BRYOZOA: CHEILOSTOMATA Interim classification for Treatise ...

BRYOZOA: CHEILOSTOMATA 

Interim classification for Treatise) 

Compiled by 

Dennis P. Gordon 

1

2 

# genera 

/subgenera 

Suborder MALACOSTEGINA Levinsen, 1902, p. 2 [Malacostegina d'Hondt 1985, pp. 9, 11] ?KIM/TTH- 

Rec. 32 

Superfamily Membraniporoidea Busk, 1852, p. 2 [Membraniporacea Vigneaux 1949, pp. 10, 27) ?KIM/ 

TTH-Rec. 32 

Electridacea Pheophanova 1960, p. 155; Membraniporacea Medd 1979, p. 1 

Family Electridae Stach, 1937, p. 376 ?KIM/TTH-Rec. 26 

Electrinae d'Orbigny, 1851, p. 187; Electrinidae d'Orbigny, 1851, pp. 319, 329; Electridae Lagaaij, 1952, p. 20 

Arbocuspis, Arbopercula, Aspidelectra, Basslerinella, Bathypora, Bullaconopeum, Charixa, Chiplonkarina, 

Conopeum, Craspedopora, Craspedoporina, Danocella, Einhornia, Electra, Eokotosokum, Gontarella, 

Harpecia, Herpetopora, Heteroconopeum, Mychoplectra, Osburnea, Pyripora, Pyriporopsis, 

Rhammatopora, Spinicharixa, Villicharixa 

Family Corynostylidae Vigneaux, 1949, p. 12 (Corynostylusidae; nom. corr. Sandberg 1962, p. 61) ?AQT-? 

MES 1 

Corynostylus 

Family Membraniporidae Busk, 1852, p. 2; 1854, 55 ?PRB-Rec. 3 

Biflustridae d'Orbigny, 1852, p. 217; Biflustridae Smitt, 1873, p. 17; Acanthodesiidae Vigneaux, 1949, p. 11 

Biflustra, Jellyella, Membranipora 

Family Wawaliidae Dzik, 1975, p. 405 VLG-HAU 1 

Wawalia 

Incertae sedis 1 

Christinella 

Suborder INOVICELLINA Jullien, 1888, p. 8 [Inovicellatina d'Hondt 1985, pp. 9, 11] ZAN-Rec. 2 

Superfamily Aeteoidea Smitt, 1868, p. 279 [Aeteidacea Pheophanova 1960, p. 155] ZAN-Rec. 2 

Family Aeteidae Smitt, 1868, p. 279 ZAN-Rec. 2 

Aetea, Callaetea 

Suborder SCRUPARIINA Silén, 1941, p. 9 [Scrupariinina d'Hondt 1985, pp. 9, 11] MAA-Rec. 5 

Superfamily Scruparioidea Gray, 1848 p. 132 [Scruparioidea Gordon 1986, p. 13] MAA-Rec. 5 

Family Scrupariidae Gray, 1848 p. 132 (as Scupariadae, lapsus calami, vide Harmer 1923, p. 316) MAA-Rec. 2 

Scrupariadae Busk, 1852, p. 28 

Brettiopsis, Scruparia 

Family Leiosalpingidae d'Hondt & Gordon, 1996, p. 87 Rec. 2 

Astoleiosalpinx, Leiosalpinx 

Family Eucrateidae Johnston, 1847, p. 288 (Eucrateadae) Rec. 1 

Gemellariadae Busk, 1852, p. 383 

Eucratea 

Suborder NEOCHEILOSTOMINA d’Hondt, 1985, pp, 9, 11 (part) ALB-Rec. 1025 

Infraorder FLUSTRINA Smitt, 1868, p. 357 (part) ALB-Rec. 373 


Family Mourellinidae Rec. 1 

Mourellina 

Superfamily Calloporoidea Norman, 1903, p. 587 [nom. transl. Gordon 1989, p. 1324] ALB-Rec. 136 

Antroporacea Vigneaux 1949, p. 37 

Family Calloporidae Norman, 1903, p. 587 ALB-Rec. 81 

Flustrellaradae d'Orbigny, 1852 (nom. nud.), pp. 239, 339; 1853, p. 504; ?Escharinellidae d'Orbigny, 1852, pp. 339, 

425; ?Flustrellidae d'Orbigny, 1852, p. 339; 1853, p. 559; Flustrinidae d'Orbigny, 1852, p. 339; 1853, p. 474; 

Alderinidae Canu & Bassler, 1925, p.14; 1927, p. 3; Allantoporidae Vigneaux, p. 11; Tegellidae Vigneaux, 1949, p. 

12; Ellisinidrinae Vigneaux, 1949, p. 33; Ellisininae Vigneaux, 1949, p. 36; Ogivalininae Vigneaux, 1949, p. 40

Acanthoporella, Acanthoporidra, Adenifera, Alderina, Allantopora, Allantocallopora, Ammatophora, 

Amphiblestrum, Aplousina, Aviculamphiblestrum, Berthelsenia, Biaviculigera, Bidenkapia, Bryocalyx, 

Callopora, Caulorhamphus, Clavodesia, Concertina, Copidozoum, Corbulella, Cranosina, Crassicellepora, 

Crassimarginatella, Crepis, Cymulopora, Dactylostega, Daisyella, Dermatopora, Dionella, Distelopora, 

Ellisina, Flustrellaria, Gilbertopora, Glenelgia, Hapsidopora, Hemiseptella, Kristerina, Kunradocella, 

Lamourouxia, Leptinatella, Marginaria, Marssonopora, Megapora, Membraniporidra, Mystriopora, 

Ogivalina, Onychoblestrum, Ornatella, Pachybaktropora, Parellisina, Periporosella, Pithodella, 

Planicellaria, Pseudoallantopora, Punctocellaria, Pyriflustrina, Pyriporella, Pyriporoides, Ramphonotus, 

Reptoflustrella, Reptoporina, Retevirgula, Semiflustrella, Semiflustrina, Septentriopora, Solenophragma, 

Stamenocella, Tegella, Tremogasterina, Treptopora, Tylopora, Tyloporella, Unidistelopora, 

Valdemunitella, Wilbertopora, Xylochotridens 

Incertae sedis 

2 

Filiflustrina, Fissuricella 


Austroflustra, Klugeflustra, Neoflustra 

Family Antroporidae Vigneaux, 1949, p. 12 CEN/TUR-Rec. 6 

Akatopora, Antropora, Lateroflustrella, Parantropora, Rosseliana, Samudraka 

Family Chaperiidae Jullien, 1888, p. 61 (Chaperidae) MAA-Rec. 13 

Chapperiidae Bassler, 1935, p. 38; Chaperiellidae Harmer, 1957, p. 641; Chaperinidae Balavoine, 1959, p. 265 

Amphiblestrella, Bryochaperia, Catenariopsis, Chaperia, Chaperiopsis, Clipeochaperia, Exallozoon, 

Exostesia, Icelozoon, Larnacicus, Notocoryne, Patsyella, Pyrichaperia 

Family Doryporellidae Grischenko, Taylor & Mawatari, 2004, p. 327 Rec. 2 

Doryporella, Doryporellina 

Family Foveolariidae Gordon & Winston in Winston, 2005, p. 12 RUP-Rec. 3 

Amplexicamera, Foveolaria, Odontionella 

Family Hiantoporidae Gregory, 1893, p. 222 DAN-Rec. 2 

Hiantoporidae MacGillivray, 1895, p. 60 

Hiantopora, Tremopora 

Family Quadricellariidae Gordon, 1984, p. 24 CEN-Rec. 7 

Cellarinidra, Curvacella, Mediosola, Nellia, Nelliella, Quadricellaria, Vincularina 

Family Bryopastoridae d’Hondt & Gordon, 1999, p. 

MAA-Rec. 6 

Bryopastor, Dioptropora, Monticellaria, Omoiosia, Pseudothyracella, Thyracella 

Family Vinculariidae Busk, 1852, p. 2; 1854, p. 95 (Vinculariadae) LUT-Rec. 1 

Vincularia 

Family Farciminariidae Busk, 1852, pp. 2, 32 (Farciminariadae) Rec. 4 

Columnella, Didymozoum, Farciminaria, Farciminellum 

Family Tendridae Vigneaux, 1949, p. 10 Rec. 2 

Heterooecium, Tendra 

Family Heliodomidae Vigneaux, 1949, p. 37 (Heliodominae) DAN-Rec. 2 

Setosellinidae Hayward & Cook, 1979, p. 45 

Heliodoma, Setosellina 

Family Cupuladriidae Lagaaij, 1952, p. 31 ?DAN/?THA-Rec. 5 

Porellidae d'Orbigny, 1852, pp. 339, 472; Discoporellidae Baluk & Radwanski, 1984, p. 24 

Biselenaria, Cupuladria, Discoporella, Reussirella, Vibracellina 

3


Parastichopora, Pseudolunularia, Stichopora 

Superfamily Flustroidea Fleming, 1828, p. 534 [Flustroidea Alvarez, 1991, pp. (10), 49] Rec. 17 

Family Flustridae Fleming, 1828, p. 534 (Flustradae) Rec. 17 

Hincksinae Canu & Bassler, 1925, p. 14; Hincksinidae Canu & Bassler, 1927, p. 3 

Carbasea, Chartella, Flustra, Gregarinidra, Hincksina, Hincksinoflustra, Isosecuriflustra, Kenella, 

Membraniporopsis, Nematoflustra, Retiflustra, Sarsiflustra, Securiflustra, Serratiflustra, Sinoflustra, 

Spiralaria, Terminoflustra 

Superfamily Buguloidea Gray, 1848, p. 110 [Bugulidacea Pheophanova 1960, p. 162] MAA-Rec. 67 

Family Bugulidae Gray, 1848, p. 110 Rec. 31 

Acamarchisidae d'Orbigny, 1852, p. 319; Cellulariadae Busk, 1852, p. 368; Bicellariadae Busk, 1852, p. 373; 

Bicellariellidae Levinsen, 1909, p. 431; Halophilidae Vigneaux, 1949, p. 10; Sadkoidae Kluge, 1962, p. 362 

Beanodendria, Bicellariella, Bicellarina, Brettiella, Bugula, Bugularia, Bugulella, Calyptozoum, 

Camptoplites, Caulibugula, Chalastobugula, Cornucopina, Corynoporella, Cuneiforma, Dendrobeania, 

Dimetopia, Falsibugulella, Farciminellopsis, Halophila, Himantozoum, Himantozoumella, Kinetoskias, 

Klugella, Luguba, Microcorcillum, Nordgaardia, Semidendrobeania, Semikinetoskias, Sessibugula, 

Thaminozoum, Uschakovia 


Brettia 

Family Rhabdozoidae MacGillivray, 1887, p. 196 Rec. 1 

Rhabdozoum 

Family Beaniidae Canu & Bassler, 1927, p. 14 ?LUT/?BRT-Rec. 3 

Amphibiobeania, Beania, Stolonella 

Family Epistomiidae Gregory, 1893, p. 222 ?YPR/AQT/BUR/LAN-Rec. 2 

Notamiidae Hincks, 1879, p. 155 

Epistomia, Synnotum 

Family Euoplozoidae Harmer, 1926, p. 390 Rec. 1 

Euoplozoum 

Family Candidae d'Orbigny, 1851, pp. 22, 25, 51 MAA-Rec. 27 

Cabereadae Busk, 1852, p. 376; Cabereidae Jullien, 1883, p. 528; Scrupocellariidae Levinsen, 1909, p. 130; 

Synaptacellidae Maplestone, 1911, p. 272; Menipeidae Vigneaux, 1949, p. 10; Pseudoporicellariidae d'Hondt, 1987, 

p. 544 

Amastigia, Bactrellaria, Bobinella, Bugulopsis, Caberea, Cabereopsis, Canda, Candomenipea, 

Candoscrupocellaria, Diplobicellariella, Emma, Eoscrupocellaria, Eupaxia, Hoplitella, Maplestonia, 

Menipea, Monartron, Notoplites, Notoplitesigia, Penemia, Pseudoporicellaria, Retiscrupocellaria, 

Scrupocellaria, Scrupocellarinella, Semibugula, Synaptacella, Tricellaria 

Family Jubellidae Reverter-Gil & Fernández-Pulpeiro, 2001 Rec. 1 

Jubella 

Superfamily Microporoidea Gray, 1848, p. 115 [Microporacea Vigneaux 1949, p. 13] CEN-Rec. 110 

Steganoporellacea Vigneaux 1949, p. 12 

Family Microporidae Gray, 1848, p. 115 CEN-Rec. 31 

Eschariporidae d'Orbigny, 1852, pp. 217, 339; 1853, p. 478; ?Escharellidae d'Orbigny, 1852, pp. 339, 459; 

Opesiulidae Jullien, 1888, p. 79; Calpensiidae Canu & Bassler, 1923, p. 67; Monsellinae Vigneaux, 1949, p. 43 

Andreella, Apiophragma, Aviculiera, Calpensia, Cianotremella, Dimorphostylus, Escharipora, Flustrapora, 

Gargantua, Hoplitaechmella, Manzonella, Micropora, Microporina, Mollia, Monsella, Nematoporella, 

Opaeophora, Poropeltarion, Promicroa, Puncturiella, Puncturiellina, Rectimicropora, Reptescharinella, 

Reussinella, Rimosocella, Scriblitopora, Setosinella, Steraechmella, Stichomicropora, Stictostega, 

Verminaria 

Family Monoporellidae Hincks, 1882, p. 123; ??-Rec. 1 

4

Monoporella 

Family Macroporidae Uttley, 1949, p. 175 ?MAA/?THA-Rec. 1 

Macropora 

Family Calescharidae Bock & Cook, 

1998 ??-Rec. 4 Caleschara, Coronellina, 

Hagenowinella, Tretosina 

Family Setosellidae Levinsen, 1909, p. 196 THA-Rec. 1 

Setosella 

Family Lunulitidae Gregory, 1893, 224 (Lunulitinae) 

SAN-Rec.? 7 

Lunulitinae Buge, 1952, p.718; Lunulitidae Lagaaij, 1952, p. 43 

Discoflustrellaria, Heteractis, Luganella, Lunulites, Oligotresium, Pavolunulites, Trochopora 

Family Lunulariidae Levinsen, 1909, p.155 ? 

BUR-Rec 1 

Lunularia 

Family Otionellidae Bock & Cook, 1998, p. ? 

RUP-Rec 5 Helixotionella, Kausiaria, Otionella, Otionellina, Petasosella 

Family Selenariidae Busk, 1852, p. 2 (nom. nud.); 1854, p. 97 (Selenariadae) ?BUR-Rec. 2 

Discovibracella, Selenaria 


Vibracella, Volviflustrellaria 

Family Onychocellidae Jullien, 1882, p. 276 CEN-Rec. 29 

Euritininae Vigneaux, 1949, p. 41 

Aechmella, Cheethamia, Chondriovelum, Collura, Dictuonia, Euritina, Floridina, Floridinella, Holsacella, 

Inversaria, Latereschara, Matyashegyella, Nudicella, Nudonychocella, Ogiva, Onychocella, Onychocellaria, 

Otiochmella, Pseudolunulites, Radulopora, Rectonychocella, Reptolunulites, Rhebasia, Semieschara, 

Smittipora, Solenonychocella, Tornipora, Virgocella, Woodipora 


Acanthodesiomorpha, Ogivalia 

Family Aspidostomatidae Jullien, 1888, p. 77 (Aspidostomidae) CEN-Rec. 6 

Aspidostoma, Crateropora, Entomaria, Lagarozoum, Larvapora, Taphrostoma 

Family Steginoporellidae Hincks, 1884, p. 358 (Steganoporellidae, nom. emend. Bassler 1953, p. 171) LUT- 

Rec. 6 

Labioporellidae Harmer, 1926, p. 280; Siphonoporellinae Vigneaux, 1949, p. 42 

Dimorphomicropora, Gaudryanella, Labioporella, Reniporella, Siphonoporella, Steginoporella 

Family Thalamoporellidae Levinsen, 1902, p. 175 LUT-Rec. 6 

Hesychoxeniinae Gordon & Parker, 1991, p. 1377 

Dibunostoma, Diploporella, Hesychoxenia, Marsupioporella, Thairopora, Thalamoporella 

Family Chlidoniidae Busk, 1884, p. 8 (Chlidoniadae) BRT-Rec. 1 

Cothurnicellidae Bassler, 1935, p. 27 

Chlidonia 

Family Poricellariidae Harmer, 1926, p. 313 MAA-Rec. 1 

Poricellaria 

Family Alysidiidae Levinsen, 1909, p. 201 Rec. 3 

Alysidium, Catenicula, Paracatenicula 

5

Superfamily Cellarioidea Fleming, 1828, p. 539 [Cellariidacea Pheophanova 1960, p. 7] SAN-Rec. 28 

Family Cellariidae Fleming, 1828, p. 539 (Cellariadae) SAN-Rec. 22 

Salicornariadae Busk, 1852, p. 366; Meliceritidae Jullien, 1888, p. 67; Euginomidae Vigneaux, 1949, p. 12; 

Meliceritidae Vigneaux, 1949, p. 12 

Acerinucleus, Atelestozoum, Cellaria, Cryptostomaria, Dimorphocellaria, Dubiocellaria, Erinella, 

Escharicellaria, Euginoma, Formosocellaria, Hemistylus, Henrimilnella, Melicerita, Melychocella, 

Mesostomaria, Paracellaria, Pseudocellaria, Smitticellaria, Steginocellaria, Stomhypselosaria, Swanomia, 

Syringotrema 

Family Membranicellariidae Levinsen, 1902, p. 22 Rec. 2 

Cookinella, Membranicellaria 

Family Coscinopleuridae Canu, 1913, p. 145 SAN-THA 4 

Acoscinopleura, Coscinopleura, Escharifora, Tremocoscinopleura 

INCERTAE SEDIS 12 

Family Fusicellariidae Canu, 1900, p. 458 TUR-MAA 2 

Encicellaria, Fusicellaria 

Family Skyloniidae Sandberg, 1963, p. 16 DAN-?MES/?ZAN 4 

Calvina, Kylonisa, Skylonia, Sylonika 

Family Bicorniferidae Keij, 1977, p. 233 DAN-Rec. 4 

Babickella, Bicornifera, Bifissurinella, Voorthuyseniella 


Hoeverella, Taeniocellaria 

Infraorder ASCOPHORINA Levinsen, 1909, p. 213 (Ascophora) [nom. transl. Cuffey 1973, p. 561] CEN-Rec. 

652 

“Grade” ACANTHOSTEGA Levinsen, 1902, p. 2 (Acanthostega) [see Cheetham 1963, p. 60] nom. transl. CEN- 

Rec. 179 

Cribrimorpha Harmer, 1926, p. 191; Cribriomorpha, nom. transl. Gordon 1989, p. 15 

Superfamily Cribrilinoidea Hincks, 1879, p. 156 [Costulacea Vigneaux, 1949, p. 13; Cribrilinidacea 

Pheophanova 1960, p. 163; Cribrilinoidea Gordon 1984, p. 60] CEN-Rec. 120 

Family Cribrilinidae Hincks, 1879, p. 156 CEN-Rec. 116 

**Steginoporidae d'Orbigny 1852, p. 339; 1853, p. 497; ?Escharellidae d'Orbigny, 1852, pp. 459, 462; Costulidae 

Jullien, 1886, p. 602; Pelmatoporidae Lang, 1916, p. 83; Francoporinae Lang, 1916, p. 84; Opisthornithoporinae 

Lang, 1916, p. 84; Kelestominae Lang, 1916, p. 85; Tricephaloporinae Lang, 1916, p. 85; Pnictoporinae Lang, 1916, 

p. 143; Castoporinae Lang, 1916, p. 93; Diacanthoporinae Lang, 1916, p. 100; Andrioporidae Lang, 1916, p. 381; 

Pliophloeinae Lang, 1916, p. 391; Lagynoporidae Lang, 1916, p. 393; Schistacanthoporinae Lang, 1916, p. 393; 

Leptocheiloporinae Lang, 1916, p. 394; Rhacheoporidae Lang, 1916, p. 397; Disheloporinae Lang, 1916, p. 400 

(Disheloporidae, nom. transl. Lang 1921, p. 215); Otoporidae Lang, 1916, p. 401; Calpidoporidae Lang, 1916, p. 198; 

Ctenoporidae Lang, 1916, p. 403; Myagroporidae Lang, 1916, p. 406; Taractoporidae Lang, 1916, p. 406; 

Thoracoporidae Lang, 1916, p. 407; Puellinidae O’Donoghue & de Watteville, 1939, p. 19. 

Abdomenopora, Acanthobaktron, Aeolopora, Anaptopora, Anaskopora, Andriopora, Angelopora, 

Anornithopora, Anotopora, Argopora, Ascancestor, Auchenopora, Baptopora, Batrachopora, 

Callistopora, Calpidopora, Canupora, Carydiopora, Castanopora, Castanoporina, Cillia, Coelopora, 

Collarina, Confusocella, Corbulipora, Corbuliporina, Corymboporella, Costula, Craticulacella, Cribralaria, 

Cribrendoecium, Cribrilaria, Cribrilina, Ctenopora, Decurtaria, Dendroperistoma, Diacanthopora, 

Diancopora, Diceratopora, Dishelopora, Distansescharella, Disteginopora, Eucheilopora, Figularia, 

Filaguria, Francopora, Geisopora, Gephyrotes, Gordoniella, Glabrilaria, Graptoporella, Haplocephalopora, 

Hayamiellina, Hesperopora, Hexacanthopora, Hippiopora, Holostegopora, Hybopora, Hystricopora, 

Ichnopora, Inversiscaphos, Jolietina, Jullienula, Kankopora, Kelestoma, Keratostoma, Klugerella, 

Lagynopora, Lepralina, Leptocheilopora, Membraniporella, Metracolposa, Monoceratopora, 

Morphasmopora, Multescharipora, Mumiella, Murinopsia, Myagropora, Nannopora, Oligotopora, 

Opisthornithopora, Otopora, Pachydera, Pancheilopora, Parafigularia, Pelmatopora, Phractoporella, 

Phrynopora, Pleuroschizella, Pliophloea, Pnictopora, Polycephalopora, Polyceratopora, Prodromopora, 

Prosotopora, Puellina, Reginella, Reginelloides, Reptescharella, Reptescharipora, Reptoporella, 

6

Rhabdopora, Rhacheopora, Rhiniopora, Sandalopora, Schistacanthopora, Semiescharipora, Steginopora, 

Stichocados, Taractopora, Thoracopora, Tricephalopora, Turnerellina, Ubaghsia, Vavropora, Vicariopora 

Family Lekythoglenidae Marsson, 1887, p. 90 (Lekythoglenidea) CMP 1 

Lekythoglena 

Family Euthyroididae Levinsen, 1909, p. 264 (Euthyroidae) Rec. 1 

Euthyroides 

Family Polliciporidae Moyano, 2000, p. 301 

Rec. 1 

Pollicipora 

Family Belluloporidae nov. Rec. 1 

Bellulopora 

Superfamily Bifaxarioidea Busk, 1884, p. 79 [nom. transl. d'Hondt 1985, pp. 9, 11] TUR-Rec. 10 

Family Bifaxariidae Busk, 1884, p. 79 (Bifaxariadae) YPR-Rec. 6 

Aberrodomus, Bifaxaria, Diplonotos, Domosclerus, Raxifabia, Smithsonius 

Family Platyglenidae Marsson, 1887, p. 89 (Platyglenidea) TUR-DAN 3 

Platyglena, Pnictoporopsis, Systenostoma 

Family Mixtopeltidae Gordon, 1994, p. 66 Rec. 1 

Mixtopelta 

Superfamily Scorioporoidea Gordon, 2002, p. 123 CMP-DAN 9 

Family Scorioporidae Gordon, 2002, p. 123 

CMP-DAN 6 

Bathystomella, Beisselinopsis, Columnotheca, Diplobeisselina, Rotiporina, Scoriopora 

Family Nephroporidae Marsson, 1887, p. 90 (Nephroporidea) 

MAA 1 

Nephropora 


Ochetosellina, Taenioporella 

Superfamily Taenioporinoidea nov. MAA 1 

Family Taenioporinidae nov. MAA 1 

Taenioporina 

Superfamily Catenicelloidea Busk, 1852, p. 352 [nom. transl. Gordon 1984, p. 64] MAA-Rec. 39 

Family Catenicellidae Busk, 1852, p. 352 MAA-Rec. 29 

Catenariidae Levinsen, 1909, p. 213; Vittaticellinae Stach, 1933, p. 87; Scuticellinae Stach, 1934, p. 21; 

Cornuticellinae Stach, 1935, p. 393; Ditaxiporinae Stach, 1935, p. 394; Claviporellinae Correa, 1947, p. 5; 

Vitatticellidae Harmer, 1957, p. 765; Ditaxiporinidae Cheetham, 1963, p. 489; Vasignyellinae Gordon & Braga, 1994, 

p. 76 

Ahcheethamia, Bryosartor, Caberoides, Caloporella, Calpidium, Catenicella, Catenicellopsis, Claviporella, 

Cornuticella, Cornuticellina, Costaticella, Costatimorpha, Cribricellina, Digenopora, Ditaxipora, 

Ditaxiporina, Geminella, Orthoscuticella, Paracribricellina, Plagiopora, Pterocella, Scalicella, Scuticella, 

Stenostomaria, Strongylopora, Strophipora, Talivittaticella, Terminocella, Vasignyella 

Family Concatenellidae Bock & Cook, 1996, p. 48 PRB-AQT 1 

Concatenella 

Family Eurystomellidae Levinsen, 1909, p. 314 AQT-Rec. 4 

Australianidae Powell, 1966, p. 19 

Eurystomella, Integripelta, Selenariopsis, Zygopalme 

Family Savignyellidae Levinsen, 1909, p. 273 YPR-Rec. 2 

7

Catenaridae d'Orbigny, 1851, p. 22 

Halysisis, Savignyella 

Family Petalostegidae Gordon, 1984, p. 46 LAN-Rec. 2 

Chelidozoum, Petalostegus 


Dicasignatella 

“Grade” HIPPOTHOOMORPHA Gordon, 1989, p. 25 CON-Rec. 32 

Superfamily Hippothooidea Busk, 1859, p. v, 24 [nom. transl. Gordon 1984, p. 108] CON-Rec. 30 

Family Hippothoidae Busk, 1859, p. v, 24 CON-Rec. 9 

Diazeuxidae Jullien, 1888, p. 8; Celleporellidae Norman, 1894, p. 129 

Antarctothoa, Austrothoa, Celleporella, Dacryoporella, Haplota, Hippothoa, Laterotecatia, Neothoa, 

Plesiothoa 

Family Chorizoporidae Vigneaux, 1949, p. 18 SRV-Rec. 2 

Chorizoporidae Harmer, 1957, p. 946 

Chorizopora, Costulostega 

Family Trypostegidae Gordon, Tilbrook & Winston in Winston, 2005 SAN-Rec. 12 

Boreas, Boreasina, Diplotresis, Eohippothoa, Grammothoa, Kronothoa, Kunradina, Mosathoa, 

Rectoporina, Tricolpopora, Trilophopora, Trypostega 

Family Haplopomidae nov. PIA-Rec. 1 

Haplopoma 

Family Pasytheidae Davis, 1934, p. 232 YPR-Rec. 6 

Epicaulidiidae Hincks, 1881, p. 156; Liriozoidae Levinsen, 1909, p. 313; Gemelliporidae Vigneaux, 1949, p. 62 

Dittosaria, Eutaleola, Gemellipora, Pasythea, Tecatia, Unifissurinella 

Superfamily Dysnoetoporoidea Voigt, 1971, p. 95 [nov.] CMP-MAA 2 

Family Dysnoetoporidae Voigt, 1971, p. 95 CMP-MAA 1 

Dysnoetopora 


Dysnoetocella 

“Grade” UMBONULOMORPHA Gordon, 1989, p. 28 [cf. Celleporina Smitt, 1868, p. 457] SAN-Rec. 138 

Superfamily Arachnopusioidea Jullien, 1888, p. 62 [Arachnopusiacea, nom. transl. Moyano 1970, p. 273] 

SAN-Rec. 12 

Family Arachnopusiidae Jullien, 1888, p. 62 (Arachnopusidae) SAN-Rec. 8 

Arachnopusia, Brendella, Briarachnia, Poricella, Ramicosticella, Staurosteginopora, Trichinopolia, 

Trilaminopora 

Family Exechonellidae Harmer, 1957, p. 651 ?YPR/?LUT-Rec. 4 

Stephanoporinae Gordon, 1993, p. 216 

Anexechona, Exechonella, Stephanopora, Xynexecha 

Superfamily Adeonoidea Busk, 1884, p. 177 [nom. transl. Gordon 1989, p. 34] YPR-Rec. 20 

Family Adeonidae Busk, 1884, p. 177 (Adeoneae) YPR-Rec. 15 

Adeonellidae Gregory, 1893, p. 225; Teichoporinae Gregory, 1893, p. 225; Meniscoporidae Canu, 1907, p. 153; 

Smittistomidae Vigneaux, 1949, p. 16; Anarthroporinae Vigneaux, 1949, p. 68; Dimorphocellinae Vigneaux, 1949, p. 

71 

Adeona, Adeonellopsis, Anarthropora, Bracebridgia, Dimorphocella, Kubaninella, Meniscopora, 

Ovaticella, Poristoma, Reptadeonella, Schizostomella, Smittistoma, Teichopora, Triporula, Trypocella 

Family Adeonellidae Gregory, 1893, p. 225 

PRB-Rec. 2 

Adeonellidae Vigneaux, 1949, p. 16; Laminoporinae Vigneaux, 1949, p. 68 

Adeonella, Laminopora 

8

Family Inversiulidae Vigneaux, 1949, p. 16 RUP-Rec. 1 

Inversiulidae Harmer, 1957, p. 955 

Inversiula 


Cucullipora, Goodonia 

Superfamily Pseudolepralioidea Silén, 1942, p. 51 [nom. transl. Gordon 1993, p. 218] CMP-Rec. 1 

Family Pseudolepraliidae Silén, 1942, p. 51 Rec. 1 

Pseudolepralia 

Superfamily Lepralielloidea Vigneaux, 1949, p. 68 (Lepraliellacea) SAN-Rec. 100 

Family Lepraliellidae Vigneaux, 1949, p. 17 SAN-Rec. 18 

Holoporellidae Levinsen, 1909, p. 347; Kleidionellidae Vigneaux, 1949, p. 17; Celleporariidae Harmer, 1957, p. 662; 

Buchneriidae Harmer, 1957, p. 876 

Acanthophragma, Balantiostoma, Buchneria, Celleporaria, Ceratopora, Cryptostomella, Dennisia, 

Drepanophora, Frurionella, Kladapheles, Kleidionella, Leiosellina, Lepraliella, Multescharellina, 

Pseudobeisselina, Schizemiellopsis, Sinuporaria, Sphaeropora 

Family Dhondtiscidae Gordon, 1989, p. 256 Rec. 1 

Dhondtiscus 

Family Bryocryptellidae Vigneaux, 1949 YPR-Rec. 10 

Bryocryptellidae Vigneaux, 1949, p. 19; Porellinae*** Vigneaux, 1949, p. 91 

Bryocryptella, Cyclocolposa, Cystisella, Marguetta, Palmiskenea, Porella, Porelloides, Reussia, 

Rhamphosmittina, Simibryocryptella 

Family Cheilhorneropsidae Annoscia et al., 1984, p. 108 PRB 1 

Cheilhorneropsis 

Family Romancheinidae Jullien, 1888, p. 60 CMP-Rec. 32 

Mucronellinae MacGillivray, 1887, pp. 191, 194, 212; Mucronellidae Levinsen, 1902, p. 26; Escharellidae Levinsen, 

1909, p. 314; Peristomellae Canu & Bassler, 1917; Exochellinae Bassler, 1935, p. 33; Perigastrellidae Vigneaux, 1949, 

p. 17; Hemicycloporinae Vigneaux, 1949, p. 83; Pleurolyrulinae Vigneaux, 1949, p. 105; Phylactelliporidae Bassler, 

1953, p. 217; Peristomellidae Kluge, 1962, p. 564 

Allerescha, Antarcticaetos, Arctonula, Bathosella, Biporochella, Bostrychopora, Bulbipora, Cheilonella, 

Elleschara, Escharella, Escharoides, Exochella, Hellerasca, Hemicyclopora, Hemiphylactella, Hippolyrula, 

Hippopleurifera, Hoplocheilina, Lageneschara, Metrarabdotomorpha, Neolagenipora, Obliquostoma, 

Ochetosella, Opaeomorpha, Pachykraspedon, Pleurolyrula, Pleuromucrum, Pseudosclerodomus, 

Psilopsella, Ragionula, Romancheina, Temachia 

Family Peedeesellidae CMP 1 

Peedeesella 

Family Sfeniellidae Gordon, 2006, p. 67 SRV-ZAN 2 

Sfeniella, Stephanollina 

Family Umbonulidae Canu, 1904, p. 18 LUT-Rec. 10 

?Escharellinidae d'Orbigny, 1852, pp. 339, 445; ?Porellinidae d'Orbigny, 1852, pp. 339, 475; Rhamphostomellidae 

Kluge, 1962, p. 534; Desmacystidae Dick & Ross, 19988, p. 30 

Aegyptopora, Astochoporella, Desmacystis, Escharopsis, Oshurkovia, Posterula, Rhamphostomella, 

Scorpiodina, Trigonopora, Umbonula 

Family Tessaradomidae Jullien & Calvet, 1903, pl. 14, p. 53 DAN-Rec. 6 

Beisselina, Pachydermopora, Pachythecella, Pavobeisselina, Pseudobathystomella, Tessaradoma 

Family Hincksiporidae Powell, 1968, p. 2293 Rec. 1 

Hincksipora 

Family Sclerodomidae Levinsen, 1909, p. 301 PIA-Rec. 4 

9

Cellarinellidae Moyano, 1970, p. 

Cellarinella, Cellarinelloides, Sclerodomus, Systenopora 

Family Metrarabdotosidae Vigneaux, 1949, p. 90 (Metrarabdotosinae) [nom. transl. Cheetham 1968, p. 61) 

PRB-Rec. 7 

Aequilumina, Biavicularium, Metrarabdotos, Polirhabdotos, Porometra, Rhabdotometra, Uniavicularium 

Superfamily Chlidoniopsoidea Harmer, 1957, p. 655 [nom. transl. d'Hondt 1985, p. 9, 11 (Chlidoniopsioidea 

sic)] PRB-Rec. 1 

Family Chlidoniopsidae Harmer, 1957, p. 655 PRB-Rec. 1 

Chlidoniopsis 

Superfamily incertae sedis 4 

Family Brydonellidae Taylor, Casadío & Gordon, 2008 CMP-DAN 4 

Brydonella, Bubnoffiella, Homalostega, Uharella 

”Grade” LEPRALIOMORPHA” Gordon, 1989, p. 34 [cf. Escharina Smitt, p. 453] DAN-Rec. 304 

Superfamily Smittinoidea Levinsen, 1909, p. 335 [nom. transl. Vigneaux 1949, p. 90] DAN-Rec. 43 

Family Smittinidae Levinsen, 1909, p. 335 YPR-Rec. 25 

Smittidae Jullien, 1888, p. 52; **Schismoporellineae Gregory, 1893, p. 225; Phylactellidae Canu & Bassler, 1917, p. 66 

Aspericreta, Dakariella, Dengordonia, Dittomesia, Hemismittoidea, Hippophylactella, Houzeauina, 

Parasmittina, Pemmatoporella, Phylactella, Phylactellina, Plagiosmittia, Platychelyna, Pleurocodonellina, 

Porismittina, Prenantia, Pseudoflustra, Raymondcia, Rimulostoma, Schismoporella, Smittina, Smittinella, 

Smittoidea, Thrypticocirrus, Tracheloptyx 

Family Bitectiporidae MacGillivray, 1895, p. 106 YPR-Rec. 13 

Hippoporinidae Brown, 1952, p. 267 

Bitectipora, Cribella, Hippomonavella, Hippoporina, Hippothyris, Kermadecazoon, Metroperiella, 

Neodakaria, Nigrapercula, Parkermavella, Pentapora, Schizomavella, Schizosmittina 


Tubescharina 

Family Watersiporidae Vigneaux, 1949, p. 15 ?MES-Rec. 4 

Dakariinae Vigneaux, 1949, p. 77 

Pachycleithonia, Uscia, Veleroa, Watersipora 

Superfamily Schizoporelloidea Jullien, 1883, p. 527 [Schizoporellacea Vigneaux 1949, p. 55] 151 

Family Schizoporellidae Jullien, 1883, p. 527 YPR-Rec. 7 

Enoplostomella, Ferganula, Hipposera, Schizoporella, Schizobrachiella, Schizolepralia, Stylopoma 

Family Stomachetosellidae Canu & Bassler, 1917, p. 44 ?LUT/?BAR-Rec. 12 

Metrocryptidae Vigneaux, 1949, p. 20; Tremoschizodininae Vigneaux, 1949, p. 56 

Cycloperiella, Fatkullina, Junerossia, Lepralioides, Pachyegis, Marcusadorea?, Metradolium, Metrocrypta, 

Schizemiella, Stephanotrema, Stomachetosella, Tremoschizodina 

Family Tetraplariidae Harmer, 1957, p. 1052 LUT-Rec. 2 

Bigemellariidae MacGillivray, 1895, p. 7 

Tetraplaria, Tychinella 

Family Bryobaculidae Rosso, 2002, p. 275 

PLI 1 

Bryobaculum 

Family Phorioppniidae Gordon & d'Hondt, 1997, p. ?RUP/?CHT-Rec. 4 

Oppiphorina, Phorioppnia, Punctiscutella, Quadriscutella 

Family Porinidae d'Orbigny, 1852, p. 431 ?YPR-Rec. 7 

Acroporidae Canu, 1913, p. ?; Gastropellinae Vigneaux, 1949, p. 76; Spiroporinidae Harmer, 1957, p. 845; 

Haswellinidae Gordon & Ballantine, 1977, p. 126 

Cyphonella, Gastropella, Haswelliporina, Mosaicoporina, Porina, Semihaswellia, Tremotoichos 

10

Family Margarettidae Harmer, 1957, p. 824 YPR-Rec. 2 

Tubucellariadae Busk, 1884, p. 98 

Margaretta, Tubucella 

Family Myriaporidae Gray, 1841, p. 150 PRB-Rec. 4 

Myriozoumidae d'Orbigny, 1849, p. 503; Myriozoidae Smitt, 1868, p. 454; Myriozoumidae Calvet in Jullien & Calvet, 

1903, p. 129 

Gemelliporina, Leieschara, Myriapora, Myriozoella 

Family Hippopodinidae Levinsen, 1909, p. 353 LUT-Rec. 5 

Archoporella, Hippopodina, Saevitella, Thornelya, Trilochites 

Family Pacificincolidae Lu, 1999, p. 338 Rec. 

2 

Pacificincola, Primavelans 

Family Hippaliosinidae Winston, 2005, p. 74 Rec. 1 

Hippaliosina 

Family Duvergieriidae Vigneaux, 1949, p. 68 (Duvergieriinae) LUT-BUR 3 

Schizotremoporinae Vigneaux, 1949, p. 69 

Duvergieria, Calvetina, Schizotremopora 

Family Gigantoporidae Bassler, 1935, p. 32 ?PRB-Rec. 5 

Aptonella, Cosciniopsis, Gephyrophora, Gigantopora, Hemicosciniopsis 

Family Lanceoporidae Harmer, 1957, p. 983 PRB-Rec. 3 

Parmulariidae Canu & Bassler, 1927, p. 35 

Calyptotheca, Emballotheca, Lanceopora 

Family Cheiloporinidae Bassler, 1936, p. 161 ?PRB-Rec. 7 

Cheiloporinidae Vigneaux, 1949, p. 17 

Cheilopora, Cheiloporina, Crustoporina, Cyttaridium, Hagiosynodos, Pachystomaria, Retelepralia 

Family Cryptosulidae Vigneaux, 1949, p. 14 TOR-Rec. 2 

Cryptosula, Harmeria 

Family Actisecidae Harmer, 1957, p. 854 Rec. 1 

Actisecos 

Family Teuchoporidae Neviani, 1895, p. 116 (Teuchoporinae) [nom. transl. Harmer 1957, p. 896] ?ZAN-Rec. 3 

Lagenicellidae Banta & Santagata, 1996, p. 308 

Coleopora, Lagenicella, Teuchopora 

Family Echinovadomidae Tilbrook, Hayward & Gordon, 2001, p. 83 

Rec. 1 

Echinovadoma 

Family Phoceanidae Vigneaux, 1949, p. 18 ?ZAN-Rec. 2 

Phoceana, Sertulipora 

Family Mawatariidae Gordon 1990, p. 471 Rec. 1 

Mawatarius 

Family Vicidae Gordon, 1988, pp. 284, 289 ?ZAN-Rec. 3 

Cyclostomella, Vibraculina, Vix 

Family Robertsonidridae Rosso, Sciuto & Sinagra, 2010 ?PIA-Rec. 2 

Bertorsonidra, Robertsonidra 

11


Tshokrakopora 

Family Microporellidae Hincks, 1879, p. 156 AQT-Rec. 12 

Fenestrulinidae Jullien, 1888, p. 8 

Adelascopora, Calloporina, Chronocerastes, Cribriporella, Diporula, Fenestrulina, Fenestruloides, 

Flustramorpha, Microporella, Microporelloides, Pseudoadelascopora, Tenthrenulina 

Family Calwelliidae MacGillivray, 1887, pp. 193, 198 Rec. 7 

Onchoporidae Busk, 1884, p. 102 

Calwellia, Ichthyaria, Ijimaiaellia, Malakosaria, Onchoporella, Onchoporoides, Wrigiana 

Family Petraliidae Levinsen, 1909, p. 350 Rec. 1 

Petralia 

Family Petraliellidae Harmer, 1957, p. 692 YPR-Rec. 10 

Discoporidae Smitt, 1868, p. 457 

Hippopetraliella, Mobunula, Mucropetraliella, Orbiculipora, Petraliella, Poripetraliella, Riscodopa, 

Serripetraliella, Sinupetraliella, Utinga 

Family Cyclicoporidae Hincks, 1884, p. 279 BRT-Rec. 1 

Cyclicopora 

Family Lacernidae Jullien, 1888, p. 48 PRB-Rec. 14 

Arthropoma, Clithriellum, Cribellopora, Cylindroporella, Lacerna, Nimba, Nimbella, Phonicosia, Ralepria, 

Rogicka, Schizaropsis, Toretocheilum, Vitrius 

Family Escharinidae Tilbrook, 2006, p. 250 LUT-Rec. 10 

Hippozeugosellidae Vigneaux, 1949, p. 15; Schizobathysellidae Vigneaux, 1949, p. 20; Mastigophorinae Vigneaux, 

1949, p. 83 

Bactridium, Bryopesanser, Chiastosella, Dightonia, Escharina, Herentia, Hippomenella, Phaeostachys, 

Schizobathysella, Schizocoryne, Therenia 

Family Acoraniidae López-Fé, 2006, 1809 Rec. 1 

Acorania 


Hippadenella, Julianca, Kymella, Schizoporellopsis 

Family Buffonellodidae Gordon & d'Hondt, 1997, p. 41 5 

Buffonellidae Jullien, 1888, p. 47 

Aimulosia, Buffonellodes, Chataimulosia, Ipsibuffonella, Maiabuffonella 

Family Jaculinidae Zabala, 1986, p. 186 1 

Jaculina 

Family Eminooeciidae Hayward & Thorpe, 1988, p. 4 Rec. 3 

Eminooecia, Isoschizoporella, Macrocamera 

Superfamily Urceoliporoidea Bassler, 1936, p. 160 [nom. transl. Gordon 1990, pp. 470] BUR-Rec. 4 

Family Urceoliporidae Bassler, 1936, p. 160 BUR-Rec. 3 

Urceoliporidae Vigneaux, 1949, p. 14 

Cureolipora, Reciprocus, Urceolipora 

Family Prostomariidae MacGillivray, 1895, p. 105 LAN 1 

Prostomaria 

Superfamily Didymoselloidea Brown, 1952, p. 194 [nom. transl. d'Hondt 1985, pp. 9, 11] ?LUT/?BRT-Rec. 3 

Family Didymosellidae Brown, 1952, p. 194 ?LUT/?BRT-Rec. 2 

Didymosella, Tubiporella 

12


Bimicroporella 

Superfamily Euthyriselloidea Bassler, 1953, p. 226 [nom. transl. d'Hondt 1985, pp. 9, 11] Rec. 5 

Family Euthyrisellidae Bassler, 1953 Rec. 4 

Euthyridae Levinsen, 1909, p. 269; Euthyrisellidae Harmer, 1957, p. 874 

Euthyrisella, Pleurotoichus, Pseudoplatyglena, Tropidozoum 

Family Neoeuthyridae Cook, nov. Rec. 1 

Neoeuthyris 

Superfamily Siphonicytaroidea Harmer, 1957, p. 892 [nov.] YPR-Rec. 1 

Family Siphonicytaridae Harmer, 1957, p. 892 YPR-Rec. 1 

Siphonicytara 

Superfamily Mamilloporoidea Canu & Bassler, 1927, p. 9 [nom. transl. d'Hondt 1985, pp. 9, 11] YPR- 

Rec. 15 

Family Mamilloporidae Canu & Bassler, 1927, p. 9 YPR-Rec. 2 

Anoteropora, Mamillopora 

Family Crepidacanthidae Levinsen, 1909, p. 266 SRV-Rec. 1 

Crepidacantha 

Family Cleidochasmatidae Cheetham & Sandberg, 1964, p. 1032 LAN-Rec. 6 

Anchicleidochasma, Calyptooecia, Characodoma, Cleidochasmidra, Fedorella, Yrbozoon 

Family Ascosiidae Jullien, 1883, p. 526 (Ascosidae) LUT-Rec. 4 

Ascosia, Fedora, Kionidella, Stenosipora 


Prattia, Schizorthosecos 

Superfamily Celleporoidea Johnston, 1838, p. 247 [nom. transl. Gordon 1984, p. 114] DAN-Rec. 55 

Family Colatooeciidae Rec. 3 

Cigclisula, Colatooecia,Trematooecia, 

Family Celleporidae Johnston, 1838, p. 247 PRB-Rec. 23 

Lageniporidae Jullien, 1883, p. 526; Osthimosidae Jullien, 1888, p. 63; Schismoporinae Gregory, 1893, p. 224; 

Galeopsidae Jullien & Calvet, 1903, pl. 14, p. 53; Tegminulidae Vigneaux, 1949, p. 20; Schismoporidae Vigneaux, 

1949, p. 86; Celleporinidae Harmer, 1957, p. 899 

Buffonellaria, Buskea, Calvipelta, Cellepora, Celleporina, Chasmatooecium, Chasmazoon, Favosthimosia, 

Galeopsis, Lagenipora, Omalosecosa, Orthoporidroides, Osthimosia, Palmicellaria, Pourtalesella, 

Predanophora, Pseudocelleporina, Richbunea, Scoriodinipora, Sinuporina, Spigaleos, Tegminula, 

Turbicellepora 

Family Torquatellidae Tilbrook, 2006, p. 286 Rec. 1 

Torquatella 

Family Hippoporidridae Vigneaux, 1949, p. 80 (Hippoporidrinae, nom. transl. Pouyet 1973, p. 54) BUR-Rec. 4 

Hippoponellidae Kluge, 1962, p. 530 

Hippoporella, Hippoporidra, Hippotrema, Odontoporella 

Family Phidoloporidae Gabb & Horn, 1862, p. 138? DAN-Rec. 24 

Reteporidae Smitt, 1868, p. 458; Schizoreteporinae Gregory, 1893, p. 224; Sertellidae Jullien in Jullien & Calvet, 1903, 

p. 57; Hippoporininae Bassler, 1935, p. 33; Hippellozo[on]idae Vigneaux, 1949, p. 17; Rhynchozo[on]inae Vigneaux, 

1949, p. 102; Hippoporinidae Osburn, 1952, p. 343 

Bryorachis, Chevron, Dentiporella, Dictyochasma, Fodinella, Hippellozoon, Hippopozoon, Iodictyum, 

Lifuella, Malleatia, Metacleidochasma, Phidolopora, Plesiocleidochasma, Psileschara, Psilosecos, 

Reteporella, Reteporellina, Rhynchozoon, Schedocleidochasma, Schizoretepora, Schizotheca, 

Sparsiporina, Stephanollona, Triphyllozoon 

13


Pseudostega 

Superfamily Conescharellinoidea Levinsen, 1909, p. 308 [nom. transl. d’Hondt 1985, pp. 9, 11] YPR-Rec. 

27 

Family Batoporidae Neviani, 1900, p. 220 (Batoporideae, nom. transl.) YPR-Rec. 2 

Batopora, Lacrimula, 

Family Conescharellinidae Levinsen, 1909, p. 308 YPR-Rec. 7 

Bipora, Conescharellina, Crucescharellina, Flabellopora, Ptoboroa, Trochosodon, Zeuglopora 

Family Lekythoporidae Levinsen, 1909, p. 313 CHT-Rec. 9 

Aulopocellidae Vigneaux, 1949, p. 20 

Aulopocella, Catadysis, Harpagozoon, Jugescharellina, Lekythopora, Orthoporidra, Poecilopora, 

Terataulopocella, Turritigera 

Family Orbituliporidae Canu & Bassler, 1923, p. 186 YPR-Rec. 8 

Atactoporidra, Batoporella, Bicupularia, Conescharellinopsis, Orbitulipora, Sphaerulobryozoon, 

Stichoporina, Xaveropora 


Family Cuvillieridae Annoscia, 1965, p. 6 LUT 1 

Cuvilliera 

SUMMARY OF GENERIC/SUBGENERIC DIVERSITY 

ORDER CHEILOSTOMATA 1064 

Suborder Malacostegina 32 

Suborder Inovicellina 2 

Suborder Scrupariina 5 

Suborder Neocheilostomina 1025 

Infraorder Flustrina 372 

Infraorder Ascophorina 653 

“Grade” Acanthostega 179 

“Grade” Hippothoomorpha” 32 

"Grade” Umbonulomorpha" 138 

"Grade” Lepraliomorpha" 304 

RATIO OF “ANASCA” (Malacostegina, Inovicellina, Scrupariina, Flustrina) : ASCOPHORINA = 

411 : 653 

1. The above list is only an ongoing attempt to sort order out of chaos. Consequently you 

will probably see immediate areas for improvement. Good. Please send me corrections and 

improvements. Please understand that the classification is evolving and a rigorous scheme is 

dependent on phylogenetic analysis. 

2. One of the benefits of this list is to give an indication of relative diversity of various 

groupings, e.g., anascans : ascophorans, or proportions of frontal-shield morphologies 

among the Ascophorina. 

3. At first glance it appears there are a disproportionate number of small, even monogeneric, 

families (especially among the lepralioid ascophorines). Is this a consequence of oversplitting 

and too much weight accorded to single characters? On the other hand, some families are 

rather large, with a range of disparate morphologies, and appear to need splitting, e.g., 

Calloporidae. The Cribrilinidae needs splitting too, there being several major lineages, and 

14

some of Lang's family names will be retainable. For the moment it is worth noting that the 

oldest-available valid family-group name for any of the Cribrilinoidea is the Steginoporidae 

d'Orbigny, 1853, which Lang for some curious reason chose to ignore while accepting the 

genus Steginopora (see also Bassler 1953). 

4. I plan on expanding the list of generic synonyms to include those already in Bassler. I 

haven't done this yet. 

5. The suggested new families and superfamilies are just that — suggestions. If others agree 

that these are appropriate, I don't care who formally establishes any of them, the main job is 

make progress. 

6. The suborder Protocheilostomat[in]a Silén, 1942 has been deleted from this compilation. 

Its sole genus Labiostomella is uncalcified (Silén 1944, pp. 7, 8), has a ctenostomatous mode 

of reproduction and closing apparatus, and nests with the Benedeniporidae in cladograms. All 

available evidence therefore indicates that Labiostomella is a ctenostome and not a 

cheilostome. It needs to be ascertained if Benedeniporidae Delage & Hérouard, 1897 and 

Labiostomellidae Silén, 1942 are confamilial. Two superfamilies were proposed ⎯⎯ 

Labiostomellacea (Phaeophanova 1960, p. 155) and Benedeniporoidea (Jebram 1973, p. 34). 

7. Priority of taxon names. Article 23.1. [Statement of the Principle of Priority] in the fourth 

edition of the ICZN (1999) reads: “The valid name of a taxon is the oldest available name 

applied to it, unless that name has been invalidated or another name is given precedence by 

any provision of the Code or by any ruling of the Commission.” Note also Article 23.9. 

[Reversal of precedence] which reverses the Principle of Priority if the senior synonym does 

not now satisfy two conditions: 1) it has not been used as a valid name after 1899, and 2) the 

junior synonym has been used in at least 25 works published by at least 10 authors in the 

immediately preceding 50 years and encompassing a span of not less than 10 years. 

8. Note Article 35.5. Precedence for names in use at higher rank: “If after 1999 a name in use 

for a family-group taxon (e.g. for a subfamily [or superfamily]) is found to be older than a 

name in prevailing usage for a taxon at higher rank in the same family-group taxon … the 

older name is not to displace the younger name.” This helpful ruling affects some names of 

family-group taxa in recent versions of the cheilostome classification scheme that have not 

yet been formally established through publication. For example, when I proposed the 

superfamily Calloporoidea (Gordon 1989), I chose not to adopt the Antroporacea of Vigneaux 

(1949) because the family Antroporidae Vigneaux, 1949 was not in current use, Antropora 

being traditionally classified with calloporids. In his discussion of the familial relationships of 

Antropora, Tilbrook (1998) remarked on the heterogeneous composition of the Calloporidae 

(see also note 3 above) and suggested that adoption of the Antroporidae, inter alia, would 

ease the problem. He did not comment on the use of Antroporacea (Antroporoidea). The 

classification given herein uses the Antroporidae following Tilbrook’s suggestion but, 

following Article 35.5, it is not necessary to overturn Calloporoidea in favour of 

Antroporoidea. 

Taxa added 15 September 2011: Arbocuspis, Arbopercula, Osburnea 

Taxa added 25 August 2011: Favosthimosia, Mourellina, Mourellinidae, Rectimicropora, 

Schizocoryne Scriblitopora; Crepis transferred to Calloporidae 

Taxa added 15 December 2010: Corbuliporina, Eutaleola (replaces Euteleia), Uharella 

Taxon reinstated 29 June 2010: Scorpiodinipora 

Taxon reinstated 30 April 2010: Rhammatopora 

15

Taxon deleted 1 March 2010: Alismittina 

Taxon added 1 February 2010: Marcusadorea 

Taxon added 3 August 2009: Microcorcillum 

Taxa added 16 June 2009: Harpagozoon [replaces Harpago], Ijimaiellia [replaces Ijimaia], 

Reussinella [replaces Reussina], Leiosellina [replaces Leiosella] 

Taxa added 3 February 2009: Callaetea, Einhornia, Elleschara (reinstated, substituting for 

Haywardipora in previous list] 

Taxon composition changed 16 December 2008: Colatooeciidae is suggested also to include 

Cigclisula and Trematooecia, both transferred from Stomachetosellidae 

Taxon added 21 November 2008: Turnerellina [replaces Turnerella] 

Taxon deleted 11 October 2007: Strophiella (junior subjective synonym of Schizotheca) [see 

Reverter-Gil & Fernández-Pulpeiro 2007] 

Taxon added 11 October 2007: Amphibiobeania 

Taxon added 13 September 2007: Septentriopora; Rosseliana transferred to Antroporidae 

Taxon deleted 5 September 2007: Scorpiodinipora 

Taxon added 17 August 2007: Primavelans 

Taxon added 19 June 2007: Oshurkovia 

Taxon added 14 February 2007: Junerossia 

Taxa added 8 January 2007: Acorania, Acoraniidae, Sfeniella, Sfeniellidae 

Taxa added 28 September 2006: Archoporella, Orbiculipora 

Taxa added 25 August 2006: Escharinidae, Peedeesellidae, Torquatellidae 

16 

Dennis P. Gordon 

15 September 2011

BRYOZOA: CHEILOSTOMATA Interim classification for Treatise ...

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