polychaetous annelids from the south china sea - Raffles Museum of ...

THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8: 209-232 

© National University of Singapore 

POLYCHAETOUS ANNELIDS FROM THE SOUTH CHINA SEA 

Hannelore Paxton 

Department ofBiological Sciences. Macquarie University. NS. W 2109, Australia 

L. M. Chou 

Department ofBiological Sciences, National University of Singapore, Kent Ridge 119260, 

Republic of Singapore 

ABSTRACT. - A checklist of polychaetous anneIids from the South China Sea is 

generated from available literature. On record are 661 species belonging to 54 families. 

Biodiversity distribution appears linked to collecting efforts rather than a reflection 

of the real situation. The potential for discovery of new species remains high. 

INTRODUCTION 

This check-list, based mainly on literature reports, lists 661 species ofpolychaetous annelids 

from the South China Sea. The apparently highest biodiversity ranked as mainland China, 

Vietnam, Hong Kong, Taiwan, Singapore, Philippines, Thailand, Malaysia, is due mainly 

to the collecting efforts of residing specialists or expeditions rather than true biodiversity. 

The polychaetes listed belong to 54 families. The families are grouped according to Hartman 

(1959), except for the Eunicidae and related families. We are emphasising that this list is 

primarily based on literature rather than personal knowledge and are presenting references 

for each record. This list is only a baseline; we expect it to change considerably with the 

removal of names as families are revised, and the addition of new species, as the areas of 

the South China Sea become studied more extensively. This is publication number 300 of 

the Commonwealth Key Centre for Biodiversity and Bioresources, Macquarie University. 

NAME 

Aphroditidae 

Aphrodita talpa 

Hermonia hystrix 

Pontogenia indica 

Pontogenia lichaucoi 

AUTHOR 

Quatrefages, 1865 

Hartman, 1959 

Grube, 1878 

Pillai, 1965 

209 

COUNTRY 

China 

Philippines 

China 

Philippines 

Singapore 

Philippines 

REFERENCE 

Yang & Sun 1988:133 

Pillai 1965:112 

Yang & Sun 1988: 133 

Grube 1878:19 

Tan & Chou 1993:28I 

Pillai 1965:112

Paxton & Chou: Polychaetous annelids from the South China Sea 

NAME AUTHOR COUNTRY REFERENCE 

Pontogenia nuda Horst, 1917 China Yang & Sun 1988:135 

Polynoidae 

Eulagisca sp. Hong Kong Mak 1982:599 

Gastrolepidia clavigera Sehmarda, 1861 China Wu et al. 1980:114 

Halosydna brevisetosa Kinberg, 1855 Hong Kong Mak 1982:599 

Harmothoe ampullijera (Grube, 1878) Philippines Pillai 1965: 117 

Harmothoe dictyophora (Grube, 1878) S. Vietnam Gallardo 1968:47 

China Yang & Sun 1988:148 

Harmothoe extenuata (Grube, 1840) Philippines Natividad & Palpal-Latoe 1986:296 

Harmothoe imbricata (Linnaeus, 1767) Hong Kong Shin 1982: 162 

Harmothoe minuta (Potts, 1910) China Wu et al. 1980:114 

Harmothoe nigricans Horst, 1915 China Wu et al. 1980:114 

Harmothoe (Lagisca) waahli Kinberg, 1855 Philippines Natividad & Palpa1-Latoe 1986:294 

Hermenia acantholepis (Grube, 1876) China Wu et al. 1980:114 

Hololapidella commensalis Willey,1905 China Wu et al. 1980:114 

Intoshella aphthalma Gallardo, 1968 S. Vietnam Gallardo 1968:47 

?Intoshella holothuricola (Izuka, 1912) S. Vietnam Gallardo 1968:48 

Iphione muricata (Savigny, 1818) China Wu et al. 1980:114 

Hong Kong Mak 1982:599 

Singapore Tan & Chou 1993:281 

Lepidonotus carinulatus Grube, 1870 Philippines Pillai 1965:115 

Lepidonotus cristatus (Grube, 1876) China Wu et al. 1980:114 

Lepidonotus hedleyi Benham, 1915 Philippines Natividad & Palpal-Latoe 1986:290 

Lepidonotus jacksoni Kinberg, 1855 Philippines Natividad & Palpal-Latoe 1986:286 

Lepidonotus squamatus (Linnaeus, 1767) Hong Kong Mak 1980:599 

Lepidonotus tenuisetosus (Gravier, 1901) Hong Kong Shin 1982:162 

Lepidasthenia izukai Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:46 


Lepidasthenia microlepis Potts, 1910 China Wu et al. 1980:114 

Lucopia magnicirra Pillai, 1965 Philippines Pillai 1965: 117 

Parahalosydnopsis hartmanae Pettibone, 1977 Hong Kong Shin 1982:162 

Paralepidonotus ampullijerus (Grube, 1878) Singapore Tan & Chou 1993:281 


Scalisetosus jragilis (Claparede, 1868) China Wu & Wu 1987:79 

Scalisetosus longicirrus (Sehmarda, 1861) China Wu et al. 1980:114 

Thormora jukesii Baird, 1865 China Wu et al. 1980:114 

Philippines Natividad & Palpa1-Latoe 1986:285 

Polyodontidae 

Eupanthalis maculosa (Treadwell, 1931) China Wu et al. 1980:114 

Neopolyodontes lepidus Shen & Wu, 1993 China Shen & Wu 1993:537 

Neopanthalis muricatus Shen & Wu, 1993 China Shen & Wu 1993:537 

Panthalis jogasimae Izuka, 1912 China Wu et al. 1980:114 

Polyodontes gracilis Pflugfelder, 1932 S. Vietnam Gallardo 1968:49 

Polyodontes maxillosus (Ranzani, 1817) China Wu et al. 1980:114 

Polyodontes melanonotus (Grube, 1876) Hong Kong Shin 1982:162 


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THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8 


Sigalionidae 

Ehlersileanira incisa (Grube, 1877) China Yang & Sun 1988:154 

Euleanira ehlersi Horst, 1916 S. Vietnam Gallardo 1968:50 

Mayel/a sp. Taiwan Hsieh, pers. comm. 

Psammolyce fijiensis Mclntosh, 1885 S. Vietnam Gallardo 1968:50 

Psammolyce malayana Horst, 1913 Philippines Natividad & Pa1pal-Latoc 1986:301 

Sigalion sp. Taiwan Hsieh, pers. comm. 

Sthenelais boa (Johnston, 1839) Hong Kong Shin 1982: 162 

Sthenelais mitsuii (Okuda, 1938) S. Vietnam Gallardo 1968:51 

Sthenelais nami Gallardo, 1968 S. Vietnam Gallardo 1968:51 

Augener, 1927 S. Vietnam Gallardo 1968:52 

Sthenelais japonica (Mclntosh, 1885) S. Vietnam Gallardo 1968:52 

Sthenolepis izuensis (Takahashi, 1938) China Wu et a1. 1980:115 

Sthenolepis tentaculata (Horst, 1917) China Wu et a1. 1980: 115 

Thalenessa oculata (Peters, 1854) China Wu & Wu 1987:79 

Thalenessa cf. spinosa asiatica Uschakov & Wu, 1959 Taiwan Hsieh, pers. comm. 

Thalenessa stylolepis Willey, 1905 S. Vietnam Gallardo 1968:53 

Thalenessa tropica Hartman, 1954 S. Vietnam Gallardo 1968:53 

Eulepethidae 

Pareulepis malayana (Horst, 1913) S. Vietnam Gallardo 1968:54 

Chrysopetalidae 

Bhawania brevis Gallardo, 1968 S. Vietnam Gallardo 1968:54 

Hong Kong Shin 1982: 162 

Bhawania cryptocephala Gravier, 1901 China Yang & Sun 1988:158 

Bhawania goodei Webster, 1884 Hong Kong Mak 1982:599 

Philippines Natividad & Palpal-Latoc 1986:304 

China Wu & Wu 1987:79 

Chrysopetalum ehlersi Gravier, 1901 China Yang & Sun 1988:160 

Paleanotus bel/is (Johnson, 1897) Hong Kong Mak 1982:599 

Paleanotus chrysolepis Schmarda, 1861 China Yang & Sun 1988:161 

Paleanotus debilis (Grube, 1855) China Wu et a1. 1980:115 

Amphinomidae 

Amphinome pulchra Horst, 1912 China Wu et a1. 1980:115 

Amphinome rostrata (Pallas, 1766) China Yang & Sun 1988:164 

Philippines Natividad & Pa1pal-Latoc 1986:321 

Chloeia flava (Pallas, 1766) S. Vietnam Gallardo 1968:56 

Hong Kong Shin 1982:163 




Chloeia fusca Mclntosh, 1885 China Yang & Sun 1988:165 

Chloeia entypa Chamberlin, 1919 Philippines Natividad & Palpal-Latoc 1986:321 

Chloeia inermis Quatrefages, 1865 China Wu & Wu 1987:79 

Chloeia parva Baird, 1870 S. Vietnam Gallardo 1968:56 


Chloeia rosea Pous, 1909 Philippines Natividad & Pa1pa1-Latoc 1986:324 

211



Chloeia violacea Horst, 1910 S. Vietnam Gallardo 1968:56 

China Wu et at. 1980:115 

Eurythoe complanata (pallas, 1766) Philippines Pillai 1965:119 



Eurythoe parvecarunculata Horst, 1912 China Wu et at. 1980:115 

Unopherus hirsuta (Wesenberg-Lund, 1949) Hong Kong Shin 1982: 163 

Notopygos gigas Horst, 1911 China Wu et at. 1980:115 

Notopygos labiatus Fauvel, 1932 Philippines Natividad & Palpal-Latoe 1986:326 

Notopygos ornata Grube, 1856 Philippines Natividad & Palpal-Latoe 1986:326 

Notopygos sibogae Horst, 1911 China Wu et at. 1980:115 

Notopygos supragigas Usehakov & Wu, 1962 China Wu et a.11980:115 

Notopygos variabilis Grube, 1855 Philippines Pillai 1965: 120 

Paramphinome indica Fauvel, 1932 China Wu et at. 1980:115 

Pareurythoe borealis (Sars, 1862) China Yang & Sun 1988:170 

Pherecardia striata (Kinberg, 1857) Philippines Pillai 1965:121 


Pseudeurythoe hirsuta Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:57 

Pseudeurythoe oligobranchia Wu et aI, China Wu et at. 1980:115 

Euphrosinidae 

Euphrosyne foliosa Audouin & M. Edw., 1833 China Sun et at. 1978:161 

Euphrosine myrtosa Savigny, 1818 Hong Kong Mak 1982:599 


Phyllodocidae 

Anaitides madeirensis (Langerhans, 1880) China Sun et at. 1978:135 as P.(Anait.)mad. 



Anaitides tenuissima (Grube, 1878) Philippi nes Grube 1878:95 

Anaitides ?tenuissima S. Vietnam Gallardo 1968:57 

Eteone (Mysta) siphodonta (Delle Chiaje, 1822) Singapore Tan & Chou 1993:282 

Eteone (Mysta) tchangsii Usehakov & Wu, 1959 China Yang & Sun 1988:109 

Eulalia (Pterocirrus) magalhaensis Kinberg, 1866 Singapore Tan & Chou 1993:282 

Eulalia viridis (Linnaeus, 1767) Hong Kong Mak 1982:599 

China Yang & Sun 1988: 105 

Eumida sanguinea (Marenzeller, 1879) Hong Kong Mak 1982:599 

Genetyllis castanea (Marenzeller, 1879) Hong Kong Mak 1982:599 

China Yang & Sun 1988: 108 

Philippines Natividad & Palpal-Latoe 1986:275 

Genetyllis sp. Taiwan Hsieh, pers. eomm. 

Mysta ornata (Grube, 1878) S. Vietnam Gallardo 1968:58 


Notophyllum splendens (Sehmarda, 1861) China Wu et at. 1980:115 

Paranaitis zeylanica (Willey, 1905) S. Vietnam Gallardo 1968:58 

Phyllodoce (Anaitides) dissotyla Willey, 1905 China Wu & Sun 1978:176 

Phyllodoce (Anait.) madeirensis Langerhans, 1880 China Wu et at. 1980:115 

Phyllodoce fristedti Bergstriim, 1914 China Wu et at. 1980:115 

212



Phyllodoce ?laminosa Savigny, 1818 S. Vietnam Gallardo 1968:58 

Phyllodoce malmgreni Gravier, 1900 China Yang & Sun 1988:110 

Pterocirrus macroceros (Grube, 1860) China Yang & Sun 1988:111 

Alciopidae 

Alciopina parasitica Claparecte & Panceri, 1867 China Shen 1978:215 

Krohnia lepidota (Krohn, 1845) China Sun & Wu 1979:64 

Naiades cantrainii Delle Chiaje, 1830 China Shen 1978:211 

Plotohelmis capitata (Greeff, 1876) China Shen 1978:214 

Rhynchonerella petersii (Langerhans, 1880) China Sun & Wu 1979:63 

Rhynchonerella gracilis Costa, 1864 China Wu et at. 1980:116 

Rhynchonerella xishaensis Shen, 1978 China Shen 1978:214 

Vanadis crystallina Greeff, 1876 China Shen 1978:212 

Vanadis juscapunctata Treadwell, 1906 China Shen 1978:212 

Vanadis minuta Treadwell, 1906 China Shen 1978:212 


Lopadorrhynchidae 

Lopadorrynchus brevis Grube, 1855 China Shen 1978:203 

Lopadorrynchus krohni (Claparecte, 1870) China Shen 1978:205 

Lopadorrhynchus tuberculus Shen, 1978 China Shen 1978:205 

Lopadorrhynchus uncinatus Fauvel, 1915 China Shen 1978:203 

Maupasia caeca Vignier, 1886 China Shen 1978:207 

Pedinosoma curtum Reibisch, 1895 China Shen 1978:207 

Pelagobia longicirrata Greeff, 1879 China Shen 1978:206 

Lacydoniidae 

Paralacydonia paradoxa Fauvel, 1913 S. Vietnam Gal1ardo 1968:59 



Iospilidae 

Phalacrophorus pietus Greeff, 1879 China Shen 1978:208 

Phalocrophorus uniformis Reibisch, 1895 China Shen 1978:209 

Pontodoridae 

Pontodora pelagica Greef, 1879 China Shen 1978:209 

Typhloscolecidae 

Sagitella kowalevskii Wagner, 1872 China Shen 1978:222 

Travisiopsis dubia Stop-Bowitz, 1948 China Wu et at. 1980:117 

Travisiopsis levinseni Southern, 1910 China Shen & Wu 1978:105 

Travisiopsis lobifera Levinsen, 1885 China Shen 1978:223 

Typhloscolex muelleri Busch, 1851 China Shen 1978:221 

Tomopteridae 

Tomopteris apsteini Rosa, 1908 China Sun & Wu 1979:65 

Tomopteris cavalii Rosa, 1908 China Shen 1978:218 

Tomopteris duccii Rosa, 1908 China Shen 1978:220 

213



Tomopteris dunckeri Rosa, 1908 China Wu et al. 1980: 117 

Tomopteris elegans Chun, 1888 China Shen 1978:217 

Tomopteris ligulata Rosa, 1908 China Shen 1978:220 

Tomopteris mariana Greef, 1885 China Wu et al. 1980:117 

Tomopteris planktonis Apstein, 1900 China Shen 1978:218 

Tomopteris rolasi Greef,1885 China Shen & Wu 1978:103 

Tomopteris septentrionalis Steenstrup, 1849 China Sun & Wu 1979:67 

Hesionidae 

Bonuania parva Pillai, 1965 Phillippines Pillai 1965: 123 

Gyptis labatus (Hessle, 1925) China Yang & Sun 1988:80 

Gyptis cf. maraunibinae Taiwan Hsieh, pers. eomm. 

Hesione genetta Grube, 1866 China Wu et al. 1980: 117 

Hesione intertexta Grube, 1878 Philippines Grube 1878: 

China Wu et al. 1980:117 

Hesione splendida Savigny, 1818 China Wu et al. 1980:117 

Leocrates chinensis Kinberg, 1866 Hong Kong Mak 1982:599 


Leocrates claparedii (Costa in C1aparede, 1868) Singapore Tan & Chou 1993:282 

Ophiodromus berrisfordi Day, 1967 China Wu & Wu 1987:79 

Ophiodromus pugettensis (Johnson, 1901) Hong Kong Shin 1982:164 

Podarke latifrons (Grube 1878) Singapore Tan & Chou 1993:282 

Podarke cf. obscura (Verrill, 1837) China Yang & Sun, 1988:84 

Sinohesione genitaliphora Westheide et aI., 1994 Thailand Westheide et al. 1994:95 

Pilargidae 

Ancistargis sp. Taiwan Hsieh, pers. eomm. 

Loandalia indica Thomas, 1963 S. Vietnam Gallardo 1968:60 

Pilargis mohri Gallardo, 1968 S. Vietnam Gallardo 1968:60 

Pilargis verrucosa pacific Usehakov, 1955 China Yang & Sun 1988:77 

Sigambra cf. bassi Taiwan Hsieh, pers. eomm. 

Sigambra hanaokai (Kitamori, 1960) S. Vietnam Gallardo 1968:61 

Sigambra tentaculata (Treadwell, 1941) Hong Kong Shin 1982:164 

Synelmis albini (Langerhans, 1881) S. Vietnam Gallardo 1968:61 



Synelmis annamita Gallardo, 1968 S. Vietnam Gallardo 1968:62 

Syllidae 

Autolytus robustisetus Wu & Sun, 1978 China Sun et al. 1978:137 

Autolytus setoensis 1majima, 1966 China Yang & Sun 1988:55 

Autolytus sp. Taiwan Hsieh, pers. comm. 

Ehlersia cornuta (Rathke, 1843) Hong Kong Shin 1982: 164 


Ehlersia ef. rosea Imajima, 1966 China Yang & Sun 1988:66 

Eusyllis edenticulata Pillai, 1965 Philippines Pillai 1965:125 

Eusyllis ef. irregulata Imajima, 1966 China Yang & Sun, 1988:57 

214



Exogone verugera (Claparede, 1868) China Sun & Wu 1981:621 

Haplosyllis spongicola (Grube, 1855) China Yang & Sun 1988:67 

Haplosyllis spongicola tentaculata (Marion, 1879) China Yang & Sun 1988:67 

Langerhansia cornuta (Rathke, 1843) S. Vietnam Gallardo 1968:62 

Myrianida cf. pachycera (Augener, 1913) China Yang & Sun 1988:56 

Odontosyllis gibba Claparede, 1863 China Sun et al. 1978:140 

Odontosyllis rubrofasciata Grube, 1879 China Wu & Sun 1978:178 

Parautolytus luzonensis Pillai, 1965 Philippines Pillai 1965:123 

Sphaerosyllis hystrix Claparede, 1863 China Sun & Wu 1981:622 

Streptosyllis baolingi Ding & Westheide, 1994 China Ding & Westheide 1994:304 

Streptosyllis hainanensis Ding & Westheide, 1994 China Ding & Westheide 1994:308 

Syllis amica Quatrefages, 1865 China Yang & Sun 1988:68 

Syllis gracilis Grube, 1840 China Yang & Sun 1988:69 


Trypanosyllis taeniaformis (Haswell, 1886) Hong Kong Mak 1982:599 

China Yang & Sun 1988:71 as 

T. (Tryp. )taen. 

Trypanosyllis zebra (Grube, 1860) China Wu et al. 1980:118 

Typosyllis aciculata orientalis Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:63 


Typosyllis alternata (Moore, 1908) China Wu & Sun 1978:179 

Typosyllis armillaris (Muller, 1771) China Wu et al. 1980:118 

T)posyllis exilis Gravier, 1900 Philippines Natividad & Palpal-Latoc 1986:310 

Typosyllis maculata Imajima, 1966 China Yang & Sun 1988:74 

Typosyllis magnipectinis (Storch, 1967) China Licher et al. 1995: 107 

Typosyllis monilata Imajima, 1966 China Sun et al. 1978:141 

Typosyllis prolifera (Krohn, 1852) China Wu et al. 1980:118 

Typosyllis variegata (Grube, J860) S. Vietnam Gallardo 1968:63 

China Sun et al. 1978: 140 


Nereididae 

Ceratonereis anchylochaeta (Hors!, 1924) China Wu et al. 1981:Table 1 

Ceratonereis cf. burmensis Monro, 1937 China Sun et al. 1978: 148 

Ceratonereis costae (Grube, 1840) China Sun et al. 1978:146 


Ceratocephale fauveli (Pillai, 1961) S. Vietnam Gallardo 1968:63 

Ceratonereis erythraeensis Fauvel, 1918 China Wu et al. 1981:Table 1 

Ceratonereis hircinicola (Eisig, 1870) China Wu et al. 1981:Table I 

Ceratonereis ?hircinicola Singapore Tan & Chou 1993:283 

Ceratonereis japonica Imajima, 1972 China Sun et al. 1978:145 

Ceratonereis marmorata Horst, 1924 China Wu et al. 1981:Table 1 

Ceratonereis mirabilis Kinberg, 1866 China Sun et al. 1978:144 



Ceratonereis pachychaeta Fauvel,1918 China Sun et al. 1978:147 

Ceratonereis tripartita Horst, 1918 China Wu et al. 1980:118 

215



Ceratonereis sp. Taiwan Hsieh, pers. comm. 

Gymnonereis fauveli (Pillai, 1961) Thailand Hylleberg & Nateewathana 1988:8 

Dendronereis pinnaticirris Grube, 1878 Philippines Pillai 1965:127 

China Wu et al. 1981:Tb1e 1 

Taiwan Hsieh, pers. comm. 

Leonnates decipiens Fauve1, 1929 China Wu et al. 1981 :Tab1e 1 


Leonnates decipiens manilensis Pillai, 1965 Philippines Pillai 1965:144 

Leonnates jousseaumei Gravier, 1911 China Wu et al. 1981:Table 1 


Leonnates nipponicus 1majima, 1972 Hong Kong Mak 1982:599 

Leonnates persica Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:64 

China Wu et al. 1981 :Tab1e 1 


Leonnates sp. Taiwan Hsieh, pers. comm. 

Namalycastis aibiuma (Muller in Grube, 1881) China Wu et al. 1981:Tab1e 1 


Namalycastis rigida Pillai, 1965 Philippines Pillai 1965:133 

Neanthes arenaceodentata (Moore, 1903) S. Vietnam Gallardo 1968:65 

Neanthes bongcoi Pillai, 1965 Philippines Pillai 1965:142 

Neanthes diversicolor (Muller, 1776) Philippines Natividad & Pa1pa1-Latoc 1986:315 

Neanthes glandicincta (Southern, 1921) China Wu et al. 198I:Tab1e 1 


Neanthes maculata Wu et aI., 1981 China Wu et al. 1981:66 

Neanthes manatensis Pillai, 1965 Philippines Pillai 1965:139 

Neanthes nanhaiensis Wu et aI., 1981 China Wu et al. 1981:68 

Neanthes unifilSciata (Willey, 1905) China Wu et al. 1980:119 


Nectoneanthes ijimai (Izuka, 1912 ) China Wu et al. 1981:Tab1e 1 

Nectoneanthes multignatha Wu et aI., 1981 China Wu et al. 1981:73 

Nectoneanthes oxypoda (Marenzeller, 1879) China Wu et al. 1981:Tab1e I 

Nectoneanthes cf. oxypoda Hong Kong Shin 1982: 164 

Nereis coutieri Gravier, 1899 China Wu et al. 1981 :Table I 

Ne reis dayana Sun & Shen, 1978 China Sun et al. 1978:155 

Nereis denhamensis Augener, 1913 China Sun et al. 1978:151 

Nereis falcaria (Willey, 1905) China Wu et al. 1980:119 

Nereis guangdongensis Wu et aI., 1981 China Wu et al. 1981:75 

Nereis heteromorpha Horst, 1924 China Sun et al. 1978:157 

Nereis jacksoni Kinberg, 1866 China Wu et al. 1981:Tab1e I 

Nereis multignatha Imajima & Hartman, 1964 China Wu et al. 1981 :Tab1e 1 

Nereis nichollsi Kotl, 1951 China Wu et al. 1980:]]9 

Nereis pers;c'a Fauve1, 1911 China Wu et al. 1981:Table 1 


Nereis surugaense nanhaiensis Sun & Shen, 1978 China Sun et al. 1978:153 

Nereis trifasciata Grube, 1878 China Wu et al. 1980:119 

Nereis unifasciata Willey,1905 China Sun et al. 1978:154 

Nereis zhongshaensis Shen & Sun, 1978 China Sun et al. 1978:158 

216



Nereis zonata Malmgren, 1867 China Wu et al. 1980:119 

Nereis cf. zonata Taiwan Hsieh, pers. comm. 

Nicon japonica 1majima, 1972 China Wu & Sun 1979:104 

Nicon maculata Kinberg, 1866 S. Vietnam Gallardo 1968:65 as N. maculatus 

China Wu & Sun 1979:101 

Nicon sinica Wu & Sun, 1979 China Wu & Sun 1979:99 

Paraleonnates uschakovi Chlebovitsch & Wu, 1962 China Wu et al. 1981:Table I 

Perinereis aibuhitensis Grube, 1878 China Wu et al. 1981:Table 1 


Perinereis barbara Momo, 1926 Singapore Tan & Chou 1993:283 

Perinereis camiguinoides Augener, 1922 China Wu et al. 1981:Table 1 

Perinereis cavifrons Ehlers, 1920 China Wu et al. 1981:Table I 

Perinereis cultrifera cultrifera (Grube, 1840) China Wu et al. 1981:Table I 



Perinereis cultrifera jloridana Ehlers, 1868 China Wu et al. 1985:218 

Perinereis cultrifera helleri Grube, 1878 China Wu et al. 1985:217 

Perinereis cultrifera obfuscata Grube, 1878 China Wu et al. 1985:219 

Perinereis cultrifera perspicillata Grube, 1878 Singapore Tan & Chou 1993:283 

Perinereis cultrifera striolata Grube, 1878 Singapore Tan & Chou 1993:284 

Perinereis nuntia (Savigny, 1818) China Wu et al. 1981:Table 1 

Perinereis nuntia brevicirris (Grube, 1878) Singapore Tan & Chou 1993:284 


Perinereis nuntia majungaensis Fauvel, 1918 China Wu et al. 1985:213 

Perinereis nuntia vallata (Grube, 1857) China Wu et al. 1985:210 

Perinereis quatrefagesi (Grube, 1878) Thailand Hylleberg et al. 1986:5 

Perinereis rhombodonta Wu et aI., 1981 China Wu et al. 1981:83 

Perinereis singaporiensis (Grube, 1878) Singapore Grube 1878:84 

Perinereis suluana Horst, 1924 China Sun et al. 1978: 150 

Perinereis vancaurica (Ehlers, 1868) China Wu et al. 1981:Table 1 


Perinereis weijhouensis Wu et aI., 1981 China Wu et al. 1981:83 

Perinereis cf. weijhouensis Taiwan Hsieh, pers. comm. 

Platynereis abnormis (Horst, 1924) China Wu et al. 1980:119 

Platynereis bicanaliculata (Baird, 1863) China Wu et al. 1981:Table 1 

Platynereis dumerilli (Audouin & M. Edw., 1833) China Wu et al. 1980:119 

Platynereis pulchella Gravier, 1902 S. Vietnam Gallardo 1968:65 

China Wu et al. 1981:Table 1 

Platynereis sinica Sun & Shen, 1978 China Sun et al. 1978: 148 

Pseudonereis anomala Gravier, 1902 China Wu et al. 1980:119 

Pseudonereis gallapagensis Kinberg, 1866 China Wu et al. 198\ :Table 1 

Pseudonereis variegata (Grube, 1857) China Wu et al. 1981:Table 1 

Rullierinereis misakiensis (Imajima & Hayashi, 1972) China Wu & Sun 1979:106 

Sinonereis heteropoda Wu & Sun, 1979 China Wu & Sun 1979:96 

Tambalagamia fauveli Pillai, 1961 China Yang & Sun 1988:28 


Tylonereis bogoyawleskyi Fauvel, 1911 China Wu et al. 1981:Table 1 

217



Tylonereis heterochaeta Tan & Chou, 1994 Singapore Tan & Chou 1994:665 

Tylorrhynchus heterochaetus (Quatrefages, 1865) China Wu et al. 1981:Tab1e 1 

Nephtyidae 

Aglaophamus dibranchis (Gmbe, 1877) Singapore Tan & Chou 1993:284 

Aglaophamus dicirroides Fauehald, 1968 S. Vietnam Faueha1d 1968: 10 

Aglaophamus ef. jeffreysii (McIntosh, 1885) China Wu et al. 1980:120 

Aglaophamus lyrochaeta (Fauve1, 1902) Hong Kong Shin 1982: 165 

Aglaophamus munamaorii Gibbs, 1971 China Wu et al. 1980:120 

Aglaophamus orientalis Faueha1d, 1968 S. Vietnam Faucha1d 1968:12 

Aglaophamus sinensis (Fauve1, 1932) S. Vietnam Fauchald 1968:13 


Aglaophamus vietnamensis Faucha1d, 1968 S. Vietnam Faucha1d 1968:13 

Aglaophamus sp. Taiwan Hsieh, pers. comm. 

Inermonephtys gallardi Faueha1d, 1968 S. Vietnam Faucha1d 1968: 15 

Inermonephtys inermis (EhIers, 1887) S. Vietnam FauchaId 1968:16 


Micronephthys sphaerocirrata (Wesenberg-Lund, 1949) S. Vietnam Fauchald 1968: 18 


Micronephthys sp. Taiwan Hsieh, pers. comm. 

Nephtys cf. califomiensis Taiwan Hsieh, pers. comm. 

Nephtys oligobranchia Southern, 1921 S. Vietnam Faucha1d 1968: 18 

Nephtys polybranchia Southern, 1921 S. Vietnam FauchaId 1968:19 


Sphaerodoridae 

Sphaerodorum sp. S. Vietnam Gallardo 1968:67 

Glyceridae 

Glycera alba (Muller, 1788) S. Vietnam Gallardo 1968:68 



Glycera ?capitata Oersted, 1843 S. Vietnam Gallardo 1968:68 

Glycera chirori lzuka, 1912 China Yang & Sun 1988:87 

Glycera convoluta Keferstein, 1862 Singapore Tan & Chou 1993:284 

Glycera cf. convoluta Taiwan Hsieh, pers. comm. 

Glycera lancadivae Schmarda, 1861 China Sun et al. 1978:136 

Glycera nicobarica Grube, 1868 S. Vietnam Gallardo 1968:69 

Glycera onomichiensis Izuka, 1912 S. Vietnam Gallardo 1968:69 


Glycera posterobranchia Hoagland, 1920 Philippines Hoag1and 1920:620 

S. Vietnam Gallardo 1968:70 

Glycera parashadi Fauve1, 1932 China Wu et al. 1980:120 

Glycera rouxii Audouin & M. Edw., 1833 China Wu & Wu 1987:79 


Glycera cf. subaenea Taiwan Hsieh, pers. comm. 

Glyera cf. tenuis Taiwan Hsieh, pers. comm. 

Glycera tesselata Gmbe, 1863 S. Vietnam Gallardo 1968:70 


218



Goniadidae 

Glyeinde oligodon Southern, 1921 S. Vietnam Gallardo 1968:71 

Goniada annulata Moore, 1905 China Wu et al. 1980:119 

Goniada emerita Audouin & M. Edw., 1833 China Wu et al. 1980:119 

Goniada cf. emerita Taiwan Hsieh, pers. comm. 

Goniada cf. maeulata Oersted, 1843 China Wu et al. 1980:119 



Goniada multidentata Arwidsson, 1899 S. Vietnam Gallardo 1968:72 

Goniada paueidens Grube, 1878 Hong Kong Mak 1982:599 

Goniada tridens Gallardo, 1968 S. Vietnam Gallardo 1968:73 

Goniada uneinigera EhJers, 1901 S. Vietnam Gallardo 1968:73 

Goniada cf. uneinigera Taiwan Hsieh, pers. comm. 

Goniadopsis sp. Taiwan Hsieh, pers. comm. 

Dorvilleidae 

Dorvillea australiensis (McIntosh, 1885) China Wu et al. 1980:121 as Papilliod.aust. 

Sehistomeringos ineerta (Schmarda, 1861) S. Vietnam Gallardo 1968:91 as Dorvillea ine. 


Eunicidae 

Euniee afra Peters, 1854 China Wu & Wu 1987:79 

Euniee antennata (Savigny, 1820) Philippines Pi1lai 1965: 148 




Euniee aphroditois (Pallas, 1788) China Wu & Wu 1987:79 


Euniee ?australis Quatrefages, 1865 S. Vietnam Gallardo 1968:77 

Euniee australis Hong Kong Mak 1982:599 


Euniee eineta (Kinberg, 1865) Hong Kong Mak 1982:599 

Euniee eoeeinea Grube, 1878 China Wu et al. 1980:121 

Euniee eoeeinioides Augener, 1922 Singapore Tan & Chou 1993:285 

Euniee grubei Gravier, 1900 Singapore Tan & Chou 1993:286 

Euniee hirsehi Fauchald, 1992 Singapore Tan & Chou 1993:286 

Euniee indiea Kinberg, 1865 S. Vietnam Gallardo 1968:78 

Philippines Gallardo 1968:78 


Euniee lueei Grube, 1856 Singapore Tan & Chou 1993:286 

Euniee marenzelleri Gravier, 1900 China Wu et al. 1980:121 

Euniee medicina Moore, 1903 S. Vietnam Gallardo 1968:78 

Euniee nesiotes (Chamberlin, 1919) Singapore Tan & Chou 1993:286 

Euniphysa aeuleata Wesenberg-Lund, 1949 China Shen & Wu 1991:131 


Euniphysa oeulata Wu & Sun in Wu et aI., 1979 China Wu et al. 1979:89 

Euniphysa unieusa Shen & Wu 1991 China Shen & Wu 1991:131 

Heterophysa tridontesa Shen & Wu, 1991 China Shen & Wu 1991:138 

219



Lysidice collaris Grube, 1870 S. Vietnam Gallardo 1968:79 

Philippines Gallardo 1968:79 

China Sun et al. 1978: 162 as L.nin.collaris 



Lysidice ninetta Audouin & M. Edw., 1833 Hong Kong Mak 1982:599 


Marphysa adenensis Gravier, 1900 Hong Kong Mak 1982:600 

Marphysa bellii (Aud. & M. Edwards,1833) S. Vietnam Gallardo 1968:80 

Thailand Gallardo 1968:80 


Marphysa bifurcata Kott, 1951 S. Vietnam Gallardo 1968:80 

Marphysa depressa (Schmarda, 1861) China Yang & Sun 1988:186 

Marphysa disjuncta Hartman, 1961 Singapore Tan & Chou 1993:286 

Marphysa gravelyi Southern, 1921 Philippines Pillai 1965: 150 

Marphysa macintoshi Crossland, 1903 China Wu et al. 1980:121 


Marphysa mossambica (Peters, 1854) China Wu & Wu 1987:79 


Marphysa sanguinea (Montagu, 1815) Hong Kong Shin 1982: 166 


Marphysa stragulum (Grube, 1878) China Wu et al. 1980:121 

Nauphanta sp. Tan & Chou, in press Singapore Tan & Chou, in press 

Nematonereis unicornis (Grube, 1840) S. Vietnam Gallardo 1968:81 


Palola siciliensis (Grube, 1840) China Wu et al. 1980:121 


Paraeuniphysa falciseta Shen & Wu, 1991 China Shen & Wu 1991:134 

Paraeuniphysa spinea (Miura, 1977) China Shen & Wu 1991:136 

Paraeuniphysa taiwanensis Wu & He, 1988 Taiwan Shen & Wu 1991:134 

Paramarphysa sp. Hong Kong Mak 1982:600 

Lumbrineridae 

Lumbrineris acutiformis Gallardo, 1968 S. Vietnam Gallardo 1968:82 

Lumbrineris ? amboinensis (Grube, 1877) S. Vietnam Gallardo 1968:82 

Lumbrineris bifurcata (Mclntosh, 1885) China Wu et al. 1980:121 

Lumbrineris caudaensis Gallardo, 1968 S. Vietnam Gallardo 1968:83 

Lumbrineris cf. cruzensis Hartman, 1944 China Wu et al. 1980:121 


Lumbrineris hartmani (Day, 1953) Singapore Tan & Chou 1993:287 

Lumbrineris heteropoda (Marenzeller, 1879) S. Vietnam Gal1ardo 1968:84 


Lumbrineris inflata Moore, 1911 Hong Kong Mak 1982:600 

Lumbrineris japonica (Marenzeller, 1879) S. Vietnam Gallardo 1968:84 

Lumbrineris latreilli Audouin & M. Edw., 1834 China Wu et al. 1980:121 


Lumbrineris mucronata (Ehlers, 1908) S. Vietnam Gallardo 1968:85 

Lumbrineris nagae Gallardo, 1968 S. Vietnam Gallardo 1968:85 


220



Lumbrineris ?notocirrata (Fauvel, 1932) S. Vietnam Gallardo 1968:86 

Lumbrineris pterignatha Gallardo, 1968 S. Vietnam Gallardo 1968:86 

Lumbrineris shiinoi Gallardo, 1968 S. Vietnam Gallardo 1968:87 


Lumbrineris simplex Southern, 1921 China Wuetal.1980:121 

Lumbrineris sphaerocephala (Sehmarda, 1861) China Wu et al. 1980: 121 

Ninoe bruuni Gallardo, 1968 S. Vietnam Gallardo 1968:88 

Ninoe mando (Crossland, 1924) S. Vietnam Gallardo 1968:88 

Oenonidae 

Arabella iricolor (Montague, 1804) Singapore Tan & Chou 1993:287 

Arabella mu tans (Chamberlin, 1919) China Wu & Wu 1987:79 

Arabella novecrinita Crossland, 1924 S. Vietnam Gallardo 1968:89 

Drilonereis logani Crossland, 1924 S. Vietnam Gallardo 1968:89 


Drilonereis ef. logani Taiwan Hsieh, pers. eomm. 

Drilonereis ?paucidentata Treadwell, 1922 S. Vietnam Gallardo 1968:90 

Notocirrus biaciculus Sehmarda, 1861 S. Vietnam Gallardo 1968:90 

Notocirrus japonicus (Okuda, 1939) S. Vietnam Gallardo 1968:91 

Oenone julgida (Savigny, 1818) Hong Kong Mak 1982:600 


Onuphidae 

Diopatra bulohensis Tan & Chou, 1996 Singapore Tan & Chou 1996:359 

Diopatra neapolitana Delle Chiaje, 1841 Singapore Tan & Chou 1993:285 

Diopatra sugokai lzuka, 1907 China Yang & Sun 1988:175 

as D. chiliensis 

Taiwan Hsieh & Chang 1991 :331 

as D. bilobata 

Diopatra variabilis Southern, 1921 S. Vietnam Gallardo 1968:74 



Hyalinoecia tubicola (MUller, 1788) China Wu et al. 1980: 120 

Onuphis punggolensis Tan & Chou, in press Singapore Tan & Chou, in press 

Onuphis sp. Taiwan Hsieh, pers. eomm. 

Nothria conchylega (Sars, 1835) S. Vietnam Gallardo 1968:75 

Nothria holobranchiata (Marenzeller, 1879) S. Vietnam Gallardo 1968:75 

Nothria nr. atlantisa Hartman, 1965 S. Vietnam Gallardo 1968:75 

Onuphis eremita Audouin & M. Edw. 1833 S. Vietnam Gallardo 1968:76 


Onuphis pseudodibranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:76 


Onuphis variolata Shen, 1987 China Shen 1987:222 

Paradiopatra willemoesii (Mclntosh, 1885) China Wu et al. 1980:120 

Orbiniidae 

Haploscoloplos elongatus (Johnson, 1901) S. Vietnam Gallardo 1968:92 


Haploscoloplos ef. jragilis (Verrill, 1873) S. Vietnam Gallardo 1968:92 

221



Leitoscoloplos elongatus Taiwan Hsieh, pers. eomm. 

Leitoscoloplos ef. kerguelensis (McIntosh, 1885) Hong Kong Shin 1982:167 

Naineris hainanensis Q. WU China Wu & Wu 1987:79 

Orbinia ef. dicrochaeta Taiwan Hsieh, pers. eomm. 

Orbinia vietnamensis Gallardo, 1968 S. Vietnam Gallardo 1968:93 


Phylo kupfferi (Eh1ers, 1875) Hong Kong Shin 1982: 167 

Phylo ornatus (Verrill, 1873) China Yang & Sun 1988:206 

Scoloplos cf. acmeceps Taiwan Hsieh, pers. eomm. 

Scoloplos ef. armiger (Muller, 1776) China Wu et al. 1980:122 

Scoloplos (Leodamas) ?dubia Tebble, 1955 S. Vietnam Gallardo 1968:94 

Scoloplos (L.) gracilis Pillai, 1961 S. Vietnam Gallardo 1968:94 

Scoloplos (L.) rubra (Webster, 1879) China Yang & Sun 1988:208 

Scoloplos (L.) rubra orientalis Gallardo, 1968 S. Vietnam Gallardo 1968:95 

Scoloplos cf. rubra Taiwan Hsieh, pers. eomm. 

Scoloplos (Scoloplos) marsupialis Southern, 1921 China Yang & Sun 1988:209 

Scoloplos (Scoloplos) spiniferus Gallardo, 1968 S. Vietnam Gallardo 1968:95 

Scoloplos tumidus Maekie, 1991 Hong Kong Maekie 1991:38 

S. Vietnam Gallardo 1968:95 as S. marsupialis 

Scoloplos cf. tumidus Taiwan Hsieh, pers. eomm. 

Paraonidae 

Aricidea cf. fragilis caeca Taiwan Hsieh, pers. eomm. 

Paradoneis lyra (Southern, 1914) S. Vietnam Gallardo 1968:96 

Spionidae 

Aonides sp. China Wu & Sun 1978:187 

Aonides ef. oxycephala Taiwan Hsieh, pers. eomm. 

Dispio ef. uncinata Taiwan Hsieh, pers. eomm. 

Laonice cirrata (M. Sars, 1851) S. Vietnam Gallardo 1968:97 

China Wu & Sun 1978:192 


Malacoceros (Rh.) areninc.asiaticus (Ch1ebovitseh, 1959 ) China Wu & Sun 1978:189 

Malacoceros indicus (Fauve1, 1928) China Wu et al. 1980:122 

Taiwan Hsieh, pers. eomm. 

Microspio sp. China Wu & Sun 1978:191 

Minuspio cirrifera (Wiren, 1883) Hong Kong Shin 1982:167 

Nerine faughni Gallardo, 1968 S. Vietnam Gallardo 1968:97 

Paraprionospio pinnata (Ehlers, 1901) Hong Kong Shin 1982: 167 


Paraprionospio sp. Taiwan Hsieh, pers. eomm. 

Polydorella novaegeorgiae Gibbs, 1971 Hong Kong Shin 1982: 168 

Polydora cf. pilikia Ward, 1981 China Yang & Sun 1988:213 

Polydora cf. socialis (Sehmarda, 1861) Hong Kong Shin 1982: 167 

Polydora ef. tentaculata Blake & Kudenov,1978 Hong Kong Shin 1982: 168 


Polydora cavitensis Pillai, 1965 Philippines Pillai 1965:152 

222



Polydora glycymerica Radashevsky, 1993 Manchenko & Radashevsky 

1994:767 

Polydora ligni Webster, 1879 Taiwan Hsieh, pers. comm. 

Polydora vulgaris Mohammad, 1972 Manchenko & Radashevsky 

1994:767 

Prionmpio africana Augener, 1918 S. Vietnam Gallardo 1968:98 

Prionospio depauperata Taiwan Hsieh, pers. comm. 

Prionospio ehlersi Fauvel, 1928 S. Vietnam Gallardo 1968:98 

China Wu et al. 1980: 122 


Prionospio japonica Taiwan Hsieh, pers. comm. 

Prionospio komaeti Hylleberg & Nateew., 1991 Singapore Tan & Chou 1993:288 

Prionospio malmgreni Claparede, 1870 China Wu & Sun 1978:189 


Prionospio malayensis Caullery, 1914 S. Vietnam Gallardo 1968:99 


Prionospio paradisea Taiwan Hsieh, pers. comm. 

Prionospio pinnata Eh1ers, 1901 S. Vietnam Gallardo 1968:99 

Prionospio cf. pulchra Taiwan Hsieh, pers. comm. 

Prionospio saccifera Mackie & H. 1990 Hong Kong Mackie & Hart1ey 1990:366 

Prionospio sishaensis Wu & Chen, 1964 China Wu & Wu 1987:79 

Pseudopolydora antennata Claparede, 1870 China Wu & Sun 1978:183 

Pseudopolydora diopatra Hsieh, 1992 Taiwan Hsieh 1992:630 

Pseudopolydora cf. kempi Taiwan Hsieh, pers. comm. 

Pygospio elegans C1aparede, 1863 China Wu & Sun 1978:186 

Scolelepis ? indica Fauvel, 1928 S. Vietnam Gallardo 1968:100 

Scolelepis sp. Taiwan Hsieh, pers. comm. 

Spiophanes bombyx (C1aparede, 1870) China Wu & Sun 1978:185 

Spiophanes malayensis Caullery, 1915 China Wu et al. 1980:122 

Spiophanes sp. Taiwan Hsieh, pers. comm. 

Heterospionidae 

Heterospio sinica Wu & Chen, 1966 China Yang & Sun 1988:221 

Magelonidae 

Magelona cf. cincta Taiwan Hsieh, pers. comm. 

Magelona crenulifrons Gallardo, 1968 S. Vietnam Gallardo 1968: 100 


Magelona japonica Okuda, 1937 S. Vietnam Gallardo 1968:101 

China Wu et a1 1980:122 

Magelone koreana Okuda, 1937 S. Vietnam Gallardo 1968:101 

Magelona lenticulata Gallardo, 1968 S. Vietnam Gallardo 1968: 101 

Magelona cf. pacifica Taiwan Hsieh, pers. comm. 

Magelona sp. China Wu & Sun 1978:195 

Cossuridae 

Cossura coasta Kitamori, 1960 Hong Kong Shin 1982: 169 

Cossurella cf. diamorpha Taiwan Hsieh, pers. comm. 

223



Trochochaetidae 

Trochochaeta diverapoda (Hoagland, 1920) Hong Kong Mackie 1990:338 

Trochochaeta cf. orissae (Fauvel, 1932) Hong Kong Shin 1982:168 

Poecilochaetidae 

Poecilochaetus gallardoi Pilato & Cantone, 1976 S. Vietnam Pilato & Cantone 1976:43 

S. Vietnam Gallardo 1968: 103 as P. sp. A 

Poecilochaetus hystricosus Mackie, 1990 Hong Kong Mackie 1990: 351 

Poecilochaetus paratropicus Gallardo, 1968 S. Vietnam Gallardo 1968: 102 

Poecilochaetus serpens Alien, 1904 China Wu et aI 1980:122 


Poecilochaetus spinulosus Mackie, 1990 Hong Kong Mackie 1990:342 

Poecilochaetus tricirratus Mackie, 1990 Hong Kong Mackie 1990:346 

Poecilochaetus tropicus Okuda, 1937 China Wu & Sun 1978:180 

Poecilochaetus vietnamita Gallardo, 1968 S. Vietnam Gallardo 1968: 103 

Poecilochaetus sp. Taiwan Hsieh, pers. comm. 

Chaetopteridae 

Chaetopterus variopedatus (Renier, 1804) Singapore Tan & Chou 1993:288 


Sasekumaria se/angora Rullier, 1976 Malaysia Ng & Sasekumar 1993:467 

Spiochaetopterus costarum (Claparecte, 1868) Hong Kong Shin 1982: 168 

costarum 

Cirratulidae 

Caulleriella glabra Gallardo, 1968 S. Vietnam Gallardo 1968: 104 

Caulleriella cf. typhlops (Willey, 1905) China Sun et al 1978: 163 

Chaetozone flagellifera Gallardo, 1968 S. Vietnam Gallardo 1968: 105 

Chaetozone maotienae Gallardo, 1968 S. Vietnam Gallardo 1968:105 

Cirratulus annamensis Gallardo, 1968 S. Vietnam Gallardo 1968: 105 

Cirratulus cirratulus Muller, 1776 Singapore Tan & Chou 1993:288 

Cirratulus filiformis Keferstein, 1862 China Yang & Sun 1988:234 

Cirriformia chrysodermoides Pill ai, 1965 Philippines Pillai 1965:156 

Cirriformia cf. filiformis Taiwan Hsieh, pers. comm. 

Cirriformia jiligera (Delle Chiaje, 1828) China Wu 1987:79 

Cirriformia punctata (Grube, 1859) China Wu et al. 1980:123 

Cirriformia semicincta (Ehlers, 1905) S. Vietnam Gallardo 1968: 106 

Cirriformia tentaculata (Montagu, 1808) China Yang & Sun 1988:235 

Dodecaceria jewkesi Berkeley & Berk., 1954 China Yang & Sun 1988:236 

Tharyx filibranchia Day, 1961 Hong Kong Shin 1982: 169 

Tharyx marioni (Saint-Joseph, 1894) S. Vietnam Gallardo 1968:1 06 

Tharyx sp. Taiwan Hsieh, pers. comm. 

Flabelligeridae 

Brada jerruginea Gallardo, 1968 S. Vietnam Gallardo 1968:107 

Diplocirrus erythroporus Gallardo, 1968 S. Vietnam Gallardo 1968: 108 

Flabelliderma sp. Taiwan Hsieh, pers. comm. 

224



Pherusa parmata (Grube, 1877) Hong Kong Mak 1982:600 


Pherusa curvisetis (Caullery, 1944) Philippines Gallardo 1968:108 

S. Vietnam Gallardo 1968: 108 

Pherusa eruca indica (Fauvel, 1928) S. Vietnam Gallardo 1968:109 

Pherusa monroi (Day, 1957) Singapore Tan & Chou 1993:289 

Piromis congoensis (Grube, 1877) S. Vietnam Gallardo 1968:109 

Scalibregmidae 

Scalibregma inflatum Rathke, 1843 S. Vietnam Gallardo 1968:110 


Opheliidae 

Ammotrypane aulogaster Rathke, 1843 Singapore Tan & Chou 1993:289 

Ammotrypane ?dubia Caullery, 1944 S. Vietnam Gallardo 1968: 111 

Ammotrypane ?ehlersi Horst, 1919 S. Vietnam Gallardo 1968:111 

Ammotrypane ?fauveli Caullery, 1944 S. Vietnam Gallardo 1968:111 

Ammotrypane filobranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:112 

Ammotrypane grandis Pillai, 1961 S. Vietnam Gallardo 1968:112 

Armandia ?intermedia Fauvel, 1902 S. Vietnam Gallardo 1968: 113 

Armandia intermedia China Yang & Sun 1988:246 

Armandia lanceolata Willey, 1905 S. Vietnam Gallardo 1968: 113 

Armandia leptocirris (Grube, 1878) Philippines Grube 1878:194 

S. Vietnam Gallardo 1968: 113 


Armandia ef. leptocirris Taiwan Hsieh, pers. eomm. 

Armandia longicaudata (Caullery, 1944) S. Vietnam Gallardo 1968:114 

Ophelina acuminata Oersted, 1843 China Yang & Sun 1988:248 

Ophelina ef. acuminata Taiwan Hsieh, pers. eomm. 

Polyophthalmus pictus (Dujardin, 1839) Philippines Pillai 1965:157 

China Sun et al. 1978: 163 

Sternaspidae 

Sternaspis scutata (Renier, 1807) S. Vietnam Gallardo 1968:114 




Capitellidae 

Capitella capitata (Fabrieius, 1780) Singapore Tan & Chou 1993:289 

Capitella sp. Taiwan Hsieh, pers. eomm. 

Capitellides sp. Taiwan Hsieh, pers. eomm. 

Capitellethus branchiferus Gallardo, 1968 S. Vietnam Gallardo 1968:115 

Dasybranchus caducus (Grube, 1846) S. Vietnam Gallardo 1968:116 


Heteromastus filiformis (Claparede, 1864) China Yang & Sun 1988:254 

Heteromastus similis Southern, 1921 S. Vietnam Gallardo 1968: 117 

Mastobranchus loii Gallardo, 1968 S. Vietnam Gallardo 1968:118 

Mediomastus sp. Taiwan Hsieh, pers. eomm. 

225



Branchiomma cf. boholense (Grube, 1878) Taiwan Hsieh, pers. comm. 

Branchiomma cingulata (Grube, 1870) Philippines Pillai 1965:163 



Branchiomma cf. conspersa Ehlers, 1887) Taiwan Hsieh, pers. comm. 

Chone ?infundibuliformis Krayer, 1856 S. Vietnam Gallardo 1968:130 

Chone trilobata Gallardo, 1968 S. Vietnam Gallardo 1968:130 

Hypsicomus phaeotaenia (Schmarda, 1861) China Wu et al. 1980:124 


lasmineira caducibranchiata Willey, 1905 S. Vietnam Gallardo 1968:131 

Laonome albicingillum Hsieh, 1995 Taiwan Hsieh 1995:131 

Megalomma intermedium (Beddard, 1889) Philippines Pillai 1965: 164 

Megalomma vesiculosa (Montagu, 1815) Hong Kong Mak 1982:600 


Megalomma sp. Taiwan Hsieh, pers. comm. 

Myxicola infundibulum (Renier, 1804) Hong Kong Mak 1982:600 

Notaulax cL pyrroghaster Taiwan Hsieh, pers. comm. 

Potamilla leptochaeta Southern, 1921 Singapore Tan & Chou 1993:290 

Potamilla ocalea Pillai, 1965 Philippines Pillai 1965: 164 

Potamilla torelli Ma1mgren, 1866 Hong Kong Shin 1982: 170 

Pseudobranchiomma sp. Taiwan Hsieh, pers. comm. 

Pseudopotamilla reniformis (Muller, 1771) Hong Kong Mak 1982:600 

Singapore Tan & Chou 1993;600 

Pseudopotamilla sp. Taiwan Hsieh, pers. comm. 

Sabella penicillus Linneus, 1767 China Wu et al. 1980:124 

Sabella sp. Hong Kong Mak 1982:600 

Sabellastarte indica (Savigny, 1818) S. Vietnam Gallardo 1968:131 




Sabellastarte cf. samoensis (Grube, 1870) Taiwan Hsieh, pers. comm. 

Sabellastarte sanctijosephi (Gravier, 1906) China Wu et a1 1980:124 

Serpulidae 

Apomatus enosimae Marenzeller, 1884 China Yang & Sun 1988:308 

Circeis spirillum (Linnaeus, 1758) China Yang & Sun 1988:318 

Dexiospira foraminosus Busch, 1904 China Yang & Sun 1988:319 

Dexiospira sp. Taiwan Cheng & Chen 1994: 

Hydroides albiceps Grube, 1869 Hong Kong Mak 1982:601 


Hydroides centrospina Wu & Chen, 1981 China Wu & Chen 1981:354 

Hydroides dirampha March, 1863 Hong Kong Mak 1982:602 

Hydroides elegans (Haswell, 1883) Hong Kong Mak 1982:602 


Hydroides cf. elegans Taiwan Hsieh, pers. comm. 

Hydroides exaltata (Marenzeller, 1884) China Wu et al. 1980: 124 


Hydroides fusca Imajima, 1976 China Wu et al. 1980:124 

228



Hydroides cf. jusicola March, 1863 Hong Kong Mak 1982:603 

Hydroides grubei Pillai, 1965 Philippines Pillai 1965:166 

Hydroides inornata Pillai, 1960 Hong Kong Mak 1982:605 

Hydroides longistylaris Chen & Wu, 1980 China Chen & Wu 1980:249 

Hydroides nanhaiensis Wu & Chen, 1981 China Wu & Chen 1981:355 

Hydroides norvegica Gunnereus, 1768 Philippines Pillai 1965: 168 

Hydroides rhombobulus Chen & Wu,1980 China Chen & Wu 1980:247 


Hydroides trilobulus Wu & Chen, China Wu et al. 1980:125 

Metavermilia acanthophora (Augener, 1914) China Wu et al. 1980:125 


Metavermilia annobonensis Zibrowius, 1973 China Wu et al. 1980:125 

Metavermilia cf. inflata 1majima, 1977 Hong Kong Mak 1982:607 

Metavermilia cf. taenia Zibrowius, 1973 China Wu et al. 1980:125 

Neodexiospira cf. joraminosa Taiwan Hsieh, pers. comm. 

Neodexiospira cf. preacuta (Bush, 1904) Taiwan Hsieh, pers. comm. 

Neodexiospira cf. knightjonesi Taiwan Hsieh, pers. comm. 

Neodexiospira cf. steueri (Sterzinger, 1909) Taiwan Hsieh, pers. comm. 

Neopomatus uschakovi Pillai, 1965 Philippines Pillai 1965:170 

lingayanensis 

Pileolaria cf. militaris C1aparecte, 1870 Taiwan Hsieh, pers. comm. 

Pileolaria cf. pettibonei Taiwan Hsieh, pers. comm. 

Pileospira koehleri (Caullery & Mesni1, 1897) China Wu et al. 1980: 125 

Pomatoleios kraussii (Baird, 1865) Hong Kong Mak 1982:607 

Thailand Becker 1993 

Pomatoleios kraussii manilensis Pillai, 1965 Philippines Pillai 1965:169 

Pomatostegus stellatus (Abi1dgaard, 1789) Hong Kong Mak 1982:608 

Protohydroides elegans (Haswell, 1883) Hong Kong Shin 1982: 170 

Protula tubularia tubularia (Montagu, 1803) China Wu et al. 1980:125 

Protula sp. Taiwan Hsieh, pers. comm. 

Salmacina cf. dysteri (Hyx1ey, 1955) Taiwan Hsieh, pers. comm. 

Serpula (Paraserpula) sinica Wu & Chen in Wu et al.,1979 China Wu et al. 1979:92 

Serpula vermicularis Linnaeus, 1767 Hong Kong Mak 1982:608 


Serpula watsoni Willey, 1905 Hong Kong Mak 1982:609 

Simplicaria pseudomilitaris (Thiriot-Quievreux, 1965) Taiwan Hsieh, pers. comm. 

Spirobranchus giganteus (Pallas, 1766) China Wu et al. 1980: 125 

Spirobranch. giganteus corniculatus ten Hove Taiwan Dai & Yang 1995: 

Spirobranchus jousseaumei (Gravier, 1908) China Yang & Sun 1988:314 

Spirobranchus latiscapus (Marenzeller, 1884) China Wu et al. 1980:125 

Spirobranchus maldivensis Pixell, 1913 China Yang & Sun 1988:315 

Spirobranchus semperi March, 1863 Hong Kong Mak 1982:609 


Spirobranchus tricornis (March, 1863) Hong Kong Mak 1982:609 


Spirorbis (Dexiospira) treadwelli Pillai, 1965 Philippines Pillai 1965: 172 

Vermiliopsis glandigerous Gravier, 1908 China Wu et al. 1980:125 

229



Vermiliopsis infundibulum (Linneus, 1788) China Wu et al. 1980:125 

Vermiliopsis infundibul. glandigera Imajima, 1967 Hong Kong Mak 1982:610 

Vermiliopsis pygidialis (Willey, 1905) China Wu et al. 1980:125 

Nerillidae 

Leptonerilla diplocirrata Westheide & Purschke 1996 China 

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