Download the Nearctic Scarabaeoidea checklist PDF file

Checklist and Nomenclatural Authority File of 

the Scarabaeoidea of the Nearctic Realm 

including Canada, the continental United States, and the northern Mexican states of 

Baja California, Baja California Sur, Chihuahua, Coahuila de Zaragoza, Durango, 

Nuevo Leon, Sinaloa, Sonora, Tamaulipas, and Zacatecas. 

SCARABAEOIDEA Latreille, 1802 

PLEOCOMIDAE LeConte, 1861 

Pleocoma LeConte, 1856 

Version 4 - released 22 April 2009 

Andrew B. T. Smith 

Canadian Museum of Nature 

P.O. Box 3443, Station D 

Ottawa, ON, K1P 6P4 

CANADA 

asmith@mus-nature.ca 

Pleocoma australis Fall, 1911 (Pleocoma) 

Pleocoma badia badia Fall, 1917 (Pleocoma) 

Pleocoma badia hirsuta Davis, 1934 (Pleocoma) 

Pleocoma nitida Linsley, 1941 

Pleocoma venturae Linsley, 1945 

Pleocoma behrensii LeConte, 1874 (Pleocoma) 

Pleocoma bicolor Linsley, 1935 (Pleocoma) 

Pleocoma blaisdelli Linsley, 1938 (Pleocoma) 

Pleocoma carinata Linsley, 1938 (Pleocoma) 

Pleocoma conjungens conjungens Horn, 1888 (Pleocoma) 

Pleocoma conjungens lucia Linsley, 1941 (Pleocoma) 

Pleocoma crinita Linsley, 1938 (Pleocoma) 

Pleocoma dubitabilis dubitabilis Davis, 1935 (Pleocoma) 

Pleocoma dubitabilis leachi Linsley, 1938 (Pleocoma) 

Pleocoma fimbriata LeConte, 1856 (Pleocoma) 

Pleocoma hirticollis hirticollis Schaufuss, 1870 (Pleocoma) 

Pleocoma hirticollis reflexa Hovore, 1972 (Pleocoma) 

Pleocoma hirticollis vandykei Linsley, 1938 (Pleocoma) 

Pleocoma hoppingi Fall, 1906 (Pleocoma) 

Pleocoma hovorei La Rue, 2007 (Pleocoma) 

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Pleocoma linsleyi Hovore, 1971 (Pleocoma) 

Pleocoma marquai Hovore, 1972 (Pleocoma) 

Pleocoma minor Linsley, 1938 (Pleocoma) 

Pleocoma octopagina Robertson, 1970 (Pleocoma) 

Pleocoma oregonensis Leach, 1933 (Pleocoma) 

Pleocoma remota Davis, 1934 

Pleocoma puncticollis Rivers, 1889 (Pleocoma) 

Pleocoma rickseckeri Horn, 1888 (Pleocoma) 

Pleocoma rubiginosa rubiginosa Hovore, 1972 (Pleocoma) 

Pleocoma rubiginosa transsierrae Hovore, 1977 (Pleocoma) 

Pleocoma shastensis Van Dyke, 1933 (Pleocoma) 

Pleocoma simi Davis, 1934 (Pleocoma) 

Pleocoma sonomae Linsley, 1935 (Pleocoma) 

Pleocoma staff Schaufuss, 1870 (Pleocoma) 

Pleocoma adjuvans Crotch, 1873 

Pleocoma edwardsii LeConte, 1874 

Pleocoma ulkei Horn, 1888 

Pleocoma trifoliata Linsley, 1938 (Pleocoma) 

Pleocoma tularensis Leach, 1933 (Pleocoma) 

GEOTRUPIDAE Latreille, 1802 

BOLBOCERATINAE Mulsant, 1842 

ATHYREINI Howden and Martínez, 1963 

Neoathyreus Howden and Martínez, 1963 

Neoathyreus fissicornis (Harold, 1880) (Athyreus) 

Neoathyreus mixtus Howden, 1964 (Neoathyreus) 

BOLBOCERATINI Mulsant, 1842 

Bolbelasmus Boucomont, 1911 

Kolbeus Boucomont, 1911 

Bolbelasmus arcuatus (Bates, 1887) (Bolboceras) 

Bolbelasmus bajaensis Howden, 1964 (Bolbelasmus) 

Bolbelasmus hornii (Rivers, 1886) (Bradycinetus) 

Bolbelasmus minor (Linell, 1896) (Bradycinetus) 

Bolbelasmus variabilis Howden, 1964 (Bolbelasmus) 

Bolbocerastes Cartwright, 1953 

Bolbocerastes imperialis imperialis Cartwright, 1953 (Bolbocerastes) 

Bolbocerastes imperialis kansanus Cartwright, 1953 (Bolbocerastes) 

Bolbocerastes peninsularis Schaeffer, 1906 (Bradycinetus) 

Bolbocerastes regalis Cartwright, 1953 (Bolbocerastes) 

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Bolbocerastes serratus (LeConte, 1854) (Athyreus) 

Bolbocerosoma Schaeffer, 1906 

Bolbocerosoma (Bolbocerosoma) Schaeffer, 1906 

Bolbocerosoma (Bolbocerosoma) biplagiatum Dawson and McColloch, 1924 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) bruneri Dawson and McColloch, 1924 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) cartwrighti Howden, 1955 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) confusum Brown, 1928 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) elongatum Howden, 1955 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) farctum (Fabricius, 1775) (Scarabaeus) 

Scarabaeus cephus Fabricius, 1775 

Bolbocerosoma (Bolbocerosoma) hamatum Brown, 1929 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) lepidissimum Brown, 1928 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) mexicanum Howden, 2005 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) pusillum pusillum Dawson and McColloch, 1924 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) pusillum townesi Howden, 1955 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) quadricornum Robinson, 1941 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) ritcheri Howden, 1955 (Bolbocerosoma) 

Bolbocerosoma (Bolbocerosoma) tumefactum (Palisot de Beauvois, 1809) (Scarabaeus) 

Bolborhombus Cartwright, 1953 

Bolborhombus angulus (Robinson, 1947) (Bolboceras) 

Bolborhombus magnus Howden, 1964 (Bolborhombus) 

Bolborhombus parvulus Cartwright, 1953 (Bolborhombus) 

Bolborhombus sallaei (Bates, 1887) (Bolboceras) 

Bradycinetus carinatus Schaeffer, 1906 

Bolboceras schaefferi Boucomont, 1911 

Bradycinetulus Cockerell, 1906 

Amechanus Horn, 1870 

Bradycinetus Horn, 1871 

Bradycinetulus ferrugineus (Palisot de Beauvois, 1809) (Scarabaeus) 

Bolboceras furcicollis Laporte, 1840 

Bradycinetulus fossatus (Haldeman, 1853) (Bolboceras) 

Bradycinetulus rex Cartwright, 1953 (Bradycinetulus) 

Eucanthus Westwood, 1848 

Eucanthus alutaceus Cartwright, 1944 (Eucanthus) 

Eucanthus greeni Robinson, 1948 (Eucanthus) 

Eucanthus impressus Howden, 1964 (Eucanthus) 

Eucanthus lazarus (Fabricius, 1775) (Scarabaeus) 

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Scarabaeus meliboeus Fabricius, 1792 

Eucanthus mexicanus Howden, 1964 (Eucanthus) 

Eucanthus subtropicus Howden, 1955 (Eucanthus) 

Odonteus Samouelle, 1819 

Odonteus alabamensis Wallis, 1929 (Odonteus) 

Odonteus cornigerus (Melsheimer, 1845) (Bolboceras) 

Odonteus darlingtoni Wallis, 1928 (Odonteus) 

Odonteus falli Wallis, 1928 (Odonteus) 

Odonteus filicornis (Say, 1823) (Geotrupes) 

Odonteus floridensis Wallis, 1928 (Odonteus) 

Odonteus liebecki Wallis, 1928 (Odonteus) 

Odonteus obesus LeConte, 1859 (Odonteus) 

Odonteus simi Wallis, 1928 (Odonteus) 

Odonteus thoracicornis Wallis, 1928 (Odonteus) 

GEOTRUPINAE Latreille, 1802 

Ceratophyus Fischer von Waldheim, 1823 

Ceratophyus gopherinus Cartwright, 1966 (Ceratophyus) 

Ceratotrupes Jekel, 1865 

Ceratotrupes bolivari Halffter and Martínez, 1962 (Ceratotrupes) 

Geotrupes Latreille, 1797 

Geotrupes (Anoplotrupes) Jekel, 1865 

Melanotrupes Blanchard, 1888 

Geotrupes (Cnemotrupes) Jekel, 1865 

Onychotrupes Jekel, 1865 

Geotrupes (Geohowdenius) Zunino, 1984 

Geotrupes (Geotrupes) Latreille, 1797 

Geotrupes (Megatrupes) Zunino, 1984 

Geotrupes (Anoplotrupes) balyi Jekel, 1865 (Geotrupes) 

Geotrupes similis Jekel, 1865 

Geotrupes (Cnemotrupes) blackburnii blackburnii (Fabricius, 1781) (Scarabaeus) 

Geotrupes conicollis Jekel, 1865 

Geotrupes jekelii Horn, 1868 

Geotrupes (Cnemotrupes) blackburnii excrementi Say, 1823 (Geotrupes) 

Geotrupes (Megatrupes) cavicollis Bates, 1887 (Geotrupes) 

Geotrupes (Geohowdenius) egeriei Germar, 1824 (Geotrupes) 

Geotrupes lecontei Jekel, 1865 

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Geotrupes (Anoplotrupes) hornii Blanchard, 1888 (Geotrupes) 

Geotrupes (Geohowdenius) opacus Haldeman, 1853 (Geotrupes) 

Geotrupes haldemani Jekel, 1865 

Geotrupes chevrolati Jekel, 1865 

Geotrupes (Cnemotrupes) semiopacus Jekel, 1865 (Geotrupes) 

Geotrupes melsheimeri Jekel, 1865 

Geotrupes ovalipennis Jekel, 1865 

Geotrupes (Cnemotrupes) splendidus miarophagus Say, 1823 (Geotrupes) 

Geotrupes (Cnemotrupes) splendidus splendidus (Fabricius, 1775) (Scarabaeus) 

Geotrupes gilnickii Jekel, 1865 

Geotrupes starki Jekel, 1865 

Geotrupes mixtus Horn, 1868 

Geotrupes (Geotrupes) stercorarius (Linnaeus, 1758) (Scarabaeus) 

Geotrupes (Cnemotrupes) ulkei Blanchard, 1888 (Geotrupes) 

Mycotrupes LeConte, 1866 

Mycotrupes cartwrighti Olson and Hubbell, 1954 (Mycotrupes) 

Mycotrupes gaigei Olson and Hubbell, 1954 (Mycotrupes) 

Mycotrupes lethroides (Westwood, 1837) (Geotrupes) 

Mycotrupes pedester Howden, 1954 (Mycotrupes) 

Mycotrupes retusus (LeConte, 1866) (Geotrupes) 

Geotrupes aeneus Felsche, 1909 

Peltotrupes Blanchard, 1888 

Peltotrupes profundus (Howden, 1952) (Geotrupes) 

Geotrupes chalybaeus LeConte, 1878 

Peltotrupes dubius Howden, 1955 

Peltotrupes youngi Howden, 1955 (Peltotrupes) 

PASSALIDAE Leach, 1815 

PASSALINAE Leach, 1815 

PROCULINI Kaup, 1868 

Heliscus Zang, 1906 

Heliscus tropicus (Percheron, 1835) (Passalus) 

Odontotaenius Kuwert, 1896 

Odontotaenius disjunctus (Illiger, 1800) (Passalus) 

Passalus cornutus Fabricius, 1801 

Passalus distinctus Weber, 1801 

Passalus bos Kuwert, 1891 

Odontotaenius floridanus Schuster, 1994 (Odontotaenius) 

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Odontotaenius striatopunctatus (Percheron, 1835) (Passalus) 

PASSALINI Leach, 1815 

Passalus Fabricius, 1792 

Pertinax Kaup, 1869 

Neleides Kaup, 1869 

Petrejus Kaup, 1869 

Mitrorhinus Kaup, 1871 

Rhodocanthopus Kaup, 1871 

Epiphanus Kaup, 1871 

Rhagonocerus Kaup, 1871 

Valerius Kuwert, 1891 

Cassius Kuwert, 1891 

Lucilius Kuwert, 1891 

Pertinacides Kuwert, 1891 

Epipertinax Kuwert, 1891 

Ninoides Kuwert, 1891 

Neleurops Kuwert, 1891 

Manlius Kuwert, 1891 

Phoronaeosomus Kuwert, 1891 

Tetraracus Kuwert, 1891 

Lophocephalus Kuwert, 1896 

Aponelides Kuwert, 1896 

Parapertinax Kuwert, 1896 

Morosophus Kuwert, 1896 

Severus Kuwert, 1896 

Polyacanthopus Kuwert, 1896 

Synesius Kuwert, 1896 

Oeneus Kuwert, 1896 

Ptychotrichus Kuwert, 1896 

Calidas Kuwert, 1896 

Toxeutotaenius Kuwert, 1896 

Gnomon Zang, 1905 

Lasioperix Zang, 1905 

Phaulothorax Zang, 1905 

Scalmus Zang, 1905 

Psilomus Zang, 1905 

Macrolobus Zang, 1905 

Alocerus Zang, 1905 

Epipleurothrix Zang, 1905 

Zosterothrix Zang, 1905 

Procululus Zang, 1905 

Epiphoroneus Arrow, 1907 

Passalus punctatostriatus Percheron, 1835 (Passalus) 

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Passalus punctiger LePeletier and Serville, 1825 (Passalus) 

Passalus punctatissimus Eschscholtz, 1829 

Passalus striolatus Eschscholtz, 1829 

Neleus laborator Kaup, 1869 

Neleus transvalensis Kuwert, 1891 

Neleus guatemalae Kuwert, 1891 

Neleus difficilis Kuwert, 1891 

Neleus distinguendus Kuwert, 1891 

Neleus laeviclypeatus Kuwert, 1891 

Neleus scurra Kuwert, 1891 

Neleus sanio Kuwert, 1891 

Neleus chilensis Kuwert, 1891 

Neleus nicaraguae Kuwert, 1891 

Neleus scelus Kuwert, 1891 

Neleus vagans Kuwert, 1891 

Neleus scepticus Kuwert, 1891 

Neleus sulcicornis Kuwert, 1898 

Neleus arcuatotaeniatus Kuwert, 1898 

Neleus distinctus Kuwert, 1898 

Neleus intermissus Kuwert, 1898 

Neleus subcarinaefrons Kuwert, 1898 

Neleus subcarinatus Kuwert, 1898 

Neleus taeniolatus Kuwert, 1898 

Neleus dislocandus Kuwert, 1898 

Neleus scutellosulcatus Kuwert, 1898 

Neleus durangi Kuwert, 1898 

Neleus obtusecornutus Kuwert, 1898 

Neleus altidens Kuwert, 1898 

Neleus carinaefrons Kuwert, 1898 

Neleus acarinatus Kuwert, 1898 

Neleus dilatipunctatus Kuwert, 1898 

Neleus dispar Kuwert, 1898 

Neleus dilatus Kuwert, 1898 

Neleus aequatoris Kuwert, 1898 

Neleus difformis Kuwert, 1898 

Neleus argentinus Kuwert, 1898 

Neleus boliviensis Kuwert, 1898 

Neleus dilatidentatus Kuwert, 1898 

Neleus disjunctus Kuwert, 1898 

Neleus approximatidentatus Kuwert, 1898 

Neleus dispositus Kuwert, 1898 

Neleus labroexcisus Kuwert, 1898 

Neleus scurroides Kuwert, 1898 

Neleus arrogans Kuwert, 1898 

Neleus festae Rosmini, 1902 

Neleus cognettii Pangella, 1905 

7

Passalus riograndensis Luederwaldt, 1931 

Ptichopus Kaup, 1869 

Ptichopus angulatus (Percheron, 1835) (Passalus) 

Passalus nodus Apetz, 1837 

Passalus thoracicus Smith, 1852 

Ptichopus inca Kuwert, 1891 

Ptichopus nitidus Kuwert, 1891 

Ptichopus montezuma Kuwert, 1891 

Ptichopus aberrator Kuwert, 1891 

Ptichopus inflatus Kuwert, 1898 

Ptichopus borellii Rosmini, 1902 

TROGIDAE MacLeay, 1819 

Omorgus Erichson, 1847 

Chesas Burmeister, 1876 

Lagopelas Burmeister, 1876 

Megalotrox De Borre, 1886 

Omorgus asper LeConte, 1854 (Omorgus) 

Omorgus carinatus (Loomis, 1922) (Trox) 

Omorgus fuliginosus (Robinson, 1941) (Trox) 

Omorgus howelli (Howden and Vaurie, 1957) (Trox) 

Omorgus inflatus (Loomis, 1922) (Trox) 

Omorgus monachus (Herbst, 1790) (Trox) 

Trox tuberculatus Palisot de Beauvois, 1818 

Omorgus pustulatus LeConte, 1854 

Omorgus nodosus (Robinson, 1940) (Trox) 

Omorgus punctatus (Germar, 1824) (Trox) 

Omorgus integer LeConte, 1854 

Omorgus morsus LeConte, 1854 

Omorgus rubricans (Robinson, 1946) (Trox) 

Omorgus scabrosus (Palisot de Beauvois, 1818) (Trox) 

Omorgus scutellaris (Say, 1824) (Trox) 

Omorgus suberosus (Fabricius, 1775) (Trox) 

Trox crenatus Olivier, 1789 

Trox gibbus Olivier, 1789 

Trox ovatus Palisot de Beauvois, 1818 

Trox denticulatus Palisot de Beauvois, 1818 

Trox alternatus Say, 1831 

Trox nobilis Wollaston, 1867 

Trox tricolor Blackburn, 1904 

Trox manilensis Schultze, 1915 

Trox torressalai Baguena, 1959 

8

Trox triestinae Pittino, 1987 

Omorgus tesselatus LeConte, 1854 (Omorgus) 

Trox nigrescens Robinson, 1941 

Omorgus texanus LeConte, 1854 (Omorgus) 

Omorgus suturalis LeConte, 1854 

Trox oligonus Loomis, 1922 

Omorgus tomentosus (Robinson, 1941) (Trox) 

Omorgus tytus (Robinson, 1941) (Trox) 

Omorgus umbonatus LeConte, 1854 (Omorgus) 

Trox platycyphus Loomis, 1922 

Trox confluentus Robinson, 1941 

Trox Fabricius, 1775 

Pseudotrox Robinson, 1948 

Trox aequalis Say, 1832 (Trox) 

Trox affinis Robinson, 1940 (Trox) 

Trox atrox LeConte, 1854 (Trox) 

Trox capillaris Say, 1824 (Trox) 

Trox contractus Robinson, 1940 (Trox) 

Trox fascifer LeConte, 1854 (Trox) 

Trox floridanus Howden and Vaurie, 1957 (Trox) 

Trox foveicollis Harold, 1872 (Trox) 

Trox aciculeatus Robinson, 1940 

Trox frontera Vaurie, 1955 (Trox) 

Trox gemmulatus Horn, 1874 (Trox) 

Trox hamatus Robinson, 1940 (Trox) 

Trox laticollis LeConte, 1854 (Trox) 

Trox plicatus Robinson, 1940 (Trox) 

Trox robinsoni Vaurie, 1955 (Trox) 

Trox scaber (Linnaeus, 1767) (Silpha) 

Trox barbosus von Laicharting, 1781 

Trox arenarius Fabricius, 1787 

Trox arenosus Gmelin, 1790 

Trox hispidus von Paykull, 1798 

Trox trisulcatus Curtis, 1844 

Trox niponensis Lewis, 1895 

Trox sonorae LeConte, 1854 (Trox) 

Trox alternans LeConte, 1854 

Trox lecontei Harold, 1872 

Trox sordidus LeConte, 1854 (Trox) 

Trox spinulosus dentibius Robinson, 1940 (Trox) 

Trox spinulosus spinulosus Robinson, 1940 (Trox) 

Trox spinulosus simi Robinson, 1940 (Trox) 

Trox striatus Melsheimer, 1845 (Trox) 

Trox terrestris Say, 1825 (Trox) 

9

Trox tuberculatus (De Geer, 1774) (Scarabaeus) 

Trox serrulatus Palisot de Beauvois, 1818 

Trox canaliculatus Say, 1824 

Trox unistriatus Palisot de Beauvois, 1818 (Trox) 

Trox porcatus Say, 1825 

Trox variolatus Melsheimer, 1845 (Trox) 

Trox erinaceus LeConte, 1854 

Trox monticola Robinson, 1940 

GLARESIDAE Kolbe, 1905 

Glaresis Erichson, 1848 

Eoglaresis Semenov and Medvedev, 1932 

Eremoglaresis Semenov and Medvedev, 1932 

Afroglaresis Petrovitz, 1968 

Glaresis arenata Gordon, 1974 (Glaresis) 

Glaresis canadensis Brown, 1928 (Glaresis) 

Glaresis cartwrighti Gordon, 1970 (Glaresis) 

Glaresis clypeata Van Dyke, 1928 (Glaresis) 

Glaresis confusa Brown, 1928 (Glaresis) 

Glaresis dakotensis Gordon, 1970 (Glaresis) 

Glaresis ecostata Fall, 1907 (Glaresis) 

Glaresis gordoni Warner, 1995 (Glaresis) 

Glaresis howdeni Gordon, 1970 (Glaresis) 

Glaresis inducta Horn, 1885 (Glaresis) 

Glaresis knausi Brown, 1928 

Glaresis medialis Gordon, 1969 (Glaresis) 

Glaresis mendica Horn, 1885 (Glaresis) 

Glaresis phoenicis Fall, 1907 (Glaresis) 

Glaresis texana Gordon, 1970 (Glaresis) 

Glaresis zvirgzdinsi Warner, 1995 (Glaresis) 

DIPHYLLOSTOMATIDAE Holloway, 1972 

Diphyllostoma Fall, 1901 

Phyllostoma Fall, 1901 

Diphyllostoma fimbriatum (Fall, 1901) (Phyllostoma) 

Diphyllostoma linsleyi Fall, 1932 (Diphyllostoma) 

Diphyllostoma nigricollis Fall, 1912 (Diphyllostoma) 

LUCANIDAE Latreille, 1804 

AESALINAE MacLeay, 1819 

NICAGINI LeConte, 1861 

10

Nicagus LeConte, 1861 

Nicagus occultus Paulsen and Smith, 2005 (Nicagus) 

Nicagus obscurus (LeConte, 1847) (Ochodaeus) 

LUCANINAE Latreille, 1804 

LUCANINI Latreille, 1804 

Dorcus MacLeay, 1819 

Dorcus brevis (Say, 1825) (Lucanus) 

Dorcus caucasicus Ganglbauer, 1886 

Dorcus parallelus (Say, 1824) (Lucanus) 

Lucanus oblongus de Charpentier, 1825 

Dorcus costatus LeConte, 1866 

Dorcus nanus Casey, 1909 

Dorcus carnochani Angell, 1916 

Lucanus Scopoli, 1763 

Hexaphyllus Mulsant, 1839 

Pseudolucanus Hope, 1845 

Lucanus capreolus (Linnaeus, 1764) (Scarabaeus) 

Lucanus dama Fabricius, 1775 

Lucanus muticus Thunberg, 1806 

Lucanus trigonus Thunberg, 1806 

Lucanus elaphus Fabricius, 1775 (Lucanus) 

Lucanus carlengi Angell, 1916 

Lucanus mazama (LeConte, 1861) (Dorcus) 

Pseudolucanus bicostatus Angell, 1917 

Lucanus placidus Say, 1825 (Lucanus) 

Lucanus lentus Laporte, 1840 

PLATYCERINI Mulsant, 1842 

Platycerus Geoffroy, 1762 

Systenocerus Weise, 1883 

Systenus Sharp and Muir, 1912 

Platycerus depressus LeConte, 1850 (Platycerus) 

Platycerus marginalis Casey, 1897 (Platycerus) 

Platycerus cribripennis Van Dyke, 1928 

Platycerus oregonensis Westwood, 1846 (Platycerus) 

Platycerus coerulescens LeConte, 1869 

Platycerus chalybaeus Casey, 1897 

Platycerus virescens (Fabricius, 1775) (Lucanus) 

11

Lucanus quercus Weber, 1801 

Platycerus securidens Say, 1824 

Platycerus angustus Casey, 1909 

Platycerus iowanus Casey, 1909 

Platycerus peregrinus Casey, 1909 

PLATYCEROIDINI Paulsen and Hawks, 2008 

Platyceroides Benesh, 1946 

Platyceroides aeneus (Van Dyke, 1928) (Platycerus) 

Platyceroides agassii (LeConte, 1861) (Platycerus) 

Platycerus parvicollis Casey, 1889 

Platycerus pacificus Casey, 1889 

Platycerus californicus Casey, 1885 

Platycerus pygmaeus Van Dyke, 1946 

Platyceroides laticollis (Casey, 1914) (Platycerus) 

Platyceroides latus (Fall, 1901) (Platycerus) 

Platyceroides opacus (Fall, 1906) (Platycerus) 

Platyceroides thoracicus (Casey, 1895) (Platycerus) 

Platyceroides viriditinctus (Benesh, 1942) (Platycerus) 

Platyceropsis Benesh, 1946 

Platyceropsis keeni (Casey, 1895) (Platycerus) 

Platycerus pedicellaris Möllenkamp, 1911 

SYNDESINAE MacLeay, 1819 

Ceruchus MacLeay, 1819 

Ceruchus piceus (Weber, 1801) (Lucanus) 

Platycerus balbi Laporte, 1840 

Ceruchus virginiensis Casey, 1914 

Ceruchus punctatus LeConte, 1869 (Ceruchus) 

Ceruchus striatus LeConte, 1859 (Ceruchus) 

Sinodendron Hellwig, 1791 

Ligniperda Fabricius, 1790 

Sinodendron rugosum Mannerheim, 1843 (Sinodendron) 

OCHODAEIDAE Mulsant and Rey, 1871 

OCHODAEINAE Mulsant and Rey, 1871 

OCHODAEINI Mulsant and Rey, 1871 

12

Codocera Eschscholtz, 1821 

Stomphax Fischer von Waldheim, 1823 

Codocera gnatho (Fall, 1907) (Ochodaeus) 

Ochodaeus nimius Fall, 1907 

Cucochodaeus Paulsen, 2007 

Cucochodaeus sparsus (LeConte, 1868) (Ochodaeus) 

Ochodaeus mandibularis Linell, 1896 

Neochodaeus Nikolayev, 1995 

Neochodaeus frontalis (LeConte, 1863) (Ochodaeus) 

Ochodaeus complex LeConte, 1868 

Neochodaeus praesidii (Bates, 1887) (Ochodaeus) 

Neochodaeus repandus (Fall, 1909) (Ochodaeus) 

Neochodaeus striatus (LeConte, 1854) (Ochodaeus) 

Parochodaeus Nikolayev, 1995 

Parochodaeus biarmatus (LeConte, 1868) (Ochodaeus) 

Parochodaeus californicus (Horn, 1895) (Ochodaeus) 

Parochodaeus duplex (LeConte, 1868) (Ochodaeus) 

Ochodaeus kansanus Fall, 1909 

Parochodaeus howdeni (Carlson, 1975) (Ochodaeus) 

Parochodaeus inarmatus (Schaeffer, 1906) (Ochodaeus) 

Parochodaeus pectoralis (LeConte, 1868) (Ochodaeus) 

Parochodaeus peninsularis (Horn, 1895) (Ochodaeus) 

Parochodaeus ritcheri (Carlson, 1975) (Ochodaeus) 

Xenochodaeus Paulsen, 2007 

Xenochodaeus americanus (Westwood, 1852) (Ochodaeus) 

Ochodaeus opacus LeConte, 1868 

Xenochodaeus luscinus (Howden, 1968) (Ochodaeus) 

Xenochodaeus musculus (Say, 1835) (Odontaeus) 

Xenochodaeus planifrons (Schaeffer, 1906) (Ochodaeus) 

Xenochodaeus simplex (LeConte, 1854) (Ochodaeus) 

Xenochodaeus ulkei (Horn, 1876) (Ochodaeus) 

CHAETOCANTHINAE Scholtz, 1988 

PSEUDOCHODAEINI Scholtz, 1988 

Pseudochodaeus Carlson and Ritcher, 1974 

13

Pseudochodaeus estriatus (Schaeffer, 1906) (Ochodaeus) 

HYBOSORIDAE Erichson, 1847 

HYBOSORINAE Erichson, 1847 

Hybosorus MacLeay, 1819 

Hybosorus illigeri Reiche, 1853 (Hybosorus) 

Scarabaeus arator Illiger, 1803 

Hybosorus pinguis Westwood, 1845 

Hybosorus roei Westwood, 1845 

Hybosorus carolinus LeConte, 1848 

Hybosorus nitidus Lansberge, 1882 

Hybosorus nossibianus Fairmaire, 1895 

Hybosorus palearcticus Endrödi, 1957 

CERATOCANTHINAE Martínez, 1968 

CERATOCANTHINI Martínez, 1968 

Ceratocanthus White, 1842 

Acanthocerus MacLeay, 1819 

Sphaeromorphus Germar, 1843 

Gymnoropterus Gestro, 1899 

Ceratocanthus aeneus (MacLeay, 1819) (Acanthocerus) 

Sphaeromorphus volvox Erichson, 1843 

Germarostes Paulian, 1982 

Germarostes (Germarostes) Paulian, 1982 

Germarostes (Germarostes) aphodioides (Illiger, 1800) (Melolontha) 

Scarabaeus latipes Germar, 1824 

Trox splendidus Say, 1835 

Acanthocerus laevistriatus Laporte, 1840 

Cloeotus prionomus Bates, 1887 

Germarostes (Germarostes) globosus (Say, 1835) (Trox) 

PACHYPLECTRINAE Ocampo, 2006 

Pachyplectrus LeConte, 1874 

Pachyplectrus laevis LeConte, 1874 (Pachyplectrus) 

GLAPHYRIDAE MacLeay, 1819 

GLAPHYRINAE MacLeay, 1819 

14

Lichnanthe Burmeister, 1844 

Dasydera LeConte, 1861 

Lichnanthe albipilosa Carlson, 1980 (Lichnanthe) 

Lichnanthe apina Carlson, 1980 (Lichnanthe) 

Lichnanthe brachyselis Carlson, 1980 (Lichnanthe) 

Lichnanthe cooperi (Horn, 1867) (Dasydera) 

Lichnanthe lupina LeConte, 1856 (Lichnanthe) 

Lichnanthe rathvoni (LeConte, 1863) (Dasydera) 

Lichnanthe canina Horn, 1867 

Lichnanthe edwardsi Horn, 1870 

Lichnanthe ursina (LeConte, 1861) (Dasydera) 

Lichnanthe vulpina (Hentz, 1827) (Amphicoma) 

SCARABAEIDAE Latreille, 1802 

APHODIINAE Leach, 1815 

AEGIALIINI Laporte, 1840 

Aegialia Latreille, 1807 

Aegialia (Aegialia) Latreille, 1807 

Aegialia (Psammoporus) Thomson, 1863 

Dimalia Mulsant and Rey, 1870 

Aegialia (Aegialia) amplipunctata Gordon and Cartwright, 1988 (Aegialia) 

Aegialia (Aegialia) arenaria (Fabricius, 1787) (Scarabaeus) 

Scarabaeus globosus Kugelann, 1794 

Aegialia (Aegialia) blanchardi Horn, 1887 (Aegialia) 

Aegialia (Aegialia) carri Gordon and Cartwright, 1988 (Aegialia) 

Aegialia (Aegialia) cartwrighti Stebnicka, 1977 (Aegialia) 

Aegialia (Aegialia) concinna Gordon and Cartwright, 1977 (Aegialia) 

Aegialia (Aegialia) conferta Horn, 1871 (Aegialia) 

Aegialia (Aegialia) convexa Fall, 1932 (Aegialia) 

Aegialia (Aegialia) crassa LeConte, 1857 (Aegialia) 

Aegialia insularis Brown, 1931 

Aegialia (Aegialia) crescenta Gordon and Cartwright, 1977 (Aegialia) 

Aegialia (Psammoporus) criddlei Brown, 1931 (Aegialia) 

Aegialia (Psammoporus) cylindrica (Eschscholtz, 1822) (Psammodius) 

Aegialia (Psammoporus) exarata Mannerheim, 1853 (Aegialia) 

Aegialia (Aegialia) hardyi Gordon and Cartwright, 1977 (Aegialia) 

Aegialia (Aegialia) kelsoi Gordon and Cartwright, 1988 (Aegialia) 

Aegialia (Aegialia) knighti Gordon and Rust, 1997 (Aegialia) 

Aegialia (Psammoporus) lacustris LeConte, 1850 (Aegialia) 

Aegialia (Aegialia) latispina LeConte, 1878 (Aegialia) 

Aegialia (Aegialia) magnifica Gordon and Cartwright, 1977 (Aegialia) 

15

Aegialia (Aegialia) mcclevei Gordon, 1990 (Aegialia) 

Aegialia (Psammoporus) nana Brown, 1931 (Aegialia) 

Aegialia (Aegialia) nigrella Brown, 1931 (Aegialia) 

Aegialia (Psammoporus) opaca Brown, 1931 (Aegialia) 

Aegialia (Aegialia) opifex Horn, 1887 (Aegialia) 

Aegialia (Aegialia) punctata Brown, 1931 (Aegialia) 

Aegialia (Aegialia) spinosa Gordon and Cartwright, 1988 (Aegialia) 

Aegialia (Psammoporus) terminalis Brown, 1931 (Aegialia) 

Caelius Lewis, 1895 

Leptaegialia Brown, 1931 

Caelius browni (Saylor, 1934) (Aegialia) 

Caelius humeralis (Brown, 1931) (Aegialia) 

Caelius montana (Brown, 1931) (Aegialia) 

Caelius rufescens (Horn, 1887) (Aegialia) 

Aegialia rufa LeConte, 1878 

Micraegialia Brown, 1931 

Micraegialia pusilla (Horn, 1887) (Aegialia) 

Rhysothorax Bedel, 1911 

Anomalaegialia Brown, 1931 

Rhysothorax rufus (Fabricius, 1792) (Scarabaeus) 

Aegialia spissipes LeConte, 1878 

Aegialia rufina Silfverberg, 1977 

APHODIINI Leach, 1815 

Acrossus Mulsant, 1842 

Hemicyclorhinus Seabra, 1909 

Acrossus rubripennis (Horn, 1870) (Aphodius) 

Acrossus rufipes (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus oblongus Scopoli, 1763 

Scarabaeus capitatus De Geer, 1774 

Aphodius muticus Stephens, 1830 

Aphodius juvenilis Mulsant, 1842 

Aphodius boum Gistel, 1857 

Aphodius capicola Harold, 1862 

Aphodius rufotestaceus Dalla Torre, 1879 

Aphodius matsuzawai Yawata, 1943 

Agrilinellus Dellacasa, Dellacasa, and Gordon, 2008 

16

Agrilinellus nuriae Dellacasa, Dellacasa, and Gordon, 2008 (Agrilinellus) 

Agrilinellus ornatus (Schmidt, 1911) (Aphodius) 

Aphodius magnopunctatus Hinton, 1934 

Agoliinus Schmidt, 1913 

Agoliinus albertanus (Brown, 1928) (Aphodius) 

Agoliinus aleutus (Eschscholtz, 1822) (Aphodius) 

Aphodius ursinus Motschulsky, 1845 

Agoliinus anthracus Gordon and Skelley, 2007 (Agoliinus) 

Aphodius anthracinus LeConte, 1878 

Agoliinus aquilonarius (Brown, 1928) (Aphodius) 

Agoliinus ashworthi (Gordon, 2006) (Aphodius) 

Agoliinus bidentatus (Schmidt, 1907) (Aphodius) 

Aphodius bidens LeConte, 1878 

Agoliinus canadensis (Garnett, 1920) (Aphodius) 

Aphodius phaeopterus LeConte, 1878 

Aphodius haywardi Brown, 1928 

Aphodius russelli Hatch, 1971 

Agoliinus congregatus (Mannerheim, 1853) (Aphodius) 

Aphodius arcticus Harold, 1863 

Aphodius richardsonae Hatch, 1971 

Agoliinus corruptor (Brown, 1929) (Aphodius) 

Aphodius explanatus Brown, 1928 

Aphodius yukonensis Robinson, 1948 

Aphodius calgaryensis Balthasar, 1970 

Agoliinus cruentatus (LeConte, 1878) (Aphodius) 

Aphodius snowi Brown, 1929 

Agoliinus explanatus (LeConte, 1878) (Aphodius) 

Agoliinus guttatus (Eschscholtz, 1823) (Aphodius) 

Agoliinus hatchi Gordon and Skelley, 2007 (Agoliinus) 

Aphodius angustus Hatch, 1971 

Agoliinus idahoensis Gordon and Skelley, 2007 (Agoliinus) 

Agoliinus incommunis (Fall, 1932) (Aphodius) 

Agoliinus ingenursus Gordon and Skelley, 2007 (Agoliinus) 

Agoliinus leopardus (Horn, 1870) (Aphodius) 

Agoliinus malkini (Hatch, 1971) (Aphodius) 

Agoliinus manitobensis (Brown, 1928) (Aphodius) 

Agoliinus piceatus (Robinson, 1946) (Aphodius) 

Aphodius rainieri Hatch, 1971 

Agoliinus plutonicus (Fall, 1907) (Aphodius) 

Agoliinus poudreus Gordon and Skelley, 2007 (Agoliinus) 

Agoliinus praealtus Gordon and Skelley, 2007 (Agoliinus) 

Agoliinus sigmoideus (Van Dyke, 1918) (Aphodius) 

Agoliinus wickhami (Brown, 1928) (Aphodius) 

17

Aphodius Illiger, 1798 

Aphodius fimetarius (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus pedellus De Geer, 1774 

Scarabaeus bicolor Fourcroy, 1785 

Scarabaeus autumnalis Naezen, 1792 

Aphodius orophilus de Charpentier, 1825 

Aphodius nodifrons Randall, 1838 

Ballucus Gordon and Skelley, 2007 

Ballucus barri (Gordon, 2006) (Aphodius) 

Ballucus bruneaui Gordon and Skelley, 2007 (Ballucus) 

Ballucus oregonus Gordon and Skelley, 2007 (Ballucus) 

Ballucus obtusus (LeConte, 1878) (Aphodius) 

Aphodius utahensis Robinson, 1938 

Blackburneus Schmidt, 1913 

Blackburneus aegrotus (Horn, 1870) (Aphodius) 

Aphodius campestris Blatchley, 1912 

Blackburneus cavidomus (Brown, 1927) (Aphodius) 

Blackburneus cynomysi (Brown, 1927) (Aphodius) 

Blackburneus fordi (Gordon, 1974) (Aphodius) 

Blackburneus geomysi (Cartwright, 1939) (Aphodius) 

Blackburneus lentus (Horn, 1870) (Aphodius) 

Blackburneus rubeolus (Palisot de Beauvois, 1809) (Scarabaeus) 

Aphodius copronymus Melsheimer, 1845 

Blackburneus saylorea (Robinson, 1940) (Aphodius) 

Blackburneus stercorosus (Melsheimer, 1845) (Aphodius) 

Blackburneus tenuistriatus (Horn, 1887) (Aphodius) 

Blackburneus troglodytes (Hubbard, 1894) (Aphodius) 

Calamosternus Motschulsky, 1859 

Megalisus Mulsant and Rey, 1870 

Calamosternus granarius (Linnaeus, 1767) (Scarabaeus) 

Scarabaeus quadrituberculatus Fabricius, 1798 

Aphodius niger Creutzer, 1799 

Aphodius carbonarius Sturm, 1805 

Aphodius haemorrhous Stephens, 1830 

Aphodius lucens Stephens, 1830 

Aphodius emarginatus Stephens, 1830 

Aphodius melanopus Stephens, 1830 

Aphodius elongatus Ménétries, 1832 

18

Aphodius retusus Waltl, 1835 

Aphodius suturalis Faldermann, 1835 

Aphodius parcepunctatus Mulsant, 1842 

Aphodius cribratus Mulsant, 1842 

Aphodius rugosulus Mulsant, 1842 

Aphodius concolor Mulsant, 1842 

Aphodius aterrimus Melsheimer, 1845 

Aphodius adelaidae Hope, 1847 

Aphodius metallicus Haldeman, 1848 

Aphodius spretus Haldeman, 1848 

Aphodius thoracicus Dalla Torre, 1879 

Aphodius inutilis Horn, 1887 

Aphodius brunnescens Reitter, 1892 

Aphodius ragusanus Reitter, 1906 

Aphodius nama Kolbe, 1908 

Aphodius signatus Schmidt, 1916 

Aphodius pirinensis Balthasar 1946 

Caligodorus Gordon and Skelley, 2007 

Caligodorus opacus (LeConte, 1872) (Aphodius) 

Aphodius columbiensis Wickham, 1913 

Caligodorus tuberosus (Barrett, 1932) (Aphodius) 

Aphodius schuhi Gordon, 1975 

Caligodorus vandykei (Barrett, 1931) (Aphodius) 

Cephalocyclus Dellacasa, Gordon, and Dellacasa, 1998 

Cephalocyclus bordati Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Cephalocyclus carmenae Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Cephalocyclus durangoensis (Bates, 1887) (Aphodius) 

Aphodius pugil Balthasar, 1946 

Cephalocyclus fuliginosus (Harold, 1863) (Aphodius) 

Cephalocyclus gravidus (Harold, 1863) (Aphodius) 

Cephalocyclus hogei (Bates, 1887) (Aphodius) 

Aphodius transversus Robinson, 1940 

Cephalocyclus howdenorum Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Cephalocyclus lagoi Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Cephalocyclus luteolus (Horn, 1887) (Aphodius) 

Cephalocyclus mexicanus (Harold, 1862) (Aphodius) 

Aphodius omiltemius Bates, 1889 

Cephalocyclus potosinus Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Cephalocyclus pullatus (Schmidt, 1913) (Aphodius) 

Cephalocyclus rockefelleri Dellacasa, Dellacasa, and Gordon, 2007 (Cephalocyclus) 

Chilothorax Motschulsky, 1859 

19

Chilothorax distinctus (Müller, 1776) (Scarabaeus) 

Scarabaeus inquinatus Herbst, 1783 

Scarabaeus centrolineatus Panzer, 1798 

Scarabaeus nubilus Panzer, 1798 

Scarabaeus attaminatus Marsham, 1802 

Scarabaeus foedatus Marsham, 1802 

Aphodius anxius Mulsant, 1842 

Aphodius auctus Mulsant, 1842 

Aphodius baseolus Mulsant, 1842 

Aphodius fumosus Mulsant, 1842 

Aphodius hemicyclus Mulsant, 1842 

Aphodius interruptus Mulsant, 1842 

Aphodius lunatus Mulsant, 1842 

Aphodius ophthalmicus Mulsant, 1842 

Aphodius pauper Mulsant, 1842 

Aphodius scutellaris Mulsant, 1842 

Aphodius subcinctus Mulsant, 1842 

Aphodius maculipennis Melsheimer, 1845 

Aphodius hypocoprus Gistel, 1857 

Aphodius confluens Schilsky, 1888 

Aphodius inquinatulus Reitter, 1898 

Aphodius striatulus Eichler, 1922 

Aphodius humeralis Báguena, 1927 

Aphodius lopezromeui Báguena, 1927 

Aphodius centrolineatoides Strand and Gulbis, 1957 

Aphodius distinctoides Strand and Gulbis, 1957 

Aphodius nubiloides Strand and Gulbis, 1957 

Aphodius pseudonubilus Strand and Gulbis, 1957 

Aphodius pulcheroides Strand and Gulbis, 1957 

Aphodius subconfluens Strand and Gulbis, 1957 

Aphodius trifasciatus Strand and Gulbis, 1957 

Aphodius libyanus Balthasar, 1961 

Cinacanthus Schmidt, 1913 

Cinacanthus coquilletti (Linell, 1896) (Aphodius) 

Cinacanthus crenicollis (Fall, 1932) (Aphodius) 

Aphodius spermophili Gordon, 1973 

Cinacanthus crescentus Gordon and Skelley, 2007 (Cinacanthus) 

Cinacanthus hirsutus (Brown, 1928) (Aphodius) 

Cinacanthus militaris (LeConte, 1858) (Aphodius) 

Cinacanthus ulkei (Gordon, 2006) (Aphodius) 

Coelotrachelus Schmidt, 1913 

20

Coelotrachelus kuntzeni (Schmidt, 1913) (Aphodius) 

Aphodius amplinotum Gordon and Howden, 1973 

Aphodius michiliensis Deloya, 1991 

Coelotrachelus rudis (LeConte, 1878) (Aphodius) 

Aphodius fenyesi Van Dyke, 1947 

Coelotrachelus symbius (Gordon and Howden, 1973) (Aphodius) 

Colobopterus Mulsant, 1842 

Colobopterus erraticus (Linnaeus, 1758) (Scarabaeus) 

Aphodius striolatus Gebler, 1832 

Colobopterus fumigatus Mulsant, 1842 

Colobopterus nebulosus Mulsant, 1842 

Colobopterus submaculatus Mulsant, 1842 

Aphodius gonagricus Fischer von Waldheim, 1844 

Aphodius pensvallensis Melsheimer, 1845 

Aphodius capraram Gistel, 1857 

Aphodius bistrigatus Dalla Torre, 1879 

Aphodius lineatus Dalla Torre, 1879 

Aphodius melancholicus Dalla Torre, 1879 

Aphodius unicolor Dalla Torre, 1879 

Aphodius taeniatus Schilsky, 1888 

Aphodius zighanicus Eichler, 1922 

Aphodius limitinus Schmidt, 1922 

Aphodius korbeli Roubal, 1935 

Cryptoscatomaseter Gordon and Skelley, 2007 

Cryptoscatomaseter acuminatus (Cartwright, 1944) (Aphodius) 

Cryptoscatomaseter brevicollis (LeConte, 1878) (Aphodius) 

Cryptoscatomaseter browni (Hinton, 1934) (Aphodius) 

Aphodius smithi Brown, 1930 

Cryptoscatomaseter caccabatus (Warner and Skelley, 2006) (Aphodius) 

Cryptoscatomaseter captivus (Cartwright, 1944) (Aphodius) 

Cryptoscatomaseter criddlei (Brown, 1928) (Aphodius) 

Cryptoscatomaseter depressiusculus (Schmidt, 1907) (Aphodius) 

Aphodius marginatus LeConte, 1878 

Cryptoscatomaseter durangoi (Gordon and Howden, 1973) (Aphodius) 

Cryptoscatomaseter ejectus Gordon and Skelley, 2007 (Cryptoscatomaseter) 

Cryptoscatomaseter epulus (Gordon and Howden, 1973) (Aphodius) 

Cryptoscatomaseter excavatus (Robinson, 1940) (Aphodius) 

Cryptoscatomaseter gambrinus (Skelley and Gordon, 2002) (Aphodius) 

Cryptoscatomaseter haldemani (Horn, 1887) (Aphodius) 

Aphodius politus Horn, 1870 

Cryptoscatomaseter hubbelli (Skelley and Woodruff, 1991) (Aphodius) 

Cryptoscatomaseter incomptus Gordon and Skelley, 2007 (Cryptoscatomaseter) 

21

Cryptoscatomaseter iowensis (Wickham, 1913) (Aphodius) 

Cryptoscatomaseter magnificens (Robinson, 1940) (Aphodius) 

Cryptoscatomaseter ochreipennis (Horn, 1871) (Aphodius) 

Cryptoscatomaseter oklahomensis (Brown, 1928) (Aphodius) 

Aphodius atwateri Cartwright, 1944 

Cryptoscatomaseter paulseni (Skelley, 2006) (Aphodius) 

Cryptoscatomaseter pholetus (Skelley and Woodruff, 1991) (Aphodius) 

Cryptoscatomaseter platypleurus (Skelley and Woodruff, 1991) (Aphodius) 

Cryptoscatomaseter punctissimus (Brown, 1928) (Aphodius) 

Aphodius socialis Brown, 1928 

Cryptoscatomaseter rossi (Cartwright, 1944) (Aphodius) 

Cryptoscatomaseter salsburyi (Gordon, 2006) (Aphodius) 

Cryptoscatomaseter talpoidesi (Brown, 1928) (Aphodius) 

Cryptoscatomaseter umbricollis (Fall, 1907) (Aphodius) 

Cryptoscatomaseter utopensis (Warner and Skelley, 2006) (Aphodius) 

Cryptoscatomaseter yavapai (Robinson, 1946) (Aphodius) 

Dellacasiellus Gordon and Skelley, 2007 

Dellacasiellus carpinterius Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus claudus (Fall, 1932) (Aphodius) 

Dellacasiellus concavus (Say, 1823) (Aphodius) 

Aphodius cressoni Robinson, 1946 

Dellacasiellus fucosus (Schmidt, 1922) (Aphodius) 

Aphodius rubidus LeConte, 1857 

Dellacasiellus glamisensis Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus griffini Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus humboldti Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus kirni (Cartwright, 1944) (Aphodius) 

Dellacasiellus laevigatus (Haldeman, 1848) (Aphodius) 

Aphodius goffi Cartwright, 1939 

Dellacasiellus milleri Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus pseudofucosus Gordon and Skelley, 2007 (Dellacasiellus) 

Dellacasiellus ruficlarus (Fall, 1932) (Aphodius) 

Aphodius subcarinatus Robinson, 1946 

Dialytellus Brown, 1929 

Dialytellus dialytoides (Fall, 1907) (Aphodius) 

Dialytellus tragicus (Schmidt, 1916) (Aphodius) 

Aphodius humeralis LeConte, 1878 

Aphodius lecontei Balthasar, 1941 

Dialytes Harold, 1869 

Dialytes criddlei Brown, 1929 (Dialytes) 

22

Dialytes striatulus (Say, 1825) (Trox) 

Rhyssemus cribrosus LeConte, 1850 

Dialytes truncatus (Melsheimer, 1845) (Aphodius) 

Aphodius corvinus Haldeman, 1848 

Dialytes ulkei Horn, 1875 (Dialytes) 

Dialytodius Gordon and Skelley, 2007 

Dialytodius carri (Brown, 1929) (Aphodius) 

Dialytodius decipiens (Horn, 1887) (Aphodius) 

Dialytodius rogersi (Hatch, 1971) (Aphodius) 

Dialytodius olympicus Gordon and Skelley, 2007 (Aphodius) 

Diapterna Horn, 1887 

Diapterna hamata (Say, 1824) (Aphodius) 

Aphodius occidentalis Horn, 1870 

Aphodius garmani Brown, 1929 

Diapterna hyperborea (LeConte, 1850) (Aphodius) 

Aphodius angularis LeConte, 1850 

Aphodius sagittarius Harold, 1860 

Diapterna omissa (LeConte, 1850) (Aphodius) 

Aphodius torpidus Horn, 1870 

Diapterna pinguella (Brown, 1929) (Aphodius) 

Diapterna pinguis (Haldeman, 1848) (Aphodius) 

Aphodius validus Horn, 1870 

Drepanocanthoides Schmidt, 1913 

Drepanocanthoides larreae (Horn, 1887) (Aphodius) 

Aphodius bifrons Bates, 1887 

Drepanocanthoides walshii (Horn, 1870) (Aphodius) 

Eupleurus Mulsant, 1842 

Eupleurus subterraneus (Linnaeus, 1758) (Scarabaeus) 

Flaviellus Gordon and Skelley, 2007 

Flaviellus aggregatus (Gordon, 1977) (Aphodius) 

Flaviellus gordoni (Ratcliffe, 1988) (Aphodius) 

Flaviellus perfimbriatus (Gordon, 1977) (Aphodius) 

Flaviellus phalerioides (Horn, 1870) (Aphodius) 

Flaviellus phalerius Gordon and Skelley, 2007 (Flaviellus) 

Flaviellus subtruncatus (LeConte, 1878) (Aphodius) 

Aphodius rehni Robinson, 1938 

23

Geomyphilus Gordon and Skelley, 2007 

Geomyphilus alabama (Skelley and Gordon, 1995) (Aphodius) 

Geomyphilus essigi (Saylor, 1935) (Aphodius) 

Geomyphilus geronimo Gordon and Skelley, 2007 (Geomyphilus) 

Geomyphilus insolitus (Brown, 1928) (Aphodius) 

Geomyphilus kiowensis (Gordon and Salsbury, 1999) (Aphodius) 

Geomyphilus leptotarsis (Brown, 1928) (Aphodius) 

Geomyphilus pinolus Gordon and Skelley, 2007 (Geomyphilus) 

Geomyphilus rubiginosus (Horn, 1870) (Aphodius) 

Geomyphilus russeus (Brown, 1928) (Aphodius) 

Geomyphilus thomomysi (Brown, 1928) (Aphodius) 

Geomyphilus ungulatus (Fall, 1901) (Aphodius) 

Geomyphilus viceversus (Paulsen, 2006) (Aphodius) 

Haroldiellus Gordon and Skelley, 2007 

Haroldiellus sallei (Harold, 1863) (Aphodius) 

Aphodius freyi Balthasar, 1941 

Hornietus Stebnicka, 2000 

Hornietus ventralis (Horn, 1887) (Pleurophorus) 

Ataenius braunarum Dury, 1902 (Ataenius) 

Irrasinus Gordon and Skelley, 2007 

Irrasinus dumasi Gordon and Skelley, 2007 (Irrasinus) 

Irrasinus stupidus (Horn, 1870) (Aphodius) 

Labarrus Mulsant and Rey, 1870 

Pseudocalamosternus Balthasar, 1936 

Labarrus lividus (Olivier, 1789) (Scarabaeus) 

Scarabaeus suturalis Fabricius, 1792 

Aphodius obsoletus Fabricius, 1801 

Scarabaeus biliteratus Marsham, 1802 

Aphodius discus Wiedemann, 1823 

Aphodius cincticulus Hope, 1847 

Aphodius spilopterus Germar, 1848 

Aphodius scutellaris Roth, 1851 

Aphodius sequens Walker, 1858 

Aphodius maculicollis Montrouzier, 1860 

Aphodius matusitai Nakane, 1961 

Labarrus pseudolividus Balthasar, 1941 (Aphodius) 

24

Lechorodius Gordon and Skelley, 2007 

Lechorodius lutulentus (Haldeman, 1843) (Aphodius) 

Lechorodius terminalis (Say, 1823) (Aphodius) 

Liothorax Motschulsky, 1859 

Ataeniomorphus Balthasar, 1953 

Liothorax alternatus (Horn, 1870) (Aphodius) 

Aphodius inermis Robinson, 1940 

Aphodius moclipsi Hatch, 1971 

Liothorax consociatus (Horn, 1887) (Aphodius) 

Liothorax innexus (Say, 1835) (Aphodius) 

Aphodius flavocinctus Harold, 1860 

Liothorax subaeneus (LeConte, 1857) (Aphodius) 

Aphodius gentneri Hatch, 1971 

Aphodius perpolitus Gordon, 1973 

Luxolinus Gordon and Skelley, 2007 

Luxolinus luxatus (Horn, 1887) (Aphodius) 

Maculaphodius Gordon and Skelley, 2007 

Maculaphodius conspersus (Horn, 1887) (Aphodius) 

Melinopterus Mulsant, 1842 

Melinopterus consentaneus (LeConte, 1850) (Aphodius) 

Melinopterus femoralis (Say, 1823) (Aphodius) 

Melinopterus prodromus (Brahm, 1790) (Scarabaeus) 

Aphodius rapax Faldermann, 1835 

Aphodius angustatus Mulsant, 1842 

Aphodius flavogriseus Mulsant, 1842 

Aphodius griseolus Mulsant, 1842 

Aphodius obliquus Mulsant, 1842 

Aphodius restrictus Mulsant, 1842 

Aphodius semilunus Mulsant, 1842 

Aphodius discoidalis Gistel, 1857 

Aphodius syriacus Mulsant and Rey, 1870 

Aphodius semipellitus Solsky, 1876 

Aphodius griseus Dalla Torre, 1879 

Aphodius mulsanti d’Orbigny, 1896 

Aphodius weberi Reitter, 1906 

Aphodius gisteli Strand, 1917 

25

Aphodius abscurus Eichler, 1922 

Aphodius lunulatus Eichler, 1922 

Mendidius Harold, 1868 

Cnemargus Harold, 1866 

Aphodosugrames Endrödi, 1964 

Mendidius aculeatus (Robinson, 1940) (Aphodius) 

Merogyrus Gordon and Skelley, 2007 

Merogyrus rotundiceps (Fall, 1907) (Aphodius) 

Neotrichonotulus Dellacasa, Gordon, and Dellacasa, 2004 

Neotrichonotulus inurbanus (Gordon and Howden, 1973) (Aphodius) 

Nialaphodius Kolbe, 1908 

Nialaphodius nigritus (Fabricius, 1801) (Aphodius) 

Aphodius nigritulus Boheman, 1857 

Aphodius cuniculus Chevrolat, 1864 

Aphodius paivanus Wollaston, 1867 

Aphodius vestiarius Horn, 1870 

Aphodius expertus Harold, 1871 

Aphodius guadeloupensis Fleutiaux and Sallé, 1889 

Aphodius tuberifrons Fairmaire, 1903 

Aphodius binominatus Schmidt, 1907 

Aphodius pervicax Balthasar, 1942 

Oscarinus Gordon and Skelley, 2007 

Oscarinus abusus (Fall, 1907) (Aphodius) 

Oscarinus bottimeri (Cartwright, 1957) (Aphodius) 

Oscarinus brimleyi (Cartwright, 1939) (Aphodius) 

Oscarinus crassuloides (Fall, 1907) (Aphodius) 

Oscarinus crassulus (Horn, 1870) (Aphodius) 

Oscarinus floridanus (Robinson, 1947) (Aphodius) 

Oscarinus lodingi (Cartwright, 1957) (Aphodius) 

Oscarinus matiganae (Paulsen, 2006) (Aphodius) 

Oscarinus odocoilis (Robinson, 1939) (Aphodius) 

Oscarinus pseudabusus (Cartwright, 1957) (Aphodius) 

Oscarinus rusicola (Melsheimer, 1845) (Aphodius) 

Aphodius curtus Haldeman, 1848 

Aphodius aurelianus Harold, 1863 

Aphodius mcdonaldi Robinson, 1946 

26

Oscarinus silvanicus (Cartwright, 1972) (Aphodius) 

Oscarinus stuessyi Gordon and Skelley, 2007 (Oscarinus) 

Oscarinus texensis (Cartwright, 1972) (Aphodius) 

Oscarinus welderi Gordon and Skelley, 2007 (Oscarinus) 

Oscarinus windsori (Cartwright, 1939) (Aphodius) 

Otophorus Mulsant, 1842 

Otophorus haemorrhoidalis (Linnaeus, 1758) (Scarabaeus) 

Oxyomus Dejean, 1833 

Amidorinus Koshantschikov, 1912 

Oxyomus silvestris (Scopoli, 1763) (Scarabaeus) 

Scarabaeus porcatus Fabricius, 1775 

Scarabaeus fenestralis Schrank, 1781 

Scarabaeus foveolatus Moll, 1785 

Oxyomus opacifrons Horn, 1871 

Pardalosus Gordon and Skelley, 2007 

Pardalosus neodistinctus (Brown, 1928) (Aphodius) 

Pardalosus pardalis (LeConte, 1857) (Aphodius) 

Pardalosus pseudopardalis Gordon and Skelley, 2007 (Pardalosus) 

Pardalosus pumilio (Schmidt, 1907) (Aphodius) 

Aphodius pumilus Horn, 1887 

Pardalosus sayi (Gordon, 2006) (Aphodius) 

Pardalosus serval (Say, 1835) (Aphodius) 

Aphodius steinheili Harold, 1869 

Aphodius pennsylvanicus Robinson, 1940 

Pardalosus slevini (Van Dyke, 1928) (Aphodius) 

Aphodius davisi Fall, 1930 

Phaeaphodius Reitter, 1892 

Limaroides Balthasar, 1932 

Phaeaphodius rectus (Motschulsky, 1866) (Aphodius) 

Aphodius vitta Motschulsky, 1866 

Aphodius solskyi Harold, 1871 

Aphodius castaneipennis Waterhouse, 1875 

Aphodius diversus Waterhouse, 1875 

Aphodius lividipennis Waterhouse, 1875 

Aphodius biformis Reitter, 1892 

Aphodius semicolor Reitter, 1892 

Aphodius zichyi Csiki, 1901 

Aphodius controversus Balthasar, 1932 

27

Aphodius diabolicus Balthasar, 1932 

Aphodius agraphus Mader, 1953 

Aphodius edai Balthasar, 1956 

Aphodius tshuensis Protsenko, 1979 

Planolinellus Dellacasa and Dellacasa, 2005 

Planolinellus vittatus (Say, 1825) (Aphodius) 

Aphodius sellatus Mannerheim, 1852 

Aphodius semiruber Motschulsky, 1860 

Aphodius niger Cockerell, 1888 

Aphodius mundus Reitter, 1892 

Aphodius rufoplagiatus Reitter, 1892 

Aphodius vitiosus Reitter, 1892 

Aphodius cockerelli Leng, 1918 

Aphodius tjanshanicus Balthasar, 1956 

Planolinoides Dellacasa and Dellacasa, 2005 

Planolinoides aenictus (Cooper and Gordon, 1987) (Aphodius) 

Planolinoides borealis (Gyllenhal, 1827) (Aphodius) 

Aphodius errans Brown, 1930 

Planolinoides duplex (LeConte, 1878) (Aphodius) 

Planolinoides neotomae (Fall, 1907) (Aphodius) 

Planolinoides pectoralis (LeConte, 1857) (Aphodius) 

Planolinus Mulsant and Rey, 1870 

Planolinus tenellus (Say, 1823) (Aphodius) 

Pseudagolius Schmidt, 1913 

Pseudagolius bicolor (Say, 1823) (Aphodius) 

Aphodius comanchi Robinson, 1940 

Pseudagolius coloradensis (Horn, 1870) (Aphodius) 

Aphodius flohri Bates, 1887 

Pseudagolius warneri Gordon and Skelley, 2007 (Pseudagolius) 

Rugaphodius Gordon and Skelley, 2007 

Rugaphodius rugatus (Schmidt, 1907) (Aphodius) 

Aphodius rugifrons Horn, 1871 

Scabrostomus Gordon and Skelley, 2007 

Scabrostomus baileyi (Skelley and Gordon, 2002) (Aphodius) 

28

Scabrostomus bakeri (Skelley and Gordon, 2002) (Aphodius) 

Scabrostomus dyspistus (Skelley and Woodruff, 1991) (Aphodius) 

Scabrostomus kerni Gordon and Skelley, 2007 (Scabrostomus) 

Scabrostomus parapeculiosus Gordon and Skelley, 2007 (Scabrostomus) 

Scabrostomus peculiosus (Schmidt, 1922) (Aphodius) 

Aphodius longitarsis Fall, 1907 

Aphodius awemeanus Brown, 1928 

Scabrostomus sepultus (Cartwright, 1944) (Aphodius) 

Scabrostomus siccatus Gordon and Skelley, 2007 (Scabrostomus) 

Scabrostomus skillmani (Warner and Skelley, 2006) (Aphodius) 

Scabrostomus tanytarsus (Skelley and Woodruff, 1991) (Aphodius) 

Schaefferellus Gordon and Skelley, 2007 

Schaefferellus arizonensis (Schaeffer, 1907) (Aphodius) 

Setodius Gordon and Skelley, 2007 

Setodius bryanti (Brown, 1929) (Aphodius) 

Setodius diogenes Gordon and Skelley, 2007 (Setodius) 

Aphodius suspectus Mann, 1911 

Setodius edmontonus (Brown, 1929) (Aphodius) 

Stenotothorax Schmidt, 1913 

Stenobronchus Schmidt, 1916 

Stenotothorax badipes (Melsheimer, 1845) (Aphodius) 

Aphodius oblongus Say, 1823 

Stenotothorax cadaverinus (Mannerheim, 1843) (Oxyomus) 

Stenotothorax comosus Gordon and Skelley, 2007 (Stenotothorax) 

Stenotothorax cribratulus (Schmidt, 1922) (Aphodius) 

Aphodius cribratus LeConte, 1878 

Stenotothorax dilaticollis (Saylor, 1935) (Aphodius) 

Stenotothorax gardneri (Gordon, 2006) (Aphodius) 

Stenotothorax gentilis (Horn, 1887) (Aphodius) 

Stenotothorax gravis (Fall, 1927) (Aphodius) 

Stenotothorax lanei (Saylor, 1940) (Aphodius) 

Aphodius caseyi Saylor, 1940 

Aphodius pullmani Saylor, 1940 

Stenotothorax linsleyi (Van Dyke, 1933) (Aphodius) 

Stenotothorax martini (Van Dyke, 1928) (Aphodius) 

Stenotothorax mcpeaki Gordon and Skelley, 2007 (Stenotothorax) 

Stenotothorax micellus Gordon and Skelley, 2007 (Stenotothorax) 

Stenotothorax nevadensis (Horn, 1870) (Aphodius) 

Stenotothorax oriens Gordon and Skelley, 2007 (Stenotothorax) 

Stenotothorax oviformis (Robinson, 1946) (Aphodius) 

29

Aphodius inyoensis Van Dyke, 1947 

Stenotothorax ovipennis (Horn, 1870) (Aphodius) 

Stenotothorax parapyriformis Gordon and Skelley, 2007 (Stenotothorax) 

Stenotothorax pyriformis (Brown, 1929) (Aphodius) 

Stenotothorax rugoclypeus (Hinton, 1934) (Aphodius) 

Stenotothorax sparsus (LeConte, 1878) (Aphodius) 

Aphodius blaisdelli Fall, 1905 

Aphodius sheldoni Barrett, 1932 

Stenotothorax washtucna (Robinson, 1938) (Aphodius) 

Aphodius constrictus Robinson, 1946 

Stenotothorax woodleyi Gordon, 2006 (Stenotothorax) 

Strigodius Gordon and Skelley, 2007 

Strigodius robinsoni (Cartwright, 1939) (Aphodius) 

Tetraclipeoides Schmidt, 1913 

Tetraclipeoides acutissimus (Gordon, 1976) (Aphodius) 

Tetraclipeoides aemulus (Horn, 1887) (Aphodius) 

Aphodius fortunus Fall, 1932 

Tetraclipeoides anomaliceps (Brown, 1929) (Aphodius) 

Tetraclipeoides denticulatus (Haldeman, 1848) (Aphodius) 

Tetraclipeoides dentiger (LeConte, 1858) (Aphodius) 

Tetraclipeoides dentigerulus (Brown, 1929) (Aphodius) 

Tetraclipeoides fimbripes (Brown, 1928) (Aphodius) 

Tetraclipeoides formidatus (Gordon, 1976) (Aphodius) 

Tetraclipeoides giulianii (Gordon, 1977) (Aphodius) 

Tetraclipeoides henryi (Gordon, 1976) (Aphodius) 

Tetraclipeoides moquinus (Fall, 1907) (Aphodius) 

Tetraclipeoides testaceiventris (Fall, 1932) (Aphodius) 

Tetraclipeoides texanus Gordon and Skelley, 2007 (Tetraclipeoides) 

Teuchestes Mulsant, 1842 

Teuchestes fossor (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus triangulus Piller and Mitterpacher, 1783 

Teuchestes brunneus Mulsant, 1842 

Trichonotulus Bedel, 1911 

Trichonotus Mulsant, 1842 

Trichonotulus scrofa (Fabricius, 1787) (Scarabaeus) 

Scarabaeus minutus Herbst, 1789 

Scarabaeus tomentosus Kugelann, 1792 

Scarabaeus fuscus Rossi, 1792 

30

Trichonotus setiger Mulsant, 1842 

Aphodius stercoris Gistel, 1857 

Aphodius cinereus Mulsant and Rey, 1870 

Xeropsamobeus Saylor, 1937 

Xeropsamobeus acerbus (Horn, 1887) (Aphodius) 

Xeropsamobeus ambiguus (Fall, 1907) (Psammodius) 

Xeropsamobeus arenicolus Gordon and Skelley, 2007 (Xeropsamobeus) 

Xeropsamobeus asellus (Schmidt, 1907) (Aphodius) 

Aphodius nanus Horn, 1887 

Xeropsamobeus brighti Gordon and Skelley, 2007 (Xeropsamobeus) 

Xeropsamobeus desertus (Van Dyke, 1918) (Psammodius) 

Xeropsamobeus doyeni Gordon and Skelley, 2007 (Xeropsamobeus) 

Xeropsamobeus mohavei Gordon and Skelley, 2007 (Xeropsamobeus) 

Xeropsamobeus padrei Gordon and Skelley, 2007 (Xeropsamobeus) 

Xeropsamobeus scabriceps (LeConte, 1878) (Aphodius) 

DIDACTYLIINI Pittino, 1985 

Aidophus Balthasar, 1963 

Aidophus parcus (Horn, 1887) (Aphodius) 

Aphodius kansanus Wickham, 1905 

Aphodius knausii Fall, 1905 

Aphodius wolcotti Blatchley, 1910 

Aidophus skelleyi Harpootlian and Gordon, 2002 (Aidophus) 

EUPARIINI Schmidt, 1910 

Annegialia Howden, 1971 

Annegialia ataeniformis Howden, 1971 (Annegialia) 

Aphotaenius Cartwright, 1952 

Aphotaenius carolinus (Van Dyke, 1928) (Ataenius) 

Aphotaenius howdeni Cartwright, 1963 (Aphotaenius) 

Ataeniopsis Petrovitz, 1973 

Ataeniopsis duncani (Cartwright, 1974) (Ataenius) 

Ataeniopsis edistoi (Cartwright, 1974) (Ataenius) 

Ataeniopsis figurator (Harold, 1874) (Ataenius) 

Ataeniopsis parkeri (Cartwright, 1974) (Ataenius) 

Ataeniopsis rugopygus (Cartwright, 1974) (Ataenius) 

31

Ataeniopsis saxatilis (Cartwright, 1944) (Ataenius) 

Ataenius Harold, 1867 

Aphodinus Motschulsky, 1862 

Hexalus Mulsant and Rey, 1870 

Pseudammoecius Schmidt, 1913 

Placopterus Chalumeau, 1983 

Ataenius abditus (Haldeman, 1848) (Aphodius) 

Ataenius aequalis Harold, 1880 (Ataenius) 

Ataenius insulicola Chapin, 1940 

Ataenius titschacki Balthasar, 1941 

Ataenius alternatus (Melsheimer, 1845) (Oxyomus) 

Ataenius apicalis Hinton, 1937 (Ataenius) 

Ataenius barberi Cartwright, 1974 (Ataenius) 

Ataenius brevicollis (Wollaston, 1854) (Oxyomus) 

Auperia sulcatula Chevrolat, 1864 

Ataenius frankenbergeri Balthasar, 1938 

Ataenius lherminieri Paulian, 1947 

Ataenius brevis Fall, 1930 (Ataenius) 

Ataenius californicus Horn, 1887 (Ataenius) 

Ataenius carinator Harold, 1874 (Ataenius) 

Ataenius vincentiae Arrow, 1903 

Ataenius castaniellus Bates, 1887 (Ataenius) 

Ataenius cognatus (LeConte, 1858) (Euparia) 

Ataenius confertus Fall, 1909 (Ataenius) 

Ataenius cribratus Van Dyke, 1928 

Ataenius crenulatus Schmidt, 1910 (Ataenius) 

Ataenius rickardsi Hinton, 1938 

Ataenius cribrithorax Bates, 1887 (Ataenius) 

Ataenius cylindrus Horn, 1871 (Ataenius) 

Ataenius horni Harold, 1874 

Ataenius desertus Horn, 1871 (Ataenius) 

Ataenius ecruensis Stebnicka and Lago, 2005 (Ataenius) 

Ataenius erratus Fall, 1930 (Ataenius) 

Ataenius exiguus Brown, 1932 (Ataenius) 

Ataenius fattigi Cartwright, 1948 (Ataenius) 

Ataenius glaseri Cartwright, 1974 (Ataenius) 

Ataenius gracilis (Melsheimer, 1845) (Oxyomus) 

Saprosites nocturnus Nomura, 1943 

Ataenius heinekeni (Wollaston, 1854) (Oxyomus) 

Auperia rhyticephalus Chevrolat, 1864 

Ataenius vexator Harold, 1869 

Ataenius solitarius Blatchley, 1928 

Ataenius floridanus Brown, 1930 

Ataenius hesperius Cartwright, 1974 (Ataenius) 

32

Ataenius hirsutus Horn, 1871 (Ataenius) 

Ataenius imbricatus (Melsheimer, 1845) (Aphodius) 

Ataenius sordidus Harold, 1869 

Ataenius inquisitus Horn, 1887 (Ataenius) 

Ataenius insculptus Horn, 1887 (Ataenius) 

Ataenius languidus Schmidt, 1911 (Ataenius) 

Ataenius linelli Cartwright, 1944 

Ataenius liogaster Bates, 1887 (Ataenius) 

Ataenius orbicularis Schmidt, 1914 

Ataenius edwardsi Chapin, 1940 

Ataenius nitidulus Nomura, 1943 

Ataenius kelatianus Balthasar, 1965 

Ataenius hoguei Cartwright and Spangler, 1981 

Ataenius lobatus Horn, 1871 (Ataenius) 

Ataenius laeviventris Horn, 1887 

Ataenius miamii Cartwright, 1934 (Ataenius) 

Ataenius oklahomensis Brown, 1930 (Ataenius) 

Ataenius opatrinus Harold, 1867 (Ataenius) 

Ataenius narialis Petrovitz, 1973 

Ataenius woodruffi Cartwright, 1974 

Ataenius paraperforatus Deloya and Ibéñez-Bernal, 2000 

Ataenius ovatulus Horn, 1871 (Ataenius) 

Ataenius lecontei Harold, 1874 

Ataenius peregrinator Harold, 1877 (Ataenius) 

Ataenius hieronymi Bates, 1887 

Ataenius jalapensis Bates, 1887 

Ataenius nunenmacheri Cartwright, 1974 

Ataenius stroheckeri Cartwright, 1974 

Ataenius picinus Harold, 1867 (Ataenius) 

Ataenius duplopunctatus Lea, 1923 

Ataenius salutator Fall, 1930 

Ataenius queirosii Paulian, 1934 

Ataenius darlingtoni Hinton, 1937 

Ataenius boucomontii Paulian, 1937 

Ataenius alegrus Balthasar, 1947 

Saprosites rugosus Richards, 1959 

Ataenius paracognatus Balthasar, 1961 

Ataenius platensis (Blanchard, 1847) (Ataenius) 

Ataenius integer Harold, 1868 (Ataenius) 

Ataenius anticus Fall, 1930 

Ataenius granchaocensis Balthasar, 1939 

Ataenius histrionicus Balthasar, 1947 

Ataenius heyrovskyi Balthasar, 1960 

Ataenius degallieri Chalumeau, 1990 

Ataenius robustus Horn, 1871 (Ataenius) 

Ataenius scabrelloides Petrovitz, 1962 (Ataenius) 

33

Ataenius superficialis Cartwright, 1974 

Ataenius scabrellus Schmidt, 1909 (Ataenius) 

Ataenius havanensis Balthasar, 1938 

Ataenius fleutiauxi Paulian, 1947 

Ataenius sculptor Harold, 1868 (Ataenius) 

Ataenius oblongus Horn, 1871 

Ataenius setiger Bates, 1887 (Ataenius) 

Ataenius pseudohirsutus Cartwright, 1974 

Ataenius spretulus (Haldeman, 1848) (Aphodius) 

Ataenius consors Fall, 1930 

Ataenius falli Hinton, 1934 

Ataenius stephani Cartwright, 1974 (Ataenius) 

Ataenius strigatus (Say, 1823) (Aphodius) 

Ataenius texanus Harold, 1874 (Ataenius) 

Ataenius punctifrons Cartwright, 1974 

Ataenius utahensis Cartwright, 1974 (Ataenius) 

Ataenius vandykei Cartwright, 1974 (Ataenius) 

Ataenius waltherhorni Balthasar, 1938 (Ataenius) 

Ataenius wenzelii Horn, 1887 (Ataenius) 

Ataenius ludovicianus Fall, 1930 

Ataenius rudellus Fall, 1930 

Auperia Jacquelin du Val, 1856 

Phalangochaeta Martínez, 1952 

Auperia denominata Chevrolat, 1864 (Auperia) 

Ataenius arator Harold, 1869 

Ataenius euglyptus Bates, 1887 

Ataenius brevinotus Chapin, 1940 

Ataenius benjaminbanderai Islas, 1955 

Ataenius sciurus Cartwright, 1974 

Phalangochaeta grandis Petrovitz, 1973 

Euparia LePeletier and Serville, 1828 

Euparia castanea LePeletier and Serville, 1828 (Euparia) 

Euparixia Brown, 1927 

Euparixia duncani Brown, 1927 (Euparixia) 

Euparixia mexicana Gordon and McCleve, 2003 (Euparixia) 

Euparixia moseri Woodruff and Cartwright, 1967 (Euparixia) 

Haroldiataenius Chalumeau, 1981 

Haroldiataenius Chalumeau, 1981 

34

Sayloria Chalumeau, 1981 

Haroldiataenius buvexus Stebnicka and Skelley, 2009 (Haroldiataenius) 

Haroldiataenius convexus (Robinson, 1940) (Ataenius) 

Haroldiataenius griffini (Cartwright, 1974) (Ataenius) 

Haroldiataenius hintoni (Saylor, 1933) (Ataenius) 

Haroldiataenius limbatus (Bates, 1887) (Ataenius) 

Haroldiataenius lucanus (Horn, 1871) (Ataenius) 

Ataenius sabinoi Cartwright, 1974 

Haroldiataenius mariarum (Bates, 1887) (Ataenius) 

Psammodius perpolitus Schmidt, 1916 

Haroldiataenius saramari (Cartwright, 1939) (Ataenius) 

Haroldiataenius semipilosus (Van Dyke, 1928) (Ataenius) 

Martineziana Chalumeau and Özdikmen, 2006 

Martineziella Chalumeau, 1987 

Martinezia Chalumeau, 1983 

Martineziana dutertrei (Chalumeau, 1983) (Martinezia) 

Martineziana vandykei (Hinton, 1936) (Euparia) 

Parataenius Balthasar, 1961 

Brancoataenius Paulian, 1979 

Parataenius simulator (Harold, 1868) (Ataenius) 

Psammodius schwarzi Linell, 1896 

Parataenius granuliceps Petrovitz, 1971 

Ataenius lusitanicus Paulian, 1979 

Pseudataenius Brown, 1927 

Ataenioides Petrovitz, 1973 

Pseudataenius contortus Cartwright, 1974 (Pseudataenius) 

Pseudataenius socialis (Horn, 1871) (Ataenius) 

PROCTOPHANINI Stebnicka and Howden, 1995 

Australaphodius Balthasar, 1942 

Phaeaphodius Schmidt, 1922 

Australaphodius frenchi (Blackburn, 1892) (Aphodius) 

Aphodius ambiguus Boheman, 1858 

Aphodius brevitarsis Peringuey, 1901 

Aphodius tarsalis Schmidt, 1907 

Aphodius melbournicus Balthasar, 1942 

Aphodius catulus Balthasar, 1946 

35

PSAMMODIINI Mulsant, 1842 

Geopsammodius Gordon and Pittino, 1992 

Geopsammodius fuscus Skelley, 2006 (Geopsammodius) 

Geopsammodius hydropicus (Horn, 1887) (Psammodius) 

Geopsammodius morrisi Skelley, 2006 (Geopsammodius) 

Geopsammodius ohoopee Skelley, 2006 (Geopsammodius) 

Geopsammodius relictillus (Deyrup and Woodruff, 1991) (Psammodius) 

Geopsammodius rileyi Skelley, 2006 (Geopsammodius) 

Geopsammodius subpedalis Skelley, 2006 (Geopsammodius) 

Geopsammodius unsidensis Skelley, 2006 (Geopsammodius) 

Geopsammodius withlacoochee Skelley, 2006 (Geopsammodius) 

Leiopsammodius Rakovic, 1981 

Leiopsammodius acei Harpootlian, Gordon, and Woodruff, 2000 (Leiopsammodius) 

Leiopsammodius deyrupi Harpootlian, Gordon, and Woodruff, 2000 (Leiopsammodius) 

Leiopsammodius malkini (Cartwright, 1946) (Psammodius) 

Leiopsammodius ocmulgeei Harpootlian, Gordon, and Woodruff, 2000 (Leiopsammodius) 

Neopsammodius Rakovic, 1986 

Neopsammodius blandus (Fall, 1932) (Psammodius) 

Neopsammodius interruptus (Say, 1835) (Psammodius) 

Neopsammodius mimeticus (Fall, 1932) (Psammodius) 

Neopsammodius quinqueplicatus (Horn, 1871) (Psammodius) 

Neopsammodius saltilloensis (Cartwright, 1955) (Psammodius) 

Neopsammodius werneri (Cartwright, 1955) (Psammodius) 

Odontopsammodius Gordon and Pittino, 1992 

Odontopsammodius armaticeps (Fall, 1932) (Psammobius) 

Odontopsammodius cruentus (Harold, 1867) (Psammodius) 

Psammodius shermani Cartwright, 1946 

Parapsammodius Gordon and Pittino, 1992 

Parapsammodius bidens (Horn, 1871) (Psammodius) 

Parapsammodius puncticollis (LeConte, 1858) (Euparia) 

Ataenius inops Horn, 1887 

Platytomus Mulsant, 1842 

Platytomus atlanticus (Cartwright, 1948) (Pleurophorus) 

36

Platytomus caelicollis (Cartwright, 1948) (Pleurophorus) 

Platytomus longulus (Cartwright, 1948) (Pleurophorus) 

Platytomus micros (Bates, 1887) (Psammodius) 

Pleurophorus batesi Arrow, 1903 

Diastictus californicus Petrovitz 1963 

Platytomus notialis (Cartwright, 1948) (Pleurophorus) 

Platytomus parvulus (Chevrolat, 1864) (Psammodius) 

Platytomus tibialis (Fabricius, 1798) (Scarabaeus) 

Psammodius grundeli Van Dyke, 1918 

Pleurophorus Mulsant, 1842 

Pleurophorus caesus (Panzer, 1796) (Scarabaeus) 

Aphodius angustus Philippi and Philippi, 1864 

Ataenius rugiceps Dury, 1902 

Psammodius Fallén, 1807 

Psammobius Heer, 1841 

Psammodius basalis (Mulsant and Rey, 1870) (Psammobius) 

Psammodius laevipennis Costa, 1844 (Psammodius) 

Psammodius aegialioides Haldeman, 1848 

Psammodius pierottii Pittino, 1979 (Psammodius) 

Rhyssemus Mulsant, 1842 

Boucardius Petrovitz, 1967 

Rhyssemus brownwoodi Gordon and Cartwright, 1980 (Rhyssemus) 

Rhyssemus californicus Horn, 1871 (Rhyssemus) 

Rhyssemus germanus (Linnaeus, 1767) (Ptinus) 

Scarabaeus asper Fabricius, 1775 

Rhyssemus aspericeps Chevrolat, 1861 

Rhyssemus obsoletus Rey, 1890 

Rhyssemus parallelus Reitter, 1892 

Rhyssemus puncticollis Brown, 1929 

Rhyssemus neglectus Brown, 1929 (Rhyssemus) 

Rhyssemus scaber Haldeman, 1848 (Rhyssemus) 

Rhyssemus sonatus LeConte, 1881 (Rhyssemus) 

Tesarius Rakovic, 1981 

Tesarius caelatus (LeConte, 1857) (Aegialia) 

Tesarius doyeni (Cartwright, 1977) (Psammodius) 

Tesarius mcclayi (Cartwright, 1955) (Psammodius) 

Tesarius oregonensis (Cartwright, 1955) (Psammodius) 

37

Trichiorhyssemus Clouët, 1901 

Trichiorhyssemus alternatus Hinton, 1938 (Trichiorhyssemus) 

Trichiorhyssemus cristatellus (Bates, 1887) (Rhyssemus) 

Trichiorhyssemus riparius (Horn, 1871) (Rhyssemus) 

Rhyssemus caelatus LeConte, 1881 

SCARABAEINAE Latreille, 1802 

CANTHONINI van Lansberge, 1874 

Canthon Hoffmannsegg, 1817 

Canthon (Boreocanthon) Halffter, 1958 

Canthon (Canthon) Hoffmannsegg, 1817 

Coprobius Latreille, 1829 

Coeloscelis Reiche, 1841 

Paedhyboma Kolbe, 1893 

Canthomoechus Pereira and Martínez, 1960 

Canthon (Glaphyrocanthon) Martínez, 1948 

Coprocanthon Martínez, 1948 

Geocanthon Pereira and Martínez, 1956 

Canthon bisignatus Balthasar, 1939 (Canthon) 

Canthon (Canthon) chalcites (Haldeman, 1843) (Coprobius) 

Canthon (Boreocanthon) coahuilensis Howden, 1966 (Canthon) 

Canthon (Canthon) cyanellus LeConte, 1859 (Canthon) 

Canthon (Boreocanthon) depressipennis LeConte, 1859 (Canthon) 

Canthon (Boreocanthon) ebenus (Say, 1823) (Ateuchus) 

Canthon (Glaphyrocanthon) femoralis (Chevrolat, 1834) (Coprobius) 

Canthon balsensis Martínez and Halffter, 1972 

Canthon (Boreocanthon) forreri Bates, 1887 (Canthon) 

Canthon (Canthon) humectus alvarengai Halffter, 1961 (Canthon) 

Canthon (Canthon) humectus blumei Halffter and Halffter, 2003 (Canthon) 

Canthon (Canthon) humectus hidalgoensis Bates, 1887 (Canthon) 

Canthon (Canthon) humectus humectus (Say, 1832) (Ateuchus) 

Canthon gagatinus Harold, 1863 

Canthon amethystinum Harold, 1863 

Canthon (Canthon) imitator Brown, 1946 (Canthon) 

Canthon laevis Bates, 1887 

Canthon floridanus Brown, 1946 

Canthon (Canthon) indigaceus LeConte, 1866 (Canthon) 

Canthon chevrolati Harold, 1868 

Canthon (Boreocanthon) lecontei Harold, 1868 (Canthon) 

Canthon (Boreocanthon) melanus Robinson, 1948 (Canthon) 

Canthon (Boreocanthon) mixtus Robinson, 1948 (Canthon) 

Canthon (Canthon) obliquus Horn, 1894 (Canthon) 

38

Canthon (Canthon) pilularius (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus laevis Drury, 1773 

Scarabaeus hudsonias Forster, 1771 

Scarabaeus volvens Fabricius, 1792 

Coprobius obtusidens Ziegler, 1844 

Canthon viridescens Horn, 1870 

Canthon (Boreocanthon) praticola LeConte, 1859 (Canthon) 

Canthon vetustus Pierce, 1946 

Canthon (Boreocanthon) probus (Germar, 1824) (Ateuchus) 

Canthon minor Sturm, 1843 

Canthon abrasus LeConte, 1859 

Canthon (Boreocanthon) puncticollis LeConte, 1866 (Canthon) 

Canthon nyctelius Bates, 1887 

Canthon integricollis Schaeffer, 1915 

Canthon (Boreocanthon) simplex LeConte, 1857 (Canthon) 

Canthon corvinus Harold, 1868 

Canthon militaris Horn, 1870 

Canthon humeralis Horn, 1870 

Canthon antiquus Pierce, 1946 

Canthon (Canthon) vigilans LeConte, 1858 (Canthon) 

Canthon (Glaphyrocanthon) viridis (Palisot de Beauvois, 1805) (Copris) 

Ateuchus obsoletus Say, 1823 

Onthophagus viridicatus Say, 1835 

Canthon metallicus Sturm, 1843 

Deltochilum Eschscholtz, 1822 

Anamnesis Vigors, 1826 

Hyboma LePeletier and Serville, 1828 

Meghyboma Kolbe, 1893 

Deltochilum gibbosum gibbosum (Fabricius, 1775) (Scarabaeus) 

Deltochilum scabriusculum montanum Howden, 1966 (Deltochilum) 

Deltochilum scabriusculum scabriusculum Bates, 1887 (Deltochilum) 

Deltochilum tumidum Howden, 1966 (Deltochilum) 

Malagoniella Martínez, 1961 

Malagoniella astyanax yucateca (Harold, 1863) (Megathopa) 

Melanocanthon Halffter, 1958 

Melanocanthon bispinatus (Robinson, 1941) (Canthon) 

Melanocanthon granulifer (Schmidt, 1920) (Canthon) 

Melanocanthon nigricornis (Say, 1823) (Ateuchus) 

Melanocanthon punctaticollis (Schaeffer, 1915) (Canthon) 

39

Pseudocanthon Bates, 1887 

Opiocanthon Paulian, 1947 

Pseudocanthon perplexus (LeConte, 1847) (Canthon) 

COPRINI Leach, 1815 

Canthidium Erichson, 1847 

Canthidium (Eucanthidium) Martínez and Halffter, 1986 

Canthidium (Eucanthidium) laetum Harold, 1867 (Canthidium) 

Canthidium (Eucanthidium) macclevei Kohlmann and Solís, 2006 (Canthidium) 

Canthidium (Eucanthidium) pseudopuncticolle Solís and Kohlmann, 2004 (Canthidium) 

Copris Geoffroy, 1762 

Litocopris Waterhouse, 1891 

Paracopris Balthasar, 1939 

Microcopris Balthasar, 1958 

Copris arizonensis Schaeffer, 1906 (Copris) 

Copris fricator (Fabricius, 1787) (Scarabaeus) 

Scarabaeus tullius Olivier, 1789 

Copris anaglypticus Say, 1823 

Copris cartwrighti Robinson, 1941 

Copris gopheri Hubbard, 1894 (Copris) 

Copris incertus Say, 1835 (Copris) 

Copris procidua Say, 1835 

Copris inemarginatus Blatchley, 1918 (Copris) 

Copris howdeni Matthews and Halffter, 1959 (Copris) 

Copris klugi sierrensis Matthews, 1962 (Copris) 

Copris laeviceps Harold, 1862 (Copris) 

Copris lecontei lecontei Matthews, 1962 (Copris) 

Copris lugubris Boheman, 1858 (Copris) 

Copris macclevei Warner, 1990 (Copris) 

Copris martinezi Matthews and Halffter, 1968 (Copris) 

Copris minutus (Drury, 1773) (Scarabaeus) 

Scarabaeus silenus Fabricius, 1775 

Scarabaeus ammon Fabricius, 1781 

Scarabaeus lar Fabricius, 1787 

Copris reflexus Panzer, 1794 

Copris moechus LeConte, 1854 (Copris) 

Copris clavicornis Matthews and Halffter, 1959 

Copris remotus dicyrtus Matthews and Halffter, 1959 (Copris) 

Copris remotus remotus LeConte, 1866 (Copris) 

40

Dichotomius Hope, 1838 

Homocopris Burmeister, 1842 

Brachycopris Haldeman, 1846 

Pinotus Erichson, 1847 

Dichotomius carolinus (Linnaeus, 1767) (Copris) 

Dichotomius colonicus (Say, 1835) (Copris) 

Ontherus Erichson, 1847 

Ontherus (Caelontherus) Génier, 1996 

Ontherus (Caelontherus) mexicanus Harold, 1868 (Ontherus) 

Uroxys Westwood, 1842 

Uroxys deavilai Delgado and Kohlmann, 2007 (Uroxys) 

ATEUCHINI Laporte, 1840 

Ateuchus Weber, 1801 

Choeridium LePeletier and Serville, 1828 

Ateuchus floridensis Génier, 2000 (Ateuchus) 

Ateuchus histeroides Weber, 1801 (Ateuchus) 

Ateuchus capistratus Fabricius, 1801 

Choeridium punctatus Robinson, 1948 

Ateuchus illaesum (Harold, 1868) (Choeridium) 

Ateuchus lecontei (Harold, 1868) (Choeridium) 

Ateuchus rodriguezi (Preudhomme de Borre, 1886) (Uroxys) 

Ateuchus texanus (Robinson, 1948) (Choeridium) 

ONITICELLINI Kolbe, 1905 

Euoniticellus Janssens, 1953 

Euoniticellus cubiensis (Laporte, 1840) (Oniticellus) 

Euoniticellus intermedius (Reiche, 1849) (Oniticellus) 

Liatongus Reitter, 1893 

Liatongus californicus (Horn, 1882) (Oniticellus) 

Oniticellus LePeletier and Serville, 1828 

Oniticellus rhinocerulus Bates, 1889 (Oniticellus) 

41

ONITINI Laporte, 1840 

Onitis Fabricius, 1798 

Onitis alexis Klug, 1835 (Onitis) 

ONTHOPHAGINI Burmeister, 1846 

Onthophagus Latreille, 1802 

Onthophagus (Onthophagus) Latreille, 1802 

Chalcoderus Erichson, 1848 

Monapus Erichson, 1848 

Psilax Erichson, 1848 

Gonocyphus Lansberge, 1885 

Tauronthophagus Shipp, 1895 

Onthophagus (Digitonthophagus) Balthasar, 1959 

Onthophagus (Onthophagus) aciculatulus Blatchley, 1928 (Onthophagus) 

Onthophagus alutaceus Blatchley, 1916 

Onthophagus (Onthophagus) alluvius Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) antivagans Howden and Génier, 2004 (Onthophagus) 

Onthophagus (Onthophagus) arnetti Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) batesi Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) brevifrons Horn, 1881 (Onthophagus) 

Onthophagus (Onthophagus) browni Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) canelasensis Howden and Génier, 2004 (Onthophagus) 

Onthophagus (Onthophagus) cartwrighti Howden, 1973 (Onthophagus) 

Onthophagus (Onthophagus) cavernicollis Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) chevrolati longecarinatus Zunino and Halffter, 1988 (Onthophagus) 

Onthophagus (Onthophagus) coahuilae Zunino and Halffter, 1988 (Onthophagus) 

Onthophagus (Onthophagus) cochisus Brown, 1927 (Onthophagus) 

Onthophagus (Onthophagus) concinnus Laporte, 1840 (Onthophagus) 

Onthophagus protensus Melsheimer, 1845 

Onthophagus (Onthophagus) coproides Horn, 1881 (Onthophagus) 

Onthophagus cuboidalis Bates, 1887 

Onthophagus (Onthophagus) cuevensis Howden, 1973 (Onthophagus) 

Onthophagus (Onthophagus) cyanellus Bates, 1887 (Onthophagus) 

Onthophagus (Onthophagus) cynomysi Brown, 1927 (Onthophagus) 

Onthophagus (Onthophagus) depressus Harold, 1871 (Onthophagus) 

Onthophagus carteri Blackburn, 1904 

Onthophagus (Onthophagus) fuscus fuscus Boucomont, 1932 (Onthophagus) 

Onthophagus (Digitonthophagus) gazella (Fabricius, 1787) (Scarabaeus) 

Scarabaeus dorcas Olivier, 1789 

Copris antilope Fabricius, 1798 

42

Copris metallicus Fabricius, 1798 

Onthophagus intermedius Reiche, 1840 

Onthophagus (Onthophagus) hecate blatchleyi Brown, 1929 (Onthophagus) 

Onthophagus (Onthophagus) hecate hecate (Panzer, 1794) (Scarabaeus) 

Copris hastator Fabricius, 1798 

Copris latebrosus Fabricius, 1801 

Copris obtectus Palisot de Beauvois, 1805 

Onthophagus scabricollis Kirby, 1837 

Onthophagus sayi Laporte, 1840 

Onthophagus (Onthophagus) hidalgus Zunino and Halffter, 1988 (Onthophagus) 

Onthophagus (Onthophagus) hoepfneri Harold, 1869 (Onthophagus) 

Onthophagus arizonensis Schaeffer, 1909 

Onthophagus (Onthophagus) incensus Say, 1835 (Onthophagus) 

Onthophagus curvicornis Latreille, 1812 

Onthophagus (Onthophagus) knausi Brown, 1927 (Onthophagus) 

Onthophagus (Onthophagus) knulli Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) landolti landolti Harold, 1880 (Onthophagus) 

Onthophagus (Onthophagus) landolti texanus Schaeffer, 1914 (Onthophagus) 

Onthophagus (Onthophagus) lecontei Harold, 1871 (Onthophagus) 

Onthophagus (Onthophagus) mcclevei Howden and Génier, 2004 (Onthophagus) 

Onthophagus (Onthophagus) medorensis Brown, 1929 (Onthophagus) 

Onthophagus (Onthophagus) mextexus Howden and Cartwright, 1970 (Onthophagus) 

Onthophagus monticolus Howden and Cartwright, 1963 

Onthophagus (Onthophagus) nuchicornis (Linnaeus, 1758) (Scarabaeus) 

Onthophagus rhinoceros Melsheimer, 1845 

Onthophagus (Onthophagus) oklahomensis Brown, 1927 (Onthophagus) 

Onthophagus (Onthophagus) orpheus canadensis (Fabricius, 1801) (Copris) 

Onthophagus (Onthophagus) orpheus orpheus (Panzer, 1794) (Scarabaeus) 

Onthophagus (Onthophagus) orpheus pseudorpheus Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 (Onthophagus) 

Onthophagus falcipes Harold, 1871 

Onthophagus (Onthophagus) polyphemi polyphemi Hubbard, 1894 (Onthophagus) 

Onthophagus (Onthophagus) polyphemi sparsisetosus Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) pseudofuscus Zunino and Halffter, 1988 (Onthophagus) 

Onthophagus (Onthophagus) schaefferi Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) striatulus floridanus Blatchley, 1928 (Onthophagus) 

Onthophagus nigrescens Blatchley, 1916 

Onthophagus (Onthophagus) striatulus striatulus (Palisot de Beauvois, 1809) (Copris) 

Scarabaeus janus Panzer, 1794 

Onthophagus cervicornis Kirby, 1825 

Onthophagus castaneus Melsheimer, 1845 

Onthophagus viridicollis Harold, 1869 

Onthophagus scabricollis Horn, 1875 

Onthophagus canadensis Horn, 1875 

Onthophagus subaeneus Horn, 1875 

Onthophagus concinnus Horn, 1875 

43

Onthophagus protensus Horn, 1875 

Onthophagus (Onthophagus) subaeneus (Palisot de Beauvois, 1811) (Copris) 

Onthophagus cribricollis Horn, 1881 

Onthophagus (Onthophagus) subopacus Robinson, 1940 (Onthophagus) 

Onthophagus (Onthophagus) subtropicus Howden and Cartwright, 1963 (Onthophagus) 

Onthophagus (Onthophagus) taurus (Schreber, 1759) (Scarabaeus) 

Scarabaeus rugosus Poda von Neuhaus, 1761 

Copris corniger Geoffroy, 1785 

Scarabaeus recticornis Leske, 1785 

Scarabaeus capra Fabricius, 1787 

Scarabaeus quadrum Kugelann, 1792 

Scarabaeus cruoreus Schrank, 1798 

Onthophagus morio Brullé, 1832 

Onthophagus bos Villa and Villa, 1833 

Onthophagus bovillus Mulsant, 1842 

Onthophagus capreolus Mulsant, 1842 

Onthophagus femineus Mulsant, 1842 

Onthophagus fuscipennis Mulsant, 1842 

Onthophagus mendax Mulsant, 1842 

Onthophagus nigrovirescens Mulsant, 1842 

Onthophagus glareanus Gistel, 1857 

Onthophagus alternans d’Orbigny, 1898 

Onthophagus alternatus d’Orbigny, 1898 

Onthophagus castanonotus Seabra, 1907 

Onthophagus saharae Bedel, 1911 

Onthophagus circumcinctus Escalera, 1914 

Onthophagus rubripennis Goidanich, 1924 

Onthophagus athenae Goidanich, 1925 

Onthophagus reichardti Medvedev, 1930 

Onthophagus (Onthophagus) tuberculifrons Harold, 1871 (Onthophagus) 

Onthophagus (Onthophagus) velutinus Horn, 1875 (Onthophagus) 

PHANAEINI Kolbe, 1905 

Coprophanaeus D’Olsoufieff, 1924 

Coprophanaeus (Coprophanaeus) D’Olsoufieff, 1924 

Coprophanaeus (Coprophanaeus) pluto noguerai Arnaud, 2002 (Coprophanaeus) 

Phanaeus MacLeay, 1819 

Phanaeus (Phanaeus) MacLeay, 1819 

Lonchophorus Germar, 1824 

Onthurgus Gistel, 1857 

Palaeocopris Pierce, 1946 

44

Phanaeus (Phanaeus) adonis Harold, 1863 (Phanaeus) 

Phanaeus (Phanaeus) amithaon Harold, 1875 (Phanaeus) 

Phanaeus (Phanaeus) amethystinus amethystinus Harold, 1863 (Phanaeus) 

Phanaeus martinezi Halffter, 1955 

Phanaeus (Phanaeus) difformis LeConte, 1847 (Phanaeus) 

Phanaeus magnificens Robinson, 1948 

Phanaeus (Phanaeus) flohri Nevinson, 1892 (Phanaeus) 

Phanaeus (Phanaeus) furiosus Bates, 1887 (Phanaeus) 

Phanaeus furcosus Felsche, 1901 

Phanaeus (Phanaeus) genieri Arnaud, 2001 (Phanaeus) 

Phanaeus (Phanaeus) igneus MacLeay, 1919 (Phanaeus) 

Phanaeus floridanus D’Olsoufieff, 1924 

Phanaeus (Phanaeus) palliatus Sturm, 1843 (Phanaeus) 

Phanaeus (Phanaeus) quadridens (Say, 1835) (Copris) 

Phanaeus violaceus Laporte, 1840 

Phanaeus laevipennis Sturm, 1843 

Phanaeus borealis D’Olsoufieff, 1924 

Phanaeus (Phanaeus) triangularis texensis Edmonds, 1994 (Phanaeus) 

Phanaeus (Phanaeus) triangularis triangularis (Say, 1823) (Copris) 

Phanaeus torrens LeConte, 1847 

Phanaeus niger D’Olsoufieff, 1924 

Phanaeus goidanichi Balthasar, 1939 

Phanaeus (Phanaeus) vindex MacLeay, 1819 (Phanaeus) 

Phanaeus cyanellus Robinson, 1938 

Phanaeus rubervirens Robinson, 1948 

Phanaeus (Phanaeus) yecoraensis Edmonds, 2004 (Phanaeus) 

SISYPHINI Mulsant, 1842 

Sisyphus Latreille, 1807 

Sisyphus submonticolus Howden, 1965 (Sisyphus) 

MELOLONTHINAE Leach, 1819 

ONCERINI LeConte, 1861 

Nefoncerus Saylor, 1938 

Nefoncerus convergens (Horn, 1894) (Oncerus) 

Oncerus LeConte, 1856 

Oncerus floralis LeConte, 1856 (Oncerus) 

PODOLASIINI Howden, 1997 

45

Podolasia Harold, 1869 

Lasiopus LeConte, 1856 

Podolasia emarginata Howden, 1954 (Podolasia) 

Podolasia ferruginea (LeConte, 1856) (Lasiopus) 

Podolasia involucris Howden, 1998 (Podolasia) 

Podolasia lavignei Howden, 1997 (Podolasia) 

Podolasia longipenis Howden, 1997 (Podolasia) 

Podolasia peninsularis Howden, 1954 (Podolasia) 

Podolasia saylori Howden, 1954 

Podolasia pilosa Howden, 1954 (Podolasia) 

Podolasia rotundipenis Howden, 1997 (Podolasia) 

Podolasia stillwellorum Howden, 1997 (Podolasia) 

Podolasia varicolor Saylor, 1948 (Podolasia) 

Podostena Howden, 1997 

Podostena bottimeri (Howden, 1958) (Podolasia) 

Podostena litoralis Howden, 1997 (Podostena) 

Podostena rileyi Howden, 1997 (Podostena) 

Podostena sleeperi Howden, 1997 (Podostena) 

SERICINI Kirby, 1837 

Maladera Mulsant and Rey, 1871 

Maladera (Cephaloserica) Brenske, 1900 

Maladera (Cephaloserica) castanea castanea (Arrow, 1913) (Autoserica) 

Serica korgei Petrovitz, 1967 

Nipponoserica Nomura, 1973 

Pseudomaladera Nikolajev, 1980 

Nipponoserica peregrina (Chapin, 1938) (Serica) 

Serica MacLeay, 1819 

Stilbolemma Harris, 1827 

Camptorhina Kirby, 1837 

Paramaladera Nikolajev, 1979 

Serica abdita Dawson, 1921 (Serica) 

Serica acicula Dawson, 1932 (Serica) 

Serica acontia Dawson, 1933 (Serica) 

Serica adversa Dawson, 1967 (Serica) 

46

Serica aemula Dawson, 1947 (Serica) 

Serica alabama Dawson, 1967 (Serica) 

Serica alleni Saylor, 1939 (Serica) 

Serica searli Saylor, 1939 

Serica alternata alternata LeConte, 1856 (Serica) 

Serica alternata exolita Dawson, 1933 (Serica) 

Serica alternata patruela Dawson, 1933 (Serica) 

Serica ammomenisco Hardy, 1987 (Serica) 

Serica anthracina LeConte, 1856 (Serica) 

Serica frontalis LeConte, 1856 

Serica robusta LeConte, 1856 

Serica valida Harold, 1869 

Serica crassata Walker, 1866 

Serica apatela Dawson, 1922 (Serica) 

Serica arkansana Dawson, 1947 (Serica) 

Serica aspera Dawson, 1922 (Serica) 

Serica atracapilla (Kirby, 1837) (Camptorhina) 

Serica cucullata Dawson, 1919 

Serica atratula atratula LeConte, 1856 (Serica) 

Serica atratula monita Dawson, 1947 (Serica) 

Serica aviceps Dawson, 1967 (Serica) 

Serica barri Dawson, 1967 (Serica) 

Serica blatchleyi Dawson, 1932 (Serica) 

Serica bruneri Dawson, 1967 (Serica) 

Serica caliginosa Dawson, 1932 (Serica) 

Serica campestris Dawson, 1919 (Serica) 

Serica carolina Dawson, 1920 (Serica) 

Serica catalina Dawson, 1947 (Serica) 

Serica chaetosoma Dawson, 1932 (Serica) 

Serica chicoensis Saylor, 1939 (Serica) 

Serica coalinga Dawson, 1952 (Serica) 

Serica concinna Dawson, 1947 (Serica) 

Serica contorta Dawson, 1947 (Serica) 

Serica craighead Saylor, 1939 (Serica) 

Serica craigheadi Dawson, 1952 

Serica cruzi Saylor, 1939 (Serica) 

Serica curvata LeConte, 1856 (Serica) 

Serica cuyamaca Saylor, 1939 (Serica) 

Serica delicata Dawson, 1922 (Serica) 

Serica deserticola Dawson, 1952 (Serica) 

Serica diablo Dawson, 1967 (Serica) 

Serica egregia Dawson, 1921 (Serica) 

Serica elmontea Saylor, 1939 (Serica) 

Serica elongatula Horn, 1870 (Serica) 

Serica elusa Dawson, 1919 (Serica) 

Serica ensenada Saylor, 1948 (Serica) 

47

Serica evidens Blatchley, 1919 (Serica) 

Serica carinata Blatchley, 1910 

Serica falcata Dawson, 1933 (Serica) 

Serica falli Dawson, 1932 (Serica) 

Serica fimbriata LeConte, 1856 (Serica) 

Serica michelbacheri Saylor, 1948 

Serica floridana Dawson, 1967 (Serica) 

Serica frosti Dawson, 1967 (Serica) 

Serica georgiana georgiana Leng, 1911 (Serica) 

Serica georgiana lecontei Dawson, 1921 (Serica) 

Serica heteracantha Dawson, 1967 (Serica) 

Serica howdeni Dawson, 1967 (Serica) 

Serica humboldti Gordon, 1975 (Serica) 

Serica imitans Chapin, 1931 (Serica) 

Omaloplia trogiformis Uhler, 1855 

Serica intermixta Blatchley, 1910 (Serica) 

Serica iricolor (Say, 1824) (Melolontha) 

Serica laguna Saylor, 1935 (Serica) 

Serica ligulata ligulata Dawson, 1932 (Serica) 

Serica ligulata praetermissa Dawson, 1932 (Serica) 

Serica lodingi Dawson, 1952 (Serica) 

Serica loxia Dawson, 1920 (Serica) 

Serica mckenziei Saylor, 1935 (Serica) 

Serica mixta LeConte, 1856 (Serica) 

Serica mystaca Dawson, 1922 (Serica) 

Serica ochrosoma Dawson, 1919 (Serica) 

Serica oliver Saylor, 1939 (Serica) 

Serica joaquinella Saylor, 1939 

Serica oliveri Dawson, 1952 

Serica opposita Dawson, 1921 (Serica) 

Serica panda Dawson, 1952 (Serica) 

Serica parallela Casey, 1884 (Serica) 

Serica parvula (Blanchard, 1850) (Omaloplia) 

Serica pavonia Dawson, 1932 (Serica) 

Serica peleca Dawson, 1952 (Serica) 

Serica perigonia eremicola Dawson, 1967 (Serica) 

Serica perigonia perigonia Dawson, 1920 (Serica) 

Serica pilifera Horn, 1894 (Serica) 

Serica porcula Casey, 1884 (Serica) 

Serica prava Dawson, 1933 (Serica) 

Serica prunipennis Saylor, 1936 (Serica) 

Serica pruinosa Saylor, 1935 

Serica mendota Saylor, 1939 

Serica psammobuna Hardy, 1987 (Serica) 

Serica pullata Dawson, 1967 (Serica) 

Serica pusilla Dawson, 1922 (Serica) 

48

Serica errans Blatchley, 1929 

Serica repanda Dawson, 1933 (Serica) 

Serica rhypha Dawson, 1952 (Serica) 

Serica rossi Saylor, 1948 (Serica) 

Serica sandiegensis Saylor, 1939 (Serica) 

Serica satrapa Dawson, 1947 (Serica) 

Serica scaphia Dawson, 1952 (Serica) 

Serica sculptilis Dawson, 1922 (Serica) 

Serica senta Dawson, 1933 (Serica) 

Serica serensia Saylor, 1948 (Serica) 

Serica sericea (Illiger, 1802) (Melolontha) 

Serica sericeoides Dawson, 1967 (Serica) 

Serica serotina LeConte, 1856 (Serica) 

Serica solita Dawson, 1922 (Serica) 

Serica spicula Dawson, 1921 (Serica) 

Serica sponsa Dawson, 1919 (Serica) 

Serica stygia Dawson, 1933 (Serica) 

Serica subnisa Dawson, 1947 (Serica) 

Serica tantula Dawson, 1922 (Serica) 

Serica texana LeConte, 1856 (Serica) 

Serica tristis LeConte, 1850 (Serica) 

Serica trociformis Burmeister, 1844 (Serica) 

Melolontha aphodiina Billberg, 1820 

Serica ventura dorsalis Dawson, 1952 (Serica) 

Serica ventura personata Dawson, 1932 (Serica) 

Serica ventura ventura Dawson, 1932 (Serica) 

Serica vespertina accola Dawson, 1921 (Serica) 

Serica vespertina vespertina (Gyllenhal, 1817) (Melolontha) 

Serica watson Saylor, 1939 (Serica) 

Serica watsoni Dawson, 1952 

MELOLONTHINI Leach, 1819 

Amblonoxia Reitter, 1902 

Parathyce Hardy, 1974 

Amblonoxia bidentata (Fall, 1932) (Thyce) 

Amblonoxia carpenteri (LeConte, 1876) (Plectrodes) 

Thyce pistoria Casey, 1895 

Thyce crinicollis Casey, 1914 

Amblonoxia fieldi (Fall, 1908) (Thyce) 

Thyce angusticollis Casey, 1914 

Thyce simplicipes Casey, 1914 

Amblonoxia harfordi (Casey, 1889) (Thyce) 

Thyce squamosa Casey, 1891 

Thyce longipalpis Casey, 1914 

49

Thyce nanella Casey, 1914 

Amblonoxia palpalis (Horn, 1880) (Plectrodes) 

Thyce fossiger Casey, 1889 

Thyce pulverea Casey, 1889 

Thyce blaisdelli Casey, 1891 

Microphylla quedenfeldtii Kraatz, 1894 

Thyce angustula Casey, 1914 

Thyce aperta Casey, 1914 

Thyce brevitarsis Casey, 1914 

Thyce nitidula Casey, 1914 

Thyce ochreata Casey, 1914 

Thyce rotundicauda Casey, 1914 

Thyce vestita Casey, 1914 

Thyce aurata Van Dyke, 1943 

Thyce cinerea Van Dyke, 1943 

Thyce clypeata Van Dyke, 1943 

Amblonoxia riversi (Casey, 1895) (Thyce) 

Amphimallon Latreille, 1825 

Amphimallum Agassiz, 1847 

Amphimallon majale (Razoumowski, 1789) (Scarabaeus) 

Melolontha rufescens Latreille, 1804 

Melolontha semirufa Gyllenhal, 1817 

Amphimallon korbi Reitter, 1894 

Amphimallon pellitulum Reitter, 1902 

Dinacoma Casey, 1889 

Dinacoma caseyi Blaisdell, 1930 (Dinacoma) 

Dinacoma marginata (Casey, 1886) (Thyce) 

Fossocarus Howden, 1961 

Fossocarus creoleorum Howden, 1961 (Fossocarus) 

Gronocarus Schaeffer, 1927 

Gronocarus autumnalis Schaeffer, 1927 (Gronocarus) 

Gronocarus multispinosus Howden, 1961 

Gronocarus inornatus Skelley, 2005 (Gronocarus) 

Hypothyce Howden, 1968 

Hypothyce burnei Skelley, 2005 (Hypothyce) 

Hypothyce mixta Howden, 1968 (Hypothyce) 

50

Hypothyce osburni (Cartwright, 1967) (Thyce) 

Hypotrichia LeConte, 1861 

Hypotrichia spissipes LeConte, 1861 (Hypotrichia) 

Phyllophaga Harris, 1827 

Phyllophaga (Chlaenobia) Blanchard, 1851 

Phyllophaga (Cnemarachis) Saylor, 1942 

Abcrana Saylor, 1942 

Clemora Saylor, 1942 

Phyllophaga (Listrochelus) Blanchard, 1851 

Phyllophaga (Phyllophaga) Harris, 1827 

Lachnosterna Hope, 1837 

Trichesthes Erichson, 1847 

Endrosa LeConte, 1856 

Eugastra LeConte, 1856 

Gynnis LeConte, 1856 

Phyllophaga (Phytalus) Erichson, 1847 

Phyllophaga (Tostegoptera) Blanchard, 1851 

Phyllophaga (Triodonyx) Saylor, 1942 

Phyllophaga (Phyllophaga) aemula (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) aequalis (LeConte, 1854) (Tostegoptera) 

Lachnosterna exorata Horn, 1887 

Phyllophaga (Phyllophaga) affabilis (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) albina (Burmeister, 1855) (Ancylonycha) 

Phyllophaga (Listrochelus) almada Saylor, 1941 (Phyllophaga) 

Phyllophaga (Phyllophaga) anomaloides (Bates, 1888) (Lachnosterna) 

Phyllophaga (Phyllophaga) antennata (Smith, 1889) (Lachnosterna) 

Phyllophaga (Phyllophaga) anxia (LeConte, 1850) (Lachnosterna) 

Ancylonycha brevicollis Blanchard, 1851 

Ancylonycha guadulpensis Blanchard, 1851 

Ancylonycha puncticollis Blanchard, 1851 

Lachnosterna cephalica LeConte, 1856 

Ancylonycha uninotata Walker, 1866 

Lachnosterna dubia Smith, 1888 

Lachnosterna insperata Smith, 1889 

Lachnosterna alpina Linell, 1896 

Phyllophaga (Phyllophaga) apicata Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) arcta (Horn, 1887) (Lachnosterna) 

Phyllophaga opacita Reinhard, 1939 

Phyllophaga (Listrochelus) arenicola Howden, 1960 (Phyllophaga) 

Phyllophaga (Listrochelus) arizona Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) arkansana (Schaeffer, 1906) (Lachnosterna) 

51

Lachnosterna lenta Fall, 1908 

Phyllophaga (Phyllophaga) aurea Luginbill and Painter, 1941 (Phyllophaga) 

Phyllophaga (Listrochelus) babicora Morón, 2004 (Phyllophaga) 

Phyllophaga (Phyllophaga) balia (Say, 1825) (Melolontha) 

Ancylonycha comata Burmeister, 1855 

Phyllophaga (Phyllophaga) barda (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) beckeri (Moser, 1921) (Lachnosterna) 

Phyllophaga inopia Sanderson, 1942 

Phyllophaga (Triodonyx) bellamyi Warner and Morón, 1992 (Phyllophaga) 

Phyllophaga (Phytalus) bilobatata Saylor, 1939 (Phyllophaga) 

Phytalus cephalicus Horn, 1885 

Phyllophaga (Phyllophaga) bipartita (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) blanchardi (Arrow, 1933) (Lachnosterna) 

Ancylonycha cribricollis Blanchard, 1851 

Phyllophaga (Phytalus) blanda Sanderson, 1958 (Phyllophaga) 

Phyllophaga (Listrochelus) bottimeri Reinhard, 1950 (Phyllophaga) 

Phyllophaga (Phyllophaga) brevidens (Bates, 1888) (Lachnosterna) 

Phyllophaga (Cnemarachis) bruneri Chapin, 1932 (Phyllophaga) 

Phyllophaga (Listrochelus) bueta Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) cahitana Morón, 2001 (Phyllophaga) 

Phyllophaga (Phyllophaga) calceata (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) calculiventris Saylor, 1935 (Phyllophaga) 

Phyllophaga (Phyllophaga) callosiventris (Moser, 1921) (Lachnosterna) 

Phyllophaga (Phyllophaga) canipolea Saylor, 1948 (Phyllophaga) 

Phyllophaga (Listrochelus) carminator (Horn, 1894) (Listrochelus) 

Phyllophaga (Listrochelus) cavata (Bates, 1888) (Listrochelus) 

Listrochelus durangoensis Bates, 1888 

Phyllophaga (Phyllophaga) cazieriana Saylor, 1938 (Phyllophaga) 

Phyllophaga cazieri Saylor, 1936 

Phyllophaga (Phyllophaga) certanca Saylor, 1943 (Phyllophaga) 

Phyllophaga (Listrochelus) chapini Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) chippewa Saylor, 1939 (Phyllophaga) 

Phyllophaga (Phytalus) ciudadensis (Bates, 1888) (Phytalus) 

Phyllophaga (Phyllophaga) clemens (Horn, 1887) (Lachnosterna) 

Phyllophaga howei Sanderson, 1937 

Phyllophaga (Phyllophaga) clypeata (Horn, 1887) (Lachnosterna) 

Lachnosterna integra LeConte, 1856 

Phyllophaga (Listrochelus) cochisa Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) congrua (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) contaminata Fall, 1932 (Phyllophaga) 

Phyllophaga extranea Fall, 1932 

Phyllophaga (Phyllophaga) corrosa (LeConte, 1856) (Lachnosterna) 

Lachnosterna affinis LeConte, 1856 

Phyllophaga (Phyllophaga) crassissima (Blanchard, 1851) (Ancylonycha) 

Lachnosterna obesa LeConte, 1856 

Lachnosterna robusta LeConte, 1856 

52

Lachnosterna generosa Horn, 1887 

Phyllophaga (Phyllophaga) crenulata (Frölich, 1792) (Melolontha) 

Melolontha georgicana Gyllenhal, 1817 

Phyllophaga (Phyllophaga) cribrosa (LeConte, 1853) (Tostegoptera) 

Tostegoptera ventricosa LeConte, 1854 

Phyllophaga (Phyllophaga) crinita (Burmeister, 1855) (Trichestes) 

Lachnosterna glabripennis LeConte, 1856 

Listrochelus longiclavus Fall, 1922 

Phyllophaga (Listrochelus) cristagalli (Arrow, 1933) (Listochelus) 

Phyllophaga (Phyllophaga) cupuliformis Langston, 1924 (Phyllophaga) 

Phyllophaga (Phyllophaga) curialis Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Listrochelus) cushmani Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) cylindrica (Burmeister, 1855) (Trichestes) 

Phyllophaga emusa Saylor, 1937 

Phyllophaga (Phyllophaga) davisi Langston, 1927 (Phyllophaga) 

Phyllophaga (Phyllophaga) debilis (LeConte, 1856) (Gynnis) 

Phyllophaga austricolia Fall, 1929 

Phyllophaga (Phyllophaga) delata (Horn, 1887) (Lachnosterna) 

Phyllophaga (Listrochelus) densicollis (LeConte, 1863) (Listrochelus) 

Phyllophaga (Phyllophaga) dentex (Bates, 1888) (Lachnosterna) 

Phyllophaga (Phyllophaga) diffinis (Blanchard, 1851) (Ancylonycha) 

Trichestes comans Burmeister, 1855 

Lachnosterna sororia LeConte, 1856 

Lachnosterna rufiola LeConte, 1856 

Phyllophaga (Phyllophaga) dispar (Burmeister, 1855) (Trichestes) 

Lachnosterna boops Horn, 1887 

Phyllophaga (Listrochelus) disparilis (Horn, 1878) (Listrochelus) 

Phyllophaga (Phyllophaga) divertens (Bates, 1888) (Lachnosterna) 

Lachnosterna setipennis Bates, 1888 

Phyllophaga (Phyllophaga) drakii (Kirby, 1837) (Rhizotrogus) 

Lachnosterna consimilis LeConte, 1850 

Lachnosterna grandis Smith, 1888 

Phyllophaga (Listrochelus) duncani (Barrett, 1933) (Listrochelus) 

Phyllophaga (Listrochelus) durango Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) durangoana (Moser, 1921) (Listrochelus) 

Phyllophaga (Phyllophaga) durangosa Saylor, 1943 (Phyllophaga) 

Phyllophaga (Listrochelus) eligia Sanderson, 1958 (Phyllophaga) 

Phyllophaga (Cnemarachis) ecostata (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) elizoria Saylor, 1937 (Phyllophaga) 

Lachnosterna pygidialis Schaeffer, 1906 

Phyllophaga (Phyllophaga) elongata (Linell, 1896) (Lachnosterna) 

Phyllophaga murrea Sanderson, 1950 

Phyllophaga (Phyllophaga) eniba Saylor, 1943 (Phyllophaga) 

Phyllophaga (Phyllophaga) ephilida ephilida (Say, 1825) (Melolontha) 

Ancylonycha laticeps Blanchard, 1851 

Lachnosterna burmeisteri LeConte, 1856 

53

Phyllophaga (Phyllophaga) ephilida virilis Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) epigaea (Wickham, 1903) (Lachnosterna) 

Phyllophaga (Phyllophaga) errans (LeConte, 1860) (Lachnosterna) 

Phyllophaga (Listrochelus) estacea Saylor, 1943 (Phyllophaga) 

Listrochelus testaceipennis Moser, 1921 

Phyllophaga ponerosa Blackwelder, 1944 

Phyllophaga (Listrochelus) falsa (LeConte, 1856) (Listrochelus) 

Listrochelus langeri Chapin, 1934 

Phyllophaga (Phyllophaga) falta Sanderson, 1950 (Phyllophaga) 

Phyllophaga (Phyllophaga) farcta (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) fervida (Fabricius, 1775) (Melolontha) 

Melolontha quercina Knoch, 1801 

Lachnosterna arcuata Smith, 1888 

Phyllophaga (Listrochelus) fimbripes (LeConte, 1856) (Listrochelus) 

Phyllophaga (Listrochelus) flavipennis (Horn, 1885) (Listrochelus) 

Phyllophaga (Phyllophaga) floridana Robinson, 1938 (Phyllophaga) 

Phyllophaga (Listrochelus) foralita Saylor, 1938 (Phyllophaga) 

Phyllophaga (Phyllophaga) forbesi Glasgow, 1916 (Phyllophaga) 

Phyllophaga (Phyllophaga) forsteri (Burmeister, 1855) (Ancylonycha) 

Lachnosterna semicribrata LeConte, 1856 

Lachnosterna lugubris LeConte, 1856 

Lachnosterna lutescens LeConte, 1856 

Lachnosterna politula Horn, 1887 

Lachnosterna nova Smith, 1889 

Phyllophaga (Phyllophaga) foxii Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) fraterna fraterna Harris, 1842 (Phyllophaga) 

Ancylonycha cognata Burmeister, 1855 

Phyllophaga (Phyllophaga) fraterna mississippiensis Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) fucata (Horn, 1887) (Lachnosterna) 

Phyllophaga linsleyi Saylor, 1936 

Phyllophaga (Phyllophaga) fulviventris (Moser, 1918) (Lachnosterna) 

Phyllophaga (Phyllophaga) fusca (Frölich, 1792) (Melolontha) 

Melolontha fervens Gyllenhal, 1817 

Phyllophaga (Phyllophaga) futilis (LeConte, 1850) (Lachnosterna) 

Ancylonycha gibbosa Burmeister, 1855 

Lachnosterna decidua LeConte, 1856 

Lachnosterna serricornis LeConte, 1856 

Phyllophaga gaigei Sanderson, 1948 (Phyllophaga) 

Phyllophaga (Phyllophaga) galeanae Saylor, 1943 (Phyllophaga) 

Phyllophaga (Listrochelus) gentryi (Saylor, 1936) (Listrochelus) 

Phyllophaga (Phytalus) georgiana (Horn, 1885) (Phytalus) 

Phyllophaga (Phyllophaga) ghentata (Saylor, 1937) (Lachnosterna) 

Phyllophaga (Triodonyx) gigantissima Saylor, 1935 (Phyllophaga) 

Phyllophaga (Phyllophaga) glaberrima (Blanchard, 1851) (Ancylonycha) 

Lachnosterna parva Linell, 1896 

Phyllophaga pagilis Saylor, 1937 

54

Phyllophaga (Phyllophaga) glabricula (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) gracilis (Burmeister, 1855) (Trichesthes) 

Endrosa volvula LeConte, 1856 

Lachnosterna inana LeConte, 1856 

Phyllophaga angulata Glasgow, 1925 

Phyllophaga (Listrochelus) granti Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) gravidula (Moser, 1921) (Lachnosterna) 

Phyllophaga perita Sanderson, 1937 

Phyllophaga (Phyllophaga) hamata (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) hirsuta (Knoch, 1801) (Melolontha) 

Phyllophaga (Phyllophaga) hirticula comosa Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) hirticula hirticula (Knoch, 1801) (Melolontha) 

Phyllophaga (Phyllophaga) hirtiventris (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) hornii (Smith, 1889) (Lachnosterna) 

Phyllophaga (Listrochelus) huachuca Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) hubbelli Cartwright, 1946 (Phyllophaga) 

Phyllophaga (Phyllophaga) idonea Sanderson, 1948 (Phyllophaga) 

Phyllophaga (Phyllophaga) ignava (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) ilicis (Knoch, 1801) (Melolontha) 

Melolontha porcina Hentz, 1830 

Ancylonycha fimbriata Burmeister, 1855 

Lachnosterna ciliata LeConte, 1856 

Phyllophaga jonesi Sanderson, 1939 

Phyllophaga (Phyllophaga) impar Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) implicita (Horn, 1887) (Lachnosterna) 

Lachnosterna minor Linell, 1896 

Phyllophaga linelli Saylor, 1937 

Phyllophaga (Phyllophaga) incuria Sanderson, 1942 (Phyllophaga) 

Phyllophaga texana Sanderson, 1942 

Phyllophaga (Phyllophaga) inepta (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) infidelis (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) inflexa Barrett, 1935 (Phyllophaga) 

Phyllophaga (Chlaenobia) instabilis Blackwelder, 1944 (Phyllophaga) 

Phyllophaga arrowi Chapin, 1935 

Phyllophaga (Phyllophaga) inversa (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) invisa Riley and Wolfe, 1997 (Phyllophaga) 

Phyllophaga (Phyllophaga) juvenilis (Fall, 1932) (Listrochelus) 

Phyllophaga seri Morón, 2002 

Phyllophaga (Phyllophaga) karlsioei (Linell, 1898) (Lachnosterna) 

Phyllophaga (Phyllophaga) kentuckiana Ritcher, 1937 (Phyllophaga) 

Phyllophaga (Listrochelus) knausii (Schaeffer, 1907) (Listrochelus) 

Phyllophaga (Phyllophaga) knochii (Schönherr and Gyllenhal, 1817) (Melolontha) 

Phyllophaga (Listrochelus) koehleriana Saylor, 1940 (Phyllophaga) 

Phyllophaga (Triodonyx) lalanza Saylor, 1941 (Phyllophaga) 

Phyllophaga (Tostegoptera) lanceolata (Say, 1824) (Melolontha) 

Phyllophaga arizonae von Bloeker, 1936 

55

Phyllophaga cazieri von Bloeker, 1936 

Phyllophaga grisiana von Bloeker, 1936 

Phyllophaga (Listrochelus) laportaei (Blanchard, 1851) (Listrochelus) 

Phyllophaga (Phyllophaga) latidens (Schaeffer, 1906) (Lachnosterna) 

Phyllophaga (Phyllophaga) latifrons (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) lenis (Horn, 1887) (Lachnosterna) 

Lachnosterna anodentata Bates, 1888 

Phyllophaga (Phyllophaga) lobata (Fall, 1908) (Lachnosterna) 

Phyllophaga (Phyllophaga) lodingi Sanderson, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) longispina (Smith, 1889) (Lachnosterna) 

Phyllophaga (Phyllophaga) longitarsa (Say, 1824) (Melolontha) 

Lachnosterna frontalis LeConte, 1856 

Phyllophaga (Phyllophaga) lota Luginbill, 1928 (Phyllophaga) 

Phyllophaga (Phyllophaga) luctuosa (Horn, 1887) (Lachnosterna) 

Lachnosterna rugosioides Linell, 1896 

Phyllophaga (Listrochelus) luginbilli Saylor, 1941 (Phyllophaga) 

Phyllophaga (Listrochelus) macgregori Morón, 2004 (Phyllophaga) 

Phyllophaga (Listrochelus) macmurryi Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) maculicollis (LeConte, 1863) (Lachnosterna) 

Lachnosterna nitidula LeConte, 1863 

Phyllophaga (Phyllophaga) magnicornis (Moser, 1921) (Lachnosterna) 

Phyllophaga (Phyllophaga) marginalis insolita Cartwright, 1944 (Phyllophaga) 

Phyllophaga (Phyllophaga) marginalis marginalis (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) mariana Fall, 1929 (Phyllophaga) 

Phyllophaga (Listrochelus) meadei Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) micans (Knoch, 1801) (Melolontha) 

Phyllophaga (Listrochelus) michelbacheri Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) microcera (Arrow, 1933) (Lachnosterna) 

Lachnosterna cribricollis Bates, 1888 

Phyllophaga (Listrochelus) micros (Bates, 1888) (Listrochelus) 

Phyllophaga (Listrochelus) mimicana Saylor, 1938 (Phyllophaga) 

Phyllophaga (Listrochelus) miraflora Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) misteca (Bates, 1888) (Lachnosterna) 

Phyllophaga (Phyllophaga) molopia (Bates, 1888) (Lachnosterna) 

Phyllophaga (Listrochelus) monstrosa (Saylor, 1935) (Listochelus) 

Phyllophaga (Listrochelus) mucorea (LeConte, 1856) (Listrochelus) 

Listrochelus obtusus LeConte, 1856 

Listrochelus texanus LeConte, 1856 

Phyllophaga (Listrochelus) neomexicana Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) nitida (LeConte, 1856) (Lachnosterna) 

Lachnosterna limula Horn, 1887 

Lachnosterna innominata Smith, 1889 

Phyllophaga (Listrochelus) nogales Saylor, 1940 (Phyllophaga) 

Phyllophaga (Listrochelus) nogueirana Morón, 2002 (Phyllophaga) 

Phyllophaga (Phytalus) nosa Blackwelder, 1944 (Phyllophaga) 

Phytalus aegrotus Bates, 1888 

56

Phyllophaga (Phytalus) obsoleta (Blanchard, 1851) (Phytalus) 

Phytalus laevigatus Blanchard, 1851 

Ancylonycha longicornis Burmeister, 1855 

Phytalus longicornis Burmeister, 1855 

Phytalus oxypygus Burmeister, 1855 

Phyllophaga mecocera Blackwelder, 1944 

Phytalus vanalleri Schaeffer, 1927 

Phyllophaga (Phyllophaga) okeechobea Robinson, 1948 (Phyllophaga) 

Phyllophaga (Phytalus) omani Sanderson, 1937 (Phyllophaga) 

Phyllophaga (Phyllophaga) opaca (Moser, 1918) (Lachnosterna) 

Phyllophaga iroides Fall, 1929 

Phyllophaga (Listrochelus) opacicollis (Horn, 1878) (Listrochelus) 

Phyllophaga (Phyllophaga) ovalis Cartwright, 1939 (Phyllophaga) 

Phyllophaga (Phytalus) pallida (Horn, 1885) (Phytalus) 

Phyllophaga (Phyllophaga) panorpa Sanderson, 1950 (Phyllophaga) 

Phyllophaga (Listrochelus) parilis (Bates, 1888) (Listrochelus) 

Phyllophaga (Phyllophaga) parvidens hysteropyga Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) parvidens parvidens (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) paternoi Glasgow, 1925 (Phyllophaga) 

Phyllophaga (Phyllophaga) pearliae Davis, 1920 (Phyllophaga) 

Phyllophaga (Listrochelus) peninsularis Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phytalus) pentaphylla (Bates, 1888) (Phytalus) 

Phyllophaga (Phyllophaga) perlonga Davis, 1920 (Phyllophaga) 

Phyllophaga (Listrochelus) pilosipes Saylor, 1940 (Phyllophaga) 

Listrochelus puberulus LeConte, 1863 

Phyllophaga (Chlaenobia) pilositarsis Blackwelder, 1944 (Phyllophaga) 

Phyllophaga dissimilis Chapin, 1935 

Phyllophaga (Listrochelus) planeta Reinhard, 1950 (Phyllophaga) 

Phyllophaga (Listrochelus) plena (Fall, 1932) (Listrochelus) 

Phyllophaga (Phyllophaga) pleroma Reinhard, 1940 (Phyllophaga) 

Phyllophaga plena Reinhard, 1939 

Phyllophaga (Phyllophaga) nebulosa Polihronakis, 2007 (Phyllophaga) 

Phyllophaga (Phyllophaga) postrema (Horn, 1887) (Lachnosterna) 

Lachnosterna quadrata Smith, 1889 

Phyllophaga (Listrochelus) praesidii (Bates, 1888) (Listochelus) 

Phyllophaga (Phyllophaga) praetermissa (Horn, 1887) (Lachnosterna) 

Lachnosterna definita Smith, 1889 

Phyllophaga (Phyllophaga) profunda (Blanchard, 1851) (Ancylonycha) 

Lachnosterna biimpressa Smith, 1889 

Lachnosterna grandior Linell, 1896 

Phyllophaga (Phytalus) pruinosa (Blanchard, 1851) (Phytalus) 

Phyllophaga (Phyllophaga) prunina (LeConte, 1856) (Lachnosterna) 

Ancylonycha pruinosa Melsheimer, 1845 

Phyllophaga (Phyllophaga) prununculina (Burmeister, 1855) (Trichesthes) 

Lachnosterna cerasina LeConte, 1856 

Phyllophaga (Phyllophaga) pseudofloridana Woodruff and Beck, 1989 (Phyllophaga) 

57

Phyllophaga (Phyllophaga) psiloptera Sanderson, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) pudorosa Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Listrochelus) pulcher (Linell, 1896) (Listrochelus) 

Phyllophaga (Phyllophaga) pusillidens Fall, 1937 (Phyllophaga) 

Phyllophaga microdon Fall, 1929 

Phyllophaga (Phyllophaga) querca (Knoch, 1801) (Melolontha) 

Phyllophaga (Phyllophaga) quetzala Morón, 2001 (Phyllophaga) 

Phyllophaga (Phyllophaga) ravida (Blanchard, 1851) (Ancylonycha) 

Lachnosterna heterodoxa Horn, 1887 

Lachnosterna guatemalica Moser, 1918 

Lachnosterna longiclava Moser, 1918 

Phyllophaga (Phyllophaga) reevesi Saylor, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) regiomontana Morón, 2001 (Phyllophaga) 

Phyllophaga (Listrochelus) reinhardi Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) renodis Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) riviera Reinhard, 1950 (Phyllophaga) 

Phyllophaga (Phyllophaga) rolbakeri Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) rolstoni Riley and Wolfe, 1997 (Phyllophaga) 

Phyllophaga (Phyllophaga) rossi Saylor, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) rubiginosa (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) rubricosa Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) rugipennis (Schaufuss, 1858) (Ancylonycha) 

Phyllophaga (Phyllophaga) rugosa (Melsheimer, 1845) (Ancylonycha) 

Phyllophaga (Phyllophaga) sacoma Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) saylori Sanderson, 1965 (Phyllophaga) 

Phyllophaga (Phyllophaga) schaefferi Saylor, 1937 (Phyllophaga) 

Lachnosterna georgiana Schaeffer, 1909 

Phyllophaga duvala Robinson, 1938 

Phyllophaga (Phyllophaga) schenklingi Moser, 1918 (Lachnosterna) 

Phyllophaga (Phyllophaga) scitula (Horn, 1887) (Lachnosterna) 

Phyllophaga (Listrochelus) scoparia (LeConte, 1856) (Listrochelus) 

Phyllophaga (Listrochelus) scuticeps (Bates, 1888) (Listrochelus) 

Phyllophaga (Phyllophaga) segregans (Bates, 1888) (Lachnosterna) 

Phyllophaga (Listrochelus) senex (Horn, 1878) (Listrochelus) 

Phyllophaga (Phyllophaga) sequoiana Saylor, 1936 (Phyllophaga) 

Phyllophaga (Phyllophaga) setifera (Burmeister, 1855) (Ancylonycha) 

Phyllophaga (Phytalus) sinaloana Saylor, 1935 (Phyllophaga) 

Phyllophaga (Phyllophaga) sinuaticeps (Moser, 1921) (Lachnosterna) 

Phyllophaga (Phyllophaga) skelleyi Woodruff and Beck, 1989 (Phyllophaga) 

Phyllophaga (Listrochelus) snowi Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) sociata (Horn, 1878) (Lachnosterna) 

Phyllophaga (Phyllophaga) sodalis Reinhard, 1940 (Phyllophaga) 

Phyllophaga (Phytalus) sonora Saylor, 1939 (Phyllophaga) 

Phytalus debilis Horn, 1885 

Phyllophaga (Phyllophaga) soror Davis, 1920 (Phyllophaga) 

Phyllophaga (Phyllophaga) spreta (Horn, 1887) (Lachnosterna) 

58

Phyllophaga (Phyllophaga) squamipilosa Saylor, 1937 (Phyllophaga) 

Phyllophaga (Phyllophaga) stohleri Saylor, 1938 (Phyllophaga) 

Phyllophaga (Phyllophaga) submucida (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) subpruinosa (Casey, 1884) (Lachnosterna) 

Phyllophaga deanii Luginbill, 1928 

Phyllophaga (Phyllophaga) subtonsa (LeConte, 1856) (Lachnosterna) 

Phyllophaga (Phyllophaga) sylvatica Sanderson, 1942 (Phyllophaga) 

Phyllophaga (Listrochelus) tarsalis (Schaeffer, 1908) (Listochelus) 

Phyllophaga (Phyllophaga) taxodii Langston, 1924 (Phyllophaga) 

Phyllophaga (Phyllophaga) tecta Cartwright, 1944 (Phyllophaga) 

Phyllophaga (Listrochelus) temaxa Saylor, 1940 (Phyllophaga) 

Lachnosterna parilis Bates, 1888 

Phyllophaga (Phyllophaga) temora Saylor, 1943 (Phyllophaga) 

Phyllophaga (Listrochelus) texensis Saylor, 1940 (Phyllophaga) 

Phyllophaga (Listrochelus) timida (Horn, 1878) (Listochelus) 

Phyllophaga (Phyllophaga) torta (LeConte, 1856) (Lachnosterna) 

Lachnosterna dampfi Arrow, 1933 

Phyllophaga (Phytalus) trichodes (Bates, 1890) (Phytalus) 

Phytalus robustus Horn, 1885 

Phyllophaga sandersonia Saylor, 1939 

Phyllophaga (Phyllophaga) tristis amplicornis Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) tristis suttonana Reinhard, 1939 (Phyllophaga) 

Phyllophaga (Phyllophaga) tristis tristis (Fabricius, 1781) (Melolontha) 

Melolontha pilosicollis Knoch, 1801 

Phyllophaga (Listrochelus) trochanter Saylor, 1940 (Phyllophaga) 

Listrochelus gracilis Horn, 1885 

Phyllophaga (Phyllophaga) tusa (Horn, 1887) (Lachnosterna) 

Phyllophaga (Phyllophaga) ulkei (Smith, 1889) (Lachnosterna) 

Phyllophaga (Phyllophaga) uniformis (Blanchard, 1851) (Ancylonycha) 

Lachnosterna carolina Fall, 1912 

Phyllophaga (Phyllophaga) vehemens (Horn, 1887) (Lachnosterna) 

Phyllophaga (Listrochelus) venodiola Saylor, 1948 (Phyllophaga) 

Phyllophaga (Phyllophaga) verruciventris (Moser, 1918) (Lachnosterna) 

Phyllophaga (Phyllophaga) vetula (Horn, 1887) (Lachnosterna) 

Lachnosterna longipilosa Bates, 1888 

Phyllophaga (Chlaenobia) vexata vexata (Horn, 1885) (Phytalus) 

Phytalus cavifrons Linell, 1896 

Phyllophaga (Phyllophaga) vilifrons (LeConte, 1856) (Lachnosterna) 

Lachnosterna hirticeps LeConte, 1856 

Phyllophaga (Listrochelus) wickhami Saylor, 1940 (Phyllophaga) 

Phyllophaga (Phyllophaga) xanthe (Bates, 1888) (Lachnosterna) 

Phyllophaga (Phyllophaga) xerophila Saylor, 1939 (Phyllophaga) 

Phyllophaga (Listrochelus) yaqui Saylor, 1940 (Phyllophaga) 

Listrochelus debilis Arrow, 1933 

Phyllophaga (Phyllophaga) yemasseei Cartwright, 1944 (Phyllophaga) 

Phyllophaga (Cnemarachis) youngi Cartwright, 1935 (Phyllophaga) 

59

Phyllophaga bahama Saylor, 1940 

Phyllophaga (Phyllophaga) zavalana Reinhard, 1946 (Phyllophaga) 

Plectrodes Horn, 1867 

Plectrodes pubescens Horn, 1867 (Plectrodes) 

Polyphylla Harris, 1841 

Polyphylla (Polyphylla) Harris, 1841 

Macranoxia Crotch, 1873 

Polylamina Hardy, 1974 

Polyphylla (Polyphylla) aeola La Rue, 1998 (Polyphylla) 

Polyphylla (Polyphylla) anteronivea Hardy, 1978 (Polyphylla) 

Polyphylla (Polyphylla) arguta Casey, 1914 (Polyphylla) 

Polyphylla (Polyphylla) avittata Hardy, 1978 (Polyphylla) 

Polyphylla (Polyphylla) barbata Cazier, 1938 (Polyphylla) 

Polyphylla (Polyphylla) brownae Young, 1986 (Polyphylla) 

Polyphylla (Polyphylla) cavifrons LeConte, 1854 (Polyphylla) 

Polyphylla (Polyphylla) comes Casey, 1914 (Polyphylla) 

Polyphylla (Polyphylla) crinita LeConte, 1856 (Polyphylla) 

Polyphylla pacifica Casey, 1895 

Polyphylla incolumis Casey, 1914 

Polyphylla mystica Casey, 1914 

Polyphylla perversa Casey, 1914 

Polyphylla ona von Bloeker, 1939 

Polyphylla santarosae von Bloeker, 1939 

Polyphylla (Polyphylla) decemlineata (Say, 1824) (Melolontha) 

Polyphylla castanea Casey, 1914 

Polyphylla laticauda Casey, 1914 

Polyphylla matrona Casey, 1914 

Polyphylla oregona Casey, 1914 

Polyphylla parilis Casey, 1914 

Polyphylla potosiana Casey, 1914 

Polyphylla reducta Casey, 1914 

Polyphylla ruficollis Casey, 1914 

Polyphylla squamotecta Casey, 1914 

Polyphylla comstockiana von Bloeker, 1939 

Polyphylla (Polyphylla) devestiva Young, 1966 (Polyphylla) 

Polyphylla (Polyphylla) diffracta Casey, 1891 (Polyphylla) 

Polyphylla fuscula Fall, 1908 

Polyphylla adusta Casey, 1914 

Polyphylla laevicauda Casey, 1914 

Polyphylla opposita Casey, 1914 

Polyphylla uteana Tanner, 1928 

60

Polyphylla alleni Cazier, 1939 

Polyphylla arida Van Dyke, 1947 

Polyphylla (Polyphylla) donaldsoni Skelley, 2005 (Polyphylla) 

Polyphylla (Polyphylla) erratica Hardy, 1978 (Polyphylla) 

Polyphylla (Polyphylla) gracilis Horn, 1881 (Polyphylla) 

Polyphylla (Polyphylla) hammondi LeConte, 1856 (Polyphylla) 

Polyphylla subvittata LeConte, 1856 

Polyphylla speciosa Casey, 1889 

Polyphylla acomana Casey, 1914 

Polyphylla bisinuata Casey, 1914 

Polyphylla diffusa Casey, 1914 

Polyphylla impigra Casey, 1914 

Polyphylla latifrons Casey, 1914 

Polyphylla molesta Casey, 1914 

Polyphylla oblita Casey, 1914 

Polyphylla pimalis Casey, 1914 

Polyphylla proba Casey, 1914 

Polyphylla sejuncta Casey, 1914 

Polyphylla squamicauda Casey, 1914 

Polyphylla verecunda Casey, 1914 

Polyphylla oklahomensis Hatch, 1926 

Polyphylla rufescenta Tanner, 1928 

Polyphylla (Polyphylla) hirsuta Van Dyke, 1933 (Polyphylla) 

Polyphylla (Polyphylla) mescalerensis Young, 1988 (Polyphylla) 

Polyphylla (Polyphylla) modulata Casey, 1914 (Polyphylla) 

Polyphylla (Polyphylla) monahansensis Hardy, 1978 (Polyphylla) 

Polyphylla (Polyphylla) multimaculata Hardy, 1981 (Polyphylla) 

Polyphylla (Polyphylla) nigra Casey, 1914 (Polyphylla) 

Polyphylla relicta Casey, 1914 

Polyphylla robustula Casey, 1914 

Polyphylla martini von Bloeker, 1939 

Polyphylla santacruzae von Bloeker, 1939 

Polyphylla (Polyphylla) nubila Van Dyke, 1947 (Polyphylla) 

Polyphylla (Polyphylla) occidentalis (Linnaeus, 1767) (Scarabaeus) 

Polyphylla (Polyphylla) petitii (Guérin-Méneville, 1844) (Melolontha) 

Melolontha leucogramma Blanchard, 1851 

Polyphylla dubia Hatch, 1971 

Polyphylla (Polyphylla) pottsorum Hardy, 1978 (Polyphylla) 

Polyphylla (Polyphylla) pubescens Cartwright, 1939 (Polyphylla) 

Polyphylla (Polyphylla) ratcliffei Young, 1986 (Polyphylla) 

Polyphylla (Polyphylla) rugosipennis Casey, 1914 (Polyphylla) 

Polyphylla peninsularis Van Dyke, 1947 

Polyphylla (Polyphylla) sobrina Casey, 1914 (Polyphylla) 

Polyphylla (Polyphylla) squamiventris Cazier, 1939 (Polyphylla) 

Polyphylla (Polyphylla) stellata Young, 1986 (Polyphylla) 

Polyphylla (Polyphylla) variolosa (Hentz, 1830) (Melolontha) 

61

Polyphylla (Polyphylla) woodruffi Skelley, 2005 (Polyphylla) 

Thyce LeConte, 1856 

Thyce deserta Hardy, 1974 (Thyce) 

Thyce squamicollis LeConte, 1856 (Thyce) 

DIPLOTAXINI Kirby, 1837 

Diplotaxis Kirby, 1837 

Orsonyx LeConte, 1856 

Alobus LeConte, 1856 

Diazus LeConte, 1859 

Diplotaxis abnormis Fall, 1909 (Diplotaxis) 

Diplotaxis academia Vaurie, 1960 (Diplotaxis) 

Diplotaxis aenea Blanchard, 1851 (Diplotaxis) 

Liogenys pauperata Burmeister, 1855 

Diplotaxis aequalis Cazier, 1940 (Diplotaxis) 

Diplotaxis ambigua Fall, 1909 (Diplotaxis) 

Diplotaxis angularis LeConte, 1856 (Diplotaxis) 

Diplotaxis producta Fall, 1932 

Diplotaxis anthracina Fall, 1909 (Diplotaxis) 

Diplotaxis anxia (LeConte, 1856) (Orsonyx) 

Diplotaxis arctifrons Bates, 1888 (Diplotaxis) 

Diplotaxis arizonica Schaeffer, 1907 (Diplotaxis) 

Diplotaxis atlantis Fall, 1909 (Diplotaxis) 

Diplotaxis nigra Cartwright, 1939 

Diplotaxis atratula LeConte, 1856 (Diplotaxis) 

Diplotaxis aulacochela Cazier, 1940 (Diplotaxis) 

Diplotaxis australis Vaurie, 1960 (Diplotaxis) 

Diplotaxis bakeri Cazier, 1940 (Diplotaxis) 

Diplotaxis volatica Cazier, 1940 

Diplotaxis barbarae Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis basalis Fall, 1909 (Diplotaxis) 

Diplotaxis belfragei Fall, 1909 (Diplotaxis) 

Diplotaxis sinuata Fall, 1909 

Diplotaxis protensa Fall, 1932 

Diplotaxis cartwrighti Cazier, 1940 

Diplotaxis stabilis Cazier, 1940 

Diplotaxis beyeri Schaeffer, 1907 (Diplotaxis) 

Diplotaxis bidentata LeConte, 1856 (Diplotaxis) 

Diplotaxis blanchardi Vaurie, 1956 (Diplotaxis) 

Diplotaxis boops Bates, 1888 (Diplotaxis) 

Diplotaxis bowditchi Fall, 1909 (Diplotaxis) 

Diplotaxis brachyptera Vaurie, 1960 (Diplotaxis) 

62

Diplotaxis brevicollis LeConte, 1856 (Diplotaxis) 

Ancylonycha nigropicea Walker, 1866 

Diplotaxis transversicollis Moser, 1918 

Diplotaxis brevicornis Cazier, 1940 (Diplotaxis) 

Diplotaxis brevidens LeConte, 1856 (Diplotaxis) 

Diplotaxis laeviscutata Moser, 1918 

Diplotaxis brevisetosa Linell, 1896 (Diplotaxis) 

Diplotaxis caelestis Delgado and Capistran, 1993 (Diplotaxis) 

Diplotaxis carbonata LeConte, 1856 (Diplotaxis) 

Diplotaxis levicula Casey, 1885 

Diplotaxis arcuata Fall, 1909 

Diplotaxis catarinas Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis chiricahuae Fall, 1909 (Diplotaxis) 

Diplotaxis circulans Vaurie, 1960 (Diplotaxis) 

Diplotaxis coenonychoides Davidson and Davidson, 2006 (Diplotaxis) 

Diplotaxis commixta Vaurie, 1960 (Diplotaxis) 

Diplotaxis completa Cazier, 1940 (Diplotaxis) 

Diplotaxis conformis Fall, 1909 (Diplotaxis) 

Diplotaxis confusa Fall, 1909 (Diplotaxis) 

Diplotaxis connata Schaeffer, 1905 (Diplotaxis) 

Diplotaxis contracta Bates, 1888 (Diplotaxis) 

Diplotaxis convexilabra Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis corbula Vaurie, 1960 (Diplotaxis) 

Diplotaxis coriacea Bates, 1888 (Diplotaxis) 

Diplotaxis corvina LeConte, 1856 (Diplotaxis) 

Diplotaxis costanera Vaurie, 1958 (Diplotaxis) 

Diplotaxis cribatella Bates, 1889 (Diplotaxis) 

Diplotaxis ennea Vaurie and Cazier, 1955 

Diplotaxis cribulosa cribulosa LeConte, 1856 (Diplotaxis) 

Diplotaxis popino Casey, 1885 

Diplotaxis cribulosa sinaloa Vaurie, 1958 (Diplotaxis) 

Diplotaxis curvaticeps Fall, 1909 (Diplotaxis) 

Diplotaxis dahli Cazier, 1940 (Diplotaxis) 

Diplotaxis decima Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis dentella Fall, 1909 (Diplotaxis) 

Diplotaxis denticeps Bates, 1887 (Diplotaxis) 

Diplotaxis deserta Fall, 1932 (Diplotaxis) 

Diplotaxis dissona Vaurie, 1960 (Diplotaxis) 

Diplotaxis dubia LeConte, 1856 (Diplotaxis) 

Diplotaxis durango Vaurie, 1960 (Diplotaxis) 

Diplotaxis elongata Vaurie, 1960 (Diplotaxis) 

Diplotaxis errans Fall, 1909 (Diplotaxis) 

Diplotaxis ungula Cazier, 1940 

Diplotaxis exstans Vaurie, 1960 (Diplotaxis) 

Diplotaxis fimbriata Fall, 1909 (Diplotaxis) 

Diplotaxis arta Cazier, 1940 

63

Diplotaxis fissilabris Fall, 1909 (Diplotaxis) 

Diplotaxis fissilis Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis flexa Vaurie, 1960 (Diplotaxis) 

Diplotaxis fossipalpa Fall, 1909 (Diplotaxis) 

Diplotaxis villosipes Fall, 1932 

Diplotaxis frondicola (Say, 1825) (Melolontha) 

Diplotaxis testacea Burmeister, 1855 

Diplotaxis fulva (LeConte, 1856) (Alobus) 

Diplotaxis glabrimargo Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis harperi Blanchard, 1851 (Diplotaxis) 

Diplotaxis haydenii LeConte, 1856 (Diplotaxis) 

Diplotaxis innoxia LeConte, 1856 

Diplotaxis levicoxa Fall, 1909 

Diplotaxis impressifrons Cazier, 1940 

Diplotaxis hebes Bates, 1888 (Diplotaxis) 

Diplotaxis sericeiventris Moser, 1918 

Diplotaxis hirsuta Vaurie, 1958 (Diplotaxis) 

Diplotaxis illustris Fall, 1909 (Diplotaxis) 

Diplotaxis impar Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis incuria Fall, 1932 (Diplotaxis) 

Diplotaxis indigena Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis ingenua Fall, 1909 (Diplotaxis) 

Diplotaxis rufocastanea Moser, 1918 

Diplotaxis fusca Moser, 1918 

Diplotaxis futilis Fall, 1932 

Diplotaxis insignis LeConte, 1861 (Diplotaxis) 

Diplotaxis falli Saylor, 1935 

Diplotaxis knausii Schaeffer, 1907 (Diplotaxis) 

Diplotaxis kuschei Vaurie, 1960 (Diplotaxis) 

Diplotaxis languida LeConte, 1878 (Diplotaxis) 

Diplotaxis lengii Fall, 1909 (Diplotaxis) 

Diplotaxis levicosta Fall, 1909 (Diplotaxis) 

Diplotaxis liberta (Germar, 1824) (Melolontha) 

Melolontha moesta Say, 1825 

Diplotaxis georgiae Blanchard, 1851 

Diplotaxis ebenina Blanchard, 1851 

Diplotaxis macronycha Fall, 1909 (Diplotaxis) 

Diplotaxis magna Bates, 1888 (Diplotaxis) 

Diplotaxis costulata Fall, 1909 

Diplotaxis impressicollis Moser, 1918 

Diplotaxis marginicollis Fall, 1909 (Diplotaxis) 

Diplotaxis martinezi Delgado and Capistran, 1993 (Diplotaxis) 

Diplotaxis mascula Vaurie, 1960 (Diplotaxis) 

Diplotaxis maura Fall, 1909 (Diplotaxis) 

Diplotaxis megapleura Vaurie, 1958 (Diplotaxis) 

Diplotaxis mentalis Fall, 1909 (Diplotaxis) 

64

Diplotaxis mima Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis mimosae Fall, 1909 (Diplotaxis) 

Diplotaxis misella Fall, 1909 (Diplotaxis) 

Diplotaxis missionaria Vaurie, 1960 (Diplotaxis) 

Diplotaxis moerens moerens LeConte, 1856 (Diplotaxis) 

Diplotaxis moerens peninsularis Fall, 1909 (Diplotaxis) 

Diplotaxis muricata Schaeffer, 1907 (Diplotaxis) 

Diplotaxis sparsesetosa Schaeffer, 1907 

Diplotaxis hispida Fall, 1909 

Diplotaxis mus Fall, 1932 (Diplotaxis) 

Diplotaxis nigriventris Bates, 1887 (Diplotaxis) 

Diplotaxis obregon Vaurie, 1958 (Diplotaxis) 

Diplotaxis obscura LeConte, 1859 (Diplotaxis) 

Ancylonycha consequens Walker, 1866 

Diplotaxis prominens Fall, 1909 

Diplotaxis compacta Fall, 1909 

Diplotaxis ohausi Moser, 1921 (Diplotaxis) 

Diplotaxis pacata LeConte, 1856 (Diplotaxis) 

Diplotaxis pala Vaurie, 1960 (Diplotaxis) 

Diplotaxis parallela Fall, 1909 (Diplotaxis) 

Diplotaxis parkeri Cazier, 1940 

Diplotaxis parpolita Vaurie, 1960 (Diplotaxis) 

Diplotaxis parvicollis Fall, 1909 (Diplotaxis) 

Diplotaxis labrata Moser, 1918 

Diplotaxis acononica Cazier, 1940 

Diplotaxis persisae Cazier, 1940 (Diplotaxis) 

Diplotaxis pilipennis Moser, 1918 (Diplotaxis) 

Diplotaxis pinalica Fall, 1932 (Diplotaxis) 

Diplotaxis microps Cazier, 1940 

Diplotaxis recticantha Cazier, 1940 

Diplotaxis planidens Fall, 1909 (Diplotaxis) 

Diplotaxis polita Fall, 1909 (Diplotaxis) 

Diplotaxis profunda Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis puberula LeConte, 1863 (Diplotaxis) 

Diplotaxis villosa Fall, 1909 

Diplotaxis pubipes Schaeffer, 1907 (Diplotaxis) 

Diplotaxis pumila Fall, 1909 (Diplotaxis) 

Diplotaxis punctata LeConte, 1856 (Diplotaxis) 

Diplotaxis acerba Fall, 1909 

Diplotaxis fulgida Cazier, 1940 

Diplotaxis punctatorugosa Blanchard, 1851 (Diplotaxis) 

Diplotaxis excavata LeConte, 1856 

Diplotaxis frontalis LeConte, 1856 

Diplotaxis densicollis Fall, 1909 

Diplotaxis puncticeps Moser, 1918 (Diplotaxis) 

Diplotaxis punctipennis LeConte, 1856 (Diplotaxis) 

65

Diplotaxis punctulata Horn, 1894 (Diplotaxis) 

Diplotaxis pygidialis Vaurie, 1960 (Diplotaxis) 

Diplotaxis residua Fall, 1909 (Diplotaxis) 

Diplotaxis rex Vaurie, 1958 (Diplotaxis) 

Diplotaxis robertmarki Davidson and Davidson, 2006 (Diplotaxis) 

Diplotaxis rockefelleri Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis rosae Vaurie and Cazier, 1955 (Diplotaxis) 

Diplotaxis rotunda Vaurie, 1960 (Diplotaxis) 

Diplotaxis rudis (LeConte, 1859) (Diazus) 

Diplotaxis rufa Linell, 1895 (Diplotaxis) 

Diplotaxis rufiola Fall, 1909 (Diplotaxis) 

Diplotaxis fulva Robinson, 1946 

Diplotaxis saltensis Vaurie, 1960 (Diplotaxis) 

Diplotaxis saylori Cazier, 1940 (Diplotaxis) 

Diplotaxis schaefferi Fall, 1909 (Diplotaxis) 

Diplotaxis semifoveata Fall, 1909 (Diplotaxis) 

Diplotaxis sierrae Fall, 1909 (Diplotaxis) 

Diplotaxis simplex Blanchard, 1851 (Diplotaxis) 

Diplotaxis sinuaticeps Bates, 1888 

Diplotaxis cephalotes Fall, 1909 

Diplotaxis sonora Vaurie, 1960 (Diplotaxis) 

Diplotaxis sordida (Say, 1825) (Melolontha) 

Diplotaxis carbonaria Burmeister, 1855 

Diplotaxis rugosioides Schaeffer, 1907 

Diplotaxis sparsa Vaurie, 1960 (Diplotaxis) 

Diplotaxis statura Cazier, 1940 (Diplotaxis) 

Diplotaxis subangulata LeConte, 1856 (Diplotaxis) 

Diplotaxis californica Schaeffer, 1907 

Diplotaxis subcostata Blanchard, 1851 (Diplotaxis) 

Diplotaxis castanea Burmeister, 1855 

Diplotaxis subrugata Moser, 1918 (Diplotaxis) 

Diplotaxis sulcatula Fall, 1909 (Diplotaxis) 

Diplotaxis superflua Vaurie, 1960 (Diplotaxis) 

Diplotaxis tarsalis Schaeffer, 1907 (Diplotaxis) 

Diplotaxis mexicana Moser, 1918 

Diplotaxis tenebrosa Fall, 1909 (Diplotaxis) 

Diplotaxis tenuis LeConte, 1856 (Diplotaxis) 

Diplotaxis texana LeConte, 1856 (Diplotaxis) 

Diplotaxis thoracica Fall, 1909 (Diplotaxis) 

Diplotaxis pinguescens Fall, 1909 

Diplotaxis trapezifera Bates, 1887 (Diplotaxis) 

Diplotaxis trementina Fall, 1909 (Diplotaxis) 

Diplotaxis tristis Kirby, 1837 (Diplotaxis) 

Diplotaxis corpulenta Burmeister, 1855 

Diplotaxis truncatula LeConte, 1856 (Diplotaxis) 

Diplotaxis consors LeConte, 1856 

66

Diplotaxis morula LeConte, 1856 

Diplotaxis ungulata Cazier, 1940 (Diplotaxis) 

Diplotaxis urbana Vaurie, 1960 (Diplotaxis) 

Diplotaxis vandykei Vaurie, 1958 (Diplotaxis) 

PACHYDEMINI Burmeister, 1855 

Phobetus LeConte, 1856 

Phobetus chearyi Hardy, 1973 (Phobetus) 

Phobetus ciliatus Barrett, 1935 (Phobetus) 

Phobetus comatus LeConte, 1856 (Phobetus) 

Phobetus centralis Casey, 1909 

Phobetus sloopi Barrett, 1933 

Phobetus leachi Barrett, 1935 

Phobetus robinsoni Saylor, 1939 

Phobetus desertus Blom and Clark, 1984 (Phobetus) 

Phobetus humeralis Cazier, 1937 (Phobetus) 

Phobetus mojavus Barrett, 1933 (Phobetus) 

Phobetus palpalis Saylor, 1936 (Phobetus) 

Phobetus panamintensis Hardy, 1978 (Phobetus) 

Phobetus saylori Cazier, 1937 (Phobetus) 

Phobetus sleeperi Hardy, 1978 (Phobetus) 

Phobetus testaceus LeConte, 1861 (Phobetus) 

Rhizotrogus collocatus Walker, 1866 

Warwickia Smith and Evans, 2005 

Benedictia Sanderson, 1939 

Warwickia pilosa (Sanderson, 1939) (Benedictia) 

DICHELONYCHINI Burmeister, 1855 

Coenonycha Horn, 1876 

Coenonycha acuta Cazier, 1943 (Coenonycha) 

Coenonycha ampla Cazier, 1943 (Coenonycha) 

Coenonycha barri Cazier, 1943 (Coenonycha) 

Coenonycha bowlesi Cazier, 1943 (Coenonycha) 

Coenonycha clementina Casey, 1909 (Coenonycha) 

Coenonycha clypeata McClay, 1943 (Coenonycha) 

Coenonycha crispata McClay, 1943 (Coenonycha) 

Coenonycha dimorpha Evans, 1986 (Coenonycha) 

Coenonycha fuga Cazier, 1943 (Coenonycha) 

Coenonycha fulva McClay, 1943 (Coenonycha) 

Coenonycha fusca McClay, 1943 (Coenonycha) 

67

Coenonycha globosa McClay, 1943 (Coenonycha) 

Coenonycha hageni Cazier, 1943 (Coenonycha) 

Coenonycha lurida Cazier, 1943 (Coenonycha) 

Coenonycha mediata Cazier, 1943 (Coenonycha) 

Coenonycha ochreata Evans, 1986 (Coenonycha) 

Coenonycha ovatis McClay, 1943 (Coenonycha) 

Coenonycha ovipennis Horn, 1876 (Coenonycha) 

Coenonycha pallida Cazier, 1943 (Coenonycha) 

Coenonycha parvula Fall, 1901 (Coenonycha) 

Coenonycha pascuensis Potts, 1945 (Coenonycha) 

Coenonycha purshiae Cazier, 1943 (Coenonycha) 

Coenonycha pygmaea Smith, 1986 (Coenonycha) 

Coenonycha rotundata (LeConte, 1856) (Dichelonyx) 

Coenonycha rufobrunnea Howden, 1969 

Coenonycha rubida McClay, 1943 (Coenonycha) 

Coenonycha santacruzae Evans, 1986 (Coenonycha) 

Coenonycha saylori Cazier, 1943 (Coenonycha) 

Coenonycha scotti McClay, 1943 (Coenonycha) 

Coenonycha sleeperi Evans, 1988 (Coenonycha) 

Coenonycha socialis Horn, 1876 (Coenonycha) 

Coenonycha stohleri Saylor, 1935 (Coenonycha) 

Coenonycha testacea Cazier, 1937 (Coenonycha) 

Coenonycha tingi Cazier, 1937 (Coenonycha) 

Coenonycha utahensis McClay, 1943 (Coenonycha) 

Dichelonyx Harris, 1827 

Philochloenia Dejean, 1833 

Anaeretes Dejean, 1836 

Dichelonyx albicollis Burmeister, 1855 (Dichelonyx) 

Dichelonyx backii Kirby, 1837 (Dichelonyx) 

Dichelonyx fulgida LeConte, 1856 

Dichelonyx crotchii Horn, 1876 

Dichelonyx mormona Fall, 1907 

Dichelonyx oregona Van Dyke, 1918 

Dichelonyx arizonica Barrett, 1933 

Dichelonyx canadensis Horn, 1876 (Dichelonyx) 

Dichelonyx clypeata Horn, 1876 (Dichelonyx) 

Dichelonyx crumbi Hatch, 1971 (Dichelonyx) 

Dichelonyx decolorata Fall, 1901 (Dichelonyx) 

Dichelonyx diluta Fall, 1901 (Dichelonyx) 

Dichelonyx elongatula (Schönherr, 1817) (Melolontha) 

Melolontha elongata Fabricius, 1793 

Melolontha linearis Gyllenhal, 1817 

Melolontha hexagona Germar, 1824 

Dichelonyx virescens Kirby, 1837 

68

Dichelonyx testacea Kirby, 1837 

Dichelonyx fuscula LeConte, 1856 (Dichelonyx) 

Dichelonyx kirbyi Brown, 1946 (Dichelonyx) 

Dichelonyx lateralis Fall, 1901 (Dichelonyx) 

Dichelonyx muscula Fall, 1901 (Dichelonyx) 

Dichelonyx nana Fall, 1901 (Dichelonyx) 

Dichelonyx pallens LeConte, 1856 (Dichelonyx) 

Dichelonyx longiclava Fall, 1901 

Dichelonyx picea Horn, 1894 (Dichelonyx) 

Dichelonyx pusilla LeConte, 1856 (Dichelonyx) 

Dichelonyx robusta Fall, 1901 (Dichelonyx) 

Dichelonyx subvittata LeConte, 1856 (Dichelonyx) 

Dichelonyx sulcata LeConte, 1856 (Dichelonyx) 

Dichelonyx arizonensis Saylor, 1933 

Dichelonyx testaceipennis Fall, 1907 (Dichelonyx) 

Dichelonyx truncata LeConte, 1856 (Dichelonyx) 

Dichelonyx vaga Fall, 1901 (Dichelonyx) 

Dichelonyx valida LeConte, 1856 (Dichelonyx) 

Dichelonyx vandykei Saylor, 1932 (Dichelonyx) 

Dichelonyx vicina Fall, 1901 (Dichelonyx) 

Dichelonyx deserta Hopping, 1931 

Dichelonyx columbiana Hopping, 1931 

Gymnopyge Linell, 1896 

Gymnopyge coquilletti Linell, 1896 (Gymnopyge) 

Gymnopyge hirsuta Cazier, 1939 (Gymnopyge) 

Gymnopyge hopliaeformis Linell, 1896 (Gymnopyge) 

Gymnopyge pygmaea Linell, 1896 (Gymnopyge) 

MACRODACTYLINI Kirby, 1837 

Isonychus Mannerheim, 1829 

Colporhina Curtis, 1844 

Isonychus arizonensis Howden, 1959 (Isonychus) 

Macrodactylus Dejean, 1821 

Stenothorax Harris, 1827 

Chremastodus Solier, 1851 

Macrodactylus angustatus (Palisot de Beauvois, 1805) (Melolontha) 

Macrodactylus setulosus LeConte, 1856 

Macrodactylus fulvescens Bates, 1887 (Macrodactylus) 

Macrodactylus nigritarsis Bates, 1887 

Macrodactylus rhomboderus Bates, 1887 

69

Macrodactylus impressus Bates, 1887 (Macrodactylus) 

Macrodactylus infuscatus Bates, 1887 (Macrodactylus) 

Macrodactylus vicinus Bates, 1887 

Macrodactylus lineaticollis Bates, 1887 (Macrodactylus) 

Macrodactylus mexicanus Burmeister, 1855 (Macrodactylus) 

Macrodactylus murinus Bates, 1887 (Macrodactylus) 

Macrodactylus nigripes Bates, 1887 (Macrodactylus) 

Macrodactylus rufescens Bates, 1887 (Macrodactylus) 

Macrodactylus silaonus Bates, 1887 (Macrodactylus) 

Macrodactylus subspinosus (Fabricius, 1775) (Melolontha) 

Melolontha elongata Herbst, 1790 

Macrodactylus polyphagus Burmeister, 1855 

Macrodactylus angustatus LeConte, 1856 

Macrodactylus barbatus Fitch, 1863 

Macrodactylus uniformis Horn, 1876 (Macrodactylus) 

Macrodactylus variipes Bates, 1887 (Macrodactylus) 

Macrodactylus virens Bates, 1887 (Macrodactylus) 

Plectris LePeletier and Serville, 1828 

Pseudoserica Guérin-Méneville, 1838 

Euryaspis Blanchard, 1851 

Trichoderma Nonfried, 1894 

Junkia Dalla Torre, 1913 

Plectris aliena Chapin, 1934 (Plectris) 

HOPLIINI Latreille, 1829 

Hoplia Illiger, 1803 

Hoplia (Hoplia) Illiger, 1803 

Hopliatus Rafinesque, 1815 

Diphydactylus Thomson, 1858 

Hoplia (Hoplia) callipyge LeConte, 1856 (Hoplia) 

Hoplia convexula LeConte, 1856 

Hoplia irrorata LeConte, 1856 

Hoplia oregona LeConte, 1856 

Hoplia pubicollis LeConte, 1856 

Hoplia mutata Harold, 1869 

Hoplia lecontei Dalla Torre, 1913 

Hoplia cazieri Boyer, 1940 

Hoplia deserticola Boyer, 1940 

Hoplia humboltensis Boyer, 1940 

Hoplia utahensis Boyer, 1940 

Hoplia (Hoplia) cinereonebulosa Nonfried, 1895 (Hoplia) 

70

Hoplia (Hoplia) dispar LeConte, 1880 (Hoplia) 

Hoplia (Hoplia) equina LeConte, 1880 (Hoplia) 

Hoplia (Hoplia) floridana Fisher, 1918 (Hoplia) 

Hoplia (Hoplia) gilleti Hardy, 1977 (Hoplia) 

Hoplia (Hoplia) hirta LeConte, 1880 (Hoplia) 

Hoplia (Hoplia) laticollis LeConte, 1856 (Hoplia) 

Hoplia (Hoplia) modesta Haldeman, 1843 (Hoplia) 

Hoplia singularis Burmeister, 1844 

Hoplia monticola Melsheimer, 1845 

Hoplia helvola Melsheimer, 1845 

Hoplia barbata Blatchley, 1910 

Hoplia (Hoplia) mucorea (Germar, 1824) (Melolontha) 

Hoplia meridionalis Boyer, 1940 

Hoplia (Hoplia) sackenii LeConte, 1880 (Hoplia) 

Hoplia (Hoplia) trifasciata Say, 1825 (Hoplia) 

Hoplia primaria Burmeister, 1844 

Hoplia tristis Melsheimer, 1845 

Hoplia ferrisi Boyer, 1940 

Hoplia (Hoplia) trivialis Harold, 1869 (Hoplia) 

Hoplia debilis LeConte, 1856 

INCERTA SEDIS 

Acoma Casey, 1889 

Acoma arizonica Brown, 1929 (Acoma) 

Acoma brunnea Casey, 1889 (Acoma) 

Acoma cazieri Saylor, 1948 (Acoma) 

Acoma confusa Van Dyke, 1928 (Acoma) 

Acoma conjuncta Howden, 1962 (Acoma) 

Acoma dilemma Saylor, 1948 (Acoma) 

Acoma diminiata Howden, 1958 (Acoma) 

Acoma evansi Howden, 1962 (Acoma) 

Acoma gibsoni Howden, 1962 (Acoma) 

Acoma glabrata Cazier, 1953 (Acoma) 

Acoma nigrita Cazier, 1953 

Acoma granulifrons Howden, 1958 (Acoma) 

Acoma incognita Howden, 1958 (Acoma) 

Acoma knulli Howden, 1958 (Acoma) 

Acoma leechi Cazier, 1953 (Acoma) 

Acoma martini Howden, 1962 (Acoma) 

Acoma mimica Howden, 1962 (Acoma) 

Acoma minuta Cazier, 1953 (Acoma) 

Acoma mixta Howden, 1958 (Acoma) 

Acoma ochlera Howden, 1958 (Acoma) 

Acoma parva Howden, 1958 (Acoma) 

71

Acoma robusta Van Dyke, 1928 (Acoma) 

Acoma rossi Saylor, 1948 (Acoma) 

Acoma rufula Howden, 1958 (Acoma) 

Acoma seticollis Howden, 1958 (Acoma) 

Acoma sexfoliata Saylor, 1948 (Acoma) 

Acoma stathamae Cazier, 1953 (Acoma) 

Chaunocolus Saylor, 1937 

Chaunocolus cornutus Saylor, 1937 (Chaunocolus) 

Chnaunanthus Burmeister, 1844 

Acratus Horn, 1867 

Pseudacratus Dalla Torre, 1912 

Chnaunanthus chapini Saylor, 1937 (Chnaunanthus) 

Chnaunanthus discolor Burmeister, 1844 (Chnaunanthus) 

Chnaunanthus flavipennis (Horn, 1867) (Acratus) 

Chnaunanthus palmeri Horn, 1894 

RUTELINAE MacLeay, 1819 

RUTELINI MacLeay, 1819 

Calomacraspis Bates, 1888 

Calomacraspis splendens (Burmeister, 1844) (Macraspis) 

Chrysina Kirby, 1828 

Plusiotis Burmeister, 1844 

Pelidnotopsis Ohaus, 1915 

Plusiotina Casey, 1915 

Chrysina adelaida (Hope, 1840) (Pelidnota) 

Pelidnota ornatissima Sturm, 1843 

Chrysina beckeri Bates, 1889 (Chrysina) 

Chrysina beyeri (Skinner, 1905) (Plusiotis) 

Plusiotis ampliata Casey, 1915 

Plusiotis ocularis Casey, 1915 

Chrysina brevis (Rothschild and Jordan, 1894) (Plusiotis) 

Chrysina erubescens Bates, 1889 (Chrysina) 

Chrysina flohri (Ohaus, 1905) (Plusiotis) 

Chrysina gloriosa (LeConte, 1854) (Plusiotis) 

Chrysina lecontei (Horn, 1882) (Plusiotis) 

Plusiotina aeruginis Casey, 1915 

Plusiotina angusta Casey, 1915 

Plusiotina sonorica Casey, 1915 

72

Plusiotina subenodis Casey, 1915 

Chrysina plusiotina (Ohaus, 1912) (Pelidnota) 

Chrysina psittacina (Sturm, 1843) (Pelidnota) 

Pelidnota laeta Sturm, 1843 

Chrysina woodii (Horn, 1885) (Plusiotis) 

Cotalpa Burmeister, 1844 

Ciocotalpa Saylor, 1940 

Cotalpa ashleyae La Rue, 1986 (Cotalpa) 

Cotalpa conclamara Young, 2002 (Cotalpa) 

Cotalpa consobrina Horn, 1871 (Cotalpa) 

Cotalpa flavida Horn, 1878 (Cotalpa) 

Cotalpa lanigera (Linnaeus, 1758) (Scarabaeus) 

Cotalpa tau Wickham, 1905 

Cotalpa molaris Casey, 1915 

Cotalpa obesa Casey, 1915 

Cotalpa vernicata Casey, 1915 

Cotalpa subcribrata Wickham, 1905 (Cotalpa) 

Ectinoplectron Ohaus, 1915 

Ectinoplectron oryctoides (Ohaus, 1905) (Homonyx) 

Pelidnota howdeni Hardy, 1975 

Homoiosternus Ohaus, 1901 

Homoiosternus beckeri Ohaus, 1901 (Homoiosternus) 

Parabyrsopolis Ohaus, 1915 

Byrsopolis Burmeister, 1844 

Parareoda Casey, 1915 

Parabyrsopolis chihuahuae (Bates, 1888) (Byrsopolis) 

Byrsopolis lanigera Bates, 1888 

Byrsopolis arizonae Ohaus, 1912 

Cotalpa batesi Ohaus, 1915 

Parareoda rufobrunnea Casey, 1915 

Parabyrsopolis villadealendensis Morón, 1976 

Parachrysina Bates, 1888 

Parachrysina borealis Jameson, 1991 (Parachrysina) 

Paracotalpa Ohaus, 1915 

Pocalta Casey, 1915 

73

Paracotalpa deserta Saylor, 1940 (Paracotalpa) 

Paracotalpa granicollis (Haldeman, 1852) (Cotalpa) 

Pocalta pubicollis Casey, 1915 

Paracotalpa puncticollis (LeConte, 1863) (Cotalpa) 

Paracotalpa ursina (Horn, 1867) (Cotalpa) 

Pocalta brevis Casey, 1915 

Pocalta laevicauda Casey, 1915 

Pocalta nigripennis Casey, 1915 

Pocalta rotunda Casey, 1915 

Pocalta rubripennis Casey, 1915 

Pocalta seriata Casey, 1915 

Cotalpa leonina Fall, 1932 

Paracotalpa piceola Saylor, 1940 

Parastasia Westwood, 1842 

Barymorpha Guérin-Méneville, 1843 

Polymoechus LeConte, 1856 

Urleta Westwood, 1875 

Echmatophorus Waterhouse, 1895 

Ohkubous Sawada, 1938 

Parastasia brevipes (LeConte, 1856) (Polymoechus) 

Polymoechus conicicollis Casey, 1915 

Polymoechus discernens Casey, 1915 

Pelidnota MacLeay, 1819 

Odontognathus Laporte, 1840 

Strigidia Burmeister, 1844 

Chalcoplethis Burmeister, 1844 

Aglycoptera Sharp, 1885 

Epichalcoplethis Bates, 1904 

Delipnia Casey, 1915 

Gananota Ohaus, 1915 

Pelidnotidia Casey, 1915 

Pelidnota lucae LeConte, 1863 (Pelidnota) 

Pelidnota lugubris LeConte, 1874 (Pelidnota) 

Pelidnota perplexa Hardy, 1975 (Pelidnota) 

Pelidnota punctata (Linnaeus, 1758) (Scarabaeus) 

Melolontha lutea Olivier, 1789 

Pelidnota brevicollis Casey, 1915 

Pelidnota brevis Casey, 1915 

Pelidnota debiliceps Casey, 1915 

Pelidnota hudsonica Casey, 1915 

Pelidnota oblonga Casey, 1915 

74

Pelidnota pallidipes Casey, 1915 

Pelidnota ponderella Casey, 1915 

Pelidnota strenua Casey, 1915 

Pelidnota tarsalis Casey, 1915 

Pelidnota texensis Casey, 1915 

Pelidnota strigosa Laporte, 1840 (Pelidnota) 

Pelidnota alutacea Bates, 1888 

Pelidnota cuprascens Casey, 1915 

Pelidnota obscurella Casey, 1915 

Pelidnota refulgens Casey, 1915 

Pelidnota virescens Burmeister, 1844 (Pelidnota) 

Pelidnota planipennis Ohaus, 1918 

Pelidnota permicans Casey, 1915 

Plesiosternus Morón, 1983 

Plesiosternus setosus Morón, 1983 (Plesiosternus) 

Pseudocotalpa Hardy, 1971 

Pseudocotalpa andrewsi Hardy, 1971 (Pseudocotalpa) 

Pseudocotalpa giulianii Hardy, 1974 (Pseudocotalpa) 

Pseudocotalpa sonorica Hardy, 1974 (Pseudocotalpa) 

Rutela Latreille, 1802 

Rutela formosa Burmeister, 1844 (Rutela) 

Rutelisca Bates, 1888 

Rutelisca durangoana Ohaus, 1905 (Rutelisca) 

ANOMALINI Streubel, 1839 

ANOMALINA Streubel, 1839 

Anomala Samouelle, 1819 

Phyllopertha Stephens, 1830 

Spilota Burmeister, 1844 

Callistethus Blanchard, 1851 

Pachystethus Blanchard, 1851 

Peripopillia Kolbe, 1894 

Blitopertha Reitter, 1903 

Cyriopertha Reitter, 1903 

Exomala Reitter, 1903 

Megapertha Reitter, 1903 

Pleopertha Reitter, 1903 

75

Trichopertha Reitter, 1903 

Apleopertha Semenov, 1903 

Anomalepta Casey, 1915 

Anomalopus Casey, 1915 

Hemispilota Casey, 1915 

Oliganomala Casey, 1915 

Paranomala Casey, 1915 

Rhombonalia Casey, 1915 

Anomalopides Strand, 1928 

Asiopertha Medvedev, 1949 

Anomala adscita (Robinson, 1941) (Rhombonalia) 

Anomala antennata Schaeffer, 1906 (Anomala) 

Anomala arida Casey, 1915 (Anomala) 

Anomala beckeri Ohaus, 1897 (Anomala) 

Anomala binotata (Gyllenhal, 1817) (Melolontha) 

Melolontha unifasciata Say, 1825 

Euchlora maculata Laporte, 1840 

Anomala marginella LeConte, 1854 

Anomala compacta Casey, 1915 

Anomala butleri (Howden, 1955) (Rhombonalia) 

Anomala carinifrons Bates, 1888 (Anomala) 

Anomala camancha Wickham, 1913 

Anomala carlsoni Hardy, 1976 (Anomala) 

Anomala castaniceps Bates, 1888 (Anomala) 

Anomala cavifrons LeConte, 1867 (Anomala) 

Anomala apacheana Wickham, 1913 

Rhombonalia cochiseana Casey, 1915 

Rhombonalia comes Casey, 1915 

Rhombonalia transversa Casey, 1915 

Anomala centralis LeConte, 1863 (Anomala) 

Anomala raydoma Saylor, 1948 

Anomala chevrolati Bates, 1888 (Anomala) 

Anomala cincta cincta Say, 1835 (Anomala) 

Anomala latreillei Blanchard, 1851 

Anomala crinicollis Ohaus, 1902 (Anomala) 

Anomala delicata Casey, 1915 (Anomala) 

Anomala moquina Casey, 1915 

Anomala papagoana Casey, 1915 

Anomala diabla Potts, 1976 (Anomala) 

Anomala digressa Casey, 1915 (Anomala) 

Anomala moquina Casey, 1915 

Anomala sagax Casey, 1915 

Anomala discoidalis Bates, 1888 (Anomala) 

Anomala dubia (Scopoli, 1763) (Scarabaeus) 

Anomala ellipsis Casey, 1915 (Anomala) 

76

Anomala exigua (Schwarz, 1878) (Strigoderma) 

Anomala eximia Potts, 1976 (Anomala) 

Anomala flavilla coachellae Potts, 1977 (Anomala) 

Anomala flavilla flavilla Bates, 1888 (Anomala) 

Anomala tenera Casey, 1915 

Anomala flavipennis aransas Potts, 1977 (Anomala) 

Anomala flavipennis flavipennis Burmeister, 1844 (Anomala) 

Anomala dichroa Melsheimer, 1845 

Anomala luteipennis LeConte, 1854 

Anomala amissa Casey, 1915 

Anomala modulata Casey, 1915 

Anomala stigmatella Casey, 1915 

Anomala kansana Hayes and McColloch, 1924 

Anomala flavipennis okaloosensis Potts, 1977 (Anomala) 

Anomala flavipennis subquadrata Casey, 1915 (Anomala) 

Anomala flohri Ohaus, 1897 (Anomala) 

Anomala foraminosa Bates, 1888 (Anomala) 

Anomala hardyorum Potts, 1976 (Anomala) 

Anomala histrionella Bates, 1888 (Anomala) 

Anomala imperialae Potts, 1976 (Anomala) 

Anomala innuba (Fabricius, 1787) (Melolontha) 

Anomala minuta Burmeister, 1844 

Anomala medorensis Casey, 1915 

Anomala piceola Casey, 1915 

Anomala insitiva Robinson, 1938 (Anomala) 

Anomala kanei Potts, 1976 (Anomala) 

Anomala lucicola (Fabricius, 1798) (Melolontha) 

Melolontha atrata Fabricius, 1801 

Melolontha moerens Fabricius, 1801 

Melolontha quadrimaculata Drapiez, 1820 

Euchlora nigritula Laporte, 1840 

Anomala cinctella Burmeister, 1844 

Anomala pinicola Melsheimer, 1845 

Anomala phylloperthoides Nonfried, 1894 

Anomala alterata Arrow, 1915 

Anomala ludoviciana Schaeffer, 1906 (Anomala) 

Anomala pubescens Blatchley, 1910 

Anomala marginata (Fabricius, 1793) (Melolontha) 

Melolontha annulata Germar, 1824 

Spilota incolumis Casey, 1915 

Anomala mendica Casey, 1915 (Anomala) 

Anomala nimbosa Casey, 1915 (Anomala) 

Anomala oblivia Horn, 1884 (Anomala) 

Spilota maritima Casey, 1915 

Anomala orientalis (Waterhouse, 1875) (Phyllopertha) 

Phyllopertha xanthogastra Harold, 1881 

77

Blitopertha flavipennis Reitter, 1903 

Phyllopertha tanbaensis Niijima and Kinoshita, 1923 

Anomala parvula Burmeister, 1844 (Anomala) 

Anomala peninsularis Schaeffer, 1906 (Anomala) 

Anomala robinsoni Potts, 1974 (Anomala) 

Pachystethus floridana Robinson, 1941 

Anomala sabinae Potts, 1976 (Anomala) 

Anomala semilivida LeConte, 1878 (Anomala) 

Anomalepta flaccida Casey, 1915 

Anomala suavis Potts, 1976 (Anomala) 

Anomala tibialis Schaeffer, 1906 (Anomala) 

Anomala umbra Casey, 1915 (Anomala) 

Anomala servilis Casey, 1915 

Anomala undulata Melsheimer, 1845 (Anomala) 

Anomala varians Burmeister, 1844 

Anomala dubia Blatchley, 1910 

Anomala canadensis Casey, 1915 

Anomala floridana Casey, 1915 

Anomala nigropicta Casey, 1915 

Anomala saginulata Casey, 1915 

Strigoderma Burmeister, 1844 

Alamona Casey, 1915 

Strigodermella Casey, 1915 

Strigoderma arbicola (Fabricius, 1793) (Melolontha) 

Strigoderma testaceipennis Nonfried, 1893 

Strigoderma angustula Casey, 1915 

Strigoderma irregularis Casey, 1915 

Strigoderma obesula Casey, 1915 

Strigoderma puritana Casey, 1915 

Strigoderma quaternaria Casey, 1915 

Strigoderma texana Casey, 1915 

Strigoderma virginica Casey, 1915 

Strigoderma costulipennis Bates, 1888 (Strigoderma) 

Strigoderma knausi (Brown, 1925) (Strigodermella) 

Strigodermella oklahomensis Hatch, 1930 

Strigoderma pimalis Casey, 1884 (Strigoderma) 

Strigoderma prolixella Casey, 1915 

Strigoderma sonorica Casey, 1915 

Strigoderma subrutilans Casey, 1915 

Strigoderma viridicollis Schaeffer, 1907 

Strigoderma protea Burmeister, 1844 (Strigoderma) 

Strigoderma nigrita Blanchard, 1851 

Strigoderma pygmaea (Fabricius, 1798) (Melolontha) 

Melolontha minima Drapiez, 1819 

78

Strigoderma floridana Ohaus, 1905 

Strigoderma sulcipennis Burmeister, 1844 (Strigoderma) 

Strigoderma fastuosa Burmeister, 1944 

Strigoderma nigripes Burmeister, 1944 

Strigoderma imitatrix Nonfried, 1893 

Strigoderma impressicollis Nonfried, 1893 

Strigoderma metallica Nonfried, 1893 

Strigoderma aterrima Casey, 1915 

Strigoderma convergens Casey, 1915 

Strigoderma mediocris Casey, 1915 

Strigoderma morelosana Casey, 1915 

Strigoderma radula Casey, 1915 

Strigoderma teapensis Bates, 1888 (Strigoderma) 

Strigoderma latitibia Schaeffer, 1907 

Alamona parviceps Casey, 1915 

Strigoderma vestita Burmeister, 1844 (Strigoderma) 

ANISOPLIINA Burmeister, 1844 

Anomalacra Casey, 1915 

Anomalacra clypealis (Schaeffer, 1907) (Anomala) 

Anomalacra cuneata Casey, 1915 

Anomalacra werneri Howden, 1956 

POPILLIINA Ohaus, 1918 

Popillia Dejean, 1821 

Calopopillia Kolbe, 1894 

Eupopillia Kolbe, 1894 

Metapopillia Kolbe, 1894 

Pseudopopillia Kolbe, 1894 

Godschama Reitter, 1903 

Xenopopillia Kolbe, 1910 

Castapopillia Benderitter, 1922 

Popillia japonica Newman, 1841 (Popillia) 

Popillia plicatipennis Burmeister, 1844 

LEPTOHOPLIINA Potts, 1974 

Leptohoplia Saylor, 1935 

Leptohoplia testaceipennis Saylor, 1935 (Leptohoplia) 

DYNASTINAE MacLeay, 1819 

79

CYCLOCEPHALINI Laporte, 1840 

Ancognatha Erichson, 1847 

Barotheus Bates, 1891 

Pseudancognatha Otoya, 1945 

Ancognatha manca (LeConte, 1866) (Cyclocephala) 

Ancognatha aequata Bates, 1888 

Ancognatha laevigata Bates, 1888 

Ancognatha durangoana Casey, 1915 

Ancognatha perspicua Casey, 1915 

Ancognatha zuniella Casey, 1915 

Cyclocephala Dejean, 1821 

Aclinidia Casey, 1915 

Halotosia Casey, 1915 

Diapatalia Casey, 1915 

Dichromina Casey, 1915 

Graphalia Casey, 1915 

Homochromina Casey, 1915 

Isocoryna Casey, 1915 

Mononidia Casey, 1915 

Ochrosidia Casey, 1915 

Plagiosalia Casey, 1915 

Spilosota Casey, 1915 

Stigmalia Casey, 1915 

Aspidolites Höhne, 1922 

Aspidolella Prell, 1936 

Albridarollia Bolívar y Pieltain, Jiménez-Asúa, and Martínez, 1963 

Paraclinidia Martínez, 1965 

Cyclocephala aravaipensis Ratcliffe, 1992 (Cyclocephala) 

Cyclocephala arenosa Howden and Endrödi, 1966 (Cyclocephala) 

Cyclocephala barrerai Martínez, 1969 (Cyclocephala) 

Cyclocephala borealis Arrow, 1911 (Cyclocephala) 

Cyclocephala villosa Burmeister, 1847 

Cyclocephala caelestis Delgado and Ratcliffe, 1990 (Cyclocephala) 

Cyclocephala comata Bates, 1888 (Cyclocephala) 

Cyclocephala deceptor (Casey, 1915) (Stigmalia) 

Stigmalia cuernavacana Casey, 1915 

Stigmalia deficiens Casey, 1915 

Stigmalia fallaciosa Casey, 1915 

Cyclocephala forcipulata Howden and Endrödi, 1966 (Cyclocephala) 

Cyclocephala freudei Endrödi, 1963 (Cyclocephala) 

Cyclocephala hirta LeConte, 1861 (Cyclocephala) 

Spilosota inconspicua Casey, 1915 

80

Spilosota magister Casey, 1915 

Spilosota nubeculina Casey, 1915 

Spilosota pallidissima Casey, 1915 

Cyclocephala knobelae (Brown, 1934) (Ochrosidia) 

Cyclocephala longula LeConte, 1863 (Cyclocephala) 

Ochrosidia abrupta Casey, 1915 

Ochrosidia ambiens Casey, 1915 

Ochrosidia marcida Casey, 1915 

Ochrosidia modulata Casey, 1915 

Ochrosidia obesula Casey, 1915 

Ochrosidia oblongula Casey, 1915 

Ochrosidia phasma Casey, 1915 

Ochrosidia prona Casey, 1915 

Ochrosidia reflexa Casey, 1915 

Ochrosidia rugulifrons Casey, 1915 

Ochrosidia rustica Casey, 1915 

Ochrosidia californica Arrow, 1937 

Cyclocephala lunulata Burmeister, 1847 (Cyclocephala) 

Cyclocephala nubeculosa Burmeister, 1847 

Ochrosidia oblita Casey, 1915 

Cyclocephala lurida coahuilae Bates, 1888 (Cyclocephala) 

Cyclocephala lurida lurida Bland, 1863 (Cyclocephala) 

Ochrosidia pagana Casey, 1915 

Ochrosidia protenta Casey, 1915 

Ochrosidia rufifrons Casey, 1915 

Ochrosidia tenuicutis Casey, 1915 

Cyclocephala mafaffa Burmeister, 1847 (Cyclocephala) 

Cyclocephala grandis Burmeister, 1847 

Stigmalia histrionica Casey, 1915 

Cyclocephala melanocephala (Fabricius, 1775) (Melolontha) 

Chalepus leucophthalma Fischer von Waldheim, 1823 

Cyclocephala dimidiata Burmeister, 1847 

Cyclocephala ventralis Erichson, 1847 

Cyclocephala elegans Horn, 1871 

Dichromina ocularis Casey, 1915 

Cyclocephala miamiensis Howden and Endrödi, 1966 (Cyclocephala) 

Cyclocephala nigricollis Burmeister, 1847 (Cyclocephala) 

Cyclocephala robusta LeConte, 1863 

Ochrosidia subvittata Brown, 1930 

Cyclocephala parallela (Casey, 1915) (Ochrosidia) 

Cyclocephala pasadenae (Casey, 1915) (Ochrosidia) 

Ochrosidia arizonica Casey, 1915 

Ochrosidia facilis Casey, 1915 

Ochrosidia melina Casey, 1915 

Ochrosidia ovulata Casey, 1915 

Ochrosidia pusilla Casey, 1915 

81

Ochrosidia validiceps Casey, 1915 

Cyclocephala mexica Martínez, 1969 

Cyclocephala pilosicollis Saylor, 1936 (Cyclocephala) 

Cyclocephala puberula LeConte, 1863 (Cyclocephala) 

Cyclocephala regularis Casey, 1915 (Cyclocephala) 

Cyclocephala sanguinicollis Burmeister, 1847 (Cyclocephala) 

Cyclocephala seditiosa LeConte, 1863 (Cyclocephala) 

Cyclocephala sinaloae Howden and Endrödi, 1966 (Cyclocephala) 

Cyclocephala sororia Bates, 1888 (Cyclocephala) 

Cyclocephala sparsa Arrow, 1902 (Cyclocephala) 

Cyclocephala landini Endrödi, 1964 

Cyclocephala virkkii Howden and Endrödi, 1966 

Cyclocephala testacea Burmeister, 1847 (Cyclocephala) 

Cyclocephala ovulum Bates, 1888 

Cyclocephala variabilis Burmeister, 1847 (Cyclocephala) 

Cyclocephala wandae Hardy, 1974 (Cyclocephala) 

Dyscinetus Harold, 1869 

Chalepus MacLeay, 1819 

Palechus Casey, 1915 

Dyscinetus laevicollis Arrow, 1937 (Dyscinetus) 

Dyscinetus morator (Fabricius, 1798) (Melolontha) 

Chalepus trachypygus Burmeister, 1847 

Dyscinetus bitumorosus Casey, 1915 

Dyscinetus borealis Casey, 1915 

Dyscinetus discedens Casey, 1915 

Dyscinetus picipes (Burmeister, 1847) (Chalepus) 

Chalepus obsoletus LeConte, 1854 

Chalepus geminatus Jacquelin du Val, 1856 

Dyscinetus punctipes Bates, 1888 

Dyscinetus puncticauda Casey, 1909 

Dyscinetus ebeninus Casey, 1915 

Dyscinetus gilanus Casey, 1915 

Dyscinetus laevissimus Casey, 1915 

Dyscinetus obsidianus Casey, 1915 

Dyscinetus subquadratus Casey, 1915 

PENTODONTINI Mulsant, 1842 

Anoplognatho Rivers, 1888 

Aphonides Rivers, 1889 

Anoplognatho dunnianus Rivers, 1888 (Anoplognatho) 

Aphonus LeConte, 1856 

82

Aphonus brevicruris Cartwright, 1944 (Aphonus) 

Aphonus castaneus (Melsheimer, 1845) (Bothynus) 

Podalgus obesus Burmeister, 1847 

Aphonus cubiformis Casey, 1915 

Aphonus saginatus Casey, 1915 

Aphonus trapezicollis Casey, 1915 

Aphonus densicauda Casey, 1915 (Aphonus) 

Aphonus texanus Gill and Howden, 1985 (Aphonus) 

Aphonus tridentatus (Say, 1823) (Scarabaeus) 

Aphonus frater LeConte, 1856 

Aphonus aterrimus Casey, 1915 

Aphonus congestus Casey, 1915 

Aphonus elongatus Casey, 1915 

Aphonus modulatus Casey, 1915 

Aphonus politus Casey, 1915 

Aphonus scutellaris Casey, 1924 

Aphonus variolosus (LeConte, 1847) (Bothynus) 

Aphonus hydropicus LeConte, 1856 

Aphonus ingens Casey, 1924 

Coscinocephalus Prell, 1936 

Anoplocephalus Schaeffer, 1906 

Coscinocephalus cribrifrons (Schaeffer, 1906) (Anoplocephalus) 

Euetheola Bates, 1888 

Euetheola humilis (Burmeister, 1847) (Heteronychus) 

Ligyrus rugiceps LeConte, 1856 

Euetheola hondurana Casey, 1915 

Dyscinetus parvus Casey, 1915 

Euetheola subglabra (Schaeffer, 1909) (Ligyrus) 

Gillaspytes Howden, 1980 

Gillaspytes janzeni Howden, 1980 (Gillaspytes) 

Orizabus Fairmaire, 1878 

Aztecalius Casey, 1915 

Pseudaphonus Casey, 1915 

Orizabus batesi Prell, 1914 (Orizabus) 

Orizabus brevicollis Prell, 1914 (Orizabus) 

Orizabus clunalis (LeConte, 1856) (Aphonus) 

Orizabus cultripes Fairmaire, 1878 

83

Orizabus marginatus Fairmaire, 1878 

Orizabus sallei Fairmaire, 1878 

Orizabus ligyrodes Horn, 1885 

Orizabus snowi Horn, 1885 

Orizabus verticalis Fall, 1905 

Orizabus fontinalis Casey, 1915 

Orizabus parvitarsis Casey, 1915 

Orizabus ponderosus Casey, 1915 

Orizabus isodonoides Fairmaire, 1878 (Orizabus) 

Orizabus pyriformis (LeConte, 1847) (Bothynus) 

Pseudaphonus debiliceps Casey, 1915 

Pseudaphonus lucidus Casey, 1915 

Pseudaphonus ovalis Casey, 1915 

Pseudaphonus repens Casey, 1915 

Pseudaphonus puncticollis Casey, 1924 

Orizabus ratcliffei Delgado, 2008 (Orizabus) 

Orizabus rubricollis Prell, 1914 (Orizabus) 

Oxygrylius Casey, 1915 

Oxygrylius peninsularis Casey, 1915 (Oxygrylius) 

Oxygrylius ruginasus (LeConte, 1856) (Ligyrus) 

Oxygrylius pimalis Casey, 1915 

Tomarus Erichson, 1847 

Ligyrus Burmeister, 1847 

Anagrylius Casey, 1915 

Ligyrellus Casey, 1915 

Ligyrodes Casey, 1915 

Tomarus cuniculus (Fabricius, 1801) (Geotrupes) 

Scarabaeus antillarum Palisot de Beauvois, 1811 

Heteronychus tumulosus Burmeister, 1847 

Tomarus gibbosus (De Geer, 1774) (Scarabaeus) 

Scarabaeus juvencus Fabricius, 1775 

Bothynus morio LeConte, 1847 

Bothynus obsoletus LeConte, 1847 

Podalgus variolosus Burmeister, 1847 

Ligyrus californicus Casey, 1909 

Ligyrus spissipes Casey, 1909 

Ligyrus arizonensis Casey, 1915 

Ligyrus bicorniculatus Casey, 1915 

Ligyrus brevipes Casey, 1915 

Ligyrus breviusculus Casey, 1915 

Ligyrus curtipennis Casey, 1915 

Ligyrus effetus Casey, 1915 

84

Ligyrus farctus Casey, 1915 

Ligyrus lacustris Casey, 1915 

Ligyrus laetulus Casey, 1915 

Ligyrus laevicauda Casey, 1915 

Ligyrus laticauda Casey, 1915 

Ligyrus laticollis Casey, 1915 

Ligyrus longulus Casey, 1915 

Ligyrus lucublandus Casey, 1915 

Ligyrus parallelus Casey, 1915 

Ligyrus puncticauda Casey, 1915 

Ligyrus remotus Casey, 1915 

Ligyrus rubidus Casey, 1915 

Ligyrus scitulus Casey, 1915 

Ligyrus texanus Casey, 1915 

Ligyrus virginicus Casey, 1915 

Tomarus laevicollis (Bates, 1888) (Ligyrus) 

Ligyrus bryanti Rivers, 1891 

Tomarus nasutus (Burmeister, 1847) (Podalgus) 

Ligyrus pygidialis Bates, 1888 

Tomarus neglectus (LeConte, 1847) (Bothynus) 

Tomarus relictus (Say, 1825) (Scarabaeus) 

Ligyrodes clypealis Casey, 1915 

Ligyrodes parviceps Casey, 1915 

Ligyrodes quadripennis Casey, 1915 

Ligyrodes vernicicollis Casey, 1915 

Ligyrodes dawsoni Casey, 1924 

Tomarus sallaei (Bates, 1888) (Ligyrus) 

Ligyrodes aztecus Casey, 1915 

Ligyrodes propinquus Casey, 1915 

Tomarus selanderi (Cartwright, 1959) (Bothynus) 

Tomarus subtropicus (Blatchley, 1922) (Ligyrus) 

Ligyrus blatchleyi Cartwright, 1944 

ORYCTINI Mulsant, 1842 

Enema Hope, 1837 

Enema endymion Chevrolat, 1843 (Enema) 

Enema lupercus Burmeister, 1847 

Enema paniscus Burmeister, 1847 

Enema gibbicollis Sternberg, 1908 

Strategus Kirby, 1828 

Anastrategus Casey, 1915 

Strategodes Casey, 1915 

Strategopsis Chapin, 1932 

85

Strategus aloeus (Linnaeus, 1758) (Scarabaeus) 

Strategus julianus Burmeister, 1847 

Strategus arizonicus Schaeffer, 1915 

Strategus gaillardi Casey, 1915 

Strategus roosevelti Casey, 1915 

Strategus tarsalis Casey, 1915 

Strategus antaeus (Drury, 1773) (Scarabaeus) 

Scarabaeus maimon Fabricius, 1775 

Strategus atrolucens Casey, 1915 

Strategus divergens Casey, 1915 

Strategus pinorum Casey, 1915 

Strategus semistriatus Casey, 1915 

Strategus septentrionis Casey, 1915 

Strategus sinuatus Casey, 1915 

Strategus houstonensis Knaus, 1925 

Strategus cessus LeConte, 1866 (Strategus) 

Strategus beckeri Kolbe, 1906 

Anastrategus cavicauda Casey, 1915 

Anastrategus durangoensis Casey, 1915 

Anastrategus inflatus Casey, 1915 

Anastrategus tantalus Casey, 1915 

Strategus craigi Ratcliffe, 1976 (Strategus) 

Strategus howdeni Ratcliffe, 1976 (Strategus) 

Strategus longichomperus Ratcliffe, 1976 (Strategus) 

Strategus mormon Burmeister, 1847 (Strategus) 

Strategus splendens (Palisot de Beauvois, 1809) (Scarabaeus) 

Scarabaeus boscii Palisot de Beauvois, 1809 

Anastrategus carolinensis Casey, 1915 

Anastrategus cognatus Casey, 1915 

Strategus temoltzin Morón and Nogueira, 2008 (Strategus) 

Xyloryctes Hope, 1837 

Xyloryctes ensifer Bates, 1888 (Xyloryctes) 

Xyloryctes laevipennis Sternberg, 1908 

Xyloryctes perpendicularis Sternberg, 1908 

Xyloryctes jamaicensis (Drury, 1773) (Scarabaeus) 

Scarabaeus satyrus Fabricius, 1775 

Scarabaeus americanus Palisot de Beauvois, 1807 

Xyloryctes faunus Casey, 1895 

Xyloryctes hebes Casey, 1915 

Xyloryctes lacustris Casey, 1915 

Xyloryctes obsolescens Casey, 1915 

Xyloryctes tenuicornutus Casey, 1915 

Xyloryctes telephus Burmeister, 1847 (Xyloryctes) 

86

Xyloryctes thestalus borealis Endrödi, 1975 (Xyloryctes) 

Xyloryctes thestalus thestalus Bates, 1888 (Xyloryctes) 

Xyloryctes intermedia Sternberg, 1908 

PHILEURINI Burmeister, 1847 

Archophileurus Kolbe, 1910 

Amblyphileurus Kolbe, 1910 

Periphileurus Kolbe, 1910 

Anisophileurus Prell, 1914 

Archophileurus cribrosus (LeConte, 1854) (Phileurus) 

Archophileurus brevis Casey, 1915 

Archophileurus bullatus Casey, 1915 

Archophileurus longulus Casey, 1915 

Hemiphileurus Kolbe, 1910 

Epiphileurus Kolbe, 1910 

Hemiphileurus illatus (LeConte, 1854) (Phileurus) 

Phileurus vitulus LeConte, 1863 

Phileurus phoenicus Casey, 1915 

Phileurus puncticollis Casey, 1915 

Phileurus Latreille, 1807 

Phileurus truncatus (Palisot de Beauvois, 1806) (Scarabaeus) 

Phileurus recurvatus Casey, 1915 

Phileurus valgus (Olivier, 1789) (Scarabaeus) 

Phileurus castaneus Haldeman, 1843 

Phileurus capra Bates, 1888 

Epiphileurus septentrionis Kolbe, 1910 

Phileurus carolinae Casey, 1915 

Phileurus floridanus Casey, 1915 

Phileurus sulcifer Casey, 1915 

Phileurus texensis Casey, 1915 

DYNASTINI MacLeay, 1819 

Dynastes MacLeay, 1819 

Theogenes Burmeister, 1847 

Dynastes granti Horn, 1870 (Dynastes) 

Dynastes hyllus Chevrolat, 1843 (Dynastes) 

Dynastes tityus (Linnaeus, 1767) (Scarabaeus) 

Scarabaeus marianus Linnaeus, 1767 

87

Scarabaeus pennsylvanicus De Geer, 1774 

Dynastes corniger Sternberg, 1910 

Golofa Hope, 1837 

Golofa (Golofa) Hope, 1837 

Asseroder Maunders, 1848 

Golofa (Golofa) imperialis Thomson, 1858 (Golofa) 

Golofa (Golofa) xiximeca Morón, 1995 (Golofa) 

Megasoma Kirby, 1825 

Megalosoma Burmeister, 1847 

Lycophontes Bruch, 1910 

Megasominus Casey, 1915 

Megasoma cedrosum Hardy, 1972 (Megasoma) 

Megasoma lecontei Hardy, 1972 (Megasoma) 

Megasoma lenczyi Cartwright, 1976 (Megasoma) 

Megasoma occidentalis Bolívar y Pieltain, Jiménez-Asúa, and Martínez, 1963 (Megasoma) 

Megasoma pachecoi Cartwright, 1963 (Megasoma) 

Megasoma punctulatum Cartwright, 1952 (Megasoma) 

Megasoma sleeperi Hardy, 1972 (Megasoma) 

Megasoma thersites LeConte, 1861 (Megasoma) 

Megasoma vogti Cartwright, 1963 (Megasoma) 

CETONIINAE Leach, 1815 

VALGINI Mulsant, 1842 

Valgus Scriba, 1790 

Acanthurus Kirby, 1827 

Homovalgus Kolbe, 1897 

Valgus californicus Horn, 1870 (Valgus) 

Valgus canaliculatus (Olivier, 1789) (Cetonia) 

Trichius variegatus Palisot de Beauvois, 1805 

Valgus serricollis Fitch, 1858 

Valgus minutus Casey, 1909 

Valgus hemipterus (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus variegatus Scopoli, 1763 

Valgus rufosquamatus Dalla Torre, 1879 

Valgus rubi Baguena, 1955 

Valgus seticollis (Palisot de Beauvois, 1805) (Trichius) 

Trichius squamiger Palisot de Beauvois, 1805 

TRICHIINI Fleming, 1821 

88

Apeltastes Howden, 1968 

Apeltastes elongatus Howden, 1968 (Apeltastes) 

Gnorimella Casey, 1915 

Gnorimella maculosa (Knoch, 1801) (Cetonia) 

Trichius bigsbii Kirby, 1827 

Gnorimus dissimilis Gory and Percheron, 1833 

Trichiotinus Casey, 1915 

Trichinus Kirby, 1827 

Trichiotinus affinis (Gory and Percheron, 1833) (Trichius) 

Trichius variabilis Burmeister and Schaum, 1841 

Trichius mutabilis Schaum, 1844 

Trichiotinus venticosus Casey, 1915 

Trichiotinus parvulus Casey, 1915 

Trichiotinus assimilis (Kirby, 1837) (Trichius) 

Trichius bistriga Newman, 1838 


Trichiotinus bibens (Fabricius, 1775) (Trichius) 

Trichiotinus lunulatus (Fabricius, 1775) (Trichius) 

Trichius viridulus Fabricius, 1775 

Trichius virens Gmelin, 1790 

Trichius mutabilis Schaum, 1844 

Trichius semiviridis Casey, 1914 

Trichius carolinensis Casey, 1914 

Trichius rasilicaudus Casey, 1915 

Trichius rufiventris Casey, 1915 

Trichiotinus piger (Fabricius, 1775) (Trichius) 

Trichius drummond Gory and Percheron, 1833 

Trichius rotundicollis Kirby, 1837 

Trichiotinus reductus Casey, 1915 

Trichiotinus rufobrunneus (Casey, 1914) (Trichius) 

Trichius obesulus Casey, 1914 

Trichiotinus texanus (Horn, 1876) (Trichius) 

Trichiotinus monticola Casey, 1915 

Trichiotinus intermedius Casey, 1915 

Trichiotinus viridans (Kirby, 1837) (Trichius) 


Trigonopeltastes Burmeister, 1840 

Archimedius Kirby, 1827 

Euclidius Kirby, 1827 

89

Roplisa Casey, 1909 

Trigonopeltastes archimedes Schaum, 1841 (Trigonopeltastes) 

Trigonopeltastes delta (Forster, 1771) (Scarabaeus) 

Trigonopeltastes discrepans Howden, 1968 (Trigonopeltastes) 

Trigonopeltastes floridanus (Casey, 1909) (Roplisa) 

Trigonopeltastes frontalis Bates, 1889 (Trigonopeltastes) 

Trigonopeltastes sallaei sinaloensis Howden, 1968 (Trigonopeltastes) 

Trigonopeltastes truncatus Howden, 1968 (Trigonopeltastes) 

INCAINI Burmeister, 1842 

Inca LePeletier and Serville, 1828 

Inca clathrata sommeri Westwood, 1845 (Inca) 

OSMODERMATINI Schenkling, 1922 

Osmoderma LePeletier and Serville, 1828 

Gymnodus Kirby, 1827 

Osmoderma eremicola (Knoch, 1801) (Cetonia) 

Osmoderma scabra (Palisot de Beauvois, 1805) (Trichius) 

Trichius foveatus Kirby, 1837 

Trichius rugosus Kirby, 1837 

Osmoderma beauvoisii Bainbridge, 1841 

Osmoderma delicatula Casey, 1915 

Osmoderma gracilipes Casey, 1915 

Osmoderma caviceps Casey, 1915 

Osmoderma lacustrina Casey, 1915 

Osmoderma subplanata Casey, 1915 (Osmoderma) 

Osmoderma montana Wickham, 1920 

CREMASTOCHEILINI Burmeister and Schaum, 1841 

CREMASTOCHEILINA Burmeister and Schaum, 1841 

Centrochilus Krikken, 1976 

Centrochilus howdeni Krikken, 1976 (Centrochilus) 

Cremastocheilus Knoch, 1801 

Cremastocheilus (Anatrinodia) Casey, 1915 

Cremastocheilus (Cremastocheilus) Knoch, 1801 

Cremastocheilus (Macropodina) Knoch, 1801 

Cremastocheilus (Myrmecotonus) Mann, 1914 

90

Myrmeceicon Mann, 1914 

Cremastocheilus (Trinodia) Knoch, 1801 

Cremastocheilus (Trinodia) academicus Krikken, 1982 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) angularis LeConte, 1857 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) armatus armatus Walker, 1866 (Cremastocheilus) 

Cremastocheilus pilosicollis Horn, 1871 

Cremastocheilus (Myrmecotonus) armatus cribripennis Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) armatus maritimus Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) armatus montanus Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Macropodina) beameri Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) canaliculatus Kirby, 1827 (Cremastocheilus) 

Cremastocheilus hentzii Harris, 1827 

Cremastocheilus (Cremastocheilus) castaneae brevisetosus Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) castaneae castaneae Knoch, 1801 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) castaneae lecontei Westwood, 1874 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) castaneae pocularis Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) chapini Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) congener Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Trinodia) constricollis Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) crinitus bifoveatus Van Dyke, 1918 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) crinitus crinitus LeConte, 1874 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) crinitus pugetanus Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Trinodia) excavatus Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) harrisii Kirby, 1827 (Cremastocheilus) 

Cremastocheilus (Trinodia) hirsutus Van Dyke, 1918 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) knochii LeConte, 1853 (Cremastocheilus) 

Cremastocheilus crenicollis Westwood, 1874 

Cremastocheilus gracilipes Casey, 1915 

Cremastocheilus areolatus Casey, 1915 

Cremastocheilus (Trinodia) lengi Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Trinodia) mentalis Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) mexicanus Schaum, 1841 (Cremastocheilus) 

Cremastocheilus villadae Dugés, 1879 

Cremastocheilus (Cremastocheilus) nitens LeConte, 1853 (Cremastocheilus) 

Cremastocheilus (Trinodia) opaculus Horn, 1894 (Cremastocheilus) 

Cremastocheilus (Macropodina) planatus LeConte, 1863 (Cremastocheilus) 

Cremastocheilus (Trinodia) planipes Horn, 1885 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) pulverulentus Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Macropodina) puncticollis Cazier, 1940 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) quadratus Fall, 1912 (Cremastocheilus) 

Cremastocheilus (Trinodia) quadricollis (Casey, 1915) (Trinodia) 

Cremastocheilus (Cremastocheilus) retractus incisus Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) retractus retractus LeConte, 1874 (Cremastocheilus) 

Cremastocheilus walshii Westwood, 1874 

Cremastocheilus (Myrmecotonus) robinsoni Cazier, 1940 (Cremastocheilus) 

91

Cremastocheilus (Trinodia) saucius LeConte, 1858 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) schaumii schaumii LeConte, 1853 (Cremastocheilus) 

Cremastocheilus crassipes Westwood, 1874 

Cremastocheilus (Myrmecotonus) schaumii tibialis Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Trinodia) setosifrons (Casey, 1915) (Trinodia) 

Cremastocheilus (Trinodia) spinifer Horn, 1885 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) squamulosus LeConte, 1858 (Cremastocheilus) 

Cremastocheilus junior Westwood, 1874 

Cremastocheilus (Trinodia) stathamae Cazier, 1961 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) tomentosus Warner, 1985 (Cremastocheilus) 

Cremastocheilus (Cremastocheilus) variolosus Kirby, 1826 (Cremastocheilus) 

Cremastocheilus sayi Harris, 1827 

Cremastocheilus cicatricosus Westwood, 1874 

Cremastocheilus percheronii Westwood, 1874 

Cremastocheilus (Myrmecotonus) westwoodi tridens Casey, 1915 (Cremastocheilus) 

Cremastocheilus (Myrmecotonus) westwoodi westwoodi Horn, 1880 (Cremastocheilus) 

Cremastocheilus (Anatrinodia) wheeleri LeConte, 1876 (Cremastocheilus) 

Genuchinus Westwood, 1874 

Genuchinus ineptus (Horn, 1885) (Cremastocheilus) 

Genuchinus angustus Casey, 1915 

Lissomelas Bates, 1889 

Lissomelas flohri Bates, 1889 (Lissomelas) 

Genuchinus grandis Van Dyke, 1952 

Psilocnemis Burmeister, 1842 

Psilocnemis leucosticta Burmeister, 1842 (Psilocnemis) 

CETONIINI Leach, 1815 

CETONIINA Leach, 1815 

Chlorixanthe Bates, 1889 

Chlorixanthe propinqua (Gory and Percheron, 1833) (Cetonia) 

Chlorixanthe chapini Cartwright, 1939 

Euphoria Burmeister, 1842 

Erirhipis Burmeister, 1842 

Stephanucha Burmeister, 1842 

Goraqua Péringuey, 1907 

Anatropis Casey, 1915 

Erirhipidia Casey, 1915 

92

Euphorhipis Casey, 1915 

Euphoriaspis Casey, 1915 

Euphoriopsis Casey, 1915 

Haplophoria Casey, 1915 

Isorhipina Casey, 1915 

Parisorhipis Casey, 1915 

Rhipiphoria Casey, 1915 

Euphoria anneae (Howden, 1955) (Stephanucha) 

Euphoria areata (Fabricius, 1775) (Cetonia) 

Stephanucha thoracica Casey, 1915 

Euphoria basalis (Gory and Percheron, 1833) (Cetonia) 

Euphoria crinicauda Casey, 1915 

Euphoria biguttata (Gory and Percheron, 1833) (Cetonia) 

Euphoria binoculata Casey, 1915 

Euphoria biplagiata Casey, 1915 

Euphoria bispinis (Bates, 1889) (Stephanucha) 

Euphoria canescens (Gory and Percheron, 1833) (Cetonia) 

Euphoria solidula Casey, 1915 

Euphoria casselberryi Robinson, 1937 (Euphoria) 

Euphoria devulsa Horn, 1880 (Euphoria) 

Euphoria discicollis (Thomson, 1878) (Stephanucha) 

Stephanucha atrata Thomson, 1878 

Euphoria aestuosa Horn, 1880 

Euphoria fascifera fascifera (LeConte, 1861) (Euryomia) 

Euphoria fascifera trapezium Casey, 1915 (Euphoria) 

Euphoria fulgida fulgida (Fabricius, 1775) (Cetonia) 

Scarabaeus turchesinus Voet, 1778 

Euphoria fulgida fuscocyanea Casey, 1915 (Euphoria) 

Euphoria fulgida limbalis Fall, 1905 (Euphoria) 

Euphoria fulgida holochloris Fall, 1905 (Euphoria) 

Euphoria geminata (Chevrolat, 1835) (Cetonia) 

Euphoria herbacea (Olivier, 1789) (Cetonia) 

Cetonia antennata Gory and Percheron, 1833 

Cetonia pubera Gyllenhal, 1817 

Erirhipis occidentalis Knaus, 1916 

Euphoria hirtipes Horn, 1880 (Euphoria) 

Euphoria histrionica Thomson, 1878 (Euphoria) 

Euphoria sonorae Bates, 1889 

Euphoria scabiosa Casey, 1915 

Euphoria inda (Linnaeus, 1758) (Scarabaeus) 

Cetonia marilandica Frölich, 1792 

Cetonia barbata Say, 1824 

Cetonia brunnea Gory and Percheron, 1833 

Euphoria nigripennis Klages, 1894 

Goraqua smithsana Péringuey, 1907 

93

Euphoria rufobrunnea Casey, 1915 

Euphoria kernii Haldeman, 1852 (Euphoria) 

Erirhipis clarkii LeConte, 1854 

Euphoria texana Schaufuss, 1863 

Euphoria connivens Casey, 1915 

Euphoria retusa Casey, 1915 

Euphoria wichitana Casey, 1915 

Euphoria leucopyge Bates, 1889 (Euphoria) 

Euphoria lineoligera (Blanchard, 1850) (Cetonia) 

Euphoria xanthomelas Thomson, 1878 

Euphoria monticola Bates, 1889 (Euphoria) 

Euphoria pilipennis (Kraatz, 1883) (Stephanucha) 

Euphoria pulchella (Gory and Percheron, 1833) (Cetonia) 

Cetonia childrenii Gory and Percheron, 1833 

Cetonia ferrugata Gory and Percheron, 1833 

Cetonia montesuma Gory and Percheron, 1833 

Cetonia thelasco Gory and Percheron, 1833 

Euphoria quadricollis Bates, 1889 (Euphoria) 

Euphoria arizonica Schaeffer, 1907 

Euphoria schottii (LeConte, 1854) (Erirhipis) 

Euphoria sepulcralis leucographa (Gory and Percheron, 1833) (Cetonia) 

Euphoria limatula Janson, 1881 

Euphoria yucateca Bates, 1889 

Euphoria submetallica Casey, 1915 

Euphoria sepulcralis nitens Casey, 1915 (Euphoria) 

Euphoria sepulcralis rufina (Gory and Percheron, 1833) (Cetonia) 

Euphoria histrionella Schaeffer, 1907 

Euphoria testacea Casey, 1915 

Euphoria sepulcralis sepulcralis (Fabricius, 1801) (Cetonia) 

Cetonia reichii Gory and Percheron, 1833 

Euphoria appalachia Casey, 1915 

Euphoria crinitula Casey, 1915 

Euphoria cuprascens Casey, 1915 

Euphoria floridana Casey, 1915 

Euphoria kansana Casey, 1915 

Euphoria oxysternum Casey, 1915 

Euphoria scolopacea Casey, 1915 

Euphoria verticalis Horn, 1880 (Euphoria) 

Protaetia Burmeister, 1842 

Protaetia (Heteroprotaetia) Miksic, 1963 

Protaetia (Heteroprotaetia) fusca (Herbst, 1790) (Cetonia) 

Cetonia mandarina Lichtenstein, 1796 

Cetonia atomaria Fabricius, 1801 

94

Cetonia fictilis Newman, 1838 

Protaetia taiwana Niijima and Kinoshita, 1923 

Protaetia bourgoini Paulian, 1960 

GYMNETINI Kirby, 1827 

BLAESIINA Schoch, 1895 

Halffterinetis Morón and Nogueira, 2007 

Halffterinetis gonzaloi Morón and Nogueira, 2007 (Halffterinetis) 

Halffterinetis violetae Morón and Nogueira, 2007 (Halffterinetis) 

GYMNETINA Kirby, 1827 

Cotinis Burmeister, 1842 

Cotinis (Cotinis) Burmeister, 1842 

Latemnis Thomson, 1880 

Cotinorrhina Schoch, 1895 

Cotinis (Criniflava) Goodrich, 1966 

Cotinis (Cotinis) aliena Woodruff, 2008 (Cotinis) 

Cotinis (Cotinis) antonii (Dugés, 1878) (Gymnetis) 

Cotinis senex Janson, 1880 

Cotinis (Criniflava) boylei Goodrich, 1966 (Cotinis) 

Cotinis (Cotinis) impia (Fall, 1905) (Gymnetis) 

Cotinis (Cotinis) laticornis Bates, 1889 (Cotinis) 

Cotinis latifrons Schoch, 1895 

Cotinis (Cotinis) mutabilis (Gory and Percheron, 1833) (Gymnetis) 

Gymnetis atrata Gory and Percheron, 1833 

Gymnetis nigrorubra Gory and Percheron, 1833 

Gymnetis mexicana Gory and Percheron, 1833 

Gymnetis palliata Gory and Percheron, 1833 

Gymnetis sobrina Gory and Percheron, 1833 

Cotinis cabira Burmeister, 1842 

Cotinis malinus Janson, 1889 

Cotinis cuprascens Bates, 1889 

Cotinis subcastanca Bates, 1889 

Cotinis intergenea Bates, 1889 

Cotinis aurantica Bates, 1889 

Cotinis robusta Bates, 1889 

Cotinis schafraneki Nonfried, 1894 

Cotinis piciventris Kraatz, 1898 

Cotinis nigrovariegata Kraatz, 1898 

Cotinis cuprea Kraatz, 1898 

Cotinis atropurpurea Kraatz, 1898 

95

Cotinis atra Kraatz, 1898 

Cotinis texana Casey, 1915 

Cotinis arizonica Casey, 1915 

Cotinis abdominalis Casey, 1915 

Cotinis discolor Casey, 1915 

Cotinis obliqua Casey, 1915 

Cotinis coahuilae Casey, 1915 

Cotinis viridicauda Casey, 1915 

Cotinis commiscens Casey, 1915 

Cotinis ovicornuta Casey, 1915 

Cotinis capito Casey, 1915 

Cotinis (Cotinis) nitida (Linnaeus, 1758) (Scarabaeus) 

Scarabaeus flagranticeps Voet, 1766 

Cotinis debiliceps Casey, 1915 

Cotinis longula Casey, 1915 

Cotinis angustula Casey, 1915 

Cotinis parvula Casey, 1915 

Cotinis ornata Casey, 1915 

Cotinis tibialis Casey, 1915 

Cotinis pygidialis Casey, 1915 

Cotinis longitarsus Casey, 1915 

Cotinis seriella Casey, 1915 

Cotinis brunnea Hatch, 1930 

Cotinis (Cotinis) orientalis Deloya and Ratcliffe, 1988 (Cotinis) 

Cotinis (Criniflava) producta Bates, 1889 (Cotinis) 

Cotinis (Cotinis) rufipennis Bates, 1889 (Cotinis) 

Cotinis (Cotinis) sinitoc Deloya, Ibáñez-Bernal, and Nogueira, 2000 (Cotinis) 

Gymnetina Casey, 1915 

Gymnetina cretacea (LeConte, 1866) (Gymnetis) 

Gymnetis MacLeay, 1819 

Gymnetoides Martínez, 1949 

Gymnetosoma Martínez, 1949 

Gymnetis caseyi Antoine, 2001 (Gymnetis) 

Hologymnetis Martínez, 1949 

Cineretis Krikken, 1984 

Hologymnetis argenteola (Bates, 1889) (Gymnetis) 

Gymnetis laetula Casey, 1915 

Hologymnetis cinerea (Gory and Percheron, 1833) (Gymnetis) 

Gymnetis punctata Blanchard, 1850 

Gymnetis uniformis Blanchard, 1850 

96

Gymnetis aequalis Casey, 1915 

Gymnetis cuneata Casey, 1915 

Gymnetis lobiculata Casey, 1915 

Gymnetis simulans Casey, 1915 

97

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