistes unifoliata (C. Schweinf.) Carnevali & I. Ramírez 

Clowesia warczewiczii (Lindl. & Paxton) Dodson 

Cochleanthes flabelliformis (Sw.) R.E. Schult. & Garay 

Cochleanthes guianensis Lafontaine, G. Gerlach & Senghas 

Comparettia falcata Poepp. & Endl. 

Coryanthes albertinae H. Karst. 

Coryanthes cataniapoënsis G.A. Romero & Carnevali 

Coryanthes feildingii Lindl. 

Coryanthes gomezii G.A. Romero & G. Gerlach 

Coryanthes macrantha (Hook.) Hook. 

Coryanthes maculata Hook. 

Coryanthes pegiae G.A. Romero 

Coryanthes punctata Beer 

Coryanthes rutkisii Foldats 

Coryanthes speciosa (Hook.) Hook. 

Corymborkis sp. 

Cranichis diphylla Sw. 

Cranichis sp. 

Cryptarrhena guatemalensis Schltr. 

Cryptarrhena kegelii Rchb. f. 

Cryptocentrum dunstervilleorum Carnevali & G.A. Romero 

Cryptocentrum peruvianum (Cogn.) C. Schweinf. 

Cryptocentrum peruvianum subsp. peruvianum 

Cryptocentrum sp. nov. [Cryptocentrum peruvianum auct., Cryptocentrum 

gracillimum auct.] 

Cyclopogon cranichoides (Griseb.) Schltr. 

Cyclopogon elatus (Sw.) Schltr. 

Cyclopogon olivaceus (Rolfe) Schltr. 

Cycnoches chlorochilon Klotzsch 

Cycnoches haagii Barb. Rodr. 

Cycnoches loddigesii Lindl. 

Cycnoches lusiae G.A. Romero & Garay 

Cycnoches thurstonorum Dodson 

Cyrtopodium andersonii (Lamb. ex Andrews) R. Br. 

Cyrtopodium cristatum Lindl. 

Cyrtopodium fowliei L.C. Menezes 

Cyrtopodium graniticum G.A. Romero & Carnevali 

Cyrtopodium intermedium Brade 

Cyrtopodium paniculatum (Ruiz & Pav.) Garay 

Cyrtopodium parviflorum Lindl. 

Degranvillea dermaptera Determann 

Diadenium barkeri (Lindl.) Benth. & Hook. f. 

Dichaea ancoraelabia C. Schweinf. 

Dichaea angustifolia H.G. Jones 

Dichaea brevicaulis Cogn. 

Dichaea bryophila Rchb. f. 

Dichaea camaridioides Schltr. [Mt. Roraima] 

Dichaea cornuta S. Moore 

Dichaea histrio Rchb. f. 

Dichaea hookeri Garay & H.R. Sweet 

Dichaea hystricina Rchb. f. 

Dichaea kegelii Rchb. f. 

Dichaea latifolia Lindl. 

Dichaea latifolia var. longa (Schltr.) Folsom 

BO DA 

GU FG 

BO 

BO GU 

SU FG 



AM BO 

AM BO 

BO 

GU SU FG 

BO SU 

AM BO 

BO 

GU? SU? 

GU SU FG 

AM BO FG 

AM 

AM 

BO GU SU FG 

AM 

BO GU 


AM 

BO 

BO 

GU 

GU 

SU FG 

GU SU FG 

GU SU? 

BO SU 

SU FG 

GU SU FG 

BO GU SU FG 

AM 

AM 

BO GU 

GU 

BO 

BO SU 

GU SU FG 

BO GU SU? FG 


BO GU SU FG 

BO 

AM 



BO GU SU 

AM 

SU 

GU SU FG 

BO GU SU FG 

FG 

BO 

AM GU SU FG 

GU 

FG? 

FG? 

AM GU? 

SU 

GU? SU? 


AM BO GU 

BO GU SU FG 

BO 

BO


Dichaea morrisii Fawc. & Rendle 

Dichaea ochracea Lindl. 

Dichaea panamensis Lindl. 

Dichaea pendula (Aubl.) Cogn. 

Dichaea picta Rchb. f. 

Dichaea rendlei Gleason 

Dichaea robusta Schltr. 

Dichaea splitgerberi Rchb. f. 

Dichaea tachirensis G.A. Romero & Carnevali 

Dichaea trinitensis Gleason 

Dichaea trulla Rchb. f. 

Dichaea venezuelensis Carnevali & I. Ramírez 

Dichaea weigeltii Rchb. f. 

Dimerandra elegans (H. Focke) Siegerist 

Dimerandra emarginata (G. Mey.) Hoehne 

Dipteranthus planifolius (Rchb. f.) Garay 

Discyphus scopulariae (Rchb. f.) Schltr. 

Dryadella lueriana Carnevali & G.A. Romero 

Duckeella alticola C. Schweinf. [expected in GU] 

Duckeella pauciflora Garay 

Dunstervillea mirabilis Garay 

Elleanthus arpophyllostachys (Rchb. f.) Rchb. f. 

Elleanthus brasiliensis (Lindl.) Rchb. f. 

Elleanthus caravata (Aubl.) Rchb. f. 

Elleanthus cephalotus Garay & H.R. Sweet 

Elleanthus columnaris (Lindl.) Rchb. f. 

Elleanthus confusus Garay 

Elleanthus crinipes Rchb. f. 

Elleanthus furfuraceus (Lindl.) Rchb. f. 

Elleanthus gracilis (Rchb. f.) Rchb. f. 

Elleanthus graminifolius (Barb. Rodr.) Lrjtnant 

Elleanthus linifolius C. Presl 

Elleanthus malpighiiflorus Carnevali & G.A. Romero [expected in AM; 

Serra da Neblina] 

Elleanthus norae Garay & Dunst. 

Elleanthus sphaerocephalus Schltr. 

Elleanthus trilobatus Ames & C. Schweinf. 

Elleanthus wageneri (Rchb. f.) Rchb. f. 

Eloyella panamensis (Dressler) Dodson 

Eltroplectris calcarata (Sw.) Garay & H.R. Sweet 

Encyclia auyantepuiensis Carnevali & I. Ramírez 

Encyclia chloroleuca (Hook.) Neumann 

Encyclia cochleata (L.) Lemée 

Encyclia conchaechila (Barb. Rodr.) Porto & Brade 

Encyclia cordigera (Kunth) Dressler 

Encyclia dichroma (Lindl.) Schltr. 

Encyclia diurna (Jacq.) Schltr. 

Encyclia granitica (Bateman ex Lindl.) Schltr. 

Encyclia guianensis Carnevali & G.A. Romero 

Encyclia ionosma (Lindl.) Schltr. 

Encyclia ivonae Carnevali & G.A. Romero 

Encyclia leucantha Schltr. 

Encyclia longifolia Barb. Rodr. 

Encyclia odoratissima (Lindl.) Schltr. 

Encyclia pachyantha (Lindl.) Hoehne 

Encyclia picta (Lindl.) Hoehne 

Encyclia pilosa (C. Schweinf.) Carnevali & I. Ramírez 

Encyclia remotiflora (C. Schweinf.) Carnevali & I. Ramírez 

Encyclia selligera (Bateman ex Lindl.) Schltr. [not in flora area, fide E. 

Christenson, specimens =E. granitica] 

Encyclia tarumana Schltr. 

Epidendrum agathosmicum Rchb. f. 

Epidendrum alsum Ridl. ex Thurn 

Epidendrum althausenii A.D. Hawkes 

GU? 

GU 

AM BO DA 

AM BO GU FG 

AM BO DA GU 

GU SU FG 

AM BO DA 

AM 

AM 


AM GU SU FG 


AM BO GU 

GU SU 


BO 

BO 

BO 

BO 

DA GU SU FG 

AM 

AM 

BO 

BO 

AM BO GU 

FG 


GU SU FG 

AM BO 

AM BO 

FG 

GU SU? 

AM BO 

AM BO GU SU 

SU 

FG 

AM BO 

AM BO 

GU 

BO 

BO GU 

BO SU 

BO 


GU SU? FG 

AM GU 

BO GU FG 

SU 

BO GU SU FG 


BO GU SU 

GU SU? FG 

BO GU 

AM BO 

SU? 

GU FG 

DA GU SU? FG 

GU 

AM GU 

AM 

SU? 

FG 

AM 

BO GU 

AM DA GU SU FG

122 


Epidendrum amazonicoriifolium Hágsater 

Epidendrum anceps Jacq. 

Epidendrum apaganum Mansf. 

Epidendrum apuahuense Mansf. 

Epidendrum attenuatum Lindl. 

Epidendrum bakrense Hágsater & Cremers 

Epidendrum calanthum Rchb. f. & Warsz. 

Epidendrum carpophorum Barb. Rodr. 

Epidendrum caurense Carnevali & G.A. Romero 

Epidendrum chimantense Hágsater & Carnevali 

Epidendrum churunense Garay & Dunst. 

Epidendrum ciliare L. 

Epidendrum ciliare var. squamatum Schnee 

Epidendrum commelinispathum Carnevali & I. Ramírez [expected in AM; 

Serra da Neblina] 

Epidendrum compressum Griseb. 

Epidendrum congestoides Ames & C. Schweinf. 

Epidendrum cooperianum Bateman 

Epidendrum coronatum Ruiz & Pav. 

Epidendrum cristatum Ruiz & Pav. 

Epidendrum densiflorum Hook. 

Epidendrum dichaeoides Carnevali & G.A. Romero 

Epidendrum difforme Jacq. 

Epidendrum diffusum Sw. 

Epidendrum diffusum var. depauperatum Cogn. [=Epidendrum compressum?] 

Epidendrum dipus Lindl. 

Epidendrum durum Lindl. 

Epidendrum ecostatum Pabst ex Angley 

Epidendrum flexuosum G. Mey. 

Epidendrum fraudulentum Rchb. f. 

Epidendrum geminiflorum Kunth 

Epidendrum gracillimum Lindl. 

Epidendrum gramineum Lindl. 

Epidendrum gratissimum (Rchb. f.) Hágsater 

Epidendrum hombersleyi Summerh. 

Epidendrum ibaguense Kunth 

Epidendrum ibaguense Kunth x secundum Jacq. 

Epidendrum imitans Schltr. 

Epidendrum imthurnii Ridl. ex Thurn 

Epidendrum isomerum Schltr. 

Epidendrum klotzscheanum Rchb. f. 

Epidendrum latilabre Lindl. 

Epidendrum leucochilum Klotzsch 

Epidendrum longicolle Lindl. 

Epidendrum macrocarpum Rich. 

Epidendrum magnicallosum C. Schweinf. 

Epidendrum micronocturnum Carnevali & G.A. Romero 

Epidendrum microphyllum Lindl. 

Epidendrum miserrimum Rchb. f. 

Epidendrum montigenum Ridl. ex Thurn 

Epidendrum myrmecophorum Barb. Rodr. 

Epidendrum nocturnum Jacq. 

Epidendrum norae Carnevali & G.A. Romero 

Epidendrum nuriense Carnevali & Hágsater 

Epidendrum nutans Sw. 

Epidendrum oldemanii Christenson 

Epidendrum orchidiflorum Salzm. 

Epidendrum pachyphyton Garay 

Epidendrum paniculatum Ruiz & Pav. [=Epidendrum densiflorum?] 

Epidendrum paruimense G.A. Romero & Carnevali [expected in BO] 

Epidendrum patens Sw. 

Epidendrum prostratum (Lindl.) Cogn. 

Epidendrum pseudoramosum Schltr. 

AM BO GU 

BO GU SU FG 

SU 

AM 

BO 

FG 

BO GU? 


BO 

BO 

BO GU 

BO GU SU FG 

BO 


DA 

GU 

AM BO SU 

BO GU SU 

BO GU SU FG 

AM 

GU SU FG 

GU SU 

GU SU 

FG 

AM BO GU 

GU FG 


GU? 

SU 

SU? 

SU? 

GU 

AM 

AM BO GU SU? FG 

BO 

GU FG 

AM BO GU FG 

GU 

AM 

GU SU? FG 

BO 

AM BO DA GU 


AM 

BO 


AM BO GU SU 

BO GU 

AM BO 


AM BO GU 

BO 

SU 

FG 


AM BO GU 

GU SU FG 

GU 

GU? 

BO GU 

BO GU


Epidendrum purpurascens H. Focke 

Epidendrum ramosum Jacq. 

Epidendrum repens Cogn. 

Epidendrum revertianum (Stehlé) Hágsater 

Epidendrum rigidum Jacq. 

Epidendrum rostratum Garay & Dunst. 

Epidendrum saxatile Lindl. 

Epidendrum schlechterianum Ames 

Epidendrum schlimii Rchb. f. 

Epidendrum sculptum Rchb. f. 

Epidendrum sculptum var. linearifolium Rchb. f. 

Epidendrum sculptum var. sculptum 

Epidendrum secundum Jacq. 

Epidendrum sertorum Garay & Dunst. 

Epidendrum smaragdinum Lindl. 

Epidendrum sp. A 

Epidendrum splendens Schltr. 

Epidendrum stalkyi Carnevali & G.A. Romero 

Epidendrum stamfordianum Bateman 

Epidendrum stiliferum Dressler 

Epidendrum strobiliferum Rchb. f. 

Epidendrum strobiloides Garay & Dunst. 

Epidendrum tenuifolium Lindl. 

Epidendrum tridens Poepp. & Endl. 

Epidendrum tumuc-humaciense (Veyret) Carnevali & G.A. Romero 

Epidendrum ulei Schltr. 

Epidendrum unguiculatum (C. Schweinf.) Garay & Dunst. 

Epidendrum urbanianum Cogn. 

Epidendrum urichianum Carnevali, Foldats & I. Ramírez 

Epidendrum vincentinum Lindl. 

Epidendrum violascens Ridl. ex Thurn 

Epidendrum viviparum Lindl. 

Epidendrum xanthium Lindl. 

Epistephium amplexicaule (Ruiz & Pav.) Poepp. & Endl. 

Epistephium duckei Huber 

Epistephium elatum Kunth 

Epistephium ellipticum R.O. Williams & Summerh. 

Epistephium hernandii Garay 

Epistephium lucidum Cogn. 

Epistephium parviflorum Lindl. 

Epistephium sclerophyllum Lindl. 

Epistephium subrepens Hoehne 

Eriopsis biloba Lindl. 

Eriopsis rutidobulbon Hook. 

Eriopsis sceptrum Rchb. f. & Warsz. 

Erycina glossomystax (Rchb. f.) N.H. Williams & M.W. Chase 

Erycina pumilio (Rchb. f.) N.H. Williams & M.W. Chase 

Erycina pusilla (L.) N.H. Williams & M.W. Chase 

Erythrodes arietina (Rchb. f. & Warm.) Ames 

Erythrodes hirtella (Sw.) Fawc. & Rendle 

Erythrodes major (Presl) Ames 

Erythrodes paleacea (Schltr.) Ames 

Erythrodes pumila (Cogn.) Pabst 

Eulophia alta (L.) Fawc. & Rendle 

Eurystyles cotyledon Wawra 

Galeandra badia Garay & G.A. Romero 

Galeandra baueri Lindl. 

Galeandra baueri var. baueri 

Galeandra baueri var. lutea S. Watson 

Galeandra beyrichii Rchb. f. 

Galeandra carnevaliana G.A. Romero & Warford 

Galeandra devoniana R.H. Schomb. ex Lindl. 

Galeandra dives Rchb. f. 



AM BO GU 

GU 


FG 

AM BO GU 

GU SU 

BO 

DA GU SU FG 

SU 

DA GU SU FG 

AM BO GU 

AM 


BO GU 

BO 

BO 

AM BO 

SU? 


BO GU SU FG 

GU 

GU SU 


AM BO GU 

AM SU FG 

AM BO 

AM BO GU 

AM BO GU 

AM BO GU 

GU FG? 

GU? 

GU 


AM BO 

AM BO GU 

AM 

GU 


AM SU? 

AM BO GU 


BO 



SU FG 


AM BO 

GU SU? FG 

GU SU 

AM BO GU 

SU 


BO 

BO GU SU FG 

FG 

FG 

FG 

BO GU SU FG 

BO 

AM GU 

GU SU FG

124 


Galeandra duidensis Garay & G.A. Romero 

Galeandra lagoensis Rchb. f. & Warm. 

Galeandra macroplectra G.A. Romero & Warford 

Galeandra magnicolumna G.A. Romero & Warford 

Galeandra minax Rchb. f. 

Galeandra stangeana Rchb. f. 

Galeandra stylomisantha (Vell.) Hoehne 

Galeottia burkei (Rchb. f.) Dressler & Christenson 

Galeottia jorisiana (Rolfe) Schltr. 

Gomphichis adnata (Ridl. ex Thurn) Schltr. [=Gomphichis viscosa?] 

Gomphichis costaricensis (Schltr.) Ames, Hubb. & C. Schweinf. 

Gomphichis viscosa (Rchb. f.) Schltr. 

Gongora atropurpurea Hook. 

Gongora histrionica Rchb. f. 

Gongora maculata Lindl. 

Gongora nigrita Lindl. 

Gongora pleiochroma Rchb. f. 

Govenia utriculata (Sw.) Lindl. 

Guanchezia maguirei (C. Schweinf.) G.A. Romero & Carnevali 

Habenaria abortiens Lindl. 

Habenaria alata Hook. 

Habenaria alterosula Snuv. & Westra 

Habenaria amambayensis Schltr. [expected in BO and AM] 

Habenaria armata Rchb. f. 

Habenaria ayangannensis Renz [expected in BO] 

Habenaria caldensis Kraenzl. 

Habenaria distans Griseb. 

Habenaria dusenii Schltr. 

Habenaria entomantha (La Llave & Lex.) Lindl. 

Habenaria ernestii Schltr. 

Habenaria floribunda Lindl. 

Habenaria heptadactyla Rchb. f. 

Habenaria huberi Carnevali & Morillo 

Habenaria lehmanniana Kraenzl. 

Habenaria leprieuri Rchb. f. 

Habenaria longicauda Hook. 

Habenaria longicauda subsp. ecalcarata Snuv. & Westra 

Habenaria longicauda subsp. longicauda 

Habenaria macilenta (Lindl.) Rchb. f. 

Habenaria mesodactyla Griseb. 

Habenaria monorrhiza (Sw.) Rchb. f. [expected in the Guianas] 

Habenaria montis-wilhelminae Renz 

Habenaria nilssonii Foldats 

Habenaria obtusa Lindl. 

Habenaria odontopetala Rchb. f. 

Habenaria parvidens Lindl. 

Habenaria parviflora Lindl. 

Habenaria platydactyla Kraenzl. 

Habenaria pratensis (Lindl.) Rchb. f. 

Habenaria pratensis var. parviflora Cogn. 

Habenaria pygmaea C. Schweinf. & R.E. Schult. 

Habenaria quadrata Lindl. 

Habenaria quinqueseta (Michx.) A. Eaton 

Habenaria repens Nutt. 

Habenaria rodeiensis Barb. Rodr. 

Habenaria rodriquezii Cogn. 

Habenaria roraimensis Rolfe [Mt. Roraima] 

Habenaria santensis Barb. Rodr. 

Habenaria schomburgkii Lindl. ex Benth. 

Habenaria setacea Lindl. 

Habenaria setacea var. setacea 

Habenaria seticauda Lindl. ex Benth. 

Habenaria sp. A 

Habenaria sp. B 

AM 

SU 

AM 

AM 

AM 

AM 


AM BO GU SU? 

AM BO GU 

BO 

BO 

GU 

GU 


GU? SU FG? 

GU FG? 

GU SU? 

AM 

AM 

BO DA GU SU 

SU? 

SU? 

FG 

FG? 

GU SU FG 

GU FG 

AM BO GU 

GU 

BO 

BO 

GU SU FG? 

AM BO SU FG 

GU SU FG? 

AM BO GU FG 

BO GU 

AM 

AM 

BO GU SU 

BO GU FG 



GU SU FG 

GU SU 

BO 

BO 

BO 

GU SU 

SU 

FG 

BO GU 

GU 

SU 

SU? 

FG? 

BO GU SU? FG? 

GU SU FG 

BO GU? SU FG 

AM 

BO GU? SU FG 

AM BO 

BO GU SU FG? 


GU FG 

FG 

AM BO GU 

SU? 

BO GU SU 

BO GU FG 

BO GU FG 

AM 

AM 

GU FG

Habenaria sprucei Cogn. 

Habenaria trifida Kunth 

Helonoma americana (C. Schweinf. & Garay) Garay 

Helonoma bifida (Ridl. ex Thurn) Garay 

Helonoma chiropterae (Szlach.) Carnevali & G.A. Romero 

Heterotaxis proboscidea (Rchb. f.) F. Barros 

Heterotaxis superflua (Rchb. f.) F. Barros 

Heterotaxis villosa (Barb. Rodr.) F. Barros 

Heterotaxis violaceopunctata (Rchb. f.) F. Barros 

Hexisea bidentata Lindl. 

Hexisea imbricata (Lindl.) Rchb. f. 

Houlletia odoratissima Linden ex Lindl. 

Houlletia roraimensis Rolfe 

Huntleya lucida (Rolfe) Rolfe 

Huntleya meleagris Lindl. 

Huntleya sessiliflora Bateman ex Lindl. 

Hylaeorchis petiolaris (Schltr.) Carnevali & G.A. Romero 

Ionopsis satyrioides (Sw.) Rchb. f. 

Ionopsis utricularioides (Sw.) Lindl. 

Isochilus linearis (Jacq.) R. Br. 

Jacquiniella globosa (Jacq.) Schltr. 

Jacquiniella leucomelana (Rchb. f.) Schltr. 

Jacquiniella steyermarkii Carnevali & Dressler 

Jacquiniella teretifolia (Sw.) Britton & P. Wilson 

Kefersteinia lafontainei Senghas & G. Gerlach 

Kegeliella houtteana (Rchb. f.) L.O. Williams 

Kegeliella orientalis G. Gerlach 

Koellensteinia carraoënsis Garay & Dunst. 

Koellensteinia graminea (Lindl.) Rchb. f. 

Koellensteinia hyacinthoides Schltr. 

Koellensteinia kellneriana Rchb. f. 

Koellensteinia lilijae Foldats 

Koellensteinia roraimae Schltr. 

Koellensteinia tricolor (Lindl.) Rchb. f. 

Leochilus labiatus (Sw.) Kuntze 

Lepanthes aithalos Carnevali & I. Ramírez 

Lepanthes cercion Luer & Escobar 

Lepanthes duidensis Ames & C. Schweinf. 

Lepanthes dussii Urb. 

Lepanthes exilis C. Schweinf. 

Lepanthes helicocephala Rchb. f. 

Lepanthes marahuacensis Carnevali & I. Ramírez 

Lepanthes pariaënsis Foldats 

Lepanthes pectinata Luer 

Lepanthes ruscifolia Rchb. f. 

Lepanthes unitrinervis Carnevali & I. Ramírez 

Lepanthopsis floripecten (Rchb. f.) Ames 

Lepanthopsis obliquipetala (Ames & C. Schweinf.) Luer 

Lepanthopsis pulchella Garay & Dunst. 

Lepanthopsis vinacea C. Schweinf. 

Leucohyle mutica (Lindl.) Schltr. 

Leucohyle subulata (Sw.) Schltr. 

Ligeophila amazonica Garay 

Ligeophila clavigera (Rchb. f.) Garay 

Ligeophila juruensis (Hoehne) Garay 

Ligeophila peterana (Cogn.) Garay 

Ligeophila rosea (Lindl.) Garay 

Ligeophila stigmatoptera (Rchb. f.) Garay 

Liparis jamaicensis Lindl. ex Griseb. 

Liparis nervosa (Thunb.) Lindl. subsp. A 

Liparis nervosa subsp. nervosa 

Liparis verticillata G.A. Romero & Garay 

Lockhartia acuta (Lindl.) Rchb. f. 

Lockhartia biserra (Rich.) Christenson & Garay 


BO GU SU FG 


AM BO GU 

AM BO GU 

AM 

AM 





AM BO GU SU 

AM BO GU SU? 

AM BO GU 

GU 

BO GU 

GU 

AM BO GU 



BO GU SU FG 


GU 

AM BO GU 


GU? SU FG 

SU FG 

BO 

BO GU SU FG 


AM SU? FG? 

GU SU FG 

AM 

BO 

AM BO GU 

BO 

BO 

BO 

AM BO 

AM BO 

BO 


AM 

AM BO 

AM BO 

SU FG 

BO 

AM BO GU FG 

AM BO 

BO 

BO 

BO DA GU SU 

GU 

AM 

GU SU FG 

AM BO SU 

SU FG 

SU? 


BO GU 

AM BO 


BO 

DA 

GU

126 


Lockhartia imbricata (Lam.) Hoehne 

Lockhartia latilabris C. Schweinf. 

Lockhartia parthenocomos Rchb. f. 

Lueddemannia pescatorei (Lindl.) Linden & Rchb. f. 

Lycaste fulvescens Hook. 

Lycaste macrophylla (Poepp. & Endl.) Lindl. 

Lyroglossa grisebachii (Cogn.) Schltr. 

Macradenia brassavolae Rchb. f. 

Macradenia loxoglottis H. Focke & Rchb. f. ex Rchb. f. 

Macradenia lutescens R. Br. 

Macradenia rubescens Barb. Rodr. 

Macroclinium mirabile (C. Schweinf.) Dodson 

Macroclinium wullschlaegelianum (H. Focke) Dodson 

Malaxis excavata (Lindl.) Kuntze 

Malaxis maguirei C. Schweinf. 

Malaxis umbellifera Sw. 

Masdevallia cuprea Lindl. 

Masdevallia guayanensis Lindl. 

Masdevallia infracta Lindl. 

Masdevallia manarana Carnevali & I. Ramírez 

Masdevallia minuta Lindl. 

Masdevallia norae Luer 

Masdevallia picturata Rchb. f. 

Masdevallia sprucei Rchb. f. 

Masdevallia vargasii C. Schweinf. 

Masdevallia wendlandiana Rchb. f. 

Maxillaria acutifolia Lindl. 

Maxillaria alba (Hook.) Lindl. 

Maxillaria albiflora Ames & C. Schweinf. 

Maxillaria alpestris Lindl. 

Maxillaria alticola C. Schweinf. 

Maxillaria aurea (Poepp. & Endl.) L.O. Williams 

Maxillaria auyantepuiensis Foldats 

Maxillaria bolivarensis C. Schweinf. 

Maxillaria brachybulbon Schltr. 

Maxillaria caespitifica Rchb. f. 

Maxillaria camaridii Rchb. f. 

Maxillaria candida Lodd. ex Lindl. 

Maxillaria chlorantha Lindl. 

Maxillaria christobalensis Rchb. f. 

Maxillaria colemanii Carnevali & W. Fritz [expected in GU] 

Maxillaria conferta (Griseb.) C. Schweinf. ex León 

Maxillaria connellii Rolfe 

Maxillaria crassifolia (Lindl.) Rchb. f. 

Maxillaria cryptobulbon Carnevali & J.T. Atwood 

Maxillaria desvauxiana Rchb. f. 

Maxillaria discolor (Lodd. ex Lindl.) Rchb. f. 

Maxillaria x dunstervillei Carnevali & I. Ramírez [natural hybrid of M. 

meridensis Lindl. and M. quelchii Rolfe] 

Maxillaria eburnea Lindl. 

Maxillaria equitans (Schltr.) Garay 

Maxillaria foldatsiana Carnevali & I. Ramírez 

Maxillaria grobyoides Garay & Dunst. 

Maxillaria guadalupensis Cogn. 

Maxillaria guareimensis Rchb. f. 

Maxillaria imbricata Barb. Rodr. 

Maxillaria jenischiana (Rchb. f.) C. Schweinf. 

Maxillaria kegelii Rchb. f. 

Maxillaria lasallei Foldats 

Maxillaria leucaimata Barb. Rodr. 

Maxillaria liparophylla Summerh. 

Maxillaria longipetiolata Ames & C. Schweinf. 

Maxillaria loretoensis C. Schweinf. 

Maxillaria luteoalba Lindl. 


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Maxillaria mapiriensis (Kraenzl.) L.O. Williams 

Maxillaria marmoliana Dodson 

Maxillaria melina Lindl. 

Maxillaria meridensis Lindl. 

Maxillaria monacensis Kraenzl. 

Maxillaria nasuta Rchb. f. 

Maxillaria notylioglossa Rchb. f. 

Maxillaria nuriensis Carnevali & I. Ramírez 

Maxillaria ochroleuca Lodd. ex Lindl. 

Maxillaria ochroleuca var. longipes Sander 

Maxillaria pannieri Foldats 

Maxillaria parkeri Hook. 

Maxillaria parviflora (Poepp. & Endl.) Garay 

Maxillaria pauciflora Barb. Rodr. 

Maxillaria ponerantha Rchb. f. 

Maxillaria porrecta Lindl. 

Maxillaria prolifera Ruiz & Pav. 

Maxillaria pterocarpa Barb. Rodr. 

Maxillaria pumila Hook. 

Maxillaria quelchii Rolfe 

Maxillaria ramosa Ruiz & Pav. 

Maxillaria reichenheimiana Endres & Rchb. f. 

Maxillaria rigida Barb. Rodr. 

Maxillaria rufescens Lindl. 

Maxillaria santanae Carnevali & I. Ramírez 

Maxillaria schlechteri Foldats 

Maxillaria scorpioidea Kraenzl. 

Maxillaria setigera Lindl. 

Maxillaria splendens Poepp. & Endl. 

Maxillaria squamata Barb. Rodr. 

Maxillaria stenophylla Rchb. f. 

Maxillaria tenuis C. Schweinf. 

Maxillaria uncata Lindl. 

Maxillaria venezuelana (C. Schweinf.) Carnevali ined 

Maxillaria xylobiiflora Schltr. 

Mesadenella angustisegmenta Garay 

Mesadenella cuspidata (Lindl.) Garay 

Miltoniopsis santanaei Garay & Dunst. 

Mormodes atropurpurea Lindl. 

Mormodes buccinator Lindl. 

Mormodes carnevaliana Salazar & G.A. Romero 

Mormodes cucumerina Pabst 

Mormodes igneum Lindl. & Paxton 

Mormodes orinocoensis Salazar & G.A. Romero 

Mormodes vernixioidea Pabst 

Mormodes vernixioidea subsp. autanensis G.A. Romero & Salazar 

Mormodes vernixium Rchb. f. 

Myoxanthus octomeriae (Schltr.) Luer 

Myoxanthus parahybunensis (Barb. Rodr.) Luer 

Myoxanthus parvilabius (C. Schweinf.) Luer 

Myoxanthus reymondii (H. Karst.) Luer 

Myoxanthus simplicicaulis (C. Schweinf.) Luer 

Myoxanthus speciosus (Luer) Luer 

Myoxanthus trachychlamys (Schltr.) Luer 

Nidema ottonis (Rchb. f.) Britton & Millsp. 

Notylia angustifolia Cogn. 

Notylia aromatica Barker ex Lindl. 

Notylia bungerothii Rchb. f. 

Notylia durandiana Cogn. 

Notylia fragrans H. Focke 

Notylia laxa Rchb. f. 

Notylia longispicata Hoehne & Schltr. 

Notylia micrantha Lindl. 

Notylia microchila Cogn. 

AM BO GU SU 

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AM BO GU 

AM BO 

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AM BO 

AM BO 

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AM GU 

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BO GU SU FG 


AM GU 


AM BO GU FG 

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AM BO 

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AM GU SU FG 

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GU 

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128 

Notylia pentachne Rchb. f. 

Notylia peruviana (Schltr.) C. Schweinf. 

Notylia platyglossa Schltr. 

Notylia rhombilabia C. Schweinf. 

Notylia sagittifera (Kunth) Link, Klotzsch & Otto 

Notylia yauaperyensis Barb. Rodr. 

Octomeria amazonica Pabst 

Octomeria anomala Garay & Dunst. 

Octomeria apiculata (Lindl.) Garay & H.R. Sweet 

Octomeria connellii Rolfe 

Octomeria cordilabia C. Schweinf. 

Octomeria dentifera C. Schweinf. 

Octomeria erosilabia C. Schweinf. 

Octomeria exigua C. Schweinf. 

Octomeria filifolia C. Schweinf. 

Octomeria flaviflora C. Schweinf. 

Octomeria gemmula Carnevali & I. Ramírez 

Octomeria graminifolia R. Br. 

Octomeria heleneana Carnevali & Delascio 

Octomeria integrilabia C. Schweinf. 

Octomeria lancipetala C. Schweinf. 

Octomeria minor C. Schweinf. 

Octomeria monticola C. Schweinf. 

Octomeria nana C. Schweinf. 

Octomeria oxychela Barb. Rodr. 

Octomeria parvifolia Rolfe 

Octomeria parvula C. Schweinf. 

Octomeria petulans Rchb. f. 

Octomeria rhizomatosa C. Schweinf. 

Octomeria romerorum Carnevali & I. Ramírez 

Octomeria sarthouae Luer 

Octomeria scirpoidea (Poepp. & Endl.) Rchb. f. 

Octomeria sp. A 

Octomeria spathulata Rchb. f. 

Octomeria splendida Garay & Dunst. 

Octomeria steyermarkii Garay & Dunst. 

Octomeria surinamensis H. Focke 

Octomeria tapiricataractae G.A. Romero & C.A. Luer 

Octomeria taracuana Schltr. 

Octomeria yauaperyensis Barb. Rodr. 

Odontoglossum naevium Lindl. 

Oeceoclades maculata (Lindl.) Lindl. 

Oliveriana egregia Rchb. f. 

Oncidium baueri Lindl. 

Oncidium boothianum Rchb. f. 

Oncidium citrinum Lindl. 

Oncidium dactyliferum Garay & Dunst. 

Oncidium heteranthum Poepp. & Endl. 

Oncidium luridum Lindl. 

Oncidium nigratum Lindl. 

Oncidium orthostates Ridl. ex Thurn 

Oncidium pirarense Rchb. f. 

Oncidium sphacelatum Lindl. 

Oncidium tenellum F. Gérard 

Oncidium warmingii Rchb. f. 

Ondidium pardalis Rchb. f. 

Ophidion pleurothallopsis (Kraenzl.) Luer 

Orleanesia amazonica Barb. Rodr. 

Orleanesia maculata Garay 

Orleanesia yauaperyensis (Schltr.) Barb. Rodr. 

Ornithocephalus bicornis Lindl. 

Ornithocephalus bonplandii Rchb. f. 

Ornithocephalus brachyceras G.A. Romero & Carnevali 

Ornithocephalus ciliatus Lindl. 


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Ornithocephalus dasyrhizus Kraenzl. 

Ornithocephalus falcatus H. Focke 

Ornithocephalus gladiatus Hook. 

Ornithocephalus kruegeri Rchb. f. 

Otoglossum arminii (Rchb. f.) Garay & Dunst. 

Otoglossum globuliferum (Kunth) N.H. Williams & M.W. Chase 

Otoglossum scansor (Rchb. f.) Carnevali & I. Ramirez 

Otostylis brachystalix (Rchb. f.) Schltr. 

Otostylis lepida (Linden & Rchb. f.) Schltr. 

Palmorchis guianensis (Schltr.) C. Schweinf. & Correll 

Palmorchis kuhlmannii (Schltr.) L.O. Williams 

Palmorchis lobulata (Mansf.) C. Schweinf. & Correll 

Palmorchis pabstii Veyret 

Palmorchis prospectorum Veyret 

Palmorchis puber (Cogn.) Garay 

Palmorchis pubescens Barb. Rodr. 

Palmorchis trilobulata L.O. Williams 

Paphinia cristata (Lindl.) Lindl. 

Paphinia dunstervillei Dodson & G.A. Romero 

Paphinia lindeniana Rchb. f. 

Pelexia callifera (C. Schweinf.) Garay 

Pelexia delicatula Szlach. & Veyret 

Pelexia goninensis (Pulle) Schltr. 

Pelexia laxa (Poepp. & Endl.) Lindl. 

Peristeria aspersa Rolfe 

Peristeria cerina Lindl. 

Peristeria pendula Hook. 

Peristeria rossiana Rchb. f. 

Peristeria selligera Rchb. f. 

Phragmipedium caudatum (Lindl.) Rolfe 

Phragmipedium klotzschianum (Rchb. f.) Rolfe 

Phragmipedium lindleyanum (R.H. Schomb. ex Lindl.) Rolfe 

Pinelianthe alticola (Garay & Dunst.) S. Rauschert 

Platystele johnstonii (Ames) Garay 

Platystele ovalifolia (H. Focke) Garay & Dunst. 

Platystele oxyglossa (Schltr.) Garay 

Platystele stenostachya (Rchb. f.) Garay 

Platythelys maculata (Hook.) Garay 

Plectrophora cultrifolia (Barb. Rodr.) Cogn. 

Plectrophora iridifolia H. Focke 

Pleurothallis aondae Carnevali & G.A. Romero 

Pleurothallis arachnopsis H.G. Jones 

Pleurothallis archidiaconi Ames 

Pleurothallis arenicola (Carnevali & I. Ramírez) Carnevali & I. Ramírez 

Pleurothallis aristata Hook. 

Pleurothallis aspasicensis Rchb. f. 

Pleurothallis barboselloides Schltr. 

Pleurothallis barbulata Lindl. 

Pleurothallis barthelemyi Luer 

Pleurothallis biglandulosa Schltr. 

Pleurothallis brevipes H. Focke 

Pleurothallis cabellensis Rchb. f. 

Pleurothallis callifera C. Schweinf. 

Pleurothallis ciliolata Schltr. 

Pleurothallis coriacardia Rchb. f. 

Pleurothallis corniculata Lindl. 

Pleurothallis curvifructa H.G. Jones [possibly =P. archidiaconi] 

Pleurothallis deborana Carnevali & I. Ramírez 

Pleurothallis determannii Luer 

Pleurothallis discoidea Lindl. 

Pleurothallis discophylla Luer & Carnevali 

Pleurothallis elvirana Carnevali & I. Ramírez 

Pleurothallis ephemera Lindl. 

Pleurothallis erebatensis Carnevali & G.A. Romero 

FG? 

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130 

Pleurothallis erinacea Rchb. f. 

Pleurothallis exigua Cogn. 

Pleurothallis floribunda Poepp. & Endl. 

Pleurothallis fockei Lindl. 

Pleurothallis funerea (Barb. Rodr.) Cogn. 

Pleurothallis galeata Lindl. 

Pleurothallis garciae Luer 

Pleurothallis geographica Luer 

Pleurothallis glandulosa Ames 

Pleurothallis granitica Luer & G.A. Romero 

Pleurothallis grobyi Bateman ex Lindl. 

Pleurothallis hexandra Garay & Dunst. 

Pleurothallis hilariana Carnevali & G.A. Romero 

Pleurothallis hitchcockii Ames 

Pleurothallis holstii Carnevali & I. Ramírez 

Pleurothallis humilis C. Schweinf. 

Pleurothallis hymenantha Lindl. 

Pleurothallis imraei Lindl. 

Pleurothallis ionantha Rchb. f. 

Pleurothallis kerrii Braga 

Pleurothallis lanceana Lodd. 

Pleurothallis lancipetala (H. Karst.) Schltr. 

Pleurothallis lappiformis A.H. Heller & L.O. Williams 

Pleurothallis longisepala Barb. Rodr. 

Pleurothallis loranthophylla Rchb. f. 

Pleurothallis maguirei Luer 

Pleurothallis mentosa Cogn. 

Pleurothallis minima C. Schweinf. 

Pleurothallis miqueliana (H. Focke) Lindl. 

Pleurothallis monophylla (Hook.) Fawc. & Rendle 

Pleurothallis morilloi Carnevali & I. Ramírez 

Pleurothallis moritzii Rchb. f. [Mt. Roraima] 

Pleurothallis nanifolia Foldats 

Pleurothallis obovata (Lindl.) Lindl. 

Pleurothallis omissa Luer 

Pleurothallis omoglossa Luer 

Pleurothallis parvifolia Lindl. 

Pleurothallis pemonum Carnevali & I. Ramírez 

Pleurothallis phoenicoptera Carnevali & G.A. Romero 

Pleurothallis picta Lindl. 

Pleurothallis polygonoides Griseb. 

Pleurothallis prolifera Herb. ex Lindl. 

Pleurothallis pruinosa Lindl. 

Pleurothallis pubescens Lindl. 

Pleurothallis revoluta (Ruiz & Pav.) Garay 

Pleurothallis rhombipetala Rolfe 

Pleurothallis ruscifolia (Jacq.) R. Br. 

Pleurothallis samacensis Ames 

Pleurothallis sandaliorum G.A. Romero & Carnevali 

Pleurothallis sarcosepala Carnevali & I. Ramírez 

Pleurothallis sclerophylla Lindl. 

Pleurothallis secunda Poepp. & Endl. 

Pleurothallis semperflorens Lindl. 

Pleurothallis seriata Lindl. 

Pleurothallis sp. A 

Pleurothallis spiculifera Lindl. 

Pleurothallis steinbuchiae Carnevali & G.A. Romero 

Pleurothallis stenocardium Schltr. 

Pleurothallis suspensa Luer 

Pleurothallis tepuiensis Carnevali & I. Ramírez 

Pleurothallis testifolia (Sw.) Lindl. 

Pleurothallis tridentata Klotzsch 

Pleurothallis trinitensis (Griseb.) Carnevali & G.A. Romero 

Pleurothallis uniflora Lindl. 


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GU 

AM BO 

AM BO GU SU 

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Pleurothallis velaticaulis Rchb. f. 

Pleurothallis wilsonii Lindl. 

Pleurothallis yauaperyensis Barb. Rodr. 

Pleurothallis zephyrina Rchb. f. 

Polycycnis surinamensis C. Schweinf. 

Polyotidium huebneri (Mansf.) Garay 

Polystachya amazonica Schltr. 

Polystachya cavanayensis Garay & Dunst. 

Polystachya concreta (Jacq.) Garay & H.R. Sweet 

Polystachya d’agremondiana Kraenzl. [dubious species] 

Polystachya foliosa (Lindl.) Rchb. f. 

Ponthieva diptera Linden & Rchb. f. 

Ponthieva ovatilabia C. Schweinf. 

Prescottia auyantepuiensis Carnevali & G.A. Romero 

Prescottia carnosa C. Schweinf. 

Prescottia oligantha (Sw.) Lindl. 

Prescottia stachyodes (Sw.) Lindl. 

Prescottia tepuyensis Carnevali & C.A. Vargas 

Prosthechea aemula (Lindl.) W.E. Higgins 

Prosthechea calamaria (Lindl.) W.E. Higgins 

Prosthechea crassilabia (Poepp. & Endl.) Carnevali & I. Ramírez 

Prosthechea fragrans (Sw.) W.E Higgins [in Guianas records may 

=Prosthechea aemula] 

Prosthechea jauana (Carnevali & I. Ramírez) W.E. Higgins 

Prosthechea pygmaea (Hook.) W.E. Higgins 

Prosthechea tigrina (Linden ex Lindl.) W.E. Higgins 

Prosthechea vespa (Vell.) W.E. Higgins 

Psilochilus dusenianus Kraenzl. ex Garay & Dunst. 

Psilochilus macrophyllus (Lindl.) Ames 

Psilochilus physurifolius (Rchb. f.) Lojtnant 

Psychopsis kramerianum (Rchb. f.) H.G. Jones 

Psychopsis papilio (Lindl.) H.G. Jones 

Psychopsis versteegianum (Pulle) Lückel & Braem 

Pterichis galeata Lindl. 

Quekettia microscopica Lindl. 

Quekettia papillosa Garay 

Quekettia vermeuleniana Determann 

Reichenbachanthus reflexus (Lindl.) Brade 

Restrepiopsis tubulosa (Lindl.) Luer 

Rodriguezia batemanii Poepp. & Endl. 

Rodriguezia candida (Bateman ex Lindl.) Christenson 

Rodriguezia cinnabarina H.G. Jones 

Rodriguezia flavida Garay & Dunst. 

Rodriguezia lanceolata Ruiz & Pav. 

Rodriguezia leeana Rchb. f. 

Rodriguezia luteola N.E. Br. 

Rudolfiella aurantiaca (Lindl.) Hoehne 

Rudolfiella bicornaria (Rchb. f.) Hoehne 

Sacoila hassleri (Cogn.) Garay 

Sacoila lanceolata (Aubl.) Garay 

Sarcoglottis acaulis (Sm.) Schltr. 

Sarcoglottis amazonica Pabst 

Sarcoglottis aphylla (Ridl. ex Thurn) Schltr. 

Sarcoglottis degranvillei Szlach. & Veyret 

Sarcoglottis grandiflora (Lindl.) Klotzsch 

Sarcoglottis metallica (Rolfe) Schltr. 

Sarcoglottis rupestris (Lindl.) Barb. Rodr. 

Sarcoglottis sagittata (Rchb. f. & Warm.) Schltr. 

Sarcoglottis simplex (Griseb.) Schltr. 

Sarcoglottis stergiosii Carnevali & I. Ramírez 

Scaphosepalum breve (Rchb. f.) Rolfe 

Scaphyglottis bifida (Rchb. f.) C. Schweinf. 

Scaphyglottis boliviensis (Rolfe) B.R. Adams 

Scaphyglottis dunstervillei (Garay) Foldats 

AM 

AM 

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BO 

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AM BO SU FG 

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132 


Scaphyglottis felskyi (Rchb. f.) Schltr. 

Scaphyglottis fusiformis (Griseb.) R.E. Schult. 

Scaphyglottis graminifolia (Ruiz & Pav.) Poepp. & Endl. 

Scaphyglottis grandiflora Ames & C. Schweinf. 

Scaphyglottis leucantha Rchb. f. 

Scaphyglottis lindeniana (A. Rich. & Galeotti) L.O. Williams 

Scaphyglottis michelangeliorum Carnevali & Steyerm. 

Scaphyglottis modesta (Rchb. f.) Schltr. 

Scaphyglottis ochroleuca Schltr. 

Scaphyglottis prolifera Cogn. 

Scaphyglottis propinqua C. Schweinf. 

Scaphyglottis sessilis (Rchb. f.) Foldats 

Scaphyglottis sickii Pabst 

Scaphyglottis stellata Lodd. ex Lindl. 

Scelochilus ecalcaratus Determann 

Scelochilus ottonis Klotzsch 

Scelochilus paraguaensis Garay & Dunst. 

Schlimia alpina Rchb. f. & Warsz. 

Schomburgkia crispa Lindl. [records in Guianas may =S. marginata] 

Schomburgkia gloriosa Rchb. f. 

Schomburgkia heidii Carnevali 

Schomburgkia marginata Lindl. 

Schomburgkia rosea Linden ex Lindl. 

Schomburgkia undulata Lindl. 

Scuticaria hadwenii hort. ex Planch. 

Scuticaria hadwenii hort. var. dodgsoni (B.S. Wms.) Stein 

Scuticaria steelei (Hook.) Lindl. 

Selenipedium palmifolium (Lindl.) Rchb. f. 

Selenipedium steyermarkii Foldats 

Sievekingia jenmanii Rchb. f. 

Sigmatostalix amazonica Schltr. 

Sigmatostalix huebneri Mansf. 

Sobralia candida (Poepp. & Endl.) Rchb. f. 

Sobralia crocea (Poepp. & Endl.) Rchb. f. 

Sobralia dichotoma Ruiz & Pav. 

Sobralia fimbriata Poepp. & Endl. 

Sobralia fragrans Lindl. 

Sobralia granitica G.A. Romero & Carnevali 

Sobralia infundibuligera Garay & Dunst. 

Sobralia liliastrum Lindl. 

Sobralia macrophylla Rchb. f. 

Sobralia oliva-estevae Carnevali & I. Ramírez 

Sobralia sessilis Lindl. 

Sobralia speciosa C. Schweinf. 

Sobralia stenophylla Lindl. 

Sobralia suaveolens Rchb. f. 

Sobralia valida Rolfe 

Sobralia violacea Linden ex Lindl. 

Solenidium lunatum (Lindl.) Kraenzl. 

Spiranthes tenuis Lindl. 

Spiranthes torta (Thunb.) Garay & H.R. Sweet 

Stanhopea candida Barb. Rodr. 

Stanhopea grandiflora (Lodd.) Lindl. 

Stanhopea wardii Lodd. ex Lindl. 

Stelis alata Lindl. 

Stelis aprica Lindl. 

Stelis argentata Lindl. 

Stelis aviceps Lindl. 

Stelis bangii Rolfe 

Stelis cucullata Ames 

Stelis cuprea Lindl. [dubious record] 

Stelis effusa Schltr. 

Stelis fendleri Lindl. 

Stelis fraterna Lindl. 

BO GU SU 



AM BO GU FG 

BO 

GU SU 

AM 


SU FG 

BO GU SU FG 

SU 

AM BO GU 



FG 

AM 

BO 

GU 

GU? SU? FG? 

GU SU? 

AM BO 

BO GU SU FG 

GU 

BO 

GU 

GU 


BO GU SU FG 

BO GU 


AM SU FG 

AM 

BO 

SU? 

AM 

AM BO SU FG 


AM BO 

AM BO DA GU 



BO GU 


AM 

BO GU SU FG 

BO GU SU FG 

AM BO DA GU 

AM BO SU? 

AM BO GU 

GU SU FG 

FG? 

AM GU? 


BO 

AM BO GU 

GU SU 


AM BO GU 

AM BO GU 

AM BO 

FG 

BO 

BO 

AM BO GU


Stelis garayi (Dunst.) Carnevali & I. Ramírez 

Stelis grandiflora Lindl. 

Stelis guianensis Rolfe 

Stelis intermedia Poepp. & Endl. 

Stelis latisepala C. Schweinf. 

Stelis maxima Lindl. 

Stelis minimiflora Schltr. 

Stelis muscifera Lindl. 

Stelis obovata C. Schweinf. 

Stelis ophioglossoides (Jacq.) Jacq. 

Stelis papaquerensis Rchb. f. 

Stelis perparva C. Schweinf. 

Stelis pusilla Kunth 

Stelis pygmaea Cogn. 

Stelis santiagoensis Mansf. 

Stelis schomburgkii Fawc. & Rendle 

Stelis tolimensis Schltr. 

Stelis tridentata Lindl. 

Stelis tristyla Lindl. 

Stelis zonata Rchb. f. [=Stelis muscifera?] 

Stellilabium alticolum Dodson & R. Escobar 

Stenia pallida Lindl. 

Stictophyllorchis pygmaea (Cogn.) Carnevali & Dodson 

Teuscheria dodsonii Dressler 

Teuscheria wageneri (Rchb. f.) Garay 

Trichocentrum carthagenense (Jacq.) M.W. Chase & N.H. Williams 

Trichocentrum cebolleta (Jacq.) M.W. Chase & N.H. Williams 

Trichocentrum fuscum Lindl. 

Trichocentrum lanceanum (Lindl.) M.W. Chase & N.H. Williams 

Trichocentrum nanum (Lindl.) M.W. Chase & N.H. Williams 

Trichocentrum purpureum Lindl. ex Rchb. f. 

Trichocentrum recurvum Lindl. 

Trichopilia aenigma Garay 

Trichopilia oicophylax Rchb. f. 

Trichopilia wageneri Rchb. f. 

Trichosalpinx cedralensis (Ames) Luer 

Trichosalpinx ciliaris (Lindl.) Luer 

Trichosalpinx dura (Lindl.) Luer 

Trichosalpinx egleri (Pabst) Luer 

Trichosalpinx intricata (Lindl.) Luer 

Trichosalpinx memor (Rchb. f.) Luer 

Trichosalpinx orbicularis (Lindl.) Luer 

Trichosalpinx oxychilos Carnevali & G.A. Romero 

Trichosalpinx patula Luer 

Trichosalpinx pumila (Luer) Luer 

Trichosalpinx pusilla (Kunth) Luer 

Trichosalpinx roraimensis (Rolfe) Luer 

Trichosalpinx steyermarkii Luer 

Trigonidium acuminatum Bateman ex Lindl. 

Trigonidium egertonianum Bateman ex Lindl. 

Trigonidium macranthum Barb. Rodr. 

Trigonidium obtusum Lindl. 

Triphora amazonica Schltr. 

Triphora surinamensis (Lindl. ex Benth.) Britton & P. Wilson 

Trisetella triglochin (Rchb. f.) Luer 

Trizeuxis falcata Lindl. 

Uleiorchis liesneri Carnevali & I. Ramírez 

Uleiorchis ulaei (Cogn.) Handro 

Vanilla acuta Rolfe 

Vanilla appendiculata Rolfe 

Vanilla barrereana Veyret & Szlach. 

Vanilla bicolor Lindl. 

Vanilla chamissonis Klotzsch 

Vanilla cristato-callosa Hoehne 

AM BO 

BO 

GU 

GU FG 

BO 

BO 

GU SU 

AM 

BO 

GU 

AM BO 

BO GU SU? FG 

BO 

GU 

SU? FG 

BO FG 

AM BO DA FG 

AM BO GU 

AM 

AM 

BO 

GU SU 

BO 

BO 

GU 

BO DA GU 

AM BO 

SU? 

BO 

AM 


BO SU FG 


AM BO GU 

GU 

FG 

BO 

BO 

GU SU 

GU SU 

AM BO 

GU SU FG 

AM BO SU 

BO 

BO 

DA GU SU FG 

AM BO GU SU 

AM 

AM 

BO 

BO 

BO 

BO 

GU SU FG 

AM BO 

BO 

GU 

AM BO DA GU 

GU 

GU 

SU FG 


GU SU? FG 


AM SU FG 

AM 

BO DA GU 

AM BO GU FG 

GU 

GU 

SU FG 

FG 

GU 

GU 

SU FG 

FG

134 


Vanilla fimbriata Rolfe 

Vanilla gardneri Rolfe 

Vanilla grandiflora Lindl. 

Vanilla grandifolia Lindl. 

Vanilla guianensis Splitg. [name of uncertain application; =V. mexicana? 

V. grandiflora?] 

Vanilla hostmanni Rolfe 

Vanilla latisegmenta Ames & C. Schweinf. 

Vanilla leprieurii PortPres 

Vanilla marowynensis Pulle 

Vanilla mexicana Mill. 

Vanilla odorata C. Presl 

Vanilla ovata Rolfe 

Vanilla palmarum (Salzm. ex Lindl.) Lindl. 

Vanilla penicillata Garay & Dunst. 

Vanilla planifolia Andrews 

Vanilla porteresiana Veyret & Szlach. 

Vanilla surinamensis Rchb. f. 

Vanilla wrightii Rchb. f. 

Vargasiella venezuelana C. Schweinf. 

Wullschlaegelia calcarata Benth. 

Xerorchis amazonica Schltr. 

Xerorchis trichorhiza (Kraenzl.) Garay 

Xylobium chapadense (Barb. Rodr.) Cogn. 

Xylobium colleyi (Bateman ex Lindl.) Rolfe 

Xylobium foveatum (Lindl.) G. Nicholson 

Xylobium pallidiflorum (Hook.) G. Nicholson 

Xylobium palmifolium (Sw.) Benth. ex Fawc. 

Xylobium variegatum (Ruiz & Pav.) Garay & Dunst. 

Zygosepalum angustilabium (C. Schweinf.) Garay 

Zygosepalum kegelii (Rchb. f.) Rchb. f. 

Zygosepalum labiosum (Rich.) Garay 

Zygosepalum lindeniae (Rolfe) Garay & Dunst. 

Zygosepalum tatei (Ames & C. Schweinf.) Garay & Dunst. 

Zygostates sp. nov. 

AM BO 

AM BO 


Acianthera fockei (Lindl.) Pridgeon & M.W. Chase =Pleurothallis fockei 

Acianthera garciae (Luer) Pridgeon & M.W. Chase =Pleurothallis garciae 

Acianthera lanceana Pridgeon & M.W. Chase =Pleurothallis lanceana 

Acianthera miqueliana (H. Focke) Pridgeon & M.W. Chase =Pleurothallis miqueliana 

Acianthera omissa (Luer) Pridgeon & M.W. Chase =Pleurothallis omissa 

Acianthera prolifera (Herb. ex Lindl.) Pridgeon & M.W. Chase =Pleurothallis prolifera 

Acianthera wilsonii (Lindl.) Pridgeon & M.W. Chase =Pleurothallis wilsonii 

Acianthera yauaperyensis (Barb. Rodr.) Pridgeon & M.W. Chase =Pleurothallis yauaperyensis 

Acoidium fuscum Lindl. [nomen nudum] =Trichocentrum fuscum 

Adeneleutherophora graminifolia Barb. Rodr. =Elleanthus graminifolius 

Adipe longicornis (Lindl.) M. Wolfe =Bifrenaria longicornis 

Aeranthus pachyrrhizus Rchb. f. =Campylocentrum pachyrrhizum 

Aganisia alba Ridl. ex Thurn =Otostylis lepida 

Aganisia brachypoda Schltr. =Aganisia pulchella 

Aganisia cyanea (Lindl.) Rchb. f. =Acacallis cyanea 

Aganisia fimbriata Rchb. f. =Acacallis fimbriata 

Aganisia lepida Linden & Rchb. f. =Otostylis lepida 

Aganisia venusta (Ridl. ex Oliv.) Rolfe =Otostylis lepida 

Anacheilium calamarium (Lindl.) Pabst, Moutinho & Pinto =Prosthechea calamaria 

Anathallis barbulata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis barbulata 

Anathallis brevipes (H. Focke) Pridgeon & M.W. Chase =Pleurothallis brevipes 

Anathallis maguirei (Luer) Pridgeon & M.W. Chase =Pleurothallis maguirei 

Anathallis sclerophylla (Lindl.) Pridgeon & M.W. Chase =Pleurothallis sclerophylla 

Angraecum fasciola Lindl. =Campylocentrum fasciola 

Angraecum lansbergii Rchb. f. =Campylocentrum lansbergii 

Angraecum maculatum Lindl. =Oeceoclades maculata 

Angraecum micranthum Lindl. =Campylocentrum micranthum 

GU 

GU SU FG? 

SU 

SU 

GU 

FG 

AM BO DA GU 

SU 

SU FG 

AM BO FG 

GU? FG 


AM BO GU 

AM BO GU SU 

SU 

FG 

AM BO 

BO 

GU SU FG 


AM BO SU 

BO GU 

GU 

FG 

BO GU SU FG 

GU SU FG 

BO 

GU 

SU 

SU FG 

BO GU 

SU 


AM GU FG 

AM BO GU 

SU


Angraecum poeppigii Rchb. f. =Campylocentrum poeppigii 

Anthereon mentosus (Cogn.) Pridgeon & M.W. Chase =Pleurothallis mentosa 

Apatostelis alata (Lindl.) Garay =Stelis alata 

Apatostelis garayi Dunst. =Stelis garayi 

Apatostelis minimiflora (Schltr.) Garay =Stelis minimiflora 

Aspegrenia scirpoidea Poepp. & Endl. =Octomeria scirpoidea 

Aulizia ciliaris (L.) Salisb. =Epidendrum ciliare 

Barbosella kegelii (Rchb. f.) Schltr. =Pleurothallis hexandra 

Baskervilla venezuelana Garay & Dunst. [not in flora area] 

Beadlea bifida (Ridl.) Garay & Dunst. =Helonoma chiropterae 

Beadlea bifida (Ridl. ex Oliv.) Garay & Dunst. =Helonoma bifida 

Beadlea cranichoides (Griseb.) Small =Cyclopogon cranichoides 

Beadlea elata (Sw.) Small ex Britton =Cyclopogon elatus 

Beloglottis americana (C. Schweinf. & Garay) Szlach. =Helonoma americana 

Beloglottis bifida (Ridl. ex Oliv.) Szlach. =Helonoma bifida 

Beloglottis chiropterae Szlach. =Helonoma chiropterae 

Bifrenaria aurantiaca Lindl. =Rudolfiella aurantiaca 

Bifrenaria bicornaria Rchb. f. =Rudolfiella bicornaria 

Bifrenaria maguirei C. Schweinf. =Guanchezia maguirei 

Bifrenaria minuta Garay =Hylaeorchis petiolaris 

Bifrenaria rudolfii (Hoehne) Carnevali & G.A. Romero =Hylaeorchis petiolaris 

Bifrenaria sabulosa Barb. Rodr. =Bifrenaria longicornis 

Bifrenaria wageneri Rchb. f. =Teuscheria wageneri 

Bletia verecunda R. Br. =Bletia purpurea 

Bolbophyllaria oerstedii Rchb. f. =Bulbophyllum oerstedii 

Bollea cardonae Schnee =Bollea hemixantha 

Bollea guyanensis Klotzsch [nomen nudum] 

Bonatea macilenta (Lindl.) Rchb. f. =Habenaria macilenta 

Bonatea pauciflora Lindl. =Habenaria trifida 

Bonatea pratensis Lindl. =Habenaria pratensis 

Brassavola angustata Lindl. =Brassavola martiana 

Brassavola surinamensis H. Focke =Brassavola martiana 

Brassia attenuata H.G. Jones =Brassia chloroleuca 

Brassia cochleata Knowles & Westc. =Brassia lawrenceana 

Brassia verrucosa auct., non Lindl. =Brassia spp. 

Bulbophyllum correae Pabst =Bulbophyllum setigerum 

Bulbophyllum pachyrrhachis auct. non (A. Rich.) Griseb. 1864, sensu Dunst. & Garay =Bulbophyllum oerstedii 

Burlingtonia candida Bateman ex Lindl. =Rodriguezia candida 

Camaridium album (Hook.) Hoehne =Maxillaria alba 

Camaridium amazonicum Schltr. =Maxillaria camaridii 

Camaridium equitans Schltr. =Maxillaria equitans 

Camaridium iguapensis (Hoehne & Schltr.) Hoehne =Maxillaria imbricata 

Camaridium imbricatum (Barb. Rodr.) Hoehne =Maxillaria imbricata 

Camaridium ochroleucum Lindl. =Maxillaria camaridii 

Camaridium pendulum Barb. Rodr. =Maxillaria rigida 

Camaridium squamatum (Barb. Rodr.) Hoehne =Maxillaria squamata 

Camaridium uncatum (Lindl.) Hoehne =Maxillaria uncata 

Camaridium vandiforme Schltr. =Maxillaria equitans 

Campylocentrum burchellii Cogn. [not in flora area] 

Campylocentrum micranthum auct. non (Lindl.) Rolfe 1903: sensu Foldats in Lasser =Campylocentrum panamense p.p., C. 

lansbergii p.p. 

Campylocentrum steyermarkii Foldats =Campylocentrum hondurense 

Campylocentrum uroplectron Pabst =Campylocentrum huebneri 

Catasetum apertum Rolfe =Catasetum ×tapiriceps 

Catasetum bicallosum auct. non Cogn. 1894: sensu Dunst. & Garay =Catasetum parguazense 

Catasetum bungerothii N.E. Br. =Catasetum pileatum 

Catasetum chloranthum Cogn. =Catasetum planiceps 

Catasetum comosum Cogn. =Catasetum barbatum 

Catasetum discolor var. roseo-album (Hook.) Mansf. =Catasetum x roseo-album 

Catasetum fuliginosum Lindl. ? =Catasetum callosum var. carunculatum 

Catasetum gongoroides Kraenzl. =Catasetum bicolor 

Catasetum recurvatum Link =Catasetum planiceps 

Catasetum scurra Rchb. f. =Clowesia warczewiczii 

Catasetum splendens Cogn. =Catasetum ×tapiriceps

136 


Catasetum stenoglossum Pabst =Catasetum bicallosum 

Catasetum warczewiczii Lindl. & Paxton =Clowesia warczewiczii 

Cattleya liliastrum (Lindl.) Beer =Sobralia liliastrum 

Cattleya pumila R.H. Schomb. =Cattleya lawrenceana 

Cattleya schomburgkii Lodd. ex Lindl. =Cattleya violacea 

Cattleya superba R.H. Schomb. ex Lindl. =Cattleya violacea 

Cattleya violacea (Linden ex Lindl.) Beer =Sobralia violacea 

Caularthron indivisum (Bradf. ex Griseb.) Garay & Dunst =Caularthron bilamellatum f. indivisa 

Centrogenium calcaratum (Sw.) Schltr. =Eltroplectris calcarata 

Centrogenium setaceum (Lindl.) Schltr. =Eltroplectris calcarata 

Centroglossa peruviana Cogn. =Cryptocentrum peruvianum 

Ceratochilus grandiflorus Lodd. =Stanhopea grandiflora 

Chaenanthe barkeri Lindl. =Diadenium barkeri 

Chamelophyton kegelii (Rchb. f.) Garay =Pleurothallis hexandra 

Chaubardia tigrina Garay & Dunst. =Chaubardiella tigrina 

Cheiradenia imthurnii Cogn. =Cheiradenia cuspidata 

Cladobium violaceum Lindl. =Scaphyglottis graminifolia 

Cleistes moritzii (Rchb. f.) Garay & Dunst. =Cleistes lepida 

Cochleanthes amazonica (Rchb. f.) Schult. & Garay [not in flora area] 

Cohniella cebolleta (Jacq.) Christenson =Trichocentrum cebolleta 

Coryanthes speciosa var. punctata (Beer) G. Gerlach =Coryanthes punctata 

Cranichis alfredi Schltr. =Cranichis diphylla 

Cranichis guatemalensis Schltr. =Cranichis diphylla 

Cranichis monophylla Lindl. =Cranichis diphylla 

Cranichis nigrescens Schltr. =Cranichis diphylla 

Cranichis oligantha Sw. =Prescottia oligantha 

Cranichis ovatilabia Schltr. =Cranichis diphylla 

Cranichis stachyodes Sw. =Prescottia stachyodes 

Cranichis stictophylla Schltr. =Cranichis diphylla 

Cryptarrhena acrensis Schltr. =Cryptarrhena guatemalensis 

Cryptarrhena ghillanyi Pabst =Cryptarrhena guatemalensis 

Cryptarrhena lunata auct., non R. Br. =Cryptarrhena kegelii 

Cryptocentrum gracillimum auct. non Ames & C. Schweinf. 1925: sensu Dunst. & Garay =Cryptocentrum dunstervilleorum 

Cryptocentrum hoppii Schltr. =Cryptocentrum peruvianum subsp. peruvianum 

Cryptophoranthus pleurothallopsis Kraenzl. =Ophidion pleurothallopsis 

Cyathoglottis candida Poepp. & Endl. =Sobralia candida 

Cyclopogon bifidus (Ridl. ex Oliv.) Schltr. =Helonoma bifida 

Cyclopogon stictophyllus Schltr. =Cyclopogon olivaceus 

Cycnoches cucullata Lindl. =Cycnoches loddigesii 

Cycnoches pescatorei Lindl. =Lueddemannia pescatorei 

Cycnoches ventricosum var. chlorochilon (Klotzsch) P. Allen =Cycnoches chlorochilon 

Cycnoches versicolor Rchb. f. =Cycnoches haagii 

Cymbidium andersonii Lamb. ex Andrews =Cyrtopodium andersonii 

Cymbidium cordigerum Kunth =Encyclia cordigera 

Cymbidium diurnum (Jacq.) Sw. =Encyclia diurna 

Cymbidium ochroleucum (Lindl.) Lindl. =Maxillaria camaridii 

Cymbidium violaceum Kunth =Cattleya violacea 

Cypripedium klotzschianum Rchb. f. =Phragmipedium klotzschianum 

Cypripedium lindleyanum R.H. Schomb. ex Lindl. =Phragmipedium lindleyanum 

Cypripedium palmifolium Lindl. =Selenipedium palmifolium 

Cypripedium schomburgkianum Klotzsch ex M.R. Schomb. =Phragmipedium klotzschianum 

Cyrtopera longifolia Rchb. f. =Eulophia alta 

Cyrtopodium broadwayi Ames =Cyrtopodium parviflorum 

Cyrtopodium eburnea Barb. Rodr. =Koellensteinia tricolor 

Cyrtopodium flavescens Cogn. =Cyrtopodium andersonii 

Cyrtopodium glutiniferum Raddi [dubious in Guianas] 

Cyrtopodium punctatum (L.) Lindl. [dubious in Guianas] 

Dendrobium album Hook. =Maxillaria alba 

Dendrobium cebolleta Jacq. =Trichocentrum cebolleta 

Diacrium amazonicum Schltr. =Caularthron bicornutum 

Diacrium bicornutum (Hook.) Benth. =Caularthron bicornutum 

Diacrium bilamellatum (Rchb. f.) Hemsl. =Caularthron bilamellatum 

Diacrium venezuelanum Schltr. =Caularthron bilamellatum 

Dichaea ancoraelabia C. Schweinf. [not in flora area. Peruvian taxon]


Dichaea brachypoda auct. non Rchb. f 1866: sensu Foldats in Lasser =Dichaea camaridioides 

Dichaea brachypoda Rchb. f. [Costa Rican taxon, name widely misapplied] 

Dichaea echinocarpa (Sw.) Lindl. [nom. illeg.] =Dichaea pendula 

Dichaea graminoides (Sw.) Lindl. [name (mis?)applied to Dichaea rendlei in Guianas] 

Dichaea histrio auct. non Rchb. f. 1859: sensu Dunst. & Garay =Dichaea tachirensis 

Dichaea kegelii auct. non Rchb. f. 1877: sensu Dunst. and Garay =Dichaea splitgerberi 

Dichaea longa Schltr. =Dichaea latifolia var. longa 

Dichaea morrisii auct. non Fawc. & Rendle 1910: sensu Dunst. and Garay =Dichaea kegelii, p.p., Dichaea robusta, p.p. 

Dichaea muricata auct. non (Sw.) Lindl. 1833: sensu Dunst. and Garay =Dichaea latifolia var. longa 

Dichaea muricata auct. non (Sw.) Lindl. 1833: sensu J. Folsom =Dichaea robusta 

Dichaea muricata (Sw.) Lindl. [rejected name] 

Dichaea pendula var. swartzii C. Schweinf. [nom. illeg.] =Dichaea pendula 

Dichaea rendlei var. rendlei auct. non Gleason 1927: sensu Foldats in Lasser =Dichaea trinitensis 

Dichaea rendlei var. trinitensis auct. non (Gleason) Foldats 1959: sensu Dunst. and G =Dichaea hookeri 

Dichaea rendlei var. trinitensis (Gleason) Foldats =Dichaea trinitensis 

Dichaea swartzii (C. Schweinf.) Garay & H.R. Sweet =Dichaea pendula 

Dicrypta bicolor Paxton ex Planch. =Maxillaria discolor 

Dicrypta discolor Lodd. ex Lindl. =Maxillaria discolor 

Dicrypta longifolia Barb. Rodr. =Heterotaxis superflua 

Dicrypta villosa Barb. Rodr. =Heterotaxis villosa 

Didactyle exaltata (Lindl.) Lindl. =Bulbophyllum exaltatum 

Didactyle meridensis (Rchb. f.) Lindl. =Bulbophyllum meridense 

Dikylikostigma preusii Kraenzl. =Discyphus scopulariae 

Dimerandra isthmii Schltr. =Dimerandra elegans 

Dimerandra stenopetala auct., non (Hook.) Schltr. [name widely misapplied] =Dimerandra spp. 

Diothonea gratissima Rchb. f. =Epidendrum gratissimum 

Diothonea imbricata Lindl. =Hexisea imbricata 

Dipteranthus peruvianus Schltr. =Notylia peruviana 

Duboisia reymondii H. Karst. =Myoxanthus reymondii 

Duckeella adolphii auct. non Porto et Brade 1940: sensu Foldats in Lasser =Duckeella pauciflora 

Echinella aspasicensis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis aspasicensis 

Echinella lappiformis (A.H. Heller & L.O. Williams) Pridgeon & M. W. Chase =Pleurothallis lappiformis 

Echinosepala aspasicensis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis aspasicensis 

Echinosepala lappiformis (A.H. Heller & L.O. Williams) Pridgeon & M. W. Chase =Pleurothallis lappiformis 

Elleanthus attenuatus J.R. Johnst. =Elleanthus arpophyllostachys 

Elleanthus kermesinus auct. non (Lindl.) Rchb. f. 1862: sensu Foldats in Lasser =Elleanthus confusus 

Elleanthus linifolius auct., non C. Presl =Elleanthus graminifolius 

Elleanthus puber Cogn. =Palmorchis puber 

Elleanthus pusillus Schltr. =Elleanthus graminifolius 

Elleanthus strobilifer (Poepp. & Endl.) Rchb. f. [not in flora area] 

Elleanthus virgatus auct. non (Rchb. f.) C. Schweinf. 1938: sensu Dunst. & Garay =Elleanthus confusus 

Encyclia abbreviata auct., non (Schltr.) Dressler =Prosthechea calamaria 

Encyclia acuta auct. non Schltr. 1825: sensu Carnevali & I. Ramírez =Encyclia chloroleuca 

Encyclia aemula (Lindl.) Carnevali & I. Ramírez =Prosthechea aemula 

Encyclia amicta auct. non (Linden & Rchb.) Schltr. 1919: sensu Carnevali & I. Ra =Encyclia pilosa p.p., Encyclia 

conchaechila p.p. 

Encyclia atropurpurea Schltr. =Encyclia cordigera 

Encyclia calamaria (Lindl.) Pabst =Prosthechea calamaria 

Encyclia chloroleuca (Hook.) Schltr. =Encyclia chloroleuca 

Encyclia ciliaris (L.) Lemée =Epidendrum ciliare 

Encyclia crassilabia (Poepp. & Endl.) Dressler =Prosthechea crassilabia 

Encyclia fragrans subsp. aemula (Lindl.) Dressler & G.E. Pollard =Prosthechea aemula 

Encyclia jauana Carnevali & I. Ramírez =Prosthechea jauana 

Encyclia latipetala (C. Schweinf.) Pabst =Encyclia pachyantha 

Encyclia maravalensis Withner =Encyclia chloroleuca 

Encyclia oncidioides auct. non (Lindl.) Schltr. 1833: sensu Carnevali & I. Ramírez =Encyclia granitica 

Encyclia oncidioides (Lindl.) Schltr. [not in flora area] 

Encyclia palacea (Rchb. f.) Lemée [not in flora area] =Nidema boothii 

Encyclia pamplonense (Rchb. f.) Carnevali =Prosthechea tigrina 

Encyclia picta auct. non (Lindl.) Hoehne 1952: sensu Carnevali & I. Ramírez =Encyclia guianensis 

Encyclia pygmaea (Hook.) Dressler =Prosthechea pygmaea 

Encyclia recurvata auct. non Schltr. 1919: sensu Dunst. & Garay =Encyclia ivonae 

Encyclia recurvata Carnevali & I. Ramírez =Encyclia ivonae 

Encyclia tigrina (Linden ex Lindl.) Carnevali & I. Ramírez =Prosthechea tigrina 

Encyclia tumuc-humaciense (Veyret) Hágsater =Epidendrum tumuc-humaciense

138 


Encyclia vespa (Vell.) Dressler =Prosthechea vespa 

Encyclia wageneri (Klotzsch, Otto & Dietr.) Schltr. =Encyclia diurna 

Epidendropsis vincentina (Lindl.) Garay & Dunst. =Epidendrum vincentinum 

Epidendropsis violascens (Ridl. ex Thurn) Garay & Dunst. =Epidendrum violascens 

Epidendrum aemulum Lindl. =Prosthechea aemula 

Epidendrum alsum Ridl. [nom. superfl.] =Epidendrum alsum 

Epidendrum amazonicum Schltr. =Epidendrum coronatum 

Epidendrum amictum auct. non Linden & Rchb. f. 1855: sensu Dunst. & Garay =Encyclia leucantha 

Epidendrum atropurpureum auct. non Willd. 1805: sensu Dunst. & Garay =Encyclia cordigera 

Epidendrum bathyschistum Schltr. =Epidendrum cristatum 

Epidendrum bicornutum Hook. =Caularthron bicornutum 

Epidendrum bifidum Aubl. [not in flora area] 

Epidendrum bilamellatum Rchb. f. =Caularthron bilamellatum 

Epidendrum boothii (Lindl.) L.O. Williams [not in flora area] =Nidema boothii 

Epidendrum caespitosum Barb. Rodr. =Prosthechea pygmaea 

Epidendrum calamarium Lindl. =Prosthechea calamaria 

Epidendrum carnosum Lindl. =Epidendrum durum 

Epidendrum carthagenense Jacq. =Trichocentrum carthagenense 

Epidendrum caudatum L. =Brassia caudata 

Epidendrum chloranthum Lindl. =Encyclia chloroleuca 

Epidendrum chloroleucum Hook. =Encyclia chloroleuca 

Epidendrum clavatum Lindl. =Epidendrum purpurascens 

Epidendrum cochleatum L. =Encyclia cochleata 

Epidendrum conchaechilum Barb. Rodr. =Encyclia conchaechila 

Epidendrum concretum Jacq. =Polystachya concreta 

Epidendrum cordigerum (Kunth) Foldats =Encyclia cordigera 

Epidendrum coriifolium auct. non Lindl. 1851: sensu Dunst. & Garay =Epidendrum amazonicoriifolium 

Epidendrum coriifolium Lindl. [all material probably referable to E. imitans in flora area] 

Epidendrum crassifolium Lindl. =Epidendrum secundum 

Epidendrum crassilabium Poepp. & Endl. =Prosthechea crassilabia 

Epidendrum cucullatum L. =Brassavola cucullata 

Epidendrum decipiens auct., non Lindl. =Epidendrum ibaguense 

Epidendrum dendrobioides auct. non Thunb. 1818: sensu Dunst. & Garay =Epidendrum ulei 

Epidendrum dendrobioides Thunb. [not in flora area] 

Epidendrum dichotomum C. Presl =Epidendrum secundum 

Epidendrum diurnum (Jacq.) Rchb. f. =Encyclia diurna 

Epidendrum diurnum auct. non (Jacq.) Rchb. f. 1866 [1867]: sensu Foldats in Lasser =Encyclia spp. 

Epidendrum diurnum (Jacq.) Cogn. [nom. illeg.] =Encyclia diurna 

Epidendrum duckei Porto & Brade =Epidendrum hombersleyi 

Epidendrum durum auct. non Lindl. 1841: sensu Foldats in Lasser =Epidendrum alsum p.p.; Epidendrum ulei p.p. 

Epidendrum echinocarpum Sw. [nom. superfl.] =Dichaea pendula 

Epidendrum ellipticum Graham ex Hook. =Epidendrum secundum 

Epidendrum elongatum Jacq. =Epidendrum secundum 

Epidendrum ernstii Schltr. =Epidendrum klotzscheanum 

Epidendrum fastigiatum auct. non Lindl. 1853: sensu Dunst. & Garay =Epidendrum unguiculatum 

Epidendrum flabelliformis Sw. =Cochleanthes flabelliformis 

Epidendrum floribundum Kunth =Epidendrum densiflorum 

Epidendrum florijugum Barb. Rodr. =Epidendrum sculptum 

Epidendrum fragrans auct. non Sw. 1788: sensu Foldats in Lasser =Prosthechea aemula 

Epidendrum fragrans Sw. [Guianas records =Prosthechea aemula] =Prosthechea fragrans 

Epidendrum fragrans var. aemulum (Lindl.) Barb. Rodr. =Prosthechea aemula 

Epidendrum frigidum var. stenophyton auct. non (Schltr.) C. Schweinf. 1944: sensu Foldats =Epidendrum imthurnii 

Epidendrum fulgens Brongn. ? =Epidendrum ibaguense 

Epidendrum fuscatum Sm. =Epidendrum anceps 

Epidendrum globosum Jacq. =Jacquiniella globosa 

Epidendrum graniticum Bateman ex Lindl. =Encyclia granitica 

Epidendrum huebneri auct. non Schltr. 1925: sensu Dunst. & Garay =Epidendrum orchidiflorum 

Epidendrum huebneri Schltr. =Epidendrum myrmecophorum 

Epidendrum ibaguense var. confluens (Lindl.) C. Schweinf. =Epidendrum calanthum 

Epidendrum ibaguense var. schomburgkii (Lindl.) C. Schweinf. =Epidendrum macrocarpum 

Epidendrum imatophyllum Lindl. =Epidendrum flexuosum 

Epidendrum imbricatum Lam. =Lockhartia imbricata 

Epidendrum imitans auct. non Schltr. 1921: sensu G.A. Romero & Carnevali =Epidendrum amazonicoriifolium 

Epidendrum incisum Vell. =Epidendrum macrocarpum


Epidendrum inconstans Ames =Epidendrum secundum 

Epidendrum indivisum Bradf. ex Griseb. =Caularthron bilamellatum f. indivisa 

Epidendrum ionosmum Lindl. =Encyclia ionosma 

Epidendrum labiatum Sw. =Leochilus labiatus 

Epidendrum labiosum Rich. =Zygosepalum labiosum 

Epidendrum latifolium (Lindl.) Garay & H.R. Sweet =Epidendrum carpophorum 

Epidendrum latipetalum C. Schweinf. =Encyclia pachyantha 

Epidendrum laxum Poepp. & Endl. =Epidendrum compressum 

Epidendrum lechleri auct. non Rchb. f. 1876: sensu Foldats in Lasser =Epidendrum chimantense 

Epidendrum leucanthum (Schltr.) Schnee =Encyclia leucantha 

Epidendrum leucanthum (Schltr.) C. Schweinf. =Encyclia leucantha 

Epidendrum lindenii Lindl. =Epidendrum secundum 

Epidendrum lineare Jacq. =Isochilus linearis 

Epidendrum maculatum Aubl. =Oncidium luridum 

Epidendrum miersii Lindl. =Epidendrum saxatile 

Epidendrum moyobambae Kraenzl. =Epidendrum coronatum 

Epidendrum musciferum Lindl. =Epidendrum anceps 

Epidendrum nocturnum var. latifolium Lindl. =Epidendrum carpophorum 

Epidendrum nocturnum var. minus Cogn. =Epidendrum longicolle 

Epidendrum nocturnum var. tumuc-humacience Veyret =Epidendrum tumuc-humaciense 

Epidendrum odoratissimum Lindl. =Encyclia odoratissima 

Epidendrum oncidioides auct. non Lindl. 1833: sensu Dunst. & Garay =Encyclia auyantepuiensis 

Epidendrum oncidioides Lindl. =Encyclia oncidioides 

Epidendrum ophioglossoides Jacq. =Stelis ophioglossoides 

Epidendrum ottonis Rchb. f. =Nidema ottonis 

Epidendrum pachyanthum Lindl. =Encyclia pachyantha 

Epidendrum palmarum Salzm. ex Lindl. =Vanilla palmarum 

Epidendrum pamplonense Rchb. f. =Prosthechea tigrina 

Epidendrum paniculatum auct. non Ruiz & Pav. 1798: sensu Foldats in Lasser =Epidendrum unguiculatum p.p., E. 

densiflorum p.p. 

Epidendrum paniculatum var. unguiculatum C. Schweinf. =Epidendrum unguiculatum 

Epidendrum pictum Lindl. =Encyclia picta 

Epidendrum pilosum C. Schweinf. =Encyclia pilosa 

Epidendrum pusillum L. =Erycina pusilla 

Epidendrum pygmaeum Hook. =Prosthechea pygmaea 

Epidendrum pygmaeum Hook. [nom. superfl.] =Prosthechea pygmaea 

Epidendrum raniferum auct., non Lindl. =Epidendrum cristatum 

Epidendrum rectopedunculatum C. Schweinf. =Epidendrum myrmecophorum 

Epidendrum rectopedunculatum C. Schweinf. f. denticulatum C. Schweinf. =Epidendrum orchidiflorum 

Epidendrum remotiflorum C. Schweinf. =Encyclia remotiflora 

Epidendrum ruscifolium Jacq. =Pleurothallis ruscifolia 

Epidendrum satyrioides Sw. =Ionopsis satyrioides 

Epidendrum schlechterianum auct. non Ames 1924: sensu Foldats in Lasser =Epidendrum congestoides 

Epidendrum schomburgkii Lindl. =Epidendrum macrocarpum 

Epidendrum schomburgkii var. confluens Lindl. =Epidendrum calanthum 

Epidendrum secundum auct. non Jacq. 1760: sensu Dunst. & Garay =Epidendrum anceps 

Epidendrum selligerum auct. non Bateman ex Lindl. 1838: sensu Dunst. & Garay =Encyclia granitica 

Epidendrum serricardium Schltr. =Epidendrum vincentinum 

Epidendrum spilotum Garay & Dunst. =Epidendrum myrmecophorum 

Epidendrum spruceanum Lindl. =Epidendrum nocturnum 

Epidendrum stenopetalum Hook. [name misapplied] =Dimerandra stenopetala (Hook.) Schltr. 

Epidendrum teretifolium Sw. =Jacquiniella teretifolia 

Epidendrum tigrinum Linden ex Lindl. =Prosthechea tigrina 

Epidendrum trichorhizum Kraenzl. =Xerorchis trichorhiza 

Epidendrum trigoniflorum Sw. =Stelis ophioglossoides 

Epidendrum uniflorum Barb. Rodr. =Epidendrum althausenii 

Epidendrum utricularioides Sw. =Ionopsis utricularioides 

Epidendrum variegatum Hook. =Prosthechea vespa 

Epidendrum vespa Vell. =Prosthechea vespa 

Epidendrum wageneri Klotzsch, Otto & Dietr. =Encyclia diurna 

Epidendrum weddellii Lindl. =Epidendrum saxatile 

Epidendrum xanthinum Lindl. =Epidendrum secundum 

Epidendrum yatapuense Barb. Rodr. =Epidendrum compressum 

Epistephium cruegeri Rchb. f. ex Griseb. [nomen nudum] =Epistephium subrepens

140 


Epistephium petiolatum Huber =Epistephium parviflorum 

Eriopsis colombiana Schltr. =Eriopsis rutidobulbon 

Eriopsis grandibulbosa Ames & C. Schweinf. =Eriopsis biloba 

Eriopsis schomburgkii (Rchb. f.) Rchb. f. =Eriopsis biloba 

Eriopsis sprucei Rchb. f. =Eriopsis sceptrum 

Erythrodes confusa C. Schweinf. =Aspidogyne confusa 

Erythrodes petersiana (Cogn.) Ames =Ligeophila peterana 

Erythrodes plantaginea (L.) Fawc. & Rendle [dubious in Guianas] 

Erythrodes pumilus (Cogn.) Pabst =Aspidogyne pumila 

Erythrodes robusta C. Schweinf. =Aspidogyne robusta 

Erythrodes santensis (Kraenzl.) C. Schweinf. =Ligeophila juruensis 

Erythrodes steyermarkii (Carnevali & Foldats) Carnevali & Dodson =Aspidogyne steyermarkii 

Erythrodes stigmatoptera (Rchb. f.) Pabst =Ligeophila stigmatoptera 

Eulophidium maculatum (Lindl.) Pfitzer =Oeceoclades maculata 

Evelyna arpophyllostachys Rchb. f. =Elleanthus arpophyllostachys 

Evelyna columnaris Lindl. =Elleanthus columnaris 

Evelyna gracilis Rchb. f. =Elleanthus gracilis 

Evelyna wageneri Rchb. f. =Elleanthus wageneri 

Fernandezia acuta Lindl. =Lockhartia acuta 

Fernandezia graminifolia Ruiz & Pav. =Scaphyglottis graminifolia 

Fractiunguis reflexa Schltr. =Reichenbachanthus reflexus 

Galeandra juncea Lindl. =Galeandra stylomisantha 

Galeandra lacustris Barb. Rodr. =Galeandra minax 

Galeandra montana Barb. Rodr. [dubious in Guianas] =Galeandra lagoensis Reichb.f. & Warm. 

Garayella hexandra (Garay & Dunst.) Brieger [nom. inval.] =Pleurothallis hexandra 

Gongora macrantha Hook. =Coryanthes macrantha 

Gongora quinquenervis Ruiz & Pav. [not in flora area] 

Goodyera guayanensis Lindl. =Brachystele guayanensis 

Habenaria amalfitana F. Lehm. & Kraenzl. =Habenaria parviflora 

Habenaria amazonica Schltr. =Habenaria platydactyla 

Habenaria arecunarum Schltr. =Habenaria parviflora 

Habenaria caldensis auct. non Kraenzl.: sensu Dunst. & Garay =Habenaria ernestii 

Habenaria caldensis auct. non Kraenzl. 1892: sensu Foldats in Lasser =Habenaria lehmanniana 

Habenaria culmiformis Schltr. =Habenaria leprieuri 

Habenaria demerarensis Rchb. f. ex M.R. Schomb. =Habenaria parviflora 

Habenaria enthomantha auct. non (La Llave & Lex) Lindl. 1835: sensu Foldats in Lasser =Habenaria armata 

Habenaria ernestii auct. non Schltr. 1914: sensu Foldats in Lasser =Habenaria caldensis 

Habenaria fastor auct., non Warm. =Habenaria longicauda 

Habenaria georgii Schltr. =Habenaria pratensis var. parviflora 

Habenaria goyazensis Cogn. [as circumscribed by Renz, not in flora area] 

Habenaria guanchezii Carnevali & I. Ramírez =Habenaria pygmaea 

Habenaria leaoana Schltr. =Habenaria sprucei 

Habenaria leprieuri auct. non Rchb. f. 1846: sensu Foldats in Lasser =Habenaria heptadactyla 

Habenaria leprieuri var. heptadactyla (Rchb. f.) R.E. Schult. =Habenaria heptadactyla 

Habenaria macroceratitis Willd. =Habenaria quinqueseta 

Habenaria maculosa Lindl. =Habenaria monorrhiza 

Habenaria moritzii Ridl. =Habenaria armata 

Habenaria muelleriana Cogn. =Habenaria macilenta 

Habenaria pauciflora (Lindl.) Rchb. f. =Habenaria trifida 

Habenaria petalodes auct. non Lindl. 1835: sensu Dunst. & Garay =Habenaria floribunda 

Habenaria rodeiensis auct. non Barb. Rodr.: sensu Dunst. & Garay =Habenaria ernestii 

Habenaria sartor auct. non Lindl. 1843: sensu Foldats in Lasser =Habenaria longicauda 

Habenaria savannensis Renz =Habenaria parviflora 

Habenaria speciosa Poepp. & Endl. =Habenaria monorrhiza 

Habenaria staminodiata Schltr. =Habenaria macilenta 

Habenaria viridiaurea Lindl. ex Kraenzl. =Habenaria heptadactyla 

Hexadesmia bifida Rchb. f. =Scaphyglottis bifida 

Hexadesmia boliviensis Rolfe =Scaphyglottis boliviensis 

Hexadesmia dunstervillei Garay =Scaphyglottis dunstervillei 

Hexadesmia fusiformis Griseb. =Scaphyglottis fusiformis 

Hexadesmia sessilis Rchb. f. =Scaphyglottis sessilis 

Hexisea bidentata var. imbricata (Lindl.) C. Schweinf. =Hexisea imbricata 

Hexisea reflexa (Lindl.) Rchb. f. ex Griseb. =Reichenbachanthus reflexus 

Hoehneella santos-nevesii Ruschi =Chaubardia surinamensis


Houlletia vittata Lindl. =Braemia vittata 

Huebneria yauaperyensis Schltr. =Orleanesia yauaperyensis 

Huebneria yauaperyensis auct. non Schltr. 1925: sensu Hoehne =Epidendrum sculptum 

Humboldtia aspasicensis (Rchb. f.) Kuntze =Pleurothallis aspasicensis 

Humboldtia biflora (H. Focke) Kuntze =Trichosalpinx orbicularis 

Humboldtia ciliata (Knowles & Westc.) Kuntze =Pleurothallis lanceana 

Humboldtia dura (Lindl.) Kuntze =Trichosalpinx dura 

Humboldtia fockei (Lindl.) Kuntze =Pleurothallis fockei 

Humboldtia foliata (Griseb.) Kuntze =Trichosalpinx dura 

Humboldtia grobyi (Lindl.) Kuntze =Pleurothallis grobyi 

Humboldtia memor (Rchb. f.) Kuntze =Trichosalpinx memor 

Humboldtia miqueliana (H. Focke) Kuntze =Pleurothallis miqueliana 

Humboldtia orbicularis (Lindl.) Kuntze =Trichosalpinx orbicularis 

Humboldtia pruinosa (Lindl.) Kuntze =Pleurothallis pruinosa 

Humboldtia revoluta Ruiz & Pav. =Pleurothallis revoluta 

Humboldtia zephyrina (Rchb. f.) Kuntze =Pleurothallis zephyrina 

Huntleya albido-fulva Lem. =Huntleya meleagris 

Huntleya violacea Lindl. =Bollea violacea 

Hybochilus huebneri Mansf. =Polyotidium huebneri 

Ionopsis paniculata Lindl. =Ionopsis utricularioides 

Ionopsis pygmaea Cogn. =Stictophyllorchis pygmaea 

Ionopsis teres Lindl. =Ionopsis satyrioides 

Isochilus elegans H. Focke =Dimerandra elegans 

Kefersteinia pusilla auct., non (C. Schweinf.) C. Schweinf. =Kefersteinia lafontainei 

Kegelia houtteana Rchb. f. =Kegeliella houtteana 

Koellensteinia alba (Ridl. ex Thurn) Schltr. =Otostylis lepida 

Koellensteinia eburnea (Barb. Rodr.) Schltr. =Koellensteinia tricolor 

Laelia crispa (Lindl.) Lemée =Schomburgkia crispa 

Laelia marginata (Lindl.) L.O. Williams =Schomburgkia marginata 

Laelia undulata (Lindl.) L.O. Williams =Schomburgkia undulata 

Lanium microphyllum (Lindl.) Benth. =Epidendrum microphyllum 

Lanium subulatum Rolfe =Epidendrum stiliferum 

Leaoa reedii (Rchb. f.) Garay =Scaphyglottis sessilis 

Lepanthes funerea Barb. Rodr. =Pleurothallis funerea 

Lepanthes orbiculata Lindl. ex Rchb. f. =Trichosalpinx orbicularis 

Lepanthes yauaperyensis Barb. Rodr. =Pleurothallis mentosa 

Leptorchis kappleri (Rchb. f.) Kuntze =Liparis nervosa 

Limodorum altum L. =Eulophia alta 

Limodorum diurnum Jacq. =Encyclia diurna 

Limodorum grandiflorum Aubl. =Cleistes grandiflora 

Limodorum lanceolatum Aubl. =Sacoila lanceolata 

Limodorum pendulum Aubl. =Dichaea pendula 

Lindleyella aurantiaca (Lindl.) Schltr. =Rudolfiella aurantiaca 

Liparis elata Lindl. =Liparis nervosa subsp. nervosa 

Liparis elliptica (Rchb. f.) Rchb. f. [non Wight] =Liparis jamaicensis 

Liparis kappleri (Rchb. f.) Rchb. f. =Liparis nervosa 

Liparis nervosa f. kappleri (Rchb. f.) Christenson & Carnevali =Liparis nervosa 

Liparis vexillifera auct. non (La Llave & Lex.) Cogn.: sensu Dunst. & Garay =Liparis jamaicensis 

Lockhartia elegans Hook. =Lockhartia imbricata 

Lockhartia lasseri Schnee =Lockhartia acuta 

Lockhartia micrantha Rchb. f. [not in flora area] 

Lockhartia pallida Rchb. f. =Lockhartia acuta 

Lockhartia weigeltii Rchb. & Rchb. f. ex Rchb. f. =Lockhartia imbricata 

Lophiaris carthagenensis (Jacq.) Braem =Trichocentrum carthagenense 

Lophiaris fragrans Raf. =Trichocentrum lanceanum 

Lophiaris lanceana (Lindl.) Braem =Trichocentrum lanceanum 

Lophiaris nana (Lindl.) Braem =Trichocentrum nanum 

Lycaste lindeniana (Rchb. f.) G. Nicholson =Paphinia lindeniana 

Lycomormium minus Kraenzl. =Peristeria cerina 

Macradenia modesta (Rolfe) Rolfe =Macradenia brassavolae 

Macradenia mutica Lindl. =Leucohyle mutica 

Macradenia surinamensis Rchb. f. & Wullschl. =Macradenia lutescens 

Macradenia triandra Lindl. =Macradenia lutescens 

Macroclinium norae (Garay) Dodson =Macroclinium mirabile

142 


Malaxis excavata auct. non (Lindl.) Kuntze 1891: sensu Foldats in Lasser =Malaxis maguirei 

Malaxis umbelliflora Sw. [not in flora area] 

Manniella americana C. Schweinf. & Garay =Helonoma americana 

Marsupiaria equitans Schltr. =Maxillaria equitans 

Marsupiaria mattogrossensis (Brade) Hoehne =Maxillaria equitans 

Masdevallia brevis Rchb. f. =Scaphosepalum breve 

Masdevallia cayennensis Rchb. f. =Masdevallia cuprea 

Masdevallia guttulata auct. non Rchb. f. 1877: sensu Dunst. & Garay =Masdevallia norae 

Masdevallia huebneri Schltr. =Trisetella triglochin 

Masdevallia surinamensis H. Focke =Masdevallia minuta 

Masdevallia triaristella auct. non Rchb. f. 1876: sensu Foldats in Lasser =Trisetella triglochin 

Masdevallia triglochin Rchb. f. =Trisetella triglochin 

Maxillaria amazonica Schltr. =Maxillaria kegelii 

Maxillaria amazonica auct. non Schltr. 1925: sensu Dunst. & Garay =Maxillaria porrecta 

Maxillaria anatamorum auct. non Rchb. f.: sensu Foldats in Lasser =Maxillaria albiflora 

Maxillaria arachnitis auct non Rchb. f.: sensu Foldats in Lasser =Maxillaria marmoliana 

Maxillaria attenuata Ames & C. Schweinf. =Maxillaria guadalupensis 

Maxillaria auyantepuiensis auct. non Foldats sensu: Foldats in Lasser =Maxillaria longipetiolata 

Maxillaria auyantepuiensis subsp. epiphytica Carnevali & I. Ramírez =Maxillaria longipetiolata 

Maxillaria brevipedunculata Ames & C. Schweinf. =Maxillaria nasuta 

Maxillaria brunnea auct. non Linden & Rchb. f. 1854: sensu Dunst. & Garay =Maxillaria cryptobulbon 

Maxillaria brunnea Linden & Rchb. f. =Maxillaria porrecta 

Maxillaria buchtienii Schltr. =Maxillaria splendens 

Maxillaria colleyi Bateman ex Lindl. =Xylobium colleyi 

Maxillaria condensata C. Schweinf. =Maxillaria mapiriensis 

Maxillaria coriacea Barb. Rodr. =Maxillaria desvauxiana 

Maxillaria cozieriana H.G. Jones [possibly distinct in GU] =Maxillaria camaridii 

Maxillaria cristata Lindl. =Paphinia cristata 

Maxillaria foveata Lindl. =Xylobium foveatum 

Maxillaria graminea Lindl. =Koellensteinia graminea 

Maxillaria grandiflora auct. non (Kunth) Lindl. 1832: sensu Foldats in Lasser =Maxillaria eburnea 

Maxillaria iguapensis Hoehne & Schltr. =Maxillaria imbricata 

Maxillaria lactea Schltr. =Maxillaria alba 

Maxillaria lactea Schltr., non Schltr. 1923 =Maxillaria kegelii 

Maxillaria lactiflora Pabst =Maxillaria kegelii 

Maxillaria longifolia (Barb. Rodr.) Cogn. =Heterotaxis superflua 

Maxillaria macrophylla Poepp. & Endl. =Lycaste macrophylla 

Maxillaria melina auct. non Lindl.: sensu Foldats in Lasser =Maxillaria albiflora 

Maxillaria nana Hook. =Maxillaria uncata 

Maxillaria nasuta auct. non Rchb. f. 1866: sensu Dunst. & Garay =Heterotaxis proboscidea 

Maxillaria pallidiflora Hook. =Xylobium pallidiflorum 

Maxillaria pendens Pabst =Maxillaria rigida 

Maxillaria perparva Garay & Dunst. =Hylaeorchis petiolaris 

Maxillaria petiolaris Schltr. =Hylaeorchis petiolaris 

Maxillaria petiolaris A. Rich. [nomen nudum] =Maxillaria desvauxiana 

Maxillaria proboscidea Rchb. f. =Heterotaxis proboscidea 

Maxillaria purpurea auct. non (Spreng.) Ames & Correll 1943: sensu Foldats in Lasser =Maxillaria conferta 

Maxillaria ringens auct. non Rchb. f. 1863: sensu Dunst. & Garay =Maxillaria porrecta 

Maxillaria riopalenquensis Dodson =Maxillaria longipetiolata 

Maxillaria rudolfii Hoehne [nomen nudum] =Hylaeorchis petiolaris 

Maxillaria rufescens auct. non Lindl. 1835 [1836]: sensu Foldats in Lasser =Maxillaria acutifolia 

Maxillaria rugosa Schltr. =Maxillaria schlechteri 

Maxillaria simulans Ames & C. Schweinf. =Maxillaria conferta 

Maxillaria spilotantha auct. non Rchb. f. 1854: sensu Foldats in Lasser =Maxillaria guareimensis 

Maxillaria steelei Hook. =Scuticaria steelei 

Maxillaria superflua Rchb. f. =Heterotaxis superflua 

Maxillaria surinamensis H. Focke ex Rchb. f. =Maxillaria conferta 

Maxillaria tafallae (Rchb. f.) C. Schweinf. =Maxillaria ramosa 

Maxillaria taracuana Schltr. =Maxillaria kegelii 

Maxillaria tarumaensis Hoehne =Heterotaxis superflua 

Maxillaria trinitatis Ames ex Broadway [nom. superfl.] =Maxillaria porrecta 

Maxillaria trinitatis auct. non Ames 1923: Dunst. & Garay =Maxillaria cryptobulbon 

Maxillaria trinitatis Ames =Maxillaria porrecta 

Maxillaria uncata auct. non Lindl. 1837: sensu Foldats in Lasser =Maxillaria squamata


Maxillaria vandiformis (Schltr.) C. Schweinf. [possibly distinct species] =Maxillaria equitans 

Maxillaria variabilis auct., non Bateman ex Lindl. =Maxillaria caespitifica 

Maxillaria verrucifera C. Schweinf. =Maxillaria desvauxiana 

Maxillaria vestita (Sw.) Lemée =Maxillaria parviflora 

Maxillaria villosa (Barb. Rodr.) Cogn. =Heterotaxis villosa 

Maxillaria violaceopunctata Rchb. f. =Heterotaxis violaceopunctata 

Medadenium labiosum (Rich.) Cogn. =Zygosepalum labiosum 

Medadenium lindeniae (Rolfe) Cogn. =Zygosepalum lindeniae 

Menadenium kegelii (Rchb. f.) Cogn. =Zygosepalum kegelii 

Menadenium labiosum (Rich.) Cogn. =Zygosepalum labiosum 

Mendoncella burkei (Rchb. f.) Garay =Galeottia burkei 

Mendoncella jorisiana (Rolfe) A.D. Hawkes =Galeottia jorisiana 

Mesadenella esmeraldae (Lindl. & Rchb. f.) Pabst & Garay =Mesadenella cuspidata 

Monachanthus discolor Lindl. =Catasetum discolor 

Monachanthus roseo-albus Hook. =Catasetum ×roseo-album 

Myanthus barbatus Lindl. =Catasetum barbatum 

Myanthus deltoideus Lindl. =Catasetum deltoideum 

Myoxanthus aspasicensis (Rchb. f.) Luer =Pleurothallis aspasicensis 

Myoxanthus aspasicensis subsp. arenicola Carnevali & I. Ramírez =Pleurothallis arenicola 

Myoxanthus lappiformis (A.H. Heller & L.O. Williams) Luer =Pleurothallis lappiformis 

Nanodes discolor Lindl. =Epidendrum schlechterianum 

Neobartlettia guianensis Schltr. =Palmorchis guianensis 

Neolehmannia apagana (Mansf.) Garay & Dunst. =Epidendrum apaganum 

Neolehmannia barbeyana auct. non (Kraenzl.) Garay & Dunst. 1976: sensu Dunst. & Garay =Epidendrum althausenii 

Neolehmannia pabstii Braga =Epidendrum dichaeoides 

Neottia acaulis Sm. =Sarcoglottis acaulis 

Neottia calcarata Sw. =Eltroplectris calcarata 

Nidema boothii (Lindl.) Schltr. [not in flora area] 

Notylia amesii L.B. Sm. & S.K. Harris =Notylia microchila 

Notylia amesii L.B. Sm. & S.K. Harris =Notylia microchila 

Notylia bicolor Lindl. [not in flora area] =Macroclinium bicolor (Lindl.) Dodson 

Notylia mirabilis C. Schweinf. =Macroclinium mirabile 

Notylia nana Cogn. =Notylia angustifolia 

Notylia norae Garay =Macroclinium mirabile 

Notylia trullifera Brade =Notylia longispicata 

Notylia venezuelana Schltr. =Notylia fragrans 

Notylia wullschlaegeliana H. Focke =Macroclinium wullschlaegelianum 

Octomeria brevifolia Cogn. =Octomeria scirpoidea 

Octomeria complanata C. Schweinf. =Octomeria yauaperyensis 

Octomeria deltoglossa Garay =Octomeria exigua 

Octomeria exigua var. elata C. Schweinf. =Octomeria exigua 

Octomeria grandiflora Lindl. [not in flora area] 

Octomeria kestrochila Garay & Dunst. =Octomeria exigua 

Octomeria pusilla Lindl. [not in flora area] 

Odontoglossum arminii Rchb. f. =Otoglossum arminii 

Odontoglossum brevifolium auct. non. Lindl. 1844: sensu Foldats in Lasser =Otoglossum arminii 

Oncidium altissimum auct., non (Jacq.) Sw. =Oncidium baueri 

Oncidium ampliatum Lindl. =Chelyorchis ampliata 

Oncidium bernouillanum Kraenzl. =Chelyorchis ampliata 

Oncidium carthagenense (Jacq.) Sw. =Trichocentrum carthagenense 

Oncidium cebolleta (Jacq.) Sw. =Trichocentrum cebolleta 

Oncidium convolvulaceum Lindl. =Otoglossum globuliferum 

Oncidium emarginatum G. Mey. =Dimerandra emarginata 

Oncidium globuliferum auct. non Kunth 1815 [1816]: sensu Dunst. & Garay =Otoglossum scansor 

Oncidium globuliferum Kunth =Otoglossum globuliferum 

Oncidium glossomystax Rchb. f. =Erycina glossomystax 

Oncidium guianense (Aubl.) Garay [not in flora area] 

Oncidium kappleri Rchb. f. ex Lindl. =Oncidium baueri 

Oncidium lanceanum Lindl. =Trichocentrum lanceanum 

Oncidium lunatum Lindl. =Solenidium lunatum 

Oncidium maculatum Urb. =Oncidium luridum 

Oncidium nanum Lindl. =Trichocentrum nanum 

Oncidium papilio Lindl. =Psychopsis papilio 

Oncidium pusillum (L.) Rchb. f. =Erycina pusilla

144 


Oncidium sancti-pauli auct. non Kaenzl. 1922: sensu Foldats in Lasser =Otoglossum globuliferum 

Oncidium scansor Rchb. f. =Otoglossum scansor 

Oncidium uaipanense Schnee =Oncidium nigratum 

Oncidium ultrajectinum Pulle =Trichocentrum cebolleta 

Oncidium vagans C. Schweinf. =Oncidium warmingii 

Oncidium versteegianum Pulle =Psychopsis versteegianum 

Onychium flavescens Blume =Polystachya concreta 

Ophrys guianensis Aubl. [not in flora area] =Oncidium guianense (Aubl.) Garay 

Ophrys nervosa Thunb. =Liparis nervosa subsp. nervosa 

Ophrys nervosa Thunb. =Liparis nervosa 

Ophrys peruviana Aubl. =Spiranthes torta 

Orchis monorrhiza Sw. =Habenaria monorrhiza 

Orchis quinqueseta Michx. =Habenaria quinqueseta 

Orchis stylomisantha Vell. =Galeandra stylomisantha 

Ornithidium aureum Poepp. & Endl. =Maxillaria aurea 

Ornithidium chloroleucum Barb. Rodr. =Maxillaria conferta 

Ornithidium confertum Griseb. =Maxillaria conferta 

Ornithidium mapiriense Kraenzl. =Maxillaria mapiriensis 

Ornithidium serrulatum Lindl. [non Maxillaria serrulata Ames & Correll 1943] =Maxillaria alticola 

Ornithidium squamatum (Barb. Rodr.) Barb. Rodr. =Maxillaria squamata 

Ornithidium vestitum Rchb. f. =Maxillaria conferta 

Ornithocephalus avicula Rchb. f. =Ornithocephalus kruegeri 

Ornithocephalus bicornis auct. non Lindl. 1846: sensu Garay & Dunst. =Ornithocephalus brachyceras 

Ornithocephalus planifolius Rchb. f. =Dipteranthus planifolius 

Otostylis alba (Ridl. ex Thurn) Summerh. =Otostylis lepida 

Otostylis paludosa (Cogn.) Schltr. =Otostylis brachystalix 

Otostylis venusta (Ridl. ex Oliv.) Schltr. =Otostylis lepida 

Paphiopedilum klotzschianum Pfitzer =Phragmipedium klotzschianum 

Paphiopedilum lindleyanum Pfitzer =Phragmipedium lindleyanum 

Pelexia aphylla Ridl. ex Thurn =Sarcoglottis aphylla 

Pelexia aphylla Ridl. ex Oliv. [nom. superfl.] =Sarcoglottis aphylla 

Pelexia cranichoides Griseb. =Cyclopogon cranichoides 

Pelexia maculata Rolfe =Pelexia laxa 

Pelexia setacea Lindl. =Eltroplectris calcarata 

Peristeria guttata Knowles & Westc. =Peristeria cerina 

Phragmipedium kaieteurum (N.E. Br.) Garay =Phragmipedium lindleyanum 

Phymatidium panamense Dressler =Eloyella panamensis 

Physinga prostrata Lindl. =Epidendrum prostratum 

Physosiphon obliquipetalus Ames & C. Schweinf. =Lepanthopsis obliquipetala 

Physurus arietinus Rchb. f. & Warm. =Erythrodes arietina 

Physurus juruensis Hoehne =Ligeophila juruensis 

Physurus paleaceus Schltr. =Erythrodes paleacea 

Physurus pumilus Cogn. =Aspidogyne pumila 

Physurus santensis Kraenzl. =Ligeophila juruensis 

Physurus stigmatopterus Rchb. f. =Ligeophila stigmatoptera 

Pilumna wageneri Rchb. f. =Trichopilia wageneri 

Pinelia alticola Garay & Dunst. =Pinelianthe alticola 

Platystele lancilabris auct. non (Rchb. f.) Schltr. 1923: sensu Garay & Dunst. =Platystele oxyglossa 

Plectrophora cultrifolia auct. non (Barb. Rodr.) Cogn. 1904: sensu Garay & Dunst. =Plectrophora iridifolia 

Pleurothallis acuminata H. Focke =Pleurothallis semperflorens 

Pleurothallis acutissima Lindl. =Pleurothallis spiculifera 

Pleurothallis acutissima auct. non Lindl. 1856: sensu Foldats in Lasser =Pleurothallis trinitensis 

Pleurothallis amygdalina Rolfe ex Schltr. [nomen nudum] =Trichosalpinx dura 

Pleurothallis anomala Hoehne =Trichosalpinx dura 

Pleurothallis apiculata Lindl. =Octomeria apiculata 

Pleurothallis barbata H. Focke =Pleurothallis barbulata 

Pleurothallis barberiana Rchb. f. =Pleurothallis aristata 

Pleurothallis biflora H. Focke =Trichosalpinx orbicularis 

Pleurothallis blaisdellii S. Watson [not in flora area] =Trichosalpinx blaisdellii (S. Watson) Luer 

Pleurothallis brevis Schltr. =Trichosalpinx memor 

Pleurothallis breviscapa C. Schweinf. =Pleurothallis funerea 

Pleurothallis broadwayi Ames =Trichosalpinx dura 

Pleurothallis broadwayi subsp. anomala (Hoehne) Garay =Trichosalpinx dura 

Pleurothallis cedralensis Ames =Trichosalpinx cedralensis


Pleurothallis ciliaris auct. non (Lindl.) L.O. Williams 1842: sensu Foldats in Lasser =Trichosalpinx memor 

Pleurothallis ciliaris (Lindl.) L.O. Williams =Trichosalpinx ciliaris 

Pleurothallis ciliata Knowles & Westc. =Pleurothallis lanceana 

Pleurothallis ciliata var. elongata C. Schweinf. =Pleurothallis lanceana 

Pleurothallis ciliolata auct. non Schltr. 1926: sensu Foldats in Lasser =Pleurothallis funerea 

Pleurothallis coffeicola auct. non Schltr. 1929: sensu Foldats in Lasser =Pleurothallis discophylla p.p., P. sandaliorum p.p. 

Pleurothallis connata Luer =Lepanthopsis obliquipetala 

Pleurothallis consimilis Ames =Pleurothallis yauaperyensis 

Pleurothallis consimilis auct. non Ames 1922: sensu Garay & Dunst. =Pleurothallis garciae 

Pleurothallis corazonica F. Lehm. & Kraenzl. =Trichosalpinx dura 

Pleurothallis delascioi Carnevali & I. Ramírez =Pleurothallis kerrii 

Pleurothallis diffusiflora C. Schweinf. =Pleurothallis seriata 

Pleurothallis dura Lindl. =Trichosalpinx dura 

Pleurothallis egleri Pabst =Trichosalpinx egleri 

Pleurothallis elegans auct. non (Kunth) Lindl. 1842: sensu Foldats in Lasser =Pleurothallis moritzii 

Pleurothallis exasperata auct. non Lindl. 1859: sensu Foldats in Lasser =Myoxanthus speciosus 

Pleurothallis fimbriata Lindl. =Pleurothallis miqueliana 

Pleurothallis floripecten Rchb. f. =Lepanthopsis floripecten 

Pleurothallis foliata Griseb. =Trichosalpinx dura 

Pleurothallis gnomonifera Ames =Trichosalpinx memor 

Pleurothallis guadalupensis Cogn. =Trichosalpinx dura 

Pleurothallis huebneri Schltr. =Pleurothallis lanceana 

Pleurothallis intricata Lindl. =Trichosalpinx intricata 

Pleurothallis johnstonii Ames =Platystele johnstonii 

Pleurothallis kegelii Rchb. f. =Pleurothallis lanceana 

Pleurothallis lancifera Schltr. =Trichosalpinx orbicularis 

Pleurothallis lancilabris var. oxyglossa (Schltr.) C. Schweinf. =Platystele oxyglossa 

Pleurothallis lansbergii auct. non Regel 1856: sensu Dunst. & Garay =Pleurothallis coriacardia 

Pleurothallis lepanthopsis Schltr. =Trichosalpinx dura 

Pleurothallis leptopetala Cogn. =Pleurothallis discoidea 

Pleurothallis linguifera auct. non Lindl. 1859: sensu Foldats in Lasser =Pleurothallis coriacardia 

Pleurothallis longirostris H. Focke =Pleurothallis ephemera 

Pleurothallis marginalis Rchb. f. =Pleurothallis grobyi 

Pleurothallis memor Rchb. f. =Trichosalpinx memor 

Pleurothallis minutipetala Ames & C. Schweinf. =Trichosalpinx memor 

Pleurothallis monocardia auct. non Rchb. f. 1855: sensu Garay & Dunst. =Pleurothallis archidiaconi 

Pleurothallis myrmecophila Hoehne =Pleurothallis fockei 

Pleurothallis orbicularis (Lindl.) Lindl. =Trichosalpinx orbicularis 

Pleurothallis ovalifolia (H. Focke) Rchb. f. =Platystele ovalifolia 

Pleurothallis oxyglossa Schltr. =Platystele oxyglossa 

Pleurothallis papillosa auct. non. Lindl. 1836: sensu Dunst. & Garay =Pleurothallis granitica 

Pleurothallis parvilabia C. Schweinf. =Myoxanthus parvilabius 

Pleurothallis peduncularis Lindl. =Myoxanthus parahybunensis 

Pleurothallis penduliflora auct. non Kraenzl. 1905: sensu Garay & Dunst. =Pleurothallis suspensa 

Pleurothallis pertenuis C. Schweinf. =Pleurothallis glandulosa 

Pleurothallis plumosa Lindl. =Pleurothallis lanceana 

Pleurothallis pluriracemosa Garay =Pleurothallis floribunda 

Pleurothallis pulchella auct. non (Kunth) Lindl. 1825: sensu Luer =Pleurothallis tepuiensis 

Pleurothallis pumila Luer =Trichosalpinx pumila 

Pleurothallis revoluta auct. non (Ruiz & Pav.) Garay 1962: sensu Foldats in Lasser =Pleurothallis suspensa 

Pleurothallis reymondii (H. Karst.) Rchb. f. =Myoxanthus reymondii 

Pleurothallis ringens C. Schweinf. =Pleurothallis hitchcockii 

Pleurothallis roraimensis Rolfe =Trichosalpinx roraimensis 

Pleurothallis scandens var. simplicicaulis C. Schweinf. =Myoxanthus simplicicaulis 

Pleurothallis sertularioides var. trinitensis Griseb. =Pleurothallis trinitensis 

Pleurothallis setigera Lindl. =Pleurothallis zephyrina 

Pleurothallis simplicicaulis (C. Schweinf.) Luer =Myoxanthus simplicicaulis 

Pleurothallis speciosus Luer =Myoxanthus speciosus 

Pleurothallis spiculifera auct. non Lindl. 1859: sensu Dunst. & Garay =Pleurothallis trinitensis 

Pleurothallis stenopetala Lodd. ex Lindl. =Pleurothallis sclerophylla 

Pleurothallis striata H. Focke =Pleurothallis uniflora 

Pleurothallis tepuiensis auct. non Carnevali & I. Ramírez 1993: sensu G.A. Romero & Carne =Pleurothallis moritzii 

Pleurothallis trachyclamys Schltr. =Myoxanthus trachychlamys 

Pleurothallis trachytheca F. Lehm. & Kraenzl. =Trichosalpinx orbicularis

146 


Pleurothallis tricarinata H. Focke =Pleurothallis fockei 

Pleurothallis tubulosa Lindl. =Restrepiopsis tubulosa 

Pleurothallis uncinata auct. non Fawc. 1895: sensu Foldats in Lasser =Pleurothallis arenicola 

Pleurothallis uncinata auct. non Fawc. 1895, sensu Foldats p.p., Dunst. & Garay p.p. =Pleurothallis aspasicensis 

Pleurothallis uniflora auct non Lindl. 1836: sensu Dunst. & Garay =Pleurothallis corniculata 

Pleurothallis vaginata Schltr. =Pleurothallis imraei 

Pleurothallis velaticaulis auct. non Rchb. f. 1849: sensu Foldats in Lasser =Pleurothallis floribunda 

Pleurothallis velatipes Rchb. f. =Pleurothallis velaticaulis 

Pleurothallis villosa Knowles & Westc. =Trichosalpinx egleri 

Pleurothallis viridula Lindl. =Restrepiopsis tubulosa 

Pleurothallis vitatta auct. non Lindl. 1838: sensu Foldats in Lasser =Pleurothallis sarcosepala 

Pleurothallis vittariifolia Schltr. =Pleurothallis glandulosa 

Pleurothallis wendlandiana auct. non Rchb. f. 1888: sensu Dunst. & Garay =Pleurothallis ruscifolia 

Pleurothallis williamsii Ames =Trichosalpinx dura 

Pogonia grandiflora Rchb. f. =Cleistes grandiflora 

Pogonia lepida Rchb. f. =Cleistes lepida 

Pogonia macrophylla Lindl. =Psilochilus macrophyllus 

Pogonia moritzii Rchb. f. =Cleistes lepida 

Pogonia paludosa (Rchb. f.) Rchb. f. =Cleistes paludosa 

Pogonia parviflora (Lindl.) Rchb. f. =Cleistes parviflora 

Pogonia physuraefolia Rchb. f. =Psilochilus macrophyllus 

Pogonia physurifolia Rchb. f. =Psilochilus physurifolius 

Pogonia rosea (Lindl.) Rchb. f. =Cleistes rosea 

Pogonia stricta C. Schweinf. =Cleistes stricta 

Pogonia surinamensis Lindl. =Triphora surinamensis 

Pogonia tenuis Rchb. f. =Cleistes tenuis 

Pogonia triflora C. Schweinf. =Cleistes triflora 

Pogonia unifoliata C. Schweinf. =Cleistes unifoliata 

Polycycnis vittata (Lindl.) Rchb. f. =Braemia vittata 

Polystachya estrellensis Rchb. f. [not in flora area] 

Polystachya flavescens (Blume) J.J. Sm. =Polystachya concreta 

Polystachya geraensis Barb. Rodr. [not in flora area] 

Polystachya luteola (Sw.) Hook. =Polystachya concreta 

Polystachya minuta Britton [not in flora area] 

Polystachya nana Poepp. & Endl. [not in flora area] 

Polystachya stenophylla Schltr. =Polystachya amazonica 

Ponera amethystina Rchb. f. =Scaphyglottis stellata 

Ponera felskyi Rchb. f. =Scaphyglottis felskyi 

Ponera leucantha (Rchb. f.) Rchb. f. =Scaphyglottis leucantha 

Prescottia orchioides auct. non Lindl. 1845: sensu Foldats in Lasser =Prescottia auyantepuiensis 

Promenaea graminea (Lindl.) Lindl. =Koellensteinia graminea 

Prosthechea pamplonensis (Rchb. f.) W.E. Higgins =Prosthechea tigrina 

Pseuderiopsis schomburgkii Rchb. f. =Eriopsis biloba 

Pseudomaxillaria chloroleuca (Barb. Rodr.) Hoehne =Maxillaria conferta 

Psychopsis picta Raf. =Psychopsis papilio 

Psygmorchis glossomystax (Rchb. f.) Dodson & Dressler =Erycina glossomystax 

Psygmorchis pusilla (L.) Dodson & Dressler =Erycina pusilla 

Quekettia pygmaea (Cogn.) Garay & R.E. Schult. =Stictophyllorchis pygmaea 

Reichenbachanthus modestus Barb. Rodr. =Reichenbachanthus reflexus 

Restrepia kegelii Rchb. f. [non Pleurothallis kegelii Rchb. f. 1877] =Pleurothallis hexandra 

Restrepiella viridula (Lindl.) Garay & Dunst. =Restrepiopsis tubulosa 

Restrepiopsis viridula (Lindl.) Luer =Restrepiopsis tubulosa 

Rhynchopera punctata H. Karst. =Pleurothallis loranthophylla 

Rodriguezia secunda Kunth =Rodriguezia lanceolata 

Sarcoglottis acaulis auct. non (Sm.) Schltr. 1919: sensu Foldats in Lasser =Sarcoglottis metallica 

Sarcoglottis grandiflora auct. non (Lindl.) Klotzsch 1842: sensu Dunst. & Garay =Sarcoglottis acaulis 

Sarcoglottis maasorum Pabst =Sarcoglottis metallica 

Sarcoglottis picta auct. non Klotzsch 1842: sensu Garay & Dunst. =Sarcoglottis metallica 

Satyrium elatum Sw. =Cyclopogon elatus 

Satyrium orchioides Sw. =Sacoila lanceolata 

Scaphosepalum ochthodes Pfitzer [not in flora area] 

Scaphosepalum verrucosum auct. non (Rchb. f.) Pfitzer 1888: sensu Dunst. & Garay =Scaphosepalum breve 

Scaphyglottis amethystina (Rchb. f.) Schltr. =Scaphyglottis stellata 

Scaphyglottis cuneata Schltr. =Scaphyglottis prolifera


Scaphyglottis huebneri Schltr. =Scaphyglottis boliviensis 

Scaphyglottis propinqua auct. non C. Schweinf. 1955; Foldats in Lasser =Scaphyglottis sickii 

Scaphyglottis reedii (Rchb. f.) Ames =Scaphyglottis sessilis 

Scaphyglottis reflexa Lindl. =Reichenbachanthus reflexus 

Scaphyglottis tafallae Rchb. f. =Maxillaria ramosa 

Scaphyglottis violacea (Lindl.) Lindl. =Scaphyglottis graminifolia 

Scaphyglottis wercklei var. major C. Schweinf. =Scaphyglottis leucantha 

Schlechterella aurantiaca (Lindl.) Hoehne =Rudolfiella aurantiaca 

Schomburgkia crispa auct. non Lindl. 1838: sensu Foldats in Lasser =Schomburgkia marginata 

Schomburgkia rosea auct. non Linden ex Lindl. 1845: sensu Foldats in Lasser =Schomburgkia heidii 

Schomburgkia weberbaueriana auct. non Kraenzl. 1906: sensu Garay & Dunst. =Schomburgkia marginata 

Selenipedium kaieteurum N.E. Br. =Phragmipedium lindleyanum 

Selenipedium lindleyanum Rchb. f. =Phragmipedium lindleyanum 

Selenipedium schomburgkianum Rchb. f. =Phragmipedium klotzschianum 

Serapias caravata Aubl. =Elleanthus caravata 

Serrastylis modesta Rolfe =Macradenia brassavolae 

Sievekingia dunstervilleorum Foldats =Sievekingia jenmanii 

Sobralia biflora auct. non Ruiz & Pav. 1798: sensu Schweinf. =Sobralia valida 

Sobralia cattleya Rchb. f. =Sobralia dichotoma 

Sobralia elisabethae R.H. Schomb. =Sobralia liliastrum 

Sobralia liliastrum var. rosea Lindl. =Sobralia infundibuligera 

Sobralia sclerophylla auct. non Lindley =Sobralia stenophylla 

Sobralia sessilis auct. non Lindl. 1841: sensu Foldats in Lasser =Sobralia valida 

Sobralia violacea auct. non Linden & Rchb. f. 1846: sensu Garay & Dunst. =Sobralia sessilis 

Sobralia yauaperyensis Barb. Rodr. =Sobralia sessilis 

Sobralia yauaperyensis auct. non Barb. Rodr. 1881: sensu Foldats in Lasser =Sobralia violacea 

Spathiger rigidus (Jacq.) Small =Epidendrum rigidum 

Specklinia aristata (Hook.) Pridgeon & M.W. Chase =Pleurothallis aristata 

Specklinia grobyi (Bateman ex Lindl.) Pridgeon & M.W. Chase =Pleurothallis grobyi 

Specklinia miqueliana H. Focke =Pleurothallis miqueliana 

Specklinia orbicularis Lindl. =Trichosalpinx orbicularis 

Specklinia parvifolia (Lindl.) Pridgeon & M.W. Chase =Pleurothallis parvifolia 

Specklinia picta (Lindl.) Pridgeon & M.W. Chase =Pleurothallis picta 

Specklinia samacensis (Ames) Pridgeon & M.W. Chase =Pleurothallis samacensis 

Specklinia seriata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis seriata 

Specklinia spiculifera (Lindl.) Pridgeon & M.W. Chase =Pleurothallis spiculifera 

Specklinia vittariifolia (Schltr.) Pridgeon & M.W. Chase =Pleurothallis glandulosa 

Spiranthes acaulis (Sm.) Cogn. =Sarcoglottis acaulis 

Spiranthes aphylla (Ridl. ex Thurn) Cogn. =Sarcoglottis aphylla 

Spiranthes bifida Ridl. ex Oliv. [nom. superfl.] =Helonoma bifida 

Spiranthes bifida Ridl. ex Thurn =Helonoma bifida 

Spiranthes callifera C. Schweinf. =Pelexia callifera 

Spiranthes cranichoides (Griseb.) Cogn. =Cyclopogon cranichoides 

Spiranthes elata (Sw.) Rich. =Cyclopogon elatus 

Spiranthes goninensis (Pulle) C. Schweinf. =Pelexia goninensis 

Spiranthes grisebachii Cogn. =Lyroglossa grisebachii 

Spiranthes guayanensis (Lindl.) Cogn. =Brachystele guayanensis 

Spiranthes lanceolata (Aubl.) León =Sacoila lanceolata 

Spiranthes laxa (Poepp. & Endl.) C. Schweinf. =Pelexia laxa 

Spiranthes maculata (Rolfe) C. Schweinf. =Pelexia laxa 

Spiranthes metallica Rolfe =Sarcoglottis metallica 

Spiranthes orchioides (Sw.) A. Rich. =Sacoila lanceolata 

Spiranthes quinqueloba (Poir.) Urb. =Spiranthes torta 

Spiranthes scopulariae Rchb. f. =Discyphus scopulariae 

Spiranthes simplex Griseb. =Sarcoglottis simplex 

Stanhopea aurea Lodd. ex Lindl. =Stanhopea wardii 

Stanhopea eburnea Lindl. =Stanhopea grandiflora 

Stanhopea randii Rolfe =Stanhopea candida 

Stanhopea venusta Lindl. =Stanhopea wardii 

Stelis aprica auct. non Lindl. 1836: sensu Dunst. & Garay =Stelis schomburgkii 

Stelis braccata auct. non Rchb. f. & Warsz. 1854: sensu Foldats in Lasser =Stelis minimiflora 

Stelis cupuligera auct. non Rchb. f. 1854: sensu Dunst. & Garay =Stelis aviceps 

Stelis drosophila Barb. Rodr. =Stelis intermedia 

Stelis flavida H. Focke =Pleurothallis pruinosa

148 


Stelis floribunda (Poepp. & Endl.) Pridgeon & M.W. Chase =Pleurothallis floribunda 

Stelis foliosa Lindl. =Polystachya foliosa 

Stelis galeata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis galeata 

Stelis grossilabris auct. non Rchb. f. 1881: sensu Foldats in Lasser =Stelis zonata 

Stelis imraei (Lindl.) Pridgeon & M.W. Chase =Pleurothallis imraei 

Stelis maderoi auct. non Schltr. 1920: sensu G.A. Romero & Carnevali =Stelis obovata 

Stelis moritzii (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis moritzii 

Stelis obliquipetala (Ames & C. Schweinf.) L.O. Williams =Lepanthopsis obliquipetala 

Stelis ovalifolia H. Focke =Platystele ovalifolia 

Stelis papaquerensis auct. non Rchb. f. 1849: sensu Foldats in Lasser =Stelis fraterna 

Stelis tenuilabris Lindl. [possibly distinct species] =Stelis alata 

Stelis trichorrachis auct. non Rchb. f. 1855: sensu Dunst. & Garay =Stelis pygmaea 

Stelis trigoniflora (Sw.) Garay =Stelis ophioglossoides 

Stelis tristyla auct. non Lindl. 1838: sensu Foldats in Lasser =Stelis bangii 

Stelis velaticaulis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis velaticaulis 

Stelis vulgaris Pridgeon & M.W. Chase =Pleurothallis floribunda 

Stenia tigrina (Garay & Dunst.) Foldats =Chaubardiella tigrina 

Stenocoryne longicornis (Lindl.) Lindl. =Bifrenaria longicornis 

Stenoptera adnata Ridl. ex Oliv. [nom. superfl.] =Gomphichis adnata 

Stenoptera adnata Ridl. ex Thurn =Gomphichis adnata 

Stenoptera costaricensis Schltr. =Gomphichis costaricensis 

Stenoptera roehlii Schnee =Eurystyles cotyledon 

Stenorrhynchos goninensis Pulle =Pelexia goninensis 

Stenorrhynchos lanceolatum (Aubl.) Rich. ex Spreng. =Sacoila lanceolata 

Stenorrhynchos laxus Poepp. & Endl. =Pelexia laxa 

Stictophyllum pygmaeum (Cogn.) Dodson & M.W. Chase =Stictophyllorchis pygmaea 

Sturmia elliptica Rchb. f. =Liparis jamaicensis 

Sturmia kappleri Rchb. f. =Liparis nervosa 

Tetragamestus modestus Rchb. f. =Scaphyglottis modesta 

Teuscheria venezuelana Garay =Teuscheria wageneri 

Triaristella triglochin (Rchb. f.) Luer =Trisetella triglochin 

Triaristellina triglochin (Rchb. f.) Rauschert =Trisetella triglochin 

Trichocentrum cornucopiae Linden & Rchb. f. =Trichocentrum fuscum 

Trichocentrum hartii Rolfe =Trichocentrum fuscum 

Trichopilia albida auct. non H. Wendl. 1851: sensu Dunst. & Garay =Trichopilia aenigma 

Trichopilia fragrans auct. non (Lindl.) Rchb. f. 1858: sensu Foldats in Lasser =Trichopilia aenigma 

Trichopilia mutica (Lindl.) Rchb. f. & Wullschl. ex Rchb. f. =Leucohyle mutica 

Trichosalpinx blaisdellii auct., non (S. Watson) Luer =Trichosalpinx egleri 

Trichosalpinx foliata (Griseb.) Luer =Trichosalpinx dura 

Trichosalpinx greenwoodiana Soto Arenas =Trichosalpinx memor 

Trichosalpinx minutipetala (Ames & Schweinf.) Luer =Trichosalpinx memor 

Trichosalpinx nageliana Soto Arenas =Trichosalpinx memor 

Trichosalpinx obliquipetala (Ames & C. Schweinf.) Luer =Lepanthopsis obliquipetala 

Trichosalpinx patula Luer [nom. inval.] =Trichosalpinx patula 

Trichosalpinx zephyrina (Rchb. f.) Luer =Pleurothallis zephyrina 

Trigonidium tenue Lodd. ex Lindl. =Trigonidium acuminatum 

Trisetella huebneri (Schltr.) Luer =Trisetella triglochin 

Trizeuxis pygmaea (Cogn.) Schltr. =Stictophyllorchis pygmaea 

Uleiorchis cogniauxiana f. maior Hoehne =Uleiorchis ulaei 

Vanilla aromatica Sw. =Vanilla mexicana 

Vanilla ensifolia Rolfe =Vanilla odorata 

Vanilla inodora Schiede =Vanilla mexicana 

Vanilla pompona auct., non Schiede =Vanilla grandiflora 

Veyretia aphylla (Ridl. ex Thurn) Szlach. =Sarcoglottis aphylla 

Veyretia simplex (Griseb.) Szlach. =Sarcoglottis simplex 

Wullschlaegelia aphylla (Sw.) Rchb. f. [not in flora area] 

Wullschlaegelia ulaei Cogn. =Uleiorchis ulaei 

Xylobium steyermarkii Foldats =Bifrenaria steyermarkii 

Zygopetalum brachystalix Rchb. f. =Otostylis brachystalix 

Zygopetalum burkei Rchb. f. =Galeottia burkei 

Zygopetalum jorisianum Rolfe =Galeottia jorisiana 

Zygopetalum kegelii Rchb. f. =Zygosepalum kegelii 

Zygopetalum lindeniae Rolfe =Zygosepalum lindeniae 

Zygopetalum paludosum Cogn. =Otostylis brachystalix


Zygopetalum rostratum Hook. =Zygosepalum labiosum 

Zygopetalum tatei Ames & C. Schweinf. =Zygosepalum tatei 

Zygopetalum tricolor Lindl. =Koellensteinia tricolor 

Zygopetalum venustum Ridl. ex Oliv. =Otostylis lepida 

Zygosepalum tatei var. angustilabium C. Schweinf. =Zygosepalum angustilabium 

REFERENCES 

Ames, O., and C. Schweinfurth. 1931. Orchidaceae. In Botanical Results of the Tyler-Duida expedition, by A. Gleason. Bull. 

Torrey Bot. Club 68: 345–353. 

Anonymous. 1887. Catasetum bungerothii. Gard. Chron. ser. 3, 1: 139–140. 

Carnevali, G, I.M. Ramírez-Morillo, G.A. Romero-González, C.A. Vargas, E. Foldats. 2003. Orchidaceae. In J.A. Steyermark, 

P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 7: 200-619. 

Christenson, E.A. 1997. Orchidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 

1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. 

Memoirs of the New York Botanical Garden 76, pt. 1: 286-342. 

Cremers, G., M. Hoff, D. Barthelemy, E.A. Christenson, Ly. & L. Leneveu, & Ch. & Cl. Pawilowski. 1992. Inventaire taxonomique 

des plantes de la Guyane française II - les Orchidacées. Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Sér. Patr. Génétique) 7: 1-144. 

Dunsterville, G.C.K., and L.A. Garay. 1959–1976. Venezuelan Orchids Illustrated. 6 volumes. London: Andre Deutsch. 

Dunsterville, G.C.K., and L.A. Garay. 1979. Venezuelan Orchids: An Illustrated Field Guide. 3 volumes. Cambridge: Harvard 

University Press. 

Dunsterville, G.C.K., and E. Dunsterville 1982. Auyán-tepui: Reminiscences of an orchid search. In Orchid Botany: Reviews 

and Perspectives, edited by J. Arditti, 19–38. Ithaca: Cornell University Press. 

Foldats, E. 1969–1970. Orchidaceae. In Flora de Venezuela, edited by T. Lasser, volume 15, parts 1–5. Caracas: Instituto Botánico. 

Gleason, H.A. 1931. Botanical results of the Tyler-Duida expedition. Bull. Torrey Bot. Club 58: 277–506. 

Im Thurn, E.F. 1884a. Cattleya lawrenceana. Gard. Chron. n.s. 25: 168. 

Im Thurn, E.F. 1884b. New plants from Guiana. Timehri 3: 156–158. 

Im Thurn, E.F. 1885a. The first ascent of Roraima. Timehri 4: 1–48. 

Im Thurn, E.F. 1885b. Roraima. Timehri 4: 255–267. 

Im Thurn, E.F. 1886. Notes on the plants observed during the Roraima expedition of 1884. Timehri 5: 147–223. 

Jirak, I.L. 1968. Angel Falls Expedition 1968. Mimeo. 

Kromer, E. 1911–1912. An orchid collector’s travels through British Guiana to Brazil. Orchid World 2: 55–57, 88–90, 107–110, 

136–139, 175–177. 

Linden, L., A. Cogniaux, and G. Grignan. 1894. Les Orchidées Exotiques. Gand: Eug. Vander Haeghen. 

Reichenbach, H.G. 1885. Cattleya lawrenceana. Gard. Chron. n.s. 23: 338. 

Renz, J. 1992. The genus Habenaria in the Guianas. Candollea 47: 483-512. 

Rolfe, R.A. 1889. Catasetum bungerothii var. aureum. Gard. Chron. ser. 3, 6: 466. 

Rolfe, R.A. 1891. Peristeria aspersa. Lindenia 6: 57–58, t. 267. 

Romero-González, G.A., and G. Carnevali Fernández-Concha. 2000. Orchids of Venezuela: An Illustrated Field Guide. Second 

edition. Caracas, Venezuela: Armitano Editores. 

Sander, F. 1888. Reichenbachia. Volume 1. London: H. Sotheran. 

Schomburgk, R.H. 1931. Travels in Guiana and on the Orinoco. Translated by W.E. Roth. Georgetown, Guyana: Argosy. 

Schweinfurth, C. 1967. Orchidaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 69-214. 

Seidel, E. 1890. Voyage aux montagnes de Roraima. Orchidophile 10: 5–7, 76–77. 

Szlachetko, D.L. & Y. Veyret. 1994. Two new species of Spiranthinae (Orchidaceae) from French Guyana. Fragm. Flor. Geobot. 

39(2): 471-474. 

Ule, E. 1914. Die vegetation des Roraima. Bot. Jahrb. Syst. 52(Beibl. 115): 42–53. 

Veitch, J.H. 1906. Hortus Veitchii. London: Gilbert & Rivington. 

Werkhoven, M.C.M. 1986. Orchids of Suriname. Uitgeversmaatschappij: Paramaribo, Suriname. 

POACEAE 

G. Davidse, E.J. Judziewicz, F.O. Zuloaga 

Acroceras excavatum (Henrard) Zuloaga & Morrone 

Acroceras macrum Stapf [naturalized] 

Acroceras zizanioides (Kunth) Dandy [naturalized] 

Aegopogon cenchroides Humb. & Bonpl. ex Willd. [Mt. Roraima] 

Andropogon angustatus (J. Presl) Steud. 

Andropogon bicornis L. 

Andropogon bicornis var. bicornis 

Andropogon bicornis var. burchellii Hack. 

Andropogon carinatus Nees 

Andropogon diuturnus Sohns 

Andropogon fastigiatus Sw. 

Andropogon gayanus Kunth [cultivated, naturalized] 

BO 

GU SU FG? 

AM BO 

BO 

DA GU SU FG 

GU SU 


AM 

AM 

AM 


AM BO 

AM BO GU 

SU

150 


Andropogon glomeratus (Walter) E. Britton, Stearns & Poggenb. 

Andropogon hypogynus Hack. 

Andropogon indetonsus Sohns 

Andropogon insolitus Sohns 

Andropogon leucostachyus Kunth [expected in DA] 

Andropogon longiramosus Sohns 

Andropogon selloanus (Hack.) Hack. [expected in DA] 

Andropogon vetus Sohns 

Andropogon virgatus Desv. [expected in DA] 

Anthephora hermaphrodita (L.) Kuntze 

Arberella aff. bahiensis Soderstr. & Zuloaga 

Arberella flaccida (Döll) Soderstr. & C.E. Calderón 

Arberella venezuelae Judz. & Davidse 

Aristida adscensionis L. 

Aristida capillacea Lam. 

Aristida gibbosa (Nees) Kunth 

Aristida longifolia Trin. 

Aristida moritzii Henrard 

Aristida recurvata Kunth 

Aristida riparia Trin. 

Aristida setifolia Kunth 

Aristida ternipes Cav. 

Aristida torta (Nees) Kunth 

Arthrostylidium pubescens Rupr. 

Arthrostylidium scandens McClure 

Arthrostylidium schomburgkii (Benn.) Munro 

Arthrostylidium sp. A 

Arthrostylidium sp. B 

Arthrostylidium sp. C 

Arthrostylidium sp. D 

Arthrostylidium venezuelae (Steud.) McClure 

Arundinella hispida (Humb. & Bonpl. ex Willd.) Kuntze 

Arundo donax L. [cultivated and naturalized] 

Atractantha amazonica Judz. & L.G. Clark 

Aulonemia chimantaensis Judz. & Davidse 

Aulonemia deflexa (N.E. Br.) McClure 

Aulonemia jauaensis Judz. & Davidse 

Aulonemia nitida Judz. 

Aulonemia sp. A 

Aulonemia sp. B 

Aulonemia sp. C 

Aulonemia aff. subpectinata (Kuntze) McClure 

Axonopus anceps (Mez) Hitchc. 

Axonopus arundinaceus G.A. Black 

Axonopus aureus P. Beauv. 

Axonopus canescens (Nees ex Trin.) Pilg. 

Axonopus capillaris (Lam.) Chase 

Axonopus casiquiarensis Davidse 

Axonopus casiquiarensis var. A 

Axonopus caulescens (Mez) Henrard 

Axonopus chimantensis Davidse 

Axonopus chrysoblepharis (Lag.) Chase [expected in BO] 

Axonopus chrysostachyus (Schrad.) Pilg. 

Axonopus compressus (Sw.) P. Beauv. 

Axonopus cuatrecasasii G.A. Black 

Axonopus eminens (Nees) G.A. Black 

Axonopus equitans Hitchc. & Chase 

Axonopus excavatus (Nees ex Trin.) Henrard 

Axonopus fissifolius (Raddi) Kuhlm. 

Axonopus flabelliformis Swallen 

Axonopus gracilis G.A. Black 

Axonopus iridifolius (Poepp.) G.A. Black 

Axonopus leptostachyus (Flüggé) Hitchc. 

Axonopus longispicus (Döll) Kuhlm. 

GU? 

AM BO 

BO 

BO 

AM 

AM 

BO GU SU FG 


AM BO 


AM 

BO FG 

SU FG 

AM SU FG 

AM BO GU? 


BO GU 

AM BO 

BO 

GU SU FG 



BO 

BO 

GU SU 

AM BO 

BO 

GU SU FG 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

GU SU 

AM BO GU 


AM 

BO 

SU FG? 

AM 

AM 

AM 

BO 

BO 

GU 

GU 

AM BO 

AM 

AM 




AM 

AM 

AM 

BO GU SU FG 

BO 

BO 

GU 

SU 

FG 

AM 

AM 


BO GU SU 

AM GU SU FG 

AM BO FG 


AM 

AM 

BO GU SU FG 

AM BO GU 


AM BO GU SU FG


Axonopus magallanesiae Giraldo-CaZas 

Axonopus oiapocensis G.A. Black 

Axonopus passourae G.A. Black 

Axonopus pennellii G.A. Black 

Axonopus pubivaginatus Henrard 

Axonopus purpusii (Mez) Chase 

Axonopus ramosus Swallen [expected in GU and adjacent Brazil] 

Axonopus schultesii G.A. Black 

Axonopus senescens (Döll) Henrard 

Axonopus sp. A 

Axonopus sp. B 

Axonopus sp. C 

Axonopus steyermarkii Swallen 

Axonopus suffultiformis G.A. Black 

Axonopus suffultiformis var. A 

Axonopus surinamensis (Hochst. ex Steud.) Henrard 

Axonopus triglochinoides (Mez) Dedecca 

Axonopus villosus Swallen 

Axonopus yutajensis G.A. Black 

Bothriochloa bladhii (Retz.) S.T. Blake [naturalized] 

Bouteloua americana (L.) Scribn. [introduced? no recent collections] 

Brachiaria arrecta (Hack. ex T. Durand & Schinz) Stent [culitvated, 

naturalized?] 

Brachiaria decumbens Stapf 

Brachiaria fasciculata (Sw.) Parodi 

Brachiaria humidicola (Rendle) Schweick. 

Brachiaria mollis (Sw.) Parodi 

Brachiaria mutica (Forssk.) Stapf [naturalized] 

Brachiaria plantaginea (Link) Hitchc. 

Briza minor L. 

Cenchrus brownii Roem. & Schult. 

Cenchrus echinatus L. 

Chloris barbata Sw. 

Chloris elata Desv. 

Chloris radiata (L.) Sw. 

Chusquea linearis N.E. Br. 

Chusquea sp. A 

Coelorachis aurita (Steud.) A. Camus 

Coix lacryma-jobi L. [cultivated, naturalized] 

Cortaderia roraimensis (N.E. Br.) Pilg. 

Cryptochloa capillata (Trin.) Soderstr. 

Ctenium cirrosum (Nees) Kunth 

Cymbopogon citratus (DC.) Stapf [cultivated, naturalized] 

Cynodon dactylon (L.) Pers. 

Dactyloctenium aegyptium (L.) Willd. 

Dichanthelium aequivaginatum (Swallen) Zuloaga 

Dichanthelium davidsei (Zuloaga & Morrone) Zuloaga 

Dichanthelium hebotes (Trin.) Zuloaga 

Dichanthelium pycnoclados (Tutin) Davidse 

Dichanthelium sciurotoides (Zuloaga & Morrone) Davidse 

Dichanthelium telmatum (Swallen) Zuloaga [expected in GU] 

Dichanthium annulatum (Forssk.) Stapf [cultivated, naturalized] 

Dichanthium aristatum (Poir.) C.E. Hubb. [cultivated, naturalized] 

Digitaria ciliaris (Retz.) Koeler 

Digitaria decumbens Stent [cultivated, naturalized] 

Digitaria fragilis (Steud.) Luces 

Digitaria fuscescens (J. Presl) Henrard [naturalized] 

Digitaria horizontalis Willd. 

Digitaria insularis (L.) Fedde 

Digitaria lanuginosa (Nees) Henrard 

Digitaria laxa (Rchb. ex A. Spreng.) Parodi 

Digitaria longiflora (Retz.) Pers. [naturalized?] 

Digitaria nervalis Henrard 

Digitaria nuda Schumach. [naturalized?] 

BO 

BO 

FG 

FG 

GU SU FG 


AM SU FG 

AM 

FG 

AM BO 

AM 

BO 

AM 

AM BO GU 

AM 

BO GU SU FG 

AM 

AM 

AM 

GU SU 

BO GU 

BO GU SU FG 

AM GU SU 

BO GU SU FG 

BO GU SU FG 

BO GU FG 


DA FG 

BO 

BO GU SU FG 



BO GU SU 


AM BO GU 

AM 



AM BO GU 

FG 

SU 

GU SU FG 


BO GU SU FG 


BO 

BO 

BO GU SU 

BO GU FG 

BO 

GU 

GU 


GU SU 

GU 



BO GU SU FG 

GU FG 

GU SU FG 

GU SU 

AM 

AM BO GU SU FG

152 


Digitaria setigera Roth ex Roem. & Schult. [naturalized] 

Digitaria swazilandensis Stent [cultivated, naturalized] 

Digitaria tenuis (Nees) Henrard 

Digitaria venezuelae Henrard 

Digitaria violascens Link [cultivated, naturalized] 

Echinochloa colona (L.) Link 

Echinochloa crus-pavonis (Kunth) Schult. 

Echinochloa polystachya (Kunth) Hitchc. 

Echinochloa pyramidalis (Lam.) Hitchc. & Chase [cultivated, naturalized] 

Echinolaena gracilis Swallen 

Echinolaena inflexa (Poir.) Chase 

Eleusine indica (L.) Gaertn. [naturalized] 

Elionurus muticus (Spreng.) Kuntze 

Enteropogon mollis (Nees) Clayton 

Eragrostis acutiflora (Kunth) Nees 

Eragrostis airoides Nees [expected in GU] 

Eragrostis amabilis (L.) Wight & Arn. ex Nees [naturalized] 

Eragrostis atrovirens (Desf.) Trin. ex Steud. 

Eragrostis ciliaris (L.) R. Br. [naturalized] 

Eragrostis ciliaris var. ciliaris 

Eragrostis gangetica (Roxb.) Steud. 

Eragrostis guianensis Hitchc. 

Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 

Eragrostis japonica (Thunb.) Trin. [naturalized] 

Eragrostis maypurensis (Kunth) Steud. 

Eragrostis pectinacea (Michx.) Nees 

Eragrostis pectinacea var. pectinacea 

Eragrostis pilosa (L.) P. Beauv. [naturalized] 

Eragrostis plurinodis Swallen ex Luces 

Eragrostis polytricha Nees 

Eragrostis refracta (Muhl. ex Elliott) Scribn. 

Eragrostis riobrancensis Judz. & P.M. Peterson 

Eragrostis scaligera Salzm. ex Steud. 

Eragrostis secundiflora J. Presl [expected in SU] 

Eragrostis tephrosanthos Schult. 

Eragrostis unioloides (Retz.) Nees ex Steud. [naturalized] 

Eragrostis viscosa (Retz.) Trin. 

Eriochloa distachya Kunth 

Eriochloa polystachya Kunth 

Eriochloa procera (Retz.) C.E. Hubb. [naturalized] 

Eriochloa punctata (L.) Desv. ex Ham. 

Eriochrysis cayennensis P. Beauv. 

Eriochrysis aff. holcoides (Nees) Kuhlm. 

Froesiochloa boutelouoides G.A. Black 

Gouinia latifolia (Griseb.) Vasey 

Guadua angustifolia Kunth 

Guadua ciliata LondoZo & Davidse 

Guadua fascicularis Döll 

Guadua aff. glomerata Munro 

Guadua latifolia (Bonpl.) Kunth 

Guadua macrostachya Rupr. [expected in SU] 

Guadua paniculata Munro [expected in GU] 

Guadua superba Huber 

Guadua venezuelae Munro 

Guadua weberbaueri Pilg. 

Gymnopogon fastigiatus Nees 

Gymnopogon foliosus (Willd.) Nees 

Gymnopogon spicatus (Spreng.) Kuntze 

Gynerium sagittatum (Aubl.) P. Beauv. 

Hackelochloa granularis (L.) Kuntze [naturalized] 

Hemarthria altissima (Poir.) Stapf & C.E. Hubb. [naturalized] 

Heteropogon contortus (L.) P. Beauv. ex Roem. & Schult. 

Heteropogon melanocarpus (Elliott) Elliott ex Benth. [expected in GU] 

Homolepis aturensis (Kunth) Chase [naturalized?] 

GU SU FG 

SU 

BO GU 

BO SU FG 

BO GU SU FG 




FG 

AM BO 




AM 



BO 


AM BO DA 

BO 

AM BO GU FG 


AM BO DA GU 


AM BO SU FG 

AM BO SU FG 

AM BO GU FG 

GU 

BO GU 

SU 

GU 

FG 

GU 

GU SU FG 

BO GU SU FG 

BO 

AM BO 

GU SU FG 

GU SU 



AM BO 

FG 

BO 

AM GU 

AM 

BO 

BO DA GU FG 


FG 

GU SU 

BO DA 

AM GU 

AM BO 


BO GU 



GU FG 

BO GU 

FG 

AM BO GU SU FG


Homolepis glutinosa (Sw.) Zuloaga & Soderstr. 

Homolepis isocalycia (G. Mey.) Chase 

Hymenachne amplexicaulis (Rudge) Nees 

Hymenachne donacifolia (Raddi) Chase 

Hyparrhenia bracteata (Willd.) Stapf 

Hyparrhenia rufa (Nees) Stapf [cultivated, naturalized] 

Ichnanthus breviscrobs Döll 

Ichnanthus calvescens (Nees) Döll 

Ichnanthus dasycoleus Tutin 

Ichnanthus ephemeroblepharis G.A. Black & Fróes ex G.A. Black & Pires 

Ichnanthus hoffmannseggii (Roem. & Schult.) Döll 

Ichnanthus lancifolius Mez 

Ichnanthus lancifolius var. weberbaueri (Mez) Stieber 

Ichnanthus leiocarpus (Spreng.) Kunth 

Ichnanthus nemoralis (Schrad. ex Schult.) Hitchc. & Chase 

Ichnanthus nemorosus (Sw.) Döll [Sierra de la Neblina] 

Ichnanthus pallens (Sw.) Munro ex Benth. 

Ichnanthus panicoides P. Beauv. 

Ichnanthus procurrens (Nees ex Trin.) Swallen 

Ichnanthus ruprechtii Döll 

Ichnanthus tenuis (J. Presl) Hitchc. & Chase 

Imperata brasiliensis Trin. 

Imperata contracta (Kunth) Hitchc. 

Isachne polygonoides (Lam.) Döll 

Isachne rigens (Sw.) Trin. 

Ischaemum arenosum Sohns 

Ischaemum guianense Kunth ex Hack. 

Ischaemum indicum (Houtt.) Merr. [cultivated, naturalized] 

Ischaemum latifolium (Spreng.) Kunth 

Ischaemum rugosum Salisb. [naturalized] 

Ischaemum timorense Kunth [cultivated, naturalized] 

Ixophorus unisetus (J. Presl) Schltdl. [cultivated but expected in GU] 

Lasiacis anomala Hitchc. 

Lasiacis ligulata Hitchc. & Chase 

Lasiacis nigra Davidse 

Lasiacis procerrima (Hack.) Hitchc. 

Lasiacis ruscifolia (Kunth) Hitchc. 

Lasiacis scabrior Hitchc. 

Lasiacis sloanei (Griseb.) Hitchc. 

Lasiacis sorghoidea (Desv. ex Ham.) Hitchc. & Chase 

Leersia hexandra Sw. 

Leersia ligularis Trin. [expected in GU] 

Leptochloa filiformis (Lam.) P. Beauv. 

Leptochloa scabra Nees 

Leptochloa virgata (L.) P. Beauv. 

Leptocoryphium lanatum (Kunth) Nees 

Lithachne pauciflora (Sw.) P. Beauv. [expected in GU] 

Luziola bahiensis (Steud.) Hitchc. 

Luziola doelliana Prodoehl 

Luziola subintegra Swallen 

Melinis minutiflora P. Beauv. [cultivated, naturalized] 

Merostachys maguireorum McClure 

Merostachys aff. retrorsa McClure 

Mesosetum cayennense Steud. 

Mesosetum chaseae Luces 

Mesosetum chlorostachyum (Döll) Chase 

Mesosetum filifolium Hubb. 

Mesosetum loliiforme (Hochst. ex Steud.) Chase 

Mesosetum rottboellioides (Kunth) Hitchc. 

Microchloa indica (L. f.) P. Beauv. [introduced species expected in GU] 

Myriocladus cardonae Swallen 

Myriocladus churunensis Swallen 

Myriocladus distantiflorus Swallen [other species of this genus expected as 

well] 

BO GU FG? 




BO GU SU 




GU SU FG 

AM BO 

FG 

AM 

AM 

GU SU FG 


AM 



BO 

BO GU SU FG 



BO GU SU FG 


AM BO GU 

BO 


GU SU 

AM 

AM GU SU FG 

GU SU FG 



BO GU 

AM BO GU 

AM GU 

AM 

BO 



BO 

BO GU 




BO SU FG 

BO GU SU FG 

BO 


AM BO GU SU 

BO 

AM BO GU 

BO GU SU FG 

BO GU 

AM 

AM BO 



BO 

BO 

BO GU

154 

Myriocladus exsertus Swallen 

Myriocladus grandifolius Swallen 

Myriocladus involutus Judz. & Davidse 

Myriocladus longiramosus Swallen 

Myriocladus neblinaensis Swallen 

Myriocladus paludicolus Swallen [Sierra de la Neblina] 

Myriocladus simplex Swallen 

Myriocladus sp. A 

Myriocladus steyermarkii Swallen 

Myriocladus virgatus Swallen 

Neurolepis angusta Swallen 

Neurolepis diversiglumis Soderstr. [Sierra de la Neblina] 

Neurolepis glomerata Swallen 

Neurolepis pittieri McClure 

Neurolepis sp. A 

Olyra caudata Trin. 

Olyra ciliatifolia Raddi 

Olyra ecaudata Döll 

Olyra latifolia L. 

Olyra longifolia Kunth 

Olyra obliquifolia Steud. 

Olyra standleyi Hitchc. 

Olyra wurdackii Swallen 

Oplismenus burmannii (Retz.) P. Beauv. 

Oplismenus hirtellus (L.) P. Beauv. 

Orthoclada laxa (Rich.) P. Beauv. 

Oryza grandiglumis (Döll) Prodoehl 

Oryza latifolia Desv. 

Oryza rufipogon Griff. [naturalized] 

Oryza sativa L. [cultivated, occasionally escaping] 

Otachyrium grandiflorum Send. & Soderstr. 

Otachyrium succisum (Swallen) Send. & Soderstr. 

Otachyrium versicolor (Döll) Henrard 

Panicum altum Hitchc. & Chase 

Panicum arctum Swallen 

Panicum bergii Arechav. 

Panicum caricoides Nees 

Panicum cayennense Lam. 

Panicum cervicatum Chase [expected in GU] 

Panicum chnoodes Trin. 

Panicum cordovense E. Fourn. 

Panicum cyanescens Nees ex Trin. 

Panicum deciduum Swallen 

Panicum dichotomiflorum Michx. 

Panicum discrepans Döll 

Panicum elephantipes Nees ex Trin. 

Panicum eligulatum N.E. Br. 

Panicum fontanale Swallen 

Panicum fonticolum Swallen 

Panicum grande Hitchc. & Chase 

Panicum granuliferum Kunth 

Panicum haenkeanum J. Presl 

Panicum hirsutum Sw. 

Panicum hirtum Lam. 

Panicum hispidifolium Swallen 

Panicum hylaeicum Mez 

Panicum ichunense Swallen 

Panicum jauanum Davidse 

Panicum ligulare Nees ex Trin. 

Panicum maximum Jacq. [naturalized] 

Panicum mertensii Roth 

Panicum micranthum Kunth 

Panicum miliaceum L. [cultivated; naturalized?] 

Panicum millegrana Poir. 


AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

AM BO 

BO 

AM BO 

BO GU 

AM 

AM 

BO 

BO 

BO 

BO GU SU FG 

BO GU FG 




AM 

AM 

BO 

SU FG 



FG 



AM 

AM 

BO DA 

GU SU FG 

AM BO GU 


BO GU 

GU 

SU FG 



AM BO 

AM BO GU 

BO GU SU FG 

AM 

AM 

BO GU SU FG 

DA GU SU FG 



AM 

AM 

BO 

BO 

GU 


AM BO 

BO 

GU FG 

DA GU 

AM BO 

BO 

GU SU FG 

AM 

BO 

BO 

BO 

GU SU FG 




GU FG 

BO GU SU FG

Panicum nervosum Lam. 

Panicum olyroides Kunth 

Panicum orinocanum Luces 

Panicum pandum Swallen 

Panicum pantrichum Hack. [expected in GU] 

Panicum parvifolium Lam. 

Panicum petrense Swallen 

Panicum pilosum Sw. 

Panicum polycomum Trin. 

Panicum polygonatum Schrad. 

Panicum pulchellum Raddi 

Panicum pyrularium Hitchc. & Chase 

Panicum rivale Swallen 

Panicum rudgei Roem. & Schult. 

Panicum scabridum Döll 

Panicum schwackeanum Mez 

Panicum sellowii Nees 

Panicum sipapoense Swallen 

Panicum stenodes Griseb. 

Panicum steyermarkii Swallen 

Panicum stoloniferum Poir. 

Panicum tepuianum Davidse & Zuloaga 

Panicum trichanthum Nees 

Panicum trichocladum Hack. ex K. Schum. [naturalized] 

Panicum trichoides Sw. 

Panicum tricholaenoides Steud. 

Panicum yavitaense Swallen 

Pappophorum mucronulatum Nees 

Pappophorum pappiferum (Lam.) Kuntze [naturalized] 

Paratheria prostrata Griseb. 

Pariana campestris Aubl. 

Pariana radiciflora Sagot ex Döll 

Parodiolyra lateralis (J. Presl ex Nees) Soderstr. & Zuloaga 

Parodiolyra luetzelburgii (Pilg.) Soderstr. & Zuloaga 

Parodiolyra micrantha (Kunth) Davidse & Zuloaga 

Paspalidium geminatum (Forssk.) Stapf 

Paspalum albidulum Henrard 

Paspalum altsonii Chase 

Paspalum ammodes Trin. 

Paspalum apiculatum Döll 

Paspalum arenarium Schrad. 

Paspalum arundinaceum Poir. 

Paspalum aspidiotes Trin. 

Paspalum atabapense Davidse & Zuloaga 

Paspalum bifidifolium Soderstr. 

Paspalum boscianum Flüggé 

Paspalum carinatum Humb. & Bonpl. ex Flüggé 

Paspalum chaffanjonii Maury 

Paspalum cinerascens (Döll) A.G. Burm. & M.N.C. Bastos 

Paspalum clavuliferum C. Wright 

Paspalum conjugatum P.J. Bergius 

Paspalum conspersum Schrad. 

Paspalum convexum Humb. & Bonpl. ex Flüggé 

Paspalum corcovadense Raddi 

Paspalum coryphaeum Trin. 

Paspalum decumbens Sw. 

Paspalum delicatum Swallen 

Paspalum densum Poir. 

Paspalum dilatatum Poir. [cultivated, naturalized] 

Paspalum distichum L. 

Paspalum exaltatum J. Presl 

Paspalum fasciculatum Willd. ex Flüggé [cultivated; naturalized] 

Paspalum fimbriatum Kunth [introduced, no recent collections] 

Paspalum gardnerianum Nees 



BO GU SU FG 

AM 

AM BO GU 

BO FG 


AM 



AM BO GU FG 

AM 


BO GU SU FG 



GU SU FG 

BO GU 

AM BO 


BO 


AM 

GU SU FG 

SU 

BO GU SU FG 

AM 

AM 

BO 

GU 

AM GU FG 

GU? SU FG 

AM GU SU FG 

AM BO GU SU 



BO GU SU FG 

SU 

AM BO GU SU 

BO GU 

BO 

BO GU SU FG 

FG 

AM BO GU 

AM 

GU 

GU SU FG 

AM BO GU SU 

BO 

SU FG 

BO GU 


AM GU SU FG 


BO GU 



AM FG 


GU SU 

GU SU FG? 

GU 

AM DA SU FG 

GU? 

AM BO DA GU SU FG

156 


Paspalum goeldii Swallen 

Paspalum hyalinum Nees ex Trin. 

Paspalum intermedium Munro ex Morong & Britton 

Paspalum lacustre Swallen [records erroneous, but perhaps expected] 

Paspalum lanciflorum Trin. 

Paspalum laxum Lam. 

Paspalum lineare Trin. 

Paspalum loefgrenii Ekman 

Paspalum maculosum Trin. 

Paspalum maritimum Trin. 

Paspalum melanospermum Desv. ex Poir. 

Paspalum millegrana Schrad. 

Paspalum morichalense Davidse, Zuloaga & Filg. [specimens previously 

identified as Paspalum lacustre] 

Paspalum multicaule Poir. 

Paspalum multinervium A.G. Burm. 

Paspalum notatum Flüggé 

Paspalum nudatum Luces 

Paspalum nutans Lam. 

Paspalum oligostachyum Salzm. ex Steud. 

Paspalum orbiculatum Poir. 

Paspalum paniculatum L. 

Paspalum parviflorum Rhode ex Flüggé 

Paspalum pectinatum Nees ex Trin. 

Paspalum petilum Chase 

Paspalum pictum Ekman 

Paspalum pilosum Lam. 

Paspalum plicatulum Michx. 

Paspalum pulchellum Kunth 

Paspalum pumilum Nees 

Paspalum repens P.J. Bergius 

Paspalum riparium Nees 

Paspalum saccharoides Nees ex Trin. 

Paspalum scrobiculatum L. [cultivated; naturalized?] 

Paspalum serpentinum Hochst. ex Steud. 

Paspalum stellatum Humb. & Bonpl. ex Flüggé [expected in GU] 

Paspalum subciliatum Chase 

Paspalum subfalcatum (Döll) Tutin 

Paspalum tillettii Davidse & Zuloaga 

Paspalum trinii Swallen 

Paspalum vaginatum Sw. 

Paspalum virgatum L. 

Paspalum wrightii Hitchc. & Chase 

Pennisetum ciliare (L.) Link [cultivated; naturalized?] 

Pennisetum polystachion (L.) Schult. 

Pennisetum purpureum Schumach. [cultivated, naturalized] 

Pharus lappulaceus Aubl. 

Pharus latifolius L. 

Pharus mezii Prodoehl 

Pharus parvifolius Nash 

Pharus parvifolius subsp. elongatus Judz. 

Pharus parvifolius subsp. parvifolius 

Pharus virescens Döll 

Phragmites australis (Cav.) Trin. ex Steud. 

Piresia goeldii Swallen 

Piresia macrophylla Soderstr. 

Piresia sympodica (Döll) Swallen 

Pseudechinolaena polystachya (Kunth) Stapf 

Raddia guianensis (Brongn.) Hitchc. 

Raddiella esenbeckii (Steud.) C.E. Calderón & Soderstr. 

Raddiella aff. kaieteurana Soderstr. 

Raddiella aff. potaroensis Soderstr. 

Rehia nervata (Swallen) Fijten 

Reimarochloa aberrans (Döll) Chase 

GU SU FG 


BO 

AM BO GU SU 

FG 

AM BO 

GU 

GU 

GU SU 

GU SU FG 



AM BO SU 


FG 

GU SU FG 

BO GU SU 


SU FG 





AM BO GU 

AM GU 




GU SU FG 

GU SU FG 

GU SU FG 

BO 

SU FG 

GU SU FG 

BO 

BO GU FG 

AM 

AM 

GU 

DA GU SU FG 


GU 

GU 


AM GU FG 



BO 

DA GU SU FG 

GU 

DA GU SU FG 


BO GU SU FG 

SU FG 

FG 

BO GU SU FG 

GU 

SU FG 


BO GU SU 

BO GU 

GU SU FG 

FG


Reimarochloa acuta (Flüggé) Hitchc. 

Rhipidocladum aff. maxonii (Hitchc.) McClure 

Rhipidocladum aff. racemiflorum (Steud.) McClure 

Rhipidocladum sibilans Davidse, Judz. & L.G. Clark [Mt. Roraima] 

Rhipidocladum sp. A 

Rhipidocladum sp. B 

Rhynchelytrum repens (Willd.) C.E. Hubb. [naturalized] 

Rhytachne gonzalezii Davidse 

Rhytachne guianensis (Hitchc.) Clayton 

Rhytachne rottboellioides Desv. ex Ham. 

Rhytachne subgibbosa (Winkl. ex Hack.) Clayton 

Rottboellia cochinchinensis (Lour.) Clayton [naturalized] 

Saccharum asperum (Nees) Steud. [expected in GU] 

Saccharum trinii (Hack.) Renvoize 

Sacciolepis angustissima (Hochst. ex Steud.) Kuhlm. 

Sacciolepis indica (L.) Chase [naturalized?] 

Sacciolepis myuros (Lam.) Chase 

Sacciolepis otachyrioides Judz. 

Sacciolepis striata (L.) Nash 

Sacciolepis vilvoides (Trin.) Chase 

Schizachyrium brevifolium (Sw.) Nees ex Büse 

Schizachyrium condensatum (Kunth) Nees 

Schizachyrium maclaudii (Jacq.-Fél.) S.T. Blake 

Schizachyrium sanguineum (Retz.) Alston 

Schizachyrium tenerum Nees 

Setaria barbata (Lam.) Kunth [naturalized] 

Setaria italica (L.) P. Beauv. [naturalized?] 

Setaria macrostachya Kunth 

Setaria magna Griseb. 

Setaria palmifolia (J. König) Stapf [cultivated, native to tropical S. Asia] 

Setaria parviflora (Poir.) Kerguélen 

Setaria setosa (Sw.) P. Beauv. [naturalized?] 

Setaria sulcata Raddi 

Setaria tenacissima Schrad. ex Schult. 

Setaria tenax (Rich.) Desv. 

Setaria vulpiseta (Lam.) Roem. & Schult. 

Sorghastrum setosum (Griseb.) Hitchc. 

Sorghastrum stipoides (Kunth) Nash 

Sorghum bicolor (L.) Moench [cultivated, sparingly naturalized] 

Sorghum halepense (L.) Pers. [cultivated, naturalized] 

Spartina alterniflora Loisel. 

Spheneria kegelii (Müll. Berol.) Pilg. 

Sporobolus ciliatus J. Presl 

Sporobolus cubensis Hitchc. 

Sporobolus jacquemontii Kunth 

Sporobolus pyramidatus (Lam.) Hitchc. 

Sporobolus temomairemensis Judz. & P.M. Peterson 

Sporobolus tenuissimus (Mart. ex Schrank) Kuntze 

Sporobolus virginicus (L.) Kunth 

Steinchisma decipiens (Nees ex Trin.) W.V. Br. 

Steinchisma laxa (Sw.) Zuloaga 

Steinchisma stenophylla (Hack.) Zuloaga & Morrone 

Steirachne barbata (Trin.) Renvoize 

Stenotaphrum secundatum (Walter) Kuntze 

Steyermarkochloa angustifolia (Spreng.) Judz. 

Streptochaeta spicata Schrad. ex Nees [expected in GU] 

Streptogyna americana C.E. Hubb. 

Streptostachys asperifolia Desv. 

Thrasya achlysophila Soderstr. 

Thrasya axillaris (Swallen) A.G. Burm. ex Judz. [expected in GU] 

Thrasya granitica A.G. Burm. 

Thrasya hitchcockii Chase 

Thrasya paspaloides Kunth 

Thrasya petrosa (Trin.) Chase 

GU SU FG 

GU 

AM BO GU? SU 

BO 

AM BO 

BO 

AM BO DA GU 

SU 

AM BO GU SU 

BO 

AM BO 

SU FG 

BO 

BO GU SU 


GU 


AM GU 

DA GU SU FG 

DA GU SU FG 



GU SU FG 


AM BO GU 

FG 

FG 

BO 

SU FG 

GU SU 


FG 

BO GU SU FG 

BO 


BO GU SU FG 

AM BO GU FG 

GU SU FG 

AM GU SU FG 

GU FG 

DA GU SU FG 

GU SU 

GU SU FG 



SU FG 

SU 

BO GU SU FG 

GU SU FG 

BO 


AM SU 

BO GU 

GU SU FG 

AM 


BO GU SU FG 

GU 

AM BO SU 

SU 

FG 

AM BO 

AM BO GU SU

158 


Thrasya robusta Hitchc. & Chase 

Thrasya scandens (Tutin) Soderstr. ex A.G. Burm. 

Thrasya stricta A.G. Burm. 

Thrasya trinitensis Mez 

Trachypogon spicatus (L. f.) Kuntze 

Tridens flaccidus (Döll) Parodi 

Tripogon spicatus (Nees) Ekman [expected in GU] 

Tripsacum andersonii J.R. Gray [introduced] 

Tripsacum australe H.C. Cutler & E.S. Anderson 

Tripsacum dactyloides (L.) L. 

Urochloa reptans (L.) Stapf [naturalized] 

Vetiveria zizanioides (L.) Nash [cultivated; naturalized?] 

Zea luxurians (Durieu & Asch.) Bird [cultivated or escaped from cultivation] 

Zeugites sp. A 


Acicarpa sacchariflora Raddi =Digitaria insularis 

Aegopogon cenchroides Kunth =Aegopogon cenchroides 

Agrosticula muralis Raddi =Sporobolus tenuissimus 

Agrostis brasiliensis Spreng. =Reimarochloa acuta 

Agrostis compressa (Sw.) Poir. =Axonopus compressus 

Agrostis littoralis Lam. =Sporobolus virginicus 

Agrostis minutiflora (Trin.) Desf. ex Steud. =Sporobolus tenuissimus 

Agrostis radiata L. =Chloris radiata 

Agrostis villosa Rchb. ex A. Spreng. =Sporobolus ciliatus 

Aira laxa Rich. =Orthoclada laxa 

Anastrophus capillaris (Lam.) Nash =Axonopus capillaris 

Anastrophus platycaulis (Poir.) Nash ex Small =Axonopus compressus 

Anatherum holcoides Nees =Eriochrysis holcoides 

Anatherum zizanioides (L.) Hitchc. & Chase =Vetiveria zizanioides 

Andropogon adustus Trin. =Elionurus muticus 

Andropogon angustifolius Kunth =Trachypogon spicatus 

Andropogon barbatus L. =Chloris elata 

Andropogon barbatus L., 1771 =Chloris barbata 

Andropogon bracteatus Humb. & Bonpl. ex Willd. =Hyparrhenia bracteata 

Andropogon brevifolius Sw. =Schizachyrium brevifolium 

Andropogon canescens (Nees) Kunth =Trachypogon spicatus 

Andropogon condensatus Kunth =Schizachyrium condensatum 

Andropogon contortus L. =Heteropogon contortus 

Andropogon crassus Sohns =Andropogon vetus 

Andropogon crucianus Renvoize =Andropogon insolitus 

Andropogon dactyloides Steud. =Trachypogon spicatus 

Andropogon dissolutus (Nees) Steud. =Trachypogon spicatus 

Andropogon hirtiflorus (Nees) Kunth =Schizachyrium sanguineum 

Andropogon hispidus Humb. & Bonp. ex Willd. =Arundinella hispida 

Andropogon insularis L. =Digitaria insularis 

Andropogon latifolius Spreng. =Ischaemum latifolium 

Andropogon leucostachyus subsp. selloanus Hack. =Andropogon selloanus 

Andropogon ligularis (Nees) Kunth =Trachypogon spicatus 

Andropogon macroglossus (Trin.) Steud. =Trachypogon spicatus 

Andropogon microstachyum Desv. =Schizachyrium condensatum 

Andropogon mollis (Nees) Kunth =Trachypogon spicatus 

Andropogon montufarii Kunth =Trachypogon spicatus 

Andropogon nodosus (Willemet) Nash =Dichanthium aristatum 

Andropogon perdignus Sohns =Andropogon vetus 

Andropogon plumosus Humb. & Bonpl. ex Willd. =Trachypogon spicatus 

Andropogon polydactylon L. [nom. superfl.] =Chloris elata 

Andropogon riedelii Trin. =Schizachyrium sanguineum 

Andropogon secundus (J. Presl) Kunth =Trachypogon spicatus 

Andropogon segetum (Trin.) Steud. =Ischaemum rugosum 

Andropogon semiberbis (Nees) Kunth =Schizachyrium sanguineum 

Andropogon setosus Griseb. =Sorghastrum setosum 

Andropogon sorghum (L.) Brot. =Sorghum bicolor 

Andropogon spicatus Steud. =Trachypogon spicatus 

BO SU FG 

GU SU FG 

AM 

AM BO GU 


BO GU 

AM BO 

GU SU FG 

AM GU SU FG 

GU SU FG 

GU FG 

GU SU FG 

FG 

AM BO


Andropogon squarrosus sensu Pulle non L. f. =Vetiveria zizanioides 

Andropogon stipoides (J. Presl) Kunth =Trachypogon spicatus 

Andropogon tener (Nees) Kunth =Schizachyrium tenerum 

Andropogon trichospirus Hack. =Trachypogon spicatus 

Andropogon truncatus (Nees) Steud. =Trachypogon spicatus 

Andropogon vestitus Steud. =Trachypogon spicatus 

Anthaenantia lanata (Kunth) Benth. =Leptocoryphium lanatum 

Apluda zeugites sensu Aubl. non L. =Ichnanthus pallens 

Aristida adscensionis var. bromoides (Kunth) Henrard =Aristida adscensionis 

Aristida adscensionis subsp. bromoides (Kunth) Henrard =Aristida adscensionis 

Aristida adscensionis var. coarctata (Kunth) Kuntze =Aristida adscensionis 

Aristida americana L. =Bouteloua americana 

Aristida arenaria Trin. =Aristida setifolia 

Aristida bromoides Kunth =Aristida adscensionis 

Aristida coarctata Kunth =Aristida adscensionis 

Aristida dispersa var. humilis (Kunth) Trin. & Rupr. =Aristida adscensionis 

Aristida doelliana Henrard =Aristida setifolia 

Aristida elegans Rudge =Aristida capillacea 

Aristida humilis Kunth =Aristida adscensionis 

Aristida implexa var. aequa Trin. & Rupr. =Aristida riparia 

Aristida marginalis Ekman =Aristida gibbosa 

Aristida neesiana Trin. & Rupr. =Aristida recurvata 

Aristida orizabensis E. Fourn. =Aristida gibbosa 

Aristida pittieri Henrard =Aristida moritzii 

Aristida planifolia Swallen =Aristida riparia 

Aristida riedeliana Trin. & Rupr. =Aristida recurvata 

Aristida scabra (Kunth) Kunth =Aristida ternipes 

Aristida setifolia var. arenaria (Trin.) Trin. & Rupr. =Aristida setifolia 

Aristida setifolia var. grandiflora Trin. & Rupr. =Aristida setifolia 

Aristida setifolia var. intermedia Trin. & Rupr. =Aristida setifolia 

Aristida tarapotana Mez =Aristida setifolia 

Aristida tenuis (Kunth) Kunth =Aristida ternipes 

Aristida tincta Trin. & Rupr. =Aristida torta 

Arthrostylidium cacuminis McClure =Arthrostylidium scandens 

Arthrostylidium capillifolium Griseb. [West Indian species, not in flora area] =Arthrostylidium farctum 

Arthrostylidium farctum (Aubl.) Soderstr. & Lourteig [West Indian species, not in flora area] 

Arthrostylidium racemiflorum Steud. =Rhipidocladum racemiflorum 

Arthrostylidium steyermarkii McClure =Aulonemia deflexa 

Arthrostylidium subpectinatum Kuntze =Aulonemia subpectinata 

Arunda farcta Aubl. [West Indian species, not in flora area] =Arthrostylidium farctum 

Arundinaria deflexa N.E. Br. =Aulonemia deflexa 

Arundinaria schomburgkii Benn. =Arthrostylidium schomburgkii 

Arundinella confinis (Schult.) Hitchc. & Chase =Arundinella hispida 

Arundo altissima Benth. =Phragmites australis 

Arundo australis Cav. =Phragmites australis 

Arundo phragmites L. =Phragmites australis 

Arundo roraimensis N.E. Br. =Cortaderia roraimensis 

Aulonemia steyermarkii (McClure) McClure =Aulonemia deflexa 

Axonopus “perpusillus Swallen” [invalid name, error for Axonopus purpurellus Swallen] 

Axonopus affinis sensu Lindeman & Görts non Chase =Axonopus purpusii 

Axonopus affinis Chase =Axonopus fissifolius 

Axonopus anomalus Swallen =Axonopus purpusii 

Axonopus appendiculatus (J. Presl) Hitchc. & Chase =Axonopus excavatus 

Axonopus ater Chase =Axonopus fissifolius 

Axonopus attenuatus non (J. Presl) Hitchc., sensu Hitchc. =Axonopus flabelliformis 

Axonopus aturensis Luces =Axonopus anceps 

Axonopus aureus var. pilosus (Döll) Henrard =Axonopus chrysoblepharis 

Axonopus canescens var. psilachne (Döll) G.A. Black =Axonopus canescens 

Axonopus caracarahyensis G.A. Black & Fróes =Axonopus anceps 

Axonopus carinato-vaginatus (Mez) H. Scholz =Axonopus aureus 

Axonopus carinato-vaginatus var. sprucei (G.A. Black) H. Scholz =Axonopus canescens 

Axonopus caulescens sensu Swallen non (Mez) Henrard =Axonopus ramosus 

Axonopus chrysites (Steud.) Kuhlm. =Axonopus aureus 

Axonopus chrysodactylus (Trin.) Kuhlm. =Axonopus canescens

160 


Axonopus compressus var. affinis (Chase) M.R. Hend. =Axonopus fissifolius 

Axonopus compressus var. australis G.A. Black =Axonopus compressus 

Axonopus compressus var. macropodius (Steud.) G.A. Black =Axonopus compressus 

Axonopus erectus Swallen =Axonopus anceps 

Axonopus exasperatus (Nees ex Steud.) G.A. Black =Axonopus aureus 

Axonopus extenuatus (Nees) Kuhlm. =Axonopus capillaris 

Axonopus fissifolius sensu Amshoff & Henrard non (Raddi) Kuhlm. =Axonopus purpusii 

Axonopus fissifolius var. coronatus G.A. Black =Axonopus fissifolius 

Axonopus flabelliformis var. camporum G.A. Black =Axonopus flabelliformis 

Axonopus flabelliformis var. decipiens G.A. Black =Axonopus flabelliformis 

Axonopus flexilis (Mez) Henrard =Axonopus purpusii 

Axonopus fockei (Mez) Henrard =Axonopus surinamensis 

Axonopus fockei sensu Amshoff & Henrard non (Mez) Henrard, =Axonopus equitans 

Axonopus gentilis Henrard =Axonopus eminens 

Axonopus hirsutus G.A. Black =Axonopus fissifolius 

Axonopus hitchcockii G.A. Black =Axonopus longispicus 

Axonopus immersus (Nees) Kuhlm. =Axonopus chrysoblepharis 

Axonopus kaietukensis Swallen =Axonopus flabelliformis 

Axonopus laxus Luces =Axonopus capillaris 

Axonopus leptostachyus non (Flüggé) Hitchc. sensu Hitchc. =Axonopus longispicus 

Axonopus macrostachyus Hitchc. & Chase =Axonopus leptostachyus 

Axonopus maguirei G.A. Black [not in flora area] 

Axonopus minutus Luces =Axonopus aureus 

Axonopus multipes Swallen =Axonopus compressus 

Axonopus paranaensis Parodi =Axonopus leptostachyus 

Axonopus paucisetosus G.A. Black =Axonopus canescens 

Axonopus piccae Giraldo-CaZas =Axonopus iridifolius 

Axonopus pruinosus Henrard =Axonopus anceps 

Axonopus pulcher (Nees) Kuhlm. =Axonopus canescens 

Axonopus purpurellus Swallen =Axonopus flabelliformis 

Axonopus rivularis G.A. Black =Axonopus longispicus 

Axonopus scoparius (Flüggé) Hitchc. sensu Hitchc. =Axonopus pubivaginatus 

Axonopus sprucei G.A. Black =Axonopus canescens 

Axonopus sprucei var. glabratus (Döll) G.A. Black =Axonopus canescens 

Axonopus stragulus Chase =Axonopus fissifolius 

Axonopus stragulus sensu Amshoff & Henrard non Chase =Axonopus ramosus 

Axonopus tamayonis Luces =Axonopus flabelliformis 

Bambusa guadua Bonpl. =Guadua angustifolia 

Bambusa latifolia Bonpl. =Guadua latifolia 

Bambusa macrostachya (Rupr.) McClure =Guadua macrostachya 

Bambusa surinamensis Rupr. [cultivated] =Bambusa vulgaris 

Bambusa venezuelae (Munro) McClure =Guadua venezuelae 

Bambusa vulgaris Schrad. ex J.C. Wendl. [cultivated] 

Bothriochloa ischaemum sensu Hitchc. non (L.) Keng =Bothriochloa bladhii 

Bothriochloa pertusa sensu Hitchc. non (L.) Willd. =Bothriochloa bladhii 

Brachiaria purpurascens (Raddi) Henrard =Brachiaria mollis 

Bromus spicatus Nees =Tripogon spicatus 

Bulbulus nervatus Swallen =Rehia nervata 

Cabrera chrysoblepharis Lag. =Axonopus chrysoblepharis 

Capriola dactylon (L.) Kuntze =Cynodon dactylon 

Caryochloa bahiensis Steud. =Luziola bahiensis 

Cenchrus ciliaris L. =Pennisetum ciliare 

Cenchrus dactylolepis Steud. =Cenchrus brownii 

Cenchrus granularis L. =Hackelochloa granularis 

Cenchrus inflexus Poir. =Echinolaena inflexa 

Cenchrus marginalis Rudge =Echinolaena inflexa 

Cenchrus parviflorus Poir. =Setaria parviflora 

Cenchrus setosus Sw. =Pennisetum polystachion 

Cenchrus viridis Spreng. =Cenchrus brownii 

Chaetaria adscensionis (L.) P. Beauv. =Aristida adscensionis 

Chaetaria bromoides (Kunth) Roem. & Schult. =Aristida adscensionis 

Chaetaria capillacea (Lam.) P. Beauv. =Aristida capillacea 

Chaetaria coarctata (Kunth) Roem. & Schult. =Aristida adscensionis 

Chaetaria gibbosa Nees =Aristida gibbosa


Chaetaria humilis (Kunth) Roem. & Schult. =Aristida adscensionis 

Chaetaria recurvata (Kunth) Roem. & Schult. =Aristida recurvata 

Chaetaria setifolia (Kunth) Nees =Aristida setifolia 

Chaetaria torta Nees =Aristida torta 

Chaetochloa geniculata (Lam.) Millsp. & Chase =Setaria parviflora 

Chaetochloa poiretiana (Schult.) Hitchc. =Setaria poiretiana 

Chaetochloa tenax (Rich.) Hitchc. =Setaria tenax 

Chloris barbata (L.) Nash =Chloris elata 

Chloris dandyana C.D. Adams [nom. illeg.] =Chloris elata 

Chloris foliosa Willd. =Gymnopogon foliosus 

Chloris inflata Link =Chloris barbata 

Chloris mollis (Nees) Swallen =Enteropogon mollis 

Chloris polydactyla (L.) Sw. [nom. illeg.] =Chloris elata 

Chusquea venezuelae Steud. =Arthrostylidium venezuelae 

Coelorachis loricata (Trin.) Nash =Rhytachne rottboellioides 

Cryptochloa concinna sensu Swallen non (Hook. f.) Swallen =Piresia goeldii 

Cymbopogon bracteatus (Willd.) Hitchc. =Hyparrhenia bracteata 

Cynosurus aegyptius L. =Dactyloctenium aegyptium 

Cynosurus domingensis Jacq. =Leptochloa virgata 

Cynosurus indicus L. =Eleusine indica 

Cynosurus virgatus L. =Leptochloa virgata 

Diandrochloa diplachnoides (Steud.) A.N. Henry =Eragrostis japonica 

Diandrochloa glomerata (Walter) Burkart =Eragrostis japonica 

Diandrochloa japonica (Thunb.) A.N. Henry =Eragrostis japonica 

Diectomis fastigiata (Sw.) P. Beauv. =Andropogon fastigiatus 

Digitaria adscendens (Kunth) Henrard =Digitaria ciliaris 

Digitaria argillacea sensu Swallen non (Hitchc. & Chase) Fernald =Digitaria fragilis 

Digitaria aurea (P. Beauv.) Spreng. =Axonopus aureus 

Digitaria glabriculmis Swallen =Digitaria venezuelae 

Digitaria insularis Mez ex Ekman =Digitaria insularis 

Digitaria malacophylla (Hitchc.) Henrard =Digitaria lanuginosa 

Digitaria microbachne (J. Presl) Henrard =Digitaria setigera 

Digitaria platicaulis (Poir.) Desv. =Axonopus compressus 

Digitaria sacchariflora (Raddi) Henrard =Digitaria insularis 

Digitaria sanguinalis auct. non (L.) Scop. 1771: sensu Luces =Digitaria ciliaris 

Digitaria uniflora Salzm. ex Steud. =Axonopus compressus 

Elionurus adustus (Trin.) Ekman =Elionurus muticus 

Elionurus planifolius Renvoize =Elionurus muticus 

Eragrostis acuminata Döll =Eragrostis maypurensis 

Eragrostis acutiflora var. humilior J. Presl =Eragrostis acutiflora 

Eragrostis amabilis var. plumosa (Retz.) E.G. Camus & A. Camus =Eragrostis amabilis 

Eragrostis amoena J. Presl =Eragrostis maypurensis 

Eragrostis aturensis (Kunth) Trin. ex Steud. =Eragrostis japonica 

Eragrostis bahiensis sensu Lemée non Schrad. ex Schult. =Eragrostis scaligera 

Eragrostis barbata Trin. =Steirachne barbata 

Eragrostis capillaris sensu Lemée non (L.) Nees =Eragrostis pilosa 

Eragrostis chariis (Schult.) Hitchc. =Eragrostis atrovirens 

Eragrostis ciliaris var. patens Chapm. ex Beal =Eragrostis amabilis 

Eragrostis conferta (Elliott) Trin. =Eragrostis japonica 

Eragrostis diplachnoides Steud. =Eragrostis japonica 

Eragrostis elegantula (Kunth) Nees ex Steud. =Eragrostis gangetica 

Eragrostis filiformis Link =Eragrostis pilosa 

Eragrostis flamignii De Wild. =Eragrostis gangetica 

Eragrostis floridana Hitchc. =Eragrostis polytricha 

Eragrostis fragilis Swallen =Eragrostis polytricha 

Eragrostis glomerata (Walter) L.H. Dewey =Eragrostis japonica 

Eragrostis hapalantha Trin. =Eragrostis japonica 

Eragrostis interrupta var. diplachnoides (Steud.) Stapf =Eragrostis japonica 

Eragrostis interrupta var. laxiflora Döll =Eragrostis japonica 

Eragrostis interrupta var. parviflora Döll =Eragrostis japonica 

Eragrostis lindeniana Steud. =Eragrostis maypurensis 

Eragrostis linkii (Kunth) Steud. =Eragrostis pilosa 

Eragrostis maypurensis var. densiuscula Pilg. =Eragrostis maypurensis 

Eragrostis maypurensis var. meratiana (Kunth) Pilg. =Eragrostis maypurensis

162 


Eragrostis meratiana (Kunth) Steud. =Eragrostis maypurensis 

Eragrostis mexicana auct., non (Lag.) Link =Eragrostis pilosa 

Eragrostis namaquensis var. diplachnoides (Steud.) Clayton =Eragrostis japonica 

Eragrostis pallida Vasey =Eragrostis japonica 

Eragrostis panamensis J. Presl =Eragrostis maypurensis 

Eragrostis plumosa (Retz.) Link =Eragrostis amabilis 

Eragrostis poaeoides sensu Pulle non P. Beauv. =Eragrostis tephrosanthos 

Eragrostis polytricha var. glabrior Döll =Eragrostis polytricha 

Eragrostis polytricha var. hirsutior Döll =Eragrostis polytricha 

Eragrostis purpusii Jedwabn. =Eragrostis polytricha 

Eragrostis reptans (Michx.) Nees =Eragrostis hypnoides 

Eragrostis reptans var. contracta Döll =Eragrostis hypnoides 

Eragrostis reptans var. laxior Döll =Eragrostis hypnoides 

Eragrostis reptans var. pygmaea Döll =Eragrostis hypnoides 

Eragrostis rufescens var. laurentiana Benoist =Eragrostis scaligera 

Eragrostis stenoclada J. Presl =Eragrostis gangetica 

Eragrostis tenella (L.) P. Beauv. ex Roem. & Schult. =Eragrostis amabilis 

Eragrostis tenella var. japonica (Thunb.) Roem. & Schult. =Eragrostis japonica 

Eragrostis tenella var. plumosa (Retz.) Stapf =Eragrostis amabilis 

Eragrostis tenella var. viscosa (Retz.) Stapf =Eragrostis viscosa 

Eragrostis trichocolea Hack. & Arechav. =Eragrostis polytricha 

Eragrostis trichocolea var. floridana (Hitchc.) Witherspoon =Eragrostis polytricha 

Eragrostis vahlii (Roem. & Schult.) Nees =Eragrostis maypurensis 

Eragrostis vahlii var. polyantha Döll =Eragrostis maypurensis 

Eragrostis viscosa var. pilosissima (Hochst. ex A. Rich.) Hochst. =Eragrostis viscosa 

Eragrostis warmingii Hack. =Eragrostis viscosa 

Erianthus asper Nees =Saccharum asperum 

Erianthus saccharoides var. ß trinii Hack. =Saccharum trinii 

Erianthus trinii (Hack.) Hack. =Saccharum trinii 

Eriochloa kunthii G. Mey. =Eriochloa punctata 

Eriochloa ramosa (Retz.) Kuntze =Eriochloa procera 

Eriochloa subglabra (Nash) Hitchc. =Eriochloa polystachya 

Erosion hypnoides (Lam.) Lunell =Eragrostis hypnoides 

Euchlaena luxurians Durieu & Asch. =Zea luxurians 

Festuca filiformis Lam. =Leptochloa filiformis 

Guadua dioica Steud. =Guadua macrostachya 

Guadua tagoara sensu Pulle non (Nees) Kunth =Guadua latifolia 

Gymnopogon mollis Nees =Enteropogon mollis 

Gynerium saccharoides Humb. =Gynerium sagittatum 

Helopus cognatus Steud. =Eriochloa punctata 

Helopus kunthii (G. Mey.) Trin. ex Steud. =Eriochloa punctata 

Heteropogon secundus J. Presl =Trachypogon spicatus 

Heteropogon stipoides J. Presl =Trachypogon spicatus 

Heteropogon truncatus Nees =Trachypogon spicatus 

Holcus bicolor L. =Sorghum bicolor 

Holcus sorghum L. =Sorghum bicolor 

Holcus sorghum var. sudanensis (Piper) Hitchc. =Sorghum halepense 

Holcus striatus L. =Sacciolepis striata 

Homalocenchrus hexandrus (Sw.) Kuntze =Leersia hexandra 

Hymenachne auriculata (Spreng.) Chase =Hymenachne donacifolia 

Hypogynium virgatum (Desv.) Dandy =Andropogon virgatus 

Ichnanthus acuminatus Swallen =Ichnanthus calvescens 

Ichnanthus amplus Swallen =Ichnanthus calvescens 

Ichnanthus angustifolius Swallen =Ichnanthus tenuis 

Ichnanthus angustus Swallen =Ichnanthus ephemeroblepharis 

Ichnanthus areolatus K.E. Rogers =Ichnanthus pallens 

Ichnanthus axillaris (Nees) Hitchc. & Chase =Ichnanthus pallens 

Ichnanthus candicans sensu Lemée non (Nees) Döll =Ichnanthus tenuis 

Ichnanthus candicans var. virescens Döll =Ichnanthus tenuis 

Ichnanthus chaseae Swallen =Ichnanthus calvescens 

Ichnanthus confertus K.E. Rogers =Ichnanthus pallens 

Ichnanthus duidensis Swallen =Ichnanthus lancifolius var. weberbaueri 

Ichnanthus hispidus Swallen =Ichnanthus calvescens 

Ichnanthus hitchcockii K.E. Rogers =Ichnanthus pallens


Ichnanthus ichnodes (Griseb.) Hitchc. & Chase =Ichnanthus calvescens 

Ichnanthus longifolius Swallen =Ichnanthus ephemeroblepharis 

Ichnanthus neblinaensis Swallen =Ichnanthus ephemeroblepharis 

Ichnanthus nubilis Chase =Ichnanthus nemorosus 

Ichnanthus pallidus Swallen =Ichnanthus calvescens 

Ichnanthus petiolatus var. lanceolatus Döll =Ichnanthus nemoralis 

Ichnanthus polythyrsus (Nees ex Steud.) K.E. Rogers =Ichnanthus tenuis 

Ichnanthus riedelii sensu Hitchc. non (Trin.) Döll =Ichnanthus nemoralis 

Ichnanthus sabulosus K.E. Rogers =Ichnanthus tenuis 

Ichnanthus serratus Swallen =Ichnanthus ephemeroblepharis 

Ichnanthus sucrensis K.E. Rogers =Ichnanthus tenuis 

Ichnanthus tamayonis Chase =Ichnanthus tenuis 

Ichnanthus tectus Swallen =Ichnanthus ephemeroblepharis 

Ichnanthus tenuifolius K.E. Rogers =Ichnanthus nemorosus 

Ichnanthus venezuelanus Mez =Ichnanthus tenuis 

Ichnanthus vestitus Swallen =Ichnanthus calvescens 

Ichnanthus weberbaueri Mez =Ichnanthus lancifolius var. weberbaueri 

Imperata arundinacea var. americana L. Andersson =Imperata brasiliensis 

Imperata caudata (G. Mey.) Trin. =Imperata contracta 

Imperata flexuosa Swallen =Imperata contracta 

Isachne arundinacea auct. non (Sw.) Griseb. 1864: sensu Luces =Isachne rigens 

Isachne ligulata Swallen =Isachne rigens 

Ischaemopogon latifolius (Spreng.) Griseb. =Ischaemum latifolium 

Ischaemum ciliare Retz. =Ischaemum indicum 

Ischaemum guianense var. schomburgkii Hack. =Ischaemum guianense 

Ischaemum latifolium subsp. hirtivaginum Hitchc. =Ischaemum latifolium 

Ischaemum latifolium var. minus E. Fourn. =Ischaemum latifolium 

Ischaemum rugosum var. segetum (Trin.) Hack. =Ischaemum rugosum 

Ischaemum segetum Trin. =Ischaemum rugosum 

Lasiacis acuminata Swallen =Lasiacis sorghoidea 

Lasiacis compacta (Sw. ex Wikstr.) Hitchc. =Lasiacis ruscifolia 

Lasiacis divaricata sensu Amshoff & Henrard non (L.) Hitchc. =Lasiacis ligulata 

Lasiacis glabra Swallen =Lasiacis ruscifolia 

Lasiacis guaraniticum (Speg.) Parodi =Lasiacis sorghoidea 

Lasiacis liebmanniana (E. Fourn.) Hitchc. =Lasiacis ruscifolia 

Lasiacis patentiflora Hitchc. & Chase =Lasiacis sorghoidea 

Lasiacis sorghoidea var. patentiflora (Hitchc. & Chase) Davidse =Lasiacis sorghoidea 

Lasiacis swartziana (Hitchc.) Hitchc. =Lasiacis sorghoidea 

Leersia distichophylla Balansa & Poitr. =Leersia ligularis 

Leersia grandiflora (Döll) Prodoehl =Leersia ligularis 

Leersia ligularis var. glabriflora Pyrah =Leersia ligularis 

Leersia ligularis var. grandiflora (Döll) Pyrah =Leersia ligularis 

Leptochloa domingensis (Jacq.) Trin. =Leptochloa virgata 

Leptochloa mutica Steud. =Leptochloa virgata 

Leptochloa virgata var. mutica (Steud.) Döll =Leptochloa virgata 

Leptostachys domingensis G. Mey. =Leptochloa virgata 

Leptostachys filiformis G. Mey. =Leptochloa filiformis 

Leptostachys virgata G. Mey. =Leptochloa virgata 

Luziola pittieri Luces =Luziola doelliana 

Luziola spruceana auct. non Benth. ex Döll 1871: sensu Hitchc. =Luziola subintegra 

Lycurus muticus Spreng. =Elionurus muticus 

Manisuris aurita (Steud.) Kuntze =Coelorachis aurita 

Manisuris granularis (L.) Naezén =Hackelochloa granularis 

Manisuris guianensis Hitchc. =Rhytachne guianensis 

Manisuris loricata (Trin.) Kuntze =Rhytachne rottboellioides 

Megastachya acutiflora (Kunth) Roem. & Schult. =Eragrostis acutiflora 

Megastachya breviflora E. Fourn. =Eragrostis hypnoides 

Megastachya corymbifera E. Fourn. =Eragrostis hypnoides 

Megastachya gouinii E. Fourn. =Eragrostis hypnoides 

Megastachya hypnoides (Lam.) P. Beauv. =Eragrostis hypnoides 

Megastachya maypurensis (Kunth) Roem. & Schult. =Eragrostis maypurensis 

Melinis repens (Willd.) Zizka =Rhynchelytrum repens 

Meoschium rugosum (Salisb.) Nees =Ischaemum rugosum 

Mesosetum cardonum Luces =Mesosetum chaseae

164 


Mesosetum elongatum Mez =Mesosetum loliiforme 

Mesosetum tenuifolium Swallen =Mesosetum loliiforme 

Milium compressum Sw. =Axonopus compressus 

Milium punctatum L. =Eriochloa punctata 

Mnesithea aurita (Steud.) de Koning & Sosef =Coelorachis aurita 

Mnesithea subgibbosa (Winkl. ex Hack.) de Koning & Sosef =Rhytachne subgibbosa 

Myriocladus affinis Swallen =Myriocladus longiramosus 

Myriocladus confertus Swallen =Myriocladus distantiflorus 

Myriocladus gracilis Swallen =Myriocladus steyermarkii 

Myriocladus maguirei Swallen =Myriocladus virgatus 

Myriocladus paraquensis Swallen =Myriocladus grandifolius 

Myriocladus paruensis Swallen =Myriocladus longiramosus 

Myriocladus purpureus Swallen =Myriocladus distantiflorus 

Myriocladus variabilis Swallen =Myriocladus distantiflorus 

Myriocladus wurdackii Swallen =Myriocladus distantiflorus 

Neeragrostis hypnoides (Lam.) Bush =Eragrostis hypnoides 

Neurolepis densiflora Swallen =Neurolepis glomerata 

Neurolepis nigra Swallen =Neurolepis glomerata 

Olyra cordifolia Kunth =Olyra latifolia 

Olyra dimidiata Hochst. ex Steud. =Olyra caudata 

Olyra fasciculata auct. non Trin. 1834 =Olyra standleyi 

Olyra floribunda var. microphylla Döll =Raddia guianensis 

Olyra kegelii Mez =Olyra longifolia 

Olyra lateralis (J. Presl ex Nees) Chase =Parodiolyra lateralis 

Olyra lateralis sensu Amshoff & Henrard non (J. Presl ex Nees) Chase =Parodiolyra luetzelburgii 

Olyra latifolia var. pubescens (Raddi) Döll =Olyra latifolia 

Olyra latifolia var. vestita Henrard =Olyra latifolia 

Olyra luetzelburgii Pilg. =Parodiolyra luetzelburgii 

Olyra micrantha Kunth =Parodiolyra micrantha 

Olyra nana Döll [nom. superfl.] =Raddiella esenbeckii 

Olyra ovata Desv. =Ichnanthus panicoides 

Olyra pauciflora Sw. =Lithachne pauciflora 

Olyra pubescens Raddi =Olyra latifolia 

Olyra sarmentosa Döll =Parodiolyra lateralis 

Olyra scabra Nees =Olyra latifolia 

Olyra speciosa Mez =Olyra caudata 

Olyra surinamensis Hochst. ex Steud. =Olyra latifolia 

Olyra surinamensis auct. non Hochst. ex Steud. =Olyra longifolia 

Olyra sympodica Döll =Piresia sympodica 

Oplismenus crus-pavonis Kunth =Echinochloa crus-pavonis 

Oplismenus polystachyus Kunth =Echinochloa polystachya 

Oplismenus pseudo-undulatifolius (Roem. & Schult.) Kunth =Panicum stoloniferum 

Oplismenus tenuis J. Presl =Ichnanthus tenuis 

Oplismenus velutinus (G. Mey.) Schult. =Oplismenus hirtellus 

Orthoclada rariflora (Lam.) P. Beauv. =Orthoclada laxa 

Orthoclada rariflora var. lanceolata Döll =Orthoclada laxa 

Orthoclada rariflora var. sesquiflora Döll =Orthoclada laxa 

Oryza alta Swallen =Oryza latifolia 

Oryza glumaepatula Hochst. ex Steud. =Oryza rufipogon 

Oryza monandra var. grandiflora Döll =Leersia ligularis 

Oryza perennis Moench [nom. confus.] =Oryza rufipogon 

Panicum aequivaginatum Swallen =Dichanthelium aequivaginatum 

Panicum agglutinans Kunth =Lasiacis sorghoidea 

Panicum albociliatum Swallen =Dichanthelium pycnoclados 

Panicum altissimum G. Mey. =Panicum mertensii 

Panicum amplexicaule Rudge =Hymenachne amplexicaulis 

Panicum amplifolium Steud. =Setaria vulpiseta 

Panicum angulosum Swallen =Panicum rivale 

Panicum angustissimum Hochst. ex Steud. =Sacciolepis angustissima 

Panicum appressifolium Swallen =Dichanthelium aequivaginatum 

Panicum appressum (Lam.) Döll =Paspalidium geminatum 

Panicum arrectum Hack. ex T. Durand & Schinz =Brachiaria arrecta 

Panicum asperifolium (Desv.) Hitchc. =Streptostachys asperifolia 

Panicum aturense Kunth =Homolepis aturensis


Panicum auyanense Swallen =Panicum eligulatum 

Panicum axillare Nees =Ichnanthus pallens 

Panicum axipilium Steud. =Panicum hirtum 

Panicum blackii Swallen =Panicum micranthum 

Panicum breviscrobs (Döll) Pilg. =Ichnanthus breviscrobs 

Panicum burmannii Retz. =Oplismenus burmannii 

Panicum calvescens Nees =Ichnanthus calvescens 

Panicum caricoides sensu Lemée non Nees =Panicum stenodes 

Panicum caricoides var. pilosum Döll =Panicum caricoides 

Panicum caroniense Luces =Steinchisma laxa 

Panicum caudatum Lam. =Setaria setosa 

Panicum cayennense var. curtatum Döll =Panicum cayennense 

Panicum cayennense var. divaricatum Döll =Panicum rudgei 

Panicum cayennense var. patulum Döll =Panicum cayennense 

Panicum chlorostachyum Döll =Mesosetum chlorostachyum 

Panicum chloroticum Nees =Panicum dichotomiflorum 

Panicum chrysites Steud. =Axonopus aureus 

Panicum chrysoblephare (Lag.) Steud. =Axonopus chrysoblepharis 

Panicum chrysodactylon Trin. =Axonopus canescens 

Panicum churunense Swallen =Panicum chnoodes 

Panicum ciliare Retz. =Digitaria ciliaris 

Panicum coenosum Döll =Panicum pilosum 

Panicum colonum L. =Echinochloa colona 

Panicum commelinaefolium Rudge =Panicum nervosum 

Panicum compactum Sw. ex Wikstr. =Lasiacis ruscifolia 

Panicum compositum auct., non L. =Oplismenus hirtellus 

Panicum cordovense sensu Amshoff & Henrard non E. Fourn. =Dichanthelium pycnoclados 

Panicum cowanii Swallen =Panicum chnoodes 

Panicum crus-galli var. aristatum G. Mey. =Echinochloa crus-pavonis 

Panicum curvifolium Swallen =Panicum chnoodes 

Panicum cyanescens var. lamarckianum Döll =Panicum cyanescens 

Panicum dactylon L. =Cynodon dactylon 

Panicum davidsei Zuloaga & Morrone =Dichanthelium davidsei 

Panicum decipiens Nees ex Trin. =Steinchisma decipiens 

Panicum densifolium Swallen =Panicum micranthum 

Panicum distichum var. lancifolium Griseb. =Panicum pilosum 

Panicum distichum var. lancifolium Griseb. ex Hitchc. =Panicum pilosum 

Panicum distichum var. luxurians G. Mey. =Panicum pilosum 

Panicum divaricatum var. lanatum Schltdl. & Cham. =Lasiacis sorghoidea 

Panicum divaricatum var. latifolium Schltdl. & Cham. =Lasiacis sorghoidea 

Panicum donacifolium Raddi =Hymenachne donacifolia 

Panicum echinolaena Nees =Echinolaena inflexa 

Panicum elatior Kunth =Panicum mertensii 

Panicum errabundum Hitchc. =Panicum schwackeanum 

Panicum esenbeckii Nees ex Steud. =Raddiella esenbeckii 

Panicum exasperatum Nees ex Steud. =Axonopus aureus 

Panicum excavatum Henrard =Acroceras excavatum 

Panicum fasciculatum Sw. =Brachiaria fasciculata 

Panicum fistulosum Hochst. ex Steud. =Panicum elephantipes 

Panicum frondescens G. Mey. =Panicum stoloniferum 

Panicum funckianum Steud. =Panicum olyroides 

Panicum fusciflorum Steud. =Leptocoryphium lanatum 

Panicum fuscum Sw. =Brachiaria fasciculata 

Panicum geminatum Forssk. =Paspalidium geminatum 

Panicum geniculatum Willd. =Setaria parviflora 

Panicum glutinosum Lam. =Lasiacis sorghoidea 

Panicum glutinosum Sw. =Homolepis glutinosa 

Panicum gracilissimum Swallen =Panicum polycomum 

Panicum graniticum Swallen =Panicum pandum 

Panicum guaraniticum Speg. =Lasiacis sorghoidea 

Panicum guianense Hitchc. =Panicum hylaeicum 

Panicum hebotes Trin. =Dichanthelium hebotes 

Panicum hirtellum L. =Oplismenus hirtellus 

Panicum hoffmannseggii Roem. & Schult. =Ichnanthus hoffmannseggii

166 


Panicum horizontale (Willd.) G. Mey. =Digitaria horizontalis 

Panicum humidicolum Rendle =Brachiaria humidicola 

Panicum imberbe Poir. =Setaria parviflora 

Panicum immersum (Nees) Trin. =Axonopus chrysoblepharis 

Panicum insulare (L.) G. Mey. =Digitaria insularis 

Panicum inversum Swallen =Panicum chnoodes 

Panicum isocalycium G. Mey. =Homolepis isocalycia 

Panicum kaietukense Tutin =Panicum parvifolium 

Panicum kappleri Steud. =Panicum micranthum 

Panicum kavanayense Swallen =Panicum chnoodes 

Panicum lanatum Sw. =Lasiacis sorghoidea 

Panicum lanatum var. sorghoideum (Desv. ex Ham.) Griseb. =Lasiacis sorghoidea 

Panicum laterale C. Presl. ex Nees =Parodiolyra lateralis 

Panicum latifolium Ham. =Lasiacis sloanei 

Panicum laxum Sw. =Steinchisma laxa 

Panicum leprieurii Steud. =Panicum stoloniferum 

Panicum leptachyrium Döll =Paratheria prostrata 

Panicum liebmannianum E. Fourn. =Lasiacis ruscifolia 

Panicum loliaceum auct., non Lam. =Oplismenus hirtellus 

Panicum loliiforme Hochst. ex Steud. =Mesosetum loliiforme 

Panicum longepaniculatum Benoist =Setaria poiretiana 

Panicum luticola Hitchc. =Steinchisma laxa 

Panicum macrostachyum (Kunth) Döll =Setaria vulpiseta 

Panicum magnum Hitchc. =Ichnanthus breviscrobs 

Panicum maguirei Swallen =Panicum chnoodes 

Panicum manacalense Swallen =Panicum scabridum 

Panicum manacalensis Swallen =Panicum scabridum 

Panicum mauryi Swallen =Panicum micranthum 

Panicum megiston Schult. =Panicum mertensii 

Panicum micranthum var. hirtum Maury =Panicum micranthum 

Panicum milleflorum Hitchc. & Chase =Panicum pilosum 

Panicum mirandum Luces =Dichanthelium hebotes 

Panicum molle Sw. =Brachiaria mollis 

Panicum monostachyum Kunth =Panicum pilosum 

Panicum multiflorum sensu Pulle non Poir. =Panicum grande 

Panicum musei Steud. =Echinochloa colona 

Panicum muticum Forssk. =Brachiaria mutica 

Panicum myosurus Rich. =Sacciolepis myuros 

Panicum myrianthum Mez =Panicum grande 

Panicum myuros Lam. =Sacciolepis myuros 

Panicum nemorale Schrad. ex Schult. =Ichnanthus nemoralis 

Panicum nemorosum Sw. =Ichnanthus nemorosus 

Panicum numidianum sensu Pulle non Lam. =Brachiaria mutica 

Panicum obovatum Döll =Panicum polycomum 

Panicum pallens Sw. =Ichnanthus pallens 

Panicum pedunculare Willd. ex Steud. =Panicum cayennense 

Panicum perpusillum Benoist =Panicum polycomum 

Panicum petrosum Trin. =Thrasya petrosa 

Panicum pilcomayense Hack. =Panicum bergii 

Panicum pilisparsum G. Mey. =Panicum pilosum 

Panicum pilosissimum Roth =Panicum rudgei 

Panicum pilosum var. lancifolium (Griseb.) R. Pohl =Panicum pilosum 

Panicum plantagineum Link =Brachiaria plantaginea 

Panicum platycaulon (Poir.) Kuntze =Axonopus compressus 

Panicum poiretianum Schult. =Setaria poiretiana 

Panicum politii Swallen =Panicum granuliferum 

Panicum polygonoides Lam. =Isachne polygonoides 

Panicum polystachion L. =Pennisetum polystachion 

Panicum praegnans Steud. =Lasiacis sorghoidea 

Panicum prieurii Mez =Panicum scabridum 

Panicum procerrimum Hack. =Lasiacis procerrima 

Panicum procurrens Nees ex Trin. =Ichnanthus procurrens 

Panicum proximum Steud. =Panicum mertensii 

Panicum pseudo-undulatifolium Roem. & Schult. =Panicum stoloniferum


Panicum purpurascens Raddi =Brachiaria mutica 

Panicum pycnoclados Tutin =Dichanthelium pycnoclados 

Panicum quetameense Mez =Panicum pyrularium 

Panicum racemiferum Steud. ? =Setaria poiretiana 

Panicum ramuliflorum Hochst. ex Steud. =Steinchisma laxa 

Panicum rariflorum Lam. =Orthoclada laxa 

Panicum recalvum sensu Pulle non (Nees) Kunth =Digitaria laxa 

Panicum reptans L. =Urochloa reptans 

Panicum retroflexum Steud. ? =Setaria sp. 

Panicum rigens Sw. =Isachne rigens 

Panicum rottboellioides Kunth =Mesosetum rottboellioides 

Panicum rugulosum var. subvelutinum Döll =Panicum millegrana 

Panicum ruscifolium Kunth =Lasiacis ruscifolia 

Panicum sanguinale sensu Pulle non L. =Digitaria ciliaris 

Panicum savannarum Soderstr. =Panicum cyanescens 

Panicum savannarum Schltdl. ex G.A. Black =Axonopus chrysoblepharis 

Panicum sciurotis auct., non Trin. =Dichanthelium sciurotoides 

Panicum sciurotoides Zuloaga & Morrone =Dichanthelium sciurotoides 

Panicum scoparium Rudge =Panicum rudgei 

Panicum sellowii var. subvelutinum (Döll) Henrard =Panicum millegrana 

Panicum setarioides Steud. =Echinochloa pyramidalis 

Panicum siccaneum Trin. =Panicum polycomum 

Panicum sloanei Griseb. =Lasiacis sloanei 

Panicum sorghoideum Desv. ex Ham. =Lasiacis sorghoidea 

Panicum spectabile Nees ex Trin. =Echinochloa polystachya 

Panicum spissifolium Swallen =Panicum granuliferum 

Panicum stenodoides F.T. Hubb. =Panicum caricoides 

Panicum stenophyllum Hack. =Steinchisma stenophylla 

Panicum subcordatum Swallen =Panicum granuliferum 

Panicum subfalcatum Döll =Paspalum subfalcatum 

Panicum subinclusum Swallen =Panicum granuliferum 

Panicum sulcatum Aubl. =Setaria palmifolia 

Panicum supernum Swallen =Panicum micranthum 

Panicum surinamense Hochst. ex Steud. =Axonopus surinamensis 

Panicum swartzianum Hitchc. =Lasiacis sorghoidea 

Panicum tamayonis Luces =Panicum polycomum 

Panicum tatei Swallen =Panicum chnoodes 

Panicum telmatum Swallen =Dichanthelium telmatum 

Panicum tenax Rich. =Setaria tenax 

Panicum tenuiculmum G. Mey. =Steinchisma laxa 

Panicum tenuissimum Mart. ex Schrank =Sporobolus tenuissimus 

Panicum tiricaense Swallen =Dichanthelium pycnoclados 

Panicum tiricaoides Swallen =Dichanthelium pycnoclados 

Panicum tropidoblephare Tutin =Panicum chnoodes 

Panicum vannum Swallen =Panicum chnoodes 

Panicum velutinum G. Mey. =Oplismenus hirtellus 

Panicum versicolor Döll =Otachyrium versicolor 

Panicum vigoratum Swallen =Panicum altum 

Panicum vilvoides Trin. =Sacciolepis vilvoides 

Panicum vinaceum Swallen =Panicum cervicatum 

Panicum vulpisetum Lam. =Setaria vulpiseta 

Panicum wurdackii Swallen =Panicum eligulatum 

Panicum zizanioides Kunth =Acroceras zizanioides 

Pariana angustifolia Spreng. =Steyermarkochloa angustifolia 

Pariana debilis Tutin =Pariana radiciflora 

Pariana gleasonii Hitchc. =Pariana radiciflora 

Pariana inaequalis Miq. =Pariana campestris 

Pariana longiflora Tutin =Pariana radiciflora 

Pariana lunata Nees =Pariana campestris 

Pariana maynensis Huber =Pariana radiciflora 

Pariana mollis Nees =Pariana campestris 

Pariana obtusa Swallen =Pariana radiciflora 

Pariana pallida Swallen =Pariana radiciflora 

Pariana pulcherrima Tutin =Pariana radiciflora

168 


Pariana trichosticha Tutin =Pariana radiciflora 

Pariana violascens Swallen =Pariana radiciflora 

Pariana vulgaris Tutin =Pariana radiciflora 

Pariana zingiberina Rich. ex Döll =Pariana radiciflora 

Paspalum abrahamii Chase =Paspalum maritimum 

Paspalum abstrusum Trin. =Paspalum hyalinum 

Paspalum anceps Mez =Axonopus anceps 

Paspalum appendiculatum J. Presl =Axonopus excavatus 

Paspalum arenicolum Müll. Hal. =Paspalum pulchellum 

Paspalum argyrocondylon Steud. =Paspalum serpentinum 

Paspalum aureolatum Swallen =Paspalum lanciflorum 

Paspalum aureum (P. Beauv.) Kunth =Axonopus aureus 

Paspalum axillare Swallen =Thrasya axillaris 

Paspalum bistipulatum Hochst. ex Steud. =Paspalum repens 

Paspalum caespitosum sensu Pulle non Flüggé =Paspalum maritimum 

Paspalum caespitosum Steud. =Paspalum melanospermum 

Paspalum canum Sohns =Paspalum ammodes 

Paspalum capillare Lam. =Axonopus capillaris 

Paspalum carinato-vaginatum Mez =Axonopus aureus 

Paspalum carinatum var. kappleri (Steud.) Döll =Paspalum carinatum 

Paspalum caulescens Mez =Axonopus caulescens 

Paspalum chrysites (Steud.) Döll =Axonopus aureus 

Paspalum chrysoblephare (Lag.) Döll =Axonopus chrysoblepharis 

Paspalum chrysodactylon (Trin.) Döll =Axonopus canescens 

Paspalum chrysodactylon var. glabratum Döll =Axonopus canescens 

Paspalum chrysodactylon var. psilachne Döll =Axonopus canescens 

Paspalum chrysodactylon var. villosum Döll =Axonopus canescens 

Paspalum ciliatum Lam. =Paspalum conjugatum 

Paspalum compressum (Sw.) Raspail =Axonopus compressus 

Paspalum compressum var. arenarium Bertoni =Axonopus fissifolius 

Paspalum conjugatum var. pubescens Döll =Paspalum conjugatum 

Paspalum conjugatum var. subcordatum Griseb. =Axonopus compressus 

Paspalum contractum Pilg. =Paspalum lanciflorum 

Paspalum densiflorum sensu Pulle non Döll =Paspalum maritimum 

Paspalum densum var. ciliatum Döll =Paspalum densum 

Paspalum depressum Steud. =Axonopus compressus 

Paspalum dispar var. marahuacense H. Rodr. =Paspalum petilum 

Paspalum echinotrichum Mez =Paspalum lanciflorum 

Paspalum elatum Rich. ex Döll =Paspalum arundinaceum 

Paspalum eminens Nees =Axonopus eminens 

Paspalum erectifolium Swallen =Paspalum aspidiotes 

Paspalum excavatum Nees ex Trin. =Axonopus excavatus 

Paspalum extenuatum Nees =Axonopus capillaris 

Paspalum familiare Steud. =Paspalum coryphaeum 

Paspalum filostachyum A. Rich. ex Steud. =Axonopus compressus 

Paspalum fissifolium Raddi =Axonopus fissifolius 

Paspalum flexile Mez =Axonopus purpusii 

Paspalum fockei Mez =Axonopus surinamensis 

Paspalum foveolatum Steud. =Paspalum melanospermum 

Paspalum froesianum Swallen =Paspalum loefgrenii 

Paspalum furcatum sensu Pulle non Flüggé =Axonopus surinamensis 

Paspalum furcatum var. fissum Döll =Axonopus compressus 

Paspalum furcatum var. parviflorum Döll =Axonopus compressus 

Paspalum fuscescens J. Presl =Digitaria fuscescens 

Paspalum geniculatum Steud. =Paspalum orbiculatum 

Paspalum gnaphalioideum Müll. Hal. =Axonopus chrysostachyus 

Paspalum gossypinum Mez =Paspalum hyalinum 

Paspalum gracile Rudge =Paspalum repens 

Paspalum guadaloupense Steud. =Axonopus compressus 

Paspalum hirtigluma Steud. =Paspalum maritimum 

Paspalum humile Steud. =Paspalum melanospermum 

Paspalum immersum Nees =Axonopus chrysoblepharis 

Paspalum immersum var. pilosum Döll =Axonopus chrysoblepharis 

Paspalum indutum Luces =Paspalum coryphaeum


Paspalum iridifolium Poepp. =Axonopus iridifolius 

Paspalum kappleri Hochst. ex Steud. =Paspalum carinatum 

Paspalum kegelii Müll. Hal. =Spheneria kegelii 

Paspalum lacustre sensu Judz., non Swallen =Paspalum morichalense 

Paspalum lanatum Kunth =Leptocoryphium lanatum 

Paspalum laticulmum Spreng. =Axonopus compressus 

Paspalum leptostachyum Flüggé =Axonopus leptostachyus 

Paspalum longebrachiatum Steud. =Paspalum arundinaceum 

Paspalum longispica Döll =Axonopus longispicus 

Paspalum longispicum Döll =Axonopus longispicus 

Paspalum longissimum Hochst. ex Steud. =Paspalum conjugatum 

Paspalum lucidulum Swallen =Paspalum altsonii 

Paspalum macropodium Steud. =Axonopus compressus 

Paspalum maculosum var. rotundiflorum Döll =Paspalum serpentinum 

Paspalum meyerianum Spreng. =Paspalum exaltatum 

Paspalum minutum Trin. =Axonopus capillaris 

Paspalum montanum Henrard =Paspalum oligostachyum 

Paspalum orbiculatum var. lanuginosum Henrard =Paspalum orbiculatum 

Paspalum orbiculatum var. potaroense Chase =Paspalum orbiculatum 

Paspalum pectinatum sensu Hitchc. non Nees ex Trin. =Paspalum lanciflorum 

Paspalum pedunculatum Poir. =Paspalum decumbens 

Paspalum piligerum Swallen =Paspalum lanciflorum 

Paspalum platycaulon Poir. =Axonopus compressus 

Paspalum platycaulon var. gracilius Döll =Axonopus purpusii 

Paspalum platycaulon var. parviflorum Döll =Axonopus purpusii 

Paspalum plicatulum var. microspermum Döll =Paspalum melanospermum 

Paspalum polychaetum Mez =Paspalum hyalinum 

Paspalum pulchrum Nees =Axonopus canescens 

Paspalum pulchrum var. angustifolium Döll =Axonopus aureus 

Paspalum purpusii Mez =Axonopus purpusii 

Paspalum pusillum Vent. ex Flüggé =Paspalum orbiculatum 

Paspalum raunkiaerii Mez =Axonopus compressus 

Paspalum scandens Tutin =Thrasya scandens 

Paspalum scoparium var. parviflorum Döll =Axonopus anceps 

Paspalum senescens Döll =Axonopus senescens 

Paspalum setifolium Döll =Spheneria kegelii 

Paspalum setiglume Chase =Paspalum aspidiotes 

Paspalum triglochinoides Mez =Axonopus triglochinoides 

Paspalum tristachyon Lam. =Axonopus compressus 

Paspalum vaginiflorum Steud. =Paspalum decumbens 

Paspalum wagenerianum Schltdl. =Paspalum stellatum 

Paspalum wullschlaegelii Mez =Paspalum maritimum 

Pennisetum dasistachyum Desv. =Pennisetum polystachion 

Pennisetum richardii Kunth =Pennisetum polystachion 

Pennisetum setosum (Sw.) Rich. =Pennisetum polystachion 

Pharus glaber Kunth =Pharus lappulaceus 

Pharus glaber sensu Lemée non Kunth =Pharus latifolius 

Pharus scaber var. pictus Döll =Pharus latifolius 

Phragmites communis Trin. =Phragmites australis 

Piptatherum confine Schult. =Arundinella hispida 

Poa acutiflora Kunth =Eragrostis acutiflora 

Poa amabilis L. =Eragrostis amabilis 

Poa atrovirens Desf. =Eragrostis atrovirens 

Poa aturensis Kunth =Eragrostis japonica 

Poa chariis Schult. =Eragrostis atrovirens 

Poa ciliaris L. =Eragrostis ciliaris 

Poa conferta Elliott =Eragrostis japonica 

Poa elegantula Kunth =Eragrostis gangetica 

Poa gangetica Roxb. =Eragrostis gangetica 

Poa glomerata Walter =Eragrostis japonica 

Poa hypnoides Lam. =Eragrostis hypnoides 

Poa japonica Thunb. =Eragrostis japonica 

Poa linkii Kunth =Eragrostis pilosa 

Poa maypurensis Kunth =Eragrostis maypurensis

170 


Poa meratiana Kunth =Eragrostis maypurensis 

Poa pectinacea Michx. =Eragrostis pectinacea 

Poa pilosa L. =Eragrostis pilosa 

Poa plumosa Retz. =Eragrostis amabilis 

Poa polytricha (Nees) Kunth =Eragrostis polytricha 

Poa racemosa Vahl =Eragrostis maypurensis 

Poa tenella L. =Eragrostis amabilis 

Poa unioloides Retz. =Eragrostis unioloides 

Poa vahlii Roem. & Schult. =Eragrostis maypurensis 

Poa viscosa Retz. =Eragrostis viscosa 

Poa viscosa var. pilosissima Hochst. ex A. Rich. =Eragrostis viscosa 

Polypogon spicatus Spreng. =Gymnopogon spicatus 

Raddia capillata (Trin.) Hitchc. =Cryptochloa capillata 

Raddia malmeana sensu Hitchc. non (Ekman) Hitchc. =Raddiella kaieteurana 

Raddia nana (Döll) Chase =Raddiella esenbeckii 

Raddia sympodica (Döll) Hitchc. =Piresia sympodica 

Raddiella maipuriensis Soderstr. =Raddiella potaroensis 

Raddiella malmeana sensu Hitchc. non (Ekman) Swallen =Raddiella kaieteurana 

Raddiella nana (Döll) Swallen =Raddiella esenbeckii 

Raddiella truncata Swallen =Parodiolyra lateralis 

Reimaria aberrans Döll =Reimarochloa aberrans 

Reimaria acuta Flüggé =Reimarochloa acuta 

Reimaria laxa Rchb. ex A. Spreng. =Digitaria laxa 

Reimarochloa brasiliensis (Spreng.) Hitchc. =Reimarochloa acuta 

Rhynchelytrum roseum (Nees) Stapf & C.E. Hubb. =Rhynchelytrum repens 

Roshevitzia glomerata (Walter) Tzvelev =Eragrostis japonica 

Roshevitzia japonica (Thunb.) Tzvelev =Eragrostis japonica 

Rottboellia aurita Steud. =Coelorachis aurita 

Rottboellia compressa L. f. =Hemarthria altissima 

Rottboellia exaltata L. f. =Rottboellia cochinchinensis 

Rottboellia loricata Trin. =Rhytachne rottboellioides 

Rottboellia loricata subsp. glaberrima Hack. =Rhytachne subgibbosa 

Rottboellia loricata subsp. subgibbosa Winkl. ex Hack. =Rhytachne subgibbosa 

Rottboellia sanguinea Retz. =Schizachyrium sanguineum 

Saccharum caudatum G. Mey. =Imperata contracta 

Saccharum cayennense (P. Beauv.) Benth. =Eriochrysis cayennensis 

Saccharum contractum Kunth =Imperata contracta 

Saccharum giganteum Trin. =Saccharum trinii 

Saccharum repens Willd. =Rhynchelytrum repens 

Saccharum sagittatum Aubl. =Gynerium sagittatum 

Sacciolepis pungens Swallen =Sacciolepis angustissima 

Schizachyrium hirtiflorum Nees =Schizachyrium sanguineum 

Schizachyrium microstachyum (Desv. ex Ham.) Roseng., B.R. Arrill. & Izag. =Schizachyrium condensatum 

Schizachyrium riedelii (Trin.) A. Camus =Schizachyrium sanguineum 

Schizachyrium riedelii var. multirameum (Hack.) Henrard =Schizachyrium sanguineum 

Schizachyrium semiberbe Nees =Schizachyrium sanguineum 

Setaria geniculata (Willd.) P. Beauv. [nom. illeg.] =Setaria parviflora 

Setaria gracilis Kunth =Setaria parviflora 

Setaria poiretiana (Schult.) Kunth = Setaria sulcata Raddi 

Setaria sulcata (Aubl.) A. Camus =Setaria palmifolia 

Sorghastrum parviflorum (Desv.) Hitchc. & Chase =Sorghastrum setosum 

Sorghum parviflorum Desv. =Sorghastrum setosum 

Sorghum vulgare Pers. =Sorghum bicolor 

Spartina brasiliensis Raddi =Spartina alterniflora 

Spheneria setifolia (Döll) Kuhlm. =Spheneria kegelii 

Sporobolus aeneus sensu Hitchc. non (Trin.) Kunth =Sporobolus cubensis 

Sporobolus effusus Desv. =Sporobolus tenuissimus 

Sporobolus indicus auct., non (L.) R. Br. =Sporobolus jacquemontii 

Sporobolus indicus var. pyramidalis (P. Beauv.) Veldkamp =Sporobolus jacquemontii 

Sporobolus littoralis (Lam.) Kunth =Sporobolus virginicus 

Sporobolus minutiflorus (Trin.) Link =Sporobolus tenuissimus 

Sporobolus muralis (Raddi) Hitchc. & Chase =Sporobolus tenuissimus 

Sporobolus poiretii auct., non (Roem. & Schult.) Hitchc. =Sporobolus jacquemontii 

Sporobolus pyramidalis P. Beauv. =Sporobolus jacquemontii 

Sporobolus pyramidalis var. jacquemontii (Kunth) Jovet & GuédPs =Sporobolus jacquemontii


Sporobolus villosus (A. Spreng.) Kunth =Sporobolus ciliatus 

Steirachne diandra sensu Swallen non Ekman =Steirachne barbata 

Stenotaphrum glabrum var. americanum (Schrank) Döll =Stenotaphrum secundatum 

Steyermarkochloa unifolia Davidse & R.P. Ellis =Steyermarkochloa angustifolia 

Stipa spicata L. f. =Trachypogon spicatus 

Strephium guianense Brongn. =Raddia guianensis 

Streptachne scabra Kunth =Aristida ternipes 

Streptachne tenuis Kunth =Aristida ternipes 

Streptogyna crinita auct., non P. Beauv. =Streptogyna americana 

Streptostachys hirsuta P. Beauv. =Streptostachys asperifolia 

Streptostachys vaginaeflorum Steud. =Streptostachys asperifolia 

Syntherisma chinensis (Nees) Hitchc. =Digitaria violascens 

Syntherisma cuyabensis (Trin.) Hitchc. =Digitaria lanuginosa 

Syntherisma digitata sensu Hitchc. non (Sw.) Hitchc. =Digitaria horizontalis 

Syntherisma longiflora (Retz.) Skeels =Digitaria longiflora 

Syntherisma malacophylla Hitchc. =Digitaria lanuginosa 

Syntherisma sanguinalis sensu Hitchc. non (L.) Dulac =Digitaria ciliaris 

Thrasya campylostachya sensu Amshoff & Henrard non (Hack.) Chase =Thrasya granitica 

Thrasya cinerascens (Döll) Judz. =Paspalum cinerascens 

Thrasya guianensis Swallen =Thrasya scandens 

Thrasya reticulata Swallen =Paspalum multinervium 

Thrasya setosa Swallen =Thrasya paspaloides 

Trachypogon angustifolius E. Fourn. ex Hemsl. =Trachypogon spicatus 

Trachypogon angustifolius Nees ex Hack. =Trachypogon spicatus 

Trachypogon canescens Nees =Trachypogon spicatus 

Trachypogon capensis Trin. =Trachypogon spicatus 

Trachypogon capensis subvar mollis (Nees) Roberty =Trachypogon spicatus 

Trachypogon dactyloides (Steud.) E. Fourn. =Trachypogon spicatus 

Trachypogon densus Swallen =Trachypogon spicatus 

Trachypogon dissoluta Nees =Trachypogon spicatus 

Trachypogon durus Stapf =Trachypogon spicatus 

Trachypogon filifolius (Hack.) Hitchc. =Trachypogon spicatus 

Trachypogon glaucescens Pilg. =Trachypogon spicatus 

Trachypogon gouinii E. Fourn. =Trachypogon spicatus 

Trachypogon gracilis Andersson =Trachypogon spicatus 

Trachypogon gracilis var. ciliatus Andersson =Trachypogon spicatus 

Trachypogon gracilis var. hirtus Andersson =Trachypogon spicatus 

Trachypogon involutus Pilg. =Trachypogon spicatus 

Trachypogon karwinskyi (Hack.) Nash =Trachypogon spicatus 

Trachypogon ligularis Nees =Trachypogon spicatus 

Trachypogon macroglossus Trin. =Trachypogon spicatus 

Trachypogon mayaensis Wipff & S.D. Jones =Trachypogon spicatus 

Trachypogon micans Andersson =Trachypogon spicatus 

Trachypogon mollis Nees =Trachypogon spicatus 

Trachypogon montufarii (Kunth) Nees =Trachypogon spicatus 

Trachypogon montufarii var. bolivianus (Pilg.) Pilg. =Trachypogon spicatus 

Trachypogon montufarii var. grandiflorus Andersson =Trachypogon spicatus 

Trachypogon montufarii var. mollis (Nees) Andersson =Trachypogon spicatus 

Trachypogon montufarii var. pauciflorus Andersson =Trachypogon spicatus 

Trachypogon montufarii var. pilosus E. Fourn. =Trachypogon spicatus 

Trachypogon muelleri E. Fourn. =Trachypogon spicatus 

Trachypogon palmeri Nash =Trachypogon spicatus 

Trachypogon parviflorus Swallen =Trachypogon spicatus 

Trachypogon plumosus (Humb. & Bonpl. ex Willd.) Nees =Trachypogon spicatus 

Trachypogon plumosus var. montufarii (Kunth) Hack. =Trachypogon spicatus 

Trachypogon plumosus var. secundus (J. Presl) Beetle =Trachypogon spicatus 

Trachypogon plumosus subvar secundus (J. Presl) Hack. ex Henrard =Trachypogon spicatus 

Trachypogon polymorphus Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. bolivianus Pilg. =Trachypogon spicatus 

Trachypogon polymorphus var. canescens (Nees) Hack. =Trachypogon spicatus 

Trachypogon polymorphus subvar capensis (Trin.) Hack. =Trachypogon spicatus 

Trachypogon polymorphus subvar dactyloides (Steud.) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. dissolutus (Nees) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. filifolius Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. gouinii (E. Fourn.) Hack. =Trachypogon spicatus

172 


Trachypogon polymorphus subvar gracilis (Andersson) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. karwinskyi Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. ligularis (Nees) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. macroglossus (Trin.) Hack. =Trachypogon spicatus 

Trachypogon polymorphus subvar mollis (Nees) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. montufarii (Kunth) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. plumosus (Willd.) Hack. =Trachypogon spicatus 

Trachypogon polymorphus subvar secundus (J. Presl) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. thollonii Franch. =Trachypogon spicatus 

Trachypogon polymorphus var. truncatus (Nees) Hack. =Trachypogon spicatus 

Trachypogon polymorphus var. vestitus (Andersson) Hack. =Trachypogon spicatus 

Trachypogon preslii Andersson =Trachypogon spicatus 

Trachypogon preslii f. secundus (J. Presl) Andersson =Trachypogon spicatus 

Trachypogon ramosus Swallen =Trachypogon spicatus 

Trachypogon renvoizei Cat. Guerra =Trachypogon spicatus 

Trachypogon rigidifolius Swallen =Trachypogon spicatus 

Trachypogon rufus Nees =Hyparrhenia rufa 

Trachypogon thollonii (Franch.) Stapf =Trachypogon spicatus 

Trachypogon truncatus (Nees) Andersson =Trachypogon spicatus 

Trachypogon vestitus Andersson =Trachypogon spicatus 

Trichachne insularis (L.) Nees =Digitaria insularis 

Trichachne sacchariflora (Raddi) Nees =Digitaria insularis 

Trichachne tenuis Nees =Digitaria tenuis 

Tricholaena rosea Nees =Rhynchelytrum repens 

Tricuspis latifolia Griseb. =Gouinia latifolia 

Triodia flaccida (Döll) Hitchc. =Tridens flaccidus 

Tripsacum hermaphroditum L. =Anthephora hermaphrodita 

Tripsacum latifolium sensu Hitchc. (1922) non Hitchc. (1906) =Tripsacum australe 

Tripsacum laxum sensu Ostendorf non Nash =Tripsacum andersonii 

Uralepis flaccida Döll =Tridens flaccidus 

Urochloa arrecta (Hack. ex T. Durand & Schwinze) Morrone & Zuloaga =Brachiaria arrecta 

Urochloa decumbens (Stapf) R.D. Webster =Brachiaria decumbens 

Urochloa fasciculata (Sw.) R.D. Webster =Brachiaria fasciculata 

Urochloa fusca (Sw.) B.F. Hansen & Wunderlin =Brachiaria fasciculata 

Urochloa humidicola (Rendle) Morrone & Zuloaga =Brachiaria humidicola 

Urochloa maxima (Jacq.) R.D. Webster =Panicum maximum 

Urochloa mollis (Sw.) Morrone & Zuloaga =Brachiaria mollis 

Urochloa mutica (Forssk.) T.Q. Nguyen =Brachiaria mutica 

Urochloa plantaginea (Link) R.D. Webster =Brachiaria plantaginea 

Valota insularis (L.) Chase =Digitaria insularis 

Valota laxa (A. Spreng.) Hitchc. & Chase =Digitaria laxa 

Vilfa jacquemontii (Kunth) Trin. =Sporobolus jacquemontii 

Vilfa minutiflora Trin. =Sporobolus tenuissimus 

Vilfa pyramidalis (P. Beauv.) Trin. ex Steud. =Sporobolus jacquemontii 

Vilfa villifera Steud. =Sporobolus ciliatus 

REFERENCES 

(for a more extensive bibliography, see Judziewicz 1990) 

Amshoff, G.J.H. & J.T. Henrard. 1943. Gramineae. In A. Pulle, Flora of Suriname 1(1): 273-442. 

Cremers, G. & M. Hoff. 1993. Inventaire taxonomique des plantes de la Guyane française III - les Cyperaceae et les Poaceae. 

Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Sér. Patr. Génétique) 11: 1-213. 

Davidse, G., F.O. Zuloaga, & T.S. Filrueiras. 1995. Paspalum morichalense (Poaceae: Paniceae), a new aquatic species from 

South America. Novon 5(3): 234-237. 

Davidse, G. E.J. Judziewicz, & F.O. Zuloaga. 2004. Poaceae. In In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Hitchcock, A.S. 1922. Grasses of British Guiana. Contr. U.S. Natl. Herb. 22: 443-514. 

Judziewicz, E. 1990. Poaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 8: 1-725. 

Judziewicz, E. 1997. Poaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Gramineae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 343-375. 

Swallen, J. 1948. Gramineae. In B. Maguire et al., Plant explorations in Guiana in 1944, chiefly to the Tafelberg and the Kaieteur 

Plateau - I. Bull. Torrey Bot. Club 75: 81-91.


PONTEDERIACEAE 

C.N. Horn 

Eichhornia azurea (Sw.) Kunth 

Eichhornia crassipes (Mart.) Solms 

Eichhornia diversifolia (Vahl) Urb. 

Eichhornia heterosperma Alexander 

Eichhornia paniculata (Spreng.) Solms [probably naturalized] 

Heteranthera multiflora (Griseb.) C.N. Horn 

Pontederia rotundifolia L. 

Pontederia subovata (Seub.) Lowden 

Pontederia triflora (Endl. ex Seub.) G. Agostini, D. Velásquez & Velásquez 


Eichhornia polysperma [dubious name of uncertain origin] 

Eichhornia venezuelensis Velasquez =Eichhornia heterosperma 

Heteranthera diversifolia Vahl =Eichhornia diversifolia 

Heteranthera formosa Miq. =Eichhornia crassipes 

Heteranthera grandiflora Klotzsch [nomen nudum] =Eichhornia diversifolia 

Heteranthera reniformis var. multiflora Griseb. =Heteranthera multiflora 

Heteranthera zosterifolia Mart. [not in flora area, misident. for Eichhornia diversifolia] 

Pontederia azurea Sw. =Eichhornia azurea 

Pontederia cordata L. [not in flora area] 

Pontederia cordifolia Mart. ex Schult. & Schult. f. =Pontederia rotundifolia 

Pontederia crassipes Mart. =Eichhornia crassipes 

Pontederia eriantha Miq. =Pontederia rotundifolia 

Pontederia schomburgkiana Klotzsch [nomen nudum] =Pontederia subovata 

Reussia triflora Endl. ex Seub. =Pontederia triflora 

REFERENCES 

Castellanos, A. 1958. Las Pontederiaceae de Brasil. Arq. Jard. Bot. Rio de Janeiro 16: 149-216. 

Horn, C.N. 1994. Pontederiaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 15: 91-102. 

Horn, C.N. 2004. Pontederiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

8: 371-376. 

Jonker-Verhoef, A.M.E. 1953. Pontederiaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 81-90. 

Lowden, R.M. 1973. A revision of the genus Pontederia. Rhodora 75: 426-487. 

Amphiphyllum rigidum Gleason 

Cephalostemon affinis Körn. 

Cephalostemon gracilis (Poepp. & Endl.) R.H. Schomb. 

Cephalostemon microglochin Sandwith 

Duckea cyperaceoidea (Ducke) Maguire 

Duckea flava (Link) Maguire 

Duckea junciformis Maguire 

Duckea squarrosa (Willd.) Maguire 

Epidryos guayanensis Maguire 

Epidryos sp. A 

Guacamaya superba Maguire 

Kunhardtia radiata Maguire & Steyerm. 

Kunhardtia rhodantha Maguire 

Marahuacaea schomburgkii (Maguire) Maguire 

Monotrema aemulans Körn. 

Monotrema affine Maguire 

Monotrema bracteatum Maguire 

Monotrema bracteatum subsp. bracteatum 

Monotrema bracteatum subsp. major Maguire 

Monotrema xyridoides Gleason 

Phelpsiella ptericaulis Maguire 

Potarophytum riparium Sandwith 

Rapatea angustifolia Spruce ex Körn. 

Rapatea aracamuniana Steyerm. 

RAPATEACEAE 

P.E. Berry 





GU 

DA 


BO GU 

BO GU 

AM 

AM SU 

SU? 

AM 

AM 

AM 

AM 

AM BO 

BO GU 

AM 

AM 

AM 

AM BO 

AM 

AM GU 

AM 

AM BO 

AM BO 

AM 

AM 

AM 

GU 

AM 

AM

174 

Rapatea chimantensis Steyerm. 

Rapatea circasiana García-Barr. & L.E. Mora 

Rapatea fanshawei Maguire 

Rapatea fanshawei var. fanshawei 

Rapatea fanshawei var. minor Maguire 

Rapatea linearis Gleason 

Rapatea longipes Spruce ex Körn. 

Rapatea membranacea Maguire 

Rapatea paludosa Aubl. 

Rapatea paludosa var. paludosa 

Rapatea paludosa var. sessiliflora Maguire 

Rapatea saülensis B.M. Boom 

Rapatea scabra Maguire 

Rapatea spruceana Körn. 

Rapatea steyermarkii Maguire 

Rapatea ulei Pilg. 

Rapatea xiphoides Sandwith 

Rapatea yapacana Maguire 

Saxofridericia aculeata Körn. 

Saxofridericia compressa Maguire 

Saxofridericia duidae Maguire 

Saxofridericia grandis Maguire 

Saxofridericia inermis Ducke 

Saxofridericia petiolata Maguire 

Saxofridericia regalis R.H. Schomb. 

Saxofridericia sp. A 

Saxofridericia spongiosa Maguire 

Schoenocephalium cucullatum Maguire 

Schoenocephalium teretifolium Maguire 

Spathanthus bicolor Ducke 

Spathanthus unilateralis (Rudge) Desv. 

Spathanthus unilateralis var. abruptus Maguire 

Spathanthus unilateralis var. jenmanii (N.E. Br.) Maguire 

Spathanthus unilateralis var. unilateralis 

Stegolepis albiflora Steyerm. 

Stegolepis angustata Gleason 

Stegolepis breweri Maguire 

Stegolepis cardonae Maguire 

Stegolepis celiae Maguire 

Stegolepis choripetala Maguire 

Stegolepis ferruginea Baker f. 

Stegolepis gleasoniana Steyerm. 

Stegolepis grandis Maguire 

Stegolepis guianensis Klotzsch ex Körn. 

Stegolepis hitchcockii Maguire 

Stegolepis hitchcockii subsp. hitchcockii 

Stegolepis hitchcockii subsp. morichensis Maguire 

Stegolepis huberi Steyerm. 

Stegolepis humilis Steyerm. 

Stegolepis jauaensis Maguire 

Stegolepis ligulata Maguire 

Stegolepis linearis Gleason 

Stegolepis maguireana Steyerm. 

Stegolepis membranacea Maguire 

Stegolepis microcephala Maguire 

Stegolepis minor Steyerm. 

Stegolepis neblinensis Maguire [Sierra de la Neblina] 

Stegolepis parvipetala Steyerm. 

Stegolepis pauciflora Gleason 

Stegolepis perligulata Maguire 

Stegolepis ptaritepuiensis Steyerm. 

Stegolepis pulchella Maguire 

Stegolepis pungens Gleason 

Stegolepis sp. nov. [Mt. Ayanganna] 


AM 

AM 

BO 

BO GU 

BO GU 

GU 

GU 

GU 



SU 

SU FG 

AM 

AM 

BO GU 

GU FG 

GU 

AM 

GU SU FG 

AM 

AM 

AM 

AM 

AM 

BO GU SU 

AM 

AM 

AM 

AM 

AM BO 

BO GU SU FG 

GU? FG 

GU FG 

BO GU SU FG 

AM BO 

BO GU 

BO 

BO 

AM BO 

AM BO 

GU 

AM 

AM BO 

BO GU 

AM 

AM 

AM 

BO GU 

BO 

BO 

BO 

AM 

BO 

AM 

BO 

BO 

AM 

BO 

AM 

BO 

BO GU 

AM 

AM 

GU

Stegolepis squarrosa Maguire 

Stegolepis steyermarkii Maguire 

Stegolepis terramarensis Steyerm. 

Stegolepis vivipara Maguire 

Stegolepis wurdackii Maguire 

Stegolepis wurdackii subsp. chimantensis Maguire 

Stegolepis wurdackii subsp. wurdackii [Sierra de la Neblina] 

Windsorina guianensis Gleason 



Amphiphyllum schomburgkii Maguire =Marahuacaea schomburgkii 

Cephalostemon cyperaceoides Ducke =Duckea cyperaceoidea 

Cephalostemon flavus (Link) Steyerm. =Duckea flava 

Cephalostemon junciformis (Maguire) Steyerm. =Duckea junciformis 

Cephalostemon squarrosus (Willd.) Körn. =Duckea squarrosa 

Mnasium paludosum (Aubl.) Willd. =Rapatea paludosa 

Mnasium unilaterale Rudge =Spathanthus unilateralis 

Mnasium unilateralis Rudge =Spathanthus unilateralis 

Rapatea longipes var. modesta (Maguire) Maguire =Rapatea longipes 

Rapatea modesta Maguire =Rapatea longipes 

Rapatea schultesiana García-Barr. & L.E. Mora =Rapatea paludosa 

Rapatea unilateralis (Rudge) Schult. & Schult. f. =Spathanthus unilateralis 

Rapatea viscosa Gleason =Saxofridericia regalis 

Saxofridericia australis Gleason =Saxofridericia aculeata 

Saxofridericia duidae subsp. marahuacensis Maguire =Saxofridericia duidae 

Saxofridericia pandanoides Linden & André =Saxofridericia aculeata 

Saxofridericia subcordata Körn. =Saxofridericia aculeata 

Schoenocephalium coriaceum Maguire =Schoenocephalium cucullatum 

Schoenocephalium sipapoanum Maguire =Kunhardtia rhodantha 

Spathanthus jenmanii N.E. Br. =Spathanthus unilateralis var. jenmanii 

Stegolepis aemulans (Körn.) Benth. & Hook. =Monotrema aemulans 

Stegolepis grandis subsp. jauensis Maguire =Stegolepis grandis 

Stegolepis grandis subsp. phelpsiae Maguire =Stegolepis grandis 

Stegolepis parvipetala subsp. chimantensis Maguire =Stegolepis parvipetala 

REFERENCES 

Berry, P.E. 2004. Rapateaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

8: 413-472. 

Boom, B.M. 1994. A new species of Rapatea section Paludosa from French Guiana. Brittonia 46(4): 314-316. 

Boom, B.M. 1997. Rapateaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Rapateaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 326-328. 

Maguire, B. 1958. Rapateaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part 3. Mem. 


Maguire, B. 1965. Rapateaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 6. Mem. New York Bot. 

Gard. 12(3): 1-285. 

Maguire, B. 1982. Rapateaceae. In Z. Luces de Febres, Flora de Venezuela 6(2): 85-203. 

Uittien, H. 1938. Rapateaceae. In A. Pulle, Flora of Suriname 1(1): 191-196. 

Ruppia maritima L. 

RUPPIACEAE 

R. Haynes, L.B. Holm-Nielsen 

REFERENCES 

Haynes, R.R., L.B. Holm-Nielsen & D.H. Les. 1998. Ruppiaceae. Families and Genera of Vascular Plants 4: 445—448. 

Smilax cinnamommea Desf. ex DC. 

Smilax cordato-ovata Rich. 

AM 

SMILACACEAE 

P.E. Berry, R.A. DePhilipps†, J.F. Gaskin 

BO 

AM BO 

BO 

AM 

BO 

BO GU 

GU 

GU SU FG 

FG 

FG

176 

Smilax cumanensis Humb. & Bonpl. ex Willd. 

Smilax cuspidata Poir. 

Smilax immersa A.C. Sm. 

Smilax krukovii A.C. Sm. 

Smilax kunthii Killip & C.V. Morton 

Smilax lappacea Humb. & Bonpl. ex Willd. 

Smilax latipes Gleason 

Smilax longifolia Rich. 

Smilax maypurensis Humb. & Bonpl. ex Willd. 

Smilax megalophylla Duhamel 

Smilax pittieriana Steyerm. 

Smilax poeppigii Kunth 

Smilax pseudosyphilitica Kunth 

Smilax quinquenervia Vell. 

Smilax riedeliana A. DC. 

Smilax santaremensis DC. 

Smilax saülensis J.D. Mitch. 

Smilax schomburgkiana Kunth 

Smilax staminea Griseb. 

Smilax syphilitica Humb. & Bonpl. ex Willd. 

Smilax talbotiana A. DC. 

Smilax triplinervia Humb. & Bonpl. ex Willd. 



Smilax auraimensis Steyerm. =Smilax pittieriana 

Smilax chimantensis Steyerm. & Maguire =Smilax pittieriana 

Smilax duidae Steyerm. =Smilax syphilitica 

Smilax flavescens Ham. =Smilax longifolia 

Smilax floribunda Ham. ? =Smilax cuspidata 

Smilax globifera G. Mey. =Smilax cumanensis 

Smilax hostmanniana Kunth =Smilax cumanensis 

Smilax jauaensis Steyerm. & Maguire =Smilax staminea 

Smilax lasseriana Steyerm. =Smilax staminea 

Smilax papyracea Duhamel =Smilax longifolia 

Smilax pirarensis Kunth & M.R. Schomb. =Smilax cumanensis 

Smilax procera Griseb. =Smilax quinquenervia 

Smilax staminea Griseb. f. obtusata Steyerm. =Smilax staminea 

Smilax surinamensis Miq. =Smilax cumanensis 

REFERENCES 

Andreata, R.H.P. 1980. Smilax Linnaeus (Smilacaceae): Ensaio para uma revisão das especies brasileiras. Arq. Jard. Bot. Rio de 

Janeiro 24: 179-301.Lachman-White, D.A., Adams, C.D. and U.O. Trotz. 1987. A Guide to Medicinal Plants of Coastal 

Guyana: CSC Technical Publication Series No. 225. 350 pp. London: Commonwealth Science Council. (Smilax, pp. 256- 

257 

Gaskin, J.F. & P.E. Berry. 2005. Smilacaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 184-193. 

Lemée, A. 1955. Smilax. Flore de la Guyane Française 1: 321-324. Paris: Paul Lechevalier. 

Mitchell, J.D. 1994. Smilax saülensis, a new species from French Guiana. Brittonia 46(3): 225-227. 

Mitchell, J.D. 1997. Smilacaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Ostendorf, F.W. 1962. Nuttige Planten en Sierplanten in Suriname. Landbouwproefstation in Suriname Bulletin No. 79. 325 pp. 

(Smilax, pp. 216-217). 

Roosmalen, M.G.M. van. 1985. Fruits of the Guianan Flora. 483 pp. Utrecht: Institute of Systematic Botany, Utrecht University; 

Wageningen: Silvicultural Department, Wageningen Agricultural University. (Smilax, pp. 254-256) 

Sipman, H. 1979. Liliaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 442-456. 

Phenakospermum guyannense (Rich.) Endl. ex Miq. 

STRELITZIACEAE 

L.L. Andersson, P.J.M. Maas 


Ravenala guyannensis (Rich.) Petersen =Phenakospermum guyannense 

Urania guyannensis Rich. =Phenakospermum guyannense 


FG 

GU 

GU FG? 

GU SU FG 

AM 

GU SU FG 

SU? FG 

AM BO 

GU SU FG? 

AM BO GU? 

AM BO GU? 

FG 

FG 

FG 

GU SU FG 

FG 




GU 

AM 

AM BO GU SU FG


REFERENCES 

Andersson, L.L. 2005. Strelitziaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 281-283. 

Maas, P.J.M. 1985. Musaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 1: 1-28. 


Maas, P. & H. Maas-van de Kamer. 1997. Strelitziaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. 



Tacca parkeri Seem. 

TACCACEAE 

P.E. Berry, Z.S. Rogers 


Tacca parkeri var. lanceolata H. Limpr. =Tacca parkeri 

Tacca sprucei Benth. =Tacca parkeri 

Tacca ulei H. Limpr. =Tacca parkeri 

REFERENCES 

Berry, P.E. & Z.S. Rogers. 2005. Taccaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 295-296. 

Drenth, E. 1972. A revision of the family Taccaceae. Blumea 20: 367-406. 

Thurnia jenmanii Hook. f. 

Thurnia polycephala Schnee 

Thurnia sphaerocephala (Rudge) Hook. f. 

Thurnia sphaerocephala subsp. 1 

THURNIACEAE 

P.E. Berry, R. Riina, M.T. Strong 


Mnasium sphaerocephala Rudge =Thurnia sphaerocephala 

REFERENCES 

Mori, S.A. 1997. Thurniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. 



Schnee, L. 1943. El genero Thurnia (con descripción de una nueva especie). Bol. Soc. Venez. Cienc. Nat. 8(55):_241-243. 

Riina, R. & P.E. Berry. 2005. Thurniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 330-332. 

Uittien, H. 1938. Thurniaceae. In A. Pulle, Flora of Suriname 1(1): 188-190. 

Peltophyllum luteum Gardner 

Sciaphila albescens Benth. 

Sciaphila picta Miers 

Sciaphila polygyna Maas 

Sciaphila purpurea Benth. 

Sciaphila rubra Maas 

Soridium spruceanum Miers 

Triuris hexophthalma Maas 

Triuris hyalina Miers 

TRIURIDACEAE 

P.J.M. Maas, H. Maas 


Sciaphila brevipes S.F. Blake =Soridium spruceanum 

Sciaphila guianensis Sandwith =Sciaphila albescens 

Sciaphila spruceana (Miers) Engl. =Soridium spruceanum 

Sciaphila spruceana (Miers) Benth. & Hook. f. =Soridium spruceanum 

Soridium guianense (Sandwith) Giesen =Sciaphila albescens 

AM GU SU? 

GU 

AM 

AM BO GU 

GU 

SU FG 

GU 


GU? 

FG 

AM BO GU 

AM 

AM SU FG 

GU 

AM GU SU

178 


REFERENCES 

Jonker, F.P. 1943. Triuridaceae. In A. Pulle, Flora of Suriname 1(1): 461-466. 

Maas, P.J.M. & T. Rübsamen. 1986. Triuridaceae. Flora Neotropica Monogr. 40: 1-55. 

Maas, P.J.M. & H. Maas-van de Kamer. 1989. Triuridaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 5: 1-16. 

Maas, P.J.M. & H. Maas. 2005. Triuridaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 368-371. 

Maas-van de Kamer, H. & P. Maas. 1997. Triuridaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. 


Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1:367-371. 

Typha domingensis Pers. 

Typha angustifolia L. [not in flora area] 

TYPHACEAE 

L.J. Raechal, M.T. Strong 


REFERENCES 

Lanjouw, J. 1938. Typhaceae. In A. Pulle, Flora of Suriname 1(1): 252-254. 

Raechal, L.J. 2005. Typhaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 385. 

Barbacenia celiae Maguire 

Vellozia tubiflora (A. Rich.) Kunth 

VELLOZIACEAE 

P.E. Berry, J.S. Miller 


Barbacenia alexandrinae R.H. Schomb. ex Hook. =Vellozia tubiflora 

Barbacenia uaipanensis Maguire (Bromeliaceae) =Ayensua uaipanensis 

Pleurostima celiae (Maguire) Menezes =Barbacenia celiae 

Radia tubiflora A. Rich. =Vellozia tubiflora 

Vellozia alexandrinae (R.H. Schomb. ex Hook.) Goethart & Henrard =Vellozia tubiflora 

Vellozia duidae Steyerm. =Vellozia tubiflora 

Vellozia uaipanensis (Maguire) L.B. Sm. (Bromeliaceae) =Ayensua uaipanensis 

REFERENCES 

Maguire, B. 1969. Velloziaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part VIII. Mem. New York 

Bot. Gard. 18(2): 1-290. 

Miller, J.S. & P.E. Berry. 2005. Velloziaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 404-406. 

Smith, L.B. & E.S. Ayensu. 1976. A revision of American Velloziaceae. Smiths. Contrib. Bot. 30: 1-172. 

Abolboda acaulis Maguire 

Abolboda acaulis var. acaulis 

Abolboda acaulis var. scaposa Kral 

Abolboda acicularis Idrobo & L.B. Sm. 

Abolboda acicularis var. acicularis 

Abolboda acicularis var. granularis Maguire 

Abolboda americana (Aubl.) Lanj. 

Abolboda bella Maguire 

Abolboda ciliata Maguire & Wurdack 

Abolboda dunstervillei Maguire ex Kral 

Abolboda ebracteata Maguire & Wurdack 

Abolboda ebracteata var. brevifolia Maguire 

Abolboda ebracteata var. ebracteata 

Abolboda x glomerata Maguire [sterile hybrid] 

Abolboda grandis Griseb. 

Abolboda grandis var. grandis 

XYRIDACEAE 

R. Kral, M.T. Strong 


AM 

AM BO GU 

AM BO GU 

AM BO GU 

AM BO GU 

AM BO 

AM BO 

AM 


AM 

AM 

AM 

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AM BO GU SU FG

Abolboda grandis var. rigida Malme 

Abolboda killipii Lasser 

Abolboda linearifolia Maguire 

Abolboda macrostachya Spruce ex Malme 

Abolboda macrostachya var. macrostachya 

Abolboda macrostachya var. robustior Steyerm. 

Abolboda neblinae Maguire 

Abolboda paniculata Maguire 

Abolboda poarchon Seub. 

Abolboda pulchella Bonpl. 

Abolboda scabrida Kral 

Abolboda sprucei Malme 

Abolboda uniflora Maguire 

Achlyphila disticha Maguire & Wurdack 

Aratitiyopea lopezii (L.B. Sm.) Steyerm. & P.E. Berry 

Aratitiyopea lopezii var. lopezii 

Orectanthe ptaritepuiana (Steyerm.) Maguire 

Orectanthe sceptrum (Oliv.) Maguire 

Orectanthe sceptrum subsp. sceptrum 

Xyris albescens Steyerm. [Mt. Roraima] 

Xyris anceps Lam. 

Xyris aquatica Idrobo & L.B. Sm. 

Xyris aracamunae Kral 

Xyris arachnoidea Maguire & L.B. Sm. 

Xyris atriceps Malme 

Xyris atriceps subsp. atriceps 

Xyris atriceps subsp. chimantensis Maguire & L.B. Sm. 

Xyris atriceps subsp. marahuacae Kral & L.B. Sm. 

Xyris atriceps subsp. neblinensis Maguire & L.B. Sm. 

Xyris bicephala Gleason 

Xyris bicostata Maguire & L.B. Sm. 

Xyris bicostata var. bicostata 

Xyris bicostata var. tillettii (L.B. Sm.) Kral 

Xyris byssacea Kral 

Xyris carinata Maguire & L.B. Sm. 

Xyris chimantae Kral & L.B. Sm. 

Xyris concinna N.E. Br. [Mt. Roraima] 

Xyris connosepala Lanj. & Lindeman 

Xyris consolida Kral & L.B. Sm. 

Xyris contracta Maguire & L.B. Sm. 

Xyris cryptantha Maguire & L.B. Sm. 

Xyris culmenicola Steyerm. 

Xyris cyperoides Gleason 

Xyris decussata Gleason [Mt. Roraima] 

Xyris delicatula Maguire & L.B. Sm. 

Xyris esmeraldae Steyerm. 

Xyris fallax Malme 

Xyris frequens Maguire & L.B. Sm. 

Xyris frondosa Maguire & L.B. Sm. 

Xyris fuliginea Kral & L.B. Sm. 

Xyris globosa L.A. Nilsson 

Xyris gongylospica Kral 

Xyris graniticola Kral 

Xyris guianensis Steud. 

Xyris huberi Kral & L.B. Sm. 

Xyris hymenachne Mart. [Mt. Roraima] 

Xyris involucrata Nees 

Xyris juncifolia Maguire & L.B. Sm. 

Xyris jupicai Rich. 

Xyris kukenaniana Kral [expected in GU] 

Xyris lacerata Pohl ex Seub. [expected in GU] 

Xyris lanulobractea Steyerm. 

Xyris laxifolia Mart. 

Xyris laxifolia var. laxifolia 


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AM BO GU 

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AM 

AM 

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180 


Xyris liesneri Kral 

Xyris lithophila Kral & L.B. Sm. 

Xyris lomatophylla Mart. 

Xyris lugubris Malme 

Xyris malmeana L.B. Sm. 

Xyris melanovaginata Kral & L.B. Sm. 

Xyris mima L.B. Sm. & Downs 

Xyris neblinae Maguire & L.B. Sm. 

Xyris oblata Kral & L.B. Sm. 

Xyris oxylepis Idrobo & L.B. Sm. 

Xyris paraensis Poepp. ex Kunth 

Xyris paraensis var. longiceps (Malme) L.B. Sm. & Downs 

Xyris paraensis var. paraensis 

Xyris pratensis Maguire & L.B. Sm. 

Xyris prolificans Kral 

Xyris ptariana Steyerm. 

Xyris riparia Maguire & L.B. Sm. 

Xyris roraimae Malme [Mt. Roraima] 

Xyris rubrolimbata Heimerl 

Xyris savanensis Miq. 

Xyris schneeana L.B. Sm. & Steyerm. 

Xyris setigera Oliv. ex Thurn 

Xyris seubertii L.A. Nilsson [Mt. Roraima] 

Xyris spathacea Lanj. 

Xyris spruceana Malme 

Xyris stenophylloides Malme 

Xyris stenostachya Steyerm. 

Xyris subglabrata Malme 

Xyris subuniflora Malme 

Xyris sulcatifolia Kral 

Xyris surinamensis Spreng. 

Xyris tatei Malme 

Xyris teinosperma Idrobo & L.B. Sm. 

Xyris tenella Kunth 

Xyris tenella var. leprieurii Malme 

Xyris tenella var. tenella 

Xyris thysanolepis Maguire & L.B. Sm. 

Xyris thysanolepis var. sipapoa (Maguire & L.B. Sm.) Kral & L.B. Sm. 

[expected in GU] 

Xyris thysanolepis var. thysanolepis [expected in GU] 

Xyris toronoana Kral 

Xyris uleana Malme 

Xyris uleana var. angustifolia Lanj. 

Xyris uleana var. uleana 

Xyris valdeapiculata Kral 

Xyris witsenioides Oliv. ex Thurn [Mt. Roraima] 

Xyris wurdackii Maguire & L.B. Sm. 

Xyris wurdackii subsp. wurdackii 

Xyris xiphophylla Maguire & L.B. Sm. 


Abolboda americana var. imberbis (Kunth) Maguire =Abolboda americana 

Abolboda brasiliensis Kunth =Abolboda pulchella 

Abolboda chapadensis var. pauciflora Hoehne =Abolboda pulchella 

Abolboda excelsa Malme =Abolboda macrostachya var. robustior 

Abolboda gracilis Huber =Abolboda pulchella 

Abolboda grandis var. guayanensis Maguire =Abolboda grandis var. rigida 

Abolboda grandis var. minor Spruce ex Malme =Abolboda grandis 

Abolboda imberbis Kunth =Abolboda americana 

Abolboda macrostachya var. angustior Maguire =Abolboda macrostachya 

Abolboda minima Maguire =Abolboda killipii 

Abolboda pervaginata Malme =Abolboda grandis var. rigida 

Abolboda poeppigii Kunth =Abolboda americana 

Abolboda psammophila Maguire =Abolboda killipii 

AM 

BO 

AM 

AM 

AM 

AM 

AM 

AM 

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AM 

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AM BO 

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Abolboda ptaritepuiana Steyerm. =Orectanthe ptaritepuiana 

Abolboda rigida Gleason =Abolboda macrostachya var. robustior 

Abolboda rigida (Malme) Steyerm. =Abolboda grandis var. rigida 

Abolboda sceptrum Oliv. =Orectanthe sceptrum 

Abolboda schultesii Idrobo & L.B. Sm. =Abolboda sprucei 

Orectanthe sceptrum subsp. occidentalis Maguire =Orectanthe sceptrum 

Xyris acuminata Miq. ex Steud. =Xyris jupicai 

Xyris ambigua Beyr. ex Kunth =Xyris jupicai 

Xyris americana Aubl. =Abolboda americana 

Xyris anceps Pers., [nom. illeg.] =Xyris jupicai 

Xyris applanata Idrobo & L.B. Sm. =Xyris spruceana 

Xyris arenicola Miq. =Xyris jupicai 

Xyris arescens Kunth =Xyris hymenachne 

Xyris asterocephala Seub. =Xyris involucrata 

Xyris calcarata Heimerl =Xyris seubertii 

Xyris capillaris Steyerm. =Xyris subuniflora 

Xyris caroliniana var. major (Mart.) Idrobo & L.B. Sm. =Xyris laxifolia var. laxifolia 

Xyris coerulea Lam. =Abolboda americana 

Xyris communis Kunth =Xyris jupicai 

Xyris connectens Malme =Xyris uleana var. angustifolia 

Xyris dolichosperma Lanj. =Xyris fallax 

Xyris duidensis Malme =Xyris uleana var. uleana 

Xyris epicarae Kral & L.B. Sm. =Xyris cyperoides 

Xyris eriophylla Rchb. [nomen nudum] =Xyris surinamensis 

Xyris eriophylla Rchb. ex Kunth =Xyris surinamensis 

Xyris erythema Maguire & L.B. Sm. =Xyris fallax 

Xyris exserta Idrobo & L.B. Sm. =Xyris spathacea 

Xyris filiscapa Malme =Xyris guianensis 

Xyris foveolata Kral & L.B. Sm. =Xyris huberi 

Xyris fusiformis Maguire & L.B. Sm. =Xyris oxylepis 

Xyris garcia-barrigae Idrobo & L.B. Sm. =Xyris subglabrata 

Xyris gardneri Malme =Xyris guianensis 

Xyris glabrata sensu Griseb. et auct. =Xyris malmeana 

Xyris guianensis Klotzsch [nomen nudum] =Xyris jupicai 

Xyris gymnoptera Griseb. =Xyris jupicai 

Xyris jauana L.B. Sm. & Steyerm. =Xyris thysanolepis var. thysanolepis 

Xyris jupicae Michx. =Xyris jupicai 

Xyris jupicae var. brachylepis Malme =Xyris jupicai 

Xyris jupicae var. major (Mart.) L.B. Sm. & Downs =Xyris laxifolia var. laxifolia 

Xyris laxifolia var. major Mart. =Xyris laxifolia var. laxifolia 

Xyris laxifolia var. minor Mart. =Xyris jupicai 

Xyris leptostachya Malme =Xyris paraensis var. longiceps 

Xyris longiceps Malme =Xyris paraensis var. longiceps 

Xyris macrocephala auct. non Vahl =Xyris laxifolia 

Xyris ? macrocephala Vahl [nom. confus.] =Xyris laxifolia var. laxifolia 

Xyris macrocephala var. major (Mart.) L.A. Nilsson =Xyris laxifolia var. laxifolia 

Xyris macrocephala f. minor (Mart.) M. Kuhlm. & E. Kuhn =Xyris jupicai 

Xyris rugulosa Maguire & L.B. Sm. =Xyris esmeraldae 

Xyris scabridula Steyerm., [nom. illeg.] =Xyris sulcatifolia 

Xyris setigera var. elongata Steyerm. =Xyris setigera 

Xyris sipapoa Maguire & L.B. Sm. =Xyris thysanolepis var. sipapoa 

Xyris steyermarkii Maguire & L.B. Sm. =Xyris tenella var. tenella 

Xyris submontana Gleason =Xyris hymenachne 

Xyris surinamensis Miq. =Xyris jupicai 

Xyris tillettii L.B. Sm. =Xyris bicostata var. tillettii 

Xyris trisperma Kral & L.B. Sm. =Xyris mima 

Xyris vaginata Spreng. =Abolboda pulchella 

Xyris vaupesana Idrobo & L.B. Sm. =Xyris uleana var. uleana 

Xyris yapobodensis Idrobo & L.B. Sm. =Xyris spathacea 

Xyris yutajensis Maguire & L.B. Sm. =Xyris tenella var. tenella 

REFERENCES 

Kral, R. 1988. The genus Xyris in Venezuela and contiguous northern South America. Ann. Missouri Bot. Gard. 75: 522-722. 

Kral, R. 1992. A treatment of American Xyridaceae exclusive of Xyris. Ann. Missouri Bot. Gard. 79: 819-885.

182 


Kral, R. 1994. Xyridaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 15: 3-90. 

Kral, R. 2005. Xyridaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 9: 

526-574. 

Lanjouw, J. 1938. Xyridaceae. In A. Pulle, Flora of Suriname 1(1): 225-248. 

Maguire, B. 1958. Xyridaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part III. Mem. 


Maguire, B. & L.B. Smith. 1964. Xyridaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland 

part V. Mem. New York Bot. Gard. 10(5): 1-278. 

Mennega, A.M.W., J.C. Lindeman, & A.R.A. Görts-van Rijn. 1968. Xyridaceae (additions and corrections). In A. Pulle & J. 

Lanjouw, Flora of Suriname 1(2): 340-342. 

Mori, S.A. 1997. Xyridaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide 


New York Botanical Garden 76, pt. 1: 371. 

Hedychium coronarium J. Koenig [cultivated, naturalized] 

Renealmia alpinia (Rottb.) Maas 

Renealmia aromatica (Aubl.) Griseb. 

Renealmia cernua (Sw. ex Roem. & Schult.) J.F. Macbr. 

Renealmia floribunda K. Schum. 

Renealmia guianensis Maas 

Renealmia microcalyx Maas & H. Maas 

Renealmia monosperma Miq. 

Renealmia orinocensis Rusby 

Renealmia thyrsoidea (Ruiz & Pav.) Poepp. 

Renealmia thyrsoidea subsp. chrysantha Maas 

Renealmia thyrsoidea subsp. thyrsoidea 

Zingiber spp. [several species cultivated, possibly escaping] 

ZINGIBERACEAE 

H. Maas, P.J.M. Maas 


Alpinia aromatica Aubl. =Renealmia aromatica 

Alpinia latifolia Willd. ex Link (Marantaceae) =Calathea latifolia 

Alpinia spiralis Jacq. (Costaceae) =Costus spiralis var. spiralis 

Alpinia tubulata Ker Gawl. =Renealmia alpinia 

Amomum alpinia Rottb. =Renealmia alpinia 

Amomum thyrsoideum Ruiz & Pav. =Renealmia thyrsoidea 

Renealmia exaltata L. f. =Renealmia alpinia 

Renealmia gracilis Petersen =Renealmia guianensis 

Renealmia humilis Petersen =Renealmia monosperma 

Renealmia lutea J.R. Johnst. =Renealmia aromatica 

Renealmia monostachia L. (Bromeliaceae) =Guzmania monostachia 

Renealmia occidentalis (Sw.) Sweet =Renealmia aromatica 

Renealmia occidentalis var. macrocarpa Loes. =Renealmia aromatica 

Renealmia occidentalis var. occidentalis =Renealmia aromatica 

Renealmia occidentalis var. ß cylindrica Petersen =Renealmia aromatica 

Renealmia pedicellaris Gleason =Renealmia floribunda 

Renealmia pendula C.F. Gaertn. (Bromeliaceae) =Catopsis berteroniana 

Renealmia polystachia L. (Bromeliaceae) =Tillandsia polystachia 

Renealmia raja Petersen =Renealmia alpinia 

Renealmia recurvata var. ß (Bromeliaceae) =Tillandsia recurvata 

Renealmia usneoides (Bromeliaceae) =Tillandsia usneoides 

GU SU FG 


AM BO DA GU FG? 

SU 



AM 



AM GU SU 

GU SU 

AM 

REFERENCES 

Maas, P.J.M. 1977. Renealmia. Flora Neotropica Monogr. 18: 1-161. 

Maas, P.J.M. 1979. Zingiberaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 416-441. 

Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A). 

Maas, P. & H. Maas-van de Kamer. 1997. Zingiberaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. 



Maas-van de Kamer, H. & P. Maas. 2005. Zingiberaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 

the Venezuelan Guayana 9: 576-580.

Zostera marina L. [record by Lemée extremely dubious] 


ZOSTERACEAE 

REFERENCES 

Phillips, R.C. & E.G. Meñez. 1988. Seagrasses. Smithsonian Contrib. Marine Sci. 34: 1-104. 

DICOTYLEDONEA 

ACANTHACEAE 

D. Wasshausen 

Anisacanthus secundus Leonard 

Aphelandra aurantiaca (Schweidw.) Lindl. 

Aphelandra aurantiaca var. aurantiaca 

Aphelandra fasciculata Wassh. 

Aphelandra pilosa Leonard 

Aphelandra pulcherrima (Jacq.) Kunth 

Aphelandra scabra (Vahl) Sm. 

Asystasia gangetica (L.) T. Anderson [cultivated, persists as an escape] 

Barleria cristata L. 

Barleria lupulina Lindl. [introduced, possibly naturalized] 

Blechum pyramidatum (Lam.) Urb. 

Bravaisia integerrima (Spreng.) Standl. 

Dicliptera ciliaris J. Juss. 

Dicliptera granvillei Wassh. 

Dicliptera knappii Wassh. 

Dicliptera sexangularis (L.) J. Juss. [expected in GU] 

Elytraria imbricata (Vahl) Pers. 

Eranthemum pulchellum Andr. [cultivated, naturalized] 

Gynocraterium guianense Bremek. 

Hemigraphis alternata (Burm. f.) T. Anderson [cultivated, persists as an 

escape] 

Herpetacanthus rotundatus (Lindau) Bremek. 

Hygrophila costata Nees 

Hygrophila erecta (Burm. f.) Hochr. [introduced] 

Hygrophila sandwithii Bremek. 

Hygrophila surinamensis Bremek. [expected in GU] 

Justicia betonica L. [introduced] 

Justicia breteleri Wassh. 

Justicia calycina (Nees) V.A.W. Graham 

Justicia carthaginensis Jacq. 

Justicia cataractae Leonard 

Justicia cayennensis (Nees) Lindau 

Justicia cayennensis var. cayennensis 

Justicia cayennensis var. parvifolia Bremek. 

Justicia comata (L.) Lam. 

Justicia coppenamensis Wassh. [nom. nov. for Beloperone modesta Bremek.] 

Justicia delascioi Wassh. 

Justicia guianensis (N.E. Br.) Wassh. 

Justicia huberi Wassh. 

Justicia hylaea Leonard 

Justicia kunhardtii Leonard 

Justicia laevilinguis (Nees) Lindau [expected in GU] 

Justicia maguirei Wassh. 

Justicia mcdowellii Wassh. 

Justicia monachinoi Wassh. 

Justicia moritziana Wassh. 

Justicia nuriana Wassh. 

Justicia oldemanii Wassh. 

Justicia panarensis Wassh. 

Justicia parguazensis Wassh. 

FG? 

BO GU SU FG 

GU SU FG 

GU SU FG 

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AM BO 

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AM BO GU SU 

AM DA 

DA 

GU 

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184 


Justicia parimensis Wassh. 

Justicia paruana Wassh. 

Justicia pectoralis Jacq. 

Justicia polystachya Lam. [expected in GU] 

Justicia potarensis (Bremek.) Wassh. 

Justicia prevostiae Wassh. 

Justicia schomburgkiana (Nees) V.A.W. Graham 

Justicia secunda Vahl 

Justicia sprucei V.A.W. Graham 

Justicia uvida Wassh. 

Justicia wilhelminensis Wassh. 

Justicia wurdackii Leonard 

Lepidagathis alopecuroidea (Vahl) R. Br. ex Griseb. 

Lepidagathis gracilis (Bremek.) Wassh. [expected in GU] 

Lepidagathis medicaginea (Bremek.) Wassh. [expected in GU] 

Lepidagathis nickeriensis Wassh. [expected in GU] 

Lepidagathis surinamensis (Bremek.) Wassh. 

Megaskepasma erythrochlamys Lindau 

Nelsonia campestris R. Br. 

Odontonema album V.M. Baum 

Odontonema bracteolatum (Jacq.) Kuntze 

Odontonema liesneri Wassh. 

Odontonema mazarunensis Wassh. 

Odontonema schomburgkianum (Nees) Kuntze 

Pachystachys coccinea (Aubl.) Nees 

Pachystachys spicata (Ruiz & Pav.) Wassh. 

Polylychnis fulgens Bremek. 

Polylychnis radicans (Nees) Wassh. 

Pseuderanthemum maguirei Leonard 

Pulchranthus congestus (Lindau) V.M. Baum, Reveal & Nowicke 

Pulchranthus surinamensis (Bremek.) V.M. Baum, Reveal & Nowicke 

Pulchranthus variegatus (Aubl.) V.M. Baum, Reveal & Nowicke 

Ruellia bolivarensis Wassh. 

Ruellia delascioi Wassh. 

Ruellia geminiflora Kunth 

Ruellia geminiflora var. angustifolia (Nees) Griseb. 

Ruellia geminiflora var. geminiflora 

Ruellia inflata Rich. 

Ruellia liesneri Wassh. 

Ruellia longifolia Rich. 

Ruellia macrophylla Vahl 

Ruellia malaca Leonard 

Ruellia menthoides (Nees) Hiern 

Ruellia microcalyx (Nees) Lindau 

Ruellia prostrata Poir. 

Ruellia rubra Aubl. [includes several varieties without combinations in 

Ruellia] 

Ruellia saülensis Wassh. 

Ruellia sprucei Lindau 

Ruellia steyermarkii Wassh. 

Ruellia tuberosa L. [naturalized] 

Ruellia violacea Aubl. 

Ruellia wurdackii Wassh. 

Sanchezia lutea Leonard 

Sanchezia pennellii Leonard 

Staurogyne fockeana Bremek. 

Staurogyne miqueliana Kuntze 

Staurogyne repens (Nees) Kuntze 

Staurogyne spraguei Wassh. 

Staurogyne stahelii Bremek. 

Staurogyne stolonifera (Nees) Kuntze 

Staurogyne trinitensis Leonard 

Staurogyne wullschlaegeliana Bremek. 

Teliostachya cataractae Nees [=Lepidagathis sp.] 

AM 

AM 


AM BO SU FG 

GU SU FG 

FG 

AM BO DA GU 


FG 

BO 

SU 

AM 


AM BO SU 

BO SU FG 

BO SU 

BO GU SU FG 

SU 

FG? 

BO 

AM BO 

AM 

GU 

GU 

GU FG 

AM GU FG 

SU FG 

GU FG 

AM 

FG 

SU 

AM SU FG 

BO 

AM 



GU SU FG 

SU FG 

AM 

SU FG 

BO 

AM BO 

AM 

GU SU 

GU 

GU SU FG 

FG 

AM 

BO 

BO GU SU FG 

FG 

AM BO 

GU 

DA 

SU 

SU 

AM BO 

GU 

GU 

BO DA GU 

SU 

SU 

SU 

SU 

FG 

GU FG


Teliostachya coppenamensis Bremek. [=Lepidagathis sp.] 

Teliostachya lanceolata Nees [=Lepidagathis sp.] 

Teliostachya mazarunensis Bremek. [=Lepidagathis sp.] 

Thunbergia alata Bojer ex Sims [cultivated, naturalized] 

Thunbergia erecta (Benth.) T. Anderson [cultivated, naturalized] 

Thunbergia fragrans Roxb. [cultivated, naturalized] 

Thunbergia grandiflora (Roxb. ex Rottl.) Roxb. [cultivated, naturalized] 

Trichanthera gigantea (Bonpl.) Nees 

Trichanthera gigantea var. guianensis Gleason 


Adhatoda carthaginensis (Jacq.) Nees =Justicia carthaginensis 

Amphiscopia lancifolia var. minor Nees =Justicia moritziana 

Anisacanthus boliviensis (Nees) Wassh. [not in flora area; records referable to A. secundus] 

Aphelandra deppeana Schltdl. & Cham. =Aphelandra scabra 

Aphelandra inserta Leonard =Aphelandra pulcherrima 

Aphelandra paraensis Lindau =Aphelandra aurantiaca 

Aphelandra pectinata Willd. ex Nees =Aphelandra scabra 

Aphelandra tetragona (Vahl) Nees [not in flora area] 

Arrhostoxylum coccineum Nees [not in flora area] 

Arrhostoxylum rubrum (Aubl.) Nees =Ruellia rubra 

Asystasia coromandeliana Nees =Asystasia gangetica 

Asystasia nemorum Nees =Asystasia gangetica 

Barleria pyramidata Lam. =Blechum pyramidatum 

Beloperone calycina Nees =Justicia calycina 

Beloperone modesta Bremek. =Justicia coppenamensis 

Beloperone schomburgkiana Nees =Justicia schomburgkiana 

Blechum brownei J. Juss. =Blechum pyramidatum 

Blechum brownei f. puberulum Leonard =Blechum pyramidatum 

Bravaisia floribunda DC. =Bravaisia integerrima 

Chaetochlamys ciliata Lindau =Justicia sprucei 

Chaetochlamys wurdackii Leonard =Justicia parguazensis 

Dactylostegium assurgens (L.) Bremek. =Dicliptera sexangularis 

Dianthera acuminatissima (Miq.) Benoist =Justicia calycina 

Dianthera calycina (Nees) Benth. ex B.D. Jacks. =Justicia calycina 

Dianthera comata L. =Justicia comata 

Dianthera guianensis N.E. Br. =Justicia guianensis 

Dianthera pectoralis (Jacq.) J.F. Gmel., nom. illeg. =Justicia pectoralis 

Dicliptera assurgens (L.) J. Juss. =Dicliptera sexangularis 

Dicliptera vahliana Nees =Dicliptera sexangularis 

Dipteracanthus angustifolius (Nees) Bremek. =Ruellia geminiflora var. angustifolia 

Dipteracanthus canescens Nees =Ruellia geminiflora var. angustifolia 

Dipteracanthus geminiflorus (Kunth) Nees =Ruellia geminiflora 

Dipteracanthus geminiflorus var. angustifolius Nees =Ruellia geminiflora var. angustifolia 

Dipteracanthus menthoides Nees =Ruellia menthoides 

Dipteracanthus paniculatus Nees ? [not in flora area] =Ruellia sp. 

Dipteracanthus surinamensis Miq. =Ruellia microcalyx 

Dipteracanthus vindex Nees =Ruellia geminiflora var. angustifolia 

Dipteracanthus violaceus (Nees) Lemée =Ruellia violacea 

Drejera boliviensis Nees =Anisacanthus secundus p.p. 

Duvernoia potarensis Bremek. =Justicia potarensis 

Duvernoia potarensis var. tetrapora Bremek. =Justicia potarensis 

Dyspemptemorion cayennense (Nees) Bremek. =Justicia cayennensis 

Ebermaiera repens Nees =Staurogyne repens 

Ebermaiera spp. =Staurogyne spp. 

Eranthemum nervosum (Vahl) R. Br. =Eranthemum pulchellum 

Fittonia sp. [GU record =Justicia sp. nov., fide Wasshausen] 

Hygrophila guianensis Nees ex Benth. =Hygrophila costata 

Jacobinia venezuelica Lindau =Justicia guianensis 

Juruasia rotundata Lindau =Herpetacanthus rotundatus 

Justicia “divergens (Nees) Lindau” [invalid name, records based on misidentifications] 

Justicia acuminatissima (Miq.) Bremek. =Justicia calycina 

Justicia bracteolata Jacq. =Odontonema bracteolatum 

Justicia caripensis Kunth =Justicia secunda 

SU 

SU 

DA 

GU 

GU SU 

DA SU 

SU 

GU SU 

DA GU SU 

GU SU

186 


Justicia coccinea Aubl. =Pachystachys coccinea 

Justicia gangetica L. =Asystasia gangetica 

Justicia glabribracteata (Lindau) V.A.W. Graham [records apparently in error] 

Justicia imbricata Vahl =Elytraria imbricata 

Justicia martiana (Nees) Lindau =Justicia comata 

Justicia moricandiana (Nees) Lemée =Justicia secunda 

Justicia obtusifolia (Nees) Lindau =Justicia laevilinguis 

Justicia pectoralis var. latifolia Bremek. =Justicia pectoralis 

Justicia plectica Leonard =Justicia schomburgkiana 

Justicia pulcherrima Jacq. =Aphelandra pulcherrima 

Justicia scabra Vahl =Aphelandra scabra 

Justicia spicata Ruiz & Pav. =Pachystachys spicata 

Justicia variegata Aubl. =Pulchranthus variegatus 

Leptostachya martiana var. hispida Nees =Justicia comata 

Meyenia erecta Benth. =Thunbergia erecta 

Nelsonia canescens (Lam.) Spreng. =Nelsonia campestris 

Odontonema surinamense Bremek. =Pulchranthus surinamensis 

Odontonema variegatum (Aubl.) Kuntze =Pulchranthus variegatus 

Odontonema venezuelense Leonard =Odontonema bracteolatum 

Onychacanthus arboreus H. Karst. =Bravaisia integerrima 

Pachystachys riedeliana Nees =Pachystachys spicata 

Polylychnis essequibensis Bremek. =Polylychnis radicans 

Psadacocalymma spp. =Justicia spp. 

Rhacodiscus acuminatissimus (Miq.) Lindau =Justicia calycina 

Rhacodiscus calycinus (Nees) Bremek. =Justicia calycina 

Rhacodiscus intermedius (Nees) Lindau =Justicia calycina 

Rhacodiscus moricandianus (Nees) Bremek. =Justicia secunda 

Rhacodiscus secundus (Vahl) Bremek. =Justicia secunda 

Rhytiglossa acuminatissima Miq. =Justicia calycina 

Rhytiglossa laevilinguis Nees =Justicia laevilinguis 

Rhytiglossa moricandiana Nees =Justicia secunda 

Rhytiglossa secunda (Vahl) Nees =Justicia secunda 

Rhytiglossa secunda var. intermedia Nees =Justicia calycina 

Ruellia alopecuroidea Vahl =Lepidagathis alopecuroidea 

Ruellia blechum L. =Blechum pyramidatum 

Ruellia cordifolia Lindau =Ruellia inflata 

Ruellia gigantea Bonpl. =Trichanthera gigantea 

Ruellia pipericaulis Rizzini =Ruellia inflata 

Ruellia radicans (Nees) Lindau =Polylychnis radicans 

Ruellia tubiflora Kunth [not in flora area, based on misidentifications] 

Ruellia vindex (Nees) Lindau =Ruellia geminiflora var. angustifolia 

Sericographis caripensis (Kunth) Nees =Justicia secunda 

Staurogyne leptocaulis Leonard =Staurogyne spraguei 

Staurogyne linearifolia Bremek. =Staurogyne stolonifera 

Staurogyne versteegii Bremek. =Staurogyne trinitensis 

Stemonacanthus macrophyllus (Vahl) Nees =Ruellia macrophylla 

Teliostachya alopecuroidea (Vahl) Nees =Lepidagathis alopecuroidea 

Teliostachya glandulosa Bremek. =Lepidagathis nickeriensis 

Teliostachya gracilis Bremek. =Lepidagathis gracilis 

Teliostachya medicaginea Bremek. =Lepidagathis medicaginea 

Teliostachya surinamensis Bremek. =Lepidagathis surinamensis 

Thyrsacanthus bracteolatus (Jacq.) Nees =Odontonema bracteolatum 

Thyrsacanthus variegatus (Aubl.) Nees =Pulchranthus variegatus 

Ulleria angustifolia (Nees) Bremek. =Ruellia geminiflora var. angustifolia 

REFERENCES 

(see also Mendonciaceae) 

Baum, V.M. & J.L. Reveal. 1983. Pulchranthus, a new genus from northern South America. Syst. Bot. 8(2): 221-220. 

Bremekamp, C.E.B. 1938. Acanthaceae. In A. Pulle, Flora of Suriname 4(2): 166-252. 

Bremekamp, C.E.B. 1938. Notes on the Acanthaceae of Surinam. Rec. Trav. Bot. Neerl. 35: 130-171. 


Bremekamp, C.E.B. 1938. Notes on the Acanthaceae of Surinam. Rec. Trav. Bot. Neerl. 35: 130-171. 

Graham, V.A.W. 1988. Delimitation and infra-generic classification of Justicia. Kew Bull. 43(4): 551-624. 

Leonard, E.C. 1961. Acanthaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. Mem. 

New York Bot. Gard. 10(4): 47-58.


Wasshausen, D.C. 1975. The genus Aphelandra. Smiths. Contrib. Bot. 18: 1-157. 

Wasshausen, D.C. 1990. Justicia maguirei, a new species of Acanthaceae from Guyana. Mem. New York Bot. Gard. 64: 235- 

238. 

Wasshausen, D.C. 1991. New species of Dicliptera from the Guianas. Brittonia 43(1): 1-6. 

Wasshausen, D.C. 1992a. New species and new combinations of Justicia (Acanthaceae) from the Venezuelan Guayana. Novon 

2: 62-80. 

Wasshausen, D.C. 1992b. Nomenclatural changes in the Acanthaceae and two new species of Odontonema from Venezuela and 

Guyana. Novon 2: 149-152. 

Wasshausen, D.C. 1995. Acanthaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 335- 

374. 

Wasshausen, D.C. 1995. A new species, new combinations, and a new name in Acanthaceae from northern South America. 

Brittonia 47(4): 422-425. 

Wasshausen, D.C. 2002. Acanthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 

Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 

76 (2): 31-39. 

Sesuvium portulacastrum (L.) L. 

AIZOACEAE 

R.A. DeFilipps†, J.A. Steyermark† 


Sesuvium acutifolium Miq. =Sesuvium portulacastrum 

REFERENCES(see also Molluginaceae) 

Burger, W. 1983. Aizoaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212. 

Eyma, P.J. 1934. Aizoaceae. In A. Pulle, Flora of Suriname 1(1): 158-161. 

Nevling, L.I. Jr. 1961. Aizoaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. Gard. 

48(1): 80-85. 

AMARANTHACEAE 

R.A. DeFilipps†, M.H. Nee 

Achyranthes indica (L.) Mill. [naturalized] 

Alternanthera brasiliana (L.) Kuntze 

Alternanthera halimifolia (Lam.) Standl. [=Alternanthera tenella?] 

Alternanthera littoralis P. Beauv. 

Alternanthera littoralis var. maritima (Mart.) Pedersen 

Alternanthera paronychioides A. St.-Hil. [specimens misidentified = 

Alternanthera tenella in the Guianas] 

Alternanthera philoxeroides (Mart.) Griseb. 

Alternanthera pulchella Kunth 

Alternanthera sessilis (L.) R. Br. ex DC. 

Alternanthera tenella Colla 

Amaranthus australis (A. Gray) J.D. Sauer [introduced?] 

Amaranthus blitum L. [introduced?] 

Amaranthus caudatus L. [cultivated or escaped from cultivation] 

Amaranthus dubius Mart. ex Thell. 

Amaranthus hybridus L. 

Amaranthus spinosus L. [introduced?] 

Amaranthus tricolor L. [cultivated only?] 

Amaranthus viridis L. [introduced?] 

Blutaparon vermiculare (L.) Mears 

Celosia argentea L. [cultivated, naturalized] 

Celosia virgata Jacq. 

Chamissoa altissima (Jacq.) Kunth 

Chamissoa altissima var. altissima 

Chamissoa altissima var. swansonii Sohmer 

Cyathula achyranthoides (Kunth) Moq. [introduced?] 

Cyathula prostrata (L.) Blume [introduced?] 

Froelichia humboldtiana (Schult.) Seub. 

Gomphrena globosa L. [widely cultivated and escaping] 

Iresine diffusa Humb. & Bonpl. ex Willd. 

Pfaffia glabrata Mart. 

DA GU SU FG 


AM GU SU FG 

GU SU FG 

FG? 

FG? 

BO 

DA GU SU FG 

BO 

BO GU SU FG 

GU SU FG 

DA GU SU FG 


SU FG 


SU FG 

BO GU SU FG 

SU FG 


DA GU SU FG 

AM BO 

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GU SU 

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188 

Pfaffia glabrata var. rostrata Stützer 

Pfaffia glomerata (Spreng.) Pedersen 

Pfaffia grandiflora (Hook.) R.E. Fr. 

Pfaffia iresinoides (Kunth) Spreng. 


FG? 

GU SU FG 

AM BO DA GU 

BO 


Achyranthes altissima Jacq. =Chamissoa altissima var. altissima 

Achyranthes aspera var. indica L. =Achyranthes indica 

Achyranthes brasiliana (L.) Standl. =Alternanthera brasiliana 

Achyranthes corymbosa L. (Caryophyllaceae) =Polycarpaea corymbosa 

Achyranthes halimifolia Lam. =Alternanthera halimifolia 

Achyranthes philoxeroides (Mart.) Standl. = Alternanthera philoxeroides (Mart.) Griseb. 

Achyranthes prostrata L. =Cyathula prostrata 

Achyranthes sessilis (L.) Desf. ex Steud. =Alternanthera sessilis 

Acnida australis A. Gray =Amaranthus australis 

Acnida cuspidata Bertero ex Spreng. =Amaranthus australis 

Alternanthera crucis Bouldingh =Alternanthera halimifolia 

Alternanthera dentata (Moench) Scheyg. =Alternanthera brasiliana 

Alternanthera ficoidea (L.) Sm. =Alternanthera paronychioides 

Alternanthera ficoidea (L.) R. Br. ex Roem. & Schult. [name often applied to A. tenella specimens] =Alternanthera 

paronychioides 

Alternanthera flavogrisea (Urb.) Urb. =Alternanthera halimifolia 

Alternanthera flavogrisea subsp. diffusa (Mart.) Mears ex Fourn. =Alternanthera tenella 

Alternanthera iresinoides Kunth =Pfaffia iresinoides 

Alternanthera maritima (Mart.) A. St.-Hil. =Alternanthera littoralis var. maritima 

Alternanthera ramosissima R.E. Fr. =Alternanthera brasiliana 

Amaranthus gracilis Desf. =Amaranthus viridis 

Amaranthus lividus L. =Amaranthus blitum 

Amaranthus paniculatus L. =Amaranthus hybridus 

Bucholzia philoxeroides Mart. =Alternanthera philoxeroides 

Caraxeron vermicularis (L.) Raf. =Blutaparon vermiculare 

Celosia cristata L. =Celosia argentea 

Celosia paniculata L. =Iresine diffusa 

Desmochaeta achyranthoides Kunth =Cyathula achyranthoides 

Froelichia lanata (Kunth) Moq. =Froelichia humboldtiana 

Gomphrena brasiliana L. =Alternanthera brasiliana 

Gomphrena humboldtiana Schult. =Froelichia humboldtiana 

Gomphrena iresinoides (Kunth) Moq. =Pfaffia iresinoides 

Gomphrena lanata Kunth =Froelichia humboldtiana 

Gomphrena sessilis L. =Alternanthera sessilis 

Gomphrena stenophylla Spreng. =Pfaffia glomerata 

Gomphrena vermicularis L. =Blutaparon vermiculare 

Hebanthe hookeriana Hemsl. =Pfaffia grandiflora 

Illecebrum diffusum Schult. =Alternanthera pulchella 

Iresine celosia L. =Iresine diffusa 

Iresine celosioides L. =Iresine diffusa 

Iresine glomerata Spreng. =Pfaffia glomerata 

Iresine grandiflora Hook. (non R.E. Fr.) =Pfaffia grandiflora 

Iresine polymorpha Mart. =Iresine diffusa 

Iresine vermicularis (L.) Moq. =Blutaparon vermiculare 

Pfaffia glauca (Mart.) Spreng. =Pfaffia glomerata 

Pfaffia hookeriana (Hemsl.) Greenm. =Pfaffia grandiflora 

Pfaffia stenophylla (Spreng.) Stuchlik =Pfaffia glomerata 

Philoxerus vermicularis (L.) R. Br. =Blutaparon vermiculare 

Serturnera glauca Mart. =Pfaffia glomerata 

Telanthera maritima Moq. =Alternanthera littoralis var. maritima 

REFERENCES 

Burger, W. 1983. Amaranthaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 142-180. 

DeFilipps, R.A. and S.L. Maina. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 40-42. 

Duke, J.A. 1961. Amaranthaceae. In R.E. Woodson, R.W. Schery, & collaborators, Flora of Panama 4(4): 348-392. 

Kellogg, E.A. 1988. Amaranthaceae. In R.A. Howard, Flora of the Lesser Antilles.


Mears, J.A. 1977. The nomenclature and type collections of the widespread taxa of Alternanthera. Proc. Acad. Nat. Sci. Phila. 

129: 1-21. 

Morros, M.E., B. Trujillo, & M. Ponce. 1990. Descripcion del genero Amaranthus L. con 3 nuevos registros para Venezuela y 

consiguiente clave para las especies. Ernstia 58-59-60: 45-51. 

Nee, M.H. 1995. Amaranthaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 384-399. 

Pedersen, T.M. 1967. Studies in South American Amaranthaceae. Darwiniana 14(2-3): 430-462. 

Pedersen, T.M. 1976. Estudios sobre Amaranthaceae Sudamericanas, II. Darwiniana 20(1-2): 269-303. 

Scheygrond, A. 1932. Amarantaceae. In A. Pulle, Flora of Suriname 1(1): 25-44. 

Sohmer, S.H. 1977. Revision of Chamissoa. Bull. Torr. Bot. Club 104: 111-126. 

Anacardium amapaense J.D. Mitch. 

Anacardium fruticosum J.D. Mitch. & S.A. Mori 

Anacardium giganteum W. Hancock ex Engl. 

Anacardium occidentale L. 

Anacardium spruceanum Benth. ex Engl. 

Astronium fraxinifolium Schott 

Astronium graveolens Jacq. 

Astronium lecointei Ducke 

Astronium ulei Mattick 

Campnosperma gummiferum (Benth.) Marchand 

Cyrtocarpa velutinifolia (R.S. Cowan) J.D. Mitch. & Daly 

Loxopterygium sagotii Hook. f. 

Mangifera indica L. [widely cultivated] 

Spondias dulcis Parkinson 

Spondias mombin L. 

Spondias purpurea L. 

Tapirira bethanniana J.D. Mitch. 

Tapirira guianensis Aubl. 

Tapirira obtusa (Benth.) J.D. Mitch. 

Thyrsodium guianense Sagot ex Marchand 

Thyrsodium herrerense Encarnación 

Thyrsodium puberulum J.D. Mitch. & Daly 

Thyrsodium spruceanum Benth. 

ANACARDIACEAE 

J.D. Mitchell 


Anacardium microcarpum Ducke =Anacardium occidentale 

Comocladia tapaculo Kunth =Tapirira guianensis 

Cyrtospermum gummiferum Benth. =Campnosperma gummiferum 

Drepanospermum gummiferum (Benth.) Benth. =Campnosperma gummiferum 

Garuga gigantea Engl. =Thyrsodium spruceanum 

Garuga guianensis (Sagot) Engl. =Thyrsodium guianense 

Garuga schomburgkiana (Benth.) Engl. =Thyrsodium spruceanum 

Garuga spruceana (Benth.) Engl. =Thyrsodium spruceanum 

Loxopterygium gutierrezii F.A. Barkley =Cyrtocarpa velutinifolia 

Mauria multiflora Mart. ex Benth. =Tapirira guianensis 

Mauria obtusa Benth. =Tapirira obtusa 

Spondias cytherea Sonn. =Spondias dulcis 

Spondias lutea L. =Spondias mombin 

Tapirira fanshawei Sandwith =Tapirira guianensis 

Tapirira guianensis var. cuneata Engl. =Tapirira guianensis 

Tapirira guianensis var. elliptica Engl. =Tapirira guianensis 

Tapirira marchandii Engl. [nom. illeg.] =Tapirira obtusa 

Tapirira myriantha Triana & Planch. =Tapirira guianensis 

Tapirira pao-pombo var. major Marchand =Tapirira obtusa 

Tapirira peckoltiana Engl. =Tapirira obtusa 

Tapirira velutinifolia (R.S. Cowan) Marc.-Berti =Cyrtocarpa velutinifolia 

Thyrsodium dasytrichum Sandwith =Thyrsodium spruceanum 

Thyrsodium giganteum (Engl.) Engl. =Thyrsodium spruceanum 

Thyrsodium paraense Huber =Thyrsodium spruceanum 

Thyrsodium schomburgkianum Benth. =Thyrsodium spruceanum 

FG 

AM BO 

GU 

GU SU 


AM BO GU SU 

SU 

FG 

DA 

BO SU 

AM 

AM 

BO GU SU FG 

BO GU 



BO DA GU FG 


GU FG 

FG 



BO 

GU? SU FG 

SU FG 

AM BO DA GU SU FG

190 


REFERENCES 

Barkley, F.A. 1968. Anacardiaceae: Rhoideae: Astronium. Phytologia 16: 107-152. 

Jansen-Jacobs, M.J. 1976. Anacardiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 

441-444. 

Mitchell, J.D. 1992. Additions to Anacardium. Anacardium amapaënse, a new species from French Guiana and eastern Amazonian 

Brazil. Brittonia 44(3): 331-338. 

Mitchell, J.D. 1993. Tapirira obtusa comb. nov. (Anacardiaceae). Novon 3(1): 66. 

Mitchell, J.D. 1995. Anacardiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 399-412. 

Mitchell, J.D. 1997. Anacardiaceae. In A.R.A. Görts-van Rijn & M.J. Jansen-Jacobs, Flora of the Guianas (series A) 129: 1-79. 

Mitchell, J.D. 2002. Anacardiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 

eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 

43-53. 

Mitchell, J.D. & D.C. Daly. 1991. Cyrtocarpa Kunth (Anacardiaceae) in South America. Ann. Missouri Bot. Gard. 78: 184-189. 

Mitchell, J.D. & D.C. Daly. 1993. A revision of Thyrsodium. Brittonia 45(2): 115-129. 

Mitchell, J.D. & S. Mori. 1987. The cashew and its relatives (Anacardium: Anacardiaceae). Mem. New York Bot. Gard. 42: 1- 

76. 

Nannenga, E.T. 1934. Anacardiaceae. In A. Pulle, Flora of Suriname 2(1): 132-145. 

Anisophyllea guianensis Sandwith 

ANISOPHYLLEACEAE 

REFERENCES 

Juncosa, A.M. & P.B. Tomlinson. 1988. A historical and taxonomic synopsis of Rhizophoraceae and Anisophylleaceae. Ann. 

Missouri Bot. Gard. 75: 1278-1295. 

Sandwith, N.Y. 1952. Contributions to the flora of tropical America LV. Discovery of Anisophyllea in America. Kew Bull. 1952: 

303-306. 

ANNONACEAE 

P.E. Berry, D.M. Johnson, P.J.M. Maas, N.A. Murray, H. Rainer, J.A. Steyermark† 

Anaxagorea acuminata (Dunal) A. DC. 

Anaxagorea angustifolia Timmerman 

Anaxagorea brachycarpa R.E. Fr. 

Anaxagorea brevipedicellata Timmerman 

Anaxagorea brevipes Benth. 

Anaxagorea dolichocarpa Sprague & Sandwith 

Anaxagorea gigantophylla R.E. Fr. 

Anaxagorea inundata P.E. Berry & R.B. Mill. 

Anaxagorea petiolata R.E. Fr. 

Anaxagorea phaeocarpa Mart. 

Anaxagorea prinoides (Dunal) A. DC. 

Anaxagorea rheophytica Maas & Westra 

Anaxagorea rufa Timmerman 

Annona ambotay Aubl. 

Annona atabapensis Kunth 

Annona densicoma Mart. 

Annona echinata Dunal 

Annona foetida Mart. 

Annona gigantophylla (R.E. Fr.) R.E. Fr. 

Annona glabra L. 

Annona haematantha Miq. 

Annona hypoglauca Mart. 

Annona jahnii Saff. 

Annona montana Macfad. 

Annona muricata L. 

Annona paludosa Aubl. 

Annona prevostiae H. Rainer 

Annona purpurea Moç. & Sessé ex Dunal 

Annona sericea Dunal 

Annona symphyocarpa Sandwith 

Annona tenuiflora Mart. 

Annona tenuipes R.E. Fr. 

GU 

AM GU SU FG 

AM 

AM 

FG 

AM BO GU SU 


AM 

AM 

AM BO GU 

FG 

SU FG 

AM 

AM 


AM 

GU SU FG 

GU SU FG 

SU FG 

AM 

DA GU SU FG 

GU SU FG 

AM GU SU FG 

BO 



GU SU FG 

FG 

BO 


AM BO GU FG 

FG 

GU

Annona trunciflora R.E. Fr. 

Annona ulei R.E. Fr. 

Bocageopsis canescens (Spruce ex Benth.) R.E. Fr. 

Bocageopsis multiflora (Mart.) R.E. Fr. 

Cardiopetalum surinamense R.E. Fr. 

Cremastosperma brevipes (DC. ex Dunal) R.E. Fr. 

Cymbopetalum brasiliense (Vell.) Benth. ex Baill. 

Diclinanona calycina (Diels) R.E. Fr. 

Duguetia aberrans Maas 

Duguetia amplexifolia R.E. Fr. 

Duguetia argentea (R.E. Fr.) R.E. Fr. 

Duguetia cadaverica Huber 

Duguetia calycina Benoist 

Duguetia cauliflora R.E. Fr. 

Duguetia chrysea Maas 

Duguetia decurrens R.E. Fr. 

Duguetia dimorphopetala R.E. Fr. 

Duguetia eximia Diels 

Duguetia flagellaris Huber 

Duguetia granvilleana Maas 

Duguetia guianensis R.E. Fr. 

Duguetia inconspicua Sagot 

Duguetia latifolia R.E. Fr. 

Duguetia lepidota (Miq.) Pulle 

Duguetia lucida Urb. 

Duguetia macrocalyx R.E. Fr. 

Duguetia macrophylla R.E. Fr. 

Duguetia megalophylla R.E. Fr. 

Duguetia neglecta Sandwith 

Duguetia oligocarpa Maas & van Dam 

Duguetia paraensis R.E. Fr. 

Duguetia pauciflora Rusby 

Duguetia pycnastera Sandwith 

Duguetia quitarensis Benth. 

Duguetia riberensis Aristeg. ex Maas & Boon 

Duguetia rigida R.E. Fr. 

Duguetia rionegrensis van Zuilen & Maas 

Duguetia riparia Huber 

Duguetia sancticaroli Maas 

Duguetia schulzii Jans.-Jac. 

Duguetia surinamensis R.E. Fr. 

Duguetia tenuis R.E. Fr. 

Duguetia trunciflora Maas & A.H. Gentry 

Duguetia uniflora (Dunal) Mart. 

Duguetia vallicola J.F. Macbr. 

Duguetia venezuelana R.E. Fr. 

Duguetia yeshidan Sandwith 

Ephedranthus guianensis R.E. Fr. 

Fusaea longifolia (Aubl.) Saff. 

Guatteria atabapensis Aristeg. ex D.M. Johnson & N.A. Murray 

Guatteria atra Sandwith 

Guatteria axilliflora (DC.) R.E. Fr. 

Guatteria brachypoda R.E. Fr. 

Guatteria cardoniana R.E. Fr. [cf. Guatteria recurvisepala] 

Guatteria citriodora Ducke 

Guatteria clusiifolia D.M. Johnson & N.A. Murray 

Guatteria conspicua R.E. Fr. 

Guatteria discolor R.E. Fr. 

Guatteria duckeana var. subcordata R.E. Fr. 

Guatteria dura R.E. Fr. 

Guatteria elongata Benth. [Brazilian species? “Rio Negro”] 

Guatteria flexilis R.E. Fr. 

Guatteria foliosa Benth. 

Guatteria ? gracilipes R.E. Fr. 


AM BO FG 

GU 

AM 


SU FG 

FG 


AM 

AM 

SU FG 

AM 

GU SU FG 

AM GU SU FG 

AM BO GU SU 

GU 

GU 

AM 

SU FG 

AM 

FG 

GU SU 

GU SU FG 

AM GU 

AM BO SU 

AM BO DA GU 

GU 

AM 

BO DA GU 

GU SU 

FG 

GU FG 

BO DA GU 


AM GU 

BO 

AM BO GU 

AM 

SU FG 

AM 

SU 

GU SU FG 

AM GU 

AM FG 

AM 

AM 

AM 


AM GU SU FG 


AM 

GU 

FG 

GU 

AM BO GU 

GU SU FG 

GU 

GU SU FG 

FG 

AM 

AM 

GU? 

BO DA GU 

AM BO FG 

SU

192 


Guatteria guianensis (Aubl.) R.E. Fr. 

Guatteria insculpta R.E. Fr. 

Guatteria ? insignis R.E. Fr. 

Guatteria inundata Mart. 

Guatteria latipetala R.E. Fr. 

Guatteria liesneri D.M. Johnson & N.A. Murray 

Guatteria longicuspis R.E. Fr. 

Guatteria maguirei R.E. Fr. 

Guatteria maypurensis Kunth 

Guatteria megalophylla Diels 

Guatteria monticola R.E. Fr. 

Guatteria oblonga R.E. Fr. 

Guatteria obovata R.E. Fr. [Brazilian species? “Rio Negro, Lagoon Pedrero”] 

Guatteria ouregou (Aubl.) Dunal 

Guatteria ovalifolia R.E. Fr. 

Guatteria paludosa R.E. Fr. 

Guatteria procera R.E. Fr. 

Guatteria ? pteropus Benth. 

Guatteria punctata (Aubl.) R.A. Howard 

Guatteria recurvisepala R.E. Fr. [cf. Guatteria cardoniana] 

Guatteria richardii R.E. Fr. 

Guatteria riparia R.E. Fr. 

Guatteria rubrinervis R.E. Fr. 

Guatteria scandens Ducke 

Guatteria schomburgkiana Mart. 

Guatteria stenopetala R.E. Fr. 

Guatteria subsessilis Mart. 

Guatteria ? trichostemon R.E. Fr. 

Guatteria ? umbonata R.E. Fr. 

Guatteria wachenheimi Benoist 

Guatteria williamsii R.E. Fr. 

Guatteriopsis blepharophylla (Mart.) R.E. Fr. 

Heteropetalum brasiliense Benth. 

Malmea guianensis R.E. Fr. 

Malmea surinamensis Chatrou 

Mosannona discolor (R.E. Fr.) Chatrou 

Oxandra asbeckii (Pulle) R.E. Fr. 

Oxandra espintana (Spruce ex Benth.) Baill. 

Oxandra euneura Diels 

Oxandra guianensis R.E. Fr. 

Oxandra ? krukoffii R.E. Fr. 

Oxandra leucodermis (Spruce ex Benth.) Warm. 

Oxandra surinamensis Jans.-Jac. 

Oxandra xylopioides Diels 

Pseudephedranthus fragrans (R.E. Fr.) Aristeg. 

Pseudoxandra coriacea R.E. Fr. 

Pseudoxandra cuspidata Maas 

Pseudoxandra guianensis (R.E. Fr.) R.E. Fr. 

Pseudoxandra polyphleba (Diels) R.E. Fr. 

Rollinia cuspidata Mart. 

Rollinia elliptica R.E. Fr. 

Rollinia exsucca (DC. ex Dunal) A. DC. 

Rollinia mucosa (Jacq.) Baill. 

Tetrameranthus duckei R.E. Fr. 

Trigynaea caudata (R.E. Fr.) R.E. Fr. 

Trigynaea duckei (R.E. Fr.) R.E. Fr. 

Unonopsis glaucopetala R.E. Fr. 

Unonopsis grandis (Benth.) R.E. Fr. 

Unonopsis guatterioides (A. DC.) R.E. Fr. 

Unonopsis perrottetii (A. DC.) R.E. Fr. 

Unonopsis rufescens (Baill.) R.E. Fr. 

Unonopsis stipitata Diels 

Unonopsis velutina Maas 

Xylopia amazonica R.E. Fr. 

Xylopia aromatica (Lam.) Mart. 

AM 

AM 

AM 

AM BO 

AM 

AM 

AM BO GU 

GU 

GU 

GU? 

SU 

FG 

SU? FG 

FG 

AM BO DA GU 

GU 

GU SU FG 

SU FG 

GU SU FG 

GU FG 

FG 

AM 

BO DA GU 

GU SU FG 


AM 

AM GU 

SU 

GU 

GU SU FG 

AM 

BO SU FG 

AM 

GU 

SU 

GU 

AM GU SU FG 

AM BO 

AM 

GU 

SU 

AM 

SU 

AM GU SU FG 

AM 

AM 

FG 

GU FG 

AM BO GU FG 

AM GU FG 

SU FG 


BO GU SU FG 

AM 

GU SU 

GU 


AM 

BO GU SU FG 

GU FG 

GU SU FG 

AM GU SU FG 

AM BO 

AM BO SU 

AM BO DA GU SU FG

Xylopia benthamii R.E. Fr. 

Xylopia calophylla R.E. Fr. 

Xylopia cayennensis Maas 

Xylopia crinita R.E. Fr. 

Xylopia discreta (L. f.) Sprague & Hutch. 

Xylopia emarginata Mart. 

Xylopia excellens R.E. Fr. 

Xylopia frutescens Aubl. 

Xylopia ligustrifolia Humb. & Bonpl. ex Dunal 

Xylopia nervosa (R.E. Fr.) Maas 

Xylopia nitida Dunal 

Xylopia parviflora Spruce 

Xylopia plowmanii P.E. Berry & D.M. Johnson 

Xylopia pulcherrima Sandwith 

Xylopia sericea A. St.-Hil. 

Xylopia spruceana Benth. ex Spruce 

Xylopia surinamensis R.E. Fr. 

Xylopia venezuelana R.E. Fr. 



Aberemoa argentea R.E. Fr. =Duguetia argentea 

Aberemoa guianensis Aubl. =Guatteria guianensis 

Aberemoa longifolia (Aubl.) Baill. =Fusaea longifolia 

Anaxagorea mutica R.E. Fr. =Anaxagorea dolichocarpa 

Annona axilliflora DC. =Guatteria axilliflora 

Annona exsucca DC. ex Dunal =Rollinia exsucca 

Annona guaricensis Pittier =Annona jahnii 

Annona hostmannii Steud. =Guatteria schomburgkiana 

Annona jenmanii Saff. =Annona sericea 

Annona lepidota Miq. =Duguetia lepidota 

Annona longifolia Aubl. =Fusaea longifolia 

Annona manirote Kunth =Annona purpurea 

Annona mucosa Jacq. =Rollinia mucosa 

Annona palustris L. =Annona glabra 

Annona punctata Aubl. =Guatteria punctata 

Annona rhombipetala Ruiz & Pav. ex G. Don =Fusaea longifolia 

Annona uniflora Dunal =Duguetia uniflora 

Bocagea asbeckii Pulle =Oxandra asbeckii 

Bocagea espintana Spruce ex Benth. =Oxandra espintana 

Bocagea leucodermis Spruce ex Benth. =Oxandra leucodermis 

Bocagea multiflora Mart. =Bocageopsis multiflora 

Cananga ouregou Aubl. =Guatteria ouregou 

Cremastosperma guianense R.E. Fr. =Pseudoxandra guianensis 

Cremastosperma poiteaui (Diels) R.E. Fr. =Cremastosperma brevipes 

Cremastosperma polyphlebum (Diels) R.E. Fr. =Pseudoxandra polyphleba 

Cymbopetalum venosum Gleason (Lecythidaceae) =Lecythis corrugata subsp. corrugata 

Diclinanona calycina var. macrophylla R.E. Fr. =Diclinanona calycina 

Duguetia adiscandra Jans.-Jac. =Duguetia cadaverica 

Duguetia cuspidata R.E. Fr. =Duguetia calycina 

Duguetia elegans R.E. Fr. =Duguetia calycina 

Duguetia friesii Jans.-Jac. =Duguetia cadaverica 

Duguetia insculpta R.E. Fr. =Duguetia riparia 

Duguetia longifolia (Aubl.) Baill. =Fusaea longifolia 

Duguetia obovata R.E. Fr. =Duguetia yeshidan 

Duguetia rostrata Rusby =Duguetia pauciflora 

Duguetia sandwithii R.E. Fr. =Duguetia yeshidan 

Ephedranthus fragrans R.E. Fr. =Pseudephedranthus fragrans 

Froesiodendron surinamense (R.E. Fr.) R.E. Fr. =Cardiopetalum surinamense 

Fusaea decurrens R.E. Fr. =Fusaea longifolia 

Guatteria bernardii R.E. Fr. =Guatteria schomburgkiana 

Guatteria blepharophylla Mart. =Guatteriopsis blepharophylla 

Guatteria calva R.E. Fr. =Guatteria maypurensis 

Guatteria chrysopetala (Steud.) Miq. =Guatteria punctata 

Guatteria ferruginea A. St.-Hil. [not in flora area] 

Guatteria flavovirens R.E. Fr. =Guatteria schomburgkiana 


AM BO GU 

GU SU FG 

AM FG 


AM 

AM FG 


AM 

FG 


AM FG 

AM 

GU SU FG 


AM 

GU SU FG 

AM BO

194 


Guatteria heteropetala Benth. =Heteropetalum brasiliense 

Guatteria maypurensis var. attenuata R.E. Fr. =Guatteria maypurensis 

Guatteria maypurensis var. pulchra R.E. Fr. =Guatteria maypurensis 

Guatteria microsperma R.E. Fr. =Guatteria wachenheimi 

Guatteria ouregou Griseb., non Dunal [not in flora area] =Guatteria inuncta 

Guatteria ouregou Mart., non Dunal =Guatteria pteropus 

Guatteria podocarpa DC. ex Dunal =Guatteria ouregou 

Guatteria sagotiana R.E. Fr. =Guatteria punctata 

Guatteria sandwithii R.E. Fr. =Guatteria schomburgkiana 

Guatteria sessilis R.E. Fr. =Guatteria schomburgkiana 

Guatteria spruceana R.E. Fr. =Guatteria schomburgkiana 

Guatteria velezii R.E. Fr. =Guatteria maypurensis 

Guatteria wessels-boerii Jans.-Jac. =Guatteria punctata 

Heteropetalum spruceanum R.E. Fr. =Heteropetalum brasiliense 

Heteropetalum spruceanum var. longipetalum R.E. Fr. =Heteropetalum brasiliense 

Hornschuchia caudata R.E. Fr. =Trigynaea caudata 

Hornschuchia duckei R.E. Fr. =Trigynaea duckei 

Hornschuchia guianensis R.E. Fr. =Trigynaea duckei 

Malmea discolor R.E. Fr. =Mosannona discolor 

Oxandra nervosa R.E. Fr. =Xylopia nervosa 

Pseudoxandra lucida R.E. Fr. =Pseudoxandra polyphleba 

Raimondia tenuiflora (Mart.) R.E. Fr. =Annona tenuiflora 

Rollinia broadwayi var. cuneata R.E. Fr. =Rollinia exsucca 

Rollinia cardiantha Diels =Rollinia cuspidata 

Rollinia multiflora Splitg. =Rollinia exsucca 

Rollinia puberula A. DC. =Rollinia exsucca 

Rollinia pulchrinervia A. DC. =Rollinia mucosa 

Rollinia sieberi A. DC. =Rollinia mucosa 

Rollinia surinamensis R.E. Fr. =Rollinia exsucca 

Trigynaea anastomosans Rusby =Cymbopetalum brasiliense 

Trigynaea canescens Spruce ex Benth. =Bocageopsis canescens 

Trigynaea grandis Benth. =Unonopsis grandis 

Trigynaea guianensis (R.E. Fr.) R.E. Fr. =Trigynaea duckei 

Unona crassipetala Dunal =Guatteria ouregou 

Unona discreta L. f. =Xylopia discreta 

Unonopsis polyphleba Diels =Pseudoxandra polyphleba 

Uva brasiliensis (Vell.) Kuntze =Cymbopetalum brasiliense 

Uvaria aromatica Lam. =Xylopia aromatica 

Uvaria brasiliensis Vell. =Cymbopetalum brasiliense 

Uvaria guatterioides A. DC. =Unonopsis guatterioides 

Uvaria spectabilis DC. =Fusaea longifolia 

Xylopia acuminata Dunal =Anaxagorea acuminata 

Xylopia calycina Diels =Diclinanona calycina 

Xylopia cinerea Sandwith =Xylopia nitida 

Xylopia gigantophylla R.E. Fr. =Annona gigantophylla 

Xylopia grandiflora A. St.-Hil. =Xylopia aromatica 

Xylopia longifolia (Sagot) R.E. Fr. =Xylopia cayennensis 

Xylopia longifolia A. DC. =Xylopia aromatica 

Xylopia neglecta (Kuntze) R.E. Fr. =Xylopia parviflora 

Xylopia nitida var. longifolia Sagot =Xylopia cayennensis 

REFERENCES 

Chatrou, L.W. & P. He. 1999. Studies in Annonaceae XXXIII. A revision of Fusaea. (Baill.) Saff. Brittonia 51(2): 181-203. 

Fries, R.E. 1930-1939. Revision der Arten einiger Annonaceen-Gattungen. Acta Horti Berg. 10(1): 1-128 (1930), 10(2): 129-341 

(1931), 12(1): 1-220 (1934), 12(2): 221-288 (1937), 12(3): 289-577 (1939). 

Fries, R.E. 1940. Annonaceae. In A. Pulle, Flora of Suriname 2(2): 341-383. 

Jansen-Jacobs, M.J. 1976. Annonaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 658- 

687. 

Johnson, D.M. & N.A. Murray. 1995. Synopsis of the tribe Bocageeae (Annonaceae), with revisions of Cardiopetalum, 

Froesiodendron, Trigynaea, Bocagea, and Hornschuchia. Brittonia 47(3): 248-319. 

Maas, P.J.M. 1992. Rollinia. Flora Neotropica Monogr. 57: 1-192. 

Maas, P.J.M. 1996. Studies in Annonaceae XXX. Monograph of Duguetia: Preliminary notes. Bot. Jahrb. 118(2): 187-227. 

Maas, P.J.M., J. Koek-Noorman, & L.Y.Th. Westra. 1993. Studies in Annonaceae XVIII. New species from the Neotropics and 

miscellaneous notes. Bot. Jahrb. 115(1): 77-95.


Maas, P.J.M. & H Maas-van de Kamer. 2002. Annonaceae.In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. 

Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New 

York Bot. Gard. 76 (2): 53 - 67. 

Maas, P.J.M., E.A. Mennega, & L.Y.Th. Westra. 1994. Studies in Annonaceae XXI. Index to species and infraspecific taxa of 

neotropical Annonaceae. Candollea 49(2): 389-481. 

Maas, P.J.M. & L.Y.Th. Westra. 1984-1985. Studies in Annonaceae II. A monograph of the genus Anaxagorea A. St. Hil. Part 1. 

Bot. Jahrb. Syst. 105: 73-134. Part 2. Bot. Jahrb. Syst. 105: 145-204. 

Steyermark, J.A., P.J.M. Maas, P.E. Berry, D.M. Johnson, N.A. Murray, & H. Rainer. 1995. Annonaceae. In J.A. Steyermark, 

P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 413-469. 

Zuilen, C.M. van & P.J.M. Maas. 1994. Studies in Annonaceae XXII. A taxonomic revision of Duguetia A. St.-Hil., sections 

Alcmene and Xylopipetalum. Bot. Jahrb. 116(2): 221-242. 

Eryngium ebracteatum Lam. 

Eryngium foetidum L. 

Hydrocotyle rotundifolia Roxb. [introduced, no recent records] 

Hydrocotyle umbellata L. 

APIACEAE 

P.E. Berry, J.C. Lindeman 

REFERENCES 

Berry, P.E. 1995. Apiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 470-471. 

Mathias, M.E. 1936. The genus Hydrocotyle in northern South America. Brittonia 2: 201-237. 

Mori, S.A. 2002. Apiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 

2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 67-68. 

Stafleu, F.A. 1951. Umbelliferae. In A. Pulle, Flora of Suriname 3(2): 252-256. 

APOCYNACEAE 

L. Allorge, M.E. Endress, B.F. Hansen, A.J.M. Leeuwenberg, G.N. Morillo, J.L. Zarucchi 

Allamanda cathartica L. 

Allamanda doniana Müll. Arg. 

Allamanda oenotherifolia Pohl 

Allamanda schottii Pohl 

Allamanda setulosa Miq. 

Allamanda thevetiifolia Müll. Arg. 

Ambelania acida Aubl. 

Aspidosperma album (Vahl) Benoist ex Pichon 

Aspidosperma araracanga Marc.-Ferr. 

Aspidosperma carapanauba Pichon 

Aspidosperma cruentum Woodson 

Aspidosperma cuspa (Kunth) S.F. Blake ex Pittier 

Aspidosperma decussatum Woodson 

Aspidosperma discolor A. DC. 

Aspidosperma excelsum Benth. 

Aspidosperma glaucum Monach. 

Aspidosperma helstonei Donsel. 

Aspidosperma leucocymosum Kuhlm. 

Aspidosperma macrophyllum Müll. Arg. 

Aspidosperma macrophyllum subsp. macrophyllum 

Aspidosperma macrophyllum subsp. morii L. Allorge 

Aspidosperma marcgravianum Woodson 

Aspidosperma neblinae Monach. 

Aspidosperma oblongum A. DC. 

Aspidosperma pachypterum Müll. Arg. 

Aspidosperma pichonianum Woodson 

Aspidosperma sandwithianum Markgr. 

Aspidosperma schultesii Woodson 

Aspidosperma spruceanum Benth. ex Müll. Arg. 

Aspidosperma steyermarkii Woodson 

Aspidosperma ulei Markgr. 

Aspidosperma vargasii A. DC. 

Aspidosperma verruculosum Müll. Arg. 

BO 


GU? 



AM BO 

AM 

FG 

GU SU 

AM 

BO GU SU FG 


AM 

FG 


AM BO 

BO 

BO SU 


AM 

SU FG 

AM 

GU FG 

GU 

FG 


AM 

BO GU SU FG 

AM 

BO 

GU SU FG 

FG 

GU SU FG 

BO 

AM BO GU SU 

BO GU SU 

AM

196 

Catharanthus roseus (L.) G. Don [widely cultivated, escaping] 

Condylocarpon amazonicum (Markgr.) Ducke 

Condylocarpon guyanense Desf. 

Condylocarpon intermedium Müll. Arg. 

Condylocarpon intermedium var. intermedium 

Condylocarpon myrtifolium (Miq.) Müll. Arg. 

Condylocarpon pubiflorum Müll. Arg. 

Couma catingae Ducke 

Couma guianensis Aubl. 

Couma macrocarpa Barb. Rodr. 

Couma rigida Müll. Arg. 

Couma utilis (Mart.) Müll. Arg. 

Forsteronia acouci (Aubl.) A. DC. 

Forsteronia adenobasis Müll. Arg. 

Forsteronia affinis Müll. Arg. 

Forsteronia diospyrifolia Müll. Arg. 

Forsteronia duckei Markgr. 

Forsteronia gracilis (Benth.) Müll. Arg. 

Forsteronia guyanensis Müll. Arg. 

Forsteronia laurifolia (Benth.) A. DC. 

Forsteronia lucida Markgr. 

Forsteronia obtusiloba Müll. Arg. 

Forsteronia paraensis Hansen 

Forsteronia schomburgkii A. DC. 

Forsteronia umbellata (Aubl.) Woodson 

Galactophora calycina (Huber ex Ducke) Woodson 

Galactophora colellana Morillo 

Galactophora crassifolia (Müll. Arg.) Woodson 

Galactophora pulchella Woodson 

Galactophora pumila Monach. 

Galactophora schomburgkiana Woodson 

Galactophora schomburgkiana var. megaphylla Monach. 

Galactophora schomburgkiana var. schomburgkiana 

Geissospermum argenteum Woodson 

Geissospermum laeve (Thunb.) Miers 

Geissospermum reticulatum A.H. Gentry 

Geissospermum sericeum Benth. & Hook. f. ex Miers 

Himatanthus articulatus (Vahl) Woodson 

Himatanthus attenuatus (Benth.) Woodson 

Himatanthus attenuatus var. attenuatus 

Himatanthus attenuatus var. obtusifolius (Müll. Arg.) Plumel 

Himatanthus bracteatus (A. DC.) Woodson 

Himatanthus bracteatus var. bracteatus 

Himatanthus bracteatus var. revolutus (Huber) Plumel 

Himatanthus drasticus (Mart.) Plumel 

Himatanthus fallax (Müll. Arg.) Plumel 

Himatanthus obovatus (Müll. Arg.) Woodson 

Himatanthus phagedaenicus (Mart.) Woodson 

Himatanthus semilunatus Markgr. 

Himatanthus speciosus (Müll. Arg.) Plumel 

Himatanthus stenophyllus Plumel 

Himatanthus sucuuba (Spruce ex Müll. Arg.) Woodson 

Hylaea arborescens (Monach.) J.F. Morales 

Kopsia arborea Blume 

Lacmellea aculeata (Ducke) Monach. 

Lacmellea arborescens (Müll. Arg.) Markgr. 

Lacmellea floribunda (Poepp.) Benth. & Hook. f. 

Lacmellea guyanensis (Müll. Arg.) Monach. 

Lacmellea microcarpa (Müll. Arg.) Markgr. 

Lacmellea pygmaea Monach. 

Lacmellea pygmaea var. latifolia Monach. 

Lacmellea pygmaea var. pygmaea 

Lacmellea ramosissima (Müll. Arg.) Markgr. 

Lacmellea utilis (Arn.) Markgr. 

Laxoplumeria baehniana Monach. 



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Macoubea guianensis Aubl. 

Macoubea sprucei (Müll. Arg.) Markgr. 

Macropharynx spectabilis (Stadelm.) Woodson 

Macropharynx strigillosa Woodson 

Malouetia calva Markgr. 

Malouetia duckei Markgr. [expected on or near Sierra de la Neblina] 

Malouetia flavescens (Willd. ex Roem. & Schult.) Müll. Arg. 

Malouetia glandulifera Miers 

Malouetia gracilis (Benth.) A. DC. 

Malouetia grandiflora Woodson 

Malouetia molongo M.E. Endress 

Malouetia naias M.E. Endress 

Malouetia parvifolia Woodson 

Malouetia pubescens Markgr. 

Malouetia pubescens var. glabra M.E. Endress 

Malouetia tamaquarina (Aubl.) A. DC. 

Malouetia virescens Spruce ex Müll. Arg. 

Mandevilla anceps Woodson 

Mandevilla annulariifolia Woodson 

Mandevilla aracamunensis Morillo 

Mandevilla benthamii (A. DC.) K. Schum. 

Mandevilla caurensis Markgr. 

Mandevilla duidae (Woodson) Woodson 

Mandevilla filifolia Monach. 

Mandevilla hirsuta (Rich.) K. Schum. 

Mandevilla holstii Morillo 

Mandevilla huberi Morillo 

Mandevilla javitensis (Kunth) K. Schum. 

Mandevilla lancifolia Woodson 

Mandevilla lasiocarpa (A. DC.) Malme 

Mandevilla leptophylla (A. DC.) K. Schum. 

Mandevilla obtusifolia Monach. 

Mandevilla pachyphylla Woodson 

Mandevilla rugellosa (Rich.) L. Allorge 

Mandevilla rugosa (Benth.) Woodson 

Mandevilla scaberula N.E. Br. 

Mandevilla scabra (Hoffmanns. ex Roem. & Schult.) K. Schum. 

Mandevilla spruceana (Müll. Arg.) K. Schum. 

Mandevilla steyermarkii Woodson 

Mandevilla subcarnosa (Benth.) Woodson 

Mandevilla subcarnosa var. angustata Steyerm. 

Mandevilla subcarnosa var. subcarnosa 

Mandevilla subspicata (Vahl) Markgr. 

Mandevilla surinamensis (Pulle) Woodson 

Mandevilla symphitocarpa (G. Mey.) Woodson 

Mandevilla tenuifolia (Mik.) Woodson 

Mandevilla trianae Woodson 

Mandevilla turgida Woodson 

Mandevilla vanheurckii (Müll. Arg.) Markgr. 

Mesechites acutisepala Monach. 

Mesechites trifida (Jacq.) Müll. Arg. 

Microplumeria anomala (Müll. Arg.) Markgr. 

Molongum laxum (Benth.) Pichon 

Molongum lucidum (Kunth) Zarucchi 

Mucoa pantchenkoana (Markgr.) Zarucchi 

Neocouma parviflora (Markgr.) Zarucchi 

Neocouma ternstroemiacea (Müll. Arg.) Pierre 

Nerium oleander L. [cultivated, perhaps escaping] 

Odontadenia funigera Woodson 

Odontadenia geminata (Hoffmanns. ex Roem. & Schult.) Müll. Arg. 

Odontadenia glauca Woodson 

Odontadenia macrantha (Roem. & Schult.) Markgr. 

Odontadenia nitida (Vahl) Müll. Arg. 

Odontadenia perrottetii (A. DC.) Woodson 

Odontadenia puncticulosa (Rich.) Pulle 

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198 


Odontadenia sandwithiana Woodson 

Odontadenia verrucosa (Willd. ex Roem. & Schult.) K. Schum. ex Markgr. 

Pacouria guianensis Aubl. 

Parahancornia fasciculata (Poir.) Benoist ex Pichon 

Parahancornia negroensis Monach. 

Parahancornia oblonga (Benth. ex Müll. Arg.) Monach. 

Parahancornia surrogata Zarucchi 

Plumeria inodora Jacq. 

Plumeria rubra L. [cultivated & escaped] 

Prestonia acutifolia (Benth. ex Müll. Arg.) K. Schum. 

Prestonia annularis (L. f.) G. Don 

Prestonia cayennensis (A. DC.) Pichon 

Prestonia coalita (Vell.) Woodson 

Prestonia exserta (A. DC.) Standl. 

Prestonia finitima Woodson 

Prestonia guianensis Gleason 

Prestonia ipomaeifolia A. DC. 

Prestonia latifolia Benth. 

Prestonia lindleyana Woodson ex Gleason & A.C. Sm. 

Prestonia megagros (Vell.) Woodson 

Prestonia perplexa Woodson 

Prestonia quinquangularis (Jacq.) Spreng. 

Prestonia surinamensis Müll. Arg. 

Prestonia tomentosa R. Br. 

Prestonia vaupesiana Woodson 

Rauvolfia ligustrina Willd. ex Roem. & Schult. 

Rauvolfia pachyphylla Markgr. 

Rauvolfia paraensis Ducke 

Rauvolfia polyphylla Benth. 

Rauvolfia tetraphylla L. 

Rhabdadenia biflora (Jacq.) Müll. Arg. 

Rhabdadenia macrostoma (Benth.) Müll. Arg. 

Rhabdadenia pohlii Müll. Arg. 

Rhigospira quadrangularis (Müll. Arg.) Miers 

Salpinctes kalmiifolius Woodson 

Secondatia densiflora A. DC. 

Spongiosperma cataractarum Zarucchi 

Spongiosperma macrophyllum (Müll. Arg.) Zarucchi 

Spongiosperma oleifolium (Monach.) Zarucchi 

Spongiosperma riparium (Monach.) Zarucchi 

Stemmadenia grandiflora (Jacq.) Miers 

Tabernaemontana albiflora (Miq.) Pulle [=Bonafousia albiflora] 

Tabernaemontana angulata Mart. ex Müll. Arg. [=Bonafousia angulata] 

Tabernaemontana attenuata (Miers) Urb. [=Anartia meyeri] 

Tabernaemontana cerea (Woodson) Leeuwenb. [=Anartia cerea] 

Tabernaemontana cymosa Jacq. [=Peschiera cymosa] 

Tabernaemontana disticha A. DC. [=Bonafousia disticha] 

Tabernaemontana divaricata (L.) R. Br. ex Roem. & Schult. [cultivated or 

escaped from cultivation] 

Tabernaemontana flavicans Willd. ex Roem. & Schult. [=Anartia flavicans] 

Tabernaemontana heterophylla Vahl [=Stenosolen heterophyllus] 

Tabernaemontana lagenaria Leeuwenb. 

Tabernaemontana lorifera (Miers) Leeuwenb. [=Bonafousia lorifera] 

Tabernaemontana macrocalyx Müll. Arg. [=Bonafousia macrocalyx, 

Bonafousia muelleriana] 

Tabernaemontana maxima Markgr. 

Tabernaemontana palustris Markgr. 

Tabernaemontana pandacaqui Lam. [Asian species, cultivated only?] 

Tabernaemontana rupicola Benth. [=Bonafousia rupicola] 

Tabernaemontana sananho Ruiz & Pav. [=Bonafousia sananho] 

Tabernaemontana siphilitica (L. f.) Leeuwenb. [=Bonafousia siphilitica] 

Tabernaemontana undulata Vahl [=Bonafousia undulata] 

Thevetia ahouai A. DC. 

Thevetia peruviana (Pers.) K. Schum. 

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Ahouai nitida (Kunth) Pichon =Thevetia ahouai 

Ambelania cucumerina Miers =Ambelania acida 

Ambelania cuneata Müll. Arg. =Molongum lucidum 

Ambelania lanceolata Markgr. =Spongiosperma oleifolium 

Ambelania laxa (Benth.) Müll. Arg. =Molongum laxum 

Ambelania lopezii Woodson =Spongiosperma macrophyllum 

Ambelania lucida (Kunth) Markgr. =Molongum lucidum 

Ambelania macrophylla Müll. Arg. =Spongiosperma macrophyllum 

Ambelania oleifolia Monach. =Spongiosperma oleifolium 

Ambelania oleifolia var. riparia Monach. =Spongiosperma riparium 

Ambelania pantchenkoana Markgr. =Mucoa pantchenkoana 

Ambelania parviflora Markgr. =Neocouma parviflora 

Ambelania quadrangularis Müll. Arg. =Rhigospira quadrangularis 

Ambelania sagotii Müll. Arg. =Ambelania acida 

Ambelania tenuiflora Müll. Arg. =Ambelania acida 

Ambelania tenuiflora var. tenuiramea Müll. Arg. =Ambelania acida 

Ambelania ternstroemiacea (Müll. Arg.) Monach. =Neocouma ternstroemiacea 

Amblyanthera crassifolia Müll. Arg. =Galactophora crassifolia 

Amblyanthera spruceana Müll. Arg. =Mandevilla spruceana 

Anacampta albescens (Rusby) Markgr. =Tabernaemontana undulata 

Anacampta disticha (A. DC.) Markgr. =Tabernaemontana disticha 

Anacampta echinata (Aubl.) Markgr. [see Tabernaemontana echinata, Peschiera echinata] 

Anacampta macrocalyx (Müll. Arg.) Markgr. =Tabernaemontana macrocalyx 

Anacampta maxima (Markgr.) Markgr. =Tabernaemontana maxima 

Anacampta muelleriana (Mart. ex Müll. Arg.) Markgr. =Tabernaemontana macrocalyx 

Anacampta palustris (Markgr.) Markgr. =Tabernaemontana palustris 

Anacampta rigida (Miers) Markgr. [not in flora area] =Tabernaemontana muricata Link ex Roem. & Schult. 

Anacampta rupicola (Benth.) Markgr. =Tabernaemontana rupicola 

Anartia attenuata (Miers) Markgr. =Tabernaemontana attenuata 

Anartia cerea (Woodson) L. Allorge =Tabernaemontana cerea 

Anartia flavicans (Willd. ex Roem. & Schult.) Miers =Tabernaemontana flavicans 

Anartia meyeri (G. Mey. ex G. Don) Miers =Tabernaemontana attenuata 

Anartia oblongifolia (A. DC.) Markgr. =Tabernaemontana flavicans 

Anartia olivacea (Müll. Arg.) Markgr. =Tabernaemontana flavicans 

Anechites amazonica Markgr. =Condylocarpon amazonicum 

Anechites nerium Urb. [not in flora area] 

Angadenia geminata (Hoffmanns. ex Roem. & Schult.) Miers =Odontadenia geminata 

Angadenia nitida (Vahl) Miers =Odontadenia nitida 

Anisolobus cognatus (Stadelm.) Müll. Arg. =Odontadenia verrucosa 

Anisolobus puncticulosus (Rich.) Miq. =Odontadenia puncticulosa 

Apocynum acouci Aubl. =Forsteronia acouci 

Apocynum cumanense Willd. ex Schult. (Asclepiadaceae) =Matelea cumanensis 

Apocynum paniculatum Lam. =Forsteronia acouci 

Apocynum umbellatum Aubl. =Forsteronia umbellata 

Aspidosperma anomalum Müll. Arg. =Microplumeria anomala 

Aspidosperma decipiens Müll. Arg. =Aspidosperma cuspa 

Aspidosperma decussatum subsp. jauaensis Steyerm. & Maguire =Aspidosperma decussatum 

Aspidosperma desmanthum Benth. ex Müll. Arg. =Aspidosperma album 

Aspidosperma exalatum Monach. =Aspidosperma pachypterum 

Aspidosperma lucentivenium S.F. Blake =Aspidosperma cuspa 

Aspidosperma megalocarpon Müll. Arg. [not in flora area] 

Aspidosperma nitidum Benth. ex Müll. Arg. [not in flora area] 

Aspidosperma quadriovulatum Pittier =Rauvolfia pachyphylla 

Aspidosperma rhombeosignatum Markgr. =Aspidosperma pachypterum 

Aspidosperma verruculosum var. laeve Monach. =Aspidosperma verruculosum 

Aspidosperma woodsonianum Markgr. =Aspidosperma spruceanum 

Bonafousia albiflora (Miq.) Boiteau & L. Allorge =Tabernaemontana albiflora 

Bonafousia angulata (Mart. ex Müll. Arg.) Boiteau & L. Allorge =Tabernaemontana angulata 

Bonafousia attenuata Miers =Tabernaemontana attenuata 

Bonafousia disticha (A. DC.) Boiteau & L. Allorge =Tabernaemontana disticha 

Bonafousia lorifera (Miers) Boiteau & L. Allorge =Tabernaemontana lorifera 

Bonafousia macrocalyx (Müll. Arg.) Boiteau & L. Allorge =Tabernaemontana macrocalyx 

Bonafousia maxima (Markgr.) Boiteau & L. Allorge =Tabernaemontana maxima

200 


Bonafousia morettii L. Allorge =Tabernaemontana macrocalyx 

Bonafousia muelleriana (Mart. ex Müll. Arg.) Boiteau & L. Allorge =Tabernaemontana macrocalyx 

Bonafousia obliqua Miers =Tabernaemontana undulata 

Bonafousia oblongifolia (A. DC.) Miers =Tabernaemontana flavicans 

Bonafousia palustris (Markgr.) Boiteau & L. Allorge =Tabernaemontana palustris 

Bonafousia rimulosa (Woodson ex R.E. Schult.) Boiteau & L. Allorge =Tabernaemontana palustris 

Bonafousia rupicola (Benth.) Miers =Tabernaemontana rupicola 

Bonafousia rupicola var. versicolor (Woodson) L. Allorge =Tabernaemontana rupicola 

Bonafousia sananho (Ruiz & Pav.) Markgr. =Tabernaemontana sananho 

Bonafousia siphilitica (L. f.) L. Allorge =Tabernaemontana siphilitica 

Bonafousia speciosa (Poir.) Boiteau =Tabernaemontana siphilitica 

Bonafousia tetrastachya (Kunth) Markgr. =Tabernaemontana siphilitica 

Bonafousia undulata (Vahl) A. DC. =Tabernaemontana undulata 

Cameraria tamaquarina Aubl. =Malouetia tamaquarina 

Cerbera ahouai L. =Thevetia ahouai 

Cerbera nitida Kunth =Thevetia ahouai 

Cerbera peruviana Pers. =Thevetia peruviana 

Codonemma macrocalyx (Müll. Arg.) Miers =Tabernaemontana macrocalyx 

Collophora utilis Mart. =Couma utilis 

Condylocarpon breviarticulatum Müll. Arg. =Condylocarpon intermedium var. intermedium 

Condylocarpon ciliatum Müll. Arg. =Condylocarpon pubiflorum 

Condylocarpon hirtellum Ducke =Condylocarpon pubiflorum 

Condylocarpon intermedium var. brevifolium Müll. Arg. =Condylocarpon intermedium var. intermedium 

Condylocarpon longii Standl. & L.O. Williams =Condylocarpon intermedium var. intermedium 

Condylocarpon obtusiusculum Müll. Arg. =Condylocarpon intermedium var. intermedium 

Condylocarpon occidentale Markgr. =Condylocarpon amazonicum 

Condylocarpon rauwolfiae Müll. Arg. [not in flora area] =Condylocarpon isthmicum (Vell.) A. DC. 

Condylocarpon reticulatum Ducke =Condylocarpon amazonicum 

Conoria cuspa Kunth =Aspidosperma cuspa 

Couma capiron Pittier =Couma macrocarpa 

Couma caurensis Pittier =Couma macrocarpa 

Couma fasciculata (Poir.) Benoist =Parahancornia fasciculata 

Couma multinervis Monach. =Couma utilis 

Couma oblonga Benth. ex Müll. Arg. =Parahancornia oblonga 

Couma sapida Pittier =Couma macrocarpa 

Cylindrosperma anomalum (Müll. Arg.) Ducke =Microplumeria anomala 

Dipladenia calycina Huber ex Ducke =Galactophora calycina 

Dipladenia surinamensis Pulle =Mandevilla surinamensis 

Dipladenia upatae Woodson =Mandevilla surinamensis 

Echites angustifolia Benth. =Mandevilla benthamii 

Echites benthamii A. DC. =Mandevilla benthamii 

Echites biflora Jacq. =Rhabdadenia biflora 

Echites brachystachya Benth. =Mandevilla scabra 

Echites cognata Stadelm. =Odontadenia verrucosa 

Echites elegans Benth. =Odontadenia geminata 

Echites geminata Hoffmanns. ex Roem. & Schult. =Odontadenia geminata 

Echites grandiflora G. Mey. =Odontadenia macrantha 

Echites hirsuta Rich. =Mandevilla hirsuta 

Echites hoffmannseggiana Steud. =Odontadenia macrantha 

Echites javitensis Kunth =Mandevilla javitensis 

Echites laurentiae-disca Rusby =Prestonia acutifolia 

Echites leptophylla A. DC. =Mandevilla leptophylla 

Echites macrantha Roem. & Schult. =Odontadenia macrantha 

Echites macrostoma Benth. =Rhabdadenia macrostoma 

Echites megagros Vell. =Prestonia megagros 

Echites nitida Vahl =Odontadenia nitida 

Echites puncticulosa Rich. =Odontadenia puncticulosa 

Echites quinquangularis Jacq. =Prestonia quinquangularis 

Echites rugellosa Rich. =Mandevilla rugellosa 

Echites scabra Hoffmanns. ex Roem. & Schult. =Mandevilla scabra 

Echites siphilitica L. f. =Tabernaemontana siphilitica 

Echites spectabilis Stadelm. =Macropharynx spectabilis 

Echites subcarnosa Benth. =Mandevilla subcarnosa 

Echites subspicata Vahl =Mandevilla subspicata


Echites symphytocarpa G. Mey. =Mandevilla symphitocarpa 

Echites trifida Jacq. =Mesechites trifida 

Echites tubulosa Benth. =Mesechites trifida 

Echites verrucosa Roem. & Schult. =Odontadenia verrucosa 

Ervatamia coronaria (Jacq.) Stapf =Tabernaemontana divaricata 

Ervatamia cumingiana (A. DC.) Markgr. =Tabernaemontana pandacaqui 

Exothostemon javitense (Kunth) G. Don =Mandevilla javitensis 

Forsteronia benthamiana Müll. Arg. =Forsteronia acouci 

Forsteronia corymbifera (Miers) Sandwith =Forsteronia acouci 

Forsteronia mediocris Woodson =Forsteronia affinis 

Forsteronia schomburgkii (Benth.) Müll. Arg. =Forsteronia acouci 

Forsteronia viridescens S.F. Blake =Forsteronia acouci 

Haemadictyon acutifolium Benth. ex Müll. Arg. =Prestonia acutifolia 

Haemadictyon cayennense A. DC. =Prestonia cayennensis 

Haemadictyon exsertum A. DC. =Prestonia exserta 

Haemadictyon marginatum Benth. =Prestonia cayennensis 

Hancornia amapa Huber =Parahancornia fasciculata 

Hancornia laxa (Benth.) A. DC. =Molongum laxum 

Himatanthus rigida Willd. ex Roem. & Schult. =Himatanthus articulatus 

Hortsmania myrtifolia Miq. =Condylocarpon myrtifolium 

Lacmellea lucida (Kunth) Miers =Molongum lucidum 

Lacmellea ramosissima (Müll. Arg.) Monach. =Lacmellea ramosissima 

Lacmellea utilis (Arn.) Monach. =Lacmellea utilis 

Landolphia guianensis (Aubl.) Pulle =Pacouria guianensis 

Lochnera rosea (L.) Rchb. =Catharanthus roseus 

Macaglia alba Vahl =Aspidosperma album 

Macaglia excelsa (Benth.) Kuntze =Aspidosperma excelsum 

Macaglia oblonga (A. DC.) Kuntze =Aspidosperma oblongum 

Macaglia verruculosa (Müll. Arg.) Kuntze =Aspidosperma verruculosum 

Macoubea fasciculata (Poir.) Lemée =Parahancornia fasciculata 

Macoubea guianensis var. reticulata (A. DC.) L. Allorge =Macoubea guianensis 

Macoubea paucifolia (Spruce ex Müll. Arg.) Markgr. ex Ll. Williams =Macoubea guianensis 

Macoubea reticulata (A. DC.) Markgr. =Macoubea guianensis 

Macoubea sinuosa (Miers) Markgr. =Macoubea guianensis 

Macoubea sprucei var. paucifolia (Spruce ex Müll. Arg.) Monach. =Macoubea guianensis 

Malouetia furfuracea Spruce ex Müll. Arg. =Malouetia tamaquarina 

Malouetia guianensis (Aubl.) Miers =Malouetia tamaquarina 

Malouetia lactiflua Miers =Parahancornia fasciculata 

Malouetia mexiae Woodson =Malouetia flavescens 

Malouetia obtusiloba A. DC. =Malouetia tamaquarina 

Malouetia odorata DC. =Malouetia tamaquarina 

Malouetia retroflexa Müll. Arg. [not in flora area] =Malouetia cubana A. DC. 

Malouetia schomburgkii Müll. Arg. =Malouetia flavescens 

Malouetia tetrastachya (Kunth) Miers =Tabernaemontana siphilitica 

Mandevilla angustissima Markgr. =Mandevilla lancifolia 

Mandevilla dielsiana Markgr. =Mandevilla subcarnosa var. subcarnosa 

Mandevilla glabra N.E. Br. =Mandevilla vanheurckii 

Mandevilla lancifolia var. calva Monach. =Mandevilla lancifolia 

Mandevilla linearis N.E. Br. =Mandevilla leptophylla 

Merizadenia amplifolia Miers =Macoubea guianensis 

Merizadenia sananho (Ruiz & Pav.) Miers =Tabernaemontana sananho 

Mesechites syphilitica (L. f.) Lemée =Tabernaemontana siphilitica 

Microplumeria anomala var. parvifolia Monach. =Microplumeria anomala 

Microplumeria sprucei Baill. =Microplumeria anomala 

Mitozus leptophyllus (A. DC.) Miers =Mandevilla leptophylla 

Molongum cuneatum (Müll. Arg.) Pichon =Molongum lucidum 

Molongum macrophyllum (Müll. Arg.) Pichon =Spongiosperma macrophyllum 

Odontadenia “schomburgkiana (Nees) Kr.” [invalid name, error for Odontonema schomburgkiana (Acanthaceae)?] 

Odontadenia amazonica (Stadelm.) Malme =Odontadenia verrucosa 

Odontadenia cognata (Stadelm.) Woodson =Odontadenia verrucosa 

Odontadenia cururu (Mart.) K. Schum. =Odontadenia puncticulosa 

Odontadenia grandiflora (G. Mey.) Kuntze =Odontadenia macrantha 

Odontadenia hoffmannseggiana (Steud.) Woodson =Odontadenia macrantha 

Odontadenia kochii Pilg. =Odontadenia puncticulosa

202 


Odontadenia neglecta Woodson =Odontadenia verrucosa 

Odontadenia silvestris (A. DC.) Müll. Arg. =Odontadenia macrantha 

Odontadenia speciosa Benth. =Odontadenia macrantha 

Odontadenia spoliata Malme =Odontadenia puncticulosa 

Odontadenia surinamensis (Pulle) Woodson =Odontadenia verrucosa 

Orelia grandiflora Aubl. =Allamanda cathartica 

Parahancornia amapa (Huber) Ducke =Parahancornia fasciculata 

Parahancornia tabernaemontana Woodson =Macoubea guianensis 

Peschiera cymosa (Jacq.) Dugand =Tabernaemontana cymosa 

Peschiera echinata (Aubl.) A. DC. [name misapplied to Tabernaemontana lagenaria] 

Peschiera heterophylla (Vahl) Miers =Tabernaemontana heterophylla 

Peschiera psychotriifolia (Kunth) Miers =Tabernaemontana cymosa 

Peschiera umbrosa (Kunth) Miers =Tabernaemontana cymosa 

Plumeria alba var. inodora (Jacq.) G. Don =Plumeria inodora 

Plumeria articulata Vahl =Himatanthus articulatus 

Plumeria attenuata Benth. =Himatanthus attenuatus 

Plumeria attenuata var. obtusifolia Müll. Arg. =Himatanthus attenuatus var. obtusifolius 

Plumeria bracteata A. DC. =Himatanthus bracteatus 

Plumeria mollis Kunth =Plumeria rubra 

Plumeria phagedaenica Mart. =Himatanthus phagedaenicus 

Plumeria sucuuba Spruce ex Müll. Arg. =Himatanthus sucuuba 

Plumeriopsis ahouai (L.) Rusby & Woodson =Thevetia ahouai 

Prestonia arborescens Monach. =Hylaea arborescens 

Prestonia marginata (Benth.) Woodson =Prestonia cayennensis 

Quadricasaea inaequilateralis Woodson =Tabernaemontana macrocalyx 

Rauvolfia hirsuta Jacq. =Rauvolfia tetraphylla 

Rauvolfia lamarckii DC. =Rauvolfia tetraphylla 

Rauvolfia parvifolia Bertero ex Spreng. =Rauvolfia ligustrina 

Rauvolfia ternifolia Kunth =Rauvolfia ligustrina 

Rhigospira paucifolia (Spruce ex Müll. Arg.) Miers =Macoubea guianensis 

Rhigospira sinuosa Miers =Macoubea guianensis 

Rhigospira sprucei (Müll. Arg.) Miers =Macoubea sprucei 

Rhigospira ternstroemiacea (Müll. Arg.) Miers =Neocouma ternstroemiacea 

Rhigospira venulosa Miers =Spongiosperma macrophyllum 

Rhipidia amazonica (Markgr.) Markgr. =Condylocarpon amazonicum 

Salpinctes duidae Woodson =Mandevilla duidae 

Stemmadenia cerea Woodson =Tabernaemontana cerea 

Stenosolen heterophyllus (Vahl) Markgr. =Tabernaemontana heterophylla 

Stenosolen holothuria Markgr. =Tabernaemontana heterophylla 

Taberna albiflora (Miq.) Markgr. =Tabernaemontana albiflora 

Taberna cymosa (Jacq.) Miers =Tabernaemontana cymosa 

Taberna disticha Miers =Tabernaemontana disticha 

Tabernaemontana albescens Rusby =Tabernaemontana undulata 

Tabernaemontana aubletii Pulle, nom. illeg. =Macoubea guianensis 

Tabernaemontana citrifolia Müll. Arg. [not in flora area] 

Tabernaemontana coronaria (Jacq.) Willd. =Tabernaemontana divaricata 

Tabernaemontana echinata Vell., non Aubl. [not in flora area] =Tabernaemontana hystrix Steud. 

Tabernaemontana fasciculata Poir. =Parahancornia fasciculata 

Tabernaemontana flavescens Willd. ex Roem. & Schult. =Malouetia flavescens 

Tabernaemontana gracilis Benth. =Malouetia gracilis 

Tabernaemontana grandiflora Jacq. =Stemmadenia grandiflora 

Tabernaemontana holothuria (Markgr.) Leeuwenb. =Tabernaemontana heterophylla 

Tabernaemontana inaequilateralis (Woodson) Pichon =Tabernaemontana macrocalyx 

Tabernaemontana laurifolia L. [not in flora area] 

Tabernaemontana laxa Benth. =Molongum laxum 

Tabernaemontana longifolia Benth. =Tabernaemontana siphilitica 

Tabernaemontana lucida Kunth =Molongum lucidum 

Tabernaemontana macrophylla Müll. Arg. [not in flora area] =Tabernaemontana muricata Link ex Roem. & Schult. 

Tabernaemontana meyeri G. Don =Tabernaemontana attenuata 

Tabernaemontana muelleriana Mart. ex Müll. Arg. =Tabernaemontana macrocalyx 

Tabernaemontana obliqua (Miers) Leeuwenb. =Tabernaemontana undulata 

Tabernaemontana oblongifolia A. DC. =Tabernaemontana flavicans 

Tabernaemontana olivacea Müll. Arg. =Tabernaemontana flavicans 

Tabernaemontana paucifolia Spruce ex Müll. Arg. =Macoubea guianensis 

Tabernaemontana psychotriifolia Kunth =Tabernaemontana cymosa


Tabernaemontana repanda E. Mey. =Tabernaemontana siphilitica 

Tabernaemontana rimulosa Woodson ex R.E. Schult. =Tabernaemontana palustris 

Tabernaemontana sessilifolia Klotzsch =Tabernaemontana rupicola 

Tabernaemontana speciosa Poir. =Tabernaemontana siphilitica 

Tabernaemontana sprucei Müll. Arg. =Macoubea sprucei 

Tabernaemontana submollis Mart. ex Müll. Arg. [not in flora area] =Tabernaemontana coriacea Link ex Roem. & Schult. 

Tabernaemontana tenuis Monach. =Tabernaemontana palustris 

Tabernaemontana ternstroemiacea Müll. Arg. =Neocouma ternstroemiacea 

Tabernaemontana tetrastachya Kunth =Tabernaemontana siphilitica 

Tabernaemontana umbrosa Kunth =Tabernaemontana cymosa 

Tabernaemontana utilis Arn. =Lacmellea utilis 

Tabernaemontana versicolor Woodson =Tabernaemontana rupicola 

Thenardia laurifolia Benth. =Forsteronia laurifolia 

Thenardia umbellata (Aubl.) Spreng. =Forsteronia umbellata 

Thyrsanthus acouci (Aubl.) Miers =Forsteronia acouci 

Thyrsanthus adenobasis (Müll. Arg.) Miers =Forsteronia adenobasis 

Thyrsanthus affinis (Müll. Arg.) Miers =Forsteronia affinis 

Thyrsanthus benthamianus (Müll. Arg.) Miers =Forsteronia acouci 

Thyrsanthus diospyrifolius (Müll. Arg.) Miers =Forsteronia diospyrifolia 

Thyrsanthus fasciculatus (Poir.) Miers =Parahancornia fasciculata 

Thyrsanthus gracilis Benth. =Forsteronia gracilis 

Thyrsanthus guyanensis (Müll. Arg.) Miers =Forsteronia guyanensis 

Thyrsanthus laurifolius (Benth.) Miers =Forsteronia laurifolia 

Thyrsanthus schomburgkii Benth. =Forsteronia acouci 

Vinca rosea L. =Catharanthus roseus 

Willughbeja acida (Aubl.) J.F. Gmel. =Ambelania acida 

Willughbeja guianensis (Aubl.) J.F. Gmel. =Pacouria guianensis 

Willughbeja scandans Willd. =Pacouria guianensis 

Zschokkea aculeata Ducke =Lacmellea aculeata 

Zschokkea microcarpa Müll. Arg. =Lacmellea microcarpa 

Zschokkea ramosissima Müll. Arg. =Lacmellea ramosissima 

Zschokkea utilis (Arn.) Hemsl. =Lacmellea utilis 

REFERENCES 

Allorge, L. 1985. Monographie des Apocynacées - Tabernaemontanoidées Américaines. Mem. Mus. Nat. Hist. Naturelle (n.s.) 

Serie B, tome 30. 

Allorge, L. 1985. Distribution, relations et intérêt biogéographique des Tabernaemontanoideae (Apocynaceae) dans la région 

néotropicale. C.R. Soc. Biogéogr. 61(4): 137-155. 

Allorge, L. 2002. Apocynaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


69-84. 

Allorge, L. & C. Poupat. 1991. Position systématique et révision du genre Aspidosperma (Apocynaceae) pour les trois Guyanes. 

Le point sur leur étude chimique. Bull. Soc. bot. Fr., Lettres bot. (4/5): 267-301. 

Allorge, L. & C. Sastre. 1990. Distribution géographique dans les Guyanes, d’Apocynaceae et d’Ochnaceae. Essai de mise en 

évidence de centres de spéciation. C.R. Soc. Biogéogr. 67(2): 77-94. 

Leeuwenberg, A.J.M. 1988. Series of revisions of Apocynaceae. XXI. Notes on Tabernaemontaneae. Agric. Univ. Wageningen 

Papers 87-5: 1-32. 

Leeuwenberg, A.J.M. 1994. A Revision of Tabernaemontana. Two. The New World Species and Stemmadenia. Royal Botanic 

Gardens, Kew, Richmond, Surrey, England. 

Markgraf, F. 1932. Apocynaceae. In A. Pulle, Flora of Suriname 4(1): 1-65. 

Markgraf, F. 1937. Apocynaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 443-467. 

Monachino, J.V. 1943 (1944). A revision of Couma and Parahancornia. Lloydia 6(4): 229-247. 

Monachino, J. 1961. Apocynaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. 

Mem. New York Bot. Gard. 10(4): 59-65. 

Plumel, M.-M. 1991. Le genre Himatanthus, révision taxonomique. Bradea 5 suppl.: 1-118. 

Woodson, R.E., Jr. 1933-1936. Studies in the Apocynaceae. IV. The American genera of Echitoideae. Ann. Missouri Bot. Gard. 

20: 605-790, 22: 153-306, 23: 169-439. 

Zarucchi, J.L., G.N. Morillo, M.E. Endress, B.F. Hansen & A.J.M. Leeuwenberg. 1995. Apocynaceae. In J.A. Steyermark, P.E. 

Berry, & B.K. Holst, Flora of the Venezuelan Guayana 2: 471-571. 

Ilex abscondita Steyerm. 

Ilex acutidenticulata Steyerm. 

AQUIFOLIACEAE 

P.E. Berry, P.-A. Loizeau, J.A. Steyermark† 

AM 

AM

204 

Ilex altiplana Steyerm. 

Ilex amazonensis Edwin 

Ilex apicidens N.E. Br. 

Ilex aracamuniana Steyerm. 

Ilex archeri Edwin 

Ilex atabapoensis T.R. Dudley 

Ilex attenuata Steyerm. [near Pico Neblina, Brazil] 

Ilex brevipedicellata Steyerm. 

Ilex cardonae Steyerm. [near Sierra Tapirapecó, Brazil] 

Ilex casiquiarensis Loes. 

Ilex ciliolata Steyerm. 

Ilex costata Edwin 

Ilex cowanii Wurdack 

Ilex culmenicola Steyerm. 

Ilex daphnogenea Reissek 

Ilex davidsei Steyerm. 

Ilex diospyroides Reissek 

Ilex divaricata Mart. ex Reissek 

Ilex duidae Gleason 

Ilex fanshawei Edwin 

Ilex glabella Steyerm. 

Ilex gleasoniana Steyerm. 

Ilex gransabanensis Steyerm. 

Ilex guaiquinimae Steyerm. 

Ilex guianensis (Aubl.) Kuntze 

Ilex holstii Steyerm. 

Ilex huachamacariana Edwin 

Ilex ignicola Steyerm. 

Ilex inundata Poepp. ex Reissek 

Ilex jauaensis Steyerm. 

Ilex jenmanii Loes. 

Ilex karuaiana Steyerm. 

Ilex lasseri Edwin 

Ilex laureola Triana 

Ilex liesneri Steyerm. 

Ilex longipilosa Steyerm. 

Ilex macarenensis Cuatrec. 

Ilex magnifructa Edwin 

Ilex maguirei Wurdack 

Ilex marahuacae Steyerm. 

Ilex marginata Edwin 

Ilex martiniana D. Don 

Ilex neblinensis Edwin 

Ilex oliveriana Loes. 

Ilex ovalifolia G. Mey. 

Ilex paruensis Steyerm. 

Ilex parvifructa Edwin 

Ilex paujiensis Steyerm. 

Ilex polita Steyerm. 

Ilex psammophila Reissek 

Ilex ptariana Steyerm. 

Ilex retusa Klotzsch ex Reissek 

Ilex savannarum Wurdack 

Ilex savannarum var. morichei Edwin 

Ilex savannarum var. savannarum 

Ilex sessilifructa Edwin 

Ilex sipapoana Edwin 

Ilex soderstromii Edwin 

Ilex soderstromii var. ovata Edwin 

Ilex soderstromii var. soderstromii 

Ilex solida Edwin 

Ilex spruceana Reissek 

Ilex steyermarkii Edwin 

Ilex subrotundifolia Steyerm. 


BO GU 

AM GU? 

BO GU 

AM 

AM 

AM 

AM FG 

AM 

AM FG 

BO 

GU 

AM 

AM 

GU FG 

AM FG 

AM BO 

AM BO 

AM 

GU 

AM 

AM 

AM BO 

BO 


AM 

AM BO 

AM BO 

GU SU FG? 

BO 

BO GU SU 

BO 

AM BO 

AM BO GU 

AM 

BO 

AM 

AM 

AM BO 

AM 

BO GU 

DA GU SU FG 

AM 

BO GU 

GU SU FG 

AM 

AM BO DA 

BO 

BO 

AM BO 

BO 

AM BO GU 

AM 

AM 

AM 

AM BO 

AM BO 

GU 

GU 

GU 

BO 

AM 

BO 

AM BO


Ilex sulcata Edwin [?later homonym of Ilex sulcata Wall., nomen nudum] 

Ilex summa Steyerm. 

Ilex tateana Steyerm. 

Ilex tepuiana Steyerm. ex Edwin 

Ilex theezans Mart. 

Ilex theezans var. riedelii Loes. 

Ilex thyrsiflora Klotzsch ex Reissek 

Ilex thyrsiflora var. schomburgkii (Klotzsch ex Reissek) Loes. 

Ilex tiricae Edwin 

Ilex uaramae Edwin 

Ilex umbellata Klotzsch ex Reissek 

Ilex vacciniifolia Klotzsch ex Reissek 

Ilex venezuelensis Steyerm. 

Ilex vismiifolia Reissek 

Ilex wurdackiana Steyerm. 

Ilex yutajensis Wurdack 


Ilex acuminata Willd. =Ilex guianensis 

Ilex bolivarensis Edwin =Ilex vacciniifolia Klotzsch ex Reissek 

Ilex brevipetiolata Steyerm. & Wurdack =Ilex subrotundifolia 

Ilex celastroides Klotzsch ex Garcke =Ilex guianensis 

Ilex congesta Reissek (Myrsinaceae) =Cybianthus lineatus 

Ilex cumanensis Turcz. =Ilex guianensis 

Ilex humirioides Reissek =Ilex umbellata 

Ilex lanceolata Klotzsch [nomen nudum] =Ilex martiniana 

Ilex laureola var. neglecta Loes. =Ilex laureola 

Ilex laurina Klotzsch [nomen nudum] =Ilex daphnogenea 

Ilex lucumifolia Reissek ex Maxim. (Sapotaceae) =Pouteria lucumifolia 

Ilex lymanii Edwin =Ilex sipapoana 

Ilex macoucoua Pers. =Ilex guianensis 

Ilex magnifructa var. minor Steyerm. =Ilex magnifructa 

Ilex neblinensis var. wurdackii Edwin =Ilex neblinensis 

Ilex retusa Klotzsch [in R.M. Schomb., nom. inval.] =Ilex retusa Klotzsch ex Reissek 

Ilex retusa f. brunnescens Loes. =Ilex retusa 

Ilex retusa f. genuina Loes. =Ilex retusa 

Ilex retusa f. glabra Steyerm. =Ilex retusa 

Ilex retusa f. glaucescens Steyerm. =Ilex retusa 

Ilex retusa f. major Steyerm. =Ilex retusa 

Ilex riparia Reissek =Ilex inundata 

Ilex schomburgkii Klotzsch ex Reissek =Ilex thyrsiflora var. schomburgkii 

Ilex spathulata Steyerm., non Koidz 1925 =Ilex atabapoensis 

Ilex spruceana var. genuina Loes. =Ilex spruceana 

Ilex spruceana var. guainiensis Loes. =Ilex spruceana 

Ilex stenophylla Steyerm. =Ilex retusa 

Ilex sulcata var. pygmaea Edwin =Ilex sulcata 

Ilex vacciniifolia Klotzsch =Ilex vacciniifolia Klotzsch ex Reissek 

Macoucoua guianensis Aubl. =Ilex guianensis 

REFERENCES 

Amshoff, G.J.H. 1951. Aquifoliaceae. In A. Pulle, Flora of Suriname 3(2): 159-165. 

Edwin, G. 1965. Aquifoliaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part VI. Mem. New York Bot. 

Gard. 12(3): 1-285. 

Steyermark, J.A. & P.E. Berry. 1995. Aquifoliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 2: 571-599. 

Wurdack, J.J. 1961. Aquifoliaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. 


Dendropanax arboreus (L.) Decne. & Planch. 

Dendropanax neblinae Maguire, Steyerm. & Frodin 

Oreopanax capitatus (Jacq.) Decne. & Planch. 

ARALIACEAE 

D.G. Frodin, J.C. Lindeman 

AM 

BO 

BO 

BO GU 

BO GU 

BO GU 

BO GU 

BO GU 

BO 

BO 

BO GU 

BO GU 

AM BO 

GU 

BO 

AM 

AM BO DA GU 

AM 

AM BO DA GU FG

206 


Oreopanax capitatus var. capitatus 

Schefflera acaropunctata Frodin 

Schefflera argophylla Frodin 

Schefflera asymmetrica Frodin 

Schefflera auyantepuiensis (Steyerm.) Maguire, Steyerm. & Frodin 

Schefflera ayangannensis Maguire, Steyerm. & Frodin 

Schefflera baculosa Frodin 

Schefflera brachypodia Frodin 

Schefflera chimantensis (Steyerm. & Maguire) Maguire, Steyerm. & Frodin 

Schefflera chimantensis subsp. chimantensis 

Schefflera chimantensis var. iluensis Frodin 

Schefflera chimantensis subsp. multiramosa (Steyerm.) Frodin 

Schefflera chimantensis var. rugosifolia (Maguire, Steyerm. & Frodin) Frodin 

[Mt. Roraima] 

Schefflera chimantensis subsp. rugosifolia (Maguire, Steyerm. & Frodin) 

Frodin 

Schefflera clausa Frodin 

Schefflera clavigera Frodin 

Schefflera clusietorum Frodin 

Schefflera concolor Frodin 

Schefflera contracta Frodin 

Schefflera coriacea (Marchal) Harms 

Schefflera cracens Frodin 

Schefflera crassilimba Frodin 

Schefflera decaphylla (Seem.) Harms 

Schefflera disparifolia Frodin 

Schefflera dissidens Frodin [Mt. Roraima] 

Schefflera duidae Steyerm. [expected in GU] 

Schefflera eximia Frodin [Sierra de la Neblina] 

Schefflera gracillima Steyerm. & Maguire 

Schefflera guanayensis Maguire, Steyerm. & Frodin 

Schefflera hitchcockii (Lasser & Maguire) Maguire, Steyerm. & Frodin 

Schefflera huachamacarii Maguire, Steyerm. & Frodin 

Schefflera huberi Frodin 

Schefflera japurensis (Mart. & Zucc. ex Marchal) Harms 

Schefflera jauaensis (Maguire, Steyerm. & Frodin) Frodin 

Schefflera longistyla Frodin 

Schefflera maguireanorum Frodin 

Schefflera marahuacensis (Maguire, Steyerm. & Frodin) Frodin 

Schefflera meiurophylla Frodin 

Schefflera monosperma Maguire, Steyerm. & Frodin [Mt. Roraima] 

Schefflera montana (Gleason) Maguire, Steyerm. & Frodin 

Schefflera morototoni (Aubl.) Maguire, Steyerm. & Frodin 

Schefflera myrioneura Frodin 

Schefflera neblinae (Maguire, Steyerm. & Frodin) Frodin [Sierra de la 

Neblina] 

Schefflera nigrescens Frodin 

Schefflera pallens Maguire, Steyerm. & Frodin 

Schefflera paruana (Maguire, Steyerm. & Frodin) Frodin 

Schefflera pauciradiata Maguire, Steyerm. & Frodin 

Schefflera pedicelligera Maguire, Steyerm. & Frodin 

Schefflera pimichinensis Maguire, Steyerm. & Frodin 

Schefflera plurispicata Maguire, Steyerm. & Frodin [expected in AM] 

Schefflera psilophylla (Harms) Maguire, Steyerm. & Frodin [Mt. Roraima] 

Schefflera quinquecarinata Steyerm. 

Schefflera quinquestylorum Steyerm. 

Schefflera sessiliflora Splithof-Heerschop ex Frodin 

Schefflera simplex Steyerm. & B. Holst 

Schefflera sipapoensis Maguire, Steyerm. & Frodin 

Schefflera spruceana (Seem.) Maguire, Steyerm. & Frodin 

Schefflera steyermarkii Frodin 

Schefflera suaveolens Frodin 

Schefflera tamatamaensis Maguire, Steyerm. & Frodin 

Schefflera tremuloidea Maguire, Steyerm. & Frodin 


AM 

AM 

AM 

BO 

GU 

AM 

BO 

BO GU 

BO 

BO 

BO 

BO GU 

AM 

AM 

AM 

BO GU 

BO 

BO 

BO 

BO GU 

BO 

GU SU FG 

BO 

BO? GU 

AM BO 

AM 

BO 

AM 

AM 

AM 

AM 

AM BO 

BO 

AM 

AM 

AM GU? 

AM 

BO GU 

AM BO 


AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO GU 

BO GU 

AM BO GU 

AM 

AM 

AM 

BO 

AM 

AM 

AM

Schefflera ulocephala Frodin 

Schefflera umbellata (N.E. Br.) R. Vig. [Mt. Roraima] 

Schefflera umbraculifera Frodin 

Schefflera varisiana Frodin [expected in AM] 

Schefflera yutajensis Steyerm. & B. Holst 

Sciadodendron excelsum Griseb. 



Aralia arborea L. =Dendropanax arboreus 

Aralia capitata Jacq. =Oreopanax capitatus 

Crepinella gracilis Marchal ex Thurn =Schefflera monosperma 

Didymopanax auyantepuiensis Steyerm. =Schefflera auyantepuiensis 

Didymopanax chimantensis Steyerm. & Maguire =Schefflera chimantensis 

Didymopanax hitchcockii Lasser & Maguire =Schefflera hitchcockii 

Didymopanax montanum Gleason =Schefflera montana 

Didymopanax morototoni (Aubl.) Decne. & Planch. =Schefflera morototoni 

Didymopanax multiramosus Steyerm. =Schefflera chimantensis subsp. multiramosa 

Didymopanax psilophyllus Harms =Schefflera psilophylla 

Didymopanax reticulatus Gleason =Schefflera sessiliflora 

Didymopanax rugosum N.E. Br. =Schefflera chimantensis subsp. rugosifolia 

Didymopanax spruceanus Seem. =Schefflera spruceana 

Didymopanax spruceanus var. cuyabensis Hoehne =Schefflera spruceana 

Gilibertia arborea (L.) Marchal =Dendropanax arboreus 

Oreopanax capitatus var. multiflorus (DC.) Marchal [Guianan material agrees with var. capitatus fide Lindeman] 

Panax morototoni Aubl. =Schefflera morototoni 

Panax splendens Kunth =Schefflera morototoni 

Panax undulata Aubl. =Schefflera morototoni 

Schefflera arborea (L.) M. Gómez =Dendropanax arboreus 

Schefflera coriacea subsp. neblinae Maguire, Steyerm. & Frodin =Schefflera neblinae 

Schefflera duidae subsp. quinquestylorum (Steyerm.) Maguire, Steyerm. & Frodin =Schefflera quinquestylorum 

Schefflera globulifera Maguire, Steyerm. & Frodin =Schefflera ulocephala 

Schefflera guanayensis subsp. paruensis Maguire, Steyerm. & Frodin =Schefflera paruana 

Schefflera guanayensis subsp. sipapoensis Maguire, Steyerm. & Frodin =Schefflera crassilimba 

Schefflera paraensis Huber ex Ducke =Schefflera decaphylla 

Schefflera reticulata (Gleason) Maguire, Steyerm. & Frodin, non Philipson =Schefflera sessiliflora 

Schefflera reticulata subsp. jauaensis Maguire, Steyerm. & Frodin =Schefflera jauaensis 

Schefflera reticulata subsp. yutajensis Maguire, Steyerm. & Frodin =Schefflera maguireanorum 

Schefflera rugosifolia Maguire, Steyerm. & Frodin =Schefflera chimantensis subsp. rugosifolia 

Schefflera splendens (Kunth) Frodin ex Lindeman =Schefflera morototoni 

Schefflera tremuloides var. marahuacensis Maguire, Steyerm. & Frodin =Schefflera marahuacensis 

Schefflera umbellata subsp. duidana Maguire, Steyerm. & Frodin =Schefflera acaropunctata 

Sciadophyllum coriaceum Marchal =Schefflera coriacea 

Sciadophyllum japurense Mart. & Zucc. ex Marchal =Schefflera japurensis 

Sciadophyllum umbellatum N.E. Br. =Schefflera umbellata 

REFERENCES 

Frodin, D.G. 1993. Studies in Schefflera, VI. New species and subordinate taxa in the Venezuelan Guayana and immediately 

adjacent areas. Novon 3: 367-403. 

Frodin, D.G. 1997. Araliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 1-31. 

Maguire, B., J.A. Steyermark, & D.G. Frodin. 1984. Araliaceae. In B. Maguire & collaborators, Botany of the Guayana Highland 

Part XII. Mem. New York Bot. Gard. 38: 1-84. 

Mori, S.A. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide 

to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 84-87. 

Muller, F.M. 1934. Araliaceae. In A. Pulle, Flora of Suriname 3(1): 160-163. 

ARISTOLOCHIACEAE 

K.A. Barringer, C. Feuillet, F.A. González G., O. Poncy 

Aristolochia acutifolia Duch. 

Aristolochia amara (Aubl.) Poncy [taxon based on sterile material, of 

uncertain application] 

Aristolochia anguicida Jacq. 

Aristolochia bukuti Poncy 

Aristolochia chamissonis Duch. 

AM 

AM 

AM 

AM BO 

AM BO 

AM 

BO GU 

BO GU 

FG 

SU FG

208 

Aristolochia consimilis Mast. 

Aristolochia cornuta Mast. 

Aristolochia cremersii Poncy 

Aristolochia daemoninoxia Mast. 

Aristolochia didyma S. Moore 

Aristolochia disticha Mast. 

Aristolochia flava Poncy 

Aristolochia guianensis Poncy 

Aristolochia guntheri O.C. Schmidt 

Aristolochia hians Willd. 

Aristolochia iquitensis O.C. Schmidt 

Aristolochia kanukuensis Feuillt, ined. 

Aristolochia leprieurii Duch. 

Aristolochia maxima Jacq. 

Aristolochia melgueiroi Barringer & Guánchez 

Aristolochia mossii S. Moore 

Aristolochia nummularifolia Kunth 

Aristolochia pannosoides Hoehne 

Aristolochia paramaribensis Duch. 

Aristolochia ringens Vahl 

Aristolochia rugosa Lam. 

Aristolochia silvatica Barb. Rodr. 

Aristolochia sprucei Mast. 

Aristolochia stahelii O.C. Schmidt 

Aristolochia surinamensis Willd. 

Aristolochia trilobata L. 

Aristolochia weddellii Duch. 

Aristolochia weddellii var. rondoniana Hoehne 



Aristolochia amesiana R.E. Schult. =Aristolochia silvatica 

Aristolochia barbata Jacq. =Aristolochia rugosa 

Aristolochia dictyantha var. schomburgkii (Klotzsch) Duch. =Aristolochia rugosa 

Aristolochia dubia Hoehne =Aristolochia mossii 

Aristolochia gabrielis Briq. =Aristolochia leprieurii 

Aristolochia glaucescens Kunth [not in flora area] 

Aristolochia longecaudata Mast. =Aristolochia weddellii var. rondoniana 

Aristolochia macrophylla Duch. =Aristolochia iquitensis 

Aristolochia macrota Duch. =Aristolochia surinamensis 

Aristolochia platyloba Garcke =Aristolochia surinamensis 

REFERENCES 

Barringer, K.A., & G.A. González G. 1997. Aristolochiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 3: 122-129. 

Feuillet, C. & O. Poncy. 1998. Aristolochiaceae. In A.R.A. Görts-van Rijn & M.J. Jansen-Jacobs, Flora of the Guianas (series A) 

20: 1-34. 

Feuillet, C. & O. Poncy. 2002. Aristolochiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 

J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. 

Gard. 76 (2): 87-89. 

Hoehne, F.C. 1942. Aristolochiaceas. In F.C. Hoehne, Flora Brasílica 14(2): 1-141. 

Poncy, O. 1988. Quatre espèces nouvelles d’Aristolochia de Guyane française, et remarques nomenclaturales concernant une 

espèce d’Aublet. Bull. Mus. Hist. Nat. Paris, sér. 4, 10 (sect. B), Adansonia 4: 337-344. 

ASCLEPIADACEAE 

G.N. Morillo 

Asclepias curassavica L. 

Asclepias fruticosa L. 

Barjonia erecta (Vell.) K. Schum. 

Blepharodon glaucescens (Decne.) Fontella 

Blepharodon grandiflorus Benth. 

Blepharodon grandiflorus subsp. crassifolius (Schltr.) Morillo 

Blepharodon julianii Morillo [expected in GU] 

Blepharodon maigualidae Morillo 

GU 

GU 

GU 

FG 

FG 

FG 

FG 

FG 

SU FG 

FG? 

GU 

GU SU FG 

GU FG 

AM BO 

AM BO 

AM BO FG 

AM BO 

BO 

GU 

GU 

SU FG 

AM BO 

BO DA GU FG 

AM 

DA 

SU FG 

GU SU 

GU SU FG 

GU SU FG 

GU SU FG 


SU 

SU 

AM BO GU 

BO GU 

BO GU 

BO 

AM


Blepharodon mucronatus (Schltdl.) Decne. 

Blepharodon nitidus (Vell.) J.F. Macbr. 

Blepharodon tillettii Morillo 

Blepharodon ulei Schltr. 

Calotropis gigantea (L.) Dryand. [cultivated, naturalized] 

Cryptostegia madagascariensis Bojer ex Decne. [escaped from cultivation?] 

Cynanchum blandum (Decne.) Sundell 

Cynanchum buntingii Morillo 

Cynanchum chimantense Morillo 

Cynanchum goertsianum Morillo [published as C. görtsianum] 

Cynanchum guanchezii Morillo 

Cynanchum hirtellum (Oliv.) V.M. Badillo 

Cynanchum huberi Morillo 

Cynanchum mirifolium (Gleason & Moldenke) R.W. Holm 

Cynanchum montevidense Spreng. 

Cynanchum neei Morillo 

Cynanchum paraquense Morillo 

Cynanchum parviflorum Sw. 

Cynanchum prevostiae Morillo 

Cynanchum sobradoi Morillo 

Cynanchum stannardii Morillo 

Cynanchum strictum R.W. Holm 

Ditassa acerifolia Lasser & Maguire 

Ditassa angustifolia Decne. 

Ditassa auyantepuiensis (Steyerm.) Morillo 

Ditassa ayangannensis Morillo 

Ditassa bolivarensis (R.W. Holm) Morillo [expected in GU] 

Ditassa carnevalii (Morillo) Morillo 

Ditassa ciliata (Moldenke) Morillo 

Ditassa colellae (Morillo) Morillo 

Ditassa consanguinea Decne. 

Ditassa duidae Gleason 

Ditassa foldatsii Morillo 

Ditassa franciscoi (Morillo) S. Liede 

Ditassa gillespieae Morillo 

Ditassa juliani (Morillo) Morillo 

Ditassa liesneri Morillo 

Ditassa lindemanii Morillo 

Ditassa multinervia (Morillo) Morillo 

Ditassa obovata (R.W. Holm) Morillo [expected in GU] 

Ditassa olivaestevae Morillo 

Ditassa ottohuberi Morillo 

Ditassa pauciflora Decne. 

Ditassa roraimensis Schltr. 

Ditassa sipapoana Morillo 

Ditassa surinamensis Morillo 

Ditassa tatei Gleason & Moldenke 

Ditassa taxifolia Decne. 

Ditassa venamensis Morillo 

Ditassa verticillata Morillo 

Fischeria stellata (Vell.) E. Fourn. 

Gonolobus aristolochioides Kunth 

Gonolobus germanianus Morillo [expected in GU] 

Gonolobus glaber Decne. [specimens not seen, status uncertain] 

Gonolobus grandiflorus R. Br. [specimens not seen, status uncertain] 

Gonolobus tobagensis Urb. 

Macroditassa carolina Morillo [Mt. Roraima] 

Macroditassa laurifolia (Decne.) Fontella 

Marsdenia altissima (Jacq.) Dugand 

Marsdenia gillespieae Morillo 

Marsdenia guanchezii Morillo 

Marsdenia macrophylla (Humb. & Bonpl. ex Schult.) E. Fourn. 

Marsdenia rubrofusca Benth. ex E. Fourn. 

Marsdenia sprucei Rothe 

AM BO 


GU 

AM BO GU 

BO GU SU 

AM BO GU 

GU SU FG 

AM 

BO 

SU FG 

AM 

BO GU 

AM 

AM BO 

BO GU 

AM 

AM 

BO GU 

FG 

AM 

AM 

AM DA 

AM 

BO GU 

BO 

GU 

BO 

AM BO 

AM 

BO 

GU SU FG 

AM 

AM 

AM 

GU 

AM 

AM 

SU 

AM 

BO 

AM 

AM 

GU 

BO GU 

AM 

SU 

AM BO GU 

BO GU 

BO GU 

AM 

BO GU 

AM BO 

BO 

SU? 

GU? 

GU 

BO GU 

GU 

BO GU SU FG 

GU 

AM BO 

AM BO GU SU 


AM

210 


Marsdenia undulata (Jacq.) Dugand 

Marsdenia weddellii (E. Fourn.) Malme 

Marsdenia xerohylica Dugand 

Matelea amazonica Morillo 

Matelea badilloi Morillo 

Matelea bolivarensis Morillo 

Matelea cayennensis Morillo 

Matelea coriacea Morillo [expected in GU] 

Matelea cremersii Morillo 

Matelea cumanensis (Willd. ex Schult.) W.D. Stevens 

Matelea delascioi Morillo 

Matelea denticulata (Vahl) Fontella & E.A. Schwarz 

Matelea fournieri Morillo 

Matelea fucata Woodson 

Matelea funkiana Morillo 

Matelea glaziovii (E. Fourn.) Morillo 

Matelea gracieae Morillo 

Matelea grenandii Morillo 

Matelea herbacea Woodson 

Matelea hildegardiana Morillo 

Matelea hirsuta (Vahl) Woodson 

Matelea hoffmanii Morillo 

Matelea holstii Morillo & Carnevali 

Matelea jansen-jacobsiae Morillo 

Matelea jenmanii (Morillo) Morillo 

Matelea liesneri Morillo 

Matelea lourteigiae Morillo 

Matelea maritima (Jacq.) Woodson 

Matelea neblinae Morillo 

Matelea oldemanii Morillo 

Matelea palustris Aubl. 

Matelea planiflora (Jacq.) Dugand 

Matelea riparia Morillo 

Matelea sastrei Morillo 

Matelea squiresii (Rusby) Morillo 

Matelea stenopetala Sandwith 

Matelea stergiosii Morillo 

Matelea suareziae Morillo 

Matelea surinamensis Morillo 

Matelea yanomamica Morillo 

Metalepis albiflora Urb. 

Metalepis prevostiae Morillo 

Nephradenia linearis Benth. ex E. Fourn. 

Oxypetalum capitatum Mart. 

Sarcostemma clausum (Jacq.) Schult. 

Stenomeria decalepis Turcz. 

Tassadia aristata (Benth. ex E. Fourn.) Fontella [expected in the Guianas] 

Tassadia berteriana (Spreng.) W.D. Stevens 

Tassadia castellanosii Fontella 

Tassadia decaisneana Miq. 

Tassadia guianensis Decne. 

Tassadia ivonae Morillo 

Tassadia kamaensis (Morillo) Morillo 

Tassadia leptobotrys Decne. 

Tassadia marahuacensis Morillo 

Tassadia martiana Decne. [expected in the Guianas] 

Tassadia medinae (Morillo) Morillo 

Tassadia obovata Decne. 

Tassadia ovalifolia (E. Fourn.) Fontella [expected in the Guianas] 

Tassadia propinqua Decne. [expected in FG] 

Tassadia trailiana (Benth.) Fontella 


Asclepias altissima Jacq. =Marsdenia altissima 

Asclepias clausa Jacq. =Sarcostemma clausum 

BO 

SU 

BO 

AM 

BO GU FG 

BO 

BO 

BO 

GU 

FG 

FG 

DA GU 

BO 

BO 

BO 

DA GU 

GU 

GU 

SU 

SU 

SU 

FG 

FG 

FG 

BO 

BO 

GU 

GU 

GU 

SU 

AM 

AM GU SU FG 

AM 

AM 

BO GU 

GU SU 

FG 

FG 

BO GU 

GU 

SU FG 

BO DA GU SU 

AM 

AM 

BO 

BO 

GU FG 

SU 

AM 

FG 

FG 

AM BO GU 

BO GU SU 

AM BO DA GU 

GU 

SU FG 

AM BO 

AM 

BO GU SU 

GU SU 

GU SU FG 

AM 

AM 

BO 

BO 

SU 

AM 


AM 

AM DA GU SU 

AM GU


Asclepias gigantea L. =Calotropis gigantea 

Asclepias macrophylla Humb. & Bonpl. ex Schult. =Marsdenia macrophylla 

Asclepias planiflora Jacq. =Matelea planiflora 

Asclepias undulata Jacq. =Marsdenia undulata 

Astephanus mucronata Schltdl. =Blepharodon mucronatus 

Blepharodon auyantepuiense Steyerm. =Blepharodon ulei 

Blepharodon bifidus Schltr. =Blepharodon grandiflorus subsp. crassifolius 

Blepharodon ciliatum Moldenke =Ditassa ciliata 

Blepharodon crassifolius Schltr. =Blepharodon grandiflorus subsp. crassifolius 

Blepharodon hitchcockii Gleason =Blepharodon ulei 

Blepharodon steudelianum (Miq.) Pulle =Blepharodon nitidus 

Blepharodon steyermarkii R.W. Holm =Blepharodon nitidus 

Blepharodon venezuelense Moldenke =Blepharodon nitidus 

Blepharodon viscidum R.W. Holm =Nephradenia linearis 

Cynanchum acerifolium (Lasser & Maguire) Morillo =Ditassa acerifolia 

Cynanchum altissimum (Jacq.) Jacq. =Marsdenia altissima 

Cynanchum angustifolium (Decne.) Morillo =Ditassa angustifolia 

Cynanchum apocynellum (Gleason & Moldenke) Spellman =Tassadia obovata 

Cynanchum auyantepuiense Steyerm. =Ditassa auyantepuiensis 

Cynanchum bolivarense R.W. Holm =Ditassa bolivarensis 

Cynanchum carnevalii Morillo =Ditassa carnevalii 

Cynanchum clausum (Jacq.) Jacq. =Sarcostemma clausum 

Cynanchum colellae Morillo =Ditassa colellae 

Cynanchum denticulatum Vahl =Matelea denticulata 

Cynanchum duidae R.W. Holm =Ditassa ciliata 

Cynanchum fimbriatum Kunth =Matelea cumanensis 

Cynanchum franciscoi Morillo =Ditassa franciscoi 

Cynanchum guianense Spreng. =Matelea denticulata 

Cynanchum hirsutum Vahl =Matelea hirsuta 

Cynanchum holmianum Steyerm. =Ditassa duidae 

Cynanchum jenmanii Morillo =Matelea jenmanii 

Cynanchum julianii Morillo =Ditassa juliani 

Cynanchum kamaense Morillo =Tassadia kamaensis 

Cynanchum maritimum Jacq. =Matelea maritima 

Cynanchum medinae Morillo =Tassadia medinae 

Cynanchum multinervium Morillo =Ditassa multinervia 

Cynanchum nitidum Vell. =Blepharodon nitidus 

Cynanchum obovatum R.W. Holm =Ditassa obovata 

Cynanchum planiflorum Jacq. =Matelea planiflora 

Cynanchum recurvum (Rusby) Spellman =Tassadia obovata 

Cynanchum revolutum R.W. Holm =Cynanchum hirtellum 

Cynanchum roraimense (Schlltr.) Morillo =Ditassa roraimensis 

Cynanchum stellatum Vell. =Fischeria stellata 

Cynanchum stenolobum (Decne.) Morillo =Tassadia berteriana 

Cynanchum tatei (Gleason & Moldenke) R.W. Holm =Ditassa tatei 

Cynanchum taxifolium (Decne.) R.W. Holm =Ditassa taxifolia 

Cynanchum tepuiense Morillo =Ditassa ciliata 

Cynanchum undulatum (Jacq.) Jacq. =Marsdenia undulata 

Cynanchum viridiflorum G. Mey. =Matelea denticulata 

Ditassa aristata Benth. ex E. Fourn. =Tassadia aristata 

Ditassa blepharodontoides Schltr. =Blepharodon glaucescens 

Ditassa glaucescens Decne. =Blepharodon glaucescens 

Esmeraldia stricta E. Fourn. =Cynanchum strictum 

Fischeria calycina Decne. =Fischeria stellata 

Fischeria subaequalis S.F. Blake =Fischeria stellata 

Funastrum clausum (Jacq.) Schltr. =Sarcostemma clausum 

Glaziostelma ovalifolium E. Fourn. =Tassadia ovalifolia 

Gonolobus adenophorus Suess. =Matelea planiflora 

Gonolobus ligustrinus Decne. [not in flora area] 

Gonolobus maritimus (Jacq.) R. Br. =Matelea maritima 

Gonolobus riparius Kunth [not in flora area] 

Gonolobus squiresii Rusby =Matelea squiresii 

Gonolobus surinamensis Jonker =Matelea lourteigiae 

Gonolobus viridiflorus (G. Mey.) Schult. =Matelea denticulata

212 


Ibatia albiflora H. Karst. =Matelea cumanensis 

Ibatia maritima (Jacq.) Decne. =Matelea maritima 

Macroscepis hirsuta (Vahl) Schltr. =Matelea hirsuta 

Macroscepis trianae Decne. ex Triana =Matelea hirsuta 

Madarosperma trailiana Benth. =Tassadia trailiana 

Marsdenia amazonica Morillo =Marsdenia sprucei 

Marsdenia caurensis Morillo =Marsdenia altissima 

Marsdenia maculata Hook. =Marsdenia macrophylla 

Marsdenia margaritaria R.C. Foster =Marsdenia undulata 

Marsdenia mollissima E. Fourn. =Marsdenia altissima 

Matelea albiflora (H. Karst.) Dugand =Matelea cumanensis 

Matelea fimbriata (Kunth) Dugand =Matelea cumanensis 

Matelea latifolia Aubl. =Matelea palustris 

Matelea mediocris Woodson =Matelea squiresii 

Matelea trianae (Decne. ex Triana) Spellman =Matelea hirsuta 

Matelea viridiflora (G. Mey.) Woodson =Matelea denticulata 

Meresaldia stricta (E. Fourn.) Bullock =Cynanchum strictum 

Metastelma berterianum (Spreng.) Decne. =Tassadia berteriana 

Metastelma brasiliense K. Schum. [not in flora area] 

Metastelma campanulatum Decne. =Cynanchum parviflorum 

Metastelma decaisneana Benth. & Hook. =Tassadia decaisneana 

Metastelma mirifolium Gleason & Moldenke =Cynanchum mirifolium 

Metastelma paraquense (Morillo) Liede =Cynanchum paraquense 

Metastelma parviflorum (Sw.) R. Br. ex Schult. =Cynanchum parviflorum 

Metastelma stenolobum Decne. =Tassadia berteriana 

Metastelma steudelianum Miq. =Blepharodon nitidus 

Metastelma strictum (E. Fourn.) Spruce ex Knuth =Cynanchum strictum 

Metastelma strictum Gleason & Moldenke =Cynanchum strictum 

Oxypetalum berterianum Spreng. =Tassadia berteriana 

Roulinia blanda Decne. =Cynanchum blandum 

Roulinia voglii Suess. =Cynanchum montevidense 

Rouliniella guianensis (Decne.) Jonker =Cynanchum blandum 

Sarcostemma cumanense Kunth =Sarcostemma clausum 

Sarcostemma pubescens Kunth =Sarcostemma clausum 

Stephanotella killipii Woodson =Marsdenia rubrofusca 

Tassadia angustifolia Malme =Tassadia propinqua 

Tassadia apocynella Gleason & Moldenke =Tassadia obovata 

Tassadia recurva Rusby =Tassadia obovata 

Telminostelma corymbosum (Decne.) Fontella & Schwarz =Cynanchum blandum 

Vincetoxicum hirtellum Oliv. =Cynanchum hirtellum 

REFERENCES 

Fontella Pereira, J. 1977. Revisao taxonômica do gênero Tassadia Decaisne. Arq. Jard. Bot. Rio de Janeiro 21: 235-392. 

Jonker, F.P. 1940. Asclepiadaceae. In A. Pulle, Flora of Suriname 4(2): 326-352. 

Morillo, G. 1986. Asclepiadaceae II. Especies nuevas o interesantes en Venezuela y Guyana. Ernstia 37: 1-13. 

Morillo, G. 1988. Asclepiadaceae III, one new species and four new records. Proc. Kon. Ned. Akad. Wet. ser. C, 91(2): 161-164. 

Morillo, G. 1989. Contribución al conocimiento de las Asclepiadaceae suramericanas, principalmente de las Guayanas. Ernstia 

51: 2-15. 

Morillo, G. 1989. Veinte Asclepiadaceae sudamericanas nuevas para la ciencia y una nueva combinación. An. Jard. Bot. Madrid 

47(2): 349-359. 

Morillo, G. 1991. Once Asclepiadaceae suramericanas nuevas para la ciencia. Ernstia 1(3): 109-120. 

Morillo, G. 1993. Asclepiadaceae nuevas o interesantes de Venezuela y Guyana. Ernstia 3(2): 61-70.Morillo, G. 1994. Seis 

Asclepiadaceae Sudamericanas nuevas para la ciencia. Ernstia 4(1-2): 3-10. 

Morillo, G.N. 1997. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 129-177. 

Morillo, G. 1988. Asclepiadaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


89-94. 

Helosis cayennensis (Sw.) Spreng. 

Helosis cayennensis var. cayennensis 

Langsdorffia hypogaea Mart. 

BALANOPHORACEAE 

B. Hansen, J.S. Miller 



AM BO GU



Caldasia brasiliensis (Schott & Endl.) Kuntze =Helosis cayennensis 

Caldasia cayennensis (Sw.) Mutis ex Steud. =Helosis cayennensis 

Cynomorium cayennense Sw. =Helosis cayennensis 

Helosis brasiliensis Schott & Endl. =Helosis cayennensis 

Helosis guyanensis Rich. [nom. illeg. superfl.] =Helosis cayennensis 

Langsdorffia rubiginosa Wedd. =Langsdorffia hypogaea 

REFERENCES 

Gracie, C.A. 2002. Balanophoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 116. 

Hansen, B. 1980. Balanophoraceae. Flora Neotropica Monogr. 23: 1-80. 

Hansen, B. 1993. Balanophoraceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 14: 40-43. 

Lanjouw, J. 1932. Balanophoraceae. In A. Pulle, Flora of Suriname 1(1): 45-46. 

Miller, J.S. 1997. Balanophoraceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 393-395. 

Impatiens balsamina L. [probably escaped from cultivation] 

BALSAMINACEAE 

BASELLACEAE 


Anredera vesicaria (Lam.) C.F. Gaertn. [probably escaped from cultivation in 

FG] 

Basella alba L. [cultivated & naturalized] 


Anredera leptostachys (Moq.) Steenis =Anredera vesicaria 

Basella rubra L. =Basella alba 

Boussingaultia leptostachya Moq. =Anredera vesicaria 

Boussingaultia leptostachys Moq. =Anredera vesicaria 

GU 

AM BO FG 

GU SU FG 

REFERENCES 

Sperling, C.R. 1987. Systematics of the Basellaceae. Ph.D. Dissertation, Harvard University. U.M.I. Dissertation Information 

Service, Ann Arbor, Michigan. 

Steyermark, J.A. 1997. Basellaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 395-397. 

Batis maritima L. 

BATACEAE 

REFERENCES 

Jonker, F.P. 1953. Batidaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 91-93. 

BEGONIACEAE 

J.C. Lindeman, J.A. Steyermark†, D. Wasshausen 

Begonia fischeri Schrank 

Begonia fuchsioides Hook. 

Begonia glabra Aubl. 

Begonia guaduensis Kunth 

Begonia heloisana Brade 

Begonia hirsuta Aubl. 

Begonia humilis Dryand. 

Begonia jenmanii Tutin 

Begonia meridensis A. DC. 

Begonia nubicola L.B. Sm. & B.G. Schub. [Sierra de la Neblina] 

Begonia prieurii A. DC. 

Begonia semiovata Liebm. 

Begonia steyermarkii L.B. Sm. & B.G. Schub. 

Begonia ulmifolia Willd. 

GU SU FG? 

BO DA GU 

AM 

AM GU SU FG 

AM BO 

GU SU FG 

FG 

AM 

AM 

BO 

BO 

GU 

GU 

SU FG 

BO GU SU FG 


BO GU 

BO GU SU FG

214 



Begonia diffusa L.B. Sm. & B.G. Schub. [not in flora area] 

Begonia filipes Benth. [not in flora area] 

Begonia guyanensis A. DC. =Begonia semiovata 

Begonia hirtella Link [not in flora area] 

Begonia kaietukensis Tutin =Begonia fischeri 

Begonia kvistii L.B. Sm. [nomen nudum] =Begonia jenmanii 

Begonia roraimensis Tutin =Begonia fischeri 

Begonia scandens Sw. =Begonia glabra 

Begonia tovarensis Klotzsch =Begonia fischeri 

REFERENCES 

Heald, S.V. 2002. Begoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


117-118. 

Lindeman, J.C. 1984. Begoniaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 576-584. 

Smith, L.B., D.C. Wasshausen, J. Golding, & C.E. Karegeannes. 1986. Begoniaceae (illustrated key and annotated species list). 

Smithsonian Contrib. Bot. 60: 1-584. 

Smith, L and D. Wasshausen. 1989. Begoniaceae. Flora de Venezuela 4(1): 5-68 

Steyermark, J.A. 1997. Begoniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 397-403. 

BIGNONIACEAE 

A.H. Gentry†, S.O. Grose 

Adenocalymna apurense (Kunth) Sandwith 

Adenocalymna impressum (Rusby) Sandwith 

Adenocalymna inundatum Mart. ex DC. 

Adenocalymna inundatum var. surinamense Bureau & K. Schum. 

Adenocalymna prancei A.H. Gentry 

Adenocalymna purpurascens Rusby 

Adenocalymna saulense A.H. Gentry 

Adenocalymna subincanum Huber 

Amphilophium aschersonii Ule 

Amphilophium paniculatum (L.) Kunth 

Amphilophium paniculatum var. imatacense A.H. Gentry 

Amphilophium paniculatum var. molle (Schltdl. & Cham.) Standl. 

Amphilophium paniculatum var. paniculatum 

Anemopaegma alatum A.H. Gentry 

Anemopaegma cf. brevipes S. Moore 

Anemopaegma chamberlaynii (Sims) Bureau & K. Schum. 

Anemopaegma chrysoleucum (Kunth) Sandwith 

Anemopaegma floridum Mart. ex DC. 

Anemopaegma foetidum Bureau & K. Schum. 

Anemopaegma granvillei A.H. Gentry 

Anemopaegma ionanthum A.H. Gentry 

Anemopaegma jucundum Bureau & K. Schum. 

Anemopaegma karstenii Bureau & K. Schum. 

Anemopaegma longidens DC. 

Anemopaegma maguirei Sandwith 

Anemopaegma oligoneuron (Sprague & Sandwith) A.H. Gentry 

Anemopaegma paraense Bureau & K. Schum. 

Anemopaegma parkeri Sprague 

Anemopaegma patelliforme A.H. Gentry 

Anemopaegma robustum Bureau & K. Schum. 

Anemopaegma salicifolium (Kunth) Sandwith 

Arrabidaea arthrerion (DC.) Bureau ex K. Schum. 

Arrabidaea bilabiata (Sprague) Sandwith 

Arrabidaea brachypoda (DC.) Bureau 

Arrabidaea candicans (Rich.) DC. 

Arrabidaea carichanensis (Kunth) Bureau & K. Schum. 

Arrabidaea chica (Bonpl.) B. Verl. 

Arrabidaea cinerea Bureau ex K. Schum. 

Arrabidaea cinerea var. cinerea 

Arrabidaea cinerea var. longipila Sandwith 

BO GU? 

AM BO SU FG 



FG 

AM BO DA GU 

FG 

FG 

AM BO 



BO 

BO 

BO 

SU 

GU 


AM 

GU SU 

FG 

GU FG 

AM BO GU 

AM BO DA GU 

FG 

GU? SU 


BO DA GU? SU FG 


AM 

AM GU FG 

AM BO 

SU? 


BO DA GU? 

BO GU SU FG 

AM BO 

AM BO DA SU? FG 

GU FG 

GU FG 

GU


Arrabidaea cinnamomea (DC.) Sandwith 

Arrabidaea conjugata (Vell.) Mart. 

Arrabidaea corallina (Jacq.) Sandwith 

Arrabidaea egensis Bureau & K. Schum. 

Arrabidaea fanshawei Sandwith 

Arrabidaea florida DC. 

Arrabidaea grosourdyana (Baill.) Sandwith 

Arrabidaea inaequalis (DC. ex Splitg.) K. Schum. 

Arrabidaea japurensis (DC.) Bureau & K. Schum. 

Arrabidaea lachnaea (Bureau ex Bureau & K. Schum.) Sandwith 

Arrabidaea mollis (Vahl) Bureau ex K. Schum. 

Arrabidaea mollissima (Kunth) Bureau & K. Schum. 

Arrabidaea nigrescens Sandwith 

Arrabidaea oligantha Bureau & K. Schum. 

Arrabidaea oxycarpa Urb. 

Arrabidaea patellifera (Schltdl.) Sandwith 

Arrabidaea prancei A.H. Gentry [Sierra de la Neblina] 

Arrabidaea pubescens (L.) A.H. Gentry 

Arrabidaea revillae A.H. Gentry 

Arrabidaea selloi (Spreng.) Sandwith 

Arrabidaea spicata Bureau & K. Schum. 

Arrabidaea trailii Sprague 

Arrabidaea triplinervia (Mart. ex DC.) Baill. ex Bureau 

Arrabidaea tuberculata DC. 

Callichlamys latifolia (Rich.) K. Schum. 

Ceratophytum tetragonolobum (Jacq.) Sprague & Sandwith 

Clytostoma binatum (Thunb.) Sandwith 

Clytostoma sciuripabulum Bureau & K. Schum. 

Crescentia amazonica Ducke 

Crescentia cujete L. [cultivated only?] 

Cuspidaria subincana A.H. Gentry 

Cybistax antisyphilitica (Mart.) Mart. ex DC. 

Cydista aequinoctialis (L.) Miers 

Cydista lilacina A.H. Gentry 

Digomphia ceratophora A.H. Gentry 

Digomphia densicoma (Mart. ex DC.) Pilg. 

Digomphia laurifolia Benth. 

Distictella arenaria A.H. Gentry 

Distictella cremersii A.H. Gentry 

Distictella cuneifolia (DC.) Sandwith 

Distictella dasytricha Sandwith 

Distictella elongata (Vahl) Urb. 

Distictella laevis (Sandwith) A.H. Gentry 

Distictella magnoliifolia (Kunth) Sandwith 

Distictella monophylla Sandwith 

Distictella obovata Sandwith 

Distictella parkeri (DC.) Sprague & Sandwith 

Distictella pauciflora A.H. Gentry 

Distictella porphyrotricha Sandwith 

Distictis granulosa Bureau & K. Schum. 

Distictis pulverulenta (Sandwith) A.H. Gentry 

Godmania aesculifolia (Kunth) Standl. 

Haplolophium rodriguesii A.H. Gentry 

Jacaranda copaia (Aubl.) D. Don 

Jacaranda copaia subsp. copaia 

Jacaranda copaia subsp. spectabilis (Mart. ex DC.) A.H. Gentry 

Jacaranda duckei Vattimo 

Jacaranda obtusifolia Bonpl. 

Jacaranda obtusifolia subsp. obtusifolia 

Jacaranda obtusifolia subsp. rhombifolia (G. Mey.) A.H. Gentry 

Jacaranda orinocensis Sandwith 

Lundia corymbifera (Vahl) Sandwith 

Lundia densiflora DC. 

Lundia erionema DC. 

AM BO SU FG 

GU FG 

BO GU SU? FG 

AM GU 



BO GU FG 



AM BO 


BO 


GU SU FG 

AM BO 


AM GU 

AM BO GU SU 

GU SU 

BO 

FG 

BO GU SU FG 

FG 



BO GU SU 


AM BO DA GU 

AM BO 


BO GU SU 

SU 



AM 

AM BO GU 

AM BO GU 

AM BO 

SU FG 

FG 

AM 

AM GU SU FG 

AM 


AM 

BO GU FG? 

GU 

BO 

BO GU 



AM BO GU 

FG 




SU FG 


AM BO 


AM BO 

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AM BO GU SU 

AM GU SU FG

216 


Lundia puberula Pittier 

Macfadyena uncata (T.F. Andrews) Sprague & Sandwith 

Macfadyena unguis-cati (L.) A.H. Gentry 

Manaosella cordifolia (DC.) A.H. Gentry 

Mansoa alliacea (Lam.) A.H. Gentry 

Mansoa hymenaea (DC.) A.H. Gentry [cultivated, escaping?] 

Mansoa kerere (Aubl.) A.H. Gentry 

Mansoa kerere var. incarnata (Aubl.) A.H. Gentry 

Mansoa kerere var. kerere 

Mansoa onohualcoides A.H. Gentry 

Mansoa sp. nov. aff. standleyi 

Mansoa standleyi (Steyerm.) A.H. Gentry 

Mansoa verrucifera (Schltdl.) A.H. Gentry 

Martinella iquitosensis A. Samp. 

Martinella obovata (Kunth) Bureau & K. Schum. 

Melloa quadrivalvis (Jacq.) A.H. Gentry 

Memora alba (Aubl.) Miers 

Memora bracteosa (DC.) Bureau ex K. Schum. 

Memora cladotricha Sandwith 

Memora flavida (DC.) Bureau & K. Schum. 

Memora flaviflora (Miq.) Pulle 

Memora heterophylla (Kraenzl.) Sandwith 

Memora moringifolia (DC.) Sandwith 

Memora patula Miers 

Memora pseudopatula A.H. Gentry 

Memora racemosa A.H. Gentry 

Memora schomburgkii (DC.) Miers 

Memora tanaeciicarpa A.H. Gentry 

Mussatia hyacinthina (Standl.) Sandwith 

Mussatia prieurei (DC.) Bureau ex K. Schum. 

Parabignonia steyermarkii Sandwith 

Paragonia pyramidata (Rich.) Bureau 

Phryganocydia corymbosa (Vent.) Bureau ex K. Schum. 

Pithecoctenium crucigerum (L.) A.H. Gentry 

Pleonotoma albiflora (Salzm. ex DC.) A.H. Gentry 

Pleonotoma clematis (Kunth) Miers 

Pleonotoma dendrotricha Sandwith 

Pleonotoma echitidea Sprague & Sandwith 

Pleonotoma exserta A.H. Gentry 

Pleonotoma jasminifolia (Kunth) Miers 

Pleonotoma moringifolia (DC.) Miers [unverified name] 

Pleonotoma variabilis (Jacq.) Miers 

Potamoganos microcalyx (G. Mey.) Sandwith 

Pseudocalymma sagotii (Bureau & K. Schum.) Sandwith [unverified name] 

Pyrostegia dichotoma Miers ex K. Schum. 

Roentgenia sordida (Bureau & K. Schum.) Sprague & Sandwith 

Schlegelia fuscata A.H. Gentry 

Schlegelia macrophylla Ducke 

Schlegelia paraensis Ducke 

Schlegelia parviflora (Oerst.) Monach. 

Schlegelia roseiflora Ducke 

Schlegelia scandens (Briq. & Spruce) Sandwith 

Schlegelia spruceana K. Schum. 

Schlegelia violacea (Aubl.) Griseb. 

Sparattosperma leucanthum (Vell.) K. Schum. 

Stizophyllum inaequilaterum Bureau & K. Schum. 

Stizophyllum riparium (Kunth) Sandwith 

Tabebuia aurea (Silva Manso) Benth. & Hook. f. ex S. Moore 

Tabebuia barbata (E. Mey.) Sandwith 

Tabebuia capitata (Bureau & K. Schum.) Sandwith 

Tabebuia chrysantha (Jacq.) G. Nicholson [cultivated?] 

Tabebuia donnell-smithii Rose 

Tabebuia fluviatilis (Aubl.) DC. 

Tabebuia guayacan (Seem.) Hemsl. 

BO 



AM 

GU SU FG 

SU FG 


AM FG 


AM SU 

FG 


AM BO DA GU 

AM 

AM GU SU FG 

BO GU 

SU FG 


AM BO 

GU SU FG 


AM GU 

GU SU FG 

BO DA 

AM BO 

GU SU FG 


AM BO GU FG 

AM BO DA GU 



GU SU FG 





AM FG 

GU SU 

AM BO 

AM BO 

FG 

AM BO SU 

AM GU SU 

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FG 

FG 

GU SU FG 

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FG 

AM SU 

AM BO GU 


BO 



SU 

AM BO 


AM BO GU 

BO 

DA GU SU FG 

BO


Tabebuia impetiginosa (Mart. ex DC.) Standl. 

Tabebuia insignis (Miq.) Sandwith 

Tabebuia insignis var. insignis 

Tabebuia insignis var. monophylla Sandwith 

Tabebuia insignis var. pacimonensis Sandwith 

Tabebuia obscura (Bureau & K. Schum.) Sandwith 

Tabebuia ochracea (Cham.) Standl. 

Tabebuia ochracea subsp. heterotricha (DC.) A.H. Gentry 

Tabebuia ochracea subsp. neochrysantha (A.H. Gentry) A.H. Gentry 

Tabebuia orinocensis (Sandwith) A.H. Gentry 

Tabebuia pilosa A.H. Gentry 

Tabebuia rosea (Bertol.) DC. [cultivated] 

Tabebuia serratifolia (Vahl) G. Nicholson 

Tabebuia sp. A 

Tabebuia stenocalyx Sprague & Stapf 

Tabebuia subtilis Sprague & Sandwith 

Tabebuia uleana (Kraenzl.) A.H. Gentry 

Tanaecium jaroba Sw. 

Tanaecium nocturnum (Barb. Rodr.) Bureau & K. Schum. 

Tecoma stans (L.) Juss. ex Kunth [cultivated?] 

Tynanthus polyanthus (Bureau) Sandwith 

Tynanthus pubescens A.H. Gentry 

Tynanthus sastrei A.H. Gentry 

Xylophragma seemannianum (Kuntze) Sandwith 


Adenocalymna bilabiatum (Sprague) Sandwith =Arrabidaea bilabiata 

Adenocalymna heterophyllum Kraenzl. =Memora heterophylla 

Adenocalymna sagotii Bureau & K. Schum. =Pseudocalymma sagotii 

Adenocalymna symmetricum Rusby =Mansoa kerere 

Amphilophium molle Schltdl. & Cham. =Amphilophium paniculatum var. molle 

Anemopaegma carrerense Arm. =Anemopaegma karstenii 

Anomoctenium granulosum (Bureau & K. Schum.) Sandwith =Distictis granulosa 

Arrabidaea obliqua (Kunth) Bureau =Arrabidaea corallina 

Arrabidaea sordida Bureau & K. Schum. =Roentgenia sordida 

Bignonia aequinoctialis L. =Cydista aequinoctialis 

Bignonia aesculifolia Kunth =Godmania aesculifolia 

Bignonia alba Aubl. =Memora alba 

Bignonia albiflora Salzm. ex DC. =Pleonotoma albiflora 

Bignonia alliacea Lam. =Mansoa alliacea 

Bignonia apurensis Kunth =Adenocalymna apurense 

Bignonia aquatilis E. Mey. =Tabebuia fluviatilis 

Bignonia aurea Silva Manso =Tabebuia aurea 

Bignonia barbata E. Mey. =Tabebuia barbata 

Bignonia binata Thunb. =Clytostoma binatum 

Bignonia brachypoda DC. =Arrabidaea brachypoda 

Bignonia candicans Rich. =Arrabidaea candicans 

Bignonia carichanensis Kunth =Arrabidaea carichanensis 

Bignonia chamberlaynii Sims =Anemopaegma chamberlaynii 

Bignonia chica Bonpl. =Arrabidaea chica 

Bignonia chrysantha Jacq. =Tabebuia chrysantha 

Bignonia chrysoleuca Kunth =Anemopaegma chrysoleucum 

Bignonia cinnamomea DC. =Arrabidaea cinnamomea 

Bignonia clematis Kunth =Pleonotoma clematis 

Bignonia copaia Aubl. =Jacaranda copaia 

Bignonia corallina Jacq. =Arrabidaea corallina 

Bignonia cordifolia DC. =Manaosella cordifolia 

Bignonia corymbifera Vahl =Lundia corymbifera 

Bignonia crucigera L. =Pithecoctenium crucigerum 

Bignonia digitata E. Mey. =Tabebuia fluviatilis 

Bignonia echinata Aubl. =Pithecoctenium crucigerum 

Bignonia echinatum Jacq. =Pithecoctenium crucigerum 

Bignonia elongata Vahl =Distictella elongata 

Bignonia filicifolia Anderson =Jacaranda obtusifolia subsp. rhombifolia 

AM BO SU FG 




AM BO DA 

AM GU FG? 

AM BO GU 

AM BO GU 

BO 

AM BO 

AM BO 


AM 

BO DA GU SU? FG 

BO GU 

AM BO GU 

AM BO DA GU 

SU FG 

GU FG 

AM FG 

AM GU FG 

FG 

AM BO DA GU SU

218 


Bignonia fluviatilis Aubl. =Tabebuia fluviatilis 

Bignonia heterotricha DC. =Tabebuia ochracea subsp. heterotricha 

Bignonia impressa Rusby =Adenocalymna impressum 

Bignonia inaequalis DC. ex Splitg. =Arrabidaea inaequalis 

Bignonia incarnata Aubl. =Mansoa kerere var. incarnata 

Bignonia jasminifolia Kunth =Pleonotoma jasminifolia 

Bignonia kerere Aubl. =Mansoa kerere 

Bignonia latifolia Rich. =Callichlamys latifolia 

Bignonia leucantha Vell. =Sparattosperma leucanthum 

Bignonia magnoliifolia Kunth =Distictella magnoliifolia 

Bignonia martinii DC. =Martinella obovata 

Bignonia microcalyx G. Mey. =Potamoganos microcalyx 

Bignonia mollis Vahl =Arrabidaea mollis 

Bignonia mollissima Kunth =Arrabidaea mollissima 

Bignonia moringaefolia DC. =Pleonotoma moringifolia 

Bignonia paniculata L. =Amphilophium paniculatum 

Bignonia patellifera Schltdl. =Arrabidaea patellifera 

Bignonia picta Kunth =Cydista aequinoctialis 

Bignonia populifolia DC. =Melloa quadrivalvis 

Bignonia prieurei DC. =Mussatia prieurei 

Bignonia pubescens L. =Arrabidaea pubescens 

Bignonia pyramidata Rich. =Paragonia pyramidata 

Bignonia quadrivalvis Jacq. =Melloa quadrivalvis 

Bignonia riparia Kunth =Stizophyllum riparium 

Bignonia salicifolia Kunth =Anemopaegma salicifolium 

Bignonia selloi Spreng. =Arrabidaea selloi 

Bignonia serratifolia Vahl =Tabebuia serratifolia 

Bignonia tetragonoloba Jacq. =Ceratophytum tetragonolobum 

Bignonia tiliaefolia Kunth =Pithecoctenium crucigerum 

Bignonia tinctoria Arruda ? =Arrabidaea chica 

Bignonia uncata Andrews =Macfadyena uncata 

Bignonia unguis-cati L. =Macfadyena unguis-cati 

Bignonia variabilis Jacq. =Pleonotoma variabilis 

Bignonia verrucifera Schltdl. =Mansoa verrucifera 

Bignonia verrucosa Kunth =Arrabidaea carichanensis 

Callichlamys riparia Miq. =Callichlamys latifolia 

Ceratophytum tobagense (Urb.) Sprague & Sandwith =Ceratophytum tetragonolobum 

Cremastus lachnaeus Bureau ex Bureau & K. Schum. =Arrabidaea lachnaea 

Cybistax donnell-smithii (Rose) Seib. =Tabebuia donnell-smithii 

Cybistax millsii Miranda =Tabebuia donnell-smithii 

Distictella monophylla var. laevis Sandwith =Distictella laevis 

Distictella pulverulenta Sandwith =Distictis pulverulenta 

Distictella racemosa (Bureau & K. Schum.) Urb. =Distictella magnoliifolia 

Distictis robinsonii J. Johnst. =Anemopaegma karstenii 

Handroanthus durus (Bureau ex K. Schum.) Mattos =Tabebuia insignis var. insignis 

Jacaranda densicoma Mart. ex A. DC. =Digomphia densicoma 

Jacaranda filicifolia D. Don =Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda filicifolia var. puberula K. Schum. =Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda obtusifolia var. rhombifolia (G. Mey.) Sandwith =Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda paraensis (Huber) Vattimo =Jacaranda copaia subsp. spectabilis 

Jacaranda rhombifolia G. Mey. =Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda spectabilis Mart. ex DC. =Jacaranda copaia subsp. spectabilis 

Jacaranda superba Pittier =Jacaranda copaia subsp. spectabilis 

Lundia stricta Bureau & K. Schum. =Lundia densiflora 

Martinella insculpta Sprague & Sandwith =Martinella obovata 

Martinella manaosiana A. Samp. =Martinella iquitosensis 

Melloa populifolia (DC.) Britton =Melloa quadrivalvis 

Memora bilabiata Sprague =Arrabidaea bilabiata 

Memora ovata (Bureau & K. Schum.) Sprague & Sandwith =Memora schomburgkii 

Pachyptera alliacea (Lam.) A.H. Gentry =Mansoa alliacea 

Pachyptera kerere (Aubl.) Sandwith =Mansoa kerere 

Pachyptera kerere var. incarnata (Aubl.) A.H. Gentry =Mansoa kerere var. incarnata 

Pachyptera standleyi (Steyerm.) A.H. Gentry =Mansoa standleyi 

Paramansoa grosourdyana Baill. =Arrabidaea grosourdyana 

Petastoma kerere (Aubl.) Schnee =Mansoa kerere


Petastoma patelliferum (Schltdl.) Miers =Arrabidaea patellifera 

Pithecoctenium echinatum (Jacq.) Baill. =Pithecoctenium crucigerum 

Pithecoctenium granulosum (Bureau & K. Schum.) Sprague & Sandwith =Distictis granulosa 

Pithecoctenium uleanum Kraenzl. =Distictis granulosa 

Pseudocalymma alliaceum (Lam.) Sandwith =Mansoa alliacea 

Pseudocalymma alliaceum var. macrocalyx Sandwith =Mansoa standleyi 

Pseudocalymma standleyi Steyerm. =Mansoa standleyi 

Pseudopaegma oligoneuron Sprague & Sandwith =Anemopaegma oligoneuron 

Roentgenia tuberculata Sprague & Sandwith [name of uncertain application, apparently a nomen nudum] 

Saldanhaea seemanniana Kuntze =Xylophragma seemannianum 

Schizopsis polyantha Bureau =Tynanthus polyanthus 

Schlegelia lilacina Miq. =Schlegelia violacea 

Schlegelia parasitica (Sw.) Miers ex Griseb. [not in flora area] 

Spathodea ? bracteosa DC. =Memora bracteosa 

Spathodea corymbosa Vent. =Phryganocydia corymbosa 

Spathodea flaviflora Miq. =Memora flaviflora 

Spathodea obovata Kunth =Martinella obovata 

Spathodea orinocensis Kunth =Phryganocydia corymbosa 

Spathodea schomburgkii DC. =Memora schomburgkii 

Tabebuia aquatilis (E. Mey.) Sprague & Sandwith =Tabebuia fluviatilis 

Tabebuia caraiba (Mart.) Bureau =Tabebuia aurea 

Tabebuia chrysotricha (Mart. ex DC.) Standl. [not in flora area] 

Tabebuia dura (Bureau & K. Schum.) Sprague & Sandwith =Tabebuia insignis var. insignis 

Tabebuia floccosa (Bureau & K. Schum.) Sprague & Sandwith =Xylophragma seemannianum 

Tabebuia glomerata Urb. =Tabebuia capitata 

Tabebuia hypolepra Sprague & Sandwith =Tabebuia capitata 

Tabebuia insignis var. orinocensis Sandwith =Tabebuia orinocensis 

Tabebuia ipe var. integra (Sprague) Sandwith =Tabebuia impetiginosa 

Tabebuia japurensis DC. =Arrabidaea japurensis 

Tabebuia latifolia DC. =Martinella obovata 

Tabebuia longipes Baker =Tabebuia insignis var. monophylla 

Tabebuia millsii (Miranda) A.H. Gentry =Tabebuia donnell-smithii 

Tabebuia moritziana (Kraenzl.) Schnee =Arrabidaea grosourdyana 

Tabebuia neochrysantha A.H. Gentry =Tabebuia ochracea subsp. neochrysantha 

Tabebuia neurophylla Miq. =Arrabidaea patellifera 

Tabebuia obscura var. schultesiana (Sandwith) Sandwith =Tabebuia obscura 

Tabebuia roraimae Oliv. =Tabebuia insignis var. insignis 

Tabebuia rufescens J. Johnst. =Tabebuia chrysantha 

Tabebuia rufinervis Hoffmanns. ex DC. =Callichlamys latifolia 

Tabebuia subtilis var. schultesiana Sandwith =Tabebuia obscura 

Tanaecium praelongum Miers =Tanaecium jaroba 

Tecoma capitata Bureau & K. Schum. =Tabebuia capitata 

Tecoma conspicua DC. =Tabebuia serratifolia 

Tecoma dura Bureau ex K. Schum. =Tabebuia insignis var. insignis 

Tecoma fluviatilis (Aubl.) Miq. =Tabebuia fluviatilis 

Tecoma guayacan Seem. =Tabebuia guayacan 

Tecoma heterotricha DC. =Tabebuia ochracea subsp. heterotricha 

Tecoma impetiginosa Mart. ex DC. =Tabebuia impetiginosa 

Tecoma insignis Miq. =Tabebuia insignis 

Tecoma meyeriana DC. =Tabebuia fluviatilis 

Tecoma moritziana Kraenzl. =Arrabidaea grosourdyana 

Tecoma obscura Bureau & K. Schum. =Tabebuia obscura 

Tecoma ochracea Cham. =Tabebuia ochracea 

Tecoma patrisiana DC. =Tabebuia serratifolia 

Tecoma roraimae (Oliv.) K. Schum. =Tabebuia insignis var. insignis 

Tecoma serratifolia (Vahl) G. Don =Tabebuia serratifolia 

Tecoma uleana Kraenzl. =Tabebuia uleana 

Tetrastichella inaequalis (DC. ex Splitg.) Pichon =Arrabidaea inaequalis 

Tynanthus hyacinthinus Standl. =Mussatia hyacinthina 

Tynanthus myrianthus Bureau & K. Schum. =Tynanthus polyanthus 

Zeyheria surinamensis Miq. =Paragonia pyramidata 

REFERENCES 

Gentry, A.H. 1978. Bignoniaceae. In B. Maguire & collaborators, 1978. Botany of the Guayana Highland part 10. Mem. New 

York Bot. Gard. 29: 245-283.

220 


Gentry, A.H. 1980. Bignoniaceae - Part I (Crescentieae and Tourretieae). Flora Neotropica Monogr. 25(I): 1-130. 

Gentry, A.H. 1982. Bignoniaceae. In Z. Luces de Febres, Flora de Venezuela 8: 1-462. 

Gentry, A.H. 1992. Bignoniaceae - Part II (Tribe Tecomeae). Flora Neotropica Monogr. 25(II): 1-335. 

Gentry, A.H. 1997. Bignoniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 403-491. 

Lohmann, L.G, J.L. Brown, & S.A. Mori. 2002. Bignoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. 

Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. 

Mem. New York Bot. Gard. 76 (2): 118-139. 

Sandwith, N.Y. 1938. Bignoniaceae. In A. Pulle, Flora of Suriname 4(2): 1-86. 

Bixa orellana L. 

Bixa urucurana Willd. [=Bixa orellana?] 

Cochlospermum orinocense (Kunth) Steud. 

Cochlospermum vitifolium (Willd.) Spreng. 

BIXACEAE 

B.K. Holst, J.A. Steyermark† 


Bixa orellana var. urucurana (Willd.) Kuntze ex Pilg. =Bixa urucurana 

Cochlospermum insigne A. St.-Hil. [not in flora area] =Cochlospermum regium (Schr.) Pilger 

Cochlospermum parkeri Planch. =Cochlospermum orinocense 

Cochlospermum parviifolium Planch. =Cochlospermum orinocense 

Cochlospermum triphyllum (S.F. Blake) Pittier =Cochlospermum vitifolium 

Cochlospermum wentii Pulle =Cochlospermum orinocense 

Maximilianea triphylla S.F. Blake =Cochlospermum vitifolium 

Orellana americana var. urucurana Kuntze =Bixa urucurana 

REFERENCES 

Heald, S.V. 2002. Bixaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 

2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 139. 

Poppendieck, H.H. 1981. Cochlospermaceae. Flora Neotropica Monogr. 27: 1-33. 

Steyermark, J.A., & B.K. Holst. 1997. Bixaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 

3: 492-495. 

Bombacopsis nervosa (Uittien) A. Robyns 

Catostemma altsonii Sandwith 

Catostemma commune Sandwith 

Catostemma durifolius W.S. Alverson 

Catostemma ebracteolatum Steyerm. 

Catostemma fragrans Benth. 

Catostemma hirsutulum Steyerm. 

Catostemma marahuacense Steyerm. 

Catostemma sancarlosianum Steyerm. 

Cavanillesia sp. A 

Ceiba pentandra (L.) Gaertn. 

Eriotheca crassa (Uittien) A. Robyns 

Eriotheca globosa (Aubl.) A. Robyns 

Eriotheca longitubulosa A. Robyns 

Eriotheca surinamensis (Uittien) A. Robyns 

Huberodendron swietenioides (Gleason) Ducke 

Matisia amplifolia Pittier 

Matisia ochrocalyx K. Schum. 

Ochroma pyramidale (Cav. ex Lam.) Urb. 

Pachira amazonica (A. Robyns) W.S. Alverson 

Pachira aquatica Aubl. 

Pachira aracamuniana (Steyerm.) W.S. Alverson 

Pachira coriacea (Mart.) W.S. Alverson 

Pachira cowanii (A. Robyns) W.S. Alverson 

Pachira dolichocalyx A. Robyns 

Pachira faroensis (Ducke) W.S. Alverson 

Pachira flaviflora (Pulle) Fern. Alonso 

BOMBACACEAE 

W.S. Alverson, L.J. Dorr, J.A. Steyermark† 

GU SU FG 




SU FG 

AM BO DA 

GU 

GU 

BO GU 

AM BO 

GU SU FG 

AM 

AM 

AM 

AM 

BO 


SU FG 


FG 

BO 

GU SU FG 

FG 


BO GU FG 

AM BO 

AM 

AM 

AM 

AM 


AM 

GU FG 

GU SU FG


Pachira fuscolepidota (Steyerm.) W.S. Alverson 

Pachira gracilis (A. Robyns) W.S. Alverson 

Pachira gracilis subsp. bolivarensis (Steyerm.) W.S. Alverson 

Pachira gracilis subsp. gracilis 

Pachira humilis Spruce ex Decne. 

Pachira insignis (Sw.) Sw. ex Savigny 

Pachira liesneri (Steyerm.) W.S. Alverson 

Pachira macrocalyx (Ducke) Fern. Alonso 

Pachira mawarinumae (Steyerm.) W.S. Alverson 

Pachira minor (Sims) Hemsl. [Sierra de la Neblina] 

Pachira nitida Kunth 

Pachira obovata (A. Robyns) W.S. Alverson [Sierra de la Neblina] 

Pachira orinocensis (A. Robyns) W.S. Alverson 

Pachira paraensis (Ducke) W.S. Alverson 

Pachira pseudofaroensis (A. Robyns) W.S. Alverson 

Pachira quinata (Jacq.) W.S. Alverson 

Pachira robynsii (Steyerm. & W.D. Stevens) W.S. Alverson [Sierra de la 

Neblina] 

Pachira rupicola (A. Robyns) W.S. Alverson 

Pachira sordida (R.E. Schult.) W.S. Alverson 

Pachira tepuiensis (Steyerm.) W.S. Alverson 

Pachira trinitensis Urb. 

Pachira yapacanae Steyerm. ex W.S. Alverson 

Pseudobombax croizatii A. Robyns 

Quararibea amazonica Ulbr. 

Quararibea duckei Huber 

Quararibea guianensis Aubl. 

Quararibea lasiocalyx (K. Schum.) Vischer 

Quararibea spatulata Ducke 

Scleronema guianense Sandwith 

Scleronema micranthum (Ducke) Ducke [Sierra de la Neblina] 


Bombacopsis amazonica A. Robyns =Pachira amazonica 

Bombacopsis coriacea A. Robyns =Pachira robynsii 

Bombacopsis cowanii A. Robyns =Pachira cowanii 

Bombacopsis fendleri (Seem.) Pittier =Pachira quinata 

Bombacopsis jaris Pittier [nomen nudum] =Pachira quinata 

Bombacopsis jenmanii (Oliv.) Lasser =Pachira minor 

Bombacopsis macrocalyx (Ducke) A. Robyns =Pachira macrocalyx 

Bombacopsis mucronulata Pittier =Pachira trinitensis 

Bombacopsis obovata A. Robyns =Pachira obovata 

Bombacopsis orinocensis A. Robyns =Pachira orinocensis 

Bombacopsis pachiroides auct. non Pittier: sensu Ll. Williams =Pachira quinata 

Bombacopsis pachiroides auct. non Pittier: Ll. Williams =Pachira rupicola 

Bombacopsis pachiroides Pittier =Pachira trinitensis 

Bombacopsis paraensis (Ducke) A. Robyns =Pachira paraensis 

Bombacopsis pseudamazonica A. Robyns =Pachira amazonica 

Bombacopsis quinata (Jacq.) Dugand =Pachira quinata 

Bombacopsis rupicola A. Robyns =Pachira rupicola 

Bombacopsis sepium Pittier =Pachira quinata 

Bombacopsis trinitensis (Urb.) A. Robyns =Pachira trinitensis 

Bombacopsis trinitensis var. mucronulata (Pittier) A. Robyns =Pachira trinitensis 

Bombacopsis wurdackii A. Robyns =Pachira amazonica 

Bombax aquaticum (Aubl.) K. Schum. =Pachira aquatica 

Bombax coriaceum Mart. =Pachira coriacea 

Bombax crassum Uittien =Eriotheca crassa 

Bombax faroense Ducke =Pachira faroensis 

Bombax flaviflorum Pulle =Pachira flaviflora 

Bombax globosum Aubl. =Eriotheca globosa 

Bombax humile (Spruce ex Decne.) Benth. ex K. Schum. =Pachira humilis 

Bombax jenmanii Oliv. =Pachira minor 

Bombax macrocarpum (Schltdl. & Cham.) K. Schum. =Pachira aquatica 

Bombax minus (Sims) Ducke =Pachira minor 

AM 

AM BO 

BO 

AM 

AM 

AM 

GU SU FG 

AM 


AM 

AM 

AM BO 

AM 

AM 

AM BO GU 

AM 

BO 

AM 

BO 

BO 

AM 

AM BO 

AM 

FG? 

FG 

SU FG 

GU SU FG 

FG 

FG 

GU

222 


Bombax nervosum Uittien =Bombacopsis nervosa 

Bombax orinocense Kunth (Bixaceae) =Cochlospermum orinocense 

Bombax paraense Ducke =Pachira paraensis 

Bombax pentandrum L. =Ceiba pentandra 

Bombax poissonianum K. Schum. =Pachira nitida 

Bombax pyramidale Cav. ex Lam. =Ochroma pyramidale 

Bombax quinatum Jacq. =Pachira quinata 

Bombax rigidifolium Ducke =Pachira aquatica 

Bombax sordidum R.E. Schult. =Pachira sordida 

Bombax spectabile Ulbr. =Pachira insignis 

Bombax surinamense Uittien =Eriotheca surinamensis 

Bombax vitifolium Willd. (Bixaceae) =Cochlospermum vitifolium 

Carolinea macrocarpa Schltdl. & Cham. =Pachira aquatica 

Carolinea minor Sims =Pachira minor 

Catostemma clarkii Steyerm. =Catostemma sancarlosianum 

Catostemma macrosperma (Sagot) Lemée =Catostemma fragrans 

Catostemma micranthum Ducke =Scleronema micranthum 

Catostemma pubistylum Steyerm. =Catostemma sancarlosianum 

Ceiba occidentalis (Spreng.) Burkill =Ceiba pentandra 

Matisia apaporiensis Cuatrec. =Matisia ochrocalyx 

Matisia lasiocalyx K. Schum. =Quararibea lasiocalyx 

Matisia muricata (Cuatrec.) Cuatrec. =Matisia ochrocalyx 

Ochroma lagopus Sw. =Ochroma pyramidale 

Pachira cardonae Cuatrec. =Pachira minor 

Pachira fendleri Seem. =Pachira quinata 

Pachira jenmanii (Oliv.) A. Terracc. =Pachira minor 

Pachira longiflora Hook. =Pachira aquatica 

Pachira macrocarpa (Schltdl. & Cham.) Walp. =Pachira aquatica 

Pachira obtusa Spruce ex K. Schum. =Pachira nitida 

Pachira pustulifera Pittier =Pachira aquatica 

Pachira villosula Pittier =Pachira aquatica 

Pochota amazonica (A. Robyns) Steyerm. & W.D. Stevens =Pachira amazonica 

Pochota aracamuniana Steyerm. =Pachira aracamuniana 

Pochota coriacea (Mart.) Steyerm. & W.D. Stevens =Pachira coriacea 

Pochota coriacea subsp. orinocensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira coriacea 

Pochota cowanii (A. Robyns) Steyerm. & W.D. Stevens =Pachira cowanii 

Pochota ewelii Steyerm. =Pachira amazonica 

Pochota flaviflora (Pulle) Steyerm. & W.D. Stevens =Pachira flaviflora 

Pochota fuscolepidota Steyerm. =Pachira fuscolepidota 

Pochota gracilis (A. Robyns) Steyerm. & W.D. Stevens =Pachira gracilis 

Pochota gracilis subsp. bolivarensis Steyerm. =Pachira gracilis subsp. bolivarensis 

Pochota humilis (Spruce ex Decne.) Steyerm. & W.D. Stevens =Pachira humilis 

Pochota liesneri Steyerm. =Pachira liesneri 

Pochota maguirei (A. Robyns) Steyerm. & W.D. Stevens =Pachira gracilis subsp. bolivarensis 

Pochota mawarinumae Steyerm. =Pachira mawarinumae 

Pochota minor (Sims) Steyerm. & W.D. Stevens =Pachira minor 

Pochota nitida (Kunth) Steyerm. & W.D. Stevens =Pachira nitida 

Pochota obovata (A. Robyns) Steyerm. & W.D. Stevens =Pachira obovata 

Pochota orinocensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira orinocensis 

Pochota pseudamazonica (A. Robyns) Steyerm. & W.D. Stevens =Pachira amazonica 

Pochota pseudofaroensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira pseudofaroensis 

Pochota quinata (Jacq.) W.D. Stevens =Pachira quinata 

Pochota redmondii Steyerm. =Pachira obovata 

Pochota robynsii Steyerm. & W.D. Stevens =Pachira robynsii 

Pochota sordida (R.E. Schult.) Steyerm. & W.D. Stevens =Pachira sordida 

Pochota tepuiensis Steyerm. =Pachira tepuiensis 

Pochota trinitensis (Urb.) Steyerm. & W.D. Stevens =Pachira trinitensis 

Pochota vulgaris Ram. Goyena =Pachira quinata 

Pochota wurdackii (A. Robyns) Steyerm. & W.D. Stevens =Pachira amazonica 

Quararibea apaporiensis (Cuatrec.) A. Robyns =Matisia ochrocalyx 

Quararibea muricata Cuatrec. =Matisia ochrocalyx 

Quararibea ochrocalyx (K. Schum.) Vischer =Matisia ochrocalyx 

Quararibea turbinata Poir. [specimens apparently referable to Quararibea duckei] 

Rhodognaphalopsis coriacea (Mart.) A. Robyns =Pachira coriacea 

Rhodognaphalopsis coriacea subsp. orinocensis A. Robyns =Pachira coriacea


Rhodognaphalopsis coriacea var. sordida (R.E. Schult.) A. Robyns =Pachira sordida 

Rhodognaphalopsis discolor A. Robyns =Pachira sordida 

Rhodognaphalopsis faroensis (Ducke) A. Robyns =Pachira faroensis 

Rhodognaphalopsis flaviflora var. flaviflora =Pachira flaviflora 

Rhodognaphalopsis flaviflora var. glabra A. Robyns =Pachira flaviflora 

Rhodognaphalopsis gracilis A. Robyns =Pachira gracilis 

Rhodognaphalopsis humilis (Spruce ex Decne.) A. Robyns =Pachira humilis 

Rhodognaphalopsis maguirei A. Robyns =Pachira gracilis subsp. bolivarensis 

Rhodognaphalopsis minor (Sims) A. Robyns =Pachira minor 

Rhodognaphalopsis minor var. amblyophylla A. Robyns =Pachira minor 

Rhodognaphalopsis minor var. minor =Pachira minor 

Rhodognaphalopsis nitida (Kunth) A. Robyns =Pachira nitida 

Rhodognaphalopsis pseudofaroensis A. Robyns =Pachira pseudofaroensis 

Scleronema neblinense Steyerm. =Scleronema micranthum 

Sophia carolina L. =Pachira aquatica 

REFERENCES 

Alverson, W.S. 1994. New species and combinations of Catostemma and Pachira (Bombacaceae) from the Venezuelan Guayana. 

Novon 4(1): 3-8. 

Alverson, W.S. & J.A. Steyermark. 1997. Bombaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 3: 496-527. 

Alverson, W.S. & S.A. Mori. 2002. Bombaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 


Gard. 76 (2): 139-145. 

Robyns, A. 1963. Essai de monographie du genre Bombax s.l. Bull. Jard. Bot. État 33: 1-311. 

Robyns, A. 1967. Bombacaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 7. Mem. New York Bot. 

Gard. 17(1): 1-439. 

Steyermark, J.A. 1987. Bombacaceae. In Flora of the Venezuelan Guayana - III. Ann. Missouri Bot. Gard. 74(3): 636-646. 

Steyermark, J.A. & W.D. Stevens. 1988. Notes on Rhodognaphalopsis and Bombacopsis in the Guayanas. Ann. Missouri Bot. 

Gard. 75: 396-398. 

Uittien, H. 1932. Bombacaceae. In A. Pulle, Flora of Suriname 3(1): 26-33. 

Archytaea angustifolia Maguire 

Archytaea triflora Mart. 

Bonnetia ahogadoi (Steyerm.) A.L. Weitzman & P.F. Stevens 

Bonnetia bolivarensis Steyerm. 

Bonnetia celiae Maguire 

Bonnetia chimantensis Steyerm. 

Bonnetia cordifolia Maguire 

Bonnetia crassa Gleason 

Bonnetia euryanthera Steyerm. 

Bonnetia fasciculata A.L. Weitzman & P.F. Stevens 

Bonnetia huberiana Steyerm. 

Bonnetia jauaensis Maguire 

Bonnetia kathleenae Lasser 

Bonnetia lanceifolia Kobuski 

Bonnetia liesneri Steyerm. 

Bonnetia maguireorum Steyerm. 

Bonnetia multinervia (Maguire) Steyerm. [expected in GU?] 

Bonnetia neblinae Maguire 

Bonnetia paniculata Spruce ex Benth. 

Bonnetia ptariensis Steyerm. 

Bonnetia roraimae Oliv. 

Bonnetia roseiflora Maguire 

Bonnetia rubicunda (Sastre) A.L. Weitzman & P.F. Stevens 

Bonnetia sessilis Benth. 

Bonnetia steyermarkii Kobuski 

Bonnetia tepuiensis Kobuski & Steyerm. 

Bonnetia tristyla Gleason 

Bonnetia wurdackii Maguire 

BONNETIACEAE 


Acopanea ahogadoi Steyerm. =Bonnetia ahogadoi 

AM 

AM BO GU 

BO 

BO 

AM 

BO 

BO 

AM 

BO 

BO 

BO 

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AM 

AM 

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BO 

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AM BO GU 

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224 


Archytaea multiflora Benth. =Archytaea triflora 

Bonnetia dinizii Huber =Bonnetia paniculata 

Bonnetia duidae Kobuski & Steyerm. (Clusiaceae) =Neotatea duidae 

Bonnetia guaiquinimae Steyerm. =Bonnetia lanceifolia 

Bonnetia longifolia Gleason (Clusiaceae) =Neotatea longifolia 

Bonnetia martiana Maguire =Bonnetia sessilis 

Bonnetia martiana var. venadoensis Maguire =Bonnetia sessilis 

Bonnetia neblinensis Steyerm. (Clusiaceae) =Neotatea neblinae 

Bonnetia parviflora Spruce ex Benth. =Bonnetia paniculata 

Bonnetia phelpsii Gleason =Bonnetia sessilis 

Bonnetia tepuiensis subsp. minor Steyerm. =Bonnetia tepuiensis 

Bonnetia toronoensis Steyerm. =Bonnetia chimantensis 

Bonnetia tristyla subsp. nervosa Steyerm. =Bonnetia tristyla 

Kieseria sessilis (Benth.) Kuntze =Bonnetia sessilis 

Neblinaria celiae Maguire =Bonnetia maguireorum 

Neblinaria rubicunda Sastre =Bonnetia rubicunda 

Neogleasonia duidae (Kobuski & Steyerm.) Maguire (Clusiaceae) =Neotatea duidae 

Neogleasonia multinervia Maguire =Bonnetia multinervia 

Neogleasonia wurdackii Maguire =Bonnetia fasciculata 

REFERENCES 

Kobuski, C.E. 1948. Studies in the Theaceae, XVII. A review of the genus Bonnetia. J. Arn. Arb. 29: 393-413. 

Maguire, B. 1972. Bonnetiaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot. 

Gard. 23: 1-832. 

Bourreria cumanensis (Loefl.) O.E. Schulz 

Cordia alliodora (Ruiz & Pav.) Oken 

Cordia bicolor DC. 

Cordia bullata (L.) Roem. & Schult. 

Cordia bullata subsp. humilis (Jacq.) Gaviria 

Cordia cabanayensis Gaviria 

Cordia collococca Sandmark ex L. 

Cordia cremersii Feuillet 

Cordia curassavica (Jacq.) Roem. & Schult. 

Cordia dentata Poir. 

Cordia exaltata Lam. 

Cordia fallax I.M. Johnst. 

Cordia fanchoniae Feuillet 

Cordia fulva I.M. Johnst. 

Cordia goeldiana Huber 

Cordia grandiflora (Desv.) Kunth 

Cordia koemarae J.S. Mill. 

Cordia laevifrons I.M. Johnst. 

Cordia lomatoloba I.M. Johnst. 

Cordia marioniae Feuillet 

Cordia multispicata Cham. 

Cordia naidophila I.M. Johnst. 

Cordia nervosa Lam. 

Cordia nodosa Lam. 

Cordia panamensis Riley 

Cordia panicularis Rudge 

Cordia polycephala (Lam.) I.M. Johnst. 

Cordia polystachya Kunth 

Cordia roraimae I.M. Johnst. 

Cordia sagotii I.M. Johnst. 

Cordia scabrifolia DC. 

Cordia schomburgkii DC. 

Cordia sericicalyx DC. 

Cordia sipapoi Gaviria 

Cordia sp. A 

Cordia sprucei Mez 

BORAGINACEAE 

C. Feuillet, J. Gaviria, R. Gómez, J.S. Miller, G. Rodríguez 

BO 



BO DA GU 

BO DA GU 

BO 

BO DA GU 

FG 


BO DA 


BO DA GU 

FG 

SU FG 

FG 

AM BO DA GU 

SU 

SU FG 

BO GU SU FG 

GU 

GU 

AM FG 

GU SU FG 


BO DA 

GU SU FG 

BO GU SU FG 

AM 

BO 

BO GU SU FG 

AM GU 

BO GU SU FG 

BO DA GU SU 

AM 

AM 

AM FG

Cordia stenostachya Killip ex Gaviria 

Cordia steyermarkii Gaviria 

Cordia tetrandra Aubl. 

Cordia tomentosa Lam. ex Roem. & Schult. 

Cordia toqueve Aubl. [expected in SU] 

Cordia umbellifera Killip ex G. Agostini 

Cordia williamsii G. Agostini ex Gaviria 

Cordia wurdackiana Feuillet 

Heliotropium angiospermum Murray 

Heliotropium curassavicum L. 

Heliotropium filiforme Lehm. 

Heliotropium indicum L. [cultivated, persists as an escape] 

Heliotropium lagoense (Warm.) Gürke 

Heliotropium polyphyllum Lehm. 

Heliotropium procumbens Mill. 

Heliotropium purdiei I.M. Johnst. 

Heliotropium ternatum Vahl 

Lepidocordia punctata Ducke 

Tournefortia bicolor Sw. 

Tournefortia candidula (Miers) I.M. Johnst. 

Tournefortia cuspidata Kunth 

Tournefortia hirsutissima L. 

Tournefortia maculata Jacq. 

Tournefortia melanochaeta DC. 

Tournefortia paniculata Cham. 

Tournefortia paniculata var. spigeliiflora (DC.) I.M. Johnst. 

Tournefortia ulei Vaupel 

Tournefortia volubilis L. 



Bourreria viridis Rusby =Cordia sericicalyx 

Cerdana alliodora Ruiz & Pav. =Cordia alliodora 

Cordia alba auct. non DC. =Cordia dentata 

Cordia alliodora (Ruiz & Pav.) Cham. ex DC. =Cordia alliodora (Ruiz & Pav.) Oken 

Cordia aubletii DC. =Cordia schomburgkii 

Cordia canescens Willd. ex Roem. & Schult. =Cordia polystachya 

Cordia carnosa Rusby =Cordia bicolor 

Cordia collococa Aubl. non L. =Cordia nodosa 

Cordia cordifolia Kunth =Cordia tetrandra 

Cordia cordifolia Willd. ex Roem. & Schult. [nom. superfl.] =Cordia tetrandra 

Cordia coriacea Sagot ex Benoist =Cordia sagotii 

Cordia corymbosa Willd. ex Roem. & Schult. [nom. superfl.] =Cordia polycephala 

Cordia corymbosa (Desv.) G. Don =Cordia polycephala 

Cordia dasycephala (Desv.) Kunth =Cordia bullata subsp. humilis 

Cordia dentata Vahl =Cordia dentata 

Cordia exaltata var. melanoneura (Klotzsch) I.M. Johnst. =Cordia exaltata 

Cordia flavescens Aubl. [name of uncertain application, referable to Lauraceae] 

Cordia formicarum Willd. ex Roem. & Schult. =Cordia nodosa 

Cordia globosa (Jacq.) Kunth =Cordia bullata subsp. humilis 

Cordia globosa var. humilis (Jacq.) I.M. Johnst. =Cordia bullata subsp. humilis 

Cordia globosa subsp. humilis (Jacq.) Borh. =Cordia bullata subsp. humilis 

Cordia guianensis Klotzsch [nomen nudum] =Cordia fallax 

Cordia gujanensis (Desv.) Roem. & Schult. =Cordia fallax 

Cordia hebecarpa DC. =Cordia toqueve 

Cordia heterophylla Willd. ex Roem. & Schult. [nom. superfl.] =Cordia toqueve 

Cordia heterophylla Poir. =Cordia toqueve 

Cordia hirta I.M. Johnst. =Cordia sagotii 

Cordia hispidissima DC. =Cordia nodosa 

Cordia humilis (Jacq.) G. Don =Cordia bullata subsp. humilis 

Cordia ierensis Britton =Cordia sericicalyx 

Cordia lockhartii Kuntze [nom. inval.] =Cordia bicolor 

Cordia lucida Splitg. ex Pulle [nomen nudum] =Cordia schomburgkii 

Cordia macrostachya (Jacq.) Roem. & Schult. =Cordia curassavica 

Cordia melanoneura Klotzsch [nomen nudum] =Cordia exaltata 

Cordia miranda DC. =Cordia nodosa 

AM 

BO 


SU FG 

BO 

BO 

BO 

BO 

GU 

GU 

FG 

GU SU FG? 



BO 

BO 

SU 

AM BO 

BO 

DA GU 

BO GU 

BO GU SU 


BO GU 


DA GU 

BO GU SU FG 

GU FG 

GU SU FG 

GU SU FG 


BO GU FG

226 


Cordia monosperma (Jacq.) Roem. & Schult. =Cordia polycephala 

Cordia opaca Rusby =Cordia sericicalyx 

Cordia pubescens Willd. ex Roem. & Schult. =Cordia toqueve 

Cordia scabrida Mart. =Cordia exaltata 

Cordia scabrida auct. non Fresen. 1857. =Cordia naidophila 

Cordia scandens Poir. (Dichapetalaceae) =Dichapetalum rugosum 

Cordia spinescens L. [not in flora area] 

Cordia tetraphylla Aubl. (Combretaceae) =Buchenavia tetraphylla 

Cordia tobagensis Urb. =Cordia schomburgkii 

Cordia tobagensis var. broadwayi Urb. =Cordia schomburgkii 

Cordia umbraculifera DC. =Cordia tetrandra 

Cordia viridis (Rusby) I.M. Johnst. =Cordia sericicalyx 

Cordia volubilis Pittier =Cordia nodosa 

Crematomia cumanensis (Loefl.) Miers =Bourreria cumanensis 

Heliotropium foliosum Roem. & Schult. =Heliotropium polyphyllum 

Heliotropium fumana (Fresen.) Gürke =Heliotropium ternatum 

Heliotropium ottonis Lehm. =Heliotropium ternatum 

Heliotropium scorpioides Kunth =Heliotropium angiospermum 

Heliotropium trinitensis Urb. =Heliotropium lagoense 

Lithocardium sprucei (Mez) Kuntze =Cordia sprucei 

Messerschmidia candidula Miers =Tournefortia candidula 

Schleidenia fumana Fresen. =Heliotropium ternatum 

Schleidenia lagoensis Warm. =Heliotropium lagoense 

Schleidenia polyphylla (Lehm.) Fresen. =Heliotropium polyphyllum 

Tournefortia aubletii J.F. Macbr. =Tournefortia bicolor 

Tournefortia bicolor var. bicolor =Tournefortia bicolor 

Tournefortia bicolor var. calycosa J.D. Sm. =Tournefortia bicolor 

Tournefortia hirsutissima auct. non L.: sensu Pulle =Tournefortia cuspidata 

Tournefortia hirsutissima L. f. bicolor (Sw.) D.L. Nash =Tournefortia bicolor 

Tournefortia syringifolia Vahl =Tournefortia maculata 

Tournefortia tenuiflora Willd. ex Roem. & Schult. (Rubiaceae) =Declieuxia tenuiflora 

Varronia bullata L. =Cordia bullata 

Varronia corymbosa Desv. [nomen nudum] =Cordia polycephala 

Varronia curassavica Jacq. =Cordia curassavica 

Varronia dasycephala Desv. =Cordia bullata subsp. humilis 

Varronia globosa Jacq. =Cordia bullata subsp. humilis 

Varronia grandiflora Desv. =Cordia grandiflora 

Varronia guianensis Desv. =Cordia curassavica 

Varronia humilis Jacq. =Cordia bullata subsp. humilis 

Varronia lineata auct. non L. [nom. illeg.] =Cordia polycephala 

Varronia macrostachya Jacq. =Cordia curassavica 

Varronia monosperma Jacq. =Cordia polycephala 

Varronia polycephala Lam. =Cordia polycephala 

REFERENCES 

Diane N., H. Förther & H.H. Hilger. 2002. A systematic analysis of Heliotropium, Tournefortia, and allied taxa of the 

Heliotropiaceae (Boraginales) based on ITS1 sequences and morphological data. Am. J. Bot. 89: 287-295. 

Feuillet, C. 2002. Boraginaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


145-151. 

Johnston, I.M. 1928. Studies in the Boraginaceae VII. 1. The South American species of Heliotropium. Contrib. Gray Herb. 81: 

3-72. 

Johnston, I.M. 1935. Studies in Boraginaceae X. The Boraginaceae of northeastern South America. J. Arn. Arb. 16: 1-64. 

Johnston, I.M. 1936. Boraginaceae. In A. Pulle, Flora of Suriname 4(1): 306-333. 

Miller, J.S., J. Gaviria, R. Gómez, & G. Rodríguez. 1997. Boraginaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of 


Uittien, H. 1937. Boraginaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 496-497. 

Brassica juncea (L.) Czern. & Coss. [introduced] 

Cardamine flexuosa With. 

Coronopus didymus (L.) Sm. [probably introduced] 

BRASSICACEAE 

J.C. Lindeman, L.J. Raechal 

BO 

FG 

FG


Lepidium virginicum L. [introduced] 

Rorippa dubia (Pers.) H. Hara [introduced] [=Rorippa indica (L.) Hiern] 

Rorippa nasturtium-aquaticum (L.) Hayek [introduced] [=Nasturtium 

officinale R. Br.] 

Rorippa palustris (L.) Besser [introduced] [=Rorippa islandica (Leder) 

Borbás] 

Sinapis arvensis L. [introduced] 


Lepidium micropterum Miq. =Lepidium virginicum 

Nasturtium palustre (L.) DC., non Crantz =Rorippa palustris 

Rorippa heterophylla (Blume) R.O. Williams =Rorippa dubia 

REFERENCES 

Al-Shehbaz, I. 1988. Cruciferae. In R.A. Howard, Flora of the Lesser Antilles 4: 276-292. 

Mitchell, J.D. 2002 Brassicaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


151. 

Raechal, L.J. 1997. Brassicaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 547-548. 

BRUNELLIACEAE 

J.S. Miller 

Brunellia comocladifolia Bonpl. 

Brunellia comocladifolia subsp. ptariana (Steyerm.) Cuatrec. [Mt. Roraima] 

Brunellia neblinensis Steyerm. & Cuatrec. 


Brunellia integrifolia Szyszyl. var. ptariana Steyerm. =Brunellia comocladifolia subsp. ptariana 

REFERENCES 

Cuatrecasas, J. 1970. Brunelliaceae. Flora Neotropica Monogr. 2: 1-188. 

Miller, J.S. 1997. Brunelliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 676-688. 

Bursera karsteniana Engl. 

Bursera simaruba (L.) Sarg. 

Bursera tomentosa (Jacq.) Triana & Planch. 

Commiphora leptophloeos (Mart.) J.B. Gillett 

Crepidospermum goudotianum (Tul.) Triana & Planch. 

Crepidospermum prancei Daly 

Crepidospermum rhoifolium (Benth.) Triana & Planch. 

Dacryodes belemensis Cuatrec. 

Dacryodes chimantensis Steyerm. & Maguire 

Dacryodes cuspidata (Cuatrec.) Daly 

Dacryodes glabra (Steyerm.) Cuatrec. 

Dacryodes microcarpa Cuatrec. 

Dacryodes negrensis Daly & M.C. Martínez 

Dacryodes nitens Cuatrec. 

Dacryodes peruviana (Loes.) H.J. Lam 

Dacryodes roraimensis Cuatrec. 

Dacryodes sclerophylla Cuatrec. 

Dacryodes sp. A 

Dacryodes steyermarkii Sandwith 

Protium altsonii Sandwith 

Protium amazonicum (Cuatrec.) Daly 

Protium apiculatum Swart 

Protium aracouchini (Aubl.) Marchand 

Protium boomii Daly 

Protium carolense Daly 

Protium crassipetalum Cuatrec. 

Protium crenatum Sandwith 

Protium cuneatum Swart 

BURSERACEAE 

D.C. Daly 

AM 

BO 

BO 

SU FG 

GU SU 

FG 

GU 

SU 

BO 

AM BO GU FG 

BO 

AM BO 

GU FG 

AM 


SU? 

AM BO GU 

AM FG 

AM BO 

AM BO 

AM BO 


AM BO GU FG 

BO GU 

BO 

AM BO 

BO GU 


AM BO FG 

AM SU FG 


BO GU 

AM 

AM BO GU SU? 

AM BO DA GU 

BO GU SU? FG

228 

Protium decandrum (Aubl.) Marchand 

Protium demerarense Swart 

Protium divaricatum Engl. 

Protium divaricatum subsp. divaricatum 

Protium divaricatum subsp. fumarium Daly 

Protium elegans Engl. 

Protium ferrugineum Engl. 

Protium gallicum Daly 

Protium giganteum Engl. 

Protium giganteum var. crassifolium (Engl.) Daly 

Protium giganteum var. giganteum 

Protium glabrescens Swart 

Protium grandifolium Engl. 

Protium guianense (Aubl.) Marchand 

Protium heptaphyllum (Aubl.) Marchand 

Protium heptaphyllum subsp. heptaphyllum 

Protium heptaphyllum subsp. ulei (Swart) Daly 

Protium inodorum Daly 

Protium llanorum Cuatrec. 

Protium macrocarpum Cuatrec. 

Protium macrosepalum Swart 

Protium melinonis Engl. 

Protium morii Daly 

Protium nitidifolium (Cuatrec.) Daly 

Protium nodulosum Swart 

Protium occultum Daly 

Protium opacum Swart 

Protium opacum subsp. exaggeratum Daly 

Protium opacum subsp. opacum 

Protium opacum subsp. rabelianum Daly 

Protium pallidum Cuatrec. 

Protium paniculatum Engl. 

Protium paniculatum var. A 

Protium paniculatum var. paniculatum 

Protium paniculatum var. riedelianum (Engl.) Daly 

Protium pilosum (Cuatrec.) Daly 

Protium plagiocarpium Benoist 

Protium poeppigianum Swart 

Protium polybotryum (Turcz.) Engl. 

Protium polybotryum subsp. blackii (Swart) Daly 

Protium polybotryum subsp. polybotryum 

Protium ptarianum Steyerm. 

Protium pullei Swart [unverified name] 

Protium reticulatum (Engl.) Engl. 

Protium robustum (Swart) D.M. Porter 

Protium sagotianum Marchand 

Protium sp. A 

Protium sp. B 

Protium sp. C 

Protium sp. D 

Protium sp. E 

Protium sp. F 

Protium spruceanum (Benth.) Engl. 

Protium strumosum Daly 

Protium subserratum (Engl.) Engl. 

Protium subserratum subsp. A 

Protium subserratum subsp. subserratum 

Protium tenuifolium (Engl.) Engl. 

Protium trifoliolatum Engl. 

Protium unifoliolatum Spruce ex Engl. 

Tetragastris altissima (Aubl.) Swart 

Tetragastris hostmannii (Engl.) Kuntze 

Tetragastris panamensis (Engl.) Kuntze 

Trattinnickia boliviana (Swart) Daly 



GU FG 

AM BO FG 

AM BO FG 

FG 

AM FG? 

AM BO 

FG 

AM GU SU FG 

SU FG 

AM GU SU FG 

SU? 

AM BO FG 




AM BO 

FG 

AM BO GU 

AM BO 

AM 

FG 

FG 

SU 

FG 

FG 

AM BO GU FG 

AM BO GU? 

AM BO GU 

GU FG 

FG 

AM 

AM 

AM 

AM 

FG 

GU? SU? FG 

FG 

GU SU FG 

GU 

GU SU FG 

AM BO 

SU 

AM 

GU SU FG 


AM BO 

AM BO 

BO GU 

AM 

BO 

AM 

AM BO 

GU FG 

AM GU SU FG 

AM 

AM SU FG 



AM BO GU 


BO GU SU FG 


AM BO FG?

Trattinnickia burserifolia Mart. 

Trattinnickia demerarae Sandwith 

Trattinnickia glaziovii Swart 

Trattinnickia rhoifolia Willd. 



Bursera aracouchili (Aubl.) Baill. =Protium aracouchini 

Bursera bonairensis Bold. =Bursera simaruba 

Bursera caudata Turcz. [nom. confus.] =Protium sagotianum 

Bursera decandra (Aubl.) Baill. =Protium decandrum 

Bursera gummifera L. =Bursera simaruba 

Bursera integerrima (Tul.) Triana & Planch. =Bursera simaruba 

Bursera leptophloeos Mart. =Commiphora leptophloeos 

Bursera martiana Engl. =Commiphora leptophloeos 

Bursera orinocensis Engl. =Commiphora leptophloeos 

Bursera subpubescens (Rose) Engl. =Bursera simaruba 

Bursera trinitensis R.O. Williams =Bursera karsteniana 

Bursera velutinifolia R.S. Cowan (Anacardiaceae) =Cyrtocarpa velutinifolia 

Burseria gummifera (L.) Jacq. =Bursera simaruba 

Dacryodes microcarpa var. lanceolata Cuatrec. =Dacryodes microcarpa 

Dacryodes peruviana var. caroniensis Cuatrec. =Dacryodes peruviana 

Elaphrium decandrum (Aubl.) Spreng. ex D. Dietr. =Protium decandrum 

Elaphrium enneandrum Spreng. ex D. Dietr. =Protium decandrum 

Elaphrium gujanense (Aubl.) Spreng. ex D. Dietr. =Protium guianense 

Elaphrium heptaphyllum (Aubl.) Spreng. ex D. Dietr. =Protium heptaphyllum 

Elaphrium heterophyllum (Willd.) Spreng. ex D. Dietr. =Protium aracouchini 

Elaphrium integerrimum Tul. =Bursera simaruba 

Elaphrium jacquinianum Kunth =Bursera tomentosa 

Elaphrium simaruba (L.) Rose =Bursera simaruba 

Elaphrium subpubescens Rose =Bursera simaruba 

Elaphrium tomentosum Jacq. =Bursera tomentosa 

Elaphrium trinitense (R.O. Williams) Rose =Bursera karsteniana 

Hedwigia hostmannii Engl. =Tetragastris hostmannii 

Hedwigia panamensis Engl. =Tetragastris panamensis 

Hedwigia rhoifolia Benth. =Crepidospermum rhoifolium 

Hedwigia rhoifolia (Willd.) Baill. =Trattinnickia rhoifolia 

Hemicrepidospermum rhoifolium (Benth.) Swart =Crepidospermum rhoifolium 

Icica altissima Aubl. =Tetragastris altissima 

Icica aracouchini Aubl. =Protium aracouchini 

Icica decandra Aubl. =Protium decandrum 

Icica enneandra Aubl. =Protium decandrum 

Icica guianensis Aubl. =Protium guianense 

Icica heptaphylla Aubl. =Protium heptaphyllum 

Icica heterophylla Willd. [nom. illeg.] =Protium aracouchini 

Icica hostmanni Miq. =Protium guianense 

Icica insignis Triana & Planch. =Protium sagotianum 

Icica paniculata (Engl.) Villafranca =Protium paniculatum 

Icica polybotrya Turcz. =Protium polybotryum 

Icica pubescens Benth. =Protium unifoliolatum 

Icica spruceana Benth. =Protium spruceanum 

Icica surinamensis Miq. =Protium heptaphyllum 

Icica viridiflora Lam. [nom. illeg.] =Protium guianense 

Icicopsis caudata (Turcz.) Engl. =Protium sagotianum 

Icicopsis ferruginea (Engl.) Engl. =Protium ferrugineum 

Icicopsis insignis (Triana & Planch.) Engl. =Protium sagotianum 

Icicopsis reticulata Engl. =Protium reticulatum 

Icicopsis subserrata Engl. =Protium subserratum 

Icicopsis tenuifolia Engl. =Protium tenuifolium 

Pachylobus peruviana Loes. =Dacryodes peruviana 

Paraprotium amazonicum Cuatrec. =Protium amazonicum 

Paraprotium ferrugineum (Engl.) Cuatrec. =Protium ferrugineum 

Pistacia simaruba L. =Bursera simaruba 

Protium almecega Marchand =Protium spruceanum 

Protium almecega var. ulei Swart =Protium heptaphyllum subsp. ulei 

Protium angustifolium Swart =Protium heptaphyllum 


GU SU FG 

FG 

BO DA GU SU FG

230 


Protium aracouchini var. angustifolium Swart =Protium aracouchini 

Protium araguense Cuatrec. [not in flora area] 

Protium benthamii Swart =Protium unifoliolatum 

Protium crassifolium Engl. =Protium giganteum var. crassifolium 

Protium fimbriatum Swart =Protium amazonicum 

Protium firmum Swart =Protium apiculatum 

Protium heptaphyllum var. floribundum Swart =Protium heptaphyllum 

Protium heptaphyllum var. multiflorum (Engl.) Swart =Protium heptaphyllum 

Protium heptaphyllum var. puberulum Engl. =Protium guianense 

Protium heptaphyllum var. surinamense (Miq.) Swart =Protium heptaphyllum 

Protium heptaphyllum var. unifoliolatum Swart =Protium heptaphyllum 

Protium hostmannii (Miq.) Engl. =Protium guianense 

Protium hostmannii var. brasiliensis Swart =Protium heptaphyllum 

Protium insigne (Triana & Planch.) Engl. =Protium sagotianum 

Protium joannis Cuatrec. =Protium trifoliolatum 

Protium llewelyni J.F. Macbr. =Protium grandifolium 

Protium martianum Engl. =Protium trifoliolatum 

Protium multiflorum Engl. =Protium heptaphyllum 

Protium neglectum Swart =Protium tenuifolium 

Protium nitidum Engl. =Protium paniculatum var. riedelianum 

Protium octandrum Swart =Protium heptaphyllum 

Protium orinocense Rusby =Protium decandrum 

Protium paniculatum var. pentamerum Swart =Protium paniculatum var. paniculatum 

Protium paraense auct. non Swart 1966: sensu Cuatrec. =Protium altsonii 

Protium pauciflorum Swart =Protium trifoliolatum 

Protium puberulenta Steyerm. =Protium altsonii 

Protium pubescens (Benth.) Engl. [nom. illeg.] =Protium unifoliolatum 

Protium riedelianum Engl. =Protium paniculatum var. riedelianum 

Protium schomburgkianum Engl. =Protium decandrum 

Protium titubans J.F. Macbr. =Protium trifoliolatum 

Protium unifoliolatum var. inconforme Swart =Protium unifoliolatum 

Protium unifoliolatum var. macrophyllum Huber =Protium unifoliolatum 

Protium unifoliolatum var. puberulum Hoehne =Protium unifoliolatum 

Protium unifoliolatum var. subserratum Engl. =Protium unifoliolatum 

Tetragastris panamensis var. grandifolia Swart =Tetragastris panamensis 

Tetragastris panamensis var. hirtella Swart =Tetragastris panamensis 

Tetragastris paraensis Cuatrec. =Tetragastris panamensis 

Tetragastris phanerosepala Sandwith =Tetragastris altissima 

Tetragastris pilosa Cuatrec. =Protium pilosum 

Tetragastris stevensonii Standl. =Tetragastris panamensis 

Tetragastris trifoliolata (Engl.) Cuatrec. =Protium trifoliolatum 

Tetragastris unifoliolata (Engl.) Cuatrec. =Protium unifoliolatum 

Tingulonga almecega (Marchand) Kuntze =Protium spruceanum 

Tingulonga aracuchini (Aubl.) Kuntze =Protium aracouchini 

Tingulonga caudata (Turcz.) Kuntze =Protium sagotianum 

Tingulonga divaricata (Engl.) Kuntze =Protium divaricatum 

Tingulonga enneandra (Aubl.) Kuntze =Protium decandrum 

Tingulonga ferruginea (Engl.) Kuntze =Protium ferrugineum 

Tingulonga grandifolia (Engl.) Kuntze =Protium grandifolium 

Tingulonga guianensis (Aubl.) Kuntze =Protium guianense 

Tingulonga heptaphylla (Aubl.) Kuntze =Protium heptaphyllum 

Tingulonga hostmanni (Miq.) Kuntze =Protium guianense 

Tingulonga insignis (Triana & Planch.) Kuntze =Protium sagotianum 

Tingulonga martiana (Engl.) Kuntze =Protium trifoliolatum 

Tingulonga multiflora (Engl.) Kuntze =Protium heptaphyllum 

Tingulonga nitida (Engl.) Kuntze =Protium paniculatum var. riedelianum 

Tingulonga paniculata (Engl.) Kuntze =Protium paniculatum 

Tingulonga riedeliana (Engl.) Kuntze =Protium paniculatum var. riedelianum 

Tingulonga simplicifolia Kuntze =Protium unifoliolatum 

Tingulonga spruceana (Benth.) Kuntze =Protium spruceanum 

Tingulonga subserrata (Engl.) Kuntze =Protium subserratum 

Tingulonga tenuifolia (Engl.) Kuntze =Protium tenuifolium 

Tingulonga trifoliolata Kuntze =Protium trifoliolatum 

Trattinnickia burserifolia subsp. obtusa Engl. =Trattinnickia burserifolia


Trattinnickia burserifolia subsp. quinquejuga Engl. =Trattinnickia burserifolia 

Trattinnickia cuspidata Cuatrec. =Dacryodes cuspidata 

Trattinnickia glabra Steyerm. =Dacryodes glabra 

Trattinnickia guianensis Klotzsch [nomen nudum] =Trattinnickia burserifolia 

Trattinnickia lawrancei var. boliviana Swart =Trattinnickia boliviana 

Trattinnickia rhoifolia var. lancifolia Cuatrec. =Trattinnickia boliviana 

Trattinnickia rhoifolia subsp. sprucei Engl. =Trattinnickia rhoifolia 

Trattinnickia rhoifolia subsp. willdenowii Engl. [nom. illeg.] =Trattinnickia rhoifolia 

Trattinnickia schomburgkii Klotzsch [nomen nudum] =Trattinnickia burserifolia 

Trattinnickia subchoripetala Swart =Trattinnickia burserifolia 

REFERENCES 

Daly, D. 1987. Studies in neotropical Burseraceae I. A synopsis of the genus Crepidospermum. Brittonia 39: 51-58. 

Daly, D. 1989. Studies in neotropical Burseraceae II. Generic limits in New World Protieae and Canarieae. Brittonia 41: 17-27. 

Daly, D. 1992. Studies in neotropical Burseraceae VI. New taxa and combinations in Protium Burm.f. Brittonia 44: 280-299. 

Daly, D.C. 1997. Burseraceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 688-728. 

Daly, D.C. 2002. Burseraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 

2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 151- 

165. 

Swart, J.J. 1942. A monograph of the genus Protium and some allied genera. Rec. Trav. Bot. Neerl. 34: 211-446. 

Swart, J.J. 1951. Burseraceae. In A. Pulle, Flora of Suriname 3(2): 204-251. 

Buxus citrifolia (Willd.) Spreng. 

Tricera citrifolia Willd. =Buxus citrifolia 

BUXACEAE 



REFERENCES 

Yatskievych, K. 1997. Buxaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 729-730. 

Brasenia schreberi J.F. Gmel. [possibly extirpated from GU] 

Cabomba aquatica Aubl. 

Cabomba furcata Schult. & Schult. f. 

Cabomba haynesii Wiersema [expected in GU] 

CABOMBACEAE 

J.H. Wiersema 


Cabomba piauhyensis Gardner =Cabomba furcata 

Cabomba piauhyensis var. albida Fassett =Cabomba haynesii 

Cabomba piauhyensis var. piauhyensis =Cabomba furcata 

Cabomba warmingii Casp. =Cabomba furcata 

REFERENCES 

Cramer, J. 1979. Cabombaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 385-389. 

Fassett, N.C. 1953. A monograph of Cabomba. Castanea 18: 116-128. 

Oergaard, M., H.W.E. van Bruggen, & P.J. van der Vlugt. 1992. Die Familie Cabombaceae (Cabomba und Brasenia). Aqua- 

Planta sonderheft nr. 3. 

Wiersema, J.H. 1989. A new species of Cabomba from tropical America. Ann. Missouri Bot. Gard. 76(4): 1167-1168. 

Wiersema, J.H. 1997. Cabombaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 730-732. 

CACTACEAE 

B.E. Leuenberger, B. Trujillo 

Acanthocereus tetragonus (L.) Hummelinck 

Cereus hexagonus (L.) Mill. 

Cereus repandus (L.) Mill., non Haw. 1812 

Disocactus amazonicus (K. Schum.) D.R. Hunt [expected in Venezuelan 

Guayana] 

Epiphyllum phyllanthus (L.) Haw. 

Hylocereus lemairei (Hook.) Britton & Rose 

BO 

GU? 



BO 

BO 


BO 


BO DA GU? SU? FG?

232 


Hylocereus scandens (Salm-Dyck) Backeb. 

Melocactus matanzanus León [cultivated] 

Melocactus mazelianus Ríha 

Melocactus neryi K. Schum. 

Melocactus smithii (Alexander) Buining ex G.D. Rowley 

Opuntia cochenillifera (L.) Mill. [cultivated only?] 

Opuntia elatior Mill. 

Pereskia aculeata Mill. [widely cultivated but apparently native in Guianas] 

Pereskia guamacho F.A.C. Weber 

Pilosocereus oligolepis (Vaupel) Byles & G.D. Rowley [described as having 5 

ribs] 

Rhipsalis baccifera (J.S. Muell.) Stearn 

Rhipsalis floccosa Pfeiff. 

Rhipsalis oblonga Loefgr. [possibly confused with R. occidentalis] 

Rhipsalis occidentalis Barthlott & Rauh [possibly confused with R. oblonga] 

Rhipsalis pachyptera Pfeiff. 

Selenicereus extensus (Salm-Dyck ex DC.) Leuenb. 

Selenicereus megalanthus (K. Schum. ex Vaupel) Moran [known for AM only 

by vegetative plant (#4528) at Jardín Botánico Universitario de Maracay] 

Selenicereus sp. [sect. Salmdyckia][expected in AM & BO] 

Strophocactus wittii (K. Schum.) Britton & Rose 


Acanthocereus colombianus Britton & Rose =Acanthocereus tetragonus 

Acanthocereus pentagonus (L.) Britton & Rose =Acanthocereus tetragonus 

Acanthocereus pitajaya (Jacq.) Dugand ex Croizat =Acanthocereus tetragonus 

Cactus caripensis Kunth =Rhipsalis baccifera 

Cactus hexagonus L. =Cereus hexagonus 

Cactus neryi (K. Schum.) Britton & Rose =Melocactus neryi 

Cactus pentagonus L. =Acanthocereus tetragonus 

Cactus pereskia L. =Pereskia aculeata 

Cactus phyllanthus L. =Epiphyllum phyllanthus 

Cactus pitajaya Jacq. =Acanthocereus tetragonus 

Cactus repandus L. =Cereus repandus 

Cactus smithii Alexander =Melocactus smithii 

Cactus tetragonus L. =Acanthocereus tetragonus 

Cephalocereus kanukuensis Alexander =Pilosocereus oligolepis 

Cereus atroviridis Backeb. =Cereus repandus 

Cereus caripensis (Kunth) DC. =Rhipsalis baccifera 

Cereus curtisii Otto [Guianan records dubious] 

Cereus grandiflorus Mill. [Guianan records dubious] 

Cereus grenadensis Britton & Rose =Cereus repandus 

Cereus jamacaru DC. [Guianan records dubious] 

Cereus lemairei Hook. =Hylocereus lemairei 

Cereus lepidotus Salm-Dyck =Cereus hexagonus 

Cereus longiflorus Alexander =Cereus hexagonus 

Cereus margaritensis J.R. Johnst. =Cereus repandus 

Cereus megalanthus K. Schum. ex Vaupel =Selenicereus megalanthus 

Cereus peruvianus auct. non (L.) Mill.[Guianan records dubious] =Cereus uruguayanus 

Cereus pitajaya (Jacq.) DC. =Acanthocereus tetragonus 

Cereus polyrhizus F.A.C. Weber =Hylocereus lemairei 

Cereus triangularis (L.) Haw. [Guianan records dubious] =Hylocereus triangularis 

Cereus trinitatensis Lem. & Herment =Hylocereus lemairei 

Cereus wittii K. Schum. =Strophocactus wittii 

Epiphyllum hookeri Haw. =Epiphyllum phyllanthus 

Epiphyllum phyllanthus var. hookeri (Haw.) Kimnach =Epiphyllum phyllanthus 

Hylocereus polyrhizus (F.A.C. Weber) Britton & Rose =Hylocereus lemairei 

Hylocereus triangularis (L.) Britton & Rose [Guianan records dubious] 

Hylocereus trinitatensis (Lem. & Herment) A. Berger =Hylocereus lemairei 

Hylocereus venezuelensis Britton & Rose =Hylocereus lemairei 

Mediocactus megalanthus (K. Schum. ex Vaupel) Britton & Rose =Selenicereus megalanthus 

Melocactus guaricensis Croizat =Melocactus neryi 

Melocactus schulzianus Buining & Brederoo =Melocactus neryi 

Nopalea cochenillifera (L.) Salm-Dyck =Opuntia cochenillifera 

Pereskia colombiana Britton & Rose =Pereskia guamacho 

GU SU 

AM 

AM BO 

AM BO GU SU 

GU 

GU SU FG 

BO 

BO GU SU FG 

BO 

BO GU SU FG 


SU 

SU? 

SU? 

GU? FG? 

FG? 

AM 

AM 

GU? SU? FG


Phyllocactus phyllanthus (DC.) Link =Epiphyllum phyllanthus 

Pilocereus repandus (L.) K. Schum. =Cereus repandus 

Pilosocereus kanukuensis (Alexander) Leuenb. [described as having 4 ribs, illustrations in FVG contradictory] =Pilosocereus 

oligolepis 

Pilosocereus oligolepis subsp. kanukuensis (Alexander) P.J. Braun & Esteves =Pilosocereus oligolepis 

Rhipsalis cassutha Gaertn. =Rhipsalis baccifera 

Rhipsalis minutiflora K. Schum. =Rhipsalis baccifera 

Rhodocactus colombianus (Britton & Rose) F.M. Knuth =Pereskia guamacho 

Rhodocactus guamacho (F.A.C. Weber) F.M. Knuth =Pereskia guamacho 

Selenicereus wittii (K. Schum.) G.D. Rowley =Strophocactus wittii 

Subpilocereus grenadensis (Britton & Rose) Backeb. =Cereus repandus 

Subpilocereus margaritensis (J.R. Johnst.) Backeb. =Cereus repandus 

Subpilocereus repandus (L.) Backeb. =Cereus repandus 

Wilmattea venezuelensis (Britton & Rose) Croizat =Hylocereus lemairei 

Wittiocactus amazonicus (K. Schum.) Rauschert =Disocactus amazonicus 

REFERENCES 

Hunt, D. 1992. CITES Cactaceae Checklist. Royal Botanic Gardens, Kew. 

Leuenberger, B.E. 1987. A preliminary list of Cactaceae from the Guianas and recommendations for future collecting and 

preparation of specimens. Willdenowia 16: 497-510. 

Leuenberger, B.E. 1989. Lectotypification of Cereus hexagonus (L.) Miller and taxonomic notes on related taxa from South 

America. Bot. Jahrb. Syst. 111(2): 145-164. 

Leuenberger, B.E. 1997. Cactaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 31: 1-63. 

Leuenberger, B.E. 2002. Cactaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


165-166. 

Taylor, N.P. 1991. The genus Melocactus in Central and South America. Bradleya 9: 1-80. 

Trujillo, B. 1997. Cactaceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 732-749. 

CAMPANULACEAE 

T. Lammers, J.L. Luteyn, B.A. Stein 

Burmeistera ramosa E. Wimm. 

Centropogon cornutus (L.) Druce 

Centropogon roraimanus E. Wimm. 

Hippobroma longiflora (L.) G. Don [expected in FG] 

Lobelia aquatica Cham. 

Lobelia fastigiata Kunth 

Siphocampylus reticulatus (Willd. ex Roem. & Schult.) Vatke 

Siphocampylus sp. A 

Siphocampylus tillettii Steyerm. 

Wahlenbergia perrottetii (A. DC.) Thulin [native to Africa and Brazil] 


Centropogon bonplandianus (Willd. ex Roem. & Schult.) C. Presl =Centropogon cornutus 

Centropogon solanifolius Benth. [records based on misidentifications fide B. Stein] 

Centropogon surinamensis (L.) C. Presl =Centropogon cornutus 

Cephalostigma bahiense A. DC. =Wahlenbergia perrottetii 

Isotoma longiflora (L.) C. Presl =Hippobroma longiflora 

Laurentia longiflora (L.) Endl. =Hippobroma longiflora 

Lobelia bonplandiana Willd. ex Roem. & Schult. =Centropogon cornutus 

Lobelia cornuta L. =Centropogon cornutus 

Lobelia longiflora L. =Hippobroma longiflora 

Lobelia reticulata Willd. ex Roem. & Schult. =Siphocampylus reticulatus 

Lobelia surinamensis L. =Centropogon cornutus 

Siphocampylus tepuiensis Gleason =Siphocampylus reticulatus 

AM 


BO GU 



BO 

AM BO 

AM BO 

AM 

FG 

REFERENCES (incl. Lobeliaceae. See also Sphenocleaceae) 

Gracie, C.A. 2002. Campanulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


183-184. 

Lanjouw, J. 1934. Campanulaceae. In A. Pulle, Flora of Suriname 4(1): 302-305. 

Lanjouw, J. 1937. Campanulaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 494-495. 

Stein, B.A. 1998. Campanulaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 122-129.

234 

Cinnamodendron tenuifolium Uittien 


CANELLACEAE 

REFERENCES 

Brown, J.L. & S.A. Mori. 2002. Canellaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 184-186. 

Uittien, H. 1935. Canellaceae. In A. Pulle, Flora of Suriname 3(1): 304-305. 

Wilson, T.K. 1960. The comparative morphology of the Canellaceae. I. Synopsis of genera and wood anatomy. Trop. Woods 112: 

1-27. 

CAPPARACEAE 

H.H. Iltis, J.C. Lindeman, T. Ruiz-Zapata 

Capparis amplissima Lam. 

Capparis detonsa Triana & Planch. 

Capparis flexuosa (L.) L. 

Capparis flexuosa subsp. polyantha (Triana & Planch.) H.H. Iltis 

Capparis frondosa Jacq. [=Capparis baducca L.?] 

Capparis grandipetala Maguire & Steyerm. [Serranía de la Neblina , expected 

in AM] 

Capparis guaguaensis Steyerm. [expected in GU] 

Capparis hastata Jacq. 

Capparis hastata f. coccolobifolia (Mart. ex Eichler) H.H. Iltis 

Capparis leprieurii Briq. 

Capparis maroniensis Benoist 

Capparis muco H.H. Iltis, L. Cumana, R. Delgado & Aymard 

Capparis odoratissima Jacq. 

Capparis osmantha Diels 

Capparis pulcherrima Jacq. 

Capparis sola J.F. Macbr. 

Capparis sp. A 

Capparis verrucosa Jacq. 

Capparis yco (Mart.) Mart. & Eichler 

Capparis yco var. guianensis Sagot 

Cleome aculeata L. 

Cleome frutescens Aubl. [=Cleome spinosa?] 

Cleome guianensis Aubl. 

Cleome gynandra L. [naturalized] 

Cleome latifolia Vahl ex DC. 

Cleome parviflora Kunth 

Cleome pilosa Benth. 

Cleome rutidosperma DC. [naturalized] 

Cleome serrata Jacq. 

Cleome speciosa Raf. 

Cleome spinosa Jacq. 

Cleome stenophylla Klotzsch ex Urb. 

Cleome viridiflora Schreb. 

Crateva tapia L. 

Morisonia americana L. 

Steriphoma ellipticum (DC.) Spreng. 

BO DA GU? SU 

BO 

BO SU 

BO SU 


Capparidastrum baducca (L.) Hutch. =Capparis frondosa 

Capparis baducca L. [Capparis frondosa Jacq. synonymized here by some authors] =Capparis frondosa 

Capparis cynophallophora L. =Capparis flexuosa 

Capparis cynophallophora var. isthmensis (Eichler) H.H. Iltis ex Jans.-Jac. [nomen nudum] =Capparis flexuosa 

Capparis flexuosa f. hastata (Jacq.) Dugand =Capparis hastata 

Capparis guaguaensis Steyerm. =Capparis osmantha 

Capparis isthmensis Eichler =Capparis flexuosa 

Capparis macrophylla auct. non Kunth 1821. =Capparis osmantha 

Capparis montana (Aubl.) Lemée (Sapotaceae) [fide Lindeman] =Pradosia ptychandra 

SU FG 


BO 

BO 

FG 

GU SU FG 

BO GU SU FG 

BO 

BO 

BO DA 

FG? 

AM BO SU FG 

AM 

BO GU? 

FG 

FG 

AM GU SU FG 

FG 

BO GU SU FG 

BO SU FG 

BO GU SU FG 


AM 

FG 

GU SU 

GU SU FG 


GU 

AM FG 


GU 

BO


Capparis nitida Ruiz & Pav. ex DC. =Capparis amplissima 

Capparis pendula Triana & Planch. =Capparis amplissima 

Capparis perseifolia Iltis ex Jans.-Jac. [nomen nudum, fide Lindeman] =Capparis leprieurii 

Capparis polyantha Triana & Planch. =Capparis flexuosa subsp. polyantha 

Capparis sola var. longiracemosa Dugand =Capparis sola 

Capparis surinamensis J.C. Went =Capparis maroniensis 

Capparis trinitensis Urb. =Capparis verrucosa 

Cleome amazonica Heilb. =Cleome viridiflora 

Cleome brachypoda Briq. =Cleome parviflora 

Cleome friesii Heilb. =Cleome viridiflora 

Cleome gigantea L. =Cleome viridiflora 

Cleome hostmanni Miq. =Cleome latifolia 

Cleome humboldtii DC. =Cleome parviflora 

Cleome kerberi Briq. =Cleome viridiflora 

Cleome micrantha Desv. ex Ham. [name of uncertain application] =Cleome parviflora 

Cleome monochroma J.F. Macbr. =Cleome viridiflora 

Cleome pentaphylla L. =Cleome gynandra 

Cleome pittieri Briq. =Cleome pilosa 

Cleome polygama L. =Cleome serrata 

Cleome psoraleaefolia DC. =Cleome parviflora 

Cleome pungens Willd. [nom. confus.] =Cleome spinosa 

Cleome surinamensis Miq. =Cleome aculeata 

Crateva acuminata DC. =Crateva tapia 

Crateva benthamii Eichler =Crateva tapia 

Crateva gynandra L. =Crateva tapia 

Gynandropsis gynandra (L.) Briq. =Cleome gynandra 

Gynandropsis pentaphylla (L.) DC. =Cleome gynandra 

Gynandropsis speciosa (Kunth) DC. =Cleome speciosa 

Melidiscus gigantea (L.) Raf. =Cleome viridiflora 

Morisonia elliptica Rusby =Morisonia americana 

Morisonia flexuosa L. =Capparis flexuosa 

Morisonia johnstonii Urb. =Morisonia americana 

Neocalyptrocalyx surinamensis (J.C. Went) Hutch. =Capparis maroniensis 

Physostemon ambiguum Bong. =Cleome guianensis 

Physostemon guianense (Aubl.) Malme =Cleome guianensis 

Physostemon intermedium Moric. =Cleome guianensis 

Physostemon lanceolatum Mart. & Zucc. [fide Lindeman] =Cleome stenophylla 

Quadrella cynophallophora (L.) Hutch. =Capparis flexuosa 

Quadrella odoratissima (Jacq.) Hutch. =Capparis odoratissima 

Sinapistrum giganteum (L.) Moench. =Cleome viridiflora 

Tarenaya spinosa (Jacq.) Raf. =Cleome spinosa 

Voyara montana Aubl. (Sapotaceae) =Pradosia ptychandra 

REFERENCES 

Iltis, H.H. & S.A. Mori. 2002. Capparaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 186-188. 

Jansen-Jacobs, M.J. 1976. Capparaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 512- 

517. 

Ruiz-Zapata, T. & H.H. Iltis. Capparaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 

132-157. 

Went, J.C. 1937. Capparidaceae. In A. Pulle, Flora of Suriname 2(1): 397-405. 

CAPRIFOLIACEAE 

P.E. Berry, N. Ramírez 

Sambucus canadensis L. 

Sambucus canadensis var. laciniata A. Gray 

Sambucus simpsonii Rehder [introduced] 

Viburnum tinoides L. f. 

Viburnum tinoides var. roraimense (Killip & A.C. Sm.) Steyerm. 


Oreinotinus tinoides (L. f.) Oerst. =Viburnum tinoides 

DA 

DA 

BO GU 

BO GU 

SU

236 


Viburnum glabratum Kunth [not in flora area] 

Viburnum leptophyllum Killip & A.C. Sm. =Viburnum tinoides 

Viburnum roraimense Killip & A.C. Sm. =Viburnum tinoides var. roraimense 

Viburnum tinoides var. venezuelense (Killip & A.C. Sm.) Steyerm. [not in flora area - Andes of Venezuela] =Viburnum 

tinoides var. roraimense 

REFERENCES 

Killip, E.P. & A.C. Smith. 1929. The genus Viburnum in northwestern South America. Bull. Torrey Bot. Club 56: 265-274. 

Ramírez, N. & P.E. Berry. 1998. Caprifoliaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 

4: 158-161. 

Steyermark, J.A. 1953. Caprifoliaceae. In J.A. Steyermark & collaborators, Contributions to the Flora of Venezuela. Fieldiana 

Bot. 28(3): 618-619. 

CARICACEAE 

J.S. Miller 

Carica papaya L. [cultivated, naturalized] 

Jacaratia spinosa (Aubl.) A. DC. 

Vasconcellosia microcarpa (Jacq.) A. DC. 

Vasconcellosia microcarpa subsp. heterophylla (Poepp. & Endl.) V.M. Badillo 


Carica microcarpa subsp. heterophylla (Poepp. & Endl.) V.M. Badillo =Vasconcellosia microcarpa subsp. heterophylla 

Carica sativa Tussac =Carica papaya 

Carica spinosa Aubl. =Jacaratia spinosa 

Papaya spinosa A. DC. =Jacaratia spinosa 

REFERENCES 

Badillo, V.M. 1971. Monografia de la familia Caricaceae. Publ. Asoc. Prof., Univ. Central de Venezuela. Maracay. 

Lindeman, J.C. 1984. Caricaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 585-591. 

Miller, J.S. 1998. Caricaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 162-163. 

Mori, S.A. 2002. Caricaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


190. 

Anthodiscus mazarunensis Gilly 

Anthodiscus obovatus Benth. ex Wittm. 

Anthodiscus trifoliatus G. Mey. 

Caryocar glabrum (Aubl.) Pers. 

Caryocar glabrum subsp. album Prance & M.F. Silva 

Caryocar glabrum subsp. glabrum 

Caryocar gracile Wittm. 

Caryocar microcarpum Ducke 

Caryocar montanum Prance 

Caryocar nuciferum L. 

Caryocar pallidum A.C. Sm. 

Caryocar villosum (Aubl.) Pers. 

CARYOCARACEAE 

G.T. Prance 


Caryocar butyrosum (Aubl.) Willd. =Caryocar villosum 

Caryocar coccineum Pilg. =Caryocar glabrum 

Caryocar glabrum var. edule Wittm. =Caryocar microcarpum 

Caryocar glabrum var. pilosum Wittm. =Caryocar microcarpum 

Caryocar krukovii Gilly =Caryocar gracile 

Caryocar riparium A.C. Sm. =Caryocar microcarpum 

Caryocar tessmannii Pilg. =Caryocar glabrum 

Caryocar tomentosum Willd. [nom. illeg.] =Caryocar nuciferum 

Caryocar toxiferum Barb. Rodr. =Caryocar glabrum 

Pekea butyrosa Aubl. =Caryocar villosum 

Pekea tuberculosa Aubl. =Caryocar nuciferum 

Pekea villosa (Aubl.) Poir. =Caryocar villosum 

Rhizobolus butyrosus (Aubl.) J.F. Gmel. =Caryocar villosum 


GU SU FG 

FG 

FG 

AM BO GU SU 

AM BO 

GU 


GU 

GU SU FG 

AM 


BO GU 


AM BO GU FG? 

AM FG


Rhizobolus pekea Gaertn. =Caryocar nuciferum 

Rhizobolus tuberculosus (Aubl.) J.F. Gmel. =Caryocar nuciferum 

Saouari glabra Aubl. =Caryocar glabrum 

Saouari villosa Aubl. =Caryocar villosum 

REFERENCES 

Granville, J.J. de. 2002. Caryocaraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 191-192. 

Lanjouw, J. & P.F. van Heerdt. 1941. Caryocaraceae. In A. Pulle, Flora of Suriname 3(1): 366-372. 

Prance, G.T. 1972. Caryocaraceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot. 

Gard. 23: 1-832. 

Prance, G.T. 1998. Caryocaraceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 164-170. 

Prance, G.T. & M.F. da Silva. 1973. Caryocaraceae. Flora Neotropica Monogr. 12: 1-75. 

CARYOPHYLLACEAE 

G.A. Aymard C., N.L. Cuello A., R.A. DeFilipps† 

Drymaria cordata (L.) Willd. ex Schult. 

Drymaria cordata subsp. cordata 

Polycarpaea corymbosa (L.) Lam. 

Polycarpaea corymbosa var. brasiliensis (Cambess.) Chodat & Hassl. 

Polycarpon apurense Kunth 


Holosteum cordatum L. =Drymaria cordata 

Polycarpaea atherophora Steud. =Polycarpaea corymbosa 

Polycarpaea brasiliensis Cambess. =Polycarpaea corymbosa var. brasiliensis 

REFERENCES 

Aymard C, G.A. & N.L. Cuello A. 1998. Caryophyllaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan 

Guayana 4: 171-174. 

DeFilipps, R.A. & S.L. Maina. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 193. 

Ooststroom, S.J. van. 1934. Caryophyllaceae. In A. Pulle, Flora of Suriname 1(1): 150-153. 

CASUARINACEAE 

Casuarina equisetifolia J.R. Forst. & G. Forst. [Old World family, one species 

widely cultivated] 

REFERENCES 

Berg, C.C. 1992. Casuarinaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 11: 140. 

Cecropia angulata I.W. Bailey 

Cecropia distachya Huber 

Cecropia engleriana Snethl. 

Cecropia ficifolia Warb. ex Snethl. 

Cecropia granvilleana C.C. Berg 

Cecropia kavanayensis Cuatrec. 

Cecropia latiloba Miq. 

Cecropia membranacea Trécul 

Cecropia metensis Cuatrec. 

Cecropia obtusa Trécul 

Cecropia palmata Willd. 

Cecropia peltata L. 

Cecropia sciadophylla Mart. 

Cecropia silvae C.C. Berg 

Coussapoa angustifolia Aubl. 

CECROPIACEAE 

C.C. Berg 





AM BO DA 

BO DA GU 

AM BO FG 

AM 

AM 

SU? FG 

AM BO GU? 


AM 

AM BO GU 

GU SU FG 

SU FG 



SU FG? 

GU SU FG

238 


Coussapoa argentea Akkermans & C.C. Berg 

Coussapoa asperifolia Trécul 

Coussapoa asperifolia subsp. asperifolia 

Coussapoa asperifolia subsp. magnifolia (Trécul) Akkermans & C.C. Berg 

Coussapoa asperifolia subsp. rhamnoides (Standl.) Akkermans & C.C. Berg 

Coussapoa crassivenosa Mildbr. 

Coussapoa ferruginea Trécul 

Coussapoa latifolia Aubl. 

Coussapoa leprieurii Benoist 

Coussapoa microcephala Trécul 

Coussapoa orthoneura Standl. 

Coussapoa parvifolia Standl. 

Coussapoa trinervia Spruce ex Mildbr. 

Coussapoa villosa Poepp. & Endl. 

Coussapoa viridifolia Cuatrec. 

Pourouma acuminata Miq. 

Pourouma bicolor Mart. 

Pourouma bicolor subsp. bicolor 

Pourouma bicolor subsp. digitata (Trécul) C.C. Berg & Heusden 

Pourouma bolivarensis C.C. Berg 

Pourouma cecropiifolia Mart. 

Pourouma cucura Standl. & Cuatrec. 

Pourouma guianensis Aubl. 

Pourouma guianensis subsp. guianensis 

Pourouma melinonii Benoist 

Pourouma melinonii subsp. melinonii 

Pourouma minor Benoist 

Pourouma mollis Trécul 

Pourouma mollis subsp. mollis 

Pourouma ovata Trécul 

Pourouma saulensis C.C. Berg & Kooy 

Pourouma stipulacea C.C. Berg 

Pourouma tomentosa Mart. ex Miq. 

Pourouma tomentosa subsp. apiculata (Spruce ex Benoist) C.C. Berg & 

Heusden 

Pourouma tomentosa subsp. essequiboensis (Standl.) C.C. Berg & Heusden 

Pourouma tomentosa subsp. maroniensis (Benoist) C.C. Berg & Heusden 

Pourouma tomentosa subsp. tomentosa 

Pourouma velutina Mart. ex Miq. 

Pourouma villosa Trécul 


Cecropia auyantepuiana Cuatrec. =Cecropia kavanayensis 

Cecropia bureauiana V.A. Richt. =Cecropia palmata 

Cecropia concolor subsp. engleriana (Snethl.) C.C. Berg & P. Franco =Cecropia engleriana 

Cecropia dielsiana Snethl. =Cecropia peltata 

Cecropia juranyiana V.A. Richt. =Cecropia sciadophylla 

Cecropia orinocensis Standl. =Cecropia latiloba 

Cecropia paraensis Huber =Cecropia latiloba 

Cecropia peltata var. candida Velásquez =Cecropia metensis 

Cecropia riparia Snethl. =Cecropia distachya 

Cecropia sciadophylla var. decurrens Snethl. =Cecropia sciadophylla 

Cecropia sciadophylla var. juranyiana (V.A. Richt.) Snethl. =Cecropia sciadophylla 

Cecropia steyermarkii Cuatrec. =Cecropia kavanayensis 

Cecropia surinamensis Miq. =Cecropia peltata 

Cecropia tessmannii Mildbr. =Cecropia membranacea 

Coussapoa cardonae Pittier =Coussapoa asperifolia subsp. magnifolia 

Coussapoa cayennensis Hawkes =Coussapoa asperifolia subsp. asperifolia 

Coussapoa cornifolia Standl. =Coussapoa parvifolia 

Coussapoa cuneata Miq. =Coussapoa microcephala 

Coussapoa fagifolia Klotzsch =Coussapoa microcephala 

Coussapoa hypochlora Standl. =Coussapoa asperifolia subsp. magnifolia 

Coussapoa krukovii Standl. =Pourouma tomentosa subsp. apiculata 

Coussapoa latifolia var. obovata (Miq.) Miq. =Coussapoa latifolia 

BO GU? 


GU SU FG 

AM BO DA GU 

BO 

AM BO GU? 

SU FG 

GU SU FG 

FG 

GU SU FG 

AM 

AM GU 

AM GU 

AM 

AM BO GU 

BO 



AM GU SU FG 

BO GU? 

AM BO 

AM BO GU SU 






BO GU SU FG 

BO GU SU FG 

BO 

FG 

GU 


AM BO 

GU 

SU FG 

AM 

AM BO SU FG 

GU SU FG


Coussapoa magnifolia Trécul =Coussapoa asperifolia subsp. magnifolia 

Coussapoa microcephala subsp. cornifolia (Standl.) Akkermans & C.C. Berg =Coussapoa parvifolia 

Coussapoa obovata Miq. =Coussapoa latifolia 

Coussapoa prancei C.C. Berg =Coussapoa orthoneura 

Coussapoa rhamnoides Standl. =Coussapoa asperifolia subsp. rhamnoides 

Coussapoa steyermarkii Cuatrec. =Coussapoa crassivenosa 

Coussapoa viridifolia var. tenuifolia Cuatrec. =Coussapoa viridifolia 

Coussapoa williamsii Cuatrec. =Coussapoa orthoneura 

Pourouma acutiflora Trécul =Pourouma guianensis subsp. guianensis 

Pourouma albistipulata Cuatrec. =Pourouma tomentosa subsp. tomentosa 

Pourouma apaporiensis Cuatrec. =Pourouma melinonii subsp. melinonii 

Pourouma apiculata Spruce ex Mildbr. =Pourouma tomentosa subsp. apiculata 

Pourouma apiculata Spruce ex Benoist =Pourouma tomentosa subsp. apiculata 

Pourouma aspera Trécul =Pourouma bicolor subsp. bicolor 

Pourouma aurea Mildbr. =Pourouma minor 

Pourouma camaratana Cuatrec. =Pourouma bicolor subsp. bicolor 

Pourouma cuatrecasasii Standl. =Pourouma minor 

Pourouma digitata Trécul =Pourouma bicolor subsp. digitata 

Pourouma essequiboensis Standl. =Pourouma tomentosa subsp. essequiboensis 

Pourouma garciana Cuatrec. =Pourouma cucura 

Pourouma laevis Benoist =Pourouma villosa 

Pourouma longipendula Ducke =Pourouma ovata 

Pourouma maroniensis Benoist =Pourouma tomentosa subsp. maroniensis 

Pourouma palmata Poepp. & Endl. =Pourouma guianensis subsp. guianensis 

Pourouma populifolia Standl. =Pourouma acuminata 

Pourouma radula Benoist =Pourouma guianensis subsp. guianensis 

Pourouma steyermarkii Standl. & Cuatrec. =Pourouma velutina 

Pourouma umbellifera W.C. Burger =Pourouma minor 

REFERENCES 

Berg, C.C. 1992. Cecropiaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 11: 93-124. 

Berg, C.C. 1998. Cecropiaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 174-190. 

Berg, C.C. 2002. Cecropiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


194-198. 

Berg, C.C., R.W.A.P. Akkermans, & E.C.H. van Heusden. 1990. Cecropiaceae: Coussapoa and Pourouma. Flora Neotropica 

Monogr. 51: 1-194. 

Berg, C.C. & G.P. DeWolf. 1975. Moraceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 265-294. 

CELASTRACEAE 

D.M. Kearns 

Elaeodendron xylocarpum (Vent.) DC. [possibly expected in GU] 

Goupia glabra Aubl. 

Maytenus apiculata Steyerm. 

Maytenus ficiformis Reissek 

Maytenus floribunda Reissek 

Maytenus guyanensis Klotzsch ex Reissek 

Maytenus huberi Steyerm. 

Maytenus insculpta Steyerm. 

Maytenus kanukuensis A.C. Sm. 

Maytenus laevis Reissek 

Maytenus longistipitata Steyerm. 

Maytenus macrocarpa (Ruiz & Pav.) Briq. 

Maytenus myrsinoides Reissek 

Maytenus neblinae Steyerm. [Sierra de la Neblina] 

Maytenus oblongata Reissek 

Maytenus pittieriana Steyerm. 

Maytenus planifolia A.C. Sm. 

Maytenus pruinosa Reissek 

Maytenus pustulata Steyerm. 

Maytenus sp. A 

Maytenus tetragona Griseb. 

Zinowiewia aymardii Steyerm. 

DA 


BO 

AM BO GU 

FG 


AM 

AM 

BO GU 

AM BO 

BO 

BO 


AM 

GU SU FG 

BO 

GU 


AM 

AM 

FG 

BO DA

240 



Cassine xylocarpa Vent. =Elaeodendron xylocarpum 

Celastrus macrocarpos Ruiz & Pav. =Maytenus macrocarpa 

Celastrus myrtifolius L. (Rosaceae) =Prunus myrtifolia 

Goupia paraensis Huber =Goupia glabra 

Goupia tomentosa Aubl. =Goupia glabra 

Maytenus floribunda Pittier, non Reissek 1861 =Maytenus pittieriana 

Maytenus guianensis Klotzsch [nomen nudum] =Maytenus guyanensis 

Maytenus guianensis f. crenulata Steyerm. =Maytenus guyanensis 

Maytenus jauaensis Steyerm. =Maytenus laevis 

Maytenus reissekii Urb. =Maytenus myrsinoides 

REFERENCES 

Kearns, D.M. 1998. Celastraceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 190-197. 

Loesener, T. 1942. Celastraceae. In A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien 20b: 87-197. 

Mitchell, J.D. 2002. Celastraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


199-202. 

Ceratophyllum muricatum Cham. 

Ceratophyllum muricatum subsp. australe (Griseb.) Les 

CERATOPHYLLACEAE 

P.E. Berry 


Ceratophyllum demersum var. cristatum (Spruce) K. Schum. =Ceratophyllum muricatum 

Ceratophyllum lleranae Fassett =Ceratophyllum muricatum 

REFERENCES 

Les, D.H. 1988. The evolution of achene morphology in Ceratophyllum, II. Fruit variation and systematics of the “spiny-margined” 

group. Syst. Bot. 13(1): 73-86. 

Les, D.H. 1989. The evolution of achene morphology in Ceratophyllum, IV. Summary of proposed relationships and evolutionary 

trends. Syst. Bot. 14(2): 254-262. 

Lowden, R.M. 1978. Studies on the submerged genus Ceratophyllum in the Neotropics. Aquatic Botany 4: 127-142. 

Wilmot-Dear, 1985. Ceratophyllum revised, a study in leaf and fruit variation. Kew Bull. 40: 243-271. 

Chenopodium ambrosioides L. [naturalized] 

CHENOPODIACEAE 

R.A. DeFilipps†, D.M. Kearns 

REFERENCES 

Kearns, D.M. 1998. Chenopodiaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 198. 

Lanjouw, J. 1957. Chenopodiaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 291-292. 

Hedyosmum intermedium Todzia 

Hedyosmum neblinae Todzia 

Hedyosmum tepuiense Todzia 

CHLORANTHACEAE 

C.A. Todzia 


Hedyosmum brasiliense Mart. [not in flora area] 

Hedyosmum racemosum auct., non (Ruiz & Pav.) G. Don =Hedyosmum tepuiense 

DA GU SU 

DA 


AM BO GU? 

AM 

AM BO GU 

REFERENCES 

Todzia, C.A. 1988. Chloranthaceae: Hedyosmum. Flora Neotropica Monogr. 48: 1-138. 

Todzia, C.A. 1993. New species of Hedyosmum from northern South America. Novon 3: 81-85. 

Todzia, C.A. 1988. Chloranthaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 198-201.


CHRYSOBALANACEAE 

G.T. Prance 

Acioa guianensis Aubl. 

Acioa schultesii Maguire 

Acioa somnolens Maguire 

Chrysobalanus icaco L. 

Chrysobalanus venezuelanus Prance 

Couepia bracteosa Benth. 

Couepia canescens (Gleason) Prance 

Couepia canomensis (Mart.) Benth. ex Hook. f. 

Couepia caryophylloides Benoist 

Couepia chrysocalyx (Poepp.) Benth. ex Hook. f. 

Couepia cognata (Steud.) Fritsch 

Couepia cognata var. cognata 

Couepia cognata var. major Prance 

Couepia cognata var. membranacea Prance 

Couepia comosa Benth. 

Couepia elata Ducke 

Couepia excelsa Ducke 

Couepia exflexa Fanshawe & Maguire 

Couepia foveolata Prance 

Couepia guianensis Aubl. 

Couepia guianensis subsp. divaricata (Huber) Prance 

Couepia guianensis subsp. glandulosa (Miq.) Prance 

Couepia guianensis subsp. guianensis 

Couepia habrantha Standl. 

Couepia joaquinae Prance 

Couepia magnoliifolia Benth. ex Hook. f. 

Couepia maguirei Prance 

Couepia martinii Prance 

Couepia multiflora Benth. 

Couepia obovata Ducke 

Couepia paraensis (Mart. & Zucc.) Benth. 

Couepia paraensis subsp. glaucescens (Spruce ex Hook. f.) Prance 

Couepia parillo DC. 

Couepia racemosa Benth. ex Hook. f. 

Couepia rankinae Prance 

Couepia sandwithii Prance 

Couepia steyermarkii Maguire 

Exellodendron barbatum (Ducke) Prance 

Exellodendron coriaceum (Benth.) Prance 

Hirtella adderleyi Prance 

Hirtella angustissima Sandwith 

Hirtella araguariensis Prance 

Hirtella bicornis Mart. & Zucc. 

Hirtella bicornis var. bicornis 

Hirtella bicornis var. pubescens Ducke 

Hirtella brachystachya Spruce ex Hook. f. 

Hirtella bullata Benth. 

Hirtella caduca Fanshawe & Maguire 

Hirtella castilloana Prance 

Hirtella ciliata Mart. & Zucc. 

Hirtella confertiflora Prance 

Hirtella cordifolia Prance 

Hirtella couepiiflora Prance 

Hirtella cowanii Prance & Maguire 

Hirtella davisii Sandwith 

Hirtella deflexa Maguire [expected in GU] 

Hirtella duckei Huber 

Hirtella elongata Mart. & Zucc. 

Hirtella eriandra Benth. [expected in the Guianas] 

Hirtella glabrata Pilg. 

Hirtella glandistipula Ducke 

SU FG 

AM 

BO 

BO 

DA GU SU 

FG 

FG 

GU SU FG 

AM BO GU 

AM GU FG 

SU FG 

AM 

BO GU SU 

BO GU 

GU 

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AM GU FG 

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AM BO GU 

AM BO GU 

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AM 

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242 


Hirtella glandulosa Spreng. 

Hirtella guainiae Spruce ex Hook. f. 

Hirtella guyanensis (Fritsch) Sandwith 

Hirtella hispidula Miq. 

Hirtella liesneri Prance 

Hirtella longipedicellata Prance 

Hirtella macrophylla Benth. ex Hook. f. 

Hirtella macrosepala Sandwith 

Hirtella margae Prance 

Hirtella mucronata Prance 

Hirtella obidensis Ducke 

Hirtella orbicularis Prance 

Hirtella paniculata Sw. 

Hirtella physophora Mart. & Zucc. 

Hirtella pimichina Lasser & Maguire 

Hirtella piresii Prance 

Hirtella punctillata Ducke 

Hirtella racemosa Lam. 

Hirtella racemosa var. hexandra (Willd. ex Roem. & Schult.) Prance 

Hirtella racemosa var. hispida Prance 

Hirtella racemosa var. racemosa 

Hirtella scabra Benth. 

Hirtella schultesii Prance 

Hirtella silicea Griseb. 

Hirtella subscandens Spruce ex Hook. f. 

Hirtella suffulta Prance 

Hirtella tentaculata Poepp. 

Hirtella tenuifolia Prance 

Hirtella triandra Sw. 

Hirtella triandra subsp. triandra 

Hirtella ulei Pilg. 

Licania affinis Fritsch 

Licania alba (Bernoulli) Cuatrec. 

Licania albiflora Fanshawe & Maguire 

Licania amapaensis Prance 

Licania apetala (E. Mey.) Fritsch 

Licania apetala var. aperta (Benth.) Prance 

Licania apetala var. apetala 

Licania arachnoidea Fanshawe & Maguire 

Licania boyanii Tutin 

Licania buxifolia Sandwith 

Licania canescens Benoist 

Licania cardiophylla Prance 

Licania caudata Prance 

Licania compacta Fritsch [Mt. Roraima] 

Licania cordata Prance 

Licania coriacea Benth. 

Licania couepifolia Prance 

Licania crassivenia Spruce ex Hook. f. 

Licania cruegeriana Urb. 

Licania cuprea Sandwith 

Licania cyathodes Benoist 

Licania davillifolia Benoist 

Licania densiflora Kleinhoonte 

Licania discolor Pilg. 

Licania divaricata Benth. 

Licania egleri Prance 

Licania elliptica Standl. 

Licania emarginata Spruce ex Hook. f. 

Licania fanshawei Prance 

Licania foldatsii Prance 

Licania foveolata Prance 

Licania furfuracea Prance 

Licania glabriflora Prance 


AM 

BO GU 

AM 

AM 

BO 

BO 

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AM FG 


GU SU 

GU SU 

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GU SU 


AM 

AM 

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BO 

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Licania gracilipes Taub. 

Licania granvillei Prance 

Licania guianensis (Aubl.) Griseb. 

Licania heteromorpha Benth. 

Licania heteromorpha var. glabra (Mart. ex Hook. f.) Prance 

Licania heteromorpha var. heteromorpha 

Licania heteromorpha var. perplexans Sandwith 

Licania heteromorpha var. subcordata Fritsch 

Licania hirsuta Prance 

Licania hispida Prance 

Licania hitchcockii Maguire 

Licania hypoleuca Benth. 

Licania hypoleuca var. hypoleuca 

Licania imbaimadaiensis Prance 

Licania incana Aubl. 

Licania intrapetiolaris Spruce ex Hook. f. 

Licania irwinii Prance 

Licania jimenezii Prance 

Licania kunthiana Hook. f. 

Licania laevigata Prance 

Licania lanceolata Prance 

Licania lasseri Maguire 

Licania lata J.F. Macbr. 

Licania latifolia Benth. ex Hook. f. 

Licania latistipula Prance 

Licania laxiflora Fritsch 

Licania leptostachya Benth. 

Licania leucosepala Griseb. 

Licania licaniiflora (Sagot) S.F. Blake 

Licania longistyla (Hook. f.) Fritsch 

Licania macrophylla Benth. 

Licania majuscula Sagot 

Licania maxima Prance 

Licania membranacea Sagot ex Laness. 

Licania micrantha Miq. 

Licania micrantha subsp. atabapoensis Prance 

Licania micrantha subsp. micrantha 

Licania microphylla Fanshawe & Maguire 

Licania minutiflora (Sagot) Fritsch 

Licania mollis Benth. [US type from Br. Guiana! R. Schomb. 910] 

Licania oblongifolia Standl. 

Licania obtusifolia Fritsch [dubious species] 

Licania occultans Prance [expected] 

Licania octandra (Hoffmanns. ex Roem. & Schult.) Kuntze 

Licania octandra subsp. octandra 

Licania octandra subsp. pallida (Hook. f.) Prance 

Licania orbicularis Spruce ex Hook. f. 

Licania ovalifolia Kleinhoonte 

Licania pakaraimensis Prance 

Licania pallida Spruce ex Sagot 

Licania parviflora Benth. 

Licania parvifolia Huber 

Licania parvifructa Fanshawe & Maguire 

Licania persaudii Fanshawe & Maguire 

Licania piresii Prance 

Licania polita Spruce ex Hook. f. 

Licania prismatocarpa Spruce ex Hook. f. 

Licania pruinosa Benoist 

Licania pyrifolia Griseb. 

Licania reticulata Prance 

Licania robusta Sagot 

Licania rodriguesii Prance 

Licania roraimensis Standl. [Mt. Roraima] 

Licania rufescens Klotzsch ex Fritsch 

AM 

AM GU FG 

DA GU SU FG 


AM BO GU FG 


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GU 


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244 

Licania sandwithii Prance 

Licania savannarum Prance 

Licania silvae Prance 

Licania sp. A 

Licania sprucei (Hook. f.) Fritsch 

Licania steyermarkii Maguire 

Licania stricta Kleinhoonte [dubious species] 

Licania subarachnophylla Cuatrec. 

Licania tepuiensis Prance 

Licania triandra Mart. ex Hook. f. 

Licania urceolaris Hook. f. 

Licania vaupesiana Killip & Cuatrec. 

Licania wurdackii Prance 

Parinari campestris Aubl. 

Parinari excelsa Sabine 

Parinari maguirei Prance 

Parinari montana Aubl., emend. Ducke 

Parinari pachyphylla Rusby 

Parinari parvifolia Sandwith 

Parinari rodolphii Huber 

Parinari sprucei Hook. f. 


GU 

AM 

AM 

AM 

GU SU FG 

AM BO GU FG 

AM BO 

SU 

AM 

BO 

AM 

AM 

AM 

AM 



BO GU 

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AM BO 

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AM 



Balantium cordifolium Desv. ex Ham. =Parinari campestris 

Chrysobalanus icaco var. pellocarpus (G. Mey.) Hook. f. =Chrysobalanus icaco 

Chrysobalanus incanus (Aubl.) M. Gómez =Licania incana 

Chrysobalanus montanus (Aubl.) M. Gómez =Parinari montana 

Chrysobalanus pellocarpus G. Mey. =Chrysobalanus icaco 

Chrysobalanus savannarum Britton =Chrysobalanus icaco 

Chrysobalanus triandra (Sw.) Morales =Hirtella triandra 

Couepia calophlebia Standl. =Couepia racemosa 

Couepia divaricata Huber =Couepia guianensis subsp. divaricata 

Couepia duckei Huber =Couepia paraensis subsp. glaucescens 

Couepia glandulosa Miq. =Couepia guianensis subsp. glandulosa 

Couepia glaucescens Spruce ex Hook. f. =Couepia paraensis subsp. glaucescens 

Couepia leptostachya Benth. ex Hook. f. =Couepia guianensis subsp. guianensis 

Couepia myrtifolia Benth. ex Hook. f. =Couepia guianensis subsp. glandulosa 

Couepia pauciflora Huber =Couepia parillo 

Couepia steudeliana Walp. =Couepia cognata 

Couepia surinamensis Kleinhoonte =Couepia guianensis subsp. guianensis 

Couepia thyrsiflora Hook. f. =Couepia guianensis subsp. glandulosa 

Couepia versicolor Benoist =Couepia guianensis subsp. guianensis 

Couepia villosa Fanshawe & Maguire =Couepia parillo 

Ferolia amazonica Kuntze =Parinari excelsa 

Ferolia coriacea (Benth.) Kuntze =Exellodendron coriaceum 

Ferolia excelsa (Sabine) Kuntze =Parinari excelsa 

Ferolia montana (Aubl.) Kuntze =Parinari montana 

Hirtella aggregata Poir. =Hirtella paniculata 

Hirtella americana Jacq. =Hirtella triandra 

Hirtella americana L. [not in flora area] 

Hirtella americana Aubl. =Hirtella racemosa 

Hirtella americana var. hexandra (Willd. ex Roem. & Schult.) Hook. f. =Hirtella racemosa var. hexandra 

Hirtella amplexicaulis Pilg. =Hirtella bullata 

Hirtella apetala E. Mey. =Licania apetala 

Hirtella aureohirsuta Pilg. =Hirtella bullata 

Hirtella bracteata Mart. & Zucc. =Hirtella triandra 

Hirtella bracteosa Steud. =Hirtella paniculata 

Hirtella canomensis (Mart.) Spreng. =Couepia canomensis 

Hirtella castanea DC. =Hirtella triandra 

Hirtella caudata Kleinhoonte =Hirtella bicornis var. pubescens 

Hirtella cauliflora Huber =Hirtella physophora 

Hirtella cognata Steud. =Couepia cognata 

Hirtella coppenamensis Kleinhoonte =Hirtella obidensis 

Hirtella cotticaensis Kleinhoonte =Hirtella physophora


Hirtella damaziana Beauverd =Hirtella glandulosa 

Hirtella egensis Fritsch =Hirtella hispidula 

Hirtella formicaria Suess. =Hirtella duckei 

Hirtella glaberrima Steud. (Leguminosae-Caesalpinioideae) =Crudia glaberrima 

Hirtella hexandra Willd. ex Roem. & Schult. =Hirtella racemosa var. hexandra 

Hirtella hirsuta Lam. =Hirtella paniculata 

Hirtella hookeri Pilg. =Hirtella glandulosa 

Hirtella indecora Schott =Hirtella paniculata 

Hirtella jamaicensis Urb. =Hirtella triandra 

Hirtella lanceolata Kleinhoonte =Hirtella obidensis 

Hirtella manigera Kleinhoonte =Hirtella silicea 

Hirtella melinonii Benoist =Hirtella silicea 

Hirtella multiflora Urb. =Hirtella triandra 

Hirtella nemorosa Hoffmanns. ex Roem. & Schult. =Hirtella racemosa 

Hirtella octandra Hoffmanns. ex Roem. & Schult. =Licania octandra 

Hirtella paniculata Lam. =Hirtella triandra 

Hirtella peruviana Pers. =Hirtella triandra 

Hirtella praealta Sagot =Hirtella bicornis var. bicornis 

Hirtella racemosa Ruiz & Pav. =Hirtella triandra 

Hirtella rubra Benth. =Hirtella ciliata 

Hirtella scandens Hoffmanns. ex Roem. & Schult. =Hirtella racemosa 

Hirtella stipitadenia Rusby =Hirtella bullata 

Hirtella strigulosa Steud. =Hirtella racemosa 

Hirtella subsetosa Fanshawe & Maguire =Hirtella hispidula 

Hirtella velutina Pilg. =Hirtella glandulosa 

Hirtella violacea Steud. =Hirtella racemosa 

Hirtella wachenheimii Benoist =Hirtella glandulosa 

Licania anisophylla Standl. =Licania micrantha subsp. micrantha 

Licania aperta Benth. =Licania apetala var. aperta 

Licania apetala var. pendula (Benth.) Fritsch =Licania apetala var. apetala 

Licania arachnites Standl. =Licania parvifolia 

Licania aubletiana (Blume) Lemée =Acioa guianensis 

Licania axilliflora (Sagot) Hochr. =Licania leptostachya 

Licania benthamii Hook. f. =Licania heteromorpha var. heteromorpha 

Licania biglandulosa Griseb. ex Urb. =Licania heteromorpha var. heteromorpha 

Licania bothynophylla Mart. =Licania octandra subsp. octandra 

Licania campinensis Huber =Licania licaniiflora 

Licania caracasana Klotzsch ex Hook. f. =Licania apetala var. aperta 

Licania costata Spruce ex Hook. f. =Licania heteromorpha var. glabra 

Licania crassifolia Benth. =Licania incana 

Licania cymosa Fritsch [not in flora area] 

Licania dahlgrenii Standl. =Licania apetala var. apetala 

Licania egensis Fritsch =Licania octandra subsp. pallida 

Licania exiguifolia Standl. =Couepia cognata var. cognata 

Licania floribunda Benth. =Licania apetala var. apetala 

Licania galibica Benoist =Licania membranacea 

Licania glabra Mart. ex Hook. f. =Licania heteromorpha var. glabra 

Licania gracilis Kleinhoonte =Licania laxiflora 

Licania grisea Kleinhoonte =Licania hypoleuca 

Licania helvola Spruce ex Hook. f. =Licania micrantha subsp. micrantha 

Licania heteromorpha var. divaricata (Benth.) Fritsch =Licania divaricata 

Licania hookeri Fritsch =Licania octandra subsp. pallida 

Licania hookeri var. obtusa Huber =Licania octandra subsp. octandra 

Licania hostmannii Fritsch =Licania majuscula 

Licania huberiana Maguire =Licania licaniiflora 

Licania hylaea Cuatrec. =Licania apetala var. apetala 

Licania hypagyraea Malme =Licania kunthiana 

Licania incana var. axilliflora Sagot =Licania leptostachya 

Licania kanukuensis Standl. =Licania densiflora 

Licania kuntzeana Urb. =Licania apetala var. aperta 

Licania laurifolia Huber =Licania polita 

Licania laxa Fanshawe & Maguire =Licania polita 

Licania leptostachya var. axilliflora (Sagot) Fritsch =Licania leptostachya 

Licania leptostachya var. crassifolia (Benth.) Benoist =Licania incana

246 


Licania longifolia Benoist =Licania alba 

Licania lucida J.F. Macbr. =Licania mollis 

Licania micrantha var. triandra (Mart. ex Hook. f.) Fritsch =Licania triandra 

Licania microcarpa Hook. f. =Licania hypoleuca 

Licania obovata Benth. ex Hook. f. =Licania latifolia 

Licania obovatifolia Maguire =Licania licaniiflora 

Licania pachystachya Kleinhoonte =Licania robusta 

Licania pallida (Hook. f.) Kuntze =Licania octandra subsp. pallida 

Licania pallida Spruce ex Hook. f. =Licania pallida 

Licania pallidula Standl. =Licania coriacea 

Licania paniculata Fanshawe & Maguire =Licania elliptica 

Licania parinarioides Huber =Licania licaniiflora 

Licania parviflora (Blume) Lemée, nom. illeg. =Licania obtusifolia 

Licania parviflora var. conduplicata Maguire =Licania hypoleuca 

Licania parviflora var. membranacea Maguire =Licania kunthiana 

Licania parviflora var. pallida Hook. f. =Licania pallida 

Licania parvifolia Pittier =Licania hypoleuca 

Licania pendula Benth. =Licania apetala var. apetala 

Licania petrensis Prance =Licania emarginata 

Licania poeppigii Fritsch =Licania polita 

Licania pubiflora Benth. =Licania apetala var. aperta 

Licania pulchravenia Killip & Cuatrec. =Licania triandra 

Licania riparia (Gleason) Standl. =Licania minutiflora 

Licania rondonii Pilg. =Licania minutiflora 

Licania sclerophylla (Mart. ex Hook. f.) Fritsch [not in flora area] 

Licania stahelii Kleinhoonte =Licania ovalifolia 

Licania stenocarpa Standl. =Licania octandra subsp. pallida 

Licania stipata Fanshawe & Maguire [nomen nudum] 

Licania takutuensis Standl. =Licania octandra subsp. octandra 

Licania turiuva Cham. & Schltdl., nom. illeg. =Licania octandra subsp. octandra 

Licania utilis (Hook. f.) Fritsch =Licania octandra subsp. octandra 

Licania venosa Rusby =Licania alba 

Licania wilson-brownei Maguire =Licania licaniiflora 

Moquilea aubletiana Blume, nom. illeg. =Acioa guianensis 

Moquilea bothynophylla (Mart.) Hook. f. =Licania octandra subsp. octandra 

Moquilea canomensis Mart. =Couepia canomensis 

Moquilea chrysocalyx Poepp. =Couepia chrysocalyx 

Moquilea comosa (Benth.) Walp. =Couepia comosa 

Moquilea couepia Steud. =Couepia guianensis 

Moquilea floribunda (Benth.) Hook. f. =Licania apetala var. apetala 

Moquilea glandulosa (Miq.) Walp. =Couepia guianensis subsp. glandulosa 

Moquilea guianensis Aubl. =Licania guianensis 

Moquilea kuntzeana (Urb.) R.O. Williams =Licania apetala var. aperta 

Moquilea leucosepala (Griseb.) R.O. Williams =Licania leucosepala 

Moquilea licaniiflora Sagot =Licania licaniiflora 

Moquilea longistyla Hook. f. =Licania longistyla 

Moquilea macrocarpa Pittier =Licania pyrifolia 

Moquilea minutiflora Sagot =Licania minutiflora 

Moquilea multiflora (Benth.) Walp. =Couepia multiflora 

Moquilea orinocensis Rusby =Licania apetala var. apetala 

Moquilea pallida Hook. f. =Licania octandra subsp. pallida 

Moquilea paraensis Mart. & Zucc. =Couepia paraensis 

Moquilea pendula (Benth.) Hook. f. =Licania apetala var. apetala 

Moquilea pyrifolia (Griseb.) R.O. Williams =Licania pyrifolia 

Moquilea riparia Gleason =Licania minutiflora 

Moquilea sprucei Hook. f. =Licania sprucei 

Moquilea turiuva (Cham. & Schltdl.) Hook. f. =Licania octandra 

Moquilea utilis Hook. f. =Licania octandra subsp. octandra 

Parinari barbatum Ducke =Exellodendron barbatum 

Parinari brachystachya Benth. =Parinari excelsa 

Parinari canescens Gleason =Couepia canescens 

Parinari coriacea Benth. 

Parinari glazioviana Warm. =Parinari excelsa 

Parinari guyanense Fritsch =Hirtella guyanensis


Parinari guyanensis (Aubl.) Lemée (Moraceae) =Brosimum rubescens 

Parinari hostmannii Fritsch =Couepia cognata 

Parinari krukovii Gleason =Couepia guianensis subsp. divaricata 

Parinari laxiflora Ducke =Parinari excelsa 

Parinari lucidissima Standl. =Parinari rodolphii 

Parinari pilosa Standl. =Couepia canomensis 

Parinari sylvestris Kuhlm. =Parinari excelsa 

Petrocarya campestris (Aubl.) Willd. =Parinari campestris 

Petrocarya excelsa (Sabine) Steud. =Parinari excelsa 

Petrocarya montana (Aubl.) Willd. =Parinari montana 

Salmasia guianensis (Aubl.) J.F. Gmel. =Hirtella racemosa 

Salmasia racemosa Willd. =Hirtella racemosa 

Tachibota guianensis Aubl. =Hirtella racemosa 

REFERENCES 

Kleinhoonte, A. 1939. Rosaceae. In A. Pulle, Flora of Suriname 2(1): 426-456. 

Prance, G.T. 1972. Chrysobalanaceae. Flora Neotropica Monogr. 9: 1-409. 

Prance, G.T. 1981. Notes on Couepia and Hirtella. Brittonia 33: 347-356. 

Prance, G.T. 1982. Chrysobalanaceae. Flora de Venezuela 14(2): 325-487. 

Prance, G.T. 1986a. Chrysobalanaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 2: 1-146. 

Prance, G.T. 1986b. Studies on the flora of the Guianas. 19. New taxa of Chrysobalanaceae for the Flora of the Guianas. Proc. 

Kon. Ned. Akad. Wetensch., ser. C. 89: 111-116. 

Prance, G.T. 1989. Chrysobalanaceae (supplement). Flora Neotropica Monogr. 9S: 1-267. 

Prance, G.T. 1992. New species and new records of neotropical Chrysobalanaceae. Kew Bull. 47: 633-646. 

Prance, G.T. 1998. Chrysobalanaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 202- 

246. 

Prance, G.T. 2002. Chrysobalanaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 202-212. 

Prance, G.T. & A.R.A. Görts-van Rijn. 1976. Chrysobalanaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora 

of Suriname 2(2): 524-555. 

Clethra guyanensis Klotzsch ex Meisn. [Mt. Roraima] 

CLETHRACEAE 


REFERENCES 

Miller, J.S. & P.E. Berry. 1998. Clethraceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 

246-247. 

Sleumer, H. 1967. Monographia Clethracearum. Bot. Jahrb. Syst. 87: 36-175. 

Sleumer, H. 1971. Clethraceae. In T. Lasser, Flora de Venezuela 3(1): 171-183. 

Calophyllum brasiliense Cambess. 

Calophyllum longifolium Willd. 

Calophyllum pachyphyllum Planch. & Triana 

Caraipa ampla Ducke 

Caraipa densifolia Mart. 

Caraipa densifolia subsp. densifolia 

Caraipa grandifolia Mart. 

Caraipa grandifolia subsp. grandifolia 

Caraipa heterocarpa Ducke 

Caraipa llanorum Cuatrec. 

Caraipa llanorum subsp. cordifolia Kubitzki 

Caraipa llanorum subsp. llanorum 

Caraipa longipedicellata Steyerm. 

Caraipa parvielliptica Cuatrec. 

Caraipa parvifolia Aubl. 

Caraipa psilocarpa Kubitzki 

Caraipa punctulata Ducke 

BO GU 

CLUSIACEAE 

P.E. Berry, N.L. Cuello A., B.K. Holst, D.M. Kearns, K. Kubitzki, J.J. Pipoly, 

N.K.B. Robson, P.F. Stevens, A.L. Weitzman 


GU SU 

AM? 

FG 



AM BO GU 

AM BO 

AM 

AM BO GU 

AM 

AM BO 

AM BO 

AM BO 

FG 

BO GU? 

AM BO GU SU FG

248 


Caraipa racemosa Cambess. 

Caraipa richardiana Cambess. 

Caraipa savannarum Kubitzki 

Caraipa sp. A 

Caraipa spuria Barb. Rodr. 

Caraipa tereticaulis Tul. 

Chrysochlamys membranacea Planch. & Triana 

Chrysochlamys nicaraguensis (Oerst., Planch. & Triana) Hemsl. 

Chrysochlamys pauciflora Steyerm. [may be synonymous with 

Chrysochlamys weberbaueri] 

Chrysochlamys weberbaueri Engl. 

Clusia aishaltonensis Pipoly, sp. nov. ined. 

Clusia amabilis Maguire [Section Clusiastrum] 

Clusia amazonica Planch. & Triana [Section Criuvopsis] 

Clusia annularis Maguire [Section Omphalanthera] 

Clusia asymmetrica Pipoly [Section Criuva] 

Clusia aymardii Pipoly 

Clusia brachystyla Maguire 

Clusia brachystyla var. brachystyla 

Clusia brongniartiana Planch. & Triana [dubious species] 

Clusia candelabrum Planch. & Triana 

Clusia capituliflora Pipoly, sp. nov. ined. 

Clusia cardonae Maguire 

Clusia cerroana Steyerm. 

Clusia chiribiquetensis Maguire 

Clusia cochlitheca Maguire 

Clusia columnaris Engl. 

Clusia comans (Meisn.) Pipoly 

Clusia crassifolia Planch. & Triana 

Clusia cuneata Benth. 

Clusia duartei Maguire 

Clusia duidae Gleason 

Clusia fabiolae Pipoly 

Clusia flavida (Benth.) Pipoly 

Clusia flaviflora Engl. 

Clusia fockeana Miq. 

Clusia gaudichaudii Choisy 

Clusia grammadenioides Pipoly 

Clusia grandiflora Splitg. 

Clusia gratula Maguire 

Clusia guayanae Pipoly 

Clusia hammeliana Pipoly 

Clusia hexacarpa Gleason 

Clusia huberi Pipoly [Section Oedematopus] 

Clusia imbricata Steyerm. 

Clusia insignis Mart. [=Clusia grandiflora?] 

Clusia kanukuana Maguire 

Clusia leprantha Mart. [=Clusia purpurea?] 

Clusia lopezii Maguire [Section Retinostemon] 

Clusia lunanthera Maguire 

Clusia macropoda Klotzsch ex Engl. [Section Clusia] 

Clusia maguireana Pipoly [Section Criuva] 

Clusia martiana Engl. [Section Criuvopsis] 

Clusia martinii Sagot [=Clusia palmicida?] 

Clusia melchiori Gleason [Section Criuva] 

Clusia microstemon Planch. & Triana [Section Phloianthera] 

Clusia minor L. [Section Retinostemon] 

Clusia multilineata Pipoly [Section Criuva] 

Clusia mutica Maguire 

Clusia myriandra (Benth.) Planch. & Triana [Section Phloianthera] 

Clusia nemorosa G. Mey. [Section Clusia] 

Clusia obovata (Spruce ex Planch. & Triana) Pipoly 

Clusia octandra (Poepp.) Pipoly 

Clusia opaca Maguire [Section Criuva] 

GU FG 


AM BO GU 

AM 

AM 

AM BO GU 

AM BO GU FG 

GU 

BO 

BO 

BO 

GU 

GU 

AM 

AM 

AM 

GU 

AM 

BO 

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FG 

AM BO DA 

BO 

GU 

GU 

AM BO GU SU 

AM 

AM 

BO DA GU 

AM BO GU SU 

DA FG 

BO GU 

GU FG 

AM 

AM BO 

BO 

GU 


BO GU 

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SU FG 

GU SU 

AM 

AM 

AM 


BO GU 

AM GU 

AM BO 

AM BO 

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GU 

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GU FG? 

AM 

AM 

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AM 

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AM DA GU SU 

AM 

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BO GU 



AM 

AM 

BO GU FG


Clusia pachyphylla Gleason [Section Anandrogyne] 

Clusia palmicida Rich. ex Planch. & Triana [Section Clusia] 

Clusia panapanari (Aubl.) Choisy [Section Phloianthera] 

Clusia parvifolia Maguire [Section Clusiastrum] 

Clusia parvula (Maguire) Pipoly 

Clusia phelpsiae Lasser & Maguire [Section Anandrogyne] 

Clusia phelpsiana Maguire [expected in GU] 

Clusia picta Pipoly, sp. nov. ined. 

Clusia platystigma Eyma 

Clusia ptaritepuiensis (Steyerm.) Pipoly [Section Oedematopus] 

Clusia pusilla Steyerm. 

Clusia pusilla subsp. orinocensis Maguire 

Clusia pusilla subsp. pusilla [Section Clusiastrum] 

Clusia radiata Maguire & K.D. Phelps [Section Criuva] 

Clusia renggerioides Planch. & Triana [Section Phloianthera] 

Clusia robusta Eyma 

Clusia rosea Jacq. [=Clusia major L.?] 

Clusia rotundifolia Gleason [Section Anandrogyne] 

Clusia savannarum Maguire [Section Anandrogyne] 

Clusia schomburgkiana (Planch. & Triana) Benth. ex Engl. [Section 

Omphalanthera] 

Clusia schomburgkii Vesque [Section Clusiastrum] 

Clusia scrobiculata Benoist 

Clusia sipapoana (Maguire) Pipoly [Section Quapoya] 

Clusia sp. A [Section Oedematopus] 

Clusia sp. B [Section Oedematopus] 

Clusia sp. C [Section Oedematopus] 

Clusia sp. D [Section Oedematopus] 

Clusia spathulaefolia Engl. [Section Brachystemon] 

Clusia steyermarkii Maguire [Section Clusiastrum] 

Clusia stylosa Maguire 

Clusia tabulamontana Maguire [Section Clusiastrum] 

Clusia troncosii Maguire [Section uncertain] 

Clusia viscida Engl. [Section Clusia] 

Clusiella axillaris (Engl.) Cuatrec. 

Clusiella elegans Planch. & Triana 

Clusiella sp. A 

Dystovomita sp. 

Haploclathra leiantha (Benth.) Benth. 

Hypericum marahuacanum N. Robson 

Hypericum marahuacanum subsp. chimantaicum N. Robson [expected in GU] 

Hypericum marahuacanum subsp. marahuacanum 

Hypericum roraimense Gleason [Mt. Roraima] 

Hypericum thesiifolium Kunth [expected in GU] 

Lorostemon bombaciflorum Ducke 

Lorostemon colombianus Maguire 

Mahurea exstipulata Benth. 

Mahurea exstipulata subsp. duckei (Huber) Kubitzki 

Mahurea exstipulata subsp. exstipulata 

Mahurea palustris Aubl. 

Marila laxiflora Rusby 

Marila saramaccana Pulle 

Moronobea coccinea Aubl. 

Moronobea intermedia Engl. 

Moronobea jenmanii Engl. 

Moronobea ptaritepuiana Steyerm. 

Moronobea ptaritepuiana subsp. neblinensis Maguire 

Moronobea ptaritepuiana subsp. ptaritepuiana 

Moronobea riparia (Spruce) Planch. & Triana 

Neotatea duidae (Kobuski & Steyerm.) P.F. Stevens & A.L. Weitzman 

Neotatea longifolia (Gleason) Maguire 

Neotatea neblinae Maguire 

Platonia insignis Mart. 

Quapoya scandens Aubl. 

AM BO 



BO GU 

BO 

AM 

BO 

GU 

GU? SU FG 

BO 

AM BO GU SU 

AM BO 

BO GU SU 

AM 

AM GU SU 

GU SU 

BO GU SU FG 

AM 

BO GU 

AM BO GU 

BO GU 

GU SU FG 

AM GU 

AM 

BO 

AM BO 

AM BO GU 

AM 

BO 

SU 

BO GU SU 

AM 

AM GU SU 

AM GU 

GU 

BO GU 

GU 

GU 

AM BO 

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BO 

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AM 


AM GU SU FG 

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AM BO 

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BO GU SU FG 

BO GU 

BO GU FG? 

AM BO GU 

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BO 

AM GU 

AM 

AM 

AM 

AM GU SU FG 

FG

250 


Rheedia benthamiana Planch. & Triana 

Rheedia brasiliensis (Mart.) Planch. & Triana [=Garcinia brasiliensis Mart.] 

Rheedia lateriflora L. [=Garcinia humilis (Vahl) C. Adams] 

Rheedia macrophylla (Mart.) Planch. & Triana [=Garcinia macrophylla Mart.] 

Rheedia madruno (Kunth) Planch. & Triana [=Garcinia madruno (Kunth) 

Hammel] 

Rheedia martinii Maguire [=Garcinia sp.] 

Rheedia virens Planch. & Triana [=Garcinia sp.] 

Symphonia globulifera L. f. 

Thysanostemon fanshawei Maguire 

Thysanostemon pakaraimae Maguire 

Tovomita acutiflora M.S. Barros & G. Mariz 

Tovomita albiflora A.C. Sm. 

Tovomita atropurpurea Steyerm. 

Tovomita brasiliensis (Mart.) Walp. 

Tovomita aff. brasiliensis (Mart.) Walp. 

Tovomita brevistaminea Engl. 

Tovomita calodictyos Sandwith 

Tovomita caloneura A.C. Sm. 

Tovomita carinata Eyma 

Tovomita choisyana Planch. & Triana 

Tovomita clusiifolia G. Don 

Tovomita divaricata Maguire 

Tovomita eggersii Vesque 

Tovomita elliptica Engl. 

Tovomita fanshawei Maguire 

Tovomita aff. glazioviana Engl. 

Tovomita gracilipes Planch. & Triana 

Tovomita grata Sandwith 

Tovomita guianensis Aubl. 

Tovomita longifolia (Rich.) Hochr. 

Tovomita macrophylla (Poepp.) Walp. 

Tovomita melinonis Vesque 

Tovomita obovata Engl. 

Tovomita obscura Sandwith 

Tovomita pyrifolia Planch. & Triana 

Tovomita rubella Spruce ex Planch. & Triana 

Tovomita schomburgkii Planch. & Triana 

Tovomita secunda Poepp. ex Planch. & Triana 

Tovomita sp. A 

Tovomita sp. B 

Tovomita sp. C 

Tovomita sp. D 

Tovomita sp. E 

Tovomita sp. F 

Tovomita spruceana Planch. & Triana 

Tovomita tenuiflora Benth. ex Planch. & Triana 

Tovomita umbellata Benth. 

Tovomita uniflora G. Don 

Tovomita weddelliana Planch. & Triana 

Tovomitidium speciosa (Ducke) Ducke [expected in FG] 

Vismia baccifera (L.) Triana & Planch. 

Vismia baccifera subsp. dealbata (Kunth) Ewan 

Vismia cavalcantii M.E. Berg 

Vismia cayennensis (Jacq.) Pers. 

Vismia confertiflora Spruce ex Reichardt 

Vismia gracilis Hieron. 

Vismia guianensis (Aubl.) Choisy 

Vismia japurensis Reichardt 

Vismia latifolia (Aubl.) Choisy 

Vismia lauriformis (Lam.) Choisy [expected in GU] 

Vismia laxiflora Reichardt 

Vismia macrophylla Kunth 

Vismia ramuliflora Miq. 

GU SU FG 

FG 

GU 



GU SU 

GU 


GU 

GU 

BO GU 

GU 

BO GU 

GU SU FG 

AM 


GU SU 

GU 

AM GU SU FG 

GU SU FG 

FG 

GU 

AM BO GU 

AM 

AM BO GU 

AM 

AM BO GU? 

GU SU FG 

GU FG 


AM GU 

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GU 

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AM 

AM 

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AM 

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AM BO GU 

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AM BO SU? FG? 

SU 


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GU SU FG 


AM BO GU SU 

GU SU FG 

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GU SU FG

Vismia sandwithii Ewan 

Vismia schultesii N. Robson 

Vismia sessilifolia (Aubl.) Choisy 

Vismia sprucei Sprague 

Vismia steyermarkii N. Robson 

Vismia tenuinervia (M.E. Berg) N. Robson 



Androstylium fockeanum (Miq.) Miq. =Clusia fockeana 

Arrudea purpurea Splitg. =Clusia leprantha 

Calophyllum antillanum Britton =Calophyllum brasiliense 

Calophyllum brasiliense var. antillanum (Britton) Standl. =Calophyllum brasiliense 

Calophyllum lucidum Benth. =Calophyllum brasiliense 

Calophyllum madruno Kunth =Rheedia madruno 

Calophyllum piaroanum A. Castillo & C. Gil =Calophyllum brasiliense 

Caopia acuminata (Lam.) Kuntze =Vismia guianensis 

Caopia cayennensis (Jacq.) Kuntze =Vismia cayennensis 

Caopia guianensis (Aubl.) Lyons =Vismia guianensis 

Caopia latifolia (Aubl.) Kuntze =Vismia latifolia 

Caopia laxiflora (Reichardt) Kuntze =Vismia laxiflora 

Caopia sessilifolia (Aubl.) Kuntze =Vismia sessilifolia 

Caraipa angustifolia Aubl. (Chrysobalanaceae) =Couepia sp. 

Caraipa ferruginea Steyerm. =Caraipa punctulata 

Caraipa glabrata Mart. [not in flora area] =Caraipa grandifolia 

Caraipa lacerdaei Barb. Rodr. [not in flora area] =Caraipa grandifolia 

Caraipa latifolia Aubl. (Chrysobalanaceae) =Licania membranacea 

Caraipa laxiflora Benth. =Caraipa densifolia 

Caraipa leiantha Benth. =Haploclathra leiantha 

Caraipa longifolia Aubl. (Chrysobalanaceae) =Licania alba 

Caraipa paraensis Huber [not in flora area] =Caraipa grandifolia 

Caraipa psidiifolia Ducke =Caraipa richardiana 

Caraipa simplicior Sandwith ? =Caraipa tereticaulis 

Caraipa variabilis Cambess. ? =Caraipa densifolia 

Clusia axillaris Engl. =Clusiella axillaris 

Clusia brachystyla var. guaiquinimensis Maguire =Clusia brachystyla 

Clusia colorans Klotzsch ex Engl. =Clusia panapanari 

Clusia columnaris var. magdalenensis Cuatrec. =Clusia columnaris 

Clusia columnaris var. vaupesana Cuatrec. =Clusia columnaris 

Clusia cucullata Klotzsch [nomen nudum] 

Clusia galactodendron Desv. =Clusia minor 

Clusia grandifolia Engl. =Clusia candelabrum 

Clusia hexacarpa var. ptaritepuiana Steyerm. =Clusia hexacarpa 

Clusia insignis var. hoffmannseggiana Engl. =Clusia insignis 

Clusia jenmanii Engl. =Clusia myriandra 

Clusia longifolia Rich. =Tovomita longifolia 

Clusia microphylla Klotzsch ex Engl. =Clusia panapanari 

Clusia oedematopoidea Maguire =Clusia amazonica 

Clusia parvicapsula Vesque =Clusia minor 

Clusia planchoniana Engl. =Clusia schomburgkiana 

Clusia planchoniana var. oblongifolia Vesque =Clusia schomburgkiana 

Clusia purpurea (Splitg.) Engl. =Clusia leprantha 

Clusia pusilla subsp. jauaensis Maguire =Clusia pusilla subsp. pusilla 

Clusia pusilla subsp. pakaraimae Maguire =Clusia pusilla subsp. pusilla 

Clusia reducta Steyerm. =Clusia opaca 

Clusia roraimana Steyerm. =Clusia schomburgkii 

Clusia rosea var. colombiana Cuatrec. =Clusia rosea 

Clusia scariosa Lasser & Maguire =Clusia pachyphylla 

Clusia schomburgkiana var. lakei Maguire =Clusia schomburgkiana 

Clusia sessilis Klotzsch ex Engl. [latter homonym of C. sessilis G. Forst.] =Clusia cerroana 

Clusia spruceana Planch. & Triana =Clusia schomburgkiana 

Clusia stahelii Maguire =Clusia fockeana 

Clusia uleana Engl. =Clusia martiana 

Clusia venosa Jacq. =Clusia minor 

Clusia williamsii Steyerm. =Clusia columnaris 

Clusia wurdackiana Pipoly [nom. superfl.] =Clusia cerroana 

AM GU SU FG 

AM GU? 

BO GU SU FG 

AM 

BO 

AM

252 


Garcinia acuminata (Ruiz & Pav.) Hammel =Rheedia madruno 

Garcinia benthamiana (Planch. & Triana) Pipoly [nomen nudum] =Rheedia benthamiana 

Garcinia brasiliensis Mart. =Rheedia brasiliensis 

Garcinia macrophylla Mart. =Rheedia macrophylla 

Garcinia madruno (Kunth) Hammel =Rheedia madruno 

Havetia flavida Benth. =Clusia flavida 

Havetia octandra Poepp. =Clusia octandra 

Havetiopsis flavida (Benth.) Planch. & Triana =Clusia flavida 

Havetiopsis flexilis Spruce ex Planch. & Triana =Clusia flavida 

Hypericum acuminatum Lam. =Vismia guianensis 

Hypericum bacciferum L. =Vismia baccifera 

Hypericum cayennensis Jacq. =Vismia cayennensis 

Hypericum guianense Aubl. =Vismia guianensis 

Hypericum latifolium Aubl. =Vismia latifolia 

Hypericum lauriforme Lam. =Vismia lauriformis 

Hypericum reticulatum Poir. =Vismia macrophylla 

Hypericum rufescens Lam. =Vismia sessilifolia 

Hypericum sessilifolium Aubl. =Vismia sessilifolia 

Hypericum silenoides auct. non Juss. 1789. =Hypericum thesiifolium 

Hypericum uliginosum Kunth =Hypericum thesiifolium 

Mahurea casiquiarensis Benth. =Mahurea exstipulata 

Mahurea duckei Huber =Mahurea exstipulata subsp. duckei 

Mahurea linguiformis Tul. =Mahurea exstipulata subsp. exstipulata 

Mahurea sororopantepuiana Steyerm. =Mahurea exstipulata subsp. exstipulata 

Mahurea tomentosa Ducke =Mahurea exstipulata subsp. duckei 

Marialvaea brasiliensis Mart. =Tovomita brasiliensis 

Marialvaea macrophylla Poepp. =Tovomita macrophylla 

Moronobea jenmanii var. fanshawei Maguire =Moronobea jenmanii 

Moronobea jenmanii var. jenmanii =Moronobea jenmanii 

Moronobea ptaritepuiana f. rosea Steyerm. =Moronobea ptaritepuiana 

Moronobea riparia var. fimbrillata R.E. Schult. =Moronobea riparia 

Moronobea riparia var. piraparanensis R.E. Schult. =Moronobea riparia 

Oedematopus duidae Gleason =Clusia huberi 

Oedematopus obovatus Spruce ex Planch. & Triana =Clusia obovata 

Oedematopus octandrus (Poepp.) Planch. & Triana =Clusia octandra 

Oedematopus parvulus Maguire =Clusia parvula 

Oedematopus ptaritepuiensis Steyerm. =Clusia ptaritepuiensis 

Oedematopus quadratus Maguire =Clusia obovata 

Polythecandra schomburgkiana Planch. & Triana =Clusia schomburgkiana 

Quapoya acuminata (Planch. & Triana) Kuntze =Clusia hammeliana 

Quapoya bracteolata Sandwith =Clusia hammeliana 

Quapoya panapanari Aubl. =Clusia panapanari 

Quapoya peruviana (Poepp.) Kuntze =Clusia hammeliana 

Quapoya peruviana var. guayanensis Maguire =Clusia hammeliana 

Quapoya sipapoana Maguire =Clusia sipapoana 

Renggeria comans Meisn. =Clusia comans 

Renggeria montana Klotzsch [nomen nudum] =Clusia hammeliana 

Rengifa acuminata Planch. & Triana =Clusia hammeliana 

Rengifa peruviana Poepp. =Clusia hammeliana 

Rengifa scandens (Aubl.) Planch. & Triana =Quapoya scandens 

Rheedia acuminata (Ruiz & Pav.) Planch. & Triana =Rheedia madruno 

Rheedia kappleri Eyma =Rheedia madruno 

Rheedia sagotiana Engl. =Rheedia benthamiana 

Symphonia coccinea (Aubl.) Oken =Moronobea coccinea 

Symphonia microphylla R.E. Schult. =Symphonia globulifera 

Symphonia utilissima R.E. Schult. =Symphonia globulifera 

Tovomita angustata Steyerm. =Tovomita weddelliana 

Tovomita cephalostigma Vesque =Tovomita umbellata 

Tovomita duidae Steyerm. =Tovomita gracilipes 

Tovomita guianensis Planch. & Triana, non Aubl. 1775 =Tovomita brevistaminea 

Tovomita jenmanii Engl. =Tovomita brevistaminea 

Tovomita myriandra Benth. =Clusia myriandra 

Tovomita nigrescens Planch. & Triana =Tovomita umbellata 

Tovomita richardiana Planch. & Triana [nom. illeg.] =Tovomita longifolia


Tovomita stigmatosa Planch. & Triana =Tovomita umbellata 

Tovomitopsis membranacea (Planch. & Triana) D’Arcy =Chrysochlamys membranacea 

Tovomitopsis nicaraguensis Oerst., Planch. & Triana =Chrysochlamys nicaraguensis 

Tovomitopsis standleyana (L.O. Williams) D’Arcy =Chrysochlamys weberbaueri 

Verticillaria acuminata Ruiz & Pav. =Rheedia madruno 

Vismia acuminata (Lam.) Pers. =Vismia guianensis 

Vismia amazonica Ewan =Vismia gracilis 

Vismia angusta Miq. =Vismia macrophylla 

Vismia angustifolia Rusby =Vismia laxiflora 

Vismia baccifera subsp. ferruginea (Kunth) Ewan =Vismia guianensis 

Vismia caparosa Kunth =Vismia guianensis 

Vismia cayennensis var. sessilifolia (Aubl.) M.E. Berg =Vismia sessilifolia 

Vismia cayennensis var. tenuinervia M.E. Berg =Vismia tenuinervia 

Vismia confertiflora auct. non Spruce ex Reichardt 1878: sensu Ewan =Vismia baccifera subsp. dealbata 

Vismia dealbata Kunth =Vismia baccifera subsp. dealbata 

Vismia falcata Rusby =Vismia laxiflora 

Vismia ferruginea Kunth =Vismia guianensis 

Vismia glaziovii Ruhland =Vismia gracilis 

Vismia guianensis var. acuminata (Lam.) M.E. Berg =Vismia guianensis 

Vismia guianensis var. guianensis =Vismia guianensis 

Vismia lindeniana Decne. [not in flora area] 

Vismia pentagyna (Spreng.) Ewan [not in flora area] 

Vismia reticulata (Poir.) Choisy =Vismia macrophylla 

Vismia rufescens (Lam.) Pers. =Vismia sessilifolia 

Vismia schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum] 

Vismia tomentosa auct. non Ruiz & Pav. 1798 =Vismia schultesii 

REFERENCES 

Berg, M.E. van der. 1979. Revisao das espécies brasileiras do gênero Rheedia L. Acta Amaz. 9(1): 43-74. 

Ewan, J. 1962. Synopsis of the South American species of Vismia. Contrib. U.S. Nat. Herb. 35(5): 293-377. 

Eyma, P.J. 1934. Guttiferae. In A. Pulle, Flora of Suriname 3(1): 65-118. 

Kearns, D.M., P.E. Berry, P.F. Stevens, N.L. Cuello A., J.J. Pipoly III, N.K.B. Robson, B.K. Holst, K. Kubitzki, & A.L. Weitzman. 

Clusiaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4. 1998. 248-329. 

Kubitzki, K. 1978. Caraipa and Mahurea (Bonnetiaceae). In B. Maguire & collaborators, Botany of the Guayana Highland. 


Maguire, B. 1958. Guttiferae. In B. Maguire, J. Wurdack & collaborators, Botany of the Guayana Highland Part III. Mem. New 

York Bot. Gard. 10(1): 49-61. 

Maguire, B. 1961. Guttiferae. In B. Maguire, J.J. Wurdack & collaborators, Botany of the Guayana Highland part IV. Mem. New 

York Bot. Gard. 10(4): 21-32. 

Maguire, B. 1964. Guttiferae subfam. Moronoboideae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana 

Highland Part V. Mem. New York Bot. Gard. 10(5): 123-135. 

Maguire, B. 1972. Clusiaceae - Quapoya and Renggeria. In B. Maguire & collaborators, Botany of the Guayana Highland Part 

IX. Mem. New York Bot. Gard. 23: 192-195. 

Pipoly, J.J. III & M.H.G. Gustafsson. 2002. Clusiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. 


York Bot. Gard. 76 (2): 212-222. 

Robson, N.K.B. 1987. Studies in the genus Hypericum L. (Guttiferae). 7. Section 29. Brathys (part 1). Bull. Br. Mus. Nat. Hist. 

(Bot.) 16: 1-106. 

Robson, N.K.B. 1990. Studies in the genus Hypericum L. (Guttiferae). 8. Sections 29. Brathys (part 2) and 30. Trigynobrathys. 

Bull. Br. Mus. Nat. Hist. (Bot.) 20: 1-151. 

Sandwith, N.Y. 1936. Contributions to the flora of tropical America: XXVI. The genus Tovomita in British Guiana. Kew Bull. 

1936: 210-221. 

Combretaceae 

Alwan Al-Mayah, A.-R. A. & C.A. Stace. 1985. Resurrection of two Eighteenth Century names in American Combretaceae. 

Nord. J. Bot. 5(5): 447-449. 

Exell, A.W. 1935. Combretaceae. In A. Pulle, Flora of Suriname 3(1): 164-177. 

Exell, A.W. 1953. The Combretum species of the New World. J. Linn. Soc. Lond. (Bot.) 55: 103-141. 

Exell, A.W. & C.A. Stace. 1963. A revision of the genera Buchenavia and Ramatuella. Bull. Brit. Mus. (Nat. Hist.) 3: 1-46. 

Exell, A.W. & C.A. Stace. 1966. Revision of the Combretaceae. Bol. Soc. Brot. (ser. 2) 40: 5-26. 

Kawasaki, M.L. 2002. Combretaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 224-227. 

Stace, C.A., A.-R. Alwan. 1998. Combretaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 

4: 329-352.

254 

Buchenavia amazonia Alwan & Stace 

Buchenavia congesta Ducke 

Buchenavia fanshawei Exell & Maguire 

Buchenavia grandis Ducke 

Buchenavia guianensis (Aubl.) Alwan & Stace 

Buchenavia macrophylla Eichler 

Buchenavia megalophylla van Heurck & Müll. Arg. 

Buchenavia nitidissima (Rich.) Alwan & Stace 

Buchenavia ochroprumna Eichler 

Buchenavia oxycarpa (Mart.) Eichler 

Buchenavia pallidovirens Cuatrec. 

Buchenavia parvifolia Ducke 

Buchenavia pulcherrima Exell & Stace 

Buchenavia reticulata Eichler 

Buchenavia suaveolens Eichler 

Buchenavia tetraphylla (Aubl.) R.A. Howard 

Buchenavia viridiflora Ducke 

Combretum assimile Eichler 

Combretum brevistylum Eichler 

Combretum cacoucia Exell ex Sandwith 

Combretum decandrum Jacq. 

Combretum frangulifolium Kunth 

Combretum fruticosum (Loefl.) Stuntz 

Combretum glabrum DC. 

Combretum laurifolium Mart. 

Combretum laxum Jacq. 

Combretum llewelynii J.F. Macbr. 

Combretum pyramidatum Desv. 

Combretum rohrii Exell 

Combretum rotundifolium Rich. 

Combretum spinosum Bonpl. 

Conocarpus erectus L. [expected in DA] 

Laguncularia racemosa (L.) C.F. Gaertn. 

Terminalia amazonia (J.F. Gmel.) Exell 

Terminalia catappa L. [cultivated, naturalized] 

Terminalia crispialata (Ducke) Alwan & Stace 

Terminalia dichotoma G. Mey. 

Terminalia guaiquinimae Maguire & Exell 

Terminalia guyanensis Eichler 

Terminalia lucida Hoffmanns. ex Mart. 

Terminalia oblonga (Ruiz & Pav.) Steud. 

Terminalia quintalata Maguire 

Terminalia ramatuella Alwan & Stace 

Terminalia steyermarkii Alwan & Stace 

Terminalia tanibouca Rich. 

Terminalia virens (Spruce ex Eichler) Alwan & Stace 

Terminalia yapacana Maguire 

Thiloa paraguariensis Eichler 


COMBRETACEAE 

A.-R. Alwan, C.A. Stace 


Buceras bucida Crantz [not in flora area] =Bucida buceras L. 

Buchenavia capitata (Vahl) Eichler =Buchenavia tetraphylla 

Buchenavia discolor Diels =Buchenavia ochroprumna 

Buchenavia huberi Ducke =Buchenavia grandis 

Buchenavia longibracteata Fróes =Buchenavia congesta 

Buchenavia ptariensis Steyerm. =Buchenavia tetraphylla 

Buchenavia pterocarpa Exell & Stace =Buchenavia suaveolens 

Buchenavia stellae Cuatrec. =Buchenavia macrophylla 

Buchenavia vaupesana Cuatrec. =Buchenavia tetraphylla 

Bucida buceras L. [not in flora area] 

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Bucida capitata Vahl =Buchenavia tetraphylla 

Cacoucia coccinea Aubl. =Combretum cacoucia 

Chuncoa amazonia J.F. Gmel. =Terminalia amazonia 

Combretum accedens Van Heurck & Müll. Arg. =Combretum laxum 

Combretum aubletii DC. =Combretum rotundifolium 

Combretum brunnescens Gleason =Combretum laxum 

Combretum elegans Kunth =Combretum rotundifolium 

Combretum ferrugineum Hoffmanns. ex Mart. [nomen nudum] =Combretum laxum 

Combretum fulgens Gleason =Combretum laxum 

Combretum fusiforme Gleason =Combretum glabrum 

Combretum guanaiense Rusby =Combretum assimile 

Combretum guianense Miq. =Combretum rotundifolium 

Combretum jacquinii Griseb. =Combretum laxum 

Combretum latipaniculatum Rusby =Combretum decandrum 

Combretum laxum Aubl. =Combretum rotundifolium 

Combretum loeflingii Eichler =Combretum fruticosum 

Combretum magnificum Mart. =Combretum rotundifolium 

Combretum oblongifolium Rusby =Combretum laxum 

Combretum obtusifolium Rich. =Combretum laxum 

Combretum phaeocarpum Mart. =Combretum pyramidatum 

Combretum puberum Rich. =Combretum laxum 

Combretum pulchellum Mart. =Combretum laxum 

Combretum punctatum Steud. =Combretum rotundifolium 

Combretum sambuense Pittier =Combretum assimile 

Combretum sprucei Eichler =Thiloa paraguariensis 

Combretum terminalioides Steud. =Combretum laxum 

Conocarpus racemosa L. =Laguncularia racemosa 

Gaura fruticosa Loefl. =Combretum fruticosum 

Gimbernatea oblonga Ruiz & Pav. =Terminalia oblonga 

Gimbernatea obovata Ruiz & Pav. =Terminalia amazonia 

Pamea guianensis Aubl. =Buchenavia guianensis 

Ramatuela argentea Kunth =Terminalia ramatuella 

Ramatuela crispialata Ducke =Terminalia crispialata 

Ramatuela crispialata var. obtusa Maguire =Terminalia crispialata 

Ramatuela latifolia Maguire =Terminalia virens 

Ramatuela maguirei Exell & Stace =Terminalia virens 

Ramatuela obtusa (Maguire) Exell & Stace =Terminalia crispialata 

Ramatuela virens Spruce ex Eichler =Terminalia virens 

Tanibouca guianensis Aubl. =Terminalia dichotoma 

Terminalia firma Mart. =Terminalia lucida 

Terminalia nitidissima Rich. =Buchenavia nitidissima 

Terminalia obovata (Ruiz & Pav.) Steud. =Terminalia amazonia 

Terminalia odontoptera Van Heurck & Müll. Arg. =Terminalia amazonia 

Terminalia opacifolia Maguire & Exell =Terminalia guaiquinimae 

Terminalia oxycarpa Mart. =Buchenavia oxycarpa 

Terminalia pamea DC. =Buchenavia guianensis 

Terminalia virgata Maguire & Exell =Terminalia guaiquinimae 

Thiloa inundata Ducke =Thiloa paraguariensis 

Acanthospermum australe (Loefl.) Kuntze 

Acanthospermum hispidum DC. 

Achnopogon steyermarkii Aristeg. 

Achnopogon virgatus Maguire, Steyerm. & Wurdack 

Achyrocline flaccida DC. 

Achyrocline satureioides (Lam.) DC. 

Achyrocline vargasiana DC. 

Acmella brachyglossa Cass. 

Acmella ciliata (Kunth) Cass. 

Acmella radicans (Jacq.) R.K. Jansen 

Acmella radicans var. debilis (Kunth) R.K. Jansen 

COMPOSITAE 

V.A. Funk, H. Robinson, J.F. Pruski 


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256 


Acmella repens (Walter) Rich. ex Pers. 

Acmella uliginosa (Sw.) Cass. 

Ageratina ibaguensis (Sch. Bip. ex Hieron.) R.M. King & H. Rob. 

Ageratina roraimensis (N.E. Br.) R.M. King & H. Rob. [Mt. Roraima] 

Ageratum ballotifolium (Maguire, Steyerm. & Wurdack) R.M. King & H. Rob. 

Ageratum conyzoides L. [introduced] 

Ageratum houstonianum Mill. [widely cultivated, possibly naturalized] 

Ageratum scorpioideum Baker 

Ambrosia crithmifolia DC. 

Ambrosia microcephala DC. 

Ambrosia peruviana Willd. [expected in the Guianas] 

Austroeupatorium inulifolium (Kunth) R.M. King & H. Rob. 

Ayapana amygdalina (Lam.) R.M. King & H. Rob. 

Ayapana trinitensis (Kuntze) R.M. King & H. Rob. 

Ayapana triplinervis (Vahl) R.M. King & H. Rob. 

Baccharis brachylaenoides DC. 

Baccharis brachylaenoides var. brachylaenoides 

Baccharis brachylaenoides var. ligustrina (DC.) Maguire & Wurdack 

Baccharis densa (N.E. Br.) V.M. Badillo 

Baccharis lundii DC. 

Baccharis nitida (Ruiz & Pav.) Pers. 

Baccharis rufescens Spreng. 

Baccharis schomburgkii Baker 

Baccharis trinervis (Lam.) Pers. 

Baccharis varians Gardner 

Baltimora geminata (Brandegee) Stuessy 

Barrosoa metensis (B.L. Rob.) R.M. King & H. Rob. 

Bidens alba (L.) DC. 

Bidens alba var. radiata (Sch. Bip.) R.E. Ballard 

Bidens cynapiifolia Kunth 

Bidens pilosa L. 

Bidens riparia Kunth 

Bidens riparia var. refracta (Brandegee) O.E. Schulz 

Blainvillea dichotoma (Murray) Cass. 

Brickellia diffusa (Vahl) A. Gray 

Calea abelioides S.F. Blake 

Calea berteriana DC. 

Calea caleoides (DC.) H. Rob. 

Calea camani Maguire & K.D. Phelps 

Calea divaricata Benth. 

Calea esposi Maguire & K.D. Phelps 

Calea kunhardtii Maguire 

Calea linearifolia Maguire & Wurdack 

Calea lucida Maguire & Wurdack 

Calea lucidivenia Gleason & S.F. Blake 

Calea lucidivenia var. cardonae (Maguire & Wurdack) Pruski 

Calea lucidivenia var. lucidivenia 

Calea lucidivenia var. orientalis (Maguire & Wurdack) Pruski 

Calea lucidivenia var. punctata Maguire & Wurdack 

Calea nana Maguire 

Calea neblinensis (Maguire & Wurdack) Pruski 

Calea nov. sp. 1 Pruski 





Calea oliveri B.L. Rob. & Greenm. 

Calea orbiculata Maguire & Aristeg. 

Calea phelpsiae Lasser & Maguire 

Calea politii Maguire 

Calea punctata Maguire & Wurdack 

Calea sipapoana Maguire 

Calea solidaginea Kunth 

Calea solidaginea subsp. deltophylla (R.S. Cowan) Pruski 

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Calea sublantanoides V.M. Badillo 

Calea tolimana Hieron. 

Calea tricephala Maguire 

Centratherum punctatum Cass. 

Chaptalia integerrima (Vell.) Burkart 

Chaptalia nutans (L.) Pol. 

Chimantaea acopanensis Steyerm. 

Chimantaea cinerea (Gleason & S.F. Blake) Maguire, Steyerm. & Wurdack 

Chimantaea cinerea subsp. cinerea 

Chimantaea cinerea subsp. similis (Maguire, Steyerm. & Wurdack) Pruski & 

O. Huber 

Chimantaea eriocephala Maguire, Steyerm. & Wurdack 

Chimantaea espeletoidea Maguire, Steyerm. & Wurdack 

Chimantaea huberi Steyerm. 

Chimantaea humilis Maguire, Steyerm. & Wurdack 

Chimantaea lanocaulis Maguire, Steyerm. & Wurdack 

Chimantaea lanocaulis subsp. lanocaulis 

Chimantaea lanocaulis subsp. murisipanensis Pruski & O. Huber 

Chimantaea mirabilis Maguire, Steyerm. & Wurdack 

Chimantaea rupicola Maguire, Steyerm. & Wurdack 

Chionolaena latifolia (Benth.) Baker 

Chromolaena bathyphlebia (B.L. Rob.) R.M. King & H. Rob. 

Chromolaena corymbosa (Aubl.) R.M. King & H. Rob. 

Chromolaena ivifolia (L.) R.M. King & H. Rob. 

Chromolaena laevigata (Lam.) R.M. King & H. Rob. 

Chromolaena maximilianii (Schrad. ex DC.) R.M. King & H. Rob. 

Chromolaena odorata (L.) R.M. King & H. Rob. 

Chromolaena pharcidodes (B.L. Rob.) R.M. King & H. Rob. 

Chromolaena squalida (DC.) R.M. King & H. Rob. 

Chromolaena ternicapitulata Pruski 

Chromolaena thurnii (B.L. Rob.) R.M. King & H. Rob. 

Chromolaena tyleri (B.L. Rob.) R.M. King & H. Rob. 

Clibadium armanii (Balbis) Sch. Bip. ex O.E. Schulz 

Clibadium surinamense L. 

Clibadium sylvestre (Aubl.) Baill. 

Condylidium iresinoides (Kunth) R.M. King & H. Rob. 

Conyza bonariensis (L.) Cronquist 

Conyza canadensis (L.) Cronquist [naturalized] 

Conyza laevigata (Rich.) Pruski 

Conyza primulaefolia (Lam.) Cuatrec. & Lourteig 

Cosmos caudatus Kunth [widely cultivated, probably escaping] 

Cyanthillium cinereum (L.) H. Rob. [naturalized] 

Cyrtocymura scorpioides (Lam.) H. Rob. 

Dasyphyllum vepreculatum (D. Don) Cabrera 

Delilia biflora (L.) Kuntze 

Duidaea marahuacensis Steyerm. 

Duidaea pinifolia S.F. Blake 

Duidaea rubriceps S.F. Blake 

Duidaea tatei S.F. Blake 

Eclipta prostrata (L.) L. 

Egletes florida Shinners 

Egletes prostrata (Sw.) Kuntze 

Egletes viscosa (L.) Less. 

Elaphandra moriana Pruski 

Elephantopus hirtiflorus DC. 

Elephantopus mollis Kunth 

Eleutheranthera ruderalis (Sw.) Sch. Bip. 

Emilia coccinea (Sims) G. Don [African: cultivated, escaping] 

Emilia fosbergii Nicolson 

Emilia sonchifolia (L.) DC. ex Wight 

Erechtites hieracifolius (L.) Raf. ex DC. 

Erechtites valerianifolius (Link ex Spreng.) DC. 

Erigeron karwinskianus DC. 

Fleischmannia microstemon (Cass.) R.M. King & H. Rob. 

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258 


Gamochaeta americana (Mill.) Wedd. 

Gamochaeta simplicicaulis (Willd. ex Spreng.) Cabrera 

Glossarion bilabiatum (Maguire) Pruski 

Glossarion rhodanthum Maguire & Wurdack 

Gnaphalium elegans Kunth 

Gnaphalium polycaulon Pers. [Mt. Roraima; Sierra de la Neblina, Brazil] 

Gochnatia oligocephala (Gardner) Cabrera [expected in GU] 

Gongylolepis benthamiana R.H. Schomb. 

Gongylolepis bracteata Maguire 

Gongylolepis cortesii (S. Díaz) Pruski & S. Díaz [expected in AM] 

Gongylolepis erioclada S.F. Blake 

Gongylolepis fruticosa Maguire, Steyerm. & Wurdack 

Gongylolepis glaberrima S.F. Blake 

Gongylolepis huachamacari Maguire 

Gongylolepis jauaensis (Aristeg., Maguire & Steyerm.) V.M. Badillo 

Gongylolepis martiana (Baker) Steyerm. & Cuatrec. 

Gongylolepis oblanceolata Pruski 

Gongylolepis paniculata Maguire & K.D. Phelps 

Gongylolepis paruana Maguire 

Gongylolepis pedunculata Maguire 

Gongylolepis yapacana Maguire 

Guayania bulbosa (Aristeg.) R.M. King & H. Rob. 

Guayania cerasifolia (Sch. Bip. ex Baker) R.M. King & H. Rob. 

Guayania crassicaulis (Steyerm.) R.M. King & H. Rob. 

Guayania penninervata (Wurdack) R.M. King & H. Rob. 

Guayania roupalifolia (B.L. Rob.) R.M. King & H. Rob. 

Guayania yaviana (Lasser & Maguire) R.M. King & H. Rob. 

Hebeclinium macrophyllum (L.) DC. 

Heterocondylus vitalbae (DC.) R.M. King & H. Rob. 

Huberopappus maigualidae Pruski 

Ichthyothere davidsei H. Rob. 

Ichthyothere granvillei H. Rob. 

Ichthyothere terminalis (Spreng.) S.F. Blake 

Imeria memorabilis (Maguire & Wurdack) R.M. King & H. Rob. 

Imeria memorabilis var. densesericea Pruski [SW of Neblina] 

Imeria memorabilis var. memorabilis 

Koanophyllon fuscum (N.E. Br.) R.M. King & H. Rob. 

Koanophyllon tatei (B.L. Rob.) R.M. King & H. Rob. 

Lepidaploa bolivarensis (V.M. Badillo) H. Rob. 

Lepidaploa chrysotricha (Alexander) H. Rob. 

Lepidaploa cotoneaster (Willd. ex Spreng.) H. Rob. 

Lepidaploa ehretiifolia (Benth.) H. Rob. 

Lepidaploa gracilis (Kunth) H. Rob. 

Lepidaploa imeriensis (V.M. Badillo) Pruski 

Lepidaploa pari (V.M. Badillo) H. Rob. [expected in the Guianas] 

Lepidaploa remotiflora (Rich.) H. Rob. 

Lepidaploa salzmannii (DC.) H. Rob. [Mt. Roraima] 

Lessingianthus morilloi (V.M. Badillo) H. Rob. 

Lycoseris triplinervia Less. 

Melanthera nivea (L.) Small 

Mikania amazonica Baker 

Mikania banisteriae DC. 

Mikania boomii Pruski 

Mikania chaetoloba Pruski 

Mikania chlorolepis Baker 

Mikania congesta DC. 

Mikania cordifolia (L. f.) Willd. 

Mikania duidensis B.L. Rob. 

Mikania gleasonii B.L. Rob. 

Mikania guaco Bonpl. 

Mikania hookeriana DC. 

Mikania lindleyana DC. 

Mikania lucida S.F. Blake [Mt. Roraima] 

Mikania marahuacensis Steyerm. & Maguire 

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Mikania mazaruniensis W.C. Holmes & McDaniel 

Mikania michelangeliana Steyerm. 

Mikania micrantha Kunth 

Mikania microptera DC. 

Mikania neblinensis Aristeg. 

Mikania nigropunctulata Hieron. 

Mikania pannosa Baker 

Mikania parviflora (Aubl.) H. Karst. 

Mikania psilostachya DC. 

Mikania rondonensis V.M. Badillo 

Mikania simpsonii W.C. Holmes & McDaniel 

Mikania solidinervia V.M. Badillo 

Mikania sprucei Baker 

Mikania trinitaria DC. 

Mikania vitifolia DC. 

Neblinaea promontoriorum Maguire & Wurdack 

Neurolaena lobata (L.) R. Br. 

Oritrophium marahuacense Steyerm. & Maguire 

Orthopappus angustifolius (Sw.) Gleason 

Oyedaea scaberrima (Benth.) S.F. Blake 

Oyedaea tepuiana (V.M. Badillo) Pruski [expected in the Guianas] 

Oyedaea wurdackii Pruski 

Pacourina edulis Aubl. 

Parthenium hysterophorus L. 

Pectis ciliaris L. 

Pectis elongata Kunth 

Pectis elongata var. elongata 

Pectis elongata var. floribunda (A. Rich.) Keil 

Pectis humifusa Sw. 

Pectis pygmaea Kunth 

Pectis venezuelensis Steyerm. [expected in the Guianas] 

Pentacalia freemanii (Britton & Greenm.) Cuatrec. 

Pentacalia neblinensis Pruski 

Pentacalia phelpsiae (Cuatrec.) Cuatrec. 

Pentacalia ptariana (Cuatrec.) Cuatrec. 

Pentacalia yapacana (Aristeg.) Cuatrec. 

Piptocarpha auyantepuiensis Aristeg. 

Piptocarpha jauaensis Aristeg. & Steyerm. 

Piptocarpha leprosa (Less.) Baker 

Piptocarpha opaca (Benth.) Baker 

Piptocarpha opaca subsp. opaca 

Piptocarpha poeppigiana (DC.) Baker 

Piptocarpha polycephala Baker 

Piptocarpha triflora (Aubl.) Benn. ex Baker 

Piptocoma areolata (Wurdack) Pruski 

Piptocoma roraimensis (Steyerm.) Pruski 

Piptocoma schomburgkii (Sch. Bip.) Pruski 

Piptocoma spruceana (Benth.) Pruski 

Pluchea carolinensis (Jacq.) G. Don [probably cultivated only] 

Pluchea odorata (L.) Cass. 

Pluchea sagittalis (Lam.) Cabrera [expected in the Guianas] 

Porophyllum ruderale (Jacq.) Cass. 

Praxelis asperulacea (Baker) R.M. King & H. Rob. 

Praxelis diffusa (Rich.) Pruski 

Pseudelephantopus spicatus (Juss.) Rohr 

Pseudogynoxys chenopodioides (Kunth) Cabrera [expected in the Guianas] 

Pterocaulon alopecuroides (Lam.) DC. 

Quelchia bracteata Maguire, Steyerm. & Wurdack 

Quelchia cardonae Steyerm. 

Quelchia conferta N.E. Br. 

Quelchia eriocaulis Maguire, Steyerm. & Wurdack 

Quelchia x grandifolia Maguire, Steyerm. & Wurdack 

Riencourtia latifolia Gardner 

Riencourtia oblongifolia Gardner 

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260 


Riencourtia pedunculosa (Rich.) Pruski 

Rolandra fruticosa (L.) Kuntze 

Salmea scandens (L.) DC. [expected in flora area] 

Sciadocephala pakaraimae (Maguire & Wurdack) R.M. King & H. Rob. 

[expected in BO] 

Siapaea liesneri Pruski 

Sigesbeckia orientalis L. [naturalized] 

Sonchus asper (L.) Hill 

Sphagneticola brachycarpa (Baker) Pruski 

Sphagneticola trilobata (L.) Pruski 

Spilanthes nervosa Chodat 

Spilanthes urens Jacq. 

Stenocephalum apiculatum (Mart.) Sch. Bip. 

Stenopadus campestris Maguire & Wurdack 

Stenopadus chimantensis Maguire, Steyerm. & Wurdack 

Stenopadus colveei (Steyerm.) Pruski 

Stenopadus connellii (N.E. Br.) S.F. Blake 

Stenopadus cucullatus Maguire & Wurdack 

Stenopadus huachamacari Maguire 

Stenopadus jauensis Aristeg. 

Stenopadus kunhardtii Maguire 

Stenopadus megacephalus Pruski 

Stenopadus obconicus Maguire & Wurdack 

Stenopadus sericeus Maguire & Aristeg. 

Stenopadus talaumifolius S.F. Blake 

Stifftia cayennensis H. Rob. & Kahn 

Stomatochaeta acuminata Pruski 

Stomatochaeta condensata (Baker) Maguire & Wurdack 

Stomatochaeta crassifolia (S.F. Blake) Maguire & Wurdack 

Stomatochaeta cylindrica Maguire & Wurdack 

Stomatochaeta cymbifolia (S.F. Blake) Maguire & Wurdack 

Stomatochaeta steyermarkii Aristeg. 

Struchium sparganophorum (L.) Kuntze 

Symphyotrichum laeve (L.) Á. Löve & D. Löve [cultivated, naturalized] 

Synedrella nodiflora (L.) Gaertn. 

Tagetes erecta L. [cultivated, naturalized] 

Tessaria integrifolia Ruiz & Pav. 

Trichogonia arguta (Kunth) Benth. & Hook. f. ex Klatt [Mt. Roraima] 

Trichospira verticillata (L.) S.F. Blake 

Tridax procumbens L. 

Tuberculocarpus ruber (Aristeg.) Pruski 

Tyleropappus dichotomus Greenm. 

Unxia camphorata L. f. 

Unxia suffruticosa (Baker) Stuessy 

Verbesina angusta Maguire, Steyerm. & Wurdack 

Verbesina columbiana B.L. Rob. [expected in the Guianas] 

Verbesina guianensis Baker 

Verbesina ligulata (Maguire & Wurdack) Pruski [expected in the Guianas] 

Verbesina pilosa Maguire & Wurdack 

Vernonanthura brasiliana (L.) H. Rob. 

Wedelia ambigens S.F. Blake 

Wedelia calycina Rich. 

Wedelia villosa Gardner 

Wulffia baccata (L.) Kuntze 

Wulffia rubens Alexander 

Xiphochaeta aquatica Poepp. 

Youngia japonica (L.) DC. [introduced] 


Acanthospermum brasilum Schrank =Acanthospermum australe 

Acanthospermum humile var. hispidum (DC.) Kuntze =Acanthospermum hispidum 

Achnopogon quelchioides Aristeg. =Achnopogon steyermarkii 

Achyrocline satureioides var. vargasianum (DC.) Baker =Achyrocline vargasiana 

Acmella brasiliensis Spreng. =Sphagneticola trilobata 



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Acmella oppositifolia var. oppositifolia =Acmella repens 

Acmella radicans var. radicans [not in flora area] 

Adenostemma pakaraimae Maguire & Wurdack =Sciadocephala pakaraimae 

Ageratum guianense Aubl. =Hebeclinium macrophyllum 

Alomia ballotifolia Maguire, Steyerm. & Wurdack =Ageratum ballotifolium 

Ambrosia cumanensis Kunth =Ambrosia peruviana 

Ampherephis aristata Kunth =Centratherum punctatum 

Ampherephis mutica Kunth =Centratherum punctatum 

Aspilia bolivarana V.M. Badillo =Wulffia baccata 

Aspilia rubra Aristeg. =Tuberculocarpus ruber 

Aster laevis L. =Symphyotrichum laeve 

Augusta benthamiana (R.H. Schomb.) Kuntze =Gongylolepis benthamiana 

Augusta condensata (Baker) Kuntze =Stomatochaeta condensata 

Augusta martiana (Baker) Kuntze =Gongylolepis martiana 

Baccharis brasiliana L. =Vernonanthura brasiliana 

Baccharis daphnoides subsp. schomburgkii (Baker) Malag. =Baccharis schomburgkii 

Baccharis erioptera Benth. =Pterocaulon alopecuroides 

Baccharis ligustrina DC. =Baccharis brachylaenoides var. ligustrina 

Baccharis maguireana Malag. =Baccharis brachylaenoides var. brachylaenoides 

Baccharis meridensis Steyerm. =Baccharis brachylaenoides 

Baccharis ptariensis Steyerm. =Baccharis brachylaenoides var. ligustrina 

Baccharis rhexioides Kunth =Baccharis trinervis 

Baccharis roraimae M.R. Schomb. =Baccharis brachylaenoides 

Baccharis rufescens var. varians (Gardner) Baker =Baccharis varians 

Baccharis schomburgkii subsp. occidentalis Steyerm. & Maguire =Baccharis schomburgkii 

Baccharis trinervis var. rhexioides (Kunth) Baker =Baccharis trinervis 

Baccharis trinitensis Kuntze =Ayapana trinitensis 

Baccharis venulosa DC. =Baccharis brachylaenoides 

Baccharis venulosa var. cuspidibracteata Steyerm. =Baccharis brachylaenoides var. brachylaenoides 

Baccharis vitis-idaea Oliv. ex Thurn, non Kuntze =Baccharis brachylaenoides var. ligustrina 

Baccharis wurdackiana Malag. =Baccharis brachylaenoides var. ligustrina 

Baccharoides brachylepis (Sch. Bip. ex Baker) Kuntze =Centratherum punctatum 

Baccharoides muticum (Kunth) Kuntze =Centratherum punctatum 

Baccharoides punctatum (Cass.) Kuntze =Centratherum punctatum 

Baillieria aspera Aubl. =Clibadium surinamense 

Baillieria barbasco Kunth =Clibadium sylvestre 

Baillieria sylvestris Aubl. =Clibadium sylvestre 

Baltimora ovata Rusby =Baltimora geminata 

Bidens bipinnata var. cynapiifolia (Kunth) M. Gómez =Bidens cynapiifolia 

Bidens caudata (Kunth) Sch. Bip. =Cosmos caudatus 

Bidens leucantha var. pilosa (L.) Griseb. =Bidens pilosa 

Bidens nivea L. =Melanthera nivea 

Bidens pilosa f. indivisa Sherff =Bidens pilosa 

Bidens verticillata L. =Trichospira verticillata 

Buphthalmum strigosum Spreng. =Sphagneticola trilobata 

Cacalia cinerea (L.) Kuntze =Cyanthillium cinereum 

Cacalia cordifolia L. f. =Mikania cordifolia 

Cacalia porophyllum L. =Porophyllum ruderale 

Cacalia ruderalis (Jacq.) Sw. =Porophyllum ruderale 

Cacalia sonchifolia L. =Emilia sonchifolia 

Calea aspera Jacq. =Melanthera nivea 

Calea cardonae Maguire & Wurdack =Calea lucidivenia var. cardonae 

Calea cardonae subsp. neblinensis Maguire & Wurdack =Calea neblinensis 

Calea cardonae var. orientalis Maguire & Wurdack =Calea lucidivenia var. orientalis 

Calea clematidifolia Steyerm. =Calea solidaginea subsp. deltophylla 

Calea collina Klatt =Calea berteriana 

Calea congesta Rusby =Calea solidaginea 

Calea deltophylla R.S. Cowan =Calea solidaginea subsp. deltophylla 

Calea divaricata var. diffusa Maguire & Wurdack =Calea divaricata 

Calea herbert-smithii Rusby =Calea solidaginea 

Calea holtonii Rusby =Calea berteriana 

Calea huberana V.M. Badillo =Calea tolimana 

Calea lobata (L.) Sw. =Neurolaena lobata 

Calea lucidivenia f. lanceolata Steyerm. =Calea lucidivenia

262 


Calea luetzelburgii Suess. =Calea oliveri 

Calea membranacea Maguire & Wurdack =Calea lucidivenia 

Calea oliveri var. taeniotricha Steyerm. =Calea oliveri 

Calea ovalis subsp. sabanensis Cuatrec. =Calea tolimana 

Calea perforata Klatt =Calea solidaginea 

Calea prunifolia Kunth [not in flora area] 

Calea scandens Griseb. =Calea solidaginea 

Calea suffruticosa Maguire & Wurdack =Calea divaricata 

Calea suriani Cass. =Neurolaena lobata 

Calea ternifolia Oliv. ex Thurn =Calea oliveri 

Cardonaea jauaensis Aristeg., Maguire & Steyerm. =Gongylolepis jauaensis 

Carphobolus latifolius Sch. Bip. =Piptocarpha opaca subsp. opaca 

Centratherum brachylepis Sch. Bip. ex Baker =Centratherum punctatum 

Centratherum muticum (Kunth) Less. =Centratherum punctatum 

Centratherum punctatum var. parviflorum Baker =Centratherum punctatum 

Centrospermum xanthioides Kunth =Acanthospermum australe 

Ceratocephalus acmelia var. depauperata Kuntze =Acmella uliginosa 

Ceratocephalus urens (Jacq.) Kuntze =Spilanthes urens 

Chaptalia integrifolia (Cass.) Baker =Chaptalia integerrima 

Chatiakella platyglossa Cass. =Wulffia baccata 

Chatiakella stenoglossa Cass. =Wulffia baccata 

Chimantaea cinerea f. glabra Steyerm. =Chimantaea cinerea 

Chimantaea similis Maguire, Steyerm. & Wurdack =Chimantaea cinerea var. similis 

Chionolaena breweri Steyerm. & Maguire =Chionolaena latifolia 

Chuquiraga vepreculata D. Don =Dasyphyllum vepreculatum 

Chylodia sarmentosa Rich. ex Cass. =Wulffia baccata 

Clibadium asperum (Aubl.) DC. =Clibadium surinamense 

Clibadium barbasco (Kunth) DC. =Clibadium sylvestre 

Clibadium caracasanum DC. =Clibadium surinamense 

Clibadium pedunculosum (Rich.) DC. =Riencourtia pedunculosa 

Clibadium rotundifolium DC. =Clibadium armanii 

Clibadium schomburgkii Sch. Bip. ex M.R. Schomb. =Clibadium sylvestre 

Clibadium surinamense var. asperum (Aubl.) Baker =Clibadium surinamense 

Clibadium surinamense var. macrophyllum Steyerm. =Clibadium surinamense 

Clibadium vargasii DC. =Clibadium sylvestre 

Coleosanthus diffusus (Vahl) Kuntze =Brickellia diffusa 

Complaya trilobata (L.) Strother =Sphagneticola trilobata 

Conoclinium cerasifolium Sch. Bip. ex Baker =Guayania cerasifolia 

Conyza alopecuroides Lam. =Pterocaulon alopecuroides 

Conyza calophylla Willd. ex Sch. Bip. =Gochnatia oligocephala 

Conyza cayennensis Spreng. [name of uncertain application] 

Conyza chilensis Spreng. =Conyza primulaefolia 

Conyza cinerea L. =Cyanthillium cinereum 

Conyza cotoneaster Willd. ex Spreng. =Lepidaploa cotoneaster 

Conyza lobata L. =Neurolaena lobata 

Conyza riparia Kunth =Tessaria integrifolia 

Conyza sagittalis Lam. =Pluchea sagittalis 

Conyza scorpioides Lam. =Cyrtocymura scorpioides 

Conyza trinervis Lam. =Baccharis trinervis 

Coreopsis baccata L. =Wulffia baccata 

Cotula viscosa L. =Egletes viscosa 

Dermatophyllum schomburgkii Sch. Bip. ex M.R. Schomb. =Stomatochaeta condensata 

Diazeuxis latifolia D. Don =Lycoseris triplinervia 

Echinocephalum discoideum Baker =Melanthera nivea 

Echinops fruticosus L. =Rolandra fruticosa 

Echinops nodiflorus Lam. =Rolandra fruticosa 

Eclipta alba (L.) Hassk. =Eclipta prostrata 

Eclipta erecta L. [nom. illeg. superfl.] =Eclipta prostrata 

Egletes commixta Shinners =Egletes florida 

Elephantopus angustifolius Sw. =Orthopappus angustifolius 

Elephantopus carolinianus Willd. [not in flora area] 

Elephantopus carolinianus var. mollis (Kunth) Beurl. =Elephantopus mollis 

Elephantopus decemsquamatus V.M. Badillo =Elephantopus hirtiflorus 

Elephantopus martii Graham =Elephantopus mollis


Elephantopus nudiflorus Willd. =Orthopappus angustifolius 

Elephantopus pilosus Philipson =Elephantopus mollis 

Elephantopus quadriflora Less. =Orthopappus angustifolius 

Elephantopus scaber var. tomentosus Sch. Bip ex Baker =Elephantopus mollis 

Elephantopus spicatus B. Juss. =Pseudelephantopus spicatus 

Elephantopus tomentosus L. =Elephantopus mollis 

Elephantopus virgatus Ham. [name of uncertain application] 

Emilia javanica (Burm.) C.B. Rob. [not in flora area] 

Emilia sagittata (Vahl) DC. [not in flora area] 

Erigeron bonariensis L. =Conyza bonariensis 

Erigeron canadensis L. =Conyza canadensis 

Erigeron chilensis (Spreng.) G. Don =Conyza primulaefolia 

Erigeron laevigatus Rich. =Conyza laevigata 

Erigeron mucronatus DC. =Erigeron karwinskianus 

Ethulia sparganophora L. =Struchium sparganophorum 

Eupatorium amarum Vahl =Mikania parviflora 

Eupatorium amygdalinum Lam. =Ayapana amygdalina 

Eupatorium angulicaule Lasser & Maguire =Guayania yaviana 

Eupatorium asperulaceum Baker =Praxelis asperulacea 

Eupatorium bathyphlebium B.L. Rob. =Chromolaena bathyphlebia 

Eupatorium bulbosum Aristeg. =Guayania bulbosa 

Eupatorium cerasifolium Sch. Bip. ex Baker =Guayania cerasifolia 

Eupatorium conyzoides Vahl =Chromolaena odorata 

Eupatorium corymbosum Aubl. =Chromolaena corymbosa 

Eupatorium crassicaule Steyerm. =Guayania crassicaulis 

Eupatorium denticulatum Vahl =Mikania cordifolia Mikania micrantha 

Eupatorium dichotomum Sch. Bip. =Chromolaena odorata 

Eupatorium diffusum Vahl =Brickellia diffusa 

Eupatorium duidense V.M. Badillo =Koanophyllon fuscum 

Eupatorium fuscum N.E. Br. =Koanophyllon fuscum 

Eupatorium ibaguense Sch. Bip. ex Hieron. =Ageratina ibaguensis 

Eupatorium ibaguense var. indiscretum Maguire, Steyerm. & Wurdack =Ageratina ibaguensis 

Eupatorium inulifolium Kunth =Austroeupatorium inulifolium 

Eupatorium iresinoides Kunth =Condylidium iresinoides 

Eupatorium ivifolium L. =Chromolaena ivifolia 

Eupatorium ivifolium var. gracillima Baker =Chromolaena ivifolia 

Eupatorium ixodes Benth. =Ayapana amygdalina 

Eupatorium karuaiense V.M. Badillo =Praxelis asperulacea 

Eupatorium kavanayense V.M. Badillo =Koanophyllon fuscum 

Eupatorium laevigatum Lam. =Chromolaena laevigata 

Eupatorium macrophyllum L. =Hebeclinium macrophyllum 

Eupatorium martiusii DC. =Chromolaena squalida 

Eupatorium memorabile Maguire & Wurdack =Imeria memorabilis 

Eupatorium metense B.L. Rob. =Barrosoa metensis 

Eupatorium microstemon Cass. =Fleischmannia microstemon 

Eupatorium microstemon var. albiflorum Kuntze =Fleischmannia microstemon 

Eupatorium microstemon var. lilacinum Kuntze =Fleischmannia microstemon 

Eupatorium morichalanum Aristeg. =Barrosoa metensis 

Eupatorium obscurum DC. =Chromolaena ivifolia 

Eupatorium odoratum L. =Chromolaena odorata 

Eupatorium parviflorum Aubl. =Mikania parviflora 

Eupatorium pauciflorum Kunth =Praxelis diffusa 

Eupatorium penninervatum Wurdack =Guayania penninervata 

Eupatorium pharcidodes B.L. Rob. =Chromolaena pharcidodes 

Eupatorium pharcidodes var. pentanthum S.F. Blake =Chromolaena pharcidodes 

Eupatorium pharcidodes var. sororopanensis Steyerm. =Chromolaena pharcidodes 

Eupatorium psiadiifolium DC. =Chromolaena laevigata 

Eupatorium ptaretepuiense V.M. Badillo =Ageratina ibaguensis 

Eupatorium roraimensis N.E. Br. =Ageratina roraimensis 

Eupatorium roraimensis f. albisetum Steyerm. =Ageratina roraimensis 

Eupatorium roupalifolium B.L. Rob. =Guayania roupalifolia 

Eupatorium roupalifolium f. latifolium Steyerm. =Guayania roupalifolia 

Eupatorium roupalifolium var. subintegrifolium Steyerm. =Guayania roupalifolia 

Eupatorium schomburgkii Benth. =Baccharis brachylaenoides

264 


Eupatorium sieberianum DC. =Condylidium iresinoides 

Eupatorium squalidum DC. =Chromolaena squalida 

Eupatorium squalidum var. subvelutinum (DC.) Baker =Chromolaena squalida 

Eupatorium squiresii Rusby ex B.L. Rob. =Ayapana trinitensis 

Eupatorium subvelutinum DC. =Chromolaena squalida 

Eupatorium surinamense Sch. Bip. ex Miq. =Ayapana trinitensis 

Eupatorium tatei B.L. Rob. =Koanophyllon tatei 

Eupatorium tepuianum Steyerm. =Guayania roupalifolia 

Eupatorium thurnii B.L. Rob. =Chromolaena thurnii 

Eupatorium triflorum Aubl. =Piptocarpha triflora 

Eupatorium trinitense (Kuntze) Rusby & B.L. Rob. =Ayapana trinitensis 

Eupatorium triplinerve Vahl =Ayapana triplinervis 

Eupatorium tyleri B.L. Rob. =Chromolaena tyleri 

Eupatorium urticifolium L. f. =Praxelis diffusa 

Eupatorium vitalbae DC. =Heterocondylus vitalbae 

Eupatorium wageneri Hieron. =Condylidium iresinoides 

Eupatorium yavianum Lasser & Maguire =Guayania yaviana 

Eurydochus bracteatus Maguire & Wurdack =Gongylolepis wurdackii 

Eurydochus cortesii S. Díaz = Gonglolepis cortesii (S.Díaz) Pruski & S. Díaz 

Gamochaeta purpurea (L.) Cabrera =Gamochaeta simplicicaulis 

Geissopappus caleoides (DC.) Benth. ex Baker =Calea caleoides 

Glossarion rhodanthum var. bracteatum Maguire =Glossarion rhodanthum 

Gnaphalium americanum Mill. =Gamochaeta americana 

Gnaphalium purpureum var. americanum (Mill.) Klatt =Gamochaeta americana 

Gnaphalium purpureum var. simplicicaule (Willd. ex Spreng.) Klatt =Gamochaeta simplicicaulis 

Gnaphalium satureioides Lam. =Achyrocline satureioides 

Gnaphalium satureioides var. vargasianum (DC.) Kuntze =Achyrocline vargasiana 

Gnaphalium schomburgkii Sch. Bip. [nomen nudum] =Gnaphalium polycaulon 

Gnaphalium simplicicaule Willd. ex Spreng. =Gamochaeta simplicicaulis 

Gnaphalium spicatum Lam. =Gamochaeta americana 

Gnaphalium suaveolens Vell. =Pluchea sagittalis 

Gochnatia calophylla (Sch. Bip.) V.M. Badillo =Gochnatia oligocephala 

Gongylolepis benthamiana var. pubescens Maguire & Wurdack =Gongylolepis benthamiana 

Gongylolepis cabreraei V.M. Badillo =Gongylolepis benthamiana 

Gongylolepis huachamacari subsp. neblinensis Maguire & Wurdack =Gongylolepis huachamacari 

Gongylolepis maroana V.M. Badillo =Gongylolepis martiana 

Gongylolepis paniculata var. umbellata Maguire & K.D. Phelps =Gongylolepis paniculata 

Gongylolepis terramarae Steyerm. =Gongylolepis paniculata 

Greenmania boladorensis Hieron. =Unxia camphorata 

Guaicaia bilabiata Maguire =Glossarion bilabiatum 

Guayania davidsei R.M. King & H. Rob. =Guayania crassicaulis 

Heterothalamus densus N.E. Br. =Baccharis densa 

Ichthyothere cunabi Mart. =Ichthyothere terminalis 

Ichthyothere suffruticosa var. pilosa Aristeg. =Ichthyothere terminalis 

Imeria serratifolia V.M. Badillo =Imeria memorabilis var. memorabilis 

Inula primulifolia Lam. =Conyza primulaefolia 

Kleinia ruderalis Jacq. =Porophyllum ruderale 

Koanophyllon kavanayense (V.M. Badillo) R.M. King & H. Rob. =Koanophyllon fuscum 

Kuhnia arguta Kunth =Trichogonia arguta 

Lagascea mollis Cav. [not in flora area] 

Latreillea glabrata Benth. =Ichthyothere terminalis 

Lepidaploa auyantepuiensis (Aristeg.) H. Rob. =Lepidaploa ehretiifolia 

Leria integrifolia Cass. =Chaptalia integerrima 

Leucopholis latifolia Benth. =Chionolaena latifolia 

Lipochaeta scaberrima Benth. =Oyedaea scaberrima 

Lucilia breweri (Steyerm. & Maguire) V.M. Badillo =Chionolaena latifolia 

Lycoseris latifolia (D. Don) Benth. =Lycoseris triplinervia 

Lycoseris oblongifolia Rusby =Lycoseris triplinervia 

Marsea bonariensis (L.) V.M. Badillo =Conyza bonariensis 

Melampodium australe Loefl. =Acanthospermum australe 

Melampodium bonairense Bold. =Baltimora geminata 

Melampodium camphoratum (L. f.) Baker =Unxia camphorata 

Melampodium geminata Brandegee =Baltimora geminata 

Melampodium ruderale Sw. =Eleutheranthera ruderalis


Melampodium suffruticosum Baker =Unxia suffruticosa 

Melanthera aspera (Jacq.) Steud ex Small =Melanthera nivea 

Mikania acrensis B.L. Rob. =Mikania lindleyana 

Mikania amara (Vahl) Willd. =Mikania parviflora 

Mikania amara var. guaco (Humb. & Bonpl.) Baker =Mikania guaco 

Mikania anzoatiguensis V.M. Badillo =Mikania vitifolia 

Mikania argyrostigma Miq. =Mikania guaco 

Mikania aschersonii sensu Pruski, non Hieron. =Mikania chlorolepis 

Mikania aspera Miq. =Mikania guaco 

Mikania atriplicifolia Sch. Bip. ex Miq. =Mikania congesta 

Mikania auyantepuensis S.F. Blake =Mikania sprucei 

Mikania auyantepuensis var. brevipedicellata Steyerm. =Mikania sprucei 

Mikania banisteriae var. gabrieli (Baker) G.M. Barroso =Mikania banisteriae 

Mikania bolivarensis V.M. Badillo =Mikania pannosa 

Mikania capayensis V.M. Badillo =Mikania trinitaria 

Mikania denticulata (Vahl) Willd. =Mikania micrantha 

Mikania divaricata Poepp. =Mikania parviflora 

Mikania florida Aristeg. =Mikania lucida 

Mikania fockeana Miq. =Mikania psilostachya 

Mikania gabrieli Baker =Mikania banisteriae 

Mikania gonoclada DC. =Mikania cordifolia 

Mikania gracilipes B.L. Rob. [not in flora area] 

Mikania gracilis Sch. Bip. ex Miq. =Mikania vitifolia 

Mikania hexagona B.L. Rob. =Mikania microptera 

Mikania hirsutissima DC. =Mikania banisteriae 

Mikania hookeriana var. crassicaulis Steyerm. =Mikania hookeriana 

Mikania hookeriana var. platyphylla (DC.) B.L. Rob. =Mikania hookeriana 

Mikania hostmannii Miq. =Mikania cordifolia 

Mikania javitensis V.M. Badillo =Mikania nigropunctulata 

Mikania karuaiensis V.M. Badillo =Mikania psilostachya 

Mikania kavanayensis V.M. Badillo =Mikania banisteriae 

Mikania lanata Sch. Bip. [not in flora area] 

Mikania manaosensis Hieron. =Mikania sprucei 

Mikania micrantha var. congesta (DC.) B.L. Rob. =Mikania congesta 

Mikania orinocensis Kunth =Mikania micrantha 

Mikania parkeriana DC. =Mikania congesta 

Mikania phelpsii Maguire & Steyerm. =Mikania lucida 

Mikania platyphylla DC. =Mikania hookeriana 

Mikania psilostachya var. racemulosa (Benth.) Baker =Mikania psilostachya 

Mikania psilostachya var. scabra (DC.) Baker =Mikania psilostachya 

Mikania ptaretepuiensis V.M. Badillo =Mikania banisteriae 

Mikania punctata Klatt =Mikania vitifolia 

Mikania racemulosa Benth. =Mikania psilostachya 

Mikania roraimensis B.L. Rob. =Mikania sprucei 

Mikania ruiziana Poepp. =Mikania banisteriae 

Mikania santamensis V.M. Badillo =Mikania trinitaria 

Mikania scabra DC. =Mikania psilostachya 

Mikania scandens var. congesta (DC.) Baker =Mikania congesta 

Mikania scandens var. microptera (DC.) Baker =Mikania microptera 

Mikania serratifolia Sieber ex DC. =Condylidium iresinoides 

Mikania sprucei var. roraimensis (B.L. Rob.) Aristeg. =Mikania sprucei 

Mikania stictophora Steyerm. =Mikania sprucei 

Mikania stipitata Sch. Bip. ex Miq. =Mikania parviflora 

Mikania surinamensis Miq. =Mikania cordifolia 

Mikania tepuiensis V.M. Badillo =Mikania nigropunctulata 

Mikania tillettii R.M. King & H. Rob. =Mikania gleasonii 

Mikania vitrea B.L. Rob. =Mikania hookeriana 

Monanthemum cruegeri Griseb. =Piptocarpha triflora 

Neurolaena suriani (Cass.) Cass. =Neurolaena lobata 

Ogiera ruderalis (Sw.) Griseb. =Eleutheranthera ruderalis 

Oliganthes areolata Wurdack =Piptocoma areolata 

Oliganthes roraimensis Steyerm. =Piptocoma roraimensis 

Oliganthes schomburgkii Sch. Bip. =Piptocoma schomburgkii 

Oliganthes spruceana Benth. =Piptocoma spruceana

266 


Ooclinium clavatum Benth. =Chromolaena ivifolia 

Ooclinium villosum (Cass.) DC. =Praxelis diffusa 

Oyedaea blakeana Steyerm. =Oyedaea scaberrima 

Oyedaea rusbyi S.F. Blake [not in flora area] 

Pectis linifolia var. linifolia [not in flora area] 

Pentacalia gransabanensis (Aristeg.) Cuatrec. =Pentacalia freemanii 

Pentacalia nigella (V.M. Badillo) Cuatrec. =Pentacalia freemanii 

Piptocarpha cruegeri (Griseb.) Baker =Piptocarpha triflora 

Piptocarpha foliosa Cuatrec. =Piptocarpha poeppigiana 

Piptocarpha kavanayensis V.M. Badillo =Piptocarpha polycephala 

Piptocarpha opaca var. latifolia (Sch. Bip.) Baker =Piptocarpha opaca subsp. opaca 

Piptocarpha paraensis Cabrera =Piptocarpha poeppigiana 

Piptocarpha sprucei Baker =Piptocarpha leprosa 

Piptocarpha tereticaulis (DC.) Baker =Piptocarpha poeppigiana 

Piptocarpha upatensis V.M. Badillo =Gochnatia oligocephala 

Pluchea purpurascens (Sw.) DC. =Pluchea odorata 

Pluchea suaveolens (Vell.) Kuntze =Pluchea sagittalis 

Pollalesta areolata (Wurdack) Aristeg. =Piptocoma areolata 

Pollalesta faustiana Stutts =Piptocoma schomburgkii 

Pollalesta milleri (Johnst.) Aristeg. [not in flora area] 

Pollalesta roraimensis (Steyerm.) Aristeg. =Piptocoma roraimensis 

Pollalesta schomburgkii (Sch. Bip.) Aristeg. =Piptocoma schomburgkii 

Pollalesta spruceana (Benth.) Aristeg. =Piptocoma spruceana 

Polymnia stenocarpha S.F. Blake =Unxia suffruticosa 

Polymnia suffruticosa (Baker) V.M. Badillo =Unxia suffruticosa 

Porophyllum ellipticum (L.) Cass. =Porophyllum ruderale 

Porophyllum latifolium Benth. =Porophyllum ruderale 

Porophyllum macrocephalum DC. =Porophyllum ruderale 

Porophyllum porophyllum (L.) Kuntze =Porophyllum ruderale 

Porophyllum ruderale subsp. macrocephalum (DC.) R.R. Johnson =Porophyllum ruderale 

Praxelis karuaiensis (V.M. Badillo) R.M. King & H. Rob. =Praxelis asperulacea 

Praxelis pauciflora (Kunth) R.M. King & H. Rob. =Praxelis diffusa 

Praxelis villosa Cass. =Praxelis diffusa 

Pseudelephantopus spiralis (Less.) Cronquist [not in flora area] 

Pseudobaccharis densa (N.E. Br.) V.M. Badillo =Baccharis densa 

Pseudobaccharis ligustrina (DC.) I.L. Teodoro =Baccharis brachylaenoides var. ligustrina 

Pseudobaccharis rhexioides (Kunth) V.M. Badillo =Baccharis trinervis 

Pseudobaccharis trinervis (Lam.) V.M. Badillo =Baccharis trinervis 

Psila brachylaenoides (DC.) Aristeg. =Baccharis brachylaenoides 

Psila brachylaenoides var. ligustrina (DC.) Aristeg. =Baccharis brachylaenoides var. ligustrina 

Psila densa (N.E. Br.) V.M. Badillo =Baccharis densa 

Psila ligustrina (DC.) Cabrera =Baccharis brachylaenoides var. ligustrina 

Psila trinervis (Lam.) Cabrera =Baccharis trinervis 

Pterocaulon virgatum var. alopecuroides (Lam.) Griseb. =Pterocaulon alopecuroides 

Riencourtia glomerata Cass. =Riencourtia pedunculosa 

Riencourtia ovata S.F. Blake =Riencourtia latifolia 

Riencourtia pittieri S.F. Blake =Riencourtia latifolia 

Rolandra argentea Rottb. =Rolandra fruticosa 

Rolandra septans Willd. ex Less. =Trichospira verticillata 

Rolandra terminalis Spreng. =Ichthyothere terminalis 

Schomburgkia caleoides DC. =Calea caleoides 

Senecio chenopodioides Kunth =Pseudogynoxys chenopodioides 

Senecio confusus Britten =Pseudogynoxys chenopodioides 

Senecio freemanii Britton & Greenm. =Pentacalia freemanii 

Senecio gransabanensis Aristeg. =Pentacalia freemanii 

Senecio hieracifolius L. =Erechtites hieracifolius 

Senecio nigellus V.M. Badillo =Pentacalia freemanii 

Senecio phelpsiae Cuatrec. =Pentacalia phelpsiae 

Senecio ptarianus Cuatrec. =Pentacalia ptariana 

Senecio valerianifolius Wolf =Erechtites valerianifolius 

Senecio yapacanus Aristeg. =Pentacalia yapacana 

Seruneum scaberrimum (Benth.) Kuntze =Wedelia calycina 

Silphium trilobatum L. =Sphagneticola trilobata 

Sonchus oleraceus L. [not in flora area]


Sonchus oleraceus var. asper L. =Sonchus asper 

Sparganophorus sparganophora (L.) C. Jeffrey =Struchium sparganophorum 

Sparganophorus vaillantii Crantz =Struchium sparganophorum 

Spilanthes acmella (L.) Murray =Acmella sp. 

Spilanthes acmella var. uliginosa (Sw.) Baker =Acmella uliginosa 

Spilanthes barinensis Aristeg. =Spilanthes nervosa 

Spilanthes ciliata Kunth =Acmella ciliata 

Spilanthes debilis Kunth =Acmella radicans var. debilis 

Spilanthes oppositifolia (Lam.) D’Arcy =Acmella repens 

Spilanthes paniculata Wall. [not in flora area] =Acmella sp. 

Spilanthes radicans Jacq. =Acmella radicans 

Spilanthes repens Spreng. ex DC. =Spilanthes urens 

Spilanthes tenella Kunth =Acmella radicans var. debilis 

Spilanthes uliginosa Sw. =Acmella uliginosa 

Spilanthes uliginosa var. discoidea Aristeg. =Acmella uliginosa 

Spilanthes urens var. megalophylla S.F. Blake =Spilanthes urens 

Spilanthes urens var. nervosa (Chodat) Hassl. =Spilanthes nervosa 

Spiracantha cornifolia Kunth [not in flora area] 

Stenopadus affinis Maguire, Steyerm. & Wurdack =Stenopadus talaumifolius 

Stenopadus cardonae Maguire & Lasser =Stenopadus talaumifolius 

Stenopadus cinereus Gleason & S.F. Blake =Chimantaea cinerea 

Stenopadus cinereus f. glabra Steyerm. =Chimantaea cinerea 

Stenopadus condensatus (Baker) S.F. Blake =Stomatochaeta condensata 

Stenopadus crassifolius S.F. Blake [not in flora area] 

Stenopadus cymbifolius S.F. Blake =Stomatochaeta cymbifolia 

Stenopadus eurylepis S.F. Blake [not in flora area] 

Stenopadus guaiquinimensis V.M. Badillo =Stomatochaeta condensata 

Stenopadus kunhardtii var. grandifolia Maguire =Stenopadus kunhardtii 

Stenopadus neblinensis Maguire & Wurdack =Stenopadus talaumifolius 

Stenopadus steyermarkii Aristeg. =Stenopadus talaumifolius 

Stenopadus stipitatus Maguire & Wurdack =Stenopadus kunhardtii 

Stenopadus talaumifolius var. magnifolius Steyerm. =Stenopadus talaumifolius 

Stenopadus variabilis S.F. Blake =Stomatochaeta condensata 

Stenopadus ventricosus Maguire & Aristeg. =Stenopadus sericeus 

Stifftia benthamiana (R.H. Schomb.) Baker =Gongylolepis benthamiana 

Stifftia condensata Baker =Stomatochaeta condensata 

Stifftia connellii N.E. Br. =Stenopadus connellii 

Stifftia martiana Baker =Gongylolepis martiana 

Stilpnopappus aquaticus (Poepp.) M.O. Dillon =Xiphochaeta aquatica 

Stilpnopappus viridis Benth. ex Baker =Xiphochaeta aquatica 

Stomatochaeta colveei Steyerm. =Stenopadus colveei 

Tephrothamnus calophyllus Sch. Bip. =Gochnatia oligocephala 

Tessaria dentata Ruiz & Pav. =Tessaria integrifolia 

Thelechitonia brachycarpa (Baker) H. Rob. & Cuatrec. =Sphagneticola brachycarpa 

Thelechitonia muricata Cuatrec. =Sphagneticola brachycarpa 

Thelechitonia trilobata (L.) H. Rob. & Cuatrec. =Sphagneticola trilobata 

Tilesia baccata var. baccata =Wulffia baccata 

Tilesia capitata G. Mey. =Wulffia baccata 

Tilesia rubens (Alexander) Pruski =Wulffia rubens 

Trichospira biaristata Less. =Trichospira verticillata 

Trichospira menthoides Kunth =Trichospira verticillata 

Trichospira prieurei DC. =Trichospira verticillata 

Trichospira pulegium Mart. ex DC. =Trichospira verticillata 

Trinchinettia caleoides (DC.) Endl. =Calea caleoides 

Trixis pedunculosa Rich. =Riencourtia pedunculosa 

Tussilago integerrima Vell. =Chaptalia integerrima 

Tussilago nutans L. =Chaptalia nutans 

Unxia kubitzkii H. Rob. =Unxia suffruticosa 

Verbesina alata L. [not in flora area] 

Verbesina alba L. =Eclipta prostrata 

Verbesina nodiflora L. =Synedrella nodiflora 

Verbesina prostrata L. =Eclipta prostrata 

Verbesina schomburgkii subsp. ligulata Maguire & Wurdack =Verbesina ligulata 

Verbesina schomburgkii subsp. schomburgkii =Verbesina guianensis

268 


Verbesina weberbaueri Sagást. =Verbesina columbiana 

Vernonia apiculata Mart. ex DC. =Stenocephalum apiculatum 

Vernonia auyantepuiensis Aristeg. =Lepidaploa ehretiifolia 

Vernonia bolivarensis V.M. Badillo =Lepidaploa bolivarensis 

Vernonia brasiliana (L.) Druce =Vernonanthura brasiliana 

Vernonia cardonae Aristeg. =Lepidaploa ehretiifolia 

Vernonia chrysotricha Alexander =Lepidaploa chrysotricha 

Vernonia cinerea (L.) Less. =Cyanthillium cinereum 

Vernonia cotoneaster (Willd. ex Spreng.) Less. [misident. of Lepidaploa gracilis H. Rob.] =Lepidaploa cotoneaster 

Vernonia decantha Sch. Bip. =Lepidaploa ehretiifolia 

Vernonia ehretiifolia Benth. =Lepidaploa ehretiifolia 

Vernonia glandulosa V.M. Badillo =Lepidaploa bolivarensis 

Vernonia gracilis Kunth =Lepidaploa gracilis 

Vernonia guianensis V.M. Badillo =Lepidaploa salzmannii 

Vernonia imeriensis V.M. Badillo =Lepidaploa imeriensis 

Vernonia leprosa Less. =Piptocarpha leprosa 

Vernonia miersiana Gardner =Lepidaploa salzmannii 

Vernonia morilloi V.M. Badillo =Lessingianthus morilloi 

Vernonia moritziana Sch. Bip. =Lepidaploa gracilis 

Vernonia odoratissima Kunth =Vernonanthura brasiliana 

Vernonia opaca Benth. =Piptocarpha opaca 

Vernonia pari V.M. Badillo =Lepidaploa pari 

Vernonia poeppigiana DC. =Piptocarpha poeppigiana 

Vernonia remotiflora Rich. =Lepidaploa remotiflora 

Vernonia scabra Pers. =Vernonanthura brasiliana 

Vernonia schomburgkiana Sch. Bip. =Lepidaploa ehretiifolia 

Vernonia scorpioides (Lam.) Pers. =Cyrtocymura scorpioides 

Vernonia sericea Rich. [not in flora area] =Lepidaploa sericea (Rich.) H. Rob. 

Vernonia sessiliflora Willd. =Lepidaploa remotiflora 

Vernonia tereticaulis DC. =Piptocarpha poeppigiana 

Vernonia tricephala auct. non Gardner =Lepidaploa pari 

Vernonia tricholepis DC. =Lepidaploa remotiflora 

Wedelia brachycarpa Baker =Sphagneticola brachycarpa 

Wedelia brasiliensis (Spreng.) S.F. Blake [nom. illeg.] =Sphagneticola trilobata 

Wedelia brasiliensis var. villosa (Baker) S.F. Blake =Sphagneticola brachycarpa 

Wedelia caracasana DC. =Wedelia calycina 

Wedelia discoidea Less. =Eleutheranthera ruderalis 

Wedelia frutescens Jacq. =Wedelia calycina 

Wedelia fruticosa Jacq. [not in flora area. Misident. of W. calycina Rich.] 

Wedelia jacquini O.E. Schulz =Wedelia calycina 

Wedelia paludosa DC. =Sphagneticola trilobata 

Wedelia paludosa var. villosa Baker ? =Sphagneticola brachycarpa 

Wedelia parviflora Rich. =Wedelia calycina 

Wedelia psammophila Poepp. =Eclipta prostrata 

Wedelia rubra (Aristeg.) B.L. Turner =Tuberculocarpus ruber 

Wedelia scaberrima Benth. =Wedelia calycina 

Wedelia trilobata (L.) Hitchc. =Sphagneticola trilobata 

Willoughbya amazonica (Baker) Kuntze =Mikania amazonica 

Willoughbya argyrostigma (Miq.) Kuntze =Mikania guaco 

Willoughbya aspera (Miq.) Kuntze =Mikania guaco 

Willoughbya banisteriae (DC.) Kuntze =Mikania banisteriae 

Willoughbya chlorolepis (Baker) Kuntze =Mikania chlorolepis 

Willoughbya cordifolia (L. f.) Kuntze =Mikania cordifolia 

Willoughbya divaricata (Poepp.) Kuntze =Mikania parviflora 

Willoughbya gabrieli (Baker) Kuntze =Mikania banisteriae 

Willoughbya gracilis (Sch. Bip. ex Miq.) Kuntze =Mikania vitifolia 

Willoughbya guaco (Humb. & Bonpl.) Kuntze =Mikania guaco 

Willoughbya hirsutissima (DC.) Kuntze =Mikania banisteriae 

Willoughbya lindleyana (DC.) Kuntze =Mikania lindleyana 

Willoughbya micrantha (Kunth) Rusby =Mikania micrantha 

Willoughbya pannosa (Baker) Kuntze =Mikania pannosa 

Willoughbya parviflora (Aubl.) Kuntze =Mikania parviflora 

Willoughbya parviflora var. guaco (Bonpl.) Kuntze =Mikania guaco 

Willoughbya platyphylla (DC.) Kuntze =Mikania hookeriana


Willoughbya psilostachya (DC.) Kuntze =Mikania psilostachya 

Willoughbya scabra (DC.) Kuntze =Mikania psilostachya 

Willoughbya scandens var. congesta (DC.) Kuntze =Mikania congesta 

Willoughbya scandens var. orinocensis (Kunth) Kuntze =Mikania micrantha 

Willoughbya sprucei (Baker) Kuntze =Mikania sprucei 

Willoughbya stipitata (Sch. Bip. ex Miq.) Kuntze =Mikania parviflora 

Willoughbya trinitaria (DC.) Kuntze =Mikania trinitaria 

Willoughbya vitifolia (DC.) Kuntze =Mikania vitifolia 

Wulffia capitata (G. Mey.) Sch. Bip. =Wulffia baccata 

Wulffia maculata DC. =Wulffia baccata 

Wulffia platyglossa (Cass.) DC. =Wulffia baccata 

Wulffia stenoglossa (Cass.) DC. =Wulffia baccata 

Zexmenia scaberrima (Benth.) Baker =Oyedaea scaberrima 

Zexmenia tepuiana V.M. Badillo =Oyedaea tepuiana 

REFERENCES 

Aristeguieta, L. 1964. Compositae. In T. Lasser, Flora de Venezuela 10: 1-947. 

Badillo, V.M. 1994 (1995). Enumeración de las Compuestas (Asteraceae) de Venezuela. Revista de la Facultad de Agronomía 

(Maracay) 45: 1–191. 

Bremer, K. 1994. Asteraceae: Cladistics & Classification. Portland, Oregon: Timber Press. 

Funk, V.A. 1991. The Compositae of the Guianas, I: Heliantheae (Heliantheae, Tageteae, Coreopsideae). Rhodora 93(875): 256- 

267. 

King, R.M., and H. Robinson. 1987. The genera of the Eupatorieae (Asteraceae). Monogrographs in Systematic Botany Missouri 

Botanical Garden 22: 1–581. 

Koster, J.T. 1938. Compositae. In A. Pulle, Flora of Suriname 4(2): 87-165. 

Pruski, J.F. 1991 (1993). Compositae of the Guayana Highland - V. The Mutisieae of the lost world of Brazil, Colombia, and 

Guyana. Bol. Mus. Paraense Emílio Goeldi (Bot.) 7: 335-392. 

Pruski, J.F. 1994. Compositae of the Guayana Shield, with special reference to the tribe Mutisieae. Page 28 (abstract) in Compositae: 

Systematics, Biology, Utilization: Papers and Abstracts, coordinator D. J. N. Hind. Kew: Royal Botanic Gardens. 

Pruski, J.F. 1997. Asteraceae. In J.A. Steyermark, P.E. Berry, & B.K. Holst, Flora of the Venezuelan Guayana 3: 177-393. 

Pruski, J.F. 2002. Asteraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


116. 

Robinson, H. 1992. The Asteraceae of the Guianas, III: Vernonieae and restoration of the genus Xiphochaeta. Rhodora 94(880): 

348-361. 

Cnestidium guianense (G. Schellenb.) G. Schellenb. 

Connarus cordatus L.A. Vidal, Carbonó & Forero 

Connarus coriaceus G. Schellenb. 

Connarus erianthus Benth. ex Baker 

Connarus erianthus var. stipitatus Forero 

Connarus fasciculatus (DC.) Planch. 

Connarus fasciculatus subsp. fasciculatus 

Connarus incomptus Planch. 

Connarus lambertii (DC.) Sagot 

Connarus megacarpus S.F. Blake 

Connarus patrisii (DC.) Planch. 

Connarus perrottetii (DC.) Planch. 

Connarus perrottetii var. perrottetii 

Connarus perrottetii var. rufus Forero 

Connarus punctatus Planch. 

Connarus rigidus Forero 

Connarus ruber (Poepp.) Planch. 

Connarus ruber var. ruber 

Connarus ruber var. sprucei (Baker) Forero 

Connarus venezuelanus Baill. 

Connarus venezuelanus var. orinocensis Forero 

Connarus venezuelanus var. venezuelanus 

Pseudoconnarus macrophyllus (Poepp.) Radlk. 

Pseudoconnarus subtriplinervis (Radlk.) G. Schellenb. 

Rourea amazonica (Baker) Radlk. 

CONNARACEAE 

E. Forero, E. Santana 


AM 


GU SU FG? 

GU SU FG? 

SU FG 

GU 

SU FG 

AM BO DA GU 

GU 

AM GU SU FG 


GU SU FG 

BO DA GU 

BO GU SU FG 

AM BO 

AM BO 

AM BO 

AM BO 

AM BO DA 

AM BO 

BO DA 

AM BO GU 

BO GU SU 

AM GU?

270 

Rourea cuspidata Benth. ex Baker 

Rourea cuspidata var. cuspidata 

Rourea cuspidata var. multijuga Forero 

Rourea cuspidata var. pedicellata Baker 

Rourea foreroi Aymard & P.E. Berry 

Rourea frutescens Aubl. 

Rourea glabra Kunth 

Rourea glabra var. glabra 

Rourea grosourdyana Baill. 

Rourea grosourdyana var. grosourdyana 

Rourea kappleri Lanj. 

Rourea krukovii Steyerm. 

Rourea ligulata Baker 

Rourea neglecta G. Schellenb. 

Rourea neglecta var. neglecta 

Rourea pubescens (DC.) Radlk. 

Rourea pubescens var. pubescens 

Rourea pubescens var. spadicea (Radlk.) Forero 

Rourea revoluta Planch. 

Rourea revoluta var. glabra Baker 

Rourea revoluta var. revoluta 

Rourea sprucei G. Schellenb. 

Rourea sprucei var. sprucei 

Rourea sprucei var. subcoriacea Forero 

Rourea surinamensis Miq. 



Bernardinia guianensis G. Schellenb. =Cnestidium guianense 

Connarus araucanus Cuatrec. =Connarus venezuelanus var. orinocensis 

Connarus confertiflorus Baker =Connarus patrisii 

Connarus fecundus Baker =Pseudoconnarus macrophyllus 

Connarus floribundus Planch. =Connarus perrottetii 

Connarus guianensis Lamb. ex DC. =Connarus lambertii 

Connarus incomptus var. subcordata Baker =Connarus incomptus 

Connarus macrophyllus Poepp. =Pseudoconnarus macrophyllus 

Connarus neglectus G. Schellenb. =Connarus coriaceus 

Connarus opacus G. Schellenb. =Connarus coriaceus 

Connarus pubescens DC. =Rourea pubescens 

Connarus schomburgkii auct. non Planch. 1850: sensu Pittier =Connarus ruber var. sprucei 

Connarus schomburgkii Planch. =Connarus lambertii 

Connarus sprucei auct. non Baker 1871: sensu Pittier =Connarus lambertii 

Connarus sprucei Baker =Connarus ruber var. sprucei 

Omphalobium fasciculatum DC. =Connarus fasciculatus 

Omphalobium lambertii DC. =Connarus lambertii 

Omphalobium patrisii DC. =Connarus patrisii 

Omphalobium perrottetii DC. =Connarus perrottetii 

Omphalobium rubrum Poepp. =Connarus ruber 

Pseudoconnarus fecundus (Baker) Radlk. =Pseudoconnarus macrophyllus 

Pseudoconnarus krukovii Steyerm. =Pseudoconnarus macrophyllus 

Robergia frutescens (Aubl.) Willd. =Rourea frutescens 

Rourea amazonica Huber =Rourea amazonica 

Rourea cardonae Lasser & Maguire =Rourea frutescens 

Rourea glabra Kunth var. amazonica Baker =Rourea amazonica 

Rourea glabra var. coriacea Baker =R. neglecta var. neglecta p.p., R. surinamensis p.p. 

Rourea glabra var. parviflora Baker =Rourea cuspidata var. cuspidata 

Rourea granatensis Cuatrec. =Rourea glabra var. glabra 

Rourea induta var. concinna Baker =Rourea frutescens 

Rourea macrophylla (Poepp.) Baker =Pseudoconnarus macrophyllus 

Rourea spadicea Radlk. =Rourea pubescens var. spadicea 

Rourea subtriplinervis Radlk. =Pseudoconnarus subtriplinervis 

Santalodes cuspidatum (Benth. ex Baker) Kuntze =Rourea cuspidata 

Santalodes frutescens (Aubl.) Kuntze =Rourea frutescens 

Santalodes macrophyllum (Poepp.) Kuntze =Pseudoconnarus macrophyllus 

Santalodes revolutum (Planch.) Kuntze =Rourea revoluta 

AM 

AM 

AM 

AM 

AM 


AM BO 

AM BO 

BO GU 

BO GU 

SU 

BO 

GU? FG 

AM FG 

AM FG 

DA GU SU FG 

DA FG 

GU 

GU 

GU 

GU 

SU FG 

AM 

AM 

BO 

AM BO 

BO DA GU SU FG


REFERENCES 

Forero, E. 1983. Connaraceae. Flora Neotropica Monogr. 36: 1-207. 

Forero, E. 2002. Connaraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


230. 

Forero, E. & A.R.A. Görts-van Rijn. 1976. Connaraceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of 

Suriname 2(2): 654-657. 

Forero, E. & E. Santana. 1998. Connaraceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 

365-376. 

Lanjouw, J. 1940. Connaraceae. In A. Pulle, Flora of Suriname 2(2): 332-340. 

CONVOLVULACEAE 

D.F. Austin 

Aniseia cernua Choisy 

Aniseia martinicensis (Jacq.) Choisy 

Bonamia apurensis D.F. Austin 

Bonamia holtii O’Donell 

Bonamia maripoides Hallier f. 

Calycobolus glaber (Kunth) House 

Convolvulus nodiflorus Desr. 

Dicranostyles ampla Ducke 

Dicranostyles ampla var. ampla 

Dicranostyles ampla var. castanea D.F. Austin 

Dicranostyles globostigma D.F. Austin 

Dicranostyles guianensis Mennega 

Dicranostyles holostyla Ducke 

Dicranostyles integra Ducke 

Dicranostyles longifolia Ducke 

Dicranostyles scandens Benth. 

Dicranostyles sericea Gleason 

Dicranostyles villosus Ducke 

Evolvulus alsinoides (L.) L. 

Evolvulus cardiophyllus Schltdl. 

Evolvulus convolvuloides (Willd.) Stearn 

Evolvulus elegans Moric. 

Evolvulus filipes Mart. 

Evolvulus frankenioides Moric. 

Evolvulus glomeratus Nees & Mart. 

Evolvulus nummularius (L.) L. 

Evolvulus ovatus Fernald 

Evolvulus paniculatus (Bonpl.) Spreng. 

Evolvulus pterocaulon Moric. 

Evolvulus sericeus Sw. 

Evolvulus sericeus var. holosericeus (Kunth) Ooststr. 

Evolvulus sericeus var. sericeus 

Evolvulus tenuis Mart. ex Choisy 

Evolvulus tenuis subsp. sericatus (House) Ooststr. 

Evolvulus villosissimus Ooststr. 

Ipomoea alba L. 

Ipomoea angulata Mart. ex Choisy 

Ipomoea anisomeres B.L. Rob. & Bartlett 

Ipomoea aquatica Forssk. 

Ipomoea argentea Meisn. 

Ipomoea asarifolia (Desr.) Roem. & Schult. 

Ipomoea batatas (L.) Lam. [cultivated and frequently escaping] 

Ipomoea batatoides Choisy 

Ipomoea bignonioides Lindl. 

Ipomoea cairica (L.) Sweet 

Ipomoea carnea Jacq. 

Ipomoea carnea subsp. fistulosa (Mart. ex Choisy) D.F. Austin 

Ipomoea discolor (Kunth) G. Don 

Ipomoea distans Choisy 

Ipomoea fimbriosepala Choisy 



AM 

AM BO 



AM BO 


GU SU? 

GU SU? 

AM 


AM GU? 

FG 

BO 

AM GU? 

AM 

SU FG 


BO GU SU FG 

GU SU? FG? 

AM BO 


BO 

BO GU SU FG 

BO GU SU FG 

BO 

AM 

AM BO 

BO GU SU 

BO 

BO GU SU 

AM 

AM 

BO 


FG? 

GU SU FG? 

GU SU FG 


BO GU? 


GU SU FG 

FG 

GU 

BO DA GU SU? 

BO DA GU SU? 

BO 

FG 

DA FG?

272 

Ipomoea gabrieli Choisy 

Ipomoea hederifolia L. 

Ipomoea horsfalliae Hook. 

Ipomoea imperati (Vahl) Griseb. 

Ipomoea incarnata (Vahl) Choisy 

Ipomoea indica (Burm.) Merr. 

Ipomoea killipiana O’Donell 

Ipomoea leprieurii D.F. Austin 

Ipomoea mauritiana Jacq. 

Ipomoea meyeri (Spreng.) G. Don 

Ipomoea minutiflora (M. Martens & Galeotti) House 

Ipomoea nil (L.) Roth 

Ipomoea pes-caprae (L.) R. Br. 

Ipomoea phyllomega (Vell.) House 

Ipomoea piurensis O’Donell 

Ipomoea quamoclit L. 

Ipomoea reticulata O’Donell 

Ipomoea rubens Choisy 

Ipomoea schomburgkii Choisy 

Ipomoea setifera Poir. 

Ipomoea squamosa Choisy 

Ipomoea subrevoluta Choisy 

Ipomoea tiliacea (Willd.) Choisy 

Ipomoea trifida (Kunth) G. Don 

Ipomoea turbinata Lag. 

Ipomoea violacea L. [expected in DA] 

Iseia luxurians (Moric.) O’Donell 

Jacquemontia agrestis (Choisy) Meisn. 

Jacquemontia ciliata Sandwith 

Jacquemontia densiflora (Meisn.) Hallier f. 

Jacquemontia gracillima (Choisy) Hallier f. 

Jacquemontia guyanensis (Aubl.) Meisn. 

Jacquemontia pentantha (Jacq.) G. Don 

Jacquemontia sphaerostigma (Cav.) Rusby 

Jacquemontia tamnifolia (L.) Griseb. 

Jacquemontia velutina Choisy 

Lysiostyles scandens Benth. 

Maripa densiflora Benth. 

Maripa elongata Ducke 

Maripa glabra Choisy 

Maripa longifolia Sagot ex Hallier f . 

Maripa paniculata Barb. Rodr. 

Maripa pauciflora D.F. Austin 

Maripa repens Rusby 

Maripa reticulata Ducke 

Maripa reticulata var. reticulata 

Maripa reticulata var. rugosa (Ducke) D.F. Austin 

Maripa scandens Aubl. 

Maripa violacea (Aubl.) Ooststr. ex Lanj. & Uittien 

Merremia aegyptia (L.) Urb. 

Merremia aturensis (Kunth) Hallier f. 

Merremia cissoides (Lam.) Hallier f. 

Merremia digitata (Spreng.) Hallier f. 

Merremia dissecta (Jacq.) Hallier f. 

Merremia macrocalyx (Ruiz & Pav.) O’Donell 

Merremia maypurensis Hallier f. 

Merremia quinquefolia (L.) Hallier f. 

Merremia ternifoliola Pittier 

Merremia umbellata (L.) Hallier f. 

Merremia wurdackii D.F. Austin & Staples 

Odonellia hirtiflora (M. Martens & Galeotti) K.R. Robertson 

Operculina hamiltonii (G. Don) D.F. Austin & Staples 

Operculina sericantha (Miq.) Ooststr. 


FG 

BO GU SU 

SU 

DA GU SU FG 

BO 

BO DA GU? 

AM BO 

BO 

BO 

DA GU SU 

FG 

FG 

FG 


DA GU SU FG 



BO 

BO 

GU SU FG 

DA GU SU FG 


DA GU SU FG 


DA GU SU FG 


AM BO 

BO 

GU? 

GU SU FG 

BO DA GU? 

AM BO GU 

BO 

BO 

DA GU SU FG 


AM BO FG 

BO GU 


GU? SU? 


AM 

AM 

BO GU SU FG 

GU SU FG 

FG 

AM 

AM 

GU SU? FG 

BO DA 

AM GU SU FG 

AM 

AM 

GU SU FG 



GU SU FG 


AM BO DA GU 

GU 

SU FG 

BO GU SU FG 

AM 

AM 


BO GU 

BO GU 



AM BO 

BO GU SU FG 

BO DA GU SU FG


Operculina turpethum (L.) Silva Manso [var. turpethum?] [introduced, native 

to Asia] 

Operculina turpethum var. ventricosa (Bertero) Staples & D.F. Austin 

[introduced, native to Asia] 

Tetralocularia pennellii O’Donell 


Aniseia cernua Moric. =Aniseia martinicensis 

Aniseia ensifolia Choisy =Aniseia martinicensis 

Aniseia ensifolia var. minor Choisy =Aniseia cernua 

Aniseia gracillima Choisy =Jacquemontia gracillima 

Aniseia hastata Meisn. =Ipomoea fimbriosepala 

Aniseia minor (Choisy) J.A. McDonald =Aniseia cernua 

Aniseia trichantha Pittier =Iseia luxurians 

Batatas crassicaulis Benth. =Ipomoea carnea subsp. fistulosa 

Batatas edulis (Thunb. ex Murray) Choisy =Ipomoea batatas 

Batatas villosa Choisy =Ipomoea argentea 

Calonyction aculeatum (L.) House =Ipomoea alba 

Calonyction bona-nox (L.) Bojer =Ipomoea alba 

Calonyction muricatum (L.) G. Don. =Ipomoea turbinata 

Calonyction tuba (Schltdl.) Colla =Ipomoea violacea 

Calycobolus pulchellus Willd. ex Roem. & Schult. =Calycobolus glaber 

Calycobolus spectabilis (Choisy) House =Bonamia maripoides 

Calystegia setifera (Poir.) Meisn. =Ipomoea setifera 

Cladostyles paniculatus Bonpl. =Evolvulus paniculatus 

Convolvulus aculeatus L. =Ipomoea alba 

Convolvulus acuminatus Vahl =Ipomoea indica 

Convolvulus agrestis Choisy =Jacquemontia agrestis 

Convolvulus alatus Ham. =Operculina hamiltonii 

Convolvulus alsinoides L. =Evolvulus alsinoides 

Convolvulus asarifolius Desr. =Ipomoea asarifolia 

Convolvulus aturensis Kunth =Merremia aturensis 

Convolvulus azureus Desr. =Jacquemontia pentantha 

Convolvulus batatas L. =Ipomoea batatas 

Convolvulus batatilla Kunth =Ipomoea carnea subsp. fistulosa 

Convolvulus caracassanus Roem. & Schult. =Merremia umbellata 

Convolvulus cissoides Lam. =Merremia cissoides 

Convolvulus discolor Kunth =Ipomoea discolor 

Convolvulus edulis Thunb. ex Murray =Ipomoea batatas 

Convolvulus glaber Aubl. =Merremia macrocalyx 

Convolvulus guyanensis Aubl. =Jacquemontia guyanensis 

Convolvulus imperati Vahl =Ipomoea imperati 

Convolvulus incarnatus Vahl =Ipomoea incarnata 

Convolvulus indicus Burm. =Ipomoea indica 

Convolvulus littoralis L. =Ipomoea imperati 

Convolvulus macrocalyx Ruiz & Pav. =Merremia macrocalyx 

Convolvulus martinicensis Jacq. =Aniseia martinicensis 

Convolvulus meyeri Spreng. =Ipomoea meyeri 

Convolvulus minutiflorus M. Martens & Galeotti =Ipomoea minutiflora 

Convolvulus mollis Kunth =Ipomoea indica 

Convolvulus muricatus L. =Ipomoea turbinata 

Convolvulus nil L. =Ipomoea nil 

Convolvulus nummularius L. =Evolvulus nummularius 

Convolvulus paniculatus L. =Ipomoea mauritiana 

Convolvulus pentanthus Jacq. =Jacquemontia pentantha 

Convolvulus pes-caprae L. =Ipomoea pes-caprae 

Convolvulus phillomega Vell. =Ipomoea phyllomega 

Convolvulus pinnatus Desr. =Ipomoea quamoclit 

Convolvulus quinquefolius (L.) L. =Merremia quinquefolia 

Convolvulus riparius Kunth =Merremia cissoides 

Convolvulus ruber Vahl =Ipomoea setifera 

Convolvulus sphaerostigma Cav. =Jacquemontia sphaerostigma 

Convolvulus stoloniferus Cirillo =Ipomoea imperati 

Convolvulus tamnifolius (L.) G. Mey. =Jacquemontia tamnifolia 

SU FG 

SU 

FG

274 


Convolvulus tiliaceus Willd. =Ipomoea tiliacea 

Convolvulus trifidus Kunth =Ipomoea trifida 

Convolvulus umbellatus L. =Merremia umbellata 

Convulvulus dissectus Jacq. =Merremia dissecta 

Dicranostyles imatacensis Steyerm. =Dicranostyles guianensis 

Dufourea glabra Kunth =Calycobolus glaber 

Evolvulus alsinoides var. debilis (Kunth) Ooststr. =Evolvulus alsinoides 

Evolvulus angustissimus Kunth =Evolvulus sericeus var. sericeus 

Evolvulus arenicola J.R. Johnst. =Evolvulus sericeus var. holosericeus 

Evolvulus commelinifolius Fernald =Evolvulus cardiophyllus 

Evolvulus debilis Kunth =Evolvulus alsinoides 

Evolvulus glomeratus f. strictus (Benth.) Ooststr. =Evolvulus glomeratus 

Evolvulus holosericeus Kunth =Evolvulus sericeus var. holosericeus 

Evolvulus linifolius auct. non L. 1762. =Evolvulus filipes 

Evolvulus sericatus House =Evolvulus tenuis subsp. sericatus 

Evolvulus strictus Benth. =Evolvulus glomeratus 

Ipomoea acuminata (Vahl) Roem. & Schult. =Ipomoea indica 

Ipomoea altissima Mart. ex Choisy =Operculina hamiltonii 

Ipomoea alulata Miq. =Operculina hamiltonii 

Ipomoea aturensis (Kunth) G. Don =Merremia aturensis 

Ipomoea batatilla (Kunth) G. Don =Ipomoea carnea subsp. fistulosa 

Ipomoea bona-nox L. =Ipomoea alba 

Ipomoea cardiosepala Meisn. =Ipomoea phyllomega 

Ipomoea cathartica Poir. =Ipomoea indica 

Ipomoea cissoides f. viscidula Meisn. =Merremia cissoides 

Ipomoea coccinea auct. non L. 1753. =Ipomoea hederifolia 

Ipomoea comosa House =Ipomoea argentea 

Ipomoea crassicaulis (Benth.) B.L. Rob. =Ipomoea carnea subsp. fistulosa 

Ipomoea demerariana Choisy =Ipomoea phyllomega 

Ipomoea digitata L. [not in flora area] 

Ipomoea digitata auct. non L. 1753. =Ipomoea mauritiana 

Ipomoea fistulosa Mart. ex Choisy =Ipomoea carnea subsp. fistulosa 

Ipomoea fragrans (Bojer) Bojer ex Choisy =Ipomoea rubens 

Ipomoea glabra (Aubl.) Choisy =Merremia macrocalyx 

Ipomoea guyanensis (Aubl.) Choisy =Jacquemontia guyanensis 

Ipomoea hamiltonii G. Don =Operculina hamiltonii 

Ipomoea hirtiflora M. Martens & Galeotti =Odonellia hirtiflora 

Ipomoea hostmanni Meisn. =Merremia macrocalyx 

Ipomoea indica var. acuminata (Vahl) Fosberg =Ipomoea indica 

Ipomoea juncea Choisy =Merremia aturensis 

Ipomoea littoralis (L.) Boiss. =Ipomoea imperati 

Ipomoea luxurians Moric. =Iseia luxurians 

Ipomoea macrantha Roem. & Schult. =Ipomoea violacea 

Ipomoea macrocalyx (Ruiz & Pav.) Choisy =Merremia macrocalyx 

Ipomoea martinicensis (Jacq.) G. Mey. =Aniseia martinicensis 

Ipomoea mollicoma Miq. =Merremia umbellata 

Ipomoea mollis (Kunth) G. Don =Ipomoea indica 

Ipomoea muricata (L.) Jacq. =Ipomoea turbinata 

Ipomoea paniculata (L.) R. Br. =Ipomoea mauritiana 

Ipomoea parkeri Choisy =Ipomoea rubens 

Ipomoea parkeri var. subsericea Meisn. =Ipomoea rubens 

Ipomoea perryana Duchass. & Walp. =Odonellia hirtiflora 

Ipomoea polyanthes Roem. & Schult. =Merremia umbellata 

Ipomoea pterodes Choisy =Operculina hamiltonii 

Ipomoea quinquefolia L. =Merremia quinquefolia 

Ipomoea reptans (L.) Poir. =Ipomoea aquatica 

Ipomoea riedelii Meisn. =Ipomoea batatoides 

Ipomoea riparia G. Don =Ipomoea rubens 

Ipomoea rubra (Vahl) Millsp. =Ipomoea setifera 

Ipomoea sericantha Miq. =Operculina sericantha 

Ipomoea solanifolia L. [misapplied name, not in flora area] 

Ipomoea squamisepala O’Donell =Ipomoea angulata 

Ipomoea stolonifera (Cirillo) J.F. Gmel. =Ipomoea imperati


Ipomoea tamnifolia L. =Jacquemontia tamnifolia 

Ipomoea trichocarpa auct. non Elliott 1817. =Ipomoea trifida 

Ipomoea triloba auct. non L. 1753. =Ipomoea trifida 

Ipomoea tuba (Schltdl.) G. Don =Ipomoea violacea 

Ipomoea tuberosa L. [not in flora area] =Merremia tuberosa (L.) Rendle 

Ipomoea villosa (Choisy) Meisn. =Ipomoea argentea 

Ipomoea villosa var. argentea (Meisn.) Hallier f. =Ipomoea argentea 

Jacquemontia azurea (Desr.) Choisy =Jacquemontia pentantha 

Jacquemontia densiflora Rusby =Jacquemontia densiflora 

Jacquemontia hirtiflora (M. Martens & Galeotti) O’Donell =Odonellia hirtiflora 

Jacquemontia lactescens Seem. =Odonellia hirtiflora 

Jacquemontia luxurians (Moric.) Hallier f. =Iseia luxurians 

Jacquemontia nodiflora (Desr.) G. Don =Convolvulus nodiflorus 

Jacquemontia rusbyana Standl. =Jacquemontia densiflora 

Jacquemontia violacea (Vahl) Choisy =Jacquemontia pentantha 

Jacquemontia violacea var. densiflora Meisn. =Jacquemontia densiflora 

Kuhlmaniella holostyla (Ducke) Barroso =Dicranostyles holostyla 

Lysiostyles cayennensis (Meisn.) Roberty =Maripa scandens 

Lysiostyles epiphytica Pittier =Dicranostyles scandens 

Lysiostyles pubescens Gleason =Lysiostyles scandens 

Lysiostyles scandens var. latifolia Benth. =Lysiostyles scandens 

Maripa cayennensis Meisn. =Maripa scandens 

Maripa passifloroides Benth. ex Meisn. =Maripa violacea 

Maripa rugosa Ducke =Maripa reticulata var. rugosa 

Maripa spectabilis Choisy =Bonamia maripoides 

Maripa volubilis Pittier =Odonellia hirtiflora 

Merremia glabra (Aubl.) Hallier f. =Merremia macrocalyx 

Merremia parviflora Pittier =Merremia quinquefolia 

Mouroucoa violacea Aubl. =Maripa violacea 

Murucoa cayennensis (Meisn.) Kuntze =Maripa scandens 

Murucoa densiflora (Benth.) Kuntze =Maripa densiflora 

Murucoa scandens (Aubl.) Kuntze =Maripa scandens 

Operculina alata (Ham.) Urb. =Operculina hamiltonii 

Operculina altissima (Mart. ex Choisy) Meisn. =Operculina hamiltonii 

Operculina passifloroides (Benth. ex Meisn.) Ducke =Maripa violacea 

Operculina pterodes (Choisy) Meisn. =Operculina hamiltonii 

Operculina surinamensis Meisn. =Operculina sericantha 

Operculina triquetra (Vahl) Hallier f. =Operculina turpethum 

Operculina ventricosa Peter =Operculina turpethum var. ventricosa 

Operculina violacea (Aubl.) Barb. Rodr. =Maripa violacea 

Pharbitis acuminatus (Vahl) Choisy =Ipomoea indica 

Pharbitis cathartica (Poir.) Choisy =Ipomoea indica 

Pharbitis mollis (Kunth) Choisy =Ipomoea indica 

Prevostea amazonicus Choisy =Calycobolus glaber 

Prevostea spectabilis (Choisy) Meisn. =Bonamia maripoides 

Quamoclit coccinea var. hederifolia (L.) House =Ipomoea hederifolia 

Quamoclit hederifolia (L.) G. Don =Ipomoea hederifolia 

Quamoclit pinnata (Desr.) Bojer =Ipomoea quamoclit 

Quamoclit quamoclit (L.) Britton =Ipomoea quamoclit 

Quamoclit vulgaris Choisy =Ipomoea quamoclit 

Thyella tamnifolia (L.) Raf. =Jacquemontia tamnifolia 

REFERENCES 

Austin, D.F. 1982. Convolvulaceae. In T. Lasser, Flora de Venezuela. 

Austin, D.F. 1982. Convolvulaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 377-424. 

Austin, D.F. 2002. Convolvulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


231-236. 

Austin, D.F. & P.B. Cavalcante. 1982. Convolvuláceas da Amazônia. Publ. Avulsas Mus. Paraense E. Goeldi 36: 1-134. 

Austin, D.F. & G.W. Staples III. 1981. Convolvulaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part XI. 

Mem. N.Y. Bot. Gard. 32: 1-391. 

Ooststroom, S.J. van. 1932. Convolvulaceae. In A. Pulle, Flora of Suriname 4(1): 66-102. 

Ooststroom, S.J. van. 1937. Convolvulaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 468-471.

276 


CRASSULACEAE 

D.M. Kearns 

Kalanchoe crenata (Andrews) Haw. [cultivated or escaped from cultivation] 

Kalanchoe pinnata (Lam.) Pers. [cultivated, naturalized] 


Bryophyllum calycinum Salisb. =Kalanchoe pinnata 

Bryophyllum pinnatum (Lam.) Kurz [nom. Illeg.] =Kalanchoe pinnata 

Cotyledon pinnata Lam. =Kalanchoe pinnata 

Kalanchoe brasiliensis Cambess. =Kalanchoe crenata 

REFERENCES 

Kearns, D.M. 1998. Crassulaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 430-431. 

Stoffers, A.L. 1984. Crassulaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname V(1): 573-575. 

CUCURBITACEAE 

C. Jeffrey, D.M. Kearns, M. Nee 

Cayaponia angustiloba (Cogn.) Cogn. 

Cayaponia botryocarpa C. Jeffrey 

Cayaponia coriacea Cogn. 

Cayaponia cruegeri (Naudin) Cogn. 

Cayaponia granatensis Cogn. 

Cayaponia guianensis C. Jeffrey 

Cayaponia jenmanii C. Jeffrey 

Cayaponia macrocalyx Harms [Sierra de la Neblina] 

Cayaponia metensis Cuatrec. 

Cayaponia ophthalmica R.E. Schult. 

Cayaponia racemosa (Mill.) Cogn. 

Cayaponia rigida (Cogn.) Cogn. 

Cayaponia selysioides C. Jeffrey 

Cayaponia sp. A 

Cayaponia tessmannii Harms 

Cayaponia triangularis (Cogn.) Cogn. 

Ceratosanthes palmata (L.) Urb. 

Citrullus lanatus (Thunb.) Matsum. & Nakai [cultivated, occasionally 

escaping] 

Coccinia grandis (L.) Voigt [cultivated, occasionally escaping] 

Cucumis anguria L. [cultivated, naturalized] 

Cucumis anguria var. anguria 

Cucumis melo L. [cultivated, occasionally escaping] 

Cucurbita moschata Duchesne [cultivated, occasionally escaping] 

Doyerea emetocathartica Grosourdy 

Fevillea cordifolia L. 

Fevillea trilobata L. 

Gurania acuminata Cogn. 

Gurania bignoniacea (Poepp. & Endl.) C. Jeffrey 

Gurania brevipedunculata Cogn. 

Gurania huberi Cogn. 

Gurania lobata (L.) Pruski 

Gurania nigrescens C. Jeffrey 

Gurania oxyphylla C. Jeffrey 

Gurania reticulata Cogn. 

Gurania robusta Suess. 

Gurania simplicifolia (Steyerm.) C. Jeffrey 

Gurania spruceana Cogn. 

Gurania subumbellata (Miq.) Cogn. 

Gurania trialata Cogn. 

Helmontia cardiophylla Harms 

Helmontia leptantha (Schltdl.) Cogn. 

Kedrostis foetidissima (Jacq.) Cogn. [naturalized?] 

Lagenaria siceraria (Molina) Standl. [cultivated, occasionally escaping] 

Luffa cylindrica (L.) M. Roem. [cultivated, occasionally escaping] 

Luffa sepium (G. Mey.) C. Jeffrey 

GU 


BO GU SU 

BO 

AM GU SU FG 


AM BO 

AM BO DA GU 

BO GU SU FG 

BO 

BO DA 

DA GU SU FG 


GU SU FG 


AM? BO? 

FG? 

GU SU FG 

BO GU SU FG 

AM GU SU FG 

BO 

GU FG 

DA GU? SU FG 

DA 

GU? SU FG? 

GU SU FG 

DA FG 

FG? 



GU FG 



BO GU 

GU SU FG 

FG 

GU? SU FG 

BO GU 

AM 


AM 

FG 


SU 



BO GU SU FG

Melothria cucumis Vell. 

Melothria pendula L. 

Melothria trilobata Cogn. 

Momordica charantia L. [cultivated, naturalized] 

Posadaea sphaerocarpa Cogn. 

Psiguria racemosa C. Jeffrey 

Psiguria triphylla (Miq.) C. Jeffrey 

Psiguria umbrosa (Kunth) C. Jeffrey 

Rytidostylis amazonica (C. Mart. ex Cogn.) Spruce ex Kuntze 

Selysia prunifera (Poepp. & Endl.) Cogn. 

Siolmatra pentaphylla Harms 

Wilbrandia hibiscoides Silva Manso 



Anguria bignoniacea Poepp. & Endl. =Gurania bignoniacea 

Anguria cissoides Benth. =Gurania bignoniacea 

Anguria elliptica Britton =Psiguria triphylla 

Anguria guianensis Klotzsch ex Schltdl. =Gurania bignoniacea 

Anguria kegeliana Schltdl. =Gurania lobata 

Anguria leptantha Schltdl. =Helmontia leptantha 

Anguria leptantha var. brevipes Schltdl. =Helmontia leptantha 

Anguria lobata L. =Gurania lobata 

Anguria longispica Sagot [name of uncertain application fide C. Jeffrey] 

Anguria multiflora Miq. =Gurania lobata 

Anguria simplicifolia Steyerm. =Gurania simplicifolia 

Anguria spinulosa Poepp. & Endl. =Gurania lobata 

Anguria subumbellata Miq. =Gurania subumbellata 

Anguria treslingiana Pulle =Psiguria triphylla 

Anguria trilobata sensu Pulle non (L.) Jacq. =Gurania lobata 

Anguria triphylla Miq. =Psiguria triphylla 

Anguria umbrosa Kunth =Psiguria umbrosa 

Anguria umbrosa auct. non Kunth =Psiguria triphylla 

Anguria vogeliana Suess. =Psiguria triphylla 

Anguriopsis margaritensis J.R. Johnst. =Doyerea emetocathartica 

Apodanthera spp. [no records verified for Guianas] 

Arkezostis granatensis (Cogn.) Kuntze =Cayaponia granatensis 

Bryonia guadalupensis Spreng. =Melothria pendula 

Bryonia latebrosa Aiton =Cayaponia racemosa 

Bryonia racemosa Mill. =Cayaponia racemosa 

Cayaponia andreana Cogn. =Cayaponia cruegeri 

Cayaponia bonplandii (Cogn.) Cogn. =Cayaponia coriacea 

Cayaponia calycina Cogn. =Cayaponia cruegeri 

Cayaponia latebrosa (Aiton) Cogn. =Cayaponia racemosa 

Cayaponia sclerocarpa Sagot [name of uncertain application fide C. Jeffrey] 

Cayaponia simplicifolia (Naudin) Cogn. [not in flora area] 

Ceratosanthes corniculata (Lam.) Cogn. =Ceratosanthes palmata 

Ceratosanthes latiloba var. acutiloba Cogn. =Ceratosanthes palmata 

Ceratosanthes tuberosa J.F. Gmel. =Ceratosanthes palmata 

Citrullus vulgaris Schrad. ex Eckl. & Zeyh. =Citrullus lanatus 

Corallocarpus emetocatharticus (Grosourdy) Cogn. =Doyerea emetocathartica 

Cucumis sepium G. Mey. =Luffa sepium 

Cucurbita lagenaria L. =Lagenaria siceraria 

Cucurbita siceraria Molina =Lagenaria siceraria 

Doyerea angosturensis Grosourdy =Doyerea emetocathartica 

Elaterium amazonicum Mart. ex Cogn. =Rytidostylis amazonica 

Elaterium carthaginense Jacq. [not in flora area] =Rytidostylis carthaginensis (Jacq.) Kuntze 

Fevillea javilla Kunth =Fevillea cordifolia 

Fevillea karstenii Cogn. =Fevillea cordifolia 

Gurania breviflora Cogn. =Gurania bignoniacea 

Gurania cissoides (Benth.) Cogn. =Gurania bignoniacea 

Gurania diffusa Cogn. =Gurania huberi 

Gurania diversifolia Cogn. =Gurania acuminata 

Gurania durandii Cogn. =Gurania lobata 

Gurania guianensis (Klotzsch ex Schltdl.) Cogn. =Gurania bignoniacea 

Gurania kegeliana (Schltdl.) Cogn. =Gurania lobata 

GU? 





GU? 


BO 

AM BO FG 

GU? SU FG 

GU 

GU?

278 


Gurania klotzschiana Cogn. =Gurania subumbellata 

Gurania levyana var. glabra Cogn. =Gurania robusta 

Gurania multiflora (Miq.) Cogn. =Gurania lobata 

Gurania sagotiana Cogn. =Gurania reticulata 

Gurania spinulosa (Poepp. & Endl.) Cogn. =Gurania lobata 

Helmontia simplicifolia Cogn. =Helmontia leptantha 

Lagenaria lagenaria (L.) Cockerell =Lagenaria siceraria 

Lagenaria vulgaris Ser. =Lagenaria siceraria 

Luffa aegyptiaca Mill. =Luffa cylindrica 

Luffa operculata (L.) Cogn. [in S. Am. Records =L. sepium. L. operculata valid taxon for Cent. Am. & Mex. Species] =Luffa 

sepium 

Luffa purgans (Mart.) Mart. =Luffa sepium 

Melothria fluminensis Gardner =Melothria pendula 

Melothria guadalupensis (Spreng.) Cogn. =Melothria pendula 

Melothria heteroloba Pittier =Melothria trilobata 

Momordica balsamina L. [dubious, probably based on misidentification fide C. Jeffrey] 

Momordica cylindrica L. =Luffa cylindrica 

Momordica lanata Thunb. =Citrullus lanatus 

Momordica operculata L. =Luffa sepium 

Momordica purgans Mart. =Luffa sepium 

Nhandiroba cordifolia (L.) Kuntze =Fevillea cordifolia 

Perianthopodus bonplandii Cogn. =Cayaponia coriacea 

Rytidostylis carthaginensis (Jacq.) Kuntze [not in flora area] 

Trianosperm rigida Cogn. =Cayaponia rigida 

Trianosperma angustiloba Cogn. =Cayaponia angustiloba 

Trianosperma cruegeri Naudin =Cayaponia cruegeri 

Trianosperma triangularis Cogn. =Cayaponia triangularis 

Trichosanthes corniculata Lam. =Ceratosanthes palmata 

Trichosanthes palmata L. =Ceratosanthes palmata 

REFERENCES 

Jeffrey, C. 1978. Further notes on Cucurbitaceae: IV. Some New World taxa. Kew Bull. 33: 347-380. 

Jeffrey, C. 1984. Cucurbitaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 457-518. 

Jeffrey, C. & B. Trujillo. 1992. Cucurbitaceae. In G. Morillo, Flora de Venezuela 5(1): 11-201. 

Kearns, D.M. 1998. Cucurbitaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 431-461. 

Nee, M. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to 

the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 236-246. 

CUNONIACEAE 

P. Berry, J.C. Bradford 

Weinmannia balbisiana Kunth 

Weinmannia brachystachya Willd. ex Engl. [expected in GU][Mt. Roraima] 

Weinmannia corocoroensis J.C. Bradford & P.E. Berry 

Weinmannia elliptica Kunth 

Weinmannia fagaroides Kunth [Mt. Roraima] 

Weinmannia glabra L. f. [Mt. Roraima] 

Weinmannia guyanensis Klotzsch ex Engl. 

Weinmannia guyanensis var. guyanensis [Mt. Roraima] 

Weinmannia ilutepuiensis P.E. Berry & J.C. Bradford 

Weinmannia laxiramea Killip & A.C. Sm. [Mt. Roraima] 

Weinmannia pubescens Kunth 

Weinmannia sorbifolia Kunth 

Weinmannia sorbifolia var. sorbifolia 

Weinmannia sp. A [Distinct from Weinmannia fagaroides] 

Weinmannia velutina O.C. Schmidt 


Weinmannia baccariniana var. caracasana Pamp. =Weinmannia fagaroides 

Weinmannia balbisiana var. moritzii (Engl.) Cuatrec. =Weinmannia balbisiana 

Weinmannia balbisiana var. ptariana Cuatrec. =Weinmannia elliptica 

Weinmannia balbisiana var. roraimensis Bernardi =Weinmannia elliptica 

Weinmannia cardonae Cuatrec. =Weinmannia elliptica 

Weinmannia caripensis Kunth =Weinmannia glabra 

AM BO GU? 

BO 

AM 

AM BO 

AM BO GU 

AM BO GU 

BO GU 

BO GU 

BO 

BO GU 

AM 

AM 

AM 

AM 

AM BO GU


Weinmannia glabra var. caripensis (Kunth) Engl. =Weinmannia glabra 

Weinmannia microphylla Kunth =Weinmannia brachystachya 

Weinmannia microphylla var. caracasana (Pamp.) Cuatrec. =Weinmannia fagaroides 

Weinmannia microphylla var. fagaroides (Kunth) Cuatrec. =Weinmannia fagaroides 

Weinmannia moritzii Engl. =Weinmannia balbisiana 

Weinmannia neblinensis Maguire & Steyerm. =Weinmannia pubescens 

Weinmannia ovalis var. elliptica (Kunth) Cuatrec. =Weinmannia elliptica 

Weinmannia ovalis var. roraimensis Pamp. =Weinmannia elliptica 

Weinmannia pinnata L. =Weinmannia glabra 

Weinmannia pinnata var. caripensis (Kunth) Cuatrec. =Weinmannia glabra 

Weinmannia pinnata var. ptaritepuiana Cuatrec. =Weinmannia glabra 

Weinmannia roraimensis Cuatrec. =Weinmannia fagaroides 

Weinmannia steyermarkii Cuatrec. =Weinmannia velutina 

Weinmannia tamana Cuatrec. =Weinmannia fagaroides 

Weinmannia velutina f. steyermarkii (Cuatrec.) Bernardi =Weinmannia velutina 

REFERENCES 

Bradford, J.C. & P.E. Berry. 1998. Cunoniaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 

4: 462-469. 

Killip, E.P. & A.C. Smith. 1929. The genus Weinmannia in northern South America. Bull. Torrey Bot. Club 56: 361-377. 

Cuscuta americana L. 

Cuscuta campestris Yunck. 

Cuscuta costaricensis Yunck. 

Cuscuta indecora Choisy 

Cuscuta umbellata Kunth 

CUSCUTACEAE 

D.F. Austin 

REFERENCES 

Austin, D.F. 1982. Convolvulaceae. In Flora de Venezuela. 

Austin, D.F. 1998. Cuscutaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 469-470. 

Yuncker, T.G. 1923. Revision of the South American species of Cuscuta. I. Am. J. Bot. 9: 557-575; II. Am. J. Bot. 10: 1-17. 

Cyrilla racemiflora L. 

Purdiaea nutans Planch. 

CYRILLACEAE 

J.S. Miller 


Alloiosepalum weberbaueri Gilg =Purdiaea nutans 

Cyrilla antillana Michx. =Cyrilla racemiflora 

Cyrilla brevifolia N.E. Br. =Cyrilla racemiflora 

Cyrilla perrottetii Briq. =Cyrilla racemiflora 

Cyrilla racemiflora var. brevifolia (N.E. Br.) Maguire & Wurdack ex Steyerm. =Cyrilla racemiflora 

Itea cyrilla L’Hér. =Cyrilla racemiflora 

Purdiaea weberbaueri (Gilg) Mattick =Purdiaea nutans 

REFERENCES 

Miller, J.S. 1998. Cyrillaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 663-665. 

Thomas, J.L. 1960. A monographic study of the Cyrillaceae. Contr. Gray Herb. 186: 1-114. 

Dichapetalum froesii Prance 

Dichapetalum latifolium Baill. 

Dichapetalum odoratum Baill. 

Dichapetalum pedunculatum (DC.) Baill. 

Dichapetalum rugosum (Vahl) Prance 

Dichapetalum schulzii Prance 

Tapura amazonica Poepp. 

DICHAPETALACEAE 

G.T. Prance 

BO 

BO 

BO 

SU 

GU SU 


AM BO 

AM 

AM 

AM BO 


BO GU SU FG 

SU 

AM GU SU FG

280 

Tapura amazonica var. amazonica 

Tapura capitulifera Spruce ex Baill. 

Tapura guianensis Aubl. 

Tapura singularis Ducke 



Chailletia flavicans Klotzsch =Dichapetalum rugosum 

Chailletia parkeri Planch. ex Triana & Planch. [nomen nudum] =Dichapetalum rugosum 

Chailletia pedunculata DC. =Dichapetalum pedunculatum 

Chailletia sessiliflora DC. =Tapura guianensis 

Chailletia vestita Benth. =Dichapetalum rugosum 

Dichapetalum amazonicum K. Krause =Dichapetalum latifolium 

Dichapetalum flavicans (Klotzsch) Engl. =Dichapetalum rugosum 

Dichapetalum glabrum (Vahl) Prance =Dichapetalum pedunculatum 

Dichapetalum scandens (Poir.) I.M. Johnst. =Dichapetalum rugosum 

Dichapetalum scandens var. cinerascens (Baill.) J.F. Macbr. =Dichapetalum latifolium 

Dichapetalum vestitum (Benth.) Baill. =Dichapetalum rugosum 

Dichapetalum vestitum var. cinerascens Baill. =Dichapetalum latifolium 

Rohria petioliflora Willd. =Tapura guianensis 

Rohria schreberi J.F. Gmel. =Tapura guianensis 

Rohria tapura J.F. Gmel. =Tapura guianensis 

Symphyllanthus glaber Vahl =Dichapetalum pedunculatum 

Symphyllanthus rugosus Vahl =Dichapetalum rugosum 

Tapura amazonica var. ciliata (Gardner) Baill. =Tapura amazonica 

Tapura amazonica var. cuspidata Baill. =Tapura amazonica 

Tapura amazonica var. dasyphylla Baill. =Tapura amazonica 

Tapura amazonica var. sublanceolata Baill. =Tapura amazonica 

Tapura ciliata Gardner =Tapura amazonica 

Tapura cucullata Benth. =Tapura guianensis 

Tapura negrensis Suess. =Tapura guianensis 

REFERENCES 

Prance, G.T. 1972. Dichapetalaceae. Flora Neotropica Monogr. 10: 1-84. 

Prance, G.T. 1979. A new species of Dichapetalum from Suriname. Bull. Torrey Bot. Club 106(4): 309-312. 

Prance, G.T. 1983. Additions to Neotropical Dichapetalaceae. Brittonia 35(1): 49-54. 

Prance, G.T. 1998. Dichapetalaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 666-671. 

Prance, G.T. 2002. Dichapetalaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


247-249. 

Stafleu, F.A. 1951. Dichapetalaceae. In A. Pulle, Flora of Suriname 3(2): 166-172. 

Curatella americana L. 

Davilla alata (Vent.) Briq. 

Davilla kunthii A. St.-Hil. 

Davilla lacunosa Mart. 

Davilla nitida (Vahl) Kubitzki 

Davilla rugosa Poir. 

Davilla rugosa var. rugosa 

Davilla steyermarkii Kubitzki [expected in GU] 

Doliocarpus amazonicus Sleumer 

Doliocarpus amazonicus subsp. amazonicus 

Doliocarpus areolatus Kubitzki 

Doliocarpus brevipedicellatus Garcke 

Doliocarpus brevipedicellatus subsp. brevipedicellatus 

Doliocarpus carnevaliorum Aymard 

Doliocarpus dentatus (Aubl.) Standl. 

Doliocarpus dentatus subsp. dentatus 

Doliocarpus dentatus subsp. esmeraldae (Steyerm.) Kubitzki 

Doliocarpus dentatus subsp. ferrugineus (Rusby) Kubitzki 

Doliocarpus dentatus subsp. lanceolatus Kubitzki 

Doliocarpus dentatus subsp. latifolius Kubitzki 

DILLENIACEAE 

G.A. Aymard C., C.L. Kelloff 

AM GU SU FG 



FG 


GU SU FG 


FG 




BO 

SU 

SU 

AM 



AM 




AM 

AM 

AM BO

Doliocarpus dentatus subsp. obovatus Aymard 

Doliocarpus dentatus subsp. undulatus (Eichler) Kubitzki 

Doliocarpus gracilis Kubitzki 

Doliocarpus guianensis (Aubl.) Gilg 

Doliocarpus hispidobaccatus Aymard 

Doliocarpus leiophyllus Kubitzki 

Doliocarpus liesneri Aymard 

Doliocarpus macrocarpus Mart. ex Eichler 

Doliocarpus major J.F. Gmel. 

Doliocarpus major subsp. major 

Doliocarpus multiflorus Standl. 

Doliocarpus novogranatensis Kubitzki 

Doliocarpus ortegae Aymard 

Doliocarpus paraensis Sleumer 

Doliocarpus paucinervis Kubitzki 

Doliocarpus prancei Kubitzki 

Doliocarpus sagotianus Kubitzki 

Doliocarpus savannarum Sandwith 

Doliocarpus spraguei Cheesman 

Doliocarpus verruculosus Kubitzki 

Neodillenia venezuelana Aymard 

Pinzona coriacea Mart. & Zucc. 

Tetracera asperula Miq. 

Tetracera costata Mart. ex Eichler 

Tetracera costata subsp. costata 

Tetracera costata subsp. rotundifolia (Sm.) Kubitzki 

Tetracera maguirei Aymard & B.M. Boom 

Tetracera surinamensis Miq. 

Tetracera tigarea DC. 

Tetracera volubilis L. 

Tetracera volubilis subsp. volubilis 

Tetracera willdenowiana Steud. 

Tetracera willdenowiana subsp. emarginata Kubitzki 

Tetracera willdenowiana subsp. willdenowiana 



Calinea scandens Aubl. =Doliocarpus spraguei 

Curatella americana var. pentagyna Donn. Sm. =Curatella americana 

Curatella cambaiba A. St.-Hil. =Curatella americana 

Curatella grisebachiana Eichler =Curatella americana 

Davilla aspera (Aubl.) Benoist =Davilla kunthii 

Davilla multiflora (DC.) A. St.-Hil. =Davilla nitida 

Davilla pilosa Miq. =Davilla rugosa 

Davilla surinamensis Miq. =Davilla kunthii 

Davilla tenuis Eichler =Davilla kunthii 

Davilla wormiaefolia Baill. =Davilla alata 

Doliocarpus belizensis Lundell =Pinzona coriacea 

Doliocarpus brevipedicellatus var. densiflorus Eichler =Doliocarpus dentatus subsp. undulatus 

Doliocarpus calinea J.F. Gmel. =Doliocarpus spraguei 

Doliocarpus coriaceus (Mart. & Zucc.) Gilg =Pinzona coriacea 

Doliocarpus densiflorus (Eichler) Steyerm. =Doliocarpus dentatus subsp. undulatus 

Doliocarpus esmeraldae Steyerm. =Doliocarpus dentatus subsp. esmeraldae 

Doliocarpus ferrugineus Rusby =Doliocarpus dentatus subsp. ferrugineus 

Doliocarpus ptariensis Steyerm. =Doliocarpus savannarum 

Doliocarpus rolandri J.F. Gmel. =Doliocarpus major 

Doliocarpus scandens Poir. =Doliocarpus major 

Doliocarpus scandens (Aubl.) Gilg =Doliocarpus spraguei 

Doliocarpus surinamensis Lanj. =Doliocarpus paraensis 

Doliocarpus undulatus Eichler =Doliocarpus dentatus subsp. undulatus 

Doliocarpus virgatus Sagot =Doliocarpus macrocarpus 

Soramia guianensis Aubl. =Doliocarpus guianensis 

Tetracera aspera (Aubl.) Willd. =Tetracera tigarea 

Tetracera multiflora DC. =Davilla nitida 

Tetracera nitida Vahl =Davilla nitida 

AM 

AM 

SU FG 

AM 

AM 

AM 

AM 



AM GU SU FG 

AM 

AM 

AM 

AM 

GU SU FG 

AM 

BO 

GU SU FG 

FG 


AM 

AM 

AM 

BO GU SU FG 


GU SU FG 

BO GU SU FG 

BO GU SU FG 

BO GU 

GU 

FG 






AM FG 

AM BO GU SU

282 


Tetracera ovalifolia auct. non DC. =Tetracera asperula 

Tetracera ovalifolia DC. =Tetracera costata subsp. rotundifolia 

Tetracera rotundifolia auct. non Sm. =Tetracera asperula 

Tetracera rotundifolia Sm. =Tetracera costata subsp. rotundifolia 

Tetracera trinitensis Sprague & R.O. Williams =Tetracera costata 

Tigarea aspera Aubl. =Tetracera tigarea 

Tigarea dentata Aubl. =Doliocarpus dentatus 

REFERENCES 

Aymard, G. 1998. Dilleniaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 671-685. 

Aymard, G. & J. Miller. 1994. Dilleniaceae novae Neotropicae III. Sinopsis y adiciones a las Dilleniaceae del Perú. Candollea 

49(1): 169-182. 

Aymard, G. & S.A. Mori. 2002. Dilleniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 250-253. 

Kubitzki, K. 1970. Die gattung Tetracera. Mitt. Bot. Münch. 8: 1-98. 

Kubitzki, K. 1971. Doliocarpus, Davilla und verwandte gattungen. Mitt. Bot. Münch. 9: 1-105. 

Lanjouw, J. & P.F. van Heerdt. 1941. Dilleniaceae. In A. Pulle, Flora of Suriname 3(1): 386-408. 

DROSERACEAE 

A. Culham, R.D. de Stefano 

Drosera arenicola Steyerm. 

Drosera biflora Willd. ex Roem. & Schult. 

Drosera capillaris Poir. 

Drosera cayennensis Sagot ex Diels 

Drosera communis A. St.-Hil. 

Drosera esmeraldae (Steyerm.) Maguire & Wurdack 

Drosera felix Steyerm. & L.B. Sm. 

Drosera hirticalyx Duno & Culham 

Drosera intermedia Hayne 

Drosera kaieteurensis Brumm.-Ding. 

Drosera meristocaulis Maguire & Wurdack 

Drosera roraimae (Klotzsch ex Diels) Maguire & J.R. Laundon 

Drosera sessilifolia A. St.-Hil. 

Drosera yutajensis Duno & Culham 


Drosera americana Willd. =Drosera intermedia 

Drosera arenicola var. occidentalis Maguire & Wurdack =Drosera arenicola 

Drosera colombiana A. Fernández =Drosera cayennensis 

Drosera dentata Benth. =Drosera sessilifolia 

Drosera montana var. robusta Diels =Drosera roraimae 

Drosera montana var. roraimae Klotzsch ex Diels =Drosera roraimae 

Drosera panamensis M.D. Correa & A.S. Taylor =Drosera cayennensis 

Drosera pusilla Kunth =Drosera biflora 

Drosera sanariapoana Steyerm. =Drosera cayennensis 

Drosera tenella Willd. ex Roem. & Schult. =Drosera capillaris 

Drosera tenella var. esmeraldae Steyerm. =Drosera esmeraldae 

REFERENCES 

Brummer-Dinger, C.A. 1955. Notes on Guiana Droseraceae. Acta Bot. Neerl. 4(1): 136-138. 

Duno de Stefano, R. & A. Culham. 1998. Droseraceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan 

Guayana 4: 697-703. 

Maguire, B. & J.J. Wurdack. 1957. Droseraceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland 

part 2. Mem. New York Bot. Gard. 9(3): 331-336. 

Diospyros artanthifolia Mart. ex Miq. 

Diospyros capreifolia Mart. ex Hiern 

Diospyros carbonaria Benoist 

Diospyros cauliflora Mart. 

EBENACEAE 

P.E. Berry, C. Sothers 

AM BO 

AM GU SU 


AM BO GU FG 

AM BO 

AM GU 

BO 

AM 

BO GU SU 

BO GU 

AM 

AM BO GU 

AM BO GU 

AM 

AM GU SU FG 


FG 

FG

Diospyros cauligera F. White 

Diospyros cavalcantei Sothers 

Diospyros cayennensis A. DC. 

Diospyros dichroa Sandwith 

Diospyros discolor Willd. [cultivated; naturalized?] 

Diospyros duckei Sandwith 

Diospyros glomerata Spruce ex Hiern 

Diospyros guianensis (Aubl.) Gürke 

Diospyros guianensis subsp. akaraiensis (A.C. Sm.) F. White 

Diospyros guianensis subsp. coriacea F. White 

Diospyros guianensis subsp. guianensis 

Diospyros inconstans Jacq. 

Diospyros lissocarpoides Sandwith 

Diospyros martinii Benoist ex Amshoff 

Diospyros matheriana A.C. Sm. 

Diospyros poeppigiana A. DC. 

Diospyros ropourea B. Walln. 

Diospyros santaremnensis Sandwith [perhaps expected in FG] 

Diospyros sericea A. DC. 

Diospyros sp. A 

Diospyros sprucei Hiern 

Diospyros tenuiflora A.C. Sm. 

Diospyros tetrandra Hiern 

Diospyros vestita Benoist 



Diospyros akaraiensis A.C. Sm. =Diospyros guianensis subsp. akaraiensis 

Diospyros anzoateguiensis Steyerm. =Diospyros inconstans 

Diospyros delgadoi Standl. =Diospyros inconstans 

Diospyros emarginata Hiern =Diospyros poeppigiana 

Diospyros ierensis Britton =Diospyros cayennensis 

Diospyros liriosmoides A.C. Sm. =Diospyros poeppigiana 

Diospyros longifolia (Spruce ex Engl.) Sleumer & F. White =Diospyros tenuiflora 

Diospyros melinonii (Hiern) A.C. Sm. =Diospyros capreifolia 

Diospyros paralea Steud. =Diospyros guianensis 

Diospyros polyandra Spruce ex Hiern =Diospyros artanthifolia 

Diospyros praetermissa Sandwith =Diospyros vestita 

Diospyros subrotata Hiern =Diospyros artanthifolia 

Maba inconstans (Jacq.) Griseb. =Diospyros inconstans 

Maba mellinoni Hiern =Diospyros capreifolia 

Paralea guianensis Aubl. =Diospyros guianensis 

Ropourea guianensis Aubl. =Diospyros ropourea 

AM 


FG 

FG 

GU SU FG 

FG 

FG? 

AM FG 


GU? FG 

BO GU 

AM 

BO 

DA GU SU FG 


SU FG 

AM 

GU SU FG 

FG 

REFERENCES 

Cavalcante, P.B. 1963. Contribuçao ao conhecimento do gênero Diospyros Dalech. na Amazônia. Bol. Mus. Goeldi n.s. 20: 1- 

53. 

Sothers, C. & P.E. Berry. 1998. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 704-712. 

Wallnöfer, B. 1999. Neue Diospyros-Arten (Ebenaceae) aus Südamerika. Ann. Naturhist. Mus. Wien, B, 101: 565-592. 

Wallnöfer, B. 2000. Neue Diospyros-Arten (Ebenaceae) aus Südamerika - II. Ann. Naturhist. Mus. Wien, B, 102: 417-433. 

Wallnöfer, B. 2001. Lectotypification of Diospyros cayennensis A. DC. (Ebenaceae). Taxon 50: 887-889 (see Erratum in Taxon 

50 (4): 1319). 

Wallnöfer, B. 2001. The biology and systematics of Ebenaceae: a review. Ann Naturhist. Mus. Wein 103B: 485-512. 

Wallnöfer, B. 2003. A new species of Diospyros from southwestern Amazonia. Ann. Naturhist. Mus. Wien, B, 104: 563-566. 

Wallnöfer, B. 2004. Ebenaceae. In: Kubitzki, K. (ed.): The families and genera of vascular plants. Vol. 6: 125-130. Springer 

Verlag, Berlin. 

Wallnöfer, B. 2004. Lissocarpaceae. In: Kubitzki, K. (ed.): The families and genera of vascular plants. Vol. 6: 236-238. Springer 

Verlag, Berlin. 

Wallnöfer, B. 2004. A revision of Lissocarpa Benth. (Ebenaceae subfam. Lissocarpoideae (Gilg in Engler) B.Walln.). Ann. 

Naturhist. Mus. Wien, B, 105: 515-564. 

Wallnöfer, B. & S.A. Mori. 2002. Ebenaceae. In Mori S.A., Cremers G., Gracie C.A., Granville J.-J. de, Heald S.V., Hoff M. & 

Mitchell J.D., eds: Guide to the vascular plants of central French Guiana. Part 2. Dicotyledons. Mem. New York Bot. Gard. 

76 (2): 254-257. 

White, F. 1981. Ebenaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 11. Mem. New York Bot. Gard. 

32: 1-391. 

AM 

AM 

BO 

GU 

SU FG 

BO SU FG

284 

Sloanea bolivarensis Steyerm. 

Sloanea brevipes Benth. 

Sloanea carrenoi Steyerm. 

Sloanea cataniapensis Steyerm. 

Sloanea caudata Steyerm. 

Sloanea cavicola Steyerm. 

Sloanea crassifolia Earle Sm. 

Sloanea davidsei Steyerm. 

Sloanea durissima Spruce ex Benth. 

Sloanea echinocarpa Uittien 

Sloanea eichleri K. Schum. 

Sloanea floribunda Spruce ex Benth. 

Sloanea garckeana K. Schum. 

Sloanea gracilis Uittien 

Sloanea grandiflora Sm. 

Sloanea guianensis (Aubl.) Benth. 

Sloanea latifolia (Rich.) K. Schum. 

Sloanea laurifolia (Benth.) Benth. 

Sloanea laxiflora Spruce ex Benth. 

Sloanea longiaristata Steyerm. 

Sloanea macrophylla Benth. ex Turcz. 

Sloanea megacarpa Steyerm. & Marc.-Berti 

Sloanea merevariensis Pittier ex Steyerm. 

Sloanea nitida G. Don 

Sloanea obtusifolia (Moric.) K. Schum. 

Sloanea parviflora Planch. ex Benth. 

Sloanea parvifructa Steyerm. 

Sloanea picapica Standl. 

Sloanea picapica var. hirsuta C.E. Sm. 

Sloanea pittieriana Steyerm. 

Sloanea ptariana Steyerm. 

Sloanea pubescens Benth. 

Sloanea robusta Uittien 

Sloanea rufa Planch. ex Benth. 

Sloanea schomburgkii Spruce ex Benth. 

Sloanea sinemariensis Aubl. 

Sloanea sipapoana Steyerm. 

Sloanea sp. A 

Sloanea sp. B [Sierra de la Neblina] 

Sloanea sp. C 

Sloanea sp. D 

Sloanea spathulata Earle Sm. 

Sloanea steyermarkii Earle Sm. 

Sloanea steyermarkii subsp. autanae (Steyerm.) Steyerm. 

Sloanea steyermarkii subsp. jauaensis (Steyerm.) Steyerm. 

Sloanea steyermarkii subsp. steyermarkii 

Sloanea stipitata Spruce ex Benth. 

Sloanea subpsilocarpa Steyerm. 

Sloanea synandra Spruce ex Benth. 

Sloanea terniflora (Moç. & Sessé ex DC.) Standl. 

Sloanea trichosticha R.O. Williams & Sandwith 

Sloanea tuerckheimii Donn. Sm. 

Sloanea venezuelana Steyerm. 

Sloanea wurdackii Steyerm. 


ELAEOCARPACEAE 

L.J. Dorr, D.A. Smith, J.A. Steyermark† 


Ablania guianensis Aubl. =Sloanea guianensis 

Adenobasium obtusifolium Moric. =Sloanea obtusifolia 

Courimari guianensis Aubl. [name of uncertain application] =Sloanea sp. 

Dasynema laurifolium Benth. =Sloanea laurifolia 

Dasynema pubescens Poepp. =Sloanea pubescens 

BO 

AM GU SU? FG 

BO 

AM 

BO 

BO 

BO GU 

AM 

GU 

GU SU FG 

BO GU SU FG 

AM 

GU SU FG 

SU 



GU FG 

AM GU FG? 


AM 

AM GU FG? 

DA SU 

BO 

GU FG 

AM DA FG? 

GU SU FG 

AM 

GU 

GU 

BO 

BO 

AM BO SU? FG 

AM BO GU SU 

AM GU SU FG 

GU FG 

AM GU SU FG 

AM 

BO 

AM 

AM 

AM 

AM 

AM BO 

AM 

AM BO 

BO 

AM BO 

BO DA 

AM BO GU FG 

AM SU FG 

GU 

AM GU SU FG 

BO 

AM BO


Sloanea acutiflora Uittien =Sloanea laxiflora 

Sloanea amplifrons Johnst. =Sloanea grandiflora 

Sloanea autanae Steyerm. =Sloanea steyermarkii subsp. autanae 

Sloanea brachytepala Ducke [considered distinct by D.A. Sm. and Steyerm., in AM] =Sloanea laurifolia 

Sloanea bracteosa Ducke =Sloanea nitida 

Sloanea breviseta Steyerm. =Sloanea guianensis 

Sloanea corymbiflora DC. =Sloanea latifolia 

Sloanea dentata L. [not in flora area] 

Sloanea eriostemon Sprague & L. Riley =Sloanea tuerckheimii 

Sloanea jauaensis Steyerm. =Sloanea steyermarkii subsp. jauaensis 

Sloanea jauaensis var. minor Steyerm. =Sloanea steyermarkii subsp. jauaensis 

Sloanea maroana Steyerm. =Sloanea floribunda 

Sloanea massoni auct. non Sw. 1788: sensu Pittier et al. =Sloanea macrophylla 

Sloanea obtusa (Splitg.) K. Schum. =Sloanea parviflora 

Sloanea oppositifolia Spruce ex Benth. =Sloanea laurifolia 

Sloanea paniculata Spruce ex Benth. =Sloanea floribunda 

Sloanea sinemariensis auct. non Aubl. sensu Pittier et al. (as Sloanea surinamensis) =Sloanea eichleri 

Sloanea usurpatrix Sprague & L. Riley =Sloanea tuerckheimii 

Sloanea yapacanae Steyerm. =Sloanea steyermarkii subsp. jauaensis 

REFERENCES 

Heald, S.V., A. Vicentini, & D.A. Smith. 2002. Elaeocarpaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. 


Mem. New York Bot. Gard. 76 (2): 258-261. 

Jansen-Jacobs, M.J. 1986. Elaeocarpaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 3(1- 

2): 301-305. 

Smith, C.E., Jr. 1954. The New World species of Sloanea. Contr. Gray Herb. 175: 1-114. 

Smith, D.A. & J.A. Steyermark. 1998. Elaeocarpaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan 

Guayana 4: 712-730. 

Uittien, H. 1932. Elaeocarpaceae. In A. Pulle, Flora of Suriname 3(1): 58-64. 

Antidaphne antidaphneoides (Rizzini) Kuijt 

Antidaphne viscoidea Poepp. & Endl. 

Eubrachion gracile Kuijt 

EREMOLEPIDACEAE 

J. Kuijt 


Antidaphne fendleri (Tiegh.) Engl. =Antidaphne viscoidea 

Antidaphne fendleri var. pseudoviscoidea Rizzini =Antidaphne viscoidea 

Antidaphne orbicularis Kuijt =Antidaphne viscoidea 

Antidaphne viscoidea var. alpestris Rizzini =Antidaphne viscoidea 

Eremolepis mirocalyx Ernst =Antidaphne viscoidea 

Ixidium antidaphneoides Rizzini =Antidaphne antidaphneoides 

Stachyphyllum fendleri Tiegh. =Antidaphne viscoidea 

REFERENCES 

Kuijt, J. 1988. Monograph of the Eremolepidaceae. Syst. Bot. Monogr. 18: 1-60. 

Kuijt, J. 1988. Eremolepidaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 731-734. 

Agarista duckei (Huber) Judd [expected in GU] 

Bejaria aestuans L. [primarily Andean taxon] 

Bejaria imthurnii N.E. Br. [Mt. Roraima] 

Bejaria neblinensis Maguire, Steyerm. & Luteyn 

Bejaria sprucei Meisn. 

Cavendishia callista Donn. Sm. 

Cavendishia neblinae Maguire, Steyerm. & Luteyn 

Cavendishia aff. pubescens (Kunth) Hemsl. 

Ceratostema glandulifera Maguire, Steyerm. & Luteyn 

Disterigma cuspidatum (Planch. ex Wedd.) Nied. 

ERICACEAE 

J.L. Luteyn 

BO 

BO GU 

BO 

AM BO 

AM BO 

BO GU? 

AM 

AM BO GU 


AM 

AM 

GU 

AM

286 


Disterigma humboldtii (Klotzsch) Nied. 

Gaultheria alnifolia (Dunal) A.C. Sm. 

Gaultheria alnifolia var. alnifolia 

Gaultheria erecta Vent. [Mt. Roraima] 

Gaultheria setulosa N.E. Br. [Mt. Roraima] 

Gaylussacia buxifolia Kunth 

Ledothamnus atroadenus Maguire, Steyerm. & Luteyn 

Ledothamnus decumbens Maguire, Steyerm. & Luteyn 

Ledothamnus guyanensis Meisn. [Mt. Roraima] 

Ledothamnus jauaensis Maguire, Steyerm. & Luteyn 

Ledothamnus luteus Maguire, Steyerm. & Luteyn 

Ledothamnus parviflorus Gleason 

Ledothamnus sessiliflorus N.E. Br. [Mt. Roraima] 

Mycerinus chimantensis Maguire, Steyerm. & Luteyn 

Mycerinus sclerophyllus A.C. Sm. 

Mycerinus viridiflorus Steyerm. & Maguire 

Notopora auyantepuiensis Steyerm. 

Notopora cardonae A.C. Sm. 

Notopora chimantensis Steyerm. & Maguire 

Notopora schomburgkii Hook. f. 

Notopora smithiana Steyerm. & Maguire 

Orthaea apophysata (Griseb.) A.C. Sm. 

Orthaea crinita A.C. Sm. 

Orthaea hispida A.C. Sm. 

Orthaea merumensis Maguire, Steyerm. & Luteyn 

Orthaea paruensis Maguire, Steyerm. & Luteyn 

Orthaea thibaudioides Maguire, Steyerm. & Luteyn 

Orthaea venamensis Maguire, Steyerm. & Luteyn 

Orthaea wurdackii Maguire, Steyerm. & Luteyn 

Pernettya marginata N.E. Br. [Mt. Roraima] 

Psammisia guianensis Klotzsch 

Psammisia hookeriana Klotzsch 

Psammisia urichiana (Britton) A.C. Sm. 

Satyria carnosiflora Lanj. 

Satyria cerander (Dunal) A.C. Sm. 

Satyria panurensis (Benth. ex Meisn.) Benth. & Hook. f. ex Nied. 

Sphyrospermum buxifolium Poepp. & Endl. 

Sphyrospermum cordifolium Benth. 

Sphyrospermum ellipticum Sleumer 

Sphyrospermum klotzschianum (Mansf.) A.C. Sm. 

Tepuia cardonae A.C. Sm. 

Tepuia intermedia Steyerm. 

Tepuia multiglandulosa Steyerm. & Maguire 

Tepuia speciosa A.C. Sm. 

Tepuia tatei Camp 

Tepuia vareschii Steyerm. 

Tepuia venusta Camp 

Thibaudia breweri Steyerm. & Maguire 

Thibaudia carrenoi Steyerm. & Maguire 

Thibaudia cupatensis Huber 

Thibaudia dolichandra Maguire, Steyerm. & Luteyn 

Thibaudia formosa (Klotzsch) Hoerold 

Thibaudia glandulifera A.C. Sm. 

Thibaudia involucrata A.C. Sm. 

Thibaudia longipes Maguire, Steyerm. & Luteyn 

Thibaudia nutans Klotzsch ex Mansf. 

Thibaudia smithiana Maguire, Steyerm. & Luteyn 

Thibaudia truncata A.C. Sm. 

Vaccinium chimantense Maguire, Steyerm. & Luteyn 

Vaccinium corymbodendron Dunal 

Vaccinium euryanthum A.C. Sm. 

Vaccinium puberulum Klotzsch ex Meisn. 

Vaccinium puberulum var. cardonae (A.C. Sm.) Maguire, Steyerm. & Luteyn 

Vaccinium puberulum var. hitchcockii (Camp) Maguire, Steyerm. & Luteyn 

AM BO GU 

BO 

BO 

AM BO GU 

AM BO GU? 

AM 

BO 

BO 

AM BO GU? 

BO 

BO 

AM 

BO GU? 

BO 

AM 

BO 

BO 

BO 

BO 

BO GU 

BO 

GU 

BO GU 

BO GU 

BO GU 

AM 

AM 

BO GU 

AM 

AM BO 

AM BO GU 

AM BO 

AM BO GU 

AM BO GU SU 

SU? FG 


AM BO GU FG 


GU 

AM BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

AM 

AM 

AM BO GU 

AM 

AM BO 

AM 

AM BO GU SU 

AM 

AM 

BO 

BO GU 

BO GU 

AM BO GU 

BO 

AM

Vaccinium puberulum var. jauaense Maguire, Steyerm. & Luteyn 

Vaccinium puberulum var. puberulum 


Vaccinium puberulum var. spathulatum (Camp & A.C. Sm.) Maguire, Steyerm. 

& Luteyn 

Vaccinium puberulum var. subcrenulatum (Klotzsch ex Meisn.) Maguire, 

Steyerm. & Luteyn 

Vaccinium puberulum var. tepuiense (Camp & A.C. Sm.) Maguire, Steyerm. & 

Luteyn 

Vaccinium roraimense N.E. Br. [Mt. Roraima] 

Vaccinium steyermarkii Luteyn 


Befaria spp. =Bejaria spp., nom. cons. prop. “Befaria” 

Bejaria coarctata Bonpl. =Bejaria aestuans 

Bejaria glauca Bonpl. =Bejaria aestuans 

Bejaria glauca var. coarctata (Bonpl.) Mansf. & Sleumer =Bejaria aestuans 

Bejaria glauca var. setosa Mansf. & Sleumer =Bejaria aestuans 

Bejaria glauca var. tomentella Mansf. & Sleumer =Bejaria aestuans 

Bejaria guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Bejaria sprucei 

Bejaria guianensis Klotzsch ex N.E. Br. =Bejaria sprucei 

Bejaria hispida Poepp. & Endl. =Bejaria aestuans 

Bejaria puberula Klotzsch ex Mansf. & Sleumer =Bejaria sprucei 

Bejaria schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum] =Bejaria sprucei 

Bejaria schomburgkiana Klotzsch ex Mansf. & Sleumer =Bejaria sprucei 

Bejaria tatei Gleason =Bejaria imthurnii 

Bejaria variabilis A.C. Sm. =Bejaria sprucei 

Brossea lindeniana (Planch.) Kuntze =Gaultheria alnifolia var. alnifolia 

Brossea roraimae (Klotzsch ex Meisn.) Kuntze =Gaultheria erecta 

Cavendishia duidae A.C. Sm. =Cavendishia callista 

Cavendishia gleasoniana A.C. Sm. =Cavendishia callista 

Cavendishia phelpsiae Camp =Cavendishia callista 

Cavendishia salicifolia Maguire, Steyerm. & Luteyn =Cavendishia callista 

Cavendishia urichiana Britton =Psammisia urichiana 

Ceratostema ulei Mansf. =Thibaudia nutans 

Chupalon pubescens (Kunth) Kuntze =Cavendishia aff. pubescens 

Epigaea cordifolia Sw. [not in flora area] =Gaultheria domingensis 

Gaultheria cordifolia Kunth =Gaultheria erecta 

Gaultheria lepida A.C. Sm. =Gaultheria erecta 

Gaultheria lindeniana Planch. =Gaultheria alnifolia var. alnifolia 

Gaultheria odorata Bredem. ex Willd. =Gaultheria erecta 

Gaultheria odorata Kunth =Gaultheria erecta 

Gaultheria roraimae Klotzsch ex Meisn. =Gaultheria erecta 

Gaultheria scabra Willd. =Gaultheria erecta 

Gaultheria tatei A.C. Sm. =Gaultheria erecta 

Gaylussacia cacuminis A.C. Sm. =Gaylussacia buxifolia 

Ledothamnus guyanensis var. nitidus A.C. Sm. =Ledothamnus guyanensis 

Ledothamnus guyanensis subsp. tatei (A.C. Sm.) Maguire, Steyerm. & Luteyn =Ledothamnus guyanensis 

Ledothamnus stenopetalus Maguire, Steyerm. & Luteyn =Ledothamnus guyanensis 

Ledothamnus steyermarkii A.C. Sm. =Ledothamnus guyanensis 

Ledothamnus steyermarkii subsp. longisetus Maguire, Steyerm. & Luteyn =Ledothamnus guyanensis 

Ledothamnus tatei A.C. Sm. =Ledothamnus guyanensis 

Ledothamnus tatei var. puberulus A.C. Sm. =Ledothamnus guyanensis 

Leucothoe duckei Huber =Agarista duckei 

Leucothoe venezuelensis A.C. Sm. =Agarista duckei 

Orthaea pendula Tutin =Orthaea apophysata 

Orthaea pubifolia Maguire, Steyerm. & Luteyn =Orthaea hispida 

Proclesia pubescens (Kunth) Klotzsch =Cavendishia aff. pubescens 

Psammisia breweri Steyerm. & Maguire =Psammisia urichiana 

Psammisia coriacea N.E. Br. =Thibaudia nutans 

Psammisia formosa Klotzsch =Thibaudia formosa 

Psammisia leucostoma Benth. ex Meisn. =Psammisia guianensis 

Riedelia panurensis (Benth. ex Meisn.) Kuntze =Satyria panurensis 

Satyria maguirei Camp =Satyria carnosiflora 

Sophoclesia cordifolia (Benth.) Klotzsch =Sphyrospermum cordifolium 

BO 

AM BO GU 

BO 

AM BO GU 

AM BO GU 

AM BO 

BO

288 


Sophoclesia klotzschiana Mansf. =Sphyrospermum klotzschianum 

Sophoclesia major (Griseb.) Benth. & Hook. f. =Sphyrospermum cordifolium 

Sophoclesia nummulariaefolia Klotzsch =Sphyrospermum buxifolium 

Sophoclesia ovata Klotzsch =Sphyrospermum cordifolium 

Sophoclesia subscandens Klotzsch =Sphyrospermum cordifolium 

Sphyrospermum majus Griseb. =Sphyrospermum cordifolium 

Sphyrospermum roraimae Klotzsch =Sphyrospermum buxifolium 

Tepuia floribunda Camp =Tepuia venusta 

Tepuia tatei var. chimantana Steyerm. & Maguire =Tepuia tatei 

Thibaudia alnifolia Dunal =Gaultheria alnifolia 

Thibaudia jauaensis Steyerm. & Maguire =Thibaudia involucrata 

Thibaudia panurensis Benth. ex Meisn. =Satyria panurensis 

Thibaudia pubescens Kunth =Cavendishia aff. pubescens 

Thibaudia roraimae Mansf. =Thibaudia formosa 

Thibaudia ulei (Mansf.) A.C. Sm. =Thibaudia nutans 

Vaccinium cardonae A.C. Sm. =Vaccinium puberulum var. cardonae 

Vaccinium cuspidatum Planch. ex Wedd. =Disterigma cuspidatum 

Vaccinium hitchcockii Camp =Vaccinium puberulum var. hitchcockii 

Vaccinium humboldtii Klotzsch =Disterigma humboldtii 

Vaccinium spathulatum Camp & A.C. Sm. =Vaccinium puberulum var. spathulatum 

Vaccinium subcrenulatum Klotzsch ex Meisn. =Vaccinium puberulum var. subcrenulatum 

Vaccinium tepuiense Camp & A.C. Sm. =Vaccinium puberulum var. tepuiense 

REFERENCES 

Lanjouw, J. 1934. Ericaceae. In A. Pulle, Flora of Suriname 4(1): 299-301. 

Lanjouw, J. 1937. Ericaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 492-493. 

Luteyn, J.L. 1983. Ericaceae Part I. Cavendishia. Flora Neotrop. Monogr. 35: 1-290. 

Luteyn, J.L. 1987. Orthaea: new species and redefinition of the genus. Nord. J. Bot. 7(1): 31-37. 

Luteyn, J.L. (ed.). 1995. Ericaceae - Part II. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae, 

and Vaccinioideae p.p.). Flora Neotropica Monogr. 66: 1-560. 

Luteyn, J.L. 1998. Ericaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 735-769. 

Luteyn, J.L. 2002. Ericaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


263. 

Maguire, B., J.A. Steyermark, & J.L. Luteyn. 1978. Ericaceae. In B. Maguire & collaborators, Botany of the Guayana Highland. 


Sleumer, H.O. 1936. Die Arten der Gattung Vaccinium L. in Zentral- und Südamerika. Notizbl. Bot. Gart. Berlin-Dahlem 13: 

111-140. 

Smith, A.C. 1932. The American species of Thibaudieae. Contr. U.S. Nat. Herb. 28: 311-547. 

Smith, A.C. 1933. The genera Sphyrospermum and Disterigma. Brittonia 1: 203-232. 

Erythroxylum amazonicum Peyr. 

Erythroxylum amplum Benth. 

Erythroxylum brevipes DC. 

Erythroxylum campestre A. St.-Hil. 

Erythroxylum cataractarum Spruce ex Peyr. 

Erythroxylum citrifolium A. St.-Hil. 

Erythroxylum coelophlebium Mart. 

Erythroxylum cumanense Kunth 

Erythroxylum divaricatum Peyr. 

Erythroxylum fimbriatum Peyr. 

Erythroxylum foetidum Plowman 

Erythroxylum gracilipes Peyr. 

Erythroxylum grisebachii Peyr. 

Erythroxylum guanchezii Plowman 

Erythroxylum havanense Jacq. 

Erythroxylum hypoleucum Plowman 

Erythroxylum impressum O.E. Schulz 

Erythroxylum kapplerianum Peyr. 

Erythroxylum lenticellosum Huber 

Erythroxylum leptoneurum O.E. Schulz 

ERYTHROXYLACEAE 

P.E. Berry, T. Plowman† 


GU SU FG 

GU? 

GU? 

AM 


SU 

GU SU FG 

AM BO GU 

BO FG 

AM 


SU 

AM 

BO GU 

AM 

AM BO GU 


AM 

FG


Erythroxylum ligustrinum DC. 

Erythroxylum lindemanii Plowman 

Erythroxylum lineolatum DC. 

Erythroxylum macrophyllum Cav. 

Erythroxylum macrophyllum var. macrophyllum 

Erythroxylum macrophyllum var. savannarum Plowman 

Erythroxylum mucronatum Benth. 

Erythroxylum oreophilum (O.E. Schulz ex Pilg.) Steyerm. & Maguire 

Erythroxylum orinocense Kunth 

Erythroxylum roraimae Klotzsch ex O.E. Schulz 

Erythroxylum rufum Cav. 

Erythroxylum schomburgkii Peyr. 

Erythroxylum spruceanum Peyr. 

Erythroxylum squamatum Sw. 

Erythroxylum steyermarkii Plowman 

Erythroxylum suberosum A. St.-Hil. 

Erythroxylum aff. subrotundum A. St.-Hil. 

Erythroxylum vernicosum O.E. Schulz 

Erythroxylum williamsii Standl. ex Plowman 


Erythroxylum acutifolium Steud. ex Peyr. =Erythroxylum citrifolium 

Erythroxylum albertianum Kuhlm. & W.A. Rodrigues =Erythroxylum mucronatum 

Erythroxylum areolatum Vell. =Erythroxylum suberosum 

Erythroxylum aristigerum Peyr. =Erythroxylum squamatum 

Erythroxylum aristigerum var. bahiense (Peyr.) O.E. Schulz =Erythroxylum squamatum 

Erythroxylum bahiense Peyr. =Erythroxylum squamatum 

Erythroxylum christii O.E. Schulz [nomen nudum] =Erythroxylum aff. subrotundum 

Erythroxylum coelophlebium var. grisebachii (Peyr.) O.E. Schulz =Erythroxylum grisebachii 

Erythroxylum comosum O.E. Schulz =Erythroxylum mucronatum 

Erythroxylum cuatrecasasii W.A. Gentner =Erythroxylum gracilipes 

Erythroxylum duckei Huber =Erythroxylum citrifolium 

Erythroxylum ectinocalyx Klotzsch =Erythroxylum divaricatum 

Erythroxylum filipes Huber =Erythroxylum macrophyllum 

Erythroxylum floribundum Mart. =Erythroxylum macrophyllum 

Erythroxylum gomphioides Planch. & Linden ex Triana & Planch. =Erythroxylum citrifolium 

Erythroxylum havanense var. continentis O.E. Schulz =Erythroxylum havanense 

Erythroxylum kirkianum O.E. Schulz =Erythroxylum mucronatum 

Erythroxylum laurinum Planch. & Linden ex Triana & Planch. =Erythroxylum macrophyllum 

Erythroxylum ligustrinum var. grandifolium Sagot =Erythroxylum kapplerianum 

Erythroxylum lucidum Kunth =Erythroxylum macrophyllum 

Erythroxylum mapuerae Huber =Erythroxylum kapplerianum 

Erythroxylum micranthum Bong. ex Peyr. =Erythroxylum citrifolium 

Erythroxylum novogranatense var. macrophyllum O.E. Schulz =Erythroxylum gracilipes 

Erythroxylum obtusum DC. =Erythroxylum havanense 

Erythroxylum ovatum Cav. =Erythroxylum havanense 

Erythroxylum paraense Peyr. =Erythroxylum citrifolium 

Erythroxylum recurrens Huber =Erythroxylum gracilipes 

Erythroxylum squamatum var. emarginatum Peyr. =Erythroxylum vernicosum 

Erythroxylum squamatum var. microcarpum O.E. Schulz =Erythroxylum squamatum 

Erythroxylum squamatum var. orinocense Kuntze =Erythroxylum impressum 

Erythroxylum squarrosum Klotzsch [nomen nudum] =Erythroxylum rufum 

Erythroxylum testaceum Peyr. =Erythroxylum suberosum 

Erythroxylum trinerve Huber =Erythroxylum squamatum 

Erythroxylum venezuelense Steyerm. =Erythroxylum mucronatum 

Erythroxylum vernicosum var. oreophilum O.E. Schulz ex Pilg. =Erythroxylum oreophilum 

AM SU FG 

AM GU SU 




AM BO 


AM BO GU 

AM BO GU 


AM BO GU FG 

BO GU 

AM 


BO 

BO GU FG 

BO 

AM BO GU FG? 

AM BO 

REFERENCES 

Heald, S.V. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide 


Plowman, T. & P.E. Berry. 1999. Erythroxylaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Schulz, O.E. 1907. Erythroxylaceae. In A. Engler, Das Pflanzenreich IV(134): 1-176. 

Westhoff, V. 1942. Erythroxylaceae. In A. Pulle, Flora of Suriname 3(2): 1-12.

290 


EUPHORBIACEAE 

W.S. Armbruster, P.E. Berry, H.-J. Esser, L.J. Gillespie, W.J. Hayden, 

S.V. Heald, G.A. Levin, W. Punt, R. de S. Secco, G.L. Webster† 

Acalypha alopecuroides Jacq. 

Acalypha arvensis Poepp. 

Acalypha cuneata Poepp. 

Acalypha diversifolia Jacq. 

Acalypha indica L. [naturalized] 

Acalypha lanceolata Willd. [naturalized] 

Acalypha macrostachya Jacq. 

Acalypha poiretii Spreng. 

Acalypha scandens Benth. 

Acalypha schiedeana Schltdl. 

Acalypha setosa A. Rich. 

Acalypha villosa Jacq. 

Acidoton nicaraguensis (Hemsl.) G.L. Webster 

Actinostemon amazonicus Pax & K. Hoffm. 

Actinostemon schomburgkii (Klotzsch) Hochr. 

Adelia triloba (Müll. Arg.) Hemsl. 

Adenophaedra grandifolia (Klotzsch) Müll. Arg. 

Alchornea castaneifolia (Willd.) A. Juss. 

Alchornea discolor Poepp. 

Alchornea fluviatilis Secco 

Alchornea glandulosa Poepp. 

Alchornea grandiflora Müll. Arg. 

Alchornea triplinervia (Spreng.) Müll. Arg. 

Alchorneopsis floribunda (Benth.) Müll. Arg. 

Amanoa almerindae Leal 

Amanoa congesta W.J. Hayden 

Amanoa cupatensis Huber 

Amanoa glaucophylla Müll. Arg. 

Amanoa guianensis Aubl. 

Amanoa neglecta W.J. Hayden 

Amanoa oblongifolia Müll. Arg. 

Amanoa steyermarkii Jabl. 

Antidesma ghaesembilla Gaertn. [cultivated, naturalized] 

Aparisthmium cordatum (A. Juss.) Baill. 

Astrococcus cornutus Benth. 

Bernardia amazonica Croizat [specimens not seen] 

Bernardia sidoides (Klotzsch) Müll. Arg. 

Caperonia castaneifolia (L.) A. St.-Hil. 

Caperonia corchoroides Müll. Arg. [=Caperonia palustris?] 

Caperonia paludosa Klotzsch [=Caperonia castaneifolia in FVG] 

Caperonia palustris (L.) A. St.-Hil. 

Caperonia stenophylla Müll. Arg. 

Celianella montana Jabl. 

Chaetocarpus schomburgkianus (Kuntze) Pax & K. Hoffm. 

Cnidoscolus urens (L.) Arthur 

Conceveiba guianensis Aubl. 

Conceveiba latifolia Benth. 

Conceveiba martiana Baill. 

Conceveiba ptariana (Steyerm.) Jabl. 

Croton argenteus L. 

Croton argyrophylloides Müll. Arg. 

Croton argyrophyllus Kunth 

Croton bolivarensis Croizat 

Croton cajucara Benth. 

Croton conduplicatus Kunth 

Croton cuneatus Klotzsch 

Croton aff. draconoides Müll. Arg. 

Croton essequiboensis Klotzsch [=Croton populifolius P. Mill.?] 

Croton fragrans Kunth 

Croton galeopsifolius Lanj. 

AM BO 

AM GU SU FG 

AM BO 


GU FG 

GU 


FG 

BO DA GU SU 

BO 

AM 

AM BO GU 

BO GU 

AM 


AM 

BO GU FG 

AM BO DA 


GU 

AM BO 

DA 



AM 

FG 

AM 

AM 


SU FG 

AM 

AM BO 

GU 


AM 

AM BO FG 

GU 


SU FG 

GU 

GU SU FG 

GU SU FG? 

AM BO 




AM 

AM GU FG 

AM BO 

BO 

FG 

AM BO GU 

BO DA 

AM BO GU 

BO GU 


FG 

BO GU 

AM BO 

GU


Croton gossypiifolius Vahl 

Croton guaiquinimae Steyerm. 

Croton guianensis Aubl. 

Croton hirtus L’Hér. 

Croton hostmannii Miq. ex Schltdl. 

Croton icabarui Jabl. 

Croton kaieteuri Jabl. [probably conspecific with Croton cuneatus] 

Croton kavayanensis Steyerm. 

Croton lobatus L. 

Croton longiradiatus Lanj. 

Croton macradenis Görts & Punt 

Croton matourensis Aubl. 

Croton megalodendron Müll. Arg. 

Croton mollis Benth. 

Croton neblinae Jabl. 

Croton nepetifolius Baill. 

Croton nuntians Croizat 

Croton orinocensis Müll. Arg. 

Croton palanostigma Klotzsch 

Croton parodianus Croizat 

Croton polypleurus Croizat 

Croton potaroensis Lanj. 

Croton pullei Lanj. 

Croton pullei var. glabrior Lanj. 

Croton pullei var. pullei 

Croton rhamnifolius Kunth 

Croton roraimensis Croizat 

Croton roraimensis var. roraimensis 

Croton rottlerifolius Baill. 

Croton schiedeanus Schltdl. 

Croton scutatus P.E. Berry & J.F. Gaskin 

Croton sipaliwinensis Lanj. 

Croton sp. A 

Croton sp. B 

Croton sp. C 

Croton sp. D 

Croton sp. E 

Croton sp. F 

Croton sp. G 

Croton sp. H 

Croton spiraeifolius Jabl. 

Croton spruceanus Benth. 

Croton stahelianus Lanj. 

Croton subcoriaceus Jabl. 

Croton subincanus Müll. Arg. 

Croton subserratus Jabl. 

Croton trinitatis Millsp. 

Croton umbratilis Kunth 

Croton vergarenae (Jabl.) L.J. Gillespie 

Croton yavitensis Croizat 

Dalechampia affinis Müll. Arg. 

Dalechampia attenuistylus Armbr. 

Dalechampia brevicolumna Armbr. 

Dalechampia brownsbergensis G.L. Webster & Armbr. 

Dalechampia aff. cissifolia Poepp. 

Dalechampia dioscoreifolia Poepp. 

Dalechampia fragrans Armbr. 

Dalechampia heterobractea Armbr. [often confused with Dalechampia 

parvibracteata] 

Dalechampia liesneri Huft 

Dalechampia magnoliifolia Müll. Arg. 

Dalechampia megacarpa Armbr. 

Dalechampia micrantha Poepp. 

Dalechampia papillistigma Armbr. 

BO 

BO 

BO GU SU FG 


GU SU FG 

BO 

GU 

FG? 


SU FG 

GU SU FG 


BO 

AM 

AM 

BO 

DA GU FG 

AM GU 

AM BO GU FG 

AM GU 

GU 

AM BO GU 

AM GU? SU FG 

SU FG 

GU? SU 

GU 

BO GU 

GU 

FG? 

BO GU SU FG 

AM 

AM BO GU SU 

AM 

AM 

AM 

AM 

BO 

AM 

AM 

BO 

AM 

AM 

SU 

AM 

BO GU SU FG 

AM BO 


BO 

BO 

AM 


BO SU FG 

SU? FG 

BO DA SU 

GU 

AM BO FG 

SU FG 

BO GU SU FG 

AM 

AM BO GU FG 

AM BO 

BO SU FG 

BO

292 


Dalechampia parvibracteata Lanj. 

Dalechampia scandens L. 

Dalechampia stipulacea Müll. Arg. 

Dalechampia tenuiramea Müll. Arg. 

Dalechampia tiliifolia Lam. 

Dalechampia aff. triphylla Lam. 

Dendrothrix multiglandulosa Esser 

Dendrothrix yutajensis (Jabl.) Esser 

Discocarpus essequeboensis Klotzsch [=”essequiboensis”] 

Discocarpus gentryi S.M. Hayden 

Discocarpus spruceanus Müll. Arg. 

Ditaxis polygama (Jacq.) L.C. Wheeler 

Dodecastigma amazonicum Ducke 

Dodecastigma integrifolium (Lanj.) Lanj. & Sandwith 

Drypetes fanshawei Sandwith 

Drypetes monachinoi Jabl. 

Drypetes variabilis Uittien 

Euphorbia bahiensis (Klotzsch & Garcke) Boiss. 

Euphorbia blodgettii Engelm. [adventitious] 

Euphorbia comosa Vell. 

Euphorbia cotinifolia L. 

Euphorbia cotinifolia subsp. cotinifolia 

Euphorbia cotinifolia subsp. cotinoides (Miq.) Christenh. 

Euphorbia dioeca Kunth 

Euphorbia graminea Jacq. [adventitious] 

Euphorbia heterophylla L. 

Euphorbia heterophylla var. cyathophora (Murray) Griseb. [native] 

Euphorbia heterophylla var. heterophylla [native] 

Euphorbia hirta L. 

Euphorbia humayensis Brandegee 

Euphorbia hypericifolia L. 

Euphorbia hyssopifolia L. 

Euphorbia hyssopifolia var. pubescenticocca Christenh. 

Euphorbia aff. laurifolia Juss. 

Euphorbia mesembrianthemifolia Jacq. [adventitous] 

Euphorbia neriifolia L. [cultivated, naturalized] 

Euphorbia oerstediana (Klotzsch & Garcke) Boiss. 

Euphorbia prostrata Aiton 

Euphorbia pulcheririma Willd. ex Klotzsch [cultivated, naturalized] 

Euphorbia serpens Kunth 

Euphorbia spruceana Boiss. 

Euphorbia thymifolia L. 

Euphorbia tirucalli L. [cultivated, naturalized] 

Gavarretia terminalis Baill. 

Glycydendron amazonicum Ducke 

Gymnanthes hypoleuca Benth. 

Haematostemon coriaceus (Baill.) Pax & K. Hoffm. 

Haematostemon guianensis Sandwith 

Hevea benthamiana Müll. Arg. 

Hevea brasiliensis (Willd. ex A. Juss.) Müll. Arg. [widely cultivated, possibly 

indigenous in FG] 

Hevea guianensis Aubl. 

Hevea guianensis var. guianensis 

Hevea guianensis var. lutea (Spruce ex Benth.) Ducke & R.E. Schult. 

Hevea microphylla Ule 

Hevea pauciflora (Spruce ex Benth.) Müll. Arg. 

Hevea pauciflora var. coriacea Ducke 

Hieronyma alchorneoides Allemno 

Hieronyma alchorneoides var. alchorneoides 

Hieronyma alchorneoides var. stipulosa Franco 

Hieronyma oblonga (Tul.) Müll. Arg. 

Hura crepitans L. 

Jablonskia congesta (Benth. ex Müll. Arg.) G.L. Webster 

Jatropha curcas L. [cultivated, naturalized] 

BO GU 


FG 

AM 

AM BO GU FG 

SU FG 

AM 

AM BO 

AM GU SU FG 

AM 

AM SU 

BO 

GU 

GU FG 

DA GU 

BO 


FG 

BO SU? 



BO GU SU FG 

BO GU FG 

FG 


SU FG 

AM BO GU SU 


BO 

BO GU SU FG 


GU 

GU 

FG? 

GU SU 


GU FG 

BO 


AM BO GU 

AM GU SU FG 

AM GU FG 

AM 

GU 

AM 

FG? 




AM 

AM BO GU SU 

AM BO GU SU 


GU SU FG 

DA FG 


BO GU SU FG 

AM GU SU FG? 

BO GU SU FG


Jatropha gossypiifolia L. 

Mabea angularis Hollander 

Mabea anomala Müll. Arg. 

Mabea arenicola Esser 

Mabea biglandulosa Baill. ex Müll. Arg. 

Mabea frutescens Jabl. 

Mabea linearifolia Jabl. 

Mabea longibracteata Esser 

Mabea montana Müll. Arg. 

Mabea nitida Spruce ex Benth. 

Mabea occidentalis Benth. [not in flora area] 

Mabea pakaraimaea Esser, sp. nov. ined. 

Mabea piriri Aubl. 

Mabea pulcherrima Müll. Arg. 

Mabea rubicunda Jabl. 

Mabea salicoides Esser 

Mabea sp. A 

Mabea sp. B [expected in BO. Base of Roraima on Guyana side] 

Mabea speciosa Müll. Arg. 

Mabea speciosa subsp. A 

Mabea speciosa subsp. B 

Mabea speciosa subsp. speciosa 

Mabea subsessilis Pax & K. Hoffm. 

Mabea taquari Aubl. 

Mabea trianae Pax 

Manihot anomala Pohl 

Manihot anomala subsp. pubescens D.J. Rogers & Appan 

Manihot brachyloba Müll. Arg. 

Manihot carthaginensis (Jacq.) Müll. Arg. 

Manihot esculenta Crantz [cultivated, naturalized] 

Manihot aff. quinquepartita Huber ex D.J. Rogers & Appan 

Manihot sprucei Pax 

Manihot surinamensis D.J. Rogers & Appan 

Manihot tristis Müll. Arg. 

Manihot tristis subsp. saxicola (Lanj.) D.J. Rogers & Appan 

Manihot tristis subsp. tristis [not in flora area] 

Maprounea amazonica Esser 

Maprounea guianensis Aubl. 

Margaritaria nobilis L. f. 

Micrandra brownsbergensis Lanj. [probably conspecific with Micrandra 

elata] 

Micrandra glabra (R.E. Schult.) R.E. Schult. 

Micrandra gleasoniana (Croizat) R.E. Schult. 

Micrandra inundata P.E. Berry & Wiedenhoeft 

Micrandra lopezii R.E. Schult. 

Micrandra minor Benth. 

Micrandra rossiana R.E. Schult. 

Micrandra siphonioides Benth. 

Micrandra sp. A 

Micrandra spruceana (Baill.) R.E. Schult. [record probably misident.] 

Micrandra sprucei (Müll. Arg.) R.E. Schult. 

Microstachys corniculata (Vahl) Griseb. 

Omphalea diandra L. 

Pausandra fordii Secco 

Pausandra martinii Baill. 

Pausandra trianae (Müll. Arg.) Baill. [specimens not seen] 

Pedilanthus tithymaloides (L.) Poit. 

Pedilanthus tithymaloides subsp. tithymaloides [native to GU, SU; cultivated 

in all] 

Pera bicolor (Klotzsch) Müll. Arg. 

Pera citriodora Baill. 

Pera decipiens (Müll. Arg.) Müll. Arg. 

Pera distichophylla (Mart.) Baill. 

Pera glabrata (Schott) Poepp. ex Baill. 


GU 

AM 

AM 

GU 

AM 

AM 

AM 

AM BO GU 

AM BO 

AM 

GU 


BO GU SU FG 

GU 

FG 

AM 

GU 

GU SU FG 

GU SU FG 

FG 

GU SU FG 



AM BO 

BO 

BO 


BO 

GU SU FG 

FG 

FG 



BO GU SU FG 

AM BO 

AM 



GU SU FG 

BO GU 

GU 

SU 

AM 

GU? 

AM BO 

AM BO GU 

AM 

AM 

BO GU FG 

AM GU? SU 

AM GU 


DA GU SU FG 

FG 


FG 

BO GU SU FG 

BO GU SU FG 


AM 

AM BO GU SU 

AM BO 

AM BO DA GU SU FG

294 


Pera tomentosa (Benth.) Müll. Arg. 

Phyllanthus acuminatus Vahl 

Phyllanthus adiantoides Klotzsch 

Phyllanthus amarus Schumach. & Thonn. 

Phyllanthus atabapoensis Jabl. 

Phyllanthus attenuatus Miq. 

Phyllanthus bolivarensis Steyerm. 

Phyllanthus borjaensis Jabl. 

Phyllanthus brasiliensis (Aubl.) Poir. 

Phyllanthus caribaeus Urb. 

Phyllanthus caroliniensis Walter 

Phyllanthus caroliniensis subsp. caroliniensis 

Phyllanthus caroliniensis subsp. guianensis (Klotzsch) G.L. Webster 

Phyllanthus caroliniensis subsp. stenopterus (Müll. Arg.) G.L. Webster 

Phyllanthus carrenoi Steyerm. 

Phyllanthus chimantae Jabl. 

Phyllanthus debilis Klein ex Willd. [naturalized] 

Phyllanthus duidae Gleason 

Phyllanthus elsiae Urb. 

Phyllanthus fluitans Benth. ex Müll. Arg. 

Phyllanthus gallinetae Jabl. [included in Phyllanthus myrsinites?] 

Phyllanthus hyssopifolioides Kunth 

Phyllanthus jablonskianus Steyerm. & Luteyn 

Phyllanthus jauaensis Jabl. 

Phyllanthus juglandifolius Willd. 

Phyllanthus juglandifolius subsp. cornifolius (Kunth) G.L. Webster 

Phyllanthus juglandifolius subsp. juglandifolius 

Phyllanthus lediformis Jabl. 

Phyllanthus lindbergii Müll. Arg. 

Phyllanthus longistylus Jabl. 

Phyllanthus maguirei Jabl. 

Phyllanthus majus Steyerm. 

Phyllanthus microphyllus Kunth 

Phyllanthus minutifolius Jabl. 

Phyllanthus minutulus Müll. Arg. 

Phyllanthus myrsinites Kunth 

Phyllanthus myrsinites subsp. francavillanus (Müll. Arg.) G.L. Webster 

Phyllanthus myrsinites subsp. myrsinites [=includes Phyllanthus gallinetae?] 

Phyllanthus neblinae Jabl. 

Phyllanthus niruri L. 

Phyllanthus obfalcatus Lasser & Maguire 

Phyllanthus orbiculatus Rich. 

Phyllanthus orinocensis Steyerm. 

Phyllanthus paezensis Jabl. 

Phyllanthus paraqueensis Jabl. 

Phyllanthus piscatorum Kunth [specimens not seen] 

Phyllanthus pseudoconami Müll. Arg. 

Phyllanthus pycnophyllus Müll. Arg. 

Phyllanthus rupestris Kunth 

Phyllanthus stipulatus (Raf.) G.L. Webster 

Phyllanthus strobilaceus Jabl. 

Phyllanthus subapicalis Jabl. 

Phyllanthus subapicalis Jabl. subsp. A 

Phyllanthus subapicalis Jabl. subsp. sequoiifolius (Jabl.) G.L. Webster 

Phyllanthus subapicalis Jabl. subsp. subapicalis 

Phyllanthus subglomeratus Poir. [cultivated only?] 

Phyllanthus tenellus Roxb. [naturalized] 

Phyllanthus tepuicola Steyerm. 

Phyllanthus urinaria L. [naturalized] 

Phyllanthus vacciniifolius (Müll. Arg.) Müll. Arg. 

Phyllanthus ventuarii Jabl. 

Piranhea longepedunculata Jabl. 

Piranhea trifoliata Baill. 

Plukenetia brachybotrya Müll. Arg. [group, vide also P. buchtienii, P. 

loretensis] 

AM 

GU SU 

GU SU 

AM 

AM 


AM BO 

BO 

GU SU FG 

AM BO 

BO GU SU FG 

BO DA GU SU 


DA FG 

BO 

BO 

BO 

DA GU SU FG 

FG 

AM 

GU SU FG 

BO DA 

DA 

GU 

GU 

SU 

AM 

AM 

BO 

BO 

SU? FG 

BO 

BO 

GU FG 

AM 

AM 

GU FG 

AM 

AM 

BO 

AM BO GU 

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Plukenetia loretensis Ule 

Plukenetia multiglandulosa Jabl. 

Plukenetia penninervia Müll. Arg. 

Plukenetia polyadenia Müll. Arg. 

Plukenetia supraglandulosa L.J. Gillespie 

Plukenetia verrucosa Sm. 

Plukenetia volubilis L. 

Podocalyx loranthoides Klotzsch 

Pogonophora schomburgkiana Miers ex Benth. 

Richeria grandis Vahl 

Ricinus communis L. [cultivated, naturalized] 

Sagotia brachysepala (Müll. Arg.) Secco 

Sagotia racemosa Baill. 

Sandwithia guyanensis Lanj. 

Sandwithia heterocalyx Secco 

Sapium argutum (Müll. Arg.) Huber 

Sapium ciliatum Hemsl. 

Sapium contortum Croizat 

Sapium glandulosum (L.) Morong 

Sapium jenmanii Hemsl. 

Sapium paucinervium Hemsl. 

Savia sessiliflora (Sw.) Willd. 

Sebastiania bidentata (Mart.) Pax 

Sebastiania guianensis (Müll. Arg.) Müll. Arg. 

Senefeldera inclinata Müll. Arg. 

Senefeldera macrophylla Ducke 

Senefelderopsis chiribiquetensis (R.E. Schult. & Croizat) Steyerm. 

Senefelderopsis croizatii Steyerm. 

Tetrorchidium rubrivenium Poepp. 

Tragia fallax Müll. Arg. 

Tragia fendleri Müll. Arg. 

Tragia guayanensis L.J. Gillespie 

Tragia lessertiana (Baill.) Müll. Arg. 

Tragia tabulaemontana L.J. Gillespie 

Tragia volubilis L. 


Acalypha cuneata var. obovata (Benth.) Müll. Arg. =Acalypha cuneata 

Acalypha diversifolia var. carpinifolia (Müll. Arg.) Müll. Arg. =Acalypha diversifolia 

Acalypha diversifolia var. leptostachya (Kunth) Müll. Arg. =Acalypha diversifolia 

Acalypha hirsutissima Willd. =Acalypha macrostachya 

Acalypha karsteniana Pax & K. Hoffm. =Acalypha villosa 

Acalypha leptostachya Kunth =Acalypha diversifolia 

Acalypha leptostachya var. carpinifolia Müll. Arg. =Acalypha diversifolia 

Acalypha macrophylla Kunth =Acalypha macrostachya 

Acalypha macrostachya var. hirsutissima (Willd.) Müll. Arg. =Acalypha macrostachya 

Acalypha macrostachya var. macrophylla (Kunth) Müll. Arg. =Acalypha macrostachya 

Acalypha macrostachya var. tristis (Poepp.) Müll. Arg. =Acalypha macrostachya 

Acalypha obovata Benth. =Acalypha cuneata 

Acalypha persimilis var. corchorifolia (A. Rich.) Müll. Arg. [not in flora area] 

Acalypha samydifolia Poepp. =Acalypha diversifolia 

Acalypha schiedeana var. angustifolia (Müll. Arg.) Pax & K. Hoffm. =Acalypha schiedeana 

Acalypha schiedeana f. angustifolia Müll. Arg. =Acalypha schiedeana 

Acalypha tristis Poepp. =Acalypha macrostachya 

Acalypha villosa var. intermedia Müll. Arg. =Acalypha villosa 

Acalypha villosa var. paniculata Müll. Arg. =Acalypha villosa 

Acalypha villosa var. tomentosa Müll. Arg. =Acalypha villosa 

Acidoton venezolanus (Pax & K. Hoffm.) G.L. Webster =Acidoton nicaraguensis 

Actinostemon depauperatus Pax & K. Hoffm. =Actinostemon schomburgkii 

Actinostemon guyanensis Pax & K. Hoffm. ? =Sandwithia guyanensis 

Actinostemon parvifolius Pittier =Actinostemon schomburgkii 

Alchornea brachygyne Pax & K. Hoffm. =Alchornea discolor 

Alchornea cordata (A. Juss.) Müll. Arg. =Aparisthmium cordatum 

Alchornea glandulosa var. floribunda Benth. =Alchorneopsis floribunda 

Alchornea orinocensis Croizat =Aparisthmium cordatum 

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296 


Alchornea schomburgkii Klotzsch =Alchornea discolor 

Alchorneopsis trimera Lanj. =Alchorneopsis floribunda 

Alectoroctonum caracasanum Klotzsch & Garcke =Euphorbia cotinifolia 

Amanoa grandiflora (Müll. Arg.) Müll. Arg. =Amanoa guianensis 

Amanoa guianensis var. grandiflora Müll. Arg. =Amanoa guianensis 

Amanoa pubescens Steyerm. =Amanoa almerindae 

Angostyles tabulamontana Croizat [dubious species of uncertain placement] 

Antidesma triplinervia Spreng. =Alchornea triplinervia 

Apodandra brachybotrya (Müll. Arg.) J.F. Macbr. =Plukenetia brachybotrya 

Apodandra loretensis (Ule) Pax & K. Hoffm. =Plukenetia loretensis 

Argythamnia polygama (Jacq.) Kuntze =Ditaxis polygama 

Astrococcus coriaceus Baill. =Haematostemon coriaceus 

Ateramnus hypoleucus (Benth.) Rothm. =Gymnanthes hypoleuca 

Chaetocarpus stipularis Gleason =Chaetocarpus schomburgkianus 

Chaetocarpus williamsii Steyerm. =Chaetocarpus schomburgkianus 

Chamaesyce adenoptera (Bertol.) Small =Euphorbia dioeca 

Chamaesyce dioeca (Kunth) Millsp. =Euphorbia dioeca 

Chamaesyce glomerifera Millsp. =Euphorbia hypericifolia 

Chamaesyce hirta (L.) Millsp. =Euphorbia hirta 

Chamaesyce hypericifolia (L.) Millsp. =Euphorbia hypericifolia 

Chamaesyce hyssopifolia (L.) Small = Euphorbia hyssopifolia (L.) 

Chamaesyce mesembrianthemifolia (Jacq.) Dugand =Euphorbia mesembrianthemifolia 

Chamaesyce prostrata (Aiton) Small =Euphorbia prostrata 

Chamaesyce serpens (Kunth) Small =Euphorbia serpens 

Chamaesyce thymifolia (L.) Millsp. =Euphorbia thymifolia 

Cleidion sp. [not in flora area] 

Cleidion? nicaraguense Hemsl. =Acidoton nicaraguensis 

Clusiophyllum sprucei Müll. Arg. =Micrandra sprucei 

Conami brasiliensis Aubl. =Phyllanthus brasiliensis 

Conceveiba cordatum A. Juss. =Aparisthmium cordatum 

Conceveiba hostmannii Benth. =Conceveiba guianensis 

Conceveiba terminalis (Baill.) Müll. Arg. =Gavarretia terminalis 

Conceveibastrum martianum (Baill.) Pax & K. Hoffm. =Conceveiba martiana 

Conceveibastrum ptarianum Steyerm. =Conceveiba ptariana 

Conceveibum cordatum A. Juss. =Aparisthmium cordatum 

Croton barlettii Lanj. =Croton potaroensis 

Croton bartlettii Lanj. =Croton potaroensis 

Croton benthamianus Müll. Arg. =Croton palanostigma 

Croton cardonei Croizat =Croton potaroensis 

Croton castaneifolius L. =Caperonia castaneifolia 

Croton crenatus Müll. Arg. =Croton mollis 

Croton glabellus L. [not in flora area] 

Croton glandulosus L. [not in flora area] 

Croton glandulosus subsp. hirtus (L’Hér.) Croizat =Croton hirtus 

Croton glandulosus var. hirtus (L’Hér.) Müll. Arg. =Croton hirtus 

Croton glandulosus subsp. hirtus (L’Hér.) Croizat, Bull. =Croton hirtus 

Croton kavanayensis Steyerm. =Croton matourensis 

Croton lanjouwensis Jabl. =Croton matourensis 

Croton matourensis var. benthamianus Müll. Arg. =Croton matourensis 

Croton micans Sw. =Croton argyrophyllus 

Croton miquelensis Ferguson ex Lanj. =Croton trinitatis 

Croton miquelianus Ferguson =Croton trinitatis 

Croton monachinoensis Jabl. =Croton cuneatus 

Croton nervosus Klotzsch [nom. illeg.] =Croton argyrophyllus 

Croton pakaraimae Jabl. =Croton matourensis 

Croton polygamus Jacq. =Ditaxis polygama 

Croton populifolius Mill. [see Croton essequiboensis] 

Croton populifolius var. essequiboensis (Klotzsch) Müll. Arg. =Croton essequiboensis 

Croton roraimensis var. subinteger Steyerm. =Croton roraimensis 

Croton sessiliflorum Sw. =Savia sessiliflora 

Croton surinamensis Müll. Arg. =Croton cuneatus 

Croton tafelbergicus Croizat =Croton guianensis 

Croton xanthochloros Croizat =Croton megalodendron 

Cunuria ? casiquiarensis Croizat =Podocalyx loranthoides


Cunuria crassipes Müll. Arg. =Micrandra sprucei 

Cunuria glabra R.E. Schult. ex Baldwin & R.E. Schult. =Micrandra glabra 

Cunuria gleasoniana Croizat =Micrandra gleasoniana 

Cunuria spruceana Baill. =Micrandra spruceana 

Dactylostemon schomburgkii Klotzsch =Actinostemon schomburgkii 

Dalechampia brevipes Briq. =Dalechampia scandens 

Dalechampia colorata L. f. =Dalechampia scandens 

Dalechampia cynanchoides S. Moore =Dalechampia tenuiramea 

Dalechampia dioscoreifolia Pulle [latter homonym] =Dalechampia affinis 

Dalechampia heterophylla Vahl =Dalechampia tiliifolia 

Dalechampia latifolia Lam. =Dalechampia scandens 

Dalechampia mollis Vahl =Dalechampia scandens 

Dalechampia peruviana Lam. =Dalechampia tiliifolia 

Dalechampia pruriens Griseb. =Dalechampia tiliifolia 

Dalechampia roezliana var. amazonica Ule =Dalechampia magnoliifolia 

Dalechampia rubiformis Spreng. =Dalechampia scandens 

Dalechampia ruboides Kunth =Dalechampia scandens 

Dalechampia sidaefolia Kunth =Dalechampia scandens 

Dalechampia spathulata (Scheidw.) Baill. [not in flora area] 

Dalechampia villosa Lam. =Dalechampia scandens 

Discocarpus brasiliensis Klotzsch ex Müll. Arg. =Discocarpus essequeboensis 

Discocarpus mazarunensis Croizat =Chaetocarpus schomburgkianus 

Dodecastigma mazarunense Croizat =Dodecastigma integrifolium 

Drypetes maguireana Monach. =Chaetocarpus schomburgkianus 

Drypetes spruceana Müll. Arg. =Chaetocarpus schomburgkianus 

Elaeophora abutifolia Ducke =Plukenetia polyadenia 

Elaeophora polyadenia (Müll. Arg.) Ducke =Plukenetia polyadenia 

Euphorbia adenoptera Bertol. =Euphorbia dioeca 

Euphorbia brasiliensis Lam. =Euphorbia hyssopifolia 

Euphorbia buxifolia Lam. =Euphorbia mesembrianthemifolia 

Euphorbia caracasana (Klotzsch & Garcke) Boiss. =Euphorbia cotinifolia 

Euphorbia cotinoides Miq. =Euphorbia cotinifolia subsp. cotinoides 

Euphorbia cyathophora Murray =Euphorbia heterophylla var. cyathophora 

Euphorbia globulifera Kunth =Euphorbia hirta 

Euphorbia morisoniana Klotzsch =Euphorbia heterophylla var. heterophylla 

Euphorbia multinodis Urb. [not in flora area] 

Euphorbia myrtillifolia L. [not in flora area] 

Euphorbia surinamensis Lanj. =Euphorbia oerstediana 

Euphorbia tenella Kunth =Euphorbia serpens 

Euphorbia tithymaloides L. =Pedilanthus tithymaloides 

Gaedawakka schomburgkiana Kuntze =Chaetocarpus schomburgkianus 

Gaedawakka schomburgkianus Kuntze =Chaetocarpus schomburgkianus 

Gitara venezolana Pax & K. Hoffm. =Acidoton nicaraguensis 

Glochidion vacciniifolium Müll. Arg. =Phyllanthus vacciniifolius 

Guarania laurifolia Baill. =Richeria grandis 

Gymnanthes guyanensis Müll. Arg. =Sebastiania guianensis 

Hermesia castaneifolia Willd. =Alchornea castaneifolia 

Hevea confusa Hemsl. =Hevea pauciflora var. coriacea 

Hevea lutea (Spruce ex Benth.) Müll. Arg. =Hevea guianensis var. lutea 

Hevea minor Hemsl. =Hevea pauciflora var. coriacea 

Hevea paludosa Ule =Hevea pauciflora var. coriacea 

Hieronyma laxiflora (Tul.) Müll. Arg. =Hieronyma alchorneoides var. alchorneoides 

Hippomane biglandulosa L. [nom. illeg.] =Sapium glandulosum 

Hippomane glandulosa L. =Sapium glandulosum 

Jatropha carthaginensis Jacq. =Manihot carthaginensis 

Jatropha manihot L. =Manihot esculenta 

Jatropha urens L. =Cnidoscolus urens 

Julocroton argenteus (L.) Didr. =Croton argenteus 

Julocroton vergarenae Jabl. =Croton vergarenae 

Mabea argutissima Croizat =Mabea subsessilis 

Mabea atroviridis Pax & K. Hoffm. =Mabea montana 

Mabea caudata Pax & K. Hoffm. =Mabea speciosa subsp. speciosa 

Mabea caudata var. concolor Lanj. =Mabea speciosa subsp. A 

Mabea ciliata Gleason =Mabea montana

298 


Mabea costata Pax & K. Hoffm. =Mabea montana 

Mabea eximia Ducke =Mabea pulcherrima 

Mabea floribunda Jabl. =Mabea montana 

Mabea longepedicellata Pittier =Mabea montana 

Mabea lucida Pax & K. Hoffm. =Mabea montana 

Mabea maynensis Spruce ex Müll. Arg. =Mabea piriri 

Mabea montana subsp. biglandulosa (Müll. Arg.) Hollander =Mabea biglandulosa 

Mabea montana subsp. lucida (Pax & K. Hoffm.) Hollander =Mabea montana 

Mabea muricata Jabl. =Mabea nitida 

Mabea nitida var. albiflora Müll. Arg. =Mabea nitida 

Mabea occidentalis var. laevigata (Müll. Arg.) Müll. Arg. =Mabea biglandulosa 

Mabea occidentalis var. obovata (Müll. Arg.) Müll. Arg. =Mabea speciosa subsp. B 

Mabea occidentalis var. purpurascens (Müll. Arg.) Müll. Arg. =Mabea speciosa subsp. B 

Mabea orbiculata Jabl. =Mabea frutescens 

Mabea pallida Müll. Arg. =Mabea occidentalis 

Mabea parguazae Jabl. =Mabea montana 

Mabea parvifolia Pax & K. Hoffm. =Mabea trianae 

Mabea piriri var. laevigata Müll. Arg. =Mabea biglandulosa 

Mabea piriri var. obovata Müll. Arg. =Mabea speciosa subsp. B 

Mabea piriri var. purpurascens Müll. Arg. =Mabea speciosa subsp. B 

Mabea saramaccensis Croizat =Mabea speciosa subsp. A 

Mabea schomburgkii Benth. =Mabea taquari 

Mabea speciosa subsp. concolor (Müll. Arg.) Hollander =Mabea piriri 

Mabea taquari var. angustifolia Müll. Arg. =Mabea taquari 

Mabea verrucosa Pax & K. Hoffm. =Mabea montana 

Manihot diffusa Pohl =Manihot esculenta 

Manihot dulcis Pax =Manihot esculenta 

Manihot glaziovii Müll. Arg. [not in flora area] 

Manihot maguireana D.J. Rogers & Appan =Manihot tristis subsp. saxicola 

Manihot manihot (L.) Cockerell =Manihot esculenta 

Manihot melanobasis Müll. Arg. [not in flora area] 

Manihot orinocensis Croizat =Manihot tristis subsp. tristis 

Manihot rusbyi Britton =Manihot brachyloba 

Manihot saxicola Lanj. =Manihot tristis subsp. saxicola 

Manihot utilissima Pohl =Manihot esculenta 

Maprounea guianensis var. guianensis f. obtusata Müll. Arg. =Maprounea amazonica 

Maprounea guianensis var. obtusata (Müll. Arg.) Müll. Arg. =Maprounea amazonica 

Meborea guianensis Aubl. [name of uncertain application] =Phyllanthus sp. 

Megalostylis poeppigii S. Moore =Dalechampia micrantha 

Micrandra cunuri Baill. ex Müll. Arg. =Micrandra spruceana 

Micrandra elata (Didr.) Müll. Arg. [see Micrandra brownsbergensis] 

Micrandra siphonioides var. minor (Benth.) Müll. Arg. =Micrandra minor 

Microstachys guianensis Klotzsch =Microstachys corniculata 

Moeroris stipulata Raf. =Phyllanthus stipulatus 

Omphalea elaeophoroides Steyerm. ex Standl. =Omphalea diandra 

Omphalea megacarpa Hemsl. =Omphalea diandra 

Pausandra flagellorachis Lanj. =Pausandra martinii 

Pausandra flagellorhachis Lanj. =Pausandra martinii 

Pausandra integrifolia Lanj. =Dodecastigma integrifolium 

Pera echinocarpa Baill. [not in flora area] 

Pera ferruginea (Schott) Müll. Arg. =Pera glabrata 

Pera nitida (Benth.) Jabl. =Pera decipiens 

Pera obovata Baill. [not in flora area] 

Pera schomburgkiana (Klotzsch) Müll. Arg. =Pera bicolor 

Peridium bicolor Klotzsch =Pera bicolor 

Peridium bicolor var. nitidum Benth. =Pera decipiens 

Peridium bicolor var. tomentosum Benth. =Pera tomentosa 

Peridium decipiens Müll. Arg. =Pera decipiens 

Peridium ferrugineum Schott =Pera glabrata 

Peridium glabratum Schott =Pera glabrata 

Peridium schomburgkianum Klotzsch =Pera bicolor 

Phyllanthus adenophyllus Müll. Arg. =Phyllanthus myrsinites 

Phyllanthus anadenus Jabl. =Phyllanthus subapicalis subsp. subapicalis


Phyllanthus brachycladus Müll. Arg. =Phyllanthus rupestris 

Phyllanthus conami Sw. =Phyllanthus brasiliensis 

Phyllanthus congestus Benth. ex Müll. Arg. =Jablonskia congesta 

Phyllanthus corcovadensis Müll. Arg. =Phyllanthus tenellus 

Phyllanthus cornifolius Kunth =Phyllanthus juglandifolius subsp. cornifolius 

Phyllanthus croizatii Steyerm. =Phyllanthus urinaria 

Phyllanthus delicatissimus Jabl. =Phyllanthus rupestris 

Phyllanthus diffusus Klotzsch =Phyllanthus stipulatus 

Phyllanthus dinizii Huber =Phyllanthus myrsinites 

Phyllanthus epiphyllanthus L. [not in flora area] 

Phyllanthus francavillanus Müll. Arg. =Phyllanthus myrsinites subsp. francavillanus 

Phyllanthus glaucoviridis Jabl. =Phyllanthus myrsinites 

Phyllanthus grandifolius L. [not in flora area] 

Phyllanthus guianensis Klotzsch =Phyllanthus caroliniensis subsp. guianensis 

Phyllanthus guyanensis Müll. Arg. =Phyllanthus attenuatus 

Phyllanthus ichthyomethius Rusby =Phyllanthus piscatorum 

Phyllanthus lathyroides Kunth [not in flora area] 

Phyllanthus microcarpoides Croizat =Phyllanthus rupestris 

Phyllanthus monocladus Urb. =Phyllanthus hyssopifolioides 

Phyllanthus nobilis (L. f.) Müll. Arg. =Margaritaria nobilis 

Phyllanthus pentrandrus Schumach. & Thonn. [not in flora area] 

Phyllanthus pimichinianus Jabl. =Phyllanthus myrsinites subsp. francavillanus 

Phyllanthus pseudoconami Müll. Arg. =Phyllanthus piscatorum 

Phyllanthus pseudoconami var. pubescens Müll. Arg. =Phyllanthus piscatorum 

Phyllanthus schomburgkianus Müll. Arg. =Phyllanthus caroliniensis subsp. guianensis 

Phyllanthus sequoiifolius Jabl. =Phyllanthus subapicalis subsp. sequoiifolius 

Phyllanthus vacciniifolius subsp. vinillaensis Steyerm. =Phyllanthus vacciniifolius 

Plukenetia abutifolia (Ducke) Pax & K. Hoffm. =Plukenetia polyadenia 

Plukenetia buchtienii Pax =Plukenetia brachybotrya 

Plukenetia macrostyla Ule =Plukenetia volubilis 

Pogonophora schomburgkiana f. elliptica Pax & K. Hoffm. =Pogonophora schomburgkiana 

Pogonophora schomburgkiana var. longifolia Miers ex Benth. =Pogonophora schomburgkiana 

Poinsettia heterophylla (L.) Klotzsch & Garcke =Euphorbia heterophylla var. heterophylla 

Poinsettia oerstediana Klotzsch & Garcke =Euphorbia oerstediana 

Poraresia anomala Gleason =Pogonophora schomburgkiana 

Pseudosenefeldera inclinata (Müll. Arg.) Esser =Senefeldera inclinata 

Rhodothyrsus macrophyllus (Ducke) Esser =Senefeldera macrophylla 

Rhopalostylis buettnerioides Klotzsch ex Baill. =Dalechampia micrantha 

Richeria grandis var. divaricata Müll. Arg. =Richeria grandis 

Richeria grandis var. genuina Müll. Arg. =Richeria grandis 

Richeria grandis var. obovata Müll. Arg. =Richeria grandis 

Richeria grandis var. racemosa Müll. Arg. =Richeria grandis 

Richeria laurifolia (Baill.) Baill. =Richeria grandis 

Richeria loranthoides (Klotzsch) Müll. Arg. =Podocalyx loranthoides 

Richeria obovata (Müll. Arg.) Pax & K. Hoffm. =Richeria grandis 

Richeria racemosa (Müll. Arg.) Pax & K. Hoffm. =Richeria grandis 

Ricinella triloba Müll. Arg. =Adelia triloba 

Sagotia racemosa var. brachysepala Müll. Arg. =Sagotia brachysepala 

Sagotia racemosa var. genuina Müll. Arg. =Sagotia racemosa 

Sagotia racemosa var. ligularis Müll. Arg. =Sagotia racemosa 

Sagotia racemosa var. microsepala Müll. Arg. =Sagotia racemosa 

Sagotia tafelbergii Croizat =Sagotia racemosa 

Sapium albomarginatum Pax & K. Hoffm. =Sapium glandulosum 

Sapium aubletianum (Müll. Arg.) Huber [nom. illeg.] =Sapium glandulosum 

Sapium aucuparium Jacq. [nom. illeg.] [see Sapium glandulosum] =Sapium glandulosum 

Sapium biglandulosum (L.) Müll. Arg. [nom. illeg.] [see Sapium glandulosum] =Sapium glandulosum 

Sapium biglandulosum var. aubletianum Müll. Arg. [nom. illeg.] =Sapium glandulosum 

Sapium biglandulosum var. klotzschianum Müll. Arg. =Sapium glandulosum 

Sapium biglandulosum var. lanceolatum Müll. Arg. =Sapium glandulosum 

Sapium cladogyne Hutch. =Sapium jenmanii 

Sapium ellipticum Hemsl. [see Sapium glandulosum] 

Sapium guaricense Pittier =Sapium glandulosum 

Sapium hemsleyanum Huber =Sapium glandulosum

300 


Sapium hippomane G. Mey. [nom. illeg.] =Sapium glandulosum 

Sapium klotzschianum (Müll. Arg.) Huber =Sapium glandulosum 

Sapium lanceolatum (Müll. Arg.) Huber =Sapium glandulosum 

Sapium microdentatum Lanj. =Sapium paucinervium 

Sapium montanum Lanj. =Sapium argutum 

Sapium moritzianum Klotzsch =Sapium glandulosum 

Sapium naiguatense Pittier =Sapium glandulosum 

Sapium obtusilobum Müll. Arg. =Sapium glandulosum 

Sapium paucistamineum Pittier =Sapium glandulosum 

Sapium prunifolium Klotzsch =Sapium glandulosum 

Sapium yutajense Jabl. =Dendrothrix yutajensis 

Sebastiania corniculata (Vahl) Müll. Arg. =Microstachys corniculata 

Sebastiania corniculata var. guyanensis (Klotzsch) Pax & K. Hoffm. =Microstachys corniculata 

Sebastiania linearifolia Lanj. =Sebastiania bidentata 

Sebastiania micrantha (Benth.) Lanj. =Microstachys corniculata 

Securinega congesta (Benth. ex Müll. Arg.) Müll. Arg. =Jablonskia congesta 

Senefeldera chiribiquetensis R. Schult. & Croizat =Senefelderopsis chiribiquetensis 

Senefeldera karsteniana Pax & K. Hoffm. =Senefeldera inclinata 

Senefeldera multiflora var. acutifolia Müll. Arg. =Senefeldera inclinata 

Senefeldera yutajensis (Jabl.) G.L. Webster =Dendrothrix yutajensis 

Senefelderopsis sipapoensis Jabl. =Senefelderopsis croizatii 

Senefelderopsis venamoensis Jabl. =Senefelderopsis croizatii 

Siphonia lutea Spruce ex Benth. =Hevea guianensis var. lutea 

Siphonia pauciflora Spruce ex Benth. =Hevea pauciflora 

Spixia distichophylla Mart. =Pera distichophylla 

Stilaginella laxiflora Tul. =Hieronyma alchorneoides var. alchorneoides 

Stilaginella oblonga Tul. =Hieronyma oblonga 

Traganthus sidoides Klotzsch =Bernardia sidoides 

Tragia corniculata Vahl =Microstachys corniculata 

Tragia grandifolia Klotzsch =Adenophaedra grandifolia 

Tragia japurensis Müll. Arg. =Tragia fendleri 

Tragiopsis fruticulosa H. Karst =Microstachys corniculata 

Veconcibea latifolia (Benth.) Pax & K. Hoffm. =Conceveiba latifolia 

REFERENCES 

Burger, W. & M. Huft. Euphorbiaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 36: 1-169. 

Den Hollander, G. & C.C. Berg. 1986. Studies on the flora of the Guianas 21. Mabea species of the Guianas - a precursor. Proc. 

Kon. Ned. Akad. Wet. ser. C, 89(2): 147-157. 

Franco, P. 1990. The genus Hyeronima in South America. Bot. Jahrb. Syst. 111: 297-346. 

Gillespie, L.J. 1993. Euphorbiaceae of the Guianas: Annotated species checklist and key to the genera. Brittonia 45(1): 56-94. 

Gillespie, L.J. 1993. A synopsis of the Neotropical Plukenetia including two new species. Syst. Bot. 18(4): 575-592. 

Gillespie, L.J. 1994. A new section and two new species of Tragia from Venezuelan Guayana and French Guiana. Novon 4(4): 

330-338. 

Gillespie, L.J. 2002. Euphorbiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


266-298. 

Gillespie, L.J. & W.S. Armbruster. 1997. A contribution to the Guianan Flora: Dalechampia, Haematostemon, Omphalea, Pera, 

Plukenetia, and Tragia, with notes on subfamily Acalyphoideae. Smithsonian Contributions to Botany 86. 

Görts-van Rijn, A.R.A. 1976. Euphorbiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 

387-424. 

Hayden, W.J. 1990. Notes on Neotropical Amanoa. Brittonia 42(4): 260-270. 

Jablonski, E. 1965. Euphorbiaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part 6. Mem. New York 

Bot. Gard. 12(3): 1-285. 

Jablonski, E. 1967. Euphorbiaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part 7. Mem. New York 

Bot. Gard. 17(1): 1-439. 

Lanjouw, J. 1932. Euphorbiaceae. In A. Pulle, Flora of Suriname 2(1): 1-101. 

Lanjouw, J. 1939. Euphorbiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 457-470. 

Rogers, D.J. & S.G. Appan. 1973. Manihot and Manihotoides. Flora Neotropica Monogr. 13: 1-272. 

Webster, G.L. & W.S. Armbruster. 1991. A synopsis of the neotropical species of Dalechampia. J. Linn. Soc. Bot. 105: 137-177. 

Webster, G.L., P.E. Berry, W.S. Armbruster, H.-J. Esser, L.J. Gillespie, W.J. Hayden, G.A. Levin, R. de S. Secco, & S.V. Heald. 

1999. Euphorbiaceae.In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 5: 72- 

228.


EUPHRONIACEAE 

L. Marcano-Berti, J.A. Steyermark† 

Euphronia acuminatissima Steyerm. 

Euphronia guianensis (R.H. Schomb.) Hallier f. [=Euphronia hirtelloides s.l.] 

Euphronia hirtelloides Mart. ex Mart. & Zucc. 


Euphronia licanioides (Spruce ex Warm.) Hallier f. =Euphronia hirtelloides 

Lightia guianensis R.H. Schomb. =Euphronia guianensis 

Lightia licanioides Spruce ex Warm. =Euphronia hirtelloides 

REFERENCES 

Lleras, E. 1976. Revision and taxonomic position of the genus Euphronia Martius ex Martius & Zuccarini (Vochysiaceae). Acta 

Amaz. 6: 43-47. 

Marcano-Berti, L. 1989. Euphroniaceae: una nueva familia. Pittieria 18: 15-17. 

Marcano-Berti, L. 1998. Euphroniaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 21: 45-48. 

Steyermark, J.A. & L. Marcano-Berti. 1999. Euphroniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora 

of the Venezuelan Guayana 5: 228-230. 

FLACOURTIACEAE 

G.A. Aymard C., P.E. Berry, R.W. Kiger, M. Olson 

Banara guianensis Aubl. 

Banara nitida Spruce ex Benth. 

Banara orinocensis (Cuatrec.) Sleumer 

Carpotroche crispidentata Ducke 

Carpotroche grandiflora Spruce ex Benth. 

Carpotroche longifolia (Poepp.) Benth. 

Carpotroche surinamensis Uittien 

Casearia aculeata Jacq. 

Casearia acuminata DC. 

Casearia arborea (Rich.) Urb. 

Casearia celastroides Klotzsch [nomen nudum] 

Casearia combaymensis Tul. [Guianan material often det. C. bracteifera or C. 

singularis] 

Casearia commersoniana Cambess. 

Casearia decandra Jacq. 

Casearia duckeana Sleumer 

Casearia fasciculata (Ruiz & Pav.) Sleumer 

Casearia grandiflora Cambess. 

Casearia guianensis (Aubl.) Urb. 

Casearia hirsuta Sw. 

Casearia javitensis Kunth 

Casearia mariquitensis Kunth 

Casearia mollis Kunth 

Casearia neblinae Sleumer 

Casearia negrensis Eichler 

Casearia pitumba Sleumer 

Casearia prunifolia Kunth 

Casearia resinifera Spruce ex Eichler 

Casearia rusbyana Briq. 

Casearia spinescens (Sw.) Griseb. 

Casearia spruceana Benth. ex Eichler 

Casearia sylvestris Sw. 

Casearia tremula (Griseb.) Griseb. ex Wright 

Casearia ulmifolia Vahl ex Vent. 

Casearia zizyphoides Kunth 

Euceraea nitida Mart. 

Euceraea rheophytica P.E. Berry & M. Olson 

Euceraea sleumeriana Steyerm. & Maguire 

Hasseltia floribunda Kunth 

Hecatostemon completus (Jacq.) Sleumer 

Homalium guianense (Aubl.) Oken 

AM BO 

BO GU 

AM 


BO DA 

AM BO DA 

AM FG 

AM SU 

GU SU FG 

GU SU FG 


GU? SU FG 


GU 




GU 

GU? 



AM BO SU 



BO 

AM 

SU FG 


AM FG 

FG 


BO DA GU 

AM 


BO GU 



AM BO GU SU 

AM 

BO 

FG? 

BO GU 

BO DA GU SU FG

302 

Homalium racemosum Jacq. 

Laetia americana L. 

Laetia coriacea Spruce ex Benth. 

Laetia cupulata Spruce ex Benth. 

Laetia procera (Poepp.) Eichler 

Laetia suaveolens (Poepp.) Benth. 

Lindackeria latifolia Benth. 

Lindackeria paludosa (Benth.) Gilg 

Lunania parviflora Spruce ex Benth. 

Mayna grandifolia (H. Karst.) Warb. 

Mayna odorata Aubl. 

Neoptychocarpus apodanthus (Kuhlm.) Buchheim 

Neoptychocarpus killipii (Monach.) Buchheim 

Neosprucea paterna Alford, sp. nov. ined. 

Prockia crucis P. Browne ex L. [expected in the Guianas] 

Ryania angustifolia (Turcz.) Monach. 

Ryania dentata (Kunth) Miq. 

Ryania dentata var. dentata 

Ryania pyrifera (Rich.) Uittien & Sleumer 

Ryania speciosa Vahl 

Ryania spruceana Monach. 

Xylosma benthamii (Tul.) Triana & Planch. 

Xylosma ciliatifolium (Clos) Eichler 

Xylosma tessmannii Sleumer 

Zuelania guidonia (Sw.) Britton & Millsp. 



Athenaea guianensis (Aubl.) J.F. Gmel. =Casearia guianensis 

Azara umbellata C. Presl =Laetia americana 

Banara glandulosa (Desv. ex Ham.) Speg. =Banara guianensis 

Banara guianensis auct. non Aubl. 1775 =Banara nitida 

Banara guianensis var. mollis (Poepp.) Eichler =Banara guianensis 

Banara mollis (Poepp.) Tul. =Banara guianensis 

Banara sacupanensis Aristeg. =Banara orinocensis 

Carpotroche amazonica Mart. =Carpotroche grandiflora 

Carpotroche amazonica auct. non Mart. 1871 =Mayna grandifolia (p.p.) 

Carpotroche laxiflora (Benth.) Walp. =Lindackeria paludosa 

Carpotroche paludosa (Benth.) Walp. =Lindackeria paludosa 

Carpotroche zuliana Pittier =Mayna grandifolia 

Casearia aculeata auct. non Jacq. 1760 =Casearia spinescens 

Casearia arguta auct. non Kunth 1821 [1823], sensu Pittier et al. =Casearia mollis 

Casearia arguta Kunth [not in flora area] 

Casearia arguta auct. non Kunth 1821 [1823], sensu Steyerm. =Casearia guianensis 

Casearia bracteifera Sagot =Casearia combaymensis 

Casearia cambessedesii Eichler =Casearia ulmifolia 

Casearia carpinifolia Benth. =Casearia sylvestris 

Casearia celtidifolia Poepp. ex Eichler non Kunth 1823 =Casearia ulmifolia 

Casearia celtidifolia Kunth =Casearia ulmifolia 

Casearia cotticensis Uittien =Casearia ulmifolia 

Casearia densiflora Benth. =Casearia commersoniana 

Casearia dentata (Aubl.) Sessé & Moç. =Casearia commersoniana 

Casearia guidonia (Sw.) Lundell non Benth. =Zuelania guidonia 

Casearia inaequilatera auct. non Cambess. 1829 [1830] =Casearia sylvestris 

Casearia laevigata Willd. ex Klotzsch [nomen nudum] =Casearia zizyphoides 

Casearia lasiophylla auct. non Eichler 1871 =Casearia mollis 

Casearia laurifolia Benth. =Casearia commersoniana 

Casearia lingua Cambess. =Casearia sylvestris 

Casearia macrophylla Vahl =Casearia pitumba 

Casearia mollis auct. non Kunth 1821 [1823] =Casearia hirsuta 

Casearia negrensis auct. non Eichler 1871 =Laetia suaveolens 

Casearia nitida Jacq. [not in flora area] 

Casearia petraea Benth. =Casearia ulmifolia 

Casearia singularis Eichler =Casearia combaymensis 

Casearia spinescens auct. non (Sw.) Griesb. 1863, sensu Pittier et al. =Casearia guianensis 


BO 

AM BO 

AM GU 


AM GU 

AM GU SU 

AM BO GU FG? 

AM GU 

AM 

GU SU FG 

SU FG 

FG 

GU FG? 

AM 

AM 

AM 

BO 

GU SU FG 

AM 

AM 

BO GU SU FG 

BO GU SU FG 

GU FG? 

GU FG 

AM BO


Casearia sylvestris var. carpinifolia (Benth.) Briq. =Casearia sylvestris 

Casearia sylvestris var. lingua (Cambess.) Eichler =Casearia sylvestris 

Casearia sylvestris var. sylvestris =Casearia sylvestris 

Casinga procera (Poepp.) Griseb. =Laetia procera 

Casinga suaveolens (Poepp.) Griseb. ex Benth. =Laetia suaveolens 

Chaetocrater javitensis (Kunth) Raf. =Casearia javitensis 

Dendrostylis grandifolia H. Karst. =Mayna grandifolia 

Flacourtia benthamii Tul. =Xylosma benthamii 

Flacourtia nitida Benth. =Xylosma benthamii 

Guidonia apetala (Jacq.) Kuntze =Laetia americana 

Guidonia calophylla (Eichler) Kuntze =Laetia suaveolens 

Guidonia coriacea (Spruce ex Benth.) Kuntze =Laetia coriacea 

Guidonia cupulata (Spruce ex Benth.) Kuntze =Laetia cupulata 

Guidonia hirsuta (Sw.) M. Gómez =Casearia hirsuta 

Guidonia procera (Poepp.) Kuntze =Laetia procera 

Guidonia spinescens (Sw.) Griseb. =Casearia spinescens 

Guidonia suaveolens (Poepp.) Kuntze =Laetia suaveolens 

Guidonia tremula (Griseb.) Maza =Casearia tremula 

Guidonia ulmifolia (Vahl ex Vent.) Baill. =Casearia ulmifolia 

Hecatostemon dasygynus S.F. Blake =Hecatostemon completus 

Hecatostemon guazumifolius (Kunth) Sleumer =Hecatostemon completus 

Hisingera benthamii (Tul.) Clos =Xylosma benthamii 

Homalium anzoateguiense Steyerm. =Homalium racemosum 

Homalium densiflorum Spruce ex Benth. =Homalium guianense 

Homalium eleuthrostylum S.F. Blake =Homalium racemosum 

Homalium eurypetalum S.F. Blake =Homalium racemosum 

Homalium mituense Cuatrec. =Homalium racemosum 

Homalium napimoga Spreng. =Homalium guianense 

Homalium pedicellatum Spruce ex Benth. =Homalium racemosum 

Homalium pittieri S.F. Blake =Homalium racemosum 

Homalium puberulum Klotzsch =Homalium guianense 

Homalium racoubea Sw. =Homalium guianense 

Iroucana guianensis Aubl. =Casearia guianensis 

Kuhlia mollis Poepp. =Banara guianensis 

Laetia acuminata Bonpl. ex Triana & Planch. =Laetia americana 

Laetia apetala Jacq. =Laetia americana 

Laetia calophylla Eichler =Laetia suaveolens 

Laetia casearioides Sagot ex Benth. =Laetia procera 

Laetia completa Jacq. =Hecatostemon completus 

Laetia guazumaefolia Kunth =Hecatostemon completus 

Laetia guidonia Sw. =Zuelania guidonia 

Laetia hirtella Kunth =Hecatostemon completus 

Laetia obtusifolia Eichler ex Benoist [nomen nudum] =Laetia procera 

Lindackeria latifolia auct. non Benth. 1851 =Lindackeria paludosa 

Lindackeria maynensis var. laxiflora (Benth.) Mart. ex Pittier et al. =Lindackeria paludosa 

Mayna denticulata Benth. =Mayna odorata 

Mayna grandiflora (Spruce ex Benth.) R.E. Schult. =Carpotroche grandiflora 

Mayna latifolia (Benth.) Benth. =Lindackeria latifolia 

Mayna laxiflora Benth. =Lindackeria paludosa 

Mayna paludosa Benth. =Lindackeria paludosa 

Mayna zuliana (Pittier) A. Robyns =Mayna grandifolia 

Myroxylon benthamii (Tul.) Kuntze =Xylosma benthamii 

Napimoga guianensis Aubl. =Homalium guianense 

Oncoba latifolia (Benth.) Eichler =Lindackeria latifolia 

Oncoba maynensis var. laxiflora (Benth.) Eichler =Lindackeria paludosa 

Patrisia acuminata (Spruce ex Eichler) Kuntze =Ryania angustifolia 

Patrisia affinis Kunth =Ryania dentata 

Patrisia bicolor DC. =Ryania speciosa 

Patrisia dentata Kunth =Ryania dentata 

Patrisia parviflora DC. =Ryania speciosa 

Patrisia pyrifera Rich. =Ryania pyrifera 

Patrisia speciosa (Vahl) Monach. =Ryania speciosa 

Patrisia spruceana (Monach.) Lemée [not in flora area] =Ryania spruceana 

Patrisia tomentosa (Miq.) M. Roem. =Ryania speciosa

304 


Piparea dentata Aubl. =Casearia commersoniana 

Pitumba guianensis Aubl. =Casearia pitumba 

Prockia orinocensis Cuatrec. =Banara orinocensis 

Ptychocarpus apodanthus Kuhlm. =Neoptychocarpus apodanthus 

Racoubea guianensis Aubl. =Homalium guianense 

Ryania acuminata Spruce ex Eichler =Ryania angustifolia 

Ryania bicolor (DC.) Eichler =Ryania speciosa 

Ryania candollei Miq. =Ryania speciosa 

Ryania casiquiarensis Steyerm. =Ryania speciosa 

Ryania kunthii Miq. =Ryania dentata 

Ryania mansoana auct. non Eichler 1871 =Ryania speciosa 

Ryania parviflora (DC.) Griseb. =Ryania speciosa 

Ryania pyrifera auct. non (Rich.) Uitten & Sleumer 1935, sensu J.F. Macbr. =Ryania speciosa 

Ryania pyrifera var. subuliflora Sandwith =Ryania speciosa 

Ryania pyrifera var. tomentosa (Miq.) Sleumer =Ryania speciosa 

Ryania sagotiana Eichler =Ryania pyrifera 

Ryania sagotina auct. non Eichler 1871 =Ryania dentata 

Ryania speciosa var. bicolor (A. DC.) Monach. =Ryania speciosa 

Ryania speciosa var. minor Monach. =Ryania speciosa 

Ryania speciosa var. stipularis (Linden & Planch.) Monach. =Ryania speciosa 

Ryania speciosa var. subuliflora (Sandwith) Monach. =Ryania speciosa 

Ryania speciosa var. tomentosa (Miq.) Monach. =Ryania speciosa 

Ryania stipularis auct. non Linden & Planch. 1863 =Ryania spruceana 

Ryania stipularis Linden & Planch. =Ryania speciosa 

Ryania tomentosa Miq. =Ryania speciosa 

Samyda arborea Rich. =Casearia arborea 

Samyda guidonia L. (Meliaceae) =Guarea guidonia 

Samyda hirsuta (Sw.) Poir. =Casearia hirsuta 

Samyda petiolaris Spruce ex Eichler =Laetia suaveolens 

Samyda pitumba Poir. =Casearia pitumba 

Samyda procera Poepp. =Laetia procera 

Samyda silvestris (Sw.) Poir. =Casearia sylvestris 

Samyda spinescens Sw. =Casearia spinescens 

Samyda suaveolens Poepp. =Laetia suaveolens 

Tetracoryne angustifolia Turcz. =Ryania angustifolia 

Xyladenius gladulosus Desv. ex Ham. =Banara guianensis 

Xylosma nitidum (Hell.) A. Gray ex Griseb. [not in flora area] 

Xylosma pallidifolium Sleumer ex Steyerm. & O. Huber =Xylosma benthamii 

Zuelania tremula Griseb. =Casearia tremula 

REFERENCES 

Mori, S.A. & B. Fischer. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


328. 

Olson, M., P.E. Berry, & G.A. Aymard C. 1999. Flacourtiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Sleumer, H.O. 1980. Flacourtiaceae. Flora Neotropica Monogr. 22: 1-499. 

Uittien, H. & H.O. Sleumer, 1935. Flacourtiaceae. In A. Pulle, Flora of Suriname 3(1): 282-303. 

GENTIANACEAE 

V.A. Albert, M.J. Jansen-Jacobs, P.J.M. Maas, O. Pihlar, L.Struwe 

Adenolisianthus arboreus (Spruce ex Progel) Gilg 

Celiantha bella Maguire & Steyerm. 

Celiantha chimantensis (Steyerm. & Maguire) Maguire 

Celiantha imthurniana (Oliv.) Maguire [Mt. Roraima] 

Chelonanthus alatus (Aubl.) Pulle 

Chelonanthus albus (Spruce ex Progel) V.M. Badillo 

Chelonanthus angustifolius (Kunth) Gilg 

Chelonanthus longistylus (J.G.M. Pers. & Maas) Struwe & V.A. Albert 

Chelonanthus purpurascens (Aubl.) Struwe, S. Nilsson & V.A. Albert 

Chelonanthus schomburgkii (Griseb.) Gilg 

Chelonanthus viridiflorus (Mart.) Gilg 

Chorisepalum carnosum Ewan 

AM 

AM 

BO 

BO GU 

AM 

AM 


AM BO GU 

BO GU SU FG 


AM GU 

GU 

BO GU


Chorisepalum ovatum Gleason 

Chorisepalum psychotrioides Ewan 

Chorisepalum rotundifolium Ewan 

Chorisepalum sipapoanum (Maguire) Struwe & V.A. Albert 

Coutoubea humilis Sandwith 

Coutoubea minor Kunth 

Coutoubea ramosa Aubl. 

Coutoubea reflexa Benth. 

Coutoubea spicata Aubl. 

Curtia conferta (Mart.) Knobl. 

Curtia obtusifolia (Benth.) Knobl. 

Curtia tenuifolia (Aubl.) Knobl. 

Enicostema verticillatum (L.) Engl. ex Gilg 

Irlbachia cardonae (Gleason) Maguire 

Irlbachia nemorosa (Willd. ex Roem. & Schult.) Merr. 

Irlbachia phelpsiana Maguire 

Irlbachia plantaginifolia Maguire 

Irlbachia poeppigii (Griseb.) L. Cobb & Maas 

Irlbachia pratensis (Kunth) L. Cobb & Maas 

Irlbachia pumila (Benth.) Maguire 

Irlbachia tatei (Gleason) Maguire 

Macrocarpaea autanae Weaver 

Macrocarpaea ayangannae J.R. Grant, Struwe & J.K. Boggan [Mt. 

Ayanganna] 

Macrocarpaea marahuacae Struwe & V.A. Albert 

Macrocarpaea neblinae Maguire & Steyerm. 

Macrocarpaea piresii Maguire 

Macrocarpaea rugosa Steyerm. 

Neblinantha neblinae Maguire 

Neblinantha parvifolia Maguire 

Neurotheca loeselioides (Spruce ex Progel) Baill. 

Neurotheca loeselioides subsp. loeselioides 

Potalia amara Aubl. 

Potalia elegans Struwe & V.A. Albert 

Potalia maguireorum Struwe & V.A. Albert 

Potalia resinifera Mart. 

Rogersonanthus arboreus (Britton) Maguire & B.M. Boom 

Rogersonanthus coccineus Steyerm. ex Struwe, S. Nilsson & V.A. Albert 

Rogersonanthus quelchii (N.E. Br.) Maguire & B.M. Boom 

Saccifolium bandeirae Maguire & Pires 

Schultesia benthamiana Klotzsch ex Griseb. 

Schultesia brachyptera Cham. 

Schultesia guianensis (Aubl.) Malme 

Schultesia pohliana Progel 

Schultesia subcrenata Klotzsch ex Griseb. 

Sipapoantha ostrina Maguire & B.M. Boom 

Symbolanthus argyreus (Maguire) Struwe & K. Gould 

Symbolanthus aureus Struwe & V.A. Albert 

Symbolanthus camanensis Maguire & B.M. Boom 

Symbolanthus elisabethae (M.R. Schomb.) Gilg 

Symbolanthus huachamacariensis Steyerm. 

Symbolanthus rosmarinifolius Struwe & V.A. Albert 

Symbolanthus sessilis Steyerm. & Maguire 

Symbolanthus yaviensis Steyerm. 

Tachia gracilis Benth. 

Tachia grandiflora Maguire & Weaver 

Tachia grandifolia Maguire & Weaver 

Tachia grandifolia var. grandifolia 

Tachia grandifolia var. orientalis Maguire & Weaver 

Tachia guianensis Aubl. 

Tachia longipes L. Cobb & Maas 

Tachia schomburgkiana Benth. 

Tachia smithii Maguire & Weaver 

Tapeinostemon breweri Steyerm. & Maguire 

AM BO 

BO GU 

BO GU 

AM SU 

GU 

AM BO 


AM BO GU 


AM 

AM 


BO GU 

AM BO GU 

AM BO GU 

AM 

AM 

BO 

AM 

AM 

AM BO GU 

AM 

GU 

AM 

AM 

AM 

BO GU 

AM 

AM 

AM GU SU FG 

GU SU 

SU FG 

AM 

AM 

AM 

AM BO 

AM 

AM GU 

AM 

BO GU SU? 


BO GU SU FG 

AM BO GU SU 

BO GU 

AM 

AM 

BO 

AM 

AM BO GU 

AM 

AM 

BO 

AM BO 

AM GU 

FG 

AM BO 

AM BO 

AM BO 

GU SU FG 

SU 

BO GU 

GU 

AM BO

306 


Tapeinostemon jauaensis Steyerm. & Maguire 

Tapeinostemon longiflorum Maguire & Steyerm. 

Tapeinostemon longiflorum var. australe Maguire & Steyerm. [Sierra de la 

Neblina] 

Tapeinostemon longiflorum var. longiflorum [Sierra de la Neblina] 

Tapeinostemon rugosum Maguire & Steyerm. [Macizo del Chimantá] 

Tapeinostemon sessiliflorum (Humb. & Bonpl. ex Schult.) Pruski & S.F. Sm. 

Tapeinostemon spenneroides Benth. 

Tetrapollinia caerulescens (Aubl.) Maguire & B.M. Boom 

Voyria acuminata Benth. 

Voyria aphylla (Jacq.) Pers. 

Voyria aurantiaca Splitg. 

Voyria caerulea Aubl. 

Voyria chionea Benth. 

Voyria clavata Splitg. 

Voyria corymbosa Splitg. 

Voyria corymbosa subsp. alba (Standl.) Ruyters & Maas 

Voyria corymbosa subsp. corymbosa 

Voyria flavescens Griseb. 

Voyria pittieri (Standl.) L.O. Williams 

Voyria rosea Aubl. 

Voyria spruceana Benth. 

Voyria tenella Hook. 

Voyria tenuiflora Griseb. 

Voyriella parviflora (Miq.) Miq. 

AM 

AM 


Adenolisianthus virgatus (Progel) Gilg =Chelonanthus alatus 

Apophragma tenuifolium (Aubl.) Griseb. =Curtia tenuifolia 

Brachycodon pumilus (Benth.) Progel =Irlbachia pumila 

Brachycodon ramosissimus (Benth.) Progel =Irlbachia poeppigii 

Calolisianthus imthurnianus (Oliv.) Gleason =Celiantha imthurniana 

Calolisianthus tatei Gleason =Irlbachia tatei 

Calolisianthus tepuiensis Gleason =Rogersonanthus arboreus 

Chelonanthus acutangulus (Ruiz & Pav.) Gilg =Chelonanthus alatus 

Chelonanthus arboreus Britton =Rogersonanthus arboreus 

Chelonanthus bifidus (Kunth) Gilg =Chelonanthus angustifolius 

Chelonanthus campanuloides (Spruce ex Benth.) Gilg =Irlbachia pratensis 

Chelonanthus cardonae Gleason =Irlbachia cardonae 

Chelonanthus chelonoides (L. f.) Gilg =Chelonanthus alatus 

Chelonanthus fruticosus Maguire & B.M. Boom =Adenolisianthus arboreus 

Chelonanthus pyriformis Gleason =Chelonanthus angustifolius 

Chelonanthus uliginosus (Griseb.) Gilg =Chelonanthus purpurascens 

Chorisepalum acuminatum Steyerm. =Chorisepalum psychotrioides 

Chorisepalum breweri Steyerm. & Maguire =Chorisepalum ovatum 

Chorisepalum ovatum var. sipapoanum Maguire =Chorisepalum sipapoanum 

Chorisepalum psychotrioides var. acuminatum (Steyerm.) Maguire =Chorisepalum psychotrioides 

Claytonia nemorosa Willd. ex Roem. & Schult. =Irlbachia nemorosa 

Coutoubea racemosa G. Mey. =Coutoubea ramosa 

Coutoubea ramosa var. longifolia Benth. =Coutoubea ramosa 

Coutoubea ramosa f. racemosa (G. Mey.) Jonker =Coutoubea ramosa 

Coutoubea ramosa var. racemosa (G. Mey.) Benth. =Coutoubea ramosa 

Curtia quadrifolia Maguire =Curtia conferta 

Curtia tenella (Mart.) Cham. =Curtia tenuifolia 

Curtia tenuifolia subsp. tenella (Mart.) Grothe & Maas =Curtia tenuifolia 

Ditomaga nemorosa (Willd. ex Roem. & Schult.) Raf. =Irlbachia nemorosa 

Exacum guianense Aubl. =Schultesia guianensis 

Exacum racemosum (G. Mey.) Roem. & Schult. =Coutoubea ramosa 

Exacum spicatum (Aubl.) Vahl =Coutoubea spicata 

Exacum tenuifolium Aubl. =Curtia tenuifolia 

Gentiana aphylla Jacq. =Voyria aphylla 

Gentiana verticillata L. =Enicostema verticillatum 

Helia acutangula (Ruiz & Pav.) Kuntze =Chelonanthus alatus 

Helia alata (Aubl.) Kuntze =Chelonanthus alatus 

BO 

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BO GU SU FG 

BO GU SU FG 

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Helia alba (Spruce ex Progel) Kuntze =Chelonanthus albus 

Helia angustifolia (Kunth) Kuntze =Chelonanthus angustifolius 

Helia arborea (Spruce ex Progel) Kuntze =Adenolisianthus arboreus 

Helia bifida (Kunth) Kuntze =Chelonanthus angustifolius 

Helia caerulescens (Aubl.) Kuntze =Tetrapollinia caerulescens 

Helia campanuloides (Spruce ex Benth.) Kuntze =Irlbachia pratensis 

Helia chelonoides (L. f.) Kuntze =Chelonanthus alatus 

Helia elisabethae (M.R. Schomb.) Kuntze =Symbolanthus elisabethae 

Helia nemorosa (Willd. ex Roem. & Schult.) Kuntze =Irlbachia nemorosa 

Helia pratensis (Kunth) Kuntze =Irlbachia pratensis 

Helia pumila (Benth.) Kuntze =Irlbachia pumila 

Helia purpurascens (Aubl.) Kuntze =Chelonanthus purpurascens 

Helia ramosissima (Benth.) Kuntze =Irlbachia poeppigii 

Helia schomburgkii (Griseb.) Kuntze =Chelonanthus schomburgkii 

Helia trifidus (Kunth) Kuntze =Chelonanthus alatus 

Helia uliginosa (Griseb.) Kuntze =Chelonanthus purpurascens 

Irlbachia alata (Aubl.) Maas =Chelonanthus alatus 

Irlbachia alata subsp. alata =Chelonanthus alatus 

Irlbachia alata subsp. alba (Spruce ex Progel) J.G.M. Pers. & Maas =Chelonanthus albus 

Irlbachia alata subsp. angustifolia (Kunth) J.G.M. Pers. & Maas =Chelonanthus angustifolius 

Irlbachia alata subsp. arborea (Spruce ex Progel) J.G.M. Pers. & Maas =Adenolisianthus arboreus 

Irlbachia alata subsp. longistyla J.G.M. Pers. & Maas =Chelonanthus longistylus 

Irlbachia alata subsp. viridiflora (Mart.) J.G.M. Pers. & Maas =Chelonanthus viridiflorus 

Irlbachia bonplandiana Fenzl =Irlbachia nemorosa 

Irlbachia caerulescens (Aubl.) Griseb. =Tetrapollinia caerulescens 

Irlbachia paruana Maguire =Irlbachia cardonae 

Irlbachia purpurascens (Aubl.) Maas =Chelonanthus purpurascens 

Irlbachia quelchii (N.E. Br.) Maas =Rogersonanthus quelchii 

Irlbachia ramosissima (Benth.) Maguire =Irlbachia poeppigii 

Irlbachia recurva (Benth.) Progel =Irlbachia nemorosa 

Irlbachia subcordata (Benth.) Progel =Irlbachia nemorosa 

Leiothamnus elisabethae M.R. Schomb. =Symbolanthus elisabethae 

Leiphaimos alba Standl. =Voyria corymbosa subsp. alba 

Leiphaimos aphylla (Jacq.) Gilg =Voyria aphylla 

Leiphaimos aurantiaca (Splitg.) Miq. =Voyria aurantiaca 

Leiphaimos calycina (Splitg. ex A. DC.) Miq. =Voyria caerulea 

Leiphaimos chionea (Benth.) Gilg =Voyria chionea 

Leiphaimos flavescens (Griseb.) Gilg =Voyria flavescens 

Leiphaimos leucantha (Miq.) Gilg =Voyria tenuiflora 

Leiphaimos montana Jonker =Voyria corymbosa subsp. alba 

Leiphaimos obconica auct., non (Progel) Gilg =Voyria tenella 

Leiphaimos pittieri Standl. =Voyria pittieri 

Leiphaimos spruceana (Benth.) Gilg =Voyria spruceana 

Leiphaimos tenella (Hook.) Miq. =Voyria tenella 

Leiphaimos tenuiflora (Griseb.) Miq. =Voyria tenuiflora 

Lisianthus acutangulus Ruiz & Pav. =Chelonanthus alatus 

Lisianthus acutilobus Steyerm. =Irlbachia cardonae 

Lisianthus alatus Aubl. =Chelonanthus alatus 

Lisianthus albus Spruce ex Progel =Chelonanthus albus 

Lisianthus angustifolius Kunth =Chelonanthus angustifolius 

Lisianthus arboreus Spruce ex Progel =Adenolisianthus arboreus 

Lisianthus bifidus Kunth =Chelonanthus angustifolius 

Lisianthus caerulescens Aubl. =Tetrapollinia caerulescens 

Lisianthus campanuloides Spruce ex Benth. =Irlbachia pratensis 

Lisianthus cardonae (Gleason) Steyerm. =Irlbachia cardonae 

Lisianthus chelonoides L. f. =Chelonanthus alatus 

Lisianthus chimantensis Steyerm. & Maguire =Celiantha chimantensis 

Lisianthus elisabethae (M.R. Schomb.) Griseb. =Symbolanthus elisabethae 

Lisianthus fistulosus Poir. =Chelonanthus alatus 

Lisianthus grandiflorus Aubl. =Chelonanthus alatus 

Lisianthus imthurnianus Oliv. =Celiantha imthurniana 

Lisianthus jauaensis Steyerm. & Maguire =Irlbachia tatei 

Lisianthus pratensis Kunth =Irlbachia pratensis 

Lisianthus pumilus Benth. =Irlbachia pumila

308 


Lisianthus purpurascens Aubl. =Chelonanthus purpurascens 

Lisianthus pyriformis (Gleason) Steyerm. =Chelonanthus angustifolius 

Lisianthus quelchii N.E. Br. =Rogersonanthus quelchii 

Lisianthus ramosissimus Benth. =Irlbachia poeppigii 

Lisianthus recurvus Benth. =Irlbachia nemorosa 

Lisianthus scabridulus Steyerm. =Irlbachia tatei 

Lisianthus schomburgkii Griseb. =Chelonanthus schomburgkii 

Lisianthus subcordatus Benth. =Irlbachia nemorosa 

Lisianthus tatei (Gleason) Steyerm. =Irlbachia tatei 

Lisianthus trifidus Kunth =Chelonanthus alatus 

Lisianthus uliginosus Griseb. =Chelonanthus purpurascens 

Lisianthus virgatus Progel =Chelonanthus alatus 

Lisyanthus alatus Aubl. =Chelonanthus alatus 

Lisyanthus angustifolius (Kunth) Griseb. =Chelonanthus angustifolius 

Lisyanthus bifidus (Kunth) Griseb. =Chelonanthus angustifolius 

Lisyanthus caerulescens Aubl. =Tetrapollinia caerulescens 

Lisyanthus purpurascens Aubl. =Chelonanthus purpurascens 

Lisyanthus trifidus (Kunth) Griseb. =Chelonanthus alatus 

Lisyanthus uliginosus Griseb. =Chelonanthus purpurascens 

Macrocarpaea arborea (Britton) Ewan =Rogersonanthus arboreus 

Macrocarpaea cerronis Ewan =Rogersonanthus arboreus 

Macrocarpaea quelchii (N.E. Br.) Ewan =Rogersonanthus quelchii 

Macrocarpaea salicifolia Ewan =Rogersonanthus arboreus 

Macrocarpaea tepuiensis (Gleason) Steyerm. =Rogersonanthus arboreus 

Octopleura loeselioides Spruce ex Progel =Neurotheca loeselioides 

Pagaea nemorosa (Willd. ex Roem. & Schult.) Gilg =Irlbachia nemorosa 

Pagaea poeppigii Griseb. =Irlbachia poeppigii 

Pagaea pumila (Benth.) Gilg =Irlbachia pumila 

Pagaea ramosissima (Benth.) Ewan =Irlbachia poeppigii 

Pagaea recurva (Benth.) Benth. & Hook. f. =Irlbachia nemorosa 

Pagaea subcordata (Benth.) Benth. & Hook. f. =Irlbachia nemorosa 

Rogersonanthus tepuiensis (Gleason) Maguire & B.M. Boom =Rogersonanthus arboreus 

Schuebleria conferta Mart. =Curtia conferta 

Schuebleria obtusifolia Benth. =Curtia obtusifolia 

Schuebleria tenella Mart. =Curtia tenuifolia 

Schuebleria tenuifolia (Aubl.) G. Don =Curtia tenuifolia 

Schultesia brachyptera f. heterophylla (Miq.) Jonker =Schultesia brachyptera 

Schultesia heterophylla Miq. =Schultesia brachyptera 

Schultesia schomburgkiana Progel =Schultesia benthamiana 

Stahelia spenneroides (Benth.) Jonker =Tapeinostemon spenneroides 

Stahelia surinamensis Jonker =Tapeinostemon spenneroides 

Symbolanthus aracamuniensis Steyerm. =Symbolanthus argyreus 

Symbolanthus calygonus (Ruiz & Pav.) Griseb. [s.l.] =Symbolanthus elisabethae 

Symbolanthus elisabethae subsp. occidentalis Maguire & B.M. Boom =Symbolanthus huachamacariensis 

Symbolanthus quelchii (N.E. Br.) Gleason =Rogersonanthus quelchii 

Symbolanthus yutajensis Maguire & B.M. Boom =Symbolanthus yaviensis 

Tapeinostemon borrerioides Benth. ex Knobl. [nomen nudum] =Tapeinostemon sessiliflorum 

Tapeinostemon capitatum Benth. =Tapeinostemon sessiliflorum 

Tapeinostemon ptariense Steyerm. =Tapeinostemon spenneroides 

Tapeinostemon surinamensis (Jonker) Steyerm. =Tapeinostemon spenneroides 

Voyria araguensis H. Karst. =Voyria aphylla 

Voyria parviflora Miq. =Voyriella parviflora 

Voyriella oxycarpha Sandwith =Voyriella parviflora 

Wurdackanthus argyreus Maguire =Symbolanthus argyreus 

REFERENCES 

Grothe, E., and P.J.M. Maas. 1984. A scanning electron microscopy study of the seed coat structure of Curtia Chamisso and 

Schlechtendahl and Hockinia Gardner (Gentianaceae). Proceedings Koninklijke Nederlandse Akademie van Wetenschappen. 

Series C. 87: 33–42. 

Jonker, F.P. 1936. Gentianaceae. In A. Pulle, Flora of Suriname 4(1): 400-427. 

Kuntze, C.E.O. 1891. Revisio Generum Plantarum par. 2. Leipzig. 

Leeuwenberg, A.J.M. & P.W. Leenhouts. 1980. Taxonomy. Pages 8–96 in: Engler and Prantl’s Die Natürlichen Pflanzenfamilien, 

Angiospermae: Ordnung Gentianales, Fam. Loganiaceae. Vol. 28b (1). Leeuwenberg, A. J. M., ed. Duncker and Humblot, 

Berlin


Maas-van de Kamer, H. & P. Maas. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 328-334. 

Maas, P.J.M. 1985. Nomenclatural notes on neotropical Lisyantheae. Proc. Kon. Ned. Akad. Wetensch. (ser. c) 88(4): 405-412. 

Maas, P.J.M., S. Nilsson, A.M.C. Hollants, B.J.H. ter Welle, H. Persoon, & E.C.H. van Heusden. 1984. Systematic studies in 

neotropical Gentianaceae - the Lisianthus complex. Acta Bot. Neerl. 32: 371-374. 

Maas, P.J.M. & P. Ruyters. 1986. Voyria and Voyriella (Saprophytic Gentianaceae). Flora Neotropica Monogr. 41: 1-93. 

Machado, I.C.S., I. Sazima, and M. Sazima. 1998. Bat pollination of the terrestrial herb Irlbachia alata (Gentianaceae) in 

northeastern Brazil. Plant Systematics and Evolution 209: 231–237. 

Maguire, B. 1981. Gentianaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 11. Mem. N.Y. Bot. Gard. 

32: 1-391. 

Maguire, B. & B.M. Boom. 1989. Gentianaceae (part 3). In B. Maguire & collaborators, Botany of the Guayana Highland part 

13. Memoirs of the New York Bot. Garden 51: 1-127. 

Pringle, J.S. 1995. Gentianaceae. Pages 1–131 in: Flora of Ecuador. Vol. 53. G. Harling and L. Andersson, eds. Department of 

Systematic Botany, Göteborg University, Göteborg. 

Struwe, L., and V.A. Albert. 1998. Lisianthius (Gentianaceae), its probable homonym Lisyanthus, and the priority of Helia over 

Irlbachia as its substitute. Harvard Papers in Botany 3: 63–71. 

Struwe, L., V.A. Albert, and B. Bremer. 1994 (1995). Cladistics and family level classification of the Gentianales. Cladistics 10: 

175–206. 

Struwe, L., P.J.M. Maas, and V.A. Albert. 1997. Aripuana cullmaniorum, a new genus and species of Gentianaceae from whitesands 

of southeastern Amazonas, Brazil. Harvard Papers of Botany 2: 235–253. 

Struwe, L., P.J.M. Maas, O. Pihlar, & V.A. Albert. 1999. Gentianaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. 

Holst, Flora of the Venezuelan Guayana 5: 474-542. 

Struwe, L., M. Thiv, J. W. Kadereit, A. S-R. Pepper, T.J. Motley, P.J. White, J.H.E. Rova, K. Potgieter, & V.A. Albert. 1998. 

Saccifolium (Saccifoliaceae), an endemic of Sierra de la Neblina on the Brazilian-Venezuelan frontier, is related to a temperatealpine 

lineage of Gentianaceae. Harvard Papers in Botany 3: 199–214. 

Alloplectus cristatus (L.) Mart. 

Alloplectus cristatus var. epirotes Leeuwenb. 

Alloplectus savannarum C.V. Morton 

Besleria flavovirens Nees & Mart. 

Besleria gibbosa (Poepp.) Hanst. 

Besleria insolita C.V. Morton 

Besleria laxiflora Benth. 

Besleria parviflora L.E. Skog & Steyerm. 

Besleria patrisii DC. 

Besleria penduliflora Fritsch 

Besleria saxicola C.V. Morton 

Besleria sp. A 

Besleria sp. B 

Besleria sp. C 

Chrysothemis dichroa Leeuwenb. 

Chrysothemis pulchella (Donn ex Sims) Decne. 

Chrysothemis rupestris (Benth.) Leeuwenb. 

Chrysothemis villosa (Benth.) Leeuwenb. 

Codonanthe calcarata (Miq.) Hanst. 

Codonanthe crassifolia (H. Focke) C.V. Morton 

Codonanthopsis dissimulata (H.E. Moore) Wiehler 

Codonanthopsis ulei Mansf. 

Columnea calotricha Donn. Sm. 

Columnea guianensis C.V. Morton 

Columnea microsepala (C.V. Morton) Kvist & L.E. Skog 

Columnea oerstediana Klotzsch ex Oerst. 

Columnea sanguinea (Pers.) Hanst. 

Corytoplectus congestus (Linden ex Hanst.) Wiehler 

Corytoplectus deltoideus (C.V. Morton) Wiehler 

Cremersia platula Feuillet & L.E. Skog 

Diastema racemiferum Benth. 

Drymonia antherocycla Leeuwenb. 

Drymonia coccinea (Aubl.) Wiehler 

Drymonia psilocalyx Leeuwenb. 

GESNERIACEAE 

C. Feuillet, L.E. Skog, J.A. Steyermark† 

GU 

GU 

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AM BO 

GU FG 


AM BO GU 

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AM 

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SU FG

310 


Drymonia serrulata (Jacq.) Mart. 

Episcia fimbriata Fritsch 

Episcia reptans Mart. 

Episcia sp. A 

Episcia sp. B 

Episcia sphalera Leeuwenb. 

Episcia xantha Leeuwenb. 

Gloxinia perennis (L.) Fritsch [cultivated or escaped from cultivation] 

Gloxinia purpurascens (Rusby) Wiehler 

Koellikeria erinoides (DC.) Mansf. 

Kohleria hirsuta (Kunth) Regel 

Lampadaria rupestris Feuillet & L.E. Skog 

Lembocarpus amoenus Leeuwenb. 

Napeanthus angustifolius Feuillet & L.E. Skog 

Napeanthus jelskii Fritsch FG 

Napeanthus macrostoma Leeuwenb. 

Napeanthus rupicola Feuillet & L.E. Skog 

Nautilocalyx adenosiphon (Leeuwenb.) Wiehler 

Nautilocalyx arenarius L.E. Skog & Steyerm. 

Nautilocalyx bryogeton (Leeuwenb.) Wiehler 

Nautilocalyx cataractarum Wiehler 

Nautilocalyx chimantensis L.E. Skog & Steyerm. 

Nautilocalyx coccineus Feuillet & L.E. Skog 

Nautilocalyx cordatus (Gleason) L.E. Skog 

Nautilocalyx fasciculatus L.E. Skog & Steyerm. 

Nautilocalyx kohlerioides (Leeuwenb.) Wiehler 

Nautilocalyx maguirei L.E. Skog & Steyerm. 

Nautilocalyx mimuloides (Benth.) C.V. Morton 

Nautilocalyx pallidus (Sprague) Sprague 

Nautilocalyx pemphidius L.E. Skog 

Nautilocalyx pictus (Hook.) Sprague 

Nautilocalyx porphyrotrichus (Leeuwenb.) Wiehler 

Nautilocalyx punctatus Wiehler 

Nautilocalyx resioides (Leeuwenb.) Wiehler 

Nautilocalyx sp. A 

Nautilocalyx sp. B 

Nautilocalyx sp. C 

Nautilocalyx sp. D 

Nautilocalyx sp. E 

Nautilocalyx sp. F 

Nautilocalyx sp. G 

Nautilocalyx villosus (Kunth & Bouché) Sprague 

Paradrymonia anisophylla Feuillet & L.E. Skog 

Paradrymonia barbata Feuillet & L.E. Skog 

Paradrymonia campostyla (Leeuwenb.) Wiehler 

Paradrymonia ciliosa (Mart.) Wiehler 

Paradrymonia densa (C.H. Wright) Wiehler 

Paradrymonia longifolia (Poepp.) Wiehler 

Paradrymonia maculata (Hook. f.) Wiehler 

Paradrymonia metamorphophylla (J.D. Sm.) Wiehler 

Paradrymonia sp. A 

Paradrymonia sp. B 

Paradrymonia sp. C 

Paradrymonia sp. D 

Paradrymonia sp. E 

Rhoogeton cyclophyllus Leeuwenb. 

Rhoogeton viviparus Leeuwenb. 

Sinningia elatior (Kunth) Chautems 

Sinningia incarnata (Aubl.) D.L. Denham 

Sinningia schomburgkiana (Kunth & Bouché) Chautems 

Tylopsacas cuneatum (Gleason) Leeuwenb. 


Achimenes erinoides DC. =Koellikeria erinoides 

Alloplectus calotrichus (J.D. Sm.) Stearn =Columnea calotricha 


AM 

AM GU SU FG 

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Alloplectus circinatus Mart. =Drymonia coccinea 

Alloplectus coccineus (Aubl.) Mart. =Drymonia coccinea 

Alloplectus congestus Linden ex Hanst. =Corytoplectus congestus 

Alloplectus deltoideus C.V. Morton =Corytoplectus deltoideus 

Alloplectus meridensis Klotzsch ex Hanst. [not in flora area] 

Alloplectus metamorphophyllus J.D. Sm. =Paradrymonia metamorphophylla 

Alloplectus microsepalus C.V. Morton =Columnea microsepala 

Alloplectus pallidus Sprague =Nautilocalyx pallidus 

Alloplectus patrisii DC. =Drymonia coccinea 

Besleria bivalvis L. f. (Mendonciaceae) =Mendoncia bivalvis 

Besleria bivalvis L. f. (Mendonciaceae) =Mendoncia bivalvis 

Besleria coccinea Aubl. =Drymonia coccinea 

Besleria incarnata Aubl. =Sinningia incarnata 

Besleria lutea L. [not in flora area] 

Besleria maasii Wiehler =Besleria patrisii 

Besleria pulchella Donn ex Sims =Chrysothemis pulchella 

Besleria sanguinea Pers. =Columnea sanguinea 

Besleria serrulata Jacq. =Drymonia serrulata 

Besleria spectabilis Kunth =Drymonia serrulata 

Besleria verecunda C.V. Morton =Besleria patrisii 

Besleria violacea Aubl. (Bignoniaceae) =Schlegelia violacea 

Brachyloma rhodomallos (Hanst.) Hanst. =Kohleria hirsuta 

Centrosolenia densa (C.H. Wright) Sprague =Paradrymonia densa 

Centrosolenia hirsuta Benth. =Nautilocalyx cordatus 

Codonanthe bipartita L.B. Sm. =Codonanthe calcarata 

Codonanthe carnosa Hanst. [not in flora area] 

Codonanthe confusa Sandwith =Codonanthe crassifolia 

Codonanthe dissimulata H.E. Moore =Codonanthopsis dissimulata 

Codonanthe ulei (Mansf.) H.E. Moore =Codonanthopsis ulei 

Codonanthopsis ulei Fritsch [not in flora area] 

Columnea affinis C.V. Morton =Columnea sanguinea 

Columnea aureonitens Hook. =Columnea sanguinea 

Columnea calotricha var. austroamericana C.V. Morton =Alloplectus savannarum 

Columnea coccinea (Aubl.) Kuntze =Drymonia coccinea 

Columnea scandens L. [not in flora area] 

Columnea steyermarkii C.V. Morton =Alloplectus savannarum 

Crantzia patrisii (DC.) Fritsch =Drymonia coccinea 

Dalbergaria aureonitens (Hook.) Wiehler =Columnea sanguinea 

Dalbergaria sanguinea (Pers.) Steud. =Columnea sanguinea 

Drymonia calcarata Mart. =Drymonia serrulata 

Drymonia campostyla Leeuwenb. =Paradrymonia campostyla 

Drymonia cristata Miq. =Drymonia serrulata 

Drymonia sarmentosula Lem. =Drymonia serrulata 

Drymonia turrialvae Hanst. [not in flora area] 

Episcia adenosiphon Leeuwenb. =Nautilocalyx adenosiphon 

Episcia bryogeton Leeuwenb. =Nautilocalyx bryogeton 

Episcia ciliosa (Mart.) Hanst. =Paradrymonia ciliosa 

Episcia cordata Gleason =Nautilocalyx cordatus 

Episcia cuneata Gleason =Tylopsacas cuneatum 

Episcia densa C.H. Wright =Paradrymonia densa 

Episcia hirsuta (Benth.) Hanst. =Nautilocalyx cordatus 

Episcia maculata Hook. f. =Paradrymonia maculata 

Episcia mimuloides Benth. =Nautilocalyx mimuloides 

Episcia porphyrotricha Leeuwenb. =Nautilocalyx porphyrotrichus 

Episcia resioides Leeuwenb. =Nautilocalyx resioides 

Gesneria caracasana Otto & Dietr. =Sinningia incarnata 

Gesneria elatior Kunth =Sinningia elatior 

Gesneria gollmeriana Hanst. =Sinningia incarnata 

Gesneria hirsuta Kunth =Kohleria hirsuta 

Gesneria lindeniana Brongn. =Sinningia incarnata 

Gesneria rubricaulis Kunth & Bouché =Kohleria hirsuta 

Gesneria schomburgkiana Kunth & Bouché =Sinningia schomburgkiana 

Gesneria vargasii DC. =Sinningia incarnata 

Gloxinia trichantha Miq. =Gloxinia perennis

312 


Hypocyrta crassifolia H. Focke =Codonanthe crassifolia 

Hypocyrta gibbosa Poepp. =Besleria gibbosa 

Isoloma pycnosuzygium Donn. Sm. =Kohleria hirsuta 

Kohleria longipedunculata Hanst. =Kohleria hirsuta 

Kohleria pycnosuzygium (Donn. Sm.) V.M. Badillo =Kohleria hirsuta 

Kohleria rhodomallos (Hanst.) Hanst. =Kohleria hirsuta 

Kohleria rubricaulis (Kunth & Bouché) Hassk. =Kohleria hirsuta 

Nautilocalyx lacteus Sandwith =Nautilocalyx pictus 

Nematanthus calcaratus Miq. =Codonanthe calcarata 

Pentadenia microsepala (C.V. Morton) Wiehler =Columnea microsepala 

Rechsteineria caracasana (Otto & Dietr.) Kuntze =Sinningia incarnata 

Rechsteineria gollmeriana (Hanst.) Kuntze =Sinningia incarnata 

Rechsteineria incarnata (Aubl.) Leeuwenb. =Sinningia incarnata 

Rechsteineria schomburgkiana (Kunth & Bouché) Leeuwenb. =Sinningia schomburgkiana 

Rhoogeton leeuwenbergianus C.V. Morton =Rhoogeton viviparus 

Tussacia pulchella (Donn ex Sims) Rchb. ex Walp. =Chrysothemis pulchella 

Tussacia rupestris Benth. =Chrysothemis rupestris 

Tylosperma cuneatum (Gleason) Leeuwenb. =Tylopsacas cuneatum 

REFERENCES 

Feuillet, C. & L.E. Skog. 2002. Gesneriaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 


Gard. 76 (2): 334-344. 

Feuillet, C. & L.E. Skog. 2002 (2003). Novae Gesneriaceae Neotropicarum XI. New genera and species from the Guianas. 

Brittonia 54 (4): 344-351. 

Feuillet, C. & L.E. Skog. 2002 (2003). Novae Gesneriaceae Neotropicarum XII. New species of Gesneriaceae from the Guianas. 

Brittonia 54 (4): 352-361. 

Feuillet, C. & J.A. Steyermark. 1999. Gesneriaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Leeuwenberg, A.J.M. 1958. The Gesneriaceae of Guiana. Acta Bot. Neerl. 7: 291-444. 

Leeuwenberg, A.J.M. 1984. Gesneriaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 592-650. 

Morton, C.V. 1939. A revision of Besleria. Contr. U.S. Nat. Herb. 26: 395-474. 

Wiehler, H.J. 1978. The genera Episcia, Alsobia, Nautilocalyx, and Paradrymonia. Selbyana 5: 11-60. 

HALORAGACEAE 

L.J. Raechal 

Laurembergia tetrandra (Schott ex Spreng.) Kanitz 

Laurembergia tetrandra subsp. tetrandra [expected in the Guianas] 

Myriophyllum aquaticum (Vell.) Verdc. 


Haloragis tetrandra Schott ex Spreng. =Laurembergia tetrandra 

Myriophyllum proserpinacoides Gillies ex Hook. & Arn. =Myriophyllum aquaticum 

REFERENCES 

Orchard, A.E. 1981. A revision of South American Myriophyllum, and its repercussions on some Australian and North American 

species. Brunonia 4: 27-65. 

Raechal, L.J. 1999. Haloragaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 581-582. 

Hernandia guianensis Aubl. 

Sparattanthelium aruakorum Tutin 

Sparattanthelium guianense Sandwith 

Sparattanthelium tupiniquinorum Mart. 

Sparattanthelium uncigerum (Meisn.) Kubitzki 

Sparattanthelium wonotoboense Kosterm. 

HERNANDIACEAE 



Hernandia sonora Kosterm. =Hernandia guianensis 

Hernandia sonora L. [not in flora area - Asian] 

BO 

GU? FG? 


GU SU 

GU FG? 


BO GU SU FG 

GU SU


Sparattanthelium botocudorum Mart. [not in flora area] 

Sparattanthelium botocudorum var. uncigerum Meisn. =Sparattanthelium uncigerum 

Sparattanthelium macusiorum A.C. Sm. =Sparattanthelium wonotoboense 

Sparattanthelium melinonis Baill. ex Lemée [nomen nudum] =Sparattanthelium uncigerum 

REFERENCES 

Kostermans, A.J.G.H. 1937. Hernandiaceae. In A. Pulle, Flora of Suriname 2(1): 338-344. 

Kubitzki, K.U. 1969. Monographie der Hernandiaceen. Bot. Jahrb. Syst. 89: 78-205. 

Kubitzki, K.U. 1976. Hernandiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 485- 

486. 

Kubitzki, K.U. 1982. Hernandiaceae. In Z. Luces de Febres, Flore de Venezuela 4(2): 317-324. 

Miller, J.S. & P.E. Berry. 1999. Hernandiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Mori, S.A. & J.L. Brown. 2002. Hernandiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 


Gard. 76 (2): 344-347. 

HIPPOCRATEACEAE 

A.R.A. Görts-van Rijn, J.P. Hedin, A.M.W. Mennega† 

Anthodon decussatum Ruiz & Pav. 

Cheiloclinium anomalum Miers 

Cheiloclinium belizense (Standl.) A.C. Sm. 

Cheiloclinium cognatum (Miers) A.C. Sm. 

Cheiloclinium diffusiflorum (Miers) A.C. Sm. 

Cheiloclinium habropodum A.C. Sm. 

Cheiloclinium hippocrateoides (Peyr.) A.C. Sm. 

Cheiloclinium klugii A.C. Sm. 

Cheiloclinium obtusum A.C. Sm. 

Cuervea kappleriana (Miq.) A.C. Sm. 

Elachyptera floribunda (Benth.) A.C. Sm. 

Hippocratea volubilis L. 

Hylenaea comosa (Sw.) Miers 

Hylenaea praecelsa (Miers) A.C. Sm. [expected in the Guianas] 

Hylenaea unguiculata Mennega 

Peritassa compta Miers 

Peritassa dulcis (Benth.) Miers 

Peritassa huanucana (Loes.) A.C. Sm. 

Peritassa laevigata (Hoffmanns. ex Link) A.C. Sm. 

Peritassa peruviana (Miers) A.C. Sm. 

Peritassa pruinosa (Seem.) A.C. Sm. 

Prionostemma aspera (Lam.) Miers 

Pristimera caudata Mennega 

Pristimera holdeniana (A.C. Sm.) A.C. Sm. 

Pristimera nervosa (Miers) A.C. Sm. 

Pristimera tenuiflora (Mart. ex Peyr.) A.C. Sm. 

Pristimera verrucosa (Kunth) Miers 

Salacia amplectens A.C. Sm. 

Salacia cordata (Miers) Mennega 

Salacia duckei A.C. Sm. 

Salacia elliptica (Mart. ex Schult.) G. Don 

Salacia gigantea Loes. 

Salacia impressifolia (Miers) A.C. Sm. 

Salacia insignis A.C. Sm. 

Salacia juruana Loes. 

Salacia kanukuensis A.C. Sm. 

Salacia maburensis Mennega 

Salacia macrantha A.C. Sm. 

Salacia miqueliana Loes. 

Salacia multiflora (Lam.) DC. 

Salacia multiflora subsp. mucronata (Rusby) Mennega 

Salacia multiflora subsp. multiflora 

Salacia opacifolia (J.F. Macbr.) A.C. Sm. 

Salacia pachyphylla (Miers) Peyr. 

AM BO FG 




AM GU SU FG 

AM 


AM BO 

BO 





SU 

AM BO GU SU 

GU 



AM BO 

AM BO GU 


SU 

GU 


BO DA SU FG 

BO GU 


AM BO GU SU 

SU? FG 


AM 


SU FG 



GU 

BO 

SU FG 




AM 

GU

314 

Tontelea attenuata Miers 

Tontelea coriacea A.C. Sm. 

Tontelea corymbosa (Huber) A.C. Sm. 

Tontelea cylindrocarpa (A.C. Sm.) A.C. Sm. 

Tontelea glabra A.C. Sm. 

Tontelea laxiflora (Benth.) A.C. Sm. 

Tontelea mauritioides (A.C. Sm.) A.C. Sm. 

Tontelea nectandrifolia (A.C. Sm.) A.C. Sm. 

Tontelea ovalifolia (Miers) A.C. Sm. 

Tontelea ovalifolia subsp. ovalifolia 

Tontelea ovalifolia subsp. richardii (Peyr.) Görts & Mennega 

Tontelea sandwithii A.C. Sm. 



Amphizoma laxiflora (Benth.) Miers =Tontelea laxiflora 

Anthodon grandiflorus Benth. =Salacia impressifolia 

Anthodon laxiflorus Benth. =Tontelea laxiflora 

Anthodus ellipticus Mart. ex Schult. =Salacia elliptica 

Cheiloclinium divaricata A.C. Sm. =Cheiloclinium hippocrateoides 

Cheiloclinium glaziovii A.C. Sm. =Cheiloclinium anomalum 

Cheiloclinium gleasonianum (A.C. Sm.) A.C. Sm. =Cheiloclinium hippocrateoides 

Cheiloclinium jenmanii A.C. Sm. =Cheiloclinium belizense 

Cheiloclinium krukovii (A.C. Sm.) A.C. Sm. =Cheiloclinium hippocrateoides 

Cheiloclinium lineolatum (A.C. Sm.) A.C. Sm. =Cheiloclinium cognatum 

Cheiloclinium lucidum A.C. Sm. =Cheiloclinium diffusiflorum 

Cheiloclinium meianthemum A.C. Sm. =Cheiloclinium belizense 

Cheiloclinium minutiflorum (A.C. Sm.) A.C. Sm. [not in flora area] 

Cheiloclinium parviflorum (Miers) A.C. Sm. =Cheiloclinium diffusiflorum 

Cheiloclinium podostemmum (Sandwith) A.C. Sm. =Cheiloclinium belizense 

Cuervea granatensis Miers =Cuervea kappleriana 

Cuervea mitchellae (I.M. Johnst.) A.C. Sm. =Cuervea kappleriana 

Cuervea ovalifolia Miers =Tontelea ovalifolia 

Hippocratea aggregata Peyr. =Cuervea kappleriana 

Hippocratea ambigua Peyr. [insufficiently known taxon] 

Hippocratea aspera Lam. =Prionostemma aspera 

Hippocratea comosa Sw. =Hylenaea comosa 

Hippocratea crinita Pittier =Hylenaea comosa 

Hippocratea emarginata Rudge =Salacia multiflora subsp. multiflora 

Hippocratea flava Gleason =Cuervea kappleriana 

Hippocratea floribunda Benth. =Elachyptera floribunda 

Hippocratea granatensis Peyr. =Cuervea kappleriana 

Hippocratea holdeniana A.C. Sm. =Pristimera holdeniana 

Hippocratea huanucana Loes. =Peritassa huanucana 

Hippocratea inundata Peyr. =Cuervea kappleriana 

Hippocratea kappleriana Miq. =Cuervea kappleriana 

Hippocratea laevigata Rich. =Hippocratea volubilis 

Hippocratea lancifolia Lundell =Elachyptera floribunda 

Hippocratea lutea Gleason =Cuervea kappleriana 

Hippocratea malpighiifolia Rudge =Prionostemma aspera 

Hippocratea micrantha Cambess. =Elachyptera floribunda 

Hippocratea mitchellae I.M. Johnst. =Cuervea kappleriana 

Hippocratea multiflora Lam. =Salacia multiflora 

Hippocratea obovata Rich. =Salacia multiflora subsp. multiflora 

Hippocratea obovata var. guianense Peyr. =Salacia multiflora subsp. multiflora 

Hippocratea opacifolia J.F. Macbr. =Salacia opacifolia 

Hippocratea schomburgkii Klotzsch [nomen nudum] =Prionostemma aspera 

Hippocratea tenuiflora Mart. ex Peyr. =Pristimera tenuiflora 

Hippocratea verrucosa Kunth =Pristimera verrucosa 

Hylenaea capilliflora Miers =Hylenaea comosa 

Kippistia cognata Miers =Cheiloclinium cognatum 

Kippistia diffusiflora Miers =Cheiloclinium diffusiflorum 

Peritassa cuneata Mennega =Peritassa huanucana 

Peritassa granulata (Urb.) A.C. Sm. =Peritassa laevigata 

Peritassa laevigata subsp. obtusa Peyr. =Peritassa compta 

AM GU FG 

BO GU SU FG 

SU? 

AM SU FG 

GU SU FG 


BO GU 

GU SU FG? 



FG 

GU FG


Peritassa rorida Miers =Peritassa laevigata 

Peritassa rubicunda Miers =Peritassa laevigata 

Peritassa smaragdina Miers =Peritassa laevigata 

Prionostemma malpighiifolia (Rudge) Miers =Prionostemma aspera 

Pristimera apiculata Miers =Cuervea kappleriana 

Pristimera caribaea (Urb.) A.C. Sm. [not in flora area] 

Raddia elliptica (Mart. ex Schult.) Miers =Salacia elliptica 

Raddia grandiflora (Benth.) Miers =Salacia impressifolia 

Raddia impressifolia Miers =Salacia impressifolia 

Raddia lacunosa Miers =Salacia elliptica 

Raddia pachyphylla Miers =Salacia pachyphylla 

Salacia acreana A.C. Sm. =Salacia cordata 

Salacia affinis Peyr. =Cheiloclinium cognatum 

Salacia anomala (Miers) Peyr. =Cheiloclinium anomalum 

Salacia anomalum (Miers) Peyr. =Cheiloclinium anomalum 

Salacia attenuata (Miers) Peyr. =Tontelea attenuata 

Salacia belizensis Standl. =Cheiloclinium belizense 

Salacia brevistaminea Pittier =Anthodon decussatum 

Salacia catalinensis Rusby =Prionostemma aspera 

Salacia cognata (Miers) Peyr. =Cheiloclinium cognatum 

Salacia cognata var. egensis Peyr. =Cheiloclinium cognatum 

Salacia crassifolia Pittier =Salacia multiflora subsp. multiflora 

Salacia cylindrocarpa A.C. Sm. =Tontelea cylindrocarpa 

Salacia diffusiflora (Miers) Peyr. =Cheiloclinium diffusiflorum 

Salacia dulcis Benth. =Peritassa dulcis 

Salacia elliptica var. apiculata Peyr. =Salacia elliptica 

Salacia fluminensis Peyr. =Tontelea ovalifolia subsp. richardii 

Salacia gleasoniana A.C. Sm. =Cheiloclinium hippocrateoides 

Salacia gracilis A.C. Sm. =Salacia opacifolia 

Salacia grandiflora Peyr. =Salacia impressifolia 

Salacia granulata Urb. =Peritassa laevigata 

Salacia guianensis Klotzsch [nomen nudum] =Salacia elliptica 

Salacia guianensis (Aubl.) Lemée [incertae sedis; excluded from Hippocrateaceae by A.C. Smith] =Macahanea guianensis 

Aubl. 

Salacia hippocrateoides Peyr. =Cheiloclinium hippocrateoides 

Salacia krukovii A.C. Sm. =Cheiloclinium hippocrateoides 

Salacia lacunosa (Miers) Peyr. =Salacia elliptica 

Salacia laevigata (Hoffmanns. ex Link) DC. =Peritassa laevigata 

Salacia laevigata subsp. obtusa Peyr. =Peritassa compta 

Salacia laxiflora (Benth.) Peyr. =Tontelea laxiflora 

Salacia laxiflora var. amygdalifolia Peyr. =Tontelea laxiflora 

Salacia laxiflora var. oblongifolia Peyr. =Tontelea laxiflora 

Salacia lineolata A.C. Sm. =Cheiloclinium cognatum 

Salacia macrophylla Miq. =Salacia miqueliana 

Salacia megistophylla Standl. =Salacia cordata 

Salacia mucronata Rusby =Salacia multiflora subsp. mucronata 

Salacia nectandrifolia A.C. Sm. =Tontelea nectandrifolia 

Salacia obovata (Rich.) Peyr. =Salacia multiflora subsp. multiflora 

Salacia obovata var. guianensis Peyr. =Salacia multiflora subsp. multiflora 

Salacia parviflora Sagot ex Peyr. =Cheiloclinium diffusiflorum 

Salacia patens Triana & Planch. =Tontelea ovalifolia 

Salacia pittieriana A.C. Sm. =Salacia multiflora subsp. multiflora 

Salacia podostemma Sandwith =Cheiloclinium belizense 

Salacia pruinosa Seem. =Peritassa pruinosa 

Salacia richardii Peyr. =Tontelea ovalifolia subsp. richardii 

Salacia rugulosa Sagot [nomen nudum] =Salacia multiflora subsp. multiflora 

Salacia scabra DC. =Prionostemma aspera 

Salacia sphaerocarpa Rusby =Cheiloclinium cognatum 

Sicyomorpha nervosa Miers =Pristimera nervosa 

Sicyomorpha peruviana Miers =Peritassa peruviana 

Sicyomorpha pruinosa (Seem.) Miers =Peritassa pruinosa 

Thermophila cordata Miers =Salacia cordata 

Thermophila emarginata (Rudge) Miers =Salacia multiflora subsp. multiflora 

Thermophila macrophylla (Miq.) Miers =Salacia miqueliana

316 


Thermophila obovata (Rich.) Miers =Salacia multiflora subsp. multiflora 

Thermophila rugulosa Miers =Salacia multiflora subsp. multiflora 

Tonsella laevigata Hoffmanns. ex Link =Peritassa laevigata 

Tonsella malpighiifolia (Rudge) G. Mey. =Prionostemma aspera 

Tontelea aubletiana Miers =Tontelea sp. 

Tontelea floribunda (Benth.) Miers =Elachyptera floribunda 

Tontelea fluminensis (Peyr.) A.C. Sm. =Tontelea ovalifolia subsp. richardii 

Tontelea longifolia Miers =Tontelea attenuata 

Tontelea parviflora Miers =Cheiloclinium diffusiflorum 

Tontelea richardii (Peyr.) A.C. Sm. =Tontelea ovalifolia subsp. richardii 

Tontelea scandens Aubl. [nomen nudum] 

REFERENCES 

Görts-van Rijn, A. 2002. Hippocrateaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 347-353. 

Görts-van Rijn, A.R.A. & A.M.W. Mennega. 1994. Hippocrateaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 

16: 1-157. 

Loesener, T. 1942. Hippocrateaceae. In A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien 20b: 198-231. 

Mennega, A.M.W. & J.P. Hedin. 1999. Hippocrateaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Smith, A.C. 1940. The American species of Hippocrateaceae. Brittonia 3: 341-355. 

Hebepetalum humiriifolium (Planch.) Benth. 

Hebepetalum roraimense Secco & S.M.B. Silva 

Hebepetalum sp. A 

Roucheria calophylla Planch. 

Roucheria columbiana Hallier f. 

Roucheria laxiflora H. Winkl. 

Roucheria schomburgkii Planch. 

Roucheria sp. A 

HUGONIACEAE 

P.E. Berry, A. Jardim, N. Ramírez, D. Sabatier 


Hebepetalum parviflorum Ducke =Roucheria calophylla 

Hebepetalum punctatum Ducke =Roucheria columbiana 

Hebepetalum schomburgkii (Planch.) Ducke =Roucheria schomburgkii 

Roucheria angulata Gleason =Hebepetalum humiriifolium 

Roucheria humeriifolia Planch. =Hebepetalum humiriifolium 

Roucheria humiriifolia Planch. =Hebepetalum humiriifolium 

Roucheria parviflora (Ducke) Ducke =Roucheria calophylla 

Roucheria punctata (Ducke) Ducke =Roucheria columbiana 

REFERENCES 

Bakhuizen van den Brink f., R.C. 1941. Linaceae. In A. Pulle, Flora of Suriname 3(1): 409-411. 

Ramírez, N., P.E. Berry, & A. Jardim. 1999. Hugoniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Sabatier, D. 2002. Hugoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


353-354. 

Secco, R. de S. & S.M.B. Silva. 1990. Materiaes para a flora amazônica - VIII. Contribuçao à sistemática das Linaceae da 

Amazônia Brasileira. Bol. Mus. Paraense Emílio Goeldi (Bot.) 6(1): 113-135. 

Endopleura uchi (Huber) Cuatrec. 

Humiria balsamifera Aubl. 

Humiria balsamifera var. balsamifera 

Humiria balsamifera var. coriacea Cuatrec. 

Humiria balsamifera var. floribunda (Mart.) Cuatrec. 

Humiria balsamifera var. guaiquinimana Cuatrec. 

HUMIRIACEAE 

J. Cuatrecasas†, O. Huber, D. Sabatier 


AM GU 

AM 

AM GU SU FG? 

AM GU 

AM BO SU FG 

BO GU SU 

AM 

AM GU FG 



AM BO GU SU 


BO


Humiria balsamifera var. guianensis (Benth.) Cuatrec. 

Humiria balsamifera var. iluana Cuatrec. 

Humiria balsamifera var. imbaimadaiensis Cuatrec. 

Humiria balsamifera var. laurina (Urb.) Cuatrec. 

Humiria balsamifera var. pilosa (Steyerm.) Cuatrec. 

Humiria balsamifera var. savannarum (Gleason) Cuatrec. 

Humiria balsamifera var. stenocarpa Cuatrec. 

Humiria balsamifera var. subsessilis (Urb.) Cuatrec. 

Humiria crassifolia Mart. ex Urb. 

Humiria fruticosa Cuatrec. 

Humiria wurdackii Cuatrec. 

Humiriastrum colombianum Cuatrec. 

Humiriastrum cuspidatum (Benth.) Cuatrec. 

Humiriastrum excelsum (Ducke) Cuatrec. 

Humiriastrum liesneri Cuatrec. 

Humiriastrum obovatum (Benth.) Cuatrec. 

Humiriastrum ottohuberi Cuatrec. 

Humiriastrum piraparanense Cuatrec. 

Humiriastrum subcrenatum (Benth.) Cuatrec. 

Sacoglottis amazonica Mart. 

Sacoglottis ceratocarpa Ducke 

Sacoglottis cydonioides Cuatrec. 

Sacoglottis guianensis Benth. 

Sacoglottis guianensis var. guianensis 

Sacoglottis maguirei Cuatrec. 

Sacoglottis mattogrossensis Malme [Guianas specimens represent a new 

species, fide Sabatier] 

Schistostemon auyantepuiensis Cuatrec. 

Schistostemon densiflorum (Benth.) Cuatrec. 

Schistostemon dichotomum (Urb.) Cuatrec. 

Schistostemon fernandezii Cuatrec. 

Schistostemon oblongifolium (Benth.) Cuatrec. 

Schistostemon retusum (Ducke) Cuatrec. 

Schistostemon sylvaticum Sabatier 

Vantanea guianensis Aubl. 

Vantanea minor Benth. 

Vantanea ovicarpa Sabatier 

Vantanea parviflora Lam. 

Vantanea parviflora var. parviflora 


Houmiri balsamifera Aubl. =Humiria balsamifera 

Humiria balsamifera J. St.-Hil. =Humiria balsamifera Aubl. 

Humiria cassiquiari Suess. & Bergdolt =Humiria balsamifera var. guianensis 

Humiria densiflora Benth. =Schistostemon densiflorum 

Humiria floribunda (Mart.) Urb. =Humiria balsamifera var. floribunda 

Humiria floribunda var. guianensis Urb. =Humiria balsamifera var. guianensis 

Humiria floribunda var. laurina Urb. =Humiria balsamifera var. laurina 

Humiria floribunda var. spathulata Gleason =Humiria balsamifera var. laurina 

Humiria floribunda var. subsessilis Urb. =Humiria balsamifera var. subsessilis 

Humiria guianense Benth. =Humiria balsamifera var. guianensis 

Humiria pilosa Steyerm. =Humiria balsamifera var. pilosa 

Humiria savannarum Gleason =Humiria balsamifera var. savannarum 

Humiria subcrenata Urb. =Humiriastrum subcrenatum 

Humirium balsamiferum Benth. =Humiria balsamifera 

Humirium cuspidatum Benth. =Humiriastrum cuspidatum 

Humirium floribundum Mart. =Humiria balsamifera var. floribunda 

Humirium oblongifolium Benth. =Schistostemon oblongifolium 

Humirium obovatum Benth. =Humiriastrum obovatum 

Humirium surinamense Miq. =Humiria balsamifera var. guianensis 

Lemniscia floribunda Willd. =Vantanea guianensis 

Lemniscia guianensis (Aubl.) J.F. Gmel. =Vantanea guianensis 

Myrodendrum amplexicaule Willd. =Humiria balsamifera 

Myrodendrum balsamiferum Raeusch. [comb. illeg.] =Humiria balsamifera 

AM GU SU 

BO 

BO GU 

AM BO GU 

BO 

AM BO GU 

BO 

AM BO 

AM BO GU 

AM 

AM 

AM 

AM BO GU 

AM 

AM 

AM 

BO GU 

FG 

FG 

DA GU FG? 

AM BO 



GU SU FG 

AM 

AM BO GU 

BO 

BO 

GU SU 

GU? SU FG 

AM 

AM 

AM GU SU 

FG 

FG 

BO GU 

AM GU SU 

FG 

FG 

SU FG

318 


Sacoglottis amazonica Benth. =Sacoglottis guianensis 

Sacoglottis cuspidata (Benth.) Urb. =Humiriastrum cuspidatum 

Sacoglottis densiflora (Benth.) Urb. =Schistostemon densiflorum 

Sacoglottis dichotoma Urb. =Schistostemon dichotomum 

Sacoglottis excelsa var. colombiana Cuatrec. =Humiriastrum colombianum 

Sacoglottis guianensis f. dolichocarpa Ducke =Sacoglottis guianensis 

Sacoglottis guianensis var. hispidula Cuatrec. =Sacoglottis guianensis 

Sacoglottis guianensis var. major Ducke =Sacoglottis guianensis 

Sacoglottis guianensis f. sphaerocarpa Ducke =Sacoglottis mattogrossensis 

Sacoglottis kaboeriensis Bakh. v.d. Brink f. =Schistostemon densiflorum 

Sacoglottis oblongifolia (Benth.) Urb. =Schistostemon oblongifolium 

Sacoglottis obovata (Benth.) Urb. =Humiriastrum obovatum 

Sacoglottis retusa Ducke =Schistostemon retusum 

Sacoglottis uchi Huber =Endopleura uchi 

Vantanea cupularis Huber =Vantanea parviflora 

REFERENCES 

Bakhuizen van den Brink f., R.C. 1941. Humiriaceae. In A. Pulle, Flora of Suriname 3(1): 412-421. 

Cuatrecasas, J. 1961. A taxonomic revision of the Humiriaceae. Contrib. U.S. Nat. Herb. 35: 25-214. 

Cuatrecasas, J. & O. Huber. 1999. Humiriaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Sabatier, D. 1987. Studies on the Flora of the Guianas 12. Quelques nouveautés chez les Humiriaceae. Proc. Kon. Ned. Akad. 

Wet. ser. C, 90(2): 203-210. 

Sabatier, D. 2002. Humiriaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


355-357. 

Hydrolea elatior Schott 

Hydrolea palustris (Aubl.) Raeusch. [naturalized?] 

Hydrolea spinosa L. 

Hydrolea spinosa var. spinosa 


Hydrolea minima Brand =Hydrolea elatior 

Hydrolea multiflora Mart. ex Choisy =Hydrolea elatior 

Nama spinosa (L.) Kuntze =Hydrolea spinosa 

Sagonea palustris Aubl. =Hydrolea palustris 

HYDROPHYLLACEAE 

J.S. Miller 

REFERENCES 

Constance, L. 1982. Hydrophyllaceae. In Z. Luces de Febres, Flora de Venezuela 8(3): 7-14. 

Davenport, L.J. 1988. A monograph of Hydrolea. Rhodora 90 (862): 169-208. 

Miller, J.S. 1999. Hydrophyllaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 5: 644-646. 

ICACINACEAE 

A.C. de Roon, R. A. Howard, R.D. de Stefano 

Casimirella ampla (Miers) R.A. Howard 

Dendrobangia boliviana Rusby 

Discophora froesii Pires 

Discophora guianensis Miers 

Emmotum acuminatum (Benth.) Miers [Guianan records erroneous, but 

possibly expected] 

Emmotum celiae R.A. Howard 

Emmotum conjunctum R.A. Howard 

Emmotum fagifolium Ham. 

Emmotum floribundum R.A. Howard 

Emmotum fulvum R.A. Howard [Mt. Roraima] 

Emmotum glabrum Benth. ex Miers 

Emmotum orbiculatum (Benth.) Miers [Guianan records erroneous, but 

possibly expected] 

Emmotum yapacanum R.A. Howard 

AM BO DA GU 


FG 

FG 

GU SU FG 

AM FG 


AM 


AM 

AM 

AM 

BO GU 

GU SU FG 

BO GU? 

AM BO GU 

AM

Leretia cordata Vell. 

Pleurisanthes artocarpi Baill. 

Pleurisanthes emarginata Tiegh. 

Pleurisanthes flava Sandwith 

Pleurisanthes parviflora (Ducke) R.A. Howard 

Pleurisanthes sp. A 

Poraqueiba guianensis Aubl. 

Poraqueiba paraensis Ducke 

Poraqueiba sericea Tul. 



Asterolepidion elatum Ducke =Dendrobangia boliviana 

Clavapetalum elatum (Ducke) Ducke =Dendrobangia boliviana 

Clavapetalum surinamense Pulle =Dendrobangia boliviana 

Dendrobangia tenuis Ducke =Dendrobangia boliviana 

Discophora panamensis Standl. =Discophora guianensis 

Emmotum argenteum Gleason =Emmotum glabrum 

Emmotum nudum R.A. Howard =Emmotum acuminatum 

Emmotum ptarianum Steyerm. =Emmotum glabrum 

Humirianthera ampla (Miers) Baehni =Casimirella ampla 

Humirianthera duckei Huber =Casimirella ampla 

Kummeria brasiliensis Mart. =Discophora guianensis 

Leretia ampla Miers =Casimirella ampla 

Leretia nitida Miers =Leretia cordata 

Leretia parviflora Ducke =Pleurisanthes parviflora 

Leretia poeppigiana (Baill.) Sleumer =Leretia cordata 

Leretia vellozii Miers =Leretia cordata 

Mappia ampla (Miers) Engl. =Casimirella ampla 

Mappia cordata (Vell.) Engl. =Leretia cordata 

Mappia nitida (Miers) Engl. =Leretia cordata 

Mappia parviflora (Ducke) Baehni =Pleurisanthes parviflora 

Mappia poeppigiana Baill. =Leretia cordata 

Pleurisanthes brasiliensis Tiegh. [not in flora area] 

Pleurisanthes simpliciflora Sleumer [not in flora area] 

Pogopetalum acuminatum Benth. =Emmotum acuminatum 

Pogopetalum acutum Benth. =Emmotum fagifolium 

Pogopetalum orbiculatum Benth. =Emmotum orbiculatum 

Poraqueiba acuminata Miers =Poraqueiba sericea 

Poraqueiba surinamensis Miers =Poraqueiba guianensis 

REFERENCES 

Howard, R.A. 1942. Studies in the Icacinaceae, III. A revision of Emmotum. J. Arn. Arb. 23: 479-494. 

Howard, R.A. 1942. Studies of the Icacinaceae IV. Considerations of the New World genera. Contrib. Gray Herb. 142: 3-60. 

Howard, R.A. 1978. Icacinaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 10. Mem. New York Bot. 

Gard. 29: 62-68. 

Howard, R.A. 1992. A revision of Casimirella, including Humirianthera. Brittonia 44(2): 166-172. 

Howard, R.A. & R.D. de Stefano. 1999. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 5: 646-658. 

Jansen-Jacobs, M.J. 1979. Icacinaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 344-355. 

Roon, A.C. de. 1994. Icacinaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 16: 1-157. 

Roon, A.C. de. & S.A. Mori. 2002. Icacinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 


Gard. 76 (2): 358-362. 

Sleumer, H. 1942. Icacinaceae. In A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien 20b: 322-396. 

Cyrillopsis micrantha (Steyerm.) P.E. Berry & N. Ramírez 

Cyrillopsis paraensis Kuhlm. 

Ochthocosmus attenuatus Steyerm. & Luteyn 

Ochthocosmus barrae Hallier f. 

Ochthocosmus berryi Steyerm. 

Ochthocosmus floribundus Gleason 

Ochthocosmus longipedicellatus Steyerm. & Luteyn 

IXONANTHACEAE 

P.E. Berry, N. Ramírez, D. Sabatier 

DA GU SU FG 

GU FG 

AM SU FG 

GU 

GU SU FG 

AM 

GU SU FG 

AM BO 

AM 

BO 

AM GU FG 

BO GU 

GU? 

AM 

BO 

BO GU

320 


Ochthocosmus multiflorus Ducke 

Ochthocosmus multiflorus var. angustifolius Steyerm. & Luteyn 

Ochthocosmus multiflorus var. canaripoensis Steyerm. & Luteyn 

Ochthocosmus multiflorus var. multiflorus 

Ochthocosmus parvifolius Hallier f. [incertae sedis] 

Ochthocosmus roraimae Benth. 

Ochthocosmus roraimae var. grandifolius (Steyerm.) Steyerm. & Luteyn 

Ochthocosmus roraimae var. roraimae 


Ochthocosmus grandifolius Steyerm. =Ochthocosmus roraimae var. grandifolius 

Ochthocosmus micranthus Steyerm. =Cyrillopsis micrantha 

Ochthocosmus roraimae var. parvifolius (Hallier f.) Steyerm. & Luteyn =Ochthocosmus parvifolius 

REFERENCES 

Forman, L.L. 1965. A new genus of Ixonanthaceae with notes on the family. Kew Bull. 19: 517-526. 

Kuhlmann, J.G. 1925. Cyrillaceae, in Contribuiçao para o conhecimento de algumas plantas novas. Arq. Jard. Bot. Rio de 

Janeiro 4: 356-358. 

Ramírez, N. & P.E. Berry. 1999. Ixonanthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Steyermark, J.A. & J.L. Luteyn. 1980. Revision of the genus Ochthocosmus. Brittonia 32: 128-143. 

Secco, R. de S. & S.M.B. Silva. 1990. Materiaes para a flora amazônica - VIII. Contribuçao à sistemática das Linaceae da 

Amazônia Brasileira. Bol. Mus. Paraense Emílio Goeldi (Bot.) 6(1): 113-135. 

Juglans sp. [probably based on misidentifications] 

JUGLANDACEAE 


Juglans brasiliensis Dode [not in flora area] =Juglans australis Griseb. 

REFERENCES 

Manning, W.E. 1960. The genus Juglans in South America and the West Indies. Brittonia 12: 1-26. 

Krameria ixine Loefl. 

Krameria spartioides Klotzsch ex O. Berg 

Krameria tomentosa A. St.-Hil. 

KRAMERIACEAE 

L. Brako, B.B. Simpson 


Krameria arida O. Berg =Krameria ixine 

Krameria evolvuloides Triana & Planch. ex Bennett =Krameria spartioides 

Krameria ixina L. =Krameria ixine 

Krameria lanceolata O. Berg [nom. illeg.] =Krameria ixine 

Krameria linifolia Willd. ex Schult. & Schult. f. =Krameria ixine 

REFERENCES 

Brako, L. 1999. Krameriaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 674-676. 

Simpson, B.B. 1989. Krameriaceae. Flora Neotropica Monogr. 49: 1-108. 

Simpson, B.B. 1998. Krameriaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 21: 61-65. 

Lacistema aggregatum (P.J. Bergius) Rusby 

Lacistema grandifolium W. Schnizl. 

Lacistema polystachyum W. Schnizl. 

LACISTEMATACEAE 

L.J. Raechal 


Lacistema aggregatum var. elongatum Maguire =Lacistema aggregatum 

Lacistema coriaceum A. DC. =Lacistema aggregatum 

AM 

AM 

AM 

AM 

GU? 

BO GU 

BO 

BO GU 

BO GU 

BO GU 

GU? 

FG 


GU SU FG 

GU SU FG

Lacistema elongatum Schnizl. =Lacistema aggregatum 

Lacistema orinocense Baehni =Lacistema aggregatum 


REFERENCES 

Mori, S.A. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide 


Raechal, L.J. 1999. Lacistemaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 5: 676-677. 

Sleumer, H.O. 1980. Flacourtiaceae. Flora Neotropica Monogr. 22: 1-499. 

Vaandrager, G. 1938. Lacistemaceae. In A. Pulle, Flora of Suriname 1(1): 258-261. 

LAMIACEAE 

R.M. Harley 

Anisomeles indica (L.) Kuntze [South Asian exotic] 

Eriope crassipes Benth. 

Eriope crassipes subsp. crassipes 

Hypenia salzmannii (Benth.) Harley 

Hyptidendron arboreum (Benth.) Harley 

Hyptis atrorubens Poit. 

Hyptis brachiata Briq. 

Hyptis brevipes Poit. [frequently confused with H. lanceolata, fide Harley] 

Hyptis capitata Jacq. 

Hyptis colubrimontis Epling & Játiva [=Hyptis pyriformis var. yutajeensis] 

Hyptis conferta Pohl ex Benth. 

Hyptis conferta var. angustata (Briq.) Pool & Harley 

Hyptis dilatata Benth. 

Hyptis guanchezii Harley 

Hyptis hirsuta Kunth 

Hyptis huberi Harley 

Hyptis laciniata Benth. 

Hyptis lacustris A. St.-Hil. ex Benth. 

Hyptis lanceolata Poir. 

Hyptis lantanifolia Poit. 

Hyptis luticola Epling 

Hyptis microphylla Pohl ex Benth. 

Hyptis mutabilis (Rich.) Briq. 

Hyptis pachycephala Epling 

Hyptis parkeri Benth. 

Hyptis pectinata (L.) Poit. 

Hyptis pulegioides Pohl ex Benth. 

Hyptis pyriformis Epling & Játiva 

Hyptis pyriformis var. pyriformis 

Hyptis pyriformis var. yutajeensis Harley 

Hyptis recurvata Poit. 

Hyptis spicigera Lam. 

Hyptis suaveolens (L.) Poit. 

Leonotis nepetifolia (L.) W.T. Aiton 

Leonurus japonicus Houtt. [naturalized] 

Marsypianthes chamaedrys (Vahl) Kuntze 

Ocimum americanum L. 

Ocimum campechianum Mill. 

Ocimum gratissimum L. [naturalized] 

Scutellaria leucantha Loes. 

Scutellaria purpurascens Sw. 

Scutellaria uliginosa A. St.-Hil. ex Benth. 

Ballota suaveolens L. =Hyptis suaveolens 

Clinopodium chamaedrys Vahl =Marsypianthes chamaedrys 

Eriope nudiflora Kunth ex Benth. =Eriope crassipes 

Hyptis americana (Aubl.) Urb. =Hyptis spicigera 

Hyptis arborea Benth. =Hyptidendron arboreum 

Hyptis belizensis Lundell =Hyptis conferta var. angustata 


GU 

AM FG 

AM FG 

BO GU 

BO GU 


AM BO 

BO DA SU FG 

BO FG? 

AM 

AM BO 

AM BO 

AM BO GU 

AM 

BO GU SU 

AM 

AM BO GU 

AM BO GU 



AM 

BO GU SU 

AM BO SU FG 

FG 


BO DA GU SU? FG 

GU 

AM BO 

AM BO 

AM 


GU FG 


DA GU SU FG 

DA GU SU FG 


GU SU 


SU 

FG 

BO 

FG

322 


Hyptis brevipes var. serrata Briq. =Hyptis brevipes 

Hyptis canescens Kunth =Hyptis mutabilis 

Hyptis conferta var. angustifolia Benth. ex Epling [nom. illeg.] =Hyptis conferta var. angustata 

Hyptis constricta Briq. =Hyptis conferta var. angustata 

Hyptis crenata var. angustifolia Benth. ex Seem. =Hyptis dilatata 

Hyptis excelsa M. Martens & Galeotti =Hyptis conferta var. angustata 

Hyptis ferruginea Benth. =Hyptis conferta var. angustata 

Hyptis jurgensenii Briq. =Hyptis conferta var. angustata 

Hyptis jurgensenii var. angustata Briq. =Hyptis conferta var. angustata 

Hyptis lanicephala Epling & Játiva =Hyptis dilatata 

Hyptis lundii Benth. =Hyptis conferta var. angustata 

Hyptis micrantha Pohl ex Benth. =Hyptis mutabilis 

Hyptis mutabilis var. canescens (Kunth) Briq. =Hyptis mutabilis 

Hyptis parkeri var. verbenifolia (J.A. Schmidt ex Mart.) Epling =Hyptis parkeri 

Hyptis polystachya Kunth =Hyptis mutabilis 

Hyptis radiata Willd. [not in flora area] =Hyptis alata (Raf.) Shinners 

Hyptis recurvata var. megacephala Benth. =Hyptis recurvata 

Hyptis salzmannii Benth. =Hypenia salzmannii 

Hyptis spicata Poit. =Hyptis mutabilis 

Hyptis verbenifolia J.A. Schmidt ex Mart. =Hyptis parkeri 

Leonurus heterophyllus Sweet =Leonurus japonicus 

Leonurus sibiricus auct. non L. 1753 =Leonurus japonicus 

Marsypianthes hyptoides Mart. ex Benth. =Marsypianthes chamaedrys 

Mesosphaerum atrorubens (Poit.) Kuntze =Hyptis atrorubens 

Mesosphaerum brevipes (Poit.) Kuntze =Hyptis brevipes 

Mesosphaerum capitatum (Jacq.) Kuntze =Hyptis capitata 

Mesosphaerum confertum (Pohl ex Benth.) Kuntze =Hyptis conferta 

Mesosphaerum dilatatum (Benth.) Kuntze =Hyptis dilatata 

Mesosphaerum excelsum (M. Martens & Galeotti) Kuntze =Hyptis conferta var. angustata 

Mesosphaerum hirsutum (Kunth) Kuntze =Hyptis hirsuta 

Mesosphaerum laciniatum (Benth.) Kuntze =Hyptis laciniata 

Mesosphaerum lacustre (St.-Hil. ex Benth.) Kuntze =Hyptis lacustris 

Mesosphaerum lanceolatum (Poir.) Kuntze =Hyptis lanceolata 

Mesosphaerum lantanifolium (Poit.) Kuntze =Hyptis lantanifolia 

Mesosphaerum microphyllum (Pohl ex Benth.) Kuntze =Hyptis microphylla 

Mesosphaerum mutabile (Rich.) Kuntze =Hyptis mutabilis 

Mesosphaerum parkeri (Benth.) Kuntze =Hyptis parkeri 

Mesosphaerum pectinatum (L.) Kuntze =Hyptis pectinata 

Mesosphaerum recurvatum (Poit.) Kuntze =Hyptis recurvata 

Mesosphaerum suaveolens (L.) Kuntze =Hyptis suaveolens 

Nepeta mutabilis Rich. =Hyptis mutabilis 

Nepeta pectinata L. =Hyptis pectinata 

Ocimum micranthum Willd. =Ocimum campechianum 

Phlomis nepetifolia L. =Leonotis nepetifolia 

REFERENCES 

Epling, C. 1935-1937. Synopsis of the South American Labiatae. Repert. Sp. Nov. Beih. 85: 1-341. 

Epling, C. 1949. Revisión del género Hyptis (Labiatae). Rev. Mus. La Plata (Sect. Bot.) 7: 153-497. 

Harley, R.M. 1988. Revision of generic limits in Hyptis Jacq. (Labiatae) and its allies. Bot. J. Linn. Soc. 98: 87-95. 

Harley, R.M. 1999. Lamiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 678-700. 

Harley, R.M. 1999. Lamiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


363-369. 

Kostermans, A.J.G.H. 1936. Labiatae. In A. Pulle, Flora of Suriname 4(1): 334-353. 

Kostermans, A.J.G.H. 1937. Labiatae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 498. 

Aiouea benthamiana Mez 

Aiouea guianensis Aubl. 

Aiouea impressa (Meisn.) Kosterm. 

Aiouea laevis (Mart.) Kosterm. 

LAURACEAE 

J.G. Rohwer, H. van der Werff 

AM BO 

BO GU SU FG 

FG 

BO GU SU FG


Aiouea longipetiolata van der Werff 

Aiouea maguireana (C.K. Allen) S.S. Renner 

Aiouea myristicoides Mez 

Aiouea opaca van der Werff 

Aniba affinis (Meisn.) Mez 

Aniba burchellii Kosterm. 

Aniba canelilla (Kunth) Mez 

Aniba cinnamomiflora C.K. Allen 

Aniba citrifolia (Nees) Mez 

Aniba cylindriflora Kosterm. 

Aniba excelsa Kosterm. 

Aniba ferruginea Kubitzki 

Aniba guianensis Aubl. 

Aniba hostmanniana (Nees) Mez 

Aniba hypoglauca Sandwith 

Aniba jenmanii Mez 

Aniba kappleri Mez 

Aniba megaphylla Mez 

Aniba panurensis (Meisn.) Mez 

Aniba parviflora (Meisn.) Mez 

Aniba percoriacea C.K. Allen 

Aniba permollis (Nees) Mez 

Aniba riparia (Nees) Mez 

Aniba rosaeodora Ducke 

Aniba taubertiana Mez 

Aniba terminalis Ducke 

Aniba williamsii O.C. Schmidt 

Beilschmiedia curviramea (Meisn.) Kosterm. 

Beilschmiedia hexanthera van der Werff 

Cassytha filiformis L. 

Chlorocardium rodiei (R.H. Schomb.) Rohwer, H.G. Richt. & van der Werff 

Cinnamomum cinnamomifolium (Kunth) Kosterm. [recorded from FG in error, 

but perhaps expected in GU] 

Cinnamomum semecarpifolium (Meisn.) Kosterm. 

Cinnamomum triplinerve (Ruiz & Pav.) Kosterm. 

Cryptocarya guianensis Meisn. 

Cryptocarya maroniensis Benoist 

Dicypellium caryophyllaceum (Mart.) Nees 

Endlicheria anomala (Nees) Mez 

Endlicheria arunciflora (Meisn.) Mez 

Endlicheria bracteolata (Meisn.) C.K. Allen 

Endlicheria canescens Chanderbali 

Endlicheria chalisea Chanderbali 

Endlicheria dictifarinosa C.K. Allen 

Endlicheria directonervia C.K. Allen 

Endlicheria gracilis Kosterm. 

Endlicheria macrophylla (Meisn.) Mez 

Endlicheria multiflora (Miq.) Mez 

Endlicheria nilssonii C.K. Allen 

Endlicheria paradoxa Mez 

Endlicheria punctulata (Mez) C.K. Allen 

Endlicheria pyriformis (Nees) Mez 

Endlicheria reflectens (Nees) Mez 

Endlicheria rubriflora Mez 

Endlicheria sericea Nees 

Endlicheria aff. szyszylowiczii Mez 

Endlicheria szyszylowiczii Mez 

Endlicheria vinotincta C.K. Allen 

Kubitzkia macrantha (Kosterm.) van der Werff 

Kubitzkia mezii (Kosterm.) van der Werff 

Licaria armeniaca (Nees) Kosterm. 

Licaria aurea (Huber) Kosterm. 

Licaria aureosericea van der Werff 

Licaria brasiliensis (Nees) Kosterm. 

AM 

BO 

GU FG 

FG 

AM GU 

AM 

AM BO GU SU? 

AM 


GU 

BO DA GU 

AM 

AM GU SU FG 

AM GU SU FG 

GU SU 

GU SU FG 

DA GU SU FG 



SU FG 

SU 

GU 

GU SU 

AM GU SU FG 


FG 

AM GU? SU FG 

BO DA GU 

FG 


GU SU FG 

AM 

DA FG 

FG 

FG 

FG? 


AM 


BO GU SU 

AM GU FG 

AM BO 

AM 


AM 


BO 

SU? 

GU SU FG 

GU SU FG 

BO GU FG 

AM FG 

SU FG 

AM 

GU? 

AM 

BO 

GU SU FG 

AM GU 

GU SU 

GU 

AM BO SU?

324 


Licaria camara (R.H. Schomb.) Kosterm. 

Licaria cannella (Meisn.) Kosterm. 

Licaria cannella subsp. tenuicarpa (Kosterm. ex Rodrigues) Kurz 

Licaria chrysophylla (Meisn.) Kosterm. 

Licaria debilis (Mez) Kosterm. 

Licaria dolichantha Kurz 

Licaria guianensis Aubl. 

Licaria macrophylla (A.C. Sm.) Kosterm. 

Licaria martiniana (Mez) Kosterm. 

Licaria multiflora (Kosterm.) Kosterm. 

Licaria oppositifolia (Nees) Kosterm. 

Licaria pachycarpa (Meisn.) Kosterm. 

Licaria polyphylla (Nees) Kosterm. 

Licaria rufotomentosa van der Werff 

Licaria subbullata Kosterm. 

Licaria tomentosa van der Werff 

Licaria trinervis van der Werff 

Licaria vernicosa (Mez) Kosterm. 

Mezilaurus itauba (Meisn.) Taub. ex Mez 

Mezilaurus lindaviana Schwacke & Mez 

Mezilaurus sprucei (Meisn.) Taub. ex Mez 

Mezilaurus thoroflora van der Werff 

Nectandra amazonum Nees 

Nectandra aurea Rohwer 

Nectandra cissiflora Nees 

Nectandra cuspidata Nees 

Nectandra fulva Rohwer 

Nectandra globosa (Aubl.) Mez 

Nectandra hihua (Ruiz & Pav.) Rohwer 

Nectandra lineata (Kunth) Rohwer 

Nectandra martinicensis Mez 

Nectandra paucinervia Coe-Teix. 

Nectandra pearcei Mez 

Nectandra pichurim (Kunth) Mez 

Nectandra pulverulenta Nees 

Nectandra purpurea (Ruiz & Pav.) Mez 

Nectandra reticulata (Ruiz & Pav.) Mez 

Nectandra ruforamula Rohwer 

Nectandra sanguinea Rol. ex Rottb. 

Ocotea aciphylla (Nees & Mart. ex Nees) Mez 

Ocotea adenotrachelium (Nees) Mez 

Ocotea aff. amazonica (Meisn.) Mez [name of uncertain application] 

Ocotea argyrophylla Ducke 

Ocotea atrata C.K. Allen 

Ocotea basirecurva C.K. Allen 

Ocotea bijuga (Rottb.) Bernardi [=Ocotea neesiana?] 

Ocotea bofo Kunth 

Ocotea bracteosa (Meisn.) Mez 

Ocotea canaliculata (Rich.) Mez 

Ocotea caracasana (Nees) Mez 

Ocotea ceanothifolia (Nees) Mez 

Ocotea cernua (Nees) Mez [s.l.] 

Ocotea cinerea van der Werff 

Ocotea commutata (Nees) Mez 

Ocotea cowaniana C.K. Allen 

Ocotea crassifolia (Nees) Mez [Mt. Roraima] 

Ocotea cujumary Mart. 

Ocotea cymbarum Kunth 

Ocotea debilis Mez 

Ocotea depauperata C.K. Allen 

Ocotea diffusa van der Werff 

Ocotea discrepens C.K. Allen 

Ocotea divaricata (Nees) Mez 

Ocotea duidensis Moldenke 

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Ocotea endlicheriopsis Mez 

Ocotea erectifolia (C.K. Allen) van der Werff 

Ocotea esmeraldana Moldenke 

Ocotea fasciculata (Nees) Mez 

Ocotea fendleri (Meisn.) Rohwer 

Ocotea flavantha van der Werff 

Ocotea floribunda (Sw.) Mez 

Ocotea glabra van der Werff 

Ocotea glaucophylla Moldenke 

Ocotea glomerata (Nees) Mez 

Ocotea gracilis (Meisn.) Mez 

Ocotea guianensis Aubl. 

Ocotea guianensis var. guianensis 

Ocotea guianensis var. subsericea Kosterm. 

Ocotea hemsleyana Mez 

Ocotea huberi van der Werff 

Ocotea indirectinervia C.K. Allen 

Ocotea javitensis (Kunth) Pittier 

Ocotea julianii van der Werff 

Ocotea laticostata C.K. Allen 

Ocotea leucoxylon (Sw.) Laness. 

Ocotea liesneri van der Werff 

Ocotea longifolia Kunth 

Ocotea megacarpa van der Werff 

Ocotea mucronata (Poir.) Kosterm. 

Ocotea myriantha (Meisn.) Mez 

Ocotea neblinae C.K. Allen 

Ocotea neesiana (Miq.) Kosterm. [see also Ocotea bijuga] 

Ocotea nigra Benoist 

Ocotea nilssonii C.K. Allen 

Ocotea oblonga (Meisn.) Mez 

Ocotea pauciflora (Nees) Mez 

Ocotea percurrens Vicentini 

Ocotea perrobusta (C.K. Allen) Rohwer 

Ocotea petalanthera (Meisn.) Mez 

Ocotea puberula (Rich.) Nees 

Ocotea revoluta Moldenke 

Ocotea rhodophylla Vicentini 

Ocotea roseopedunculata van der Werff 

Ocotea rubrinervis Mez 

Ocotea rufovestita Ducke 

Ocotea sanariapensis Lasser 

Ocotea sandwithii Kosterm. 

Ocotea scabrella van der Werff 

Ocotea scandens Kosterm. 

Ocotea schomburgkiana (Nees) Mez 

Ocotea sp. A 

Ocotea splendens (Meisn.) Baill. 

Ocotea subterminalis van der Werff 

Ocotea tillettsiana C.K. Allen 

Ocotea tomentella Sandwith 

Ocotea tomentosa van der Werff 

Ocotea venosa Gleason 

Ocotea wurdackiana C.K. Allen 

Ocotea yutajensis C.K. Allen 

Persea americana Mill. [cultivated, occasionally escaping] 

Persea areolatocostae (C.K. Allen) van der Werff 

Persea caerulea (Ruiz & Pav.) Mez 

Persea croatii van der Werff 

Persea croizatii van der Werff 

Persea fastigiata Kopp 

Persea fluviatilis van der Werff 

Persea grandiflora Kopp 

Persea jenmanii Mez 


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326 


Persea julianae van der Werff 

Persea maguirei Kopp 

Persea nivea Mez 

Persea perseiphylla (C.K. Allen) van der Werff 

Persea pseudofasciculata Kopp 

Persea steyermarkii C.K. Allen 

Pleurothyrium amapaense C.K. Allen 

Rhodostemonodaphne celiana (C.K. Allen) Rohwer 

Rhodostemonodaphne crenaticupula MadriZán 

Rhodostemonodaphne elephantopus MadriZán 

Rhodostemonodaphne kunthiana (Nees) Rohwer 

Rhodostemonodaphne leptoclada MadriZán 

Rhodostemonodaphne miranda (Sandwith) Rohwer 

Rhodostemonodaphne mirecolorata (C.K. Allen) Rohwer 

Rhodostemonodaphne morii MadriZán 

Rhodostemonodaphne praeclara (Sandwith) MadriZán 

Rhodostemonodaphne revolutifolia MadriZán 

Rhodostemonodaphne rufovirgata MadriZán 

Rhodostemonodaphne saülensis MadriZán 

Rhodostemonodaphne scandens MadriZán 

Rhodostemonodaphne steyermarkiana (C.K. Allen) van der Werff 

Sextonia rubra (Mez) van der Werff 


Acrodiclidium anacardioides Meisn. =Mezilaurus itauba 

Acrodiclidium aubletii Kosterm. =Licaria guianensis 

Acrodiclidium brasiliense Nees =Licaria brasiliensis 

Acrodiclidium canella (Meisn.) Mez =Licaria cannella 

Acrodiclidium cayennense (Meisn.) Mez =Licaria cannella 

Acrodiclidium chrysophyllum Meisn. =Licaria chrysophylla 

Acrodiclidium debile Mez =Licaria debilis 

Acrodiclidium guianense Nees =Licaria polyphylla 

Acrodiclidium guianense var. caudatum Meisn. =Licaria polyphylla 

Acrodiclidium itauba Meisn. =Mezilaurus itauba 

Acrodiclidium kunthiana Nees =Rhodostemonodaphne kunthiana 

Acrodiclidium macrophyllum A.C. Sm. =Licaria macrophylla 

Acrodiclidium martinianum Mez =Licaria martiniana 

Acrodiclidium meissneri Mez =Licaria polyphylla 

Acrodiclidium oppositifolium Nees =Licaria oppositifolia 

Acrodiclidium rigidum Kosterm. =Licaria chrysophylla 

Acrodiclidium sprucei Meisn. =Mezilaurus sprucei 

Aiouea demerarensis Kosterm. =Aiouea guianensis 

Aiouea densiflora Nees =Aiouea laevis 

Aiouea pernitida C.K. Allen =Aiouea myristicoides 

Aiouea perrobusta C.K. Allen =Ocotea perrobusta 

Aiouea schomburgkii Meisn. =Aiouea guianensis 

Ampelodaphne arunciflora Meisn. =Endlicheria arunciflora 

Ampelodaphne dasyantha Meisn. =Endlicheria multiflora 

Ampelodaphne macrophylla Meisn. =Endlicheria macrophylla 

Aniba castanea C.K. Allen =Aniba jenmaniI 

Aniba duckei Kosterm. =Aniba rosaeodora 

Aniba firmula (Nees & Mart.) Mez [not in flora area] 

Aniba gonggrijpii Kosterm. =Aniba panurensis 

Aniba koumaroucapa Kosterm. =Aniba megaphylla 

Aniba mas Kosterm. =Aniba panurensis 

Aniba muca (Ruiz & Pav.) Mez [not in flora area] 

Aniba muelleriana Mez =Aniba citrifolia 

Aniba murcana C.K. Allen =Aniba williamsii 

Aniba ovalifolia Mez =Aniba hypoglauca 

Aniba pittieri O.C. Schmidt =Aniba hostmanniana 

Aniba riparia auct. non (Nees) Mez.: sensu C.K. Allen, Mem. New York Bot. Gar =Aniba kappleri 

Aniba salicifolia (Nees) Mez =Aniba guianensis 

Aniba sericea (Nees) Brooks =Endlicheria sericea 

Aniba simulans C.K. Allen =Aniba taubertiana 

Aniba trinitatis (Meisn.) Mez =Aniba citrifolia 

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Aydendron affine Meisn. =Aniba affinis 

Aydendron cannella Meisn. =Licaria cannella 

Aydendron citrifolium Nees =Aniba citrifolia 

Aydendron curvirameum Meisn. =Beilschmiedia curviramea 

Aydendron hostmannianum Nees =Aniba hostmanniana 

Aydendron panurense Meisn. =Aniba panurensis 

Aydendron trinitatis Meisn. =Aniba citrifolia 

Cassytha americana Nees =Cassytha filiformis 

Cassytha baccifera Sol. ex J.S. Muell. (Cactaceae) =Rhipsalis baccifera 

Cinnamomum areolatocostae (C.K. Allen) Kosterm. =Persea areolatocostae 

Cinnamomum filamentosum (C.K. Allen) Kosterm. =Cinnamomum triplinerve 

Clinostemon maguireanum (C.K. Allen) Kurz =Mezilaurus thoroflora 

Cryptocarya canelilla Kunth =Aniba canelilla 

Cryptocarya laevis Mart. =Aiouea laevis 

Cryptocarya moschata Mart. [not in flora area] 

Cryptocarya mucronata (Poir.) Spreng. =Ocotea mucronata 

Cryptocarya pachycarpa Gleason =Aniba citrifolia 

Endlicheria endlicheriopsis (Mez) Kosterm. =Ocotea endlicheriopsis 

Endlicheria grandis Mez =Rhodostemonodaphne praeclara 

Endlicheria impressa Mez =Aiouea impressa 

Endlicheria levelii C.K. Allen =Endlicheria multiflora 

Endlicheria maguireana C.K. Allen =Aiouea maguireana 

Endlicheria melinonii Benoist =Endlicheria multiflora 

Endlicheria wurdackiana C.K. Allen =Endlicheria rubriflora 

Evonymodaphne armeniaca Nees =Licaria armeniaca 

Goeppertia anomala Nees =Endlicheria anomala 

Goeppertia multiflora Miq. =Endlicheria multiflora 

Goeppertia reflectens Nees =Endlicheria reflectens 

Goeppertia sericea var. bracteolata Meisn. =Endlicheria bracteolata 

Gymnobalanus fendleri Meisn. =Ocotea fendleri 

Laurus caerulea Ruiz & Pav. =Persea caerulea 

Laurus canaliculata Rich. =Ocotea canaliculata 

Laurus floribunda Sw. =Ocotea floribunda 

Laurus globosa Aubl. =Nectandra globosa 

Laurus hihua Ruiz & Pav. =Nectandra hihua 

Laurus javitensis Kunth =Ocotea javitensis 

Laurus leucoxylon Sw. =Ocotea leucoxylon 

Laurus puberula Rich. =Ocotea puberula 

Laurus reticulata Ruiz & Pav. =Nectandra reticulata 

Laurus triplinervis Ruiz & Pav. =Cinnamomum triplinerve 

Licaria appelii (Mez) Kosterm. =Licaria guianensis 

Licaria cayennensis (Meisn.) Kosterm. =Licaria cannella 

Licaria cymbarum (Kunth) Pittier =Ocotea cymbarum 

Licaria foveolata (Mez) Lemée =Licaria martiniana 

Licaria maguireana C.K. Allen =Mezilaurus thoroflora 

Licaria martiniana auct. non (Mez) Kosterm.: sensu C.K. Allen, Mem. New York Bot. G =Licaria oppositifolia 

Licaria meissneri (Mez) Kosterm. =Licaria polyphylla 

Licaria rigida (Kosterm.) Kosterm. =Licaria chrysophylla 

Licaria schultesii C.K. Allen =Licaria macrophylla 

Licaria simulans C.K. Allen =Licaria vernicosa 

Licaria wilhelminensis C.K. Allen =Licaria subbullata 

Mespilodaphne ceanothifolia Nees =Ocotea ceanothifolia 

Mespilodaphne myriantha Meisn. =Ocotea myriantha 

Mespilodaphne oblonga Meisn. =Ocotea oblonga 

Mezilaurus anacardioides (Meisn.) Taub. ex Mez =Mezilaurus itauba 

Mezilaurus maguireana C.K. Allen =Mezilaurus sprucei 

Mezilaurus wurdackiana C.K. Allen =Mezilaurus lindaviana 

Misanteca maguireana (Allen) Lundell =Mezilaurus thoroflora 

Nectandra ambigua Meisn. =Nectandra amazonum 

Nectandra angustifolia (Schrad.) Nees [not in flora area] 

Nectandra barcellensis Meisn. =Ocotea cymbarum 

Nectandra bredemeyeriana Nees =Nectandra hihua 

Nectandra canescens Nees [not in flora area] 

Nectandra cymbarum (Kunth) Nees =Ocotea cymbarum

328 


Nectandra glandulifolia Lasser =Nectandra martinicensis 

Nectandra grandis (Mez) Kosterm. =Rhodostemonodaphne praeclara 

Nectandra guianensis Meisn. =Nectandra sanguinea 

Nectandra kaburiensis Kosterm. =Nectandra pearcei 

Nectandra kunthiana (Nees) Kosterm. =Rhodostemonodaphne kunthiana 

Nectandra laurel Nees [not in flora area] 

Nectandra leucantha var. attenuata Meisn. =Nectandra hihua 

Nectandra leucantha var. guianensis Meisn. =Nectandra hihua 

Nectandra lucida Nees (1833) [not in flora area] =Endlicheria paniculata (Spreng.) Macbr. 

Nectandra lucida Nees (1836) =Nectandra hihua 

Nectandra magnoliifolia Meisn. [not in flora area] =Nectandra canescens Nees 

Nectandra maguireana C.K. Allen [not in flora area] 

Nectandra meyeriana Lasser =Rhodostemonodaphne kunthiana 

Nectandra miranda Sandwith =Rhodostemonodaphne miranda 

Nectandra mollis (Kunth) Nees =Nectandra reticulata 

Nectandra neesiana Miq. =Ocotea neesiana 

Nectandra olivacea Lasser =Nectandra cuspidata 

Nectandra pallida Nees =Nectandra amazonum 

Nectandra pisi Miq. =Nectandra globosa 

Nectandra pittieri Lasser =Nectandra reticulata 

Nectandra polyphylla Nees =Licaria polyphylla 

Nectandra praeclara Sandwith =Rhodostemonodaphne praeclara 

Nectandra rhynchophylla Meisn. =Ocotea aciphylla 

Nectandra rubra (Mez) C.K. Allen =Sextonia rubra 

Nectandra schomburgkii Meisn. =Nectandra hihua 

Nectandra spumea Kubitzki =Ocotea esmeraldana 

Nectandra surinamensis Mez =Nectandra cissiflora 

Nectandra vaga Meisn. =Nectandra globosa 

Nectandra viburnoides Meisn. [not in flora area] 

Nectandra willdenoviana Nees =Nectandra hihua 

Ocotea abbreviata Schwacke & Mez [not in flora area] 

Ocotea aciphylla var. chimantaensis C.K. Allen =Ocotea aciphylla 

Ocotea acutangula (Miq.) Mez =Ocotea leucoxylon 

Ocotea barcellensis (Meisn.) Mez =Ocotea cymbarum 

Ocotea budowskiana Bernardi =Ocotea erectifolia 

Ocotea castanea C.K. Allen =Ocotea schomburgkiana 

Ocotea caudata (Nees) Mez =Ocotea cernua 

Ocotea celiana C.K. Allen =Rhodostemonodaphne celiana 

Ocotea costulata (Nees) Mez =Ocotea aciphylla 

Ocotea cuprea (Meisn.) Mez =Ocotea oblonga 

Ocotea directoramea C.K. Allen =Ocotea debilis 

Ocotea dissimilis C.K. Allen =Ocotea splendens 

Ocotea duotincta C.K. Allen =Ocotea leucoxylon 

Ocotea fallax (Miq.) Mez =Ocotea neesiana 

Ocotea ferruginea auct. non (Meisn.) Mez 1889: sensu Bernardi, Candollea 30: 261. =Rhodostemonodaphne steyermarkiana 

Ocotea finium C.K. Allen =Ocotea aciphylla 

Ocotea firmula Mart. [nomen nudum][not in flora area] =Aniba firmula (Nees) Mez 

Ocotea florulenta (Meisn.) Mez =Ocotea neesiana 

Ocotea fulvifolia C.K. Allen =Ocotea aciphylla 

Ocotea glandulosa Lasser =Ocotea fendleri 

Ocotea glaucina (Meisn.) Mez [not in flora area] =Ocotea notata (Nees) Mez 

Ocotea globifera Mez =Ocotea splendens 

Ocotea indecora (Schott) Mez [not in flora area] 

Ocotea jenmanii Mez [nomen nudum] 

Ocotea kunthiana (Nees) Mez =Rhodostemonodaphne kunthiana 

Ocotea lasseriana C.K. Allen =Ocotea leucoxylon 

Ocotea laxiflora (Meisn.) Mez =Ocotea pauciflora 

Ocotea lineata Kunth =Nectandra lineata 

Ocotea maguireana C.K. Allen =Ocotea aciphylla 

Ocotea marowynensis (Miq.) Mez =Ocotea cernua 

Ocotea martiniana (Nees) Mez =Ocotea puberula 

Ocotea mirecolorata C.K. Allen =Rhodostemonodaphne mirecolorata 

Ocotea opifera Mart. =Ocotea longifolia 

Ocotea pendula (Poepp.) Engl. [nomen nudum]


Ocotea perseiphylla C.K. Allen =Persea perseiphylla 

Ocotea persulcata C.K. Allen =Ocotea leucoxylon 

Ocotea pichurim Kunth =Nectandra pichurim 

Ocotea pubescens (Nees) Mez [not in flora area] 

Ocotea punctulata Mez =Endlicheria punctulata 

Ocotea punctulata auct. non Mez. 1889: sensu C.K. Allen, Mem. New York Bot. Gard. =Ocotea cernua 

Ocotea rhynchophylla (Meisn.) Mez =Ocotea aciphylla 

Ocotea rodiei (R.H. Schomb.) Mez =Chlorocardium rodiei 

Ocotea roraimae auct. non Mez 1920 =Licaria tomentosa 

Ocotea roraimae Mez =Ocotea aciphylla 

Ocotea rubra Mez =Sextonia rubra 

Ocotea sericiflora C.K. Allen =Ocotea aciphylla 

Ocotea simulans C.K. Allen =Endlicheria anomala 

Ocotea steyermarkiana C.K. Allen =Rhodostemonodaphne steyermarkiana 

Ocotea subalveolata C.K. Allen =Ocotea schomburgkiana 

Ocotea subglabra Benoist =Ocotea puberula 

Ocotea urophylla (Meisn.) Mez =Ocotea cernua 

Ocotea vernicosa Mez =Licaria vernicosa 

Ocotea wachenheimii Benoist =Ocotea floribunda 

Oreodaphne aciphylla Nees & Mart. ex Nees =Ocotea aciphylla 

Oreodaphne adenotrachelium Nees =Ocotea adenotrachelium 

Oreodaphne amazonica Meisn. =Ocotea amazonica 

Oreodaphne bracteosa Meisn. =Ocotea bracteosa 

Oreodaphne caudata Nees =Ocotea cernua 

Oreodaphne cernua Nees =Ocotea cernua 

Oreodaphne costulata Nees =Ocotea aciphylla 

Oreodaphne crassifolia Nees =Ocotea crassifolia 

Oreodaphne diospyrifolia Meisn. var. incompta Meisn. =Ocotea pauciflora 

Oreodaphne fasciculata Nees =Ocotea fasciculata 

Oreodaphne florulenta Meisn. =Ocotea neesiana 

Oreodaphne glomerata Nees =Ocotea glomerata 

Oreodaphne gracilis Meisn. =Ocotea gracilis 

Oreodaphne guianensis var. aurea Meisn. =Ocotea guianensis 

Oreodaphne hostmanniana Miq. =Ocotea puberula 

Oreodaphne martiniana Nees =Ocotea puberula 

Oreodaphne pauciflora Nees =Ocotea pauciflora 

Oreodaphne schomburgkiana Nees =Ocotea schomburgkiana 

Oreodaphne semecarpifolia Meisn. =Cinnamomum semecarpifolium 

Oreodaphne splendens Meisn. =Ocotea splendens 

Persea cinnamonifolia Kunth =Cinnamomum triplinerve 

Persea coriacea Kosterm. [type based on mixed collection fide van der Werff] 

Persea fastigiata var. pilosa Kopp =Persea fastigiata 

Persea lignitepala Lasser =Persea caerulea 

Persea sprucei Kosterm. =Cinnamomum semecarpifolium 

Phoebe areolatocostae C.K. Allen =Persea areolatocostae 

Phoebe cinnamomifolia (Kunth) Nees =Cinnamomum triplinerve 

Phoebe erectifolia C.K. Allen =Ocotea erectifolia 

Phoebe filamentosa C.K. Allen =Cinnamomum triplinerve 

Phoebe semecarpifolia (Meisn.) Mez =Cinnamomum semecarpifolium 

Phoebe steyermarkiana C.K. Allen =Ocotea julianii 

Pleurothyrium cowaniana C.K. Allen =Rhodostemonodaphne kunthiana 

Pleurothyrium cowanianum C.K. Allen =Rhodostemonodaphne kunthiana 

Rhodostemonodaphne grandis (Mez) Rohwer =Rhodostemonodaphne praeclara 

Strychnodaphne puberula var. truncata Meisn. =Ocotea puberula 

Systemonodaphne geminiflora Mez =Kubitzkia mezii 

Systemonodaphne macrantha Kosterm. =Kubitzkia macrantha 

Systemonodaphne mezii Kosterm. =Kubitzkia mezii 

REFERENCES 

Allen, C.K. 1964. Lauraceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part 5. Mem. New 

York Bot. Gard. 10(5): 1-278. 

Allen, C.K. 1965. Lauraceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 6. Mem. New York Bot. 

Gard. 12(3): 1-285.

330 


Jansen-Jacobs, M.J. 1976. Lauraceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 451- 

484. 

Kostermans, A.J.G.H. 1936-1938. Revision of the Lauraceae. Rec. Trav. Bot. Neerl. 33: 719-757 (I), 34: 500-609 (II), 35: 56-129 

(III), 35: 834-931 (V). 

Kostermans, A.J.G.H. 1936. Lauraceae. In A. Pulle, Flora of Suriname 2(1): 244-337. 

Kostermans, A.J.G.H. 1939. Lauraceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 481-487. 

Kubitzki, K. & S. Renner. 1982. Lauraceae I (Aniba and Aiouea). Flora Neotropica Monogr. 31: 1-124. 

Rohwer, J.G. 1986. Prodromus einer Monographie der Gattung Ocotea Aubl., sensu lato. Mitt. Inst. Allg. Bot. Hamburg 20: 3- 

278. 

Rohwer, J.G. 1993. Lauraceae: Nectandra. Flora Neotrop. Monogr. 60: 1-332. 

Rohwer, J.G., H.G. Richter, & H. van der Werff. 1991. Two new genera of neotropical Lauraceae and critical remarks on the 

generic delimitation. Ann. Missouri Bot. Gard. 78: 388-400. 

Werff, H. van der. 1986. Kubitzkia van der Werff, a new name for a genus of neotropical Lauraceae. Taxon 35(1): 164-166. 

Werff, H. van der. 1987. A revision of Mezilaurus. Ann. Missouri Bot. Gard. 74: 153-182. 

Werff, H. van der. 1991. A key to the genera of Lauraceae in the New World. Ann. Missouri Bot. Gard. 78: 377-387. 

Werff, H. van der. 1995. Two new species of Lauraceae from central French Guiana. Brittonia 47(4): 372-375. 

Werff, H. van der. 1997. Sextonia, a new genus of Lauraceae from South America. Novon 7(4):436-439. 

Werff, H. van der. 2002. Lauraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


370-384. 

Werff, H. van der & J.G. Rohwer. 1999. Lauraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Allantoma lineata (Mart. ex O. Berg) Miers 

Asteranthos brasiliensis Desf. 

Bertholletia excelsa Bonpl. 

Corythophora amapaensis Pires ex S.A. Mori & Prance 

Corythophora labriculata (Eyma) S.A. Mori & Prance 

Corythophora rimosa W.A. Rodrigues 

Corythophora rimosa subsp. rubra S.A. Mori 

Couratari calycina Sandwith 

Couratari gloriosa Sandwith 

Couratari guianensis Aubl. 

Couratari multiflora (Sm.) Eyma 

Couratari oblongifolia Ducke & R. Knuth 

Couratari oligantha A.C. Sm. 

Couratari riparia Sandwith 

Couratari sandwithii Prance 

Couratari stellata A.C. Sm. 

Couratari tenuicarpa A.C. Sm. 

Couroupita guianensis Aubl. 

Eschweilera alata A.C. Sm. 

Eschweilera apiculata (Miers) A.C. Sm. 

Eschweilera boltenii S.A. Mori 

Eschweilera bracteosa (Poepp. ex O. Berg) Miers 

Eschweilera chartaceifolia S.A. Mori 

Eschweilera collina Eyma 

Eschweilera congestiflora (Benoist) Eyma 

Eschweilera coriacea (DC.) S.A. Mori 

Eschweilera decolorans Sandwith 

Eschweilera eperuetorum Sandwith 

Eschweilera fanshawei Sandwith 

Eschweilera grandiflora (Aubl.) Sandwith 

Eschweilera laevicarpa S.A. Mori 

Eschweilera micrantha (O. Berg) Miers 

Eschweilera neblinensis S.A. Mori 

Eschweilera paniculata (O. Berg) Miers 

Eschweilera parviflora (Aubl.) Miers 

Eschweilera parvifolia Mart. ex DC. 

Eschweilera pedicellata (Rich.) S.A. Mori 

Eschweilera piresii S.A. Mori 

LECYTHIDACEAE 

S.A. Mori, G.T. Prance 

AM 

AM 

AM GU SU FG 

FG 

SU 

SU FG 

SU FG 

GU FG 

GU SU FG 



GU SU FG 

AM GU 

GU 

BO SU 

AM GU SU FG 

SU 


BO GU FG 

SU FG 

GU SU 

AM 

FG 


GU? SU FG 



GU SU 

GU 

FG 

AM BO FG 


AM 

AM 


AM BO GU 


FG

Eschweilera piresii subsp. viridipetala S.A. Mori 

Eschweilera potaroensis Sandwith 

Eschweilera praeclara Sandwith 

Eschweilera revoluta S.A. Mori 

Eschweilera rionegrense S.A. Mori 

Eschweilera roraimensis S.A. Mori 

Eschweilera sagotiana Miers 

Eschweilera simiorum (Benoist) Eyma 

Eschweilera squamata S.A. Mori 

Eschweilera subglandulosa (Steud. ex O. Berg) Miers 

Eschweilera tenuifolia (O. Berg) Miers 

Eschweilera wachenheimii (Benoist) Sandwith 

Gustavia acuminata S.A. Mori 

Gustavia augusta L. 

Gustavia coriacea S.A. Mori 

Gustavia gigantophylla Sandwith 

Gustavia hexapetala (Aubl.) Sm. 

Gustavia poeppigiana O. Berg 

Gustavia pulchra Miers 

Lecythis alutacea (A.C. Sm.) S.A. Mori 

Lecythis brancoensis (R. Knuth) S.A. Mori 

Lecythis chartacea O. Berg 

Lecythis confertiflora (A.C. Sm.) S.A. Mori 

Lecythis corrugata Poit. 

Lecythis corrugata subsp. corrugata 

Lecythis corrugata subsp. rosea (Spruce ex O. Berg) S.A. Mori 

Lecythis holcogyne (Sandwith) S.A. Mori 

Lecythis idatimon Aubl. 

Lecythis persistens Sagot 

Lecythis persistens subsp. aurantiaca S.A. Mori 

Lecythis persistens subsp. persistens 

Lecythis pneumatophora S.A. Mori 

Lecythis poiteaui O. Berg 

Lecythis schomburgkii O. Berg 

Lecythis zabucajo Aubl. 



Allantoma fagifolia (Miq. ex O. Berg) Miers =Couratari multiflora 

Bertholletia nobilis Miers =Bertholletia excelsa 

Cercophora anomala Miers =Lecythis chartacea 

Chytroma amara (Aubl.) Miers =Lecythis idatimon 

Chytroma apiculata Miers =Eschweilera apiculata 

Chytroma brancoensis R. Knuth =Lecythis brancoensis 

Chytroma chartacea (O. Berg) Miers =Lecythis chartacea 

Chytroma collina (Eyma) R. Knuth =Eschweilera collina 

Chytroma congestiflora (Benoist) R. Knuth =Eschweilera congestiflora 

Chytroma corrugata (Poit.) R. Knuth =Lecythis corrugata 

Chytroma foetida R. Knuth =Lecythis poiteaui 

Chytroma holcogyne (Sandwith) R. Knuth =Lecythis holcogyne 

Chytroma idatimon (Aubl.) Nied. =Lecythis idatimon 

Chytroma labriculata (Eyma) R. Knuth =Corythophora labriculata 

Chytroma laevicula (Miers) R. Knuth =Lecythis chartacea 

Chytroma marawynensis (O. Berg) Miers =Lecythis chartacea 

Chytroma perspicua Miers =Eschweilera pedicellata 

Chytroma praeclara (Sandwith) Kunth =Eschweilera praeclara 

Chytroma rosea (Spruce ex O. Berg) Miers =Lecythis corrugata subsp. rosea 

Chytroma rubiflora Miers =Lecythis corrugata subsp. corrugata 

Chytroma salebrosa (O. Berg) Miers =Lecythis corrugata subsp. corrugata 

Chytroma schomburgkiana (O. Berg) Miers =Lecythis schomburgkii 

Chytroma simiorum (Benoist) R. Knuth =Eschweilera simiorum 

Chytroma simiorum var. latifolia (Eyma) R. Knuth =Eschweilera simiorum 

Couratari coriacea Mart. ex O. Berg =Couratari multiflora 

Couratari fagifolia (Miq. ex O. Berg) Eyma =Couratari multiflora 

Couratari lineata Mart. ex O. Berg =Allantoma lineata 

Couratari pulchra Sandwith =Couratari guianensis 

AM 

AM 

AM 

GU 

GU 

FG 

GU SU FG 

GU SU FG 

SU FG 


AM BO 

GU SU FG 

AM 


AM 

GU 


AM BO DA GU 

AM 


GU 


GU SU FG 


DA GU SU FG 

AM BO GU 

GU SU FG 

SU FG 

GU SU? FG 

FG 

GU SU? FG 

FG 

SU FG 

GU 

AM BO DA GU SU FG

332 


Couratari reticulata A.C. Sm. =Couratari stellata 

Couroupita guianensis var. surinamensis (Mart. ex O. Berg) Eyma =Couroupita guianensis 

Couroupita surinamensis Mart. ex O. Berg =Couroupita guianensis 

Couroupita venezuelensis R. Knuth =Couroupita guianensis 

Eschweilera alutacea A.C. Sm. =Lecythis alutacea 

Eschweilera amara (Aubl.) Nied. =Lecythis idatimon 

Eschweilera chaffanjonii (Benoist) R. Knuth =Eschweilera parvifolia 

Eschweilera chartacea (O. Berg) Eyma =Lecythis chartacea 

Eschweilera conduplicata A.C. Sm. =Lecythis corrugata subsp. rosea 

Eschweilera confertiflora A.C. Sm. =Lecythis confertiflora 

Eschweilera confusa (Benoist) R. Knuth [nomen nudum] 

Eschweilera corrugata (Poit.) Miers =Lecythis corrugata 

Eschweilera flaccida Miers =Eschweilera pedicellata 

Eschweilera floribunda Eyma =Eschweilera micrantha 

Eschweilera gracilipes (Sagot) R. Knuth =Eschweilera micrantha 

Eschweilera grata Sandwith =Eschweilera parviflora 

Eschweilera holcogyne Sandwith =Lecythis holcogyne 

Eschweilera idatimon (Aubl.) Nied. =Lecythis idatimon 

Eschweilera idatimonoides (O. Berg) Miers =Eschweilera pedicellata 

Eschweilera jenmanii R. Knuth =Lecythis corrugata subsp. corrugata 

Eschweilera jucunda (Benoist) R. Knuth =Eschweilera sagotiana 

Eschweilera labriculata Eyma =Corythophora labriculata 

Eschweilera longipes (Poit.) Miers =Eschweilera pedicellata 

Eschweilera lutea (Aubl.) Miers =Lecythis idatimon 

Eschweilera macrophylla (O. Berg) Miers =Eschweilera pedicellata 

Eschweilera marawynensis (O. Berg) Nied. =Lecythis chartacea 

Eschweilera melinonis (Sagot) Knuth =Eschweilera sagotiana 

Eschweilera montana A.C. Sm. =Eschweilera parviflora 

Eschweilera obtecta (Miers) Nied. =Eschweilera tenuifolia 

Eschweilera odora (Poepp. ex O. Berg) Miers =Eschweilera coriacea 

Eschweilera ovalis R. Knuth =Lecythis persistens subsp. persistens 

Eschweilera pachycarpa Mart. [nomen nudum] 

Eschweilera pallida Miers =Eschweilera coriacea 

Eschweilera patrisii R. Knuth =Lecythis corrugata subsp. corrugata 

Eschweilera persaudii R. Knuth =Lecythis alutacea 

Eschweilera platycarpa (Poit.) Miers =Eschweilera pedicellata 

Eschweilera poiteaui (O. Berg) Nied. =Lecythis poiteaui 

Eschweilera retroflexa (Benoist) R. Knuth =Eschweilera coriacea 

Eschweilera salebrosa (O. Berg) Nied. =Lecythis corrugata subsp. corrugata 

Eschweilera schomburgkii (O. Berg) Nied. =Lecythis schomburgkii 

Eschweilera simiorum var. latifolia Eyma =Eschweilera simiorum 

Eschweilera tapuya (Benoist) R. Knuth =Eschweilera sagotiana 

Grias aubletiana Miers =Gustavia augusta 

Grias tetrapetala (Aubl.) Nied. =Gustavia augusta 

Gustavia acuta S.A. Mori =Gustavia pulchra 

Gustavia augusta var. guianensis O. Berg =Gustavia augusta 

Gustavia augusta var. verrucosa Mart. ex O. Berg =Gustavia augusta 

Gustavia fastuosa Willd. =Gustavia hexapetala 

Gustavia fastuosa var. angustisepala O. Berg =Gustavia hexapetala 

Gustavia fastuosa var. latisepala O. Berg =Gustavia hexapetala 

Gustavia tetrapetala (Aubl.) DC. =Gustavia augusta 

Gustavia urceolata Poit. =Gustavia augusta 

Gustavia yaracuyensis Pittier =Gustavia poeppigiana 

Japarandiba augusta (L.) Kuntze =Gustavia augusta 

Japarandiba fastuosa (Willd.) Kuntze =Gustavia hexapetala 

Japarandiba hexapetala (Aubl.) Kuntze =Gustavia hexapetala 

Jugastrum christii Pittier =Eschweilera tenuifolia 

Jugastrum obtectum Miers =Eschweilera tenuifolia 

Jugastrum poiteaui (O. Berg) Miers =Lecythis poiteaui 

Jugastrum sifontesii Pittier =Eschweilera tenuifolia 

Lecythis alba S.A. Mori [nomen nudum] =Lecythis persistens subsp. persistens 

Lecythis amara Aubl. =Lecythis idatimon 

Lecythis bracteosa Poepp. ex O. Berg =Eschweilera bracteosa 

Lecythis chaffanjonii Benoist =Eschweilera parvifolia


Lecythis chartacea var. calyce Sagot =Lecythis chartacea 

Lecythis cognata Miers =Lecythis corrugata subsp. corrugata 

Lecythis congestiflora Benoist =Eschweilera congestiflora 

Lecythis coriacea DC. =Eschweilera coriacea 

Lecythis crassinoda Miers =Lecythis zabucajo 

Lecythis davisii Sandwith =Lecythis zabucajo 

Lecythis davisii var. gracilipes Eyma =Lecythis zabucajo 

Lecythis fagifolia Miq. ex O. Berg =Couratari multiflora 

Lecythis gracilipes Sagot =Eschweilera micrantha 

Lecythis grandiflora Aubl. =Eschweilera grandiflora 

Lecythis jucunda Benoist =Eschweilera sagotiana 

Lecythis karuaiensis Steyerm. =Lecythis alutacea 

Lecythis laevicula Miers =Lecythis chartacea 

Lecythis longipes Poit. =Eschweilera pedicellata 

Lecythis lutea Aubl. =Lecythis idatimon 

Lecythis macrophylla O. Berg =Eschweilera pedicellata 

Lecythis marawynensis O. Berg =Lecythis chartacea 

Lecythis martinii O. Berg =Lecythis corrugata subsp. corrugata 

Lecythis melinonis Sagot =Eschweilera sagotiana 

Lecythis micrantha O. Berg =Eschweilera micrantha 

Lecythis multiflora Sm. =Couratari multiflora 

Lecythis odora Poepp. ex O. Berg =Eschweilera coriacea 

Lecythis paniculata O. Berg =Eschweilera paniculata 

Lecythis parviflora Aubl. =Eschweilera parviflora 

Lecythis parviflora Sagot ex Miers =Eschweilera sagotiana 

Lecythis pedicellata Rich. =Eschweilera pedicellata 

Lecythis platycarpa Poit. =Eschweilera pedicellata 

Lecythis racemiflora Sagot =Lecythis poiteaui 

Lecythis retroflexa (Benoist) R. Knuth =Eschweilera coriacea 

Lecythis rosea Spruce ex O. Berg =Lecythis corrugata subsp. rosea 

Lecythis rubicunda Miers =Lecythis corrugata subsp. corrugata 

Lecythis sagotiana (Miers) Sagot =Eschweilera sagotiana 

Lecythis salebrosa O. Berg =Lecythis corrugata subsp. corrugata 

Lecythis simiorum Benoist =Eschweilera simiorum 

Lecythis subglandulosa Steud. ex O. Berg =Eschweilera subglandulosa 

Lecythis tapuya Benoist =Eschweilera sagotiana 

Lecythis tenuifolia O. Berg =Eschweilera tenuifolia 

Lecythis tumefacta Miers =Lecythis zabucajo 

Lecythis venusta Miers =Lecythis corrugata subsp. corrugata 

Lecythis wachenheimii Benoist =Eschweilera wachenheimii 

Lecythis wullschlaegeliana O. Berg =Eschweilera pedicellata 

Pirigara hexapetala Aubl. =Gustavia hexapetala 

Pirigara tetrapetala Aubl. =Gustavia augusta 

REFERENCES 

(for a more extensive bibliography, see Mori & Prance 1993) 

Eyma, P.J. 1934. Lecythidaceae. In A. Pulle, Flora of Suriname 3(1): 119-155. 

Mori, S.A. 1999. Lecythidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


385-397. 

Mori, S.A. & collaborators. 1987. The Lecythidaceae of a lowland neotropical forest: La Fumée Mountain, French Guiana. 


Mori, S.A. & G.T. Prance. 1990. Lecythidaceae - part II. Flora Neotropica Monogr. 21(II): 1-373. 

Mori, S.A. & G.T. Prance. 1993. Lecythidaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 12: 1-144. 

Mori, S.A. & G.T. Prance. 1999. Lecythidaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Prance, G.T. & S.A. Mori. 1979. Lecythidaceae - part I. Flora Neotropica Monogr. 21(I): 1-270. 

LEGUMINOSAE-CAESALPINIOIDEAE 

G.A. Aymard C, R.C. Barneby†, P.E. Berry, R.S. Cowan†, M.F. da Silva, D.M. Kearns, A. Sprada Tavares, 

B. Stergios, D. Velázquez, R.P. Wunderlin, N. Xena, J.L. Zarucchi 

Apuleia leiocarpa (Vogel) J.F. Macbr. 

Barnebydendron riedelii (Tul.) J.H. Kirkbr. 

AM BO 

AM

334 


Batesia floribunda Spruce ex Benth. 

Bauhinia aculeata L. 

Bauhinia bicuspidata Benth. 

Bauhinia breviloba Benth. 

Bauhinia cinnamomea DC. 

Bauhinia confertiflora Benth. 

Bauhinia coronata Benth. 

Bauhinia cupreonitens Ducke 

Bauhinia cupulata Benth. 

Bauhinia eilertsii Pulle 

Bauhinia glabra Jacq. 

Bauhinia guianensis Aubl. 

Bauhinia kunthiana Vogel 

Bauhinia longicuspis Spruce ex Benth. 

Bauhinia microstachya (Raddi) J.F. Macbr. 

Bauhinia outimouta Aubl. [=Bauhinia guianensis?] 

Bauhinia poiteauana Vogel [=Bauhinia smilacina?] 

Bauhinia richardiana DC. 

Bauhinia rutilans Spruce ex Benth. 

Bauhinia scala-simiae Sandwith 

Bauhinia siqueiraei Ducke 

Bauhinia smilacina (Schott) Steud. 

Bauhinia surinamensis Amshoff [=Bauhinia guianensis?] 

Bauhinia ungulata L. 

Bocoa alterna (Benth.) R.S. Cowan 

Bocoa prouacensis Aubl. 

Bocoa viridiflora (Ducke) R.S. Cowan 

Brownea ariza Benth. 

Brownea coccinea Jacq. 

Brownea coccinea subsp. capitella (Jacq.) D. Velázq. & Agostini 

Brownea grandiceps Jacq. 

Brownea guianensis Klotzsch 

Brownea longipedicellata Huber 

Brownea similis R.S. Cowan 

Caesalpinia bonduc (L.) Roxb. 

Caesalpinia coriaria (Jacq.) Willd. 

Caesalpinia granadillo Pittier 

Caesalpinia pulcherrima (L.) Sw. [cultivated; naturalized?] 

Campsiandra angustifolia Spruce ex Benth. 

Campsiandra angustifolia var. angustifolia 

Campsiandra angustifolia var. rosea (Poepp.) Stergios 

Campsiandra aymardii Stergios 

Campsiandra casiquiarensis Stergios 

Campsiandra chigo-montero Stergios 

Campsiandra comosa var. comosa 

Campsiandra comosa var. surinamensis (Kleinhoonte) Stergios 

Campsiandra curaara Stergios 

Campsiandra emonensis Stergios 

Campsiandra ferruginea Stergios 

Campsiandra gomez-alvareziana Stergios 

Campsiandra gomez-alvareziana var. gomez-alvareziana 

Campsiandra gomez-alvareziana var. pusilliflora Stergios 

Campsiandra guayanensis Stergios 

Campsiandra implexicaulis Stergios 

Campsiandra laurifolia Benth. 

Campsiandra macrocarpa R.S. Cowan 

Campsiandra macrocarpa var. alveolata Stergios 

Campsiandra macrocarpa var. grandifolia Stergios 

Campsiandra macrocarpa var. macrocarpa 

Campsiandra nutans Stergios 

Campsiandra pasibensis Stergios 

Campsiandra steyermarkiana Stergios 

Campsiandra taphornii Stergios 

Campsiandra velutina Stergios 

FG 

BO GU 

GU 

GU 

SU 

SU 

SU FG 

FG 

AM GU SU FG 

AM BO 

BO GU SU 




AM FG 

AM GU 


FG 

FG 

AM BO GU? 

BO DA GU 

BO DA GU FG 

SU FG 

DA GU SU FG 

AM BO GU 

GU 

GU SU FG 

AM 

GU SU FG 

AM BO DA GU 

AM BO DA GU 

GU? 

GU 

AM BO 

AM BO 

BO 

BO 

GU SU FG 

GU SU FG 

AM BO 

AM BO 

AM 

AM 

AM 

BO 

AM GU SU 

GU SU 

AM 

AM 

AM 

AM 

AM 

AM 

GU SU 

AM BO 

AM BO DA 

AM 

AM 

AM 

AM 

AM 

GU 

AM 

AM 

AM 

BO 

AM 

AM 

BO


Campsiandra wurdackiana Stergios 

Cassia cowanii H.S. Irwin & Barneby 

Cassia cowanii var. cowanii 

Cassia cowanii var. guianensis (Sandwith) H.S. Irwin & Barneby 

Cassia fastuosa Willd. ex Benth. 

Cassia fastuosa var. fastuosa 

Cassia fistula L. [widely cultivated, occasionally escaping] 

Cassia grandis L. f. 

Cassia moschata Kunth 

Cassia spruceana Benth. 

Chamaecrista adiantifolia (Spruce ex Benth.) H.S. Irwin & Barneby 

Chamaecrista adiantifolia var. adiantifolia 

Chamaecrista adiantifolia var. pteridophylla (Sandwith) H.S. Irwin & 

Barneby [expected in SU, FG] 

Chamaecrista apoucouita (Aubl.) H.S. Irwin & Barneby 

Chamaecrista calycioides (DC. ex Collad.) Greene 

Chamaecrista calycioides var. calycioides 

Chamaecrista desvauxii (Collad.) Killip 

Chamaecrista desvauxii var. brevipes (Benth.) H.S. Irwin & Barneby 

Chamaecrista desvauxii var. mollissima (Benth.) H.S. Irwin & Barneby 

Chamaecrista desvauxii var. piptostegia H.S. Irwin & Barneby 

Chamaecrista desvauxii var. saxatilis (Amshoff) H.S. Irwin & Barneby 

Chamaecrista desvauxii var. triumviralis H.S. Irwin & Barneby 

Chamaecrista diphylla (L.) Greene 

Chamaecrista fagonioides (Vogel) H.S. Irwin & Barneby 

Chamaecrista fagonioides var. fagonioides 

Chamaecrista flexuosa (L.) Greene 

Chamaecrista flexuosa var. flexuosa 

Chamaecrista glandulosa (L.) Greene 

Chamaecrista glandulosa var. crystallina H.S. Irwin & Barneby 

Chamaecrista glandulosa var. rapidarum H.S. Irwin & Barneby 

Chamaecrista hispidula (Vahl) H.S. Irwin & Barneby 

Chamaecrista hymenaeifolia (Benth.) H.S. Irwin & Barneby 

Chamaecrista kunthiana (Schltdl. & Cham.) H.S. Irwin & Barneby 

Chamaecrista negrensis (H.S. Irwin) H.S. Irwin & Barneby 

Chamaecrista negrensis var. negrensis 

Chamaecrista nictitans (L.) Moench 

Chamaecrista nictitans var. disadena (Steud.) H.S. Irwin & Barneby 

Chamaecrista nictitans subsp. disadena (Steud.) H.S. Irwin & Barneby 

Chamaecrista nictitans var. glabrata (Vogel) H.S. Irwin & Barneby [subsp. 

patellaria] [widely naturalized] 

Chamaecrista nictitans var. paraguariensis (Chodat & Hassl.) H.S. Irwin & 

Barneby [subsp. patellaria] 

Chamaecrista nictitans var. patellaria Kartesz & Gandhi 

Chamaecrista nictitans subsp. patellaria (Collad.) H.S. Irwin & Barneby 

Chamaecrista nictitans var. pilosa (Benth.) H.S. Irwin & Barneby 

Chamaecrista nictitans var. praetexta (Vogel) H.S. Irwin & Barneby [subsp. 

patellaria] [expected in BO] 

Chamaecrista orenocensis (Spruce ex Benth.) H.S. Irwin & Barneby 

Chamaecrista pilosa (L.) Greene 

Chamaecrista pilosa var. pilosa 

Chamaecrista polystachya (Benth.) H.S. Irwin & Barneby 

Chamaecrista ramosa (Vogel) H.S. Irwin & Barneby 

Chamaecrista ramosa var. ramosa 

Chamaecrista ramosa var. ventuarensis (H.S. Irwin) H.S. Irwin & Barneby 

Chamaecrista roraimae (Benth.) Gleason 

Chamaecrista rotundifolia (Pers.) Greene [common weed expected in the 


Chamaecrista rotundifolia var. grandiflora (Benth.) H.S. Irwin & Barneby 

Chamaecrista rotundifolia var. rotundifolia 

Chamaecrista serpens (L.) Greene 

Chamaecrista serpens var. grandiflora (Benth.) H.S. Irwin & Barneby 

Chamaecrista serpens var. serpens 

Chamaecrista viscosa (Kunth) H.S. Irwin & Barneby 

AM BO 


GU SU FG 

BO DA GU SU 

AM SU FG 

AM SU FG 

GU FG 

BO GU SU FG 

AM DA GU 

FG 

AM GU 

AM 

AM GU 

GU SU FG 

BO GU FG 

BO 




BO SU 

SU FG 

AM BO 


BO GU SU FG 

BO 


AM BO DA 

AM BO 

AM BO 

AM BO 

BO GU SU FG 

AM 

AM BO 

AM 

AM 


BO GU SU FG 

BO GU SU FG 

AM GU 

BO GU 

BO GU SU FG 


SU 

GU SU FG 

AM 

BO DA 

BO DA 

BO GU 


AM 

AM 

BO 

BO GU 

AM BO GU 

AM BO GU 

AM BO 

AM BO GU 

AM BO 

BO GU 

BO GU SU

336 


Chamaecrista viscosa var. major (Benth.) H.S. Irwin & Barneby 

Chamaecrista viscosa var. viscosa [expected in GU?] 

Chamaecrista zygophylloides (Taub.) H.S. Irwin & Barneby 

Chamaecrista zygophylloides var. deamii (Britton & Rose) H.S. Irwin & 

Barneby 

Copaifera camibar Poveda, N. Zamora & P.E. Sánchez 

Copaifera epunctata Amshoff 

Copaifera guyanensis Desf. 

Copaifera langsdorffii Desf. 

Copaifera officinalis (Jacq.) L. 

Copaifera pubiflora Benth. 

Copaifera reticulata Ducke 

Crudia aequalis Ducke 

Crudia amazonica Spruce ex Benth. 

Crudia aromatica (Aubl.) Willd. 

Crudia bracteata Benth. 

Crudia glaberrima (Steud.) J.F. Macbr. 

Crudia oblonga Benth. 

Crudia spicata (Aubl.) Willd. 

Crudia tomentosa (Aubl.) J.F. Macbr. 

Cynometra bauhiniifolia Benth. 

Cynometra bauhiniifolia var. bauhiniifolia 

Cynometra hostmanniana Tul. 

Cynometra marginata Benth. 

Cynometra marginata var. guianenis Dwyer 

Cynometra marginata var. marginata 

Cynometra parvifolia Tul. 

Cynometra spruceana Benth. 

Cynometra spruceana var. procera Benth. 

Cynometra spruceana var. spruceana 

Delonix regia (Bojer ex Hook.) Raf. [cultivated only?] 

Dialium guianense (Aubl.) Sandwith 

Dicorynia guianensis Amshoff 

Dicorynia paraensis Benth. 

Dicorynia paraensis var. breviflora (Benth.) R.C. Koeppen 

Dicorynia paraensis var. paraensis 

Dicymbe altsonii Sandwith 

Dicymbe bernardii R.S. Cowan 

Dicymbe corymbosa Spruce ex Benth. 

Dicymbe duidae R.S. Cowan 

Dicymbe fraterna R.S. Cowan 

Dicymbe hymenaea Barneby 

Dicymbe jenmanii Sandwith 

Dicymbe mollis Barneby 

Dicymbe neblinensis R.S. Cowan 

Dicymbe paruensis R.S. Cowan 

Dicymbe pharangophila R.S. Cowan 

Dicymbe praeruptorum Barneby 

Dicymbe uaiparuensis R.S. Cowan [expected in GU] 

Dicymbe yutajensis R.S. Cowan 

Dimorphandra conjugata (Splitg.) Sandwith 

Dimorphandra cuprea Sprague & Sandwith 

Dimorphandra cuprea subsp. cuprea 

Dimorphandra cuprea subsp. ferruginea (Ducke) M.F. Silva 

Dimorphandra davisii Sprague & Sandwith 

Dimorphandra dissimilis R.S. Cowan 

Dimorphandra macrostachya subsp. congestiflora (Sprague & Sandwith) M.F. 

Silva 

Dimorphandra macrostachya subsp. glabrifolia (Ducke) M.F. Silva 

Dimorphandra macrostachya subsp. macrostachya 

Dimorphandra multiflora Ducke 

Dimorphandra pennigera Tul. 

Dimorphandra polyandra Benoist 

Dimorphandra pullei Amshoff 

BO GU SU 

BO 

BO GU 

BO GU 

AM 

GU 

GU 

SU 

SU FG 

BO GU? 

AM BO GU 

SU 

AM DA 

GU SU 

FG 

FG 

FG 

DA GU SU FG 


SU FG 

GU FG 



GU SU FG 


GU SU 

GU SU FG 

SU FG 

AM BO GU 

AM BO 

GU 



GU SU FG 

AM 

AM 

AM 

GU 

FG? 

AM 

BO GU 

GU 

AM 

AM 

BO 

BO 

GU 

GU 

SU 

AM GU 

AM 

AM 

AM 

BO 

GU 

GU SU 

AM GU 

GU 

AM GU 

BO 

BO 

GU 

AM BO GU 

AM BO GU FG 


GU FG 

AM 

AM GU SU FG 

SU FG


Dimorphandra unijuga Tul. 

Dimorphandra vernicosa Spruce ex Benth. 

Dimorphandra williamii M.F. Silva 

Dinizia excelsa Ducke [expected in FG] 

Elizabetha bicolor Ducke 

Elizabetha coccinea M.R. Schomb. ex Benth. 

Elizabetha coccinea var. coccinea 

Elizabetha coccinea var. oxyphylla (Harms) R.S. Cowan 

Elizabetha durissima Ducke 

Elizabetha fanshawei R.S. Cowan 

Elizabetha grahamiae R.S. Cowan 

Elizabetha leiogyne Ducke 

Elizabetha macrostachya Benth. 

Elizabetha paraensis Ducke 

Elizabetha princeps M.R. Schomb. ex Benth. 

Eperua bijuga Mart. ex Benth. 

Eperua falcata Aubl. 

Eperua glabra R.S. Cowan 

Eperua grandiflora (Aubl.) Benth. [referable to misidentified E. jenmanii ssp. 

sandwithii in Venezuela] 

Eperua grandiflora subsp. grandiflora 

Eperua grandiflora subsp. guyanensis R.S. Cowan 

Eperua jenmanii Oliv. 

Eperua jenmanii subsp. jenmanii 

Eperua jenmanii subsp. sandwithii R.S. Cowan 

Eperua leucantha Benth. 

Eperua obtusata R.S. Cowan 

Eperua purpurea Benth. 

Eperua rubiginosa Miq. 

Eperua rubiginosa var. grandiflora Pulle 

Eperua rubiginosa var. rubiginosa 

Eperua schomburgkiana Benth. 

Eperua sp. A 

Eperua venosa R.S. Cowan [expected in GU] 

Exostyles sp. 

Haematoxylum campechianum L. [cultivated only?] 

Heterostemon conjugatus Spruce ex Benth. 

Heterostemon ellipticus Mart. ex Benth. 

Heterostemon ingifolius Sandwith 

Heterostemon mazarunensis Sandwith 

Heterostemon mimosoides Desf. 

Heterostemon mimosoides var. complanatus R.S. Cowan 

Heterostemon mimosoides var. mimosoides 

Heterostemon mimosoides var. pacimoniensis R.S. Cowan 

Heterostemon otophorus Sandwith 

Hymenaea courbaril L. 

Hymenaea courbaril var. courbaril 

Hymenaea oblongifolia Huber 

Hymenaea oblongifolia var. davisii (Sandwith) Lee & Langenh. 

Hymenaea oblongifolia var. oblongifolia 

Hymenaea oblongifolia var. palustris (Ducke) Lee & Langenh. [expected in 

SU, FG] 

Hymenaea parvifolia Huber 

Jacqueshuberia brevipes Barneby 

Jacqueshuberia pustulata Stergios & P.E. Berry 

Jacqueshuberia splendens Stergios & P.E. Berry 

Lecointea amazonica Ducke 

Macrolobium acaciifolium (Benth.) Benth. 

Macrolobium acrothamnos R.S. Cowan [expected in GU] 

Macrolobium amplexans (Amshoff) R.S. Cowan 

Macrolobium angustifolium (Benth.) R.S. Cowan 

Macrolobium anomalum R.S. Cowan 

Macrolobium bifolium (Aubl.) Pers. 

Macrolobium campestre Huber 

AM 

AM 

GU 

GU 

GU SU 

GU 

GU SU 

GU 

GU 

GU 

FG? 

AM 

AM GU SU 

SU? FG? 

AM GU SU FG 

FG 


GU FG? 

GU SU FG 

AM 

AM 

AM 

AM 

SU FG 

GU FG 

BO GU SU? FG 

BO GU SU? FG 

BO GU 

BO 

GU SU FG 

GU SU 

GU SU FG 

GU SU FG 

SU 

GU SU FG 

AM 

AM GU 

GU 

GU 

AM GU SU FG 

SU 

AM GU SU FG 

AM 

GU 



GU FG? 

GU 

GU FG? 

GU 

BO 

GU 

AM 

AM 



BO 

SU 


AM 


BO GU SU FG

338 


Macrolobium campestre var. arboreum R.S. Cowan 

Macrolobium canaliculatum Spruce ex Benth. 

Macrolobium canaliculatum var. canaliculatum 

Macrolobium canaliculatum var. strigulosum R.S. Cowan 

Macrolobium cataractarum R.S. Cowan 

Macrolobium discolor Benth. 

Macrolobium discolor var. discolor 

Macrolobium discolor var. egranulosum R.S. Cowan 

Macrolobium evenulosum R.S. Cowan 

Macrolobium exfoliatum R.S. Cowan 

Macrolobium furcatum Ducke [expected in GU] 

Macrolobium gracile Spruce ex Benth. 

Macrolobium gracile var. confertum (Gleason) R.S. Cowan 

Macrolobium gracile var. gracile 

Macrolobium guianense (Aubl.) Pulle 

Macrolobium huberianum Ducke 

Macrolobium huberianum var. huberianum 

Macrolobium huberianum var. pubirachis Amshoff 

Macrolobium jenmanii (Gleason) Sandwith 

Macrolobium limbatum Spruce ex Benth. 

Macrolobium limbatum var. fanshawei (R.S. Cowan) R.S. Cowan 

Macrolobium limbatum var. limbatum 

Macrolobium longeracemosum Amshoff 

Macrolobium longipes R.S. Cowan 

Macrolobium microcalyx Ducke 

Macrolobium molle (Benth.) R.S. Cowan 

Macrolobium montanum Ducke 

Macrolobium montanum var. potaroanum R.S. Cowan 

Macrolobium multijugum (DC.) Benth. 

Macrolobium multijugum var. multijugum 

Macrolobium pendulum Willd. ex Vogel 

Macrolobium punctatum Spruce ex Benth. 

Macrolobium rubrum R.S. Cowan 

Macrolobium savannarum R.S. Cowan 

Macrolobium savannarum var. foldatsii (R.S. Cowan) P.E. Berry 

Macrolobium savannarum var. savannarum 

Macrolobium schinifolium R.S. Cowan 

Macrolobium spectabile R.S. Cowan 

Macrolobium stenopetalum Amshoff 

Macrolobium suaveolens Spruce ex Benth. 

Macrolobium suaveolens var. pakarimense R.S. Cowan 

Macrolobium suaveolens var. suaveolens 

Macrolobium taxifolium Spruce ex Benth. 

Macrolobium unifoliolatum R.S. Cowan 

Macrolobium unifoliolatum var. unifoliolatum 

Macrolobium venulosum Benth. [possibly expected in GU] 

Macrolobium wurdackii R.S. Cowan 

Martiodendron elatum (Ducke) Gleason 

Martiodendron elatum var. elatum 

Martiodendron excelsum (Benth.) Gleason 

Martiodendron mediterraneum (Mart. ex Benth.) R.C. Koeppen 

Martiodendron parviflorum (Amshoff) R. Koeppen 

Mora excelsa Benth. 

Mora gonggrijpii (Kleinhoonte) Sandwith 

Paloue brasiliensis Ducke 

Paloue guianensis Aubl. 

Paloue induta Sandwith 

Paloue riparia Pulle 

Paloveopsis emarginata R.S. Cowan 

Parkinsonia aculeata L. [cultivated only?] 

Peltogyne floribunda (Kunth) Pittier 

Peltogyne lecointei Ducke 

Peltogyne paniculata Benth. 

Peltogyne paniculata subsp. paniculata 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

GU SU FG 

AM 

AM 

AM 

SU FG 

GU FG 

GU 

GU 

GU 

FG 

AM 

AM 

GU 

GU 

BO GU SU 

AM 

AM 

AM 

AM BO 

GU 

GU 

GU SU FG 


AM 

AM 

AM 

AM 

AM 

AM 

AM 

GU SU 

SU 

FG 

AM GU 

AM GU 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

GU 

AM 

AM? 

GU 

SU FG 

BO DA GU SU 

BO DA GU SU 

SU FG 

GU 

GU 

SU FG 

GU 

GU 

SU FG 

BO SU 

BO DA GU 

AM BO GU SU 

FG? 

FG 

AM BO GU SU FG


Peltogyne paniculata subsp. pubescens (Benth.) M.F. Silva 

Peltogyne parvifolia Spruce ex Benth. 

Peltogyne parvifolia subsp. glabra M.F. Silva 

Peltogyne parvifolia subsp. parvifolia 

Peltogyne venosa (Vahl) Benth. 

Peltogyne venosa subsp. densiflora (Spruce ex Benth.) M.F. Silva 

Peltogyne venosa subsp. venosa 

Peltophorum dubium (Spreng.) Taub. 

Peltophorum pterocarpum (DC.) Backer ex K. Heyne [cultivated only?] 

Recordoxylon pulcherrimum Barneby 

Recordoxylon speciosum (Benoist) Gazel ex Barneby 

Schizolobium parahyba (Vell.) S.F. Blake 

Schizolobium parahyba var. amazonicum (Huber ex Ducke) Barneby 

Senna aculeata (Benth.) H.S. Irwin & Barneby 

Senna alata (L.) Roxb. 

Senna atomaria (L.) H.S. Irwin & Barneby 

Senna bacillaris (L. f.) H.S. Irwin & Barneby 

Senna bacillaris var. bacillaris 

Senna bacillaris var. benthamiana (J.F. Macbr.) H.S. Irwin & Barneby 

Senna bicapsularis (L.) Roxb. 

Senna bicapsularis var. bicapsularis 

Senna chrysocarpa (Desv.) H.S. Irwin & Barneby 

Senna cobanensis (Britton) H.S. Irwin & Barneby 

Senna georgica H.S. Irwin & Barneby 

Senna hirsuta (L.) H.S. Irwin & Barneby 

Senna hirsuta var. hirsuta 

Senna latifolia (G. Mey.) H.S. Irwin & Barneby 

Senna ligustrina (L.) H.S. Irwin & Barneby 

Senna lourteigiana H.S. Irwin & Barneby 

Senna macranthera (DC. ex Collad.) H.S. Irwin & Barneby [cultivated, 

perhaps escaping] 

Senna macranthera var. quadrifoliolata (Pittier) H.S. Irwin & Barneby 

Senna macrophylla (Kunth) H.S. Irwin & Barneby 

Senna macrophylla var. gigantifolia (Britton & Killip) H.S. Irwin & Barneby 

Senna multijuga (Rich.) H.S. Irwin & Barneby 

Senna multijuga subsp. lindleyana (Gardner) H.S. Irwin & Barneby 

Senna multijuga var. multijuga 

Senna multijuga subsp. multijuga 

Senna multijuga var. peregrinatrix H.S. Irwin & Barneby [ssp. lindleyana] 

Senna obtusifolia (L.) H.S. Irwin & Barneby 

Senna occidentalis (L.) Link 

Senna pallida (Vahl) H.S. Irwin & Barneby [expected] 

Senna pallida var. pallida [expected in GU] 

Senna pendula (Humb. & Bonpl. ex Willd.) H.S. Irwin & Barneby 

Senna pendula var. indecora (Kunth) Luckow 

Senna pendula var. meticola H.S. Irwin & Barneby [expected in BO] 

Senna pendula var. tenuifolia (Benth.) H.S. Irwin & Barneby 

Senna pilifera (Vogel) H.S. Irwin & Barneby 

Senna pilifera var. subglabra (S. Moore) H.S. Irwin & Barneby [expected in 

the Guianas] 

Senna polyphylla (Jacq.) H.S. Irwin & Barneby [cultivated, perhaps escaping] 

Senna quinquangulata (Rich.) H.S. Irwin & Barneby 

Senna quinquangulata var. quinquangulata 

Senna reticulata (Willd.) H.S. Irwin & Barneby 

Senna robiniifolia (Benth.) H.S. Irwin & Barneby 

Senna rupununiensis H.S. Irwin & Barneby [expected in BO] 

Senna sandwithiana H.S. Irwin & Barneby 

Senna siamea (Lam.) H.S. Irwin & Barneby [cultivated, occasionally 

escaping] 

Senna silvestris (Vell.) H.S. Irwin & Barneby 

Senna silvestris var. silvestris [expected in FG] 

Senna sophera (L.) Roxb. 

Senna spectabilis (DC.) H.S. Irwin & Barneby 

Senna spectabilis var. spectabilis 


AM 

AM 

AM 



AM GU SU FG 

GU 

BO GU SU FG 

AM 

GU FG 

BO 

BO 

BO 


BO 


BO GU SU FG 

AM 

BO GU SU FG 

BO GU SU FG 

BO GU SU FG 

AM 

FG 

GU SU FG 

GU SU FG 


GU? 

SU? FG 

BO 

BO 

AM BO 

AM BO 


BO 



BO 



BO DA GU 

BO DA 

BO 

BO 

GU 


AM BO 


BO 

GU 

BO GU 


AM BO GU SU 

GU 

BO DA 

BO DA

340 


Senna spinescens (Vogel) H.S. Irwin & Barneby 

Senna spinescens var. spinescens 

Senna splendida (Vogel) H.S. Irwin & Barneby [native? Widely cultivated] 

Senna undulata (Benth.) H.S. Irwin & Barneby 

Senna velutina (Vogel) H.S. Irwin & Barneby 

Senna wurdackii H.S. Irwin & Barneby 

Tachigali albiflora (Benoist) Zarucchi & Herend. 

Tachigali amplifolia (Ducke) Barneby 

Tachigali amplifolia (Ducke) Barneby 

Tachigali bracteolata Dwyer 

Tachigali bracteosa (Harms) Zarucchi & Pipoly 

Tachigali cavipes (Spruce ex Benth.) J.F. Macbr. 

Tachigali chrysophylla (Poepp.) Zarucchi & Herend. 

Tachigali davidsei Zarucchi & Herend. 

Tachigali dwyeri (R.S. Cowan) Zarucchi & Herend. 

Tachigali guianensis (Benth.) Zarucchi & Herend. 

Tachigali melinonii (Harms) Zarucchi & Herend. 

Tachigali micropetala (Ducke) Zarucchi & Pipoly 

Tachigali myrmecophila (Ducke) Ducke 

Tachigali odoratissima (Spruce ex Benth.) Zarucchi & Herend. 

Tachigali paniculata Aubl. 

Tachigali paniculata var. angustifolia (Miq.) Dwyer 

Tachigali paniculata var. paniculata 

Tachigali paniculata var. sulcata (Benoist) Dwyer 

Tachigali paraënsis (Huber) Barneby 

Tachigali physophora (Huber) Zarucchi & Herend. 

Tachigali pimichinensis (R.S. Cowan) Zarucchi & Herend. 

Tachigali plumbea Ducke 

Tachigali ptychophysca Spruce ex Benth. [expected in Venezuelan Guayana] 

Tachigali pubiflora Benth. 

Tachigali pulchra Dwyer 

Tachigali reticulosa (Dwyer) Zarucchi & Herend. 

Tachigali rigida Ducke 

Tachigali rusbyi Harms 

Tachigali schultesiana Dwyer [expected in Venezuelan Guayana] 

Tachigali tinctoria (Benth.) Zarucchi & Herend. 

Tamarindus indica L. [cultivated] 

Vouacapoua americana Aubl. 

Vouacapoua macropetala Sandwith 


Adipera bicapsularis (L.) Britton & Rose =Senna bicapsularis 

Adipera indecora (Kunth) Britton & Rose =Senna pendula var. indecora 

Adipera spiciflora Pittier =Senna bicapsularis 

Amphymenium mediterraneum Mart. ex Benth. =Martiodendron mediterraneum 

Apalatoa spicata Aubl. =Crudia spicata 

Apoleya leiocarpa (Vogel) Gleason =Apuleia leiocarpa 

Apoleya molaris (Spruce ex Benth.) Gleason =Apuleia leiocarpa 

Apuleia molaris Spruce ex Benth. =Apuleia leiocarpa 

Apuleia praecox Mart. =Apuleia leiocarpa 

Arouna divaricata Willd. =Dialium guianense 

Arouna guianensis Aubl. =Dialium guianense 

Bauhinia bahiensis Bong. =Bauhinia microstachya 

Bauhinia benthamiana Taub. =Bauhinia ungulata 

Bauhinia brachystachya (Benth.) Walp. =Bauhinia glabra 

Bauhinia bredemeyeri Vogel =Bauhinia aculeata 

Bauhinia chalkos R.S. Cowan =Bauhinia cupreonitens 

Bauhinia chrysophylla Vogel =Bauhinia guianensis 

Bauhinia coronata auct. non Benth.: sensu Pittier et al. =Bauhinia outimouta 

Bauhinia cumanensis Kunth =Bauhinia glabra 

Bauhinia emarginata Mill. =Bauhinia aculeata 

Bauhinia excisa (Griseb.) Hemsl. =Bauhinia guianensis 

Bauhinia glandulosa (Kunth) DC. =Bauhinia aculeata 

Bauhinia guianensis var. kunthiana (Vogel) Wunderlin =Bauhinia kunthiana 

AM BO DA 

AM BO DA 

AM BO 

GU 

GU 

AM GU 

AM BO 

BO DA GU SU 

SU? 

FG 

FG 

FG 

FG 

AM FG? 

AM 

AM 

BO 

BO 


SU FG 

GU SU 

AM 

SU FG 


SU FG 

SU FG 

FG 

AM 

AM 

AM 

SU FG 

GU 

BO 

AM 

AM 

AM BO DA GU 

FG 

BO DA GU 

GU? 

GU 

SU FG


Bauhinia guianensis var. splendens (Kunth) Amshoff =Bauhinia guianensis 

Bauhinia heterophylla Kunth =Bauhinia glabra 

Bauhinia langsdorffiana Bong. =Bauhinia microstachya 

Bauhinia langsdorffiana var. bahiensis (Bong.) Benth. =Bauhinia microstachya 

Bauhinia longipetala (Benth.) Walp. =Bauhinia glabra 

Bauhinia macrostachya Benth. =Bauhinia ungulata 

Bauhinia microstachya var. bahiensis (Bong.) J.F. Macbr. =Bauhinia microstachya 

Bauhinia miranda Pittier =Bauhinia aculeata 

Bauhinia mollicella S.F. Blake =Bauhinia aculeata 

Bauhinia mollifolia Pittier =Bauhinia aculeata 

Bauhinia platycalyx Benth. =Bauhinia guianensis 

Bauhinia punctata Jacq. [nom. illeg.] =Bauhinia glabra 

Bauhinia riparia Splitg. ex Benth. =Bauhinia outimouta 

Bauhinia rotundata Mill. =Bauhinia aculeata 

Bauhinia rubiginosa Bong. =Bauhinia outimouta 

Bauhinia schultzei Harms =Bauhinia aculeata 

Bauhinia speciosa Vogel, non Roxb. =Bauhinia outimouta 

Bauhinia spicata Vogel =Bauhinia microstachya 

Bauhinia splendens Kunth =Bauhinia guianensis 

Bauhinia splendens var. latifolia Benth. =Bauhinia guianensis 

Bauhinia sprucei var. acuminata Benth. =Bauhinia guianensis 

Bauhinia superba Steud. [nom. illeg.] =Bauhinia outimouta 

Bauhinia ungula Jacq. =Bauhinia aculeata 

Bauhinia vogeliana Steud. =Bauhinia microstachya 

Binaria cumanensis (Kunth) Raf. =Bauhinia glabra 

Brownea capitella Jacq. =Brownea coccinea subsp. capitella 

Brownea caurensis Pittier [nomen nudum] =Brownea similis 

Brownea caurensis var. major Pittier [nomen nudum] =Brownea similis 

Brownea latifolia Jacq. =Brownea coccinea subsp. capitella 

Brownea racemosa Jacq. =Brownea coccinea subsp. capitella 

Caesalpinia ferruginea Decne. =Peltophorum pterocarpum 

Caesalpinia inermis Roxb. [nomen nudum] =Peltophorum pterocarpum 

Caesalpinia parahyba (Vell.) Allemno =Schizolobium parahyba 

Campsiandra comosa Benth. [Campsiandra chigo-montero from Venezuela possibly misid. as this] =Campsiandra chigomontero 

Campsiandra comosa var. laurifolia (Benth.) Cowan =Campsiandra laurifolia 

Campsiandra rosea Poepp. =Campsiandra angustifolia var. rosea 

Cansenia ungulata (L.) Raf. =Bauhinia ungulata 

Cassia aculeata Benth. =Senna aculeata 

Cassia adiantifolia Spruce ex Benth. =Chamaecrista adiantifolia 

Cassia adiantifolia var. pteridophylla (Sandwith) Ducke =Chamaecrista adiantifolia var. pteridophylla 

Cassia aeschinomene Collad. =Chamaecrista nictitans var. glabrata 

Cassia alata L. =Senna alata 

Cassia ampliflora Steud. =Senna multijuga 

Cassia apoucouita Aubl. =Chamaecrista apoucouita 

Cassia arenaria Kunth =Chamaecrista flexuosa 

Cassia atomaria L. =Senna atomaria 

Cassia bacillaris L. f. =Senna bacillaris 

Cassia bauhiniifolia Kunth =Chamaecrista rotundifolia var. grandiflora 

Cassia bicapsularis L. =Senna bicapsularis 

Cassia bicapsularis var. indecora (Kunth) Benth. =Senna pendula var. indecora 

Cassia bicapsularis var. tenuifolia Humb. & Bonpl. ex Willd. =Senna pendula var. tenuifolia 

Cassia biflora L. [ambiguous name, probably applied to Senna pallida] 

Cassia brevipes DC. ex Collad. =Chamaecrista desvauxii var. brevipes 

Cassia calliantha G. Mey. =Senna multijuga 

Cassia calycioides DC. ex Collad. =Chamaecrista calycioides 

Cassia chrysocarpa Desv. =Senna chrysocarpa 

Cassia chrysogyne Miq. =Senna chrysocarpa 

Cassia cobanensis (Britton) Lundell =Senna cobanensis 

Cassia cultrifolia Kunth =Chamaecrista diphylla 

Cassia desvauxii Collad. =Chamaecrista desvauxii 

Cassia desvauxii var. brevipes Benth. =Chamaecrista desvauxii var. brevipes 

Cassia desvauxii var. mollissima Benth. =Chamaecrista desvauxii var. mollissima 

Cassia diphylla L. =Chamaecrista diphylla

342 


Cassia disadena Steud. =Chamaecrista nictitans subsp. disadena 

Cassia elliptica Kunth =Senna atomaria 

Cassia emarginata auct. non L. 1753 nec Isandrina emarginata auct. non (L.) Britton & Rose ex Britton & P. Wilson 1924 

=Senna atomaria 

Cassia fagonioides Vogel =Chamaecrista fagonioides 

Cassia filipes Benth. =Chamaecrista rotundifolia var. grandiflora 

Cassia flavicoma var. paraguariensis Chodat & Hassl. =Chamaecrista nictitans var. paraguariensis 

Cassia flexuosa L. =Chamaecrista flexuosa 

Cassia fruticosa Mill. [not in flora area] =Senna fruticosa (Mill.) Irwin & Barneby 

Cassia fruticosa var. benthamiana J.F. Macbr. =Senna bacillaris var. benthamiana 

Cassia glandulosa L. =Chamaecrista glandulosa 

Cassia glandulosa var. swartzii (Wikstr.) J.F. Macbr. [not in flora area] =Chamaecrista glandulosa var. swartzii 

Cassia gracilis Kunth =Chamaecrista desvauxii var. triumviralis 

Cassia hirsuta L. =Senna hirsuta 

Cassia hispidula Vahl =Chamaecrista hispidula 

Cassia hoffmannseggii Mart. ex Benth. =Senna latifolia 

Cassia hymenaeifolia Benth. =Chamaecrista hymenaeifolia 

Cassia indecora Kunth =Senna pendula var. indecora 

Cassia indecora var. advena Willd. ex Vogel =Senna pendula var. indecora 

Cassia kunthiana Schltdl. & Cham. =Chamaecrista kunthiana 

Cassia latifolia G. Mey. =Senna latifolia 

Cassia leiandra Benth. [Guianan records erroneous, but possibly expected] 

Cassia leiandra var. guianensis Sandwith =Cassia cowanii var. guianensis 

Cassia leiantha Benth. =Chamaecrista hispidula 

Cassia ligustrina L. =Senna ligustrina 

Cassia lindleyana Gardner =Senna multijuga subsp. lindleyana 

Cassia lotoides Kunth =Chamaecrista hispidula 

Cassia lucens Vogel =Senna silvestris var. silvestris 

Cassia macranthera DC. ex Collad. =Senna macranthera 

Cassia macrophylla Kunth =Senna macrophylla 

Cassia maritima Willd. ex Vogel =Senna pilifera var. subglabra 

Cassia mimosoides L. [not in flora area] =Chamaecrista mimosoides 

Cassia multijuga Rich. =Senna multijuga 

Cassia negrensis H.S. Irwin =Chamaecrista negrensis 

Cassia nictitans L. =Chamaecrista nictitans 

Cassia obtusifolia L. =Senna obtusifolia 

Cassia occidentalis L. =Senna occidentalis 

Cassia orenocensis Spruce ex Benth. =Chamaecrista orenocensis 

Cassia parahyba Vell. =Schizolobium parahyba 

Cassia paramariboensis Miq. (Leguminosae-Faboideae) =Aeschynomene sensitiva 

Cassia patellaria Collad. =Chamaecrista nictitans subsp. patellaria 

Cassia patellaria var. glabrata Vogel =Chamaecrista nictitans var. glabrata 

Cassia patellaria var. longifolia Benth. =Chamaecrista nictitans subsp. patellaria 

Cassia patellaria var. ramosa Vogel =Chamaecrista nictitans var. patellaria 

Cassia pendula Humb. & Bonpl. ex Willd. =Senna pendula 

Cassia pennelliana Amshoff [not in flora area] =Chamaecrista nictitans var. paraguariensis 

Cassia pilifera Vogel =Senna pilifera 

Cassia pilifera var. subglabra S. Moore =Senna pilifera var. subglabra 

Cassia pilosa L. =Chamaecrista pilosa 

Cassia polyphylla Jacq. =Senna polyphylla 

Cassia polystachya Benth. =Chamaecrista polystachya 

Cassia praetexta Vogel =Chamaecrista nictitans var. praetexta 

Cassia procumbens Mill. =Chamaecrista hispidula 

Cassia prostrata Humb. & Bonpl. ex Willd. =Chamaecrista serpens 

Cassia pteridophylla Sandwith =Chamaecrista adiantifolia var. pteridophylla 

Cassia pulchra Kunth =Chamaecrista desvauxii var. mollissima 

Cassia quinquangulata Rich. =Senna quinquangulata 

Cassia racemosa Mill. [not in flora area] =Senna racemosa (Mill.) Irwin & Barneby 

Cassia ramosa Vogel =Chamaecrista ramosa 

Cassia reticulata Willd. =Senna reticulata 

Cassia richardiana Kunth =Senna multijuga 

Cassia robiniifolia Benth. =Senna robiniifolia 

Cassia roraimae Benth. =Chamaecrista roraimae 

Cassia rotundifolia Pers. =Chamaecrista rotundifolia


Cassia rotundifolia var. bauhiniifolia (Kunth) Benth. =Chamaecrista rotundifolia var. grandiflora 

Cassia rotundifolia var. grandiflora Benth. =Chamaecrista rotundifolia var. grandiflora 

Cassia sagotiana Benth. =Cassia spruceana 

Cassia savannarum Miq. =Chamaecrista ramosa 

Cassia saxatilis (Amshoff) H.S. Irwin =Chamaecrista desvauxii var. saxatilis 

Cassia serpens L. =Chamaecrista serpens 

Cassia serpens var. grandiflora Benth. =Chamaecrista serpens var. grandiflora 

Cassia siamea Lam. =Senna siamea 

Cassia silvestris Vell. =Senna silvestris 

Cassia simplex Standl. =Chamaecrista nictitans var. paraguariensis 

Cassia spectabilis DC. =Senna spectabilis 

Cassia spinescens Vogel =Senna spinescens 

Cassia stenocarpa Vogel =Chamaecrista nictitans var. disadena 

Cassia tagera authors., auct. non L. 1753: sensu American =Chamaecrista kunthiana 

Cassia tarantan Kunth =Senna reticulata 

Cassia tenella Kunth =Chamaecrista serpens var. grandiflora 

Cassia tetraphylla Martyn =Chamaecrista hispidula 

Cassia tetraphylla Desv. =Chamaecrista desvauxii 

Cassia tetraphylla var. aurivilla H.S. Irwin =Chamaecrista desvauxii var. brevipes 

Cassia tetraphylla var. mollissima (Benth.) H.S. Irwin =Chamaecrista desvauxii var. mollissima 

Cassia tetraphylla var. ramosa (Vogel) Amshoff =Chamaecrista ramosa 

Cassia tetraphylla var. ventuarensis H.S. Irwin =Chamaecrista ramosa var. ventuarensis 

Cassia tora authors, auct. non L. 1762: sensu American nec Emelista tora auct. non (L.) Britton & Rose 1924: sensu 

American authors =Senna obtusifolia 

Cassia tora L. [not in flora area] 

Cassia undulata Benth. =Senna undulata 

Cassia uniflora Spreng. [dubious name, not in Guianas] 

Cassia uniflora var. ramosa (Vogel) Benth. =Chamaecrista ramosa 

Cassia velutina Vogel =Senna velutina 

Cassia viminea L. [not in flora area] =Senna viminea (L.) Irwin & Barneby 

Cassia virgata Rich. =Senna chrysocarpa 

Cassia viscosa Kunth =Chamaecrista viscosa 

Cassia viscosa var. major Benth. =Chamaecrista viscosa var. major 

Cassia viscosopilosa Steud. =Chamaecrista viscosa var. major 

Cassia zygophylloides Taub. =Chamaecrista zygophylloides 

Cassia zygophylloides var. deamii (Britton & Rose) H.S. Irwin & Barneby =Chamaecrista zygophylloides var. deamii 

Cathartocarpus moschatus (Kunth) G. Don =Cassia moschata 

Caulotretus heterophyllus (Kunth) Warb. =Bauhinia glabra 

Chamaecrista aeschinomene (Collad.) Greene =Chamaecrista nictitans var. glabrata 

Chamaecrista bauhiniifolia (Kunth) Gleason =Chamaecrista rotundifolia var. grandiflora 

Chamaecrista brevipes (DC. ex Collad.) Greene =Chamaecrista desvauxii var. brevipes 

Chamaecrista cultrifolia (Kunth) Britton & Rose ex Britton & Killip =Chamaecrista diphylla 

Chamaecrista gracilis (Kunth) Pittier =Chamaecrista desvauxii var. triumviralis 

Chamaecrista lucesiae Pittier =Chamaecrista nictitans var. praetexta 

Chamaecrista nictitans var. ramosa (Vogel) H.S. Irwin & Barneby =Chamaecrista nictitans var. patellaria 

Chamaecrista patellaria (Collad.) Greene =Chamaecrista nictitans subsp. patellaria 

Chamaecrista pulchra (Kunth) Britton & Rose =Chamaecrista desvauxii var. mollissima 

Chamaecrista ramulosa Killip & Pittier =Chamaecrista ramosa 

Chamaecrista simplex Standl. =Chamaecrista nictitans var. paraguariensis 

Chamaecrista tenella (Kunth) Pittier et al. =Chamaecrista serpens var. grandiflora 

Chamaefistula barbinervis Pittier =Senna macrophylla var. gigantifolia 

Chamaefistula fruticosa auct. non Cassia fruticosa Mill. 1768. =Senna bacillaris 

Chamaefistula gigantifolia Britton & Killip =Senna macrophylla var. gigantifolia 

Chamaefistula indecora (Kunth) G. Don =Senna pendula var. indecora 

Chamaefistula quadrifoliolata Pittier =Senna macranthera var. quadrifoliolata 

Chamaefistula quinquangulata (Rich.) Pittier =Senna quinquangulata 

Chamaefistula undulata (Benth.) Pittier =Senna undulata 

Chamaesenna aculeata (Benth.) Pittier =Senna aculeata 

Copaiba officinalis Adans. =Copaifera officinalis 

Copaifera jacquinii Desf. =Copaifera officinalis 

Copaifera martii var. pubiflora (Benth.) Benth. =Copaifera pubiflora 

Copaiva officinalis Jacq. =Copaifera officinalis 

Crudia obliqua Griseb. =Crudia glaberrima 

Crudia parivoa DC. =Crudia tomentosa

344 


Crudia pubescens Spruce ex Benth. =Crudia oblonga 

Cynometra bauhiniifolia var. grandiflora Dwyer =Cynometra bauhiniifolia var. bauhiniifolia 

Cynometra marginata var. guianensis Ducke =Cynometra marginata 

Cynometra marginata var. laevis Ducke =Cynometra marginata 

Cynometra microflora R.S. Cowan =Cynometra bauhiniifolia var. bauhiniifolia 

Dialium divaricatum (Willd.) Vahl =Dialium guianense 

Dicorynia breviflora Benth. =Dicorynia paraensis var. breviflora 

Dicorynia spruceana Benth. =Dicorynia paraensis var. breviflora 

Dimorpha falcata (Aubl.) Sm. =Eperua falcata 

Dimorphandra congestiflora Sprague & Sandwith =Dimorphandra macrostachya subsp. congestiflora 

Dimorphandra excelsa (Benth.) Baill. =Mora excelsa 

Dimorphandra ferruginea Ducke =Dimorphandra cuprea subsp. ferruginea 

Dimorphandra glabrifolia Ducke =Dimorphandra macrostachya subsp. glabrifolia 

Dimorphandra gonggrijpii Kleinhoonte =Mora gonggrijpii 

Dimorphandra guianensis (R.H. Schomb. ex Benth. & Hook. f.) Baill. =Mora excelsa 

Dimorphandra hohenkerkii Sprague & Sandwith =Dimorphandra polyandra 

Dimorphandra latifolia Tul. =Dimorphandra conjugata 

Dimorphandra macrostachya auct. non Benth.: sensu Ducke =Dimorphandra macrostachya subsp. glabrifolia 

Dimorphandra mora Benth. & Hook. f. =Mora excelsa 

Ditremexa occidentalis (L.) Britton & Rose =Senna occidentalis 

Echinocassia aculeata (Benth.) Britton & Rose =Senna aculeata 

Elizabetha oxyphylla Harms =Elizabetha coccinea var. oxyphylla 

Eperua kourouensis Benoist =Eperua grandiflora 

Eperua stipulata Kleinhoonte =Eperua jenmanii subsp. sandwithii 

Grimaldia baileyorum Britton & Rose ex L.H. Bailey =Chamaecrista fagonioides 

Grimaldia deamii Britton & Rose =Chamaecrista zygophylloides var. deamii 

Grimaldia orenocensis (Spruce ex Benth.) Pittier =Chamaecrista orenocensis 

Hermesias ariza (Benth.) Kuntze =Brownea ariza 

Hermesias capitella (Jacq.) Kuntze =Brownea coccinea subsp. capitella 

Hermesias latifolia (Jacq.) Kuntze =Brownea coccinea subsp. capitella 

Hermesias racemosa (Jacq.) Kuntze =Brownea coccinea subsp. capitella 

Heterostemon cauliflorus Pittier [nomen nudum] =Heterostemon conjugatus 

Hymenaea courbaril var. obtusifolia Ducke =Hymenaea courbaril 

Hymenaea davisii Sandwith =Hymenaea oblongifolia var. davisii 

Hymenaea floribunda Kunth =Peltogyne floribunda 

Hymenaea latifolia Hayne =Peltogyne floribunda 

Hymenaea multiflora Kleinhoonte =Hymenaea courbaril 

Hymenaea palustris Ducke =Hymenaea oblongifolia var. palustris 

Hymenaea resinifera Salisb. =Hymenaea courbaril 

Hymenaea venosa Vahl =Peltogyne venosa 

Leptolobium leiocarpum Vogel =Apuleia leiocarpa 

Libidibia granadillo (Pittier) Pittier =Caesalpinia granadillo 

Macrolobium acaciifolium var. vestitum Sandwith =Macrolobium acaciifolium 

Macrolobium bifolium var. amplexans Amshoff =Macrolobium amplexans 

Macrolobium chrysostachyum (Miq.) Benth. =Macrolobium angustifolium 

Macrolobium confertum Gleason =Macrolobium gracile var. confertum 

Macrolobium elegans Miq. =Macrolobium bifolium 

Macrolobium flexuosum var. molle Benth. =Macrolobium molle 

Macrolobium foldatsii R.S. Cowan =Macrolobium savannarum var. foldatsii 

Macrolobium hymenifolium Pittier =Macrolobium angustifolium 

Macrolobium hymenioides Willd. =Macrolobium bifolium 

Macrolobium simira (Aubl.) J.F. Gmel. (incertae sedis) =Vouapa simira 

Macrolobium stamineum G. Mey. =Macrolobium bifolium 

Macrolobium tenue Ducke =Macrolobium gracile var. gracile 

Macrolobium unijugum (Poepp. & Endl.) R.S. Cowan =Macrolobium limbatum 

Macrolobium unijugum var. fanshawei R.S. Cowan =Macrolobium limbatum var. fanshawei 

Martia excelsa (Benth.) Benth. =Martiodendron excelsum 

Martia parvifolia (Benth.) Benth. =Martiodendron mediterraneum 

Martiodendron parvifolium (Benth.) Gleason =Martiodendron mediterraneum 

Martiusia elata Ducke =Martiodendron elatum 

Martiusia excelsa Benth. =Martiodendron excelsum 

Martiusia parviflora Amshoff =Martiodendron parviflorum 

Martiusia parvifolia Benth. =Martiodendron mediterraneum 

Melanoxylon speciosum Benoist =Recordoxylon speciosum


Mora guianensis R.H. Schomb. ex Benth. & Hook. f. =Mora excelsa 

Outea acaciifolia Benth. =Macrolobium acaciifolium 

Outea guianensis Aubl. =Macrolobium guianense 

Outea multijuga DC. =Macrolobium multijugum 

Panzera falcata (Aubl.) Willd. =Eperua falcata 

Parivoa grandiflora Aubl. =Eperua grandiflora 

Parivoa tomentosa Aubl. =Crudia tomentosa 

Pauletia glandulosa Kunth =Bauhinia aculeata 

Pauletia ungulata (L.) A. Schmitz =Bauhinia ungulata 

Peiranisia macrochlamys Pittier =Senna robiniifolia 

Peiranisia robiniifolia (Benth.) Britton & Rose =Senna robiniifolia 

Peltogyne amplissima Pittier =Peltogyne paniculata subsp. pubescens 

Peltogyne densiflora Spruce ex Benth. =Peltogyne venosa subsp. densiflora 

Peltogyne micrantha Ducke =Peltogyne parvifolia subsp. parvifolia 

Peltogyne paraensis Huber =Peltogyne venosa subsp. densiflora 

Peltogyne porphyrocardia Griseb. ex Benth. [nomen nudum] =Peltogyne floribunda 

Peltogyne pubescens Benth. =Peltogyne paniculata subsp. pubescens 

Peltogyne venosa var. densiflora (Spruce ex Benth.) Amshoff =Peltogyne venosa subsp. densiflora 

Peltophorum ferrugineum (Decne.) Benth. =Peltophorum pterocarpum 

Peltophorum inerme (Roxb.) Náves ex Fern.-Vill. =Peltophorum pterocarpum 

Phyllocarpus riedelii Tul. 

Poinciana coriaria Jacq. =Caesalpinia coriaria 

Poinciana pulcherrima L. =Caesalpinia pulcherrima 

Poinciana regia Bojer ex Hook. =Delonix regia 

Pseudocassia spectabilis (DC.) Britton & Rose =Senna spectabilis 

Recordoxylon amazonicum (Ducke) Ducke =Recordoxylon speciosum 

Schizolobium amazonicum Huber ex Ducke =Schizolobium parahyba var. amazonicum 

Schizolobium excelsum Vogel =Schizolobium parahyba 

Schnella bicomata Pittier =Bauhinia guianensis 

Schnella brachystachya Benth. =Bauhinia glabra 

Schnella cumanensis (Kunth) Britton & Rose =Bauhinia glabra 

Schnella excisa Griseb. =Bauhinia guianensis 

Schnella glabra (Jacq.) Dugand =Bauhinia glabra 

Schnella heterophylla (Kunth) Benth. =Bauhinia glabra 

Schnella longipetala Benth. =Bauhinia glabra 

Schnella microstachya Raddi =Bauhinia microstachya 

Schnella rubiginosa (Bong.) Benth. =Bauhinia outimouta 

Schnella splendens (Kunth) Benth. =Bauhinia guianensis 

Sclerolobium albiflorum Benoist =Tachigali albiflora 

Sclerolobium amplifolium Ducke 

Sclerolobium aureum var. grandiflorum Dwyer =Tachigali davidsei 

Sclerolobium bracteosum Harms =Tachigali bracteosa 

Sclerolobium chrysophyllum Poepp. =Tachigali chrysophylla 

Sclerolobium dwyeri R.S. Cowan =Tachigali dwyeri 

Sclerolobium guianense Benth. =Tachigali guianensis 

Sclerolobium guianense var. guianense =Tachigali guianensis 

Sclerolobium melinonii Harms =Tachigali melinonii 

Sclerolobium micropetalum Ducke =Tachigali micropetala 

Sclerolobium odoratissimum var. latifolium Dwyer =Tachigali odoratissima 

Sclerolobium paraënsis Huber =Tachigali paraënsis 

Sclerolobium physophorum Huber =Tachigali physophora 

Sclerolobium pimichinense R.S. Cowan =Tachigali pimichinensis 

Sclerolobium reticulosum Dwyer =Tachigali reticulosa 

Sclerolobium roraimae R.H. Schomb. [nomen nudum? Mt. Roraima] 

Sclerolobium tinctorium Benth. =Tachigali tinctoria 

Senna lucens Vogel =Senna silvestris 

Senna pendula var. advena (Willd.) H.S. Irwin & Barneby =Senna pendula var. indecora 

Senna spruceana Benth. =Cassia spruceana 

Tachia paniculata (Aubl.) Pers. [orthographic variant] =Tachigali paniculata 

Tachigali carinata Gleason =Tachigali paniculata 

Tachigali glauca Tul. =Tachigali cavipes 

Tachigali paniculata var. cavipes Benth. =Tachigali cavipes 

Tachigali paniculata var. cavipes Spruce ex Benth. =Tachigali cavipes 

Tachigali purpurea Rich. (Leguminosae-Faboideae) =Diplotropis purpurea

346 


Tachigali richardiana Tul. ex Lemée [apparently a nomen nudum] 

Tachigali sulcata Benoist =Tachigali paniculata var. sulcata 

Tachigali trigona Aubl. =Tachigali paniculata 

Touchiroa aromatica Aubl. =Crudia aromatica 

Vogelocassia cobanensis Britton =Senna cobanensis 

Vouapa acaciifolia (Benth.) Baill. =Macrolobium acaciifolium 

Vouapa angustifolia Benth. =Macrolobium angustifolium 

Vouapa bifolia Aubl. =Macrolobium bifolium 

Vouapa chrysostachya Miq. =Macrolobium angustifolium 

Vouapa discolor (Benth.) Taub. =Macrolobium discolor 

Vouapa gracile (Spruce) Taub. =Macrolobium gracile 

Vouapa guianensis (Aubl.) Taub. =Macrolobium guianense 

Vouapa jenmani Gleason =Macrolobium jenmanii 

Vouapa limbata (Spruce) Taub. =Macrolobium limbatum 

Vouapa multijuga (DC.) Taub. =Macrolobium multijugum 

Vouapa punctata (Spruce) Taub. =Macrolobium punctatum 

Vouapa staminea (G. Mey.) DC. =Macrolobium bifolium 

Vouapa suaveolens (Spruce) Taub. =Macrolobium suaveolens 

Vouapa taxifolia (Spruce) Taub. =Macrolobium taxifolium 

REFERENCES 

Amshoff, G.J.H. 1939. Papilionaceae. In A. Pulle, Flora of Suriname 2(2): 1-257. 

Aymard C. 1998. Caesalpiniaceae. In J.A. Steyermark, P.E. Berry & B.K. Holst, Flora of the Venezuelan Guayana 4: 1-121. 

Barneby, R.C. & S.V. Heald. 2002. Caesalpiniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, 

& J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York 

Bot. Gard. 76 (2): 167-183. 

Barneby, R.C., B. Stergios, R.S. Cowan, P.E. Berry, J.L. Zarucchi, R.P. Wunderlin, D.M. Kearns, M.F. da Silva, A. Sprada 

Tavares, Velázquez, N. Xena, & G.A. Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators, 

Botany of the Guayana Highland part IV. Mem. New York Bot. Gard. 10(4): 65-87. 

Cowan, R.S. 1968. Swartzia. Flora Neotropica Monogr. 1: 1-228. 

Cowan, R.S. & J.C. Lindeman. 1989. Caesalpiniaceae p.p. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 7: 1-167. 

Dwyer, J.D. 1951. The Central American, West Indian, and South American species of Copaifera (Caesalpiniaceae). Brittonia 

7(3): 143-172. 

Dwyer, J.D. 1954. The tropical American genus Tachigalia Aubl. (Caesalpiniaceae). Ann. Missouri Bot. Gard. 41: 223-260. 

Dwyer, J.D. 1957. The tropical American genus Sclerolobium Vogel (Caesalpiniaceae). Lloydia 20: 67-118, suppl. 266-267. 

Dwyer, J.D. 1958. The New World species of Cynometra. Ann. Missouri Bot. Gard. 45: 313-345. 

Fonseca Vaz, A.M.S. da. 1979. Consideraçoes sobre a taxonomia do gênero Bauhinia L. sect. Tylotaea Vogel (Leguminosae - 

Caesalpinioideae) do Brasil. Rodriguésia 31(51): 127-234. 

Görts-van Rijn, A.R.A., K.U. Kramer, & J.C. Lindeman. 1976. Papilionaceae (additions and corrections). In J. Lanjouw & A.L. 

Stoffers, Flora of Suriname 2(2): 556-610. 

Irwin, H.S. & R.C. Barneby. 1982. The American Cassiinae. A synoptical revision of Leguminosae tribe Cassieae subtribe 

Cassiinae in the New World. Mem. New York Bot. Gard. 35: 1-918. 

Koeppen, R.C. 1967. Revision of Dicorynia. Brittonia 19(1): 42-61. 

Lee, Y.T. & J.H. Langenheim. 1975. Systematics of the genus Hymenaea L. Univ. Calif. Publ. Bot. 69: 1-109. 

Polhill, R.M. & P.H. Raven (ed.). 1981. Advances in Legume Systematics. Royal Botanical Garden, Kew, England. 

Silva, M.F. da. 1976. Revisao taxonômica do gênero Peltogyne Vog. Acta Amaz. 6(1) suppl.: 1-61. 

Silva, M.F. da. 1986. Dimorphandra. Flora Neotropica Monogr. 44: 1-127. 

Stergios, B. 1996. Contributions to South American Caesalpinaceae. II. A taxonomic Update of Campsiandra (Caesalpinaceae). 

Novon 6(4): 434-459. 

Velasquez, D. 1981. Revision Taxonomica del Genero Brownea Jacq. (Leguminosae-Caesalpinioideae) para Venezuela. Thesis- 

Univ. Central de Venezuela. 

LEGUMINOSAE-FABOIDEAE 

G.A. Aymard C., P.E. Berry, R.S. Cowan†, Nidia L. Cuello A., A. Delgado Salinas, P.R. Fantz, R.H. Maxwell, K.M. 

Redden,V.E. Rudd†, M. Sousa, D.R. Wind 

Abrus fruticulosus Wall. ex Wight & Arn. [introduced] 

Abrus precatorius L. [cultivated, naturalized] 

Abrus precatorius subsp. africanus Verdc. 

Abrus pulchellus Wall. ex Thwaites 

Abrus pulchellus subsp. tenuiflorus (Benth.) Verdc. 

Acosmium nitens (Vogel) Yakovlev 

Acosmium praeclarum (Sandwith) Yakovlev 

SU 


BO DA GU 

AM BO 

AM BO 


GU

Acosmium sp. A 

Aeschynomene americana L. 

Aeschynomene americana var. americana 

Aeschynomene brasiliana (Poir.) DC. 

Aeschynomene brasiliana var. brasiliana 

Aeschynomene brasiliana var. carichanica Rudd 

Aeschynomene evenia C. Wright 

Aeschynomene evenia var. serrulata Rudd 

Aeschynomene filosa Mart. ex Benth. 

Aeschynomene fluminensis Vell. 

Aeschynomene fluminensis var. fluminensis 

Aeschynomene histrix Poir. 

Aeschynomene histrix var. densiflora (Benth.) Rudd 

Aeschynomene histrix var. histrix 

Aeschynomene interrupta Benth. 

Aeschynomene paniculata Willd. ex Vogel 

Aeschynomene pratensis Small 

Aeschynomene pratensis var. caribaea Rudd 

Aeschynomene rudis Benth. 

Aeschynomene scabra G. Don 

Aeschynomene sensitiva Sw. 

Aeschynomene sensitiva var. sensitiva 

Aldina aurea R.S. Cowan 

Aldina berryi R.S. Cowan & Steyerm. 

Aldina discolor Spruce ex Benth. 

Aldina elliptica R.S. Cowan 

Aldina heterophylla Spruce ex Benth. 

Aldina insignis (Benth.) Endl. 

Aldina insignis var. insignis 

Aldina insignis var. retusa R.S. Cowan 

Aldina kunhardtiana R.S. Cowan 

Aldina latifolia Spruce ex Benth. 

Aldina latifolia var. auyantepuiensis Pittier ex H.S. Irwin 

Aldina latifolia var. latifolia [expected in GU] 

Aldina latifolia var. pubescens R.S. Cowan [expected in GU] 

Aldina macrophylla Spruce ex Benth. 

Aldina macrophylla var. macrophylla 

Aldina macrophylla var. yapacanensis (R.S. Cowan) Stergios 

Aldina paulberryi Aymard 

Aldina petiolulata R.S. Cowan 

Aldina reticulata R.S. Cowan 

Aldina sp. A 

Aldina sp. B 

Aldina sp. C 

Aldina sp. D 

Aldina sp. E 

Aldina sp. F 

Aldina sp. G 

Alexa bauhiniiflora Ducke 

Alexa canaracunensis Pittier 

Alexa confusa Pittier 

Alexa cowanii Yakovlev 

Alexa grandiflora Ducke 

Alexa herminiana N. Ramírez 

Alexa imperatricis (R.H. Schomb.) Baill. 

Alexa surinamensis Yakovlev 

Alexa wachenheimii Benoist 

Alysicarpus vaginalis (L.) DC. [introduced] 

Andira coriacea Pulle 

Andira grandistipula Amshoff 

Andira inermis (W. Wright) DC. 

Andira inermis subsp. inermis 

Andira sp. A 

Andira surinamensis (Bondt) Splitg. ex Amshoff 


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348 


Andira tervequinata R.T. Penn., Aymard & Cuello 

Andira trifoliolata Ducke 

Andira villosa Kleinhoonte 

Barbieria pinnata (Pers.) Baill. 

Bowdichia virgilioides Kunth 

Cajanus cajan (L.) Millsp. [cultivated, naturalized] 

Calopogonium caeruleum (Benth.) C. Wright 

Calopogonium mucunoides Desv. 

Calopogonium velutinum (Benth.) Amshoff 

Canavalia brasiliensis Mart. ex Benth. 

Canavalia dictyota Piper 

Canavalia dictyota var. dictyota 

Canavalia dictyota var. ferruginea Aymard & Cuello 

Canavalia ensiformis (L.) DC. [escaped from cultivation] 

Canavalia gladiata (Jacq.) DC. 

Canavalia gladiata var. gladiata [escaped from cultivation] 

Canavalia grandiflora Benth. 

Canavalia plagiosperma Piper [probably escaped from cultivation] 

Canavalia rosea (Sw.) DC. 

Canavalia sericophylla Ducke 

Candolleodendron brachystachyum (DC.) R.S. Cowan 

Centrolobium paraense Tul. 

Centrolobium paraense var. orinocense Benth. 

Centrolobium paraense var. paraense 

Centrosema angustifolium (Kunth) Benth. non (Kunth) Benth. in Mart. 

Centrosema brasilianum (L.) Benth. 

Centrosema capitatum (Rich.) Amshoff 

Centrosema macrocarpum Benth. 

Centrosema molle Mart. ex Benth. [specimens previously misidentified as 

Centrosema pubescens] 

Centrosema pascuorum Mart. ex Benth. 

Centrosema plumieri (Turpin ex Pers.) Benth. 

Centrosema tetragonolobum Schultze-Kraft & R.J. Williams 

Centrosema triquetrum (Hoffmanns. ex Benth.) Benth. 

Centrosema venosum Mart. ex Benth. 

Centrosema vexillatum Benth. 

Centrosema virginianum (L.) Benth. 

Chaetocalyx brasiliensis (Vogel) Benth. 

Chaetocalyx scandens (L.) Urb. 

Chaetocalyx scandens var. pubescens (DC.) Rudd 

Clathrotropis brachypetala (Tul.) Kleinhoonte 

Clathrotropis brunnea Amshoff 

Clathrotropis glaucophylla R.S. Cowan 

Clathrotropis macrocarpa Ducke 

Clathrotropis nitida (Benth.) Harms 

Clathrotropis paradoxa Sandwith 

Clitoria arborescens R. Br. 

Clitoria brachycalyx Harms 

Clitoria canescens Pittier ex Fantz 

Clitoria cavalcantei Fantz 

Clitoria coriacea Schery 

Clitoria dendrina Pittier 

Clitoria falcata Lam. 

Clitoria falcata var. falcata 

Clitoria guianensis (Aubl.) Benth. 

Clitoria guianensis var. guianensis 

Clitoria javitensis (Kunth) Benth. 

Clitoria javitensis var. grandifolia (Ducke) Fantz 

Clitoria javitensis var. javitensis 

Clitoria kaieteurensis Fantz 

Clitoria laurifolia Poir. 

Clitoria leptostachya Benth. 

Clitoria leptostachya var. leptostachya 

Clitoria pendens Fantz 

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Clitoria sagotii Fantz 

Clitoria sagotii var. canaliculata Fantz 

Clitoria sagotii var. sagotii 

Clitoria sagotii var. sprucei Fantz 

Clitoria simplicifolia (Kunth) Benth. 

Clitoria steyermarkii Fantz 

Clitoria ternatea L. 

Clitoria ternatea var. ternatea [cultivated or escaped from cultivation] 

Coursetia ferruginea (Kunth) Lavin 

Crotalaria goreensis Guill. & Perr. [introduced] 

Crotalaria incana L. 

Crotalaria juncea Willd. [introduced] 

Crotalaria maypurensis Kunth 

Crotalaria maypurensis var. depauperata (Mart.) Windler & S.G. Skinner 

Crotalaria maypurensis var. maypurensis 

Crotalaria micans Link 

Crotalaria nitens Kunth 

Crotalaria pallida Aiton [introduced] 

Crotalaria pilosa Mill. non Crotalaria pilosa Thunb. 1800, nec Crotalaria 

pilosa Roxb. ex Mart. 1820. 

Crotalaria quinquefolia L. [introduced] 

Crotalaria retusa L. 

Crotalaria sagittalis L. 

Crotalaria spectabilis Roth 

Crotalaria stipularia Desv. 

Crotalaria aff. velutina Benth. 

Crotalaria verrucosa L. [introduced] 

Crotalaria zanzibarica Benth. [introduced] 

Cymbosema roseum Benth. [expected in the Guianas] 

Dalbergia amazonica (Radlk. ex Köpff) Ducke 

Dalbergia ecastaphyllum (P. Browne ex L.) Taub. 

Dalbergia foliosa (Benth.) A.M. Carvalho 

Dalbergia frutescens (Vell.) Britton 

Dalbergia frutescens var. frutescens 

Dalbergia hygrophila (Mart. ex Benth.) Hoehne 

Dalbergia intermedia A.M. Carvalho 

Dalbergia inundata Spruce ex Benth. 

Dalbergia monetaria L. f. 

Dalbergia riedelii (Benth.) Sandwith 

Dalbergia sp. A 

Dalbergia sp. B 

Dalbergia sp. C 

Dalbergia sp. D 

Dalbergia spruceana Benth. 

Dalbergia subcymosa Ducke 

Derris amazonica Killip 

Derris negrensis Benth. 

Derris pterocarpus (DC.) Killip 

Desmodium adscendens (Sw.) DC. 

Desmodium affine Schltdl. 

Desmodium axillare (Sw.) DC. 

Desmodium axillare var. acutifolium (Kunze) Urb. 

Desmodium axillare var. stoloniferum (Rich. ex Poir.) B.G. Schub. 

Desmodium barbatum (L.) Benth. 

Desmodium cajanifolium (Kunth) DC. 

Desmodium campyloclados Hemsl. 

Desmodium distortum (Aubl.) J.F. Macbr. 

Desmodium guianense DC. 

Desmodium hickenianum Burkart 

Desmodium incanum DC. 

Desmodium intortum (Mill.) Urb. 

Desmodium molliculum (Kunth) DC. 

Desmodium orinocense (DC.) Cuello 

Desmodium pachyrrhizum Vogel 


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350 

Desmodium procumbens (Mill.) Hitchc. 

Desmodium procumbens var. procumbens 

Desmodium sclerophyllum Benth. 

Desmodium scorpiurus (Sw.) Desv. 

Desmodium tortuosum (Sw.) DC. [introduced] 

Desmodium triflorum (L.) DC. [introduced] 

Desmodium velutinum DC. 

Desmodium wydlerianum Urb. 

Dioclea albiflora R.S. Cowan 

Dioclea apurensis Kunth 

Dioclea coriacea Benth. 

Dioclea guianensis Benth. 

Dioclea holtiana Pittier ex R.H. Maxwell 

Dioclea huberi Ducke 

Dioclea macrantha Huber 

Dioclea macrocarpa Huber 

Dioclea malacocarpa Ducke 

Dioclea megacarpa Rolfe 

Dioclea reflexa Hook. f. 

Dioclea rigida R.S. Cowan 

Dioclea ruddiae R.H. Maxwell 

Dioclea scabra (Rich.) R.H. Maxwell 

Dioclea scabra var. brownii R.H. Maxwell 

Dioclea scabra var. scabra 

Dioclea scabra var. schulzii R.H. Maxwell 

Dioclea steyermarkii R.H. Maxwell 

Dioclea violacea Mart. ex Benth. 

Dioclea virgata (Rich.) Amshoff 

Dioclea virgata var. crenata R.H. Maxwell 

Dioclea virgata var. virgata 

Dioclea wilsonii Standl. 

Diplotropis brasiliensis (Tul.) Benth. 

Diplotropis martiusii Benth. 

Diplotropis purpurea (Rich.) Amshoff 

Diplotropis purpurea var. leptophylla (Kleinhoonte) Amshoff 

Diplotropis purpurea var. purpurea 

Diplotropis racemosa (Hoehne) Amshoff 

Diplotropis racemosa var. racemosa 

Diplotropis rigidocarpa H.C. Lima 

Diplotropis sp. A 

Diplotropis strigulosa R.S. Cowan 

Diplotropis triloba Gleason 

Dipteryx magnifica (Ducke) Ducke 

Dipteryx odorata (Aubl.) Willd. 

Dipteryx punctata (S.F. Blake) Amshoff 

Dipteryx rosea Spruce ex Benth. 

Dussia discolor (Benth.) Amshoff 

Eriosema crinitum (Kunth) G. Don 

Eriosema crinitum var. crinitum 

Eriosema crinitum var. stipulare (Benth.) Fortunato 

Eriosema rufum (Kunth) G. Don 

Eriosema rufum var. rufum 

Eriosema simplicifolium (Kunth) G. Don 

Eriosema simplicifolium var. simplicifolium 

Eriosema violaceum (Aubl.) G. Don 

Erythrina amazonica Krukoff 

Erythrina fusca Lour. 

Erythrina mitis Jacq. 

Erythrina poeppigiana (Walp.) Cook 

Erythrina rubrinervia Kunth 

Etaballia dubia (Kunth) Rudd 

Fissicalyx fendleri Benth. 

Flemingia strobilifera (L.) R. Br. [introduced] 

Galactia dubia DC. 


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Galactia gracillima Benth. 

Galactia jussiaeana Kunth 

Galactia jussiaeana var. jussiaeana 

Galactia latisiliqua Desv. 

Galactia longifolia (Jacq.) Benth. 

Galactia striata (Jacq.) Urb. 

Gliricidia sepium (Jacq.) Kunth ex Walp. [widely cultivated & naturalized] 

Hymenolobium excelsum Ducke 

Hymenolobium flavum Kleinhoonte 

Hymenolobium heterocarpum Ducke 

Hymenolobium nitidum Benth. 

Hymenolobium petraeum Ducke 

Hymenolobium pulcherrimum Ducke 

Indigofera bongardiana (Kuntze) Burkart 

Indigofera endecaphylla Jacq. 

Indigofera fruticosa L. [escaped from cultivation] 

Indigofera hirsuta L. [cultivated and escaped from cultivation] 

Indigofera lespedezioides Kunth 

Indigofera microcarpa Desv. 

Indigofera spicata Forssk. 

Indigofera suffruticosa Mill. [cultivated only?] 

Indigofera tinctoria L. [cultivated, perhaps escaped] 

Lablab purpureus (L.) Sweet [widely cultivated] 

Lonchocarpus chrysophyllus Kleinhoonte 

Lonchocarpus crassispermus Poppend. 

Lonchocarpus crucisrubierae Pittier 

Lonchocarpus densiflorus Benth. 

Lonchocarpus dipteroneurus Pittier 

Lonchocarpus fendleri Benth. 

Lonchocarpus floribundus Benth. 

Lonchocarpus hedyosmus Miq. 

Lonchocarpus heptaphyllus (Poir.) DC. 

Lonchocarpus imatacensis Poppend. 

Lonchocarpus longifolius (Benth.) Ducke 

Lonchocarpus martynii A.C. Sm. 

Lonchocarpus pictus Pittier 

Lonchocarpus punctatus Kunth 

Lonchocarpus rariflorus Mart. ex Benth. 

Lonchocarpus rufescens Benth. 

Lonchocarpus sericeus (Poir.) Kunth ex DC. [expected in the Guianas] 

Lonchocarpus spruceanus Benth. 

Lonchocarpus tubicalyx Pittier ex Poppend. 

Lonchocarpus urucu Killip & A.C. Sm. 

Lonchocarpus utilis A.C. Sm. 

Luetzelburgia sp. 

Machaerium acuminatum Kunth 

Machaerium acutifolium Vogel 

Machaerium acutifolium var. pseudacutifolium (Pittier) Rudd 

Machaerium affine Benth. 

Machaerium altiscandens Ducke 

Machaerium amazonense Hoehne 

Machaerium aristulatum (Spruce ex Benth.) Ducke 

Machaerium aureiflorum Ducke 

Machaerium dubium (Kunth) Rudd 

Machaerium ferox (Mart. ex Benth.) Ducke 

Machaerium floribundum Benth. 

Machaerium floribundum var. floribundum 

Machaerium guaremalense Pittier 

Machaerium inundatum (Mart. ex Benth.) Ducke 

Machaerium isadelphum (E. Mey.) Amshoff 

Machaerium kegelii Meisn. 

Machaerium leiophyllum (DC.) Benth. 

Machaerium leiophyllum var. crista-castrense (Mart. ex Benth.) Rudd 

Machaerium leiophyllum var. latifolium (Benth.) Rudd 

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352 

Machaerium leiophyllum var. leiophyllum 

Machaerium lunatum (L. f.) Ducke 

Machaerium macrophyllum Benth. 

Machaerium macrophyllum var. macrophyllum 

Machaerium madeirense Pittier 

Machaerium multifoliolatum Ducke 

Machaerium mutisii Killip ex Rudd 

Machaerium myrianthum Spruce ex Benth. 

Machaerium paraense Ducke 

Machaerium polyphyllum (Poir.) Benth. 

Machaerium quinatum (Aubl.) Sandwith 

Machaerium quinatum var. parviflorum (Benth.) Rudd 

Machaerium quinatum var. quinatum 

Machaerium robiniifolium (DC.) Vogel 

Machaerium sp. A 

Machaerium tovarense Pittier 

Machaerium trifoliolatum Ducke 

Macroptilium atropurpureum (Sessé & Moc. ex DC.) Urb. 

Macroptilium gracile (Poepp. ex Benth.) Urb. 

Macroptilium lathyroides (L.) Urb. 

Macroptilium lathyroides var. lathyroides 

Macroptilium longipedunculatum (Mart. ex Benth.) Urb. 

Macroptilium monophyllum (Benth.) Maréchal & Baudet 

Monopteryx angustifolia Spruce ex Benth. 

Monopteryx inpae W.A. Rodrigues 

Monopteryx sp. A 

Monopteryx uaucu Spruce ex Benth. 

Mucuna pruriens (L.) DC. [cultivated, naturalized] 

Mucuna pruriens var. utilis (Wall. ex Wight) Baker ex Burck 

Mucuna rostrata Benth. 

Mucuna sloanei Fawc. & Rendle 

Mucuna urens (L.) Medik. 

Muellera frutescens (Aubl.) Standl. 

Myrocarpus venezuelensis Rudd 

Myrospermum frutescens Jacq. 

Myroxylon balsamum (L.) Harms [cultivated?] 

Myroxylon peruiferum L. f. 

Ormosia amazonica Ducke 

Ormosia bolivarensis (Rudd) Stirton 

Ormosia cinerea Benoist 

Ormosia coarctata Jacks. 

Ormosia coccinea (Aubl.) Jacks. 

Ormosia costulata (Miq.) Kleinhoonte 

Ormosia coutinhoi Ducke 

Ormosia discolor Spruce ex Benth. 

Ormosia flava (Ducke) Rudd 

Ormosia grandiflora (Tul.) Rudd 

Ormosia guyanensis C.H. Stirt., sp. nov. ined. 

Ormosia lignivalvis Rudd 

Ormosia macrocalyx Ducke 

Ormosia macrophylla Benth. 

Ormosia maguireorum Rudd 

Ormosia melanocarpa Kleinhoonte 

Ormosia nobilis Tul. 

Ormosia nobilis var. nobilis 

Ormosia paraensis Ducke 

Ormosia smithii Rudd 

Ormosia sp. A 

Ormosia sp. B 

Ormosia steyermarkii Rudd 

Ormosia stipularis Ducke 

Ormosia subsimplex Spruce ex Benth. 

Ormosia williamsii Rudd 

Pachyrhizus erosus (L.) Urb. 


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Paramachaerium ormosioides (Ducke) Ducke 

Paramachaerium schomburgkii (Benth.) Ducke 

Phaseolus lunatus L. [cultivated, naturalized] 

Phaseolus reptans Ducke 

Phaseolus vulgaris L. [probably cultivated only] 

Phaseolus vulgaris var. vulgaris 

Piscidia carthaginensis Jacq. 

Platymiscium filipes Benth. 

Platymiscium pinnatum (Jacq.) Dugand 

Platymiscium trinitatis Benth. 

Platymiscium ulei Harms ex Harms 

Platypodium elegans Vogel 

Platypodium elegans subsp. maxonianum (Pittier) H.C. Lima 

Poecilanthe amazonica (Ducke) Ducke 

Poecilanthe effusa (Huber) Ducke 

Poecilanthe hostmannii (Benth.) Amshoff 

Poecilanthe ovalifolia Kleinhoonte 

Poiretia punctata (Willd.) Desv. 

Pterocarpus acapulcensis Rose 

Pterocarpus amazonum (Benth.) Amshoff 

Pterocarpus officinalis Jacq. 

Pterocarpus officinalis subsp. officinalis 

Pterocarpus rohrii Vahl 

Pterocarpus santalinoides L’Hér. ex DC. 

Pueraria phaseoloides (Roxb.) Benth. [naturalized] 

Rhynchosia edulis Griseb. 

Rhynchosia melanocarpa Grear [perhaps expected in the Guianas] 

Rhynchosia minima (L.) DC. 

Rhynchosia phaseoloides (Sw.) DC. 

Rhynchosia reticulata (Sw.) DC. 

Rhynchosia schomburgkii Benth. 

Sesbania emerus (Aubl.) Urb. 

Sesbania exasperata Kunth 

Sesbania grandiflora (L.) Pers. [cultivated, occasionally escaping] 

Sesbania sericea (Willd.) DC. [introduced] 

Sesbania sesban (L.) Merr. [introduced] 

Singana guianensis Aubl. [name of uncertain application] 

Soemmeringia semperflorens Mart. 

Spirotropis longifolia (DC.) Baill. 

Spirotropis sp. A 

Stylosanthes angustifolia Vogel 

Stylosanthes capitata Vogel 

Stylosanthes cayennensis Mohlenbr. [=Stylosanthes guianensis?] 

Stylosanthes guianensis (Aubl.) Sw. 

Stylosanthes guianensis var. gracilis (Kunth) Vogel 

Stylosanthes guianensis var. guianensis 

Stylosanthes hamata (L.) Taub. 

Stylosanthes hispida Rich. [=Stylosanthes guianensis?] 

Stylosanthes humilis Kunth 

Stylosanthes suffruticosa Mohlenb. 

Stylosanthes viscosa (L.) Sw. 

Swartzia alato-sericea Barneby [expected in AM] 

Swartzia amshoffiana R.S. Cowan 

Swartzia angustifoliola Schery 

Swartzia anomala R.S. Cowan 

Swartzia apiculata R.S. Cowan 

Swartzia aptera DC. 

Swartzia arborescens (Aubl.) Pittier 

Swartzia argentea Spruce ex Benth. 

Swartzia argentea var. argentea 

Swartzia aymardii Barneby 

Swartzia bannia Sandwith 

Swartzia barnebyana Cuello 

Swartzia benthamiana Miq. 

FG 

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354 


Swartzia benthamiana var. benthamiana 

Swartzia benthamiana var. yatuensis R.S. Cowan 

Swartzia brachyrachis Harms 

Swartzia brachyrachis var. glabrata R.S. Cowan 

Swartzia buntingii R.S. Cowan 

Swartzia cardiosperma Spruce ex Benth. 

Swartzia caudata R.S. Cowan 

Swartzia coarctata Jacks. 

Swartzia conferta Spruce ex Benth. 

Swartzia conferta var. conferta 

Swartzia cowanii Steyerm. 

Swartzia cupavenensis R.S. Cowan 

Swartzia cuspidata Spruce ex Benth. 

Swartzia davisii Sandwith 

Swartzia dipetala Willd. ex Vogel 

Swartzia eriocarpa Benth. 

Swartzia fanshawei R.S. Cowan 

Swartzia floribunda Spruce ex Benth. 

Swartzia grandifolia Bong. ex Benth. 

Swartzia guianensis (Aubl.) Urb. 

Swartzia hostmannii Benth. 

Swartzia invenusta Barneby 

Swartzia jenmanii Sandwith 

Swartzia kaieteurensis (R.S. Cowan) B.M. Torke, sp. nov. ined. 

Swartzia katawa R.S. Cowan 

Swartzia laevicarpa Amshoff 

Swartzia lamellata Ducke 

Swartzia lamellata var. kaieteurensis R.S. Cowan 

Swartzia lamellata var. lamellata 

Swartzia latifolia Benth. 

Swartzia latifolia var. latifolia 

Swartzia latifolia var. sylvestris R.S. Cowan 

Swartzia laurifolia Benth. [expected in GU] 

Swartzia laxiflora Bong. ex Benth. [expected in the Guianas] 

Swartzia leblondii R.S. Cowan 

Swartzia leiocalycina Benth. 

Swartzia leiogyne (Sandwith) R.S. Cowan 

Swartzia leptopetala Benth. 

Swartzia longicarpa Amshoff 

Swartzia longipedicellata Sandwith 

Swartzia macrocarpa Spruce ex Benth. 

Swartzia maguirei R.S. Cowan 

Swartzia microcarpa Spruce ex Benth. 

Swartzia oblanceolata Sandwith 

Swartzia oblonga Benth. [expected in GU] 

Swartzia oedipus Barneby 

Swartzia pachyphylla Harms [Mt. Roraima] 

Swartzia palustris Barneby 

Swartzia panacoco (Aubl.) R.S. Cowan 

Swartzia panacoco var. altsonii (Sandwith) R.S. Cowan 

Swartzia panacoco var. cardonae (R.S. Cowan) R.S. Cowan 

Swartzia panacoco var. kamarangensis R.S. Cowan 

Swartzia panacoco var. panacoco 

Swartzia panacoco var. polyanthera (Steud.) R.S. Cowan 

Swartzia panacoco var. sagotii (Sandwith) R.S. Cowan 

Swartzia panacoco var. sandwithiana (Cowan) R.S. Cowan 

Swartzia panacoco var. tepuiensis (Schery) R.S. Cowan [expected in GU] 

Swartzia parvifolia Schery [expected in GU] 

Swartzia pendula Spruce ex Benth. 

Swartzia piarensis R.S. Cowan [expected in GU] 

Swartzia picta Spruce ex Benth. 

Swartzia picta var. bolivarensis R.S. Cowan [expected in GU, possibly in SU] 

Swartzia picta var. picta 

Swartzia pinnata (Vahl) Willd. 

AM GU SU FG 

AM 

BO GU SU 

BO GU SU 

AM 

AM 

AM 

FG? 

AM BO GU 

AM BO GU 

BO 

AM 

AM 

GU 

AM BO GU 

GU 

GU 

AM 


AM GU SU FG 

SU FG 

GU 

BO GU 

GU 

GU 


GU 

GU 

GU 

GU SU 

GU 

GU SU 

AM 

GU 

AM GU 

AM BO 

AM 

AM 

AM 

AM 

BO 

GU 

FG 

SU FG 

GU SU FG 

AM 


GU 

AM BO GU 

GU 

FG 

GU SU FG 

GU? FG 

GU 

BO 

AM BO 

AM 

BO 

AM BO 

BO 

AM BO 

AM


Swartzia pittieri Schery 

Swartzia polyphylla DC. 

Swartzia recurva Poepp. [expected in GU?] 

Swartzia rediviva R.S. Cowan 

Swartzia remiger Amshoff 

Swartzia reticulata Ducke 

Swartzia roraimae Sandwith [Mt. Roraima] 

Swartzia schomburgkii Benth. 

Swartzia sericea Vogel 

Swartzia sericea var. sericea 

Swartzia sp. A 

Swartzia sp. B 

Swartzia sp. C 

Swartzia sp. D 

Swartzia sp. E 

Swartzia sp. F 

Swartzia sprucei Benth. 

Swartzia sprucei var. sprucei 

Swartzia sprucei var. tessellata R.S. Cowan 

Swartzia steyermarkii R.S. Cowan 

Swartzia stipellata R.S. Cowan [expected in GU] 

Swartzia tessmannii Harms [expected in GU] 

Swartzia tillettii R.S. Cowan 

Swartzia triptera Barneby 

Swartzia vaupesiana R.S. Cowan 

Swartzia wurdackii R.S. Cowan [expected in GU] 

Swartzia xanthopetala Sandwith 

Taralea cordata Ducke 

Taralea cordata var. cordata 

Taralea cordata var. rigida (Schery) H.C. Lima 

Taralea crassifolia (Spruce ex Benth.) Ducke 

Taralea nudipes (Tul.) Ducke 

Taralea oppositifolia Aubl. 

Taralea reticulata (Benth.) Ducke 

Tephrosia adunca Benth. 

Tephrosia candida DC. [introduced] 

Tephrosia cinerea (L.) Pers. 

Tephrosia purpurea (L.) Pers. 

Tephrosia senna Kunth 

Tephrosia sessiliflora (Poir.) Hassl. 

Tephrosia sinapou (Buc’hoz) A. Chev. 

Teramnus labialis Spreng. [introduced?] 

Teramnus uncinatus (L.) Sw. 

Teramnus uncinatus subsp. uncinatus 

Vatairea erythrocarpa (Ducke) Ducke 

Vatairea guianensis Aubl. 

Vatairea paraensis Ducke 

Vataireopsis speciosa Ducke [records may represent misidentifications] 

Vataireopsis surinamensis H.C. Lima 

Vigna adenantha (G. Mey.) Maréchal, Mascherpa & Stainier 

Vigna candida (Vell.) Maréchal, Mascherpa & Stainier 

Vigna hosei (Craib) Backer [cultivated, naturalized] 

Vigna juruana (Harms) Verdc. 

Vigna lasiocarpa (Mart. ex Benth.) Verdc. 

Vigna linearis (Kunth) Maréchal, Mascherpa & Stainier 

Vigna linearis var. linearis 

Vigna longifolia (Benth.) Verdc. 

Vigna luteola (Jacq.) Benth. 

Vigna peduncularis (Kunth) Fawc. & Rendle 

Vigna peduncularis var. peduncularis 

Vigna trichocarpa (C. Wright) A. Delgado-Salinas 

Vigna unguiculata (L.) Walp. [cultivated, naturalized] 

Vigna vexillata (L.) A. Rich. 

Vigna vexillata var. vexillata 

AM BO 


SU 

GU SU FG? 

GU SU 

BO GU 

AM BO GU SU 

AM BO GU? FG 

AM BO GU? FG 

DA GU SU 

AM 

AM 

AM 

AM 

AM 

AM BO GU 

AM BO 

BO GU 

BO GU SU? FG 

AM 

AM BO 

GU 

AM 

AM 

BO 

GU 


AM GU SU FG 

AM BO 


FG 

AM GU SU FG 

BO GU 

BO GU SU FG 

SU 


GU SU 

BO 



GU 

BO GU SU FG 

BO GU SU FG 

FG 


BO DA SU FG 

AM GU SU FG 

AM GU SU FG 


AM BO 

SU FG 





AM DA FG 

DA GU SU FG 

AM BO SU FG 

AM BO 

GU SU FG 

GU SU FG 


AM DA GU SU FG

356 

Zollernia grandifolia Schery 

Zollernia paraënsis Huber 

Zornia brasiliensis Vogel 

Zornia curvata Mohlenbr. 

Zornia diphylla (L.) Pers. 

Zornia gemella (Willd.) Vogel 

Zornia glabra Desv. 

Zornia gracilis DC. 

Zornia guanipensis Pittier 

Zornia herbacea Pittier 

Zornia lasiocarpa A.R. Molina 

Zornia latifolia Sm. 

Zornia latifolia var. latifolia 

Zornia myriadena Benth. 

Zornia pardina Mohlenbr. 

Zornia pardina var. vichadana Killip ex Mohlenbr. 

Zornia reticulata Sm. 

Zornia sericea Moric. 


BO 

AM GU SU FG 

BO 

AM BO GU 

AM BO GU 

BO GU SU FG 

GU SU FG 

FG 

BO SU 

BO 

BO 



BO 

GU 

GU 

AM BO GU 

AM BO 


Abrus minor Desv. =Abrus precatorius subsp. africanus 

Abrus tenuiflorus Benth. =Abrus pulchellus subsp. tenuiflorus 

Acouroa violacea Aubl. ? =Dalbergia sp. 

Aeschynomene biflora Fawc. & Rendle =Aeschynomene brasiliana 

Aeschynomene conferta Benth. =Aeschynomene histrix var. histrix 

Aeschynomene densiflora Benth. =Aeschynomene histrix var. densiflora 

Aeschynomene emerus Aubl. =Sesbania emerus 

Aeschynomene gracilis Miq. =Aeschynomene paniculata 

Aeschynomene guarica Pittier =Aeschynomene brasiliana 

Aeschynomene hispida Willd. [not in flora area] =Aeschynomene virginica (L.) Britton, Poggenb. & Sterns. 

Aeschynomene viscidula Michx. [not in flora area] 

Agati grandiflora (L.) Desv. =Sesbania grandiflora 

Aldina yapacanensis R.S. Cowan =Aldina macrophylla var. yapacanensis 

Alexa imperatricis auct. non R.H. Schomb. 1845, nec Baill. 1870. =Alexa confusa 

Alexa leiopetala Sandwith =Alexa imperatricis 

Alexa superba R.S. Cowan =Alexa confusa 

Alexandra imperatricis R.H. Schomb. =Alexa imperatricis 

Allania insignis Benth. =Aldina insignis 

Amerimnon brownei Jacq. =Dalbergia ecastaphyllum 

Amerimnon latifolium Willd. =Lonchocarpus heptaphyllus 

Amerimnon pinnatum Jacq. =Platymiscium pinnatum 

Amphiodon effusus Huber =Poecilanthe effusa 

Andira retusa (Poir.) Kunth =Andira surinamensis 

Anila bongardiana Kuntze =Indigofera bongardiana 

Barbieria polyphylla (Poir.) DC. =Barbieria pinnata 

Bowdichia brasiliensis (Tul.) Ducke =Diplotropis brasiliensis 

Bowdichia guianensis Ducke =Diplotropis purpurea 

Bowdichia martiusii (Benth.) Ducke =Diplotropis martiusii 

Bowdichia racemosa Hoehne =Diplotropis racemosa 

Bradburya angustifolium (Kunth) Kuntze =Centrosema brasilianum 

Bradburya brasilianum (L.) Kuntze =Centrosema brasilianum 

Bradburya macrocarpa (Benth.) Kuntze =Centrosema macrocarpum 

Bradburya pascuora (Mart. ex Benth.) Kuntze =Centrosema pascuorum 

Bradburya plumieri (Turpin ex Pers.) Kuntze =Centrosema plumieri 

Bradburya triquetra (Hoffmanns. ex Benth.) Kuntze =Centrosema triquetrum 

Bradburya venosum (Mart. ex Benth.) Kuntze =Centrosema venosum 

Bradburya virginianum (L.) Kuntze =Centrosema virginianum 

Cajanus indicus Spreng. =Cajanus cajan 

Canavalia bracteolata Merr. =Dioclea macrantha var. bracteolata 

Canavalia fendleri Piper =Canavalia brasiliensis 

Canavalia maritima Thouars =Canavalia rosea 

Canavalia obtusifolia DC. =Canavalia rosea 

Centrolobium orinocense (Benth.) Pittier =Centrolobium paraense var. orinocense 

Centrolobium patinense Pittier =Centrolobium paraense var. orinocense


Centrosema angustifolium (Kunth) Benth. in Mart., 1859 =Centrosema brasilianum 

Centrosema brasilianum var. angustifolium (Kunth) Amshoff =Centrosema brasilianum 

Centrosema latissimum Ducke =Centrosema triquetrum 

Centrosema macrocarpon auct. [orthographic error] =Centrosema macrocarpum 

Centrosema platycarpum Benth. [not in flora area] 

Centrosema pubescens auct. non Benth. 1837 [sensu Benth. in Mart., fide Fantz] =Centrosema molle 

Chaetocalyx fissa Pittier =Chaetocalyx scandens var. pubescens 

Chaetocalyx magniflora Pittier =Chaetocalyx scandens var. pubescens 

Chaetocalyx nigrescens Pittier =Chaetocalyx scandens var. pubescens 

Chaetocalyx paucifolia Pittier =Chaetocalyx scandens var. pubescens 

Chaetocalyx perglandulosa Pittier =Chaetocalyx scandens var. pubescens 

Chaetocalyx pubescens DC. =Chaetocalyx scandens var. pubescens 

Chaetocalyx vestita Standl. =Chaetocalyx scandens var. pubescens 

Clathrotropis colombiana Yakovlev =Clathrotropis brunnea 

Clathrotropis flava Ducke =Ormosia flava 

Clathrotropis surinamensis Kleinhoonte =Ormosia flava 

Clitoria amoena Miq. =Clitoria arborescens 

Clitoria angustifolia Kunth =Centrosema brasilianum 

Clitoria angustifolia auct. non Kunth 1823 [1824] =Centrosema brasilianum 

Clitoria arborea Mart. ex Benth. [not in flora area] 

Clitoria brasiliana L. =Centrosema brasilianum 

Clitoria brasiliensis L. =Centrosema brasilianum 

Clitoria cajanifolia (C. Presl.) Benth. =Clitoria laurifolia 

Clitoria capitata Rich. =Centrosema capitatum 

Clitoria cerifera R.S. Cowan =Clitoria coriacea 

Clitoria glycinoides DC. =Clitoria falcata 

Clitoria grandifolia Ducke =Clitoria javitensis var. grandifolia 

Clitoria guyanensis Benth. =Clitoria guianensis 

Clitoria guyanensis Aubl. ex Desv. =Clitoria laurifolia 

Clitoria javitensis var. glabra Sagot =Clitoria sagotii 

Clitoria parvifolia Pittier [nomen nudum] =Clitoria laurifolia 

Clitoria pinnata (Pers.) R.H. Sm. & G.P. Lewis =Barbieria pinnata 

Clitoria plumieri Turpin ex Pers. =Centrosema plumieri 

Clitoria poitaei DC. =Clitoria arborescens 

Clitoria poiteai DC. =Clitoria arborescens 

Clitoria poiteai Benth. =Clitoria arborescens 

Clitoria polyphylla Poir. =Barbieria pinnata 

Clitoria rubiginosa Juss. ex Pers. =Clitoria falcata 

Clitoria virginiana L. =Centrosema virginianum 

Collaea rosea Benth. =Galactia jussiaeana 

Coronilla scandens L. =Chaetocalyx scandens 

Coronilla sericea Willd. =Sesbania sericea 

Coublandia frutescens Aubl. =Muellera frutescens 

Coumarouna magnifica Ducke =Dipteryx magnifica 

Coumarouna odorata Aubl. =Dipteryx odorata 

Coumarouna oppositifolia (Aubl.) Taub. =Taralea oppositifolia 

Coumarouna punctata S.F. Blake =Dipteryx punctata 

Coursetia arborea Griseb. =Coursetia ferruginea 

Crotalaria anagyroides Kunth =Crotalaria micans 

Crotalaria anagyroides var. pauciflora Griseb. =Crotalaria maypurensis 

Crotalaria bracteata Schltdl. & Cham. =Crotalaria nitens 

Crotalaria cuneifolia (Forssk.) Schrank =Crotalaria retusa 

Crotalaria depauperata Mart. =Crotalaria maypurensis var. depauperata 

Crotalaria gajanensis Desv. =Clitoria laurifolia 

Crotalaria guianensis Aubl. =Clitoria guianensis 

Crotalaria leptophylla Benth. =Crotalaria maypurensis 

Crotalaria paramariboensis Miq. =Crotalaria verrucosa 

Crotalaria pilosa var. skutchii Senn =Crotalaria pilosa 

Crotalaria pterocaula Desv. =Crotalaria pilosa 

Crotalaria retusifolia Stokes =Crotalaria retusa 

Crotalaria saltiana Andrews =Crotalaria pallida 

Crotalaria schiedeana Steud. =Crotalaria nitens 

Cyclolobium amazonicum Ducke =Poecilanthe amazonica 

Cyclolobium hostmannii Benth. =Poecilanthe hostmannii

358 


Cymbosema apurense (Kunth) Pittier =Dioclea apurensis 

Cymbosema apurense (Kunth) Pittier, sensu Pitter Bol. Soc. Venez. Ci. Nat. 7: 154. =Cymbosema roseum 

Cytisus cajan L. =Cajanus cajan 

Cytisus sessiliflorus Poir. =Tephrosia sessiliflora 

Cytisus violaceous Aubl. =Eriosema violaceum 

Cytisus violaceus Aubl. =Eriosema violaceum 

Dalbergia atropurpurea Ducke =Dalbergia foliosa 

Dalbergia aturensis Pittier =Dalbergia inundata 

Dalbergia brownei (Jacq.) Urb. =Dalbergia ecastaphyllum 

Dalbergia cucullata Pittier (Leguminosae-Caesalpinioideae) =Apuleia leiocarpa 

Dalbergia glauca (Desv.) Amshoff =Dalbergia foliosa 

Dalbergia heptaphylla Poir. =Lonchocarpus heptaphyllus 

Dalbergia pachycarpa Ducke =Dalbergia riedelii 

Dalbergia pentaphylla Poir. =Lonchocarpus heptaphyllus 

Dalbergia tomentosa (Spruce ex Benth.) Ducke =Dalbergia intermedia 

Dalbergia variabilis Vogel =Dalbergia frutescens 

Deguelia scandens Aubl. =Derris pterocarpus 

Derris ernestii (Harms) Ducke =Lonchocarpus hedyosmus 

Derris floribundus (Benth.) Ducke =Lonchocarpus floribundus 

Derris glabrescens (Benth.) J.F. Macbr. =Lonchocarpus densiflorus 

Derris guianensis Benth. =Derris pterocarpus 

Derris guyanensis Benth. =Derris pterocarpus 

Derris hedyosma (Miq.) J.F. Macbr. =Lonchocarpus hedyosmus 

Derris latifolia (Kunth ex DC.) Ducke =Lonchocarpus heptaphyllus 

Derris longifolia Benth. =Lonchocarpus longifolius 

Derris nicou (Aubl.) J.F. Macbr. =Lonchocarpus utilis 

Derris scandens (Aubl.) Pittier =Derris pterocarpus 

Derris urucu (Killip & A.C. Sm.) J.F. Macbr. =Lonchocarpus urucu 

Derris utilis (A.C. Sm.) Ducke =Lonchocarpus utilis 

Desmodium adscendens var. orinocense DC. =Desmodium orinocense 

Desmodium asperum (Poir.) Desv. =Desmodium distortum 

Desmodium axillare var. sintenisii Urb. =Desmodium axillare var. stoloniferum 

Desmodium canum (J.F. Gmel.) Schinz & Thell. =Desmodium incanum 

Desmodium dubium Pittier =Desmodium campyloclados 

Desmodium flexuosum Pittier =Desmodium campyloclados 

Desmodium frutescens (Jacq.) Schindl. =Desmodium incanum 

Desmodium molle (Vahl) DC. [not in flora area] =Desmodium glabrum (Mill.) DC. 

Desmodium spectabilis Miq. =Desmodium distortum 

Desmodium supinum DC. =Desmodium incanum 

Dibrachion brasiliensis Tul. =Diplotropis brasiliensis 

Dibrachion guianense Tul. =Diplotropis purpurea 

Dibrachion riparium Spruce ex Benth. =Diplotropis martiusii 

Dioclea broadwayana Pittier [nomen nudum] =Dioclea guianensis 

Dioclea elliptica R.H. Maxwell [nomen nudum] =Dioclea scabra var. scabra 

Dioclea glabra auct. non Benth.1837 =Dioclea scabra var. scabra 

Dioclea guianensis var. villosior Benth. =Dioclea guianensis 

Dioclea holtiana Pittier [nomen nudum] =Dioclea holtiana Pittier ex R.H. Maxwell 

Dioclea lasiocarpa Mart. =Dioclea virgata 

Diplotropis brachypetala Tul. =Clathrotropis brachypetala 

Diplotropis grandiflora Tul. =Ormosia grandiflora 

Diplotropis guianensis Benth. =Diplotropis purpurea 

Diplotropis leptophylla Kleinhoonte =Diplotropis purpurea var. leptophylla 

Diplotropis nitida Benth. =Clathrotropis nitida 

Diplotropis racemosa var. kaieteurensis Amshoff =Diplotropis racemosa var. racemosa 

Dipteryx applanata Benth. =Taralea oppositifolia 

Dipteryx cordata (Ducke) R.S. Cowan =Taralea cordata 

Dipteryx crassifolia Spruce ex Benth. =Taralea crassifolia 

Dipteryx nudipes Tul. =Taralea nudipes 

Dipteryx oppositifolia (Aubl.) Willd. =Taralea oppositifolia 

Dipteryx oppositifolia var. parviflora Benth. =Taralea oppositifolia 

Dipteryx phaeophylla Steyerm. =Spirotropis longifolia 

Dipteryx reticulata Benth. =Taralea reticulata 

Dipteryx rigida (Schery) R.S. Cowan =Taralea cordata var. rigida 

Dipteryx trifoliolata Ducke =Dipteryx punctata


Dolicholus kuntzei Harms ex Kuntze =Rhynchosia reticulata 

Dolicholus phaseoloides (Sw.) DC. =Rhynchosia phaseoloides 

Dolicholus pittieri Standl. =Rhynchosia schomburgkii 

Dolichos altissmus Jacq. =Mucuna urens 

Dolichos cuneifolius Forssk. =Crotalaria retusa 

Dolichos ensiformis L. =Canavalia ensiformis 

Dolichos erosus L. =Pachyrhizus erosus 

Dolichos gladiatus Jacq. =Canavalia gladiata 

Dolichos lablab L. =Lablab purpureus 

Dolichos luteolus Jacq. =Vigna luteola 

Dolichos maritimus Aubl. =Canavalia rosea 

Dolichos minimus L. =Rhynchosia minima 

Dolichos pruriens L. =Mucuna pruriens 

Dolichos roseus Sw. =Canavalia rosea 

Dolichos scabra Rich. =Dioclea scabra 

Dolichos uncinatus L. =Teramnus uncinatus 

Dolichos unguiculatus L. =Vigna unguiculata 

Dolichos urens L. =Mucuna urens 

Dolichos virgatus Rich. =Dioclea virgata 

Drepanocarpus aristulatus Spruce ex Benth. =Machaerium aristulatum 

Drepanocarpus crista-castrensis var. latifolius Benth. =Machaerium leiophyllum var. latifolium 

Drepanocarpus dubius Kunth =Machaerium dubium 

Drepanocarpus ferox Mart. ex Benth. =Machaerium ferox 

Drepanocarpus ferox ß macrophyllum Benth. =Machaerium ferox 

Drepanocarpus frondosus Mart. ex Benth. =Machaerium leiophyllum var. latifolium 

Drepanocarpus inundatus Mart. ex Benth. =Machaerium inundatum 

Drepanocarpus isadelphus E. Mey. =Machaerium isadelphum 

Drepanocarpus lunatus (L. f.) G. Mey. =Machaerium lunatum 

Drepanocarpus macrophyllus Ducke =Machaerium macrophyllum 

Drepanocarpus ? ovalifolius Pittier =Machaerium floribundum 

Drepanocarpus paludicola Standl. =Dalbergia inundata 

Drepanocarpus venezuelensis Pittier =Machaerium floribundum 

Dussia cayennensis Harms =Dussia discolor 

Ecastaphyllum amazonicum Radlk. ex Köpff =Dalbergia amazonica 

Ecastaphyllum benthamianum Miq. =Dalbergia monetaria 

Ecastaphyllum foliosum Benth. =Dalbergia foliosa 

Ecastaphyllum glaucum Desv. =Dalbergia foliosa 

Ecastaphyllum hygrophilum Mart. ex Benth. =Dalbergia hygrophila 

Ecastaphyllum monetaria (L. f.) DC. =Dalbergia monetaria 

Ecastaphyllum monetaria var. riedelii Benth. =Dalbergia riedelii 

Ecastaphyllum pubescens DC. =Dalbergia subcymosa 

Ecastaphyllum riedeli (Benth.) Radlk. =Dalbergia riedelii 

Ecastaphyllum tomentosum Spruce ex Benth. =Dalbergia intermedia 

Eriosema edule (Griseb.) Burkhart =Rhynchosia edulis 

Eriosema flaviflorum Miq. =Eriosema violaceum 

Eriosema fusiforme Rusby [not in flora area] =Eriosema crinitum var. fusiforme (Rusby) Grear 

Eriosema lanceolatum Benth. =Eriosema simplicifolium 

Eriosema simplicifolium var. micranthum Grear =Eriosema simplicifolium 

Eriosema stipulare Benth. =Eriosema crinitum var. stipulare 

Eriosema volubile Micheli =Rhynchosia edulis 

Erythrina corallodendrum L. [not in flora area] 

Erythrina glauca Willd. =Erythrina fusca 

Etaballia guianensis Benth. =Etaballia dubia 

Galactia angustifolia Kunth =Galactia jussiaeana 

Galactia camporum Sprague =Galactia jussiaeana 

Galactia jussiaeana var. angustifolia (Kunth) Burkart =Galactia jussiaeana 

Galactia pinnata Pers. =Barbieria pinnata 

Galactia striata var. tenuiflora (Klein ex Willd.) Burkart =Galactia striata 

Galactia tenuiflora (Klein ex Willd.) Wight & Arn. pro parte, sensu Benth. in Mart. =Galactia striata 

Galactia velutina Benth. =Galactia latisiliqua 

Galega cathartica Sessé & Moc. =Tephrosia senna 

Galega cinerea L. =Tephrosia cinerea 

Galega sinapou Buc’hoz =Tephrosia sinapou 

Galega toxicaria Sw. =Tephrosia sinapou

360 


Geoffroea discolor Benth. =Dussia discolor 

Geoffroea inermis W. Wright =Andira inermis 

Geoffroea pubescens Rich. =Andira surinamensis 

Geoffroea retusa Poir. =Andira surinamensis 

Geoffroea surinamensis Bondt =Andira surinamensis 

Geoffroea violacea (Aubl.) Pers. [=Dalbergia sp.?] =Acouroa violacea 

Glycine crinita Kunth =Eriosema crinitum 

Glycine phaseoloides Sw. =Rhynchosia phaseoloides 

Glycine punctata Willd. =Poiretia punctata 

Glycine reflexa Nutt. =Rhynchosia minima 

Glycine reticulata Sw. =Rhynchosia reticulata 

Glycine rufa Kunth =Eriosema rufum 

Glycine simplicifolia Kunth =Eriosema simplicifolium 

Glycine striata Jacq. =Galactia striata 

Glycine tenuiflora Klein ex Willd. =Galactia striata 

Hecastophyllum dubium Kunth =Etaballia dubia 

Hedysarum adscendens auct. non Sw. 1788 =Desmodium orinocense 

Hedysarum adscendens Sw. =Desmodium adscendens 

Hedysarum asperum Poir. =Desmodium distortum 

Hedysarum axillare Sw. =Desmodium axillare 

Hedysarum barbatum L. =Desmodium barbatum 

Hedysarum brasilianum Poir. =Aeschynomene brasiliana 

Hedysarum cajanifolium Kunth =Desmodium cajanifolium 

Hedysarum cajanifoliun Kunth =Desmodium cajanifolium 

Hedysarum canum J.F. Gmel. =Desmodium incanum 

Hedysarum diphyllum L. =Zornia diphylla 

Hedysarum distortum Aubl. =Desmodium distortum 

Hedysarum ecastaphyllum P. Browne ex L. =Dalbergia ecastaphyllum 

Hedysarum gemellum Willd. =Zornia gemella 

Hedysarum hamatum L. =Stylosanthes hamata 

Hedysarum hamatum var. viscosum L. =Stylosanthes viscosa 

Hedysarum incanum Sw. =Desmodium incanum 

Hedysarum procumbens Mill. =Desmodium procumbens 

Hedysarum scorpiurus Sw. =Desmodium scorpiurus 

Hedysarum stoloniferum Rich. ex Poir. =Desmodium axillare var. stoloniferum 

Hedysarum tortuosum Sw. =Desmodium tortuosum 

Hedysarum triflorum L. =Desmodium triflorum 

Hedysarum vaginale L. =Alysicarpus vaginalis 

Humboldtiella ferruginea (Kunth) Harms =Coursetia ferruginea 

Indigofera anil L. =Indigofera suffruticosa 

Indigofera pascuorum Benth. =Indigofera lespedezioides 

Leptolobium costulatum Miq. =Ormosia costulata 

Leptolobium nitens Vogel =Acosmium nitens 

Leptolobium nitidum Miq. =Acosmium nitens 

Lonchocarpus benthamianus Pittier =Lonchocarpus punctatus 

Lonchocarpus boliviensis Pittier =Lonchocarpus densiflorus 

Lonchocarpus discolor Huber =Lonchocarpus heptaphyllus 

Lonchocarpus ernestii Harms =Lonchocarpus hedyosmus 

Lonchocarpus fendleri subsp. pubescens Pittier =Lonchocarpus fendleri 

Lonchocarpus glabrescens Benth. =Lonchocarpus densiflorus 

Lonchocarpus guaricensis Pittier =Piscidia carthaginensis 

Lonchocarpus killipii Ducke =Derris negrensis 

Lonchocarpus latifolius Kunth ex DC. =Lonchocarpus heptaphyllus 

Lonchocarpus margaritensis Pittier =Lonchocarpus hedyosmus 

Lonchocarpus negrensis Benth. non Derris negrensis Benth. 1860 =Derris amazonica 

Lonchocarpus nicou (Aubl.) DC. =Lonchocarpus utilis 

Lonchocarpus nicou auct. non (Aubl.) DC. sensu Killip & A.C. Sm. 1825 =Lonchocarpus utilis 

Lonchocarpus nitidulus Benth. =Lonchocarpus floribundus 

Lonchocarpus nitidus Spruce ex Benth. =Lonchocarpus floribundus 

Lonchocarpus paniculatus Ducke =Lonchocarpus hedyosmus 

Lonchocarpus pentaphyllus (Poir.) DC. =Lonchocarpus heptaphyllus 

Lonchocarpus pterocarpus DC. =Derris pterocarpus 

Lonchocarpus sanctae-marthae Pittier =Lonchocarpus fendleri 

Lonchocarpus scandens (Aubl.) Ducke =Derris pterocarpus


Lonchocarpus stenopteris Pittier =Lonchocarpus dipteroneurus 

Lonchocarpus stenurus Pittier =Lonchocarpus fendleri 

Lonchocarpus violaceus (Jacq.) DC. =Lonchocarpus punctatus 

Machaerium aculeatum Raddi [not in flora area] 

Machaerium bracteatum Benth. =Machaerium kegelii 

Machaerium caicarense Pittier =Machaerium dubium 

Machaerium compressicaule Ducke =Machaerium madeirense 

Machaerium compressicaule var. manoense Ducke =Machaerium madeirense 

Machaerium decorticans Ducke =Machaerium floribundum 

Machaerium eggersii Hoehne =Machaerium robiniifolium 

Machaerium ferrugineum (Willd.) Pers. =Machaerium quinatum 

Machaerium ferrugineum ß parviflorum Benth. =Machaerium quinatum var. parviflorum 

Machaerium frondosum (Mart. ex Benth.) Ducke =Machaerium leiophyllum var. latifolium 

Machaerium latifolium var. manoense (Ducke) Hoehne =Machaerium madeirense 

Machaerium longistipitatum Pittier =Machaerium floribundum 

Machaerium nervosum Vogel =Machaerium quinatum var. parviflorum 

Machaerium oblongifolium Vogel [not in flora area] 

Machaerium pachyphyllum Pittier =Machaerium kegelii 

Machaerium pseudacutifolium Pittier =Machaerium acutifolium var. pseudacutifolium 

Machaerium rectipes Pittier =Machaerium affine 

Machaerium rosescens Standl. =Machaerium floribundum 

Machaerium schomburgkii Benth. =Paramachaerium schomburgkii 

Machaerium sieberi Benth. =Machaerium robiniifolium 

Machaerium steinbachianum, Hoehne =Machaerium kegelii 

Machaerium surinamense Miq. =Machaerium leiophyllum 

Machaerium venezuelense (Pittier) Hoehne =Machaerium floribundum 

Machaerium woodworthii Standl. =Machaerium floribundum 

Meibomia axillaris (Sw.) Kuntze =Desmodium axillare 

Meibomia axillaris var. acutifolia Kunze =Desmodium axillare var. acutifolium 

Meibomia campyloclada (Hemsl.) Kuntze =Desmodium campyloclados 

Micropteryx poeppigiana Walp. =Erythrina poeppigiana 

Monopteryx jahnii Pittier =Fissicalyx fendleri 

Moutouchi suberosa Aubl. =Pterocarpus officinalis 

Mucuna altissima (Jacq.) DC. =Mucuna urens 

Mucuna hassjoo (Piper & Tracy) Mansf. =Mucuna pruriens var. utilis 

Mucuna utilis Wall. ex Wight =Mucuna pruriens var. utilis 

Muellera moniliformis L. f. =Muellera frutescens 

Neurocarpum cajanifolium C. Presl =Clitoria laurifolia 

Neurocarpum guianense (Aubl.) Desv. =Clitoria guianensis 

Neurocarpum javitense auct. non Kunth =Clitoria sagotii 

Neurocarpum javitense Kunth =Clitoria javitensis 

Neurocarpum simplicifolium Kunth =Clitoria simplicifolia 

Nicolsonia orinocensis (DC.) Schindl. =Desmodium orinocense 

Nissolia acuminata (Kunth) DC. =Machaerium acuminatum 

Nissolia ferruginea Willd. =Machaerium quinatum 

Nissolia leiophylla DC. =Machaerium leiophyllum 

Nissolia polyphylla Poir. =Machaerium polyphyllum 

Nissolia quinata Aubl. =Machaerium quinatum 

Nissolia robiniifolia DC. =Machaerium robiniifolium 

Ormosia chlorocalyx Ducke =Ormosia macrocalyx 

Ormosia coccinea var. subsimplex (Spruce ex Benth.) Rudd =Ormosia subsimplex 

Ormosia costulata var. trifoliata (Huber) Amshoff =Ormosia costulata 

Ormosia crassicarpa Pierce ex Pittier =Ormosia paraensis 

Ormosia cuneata Ducke =Ormosia coarctata 

Ormosia euneura Harms =Ormosia amazonica 

Ormosia fastigiata Tul. [not in flora area] 

Ormosia heterophylla Pires =Ormosia paraensis 

Ormosia micrantha Ducke =Ormosia discolor 

Ormosia microsperma Pittier =Ormosia steyermarkii 

Ormosia nobilis var. bolivarensis Rudd =Ormosia bolivarensis 

Ormosia pacimonensis Spruce ex Benth. =Vatairea guianensis 

Ormosia toledana Standl. =Ormosia macrocalyx 

Ormosia trifoliata Huber =Ormosia costulata 

Ormosiopsis flava Ducke =Ormosia flava

362 


Phaseolus adenanthus G. Mey. =Vigna adenantha 

Phaseolus appendiculatus Benth. =Vigna candida 

Phaseolus campestris Mart. ex Benth. =Macroptilium longipedunculatum 

Phaseolus candidus Vell. =Vigna candida 

Phaseolus diversifolius Pittier =Vigna lasiocarpa 

Phaseolus gracilis Poepp. ex Benth. =Macroptilium gracile 

Phaseolus juruanus Harms =Vigna juruana 

Phaseolus lasiocarpus Mart. ex Benth. =Vigna lasiocarpa 

Phaseolus lathyroides L. =Macroptilium lathyroides 

Phaseolus linearis Kunth =Vigna linearis 

Phaseolus longifolius Benth. =Vigna longifolia 

Phaseolus longipedunculatus Mart. ex Benth. =Macroptilium longipedunculatum 

Phaseolus monophyllus Benth. =Macroptilium monophyllum 

Phaseolus peduncularis Kunth =Vigna peduncularis 

Phaseolus pilosus Kunth =Vigna lasiocarpa 

Phaseolus schottii Benth. =Vigna longifolia 

Phaseolus surinamensis Miq. =Vigna adenantha 

Phaseolus trichocarpus C. Wright =Vigna trichocarpa 

Phaseolus truxillensis Kunth =Vigna adenantha 

Phaseolus vexillatus L. =Vigna vexillata 

Phellocarpus amazonum Mart. ex Benth. =Pterocarpus amazonum 

Piscidia guaricensis (Pittier) Pittier =Piscidia carthaginensis 

Platymiscium duckei Huber =Platymiscium trinitatis 

Platymiscium duckei var. nigrum (Ducke) Ducke =Platymiscium trinitatis 

Platymiscium nigrum Ducke =Platymiscium trinitatis 

Platymiscium polystachyum Benth. =Platymiscium pinnatum 

Platymiscium polystachyum var. fendleri Benth. =Platymiscium pinnatum 

Platypodium maxonianum Pittier =Platypodium elegans subsp. maxonianum 

Platysema triquetra Hoffmanns. ex Benth. =Centrosema triquetrum 

Possira arborescens Aubl. =Swartzia arborescens 

Possira triphylla Sw. =Swartzia arborescens 

Pterocarpus amazonicus Huber =Pterocarpus santalinoides 

Pterocarpus draco L. =Pterocarpus officinalis 

Pterocarpus esculentus Schumach. & Thonn. =Pterocarpus santalinoides 

Pterocarpus frutescens Vell. =Dalbergia frutescens 

Pterocarpus grandis R.S. Cowan =Pterocarpus santalinoides 

Pterocarpus lunatus L. f. =Machaerium lunatum 

Pterocarpus magnicarpa Schery =Pterocarpus rohrii 

Pterocarpus magnicarpus Schery =Pterocarpus rohrii 

Pterocarpus podocarpus S.F. Blake =Pterocarpus acapulcensis 

Pterocarpus rohrii var. rubiginosus Schery =Pterocarpus rohrii 

Pterocarpus rufescens Benth. =Pterocarpus rohrii 

Pterocarpus rupestris Pittier =Pterocarpus rohrii 

Pterocarpus suberosa (Aubl.) Pers. =Pterocarpus officinalis 

Pterocarpus ulei Harms =Pterocarpus amazonum 

Pterocarpus vernalis Pittier =Pterocarpus acapulcensis 

Rhadinocarpus brasiliensis Vogel =Chaetocalyx brasiliensis 

Rhynchosia caliensis Harms =Rhynchosia schomburgkii 

Rhynchosia crinita (Kunth) DC. =Eriosema crinitum 

Rhynchosia kuntzei Harms ex Kuntze =Rhynchosia reticulata 

Rhynchosia melanosticta Griseb. =Rhynchosia edulis 

Rhynchosia phaseoloides var. erecta Micheli =Rhynchosia melanocarpa 

Rhynchosia reticulata var. kuntzei (Harms ex Kuntze) Grear =Rhynchosia reticulata 

Rhynchosia rufa (Kunth) DC. =Eriosema rufum 

Rhynchosia simplicifolia (Kunth) DC. =Eriosema simplicifolium 

Rhynchosia violacea (Aubl.) DC. =Eriosema violaceum 

Rittera dodecandra Vahl =Swartzia arborescens 

Rittera pinnata Vahl =Swartzia pinnata 

Robinia coccinea Aubl. =Ormosia coccinea 

Robinia ferruginea Kunth =Coursetia ferruginea 

Robinia nicou Aubl. =Lonchocarpus utilis 

Robinia panacoco Aubl. =Swartzia panacoco 

Robinia sepium Jacq. =Gliricidia sepium 

Robinia sericea Poir. =Lonchocarpus sericeus


Robinia tomentosa Willd. =Swartzia panacoco 

Robinia violacea Jacq. [nom. dubium] =Lonchocarpus punctatus 

Sesbania aegyptica Poir. =Sesbania sesban 

Sphinctolobium glaucescens Miq. =Muellera frutescens 

Spirotropis candollei Tul. =Spirotropis longifolia 

Stenolobium brachycarpum (Desv.) Benth. =Calopogonium mucunoides 

Stenolobium coeruleum Benth. =Calopogonium caeruleum 

Stenolobium velutinum Benth. =Calopogonium velutinum 

Stizolobium altissimum (Jacq.) Pers. =Mucuna urens 

Stizolobium aterrimun Piper & Tracy =Mucuna pruriens var. utilis 

Stizolobium hassjo Piper & Tracy =Mucuna pruriens var. utilis 

Stizolobium pruriens (L.) Medik. =Mucuna pruriens 

Stylosanthes gracilis Kunth =Stylosanthes guianensis var. gracilis 

Swartzia acuminata Willd. ex Vogel =Swartzia polyphylla 

Swartzia acuminata var. puberula (Taub.) Glaz. =Swartzia polyphylla 

Swartzia acuminata var. tridynamia Huber =Swartzia polyphylla 

Swartzia alata Willd. =Swartzia guianensis 

Swartzia alterna Benth. (Leguminosae-Caesalpinioideae) =Bocoa alterna 

Swartzia apetala Raddi [not in flora area] 

Swartzia apetala var. acuminata Amshoff =Swartzia amshoffiana 

Swartzia bifida Steud. =Swartzia arborescens 

Swartzia brachystachya DC. =Candolleodendron brachystachyum 

Swartzia corrugata Benth. [not in flora area] 

Swartzia dicarpa Moric. ex Meisn. =Swartzia dipetala 

Swartzia discolor Poepp. =Swartzia leptopetala 

Swartzia dodecandra (Vahl) Willd. =Swartzia arborescens 

Swartzia dolichopoda R.S. Cowan =Swartzia vaupesiana 

Swartzia fugax Spruce ex Benth. =Swartzia leptopetala 

Swartzia grandifolia var. leiogyne Sandwith =Swartzia leiogyne 

Swartzia longifolia DC. =Spirotropis longifolia 

Swartzia melanoxylon Ducke =Swartzia leptopetala 

Swartzia microstylis Benth. =Swartzia dipetala 

Swartzia minutiflora Kleinhoonte (Leguminosae-Caesalpinioideae) =Bocoa prouacensis 

Swartzia opacifolia J.F. Macbr. =Swartzia polyphylla 

Swartzia parviflora DC. =Swartzia arborescens 

Swartzia platygyne Ducke =Swartzia polyphylla 

Swartzia polyanthera Steud. =Swartzia panacoco var. polyanthera 

Swartzia polycarpa Ducke =Swartzia laxiflora 

Swartzia prouacensis (Aubl.) Amshoff (Leguminosae-Caesalpinioideae) =Bocoa prouacensis 

Swartzia rariflora Hoehne =Swartzia arborescens 

Swartzia rotundata R.S. Cowan =Swartzia leptopetala 

Swartzia schomburgkii var. guayanensis R.S. Cowan =Swartzia schomburgkii 

Swartzia similis Benoist =Swartzia panacoco 

Swartzia simplex (Sw.) Spreng. [not in flora area] 

Swartzia tepuiensis Schery =Swartzia panacoco var. tepuiensis 

Swartzia tomentosa (Willd.) DC. =Swartzia panacoco 

Swartzia tomentosa var. altsoni Sandwith =Swartzia panacoco var. altsonii 

Swartzia tomentosa var. cardonae R.S. Cowan =Swartzia panacoco var. cardonae 

Swartzia tomentosa var. polyanthera (Steud.) Sandwith =Swartzia panacoco var. polyanthera 

Swartzia tomentosa var. sagotii Sandwith =Swartzia panacoco var. sagotii 

Swartzia tomentosa var. sandwithiana R.S. Cowan =Swartzia panacoco var. sandwithiana 

Swartzia triphyllata Willd. =Swartzia arborescens 

Swartzia urubuensis Ducke =Swartzia polyphylla 

Swartzia vaupesiana var. glauca R.S. Cowan =Swartzia vaupesiana 

Swartzia viridiflora Ducke (Leguminosae-Caesalpinioideae) =Bocoa viridiflora 

Sweetia nitens (Vogel) Benth. =Acosmium nitens 

Sweetia praeclara Sandwith =Acosmium praeclarum 

Taralea casiquiarensis Pittier =Dipteryx magnifica 

Taralea rigida Schery =Taralea cordata var. rigida 

Taralea steyermarkii Schery =Taralea reticulata 

Tephrosia brevipes Benth. =Tephrosia sessiliflora 

Tephrosia cathartica (Sessé & Moc.) Urb. =Tephrosia senna 

Tephrosia cinerea var. littoralis (Jacq.) Benth. =Tephrosia cinerea 

Tephrosia littoralis (Jacq.) Pers. =Tephrosia cinerea

364 


Tephrosia toxicaria (Sw.) Pers. =Tephrosia sinapou 

Ternatea arborescens (W.T. Aiton) Kuntze =Clitoria arborescens 

Ternatea javitensis (Kunth) Kuntze =Clitoria javitensis 

Ternatea simplicifolia (Kunth) Kuntze =Clitoria simplicifolia 

Toluifera balsamum L. =Myroxylon balsamum 

Touantea latifolia (Benth.) Taub. =Swartzia latifolia 

Touantea sericea (Vogel) Kuntze =Swartzia sericea 

Tounatea acuminata (Willd. ex Vogel) Taub. =Swartzia polyphylla 

Tounatea acuminata var. puberula Taub. =Swartzia polyphylla 

Tounatea aptera (DC.) Taub. =Swartzia aptera 

Tounatea arborescens (Aubl.) Britton =Swartzia arborescens 

Tounatea benthamiana (Miq.) Taub. =Swartzia benthamiana 

Tounatea dodecandra (Vahl) Taub. =Swartzia arborescens 

Tounatea guianensis Aubl. =Swartzia guianensis 

Tounatea hostmanni (Benth.) Taub. =Swartzia hostmannii 

Tounatea laxiflora (Bong. ex Benth.) Taub. =Swartzia laxiflora 

Tounatea laxiflora (Bong. ex Benth.) Kuntze =Swartzia laxiflora 

Tounatea schomburgkii (Benth.) Taub. =Swartzia schomburgkii 

Trifolium guianense Aubl. =Stylosanthes guianensis 

Vatairea surinamensis Kleinhoonte =Vatairea guianensis 

Vicia littoralis Jacq. =Tephrosia cinerea 

Vigna repens (L.) Kuntze =Vigna luteola 

Zollernia grandiflora Schery =Zollernia paraënsis 

Zollernia kanukuensis R.S. Cowan =Zollernia paraënsis 

Zollernia ulei Harms =Zollernia paraënsis 

Zornia marajoara Huber [nomen nudum] =Zornia guanipensis 

Zornia pubescens Kunth =Zornia latifolia var. latifolia 

Zornia surinamensis Miq. =Zornia latifolia var. latifolia 

Zornia tenuifolia var. latifolia Benth. =Zornia guanipensis 

REFERENCES 

Amshoff, G.J.H. 1939. Papilionaceae. In A. Pulle, Flora of Suriname 2(2): 1-257. 

Aymard C., G. & N. Cuello A. 1991. Dos nuevas especies Neotropicales del genero Canavalia (Papilionoideae - Phaseoleae - 

Diocleinae). Biollania 8: 87-92. 

Aymard C., G.A., Nidia L. Cuello A., P.E. Berry, V.E. Rudd, R.S. Cowan, P.R. Fantz, R.H. Maxwell, C.H. Stirton, H.-H. 

Poppendieck, H. Cavalcante de Lima, R.H. Fortunato, Basil Stergios, N. Xena de Enrich, D.A. Neill, R.T. Pennington, & C. 

Gil. 1999. Fabaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 5: 231- 

433. 

Barneby, R.C. & S.V. Heald. 2002. Fabaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 298-319. 

Burkart, A. 1971. El género Galactia (Legum.-Phaseoleae) en Sudamérica. Darwiniana 16: 663-796. 

Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. Mem. 


Ducke, A. 1942. Lonchocarpus, subgenus Phacelanthus Pittier, in Brazilian Amazonia. Trop. Woods 69: 2-7. 

Dwyer, J.D. 1980. Leguminosae subfamily Papilionoideae (conclusion). In R.E. Woodson, Jr., R.W. Schery, & collaborators, 

Flora of Panama. Ann. Missouri Bot. Gard. 67: 523-818. 

Görts-van Rijn, A.R.A., K.U. Kramer, & J.C. Lindeman. 1976. Papilionaceae (additions and corrections). In J. Lanjouw & A.L. 

Stoffers, Flora of Suriname 2(2): 556-610. 

Grear, J.W. 1971. A revision of the American species of Eriosema. Mem. New York Bot. Gard. 20(3): 1-98. 

Grear, J.W. 1978. A revision of the New World species of Rhynchosia. Mem. New York Bot. Gard. 31(1): 1-168. 

Krukoff, B.A. 1969. Supplementary notes on the American species of Erythrina. III. Phytologia 19: 113-175. 

Lima, H.C. de. 1982. Revisao taxonômica do género Vatairea Aublet (Leguminosae - Faboideae). Arquiv. Jard. Bot. Rio de 

Janeiro 26: 173-213. 

Lima, H.C. de. 1985. Diplotropis Bentham. Estudo dos Táxons infragenéricos. Acta Amaz. 15: 61-75. 

Maxwell, R.H. 1970. The genus Cymbosema (Leguminosae): notes and distribution. Ann. Missouri Bot. Gard. 57(2): 252-257. 

Mohlenbrock, R.H. 1958. A revision of the genus Stylosanthes. Ann. Missouri Bot. Gard. 44: 299-355. 

Mohlenbrock, R.H. 1961. A monograph of the leguminous genus Zornia. Webbia 16(1): 1-141. 

Mohlenbrock, R.H. 1963. Further considerations in Stylosanthes. Rhodora 65: 245-258. 


Ramírez, N. 1995. Revisión taxonómica del género Alexa Moq. (Fabaceae, Sophoreae). Ann. Missouri Bot. Gard. 82(4): 549- 

569. 

Rojo, J.P. 1972. Pterocarpus. Revised for the world. Phanerog. Monogr. 5: 1-119. 

Rudd, V.E. 1955. The American species of Aeschynomene. Contr. U.S. Nat. Herb. 32: 1-172.


Rudd, V.E. 1955. A revision of the genus Chaetocalyx. Contr. U.S. Nat. Herb. 32: 247-277. 

Rudd, V.E. 1963. The genus Dussia (Leguminosae). Contr. U.S. Nat. Herb. 32: 247-277. 

Rudd, V.E. 1965. The American species of Ormosia. Contr. U.S. Nat. Herb. 32: 279-384. 

Rudd, V.E. 1981. Two new species of Paramachaerium and a brief resumé of the genus. Brittonia 33(3): 435-440. 

Rudd, V.E. 1987. Studies in Machaerium (Leguminosae) VI. Phytologia 62: 282-302. 

Rudd, V.E. 1989. Studies in Machaerium VII. Section II, Lineata. Part I. Species with wingless fruit. Phytologia 64: 1-12. 

Sauer, J. 1964. Revision of Canavalia. Brittonia 16: 106-181. 

Schultze-Kraft, R.J. & L. Coradin. 1990. Biogeography of Centrosema. In R. Schultze-Kraft & R.J. Clements (ed.), Centrosema: 

Biology, Agronomy, and Utilization. 

Verdcourt, B. 1970. Studies in the Leguminosae - Papilionoideae for the ‘Flora of Tropical East Africa’: IV. Kew Bull. 24: 507- 

569. 

Williams, R.J. & R.R. Clements. 1990. Taxonomy of Centrosema. In R. Schultze-Kraft & R.J. Clements (ed.), Centrosema: 

Biology, Agronomy, and Utilization. 

Yakovlev, G.P. 1969. A review of Sweetia and Acosmium. Notes Royal Bot. Gard. Edinb. 29: 347-355. 

LEGUMINOSAE-MIMOSOIDEAE 

R.C. Barneby, P.E. Berry, D. Brunner, L. Cárdenas, E. Forero, J.W. Grimes, H.C.F. Hopkins, 

E.M. de Lamare Occhioni, G. De Martino, O. Poncy 

Abarema acreana (J.F. Macbr.) L. Rico 

Abarema adenophora (Ducke) Barneby & J.W. Grimes 

Abarema barbouriana (Standl.) Barneby & J.W. Grimes 

Abarema barbouriana var. arenaria (Ducke) Barneby & J.W. Grimes 

Abarema barbouriana var. barbouriana 

Abarema commutata Barneby & J.W. Grimes 

Abarema curvicarpa (H.S. Irwin) Barneby & J.W. Grimes 

Abarema curvicarpa var. curvicarpa 

Abarema ferruginea (Benth.) Pittier 

Abarema floribunda (Spruce ex Benth.) Barneby & J.W. Grimes 

Abarema gallorum Barneby & J.W. Grimes 

Abarema jupunba (Willd.) Britton & Killip 

Abarema jupunba var. jupunba 

Abarema jupunba var. trapezifolia (Vahl) Barneby & J.W. Grimes 

Abarema laeta (Benth.) Barneby & J.W. Grimes 

Abarema leucophylla (Spruce ex Benth.) Barneby & J.W. Grimes 

Abarema leucophylla var. leucophylla 

Abarema levelii (R.S. Cowan) Barneby & J.W. Grimes 

Abarema longipedunculata (H.S. Irwin) Barneby & J.W. Grimes [expected in 

GU] 

Abarema mataybifolia (Sandwith) Barneby & J.W. Grimes 

Abarema microcalyx (Spruce ex Benth.) Barneby & J.W. Grimes 

Abarema microcalyx var. microcalyx 

Abarema villifera (Ducke) Barneby & J.W. Grimes 

Acacia alemquerensis Huber [expected in the Guianas] 

Acacia articulata Ducke 

Acacia farnesiana (L.) Willd. 

Acacia farnesiana var. farnesiana 

Acacia floribunda (Spruce ex Benth.) Barneby & J.W. Grimes 

Acacia glomerosa Benth. 

Acacia macracantha Humb. & Bonpl. ex Willd. [introduced] 

Acacia podadenia (Britton & Killip) L. Cardenas 

Acacia polyphylla DC. 

Acacia tamarindifolia (L.) Willd. 

Acacia tenuifolia (L.) Willd. 

Acacia tenuifolia var. producta J.W. Grimes 

Acacia tenuifolia var. tenuifolia 

Adenanthera pavonina L. [cultivated, naturalized] 

Albizia barinensis L. Cardenas 

Albizia glabripetala (H.S. Irwin) G.P. Lewis & P.E. Owen 

Albizia lebbeck (L.) Benth. [cultivated, naturalized] 

Albizia niopoides (Spruce ex Benth.) Burkart 

Albizia subdimidiata (Splitg.) Barneby & J.W. Grimes 

Albizia subdimidiata var. minor Barneby & J.W. Grimes 

Albizia subdimidiata var. subdimidiata 

AM BO 

AM 


AM BO GU 

GU FG 

GU SU 

GU FG 

GU FG 

BO GU? 

AM GU? 

FG 

AM GU SU FG 

AM FG 

GU SU FG 

AM SU FG 

AM 

AM 

AM 

BO 

GU SU FG 

AM 

AM 

AM BO 

BO 

SU 

BO SU 

BO 

BO 

GU 

GU 

GU 

SU FG 

BO 

BO 

BO 

GU 

GU 

FG 

BO GU SU FG 

SU FG 

GU SU 

SU 

FG 

BO GU 

AM BO GU 

GU 

GU 

SU FG 

BO DA GU 

GU 

SU FG 

GU SU

366 


Anadenanthera peregrina (L.) Speg. [naturalized?] 

Anadenanthera peregrina var. peregrina 

Balizia pedicellaris (DC.) Barneby & J.W. Grimes 

Calliandra calothyrsus Meisn. [cultivated, perhaps escaping] 

Calliandra coriacea (Humb. & Bonpl. ex Willd.) Benth. [expected in FG] 

Calliandra cruegeri Griseb. 

Calliandra glomerulata H. Karst. 

Calliandra glomerulata var. glomerulata 

Calliandra guildingii Benth. 

Calliandra hymeniodes (Rich.) Benth. 

Calliandra laxa (Willd.) Benth. 

Calliandra laxa var. laxa 

Calliandra laxa var. stipulacea (Benth.) Barneby 

Calliandra laxa var. urimana Barneby 

Calliandra pakaraimensis R.S. Cowan 

Calliandra rigida Benth. 

Calliandra riparia Pittier 

Calliandra surinamensis Benth. 

Calliandra tergemina (L.) Benth. 

Calliandra trinervia Benth. 

Calliandra trinervia var. pilosifolia (R.S. Cowan) Barneby 

Calliandra trinervia var. trinervia 

Calliandra tsugoides R.S. Cowan 

Calliandra vaupesiana R.S. Cowan 

Calliandra vaupesiana var. oligandra Barneby 

Cedrelinga cateniformis (Ducke) Ducke 

Chloroleucon acacioides (Ducke) Barneby & J.W. Grimes 

Chloroleucon eurycyclum Barneby & J.W. Grimes 

Chloroleucon mangense (Jacq.) Britton & Rose 

Chloroleucon mangense var. tetrazyx Barneby & J.W. Grimes 

Desmanthus pernambucanus (L.) Thell. 

Desmanthus virgatus (L.) Willd. [expected in FG] 

Dichrostachys glomerata (Forssk.) Chiov. [naturalized] 

Entada polyphylla Benth. 

Entada polystachya (L.) DC. 

Enterolobium barinense L. Cardenas & H. Rodr. 

Enterolobium barnebianum Mesquita & M.F. Silva [expected in Venezuelan 

Guayana] 

Enterolobium cyclocarpum (Jacq.) Griseb. [expected in Venezuelan Guayana] 

Enterolobium oldemanii Barneby & J.W. Grimes 

Enterolobium schomburgkii (Benth.) Benth. 

Hydrochorea corymbosa (Rich.) Barneby & J.W. Grimes 

Hydrochorea gonggrijpii (Kleinhoonte) Barneby & J.W. Grimes 

Hydrochorea marginata (Spruce ex Benth.) Barneby & J.W. Grimes 

Hydrochorea marginata var. panurensis (Spruce ex Benth.) Barneby & J.W. 

Grimes 

Hydrochorea marginata var. scheryi Barneby & J.W. Grimes 

Inga acreana Harms 

Inga acrocephala Steud. 

Inga alata Benoist 

Inga alba (Sw.) Willd. 

Inga albicoria Poncy 

Inga auristellae Harms 

Inga bijuga Schery 

Inga bourgonii (Aubl.) DC. 

Inga brachystachys Ducke 

Inga brevipes Benth. 

Inga calanthoides Amshoff 

Inga capitata Desv. 

Inga cayennensis Sagot ex Benth. 

Inga cinnamomea Spruce ex Benth. 

Inga crassiflora Ducke 

Inga cylindrica Mart. 

Inga densiflora Benth. 




AM GU SU 

AM BO GU 

AM BO GU 

AM BO GU 

AM 

GU SU FG 

AM BO GU 

BO 

AM BO GU 

BO 

BO GU 

BO GU 

BO GU 


BO GU SU FG 

AM GU FG 

AM 

AM GU FG 

AM BO 

AM 

AM 


SU FG 

BO 

BO 

BO 

GU SU 

GU SU 

FG 

AM GU SU FG 

BO GU SU FG 

AM BO DA 

GU? 

GU SU? 

FG 




AM 

AM 

AM 

SU FG 

GU SU FG 

GU FG 


GU FG 

AM GU SU FG 

BO 


SU FG 

GU SU 

SU 



SU 

FG 

FG 

SU

Inga disticha Benth. 

Inga edulis Mart. 

Inga fanchoniana Poncy 

Inga fastuosa (Jacq.) Willd. 

Inga flagelliformis Mart. 

Inga graciliflora Benth. 

Inga gracilifolia Ducke 

Inga heterophylla Willd. 

Inga huberi Ducke 

Inga inflata Ducke 

Inga ingoides (Rich.) Willd. 

Inga java Pittier 

Inga jenmanii Sandwith 

Inga lateriflora Miq. 

Inga laurina (Sw.) Willd. 

Inga leiocalycina Benth. 

Inga leptingoides Amshoff 

Inga lomatophylla (Benth.) Pittier 

Inga longipedunculata Ducke 

Inga macrophylla Humb. & Bonpl. ex Willd. 

Inga marginata Willd. 

Inga melinonis Sagot 

Inga micradenia Spruce ex Benth. 

Inga multijuga Benth. 

Inga multijuga subsp. multijuga 

Inga neblinensis L. Cardenas & De Martino 

Inga nobilis Willd. 

Inga nobilis subsp. nobilis 

Inga nouragensis Poncy 

Inga nubium Poncy 

Inga nuda Salzm. ex Benth. 

Inga paraensis Ducke 

Inga pedunculata Ducke 

Inga pezizifera Benth. 

Inga pilosula (Rich.) J.F. Macbr. 

Inga poeppigiana Benth. 

Inga retinocarpa Poncy 

Inga rhynchocalyx Sandwith 

Inga rubiginosa (Rich.) DC. 

Inga salicifolia T.D. Penn. [expected in AM, Serra da Neblina] 

Inga sapindoides Willd. 

Inga sarmentosa Glaz. ex Harms 

Inga sertulifera DC. 

Inga sertulifera subsp. leptopus (Benth.) T.D. Penn. 

Inga sertulifera subsp. sertulifera 

Inga splendens Willd. 

Inga stenocalyx Spruce ex Benth. 

Inga stenoptera Benth. 

Inga stipularis DC. 

Inga striata Benth. [=I. nuda?] 

Inga suaveolens Ducke 

Inga thibaudiana DC. 

Inga thibaudiana subsp. thibaudiana 

Inga tubiformis Benoist 

Inga ulei Harms 

Inga umbellifera (Vahl) Steud. ex DC. 

Inga umbratica Poepp. 

Inga venusta Standl. 

Inga vera Willd. 

Inga vera subsp. affinis (DC.) T.D. Penn. 

Inga vera subsp. vera 

Inga virgultosa (Vahl) Desv. 

Leucaena leucocephala (Lam.) de Wit [cultivated, naturalized] 

Leucaena leucocephala subsp. glabrata (Rose) S. Zárate 


GU SU FG 


GU? FG 


FG 

GU SU FG 



DA GU SU FG 

AM 


GU FG 

GU FG 




SU 

AM GU FG 

FG 

AM GU SU FG 

AM GU SU FG 

DA GU FG 

AM BO 

BO 

BO 

AM 



FG 

FG 

GU SU FG 

BO GU FG 

FG 



FG 

FG 

GU FG 


BO DA FG 

GU SU FG 


GU 



GU FG 

AM BO DA GU 


FG 

FG 



FG 

AM 


BO FG 

SU 



BO DA 

SU FG 

BO GU SU FG 

BO

368 


Lysiloma bahamensis Benth. [introduced, dubiously persisting] 

Macrosamanea amplissima (Ducke) Barneby & J.W. Grimes 

Macrosamanea consanguinea (R.S. Cowan) Barneby & J.W. Grimes 

Macrosamanea discolor (Humb. & Bonpl. ex Willd.) Britton & Rose ex 

Britton & Killip 

Macrosamanea kegelii (Meisn.) Kleinhoonte 

Macrosamanea pubiramea (Steud.) Barneby & J.W. Grimes 

Macrosamanea pubiramea var. lindsaeifolia (Spruce ex Benth.) Barneby & 

J.W. Grimes 

Macrosamanea simabifolia (Spruce ex Benth.) Pittier 

Macrosamanea spruceana (Benth.) Killip 

Mimosa annularis Benth. 

Mimosa annularis var. odora Barneby 

Mimosa annularis var. xinguensis (Ducke) Barneby 

Mimosa arenosa (Willd.) Poir. 

Mimosa arenosa var. arenosa 

Mimosa bimucronata (DC.) Kuntze [cultivated or escaped from cultivation] 

Mimosa brachycarpoides Barneby 

Mimosa camporum Benth. 

Mimosa casta L. [possibly introduced in FG] 

Mimosa colombiana Britton & Killip 

Mimosa corynadenia Britton & Rose [expected in SW Venezuelan Guayana] 

Mimosa debilis Humb. & Bonpl. ex Willd. 

Mimosa debilis var. debilis 

Mimosa diplotricha C. Wright ex Sauv. 

Mimosa dormiens Humb. & Bonpl. ex Willd. 

Mimosa guanchezii Barneby 

Mimosa guilandinae (DC.) Barneby 

Mimosa guilandinae var. duckei (Huber) Barneby 

Mimosa guilandinae var. guilandinae 

Mimosa hirsutissima Mart. 

Mimosa huberi Barneby 

Mimosa invisa Mart. ex Colla [expected in BO] 

Mimosa invisa var. spiciflora (H. Karst.) Barneby [expected in BO] 

Mimosa microcephala Humb. & Bonpl. ex Willd. 

Mimosa microcephala var. cataractae (Ducke) Barneby 

Mimosa microcephala var. communis Barneby 

Mimosa microcephala var. lumaria Barneby 

Mimosa microcephala var. microcephala 

Mimosa myriadenia (Benth.) Benth. 

Mimosa myriadenia var. dispersa Barneby 

Mimosa myriadenia var. myriadenia 

Mimosa orinocoënsis Barneby 

Mimosa orthocarpa Spruce ex Benth. 

Mimosa pigra L. 

Mimosa piscatorum Barneby 

Mimosa polydactyla Humb. & Bonpl. ex Willd. 

Mimosa pudica L. 

Mimosa pudica var. hispida Brenan 

Mimosa pudica var. pastoris Barneby 

Mimosa pudica var. tetrandra (Humb. & Bonpl. ex Willd.) DC. 

Mimosa pudica var. unijuga (Walp. & Duchass.) Griseb. 

Mimosa quadrivalvis L. 

Mimosa quadrivalvis var. leptocarpa (DC.) Barneby 

Mimosa rufescens Benth. [expected in SU, FG] 

Mimosa rufescens var. amnis-nigri Barneby 

Mimosa rufescens var. rufescens 

Mimosa scabrella Benth. [occasionally escapes cultivation] 

Mimosa schomburgkii Benth. 

Mimosa schrankioides Benth. 

Mimosa schrankioides var. sagotiana (Benth.) Barneby 

Mimosa schrankioides var. schrankioides 

Mimosa sensitiva L. 

Mimosa somnians Humb. & Bonpl. ex Willd. 

AM 

AM 

AM 

FG 

SU FG 


AM 

AM BO 

BO 

BO 

DA 

GU SU FG 

GU SU FG 

FG 

BO 


DA FG 

AM BO 

AM 

AM BO 

AM BO 

BO GU? SU FG 

BO 

GU SU FG 

BO DA GU? 

SU FG 

SU FG 

FG 

GU SU? 

GU SU? 


SU FG 

AM BO 

AM BO GU FG? 

AM 


SU FG 

GU SU FG 

AM DA 

AM BO 


BO 



GU SU 

GU 





AM BO DA GU 

AM 

AM BO DA 

BO 

BO GU FG 



BO GU 

AM BO 

AM BO GU FG

Mimosa somnians var. deminuta Barneby 

Mimosa somnians var. somnians 

Mimosa somnians var. viscida (Willd.) Barneby 

Mimosa surumuensis Harms 

Mimosa tarda Barneby 

Mimosa velloziana Mart. [weedy, of doubtful native range] 

Mimosa xanthocentra Mart. [expected in BO] 

Mimosa xanthocentra var. xanthocentra [expected in BO] 

Neptunia oleracea Lour. [expected in BO] 

Neptunia plena (L.) Benth. 

Parkia barnebyana H.C. Hopkins 

Parkia decussata Ducke 

Parkia discolor Spruce ex Benth. 

Parkia gigantocarpa Ducke 

Parkia igneiflora Ducke 

Parkia nitida Miq. 

Parkia panurensis Benth. ex H.C. Hopkins 

Parkia pendula (Willd.) Benth. ex Walp. 

Parkia reticulata Ducke 

Parkia truncata R.S. Cowan 

Parkia ulei (Harms) Kuhlm. 

Parkia ulei var. surinamensis Kleinhoonte 

Parkia velutina Benoist [expected in GU] 

Pentaclethra macroloba (Willd.) Kuntze 

Piptadenia floribunda Kleinhoonte 

Piptadenia imatacae Barneby [expected in GU] 

Piptadenia leucoxylon Barneby & J.W. Grimes 

Piptadenia moniliformis Benth. [expected in the Guianas] 

Piptadenia uaupensis Spruce ex Benth. 

Piptadenia viridiflora (Kunth) Benth. 

Pithecellobium consanguineum R.S. Cowan 

Pithecellobium dulce (Roxb.) Benth. [cultivated, naturalized] 

Pithecellobium roseum (Vahl) Barneby & J.W. Grimes 

Pithecellobium unguis-cati (L.) Benth. [cultivated, naturalized] 

Plathymenia reticulata Benth. 

Pseudopiptadenia psilostachya (DC.) G.P. Lewis & M.P. Lima 

Pseudopiptadenia suaveolens (Miq.) J.W. Grimes 

Pseudosamanea guachapele (Kunth) Harms 

Samanea inopinata (Harms) Barneby & J.W. Grimes 

Samanea saman (Jacq.) Merr. [cultivated, perhaps escaping] 

Stryphnodendron guianense (Aubl.) Benth. 

Stryphnodendron guianense subsp. guianense 

Stryphnodendron levelii R.S. Cowan 

Stryphnodendron microstachyum Poepp. 

Stryphnodendron moricolor Barneby & J.W. Grimes 

Stryphnodendron polystachyum (Miq.) Kleinhoonte 

Stryphnodendron pulcherrimum (Willd.) Hochr. 

Zapoteca formosa (Kunth) H.M. Hern. 

Zapoteca formosa subsp. formosa 

Zapoteca portoricensis (Jacq.) H.M. Hern. 

Zapoteca portoricensis subsp. flavida (Urb.) H.M. Hern. 

Zygia ampla (Benth.) Pittier 

Zygia basijuga (Ducke) Barneby & J.W. Grimes 

Zygia cataractae (Kunth) L. Rico 

Zygia claviflora (Spruce ex Benth.) Barneby & J.W. Grimes 

Zygia collina (Sandwith) Barneby & J.W. Grimes 

Zygia eperuetorum (Sandwith) Barneby & J.W. Grimes 

Zygia inaequalis (Humb. & Bonpl. ex Willd.) Pittier 

Zygia inundata (Ducke) H.C. Lima 

Zygia juruana (Harms) L. Rico 

Zygia latifolia (L.) Fawc. & Rendle 

Zygia latifolia var. communis Barneby & J.W. Grimes 

Zygia latifolia var. lasiopus (Benth.) Barneby & J.W. Grimes 

Zygia latifolia var. latifolia 


AM BO 

BO GU FG 

GU FG 

BO GU 

AM 

AM 

GU SU FG 


AM 

SU FG 

AM 

AM 

GU 

GU 

AM 

AM 


AM 

AM 

BO GU SU FG 

FG 

DA GU SU FG 

DA GU SU FG 

BO SU FG 


BO 

SU FG 

AM 

BO 

BO 

BO 

GU 

FG? 

GU SU? FG 

BO GU 

BO GU 

SU 

FG 

GU SU FG 

AM BO 

BO 

DA GU SU FG 

BO GU 

GU 

BO GU SU FG 

AM 

BO GU FG 


GU FG 


BO 

BO 

BO 

GU SU FG 


BO 

BO 

DA GU 

GU 


AM GU SU FG 

AM BO DA GU 

GU 

AM BO GU 

GU 

SU FG 

FG 




DA GU SU FG

370 

Zygia palustris Barneby & J.W. Grimes 

Zygia potaroensis Barneby & J.W. Grimes 

Zygia racemosa (Ducke) Barneby & J.W. Grimes 

Zygia ramiflora (Benth.) Barneby & J.W. Grimes 

Zygia sabatieri Barneby & J.W. Grimes 

Zygia tetragona Barneby & J.W. Grimes 

Zygia unifoliolata (Benth.) Pittier [expected in GU] 



Abarema claviflora (Spruce ex Benth.) Kleinhoonte =Zygia claviflora 

Abarema racemosa (Ducke) Kleinhoonte =Zygia racemosa 

Abarema trapezifolia (Vahl) Pittier =Abarema jupunba var. trapezifolia 

Acacia alvaroi Cardenas & De Martino =Acacia alemquerensis 

Acacia arenosa Willd. =Mimosa arenosa 

Acacia chrysantha DC. [name of uncertain application] 

Acacia flexuosa Humb. & Bonpl. ex Willd. =Acacia macracantha 

Acacia formosa Kunth =Zapoteca formosa 

Acacia glandulosa (Vahl) DC. [nomen nudum] 

Acacia guachapele Kunth =Pseudosamanea guachapele 

Acacia guianensis (Aubl.) Willd. =Stryphnodendron guianense 

Acacia jupunba Willd. =Abarema jupunba 

Acacia laxa Willd. =Calliandra laxa 

Acacia leucocephala (Lam.) Link =Leucaena leucocephala 

Acacia lutea (Mill.) Britton =Acacia macracantha 

Acacia macroloba Willd. =Pentaclethra macroloba 

Acacia ouyrarema DC. [nomen nudum] 

Acacia paniculata Willd. =Acacia tenuifolia 

Acacia paniculiflora Steud. =Mimosa myriadenia 

Acacia psilostachya DC. =Pseudopiptadenia psilostachya 

Acacia pulcherrima Willd. =Stryphnodendron pulcherrimum 

Acacia subdimidiata Splitg. =Albizia subdimidiata 

Acacia viridiflora Kunth =Piptadenia viridiflora 

Acuan virgatum (L.) W. Medicus =Desmanthus virgatus 

Albizia caribaea (Urb.) Britton & Rose =Albizia niopoides 

Albizia corymbosa (Rich.) G.P. Lewis & P.E. Owen =Hydrochorea corymbosa 

Albizia guachapele (Kunth) Dugand =Pseudosamanea guachapele 

Albizia inopinata (Harms) G.P. Lewis =Samanea inopinata 

Albizia longepedata (Pittier) Britton & Rose =Pseudosamanea guachapele 

Anneslia calothyrsa (Meisn.) Kleinhoonte =Calliandra calothyrsus 

Anneslia fasciculata (Willd.) Kleinhoonte =Calliandra surinamensis 

Anneslia tergemina (L.) Britton & Rose =Calliandra tergemina 

Arthrosamanea corymbosa (Rich.) Kleinhoonte =Hydrochorea corymbosa 

Arthrosamanea gonggrijpii (Kleinhoonte) Kleinhoonte =Hydrochorea gonggrijpii 

Arthrosamanea multiflora (Kunth) Kleinhoonte [sensu Kleinhoonte] =Albizia subdimidiata 

Arthrosamanea panurensis (Spruce ex Benth.) Pittier =Hydrochorea marginata var. panurensis 

Arthrosamanea pullei (Kleinhoonte) Kleinhoonte =Hydrochorea gonggrijpii 

Calliandra affinis Pittier =Calliandra cruegeri 

Calliandra anthoniae J.W. Grimes =Calliandra coriacea 

Calliandra caripensis Willd. =Calliandra tergemina 

Calliandra flavida Urb. =Zapoteca portoricensis subsp. flavida 

Calliandra formosa (Kunth) Benth. =Zapoteca formosa 

Calliandra latifolia (L.) Griseb. =Zygia latifolia 

Calliandra marginata Griseb. ex R.O. Williams =Zapoteca formosa subsp. formosa 

Calliandra patrisii Sagot ? =Calliandra hymeniodes 

Calliandra pilosifolia R.S. Cowan =Calliandra trinervia var. pilosifolia 

Calliandra portoricensis (Jacq.) Benth. =Zapoteca portoricensis 

Calliandra resupina R.S. Cowan =Calliandra pakaraimensis 

Calliandra stipulacea Benth. =Calliandra laxa var. stipulacea 

Calliandra tenuiflora Benth. =Calliandra surinamensis 

Calliandra trijugata Schery =Calliandra laxa var. laxa 

Cathormion corymbosum (Rich.) Burkart =Hydrochorea corymbosa 

Entada gigas Fawc. & Rendle [not in flora area] 

Entada myriadenia Benth. =Mimosa myriadenia 

Entadopsis polyphylla (Benth.) Britton =Entada polyphylla 

AM GU 

BO GU 


AM 

FG 

FG 

AM BO


Entadopsis polystachya (L.) Britton =Entada polystachya 

Enterolobium racemosum Ducke [apparently a nomen nudum] 

Feuillea sapindoides (Willd.) Kuntze =Inga sapindoides 

Feuilleea cayennensis (Sagot ex Benth.) Kuntze =Inga cayennensis 

Feuilleea edulis (Mart.) Kuntze =Inga edulis 

Feuilleea fagifolia (L.) Kuntze =Inga laurina 

Feuilleea heterophylla (Willd.) Kuntze =Inga heterophylla 

Feuilleea ingoides (Rich.) Kuntze =Inga ingoides 

Feuilleea laurina (Sw.) Kuntze =Inga laurina 

Feuilleea lindeniana (Benth.) Kuntze =Inga sapindoides 

Feuilleea marginata (Willd.) Kuntze =Inga marginata 

Feuilleea myriantha (Poepp.) Kuntze =Inga umbellifera 

Feuilleea pezizifera (Benth.) Kuntze =Inga pezizifera 

Feuilleea pilosula (Rich.) Kuntze =Inga pilosula 

Feuilleea sciadon (Steud.) Kuntze =Inga umbellifera 

Feuilleea setifera (DC.) Kuntze =Inga pilosula 

Feuilleea splendens (Willd.) Kuntze =Inga splendens 

Feuilleea umbellifera (Vahl) Kuntze =Inga umbellifera 

Feuilleea umbratica (Poepp.) Kuntze =Inga umbratica 

Folianthera guianensis (Aubl.) Raf. =Stryphnodendron guianense 

Hydrochorea acreana (J.F. Macbr.) Barneby & J.W. Grimes =Abarema acreana 

Inga acuminata Benth. [material misidentified ] = I. rhynchocalyx 

Inga adiantifolia Kunth =Macrosamanea discolor 

Inga affinis DC. =Inga vera subsp. affinis 

Inga affinis Steud. =Inga pilosula 

Inga aggregata G. Don =Inga alba 

Inga amazonica L. Cardenas =Inga lomatophylla 

Inga amazonica var. bracteifera (Ducke) L. Cardenas =Inga lomatophylla 

Inga amazonica var. lomatophylla (Benth.) L. Cardenas [nom. superfl.] =Inga lomatophylla 

Inga amazonica var. membranacea (Ducke) L. Cardenas =Inga lomatophylla 

Inga angustifolia Willd. [not in flora area] 

Inga antioquensis Britton & Killip =Inga sapindoides 

Inga brachyptera Benth. =Inga macrophylla 

Inga bracteosa Benth. =Inga macrophylla 

Inga calocephala Poepp. =Inga macrophylla 

Inga camuriensis Pittier =Inga sapindoides 

Inga capitata Miq. ex Benth. =Abarema laeta 

Inga caracasana Pittier =Inga sapindoides 

Inga carachensis Pittier =Inga alba 

Inga cataractae Kunth =Zygia cataractae 

Inga cauliflora Willd. =Zygia latifolia var. communis 

Inga commewijnensis Miq. =Inga nuda 

Inga conglomerata Benoist =Inga nobilis 

Inga coriacea Humb. & Bonpl. ex Willd. =Calliandra coriacea 

Inga coriacea (Pers.) Desv. =Inga sertulifera subsp. sertulifera 

Inga corymbifera Benth. =Inga nobilis 

Inga crevauxii Sagot =Inga disticha 

Inga cyclocarpa Ducke =Inga melinonis 

Inga discolor Humb. & Bonpl. ex Willd. =Macrosamanea discolor 

Inga disticha var. negrensis (Spruce ex Benth.) Ducke =Inga micradenia 

Inga dysantha Benth. =Inga cayennensis 

Inga fagifolia (L.) Willd. ex Benth. =Inga laurina 

Inga floribunda Benth. =Inga splendens 

Inga forfex Kunth =Pithecellobium roseum 

Inga gladiata Desv. =Inga thibaudiana subsp. thibaudiana 

Inga glomerata DC. =Zygia cataractae 

Inga grandifolia Pittier =Inga sapindoides 

Inga guaremalensis Pittier =Inga fastuosa 

Inga inaequalis Humb. & Bonpl. ex Willd. =Zygia inaequalis 

Inga inga (Vell.) J. Moore =Inga edulis 

Inga laeta (Benth.) Poepp. =Abarema laeta 

Inga latifolia (L.) Willd. =Zygia latifolia 

Inga lawrenceana Britton & Killip =Inga umbellifera 

Inga lindeniana Benth. =Inga sapindoides

372 


Inga longiflora Spruce ex Benth. [not in flora area] 

Inga macradenia Mart. ex Benth. [nomen nudum] =Inga thibaudiana subsp. thibaudiana 

Inga macrophylla Hook. =Inga pilosula 

Inga meissneriana Miq. =Inga vera subsp. affinis 

Inga microphylla Humb. & Bonpl. ex Willd. =Pithecellobium unguis-cati 

Inga multiflora Benth. =Inga leiocalycina 

Inga myriantha Poepp. =Inga umbellifera 

Inga negrensis Spruce ex Benth. =Inga micradenia 

Inga niopo Humb. & Bonpl. ex Willd. =Anadenanthera peregrina 

Inga nitida Willd. =Inga pilosula 

Inga panurensis Spruce ex Benth. [not in flora area] 

Inga pedicellaris DC. =Balizia pedicellaris 

Inga pendula Willd. =Parkia pendula 

Inga perrottetii Sagot =Inga nuda 

Inga pilosiuscula Desv. =Inga pilosula 

Inga prieurei Sagot =Inga nuda 

Inga protracta Steud. =Inga heterophylla 

Inga pterocarpa DC. (Leguminosae-Caesalpinioideae) =Peltophorum pterocarpum 

Inga pubescens DC. =Pithecellobium roseum 

Inga pubiramea Steud. =Macrosamanea pubiramea 

Inga racemiflora Ducke =Inga ulei 

Inga ramiflora Steud. =Zygia latifolia var. lasiopus 

Inga ricardorum Bernardi & Spichiger [Guianas material identified as Inga java] 

Inga riparia Pittier =Inga vera subsp. vera 

Inga rosea (Vahl) Steud. =Pithecellobium roseum 

Inga scabriuscula Benth. =Inga edulis 

Inga sciadion Steud. =Inga umbellifera 

Inga sciandion Steud. =Inga umbellifera 

Inga semialata (Vell.) Mart. =Inga marginata 

Inga setifera DC. =Inga pilosula 

Inga speciosa Spruce ex Benth. =Inga lomatophylla 

Inga speciosa var. bracteifera Ducke =Inga lomatophylla 

Inga speciosa var. lomatophylla Benth. =Inga lomatophylla 

Inga speciosa var. membranacea Ducke =Inga lomatophylla 

Inga spuria Humb. & Bonpl. ex Willd. =Inga vera subsp. vera 

Inga strigillosa Spruce ex Benth. =Inga leiocalycina 

Inga tenuiflora Salzm. ex Benth. =Inga thibaudiana subsp. thibaudiana 

Inga umbellata G. Don =Inga heterophylla 

Inga unguis-cati (L.) Willd. =Pithecellobium unguis-cati 

Inga unijuga Poepp. & Endl. (Leguminosae-Caesalpinioideae) =Macrolobium limbatum var. limbatum 

Inga urnifera Kleinhoonte =Inga pezizifera 

Inga velutina (Poir.) Willd. [not in flora area] 

Inga venosa Griseb. ex Benth. =Inga fastuosa 

Inga vera Kunth =Inga edulis 

Jupunba trapezifolia (Vahl) Moldenke =Abarema jupunba var. trapezifolia 

Klugiodendron laetum (Benth.) Britton & Killip =Abarema laeta 

Leptoglottis leptocarpa (DC.) Standl. =Mimosa quadrivalvis var. leptocarpa 

Leucaena glabrata Rose =Leucaena leucocephala subsp. glabrata 

Leucaena glauca (L.) Benth. [not in flora area] 

Leucaena ulei Harms =Parkia ulei 

Lysiloma guachapele (Kunth) Benth. =Pseudosamanea guachapele 

Macrosamanea aquatica Pittier =Macrosamanea discolor 

Macrosamanea basijuga (Ducke) Dugand =Zygia basijuga 

Macrosamanea pedicellaris (DC.) Kleinhoonte =Balizia pedicellaris 

Manganaroa alemquerensis (Huber) Speg. =Acacia alemquerensis 

Manganaroa articulata (Ducke) Speg. =Acacia articulata 

Marmaroxylon claviflorum (Benth.) L. Rico =Zygia claviflora 

Marmaroxylon collinum (Sandwith) L. Rico =Zygia collina 

Marmaroxylon eperuetorum (Sandwith) L. Rico =Zygia eperuetorum 

Marmaroxylon racemosum (Ducke) Killip ex Record =Zygia racemosa 

Marmaroxylon ramiflorum (Benth.) L. Rico =Zygia ramiflora 

Mimosa acacioides Benth. [cultivated] =Anadenanthera peregrina (L.) Speg. 

Mimosa adhaerens Kunth =Mimosa debilis 

Mimosa alba Sw. =Inga alba


Mimosa asperata auct. non L. 1759. =Mimosa pigra 

Mimosa asperata L. [not in flora area] 

Mimosa bauhiniaefolia H. Karst. =Mimosa colombiana 

Mimosa bipinnata Aubl. =Entada polystachya 

Mimosa bourgoni Aubl. =Inga bourgonii 

Mimosa bourgonii Aubl. =Inga bourgonii 

Mimosa brevispica Harms =Mimosa invisa var. spiciflora 

Mimosa cataractae Ducke =Mimosa microcephala var. cataractae 

Mimosa cauliflora (Willd.) Poir. =Zygia latifolia var. communis 

Mimosa ceratonia L. [not in flora area] 

Mimosa coriacea Pers. =Inga sertulifera subsp. sertulifera 

Mimosa corymbosa Rich. =Hydrochorea corymbosa 

Mimosa fagifolia L. =Inga laurina 

Mimosa fagifolia Jacq. =Inga laurina 

Mimosa farnesiana L. =Acacia farnesiana 

Mimosa fasciculata var. ernestiana Kuntze =Mimosa arenosa 

Mimosa fastuosa Jacq. =Inga fastuosa 

Mimosa flavescens Splitg. =Mimosa camporum 

Mimosa flaviseta Benth. =Mimosa camporum 

Mimosa flexuosa (Humb. & Bonpl. ex Willd.) Poir. =Acacia macracantha 

Mimosa guianensis Aubl. =Stryphnodendron guianense 

Mimosa hispidula Kunth =Mimosa pudica var. tetrandra 

Mimosa hostmannii Benth. =Mimosa debilis 

Mimosa humilis Humb. & Bonpl. ex Willd. =Mimosa dormiens 

Mimosa hymenaeodes Rich. =Calliandra hymeniodes 

Mimosa inga Vell. =Inga edulis 

Mimosa inga L. =Inga vera 

Mimosa ingoides Rich. =Inga ingoides 

Mimosa intermedia Kunth =Mimosa dormiens 

Mimosa jiramenensis H. Karst. =Mimosa velloziana 

Mimosa latifolia L. =Zygia latifolia 

Mimosa laurina Sw. =Inga laurina 

Mimosa leucocephala Lam. =Leucaena leucocephala 

Mimosa lucida Vahl =Inga pilosula 

Mimosa lutea Mill. =Acacia macracantha 

Mimosa macracantha (Humb. & Bonpl. ex Willd.) Poir. =Acacia macracantha 

Mimosa macrophylla (Humb. & Bonpl. ex Willd.) Poir. =Inga macrophylla 

Mimosa mangensis Jacq. =Chloroleucon mangense 

Mimosa martensis Britton & Killip =Mimosa camporum 

Mimosa micracantha Benth. =Mimosa rufescens var. amnis-nigri 

Mimosa micrantha var. plurijuga Ducke =Mimosa rufescens var. rufescens 

Mimosa microcephala Humb. & Bonpl. ex Willd. =Mimosa microcephala var. microcephala 

Mimosa multijuga (Benth.) Kuntze =Inga multijuga 

Mimosa natans L. f. =Neptunia oleracea 

Mimosa nitida (Willd.) Poir. =Inga pilosula 

Mimosa notata Steud. =Mimosa debilis 

Mimosa obovata Benth. [not in flora area] =Mimosa laticifera Rizzini & Mattos Filho 

Mimosa palpitans Humb. & Bonpl. ex Willd. =Mimosa somnians 

Mimosa panamensis (Benth.) Standl. =Mimosa debilis 

Mimosa paniculata (Willd.) Poir. =Acacia tenuifolia 

Mimosa paniculata Benth. =Mimosa annularis var. odora 

Mimosa parae Poir. =Inga heterophylla 

Mimosa pellita Humb. & Bonpl. ex Willd. =Mimosa pigra 

Mimosa peregrina L. =Anadenanthera peregrina 

Mimosa pigra auct. non L. 1755. =Mimosa pigra 

Mimosa pilosula Rich. =Inga pilosula 

Mimosa plena L. =Neptunia plena 

Mimosa plumiifolia Kleinhoonte =Mimosa microcephala var. cataractae 

Mimosa polystachia L. =Entada polystachya 

Mimosa portoricensis Jacq. =Zapoteca portoricensis 

Mimosa prostrata Lam. [nom. illeg.] =Neptunia oleracea 

Mimosa pulcherrima (Willd.) Poir. =Stryphnodendron pulcherrimum 

Mimosa pusilla Benth. =Mimosa camporum 

Mimosa rosea Vahl =Pithecellobium roseum

374 


Mimosa rubiginosa Rich. =Inga rubiginosa 

Mimosa sagotiana Benth. =Mimosa schrankioides var. sagotiana 

Mimosa saman Jacq. =Samanea saman 

Mimosa santanderensis Britton & Rose ex Britton & Killip =Mimosa schrankioides var. sagotiana 

Mimosa sapindoides (Willd.) Poir. =Inga sapindoides 

Mimosa somniculosa Kunth =Mimosa somnians 

Mimosa spiciflora H. Karst. =Mimosa invisa var. spiciflora 

Mimosa splendens (Willd.) Poir. =Inga splendens 

Mimosa spuria (Humb. & Bonpl. ex Willd.) Poir. =Inga vera subsp. vera 

Mimosa tamarindifolia L. =Acacia tamarindifolia 

Mimosa tergemina L. =Calliandra tergemina 

Mimosa tetrandra Humb. & Bonpl. ex Willd. =Mimosa pudica var. tetrandra 

Mimosa tobagensis Urb. =Mimosa somnians 

Mimosa tomentosa Humb. & Bonpl. ex Willd. =Mimosa hirsutissima 

Mimosa trapezifolia Vahl =Abarema jupunba var. trapezifolia 

Mimosa umbellifera Vahl =Inga umbellifera 

Mimosa unguis-cati L. =Pithecellobium unguis-cati 

Mimosa unijuga Walp. & Duchass. =Mimosa pudica var. unijuga 

Mimosa velloziana var. jiramenensis (H. Karst.) Benth. =Mimosa velloziana 

Mimosa viva Vell. [nom. illeg.] =Mimosa velloziana 

Mimosa viva L. [not in flora area] 

Mimosa xantholasia Benth. =Mimosa arenosa 

Mimosa xinguensis Ducke =Mimosa annularis var. xinguensis 

Neptunia natans (L. f.) Druce =Neptunia oleracea 

Neptunia prostrata (Lam.) Baill. =Neptunia oleracea 

Neptunia surinamensis Steud. =Neptunia plena 

Newtonia psilostachya (DC.) Brenan =Pseudopiptadenia psilostachya 

Newtonia suaveolens (Miq.) Brenan =Pseudopiptadenia suaveolens 

Niopa peregrina (L.) Britton & Rose =Anadenanthera peregrina 

Parkia auriculata Spruce ex Benth. =Parkia discolor 

Parkia oppositifolia Spruce ex Benth. =Parkia nitida 

Parkia pectinata auct. non Benth. 1875: sensu Benth. =Parkia panurensis 

Pentaclethra filamentosa Benth. =Pentaclethra macroloba 

Piptadenia biuncifera Benth. =Piptadenia viridiflora 

Piptadenia cateniformis Ducke =Cedrelinga cateniformis 

Piptadenia guianensis (Aubl.) Benth. =Stryphnodendron guianense 

Piptadenia niopo (Humb. & Bonpl. ex Willd.) Spruce =Anadenanthera peregrina 

Piptadenia peregrina (L.) Benth. =Anadenanthera peregrina 

Piptadenia polystachya Miq. =Stryphnodendron polystachyum 

Piptadenia psilostachya (DC.) Benth. =Pseudopiptadenia psilostachya 

Piptadenia rubescens Pittier =Piptadenia viridiflora 

Piptadenia speciosa Britton & Killip =Piptadenia viridiflora 

Piptadenia suaveolens Miq. =Pseudopiptadenia suaveolens 

Piptadenia tocantina Ducke =Stryphnodendron polystachyum 

Pithecellobium acacioides Ducke =Chloroleucon acacioides 

Pithecellobium adiantifolium (Kunth) Benth. [not in flora area] =Macrosamanea discolor 

Pithecellobium amplissimum Ducke =Macrosamanea amplissima 

Pithecellobium aquaticum (Pittier) R.S. Cowan =Macrosamanea discolor 

Pithecellobium arenarium Ducke =Abarema barbouriana var. arenaria 

Pithecellobium arenicola R.S. Cowan =Macrosamanea discolor 

Pithecellobium barbourianum Standl. =Abarema barbouriana 

Pithecellobium basijugum Ducke [not in flora area] =Zygia basijuga 

Pithecellobium bijugatum Schery =Hydrochorea marginata var. scheryi 

Pithecellobium catenaeformis (Ducke) L. Cardenas =Cedrelinga cateniformis 

Pithecellobium cauliflorum (Willd.) Mart. =Zygia latifolia var. communis 

Pithecellobium collinum Sandwith =Zygia collina 

Pithecellobium corymbosum (Rich.) Benth. =Hydrochorea corymbosa 

Pithecellobium curvicarpum H.S. Irwin =Abarema curvicarpa 

Pithecellobium dinizii Ducke =Zygia ramiflora 

Pithecellobium discolor (Humb. & Bonpl. ex Willd.) J.F. Macbr. =Macrosamanea discolor 

Pithecellobium divaricatum Benth. =Zygia cataractae 

Pithecellobium elegans Ducke [not in flora area] =Balizia elegans (Ducke) Barneby & Grimes 

Pithecellobium eperuetorum Sandwith =Zygia eperuetorum 

Pithecellobium fanshawei Sandwith =Abarema barbouriana


Pithecellobium ferrugineum Benth. =Abarema ferruginea 

Pithecellobium floribundum Spruce ex Benth. =Abarema floribunda 

Pithecellobium forfex (Kunth) Benth. =Pithecellobium roseum 

Pithecellobium glabripetalum H.S. Irwin =Albizia glabripetala 

Pithecellobium glomeratum (DC.) Benth. =Zygia cataractae 

Pithecellobium gonggrijpii Kleinhoonte =Hydrochorea gonggrijpii 

Pithecellobium guachapele (Kunth) J.F. Macbr. =Pseudosamanea guachapele 

Pithecellobium guaricense Pittier =Pithecellobium roseum 

Pithecellobium huberi Ducke =Zygia latifolia 

Pithecellobium inaequale (Humb. & Bonpl. ex Willd.) Benth. =Zygia inaequalis 

Pithecellobium inopinatum (Harms) Ducke =Samanea inopinata 

Pithecellobium jupunba (Willd.) Urb. =Abarema jupunba 

Pithecellobium kegelii Meisn. =Macrosamanea kegelii 

Pithecellobium laetum Benth. =Abarema laeta 

Pithecellobium laetum Benth. =Abarema laeta 

Pithecellobium lanceolatum Benth. [not in flora area] 

Pithecellobium larensis L. Cardenas =Pithecellobium unguis-cati 

Pithecellobium lasiopus Benth. =Zygia latifolia var. lasiopus 

Pithecellobium latifolium (L.) Benth. =Zygia latifolia 

Pithecellobium leucophyllum Spruce ex Benth. =Abarema leucophylla 

Pithecellobium levelii R.S. Cowan =Abarema levelii 

Pithecellobium lindseifolium Spruce ex Benth. =Macrosamanea pubiramea var. lindsaeifolia 

Pithecellobium longepedatum Pittier =Pseudosamanea guachapele 

Pithecellobium longipedunculatum H.S. Irwin =Abarema longipedunculata 

Pithecellobium mangense (Jacq.) J.F. Macbr. [not in flora area] =Chloroleucon mangense (Jacq.) Britton & Rose 

Pithecellobium marginatum Spruce ex Benth. =Hydrochorea marginata 

Pithecellobium mataybifolium Sandwith =Abarema mataybifolia 

Pithecellobium microcalyx Spruce ex Benth. =Abarema microcalyx 

Pithecellobium microchlamys Pittier =Pithecellobium unguis-cati 

Pithecellobium microphyllum (Humb. & Bonpl. ex Willd.) Benth. =Pithecellobium unguis-cati 

Pithecellobium multiflorum (Kunth) Benth. [sensu Benth. 1944] =Albizia subdimidiata 

Pithecellobium niopoides (Benth.) Spruce ex Benth. =Albizia niopoides 

Pithecellobium nuriense H.S. Irwin =Samanea inopinata 

Pithecellobium panurense Spruce ex Benth. =Hydrochorea marginata var. panurensis 

Pithecellobium parviflorum Pittier =Pithecellobium roseum 

Pithecellobium pedicellare (DC.) Benth. =Balizia pedicellaris 

Pithecellobium pilosulum Pittier =Zygia inaequalis 

Pithecellobium polycephalum Benth. [not in flora area] =Albizia polycephala 

Pithecellobium pubescens (DC.) Benth. =Pithecellobium roseum 

Pithecellobium pulchellum Pittier =Pithecellobium unguis-cati 

Pithecellobium pullei Kleinhoonte =Hydrochorea gonggrijpii 

Pithecellobium racemosum Ducke =Zygia racemosa 

Pithecellobium ramiflorum Benth. =Zygia ramiflora 

Pithecellobium saman (Jacq.) Benth. =Samanea saman 

Pithecellobium samanigua (Pittier) J.F. Macbr. =Pseudosamanea guachapele 

Pithecellobium schomburgkii Benth. =Enterolobium schomburgkii 

Pithecellobium simabifolium Spruce ex Benth. =Macrosamanea simabifolia 

Pithecellobium spruceanum Benth. =Macrosamanea spruceana 

Pithecellobium subaquaticum Schery =Macrosamanea discolor 

Pithecellobium trapezifolium (Vahl) Benth. =Abarema jupunba var. trapezifolia 

Pithecellobium unifoliolatum Benth. =Zygia unifoliolata 

Pithecellobium villiferum Ducke =Abarema villifera 

Pithecolobium acreana J.F. Macbr. =Abarema acreana 

Pithecolobium adenophorum Ducke =Abarema adenophora 

Pithecolobium basijugum Ducke =Zygia basijuga 

Pithecolobium cauliflorum (Willd.) Mart. =Zygia latifolia var. communis 

Pithecolobium claviflorum Spruce ex Benth. =Zygia claviflora 

Pithecolobium racemosum Ducke =Zygia racemosa 

Pityrocarpa viridiflora (Kunth) Brenan =Piptadenia viridiflora 

Poponax canescens Britton =Acacia macracantha 

Poponax farnesiana (L.) Raf. =Acacia farnesiana 

Poponax flexuosa (Humb. & Bonpl. ex Willd.) Britton & Rose =Acacia macracantha 

Poponax lutea (Mill.) Britton & Rose =Acacia macracantha 

Poponax macracantha (Humb. & Bonpl. ex Willd.) Killip =Acacia macracantha

376 


Samanea adiantifolia (Kunth) Pittier =Macrosamanea discolor 

Samanea corymbosa (Rich.) Pittier =Hydrochorea corymbosa 

Samanea pedicellaris (DC.) Killip ex Record =Balizia pedicellaris 

Samanea polycephala (Benth.) Pittier [not in flora area] =Albizia polycephala 

Samanea samanigua Pittier =Pseudosamanea guachapele 

Samanea simabifolia (Spruce ex Benth.) Pittier =Macrosamanea simabifolia 

Schrankia leptocarpa DC. =Mimosa quadrivalvis var. leptocarpa 

Senegalia glomerosa (Benth.) Britton & Rose =Acacia glomerosa 

Senegalia paniculata (Willd.) Killip =Acacia tenuifolia 

Senegalia podadenia Britton & Killip =Acacia podadenia 

Senegalia tamarindifolia (L.) Britton & Rose =Acacia tamarindifolia 

Serianthes inopinata Harms =Samanea inopinata 

Stryphnodendron angustum Benth. =Stryphnodendron pulcherrimum 

Stryphnodendron flammatum Kleinhoonte ? =Dimorphandra sp. 

Stryphnodendron floribundum Benth. =Stryphnodendron pulcherrimum 

Stryphnodendron guianense f. floribundum (Benth.) Ducke =Stryphnodendron pulcherrimum 

Stryphnodendron inaequale Benth. =Stryphnodendron microstachyum 

Stryphnodendron melinonis Sagot =Stryphnodendron pulcherrimum 

Stryphnodendron purpureum Ducke =Stryphnodendron microstachyum 

Vachellia farnesiana (L.) Wight & Arn. =Acacia farnesiana 

Zygia cauliflora (Willd.) Killip ex Record =Zygia latifolia 

Zygia divaricata (Benth.) Pittier =Zygia cataractae 

Zygia glomerata (DC.) Pittier =Zygia cataractae 

Zygia trunciflorum (Ducke) L. Rico =Zygia juruana 

Zygia unguis-cati (L.) Sudw. =Pithecellobium unguis-cati 

REFERENCES 

Barneby, R.C. 1991. Sensitivae Censitae. A description of the genus Mimosa (Mimosaceae) in the New World. Mem. New York 

Bot. Gard. 65: 1-835. 

Barneby, R.C. & J. Grimes (in press). Mimosaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A). 

Barneby, R.C., J.W. Grimes, P.E. Berry, D. Brunner, E. Forero, L. Cárdenas, G. De Martino, H.C.F. Hopkins & E.M. de Lamare 

Occhioni. 2001. Mimosaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 580-686. 

Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. Mem. 


Grimes, J.W. 2002. Mimosaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


484-510. 

Hopkins, H.C.F. 1986. Parkia. Flora Neotropica Monogr. 43: 1-123. 

Jansen-Jacobs, M.J. 1976. Mimosaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 611- 

653. 

Kleinhoonte, A. 1940. Mimosaceae. In A. Pulle, Flora of Suriname 2(2): 258-331. 


Poncy, O. 1985. Le genre Inga (Légumineuses, Mimosoideae) en Guyane française. Systématique, morphologie des formes 

juvéniles, écologie. Mém. Mus. Nat. Hist. Nat. (n.s.), Sér. B 31: 1-153. 

Poncy, O. 1991. Le genre Inga dans les Guyanes: notes phytogeographiques. C.R. Soc. Biogéogr. 67(2): 95-111. 

Poncy, O. 1991. Deux nouvelles espèces de Inga (Mimosaceae) et notes nomenclaturales et taxonomiques sur trois autres espèces. 

Bull. Mus. natl. Hist. nat., Paris, sér. 4, 13, sect. B, Adansonia, 3-4: 147-154. 

Genlisea filiformis A. St.-Hil. 

Genlisea glabra P. Taylor 

Genlisea guianensis N.E. Br. 

Genlisea pygmaea A. St.-Hil. 

Genlisea repens Benj. 

Genlisea roraimensis N.E. Br. 

Genlisea sanariapoana Steyerm. 

Utricularia adpressa Salzm. ex A. St.-Hil. & Girard 

Utricularia alpina Jacq. 

Utricularia amethystina Salzm. ex A. St.-Hil. & Girard 

Utricularia aureomaculata Steyerm. 

Utricularia benjaminiana Oliv. 

LENTIBULARIACEAE 

P.G. Taylor 


BO 

AM BO GU 


AM BO GU SU 

AM BO GU 

AM 

BO GU SU FG 

AM BO GU SU 


BO 

AM BO GU SU FG

Utricularia breviscapa C. Wright ex Griseb. 

Utricularia calycifida Benj. 

Utricularia campbelliana Oliv. 

Utricularia chiribiquitensis A. Fern. 

Utricularia choristotheca P. Taylor 

Utricularia costata P. Taylor 

Utricularia cucullata A. St.-Hil. & Girard 

Utricularia determannii P. Taylor 

Utricularia erectiflora A. St.-Hil. & Girard 

Utricularia fimbriata Kunth 

Utricularia foliosa L. 

Utricularia gibba L. 

Utricularia guyanensis A. DC. 

Utricularia heterochroma Steyerm. 

Utricularia hispida Lam. 

Utricularia humboldtii R.H. Schomb. 

Utricularia hydrocarpa Vahl 

Utricularia jamesoniana Oliv. 

Utricularia juncea Vahl 

Utricularia lloydii Merl 

Utricularia longeciliata A. DC. 

Utricularia mirabilis P. Taylor 

Utricularia myriocista A. St.-Hil. & Girard 

Utricularia nana A. St.-Hil. & Girard 

Utricularia naviculata P. Taylor 

Utricularia neottioides A. St.-Hil. & Girard 

Utricularia nervosa Weber ex Benj. 

Utricularia olivacea C. Wright ex Griseb. 

Utricularia oliveriana Steyerm. 

Utricularia pubescens Sm. 

Utricularia pusilla Vahl 

Utricularia quelchii N.E. Br. 

Utricularia sandwithii P. Taylor 

Utricularia schultesii A. Fern. 

Utricularia simulans Pilg. 

Utricularia spruceana Benth. ex Oliv. 

Utricularia steyermarkii P. Taylor 

Utricularia subulata L. 

Utricularia tenuissima Tutin 

Utricularia trichophylla Spruce ex Oliv. 

Utricularia tricolor A. St.-Hil. 

Utricularia triloba Benj. 

Utricularia viscosa Spruce ex Oliv. 



Akentra inflata Benj. =Utricularia benjaminiana 

Aranella fimbriata (Kunth) Barnhart =Utricularia fimbriata 

Avesicaria neottioides (A. St.-Hil. & Girard) Barnhart =Utricularia neottioides 

Biovularia olivacea (C. Wright ex Griseb.) Kamienski =Utricularia olivacea 

Calpidisca amethystina (Salzm. ex A. St.-Hil. & Girard) Barnhart =Utricularia amethystina 

Calpidisca calycifida (Benj.) Gleason =Utricularia calycifida 

Calpidisca hispida (Lam.) Barnhart =Utricularia hispida 

Calpidisca humboldtii (R.H. Schomb.) Gleason =Utricularia humboldtii 

Calpidisca modesta (A. DC.) Barnhart =Utricularia amethystina 

Calpidisca roraimensis (N.E. Br.) Gleason =Utricularia amethystina 

Genlisea anfractuosa Tutin =Genlisea filiformis 

Genlisea esmeraldae Steyerm. =Genlisea pygmaea 

Genlisea nigrocaulis Steyerm. [based on mixed collection] =Genlisea pygmaea and Genlisea repens 

Genlisea pulchella Tutin =Genlisea repens 

Lecticula spruceana (Benth. ex Oliv.) Barnhart =Utricularia spruceana 

Orchyllium alpinum (Jacq.) Barnhart =Utricularia alpina 

Orchyllium humboldtii (R.H. Schomb.) Barnhart =Utricularia humboldtii 

Orchyllium quelchii (N.E. Br.) Gleason =Utricularia quelchii 

Orchyllium schimperi (Schenck) Barnhart =Utricularia jamesoniana 

Polypompholyx laciniata Benj. =Utricularia simulans 

AM BO GU 


AM BO GU 

AM BO 

SU FG 

BO 


SU 

BO GU SU 

AM BO 




BO 


AM BO GU 




AM SU FG 


AM 



AM 

AM BO 

AM BO 

AM BO GU SU 

AM BO 

AM BO GU FG 

BO GU SU FG 

AM BO GU 

AM BO GU SU 

AM 


AM 

BO 



BO GU SU FG 

AM 


AM BO GU SU FG

378 


Polypompholyx schomburgkii Klotzsch =Utricularia longeciliata 

Setiscapella pusilla (Vahl) Barnhart ex Britton & P. Wilson =Utricularia pusilla 

Setiscapella subulata (L.) Barnhart =Utricularia subulata 

Stomoisia juncea (Vahl) Barnhart =Utricularia juncea 

Stomoisia spicata (Sylvén) Gleason =Utricularia erectiflora 

Utricularia alutacea Tutin =Utricularia amethystina 

Utricularia amazonasana Steyerm. =Utricularia hydrocarpa 

Utricularia amethystina f. alutacea (Tutin) Steyerm. =Utricularia amethystina 

Utricularia angulosa Poir. =Utricularia juncea 

Utricularia angustifolia Benj. =Utricularia hispida 

Utricularia arenicola Tutin =Utricularia nana 

Utricularia arenicola var. kavanayena Steyerm. =Utricularia nana 

Utricularia aureolimba Steyerm. =Utricularia adpressa 

Utricularia ayacuchae Steyerm. =Utricularia cucullata 

Utricularia bolivarana Steyerm. =Utricularia amethystina 

Utricularia campbelliana var. minor Steyerm. =Utricularia campbelliana 

Utricularia coccinea Benj. =Utricularia hydrocarpa 

Utricularia concinna N.E. Br. =Utricularia jamesoniana 

Utricularia congesta Steyerm. =Utricularia simulans 

Utricularia congesta f. deminutiva Steyerm. =Utricularia simulans 

Utricularia connellii N.E. Br. =Utricularia pubescens 

Utricularia cuspidata Steyerm. =Utricularia calycifida 

Utricularia dubia Benj. =Utricularia nana 

Utricularia flaccida auct. non A. DC.: sensu P. Taylor, Mem. New York Bot. Gard. 17(1) =Utricularia nervosa 

Utricularia flaccida A. DC. [not in flora area] 

Utricularia fockeana Miq. =Utricularia hydrocarpa 

Utricularia guianensis Splitg. ex de Vries =Utricularia foliosa 

Utricularia hirtella A. St.-Hil. & Girard =Utricularia amethystina 

Utricularia humboldtii f. cuneata Steyerm. =Utricularia humboldtii 

Utricularia inflata Walter [sensu stricto, not in flora area] 

Utricularia kaieteurensis Steyerm. =Utricularia amethystina 

Utricularia leptantha Benj. =Utricularia pusilla 

Utricularia magnifica Pilg. =Utricularia myriocista 

Utricularia maguirei Steyerm. =Utricularia calycifida 

Utricularia modesta A. DC. =Utricularia amethystina 

Utricularia montana Jacq. =Utricularia alpina 

Utricularia muscosa Benj. =Utricularia calycifida 

Utricularia obovata Miq. =Utricularia amethystina 

Utricularia obtusa Sw. =Utricularia gibba 

Utricularia oligosperma A. St.-Hil. =Utricularia foliosa 

Utricularia orinocensis Steyerm. =Utricularia simulans 

Utricularia parkeriana A. DC. =Utricularia gibba 

Utricularia peckii S.F. Blake =Utricularia guyanensis 

Utricularia pectinata Splitg. ex de Vries =Utricularia longeciliata 

Utricularia peltata Spruce ex Oliv. =Utricularia pubescens 

Utricularia praelonga A. St.-Hil. & Girard [not in flora area] 

Utricularia punctifolia Benj. =Utricularia amethystina 

Utricularia purpurea Walter [not in flora area] 

Utricularia resupinata P. Taylor =Utricularia spruceana 

Utricularia roraimensis N.E. Br. =Utricularia amethystina 

Utricularia rubricaulis Tutin =Utricularia guyanensis 

Utricularia schimperi Schenck =Utricularia jamesoniana 

Utricularia sciaphila Tutin =Utricularia pubescens 

Utricularia spatulata Splitg. ex de Vries =Utricularia amethystina 

Utricularia spatulifolia Pilg. =Utricularia amethystina 

Utricularia spicata Sylvén =Utricularia erectiflora 

Utricularia stricta G. Mey. =Utricularia juncea 

Utricularia subpeltata Steyerm. =Utricularia pubescens 

Utricularia tenuifolia Benj. =Utricularia gibba 

Utricularia tenuiscapa Pilg. =Utricularia subulata 

Utricularia tepuiana Steyerm. =Utricularia amethystina 

Utricularia turumiquirensis Steyerm. =Utricularia amethystina 

Utricularia venezuelana Steyerm. =Utricularia pubescens 

Utricularia versicolor Benj. =Utricularia amethystina


Vesiculina cucullata (A. St.-Hil. & Girard) Barnhart =Utricularia cucullata 

REFERENCES 

Smith, N.P. 2002. Lentibulariaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


397-398. 

Taylor, P. 1967. Lentibulariaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 7. Mem. New York Bot. 

Gard. 17(1): 1-439. 

Taylor, P. 1989. The genus Utricularia - a taxonomic monograph. Kew Bull. Add. Ser. XIV: 1-724. 

Taylor, P. 1999. Lentibulariaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

5: 782-803. 

Lissocarpa benthamii Gürke 

Lissocarpa guianensis Gleason 

Lissocarpa stenocarpa Steyerm. 

LISSOCARPACEAE 

P.E. Berry 

REFERENCES 

Berry, P.E. 2001. Lissocarpaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 19-21. 

Klaprothia mentzelioides Kunth 

Mentzelia aspera L. 

LOASACEAE 

J.S. Miller 

REFERENCES 

Miller, J.S. 2001. Loasaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 6: 

22. 

LOGANIACEAE 

P.E. Berry, A.E. Brant, A.J.M. Leeuwenberg 

Antonia ovata Pohl 

Bonyunia aquatica Ducke 

Bonyunia minor N.E. Br. 

Bonyunia superba M.R. Schomb. ex Progel [Mt. Roraima] 

Mitreola petiolata (J.F. Gmel.) Torr. & A. Gray 

Mostuea surinamensis Benth. [expected in FG] 

Polypremnum procumbens L. 

Spigelia anthelmia L. 

Spigelia flemmingiana Cham. & Schltdl. 

Spigelia gracilis A. DC. 

Spigelia guianensis (Aubl.) Lemée [incertae sedis; excluded from Speigelia 

and Loganiaceae by Hurley] 

Spigelia hamelioides Kunth 

Spigelia multispica Steud. 

Spigelia polystachya Klotzsch ex Progel 

Strychnos barnhartiana Krukoff 

Strychnos brachiata Ruiz & Pav. 

Strychnos bredemeyeri (Schult. & Schult. f.) Sprague & Sandwith 

Strychnos castelnaeana Wedd. 

Strychnos cayennensis Krukoff & Barneby 

Strychnos cogens Benth. 

Strychnos davidsei Krukoff & Barneby 

Strychnos diaboli Sandwith 

Strychnos divaricans Ducke 

Strychnos erichsonii M.R. Schomb. ex Progel 

Strychnos eugeniifolia Monach. 

Strychnos fendleri Sprague & Sandwith 

Strychnos glabra Sagot ex Progel 

AM GU 

GU 

AM 

AM 

FG? 


AM 

AM BO GU 

BO GU 

SU FG? 

SU 

SU? 


GU 

AM 

AM BO GU FG 

AM 

FG 

BO GU SU FG 

BO GU 

BO 


FG? 

FG 


BO 

AM BO GU 

SU FG 


SU FG 

BO 

GU SU FG

380 

Strychnos guianensis (Aubl.) Mart. 

Strychnos hirsuta Spruce ex Benth. 

Strychnos jobertiana Baill. 

Strychnos mattogrossensis S. Moore 

Strychnos medeola Sagot ex Progel 

Strychnos melinoniana Baill. 

Strychnos mitscherlichii M.R. Schomb. 

Strychnos mitscherlichii var. mitscherlichii 

Strychnos oiapocensis Fróes 

Strychnos panamensis Seem. 

Strychnos panurensis Sprague & Sandwith 

Strychnos peckii B.L. Rob. 

Strychnos ramentifera Ducke 

Strychnos rondeletioides Spruce ex Benth. 

Strychnos subcordata Spruce ex Benth. [expected in FG] 

Strychnos tomentosa Benth. 

Strychnos toxifera R.H. Schomb. ex Benth. 



Antonia ovata var. excelsa Paula =Antonia ovata 

Antonia ovata var. pilosa (Hook.) Progel =Antonia ovata 

Antonia pilosa Hook. =Antonia ovata 

Antonia pubescens Bong. =Antonia ovata 

Bonyunia cinchonoides Gleason & Standl. =Bonyunia minor 

Lasiostoma bredemeyeri Schult. & Schult. f. =Strychnos bredemeyeri 

Lasiostoma curare Kunth =Strychnos guianensis 

Montira guianensis Aubl. [excluded from Loganiaceae by Hurley] =Spigelia guianensis 

Mostuea brasiliensis Huber =Mostuea surinamensis 

Rouhamon bredemeyeri (Schult. & Schult. f.) A. DC. =Strychnos bredemeyeri 

Rouhamon guianensis Aubl. =Strychnos guianensis 

Rouhamon pedunculatum A. DC. =Strychnos bredemeyeri 

Spigelia filipes Rusby =Spigelia multispica 

Spigelia humboldtiana Cham. & Schltdl. [not in flora area] =Spigelia scabra Cham. & Schlechtend. 

Spigelia humilis Benth. =Spigelia guianensis 

Spigelia nervosa Steud. =Spigelia anthelmia 

Spigelia schomburgkiana Benth. =Spigelia flemmingiana 

Strychnos blackii Ducke =Strychnos brachiata 

Strychnos bovetiana Pires =Strychnos erichsonii 

Strychnos curare (Kunth) Benth. =Strychnos guianensis 

Strychnos darienensis Seem. =Strychnos bredemeyeri 

Strychnos lanceolata Spruce ex Benth. =Strychnos guianensis 

Strychnos pedunculata (A. DC.) Benth. =Strychnos bredemeyeri 

Strychnos rouhamon Benth. =Strychnos guianensis 

Strychnos ruizii Sprague & Sandwith =Strychnos brachiata 

Strychnos schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum] =Strychnos bredemeyeri 

Strychnos smilacina Benth. =Strychnos mitscherlichii 

Strychnos smithiana Krukoff =Strychnos rondeletioides 

Strychnos solerederi Gilg =Strychnos mitscherlichii var. mitscherlichii 

Strychnos syntoxica Sprague & Sandwith =Strychnos toxifera 

Strychnos trichostyla Ducke =Strychnos jobertiana 

Strychnos trinitensis Griseb. =Strychnos bredemeyeri 

Strychnos urbanii Barb. Rodr. =Strychnos erichsonii 


GU FG 

AM BO FG 

AM BO DA 

SU FG 

GU SU FG 



SU FG 

AM BO GU FG 

AM BO FG 


SU FG? 

AM BO SU FG 

GU 

GU SU FG 


REFERENCES 

Bordenave, B. 2002. Loganiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


398-405. 

Hurley, H.H. 1967. A taxonomic revision of the genus Spigelia. George Washington University, Washington, DC. (Unpublished 

Ph.D. thesis) 

Krukoff, B.A. & J. Monachino. 1942. The American species of Strychnos. Brittonia 4: 248-322. (See also the subsequent series 

of supplementary notes in various journals.) 

Leeuwenberg, A.J.M. 1961. The Loganiaceae of Africa II. A revision of Mostuea Didr. Meded. Landb. Wag. 61(4): 1-31. 

Leeuwenberg, A.J.M. 1969. Notes on American Loganiaceae. III. Revision of Bonyunia Rich. Schomb. Acta Bot. Neerl. 18(1): 

152-158. 

Leeuwenberg, A.J.M. 1971. Notes on American Loganiaceae. V. Key to the genera. Acta Bot. Neerl. 20(5): 539-542.


Leeuwenberg, A.J.M. 1975. The Loganiaceae of Africa XII. A revision of Mitreola L. Meded. Landb. Wag. 74(23): 1-28. 

Leeuwenberg, A.J.M. (ed.). 1980. Loganiaceae. In Engler & Prantl, Die Natürlichen Pflanzenfamilian 28bI: 1-255. 

Leeuwenberg, A.J.M., P.E. Berry, A.E. Brant. 2001. Loganiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Raalte, M.H. van. 1932. Loganiaceae. In A. Pulle, Flora of Suriname 4(1): 103-110. 

Smith, L.B. 1960. Notes on South American phanerogams - III. Wrightia 2(2): 90-102. 

Uittien, H. 1937. Loganiaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 472-474. 

Cladocolea elliptica Kuijt 

Cladocolea intermedia (Rizzini) Kuijt 

Cladocolea micrantha (Eichler) Kuijt 

Cladocolea nitida Kuijt 

Cladocolea roraimensis (Steyerm.) Kuijt 

Cladocolea sandwithii (Maguire) Kuijt 

Gaiadendron punctatum (Ruiz & Pav.) G. Don 

Ixocactus clandestinus (Mart.) Kuijt 

Oryctanthus alveolatus (Kunth) Kuijt 

Oryctanthus florulentus (Rich.) Tiegh. 

Oryctanthus occidentalis (L.) Eichler 

Oryctanthus occidentalis subsp. continentalis Kuijt 

Oryctanthus phthirusoides Rizzini 

Oryctanthus spicatus (Jacq.) Eichler 

Oryctina myrsinites (Eichler) Kuijt 

Phthirusa bisexualis Rizzini 

Phthirusa coarctata A.C. Sm. 

Phthirusa disjectifolia (Rizzini) Kuijt 

Phthirusa guyanensis Eichler 

Phthirusa monetaria Sandwith 

Phthirusa nitens (Mart.) Eichler 

Phthirusa podoptera (Cham. & Schltdl.) Kuijt 

Phthirusa pycnostachya Eichler 

Phthirusa pyrifolia (Kunth) Eichler 

Phthirusa rufa (Mart.) Eichler 

Phthirusa stelis (L.) Kuijt 

Phthirusa stenophylla Eichler 

Psittacanthus acinarius (Mart.) Mart. 

Psittacanthus biternatus (Hoffmanns.) Blume 

Psittacanthus brachynema Eichler 

Psittacanthus cinctus (Mart.) Mart. 

Psittacanthus clusiifolius (Willd.) Eichler 

Psittacanthus collum-cygni Eichler 

Psittacanthus cordatus (Hoffmanns.) Blume 

Psittacanthus corynocephalus Eichler 

Psittacanthus cucullaris (Lam.) Blume 

Psittacanthus julianus Rizzini 

Psittacanthus lamprophyllus Eichler 

Psittacanthus lasianthus Sandwith 

Psittacanthus leptanthus Sandwith 

Psittacanthus melinonii (Tiegh.) Engl. 

Psittacanthus montis-neblinae Rizzini 

Psittacanthus peronopetalus Eichler 

Psittacanthus plagiophyllus Eichler 

Psittacanthus redactus Rizzini 

Psittacanthus rhynchanthus (Benth.) Kuijt 

Psittacanthus robustus (Mart.) Mart. 

Struthanthus concinnus Mart. 

Struthanthus confertus Mart. 

Struthanthus dichotrianthus Eichler 

Struthanthus eichlerianus Rizzini 

Struthanthus elegans Mart. 

Struthanthus gracilis (Gleason) Steyerm. & Maguire 

LORANTHACEAE 

J. Kuijt 

SU 

BO 

BO 

BO 

GU 

GU 

GU 

SU FG 

AM BO GU 

GU 



AM GU SU 

BO 

GU 

GU 

SU 

AM 

AM 

GU 

GU 

SU FG 

AM BO GU SU 

AM GU 

GU 

SU FG 

AM 

AM 

BO GU 

GU FG 


GU SU 


AM GU 

AM 

AM 

AM 

AM 

BO DA 

AM GU SU? 

AM BO SU 

GU FG 

AM FG 

AM 

GU? SU FG 

FG 

AM 

AM 

GU 

GU 

SU 

FG 

AM FG 

BO 

GU SU FG 

FG 

AM GU 

GU SU 

SU? 

AM BO 

BO 

DA GU 

GU 

SU FG 

AM BO GU

382 


Struthanthus marginatus (Desr.) Blume 

Struthanthus phyllyraeoides (Kunth) Blume 

Struthanthus syringifolius (Mart.) Mart. 

Struthanthus vulgaris Eichler 

Tripodanthus acutifolius (Ruiz & Pav.) Tiegh. [expected in the Guianas] 

GU 

GU 

BO GU SU FG 

BO GU 

BO 


Apodina cucullaris (Lam.) Tiegh. =Psittacanthus cucullaris 

Cladocolea apiculata Kuijt =Cladocolea intermedia 

Cladocolea clandestina (Mart.) Kuijt =Ixocactus clandestinus 

Furarium disjectifolium Rizzini =Phthirusa disjectifolia 

Gaiadendron lanceolatum (Ruiz & Pav.) Baehni =Gaiadendron punctatum 

Gaiadendron poasense Donn. Sm. =Gaiadendron punctatum 

Gaiadendron punctatum (Ruiz & Pav.) G. Don var. puracense (Kunth) Steyerm. =Gaiadendron punctatum 

Gaiadendron puracensis (Kunth) G. Don =Gaiadendron punctatum 

Gaiadendron revolutum Rizzini =Gaiadendron punctatum 

Gaiadendron tagua (Kunth) G. Don =Gaiadendron punctatum 

Gaiadendron tagua var. reductum Rizzini =Gaiadendron punctatum 

Gaiadendron tagua var. revolutum (Rizzini) Rizzini =Gaiadendron punctatum 

Loranthus acinarius Mart. =Psittacanthus acinarius 

Loranthus acutifolius Ruiz & Pav. =Tripodanthus acutifolius 

Loranthus aduncus G. Mey. =Phthirusa stelis 

Loranthus alveolatus Kunth =Oryctanthus alveolatus 

Loranthus amplexicaulis Kunth =Oryctanthus alveolatus 

Loranthus biternatus Hoffmanns. =Psittacanthus biternatus 

Loranthus cinctus Mart. =Psittacanthus cinctus 

Loranthus conduplicatus Kunth =Phthirusa stelis 

Loranthus cucullaris Lam. =Psittacanthus cucullaris 

Loranthus cupulifer Kunth =Psittacanthus cucullaris 

Loranthus eugenioides Kunth =Tripodanthus acutifolius 

Loranthus falcifrons Mart. =Psittacanthus cucullaris 

Loranthus florulentus Rich. =Oryctanthus florulentus 

Loranthus formosus Cham. & Schltdl. =Psittacanthus robustus 

Loranthus guadalupensis DC. (Rubiaceae) =Notopleura guadalupensis 

Loranthus lanceolatum Ruiz & Pav. =Gaiadendron punctatum 

Loranthus magdalenae Cham. & Schltdl. =Phthirusa stelis 

Loranthus nitens Mart. =Phthirusa nitens 

Loranthus occidentalis L. =Oryctanthus occidentalis 

Loranthus orinocensis Spreng. =Phthirusa stelis 

Loranthus paniculatus Kunth =Phthirusa stelis 

Loranthus patrisii DC. (Combretaceae) =Combretum sp. 

Loranthus podopterus Cham. & Schltdl. =Phthirusa podoptera 

Loranthus pterygopus Mart. =Phthirusa podoptera 

Loranthus punctatus Ruiz & Pav. =Gaiadendron punctatum 

Loranthus puracensis Kunth =Gaiadendron punctatum 

Loranthus pyrifolius Kunth =Phthirusa pyrifolia 

Loranthus retroflexus Ruiz & Pav. =Phthirusa stelis 

Loranthus rhynchanthus Benth. =Psittacanthus rhynchanthus 

Loranthus robustus Mart. =Psittacanthus robustus 

Loranthus ruficaulis Poepp. & Endl. =Oryctanthus florulentus 

Loranthus speciosus Pohl =Psittacanthus robustus 

Loranthus stelis L. =Phthirusa stelis 

Loranthus syringifolius Mart. =Struthanthus syringifolius 

Loranthus tagua Kunth =Gaiadendron punctatum 

Notanthera lanceolata (Ruiz & Pav.) G. Don =Gaiadendron punctatum 

Oryctanthus alveolatus var. amplexicaulis (Kunth) Rizzini =Oryctanthus alveolatus 

Oryctanthus alveolatus var. kuijtii Rizzini =Oryctanthus alveolatus 

Oryctanthus amplexicaulis (Kunth) Eichler =Oryctanthus alveolatus 

Oryctanthus botryostachys Eichler =Oryctanthus alveolatus 

Oryctanthus laceratus Kuijt =Oryctanthus alveolatus 

Oryctanthus ruficaulis (Poepp. & Endl.) Eichler =Oryctanthus florulentus 

Passovia suaveolens Karst. =Phthirusa stelis 

Passowia guyanensis (Eichler) Tiegh. =Phthirusa guyanensis 

Passowia micrantha (Eichler) Tiegh. =Cladocolea micrantha


Passowia myrsinites (Eichler) Tiegh. =Oryctina myrsinites 

Passowia orinocensis (Spreng.) Tiegh. =Phthirusa stelis 

Passowia pyrifolia (Kunth) Tiegh. =Phthirusa pyrifolia 

Phoradendron karuaianum Steyerm. =Dendrophthora karuaiana 

Phrygilanthus acutifolius (Ruiz & Pav.) Eichler =Tripodanthus acutifolius 

Phrygilanthus acutifolius subsp. coriaceus Rizzini =Tripodanthus acutifolius 

Phrygilanthus acutifolius subsp. eugenioides lateovatus Rizzini =Tripodanthus acutifolius 

Phrygilanthus acutifolius subsp. eugenioides vulgaris Rizzini =Tripodanthus acutifolius 

Phrygilanthus lanceolatum (Ruiz & Pav.) Eichler =Gaiadendron punctatum 

Phrygilanthus megathermicus Rizzini =Phthirusa stenophylla 

Phrygilanthus punctatus (Ruiz & Pav.) Eichler =Gaiadendron punctatum 

Phthirusa adunca (G. Mey.) Maguire =Phthirusa stelis 

Phthirusa adunca var. magdalenae (Cham. & Schltdl.) Rizzini =Phthirusa stelis 

Phthirusa adunca var. orinocensis (Spreng.) Steyerm. =Phthirusa stelis 

Phthirusa adunca var. rigidifolia Rizzini =Phthirusa stelis 

Phthirusa anastyla Rizzini =Phthirusa stelis 

Phthirusa angulata K. Krause =Phthirusa stelis 

Phthirusa bernardiana Rizzini =Cladocolea micrantha 

Phthirusa calloso-albida Rizzini =Phthirusa guyanensis 

Phthirusa castillana Rizzini =Phthirusa guyanensis 

Phthirusa cothurnata Rizzini =Phthirusa stelis 

Phthirusa gracilis Gleason =Struthanthus gracilis 

Phthirusa hippocrateoides Steyerm. & Maguire =Struthanthus syringifolius 

Phthirusa huberi Rizzini =Phthirusa stenophylla 

Phthirusa intermedia Rizzini =Cladocolea intermedia 

Phthirusa maculata Rizzini =Oryctanthus phthirusoides 

Phthirusa maritima Rizzini =Phthirusa stelis 

Phthirusa megathermica (Rizzini) Rizzini =Phthirusa stenophylla 

Phthirusa micrantha Eichler =Cladocolea micrantha 

Phthirusa micrantha var. bolivariensis Rizzini =Cladocolea micrantha 

Phthirusa myrsinites Eichler =Oryctina myrsinites 

Phthirusa myrsinites var. savannarum (Maguire) Rizzini =Oryctina myrsinites 

Phthirusa orinocensis (Spreng.) Eichler =Phthirusa stelis 

Phthirusa paniculata (Kunth) J.F. Macbr. =Phthirusa stelis 

Phthirusa pauciflora Eichler [not in flora area] =Dendropemon pauciflorus (Swartz) Tiegh. 

Phthirusa percrassa Rizzini =Phthirusa stenophylla 

Phthirusa perdivergens Rizzini =Phthirusa stelis 

Phthirusa perforata Rizzini =Phthirusa guyanensis 

Phthirusa phaeocladus Eichler =Phthirusa guyanensis 

Phthirusa ptariana Steyerm. =Struthanthus syringifolius 

Phthirusa punctata Gleason =Phthirusa stelis 

Phthirusa pyramidalis Rizzini =Phthirusa stelis 

Phthirusa retroflexa (Ruiz & Pav.) Kuijt =Phthirusa stelis 

Phthirusa roraimense Steyerm. =Cladocolea roraimensis 

Phthirusa rubromicans Rizzini =Phthirusa stelis 

Phthirusa sandwithii Maguire =Cladocolea sandwithii 

Phthirusa savannarum Maguire =Oryctina myrsinites 

Phthirusa seitzii Krug & Urb. =Phthirusa stelis 

Phthirusa squamulosa (Pohl) Klotzsch ex Eichler =Phthirusa rufa 

Phthirusa theobromae (Willd.) Eichler =Phthirusa stelis 

Psittacanthus calyculatus var. wurdackii Rizzini =Psittacanthus rhynchanthus 

Psittacanthus chrismarii Urb. =Psittacanthus rhynchanthus 

Psittacanthus cinctus var. guainiae Rizzini =Psittacanthus cinctus 

Psittacanthus clusiifolius var. pseudojulianus Rizzini =Psittacanthus clusiifolius 

Psittacanthus cupulifer (Kunth) Klotzsch =Psittacanthus cucullaris 

Psittacanthus falcifrons (Mart.) Mart. =Psittacanthus cucullaris 

Psittacanthus intermedius Rizzini =Psittacanthus robustus 

Psittacanthus rhynchanthus subsp. wurdackii (Rizzini) Kuijt =Psittacanthus rhynchanthus 

Psittacanthus semiarticulatus Rizzini =Psittacanthus rhynchanthus 

Psittacanthus warmingii Eichler =Psittacanthus acinarius 

Struthanthus chimantensis Steyerm. & Maguire =Struthanthus gracilis 

Struthanthus cupulifer Rizzini =Struthanthus gracilis 

Struthanthus dichotrianthus var. dichotrianthus =Struthanthus dichotrianthus 

Struthanthus dichotrianthus var. lasserianus Rizzini =Struthanthus dichotrianthus

384 


Struthanthus dissimilis Rizzini =Struthanthus dichotrianthus 

Struthanthus gracilis var. mucronatus (Steyerm.) Rizzini =Struthanthus gracilis 

Struthanthus granulatus Rizzini =Struthanthus dichotrianthus 

Struthanthus micranthus (Eichler) Baehni & J.F. Macbr. =Cladocolea micrantha 

Struthanthus mucronatus Steyerm. =Struthanthus gracilis 

Struthanthus nitens (Mart.) Mart. =Phthirusa nitens 

Struthanthus pseudolepidotus Rizzini =Phthirusa nitens 

Struthanthus pterygopus (Mart.) Mart. =Phthirusa podoptera 

Struthanthus retroflexa (Ruiz & Pav.) Kuijt =Phthirusa stelis 

Struthanthus squamulosus Klotzsch ex Eichler =Phthirusa rufa 

Struthanthus terniflorus Eichler =Struthanthus dichotrianthus 

Struthanthus translucens Rizzini =Struthanthus dichotrianthus 

Struthanthus umbellatus Kuijt =Phthirusa pycnostachya 

Struthanthus yavitensis Rizzini =Struthanthus dichotrianthus 

Taguaria puracense (Kunth) Raf. =Gaiadendron punctatum 

Tripodanthus eugenioides (Kunth) Tiegh. =Tripodanthus acutifolius 

REFERENCES 

Krause, K. 1932. Loranthaceae. In A. Pulle, Flora of Suriname 1(1): 4-24. 

Kuijt, J. 1976. Revision of the genus Oryctanthus. Bot. Jahrb. Syst. 95: 478-534. 

Kuijt, J. 1985. The genus Cladocolea. J. Arn. Arb. 56: 265-335. 

Kuijt, J. 2001. Loranthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 6: 

37-59. 

Kuijt, J. 2002. Loranthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


408. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Loranthaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 295-300. 

Rizzini, C.T. 1978. Loranthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 10. Mem. New York Bot. 

Gard. 29: 23-36. 

Rizzini, C.T. 1982. Loranthaceae. In Z. Luces de Febres, Flora de Venezuela 4(2): 7-316. 

Ammannia auriculata Willd. [expected in the Guianas] 

Ammannia coccinea Rottb. [expected in the Guianas] 

Ammannia latifolia L. 

Crenea maritima Aubl. 

Cuphea anagalloidea A. St.-Hil. 

Cuphea anisoclada Lourteig 

Cuphea annulata Koehne 

Cuphea antisyphilitica Kunth 

Cuphea antisyphilitica var. acutifolia Benth. 

Cuphea antisyphilitica var. antisyphilitica 

Cuphea blackii Lourteig 

Cuphea bolivariensis Lourteig 

Cuphea cardonae Lourteig 

Cuphea carthagenensis (Jacq.) J.F. Macbr. 

Cuphea cataractarum Spruce ex Koehne 

Cuphea curiosa Lourteig 

Cuphea curiosa var. curiosa 

Cuphea curiosa var. oresbia Lourteig 

Cuphea dactylophora Koehne 

Cuphea distichophylla Lourteig 

Cuphea elliptica Koehne 

Cuphea galeato-calcarata Lourteig 

Cuphea gracilis Kunth 

Cuphea insolita Lourteig 

Cuphea maigualidensis Lourteig 

Cuphea melvilla Lindl. 

Cuphea micrantha Kunth 

Cuphea odonellii Lourteig 

Cuphea pleiantha Lourteig 

LYTHRACEAE 

A. Lourteig† 

GU FG 

DA GU 

GU 

SU FG 

AM BO 

AM BO 


AM BO GU 


FG 

BO 

BO 

GU 

AM 

BO GU SU FG 

AM BO 

AM BO 

AM BO 

AM 

AM 

BO GU 

AM 

AM 

BO DA 

AM GU 

GU 

AM BO 


BO GU 

AM 

AM 

BO


Cuphea repens Koehne [expected in the Guianas] 

Cuphea rhodocalyx Lourteig 

Cuphea rigidula Benth. 

Cuphea tenuissima Koehne 

Lafoensia vandelliana Cham. & Schltdl. [expected in the Guianas] 

Lagerstroemia speciosa (L.) Pers. [cultivated or escaped from cultivation] 

Lawsonia inermis L. 

Pehria compacta (Rusby) Sprague 

Rotala mexicana Cham. & Schltdl. 

Rotala ramosior (L.) Koehne [expected in the Guianas] 


Ammannia ramosior L. =Rotala ramosior 

Crenea surinamensis (L. f.) Koehne =Crenea maritima 

Cuphea antisyphilitica f. gracillima Koehne =Cuphea antisyphilitica var. acutifolia 

Cuphea antisyphilitica f. subhirsuta Koehne =Cuphea antisyphilitica 

Cuphea balsamona Cham. & Schltdl. =Cuphea carthagenensis 

Cuphea gracilis auct. non Kunth 1824, nec Seem. 1854. =Cuphea antisyphilitica 

Cuphea gracilis var. major Koehne =Cuphea antisyphilitica var. acutifolia 

Cuphea gracilis Kunth var. media Koehne =Cuphea gracilis 

Cuphea mimuloides Cham. & Schltdl. var. guianensis Koehne =Cuphea anagalloidea 

Cuphea multicaulis Koehne =Cuphea gracilis 

Cuphea pauciflora Koehne =Cuphea gracilis 

Cuphea rhodocalyx var. setosa Lourteig =Cuphea cardonae 

Cuphea speciosa (A. Anderson) Kuntze =Cuphea melvilla 

Dodecas maritimus (Aubl.) Griseb. =Crenea maritima 

Dodecas surinamensis L. f. =Crenea maritima 

Grislea compacta Rusby =Pehria compacta 

Grislea secunda Loefl. =Pehria compacta 

Hypobrichia spruceana Benth. =Rotala mexicana 

Hypobrichia spruceana var. tenuifolia Griseb. =Rotala mexicana 

Lafoensia pacari A. St.-Hil. [not in flora area] 

Lawsonia alba Lam. =Lawsonia inermis 

Lawsonia speciosa L. =Lawsonia inermis 

Lythrum carthagenense Jacq. =Cuphea carthagenensis 

Melvilla speciosa A. Anderson [nomen nudum] =Cuphea melvilla 

Rotala apetala F. Muell. =Rotala mexicana 

Rotala mexicana var. apetala (F. Muell.) Lourteig =Rotala mexicana 

Rotala mexicana var. spruceana (Benth.) Koehne =Rotala mexicana 

REFERENCES 

Cook, C.D.K. 1979. A revision of the genus Rotala. Boissiera 29: 1-156. 

Graham, S.A. 1985. A revision of Ammannia in the western hemisphere. J. Arnold Arb. 56: 265-335. 

Jonker, F.P. 1941. Lythraceae. In A. Pulle, Flora of Suriname 3(1): 422-432. 

Kawasaki, M.L. 2002. Lythraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


408-409. 

Lourteig, A. 1964. Lythraceae Austroamericanae, addenda et corrigenda. Sellowia 16: 119-162. 

Lourteig, A. 1986. Revision del genero Lafoensia Vandelli (Litraceas). Mem. Soc. Cienc. Nat. La Salle 45(123): 115-157. 

Lourteig, A. 1986. Revision de dos secciones del genero Cuphea P. Browne. Phytologia 60(1): 17-55. 

Lourteig, A. 1987. Lythraceae Austroamericanae, addenda et corrigenda II. Sellowia 39: 15-48. 

Lourteig, A. 2001. Lythraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 6: 

59-80. 

MAGNOLIACEAE 


Dugandiodendron chimantense (Steyerm. & Maguire) Lozano 

Dugandiodendron ptaritepuianum (Steyerm.) Lozano [Mt. Roraima] 


Magnolia chimantensis Steyerm. & Maguire =Dugandiodendron chimantense 

Magnolia ptaritepuiana Steyerm. =Dugandiodendron ptaritepuianum 

Magnolia roraimae Steyerm. =Dugandiodendron ptaritepuianum 

AM 

BO GU 

BO 

GU 

BO DA GU SU 

BO 


BO DA 

BO 

BO GU

386 


REFERENCES 

Berry, P.E. and J.S. Miller. 2001. Magnoliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Lozano Contreras, G. 1994. Dugandiodendron y Talauma (Magnoliaceae) en el Neotrópica. Academia de Ciencias Exactas, 

Fisicas y Naturales Coleccion Jorge Alvarez Lleras No. 3. 

Steyermark, J.A. 1951. Magnoliaceae. In Botanical Exploration in Venezuela - I. Fieldiana Bot. 28(1): 233-234. 

Banisteriopsis caapi (Griseb.) C.V. Morton 

Banisteriopsis carolina W.R. Anderson 

Banisteriopsis cinerascens (Benth.) B. Gates 

Banisteriopsis cristata (Griseb.) Cuatrec. 

Banisteriopsis krukoffii B. Gates 

Banisteriopsis lucida (Rich.) Small 

Banisteriopsis lyrata B. Gates 

Banisteriopsis maguirei B. Gates 

Banisteriopsis martiniana (A. Juss.) Cuatrec. 

Banisteriopsis martiniana var. martiniana 

Banisteriopsis martiniana var. subenervia Cuatrec. 

Banisteriopsis mathiasiae (W.R. Anderson) W.R. Anderson 

Banisteriopsis muricata (Cav.) Cuatrec. 

Banisteriopsis nummifera (A. Juss.) B. Gates 

Banisteriopsis pulcherrima (Sandwith) B. Gates 

Banisteriopsis schwannioides (Griseb.) B. Gates 

Banisteriopsis wurdackii B. Gates 

Blepharandra angustifolia (Kunth) W.R. Anderson 

Blepharandra fimbriata MacBryde 

Blepharandra heteropetala W.R. Anderson 

Blepharandra hypoleuca (Benth.) Griseb. 

Bunchosia apiculata Huber 

Bunchosia argentea (Jacq.) DC. 

Bunchosia armeniaca (Cav.) DC. 

Bunchosia decussiflora W.R. Anderson 

Bunchosia glandulifera (Jacq.) Kunth 

Bunchosia mollis Benth. 

Bunchosia petraea W.R. Anderson 

Bunchosia polystachia (Andrews) DC. 

Burdachia prismatocarpa A. Juss. 

Burdachia sphaerocarpa A. Juss. 

Byrsonima aerugo Sagot 

Byrsonima altissima DC. 

Byrsonima arthropoda A. Juss. 

Byrsonima baccae W.R. Anderson 

Byrsonima basiliana W.R. Anderson 

Byrsonima bronweniana W.R. Anderson 

Byrsonima carraoana Steyerm. 

Byrsonima chalcophylla Nied. 

Byrsonima christianeae W.R. Anderson [expected in BO] 

Byrsonima chrysophylla Kunth 

Byrsonima coccolobifolia Kunth 

Byrsonima concinna Benth. 

Byrsonima coniophylla A. Juss. 

Byrsonima cowanii W.R. Anderson 

Byrsonima crassifolia (L.) Kunth 

Byrsonima crispa A. Juss. 

Byrsonima cuprea Griseb. 

Byrsonima densa (Poir.) DC. 

Byrsonima dubia W.R. Anderson 

Byrsonima duidana W.R. Anderson 

Byrsonima eugeniifolia Sandwith 

Byrsonima euryphylla Pilg. 

Byrsonima fanshawei W.R. Anderson 

MALPIGHIACEAE 

C. Anderson, W.R. Anderson 

AM 

FG 

BO 

GU 

GU SU 

AM BO 


BO DA 

AM BO 


AM 

BO DA GU 

GU 

SU FG 

BO GU FG 

FG 

BO GU 

FG 

AM 

BO FG 

AM 

BO GU 

AM BO GU 

FG 

BO GU SU? FG 

BO DA 

AM GU FG 

AM SU FG 

AM 

BO GU SU? 

FG 

AM BO 

AM GU 


GU FG 

AM 

AM 

AM 

GU SU 

BO GU 

BO GU 

GU 

AM BO SU 

BO DA GU SU 

AM 

AM 

AM 

BO GU 


AM 

AM 

BO FG 

GU SU FG 

AM 

BO GU 

GU 

GU 

SU 

SU


Byrsonima fernandezii Cuatrec. 

Byrsonima flexipes W.R. Anderson 

Byrsonima formosa W.R. Anderson 

Byrsonima frondosa A. Juss. 

Byrsonima gymnocalycina A. Juss. 

Byrsonima huberi W.R. Anderson 

Byrsonima incarnata Sandwith 

Byrsonima japurensis A. Juss. 

Byrsonima kariniana W.R. Anderson 

Byrsonima krukoffii W.R. Anderson 

Byrsonima laevigata (Poir.) DC. 

Byrsonima laevis Nied. 

Byrsonima leucophlebia Griseb. 

Byrsonima linguifera Cuatrec. 

Byrsonima luetzelburgii Steyerm. 

Byrsonima macrostachya W.R. Anderson 

Byrsonima maguirei W.R. Anderson [Sierra de la Neblina] 

Byrsonima nitidissima Kunth 

Byrsonima pachypoda W.R. Anderson 

Byrsonima poeppigiana A. Juss. 

Byrsonima punctulata A. Juss. 

Byrsonima rubrobracteata W.R. Anderson [expected in BO] 

Byrsonima schomburgkiana Benth. 

Byrsonima sericea DC. 

Byrsonima spicata (Cav.) DC. 

Byrsonima steyermarkii W.R. Anderson 

Byrsonima stipulacea A. Juss. 

Byrsonima surinamensis W.R. Anderson 

Byrsonima tillettii W.R. Anderson 

Byrsonima verbascifolia (L.) DC. 

Byrsonima wurdackii W.R. Anderson 

Camarea affinis A. St.-Hil. 

Clonodia complicata (Kunth) W.R. Anderson 

Coleostachys genipifolia A. Juss. 

Diacidia cordata (Maguire) W.R. Anderson [Sierra de la Neblina] 

Diacidia ferruginea (Maguire & K.D. Phelps) W.R. Anderson 

Diacidia galphimioides Griseb. 

Diacidia glaucifolia (Maguire) W.R. Anderson 

Diacidia hypoleuca (Maguire) W.R. Anderson 

Diacidia kunhardtii (Maguire) W.R. Anderson 

Diacidia rufa (Maguire) W.R. Anderson [Sierra de la Neblina] 

Diacidia steyermarkii (Maguire) W.R. Anderson 

Diacidia stipularis (Maguire & K.D. Phelps) W.R. Anderson 

Diacidia vestita (Benth.) B.D. Jacks. 

Dicella julianii (J.F. Macbr.) W.R. Anderson [expected in AM; Serra da 

Neblina] 

Diplopterys cabrerana (Cuatrec.) B. Gates 

Diplopterys pauciflora (G. Mey.) Nied. 

Excentradenia adenophora (Sandwith) W.R. Anderson 

Excentradenia propinqua (W.R. Anderson) W.R. Anderson 

Glandonia williamsii Steyerm. 

Heteropterys alata (W.R. Anderson) W.R. Anderson 

Heteropterys atabapensis W.R. Anderson 

Heteropterys ayacuchensis W.R. Anderson 

Heteropterys biglandulosa A. Juss. 

Heteropterys cristata Benth. 

Heteropterys cuatrecasasii W.R. Anderson 

Heteropterys dichromocalyx W.R. Anderson [expected in AM; Pico da 

Neblina] 

Heteropterys guianensis W.R. Anderson 

Heteropterys hoffmanii W.R. Anderson 

Heteropterys huberi W.R. Anderson 

Heteropterys leona (Cav.) Exell 

Heteropterys macradena (DC.) W.R. Anderson 

AM 

AM 

AM 

AM 

AM BO 

AM 

AM 

GU 

GU SU? FG? 

GU 

GU 

GU SU FG 

BO 

AM 

AM 

AM 

AM 

AM BO 

BO GU 

AM 

AM 

AM BO 

GU 

GU 


FG 

FG 

AM BO 

BO DA GU SU 

SU 

FG 

BO GU 

AM 

AM 

BO GU SU FG 

GU SU 

AM 

AM 

AM 

BO DA 

FG 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

AM 

GU FG 

BO DA GU 

GU SU FG 

AM 

AM 

AM 

AM 

BO 

AM BO GU 

AM 

AM 

GU 

GU 

FG 

DA GU SU FG 

AM BO GU SU FG

388 


Heteropterys macrostachya A. Juss. 

Heteropterys maguirei W.R. Anderson 

Heteropterys mathewsana A. Juss. 

Heteropterys megaptera A. Juss. 

Heteropterys molesta W.R. Anderson 

Heteropterys neblinensis W.R. Anderson [Sierra de la Neblina] 

Heteropterys nervosa A. Juss. 

Heteropterys oblongifolia Gleason 

Heteropterys oligantha W.R. Anderson 

Heteropterys orinocensis (Kunth) A. Juss. 

Heteropterys quetepensis Steyerm. 

Heteropterys siderosa Cuatrec. 

Heteropterys steyermarkii W.R. Anderson 

Heteropterys subhelicina Nied. 

Heteropterys velutina W.R. Anderson 

Hiraea affinis Miq. 

Hiraea apaporiensis Cuatrec. 

Hiraea celiana W.R. Anderson 

Hiraea fagifolia (DC.) A. Juss. 

Hiraea faginea (Sw.) Nied. 

Hiraea fimbriata W.R. Anderson 

Hiraea gracieana W.R. Anderson 

Hiraea longipes W.R. Anderson 

Hiraea longipilifera W.R. Anderson 

Hiraea morii W.R. Anderson 

Hiraea neblinensis W.R. Anderson 

Hiraea quapara (Aubl.) Sprague 

Hiraea steyermarkii W.R. Anderson 

Hiraea tepuiensis Steyerm. 

Hiraea ternifolia (Kunth) A. Juss. 

Jubelina grisebachiana W.R. Anderson 

Jubelina magnifica W.R. Anderson 

Jubelina riparia A. Juss. 

Jubelina rosea (Miq.) Nied. 

Lophanthera longifolia (Kunth) Griseb. 

Lophopterys euryptera Sandwith 

Lophopterys inpana W.R. Anderson 

Lophopterys splendens A. Juss. 

Lophopterys surinamensis (Kosterm.) Sandwith 

Malpighia emarginata DC. 

Malpighia glabra L. 

Mascagnia arenicola C. Anderson 

Mascagnia castanea (Cuatrec.) W.R. Anderson 

Mascagnia cynanchifolia Griseb. 

Mascagnia dissimilis C.V. Morton & Moldenke 

Mascagnia divaricata (Kunth) Nied. 

Mascagnia eggersiana (Nied.) W.R. Anderson 

Mascagnia glandulifera Cuatrec. 

Mascagnia guianensis W.R. Anderson 

Mascagnia liesneri W.R. Anderson 

Mascagnia macrodisca (Triana & Planch.) Nied. 

Mascagnia microcarpa (Sandwith) W.R. Anderson 

Mascagnia ovatifolia (Kunth) Griseb. 

Mascagnia poeppigiana (A. Juss.) W.R. Anderson 

Mascagnia schunkei W.R. Anderson 

Mascagnia sepium (A. Juss.) Griseb. 

Mascagnia sinemariensis (Aubl.) Griseb. 

Mascagnia stannea (Griseb.) Nied. 

Mascagnia surinamensis (Kosterm.) W.R. Anderson 

Mezia angelica W.R. Anderson [expected in SU] 

Mezia curranii W.R. Anderson 

Mezia huberi W.R. Anderson 

Mezia includens (Benth.) Cuatrec. 

Mezia rufa W.R. Anderson 


BO GU 

SU 

BO GU 

BO 

AM 

AM GU SU FG 

AM 

GU 

AM BO 

AM BO 

AM GU FG 

AM 

GU 

GU 

DA GU SU FG 

AM 

AM 

BO GU SU FG 


AM BO 

GU SU FG 

FG 

GU 

FG 

AM 

FG 

BO 

AM BO 

AM BO 

AM 

AM 

FG 

SU FG 

AM 

BO DA GU 

AM 

FG 

SU 

BO DA FG 

BO 

GU SU FG 

AM 

AM 

AM 

DA SU FG 

AM BO 

AM 

GU SU FG 

AM 

AM BO GU SU 

GU 

BO 

AM GU 

AM FG 


AM BO GU FG 

AM 

AM GU SU 

GU FG 

AM 

AM 

AM BO GU FG 

AM

Pterandra flavescens Maguire 

Pterandra guianensis W.R. Anderson [expected in BO] 

Pterandra sericea W.R. Anderson 

Spachea elegans (G. Mey.) A. Juss. 

Stigmaphyllon adenodon A. Juss. 

Stigmaphyllon adenodon var. adenodon 

Stigmaphyllon bannisterioides (L.) C.E. Anderson 

Stigmaphyllon ciliatum (Lam.) A. Juss. 

Stigmaphyllon convolvulifolium A. Juss. [expected in DA] 

Stigmaphyllon palmatum (Cav.) A. Juss. 

Stigmaphyllon puberum (Rich.) A. Juss. 

Stigmaphyllon sinuatum (DC.) A. Juss. 

Tetrapterys acutifolia Cav. 

Tetrapterys anomala W.R. Anderson 

Tetrapterys aristeguietae W.R. Anderson 

Tetrapterys calophylla A. Juss. 

Tetrapterys crispa A. Juss. 

Tetrapterys discolor (G. Mey.) DC. 

Tetrapterys fimbripetala A. Juss. 

Tetrapterys glabrifolia (Griseb.) Small 

Tetrapterys gracilis W.R. Anderson 

Tetrapterys maranhamensis A. Juss. 

Tetrapterys megalantha W.R. Anderson 

Tetrapterys mucronata Cav. 

Tetrapterys oleifolia (Benth.) Griseb. 

Tetrapterys pusilla Steyerm. 

Tetrapterys rhodopteron Oliv. 

Tetrapterys styloptera A. Juss. 


AM 

BO 

GU 

GU 

BO DA 

DA 

DA 

GU SU FG 

DA GU 

GU? 

SU FG 

GU SU FG 

FG 

DA GU SU FG 


BO 

GU 

GU 

SU FG 

GU SU FG 




AM 

SU FG 

BO GU 

GU 

SU? 


BO GU 

BO GU 

BO GU 



Alcoceratothrix rugosa (Benth.) Nied. =Byrsonima stipulacea 

Alcoceratothrix stipulacea (A. Juss.) Nied. =Byrsonima stipulacea 

Banisteria argentea (Kunth) Spreng. =Banisteriopsis muricata 

Banisteria caapi Griseb. =Banisteriopsis caapi 

Banisteria cristata Griseb. =Banisteriopsis cristata 

Banisteria fagifolia DC. =Hiraea fagifolia 

Banisteria heterophylla Willd. [non Stigmaphyllon heterophyllum Hook. 1843] =Stigmaphyllon sinuatum 

Banisteria leona Cav. =Heteropterys leona 

Banisteria leptocarpa Benth. =Banisteriopsis martiniana 

Banisteria lucida Rich. =Banisteriopsis lucida 

Banisteria macradena DC. =Heteropterys macradena 

Banisteria martiniana A. Juss. =Banisteriopsis martiniana 

Banisteria metallicolor A. Juss. =Banisteriopsis muricata 

Banisteria mossii C.V. Morton =Heteropterys nervosa 

Banisteria multiflora DC. =Heteropterys leona 

Banisteria muricata Cav. =Banisteriopsis muricata 

Banisteria orbicularis Nied. =Banisteriopsis cristata 

Banisteria orinocensis Kunth =Heteropterys orinocensis 

Banisteria ovata Cav. =Stigmaphyllon bannisterioides 

Banisteria pubera Rich. =Stigmaphyllon puberum 

Banisteria quapara Aubl. =Hiraea quapara 

Banisteria reticulata (Poir.) C.B. Rob. =Heteropterys leona 

Banisteria schomburgkiana Benth. =Banisteriopsis muricata 

Banisteria sinemariensis Aubl. =Mascagnia sinemariensis 

Banisteria sinuata DC. =Stigmaphyllon sinuatum 

Banisteria splendens DC. [nom. superfl.] =Stigmaphyllon sinuatum 

Banisteriopsis argentea (Kunth) C.B. Rob. =Banisteriopsis muricata 

Banisteriopsis cabrerana Cuatrec. =Diplopterys cabrerana 

Banisteriopsis elegans subsp. cordata (Nied.) Sandwith [not in flora area] 

Banisteriopsis elegans var. pulcherrima Sandwith =Banisteriopsis pulcherrima 

Banisteriopsis gardneriana (A. Juss.) W.R. Anderson & B. Gates [reported from FG in error, fide W. Anderson] 

Banisteriopsis inebrians C.V. Morton =Banisteriopsis caapi 

Banisteriopsis leptocarpa (Benth.) R.O. Williams =Banisteriopsis martiniana 

Banisteriopsis martiniana var. laevis Cuatrec. =Banisteriopsis martiniana var. subenervia

390 


Banisteriopsis metallicolor (A. Juss.) O’Donell & Lourteig =Banisteriopsis muricata 

Banisteriopsis schomburgkiana (Benth.) C.B. Rob. =Banisteriopsis muricata 

Blepharandra cretacea (Gleason) Steyerm. =Blepharandra hypoleuca 

Blepharandra ptariana Steyerm. =Blepharandra hypoleuca 

Brachypterys borealis A. Juss. [nom. superfl.] =Stigmaphyllon bannisterioides 

Brachypterys ovata (Cav.) Small =Stigmaphyllon bannisterioides 

Bunchosia maritima (Vell.) J.F. Macbr. [not in flora area] 

Bunchosia nitida (Jacq.) Rich. [not in flora area] 

Bunchosia rhombifolia Turcz. =Bunchosia mollis 

Bunchosia schomburgkiana Nied. =Bunchosia mollis 

Bunchosia squarrosa Griseb. =Tetrapterys styloptera 

Bunchosia tuberculata DC. [dubious name of uncertain status, fide W. Anderson] 

Burdachia atractoides Nied. =Burdachia sphaerocarpa 

Burdachia prismatocarpa var. sphaerocarpa (A. Juss.) Nied. =Burdachia sphaerocarpa 

Burdachia sphaerocarpa var. glandifera (Gleason) W.R. Anderson =Burdachia sphaerocarpa 

Burdachia williamsii Steyerm. =Burdachia prismatocarpa 

Byrsonima angustifolia Kunth =Blepharandra angustifolia 

Byrsonima aubletii Kosterm. =Byrsonima altissima 

Byrsonima barkleyana Cuatrec. =Byrsonima punctulata 

Byrsonima bolivarana Steyerm. =Byrsonima carraoana 

Byrsonima bracteolaris Benth. =Byrsonima concinna 

Byrsonima carmeniana Cuatrec. =Byrsonima crispa 

Byrsonima chalcophylla var. carraoana (Steyerm.) W.R. Anderson =Byrsonima carraoana 

Byrsonima coriacea (Sw.) DC. [not in flora area] 

Byrsonima coriacea var. spicata (Cav.) Nied. [nom. superfl.] =Byrsonima spicata 

Byrsonima cretacea Nied. =Byrsonima luetzelburgii 

Byrsonima cretacea Gleason =Blepharandra hypoleuca 

Byrsonima cydoniifolia A. Juss. [not in flora area] 

Byrsonima discolor Pilg. =Byrsonima altissima 

Byrsonima ferruginea Kunth =Byrsonima crassifolia 

Byrsonima ferruginea var. macrophylla Benth. =Byrsonima aerugo 

Byrsonima fluminensis Nied. =Byrsonima japurensis 

Byrsonima inundata Benth. =Byrsonima japurensis 

Byrsonima japurensis subsp. fluminensis (Nied.) Cuatrec. =Byrsonima japurensis 

Byrsonima japurensis subsp. silvatica Cuatrec. =Byrsonima japurensis 

Byrsonima laurifolia Kunth =Byrsonima crassifolia 

Byrsonima obversa Miq. =Byrsonima laevigata 

Byrsonima propinqua Benth. =Byrsonima spicata 

Byrsonima reticulata (Poir.) DC. =Heteropterys leona 

Byrsonima rugosa Benth. =Byrsonima stipulacea 

Byrsonima sessilifolia Benth. =Byrsonima coccolobifolia 

Byrsonima uvulifera Spruce ex Nied. =Byrsonima japurensis 

Byrsonima verbascifolia var. denudata Cuatrec. =Byrsonima verbascifolia 

Clonodia racemosa var. orinocensis Nied. =Clonodia complicata 

Coleostachys hypoleuca Benth. =Blepharandra hypoleuca 

Coleostachys vestita Benth. =Diacidia vestita 

Diacidia duckeana Maguire =Diacidia galphimioides 

Diacidia parvifolia Cuatrec. =Diacidia galphimioides 

Dicella amazonica Pires =Dicella julianii 

Diplopterys includens (Benth.) Nied. =Mezia includens 

Diplopterys involuta (Turcz.) Nied. =Mezia includens 

Diplopterys involuta var. ovata Nied. =Mezia curranii 

Diplopterys microcarpa Sandwith =Mascagnia microcarpa 

Diplopterys riparia (A. Juss.) Nied. =Jubelina riparia 

Diplopterys rosea (Miq.) Nied. =Jubelina rosea 

Diplopterys spruceana (Nied.) Nied. [not in flora area] =Mascagnia bracteosa Griseb. 

Dolichopterys surinamensis Kosterm. =Lophopterys surinamensis 

Galphimia longifolia Kunth =Lophanthera longifolia 

Heteropterys acutifolia A. Juss. =Heteropterys orinocensis 

Heteropterys africana A. Juss. [nom. superfl.] =Heteropterys leona 

Heteropterys argentea Kunth =Banisteriopsis muricata 

Heteropterys beecheyana var. alata W.R. Anderson =Heteropterys alata 

Heteropterys carinata Benth. =Heteropterys cristata 

Heteropterys castanea Cuatrec. =Mascagnia castanea


Heteropterys catoptera W.R. Anderson =Heteropterys subhelicina 

Heteropterys helicina Griseb. =Heteropterys orinocensis 

Heteropterys lasseri W.R. Anderson =Heteropterys megaptera 

Heteropterys mossii (C.V. Morton) Cuatrec. =Heteropterys nervosa 

Heteropterys multiflora (DC.) Hochr. =Heteropterys leona 

Heteropterys oleifolia (Benth.) Griseb. =Tetrapterys oleifolia 

Heteropterys reticulata (Poir.) Nied. =Heteropterys leona 

Heteropterys stannea Griseb. =Mascagnia stannea 

Heteropterys suberosa Griseb. =Heteropterys nervosa 

Hiraea adenophora Sandwith =Excentradenia adenophora 

Hiraea bifurcata W.R. Anderson =Hiraea ternifolia 

Hiraea chrysophylla A. Juss. =Hiraea faginea 

Hiraea complicata Kunth =Clonodia complicata 

Hiraea demerarensis (A. Juss.) Nied. =Hiraea faginea 

Hiraea divaricaa Kunth =Mascagnia divaricata 

Hiraea fulgens A. Juss. =Hiraea faginea 

Hiraea fulgens var. demerarensis A. Juss. =Hiraea faginea 

Hiraea gaudichaudiana A. Juss. [not in flora area] 

Hiraea gracilis Benth. =Tetrapterys maranhamensis 

Hiraea laurifolia A. Juss. =Mascagnia stannea 

Hiraea macrodisca Triana & Planch. =Mascagnia macrodisca 

Hiraea multiradiata A. Juss. =Hiraea quapara 

Hiraea nitida Kunth =Clonodia complicata 

Hiraea oblongifolia DC. =Mascagnia divaricata 

Hiraea oleifolia Benth. =Tetrapterys oleifolia 

Hiraea ovatifolia Kunth =Mascagnia ovatifolia 

Hiraea platytriphylla Hochr. =Hiraea ternifolia 

Hiraea poeppigiana A. Juss. =Mascagnia poeppigiana 

Hiraea propinqua W.R. Anderson =Excentradenia propinqua 

Hiraea schizoptera Turcz. =Mascagnia sinemariensis 

Hiraea sepium A. Juss. =Mascagnia sepium 

Hiraea swartziana A. Juss. [nom. superfl.] =Hiraea faginea 

Jubelina bracteosa auct. non Mascagnia bracteosa Griseb. =Jubelina grisebachiana 

Lophopterys splendens var. obovata Nied. =Lophopterys euryptera 

Malpighia altissima Aubl. =Byrsonima altissima 

Malpighia argentea Jacq. =Bunchosia argentea 

Malpighia armeniaca Cav. =Bunchosia armeniaca 

Malpighia bannisterioides L. =Stigmaphyllon bannisterioides 

Malpighia crassifolia L. =Byrsonima crassifolia 

Malpighia elegans G. Mey. =Spachea elegans 

Malpighia faginea Sw. =Hiraea faginea 

Malpighia glandulifera Jacq. =Bunchosia glandulifera 

Malpighia moureila Aubl. =Byrsonima crassifolia 

Malpighia punicifolia L. =Malpighia glabra 

Malpighia punicifolia auct. non L. 1762 =Malpighia emarginata 

Malpighia reticulata Poir. =Heteropterys leona 

Malpighia spicata Cav. =Byrsonima spicata 

Malpighia ternifolia Kunth =Hiraea ternifolia 

Malpighia verbascifolia L. =Byrsonima verbascifolia 

Malpighia volubilis Sims =Mascagnia sinemariensis 

Mascagnia anisopetala (A. Juss.) Griseb. [not in flora area] 

Mascagnia anisopetala var. macrodisca (Triana & Planch.) Kosterm. =Mascagnia macrodisca 

Mascagnia benthamiana (Griseb.) W.R. Anderson =Mascagnia stannea 

Mascagnia complicata (Kunth) Nied. =Clonodia complicata 

Mascagnia cordifolia var. peruviana J.F. Macbr. =Mascagnia schunkei 

Mascagnia hondensis C.V. Morton =Mascagnia sinemariensis 

Mascagnia lehmanniana Nied. =Clonodia complicata 

Mascagnia leucanthele Griseb. [not in flora area] 

Mascagnia materdei Cuatrec. =Mascagnia poeppigiana 

Mascagnia multiglandulosa Nied. [not in flora area] 

Mascagnia multiglandulosa var. surinamensis Kosterm. =Mascagnia surinamensis 

Mascagnia nervosa Nied. =Mascagnia ovatifolia 

Mascagnia nitida (Kunth) Nied. =Clonodia complicata 

Mascagnia pauciflora (G. Mey.) Cuatrec. =Diplopterys pauciflora

392 


Mascagnia schizoptera (Turcz.) Cuatrec. =Mascagnia sinemariensis 

Mascagnia sericans Nied. =Mascagnia stannea 

Mascagnia subsericea Cuatrec. =Mascagnia poeppigiana 

Mascagnia tenuifolia var. eggersiana Nied. =Mascagnia eggersiana 

Mascagnia volubilis (Sims) Nied. =Mascagnia sinemariensis 

Sipapoa cordata Maguire =Diacidia cordata 

Sipapoa ferruginea Maguire & K.D. Phelps =Diacidia ferruginea 

Sipapoa glaucifolia Maguire =Diacidia glaucifolia 

Sipapoa hypoleuca Maguire =Diacidia hypoleuca 

Sipapoa kunhardtii Maguire =Diacidia kunhardtii 

Sipapoa rufa Maguire =Diacidia rufa 

Sipapoa steyermarkii Maguire =Diacidia steyermarkii 

Sipapoa stipularis Maguire & K.D. Phelps =Diacidia stipularis 

Sipapoa vestita (Benth.) Maguire =Diacidia vestita 

Skoliopterys lehmanniana (Nied.) Cuatrec. =Clonodia complicata 

Stenocalyx involutus Turcz. =Mezia includens 

Stigmaphyllon angustilobum A. Juss. [not in flora area] 

Stigmaphyllon brachiatum Triana & Planch. =Stigmaphyllon sinuatum 

Stigmaphyllon fulgens A. Juss. [nom. superfl.] =Stigmaphyllon sinuatum 

Stigmaphyllon grenadense Nied. =Stigmaphyllon adenodon 

Stigmaphyllon hypoleucum Miq. =Stigmaphyllon sinuatum 

Stigmaphyllon latifolium Benth. =Stigmaphyllon convolvulifolium 

Stigmaphyllon martianum A. Juss. =Stigmaphyllon sinuatum 

Stigmaphyllon mathiasiae W.R. Anderson =Banisteriopsis mathiasiae 

Stigmaphyllon monancistrum Nied. =Stigmaphyllon sinuatum 

Stigmaphyllon ovatum (Cav.) Nied. =Stigmaphyllon bannisterioides 

Stigmaphyllon periplocifolium A. Juss. [not in flora area] 

Stigmaphyllon purpureum Benth. =Stigmaphyllon sinuatum 

Stigmaphyllon sagittatum A. Juss. =Stigmaphyllon palmatum 

Stigmaphyllon splendens Cuatrec. [nom. superfl.] =Stigmaphyllon sinuatum 

Stigmaphyllon tiliifolium Nied. [not in flora area] 

Tetrapodenia glandifera Gleason =Burdachia sphaerocarpa 

Tetrapterys benthamiana Griseb. =Mascagnia stannea 

Tetrapterys boliviensis Nied. =Tetrapterys styloptera 

Tetrapterys crebriflora A. Juss. =Tetrapterys mucronata 

Tetrapterys glaberrima Benth. =Tetrapterys mucronata 

Tetrapterys huachamacariensis W.R. Anderson =Tetrapterys fimbripetala 

Tetrapterys includens Benth. =Mezia includens 

Tetrapterys julianii J.F. Macbr. =Dicella julianii 

Tetrapterys mucronata var. crebriflora (A. Juss.) J.F. Macbr. =Tetrapterys mucronata 

Tetrapterys mucronata subsp. crebriflora (A. Juss.) Kosterm. =Tetrapterys mucronata 

Tetrapterys phylladenophora Maguire & Steyerm. =Tetrapterys rhodopteron 

Tetrapterys poeppigiana (A. Juss.) Griseb. =Mascagnia poeppigiana 

Tetrapterys silvatica Cuatrec. =Tetrapterys mucronata 

Tetrapterys squarrosa (Griseb.) Griseb. =Tetrapterys styloptera 

Triopterys discolor G. Mey. =Tetrapterys discolor 

REFERENCES 

Anderson, C. 1993. The identities of the sericeous-leaved species of Stigmaphyllon in the Amazon region. Contr. Univ. Mich. 

Herb. 19: 393-413. 

Anderson, C. 1997. Monograph of Stigmaphyllon (Malpighiaceae). Systematic Botany Monographs 51: 1-313. 

Anderson, W.R. 1981. Malpighiaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 11. Mem. N.Y. Bot. 

Gard. 32: 1-391. 

Anderson, W.R. 1990. The taxonomy of Jubelina. Contr. Univ. Mich. Herb. 17: 21-37. 

Anderson, W.R. 1993. Notes on Neotropical Malpighiaceae - IV. Contr. Univ. Mich. Herb. 19: 355-392. 

Anderson, W.R. 1994. New species of Hiraea from the Guianas and adjacent Brazil. Brittonia 46(2): 126-133. 

Anderson, W.R. 1993. Notes on Neotropical Malpighiaceae - VI. Contr. Univ. Mich. Herb. 21: 37-84. 

Anderson, W.R. 2001. Malpighiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 82-185. 

Anderson, W.R. 2001. Malpighiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 410-428. 

Gates, B. 1982. Banisteriopsis, Diplopterys. Flora Neotropica Monogr. 30: 1-237. 

Görts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 1976. Malpighiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, 

Flora of Suriname 2(2): 445-450.


Jonker, F.P. 1939. Malpighiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 478-480. 

Kostermans, A.J.G.H. 1936. Malpighiaceae. In A. Pulle, Flora of Suriname 2(1): 146-243. 

Niedenzu, F. 1928. Malpighiaceae. In A. Engler, Das Pflanzenreich IV. 141: 1-870. 

MALVACEAE 

L.J. Dorr, P.A. Fryxell, M.J. Jansen-Jacobs 

Abelmoschus moschatus Medik. [cultivated] 

Abelmoschus spp. [cultivated] 

Abutilon indicum (L.) Sweet [cultivated] 

Allosidastrum pyramidatum (Desp. ex Cav.) Krapov., Fryxell & D.M. Bates 

Bastardia viscosa (L.) Kunth [introduced] 

Briquetia spicata (Kunth) Fryxell 

Cienfuegosia affinis (Kunth) Hochr. 

Cienfuegosia heterophylla (Vent.) Garcke 

Gaya gaudichaudiana A. St.-Hil. 

Gaya occidentalis (L.) Sweet 

Gossypium barbadense L. [cultivated only?] 

Gossypium hirsutum L. [cultivated only?] 

Herissantia crispa (L.) Brizicky 

Hibiscus bifurcatus Cav. 

Hibiscus dimidiatus Schrank 

Hibiscus furcellatus Desr. 

Hibiscus guianensis Aubl. [name of uncertain application] 

Hibiscus mutabilis L. [cultivated?] 

Hibiscus peruvianus R.E. Fr. 

Hibiscus phoeniceus Jacq. [cultivated?] 

Hibiscus prunifolius F. Dietr. 

Hibiscus radiatus Cav. [cultivated] 

Hibiscus rosa-sinensis L. [cultivated] 

Hibiscus sabdariffa L. [cultivated] 

Hibiscus schizopetalus (Dyer) Hook. f. [cultivated] 

Hibiscus sororius L. 

Hibiscus striatus Cav. 

Hibiscus striatus subsp. lambertianus (Kunth) O.J. Blanch. ex Proctor 

Hibiscus trilobus Aubl. 

Hibiscus trilobus var. ingratus (Miq.) O.J. Blanch. & Fryxell 

Hibiscus varians Splitg. 

Hibiscus verbasciformis Klotzsch ex Hochr. 

Malachra alceifolia Jacq. 

Malachra fasciata Jacq. 

Malachra palmata Moench 

Malachra radiata (L.) L. 

Malvastrum coromandelianum (L.) Garcke 

Pavonia angustifolia Benth. 

Pavonia cancellata (L.) Cav. 

Pavonia castaneifolia A. St.-Hil. & Naudin 

Pavonia dasypetala Turcz. 

Pavonia flavispina Miq. 

Pavonia fruticosa (Mill.) Fawc. & Rendle 

Pavonia geminiflora Moric. 

Pavonia imatacensis Steyerm. 

Pavonia malacophylla (Link & Otto) Garcke 

Pavonia paludicola Nicolson ex Fryxell 

Pavonia paniculata Cav. 

Pavonia piptocalyx Krapov. 

Pavonia schiedeana Steud. 

Pavonia sidifolia Kunth 

Peltaea riedelii (Gürke) Standl. 

Peltaea sessiliflora (Kunth) Standl. 

Peltaea speciosa (Kunth) Standl. 

Peltaea surumuensis (Ulbr.) Krapov. & Cristóbal [possibly expected in the 


Peltaea trinervis (C. Presl) Krapov. & Cristóbal 

BO DA 

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GU SU 

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BO GU 

BO 

BO 

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GU SU FG 


BO GU 


AM GU 


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GU? 

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BO 

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394 


Pseudabutilon umbellatum (L.) Fryxell 

Sida acuta Burm. f. 

Sida aggregata C. Presl 

Sida angustissima A. St.-Hil. 

Sida ciliaris L. 

Sida cordata (Burm. f.) Borss. Waalk. 

Sida cordifolia L. 

Sida glomerata Cav. 

Sida jamaicensis L. 

Sida linifolia Juss. ex Cav. 

Sida poeppigiana (K. Schum.) Fryxell 

Sida rhombifolia L. 

Sida santaremensis Monteiro 

Sida serrata Willd. ex Spreng. 

Sida setosa Mart. ex Colla 

Sida spinosa L. 

Sida surumuensis Ulbr. 

Sida urens L. 

Sida viarum A. St.-Hil. 

Sidastrum micranthum (A. St.-Hil.) Fryxell 

Sidastrum quinquenervium (Duchass. ex Triana & Planch.) Baker f. 

Talipariti tiliaceum (L.) Fryxell 

Talipariti tiliaceum var. pernambucense (Arruda) Fryxell 

Thespesia lampas (Cav.) Dalzell & Gibson [cultivated] 

Thespesia populnea (L.) Sol. ex CorrLa [cultivated or escaped from 

cultivation] 

Urena lobata L. 

Urena sinuata L. 

Wissadula contracta (Link) R.E. Fr. 

Wissadula excelsior (Cav.) C. Presl 

Wissadula hernandioides (L’Hér.) Garcke 

Wissadula hirsuta C. Presl 

Wissadula periplocifolia (L.) C. Presl ex Thwaites 


Abutilon crispum (L.) Medik. =Herissantia crispa 

Abutilon patens A. St.-Hil. =Wissadula excelsior 

Abutilon spicatum Kunth =Briquetia spicata 

Althaea racemosa Sw. =Pavonia paludicola 

Bogenhardia crispa (L.) Kearney =Herissantia crispa 

Cienfuegosia phlomidifolia (A. St.-Hil.) Garcke =Cienfuegosia affinis 

Fugosia campestris Benth. =Cienfuegosia affinis 

Fugosia guianensis Klotzsch ex M.R. Schomb. =Cienfuegosia affinis 

Fugosia phlomidifolia A. St.-Hil. =Cienfuegosia affinis 

Fugosia retusa Turcz. =Cienfuegosia affinis 

Gayoides crispum (L.) Small =Herissantia crispa 

Gossypium latifolium Murr. =Gossypium hirsutum 

Gossypium peruvianum Cav. =Gossypium barbadense 

Gossypium punctatum Schumach. & Thonn. =Gossypium hirsutum 

Gossypium religiosum L. =Gossypium hirsutum 

Gossypium vitifolium Lam. =Gossypium barbadense 

Hibiscus abelmoschus L. =Abelmoschus moschatus 

Hibiscus affinis Kunth =Cienfuegosia affinis 

Hibiscus bracteosus DC. =Talipariti tiliaceum var. pernambucense 

Hibiscus brasiliensis L. [not in flora area] 

Hibiscus cancellatus L. =Pavonia cancellata 

Hibiscus cubensis A. Rich. =Hibiscus striatus subsp. lambertianus 

Hibiscus elatus Sw. =Talipariti tiliaceum 

Hibiscus esculentus L. [cultivated] =Abelmoschus esculentus (L.) Moench 

Hibiscus fraternus L. =Hibiscus sabdariffa 

Hibiscus ingratus Miq. =Hibiscus trilobus var. ingratus 

Hibiscus lambertianus Kunth =Hibiscus striatus subsp. lambertianus 

Hibiscus lampas Cav. =Thespesia lampas 

Hibiscus moschatus (Medik.) Salisb. =Abelmoschus moschatus 

Hibiscus pernambucensis Arruda =Talipariti tiliaceum var. pernambucense 

BO 


BO GU 

BO 

BO DA GU 

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BO 


GU 


GU 



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GU 

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DA GU SU FG 

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BO GU 

BO GU SU 

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Hibiscus salviifolius A. St.-Hil. =Hibiscus striatus subsp. lambertianus 

Hibiscus spathulatus Garcke =Hibiscus verbasciformis 

Hibiscus tiliaceus L. =Talipariti tiliaceum 

Lopimia dasypetala (Turcz.) Standl. =Pavonia dasypetala 

Lopimia malacophylla (Link & Otto) Mart. =Pavonia malacophylla 

Malache chiapensis Standl. =Pavonia sidifolia 

Malache scabra B. Vogel =Pavonia paludicola 

Malachra alceifolia var. fasciata (Jacq.) A. Robyns =Malachra fasciata 

Malachra alceifolia var. rotundifolia (Schrank) Guerke =Malachra alceifolia 

Malachra capitata L. =Malachra alceifolia 

Malachra heptaphylla Fischer =Malachra palmata 

Malachra kegeliana Garcke =Malachra fasciata 

Malachra trinervis C. Presl =Peltaea trinervis 

Pavonia chiapensis (Standl.) Standl. =Pavonia sidifolia 

Pavonia guanacastensis Standl. =Pavonia cancellata 

Pavonia julianae Uittien =Peltaea riedelii 

Pavonia racemosa (Sw.) Sw. =Pavonia paludicola 

Pavonia rosea Schltdl. =Pavonia schiedeana 

Pavonia scabra (B. Vogel) Cif. =Pavonia paludicola 

Pavonia sessiliflora Kunth =Peltaea sessiliflora 

Pavonia speciosa Kunth =Peltaea speciosa 

Pavonia spicata Cav. [nom. illeg.] =Pavonia paludicola 

Pavonia spinifex Cav. [not in flora area] 

Pavonia surumuensis Ulbr. =Peltaea surumuensis 

Pavonia trinervis (C. Presl) A. Robyns =Peltaea trinervis 

Pavonia typhalaea Cav. =Pavonia fruticosa 

Pavonia ulbrichiana Kearney =Hibiscus dimidiatus 

Peltaea aliculata Krapov. & Cristóbal =Peltaea surumuensis 

Pseudabutilon spicatum (Kunth) R.E. Fr. =Briquetia spicata 

Redutea heterophylla Vent. =Cienfuegosia heterophylla 

Sida carpinifolia L. f. =Sida acuta 

Sida contracta Link =Wissadula contracta 

Sida crispa L. =Herissantia crispa 

Sida excelsior Cav. =Wissadula excelsior 

Sida fruticosa Mill. =Pavonia fruticosa 

Sida guianensis K. Schum. =Sidastrum quinquenervium 

Sida hernandioides L’Hér. =Wissadula hernandioides 

Sida malacophylla Link & Otto =Pavonia malacophylla 

Sida micrantha A. St.-Hil. =Sidastrum micranthum 

Sida periplocifolia L. =Wissadula periplocifolia 

Sida quinquenervia Duchass. ex Triana & Planch. =Sidastrum quinquenervium 

Sida radiata L. =Malachra radiata 

Sida savannarum K. Schum. =Sida aggregata 

Sida setifera C. Presl =Sida aggregata 

Sida surinamensis Miq. =Sida setosa 

Sida umbellata L. =Pseudabutilon umbellatum 

Sida veronicifolia Lam. =Sida cordata 

Urena lobata var. sinuata (L.) Hochr. [s.l.] =Urena sinuata 

Urena moenchii Gómez =Malachra palmata 

Urena reticulata Cav. =Urena lobata 

Wissadula amplissima (L.) R.E. Fr. [non Sida amplissima L.] =Wissadula hernandioides 

Wissadula patens (A. St.-Hil.) Garcke =Wissadula excelsior 

Wissadula spicata (Kunth) C. Presl =Briquetia spicata 

Wissadula zeylanica Medik. =Wissadula periplocifolia 

REFERENCES 

Clement, I.D. 1957. Studies in Sida. I. Contrib. Gray Herb. 180: 1-91. 

Dorr, L.J. 2002. Malvaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


431. 

Fryxell, P.A. 2001. Malvaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 186-219. 

Jansen-Jacobs, M.J. 1986. Malvaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 3(1-2): 

259-276. 

Kearney, T.H. 1957. A tentative key to the South American species of Hibiscus L. Leafl. W. Bot. 8(7): 161-168.

396 


Kearney, T.H. 1958. A tentative key to the South American species of Pavonia Cav. Leafl. W. Bot. 8(10): 225-248. 

Kearney, T.H. 1958. A tentative key to the South American species of Sida L. Leafl. W. Bot. 8(11): 249-270. 

Krapovickas, A. & C.L. Cristobal. 1965. Revisión del genero Peltaea. Kurtziana 2: 135-216. 

Uittien, H. 1932. Malvaceae. In A. Pulle, Flora of Suriname 3(1): 1-25. 

Uittien, H. 1941. Malvaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 433-435. 

MARCGRAVIACEAE 

A.C. de Roon, S. Dressler 

Marcgravia coriacea Vahl 

Marcgravia eichleriana Wittm. 

Marcgravia magnibracteata Lanj. & Heerdt 

Marcgravia maguirei de Roon 

Marcgravia patellulifera de Roon 

Marcgravia pedunculosa Triana & Planch. 

Marcgravia punctifolia S. Dressler 

Marcgravia purpurea I.W. Bailey 

Marcgravia sororopaniana Steyerm. 

Marcgravia sprucei (Wittm.) Gilg 

Marcgraviastrum mixtum (Triana & Planch.) Bedell ex de Roon & S. Dressler 

Marcgraviastrum pendulum (Lanj. & Heerdt) Bedell 

Norantea guianensis Aubl. 

Norantea guianensis subsp. guianensis 

Norantea guianensis subsp. japurensis (Mart.) Bedell 

Sarcopera flammifera de Roon & Bedell 

Sarcopera tepuiensis (de Roon) Bedell 

Sarcopera tepuiensis subsp. coccinea de Roon & S. Dressler 

Sarcopera tepuiensis subsp. tepuiensis 

Souroubea dasystachya Gilg & Werderm. 

Souroubea guianensis Aubl. 

Souroubea guianensis subsp. cylindrica (Wittm.) de Roon 

Souroubea guianensis subsp. guianensis 


Marcgravia cuyuniensis I.W. Bailey =Marcgravia coriacea 

Marcgravia dasystachya Gilg ex Ule [nomen nudum] 

Marcgravia dubia Kunth =(Araceae) Monstera dubia 

Marcgravia gracilis Sagot =Marcgravia pedunculosa 

Marcgravia parviflora Rich. ex Wittm. =Marcgravia pedunculosa 

Marcgravia parviflora var. macrophylla Wittm. =Marcgravia pedunculosa 

Marcgravia parviflora var. sprucei Wittm. =Marcgravia sprucei 

Marcgravia picta Willd. [dubious species, misident. fide de Roon] 

Marcgravia roraimae Gilg & Werderm. =Marcgravia coriacea 

Marcgravia roraimensis Gilg ex Ule [nomen nudum] 

Marcgravia steyermarkii de Roon =Marcgravia sororopaniana 

Marcgravia umbellata L. [Guianan records misident., fide de Roon] 

Norantea guianensis var. gracilis Wittm. =Norantea guianensis subsp. japurensis 

Norantea japurensis Mart. =Norantea guianensis subsp. japurensis 

Norantea mixta Triana & Planch. =Marcgraviastrum mixtum 

Norantea paraensis Mart. =Norantea guianensis subsp. japurensis 

Norantea peduncularis Poepp. ex Wittm. =Marcgraviastrum mixtum 

Norantea pendula Lanj. & Heerdt =Marcgraviastrum pendulum 

Norantea tepuiensis de Roon =Sarcopera tepuiensis 

Souroubea guianensis var. cylindrica Wittm. =Souroubea guianensis subsp. cylindrica 

Souroubea guianensis var. tomentella Steyerm. =Souroubea guianensis subsp. cylindrica 


AM 

GU SU FG 

AM BO FG 

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BO GU FG? 

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REFERENCES 

Bailey, I.W. 1922. The pollination of Marcgravia: a classical case of ornithophily? Amer. J. Bot. 9: 370-384. 

De Roon, A.C. 1970. A new species of Marcgravia (Marcgraviaceae) from Venezuela. Acta Bot. Neerl. 19 (6): 799-803. 

De Roon, A.C. & S. Dressler. 1999. New taxa of Sarcopera and Marcgraviastrum (Marcgraviaceae) from the Guayana Shield. 

Novon 9: 248-252. 

Dressler, S. 2001. Marcgraviaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 248-260. 

Heald, S.V., A.C. De Roon, & S. Dressler. Marcgraviaceae. 1999. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. 


Mem. New York Bot. Gard. 76 (2): 431-437.


Lanjouw, J. & P.F. van Heerdt. 1941. Marcgraviaceae. In A. Pulle, Flora of Suriname 3(1): 373-385. 

MELASTOMATACEAE 

F. Almeda, P.E. Berry, A. Freire-Fierro, A. Gröger, B.K. Holst, N.G. Luckana, F.A. Michelangeli, 

T. Morley, D. Penney, S.S. Renner, O.R. Robinson, J.J. Wurdack†, K. Yatskievych 

Acanthella pulchra Gleason 

Acanthella sprucei Hook. f. 

Aciotis acuminifolia (Mart. ex DC.) Triana 

Aciotis annua (Mart. ex DC.) Triana [expected in DA] 

Aciotis circaeifolia (Bonpl.) Triana 

Aciotis indecora (Bonpl.) Triana 

Aciotis olivieriana Freire-Fierro 

Aciotis ornata (Miq.) Gleason 

Aciotis polystachya (Bonpl.) Triana 

Aciotis purpurascens (Aubl.) Triana 

Aciotis rubricaulis (Schrank & Mart. ex DC.) Triana 

Aciotis viscida (Benth.) Freire-Fierro 

Acisanthera bivalvis (Aubl.) Cogn. 

Acisanthera crassipes (Naudin) Wurdack 

Acisanthera hedyotidea (C. Presl) Triana 

Acisanthera limnobios (DC.) Triana 

Acisanthera nana Ule 

Acisanthera quadrata Pers. 

Acisanthera rosulans Huber 

Acisanthera tetraptera (Cogn.) Gleason 

Acisanthera uniflora (Vahl) Gleason 

Adelobotrys adscendens (Sw.) Triana 

Adelobotrys ayangannensis Wurdack 

Adelobotrys barbata Triana 

Adelobotrys ciliata (Naudin) Triana 

Adelobotrys duidae (Gleason) Wurdack 

Adelobotrys fruticosa Wurdack 

Adelobotrys linearifolia L. Uribe 

Adelobotrys monticola Gleason 

Adelobotrys permixta Wurdack 

Adelobotrys rotundifolia Triana 

Adelobotrys saxosa Wurdack 

Adelobotrys scandens (Aubl.) DC. 

Adelobotrys spruceana Cogn. 

Adelobotrys stenophylla Wurdack 

Appendicularia thymifolia (Bonpl.) DC. 

Bellucia grossularioides (L.) Triana 

Bellucia huberi (Wurdack) S.S. Renner 

Bellucia pentamera Naudin 

Bertolonia repens F. Michelangeli, sp. nov. ined. 

Bertolonia venezuelensis Wurdack 

Blakea rosea (Ruiz & Pav.) D. Don 

Boyania ayangannae Wurdack 

Centronia neblinae Wurdack 

Chaetolepis anisandra Naudin 

Chaetolepis phelpsiae Gleason 

Chaetolepis phelpsiae subsp. chimantensis Wurdack 

Chaetolepis phelpsiae subsp. phelpsiae 

Clidemia acurensis Wurdack 

Clidemia alternifolia Wurdack 

Clidemia andersonii Wurdack 

Clidemia anisophylla DC. 

Clidemia aphanantha (Naudin) Sagot 

Clidemia attenuata (Naudin) Cogn. 

Clidemia ayangannensis Wurdack 

Clidemia bernardii Wurdack 

Clidemia bullosa DC. 

Clidemia buntingii Wurdack 

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398 

Clidemia capillipes (Triana) Cogn. 

Clidemia capitata Benth. 

Clidemia capitellata (Bonpl.) D. Don 

Clidemia capitellata var. capitellata 

Clidemia capitellata var. dependens (D. Don) J.F. Macbr. 

Clidemia capitellata var. levelii Wurdack 

Clidemia charadrophila Tutin 

Clidemia conglomerata DC. 

Clidemia debilis Crueg. [expected in GU] 

Clidemia dentata D. Don 

Clidemia duidae Gleason 

Clidemia epibaterium DC. 

Clidemia epiphytica (Triana) Cogn. 

Clidemia granvillei Wurdack 

Clidemia heptamera Wurdack 

Clidemia heteroneura (DC.) Cogn. 

Clidemia hirta (L.) D. Don 

Clidemia hirta var. elegans (Aubl.) Griseb. 

Clidemia hirta var. hirta 

Clidemia hirta var. tiliifolia (DC.) J.F. Macbr. 

Clidemia involucrata DC. 

Clidemia japurensis DC. 

Clidemia japurensis var. heterobasis (DC.) Wurdack 

Clidemia japurensis var. japurensis 

Clidemia juruensis (Pilg.) Gleason 

Clidemia kappleri Cogn. 

Clidemia laevifolia Gleason 

Clidemia linearis (Gleason) Wurdack 

Clidemia marahuacensis Wurdack 

Clidemia micrantha Sagot 

Clidemia microthyrsa R.O. Williams 

Clidemia minutiflora (Triana) Cogn. 

Clidemia morichensis Wurdack 

Clidemia neblinae Wurdack 

Clidemia novemnervia (DC.) Triana 

Clidemia octona (Bonpl.) L.O. Williams 

Clidemia octona subsp. guayanensis Wurdack 

Clidemia octona subsp. octona 

Clidemia ostentata Wurdack 

Clidemia pakaraimae Wurdack 

Clidemia petiolata DC. 

Clidemia piperifolia Gleason 

Clidemia pustulata DC. 

Clidemia pycnaster Tutin 

Clidemia pycnaster subsp. pycnaster 

Clidemia pycnaster subsp. robusta Wurdack 

Clidemia rubra (Aubl.) Mart. 

Clidemia sandwithii Wurdack 

Clidemia saülensis Wurdack 

Clidemia septuplinervia Cogn. 

Clidemia sericea D. Don 

Clidemia sessiliflora (Naudin) Cogn. 

Clidemia siapensis Wurdack 

Clidemia silvicola Gleason 

Clidemia simpsonii Wurdack 

Clidemia stellipilis (Gleason) Wurdack 

Clidemia strigillosa (Sw.) DC. 

Clidemia tepuiensis Wurdack 

Clidemia tococoidea (DC.) Gleason 

Clidemia trinitensis (Crueg.) Griseb. 

Clidemia ulei Pilg. 

Clidemia urceolata DC. 

Clidemia venosa (Gleason) Wurdack 

Comolia ayangannae Wurdack 


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Comolia coriacea Gleason 

Comolia latifolia (Aubl.) Cogn. [dubious species] 

Comolia leptophylla (Bonpl.) Naudin 

Comolia microphylla Benth. 

Comolia montana Gleason 

Comolia nummularioides (Bonpl.) Naudin 

Comolia prostrata Wurdack 

Comolia serpyllacea Wurdack 

Comolia smithii Wurdack 

Comolia vernicosa (Benth.) Triana 

Comolia villosa (Aubl.) Triana 

Comoliopsis neblinae Wurdack [Sierra de la Neblina] 

Conostegia superba Naudin [Sierra de la Neblina] 

Desmoscelis villosa (Aubl.) Naudin 

Ernestia blackii Brade & Markgr. 

Ernestia cataractae Tutin 

Ernestia confertiflora Wurdack 

Ernestia cordifolia O. Berg ex Triana 

Ernestia glandulosa Gleason 

Ernestia granvillei Wurdack 

Ernestia maguirei Wurdack 

Ernestia minor Gleason 

Ernestia pullei Gleason 

Ernestia rubra Pulle 

Ernestia subglabra Wurdack 

Ernestia tenella (Bonpl.) DC. 

Ernestia tenella var. sprucei Cogn. 

Ernestia tenella var. tenella 

Graffenrieda caryophyllea Triana 

Graffenrieda caudata Wurdack 

Graffenrieda cinnoides Gleason 

Graffenrieda fantastica R.E. Schult. & L.B. Sm. 

Graffenrieda fruticosa Wurdack 

Graffenrieda hitchcockii Gleason 

Graffenrieda intermedia Triana 

Graffenrieda irwinii Wurdack 

Graffenrieda jauana Wurdack 

Graffenrieda kralii Wurdack 

Graffenrieda lanceolata Gleason 

Graffenrieda miconioides Naudin 

Graffenrieda obliqua Triana 

Graffenrieda patens Triana 

Graffenrieda pedunculata Gleason 

Graffenrieda polymera Gleason 

Graffenrieda polymera subsp. neblinensis Wurdack 

Graffenrieda polymera subsp. polymera 

Graffenrieda reticulata Wurdack 

Graffenrieda rotundifolia (Bonpl.) DC. 

Graffenrieda rufa Wurdack 

Graffenrieda rupestris Ducke 

Graffenrieda sessilifolia Triana 

Graffenrieda sessilifolia subsp. A 

Graffenrieda sessilifolia subsp. cardonae Wurdack 

Graffenrieda sessilifolia subsp. occidentalis Wurdack 

Graffenrieda sessilifolia subsp. sessilifolia 

Graffenrieda sipapoana Wurdack 

Graffenrieda steyermarkii Wurdack 

Graffenrieda tricalcarata Gleason 

Graffenrieda versicolor Gleason 

Graffenrieda weddellii Naudin 

Henriettea granulata O. Berg ex Triana 

Henriettea horridula Pilg. 

Henriettea maroniensis Sagot 

Henriettea martiusii (DC.) Naudin 


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400 

Henriettea mucronata (Gleason) Renner 

Henriettea multiflora Naudin 

Henriettea patrisiana DC. 

Henriettea ramiflora (Sw.) DC. 

Henriettea spruceana Cogn. 

Henriettea stellaris O. Berg ex Triana 

Henriettea succosa (Aubl.) DC. 

Henriettella caudata Gleason 

Henriettella duckeana Hoehne 

Henriettella flavescens (Aubl.) Triana 

Henriettella heteroneura Gleason 

Henriettella ininiensis Wurdack 

Henriettella ovata Cogn. 

Henriettella prancei Wurdack 

Henriettella steyermarkii Wurdack 

Leandra agrestis (Aubl.) Raddi 

Leandra aristigera (Naudin) Cogn. 

Leandra candelabrum (J.F. Macbr.) Wurdack 

Leandra chaetodon (DC.) Cogn. 

Leandra clidemioides (Naudin) Wurdack 

Leandra cremersii Wurdack 

Leandra cuneata (Mart.) Cogn. 

Leandra divaricata (Naudin) Cogn. 

Leandra edentula Gleason 

Leandra francavillana Cogn. 

Leandra glandulifera (Triana) Cogn. 

Leandra gorzulae Wurdack 

Leandra intermedia (DC.) Cogn. 

Leandra lindeniana (Naudin) Cogn. 

Leandra longisepala Wurdack 

Leandra maguirei Wurdack [Sierra de la Neblina] 

Leandra micrantha Wurdack 

Leandra micropetala (Naudin) Cogn. 

Leandra neblinensis Wurdack 

Leandra paleacea Wurdack 

Leandra phelpsiae Gleason 

Leandra polyadena Ule 

Leandra procumbens Ule 

Leandra purpurea Gleason 

Leandra rufescens (DC.) Cogn. 

Leandra sanguinea Gleason 

Leandra sanguinea subsp. sanguinea 

Leandra sanguinea subsp. tepuiensis Wurdack 

Leandra secunda (D. Don) Cogn. 

Leandra solenifera Cogn. 

Leandra steyermarkii Wurdack 

Leandra verticillata (DC.) Cogn. 

Loreya arborescens (Aubl.) DC. 

Loreya mespiloides Miq. 

Loreya ovata O. Berg ex Triana 

Loreya spruceana Benth. ex Triana 

Loreya strigosa Gleason 

Loreya subrotundifolia (Wurdack) S.S. Renner 

Macairea axilliflora Wurdack 

Macairea cardonae Wurdack 

Macairea chimantensis Wurdack 

Macairea duidae Gleason 

Macairea lanata Gleason 

Macairea lasiophylla (Benth.) Wurdack 

Macairea linearis Gleason 

Macairea maroana Wurdack 

Macairea multinervia Benth. 

Macairea neblinae Wurdack 

Macairea pachyphylla Benth. 


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Macairea parvifolia Benth. 

Macairea rigida Benth. 

Macairea rufescens DC. 

Macairea spruceana O. Berg ex Triana 

Macairea stylosa Triana 

Macairea thyrsiflora DC. 

Macrocentrum anfractum Wurdack 

Macrocentrum angustifolium Gleason 

Macrocentrum anychioides Gleason 

Macrocentrum brevipedicellatum Wurdack 

Macrocentrum chimantense Wurdack 

Macrocentrum cristatum (DC.) Triana 

Macrocentrum cristatum var. cristatum 

Macrocentrum cristatum var. microphyllum Cogn. 

Macrocentrum cristatum var. parviflorum (DC.) Cogn. 

Macrocentrum droseroides Triana 

Macrocentrum fasciculatum (Rich. ex DC.) Triana 

Macrocentrum fruticosum Gleason 

Macrocentrum gesneriaceum Sandwith 

Macrocentrum gracile Wurdack 

Macrocentrum huberi Wurdack 

Macrocentrum latifolium Wurdack 

Macrocentrum longidens (Gleason) Wurdack 

Macrocentrum maguirei Wurdack 

Macrocentrum minus Gleason 

Macrocentrum neblinae Wurdack 

Macrocentrum parvulum Gleason 

Macrocentrum repens (Gleason) Wurdack 

Macrocentrum rubescens Gleason 

Macrocentrum steyermarkii Wurdack 

Macrocentrum stipulaceum Wurdack 

Macrocentrum vestitum Sandwith 

Maguireanthus ayangannae Wurdack 

Maieta guianensis Aubl. 

Maieta neblinensis Wurdack 

Maieta poeppigii Mart. ex Cogn. 

Mallophyton chimantense Wurdack 

Marcetia taxifolia (A. St.-Hil.) DC. 

Meriania broccha Wurdack 

Meriania crassiramis (Naudin) Wurdack 

Meriania neblinensis Wurdack 

Meriania ornata Wurdack 

Meriania sclerophylla (Naudin) Triana 

Meriania urceolata Triana 

Miconia abbreviata Markgr. 

Miconia abysmophila Wurdack 

Miconia acinodendron (L.) Sweet 

Miconia acuminata (Steud.) Naudin 

Miconia acutifolia Ule 

Miconia aeruginosa Naudin 

Miconia affinis DC. 

Miconia aguitensis Gleason 

Miconia alata (Aubl.) DC. 

Miconia albicans (Sw.) Triana 

Miconia alborufescens Naudin 

Miconia aliquantula Wurdack 

Miconia alternans Naudin 

Miconia amacurensis Wurdack 

Miconia ampla Triana 

Miconia aplostachya (Bonpl.) DC. 

Miconia argyrophylla DC. 

Miconia argyrophylla subsp. argyrophylla 

Miconia argyrophylla subsp. gracilis Wurdack 

Miconia aristata Gleason 


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402 

Miconia biglandulosa Gleason 

Miconia biglomerata (Bonpl.) DC. 

Miconia borjensis Wurdack 

Miconia brachybotrya Triana 

Miconia bracteata (DC.) Triana 

Miconia brevipes Benth. 

Miconia bubalina (D. Don) Naudin 

Miconia buntingii Wurdack 

Miconia cacatin (Aubl.) S.S. Renner 

Miconia cacumina Wurdack 

Miconia campestris (Benth.) Triana 

Miconia carassana Cogn. 

Miconia cautis Wurdack 

Miconia centrodesma Naudin 

Miconia ceramicarpa (DC.) Cogn. 

Miconia ceramicarpa var. candolleana Cogn. 

Miconia ceramicarpa var. ceramicarpa 

Miconia ceramicarpa var. crozieriae Cremers & C.V. Martin 

Miconia chrysophylla (Rich.) Urb. 

Miconia ciliata (Rich.) DC. 

Miconia cowanii Wurdack [perhaps expected in FG] 

Miconia crassinervia Cogn. Cogn. 

Miconia curta (Gleason) Wurdack 

Miconia curta subsp. curta 

Miconia curta subsp. ptariensis Wurdack 

Miconia decurrens Cogn. 

Miconia diaphanea Gleason 

Miconia dioica Wurdack 

Miconia dispar Benth. 

Miconia dodecandra Cogn. 

Miconia dolichorrhyncha Naudin 

Miconia egensis Cogn. 

Miconia elaeoides Naudin 

Miconia elata (Sw.) DC. 

Miconia eriodonta DC. 

Miconia eugenioides Triana 

Miconia fallax DC. 

Miconia fanshawei Wurdack 

Miconia fragilis Naudin 

Miconia fragrans Cogn. 

Miconia francavillana Cogn. 

Miconia gratissima Benth. ex Triana 

Miconia grossidentata Wurdack 

Miconia guaiquinimae Wurdack 

Miconia holosericea (L.) DC. 

Miconia huberi Wurdack 

Miconia hypoleuca (Benth.) Triana 

Miconia ibaguensis (Bonpl.) Triana 

Miconia iluensis Wurdack 

Miconia impetiolaris (Sw.) D. Don ex DC. 

Miconia impetiolaris var. spruceana Cogn. 

Miconia inaequalifolia Triana 

Miconia kappleri Naudin 

Miconia kavanayensis Wurdack 

Miconia laevigata (L.) DC. 

Miconia lappacea (DC.) Triana 

Miconia lasseri Gleason 

Miconia lateriflora Cogn. 

Miconia lateriflora subsp. lateriflora 

Miconia lateriflora subsp. monticellensis Wurdack 

Miconia lepidota DC. 

Miconia longifolia (Aubl.) DC. 

Miconia longispicata Triana 

Miconia loreyoides Triana 


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Miconia lourteigiana Wurdack 

Miconia macrothyrsa Benth. 

Miconia macrotis Cogn. 

Miconia maguirei Gleason 

Miconia marginata Triana 

Miconia mariae Wurdack 

Miconia maroana Wurdack 

Miconia matthaei Naudin 

Miconia melinonis Naudin 

Miconia minutiflora (Bonpl.) DC. 

Miconia mirabilis (Aubl.) L.O. Williams 

Miconia mituana Wurdack 

Miconia mutabilis (DC.) Triana 

Miconia myriantha Benth. 

Miconia navioensis Wurdack [expected in FG] 

Miconia neblinensis Wurdack [expected in AM, Serra da Neblina] 

Miconia nervosa (Sm.) Triana 

Miconia oldemanii Wurdack 

Miconia perobscura Wurdack [Sierra de la Neblina] 

Miconia perturbata Wurdack 

Miconia phaeophylla Triana 

Miconia pileata DC. 

Miconia pilgeriana Ule 

Miconia platypoda Gleason 

Miconia plukenetii Naudin 

Miconia poeppigii Triana 

Miconia polita Gleason 

Miconia prasina (Sw.) DC. 

Miconia pseudoaplostachya Cogn. 

Miconia pseudocapsularis Wurdack 

Miconia pubipetala Miq. 

Miconia punctata (Desr.) D. Don ex DC. 

Miconia pyrifolia Naudin 

Miconia racemosa (Aubl.) DC. 

Miconia radulaefolia (Benth.) Naudin 

Miconia regelii Cogn. 

Miconia rhytidophylla Naudin 

Miconia roraimensis Ule [Mt. Roraima] 

Miconia rubiginosa (Bonpl.) DC. 

Miconia rufescens (Aubl.) DC. 

Miconia ruficalyx Gleason 

Miconia rugosa Triana 

Miconia rupestris Ule 

Miconia rupticalyx Wurdack 

Miconia sagotiana Cogn. 

Miconia sarmentosa Cogn. 

Miconia sastrei Wurdack 

Miconia serialis DC. 

Miconia serrulata (DC.) Naudin 

Miconia silicicola Gleason 

Miconia solmsii Cogn. 

Miconia splendens (Sw.) Griseb. 

Miconia sprucei Triana 

Miconia stenostachya DC. 

Miconia stephananthera Ule 

Miconia steyermarkii Gleason 

Miconia strigosa (Triana) Wurdack 

Miconia superba Ule 

Miconia tetraspermoides Wurdack 

Miconia theizans (Bonpl.) Cogn. 

Miconia tillettii Wurdack 

Miconia tinifolia Naudin 

Miconia tomentosa (Rich.) D. Don ex DC. 

Miconia traillii Cogn. 

BO 


BO 

GU 

AM BO GU 

BO GU 

AM 


FG 



AM 

FG 


GU? 


FG 

AM 

AM BO 


AM BO FG 

AM BO 

GU SU 



BO GU 


AM BO GU FG 

AM BO GU 





AM GU 

GU FG? 

AM GU? SU? 

AM BO GU? 

AM BO GU SU 



AM BO GU SU 

AM BO GU 

AM GU 

GU SU FG 

SU FG 

FG 



AM BO GU 

AM BO 


AM 

AM BO GU SU 


AM 

AM 

BO GU 

BO GU SU FG 



AM BO GU 


AM GU SU FG

404 

Miconia trimera Wurdack 

Miconia trinervia (Sw.) D. Don ex Loudon 

Miconia truncata Triana 

Miconia tschudyoides Cogn. 

Miconia undata Triana 

Miconia wittii Ule 

Miconia yatuensis Wurdack 

Microlicia benthamiana Triana ex Cogn. 

Microlicia guanayana Wurdack 

Monochaetum bonplandii (Kunth) Naudin 

Mouriri acutiflora Naudin 

Mouriri angulicosta Morley 

Mouriri angustifolia Spruce ex Triana 

Mouriri brevipes Hook. 

Mouriri cauliflora Mart. ex DC. 

Mouriri collocarpa Ducke 

Mouriri crassifolia Sagot 

Mouriri densifoliata Ducke 

Mouriri duckeana Morley 

Mouriri dumetosa Cogn. 

Mouriri ficoides Morley 

Mouriri francavillana Cogn. 

Mouriri grandiflora DC. 

Mouriri guianensis Aubl. 

Mouriri huberi Cogn. 

Mouriri latihila Morley 

Mouriri longifolia (Kunth) Morley 

Mouriri myrtifolia Spruce ex Triana [expected in GU] 

Mouriri myrtilloides (Sw.) Poir. 

Mouriri myrtilloides subsp. orinocensis Morley 

Mouriri nervosa Pilg. 

Mouriri nigra (DC.) Morley 

Mouriri oligantha Pilg. 

Mouriri pauciflora Spruce ex Cogn. 

Mouriri rhizophorifolia (DC.) Triana 

Mouriri sagotiana Triana 

Mouriri sideroxylon Sagot ex Triana 

Mouriri subumbellata Triana 

Mouriri uncitheca Morley & Wurdack 

Mouriri vernicosa Naudin 

Mouriri viridicosta Morley 

Myriaspora egensis DC. 

Neblinanthera cumbrensis Wurdack 

Nepsera aquatica (Aubl.) Naudin 

Ochthephilus repentinus Wurdack 

Opisthocentra clidemioides Hook. f. 

Ossaea coarctiflora Wurdack 

Ossaea mavacana Wurdack 

Ossaea micrantha (Sw.) Macfad. 

Ossaea quadrisulca (Naudin) Wurdack 

Pachyloma coriaceum DC. 

Pachyloma huberioides (Naudin) Triana 

Pachyloma pusillum Wurdack 

Pachyloma setosum Wurdack 

Phainantha laxiflora (Triana) Gleason 

Phainantha maguirei Wurdack 

Phainantha myrteoloides Wurdack 

Phainantha steyermarkii Wurdack 

Poteranthera pusilla Bong. 

Pterogastra divaricata (Bonpl.) Naudin 

Pterogastra minor Naudin 

Pterolepis glomerata (Rottb.) Miq. 

Pterolepis trichotoma (Rottb.) Cogn. 

Rhynchanthera dichotoma (Desr.) DC. 


SU FG 

AM BO FG 

AM 

SU FG 

AM BO 

AM 

AM 

AM BO GU 

AM 

BO GU 


BO SU FG 

AM 

AM BO GU SU 

AM 

AM GU SU FG 

GU? SU FG 

AM 

GU SU FG 

FG 

AM 

FG 

AM GU SU FG 



AM 

AM 

AM BO 

AM BO 

AM BO 

BO SU FG 


FG 

AM FG 

BO 



AM FG 

AM 

AM GU SU FG 

FG 


AM 


GU 

AM 

FG 

AM 

AM 

FG 

GU 

AM 

AM 

AM 

BO GU 

BO GU 

BO GU 

BO 

AM BO 

AM BO GU 

AM BO 

BO GU SU FG 


DA GU FG

Rhynchanthera grandiflora (Aubl.) DC. 

Rhynchanthera hispida Naudin 

Rhynchanthera serrulata (Rich.) DC. 

Salpinga glandulosa (Gleason) Wurdack [expected in GU] 

Salpinga maguirei Gleason 

Salpinga paleacea Wurdack 

Salpinga pusilla (Gleason) Wurdack 

Salpinga secunda Schrank & Mart. ex DC. 

Sandemania hoehnei (Cogn.) Wurdack [Sierra de la Neblina] 

Siphanthera cordifolia (Benth.) Gleason 

Siphanthera cowanii Wurdack 

Siphanthera dawsonii Wurdack 

Siphanthera duidae (Gleason) Wurdack 

Siphanthera fasciculata (Gleason) Almeda & O.R. Rob. 

Siphanthera foliosa (Naudin) Wurdack 

Siphanthera hostmannii Cogn. 

Tateanthus duidae Gleason 

Tibouchina aspera Aubl. 

Tibouchina aspera var. aspera 

Tibouchina aspera var. asperrima Cogn. 

Tibouchina bipenicillata (Naudin) Cogn. 

Tibouchina catharinae Pittier 

Tibouchina cryptadena Gleason 

Tibouchina dissitiflora Wurdack 

Tibouchina duidae Gleason 

Tibouchina fraterna N.E. Br. 

Tibouchina fraterna subsp. fraterna 

Tibouchina fraterna subsp. paruana Wurdack 

Tibouchina geitneriana (Schltdl.) Cogn. 

Tibouchina gracilis (Bonpl.) Cogn. 

Tibouchina huberi Wurdack 

Tibouchina kunhardtii Gleason 

Tibouchina llanorum Wurdack 

Tibouchina longifolia (Vahl) Baill. 

Tibouchina multiflora (Gardner) Cogn. 

Tibouchina sipapoana Gleason 

Tibouchina spruceana Cogn. [expected in BO] 

Tibouchina steyermarkii Wurdack 

Tibouchina striphnocalyx (DC.) Gleason 

Tococa aristata Benth. 

Tococa bolivarensis Gleason 

Tococa bolivarensis subsp. bolivarensis 

Tococa bolivarensis subsp. occidentalis Wurdack 

Tococa bullifera Triana 

Tococa caryophyllea (DC.) S.S. Renner 

Tococa ciliata Triana 

Tococa cinnamomea Triana 

Tococa cordata O. Berg ex Triana 

Tococa coronata Benth. 

Tococa desiliens Gleason 

Tococa erythrophylla (Ule) Wurdack 

Tococa guianensis Aubl. 

Tococa hirta O. Berg ex Triana 

Tococa lancifolia Spruce ex Triana 

Tococa liesneri Wurdack 

Tococa macrophysca Spruce ex Triana 

Tococa macrosperma Mart. 

Tococa nitens (Benth.) Triana 

Tococa obovata Gleason 

Tococa obovata subsp. neblinensis Wurdack 

Tococa obovata subsp. obovata 

Tococa oligantha Gleason 

Tococa pachystachya Wurdack 

Tococa pauciflora Spruce ex Triana 



GU? SU 

BO GU SU FG 

AM BO 

AM 

GU 

AM 

AM 

AM 

AM BO GU 

AM 

BO 

AM 

AM 

BO GU? 

AM BO GU SU 

AM BO 



BO GU SU FG 

AM BO 

AM 

AM BO 

AM 

AM BO 

AM BO GU 

AM BO GU 

AM 

AM BO GU 

BO GU 

AM BO 

AM 

BO 

BO GU 

FG 

AM 

AM 

BO 

AM 

BO GU SU FG? 

AM BO GU 

AM BO GU 

AM BO 

AM 

AM 

AM 

AM 

AM 

AM BO GU 

GU 

AM BO GU 


AM 

AM 

AM 

AM 

AM BO GU 

AM BO GU 

AM BO 

AM 

AM BO 

AM BO 

AM 

AM

406 

Tococa rotundifolia (Triana) Wurdack 

Tococa subciliata (DC.) Triana 

Tococa tepuiensis Wurdack 

Tococa tepuiensis subsp. glabrata Wurdack 

Tococa tepuiensis subsp. tepuiensis 

Topobea ferruginea Gleason [perhaps expected in GU] 

Topobea parasitica Aubl. 

Tryssophyton merumense Wurdack 

Votomita guianensis Aubl. 

Votomita aff. monadelpha (Ducke) Morley 

Votomita orinocensis Morley 

Votomita ventuarensis Morley 


AM 

AM GU 

AM BO 

AM 

BO 

AM BO 

AM 

AM 


Acanthella montana Gleason =Acanthella sprucei 

Acanthella plicata Gleason =Acanthella pulchra 

Aciotis acutiflora (Mart.) Triana =Aciotis indecora 

Aciotis acutiflora var. parvifolia Cogn. =Aciotis indecora 

Aciotis aequatorialis Cogn. =Aciotis acuminifolia 

Aciotis alata (Beurl.) Almeda =Aciotis rubricaulis 

Aciotis amazonica Cogn. =Aciotis acuminifolia 

Aciotis amazonica var. radicans Cogn. =Aciotis acuminifolia 

Aciotis anomala Brade =Aciotis indecora 

Aciotis aristata Ule =Aciotis rubricaulis 

Aciotis asplundii Wurdack =Aciotis rubricaulis 

Aciotis caulialata (Ruiz & Pav.) Triana =Aciotis rubricaulis 

Aciotis circaeifolia var. major Cogn. =Aciotis purpurascens 

Aciotis cordata J.F. Macbr. [not in flora area] 

Aciotis dichotoma (Benth.) Cogn. =Aciotis acuminifolia 

Aciotis dichotoma var. anomala (Miq.) Cogn. =Aciotis acuminifolia 

Aciotis dichotoma var. longifolia S. Moore =Aciotis acuminifolia 

Aciotis discolor D. Don =Aciotis purpurascens 

Aciotis dysophylla (Benth.) Triana =Aciotis annua 

Aciotis fragilis (Rich. ex DC.) Cogn. =Aciotis purpurascens 

Aciotis fragilis var. lancifolia (Steud. ex Cogn.) Cogn. =Aciotis purpurascens 

Aciotis herbacea Steud. ex Cogn. =Aciotis ornata 

Aciotis indecora var. glabrescens Cogn. =Aciotis viscida 

Aciotis indecora var. macrophylla Cogn. =Aciotis indecora 

Aciotis indecora var. sagotiana Cogn. =Aciotis indecora 

Aciotis laxa (DC.) Cogn. =Aciotis indecora 

Aciotis laxa var. A =Aciotis viscida 

Aciotis laxa var. B =Aciotis indecora 

Aciotis laxa var. kappleriana (Naudin) Cogn. =Aciotis indecora 

Aciotis laxa var. laxa =Aciotis indecora 

Aciotis laxa var. robusta Cogn. =Aciotis indecora 

Aciotis levyana Cogn. =Aciotis rubricaulis 

Aciotis longifolia (DC.) Triana =Aciotis circaeifolia 

Aciotis longifolia var. glabra Huber =Aciotis purpurascens 

Aciotis martinicensis (Naudin) Urb. =Aciotis purpurascens 

Aciotis pellucida (Rich. ex DC.) Pittier et al. =Aciotis purpurascens 

Aciotis purpurascens var. alata (Beurl.) Cogn. =Aciotis rubricaulis p.p.; Aciotis purpurascens p.p. 

Aciotis purpurascens var. longifolia Cogn. =Aciotis purpurascens 

Aciotis purpurascens var. martinensis J.F. Macbr. =Aciotis rubricaulis 

Aciotis purpurascens var. pellucida (Rich. ex DC.) Cogn. =Aciotis purpurascens 

Aciotis rostellata (Naudin) Triana =Aciotis indecora 

Aciotis sieberi (Steud.) Triana =Aciotis purpurascens 

Aciotis sileniflora (Bonpl.) Triana =Aciotis purpurascens 

Aciotis sphaeranthera (Naudin) Cogn. =Aciotis acuminifolia 

Aciotis trinitensis Cogn. =Aciotis purpurascens 

Aciotis uliginosa (Bonpl. ex Naudin) Triana =Aciotis purpurascens 

Aciotis viscosa (Bonpl. ex Naudin) Triana =Aciotis indecora 

Acisanthera adscendens C. Wright =Acisanthera quadrata 

Acisanthera beccabunga (Mart. ex Naudin) Triana =Acisanthera bivalvis 

Acisanthera boissieriana Cogn. =Acisanthera hedyotidea 

SU FG 

GU 

GU SU FG 

SU


Acisanthera brevifolia (DC.) Griseb. =Acisanthera bivalvis 

Acisanthera erecta Gleason =Macairea lasiophylla 

Acisanthera gracilis Ule =Acisanthera crassipes 

Acisanthera inundata (DC.) Triana [not in flora area] 

Acisanthera inundata var. pusilla Cogn. =Acisanthera limnobios 

Acisanthera lasiophylla (Benth.) Gleason =Macairea lasiophylla 

Acisanthera pellucida Wright ex Griseb. =Acisanthera limnobios 

Acisanthera quadrata Juss. ex Poir. =Acisanthera quadrata 

Acisanthera recurva (Rich.) Griseb. =Acisanthera uniflora 

Acisanthera salzmannii (Naudin) Cogn. =Acisanthera hedyotidea 

Acisanthera trivalvis (Aubl.) Cogn. =Acisanthera bivalvis 

Adelobotrys guianensis (DC.) Gleason =Adelobotrys adscendens 

Adelobotrys laxiflora Triana =Phainantha laxiflora 

Adelobotrys monticola subsp. occidentalis Wurdack =Adelobotrys monticola 

Apatitia blakeoides Desv. =Bellucia grossularioides 

Arthrostema nummularioides (Bonpl.) DC. =Comolia nummularioides 

Arthrostemma angusturenese (Rich.) Cogn. =Pterolepis glomerata 

Arthrostemma caulialatum Ruiz & Pav. =Aciotis rubricaulis 

Arthrostemma glomerata (Rottb.) Cham. =Pterolepis glomerata 

Arthrostemma glomeratum (Rottb.) Cham. =Pterolepis glomerata 

Arthrostemma ladanoides (Rich.) DC. =Pterolepis trichotoma 

Arthrostemma latifolium D. Don [dubious species, type not found] 

Arthrostemma villosum (Aubl.) DC. =Comolia villosa 

Axinanthera macrophylla H. Karst. =Bellucia pentamera 

Bellucia arborescens (Aubl.) Baill. =Loreya arborescens 

Bellucia aricuaizensium Pittier =Bellucia pentamera 

Bellucia aubletii Naudin =Bellucia grossularioides 

Bellucia brasiliensis Naudin =Bellucia grossularioides 

Bellucia cacatin (Aubl.) Sagot =Miconia cacatin 

Bellucia circumscissa Spruce ex Cogn. =Bellucia grossularioides 

Bellucia hostmannii Naudin =Bellucia grossularioides 

Bellucia mespiloides (Miq.) J.F. Macbr. =Loreya mespiloides 

Bellucia nervosa Raf. =Bellucia grossularioides 

Bellucia quinquenervia (Aubl.) H. Karst. =Bellucia grossularioides 

Bellucia subrotundifolia Wurdack =Loreya subrotundifolia 

Blakea cacatin (Aubl.) D. Don ex DC. =Miconia cacatin 

Blakea calycanthus Markgr. =Blakea rosea 

Blakea caudata Triana =Blakea rosea 

Blakea quinquenervia Aubl. =Bellucia grossularioides 

Blakea triplinervia L. f. =Bellucia grossularioides 

Brachypremna petiolata Gleason =Ernestia pullei 

Calophysa tococoidea DC. =Clidemia tococoidea 

Capitellaria benthamii Naudin =Clidemia capitata 

Centronia crassiramis (Naudin) Triana =Meriania crassiramis 

Chaenophora ferruginea Crueg. =Miconia hypoleuca 

Chaenopleura ferruginea (Crueg.) Griseb. =Miconia hypoleuca 

Chaenopleura fulva (Rich.) Griseb. =Miconia chrysophylla 

Chaenopleura hypoleuca Benth. =Miconia hypoleuca 

Chaenopleura longifolia (Aubl.) Griseb. =Miconia longifolia 

Chaetogastra callichaeta Benth. =Pterolepis glomerata 

Chaetogastra geitneriana Schltdl. =Tibouchina geitneriana 

Chaetogastra glomerata (Rottb.) Benth. =Pterolepis glomerata 

Chaetogastra hypericoides DC. =Desmoscelis villosa 

Chaetogastra lasiophylla Benth. =Macairea lasiophylla 

Chaetogastra rhynchanthera Benth. =Monochaetum bonplandii 

Chaetolepis citrina Gleason =Chaetolepis anisandra 

Charianthus spp. [Caribbean taxon, none in flora area] 

Chitonia bubalina D. Don =Miconia bubalina 

Chitonia fothergilla (Desr.) D. Don =Miconia mirabilis 

Clidemia affinis (Naudin) Cogn. =Clidemia pustulata 

Clidemia agrestis (Aubl.) D. Don =Leandra agrestis 

Clidemia aristigera Naudin =Leandra aristigera 

Clidemia benthamiana Miq. =Clidemia hirta var. hirta 

Clidemia berbiceana DC. =Miconia acinodendron

408 


Clidemia biformis Gleason =Clidemia anisophylla 

Clidemia bonplandii Cogn. =Clidemia debilis 

Clidemia bracteata DC. =Miconia bracteata 

Clidemia campestris Benth. =Miconia campestris 

Clidemia campestris var. pauciflora Benth. =Miconia biglomerata 

Clidemia candolleana Cogn. [nom. illeg.] =Clidemia bullosa 

Clidemia ceramicarpa DC. =Miconia ceramicarpa 

Clidemia chaetodon (DC.) Naudin =Leandra chaetodon 

Clidemia coccinea DC. =Miconia ceramicarpa var. ceramicarpa 

Clidemia coriacea (Naudin) Cogn. =Clidemia tepuiensis 

Clidemia depauperata DC. =Leandra agrestis 

Clidemia dependens D. Don =Clidemia capitellata var. dependens 

Clidemia desmantha Benth. =Miconia rhytidophylla 

Clidemia divaricata Naudin =Leandra divaricata 

Clidemia drosera Sagot ? =Clidemia micrantha 

Clidemia elegans (Aubl.) D. Don =Clidemia hirta var. elegans 

Clidemia glabrata Steud. =Miconia ibaguensis 

Clidemia heterobasis DC. =Clidemia japurensis var. heterobasis 

Clidemia heteroclita Naudin =Clidemia japurensis var. japurensis 

Clidemia lappacea DC. =Miconia lappacea 

Clidemia lutescens var. lindeniana Naudin =Leandra lindeniana 

Clidemia maculata Benth. =Miconia ibaguensis 

Clidemia manoacensis Kraenzl. =Clidemia bullosa 

Clidemia martii Naudin =Miconia centrodesma 

Clidemia miconioides Benth. =Miconia ibaguensis 

Clidemia mutabilis DC. =Miconia mutabilis 

Clidemia naevula (Naudin) Triana =Clidemia japurensis var. heterobasis 

Clidemia neglecta D. Don = Clidemia capitellata (Bonpl.) D.Don 

Clidemia novemnervia var. affinis (Naudin) Wurdack =Clidemia pustulata 

Clidemia novemnervia var. depauperata Pittier =Clidemia bullosa 

Clidemia petiolata var. angustifolia Cogn. =Clidemia petiolata 

Clidemia petiolata var. colorata Cogn. =Clidemia petiolata 

Clidemia radulaefolia Benth. =Miconia radulaefolia 

Clidemia rariflora Benth. =Clidemia bullosa 

Clidemia rhamnifolia var. macrodon Naudin =Leandra divaricata 

Clidemia rubra var. biacuta (Naudin) Cogn. =Clidemia sericea 

Clidemia secunda D. Don =Leandra secunda 

Clidemia smithii Gleason ? =Clidemia laevifolia x Clidemia hirta 

Clidemia spicata (Aubl.) DC. =Clidemia capitellata var. dependens 

Clidemia surinamensis Miq. =Clidemia novemnervia 

Clidemia tiliifolia DC. =Clidemia hirta var. tiliifolia 

Clidemia trichodes DC. =Miconia bracteata 

Clidemia tristis (Naudin) Sagot =Miconia bracteata 

Clidemia umbonata DC. =Clidemia strigillosa 

Clidemia violacea DC. =Miconia ceramicarpa var. ceramicarpa 

Clidemia virgata Pittier =Miconia ibaguensis 

Comolia affinis Hoehne =Comolia villosa 

Comolia angustifolia Gleason =Comolia villosa 

Comolia anomala Pittier =Acisanthera uniflora 

Comolia hirtella Naudin =Comolia villosa 

Comolia hoehnei Cogn. =Sandemania hoehnei 

Comolia kuhlmannii Hoehne =Comolia villosa 

Comolia lythrarioides Naudin =Comolia villosa 

Comolia neglecta Cogn. =Comolia villosa 

Comolia purpurea Miq. =Comolia villosa 

Comolia surinamensis Miq. =Comolia villosa 

Comolia tetraptera Cogn. =Acisanthera tetraptera 

Comolia veronicaefolia Benth. =Comolia villosa 

Conostegia acuminata Steud. =Miconia acuminata 

Conostegia icosandra (Sw.) Urb. [not in flora area] 

Conostegia petiolata Gleason [not in flora area] =Conostegia montana (Sw.) DC. 

Copedesma nitens Gleason =Miconia ruficalyx 

Coryphadenis abnormis (Naudin) Morley =Votomita guianensis 

Cremanium trinitatis Crueg. =Miconia minutiflora


Davya adscendens (Sw.) Griseb. =Adelobotrys adscendens 

Davya ciliata Naudin =Adelobotrys ciliata 

Davya crassiramis Naudin =Meriania crassiramis 

Davya guianensis DC. =Adelobotrys adscendens 

Davya scandens (Aubl.) Naudin =Adelobotrys scandens 

Davya sclerophylla Naudin =Meriania sclerophylla 

Decaraphe floribunda Miq. =Miconia acinodendron 

Decaraphe fockeana Miq. =Miconia acinodendron 

Decaraphe hostmannii Steud. =Miconia serrulata 

Desmoscelis mollis Pittier =Desmoscelis villosa 

Dicrananthera hedyotidea C. Presl =Acisanthera hedyotidea 

Dicrananthera salzmannii Naudin =Acisanthera hedyotidea 

Diolena longidens Gleason =Macrocentrum longidens 

Diolena repens Gleason =Macrocentrum repens 

Diplochita bubalina (D. Don) DC. =Miconia bubalina 

Diplochita florida DC. =Miconia mirabilis 

Diplochita fothergilla (Desr.) DC. =Miconia mirabilis 

Diplochita leucocephala DC. =Miconia serrulata 

Diplochita mucronata (Desr.) DC. =Miconia holosericea 

Diplochita parviflora Benth. =Miconia pubipetala 

Diplochita serrulata DC. =Miconia serrulata 

Diplochita serrulata var. latifolia DC. =Miconia serrulata 

Diplochita tomentosa (Rich.) Griseb. =Miconia tomentosa 

Ernestia lata Gleason =Ernestia tenella var. sprucei 

Eurychaenia punctata (Desr.) Griseb. =Miconia punctata 

Farringtonia fasciculata Gleason =Siphanthera fasciculata 

Fothergilla guianensis Aubl. =Miconia mirabilis 

Fothergilla mirabilis Aubl. =Miconia mirabilis 

Glossocentrum collinum Crueg. =Miconia minutiflora 

Graffenrieda candelabrum J.F. Macbr. =Leandra candelabrum 

Graffenrieda curta Gleason =Miconia curta 

Graffenrieda floribunda Triana =Graffenrieda miconioides 

Graffenrieda hitchcockii var. parvifolia Wurdack =Graffenrieda hitchcockii 

Graffenrieda ovalifolia Naudin =Graffenrieda weddellii 

Graffenrieda stenopetala Ule =Graffenrieda intermedia 

Guildingia psidioides Hook. =Mouriri rhizophorifolia 

Hartigia barbigera (DC.) Miq. =Miconia ciliata 

Hartigia oblongifolia Miq. =Miconia racemosa 

Hartigia spectabilis Miq. =Miconia racemosa 

Henriettea brunnescens Standl. =Loreya mespiloides 

Henriettea oblongifolia Naudin =Henriettea martiusii 

Henriettea orinocensis Naudin =Henriettea martiusii 

Henriettea succosa var. guianensis Gleason =Henriettea ramiflora 

Henriettea succosa var. kappleriana Steud. ex Naudin =Henriettea succosa 

Henriettea surinamensis Miq. =Henriettea ramiflora 

Henriettea trinervia Naudin =Henriettea ramiflora 

Henriettella longistyla Ule =Henriettella ovata 

Henriettella micrantha Gleason =Henriettella ovata 

Henriettella patrisiana (DC.) Naudin =Henriettea patrisiana 

Henriettella ramiflora (Sw.) Naudin =Henriettea ramiflora 

Henriettella venosa Gleason =Clidemia venosa 

Heterotrichum novemnervium DC. =Clidemia novemnervia 

Heterotrichum strigosum Triana =Miconia strigosa 

Hormocalyx hirsutus Gleason =Tococa macrosperma 

Jucunda tomentosa (Rich.) Benth. =Miconia tomentosa 

Lasiandra bipenicillata Naudin =Tibouchina bipenicillata 

Lasiandra tibouchina (Desr.) Naudin =Tibouchina aspera var. aspera 

Leandra anisophylla (DC.) Cogn. =Clidemia anisophylla 

Leandra chimantensis Wurdack =Leandra procumbens 

Leandra cuspidata (Triana) Cogn. =Miconia eugenioides 

Leandra fendleri Cogn. =Leandra lindeniana 

Leandra hylophila Gleason =Leandra clidemioides 

Leandra linearis Gleason =Clidemia linearis 

Leandra longicoma Cogn. [not in flora area]

410 


Leandra montana Gleason =Aciotis indecora 

Leandra pulverulenta (DC.) Cogn. =Miconia aliquantula 

Leandra rhamnifolia var. macrodon (Naudin) Cogn. =Leandra divaricata 

Leandra rufescens var. asperiuscula (DC.) Cogn. =Leandra rufescens 

Leandra rufescens (DC.) Cogn. var. glandulosa Cogn. =Leandra rufescens 

Leandra scabra DC. [not in flora area] =Leandra melastomoides Raddi 

Leandra stellipilis Gleason =Clidemia stellipilis 

Leiostegia vernicosa Benth. =Comolia vernicosa 

Loevigia sericea H. Karst. & Triana =Monochaetum bonplandii 

Loreya acutifolia O. Berg ex Triana =Loreya arborescens 

Loreya brunnescens (Standl.) Gleason =Loreya mespiloides 

Loreya collatata Wurdack =Loreya spruceana 

Loreya flavescens (Aubl.) Naudin =Henriettella flavescens 

Loreya huberi Wurdack =Bellucia huberi 

Loreya maguirei Wurdack =Loreya arborescens 

Loreya minor Cogn. =Loreya ovata 

Loreya mucronata Gleason =Henriettea mucronata 

Loreya quadrifolia Gleason =Loreya strigosa 

Macairea albiflora Cogn. =Macairea thyrsiflora 

Macairea aspera N.E. Br. =Macairea thyrsiflora 

Macairea calvescens Naudin =Macairea pachyphylla 

Macairea lanata subsp. eglandulosa Wurdack =Macairea lanata 

Macairea maguirei Wurdack =Macairea thyrsiflora 

Macairea schultesii Wurdack =Macairea thyrsiflora 

Macrocentrum glandulosum Gleason =Salpinga glandulosa 

Macrocentrum montanum Gleason =Macrocentrum cristatum var. microphyllum 

Macrocentrum pusillum Gleason =Salpinga pusilla 

Maieta dispar Miq. =Maieta guianensis 

Maieta glandulifera Standl. =Tococa guianensis 

Maieta juruensis Pilg. =Clidemia juruensis 

Marcetia andicola Naudin =Marcetia taxifolia 

Marcetia cordigera var. andicola (Naudin) Cogn. =Marcetia taxifolia 

Marcetia juniperina (Humb.) DC. =Marcetia taxifolia 

Meisneria cordifolia Benth. =Siphanthera cordifolia 

Meisneria microlicioides Naudin =Siphanthera cordifolia 

Melastoma acinodendron L. =Miconia acinodendron 

Melastoma adscendens Sw. =Adelobotrys adscendens 

Melastoma agrestis Aubl. =Leandra agrestis 

Melastoma alatum Aubl. =Miconia alata 

Melastoma albicans Sw. =Miconia albicans 

Melastoma aplostachyum Bonpl. =Miconia aplostachya 

Melastoma aquaticum Aubl. =Nepsera aquatica 

Melastoma arborescens Aubl. =Loreya arborescens 

Melastoma aromaticum Vahl =Tibouchina aspera var. aspera 

Melastoma biglomeratum Bonpl. =Miconia biglomerata 

Melastoma bivalve Aubl. =Acisanthera bivalvis 

Melastoma bivalvis Aubl. =Acisanthera bivalvis 

Melastoma brunnea Vahl =Henriettea succosa 

Melastoma cacatin Aubl. =Miconia cacatin 

Melastoma capitellatum Bonpl. =Clidemia capitellata 

Melastoma chrysophyllum Rich. =Miconia chrysophylla 

Melastoma ciliatum Rich. =Miconia ciliata 

Melastoma dichotomum Desr. =Rhynchanthera dichotoma 

Melastoma diffusum Desr. =Acisanthera uniflora 

Melastoma dodecandrum Desr. =Miconia dodecandra 

Melastoma elatum Sw. =Miconia elata 

Melastoma elegans Aubl. =Clidemia hirta var. elegans 

Melastoma flavescens Aubl. =Henriettella flavescens 

Melastoma fothergillum Desr. =Miconia mirabilis 

Melastoma fulvum Rich. =Miconia chrysophylla 

Melastoma grandiflorum Aubl. =Rhynchanthera grandiflora 

Melastoma grossularioides L. =Bellucia grossularioides 

Melastoma heteroneurum DC. =Clidemia heteroneura 

Melastoma hirtum L. =Clidemia hirta


Melastoma holosericea L. =Miconia holosericea 

Melastoma ibaguense Bonpl. =Miconia ibaguensis 

Melastoma impetiolaris Sw. =Miconia impetiolaris 

Melastoma laevigatum L. =Miconia laevigata 

Melastoma laevigatum Aubl. [nom. illeg., non M. laevigata L.] 

Melastoma longifolium Aubl. =Miconia longifolia 

Melastoma maietum Desr. =Maieta guianensis 

Melastoma micranthum Sw. =Ossaea micrantha 

Melastoma minutiflorum Bonpl. =Miconia minutiflora 

Melastoma mucronatum Desr. =Miconia holosericea 

Melastoma nervosum Sm. =Miconia nervosa 

Melastoma octonum Bonpl. =Clidemia octona 

Melastoma parviflorum Aubl. =Miconia prasina 

Melastoma physophorum Vahl =Tococa guianensis 

Melastoma pleurocarpum E. Mey. =Miconia acinodendron 

Melastoma prasinum Sw. =Miconia prasina 

Melastoma punctatum Desr. =Miconia punctata 

Melastoma purpurascens Aubl. =Aciotis purpurascens 

Melastoma purpurea (Aubl.) Willd. =Aciotis purpurascens 

Melastoma racemosum Aubl. =Miconia racemosa 

Melastoma ramiflorum Sw. =Henriettea ramiflora 

Melastoma rubiginosum Bonpl. =Miconia rubiginosa 

Melastoma rubrum Aubl. =Clidemia rubra 

Melastoma rufescens Aubl. =Miconia rufescens 

Melastoma scandens Aubl. =Adelobotrys scandens 

Melastoma scorpioides Schltdl. & Cham. =Miconia trinervia 

Melastoma spicatum Aubl. =Clidemia capitellata var. dependens 

Melastoma splendens Sw. =Miconia splendens 

Melastoma strigillosum Sw. =Clidemia strigillosa 

Melastoma succosum Aubl. =Henriettea succosa 

Melastoma tamonea Sw. =Miconia mirabilis 

Melastoma theizans Bonpl. =Miconia theizans 

Melastoma tibouchinum Desr. =Tibouchina aspera var. aspera 

Melastoma tococo Desr. =Tococa guianensis 

Melastoma tomentosum Rich. =Miconia tomentosa 

Melastoma trinervium Sw. =Miconia trinervia 

Melastoma trivalvis Aubl. =Acisanthera bivalvis 

Melastoma villosum Aubl. =Desmoscelis villosa 

Meliandra monadelpha (Ducke) Morley =Votomita monadelpha 

Meriania duidae Gleason =Adelobotrys duidae 

Meriania ferruginea Suess. =Adelobotrys barbata 

Meriania paraensis Ducke =Meriania urceolata 

Miconia agrestis (Aubl.) Baill. =Leandra agrestis 

Miconia amplexans (Crueg.) Cogn. =Miconia tomentosa 

Miconia anceps Naudin =Miconia trinervia 

Miconia argyrophylla var. attenuata Cogn. =Miconia argyrophylla subsp. argyrophylla 

Miconia arirambae Huber =Miconia alborufescens 

Miconia attenuata DC. =Miconia prasina 

Miconia attenuata var. subquintuplinervia DC. =Miconia prasina 

Miconia axilliflora Naudin =Miconia acuminata 

Miconia barbigera DC. =Miconia ciliata 

Miconia berbiceana (DC.) Naudin =Miconia acinodendron 

Miconia bifrons Naudin =Miconia lepidota 

Miconia bracteolaris Naudin =Miconia eriodonta 

Miconia bracteolaris var. cardiophora Naudin =Miconia eriodonta 

Miconia buchtienii Cogn. =Miconia centrodesma 

Miconia cachimbensis Brade =Miconia alborufescens 

Miconia caryophyllaea (DC.) Triana =Tococa caryophyllea 

Miconia caudiculata Pittier =Miconia trinervia 

Miconia centrandra Ule =Miconia centrodesma 

Miconia ceramicarpa var. navioensis Wurdack =Miconia ceramicarpa var. candolleana 

Miconia ceramicarpa var. violacea (DC.) Cogn. =Miconia ceramicarpa var. ceramicarpa 

Miconia chamissonis Naudin [not in flora area] 

Miconia clidemioides Steud. =Miconia acinodendron

412 


Miconia compacta Gleason =Miconia carassana 

Miconia congesta Cogn. =Miconia poeppigii 

Miconia cristulata Naudin =Miconia prasina 

Miconia cruegeriana Naudin =Miconia hypoleuca 

Miconia demerarensis Gleason =Miconia bracteata 

Miconia disparilis (Standl.) R.O. Williams =Miconia lateriflora subsp. lateriflora 

Miconia eriodonta var. oblongifolia DC. =Miconia eriodonta 

Miconia erioneura Cogn. =Tococa guianensis 

Miconia eriophora Miq. =Miconia ibaguensis 

Miconia eriopoda Benth. =Miconia acinodendron 

Miconia erythrophylla Ule =Tococa erythrophylla 

Miconia erythropila Steud. =Miconia ceramicarpa var. ceramicarpa 

Miconia fendleriana Cogn. =Miconia phaeophylla 

Miconia florida (DC.) Naudin =Miconia mirabilis 

Miconia fockeana Miq. =Miconia nervosa 

Miconia fothergilla (Desr.) Naudin =Miconia mirabilis 

Miconia fulva (Rich.) DC. =Miconia chrysophylla 

Miconia fulva var. angustifolia Cogn. =Miconia chrysophylla 

Miconia fulva var. aubletiana (Naudin) Cogn. =Miconia chrysophylla 

Miconia globuliflora (Rich.) Cham. [not in flora area] 

Miconia glossocentra Naudin =Miconia minutiflora 

Miconia guaiquinimae subsp. angustifolia Wurdack =Miconia guaiquinimae 

Miconia guianensis (Aubl.) Cogn. =Miconia mirabilis 

Miconia heterochroa Miq. =Miconia albicans 

Miconia holosericea var. mucronata (Desr.) Cogn. =Miconia holosericea 

Miconia holosericea var. subquintuplinervia Cogn. =Miconia holosericea 

Miconia hypargyrea Miq. =Miconia stenostachya 

Miconia ibaguensis var. glabrata (Steud.) Cogn. =Miconia ibaguensis 

Miconia impetiginosa DC. =Miconia rufescens 

Miconia involucrata Donn. Sm. =Miconia ampla 

Miconia iodopila Steud. =Miconia ceramicarpa var. ceramicarpa 

Miconia killipi Gleason =Miconia loreyoides 

Miconia lambertiana DC. =Miconia longifolia 

Miconia leschenaultiana (Naudin) Sagot =Miconia ceramicarpa var. ceramicarpa 

Miconia leucocephala (DC.) Naudin =Miconia serrulata 

Miconia longifolia var. aubletiana Naudin =Miconia chrysophylla 

Miconia longistyla Steud. =Miconia argyrophylla subsp. argyrophylla 

Miconia macrophylla Steud. =Miconia prasina 

Miconia macrophylla var. hostmannii (Steud.) Cogn. =Miconia serrulata 

Miconia macrophylla var. latifolia (DC.) Cogn. =Miconia serrulata 

Miconia macrophylla var. leucocephala (DC.) Cogn. =Miconia serrulata 

Miconia macrophylla var. serrulata (DC.) Cogn. =Miconia serrulata 

Miconia macrostachya DC. =Miconia aplostachya 

Miconia megaphylla Gleason =Miconia ampla 

Miconia micrantha Pilg. =Miconia wittii 

Miconia microcarpa DC. =Miconia affinis 

Miconia miqueliana Naudin =Miconia acinodendron 

Miconia mucronata (Desr.) Naudin =Miconia holosericea 

Miconia mucronulata Ule =Miconia prasina 

Miconia multinervulosa Cogn. =Miconia theizans 

Miconia naudiniana Miq. =Miconia lepidota 

Miconia nemoralis Naudin =Miconia prasina 

Miconia nitens Benth. =Tococa nitens 

Miconia obovalis Naudin =Miconia splendens 

Miconia pachypoda Gleason =Miconia superba 

Miconia pallida Gleason =Miconia elaeoides 

Miconia palmetorum Pittier =Miconia stephananthera 

Miconia panicularis Gleason =Miconia egensis 

Miconia parviflora (Benth.) Cogn. =Miconia pubipetala 

Miconia planinervia Naudin =Miconia affinis 

Miconia plebeia Naudin =Miconia brevipes 

Miconia plumosa Gleason =Miconia rugosa 

Miconia prasina var. attenuata (DC.) Cogn. =Miconia prasina 

Miconia pseudorubiginosa Cogn. =Miconia macrothyrsa


Miconia pterophora Miq. =Miconia alata 

Miconia pteropoda Benth. =Miconia prasina 

Miconia purpurascens (Aubl.) DC. =Aciotis purpurascens 

Miconia racemosa var. barbigera (DC.) Cogn. =Miconia ciliata 

Miconia racemosa var. ciliata (Rich.) Griseb. =Miconia ciliata 

Miconia revoluta Benth. =Miconia prasina 

Miconia robusta (Sagot) Cogn. =Miconia tschudyoides 

Miconia rubiginosa var. kuhlmannli Hoehne =Miconia brevipes 

Miconia rubiginosa var. platyura Naudin =Miconia rubiginosa 

Miconia rufescens var. grandifolia Cogn. =Miconia rufescens 

Miconia schomburgkii Benth. =Miconia lepidota 

Miconia scorpioides (Schltdl. & Cham.) Naudin =Miconia trinervia 

Miconia scrobiculata Cogn. =Miconia radulaefolia 

Miconia semota Markgr. =Miconia carassana 

Miconia septuplinervis Pittier =Miconia nervosa 

Miconia setimarginata Pittier =Tococa oligantha 

Miconia spondylantha DC. =Miconia nervosa 

Miconia spondylantha var. leschenaultiana Naudin =Miconia ceramicarpa var. ceramicarpa 

Miconia subciliata DC. =Tococa subciliata 

Miconia subtriloba Gleason =Miconia centrodesma 

Miconia surinamensis Gleason =Miconia poeppigii 

Miconia sylvestris Gleason =Miconia centrodesma 

Miconia tessmannii Markgr. =Miconia lateriflora subsp. lateriflora 

Miconia tinifolia var. roraimensis Wurdack =Miconia tinifolia 

Miconia tomentella Cogn. =Miconia serialis 

Miconia trichodes (DC.) Triana =Miconia bracteata 

Miconia trinitatis (Crueg.) Naudin =Miconia minutiflora 

Miconia verticilliflora Steud. ex Naudin =Miconia pubipetala 

Miconia virgulata Gleason =Miconia ciliata 

Microlicia bivalvis (Aubl.) DC. =Acisanthera bivalvis 

Microlicia brevifolia DC. =Acisanthera bivalvis 

Microlicia bryanthoides Oliv. =Microlicia benthamiana 

Microlicia limnobios DC. =Acisanthera limnobios 

Microlicia recurva (Rich.) DC. =Acisanthera uniflora 

Microlicia steyermarkii Gleason =Microlicia benthamiana 

Microphysca rotundifolia Triana =Tococa rotundifolia 

Miocarpus paludosus Naudin =Acisanthera limnobios 

Monochaetum sericeum Naudin ex Linden =Monochaetum bonplandii 

Mouriri abnormis Naudin =Votomita guianensis 

Mouriri acutiflora var. amblyodon Naudin =Mouriri acutiflora 

Mouriri acutiflora var. oligantha Gleason =Mouriri collocarpa 

Mouriri anomala Pulle =Mouriri crassifolia 

Mouriri pendulifolia Cogn. =Mouriri acutiflora 

Mouriri plasschaerti Pulle =Mouriri nigra 

Mouriri plasschaertii Pulle =Mouriri nigra 

Mouriri polyantha Miq. =Mouriri guianensis 

Mouriri princeps Naudin =Mouriri grandiflora 

Mouriri ulei Pilg. =Mouriri guianensis 

Mouriri weddellii Naudin =Mouriri guianensis 

Myriaspora decipiens Naudin =Myriaspora egensis 

Myriaspora surinamensis Steud. =Myriaspora egensis 

Myrmidone lanceolata Cogn. =Tococa macrosperma 

Myrmidone macrosperma (Mart.) Mart. =Tococa macrosperma 

Noterophila beccabunga Mart. ex Naudin =Acisanthera bivalvis 

Noterophila brevifolia (DC.) Naudin =Acisanthera bivalvis 

Noterophila limnobios (DC.) Mart. =Acisanthera limnobios 

Noterophila pusilla Naudin =Acisanthera limnobios 

Octopleura ciliata Triana =Miconia lateriflora subsp. lateriflora 

Octopleura micrantha (Sw.) Griseb. =Ossaea micrantha 

Olisbea rhizophorifolia DC. =Mouriri rhizophorifolia 

Onoctonia calcarata Naudin =Acisanthera crassipes 

Onoctonia crassipes Naudin =Acisanthera crassipes 

Onoctonia pauciflora Naudin =Acisanthera crassipes 

Osbeckia aubletiana Spreng. =Rhynchanthera grandiflora

414 


Osbeckia glomerata (Rottb.) DC. =Pterolepis glomerata 

Osbeckia striphnocalyx DC. =Tibouchina striphnocalyx 

Ossaea ciliata (Triana) Cogn. =Miconia lateriflora subsp. lateriflora 

Ossaea disparilis Standl. =Miconia lateriflora subsp. lateriflora 

Ossaea duckeana Hoehne =Clidemia epibaterium 

Ossaea epibaterium (DC.) Triana =Clidemia epibaterium 

Ossaea flavescens (Aubl.) DC. =Henriettella flavescens 

Ossaea trichopoda Gleason =Leandra chaetodon 

Ossaea trinitensis Crueg. =Clidemia trinitensis 

Oxymeris agrestis (Aubl.) Triana =Leandra agrestis 

Oxymeris anisophylla (DC.) Triana =Clidemia anisophylla 

Oxymeris aristigera (Naudin) Triana =Leandra aristigera 

Oxymeris asperiuscula (DC.) Triana =Leandra rufescens 

Oxymeris chaetodon (DC.) Triana =Leandra chaetodon 

Oxymeris cuspidata Triana =Miconia eugenioides 

Oxymeris glandulifera Triana =Leandra glandulifera 

Oxymeris lindeniana (Naudin) Triana =Leandra lindeniana 

Oxymeris pulverulenta (DC.) Triana =Miconia aliquantula 

Oxymeris rufescens (DC.) Triana =Leandra rufescens 

Oxymeris secunda (D. Don) Triana =Leandra secunda 

Pachyloma nanum Wurdack =Pachyloma pusillum 

Pachyloma scandens Ducke =Pachyloma huberioides 

Petaloma mouriri Sw. =Mouriri guianensis 

Petaloma myrtilloides Sw. =Mouriri myrtilloides 

Phyllopus martiusii DC. =Henriettea martiusii 

Platycentrum clidemioides Naudin =Leandra clidemioides 

Pleroma bipenicillata (Naudin) Triana =Tibouchina bipenicillata 

Pleroma tibouchinum (Desr.) Triana =Tibouchina aspera var. aspera 

Pogonorhynchus amplexans Crueg. =Miconia tomentosa 

Poteranthera calcarata (Naudin) Triana =Acisanthera crassipes 

Poteranthera crassipes (Naudin) Triana =Acisanthera crassipes 

Poteranthera duidae Gleason =Siphanthera duidae 

Poteranthera foliosa (Naudin) Cogn. =Siphanthera foliosa 

Poteranthera minor Gleason =Siphanthera foliosa 

Poteranthera pauciflora (Naudin) Triana =Acisanthera crassipes 

Pterogastra glabra Gleason =Pterogastra divaricata 

Pterogastra major Triana =Pterogastra divaricata 

Pterolepis hispida (Rich.) Gleason =Pterolepis trichotoma 

Pterolepis hostmannii Steud. =Pterolepis glomerata 

Pterolepis ladanoides (Rich.) Triana =Pterolepis trichotoma 

Pterolepis lasiophylla (Benth.) Triana =Macairea lasiophylla 

Pterolepis pumila (Bonpl.) Cogn. =Pterolepis trichotoma 

Pterolepis pumila var. procera Cogn. =Pterolepis trichotoma 

Pterolepis pusilla Miq. =Pterolepis trichotoma 

Pterolepis striphnocalyx (DC.) Cogn. =Tibouchina striphnocalyx 

Rhexia acisanthera L. =Acisanthera quadrata 

Rhexia angusturensis Rich. =Pterolepis glomerata 

Rhexia aquatica (Aubl.) Sw. =Nepsera aquatica 

Rhexia aspera (Aubl.) Willd. =Tibouchina aspera var. aspera 

Rhexia bicornis Rottb. =Acisanthera uniflora 

Rhexia bivalvis (Aubl.) Vahl =Acisanthera bivalvis 

Rhexia bonplandii Kunth =Monochaetum bonplandii 

Rhexia canescens Bonpl. =Monochaetum bonplandii 

Rhexia divaricata Bonpl. =Pterogastra divaricata 

Rhexia glomerata Rottb. =Pterolepis glomerata 

Rhexia gracilis Bonpl. =Tibouchina gracilis 

Rhexia hispida Rich. =Pterolepis trichotoma 

Rhexia hypericoides Willd. =Desmoscelis villosa 

Rhexia incana Spreng. =Monochaetum bonplandii 

Rhexia indecora Bonpl. =Aciotis indecora 

Rhexia juniperina Humb. =Marcetia taxifolia 

Rhexia ladanoides Rich. =Pterolepis trichotoma 

Rhexia latifolia Aubl. =Comolia latifolia 

Rhexia leptophylla Bonpl. =Comolia leptophylla


Rhexia longifolia Vahl =Tibouchina longifolia 

Rhexia nummularioides Bonpl. =Comolia nummularioides 

Rhexia polystachya Bonpl. =Aciotis polystachya 

Rhexia pumila Bonpl. =Pterolepis trichotoma 

Rhexia recurva Rich. =Acisanthera uniflora 

Rhexia rotundifolia Bonpl. =Graffenrieda rotundifolia 

Rhexia serrulata Rich. =Rhynchanthera serrulata 

Rhexia sileniflora Bonpl. =Aciotis purpurascens 

Rhexia strigosa Willd. ex Spreng. =Pterolepis trichotoma 

Rhexia taxifolia A. St.-Hil. =Marcetia taxifolia 

Rhexia tenella Bonpl. =Ernestia tenella 

Rhexia thymifolia Bonpl. =Appendicularia thymifolia 

Rhexia trichotoma Rottb. =Pterolepis trichotoma 

Rhexia trivalvis (Aubl.) Vahl =Acisanthera bivalvis 

Rhexia uniflora Vahl =Acisanthera uniflora 

Rhexia villosa Aubl. =Comolia villosa 

Rhexia villosissima Rich. =Desmoscelis villosa 

Rhynchanthera acuminata Benth. =Rhynchanthera grandiflora 

Rhynchanthera acuminata var. sublaevis Cogn. =Rhynchanthera grandiflora 

Rhynchanthera adenophora Miq. =Acisanthera uniflora 

Rhynchanthera ambigua Naudin =Rhynchanthera grandiflora 

Rhynchanthera betulifolia Cogn. =Rhynchanthera grandiflora 

Rhynchanthera cardonae Wurdack =Rhynchanthera grandiflora 

Rhynchanthera grandiflora var. microphylla Naudin =Rhynchanthera grandiflora 

Rhynchanthera intermedia Ule =Rhynchanthera grandiflora 

Rhynchanthera latifolia Cogn. [not in flora area] 

Rhynchanthera microphylla Gleason =Rhynchanthera grandiflora 

Rhynchanthera modesta Naudin =Rhynchanthera serrulata 

Rhynchanthera orinocensis Sprague =Rhynchanthera grandiflora 

Rhynchanthera parviflora Naudin =Rhynchanthera serrulata 

Rhynchanthera virgata Wurdack =Rhynchanthera hispida 

Sagraea aphanantha (Naudin) Triana =Clidemia aphanantha 

Sagraea capillipes Triana =Clidemia capillipes 

Sagraea capitata (Benth.) Triana =Clidemia capitata 

Sagraea cognata Steud. =Clidemia rubra 

Sagraea coriacea (Naudin) Triana =Clidemia tepuiensis 

Sagraea minutiflora Triana =Clidemia minutiflora 

Sagraea neurocarpa Naudin =Ossaea micrantha 

Sagraea petiolata (DC.) Triana =Clidemia petiolata 

Sagraea rubra (Aubl.) Triana =Clidemia rubra 

Salpinga cristata DC. =Macrocentrum cristatum 

Salpinga fasciculata DC. =Macrocentrum fasciculatum 

Salpinga parviflora DC. =Macrocentrum cristatum var. parviflorum 

Sandemania glandulosa Wurdack =Sandemania hoehnei 

Sarmentaria decora Naudin =Adelobotrys adscendens 

Siphanthera capitata Gleason =Siphanthera hostmannii 

Siphanthera cordifolia var. glomerata Gleason =Siphanthera cordifolia 

Siphanthera jenmanii Gleason =Siphanthera cordifolia 

Siphanthera microlicioides (Naudin) Cogn. =Siphanthera cordifolia 

Siphanthera paraensis Huber =Siphanthera hostmannii 

Siphanthera tatei Gleason =Siphanthera hostmannii 

Siphantheropsis williamii Brade =Macairea lasiophylla 

Spennera acuminifolia Mart. ex DC. =Aciotis acuminifolia 

Spennera acutiflora Mart. =Aciotis indecora 

Spennera alata Beurl. =Aciotis rubricaulis 

Spennera annua Mart. ex DC. =Aciotis annua 

Spennera anomala Miq. =Aciotis acuminifolia 

Spennera aquatica (Aubl.) Mart. ex DC. =Nepsera aquatica 

Spennera caulialata (Ruiz & Pav.) Naudin =Aciotis rubricaulis 

Spennera chaetodon DC. =Leandra chaetodon 

Spennera dichotoma Benth. =Aciotis acuminifolia 

Spennera dichotoma var. lanceolata Naudin =Aciotis acuminifolia 

Spennera dysophylla Benth. =Aciotis annua 

Spennera fragilis Rich. ex DC. =Aciotis purpurascens

416 


Spennera grandifolia Miq. =Aciotis purpurascens 

Spennera hydrophila Miq. =Nepsera aquatica 

Spennera indecora (Bonpl.) DC. =Aciotis indecora 

Spennera kappleriana Naudin =Aciotis indecora 

Spennera lancifolia Steud. ex Cogn. =Aciotis purpurascens 

Spennera laxa DC. =Aciotis indecora 

Spennera longifolia DC. =Aciotis circaeifolia 

Spennera martinicensis Naudin =Aciotis purpurascens 

Spennera ornata Miq. =Aciotis ornata 

Spennera pellucida Rich. ex DC. =Aciotis purpurascens 

Spennera polystachya (Bonpl.) Mart. ex DC. =Aciotis polystachya 

Spennera quintuplinervia Miq. =Clidemia japurensis var. japurensis 

Spennera rostellata Naudin =Aciotis indecora 

Spennera rubricaulis Schrank & Mart. ex DC. =Aciotis rubricaulis 

Spennera sieberi Steud. =Aciotis purpurascens 

Spennera sileniflora (Bonpl.) DC. =Aciotis purpurascens 

Spennera sphaeranthera Naudin =Aciotis acuminifolia 

Spennera tetraptera Miq. =Aciotis annua 

Spennera uliginosa Bonpl. ex Naudin =Aciotis purpurascens 

Spennera viscida Benth. =Aciotis viscida 

Spennera viscosa Bonpl. ex Naudin =Aciotis indecora 

Staphidiastrum aphananthum Naudin =Clidemia aphanantha 

Staphidiastrum attenuatum Naudin =Clidemia attenuata 

Staphidiastrum cognatum (Steud.) Naudin =Clidemia rubra 

Staphidiastrum coriaceum Naudin =Clidemia tepuiensis 

Staphidiastrum pulverulentum (DC.) Naudin =Miconia aliquantula 

Staphidiastrum rubrum (Aubl.) Naudin =Clidemia rubra 

Staphidiastrum rubrum var. biacutum Naudin =Clidemia sericea 

Staphidium affine Naudin =Clidemia pustulata 

Staphidium agreste (Aubl.) Naudin =Leandra agrestis 

Staphidium benthamianum (Miq.) Naudin =Clidemia hirta var. hirta 

Staphidium bracteosum Naudin =Clidemia capitellata var. dependens 

Staphidium conglomeratum (DC.) Naudin =Clidemia conglomerata 

Staphidium elegans (Aubl.) Naudin =Clidemia hirta var. elegans 

Staphidium glabratum (Steud.) Naudin =Miconia ibaguensis 

Staphidium hostmannii Naudin =Clidemia dentata 

Staphidium involucratum (DC.) Naudin =Clidemia involucrata 

Staphidium lappaceum (DC.) Naudin =Miconia lappacea 

Staphidium mutabile (DC.) Naudin =Miconia mutabilis 

Staphidium petiolatum (DC.) Naudin =Clidemia petiolata 

Staphidium sessiliflorum Naudin =Clidemia sessiliflora 

Staphidium spicatum (Aubl.) Naudin =Clidemia capitellata var. dependens 

Staphidium triste Naudin =Miconia bracteata 

Tamonea guianensis Aubl. =Miconia mirabilis 

Tetrameris trivalvis Naudin =Macairea multinervia 

Tetrameris villosa (Aubl.) Naudin =Comolia villosa 

Tetrazygia discolor (L.) DC. [not in flora area] 

Tibouchina adscendens (C. Wright) M. Gómez =Acisanthera quadrata 

Tibouchina brachyanthera Pittier =Tibouchina aspera var. asperrima 

Tibouchina impressa Gleason =Tibouchina geitneriana 

Tibouchina lasiophylla (Benth.) Cogn. =Macairea lasiophylla 

Tibouchina multinervia (Benth.) Baill. =Macairea multinervia 

Tibouchina pseudomollis Gleason =Tibouchina geitneriana 

Tibouchina yavitensis Pittier =Tibouchina striphnocalyx 

Tococa acuminata Benth. =Tococa guianensis 

Tococa aubletii D. Don =Tococa guianensis 

Tococa cardiophylla Naudin =Tococa guianensis 

Tococa castrata Naudin =Tococa coronata 

Tococa coriaceae S. Moore =Tococa guianensis 

Tococa didymophysca Naudin =Tococa guianensis 

Tococa discolor Pilg. =Tococa guianensis 

Tococa egensis Naudin =Tococa coronata 

Tococa erioneura (Cogn.) Wurdack =Tococa guianensis


Tococa erythrostachya Spruce ex Triana =Tococa guianensis 

Tococa ferruginea G. Nicholson =Tococa guianensis 

Tococa glandulosa Gleason =Tococa coronata 

Tococa juruensis Pilg. =Tococa guianensis 

Tococa lasiostyla Cogn. =Tococa subciliata 

Tococa longisepala Cogn. =Tococa coronata 

Tococa loretensis Ule =Tococa guianensis 

Tococa maieta D. Don =Maieta guianensis 

Tococa montana Gleason =Tococa obovata subsp. obovata 

Tococa occidentalis Naudin =Tococa guianensis 

Tococa planifolia Benth. =Tococa subciliata 

Tococa rareimi Benth. [‘Roraimae’] =Tococa guianensis 

Tococa spruceana Cogn. =Tococa bullifera 

Tococa subnuda Benth. =Tococa coronata 

Tococa traillii Cogn. =Tococa coronata 

Tococa ulei Pilg. =Tococa bullifera 

Tococa wedellii Naudin =Tococa nitens 

Topobea “guianensis Aubl.” [invalid name from herbarium records, = Topobea parasitica] 

Topobea cuspidata Gleason =Topobea parasitica 

Tricentrum leptophyllum (Bonpl.) DC. =Comolia leptophylla 

Truncaria caryophyllea DC. =Tococa caryophyllea 

Tschudya asperiuscula DC. =Leandra rufescens 

Tschudya berbiceana (DC.) Griseb. =Miconia acinodendron 

Tschudya ibaguensis (Bonpl.) Griseb. =Miconia ibaguensis 

Tschudya pulverulenta DC. =Miconia aliquantula 

Tschudya robusta Sagot =Miconia tschudyoides 

Tschudya rufescens DC. =Leandra rufescens 

Tschudya spondylantha (DC.) Griseb. =Miconia nervosa 

Tschudya strigillosa Griseb. =Leandra clidemioides 

Tschudya verticillata DC. =Leandra verticillata 

Tulasnea foliosa Naudin =Siphanthera foliosa 

Uranthera dicranophora Naudin =Acisanthera quadrata 

Uranthera hostmannii Naudin =Acisanthera uniflora 

Uranthera recurva (Rich.) Naudin =Acisanthera uniflora 

Urodesmium huberioides Naudin =Pachyloma huberioides 

Valdesia rosea Ruiz & Pav. =Blakea rosea 

Webera quinquenervia (Aubl.) J.F. Gmel. =Bellucia grossularioides 

REFERENCES 

(for a more extensive bibliography, see Wurdack et al. 1993) 

Berry, P.E., A. Gröger, B.K. Holst, T. Morley, F.A. Michelangeli, N.G. Luckana, F. Almeda, S.S. Renner, A. Freire-Fierro, O.R. 

Robinson, & K. Yatskievych. 2001. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 263-528. 

Freire-Fierro, A. 2002. Monograph of Aciotis (Melastomataceae). Systematic Botany Monographs 62: 1-99. 

Gleason, H.A. 1932. A synopsis of the Melastomataceae of British Guiana. Brittonia 1: 127-184. 

Gleason, H.A. 1935. Melastomaceae. In A. Pulle, Flora of Suriname 3(1): 178-281. 

Görts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 1986. Melastomataceae (additions and corrections). In A.L. Stoffers & J.C. 

Lindeman, Flora of Suriname 3(2): 356-429. 

Morley, T. 1976. Memecyleae (Melastomataceae). Flora Neotropica Monogr. 15: 1-295. 

Renner, S.S. 1989. Systematic studies in the Melastomataceae: Bellucia, Loreya, and Macairea. Mem. New York Bot. Gard. 50: 

1-112. 

Wurdack, J.J. 1961. Melastomataceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. 


Wurdack, J.J. 1973. Melastomataceae. In T. Lasser, Flora de Venezuela 8: 1-819. (Mouriri & Votomita contributed by T. Morley) 

Wurdack, J.J. & S. Renner. 2002. Melastomataceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, 

& J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York 

Bot. Gard. 76 (2): 437-465. 

Wurdack, J.J., T. Morley, & S. Renner. 1993. Melastomataceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 13: 1- 

424. 

Azadirachta indica A. Juss. [cultivated, naturalized] 

MELIACEAE 

K.S. Edwards, T.D. Pennington

418 


Cabralea canjerana (Vell.) Mart. 

Cabralea canjerana subsp. canjerana 

Carapa guianensis Aubl. 

Carapa procera DC. 

Cedrela fissilis Vell. 

Cedrela odorata L. 

Guarea carinata Ducke 

Guarea cinnamomea Harms 

Guarea costata A. Juss. 

Guarea glabra Vahl 

Guarea gomma Pulle 

Guarea grandifolia DC. 

Guarea guidonia (L.) Sleumer 

Guarea humaitensis T.D. Penn. 

Guarea juglandiformis T.D. Penn. 

Guarea kunthiana A. Juss. 

Guarea macrophylla Vahl 

Guarea macrophylla subsp. pachycarpa (C. DC.) T.D. Penn. 

Guarea macrophylla subsp. pendulispica (C. DC.) T.D. Penn. 

Guarea michel-moddei T.D. Penn. & S.A. Mori 

Guarea pubescens (Rich.) A. Juss. 

Guarea pubescens subsp. pubescens 

Guarea pubescens subsp. pubiflora (A. Juss.) T.D. Penn. 

Guarea scabra A. Juss. 

Guarea silvatica C. DC. 

Guarea sprucei C. DC. 

Guarea trunciflora C. DC. 

Guarea velutina A. Juss. 

Melia azedarach L. [cultivated, naturalized] 

Swietenia macrophylla King 

Trichilia acuminata (Humb. & Bonpl. ex Roem. & Schult.) A. DC. & C. DC. 

Trichilia areolata T.D. Penn. 

Trichilia cipo (A. Juss.) C. DC. 

Trichilia elegans A. Juss. 

Trichilia elegans subsp. elegans 

Trichilia euneura C. DC. 

Trichilia gamopetala T.D. Penn. 

Trichilia inaequilatera T.D. Penn. 

Trichilia lecointei Ducke 

Trichilia lepidota Mart. 

Trichilia lepidota subsp. leucastera (Sandwith) T.D. Penn. 

Trichilia martiana C. DC. 

Trichilia mazanensis J.F. Macbr. [expected in the Guianas] 

Trichilia micrantha Benth. 

Trichilia micropetala T.D. Penn. 

Trichilia moschata Sw. 

Trichilia pallida Sw. 

Trichilia pleeana (A. Juss.) C. DC. 

Trichilia quadrijuga Kunth 

Trichilia quadrijuga subsp. quadrijuga 

Trichilia rubra C. DC. 

Trichilia schomburgkii C. DC. 

Trichilia schomburgkii subsp. javariensis T.D. Penn. 

Trichilia schomburgkii subsp. schomburgkii 

Trichilia septentrionalis C. DC. 

Trichilia surinamensis (Miq.) C. DC. 

Trichilia surumuensis C. DC. 

Trichilia tetrapetala C. DC. 

Carapa surinamensis Miq. =Carapa procera 

Cedrela brownii Loefl. ex Kuntze =Cedrela odorata 

Cedrela guianensis A. Juss. =Cedrela odorata 

Granatum surinamensis (Miq.) Kuntze =Carapa procera 

Guarea adenocarpa Pittier =Trichilia pleeana 


GU 

GU 


GU? SU FG 

BO DA 


SU FG 

FG? 

SU FG 


BO GU SU FG 

AM GU SU FG 


GU 

GU 

AM GU SU FG 


AM BO SU? FG 

AM BO GU 

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AM BO SU FG? 

GU SU FG 

AM GU SU FG 

AM 

BO GU 

GU 


FG 

BO 

SU 

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BO GU 

BO GU 

GU? SU FG 

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AM BO 

GU SU 

BO GU SU 

BO GU SU 

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AM DA GU 


AM BO 



AM GU SU FG 

GU 

BO


Guarea affinis A. Juss. =Guarea pubescens subsp. pubescens 

Guarea aubletii A. Juss. =Guarea guidonia 

Guarea concinna Sandwith =Guarea pubescens subsp. pubescens 

Guarea davisii Sandwith =Guarea pubescens subsp. pubescens 

Guarea delgadoi Pittier =Trichilia pleeana 

Guarea guara (Jacq.) P. Wilson =Guarea guidonia 

Guarea pallida C. DC. =Guarea pubescens 

Guarea pedicellata C. DC. =Guarea silvatica 

Guarea pendulispica C. DC. =Guarea macrophylla subsp. pendulispica 

Guarea puberula Pittier =Guarea guidonia 

Guarea pubiflora A. Juss. [not in flora area] =Guarea pubescens (Rich.) A.Juss. subsp. pubiflora 

Guarea richardiana A. Juss. =Guarea pubescens subsp. pubescens 

Guarea schomburgkii C. DC. =Guarea glabra 

Guarea surinamensis Miq. ex C. DC. [nomen nudum] =Guarea guidonia 

Guarea trichilioides L. =Guarea guidonia 

Guarea trichilioides var. pachycarpa C. DC. =Guarea macrophylla subsp. pachycarpa 

Moschoxylum cipo A. Juss. =Trichilia cipo 

Moschoxylum pleeanum A. Juss. =Trichilia pleeana 

Moschoxylum propinquum Miq. =Trichilia quadrijuga 

Moschoxylum surinamense Miq. =Trichilia surinamensis 

Odontandra acuminata Humb. & Bonpl. ex Roem. & Schult. =Trichilia acuminata 

Odontocandra karstenii Triana & Planch. =Trichilia acuminata 

Portesia echinocarpa de Vriese =Trichilia pallida 

Surenus brownii (Loefl. ex Kuntze) Kuntze =Cedrela odorata 

Trichilia acariaeantha Harms =Trichilia micrantha 

Trichilia alternans C. DC. =Trichilia surinamensis 

Trichilia brachystachya Klotzsch ex C. DC. =Trichilia pallida 

Trichilia compacta A.C. Sm. =Trichilia quadrijuga subsp. quadrijuga 

Trichilia cuneifolia Pulle =Trichilia schomburgkii 

Trichilia davisii Sandwith =Trichilia pallida 

Trichilia echinocarpa (de Vriese) Walp. =Trichilia pallida 

Trichilia fuscescens Radlk. =Trichilia martiana 

Trichilia grandis Lasser & Maguire =Trichilia septentrionalis 

Trichilia guianensis Klotzsch ex C. DC. =Trichilia rubra 

Trichilia karstenii (Triana & Planch.) A. DC. & C. DC. =Trichilia acuminata 

Trichilia lanceolata Pittier [nom. illeg.] =Trichilia septentrionalis 

Trichilia leucastera Sandwith =Trichilia lepidota subsp. leucastera 

Trichilia maynasiana C. DC. =Trichilia surinamensis 

Trichilia montana var. fendleriana C. DC. =Trichilia pallida 

Trichilia moritzii C. DC. =Trichilia septentrionalis 

Trichilia poeppigii C. DC. =Trichilia schomburgkii 

Trichilia propinqua (Miq.) C. DC. =Trichilia quadrijuga subsp. quadrijuga 

Trichilia pubescens Rich. =Guarea pubescens 

Trichilia roraimana C. DC. =Trichilia micrantha 

Trichilia smithii C. DC. =Trichilia pleeana 

Trichilia stelligera Radlk. =Trichilia euneura 

Trichilia subsessilifolia C. DC. =Trichilia schomburgkii 

Trichilia subsimplex Steyerm. =Trichilia pallida 

Trichilia trinitensis A. Juss. =Trichilia pallida 

REFERENCES 

Pennington, T.D. 2002. Meliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


465-472. 

Pennington, T.D., B.T. Styles, & D.A.H. Taylor. 1981. Meliaceae. Flora Neotropica Monogr. 28: 1-470. 

Pennington, T.D. & K.S. Edwards. 2001In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 528-549. 

Pennington, T.D. & A.R.A. Görts-van Rijn. 1984. Meliaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 519- 

569. 

MENDONCIACEAE 

D. Wasshausen 

Mendoncia aspera (Ruiz & Pav.) Nees GU SU FG

420 

Mendoncia bivalvis (L. f.) Merr. 

Mendoncia cardonae Leonard 

Mendoncia coriacea Wassh. 

Mendoncia crenata Lindau 

Mendoncia glabra (Poepp. & Endl.) Nees 

Mendoncia hoffmannseggiana Nees 

Mendoncia neblinensis Wassh. 

Mendoncia obovata Lindau 

Mendoncia phalacra Leonard 

Mendoncia sprucei Lindau 

Mendoncia squamuligera Nees 

Mendoncia steyermarkii Wassh. 

Mendoncia williamsii Wassh. 



Mendoncia hirsuta (Poepp. & Endl.) Nees =Mendoncia bivalvis 

Mendoncia perrottetiana Nees =Mendoncia bivalvis 

Mendoncia pubescens Hoffsgg. ex Nees =Mendoncia hoffmannseggiana 

Mendoncia schomburgkiana Nees =Mendoncia hoffmannseggiana 

Mendozia hirsuta Poepp. & Endl. =Mendoncia bivalvis 

REFERENCES 


Wasshausen, D.C. 1989. New species of Mendoncia from the Venezuelan Guayana. Ann. Missouri Bot. Gard. 76: 1118-1124. 

Wasshausen, D.C. 2001. Mendonciaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 550-554. 

Wasshausen, D.C. 2002. Mendonciaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 472-474. 

MENISPERMACEAE 

R.C. Barneby†, P. Hiepko 

Abuta barbata Miers 

Abuta bullata Moldenke 

Abuta candollei Triana & Planch. 

Abuta grandifolia (Mart.) Sandwith 

Abuta grisebachii Triana & Planch. 

Abuta imene (Mart.) Eichler 

Abuta obovata Diels 

Abuta pahnii (Mart.) Krukoff & Barneby 

Abuta rufescens Aubl. 

Abuta sandwithiana Krukoff & Barneby 

Abuta solimoesensis Krukoff & Barneby 

Abuta velutina Gleason 

Anomospermum chloranthum Diels 

Anomospermum chloranthum subsp. chloranthum 

Anomospermum chloranthum subsp. confusum Krukoff & Barneby 

Anomospermum grandifolium Eichler 

Anomospermum steyermarkii Krukoff & Barneby 

Borismene japurensis (Mart.) Barneby 

Caryomene foveolata Barneby & Krukoff 

Caryomene glaucescens (Moldenke) Barneby & Krukoff 

Caryomene olivascens Barneby & Krukoff 

Cissampelos andromorpha DC. 

Cissampelos fasciculata Benth. 

Cissampelos ovalifolia DC. 

Cissampelos pareira L. 

Cissampelos tropaeolifolia DC. 

Curarea candicans (Rich. ex DC.) Barneby & Krukoff 

Curarea toxicofera (Wedd.) Barneby & Krukoff 

Disciphania ernstii Eichler 

Disciphania ernstii var. ernstii 

Disciphania moriorum Barneby 

BO GU SU FG 

AM BO 

BO 

FG 

GU FG 


AM 

AM 

AM BO 

AM BO 

GU SU FG 

BO 

AM 

GU SU FG 

GU FG 

GU SU FG 


AM BO FG? 



AM BO 


GU SU FG 

FG 

AM BO GU? 

FG 

FG 

FG 

GU 

AM BO FG 

BO DA SU? 

SU 

FG 

AM FG 


GU SU FG? 


BO GU SU 

BO 


AM 

AM? 

AM? 

FG


Disciphania unilateralis Barneby 

Elephantomene eburnea Barneby & Krukoff 

Hyperbaena domingensis (DC.) Benth. 

Odontocarya mallosperma Barneby 

Odontocarya sp. A 

Odontocarya tamoides (DC.) Miers 

Odontocarya tamoides var. canescens (Miers) Barneby 

Odontocarya tamoides var. tamoides 

Odontocarya wullschlaegelii (Eichler) Barneby 

Orthomene hirsuta (Krukoff & Moldenke) Barneby & Krukoff 

Orthomene prancei Barneby & Krukoff 

Orthomene schomburgkii (Miers) Barneby & Krukoff 

Orthomene verruculosa (Krukoff & Barneby) Barneby & Krukoff 

Sciadotenia cayennensis Benth. 

Sciadotenia duckei Moldenke 

Sciadotenia eichleriana Moldenke 

Sciadotenia sagotiana (Eichler) Diels 

Sciadotenia sprucei Diels in Engl. 

Telitoxicum duckei (Diels) Moldenke 

Telitoxicum inopinatum Moldenke 

Telitoxicum krukovii Moldenke 

Telitoxicum minutiflorum (Diels) Moldenke 


Abuta amara Aubl. (Aristolochiaceae) =Aristolochia amara 

Abuta candicans Rich. ex DC. =Curarea candicans 

Abuta concolor Poepp. & Endl. =Abuta grandifolia 

Abuta guyanensis Eichler =Abuta grandifolia 

Abuta klugii Moldenke =Abuta imene 

Abuta limaciifolia Diels =Curarea candicans 

Abuta macrocarpa Moldenke =Anomospermum grandifolium 

Abuta pullei Diels =Curarea candicans 

Abuta splendida Krukoff & Moldenke =Abuta rufescens 

Abuta tomentosa Sagot =Sciadotenia sagotiana 

Abuta umbellata Sagot =Sciadotenia cayennensis 

Abuta verruculosa Krukoff & Barneby =Orthomene verruculosa 

Abuta wilson-brownei R.S. Cowan =Abuta rufescens 

Anelasma domingense (DC.) Miers =Hyperbaena domingensis 

Anomospermum hirsutum Krukoff & Moldenke =Orthomene hirsuta 

Anomospermum japurense (Mart.) Eichler =Borismene japurensis 

Anomospermum schomburgkii Miers =Orthomene schomburgkii 

Chondrodendron candicans (Rich. ex DC.) Sandwith =Curarea candicans 

Chondrodendron limaciifolium (Diels) Moldenke =Curarea candicans 

Chondrodendron toxicoferum (Wedd.) Krukoff & Moldenke =Curarea toxicofera 

Cissampelos ramiflora Miers =Cissampelos andromorpha 

Cocculus domingensis DC. =Hyperbaena domingensis 

Cocculus grandifolius Mart. =Abuta grandifolia 

Cocculus imene Mart. =Abuta imene 

Cocculus japurensis Mart. =Borismene japurensis 

Cocculus pahnii Mart. =Abuta pahnii 

Cocculus pauper Griseb. =Odontocarya tamoides var. canescens 

Cocculus tamoides DC. =Odontocarya tamoides 

Cocculus toxicoferus Wedd. =Curarea toxicofera 

Hyperbaena cuatrecasasii Moldenke =Borismene japurensis 

Hyperbaena graciliflora Miers =Hyperbaena domingensis 

Odontocarya hederifolia var. canescens Miers =Odontocarya tamoides var. canescens 

Odontocarya paupera (Griseb.) Diels =Odontocarya tamoides var. canescens 

Pachygone domingensis (DC.) Eichler =Hyperbaena domingensis 

Pachygone graciliflora (Miers) Sagot =Hyperbaena domingensis 

Sciadotenia candicans Diels =Curarea candicans 

Somphoxylon wullschlaegelii Eichler =Odontocarya wullschlaegelii 

Stephania elliptica DC. (Capparaceae) =Steriphoma ellipticum 

FG 

SU FG 


BO 

AM 



DA GU SU FG 

GU SU FG 

AM 

FG 


AM FG 

AM GU SU FG 

FG 

FG 

GU SU FG 

AM 

FG 

AM GU SU FG 

GU SU 

GU? FG? 

REFERENCES 

Barneby, R.C. 1970. Revision of Neotropical Menispermaceae tribe Tinosporeae. Mem. New York Bot. Gard. 20(2): 81-158.

422 


Barneby, R.C. 2001. Menispermaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 554-578. 

Barneby, R.C. 2002. Menispermaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 474-483. 

Barneby, R.C. & B.A. Krukoff. 1971. Supplementary notes in American Menispermaceae. VIII. A generic survey of the American 

Triclisieae and Anomospermeae. Mem. New York Bot. Gard. 22(2): 1-89. 

Diels, L. 1934. Menispermaceae. In A. Pulle, Flora of Suriname 2(1): 123-131. 

Jansen-Jacobs, M.J. 1976. Menispermaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 

430-440. 

Jonker, F.P. 1939. Menispermaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 476-477. 

Krukoff, B.A. & R.C. Barneby. 1938. Studies of American Menispermaceae, with special reference to species used in preparation 

of arrow-poisons. Brittonia 3: 1-74. (See also numerous articles by R.C. Barneby, B.A. Krukoff, and H.N. Moldenke in a 

series of papers published from 1941 through 1971 in Bull. Torrey Bot. Club and Mem. New York Bot. Gard.) 

Nymphoides indica (L.) Kuntze 

MENYANTHACEAE 



Menyanthes indica L. =Nymphoides indica 

Nymphoides humboldtiana (Kunth) Kuntze =Nymphoides indica 

Nymphoides indica subsp. occidentalis A. Raynal =Nymphoides indica 

Villarsia humboldtiana Kunth =Nymphoides indica 

REFERENCES 

Jonker, F.P. 1936. Menyanthaceae. In A. Pulle, Flora of Suriname 4(1): 428-430. 

Yatskievych, K. 2001. Menyanthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 578-579. 

Glinus radiatus (Ruiz & Pav.) Rohrb. 

Mollugo verticillata L. 

Trianthema portulacastrum L. [pantropical weed, expected] 

MOLLUGINACEAE 



Mollugo radiata Ruiz & Pav. =Glinus radiatus 

REFERENCES 

Burger, W. 1983. Aizoaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212. 

Eyma, P.J. 1934. Aizoaceae. In A. Pulle, Flora of Suriname 1(1): 158-161. 

Nevling, L.I. Jr. 1961. Aizoaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. Gard. 

48(1): 80-85. 

Steyermark, J.A. 2001. MolluginaceaeIn J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 686-688. 

Mollinedia grazielae Peixoto 

Mollinedia killipii J.F. Macbr. 

Mollinedia ovata Ruiz & Pav. 

Mollinedia repanda Ruiz & Pav. 

MONIMIACEAE 

P.E. Berry, A.L. Peixoto, S.S. Renner 


Mollinedia glabricaulis Maguire & Steyerm. =Mollinedia ovata 

Mollinedia laurina Tul. =Mollinedia ovata 

Mollinedia neblinensis Maguire & Steyerm. =Mollinedia ovata 

Mollinedia ptariensis Steyerm. =Mollinedia ovata 

Mollinedia roraimensis Steyerm. =Mollinedia repanda 

Mollinedia yaviensis Lasser & Maguire =Mollinedia repanda 

BO GU SU FG 



AM GU SU 

AM 


AM BO GU


REFERENCES 

Berry, P.E. & A.L. Peixoto. 2001. Monimiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Jansen-Jacobs, M.J. 1976. Monimiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 425- 

426. 

Lanjouw, J. 1939. Monimiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 471-472. 

Mitchell, J.D. 2002. Monimiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


510-515. 

Peixoto, A.L. 1987. Nota sobre as espécies de Mollinedia (Monimiaceae, Monimioideae) das Guianas. Bradea 4(41): 331-336. 

Perkins, J. & E. Gilg. 1901. Monimiaceae. In A. Engler, Das Pflanzenreich IV(101): 1-122. 

Petter, H.F.M. 1932. Monimiaceae. In A. Pulle, Flora of Suriname 2(1): 107-112. 

Steyermark, J.A. 1951. Monimiaceae. In Botanical Exploration in Venezuela - I. Fieldiana Bot. 28(1): 235-236. 

MONOTACEAE 

J.S. Miller 

Pakaraimaea dipterocarpacea Maguire & P.S. Ashton 

Pakaraimaea dipterocarpacea subsp. dipterocarpacea 

Pakaraimaea dipterocarpacea subsp. nitida Maguire & Steyerm. 

REFERENCES 

Maguire, B. & P.S. Ashton. 1977. Pakaraimoideae, Dipterocarpaceae of the Western Hemisphere. II. Systematic, geographic, 

and phyletic considerations. Taxon 26(4): 343-368. 

Maguire, B. & P.S. Ashton. 1980. Pakaraimaea dipterocarpacea II. Taxon 29(2/3): 225-231. 

Maguire, B. & J.A. Steyermark. 1981. Pakaraimaea dipterocarpacea - III. In B. Maguire & collaborators, Botany of the Guayana 

Highland part 11. Mem. New York Bot. Gard. 32: 1-391. 

Miller, J.S. 2001. Monotaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 691-692. 

MORACEAE 

C.C. Berg 

Bagassa guianensis Aubl. 

Batocarpus amazonicus (Ducke) Fosberg 

Brosimum acutifolium Huber 

Brosimum acutifolium subsp. acutifolium 

Brosimum acutifolium subsp. obovatum (Ducke) C.C. Berg 

Brosimum alicastrum Sw. 

Brosimum alicastrum subsp. bolivarense (Pittier) C.C. Berg 

Brosimum guianense (Aubl.) Huber 

Brosimum lactescens (S. Moore) C.C. Berg 

Brosimum melanopotamicum C.C. Berg 

Brosimum parinarioides Ducke 

Brosimum parinarioides subsp. parinarioides 

Brosimum potabile Ducke 

Brosimum rubescens Taub. 

Brosimum utile (Kunth) Pittier 

Brosimum utile subsp. longifolium (Ducke) C.C. Berg 

Brosimum utile subsp. ovatifolium (Ducke) C.C. Berg 

Castilla elastica Sessé 

Castilla elastica subsp. elastica 

Clarisia biflora Ruiz & Pav. 

Clarisia ilicifolia (Spreng.) Lanj. & Rossberg 

Clarisia racemosa Ruiz & Pav. 

Dorstenia brasiliensis Lam. 

Dorstenia cayapia subsp. asaroides (Hook.) C.C. Berg [expected in SU] 

Dorstenia contrajerva L. [expected in GU] 

Ficus albert-smithii Standl. 

Ficus amazonica (Miq.) Miq. 

Ficus americana Aubl. 

Ficus blepharophylla Vázq. Avila [Mt. Roraima] 

Ficus broadwayi Urb. 

BO GU 

GU 

BO 

GU SU FG 

FG 

GU SU FG 

SU FG 

GU FG 

BO GU SU? FG 

BO GU 



AM GU 

GU SU FG 

GU SU FG 

AM 



AM 


BO 

BO 

AM 


AM BO DA SU 

AM BO GU FG 



AM BO GU SU 

BO GU SU FG

424 

Ficus caballina Standl. 

Ficus catappifolia Kunth & Bouché 

Ficus cremersii C.C. Berg 

Ficus crocata (Miq.) Miq. 

Ficus dendrocida Kunth 

Ficus donnell-smithii Standl. 

Ficus eximia Schott 

Ficus gomelleira Kunth & Bouché 

Ficus greiffiana Dugand 

Ficus guianensis Desv. ex Ham. 

Ficus hebetifolia Dugand 

Ficus insipida Willd. 

Ficus insipida subsp. insipida 

Ficus insipida subsp. scabra C.C. Berg 

Ficus krukovii Standl. 

Ficus leiophylla C.C. Berg 

Ficus malacocarpa Standl. 

Ficus maroniensis Benoist 

Ficus mathewsii (Miq.) Miq. 

Ficus matiziana Dugand 

Ficus maxima Mill. 

Ficus mollicula Pittier 

Ficus nymphaeifolia Mill. 

Ficus obtusifolia Kunth 

Ficus padifolia Kunth 

Ficus pakkensis Standl. 

Ficus paludica Standl. 

Ficus panurensis Standl. 

Ficus paraensis (Miq.) Miq. 

Ficus pertusa L. f. 

Ficus piresiana Vázq. Avila & C.C. Berg 

Ficus pulchella Schott 

Ficus roraimensis C.C. Berg 

Ficus schumacheri (Liebm.) Griseb. 

Ficus sphenophylla Standl. 

Ficus subapiculata (Miq.) Miq. 

Ficus tepuiensis C.C. Berg & Simonis [expected in GU] 

Ficus trigona L. f. 

Ficus trigonata L. 

Ficus yoponensis Desv. 

Helicostylis elegans (J.F. Macbr.) C.C. Berg 

Helicostylis pedunculata Benoist 

Helicostylis scabra (J.F. Macbr.) C.C. Berg 

Helicostylis tomentosa (Poepp. & Endl.) Rusby 

Maclura tinctoria (L.) D. Don ex Steud. 

Maclura tinctoria subsp. tinctoria 

Maquira calophylla (Poepp. & Endl.) C.C. Berg 

Maquira coriacea (H. Karst.) C.C. Berg 

Maquira guianensis Aubl. 

Maquira guianensis subsp. costaricana (Standl.) C.C. Berg 

Maquira guianensis subsp. guianensis 

Maquira sclerophylla (Ducke) C.C. Berg 

Naucleopsis glabra Spruce ex Pittier 

Naucleopsis guianensis (Mildbr.) C.C. Berg 

Naucleopsis oblongifolia (Kuhlm.) Carauta 

Naucleopsis ulei subsp. ulei [expected in FG] 

Perebea guianensis Aubl. 

Perebea guianensis subsp. guianensis 

Perebea mollis (Poepp. & Endl.) Huber 

Perebea mollis subsp. mollis 

Perebea rubra (Trécul) C.C. Berg 

Poulsenia sp. 

Pseudolmedia laevigata Trécul 

Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr. 




FG 

BO 

DA 


AM DA GU 


GU FG 


AM FG 


BO 


AM SU FG 

SU FG 


GU SU FG 

AM BO GU SU 

AM GU FG? 


AM BO 


AM GU? 

GU? 

BO GU SU FG 

DA GU SU FG 




AM GU FG 

FG? 

GU FG? 


AM BO 

GU? SU? FG 

BO 


GU? SU? 

BO 

AM BO 

SU FG 

AM 


BO GU SU FG 

BO GU 

AM FG 

AM BO SU 

BO GU SU FG 

BO 

BO GU SU FG 

AM GU SU FG 

SU FG 

GU SU FG 

AM 

AM GU SU FG 

AM GU SU FG 

SU 

SU 

SU FG 

FG 

AM BO GU FG 

AM BO DA GU SU FG


Sorocea muriculata Miq. 

Sorocea muriculata subsp. muriculata 

Sorocea muriculata subsp. uaupensis (Baill.) C.C. Berg 

Sorocea pubivena Hemsl. 

Sorocea pubivena subsp. hirtella (Mildbr.) C.C. Berg 

Sorocea pubivena subsp. oligotricha (Akkermans & C.C. Berg) C.C. Berg 

Sorocea sprucei (Baill.) J.F. Macbr. 

Sorocea sprucei subsp. sprucei 

Trymatococcus amazonicus Poepp. & Endl. 

Trymatococcus oligandrus (Benoist) Lanj. 


Acanthinophyllum ilicifolia (Spreng.) W. Burger =Clarisia ilicifolia 

Acanthinophyllum spruceana (Lanj.) W. Burger =Clarisia ilicifolia 

Acanthinophyllum strepitans Allemno =Clarisia ilicifolia 

Acanthosphaera amara (Ducke) Ducke =Naucleopsis ulei 

Alicastrum guianense (Aubl.) Kuntze =Brosimum guianense 

Aliteria sagotii Benoist =Clarisia ilicifolia 

Anonocarpus amazonicus Ducke =Batocarpus amazonicus 

Bagassa sagotiana Bureau ex Benth. & Hook. f. =Bagassa guianensis 

Bagassa tiliifolia (Ham.) Benoist =Bagassa guianensis 

Brosimopsis acutifolia (Huber) Ducke =Brosimum acutifolium 

Brosimopsis lactescens S. Moore =Brosimum lactescens 

Brosimopsis obovata Ducke =Brosimum acutifolium subsp. obovatum 

Brosimum angustifolium Ducke =Brosimum rubescens 

Brosimum aubletii Poepp. & Endl. =Brosimum guianense 

Brosimum bolivarense (Pittier) Romero =Brosimum alicastrum subsp. bolivarense 

Brosimum discolor Schott =Brosimum guianense 

Brosimum lanciferum Ducke =Brosimum rubescens 

Brosimum latifolium Standl. =Brosimum alicastrum subsp. bolivarense 

Brosimum lemeei (Benoist) Lemée =Brosimum guianense 

Brosimum longifolium Ducke =Brosimum utile subsp. longifolium 

Brosimum mello-barretoi Standl. =Naucleopsis oblongifolia 

Brosimum myristicoides Standl. =Brosimum potabile 

Brosimum ovatifolium Ducke =Brosimum utile subsp. ovatifolium 

Brosimum palmarum Standl. =Brosimum guianense 

Brosimum paraense Huber =Brosimum rubescens 

Brosimum rigidum Ducke =Brosimum utile subsp. ovatifolium 

Brosimum rotundatum Standl. =Brosimum guianense 

Brosimum velutinum (S.F. Blake) Ducke =Brosimum guianense 

Chlorophora tinctoria (L.) Gaudich. ex Benth. & Hook. f. =Maclura tinctoria 

Clarisia ilicifolia var. cuyunensis (Lanj.) Lanj. & Rossberg =Clarisia ilicifolia 

Clarisia ilicifolia var. guianensis (Lanj.) Lanj. & Rossberg =Clarisia ilicifolia 

Clarisia spruceana Lanj. =Clarisia ilicifolia 

Clarisia strepitans (Allemno) Lanj. =Clarisia ilicifolia 

Clarisia strepitans var. cuyunensis Lanj. =Clarisia ilicifolia 

Clarisia strepitans var. guianensis Lanj. =Clarisia ilicifolia 

Dorstenia sabanensis Cuatrec. =Dorstenia brasiliensis 

Dorstenia tubicina Ruiz & Pav. =Dorstenia brasiliensis 

Exoecaria ilicifolia Spreng. =Clarisia ilicifolia 

Ferolia guianensis Aubl. =Brosimum rubescens 

Ficus acarouaniensis Benoist =Ficus gomelleira 

Ficus anguina Benoist =Ficus nymphaeifolia 

Ficus angustifolia (Miq.) Miq. =Ficus amazonica 

Ficus arctocarpa Standl. =Ficus panurensis 

Ficus arukensis Standl. =Ficus paraensis 

Ficus citrifolia Mill. [sensu stricto, not in flora area] 

Ficus clusiifolia Schott =Ficus guianensis 

Ficus corpulenta Pittier =Ficus guianensis 

Ficus ernstiana Pittier =Ficus trigona 

Ficus erratica Standl. =Ficus guianensis 

Ficus erythrosticta (Miq.) Miq. =Ficus pertusa 

Ficus expansa Pittier =Ficus eximia 

Ficus fagifolia (Miq.) Miq. =Ficus trigona 


AM BO GU SU 

AM BO SU FG 

AM BO GU SU 

AM 

AM BO GU SU 

BO GU 

BO GU 


GU SU FG

426 


Ficus fanshawei Standl. =Ficus trigona 

Ficus foveata Pittier =Ficus eximia 

Ficus foveolata Pittier =Ficus eximia 

Ficus gardneriana (Miq.) Miq. [not in flora area] =Ficus obtusifolia 

Ficus gemina (Miq.) Miq. =Ficus pertusa 

Ficus gigantea Kunth [not in flora area] =Ficus citrifolia 

Ficus glabrata Kunth =Ficus insipida 

Ficus glandulosa Pittier =Ficus eximia 

Ficus glaucescens (Liebm.) Miq. =Ficus maxima 

Ficus gleasonii Standl. =Ficus mathewsii 

Ficus gracilis Pittier (Aquifoliaceae) =Ilex divaricata 

Ficus guanarensis Pittier =Ficus eximia 

Ficus hartii Warb. =Ficus trigonata 

Ficus hydrophila Pittier =Ficus paraensis 

Ficus involuta (Liebm.) Miq. =Ficus obtusifolia 

Ficus jacquiniifolia A. Rich. =Ficus americana 

Ficus kanukuensis Standl. =Ficus pertusa 

Ficus lanjouwii DeWolf =Ficus maroniensis 

Ficus leucosticta (Miq.) Miq. =Ficus paludica 

Ficus longifolia Schott [sensu stricto, not in flora area] 

Ficus longistipula Pittier =Ficus insipida subsp. insipida 

Ficus maguirei Standl. =Ficus panurensis 

Ficus manicariarum Standl. =Ficus paludica 

Ficus maroana Pittier =Ficus guianensis 

Ficus martinii Miq. =Ficus guianensis 

Ficus mendelsonii Britton =Ficus broadwayi 

Ficus mensalis Standl. =Ficus guianensis 

Ficus orinocensis Pittier =Ficus paraensis 

Ficus pallida Vahl [not in flora area] 

Ficus palmicida Pittier =Ficus pertusa 

Ficus parkeri Miq. =Ficus maxima 

Ficus parkeriana (Miq.) Sandwith =Ficus guianensis 

Ficus pascuorum Pittier =Ficus obtusifolia 

Ficus perforata L. =Ficus americana 

Ficus plicato-ostiolata Pittier =Ficus trigona 

Ficus prinoides Humb. & Bonpl. ex Willd. [not in flora area] =Ficus pallida Vahl 

Ficus radula Humb. & Bonpl. ex Willd. =Ficus maxima 

Ficus ramiflora Standl. =Ficus caballina 

Ficus rolanderi (Liebm.) Miq. =Ficus pertusa 

Ficus savannarum Standl. =Ficus broadwayi 

Ficus scabrida Pittier =Ficus malacocarpa 

Ficus surinamensis Miq. =Ficus amazonica 

Ficus tamatamae Pittier =Ficus caballina 

Ficus thelephora Benoist =Ficus paraensis 

Ficus tigrensis Pittier =Ficus mollicula 

Ficus turbinata Pittier =Ficus eximia 

Ficus urbaniana Warb. =Ficus nymphaeifolia 

Ficus velutina Willd. [not in flora area] 

Ficus vulpina Benoist =Ficus trigona 

Galactodendrum utile Kunth =Brosimum utile 

Greeneina affinis (Miq.) Kuntze =Helicostylis tomentosa 

Helicostylis affinis Steud. ex Miq. =Helicostylis tomentosa 

Helicostylis asperifolia Ducke =Helicostylis scabra 

Helicostylis bolivarensis Pittier =Brosimum alicastrum subsp. bolivarense 

Helicostylis lancifolia Ducke =Helicostylis elegans 

Helicostylis obtusifolia Standl. =Helicostylis tomentosa 

Helicostylis poeppigiana (Mart.) Trécul =Helicostylis tomentosa 

Lanessania oligandra Benoist =Trymatococcus oligandrus 

Lanessania turbinata Baill. =Trymatococcus amazonicus 

Laurea tiliifolia Gaudich. =Bagassa guianensis 

Maquira costaricana (Standl.) C.C. Berg =Maquira guianensis subsp. costaricana 

Morus tinctoria L. =Maclura tinctoria 

Naucleopsis amara Ducke =Naucleopsis ulei subsp. ulei 

Naucleopsis mello-barretoi (Standl.) C.C. Berg =Naucleopsis oblongifolia


Noyera mollis (Poepp. & Endl.) Ducke =Perebea mollis 

Noyera rubra Trécul =Perebea rubra 

Ogcodeia amara (Ducke) Ducke =Naucleopsis ulei subsp. ulei 

Ogcodeia glabra (Spruce ex Pittier) Mildbr. =Naucleopsis glabra 

Ogcodeia guianensis Mildbr. =Naucleopsis guianensis 

Ogcodeia oblongifolia Kuhlm. =Naucleopsis oblongifolia 

Olmedia asperula Standl. =Helicostylis tomentosa 

Olmedia calophylla Poepp. & Endl. =Maquira calophylla 

Olmedia caurensis Pittier =Pseudolmedia laevis 

Olmedia guianensis (Aubl.) Trécul =Maquira guianensis 

Olmedia laevis Ruiz & Pav. =Pseudolmedia laevis 

Olmedia maquira Steud. =Maquira guianensis 

Olmedia maxima Ducke =Maquira coriacea 

Olmedia mollis Poepp. & Endl. =Perebea mollis 

Olmedia obliqua Huber =Maquira coriacea 

Olmedia polycephala Pittier =Helicostylis tomentosa 

Olmedia tomentosa Poepp. & Endl. =Helicostylis tomentosa 

Olmedia virgata Pittier =Sorocea sprucei subsp. sprucei 

Olmedioperebea calophylla (Poepp. & Endl.) Ducke =Maquira calophylla 

Olmedioperebea sclerophylla Ducke =Maquira sclerophylla 

Olmediophaena maxima (Ducke) Ducke =Maquira coriacea 

Olmediophaena obliqua (Huber) Ducke =Maquira coriacea 

Perebea costaricana Standl. =Maquira guianensis subsp. costaricana 

Perebea elegans J.F. Macbr. =Helicostylis elegans 

Perebea laurifolia Trécul =Maquira guianensis subsp. guianensis 

Perebea lecointei Huber =Perebea mollis subsp. mollis 

Perebea mollis subsp. rubra (Trécul) C.C. Berg =Perebea rubra 

Perebea paraensis Huber =Perebea mollis subsp. mollis 

Perebea xinguana Standl. =Maquira sclerophylla 

Pharmacosycea guyanensis Miq. =Ficus maxima 

Pharmacosycea parkeriana Miq. =Ficus guianensis 

Piratinera acutifolia (Huber) Pittier =Brosimum acutifolium 

Piratinera guianensis Aubl. =Brosimum guianense 

Piratinera lancifera (Ducke) Benoist =Brosimum rubescens 

Piratinera lemeei Benoist =Brosimum guianense 

Piratinera paraensis (Huber) Benoist =Brosimum rubescens 

Piratinera scabridula S.F. Blake =Brosimum guianense 

Piratinera velutina S.F. Blake =Brosimum guianense 

Pseudolmedia coriacea H. Karst. =Maquira coriacea 

Pseudolmedia multinervis Mildbr. =Pseudolmedia laevis 

Pseudolmedia sagotii Benoist =Pseudolmedia laevis 

Pseudolmedia scabra J.F. Macbr. =Helicostylis scabra 

Pseudosorocea sprucei Baill. =Sorocea sprucei 

Pseudosorocea uaupensis Baill. =Sorocea muriculata subsp. uaupensis 

Sorocea amazonica Miq. =Sorocea muriculata subsp. muriculata 

Sorocea dentata Huber =Sorocea muriculata subsp. muriculata 

Sorocea guayanensis W.C. Burger =Sorocea muriculata subsp. uaupensis 

Sorocea hirtella Mildbr. =Sorocea pubivena subsp. hirtella 

Sorocea hirtella subsp. oligotricha Akkermans & C.C. Berg =Sorocea pubivena subsp. oligotricha 

Sorocea opima J.F. Macbr. =Sorocea pubivena subsp. hirtella 

Sorocea uaupensis (Baill.) J.F. Macbr. =Sorocea muriculata subsp. uaupensis 

Trymatococcus paraensis Ducke =Trymatococcus amazonicus 

Trymatococcus turbinatus (Baill.) Ducke =Trymatococcus amazonicus 

Urostigma amazonicum Miq. =Ficus amazonica 

Urostigma angustifolium Miq. =Ficus amazonica 

Urostigma crocatum Miq. =Ficus crocata 

Urostigma erythrostictum Miq. =Ficus pertusa 

Urostigma fagifolium Miq. =Ficus trigona 

Urostigma gardnerianum Miq. =Ficus obtusifolia 

Urostigma geminum Miq. =Ficus pertusa 

Urostigma involutum Liebm. =Ficus obtusifolia 

Urostigma leucostictum Miq. =Ficus paludica 

Urostigma mathewsii Miq. =Ficus mathewsii 

Urostigma paraense Miq. =Ficus paraensis

428 

Urostigma rolanderi Liebm. =Ficus pertusa 

Urostigma schumacheri Liebm. =Ficus schumacheri 


REFERENCES (see also Cecropiaceae) 

Berg, C.C. 1972. Olmedieae and Brosimeae (Moraceae). Flora Neotropica Monogr. 7: 1-229. 

Berg, C.C. 1992. Moraceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 11: 10-92. 

Berg, C.C. 2001. Moraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 6: 

693-729. 

Berg, C.C. 2002. Moraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


525. 

Berg, C.C. & R.W.A.P. Akkermans. 1985. Studies on the flora of the Guianas 14. New taxa and combinations in Sorocea and a 

key to its species. Proc. Kon. Ned. Akad. Wet. ser. C, 88(4): 381-394. 

Berg, C.C. & G.P. DeWolf. 1975. Moraceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 173-265. 

Berg, C.C., M. Vásquez Avila, & F. Kooy. 1984. Ficus species of Brazilian Amazonia and the Guianas. Acta Amaz. 14(1/2) 

suppl.: 159-194. 

Lindeman, J.C. 1984. Moraceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 652-654. 

MORINGACEAE 

J.S. Miller 

Moringa oleifera Lam. [Old World family, one species widely cultivated] 


Moringa pterygosperma Gaertn. =Moringa oleifera 

REFERENCES 

Miller, J.S. 2001. Moringaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

6: 729-730. 

Stoffers, A.L. 1984. Moringaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 570-572. 

Muntingia calabura L. 

MUNTINGIACEAE 

P.E. Berry 

REFERENCES 

(alt. Tiliaceae) 

MYOPORACEAE 

Bontia daphnoides L. [cultivated; dubiously native or naturalized] 

REFERENCES 

Lanjouw, J. 1938. Myoporaceae. In A. Pulle, Flora of Suriname 4(2): 255-256. 

Morella rotundata (Steyerm. & Maguire) Parra-Os. 

MYRICACEAE 

P.E. Berry, N. Ramírez 


Myrica rotundata Steyerm. & Maguire = Morella rotundata 

REFERENCES 

Ramírez, N. & P.E. Berry. 2001. Myrsinaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 6: 733-734. 

Compsoneura debilis (Spruce ex A. DC.) Warb. 

Compsoneura sprucei (A. DC.) Warb. 

MYRISTICACEAE 

G. Aymard, P.E. Berry, W.A. Rodrigues 

BO 

BO GU SU FG 

BO 

GU SU FG 

AM GU 

AM SU

Compsoneura ulei Warb. 

Iryanthera hostmannii (Benth.) Warb. 

Iryanthera juruensis Warb. 

Iryanthera laevis Markgr. 

Iryanthera lancifolia Ducke 

Iryanthera macrophylla (Benth.) Warb. 

Iryanthera obovata Ducke 

Iryanthera paradoxa (Schwacke) Warb. 

Iryanthera paraensis Huber 

Iryanthera sagotiana (Benth.) Warb. 

Iryanthera tessmannii Markgr. 

Iryanthera tricornis Ducke 

Iryanthera ulei Warb. 

Osteophloeum platyspermum (Spruce ex A. DC.) Warb. 

Virola calophylla (Spruce) Warb. 

Virola duckei A.C. Sm. 

Virola elongata (Benth.) Warb. 

Virola kwatae Sabatier 

Virola michelii Heckel 

Virola minutiflora Ducke 

Virola multicostata Ducke 

Virola parvifolia Ducke 

Virola pavonis (A. DC.) A.C. Sm. 

Virola rugulosa (Spruce) Warb. 

Virola sebifera Aubl. 

Virola sp. A 

Virola sp. B 

Virola steyermarkii W.A. Rodrigues 

Virola surinamensis (Rol. ex Rottb.) Warb. 

Virola theiodora (Spruce ex Benth.) Warb. 

Virola venosa (Benth.) Warb. 



Compsoneura costaricensis Warb. =Compsoneura sprucei 

Iryanthera congestiflora J.F. Macbr. =Iryanthera hostmannii 

Iryanthera elongata Huber =Iryanthera paraensis 

Iryanthera leptoclada Markgr. =Iryanthera hostmannii 

Iryanthera longiflora Ducke =Iryanthera paradoxa 

Iryanthera porcata A.H. Gentry =Iryanthera lancifolia 

Iryanthera sessilis Markgr. =Iryanthera paraensis 

Myristica calophylla Spruce =Virola calophylla 

Myristica cuspidata Spruce ex Benth. =Virola elongata 

Myristica debilis Spruce ex A. DC. =Compsoneura debilis 

Myristica elongata Benth. =Virola elongata 

Myristica fatua auct. non Houtt. 1774(?): sensu Sw. Prodr. 96. 1788 =Virola surinamensis 

Myristica hostmannii Benth. =Iryanthera hostmannii 

Myristica laurifolia Spruce ex A. DC. =Compsoneura sprucei 

Myristica longicuspis Spruce ex Warb. =Virola elongata 

Myristica macrophylla Benth. =Iryanthera macrophylla 

Myristica melinonii Benoist =Virola michelii 

Myristica membranacea Poepp. ex A. DC. =Virola elongata 

Myristica mexicana Hemsl. =Compsoneura sprucei 

Myristica mocoa A. DC. =Virola sebifera 

Myristica panamensis Hemsl. =Virola sebifera 

Myristica paradoxa Schwacke =Iryanthera paradoxa 

Myristica pavonis A. DC. =Virola pavonis 

Myristica platysperma Spruce ex A. DC. =Osteophloeum platyspermum 

Myristica punctata Spruce ex Benth. =Virola elongata 

Myristica rugulosa Spruce =Virola rugulosa 

Myristica sebifera (Aubl.) Sw. =Virola sebifera 

Myristica sebifera var. cordifolia A. DC. =Virola sebifera 

Myristica sebifera var. curvinervia A. DC. =Virola sebifera 

Myristica sebifera var. longifolia Lam. =Virola surinamensis 

Myristica sprucei A. DC. =Compsoneura sprucei 

GU FG 



AM BO 


AM GU 

AM 

AM BO 


GU SU FG 

SU FG? 

BO 

GU 


AM GU SU 

AM 


FG 

AM GU SU FG 

AM 

FG 

AM 

AM 

AM 


AM 

AM 

AM BO 


AM BO 

AM SU

430 


Myristica surinamensis Rol. ex Rottb. =Virola surinamensis 

Myristica theiodora Spruce ex Benth. =Virola theiodora 

Myristica uaupensis Spruce ex A. DC. =Virola elongata 

Myristica venosa Benth. =Virola venosa 

Myristica venosa var. pavonis (A. DC.) Warb. =Virola pavonis 

Myristica venosa var. poeppigii A. DC. =Virola venosa 

Osteophloeum sulcatum Little =Osteophloeum platyspermum 

Palala calophylla (Spruce) Kuntze =Virola calophylla 

Palala cuspidata (Spruce ex Benth.) Kuntze =Virola elongata 

Palala debilis (Spruce ex A. DC.) Kuntze =Compsoneura debilis 

Palala elongata (Benth.) Kuntze =Virola elongata 

Palala macrophylla (Benth.) Kuntze =Iryanthera macrophylla 

Palala membranacea (Poepp. ex A. DC.) Kuntze =Virola elongata 

Palala mexicana (Hemsl.) Kuntze =Compsoneura sprucei 

Palala mocoa (A. DC.) Kuntze =Virola sebifera 

Palala pavonis (A. DC.) Kuntze =Virola pavonis 

Palala platysperma (Spruce ex A. DC.) Kuntze =Osteophloeum platyspermum 

Palala punctata (Spruce ex Benth.) Kuntze =Virola elongata 

Palala rugulosa (Spruce) Kuntze =Virola rugulosa 

Palala sprucei (A. DC.) Kuntze =Compsoneura sprucei 

Palala surinamensis (Rol. ex Rottb.) Kuntze =Virola surinamensis 

Palala theiodora (Spruce ex Benth.) Kuntze =Virola theiodora 

Palala uaupensis (Spruce ex A. DC.) Kuntze =Virola elongata 

Palala venosa (Benth.) Kuntze =Virola venosa 

Virola boliviensis Warb. =Virola sebifera 

Virola cuspidata (Spruce ex Benth.) Warb. =Virola elongata 

Virola cuspidata var. membranacea (Poepp. ex A. DC.) Warb. =Virola elongata 

Virola elliptica A.C. Sm. =Virola pavonis 

Virola elongata var. longicuspis Warb. =Virola elongata 

Virola elongata var. punctata (Spruce ex Benth.) Warb. =Virola elongata 

Virola elongata var. subcordata Warb. =Virola elongata 

Virola glaziovii Warb. =Virola surinamensis 

Virola incolor Warb. =Virola calophylla 

Virola melinonii (Benoist) A.C. Sm. =Virola michelii 

Virola mocoa (A. DC.) Warb. =Virola sebifera 

Virola mycetis Pulle =Virola sebifera 

Virola panamensis (Hemsl.) Warb. =Virola sebifera 

Virola peruviana var. tomentosa Warb. =Virola sebifera 

Virola rufula Warb. =Virola theiodora 

Virola venezuelensis Warb. =Virola sebifera 

Virola venosa var. poeppigii (A. DC.) Warb. =Virola venosa 

Virola warburgii Pittier =Virola sebifera 

REFERENCES 

Jansen-Jacobs, M.J. 1976. Myristicaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 

427-429. 

Mitchell, J.J. 2002. Myristicaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


525-532. 

Ooststroom, S.J. van. 1934. Myristicaceae. In A. Pulle, Flora of Suriname 2(1): 113-122. 

Ooststroom, S.J. van. 1939. Myristicaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 473-475. 

Rodrigues, W.A. 1980. Revisao taxonômica das espécies de Virola Aublet do Brasil. Acta Amaz. 10(1) suppl. 1-127. 

Rodrigues, W.A., G. Aymard & P.E. Berry. 2001. Myristicaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Smith, A.C. & R.P. Wodehouse. 1937. The American species of Myristicaceae. Brittonia 2(5): 393-510. 

Ardisia elliptica Thunb. 

Ardisia guianensis (Aubl.) Mez 

Ardisia semicrenata Mart. 

Ctenardisia stenobotrys (Standl.) Lundell & Pipoly 

Cybianthus agostinianus Pipoly 

MYRSINACEAE 

J.J. Pipoly, III, J.M. Ricketson 

GU 


FG? 

AM 

AM


Cybianthus amplus (Mez) G. Agostini 

Cybianthus apiculatus (Steyerm.) G. Agostini 

Cybianthus breweri G. Agostini 

Cybianthus cardonae G. Agostini 

Cybianthus crotonoides (M.R. Schomb. ex Mez) G. Agostini 

Cybianthus deltatus Pipoly 

Cybianthus detergens Mart. 

Cybianthus duidae (Gleason & Moldenke) G. Agostini 

Cybianthus fabiolae Pipoly 

Cybianthus fulvopulverulentus (Mez) G. Agostini 

Cybianthus fulvopulverulentus subsp. fulvopulverulentus 

Cybianthus fulvopulverulentus subsp. magnoliifolius (Mez) Pipoly 

Cybianthus fuscus Mart. 

Cybianthus gardneri (A. DC.) G. Agostini 

Cybianthus grandifolius (Mez) G. Agostini 

Cybianthus guyanensis (A. DC.) Miq. 

Cybianthus guyanensis subsp. guyanensis 

Cybianthus guyanensis subsp. multipunctatus (A. DC.) Pipoly 

Cybianthus guyanensis subsp. pseudoicacoreus (Miq.) Pipoly 

Cybianthus holstii Pipoly 

Cybianthus huberi Pipoly 

Cybianthus julianii Pipoly 

Cybianthus lepidotus (Gleason) G. Agostini 

Cybianthus leprieurii G. Agostini 

Cybianthus liesneri Pipoly & Ricketson 

Cybianthus lineatus (Benth.) Pipoly 

Cybianthus longifolius Miq. 

Cybianthus maguirei G. Agostini ex Pipoly 

Cybianthus microbotrys A. DC. 

Cybianthus multicostatus Miq. 

Cybianthus penduliflorus Mart. 

Cybianthus piresii Pipoly 

Cybianthus plowmanii Pipoly 

Cybianthus potiaei (Mez) G. Agostini [=Cybianthus microbotrys?] 

Cybianthus prevostae Pipoly 

Cybianthus prieurii A. DC. 

Cybianthus ptariensis (Steyerm.) Pipoly 

Cybianthus punctatus (Mez) G. Agostini 

Cybianthus quelchii (N.E. Br.) G. Agostini 

Cybianthus resinosus Mez [not in flora area] 

Cybianthus reticulatus (Benth. ex Mez) G. Agostini 

Cybianthus roraimae (Steyerm.) G. Agostini 

Cybianthus sipapoensis Pipoly & G. Agostini 

Cybianthus sp. nov. 1 

Cybianthus sp. nov. 2 

Cybianthus spathulifolius G. Agostini ex Pipoly 

Cybianthus spicatus (Kunth) G. Agostini 

Cybianthus steyermarkianus (G. Agostini) G. Agostini 

Cybianthus surinamensis (Spreng.) G. Agostini 

Cybianthus venezuelanus Mez 

Cybianthus wurdackii G. Agostini ex Pipoly 

Myrsine coriacea (Sw.) R. Br. ex Roem. & Schult. 

Myrsine coriacea subsp. coriacea 

Myrsine coriacea subsp. reticulata (Steyerm.) Pipoly 

Myrsine guianensis (Aubl.) Kuntze 

Myrsine macrocarpa Pipoly 

Myrsine maguireana Pipoly 

Myrsine minima (Steyerm.) Pipoly 

Myrsine nitida (Mez) Pipoly 

Myrsine pellucida (Ruiz & Pav.) Spreng. 

Myrsine perpauciflora Pipoly 

Myrsine picturata Pipoly 

Myrsine resinosa (A.C. Sm.) Pipoly [Mt. Roraima] 

Stylogyne atra Mez 

AM BO GU FG 

AM GU 

BO 

AM BO 

BO GU 

AM 

GU 

AM BO GU 

GU 


AM BO GU SU 


SU FG? 

FG 

BO GU FG 


AM GU FG 

BO GU SU FG 

AM GU SU FG 

AM 

BO 

AM 

AM BO GU 

FG 

AM 

AM BO GU 

AM 

AM 

GU SU FG 

AM 

AM 

AM 

AM 

BO GU SU FG 

FG 


BO GU 

BO GU 

AM BO GU 

AM 

AM 

BO GU 

AM 

GU 

GU 

AM 

AM BO GU 

BO 



BO 

AM BO GU 

AM BO 

BO 


AM 

AM GU 

BO GU 

BO GU 

BO 

AM 

AM BO 

BO GU 

AM SU

432 

Stylogyne depauperata Mez 

Stylogyne incognita Pipoly 

Stylogyne lasseri (Lundell) Pipoly 

Stylogyne longifolia (Mart. ex Miq.) Mez 

Stylogyne micrantha (Kunth) Mez 

Stylogyne orinocensis (Kunth) Mez 

Stylogyne schomburgkiana (A. DC.) Mez 

Stylogyne viridis (Lundell) Ricketson & Pipoly 



Ardisia acuminata Willd. =Ardisia guianensis 

Ardisia acuminifolia Lundell =Ardisia guianensis 

Ardisia amanuensis Lundell =Ardisia guianensis 

Ardisia cookii Lundell =Ardisia guianensis 

Ardisia hostmannii Miq. =Stylogyne orinocensis 

Ardisia lasseri Lundell =Stylogyne lasseri 

Ardisia longicaudata Lundell =Ardisia guianensis 

Ardisia micrantha Kunth =Stylogyne micrantha 

Ardisia nigricans Miq. =Stylogyne atra 

Ardisia nigricans Miq. =Stylogyne orinocensis 

Ardisia orinocensis Kunth =Stylogyne orinocensis 

Ardisia pseudoicacorea Miq. =Cybianthus guyanensis subsp. pseudoicacoreus 

Ardisia quelchii N.E. Br. =Cybianthus quelchii 

Ardisia schomburgkiana (A. DC.) Miq. =Stylogyne schomburgkiana 

Ardisia stenobotrys Standl. =Ctenardisia stenobotrys 

Ardisia surinamensis Miq. =Stylogyne orinocensis 

Badula schomburgkiana A. DC. =Stylogyne schomburgkiana 

Caballeria ferruginea Ruiz & Pav. =Myrsine coriacea subsp. coriacea 

Caballeria pellucida Ruiz & Pav. =Myrsine pellucida 

Conomorpha ampla Mez =Cybianthus amplus 

Conomorpha apiculata Steyerm. =Cybianthus apiculatus 

Conomorpha candolleana Mez =Cybianthus spicatus 

Conomorpha crotonoides M.R. Schomb. ex Mez =Cybianthus crotonoides 

Conomorpha curvivenia Gleason =Cybianthus lepidotus 

Conomorpha depressa Steyerm. =Cybianthus quelchii 

Conomorpha duidae Gleason & Moldenke =Cybianthus duidae 

Conomorpha fulvopulverulenta Mez =Cybianthus fulvopulverulentus 

Conomorpha glaucorubens Mez =Cybianthus spicatus 

Conomorpha gracilis A.C. Sm. =Cybianthus spicatus 

Conomorpha grandiflora Mez =Cybianthus spicatus 

Conomorpha guyanensis A. DC. =Cybianthus guyanensis 

Conomorpha heterantha Benth. ex Miq. =Cybianthus guyanensis subsp. guyanensis 

Conomorpha latifolia (Miq.) Mez =Cybianthus spicatus 

Conomorpha laxiflora A. DC. =Cybianthus spicatus 

Conomorpha laxiflora var. latifolia Miq. =Cybianthus spicatus 

Conomorpha lepidota Gleason =Cybianthus lepidotus 

Conomorpha lepidota f. acutata Steyerm. =Cybianthus lepidotus 

Conomorpha leprosa Steyerm. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus 

Conomorpha lithophyta R.E. Schult. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus 

Conomorpha macrophylla Mart. =Cybianthus amplus 

Conomorpha madeirensis A.C. Sm. =Cybianthus spicatus 

Conomorpha magnoliifolia Mez =Cybianthus fulvopulverulentus subsp. magnoliifolius 

Conomorpha multipunctata (A. DC.) Miq. =Cybianthus guyanensis subsp. pseudoicacoreus 

Conomorpha peruviana var. guyanensis (A. DC.) Mez =Cybianthus guyanensis 

Conomorpha ptariensis Steyerm. =Cybianthus duidae 

Conomorpha punctata Mez =Cybianthus punctatus 

Conomorpha reticulata Benth. ex Miq. =Cybianthus reticulatus 

Conomorpha rigida Mez =Cybianthus fulvopulverulentus subsp. magnoliifolius 

Conomorpha riparia R.E. Schult. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus 

Conomorpha roraimae Steyerm. =Cybianthus roraimae 

Conomorpha sessilis A.C. Sm. =Cybianthus punctatus 

Conomorpha spicata (Kunth) Mez =Cybianthus spicatus 

Conomorpha steyermarkiana G. Agostini =Cybianthus steyermarkianus 

Conomorpha utiarityi Hoehne =Cybianthus amplus 

FG 

FG 

BO GU 

GU SU FG 



AM 

GU FG


Ctenardisia stenobotrys (Standl.) Fonnegra & S.L. Jung [nom. superfl.] =Ctenardisia stenobotrys 

Cybianthus brownii Gleason =Cybianthus venezuelanus 

Cybianthus comatus Mez =Cybianthus prieurii 

Cybianthus egensis Mez =Cybianthus venezuelanus 

Cybianthus foliosus Rusby =Stylogyne micrantha 

Cybianthus gracillimus Mez [GU records misident., fide Pipoly] 

Cybianthus lithophytus (R.E. Schult.) G. Agostini =Cybianthus fulvopulverulentus subsp. fulvopulverulentus 

Cybianthus magnoliifolius (Mez) G. Agostini =Cybianthus fulvopulverulentus subsp. magnoliifolius 

Cybianthus multipunctatus A. DC. =Cybianthus guyanensis subsp. multipunctatus 

Cybianthus nitidus Miq. =Cybianthus prieurii 

Cybianthus pseudoicacoreus (Miq.) G. Agostini =Cybianthus guyanensis subsp. pseudoicacoreus 

Cybianthus subspicatus Benth. ex Miq. =Cybianthus prieurii 

Cybianthus sylvaticus (Gleason) G. Agostini =Cybianthus grandifolius 

Cybianthus viridiflorus A.C. Sm. =Cybianthus prieurii 

Grammadenia lineata Benth. =Cybianthus lineatus 

Grammadenia ptariensis Steyerm. =Cybianthus ptariensis 

Grammadenia ptariensis subsp. auyantepuiensis G. Agostini =Cybianthus ptariensis 

Icacorea acuminata (Willd.) Lundell [not in flora area] =Ardisia guianensis 

Icacorea acuminifolia (Lundell) Lundell =Ardisia guianensis 

Icacorea amanuensis (Lundell) Lundell =Ardisia guianensis 

Icacorea cookii (Lundell) Lundell =Ardisia guianensis 

Icacorea guianensis Aubl. =Ardisia guianensis 

Icacorea longicaudata (Lundell) Lundell =Ardisia guianensis 

Manglilla ferruginea (Ruiz & Pav.) Roem. & Schult. =Myrsine coriacea subsp. coriacea 

Manglilla pellucida (Ruiz & Pav.) Roem. & Schult. =Myrsine pellucida 

Myrsine ferruginea (Ruiz & Pav.) Spreng. =Myrsine coriacea subsp. coriacea 

Myrsine myricoides Schltdl. =Myrsine coriacea subsp. coriacea 

Myrsine ovalifolia Miq. =Myrsine guianensis 

Myrsine roraimensis (A.C. Sm.) Pipoly =Myrsine nitida 

Myrsine schomburgkiana Miq. (Sapotaceae) =Elaeoluma schomburgkiana 

Myrsine spicata Kunth =Cybianthus spicatus 

Parathesis viridis Lundell =Stylogyne viridis 

Peckia guyanensis (A. DC.) Kuntze =Cybianthus guyanensis 

Peckia longifolia (Miq.) Kuntze =Cybianthus longifolius 

Peckia penduliflora (Mart.) Kuntze =Cybianthus penduliflorus 

Peckia prieureii (A. DC.) Kuntze =Cybianthus prieurii 

Peckia purpurea Rusby =Cybianthus venezuelanus 

Peckia surinamensis (Spreng.) Kuntze =Cybianthus surinamensis 

Rapanea ambigua Mez =Myrsine coriacea subsp. coriacea 

Rapanea coriacea (Sw.) Mez =Myrsine coriacea 

Rapanea ferruginea (Ruiz & Pav.) Mez =Myrsine coriacea subsp. coriacea 

Rapanea guianensis Aubl. =Myrsine guianensis 

Rapanea minima Steyerm. =Myrsine minima 

Rapanea myricoides (Schltdl.) Lundell =Myrsine coriacea subsp. coriacea 

Rapanea nitida Mez =Myrsine nitida 

Rapanea oblonga Pohl ex Miq. =Myrsine guianensis 

Rapanea ovalifolia (Miq.) Mez =Myrsine guianensis 

Rapanea pellucida (Ruiz & Pav.) Mez =Myrsine pellucida 

Rapanea perforata Mez =Myrsine pellucida 

Rapanea resinosa A.C. Sm. =Myrsine resinosa 

Rapanea reticulata Steyerm. =Myrsine coriacea subsp. reticulata 

Rapanea roraimensis A.C. Sm. =Myrsine nitida 

Salvadora surinamensis Spreng. =Cybianthus surinamensis 

Samara coriacea Sw. =Myrsine coriacea 

Stylogyne amazonica Mez =Stylogyne orinocensis 

Stylogyne kappleri Mez =Stylogyne orinocensis 

Stylogyne latifolia A.C. Sm. =Stylogyne micrantha 

Stylogyne micans Mez =Stylogyne orinocensis 

Stylogyne spruceana Mez =Stylogyne atra 

Stylogyne surinamensis (Miq.) Mez =Stylogyne orinocensis 

Stylogyne turbacensis Mez [reported from FG in error, fide Pipoly] 

Stylogyne venezuelana Mez =Stylogyne micrantha 

Tinus guianensis (Aubl.) Kuntze =Ardisia guianensis 

Tinus hostmannii (Miq.) Kuntze =Stylogyne orinocensis

434 


Tinus orinocensis (Kunth) Kuntze =Stylogyne orinocensis 

Tinus schomburgkiana (A. DC.) Kuntze =Stylogyne schomburgkiana 

Tinus surinamensis (Miq.) Kuntze =Stylogyne orinocensis 

Walleria laxiflora Mart. =Cybianthus spicatus 

Weigeltia grandifolia Mez =Cybianthus grandifolius 

Weigeltia grandifolius Mez =Cybianthus grandifolius 

Weigeltia longifolia (Miq.) Benth. =Cybianthus longifolius 

Weigeltia microbotrys (DC.) Mez =Cybianthus microbotrys 

Weigeltia parviflora Mez =Cybianthus leprieurii 

Weigeltia potiaei Mez =Cybianthus potiaei 

Weigeltia quelchii (N.E. Br.) Mez =Cybianthus quelchii 

Weigeltia surinamensis (Spreng.) Mez =Cybianthus surinamensis 

Weigeltia sylvatica Gleason =Cybianthus grandifolius 

REFERENCES 

Agostini, G. 1980. Una nueva clasificacion del genero Cybianthus (Myrsinaceae). Acta Biol. Venez. 10: 129-185. 

Bottelier, H.P. 1936. Myrsinaceae. In A. Pulle, Flora of Suriname 4(1): 431-442. 

Pipoly, J.J. 1983. Contributions toward a monograph of Cybianthus, III: a revision of subgenus Laxiflorus. Brittonia 35(1): 61- 

80. 

Pipoly, J.J. 1992. The genus Cybianthus subgenus Conomorpha in Guayana. Ann. Missouri Bot. Gard. 79(4): 908-957. 

Pipoly, J.J. 2002. Myrsinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


539. 

Pipoly, J.J. & J.M. Ricketson. 2001. Myrsinaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Blepharocalyx eggersii (Kiaersk.) Landrum 

Calycolpus alternifolius (Gleason) Landrum 

Calycolpus bolivarensis Landrum 

Calycolpus calophyllus (Kunth) O. Berg 

Calycolpus cochleatus McVaugh 

Calycolpus goetheanus (DC.) O. Berg 

Calycolpus revolutus (Schauer) O. Berg 

Calycolpus roraimensis Steyerm. 

Calycolpus schomburgkianus O. Berg 

Calycolpus surinamensis McVaugh 

Calycorectes batavorum McVaugh 

Calycorectes bergii Sandwith 

Calycorectes enormis McVaugh 

Calycorectes grandifolius O. Berg 

Calycorectes guyanensis Mattos 

Calycorectes maximus McVaugh 

Calyptranthes amshoffae McVaugh 

Calyptranthes bipennis O. Berg 

Calyptranthes bracteata M.L. Kawas. & B. Holst 

Calyptranthes clusiifolia O. Berg 

Calyptranthes conduplicata B. Holst 

Calyptranthes crebra McVaugh 

Calyptranthes fasciculata O. Berg 

Calyptranthes forsteri O. Berg 

Calyptranthes lepida McVaugh 

Calyptranthes lucida Mart. ex DC. 

Calyptranthes macrophylla O. Berg 

Calyptranthes multiflora O. Berg 

Calyptranthes nigrescens B. Holst 

Calyptranthes ovoidea McVaugh 

Calyptranthes pulchella DC. 

Calyptranthes pullei Burret ex Amshoff 

Calyptranthes pullei var. immaculata McVaugh 

Calyptranthes ruiziana O. Berg 

Calyptranthes smithii McVaugh 

MYRTACEAE 

F. Grifo, P. Hiepko, B.K. Holst, L. Landrum 

AM BO DA GU 

AM BO 

BO 

AM 

GU 


GU SU FG 

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Calyptranthes speciosa Sagot 

Calyptranthes speciosa var. speciosa 

Calyptranthes spicata Amshoff 

Calyptranthes vexata McVaugh 

Campomanesia aromatica (Aubl.) Griseb. 

Campomanesia grandiflora (Aubl.) Sagot 

Eugenia albicans (O. Berg) Urb. 

Eugenia amblyosepala McVaugh 

Eugenia amshoffae McVaugh 

Eugenia anastomosans DC. 

Eugenia arawakorum Sandwith 

Eugenia argyrophylla B. Holst & M.L. Kawas. 

Eugenia armeniaca Sagot 

Eugenia baileyi Britton 

Eugenia biflora (L.) DC. 

Eugenia brownsbergii Amshoff 

Eugenia cachoeirensis O. Berg 

Eugenia callichroma McVaugh 

Eugenia chrysophyllum Poir. 

Eugenia citrifolia Poir. 

Eugenia coffeifolia DC. 

Eugenia conduplicata B. Holst 

Eugenia conjuncta Amshoff 

Eugenia cowanii McVaugh 

Eugenia cribrata McVaugh 

Eugenia cuaoensis McVaugh 

Eugenia cucullata Amshoff 

Eugenia cupulata Amshoff 

Eugenia cupulata var. cupulata 

Eugenia denigrata McVaugh 

Eugenia dentata (O. Berg) Nied. 

Eugenia diplocampta Diels 

Eugenia dittocrepis O. Berg 

Eugenia egensis DC. 

Eugenia emarginata (Kunth) DC. 

Eugenia essequiboensis Sandwith 

Eugenia eurycheila O. Berg 

Eugenia exaltata Rich. ex O. Berg 

Eugenia excelsa O. Berg 

Eugenia fasciculiflora O. Berg 

Eugenia feijoi O. Berg 

Eugenia ferreiraeana O. Berg 

Eugenia flavescens DC. 

Eugenia florida DC. 

Eugenia foetida Pers. [unverified name] 

Eugenia gomesiana O. Berg 

Eugenia gongylocarpa M.L. Kawas. & B. Holst 

Eugenia griseiflora McVaugh 

Eugenia incanescens Benth. 

Eugenia kaieteurensis Amshoff 

Eugenia lambertiana DC. 

Eugenia lambertiana var. hispidula McVaugh 

Eugenia lambertiana var. lambertiana 

Eugenia latifolia Aubl. 

Eugenia ligustrina (Sw.) Willd. 

Eugenia ligustrina var. hebecarpa Amshoff 

Eugenia ligustrina var. ligustrina 

Eugenia longiracemosa Kiaersk. 

Eugenia luciae Amshoff 

Eugenia macrocalyx (Rusby) McVaugh 

Eugenia magna B. Holst 

Eugenia marlierioides Rusby 

Eugenia marowynensis Miq. 

Eugenia mimus McVaugh 


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436 


Eugenia minuscula McVaugh 

Eugenia monticola (Sw.) DC. 

Eugenia morii B. Holst & M.L. Kawas. 

Eugenia moritziana H. Karst. 

Eugenia muricata DC. 

Eugenia omissa McVaugh 

Eugenia pachystachya McVaugh 

Eugenia patens Poir. 

Eugenia patrisii Vahl 

Eugenia polystachya Rich. 

Eugenia polystachyoides Amshoff 

Eugenia producta DC. 

Eugenia protenta McVaugh 

Eugenia pseudopsidium Jacq. 

Eugenia pubescens (Kunth) DC. 

Eugenia punicifolia (Kunth) DC. 

Eugenia quadriovulata Amshoff 

Eugenia ramiflora Desv. 

Eugenia roseiflora McVaugh 

Eugenia sigillata McVaugh 

Eugenia sp. A 

Eugenia sp. B 

Eugenia spruceana O. Berg 

Eugenia stipitata McVaugh [cultivated in southern Amazonas] 

Eugenia tafelbergica Amshoff 

Eugenia tapacumensis O. Berg 

Eugenia tepuiensis Steyerm. 

Eugenia tetramera (McVaugh) M.L. Kawas. & B. Holst 

Eugenia trinervia Vahl 

Eugenia tumulescens McVaugh 

Eugenia umbonata McVaugh 

Eugenia undulata Aubl. 

Eugenia uniflora L. [cultivated; naturalized?] 

Eugenia wullschlaegeliana Amshoff 

Eugenia yatuae (McVaugh) B. Holst 

Marlierea bipennis (O. Berg) McVaugh 

Marlierea biptera Amshoff 

Marlierea buxifolia Amshoff 

Marlierea caesariata McVaugh 

Marlierea cana McVaugh 

Marlierea caudata McVaugh 

Marlierea convexivenia B. Holst 

Marlierea cuprea Amshoff 

Marlierea ensiformis McVaugh [Serra da Neblina] 

Marlierea ferruginea (Poir.) McVaugh 

Marlierea foveolata B. Holst 

Marlierea guildingiana (Griseb.) Krug & Urb. 

Marlierea insignis McVaugh 

Marlierea karuaiensis (Steyerm.) McVaugh 

Marlierea ligustrina McVaugh [probably restricted to Venezuela, fide Holst] 

Marlierea lituatinervia (O. Berg) McVaugh 

Marlierea macrophylla Amshoff 

Marlierea maguirei McVaugh 

Marlierea mcvaughii B. Holst 

Marlierea montana (Aubl.) Amshoff 

Marlierea multiglomerata Amshoff 

Marlierea pudica McVaugh 

Marlierea rugosior McVaugh 

Marlierea salticola Amshoff 

Marlierea schomburgkiana O. Berg 

Marlierea scytophylla Diels 

Marlierea sp. A 

Marlierea spruceana O. Berg 

Marlierea subcordata B. Holst 

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Marlierea suborbicularis McVaugh 

Marlierea subulata McVaugh 

Marlierea summa McVaugh 

Marlierea umbraticola (Kunth) O. Berg 

Marlierea uniflora McVaugh 

Marlierea ventuarensis B. Holst 

Marlierea vicina McVaugh 

Melaleuca leucadendron L. [cultivated; naturalized?] 

Melaleuca quinquenervia (Cav.) S.T. Blake [cultivated? naturalized?] 

Myrcia albidotomentosa (Amshoff) McVaugh 

Myrcia aliena McVaugh 

Myrcia amazonica DC. 

Myrcia bolivarensis (Steyerm.) McVaugh 

Myrcia bonnetiasylvestris (Steyerm.) Steyerm. 

Myrcia bracteata (Rich.) DC. 

Myrcia calycampa Amshoff 

Myrcia citrifolia (Aubl.) Urb. 

Myrcia clausa McVaugh [expected in GU] 

Myrcia clusiifolia (Kunth) DC. 

Myrcia compta McVaugh 

Myrcia coumeta (Aubl.) DC. 

Myrcia crispa McVaugh 

Myrcia cuprea (O. Berg) Kiaersk. 

Myrcia decorticans DC. 

Myrcia deflexa (Poir.) DC. 

Myrcia dichasialis McVaugh 

Myrcia ehrenbergiana (O. Berg) McVaugh 

Myrcia exploratoris McVaugh 

Myrcia extranea McVaugh 

Myrcia fallax (Rich.) DC. 

Myrcia fenestrata DC. 

Myrcia fenzliana O. Berg 

Myrcia gentryi B. Holst 

Myrcia gigas McVaugh 

Myrcia gonini McVaugh 

Myrcia graciliflora Sagot 

Myrcia grandis McVaugh 

Myrcia guianensis (Aubl.) DC. 

Myrcia guianensis var. guianensis 

Myrcia imperfecta McVaugh 

Myrcia inaequiloba (DC.) D. Legrand 

Myrcia induta McVaugh 

Myrcia intonsa (McVaugh) B. Holst 

Myrcia kylistophylla B. Holst 

Myrcia liesneri B. Holst 

Myrcia lucida McVaugh 

Myrcia magnoliifolia DC. 

Myrcia minutiflora Sagot 

Myrcia multiflora (Lam.) DC. 

Myrcia myriantha McVaugh 

Myrcia nigrescens DC. 

Myrcia nubicola McVaugh 

Myrcia paivae O. Berg 

Myrcia pistrinalis McVaugh 

Myrcia platyclada DC. 

Myrcia porphyrea McVaugh 

Myrcia ptariensis (Steyerm.) McVaugh 

Myrcia pyrifolia (Desv. ex Ham.) Nied. 

Myrcia quitarensis (Benth.) Sagot 

Myrcia revolutifolia McVaugh 

Myrcia rotundata (Amshoff) McVaugh 

Myrcia rotundata var. atrans McVaugh 

Myrcia rotundata var. rotundata 

Myrcia rufipila McVaugh 

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438 


Myrcia rupta M.L. Kawas. & B. Holst 

Myrcia salticola (Steyerm.) McVaugh 

Myrcia saxatilis (Amshoff) McVaugh 

Myrcia servata McVaugh 

Myrcia sessiliflora McVaugh 

Myrcia sipapensis McVaugh 

Myrcia sororopanensis Steyerm. 

Myrcia sp. A 

Myrcia sp. B 

Myrcia sp. C 

Myrcia sp. D 

Myrcia speciosa (Amshoff) McVaugh 

Myrcia splendens (Sw.) DC. 

Myrcia subobliqua (Benth.) Nied. 

Myrcia subsessilis O. Berg 

Myrcia subsessilis var. ovalis (O. Berg) McVaugh 

Myrcia subsessilis var. subsessilis 

Myrcia sylvatica (G. Mey.) DC. 

Myrcia tafelbergica Amshoff 

Myrcia tepuiensis Steyerm. 

Myrcia tomentosa (Aubl.) DC. 

Myrcianthes fragrans (Sw.) McVaugh 

Myrcianthes prodigiosa McVaugh 

Myrcianthes rhopaloides (Kunth) McVaugh 

Myrciaria amazonica O. Berg [=Myrciaria floribunda, fide Sobral] 

Myrciaria cordata O. Berg 

Myrciaria dubia (Kunth) McVaugh 

Myrciaria floribunda (West ex Willd.) O. Berg 

Myrciaria puberulenta B. Holst 

Myrciaria tenella (DC.) O. Berg 

Myrciaria vismeifolia (Benth.) O. Berg 

Myrteola nummularia (Poir.) O. Berg 

Plinia costata Amshoff 

Plinia involucrata (O. Berg) McVaugh 

Plinia pinnata L. 

Plinia rivularis (Cambess.) Rotman 

Plinia sp. A 

Plinia sp. B 

Pseudanamomis umbellulifera (Kunth) Kausel 

Psidium acutangulum DC. 

Psidium cinereum DC. 

Psidium densicomum DC. 

Psidium grandifolium Mart. ex DC. 

Psidium guajava L. [cultivated, naturalized] 

Psidium guineense Sw. 

Psidium laruotteanum Cambess. 

Psidium maribense DC. 

Psidium salutare (Kunth) O. Berg 

Psidium sartorianum (O. Berg) Nied. 

Psidium striatulum DC. 

Siphoneugena dussii (Krug & Urb.) Proença 

Syzygium cumini (L.) Skeels [cultivated; naturalized?] 

Syzygium jambos (L.) Alston [cultivated, frequently escaping] 

Syzygium malaccense (L.) Merr. & Perry [cultivated, frequently escaping] 

Ugni myricoides (Kunth) O. Berg 


Acrandra guianensis Amshoff =Eugenia amshoffae 

Aulomyrcia albidotomentosa Amshoff =Myrcia albidotomentosa 

Aulomyrcia amshoffiana Steyerm. =Myrcia albidotomentosa 

Aulomyrcia bolivarensis Steyerm. =Myrcia bolivarensis 

Aulomyrcia chilensis O. Berg =Myrcia aliena 

Aulomyrcia citrifolia (Aubl.) Amshoff =Myrcia citrifolia 

Aulomyrcia divaricata O. Berg =Myrcia rufipila 

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Aulomyrcia dumosa O. Berg =Myrcia platyclada 

Aulomyrcia edulis O. Berg =Myrcia platyclada 

Aulomyrcia grandiflora O. Berg [not in flora area] 

Aulomyrcia hostmanniana O. Berg =Myrcia amazonica 

Aulomyrcia inaequiloba var. paniculata (O. Berg) Amshoff =Myrcia inaequiloba 

Aulomyrcia karuaiensis Steyerm. =Marlierea karuaiensis 

Aulomyrcia obtusa (Schauer) O. Berg =Myrcia guianensis var. guianensis 

Aulomyrcia paniculata O. Berg =Myrcia inaequiloba 

Aulomyrcia parvifolia Steyerm. =Myrcia rotundata var. rotundata 

Aulomyrcia pirarensis O. Berg =Myrcia inaequiloba 

Aulomyrcia platyclada (DC.) Amshoff =Myrcia platyclada 

Aulomyrcia platyclada var. kaieteurensis Amshoff =Myrcia platyclada 

Aulomyrcia ptariensis Steyerm. =Myrcia ptariensis 

Aulomyrcia pyrifolia (Desv. ex Ham.) O. Berg =Myrcia pyrifolia 

Aulomyrcia roraimae (Oliv. ex Thurn) Steyerm. =Myrcia guianensis var. guianensis 

Aulomyrcia roraimensis O. Berg =Myrcia guianensis var. guianensis 

Aulomyrcia rosulans O. Berg =Myrcia tomentosa 

Aulomyrcia rotundata Amshoff =Myrcia rotundata 

Aulomyrcia salticola Steyerm. =Myrcia salticola 

Aulomyrcia schaueriana (Miq.) Amshoff =Myrcia graciliflora 

Aulomyrcia spp. =Myrcia spp. 

Aulomyrcia spruceana O. Berg =Myrcia amazonica 

Aulomyrcia tetramera Amshoff ? =Myrcia decorticans 

Aulomyrcia tomentosa (Aubl.) Amshoff =Myrcia tomentosa 

Aulomyrcia triflora O. Berg =Myrcia citrifolia 

Aulomyrcia vinacea Steyerm. =Myrcia multiflora 

Calycampe angustifolia O. Berg =Myrcia calycampa 

Calycampe latifolia O. Berg =Myrcia calycampa 

Calycolpus calophyllus var. angustifolius Steyerm. =Calycolpus calophyllus 

Calycolpus chnoiophyllus L. Riley =Calycolpus revolutus 

Calycolpus glaber (Benth.) O. Berg =Calycolpus goetheanus 

Calycolpus pyrifer L. Riley =Calycolpus revolutus 

Calycorectes macrocalyx Rusby =Eugenia macrocalyx 

Calycorectes yatuae McVaugh =Eugenia yatuae 

Calyptranthes apoda McVaugh [nom. illeg.] =Calyptranthes vexata 

Calyptranthes florifera McVaugh =Calyptranthes multiflora 

Calyptranthes obtusa Benth. =Marlierea montana 

Calyptranthes sericea Griseb. =Calyptranthes fasciculata 

Campomanesia coaetanea O. Berg =Campomanesia aromatica 

Campomanesia glabra Benth. =Calycolpus goetheanus 

Campomanesia tetramera McVaugh =Eugenia tetramera 

Catinga aromatica Aubl. =Eugenia feijoi 

Catinga moschata Aubl. =Calycorectes bergii 

Eugenia acetosans Poir. =Myrcia citrifolia 

Eugenia adenocalyx DC. =Eugenia citrifolia 

Eugenia atropunctata Steud. =Eugenia florida 

Eugenia benthamii O. Berg =Eugenia punicifolia 

Eugenia brachybotrya DC. (Melastomataceae) =Mouriri guianensis 

Eugenia bracteata Rich. =Myrcia bracteata 

Eugenia caurensis Steyerm. =Eugenia biflora 

Eugenia comete Aubl. =Myrcia coumeta 

Eugenia compta O. Berg =Eugenia pseudopsidium 

Eugenia congestissima Diels =Eugenia dittocrepis 

Eugenia coriacea O. Berg =Eugenia polystachyoides 

Eugenia cryptadena var. gracilis Amshoff =Eugenia cowanii 

Eugenia cumini (L.) Druce =Syzygium cumini 

Eugenia deflexa Poir =Myrcia deflexa 

Eugenia fallax Rich. =Myrcia fallax 

Eugenia ferruginea Poir. =Marlierea ferruginea 

Eugenia flavescens var. guianensis Sagot =Eugenia flavescens 

Eugenia floribunda West ex Willd. =Myrciaria floribunda 

Eugenia fulvipes Sagot =Eugenia ramiflora 

Eugenia guianensis Aubl. =Myrcia guianensis 

Eugenia helinovis Sagot =Eugenia coffeifolia

440 


Eugenia inaequiloba DC. =Myrcia inaequiloba 

Eugenia jambos L. =Syzygium jambos 

Eugenia kunthiana DC. =Eugenia punicifolia 

Eugenia lingua O. Berg =Eugenia cachoeirensis 

Eugenia llewelynii Steyerm. =Eugenia chrysophyllum 

Eugenia malaccensis L. =Syzygium malaccense 

Eugenia melinonis Sagot =Eugenia coffeifolia 

Eugenia mini Aubl. =Eugenia biflora 

Eugenia montana Aubl. =Marlierea montana 

Eugenia monticola var. latifolia Krug & Urb. =Eugenia monticola 

Eugenia monticola var. racemosa Amshoff =Eugenia monticola 

Eugenia multiflora Lam. =Myrcia multiflora 

Eugenia nigra DC. (Melastomataceae) =Mouriri nigra 

Eugenia nitida Benth. =Myrcia inaequiloba 

Eugenia ochra O. Berg =Eugenia tapacumensis 

Eugenia ottonis O. Berg =Eugenia pubescens 

Eugenia peregrina McVaugh =Eugenia baileyi 

Eugenia prieurii Jacq. =Eugenia pseudopsidium 

Eugenia producta auct. non DC. 1828: sensu McVaugh =Eugenia sp. B 

Eugenia prosoneura O. Berg =Eugenia gomesiana 

Eugenia protracta O. Berg [non Steud. 1843] =Eugenia protenta 

Eugenia protracta Steud. =Myrciaria floribunda 

Eugenia psidioides DC. =Eugenia pseudopsidium 

Eugenia ptariensis Steyerm. =Eugenia anastomosans 

Eugenia punicifolia var. brachypoda (DC.) Krug & Urb. =Eugenia ramiflora 

Eugenia pyrifolia Desv. ex Ham. =Myrcia pyrifolia 

Eugenia quitarensis Benth. =Myrcia quitarensis 

Eugenia racemifera Sagot =Eugenia florida 

Eugenia ramiflora var. montana Amshoff =Eugenia ferreiraeana 

Eugenia rivularis Cambess. =Plinia rivularis 

Eugenia rondonensis Steyerm. =Myrcianthes fragrans 

Eugenia roraimana O. Berg =Eugenia tapacumensis 

Eugenia salzmannii Benth. =Myrciaria floribunda 

Eugenia schomburgkii Benth. =Eugenia lambertiana var. lambertiana 

Eugenia sinemariensis Aubl. [name of uncertain application] =Eugenia sp. 

Eugenia steyermarkii Standl. =Myrcianthes fragrans 

Eugenia subalterna Benth. =Eugenia punicifolia 

Eugenia subobliqua Benth. =Myrcia subobliqua 

Eugenia tenella DC. =Myrciaria tenella 

Eugenia tomentosa Aubl. =Myrcia tomentosa 

Eugenia turumiquirensis Steyerm. =Myrcianthes fragrans 

Eugenia vismeaefolia Benth. =Myrciaria vismeifolia 

Eugenia wentii Amshoff =Eugenia macrocalyx 

Gomidesia bonnetiasylvestris Steyerm. =Myrcia bonnetiasylvestris 

Gomidesia lindeniana O. Berg =Myrcia fenzliana 

Jambosa vulgaris DC. =Syzygium jambos 

Krugia ferruginea (Poir.) Urb. =Marlierea ferruginea 

Marlierea acuminata O. Berg =Marlierea ferruginea 

Marlierea dussii Krug & Urb. =Siphoneugena dussii 

Marlierea gleasonii McVaugh =Myrcia decorticans 

Marlierea intonsa McVaugh =Myrcia intonsa 

Marlierea obtusa (Benth.) O. Berg =Marlierea montana 

Marlierea uaupensis O. Berg =Marlierea spruceana 

Mitranthes sartoriana O. Berg =Psidium sartorianum 

Myrcia acuminata (Kunth) DC. =Myrcia fallax 

Myrcia aguitensis Gleason =Myrcia fallax 

Myrcia berberis DC. =Myrcia fallax 

Myrcia bipennis (O. Berg) McVaugh =Marlierea bipennis 

Myrcia divaricata O. Berg =Myrcia rufipila 

Myrcia dumosa (O. Berg) Krug & Urb. =Myrcia platyclada 

Myrcia edulis (O. Berg) Krug & Urb. =Myrcia platyclada 

Myrcia ferruginea (Poir.) DC. =Marlierea ferruginea 

Myrcia grandiflora (O. Berg) Lemée [not in flora area] =Aulomyrcia grandiflora 

Myrcia granulata R.O. Williams =Plinia rivularis


Myrcia hirtellaefolia Gleason =Myrcia bracteata 

Myrcia hostmanniana Kiaersk. =Myrcia amazonica 

Myrcia humboldtiana DC. =Myrcia deflexa 

Myrcia kegeliana O. Berg =Myrcia fallax 

Myrcia laevis O. Berg =Myrcia lucida 

Myrcia leptoclada DC. =Myrcia amazonica 

Myrcia lucida var. attenuata McVaugh =Myrcia lucida 

Myrcia obtusa Schauer =Myrcia guianensis var. guianensis 

Myrcia paniculata (Jacq.) Krug & Urb. =Myrcia citrifolia 

Myrcia planifolia McVaugh =Myrcia revolutifolia 

Myrcia roraimae Oliv. ex Thurn =Myrcia guianensis var. guianensis 

Myrcia schomburgkiana O. Berg [nom. illeg.] =Myrcia servata 

Myrcia spruceana O. Berg =Myrcia fallax 

Myrcia subsessilis ß ovalis O. Berg =Myrcia subsessilis var. ovalis 

Myrcia subsessilis var. subcordata O. Berg =Myrcia subsessilis var. subsessilis 

Myrcia tetramera Amshoff ? =Myrcia decorticans 

Myrcia triflora Cambess. =Myrcia citrifolia 

Myrcianthes umbellulifera (Kunth) Alain =Pseudanamomis umbellulifera 

Myrciaria bipennis O. Berg =Marlierea bipennis 

Myrciaria caurensis Steyerm. =Myrciaria dubia 

Myrciaria ehrenbergiana O. Berg =Myrcia ehrenbergiana 

Myrciaria involucrata O. Berg =Plinia involucrata 

Myrciaria lituatinervia O. Berg =Marlierea lituatinervia 

Myrciaria nitida (Benth.) O. Berg =Myrcia inaequiloba 

Myrciaria paraensis O. Berg =Myrciaria dubia 

Myrciaria protracta (Steud.) O. Berg =Myrciaria floribunda 

Myrciaria quitarensis (Benth.) O. Berg =Myrcia quitarensis 

Myrciaria uliginosa O. Berg =Myrciaria floribunda 

Myrciaria verticillata O. Berg =Myrciaria floribunda 

Myrtus alternifolia Gleason =Calycolpus alternifolius 

Myrtus biflora L. =Eugenia biflora 

Myrtus calophylla Kunth =Calycolpus calophyllus 

Myrtus citrifolia Aubl. =Myrcia citrifolia 

Myrtus clusiifolia Kunth =Myrcia clusiifolia 

Myrtus cumini L. =Syzygium cumini 

Myrtus deflexa (Poir.) Kunth =Myrcia deflexa 

Myrtus emarginata Kunth =Eugenia emarginata 

Myrtus erythroxyloides Kunth =Pseudanamomis umbellulifera 

Myrtus fragrans Sw. =Myrcianthes fragrans 

Myrtus goetheana DC. =Calycolpus goetheanus 

Myrtus latifolia (O. Berg) V.M. Badillo =Myrcia calycampa 

Myrtus ligustrina Sw. =Eugenia ligustrina 

Myrtus longifolia Kunth (Melastomataceae) =Mouriri longifolia 

Myrtus monticola Sw. =Eugenia monticola 

Myrtus myricoides Kunth =Ugni myricoides 

Myrtus myricoides var. roraimensis (N.E. Br.) Steyerm. =Ugni myricoides 

Myrtus myricoides var. stenophylla (Oliv.) Steyerm. =Ugni myricoides 

Myrtus myricoides var. turumiquirensis Steyerm. =Ugni myricoides 

Myrtus nummularia Poir =Myrteola nummularia 

Myrtus pubescens Kunth =Eugenia pubescens 

Myrtus punicifolia Kunth =Eugenia punicifolia 

Myrtus rhopaloides Kunth =Myrcianthes rhopaloides 

Myrtus roraimensis N.E. Br. =Ugni myricoides 

Myrtus salutaris Kunth =Psidium salutare 

Myrtus splendens Sw. =Myrcia splendens 

Myrtus stenophylla Oliv. =Ugni myricoides 

Myrtus sylvatica G. Mey. =Myrcia sylvatica 

Myrtus umbellata Desv. ex Ham. (Melastomataceae) =Mouriri guianensis 

Myrtus umbellulifera Kunth =Pseudanamomis umbellulifera 

Myrtus umbraticola Kunth =Marlierea umbraticola 

Plinia dussii (Krug & Urb.) Urb. =Siphoneugena dussii 

Plinia fruticosa Steyerm. =Siphoneugena dussii 

Psidium aquaticum Benth. =Psidium striatulum 

Psidium araca Raddi =Psidium guineense

442 


Psidium aromaticum Aubl. =Campomanesia aromatica 

Psidium australe Cambess. =Psidium cinereum 

Psidium benthamianum O. Berg =Psidium guineense 

Psidium dubium Kunth =Myrciaria dubia 

Psidium glaucescens O. Berg =Psidium laruotteanum 

Psidium grandiflorum Aubl. =Campomanesia grandiflora 

Psidium guianense sensu Amshoff, not Pers. =Psidium acutangulum 

Psidium guildingianum Griseb. =Marlierea guildingiana 

Psidium guyanense Pers. =Psidium guineense 

Psidium ovatifolium O. Berg =Psidium densicomum 

Psidium ovatifolium var. glabrum Amshoff =Psidium densicomum 

Psidium parviflorum Benth. =Psidium striatulum 

Psidium persoonii McVaugh [If distinct, known from BO, GU, SU, FG] =Psidium acutangulum 

Psidium quinquedentatum Amshoff =Psidium laruotteanum 

Psidium sprucei O. Berg =Psidium guineense 

Psidium submetrale McVaugh =Psidium cinereum 

Siphoneugena densiflora auct. non O. Berg 1857: McVaugh =Siphoneugena dussii 

Siphoneugena densiflora var. salicifolia McVaugh =Siphoneugena dussii 

Siphoneugena densiflora var. salicifolia McVaugh =Siphoneugena dussii 

Siphoneugena densiflora var. tepuiensis Steyerm. =Siphoneugena dussii 

Siphoneugena dussii var. salicifolia (McVaugh) Proença =Siphoneugena dussii 

Stenocalyx patrisii (Vahl) O. Berg =Eugenia patrisii 

Ugni angustifolia Burret =Ugni myricoides 

Ugni myricoides f. bifaria McVaugh =Ugni myricoides 

Ugni myricoides f. grossa McVaugh =Ugni myricoides 

Ugni myricoides f. oligandra McVaugh =Ugni myricoides 

Ugni myricoides var. roraimensis (N.E. Br.) McVaugh =Ugni myricoides 

Ugni myricoides f. stenophylla (Oliv.) McVaugh =Ugni myricoides 

Ugni roraimensis (N.E. Br.) Burret =Ugni myricoides 

Ugni stenophylla (Oliv.) Burret =Ugni myricoides 

Ugni vaccinioides Burret =Ugni myricoides 

REFERENCES 

Amshoff, G.J.H. 1951. Myrtaceae. In A. Pulle, Flora of Suriname 3(2): 56-158. 

Holst, B.K. & M.L. Kawasaki. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 539-551. 

Holst, B.K., L, Landrum, & F. Grifo. 2003. Myrtaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 

Venezuelan Guayana 7:1-99. 

Kawasaki, M.L. & B.K. Holst. 1994. New species and a new combination in Myrtaceae from northeastern South America. 

Brittonia 46(2): 137-143. 

Landrum, L.R. 1986. Campomanesia, Pimenta, Blepharocalyx, Legrandia, Acca, Myrrhinium, and Luma. Flora Neotropica 

Monogr. 45: 1-178. 

McVaugh, R. 1958. Myrtaceae (Calyptranthes and Marlierea). In B. Maguire, J.J. Wurdack, & collaborators, Botany of the 

Guayana Highland part 3. Mem. New York Bot. Gard. 10(1): 1-156. 

McVaugh, R. 1969. Myrtaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 8. Mem. New York Bot. 

Gard. 18(2): 1-290. 

Sobral, M. 1993. Sinopse de Myrciaria. Napaea 9: 13-41. 

Nelumbo nucifera Gaertn. [cultivated, naturalized] 

NELUMBONACEAE 

REFERENCES 

Cramer, J. 1979. Nelumbonaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 390-392. 

Boerhavia coccinea Mill. 

Boerhavia diffusa L. 

Bougainvillea buttiana Holttum & Standl. 

Bougainvillea spectabilis Willd. 

NYCTAGINACEAE 

G.A. Aymard C., R.A. DeFilipps†, J.A. Steyermark† 

GU SU FG? 

AM? BO 


BO 

BO

Bougainvillea spp. [several species cultivated] 

Guapira amacurensis Steyerm. 

Guapira bolivarensis Steyerm. 

Guapira cuspidata (Heimerl) Lundell 

Guapira eggersiana (Heimerl) Lundell 

Guapira ferruginea (Klotsch ex Choisy) Lundell 

Guapira fragrans (Dum.-Cours.) Little 

Guapira glabriflora Steyerm. 

Guapira kanukuensis (Standl.) Lundell 

Guapira marcano-bertii Steyerm. 

Guapira microphylla (Heimerl) Lundell 

Guapira neblinensis Maguire & Steyerm. 

Guapira rusbyana (Heimerl ex Standl.) Lundell 

Guapira salicifolia (Heimerl) Lundell 

Guapira sancarlosiana Steyerm. 

Guapira sipapoana Steyerm. 

Mirabilis jalapa L. [cultivated, escaping and naturalized] 

Neea amaruayensis Steyerm. 

Neea bernardii Steyerm. 

Neea bracteosa Steyerm. 

Neea brevipedunculata Steyerm. 

Neea cedenensis Steyerm. 

Neea clarkii Steyerm. 

Neea constricta Spruce ex J.A. Schmidt 

Neea davidsei Steyerm. 

Neea floribunda Poepp. & Endl. 

Neea grandis Maguire & Steyerm. 

Neea guaiquinimae Steyerm. 

Neea huachamacarae Steyerm. 

Neea ignicola Steyerm. 

Neea liesneri Steyerm. 

Neea mapourioides Steyerm. 

Neea marahuacae Steyerm. 

Neea mollis Spruce ex J.A. Schmidt 

Neea neblinensis Maguire & Steyerm. 

Neea obovata Spruce ex Heimerl 

Neea ovalifolia Spruce ex J.A. Schmidt 

Neea parimensis Steyerm. 

Neea robusta Steyerm. 

Neea sebastianii Steyerm. 

Neea spruceana Heimerl 

Neea subglabrata Steyerm. 

Neea tepuiensis Steyerm. 

Neea tristis Heimerl 

Pisonia aculeata L. 

Pisonia macranthocarpa (Donn. Sm.) Donn. Sm. 



Boerhavia caribaea Jacq. =Boerhavia coccinea 

Boerhavia paniculata Rich. =Boerhavia diffusa 

Boerhavia surinamensis Miq. =Boerhavia diffusa 

Guapira ayacuchae Steyerm. =Guapira cuspidata 

Guapira davidsei Steyerm. =Guapira ferruginea 

Guapira glabra (Heimerl) Steyerm. =Guapira eggersiana 

Guapira graciliflora sensu Mennega et al. (1988), non (Mart.) Lundell =Guapira salicifolia 

Guapira guianensis Aubl. ? =Guapira eggersiana 

Guapira heimerliana (Standl.) Lundell =Guapira salicifolia 

Guapira olfersiana (Link, Klotzsch & Otto) Lundell [not in flora area] 

Guapira pacurero (Kunth) Little [not in flora area] 

Guapira riedeliana (Fisch. ex Heimerl) Lundell [not in flora area] 

Guapira schomburgkiana (Heimerl) Lundell =Guapira cuspidata 

Neea cauliflora Heimerl =Neea floribunda 

Neea nigricans (Sw.) Heimerl [not in flora area] 

Pisonia aculeata var. macranthocarpa Donn. Sm. =Pisonia macranthocarpa 

BO 

BO 



BO DA 

AM BO 

AM 

GU 

BO DA 

BO 

AM 

BO DA 

GU SU FG 

AM 

AM 


BO 

BO 

BO 

AM BO 

BO 

AM 

GU FG 

BO DA 


AM 

BO 

AM 

AM 

AM 

AM 

AM 

GU FG 

AM 

AM 

AM GU SU FG 

AM 

AM 

AM 

GU FG 

AM BO GU 

BO 

AM 

AM BO 

GU

444 


Pisonia albiflora Heimerl =Guapira salicifolia 

Pisonia cuspidata Heimerl =Guapira cuspidata 

Pisonia eggersiana Heimerl =Guapira eggersiana 

Pisonia eucalyptifolia Kunth =Guapira fragrans 

Pisonia ferruginea Klotzsch ex Choisy =Guapira ferruginea 

Pisonia fragrans Dum.-Cours. =Guapira fragrans 

Pisonia glabra Heimerl =Guapira eggersiana 

Pisonia guianensis (Aubl.) R.A. Howard ? =Guapira eggersiana 

Pisonia inermis Jacq. [not in flora area] =Guapira fragrans 

Pisonia microphylla Heimerl =Guapira microphylla 

Pisonia nitida Mart. ex J.A. Schmidt =Guapira eggersiana 

Pisonia olfersiana sensu Maguire (1948), non Link, Klotzsch & Otto =Guapira eggersiana 

Pisonia pacurero sensu Pulle (1906), non Kunth =Guapira eggersiana 

Pisonia salicifolia Heimerl =Guapira salicifolia 

Pisonia schomburgkiana Heimerl =Guapira cuspidata 

Torrubia cepalantha Standl. ex Pittier =Guapira ferruginea 

Torrubia cuspidata (Heimerl) Standl. =Guapira cuspidata 

Torrubia ferruginea (Klotsch ex Choisy) Standl. =Guapira ferruginea 

Torrubia fragrans (Dum.-Cours.) Standl. =Guapira fragrans 

Torrubia heimerliana Standl. =Guapira salicifolia 

Torrubia kanukuensis Standl. =Guapira kanukuensis 

Torrubia laurina Standl. =Guapira eggersiana 

Torrubia microphylla (Heimerl) Standl. =Guapira microphylla 

Torrubia olfersiana (Link, Klotzsch & Otto) Standl. [not in flora area] 

Torrubia rusbyana Heimerl ex Standl. =Guapira rusbyana 

Torrubia salicifolia (Heimerl) Standl. =Guapira salicifolia 

Torrubia schomburgkiana (Heimerl) Standl. =Guapira cuspidata 

REFERENCES 

DeFilipps, R.A. & S.L. Maina. 2002. Nyctaginaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. 


York Bot. Gard. 76 (2): 551-553. 

DeFilipps, R.A. & S.L. Maina. 2003. Phytolaccaceae, (+ remarks on Achatocarpaceae), Nyctaginaceae, Aizoaceae, 

Chenopodiaceae, Amaranthaceae, Portulacaceae, Basellaceae, Molluginaceae, Caryophyllaceae. In M.J. Jansen-Jacobs, ed., 

Flora of the Guianas, Fascicle 22 (series A). Royal Botanic Gardens, Kew, U.K. 

Steyermark, J.A. 1987. Nyctaginaceae. In Flora of the Venezuelan Guayana - III. Ann. Missouri Bot. Gard. 74(3): 614-635. 

Standley, P.C. 1931. The Nyctaginaceae of northwestern South America. Field Mus. Bot. 11: 73-114. 

Steyermark J.A. & G.A. Aymard C. 2003. Nyctaginaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Nymphaea amazonum Mart. & Zucc. 

Nymphaea amazonum subsp. amazonum 

Nymphaea ampla (Salisb.) DC. 

Nymphaea conardii Wiersema 

Nymphaea gardneriana Planch. 

Nymphaea glandulifera Rodschied 

Nymphaea jamesoniana Planch. 

Nymphaea lingulata Wiersema 

Nymphaea lotus L. [cultivated, naturalized] 

Nymphaea odorata Aiton 

Nymphaea potamophila Wiersema 

Nymphaea pulchella DC. 

Nymphaea rubra Roxb. ex Salisb. [cultivated, naturalized] 

Nymphaea rudgeana G. Mey. 

Nymphaea tenerinervia Casp. 

Victoria amazonica (Poepp.) Sowerby 

NYMPHAEACEAE 

J.H. Wiersema 


Castalia blanda (G. Mey.) G. Lawson =Nymphaea glandulifera 

Castalia spp. =Nymphaea spp. 

Leuconymphaea amazonum (Mart. & Zucc.) Kuntze =Nymphaea amazonum 

DA GU SU FG 

DA GU SU FG 

GU SU 

BO GU 

BO GU 


GU 

GU 

GU 

GU SU 

AM BO DA GU 

DA GU SU FG 

GU SU 


GU? 

GU


Leuconymphaea blanda (G. Mey.) Kuntze =Nymphaea glandulifera 

Nymphaea ampla auct. non (Salisb.) DC.: sensu South American authors =Nymphaea pulchella 

Nymphaea blanda G. Mey. [nom. superfl.] =Nymphaea glandulifera 

Nymphaea blanda var. amazonum (Mart. & Zucc.) Planch. =Nymphaea amazonum 

Nymphaea odorata var. gigantea Tricker =Nymphaea odorata 

Nymphaea parkeriana Lehm. =Nymphaea odorata 

Nymphaea rudgeana var. amazonum (Mart. & Zucc.) Griseb. =Nymphaea amazonum 

Victoria regia Lindl. =Victoria amazonica 

REFERENCES 

(see also Cabombaceae, Ceratophyllaceae, Nelumbonaceae) 

Conard, H.S. 1905. The Waterlilies: A Monograph of the Genus Nymphaea. Carnegie Institution of Washington. 

Cramer, J. 1979. Nymphaeaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 370-384. 

Mori, S.A. 2002. Nymphaeaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


554. 

Wiersema, J.H. 1987. A monograph of Nymphaea subgenus Hydrocallis. Syst. Bot. Monogr. 16: 1-112. 

Wiersema, J.H. 2003. Nymphaeaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 7: 118-124. 

Adenanthe bicarpellata Maguire, Steyerm. & Wurdack 

Adenanthe ciliata Sastre 

Adenarake macrocarpa Sastre 

Adenarake muriculata Maguire & Wurdack 

Blastemanthus gemmiflorus (Mart.) Planch. 

Blastemanthus gemmiflorus subsp. gemmiflorus 

Blastemanthus gemmiflorus subsp. sprucei (Tiegh.) Sastre 

Cespedesia spathulata (Ruiz & Pav.) Planch. 

Elvasia brevipedicellata Ule 

Elvasia calophyllea DC. 

Elvasia canescens (Tiegh.) Gilg 

Elvasia elvasioides (Planch.) Gilg 

Elvasia essequibensis Engl. 

Elvasia macrostipularis Sastre & Lescure 

Elvasia quinqueloba Spruce ex Engl. 

Elvasia sphaerocarpa R.S. Cowan 

Ouratea acuminata (DC.) Engl. 

Ouratea angulata Tiegh. 

Ouratea aquatica (Kunth) Engl. 

Ouratea arbobrevicalyx Sastre 

Ouratea articulata Sastre 

Ouratea asisae Maguire & Steyerm. 

Ouratea attenuata Tiegh. 

Ouratea bipartita Sastre 

Ouratea brevicalyx Maguire & Steyerm. 

Ouratea brevipedicellata Maguire & Steyerm. 

Ouratea candollei (Planch.) Tiegh. 

Ouratea cardiosperma (Lam.) Engl. 

Ouratea castaneifolia (DC.) Engl. 

Ouratea cataractarum Sandwith 

Ouratea cerebroidea Sastre 

Ouratea cernuiflora Sandwith 

Ouratea chaffanjonii (Tiegh.) Sastre 

Ouratea clarkii Sastre 

Ouratea coccinea Mart. ex Engl. 

Ouratea croizatii Maguire & Steyerm. 

Ouratea culminicola Maguire & Steyerm. 

Ouratea davidsii Sastre 

Ouratea decagyna Maguire 

Ouratea deminuta Maguire & Steyerm. 

Ouratea discophora Ducke 

OCHNACEAE 

C. Sastre 

BO GU 

GU 

AM 

AM 

AM GU 

AM 

AM 

GU 

AM BO DA GU 

BO GU 

AM 

BO? GU? 


GU SU 

FG 

AM GU 

GU 

GU? 

AM 

AM 

AM 

AM 

BO 

BO 

DA GU? SU FG 

AM 

AM 

AM 

GU 

GU 

SU FG 

GU? SU FG 

AM BO DA GU 

GU 

SU 

GU 

GU 

FG 

AM 

AM 

AM 

AM 

BO 

BO 

BO 

AM 

GU? SU FG 

AM BO

446 

Ouratea duidae Steyerm. 

Ouratea engleri Tiegh. 

Ouratea erecta Sastre 

Ouratea evoluta Maguire & Steyerm. 

Ouratea fasciculata Maguire & Steyerm. 

Ouratea ferruginea Engl. 

Ouratea francinae Sastre 

Ouratea frontium Sleumer 

Ouratea fusiformis Sastre 

Ouratea gigantophylla (Ehrh.) Engl. 

Ouratea gillyana (Dwyer) Sandwith & Maguire 

Ouratea grandiflora (DC.) Engl. 

Ouratea grosourdyi (Tiegh.) Steyerm. 

Ouratea guaiquinimensis Sastre 

Ouratea guianensis Aubl. 

Ouratea guildingii (Planch.) Urb. 

Ouratea guriensis Sastre 

Ouratea heterobracteata Sastre 

Ouratea huberi Maguire & Steyerm. 

Ouratea impressa (Tiegh.) Lemée 

Ouratea kanukuensis Sastre 

Ouratea lajaensis Sastre 

Ouratea leblondii (Tiegh.) Lemée 

Ouratea leprieurii Tiegh. 

Ouratea liesneri Sastre 

Ouratea longifolia (Lam.) Engl. 

Ouratea longistyla Maguire & Steyerm. 

Ouratea maasorum Sastre 

Ouratea macrocarpa Sastre 

Ouratea maguirei Sastre 

Ouratea maigualidae Sastre 

Ouratea marahuacensis Maguire & Steyerm. 

Ouratea mazaruniensis A.C. Sm. & Dwyer 

Ouratea medinae Maguire & Steyerm. 

Ouratea megaphylla Sastre 

Ouratea melinonii (Tiegh.) Lemée 

Ouratea microcalyx (Engl.) Sastre 

Ouratea multibracteata Steyerm. ex Sastre 

Ouratea obovata Sastre 

Ouratea occultinervis Sastre 

Ouratea oligantha Steyerm. ex Sastre 

Ouratea olivaeformis (A. St.-Hil.) Engl. [unverified name] 

Ouratea orisina Sastre 

Ouratea ornata Maguire & Steyerm. 

Ouratea papillata Maguire & Steyerm. 

Ouratea paratatei Sastre 

Ouratea paruensis Maguire & Steyerm. 

Ouratea pendula (Poepp.) Engl. 

Ouratea pisiformis Engl. 

Ouratea polyantha (Triana & Planch.) Engl. 

Ouratea pseudoguildingii Sastre 

Ouratea pseudotatei Maguire & Steyerm. 

Ouratea ptaritepuiensis Steyerm. 

Ouratea pulverulenta Sastre 

Ouratea ramosissima Maguire & Steyerm. 

Ouratea rigida Engl. 

Ouratea riparia Sleumer 

Ouratea roraimae Engl. 

Ouratea rorida Sastre 

Ouratea rotundipetala Maguire & Steyerm. 

Ouratea rupununiensis Klotzsch ex Engl. 

Ouratea saldariagae Sastre 

Ouratea sastrei Zarucchi, nom. nov. ined. 

Ouratea saülensis Sastre 


AM 

AM FG 

SU FG 

AM 

GU 

AM BO GU FG 

FG 

GU 

AM 

SU 

GU 

AM 

BO GU 

BO 


GU 

BO 

AM 

AM 

FG 

GU 

AM 


FG 

AM 

GU FG 

AM 

GU 

GU SU FG 

AM 

AM 

AM 

GU 

AM 

AM 

SU FG 

GU 

AM 

AM 

SU FG 

BO 

GU 

BO 

AM 

AM 

BO 

AM 

SU FG 

AM BO GU 

AM BO 

BO 

GU 

BO 

AM 

AM BO 

BO GU 

SU FG 

AM BO GU 

AM BO 

AM 

GU 

AM 

GU 

FG


Ouratea schomburgkii (Planch.) Engl. 

Ouratea scottii Sastre 

Ouratea scottii subsp. scottii 

Ouratea sculpta (Tiegh.) Sastre 

Ouratea sipapoensis Maguire & Steyerm. 

Ouratea soderstromii Sastre 

Ouratea spruceana Engl. 

Ouratea squamata Sastre 

Ouratea steyermarkii Sastre 

Ouratea subamplexicaulis Maguire & Steyerm. 

Ouratea superba Engl. 

Ouratea tatei Gleason 

Ouratea thyrsoidea Engl. 

Ouratea timehriensis Sastre 

Ouratea vaccinioides (A. St.-Hil. & Tul.) Engl. 

Ouratea verruculosa Engl. 

Ouratea yapacanae Sastre 

Perissocarpa steyermarkii (Maguire) Steyerm. & Maguire 

Perissocarpa umbellifera Steyerm. & Maguire 

Philacra auriculata Dwyer 

Philacra duidae (Gleason) Dwyer 

Philacra longifolia (Gleason) Dwyer 

Philacra steyermarkii Maguire 

Poecilandra pumila Steyerm. 

Poecilandra retusa Tul. 

Poecilandra retusa var. retusa 

Poecilandra retusa var. sclerophylla (Ule) Sastre 

Sauvagesia aliciae Sastre 

Sauvagesia aliciae subsp. aliciae 

Sauvagesia aliciae subsp. aratayensis Sastre 

Sauvagesia amoena Ule 

Sauvagesia angustifolia Ule 

Sauvagesia deflexifolia Gardner 

Sauvagesia elata Benth. 

Sauvagesia erecta L. 

Sauvagesia erecta var. erecta 

Sauvagesia erioclada Maguire & K.D. Phelps 

Sauvagesia falcisepala Sastre 

Sauvagesia fruticosa Mart. 

Sauvagesia guianensis (Eichler) Sastre 

Sauvagesia guianensis subsp. guianensis 

Sauvagesia imthurniana (Oliv.) Dwyer 

Sauvagesia linearifolia A. St.-Hil. 

Sauvagesia linearifolia subsp. venezuelensis Maguire & Wurdack 

Sauvagesia longifolia Eichler 

Sauvagesia longipes Steyerm. 

Sauvagesia nudicaulis Maguire & Wurdack 

Sauvagesia pulchella Planch. 

Sauvagesia ramosa (Gleason) Sastre 

Sauvagesia ramosissima Spruce ex Eichler 

Sauvagesia roraimensis Ule 

Sauvagesia rubiginosa A. St.-Hil. 

Sauvagesia sprengelii A. St.-Hil. 

Sauvagesia tafelbergensis Sastre 

Sauvagesia tenella Lam. 

Tyleria apiculata Sastre 

Tyleria aristata Maguire & Wurdack 

Tyleria breweriana Steyerm. 

Tyleria floribunda Gleason 

Tyleria grandiflora Gleason 

Tyleria linearis Gleason 

Tyleria pendula Maguire & Wurdack 

Tyleria phelpsiana Maguire & Steyerm. 

Tyleria silvana Maguire 

AM 

GU SU 

GU SU FG 

GU SU FG 

GU SU FG 

BO GU 

AM BO GU SU? 

BO 

AM 

BO 

AM BO GU SU 

BO GU 

AM 

GU 

GU? 

FG 

AM 

AM GU 

AM BO 

AM 

AM 

AM 

BO 

BO GU 

AM BO GU 

AM BO GU 

BO 

AM FG 

AM 

FG 

BO GU? 

BO GU SU 

AM 




AM 

AM 

AM BO 

AM BO GU 

AM BO GU 

AM BO GU 

AM 

AM 

AM BO GU FG 

BO GU 

AM 

GU SU 

AM BO 


AM BO GU SU 


BO GU SU FG 

SU FG 


AM BO 

AM 

BO 

AM BO 

AM 

AM 

AM 

BO GU 

AM

448 

Tyleria spathulata Gleason 

Tyleria spectabilis Maguire & Wurdack 

Tyleria terrae-humilis Sastre 

Tyleria tremuloidea Maguire & Wurdack 

Wallacea insignis Spruce ex Benth. & Hook. f. 

Wallacea multiflora Ducke 



Blastemanthus densiflorus Hallier =Blastemanthus gemmiflorus subsp. sprucei 

Blastemanthus paniculatus Tiegh. =Blastemanthus gemmiflorus subsp. gemmiflorus 

Blastemanthus sprucei Tiegh. =Blastemanthus gemmiflorus subsp. sprucei 

Camptouratea leblondii Tiegh. =Ouratea leblondii 

Camptouratea sagotii Tiegh. =Ouratea leblondii 

Cercouratea chaffanjonii Tiegh. =Ouratea chaffanjonii 

Cercouratea ferruginea (Engl.) Tiegh. =Ouratea ferruginea 

Cercouratea grosourdyi Tiegh. =Ouratea grosourdyi 

Cercouratea laxa Tiegh. =Ouratea superba 

Cespedesia amazonica Huber =Cespedesia spathulata 

Cespedesia bonplandii Goudot =Cespedesia spathulata 

Cespedesia sprucei Tiegh. =Cespedesia spathulata 

Elvasia caurensis Pittier =Elvasia elvasioides 

Elvasia hostmannia Planch. =Elvasia elvasioides 

Elvasia sagotii (Tiegh.) Lemée =Elvasia elvasioides 

Elvasia schomburgkii Tiegh. =Elvasia calophyllea 

Godoya gemmiflora Mart. =Blastemanthus gemmiflorus 

Godoya spathulata Ruiz & Pav. =Cespedesia spathulata 

Gomphia aquatica Kunth =Ouratea aquatica 

Gomphia candollei Planch. =Ouratea candollei 

Gomphia castaneifolia DC. =Ouratea castaneifolia 

Gomphia grandiflora DC. =Ouratea grandiflora 

Gomphia polyantha Triana & Planch. =Ouratea polyantha 

Gomphia surinamensis Planch. =Ouratea cardiosperma 

Hostmannia elvasioides Planch. =Elvasia elvasioides 

Hostmannia sagotii Tiegh. =Elvasia elvasioides 

Kaieteurea gillyana Dwyer =Ouratea gillyana 

Leitgebia colombiana R. Schult. =Sauvagesia fruticosa 

Leitgebia gleasoniana Lasser =Sauvagesia guianensis subsp. guianensis 

Leitgebia guianensis Eichler =Sauvagesia guianensis 

Leitgebia imthurniana Oliv. =Sauvagesia imthurniana 

Luxemburgia duidae Gleason =Philacra duidae 

Luxemburgia longifolia Gleason =Philacra longifolia 

Ouratea ayacuchae Maguire & Steyerm. =Ouratea chaffanjonii 

Ouratea coccinea auct. non Engl. 1876: sensu Maguire & Steyerm. =Ouratea fusiformis p.p., Ouratea maguirei p.p. 

Ouratea crenata Spruce ex Tiegh. =Ouratea pisiformis 

Ouratea discophora subsp. pervenulosa Maguire & Steyerm. =Ouratea discophora 

Ouratea elongata Sastre [non O. elongata (Oliv.) Engl.] =Ouratea sastrei 

Ouratea guianensis auct. non Aubl. 1775: sensu Maguire & Steyerm. =Ouratea rorida p.p. 

Ouratea inconformis Maguire & Steyerm. =Ouratea superba 

Ouratea integrifolia (DC.) Sagot [nomen nudum] 

Ouratea inundata Spruce ex Engl. =Ouratea superba 

Ouratea longifolia auct. non (Lam.) Engl.: sensu Maguire & Steyerm. =Ouratea guianensis 

Ouratea macurucoensis Maguire & Steyerm. =Ouratea steyermarkii 

Ouratea odora Poepp. ex Engl. =Ouratea superba 

Ouratea pittieri Sleumer =Ouratea grosourdyi 

Ouratea polygyna Engl. [not in flora area] 

Ouratea propingua Maguire & Steyerm. =Ouratea ptaritepuiensis 

Ouratea pumila Maguire & Steyerm. =Ouratea aquatica 

Ouratea pumila var. heterodoxa Maguire & Steyerm. =Ouratea aquatica 

Ouratea racemiformis Ule =Ouratea schomburgkii 

Ouratea sagotii (Tiegh.) Lemée =Ouratea leblondii 

Ouratea surinamensis (Planch.) Wehlb. =Ouratea cardiosperma 

Ouratea vasivae Spruce ex Engl. =Ouratea grandiflora 

Ouratea venezuelensis Steyerm. =Ouratea thyrsoidea 

Ouratea wurdackiana Maguire & Steyerm. =Ouratea superba 

Pentaspatella ramosa Gleason =Sauvagesia ramosa 

AM 

AM 

AM 

AM 

AM 

AM


Poecilandra sclerophylla Ule =Poecilandra retusa var. sclerophylla 

Roraimanthus imthurnianus (Oliv.) Gleason =Sauvagesia imthurniana 

Roraimanthus imthurnianus (Oliv.) Gleason =Sauvagesia imthurniana 

Sauvagesia deficiens A.C. Sm. =Sauvagesia roraimensis 

Sauvagesia duckei Sleumer =Sauvagesia ramosa 

Sauvagesia duidae auct. non Steyerm. 1952: sensu Steyerm. =Sauvagesia aliciae 

Sauvagesia duidae Steyerm. =Sauvagesia erioclada 

Sauvagesia elata subsp. occidentalis Maguire & Wurdack =Sauvagesia elata 

Sauvagesia erecta Aubl. [non L.] =Sauvagesia sprengelii 

Sauvagesia erecta var. rubiginosa (A. St.-Hil.) Eichler =Sauvagesia rubiginosa 

Sauvagesia erioclada var. grandiflora Maguire & Wurdack =Sauvagesia erioclada 

Sauvagesia guianensis subsp. gleasoniana (Lasser) Steyerm. =Sauvagesia guianensis subsp. guianensis 

Sauvagesia guianensis subsp. guaiquinimensis Steyerm. =Sauvagesia guianensis subsp. guianensis 

Sauvagesia guianensis subsp. sipapoensis Steyerm. =Sauvagesia guianensis subsp. guianensis 

Sauvagesia imthurniana subsp. chimantensis Maguire, Steyerm. & Wurdack =Sauvagesia imthurniana 

Sauvagesia inconspicua Dwyer =Sauvagesia tenella 

Sauvagesia kappleri Miq. =Sauvagesia sprengelii 

Sauvagesia laxa Mart. =Sauvagesia rubiginosa 

Sauvagesia marahuacensis Steyerm. =Sauvagesia guianensis subsp. guianensis 

Sauvagesia miniata Steyerm. =Sauvagesia ramosissima 

Sauvagesia rosacea Ule =Sauvagesia fruticosa 

Sauvagesia salicifolia Ging. =Sauvagesia erecta 

Sauvagesia serpyllifolia Mart. =Sauvagesia sprengelii 

Sauvagesia smithiana Dwyer =Sauvagesia roraimensis 

Sauvagesia striata Gleason & Dwyer =Sauvagesia angustifolia 

Sauvagesia surinamensis Miq. =Sauvagesia rubiginosa 

Trichovaselia canescens Tiegh. =Elvasia canescens 

Vaselia quinqueloba (Spruce ex Engl.) Tiegh. =Elvasia quinqueloba 

Villouratea spiciformis Tiegh. =Ouratea rigida 

REFERENCES 

Allorge, L. & C. Sastre. 1990. Distribution géographique dans les Guyanes, d’Apocynaceae et d’Ochnaceae. Essai de mise en 

évidence de centres de spéciation. C.R. Soc. Biogéogr. 67(2): 77-94. 

Dwyer, J.D. 1943. The taxa of the monogeneric tribe Elvasieae (Ochnaceae). Bull. Torrey Bot. Club 70: 42-49. 

Dwyer, J.D. 1945. The taxonomy of the genus Sauvagesia (Ochnaceae). Bull. Torrey Bot. Club 72(6): 521-540. 

Maguire, B., & J.J. Wurdack. 1961. Ochnaceae. In Botany of the Guayana Highland part IV. Mem. New York Bot. Gard. 10(4): 

6-21. 

Maguire, B. & J.A. Steyermark. 1989. Ouratea in Guayana and adjacent Amazonian hylaea. In B. Maguire & collaborators, 

Botany of the Guayana Highland part XIII. Mem. New York Bot. Gard. 51: 56-102. 

Sastre, C. 1971. Distribution géographique des espèces de Sauvagesia L. C.R. Soc. Biogéogr. 421: 47-59. 

Sastre, C. 1984. Etude sur le genre Sauvagesia L. en Guyane française. Saussurea 15: 111-118. 

Sastre, C. 1986. Studies on the Flora of the Guianas 23. Cinq espèces nouvelles d’Ouratea Aublet (Ochnaceae) des Guyanes. 

Bull. Mus. Nat. Hist. ser. 4, 8, sect. B, Adansonia 4: 413-422. 

Sastre, C. 1987. Studies on the Flora of the Guianas 30. Considérations phytogéographiques sur les Ochnacées guyanaises. C.R. 

Soc. Biog. 63(3): 89-97. 

Sastre, C. 1988. Synopsis generis Ouratea Aubl. Bull. Mus. natn. Hist. nat. Paris, sér. 4, 10, sect. B, Adansonia 1: 47-67. 

Sastre, C. 1991. Duae novae species Ourateae (Ochnaceae) in Guianis collectae. Studies on the Flora of the Guianas. 59. Candollea 

46: 395-398. 

Sastre, C. 1994. Three new taxa of Ouratea (Ochnaceae) from South America. Brittonia 46(4): 309-313. 

Sastre, C. 1995. Novelties in the Neotropical genus Ouratea Aublet (Ochnaceae). Novon 5(2): 193-200. 

Sastre, C. 1998. Novelties in Sauvagesioideae (Ochnaceae) from Venezuela and Guyana. Novon 8(3): 301-303. 

Sastre, C. 2001. New Ouratea species (Ochnaceae) from Venezuela and adjacent countries. Novon 11(1):105-118. 

Sastre, C. 2001. Ochnaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 7: 

124-161. 

Sastre, C. 2002. Ochnaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


559. 

Wehlburg, C. 1937. Ochnaceae. In A. Pulle, Flora of Suriname 3(1): 328-341. 

Aptandra liriosmoides Spruce ex Miers 

Aptandra tubicina (Poepp.) Benth. ex Miers 

OLACACEAE 

P. Hiepko, J.M. MacDougal 

AM GU 

BO

450 

Brachynema axillare R. Duno & P.E. Berry 

Cathedra acuminata (Benth.) Miers 

Chaunochiton angustifolium Sleumer 

Chaunochiton kappleri (Sagot ex Engl.) Ducke 

Chaunochiton loranthoides Benth. 

Dulacia candida (Poepp.) Kuntze 

Dulacia crassa (Monach.) Sleumer [expected in BO] 

Dulacia cyanocarpa Sleumer 

Dulacia guianensis (Engl.) Kuntze 

Dulacia inopiflora (Miers) Kuntze 

Dulacia macrophylla (Benth.) Kuntze 

Dulacia redmondii Steyerm. 

Dulacia tepuiensis (Steyerm.) Sleumer & Steyerm. 

Heisteria acuminata (Bonpl.) Engl. 

Heisteria barbata Cuatrec. 

Heisteria cauliflora Sm. 

Heisteria densifrons Engl. 

Heisteria duckei Sleumer 

Heisteria insculpta Sleumer 

Heisteria maguirei Sleumer 

Heisteria maytenoides Spruce ex Engl. 

Heisteria ovata Benth. 

Heisteria pentandra (Benth. ex Reissek) Engl. 

Heisteria scandens Ducke 

Heisteria spruceana Engl. 

Maburea trinervis Maas 

Minquartia guianensis Aubl. 

Ptychopetalum olacoides Benth. 

Schoepfia brasiliensis A. DC. 

Schoepfia clarkii Steyerm. 

Schoepfia lucida Pulle 

Schoepfia tepuiensis Steyerm. 

Ximenia americana L. 

Ximenia americana var. americana 



Aptandra benthamiana Miers =Aptandra liriosmoides 

Aptandra spruceana Miers =Aptandra tubicina 

Cathedra caurensis Pittier =Cathedra acuminata 

Cathedra crassifolia Benth. ex Miers =Cathedra acuminata 

Cathedra guianensis Sandwith =Cathedra acuminata 

Chaunochiton mouriroides A.C. Sm. =Aptandra tubicina 

Chaunochiton tubicinum (Poepp.) Lemée =Aptandra tubicina 

Diplocrater acuminatus Benth. =Cathedra acuminata 

Dulacia adhaerens var. stenopoda Cuatrec. =Dulacia candida 

Heisteria acuta Engl. =Heisteria pentandra 

Heisteria coccinea Jacq. [not in flora area] 

Heisteria fatoensis Standl. =Heisteria scandens 

Heisteria flexuosa auct. non Engl. 1892: sensu V.M. Badillo =Heisteria pentandra 

Heisteria flexuosa Engl. =Heisteria ovata 

Heisteria guianensis V.M. Badillo =Heisteria acuminata 

Heisteria kappleri Sagot ex Engl. =Chaunochiton kappleri 

Heisteria longifolia Spruce ex Engl. (Ebenaceae) =Diospyros tenuiflora 

Heisteria longipes Standl. =Heisteria acuminata 

Heisteria microcalyx Sagot =Heisteria densifrons 

Heisteria microcarpa Spruce ex Engl. =Heisteria densifrons 

Heisteria nitida auct. non Spruce ex Engl.: sensu V.M. Badillo =Heisteria ovata 

Heisteria surinamensis Amshoff =Heisteria ovata 

Heisteria tubicina Poepp. =Aptandra tubicina 

Heymassoli inermis Aubl. =Ximenia americana 

Heymassoli spinosa Aubl. =Ximenia americana 

Liriosma candida Poepp. =Dulacia candida 

Liriosma crassa Monach. =Dulacia crassa 

Liriosma guianensis Engl. =Dulacia guianensis 

Liriosma inopiflora Miers [not in flora area] =Dulacia inopiflora 

AM 


AM BO 

GU SU FG 

AM 

AM BO 

GU 

AM 


AM 

AM 

AM GU 

BO 

AM BO DA 

AM FG 

GU SU FG 

AM GU SU FG 

AM 

AM 

BO DA GU 

AM 


AM 


AM 

GU 


GU SU FG 

BO 

AM 

AM SU 

BO 


AM BO GU SU FG


Liriosma macrophylla (Benth.) Roem. =Dulacia macrophylla 

Liriosma micrantha Spruce ex Engl. [nomen nudum] =Dulacia inopiflora 

Liriosma tepuiensis Steyerm. =Dulacia tepuiensis 

Olax macrophylla Benth. =Dulacia macrophylla 

Olax schomburgkii Klotzsch [nomen nudum] =Dulacia guianensis 

Rhaptostylum acuminatum Bonpl. =Heisteria acuminata 

Rhaptostylum pentandrum Benth. ex Reissek =Heisteria pentandra 

Secretania loranthacea Müll. Arg. =Minquartia guianensis 

REFERENCES 

Amshoff, G.J.H. 1938. Olacaceae. In A. Pulle, Flora of Suriname 1(1): 262-272. 

Hiepko, P. 1993. Olacaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 14: 3-35. 

Hiepko, P. 2002. Olacaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


562. 

MacDougal, J.M. 2003. Olacaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 7: 162-186. 

Sleumer, H.O. 1984. Olacaceae. Flora Neotropica Monogr. 38: 1-159. 

Chionanthus compactus Sw. 

Chionanthus guianensis (Aubl.) Pers. 

Chionanthus implicatus (Rusby) P.S. Green 

Jasminum spp. [cultivated, perhaps escaping] 

OLEACEAE 

J.S. Miller, G.A. Aymard C. 


Chionanthus caribea Jacq. =Chionanthus compactus 

Chionanthus guianensis (Aubl.) Stearn [comb. superfl.] 

Chionanthus mayepea Vahl =Chionanthus guianensis 

Chionanthus tetrandra Vahl =Chionanthus guianensis 

Linociera caribea (Jacq.) Knobl. =Chionanthus compactus 

Linociera tetrandra (Vahl) Roem. & Schult. =Chionanthus guianensis 

Mayapea implicata Rusby =Chionanthus implicatus 

Mayepea caribea (Jacq.) Kuntze =Chionanthus compactus 

Mayepea guianensis Aubl. =Chionanthus guianensis 

REFERENCES 

Green, P.S. 1994. A revision of Chionanthus in S. America and the description of Priogymnanthus, gen. nov. Kew Bull. 49(2): 

261-286. 

Miller, J.S. & G.A. Aymard C. 2003. Oleaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Stearn, W.T. 1976. Union of Chionanthus and Linociera. Ann. Missouri Bot. Gard. 63: 355-357. 

Ludwigia affinis (DC.) H. Hara 

Ludwigia decurrens Walter 

Ludwigia densiflora (Micheli) H. Hara 

Ludwigia dodecandra (DC.) Zardini & P.H. Raven 

Ludwigia elegans (Cambess.) H. Hara 

Ludwigia erecta (L.) H. Hara 

Ludwigia foliobracteolata (Munz) H. Hara 

Ludwigia helminthorrhiza (Mart.) H. Hara 

Ludwigia hyssopifolia (G. Don) Exell 

Ludwigia inclinata (L. f.) M. Gómez 

Ludwigia latifolia (Benth.) H. Hara 

Ludwigia leptocarpa (Nutt.) H. Hara 

Ludwigia mexiae (Munz) H. Hara 

Ludwigia nervosa (Poir.) H. Hara 

Ludwigia octovalvis (Jacq.) P.H. Raven 

Ludwigia peruviana (L.) H. Hara 

ONAGRACEAE 

P.H. Raven, E.M. Zardini 

AM BO 

AM 

FG 



BO DA 

GU FG 

SU 


BO DA GU FG 

BO DA GU SU 


AM BO GU SU 



SU 



BO

452 

Ludwigia quadrangularis (Micheli) H. Hara 

Ludwigia rigida (Miq.) Sandwith 

Ludwigia sedoides (Bonpl.) H. Hara 

Ludwigia sericea (Cambess.) H. Hara 

Ludwigia torulosa (Arn.) H. Hara 



Jussiaea affinis DC. =Ludwigia affinis 

Jussiaea affinis var. dodecandra (DC.) Munz =Ludwigia dodecandra 

Jussiaea affinis var. multiflora Jonker =Ludwigia affinis 

Jussiaea aluligera Miq. =Ludwigia leptocarpa 

Jussiaea decurrens (Walter) DC. =Ludwigia decurrens 

Jussiaea densiflora Micheli =Ludwigia densiflora 

Jussiaea dodecandra DC. =Ludwigia dodecandra 

Jussiaea elegans Cambess. =Ludwigia elegans 

Jussiaea erecta L. =Ludwigia erecta 

Jussiaea erecta var. sebana DC. =Ludwigia erecta 

Jussiaea foliobracteolata Munz =Ludwigia foliobracteolata 

Jussiaea helminthorrhiza Mart. =Ludwigia helminthorrhiza 

Jussiaea hexamera Miq. =Ludwigia affinis 

Jussiaea hyssopifolia G. Don =Ludwigia hyssopifolia 

Jussiaea inclinata L. f. =Ludwigia inclinata 

Jussiaea latifolia Benth. =Ludwigia latifolia 

Jussiaea leptocarpa Nutt. =Ludwigia leptocarpa 

Jussiaea leptocarpa var. aluligera (Miq.) Jonker =Ludwigia leptocarpa 

Jussiaea leptocarpa var. genuina Munz =Ludwigia leptocarpa 

Jussiaea leptocarpa var. meyeriana (Kuntze) Munz =Ludwigia leptocarpa 

Jussiaea linifolia auct. non Vahl: sensu Hutch. & Dalziel =Ludwigia erecta 

Jussiaea linifolia Vahl =Ludwigia hyssopifolia 

Jussiaea lithospermifolia Micheli =Ludwigia rigida 

Jussiaea lithospermifolia var. pubescens Munz =Ludwigia rigida 

Jussiaea lithospermifolia var. typica Munz =Ludwigia rigida 

Jussiaea maypurensis Kunth =Ludwigia nervosa 

Jussiaea mexiae Munz =Ludwigia mexiae 

Jussiaea micrantha Kunze =Ludwigia hyssopifolia 

Jussiaea mollis Kunth =Ludwigia peruviana 

Jussiaea natans Bonpl. =Ludwigia helminthorrhiza 

Jussiaea nervosa Poir. =Ludwigia nervosa 

Jussiaea nervosa var. pubescens Micheli =Ludwigia nervosa 

Jussiaea nervosa var. typica Munz =Ludwigia nervosa 

Jussiaea octonervia Lam. =Ludwigia octovalvis 

Jussiaea palustris G. Mey. =Ludwigia decurrens 

Jussiaea peruviana L. =Ludwigia peruviana 

Jussiaea peruviana var. glaberrima J.D. Sm. =Ludwigia peruviana 

Jussiaea peruviana var. typica Munz =Ludwigia peruviana 

Jussiaea pterophora Miq. =Ludwigia decurrens 

Jussiaea quadrangularis Micheli =Ludwigia quadrangularis 

Jussiaea rigida Miq. =Ludwigia rigida 

Jussiaea sedoides Bonpl. =Ludwigia sedoides 

Jussiaea suffruticosa L. =Ludwigia octovalvis 

Jussiaea surinamensis Miq. =Ludwigia leptocarpa 

Jussiaea torulosa Arn. =Ludwigia torulosa 

Jussiaea variabilis var. affinis (DC.) Kuntze =Ludwigia affinis 

Jussiaea variabilis var. meyeriana Kuntze =Ludwigia leptocarpa 

Ludwigia affinis var. dodecandra (DC.) Munz =Ludwigia dodecandra 

Ludwigia lithospermifolia (Micheli) H. Hara =Ludwigia rigida 

Ludwigia micrantha (Kunze) H. Hara =Ludwigia hyssopifolia 

Oenothera octovalvis Jacq. =Ludwigia octovalvis 

Oocarpon torulosum (Arn.) Urb. =Ludwigia torulosa 

BO GU 

AM BO GU SU 

AM BO DA GU 

GU 


REFERENCES 

Jonker, F.P. 1942. Oenotheraceae. In A. Pulle, Flora of Suriname 3(2): 13-34. 

Kawasaki, M.L. 2002. Onagraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


562-564.


Zardini, E.M. & P.H. Raven. 1991. Onagraceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 10: 1-44. 

Zardini, E.M. & P.H. Raven. 2003. Onagraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Agonandra brasiliensis Miers ex Benth. & Hook. f. 

Agonandra brasiliensis subsp. brasiliensis 

Agonandra brasiliensis subsp. racemigera Hiepko 

Agonandra silvatica Ducke 

OPILIACEAE 

P. Hiepko, J.S. Miller 

REFERENCES 

Hiepko, P. 1993. Opiliaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 14: 36-39. 

Miller, J.S. 2003. Opiliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 7: 

198-199. 

OXALIDACEAE 

G.A. Aymard C., P.E. Berry, A. Lourteig† 

Biophytum calophyllum (Progel) Guillaumin 

Biophytum cardonaei Pittier 

Biophytum kayae Aymard & P.E. Berry 

Biophytum lourteigiae Aymard & P.E. Berry 

Biophytum ottohuberi Aymard & P.E. Berry 

Oxalis barrelieri L. 

Oxalis corniculata L. [naturalized] 

Oxalis debilis Kunth 

Oxalis debilis var. corymbosa (DC.) Lourteig 

Oxalis frutescens L. 

Oxalis frutescens subsp. borjensis (Kunth) Lourteig 

Oxalis frutescens subsp. frutescens 

Oxalis juruensis Diels 

Oxalis mucronulata Norlind 

Ruptiliocarpon aff. caracolita Hammel & N. Zamora [referred to segregate 

family Lepidobotryaceae by Hammel & Zamora] 


Acetosella schomburgkiana (Progel) Kuntze =Oxalis frutescens 

Biophytum casiquiarense R. Knuth =Biophytum calophyllum 

Biophytum passargei R. Knuth =Biophytum calophyllum 

Oxalis amazonica Progel =Oxalis barrelieri 

Oxalis borjensis Kunth =Oxalis frutescens subsp. borjensis 

Oxalis calophylla Progel =Biophytum calophyllum 

Oxalis pentantha Jacq. =Oxalis frutescens 

Oxalis schomburgkiana Progel =Oxalis frutescens 

Oxalis schomburgkiana var. lasiocarpa Progel =Oxalis frutescens 

Oxalis schomburgkiana var. leiocarpa Progel =Oxalis frutescens 

Oxalis sepium A. St.-Hil. [not in flora area] 



BO 

AM GU SU FG 

AM BO 

BO GU 

AM 

AM 

BO 

BO GU SU FG 

GU SU FG 

GU 

GU 


AM BO 

BO 

AM GU SU FG 

GU 

SU 

REFERENCES 

(incl. Lepidobotryaceae) 

Aymard C., G.A. & P.E. Berry. 2003. Oxalidaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Hammel, B.E. & N.A. Zamora. 1993. Ruptiliocarpon (Lepidobotryaceae): a new arborescent genus and tropical American link 

to Africa, with a reconsideration of the family. Novon 3(4): 408-417. 

Jonker, F.P. 1942. Oxalidaceae. In A. Pulle, Flora of Suriname 3(2): 44-55. 

Lourteig, A. 1980. Oxalidaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. Gard. 

67: 823-850. 

Lourteig, A. 1994. Oxalis L. subgénero Thamnoxys (Endl.) Reiche emend. Lourt. Bradea 7(1): 1-199. 

Mitchell, J.D. 2002. Oxalidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


564-566.

454 

Argemone mexicana L. [cultivated, naturalized] 


PAPAVERACEAE 

R.W. Kiger 

REFERENCES 

Stoffers, A.L. 1979. Papaveraceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 331-334. 

Ancistrothyrsus hirtellus A.H. Gentry 

Ancistrothyrsus cf. tessmannii Harms 

Dilkea acuminata Mast. 

Dilkea johannesii Barb. Rodr. 

Dilkea parviflora Killip 

Dilkea retusa Mast. 

Dilkea sp. A Feuillet, ined. 

Dilkea sp. B Feuillet, ined. 

Dilkea wallisii Mast. 

Mitostemma glaziovii Mast. 

Mitostemma jenmanii Mast. 

Passiflora acuminata DC. 

Passiflora adenopoda DC. 

Passiflora aimae Annonay & Feuillet 

Passiflora ambigua Hemsl. 

Passiflora amicorum Wurdack 

Passiflora amoena L.K. Escobar 

Passiflora arta Feuillet, ined. 

Passiflora ascidia Feuillet 

Passiflora auriculata Kunth 

Passiflora balbis Feuillet 

Passiflora bicornis Houst. ex Mill. 

Passiflora biflora Lam. 

Passiflora bomareifolia Steyerm. & Maguire 

Passiflora candida (Poepp. & Endl.) Mast. 

Passiflora capparidifolia Killip 

Passiflora cardonae Killip 

Passiflora cauliflora Harms 

Passiflora cerasina Annonay & Feuillet 

Passiflora ceratocarpa F. Silveira 

Passiflora cirrhiflora Juss. 

Passiflora citrifolia (Juss.) Mast. 

Passiflora coccinea Aubl. 

Passiflora compar Feuillet, ined. 

Passiflora coriacea Juss. 

Passiflora costata Mast. 

Passiflora crenata Feuillet & Cremers 

Passiflora cuneata Willd. 

Passiflora cyanea Mast. 

Passiflora deficiens Mast. 

Passiflora edulis Sims 

Passiflora edulis f. flavicarpa Degener [cultivated, escaping] 

Passiflora exura Feuillet 

Passiflora fanchonae Feuillet 

Passiflora filipes Benth. 

Passiflora foetida L. 

Passiflora foetida var. foetida 

Passiflora foetida var. gossypiifolia (Desv. ex Ham.) Mast. 

Passiflora foetida var. hispida (DC.) Killip 

Passiflora foetida var.. orinocensis (Killip) Feuillet, ined. 

Passiflora fuchsiiflora Hemsl. 

Passiflora gabriellana Vanderplank 

Passiflora garckei Mast. 

PASSIFLORACEAE 

C. Feuillet, S.S. Tillett 

SU FG 

AM 

GU FG 

AM BO DA FG 

FG 

GU 

SU 

SU FG 

GU 

GU 

GU 

GU 

FG 

BO? GU FG 

DA 

FG 

AM 

BO GU 

GU 

GU 

SU FG 

BO GU 


AM BO 

BO 

DA 

GU 

AM BO 

BO GU SU FG 

BO GU 

AM 

AM 

BO GU 

SU FG 

GU FG 

BO GU SU FG 

FG 


GU FG 

DA GU 


FG 

BO DA 

BO DA 

GU 



FG 

BO? 

GU FG 



BO DA 


AM BO DA 

GU SU FG 

FG 

BO GU SU FG


Passiflora glandulosa Cav. 

Passiflora gleasonii Killip 

Passiflora gracilis J. Jacq. ex Link 

Passiflora guazumaefolia Juss. 

Passiflora holtii Killip 

Passiflora involucrata (Mast.) A.H. Gentry 

Passiflora kawensis Feuillet 

Passiflora laurifolia L. 

Passiflora leptopoda Harms 

Passiflora longicuspis Vanderplank & S. Vanderplank 

Passiflora longiracemosa Ducke 

Passiflora maguirei Killip 

Passiflora maliformis L. [cultivated] 

Passiflora micropetala Mast. 

Passiflora misera Kunth 

Passiflora moritziana Planch. [expected in GU] 

Passiflora multiformis Jacq. 

Passiflora nitida Kunth 

Passiflora nuriensis Steyerm. 

Passiflora ovata Martin ex DC. 

Passiflora pachyantha Killip 

Passiflora pedata L. 

Passiflora pedata var. pedata 

Passiflora pedata var. stipularis (Killip) Killip 

Passiflora phaeocaula Killip 

Passiflora phellos Feuillet 

Passiflora picturata Ker Gawl. 

Passiflora plumosa Feuillet & Cremers 

Passiflora pulchella Kunth 

Passiflora pyrrhantha Harms 

Passiflora quadrangularis L. [cultivated or escaped from cultivation] 

Passiflora quadriglandulosa Rodschied 

Passiflora quelchii N.E. Br. 

Passiflora retipetala Mast. 

Passiflora riparia Mart. ex Mast. 

Passiflora rubra L. 

Passiflora rufostipulata Feuillet 

Passiflora saulensis Feuillet 

Passiflora sclerophylla Harms 

Passiflora securiclata Mast. 

Passiflora seemannii Griseb. 

Passiflora serrato-digitata L. 

Passiflora serrulata Jacq. 

Passiflora sp. A 

Passiflora sp. B 

Passiflora sp. C 

Passiflora sp. D Feuillet & J.M. MacDougal, ined. 

Passiflora sp. E Feuillet, ined. 

Passiflora sp. F Feuillet, ined. 

Passiflora sp. G Feuillet, ined. 

Passiflora sp. H Feuillet & J.M. MacDougal, sp. nov. ined. 

Passiflora spicata Mast. 

Passiflora spinosa (Poepp. & Endl.) Mast. 

Passiflora stipulata Aubl. 

Passiflora suberosa L. 

Passiflora subpeltata Ortega 

Passiflora trialata Feuillet & J.M. MacDougal 

Passiflora tuberosa Jacq. 

Passiflora variolata Poepp. & Endl. 

Passiflora vespertilio L. 

Passiflora vitifolia Kunth 


Dilkea johannesii var. parvifolia Hoehne =Dilkea acuminata 

Dilkea magnifica Steyerm. =Dilkea acuminata 


BO? GU 

BO 

BO 

AM 

AM 

GU FG 


GU SU FG 

FG 

AM BO GU 

AM GU 

BO GU 

SU 

SU FG 

BO SU FG 

AM 

AM GU SU FG 

BO 

FG 

GU SU 


GU SU 

AM 

AM 

AM 

BO GU 

BO SU 

FG 

AM BO 

AM BO 

DA GU SU FG 

AM BO DA GU 

GU 

BO GU SU FG 

AM GU 

GU SU FG 

FG 

FG 

AM BO GU 

AM 

AM 

BO GU 

GU SU FG 

AM 

AM 

AM 

BO 

BO 

GU 

GU 

GU 

SU 

FG 

FG 

FG 

FG 

BO 

FG 

BO DA 

DA 

SU 

FG 

DA GU 

AM BO DA FG 


FG?

456 


Passiflora capsularis L. [not in flora area] 

Passiflora foetida var. moritziana (Planch.) Killip =Passiflora moritziana 

Passiflora gossypifolia Desv. ex Ham. =Passiflora foetida var. gossypiifolia 

Passiflora hispida DC. ex Triana & Planch. =Passiflora foetida var. hispida 

Passiflora kegeliana Garcke =Passiflora auriculata 

Passiflora lonchophora Harms =Passiflora retipetala 

Passiflora murucuia L. [not in flora area] 

Passiflora pallida L. =Passiflora suberosa 

Passiflora pedata subsp. stipularis Killip =Passiflora pedata var. stipularis 

Passiflora retrorsa Killip =Passiflora securiclata 

Passiflora rotundifolia L. [not in flora area] 

Passiflora serratifolia L. [not in flora area] 

Tacsonia spinosa Poepp. & Endl. =Passiflora spinosa 

REFERENCES 

Escobar, L.K. 1994. Two new species and a key to Passiflora subg. Astrophea. Syst. Bot. 19(2): 203-210. 

Feuillet, C. 1986. Deux Passifloraceae nouvelles et quelques espèces rares en Guyane française. Candollea 41: 173-178. 

Feuillet, C. 1989. Diversity and distribution on Guianan Passifloraceae. In L.B. Holm-Nielsen, I.C. Nielsen & H. Balslev, 

Tropical Forests, Botanical Dynamics, Speciation and Diversity. Academic Press, London. 

Feuillet, C. 1994. Two new species of Passiflora (Passifloraceae) from French Guiana. Novon 4(3): 236-241. 

Feuillet, C. 2002. A new series and three new species of Passiflora subgenus Astrophea from the Guianas. Brittonia 54(1): 18-29. 

Feuillet, C. 2004. Passifloraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


566-570. 

Feuillet, C. 2004. Passiflora phellos, a new species of subgenus Passiflora (Passifloraceae) from northern South America. Novon 

14 (3): 285-287. 

Feuillet, C. & G. Cremers. 1984. Passifloraceae nouvelles ou méconnues de Guyane française. Proc. Kon. Ned. Akad. Wet. ser. 

C, 87: 377-386. 

Gentry, A.H. 1981. Distributional patterns and an additional species of the Passiflora vitifolia complex: Amazonian species 

diversity due to edaphically differentiated communities. Pl. Syst. Evol. 137: 95–105. 

Killip, E.P. 1937. Passifloraceae. In A. Pulle, Flora of Suriname 3(1): 306-327. 

Killip, E.P. 1938. The American species of Passifloraceae. Publ. Field Mus. Nat. Hist., Bot. Ser., 19: 1-613. 

Tillett, S.S. 1988. Passionis passifloris II. Terminología. Ernstia 48: 1–40. 

Tillett. S.S. 2003. Passifloraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

7: 625-667. 

Craniolaria annua L. [expected in SU] 

Sesamum orientale L. [cultivated & escaping] 

Sesamum indicum L. =Sesamum orientale 

PEDALIACEAE 

J.S. Miller 


REFERENCES 

(incl. Martyniaceae) 

Eseltine, G.P. van. 1929. A preliminary study of the unicorn plants (Martyniaceae). N.Y. State Ag. Expt. Sta. (Geneva) Tech. 

Bull. 149: 1-41. 

Miller, J.S. 2003. Pedaliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 7: 

667-670. 

Pulle, A. 1932. Pedaliaceae. In A. Pulle, Flora of Suriname 4(1): 111-112. 

Seegeler, C.J.P. 1989. Sesamum orientale L. (Pedaliaceae): sesame’s correct name. Taxon 38(4): 656-659. 

Peridiscus lucidus Benth. 

Whittonia guianensis Sandwith 

PERIDISCACEAE 

B.K. Holst 

AM BO GU 


AM BO 

REFERENCES 

Holst, B.K. 2003. Peridiscaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

7: 670-671. 

Sandwith, N.Y. 1962. Contributions to the flora of tropical America: LXIX. A new genus of Peridiscaceae. Kew Bull. 15: 467- 

471. 

GU

Microtea debilis Sw. 

Microtea glochidiata Moq. [Serra Neblina, expected in AM] 

Microtea maypurensis (Kunth) G. Don 

Petiveria alliacea L. 

Phytolacca rivinoides Kunth & Bouché 

Phytolacca rugosa A. Braun & Bouché 

Phytolacca thyrsiflora Fenzl ex J.A. Schmidt 

Rivina humilis L. 

Rivina humilis var. humilis 

Seguieria aculeata Jacq. 

Seguieria americana L. 

Seguieria macrophylla Benth. 

Trichostigma octandrum (L.) H. Walter 


PHYTOLACCACEAE 

G.A. Aymard C., N.L. Cuello A., R.A. DeFilipps† 


Ancistrocarpus maypurensis Kunth =Microtea maypurensis 

Ceratoccoca maypurensis (Kunth) Willd. ex Schult. =Microtea maypurensis 

Microtea maypurensis var. angustifolia Moq. =Microtea maypurensis 

Petiveria foetida Salisb. =Petiveria alliacea 

Phytolacca americana L. [not in flora area] 

Rivina octandra L. =Trichostigma octandrum 

Seguieria cordata Britton =Seguieria macrophylla 

Seguieria foliosa Benth. =Seguieria americana 

REFERENCES 

Aymard C., G.A. & N.L. Cuello A. Phytolaccaceae. 2003. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Burger, W. 1983. Phytolaccaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212. 

DeFilipps, R.A. & S.L. Maina. 2003. Phytolaccaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. 


York Bot. Gard. 76 (2): 571-574. 

Raeder, K. 1961. Phytolaccaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. 

Gard. 48(1): 66-79. 

Rohwer, J. 1982. A taxonomic revision of the genera Seguieria Loefl. and Gallesia Casar. Mitt. Bot. Staats. München 18: 231- 

288. 

Stoffers, A.L. 1957. Phytolaccaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 209-217. 

Walter, H. 1909. Phytolaccaceae. In A. Engler, Das Pflanzenreich 4(83): 1-154. 

Picramnia guianensis (Aubl.) Jans.-Jac. [expected in BO] 

Picramnia juniniana J.F. Macbr. 

Picramnia latifolia Tul. 

Picramnia magnifolia J.F. Macbr. 

Picramnia nuriensis Steyerm. 

Picramnia pentandra Sw. 

Picramnia sellowii Planch. 

Picramnia spruceana Engl. 

Picrasma excelsa (Sw.) Planch. 

PICRAMNIACEAE 

J.R. Pirani 


Picramnia longissima Tul. =Picramnia latifolia 

Picramnia macrostachys Klotzsch ex Engl. =Picramnia latifolia 

Picramnia platystachya Killip & Cuatrec. =Picramnia magnifolia 

Picramnia sellowii subsp. spruceana (Engl.) Pirani =Picramnia spruceana 

Picramnia tariri DC. =Picramnia guianensis 

Picramnia tristamina Steyerm. =Picramnia juniniana 

Tariri guianensis Aubl. =Picramnia guianensis 


AM GU SU 



BO 

BO GU FG 


BO DA 

BO FG 

GU SU FG 

BO DA GU FG 

DA GU FG? 

GU SU FG 

AM GU 


AM GU 

BO GU 

BO 

GU SU FG 


FG?

458 


PIPERACEAE 

R. Callejas-Posada, A.R.A. Görts-van Rijn, J.A. Steyermark† 

Peperomia acuminata Ruiz & Pav. [Mt. Roraima] 

Peperomia alata Ruiz & Pav. 

Peperomia alpina (Sw.) A. Dietr. [Mt. Roraima] 

Peperomia angularis C. DC. 

Peperomia angustata Kunth 

Peperomia blanda (Jacq.) Kunth 

Peperomia celiae Yunck. 

Peperomia circinnata Link 

Peperomia cladara Yunck. 

Peperomia delascioi Steyerm. [Mt. Roraima] 

Peperomia dendrophila Schltdl. & Cham. 

Peperomia distachya (L.) A. Dietr. [expected in GU] 

Peperomia duidana Trel. ex Gleason 

Peperomia elongata Kunth 

Peperomia elongata var. elongata 

Peperomia elongata var. guianensis Yunck. 

Peperomia elongata var. piliramea Trel. & Yunck. 

Peperomia emarginella (Sw. ex Wikstr.) C. DC. 

Peperomia enantiostachya C. DC. 

Peperomia fundacionensis Steyerm. 

Peperomia galioides Kunth [Mt. Roraima] 

Peperomia gentryi Steyerm. 

Peperomia glabella (Sw.) A. Dietr. 

Peperomia glabella var. glabella 

Peperomia glabella var. nervulosa (C. DC.) Yunck. 

Peperomia gracieana Görts 

Peperomia guaiquinimana Trel. & Yunck. 

Peperomia haematolepis Trel. 

Peperomia hernandiifolia (Vahl) A. Dietr. 

Peperomia jamesoniana C. DC. 

Peperomia lanceolato-peltata C. DC. 

Peperomia lancifolia Hook. 

Peperomia loxensis Kunth 

Peperomia macrostachya (Vahl) A. Dietr. 

Peperomia macrostachya var. macrostachya 

Peperomia macrostachya var. nematostachya (Link) Trel. & Yunck. 

Peperomia maculosa (L.) Hook. 

Peperomia magnoliifolia (Jacq.) A. Dietr. 

Peperomia maguirei Yunck. 

Peperomia manarae Steyerm. 

Peperomia marahuacensis Steyerm. 

Peperomia maypurensis Kunth 

Peperomia neblinana Yunck. 

Peperomia obtusifolia (L.) A. Dietr. 

Peperomia obtusifolia var. obtusifolia 

Peperomia ouabianae C. DC. 

Peperomia pellucida (L.) Kunth 

Peperomia pernambucensis Miq. 

Peperomia pilicaulis C. DC. 

Peperomia purpurinervis C. DC. [Mt. Roraima] 

Peperomia quadrangularis (J.V. Thomps.) A. Dietr. 

Peperomia quadrifolia (L.) Kunth 

Peperomia quaesita Trel. 

Peperomia rhombea Ruiz & Pav. 

Peperomia rotundata Kunth [Mt. Roraima] 

Peperomia rotundata var. rotundata 

Peperomia rotundata var. trinervula (C. DC.) Steyerm. [Mt. Roraima] 

Peperomia rotundifolia (L.) Kunth 

Peperomia schomburgkii C. DC. 

Peperomia serpens (Sw.) Loudon 

Peperomia spruceana Benth. ex C. DC. 

AM BO GU 


BO FG 

GU 


BO GU? SU 

AM 

SU? 

AM BO 

AM BO GU 

AM BO 

AM SU FG 

AM BO GU 


GU SU FG 


AM GU SU 


AM 

AM 

AM BO 

AM BO 




FG 

BO 

AM 

AM BO GU 

AM BO 

BO 

AM BO GU 

AM 



BO GU FG 

AM BO 


GU SU FG 

GU 

AM 

AM BO 

AM BO 


GU SU FG 



BO GU SU FG 

GU FG 

BO GU? FG 


GU 

AM BO 

GU 

AM BO 

AM 

AM BO 


GU 


AM

Peperomia striata Ruiz & Pav. 

Peperomia tenella (Sw.) A. Dietr. 

Peperomia tenella var. tenella 

Peperomia tenella var. tyleri (Trel.) Steyerm. 

Peperomia tetraphylla (G. Forst.) Hook. & Arn. 

Peperomia transparens Miq. 

Peperomia uaupesensis Yunck. 

Peperomia urocarpa Fisch. & C.A. Mey. 

Peperomia venezueliana C. DC. 

Peperomia venusta Yunck. 

Peperomia yatuensis Steyerm. 

Peperomia yutajensis Steyerm. 

Piper abellinum Trel. & Yunck. 

Piper adenandrum (Miq.) C. DC. 

Piper aduncum L. 

Piper aduncum var. aduncum 

Piper aduncum var. cordulatum (C. DC.) Yunck. 

Piper aduncum var. garcia-barrigae Trel. & Yunck. 

Piper aequale Vahl 

Piper alatabaccum Trel. & Yunck. 

Piper amalago L. 

Piper amapense Yunck. 

Piper amplectenticaule Trel. & Yunck. 

Piper amplum Kunth 

Piper angustifolium Lam. 

Piper anonifolium (Kunth) C. DC. 

Piper anonifolium var. anonifolium 

Piper anonifolium var. parkerianum (C. DC.) Steyerm. 

Piper arboreum Aubl. 

Piper arboreum var. arboreum 

Piper arboreum var. hirtellum Yunck. [subsp. arboreum] 

Piper augustum Rudge 

Piper augustum var. augustum 

Piper augustum var. pubinerve Trel. & Yunck. 

Piper auritum Kunth 

Piper avellanum (Miq.) C. DC. 

Piper avellanum var. angustifolium Trel. & Yunck. 

Piper avellanum var. avellanum 

Piper baccans (Miq.) C. DC. 

Piper baccans var. sancarlosianum (C. DC.) Steyerm. 

Piper bartlingianum (Miq.) C. DC. 

Piper bolivaranum Yunck. 

Piper brasiliense C. DC. 

Piper brownsbergense Yunck. 

Piper cernuum Vell. 

Piper cernuum var. cernuum 

Piper cernuum var. perlongispicum (Yunck.) Steyerm. 

Piper cililimbum Yunck. 

Piper ciliomarginatum Görts & Christenh. 

Piper consanguineum (Kunth) C. DC. 

Piper coruscans Kunth 

Piper crassinervium Kunth 

Piper cumanense Kunth 

Piper cuyunianum Steyerm. 

Piper cyrtopodon (Miq.) C. DC. 

Piper demeraranum (Miq.) C. DC. 

Piper dilatatum Rich. 

Piper divaricatum G. Mey. 

Piper duckei C. DC. 

Piper dumosum Rudge 

Piper dunstervilleorum Steyerm. 

Piper eucalyptifolium Rudge 

Piper fanshawei Yunck. 

Piper flexicaule (Miq.) C. DC. 


AM BO 

AM BO GU 

AM BO GU 

AM BO 

AM BO GU? 

SU FG 

AM BO 

BO GU 

AM 

AM 

AM 

AM 

BO 

AM GU SU 

FG 

FG 


AM 

AM 

BO 

BO 



BO GU SU FG 

GU? FG 

FG 

FG 

FG 



BO GU SU FG 





GU SU FG 

GU FG 

SU FG 

AM BO GU SU 

SU 

FG 

GU SU FG 

AM 

AM 

GU 


BO GU 

GU 

SU FG 

AM BO GU? FG 

AM BO 

BO 

GU? FG 

AM BO GU 

GU 

BO GU SU FG 


AM BO 

BO 

FG 

BO GU 

GU SU FG 



DA GU 

GU 

SU FG 

BO 

GU SU FG 

GU 

GU 

GU 

FG

460 


Piper foveolatum Kunth ex C. DC. 

Piper francovilleanum C. DC. 

Piper fuligineum Kunth 

Piper gabrielianum (Miq.) C. DC. 

Piper gentryi Steyerm. 

Piper glabrescens (Miq.) C. DC. 

Piper glabrescens var. caparonum (C. DC.) Yunck. 

Piper glabrescens var. glabrescens 

Piper guianense (Klotzsch) C. DC. 

Piper hermannii Trel. & Yunck. 

Piper hippocrepiforme Steyerm. 

Piper hirtilimbum Trel. & Yunck. 

Piper hispidum Sw. 

Piper holtii Trel. & Yunck. 

Piper hostmannianum (Miq.) C. DC. 

Piper humistratum Görts & K.U. Kramer 

Piper inaequale C. DC. 

Piper insigne (Kunth) C. DC. 

Piper jauaense Steyerm. 

Piper javitense Kunth 

Piper julianii Callejas 

Piper kappleri C. DC. 

Piper kegelianum (Miq.) C. DC. 

Piper kegelii C. DC. 

Piper laevigatum Kunth 

Piper lemaense Yunck. 

Piper liesneri Steyerm. 

Piper marginatum Jacq. 

Piper marginatum var. marginatum 

Piper microstachyum Vahl [=Piper angustifolium?] 

Piper modestum (Miq.) C. DC. [insufficiently known taxon] 

Piper morilloi Steyerm. 

Piper mosaicum Steyerm. 

Piper neblinanum Yunck. 

Piper nematanthera (Miq.) C. DC. 

Piper nematanthera var. nematanthera 

Piper nematanthera var. sagotii (C. DC.) Trel. & Yunck. 

Piper nigrispicum C. DC. 

Piper nigrum L. [cultivated, perhaps escaping] 

Piper obliquum Ruiz & Pav. 

Piper oblongifolium (Klotzsch) C. DC. 

Piper oblongifolium var. glabrum C. DC. 

Piper oblongifolium var. oblongifolium 

Piper otto-huberi Steyerm. 

Piper otto-huberi var. eciliatum Steyerm. 

Piper otto-huberi var. otto-huberi 

Piper ovatum Vahl [expected in GU] 

Piper papilliferum Steyerm. 

Piper paramaribense C. DC. 

Piper parapeltobryon Trel. 

Piper peltatum L. [=Pothomorphe peltata; =Lepianthes peltatum] 

Piper perciliatum Trel. & Yunck. 

Piper perstipulare Steyerm. 

Piper piscatorum Trel. & Yunck. 

Piper politii Yunck. 

Piper politii subsp. politii 

Piper politii subsp. sipapoense Steyerm. 

Piper politii subsp. toronoense Steyerm. 

Piper pseudoacreanum Steyerm. 

Piper pseudoglabrescens Trel. & Yunck. 

Piper pubivaginatum Steyerm. 

Piper pulleanum Yunck. 

Piper reticulatum L. 

Piper rudgeanum (Miq.) C. DC. 


AM BO 

GU SU FG 

FG 

BO 

GU SU 

GU 

GU SU 

AM GU FG? 

AM 

BO 

GU 


AM 


SU FG 

GU FG 

GU SU FG 

AM BO 

AM 

BO 

GU SU FG 

BO GU SU FG 

SU 

FG? 

AM BO 

AM GU 


AM BO DA 

FG 

FG 

AM 

AM 

AM BO 

GU SU FG 

GU SU FG 

FG 

GU SU FG 

DA 


GU SU FG 

GU 

GU SU FG 

AM 

AM 

AM 

BO 

AM 

GU SU FG 

AM 


AM 

AM BO GU 

AM BO GU FG 

AM BO 

AM BO 

AM 

BO 

AM 

AM BO DA GU 

AM 

GU SU FG 


SU FG?


Piper rupununianum Trel. & Yunck. 

Piper sabanaense Yunck. 

Piper salicifolium Vahl 

Piper schiedeanum Steud. 

Piper steyermarkii Yunck. 

Piper subalpinum Yunck. 

Piper subciliatum C. DC. 

Piper subduidaense Yunck. 

Piper tamayoanum Steyerm. 

Piper tectonifolium Kunth 

Piper tenue Kunth 

Piper tepuiense Steyerm. 

Piper toronotepuiense Steyerm. 

Piper tortivenulosum Yunck. 

Piper trichoneuron (Miq.) C. DC. 

Piper trigonum C. DC. 

Piper tuberculatum Jacq. 

Piper umbellatum L. [=Pothomorphe umbellata; =Lepianthes umbellatum] 

Piper venamoense Steyerm. 

Piper vitaceum Yunck. 

Piper vitaceum var. venezuelense Steyerm. 

Piper wachenheimii Trel. 


Acrocarpidium majus Miq. =Peperomia urocarpa 

Acrocarpidium tenellum (Sw.) Miq. =Peperomia tenella 

Acrocarpidium urocarpum (Fischer & C. Mey.) Miq. =Peperomia urocarpa 

Artanthe adenandra Miq. =Piper adenandrum 

Artanthe aduncum (L.) Miq. =Piper aduncum 

Artanthe amazonica Miq. =Piper coruscans 

Artanthe anonifolia (Kunth) Miq. =Piper anonifolium 

Artanthe apiculata Klotzsch =Piper anonifolium 

Artanthe asperifolia (Ruiz & Pav.) Miq. =Piper hispidum 

Artanthe augusta (Rudge) Miq. =Piper augustum 

Artanthe avellana Miq. =Piper avellanum 

Artanthe baccans Miq. =Piper baccans 

Artanthe bartlingianum Miq. =Piper bartlingianum 

Artanthe berbicense Miq. =Piper hostmannianum 

Artanthe caripensis (Kunth) Miq. =Piper aequale 

Artanthe caudata (Vahl) Miq. =Piper marginatum 

Artanthe consanguinea (Kunth) Miq. =Piper consanguineum 

Artanthe demerarana Miq. =Piper demeraranum 

Artanthe eximia (Kunth) Miq. =Piper cernuum 

Artanthe flagellaris (Kunth) Miq. =Piper obliquum 

Artanthe guianensis Klotzsch =Piper guianense 

Artanthe hirsuta (Sw.) Miq. =Piper hispidum 

Artanthe hostmanniana Miq. =Piper hostmannianum 

Artanthe javitensis (Kunth) Miq. =Piper javitense 

Artanthe kegeliana Miq. =Piper kegelianum 

Artanthe luschnathiana (Kunth) Miq. =Piper arboreum var. hirtellum 

Artanthe magnifica Miq. [nom. illeg.] =Piper obliquum 

Artanthe marginata (Jacq.) Miq. =Piper marginatum 

Artanthe obrumbata Miq. =Piper arboreum var. hirtellum 

Artanthe parkeriana Miq. =Piper anonifolium var. parkerianum 

Artanthe peduncularis Miq. =Piper pedunculare 

Artanthe pseudochurumayu (Kunth) Miq. =Piper coruscans 

Artanthe ramiflora Miq. ex C. DC. =Piper hostmannianum 

Artanthe schomburgkii Klotzsch =Piper pedunculare 

Artanthe spectabilis Miq. =Piper cernuum 

Artanthe superba Miq. =Piper obliquum 

Enckea dubia Kunth =Piper foveolatum 

Heckeria peltata (L.) Kunth =Piper peltatum 

Heckeria umbellata (L.) Kunth =Piper umbellatum 

Lepianthes peltatum (L.) Raf. =Piper peltatum 

AM GU 

BO 

AM 

AM 

BO 

BO 

BO 

BO DA 

BO 

BO 

SU 

GU? SU? 

FG 

GU? FG 

GU SU FG 

AM BO GU 


BO GU 

BO GU 

BO 

BO 

BO GU SU FG

462 


Lepianthes umbellatum (L.) Raf. =Piper umbellatum 

Ottonia ovata (Vahl) Trel. =Piper ovatum 

Peperomia alata var. angustifolia C. DC. =Peperomia alata 

Peperomia angulata Kunth =Peperomia quadrangularis 

Peperomia atabapoensis Steyerm. =Peperomia quadrangularis 

Peperomia bartlettii C. DC. =Peperomia rotundifolia 

Peperomia blanda var. langsdorffii (Miq.) Henschen =Peperomia blanda 

Peperomia boomii Steyerm. =Peperomia lanceolato-peltata 

Peperomia caulibarbis Miq. =Peperomia glabella var. glabella 

Peperomia chimantana Yunck. =Peperomia jamesoniana 

Peperomia choroniana var. puberulenta Yunck. =Peperomia hernandiifolia 

Peperomia ciliata Kunth =Peperomia blanda 

Peperomia cladara f. ciliata Yunck. =Peperomia cladara 

Peperomia cladara f. perglabra Yunck. =Peperomia cladara 

Peperomia cryptotricha Trel. ex V.M. Badillo [nomen nudum] =Peperomia tetraphylla 

Peperomia cuneata Miq. =Peperomia obtusifolia 

Peperomia decipiens C. DC. =Peperomia blanda 

Peperomia diffusa C. DC. =Peperomia loxensis 

Peperomia distachya f. geminispica (Trel. & Yunck.) Steyerm. =Peperomia distachya 

Peperomia emarginata Ruiz & Pav. =Peperomia obtusifolia 

Peperomia emarginella var. glabrior C. DC. =Peperomia emarginella 

Peperomia epilobioides Trel. & Yunck. =Peperomia rotundata var. rotundata 

Peperomia erasmiaeformis Trel. =Peperomia lancifolia 

Peperomia ernstiana C. DC. =Peperomia venezueliana 

Peperomia foveolata Steyerm. =Peperomia distachya 

Peperomia geminispica Trel. & Yunck. =Peperomia distachya 

Peperomia glabella var. melanostigma (Miq.) Dahlst. =Peperomia glabella var. nervulosa 

Peperomia guaiquinimana Trel. ex V.M. Badillo [nomen nudum] =Peperomia guaiquinimana 

Peperomia hederacea Miq. =Peperomia urocarpa 

Peperomia horrescens Trel. ex V.M. Badillo [nomen nudum] =Peperomia venezueliana 

Peperomia ionophylla Griseb. =Peperomia serpens 

Peperomia lancifolia subsp. erasmiaeformis (Trel.) Steyerm. =Peperomia lancifolia 

Peperomia lanjouwii Yunck. =Peperomia rotundifolia 

Peperomia longemucronata C. DC. =Peperomia alpina 

Peperomia longemucronata f. longirostrata (Yunck.) Steyerm. =Peperomia alpina 

Peperomia longifolia C. DC. =Peperomia pernambucensis 

Peperomia longirostrata Yunck. =Peperomia alpina 

Peperomia macaroana Trel. ex V.M. Badillo [nomen nudum] =Peperomia blanda 

Peperomia magnoliifolia var. emarginulata C. DC. =Peperomia obtusifolia 

Peperomia major C. DC. =Peperomia urocarpa 

Peperomia melanostigma var. nervulosa C. DC. =Peperomia glabella var. nervulosa 

Peperomia microreticulata Steyerm. =Peperomia alata 

Peperomia moritzii C. DC. =Peperomia rotundata var. trinervula 

Peperomia myosuroides (Rudge) A. Dietr. =Peperomia macrostachya 

Peperomia myriocarpa Miq. =Peperomia elongata var. elongata 

Peperomia nematostachya Link =Peperomia macrostachya var. nematostachya 

Peperomia nilssonii Yunck. =Peperomia alata 

Peperomia nummularifolia Kunth =Peperomia rotundifolia 

Peperomia obtusifolia var. cuneata (Miq.) Griseb. =Peperomia obtusifolia 

Peperomia obtusifolia var. emarginata (Ruiz & Pav.) Dahlst. =Peperomia obtusifolia 

Peperomia obtusifolia var. emarginulata (C. DC.) Trel. & Yunck. =Peperomia obtusifolia 

Peperomia omnicola C. DC. =Peperomia striata 

Peperomia omnicola var. oblanceolata Trel. =Peperomia striata 

Peperomia ornata Yunck. =Peperomia maypurensis 

Peperomia paniculata Regel =Peperomia pernambucensis 

Peperomia parkeriana Miq. =Peperomia macrostachya var. macrostachya 

Peperomia peltata (L.) A. Dietr. =Piper peltatum 

Peperomia pennellii Trel. & Yunck. =Peperomia dendrophila 

Peperomia pseudojamesoniana Steyerm. =Peperomia jamesoniana 

Peperomia purpurascens Moritz ex C. DC. =Peperomia rotundata var. trinervula 

Peperomia pyrifolia Kunth =Peperomia acuminata 

Peperomia reflexa (L. f.) A. Dietr. [nom, illeg., non Kunth] =Peperomia tetraphylla 

Peperomia reflexa f. americana Miq. =Peperomia tetraphylla 

Peperomia reflexa var. americana (Miq.) Dahlst. =Peperomia tetraphylla


Peperomia reptans C. DC. =Peperomia serpens 

Peperomia roraimana C. DC. =Peperomia rotundata var. trinervula 

Peperomia rotundifolia f. ovata Dahlst. =Peperomia rotundifolia 

Peperomia rotundifolia var. ovata (Dahlst.) C. DC. =Peperomia rotundifolia 

Peperomia saginans Trel. ex V.M. Badillo [nomen nudum] =Peperomia quadrangularis 

Peperomia salvaje C. DC. =Peperomia blanda 

Peperomia silvestris C. DC. =Peperomia ouabianae 

Peperomia surinamensis C. DC. =Peperomia elongata var. elongata 

Peperomia tafelbergensis Yunck. =Peperomia ouabianae 

Peperomia tetraphylla var. americana (Miq.) Yunck. =Peperomia tetraphylla 

Peperomia trifolia (L.) A. Dietr. [not in flora area] 

Peperomia trinervula C. DC. =Peperomia rotundata var. trinervula 

Peperomia trinervula var. suboppositifolia Trel. & Yunck. =Peperomia rotundata var. rotundata 

Peperomia tyleri Trel. =Peperomia tenella var. tyleri 

Peperomia umbellata (L.) Kunth =Piper umbellatum 

Peperomia victoriana C. DC. =Peperomia angustata 

Peperomia victoriana var. marginata C. DC. =Peperomia angustata 

Peperomia viridispica Trel. =Peperomia angustata 

Peperomia wurdackii Yunck. =Peperomia ouabianae 

Piper acarouanyanum C. DC. =Piper avellanum 

Piper acutissimum Trel. =Piper trigonum 

Piper adenophorum C. DC. =Piper dumosum 

Piper aduncum var. brachyarthrum (Trel.) Yunck. =Piper aduncum var. cordulatum 

Piper aequale var. santa-martanum (C. DC.) Trel. & Yunck. =Piper aequale 

Piper affine C. DC. =Piper paramaribense 

Piper aggregatum P.J. Bergius (Lacistemataceae) =Lacistema aggregatum 

Piper agostinii Yunck. =Piper kegelianum 

Piper alpinum Sw. =Peperomia alpina 

Piper amalago f. ceanothifolium (Kunth) Steyerm. [var. medium] =Piper amalago 

Piper amalago var. medium (Jacq.) Yunck. =Piper amalago 

Piper amazonasense Trel. & Yunck. =Piper baccans var. sancarlosianum 

Piper amazonicum (Miq.) C. DC. =Piper coruscans 

Piper amplectens (Miq.) C. DC. =Piper fuligineum 

Piper angremondii DC. =Piper hispidum 

Piper angustatum (Kunth) Poir. =Peperomia angustata 

Piper angustifolium var. cordulatum C. DC. =Piper aduncum var. cordulatum 

Piper angustum Rudge ex Spreng. [nom. superfl.] =Piper augustum 

Piper anisatum Kunth =Piper marginatum 

Piper anonifolium Kunth [nom. invalid] =Piper anonifolium 

Piper arboreum var. falcifolium (Trel.) Yunck. =Piper arboreum 

Piper arboreum var. latifolium (C. DC.) Yunck. =Piper arboreum var. hirtellum 

Piper arboreum subsp. tuberculatum (Jacq.) Tebbs =Piper tuberculatum 

Piper arieianum C. DC. =Piper trigonum 

Piper arieianum var. chimantanum (Yunck.) Steyerm. =Piper trigonum 

Piper arieianum var. duidaense (Trel.) Steyerm. =Piper trigonum 

Piper aristeguietae Yunck. =Piper obliquum 

Piper asperifolium Rich. =Piper hispidum 

Piper asperifolium Ruiz & Pav. =Piper hispidum 

Piper bartlingianum var. parimanum (Trel.) Steyerm. =Piper bartlingianum 

Piper benoistii Trel. ex Lemée [apparently a nomen nudum] 

Piper blandum Jacq. =Peperomia blanda 

Piper brachystachyum C. DC. =Piper eucalyptifolium 

Piper cabellense C. DC. =Piper cernuum 

Piper caracasanum Bredem. =Piper obliquum 

Piper caripense Kunth =Piper aequale 

Piper carniconnectivum C. DC. [not in flora area] 

Piper caudatum Vahl =Piper marginatum 

Piper cauranum Trel. =Piper demeraranum 

Piper ceanothifolium Kunth =Piper amalago 

Piper celtidifolium Kunth =Piper aduncum 

Piper chimantanum Yunck. =Piper trigonum 

Piper citrifolium Lam. =Piper anonifolium var. anonifolium 

Piper citrifolium auct. non Lam.: sensu Trel. & Yunck. =Piper anonifolium 

Piper citrifolium var. parkerianum (Miq.) Trel. & Yunck. =Piper anonifolium var. parkerianum

464 


Piper colubrinum Link =Piper divaricatum 

Piper consanguineum Kunth [pub. as synonym] =Piper consanguineum 

Piper coruscans var. membranaceum (C. DC.) Steyerm. =Piper coruscans 

Piper corylifolium Kunth =Piper dilatatum 

Piper crassum Vell. =Piper divaricatum 

Piper demeraranum var. rubescens (C. DC.) Trel. & Yunck. =Piper demeraranum 

Piper deminutum Yunck. =Piper bolivaranum 

Piper diospyrifolium Kunth [not in flora area] 

Piper distachyon L. =Peperomia distachya 

Piper duidaense Trel. =Piper trigonum 

Piper duidaense var. eciliatum Trel. & Yunck. =Piper trigonum 

Piper elongatum var. brachyarthrum Trel. =Piper aduncum var. cordulatum 

Piper elongatum var. cordulatum (C. DC.) C. DC. =Piper aduncum var. cordulatum 

Piper emarginatum (Ruiz & Pav.) Vahl =Peperomia obtusifolia 

Piper emarginellum Sw. ex Wikstr. =Peperomia emarginella 

Piper esmeraldanum Trel. =Piper demeraranum 

Piper eximium Kunth =Piper cernuum 

Piper falcifolium Trel. =Piper arboreum 

Piper flagellare Kunth =Piper obliquum 

Piper flexuosum Rudge =Piper flexicaule 

Piper fulgidum Yunck. [not in flora area] 

Piper funckii C. DC. =Piper aequale 

Piper geayi Trel. ex Lemée [apparently a nomen nudum] 

Piper geniculatum Sw. =Piper arboreum 

Piper geniculatum var. latifolium C. DC. =Piper arboreum var. hirtellum 

Piper geniculatum f. puberula C. DC. =Piper tuberculatum 

Piper geniculatum var. ß C. DC. =Piper arboreum var. hirtellum 

Piper gigantifolium C. DC. =Piper cernuum 

Piper glabellum (Miq.) C. DC. [nom. illeg., non Piper glabellum Sw.] =Piper divaricatum 

Piper glabellum Sw. =Peperomia glabella 

Piper glabrilimbum C. DC. =Piper divaricatum 

Piper gleasonii Yunck. =Piper trichoneuron 

Piper godebertianum Trel. ex Lemée [apparently a nomen nudum] 

Piper hernandiifolium Vahl =Peperomia hernandiifolia 

Piper hirsutum Sw. =Piper hispidum 

Piper hispidum Kunth [nom. illeg., non Piper hispidum Sw.] =Piper hispidum 

Piper hispidum var. obliquum Trel. & Yunck. =Piper hispidum 

Piper hohenackeri C. DC. =Piper anonifolium 

Piper hostmannianum var. berbicense (Miq.) C. DC. =Piper hostmannianum 

Piper hostmannianum var. glabrirameum Trel. & Yunck. =Piper hostmannianum 

Piper hostmannianum var. ramiflorum C. DC. =Piper hostmannianum 

Piper insipiens Trel. & Yunck. =Piper divaricatum 

Piper johnstonii C. DC. =Piper aequale 

Piper lapathifolium Steud. [not in flora area] 

Piper latum Kunth =Piper reticulatum 

Piper lenormandianum C. DC. =Piper kappleri 

Piper leprieuri Pulle =Piper consanguineum 

Piper loxense (Kunth) Poir. =Peperomia loxensis 

Piper lucaeanum var. magnifolium C. DC. =Piper trigonum 

Piper macrophyllum Sw. =Piper arboreum 

Piper macrostachyon Vahl =Peperomia macrostachya 

Piper maculosum L. =Peperomia maculosa 

Piper magnoliifolium Jacq. =Peperomia magnoliifolia 

Piper maguirei Yunck. =Piper paramaribense 

Piper marginatum var. anisatum (Kunth) C. DC. =Piper marginatum 

Piper marowinianum C. DC. =Piper kappleri 

Piper marowynense C. DC. =Piper avellanum 

Piper maypurense (Kunth) Poir. =Peperomia maypurensis 

Piper medium Jacq. =Piper amalago 

Piper miquelii C. DC. =Piper eucalyptifolium 

Piper mollipilosum C. DC. [not in flora area] 

Piper moraballianum Trel. =Piper avellanum 

Piper myosuroides Rudge =Peperomia macrostachya 

Piper nitidum Vahl [nom. illeg., non Piper nitidum Sw] =Piper divaricatum


Piper nitidum Sw. =Piper arboreum 

Piper nummularifolium Sw. =Peperomia rotundifolia 

Piper obliquum var. eximium (Kunth) C. DC. =Piper cernuum 

Piper obrumbatum (Miq.) C. DC. =Piper arboreum var. hirtellum 

Piper obtusifolium L. =Peperomia obtusifolia 

Piper oparumaense Yunck. =Piper obliquum 

Piper orenocanum C. DC. =Piper coruscans 

Piper orenocanum var. koriaboense Trel. & Yunck. =Piper coruscans 

Piper parakegelianum Sagot ex Lemée [apparently a nomen nudum] 

Piper parimanum Trel. =Piper bartlingianum 

Piper parkerianum (Miq.) C. DC. =Piper anonifolium var. parkerianum 

Piper pavasense Steyerm. =Piper julianii 

Piper pedunculare (Miq.) C. DC. [comb. illeg.] 

Piper pellucidum L. =Peperomia pellucida 

Piper perlongispicum Yunck. =Piper cernuum var. perlongispicum 

Piper perlongivillosum Yunck. =Piper francovilleanum 

Piper piresii Yunck. [not in flora area] 

Piper poiteanum Kunth =Piper foveolatum 

Piper pruinosum Kunth =Piper peltatum 

Piper pseudochurumayu var. membranaceum C. DC. =Piper coruscans 

Piper pseudomarowynense Trel. ex Lemée [apparently a nomen nudum] 

Piper purpurascens Desf. ? =Piper angustifolium 

Piper quadrangulare J.V. Thomps. =Peperomia quadrangularis 

Piper reflexum L. f. =Peperomia tetraphylla 

Piper rhododendrifolium Kunth =Piper anonifolium 

Piper rhynchostachyum C. DC. =Piper eucalyptifolium 

Piper rio-paraguanum Trel. ex V.M. Badillo [nomen nudum] =Piper hostmannianum 

Piper romboutsii Yunck. =Piper divaricatum 

Piper rotundifolium L. =Peperomia rotundifolia 

Piper rubescens C. DC. =Piper demeraranum 

Piper rugosum Vahl =Piper anonifolium 

Piper ruizianum D. Dietr. =Piper obliquum 

Piper sagoti C. DC. =Piper nematanthera var. sagotii 

Piper saltuum C. DC. =Piper trigonum 

Piper sancarlosianum C. DC. =Piper baccans var. sancarlosianum 

Piper santa-martanum C. DC. =Piper aequale 

Piper saramaccanum Yunck. =Piper obliquum 

Piper scabrum Sw. =Piper hispidum 

Piper serpens Sw. =Peperomia serpens 

Piper smilacifolium Kunth =Piper reticulatum 

Piper speciosum Kunth =Piper peltatum 

Piper statarium Trel. & Yunck. =Piper avellanum 

Piper subcrassifolium Yunck. =Piper hostmannianum 

Piper submelanostictum var. amelanostictum Yunck. =Piper obliquum 

Piper submelanostictum var. submelanostictum =Piper obliquum 

Piper surinamense C. DC. =Piper consanguineum 

Piper tenellum Sw. =Peperomia tenella 

Piper tetraphyllum G. Forst. =Peperomia tetraphylla 

Piper tiliifolium Ham. (Moraceae) =Bagassa guianensis 

Piper tonoroanum Trel. & Yunck. =Piper avellanum 

Piper tuberculatum var. minus C. DC. =Piper tuberculatum 

Piper tuberculatum var. scandens Trel. & Yunck. =Piper tuberculatum 

Piper variegatum Kunth =Piper cumanense 

Piper verrucosum Sw. =Piper arboreum 

Piper warakabacoura (Miq.) C. DC. =Piper bartlingianum 

Piper wurdackii Yunck. =Piper coruscans 

Piper zediogodiense Trel. ex L. Williams =Piper aequale 

Pothomorphe peltata (L.) Miq. =Piper peltatum 

Pothomorphe umbellata (L.) Miq. =Piper umbellatum 

Quebitea guianensis Aubl. =Piper humistratum 

Schilleria aequalis (Vahl) Kunth =Piper aequale 

Schilleria caudata (Vahl) Kunth =Piper marginatum 

Schilleria marginata (Jacq.) Kunth =Piper marginatum 

Steffensia adunca (L.) Kunth =Piper aduncum

466 


Steffensia anonifolia Kunth =Piper anonifolium 

Steffensia consanguinea Kunth =Piper consanguineum 

Steffensia eximia Kunth =Piper cernuum 

Steffensia hirsuta (Sw.) Kunth =Piper hispidum 

Steffensia insignis Kunth =Piper insigne 

Steffensia luschnathiana Kunth =Piper arboreum var. hirtellum 

Steffensia obliqua (Ruiz & Pav.) Kunth =Piper obliquum 

Steffensia pseudochurumayu Kunth =Piper coruscans 

REFERENCES 

Kramer, K.U. & A.R.A. Görts-van Rijn. 1968. Piperaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 

1(2): 415-421. 

Görts-van Rijn, A. 2002. Piperaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 574-584. 

Steyermark, J.A. 1984. Piperaceae. Flora de Venezuela II(2): 1-619. 

Steyermark, J.A. & R. Callejas-Posada. 2003. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 7: 681-738. 

Tebbs, M.C. 1989. Revision of Piper in the New World. 1. Review of characters and taxonomy of Piper section Macrostachys. 

Bull. Brit. Mus. Nat. Hist. (Bot.) 19: 117-158. 

Tebbs, M.C. 1990. Revision of Piper in the New World. 2. The taxonomy of Piper section Churumayu. Bull. Brit. Mus. Nat. Hist. 

(Bot.) 20(2): 193-236. 

Tebbs, M.C. 1993. Revision of Piper in the New World. 3. The taxonomy of Piper sections Lepianthes and Radula. Bull. Nat. 

Hist. Mus. 23(1): 1-50. 

Trelease, W. & T.G. Yuncker. 1950. The Piperaceae of Northern South America. Urbana, Illinois: University of Illinois Press. 

Yuncker, T.G. 1957. Piperaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 218-290. 

Yuncker, T.G. 1972. The Piperaceae of Brazil. Hoehnea 2: 19-366. 

Plantago lanceolata L. [introduced] 

Plantago major L. [introduced] 

PLANTAGINACEAE 

G. Rodríguez 

REFERENCES 

D’Arcy, W.G. 1971. Plantaginaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. 

Gard. 58(3): 363-369. 

Lanjouw, J. 1938. Plantaginaceae. In A. Pulle, Flora of Suriname 4(2): 253-254. 

Rodríguez, G. 2003. Plantaginaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 7: 739. 

PLUMBAGINACEAE 

C.A. Vargas 

Plumbago auriculata Lam. [widely cultivated, perhaps escaping] 

Plumbago zeylanica L. [naturalized] 


Plumbago scandens L. =Plumbago zeylanica 

REFERENCES 

Vargas, C.A. 2003. Plumbaginaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 7: 740-741. 

Apinagia arminensis P. Royen 

Apinagia brevicaulis Mildbr. 

Apinagia corymbosa (Tul.) Engl. 

Apinagia corymbosa var. corymbosa 

Apinagia crispa P. Royen 

Apinagia digitata P. Royen 

Apinagia divertens Went 

PODOSTEMACEAE 

P.E. Berry 

FG 


BO DA GU? SU FG 

SU FG 

BO 

BO GU SU FG 

BO GU SU 

SU 

FG 

FG 

SU FG

Apinagia exilis (Tul.) P. Royen 

Apinagia flexuosa (Tul.) P. Royen 

Apinagia goejei Went 

Apinagia guyanensis (Pulle) P. Royen 

Apinagia hulkiana (Went) P. Royen 

Apinagia imthurnii (Goebel) P. Royen 

Apinagia itanensis Schnell 

Apinagia kochii (Engl.) P. Royen 

Apinagia latifolia (Goebel) P. Royen 

Apinagia leptophylla (Goebel) P. Royen 

Apinagia longifolia (Tul.) P. Royen 

Apinagia marowynensis (Went) P. Royen 

Apinagia multibranchiata (Matthiesen) P. Royen 

Apinagia nana Went [=Apinagia pilgeri?] 

Apinagia penicillata (P. Royen) P. Royen 

Apinagia petiolata Hollander 

Apinagia pilgeri Mildbr. 

Apinagia pusilla Tul. 

Apinagia richardiana (Tul.) P. Royen 

Apinagia ruppioides (Kunth) Tul. 

Apinagia secundiflora (Tul.) Pulle 

Apinagia staheliana (Went) P. Royen 

Apinagia surumuensis (Engl.) P. Royen 

Apinagia treslingiana (Went) P. Royen 

Apinagia versteegiana (Went) P. Royen 

Jenmaniella ceratophylla Engl. 

Jenmaniella ceratophylla var. ceratophylla 

Jenmaniella ceratophylla var. hexandra Engl. 

Jenmaniella ceratophylla var. parva P. Royen 

Jenmaniella guianensis Engl. 

Jenmaniella isoetifolia P. Royen 

Jenmaniella jenmanii (Engl.) P. Royen 

Jenmaniella tridactylitifolia Engl. 

Jenmaniella varians Engl. 

Marathrum aeruginosum P. Royen 

Marathrum capillaceum (Pulle) P. Royen 

Marathrum pauciflorum Tul. 

Marathrum pauciflorum var. heterophyllum P. Royen 

Marathrum pauciflorum var. pauciflorum 

Marathrum squamosum Wedd. 

Marathrum squamosum var. squamosum 

Marathrum utile Tul. 

Mourera fluviatilis Aubl. 

Oserya flabellifera Tul. & Wedd. [not in flora area?] 

Oserya minima P. Royen 

Oserya perpusilla (Went) P. Royen 

Oserya sphaerocarpa Tul. & Wedd. 

Podostemum dichotomum Klotzsch [dubious species] 

Rhyncholacis apiculata P. Royen 

Rhyncholacis applanata K.I. Goebel 

Rhyncholacis applanata var. applanata 

Rhyncholacis applanata var. laxipinnata P. Royen 

Rhyncholacis brassicifolia P. Royen 

Rhyncholacis brevistamina P. Royen 

Rhyncholacis coronata P. Royen 

Rhyncholacis cristata P. Royen 

Rhyncholacis dentata P. Royen 

Rhyncholacis divaricata Matthiesen 

Rhyncholacis flagellifolia P. Royen 

Rhyncholacis guyanensis P. Royen 

Rhyncholacis hydrocichorium Tul. 

Rhyncholacis jenmanii Engl. 

Rhyncholacis linearis Tul. 

Rhyncholacis macrocarpa Tul. [dubious species] 


BO GU SU 

GU SU FG 

SU 

BO SU FG 

GU SU 

GU SU FG? 

FG 

BO FG 

GU 

GU 


SU FG? 

AM BO 

SU 

SU 

SU 

SU 

GU 


AM BO 

GU SU FG 

AM BO GU SU 

GU 

SU 

SU 

BO GU 

GU 

GU 

BO 

GU 

GU 

GU 

GU 

GU 

AM 

BO SU FG 

GU 

GU 

GU 

AM FG 

AM 

AM 


SU 

SU 

BO GU SU FG 

GU 

GU 

GU 

AM BO GU 

AM BO GU 

GU 

GU? 

GU 

BO GU 

SU 

SU 

AM BO 

BO 

GU 

BO GU 

GU 

GU 

GU SU?

468 

Rhyncholacis oligandra Wedd. 

Rhyncholacis oligandra var. oligandra 

Rhyncholacis oligandra var. tenella P. Royen 

Rhyncholacis palmettifolia P. Royen 

Rhyncholacis palmettifolia P. Royen var. palmettifolia 

Rhyncholacis palmettifolia var. rosea P. Royen 

Rhyncholacis penicillata Matthiesen 

Tristicha trifaria (Bory ex Willd.) Spreng. 

Weddellina squamulosa Tul. 



Apinagia flabellifera (Tul. & Wedd.) Went =Oserya flabellifera 

Apinagia perpusilla Went =Oserya perpusilla 

Dufourea trifaria Bory ex Willd. =Tristicha trifaria 

Ligea flexuosa Tul. =Apinagia flexuosa 

Ligea longifolia (Tul.) Tul. =Apinagia longifolia 

Ligea richardiana Tul. =Apinagia richardiana 

Ligea richardiana var. corymbosa Tul. =Apinagia corymbosa 

Ligea richardiana var. exilis Tul. =Apinagia exilis 

Ligea secundiflora Tul. =Apinagia secundiflora 

Lophogyne capillacea Pulle =Marathrum capillaceum 

Marathrum squamosum var. spruceanum Wedd. =Marathrum squamosum var. squamosum 

Neolacis corymbosa (Tul.) Wedd. =Apinagia corymbosa 

Neolacis corymbosa var. exilis Tul. =Apinagia exilis 

Neolacis secundiflora (Tul.) Wedd. =Apinagia secundiflora 

Oenone flexuosa (Tul.) Wedd. =Apinagia flexuosa 

Oenone guyanensis Pulle =Apinagia guyanensis 

Oenone hulkiana Went =Apinagia hulkiana 

Oenone imthurnii Goebel =Apinagia imthurnii 

Oenone kochii Engl. =Apinagia kochii 

Oenone longifolia Tul. =Apinagia longifolia 

Oenone marowynensis Went =Apinagia marowynensis 

Oenone multibranchiata Matthiesen =Apinagia multibranchiata 

Oenone penicillata P. Royen =Apinagia penicillata 

Oenone richardiana (Tul.) Warm. =Apinagia richardiana 

Oenone secundiflora (Tul.) Engl. =Apinagia secundiflora 

Oenone staheliana Went =Apinagia staheliana 

Oenone treslingiana Went =Apinagia treslingiana 

Oenone versteegiana Went =Apinagia versteegiana 

Podostemum ruppioides Kunth =Apinagia ruppioides 

Tristicha hypnoides (A. St.-Hil.) Spreng. =Tristicha trifaria 

REFERENCES 

Berry, P.E. 2004. Podostemaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

8: 301-316. 

Den Hollander, G. & C.C. Berg. 1983. Studies on the flora of the Guianas 4. A new species of Apinagia and a key to the Apinagia 

species in Suriname. Acta Bot. Neerl. 32(3): 223-226. 

Philbrick, C.T. 2002. Podostemaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 585. 

Royen, P. van. 1951-1954. The Podostemaceae of the New World. Med. Bot. Mus. Herb. Rijks Univ. Utrecht 107: 1-151 (1951); 

115: 1-21 (1953); 119: 215-263. 

Schnell, R. 1969. Contribution à l’étude des Podostémacées de Guyane. Adansonia ser. 2 9(2): 249-271. 

Barnhartia floribunda Gleason 

Bredemeyera altissima (Poepp.) A.W. Benn. 

Bredemeyera bracteata Klotzsch ex Hassk. 

Bredemeyera densiflora A.W. Benn. 

Bredemeyera densiflora var. densiflora 

Bredemeyera densiflora var. glabra A.W. Benn. 

Bredemeyera floribunda Willd. 

POLYGALACEAE 

G.A. Aymard C., P.E. Berry, B. Eriksen 

AM GU 

AM GU 

GU 

GU 

GU 

GU 

AM BO 

BO DA GU FG 



AM BO GU 

GU 

AM BO GU SU 

GU SU? 

AM BO GU SU 

AM BO GU

Bredemeyera lucida (Benth.) Klotzsch ex Hassk. 

Bredemeyera myrtifolia A.W. Benn. 

Bredemeyera sp. A 

Diclidanthera bolivarensis Pittier 

Diclidanthera octandra Gleason 

Diclidanthera sp. A 

Diclidanthera wurdackiana Aymard & P.E. Berry 

Monnina cacumina N.E. Br. 

Moutabea aculeata (Ruiz & Pav.) Poepp. & Endl. 

Moutabea aff. chodatiana Huber 

Moutabea guianensis Aubl. 

Moutabea sp. A 

Moutabea sp. B 

Polygala adenophora DC. 

Polygala appressa Benth. 

Polygala asperuloides Kunth 

Polygala blakeana Steyerm. 

Polygala brevialata Chodat 

Polygala caracasana Kunth 

Polygala coelosioides Mart. 

Polygala echinosperma Görts 

Polygala galioides Poir. 

Polygala glochidiata Kunth 

Polygala glochidiata var. glochidiata 

Polygala hebeclada DC. 

Polygala hygrophila Kunth [expected in FG] 

Polygala longicaulis Kunth 

Polygala martiana A.W. Benn. 

Polygala membranacea (Miq.) Görts 

Polygala microspora S.F. Blake 

Polygala paniculata L. 

Polygala planellasi Molinet & M. Gómez 

Polygala pseudocoelosioides Chodat 

Polygala sanariapoana Steyerm. 

Polygala savannarum Chodat 

Polygala sipapoana Wurdack 

Polygala spectabilis DC. 

Polygala spruceana A.W. Benn. 

Polygala subtilis Kunth 

Polygala tenella Willd. 

Polygala timoutoides Chodat 

Polygala timoutoides var. maguirei (Wurdack) Marques 

Polygala timoutou Aubl. 

Polygala trichosperma L. 

Polygala violacea Aubl., emend. Marques 

Securidaca bialata Benth. 

Securidaca cacumina Wurdack 

Securidaca cayennensis S.F. Blake 

Securidaca coriacea Bonpl. 

Securidaca divaricata Nees & Mart. 

Securidaca diversifolia (L.) S.F. Blake 

Securidaca fruticans Wurdack 

Securidaca lateralis A.W. Benn. 

Securidaca longifolia Poepp. 

Securidaca macrophylla (Benth.) Wurdack 

Securidaca maguirei Wurdack 

Securidaca marginata Benth. 

Securidaca paniculata Rich. 

Securidaca paniculata var. lasiocarpa Oort 

Securidaca paniculata var. paniculata 

Securidaca pendula Bonpl. 

Securidaca prancei Wurdack 

Securidaca pubescens DC. 

Securidaca pubiflora Benth. 



AM 

BO 

AM BO 

AM 

AM 

AM BO 

AM BO GU 

AM BO GU FG 

AM BO 


AM BO DA 

AM 



BO 


AM BO 

BO 

GU? 

SU FG 

BO GU SU FG 

BO GU 

BO GU 

GU? 

AM BO GU SU 


FG 

SU FG 

AM BO GU 

BO GU 

FG? 

GU 

AM 

AM BO GU 

AM 


AM BO GU 

AM BO GU SU 


AM BO GU 

AM BO GU 




AM BO DA 

AM BO 

FG 


AM GU SU FG 


AM 

GU 

AM GU 

FG 

AM 

BO GU 




AM BO GU 

AM BO 


GU

470 

Securidaca pyramidalis Sprague ex Sandwith 

Securidaca retusa Benth. 

Securidaca savannarum Wurdack 

Securidaca scandens Jacq. 

Securidaca speciosa Wurdack 

Securidaca spinifex Sandwith 

Securidaca tenuifolia Chodat 

Securidaca tomentosa A. St.-Hil. & Moq. 

Securidaca uniflora Oort 

Securidaca volubilis L., emend. Oort 

Securidaca warmingiana Chodat 



Acosta aculeata Ruiz & Pav. =Moutabea aculeata 

Bredemeyera lucida (Benth.) A.W. Benn. =Bredemeyera lucida 

Bredemeyera parviflora A.W. Benn. =Bredemeyera myrtifolia 

Catocoma altissima Poepp. & Endl. =Bredemeyera altissima 

Catocoma floribunda Benth. =Bredemeyera floribunda 

Catocoma lucida Benth. =Bredemeyera lucida 

Diclidanthera orinocensis Pittier =Diclidanthera bolivarensis 

Monnina cacumina var. duidae (S.F. Blake) Steyerm. =Monnina cacumina 

Monnina duidae S.F. Blake =Monnina cacumina 

Monnina membranacea Miq. =Polygala membranacea 

Monnina uaipanensis Wurdack =Monnina cacumina 

Moutabea longifolia Poepp. & Endl. =Moutabea aculeata 

Polygala adenophora var. gracilis Chodat =Polygala adenophora 

Polygala adenophora var. robusta Chodat =Polygala adenophora 

Polygala alopecurus Chodat =Polygala tenella 

Polygala angustifolia Kunth =Polygala violacea 

Polygala angustifolia var. latifolia A. St.-Hil. ex Chodat =Polygala violacea 

Polygala appressa subsp. cumbrensis Wurdack =Polygala appressa 

Polygala appressa var. gracillima (S. Watson) A.W. Benn. =Polygala appressa 

Polygala appressa var. kavanayena Steyerm. =Polygala appressa 

Polygala aristeguietae Wurdack =Polygala brevialata 

Polygala brizoides A. St.-Hil. =Polygala violacea 

Polygala camporum Benth. =Polygala violacea 

Polygala columbica Chodat =Polygala caracasana 

Polygala diversifolia L. =Securidaca diversifolia 

Polygala fendleri Chodat =Polygala exigua var. fendleri 

Polygala funckii Chodat =Polygala tenella 

Polygala gracilis Kunth =Polygala tenella 

Polygala incarnata Aubl. =Polygala adenophora 

Polygala leptocaulis Torr. & A. Gray =Polygala tenella 

Polygala maguirei Wurdack =Polygala timoutoides var. maguirei 

Polygala mollis Kunth =Polygala violacea 

Polygala monticola Kunth =Polygala violacea 

Polygala monticola var. brizoides (A. St.-Hil.) Steyerm. =Polygala violacea 

Polygala paludosa A. St.-Hil. & Moq. [not in flora area] =Polygala tenella 

Polygala peduncularis A. Rich. =Polygala planellasi 

Polygala retifera S.F. Blake =Polygala caracasana 

Polygala sororia Miq. =Polygala trichosperma 

Polygala stipulata Chodat =Polygala membranacea 

Polygala subsecunda S.F. Blake =Polygala brevialata 

Polygala variabilis Kunth =Polygala trichosperma 

Polygala violacea Vahl, non Aubl. =Polygala martiana 

Securidaca complicata Kunth =Securidaca pendula 

Securidaca cordata J.R. Johnst. =Securidaca scandens 

Securidaca corytholobium A.W. Benn. =Securidaca macrophylla 

Securidaca densiflora Oort =Securidaca pubescens 

Securidaca ecristata Wurdack =Securidaca cacumina 

Securidaca fendleri Chodat =Securidaca diversifolia 

Securidaca hostmannii Miq. =Securidaca paniculata var. lasiocarpa 

Securidaca latifolia Benth. =Securidaca pubiflora 

Securidaca major Sagot ex A.W. Benn. =Securidaca pubescens 

AM BO GU 

AM BO GU FG 

AM 

BO 

AM BO 

GU SU FG 

BO 

FG? 

AM GU SU FG 


AM GU?


Securidaca mollis Kunth =Securidaca coriacea 

Securidaca orinocensis Rusby =Securidaca pendula 

Securidaca rivinifolia A. St.-Hil. & Moq. =Securidaca divaricata 

Securidaca rivinifolia var. parvifolia A.W. Benn. =Securidaca divaricata 

Securidaca venosa Rusby =Securidaca diversifolia 

REFERENCES 

Aymard C.,G.A, P.E. Berry, and B. Eriksen. 2004. Polygalaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 

Flora of the Venezuelan Guayana 8: 316-347 

Chodat, R. 1893. Monographia Polygalacearum. Mém. Soc. phys. et d’hist. nat. Genève 31 part 2(2): 1-500. 

Görts-van Rijn, A.R.A. 1974. Notes on Polygalaceae from Suriname. Acta Bot. Neerl. 23(2): 189-191. 

Görts-van Rijn, A.R.A. 1976. Polygalaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 

518-523. 

Jacobs-Brouwer, A. 2002. Polygalaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 585-589. 

Marques, M.d.C.M. 1979. Revisao das espécies do gênero Polygala L. (Polygalaceae) do estado do Rio de Janeiro. Rodriguésia 

31(48): 69-339. 

Marques, M.d.C.M. 1980. Revisao das espécies do gênero Bredemeyera Willd. (Polygalaceae) do Brasil. Rodriguésia 32(54): 

269-321. 

Oort, A.J.P. 1939. Polygalaceae. In A. Pulle, Flora of Suriname 2(1): 406-425. 

Antigonon leptopus Hook. & Arn. [cultivated, naturalized] 

Coccoloba acuminata Kunth 

Coccoloba ascendens Duss ex Lindau 

Coccoloba caracasana Meisn. 

Coccoloba charitostachya Standl. 

Coccoloba conduplicata Maguire 

Coccoloba coronata Jacq. 

Coccoloba cruegeri Lindau 

Coccoloba declinata (Vell.) Mart. 

Coccoloba densifrons Mart. ex Meisn. 

Coccoloba dugandiana A. Fernández 

Coccoloba excelsa Benth. 

Coccoloba fallax Lindau 

Coccoloba gymnorrhachis Sandwith 

Coccoloba latifolia Poir. 

Coccoloba llewelynii R.A. Howard 

Coccoloba lucidula Benth. 

Coccoloba marginata Benth. 

Coccoloba mollis Casar. 

Coccoloba obtusifolia Jacq. 

Coccoloba ochreolata Wedd. 

Coccoloba orinocana R.A. Howard 

Coccoloba ovata Benth. 

Coccoloba padiformis Meisn. 

Coccoloba savannarum Standl. 

Coccoloba schomburgkii Meisn. 

Coccoloba sp. A 

Coccoloba sp. B 

Coccoloba sp. C 

Coccoloba sp. D 

Coccoloba spruceana Lindau 

Coccoloba striata Benth. 

Coccoloba uvifera (L.) L. 

Coccoloba wurdackii R.A. Howard 

Polygonum acuminatum Kunth 

Polygonum ferrugineum Wedd. 

Polygonum glabrum Willd. 

Polygonum hydropiperoides Michx. 

Polygonum meisnerianum Cham. & Schltdl. 

POLYGONACEAE 

Aymard C., G.A. J.Brandbyge, R.A. Howard 


BO GU 


AM 

BO GU 

SU 

BO 

GU SU 

AM BO DA SU FG 

BO GU 

AM BO 


AM BO DA 

BO GU SU FG 


AM BO 

BO GU SU FG 


AM BO SU FG 

AM BO 

AM BO 

AM BO 


GU? 

GU 

AM BO GU 

AM 

AM 

AM 

BO 

AM DA 

AM BO GU 

AM GU SU FG 

AM 


DA FG 



FG

472 


Polygonum meisnerianum var. beyrichianum (Cham. & Schltdl.) Meisn. 

Polygonum punctatum Elliott 

Ruprechtia brachysepala Meisn. 

Ruprechtia brachystachya Meisn. 

Ruprechtia cruegeri Griseb. ex Lindau [expected in Guyana] 

Ruprechtia howardiana Aymard & P.E. Berry 

Ruprechtia laxiflora Meisn. 

Ruprechtia ramiflora (Jacq.) C.A. Mey. 

Ruprechtia tangarana Standl. 

Ruprechtia tenuiflora Benth. 

Symmeria paniculata Benth. 

Triplaris americana L. 

Triplaris caracasana Cham. 

Triplaris cumingiana Fisch. & C.A. Mey. ex C.A. Mey. 

Triplaris weigeltiana (Rchb.) Kuntze 


Blochmannia weigeltiana Rchb. =Triplaris weigeltiana 

Coccoloba bolivarana Ll. Williams [nomen nudum] =Coccoloba llewelynii 

Coccoloba bracteolosa Meisn. =Coccoloba excelsa 

Coccoloba caribaea Urb. =Coccoloba coronata 

Coccoloba caurana Standl. =Coccoloba fallax 

Coccoloba cyclophylla S.F. Blake =Coccoloba caracasana 

Coccoloba excoriata L. [not in flora area] =Coccoloba tenuifolia L. 

Coccoloba grandifolia Jacq. [not in flora area] =Coccoloba pubescens L. 

Coccoloba grandis Benth. =Coccoloba latifolia 

Coccoloba guianensis Meisn. =Coccoloba marginata 

Coccoloba micropunctata Eyma =Coccoloba excelsa 

Coccoloba novogranatensis Lindau =Coccoloba coronata 

Coccoloba paraensis Meisn. =Coccoloba excelsa 

Coccoloba parimensis Benth. =Coccoloba excelsa 

Coccoloba parimensis var. schomburgkii Meisn. =Coccoloba excelsa 

Coccoloba pittieri R. Knuth ex Pittier =Coccoloba striata 

Coccoloba polystachya Wedd. =Coccoloba mollis 

Coccoloba sagotii Lindau =Coccoloba lucidula 

Coccoloba sparsifolia Lindau =Coccoloba ascendens 

Coccoloba trinitatis Lindau =Coccoloba marginata 

Persicaria acuminata (Kunth) M. Gómez =Polygonum acuminatum 

Persicaria densiflora (Meisn.) Moldenke [s.l.] =Polygonum glabrum 

Persicaria glabra (Willd.) M. Gómez =Polygonum glabrum 

Persicaria hydropiperoides (Michx.) Small =Polygonum hydropiperoides 

Persicaria meisneriana (Cham. & Schltdl.) M. Gómez =Polygonum meisnerianum 

Persicaria punctata (Elliott) Small =Polygonum punctatum 

Polygonum acuminatum var. glabrecens Meisn. =Polygonum acuminatum 

Polygonum acuminatum var. weddelii Meisn. =Polygonum acuminatum 

Polygonum declinatum Vell. =Coccoloba declinata 

Polygonum densiflorum Meisn. =Polygonum glabrum 

Polygonum portoricense Bertero ex Small [nom. illeg.] =Polygonum glabrum 

Ruprechtia amentacea Meisn. =Ruprechtia tenuiflora 

Ruprechtia concina Pittier =Ruprechtia ramiflora 

Ruprechtia coriacea (H. Karst.) S.F. Blake =Ruprechtia ramiflora 

Ruprechtia hamanii S.F. Blake =Ruprechtia ramiflora 

Ruprechtia marowynensis Eyma =Ruprechtia brachysepala 

Triplaris americana Aubl., non L. =Triplaris weigeltiana 

Triplaris caracasana var. vargasii Meisn. =Triplaris caracasana 

Triplaris coriacea H. Karst. =Ruprechtia ramiflora 

Triplaris felipensis Wedd. =Triplaris americana 

Triplaris pavonii Meisn. =Triplaris americana 

Triplaris ramiflora Jacq. =Ruprechtia ramiflora 

Triplaris schomburgkiana Benth. =Triplaris americana 

Triplaris surinamensis Cham. =Triplaris weigeltiana 

Triplaris surinamensis var. benthamiana Meisn. =Triplaris weigeltiana 

Triplaris surinamensis var. chamissoana Meisn. =Triplaris weigeltiana 

Triplaris surinamensis var. crassifolia Benth. =Triplaris weigeltiana 

FG 


GU SU FG 

GU 

BO 

AM 

BO GU 

AM BO 

BO DA GU 


AM 

AM 

BO 

BO 

GU SU FG 

BO DA GU SU FG


Triplaris vahliana Fisch. & C.A. Mey. ex C.A. Mey. =Triplaris weigeltiana 

REFERENCES 

Aymard C.,G.A. and R.A. Howard. 2004. Polygonaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Brandbyge, J. 1986. A revision of the genus Triplaris (Polygonaceae). Nord. J. Bot. 6(5): 545-570. 

Brandbyge, J. 2002. Polygonaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


590-592. 

Cocucci, A.E. 1961. Revision del genero Ruprechtia (Polygonaceae). Kurtziana 1: 217-269. 

Eyma, P.J. 1934. Polygonaceae. In A. Pulle, Flora of Suriname 1(1): 49-71. 

Howard, R.A. 1960. Studies in the genus Coccoloba, IX. A conspectus of the South American species. J. Arn. Arb. 41: 213-258, 

357-390. 

Howard, R.A. 1961. Studies in the genus Coccoloba, X. New species and a summary of distribution in South America. J. Arn. 

Arb. 42: 87-95. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Polygonaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 303-309. 

Wilson, K.L. 1990. Some widespread species of Persicaria and their allies. Kew Bull. 45: 621-636. 

Portulaca elatior Mart. ex Rohrb. 

Portulaca halimoides L. 

Portulaca insignis Steyerm. 

Portulaca mucronata Link 

Portulaca oleracea L. [introduced] 

Portulaca pilosa L. 

Portulaca pusilla Kunth 

Portulaca pygmaea Steyerm. 

Portulaca quadrifida L. [introduced?] 

Portulaca rubricaulis Kunth 

Portulaca sedifolia N.E. Br. 

Portulaca teretifolia Kunth 

Portulaca umbraticola Kunth 

Talinum fruticosum (L.) Juss. 

Talinum paniculatum (Jacq.) Gaertn. 

PORTULACACEAE 

J.A. Steyermark† 


Portulaca cayennensis D. Legrand =Portulaca sedifolia 

Portulaca cinerea Poelln. =Portulaca teretifolia 

Portulaca fruticosa L. =Talinum fruticosum 

Portulaca lanata Rich. =Portulaca pilosa 

Portulaca marginata Kunth =Portulaca oleracea 

Portulaca paniculata Jacq. =Talinum paniculatum 

Portulaca portulacastrum L. (Aizoaceae) =Sesuvium portulacastrum 

Portulaca triangularis Jacq. =Talinum fruticosum 

Talinum triangulare (Jacq.) Willd. =Talinum fruticosum 

REFERENCES 

Legrand, C.D. 1962. Las especies americanas de Portulaca. An. Mus. Hist. Nat. Montevideo (ser. 2) 7: 1-147. 

Ooststroom, S.J. van. 1943. Portulacaceae. In A. Pulle, Flora of Suriname 1(1): 486-491. 

Steyermark, J.A. 2005. Portulacaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 376-383. 

Anagallis pumila Sw. 

PRIMULACEAE 



Centunculus pentandrus R. Br. =Anagallis pumila 

Centunculus pumilus (Sw.) Kuntze =Anagallis pumila 

Micropyxis pumila (Sw.) Duby =Anagallis pumila 

BO 

BO 

BO 

AM BO 


AM BO DA GU SU? FG 

AM BO DA 

AM 

GU 

FG? 


AM BO DA FG? 

AM GU SU FG 

BO GU? SU? 

BO GU FG 

AM BO

474 


REFERENCES 

Miller, J.S. and P.E. Berry. 2004. Primulaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 383-383. 

Euplassa chimantensis Steyerm. 

Euplassa glaziovii (Mez) Steyerm. [Mt. Roraima] 

Euplassa pinnata (Lam.) I.M. Johnst. 

Euplassa venezuelana Steyerm. 

Panopsis ornatinervia Steyerm. 

Panopsis parimensis Steyerm. 

Panopsis ptariana Steyerm. 

Panopsis rubescens (Pohl) Rusby 

Panopsis sessilifolia (Rich.) Sandwith 

Panopsis tepuiana Steyerm. 

Roupala chimantensis Steyerm. 

Roupala griotii Steyerm. 

Roupala minima Steyerm. 

Roupala montana Aubl. 

Roupala nitida Rudge 

Roupala obtusata Klotzsch 

Roupala paruensis Steyerm. 

Roupala schulzii Mennega 

Roupala sororopana Steyerm. 

Roupala suaveolens Klotzsch 

PROTEACEAE 



Adenostephanus guianensis Meisn. =Euplassa pinnata 

Andriapetalum cayennensis Klotzsch ex Meisn. =Panopsis sessilifolia 

Andriapetalum rubescens Pohl =Panopsis rubescens 

Andriapetalum sessilifolium (Rich.) Klotzsch =Panopsis sessilifolia 

Euplassa meridionalis Salisb. =Euplassa pinnata 

Panopsis cuaensis Steyerm. =Panopsis rubescens 

Panopsis hameliifolia (Rudge) Salisb. =Panopsis sessilifolia 

Panopsis rubescens var. simulans J.F. Macbr. =Panopsis rubescens 

Panopsis schomburgkii (Klotzsch) Pittier =Roupala suaveolens 

Roupala complicata Kunth =Roupala montana 

Roupala dentata R. Br. =Roupala montana 

Roupala dissimilis Pittier =Roupala montana 

Roupala hameliifolia Rudge =Panopsis sessilifolia 

Roupala laurifolia Salisb. & Knight =Roupala nitida 

Roupala macropoda Klotzsch & H. Karst. =Roupala montana 

Roupala media R. Br. =Roupala montana 

Roupala montana ß complicata (Kunth) Griseb. =Roupala montana 

Roupala montana var. dentata (R. Br.) Sleumer =Roupala montana 

Roupala pinnata Lam. =Euplassa pinnata 

Roupala pullei Mennega =Roupala suaveolens 

Roupala pyrifolia Salisb. & Knight =Roupala montana 

Roupala schomburgkii Klotzsch =Roupala suaveolens 

Roupala sessilifolia Rich. =Panopsis sessilifolia 

REFERENCES 

Mennega, A.M.W. 1934. Proteaceae. In A. Pulle, Flora of Suriname 1(1): 154-157. 

Mennega, A.M.W. 1968. Proteaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 315-320. 

Sleumer, H.O. 1954. Proteaceae americanae. Bot. Jahrb. 76: 139-211. 

Plana, V. 2002. Proteaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


594. 

Steyermark, J.A. 2004. Proteaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 384-393. 

AM 

BO 

BO 

BO 

GU SU FG 

BO 


BO GU SU FG 

BO 

BO 

AM 

BO GU 


FG 

AM GU FG 

AM 

SU 

BO 

AM BO GU


QUIINACEAE 

J.V. Schneider, G. Zizka 

Froesia gereauana J.V. Schneid. & Zizka 

Froesia tricarpa Pires 

Froesia venezuelensis Steyerm. & G.S. Bunting 

Lacunaria crenata (Tul.) A.C. Sm. [expected in lowlands of BO or AM] 

Lacunaria jenmanii (Oliv.) Ducke 

Lacunaria macrostachya (Tul.) A.C. Sm. 

Lacunaria oppositifolia Pires 

Lacunaria panamensis (Standl.) Standl. 

Lacunaria umbonata Pires 

Quiina amazonica A.C. Sm. 

Quiina attenuata J.V. Schneid & Zizka 

Quiina brevensis Pires 

Quiina congesta R.S. Cowan 

Quiina cruegeriana Griseb. 

Quiina florida Tul. 

Quiina guianensis Aubl. 

Quiina indigofera Sandwith 

Quiina integrifolia Pulle 

Quiina leptoclada Tul. 

Quiina longifolia Spruce ex Planch. & Triana 

Quiina macrophylla Tul. 

Quiina obovata Tul. 

Quiina oiapocensis Pires 

Quiina parvifolia Lanj. & Heerdt 

Quiina pteridophylla (Radlk.) Pires 

Quiina rhytidopus Tul. 

Quiina sessilis Choisy 

Quiina tinifolia Planch. & Triana 

Quiina wurdackii Pires 

Quiina yatuensis J.V. Schneid. & Zizka 

Touroulia amazonica Pires & A.S. Foster [Sierra de la Neblina] 

Touroulia guianensis Aubl. 


Lacunaria silvatica (Pulle) A.C. Sm. =Lacunaria jenmanii 

Lacunaria spruceana (Engl.) Pires =Lacunaria oppositifolia 

Quiina acutangula Ducke =Quiina pteridophylla 

Quiina albiflora A.C. Sm. =Quiina obovata 

Quiina crenata Tul. =Lacunaria crenata 

Quiina decaisneana Planch. & Triana =Lacunaria sp. jenmanii? 

Quiina gracilis A.C. Sm. =Quiina tinifolia 

Quiina macrostachya Tul. =Lacunaria macrostachya 

Quiina oblanceolata Sandwith =Quiina obovata 

Quiina poeppigiana Tul. =Quiina florida 

Quiina silvatica Pulle =Lacunaria jenmanii 

Quiina spruceana Engl. =Lacunaria oppositifolia 

Touroulia jenmanii Oliv. =Lacunaria jenmanii 

Touroulia pteridophylla Radlk. =Quiina pteridophylla 

AM 

AM 

AM 

GU? 

GU SU FG 

AM GU SU FG 

AM 

AM 

BO 

SU 

FG 

GU? 

AM 

GU 

AM 

AM 

BO GU 

SU 

SU FG 


BO DA GU 

SU FG 

SU FG 

AM 

AM FG 

BO GU SU FG 

AM BO GU FG 

AM BO GU 

SU 

SU FG 

AM 

AM 

BO GU SU 

FG 

AM BO 

AM 

AM 

AM 


REFERENCES 

Lanjouw, J. & P.F. van Heerdt. 1941. Quiinaceae. In A. Pulle, Flora of Suriname 3(1): 355-365. 

Mori, S.A. 2002. Quiianaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


598. 

Schneider, J. and G. Zizka.1997. Two new species of Quiianaceae (Quiina, Froesia) from the Venezuelan Guayana, and some 

remarks on the genus Froesia Pires. 

Schneider, J.V. & G. Zizka. 2004. Quiinaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 393-407.

476 

Apodanthes caseariae Poit. 

Apodanthes surinamensis Pulle 


RAFFLESIACEAE 

W. Meijer†, G. Yatskievych 


Apodanthes flacourtiae H. Karst. =Apodanthes caseariae 

Apodanthes roraimae Vattimo-Gil =Apodanthes caseariae 

REFERENCES 

Gentry, A.H. 1973. Rafflesiaceae. In R.E. Woodson et al., Flora of Panama. Ann. Missouri Bot. Gard. 60: 17-21. 

Gómez, L.D. 1983. Rafflesiaceae. In Flora Costaricensis, edited by W. Burger. Fieldiana, Bot. n.s. 13: 89–93. 

Meijer, W. 1993. Rafflesiaceae. In The Families and Genera of Vascular Plants, Vol. 2, edited by K. Kubitzi, J.G. Rohwer, and V. 

Bittrich, pp. 557–563. Berlin: Springer-Verlag. 

Mori, S.A. 2002. Rafflesiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


598. 

Vattimo-Gil, I. de. 1956. Notes on Apodanthes caseariae Poit. and Pilostyles calliandrae (Gardn.) R. Br. Not. Syst. 15(2): 225- 

229. 

Vattimo-Gil, I. de. 1971. Contribuiçao ao conhecimento de tribo Apodanthea (sic) R. Br. Parte I - Conspecto das espécies. 

Rodriguesia 26(38): 37-62. 

Vattimo-Gil, I. de. 1973. Notas sobre o gênero Apodanthes Poit. com descrição de duas novas espécies. Rev. Brasil. Biol. 33: 

135–141. 

Vattimo-Gil, I. de. 1978. Uma nova espécie de Apodanthes Pot. (Rafflesiaceae). Rodriguésia 29: 47–51. 1978. 

Yatskievych, G. & W. Meijer. 2004. Rafflesiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


RANUNCULACEAE 

P.E. Berry 

Clematis dioica var. brasiliana (DC.) Eichler [not in flora area, expected in FG] 

Clematis guadeloupae Pers. 

Clematis guadeloupae var. guadeloupae [Expected in GU] 

Clematis populifolia Turcz. 


Clematis caracasana Humb. & Bonpl. ex DC. =Clematis guadeloupae var. guadeloupae 

Clematis caripensis Kunth =Clematis guadeloupae var. guadeloupae 

Clematis floribunda Triana & Planch. =Clematis populifolia 

Clematis goudotiana Triana & Planch. =Clematis populifolia 

REFERENCES 

Berry, P.E. 2004. Ranunculaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

8: 410-413. 

Lourteig, A. 1952. Ranunculáceas de Sudamérica templada. Darwiniana 9: 397-609. 

Lourteig, A. 1971. Ranunculaceae. In T. Lasser, Flora de Venezuela 3(1): 29-51. 

Rhabdodendron amazonicum (Spruce ex Benth.) Huber 

RHABDODENDRACEAE 


Lecostemon amazonicum Spruce ex Benth. =Rhabdodendron amazonicum 

Lecostemon sylvestre Gleason =Rhabdodendron amazonicum 

Lecostemon terniflorum Moç. & Sessé ex DC. (Elaeocarpaceae) =Sloanea terniflora 

Rhabdodendron crassipes (Spruce ex Benth.) Huber =Rhabdodendron amazonicum 

Rhabdodendron crassipes var. cayenense Benth. =Rhabdodendron amazonicum 

Rhabdodendron sylvestre (Gleason) Maguire =Rhabdodendron amazonicum 

BO GU SU FG 

SU 

REFERENCES 

Mori, S.A. 2002. Rhabdodendraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 598-600. 

Prance, G.T. 1972. Rhabdodendraceae. Flora Neotropica Monogr. 11: 1-22. 

BO 

BO 

BO 

GU SU FG


RHAMNACEAE 

P.E. Berry, J.A. Steyermark† 

Ampelozizyphus amazonicus Ducke 

Colubrina glandulosa Perkins 

Colubrina glandulosa var. glandulosa 

Gouania blanchetiana Miq. 

Gouania canescens Rich. 

Gouania colurnifolia Reissek 

Gouania cornifolia Reissek 

Gouania discolor Benth. 

Gouania frangulifolia (Willd. ex Roem. & Schult.) Radlk. 

Gouania lupuloides (L.) Urb. [s.l.] 

Gouania polygama (Jacq.) Urb. 

Gouania striata Rich. 

Gouania velutina Reissek 

Gouania virgata Reissek [“Guiana”] 

Gouania wurdackii Steyerm. 

Rhamnus acuminata Maguire & Steyerm. [Sierra de la Neblina] 

Rhamnus chimantensis Steyerm. & Maguire 

Rhamnus longipes Steyerm. 

Rhamnus marahuacensis Steyerm. & Maguire 

Rhamnus neblinensis Maguire & Steyerm. [Sierra de la Neblina] 

Rhamnus psilocarpa Maguire & Steyerm. [Sierra de la Neblina] 

Rhamnus sipapoensis Steyerm. 

Rhamnus ulei Pilg. [Mt. Roraima] 

Ziziphus cinnamomum Triana & Planch. 

Ziziphus jujuba Mill. [cultivated; records may be referable to Ziziphus 

mauritiana] 

Ziziphus mauritiana Lam. [cultivated, naturalized] 

Ziziphus saeri Pittier 


Gouania hypochroa Reissek =Gouania discolor 

Gouania pubescens Poir. =Gouania polygama 

Gouania tomentosa Jacq. =Gouania polygama 

Gouania ulei Pilg. =Gouania discolor 

Rhamnus cumanensis Loefl. (Boraginaceae) =Bourreria cumanensis 

Rhamnus polygamus Jacq. =Gouania polygama 

Trisecus frangulaefolius Willd. ex Roem. & Schult. =Gouania frangulifolia 

Ziziphus jujuba (L.) Lam., non Mill. =Ziziphus mauritiana 

REFERENCES 

Heald, S.V. 2002. Rhamnaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


600-602. 

Johnston, M.C. 1971. Revision of Colubrina. Brittonia 23: 2-53. 

Johnston, M.C. & L.V.A. Johnston. 1978. Rhamnus. Flora Neotropica Monogr. 20: 1-96. 

Lanjouw, J. 1932. Rhamnaceae. In A. Pulle, Flora of Suriname 2(1): 102-106. 

Steyermark, J.A. and P.E. Berry. 2004. Rhamnaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Suessenguth, K. 1953. Rhamnaceae. In A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien 20d: 7-173. 

Cassipourea elliptica (Sw.) Poir. [=Cassipourea guianensis?] 

Cassipourea guianensis Aubl. 

Cassipourea lasiocalyx Alston 

Cassipourea peruviana Alston 

Cassipourea serrata Benth. 

Rhizophora x harrisonii Leechm. [putative hybrid species] 

Rhizophora mangle L. 

RHIZOPHORACEAE 



BO GU 

GU 


FG 

BO GU 

AM 

AM BO 

AM BO FG 

FG 


FG 

GU SU 

AM BO 

AM 

BO 

AM 

AM 

AM 

AM 

AM 

BO GU? 

AM GU FG 

GU SU FG 


BO 

SU FG 


GU 

GU SU 

GU 

DA GU SU FG 

DA GU SU FG

478 


Rhizophora racemosa G. Mey. 

Sterigmapetalum chrysophyllum Aymard & Cuello 

Sterigmapetalum colombianum Monach. 

Sterigmapetalum exappendiculatum Steyerm. & Liesner [perhaps expected in 

GU] 

Sterigmapetalum guianense Steyerm. 

Sterigmapetalum guianense subsp. guianense [expected in GU] 

Sterigmapetalum guianense subsp. ichunense Steyerm. & Liesner 

Sterigmapetalum resinosum Steyerm. & Liesner 


Rhizophora mangle var. racemosa (G. Mey.) Engl. =Rhizophora racemosa 

REFERENCES 

(see also Anisophylleaceae) 

Alston, A.H.G. 1925. Revision of Cassipourea. Kew Bull. 1925: 241-276. 

Breteler, F.J. 1969. The Atlantic species of Rhizophora. Acta Bot. Neerl. 18: 434-444. 

Ding Hou. 1960. A review of the genus Rhizophora. Blumea 10: 625-634. 

Jonker, F.P. 1942. Rhizophoraceae. In A. Pulle, Flora of Suriname 3(2): 35-43. 

Juncosa, A.M. & P.B. Tomlinson. 1988. A historical and taxonomic synopsis of Rhizophoraceae and Anisophylleaceae. Ann. 

Missouri Bot. Gard. 75: 1278-1295. 

Mori, S.A. 2002. Rhizophoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


603-604. 

Prance, G.T. et al. 1975. Revisao taxonômica das espécies amazonicas de Rhizóphoraceae. Acta Amazonica 5: 5-22. 

Steyermark, J.A. 2004. Rhizophoraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 8: 484-490. 

Steyermark, J.A. & R. Liesner. 1983. Revision of the genus Sterigmapetalum. Ann. Missouri Bot. Gard. 70: 179-193. 

Hesperomeles obtusifolia (Pers.) Lindl. 

Hesperomeles obtusifolia var. obtusifolia 

Prunus accumulans (Koehne) C.L. Li & Aymard 

Prunus amplifolia Pilg. 

Prunus espinozana C.L. Li 

Prunus lichoana Aymard 

Prunus myrtifolia (L.) Urb. 

Prunus myrtifolia var. myrtifolia 

Prunus wurdackii C.L. Li 

Rubus floribundus Kunth 

Rubus guyanensis Focke 

Rubus urticifolius Poir. 

ROSACEAE 

G.A. Aymard, N.L. Cuello A. 


Crataegus obtusifolia Pers. =Hesperomeles obtusifolia 

Hesperomeles heterophylla (Ruiz & Pav.) Hook. =Hesperomeles obtusifolia 

Mespilus heterophylla Ruiz & Pav. =Hesperomeles obtusifolia 

Prunus myrtifolia var. accumulans Koehne =Prunus accumulans 

Prunus sphaerocarpa Sw. =Prunus myrtifolia 

Rubus robustus C. Presl =Rubus floribundus 

Rubus schomburgkii Klotzsch [nomen nudum] =Rubus guyanensis 

DA GU SU FG 

REFERENCES 

Aymard, G.A. & N.L. Cuello A. 2004. Rosaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Kleinhoonte, A. 1939. Rosaceae. In A. Pulle, Flora of Suriname 2(1): 426-456. 

Mitchell, J.D. 2002. Rosaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


606. 

AM 

BO 

SU 

AM BO SU 

BO SU 

AM BO 

BO GU 

AM 

BO 

BO 

BO GU SU FG 

BO 

BO 

BO GU SU FG 

BO GU SU FG 

BO 

AM BO 

BO GU 

AM BO


RUBIACEAE 

E. Araujo da Anunciação, C. Bestetti Costa, R. Cortés, P.G. Delprete, J.H. Kirkbride, 

C. Persson, J.A. Steyermark†, C.M. Taylor, A. Vincentini, D. Zappi 

Alibertia bertierifolia K. Schum. 

Alibertia edulis (Rich.) A. Rich. ex DC. 

Alibertia edulis var. edulis 

Alibertia edulis var. obtusiuscula (Steyerm.) Delprete & C. Perss. 

Alibertia latifolia (Benth.) K. Schum. 

Alibertia latifolia var. latifolia 

Alseis floribunda Schott 

Alseis labatioides H. Karst. ex K. Schum. 

Alseis longifolia Ducke 

Alseis longifolia var. pentamera Bremek. 

Alseis smithii Standl. 

Amaioua brevidentata Steyerm. 

Amaioua corymbosa Kunth 

Amaioua guianensis Aubl. 

Amaioua guianensis var. guianensis 

Amaioua guianensis var. macrantha Steyerm. [Sierra de la Neblina] 

Amphidasya neblinae Steyerm. [Sierra de la Neblina] 

Antirhea triflora J.H. Kirkbr. 

Aphanocarpus steyermarkii (Standl.) Steyerm. 

Bathysa bathysoides (Steyerm.) Delprete 

Bertiera diversiramea Steyerm. 

Bertiera guianensis Aubl. 

Bertiera parviflora Spruce ex K. Schum. 

Borreria alata (Aubl.) DC. 

Borreria bolivarensis Steyerm. 

Borreria capitata (Ruiz & Pav.) DC. 

Borreria capitata var. capitata 

Borreria capitata var. suaveolens (G. Mey.) Steyerm. 

Borreria capitata var. tenella (Kunth) Steyerm. 

Borreria cataractarum Steyerm. 

Borreria confertifolia Steyerm. 

Borreria densiflora DC. 

Borreria hispida Spruce ex K. Schum. 

Borreria intricata Steyerm. 

Borreria jangouxii Steyerm. 

Borreria latifolia (Aubl.) K. Schum. 

Borreria macrocephala Standl. & Steyerm. 

Borreria ovalifolia M. Martens & Galeotti 

Borreria prostrata (Aubl.) Miq. 

Borreria pygmaea Spruce ex K. Schum. 

Borreria remota (Lam.) Bacigalupo & E.L. Cabral 

Borreria repens DC. 

Borreria verticillata (L.) G. Mey. 

Borreria wurdackii Steyerm. 

Borreria xanthophylla Bremek. 

Bothriospora corymbosa (Benth.) Hook. f. 

Botryarrhena pendula Ducke 

Botryarrhena venezuelensis Steyerm. 

Calycophyllum obovatum (Ducke) Ducke 

Calycophyllum venezuelense Steyerm. 

Capirona decorticans Spruce 

Chalepophyllum guianense Hook. f. 

Chimarrhis brevipes Steyerm. 

Chimarrhis cymosa Jacq. 

Chimarrhis microcarpa Standl. 

Chimarrhis turbinata DC. 

Chiococca alba (L.) Hitchc. 

Chiococca alba subsp. alba 

Chiococca auyantepuiensis Steyerm. 

Chiococca lucens Standl. & Steyerm. 

AM BO GU 



AM BO GU 


GU SU FG 

FG 

BO DA GU 

GU SU FG 

GU SU FG 

GU 

GU? FG? 




AM 

AM 

FG 

BO 

AM BO 

BO GU SU FG 


AM 

AM GU SU FG 

BO 



GU SU 


BO GU 

AM BO GU 

BO GU SU FG 


AM 

AM 


AM 

GU 


AM BO 




AM 

SU 

GU 

FG 

AM 

AM BO GU 

AM BO GU 

AM GU SU FG 

BO GU 

AM 

GU 


GU SU FG 

BO GU FG 

GU FG 

BO 

BO

480 


Chiococca multipedunculata Steyerm. 

Chiococca nitida Benth. 

Chiococca nitida var. amazonica Müll. Arg. 

Chiococca nitida var. chimantensis Steyerm. 

Chiococca nitida var. nitida 

Chiococca pubescens Humb. & Bonpl. ex Roem. & Schult. [expected in GU] 

Chomelia caurensis (Standl.) Steyerm. 

Chomelia delascioi Steyerm. 

Chomelia glabricalyx Steyerm. 

Chomelia malaneoides Müll. Arg. 

Chomelia monachinoi Steyerm. 

Chomelia obtusa Cham. & Schltdl. [possibly misident. of C. angustifolia] 

Chomelia polyantha S.F. Blake 

Chomelia schomburgkii Steyerm. 

Chomelia splitgerberi Bremek. 

Chomelia stergiosii Steyerm. 

Chomelia tenuiflora Benth. 

Chomelia volubilis (Standl.) Steyerm. 

Cinchonopsis amazonica (Standl.) L. Andersson 

Coccochondra laevis (Steyerm.) Rauschert 

Coccochondra laevis subsp. laevis 

Coccochondra laevis subsp. maigualidae J.H. Kirkbr. 

Coccocypselum aureum (Spreng.) Cham. & Schltdl. 

Coccocypselum condalia Pers. 

Coccocypselum guianense (Aubl.) K. Schum. 

Coccocypselum herbaceum DC. 

Coccocypselum hirsutum Bartl. ex DC. 

Coccocypselum huberi Steyerm. 

Coccocypselum lanceolatum (Ruiz & Pav.) Pers. 

Coffea arabica L. [cultivated or escaped from cultivation] 

Coffea liberica W. Bull ex Hiern [cultivated or escaped from cultivation] 

Cordiera myrciifolia (Spruce ex K. Schum.) C. Perss. & Delprete 

Cordiera triflora A. Rich. 

Coryphothamnus auyantepuiensis (Steyerm.) Steyerm. 

Cosmibuena grandiflora (Ruiz & Pav.) Rusby 

Coussarea amapaensis Steyerm. 

Coussarea brevicaulis K. Krause [nomen nudum] 

Coussarea evoluta Steyerm. [Sierra de la Neblina] 

Coussarea fanshawei Steyerm. 

Coussarea grandis Müll. Arg. 

Coussarea granvillei Delprete & B.M. Boom 

Coussarea hallei Steyerm. 

Coussarea hirticalyx Standl. 

Coussarea hirticalyx var. glabrior Steyerm. 

Coussarea krukovii Standl. 

Coussarea lasseri Steyerm. 

Coussarea leptoloba (Spreng. ex Benth. & Hook. f.) Müll. Arg. 

Coussarea leptophragma Müll. Arg. 

Coussarea liesneri Steyerm. 

Coussarea machadoana Standl. 

Coussarea mapourioides Bremek. 

Coussarea micrococca Bremek. 

Coussarea ovalis Standl. 

Coussarea paniculata (Vahl) Standl. 

Coussarea racemosa A. Rich. 

Coussarea revoluta Steyerm. 

Coussarea sp. A [Sierra de la Neblina] 

Coussarea surinamensis Bremek. 

Coussarea violacea Aubl. 

Coutarea hexandra (Jacq.) K. Schum. 

Coutarea hexandra var. hexandra 

Declieuxia fruticosa (Willd. ex Roem. & Schult.) Kuntze 

Declieuxia tenuiflora (Willd. ex Roem. & Schult.) Steyerm. & J.H. Kirkbr. 

Dendrosipanea revoluta Steyerm. 

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AM


Dendrosipanea spigelioides Ducke 

Didymochlamys connellii N.E. Br. 

Diodia apiculata (Willd. ex Roem. & Schult.) K. Schum. 

Diodia hyssopifolia (Willd. ex Roem. & Schult.) Cham. & Schltdl. 

Diodia kuntzei K. Schum. 

Diodia multiflora DC. 

Diodia ocymifolia (Willd. ex Roem. & Schult.) Bremek. 

Diodia pulchristipula Bremek. 

Diodia radula (Willd. & Hoffmanns. ex Roem. & Schult.) Cham. & Schltdl. 

Diodia radula subsp. venezuelensis Steyerm. 

Diodia sarmentosa Sw. 

Diodia spicata Miq. 

Diodia teres Walter 

Duidania montana Standl. 

Duroia amapana Steyerm. 

Duroia aquatica (Aubl.) Bremek. 

Duroia bolivarensis Steyerm. 

Duroia eriopila L. f. 

Duroia eriopila var. eriopila 

Duroia eriopila var. tafelbergensis Steyerm. 

Duroia fusifera Hook. f. ex K. Schum. 

Duroia genipoides Hook. f. ex K. Schum. 

Duroia gransabanensis Steyerm. 

Duroia kotchubaeoides Steyerm. 

Duroia longiflora Ducke 

Duroia maguirei Steyerm. 

Duroia maguirei var. maguirei 

Duroia maguirei var. patentinervia Steyerm. 

Duroia martiniana (Miers) Bremek. 

Duroia merumensis Steyerm. 

Duroia micrantha (Ladbr.) Zarucchi & J.H. Kirkbr. 

Duroia nitida Steyerm. 

Duroia paruensis Steyerm. 

Duroia plicata Benoist 

Duroia retrorsipila Steyerm. 

Duroia saccifera (Mart. ex Roem. & Schult.) Hook. f. ex K. Schum. 

Duroia spraguei Wernham 

Duroia strigosa Steyerm. 

Duroia velutina Hook. f. ex K. Schum. [expected in AM] 

Elaeagia maguirei Standl. 

Elaeagia maguirei var. maguirei 

Elaeagia maguirei var. pubens Steyerm. 

Emmeorhiza umbellata (Spreng.) K. Schum. 

Emmeorhiza umbellata subsp. septentrionalis Steyerm. 

Faramea angustifolia Spruce ex Müll. Arg. 

Faramea anisocalyx Poepp. & Endl. 

Faramea berryi Steyerm. 

Faramea boomii Steyerm. [Sierra de la Neblina] 

Faramea brevipes Steyerm. 

Faramea calyciflora A. Rich. ex DC. 

Faramea capillipes Müll. Arg. [specimens in Guianas may represent 

misidentifications] 

Faramea cardonae Standl. & Steyerm. 

Faramea corymbosa Aubl. 

Faramea costata Steyerm. 

Faramea crassifolia Benth. 

Faramea cyanea Müll. Arg. 

Faramea egregia Sandwith 

Faramea fanshawei Steyerm. 

Faramea guianensis (Aubl.) Bremek. 

Faramea irwinii Steyerm. 

Faramea lourteigiana Steyerm. 

Faramea maguirei Steyerm. 

Faramea morilloi Steyerm. 

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482 


Faramea multiflora A. Rich. ex DC. 

Faramea neblinae Steyerm. [Sierra de la Neblina] 

Faramea occidentalis (L.) A. Rich. 

Faramea occidentalis subsp. occidentalis 

Faramea orinocensis Standl. 

Faramea paniculata (Aubl.) Benth. [=Coussarea sp.?] 

Faramea parvibractea Steyerm. 

Faramea polytriadophora Bremek. 

Faramea quadricostata Bremek., emend. Steyerm. 

Faramea quinqueflora Poepp. & Endl. 

Faramea sessiliflora Aubl. 

Faramea sessiliflora var. pedunculata Bremek. 

Faramea sessiliflora var. sessiliflora 

Faramea sessilifolia (Kunth) DC. 

Faramea spathacea Müll. Arg. ex Standl. 

Faramea stenopetala Mart. 

Faramea tamberlikiana Müll. Arg. 

Faramea tamberlikiana subsp. tamberlikiana 

Faramea tinguana Müll. Arg. 

Faramea torquata Müll. Arg. 

Faramea yavitensis Steyerm. 

Faramea yutajensis Steyerm. 

Ferdinandusa boomii Steyerm. [Sierra de la Neblina] 

Ferdinandusa goudotiana K. Schum. 

Ferdinandusa goudotiana var. eciliata Steyerm. 

Ferdinandusa goudotiana var. goudotiana [Mt. Roraima] 

Ferdinandusa goudotiana var. psilocarpa Steyerm. 

Ferdinandusa guainiae Spruce ex K. Schum. 

Ferdinandusa neblinensis Steyerm. [Sierra de la Neblina] 

Ferdinandusa paraensis Ducke 

Ferdinandusa rudgeoides (Benth.) Wedd. 

Ferdinandusa sprucei K. Schum. 

Ferdinandusa uaupensis Spruce ex K. Schum. 

Galium hypocarpium (L.) Endl. ex Griseb. 

Galium surinamense Dempster 

Genipa americana L. 

Genipa americana var. americana 

Genipa americana var. caruto (Kunth) K. Schum. 

Genipa spruceana Steyerm. [=Genipa americana L.?] 

Geophila cordifolia Miq. 

Geophila cordifolia var. cordifolia 

Geophila gracilis (Ruiz & Pav.) DC. 

Geophila macropoda (Ruiz & Pav.) DC. 

Geophila orbicularis (Müll. Arg.) Steyerm. 

Geophila orbicularis var. neblinae Steyerm. [Sierra de la Neblina] 

Geophila orbicularis var. orbicularis 

Geophila repens (L.) I.M. Johnst. 

Geophila tenuis (Müll. Arg.) Standl. 

Gleasonia duidana Standl. 

Gleasonia duidana var. duidana 

Gleasonia duidana var. latifolia Steyerm. 

Gonzalagunia bunchosioides Standl. 

Gonzalagunia cornifolia (Kunth) Standl. 

Gonzalagunia dicocca Cham. & Schltdl. [var. guianensis Steyerm.] 

Gonzalagunia spicata (Lamb.) M. Gómez 

Gonzalagunia surinamensis Bremek. 

Guettarda acreana K. Krause 

Guettarda argentea Lam. 

Guettarda aromatica Poepp. & Endl. 

Guettarda divaricata (Humb. & Bonpl. ex Roem. & Schult.) Standl. 

Guettarda fanshawei Steyerm. 

Guettarda macrantha Benth. 

Guettarda malacophylla Standl. 

Guettarda pohliana Müll. Arg. 


AM 


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AM BO 

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Guettarda roupalifolia Rusby 

Guettarda spruceana Müll. Arg. 

Guettarda viburnoides Cham. & Schltdl. 

Hamelia axillaris Sw. 

Hamelia patens Jacq. 

Henriquezia jenmanii K. Schum. 

Henriquezia nitida Spruce ex Benth. 

Henriquezia nitida var. nitida 

Henriquezia verticillata Spruce ex Benth. 

Hillia foldatsii Steyerm. 

Hillia illustris (Vell.) K. Schum. 

Hillia maguirei Steyerm. 

Hillia parasitica Jacq. 

Hillia psammophila Steyerm. 

Hillia rivalis C.M. Taylor 

Hillia ulei K. Krause 

Hoffmannia megistophylla Standl. 

Holstianthus barbigularis Steyerm. 

Ibetralia surinamensis Bremek. 

Isertia bullata K. Schum., emend. Delprete 

Isertia coccinea (Aubl.) J.F. Gmel. 

Isertia hypoleuca Benth. 

Isertia parviflora Vahl 

Isertia rosea Spruce ex K. Schum. 

Isertia spiciformis DC. 

Isertia verrucosa (Bonpl.) Standl. 

Isertia wilhelminensis Steyerm. 

Ixora acuminatissima Müll. Arg. 

Ixora aluminicola Steyerm. 

Ixora coccinea L. [cultivated or escaped from cultivation] 

Ixora davisii Sandwith 

Ixora ferrea (Jacq.) Benth. 

Ixora graciliflora Benth. 

Ixora intensa K. Krause 

Ixora intropilosa Steyerm. 

Ixora panurensis Müll. Arg. 

Ixora parviflora Vahl 

Ixora piresii Steyerm. 

Ixora potaroensis Steyerm. 

Ixora pubescens Willd. ex Schult. & Schult. f. 

Ixora sandwithiana Steyerm. 

Ixora schomburgkiana Benth. 

Ixora ulei K. Krause 

Ixora versteegii Bremek. 

Ixora yavitensis Steyerm. 

Kutchubaea insignis Fisch. ex DC. 

Kutchubaea longiloba Steyerm. 

Kutchubaea micrantha Steyerm. 

Kutchubaea morilloi Steyerm. 

Kutchubaea neblinensis Steyerm. [Sierra de la Neblina] 

Kutchubaea sericantha Standl. 

Ladenbergia amazonensis Ducke 

Ladenbergia lambertiana (A. Braun ex Mart.) Klotzsch 

Ladenbergia oblongifolia (Mutis) L. Andersson 

Ladenbergia sp. A 

Limnosipanea palustris (Seem.) Hook. f. 

Limnosipanea spruceana Hook. f. 

Machaonia brasiliensis (Hoffmanns. ex Humb.) Cham. & Schltdl. 

Maguireocharis neblinae Steyerm. [Sierra de la Neblina] 

Maguireothamnus speciosus (N.E. Br.) Steyerm. 

Maguireothamnus speciosus subsp. jauaensis Steyerm. 

Maguireothamnus speciosus subsp. speciosus 

Maguireothamnus tatei (Standl.) Steyerm. 

Malanea auyantepuiensis Steyerm. 

FG 

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AM FG 

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BO GU 

BO 

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AM 

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484 


Malanea chimantensis Steyerm. 

Malanea ciliolata Steyerm. 

Malanea cruzii Steyerm. 

Malanea gabrielensis Müll. Arg. 

Malanea guaiquinimensis Steyerm. 

Malanea hypoleuca Steyerm. 

Malanea jauaensis Steyerm. 

Malanea macrophylla Griseb. 

Malanea macrophylla var. macrophylla 

Malanea microphylla Standl. & Steyerm. 

Malanea obovata Hochr. 

Malanea ptariensis Steyerm. 

Malanea sarmentosa Aubl. 

Malanea schomburgkii Steyerm. 

Malanea setulosa Steyerm. 

Malanea sipapoensis Steyerm. 

Malanea tafelbergensis Steyerm. 

Malanea ueiensis Steyerm. 

Malanea ursina Standl. 

Manettia alba (Aubl.) Wernham 

Manettia calycosa Griseb. 

Manettia calycosa var. karsteniana K. Schum. 

Manettia coccinea (Aubl.) Willd. 

Manettia reclinata Mutis ex L. 

Merumea coccocypseloides Steyerm. 

Merumea plicata Steyerm. 

Mitracarpus diffusus (Willd. ex Roem. & Schult.) Cham. & Schltdl. 

Mitracarpus frigidus (Willd. ex Roem. & Schult.) K. Schum. 

Mitracarpus frigidus var. fruticosus (Standl.) Steyerm. 

Mitracarpus frigidus var. orinocensis Steyerm. 

Mitracarpus hirtus (L.) DC. 

Mitracarpus humboldtianus Cham. & Schltdl. 

Mitracarpus microspermus K. Schum. 

Mitracarpus parvulus K. Schum. 

Mitracarpus scabrellus Benth. 

Mitracarpus sp. A 

Morinda aurantiaca (K. Krause) Steyerm. 

Morinda brachycalyx (Bremek.) Steyerm. 

Morinda calycina (Benth.) Steyerm. 

Morinda citrifolia L. [cultivated, naturalized] 

Morinda debilis (Sandwith) Steyerm. 

Morinda longiloba (Steyerm.) Steyerm. 

Morinda longipedunculata Steyerm. [Sierra de la Neblina] 

Morinda peduncularis Kunth 

Morinda sparsiflora (Steyerm.) Steyerm. 

Morinda surinamensis (Bremek.) Steyerm. [possibly =M. tenuiflora] 

Morinda tenuiflora (Benth.) Steyerm. 

Morinda venezuelensis (Steyerm.) Steyerm. 

Neblinathamnus argyreus Steyerm. [Sierra de la Neblina] 

Neblinathamnus brasiliensis Steyerm. 

Neobertiera gracilis Wernham 

Notopleura aligera (Steyerm.) C.M. Taylor 

Notopleura aneurophylla (Standl.) C.M. Taylor 

Notopleura anomothyrsa (K. Schum. & Donn. Sm.) C.M. Taylor 

Notopleura crassa (Benth.) C.M. Taylor [Mt. Roraima] 

Notopleura guadalupensis (DC.) C.M. Taylor 

Notopleura macrophylla (Ruiz & Pav.) C.M. Taylor 

Notopleura merumensis (Steyerm.) C.M. Taylor 

Notopleura microbracteata (Steyerm.) C.M. Taylor 

Notopleura multiramosa (Steyerm.) C.M. Taylor 

Notopleura perpapillifera (Steyerm.) C.M. Taylor 

Notopleura sandwithiana (Steyerm.) C.M. Taylor 

Notopleura saülensis (Steyerm.) C.M. Taylor 

Notopleura tapajozensis (Standl.) Bremek. 

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AM BO 

BO 

GU 

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BO 

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Notopleura thesceloantha (Steyerm.) C.M. Taylor 

Notopleura uliginosa (Sw.) Bremek. 

Oldenlandia corymbosa L. [=Hedyotis corymbosa (L.) Lam.] 

Oldenlandia filicaulis K. Schum. 

Oldenlandia lancifolia (Schumach.) DC. [naturalized] [=Hedyotis lancifolia 

Schumacher] 

Oldenlandia tenuis K. Schum. 

Oldenlandiopsis callitrichoides (Griseb.) Terrell & W.H. Lewis [introduced] 

Pagamea anisophylla Standl. & Steyerm. 

Pagamea capitata Benth. 

Pagamea capitata subsp. capitata 

Pagamea capitata subsp. conferta (Standl.) Steyerm. 

Pagamea capitata subsp. thibaudiifolia (Wernham) Steyerm. 

Pagamea coriacea Spruce ex Benth. 

Pagamea coriacea var. acuta Steyerm. 

Pagamea coriacea var. coriacea 

Pagamea diceras Steyerm. 

Pagamea duidana Standl. & Steyerm. 

Pagamea guianensis Aubl. 

Pagamea guianensis var. guianensis 

Pagamea hirsuta Spruce ex Benth. 

Pagamea jauaensis Steyerm. 

Pagamea magniflora Steyerm. 

Pagamea montana Gleason & Standl. 

Pagamea pauciflora Standl. & Steyerm. 

Pagamea plicata Spruce ex Benth. 

Pagamea plicatiformis Steyerm. 

Pagamea sessiliflora Spruce ex Benth. 

Pagamea standleyana Steyerm. 

Pagamea thyrsiflora Spruce ex Benth. 

Pagamea velutina Steyerm. 

Pagameopsis garryoides (Standl.) Steyerm. 

Pagameopsis maguirei Steyerm. 

Palicourea amapaensis Steyerm. 

Palicourea anceps Standl. 

Palicourea brachyloba (Müll. Arg.) B.M. Boom 

Palicourea bracteosa Standl. 

Palicourea calophylla DC. 

Palicourea cardonae Steyerm. 

Palicourea corymbifera (Müll. Arg.) Standl. 

Palicourea crocea (Sw.) Roem. & Schult. 

Palicourea croceoides Desv. ex Ham. [=Palicourea riparia] 

Palicourea fastigiata Kunth 

Palicourea foldatsii Steyerm. 

Palicourea formosa (A. Rich.) Lemée 

Palicourea glabriflora Steyerm. 

Palicourea grandiflora (Kunth) Standl. 

Palicourea grandifolia (Willd. ex Roem. & Schult.) Standl. 

Palicourea guianensis Aubl. 

Palicourea guianensis subsp. guianensis 

Palicourea huberi Steyerm. 

Palicourea irwinii Steyerm. 

Palicourea lancigera (Standl.) Steyerm. 

Palicourea longiflora DC. 

Palicourea longistipulata (Müll. Arg.) Standl. 

Palicourea longistipulata subsp. chrysorrhachis (Bremek.) Steyerm. 

Palicourea longistipulata subsp. longistipulata 

Palicourea marcgravii A. St.-Hil. 

Palicourea nitidella (Müll. Arg.) Standl. 

Palicourea obtusata K. Krause 

Palicourea ottohuberi J.H. Kirkbr. 

Palicourea pedunculosa (A. Rich.) DC. ex Lemée [dubious name] 

Palicourea pensilis J.H. Kirkbr. 

Palicourea perquadrangularis Wernham 

AM 


AM 

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BO GU 

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BO GU SU 

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486 

Palicourea perquadrangularis var. guayanensis Steyerm. 

Palicourea quadrifolia (Rudge) DC. 

Palicourea quadrifolia subsp. quadrifolia 

Palicourea rigida Kunth 

Palicourea sp. A 

Palicourea tepuicola Steyerm. 

Palicourea triphylla DC. 

Palicourea wurdackiana J.H. Kirkbr. 

Patabea tenuiflora DC. [=Psychotria sp.] 

Patima guianensis Aubl. 

Perama dichotoma Poepp. 

Perama dichotoma var. dichotoma 

Perama dichotoma var. monocephala Steyerm. 

Perama galioides (Kunth) Poir. 

Perama galioides var. galioides 

Perama hirsuta Aubl. 

Perama plantaginea (Kunth) Hook. f. 

Platycarpum decipiens Woodson & Steyerm. 

Platycarpum duckei Steyerm. 

Platycarpum eglandulosum Steyerm. 

Platycarpum maguirei Steyerm. [Sierra de la Neblina] 

Platycarpum negrense Ducke 

Platycarpum negrense var. glaucum G.K. Rogers 

Platycarpum orinocense Bonpl. 

Platycarpum orinocense var. grandiflorum Steyerm. 

Platycarpum orinocense var. orinocense 

Platycarpum rhododactylum Woodson & Steyerm. 

Platycarpum rugosum Steyerm. 

Platycarpum schultesii Steyerm. 

Platycarpum schultesii var. zarucchii G.K. Rogers 

Posoqueria coriacea M. Martens & Galeotti 

Posoqueria latifolia (Rudge) Roem. & Schult. 

Posoqueria latifolia subsp. gracilis (Rudge) Steyerm. 

Posoqueria latifolia subsp. latifolia 

Posoqueria longiflora Aubl. 

Posoqueria panamensis (Walp. & Duchass.) Walp. 

Posoqueria trinitatis DC. 

Posoqueria williamsii Steyerm. 

Psychotria acuminata Benth. 

Psychotria adderleyi Steyerm. 

Psychotria adenophora Steyerm. 

Psychotria aetantha (Sandwith) Steyerm. 

Psychotria alba Ruiz & Pav. 

Psychotria albert-smithii Standl. 

Psychotria alloantha Steyerm. 

Psychotria altsonii (Sandwith) Steyerm. 

Psychotria amplectens Benth. 

Psychotria anartiothrix Steyerm. 

Psychotria anceps Kunth 

Psychotria anceps var. anceps 

Psychotria antennaeformis Steyerm. 

Psychotria apoda Steyerm. 

Psychotria appuniana Steyerm. 

Psychotria astrellantha Wernham 

Psychotria aubletiana Steyerm. 

Psychotria ayangannensis Steyerm. 

Psychotria barnebyana Steyerm. 

Psychotria berryi Wingf. 

Psychotria berteroana DC. 

Psychotria blakei Standl. & Steyerm. 

Psychotria bolivarensis (Standl. & Steyerm.) Steyerm. 

Psychotria boliviana Standl. 

Psychotria borjensis Kunth 

Psychotria bostrychothyrsus Sandwith 


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Psychotria boyanii Steyerm. 

Psychotria bracteocardia (DC.) Müll. Arg. 

Psychotria brazaoi Steyerm. [expected in AM; Serra da Neblina] 

Psychotria bremekampiana Steyerm. 

Psychotria brevipes DC. 

Psychotria callithrix (Miq.) Steyerm. 

Psychotria callithrix var. callithrix 

Psychotria callithrix var. tontaneoides (Britton & Standl.) Steyerm. 

Psychotria campyloneura Müll. Arg. 

Psychotria campylopoda Standl. 

Psychotria capitata Ruiz & Pav. 

Psychotria capitata subsp. amplifolia (Raeusch.) Steyerm. 

Psychotria capitata subsp. capitata 

Psychotria capitata subsp. inundata (Benth.) Steyerm. 

Psychotria carapichea (Poir.) Delprete 

Psychotria cardiomorpha C.M. Taylor & A. Pool 

Psychotria carrenoi Steyerm. 

Psychotria carthagenensis Jacq. 

Psychotria casiquiaria Müll. Arg. 

Psychotria celiae Steyerm. [expected in AM; Serra da Neblina] 

Psychotria ceratantha Standl. 

Psychotria cerronis Steyerm. 

Psychotria colorata (Willd. ex Roem. & Schult.) Müll. Arg. 

Psychotria concinna Oliv. [Mt. Roraima] 

Psychotria cornigera Benth. 

Psychotria coussareoides Standl. 

Psychotria crocochlamys Sandwith 

Psychotria ctenophora Steyerm. 

Psychotria cupularis (Müll. Arg.) Standl. 

Psychotria cypellantha Steyerm. 

Psychotria deflexa DC. 

Psychotria deinocalyx Sandwith 

Psychotria duidana Standl. 

Psychotria duricoria Standl. & Steyerm. 

Psychotria duricoria var. duricoria 

Psychotria duricoria var. longiloba Steyerm. 

Psychotria durifolia Standl. 

Psychotria egensis Müll. Arg. 

Psychotria ernestii K. Krause 

Psychotria everardii Wernham 

Psychotria everardii subsp. everardii 

Psychotria fanshawei (Standl.) Steyerm. 

Psychotria farameoides Bremek. 

Psychotria franquevilleana Müll. Arg. 

Psychotria galbaoensis Steyerm. 

Psychotria glandulicalyx Steyerm. 

Psychotria gracilenta Müll. Arg. 

Psychotria granvillei Steyerm. 

Psychotria guanchezii Steyerm. 

Psychotria hemicephaelis Wernham 

Psychotria heteroneura Steyerm. [expected in AM; Serra da Neblina] 

Psychotria hoffmannseggiana (Willd. ex Roem. & Schult.) Müll. Arg. 

Psychotria horizontalis Sw. 

Psychotria humboldtiana (Cham.) Müll. Arg. 

Psychotria hyptoides Benth. 

Psychotria imthurniana Oliv. 

Psychotria iodotricha Müll. Arg. 

Psychotria irwinii Steyerm. 

Psychotria jauaensis Steyerm. 

Psychotria kaieteurensis Sandwith 

Psychotria kappleri (Miq.) Müll. Arg. ex Benoist 

Psychotria lateralis Steyerm. FG 

Psychotria leiantha Steyerm. [Sierra de la Neblina] 

Psychotria ligularis (Rudge) Steyerm. 

GU 


GU? SU FG 

FG 

GU SU FG 

SU FG 

GU FG 


AM BO GU 


GU SU FG 

GU 


FG 

AM GU SU FG 

BO 


AM BO 

AM BO GU 

BO 


AM BO GU 


AM BO GU FG 

BO GU 

SU FG 


FG 


BO GU SU FG 

AM 

AM 

AM 

AM 

AM 

AM GU? 

BO GU SU FG 

AM BO GU 

GU 

BO GU 

SU 

AM 

GU? FG 

AM BO GU 


FG 

AM 

BO GU 

GU 


AM BO GU 

AM BO 

GU 

BO GU 



AM BO 

GU 

GU SU FG 

FG 

AM 

GU SU FG

488 


Psychotria ligularis var. carapichea (Poir.) Steyerm. 

Psychotria ligularis var. ligularis 

Psychotria longicuspis Müll. Arg. 

Psychotria lourteigiana Steyerm. 

Psychotria lupulina Benth. 

Psychotria lupulina subsp. lupulina 

Psychotria lupulina subsp. rhodoleuca (Müll. Arg.) Steyerm. 

Psychotria maguireorum Steyerm. 

Psychotria mapourioides DC. 

Psychotria mapourioides var. mapourioides 

Psychotria maturacensis Steyerm. [Sierra de la Neblina] 

Psychotria mazaruniensis Standl. 

Psychotria medusula Müll. Arg. 

Psychotria microbotrys Ruiz ex Standl. 

Psychotria microdon (DC.) Urb. 

Psychotria morindoides (A. Rich.) Lemée 

Psychotria moroidea Steyerm. 

Psychotria muscosa (Jacq.) Steyerm. 

Psychotria muscosa subsp. breviloba Steyerm. 

Psychotria nana K. Krause 

Psychotria nematostachya Steyerm. [Sierra de la Neblina] 

Psychotria nivea (Sandwith) Steyerm. 

Psychotria oblita Wernham [Mt. Roraima] 

Psychotria oblonga (DC.) Steyerm. 

Psychotria occidentalis Steyerm. 

Psychotria officinalis (Aubl.) Raeusch. ex Sandwith 

Psychotria officinalis subsp. officinalis 

Psychotria officinalis subsp. wilhelminensis Steyerm. 

Psychotria oiapoquensis Steyerm. 

Psychotria pacimonica Müll. Arg. 

Psychotria pakaraimensis Steyerm. 

Psychotria pallidinervia Steyerm. [Sierra de la Neblina] 

Psychotria paniculata (Aubl.) Raeusch. 

Psychotria parimensis Steyerm. 

Psychotria parvibractea Steyerm. 

Psychotria paupertina Standl. & Steyerm. 

Psychotria pectinata Steyerm. [Sierra de la Neblina] 

Psychotria perferruginea Steyerm. 

Psychotria phaneroloma Standl. & Steyerm. 

Psychotria phaneroloma var. phaneroloma 

Psychotria phelpsiana Steyerm. 

Psychotria pilosa Ruiz & Pav. 

Psychotria platypoda DC. 

Psychotria plocamipes Wernham 

Psychotria podocephala (Müll. Arg.) Standl. 

Psychotria poeppigiana Müll. Arg. 

Psychotria poeppigiana subsp. barcellana (Müll. Arg.) Steyerm. 

Psychotria poeppigiana subsp. poeppigiana 

Psychotria polycephala Benth. 

Psychotria potaroensis (Sandwith) Steyerm. 

Psychotria prancei Steyerm. 

Psychotria prunifolia (Kunth) Steyerm. 

Psychotria pseudinundata Wernham 

Psychotria psittacina Steyerm. 

Psychotria puberulenta Steyerm. 

Psychotria pullei Bremek. 

Psychotria pungens Steyerm. 

Psychotria racemosa Rich. 

Psychotria remota Benth. 

Psychotria rosea (Benth.) Müll. Arg. 

Psychotria rosea var. rosea 

Psychotria schlechtendaliana Müll. Arg. 

Psychotria schomburgkii Benth. 

Psychotria sipapoensis Steyerm. 

SU FG 

GU SU FG 

AM GU 

AM BO 



AM BO 

AM GU 

BO GU SU FG 

BO GU SU FG 

AM GU 

GU FG 

AM GU FG 


BO GU 

FG 

SU FG 

GU SU FG 

GU SU FG 

AM 

AM 

GU 

BO GU 


DA 


GU SU FG 

SU 

FG 

AM GU FG 

GU 

AM 


AM 

BO GU 

AM BO 

AM 

FG 

AM BO GU 

GU 

BO 

AM 


GU 

AM BO GU? FG 




AM BO DA GU 

GU 

SU FG 

AM 

SU FG 

GU 

GU 

SU FG 

FG 


AM GU SU FG 

AM GU FG 

GU FG 

FG? 

AM GU 

AM


Psychotria spadicea (Pittier) Standl. & Steyerm. 

Psychotria spectabilis Steyerm. 

Psychotria speluncae Standl. & Steyerm. 

Psychotria speluncae subsp. speluncae 

Psychotria sphaerocephala Müll. Arg. 

Psychotria spiciflora Standl. 

Psychotria squamelligera Steyerm. 

Psychotria stipulosa Müll. Arg. 

Psychotria subundulata Benth. 

Psychotria surinamensis Bremek. 

Psychotria tapirapecoana Steyerm. [expected] 

Psychotria tatei Standl. 

Psychotria tepuiensis (Steyerm.) Steyerm. 

Psychotria tetramera Steyerm. 

Psychotria tillettii Steyerm. 

Psychotria transiens Wernham 

Psychotria tricholoba Müll. Arg. 

Psychotria trichophoroides Müll. Arg. 

Psychotria trichotoma M. Martens & Galeotti 

Psychotria turbinella Müll. Arg. 

Psychotria turbinella var. sororiella (Müll. Arg.) Steyerm. 

Psychotria ulviformis Steyerm. 

Psychotria urceolata Steyerm. 

Psychotria urniformis Steyerm. 

Psychotria vareschii Steyerm. 

Psychotria variegata Steyerm. 

Psychotria vasivensis (Müll. Arg.) Standl. 

Psychotria vellosiana Benth. 

Psychotria venezuelensis Steyerm. [Sierra de la Neblina] 

Psychotria ventuariana Standl. & Steyerm. 

Psychotria venulosa Müll. Arg. 

Psychotria vichadensis Standl. 

Psychotria viridibractea Steyerm. 

Psychotria viridis Ruiz & Pav. 

Psychotria wessels-boeri Steyerm. 

Psychotria wilhelminensis Steyerm. 

Psychotria wonotobensis (Bremek.) Steyerm. 

Psychotria wurdackii Steyerm. 

Psychotria yapacanensis Steyerm. 

Psychotria yavitensis Steyerm. 

Randia aculeata L. 

Randia aculeata var. dasyphylla Steyerm. 

Randia amazonasensis Steyerm. 

Randia armata (Sw.) DC. 

Randia asperifolia (Sandwith) Sandwith 

Randia brevipes Steyerm. 

Randia calycina Cham. 

Randia dioica H. Karst. 

Randia hebecarpa Benth. 

Randia pubiflora Steyerm. 

Randia venezuelensis Steyerm. 

Remijia amazonica K. Schum. 

Remijia amazonica var. amazonica 

Remijia aracamuniensis (Steyerm.) C.M. Taylor 

Remijia argentea Steyerm. 

Remijia delascioi Steyerm. 

Remijia densiflora Benth. 

Remijia densiflora subsp. densiflora 

Remijia densiflora var. densiflora 

Remijia densiflora var. minima Steyerm. 

Remijia densiflora subsp. stenopetala (Standl. & Steyerm.) Steyerm. 

Remijia firmula (Mart.) Wedd. 

Remijia globosa (Steyerm.) C.M. Taylor [Sierra de la Neblina] 

Remijia hispida Spruce ex K. Schum. 

AM BO 

AM BO GU 

GU 

AM SU 

AM 

FG 

FG 

AM 


SU FG 

AM 

AM BO GU 

GU 

GU 

AM BO GU 

GU? 

SU FG 

AM 

AM 

AM 


FG 

GU 

AM 


AM 

AM BO GU 

AM 

AM 

AM BO GU FG 

AM BO 

FG 

AM 

GU? SU 

SU 

SU 

AM 

AM 

AM 

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BO 

AM 


GU SU 

BO GU? 

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BO 

BO GU FG 

FG 

AM BO 

AM 

AM 

AM 

AM 

AM 

AM BO GU 

GU 

AM BO GU 

BO 

AM BO 

AM 

AM 

AM

490 


Remijia longifolia Benth. ex Standl. 

Remijia maguirei Steyerm. [Sierra de la Neblina] 

Remijia marahuacensis Steyerm. 

Remijia morilloi Steyerm. 

Remijia pacimonica Standl. 

Remijia pedunculata (H. Karst.) Flueck. 

Remijia pilosinervula Steyerm. 

Remijia reducta Steyerm. 

Remijia roraimae (Benth.) K. Schum. 

Remijia roraimae var. adpressa Steyerm. 

Remijia roraimae var. auyantepuiensis Steyerm. 

Remijia roraimae var. guianensis Steyerm. 

Remijia roraimae var. roraimae 

Remijia sessilis Steyerm. [Sierra de la Neblina] 

Remijia sipapoensis Steyerm. 

Remijia steyermarkii Standl. 

Remijia ulei K. Krause 

Remijia uniflora C.M. Taylor 

Remijia wurdackii Steyerm. 

Retiniphyllum chloranthum Ducke 

Retiniphyllum concolor (Spruce ex Benth.) Müll. Arg., emend. Cortés 

Retiniphyllum discolor (Spruce ex Benth.) Müll. Arg. 

Retiniphyllum guianense Steyerm. 

Retiniphyllum laxiflorum (Benth.) N.E. Br. 

Retiniphyllum laxiflorum var. laxiflorum 

Retiniphyllum maguirei Standl. 

Retiniphyllum pauciflorum Kunth ex K. Krause 

Retiniphyllum pilosum (Spruce ex Benth.) Müll. Arg. 

Retiniphyllum scabrum Benth. 

Retiniphyllum schomburgkii (Benth.) Müll. Arg. 

Retiniphyllum schomburgkii subsp. schomburgkii 

Retiniphyllum secundiflorum Bonpl. 

Retiniphyllum tepuiense Steyerm. 

Retiniphyllum truncatum Müll. Arg. 

Richardia brasiliensis Gomes 

Richardia scabra L. 

Ronabea latifolia Aubl. 

Rondeletia orinocensis Steyerm. 

Rosenbergiodendron densiflorum (K. Schum.) Fagerl. 

Rosenbergiodendron formosum (Jacq.) Fagerl. 

Rosenbergiodendron formosum var. nitidum (K. Schum.) C. Gust. 

Rosenbergiodendron longiflorum Fagerl. 

Rudgea ayangannensis Steyerm. 

Rudgea bremekampiana Steyerm. 

Rudgea cardonae Steyerm. 

Rudgea corniculata Benth. 

Rudgea cornifolia (Kunth ex Roem. & Schult.) Standl. 

Rudgea cornigera Bremek. 

Rudgea crassiloba (Benth.) B.L. Rob. 

Rudgea enervia Steyerm. 

Rudgea graciliflora Standl. 

Rudgea guyanensis (A. Rich.) Sandwith 

Rudgea hostmanniana Benth. 

Rudgea hostmanniana subsp. hostmanniana 

Rudgea hostmanniana subsp. maypurensis (Standl.) Zappi 

Rudgea klugii Standl. 

Rudgea lanceifolia Salisb. 

Rudgea merumensis Steyerm. 

Rudgea oldemanii Steyerm. 

Rudgea phaneroneura Steyerm. 

Rudgea sandemanii Sandwith 

Rudgea sclerocalyx (Müll. Arg.) Zappi 

Rudgea stipulacea (DC.) Steyerm. 

Rudgea tanaosepala Sandwith 

AM 

AM 

AM GU 

AM 

AM 

AM 

BO GU 

AM 

AM BO GU 

AM BO 

BO 

BO GU 

AM BO GU 

AM 

AM 

AM 

AM 

AM 

AM 

AM BO GU 

AM BO GU 

AM 

GU 

AM BO GU SU 

AM BO GU SU 

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AM GU 

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AM BO GU 

AM BO GU SU 

GU SU 

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AM 

AM GU 

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AM BO 


AM BO DA GU? 

AM BO DA 

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AM 

FG 


GU SU FG 


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AM 

AM GU SU FG 

AM GU SU FG 

GU 

FG 

BO 

FG 

AM 


GU FG

Rudgea woronowii Standl. 

Rudgea wurdackii Steyerm. 

Sabicea amazonensis Wernham 

Sabicea aspera Aubl. 

Sabicea bariensis Steyerm. 

Sabicea brachycalyx Steyerm. 

Sabicea cinerea Aubl. [expected in GU] 

Sabicea glabrescens (K. Schum.) Benth. 

Sabicea grandifolia Steyerm. 

Sabicea morillorum Steyerm. 

Sabicea oblongifolia (Miq.) Steyerm. 

Sabicea panamensis Wernham 

Sabicea paraensis (K. Schum.) Wernham 

Sabicea romboutsii Bremek. 

Sabicea surinamensis Bremek. 

Sabicea tillettii Steyerm. 

Sabicea velutina Benth. 

Sabicea velutina subsp. chimantensis Steyerm. 

Sabicea velutina subsp. duidensis Steyerm. 

Sabicea velutina subsp. velutina 

Sabicea venezuelensis Steyerm. 

Sabicea villosa Willd. ex Roem. & Schult. 

Sabicea villosa var. adpressa (Wernham) Standl. 

Sabicea villosa var. villosa 

Schradera bipedunculata Steyerm. 

Schradera bipendunculata Steyerm. 

Schradera brevipes Steyerm. 

Schradera hilliifolia Steyerm. [Sierra de la Neblina] 

Schradera maguirei Steyerm. 

Schradera marahuacensis Steyerm. 

Schradera nilssonii Steyerm. 

Schradera polycephala DC. 

Schradera surinamensis Standl. 

Schradera ternata Steyerm. 

Schradera yutajensis Steyerm. 

Semaphyllanthe intonsa (Steyerm.) L. Andersson 

Semaphyllanthe merumensis (Steyerm.) L. Andersson 

Semaphyllanthe spectabilis (Steyerm.) L. Andersson 

Simira ignicola Steyerm. 

Simira pisoniiformis (Baill.) Steyerm. 

Simira rubescens (Benth.) Bremek. ex Steyerm. 

Simira tinctoria Aubl. 

Sipanea ayangannensis Steyerm. 

Sipanea biflora (L. f.) Cham. & Schltdl. 

Sipanea carrenoi Steyerm. 

Sipanea cowanii Steyerm. 

Sipanea galioides Wernham 

Sipanea glaberrima (Bremek.) Steyerm. 

Sipanea gleasonii Steyerm. 

Sipanea glomerata Kunth 

Sipanea hispida Benth. ex Wernham 

Sipanea hispida var. hispida 

Sipanea micrantha Sandwith 

Sipanea ovalifolia Bremek. 

Sipanea ovalifolia var. ovalifolia 

Sipanea ovalifolia var. villosissima Steyerm. 

Sipanea pratensis Aubl. 

Sipanea pratensis var. dichotoma (Kunth) Steyerm. 

Sipanea pratensis var. pratensis 

Sipanea setacea Steyerm. 

Sipanea stahelii Bremek. 

Sipanea veris S. Moore 

Sipanea wilson-brownei R.S. Cowan 

Sipaneopsis cururuensis J.H. Kirkbr. 


AM BO 

AM 

AM SU 

GU SU FG 

AM 

AM BO GU 

SU FG 


AM 

AM 


FG? 

FG 

SU 

GU SU 

AM 


BO 

AM 


AM BO 


AM BO DA FG? 


BO 

BO 

AM 

AM 

AM 

AM 

BO GU 

BO GU SU FG 

GU SU 

GU 

AM 

GU 

GU 

GU 

BO 

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AM BO FG? 

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BO GU 

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AM BO GU FG? 

AM BO 

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GU SU FG 

BO 

SU FG? 

AM BO GU 

BO GU FG 

GU

492 


Sipaneopsis foldatsii Steyerm. 

Sipaneopsis huberi Steyerm. 

Sipaneopsis maguirei Steyerm. 

Sipaneopsis morichensis Steyerm. 

Sipaneopsis pacimoniensis Steyerm. 

Sipaneopsis rupicola (Spruce ex K. Schum.) Steyerm. 

Spermacoce glabra Michx. 

Spermacoce hexangularis Aubl. [name of uncertain application] 

Spermacoce radicans Aubl. [=Diodia sp.?] 

Sphinctanthus maculatus Spruce ex K. Schum. 

Sphinctanthus striiflorus (DC.) Hook. f. 

Stachyarrhena duckei Standl. 

Stachyarrhena penduliflora K. Schum. 

Stachyarrhena reticulata Steyerm. 

Stachyarrhena spicata Hook. f. 

Tobagoa maleolens Urb. 

Tocoyena brevifolia Steyerm. 

Tocoyena formosa (Cham. & Schltdl.) K. Schum. 

Tocoyena guianensis K. Schum. 

Tocoyena longiflora Aubl. 

Tocoyena neglecta N.E. Br. 

Tocoyena orinocensis Standl. & Steyerm. 

Tocoyena pendulina Spruce ex Standl. 

Tocoyena surinamensis Bremek. 

Uncaria guianensis (Aubl.) J.F. Gmel. 

Uncaria tomentosa (Willd. ex Roem. & Schult.) DC. 

Warszewiczia coccinea (Vahl) Klotzsch 

Warszewiczia elata Ducke 

Warszewiczia schwackei K. Schum. 

Wittmackanthus stanleyanus (R.H. Schomb.) Kuntze 


Alibertia acuminata (Benth.) Sandwith =Alibertia edulis var. edulis 

Alibertia acuminata var. acuminata =Alibertia edulis var. edulis 

Alibertia acuminata var. obtusiuscula Steyerm. =Alibertia edulis var. obtusiuscula 

Alibertia benensis Standl. =Cordiera triflora 

Alibertia davidsei Steyerm. =Alibertia edulis var. obtusiuscula 

Alibertia granulosa Rusby =Alibertia latifolia 

Alibertia hexagyna H. Karst. =Alibertia edulis var. edulis 

Alibertia lanceolata (Cham.) Mennega [nomen nudum] 

Alibertia latifolia var. pargueniana Steyerm. =Alibertia latifolia 

Alibertia latifolia var. parvifolia K. Schum. =Alibertia latifolia 

Alibertia longistipulata L. Riley =Alibertia edulis var. edulis 

Alibertia myrciifolia Spruce ex K. Schum. =Cordiera myrciifolia 

Alibertia myrciifolia var. myrciifolia =Cordiera myrciifolia 

Alibertia myrciifolia var. tepuiensis Steyerm. =Cordiera myrciifolia 

Alibertia nitidula (Bremek.) L. Andersson =Alibertia edulis var. edulis 

Alibertia panamensis L. Riley =Alibertia edulis var. edulis 

Alibertia steinbachii Standl. =Cordiera triflora 

Alibertia stenantha Standl. =Alibertia bertierifolia 

Alibertia surinamensis (Bremek.) Steyerm. =Ibetralia surinamensis 

Alibertia tenuifolia K. Krause =Cordiera triflora 

Alibertia tobagensis Sprague & R.O. Williams =Alibertia edulis var. edulis 

Alibertia triflora (A. Rich.) K. Schum. =Cordiera triflora 

Alibertia triloba Steyerm. =Cordiera myrciifolia 

Alibertia trinitatis Sprague & R.O. Williams =Alibertia edulis var. edulis 

Alibertia tutumilla Rusby =Alibertia edulis var. edulis 

Alibertia uniflora Standl. =Cordiera myrciifolia 

Alibertia utilis A. Rich. =Alibertia edulis 

Alseis leiantha S.F. Blake =Alseis labatioides 

Alseis trichocarpa Standl. & Steyerm. =Alseis labatioides 

Amaioua fusifera Spruce ex Benth. & Hook. f. [nomen nudum] =Duroia fusifera 

Amaioua genipoides Spruce ex K. Schum. [nomen nudum] =Duroia genipoides 

Amaioua peruviana Desf. =Amaioua corymbosa 

AM 

AM 

AM 

AM 

AM 

AM 

GU 

FG 

FG 

FG 

AM BO FG? 

AM 

AM BO GU? 

AM 

AM GU 

BO 

AM 

GU? 

AM GU SU FG 

FG 

BO GU 

AM BO 

AM 

SU 



AM GU SU FG 

AM 

AM 

GU


Amaioua saccifera Mart. ex Roem. & Schult. =Duroia saccifera 

Amaioua ursina Standl. =Duroia eriopila var. eriopila 

Amaioua velutina Spruce ex Benth. & Hook. [nomen nudum] =Duroia velutina 

Anisomeris caurensis Standl. =Chomelia caurensis 

Anisomeris malaneoides (Müll. Arg.) Standl. =Chomelia malaneoides 

Anisomeris obtusa (Cham. & Schltdl.) K. Schum. =Chomelia obtusa 

Anisomeris polyantha (S.F. Blake) Rusby =Chomelia polyantha 

Anisomeris tenuiflora (Benth.) Pulle =Chomelia tenuiflora 

Anisomeris volubilis Standl. =Chomelia volubilis 

Antirhea surinamensis Bremek. =Guettarda acreana 

Aphanocarpus steyermarkii f. elongatus Steyerm. =Aphanocarpus steyermarkii 

Aphanocarpus steyermarkii f. glabrior Steyerm. =Aphanocarpus steyermarkii 

Appunia angulata (Benth.) Baill. =Morinda peduncularis 

Appunia aurantiaca (K. Krause) Sandwith =Morinda aurantiaca 

Appunia brachycalyx (Bremek.) Steyerm. =Morinda brachycalyx 

Appunia calycina (Benth.) Sandwith =Morinda calycina 

Appunia debilis Sandwith =Morinda debilis 

Appunia longiloba Steyerm. =Morinda longiloba 

Appunia sparsiflora Steyerm. =Morinda sparsiflora 

Appunia surinamensis (Bremek.) Steyerm. =Morinda surinamensis 

Appunia tenuiflora (Benth.) Jacks. & Hook. f. =Morinda tenuiflora 

Appunia tenuiflora var. leiophylla Steyerm. =Morinda tenuiflora 

Appunia venezuelensis Steyerm. =Morinda venezuelensis 

Basanacantha asperifolia Sandwith =Randia asperifolia 

Basanacantha dioica (H. Karst.) Hook. f. =Randia dioica 

Basanacantha hebecarpa (Benth.) Hook. f. =Randia hebecarpa 

Basanacantha spinosa (Jacq.) K. Schum. =Randia armata 

Basanacantha spinosa f. macrophylla K. Schum. =Randia amazonasensis 

Bellardia repens Willd. =Coccocypselum guianense 

Bellardia tontanea (Kunth) Schult. & Schult. f. =Coccocypselum guianense 

Bellynkxia angulata (Benth.) Müll. Arg. =Morinda peduncularis 

Bellynkxia brachycalyx Bremek. =Morinda brachycalyx 

Bellynkxia surinamensis Bremek. =Morinda surinamensis 

Bertiera guianensis subsp. guianensis =Bertiera guianensis 

Bertiera palustris A. Rich. =Sipanea wilson-brownei 

Billardiera paniculata Vahl =Coussarea paniculata 

Borojoa lanceolata (Cham.) Cuatrec. =Alibertia edulis var. edulis 

Borreria assurgens (Ruiz & Pav.) Griseb. =Borreria remota 

Borreria capitata f. ferruginea (A. St.-Hil.) Steyerm. =Borreria capitata var. capitata 

Borreria capitata f. glabra Steyerm. =Borreria capitata var. capitata 

Borreria exilis L.O. Williams =Borreria repens 

Borreria ferruginea (A. St.-Hil.) DC. =Borreria capitata var. capitata 

Borreria fockeana Miq. =Borreria latifolia 

Borreria hispida var. glabrescens K. Schum. =Borreria hispida 

Borreria hyssopifolia (Willd. ex Roem. & Schult.) Bacigalupo & E.L. Cabral =Diodia hyssopifolia 

Borreria kappleriana Miq. =Borreria capitata var. capitata 

Borreria laevis (Lam.) Griseb. [not in flora area] =Spermacoce tenuoir L. 

Borreria laevis auct., non (Lam.) Griseb. =Borreria remota 

Borreria latifolia var. fockeana (Miq.) Bremek. =Borreria latifolia 

Borreria latifolia var. minor K. Schum. =Borreria latifolia 

Borreria latifolia f. minor (K. Schum.) Steyerm. =Borreria latifolia 

Borreria multiflora (DC.) Bacigalupo & E.L. Cabral =Diodia multiflora 

Borreria ocimoides auct., non (Burm. f.) DC. =Borreria prostrata 

Borreria ocymifolia (Willd. ex Roem. & Schult.) Bacigalupo & E.L. Cabral =Diodia ocymifolia 

Borreria oligodonta Steyerm. =Borreria verticillata 

Borreria spicata (Miq.) Bacigalupo & E.L. Cabral =Diodia spicata 

Borreria splitgerberi Bremek. =Borreria latifolia 

Borreria suaveolens G. Mey. =Borreria capitata var. suaveolens 

Borreria tenella (Kunth) Cham. & Schltdl. =Borreria capitata var. tenella 

Borreria tenella var. platyphylla K. Schum. =Borreria capitata var. capitata 

Borreria umbellata Spreng. =Emmeorhiza umbellata 

Brignolia pubigera Benth. =Isertia parviflora 

Bruinsmania isertioides Miq. =Isertia parviflora 

Buchia plantaginea Kunth =Perama plantaginea

494 


Buena triflora Benth. =Cosmibuena grandiflora 

Callicocca alba (Aubl.) J.F. Gmel. =Psychotria ulviformis 

Callicocca purpurea (Aubl.) J.F. Gmel. =Psychotria variegata 

Callicocca tomentosa (Aubl.) J.F. Gmel. =Psychotria poeppigiana 

Calycophyllum intonsum Steyerm. =Semaphyllanthe intonsa 

Calycophyllum merumense Steyerm. =Semaphyllanthe merumensis 

Calycophyllum spectabile Steyerm. =Semaphyllanthe spectabilis 

Capirona leiophloea Benoist =Capirona decorticans 

Capirona surinamensis Bremek. =Capirona decorticans 

Capirona wurdackii Steyerm. =Capirona decorticans 

Carapichea guianensis Aubl. =Psychotria ligularis var. carapichea 

Carapichea kappleri Miq. =Psychotria kappleri 

Cascarilla roraimae (Benth.) Wedd. =Remijia roraimae 

Cassupa verrucosa Bonpl. =Isertia verrucosa 

Cephaelis aetantha Sandwith =Psychotria aetantha 

Cephaelis alba (Aubl.) Willd. =Psychotria ulviformis 

Cephaelis altsonii Sandwith =Psychotria altsonii 

Cephaelis amoena Bremek. =Psychotria colorata 

Cephaelis aneurophylla Standl. =Notopleura aneurophylla 

Cephaelis axillaris Sw. =Psychotria aubletiana 

Cephaelis barcellana (Müll. Arg.) Standl. =Psychotria poeppigiana subsp. barcellana 

Cephaelis blepharophylla Standl. =Psychotria medusula 

Cephaelis bolivarensis Standl. & Steyerm. =Psychotria bolivarensis 

Cephaelis bracteocardia DC. =Psychotria bracteocardia 

Cephaelis cacuminis Steyerm. =Psychotria aubletiana 

Cephaelis callithrix Miq. =Psychotria callithrix 

Cephaelis colorata Willd. ex Roem. & Schult. =Psychotria colorata 

Cephaelis dichotoma Willd. ex Roem. & Schult. [nom. illeg.] =Psychotria hoffmannseggiana 

Cephaelis dichotoma Rudge =Psychotria platypoda 

Cephaelis duckei Standl. =Psychotria spectabilis 

Cephaelis ernestii K. Krause =Psychotria transiens 

Cephaelis fanshawei Standl. =Psychotria fanshawei 

Cephaelis glabrescens (Müll. Arg.) Standl. [not in flora area] =Psychotria glabrescens Muell. Arg. 

Cephaelis guianensis (Aubl.) Standl. =Psychotria ligularis var. carapichea 

Cephaelis hemicephaelis (Wernham) Steyerm. =Psychotria hemicephaelis 

Cephaelis hirsuta M. Martens & Galeotti =Psychotria poeppigiana 

Cephaelis hirta Miq. =Psychotria bremekampiana 

Cephaelis hoffmannseggiana Willd. ex Roem. & Schult. =Psychotria hoffmannseggiana 

Cephaelis humboldtiana Cham. =Psychotria humboldtiana 

Cephaelis humboldtiana var. caudata Müll. Arg. =Psychotria humboldtiana 

Cephaelis humboldtiana var. ornata (Müll. Arg.) Standl. =Psychotria humboldtiana 

Cephaelis jenmanii Wernham =Psychotria mazaruniensis 

Cephaelis kaieteurensis Wernham =Psychotria appuniana 

Cephaelis kappleri (Miq.) Standl. =Psychotria kappleri 

Cephaelis ligularis (Rudge) A. Rich. =Psychotria ligularis 

Cephaelis microcephala Willd. ex Roem. & Schult. =Psychotria prunifolia 

Cephaelis microcephala Miq. [nom. illeg.] =Psychotria hoffmannseggiana 

Cephaelis muscosa (Jacq.) Sw. =Psychotria muscosa 

Cephaelis nivea Sandwith =Psychotria nivea 

Cephaelis oblonga DC. =Psychotria oblonga 

Cephaelis paraensis Standl. =Psychotria ulviformis 

Cephaelis patrisii D. Dietr. =Psychotria sp. 

Cephaelis potaroensis Sandwith =Psychotria potaroensis 

Cephaelis prunifolia Kunth =Psychotria prunifolia 

Cephaelis pubescens Hoffmanns. ex Roem. & Schult. =Psychotria bracteocardia 

Cephaelis purpurea (Aubl.) Willd. =Psychotria variegata 

Cephaelis rosea Benth. =Psychotria rosea 

Cephaelis rubra Willd. ex Roem. & Schult. =Psychotria hoffmannseggiana 

Cephaelis sandwithiana Steyerm. =Notopleura sandwithiana 

Cephaelis sororiella (Müll. Arg.) Standl. =Psychotria turbinella var. sororiella 

Cephaelis sphaerocephala (Müll. Arg.) B.L. Rob. =Psychotria sphaerocephala 

Cephaelis stipulosa (Müll. Arg.) Standl. =Psychotria stipulosa 

Cephaelis surinamensis Standl. =Psychotria variegata 

Cephaelis sylvatica Bremek. =Psychotria ctenophora


Cephaelis tatei Standl. =Psychotria crocochlamys 

Cephaelis tepuiensis Steyerm. =Psychotria tepuiensis 

Cephaelis tomentosa (Aubl.) Vahl =Psychotria poeppigiana 

Cephaelis tontaneoides (Britton & Standl.) R.O. Williams & Cheesman =Psychotria callithrix var. tontaneoides 

Cephaelis trichophoroides (Müll. Arg.) Standl. =Psychotria trichophoroides 

Cephaelis violacea (Aubl.) Sw. =Psychotria apoda 

Cephalodendron aracamuniensis Steyerm. =Remijia aracamuniensis 

Cephalodendron globosum Steyerm. =Remijia globosa 

Chalepophyllum coriaceum Gleason =Maguireothamnus speciosus subsp. speciosus 

Chalepophyllum guianense var. cuneatum Steyerm. =Chalepophyllum guianense 

Chalepophyllum latifolium Standl. =Maguireothamnus tatei 

Chalepophyllum longilobum Steyerm. =Chalepophyllum guianense 

Chalepophyllum pungens Standl. & Steyerm. (Melastomataceae) =Acanthella sprucei 

Chalepophyllum speciosum N.E. Br. =Maguireothamnus speciosus 

Chalepophyllum tatei Standl. =Maguireothamnus tatei 

Chimarrhis bathysoides Steyerm. =Bathysa bathysoides 

Chimarrhis longistipulata Bremek. =Chimarrhis microcarpa 

Chiococca alba var. micrantha (J.R. Johnst.) Steyerm. =Chiococca alba 

Chiococca alba subsp. parvifolia (Wullschl. ex Griseb.) Steyerm. =Chiococca alba 

Chiococca alba f. pilosa Steyerm. =Chiococca alba 

Chiococca brachiata Ruiz & Pav. =Chiococca alba 

Chiococca cordata R.S. Cowan =Chiococca nitida var. amazonica 

Chiococca erubescens Wernham =Chiococca nitida var. nitida 

Chiococca micrantha J.R. Johnst. =Chiococca alba 

Chiococca parvifolia Wullschl. ex Griseb. =Chiococca alba 

Chiococca racemosa L. =Chiococca alba 

Chomelia angustifolia Benth. =Chomelia obtusa 

Chomelia barbellata Standl. =Chomelia tenuiflora 

Chomelia glabriuscula Steyerm. =Chomelia malaneoides 

Chondrococcus laevis Steyerm. =Coccochondra laevis 

Chytropsia astrellantha (Wernham) Bremek. =Psychotria astrellantha 

Chytropsia guianensis Bremek. =Psychotria moroidea 

Cinchona amazonica Standl. =Cinchonopsis amazonica 

Cinchona brasiliensis Hoffmanns. ex Humb. =Machaonia brasiliensis 

Cinchona firmula Mart. =Remijia firmula 

Cinchona grandiflora Ruiz & Pav. =Cosmibuena grandiflora 

Cinchona lambertiana A. Braun ex Mart. =Ladenbergia lambertiana 

Cinchona magnifolia Ruiz & Pav. =Ladenbergia oblongifolia 

Cinchona oblongifolia Mutis =Ladenbergia oblongifolia 

Cinchona pedunculata H. Karst. =Remijia pedunculata 

Cinchona roraimae Benth. =Remijia roraimae 

Coccocypselum aureum var. capitatum (Benth.) Steyerm. =Coccocypselum aureum 

Coccocypselum brevipetiolatum Steyerm. =Coccocypselum hirsutum 

Coccocypselum canescens Willd. ex Schult. & Schult. f. =Coccocypselum lanceolatum 

Coccocypselum croatii Steyerm. =Coccocypselum condalia 

Coccocypselum guianense var. patens Steyerm. =Coccocypselum guianense 

Coccocypselum tontanea Kunth =Coccocypselum guianense 

Coffea crassiloba Benth. =Rudgea crassiloba 

Coffea laurifolia Kunth [nom. illeg.] =Rudgea sclerocalyx 

Coffea paniculata Aubl. =Faramea paniculata 

Coffea sessilis Vell. =Psychotria vellosiana 

Coffea stipulacea DC. =Rudgea stipulacea 

Coffea subsessilis Benth. =Ronabea latifolia 

Coffea tenuiflora Benth. =Morinda tenuiflora 

Commianthus concolor Spruce ex Benth. =Retiniphyllum concolor 

Commianthus discolor Spruce ex Benth. =Retiniphyllum discolor 

Commianthus pilosus Spruce ex Benth. =Retiniphyllum pilosum 

Commianthus schomburgkii Benth. =Retiniphyllum schomburgkii 

Condalia lanceolata Ruiz & Pav. =Coccocypselum lanceolatum 

Condalia repens Ruiz & Pav. =Coccocypselum condalia 

Conosiphon aureus Poepp. & Endl. =Sphinctanthus striiflorus 

Cordiera acuminata Benth. =Alibertia edulis var. edulis 

Cordiera bertierifolia (K. Schum.) Kuntze =Alibertia bertierifolia 

Cordiera edulis (Rich.) Kuntze =Alibertia edulis

496 


Cordiera hexagyna (H. Karst.) Kuntze =Alibertia edulis var. edulis 

Cordiera latifolia Benth. =Alibertia latifolia 

Cordiera trifolia Steud. =Cordiera triflora 

Cosmibuena triflora (Benth.) Klotzsch =Cosmibuena grandiflora 

Coupoui aquatica Aubl. =Duroia aquatica 

Coupoui martiniana (Miers) Wernham =Duroia martiniana 

Coupoui micrantha Ladbr. =Duroia micrantha 

Coussarea multiflora (A. Rich.) Lemée =Faramea multiflora 

Coussarea paniculata (Aubl.) Lemée, nom. illeg. =Faramea paniculata 

Coussarea schomburgkiana (Benth.) Benth. & Hook. f. =Coussarea violacea 

Coutarea campanilla DC. =Coutarea hexandra 

Coutarea hexandra var. campanilla (DC.) Steyerm. =Coutarea hexandra 

Coutarea hexandra var. pubescens K. Schum. =Coutarea hexandra 

Coutarea hexandra f. pubescens (Pohl) Steyerm. =Coutarea hexandra 

Coutarea lindeniana Baill. =Coutarea hexandra 

Coutarea pubescens Pohl =Coutarea hexandra 

Coutarea speciosa Aubl. =Coutarea hexandra 

Cupirana martiniana Miers =Duroia martiniana 

Declieuxia alfredii Ernst =Declieuxia fruticosa 

Declieuxia brasiliensis (Spreng.) Müll. Arg. =Declieuxia tenuiflora 

Declieuxia chiococcoides Kunth =Declieuxia fruticosa 

Declieuxia chiococcoides var. guyanensis Müll. Arg. =Declieuxia fruticosa 

Declieuxia fruticosa var. guyanensis (Müll. Arg.) Standl. =Declieuxia fruticosa 

Declieuxia fruticosa var. mexicana (DC.) Standl. =Declieuxia fruticosa 

Declieuxia mexicana DC. =Declieuxia fruticosa 

Declieuxia roraimensis Wernham =Psychotria capitata subsp. inundata 

Dendrosipanea wurdackii Steyerm. =Dendrosipanea spigelioides 

Dicrobotryum divaricatum Humb. & Bonpl. ex Roem. & Schult. =Guettarda divaricata 

Diodella rigida (Cham. & Schltdl.) Small =Diodia apiculata 

Diodella teres (Walter) Small =Diodia teres 

Diodia hyssopifolia var. articulata (Pohl ex DC.) Steyerm. =Diodia hyssopifolia 

Diodia hyssopifolia f. glabriuscula Steyerm. =Diodia hyssopifolia 

Diodia hyssopifolia var. linearis (Willd. ex Roem. & Schult.) Steyerm. =Diodia hyssopifolia 

Diodia hyssopifolia f. psiloclada Steyerm. =Diodia hyssopifolia 

Diodia ottonis K. Schum. ex Pittier =Tobagoa maleolens 

Diodia prostrata Sw. =Diodia teres 

Diodia rigida Cham. & Schltdl. =Diodia apiculata 

Diodia setigera DC. =Diodia apiculata 

Diodia surinamensis Miq. =Diodia hyssopifolia 

Diodia teres var. angustata A. Gray =Diodia teres 

Diodia teres subsp. angustata (A. Gray) Steyerm. =Diodia teres 

Diodia teres f. latifolia Steyerm. =Diodia teres 

Diodia teres f. latior Steyerm. =Diodia teres 

Diodia teres f. leiocarpa Steyerm. =Diodia teres 

Diodia teres subsp. prostrata (Sw.) Steyerm. =Diodia teres 

Duroia eriopila f. glabra Steyerm. =Duroia eriopila var. eriopila 

Duroia genipoides Spruce ex Hook. f. [nomen nudum] =Duroia genipoides 

Duroia sprucei Rusby =Duroia micrantha 

Euosmia corymbosa Benth. =Bothriospora corymbosa 

Evea guianensis Aubl. =Faramea guianensis 

Evea tomentosa (Aubl.) Standl. =Psychotria poeppigiana 

Evea tontaneoides Britton & Standl. =Psychotria callithrix var. tontaneoides 

Faramea anisocalyx var. pulchella (Müll. Arg.) Steyerm. =Faramea anisocalyx 

Faramea leptoloba Benth. & Hook. f. =Coussarea leptoloba 

Faramea longifolia Benth. =Faramea sessilifolia 

Faramea multiflora var. epedunculata Steyerm. =Faramea multiflora 

Faramea multiflora var. salicifolia (C. Presl) Steyerm. =Faramea multiflora 

Faramea occidentalis var. meridionalis Steyerm. =Faramea occidentalis 

Faramea pachydictyon Müll. Arg. =Faramea torquata 

Faramea paludicola Steyerm. =Faramea torquata 

Faramea pulchella Müll. Arg. =Faramea anisocalyx 

Faramea salicifolia C. Presl =Faramea multiflora 

Faramea schomburgkiana Benth. =Coussarea violacea 

Faramea truncata DC. =Faramea corymbosa


Faramea umbelliflora Bremek. =Faramea quinqueflora 

Galium croceum Ruiz & Pav. =Galium hypocarpium 

Gamotopea alba (Aubl.) Bremek. =Psychotria ulviformis 

Gamotopea callithrix (Miq.) Bremek. =Psychotria callithrix 

Gamotopea oblonga (DC.) Lemée =Psychotria oblonga 

Gamotopea purpurea (Aubl.) Bremek. =Psychotria variegata 

Gamotopea surinamensis (Standl.) Bremek. =Psychotria variegata 

Garapatica edulis (Rich.) H. Karst. =Alibertia edulis 

Gardenia armata Sw. =Randia armata 

Gardenia edulis (Rich.) Poir. =Alibertia edulis 

Genipa caruto Kunth =Genipa americana var. caruto 

Genipa edulis Rich. =Alibertia edulis 

Genipa ? striiflora DC. =Sphinctanthus striiflorus 

Geophila herbacea (Jacq.) K. Schum. =Geophila repens 

Geophila paraensis Huber ex Standl. [nomen nudum] =Psychotria ulviformis 

Geophila picta Rolfe =Psychotria ulviformis 

Geophila violacea (Aubl.) DC. ? =Coccocypselum sp. 

Gleasonia duidana var. oblanceolata Steyerm. =Gleasonia duidana var. duidana 

Gonzalagunia dicocca var. guianensis Steyerm. =Gonzalagunia dicocca 

Gonzalagunia dicocca subsp. venezuelensis Steyerm. =Gonzalagunia dicocca 

Guettarda coccinea Aubl. =Isertia coccinea 

Guettarda grandiflora Steud. =Guettarda aromatica 

Guettarda leiantha Steyerm. =Guettarda acreana 

Guettarda ulei K. Krause =Guettarda acreana 

Hamelia lutea Rohr ex Sm. =Hamelia axillaris 

Hamelia patens var. glabra Oerst. =Hamelia patens 

Hedyotis lancifolia Schumach. =Oldenlandia lancifolia 

Hemidiodia ocymifolia (Willd. ex Roem. & Schult.) K. Schum. =Diodia ocymifolia 

Henriquezia aturensis Standl. =Platycarpum orinocense var. orinocense 

Henriquezia nitida var. oblonga (Spruce ex Benth.) Steyerm. =Henriquezia nitida var. nitida 

Henriquezia oblonga Spruce ex Benth. =Henriquezia nitida var. nitida 

Hillia tubiflora Cham. =Hillia illustris 

Houstonia fruticosa Willd. ex Roem. & Schult. =Declieuxia fruticosa 

Isertia commutata Miq. =Isertia spiciformis 

Isertia parviflora var. hirta Steyerm. =Isertia parviflora 

Isertia pterantha Bremek. =Isertia spiciformis 

Ixora alternifolia Jacq. (Solanaceae) =Cestrum alternifolium 

Ixora angulata Benth. =Morinda peduncularis 

Ixora duidae Standl. =Rudgea stipulacea 

Ixora floribunda (Rich.) Griseb. [not in flora area] 

Ixora gleasonii Standl. =Ixora schomburgkiana 

Ixora guianensis (Aubl.) Spreng. [see Coffea guianensis] 

Ixora mazarunensis Standl. =Ixora davisii 

Ixora occidentalis L. =Faramea occidentalis 

Ixora orinocensis Spruce ex Standl. [nomen nudum] 

Ixora podocarpa Benth. =Simira pisoniiformis 

Ixora surinamensis Bremek. =Ixora graciliflora 

Ixora tenuiflora (DC.) Lemée =Patabea tenuiflora 

Ixora xantholoba Standl. =Ixora graciliflora 

Knoxia brasiliensis Spreng. =Declieuxia tenuiflora 

Ladenbergia lucens Standl. =Ladenbergia lambertiana 

Ladenbergia magnifolia (Ruiz & Pav.) Klotzsch =Ladenbergia oblongifolia 

Ladenbergia puberula Steyerm. =Ladenbergia lambertiana 

Ladenbergia roraimae (Benth.) Klotzsch =Remijia roraimae 

Ladenbergia schomburgkii Klotzsch =Ladenbergia lambertiana 

Ladenbergia venamoensis Steyerm. =Ladenbergia lambertiana 

Limnosipanea guaricensis Pittier =Limnosipanea spruceana 

Limnosipanea kuntzei Standl. =Limnosipanea spruceana 

Limnosipanea schomburgkii Hook. f. =Limnosipanea palustris 

Limnosipanea ternifolia Pittier =Limnosipanea spruceana 

Lonicera alba L. =Chiococca alba 

Macrocnemum coccineum Vahl =Warszewiczia coccinea 

Maguireothamnus jauaensis (Steyerm.) Steyerm. =Maguireothamnus speciosus subsp. jauaensis 

Maguireothamnus jauaensis var. breweri Steyerm. =Maguireothamnus speciosus subsp. jauaensis

498 


Maguireothamnus tatei var. latifolius (Standl.) Steyerm. =Maguireothamnus tatei 

Malanea angustifolia Bartl. [nomen nudum] =Malanea obovata 

Malanea macrophylla Bartl. ex Griseb. [nomen nudum] =Malanea macrophylla 

Malanea roraimensis Wernham =Malanea obovata 

Malanea sarmentosa f. tomentosa Steyerm. =Malanea sarmentosa 

Manettia coccinea var. spraguei (Wernham) Steyerm. =Manettia coccinea 

Manettia uniflora Kunth =Manettia reclinata 

Mapouria alba (Ruiz & Pav.) Müll. Arg. =Psychotria alba 

Mapouria borjensis (Kunth) Müll. Arg. =Psychotria borjensis 

Mapouria brownei (Spreng.) Lemée [not in flora area] =Psychotria brownei Spreng. 

Mapouria chionantha (DC.) Müll. Arg. =Psychotria mapourioides 

Mapouria cupularis Müll. Arg. =Psychotria cupularis 

Mapouria ficigemma (DC.) Lemée =Psychotria carthagenensis 

Mapouria fockeana (Miq.) Bremek. =Psychotria carthagenensis 

Mapouria guianensis Aubl. =Psychotria mapourioides 

Mapouria herbacea var. orbicularis Müll. Arg. =Geophila orbicularis 

Mapouria mahuriensis Bremek. =Psychotria cupularis 

Mapouria microdon (DC.) Bremek. =Psychotria microdon 

Mapouria opaca Bremek. =Psychotria mapourioides 

Mapouria ornithophila Bremek. =Psychotria anceps 

Mapouria patrisii (DC.) Lemée =Psychotria anceps 

Mapouria podocephala Müll. Arg. =Psychotria podocephala 

Mapouria remota (Benth.) Müll. Arg. =Psychotria remota 

Mapouria sclerocalyx Müll. Arg. =Rudgea sclerocalyx 

Mapouria sororia (DC.) Lemée [dubious name] =Psychotria sp. 

Mapouria spruceana Müll. Arg. =Rudgea sclerocalyx 

Mapouria subsessilis (Benth.) Müll. Arg. =Ronabea latifolia 

Mapouria subsessilis var. [gamma] angustifolia Müll. Arg. =Ronabea latifolia 

Mapouria tenuis Müll. Arg. =Geophila tenuis 

Mapouria umbrosa Müll. Arg. =Psychotria borjensis 

Mapouria vasivensis Müll. Arg. =Psychotria vasivensis 

Mapouria wonotobensis Bremek. =Psychotria wonotobensis 

Margaritopsis astrellantha (Wernham) L. Andersson =Psychotria astrellantha 

Mattuschkea galioides Kunth =Perama galioides 

Mattuschkea hispida Kunth =Perama galioides 

Mitracarpus discolor Miq. =Mitracarpus humboldtianus 

Mitracarpus fruticosus Standl. =Mitracarpus frigidus var. fruticosus 

Mitracarpus puberulus Benth. =Borreria prostrata 

Mitracarpus rude Benth. =Mitracarpus hirtus 

Mitracarpus villosus (Sw.) DC. =Mitracarpus hirtus 

Morinda tenuiflora var. leiophylla (Steyerm.) Steyerm. =Morinda tenuiflora 

Mussaenda formosa Jacq. =Rosenbergiodendron formosum 

Mussaenda spinosa Jacq. =Randia armata 

Nacibea alba Aubl. =Manettia alba 

Nacibea coccinea Aubl. =Manettia coccinea 

Naletonia violacea (Aubl.) Bremek. =Psychotria capitata subsp. amplifolia 

Nauclea tomentosa Willd. ex Roem. & Schult. =Uncaria tomentosa 

Neblinathamnus aracamunianus Steyerm. =Neblinathamnus brasiliensis 

Neblinathamnus glabratus Steyerm. =Neblinathamnus argyreus 

Nonatelia grandiflora Kunth =Palicourea grandiflora 

Nonatelia longiflora Aubl. =Palicourea longiflora 

Nonatelia lutea Aubl. [=Palicourea longiflora?] =Palicourea lutea 

Nonatelia lutea Aubl. ex Steud. =Palicourea longiflora 

Nonatelia macrophylla Kunth =Palicourea grandifolia 

Nonatelia officinalis Aubl. =Psychotria officinalis 

Nonatelia paniculata Aubl. =Psychotria paniculata 

Nonatelia racemosa Aubl. =Psychotria racemosa 

Nonatelia secundiflora (Bonpl.) Spreng. =Retiniphyllum secundiflorum 

Nonatelia violacea Aubl. =Psychotria capitata 

Oldenlandia herbacea (L.) DC. [not in flora area] 

Oribasia racemosa (Aubl.) J.F. Gmel. =Psychotria racemosa 

Ourouparia guianensis Aubl. =Uncaria guianensis 

Ourouparia tomentosa (Willd. ex Roem. & Schult.) K. Schum. =Uncaria tomentosa 

Pagamea auyantepuiensis Steyerm. =Coryphothamnus auyantepuiensis


Pagamea brevipedunculata Steyerm. =Pagamea capitata 

Pagamea capitata f. breviloba Steyerm. =Pagamea capitata subsp. conferta 

Pagamea conferta Standl. =Pagamea capitata subsp. conferta 

Pagamea coriacea var. pubescens Steyerm. =Pagamea coriacea 

Pagamea garryoides Standl. =Pagameopsis garryoides 

Pagamea plicata var. multinervia Steyerm. =Pagamea plicata 

Pagamea reducta Steyerm. =Pagamea standleyana 

Pagamea spadicea Pittier =Psychotria spadicea 

Pagamea steyermarkii Standl. =Aphanocarpus steyermarkii 

Pagamea uniflora Steyerm. =Pagamea standleyana 

Pagameopsis maguirei var. angustifolius Steyerm. =Pagameopsis maguirei 

Pagameopsis maguirei var. glabrus Steyerm. =Pagameopsis maguirei 

Pagameopsis maguirei subsp. neblinensis Steyerm. =Pagameopsis maguirei 

Pagameopsis maguirei subsp. pusillus Steyerm. =Pagameopsis maguirei 

Palicourea “latifolia (Rudge) Roem. & Schult.” [invalid name, error for Posoqueria latifolia] 

Palicourea aculeifera Steyerm. =Palicourea longistipulata subsp. chrysorrhachis 

Palicourea cayennensis Desv. =Palicourea pedunculosa 

Palicourea chionantha DC. =Psychotria mapourioides 

Palicourea chrysorrhachis Bremek. =Palicourea longistipulata subsp. chrysorrhachis 

Palicourea crocea f. heterodoxa Steyerm. =Palicourea croceoides 

Palicourea crocea var. riparia (Benth.) Griseb. =Palicourea croceoides 

Palicourea divaricata Schltdl. =Psychotria schlechtendaliana 

Palicourea elliptica Kunth =Psychotria ernestii 

Palicourea expetens Standl. =Palicourea corymbifera 

Palicourea grandifolia var. sprucei (Müll. Arg.) Steyerm. =Palicourea grandifolia 

Palicourea guianensis f. glabra Steyerm. =Palicourea guianensis subsp. guianensis 

Palicourea guianensis subsp. occidentalis Steyerm. =Palicourea guianensis subsp. guianensis 

Palicourea guianensis var. tetramera Bremek. =Palicourea guianensis subsp. guianensis 

Palicourea guianensis var. trimera Bremek. =Palicourea guianensis subsp. guianensis 

Palicourea longiflora (Aubl.) A. Rich. =Palicourea longiflora DC. 

Palicourea nicotianifolia auct., not Cham. & Schltdl. =Palicourea calophylla 

Palicourea rigida var. hirtibacca Steyerm. =Palicourea rigida 

Palicourea riparia Benth. =Palicourea croceoides 

Palicourea roraimae Standl. =Psychotria oblita 

Palicourea sprucei (Müll. Arg.) K. Schum. =Palicourea grandifolia 

Patabea alba Kunth =Psychotria lupulina subsp. rhodoleuca 

Patabea coccinea Aubl. =Ixora davisii 

Patima laxiflora Benth. =Retiniphyllum laxiflorum 

Perama dichotoma var. hirsutula Steyerm. =Perama dichotoma var. dichotoma 

Perama dichotoma var. scaposa (Gleason & Standl.) Steyerm. =Perama dichotoma var. dichotoma 

Perama galioides f. cinera Steyerm. =Perama galioides 

Perama galioides var. densipila Steyerm. =Perama galioides 

Perama galioides var. hispida (Kunth) Steyerm. =Perama galioides 

Perama galioides var. intermedia Steyerm. =Perama galioides 

Perama galioides var. longiflora Steyerm. =Perama galioides 

Perama galioides f. macrocephala Steyerm. =Perama galioides 

Perama galioides f. quaternata Steyerm. =Perama galioides 

Perama hirsuta var. stricta (Benth.) Bremek. =Perama hirsuta 

Perama scaposa Gleason & Standl. =Perama dichotoma var. dichotoma 

Perama stricta Benth. =Perama hirsuta 

Petagomoa atricapilla Bremek. =Psychotria iodotricha 

Petagomoa glabra Bremek. =Psychotria sp. 

Petagomoa hirta (Miq.) Bremek. =Psychotria bremekampiana 

Petagomoa nigricans Bremek. =Psychotria trichophoroides 

Petagomoa tricholoba (Müll. Arg.) Bremek. =Psychotria tricholoba 

Portlandia hexandra Jacq. =Coutarea hexandra 

Posoqueria gracilis (Rudge) Roem. & Schult. =Posoqueria latifolia subsp. gracilis 

Posoqueria intermedia Miq. =Posoqueria trinitatis 

Posoqueria panamensis subsp. grandiflora (H. Karst.) Steyerm. =Posoqueria panamensis 

Pseudochimarrhis difformis Benoist =Chimarrhis sp. 

Pseudochimarrhis turbinata Benoist =Chimarrhis turbinata 

Psychotria alboviridula K. Krause =Psychotria remota 

Psychotria aligera Steyerm. =Notopleura aligera 

Psychotria amazonica Müll Arg. =Palicourea grandifolia

500 


Psychotria amplifolia Raeusch. =Psychotria capitata subsp. amplifolia 

Psychotria amplinoda Steyerm. =Notopleura tapajozensis 

Psychotria amplinoda var. ayangannae Steyerm. =Notopleura tapajozensis 

Psychotria aneurophylla (Standl.) Steyerm. =Notopleura aneurophylla 

Psychotria anomothyrsa K. Schum. & Donn. Sm. =Notopleura anomothyrsa 

Psychotria arcuata Benth. =Psychotria capitata subsp. inundata 

Psychotria arenaria Standl. & Steyerm. =Psychotria ceratantha 

Psychotria atabapoensis Standl. =Psychotria subundulata 

Psychotria atricapilla Bremek. =Psychotria iodotricha 

Psychotria aubletiana var. andina Steyerm. =Psychotria aubletiana 

Psychotria aubletiana var. cacuminis (Steyerm.) Steyerm. =Psychotria aubletiana 

Psychotria aubletiana f. tomentella Steyerm. =Psychotria aubletiana 

Psychotria auyantepuiensis Standl. =Psychotria imthurniana 

Psychotria avia Standl. & Steyerm. =Rudgea stipulacea 

Psychotria axillaris Willd. =Ronabea latifolia 

Psychotria bahiensis var. bahiensis (sensu Steyerm.) [P. bahiensis is endemic to Bahia, Brazil] =Psychotria acuminata 

Psychotria bahiensis var. cornigera (Benth.) Steyerm. =Psychotria cornigera 

Psychotria barbiflora DC. =Psychotria hoffmannseggiana 

Psychotria barcellana Müll. Arg. =Psychotria poeppigiana subsp. barcellana 

Psychotria berteroana subsp. venezuelica Steyerm. =Psychotria berteroana 

Psychotria blepharophylla (Standl.) Steyerm. =Psychotria medusula 

Psychotria brachybotrya Müll. Arg. =Psychotria gracilenta 

Psychotria brachyloba Müll. Arg. =Palicourea brachyloba 

Psychotria bracteata DC. =Psychotria pseudinundata 

Psychotria bracteata var. intermedia Müll. Arg. =Psychotria maguireorum 

Psychotria bracteata var. latifolia Müll. Arg. =Psychotria maguireorum 

Psychotria bracteata var. tenuifolia Müll. Arg. =Psychotria maguireorum 

Psychotria breweri Steyerm. =Psychotria tatei 

Psychotria cacuminis Steyerm. =Psychotria aubletiana 

Psychotria cacuminis Standl. =Psychotria everardii 

Psychotria calviflora Steyerm. =Psychotria colorata 

Psychotria capitata var. roraimensis (Wernham) Steyerm. =Psychotria capitata subsp. inundata 

Psychotria capitata var. septentrionalis Steyerm. =Psychotria capitata subsp. inundata 

Psychotria cardiomorpha subsp. perpusilla (Steyerm.) C.M. Taylor & A. Pool =Psychotria cardiomorpha 

Psychotria chionantha DC. =Psychotria mapourioides 

Psychotria chlorantha Benth. =Psychotria anceps 

Psychotria chondroloma Standl. =Psychotria schomburgkii 

Psychotria cincta Standl. =Psychotria longicuspis 

Psychotria cordifolia Kunth [nom. illeg.] =Psychotria cardiomorpha 

Psychotria cordifolia subsp. perpusilla Steyerm. [nom. illeg.] =Psychotria cardiomorpha 

Psychotria cornifolia Kunth =Rudgea cornifolia 

Psychotria corymbifera Müll. Arg. =Palicourea corymbifera 

Psychotria coussareoides var. ciliata Steyerm. =Psychotria coussareoides 

Psychotria crassa Benth. =Notopleura crassa 

Psychotria crassa f. alba Steyerm. =Notopleura crassa 

Psychotria crassa f. angustior Steyerm. =Notopleura crassa 

Psychotria crocea Sw. =Palicourea crocea 

Psychotria cubitalis Standl. & Steyerm. =Psychotria capitata subsp. inundata 

Psychotria cuspidata Bredem. ex Roem. & Schult. [not in flora area] 

Psychotria davidsei Steyerm. [nom. illeg.] =Psychotria berryi 

Psychotria deflexa subsp. campyloneura (Müll. Arg.) Steyerm. =Psychotria campyloneura 

Psychotria deflexa subsp. venulosa (Müll. Arg.) Steyerm. =Psychotria venulosa 

Psychotria dichotoma (Rudge) Bremek. [nom. illeg.] =Psychotria platypoda 

Psychotria didymocarpos (A. Rich.) Lemée =Psychotria acuminata 

Psychotria edaphothrix Steyerm. =Palicourea bracteosa 

Psychotria erecta (Aubl.) Standl. & Steyerm. =Ronabea latifolia 

Psychotria erecta f. fluctuans (Standl.) Steyerm. =Ronabea latifolia 

Psychotria erythrophylla Müll. Arg. =Psychotria hoffmannseggiana 

Psychotria everardii subsp. neblinensis Steyerm. =Psychotria everardii 

Psychotria everardii subsp. sipapoensis Steyerm. =Psychotria duricoria var. duricoria 

Psychotria everardii subsp. tatei (Standl.) Steyerm. =Psychotria tatei 

Psychotria fastigiata (Kunth) Spreng. =Palicourea fastigiata 

Psychotria fimbriata Benth. =Rudgea cornifolia 

Psychotria flavovirens Suess. =Psychotria lupulina subsp. rhodoleuca


Psychotria flexuosa Willd. [nom. superfl.] =Psychotria paniculata 

Psychotria fluctuans Standl. =Ronabea latifolia 

Psychotria fluctuans var. angustifolia (Müll. Arg.) Standl. =Ronabea latifolia 

Psychotria fockeana Miq. =Psychotria carthagenensis 

Psychotria foetidiflora Standl. =Psychotria boliviana 

Psychotria foveolata Ruiz & Pav. =Psychotria carthagenensis 

Psychotria glabrescens Müll. Arg. [not in flora area] 

Psychotria glandulicalyx var. densipubens Steyerm. =Psychotria glandulicalyx 

Psychotria glaucescens Kunth =Psychotria horizontalis 

Psychotria grandifolia Willd. ex Roem. & Schult. =Palicourea grandifolia 

Psychotria guadalupensis (DC.) R.A. Howard =Notopleura guadalupensis 

Psychotria guianensis (Aubl.) Rusby [nom. illeg., non Psychotria guianensis (Aubl.) Raeusch.] =Palicourea guianensis 

Psychotria guianensis (Aubl.) Raeusch. [nomen nudum] =Palicourea guianensis 

Psychotria hancorniifolia Benth. =Psychotria vellosiana 

Psychotria hancorniifolia var. angustifolia Müll. Arg. =Psychotria vellosiana 

Psychotria hancorniifolia var. longifolia Müll. Arg. =Psychotria vellosiana 

Psychotria heterocarpa Standl. & Steyerm. =Psychotria capitata subsp. inundata 

Psychotria hoffmannseggiana var. erythrophylla (Müll. Arg.) Steyerm. =Psychotria hoffmannseggiana 

Psychotria homoplastica Moore =Psychotria stipulosa 

Psychotria horizontalis var. glaucescens (Kunth) Steyerm. =Psychotria horizontalis 

Psychotria horizontalis var. psilophylla Steyerm. =Psychotria horizontalis 

Psychotria humboldtiana var. ornata Müll. Arg. =Psychotria humboldtiana 

Psychotria imthurniana f. ptariensis (Standl. & Steyerm.) Steyerm. =Psychotria imthurniana 

Psychotria inundata K. Krause [nom. illeg.] =Rudgea cornifolia 

Psychotria inundata Benth. =Psychotria capitata subsp. inundata 

Psychotria involucrata Sw. [nom. superfl.] =Psychotria officinalis 

Psychotria iodotricha subsp. atricapilla (Bremek.) Steyerm. =Psychotria iodotricha 

Psychotria kukenanensis Steyerm. =Psychotria boliviana 

Psychotria lancigera Standl. =Palicourea lancigera 

Psychotria longiflora Poir. [nom. illeg.] =Palicourea longiflora 

Psychotria longiflora Willd. =Palicourea longiflora 

Psychotria longistipula Benth. =Psychotria racemosa 

Psychotria longistipulata Müll. Arg. =Palicourea longistipulata 

Psychotria loretensis Standl. =Psychotria longicuspis 

Psychotria lupulina var. maypurensis (Humb. & Bonpl. ex Roem. & Schult.) Steyerm. =Psychotria lupulina subsp. 

rhodoleuca 

Psychotria lupulina f. pubens Steyerm. =Psychotria lupulina subsp. rhodoleuca 

Psychotria macrophylla Ruiz & Pav. =Notopleura macrophylla 

Psychotria macrophylla subsp. anomothyrsa (K. Schum. & Donn. Sm.) Steyerm. =Notopleura anomothyrsa 

Psychotria macropoda Ruiz & Pav. =Geophila macropoda 

Psychotria maguirei Standl. =Rudgea stipulacea 

Psychotria mapourioides var. chionantha (DC.) Steyerm. =Psychotria mapourioides 

Psychotria mapourioides var. opaca (Bremek.) Steyerm. =Psychotria mapourioides 

Psychotria mapuria Roem. & Schult. [nom. illeg.] =Psychotria mapourioides 

Psychotria martiana Müll. Arg. =Psychotria platypoda 

Psychotria maypurensis Humb. & Bonpl. ex Roem. & Schult. =Psychotria lupulina subsp. rhodoleuca 

Psychotria merumensis Steyerm. =Notopleura merumensis 

Psychotria microbracteata Steyerm. =Notopleura microbracteata 

Psychotria microcephala (Willd. ex Roem. & Schult.) Müll. Arg. [nom. illeg.] =Psychotria prunifolia 

Psychotria microcephala (Miq.) Miq. =Psychotria hoffmannseggiana 

Psychotria microcephala var. tripotamica Müll. Arg. =Psychotria prunifolia 

Psychotria microdon var. meridionalis Steyerm. =Psychotria microdon 

Psychotria multiramosa Steyerm. =Notopleura multiramosa 

Psychotria necopinata Standl. =Psychotria pacimonica 

Psychotria nervosa Benth. =Psychotria lupulina subsp. lupulina 

Psychotria nitida Willd. [nom. superfl.] =Psychotria mapourioides 

Psychotria nitidella Müll. Arg. =Palicourea nitidella 

Psychotria officinalis (Aubl.) Raeusch. [nomen nudum] =Psychotria officinalis 

Psychotria palicoureoides Müll. Arg. [nom. illeg.] =Psychotria microbotrys 

Psychotria paradoxa Müll. Arg. =Psychotria amplectens 

Psychotria patens Sw. [not in flora area] 

Psychotria pedunculosa A. Rich. =Palicourea pedunculosa 

Psychotria pendula subsp. trinitensis Urb. =Notopleura perpapillifera 

Psychotria perpapillifera Steyerm. =Notopleura perpapillifera

502 


Psychotria persimilis Müll. Arg. =Psychotria lupulina subsp. rhodoleuca 

Psychotria phaneroloma var. angustior Steyerm. =Psychotria phaneroloma 

Psychotria phaneroneura Standl. =Psychotria everardii 

Psychotria ptariensis Standl. & Steyerm. =Psychotria imthurniana 

Psychotria quadrifolia Rudge =Palicourea quadrifolia 

Psychotria racemifera Standl. =Psychotria polycephala 

Psychotria racemosa (Aubl.) Raeusch. [nom. illeg.] =Psychotria racemosa 

Psychotria ramiflora Rusby [records due to misidentifications] 

Psychotria rhodoleuca Müll. Arg. =Psychotria lupulina subsp. rhodoleuca 

Psychotria rigida Bredem. ex Willd. ex Roem. & Schult. =Palicourea rigida 

Psychotria romboutsii Bremek. =Psychotria amplectens 

Psychotria ronaldii Steyerm. =Psychotria boliviana 

Psychotria roraimensis Wernham =Psychotria capitata subsp. inundata 

Psychotria rosea f. calvescens Steyerm. =Psychotria rosea 

Psychotria rubra (Willd. ex Roem. & Schult.) Müll. Arg. =Psychotria hoffmannseggiana 

Psychotria rudgei Bremek. =Psychotria lupulina subsp. lupulina 

Psychotria sancti-caroli Standl. =Palicourea longistipulata subsp. longistipulata 

Psychotria sandwithiana (Steyerm.) Steyerm. =Notopleura sandwithiana 

Psychotria saülensis Steyerm. =Notopleura saülensis 

Psychotria sessiliflora Humb. & Bonpl. ex Schult. (Gentianaceae) =Tapeinostemon sessiliflorum 

Psychotria sessilis (Vell.) Müll. Arg. [nom. illeg.] =Psychotria vellosiana 

Psychotria sessilis var. angustifolia (Müll. Arg.) Steyerm. [nom. illeg.] =Psychotria vellosiana 

Psychotria setifera Benth. =Psychotria capitata subsp. inundata 

Psychotria sororiella Müll. Arg. =Psychotria turbinella var. sororiella 

Psychotria sororopanensis Standl. & Steyerm. =Psychotria vellosiana 

Psychotria speluncae subsp. brevicalyx Steyerm. =Psychotria speluncae 

Psychotria speluncae subsp. exserta Steyerm. =Psychotria speluncae 

Psychotria spicata Müll. Arg. =Psychotria spiciflora 

Psychotria sprucei Müll. Arg. =Palicourea grandifolia 

Psychotria steinii Steyerm. =Psychotria leiantha 

Psychotria subcuspidata Müll. Arg. =Psychotria cornigera 

Psychotria tapajozensis Standl. =Notopleura tapajozensis 

Psychotria thesceloantha Steyerm. =Notopleura thesceloantha 

Psychotria triphylla (DC.) Müll. Arg. =Palicourea triphylla 

Psychotria uliginosa Sw. =Notopleura uliginosa 

Psychotria urophylla Schltdl. =Psychotria acuminata 

Psychotria vaupesana Standl. ex Steyerm. =Psychotria spiciflora 

Psychotria vernicifolia Standl. =Psychotria everardii 

Psychotria verrucosa Müll. Arg. =Palicourea corymbifera 

Psychotria viburnoides Kunth =Psychotria anceps 

Psychotria violacea Aubl. ? [name of uncertain application] =Coccocypselum sp. 

Psychotria xanthocephala Müll. Arg. =Psychotria prunifolia 

Psychotria yutajensis Steyerm. =Psychotria ceratantha 

Randia caracasana Standl. =Randia dioica 

Randia formosa (Jacq.) K. Schum. =Rosenbergiodendron formosum 

Randia formosa var. densiflora Bartl. ex K. Schum. =Rosenbergiodendron densiflorum 

Randia formosa var. formosa =Rosenbergiodendron formosum 

Randia formosa var. nitida K. Schum. =Rosenbergiodendron formosum var. nitidum 

Randia orinocensis Rusby =Rosenbergiodendron formosum var. nitidum 

Randia ruizana DC. =Rosenbergiodendron longiflorum 

Randia spinosa (Jacq.) H. Karst. =Randia armata 

Relbunium croceum (Ruiz & Pav.) K. Schum. =Galium hypocarpium 

Relbunium glaberrimum Standl. =Galium hypocarpium 

Relbunium guadalupense (Spreng.) Urb. [not in flora area] 

Relbunium hypocarpium (L.) Hemsl. =Galium hypocarpium 

Relbunium hypocarpium subsp. hypocarpium =Galium hypocarpium subsp. hypocarpium 

Relbunium hypocarpium subsp. nitidum (Kunth) Ehrend. =Galium hypocarpium 

Relbunium nitidum (Kunth) K. Schum. =Galium hypocarpium 

Remijia berryi Steyerm. =Remijia pedunculata 

Remijia densiflora f. glabricalyx Steyerm. =Remijia densiflora var. densiflora 

Remijia spruceana Benth. ex K. Schum. =Remijia firmula 

Retiniphyllum angustiflorum K. Krause =Retiniphyllum concolor 

Retiniphyllum cearense Standl. =Retiniphyllum scabrum 

Retiniphyllum erythranthum Standl. =Retiniphyllum scabrum


Retiniphyllum laxiflorum var. longilobum Steyerm. =Retiniphyllum laxiflorum var. laxiflorum 

Retiniphyllum maguirei var. reticulatum Steyerm. =Retiniphyllum maguirei 

Retiniphyllum martianum Müll. Arg. =Retiniphyllum concolor 

Retiniphyllum pallidum Müll. Arg. =Retiniphyllum truncatum 

Retiniphyllum scabrum var. ayangannense Steyerm. = Retinuphyllum scabrum 

Retiniphyllum scabrum var. erythranthum (Standl.) Steyerm. =Retiniphyllum scabrum 

Retiniphyllum schomburgkii var. angustiflorum Huber =Retiniphyllum schomburgkii 

Retiniphyllum schomburgkii var. hirticalyx Steyerm. =Retiniphyllum schomburgkii 

Retiniphyllum schomburgkii subsp. occidentale Steyerm. =Retiniphyllum schomburgkii 

Retiniphyllum truncatum var. angustifolium Steyerm. =Retiniphyllum truncatum 

Richardsonia scabra (L.) A. St.-Hil. =Richardia scabra 

Ronabea erecta Aubl. =Ronabea latifolia 

Ronabea latifolia var. hispidula Bremek. =Ronabea latifolia 

Ronabea morindoides A. Rich. =Psychotria morindoides 

Rondeletia capitatum Benth. =Coccocypselum aureum 

Rondeletia glomerulata Spreng. [dubious name] 

Rondeletia microdon DC. =Psychotria microdon 

Rondeletia repens L. =Geophila repens 

Rondeletia rupicola Spruce ex K. Schum. =Sipaneopsis rupicola 

Rondeletia rupicola var. chiribiquetana R.E. Schult. =Sipaneopsis rupicola 

Rubia nitida Kunth =Galium hypocarpium 

Rudgea avia (Standl. & Steyerm.) Steyerm. =Rudgea stipulacea 

Rudgea berryi Steyerm. & Dwyer =Rudgea klugii 

Rudgea bolivarensis Steyerm. =Rudgea hostmanniana subsp. hostmanniana 

Rudgea carolina Standl. & Steyerm. =Rudgea lanceifolia 

Rudgea corocoroensis Steyerm. =Rudgea hostmanniana subsp. hostmanniana 

Rudgea dasyantha Bremek. =Rudgea lanceifolia 

Rudgea duidae (Standl.) Steyerm. =Rudgea stipulacea 

Rudgea fimbriata (Benth.) Standl. =Rudgea cornifolia 

Rudgea fissistipula Müll. Arg. =Rudgea lanceifolia 

Rudgea laurifolia Steyerm. =Rudgea sclerocalyx 

Rudgea maypurensis Standl. =Rudgea hostmanniana subsp. maypurensis 

Rudgea morichensis Steyerm. =Rudgea sclerocalyx 

Rudgea obscura Sandwith =Psychotria astrellantha 

Rudgea ovalifolia Salisb. =Rudgea lanceifolia 

Rudgea schomburgkiana Benth. =Rudgea crassiloba 

Rudgea sipapoensis Steyerm. =Rudgea lanceifolia 

Rudgea standleyana Steyerm. =Rudgea klugii 

Rudgea tillettii Steyerm. =Psychotria coussareoides 

Sabicea aspera var. glabrescens K. Schum. =Sabicea glabrescens 

Sabicea aspera var. velutina (Benth.) K. Schum. =Sabicea velutina 

Sabicea colombiana Wernham =Sabicea panamensis 

Sabicea edulis (Rich.) Seem. ex B.D. Jackson =Alibertia edulis 

Sabicea glabrescens var. oblongifolia (Miq.) Sandwith =Sabicea oblongifolia 

Sabicea guianensis Wernham [nom. illeg.] =Sabicea velutina 

Sabicea guianensis (Aubl.) Baill. =Patima guianensis 

Sabicea hirsuta Kunth =Sabicea villosa 

Sabicea hirsuta var. adpressa Wernham =Sabicea villosa var. adpressa 

Sabicea leucotricha K. Krause =Sabicea velutina 

Sabicea velutina var. oblongifolia Miq. =Sabicea oblongifolia 

Saldanha illustris Vell. =Hillia illustris 

Schradera ligularis Rudge =Psychotria ligularis 

Schradera spicata Spruce ex Hook. f. [nomen nudum] =Stachyarrhena spicata 

Schwenkfeldia aurea Spreng. =Coccocypselum aureum 

Semaphyllanthe obovata (Ducke) L. Andersson =Calycophyllum obovatum 

Semaphyllanthe venezuelensis (Steyerm.) L. Andersson =Calycophyllum venezuelense 

Sickingia orinocensis Spreng. =Platycarpum orinocense var. orinocense 

Sickingia pisoniiformis Baill. =Simira pisoniiformis 

Sickingia tinctoria (Aubl.) Lemée =Simira tinctoria 

Simira erythroxylon var. saxicola Steyerm. =Simira rubescens 

Sipanea acinifolia Spruce ex Sprague =Sipanea veris 

Sipanea colombiana Wernham =Sipanea pratensis var. dichotoma 

Sipanea coppenamensis Bremek. =Sipanea pratensis var. pratensis 

Sipanea dichotoma Kunth =Sipanea pratensis var. dichotoma

504 


Sipanea glomerata var. paucinervia Steyerm. =Sipanea glomerata 

Sipanea palustris (A. Rich.) J.H. Kirkbr. =Sipanea wilson-brownei 

Sipanea palustris Seem. [nom. illeg.] =Limnosipanea palustris 

Sipanea pratensis f. brachycarpa Steyerm. =Sipanea pratensis var. dichotoma 

Sipanea pratensis f. breviflora Steyerm. =Sipanea pratensis var. dichotoma 

Sipanea pratensis f. glabriloba Steyerm. =Sipanea pratensis var. dichotoma 

Sipanea pratensis f. glabrior Steyerm. =Sipanea pratensis var. dichotoma 

Sipanea pubinoda Steyerm. =Sipanea pratensis var. dichotoma 

Sipanea radicans Endl. =Sipanea biflora 

Sipanea rupicola Spruce ex K. Schum. [nomen nudum] =Sipaneopsis rupicola 

Sipanea spraguei Wernham =Sipanea veris 

Sipaneopsis wurdackii Steyerm. =Sipaneopsis maguirei 

Solena gracilis Rudge =Posoqueria latifolia subsp. gracilis 

Solena latifolia Rudge =Posoqueria latifolia 

Spermacoce alata Aubl. =Borreria alata 

Spermacoce apiculata Willd. ex Roem. & Schult. =Diodia apiculata 

Spermacoce aspera Aubl. =Mitracarpus hirtus 

Spermacoce assurgens Ruiz & Pav. =Borreria remota 

Spermacoce aturensis Kunth =Borreria capitata var. tenella 

Spermacoce caerulescens Aubl. =Borreria latifolia 

Spermacoce capitata Ruiz & Pav. =Borreria capitata 

Spermacoce capitellata Willd. ex Roem. & Schult. =Borreria remota 

Spermacoce densiflora (DC.) Alain =Borreria densiflora 

Spermacoce diffusa Willd. ex Roem. & Schult. =Mitracarpus diffusus 

Spermacoce exilis (L.O. Williams) C.D. Adams =Borreria repens 

Spermacoce ferruginea A. St.-Hil. =Borreria capitata var. capitata 

Spermacoce frigida Willd. ex Roem. & Schult. =Mitracarpus frigidus 

Spermacoce guianensis Bremek. =Borreria remota 

Spermacoce hirta L. =Mitracarpus hirtus 

Spermacoce hyssopifolia Willd. ex Roem. & Schult. =Diodia hyssopifolia 

Spermacoce latifolia Aubl. =Borreria latifolia 

Spermacoce linearis Willd. ex Roem. & Schult. =Diodia hyssopifolia 

Spermacoce longifolia Aubl. =Mitracarpus hirtus 

Spermacoce longifolia Aubl. ex Miq. =Diodia ocymifolia 

Spermacoce mauritiana Gideon =Borreria repens 

Spermacoce ocymifolia Willd. ex Roem. & Schult. =Diodia ocymifolia 

Spermacoce ocymoides Burm. f. [not in flora area, neotropical specimens = Borreria ovalifolia or prostrata] =Borreria 

prostrata 

Spermacoce orinocensis Willd. ex Roem. & Schult. =Borreria capitata var. tenella 

Spermacoce prostrata Aubl. =Borreria prostrata 

Spermacoce radula Willd. & Hoffmanns. ex Roem. & Schult. =Diodia radula 

Spermacoce remota Lam. =Borreria remota 

Spermacoce rigida Kunth [nom. illeg.] =Diodia apiculata 

Spermacoce rigida Willd. ex Roem. & Schult. [nom. illeg.] =Diodia apiculata 

Spermacoce spinosa Sw. [nom. illeg.] =Borreria densiflora 

Spermacoce tenella Kunth =Borreria capitata var. tenella 

Spermacoce verticillata L. =Borreria verticillata 

Spermacoce villosa Sw. =Mitracarpus hirtus 

Sphinctanthus rupestris Benth. =Sphinctanthus striiflorus 

Sprucea rubescens Benth. =Simira rubescens 

Stachyarrhena longifolia Hook. f. =Stachyarrhena spicata 

Stachyarrhena spicata var. multinervis K. Schum. =Stachyarrhena spicata 

Stannia grandiflora H. Karst. =Posoqueria panamensis 

Stannia panamensis Walp. & Duchass. =Posoqueria panamensis 

Strempelia fimbriata (Benth.) Bremek. =Rudgea cornifolia 

Strempelia guyanensis A. Rich. =Rudgea guyanensis 

Synisoon schomburgkianum Baill. =Retiniphyllum laxiflorum var. laxiflorum 

Tapogomea alba Aubl. =Psychotria ulviformis 

Tapogomea carapichea Poir. =Psychotria ligularis var. carapichea 

Tapogomea glabra Aubl. =Psychotria medusula 

Tapogomea purpurea Aubl. =Psychotria variegata 

Tapogomea tomentosa Aubl. =Psychotria poeppigiana 

Tapogomea violacea Aubl. =Psychotria apoda 

Tetramerium sessifolium Kunth =Faramea sessilifolia


Thieleodoxa lanceolata Cham. =Alibertia edulis var. edulis 

Thieleodoxa nitidula Bremek. =Alibertia edulis var. edulis 

Tocoyena guianensis var. communis Steyerm. =Tocoyena guianensis 

Tocoyena guianensis var. glabriuscula Steyerm. =Tocoyena guianensis 

Tontanea guianensis Aubl. =Coccocypselum guianense 

Uragoga alba (Ruiz & Pav.) Kuntze =Psychotria alba 

Uragoga alba (Aubl.) Kuntze =Psychotria ulviformis 

Uragoga glabra (Aubl.) Kuntze =Psychotria medusula 

Uragoga hirta (Miq.) Pulle =Psychotria bremekampiana 

Uragoga kappleri (Miq.) Pulle =Psychotria kappleri 

Uragoga oblonga (DC.) Kuntze =Psychotria oblonga 

Uragoga purpurea (Aubl.) Kuntze =Psychotria variegata 

Uragoga tomentosa (Aubl.) K. Schum. =Psychotria poeppigiana 

Uragoga tomentosa (Aubl.) Kuntze =Psychotria poeppigiana 

Uragoga uliginosa (Sw.) Kuntze =Notopleura uliginosa 

Uragoga violacea (Aubl.) Pulle =Psychotria apoda 

Valantia hypocarpia L. =Galium hypocarpium 

Virecta biflora L. f. =Sipanea biflora 

Warszewiczia obovata Ducke =Calycophyllum obovatum 

Warszewiczia schomburgkiana Klotzsch =Warszewiczia coccinea 

Yutajea liesneri Steyerm. =Isertia parviflora 

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Botanical Garden Press, New York.

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Kew Bull. 50: 761-771. 

RUTACEAE 

J.A. Kallunki 

Adiscanthus fusciflorus Ducke 

Amyris guianensis Aubl. 

Angostura trifoliata (Willd.) T.S. Elias 

Apocaulon carnosum R.S. Cowan 

Citrus spp. [species and cultivars cultivated and perhaps escaping] 

Conchocarpus acuminatus (Pilg.) Kallunki & Pirani 

Conchocarpus fanshawei (Sandwith) Kallunki & Pirani 

Conchocarpus guyanensis (Pulle) Kallunki & Pirani 

Conchocarpus heterophyllus (A. St.-Hil.) Kallunki & Pirani 

Conchocarpus longifolius (A. St.-Hil.) Kallunki & Pirani 

Conchocarpus ramiflorus (Spruce ex Engl.) Kallunki & Pirani 

Conchocarpus toxicarius (Engl.) Kallunki & Pirani 

Conchocarpus ucayalinus (Huber) Kallunki & Pirani 

Decagonocarpus cornutus R.S. Cowan 

Decagonocarpus oppositifolius Spruce ex Engl. 

Ertela trifolia (L.) Kuntze [=Monnieria trifolia L.] 

Erythrochiton brasiliensis Nees & Mart. 

Esenbeckia cowanii Kaastra 

Esenbeckia grandiflora Mart. 

Esenbeckia grandiflora var. grandiflora 

Esenbeckia pilocarpoides Kunth 

Esenbeckia pilocarpoides subsp. pilocarpoides 

Galipea davisii Sandwith 

Galipea trifoliata Aubl. 

Hortia coccinea Spruce ex Engl. 

Hortia neblinensis Maguire & B. Boom 

Hortia regia Sandwith 

Hortia vandelliana Groppo 

Metrodorea flavida K. Krause 

Murraya paniculata (L.) Jack [cultivated; naturalized?] 

Neoraputia paraensis (Ducke) Emmerich 

Pilocarpus demerarae Sandwith 

Pilocarpus microphyllus Stapf ex Wardleworth 

AM 

AM BO DA 

AM 

FG 

FG 

GU 

SU FG 

BO SU 

BO DA SU FG 

AM 

GU FG 

AM 

AM 

AM 

BO GU FG 



FG 





AM BO GU FG 

AM 

AM 

SU FG 

AM 

BO GU 

SU 

BO GU SU FG 

BO DA FG 

GU 

SU

508 


Pilocarpus racemosus Vahl 

Pilocarpus racemosus subsp. racemosus 

Raputia aromatica Aubl. 

Raputia brevipedunculata Kallunki 

Raputia maroana (R.S. Cowan) Kallunki 

Raputia neblinensis (R.S. Cowan) Kallunki 

Raputia szczerbanii (Steyerm.) Kallunki 

Rauia resinosa Nees & Mart. 

Rauia spicata Haye 

Rauia subtruncata Steyerm. 

Raveniopsis abyssicola R.S. Cowan 

Raveniopsis breweri Steyerm. 

Raveniopsis capitata R.S. Cowan 

Raveniopsis cowaniana Steyerm. & Luteyn 

Raveniopsis fraterna R.S. Cowan 

Raveniopsis jauaensis Steyerm. 

Raveniopsis linearis (Gleason) R.S. Cowan 

Raveniopsis nubicola R.S. Cowan 

Raveniopsis paruana (R.S. Cowan) R.S. Cowan 

Raveniopsis peduncularis Pittier & Lasser 

Raveniopsis ruellioides (Oliv.) R.S. Cowan 

Raveniopsis sericea R.S. Cowan 

Raveniopsis stelligera (R.S. Cowan) R.S. Cowan 

Raveniopsis steyermarkii R.S. Cowan 

Raveniopsis tomentosa Gleason 

Raveniopsis trifoliolata R.S. Cowan 

Rutaneblina pusilla Steyerm. & Luteyn 

Spathelia ulei (Engl. ex Harms) R.S. Cowan & Brizicky 

Spiranthera guianensis Sandwith 

Spiranthera parviflora Sandwith 

Ticorea foetida Aubl. 

Ticorea longiflora DC. 

Ticorea pedicellata DC. 

Triphasia trifolia (Burm. f.) P. Wilson [cultivated and apparently escaped] 

Zanthoxylum amapaense (Albuq.) P.G. Waterman 

Zanthoxylum apiculatum (Sandwith) P.G. Waterman 

Zanthoxylum caribaeum Lam. 

Zanthoxylum compactum (Huber ex Albuq.) P.G. Waterman 

Zanthoxylum ekmanii (Urb.) Alain 

Zanthoxylum fagara (L.) Sarg. 

Zanthoxylum flavum Vahl 

Zanthoxylum hermaphroditum Willd. [type “Guiana”] 

Zanthoxylum huberi P.G. Waterman 

Zanthoxylum juniperinum Poepp. 

Zanthoxylum martinicense (Lam.) DC. 

Zanthoxylum nigrum Mart. 

Zanthoxylum pentandrum (Aubl.) R.A. Howard 

Zanthoxylum rhoifolium Lam. 

Zanthoxylum sp. A 

Zanthoxylum sp. B 

Zanthoxylum syncarpum Tul. 


Almeidea guyanensis Pulle =Conchocarpus guyanensis 

Amyris decandra (Aubl.) Willd. (Burseraceae) =Protium decandrum 

Amyris enneandra (Aubl.) Willd. (Burseraceae) =Protium decandrum 

Amyris guyanensis (Aubl.) Willd. (Burseraceae) =Protium guianense 

Angostura acuminata (Pilg.) Albuq. =Conchocarpus acuminatus 

Angostura cuspare Roem. & Schult. [nom. illeg.] =Angostura trifoliata 

Angostura fanshawei (Sandwith) Albuq. =Conchocarpus fanshawei 

Angostura fontanesiana (A. St.-Hil.) Albuq. [not in flora area] 

Angostura pentagyna (A. St.-Hil.) Albuq. =Conchocarpus longifolius 

Angostura ramiflora (Spruce ex Engl.) Albuq. =Conchocarpus ramiflorus 

Angostura toxicaria (Spruce ex Engl.) Albuq. =Conchocarpus toxicarius 

AM 

AM 

BO FG 

BO FG 

FG 

FG 

BO 

GU SU 

BO SU 

BO GU 

AM 

AM 

AM 

AM 

AM 

AM 

BO 

BO 

BO 

BO 

BO 

AM BO GU 

AM 

AM 

AM 

BO 

BO 

BO 

GU 

AM BO GU 

BO GU 

AM BO GU 

GU SU FG 

SU FG 

GU SU FG 

GU FG 

BO DA FG 

BO GU 

BO GU 

DA GU 


BO FG? 

FG? 

BO 

DA SU 

BO DA 

BO DA GU 

SU? 

SU FG 

AM 

AM 

BO 

BO 

BO 

GU SU FG


Angostura ucayalina (Huber) Albuq. =Conchocarpus ucayalinus 

Angostura ulei (K. Krause) Gereau =Conchocarpus ucayalinus 

Bonplandia angostura Spreng. =Angostura trifoliata 

Bonplandia angostura Rich. ex B.D. Jacks. =Angostura trifoliata 

Bonplandia trifoliata Willd. =Angostura trifoliata 

Chalcas exotica (L.) Millsp. =Murraya paniculata 

Chalcas paniculata L. =Murraya paniculata 

Conchocarpus pentagynus (A. St.-Hil.) Kallunki & Pirani [nomen nudum] =Conchocarpus longifolius 

Cusparia angostura (Rich. ex B.D. Jacks.) A. Lyons =Angostura trifoliata 

Cusparia fanshawei Sandwith =Conchocarpus fanshawei 

Cusparia febrifuga Humb. =Angostura trifoliata 

Cusparia officinalis (J. Hancock) Engl. =Angostura trifoliata 

Cusparia pentagyna (A. St.-Hil.) Engl. =Conchocarpus longifolius 

Cusparia ramiflora Spruce ex Engl. =Conchocarpus ramiflorus 

Cusparia toxicaria Spruce ex Engl. =Conchocarpus toxicarius 

Cusparia trifoliata (Willd.) Engl. =Angostura trifoliata 

Cusparia ucayalina Huber =Conchocarpus ucayalinus 

Decagonocarpus cornutus var. congestus R.S. Cowan =Decagonocarpus cornutus 

Diomma fruticosa Steyerm. =Spathelia ulei 

Diomma ulei Engl. ex Harms =Spathelia ulei 

Esenbeckia obovalifolia Pittier =Esenbeckia grandiflora var. grandiflora 

Esenbeckia venezuelensis Engl. =Esenbeckia pilocarpoides subsp. pilocarpoides 

Fagara amapaensis Albuq. =Zanthoxylum amapaense 

Fagara apiculata Sandwith =Zanthoxylum apiculatum 

Fagara caribaea (Lam.) Krug & Urb. =Zanthoxylum caribaeum 

Fagara compacta Huber ex Albuq. =Zanthoxylum compactum 

Fagara ekmanii Urb. =Zanthoxylum ekmanii 

Fagara flava Krug & Urb. =Zanthoxylum flavum 

Fagara guianensis Lam. =Zanthoxylum hermaphroditum 

Fagara martinicensis Lam. =Zanthoxylum martinicense 

Fagara ocumarensis Pittier =Zanthoxylum juniperinum 

Fagara pentandra Aubl. =Zanthoxylum pentandrum 

Fagara perrottetii DC. =Zanthoxylum rhoifolium 

Fagara pterota L. =Zanthoxylum fagara 

Fagara rhoifolia (Lam.) Engl. =Zanthoxylum rhoifolium 

Fagara syncarpa (Tul.) Engl. =Zanthoxylum syncarpum 

Galipea aromatica (Aubl.) Spreng. =Raputia aromatica 

Galipea cusparia A. St.-Hil. ex DC. [nom. illeg.] =Angostura trifoliata 

Galipea febrifuga (Humb.) Baill. =Angostura trifoliata 

Galipea fissa Miq. =Galipea trifoliata 

Galipea heterophylla A. St.-Hil. =Conchocarpus heterophyllus 

Galipea longifolia A. St.-Hil. =Conchocarpus longifolius 

Galipea officinalis J. Hancock =Angostura trifoliata 

Galipea pentagyna A. St.-Hil. =Conchocarpus longifolius 

Galipea stelligera R.S. Cowan =Raveniopsis stelligera 

Lubaria szczerbanii Steyerm. =Raputia szczerbanii 

Monnieria trifolia L. =Ertela trifolia 

Murraya exotica L. =Murraya paniculata 

Myllanthus maroana R.S. Cowan =Raputia maroana 

Myllanthus neblinensis R.S. Cowan =Raputia neblinensis 

Neoraputia cowanii Emmerich =Neoraputia paraensis 

Pilocarpus latifolius A. St.-Hil. =Pilocarpus racemosus subsp. racemosus 

Raputia paraensis Ducke =Neoraputia paraensis 

Rauia prancei W.A. Rodrigues & M.F. Silva [not in flora area] 

Rauia racemosa Nees & Mart. [not in flora area] 

Rauia ulei K. Krause =Conchocarpus ucayalinus 

Ravenia linearis Gleason =Raveniopsis linearis 

Ravenia paruana R.S. Cowan =Raveniopsis paruana 

Ravenia ruellioides Oliv. =Raveniopsis ruellioides 

Ravenia ruellioides var. ptariana Steyerm. =Raveniopsis ruellioides 

Ravenia tatei Gleason =Raveniopsis ruellioides 

Raveniopsis liesneri Steyerm. =Raveniopsis stelligera 

Raveniopsis ruellioides var. angusta R.S. Cowan =Raveniopsis ruellioides 

Raveniopsis stelligera var. plicata R.S. Cowan =Raveniopsis stelligera

510 


Schinus fagara L. =Zanthoxylum fagara 

Sciuris aromatica J.F. Gmel. =Raputia aromatica 

Sciuris officinalis (J. Hancock) Oken =Angostura trifoliata 

Spathelia chimantaensis R.S. Cowan & Brizicky =Spathelia ulei 

Spathelia fruticosa (Steyerm.) R.S. Cowan & Brizicky =Spathelia ulei 

Spathelia jauaensis R.S. Cowan =Spathelia ulei 

Spathelia neblinaensis R.S. Cowan & Brizicky =Spathelia ulei 

Zanthoxylum belizense Lundell [specimens re-determined as Zanthoxylum ekmanii] 

Zanthoxylum lucidum A. St.-Hil. [specimens re-determined as Zanthoxylum pentandrum] 

Zanthoxylum microcarpum Griseb. [specimens re-determined as Zanthoxylum rhoifolium] 

Zanthoxylum ocumarense (Pittier) Steyerm. =Zanthoxylum juniperinum 

Zanthoxylum perrottetii DC. =Zanthoxylum rhoifolium 

Zanthoxylum pterota (L.) Kunth =Zanthoxylum fagara 

Zanthoxylum ruizianum (Klotzsch ex Engl.) J.F. Macbr. =Zanthoxylum rhoifolium 

REFERENCES 

Albuquerque, B.W.P. de. 1977 (“1976”). Revisao taxonômica das Rutaceae do Estado do Amazonas. Acta Amazonica suppl. 

6(3): 1-67. 

Albuquerque, B.W.P. de. 1981. Novas combinaçoes em Angosture Roemer & Schultes (Rutaceae). Acta Amazonica 11(4): 849- 

852. 

Cowan, R.S. 1960. Rutaceae. In B. Maguire & J.J. Wurdack, Botany of the Guayana Highland part IV. Mem. New York Bot. 

Gard. 10(2): 24-37. 

Cowan, R.S. 1967. Rutaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 1-14. 

Elias, T.S. 1970. The correct name for the genus Cusparia (Rutaceae). Taxon 19(4): 573-575. 

Emmerich, M. 1978. Contribuçao ao conhecimento da tribo Cusparineae (sic) (Rutaceae). Rodriguesia 45: 223-307. 

Kaastra, R.C. 1982. Pilocarpinae (Rutaceae). Flora Neotropica Monogr. 33: 1-197. 

Kallunki, J.A. 1990. An emended description of and new combinations in Raputia Aublet. Brittonia 42: 175-177. 

Kallunki, J.A. 1992. A revision of Erythrochiton s.l. Brittonia 44(2): 107-139. 

Kallunki, J.A. 1994. Revision of Raputia Aubl. (Cuspariinae, Rutaceae). Brittonia 46(4): 279-295. 

Kallunki, J.A. 2002. Rutaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


656. 

Kallunki, J.A. 2005. Rutaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 1-39. 

Porter, D.M. & T.S. Elias. 1979. Rutaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri 

Bot. Gard. 66: 123-164. 

Waterman, P.G. 1975. New combinations in Zanthoxylum L. (1753). Taxon 24: 361-366. 

SABIACEAE 

G.A. Aymard C., N.L. Cuello A. 

Meliosma gentryi Aymard & Cuello 

Meliosma herbertii Rolfe 

Meliosma herbertii var. herbertii 

Meliosma herbertii var. tepuiensis (Steyerm. & Maguire) Steyerm. & Gentry 

Meliosma aff. meridensis Lasser 

Meliosma vasquezii Gentry 

Ophiocaryon chironectes Barneby 

Ophiocaryon duckei Barneby 

Ophiocaryon heterophyllum (Benth.) Urb. [records from Guyana probably 

represent Ophiocaryon maguirei] 

Ophiocaryon maguirei Barneby 

Ophiocaryon manausense (W.A. Rodrigues) Barneby 

Ophiocaryon paradoxum R.H. Schomb. 

Ophiocaryon sp. A [expected in AM] 


Meliosma nubicola Steyerm. & Lasser =Meliosma meridensis 

Meliosma tepuiensis Steyerm. & Maguire =Meliosma herbertii var. tepuiensis 

Meliosma uberrima Idrobo & Cuatrec. =Meliosma meridensis 

Phoxanthus manausensis W.A. Rodrigues =Ophiocaryon manausense 

AM 

AM BO DA GU 

AM BO DA GU 

BO 

BO 

AM 

REFERENCES 

Aymard C., G.A. and N.L. Cuello A. 2005. Sabiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 

AM 

AM 

GU? 

GU 

FG? 

GU FG?



Barneby, R.C. 1972. Meliosmaceae - Ophiocaryon. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. 


Steyermark, J.A. & A.H. Gentry. 1992. Sabiaceae. In G. Morillo, Flora de Venezuela 5(1): 203-220. 

Thesium tepuiense Steyerm. 

SANTALACEAE 

J.S. Miller 


Austroamericum tepuiense (Steyerm.) Hendrych =Thesium tepuiense 

REFERENCES 

Miller, J.S. 2005. Santalaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 9: 

44-45. 

Steyermark, J. 1951. Fieldiana, Botany 28. 1: 225 

SAPINDACEAE 

P. Acevedo R., H.T. Beck 

Allophylus amazonicus (Mart.) Radlk. 

Allophylus edulis (A. St.-Hil.) Hieron. ex Niederl. 

Allophylus latifolius Huber 

Allophylus leucoclados Radlk. 

Allophylus parimensis Steyerm. 

Allophylus punctatus (Poepp. & Endl.) Radlk. 

Allophylus racemosus Sw. 

Allophylus robustus Radlk. 

Cardiospermum corindum L. 

Cardiospermum grandiflorum Sw. 

Cardiospermum halicacabum L. 

Cardiospermum halicacabum var. halicacabum 

Cardiospermum halicacabum var. microcarpum (Kunth) Blume 

Cupania americana L. [s.l.] 

Cupania cinerea Poepp. & Endl. 

Cupania diphylla Vahl 

Cupania hirsuta Radlk. 

Cupania hispida Radlk. 

Cupania kukenanica Steyerm. 

Cupania rubiginosa (Poir.) Radlk. 

Cupania scrobiculata Rich. 

Cupania scrobiculata var. guianensis (Miq.) Uittien 

Cupania scrobiculata var. scrobiculata 

Dilodendron costaricense (Radlk.) Gentry & Steyerm. 

Dilodendron elegans (Radlk.) Gentry & Steyerm. 

Dodonaea viscosa Jacq. 

Houssayanthus macrolophus (Radlk.) Hunz. 

Matayba adenanthera Radlk. 

Matayba arborescens (Aubl.) Radlk. 

Matayba atropurpurea Radlk. 

Matayba camptoneura Radlk. 

Matayba elegans Radlk. 

Matayba guianensis Aubl. 

Matayba inelegans Spruce ex Radlk. 

Matayba kavanayena (Steyerm.) Steyerm. 

Matayba laevigata (Miq.) Radlk. 

Matayba longipes Radlk. 

Matayba macrostylis Radlk. 

Matayba opaca Radlk. 

Matayba peruviana Radlk. 

Matayba ptariana Steyerm. 

Matayba ptariana subsp. guaiquinimae Steyerm. 

Matayba ptariana subsp. ptariana 

AM 

AM 

BO GU 

GU FG 

FG 

FG 

BO 

BO GU 

GU FG 

BO 

BO 


BO 


BO SU FG 

AM BO 

SU FG 


AM 

BO GU 

AM BO GU 


GU SU FG 

GU FG 

BO 

BO 

BO GU SU FG 

DA 

AM BO 


AM 

AM BO GU SU 

AM BO GU 

AM BO GU FG 


AM BO 

GU SU FG 

AM 

AM GU 


AM BO GU FG 

AM BO GU 

BO 

AM BO

512 


Matayba purgans (Poepp.) Radlk. 

Matayba robusta Radlk. 

Matayba spruceana (Benth.) Radlk. 

Matayba stenodictya Radlk. 

Matayba yutajensis Steyerm. 

Melicoccus bijugatus Jacq. [cultivated or escaped from cultivation] 

Melicoccus oliviformis Kunth 

Melicoccus oliviformis subsp. intermedius (Radlk.) Acev.-Rodr. 

Melicoccus pedicellaris (Sagot ex Radlk.) Acev.-Rodr. 

Nephelium lappaceum L. [cultivated] 

Paullinia acuminata Uittien 

Paullinia alata (Ruiz & Pav.) G. Don 

Paullinia alsmithii Macbr. 

Paullinia anisoptera Turcz. 

Paullinia anodonta Radlk. 

Paullinia anomophylla Radlk. 

Paullinia arenicola Gleason 

Paullinia bernhardi Uittien 

Paullinia bracteosa Radlk. 

Paullinia caloptera Radlk. 

Paullinia cambessedesii Triana & Planch. 

Paullinia capreolata (Aubl.) Radlk. 

Paullinia clathrata Radlk. 

Paullinia conduplicata Radlk. 

Paullinia cupana Kunth 

Paullinia cururu L. 

Paullinia curvicuspis Radlk. 

Paullinia dasygonia Radlk. 

Paullinia demerarae Sandwith 

Paullinia fibulata Rich. ex Juss. 

Paullinia fuscescens Kunth 

Paullinia grandifolia Benth. ex Radlk. 

Paullinia hitchcockii Gleason 

Paullinia imberbis Radlk. 

Paullinia ingifolia Rich. ex Juss. 

Paullinia isoptera Radlk. 

Paullinia latifolia Benth. ex Radlk. 

Paullinia leiocarpa Griseb. 

Paullinia lingulata Acev.-Rodr. 

Paullinia livescens Radlk. 

Paullinia micrantha Cambess. 

Paullinia novemalata Uittien 

Paullinia nuriensis Steyerm. 

Paullinia pachycarpa Benth. 

Paullinia paullinioides Spruce ex Radlk. 

Paullinia perlata Radlk. 

Paullinia pinnata L. 

Paullinia plagioptera Radlk. 

Paullinia rubiginosa Cambess. 

Paullinia rufescens Rich. ex Juss. 

Paullinia rugosa Benth. ex Radlk. 

Paullinia serjaniifolia Triana & Planch. 

Paullinia sphaerocarpa Rich. ex Juss. 

Paullinia spicata Benth. 

Paullinia stellata Radlk. 

Paullinia stenopetala Sagot 

Paullinia subnuda Radlk. 

Paullinia tetragona Aubl. 

Paullinia triantennata F. Silveira 

Paullinia tricornis Radlk. 

Paullinia trilatera Radlk. 

Paullinia venosa Radlk. 

Paullinia verrucosa Radlk. 

Paullinia vespertilio Sw. 

GU FG 

AM 

BO 

FG 

AM 

BO GU SU FG 

AM GU 

AM GU 

SU FG 

FG 

SU FG 

SU FG 

AM BO 

BO DA GU FG 

SU FG 

FG 

GU 

SU 

AM GU 

AM BO GU FG 

FG 


FG 

AM 

BO 

GU FG 

FG 

GU 

GU 

SU FG 

FG 


GU 

GU 

SU 

AM FG 

AM BO GU 

GU 

SU FG 

GU SU FG? 

AM BO GU 

GU 

FG 

BO 

BO 

DA GU SU 

SU 

FG 

AM 

GU 

GU 

FG 


GU SU FG 

AM BO FG 


AM 

AM 

GU FG 

GU? SU FG 

SU FG 

GU 

GU 

SU 

FG 

FG 

GU SU 

GU SU FG 

SU FG 

SU FG 

BO GU 

GU? 

SU FG

Paullinia xestophylla Radlk. 

Pentascyphus thyrsiflorus Radlk. 

Porocystis toulicioides Radlk. 

Pseudima frutescens (Aubl.) Radlk. 

Pseudima pallidum Radlk. 

Sapindus saponaria L. 

Sapindus saponaria var. inaequalis (DC.) Radlk. 

Scyphonichium multiflorum (Mart.) Radlk. 

Serjania adusta Radlk. 

Serjania caracasana (Jacq.) Willd. 

Serjania chartacea Radlk. 

Serjania clematidea Triana & Planch. 

Serjania communis Cambess. 

Serjania communis var. communis 

Serjania didymadenia Radlk. 

Serjania exarata Radlk. 

Serjania grandifolia Sagot ex Radlk. 

Serjania membranacea Splitg. 

Serjania oblongifolia Radlk. 

Serjania paucidentata DC. 

Serjania pedicellaris Radlk. 

Serjania pyramidata Radlk. 

Serjania rhombea Radlk. 

Serjania setulosa Radlk. 

Talisia acutifolia Radlk. 

Talisia carinata Radlk. 

Talisia caudata Steyerm. 

Talisia chartacea Acev.-Rodr. 

Talisia clathrata Radlk. 

Talisia clathrata subsp. canescens Acev.-Rodr. 

Talisia cupularis Radlk. 

Talisia eximia K.U. Kramer 

Talisia firma Radlk. 

Talisia furfuracea Sandwith 

Talisia guianensis Aubl. 

Talisia hemidasya Radlk. 

Talisia hexaphylla Vahl 

Talisia laevigata Acev.-Rodr. 

Talisia longifolia (Benth.) Radlk. 

Talisia macrophylla (Mart.) Radlk. 

Talisia megaphylla Sagot ex Radlk. 

Talisia microphylla Uittien 

Talisia mollis Kunth ex Cambess. 

Talisia nervosa Radlk. 

Talisia obovata A.C. Sm. 

Talisia pachycarpa Radlk. 

Talisia pilosula Sagot ex Radlk. 

Talisia praealta (Sagot) Radlk. 

Talisia retusa R.S. Cowan 

Talisia simaboides K.U. Kramer 

Talisia squarrosa Radlk. 

Talisia sylvatica (Aubl.) Radlk. 

Thinouia myriantha Triana & Planch. 

Toulicia elliptica Radlk. 

Toulicia guianensis Aubl. 

Toulicia patentinervis Radlk. 

Toulicia petiolulata Radlk. 

Toulicia pulvinata Radlk. 

Urvillea ulmacea Kunth 

Vouarana anomala (Steyerm.) Acev.-Rodr. 

Vouarana guianensis Aubl. 



Allophylus occidentalis (Sw.) Radlk. =Allophylus racemosus 

GU 

GU 

FG 

FG 

GU SU FG 

SU FG 

AM BO DA GU? SU FG? 

GU? SU FG? 

FG 

BO GU 

BO GU FG 

GU FG 

AM 

AM 

AM 

BO 

GU 

FG 

BO GU SU FG 


GU SU FG 


SU FG 

AM 

AM 

BO DA GU FG 

FG 

AM 

GU SU FG 

AM 

AM 

BO 

SU 

FG 

FG 

FG 

GU SU FG 

GU FG 


BO 

BO 

DA GU FG 

GU SU FG 


GU SU FG 

GU SU FG 

GU SU FG 

AM 

AM 

BO 

FG 

FG 

SU FG 

AM BO GU SU 

SU FG 

GU FG 

GU SU FG 

DA GU SU FG 

GU? SU FG? 


AM 

GU SU 


AM BO 

BO 

GU SU? FG 

GU SU FG

514 


Cardiospermum macrolophum Radlk. =Houssayanthus macrolophus 

Cardiospermum microcarpum Kunth =Cardiospermum halicacabum var. microcarpum 

Cupania kavanayena Steyerm. =Matayba kavanayena 

Cupania lachnocarpa Radlk. [not in flora area] 

Cupania lanuginosa Sagot ex Radlk. =Cupania hirsuta 

Cupania latifolia Kunth =Cupania americana s.l. 

Cupania macrophylla Mart. =Talisia macrophylla 

Cupania papillosa Radlk. =Cupania latifolia 

Cupania roraimae Steyerm. =Cupania kukenanica 

Cupania scrobiculata f. reticulata (Cambess.) Radlk. =Cupania scrobiculata var. scrobiculata 

Cupania semidentata Radlk. =Cupania latifolia 

Cupania spruceana Benth. =Matayba spruceana 

Cupania tepuiensis Steyerm. & Maguire =Cupania rubiginosa 

Dilodendron bipinnatum var. elegans Radlk. =Dilodendron elegans 

Dipterodendron costaricense Radlk. =Dilodendron costaricense 

Dipterodendron elegans (Radlk.) Radlk. =Dilodendron elegans 

Dipterodendron venezuelense Steyerm. =Dilodendron elegans 

Enourea capreolata Aubl. =Paullinia capreolata 

Matayba affinis Steyerm. =Matayba opaca var. opaca 

Matayba chimantensis Steyerm. & Maguire =Matayba peruviana 

Matayba discolor var. atropurpurea Radlk. =Matayba atropurpurea 

Matayba fallax Radlk. =Matayba opaca var. fallax 

Matayba jauaensis Steyerm. =Matayba peruviana 

Matayba longipes subsp. tepuiensis Steyerm. =Matayba longipes 

Matayba macrolepis Radlk. =Matayba macrostylis 

Matayba miquelii Uittien =Matayba laevigata 

Matayba oligandra Sandwith =Matayba peruviana 

Matayba oligandra var. occidentalis Steyerm. =Matayba peruviana 

Matayba oligandra var. ptariana (Steyerm.) Steyerm. =Matayba peruviana 

Matayba opaca var. fallax (Radlk.) Uittien =Matayba opaca 

Matayba reducta Steyerm. =Matayba camptoneura 

Matayba sororopaniana Steyerm. =Matayba inelegans 

Matayba tovarensis Radlk. =Matayba longipes 

Matayba venezuelana Steyerm. =Matayba longipes 

Melicoccus bijuga (Jacq.) L. =Melicoccus bijugatus 

Melicoccus bijugata L. =Melicoccus bijugatus 

Paullinia caracasana Jacq. =Serjania caracasana 

Paullinia fasciculata Radlk. =Paullinia alata 

Paullinia sorbilis Mart. =Paullinia cupana 

Racaria sylvatica Aubl. =Talisia sylvatica 

Robinia rubiginosa Poir. =Cupania rubiginosa 

Sapindus arborescens Aubl. =Matayba arborescens 

Sapindus frutescens Aubl. =Pseudima frutescens 

Schmidelia amazonica Mart. =Allophylus amazonicus 

Schmidelia occidentalis Sw. =Allophylus racemosus 

Schmidelia punctata Poepp. & Endl. =Allophylus punctatus 

Serjania ierensis Britton =Serjania caracasana 

Serjania mariquitensis Triana & Planch. =Serjania caracasana 

Serjania prancei Acev.-Rodr. [nomen nudum] =Serjania caracasana 

Talisia allenii Croat =Talisia macrophylla 

Talisia amaruayana Steyerm. =Talisia nervosa 

Talisia cararensis Cuatrec. =Talisia hexaphylla subsp. hexaphylla 

Talisia dwyeri Croat =Talisia nervosa 

Talisia elephantipes Sandwith =Talisia macrophylla 

Talisia glandulifera Steyerm. =Talisia hemidasya 

Talisia grandifolia Cuatrec. =Talisia nervosa 

Talisia heterodoxa Steyerm. =Talisia retusa 

Talisia medrii Guarim =Talisia acutifolia 

Talisia micrantha Radlk. =Talisia sylvatica 

Talisia olivaeformis (Kunth) Radlk. =Melicoccus oliviformis 

Talisia oliviformis (Kunth) Radlk. =Melicoccus oliviformis 

Talisia panamensis Pittier =Talisia hexaphylla subsp. hexaphylla 

Talisia pedicellaris Sagot ex Radlk. =Melicoccus pedicellaris 

Talisia pentantha Steyerm. =Talisia macrophylla


Talisia pulverulenta Radlk. =Melicoccus pedicellaris 

Talisia reticulata Radlk. =Talisia sylvatica 

Talisia sancarlosiana Steyerm. =Talisia macrophylla 

Talisia tirirensis Steyerm. & Maguire =Talisia nervosa 

Tapirocarpus talisia Sagot =Talisia mollis 

Toulicia anomala Steyerm. =Vouarana anomala 

Trichilia caudata Killip & Cuatrec. =Matayba macrostylis 

Trichilia ptariana Steyerm. =Matayba peruviana 

Urvillea macrolopha (Radlk.) Radlk. =Houssayanthus macrolophus 

REFERENCES 

Acevedo-R., P. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. 

Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 656-669. 

Acevedo-R., P. & H.T. Beck. 2005. Sapindaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Kramer, K.U. 1973 (1972). Studies in the New World Sapindaceae. I. Notes on the genus Talisia Aublet. Acta Bot. Neerl. 21: 

671-678. 

Kramer, K.U. 1976. Sapindaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 487-511. 

Radlkofer, L. 1931-1934. Sapindaceae. In A. Engler, Das Pflanzenreich IV,165 Bd. I: 1-1018, Bd. II: 1019-1539. 

Uittien, H. 1937. Sapindaceae. In A. Pulle, Flora of Suriname 2(1): 345-396. 

SAPOTACEAE 

K.S. Edwards, T.D. Pennington 

Chromolucuma baehniana Monach. 

Chromolucuma rubriflora Ducke 

Chrysophyllum amazonicum T.D. Penn. 

Chrysophyllum argenteum Jacq. 

Chrysophyllum argenteum subsp. auratum (Miq.) T.D. Penn. 

Chrysophyllum argenteum subsp. nitidum (G. Mey.) T.D. Penn. 

Chrysophyllum cainito L. [cultivated, naturalized] 

Chrysophyllum cuneifolium (Rudge) A. DC. 

Chrysophyllum eximium Ducke [expected in FG] 

Chrysophyllum lucentifolium Cronquist 

Chrysophyllum lucentifolium subsp. pachycarpum Pires & T.D. Penn. 

Chrysophyllum manaosense (Aubrév.) T.D. Penn. 

Chrysophyllum pomiferum (Eyma) T.D. Penn. 

Chrysophyllum prieurii A. DC. 

Chrysophyllum sanguinolentum (Pierre) Baehni 

Chrysophyllum sanguinolentum subsp. balata (Ducke) T.D. Penn. 

Chrysophyllum sanguinolentum subsp. sanguinolentum 

Chrysophyllum sanguinolentum subsp. spurium (Ducke) T.D. Penn. 

Chrysophyllum sparsiflorum Klotzsch ex Miq. 

Chrysophyllum ucuquirana-branca (Aubrév. & Pellegr.) T.D. Penn. 

Chrysophyllum venezuelanense (Pierre) T.D. Penn. 

Diploon cuspidatum (Hoehne) Cronquist 

Ecclinusa atabapoensis (Aubrév.) T.D. Penn. 

Ecclinusa bullata T.D. Penn. 

Ecclinusa dumetorum (Baehni) T.D. Penn. 

Ecclinusa guianensis Eyma 

Ecclinusa lanceolata (Mart. & Eichler) Pierre 

Ecclinusa orinocoensis Aubrév. 

Ecclinusa parviflora T.D. Penn. 

Ecclinusa psilophylla Sandwith 

Ecclinusa ramiflora Mart. [expected in GU] 

Ecclinusa ulei (K. Krause) Gilly ex Cronquist [Mt. Roraima] 

Elaeoluma crispa T.D. Penn. 

Elaeoluma glabrescens (Mart. & Eichler) Aubrév. 

Elaeoluma nuda (Baehni) Aubrév. [expected in GU] 

Elaeoluma schomburgkiana (Miq.) Baill. 

Manilkara bidentata (A. DC.) A. Chev. 

Manilkara bidentata subsp. bidentata 

Manilkara bidentata subsp. surinamensis (Miq.) T.D. Penn. 

Manilkara bolivarensis T.D. Penn. 

GU 

AM 

BO GU 



GU SU FG 

GU SU FG 

SU 

SU 

FG 

AM BO GU FG 

AM BO GU FG 

GU SU 


AM GU SU FG 


AM BO GU 

AM 

AM 

GU SU FG 

AM 

BO GU SU? FG 

FG 

AM 

AM 

DA GU FG 

SU FG? 


AM 

AM 

AM 

GU FG 

GU SU 

AM 

AM 

BO 

SU FG 

AM BO GU 

AM BO SU 

AM BO GU 



AM BO 

BO 

DA GU SU FG

516 


Manilkara excelsa (Ducke) Standl. 

Manilkara huberi (Ducke) A. Chev. 

Manilkara paraensis (Huber) Standl. 

Manilkara pubicarpa Monach. 

Manilkara zapota (L.) P. Royen [widely cultivated and escaping] 

Micropholis acutangula (Ducke) Eyma 

Micropholis casiquiarensis Aubrév. 

Micropholis caudata T.D. Penn. 

Micropholis cayennensis T.D. Penn. 

Micropholis egensis (A. DC.) Pierre 

Micropholis gardneriana (A. DC.) Pierre 

Micropholis guyanensis (A. DC.) Pierre 

Micropholis guyanensis subsp. duckeana (Baehni) T.D. Penn. 

Micropholis guyanensis subsp. guyanensis 

Micropholis humboldtiana (Roem. & Schult.) T.D. Penn. 

Micropholis longipedicellata Aubrév. 

Micropholis maguirei Aubrév. 

Micropholis melinoniana Pierre 

Micropholis mensalis (Baehni) Aubrév. 

Micropholis obscura T.D. Penn. 

Micropholis porphyrocarpa (Baehni) Monach. 

Micropholis sanctae-rosae (Baehni) T.D. Penn. 

Micropholis spectabilis (Steyerm.) T.D. Penn. 

Micropholis splendens Gilly ex Aubrév. 

Micropholis submarginalis Pires & T.D. Penn. 

Micropholis suborbicularis Aubrév. 

Micropholis venamoensis (Steyerm.) T.D. Penn. 

Micropholis venulosa (Mart. & Eichler) Pierre 

Mimusops spp. [cultivated or escaped from cultivation] 

Podoluma pacimoniensis Aubrév. 

Pouteria ambelaniifolia (Sandwith) T.D. Penn. 

Pouteria amygdalicarpa (Pittier) T.D. Penn. 

Pouteria anomala (Pires) T.D. Penn. 

Pouteria arcuata T.D. Penn. 

Pouteria atabapoensis (Aubrév.) T.D. Penn. 

Pouteria aubrevillei Bernardi 

Pouteria bangii (Rusby) T.D. Penn. [expected in GU] 

Pouteria benai (Aubrév. & Pellegr.) T.D. Penn. 

Pouteria bilocularis (H. Winkl.) Baehni 

Pouteria brachyandra (Aubrév. & Pellegr.) T.D. Penn. 

Pouteria caimito (Ruiz & Pav.) Radlk. 

Pouteria campanulata Baehni [possibly expected in GU] 

Pouteria canaimaensis T.D. Penn. 

Pouteria cayennensis (A. DC.) Eyma 

Pouteria cicatricata T.D. Penn. 

Pouteria cladantha Sandwith 

Pouteria congestifolia Pilz 

Pouteria coriacea (Pierre) Pierre 

Pouteria crassiflora Pires & T.D. Penn. [possibly expected in FG] 

Pouteria cuspidata (A. DC.) Baehni 

Pouteria cuspidata subsp. cuspidata 

Pouteria cuspidata subsp. dura (Eyma) T.D. Penn. 

Pouteria cuspidata subsp. robusta (Mart. & Eichler) T.D. Penn. 

Pouteria decorticans T.D. Penn. 

Pouteria deliciosa T.D. Penn. 

Pouteria durlandii (Standl.) Baehni 

Pouteria durlandii subsp. durlandii 

Pouteria egregia Sandwith 

Pouteria elegans (A. DC.) Baehni 

Pouteria engleri Eyma 

Pouteria eugeniifolia (Pierre) Baehni 

Pouteria filipes Eyma 

Pouteria fimbriata Baehni 

Pouteria franciscana Baehni 

GU? 

AM GU SU FG 

SU FG 

GU 

FG 

AM GU 

GU 

AM BO GU? 

GU 

SU 

FG 

FG 


AM GU SU FG 

AM 

AM 

BO GU SU FG 

AM 

GU SU FG 

AM BO SU FG 

AM GU SU FG 

AM GU FG 

BO 

GU FG 

FG 

AM FG 

FG 

AM BO 

BO 


AM 

AM 

AM 

AM 


BO 

FG 


FG 

AM BO GU FG 

GU SU FG 


BO 

AM BO GU FG 

FG 


FG 

AM GU SU FG 

AM 




AM BO SU FG 

SU FG 

FG 

GU 

GU 


AM GU 

BO GU SU FG 


GU SU FG 

AM BO GU FG 

FG


Pouteria gabrielensis (Gilly ex Aubrév.) T.D. Penn. 

Pouteria glomerata (Miq.) Radlk. 

Pouteria glomerata subsp. glomerata 

Pouteria glomerata subsp. stylosa (Pierre) T.D. Penn. 

Pouteria gomphiifolia (Mart. ex Miq.) Radlk. 

Pouteria gongrijpii Eyma 

Pouteria grandis Eyma 

Pouteria guianensis Aubl. 

Pouteria hispida Eyma 

Pouteria jariensis Pires & T.D. Penn. 

Pouteria kaieteurensis T.D. Penn. 

Pouteria laevigata (Mart.) Radlk. 

Pouteria latianthera T.D. Penn. [expected in FG] 

Pouteria lucumifolia (Reissek ex Maxim.) T.D. Penn. 

Pouteria macrocarpa (Mart.) Dietr. 

Pouteria macrophylla (Lam.) Eyma 

Pouteria maguirei (Aubrév.) T.D. Penn. 

Pouteria manaosensis (Aubrév. & Pellegr.) T.D. Penn. 

Pouteria melanopoda Eyma 

Pouteria aff. minima T.D. Penn. 

Pouteria oblanceolata Pires 

Pouteria orinocoensis (Aubrév.) T.D. Penn. 

Pouteria penicillata (Baehni) Aubrév. 

Pouteria pimichinensis T.D. Penn. 

Pouteria plicata T.D. Penn. 

Pouteria procera (Mart.) T.D. Penn. 

Pouteria pullei Eyma [imperfectly known species] 

Pouteria putamen-ovi T.D. Penn. 

Pouteria reticulata (Engl.) Eyma 

Pouteria reticulata subsp. reticulata [expected in GU] 

Pouteria reticulata subsp. surinamensis T.D. Penn. 

Pouteria retinervis T.D. Penn. 

Pouteria rigida (Mart. & Eichler) Radlk. 

Pouteria rigida subsp. rigida 

Pouteria rigida subsp. tomentosa (Aubrév.) T.D. Penn. 

Pouteria rigidopsis (Monach.) T.D. Penn. 

Pouteria rodriguesiana Pires & T.D. Penn. 

Pouteria rostrata (Huber) Baehni 

Pouteria sagotiana (Baill.) Eyma 

Pouteria sapota (Jacq.) H.E. Moore & Stearn [widely cultivated] 

Pouteria scrobiculata Monach. ex T.D. Penn. 

Pouteria singularis T.D. Penn. 

Pouteria sipapoensis T.D. Penn. 

Pouteria speciosa (Ducke) Baehni 

Pouteria stipitata Cronquist 

Pouteria surumuensis Baehni 

Pouteria tenuisepala Pires & T.D. Penn. 

Pouteria torta (Mart.) Radlk. 

Pouteria torta subsp. glabra T.D. Penn. 

Pouteria torta subsp. tuberculata (Sleumer) T.D. Penn. 

Pouteria trigonosperma Eyma 

Pouteria trilocularis Cronquist 

Pouteria ucuqui Pires & R.E. Schult. 

Pouteria venosa (Mart.) Baehni 

Pouteria venosa subsp. amazonica T.D. Penn. 

Pouteria virescens Baehni 

Pouteria williamii (Aubrév. & Pellegr.) T.D. Penn. 

Pradosia beardii (Monach.) T.D. Penn. 

Pradosia caracasana (Pittier) T.D. Penn. 

Pradosia cochlearia (Lecomte) T.D. Penn. 

Pradosia cochlearia subsp. cochlearia 

Pradosia grisebachii (Pierre) T.D. Penn. 

Pradosia huberi (Ducke) Ducke 

Pradosia ptychandra (Eyma) T.D. Penn. 

AM 


AM BO FG 

BO SU 

AM 

SU FG 

GU SU FG 



SU? FG 

GU 

AM FG 

AM 

GU? SU 

FG? 

FG 

AM 

GU 

SU 

SU FG 

BO 

GU FG 

AM BO 

GU 

AM 

BO DA GU 

FG 

SU 


FG 

FG 


GU SU 

FG 

AM BO GU 

AM BO GU 

AM BO GU 

GU? 

GU? 

SU FG 

GU SU FG 

AM BO GU 

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AM 

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BO 

AM GU 

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AM BO GU FG 

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GU SU 

AM 

AM 



GU FG 

FG 

AM BO 

BO 

GU 

FG 

FG 

BO DA 

FG 

GU SU FG

518 

Pradosia schomburgkiana (A. DC.) Cronquist 

Pradosia schomburgkiana subsp. schomburgkiana 

Pradosia schomburgkiana subsp. sericea T.D. Penn. 

Pradosia surinamensis (Eyma) T.D. Penn. 

Pradosia verticillata Ducke 

Sarcaulus brasiliensis (A. DC.) Eyma 

Sarcaulus brasiliensis subsp. brasiliensis 


AM BO GU SU 

AM BO GU SU 

AM 

BO GU SU FG 

FG 

GU SU FG 

GU SU FG 


Achras balata Aubl. [described from cultivated material] =Mimusops sp. 

Achras caimito Ruiz & Pav. =Pouteria caimito 

Achras venosa Mart. =Pouteria venosa 

Achras zapota L. =Manilkara zapota 

Achrouteria pomifera Eyma =Chrysophyllum pomiferum 

Amorphospermum schomburgkianum (Miq.) Baehni =Elaeoluma schomburgkiana 

Bumelia amazonica K. Krause =Pouteria cuspidata subsp. cuspidata 

Bumelia cuneifolia Rudge =Chrysophyllum cuneifolium 

Bumelia micrantha (Kunth) Willd. (Myrsinaceae) =Stylogyne micrantha 

Bumelia nervosa Vahl =Pouteria macrophylla 

Calocarpum cooperi (Cronquist) Lundell =Pouteria torta subsp. tuberculata 

Calocarpum mammosum (L.) Pierre =Pouteria sapota 

Chrysophyllum alnifolium Engl., non Baker =Pouteria engleri 

Chrysophyllum ambelaniifolium Sandwith =Pouteria ambelaniifolia 

Chrysophyllum anomalum Pires =Pouteria anomala 

Chrysophyllum aquaticum Kunth =Micropholis humboldtiana 

Chrysophyllum auratum Miq. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum auratum var. majus Miq. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum beardii Monach. =Pradosia beardii 

Chrysophyllum brasiliense A. DC. =Sarcaulus brasiliensis 

Chrysophyllum brasiliense var. minus Miq. =Sarcaulus brasiliensis subsp. brasiliensis 

Chrysophyllum cayennense A. DC. =Pouteria cayennensis 

Chrysophyllum cochlearium Lecomte =Pradosia cochlearia 

Chrysophyllum cuspidatum Hoehne =Diploon cuspidatum 

Chrysophyllum dumetorum Baehni =Ecclinusa dumetorum 

Chrysophyllum emarginatum Klotzsch ex M.R. Schomb. [name of uncertain application] 

Chrysophyllum ficoides Steyerm. =Ecclinusa ulei 

Chrysophyllum guianense (Eyma) Baehni =Ecclinusa guianensis 

Chrysophyllum guyanense Klotzsch ex Miq. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum humbodltianum Roem. & Schult. =Micropholis humboldtiana 

Chrysophyllum macoucou Aubl. [incertae sedis; excluded from Sapotaceae by Pennington] =Aubletella macoucou (Aubl.) 

Pierre 

Chrysophyllum macrophyllum Mart., non Lam. =Sarcaulus brasiliensis subsp. brasiliensis 

Chrysophyllum macrophyllum Lam. =Pouteria macrophylla 

Chrysophyllum melinonii Engl. =Micropholis guyanensis subsp. guyanensis 

Chrysophyllum monachinoanum Steyerm. =Pradosia beardii 

Chrysophyllum nitidum G. Mey. =Chrysophyllum argenteum subsp. nitidum 

Chrysophyllum obtusatum Willd. ex Roem. & Schult. =Micropholis humboldtiana 

Chrysophyllum psilophyllum A. DC. =Chrysophyllum argenteum subsp. nitidum 

Chrysophyllum psilophyllum (Sandwith) Baehni =Ecclinusa psilophylla 

Chrysophyllum pyriforme Willd. [incertae sedis; excluded from Sapotaceae by Pennington] =Aubletella macoucou (Aubl.) 

Pierre 

Chrysophyllum reticulatum Engl. =Pouteria reticulata 

Chrysophyllum rufocupreum Ducke =Chrysophyllum eximium 

Chrysophyllum schomburgkianum A. DC. =Pradosia schomburgkiana 

Chrysophyllum sericeum A. DC. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum sericeum var. acutifolium Miq. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum sericeum var. obtusifolium Miq. =Chrysophyllum argenteum subsp. auratum 

Chrysophyllum sessiliflorum Poir. =Pouteria cayennensis 

Chrysophyllum steyermarkii Monach. =Chrysophyllum sparsiflorum 

Chrysophyllum ulei K. Krause =Ecclinusa ulei 

Cynodendron nitidum (G. Mey.) Aubrév. =Chrysophyllum argenteum subsp. nitidum 

Diploon venezuelana Aubrév. =Diploon cuspidatum 

Ecclinusa balata Ducke =Chrysophyllum sanguinolentum subsp. balata 

Ecclinusa cochlearia (Lecomte) Aubrév. =Pradosia cochlearia


Ecclinusa cuneifolia (Rudge) Aubrév. =Chrysophyllum cuneifolium 

Ecclinusa grisebachii Pierre =Pradosia grisebachii 

Ecclinusa prieurii (A. DC.) Aubrév. =Chrysophyllum prieurii 

Ecclinusa sanguinolenta (Pierre) Engl. =Chrysophyllum sanguinolentum 

Ecclinusa spuria Ducke =Chrysophyllum sanguinolentum subsp. spurium 

Ecclinusa ucuquirana-branca Aubrév. & Pellegr. =Chrysophyllum ucuquirana-branca 

Englerella macrocarpa Pierre =Pouteria speciosa 

Englerella speciosa Ducke =Pouteria speciosa 

Eremoluma sagotiana Baill. =Pouteria sagotiana 

Eremoluma williamii Aubrév. & Pellegr. =Pouteria williamii 

Franchetella atabapoensis Aubrév. =Pouteria atabapoensis 

Franchetella gonggrijpii (Eyma) Aubrév. =Pouteria gongrijpii 

Franchetella neblinaensis Aubrév. =Pouteria trilocularis 

Glycoxylon maguirei Aubrév. =Pradosia schomburgkiana subsp. schomburgkiana 

Glycoxylon rubrum Aubrév. =Pradosia beardii 

Gomphiluma martiana Baill. =Pouteria gomphiifolia 

Guapeba coriacea Pierre (Tectariaceae) =Pouteria coriacea 

Guapeba stylosa Pierre =Pouteria glomerata subsp. stylosa 

Ichthyophora elegans (Miq.) Baehni =Pouteria elegans 

Ichthyophora leptocarpa (Miq.) Baehni =Pouteria elegans 

Labatia ? bilocularis H. Winkl. =Pouteria bilocularis 

Labatia laevigata Mart. =Pouteria laevigata 

Labatia parviflora Pittier =Pouteria glomerata subsp. glomerata 

Labatia pedunculata Willd. =Pouteria guianensis 

Labatia torta Mart. =Pouteria torta 

Labatia tovarensis Engl. =Pouteria guianensis 

Lucuma duckei Huber =Pouteria venosa subsp. amazonica 

Lucuma glabrescens Mart. & Eichler =Elaeoluma glabrescens 

Lucuma glomerata Miq. =Pouteria glomerata 

Lucuma gomphiifolia Mart. ex Miq. =Pouteria gomphiifolia 

Lucuma gomphiifolia var. blepharantha Mart. =Pouteria gomphiifolia 

Lucuma jenmanii Pittier =Pouteria torta subsp. glabra 

Lucuma procera Mart. =Pouteria procera 

Lucuma pulverulenta Mart. & Eichler =Pouteria cayennensis 

Lucuma rigida Mart. & Eichler =Pouteria rigida 

Lucuma rostrata Huber =Pouteria rostrata 

Lucuma sagotiana (Baill.) Engl. =Pouteria sagotiana 

Lucuma sericea K. Krause =Pouteria surumuensis 

Lucuma speciosa Ducke =Pouteria speciosa 

Lucuma temare Kunth =Pouteria caimito 

Manilkara amazonica (Huber) A. Chev. =Manilkara bidentata subsp. surinamensis 

Manilkara balata (Pierre) Dubard =Manilkara bidentata subsp. bidentata 

Manilkara surinamensis (Miq.) Dubard =Manilkara bidentata subsp. surinamensis 

Manilkara williamsii Standl. =Manilkara bidentata subsp. bidentata 

Meioluma guianensis Baill. =Micropholis venulosa 

Micropholis burchelliana Pierre =Micropholis gardneriana 

Micropholis calophylloides Pierre =Micropholis venulosa 

Micropholis cowanii Aubrév. =Micropholis guyanensis subsp. guyanensis 

Micropholis eugeniifolia Pierre =Pouteria eugeniifolia 

Micropholis melinonii (Engl.) Mennega et al. =Micropholis guyanensis subsp. guyanensis 

Micropholis mucronata Pierre =Micropholis venulosa 

Micropholis paraensis Huber ex Eyma =Micropholis acutangula 

Micropholis robusta (Mart. & Eichler) Mennega et al. =Pouteria cuspidata subsp. robusta 

Micropholis steyermarkii Monach. =Pouteria eugeniifolia 

Micropholis wurdackii Aubrév. =Micropholis egensis 

Mimusops balata (Aubl.) C.F. Gaertn. [see Achras balata] 

Mimusops balata var. gutta Pierre =Manilkara bidentata subsp. bidentata 

Mimusops balata var. melinonis Pierre =Manilkara bidentata subsp. bidentata 

Mimusops balata var. schomburgkii Pierre =Manilkara bidentata subsp. bidentata 

Mimusops bidentata A. DC. =Manilkara bidentata 

Mimusops huberi Ducke =Manilkara huberi 

Mimusops paraensis Huber =Manilkara paraensis 

Mimusops surinamensis Miq. =Manilkara bidentata subsp. surinamensis 

Myrtiluma eugeniifolia (Pierre) Aubrév. =Pouteria eugeniifolia

520 


Nemaluma engleri (Eyma) Aubrév. & Pellegr. =Pouteria engleri 

Neopometia ptychandra (Eyma) Aubrév. =Pradosia ptychandra 

Neoxythece atabapoensis Aubrév. =Ecclinusa atabapoensis 

Neoxythece cladantha (Sandwith) Aubrév. =Pouteria cladantha 

Neoxythece dura (Eyma) Aubrév. & Pellegr. =Pouteria cuspidata subsp. dura 

Neoxythece gabrielensis Gilly ex Aubrév. =Pouteria gabrielensis 

Neoxythece maguirei Aubrév. =Pouteria pimichinensis 

Neoxythece robusta (Mart. & Eichler) Aubrév. & Pellegr. =Pouteria cuspidata subsp. robusta 

Neoxythece roraimaensis Aubrév. =Pouteria rigida subsp. rigida 

Neoxythece schulzii Aubrév. =Pouteria cuspidata subsp. dura 

Neoxythece tomentosa Aubrév. =Pouteria rigida subsp. tomentosa 

Oxythece ambelaniifolia (Sandwith) Cronquist =Pouteria ambelaniifolia 

Oxythece caracasana Pittier =Pradosia caracasana 

Oxythece dura (Eyma) Gilly =Pouteria cuspidata subsp. dura 

Oxythece elegans (A. DC.) Urb. =Pouteria elegans 

Oxythece guianensis Lecomte ? =Pouteria guianensis 

Oxythece leptocarpa Miq. =Pouteria elegans 

Oxythece rigida (Mart. & Eichler) Monach. =Pouteria rigida 

Oxythece robusta var. brevipetiolasta Monach. =Pouteria cuspidata subsp. robusta 

Oxythece schomburgkiana (Miq.) Cronquist =Elaeoluma schomburgkiana 

Oxythece scrobiculata Monach. [nom. inval.] =Pouteria scrobiculata 

Oxythece steyermarkiana Monach. =Elaeoluma schomburgkiana 

Paralabatia acutangula (Ducke) Baehni =Micropholis acutangula 

Paralabatia casiquiarensis Aubrév. =Pouteria lucumifolia 

Paralabatia orinocoensis Aubrév. =Pouteria orinocoensis 

Paramicropholis acutangula (Ducke) Aubrév. & Pellegr. =Micropholis acutangula 

Passaveria lanceolata Mart. & Eichler =Ecclinusa lanceolata 

Planchonella elegans (A. DC.) Baehni =Pouteria elegans 

Planchonella guianensis P. Royen =Chrysophyllum pomiferum 

Planchonella melinonii (Engl.) Baehni =Micropholis guyanensis subsp. guyanensis 

Platyluma calophylloides (Pierre) Baill. =Micropholis venulosa 

Podoluma benai Aubrév. & Pellegr. =Pouteria benai 

Pouteria acutangula (Ducke) Baehni =Micropholis acutangula 

Pouteria auyantepuiensis Steyerm. =Elaeoluma nuda 

Pouteria benoistii Aubrév. =Pouteria hispida 

Pouteria burchelliana (Pierre) Baehni =Micropholis gardneriana 

Pouteria caimito var. strigosa Monach. =Pouteria guianensis 

Pouteria demerarae Sandwith =Pouteria guianensis 

Pouteria duckeana Baehni =Micropholis guyanensis subsp. duckeana 

Pouteria dura Eyma =Pouteria cuspidata subsp. dura 

Pouteria echinocarpa W.A. Rodrigues =Pouteria torta subsp. tuberculata 

Pouteria elegans var. micrantha (Miq.) Baehni =Pouteria cuspidata subsp. cuspidata 

Pouteria jenmanii (Pittier) Sandwith =Pouteria torta subsp. glabra 

Pouteria lucentifolia (Standl.) Baehni =Chrysophyllum venezuelanense 

Pouteria melinoniana (Pierre) Baehni =Micropholis melinoniana 

Pouteria melinonii (Engl.) Baehni =Micropholis guyanensis subsp. guyanensis 

Pouteria mensalis Baehni =Micropholis mensalis 

Pouteria minutiflora (Britton) Sandwith =Pouteria coriacea 

Pouteria multiflora (A. DC.) Eyma [not in flora area] 

Pouteria nuda Baehni =Elaeoluma nuda 

Pouteria obidensis Huber =Pouteria guianensis 

Pouteria pedunculata (Willd.) Poir. =Pouteria guianensis 

Pouteria pittieri Baehni =Pouteria glomerata subsp. glomerata 

Pouteria pomifera (Eyma) Baehni =Chrysophyllum pomiferum 

Pouteria porphyrocarpa Baehni =Micropholis porphyrocarpa 

Pouteria ptychandra Eyma =Pradosia ptychandra 

Pouteria robusta (Mart. & Eichler) Eyma =Pouteria cuspidata subsp. robusta 

Pouteria robusta var. longifolia Eyma =Pouteria cuspidata subsp. robusta 

Pouteria rufa (Mart. & Eichler ex Mart.) Lemée =Micropholis guyanensis 

Pouteria scrobiculata Monach. [nom. inval.] =Pouteria scrobiculata 

Pouteria scytalophora Eyma =Pouteria bangii 

Pouteria sericea (Aiton) Baehni [Australian species, recorded from GU in error] 

Pouteria spectabilis Steyerm. =Micropholis spectabilis 

Pouteria spuria (Ducke) Pittier et al. =Chrysophyllum sanguinolentum subsp. spurium


Pouteria steyermarkii Monach. =Pouteria eugeniifolia 

Pouteria suffulta Baehni =Pouteria durlandii subsp. durlandii 

Pouteria surinamensis Eyma =Pradosia surinamensis 

Pouteria tovarensis Klotzsch & Karst. ex Engl. & Prantl =Pouteria guianensis 

Pouteria venamoensis Steyerm. =Micropholis venamoensis 

Pouteria venulosa (Mart. & Eichler) Baehni =Micropholis venulosa 

Prieurella cuneifolia (Rudge) Pierre ex Aubrév. =Chrysophyllum cuneifolium 

Prieurella maguirei Aubrév. =Pouteria maguirei 

Prieurella prieurii (A. DC.) Aubrév. =Chrysophyllum prieurii 

Pseudocladia minutiflora (Britton) Aubrév. =Pouteria coriacea 

Pseudocladia neblinaensis Aubrév. =Pouteria bilocularis 

Pseudocladia orinocoensis Aubrév. =Pouteria lucumifolia 

Pseudocladia scytalophora (Eyma) Aubrév. =Pouteria bangii 

Pseudolabatia filipes (Eyma) Aubrév. =Pouteria filipes 

Pseudolabatia penicillata (Baehni) Aubrév. =Pouteria penicillata 

Pseudoxythece ambelaniifolia (Sandwith) Aubrév. =Pouteria ambelaniifolia 

Radlkoferella brachyandra Aubrév. & Pellegr. =Pouteria brachyandra 

Radlkoferella grandis (Eyma) Aubrév. =Pouteria grandis 

Radlkoferella trigonosperma (Eyma) Aubrév. =Pouteria trigonosperma 

Radlkoferella venosa (Mart.) Aubrév. =Pouteria venosa 

Ragala sanguinolenta Pierre =Chrysophyllum sanguinolentum 

Ragala ulei (K. Krause) Aubrév. =Ecclinusa ulei 

Richardella cayennensis (A. DC.) Aubrév. =Pouteria cayennensis 

Richardella cladantha (Sandwith) Baehni =Pouteria cladantha 

Richardella macrophylla (Lam.) Aubrév. =Pouteria macrophylla 

Richardella surumuensis (Baehni) Aubrév. =Pouteria surumuensis 

Richardella trigonosperma (Eyma) Baehni =Pouteria trigonosperma 

Sandwithiodoxa egregia (Sandwith) Aubrév. =Pouteria egregia 

Sideroxylon acutangulum Ducke =Micropholis acutangula 

Sideroxylon amygdalicarpum Pittier =Pouteria amygdalicarpa 

Sideroxylon bangii Rusby =Pouteria bangii 

Sideroxylon calophylloides (Pierre) Engl. =Micropholis venulosa 

Sideroxylon cuspidatum A. DC. =Pouteria cuspidata 

Sideroxylon cuspidatum var. crassifolium Miq. =Pouteria cuspidata subsp. dura 

Sideroxylon cuspidatum var. ellipticum Miq. =Pouteria cuspidata subsp. cuspidata 

Sideroxylon egense A. DC. =Micropholis egensis 

Sideroxylon elegans A. DC. =Pouteria elegans 

Sideroxylon elegans var. micranthum Miq. =Pouteria cuspidata subsp. cuspidata 

Sideroxylon eugeniifolium (Pierre) Engl. =Pouteria eugeniifolia 

Sideroxylon guianense (Baill.) Engl. =Micropholis venulosa 

Sideroxylon guyanense A. DC. =Micropholis guyanensis 

Sideroxylon robustum Mart. & Eichler =Pouteria cuspidata subsp. robusta 

Sideroxylon rufum Mart. & Eichler ex Mart. =Micropholis guyanensis subsp. guyanensis 

Sideroxylon rugosum var. parviflorum Miq. =Micropholis melinoniana 

Sideroxylon venulosum Mart. & Eichler =Micropholis venulosa 

Syzygiopsis sericea Cronquist =Pouteria arcuata 

Vitellaria pulverulenta (Mart. & Eichler) Radlk. =Pouteria cayennensis 

REFERENCES 

(for a more extensive bibliography, see Pennington 1990) 

Aubréville, A. 1972. Sapotaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part IX. Mem. New York Bot. 

Gard. 23: 1-832. 

Eyma, P.J. 1936. Sapotaceae. In A. Pulle, Flora of Suriname 4(1): 354-399. 

Pennington, T.D. 1990. Sapotaceae. Flora Neotropica Monograph 52: 1-770. 

Pennington, T.D. 2002. Sapotaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


669-683. 

Pennington, T.D. & K.S. Edwards. 2005. Sapotaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


SARRACENIACEAE 

P.E. Berry, K.M. Cameron, R. Riina, J.A. Steyermark† 

Heliamphora chimantensis Wistuba, Carow & Harbarth BO

522 


Heliamphora elongata Nerz 

Heliamphora folliculata Wistuba, Harbarth & Carow 

Heliamphora heterodoxa Steyerm. 

Heliamphora heterodoxa var. exappendiculata Maguire & Steyerm. 

Heliamphora heterodoxa var. heterodoxa 

Heliamphora hispida Wistuba & Nerz 

Heliamphora ionasii Maguire 

Heliamphora minor Gleason 

Heliamphora nutans Benth. 

Heliamphora sarracenioides Carow, Wistuba & Harbarth 

Heliamphora tatei Gleason 

Heliamphora tatei var. neblinae (Maguire) Steyerm. 

Heliamphora tatei var. tatei 


Heliamphora heterodoxa var. glabra Maguire =Heliamphora heterodoxa var. heterodoxa 

Heliamphora macdonaldae Gleason =Heliamphora tatei var. tatei 

Heliamphora neblinae Maguire =Heliamphora tatei var. neblinae 

Heliamphora neblinae var. viridis Maguire =Heliamphora tatei var. neblinae 

Heliamphora tatei var. macdonaldae (Gleason) Maguire =Heliamphora tatei var. tatei 

Heliamphora tatei var. parva Maguire =Heliamphora tatei var. neblinae 

Heliamphora tyleri Gleason =Heliamphora tatei var. tatei 

REFERENCES 

Berry, P.E., R. Riina, & J.A. Steyermark. 2005. Sarraceniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Maguire, B. 1978. Sarraceniaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 10. Mem. New York 

Bot. Gard. 29: 36-62. 

Achetaria guianensis Pennell 

Achetaria ocimoides (Cham. & Schltdl.) Wettst. 

Achetaria scutellarioides (Benth.) Wettst. 

Alectra aspera (Cham. & Schltdl.) L.O. Williams 

Alectra stricta Benth. 

Angelonia angustifolia Benth. [probably cultivated only] 

Angelonia biflora Benth. 

Angelonia salicariifolia Bonpl. [probably cultivated only] 

Anisantherina hispidula (Mart.) Pennell 

Bacopa aquatica Aubl. 

Bacopa bacopoides (Benth.) Pulle 

Bacopa callitrichoides (Kunth) Pennell 

Bacopa connata (Pennell) Pennell 

Bacopa gratioloides (Cham.) Chodat & Hassl. 

Bacopa innominata (M. Gómez) Alain 

Bacopa laxiflora (Benth.) Wettst. ex Edwall 

Bacopa monnieri (L.) Wettst. 

Bacopa monnierioides (Cham.) B.L. Rob. 

Bacopa repens (Sw.) Wettst. 

Bacopa reptans (Benth.) Wettst. ex Edwall 

Bacopa salzmannii (Benth.) Wettst. ex Edwall 

Bacopa serpyllifolia (Benth.) Pennell 

Bacopa sessiliflora (Benth.) Edwall 

Bacopa stricta (Schrad.) Wettst. ex Edwall 

Benjaminia reflexa (Benth.) D’Arcy 

Buchnera longifolia Kunth 

Buchnera palustris (Aubl.) Spreng. 

Buchnera pusilla Kunth 

Buchnera rosea Kunth 

Buchnera spruceana Philcox 

Buchnera ternifolia Kunth 

Buchnera weberbaueri Diels 

Capraria biflora L. 

SCROPHULARIACEAE 

N.H. Holmgren, K.A. Vincent 

AM 

AM 

AM 

AM 

BO 

BO 

BO 

BO 

BO 

BO 

BO 

BO GU 

BO 

BO GU SU FG 

GU SU FG 

BO GU FG 

BO GU FG 

AM BO 

SU FG 

GU 

BO GU SU 



SU 

AM BO 

GU 

BO GU 

BO DA 

AM BO GU SU 

FG 

BO GU 

AM BO GU 

AM GU 

AM BO GU 

AM BO GU 


BO GU 




AM BO GU 

AM BO GU SU 

AM 

BO 

AM BO DA GU 

AM BO DA GU SU FG


Castilleja arvensis Schltdl. & Cham. 

Conobea aquatica Aubl. 

Conobea scoparioides (Cham. & Schltdl.) Benth. 

Escobedia grandiflora (L. f.) Kuntze 

Lindernia brachyphylla Pennell 

Lindernia crustacea (L.) F. Muell. 

Lindernia diffusa (L.) Wettst. 

Lindernia dubia (L.) Pennell 

Lindernia microcalyx Pennell & Stehlé 

Mecardonia procumbens (Mill.) Small 

Micranthemum umbrosum S.F. Blake 

Russelia sarmentosa Jacq. 

Scoparia dulcis L. 

Stemodia foliosa Benth. 

Stemodia pratensis (Aubl.) C.P. Cowan [=Stemodia foliosa ?] 

Stemodia verticillata (Mill.) Hassl. 

Torenia fournieri Linden ex E. Fourn. [probably cultivated or escaped from 

cultivation] 

Torenia thouarsii (Cham. & Schltdl.) Kuntze [naturalized] 

Vellosiella spathacea (Oliv.) Melch. 


Achetaria scutellarioides (Benth.) Kuntze [nom. illeg.] =Achetaria scutellarioides 

Agalinis hispidula (Mart.) D’Arcy =Anisantherina hispidula 

Alectra brasiliensis Benth. =Alectra aspera 

Alectra fluminensis (Vell.) Stearn =Alectra aspera 

Bacopa chamaedryoides (Kunth) Wettst. =Mecardonia procumbens 

Bacopa ciliata (Pennell) Pennell =Bacopa serpyllifolia 

Bacopa conferta (Pennell) Standl. =Bacopa sessiliflora 

Bacopa domingensis (Spreng.) Pennell =Bacopa stricta 

Bacopa punctata Pennell =Bacopa monnierioides 

Bacopa reflexa (Benth.) Edwall =Benjaminia reflexa 

Bacopa sessiliflora (Benth.) Pulle [nom. illeg.] =Bacopa sessiliflora 

Benjaminia utriculariiformis Mart. ex Benj. =Benjaminia reflexa 

Beyrichia ocimoides Cham. & Schltdl. =Achetaria ocimoides 

Beyrichia scutellarioides Benth. =Achetaria scutellarioides 

Bramia monnieri (L.) Drake =Bacopa monnieri 

Buchnera capillaris Desv. =Lindernia crustacea 

Buchnera elongata Sw. [records may represent Buchnera pusilla] =Buchnera longifolia 

Buchnera grandiflora L. f. =Escobedia grandiflora 

Buchnera lithospermifolia Kunth =Buchnera ternifolia 

Buchnera lobelioides Cham. & Schltdl. =Buchnera ternifolia 

Buchnera macrocarpa Kunth =Buchnera longifolia 

Buchnera virgata Kunth [name dubiously applied to Guianan specimens] 

Caconapea auriculata (B.L. Rob.) Pennell =Bacopa laxiflora 

Caconapea conferta Pennell =Bacopa sessiliflora 

Caconapea debilis Pennell =Bacopa reptans 

Caconapea gratioloides Cham. =Bacopa gratioloides 

Caconapea sessiliflora (Benth.) Pennell =Bacopa sessiliflora 

Caconapea stricta (Schrad.) Britton =Bacopa stricta 

Capraria biflora f. hirsuta Loes. =Capraria biflora 

Capraria crustacea L. =Lindernia crustacea 

Capraria integerrima Miq. (Pedaliaceae) =Sesamum orientale 

Conobea innominata M. Gómez =Bacopa innominata 

Curanga bivalvis (L. f.) Druce (Mendonciaceae) =Mendoncia bivalvis 

Drupina cristata L. (Mendonciaceae) =Mendoncia bivalvis 

Erinus procumbens Mill. =Mecardonia procumbens 

Escobedia parimensis Pennell =Escobedia grandiflora 

Gerardia hispidula Mart. =Anisantherina hispidula 

Globifera umbrosum J.F. Gmel. =Micranthemum umbrosum 

Glossostylis aspera Cham. & Schltdl. =Alectra aspera 

Gratiola dubia L. =Lindernia dubia 

Gratiola repens Sw. =Bacopa repens 

Herpestis auriculata B.L. Rob. =Bacopa laxiflora 

Herpestis caprarioides Kunth =Mecardonia procumbens 

BO 


AM 

AM BO 

AM BO GU 




DA GU SU FG 

AM BO DA FG 

DA GU SU 

FG 


BO GU 

GU FG 

GU SU 

GU 

AM DA GU SU 

AM BO GU

524 


Herpestis chamaedryoides Kunth =Mecardonia procumbens 

Herpestis ciliata Pennell =Bacopa serpyllifolia 

Herpestis connata Pennell =Bacopa connata 

Herpestis domingensis Spreng. =Bacopa stricta 

Herpestis laxiflora Benth. =Bacopa laxiflora 

Herpestis procumbens (Mill.) Urb. =Mecardonia procumbens 

Herpestis ranaria Benth. =Bacopa monnierioides 

Herpestis reflexa Benth. =Benjaminia reflexa 

Herpestis reptans Benth. =Bacopa reptans 

Herpestis salzmanni Benth. =Bacopa salzmannii 

Herpestis serpyllifolia Benth. =Bacopa serpyllifolia 

Herpestis sessiliflora Benth. =Bacopa sessiliflora 

Herpestis sessiliflora Benth. =Bacopa sessiliflora 

Herpestis stricta Schrad. =Bacopa stricta 

Hydranthelium callitrichoides Kunth =Bacopa callitrichoides 

Ilysanthes dubia (L.) Barnhart =Lindernia dubia 

Ilysanthes riparia Raf. =Lindernia dubia 

Ilysanthes rotundifolia Benth. =Lindernia rotundifolia 

Lindernia barrosorum L.B. Sm. =Lindernia brachyphylla 

Lindernia rotundifolia (L.) Standl. & L.O. Williams [nom. illeg.] =Lindernia microcalyx 

Macuillamia limosa Pennell =Bacopa repens 

Macuillamia repens (Sw.) Pennell =Bacopa repens 

Matourea pratensis Aubl. =Stemodia pratensis 

Melasma brasiliensis Chodat & Hassl. =Alectra aspera 

Melasma melampyroides (Rich.) Pennell =Alectra aspera 

Melasma spathaceum Oliv. =Vellosiella spathacea 

Mella laxiflora (Benth.) Pennell =Bacopa laxiflora 

Micranthemum orbiculatum Michx. =Micranthemum umbrosum 

Micranthemum pilosum Ernst =Micranthemum umbrosum 

Monocardia humilis Pennell =Bacopa salzmannii 

Monocardia lilacina Pennell =Bacopa salzmannii 

Monocardia violacea Pennell =Bacopa salzmannii 

Naiadothrix reflexa (Benth.) Pennell =Benjaminia reflexa 

Nortenia thouarsii Cham. & Schltdl. =Torenia thouarsii 

Pedicularis melampyroides Rich. =Alectra aspera 

Picria surinamensis Spreng. (Mendonciaceae) =Mendoncia bivalvis 

Piripea palustris Aubl. =Buchnera palustris 

Ranaria monnierioides Cham. =Bacopa monnierioides 

Scrophularia fluminensis Vell. =Alectra aspera 

Sphaerotheca scoparioides Cham. & Schltdl. =Conobea scoparioides 

Stemodia parviflora Aiton =Stemodia verticillata 

Stemodia pusilla Benth. [name apparently misapplied to S. pratensis] 

Stemodia surinamensis Miq. [name of uncertain application] 

Torenia crustacea (L.) Cham. & Schltdl. =Lindernia crustacea 

Torenia thouarsii var. nivea Pennell =Torenia thouarsii 

Vandellia crustacea (L.) Benth. =Lindernia crustacea 

Vandellia diffusa L. =Lindernia diffusa 

REFERENCES 

D’Arcy, W.G. 1979. Scrophulariaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. 

Gard. 66: 173-272. 

Diaz-Miranda, D. 1991. Scrophulariaceae en la flora de Venezuela I. Introducción. Ernstia 1(3): 95-108. 

Holmgren, N.H. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. 

Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76(2): 683-685. 

Holmgren, N.H. & K.A. Vincent. 2005. Scrophulariaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Pennell, F.W. 1940. Some new species of Scrophulariaceae from Guiana. Not. Nat. Acad. Nat. Sci. Phila. 46: 1-6. 

Philcox, D. 1965. Revision of the New World species of Buchnera L. Kew Bull. 18: 275-315. 

Turner, B.L. & C.C. Cowan. 1993. Taxonomic overview of Stemodia for South America. Phytologia 75(4): 281-324. 

SIMAROUBACEAE 

E.V. Franceschinelli, W.W. Thomas 

Picrolemma sprucei Hook. f. AM GU FG

Quassia amara L. [cultivated, naturalized] 

Simaba cedron Planch. [cultivated, naturalized] 

Simaba guianensis Aubl. 

Simaba guianensis subsp. ecaudata (Cronquist) Cavalcante 

Simaba guianensis subsp. guianensis 

Simaba guianensis subsp. huberi Francesch. & W. Thomas 

Simaba monophylla (Oliv.) Cronquist 

Simaba morettii Feuillet 

Simaba obovata Spruce ex Engl. 

Simaba orinocensis Kunth 

Simaba paraensis Ducke 

Simaba polyphylla (Cavalc.) W.W. Thomas 

Simaba sp. A 

Simarouba amara Aubl. 



Aeschrion excelsa (Sw.) Kuntze (Picramniaceae) =Picrasma excelsa 

Aruba guianensis Aubl. =Simaba guianensis 

Picrolemma pseudocoffea Ducke =Picrolemma sprucei 

Quassia cedron (Planch.) Baill. =Simaba cedron 

Quassia cuspidata (Spruce ex Engl.) Noot. =Simaba guianensis 

Quassia guianensis (Aubl.) Dietr. =Simaba guianensis 

Quassia monophylla (Oliv.) Noot. =Simaba monophylla 

Quassia multiflora (A. Juss.) Noot. =Simaba orinocensis 

Quassia simarouba L. f. =Simarouba amara 

Simaba alata Maguire =Simaba orinocensis 

Simaba cuspidata Spruce ex Engl. =Simaba guianensis 

Simaba guianensis subsp. polyphylla Cavalcante =Simaba polyphylla 

Simaba multiflora A. Juss. =Simaba orinocensis 

Simaba trichilioides A. St.-Hil. [not in flora area] 

Simarouba monophylla Oliv. =Simaba monophylla 

REFERENCES 

Cavalcante, P.B. 1983. Revisao taxonômica do gênero Simaba Aublet na América do Sul. Publ. Avulsas Mus. Goeldi 37: 1-85. 

Cronquist, A. 1944. Studies in the Simaroubaceae. II. The genus Simarouba. Bull. Torrey Bot. Club 71: 226-234. 

Cronquist, A. 1944. Studies in Simaroubaceae. III. The genus Simaba. Lloydia 7: 81-92. 

Cronquist, A. 1944. Studies in Simaroubaceae. IV. Resume of the American genera. Brittonia 5: 128-147. 

Cronquist, A. 1945. Additional notes on the Simaroubaceae. Brittonia 5: 469-470. 

Feuillet, C. 1983. Etudes sur les Simaroubaceae. II. Un Simaba nouveau de Guyane française dans la section Floribundae Engl.: 

S. morettii. Candollea 38(2): 745-750. 

Jansen-Jacobs, M.J. 1979. Simaroubaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 319-330. 

Nooteboom, H.P. 1962. Generic delimitation in Simaroubaceae tribus Simaroubeae and a conspectus of the genus Quassia L. 

Blumea 11: 509-528. 

Thomas, W.W. 1984. A new species of Simaba (Simaroubaceae) from Pará, Brazil, with a key to the species north of the Amazon 

River. Brittonia 36(3): 244-247. 

Thomas, W.W. 1985. The Simaba guianensis complex in northern South America. Acta Amazonica 15(1-2) suppl.: 71-79. 

Thomas, W.W. 2002. Simaroubaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 685-689. 

Thomas, W.W. & E.V. Franceschinelli. 2005. Simaroubaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Siparuna bifida (Poepp. & Endl.) A. DC. 

Siparuna cristata (Poepp. & Endl.) A. DC. 

Siparuna cuspidata (Tul.) A. DC. 

Siparuna decipiens (Tul.) A. DC. 

Siparuna ficoides S.S. Renner & Hausner 

Siparuna guianensis Aubl. 

Siparuna obstipa J.F. Macbr. 

Siparuna pachyantha A.C. Sm. 

Siparuna poeppigii (Tul.) A. DC. 

Siparuna reginae (Tul.) A. DC. 

SIPARUNACEAE 

G. Hausner, S.S. Renner 

BO GU SU FG 


AM GU SU FG 

GU SU FG 

GU SU FG 

AM 

AM BO GU 

FG 

AM 


GU 

FG 

BO DA 


AM BO 


AM GU SU FG 


BO 


AM 

GU FG 

GU SU FG 

AM BO GU SU

526 



Citrosma amazonica (Mart.) A. DC. =Siparuna poeppigii 

Citrosma bifida Poepp. & Endl. =Siparuna bifida 

Citrosma cristata Poepp. & Endl. =Siparuna cristata 

Citrosma cuspidata Tul. =Siparuna cuspidata 

Citrosma decipiens Tul. =Siparuna decipiens 

Citrosma guianensis (Aubl.) Tul. =Siparuna guianensis 

Citrosma reginae Tul. =Siparuna reginae 

Conuleum guianense A. Rich. =Siparuna decipiens 

Siparuna amazonica (Mart.) A. DC. =Siparuna poeppigii 

Siparuna argyrochrysea Perkins =Siparuna cuspidata 

Siparuna emarginata R.S. Cowan =Siparuna pachyantha 

Siparuna micrantha A. DC. =Siparuna reginae 

Siparuna sancheziana Steyerm. =Siparuna reginae 

Siparuna sprucei A. DC. =Siparuna poeppigii 

Siparuna surinamensis Lanj. = Siparuna vequinae (Tul.) A.DC. 

REFERENCES 

Jansen-Jacobs, M.J. 1976. Monimiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 425- 

426. 

Lanjouw, J. 1939. Monimiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 471-472. 

Mitchell, J.D. 2002. Monimiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


510-515. 

Perkins, J. & E. Gilg. 1901. Monimiaceae. In A. Engler, Das Pflanzenreich IV(101): 1-122. 

Petter, H.F.M. 1932. Monimiaceae. In A. Pulle, Flora of Suriname 2(1): 107-112. 

Renner, S.S. & G. Hausner. 2005. Siparunacae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Steyermark, J.A. 1951. Monimiaceae. In Botanical Exploration in Venezuela - I. Fieldiana Bot. 28(1): 235-236. 

SOLANACEAE 

W.G. D’Arcy, C. Benítez de Rojas, M.H. Nee 

Brunfelsia guianensis Benth. 

Brunfelsia imatacana Plowman 

Brunfelsia martiana Plowman 

Brunfelsia uniflora (Pohl) D. Don 

Capsicum annuum L. 

Capsicum annuum var. annuum [cultivated, frequently escaping] 

Capsicum annuum var. glabriusculum (Dunal) Heiser & Pickersgill 

Capsicum frutescens L. 

Capsicum spp. 

Cestrum alternifolium (Jacq.) O.E. Schulz 

Cestrum glabrescens (C.V. Morton) Steyerm. & Maguire 

Cestrum latifolium Lam. 

Cestrum multiflorum Roem. & Schult. 

Cestrum neblinense D’Arcy & Benítez 

Cestrum racemosum Ruiz & Pav. 

Cestrum reflexum Sendtn. 

Cestrum schlechtendahlii G. Don 

Cestrum schulzianum Francey 

Cestrum strigilatum Ruiz & Pav. 

Cestrum subuniflorum Dunal 

Cestrum tubulosum Sendtn. 

Cyphomandra endopogon Bitter 

Cyphomandra endopogon subsp. guianensis Bohs 

Cyphomandra hartwegii (Miers) Walp. 

Cyphomandra hartwegii subsp. hartwegii 

Cyphomandra oblongifolia Bohs 

Cyphomandra tegore (Aubl.) Walp. 

Cyphomandra tobagensis Sandwith 

Datura inoxia Mill. 

Datura metel L. [cultivated or escaped from cultivation] 

Datura stramonium L. [naturalized?] 

GU SU FG 

BO 

GU SU FG 

BO GU 


GU FG 



BO 

AM BO 


FG 

AM 

AM GU? FG? 

BO 

AM BO GU FG 

AM 

AM 

AM 

AM BO 

FG 

FG 

GU SU FG 

GU SU FG 

SU 

SU FG 

BO GU 

BO 

SU FG 

FG?


Deprea orinocensis (Kunth) Raf. 

Hawkesiophyton ulei (Dammer) Hunz. 

Lycianthes geminata Bitter [expected in GU, SU] 

Lycianthes heteroclita (Sendtn.) Bitter 

Lycianthes nitida Bitter 

Lycianthes pauciflora (Vahl) Bitter 

Markea coccinea Rich. 

Markea formicarum Dammer 

Markea longiflora Miers 

Markea reticulata Steyerm. & Maguire 

Markea sessiliflora Ducke 

Melananthus ulei Carvalho 

Nicotiana tabacum L. [cultivated, perhaps escaping] 

Physalis angulata L. 

Physalis cordata Mill. 

Physalis ignota Britton [probably cultivated or escaped from cultivation] 

Physalis lagascae Roem. & Schult. 

Physalis pubescens L. 

Schwenckia americana L. 

Schwenckia elegans Carvalho 

Schwenckia glabrata Kunth 

Schwenckia grandiflora Benth. 

Schwenckia heterantha Carvalho 

Schwenckia huberi Benítez 

Schwenckia lateriflora (Vahl) Carvalho 

Schwenckia micrantha Benth. 

Schwenckia mollissima Nees & Mart. 

Solandra grandiflora Sw. 

Solandra longiflora Tussac 

Solandra paraensis Ducke 

Solanum accrescens Standl. & C.V. Morton 

Solanum acerifolium Dunal 

Solanum adhaerens Roem. & Schult. 

Solanum agrarium Sendtn. [expected in SU, FG] 

Solanum allophyllum (Miers) Standl. 

Solanum altissimum Pittier ex. Benítez 

Solanum americanum Mill. 

Solanum anceps Ruiz & Pav. 

Solanum arboreum Dunal 

Solanum asperum Rich. 

Solanum aturense Dunal 

Solanum badilloi Benítez 

Solanum bicolor Roem. & Schult. 

Solanum campaniforme Roem. & Schult. 

Solanum capsicoides All. [introduced] 

Solanum caripense Dunal 

Solanum circinatum Bohs 

Solanum cordovense Sessé & Moc. 

Solanum coriaceum Dunal 

Solanum costatum M. Nee 

Solanum crinitum Lam. 

Solanum davidsei Carvalho 

Solanum erythrotrichum Fernald 

Solanum extensum Bitter 

Solanum fulvidum Bitter 

Solanum hirtum Vahl 

Solanum jamaicense Mill. 

Solanum lanceifolium Jacq. 

Solanum lepidotum Dunal 

Solanum leucocarpon Dunal 

Solanum mammosum L. 

Solanum maroniense Poit. 

Solanum monachophyllum Dunal 

Solanum morii S. Knapp 

BO 

AM FG 

FG 

SU 

GU 


GU SU FG 

AM GU SU FG 


AM BO 


BO 



BO DA 

GU 

AM BO 



AM BO GU FG 

AM BO 


AM 

AM 

GU 

BO DA 

SU 

AM 

AM GU SU FG? 

FG 

FG? 

AM BO GU 

BO DA GU FG 

GU 

BO 

AM 

BO GU SU FG 

GU SU FG 

AM BO DA 


AM BO FG? 

AM 

AM BO GU 

BO GU FG 

GU SU 

AM 

GU 

GU 

BO GU SU FG 

GU SU FG 


BO 

GU? 

BO GU 

GU FG 

BO 


BO DA 

GU FG? 


BO GU SU FG 

FG 


GU FG

528 


Solanum nuricum M. Nee 

BO 

Solanum oocarpum Sendtn. 

BO GU SU? 

Solanum oppositifolium Ruiz & Pav. 


Solanum paludosum Moric. 

BO GU SU FG 

Solanum pensile Sendtn. 


Solanum rubiginosum Vahl 

GU SU FG 

Solanum rufistellatum Steyerm. 

BO 

Solanum rugosum Dunal 


Solanum schlechtendalianum Walp. 


Solanum schomburgkii Sendtn. 

BO GU SU FG 

Solanum seaforthianum Andrews 

AM BO DA FG 

Solanum semotum M. Nee 

GU FG 

Solanum sendtnerianum Van Heurck & Müll. Arg. 

FG 

Solanum sessiliflorum Dunal [probably cultivated or escaped from cultivation] AM BO SU FG 

Solanum stramoniifolium Jacq. 


Solanum subinerme Jacq. 


Solanum swartzianum Roem. & Schult. 

AM BO 

Solanum tepuiense S. Knapp 

BO 

Solanum torvum Sw. 

GU FG 

Solanum tricuspidatum Rich. ex Dunal 

FG 

Solanum umbratile I.M. Johnst. 

BO 

Solanum vanheurckii Müll. Arg. 

FG 

Solanum velutinum Dunal 

BO GU SU FG 

Solanum vestissimum Dunal 

AM 

Witheringia solanacea L’Hér. 

AM FG 


Bassovia macrophylla (Dunal) J. Coulter =Witheringia solanacea 

Bassovia sylvatica Aubl. =Solanum anceps 

Brachistus macrophyllus (Dunal) Miers =Witheringia solanacea 

Brunfelsia maritima Benth. [not in flora area] =Brunfelsia latifolia (Pohl) Benth. 

Brunfelsia spruceana Benth. =Brunfelsia martiana 

Capsicum conoide Mill. ? =Capsicum annuum 

Capsicum curvipes Dunal ? =Capsicum annuum 

Capsicum hispidum var. glabruiusculum Dunal =Capsicum annuum var. glabriusculum 

Capsicum macrophyllum (Dunal) Standl. =Witheringia solanacea 

Capsicum solanaceum var. macrophylla (Dunal) Kuntze =Witheringia solanacea 

Cestrum acuminatissimum Dunal =Cestrum schlechtendahlii 

Cestrum alternifolium var. mitanthum O.E. Schulz =Cestrum alternifolium 

Cestrum baenitzii Lingelsh. =Cestrum schlechtendahlii 

Cestrum grande Pittier =Cestrum racemosum 

Cestrum latifolium var. tenuiflorum (Kunth) O.E. Schulz =Cestrum latifolium 

Cestrum megalophyllum Dunal =Cestrum schlechtendahlii 

Cestrum oliganthum Dunal =Cestrum latifolium 

Cestrum oliganthum var. latifolium Dunal =Cestrum latifolium 

Cestrum prieurei Dunal =Cestrum latifolium 

Cestrum racemosum var. grande (Pittier) Francey =Cestrum racemosum 

Cestrum silvaticum Francey [not in flora area] =Cestrum microcalyx Francey 

Cestrum tenuiflorum Kunth =Cestrum latifolium 

Cestrum tenuiflorum var. glabrescens C.V. Morton =Cestrum glabrescens 

Cyphomandra allophylla (Miers) Hemsl. =Solanum allophyllum 

Cyphomandra bolivarensis Steyerm. =Cyphomandra tobagensis 

Datura fastuosa L. =Datura metel 

Ectozoma ulei Dammer =Hawkesiophyton ulei 

Franciscea uniflora Pohl =Brunfelsia uniflora 

Hawkesiophyton panamense (Standl.) Hunz. =Hawkesiophyton ulei 

Lycianthes guianensis (Dunal) Bitter =Lycianthes pauciflora 

Markea camponoti Ducke =Markea longiflora 

Markea panamensis Standl. =Hawkesiophyton ulei 

Markea porphyrobaphes Sandwith =Markea sessiliflora 

Physalis capsicifolia Dunal [name of uncertain application] =Physalis sp. 

Physalis hirsuta Dunal [name of uncertain application] =Physalis sp. 

Physalis orinocensis Kunth =Deprea orinocensis 

Physalis surinamensis Miq. =Physalis pubescens


Physalis surinamensis auct. non Miq.: sensu Steyerm. =Physalis pubescens 

Pionandra allophylla Miers =Solanum allophyllum 

Saracha solanacea (L’Hér.) Miers =Witheringia solanacea 

Schwenckia americana var. angustifolia J.A. Schmidt =Schwenckia americana 

Schwenckia americana var. hirta (Klotzsch ex Benth.) Carvalho =Schwenckia americana 

Schwenckia guianensis Benth. =Schwenckia americana 

Schwenckia guineensis Schumach. & Thonn. =Schwenckia americana 

Schwenckia hirta Klotzsch ex Benth. =Schwenckia americana 

Schwenckia patens Kunth =Schwenckia glabrata 

Schwenckia ulei Dammer =Schwenckia grandiflora 

Sicklera tetrandra A. Braun & Bouché =Witheringia solanacea 

Solanum altissimum Pittier ex. Benítez [nomen nudum] =Solanum altissimum 

Solanum americanum var. nodiflorum (Jacq.) Edmonds =Solanum americanum 

Solanum antillarum O.E. Schulz =Solanum nudum 

Solanum arcuatum Sendtn. =Solanum acerifolium 

Solanum arecunarum Pittier et al. [nomen nudum] =Solanum sessiliflorum 

Solanum asperrimum Bitter & O. Moritz =Solanum aturense 

Solanum aubletii Pulle =Solanum anceps 

Solanum boissieri Dunal =Solanum schomburgkii 

Solanum bolivarense Pittier et al. [nomen nudum] =Solanum monachophyllum 

Solanum brachyacanthum Dunal =Solanum lanceifolium 

Solanum brachybotryon Dunal =Solanum velutinum 

Solanum callicarpifolium Kunth & Bouché =Solanum bicolor 

Solanum confusum Pittier et al. [nomen nudum] =Solanum velutinum 

Solanum cuneifolium Dunal =Solanum jamaicense 

Solanum cyananthum Dunal =Solanum crinitum 

Solanum cyananthum var. jubatum (Roem. & Schult.) Dunal =Solanum crinitum 

Solanum cyrrhosum Dunal =Solanum seaforthianum 

Solanum demerarense Dunal =Solanum stramoniifolium 

Solanum demerarense var. inerme Dunal =Solanum stramoniifolium 

Solanum diphyllum L. [not in flora area] 

Solanum dolichostylum O.E. Schulz =Solanum arboreum 

Solanum donnellsmithii J. Coulter =Solanum adhaerens 

Solanum echinocarpum Bredem. ex Roem. & Schult. =Solanum hirtum 

Solanum enoplocalyx Dunal =Solanum adhaerens 

Solanum flavescens Dunal =Solanum hirtum 

Solanum flavescens var. albescens Dunal =Solanum hirtum 

Solanum foetidum Ruiz & Pav. [not in flora area] 

Solanum formosum Dunal =Solanum crinitum 

Solanum fraternorum Pittier et al. [nomen nudum] =Solanum velutinum 

Solanum geminifolium Schltdl. & Cham. =Solanum schlechtendalianum 

Solanum globiferum Dunal =Solanum mammosum 

Solanum granelianum D’Arcy =Solanum pensile 

Solanum guianense Klotzsch [nomen nudum] 

Solanum guianense Dunal =Lycianthes pauciflora 

Solanum hastatum Sendtn. =Solanum acerifolium 

Solanum heterophyllum Lam. =Solanum subinerme 

Solanum hostmannii Dunal =Solanum schomburgkii 

Solanum humboldtii Dunal =Solanum adhaerens 

Solanum hyporrhodium Pittier et al. [nomen nudum] =Solanum vestissimum 

Solanum intermedium Sendtn. [not in flora area] 

Solanum ipomoea Sendtn. =Solanum pensile 

Solanum japurense Dunal =Lycianthes sp. 

Solanum javitense Kunth [nom. superfl.] =Solanum monachophyllum 

Solanum jubatum Roem. & Schult. =Solanum crinitum 

Solanum juripeba Rich. =Solanum subinerme 

Solanum laetum Miq. =Solanum pensile 

Solanum lanceifolium var. brachyacanthum (Dunal) O.E. Schulz =Solanum lanceifolium 

Solanum lanugiflorum Pittier et al. [nomen nudum] =Solanum velutinum 

Solanum longiflorum Vahl =Solanum subinerme 

Solanum macranthum Dunal =Solanum crinitum 

Solanum manaense Sagot ex Lemée [nomen nudum] 

Solanum mesopliarthrum Pittier et al. [nomen nudum] =Solanum velutinum 

Solanum mesopliarthrum Benítez & Steyerm. =Solanum velutinum

530 


Solanum miquelii C.V. Morton =Solanum pensile 

Solanum molle Dunal =Solanum monachophyllum 

Solanum nigrum L. [misident. for S. americanum fide Nee] 

Solanum nodiflorum Jacq. =Solanum americanum 

Solanum nudum Dunal [not in flora area] 

Solanum obscurum herb. Willd. =Solanum lanceifolium 

Solanum obtusifrons Pittier et al. [nomen nudum] =Solanum hirtum 

Solanum ochraceo-ferrugineum (Dunal) Fernald [not in flora area] 

Solanum oleraceum Dunal ex Poir. =Solanum americanum 

Solanum orenocense Dunal =Solanum hirtum 

Solanum orientale Benítez =Solanum oocarpum 

Solanum orinocense Dunal =Solanum stramoniifolium 

Solanum pauciflorum Vahl =Lycianthes pauciflora 

Solanum platanifolium Hook. =Solanum mammosum 

Solanum platyphyllum Dunal =Solanum stramoniifolium 

Solanum prehensile Pittier et al. [nomen nudum] =Solanum pensile 

Solanum proteanthum Bohs =Cyphomandra oblongifolia 

Solanum prunifolium Roem. & Schult. =Solanum seaforthianum 

Solanum pterocaulon Dunal =Solanum americanum 

Solanum puberuloba Steyerm. =Solanum oppositifolium 

Solanum quinquangulare Roem. & Schult. =Solanum acerifolium 

Solanum radula Vahl =Solanum asperum 

Solanum rhamphidacanthum Bitter =Solanum velutinum 

Solanum richardii (Dunal) Lemée ? =Solanum anceps 

Solanum ripivagum Pittier et al. [nomen nudum] =Solanum arboreum 

Solanum sacupanense Rusby =Solanum monachophyllum 

Solanum salignum Roem. & Schult. =Solanum seaforthianum 

Solanum salviifolium auct. =Solanum asperum 

Solanum salzmannii Dunal =Solanum paludosum 

Solanum scabridum Dunal =Solanum vestissimum 

Solanum scandens L. f. =Solanum pensile 

Solanum scandens L. =Solanum pensile 

Solanum secundum Bitter & O. Moritz =Solanum aturense 

Solanum sempervirens Dunal =Solanum pensile 

Solanum siparunoides Ewan =Solanum aturense 

Solanum stamineum Steud. =Solanum subinerme 

Solanum stramoniifolium var. inerme (Dunal) Whalen =Solanum stramoniifolium 

Solanum styracifolium Roem. & Schult. =Solanum subinerme 

Solanum styracoides Rusby =Solanum pensile 

Solanum surinamense Steud. =Solanum leucocarpon 

Solanum sylvaticum (Aubl.) Bitter =Solanum anceps 

Solanum tegore Aubl. =Cyphomandra tegore 

Solanum timotense Pittier et al. [nomen nudum] =Solanum hirtum 

Solanum tobagense (Sandwith) Bohs =Cyphomandra tobagensis 

Solanum topirito Pittier et al. [nomen nudum] =Solanum lanceifolium 

Solanum topiro Dunal =Solanum sessiliflorum 

Solanum toxicarium Rich. =Solanum stramoniifolium 

Solanum trichocarpum Miq. =Solanum stramoniifolium 

Solanum triste var. crassipes Dunal =Solanum leucocarpon 

Solanum umbrosum Dunal =Solanum lanceifolium 

Solanum undecimangulare Roem. & Schult. =Solanum stramoniifolium 

Solanum verbascifolium L. [not in flora area] 

Witheringia macrophylla Dunal =Witheringia solanacea 

REFERENCES 

Bernardello, L.M. & A.T. Hunziker. 1987. A synoptical revision of Solandra (Solanaceae). Nord. J. Bot. 7: 639-652. 

Bitter, G. 1919. Die Gattung Lycianthes. Ahb. Naturwiss. Vereine Bremen 24(2): 292-520. 

Bohs, L. 1994. Cyphomandra (Solanaceae). Flora Neotropica Monograph 63: 1-175. 

Carvalho, L. d’A Freire de. 1978. O gênero Schwenckia D. van Rooyen ex Linnaeus no Brasil - Solanaceae. Rodriguésia 44: 

307-524. 

Cuatrecasas, J. 1958. Notes on American Solanaceae. Feddes Repert. 61(1): 74-86. 

D’Arcy, W.G. 1973. Solanaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot. Gard. 

60: 573-780. 

D’Arcy, W.G. , C. Benítez de Rojas, & M.H. Nee. 2005. Solanaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. 

Holst, Flora of the Venezuelan Guayana 9: 194-246.


Edmonds, J.M. 1972. A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America. Kew Bull. 27: 95- 

114. 

Francey, P. 1935-1936. Monographie du genre Cestrum L. Candollea 6: 46-398, 7: 1-132. 

Hawkes, J.G., R.N. Lester, M. Nee, & N. Estrada-R. 1991. Solanaceae III. Taxonomy, Chemistry, Evolution. Royal Botanic 

Gardens, Kew. 

Nee, M. 2002. Solanaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


700. 

Plowman, T. 1979. The genus Brunfelsia: a conspectus of the taxonomy and biogeography. Pp. 475-491 in J.G. Hawkes, R.N. 

Lester, & A.D. Skelding, The Biology and Taxonomy of the Solanaceae. Linnaean Society of London, Academic Press. 

Whalen, M.D. 1984. Conspectus of species groups in Solanum subgenus Leptostemonum. Gentes Herb. 12(4): 179-282. 

Whalen, M.D., D.E. Costich, & C.B. Heiser. 1981. Taxonomy of Solanum section Lasiocarpa. Gentes Herb. 12(2): 41-129. 

Sphenoclea zeylanica Gaertn. [naturalized?] 

SPHENOCLEACEAE 

B.A. Stein 

REFERENCES 

Stein, B.A. 2005. Sphenocleaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 247-248. 

Ayenia magna L. 

Ayenia praeclara Sandwith 

Ayenia tomentosa L. 

Byttneria aristeguietae Cristóbal 

Byttneria aurantiaca Mildbr. 

Byttneria caripensis Cristóbal 

Byttneria catalpifolia Jacq. 

Byttneria catalpifolia subsp. catalpifolia 

Byttneria cordifolia Sagot 

Byttneria divaricata Benth. 

Byttneria divaricata var. divaricata 

Byttneria genistella Triana & Planch. 

Byttneria morii L.C. Barnett & Dorr 

Byttneria obliqua Benth. 

Byttneria piresii Cristóbal 

Byttneria rhamnifolia Benth. 

Byttneria scabra L. 

Byttneria uaupensis Spruce ex K. Schum. 

Guazuma ulmifolia Lam. 

Guazuma ulmifolia var. tomentella K. Schum. 

Guazuma ulmifolia var. ulmifolia 

Helicteres baruensis Jacq. 

Helicteres brevispira A. St.-Hil. 

Helicteres guazumifolia Kunth 

Helicteres heptandra L.B. Sm. 

Helicteres pentandra L. 

Herrania camargoana R.E. Schult. 

Herrania kanukuensis R.E. Schult. 

Herrania lemniscata (M.R. Schomb.) R.E. Schult. 

Melochia arenosa Benth. 

Melochia graminifolia A. St.-Hil. 

Melochia lanceolata Benth. 

Melochia manducata C. Wright 

Melochia melissifolia Benth. 

Melochia nodiflora Sw. 

Melochia parvifolia Kunth 

Melochia parvifolia var. fasciculata (Benth.) Hassl. 

Melochia parvifolia var. parvifolia 

STERCULIACEAE 

P.E. Berry, C.L. Cristóbal, L.J. Dorr, J.G. Saunders 


BO 

AM BO 

GU SU 

GU SU 

FG 

BO 

AM GU 

AM GU 

SU FG 

DA GU SU FG 

DA GU SU 

AM BO GU FG 

AM 

AM 

GU FG 

AM BO 


BO GU 


AM BO DA 

AM BO 

BO GU SU 

AM GU 

AM BO 

BO 

GU 

AM 

BO GU? SU FG 

GU SU FG 



AM BO GU 

DA 

GU SU FG? 

BO 

GU SU FG 


GU SU? 

AM BO DA GU

532 


Melochia pyramidata L. 

Melochia pyramidata var. pyramidata 

Melochia simplex A. St.-Hil. 

Melochia tomentosa L. 

Melochia tomentosa var. frutescens DC. 

Melochia tomentosa var. tomentosa 

Melochia ulmifolia Benth. 

Melochia villosa (Mill.) Fawc. & Rendle 

Melochia villosa var. tomentosa (K. Schum.) Goldberg 

Melochia villosa var. villosa 

Sterculia abbreviata E.L. Taylor ex Mondragón 

Sterculia amazonica E.L. Taylor ex Mondragón 

Sterculia apetala (Jacq.) H. Karst 

Sterculia frondosa Rich. 

Sterculia guianensis Sandwith 

Sterculia kayae P.E. Berry 

Sterculia lisae E.L Taylor, sp. nov. ined. 

Sterculia multiovula E.L Taylor, sp. nov. ined. 

Sterculia pruriens (Aubl.) K. Schum. 

Sterculia pruriens var. glabrescens E.L. Taylor, var. nov. ined. 

Sterculia pruriens var. pruriens 

Sterculia rugosa R. Br. 

Sterculia speciosa K. Schum. 

Sterculia villifera Steud. 

Theobroma bicolor Bonpl. 

Theobroma cacao L. 

Theobroma cacao subsp. sphaerocarpum (A. Chev.) Cuatrec. 

Theobroma grandiflorum (Willd. ex Spreng.) K. Schum. 

Theobroma microcarpum Mart. 

Theobroma obovatum Klotzsch ex Bernoulli 

Theobroma subincanum Mart. 

Theobroma velutinum Benoist 

Uladendron codesuri Marc.-Berti 

Waltheria albicans Turcz. 

Waltheria berteroi (Spreng.) J.G. Saunders 

Waltheria carmensarae J.G. Saunders 

Waltheria collina K. Schum. 

Waltheria indica L. 

Waltheria involucrata Benth. 

Waltheria microphylla Cav. [considered part of Waltheria indica complex by 

L.J. Dorr] 

Waltheria operculata Rose 

Waltheria paniculata Benth. 

Waltheria viscosissima A. St.-Hil. 


Bubroma grandiflorum Willd. ex Spreng. =Theobroma grandiflorum 

Buttneria scabra Aubl. =Byttneria scabra 

Byttneria sparganiocarpa Uittien =Byttneria divaricata 

Byttneria venezuelensis Steyerm. =Byttneria divaricata var. divaricata 

Cacao guianensis Aubl. =Theobroma subincanum 

Cacao sylvestris Aubl. =Theobroma subincanum 

Helicteres apetala Jacq. =Sterculia apetala 

Helicteres brachypetala Leane =Helicteres heptandra 

Helicteres bracteosa Mart. =Helicteres pentandra 

Helicteres pronifolia Rich. =Helicteres pentandra 

Herrania mariae (Mart.) Decne. ex Goudot [not in flora area] 

Ivira pruriens Aubl. =Sterculia pruriens 

Lightia lemniscata M.R. Schomb. =Herrania lemniscata 

Malachra berterii Spreng. =Waltheria berteroi 

Melochia benthamii K. Schum. =Melochia ulmifolia 

Melochia carpinifolia J.C. Wendl. =Melochia nodiflora 

Melochia carthaginensis Willd. ex K. Schum. =Melochia ulmifolia 

Melochia cinerea A. St.-Hil. & Naudin =Melochia arenosa 

FG? 

FG? 

GU SU 

BO 

BO 

BO 

SU FG? 

BO GU SU FG 

AM BO 

BO 

DA GU SU FG 

AM BO DA 

AM DA 

AM BO 

BO DA 

GU 

FG 


FG 

GU FG 

AM BO DA GU 

GU 

SU FG 

GU SU FG 

AM BO GU 

GU FG 

AM 

GU? SU FG 



AM 

AM 

GU 

GU 


BO 

SU FG 

AM BO GU 

AM BO 

BO 

BO 


AM BO DA GU 

GU 

BO 

AM BO 

GU 

GU


Melochia concinna Miq. =Melochia melissifolia 

Melochia conglobata Sessé & Moc. =Melochia nodiflora 

Melochia fasciculata Benth. =Melochia parvifolia var. fasciculata 

Melochia glandulifera Standl. =Melochia manducata 

Melochia hirsuta Cav. =Melochia villosa 

Melochia hirsuta var. tomentosa K. Schum. =Melochia villosa var. tomentosa 

Melochia llanorum Cuatrec. =Melochia arenosa 

Melochia obovata Urb. =Melochia parvifolia var. parvifolia 

Melochia ramuliflora R.E. Fr. =Melochia ulmifolia 

Melochia scordiifolia Turcz. =Melochia parvifolia var. parvifolia 

Melochia spicata (L.) Fryxell [not in flora area, or = M. villosa] 

Quamoclit rodriguesii J.I. Falcno =Melochia graminifolia 

Riedlea dichotoma Turcz. =Melochia ulmifolia 

Riedlea heterotricha Turcz. =Melochia villosa 

Riedlea littoralis K. Schum. [nom. superfl.] =Melochia arenosa 

Riedlea littoralis Poepp. & Endl. =Melochia arenosa 

Riedlea ramuliflora Miq. =Melochia ulmifolia 

Riedlea urticifolia Turcz. =Melochia nodiflora 

Sida villosa Mill. =Melochia villosa 

Sitella involucrata (Benth.) L.H. Bailey =Waltheria involucrata 

Sterculia excelsa Mart. [not in flora area] 

Sterculia parviflora (Ducke) E.L. Taylor [nom. illeg.] =Sterculia kayae 

Sterculia pruriens var. parviflora Ducke =Sterculia kayae 

Theobroma alba Bernoulli (Chrysobalanaceae) =Licania alba 

Theobroma guianense (Aubl.) J.F. Gmel. =Theobroma subincanum 

Theobroma mariae (Mart.) K. Schum. [not in flora area] =Herrania mariae (Mart.) Decne. ex Goud. 

Theobroma speciosum Mart. [not in flora area] 

Theobroma sphaerocarpa A. Chev. =Theobroma cacao subsp. sphaerocarpum 

Theobroma sylvestre Mart., non Cacao sylvestris Aubl. [not in flora area] 

Theobroma sylvestris (Aubl.) G. Don =Theobroma subincanum 

Turnera valleana Standl. & L.O. Williams =Waltheria operculata 

Waltheria americana L. =Waltheria indica 

Waltheria hirsuta C. Presl =Waltheria viscosissima 

Waltheria machrisiantha L.B. Sm. =Waltheria viscosissima 

Waltheria subcordata Standl. =Waltheria berteroi 

Waltheria tubiflora Klotzsch =Waltheria viscosissima 

Xylosterculia rugosa (R. Br.) Kosterm. =Sterculia rugosa 

REFERENCES 

Barnett, L.C. & L.J. Dorr. 1990. A new, arborescent species of Byttneria from French Guiana. Brittonia 42(4): 271-275. 

Cristóbal, C.L. 1976. Estudio taxonómico del género Byttneria Loefl. Bonplandia 4: 1-428. 

Cristóbal, C.L., J.G. Saunders, & P.E. Berry. 2005. Sterculiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, 


Cuatrecasas, J. 1964. Cacao and its allies. A taxonomic revision of the genus Theobroma. Contr. U.S. Nat. Herb. 35: 379-614. 

Dorr, L.J. 2002. Sterculiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


706. 

Goldberg, A. 1967. The genus Melochia L. Contrib. United States National Herbarium 34(5): 191-363. 

Saunders, J.G. 1995. Systematics and evolution of Waltheria. Dissertation, University of Texas, Austin. 

Uittien, H. 1932. Sterculiaceae. In A. Pulle, Flora of Suriname 3(1): 34-48. 

Uittien, H. 1941. Sterculiaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 437. 

Styrax argenteus C. Presl 

Styrax argenteus var. ramirezii (Greenm.) Gonsoulin 

Styrax camporus Pohl 

Styrax discolor M.F. Silva 

Styrax duidae Steyerm. 

Styrax duidae subsp. duidae 

Styrax duidae subsp. paruae Maguire 

Styrax glaber Sw. 

Styrax glabratus Schott 

STYRACACEAE 

P.W. Fritsch, D. Sabatier, J.A. Steyermark† 

GU? 

FG 

FG 

FG 

AM 

AM 

AM 

BO GU SU 

AM SU FG

534 

Styrax guaiquinimae (Maguire & Steyerm.) P.W. Fritsch 

Styrax guanayanus Maguire & K.D. Phelps 

Styrax guyanensis A. DC. 

Styrax hypochryseus Perkins 

Styrax lancifolius Klotzsch ex Seub. 

Styrax longifolius Standl. 

Styrax longipedicellatus Steyerm. 

Styrax neblinae (Maguire) P.W. Fritsch 

Styrax oblongus (Ruiz & Pav.) A. DC. 

Styrax pallidus A. DC. 

Styrax sieberi Perkins 

Styrax sipapoanus Maguire 

Styrax tafelbergensis Maguire 

Styrax tessmannii Perkins 

Styrax wurdackiorum Steyerm. 

Styrax yutajensis (Maguire) P.W. Fritsch 



Strigilia glabrata (Schott) Miers =Styrax glabratus 

Strigilia guyanensis (A. DC.) Miers =Styrax guyanensis 

Strigilia psilophylla (A. DC.) Miers =Styrax glabratus 

Styrax costanus Steyerm. =Styrax glaber 

Styrax duidae subsp. neblinae Maguire =Styrax neblinae 

Styrax fanshawei Sandwith =Styrax glaber 

Styrax filii Maguire =Styrax longipedicellatus 

Styrax glaber var. micranthus Perkins =Styrax glaber 

Styrax guanayanus var. yutajensis Maguire =Styrax yutajensis 

Styrax guyanensis var. japurensis Seub. =Styrax guyanensis 

Styrax jauaensis Steyerm. =Styrax guanayanus 

Styrax lauraceus Perkins =Styrax glabratus 

Styrax micrasterus Perkins =Styrax glaber 

Styrax occidentalis Sw. ex Thunb. =Styrax glaber 

Styrax psilophyllus A. DC. =Styrax glabratus 

Styrax roraimae Perkins =Styrax glaber 

Styrax squamulosus M.F. Silva =Styrax glabratus 

Styrax subleprosum Klotzsch [nomen nudum] =Styrax glaber 

Styrax tepuiensis Steyerm. & Maguire =Styrax glaber 

Styrax tepuiensis subsp. guaiquinimae Maguire & Steyerm. =Styrax guaiquinimae 

Styrax tepuiensis subsp. huachamacarii Maguire & Steyerm. =Styrax longipedicellatus 

REFERENCES 

Fritsch, P.W., S.A. Mori, and J.L. Brown. 2002. Styracaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, 

M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New 

York Bot. Gard. 76 (2): 706-708. 

P.W. Fritsch & J.A. Steyermark. 2005. Styracaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Maguire, B. & Yung-Chau Huang. 1978. Styracaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 10. 


Perkins, J. 1907. Styracaceae. In A. Engler, Das Pflanzenreich IV(241): 1-111. 

Suriana maritima L. 

SURIANACEAE 

REFERENCES 

Howard, R.A. 1988. Surianaceae. In R.A. Howard, Flora of the Lesser Antilles 4(1): 574-575. 

Symplocos acananensis Steyerm. 

Symplocos bolivarana Aristeg. 

Symplocos chimantensis Steyerm. & Maguire 

SYMPLOCACEAE 

P.E. Berry, D. Sabatier, J.A. Steyermark† 

BO 

AM BO 


GU 

SU 

GU FG 

AM 

AM 

GU 

FG 

FG 

AM 

GU? SU 

GU 

BO 

AM 

BO 

BO 

BO 

FG?

Symplocos guianensis (Aubl.) Gürke 

Symplocos jauaensis Steyerm. & Maguire 

Symplocos martinicensis Jacq. 

Symplocos neblinae Maguire & Steyerm. 

Symplocos nitens (Pohl) Benth. 

Symplocos nitens var. claussenii Brand 

Symplocos pilosiuscula Brand 

Symplocos pycnophylla Sleumer [Mt. Roraima] 

Symplocos schomburgkii Klotzsch ex Brand [Mt. Roraima] 

Symplocos ulei Brand 

Symplocos yapacanensis Steyerm. 



Ciponima guianensis Aubl. =Symplocos guianensis 

Stemmatosiphon nitens Pohl =Symplocos nitens 

Symplocos roraimensis Steyerm. =Symplocos pycnophylla 

REFERENCES 

Brand, A. 1901. Symplocaceae. In A. Engler, Das Pflanzenreich IV(242): 1-100. 

Maguire, B. & Yung-Chau Huang. 1978. Symplocaceae. In B. Maguire & collaborators, Botany of the Guayana Highland. Mem. 

New York Bot. Gard. 29: 223-230. 

Mori, S.A. and J.L. Brown. 2002. Symplocaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & 


Gard. 76 (2): 708-709. 

Steyermark, J.A. & P.E. Berry. 2005. Symplocaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


TEPUIANTHACEAE 

J.S. Miller 

Tepuianthus auyantepuiensis Maguire & Steyerm. 

Tepuianthus sarisarinamensis Maguire & Steyerm. 

Tepuianthus sarisarinamensis subsp. duidensis Maguire & Steyerm. 

Tepuianthus sarisarinamensis subsp. sarisarinamensis 

Tepuianthus savannensis Maguire & Steyerm. 

Tepuianthus yapacanensis Maguire & Steyerm. 

REFERENCES 

Maguire, B. & J.A. Steyermark. 1981. Tepuianthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part XI. 


Miller, J.S. 2005. Tepuianthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 297-299. 

Freziera calophylla Triana & Planch. 

Freziera carinata A.L. Weitzman 

Freziera roraimensis Tul. 

Ternstroemia brevistyla Kobuski 

Ternstroemia browniana Kobuski [expected in BO] 

Ternstroemia campinicola B.M. Boom 

Ternstroemia candolleana Wawra 

Ternstroemia carinata A.L. Weitzman [expected in GU] 

Ternstroemia crassifolia Benth. 

Ternstroemia dehiscens Huber 

Ternstroemia delicatula Choisy [expected in BO] 

Ternstroemia dentata (Aubl.) Sw. [expected in BO] 

Ternstroemia discoidea Gleason 

Ternstroemia distyla Kobuski 

Ternstroemia duidae Gleason 

Ternstroemia dura Gleason 

Ternstroemia gleasoniana Kobuski [expected in BO] 

Ternstroemia grandiosa Kobuski [expected in BO] 

TERNSTROEMIACEAE 

P.E. Berry, A.L. Weitzman 

AM GU SU FG 

AM BO 

BO GU FG 

AM 

AM 

AM 

GU FG 

AM BO 

BO GU 

BO GU 

AM 

AM BO 

AM BO 

AM 

BO 

AM 

AM 

AM 

AM BO 

BO GU 

BO 

GU SU FG? 

AM 

AM GU SU 

BO GU 

GU SU 

FG? 

FG 

GU SU FG 

AM 

AM 

AM 

AM 

BO GU 

GU 

GU

536 

Ternstroemia guanchezii B.M. Boom 

Ternstroemia laevigata Wawra 

Ternstroemia longipes Klotzsch ex Wawra [expected in BO] 

Ternstroemia maguirei B.M. Boom 

Ternstroemia punctata (Aubl.) Sw. 

Ternstroemia pungens Gleason 

Ternstroemia retusifolia Kobuski 

Ternstroemia roraimae Klotsch [Mt. Roraima] 

Ternstroemia schomburgkiana Benth. [expected in BO] 

Ternstroemia sp. A 

Ternstroemia sp. B 

Ternstroemia tristyla Gleason 

Ternstroemia verticillata Klotzsch ex Wawra 



Eroteum calophyllum (Triana & Planch.) Kuntze =Freziera calophylla 

Eroteum guianense (Klotzsch ex Wawra) Kuntze =Freziera roraimensis 

Eurya calophylla (Triana & Planch.) Szyszyl. =Freziera calophylla 

Eurya guianensis (Klotzsch ex Wawra) Szyszyl. =Freziera roraimensis 

Eurya roraimensis (Tul.) Szyszyl. =Freziera roraimensis 

Freziera cuatrecasasii Kobuski =Freziera calophylla 

Freziera esmeraldana Little =Freziera calophylla 

Freziera guianensis Klotzsch ex Wawra =Freziera roraimensis 

Freziera hieronymi Kobuski =Freziera calophylla 

Lettsomia guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Freziera roraimensis 

Taonabo dentata Aubl. =Ternstroemia dentata 

Taonabo punctata Aubl. =Ternstroemia punctata 

Ternstroemia candolleana var. rotundata Wawra =Ternstroemia candolleana 

Ternstroemia cayennensis Planch. [nomen nudum] 

Ternstroemia circumscissilis Kobuski [not in flora area] 

Ternstroemia corymbosa Sm. [name of uncertain application, not Ternstroemia] 

Ternstroemia duidae f. latifolia Kobuski =Ternstroemia duidae 

Ternstroemia monosperma Gleason =Ternstroemia discoidea 

Ternstroemia paucifolia Gleason =Ternstroemia duidae 

Ternstroemia revoluta Splitg. =Ternstroemia punctata 

REFERENCES 

Berry, P.E. & A.L. Weitzman. 2005. Ternstroemiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Jansen-Jacobs, M.J. & J.C. Lindeman. 1979. Theaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 356-366. 

Kobuski, C.E. 1942. Studies on the Theaceae, XII. Notes on the South American species of Ternstroemia. J. Arn. Arb. 23: 298- 

343. 

Weitzman, A.L., S. Dressler & P.F. Stevens. 2004. Ternstroemiaceae. Families and Genera of Vascular Plants 6: 450-460. 

Pentamerista neotropica Maguire 

TETRAMERISTACEAE 


REFERENCES 

Miller, J.S. & P.E. Berry. 2005. Tetrameristaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Gordonia fruticosa (Schrad.) H. Keng 

THEACEAE 

A.L. Weitzman 


Gordonia semiserrata (Nees) Spreng. =Gordonia fruticosa 

Haemocharis caracasana Linden & Planch. =Gordonia fruticosa 

Haemocharis pubescens (Planch. & Linden) Linden & Planch. =Gordonia fruticosa 

Laplacea camelliaefolia Triana & Planch. =Gordonia fruticosa 

Laplacea caracasana Klotzsch & H. Karst. ex Wawra [nom. illeg.] =Gordonia fruticosa 

Laplacea fruticosa (Schrad.) Kobuski =Gordonia fruticosa 

AM 

AM 

BO GU 

GU 

BO GU SU FG 

AM BO SU 

BO 

GU? 

GU SU 

AM 

AM 

AM 

BO GU SU 

AM 

AM BO GU SU FG


Laplacea fruticosa var. chimantae Steyerm. =Gordonia fruticosa 

Laplacea pubescens Planch. & Linden ex Triana & Planch. =Gordonia fruticosa 

Laplacea pubescens var. camelliaefolia (Triana & Planch.) Kobuski =Gordonia fruticosa 

Laplacea pubescens var. minor Steyerm. =Gordonia fruticosa 

Laplacea semiserrata (Nees) Cambess. =Gordonia fruticosa 

Lindleya ? fruticosa Nees =Gordonia fruticosa 

Lindleya semiserrata Nees [nomen nudum] =Gordonia fruticosa 

Wikstroemia camelliaefolia (Triana & Planch.) S.F. Blake =Gordonia fruticosa 

Wikstroemia fruticosa Schrad. =Gordonia fruticosa 

REFERENCES 

(see also Bonnetiaceae) 

Jansen-Jacobs, M.J. & J.C. Lindeman. 1979. Theaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 356-366. 

Kobuski, C.E. 1942. Studies on the Theaceae, XII. Notes on the South American species of Ternstroemia. J. Arn. Arb. 23: 298- 

343. 

Weitzman, A.L. Theaceae. 2005. Theaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 311-313. 

Clavija imatacae B. StDhl 

Clavija lancifolia Desf. 

Clavija lancifolia subsp. chermontiana (Standl.) B. StDhl 

Clavija lancifolia subsp. lancifolia 

Clavija macrophylla (Link ex Roem. & Schult.) Miq. 

Jacquinia armillaris Jacq. 

THEOPHRASTACEAE 

J.C. Lindeman, J.S. Miller, B. Ståhl 


Clavija chermontiana Standl. =Clavija lancifolia subsp. chermontiana 

Clavija lancifolia auct. non Desf. 1832: sensu Mez =Clavija imatacae 

Jacquinia arborea auct. non Vahl: sensu Kunth =Jacquinia armillaris 

Jacquinia revoluta Jacq. =Jacquinia armillaris 

REFERENCES 

Lindeman, J.C. 1979. Theophrastaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 367-369. 

Miller, J.S. & B. Ståhl. 2005. Theophrastaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Ståhl, B. 1991. A revision of Clavija. Opera Bot. 107: 1-77. 

Ståhl, B. 2002. Theophrastaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, 


709-711. 

THYMELAEACEAE 

P.E. Berry, Z.S. Rogers, J.A. Steyermark† 

Daphnopsis americana (Mill.) J.R. Johnst. 

Daphnopsis americana subsp. caribaea (Griseb.) Nevling 

Daphnopsis dircoides Steyerm. 

Daphnopsis granitica Pruski & Barringer 

Daphnopsis guaiquinimae Steyerm. 

Daphnopsis longifolia Taub. 

Daphnopsis longipedunculata Gilg ex Ule & Domke [=Funifera sp.?] [Mt. 

Roraima] 

Daphnopsis nevlingiana Steyerm. 

Daphnopsis steyermarkii Nevling 

Goodallia guianensis Benth. 

Lasiadenia ottohuberi Plowman & Nevling 

Lasiadenia rupestris Benth. 

Lophostoma amoenum Nevling 

Schoenobiblus amazonicus Steyerm. 

Schoenobiblus daphnoides Mart. 

Schoenobiblus panamensis Standl. & L.O. Williams 

BO DA GU 



GU SU FG 

GU 

BO 

BO 

BO 

BO 

BO 

GU? 

BO GU 

BO 

AM BO GU 

GU 

AM 

AM BO GU 

AM 

AM 

FG 

FG? 

FG

538 



Daphnopsis caribea Griseb. =Daphnopsis americana subsp. caribaea 

Laurus americanus Mill. =Daphnopsis americana 

REFERENCES 

Domke, W. 1934. Untersuchungen über die systematische und geographische Gliederung der Thymelaeaceen. Bibliotheca Botanica 

111: 1-151. 

Kawasaki, M.L. 2002. Thymelaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 711-712. 

Nevling, L.I. 1959. A revision of the genus Daphnopsis. Ann. Missouri Bot. Gard. 46: 257-358. 

Rogers, Z.S., P.E. Berry, & J.A. Steyermark. 2005. Thymelaeaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. 

Holst, Flora of the Venezuelan Guayana 9: 332-342. 

TILIACEAE 

L.J. Dorr, M.J. Jansen-Jacobs, W. Meijer† 

Apeiba albiflora Ducke 

Apeiba glabra Aubl. 

Apeiba intermedia Uittien 

Apeiba macropetala Ducke 

Apeiba membranacea Spruce ex Benth. 

Apeiba petoumo Aubl. 

Apeiba schomburgkii Szyszyl. 

Apeiba tibourbou Aubl. 

Apeiba uittienii Jans.-Jac. & Westra 

Christiana africana DC. 

Christiana mennegae (Jans.-Jac. & Westra) Kubitzki 

Corchorus aestuans L. 

Corchorus hirtus L. 

Corchorus orinocensis Kunth 

Corchorus siliquosus L. 

Luehea alternifolia (Mill.) Mabb. 

Luehea candida (DC.) Mart. 

Luehea cymulosa Spruce ex Benth. 

Luehea paniculata Mart. & Zucc. 

Lueheopsis althaeiflora (Spruce ex Benth.) Burret 

Lueheopsis duckeana Burret 

Lueheopsis rosea (Ducke) Burret 

Lueheopsis rugosa (Pulle) Burret 

Mollia glabrescens Benth. 

Mollia globularis Meijer 

Mollia grandiflora Meijer 

Mollia lepidota Spruce ex Benth. 

Mollia lepidota subsp. casiquiarensis (Baehni) Meijer 

Mollia lepidota subsp. sphaerocarpa (Gleason ex Record) Meijer 

Mollia sp. A 

Mollia speciosa Mart. 

Mollia tomentosa Spruce ex Benth. 

Mollia ulei Burret 

Triumfetta althaeoides Lam. 

Triumfetta bogotensis DC. 

Triumfetta lappula L. 

Triumfetta rhomboidea Jacq. 

Triumfetta semitriloba Jacq. 

Vasivaea alchorneoides Baill. 


Alegria candida DC. =Luehea candida 

Apeiba aspera Aubl. =Apeiba glabra 

Apeiba burchellii Sprague =Apeiba glabra 

Apeiba echinata Gaertn. =Apeiba petoumo 

Apeiba echinata var. macropetala (Ducke) Ducke =Apeiba macropetala 

Apeiba hypoleuca Steud. =Apeiba petoumo 

Apeiba surinamensis Uittien =Apeiba schomburgkii 


AM SU FG 

SU 

AM BO GU 

AM BO GU 


AM BO GU SU 


AM BO 

BO GU 

SU 

BO GU SU FG 

BO GU SU 

AM BO GU 

GU 

BO GU SU FG 

BO 

AM 

SU 

AM 

BO 

AM GU SU FG 

GU SU FG 

AM BO GU 

GU 

AM 

AM GU 

AM 

GU 

AM 

AM GU 

AM 

AM BO GU 

GU SU FG 

BO 

BO SU FG 

BO GU FG 

BO GU SU? 

AM BO GU


Apeiba tibourbou var. krukofii Uittien =Apeiba albiflora 

Apeiba tibourbou var. membranacea Lockh. ex Griseb. =Apeiba schomburgkii 

Apeiba tibourbou var. rugosa Szyszyl. =Apeiba albiflora 

Asterophorum mennegae Jans.-Jac. & Westra =Christiana mennegae 

Bartramia indica L. =Triumfetta rhomboidea 

Carpodiptera ? schomburgkii Baill. =Christiana africana 

Cedrus alternifolia Mill. =Luehea alternifolia 

Christiana cordifolia Hook. f. =Christiana africana 

Christiana madagascariensis Baill. =Christiana africana 

Corchorus acutangulus Lam. =Corchorus aestuans 

Corchorus argutus Kunth =Corchorus orinocensis 

Corchorus argutus var. benthamii K. Schum. =Corchorus orinocensis 

Corchorus hirtus var. orinocensis (Kunth) K. Schum. =Corchorus orinocensis 

Corchorus pilolobus Link =Corchorus hirtus 

Guenetia macrosperma Sagot (Bombacaceae) =Catostemma fragrans 

Luehea althaeiflora Spruce ex Benth. =Lueheopsis althaeiflora 

Luehea ferruginea Turcz. =Luehea alternifolia 

Luehea flavescens Uittien =Lueheopsis rosea 

Luehea maroniensis Benoist =Lueheopsis rugosa 

Luehea nobilis Linden & Planch. =Luehea candida 

Luehea rosea Ducke =Lueheopsis rosea 

Luehea speciosa Willd. =Luehea alternifolia 

Luehea tessmannii Burret =Luehea cymulosa 

Lueheopsis flavescens (Uittien) Burret =Lueheopsis rosea 

Lueheopsis violacea Standl. =Lueheopsis duckeana 

Mollia lepidota var. casiquiarensis Baehni =Mollia lepidota subsp. casiquiarensis 

Mollia scabra Meijer =Mollia ulei 

Mollia speciosa var. alata Meijer [nomen nudum] =Mollia sp. A 

Mollia sphaerocarpa Gleason ex Record =Mollia lepidota subsp. sphaerocarpa 

Mollia stellaris Meijer =Mollia ulei 

Speirostyla tiliifolia Baker =Christiana africana 

Triumfetta bartramia L. [nom. illeg.] =Triumfetta rhomboidea 

Triumfetta bartramia sensu Lay, non L. =Triumfetta rhomboidea 

Triumfetta bolivariensis D.E. González [nomen nudum] =Triumfetta bogotensis 

Triumfetta hispida A. Rich. =Triumfetta bogotensis 

Triumfetta hostmannii Miq. =Triumfetta lappula 

Triumfetta macrophylla Vahl =Triumfetta althaeoides 

Triumfetta rubricaulis Kunth =Triumfetta semitriloba 

Triumfetta sinuosa Miq. =Triumfetta lappula 

Triumfetta surinamensis Steud. =Triumfetta althaeoides 

Triumfetta valenciensis R. Knuth [nomen nudum] =Triumfetta bogotensis 

REFERENCES 

Dorr, L.J. 2002. Tiliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


716. 

Dorr, L.J. & W. Meijer. 2005. Tiliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 343-362. 

Jansen-Jacobs, M.J. 1986. Tiliaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 3(1-2): 

293-300. 

Jansen-Jacobs, M.J. & W. Meijer (in press). Tiliaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A). 

Jansen-Jacobs, M.J. & L.Y.Th. Westra. 1995. A new species of Apeiba (Tiliaceae) from the Venezuelan-Brazilian border. Brittonia 

47(3): 335-339. 

Ko Ko Lay 1950. The American species of Triumfetta L. Ann. Missouri Bot. Gard. 37(3): 315-395. 

Kubitzki, K. 1995. Asterophorum and Tahitia congeneric with Christiana (Tiliaceae). Bot. Jahrb. Syst. 116: 537-542. 

Meijer, W. 1978. Mollia (Tiliaceae). A revision of the South American species. In B. Maguire & collaborators, Botany of the 

Guayana Highland - Part X. Mem. New York Bot. Gard. 29: 68-82. 

Uittien, H. 1932. Tiliaceae. In A. Pulle, Flora of Suriname 3(1): 49-57. 

Uittien, H. 1941. Tiliaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 438-441. 

TRIGONIACEAE 

P.E. Berry, E. Lleras-Pérez 

Trigonia candelabra Lleras SU

540 

Trigonia coppenamensis Stafleu 

Trigonia ehrendorferi Lleras 

Trigonia hypoleuca Griseb. 

Trigonia laevis Aubl. 

Trigonia microcarpa Sagot ex Warm. 

Trigonia nivea Cambess. 

Trigonia nivea var. nivea 

Trigonia reticulata Lleras 

Trigonia sericea Kunth 

Trigonia spruceana Benth. ex Warm. 

Trigonia subcymosa Benth. 

Trigonia villosa Aubl. 

Trigonia villosa var. macrocarpa (Benth.) Lleras 

Trigonia villosa var. villosa 



Trigonia hypoleuca var. pubescens Warm. =Trigonia hypoleuca 

Trigonia kaieteurensis Maguire =Trigonia microcarpa 

Trigonia laevis var. microcarpa (Sagot ex Warm.) Sagot =Trigonia microcarpa 

Trigonia macrocarpa Benth. =Trigonia villosa var. macrocarpa 

Trigonia villosa var. cuneata DC. =Trigonia villosa var. villosa 

Trigonia villosa var. oblonga DC. =Trigonia villosa var. villosa 

Trigonia villosa var. obtusata DC. =Trigonia villosa var. villosa 

Trigonia xanthopila Garcke =Trigonia hypoleuca 

REFERENCES 

Berry, P.E. 2005. Trigoniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 364-368. 

Lleras, E. 1978. Trigoniaceae. Flora Neotropica Monogr. 19: 1-73. 

Lleras, E. 1998. Trigoniaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 21: 49-60. 

Mori, S.A. 2002. Trigoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


718. 

Stafleu, F.A. 1951. Trigoniaceae. In A. Pulle, Flora of Suriname 3(2): 173-177. 

Tropaeolum orinocense P.E. Berry 

TROPAEOLACEAE 

P.E. Berry, L.J. Raechal 

REFERENCES 

Bayer, C. & O. Appel. 2003. Families and Genera of Vascular Plants. 5: 400—404. 

Berry, P.E. & L.J. Raechal. Tropaeolaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 372. 

Piriqueta cistoides (L.) Griseb. 

Piriqueta cistoides subsp. caroliniana (Walter) Arbo 

Piriqueta cistoides subsp. cistoides 

Piriqueta guianensis N.E. Br. 

Piriqueta guianensis subsp. guianensis 

Piriqueta undulata Urb. 

Piriqueta viscosa Griseb. 

Piriqueta viscosa subsp. tovarensis Urb. 

Piriqueta viscosa subsp. viscosa 

Turnera acuta Willd. ex Schult. 

Turnera annectens Arbo 

Turnera argentea Arbo 

Turnera aurantiaca Benth. 

Turnera brasiliensis Willd. ex Roem. & Schult. 

Turnera breviflora Moura 

TURNERACEAE 

C. Feuillet, M. Hoff, M.-M. Arbo 

SU 

BO 

GU SU FG 

GU FG 


AM BO GU 

AM BO GU 

BO GU 

AM 

AM 

BO 

GU 

BO GU SU FG 

GU FG? 

BO GU SU FG 

AM 


AM BO 


GU 

GU 

AM BO 


BO 


AM BO 

AM 

AM 

GU SU FG 

GU SU FG 

BO GU

Turnera caerulea Moç. & Sessé ex DC. 

Turnera caerulea var. surinamensis (Urb.) Arbo & A. Fernández 

Turnera castilloi Arbo 

Turnera cicatricosa Arbo 

Turnera curassavica Urb. 

Turnera glaziovii Urb. 

Turnera guianensis Aubl. 

Turnera huberi Arbo 

Turnera lineata Urb. 

Turnera macrophylla Urb. 

Turnera odorata Rich. 

Turnera paruana Arbo 

Turnera pumilea L. 

Turnera rupestris Aubl. 

Turnera rupestris var. frutescens (Aubl.) Urb. 

Turnera rupestris var. rupestris 

Turnera scabra Millsp. 

Turnera schomburgkiana Urb. 

Turnera steyermarkii Arbo 

Turnera subulata Sm. 

Turnera velutina Benth. 

Turnera venosa Urb. 

Turnera waltherioides Urb. 

Wormskioldia lobata Urb. [cultivated?] 



Piriqueta caroliniana (Walter) Urb. =Piriqueta cistoides subsp. caroliniana 

Piriqueta caroliniana var. integrifolia Urb. =Piriqueta cistoides subsp. caroliniana 

Piriqueta cistoides var. bracteolata Urb. =Piriqueta cistoides subsp. cistoides 

Piriqueta cistoides var. foliosa (Garcke) Urb. =Piriqueta cistoides subsp. cistoides 

Piriqueta cistoides var. genuina Urb. =Piriqueta cistoides subsp. cistoides 

Piriqueta cistoides var. latifolia Urb. =Piriqueta cistoides subsp. cistoides 

Piriqueta foliosa Garcke =Piriqueta cistoides subsp. cistoides 

Piriqueta lanceolata Benth. =Piriqueta cistoides subsp. cistoides 

Piriqueta stenophylla Klotzsch =Piriqueta cistoides subsp. cistoides 

Piriqueta tomentosa Kunth =Piriqueta cistoides subsp. cistoides 

Piriqueta villosa Aubl. =Piriqueta cistoides subsp. cistoides 

Turnera aspera Poir. =Piriqueta cistoides subsp. cistoides 

Turnera aturensis D. Dietr. =Piriqueta cistoides subsp. cistoides 

Turnera benthamiana R.H. Schomb. =Turnera velutina 

Turnera carpinifolia Kunth =Turnera acuta 

Turnera chamaedrys Klotzsch [excluded taxon] 

Turnera cistoides L. =Piriqueta cistoides 

Turnera corchorifolia Willd. ex Schult. =Turnera odorata 

Turnera corchoroides Klotzsch [excluded taxon] 

Turnera desvauxii DC. =Piriqueta cistoides subsp. cistoides 

Turnera frutescens Aubl. =Turnera rupestris var. frutescens 

Turnera grandifolia Bremek. =Turnera glaziovii 

Turnera hexandra Spreng. =Turnera odorata 

Turnera hirta Desv., non Willd. =Piriqueta cistoides subsp. cistoides 

Turnera humboldtii Spreng. =Turnera guianensis 

Turnera muricata Rusby =Turnera macrophylla 

Turnera opifera Mart. =Turnera caerulea var. surinamensis 

Turnera parviflora Benth. =Turnera odorata 

Turnera pilosula Urb. =Turnera acuta 

Turnera refracta Klotzsch =Turnera caerulea var. surinamensis 

Turnera subglabra Klotzsch =Turnera aurantiaca 

Turnera tomentosa Willd. ex Schult. =Piriqueta cistoides subsp. cistoides 

Turnera tomentosa Kunth =Turnera odorata 

Turnera ulmifolia var. elegans (Otto) Urb. =Turnera subulata 

Turnera ulmifolia var. intermedia Urb. =Turnera scabra 

Turnera ulmifolia f. serissima Killip =Turnera lineata 

Turnera ulmifolia var. surinamensis Urb. =Turnera caerulea var. surinamensis 

Waltheria caroliniana Walter =Piriqueta cistoides subsp. caroliniana 

GU SU 

GU SU 

AM 

AM BO GU 

GU? 

GU? SU FG 

AM 

AM 

BO GU SU FG 

AM 

AM 

BO GU 

AM 

AM 

BO GU SU FG 

BO GU FG 

GU SU FG 

GU SU FG 

GU SU FG 

AM BO GU FG? 

AM 

BO GU 

AM 

GU 

GU 

SU FG 

BO GU 

FG

542 


REFERENCES 

Arbo, M.M. 1990. Turneraceae: novedades para la Guayana Venezolana. Ann. Missouri Bot. Gard. 77: 340-352. 

Arbo, M.M. 1995. Turneraceae - Parte I. Piriqueta. Flora Neotropica Monogr. 67: 1-156. 

Berry, P.E. & L.J. Raechal. 2005. Turneraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Bremekamp, E. 1941. Turneraceae. In A. Pulle, Flora of Suriname 3(1): 342-354. 

Hoff, M. 2002. Turneraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


719. 

Lemée, A. 1953. Flore de la Guyane française 3: 82. Paul Lechevalier, Paris. 

Pulle, A. 1906. An enumeration of the Vascular Plants known from Surinam, together with their distribution and Synonymy. 

Leiden. 

Urban, I. 1883a. Monographie des Familie der Turneraceen. Jahrbuch des Königlichen Botanischen Gartens und des botanischen 

Museums zu Berlin, 2: 1-152. 

Urban, I. 1883b. Turneraceae. In C.F.P. Martius, Flora Brasiliensis 13(3): 88-170, tab. 31-48. 

Ampelocera edentula Kuhlm. 

Celtis iguanaea (Jacq.) Sarg. 

Celtis schippii Standl. 

Trema integerrima (Beurl.) Standl. 

Trema micrantha (L.) Blume 

Rhamnus iguanaeus Jacq. =Celtis iguanaea 

Rhamnus micranthus L. =Trema micrantha 

Sponia integerrima Beurl. =Trema integerrima 

Sponia micrantha (L.) Decne. =Trema micrantha 

ULMACEAE 

C.C. Berg, P.E. Berry, J.S. Miller 


REFERENCES 

Berg, C.C. 1992. Ulmaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 11: 3-9. 

Berg, C.C. 2002. Ulmaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


722. 

Miller, J.S. & P.E. Berry. 2005. Ulmaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 386-390. 

Ooststroom, S.J. van. 1932. Ulmaceae. In A. Pulle, Flora of Suriname 1(1): 47-48. 

Boehmeria cylindrica (L.) Sw. 

Boehmeria ramiflora Jacq. 

Laportea aestuans (L.) Chew 

Phenax sonneratii (Poir.) Wedd. [naturalized?] 

Pilea dauciodora Pav. ex Wedd. 

Pilea fendleri Killip 

Pilea imparifolia Wedd. 

Pilea involucrata (Sims) Urb. 

Pilea microphylla (L.) Liebm. 

Pilea pubescens Liebm. 

Pilea tabularis C.C. Berg 

Pouzolzia occidentalis (Liebm.) Wedd. 

Urera baccifera (L.) Gaudich. ex Wedd. 

Urera caracasana (Jacq.) Gaudich. ex Griseb. 

Urera laciniata Goudot ex Wedd. 

URTICACEAE 

C.C. Berg, A. Pool 


Fleurya aestuans (L.) Gaudich. ex Miq. =Laportea aestuans 

Leucococcus occidentalis Liebm. =Pouzolzia occidentalis 

Parietaria microphylla L. =Pilea microphylla 

Parietaria sonneratii Poir. =Phenax sonneratii 



GU 

AM GU SU 


DA SU FG 

SU FG 

DA GU SU FG 

DA FG 

AM 

DA 

GU SU FG 

BO DA 

BO GU SU FG 

AM BO 

BO 

GU SU FG 

FG 



AM SU FG


Phenax vulgaris Wedd. =Phenax sonneratii 

Pilea chrysosplenioides Wedd. =Pilea involucrata 

Pilea dauciodora var. crenata Wedd. =Pilea fendleri 

Pilea dendrophila Miq. [not in flora area] 

Pilea guyanensis Wedd. =Pilea pubescens 

Pilea leptophylla Killip =Pilea fendleri 

Pilea pubescens var. guyanensis (Wedd.) Wedd. =Pilea pubescens 

Pilea pubescens var. involucrata (Sims) Wedd. =Pilea involucrata 

Urtica aestuans L. =Laportea aestuans 

Urtica alceifolia Poir. =Urera caracasana 

Urtica baccifera L. =Urera baccifera 

Urtica caracasana Jacq. =Urera caracasana 

Urtica cylindrica L. =Boehmeria cylindrica 

Urtica involucrata Sims =Pilea involucrata 

REFERENCES 

Berg, C.C. 1992. Urticaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 11: 125-139. 

Berg, C.C. 2002. Urticaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 


725. 

Pool, A. 2005. Urticaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 9: 

391-402. 

Rooij, M.J.M. de. 1975. Urticaceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 300-318. 

VALERIANACEAE 

N.X. de Enrich 

Valeriana scandens L. 

Valeriana scandens var. candolleana (Gardner) C.A. Müll. [Mt. Roraima] 

Valeriana scandens var. subcordata C.A. Müll. 


Valeriana alpina Vell. =Valeriana scandens var. candolleana 

Valeriana candolleana Gardner =Valeriana scandens var. candolleana 

Valeriana phaseoli Braun =Valeriana scandens 

Valeriana scandens var. dentata C.A. Müll. =Valeriana scandens var. candolleana 

REFERENCES 

Xena de Enrech, N. 1992. Valerianaceae. In G. Morillo, Flora de Venezuela 5(1): 221-267. 

Xena de Enrech, N. 2005. Valerianaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 402-404. 

Aegiphila bracteolosa Moldenke 

Aegiphila deppeana Steud. 

Aegiphila elata Sw. 

Aegiphila elata var. elata 

Aegiphila elata var. macrophylla (Kunth) López-Pal. 

Aegiphila filipes Mart. & Schauer ex Schauer 

Aegiphila glandulifera Moldenke 

Aegiphila glandulifera var. pyramidata Moldenke 

Aegiphila gleasonii Moldenke [insufficiently known taxon] 

Aegiphila hoehnei Moldenke 

Aegiphila hoehnei var. venezuelensis Moldenke 

Aegiphila hystricina Aymard & Cuello 

Aegiphila integrifolia (Jacq.) B.D. Jacks. 

Aegiphila intermedia Moldenke 

Aegiphila laevis (Aubl.) J.F. Gmel. 

Aegiphila laxiflora Benth. 

Aegiphila lewisiana Moldenke 

Aegiphila lhotzkiana Cham. 

Aegiphila longifolia Turcz. 

VERBENACEAE 

G.A. Aymard C., M.J. Jansen-Jacobs 

AM BO 

AM BO 

BO 

AM GU 

FG 

AM BO GU SU 

AM 

BO GU SU 

AM BO 

AM BO 

AM 

AM 

AM 

BO 

GU 

AM BO 

AM 

AM 



BO 

BO 

GU 

GU 

SU FG

544 


Aegiphila macrantha Ducke 

Aegiphila membranacea Turcz. 

Aegiphila mollis Kunth 

Aegiphila mollis var. intermedia Moldenke 

Aegiphila mollis var. mollis 

Aegiphila mollis var. surfaceana (Moldenke) Moldenke 

Aegiphila parviflora Moldenke 

Aegiphila pendula Moldenke 

Aegiphila perplexa Moldenke 

Aegiphila racemosa Vell. 

Aegiphila roraimensis Moldenke 

Aegiphila spruceana Moldenke 

Aegiphila venezuelensis Moldenke 

Aegiphila venezuelensis var. serrata Moldenke 

Aegiphila venezuelensis var. venezuelensis 

Aegiphila villosa (Aubl.) J.F. Gmel. 

Aegiphila vitelliniflora Klotzsch ex Walp. 

Aegiphila vitelliniflora var. egleri Moldenke [insufficiently known taxon] 

Amasonia campestris (Aubl.) Moldenke 

Amasonia obovata Gleason 

Avicennia germinans (L.) L. 

Avicennia schaueriana Stapf & Leechm. ex Moldenke 

Bouchea prismatica (L.) Kuntze 

Bouchea prismatica var. longirostra Grenzeb. 

Citharexylum macrophyllum Poir. 

Citharexylum poeppigii Walp. 

Citharexylum sp. A 

Citharexylum spinosum L. 

Citharexylum venezuelense Moldenke 

Clerodendrum aculeatum (L.) Schltdl. 

Clerodendrum indicum (L.) Kuntze [cultivated, naturalized] 

Clerodendrum japonicum (Thunb.) Sweet [cultivated, naturalized] 

Clerodendrum philippinum Schauer [widely cultivated] 

Clerodendrum ternifolium Kunth 

Clerodendrum thomsonae Balf. [cultivated, naturalized] 

Cornutia pubescens C.F. Gaertn. 

Cornutia pyramidata L. 

Duranta erecta L. 

Duranta obtusifolia Kunth 

Duranta sp. A 

Duranta sp. B 

Lantana achyranthifolia Desf. 

Lantana brasiliensis Link [expected in GU] 

Lantana camara L. 

Lantana canescens Kunth 

Lantana cujabensis Schauer 

Lantana fucata Lindl. 

Lantana glutinosa Poepp. 

Lantana radula Sw. 

Lantana ruiz-teranii López-Pal. & Steyerm. 

Lantana trifolia L. 

Lippia alba (Mill.) N.E. Br. 

Lippia aristata Schauer 

Lippia micromera Schauer 

Lippia origanoides Kunth 

Petrea blanchetiana Schauer 

Petrea bracteata Steud. 

Petrea macrostachya Benth. 

Petrea pubescens Turcz. 

Petrea sulphurea Jans.-Jac. 

Petrea volubilis L. 

Phyla betulifolia (Kunth) Greene 

Priva lappulacea (L.) Pers. 

Stachytarpheta angustifolia (Mill.) Vahl 



AM BO 

AM BO 

AM BO 

AM BO 

BO 

AM 

DA 

BO GU SU FG 

AM BO GU 

AM 

AM BO GU 

AM BO GU 

AM BO GU 

GU FG 

FG? 

FG? 


AM 

DA GU SU FG 

DA GU SU FG 

BO 

BO 

DA GU SU FG 

BO DA 

AM 


BO 

GU SU FG 

GU SU FG 

GU SU 


BO 


FG 

SU 


BO 

BO 

AM 

AM BO 


BO DA GU 

AM BO 

BO 

BO DA 

BO GU 

BO 



GU 

AM BO DA GU 

BO GU 

AM BO FG 


AM GU SU FG 

AM BO 

FG 




BO GU SU FG

Stachytarpheta cayennensis (Rich.) Vahl 

Stachytarpheta jamaicensis (L.) Vahl 

Stachytarpheta lopez-palacii Moldenke 

Stachytarpheta mutabilis (Jacq.) Vahl 

Stachytarpheta sprucei Moldenke 

Tamonea spicata Aubl. 

Vitex agnus-castus L. [cultivated, naturalized] 

Vitex calothyrsa Sandwith 

Vitex capitata Vahl 

Vitex compressa Turcz. 

Vitex guianensis Moldenke 

Vitex klugii Moldenke 

Vitex orinocensis Kunth 

Vitex orinocensis var. multiflora (Miq.) Huber 

Vitex orinocensis var. orinocensis 

Vitex panshiniana Moldenke 

Vitex schomburgkiana Schauer 

Vitex sprucei Briq. 

Vitex stahelii Moldenke 

Vitex triflora Vahl 



Aegiphila cowanii Moldenke =Aegiphila venezuelensis 

Aegiphila guianensis Moldenke =Aegiphila integrifolia 

Aegiphila integrifolia var. guianensis (Moldenke) López-Pal. =Aegiphila integrifolia 

Aegiphila lutea Lam. =Aegiphila laevis 

Aegiphila macrophylla Kunth =Aegiphila elata var. macrophylla 

Aegiphila martinicensis Jacq. [not in flora area] 

Aegiphila steyermarkii Moldenke =Aegiphila roraimensis 

Aegiphila steyermarkii var. macrophylla Moldenke =Aegiphila venezuelensis 

Aegiphila steyermarkii var. obtusifolia Moldenke =Aegiphila roraimensis 

Aegiphila surfaceana Moldenke =Aegiphila mollis var. surfaceana 

Amasonia arborea Kunth =Amasonia campestris 

Amasonia campestris var. surinamensis Moldenke =Amasonia campestris 

Amasonia erecta L. f. =Amasonia campestris 

Amasonia erecta var. punicea (Vahl) Pulle =Amasonia campestris 

Amasonia integerrima Spreng. (Acanthaceae) =Bravaisia integerrima 

Amasonia lasiocaulos Mart. & Schauer ex Schauer =Amasonia campestris 

Amasonia punicea Vahl =Amasonia campestris 

Amasonia spruceana Moldenke =Amasonia campestris 

Avicennia germinans (L.) Stearn [comb. superfl.] =Avicennia germinans 

Avicennia germinans var. cumanensis (Kunth) Moldenke =Avicennia germinans 

Avicennia germinans var. guayaquilensis (Kunth) Moldenke =Avicennia germinans 

Avicennia germinans f. venezuelensis Moldenke =Avicennia germinans 

Avicennia marina (Forssk.) Vierh. [not in flora area] 

Avicennia nitida Jacq. =Avicennia germinans 

Avicennia schaueriana f. candicans Moldenke =Avicennia schaueriana 

Avicennia tomentosa var. cumanensis Kunth =Avicennia germinans 

Avicennia tomentosa var. guayaquilensis Kunth =Avicennia germinans 

Bontia germinans L. =Avicennia germinans 

Callicarpa integrifolia Jacq. =Aegiphila integrifolia 

Citharexylum amazonicum Moldenke =Citharexylum macrophyllum 

Citharexylum fruticosum L. =Citharexylum spinosum 

Citharexylum fruticosum var. brittonii Moldenke =Citharexylum spinosum 

Citharexylum poeppigii f. anomalum Moldenke =Citharexylum poeppigii 

Citharexylum scandens Briq. & Spruce (Bignoniaceae) =Schlegelia scandens 

Clerodendrum aculeatum var. guianense Moldenke =Clerodendrum aculeatum 

Clerodendrum fragrans (Vent.) Willd. =Clerodendrum philippinum 

Clerodendrum fragrans var. pleniflora Schauer =Clerodendrum philippinum 

Clerodendrum fragrans var. pleniflorum Schauer =Clerodendrum philippinum 

Cornutia cayennensis Schauer =Cornutia pubescens 

Cryptocalyx nepetifolia Benth. =Lippia betulifolia 

Duranta coriacea Hayek =Duranta obtusifolia 

Duranta plumieri var. alba Mast. =Duranta erecta 



BO 

BO GU SU FG 

AM BO GU 

BO GU SU FG 

GU SU FG 

AM 

AM BO GU SU 


GU FG 

AM 



AM 

SU FG? 

BO GU 

AM 


AM GU SU FG

546 


Duranta repens L. =Duranta erecta 

Duranta repens var. canescens Moldenke =Duranta erecta 

Duranta repens var. lopez-palacii Moldenke =Duranta erecta 

Ghinia spicata (Aubl.) Moldenke =Tamonea spicata 

Lantana aculeata L. =Lantana camara 

Lantana alba Mill. =Lippia alba 

Lantana armata Schauer =Lantana camara 

Lantana armata var. guianensis Moldenke =Lantana camara 

Lantana camara var. aculeata (L.) Moldenke =Lantana camara 

Lantana camara var. mista (L.) L.H. Bailey =Lantana camara 

Lantana camara var. moritziana (Otto & A. Dietr.) López-Pal. =Lantana camara 

Lantana geminata (Kunth) Spreng. =Lippia alba 

Lantana mista L. =Lantana camara 

Lantana moritziana Otto & A. Dietr. =Lantana camara 

Lantana trifolia f. hirsuta Moldenke =Lantana trifolia 

Lantana trifolia f. oppositifolia Moldenke =Lantana trifolia 

Lantana trifolia var. rigiduscula Briq. =Lantana trifolia 

Lantana trifolia f. rubierensis Moldenke =Lantana trifolia 

Lippia betulifolia Kunth =Phyla betulifolia 

Lippia geminata Kunth =Lippia alba 

Lippia schomburgkiana Schauer =Lippia origanoides 

Lippia stoechadifolia (L.) Kunth [not in flora area] =Phyla stoechadifolia (L.) Small 

Manabea arborescens Aubl. =Aegiphila integrifolia 

Manabea laevis Aubl. =Aegiphila laevis 

Manabea villosa Aubl. =Aegiphila villosa 

Petrea algentryi Moldenke =Petrea blanchetiana 

Petrea arborea Kunth =Petrea volubilis 

Petrea aspera Turcz. =Petrea volubilis 

Petrea glandulosa Pittier =Petrea pubescens 

Petrea kohautiana C. Presl =Petrea volubilis 

Petrea martiana Schauer =Petrea bracteata 

Petrea morii Moldenke =Petrea blanchetiana 

Petrea peruviana Moldenke =Petrea blanchetiana 

Petrea pubescens var. albicalyx Moldenke =Petrea pubescens 

Petrea rivularis Moldenke =Petrea volubilis 

Petrea rugosa Kunth [not in flora area] 

Petrea schomburgkiana Schauer =Petrea bracteata 

Petrea subserrata Cham. [not in flora area] =Petrea racemosa Nees 

Phyla nodiflora (L.) Greene [not in flora area] 

Priva lappulacea f. albiflora Moldenke =Priva lappulacea 

Stachytarpheta angustifolia var. elatior Schrad. ex Schult. =Stachytarpheta angustifolia 

Stachytarpheta angustifolia f. elatior (Schrad. ex Schult.) López-Pal. =Stachytarpheta angustifolia 

Stachytarpheta cayennensis f. albiflora Moldenke =Stachytarpheta cayennensis 

Stachytarpheta dichotoma (Ruiz & Pav.) Vahl =Stachytarpheta cayennensis 

Stachytarpheta dichotoma f. albiflora (Moldenke) Moldenke =Stachytarpheta cayennensis 

Stachytarpheta elatior Schrad. ex Schult. =Stachytarpheta angustifolia 

Stachytarpheta elatior var. jenmanii Moldenke =Stachytarpheta angustifolia 

Stachytarpheta orubica (L.) Vahl [not in flora area] 

Stachytarpheta pilosiuscula Kunth =Stachytarpheta jamaicensis 

Stachytarpheta roraimensis Moldenke =Stachytarpheta sprucei 

Stachytarpheta roraimensis var. pubescens Moldenke =Stachytarpheta sprucei 

Stachytarpheta surinamensis Miq. ex Pulle [nomen nudum] =Stachytarpheta angustifolia 

Taligalea campestris Aubl. =Amasonia campestris 

Verbena angustifolia Mill. =Stachytarpheta angustifolia 

Verbena cayennensis Rich. =Stachytarpheta cayennensis 

Verbena dichotoma Ruiz & Pav. =Stachytarpheta cayennensis 

Verbena jamaicensis L. =Stachytarpheta jamaicensis 

Verbena lappulacea L. =Priva lappulacea 

Verbena mutabilis Jacq. =Stachytarpheta mutabilis 

Verbena prismatica L. =Bouchea prismatica 

Verbena suffruticosa Aubl. [not in flora area] =Phyla stoechadifolia (L.) Small 

Vitex appunii Moldenke =Vitex schomburgkiana 

Vitex brittoniana Moldenke =Vitex compressa 

Vitex multiflora Miq. =Vitex orinocensis var. multiflora


Vitex pacimonensis Spruce ex Sandwith =Vitex calothyrsa 

Vitex spongiocarpa Ducke =Vitex sprucei 

Vitex spongiocarpa var. longidentata Moldenke =Vitex sprucei 

Vitex sprucei var. longidentata Moldenke =Vitex sprucei 

Vitex sprucei var. vaupesensis Moldenke =Vitex sprucei 

Vitex triflora f. quinquefoliolata (Moldenke) Moldenke =Vitex triflora 

Vitex triflora var. quinquefoliolata Moldenke =Vitex triflora 

Vitex wittrockiana Moldenke =Vitex capitata 

Volkameria fragrans Vent. =Clerodendrum philippinum 

REFERENCES 

Aymard C, G.A. 2005. Verbenaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 407-445. 

Jansen-Jacobs, M.J. 1988. Verbenaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 4: 1-114. 

Jansen-Jacobs, M.J. 2002. Verbenaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. 


76 (2): 725-728. 

Lopez-Palacios, S. 1977. Verbenaceae. In Flora de Venezuela. 

Moldenke, H.N. 1940a. Avicenniaceae. In A. Pulle, Flora of Suriname 4(2): 322-325. 

Moldenke, H.N. 1940b. Verbenaceae. In A. Pulle, Flora of Suriname 4(2): 357-321. 

Moldenke, H.N. 1971. A Fifth Summary of the Verbenaceae, Avicenniaceae, Stilbaceae, Dicrostylidaceae, Symphoremaceae, 

Nyctanthaceae, and Eriocaulaceae of the World. H.N. Moldenke, Wayne, N.J. 

Rueda, R.M. 1994. Systematics and evolution of the genus Petrea. Ann. Missouri Bot. Gard. 81(4): 610-652. 

VIOLACEAE 

Á. Fernández del Valle, W.H.A. Hekking 

Amphirrhox latifolia Mart. ex Eichler [=Amphirrhox longifolia s.l.] 

Amphirrhox longifolia (A. St.-Hil.) Spreng. 

Amphirrhox surinamensis Eichler [=Amphirrhox longifolia s.l.] 

Corynostylis arborea (L.) S.F. Blake 

Corynostylis carthagenensis H. Karst. 

Corynostylis pubescens S. Moore 

Corynostylis volubilis L.B. Sm. & A. Fernández 

Gloeospermum sphaerocarpum Triana & Planch. 

Hybanthus calceolaria (L.) Schulze-Menz 

Hybanthus oppositifolius (L.) Taub. 

Hybanthus phyllanthoides (Planch. & Linden) L.B. Sm. & A. Fernández 

Leonia cymosa Mart. 

Leonia glycycarpa Ruiz & Pav. 

Noisettia orchidiflora (Rudge) Ging. 

Paypayrola confertiflora Tul. 

Paypayrola grandiflora Tul. 

Paypayrola guianensis Aubl. 

Paypayrola hulkiana Pulle 

Paypayrola longifolia Tul. 

Rinorea amapensis Hekking 

Rinorea bahiensis (Moric.) Kuntze 

Rinorea brevipes (Benth.) S.F. Blake 

Rinorea camptoneura (Radlk.) Melch. 

Rinorea endotricha Sandwith 

Rinorea falcata (Mart. ex Eichler) Kuntze 

Rinorea flavescens (Aubl.) Kuntze 

Rinorea guianensis Aubl. 

Rinorea lindeniana (Tul.) Kuntze 

Rinorea lindeniana var. lindeniana 

Rinorea macrocarpa (Mart. ex Eichler) Kuntze 

Rinorea melanodonta S.F. Blake 

Rinorea neglecta Sandwith 

Rinorea ovalifolia (Britton) S.F. Blake 

Rinorea paniculata (Mart.) Kuntze 

Rinorea pectino-squamata Hekking 

Rinorea pubiflora (Benth.) Sprague & Sandwith 

Rinorea pubiflora var. grandifolia (Eichler) Hekking 

GU SU FG 


GU SU FG 


BO 

AM 

AM 

AM GU? SU FG 

BO GU SU FG 


BO 

AM BO GU 


GU SU FG 

GU SU FG 

AM GU FG 

AM GU SU FG 

SU FG 


SU FG 

FG 

AM GU SU FG 

AM 

BO DA GU 

AM BO SU FG 


GU FG 

AM BO GU SU 

AM BO GU SU 


BO GU 

GU SU FG 

AM BO 

AM BO FG 

FG 


AM BO DA GU SU FG

548 

Rinorea pubiflora var. pubiflora 

Rinorea racemosa (Mart.) Kuntze 

Rinorea riana Kuntze 

Rinorea sprucei (Eichler) Kuntze 

Rinoreocarpus ulei (Melch.) Ducke 



Alsodeia brevipes Benth. =Rinorea brevipes 

Alsodeia camptoneura Radlk. =Rinorea camptoneura 

Alsodeia falcata Mart. ex Eichler =Rinorea falcata 

Alsodeia falcata var. grandifolia Eichler =Rinorea pubiflora var. grandifolia 

Alsodeia flavescens (Aubl.) Spreng. =Rinorea flavescens 

Alsodeia guianensis (Aubl.) Eichler =Rinorea pubiflora var. pubiflora 

Alsodeia guianensis var. brevipes (Benth.) Eichler =Rinorea brevipes 

Alsodeia guianensis var. laxiflora (Benth.) Eichler =Rinorea brevipes 

Alsodeia guianensis var. lindeniana (Tul.) Eichler =Rinorea lindeniana 

Alsodeia guianensis var. parviflora Eichler =Rinorea falcata 

Alsodeia laxiflora Benth. =Rinorea brevipes 

Alsodeia lindeniana Tul. =Rinorea lindeniana 

Alsodeia longiflora Oudem. =Amphirrhox longiflora 

Alsodeia macrocarpa Mart. ex Eichler =Rinorea macrocarpa 

Alsodeia martini Turcz. =Rinorea pubiflora var. pubiflora 

Alsodeia ovalifolia Britton =Rinorea ovalifolia 

Alsodeia pallida Klotzsch ex M.R. Schomb. [nomen nudum] 

Alsodeia paniculata Mart. =Rinorea paniculata 

Alsodeia piparea Spreng. (Flacourtiaceae) =Casearia commersoniana 

Alsodeia prunifolia Spreng. =Rinorea riana 

Alsodeia pubiflora Benth. =Rinorea pubiflora 

Alsodeia racemosa Mart. =Rinorea racemosa 

Alsodeia riana (DC. ex Ging.) Turcz. =Rinorea riana 

Alsodeia sprucei Eichler =Rinorea sprucei 

Calyptrion nitidum Benth. =Corynostylis arborea 

Conohoria aubletii D. Dietr. (Flacourtiaceae) =Casearia commersoniana 

Conohoria brevipes (Benth.) Miq. =Rinorea brevipes 

Conohoria flavescens Aubl. =Rinorea flavescens 

Conohoria laxiflora (Benth.) Miq. =Rinorea brevipes 

Conohoria passoura DC. ex Ging. =Rinorea pubiflora var. pubiflora 

Conohoria riana DC. ex Ging. [nom. illeg.] =Rinorea riana 

Corynostylis hybanthus (L.) Mart. =Corynostylis arborea 

Gloeospermum ulei Melch. =Rinoreocarpus ulei 

Hybanthus circaeoides (Kunth) Baill. =Hybanthus oppositifolius 

Hybanthus ipecacuanha (L.) Baill. =Hybanthus calceolaria 

Ionidium calceolaria (L.) Vent. =Hybanthus calceolaria 

Ionidium circaeoides Kunth =Hybanthus oppositifolius 

Ionidium ipecacuanha (L.) Vent. =Hybanthus calceolaria 

Ionidium oppositifolium (L.) Roem. & Schult. =Hybanthus oppositifolius 

Ionidium phyllanthoides Planch. & Linden =Hybanthus phyllanthoides 

Leonia melinoniana (Baill.) Baill. ex Sagot =Leonia glycycarpa 

Noisettia longifolia Kunth =Noisettia orchidiflora 

Passoura guianensis Aubl. =Rinorea pubiflora var. pubiflora 

Paypayrola ventricosa Tul. =Paypayrola grandiflora 

Riana guianensis Aubl. =Rinorea riana 

Rinorea laxiflora (Benth.) Melch. =Rinorea brevipes 

Rinorea martini (Turcz.) S.F. Blake =Rinorea pubiflora var. pubiflora 

Rinorea passoura (DC. ex Ging.) Kuntze [nom. illeg.] =Rinorea pubiflora var. pubiflora 

Rinorea passoura Kuntze =Rinorea pubiflora var. pubiflora 

Rinorea passoura var. andersonii Sandwith ex Hekking =Rinorea pubiflora var. grandifolia 

Rinorea pulleana Melch. =Rinorea pubiflora var. grandifolia 

Rinorea riana (DC. ex Ging.) Kuntze [nom. illeg.] =Rinorea riana 

Rinorea surinamensis Melch. =Rinorea falcata 

Rinoreocarpus salmoneus Ducke =Rinoreocarpus ulei 

Spathularia longifolia A. St.-Hil. =Amphirrhox longifolia 

Viola arborea L. =Corynostylis arborea 

Viola calceolaria L. =Hybanthus calceolaria 


AM 


AM 

BO SU FG

Viola hybanthus Aubl. =Corynostylis arborea 

Viola hybanthus L. =Corynostylis arborea 

Viola ipecacuanha L. =Hybanthus calceolaria 

Viola itoubou Aubl. =Hybanthus calceolaria 

Viola oppositifolia L. =Hybanthus oppositifolius 


REFERENCES 

Fernández del Valle, Á. 2005. Violaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 445-464. 

Hekking, W.H.A. 1988. Violaceae part I - Rinorea and Rinoreocarpus. Flora Neotropica Monogr. 46: 1-208. 

VISCACEAE 

J. Kuijt 

Dendrophthora crispula (Rizzini) Kuijt [expected in GU] 

Dendrophthora decipiens Kuijt [Mt. Roraima] 

Dendrophthora densifrons (Ule) Kuijt [expected in the Guianas] 

Dendrophthora elliptica (Gardner) Krug & Urb. 

Dendrophthora fanshawei (Maguire) Kuijt 

Dendrophthora fendleriana (Eichler) Kuijt 

Dendrophthora intermedia (Rizzini) Kuijt 

Dendrophthora jauana Rizzini 

Dendrophthora karuaiana (Steyerm.) Kuijt 

Dendrophthora macbridei (Standl. ex J.F. Macbr.) Kuijt 

Dendrophthora microsoma Rizzini 

Dendrophthora nitidula (Rizzini) Kuijt 

Dendrophthora obliqua (C. Presl) Wiens 

Dendrophthora oligantha Kuijt 

Dendrophthora perfurcata (Rizzini) Kuijt 

Dendrophthora roraimae (Oliv.) Ule 

Dendrophthora tenuiflora (Steyerm. & Maguire) Kuijt 

Dendrophthora tepuiana (Steyerm.) Kuijt 

Phoradendron aphyllum Steyerm. 

Phoradendron bathyoryctum Eichler 

Phoradendron berryi Rizzini 

Phoradendron bilineatum Urb. 

Phoradendron chrysocladon A. Gray 

Phoradendron crassifolium (Pohl ex DC.) Eichler 

Phoradendron diminutivum E.A. Kellogg 

Phoradendron dipterum Eichler 

Phoradendron exiguum Trel. 

Phoradendron granvillei Kuijt 

Phoradendron haughtii Kuijt 

Phoradendron herbert-smithii Trel. 

Phoradendron hexastichum (DC.) Griseb. 

Phoradendron inaequidentatum Rusby 

Phoradendron kelloggii Kuijt 

Phoradendron krameri Kuijt 

Phoradendron longipetiolatum Urb. [expected in GU] 

Phoradendron mairaryense Ule 

Phoradendron membranifolium Kuijt 

Phoradendron microstachyum Kuijt 

Phoradendron morsicatum Rizzini 

Phoradendron mucronatum (DC.) Krug & Urb. 

Phoradendron nervosum Oliv. [expected in flora area] 

Phoradendron northropiae Urb. 

Phoradendron obtusissimum (Miq.) Eichler 

Phoradendron pellucidulum Eichler 

Phoradendron perrottetii (DC.) Eichler 

Phoradendron piperoides (Kunth) Trel. 

Phoradendron platycaulon Eichler 

Phoradendron poeppigii (Tiegh.) Kuijt 

Phoradendron pteroneuron Eichler 

Phoradendron pulleanum E.H.L. Krause 

AM BO 

AM BO GU 

BO GU 

AM BO GU 

GU SU? 

AM BO 

AM BO 

AM BO 

BO 

AM 

BO 

BO 

AM BO 

BO 

GU 

AM BO GU 

BO 

AM BO GU 

BO GU 

SU 

AM DA 

AM GU SU FG 

BO GU 


AM 

BO GU SU 

BO 

FG 

AM 

BO DA GU 

AM BO GU 


AM 

GU SU 

BO 

AM BO GU 

BO 

AM 

BO GU 

AM BO GU 

AM GU 

AM BO GU SU 

AM BO GU 



AM 

GU 

BO GU SU 

AM GU SU FG

550 

Phoradendron quadrangulare (Kunth) Griseb. 

Phoradendron racemosum (Aubl.) Krug & Urb. 

Phoradendron scariosum Rizzini 

Phoradendron schultesii Kuijt 

Phoradendron steyermarkii Rizzini 

Phoradendron strongyloclados Eichler 

Phoradendron triflorum E.A. Kellogg 

Phoradendron trinervium (Lam.) Griseb. 

Phoradendron tunaeforme (DC.) Eichler 

Phoradendron undulatum (Pohl ex DC.) Eichler 



Dendrophthora basiandra Kuijt =Dendrophthora macbridei 

Dendrophthora elliptica var. stenophylla Krug & Urb. =Dendrophthora elliptica 

Dendrophthora hylaeana Rizzini =Dendrophthora elliptica 

Dendrophthora inaequidentata (Rusby) Trel. =Phoradendron inaequidentatum 

Dendrophthora lacryma-jobi E.A. Kellogg =Dendrophthora crispula 

Dendrophthora parvispicata Rizzini =Dendrophthora microsoma 

Dendrophthora rubicunda Ule =Dendrophthora elliptica 

Loranthus piperoides Kunth =Phoradendron piperoides 

Loranthus quadrangulare Kunth =Phoradendron quadrangulare 

Phoradendron acinacifolium Mart. ex Eichler =Phoradendron obtusissimum 

Phoradendron acinacifolium var. surinamense Rizzini =Phoradendron obtusissimum 

Phoradendron apertum Trel. =Phoradendron trinervium 

Phoradendron appunii Trel. =Phoradendron trinervium 

Phoradendron ayliffei Philcox =Phoradendron undulatum 

Phoradendron baileyae Trel. =Phoradendron trinervium 

Phoradendron baileyae var. exile (Rizzini) Rizzini =Phoradendron trinervium 

Phoradendron bolivianum Trel. =Phoradendron hexastichum 

Phoradendron caerulesens Trel. =Phoradendron mucronatum 

Phoradendron caracasanum Urb. =Phoradendron mucronatum 

Phoradendron caracasanum var. parvifolium Rizzini =Phoradendron mucronatum 

Phoradendron carinatum Trel. =Phoradendron bilineatum 

Phoradendron cayennense Eichler =Phoradendron racemosum 

Phoradendron chaguaramasarum Trel. =Phoradendron mucronatum 

Phoradendron chrysocarpum var. stehlei Trel. & Stehlé =Phoradendron undulatum 

Phoradendron crenato-callosum Rizzini =Phoradendron herbert-smithii 

Phoradendron crispulum Rizzini =Dendrophthora crispula 

Phoradendron demerarae Trel. =Phoradendron dipterum 

Phoradendron dendrophthora Rizzini =Dendrophthora elliptica 

Phoradendron densifrons Ule =Dendrophthora densifrons 

Phoradendron dimidiatum (Miq.) Eichler =Phoradendron perrottetii 

Phoradendron duidanum Trel. =Dendrophthora fendleriana 

Phoradendron duidanum var. hymenaeifolium Rizzini =Dendrophthora obliqua 

Phoradendron dunstervillorum Rizzini =Phoradendron perrottetii 

Phoradendron ellipticum (Gardner) Eichler =Dendrophthora elliptica 

Phoradendron emarginatum Mart. ex Eichler =Phoradendron mucronatum 

Phoradendron emarginatum var. truncato-retusum Rizzini =Phoradendron mucronatum 

Phoradendron essequibense Trel. =Phoradendron strongyloclados 

Phoradendron excisum Rizzini =Phoradendron northropiae 

Phoradendron exile Rizzini =Phoradendron trinervium 

Phoradendron fanshawei Maguire =Dendrophthora fanshawei 

Phoradendron fendlerianum Eichler =Dendrophthora fendleriana 

Phoradendron flavens (Sw.) Griseb. =Phoradendron racemosum 

Phoradendron flavens var. longispicum Rizzini =Phoradendron chrysocladon 

Phoradendron glauco-lutescens Rizzini =Phoradendron piperoides 

Phoradendron granaticola Trel. =Phoradendron nervosum 

Phoradendron granaticola var. taeniicaule (Rizzini) Rizzini =Phoradendron nervosum 

Phoradendron guascanum Trel. =Dendrophthora obliqua 

Phoradendron harmsianum Ule =Phoradendron perrottetii 

Phoradendron herminieri Trel. =Phoradendron undulatum 

Phoradendron huricola Trel. =Phoradendron quadrangulare 

Phoradendron ichythystoma Rizzini =Phoradendron obtusissimum 

Phoradendron insigne Steyerm. =Phoradendron pellucidulum 

AM BO DA GU 


BO 

AM 

BO 


AM BO 


BO 

AM BO


Phoradendron intermedium Rizzini =Dendrophthora intermedia 

Phoradendron jenmanii Trel. =Phoradendron inaequidentatum 

Phoradendron johnstonii Trel. =Phoradendron strongyloclados 

Phoradendron knoopii Warb. ex Trel. =Phoradendron chrysocladon 

Phoradendron leptarthrum Rizzini =Phoradendron herbert-smithii 

Phoradendron linguiforme Rizzini =Phoradendron mairaryense 

Phoradendron longiarticulatum Rizzini =Phoradendron dipterum 

Phoradendron lyonii Trel. =Phoradendron mucronatum 

Phoradendron macbridei Standl. ex J.F. Macbr. =Dendrophthora macbridei 

Phoradendron martinicensis (DC.) Griseb. =Phoradendron crassifolium 

Phoradendron nitidulum Rizzini =Dendrophthora nitidula 

Phoradendron obliquum (C. Presl) Eichler =Dendrophthora obliqua 

Phoradendron ottonis Eichler =Phoradendron mucronatum 

Phoradendron ottonis var. oblongifolium Rizzini =Phoradendron mucronatum 

Phoradendron pachyphyllum Trel. =Phoradendron chrysocladon 

Phoradendron pennivenium (DC.) Eichler (Loranthaceae) =Oryctanthus florulentus 

Phoradendron perfurcatum Rizzini =Dendrophthora perfurcata 

Phoradendron prancei Rizzini =Phoradendron inaequidentatum 

Phoradendron pseudomucronatum Rizzini =Phoradendron pellucidulum 

Phoradendron ptarianum Steyerm. =Phoradendron longipetiolatum 

Phoradendron quadrangulare (Kunth) Krug & Urb. =Phoradendron quadrangulare 

Phoradendron roraimae Oliv. =Dendrophthora roraimae 

Phoradendron rubrum var. longipedunculatum Rizzini =Phoradendron quadrangulare 

Phoradendron rugulosum Urb. =Phoradendron pteroneuron 

Phoradendron sanctae-martae Trel. =Phoradendron trinervium 

Phoradendron scariosum Rizzini =Phoradendron perrottetii 

Phoradendron schottii A. Gray =Phoradendron piperoides 

Phoradendron semivenosum Rizzini =Phoradendron pellucidulum 

Phoradendron semivenosum var. agostinii Rizzini =Phoradendron pellucidulum 

Phoradendron semivenosum var. longipes Rizzini =Phoradendron pellucidulum 

Phoradendron spectabile Rizzini =Dendrophthora fendleriana 

Phoradendron spectabile var. altimontanum Rizzini =Dendrophthora fendleriana 

Phoradendron sulfuratum Rizzini =Phoradendron mairaryense 

Phoradendron surinamense Pulle =Phoradendron strongyloclados 

Phoradendron taeniicaule Rizzini =Phoradendron nervosum 

Phoradendron ? tatei Trel. =Dendrophthora fendleriana 

Phoradendron tenuiflorum Steyerm. & Maguire =Dendrophthora tenuiflora 

Phoradendron tepuianum Steyerm. =Dendrophthora tepuiana 

Phoradendron theloneuron Rizzini =Phoradendron trinervium 

Phoradendron tovarense Urb. =Phoradendron dipterum 

Phoradendron treleasei Rizzini =Phoradendron trinervium 

Phoradendron tubulosum Urb. =Phoradendron nervosum 

Phoradendron venezuelense Trel. =Phoradendron quadrangulare 

Phoradendron venezuelense var. filispicum Rizzini =Phoradendron quadrangulare 

Phoradendron verticillatum Fawc. & Rendle [nom. superfl.] =Phoradendron trinervium 

Phoradendron verticillatum (L.) Druce (Vitaceae) =Cissus verticillata 

Phoradendron williamsii Rizzini =Phoradendron pellucidulum 

Phoradendron zuloagae Trel. =Phoradendron trinervium 

Viscum crassifolium Pohl ex DC. =Phoradendron crassifolium 

Viscum dichotomum Bert. ex Spreng. =Phoradendron piperoides 

Viscum dimidiatum Miq. =Phoradendron perrottetii 

Viscum ellipticum Gardner =Dendrophthora elliptica 

Viscum flavens Sw. [nom. superfl.] =Phoradendron racemosum 

Viscum fockeanum Miq. =Phoradendron piperoides 

Viscum glandulosum Miq. =Phoradendron racemosum 

Viscum hexastichum DC. =Phoradendron hexastichum 

Viscum latifolium Lam. =Phoradendron racemosum 

Viscum martinicensis DC. =Phoradendron crassifolium 

Viscum mucronatum DC. =Phoradendron mucronatum 

Viscum obliquum C. Presl =Dendrophthora obliqua 

Viscum obtusissimum Miq. =Phoradendron obtusissimum 

Viscum perrottetii DC. =Phoradendron perrottetii 

Viscum piperoides (Kunth) DC. =Phoradendron piperoides 

Viscum quadrangulare (Kunth) DC. =Phoradendron quadrangulare

552 

Viscum racemosum Aubl. =Phoradendron racemosum 

Viscum trinervium Lam. =Phoradendron trinervium 

Viscum tunaeforme DC. =Phoradendron tunaeforme 

Viscum undulatum Pohl ex DC. =Phoradendron undulatum 

Viscum verticillatum L. (Vitaceae) =Cissus verticillata 


REFERENCES 

(see also Eremolepidaceae, Loranthaceae) 

Krause, K. 1932. Loranthaceae. In A. Pulle, Flora of Suriname 1(1): 4-24. 

Kuijt, J. 1961. A revision of Dendrophthora. Wentia 6: 1-145. 

Kuijt, J. 2005. Viscaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 9: 464- 

492. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. Loranthaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of 

Suriname 1(2): 295-300. 

Rizzini, C.T. 1978. Loranthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland, part 10. Mem. New York 

Bot. Gard. 29: 23-36. 

Rizzini, C.T. 1982. Loranthaceae. In Z. Luces de Febres, Flora de Venezuela 4(2): 7-316. 

Cissus alata Jacq. 

Cissus descoingsii J.A. Lombardi 

Cissus duarteana Cambess. 

Cissus erosa Rich. 

Cissus gongylodes (Baker) Planch. 

Cissus guyanensis Desc. 

Cissus haematantha Miq. 

Cissus palmata Poir. 

Cissus spinosa Cambess. 

Cissus subrhomboidea (Baker) Planch. 

Cissus surinamensis Desc. 

Cissus trianae Planch. 

Cissus ulmifolia (Baker) Planch. 

Cissus venezuelensis Steyerm. 

Cissus verticillata (L.) Nicolson & C.E. Jarvis 

Cissus verticillata subsp. verticillata 

Vitis tiliifolia Willd. ex Roem. & Schult. 

VITACEAE 

F.D. Chitty, J.A. Steyermark† 


Cissus bonariensis Hook. & Arn. =Cissus palmata 

Cissus cervii Dunaiski =Cissus gongylodes 

Cissus cordifolia L. =Cissus verticillata subsp. verticillata 

Cissus elongata Miq. =Cissus erosa 

Cissus erosa var. salutaris (Kunth) Planch. =Cissus erosa 

Cissus gibertii (Baker) Planch. =Cissus palmata 

Cissus guaranitica Chodat & Hassl. =Cissus erosa 

Cissus hassleriana Chodat =Cissus spinosa 

Cissus lucida Poir. =Cissus erosa 

Cissus martiniana Woodson & Seibert =Cissus trianae 

Cissus microcarpa Vahl =Cissus erosa 

Cissus ovata Lam. =Cissus verticillata subsp. verticillata 

Cissus palmata var. balansaeana Planch. =Cissus palmata 

Cissus paraguayensis Planch. =Cissus palmata 

Cissus parkeri (Baker) Planch. =Cissus spinosa 

Cissus pubescens Kunth =Cissus alata 

Cissus quadrialata Kunth =Cissus erosa 

Cissus rhombifolia Vahl =Cissus alata 

Cissus salutaris Kunth =Cissus erosa 

Cissus sexangularis Ernst =Cissus erosa 

Cissus sicyoides L. =Cissus verticillata subsp. verticillata 

Cissus smilacina Kunth =Cissus verticillata subsp. verticillata 

Cissus tamoides Cambess. =Cissus verticillata subsp. verticillata 

Cissus trianae var. martiniana (Woodson & Seibert) Steyerm. =Cissus trianae 

AM BO DA 

BO GU 

AM 

AM 

BO DA GU SU 

FG 

FG 

FG 


DA GU SU FG 

DA GU SU FG 

GU SU 

AM 

SU 

SU 

FG 

BO GU 



FG?


Cissus tricuspis (Baker) Planch. =Cissus gongylodes 

Cissus trifoliata (L.) L. ? =Cissus erosa 

Cissus verticillata subsp. laciniata (Baker) J.A. Lombardi =Cissus verticillata subsp. verticillata 

Vitis alata (Jacq.) Kuntze =Cissus alata 

Vitis bakeri Herter =Cissus palmata 

Vitis erosa Baker =Cissus erosa 

Vitis gibertii Baker =Cissus palmata 

Vitis gongylodes Baker =Cissus gongylodes 

Vitis miqueliana Baker =Cissus erosa 

Vitis palmata (Poir.) Baker =Cissus palmata 

Vitis parkeri Baker =Cissus spinosa 

Vitis rhombifolia (Vahl) Baker =Cissus alata 

Vitis sicyoides (L.) Morales, [nom. illeg.] =Cissus verticillata subsp. verticillata 

Vitis sicyoides (L.) Miq. =Cissus verticillata subsp. verticillata 

Vitis sicyoides (L.) Baker , [nom. illeg.] =Cissus verticillata subsp. verticillata 

Vitis sicyoides f. laciniata Baker =Cissus verticillata subsp. verticillata 

Vitis sicyoides var. smilacina (Kunth) Baker =Cissus verticillata subsp. verticillata 

Vitis spinosa (Cambess.) Baker =Cissus spinosa 

Vitis tricuspis Baker =Cissus gongylodes 

REFERENCES 

Descoings, B. 1991. Contribution à l’étude des Vitacées d’Amérique tropicale: deux Cissus nouveaux des Guyanes. Bull. Soc. 

Bot. France, Lett. Bot. 138(3): 249-256. 

Delascio Chitty, F. & J.A. Steyermark. 2005. Vitaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the 


Görts-van Rijn, A.R.A. 1979. Vitaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 335-343. 

Lombardi, J.A. 1995. Typification of names of South American Cissus (Vitaceae). Taxon 44(2): 193-206. 

Suessenguth, K. 1953. Vitaceae. In A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien 20d: 174-333. 

Erisma blancoa Marc.-Berti 

Erisma calcaratum (Link) Warm. 

Erisma floribundum Rudge 

Erisma floribundum var. floribundum 

Erisma japura Spruce ex Warm. 

Erisma laurifolium Spruce ex Warm. 

Erisma micranthum Spruce ex Warm. 

Erisma nitidum DC. 

Erisma splendens Stafleu 

Erisma uncinatum Warm. 

Qualea acuminata Spruce ex Warm. 

Qualea amapaensis Balslev & S.A. Mori 

Qualea coerulea Aubl. 

Qualea cyanea Ducke 

Qualea decorticans Ducke [expected in AM] 

Qualea dinizii Ducke 

Qualea grandiflora Mart. 

Qualea homosepala Ducke 

Qualea mori-boomii Marc.-Berti 

Qualea paraensis Ducke 

Qualea polychroma Stafleu 

Qualea psidiifolia Spruce ex Warm. 

Qualea pulcherrima Spruce ex Warm. 

Qualea rosea Aubl. 

Qualea rupicola Ducke 

Qualea schomburgkiana Warm. 

Qualea sprucei Warm. 

Qualea suprema Ducke 

Qualea themistoclesii Ducke 

Qualea tricolor Benoist 

Qualea tuberculata Stafleu 

Qualea wurdackii Marc.-Berti 

VOCHYSIACEAE 

L. Marcano-Berti 

AM 

AM GU 

AM FG 

AM FG 

AM 

AM 

AM 

GU FG 

AM 


AM BO GU? FG 

FG? 

SU FG 

AM 


SU 

BO 

FG 

AM BO GU 

BO GU 

AM FG 

AM 

SU FG 

AM 

BO GU 

AM 

AM 

AM 

FG 

AM 

AM

554 

Ruizterania albiflora (Warm.) Marc.-Berti 

Ruizterania cassiquiarensis (Spruce ex Warm.) Marc.-Berti 

Ruizterania esmeraldae (Standl.) Marc.-Berti 

Ruizterania ferruginea (Steyerm.) Marc.-Berti 

Ruizterania nitida (Stafleu) Marc.-Berti 

Ruizterania obtusata (Briq.) Marc.-Berti 

Ruizterania obtusata subsp. multivenosa Marc.-Berti 

Ruizterania obtusata subsp. obtusata 

Ruizterania retusa (Spruce ex Warm.) Marc.-Berti 

Ruizterania rigida (Stafleu) Marc.-Berti 

Ruizterania trichanthera (Spruce ex Warm.) Marc.-Berti 

Salvertia convallariodora A. St.-Hil. 

Vochysia angustifolia Ducke 

Vochysia apopetala Ule 

Vochysia bautistae Marc.-Berti 

Vochysia calophylla Spruce ex Warm. 

Vochysia cassiquiarensis Stafleu 

Vochysia catingae Ducke 

Vochysia cayennensis Warm. 

Vochysia complicata Ducke 

Vochysia costata Warm. 

Vochysia crassifolia Warm. 

Vochysia densiflora Spruce ex Warm. 

Vochysia elegans Stafleu 

Vochysia expansa Ducke 

Vochysia ferruginea Mart. 

Vochysia glaberrima Warm. 

Vochysia grandis Mart. 

Vochysia grandis var. uaupensis Warm. 

Vochysia guianensis Aubl. 

Vochysia jonkeri Marc.-Berti 

Vochysia julianensis Marc.-Berti 

Vochysia aff. laxiflora Stafleu 

Vochysia liscanoi Marc.-Berti 

Vochysia maguirei Marc.-Berti 

Vochysia neyratii Normand 

Vochysia obscura Warm. 

Vochysia ortegae Marc.-Berti & Bautista 

Vochysia pauciflora Steyerm. 

Vochysia pinkusii A.C. Sm. [expected in BO] 

Vochysia punctata Spruce ex Warm. 

Vochysia rubiginosa Stafleu 

Vochysia sabatieri Marc.-Berti 

Vochysia saccata Stafleu 

Vochysia schomburgkii Warm. 

Vochysia spathiphylla Stafleu 

Vochysia speciosa Warm. 

Vochysia splendens Spruce ex Warm. 

Vochysia steyermarkiana Marc.-Berti 

Vochysia surinamensis Stafleu 

Vochysia surinamensis var. inflata Stafleu 

Vochysia surinamensis var. surinamensis 

Vochysia tetraphylla (G. Mey.) DC. 

Vochysia tilletii Marc.-Berti 

Vochysia tomentosa (G. Mey.) DC. 

Vochysia tucanorum Mart. 

Vochysia tucanorum var. tucanorum 

Vochysia venezuelana Stafleu 

Vochysia vismiifolia Spruce ex Warm. 



Cucullaria excelsa M. Vahl =Vochysia tomentosa 

Cucullaria ferruginea (Mart.) Spreng. =Vochysia ferruginea 

Cucullaria grandis (Mart.) Spreng. =Vochysia grandis 

GU SU FG 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

AM 

BO GU 

AM 

BO GU 

SU 

AM 

BO 

AM 

AM 

AM 

AM 

FG 

AM 

AM BO GU 

AM BO GU 

GU SU FG 

AM 

AM 

AM BO GU 

AM 

AM 

AM 

BO GU SU 

GU SU FG 

BO 

AM 

AM SU 

BO 

GU 

FG 

AM 

AM 

BO 

AM 

BO 

FG 

AM 

GU 

AM 

FG 

AM 

AM 

AM BO 

BO 

DA GU SU FG 


AM 


AM BO GU SU 

SU 

SU 

FG 

AM BO 

AM BO


Cucullaria tetraphylla G. Mey. =Vochysia tetraphylla 

Cucullaria tomentosa G. Mey. =Vochysia tomentosa 

Debraea floribundum (Rudge) Roem. & Schult. =Erisma floribundum 

Debraea violacea (Mart.) Steud. =Erisma calcaratum 

Ditmaria floribunda (Rudge) Spreng. =Erisma floribundum 

Ditmaria violacea (Mart.) Spreng. =Erisma calcaratum 

Erisma pallidiflorum Ducke =Erisma floribundum 

Erisma parvifolium Gleason =Erisma floribundum 

Erisma parvifolium var. pallidiflorum (Ducke) Ducke =Erisma floribundum 

Erisma pulverulentum Poepp. ex Warm. [nomen nudum] =Erisma uncinatum 

Erisma violaceum Mart. =Erisma calcaratum 

Qualea albiflora Warm. =Ruizterania albiflora 

Qualea apodocarpa Steyerm. =Ruizterania ferruginea 

Qualea calcarata Link. =Erisma calcaratum 

Qualea cassiquiarensis Spruce ex Warm. =Ruizterania cassiquiarensis 

Qualea esmeraldae Standl. =Ruizterania esmeraldae 

Qualea ferruginea Steyerm. =Ruizterania ferruginea 

Qualea nitida Stafleu =Ruizterania nitida 

Qualea obtusata Briq. =Ruizterania obtusata 

Qualea retusa Spruce ex Warm. =Ruizterania retusa 

Qualea retusa var. coriacea Ducke =Ruizterania obtusata 

Qualea rigida Stafleu =Ruizterania rigida 

Qualea rubiginosa Stafleu =Ruizterania ferruginea 

Qualea rubiginosa var. angustior Steyerm. =Ruizterania ferruginea 

Qualea speciosa Huber =Qualea acuminata 

Qualea trichanthera Spruce ex Warm. =Ruizterania trichanthera 

Qualea verruculosa Stafleu =Ruizterania esmeraldae 

Ruizterania apodocarpa (Steyerm.) Marc.-Berti =Ruizterania ferruginea 

Ruizterania rubiginosa (Stafleu) Marc.-Berti =Ruizterania ferruginea 

Ruizterania rubiginosa var. angustior (Steyerm.) Marc.-Berti =Ruizterania ferruginea 

Ruizterania verruculosa (Stafleu) Marc.-Berti =Ruizterania esmeraldae 

Vochya ferruginea (Mart.) Standl. =Vochysia ferruginea 

Vochysia arcuata Garcke =Vochysia tetraphylla 

Vochysia curvata Klotzsch ex R.M. Schomb. [nomen nudum] =Vochysia crassifolia 

Vochysia grandis var. genuina Briq. =Vochysia grandis 

Vochysia inundata var. venosa Ducke =Vochysia saccata 

Vochysia javitensis Stafleu =Vochysia angustifolia 

Vochysia lucida Klotzsch ex R.M. Schomb. [nomen nudum] =Vochysia glaberrima 

Vochysia racemosa Poir. [name of uncertain application, not Vochysia] 

Vochysia tetraphylla (G. Mey.) H. Stone & W. Freeman =Vochysia tetraphylla 

Vochysia vismiifolia var. robusta Ducke =Vochysia splendens 

REFERENCES 

Marcano-Berti, L. 1969. Un nuevo genero de las Vochysiaceae. Pittieria 2: 3-27. 

Marcano-Berti, L. 1989. Vochysiaceae: novedades y correciones. Pittieria 18: 5-14. 

Marcano-Berti, L. 1998. Vochysiaceae. In A.R.A. Görts-van Rijn, Flora of the Guianas (series A) 21: 1-44. 

Marcano-Berti, L. 2005. Vochysiaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan 

Guayana 9: 500-524. 

Stafleu, F.A. 1948-1954. A monograph of the Vochysiaceae. I. Salvertia & Vochysia. Rec. Trav. Bot. Neerl. 41: 397-540 (1948). 

III. Qualea. Acta Bot. Neerl. 2: 144-217 (1953). IV. Erisma. Acta Bot. Neerl. 3: 459-480 (1954). 

Stafleu, F.A. 1951. Vochysiaceae. In A. Pulle, Flora of Suriname 3(2): 178-199. 

Steyermark, J.A. 1987. Vochysiaceae: a reevaluation of the genus Euphronia. In J.A. Steyermark, Flora of the Venezuelan 

Guayana - II. Ann. Missouri Bot. Gard. 74(1): 85-116. 

WINTERACEAE 

J.S. Miller 

Drimys roraimensis (A.C. Sm.) Ehrend. & Gottsb.[Mt. Roraima] 


Drimys brasiliensis var. ptaritepuiensis Steyerm. =Drimys roraimensis 

Drimys brasiliensis var. roraimensis A.C. Sm. = Drimys roraimensis 

REFERENCES 

Smith, A.C. 1943. The American species of Drimys. J. Arn. Arb. 24: 1-33. 

AM BO GU

556 


Eherendorfer, F, I. Silberbauer-Gottsberger & G. Gottsberger. 1979. Variation on the population, racial, and species level in the 

primitive relic angiosperm genus Drimys (Winteraceae) in South America. Plant Systematics and Evolution 132: 53-83. 

Miller, J.S. 2005. Winteraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of the Venezuelan Guayana 

9: 524-525. 

Guaiacum officinale L. 

Kallstroemia maxima (L.) Hook. & Arn. 

Kallstroemia pubescens (G. Don) Dandy 

Tribulus cistoides L. 

Tribulus zeyheri Sond. 

Tribulus zeyheri subsp. macranthus (Hassk.) Hadidi 

ZYGOPHYLLACEAE 

G.A. Aymard C., N.L. Cuello A., B.E. Leuenberger 


Kallstroemia caribaea Rydb. =Kallstroemia pubescens 

Tribulus macranthus Hassk. =Tribulus zeyheri subsp. macranthus 

Tribulus maximus L. =Kallstroemia maxima 

Tribulus pubescens G. Don =Kallstroemia pubescens 

BO GU SU 

BO FG 

BO 

GU SU 

BO 

BO 

REFERENCES 

Aymard C., G.A. & N.L. Cuello A. 2005. Zygophyllaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holst, Flora of 


Howard, R.A. 1988. Zygophyllaceae. In R.A. Howard, Flora of the Lesser Antilles 4(1): 551-555. 

Lasser, T. 1971. Zygophyllaceae. In T. Lasser, Flora de Venezuela 3(1): 75-89.Stafleu, F.A. 1951. Zygophyllaceae. In A. Pulle, 

Flora of Suriname 3(2): 200-203. 

Justicia wilhelmensis Wasshausen (Acanthaceae)


Passiflora kawensis Feuillet (Passifloraceae)

558 


Stifftia uniflora Ducke amd Stifftia cayennensis H.Rob. & Kahn (Compositae)


Talisia simaboides Kramer (Sapindaceae)

560 


Pinzona coriacea Mart. & Zucc. (Dilleniaceae)


Lacunaria jenmanii (Oliv.) Ducke (Quiinaceae)

562 

Page 4 

Page 16 

Page 556 

Page 557 

Page 558 

Page 559 

Page 560 

Page 561 

Page 583 


Literature citation for illustrations 

Henriettea multiflora Naudin, In: Wurdack, J. J., Morley, T., and S. Renner 1993. 

Melastomataceae. Flora of the Guianas: Series A: Phanerogams, Fascicle 13: 

425. Koenigstein, Germany. Koeltz. Illustrator: Alice Tangerini 

Guapira eggersiana (Heimerl) Lundell, In: DeFilippe, R. (in review) Nyctaginaceae. 

Flora of the Guianas: Series A: Phanerogams, Fascicle xx. Richmond, United 

Kingdom. Royal Botanical Garden, Kew. Illustrator: Alice Tangerini. 

Justicia wilhelmensis Wasshausen, In: Wasshausen, D. C. 2006. Acanthaceae. Flora 

of the Guianas: Series A: Phanerogams, Fascicle 23: 1-141. Richmond, United 

Kingdom. Royal Botanical Garden, Kew. Illustrator: Alice Tangerini. 

Passiflora kawensis Feuillet, In: Feuillet, C. 1994. Two new species of Passiflora 

(Passifloraceae) from French Guiana. Novon 4(3): 236-238. Illustrator: Cathy 

Pasquale. 

Stifftia uniflora Ducke and Stifftia cayennensis H. Robinson & B. Kahn, In: Robinson, 

H. and B. Kahn. 1985. A new species of Stifftia from Cayenne (Asteraceae: 

Mutisieae) Phytologia 58(4): 248-251. Illustrator: Alice Tangerini. 

Talisia simaboides Kramer, In: Acevedo-Rodríquez, P. 2003. Melicocceae 

(Sapindaceae): Melicoccus and Talisia. Flora Neotropica 87: 1-178. Illustrator: 

Bobbi Angell. 

Pinzona coriacea Mart. & Zucc., In: Mori, S. A., G. Cremers, C. A. Gracie, J. de 

Granville, S. V. Heald, M. Hoff and J. D. Mitchell. 2002. Guide to the Vascular 

plants of Central French Guiana, Pt. 2. Dicotyledons. Memoirs of the New York 

Botanical Garden 76: 251. Illustrator: Bobbi Angell. 

Lacunaria jenmanii (Oliv.) Ducke, In: Mori, S. A., G. Cremers, C. A. Gracie, J. de 

Granville, S. V. Heald, M. Hoff and J. D. Mitchell. 2002. Guide to the Vascular 

plants of Central French Guiana, Pt. 2. Dicotyledons. Memoirs of the New York 

Botanical Garden 76: 596. Illustrator: Bobbi Angell. 

Rhynchospora cordatachenia M.T. Strong, In: Strong, M. T. 2005. Two new species 

of Rhynchospora section Tenues (Cyperaceae) from the Guianas, South America. 

Novon 15(3): 479-483. Illustrator: Alice Tangerini.

Dr. Pedro Acevedo R. (US) 

Dr. Mac Alford (USMS) 

Dr. Lucile Allorge (P) 

Dr. Frank Almeda (CAS) 

Dr. Christiane Anderson (MICH) 

Dr. William R. Anderson (MICH) 

Dr. A. Aptroot (U) 

Dr. Scott Armbruster (ALA) 

Dr. Daniel F. Austin (Arizona-Sonora 

Desert Museum) 

Dr. Franklin Ayala (AMAZ) 

Dr. Gerardo A. Aymard C. (PORT) 

Dr. Michael J. Balick (NY) 

Dr. Harvey E. Ballard (OH) 

Dr. Julie Barcelona (PNH) 

Dr. Rupert C. Barneby † (NY) 

Dr. Lisa C. Barnett (US) 

Dr. S. Barrier (P) 

Dr. Kerry Barringer (BKL) 

Dr. Daniel Barthelemy (MPU) 

Dr. Cornelis C. Berg (BG) 

Dr. Paul Berry (MICH) 

Dr. M.H. Bischler (G) 

Mr. John K. Boggan (US) 

Dr. Brian Boom (NY) 

Mr. Jason Bradford (MO) 

Dr. John Brandbyge (AAU) 

MS Arian Jacobs-Brouwer (U) 

Dr. William R. Buck (NY) 

Dr. Ricardo Callejas (HUA) 

Dr. Koen Camelbeke (Stichting 

Arboretum Wespelaar, RUG) 

Dr. Kenneth K. Cameron (NY) 

Lic. Germán Carnevali (MO) 

Dr. Paul M. Catling (DAOM) 

Dr. Andre Chanderbali (FLAS) 

Dr. Mark Chase (MICH) 

Dr. Eric A. Christenson 

Dr. John L. Clarke (US) 

Dr. H. David Clarke (UNCA) 

Dr. M.J.E. Coode (K) 

Dr. Alain Couté (PC) 

Dr. Richard Cowan† (PERTH) 

Dr. George Cremers (P) 

Dr. Thomas B. Croat (MO) 

Dr. José Cuatrecasas† (US) 

Dr. C. Cusset (P) 

Dr. Douglas C. Daly (NY) 

Dr. Chris Davidson (Boise Botanical 

Garden) 

Dr. Garrit Davidse (MO) 

Dr. Robert DeFilipps† (US) 


APPENDIX 

Individuals who provided plant identifications. 

(† = deceased) 

Dr. Alfonso Delgado-Salinas (MEXU) 

Dr. Piero Delprete (NY) 

Dr. Paula DePriest (US) 

Dr. B.M. Descoings (MPU) 

Mr. Ron Determann (Atlanta Botanical 

Garden) 

Dr. Pierre Detienne (CTFT) 

Lic. Wilmer Díaz (GUYN) 

Dr. Laurence J. Dorr (US) 

Dr. Stefan Dressler (Leiden) 

Dr. Renski Ek (U) 

H.-J. Esser (U) 

Dr. Robert B. Faden (US) 

Dr. Paul R. Fantz (NCSC) 

Dr. Marie L. Farr (BPI) 

Dr. Christian Feuillet (US) 

Dr J.J. Floret (P) 

Dr. James P. Folsom (HNT) 

Dr. Enrique Forero (MO) 

Dr. Alina Freire-Fierro (PH) 

Dr. V.A. Funk (US) 

Dr. Alwyn H. Gentry † (MO) 

Dr. Lynn J. Gillespie (CAN) 

Dr. Tom Givnish (WIS) 

Dr. Paul Goetghebeur (GENT) 

Dr. Aaron Goldberg (US) 

Dr. Peter Goldblatt (MO) 

Dr. A.R.A. Görts-van Rijn (U) 

Dr. Eric J. Gouda (U) 

Dr. S.R. Gradstein (GOET) 

Dr. Jason R. Grant (NEU) 

Dr. Jean-Jacques de Granville (CAY) 

Dr. James Grimes (NY at the time of 

the work) 

Dr. Susan Grose (WTU) 

Dr. Micheline Guerlesquin (ANGUC) 

Dr. J. Hagemann (B) 

Dr. Eric Hagsater (AMO) 

Dr. William J. Hahn (Georgetown 

Univ.) 

Dr. Madjit Hakki (B) 

Dr. Bertel Hansen (C) 

Dr. Ray M. Harley (K) 

Dr. Robert R. Haynes (UNA) 

Dr. W.H.A. Hekking† (U) 

Dr. H.v. Helvoort (U) 

Dr. Andrew Henderson (NY) 

Dr. Terry Henkel (HSC) 

Dr. E. Hennipman (U) 

Dr. Patrick Herendeen (Geo. 

Washington Univ.) 

Dr. Paul Hiepko (B) 

Dr. Steven Hill (CLEMS) 

Dr. Michel Hoff (STR) 

Mr. Bruce Hoffman (HAW) 

Dr. Tom Hollowell (US) 

Dr. Walter C. Holmes (BAYLU) 

Dr. Noel H. Holmgren (NY) 

Dr. Lauritz B. Holm-Nielsen (AAU) 

Mr. Bruce Holst (SEL) 

Dr. Charles N. Horn (NBYC) 

Dr. Otto Huber (Venezuela) 

Dr. Marion J. Jansen-Jacobs (U) 

Dr. C. Jeffrey (L) 

Dr. J. Jéremie (P) 

Dr. A.C. Jermy (BM) 

Dr. S. Jovet-Ast (PC) 

Dr. Emmet J. Judziewicz (UWSP) 

Dr. F. Kahn (CAY) 

Dr. Jacquelyn Kallunki (NY) 

Dr. Carol L. Kelloff (US) 

Dr. Helen Kennedy (UBC) 

Dr. Robert W. Kiger (CM) 

Dr. P. Kirk (IMI) 

Dr. Joseph H. Kirkbride (BARC) 

Dr. B. B. Klitgaard (BM) 

Dr. Sandy Knapp (BM) 

Dr. Robert Kral (BRIT) 

Dr. Karl U. Kramer (Z) 

Dr. W. John Kress (US) 

Dr. R. Kruijt (U) 

Dr. Job Kuijt (UVIC) 

Dr. Klaus Kubitzki (HBG) 

Dr. Lars Peter Kvist (AAU) 

Prof. Dr. H.W. Lack (B) 

Dr. Tom Lammers (OSH) 

Dr. Leslie R. Landrum (ASU) 

Dr. A. de Lange (AMD) 

Dr. Anthony J.M. Leeuwenberg 

(WAG) 

Dr. David B. Lellinger (US) 

Dr. Blanca Leon (TEX) 

Dr. Beat Ernst Leuenberger (B) 

Dr. Gwilym Lewis (K) 

Mr. Ron Liesner (MO) 

Dr. H. C. de Lima (RB) 

Dr. Jan C. Lindeman (U) 

Dr. Eduardo Lleras-Perez (CEN) 

Dr. Pierre-André Loizeau (G) 

Dr. Júlio A. Lombardi (HRCB) 

Ms. Ximena Londoño (Cali, 

Colombia) 

Dr. Alicia Lourteig †(P) 

Dr. Carlyle Luer (MO)

564 

Dr. Eimear Nic Lughada (K) 

Dr. James L. Luteyn (NY) 

Mr. Harry E. Luther (SEL) 

Dr. Paul J.M. Maas (U) 

Dr. H. Maas-van de Kamer (U) 

Dr. Bob Magill (MO) 

Dr. Luis Marcano-Berti via (U) 

Dr. Richard H. Maxwell (JEF) 

Dr. Lucille M. McCook (MISS) 

Dr. Tim McDowell (ETSU) 

Mr. Greg McKee (US) 

Dr. J.T. de Medeiros Costa (UFP) 

Mr. Max E. Medley (DHL) 

Dr. Alan W. Meerow (FTG) 

Dr. Willem Meijer† (KY) 

Dr. L. Melendez-Howell (PC) 

Dr. A.M.W. Mennega †(U) 

Dr. Fabian Michelangeli (NY) 

Dr. James Miller (MO) 

Dr. David Minter (IMI) 

Dr. John D. Mitchell (NY) 

Dr. Reinaldo Monteiro (HRCB) 

Dr. Robbin Moran (NY) 

Dr. Luis E. Mora-Osejo (COL) 

Dr. Scott Mori (NY) 

Dr. Gilberto Morillo (HUA) 

Dr. Thomas Morley (MIN) 

Dr. Joao Murca Pires (MG) 

Dr. Jose C. Murillo (COL) 

Dr. Michael Nee (NY) 

Dr. David A. Neill (MO) 

Dr. Dan H. Nicolson (US) 

Dr. James N. Norris (US) 

Dr. B. Oellgaard (AAU) 

Mr. Darin Penneys (FLAS) 

Dr. R. Toby Pennington (E) 

Dr. T.D. Pennington (K) 

Dr. Paul Peterson (US) 

Dr. John J. Pipoly (Florida) 

Dr. Timothy Plowman † (MO) 


Dr. Roger M. Polhill (K) 

Dr. Odile Poncy (P) 

Dr. Hans-Helmut Poppendieck (HBG) 

Dr. Ghillean T. Prance (K) 

Mr. John F. Pruski (MO) 

Dr. W. Punt (U) 

Dr. J.M. Rankin (BM) 

Dr. Thomas Raus (B) 

Dr. Peter H. Raven (MO) 

Dr. Robert W. Read † (US) 

Dr. Karen M. Redden (Geo. 

Washington Univ.) 

Dr. Susanne Renner (UMSL) 

Dr. Harold Robinson (US) 

Dr. Norman K.B. Robson (BM) 

Dr. André Robyns (BR) 

Dr. William A. Rodrigues (UPCB) 

Dr. Jack D. Rogers (WSP) 

Sr. Gustavo A. Romero (AMES) 

Dr. A.C. de Roon (U) 

Dr. Velva E. Rudd† (US) 

Dr. Daniel Sabatier (MPA) 

Dr. Gary G. Samuels (BPI) 

Dr. Claude Sastre (P) 

Dr. Jakob J. Schneller (Z) 

Dr. Brian Schrire (K) 

Mr. Matthew Sewell (NY) 

Dr. M.F. da Silva (INPA) 

Dr David A. Simpson (K) 

Dr. Beryl Simpson (TEX) 

Dr. Laurence E. Skog (US) 

Dr. Flemming Skov (AAU) 

Dr. Alan R. Smith (UC) 

Dr. Lyman B Smith † (US) 

Dr. Steven F. Smith (US) 

Dr. James Solomon (MO) 

Dr. Mario Sousa Sanchez (MEXU) 

Dr. Chelsea Specht (U.CA) 

Dr. Rodolphe E. Spichiger (G) 

Dr. Clive A. Stace (LTR) 

Dr. Bertil Ståhl (SUNIV) 

Dr. Fred W. Stauffer (G) 

Dr. Basil Stergios (PORT) 

Dr. Dennis W. Stevenson (NY) 

Dr. Donald E. Stone (DUKE) 

Dr. Mark T. Strong (US) 

Dr. D.L. Szlachetko (UGDA) 

Dr. Charlotte M. Taylor (MO) 

Dr. Peter Taylor (K) 

Mr. Oswaldo Tellez Valdez (MEXU) 

Dr. Y. Therezien (P) 

Dr. W. Wayt Thomas (NY) 

Dr. Sue Thompson (CM) 

Dr. Pierre Tixier (PC) 

Mr. Ben Torke (MO) 

Ms. Erin A. Tripp (DUKE) 

Ms. Amy Trojnar (Southern Illinois 

University) 

Dr. Gordon C. Tucker (EIU) 

Dr. Yvonne Veyret (P) 

Dr. Warren L. Wagner (US) 

Dr. Warren H. Wagner Jr.† (MICH) 

Dr. Bruno Wallnofer (W) 

Dr. Dieter Wasshausen (US) 

Dr. Grady L. Webster † (DAV) 

Dr. Anna Weitzman (US) 

Dr. Tom Wendt (TEX) 

Dr. H. van der Werff (MO) 

Dr. Marga C.M. Werkhoven (BBS) 

Dr. John H. Wiersema (BARC) 

Dr. Paulo G. Windisch (HASU) 

Dr. Donald R. Windler (BALT) 

Dr. Robert Wingfield (CORO) 

Dr. John J. Wurdack† (US) 

Dr. Kenneth Wurdack (US) 

Dr. Thomas Zanoni (NY) 

Dr. D. Zappi (K) 

Dr. Elsa M. Zardini (MO) 

Dr. James Zarucchi (MO) 

Dr. Brigitte Zimmer (B)

Abarema 365, 370-372, 374-375 

Abelmoschus 393-394 

Aberemoa 193 

Abildgaardia 88, 95-96, 98, 103 

Ablania 284 

Abolboda 178-181 

Abrus 346, 356, 499 

Abuta 420-421 

Abutilon 393-394 

Acacallis 118, 134 

Acacia 365, 370, 372-376 

Acalypha 290-295 

Acanthella 397, 406, 495 

Acanthinophyllum 425 

Acanthocereus 231-232 

Acanthospermum 255, 260, 262, 264 

Acanthosphaera 425 

Acetosella 453 

Achetaria 522-523 

Achimenes 310 

Achlyphila 179 

Achnopogon 255, 260 

Achras 518-519 

Achrouteria 518 

Achyranthes 187-188 

Achyrocline 255, 260, 264 

Acianthera 134 

Acicarpa 158 

Acidoton 290, 295-297 

Acineta 118 

Acioa 241, 245-246 

Aciotis 397, 406-407, 410-411, 413- 

417 

Acisanthera 397, 406-411, 413-417 

Acmella 255-256, 260-262, 267 

Acnida 188 

Acoidium 134 

Acopanea 223 

Acosmium 346-347, 360, 363, 365 

Acosta 470 

Acouroa 356, 360 

Acrandra 438 

Acrocarpidium 461 

Acroceras 149, 165, 167 

Acrocomia 67, 69-70 

Acrodiclidium 326 

Acrostichum 50-51 

Actinostachys 52 

Actinostemon 290, 295, 297 

Acuan 370 

Ada 118 

Adelia 290, 299 

Adelobotrys 397, 407, 409-411, 415 

Adenanthe 445 

Adenanthera 365 

Adenarake 445 

Adeneleutherophora 134 

Adenobasium 284 


INDEX TO GENERA 

Adenocalymna 214, 217-218 

Adenolisianthus 304, 306-307 

Adenophaedra 290-300 

Adenostemma 261 

Adenostephanus 474 

Adhatoda 185 

Adiantopsis 22-23 

Adiantum 22-23, 51 

Adipe 134 

Adipera 340 

Adiscanthus 507 

Aechmea 73, 78-81, 83 

Aegiphila 543-546 

Aegopogon 149, 158 

Aeranthus 134 

Aeschrion 525 

Aeschynomene 342, 347, 356, 360, 

364 

Agalinis 523 

Aganisia 118, 134 

Agarista 285, 287 

Agati 356 

Ageratina 256, 263 

Ageratum 256, 261 

Agonandra 453 

Agrosticula 158 

Agrostis 158 

Ahouai 199 

Aiouea 322-323, 326-327, 330 

Aira 158 

Akentra 377 

Albizia 365, 370, 375-376 

Alchornea 290, 295-297 

Alchorneopsis 290, 295-296 

Alcoceratothrix 389 

Aldina 347, 356 

Alectoroctonum 296 

Alectra 522-524 

Alegria 538 

Alexa 347, 356 

Alexandra 356 

Alibertia 479, 492-493, 495-497, 503, 

505-506 

Alicastrum 425 

Alisma 59 

Aliteria 425 

Allamanda 195, 202 

Allania 356 

Allantoma 330-331 

Alloiosepalum 279 

Allophylus 511, 513-514 

Alloplectus 309-311 

Allosidastrum 393 

Allosorus 30 

Almeidea 508 

Alocasia 59, 64 

Alomia 261 

Alophia 111 

Alpinia 182 

Alseis 479, 492 

Alsodeia 548 

Alsophila 26-28 

Alstroemeria 112-113 

Alternanthera 187-188 

Althaea 394 

Alysicarpus 347, 360 

Amaioua 479, 492-493 

Amanoa 290, 296 

Amaranthus 187-188 

Amaryllis 113 

Amasonia 544-546 

Amauropelta 54 

Ambelania 195, 198-199, 203 

Amblyanthera 199 

Ambrosia 256, 261 

Amerimnon 356 

Ammannia 384-385 

Amomum 182 

Amorphospermum 518 

Ampelocera 542 

Ampelodaphne 326 

Ampelozizyphus 477 

Ampherephis 261 

Amphidasya 479,507 

Amphidesmium 44 

Amphilophium 214, 217-218 

Amphineuron 54 

Amphiodon 356 

Amphiphyllum 173, 175 

Amphirrhox 547-548 

Amphiscopia 185 

Amphizoma 314 

Amphymenium 340 

Amyris 507-508 

Anacampta 199 

Anacardium 189-190 

Anacheilium 134 

Anadenanthera 366, 372-374 

Anagallis 473 

Ananas 73, 79 

Ananthacorus 51, 55 

Anaphyllopsis 59, 64 

Anartia 198-199 

Anastrophus 158 

Anathallis 134 

Anatherum 158 

Anaxagorea 190, 193-195 

Ancistrocarpus 457 

Ancistrothyrsus 454 

Andira 347-348, 356, 360 

Andriapetalum 474 

Andropogon 149-150, 158-159, 161- 

162 

Androstylium 251 

Anechites 199 

Aneilema 85

566 

Anelasma 421 

Anemia 45, 52 

Anemopaegma 214, 217-219 

Anetium 50, 55 

Angadenia 199 

Angelonia 522 

Angostura 508-510 

Angostyles 296 

Angraecum 134-135 

Anguria 277 

Anguriopsis 277 

Aniba 323, 326-328, 330 

Anila 356 

Anisacanthus 183, 185 

Anisantherina 522-523 

Aniseia 271, 273-274 

Anisolobus 199 

Anisomeles 321 

Anisomeris 493 

Anisophyllea 190 

Anneslia 370 

Annona 190, 193-194 

Anomoctenium 217 

Anomospermum 420-421 

Anonocarpus 425 

Anoplophytum 79 

Anredera 213 

Anthaenantia 159 

Anthephora 150, 172 

Anthereon 135 

Anthericum 113 

Anthodiscus 236 

Anthodon 313-315 

Anthodus 314 

Anthurium 59-60, 64-66 

Antidaphne 285 

Antidesma 290, 296 

Antigonon 471 

Antirhea 479, 493 

Antonia 379-380 

Antrophyum 23, 55 

Apalanthe 111 

Apalatoa 340 

Aparisthmium 290, 295-296 

Apatitia 407 

Apatostelis 135 

Apeiba 538-539 

Aphanocarpus 479, 493, 499 

Aphelandra 183, 185-186 

Apinagia 466-468 

Apluda 159 

Apocaulon 507 

Apocynum 199 

Apodandra 296 

Apodanthera 277 

Apodanthes 476 

Apodina 382 

Apoleya 340 

Apophragma 306 

Appendicularia 397, 415 

Appunia 493 

Aptandra 449-450 

Apteria 83-84 

Apuleia 333, 340, 344, 358 


Aracamunia 118 

Arachniodes 31-33, 48, 54 

Araeococcus 73 

Aralia 11, 207 

Aranella 377 

Aratitiyopea 80, 179 

Arberella 150 

Archytaea 223-224 

Ardisia 430, 432-433 

Argemone 454 

Argythamnia 296 

Aristida 150, 159-161, 171 

Aristolochia 207-208, 421 

Arkezostis 277 

Arouna 340 

Arrabidaea 214-215, 217-219 

Arrhostoxylum 185 

Arrudea 251 

Artanthe 461 

Arthrosamanea 370 

Arthrostema 407 

Arthrostemma 407 

Arthrostylidium 150, 159, 161 

Aruba 525 

Arum 64 

Arunda 159 

Arundinaria 159 

Arundinella 150, 158-159, 169 

Arundo 150, 159 

Asclepias 208, 210-211 

Ascolepis 88, 95, 99 

Aspasia 118 

Aspegrenia 135 

Aspidium 32 

Aspidogyne 118, 140, 144 

Aspidosperma 195, 199-201 

Aspilia 261 

Asplenium 24-25, 32, 56 

Asplundia 87 

Astephanus 211 

Aster 261 

Asteranthos 330 

Asterogyne 67, 69 

Asterolepidion 319 

Asterophorum 539 

Astrocaryum 67, 69 

Astrococcus 290, 296 

Astronium 189 

Asystasia 183, 185-186 

Ateramnus 296 

Athenaea 302 

Athyrocarpus 85 

Atractantha 150 

Attalea 67, 69-72 

Aubletella 57, 518 

Augusta 261 

Aulizia 135 

Aulomyrcia 438-440 

Aulonemia 150, 159 

Austroamericum 511 

Austroeupatorium 256, 263 

Avesicaria 377 

Avicennia 544-545 

Axinanthera 407 

Axonopus 150-151, 158-161, 164-169 

Ayapana 256, 261, 263-264 

Aydendron 327 

Ayenia 531 

Ayensua 73, 178 

Azadirachta 417 

Azara 302 

Azolla 51 

Baccharis 256, 261-264, 266 

Baccharoides 261 

Bacopa 522-524 

Bactris 67-72 

Badula 432 

Bagassa 423, 425-426, 465 

Baillieria 261 

Balantium 244 

Balizia 366, 372, 374-376 

Ballota 321 

Baltimora 256, 261, 264 

Bambusa 160 

Banara 301-304 

Banisteria 389 

Banisteriopsis 386, 389-390, 392 

Barbacenia 178 

Barbieria 348, 356-357, 359 

Barbosella 118, 135 

Barjonia 208 

Barleria 183, 185 

Barnebydendron 333 

Barnhartia 468 

Barrosoa 256, 263 

Bartramia 539 

Basanacantha 493 

Basella 213 

Baskervilla 118, 135 

Bassovia 528 

Bastardia 393 

Batatas 273 

Batemannia 118 

Batesia 334 

Bathmium 53 

Bathysa 479, 495, 505 

Batis 213 

Batocarpus 423, 425 

Bauhinia 334, 340-341, 343, 345 

Beadlea 135 

Becquerelia 88, 95, 102 

Befaria 287 

Begonia 213-214 

Beilschmiedia 323, 327 

Bejaria 285, 287 

Bellardia 493 

Bellucia 397, 407, 410, 417 

Bellynkxia 493 

Beloglottis 118, 135 

Beloperone 183, 185 

Benitzia 84 

Benjaminia 522-524 

Bernardia 290, 300 

Bernardinia 270 

Bertholletia 330-331 

Bertiera 479, 493 

Bertolonia 397 

Besleria 309, 311-312

Beyrichia 523 

Bidens 256, 261 

Bifrenaria 118, 134-135, 148 

Bignonia 217-218 

Bihai 110 

Billardiera 493 

Billbergia 73, 79 

Binaria 341 

Biophytum 453 

Biovularia 377 

Bisboeckelera 88, 95, 98 

Bixa 220 

Blainvillea 256 

Blakea 397, 407, 417 

Blastemanthus 445, 448 

Blechnum 25-26, 47 

Blechum 183, 185-186 

Blepharandra 386, 390 

Blepharocalyx 434 

Blepharodon 208-209, 211-212 

Bletia 118, 135 

Blochmannia 472 

Blotiella 29, 31, 51 

Blutaparon 187-188 

Bocagea 193 

Bocageopsis 191, 193-194 

Bocoa 334, 363 

Boehmeria 542-543 

Boerhavia 442-443 

Bogenhardia 394 

Bolbitis 23, 42-43, 50-51 

Bolbophyllaria 135 

Bolboschoenus 95 

Bollea 118, 135, 141 

Bomarea 112-113 

Bombacopsis 220-222 

Bombax 221-222 

Bonafousia 198-200 

Bonamia 271, 273, 275 

Bonatea 135 

Bonnetia 223-224 

Bonpartea 79 

Bonplandia 509 

Bontia 428, 545 

Bonyunia 379-380 

Borismene 420-421 

Borojoa 493 

Borreria 479, 493, 498, 504 

Bothriochloa 151, 160 

Bothriospora 479, 496 

Botryarrhena 479 

Bouchea 544, 546 

Bougainvillea 442-443 

Bourreria 224-226, 477 

Boussingaultia 213 

Bouteloua 151, 159 

Bowdichia 348, 356 

Boyania 397 

Brachiaria 151, 160, 164-167, 172 

Brachionidium 118 

Brachistus 528 

Brachycodon 306 

Brachyloma 311 

Brachynema 450 


Brachypremna 407 

Brachypterys 390 

Brachystele 118, 140, 147 

Bradburya 356 

Braemia 118, 141, 146 

Bramia 523 

Brasenia 231 

Brassavola 118, 135, 138 

Brassia 118, 135, 138 

Brassica 226 

Bravaisia 183, 185-186, 545 

Bredemeyera 468-470 

Brewcaria 73-74, 80 

Brickellia 256, 262-263 

Brignolia 493 

Briquetia 393-395 

Briza 151 

Brocchinia 58, 74, 79 

Bromelia 74, 79-80 

Bromus 160 

Brosimopsis 425 

Brosimum 247, 423, 425-427 

Brossea 287 

Brownea 334, 341, 344 

Bruinsmania 493 

Brunellia 227 

Brunfelsia 526, 528 

Bryonia 277 

Bryophyllum 276 

Bubroma 532 

Buceras 254 

Buchenavia 226, 254-255 

Buchia 493 

Buchnera 522-524 

Bucholzia 188 

Bucida 254-255 

Buena 494 

Buforrestia 85 

Bulbophyllum 118-119, 135, 137 

Bulbostylis 88, 95-102 

Bulbulus 160 

Bumelia 518 

Bunchosia 386, 390-391 

Buphthalmum 261 

Burdachia 386, 390, 392 

Burlingtonia 135 

Burmannia 83-84 

Burmeistera 233 

Bursera 227, 229 

Burseria 229 

Buttneria 532 

Buxus 231 

Byrsonima 386-387, 389-391 

Byttneria 531-532 

Caballeria 432 

Cabomba 231 

Cabralea418 

Cabrera 160 

Cacalia 261 

Cacao 532 

Caconapea 523 

Cacoucia 255 

Cactus 232 

Caesalpinia 334, 341, 344-345 

Cajanus 348, 356, 358 

Caladium 60, 64 

Calathea 114-116, 182 

Caldasia 213 

Calea 256-257, 261-262, 264, 266-267 

Calinea 281 

Calliandra 366, 370-371, 373-374 

Callicarpa 545 

Callichlamys 215, 218-219 

Callicocca 494 

Callisia 85-86 

Calocarpum 518 

Calolisianthus 306 

Calonyction 273 

Calophyllum 247, 251 

Calophysa 407 

Calopogonium 348, 363 

Calotropis 209, 211 

Calpidisca 377 

Caluera 119 

Calycampe 439 

Calycobolus 271, 273-275 

Calycolpus 434, 439, 441 

Calycophyllum 479, 494, 503, 505 

Calycorectes 434, 439 

Calyptranthes 434-435, 439, 442 

Calyptrion 548 

Calyptrocarya 88, 95 

Calyptrostylis 95 

Calystegia 273 

Camarea 387 

Camaridium 135 

Cameraria 200 

Campelia 85 

Campnosperma 189 

Campomanesia 435, 439, 442 

Campsiandra 334-335, 341 

Camptouratea 448 

Campylocentrum 119, 134-135 

Campyloneurum 45-49 

Campylosiphon 83 

Cananga 193 

Canavalia 348, 356, 359 

Candolleodendron 348, 363 

Canna 84 

Cansenia 341 

Caopia 251 

Caperonia 290, 296 

Capirona 479, 494 

Capitellaria 407 

Capparidastrum 234 

Capparis 234-235 

Capraria 522-523 

Capriola 160 

Capsicum 526, 528 

Caraguata 79 

Caraipa 247-248, 251 

Carapa 418 

Carapichea 494 

Caraxeron 188 

Cardamine 226 

Cardiopetalum 191, 193 

Cardiospermum 511, 514 

Cardonaea 262

568 

Carex 88-89, 95-96 

Carica 236 

Carludovica 87 

Carolinea 222 

Carphobolus 262 

Carpodiptera 539 

Carpotroche 301-303 

Carptotepala 106 

Caryocar 236-237 

Caryochloa 160 

Caryomene 420 

Cascarilla 494 

Casearia 301-304, 548 

Casimirella 318-319 

Casinga 303 

Cassebeera 22, 24, 51 

Cassia 335, 341-343, 345 

Cassine 240 

Cassipourea 477 

Cassupa 494 

Cassytha 323, 327 

Castalia 444 

Castilla 423 

Castilleja 523 

Casuarina 237 

Catasetum 119, 135-136, 143, 149 

Catharanthus 196, 201, 203 

Cathartocarpus 343 

Cathedra 450 

Cathormion 370 

Catinga 439 

Catocoma 470 

Catopsis 74, 79, 81-82, 182 

Catostemma 220, 222, 539 

Cattleya 119, 136, 149 

Caularthron 136, 138-139 

Caulotretus 343 

Cavanillesia 220 

Cavendishia 285, 287-288 

Cayaponia 276-278 

Cecropia 237-238 

Cedrela 418-419 

Cedrelinga 366, 374 

Cedrus 539 

Ceiba 220, 222 

Celastrus 240 

Celianella 290 

Celiantha 304, 306-307 

Celosia 187-188 

Celtis 542 

Cenchrus 151, 160 

Centratherum 257, 261-262 

Centrogenium 136 

Centroglossa 136 

Centrolobium 348, 356 

Centronia 397, 407 

Centropogon 233 

Centrosema 348, 356-357, 362 

Centrosolenia 311 

Centrospermum 262 

Centunculus 473 

Cephaelis 494-495 

Cephalocarpus 89, 96 

Cephalocereus 232 


Cephalodendron 495 

Cephaloschoenus 96 

Cephalostemon 173, 175 

Cephalostigma 233 

Ceradenia 35-38, 47-49 

Ceratoccoca 457 

Ceratocephalus 262 

Ceratochilus 136 

Ceratophyllum 240 

Ceratophytum 215, 218 

Ceratopteris 45, 51 

Ceratosanthes 276-278 

Ceratostema 285, 287 

Cerbera 200 

Cercophora 331 

Cercouratea 448 

Cerdana 225 

Cereus 231-233 

Cespedesia 445, 448 

Cestrum 497, 526, 528 

Chaenanthe 136 

Chaenophora 407 

Chaenopleura 407 

Chaetaria 160-161 

Chaetocalyx 348, 357, 362 

Chaetocarpus 290, 296-297 

Chaetochlamys 185 

Chaetochloa 161 

Chaetocrater 303 

Chaetogastra 407 

Chaetolepis 407 

Chaetospora 96 

Chailletia 280 

Chalcas 509 

Chalepophyllum 479, 495 

Chamaecrista 335-336, 341-344 

Chamaedorea 70 

Chamaefistula 343 

Chamaesenna 343 

Chamaesyce 296 

Chamelophyton 136 

Chamissoa 187-188 

Chaptalia 257, 262, 264, 267 

Charianthus 407 

Chatiakella 262 

Chaubardia 119, 136, 140 

Chaubardiella 119, 136, 148 

Chaunochiton 450 

Cheilanthes 22-24, 29, 50-51 

Cheiloclinium 313-316 

Cheiradenia 119, 136 

Cheiroglossa 45 

Chelonanthus 304, 306-308 

Chelyorchis 120, 143 

Chenopodium 240 

Chevaliera 79 

Chimantaea 257, 262, 267 

Chimarrhis 479, 495, 499 

Chiococca 495, 497 

Chionanthus 451 

Chionolaena 257, 262, 264 

Chitonia 407 

Chloris 151, 158, 161 

Chlorocardium 7, 323, 329 

Chloroleucon 366, 373-375 

Chlorophora 425 

Chomelia 480, 493, 495 

Chondrococcus 495 

Chondrodendron 421 

Chorisepalum 304-306 

Christella 54 

Christiana 538-539 

Chromolaena 257, 263-264, 266 

Chromolucuma 515 

Chrysobalanus 241, 244 

Chrysochlamys 248, 253 

Chrysophyllum 57, 515, 518-521 

Chrysothemis 309, 311-312 

Chuncoa 255 

Chupalon 287 

Chuquiraga 262 

Chusquea 151, 161 

Chylodia 262 

Chytroma 331 

Chytropsia 495 

Cienfuegosia 393-395 

Cinchona 495 

Cinchonopsis 480, 495 

Cinnamodendron 234 

Cinnamomum 323, 327, 329 

Ciponima 535 

Cipura 111 

Cissampelos 420-421 

Cissus 551-553 

Citharexylum 544-545 

Citrosma 526 

Citrullus 276-278 

Citrus 507 

Cladium 89, 96 

Cladobium 136 

Cladocolea 381-384 

Cladostyles 273 

Clarisia 423, 425 

Clathrotropis 348, 357-358 

Clavapetalum 319 

Clavija 537 

Claytonia 306 

Cleidion 296 

Cleistes 120, 136, 141, 146 

Clematis 476 

Cleome 234-235 

Clerodendrum 544-545, 547 

Clethra 247 

Clibadium 257, 261-262 

Clidemia 397-398, 407-411, 414-416 

Clinopodium 321 

Clinostemon 327 

Clitoria 348-349, 357, 361, 364 

Clonodia 387, 390-392 

Clowesia 120, 135-136 

Clusia 248-249, 251-252 

Clusiella 249, 251 

Clusiophyllum 296 

Clytostoma 215, 217 

Cnemidaria 26-28, 48 

Cnestidium 269-270 

Cnidoscolus 290, 297 

Coccinia 276

Coccochondra 480, 495 

Coccocypselum 480, 493, 495, 497, 

502-503, 505 

Coccoloba 471-472 

Cocculus 421 

Cochleanthes 120, 136, 138 

Cochlidium 35-37, 47-49, 51 

Cochlospermum 220, 222 

Cocos 68, 70 

Codonanthe 309, 311-312 

Codonanthopsis 309-311 

Codonemma 200 

Coelorachis 151, 161, 163-164, 170 

Coeppertia not found 

Coffea 480, 495, 497 

Cohniella 136 

Coix 151 

Coleosanthus 262 

Coleostachys 387, 390 

Collaea 357 

Collophora 200 

Colocasia 60, 64 

Colubrina 477 

Columnea 309-312 

Comanthera 106 

Combretum 254-255, 382 

Commelina 85 

Commelinopsis 85 

Commianthus 495 

Commiphora 227, 229 

Comocladia 189 

Comolia 398-399, 407-408, 410, 414- 

417 

Comoliopsis 399 

Comparettia 120 

Complaya262 

Compsoneura 428-430 

Conami 296 

Conceveiba 290, 296, 300 

Conceveibastrum 296 

Conceveibum 296 

Conchocarpus 507-509 

Condalia 495 

Condylidium 257, 263-265 

Condylocarpon 196, 199-202 

Connarus 269-270 

Connellia 74, 79-81 

Conobea 523-524 

Conocarpus 254-255 

Conoclinium 262 

Conohoria 548 

Conomorpha 432 

Conoria 200 

Conosiphon 495 

Conostegia 399, 408 

Conuleum 526 

Convolvulus 271, 273-275 

Convulvulus 274 

Conyza 257, 262-264 

Copaiba 343 

Copaifera 336, 343 

Copaiva 343 

Copedesma 408 

Copernicia 68, 70 


Coptophyllum 52 

Corallocarpus 277 

Corchorus 538-539 

Cordia 224-226 

Cordiera 480, 492, 495-496 

Cordyline 58 

Coreopsis 262 

Cornutia 544-545 

Coronilla 357 

Coronopus 226 

Cortaderia 151, 159 

Coryanthes 120, 136, 140 

Corymborkis 120 

Corynostylis 547-549 

Corypha 70 

Coryphadenis 408 

Coryphothamnus 480, 498 

Corythophora 330-332 

Corytoplectus 309-311 

Cosmibuena 480, 494-496 

Cosmos 257, 261 

Costus 86, 182 

Cottendorfia 79-80 

Cotula 262 

Coublandia 357 

Couepia 241, 244-247, 251 

Couma 196, 200, 203 

Coumarouna 357 

Coupoui 496 

Couratari 330-333 

Courimari 284 

Couroupita 330, 332 

Coursetia 349, 357, 360, 362 

Coussapoa 237-239 

Coussarea 480, 482, 493, 496 

Coutarea 480, 496, 499 

Coutoubea 305-306 

Cranichis 120, 136 

Craniolaria 456 

Crantzia 311 

Crataegus 478 

Crateva 234-235 

Cremanium 4-8 

Cremastosperma 191, 193 

Cremastus 218 

Crematomia 226 

Cremersia 309 

Crenea 384-385 

Crepidospermum 227, 229 

Crepinella 207 

Crescentia 215 

Crinum 112-113 

Crotalaria 349, 357, 359 

Croton 290-291, 296-297 

Crudia 245, 336, 340, 343-346 

Cryptangium 96 

Cryptarrhena 120, 136 

Cryptocalyx 545 

Cryptocarya 323, 327 

Cryptocentrum 120, 136 

Cryptochloa 150, 161, 170 

Cryptophoranthus 136 

Cryptostegia 209 

Ctenanthe 114, 116 

Ctenardisia 430, 432-433 

Ctenitis 32-33, 52-53 

Ctenium 151 

Ctenopteris 36 

Cucullaria 554-555 

Cucumis 276-277 

Cucurbita 276-277 

Cuervea 313-315 

Culcita 31 

Cunuria 296-297 

Cupania 511, 514 

Cuphea 384-385 

Cupirana 496 

Curanga 523 

Curarea 420-421 

Curatella 280-281 

Curculigo 112-113 

Curtia 305-306, 308 

Cuscuta 279 

Cusparia 509 

Cuspidaria 215 

Cyanthillium 257, 261-262, 268 

Cyathea 26-28, 48-49 

Cyathoglottis 136 

Cyathula 187-188 

Cybianthus 205, 430-434 

Cybistax 215, 218 

Cycas 56 

Cyclanthus 87 

Cyclodium 31-34, 49 

Cyclolobium 357 

Cyclopeltis 49, 52 

Cyclopogon 120, 135-136, 144, 146- 

147 

Cyclosorus 54 

Cycnoches 120, 136 

Cydista 215, 217-218 

Cylindrosperma 200 

Cymbidium 136 

Cymbocarpa 83 

Cymbopetalum 191, 193-194 

Cymbopogon 151, 161 

Cymbosema 349, 358 

Cynanchum 209, 211-212 

Cynodendron 518 

Cynodon 151, 160, 165 

Cynometra 336, 344 

Cynomorium 213 

Cynosurus 161 

Cypella 111 

Cyperus 89-90, 95-96, 98-99, 101-102 

Cyphomandra 526, 528, 530 

Cypripedium 136 

Cyrilla 279 

Cyrillopsis 319-320 

Cyrtocarpa 189-190, 229 

Cyrtocymura 257, 262, 268 

Cyrtopera 136 

Cyrtopodium 120, 136 

Cyrtosperma 60, 64-65 

Cyrtospermum 189 

Cytisus 358 

Dacryodes 227, 229, 231 

Dactyloctenium 151, 161

570 

Dactylostegium 185 

Dactylostemon 297 

Dahlgrenia 70 

Dalbergaria 311 

Dalbergia 349, 356, 358-360, 362 

Dalechampia 291-292, 297-299 

Danaea 44, 50 

Daphnopsis 537-538 

Dasynema 284 

Dasyphyllum 257, 262 

Datura 526, 528 

Davallia 29 

Davalliopsis 39 

Davilla 280-281 

Davya 409 

Debraea 555 

Decagonocarpus 507, 509 

Decaraphe 409 

Declieuxia 226, 480, 496-497 

Degranvillea 120 

Deguelia 358 

Delilia 257 

Delonix 336, 345 

Dendrobangia 318-319 

Dendrobium 136 

Dendropanax 205, 207 

Dendropemon 383 

Dendrophthora 383, 549-551 

Dendrosipanea 480-481, 496 

Dendrostylis 303 

Dendrothrix 292, 300 

Dennstaedtia 29-31 

Deprea 527-528 

Dermatophyllum 262 

Derris 349, 358, 360 

Desmanthus 366, 370 

Desmochaeta188 

Desmodium 349-350, 358, 360-361 

Desmoncus 68, 70 

Desmoscelis 399, 407, 409, 411, 414- 

415 

Diacidia 387, 390, 392 

Diacrium 136 

Diadenium 120, 136 

Dialium 336, 340, 344 

Diandrochloa 161 

Dianella 113 

Dianthera 185 

Diastema 309 

Diazeuxis 262 

Dibrachion 358 

Dicella 387, 390, 392 

Dichaea 120-121, 136-138, 141 

Dichanthelium 151, 164-167 

Dichanthium 151, 158 

Dichapetalum 226, 279-280 

Dichorisandra 85 

Dichromena 96-97, 101, 121 

Dichrostachys 366 

Dicksonia 31 

Diclidanthera 469-470 

Diclinanona 191, 193-194 

Dicliptera 183, 185 

Dicorynia 336, 344 


Dicrananthera 409 

Dicranoglossum 46-47, 49 

Dicranopteris 34 

Dicranopygium 87 

Dicranostyles 271, 274-275 

Dicrobotryum 496 

Dicrypta 137 

Dictyocaryum 68, 70-71 

Dictyostega 83-84 

Dicymbe 336 

Dicypellium 323 

Didactyle 137 

Didymiandrum 90, 96-97 

Didymochlaena 23, 31-32 

Didymochlamys 481 

Didymoglossum 39 

Didymopanax 207 

Diectomis 161 

Dieffenbachia 60, 64-66 

Digitaria 151-152, 158, 161, 165-168, 

170-172 

Digomphia 215, 218 

Dikylikostigma 137 

Dilkea 454-455 

Dilodendron 511, 514 

Dimerandra 121, 137, 139, 141, 143 

Dimerocostus 86 

Dimorpha 344 

Dimorphandra 336-337,344 

Dinizia 337 

Dioclea 350, 356, 358-359 

Diodella 496 

Diodia 481, 492-493, 496-497, 504 

Diolena 409 

Diomma 509 

Dioscorea 103-104 

Diospyros 282-283 

Diothonea 137 

Diphasium 18 

Diplacrum 90, 97, 99, 102 

Dipladenia 200 

Diplasia 90, 97 

Diplazium 25, 54-56 

Diplochita 409 

Diplocrater 450 

Diploon 515, 518 

Diplopterygium 34 

Diplopterys 387, 389-391 

Diplotropis 345, 350, 356, 358 

Dipteracanthus 185 

Dipteranthus 121, 137, 144 

Dipterodendron 514 

Dipteryx 350, 357-358, 363 

Disciphania 420-421 

Discocarpus 292, 297 

Discophora 318-319 

Discyphus 121, 137, 147 

Disocactus 231, 233 

Disteganthus 74, 78-80, 

Disterigma 285-286, 288 

Distictella 215, 217-218 

Distictis 215, 217-219 

Ditassa 209, 211 

Ditaxis 292, 296 

Ditmaria 555 

Ditomaga 306 

Ditremexa 344 

Dodecas 385 

Dodecastigma 292, 297-298 

Dodonaea 511 

Dolicholus 359 

Dolichopterys 390 

Dolichos 359 

Doliocarpus 280-282 

Dorstenia 423, 425 

Doryopteris 22-24, 51 

Doyerea 276-277 

Dracontium 60-61, 65 

Drejera 185 

Drepanocarpus 359 

Drepanospermum 189 

Drimys 555 

Drosera 282 

Drupina 523 

Dryadella 121 

Drymaria 237 

Drymonia 309-311 

Drynaria 47 

Dryopteris 32-33 

Drypetes 292, 297 

Duboisia 137 

Duckea 97, 101, 173, 175 

Duckeella 121, 137 

Dufourea 274, 468 

Dugandiodendron 385-386 

Duguetia 191, 193 

Duidaea 257 

Duidania 481 

Dulacia 450-451 

Dunstervillea 121 

Duranta 544-546 

Duroia 481, 492-493, 496 

Dussia 350, 359-360 

Duvernoia 185 

Dyckia 80 

Dyspemptemorion 185 

Dystovomita 249 

Ebermaiera 185 

Ecastaphyllum 359 

Ecclinusa 515, 518-520 

Eccremis 112-113 

Echeandia 112-113 

Echinella 137 

Echinocassia 344 

Echinocephalum 262 

Echinochloa 152, 164-167 

Echinodorus 59 

Echinolaena 152, 160, 165 

Echinops 262 

Echinoschoenus 97 

Echinosepala 137 

Echites 200-201 

Eclipta 257, 262, 267-268 

Ectozoma 528 

Egleria 90 

Egletes 257, 262 

Eichhornia 173 

Elachyptera 313-314, 316

Elaeagia 481 

Elaeis 68 

Elaeodendron 239-240 

Elaeoluma 433, 515, 518-520 

Elaeophora 297 

Elaphandra 257 

Elaphoglossum 42-43, 45, 50-51 

Elaphrium 229 

Elaterium 277 

Eleocharis 101-102 

Elephantomene 421 

Elephantopus 257, 262-263 

Eleusine 152, 161 

Eleutheranthera 257, 264-265, 268 

Eleutherine 111 

Elionurus 152, 158, 161, 163 

Elizabetha 337, 244 

Elleanthus 121, 134, 137, 140, 147 

Elodea 110-111 

Elopium 65 

Eloyella 121, 144 

Eltroplectris 121, 136, 143-144 

Elvasia 445, 448-449 

Elytraria 183, 186 

Emilia 257, 261, 263 

Emmeorhiza 481, 493 

Emmotum 318-319 

Encholirium 80 

Enckea 461 

Encyclia 121, 136-139, 141 

Endlicheria 323, 326-329 

Endopleura 316, 318 

Englerella 519 

Enicostema 305-306 

Enourea 514 

Entada 366, 370-371, 373 

Entadopsis 370-371 

Enterolobium 366, 371, 375 

Enteropogon 152, 161-162 

Enterosora 35-37, 47-49 

Eperua 337, 344-345 

Ephedranthus 191, 193 

Ephippiorhynchium 97 

Epidendropsis 138 

Epidendrum 121-123, 135, 137-139, 

141, 143-144, 147 

Epidryos 173 

Epigaea 287 

Epiphyllum 231-233 

Episcia 310-311 

Epistephium 123, 139-140 

Eragrostis 152, 161-164, 169-170 

Eranthemum 183, 185 

Erechtites 257, 266 

Eremolepis 285 

Eremoluma 519 

Erianthus 162 

Erigeron 257, 263 

Erinus 523 

Eriocaulon 104-107 

Eriochloa 152, 162, 164 

Eriochrysis 152, 158, 170 

Eriope 321 

Eriopsis 123, 140, 146 


Eriosema 350, 358-360, 362 

Eriosorus 23-24, 36 

Eriotheca 220-222 

Erisma 553, 555 

Ernestia 399, 407, 409, 415 

Erosion 162 

Eroteum 536 

Ertela 507, 509 

Ervatamia 201 

Erycina 123, 139, 143, 146 

Eryngium 195 

Erythrina 350, 359, 361 

Erythrochiton 507 

Erythrodes 123, 140, 144 

Erythroxylum 288-289 

Eschatogramme 47 

Eschweilera 330-333 

Escobedia 523 

Esenbeckia 507, 509 

Esmeraldia 211 

Etaballia 350, 359-360 

Ethulia 263 

Eubrachion 285 

Euceraea 301 

Euchlaena 162 

Eugenia 435-436, 438-442 

Eulophia 123, 136, 141 

Eulophidium 140 

Euosmia 496 

Eupatorium 263-264 

Euphorbia 292, 296-297, 299 

Euphronia 301 

Euplassa 474 

Eurya 536, 542 

Eurychaenia 409 

Eurydochus 264 

Eurystyles 123, 148 

Euterpe 68, 70 

Evea 496 

Evelyna 140 

Everardia 91, 97 

Evodianthus 87-88 

Evolvulus 271, 273-274 

Evonymodaphne 327 

Exacum 306 

Excentradenia 387, 391 

Exellodendron 241, 244, 246 

Exochogyne 91, 97, 99 

Exoecaria 425 

Exostyles 337 

Exothostemon 201 

Fagara 509 

Faramea 481-482, 495-497, 504 

Farringtonia 409 

Feea 39 

Ferdinandusa 482 

Fernandezia 140 

Ferolia 244, 425 

Festuca 162 

Feuillea 371 

Feuilleea 371 

Fevillea 276-278 

Ficus 423-428 

Fimbristylis 91, 97-99, 101-102 

Fischeria 209, 211 

Fissicalyx 350, 361 

Fittonia 185 

Flacourtia 303 

Fleischmannia 257, 263 

Flemingia 350 

Fleurya 542 

Floscopa 85 

Folianthera 371 

Forsteronia 196, 199, 201, 203 

Fothergilla 409 

Fractiunguis 140 

Franchetella 519 

Franciscea 528 

Freziera 535, 536 

Froelichia 187-188 

Froesia 475 

Froesiochloa 152 

Froesiodendron 193 

Fugosia 394 

Fuirena 91, 98, 102 

Funastrum 211 

Furarium 382 

Furcraea 58 

Fusaea 191, 193-194 

Gaedawakka 297 

Gaiadendron 381-384 

Galactia 350-351, 357, 359-360 

Galactodendrum 426 

Galactophora 196, 199-200 

Galeandra 123-124, 140, 144 

Galega 359 

Galeottia 124, 143, 148 

Galipea 507, 509 

Galium 482, 497, 502-503, 505 

Galphimia 390 

Gamochaeta 258, 264 

Gamotopea 497 

Garapatica 497 

Garayella 140 

Garcinia 250, 252 

Gardenia 497 

Garuga 189 

Gaultheria 286-288 

Gaura 255 

Gavarretia 292, 296 

Gaya 393 

Gaylussacia 286-287 

Gayoides 394 

Geissopappus 264 

Geissospermum 196 

Genipa 482, 497, 507 

Genlisea 376-377 

Gentiana 306 

Geoffroea 360 

Geonoma 68, 70-71 

Geophila 482, 497-498, 501, 503 

Gerardia 523 

Gesneria 311 

Ghinia 546 

Gibasis 85-86 

Gilibertia 207 

Gimbernatea 255 

Gitara 297

572 

Glandonia 387 

Glaphyropteris 54 

Glaziostelma 211 

Gleasonia 482, 497 

Gleichenia 34 

Glinus 422 

Gliricidia 351, 362 

Globifera 523 

Glochidion 297 

Gloeospermum 547-548 

Glossarion 258, 264 

Glossocentrum 409 

Glossostylis 523 

Gloxinia 310-311 

Glycine 360 

Glycoxylon 519 

Glycydendron 292 

Glyphotaenium 36 

Gnaphalium 258, 264 

Gnetum 57 

Gochnatia 258, 262, 264, 266-267 

Godmania 215, 217 

Godoya 448 

Goeppertia 327 

Gomidesia 440 

Gomphia 448 

Gomphichis 124, 148 

Gomphiluma 519 

Gomphrena 187-188 

Gongora 124, 140 

Gongylolepis 258, 261-262, 264, 267 

Goniophlebium 47 

Goniopteris 54 

Gonolobus 209, 211 

Gonzalagunia 482, 497 

Goodallia 537 

Goodyera 140 

Gordonia 536-537 

Gossypium 393-394 

Gouania 477 

Gouinia 152, 172 

Goupia 239-240 

Govenia 124 

Graffenrieda 399, 409, 415 

Grammadenia 433 

Grammitis 35-38, 47-49 

Granatum 418 

Gratiola 523 

Gravisia 80 

Greeneina 426 

Greenmania 264 

Grias 332 

Grimaldia 344 

Grislea 385 

Guacamaya 173 

Guadua 152, 160, 162 

Guaiacum 556 

Guaicaia 264 

Guanchezia 124, 135 

Guapeba 519 

Guapira 443-444 

Guarania 297 

Guarea 304, 418-419 

Guatteria 191-194 


Guatteriopsis 192-193 

Guayania 258, 262-264 

Guazuma 531 

Guenetia 539 

Guettarda 482-483, 493, 496-497, 507 

Guidonia 303 

Guildingia 409 

Guilielma 71 

Gurania 276-278 

Gustavia 331-333 

Guzmania 74, 79-82, 182 

Gymnanthes 292, 296-297 

Gymnobalanus 327 

Gymnogramma 23 

Gymnogramme 55 

Gymnopogon 152, 161-162, 170 

Gymnopteris 23 

Gymnosiphon 83-84 

Gynandropsis 235 

Gynerium 152, 162, 170 

Gynestum 71 

Gynocraterium 183 

Habenaria 124-125, 135, 140, 144 

Hackelochloa 152, 160, 163 

Haemadictyon 201 

Haematostemon 292, 296 

Haematoxylum 337 

Haemocharis 536 

Haloragis 312 

Haloschoenus 98 

Hamelia 483, 497 

Hancornia 201 

Handroanthus 218 

Haploclathra 249, 251 

Haplolophium 215 

Haplostylis 98 

Hartigia 409 

Hasseltia 301 

Havetia 252 

Havetiopsis 252 

Hawkesiophyton 527-528 

Hebanthe 188 

Hebeclinium 258, 261, 263 

Hebepetalum 316 

Hecastophyllum 360 

Hecatostemon 301-303 

Hecistopteris 55 

Heckeria 461 

Hedwigia 229 

Hedychium 182 

Hedyosmum 240 

Hedyotis 485, 497 

Hedysarum 360 

Heisteria 450-451 

Helia 306-307 

Heliamphora 521-522 

Heliconia 110, 117 

Helicostylis 424, 426-427 

Helicteres 531-532 

Heliotropium 225-226 

Helmia 104 

Helmontia 276-278 

Helonoma 125, 135-136, 142, 147 

Helopus 162 

Helosis 212-213 

Hemarthria 152, 170 

Hemicarpha 98 

Hemicrepidospermum 229 

Hemidictyum 25, 56 

Hemidiodia 497 

Hemigraphis 183 

Hemionitis 23, 51 

Hemitelia 28 

Henriettea 399-400, 409-411, 414 

Henriettella 400, 409-411, 414 

Henriquezia 483, 497 

Hepetis 80 

Herissantia 393-395 

Hermesia 297 

Hermesias 344 

Hernandia 312 

Herpestis 523-524 

Herpetacanthus 183, 185 

Herrania 531-533 

Hesperomeles 478 

Heteranthera 173 

Heterocondylus 258, 264 

Heteropetalum 192, 194 

Heteropogon 152, 158, 162 

Heteropsis 61, 65-66 

Heteropterys 387-391 

Heterostemon 337, 344 

Heterotaxis 125, 137, 142-143 

Heterothalamus 264 

Heterotrichum 409 

Hevea 292, 297, 300 

Hexadesmia 140 

Hexapterella 83-84 

Hexisea 125, 137, 140 

Heymassoli 450 

Hibiscus 393-395 

Hicriopteris 34 

Hieronyma 292, 297, 300 

Hillia 483, 497, 503 

Himatanthus 196, 201-202 

Hippeastrum 112-113 

Hippobroma 233 

Hippocratea 313-314 

Hippomane 297 

Hiraea 388-389, 391 

Hirtella 241-242, 244-247 

Hisingera 303 

Histiopteris 29, 51 

Hoehneella 140 

Hoffmannia 483 

Hohenbergia 74, 79 

Holcus 162 

Holosteum 237 

Holstianthus 483 

Homalium 301-304 

Homalocenchrus 162 

Homalomena 61 

Homoeotes 39 

Homolepis 152-153, 164-166 

Hoppia 98 

Hormocalyx 409 

Hornschuchia 194 

Hortia 507

Hortsmania 201 

Hostmannia 448 

Houlletia 125, 141 

Houmiri 317 

Houssayanthus 511, 514-515 

Houstonia 497 

Huberodendron 220 

Huberopappus 258 

Huebneria 141 

Humboldtia 141 

Humboldtiella 360 

Humiria 316-317 

Humirianthera 319 

Humiriastrum 317-318 

Humirium 317 

Huntleya 125, 141 

Huperzia 17-19 

Hura 292 

Hybanthus 547-549 

Hybochilus 141 

Hydranthelium 524 

Hydrochorea 366, 370-371, 373-376 

Hydrocleys 114 

Hydrocotyle 195 

Hydrolea 318 

Hydromystria 111 

Hygrophila 183, 185 

Hylaea 196, 202 

Hylaeanthe 114, 116 

Hylaeorchis 125, 135, 142 

Hylenaea 313-314 

Hylocereus 231-233 

Hymenachne 153, 162, 164-165 

Hymenaea 337, 344 

Hymenocallis 112-113 

Hymenodium 43 

Hymenolobium 351 

Hymenophyllopsis 25, 40-41 

Hymenophyllum 38-41 

Hymenostachys 40 

Hyospathe 68, 71 

Hyparrhenia 153, 158, 161, 172 

Hypenia 321, 322 

Hyperbaena 421 

Hypericum 249, 252 

Hypobrichia 385 

Hypocyrta 312 

Hypogynium 162 

Hypolepis 23, 29 

Hypolytrum 91-92, 98, 101-102 

Hypoporum 98 

Hypoxis 112-113 

Hyptidendron 321 

Hyptis 321-322 

Ibatia 212 

Ibetralia 483, 492 

Icacorea 433 

Ichnanthus 153, 159, 162-166 

Ichthyophora 519 

Ichthyothere 258, 264, 266 

Icica 229 

Icicopsis 229 

Ilex 203-205, 426 

Illecebrum 188 


Ilysanthes 524 

Imeria 258, 263-264 

Impatiens 213 

Imperata 153, 163, 170 

Indigofera 351, 356, 360 

Inga 366-367, 371-374 

Inula 264 

Ionidium 548 

Ionopsis 125, 139, 141 

Ipomoea 271-275 

Iresine 187-188 

Iria 98 

Iriartea 68, 71 

Iriartella 68, 71 

Iris 111 

Irlbachia 305-308 

Iroucana 303 

Iryanthera 429-430 

Isachne 153, 163, 166-167 

Ischaemopogon 163 

Ischaemum 153, 158-163 

Ischnosiphon 114-116 

Iseia 272-275 

Isertia 483, 493-494, 497, 505 

Isidrogalvia 112-113 

Isochilus 125, 139,141 

Isoëtes 17 

Isolepis 98 

Isoloma 312 

Isotoma 233 

Itea 279 

Ithycaulon 30 

Ivira 532 

Ixia 111 

Ixidium 285 

Ixocactus 381-382 

Ixophorus 153 

Ixora 483, 497, 499 

Jablonskia 292, 299-300 

Jacaranda 215, 217-218 

Jacaratia 236 

Jacobinia 185 

Jacquemontia 273-275 

Jacqueshuberia 337 

Jacquinia 537 

Jacquiniella 125, 138-139 

Jambosa 440 

Jamesonia 23 

Japarandiba 332 

Jasarum 61 

Jasminum 451 

Jatropha 292-293, 297 

Jenmaniella 467 

Jessenia 71 

Jubelina 388, 390-391 

Jucunda 409 

Jugastrum 332 

Juglans 320 

Julocroton 297 

Juncus 111-112 

Jupunba 372 

Juruasia 185 

Jussiaea 452 

Justicia 183-186 

Kaieteurea 448 

Kalanchoe 276 

Kallstroemia 556 

Karatas 80 

Kedrostis 276 

Kefersteinia 125, 141 

Kegelia 141 

Kegeliella 125, 141 

Kieseria 224 

Kippistia 314 

Klaprothia 379 

Kleinia 264 

Klugiodendron 372 

Knoxia 497 

Koanophyllon 258, 263-264 

Koellensteinia 125, 136, 141-142, 146, 

149 

Koellikeria 310 

Koernickanthe 115-116 

Kohleria 310-312 

Kopsia 196 

Koyamaea 92 

Krameria 320 

Krugia 440 

Kubitzkia 329 

Kuhlia 303 

Kuhlmaniella 275 

Kuhnia 264 

Kummeria 319 

Kunhardtia 173, 175 

Kutchubaea 483 

Kyllinga 92, 96, 98-99 

Labatia 519 

Lablab 351, 359 

Lacistema 320-321, 463 

Lacmellea 196, 201, 203 

Lacostea 40 

Lacunaria 475 

Ladenbergia 483, 495, 497 

Laelia 141 

Laetia 302-304 

Lafoensia 385 

Lagascea 264 

Lagenaria 276-278 

Lagenocarpus 92, 98-99, 102 

Lagerstroemia 385 

Laguncularia 254-255 

Lampadaria 310 

Lamprococcus 80 

Landolphia 201 

Lanessania 426 

Langsdorffia 212-213 

Lanium 141 

Lantana 544, 546 

Laplacea 536-537 

Laportea 542-543 

Larentia 111 

Lasiacis 153, 163-167 

Lasiadenia 537 

Lasiandra 409 

Lasiolepis 107 

Lasiostoma 380 

Lastrea 54 

Lastreopsis 32-33, 48, 52-53

574 

Latreillea 264 

Laurea 426 

Laurembergia 312 

Laurentia 233 

Laurus 327, 538 

Lawsonia 385 

Laxoplumeria 196 

Leandra 400, 407-411, 414-417 

Leaoa 141 

Lecanium 40 

Lecointea 337 

Lecostemon 476 

Lecticula 377 

Lecythis 193, 331-333 

Ledothamnus 286-287 

Leersia 153, 162-164 

Leiostegia 410 

Leiothamnus 307 

Leiothrix 105-107 

Leiphaimos 307 

Leitgebia 448 

Lellingeria 35-38 

Lembocarpus 310 

Lemna 112 

Lemniscia 317 

Leochilus 125, 139 

Leonia 547-548 

Leonotis 321-322 

Leonurus 321-322 

Leopoldinia 68-69 

Lepanthes 125, 141 

Lepanthopsis 125, 144-145, 148 

Lepianthes 460-462 

Lepidagathis 184-186 

Lepidaploa 258, 262, 264, 268 

Lepidium 227 

Lepidocaryum 69, 71 

Lepidocordia 225 

Lepidotis 18 

Leptochilus 43 

Leptochloa 153, 161-163 

Leptocoryphium 153, 159, 165, 169 

Leptoglottis 372 

Leptolobium 344, 360 

Leptorchis 141 

Leptorhoeo 85 

Leptoschoenus 99 

Leptostachya 186 

Leptostachys 163 

Leretia 319 

Leria 264 

Lessingianthus 258, 268 

Lettsomia 536 

Leucaena 367, 370, 372-373 

Leucococcus 542 

Leucohyle 125, 141, 148 

Leuconymphaea 444, 445 

Leucopholis 264 

Leucothoe 287 

Libidibia 344 

Licania 242-246, 251, 533 

Licaria 323-324, 326-329 

Ligea 468 

Ligeophila 125, 140, 144 


Lightia 301, 532 

Limnobium 51, 110-111 

Limnocharis 114 

Limnosipanea 483, 497, 504 

Limodorum 141 

Lindackeria 302-303 

Lindernia 523-524 

Lindleya 537 

Lindleyella 141 

Lindmania 74-75, 79-80 

Lindsaea 23, 29-31 

Linociera 451 

Liparis 125, 141, 144, 148 

Lipocarpha 92, 95, 98-99, 101 

Lipochaeta 264 

Lippia 144, 545-546 

Liriosma 450-451 

Lisianthus 307-308 

Lissocarpa 379 

Lisyanthus 308 

Lithachne 153, 164 

Lithocardium 226 

Litobrochia 51 

Lobelia 233 

Lochnera 201 

Lockhartia 125-126, 138, 140-141 

Loevigia 410 

Lomagramma 43, 48 

Lomaria 26 

Lomaridium 45 

Lomariopsis 26, 43 

Lonchitis 30-31 

Lonchocarpus 351, 356, 358, 360-364 

Lonicera 497 

Lophanthera 388, 390 

Lophiaris 141 

Lophidium 52 

Lophogyne 468 

Lophopterys 388, 390-391 

Lophostoma 537 

Lophotocarpus 59 

Lopimia 395 

Loranthus 382, 550 

Loreya 400 

Lorostemon 249 

Loxopterygium 189 

Lubaria 509 

Lucilia 264 

Lucuma 519 

Ludovia 87-88 

Ludwigia 451-452 

Lueddemannia 126, 136 

Luehea 538-539 

Lueheopsis 538-539 

Luetzelburgia 351 

Luffa 276-278 

Lunania 302 

Lundia 215-218 

Luxemburgia 448 

Luziola 153, 160, 163 

Lycaste 126, 141-142 

Lycianthes 527-530 

Lycomormium 14 

Lycopodiella 18-19 

Lycopodium 18-19 

Lycoseris 258, 262, 264 

Lycurus 163 

Lygodium 43-44 

Lyroglossa 126, 147 

Lysiloma 368, 372 

Lysiostyles 272, 275 

Lythrum 385 

Maba 283 

Mabea 293, 297-298 

Maburea 450 

Macaglia 201 

Macahanea 57, 315 

Macairea 400-401, 407, 410, 414-416 

Macfadyena 216, 218 

Machaerina 92 

Machaerium 351-352, 359, 361-362 

Machaonia 483, 495 

Maclura 424-426 

Macoubea 197, 201-203 

Macoucoua 205 

Macradenia 126, 141, 147 

Macrocarpaea 305, 308 

Macrocentrum 401, 409-410, 415 

Macrochordion 80 

Macroclinium 126, 141, 143 

Macrocnemum 497 

Macroditassa 209 

Macrolobium 57, 337-338, 344-346, 

372 

Macropharynx 197, 200 

Macroptilium 352, 362 

Macrosamanea 368, 371-372, 374-376 

Macroscepis 212 

Macuillamia 524 

Madarosperma 212 

Magnolia 385 

Maguirea 65 

Maguireanthus 401 

Maguireocharis 483 

Maguireothamnus 483, 495, 497-498 

Mahurea 249, 252 

Maieta 401, 410-411, 417 

Malache 395 

Malachra 393, 395, 532 

Malanea 483-484, 498 

Malaxis 126, 142 

Mallophyton 401 

Malmea 192, 194 

Malouetia 197, 200-202 

Malpighia 388, 391 

Malvastrum 393 

Manabea 546 

Managa 57 

Manaosella 216-217 

Mandevilla 197, 199-202 

Manettia 484, 498 

Manganaroa 372 

Mangifera 189 

Manglilla 433 

Manicaria 69, 71 

Manihot 293, 297-298 

Manilkara 515-516, 518-519 

Manisuris 163

Manniella 142 

Mansoa 216-219 

Mapania 92, 97-99 

Mapaniopsis 99 

Mapouria 498 

Mappia 319 

Maprounea 293, 298 

Maquira 424, 426-427 

Marahuacaea 173, 175 

Maranta 115-116 

Marathrum 467-468 

Marattia 44 

Marcetia 401, 410, 414-415 

Marcgravia 396 

Marcgraviastrum 396 

Margaritaria 293, 299 

Margaritopsis 498 

Marginaria 47 

Marialvaea 252 

Marila 249 

Maripa 272, 275 

Mariscus 99 

Markea 527-528 

Markleya 71 

Marlierea 436-437, 439-442 

Marmaroxylon 372 

Marsdenia 209-212 

Marsea 264 

Marsilea 44 

Marsupiaria 142 

Marsypianthes 321-322 

Martia 344 

Martinella 216, 218-219 

Martiodendron 338, 340, 344 

Martiusia 344 

Mascagnia 388-392 

Masdevallia 126, 142 

Matayba 511-512, 514-515 

Matelea 199, 210-212 

Matisia 220, 222 

Matourea 524 

Mattuschkea 498 

Mauria 189 

Mauritia 69, 71 

Mauritiella 69, 71 

Maxillaria 126-127, 135-137, 142-144, 

146-147 

Maximbignya 69, 71 

Maximiliana 71 

Maximilianea 220 

Mayaca 117 

Mayapea 451 

Mayepea 451 

Mayna 302-303 

Maytenus 239-240 

Meborea 298 

Mecardonia 523-524 

Mecodium 40 

Medadenium 143 

Mediocactus 232 

Megalastrum 32-33, 49, 52-53 

Megalostylis 298 

Megaskepasma 184 

Megastachya 163 


Meibomia 361 

Meioluma 519 

Meisneria 410 

Melaleuca 437 

Melampodium 264-265 

Melananthus 527 

Melanoxylon 344 

Melanthera 258, 261-262, 265 

Melasma 524 

Melastoma 410-411 

Melia 418 

Meliandra 411 

Melicoccus 512, 514-515 

Melidiscus 235 

Melinis 153, 163 

Meliosma 510 

Mella 524 

Melloa 216, 218 

Melocactus 232 

Melochia 531-533 

Melothria 277-278 

Melpomene 35, 37-38 

Melvilla 385 

Memora 216-219 

Menadenium 143 

Mendoncella 143 

Mendoncia 311, 419-420, 423 

Mendozia 420 

Meniscium 54 

Mentzelia 379 

Menyanthes 422 

Meoschium 163 

Meresaldia 212 

Meriania 401, 407, 409, 411 

Merizadenia 201 

Merostachys 153 

Merremia 272-275 

Mertensia 34-35 

Merumea 484 

Mesadenella 127, 143 

Mesechites 197, 201 

Mesosetum 153, 163-167 

Mesosphaerum 322 

Mespilodaphne 327 

Mespilus 478 

Messerschmidia 226 

Metalepis 210 

Metastelma 212 

Metaxya 27, 32, 44, 49 

Metrodorea 507 

Meyenia 186 

Mezia 388, 390, 392 

Mezilaurus 324, 326-327 

Mezobromelia 75, 80-83 

Miconia 401-404, 407-414, 416-417 

Micrandra 293, 296-298 

Micranthemum 523-524 

Microchloa 153 

Microgonium 40 

Microgramma 46-49, 51 

Microlepia 30 

Microlicia 404, 413 

Micropapyrus 99 

Micropholis 516, 518-521 

Microphysca 413 

Microplumeria 197, 199-201 

Micropolypodium 35-37 

Micropteryx 361 

Micropyxis 473 

Microstachys 293, 298, 300 

Microtea 457 

Miersiella 83-84 

Mikania 258-259, 261, 263, 265, 268- 

269 

Milium 164 

Miltoniopsis 127 

Mimosa 368-370, 372-374, 376 

Mimusops 516, 518-519 

Minquartia 450-451 

Miocarpus 413 

Mirabilis 443 

Misanteca 327 

Mitostemma 454 

Mitozus 201 

Mitracarpus 484, 498, 504 

Mitranthes 440 

Mitreola 379 

Mitrospora 99 

Mnasium 175, 177 

Mnesithea 164 

Moeroris 298 

Mollia 538-539 

Mollinedia 422 

Mollugo 422 

Molongum 197, 199, 201-202 

Momordica 277-278 

Monachanthus 143 

Monanthemum 265 

Monnieria 507, 509 

Monnina 469-470 

Monocardia 524 

Monochaetum 404, 407, 410, 413-414 

Monogramma 37 

Monophyllanthe 115 

Monopteryx 352, 361 

Monotagma 115-116 

Monotrema 173, 175 

Monstera 61, 65-66, 396 

Montira 380 

Montrichardia 61, 64-65 

Moquilea 246 

Mora 174-175, 338, 344-345 

Moraea 111 

Morella 428 

Morinda 484, 493, 495, 497-498 

Moringa 428 

Morisonia 234-235 

Mormodes 127 

Moronobea 249, 252 

Morus 426 

Mosannona 192, 194 

Moschoxylum 419 

Mostuea 379-380 

Mourera 467 

Mouriri 404, 409, 413, 414, 417, 439- 

441 

Mouroucoa 275 

Moutabea 469-470

576 

Moutouchi 361 

Mucoa 197, 199 

Mucuna 352, 359, 361, 363 

Muellera 352, 357,361, 363 

Muntingia 428 

Murdannia 85 

Murraya 507, 509 

Murucoa 275 

Musa 81, 110, 117 

Mussaenda 498 

Mussatia 216, 218-219 

Myanthus 143 

Mycerinus 286 

Myllanthus 509 

Myoxanthus 127, 137, 143, 145 

Myrcia 437-441 

Myrcianthes 438, 441 

Myrciaria 438-442 

Myriaspora 404, 413 

Myrica 428 

Myriocladus 153-154, 164 

Myriophyllum 312 

Myristica 429-430 

Myrmidone 413 

Myrocarpus 352 

Myrodendrum 317 

Myrosma 115-116 

Myrospermum 352 

Myroxylon 303, 352, 364 

Myrsine 431-433 

Myrteola 438, 441 

Myrtiluma 519 

Myrtus 441 

Nacibea 498 

Naiadothrix 524 

Najas 117 

Naletonia 498 

Nama 318 

Nanodes 143 

Nanopteris 37 

Napeanthus 310 

Napimoga 303 

Nasturtium 227 

Nauclea 498 

Naucleopsis 424-427 

Nautilocalyx 310-312 

Navia 75-76, 80-81, 113 

Neblinaea 259 

Neblinantha 305 

Neblinanthera 404 

Neblinaria 224 

Neblinathamnus 484, 498 

Nectandra 324, 327-329 

Neea 443 

Neeragrostis 164 

Nelsonia 184, 186 

Nelumbo 442 

Nemaluma 520 

Nematanthus 312 

Neobartlettia 143 

Neobertiera 484 

Neocalyptrocalyx 235 

Neocouma 197, 199, 202-203 

Neodillenia 281 


Neogleasonia 224 

Neolacis 468 

Neolehmannia 143 

Neomarica 111 

Neopometia 520 

Neoptychocarpus 302, 304 

Neoraputia 507, 509 

Neoregelia 76, 81 

Neosprucea 302 

Neotatea 224, 249 

Neottia 143 

Neoxythece 520 

Nepeta 322 

Nephelea 28 

Nephelium 512 

Nephradenia 210-211 

Nephrodium 33 

Nephrolepis 32, 44-45, 49 

Nepsera 404, 410, 414-416 

Neptunia 369, 373-374 

Nerium 197 

Neurocarpum 361 

Neurodium 46-47 

Neurogramme 51 

Neurolaena 259, 261-262, 265 

Neurolepis 154, 164 

Neuromanes 40 

Neurophyllum 40 

Neurotheca 305, 308 

Newtonia 374 

Nhandiroba 278 

Nicolsonia 361 

Nicotiana 527 

Nidema 127, 137-139, 143 

Nidularium 81 

Nietneria 112-113 

Niopa 374 

Niphidium 46-47 

Nissolia 361 

Noisettia 547-548 

Nonatelia 498 

Nopalea 232 

Norantea 396 

Nortenia 524 

Notanthera 382 

Noterophila 413 

Notholaena 23-24 

Notopleura 382, 484-485, 494, 499- 

502, 505 

Notopora 286 

Notylia 127-128, 137, 143 

Noyera 427 

Nymphaea 444-445 

Nymphoides 422 

Nypa 69 

Ochroma 220, 222 

Ochthephilus 404 

Ochthocosmus 319-320 

Ocimum 321-322 

Ocotea 324-329 

Octomeria 128, 135, 143-144 

Octopleura 308, 413 

Odonellia 272, 274-275 

Odontadenia 197, 202 

Odontandra 419 

Odontocandra 419 

Odontocarya 421 

Odontoglossum 128, 143 

Odontoloma 31 

Odontomanes 40 

Odontonema 184-186 

Odontosoria 30 

Oeceoclades 128, 134, 140 

Oedematopus 252 

Oenocarpus 69, 71-72 

Oenone 468 

Oenothera 452 

Ogcodeia 427 

Ogiera 265 

Olax 451 

Oldenlandia 485, 497-498 

Oldenlandiopsis 485 

Oleandra 32, 44-45 

Olfersia 31, 33 

Oliganthes 265 

Olisbea 413 

Oliveriana 128 

Olmedia 427 

Olmedioperebea 427 

Olmediophaena 427 

Olyra 154, 164 

Omphalea 293, 298 

Omphalobium 270 

Oncidium 128, 139, 143-144 

Oncoba 303 

Oncostylis 99 

Ondidium 128 

Onoctonia 413 

Onychacanthus 186 

Onychium 144 

Oocarpon 452 

Ooclinium 266 

Operculina 272-275 

Ophidion 128, 136 

Ophiocaryon 510 

Ophioglossum 45 

Ophrys 144 

Opisthocentra 404 

Oplismenus 154, 164-167 

Opuntia 232 

Orbignya 72 

Orchis 144 

Orchyllium 377 

Orectanthe 179, 181 

Oreinotinus 235 

Orelia 202 

Orellana 220 

Oreobolus 92, 99 

Oreodaphne 329 

Oreopanax 205-207 

Oribasia 498 

Oritrophium 259 

Orleanesia 128, 141 

Ormoloma 30-31 

Ormopteris 51 

Ormosia 352, 357-358, 360-362 

Ormosiopsis 361 

Ornithidium 144

Ornithocephalus 128-129, 144 

Orthaea 286-287 

Orthiopteris 31 

Orthoclada 154, 158, 164, 167 

Orthomene 421 

Orthopappus 259, 262-263 

Oryctanthus 381-383 

Oryctina 381, 383 

Oryza 154, 164 

Osbeckia 413-414 

Oserya 467-468 

Osmunda 45 

Ossaea 404, 411, 413-415 

Osteophloeum 429-430 

Otachyrium 154 

Otoglossum 129, 143-144 

Otostylis 129, 134, 141, 144, 148-149 

Ottonia 462 

Ouratea 445-449 

Ourouparia 498 

Outea 345 

Oxalis 453 

Oxandra 192-194 

Oxycaryum 92, 101 

Oxymeris 414 

Oxypetalum 210, 212 

Oxythece 520 

Oyedaea 259, 264, 266, 268-269 

Pachira 220-223 

Pachygone 421 

Pachylobus 229 

Pachyloma 404, 414, 417 

Pachyptera 218 

Pachyrhizus 352, 359 

Pachystachys 184, 186 

Pacouria 198, 201 203 

Pacourina 259 

Paepalanthus 105-109 

Paesia 23, 30-31 

Pagaea 308 

Pagamea 485, 498-499 

Pagameopsis 485, 499, 506 

Pakaraimaea 423 

Palala 430 

Palhinhaea 19 

Palicourea 485, 498, 502, 506 

Palma 72 

Palmorchis 129, 137, 143 

Paloue 2, 338 

Paloveopsis 338 

Paltonium 47 

Pamea 255 

Panax 207 

Pancratium 113 

Panicum 154, 164, 167, 172 

Panopsis 474 

Panzera 345 

Papaya 236 

Paphinia 129, 141-142 

Paphiopedilum 144 

Pappophorum 155 

Parabignonia 216 

Paradrymonia 310-311 

Paragonia 215, 218-219 


Parahancornia 198, 200-203 

Paralabatia 520 

Paralea 283 

Paramachaerium 353, 361 

Paramansoa 218 

Paramicropholis 520 

Paraprotium 229 

Parascheelea 72 

Paratheria 155, 166 

Parathesis 433 

Pariana 155, 167-168 

Parietaria 54 

Parinari 244, 246-247 

Parivoa 345 

Parkeria 45 

Parkia 369, 372, 374, 376 

Parkinsonia 338 

Parodiolyra 155, 164, 166, 170 

Parthenium 259 

Paspalidium 155, 164-165 

Paspalum 155156- 167-169, 171-172 

Passaveria 520 

Passiflora 454-456 

Passoura 548 

Passovia 382 

Passowia 382-383 

Patabea 486, 497, 499 

Patima 486, 499, 503 

Patrisia 303 

Pauletia 345 

Paullinia 512-514 

Pausandra 293, 298 

Pavonia 393-395 

Paypayrola 547-548 

Peckia 433 

Pecluma 46-49 

Pectis 259, 266 

Pedicularis 524 

Pedilanthus 293, 297 

Pehria 385 

Peiranisia 345 

Pekea 236 

Pelexia 129, 144, 147-148 

Pellaea 24 

Peltaea 393, 395 

Peltapteris 43 

Peltogyne 338-339, 344-345 

Peltophorum 339, 341, 345, 372 

Peltophyllum 177 

Pennisetum 156, 160, 166, 169 

Pentacalia 259, 266 

Pentaclethra 369-370, 374 

Pentadenia 312 

Pentamerista 536 

Pentascyphus 513 

Pentaspatella 448 

Peperomia 458-459, 461-465 

Pepinia 77, 81 

Pera 293-294, 298, 300 

Perama 486, 493, 498-499 

Perebea 424, 427 

Pereskia 232-233 

Perianthopodus 278 

Peridiscus 456 

Peridium 298 

Perissocarpa 447 

Peristeria 129, 141, 144 

Peritassa 313-316 

Pernettya 286 

Persea 325-327, 329 

Persicaria 472 

Peschiera 198-199, 202 

Pessopteris 47 

Petagomoa 499 

Petaloma 414 

Petastoma 218-219 

Petiveria 457 

Petrea 544, 546 

Petrocarya 247 

Pfaffia 187-188 

Phaeosphaerion 85 

Phainantha 404, 407 

Pharbitis 275 

Pharmacosycea 427 

Pharus 156, 169 

Phaseolus 353, 362 

Phegopteris 53 

Phellocarpus 362 

Phelpsiella 173 

Phenakospermum 176 

Phenax 542-543 

Philacra 447-447 

Philodendron 61-66 

Philodice 105 

Philonotion 66 

Philotria 111 

Philoxerus 188 

Phlebodium 46-47, 49 

Phlegmariurus 19 

Phlomis 322 

Phoebe 329 

Phoenix 69 

Phoradendron 383, 549-552 

Phoxanthus 510 

Phragmipedium 129, 136, 144, 147 

Phragmites 156, 159, 169 

Phryganocydia 216, 219 

Phrygilanthus 383 

Phrynium 116 

Phthirusa 383-384 

Phyla 544, 546 

Phyllanthus 294, 296-299 

Phyllocactus 233 

Phyllocarpus 345 

Phyllodes 116 

Phyllopus 414 

Phymatidium 144 

Phymatodes 47 

Physalis 527-529 

Physinga 144 

Physosiphon 144 

Physostemon 235 

Physurus 144 

Phytarrhiza 81 

Phytolacca 457 

Picramnia 457 

Picrasma 525 

Picria 524

578 

Picrolemma 524-525 

Pilea 542-543 

Pilocarpus 507-509 

Pilocereus 233 

Pilosocereus 232-233 

Pilumna 144 

Pinelia 144 

Pinelianthe 129, 144 

Pinus 56 

Pinzona 281 

Pionandra 529 

Piparea 304 

Piper 459-466 

Piptadenia 369-370, 374-375 

Piptatherum 169 

Piptocarpha 259, 262,264-266, 268 

Piptocoma 259, 265-266 

Piranhea 294 

Piratinera 427 

Piresia 156, 161, 164, 170 

Pirigara 333 

Piripea 524 

Piriqueta 540-541 

Piscidia 353, 360, 362 

Pisonia 443-444 

Pistacia 229 

Pistia 63 

Pitcairnia 77, 80-81 

Pithecellobium 369, 371-376 

Pithecoctenium 216-219 

Pithecolobium 375 

Pitumba 304 

Pityrocarpa 375 

Pityrogramma 23-24, 45, 50-51 

Plagiogyria 45 

Planchonella 520 

Plantago 466 

Plathymenia 369 

Platonia 249 

Platyaechmea 81 

Platycarpum 486, 497, 503 

Platycentrum 414 

Platylepis 99 

Platyluma 520 

Platymiscium 353, 356, 362 

Platypodium 353, 362 

Platysema 362 

Platystele 129, 144-145, 148 

Platythelys 129 

Plectrophora 129, 144 

Pleonotoma 216-218 

Pleopeltis 37, 46-48 

Pleroma 414 

Pleurisanthes 319 

Pleurostachys 92 

Pleurostima 178 

Pleurothallis 129-131, 134-137, 139- 

141, 143-148 

Pleurothyrium, 326, 329 

Plinia 438, 440-441 

Pluchea 259, 262, 264, 266 

Plukenetia 294-297, 299 

Plumbago 466 

Plumeria 198, 202 


Plumeriopsis 202 

Poa 169-170 

Pochota 222 

Podocalyx 295-296, 299 

Podocarpus 56 

Podoluma 516, 520 

Podostemum 467-468 

Poecilandra 447, 449 

Poecilanthe 353, 356-357 

Pogonia 146 

Pogonophora 295, 299 

Pogonorhynchus 414 

Pogonostylis 99 

Pogopetalum 319 

Pogospermum 81 

Poinciana 345 

Poinsettia 299 

Poiretia 353, 360 

Pollalesta 266 

Polybotrya 31, 33, 47 

Polycarpaea 188, 237 

Polycarpon 237 

Polycycnis 131, 146 

Polygala 469-470 

Polygonum 471-472 

Polylychnis 184, 186 

Polymnia 266 

Polyotidium 131, 141 

Polypodium 46-49 

Polypogon 170 

Polypompholyx 377-378 

Polypremnum 379 

Polystachya 131, 138, 144, 146, 148 

Polystichopsis 33 

Polystichum 31-34 

Polytaenium 55 

Polythecandra 252 

Ponera 146 

Pontederia 173 

Ponthieva 131 

Poponax 375 

Poraqueiba 319 

Poraresia 299 

Porocystis 513 

Porophyllum 259, 261, 264, 266 

Portesia 419 

Portlandia 499 

Portulaca 473 

Posadaea 277 

Posoqueria 486, 499, 504 

Possira 362 

Potalia 305 

Potamoganos 216, 218 

Potarophytum 173 

Poteranthera 404, 414 

Pothomorphe 460-461, 465 

Pothos 66 

Pothuava 81 

Poulsenia 424 

Pourouma 238-239 

Pourretia 81 

Pouteria 205, 516-521 

Pouzolzia 542 

Pradosia 234-235, 517-521 

Praxelis 259, 263-264, 266 

Prescottia 131, 136, 146 

Prestoea 69-70, 72 

Prestonia 198, 200-202 

Prevostea 275 

Prieurella 521 

Prionostemma 313-316 

Pristimera 313-315 

Priva 544, 546 

Prockia 302, 304 

Proclesia 287 

Promenaea 146 

Prosthechea 131, 134, 137-139, 146 

Protium 227-230, 508 

Prunus 240, 478 

Psadacocalymma 186 

Psammisia 286-287 

Pseudabutilon 394-395 

Pseudanamomis 438, 441 

Pseudechinolaena 156 

Pseudelephantopus 259, 263, 266 

Pseudephedranthus 192-193 

Pseuderanthemum 184 

Pseuderiopsis 146 

Pseudima 513-514 

Pseudobaccharis 266 

Pseudobombax 221 

Pseudocalymma 216-217, 219 

Pseudocassia 345 

Pseudochimarrhis 499 

Pseudocladia 521 

Pseudoconnarus 269-270 

Pseudogynoxys 259, 266 

Pseudolabatia 521 

Pseudolmedia 424, 427 

Pseudolycopodiella 19 

Pseudomaxillaria 146 

Pseudopaegma 219 

Pseudopiptadenia 369-370, 374 

Pseudosamanea 369-370, 372, 375- 

376 

Pseudosenefeldera 299 

Pseudosorocea 427 

Pseudoxandra 192-194 

Pseudoxythece 521 

Psidium 438, 440-442 

Psiguria 277 

Psila 266 

Psilocarya 99 

Psilochilus 131, 146 

Psilogramme 24 

Psilotum 17, 19 

Psittacanthus 381-383 

Psychopsis 131, 143-144, 146 

Psychotria 486-489, 494-505 

Psygmorchis 146 

Pterandra 389 

Pterichis 131 

Pteridanetium 55 

Pteridium 30-31, 51 

Pteris 31, 50-51 

Pterocarpus 353, 361-362 

Pterocaulon 259, 261-262, 266 

Pterogastra 404, 414

Pterolepis 404, 407, 414-415 

Pteroscleria 99 

Pterozonium 23-24, 51 

Ptilophyllum 40 

Ptychocarpus 304 

Ptychomeria 84 

Ptychopetalum 450 

Pueraria 353 

Pulchranthus 184, 186 

Purdiaea 279 

Puya 77, 81 

Pycreus 99 

Pyrostegia 216 

Pyrrorhiza 109 

Quadrella 235 

Quadricasaea 202 

Qualea 553, 555 

Quamoclit 275 

Quapoya 249, 252 

Quararibea 221-222 

Quassia 522 

Quebitea 465 

Quekettia 131, 146 

Quelchia 259 

Quesnelia 81 

Quiina 475 

Racaria 514 

Racinaea 77, 79, 81-82 

Racoubea 304 

Raddia 156, 170-171, 315 

Raddiella 156, 164-165, 170 

Radia 178 

Radlkoferella 521 

Ragala 521 

Ragatelus 40 

Raimondia 194 

Ramatuela 255 

Ranaria 524 

Randia 489, 493, 497-498, 502 

Rapanea 433 

Rapatea 173-175 

Raphia 72 

Raputia 508-510 

Rauia 508-509 

Rauvolfia 198-199, 202 

Ravenala 176 

Ravenia 509 

Raveniopsis 508-509 

Rechsteineria 312 

Recordoxylon 339, 344-345 

Redutea 395 

Rehia 156, 160 

Reichenbachanthus 131, 140, 146-147 

Reimaria 170 

Reimarochloa 156-158, 170 

Relbunium 502 

Remijia 489-490, 494-495, 497, 502 

Remirea 92, 96, 99 

Renealmia 116, 182 

Renggeria 252 

Rengifa 252 

Restrepia 146 

Restrepiella 146 

Restrepiopsis 131, 146 


Retiniphyllum 490, 495, 498-499, 502- 

504 

Reussia 173 

Rhabdadenia 198, 200 

Rhabdodendron 476 

Rhacodiscus 186 

Rhadinocarpus 362 

Rhamnus 477, 542 

Rhaptostylum 451 

Rheedia 250-253 

Rhexia 414-415 

Rhigospira 198-199, 202 

Rhipidia 202 

Rhipidocladum 157, 159 

Rhipidopteris 43 

Rhipsalis 232-233, 327 

Rhizobolus 236-237 

Rhizophora 477-478 

Rhodocactus 233 

Rhodognaphalopsis 222-223 

Rhodospatha 63, 66 

Rhodostemonodaphne 326-329 

Rhodothyrsus 299 

Rhoeo 85 

Rhoogeton 310, 312 

Rhopalostylis 299 

Rhynchanthera 404-405, 410, 413, 415 

Rhynchelytrum 157, 163, 170, 172 

Rhynchocladium 92, 96 

Rhyncholacis 467-468 

Rhynchopera 146 

Rhynchosia 353, 359-360, 362 

Rhynchospora 92-103 

Rhytachne 157, 161, 163-164, 170 

Rhytiglossa 186 

Riana 548 

Richardella 521 

Richardia 490, 503 

Richardsonia 503 

Richeria 295, 297, 299 

Ricinella 299 

Ricinus 295 

Riedelia 287 

Riedlea 533 

Riencourtia 259-260, 262, 266-267 

Rinorea 547-549 

Rinoreocarpus 548 

Rittera 362 

Rivina 457 

Robergia 270 

Robinia 362-363, 514 

Rodriguezia 131, 135, 146 

Roentgenia 216-217, 219 

Rogersonanthus 305-308 

Rohria 280 

Rolandra 260, 262, 266 

Rollinia 192-194 

Ronabea 490, 495, 498, 500-501, 503 

Rondeletia 490, 503 

Rondonanthus 105, 107-109 

Ropourea 283 

Roraimanthus 449 

Rorippa 227 

Rosenbergiodendron 490, 498, 502 

Roshevitzia 170 

Rotala 385 

Rottboellia 157, 170 

Roucheria 316 

Rouhamon 380 

Roulinia 212 

Rouliniella 212 

Roupala 474 

Rourea 269-270 

Roystonea 69, 72 

Rubia 503 

Rubus 478 

Rudgea 490-491, 495, 497-498, 500- 

501, 503-504 

Rudolfiella 131, 135, 141, 147 

Ruellia 184-186 

Ruizterania 554-555 

Rumohra 29, 32-33, 47, 54 

Ruppia 175 

Ruprechtia 472-473 

Ruptiliocarpon 453 

Russelia 523 

Rutaneblina 508 

Ryania 302-304 

Rytidostylis 277-278 

Sabal 69, 72 

Sabicea 491, 503 

Saccharum 157, 162, 170 

Saccifolium 305, 309 

Sacciolepis 157, 162, 164-167, 170 

Saccoloma 29-31 

Sacoglottis 317-318 

Sacoila 131, 141, 146-148 

Sagittaria 59 

Sagonea 318 

Sagotia 295, 299 

Sagraea 415 

Salacia 313-316 

Saldanha 503 

Saldanhaea 219 

Salmasia 247 

Salmea 260 

Salpichlaena 26 

Salpinctes 198, 202 

Salpinga 405, 410, 415 

Salvadora 433 

Salvertia 554-555 

Salvinia 51 

Samanea 369 

Samara 433 

Sambucus 235 

Samyda 304 

Sanchezia 184 

Sandemania 405, 408, 415 

Sandwithia 295 

Sandwithiodoxa 521 

Santalodes 270 

Saouari 237 

Sapindus 513-514 

Sapium 295, 297, 299-300 

Saracha 529 

Saranthe 116 

Sarcaulus 57, 518 

Sarcoglottis 131, 143-144, 146-148

580 

Sarcopera 396 

Sarcostemma 210-212 

Sarmentaria 415 

Satyria 286-288 

Satyrium 146 

Sauvagesia 447-449 

Savia 295 

Saxofridericia 174-175 

Scaphosepalum 131, 142, 146 

Scaphyglottis 131-132, 136, 140-141, 

146-148 

Scelochilus 132 

Scheelea 69, 72 

Schefflera 206-207 

Schiekia 109 

Schilleria 465 

Schinus 510 

Schismatoglottis 63, 66 

Schistostemon 317-318 

Schizachyrium 157-159, 170 

Schizaea 50, 52 

Schizolobium 339, 341-342, 345 

Schizoloma 31 

Schizopsis 219 

Schlechterella 147 

Schlegelia 216, 219, 311, 545 

Schleidenia 226 

Schlimia 132 

Schlumbergeria 81 

Schmidelia 514 

Schnella 345 

Schoenobiblus 537 

Schoenocephalium 174-175 

Schoenoplectus 94-95, 101-102 

Schoenus 101 

Schoepfia 450 

Schomburgkia 132 

Schradera 491, 503 

Schrankia 376 

Schuebleria 308 

Schultesia 305-306, 308 

Schwenckia 527, 529 

Schwenkfeldia 503 

Sciadocephala 260-261 

Sciadodendron 207 

Sciadophyllum 207 

Sciadotenia 421 

Sciaphila 177 

Scirpidium 101 

Scirpus 101-102 

Sciuris 510 

Scleria 94-95, 98, 101-102 

Sclerolobium 345 

Scleronema 221-223 

Scoparia 523 

Scrophularia 524 

Scutellaria 321 

Scuticaria 132, 142 

Scyphonichium 513 

Sebastiania 295, 297, 300 

Secondatia 198 

Secretania 451 

Securidaca 469-471 

Securinega 300 


Seguieria 457 

Selaginella 18-21 

Selenicereus 232-233 

Selenipedium 132, 136, 147 

Selenodesmium 40 

Selysia 277 

Semaphyllanthe 491, 494, 503 

Senapea 57 

Senecio 266 

Senefeldera 295, 299-300 

Senefelderopsis 295, 299-300 

Senegalia 376 

Senna 339-346 

Serapias 147 

Serianthes 376 

Sericographis 186 

Serjania 513-514 

Serrastylis 147 

Serturnera 188 

Seruneum 266 

Sesamum 456, 523 

Sesbania 353, 356-357, 363 

Sesuvium 187, 473 

Setaria 157, 160-161, 164-167, 170 

Setiscapella 378 

Sextonia 326, 328-329 

Siapaea 260 

Sickingia 503 

Sicklera 529 

Sicyomorpha 315 

Sida 394-395 

Sidastrum 395 

Sideroxylon 521 

Sievekingia 132, 147 

Sigesbeckia 260 

Sigmatostalix 132 

Silphium 266 

Simaba 525 

Simarouba 252 

Simira 491, 497, 503-504 

Sinapis 227 

Sinapistrum 235 

Singana 353 

Sinningia 310-312 

Siolmatra 277 

Sipanea 491, 493, 503-505 

Sipaneopsis 491-492, 503-504 

Sipapoa 392 

Sipapoantha 305 

Siparuna 525-526 

Siphanthera 405, 409-410, 414-415, 

417 

Siphantheropsis 415 

Siphocampylus 233 

Siphoneugena 438, 440-442 

Siphonia 300 

Sisyrinchium 111 

Sitella 533 

Skoliopterys 392 

Sloanea 284-285, 476 

Smilax 175-176 

Sobralia 132, 136, 147 

Socratea 69, 71-72 

Soemmeringia 353 

Solandra 527 

Solanum 527-530 

Solena 504 

Solenidium 132, 143 

Somphoxylon 421 

Sonchus 266-267 

Sophia 223 

Sophoclesia 287-288 

Soramia 281 

Sorghastrum 157-158, 170 

Sorghum 157-158, 162, 170 

Soridium 177 

Sorocea 425, 427 

Souroubea 396 

Souza 111 

Spachea 389, 391 

Sparattanthelium 312-313 

Sparattosperma 216, 218 

Sparganophorus 267 

Spartina 157, 170 

Spathanthus 174-175 

Spathelia 508-510 

Spathicarpa 66 

Spathiger 147 

Spathiphyllum 63, 66 

Spathodea 219 

Spathularia 548 

Specklinia 147 

Speirostyla 539 

Spennera 415-416 

Spermacoce 492-403, 504, 506 

Spermodon 102 

Sphaeradenia 87-88 

Sphaerocionium 40 

Sphaeropteris 28 

Sphaerotheca 424 

Sphagneticola 260-262, 266-268 

Spheneria 157, 169-170 

Sphenoclea 531 

Sphinctanthus 492, 495, 497, 504 

Sphinctolobium 363 

Sphyrospermum 286-288 

Spicanta 26 

Spigelia 379-380 

Spilanthes 260, 262, 267 

Spiracantha 267 

Spiranthera 508 

Spiranthes 132, 144, 147 

Spirodela 112 

Spirotropis 353, 358, 363 

Spixia 300 

Spondias 189 

Spongiosperma 198-199, 201-202 

Sponia 542 

Sporobolus 157-158, 167, 170-172 

Sprucea 504 

Stachyarrhena 492, 503-504 

Stachyphyllum 285 

Stachytarpheta 544-546 

Stahelia 308 

Stanhopea 132, 136, 147 

Stannia 504 

Staphidiastrum 416 

Staphidium 416

Staurogyne 184-186 

Steffensia 465-466 

Stegolepis 174-175 

Steinchisma 157, 165-167 

Steirachne 157, 161, 171 

Steiropteris 54 

Stelestylis 87 

Stelis 132-133, 135, 139, 147-148 

Stellilabium 133 

Stemmadenia 198, 202 

Stemmatosiphon 535 

Stemodia 523-524 

Stemonacanthus 186 

Stenia 133, 148 

Stenocalyx 392, 442 

Stenocephalum 260, 268 

Stenochlaena 26 

Stenocoryne 148 

Stenolobium 363 

Stenoloma 31 

Stenomeria 210 

Stenopadus 260, 267 

Stenophyllus 102 

Stenoptera 148 

Stenorrhynchos 148 

Stenosolen 198, 202 

Stenospermation 63, 65-66 

Stenotaphrum 157, 171 

Stephania 421 

Stephanotella 212 

Sterculia 532-533 

Sterigmapetalum 478 

Steriphoma 234, 421 

Steyerbromelia 77, 80-81 

Steyermarkochloa 157, 167, 171 

Sticherus 34-35 

Stictophyllorchis 133, 141, 146, 148 

Stictophyllum 148 

Stifftia 260, 267 

Stigmaphyllon 389-392 

Stigmatopteris 31-34, 48 

Stilaginella 300 

Stilpnopappus 267 

Stipa 171 

Stizolobium 363 

Stizophyllum 216, 218 

Stomatochaeta 260-262, 267 

Stomoisia 378 

Stratiotes 114 

Strempelia 504 

Strephium 171 

Streptachne 171 

Streptocalyx 73, 81 

Streptochaeta 157 

Streptogyna 157, 171 

Streptostachys 157, 164, 171 

Strigilia 534 

Stromanthe 115-116 

Strophocactus 232-233 

Struchium 260, 263, 267 

Struthanthus 381-384 

Struthiopteris 26 

Strychnodaphne 329 

Strychnos 379-380 


Stryphnodendron 369-371, 373-374. 

376 

Sturmia 148 

Stylogyne 431-434, 518 

Stylosanthes 353, 360, 363-364 

Styrax 533-534 

Subpilocereus 233 

Surenus 419 

Suriana 534 

Swartzia 346, 353-355, 363-364 

Sweetia 363 

Swietenia 418 

Syagrus 69-70 

Symbolanthus 305, 307-308 

Symmeria 472 

Symphonia 250, 252 

Symphyllanthus 280 

Symphyotrichum 260-261 

Symplocos 534-535 

Synedrella 260, 267 

Syngonanthus 105-109 

Syngonium 63, 66 

Syngramma 24 

Syngrammatopsis 24 

Synisoon 504 

Syntherisma 171 

Systemonodaphne 329 

Syzygiopsis 521 

Syzygium 438-441 

Tabebuia 216-219 

Taberna 202 

Tabernaemontana 198-203 

Tacca 177 

Tachia 305, 345 

Tachibota 247 

Tachigali 340, 345-346 

Tacsonia 456 

Taenitis 49 

Tagetes 260 

Taguaria 384 

Taligalea 546 

Talinum 473 

Talipariti 394-395 

Talisia 513-515 

Tamarindus 340 

Tamonea 416, 545-546 

Tampoa 57 

Tanaecium 217, 219 

Tanibouca 255 

Taonabo 536 

Tapeinostemon 305-306, 308, 502 

Tapirira 189-190 

Tapirocarpus 515 

Tapogomea 504 

Tapura 279-280 

Taralea 355, 357-358, 363 

Tarenaya 235 

Tariri 457 

Tassadia 210-212 

Tateanthus 405 

Tecoma 217, 219 

Tectaria 32-33, 49, 52-53 

Telanthera 188 

Teliostachya 184-186 

Telitoxicum 421 

Telminostelma 212 

Tephrosia 355, 358-359, 363-364 

Tephrothamnus 267 

Tepuia 286, 288 

Tepuianthus 535 

Teramnus 355, 359 

Terminalia 254-255 

Ternatea 364 

Ternstroemia 535-536 

Terpsichore 35-38, 46-47 

Tessaria 260, 262, 267 

Tetracera 281-282 

Tetracoryne 304 

Tetragamestus 148 

Tetragastris 148 

Tetralocularia 273 

Tetrameranthus 192 

Tetrameris 416 

Tetramerium 504 

Tetrapodenia 392 

Tetrapollinia 306-308 

Tetrapterys 389-392 

Tetrastichella 219 

Tetrazygia 416 

Tetrorchidium 295 

Teuscheria 133, 135, 148 

Thalia 115-116 

Thaumatophyllum 66 

Thecophyllum 81 

Thelechitonia 267 

Thelypteris 23, 32-33, 37, 47-49, 51, 

53-55 

Thenardia 203 

Theobroma 532-533 

Thermophila 315-316 

Thesium 511 

Thespesia 394 

Thevetia 198-200 

Thibaudia 286-288 

Thieleodoxa 505 

Thiloa 254-255 

Thinouia 513 

Thismia 83 

Thoa 57 

Thoracocarpus 65, 87 

Thrasya 157-158, 166, 168, 171 

Thunbergia 185-186 

Thurnia 177 

Thyella 275 

Thymocarpus 116 

Thyrsacanthus 186 

Thyrsanthus 203 

Thyrsodium 189 

Thysanostemon 250 

Tibouchina 405, 407, 409-411, 414, 

416 

Ticorea 508 

Tigarea 282 

Tilesia 267 

Tillandsia 77-79, 81-82, 182 

Tinantia 85-86 

Tingulonga 230 

Tinus 433-434

582 

Tobagoa 492, 496 

Tococa 405-406, 409-413, 416-417 

Tocoyena 492, 505 

Tofieldia 113 

Toluifera 364 

Tonina 106 

Tonsella 316 

Tontanea 505 

Tontelea 314-316 

Topobea 406, 417 

Torenia 523-524 

Tornelia 66 

Torrubia 444 

Torulinium 102 

Touantea 364 

Touchiroa 346 

Toulicia 513, 515 

Tounatea 364 

Tournefortia 225-226 

Touroulia 475 

Tovomita 250-253 

Tovomitidium 250 

Tovomitopsis 253 

Trachypogon 158-159, 162, 171-172 

Tradescantia 85-86 

Traganthus 300 

Tragia 295, 300 

Tragiopsis 300 

Trattinnickia 228-231 

Trema 542 

Trianosperm 278 

Trianosperma 278 

Trianthema 422 

Triaristella 148 

Triaristellina 148 

Tribulus 556 

Tricentrum 417 

Tricera 231 

Trichachne 172 

Trichanthera 185-186 

Trichelostylis 102 

Trichilia 418-419, 515 

Trichipteris 28 

Trichocentrum 133-134, 136, 138, 

141, 143-144, 148 

Trichochaeta 102 

Trichogonia 260, 264 

Tricholaena 172 

Trichomanes 38-41 

Trichopilia 133, 144, 148 

Trichosalpinx 148 

Trichosanthes 133, 141, 144-148, 278 

Trichospira 260-261, 266-267 

Trichostigma 457 

Trichovaselia 449 

Tricuspis 172 

Tridax 260 

Tridens 158, 172 

Trifolium 364 

Trigonia 539-540 

Trigonidium 133, 148 

Trigynaea 192, 194 

Trilepis 95 

Trimezia 111 


Trinchinettia 267 

Triodia 172 

Triopterys 392 

Triphasia 508 

Triphora 133, 146 

Triplaris 472-473 

Triplophyllum 32-33, 53 

Tripodanthus 382-384 

Tripogandra 85-86 

Tripogon 158, 160 

Tripsacum 158, 172 

Trisecus 477 

Trisetella 133, 142, 148 

Trismeria 24 

Tristicha 468 

Trithrinax 72 

Triumfetta 538-539 

Triuris 177 

Trixis 267 

Trizeuxis 133, 148 

Tropaeolum 540 

Troschelia 109 

Truncaria 417 

Trymatococcus 425-427 

Tryssophyton 406 

Tschudya 417 

Tuberculocarpus 260-261, 268 

Tulasnea 417 

Turnera 533, 540-541 

Tussacia 82, 312 

Tussilago 267 

Tyleria 447-448 

Tyleropappus 260 

Tylopsacas 310-312 

Tylosperma 312 

Tynanthus 217, 219 

Typha 178 

Typhonium 63, 66 

Udora 111 

Ugni 438, 441-442 

Uladendron 532 

Uleiorchis 133, 148 

Ulleria 186 

Uncaria 492, 498 

Uncinia 95 

Unona 194 

Unonopsis 192, 194 

Unxia 260, 264-267 

Uragoga 505 

Uralepis 172 

Urania 176 

Uranthera 417 

Urena 394-395 

Urera 542-543 

Urochloa 158, 167, 172 

Urodesmium 417 

Urospatha 63-64, 66 

Urospathella 63-64, 66 

Urostachys 19 

Urostigma 427-428 

Urtica 543 

Urvillea 513, 515 

Utricularia 376-379 

Uva 194 

Uvaria 194 

Vaccinium 286-288 

Vachellia 376 

Valantia 505 

Valdesia 417 

Valeriana 543 

Valota 172 

Vandellia 524 

Vandenboschia 41 

Vanilla 133-134, 148 

Vantanea 317-318 

Vargasiella 134 

Varronia 226 

Vasconcellosia 236 

Vaselia 449 

Vasivaea 538 

Vatairea 355, 361, 364 

Vataireopsis 355 

Veconcibea 300 

Vellosiella 523-524 

Vellozia 178 

Verbena 546 

Verbesina 260, 267-268 

Vernonanthura 260-261, 268 

Vernonia 268 

Verticillaria 253 

Vesiculina 379 

Vetiveria 158-159 

Veyretia 148 

Viburnum 235-236 

Vicia 364 

Victoria 444-445 

Vigna 355, 356, 359, 362, 364 

Vilfa 172 

Villarsia 422 

Villouratea 449 

Vinca 203 

Vincetoxicum 212 

Viola 548-549 

Virecta 505 

Virola 429-430 

Viscum 551-552 

Vismia 250-253 

Vitellaria 521 

Vitex 545-547 

Vitis 552-553 

Vittaria 51, 55 

Vochya 555 

Vochysia 554-555 

Vogelia 84 

Vogelocassia 346 

Volkameria 547 

Votomita 406, 408, 411, 413, 417 

Vouacapoua 340 

Vouapa 57, 344, 346 

Vouarana 513, 515 

Voyara 235 

Voyria 306-309 

Voyriella 306, 308 

Vriesea 75, 78, 81-83 

Wachendorfia 109 

Wahlenbergia 233 

Wallacea 448 

Walleria 434

Waltheria 532-533, 541 

Warszewiczia 492, 497, 505 

Webera 417 

Websteria 95, 97, 101-102 

Weddellina 469 

Wedelia 260, 266, 268 

Weigeltia 434 

Weinmannia 278-279 

Werauhia 78, 81-83 

Whittonia 456 

Wikstroemia 537 

Wilbrandia 277 

Willoughbya 268-269 

Willughbeja 203 

Wilmattea 233 

Windsorina 175 

Wissadula 394-395 

Witheringia 528-530 

Wittiocactus 233 

Wittmackanthus 492 

Wittmackia 83 

Wolffia 112 

Wolffiella 112 


Wolffiopsis 112 

Wormskioldia 541 

Wulffia 260-262, 267, 269 

Wullschlaegelia 134, 148 

Wurdackanthus 308 

Wurdackia 109 

Xanthosoma 63-64, 66 

Xerorchis 134, 139 

Ximenia 450 

Xiphidium 109, 111 

Xiphochaeta 260,267 

Xiphopteris 37-38 

Xyladenius 304 

Xylobium 134, 142, 148 

Xylophragma 217, 219 

Xylopia 192-194 

Xylosma 302-304 

Xylosterculia 533 

Xyris 179-181 

Youngia 260 

Yutajea 505 

Yuyba 72 

Rhynchospora cordatachenia M.T.Strong (Cypearaceae) 

Zamia 56 

Zantedeschia 66 

Zanthoxylum 508-510 

Zapoteca 369-370, 373 

Zea 158, 162 

Zephyranthes 113 

Zeugites 158 

Zexmenia 269 

Zeyheria 219 

Zingiber 182 

Zinowiewia 239 

Ziziphus 477 

Zollernia 356, 364 

Zornia 356, 360, 364 

Zostera 183 

Zschokkea 203 

Zuelania 302-304 

Zygia 369-376 

Zygopetalum 148-149 

Zygophlebia 36-37, 49 

Zygosepalum 134, 139, 143,148-149 

Zygostates 134

584 

Acanthaceae 183 

Adiantaceae 22 

Agavaceae 58 

Aizoaceae 187 

Alismataceae 59 

Amaranthaceae 187 

Anacardiaceae 189 

Anisophylleaceae 190 

Annonaceae 190 

Apiaceae 195 

Apocynaceae 195 

Aquifoliaceae 203 

Araceae 59 

Araliaceae 205 

Arecaceae 67 

Aristolochiaceae 207 

Asclepiadaceae 208 

Aspleniaceae 24 

Balanophoraceae 212 

Balsaminaceae 213 

Basellaceae 213 

Bataceae 213 

Begoniaceae 213 

Bignoniaceae 214 

Bixaceae 220 

Blechnaceae 25 

Bombacaceae 220 

Bonnetiaceae 223 

Boraginaceae 224 

Brassicaceae 226 


Brunelliaceae 227 

Burmanniaceae 83 

Burseraceae 227 

Buxaceae 231 

Cabombaceae 231 

Cactaceae 231 

Campanulaceae 233 

Canellaceae 234 

Cannaceae 84 

Capparaceae 234 

Caprifoliaceae 235 

Caricaceae 236 

Caryocaraceae 236 

Caryophyllaceae 237 

Casuarinaceae 237 

Cecropiaceae 237 

Celastraceae 239 

Ceratophyllaceae 240 

Chenopodiaceae 240 

Chloranthaceae 240 

Chrysobalanaceae 241 

Clethraceae 247 

Clusiaceae 247 

Combretaceae 254 

Commelinaceae 85 

Compositae 255 

Connaraceae 269 

Convolvulaceae 271 

Costaceae 86 

Crassulaceae 276 

Cucurbitaceae 276 

Cunoniaceae 278 

Cuscutaceae 279 


INDEX TO FAMILIES 

Cyatheaceae 26 

Cycadaceae 56 

Cyclanthaceae 87 

Cyperaceae 88 

Cyrillaceae 279 

Davalliaceae 29 

Dennstaedtiaceae 29 

Dichapetalaceae 279 

Dicksoniaceae 31 

Dilleniaceae 280 

Dioscoreaceae 103 

Droseraceae 282 

Dryopteridaceae 31 

Ebenaceae 282 

Elaeocarpaceae 284 

Eremolepidaceae 285 

Ericaceae 285 

Eriocaulaceae 104 

Erythroxylaceae 288 

Euphorbiaceae 290 

Euphroniaceae 301 

Flacourtiaceae 301 

Gentianaceae 304 

Gesneriaceae 309 

Gleicheniaceae 34 

Gnetaceae 57 

Grammitidaceae 35 

Haemodoraceae 109 

Haloragaceae 312 

Heliconiaceae 110 

Hernandiaceae 312 

Hippocrateaceae 313 

Hugoniaceae 316 

Humiriaceae 316 

Hydrocharitaceae 110 

Hydrophyllaceae 318 

Hymenophyllaceae 38 

Hymenophyllopsidaceae 41 

Icacinaceae 318 

Iridaceae 111 

Isoëtaceae 17 

Ixonanthaceae 319 

Juglandaceae 320 

Juncaceae 111 

Krameriaceae 320 

Lacistemataceae 320 

Lamiaceae 321 

Lauraceae 322 

Lecythidaceae 330 

Leguminosae-Caesalpinioideae 

333 

Leguminosae-Faboideae 346 

Leguminosae-Mimosoideae 365 

Lemnaceae 112 

Lentibulariaceae 376 

Liliaceae 112 

Limnocharitaceae 114 

Lissocarpaceae 379 

Loasaceae 379 

Loganiaceae 379 

Lomariopsidaceae 42 

Loranthaceae 381 

Lycopodiaceae 17 

Lygodiaceae 43 

Lythraceae 384 

Magnoliaceae 385 

Malpighiaceae 386 

Malvaceae 393 

Marantaceae 114 

Marattiaceae 44 

Marcgraviaceae 396 

Marsileaceae 44 

Mayacaceae 117 

Melastomataceae 397 

Meliaceae 417 

Mendonciaceae 419 

Menispermaceae 420 

Menyanthaceae 422 

Metaxyaceae 44 

Molluginaceae 422 

Monimiaceae 422 

Monotaceae 423 

Moraceae 423 

Moringaceae 428 

Muntingiaceae 428 

Musaceae 117 

Myoporaceae 428 

Myricaceae 428 

Myristicaceae 428 

Myrsinaceae 430 

Myrtaceae 434 

Najadaceae 117 

Nelumbonaceae 442 

Nyctaginaceae 442 

Nymphaeaceae 444 

Ochnaceae 445 

Olacaceae 449 

Oleaceae 451 

Oleandraceae 44 

Onagraceae 451 

Ophioglossaceae 45 

Opiliaceae 453 

Orchidaceae 118 

Osmundaceae 45 

Oxalidaceae 453 

Papaveraceae 454 

Parkeriaceae 45 

Passifloraceae 454 

Pedaliaceae 456 

Peridiscaceae 456 

Phytolaccaceae 457 

Picramniaceae 457 

Pinaceae 56 

Piperaceae 458 

Plagiogyriaceae 45 

Plantaginaceae 466 

Plumbaginaceae 466 

Poaceae 149 

Podocarpaceae 56 

Podostemaceae 466 

Polygalaceae 468 

Polygonaceae 471 

Polypodiaceae 45 

Pontederiaceae 173 

Portulacaceae 473 

Primulaceae 473 

Proteaceae 474 

Psilotaceae 17 

Pteridaceae 50 

Quiinaceae 475 

Rafflesiaceae 476 

Ranunculaceae 476 

Rapateaceae 173 

Rhabdodendraceae 476 

Rhamnaceae 477 

Rhizophoraceae 477 

Rosaceae 478 

Rubiaceae 479 

Ruppiaceae 175 

Rutaceae 507 

Sabiaceae 510 

Salviniaceae 51 

Santalaceae 511 

Sapindaceae 511 

Sapotaceae 515 

Sarraceniaceae 521 

Schizaeaceae 52 

Scrophulariaceae 522 

Selaginellaceae 19 

Simaroubaceae 524 

Siparunaceae 525 

Smilacaceae 175 

Solanaceae 526 

Sphenocleaceae 531 

Sterculiaceae 531 

Strelitziaceae 176 

Styracaceae 533 

Surianaceae 534 

Symplocaceae 534 

Taccaceae 177 

Tectariaceae 52 

Tepuianthaceae 535 

Ternstroemiaceae 535 

Tetrameristaceae 536 

Theaceae 536 

Thelypteridaceae 53 

Theophrastaceae 537 

Thurniaceae 177 

Thymelaeaceae 537 

Tiliaceae 538 

Trigoniaceae 539 

Triuridaceae 177 

Tropaeolaceae 540 

Turneraceae 540 

Typhaceae 178 

Ulmaceae 542 

Urticaceae 542 

Valerianaceae 543 

Velloziaceae 178 

Verbenaceae 543 

Violaceae 547 

Viscaceae 549 

Vitaceae 552 

Vittariaceae 55 

Vochysiaceae 553 

Winteraceae 555 

Woodsiaceae 55 

Xyridaceae 178 

Zamiaceae 56 

Zingiberaceae 182 

Zosteraceae 183 

Zygophyllaceae 556

Checklist of the Plants of the Guiana Shield - Department of Botany ...

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