Checklist of the Plants of the Guianas - Department of Botany ...

CHECKLIST OF THE PLANTS OF THE GUIANAS 

(Guyana, Surinam, French Guiana) 

J. Boggan, V. Funk, C. Kelloff 

Smithsonian Institution 

and 

M. Hoff, G. Cremers, C. Feuillet 

ORSTOM 

2nd Edition 

Prepared under the auspices of the Centre for the Study of Biological Diversity 

University of Guyana, Georgetown, Guyana



2nd Edition 

J. Boggan, V. Funk, C. Kelloff 

Biological Diversity of the Guianas 

Department of Botany 

National Museum of Natural History 


Washington, D.C. 20560 U.S.A. 

and 

M. Hoff, G. Cremers, C. Feuillet 

Herbarium, ORSTOM 

97323 Cayenne Cedex 

French Guiana, FRANCE 

1997 

Prepared under the auspices of the Centre for the Study of Biological Diversity 

University of Guyana, Georgetown, Guyana

CHECKLIST OF THE PLANTS OF GUIANA 


2nd Edition 

1st Edition: December 1992 

2nd Edition: February 1997 

Published by the Biological Diversity of the Guianas Program, Department of 

Botany, National Museum of Natural History, Smithsonian Institution, 

Washington, D.C. 20560 U.S.A. This is publication number 30 in the 

Smithsonian's Biological Diversity of the Guianas Program publication series. 

Copies available at no charge while supplies last. 

Printed in the U.S.A. 

This publication has been printed on acid free paper 

Cover illustration: Phenakospermum guyannense (L.C. Rich.) Endl. ex Miq. (Strelitziaceae), 

drawn by Alice Tangerini, Smithsonian Institution.

CONTENTS 

INTRODUCTION 1 

Geology 2 

Vegetation types 2 

Some current programs and projects in the Guianas 5 

Explanation of checklist and editorial notes 5 

Acknowledgements 7 

References 8 

CHECKLIST 9 

Chlorophyta-Characeae (stoneworts) 9 

Hepatophyta (liverworts) 9 

Anthocerophyta (hornworts) 15 

Bryophyta (mosses) 15 

Psilotophyta (psilopsids) 22 

Lycophyta (lycophytes) 22 

Pterophyta (ferns) 24 

Spermatophyta (seed plants) 40 

INDEX 211 

APPENDIX A. Contributors and Reviewers 229 

APPENDIX B. Contributors and reviewers for the revised edition 237

PREFACE TO THE REVISED EDITION 

Since its publication in 1992, numerous 

additions and corrections have been made to the 

Checklist of the Plants of the Guianas. We 

have incorporated these changes in a revised 

edition, along with some changes in the format 

that we hope will make the checklist easier to 

use. Additions have included newly-described 

species, new records for the Guianas, and names 

overlooked or otherwise omitted from the first 

edition. Synonymy and reference lists have been 

expanded. In several cases (e.g., Nyctaginaceae, 

Turneraceae), the species list for a family has 

been completely revised. All the spermatophytes 

(seed plants) are arranged together alphabetically 

by family. The country distribution codes have 

been changed to GU, SU, and FG for Guyana, 

Surinam, and French Guiana, respectively, in 

anticipation of a checklist including the 

"Guayanan" portions of Venezuela and Brasil. 

The treatment of the various families remains 

uneven, with some families well-known (either 

with recent monographic or floristic treatments, 

or with a specialist actively identifying 

specimens) and others (e.g., Dioscoreaceae, 

Smilacaceae) very poorly known. All available 

publications dealing with Guianan plants since 

publication of the first edition have been 

consulted, in some cases resulting in substantial 

changes; in other cases, older publications not 

seen for the original checklist were consulted 

with the result that certain families are in much 

more complete (e.g., Liliaceae s.l. but excluding 

Smilacaceae). 

We have tried not to include any unpublished 

names or combinations without indicating them 

as such (i.e., "sp. nov. ined." or "comb. nov. 

ined."). However, it has not been possible to 

verify every name found on herbarium 

specimens or in the literature as validly 

published. The editors regret and take full 

responsibility for the use of any invalid names. 

by John Boggan 

iii 

The first edition of the "Checklist of the 

Plants of the Guianas" was a collaborative 

project, with contributions by many researchers 

worldwide. Authors of the family lists were 

identified as "contributed by...." Otherwise, 

family lists were compiled by the editors, and in 

some cases these were reviewed by additional 

specialists (identified as "reviewed by...."). 

Additions and corrections suggested by the 

reviewers were incorporated. Authorship of the 

revised edition is complicated by the fact that 

additions and corrections were made by the 

editors, by most (but not all) of the original 

contributors and reviewers (listed in Appendix 

A), and by new contributors and reviewers 

(listed in Appendix B). The editors have tried to 

clearly indicate the authorship of each family 

list, and hope we have given sufficient credit to 

the contributors. 

It is the nature of publications such as this to 

be obsolete as soon as they are published. 

During the time between the completion of 

editing and final publication, several new species 

have been published, and new country records 

have been discovered. In addition, numerous 

additions and corrections were received from Jan 

C. Lindeman (U) shortly before going to press. 

Unfortunately, there was not enough time to 

incorporate most of these in the revised 

checklist. The majority of those not already 

received from other sources were synonyms 

whose current status could not be verified. New 

country records indicated by Lindeman 

(primarily for Surinam) are indicated with a 

question mark. 

This is publication number 30 in the 

Smithsonian's Biological Diversity of the 

Guianas Program publication series. This 

publication was supported by the BDG program 

and copies are available from the program at no 

charge while supplies last.


This checklist is a preliminary listing of the 

known vascular plants (flowering plants, 

gnetophytes, cycads, conifers, ferns and "fern 

allies") and "bryophytes" (mosses, hornworts, 

liverworts) from the Guianas. Although our 

knowledge of the algae is insufficient to enable 

us to produce a species list at this time, we have 

included the Characeae (stoneworts) of the 

macroalgae because the information was easily 

available. This checklist excludes lichens and 

fungi, which are usually classified together 

under fungi and are not plants in the strict sense. 

A checklist of the lichens reported from the 

Guianas before 1987 has been published 

(Hekking & Sipman 1988) and hopefully, a list 

of all the fungal groups will be forthcoming. 

This checklist is an attempt to list all of the 

vascular plants and bryophytes of the Guianas in 

an effort to encourage further research. 

Checklists have many uses. They can aid in 

identifications, although new species and new 

records are always a possibility. In addition, 

checklists tell us how many organisms of a 

certain kind are in a particular area, thus giving 

us an idea of the species richness. Checklists 

can act as an indicator of endemicity within an 

area and provide information on the number of 

introduced species. They can be used in a 

comparative manner between and among areas. 

If they cover a natural area (one that has biotic or 

geologic significance) they can be used to 

formulate questions about centers of origin and 

evolutionary history. In addition, a checklist is a 

way to help standardize spelling and authors and 

to promote accurate labeling of specimens. 

Checklists, however, reflect our knowledge at a 

particular time and our current taxonomic 

opinions and so their usefulness is tempered by 

the knowledge that they are never entirely 

correct and that the list will be constantly 

changing. For instance, we are sure that this list 

contains incorrect synonomies, records based on 

misidentifications, that additional species remain 

INTRODUCTION 

by V.A. Funk 

to be added, and that many species have not yet 

been described or even discovered. However, 

this type of "snapshot" is valuable because it 

provides a way of focusing and stimulating work 

in the area. In order to help us keep this 

checklist as current and accurate as possible, 

please contact the Director, Biological Diversity 

of the Guianas Program, with any additions or 

corrections. 

The part of the northeastern shoulder of 

South America along the Atlantic coast 

commonly known as the "Guianas" consists of 

three political areas (Map 1) with a total area of 

about 500,000 sq. km. The westernmost 

political entity is Guyana, formerly known as 

British Guiana, which has 800,000 inhabitants 

and an area of 231,800 sq. km. To the east of 

Guyana is Surinam, formerly known as Dutch 

Guiana, with 400,000 inhabitants and 173,840 

sq. km. The easternmost is French Guiana, an 

overseas department of France, with 120,000 

inhabitants and 88,240 sq. km. (areas from 

McConnell and Choubert 1975). In all three of 

these areas, the population is concentrated along 

the coast and near the respective capital cities. 

The Guianas constitute only part of a 

phytogeographic or natural area called the 

Guiana Shield (see Geology section). To make 

them a natural area would require including 

Venezuela south and east of the Orinoco River 

and small parts of Colombia and Brazil. During 

the next year the "Flora of the Venezuelan 

Guayana" project of the Missouri Botanical 

Garden will produce a checklist for the 

Venezuelan Guayana. Proposed collaboration 

between the Smithsonian and Missouri will 

produce a combined checklist covering the bulk 

of the Guiana Shield a natural area to study and 

determine if the flora of the Guiana shield is a 

unique evolutionary unit. 

1 

INTRODUCTION

2 

INTRODUCTION 


For this checklist, we have recorded over 

9200 species from the three Guianas: 

approximately 6500 from Guyana, 5100 from 

Surinam, and 5400 from French Guiana. Of 

these, only about 3% are introduced and 

naturalized. The families with the largest 

number of species in the Guianas are the 

Leguminosae (Cronquist, 1981, divides the 

legumes into three segregate families but they 

are often treated as one) with more than 800 

species and the Orchidaceae with about 700. 

Other large families are the Rubiaceae and the 

Poaceae (grasses), both with more than 400 

species. 

GEOLOGY 

Underlying all of northeastern South 

America is the Guiana Shield (Map 2), which 

occupies a broad area between the Atlantic 

Ocean and the Orinoco and Amazon rivers. In 

addition to the three Guianas, the Guiana Shield 

covers a small part of eastern Colombia and the 

Venezuelan departments of Amazonas and 

Bolivar to the west and parts of the Brazilian 

departments of Roraima, Pará and Amapá to the 

south and east. Geologically, the Guiana Shield 

is an ancient Precambrian land mass (4 billion - 

590 million years old) made up of varied 

formations of sedimentary and igneous origin 

that were metamorphosed and folded 

(McConnell and Choubert 1975). Overlying the 

central portion of the Shield are the Roraima 

sediments that, although lacking in fossils, are 

believed to have been laid down on the Guiana 

Shield during the Cretaceous Period (140 - 68 

million years ago) probably as shallow marine or 

brackish water deposits (Gansser 1954, Maguire 

1970). The Roraima formation consists of pink, 

yellow and white sandstones, red quartzites, 

green, black and red shales, conglomerates and 

boulder beds (Fanshawe 1952). At an earlier 

time, the Roraima formation must have been 

considerably larger, perhaps extending south to 

the Matto Grosso highlands in Brazil (Fanshawe 

1952, Gansser 1954). Erosion decreased the size 

of the formation and the remaining sediments 

extend west from the Kaieteur escarpment in 

central Guyana, forming the Pakaraima 

Mountains, and on through parts of Venezuela 

and Colombia and south into northern Brazil 

(Leechman 1913). Within this area, erosion has 

created vertical-walled, more or less flat-topped 

peaks called "tepui". These table-like formations 

are often virtually inaccessible and are so 

unusual that in a fictional account of a scientific 

expedition, one of them was referred to as "The 

Lost World" (Doyle 1912). Their unique flora 

and fauna has resulted in the descriptor "lost 

world" being applied to all of these table 

mountains. The highest peaks of the Guianas are 

found in the Roraima Formation in western 

Guyana with Mt. Roraima reaching 2772 m., Mt. 

Ayanganna to 2134 m., and Mt. Wokumung to 

2042 m. 

The Guiana Shield can be divided into four 

natural geologic regions: 1) coastal plain; 2) 

central pediplains and rugged mountains of 

Precambrian metamorphic and granitic rocks; 3) 

high plateaus of the Pakaraima Mountains, 

formed by the tabular Roraima Formation; and 

4) southern pediplains descending to the Rio 

Negro and the Amazon basin. 

VEGETATION TYPES 

The vegetation of the Guianas is determined 

by the underlying geology, altitude, and annual 

amount of rainfall. All of the Guianas 

experience at least one dry season a year and 

during most years, two. As with most tropical 

countries, the rains sometimes come early or 

sometimes late, and occasionally a dry or rainy 

season does not occur at all. In general, there is 

sufficient rainfall for most of the Guiana area to 

be forested. The estimated percentage of forest 

cover varies from 86% for Guyana (Haman and 

Wood 1928) to 97.7% for French Guiana 

(Granville 1974). The Guianas have a 

remarkable diversity of organisms and their rich 

flora and fauna remain largely unexplored and 

unexploited. They constitute one of the few 

tropical areas worldwide that still has the 

majority of its forests intact. Estimates vary, but 

clearly only a small percentage of the Guianas 

has been deforested. 

There are nine vegetation types in the 

Guianas according to Lindeman and Mori


Map 1: The Guianas 

3 

INTRODUCTION

4 

INTRODUCTION 


Map 2: The Guiana Shield (McConnell & Choubert 1975)


(1989): 1. seasonal evergreen forest (or 

seasonal wet forest); 2. mangrove; 3. strand 

vegetation (beach vegetation); 4. marsh forests; 

5. swamp forests; 6. herbaceous swamps; 7. 

savannas; 8. montane vegetation; and 9. 

inselbergs (granitic outcrops). However, 

Lindeman and Mori's list focuses mainly on 

Surinam and French Guiana. Fanshawe (1952) 

lists 28 vegetation types in six categories for 

Guyana: 1. rain forest; 2. seasonal forest; 3. dry 

evergreen forest; 4. montane forest; 5. marsh 

forest (includes savannas); and 6. swamp forest 

(includes mangrove and herbaceous swamps 

types). Neither list covers all of the vegetation 

types with the Lindeman and Mori list lacking 

several forest types and the Fanshawe list 

missing the strand and inselberg vegetation. 

Combining the two lists gives a broader, more 

complete range of vegetation types found in this 

region. One possible way to combine the two 

lists is to use the 28 types of Fanshawe's and add 

the two missing categories from Lindeman and 

Mori. 

SOME CURRENT PROGRAMS 

AND PROJECTS IN THE 

GUIANAS 

Several programs and projects are now 

underway in the Guianas. A few that deal with 

floras are listed below. The first two 

collaborated to produce this checklist. 

Biological Diversity of the Guianas (BDG) - 

Smithsonian Institution. The BDG started in 

1983 and now operates from the "Centre for the 

Study of Biological Diversity" on the campus of 

the University of Guyana. The BDG program 

seeks to document and study the flora and fauna 

of the Guianas. Activities include collecting 

specimens to be housed at the Centre and 

training students and staff of the University as 

well as producing checklists, flora treatments, 

inventories, vegetation maps, and other 

publications such as a listing of the plants and 

animals of Kaieteur Falls National Park in 

Guyana. 

Herbarium of Cayenne - ORSTOM. The 

general herbarium was founded in 1965 by R. A. 

A. Oldeman to succeed the savanna (grass) 

collection established by J. Hoock between 1955 

and 1965. From the beginning, the activity of 

the laboratory focused on the floristic and 

ecological studies of the forests of French 

Guiana. Activities in Cayenne include collecting 

specimens in remote areas of the country, 

especially inselbergs, and producing treatments 

for flora projects. In 1988, a checklist of the 

Flowering Plants and Pteridophytes of French 

Guiana was published (Cremers et al. 1988). 

The herbarium of some 60,000 specimens is 

stored electronically (completed in 1988) in the 

data bank AUBLET. 

Flora of the Guianas. An international 

consortium of nine botanical institutions formed 

in 1983 to produce a written account of the 

plants of the Guianas. The Flora project has its 

editorial center at the University of Utrecht, The 

Netherlands. Some of the contributors to this 

checklist are also participating in the Flora of the 

Guianas project; however, this checklist is not 

part of the Flora nor is it associated with it in any 

way. 

Flora of Central French Guiana. A joint 

project by the New York Botanical Garden and 

ORSTOM to produce a flora of 50,000 hectares 

of rainforest near Saül, French Guiana. 

Iwokrama International Rainforest 

Programme. Located in the center of Guyana, 

the principal aim of the Iwokrama Programme is 

to demonstrate, through scientific research, 

methods of sustainable utilization of the forest 

for timber and other extractive resources while 

conserving biological diversity. 

Conservation International. Operates a 

"Guianas Regional Program" (consisting of 

Surinam and Guyana) to assist these countries in 

conservation matters and to conduct 

ethnobotanical studies. 

EXPLANATION OF CHECKLIST 

AND EDITORIAL NOTES 

The checklist includes the vascular plants 

and bryophytes of Guyana, Surinam and French 

Guiana. Although the checklist does not cover 

most algae, the Characeae are included. The 

lichens and fungi are not included. As this 

checklist is preliminary, the quality and coverage 

5 

INTRODUCTION

6 

INTRODUCTION 


of the families varies widely depending on the 

availability of recent floristic and/or 

monographic literature and the participation of 

specialists. However, contributing specialists 

sometimes have supplied data that could not be 

verified from the literature. At this point it isn't 

always clear what is new and/or unpublished 

information and what is based on literature the 

editors have not seen. 

Family circumscriptions of the flowering 

plants follow Cronquist (1981), except our 

recognition of the family Bonnetiaceae on the 

recommendation of A. Weitzman (personal 

communication). Pteridophytes follow the 

classification in the United States National 

Herbarium (US), as arranged by D. Lellinger. 

Other groups are as arranged by their respective 

contributors. 

Author abbreviations, when used, follow 

Stafleu and Cowan (1976-1988). "H.B.K." is 

cited throughout the checklist as "Kunth". 

A few taxon names are included in the list 

that we have been unable to verify. Some may 

be new records, most are probably 

misidentifications or synonyms and still others 

are names from the literature that the editors 

cannot verify. The "unverified names" were not 

contributed by the specialists. 

Sources of Information: Several sources 

were used for the data presented in the checklist: 

1) the literature: see references at the end of each 

family and in the General Reference section; 2) 

contributions by specialists, either as provided 

by an individual and therefore listed as 

"contributed by..." or a list compiled by the 

editors was reviewed or proofread by a specialist 

in that group, in which case it is listed as 

"reviewed by...."; 3) databases: Centre 

Technique Forestier Tropical - France, 

Herbarium of Cayenne - ORSTOM, Biological 

Diversity of the Guianas Program - Smithsonian 

Institution; and 4) herbarium records, i.e., 

specimens at: BRG, CAY, US, MO. All 

abbreviations for herbaria follow Holmgren et al. 

(1990). Families listed without contributors or 

reviewers were compiled by the editors. 

The bibliographies are not exhaustive. Only 

recent monographic and floristic literature 

(generally after 1900) relevant to the covered 

geographic area was surveyed (see references 

following the introduction and with each 

family). Family references have been added by 

the editors as an aid to the user of the checklist. 

Families without corresponding specialists have 

more extensive literature citations. If a family 

had a recent treatment (e.g., in the Flora of the 

Guianas (Görts-van Rijn 1985 - ), Flora of 

Suriname (Pulle et al. 1932-1984), Botany of the 

Guayana Highland (Maguire and Collaborators 

1953-1989), Flora Neotropica, or family 

monographs) then little additional work was 

done for the present list. 

In a few cases (Cyperaceae, Myrtaceae, some 

parts of Orchidaceae), separate lists were 

provided by more than one specialist. In all such 

cases, the lists were not 100% in agreement and 

the list presented is an amalgamation of the two. 

In the species lists in general, and the 

synonymies in particular (see below), it should 

be noted that different systematists will have 

different opinions on the status of particular 

names. 

Taxa collected on Mt. Roraima have caused a 

problem because this table-mountain is located 

on the borders of Guyana, Venezuela and Brazil. 

Specimens from this area may lack a country 

designation or it may be incorrect or unknown, 

especially in historical collections. To prevent 

confusion, taxa known in the Guianas only from 

Mt. Roraima have been listed as "Mt. Roraima" 

rather than Guyana. However, as we continue to 

explore the Pakaraima Mountains, some taxa 

previously known only from Mt. Roraima may 

also be found in Guyana. 

Included and excluded names: This checklist 

includes indigenous taxa and naturalized taxa (so 

noted when possible) as well as taxa from 

neighboring areas that may be expected in the 

Guianas. Infraspecific taxa at the level of 

subspecies and variety are included at the 

discretion of the contributors. Infraspecific taxa 

at the level of "forma" are excluded as well as 

names of varietal status not currently considered 

to be distinct from the "typical" variety. 

We have excluded taxa known only in 

cultivation in the Guianas. Although some of 

these may be found as escapees, they are


generally not well-established. DeFilipps (1992) 

has listed many of the cultivated plants in his 

recent publication. 

Synonymies and taxa not known in the 

Guianas: The synonymy is not meant to be 

exhaustive. We have included synonyms only if 

the synonymized name is found in recent 

literature. Because some of the synonymies are 

taken from the literature they may not all 

conform to the taxonomic concepts of the 

contributing specialist for that family. 

In many cases, a species has been reported 

from the Guianas either because of a broad 

species circumscription that has since been 

restricted (or as an infraspecific taxon that has 

since been raised to species status), or due to 

misidentifications or other errors. In either case, 

the name is listed at the end of the family with 

the notation "not in Guianas". 

ACKNOWLEDGMENTS 

Once again, we thank the many specialists 

who have contributed to this checklist. It would 

not have been possible without their assistance. 

Their names are listed at the beginning of the 

family list with which they assisted as well as in 

Appendix A. Sally Adkins, Keitha Dattilo, 

Susan Grose, Marilyn Hansel, Tom Hollowell, 

Mark Strong, and Fernanda Zermoglio assisted 

with the proofreading and final preparation of 

the manuscript. We also thank Robert A. 

DeFilipps and Jan C. Lindeman for their 

numerous comments on the entire checklist. We 

have received comments, corrections, and 

encouragement from many sources, and regret 

that we can not name them all. In addition we 

thank D. B. Lellinger and L. E. Skog for being 

especially helpful with the overall project 

7 

INTRODUCTION

8 

INTRODUCTION 


REFERENCES 

Cremers, G., C. Feuillet, J.J. de Granville, & M. 

Hoff. 1988. Liste des espèces de 

Phanérogames et de Ptéridophytes de 

Guyane française d'après l'Herbier du 

Centre ORSTOM de Cayenne. ORSTOM, 

Cayenne. 

Cronquist, A. 1981. An Integrated System of 

Classification of Flowering Plants. 

Columbia University Press, New York. 

DeFilipps, R.A. 1992. Ornamental Garden 

Plants of the Guianas: An historical 

perspective of selected garden plants from 

Guyana, Surinam and French Guiana. 

Department of Botany, Smithsonian 

Institution, Washington, DC. 

Doyle, A.C. 1912. The Lost World. Puffin 

Books, London. 

Fanshawe, D.B. 1952. The vegetation of British 

Guiana: A preliminary review. Imperial 

Forestry Institute paper no. 29: 1-96. 

Gansser, A. 1954. The Guiana Shield (South 

America). Eclog. Gelo. Helv. 47: 77-117. 

Görts-van Rijn, A.R.A. [ed.]. 1985- . Flora of 

the Guianas. Koeltz Scientific Books, 

Germany. 

Granville, J.J. de. 1974. Paysages végétaux de 

la Guyane. Pages 77-78 in Regards sur la 

France. Editions S.P.E.I., Paris. 

Haman, M. & B.R. Wood. 1928. The forests of 

British Guiana. Trop. Woods 15: 2-13. 

Hekking, W.H.A. & H.J.M. Sipman. 1988. The 

lichens reported from the Guianas before 

1987. Willdenowia 17: 193-228. 

Holmgren, P.K., N.H. Holmgren, & L.C. 

Barnett. 1990. Index Herbariorum Part I: 

The Herbaria of the World. International 

Association for Plant Taxonomy. 

Leechman, A. 1913. The British Guiana 

Handbook. "The Argosy" Co., Ltd., 

Georgetown, British Guiana and Dulau & 

Co., London. 

Lindeman, J.C. & S.A. Mori. 1989. The 

Guianas. In D.G. Campbell and H.D. 

Hammond [ed.], Floristic Inventory of 

Tropical Countries. New York Botanical 

Garden, New York. 

Maguire, B. 1970. On the flora of the Guayana 

Highland. Biotropica 2: 85-100. 

Maguire, B. & collaborators. 1953-1989. 

Botany of the Guayana Highland. Parts I- 

XIII. Mem. New York Bot. Gard. 

McConnell, R.B. & B. Choubert. 1975. Guiana 

Shield-Regional Survey. In R.W. Fairbridge 

[ed.], Encyclopedia of World Regional 

Geology, Part I: Western Hemisphere. 

Dowden, Hutchinson, & Ross, Inc., 

Stroudsburg, PA. 

Pulle, A., J. Lanjouw, A.L. Stoffers, & J.C. 

Lindeman. 1932-1984. Flora of Suriname. 

J.H. deBussy, Amsterdam; E.J. Brill, Leiden. 

Stafleu, F.A. & R.S. Cowan. 1976-1988. 

Taxonomic Literature (second edition). 

Bohn, Scheltema, & Holkema, Utrecht.


CHLOROPHYTA - CHARACEAE 

contributed by M. Guerlesquin (1992); reviewed 

by M. Guerlesquin (1996) 

CHARACEAE REFERENCES: 

Guerlesquin, M. 1981. Contribution à la 

connaissance des Characées d'Amérique due 

Sud (Bolivie, Equateur, Guyane française). 

Rev. Hydrobiol. trop. 14(4): 381-406. 

Guerlesquin, M. 1983. Contribution à l'étude des 

Petites Antilles. Rev. Hydrobiol. trop. 16(3): 

213-233. 

Guerlesquin, M. 1985. Bilan et répartition de la 

flore des Charophycées dans les Antilles. 

Verh. Internat. Verein. Limnol. 22(5): 2879- 

2884. 

Guerlesquin, M., A. Couté, & A. Vaquer. 1991. 

Studies about the family of the Characeae 

from the Guyanas. Verh. Internat. Verein. 

Limnol. 24: 2673-2677. 

CHARACEAE 

Chara flaccida (A. Braun) Zaneveld [SU FG] 

Chara hydropitys Reich. [SU FG] 

Chara oerstediana A. Braun [FG] 

Chara robbinsii Halst. [FG] 

Nitella acuminata A. Braun ex Wallm. [GU] 

Nitella dictyosperma H. & J. Groves [FG] 

Nitella divaricata J. Groves & Steph. [FG] 

Nitella flagellifera J. Groves & G.O. Allen [FG] 

Nitella furcata Roxb. ex Bruz. [FG] 

Nitella glaziovii Zell. [FG] 

Nitella leptodactyla J. Groves [FG] 

Nitella megacarpa T.F. Allen [FG] 

Nitella microcarpa A. Braun [SU FG] 

Nitella mucronata A. Braun [FG] 

HEPATOPHYTA 

contributed by S.R. Gradstein (1992); additions 

and corrections by S.R. Gradstein (1996) 

HEPATOPHYTA REFERENCES: 

Cornelissen, J.H.C. & S.R. Gradstein. 1990. On 

the occurrence of bryophytes and 

macrolichens in different lowland rain forest 

types at Mabura Hill, Guyana. Tropical 

Bryology 3: 29-35. 

CHECKLIST 

Gradstein, S.R. 1975. A taxonomic monograph 

of the genus Acrolejeunea. Bryoph. Biblioth. 

4: 1-162. 

Gradstein, S.R. 1985. A revision of the genus 

Stictolejeunea (Spruce) Schiffner. Beih. Nova 

Hedwigia 80: 195-219. 

Gradstein, S.R. 1994. Lejeuneaceae: 

Ptychantheae, Brachiolejeuneae. Flora 

Neotropica Monogr. 62: 1-216. 

Gradstein, S.R. 1995. A Guide to the 

Bryophytes of Tropical America. 1. 

Liverworts and Hornworts. Draft version, 

Brussels, 255 pp. 

Gradstein, S.R. & G.M.C. Buskes. 1985. A 

revision of Neotropical Archilejeunea 

(Spruce) Schiffn. Beih. Nova Hedwigia 80: 

89-112. 

Gradstein, S.R. & J. Florschütz-de Waard. 1989. 

Results of a botanical expedition to Mount 

Roraima, Guyana. I. Bryophytes. Tropical 


Gradstein, S.R. & J. van Beek. 1985. A revision 

of the genus Symbiezidium Trevis. Beih. 

Nova Hedwigia 80: 2212-248. 

Gradstein, S.R. & W.H.A. Hekking. 1989. A 

catalogue of the bryophytes of the Guianas. I. 

Hepaticae and Anthocerotae. Journ. Hattori 

Bot. Lab. 66: 197-230. 

Grolle, R. 1989. Adelanthus am Mt. Roraima. J. 

Hattori Bot.Lab. 67: 243-247. 

Grolle, R. 1989. Marsupidium in Guayana. J. 

Hattori Bot. Lab. 66: 337-342. 

Grolle, R. & S.R. Gradstein. 1988. Haesselia, a 

new genus of Cephaloziaceae (Hepaticae) 

from Mt. Roraima, Guyana. Journ. Hattori 

Bot. Lab. 64: 327-334. 

Kuwahara, Y. 1986. The Metzgeriaceae of the 

Neotropics. Bryophytorum Bibliotheca 28: 1- 

254. 

Tixier, P. 1991. Bryophyta exotica - 9. 

Quelques Lejeuneaceae (Hépatiques) 

nouvelles pour l'Amerique du Sud. Candollea 

46(2): 267-302. 

ACROBOLBACEAE 

Marsupidium gradsteinii Grolle [GU FG] 

Tylimanthus laxus (Lindenb.) Spruce [GU] 

Tylimanthus striolatus Steph. [GU] 

Tylimanthus subtilis Steph. [GU] 

ADELANTHACEAE 

Adelanthus squarrosus Grolle [GU] 

CHLOROPHYTA, HEPATOPHYTA 

9

10 

HEPATOPHYTA 


ANEURACEAE 

Aneura pinguis (L.) Dumb. [FG] 

Aneura sp. [GU] 

Riccardia amazonica (Spruce) Schiffn. ex Gradst. & Hekking 

[GU SU FG] 

Riccardia andina (Spruce) Herz. [GU] 

Riccardia cervicornis (Spruce) Herz. ex Gradst. & Hekking 

[GU] 

Riccardia digitiloba (Spruce) Pagan [SU] 

Riccardia fendleri (Steph.) Pagan [FG] 

Riccardia fucoidea (Swartz) Mass. [GU] 

Riccardia plumaeformis (Spruce) Meenks [GU] 

Riccardia schwaneckei (Steph.) Pagan [GU] 

Riccardia sprucei (Steph.) Meenks & de Jong [FG] 

AYTONIACEAE 

Asterella venosa (Lehm. & Lindenb.) Evans [SU] 

BALANTIOPSACEAE 

Isotachis multiceps Gott. [GU] 

Neesioscyphus argillaceus (Nees) Grolle [GU] 

CALYPOGEIACEAE 

Calypogeia caespitosa (Spruce) Steph. [GU] 

Calypogeia cellulosa (Spreng.) Steph. [GU] 

Calypogeia crenulata Bischler [GU] 

Calypogeia laxa Gott. & Lindenb. [GU SU] 

Calypogeia lechleri (Steph.) Steph. [GU SU FG] 

Calypogeia lechleri (Steph.) Steph. var. densifolia (Steph.) 

Bischler [SU] 

Calypogeia miquelii Mont. [GU SU FG] 

Calypogeia peruviana Nees & Mont. [GU] 

Calypogeia venezuelana Fulford [GU] 

CEPHALOZIACEAE 

Alobiella husnotii (Gott.) Schiffn. [GU] 

Anomoclada portoricensis (Hampe & Gott.) Vána [GU SU] 

Cephalozia crassifolia (Lindenb. & Gott.) Fulford [GU FG] 

Haesselia acuminata Gradst. [GU] 

Haesselia roraimensis Grolle & Gradst. [GU] 

Nowellia evansii Grolle [GU] 

Odontoschisma atropurpureum Steph. [GU] 

Odontoschisma brasiliense Steph. [SU] 

Odontoschisma denudatum (Nees) Dum. [GU FG] 

Odontoschisma falcifolium Steph. [GU SU FG] 

Trabacellula tumidula Fulford [GU] 

CEPHALOZIACEAE - SYNONYMS AND 

MISIDENTIFICATIONS: 

Anomoclada mucosa Spruce =Anomoclada portoricensis 

CEPHALOZIELLACEAE 

Cylindrocolea planifolia (Steph.) Schust. [GU FG] 

FOSSOMBRONIACEAE 

Fossombronia sp. [GU] 

FRULLANIACEAE 

Frullania apiculata (Reinw. et al.) Dum. var. guianensis 

Lindenb. & Gott. [GU SU] 

Frullania apiculata (Reinw. et al.) Dum. [GU SU FG] 

Frullania arecae (Spreng.) Gott. [FG] 

Frullania bicornistipula Spruce [GU] 

Frullania brasiliensis Raddi [GU FG] 

Frullania breuteliana Gott. [FG] 

Frullania caulisequa (Nees) Nees [GU SU FG] 

Frullania degelii S. Arn. [FG] 

Frullania ericoides (Nees) Mont. [SU FG] 

Frullania exilis Tayl. [GU FG] 

Frullania gaudichaudii (Nees & Mont.) Nees & Mont. [GU] 

Frullania gibbosa Nees [GU SU FG] 

Frullania kunzei (Lehm. & Lindenb.) Lehm. & Lindenb. [GU 

FG] 

Frullania montagnei Gott. [FG] 

Frullania mucronata (Lehm. & Lindenb.) Lehm. & Lindenb. 

[GU SU] 

Frullania neesii Lindenb. [GU SU FG] 

Frullania nodulosa (Reinw. et al.) Nees [GU SU FG] 

Frullania parasitica Hampe ex Lehm. & Lindenb. [FG] 

Frullania pendulostyla Steph. [GU] 

Frullania riojaneirensis (Raddi) Ångstr. [GU SU FG] 

Frullania serrata Gott. [FG?] 

Frullania subtilissima Lindenb. [FG] 

FRULLANIACEAE - SYNONYMS AND 


Frullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. 

[specimens identified as F. caulisequa] 

Frullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. 

var. armata Lindenb. & Gott. [specimens identified as F. 

caulisequa] 

GEOCALYCACEAE 

Heteroscyphus contortuplicatus (Mont. & Nees) Grolle [SU] 

Leptoscyphus cuneifolius (Hook.) Mitt. [GU]


Leptoscyphus gibbosus (Tayl.) Mitt. [GU] 

Leptoscyphus ovatus (Spruce) Grolle [GU] 

Leptoscyphus porphyrius (Nees) Grolle [GU SU FG] 

Lophocolea bidentata (L.) Dum. [SU FG] 

Lophocolea bidentula (Nees) Fulford [FG] 

Lophocolea connata (Swartz) Nees [FG] 

Lophocolea liebmanniana Gott. [GU SU FG] 

Lophocolea martiana Nees [GU SU FG] 

Lophocolea perissodonta (Spruce) Steph. [GU SU] 

Lophocolea trapezoidea Mont. [GU FG] 

GEOCALYCACEAE - SYNONYMS AND 


Lophocolea coadunata (Swartz) Nees =Lophocolea bidentata 

HERBERTACEAE 

Herbertus divergens (Steph.) Herz. [GU SU FG] 

Herbertus juniperoideus (Swartz) Grolle [GU FG] 

Herbertus pensilis (Tayl.) Spruce [GU] 

JUNGERMANNIACEAE 

Jamesoniella rubricaulis (Nees) Grolle [GU] 

Stenorrhipis grollei Gradst. [GU] 

Syzygiella perfoliata (Swartz) Bruce [GU] 

Vanaea plagiochiloides (Inoue & Gradst.) Inoue & Gradst. 

[GU] [Mt. Roraima] 

JUNGERMANNIACEAE - SYNONYMS AND 


Anastrophyllum plagiochiloides Inoue & Gradst. =Vanaea 

plagiochiloides 

LEJEUNEACEAE 

Acanthocoleus aberrans (Lindenb. & Gott.) Kruijt var. 

aberrans [FG] 

Acrolejeunea emergens (Mitt.) Steph. [GU SU FG] 

Acrolejeunea torulosa (Lehm. & Lindenb.) Schiffn. [GU SU 

FG] 

Anoplolejeunea conferta (Meissn.) Evans [GU FG] 

Aphanolejeunea diaphana (Evans) Schust. [GU FG] 

Aphanolejeunea exigua Evans [FG] 

Aphanolejeunea gracilis Jovet-Ast [FG] 

Aphanolejeunea sicaefolia (Gott. ex Steph.) Evans [SU] 

Archilejeunea auberiana (Mont.) Evans [SU FG] 

Archilejeunea badia (Spruce) Steph. [GU] 

Archilejeunea crispistipula (Spruce) Steph. [GU] 

Archilejeunea fuscescens (Hampe ex Lehm.) Fulford [GU SU 

FG] 

Archilejeunea parviflora (Nees) Schiffn. [GU SU FG] 

Brachiolejeunia conduplicata (Steph.) Gradst. [SU FG] 

Bryopteris diffusa (Swartz) Nees [GU SU? FG] 

Bryopteris filicina (Swartz) Nees s.l. [GU SU FG] 

Caudalejeunea lehmanniana (Gott.) Evans [GU SU] 

Ceratolejeunea amazonica (Steph.) Steph. [SU] 

Ceratolejeunea anomala (Lindenb. & Gott.) Steph. [SU] 

Ceratolejeunea brasiliensis (Gott.) Schiffn. [FG] 

Ceratolejeunea caducifolia (Spruce) Steph. [FG] 

Ceratolejeunea confusa Schust. [GU] 

Ceratolejeunea cornuta (Lindenb.) Schiffn. [GU SU FG] 

Ceratolejeunea cubensis (Mont.) Schiffn. [SU FG] 

Ceratolejeunea dentatocornuta Steph. [FG] 

Ceratolejeunea desciscens (Sande Lac.) Schiffn. [GU SU FG] 

Ceratolejeunea flagelliformis (Steph.) Fulford [FG] 

Ceratolejeunea grandibracteata Fulford [FG] 

Ceratolejeunea guianensis (Nees & Mont.) Steph. [SU FG] 

Ceratolejeunea involvens (Nees & Mont.) Steph. [GU SU FG] 

Ceratolejeunea kegelii (Lindenb. & Gott.) Steph. [SU] 

Ceratolejeunea laetefusca (Aust.) Schust. [GU FG] 

Ceratolejeunea longicornis (Gott.) Steph. [FG] 

Ceratolejeunea maritima (Spruce) Steph. [GU SU FG] 

Ceratolejeunea megalophysa (Spruce) Steph. [GU SU FG] 

Ceratolejeunea parisii Steph. [FG] 

Ceratolejeunea patentissima (Hampe & Gott.) Evans [GU SU 

FG] 

Ceratolejeunea patulistipa Steph. [SU FG] 

Ceratolejeunea plumula (Spruce) Steph. [GU SU FG] 

Ceratolejeunea poeppigiana (Nees) Steph. [FG] 

Ceratolejeunea remotistipula Steph. [SU] 

Ceratolejeunea rubiginosa Steph. [GU] 

Ceratolejeunea rufo-pellucida (Spruce) Steph. [FG] 

Ceratolejeunea spinosa (Gott.) Steph. [GU SU FG] 

Ceratolejeunea splitgerberiana (Mont.) Steph. [SU FG] 

Ceratolejeunea valida Evans [FG] 

Ceratolejeunea variabilis (Lindenb.) Schiffn. [FG] 

Cheilolejeunea acutangula (Nees) Grolle [FG] 

Cheilolejeunea adnata (Kunze) Grolle [GU SU FG] 

Cheilolejeunea clausa (Nees & Mont.) Schust. [SU FG] 

Cheilolejeunea discoidea (Lehm. & Lindenb.) Kachroo & 

Schust. [FG] 

Cheilolejeunea fragrantissima (Spruce) Schust. [GU] [Mt. 

Roraima] 

Cheilolejeunea inflexa (Hampe ex Lehm. & Lindenb.) Grolle 

[SU FG] 

Cheilolejeunea longiflora (Tayl.) Schust. [FG] 

Cheilolejeunea rigidula (Nees & Mont.) Schust. [GU SU FG] 

Cheilolejeunea trifaria (Reinw. et al.) Mizut. [GU SU FG] 

Cololejeunea bekkerae Tixier [SU] 

Cololejeunea cardiocarpa (Mont.) Evans [FG] 

Cololejeunea cremersii Tixier [FG] 

Cololejeunea ensifolia (Spruce) Bonner [FG] 

Cololejeunea kegelii Steph. [SU] 

Cololejeunea obliqua (Nees & Mont.) Schiffn. [SU FG] 

Cololejeunea platyneura (Spruce) Evans [SU FG] 

Cololejeunea sigmoidea Jovet-Ast & Tixier [FG] 

Cololejeunea subcardiocarpa Tixier [FG] 

Cololejeunea surinamensis Tixier [SU FG] 

Colura clavigera Gott. ex Jovet-Ast. [GU SU FG] 

Colura cylindrica Herz. [GU SU FG] 

Colura greig-smithii Jovet-Ast. [SU FG] 

Colura lyrata Steph. [GU] 

Colura rhynchophora Jovet-Ast. [GU SU] 

Colura tenuicornis (Evans) Steph. [FG] 

Colura tortifolia (Nees & Mont.) Trevis. [GU SU FG] 

Colura valida Evans [FG] 

HEPATOPHYTA 

11

12 

HEPATOPHYTA 


Crossotolejeunea boryana (Mont.) Schiffn. [GU SU FG] 

Crossotolejeunea crenata (Nees & Mont.) Lacouture [GU SU 

FG] 

Crossotolejeunea paucispina (Spruce) Steph. [GU SU] 

Cyclolejeunea accedens (Gott.) Evans [FG] 

Cyclolejeunea chitonia (Tayl.) Evans [GU SU FG] 

Cyclolejeunea convexistipa (Lehm. & Lindenb.) Evans [GU 

SU FG] 

Cyclolejeunea dentistipula Steph. [FG] 

Cyclolejeunea foliorum (Nees) Grolle [FG] 

Cyclolejeunea luteola (Nees) Grolle [GU SU FG] 

Cyclolejeunea peruviana (Lehm. & Lindenb.) Evans [GU SU 

FG] 

Cyrtolejeunea holostipa (Spruce) Evans [GU SU FG] 

Cystolejeunea lineata (Lehm. & Lindenb.) Evans [GU FG] 

Dicranolejeunea axillaris (Nees & Mont.) Schiffn. [GU] 

Diplasiolejeunea brunnea Steph. [FG] 

Diplasiolejeunea cavifolia Steph. [GU SU FG] 

Diplasiolejeunea cobrensis Gott. ex Steph. [GU] 

Diplasiolejeunea galloana Jovet-Ast. [SU] 

Diplasiolejeunea lacostei Tixier [GU] 

Diplasiolejeunea latipuensis Tixier [GU] 

Diplasiolejeunea pellucida (Meissn. ex Spreng.) Schiffn. [GU 

SU FG] 

Diplasiolejeunea reyesiana Tixier [GU] 

Diplasiolejeunea rudolphiana Steph. [GU SU FG] 

Diplasiolejeunea unidentata (Lehm. & Lindenb.) Schiffn. [FG] 

Diplasiolejeunea unidentata (Lehm. & Lindenb.) Schiffn. var. 

minor Lindenb. & Gott. [SU] 

Diplasiolejeunea unidentata (Lehm. & Lindenb.) Schiffn. var. 

bidentula Lindenb. & Gott. [SU] 

Drepanolejeunea araucariae Steph. [FG] 

Drepanolejeunea bidens Steph. [FG] 

Drepanolejeunea bispinulosa Evans [SU] 

Drepanolejeunea crucianella (Tayl.) Evans [GU SU FG] 

Drepanolejeunea fragilis Bischler [GU SU FG] 

Drepanolejeunea inchoata (Meissn.) Steph. [SU FG] 

Drepanolejeunea integribracteata Bischler [FG] 

Drepanolejeunea lichenicola (Spruce) Steph. [GU FG] 

Drepanolejeunea mosenii (Steph.) Bischler [SU FG] 

Drepanolejeunea orthophylla (Nees & Mont.) Bischler [SU 

FG] 

Drepanolejeunea palmifolia (Nees) Steph. [GU] 

Drepanolejeunea pinnatiloba Schiffn. [GU FG] 

Echinocolea asperrima (Spruce) Schust. [SU FG] 

Echinocolea sp. [GU] [Mt. Roraima] 

Frullanoides corticalis (Lehm. & Lindenb.) van Slag. [GU SU 

FG] 

Frullanoides liebmanniana (Lindenb. & Gott.) van Slag. [GU 

SU? FG?] 

Frullanoides tristis (Steph.) van Slag. [GU? SU FG] 

Haplolejeunea cucullata (Steph.) Grolle [GU SU] 

Harpalejeunea heterodonta Evans [GU FG] 

Harpalejeunea oxyphylla (Nees & Mont.) Steph. [SU FG] 

Harpalejeunea stricta (Lindenb. & Gott.) Steph. [GU SU FG] 

Harpalejeunea uncinata Steph. [GU FG] 

Lejeunea angusta (Lehm. & Lindenb.) Mont. [FG] 

Lejeunea caespitosa Lindenb. [GU FG] 

Lejeunea cancellata Nees & Mont. [SU] 

Lejeunea cerina (Lehm. & Lindenb.) Gott. et al. [GU] 

Lejeunea cladogyna Evans [FG] 

Lejeunea dimorphophylla Schust. [GU SU] 

Lejeunea elegans Gott. [GU] 

Lejeunea exilis (Reinw. et al.) Grolle [FG] 

Lejeunea flava (Swartz) Nees [GU SU FG] 

Lejeunea flava (Swartz) Nees var. pellucida Lindenb. & Gott. 

[SU] 

Lejeunea glaucescens Gott. [GU SU FG] 

Lejeunea laeta (Lehm. & Lindenb.) Gott. et al. [SU] 

Lejeunea laetevirens Nees & Mont. [GU SU FG] 

Lejeunea reflexistipula (Lehm. & Lindenb.) Gott. et al. [GU 

FG] 

Lejeunea remotifolia Lindenb. & Steph. [FG] 

Lejeunea trachygona Spruce [GU] [Mt. Roraima] 

Lejeunea trinitensis Lindenb. [SU FG] 

Lejeunea weigeltii Lindenb. [SU] 

Lepidolejeunea involuta (Gott.) Grolle [GU SU FG] 

Lepidolejeunea ornata (Robins.) Schust. [GU SU] 

Leptolejeunea elliptica (Lehm. & Lindenb.) Schiffn. [GU SU 

FG] 

Leptolejeunea exocellata (Spruce) Evans [SU FG] 

Leptolejeunea foraminulosa Steph. [SU] 

Leptolejeunea mexicana Bischler [SU FG] 

Leptolejeunea moniliata Steph. [SU FG] 

Leptolejeunea obfuscata (Spruce) Steph. [SU FG] 

Leptolejeunea radicosa (Nees & Mont.) Grolle [FG] 

Leucolejeunea unciloba (Lindenb.) Evans [FG] 

Leucolejeunea xanthocarpa (Lehm. & Lindenb.) Evans [GU 

SU FG] 

Lopholejeunea eulopha (Tayl.) Schiffn. [GU SU?] 

Lopholejeunea nigricans (Lindenb.) Schiffn. [GU SU FG] 

Lopholejeunea quelchii Steph. [GU SU FG?] 

Lopholejeunea subfusca (Nees) Schiffn. [GU SU FG] 

Marchesinia brachiata (Swartz) Schiffn. [SU FG] 

Marchesinia robusta (Mitt.) Schiffn. [FG?] 

Mastigolejeunea auriculata (Wils.) Schiffn. [GU SU FG] 

Mastigolejeunea innovans (Spruce) Steph. [FG] 

Mastigolejeunea plicatiflora (Spruce) Schiffn. [SU FG] 

Microlejeunea acutifolia Steph. [GU SU FG] 

Microlejeunea aphanella (Spruce) Steph. [FG] 

Microlejeunea aspera Steph. [FG] 

Microlejeunea bullata (Tayl.) Evans [SU FG] 

Microlejeunea crenulifolia (Gott.) Steph. [SU FG] 

Microlejeunea cystifera Herz. [FG] 

Microlejeunea epiphylla Bischler [FG] 

Microlejeunea monoica Bischler [FG] 

Microlejeunea valenciana Steph. [FG] 

Neurolejeunea breutelii (Gott.) Evans [FG] 

Neurolejeunea seminervis (Spruce) Schiffn. [GU SU FG] 

Odontolejeunea decemdentata (Spruce) Steph. [FG] 

Odontolejeunea lunulata (Weber) Schiffn. [GU SU FG] 

Odontolejeunea rhomalea (Spruce) Steph. [GU SU FG] 

Omphalanthus filiformis (Swartz) Nees [GU] 

Omphalanthus paramicola (Herz.) Gradst. [GU] 

Pictolejeunea picta (Gott. ex Steph.) Grolle [GU SU FG] 

Prionolejeunea aemula (Gott.) Evans [FG] 

Prionolejeunea denticulata (F. Web.) Grolle [FG] 

Prionolejeunea elegans Spruce [GU] 

Prionolejeunea eroso-dentata Steph. [SU] 

Prionolejeunea galliotii Steph. [FG] 

Prionolejeunea gaudalupensis (Lindenb.) Steph. [FG] 

Prionolejeunea innovata Evans [GU] 

Prionolejeunea macrocardia (Spruce) Steph. [FG] 

Prionolejeunea microdonta (Gott.) Steph. [FG] 

Prionolejeunea obscura Spruce ex Steph. [FG] 

Prionolejeunea subobscura Spruce ex Steph. [FG] 

Prionolejeunea trachyodes (Spruce) Steph. [GU FG] 

Pycnolejeunea callosa (Lindenb.) Steph. [GU] 

Pycnolejeunea contigua (Nees) Grolle [GU SU FG] 

Pycnolejeunea decurviloba Steph. [GU SU]


Pycnolejeunea macroloba (Nees & Mont.) Schiffn. [GU SU 

FG] 

Pycnolejeunea surinamensis Steph. [SU] 

Rectolejeunea berteroana (Gott.) Evans [GU SU FG] 

Rectolejeunea brittoniae Evans [GU SU FG] 

Rectolejeunea emarginuliflora (Gott. ex Schiffn.) Evans [FG] 

Rectolejeunea flagelliformis Evans [GU SU FG] 

Rectolejeunea kegelii (Steph.) Steph. [SU] 

Rectolejeunea mandioccana (Steph.) Steph. [SU] 

Rectolejeunea phyllobola (Nees & Mont.) Evans [SU FG] 

Rhaphidolejeunea polyrhiza (Nees) Bischler [FG] 

Schiffneriolejeunea amazonica Gradst. [SU FG] 

Schiffneriolejeunea polycarpa (Nees) Gradst. [GU SU FG?] 

Schusterolejeunea inundata (Spruce) Grolle [SU FG] 

Stictolejeunea balfourii (Mitt.) E.W. Jones var. bekkeri Gradst. 

[SU FG] 

Stictolejeunea squamata (Willd. ex Weber) Schiffn. [GU SU 

FG] 

Symbiezidium barbiflorum (Lindenb. & Gott.) Evans [GU SU 

FG] 

Symbiezidium transversale (Swartz) Trevis. var. transversale 

[GU? SU FG] 

Symbiezidium transversale (Swartz) Trevis. var. hookerianum 

(Gott. et al.) Gradst. & van Beek [GU SU FG] 

Taxilejeunea asthenica (Spruce) Steph. [FG] 

Taxilejeunea debilis (Lehm. & Lindenb.) Steph. [FG] 

Taxilejeunea fissistipula Steph. [FG] 

Taxilejeunea sulphurea (Lehm. & Lindenb.) Schiffn. [FG] 

Taxilejeunea surinamensis (Lindenb. & Gott.) Steph. [SU FG] 

Taxilejeunea uleana Steph. [SU] 

Thysananthus amazonicus (Spruce) Schiffn. [GU SU FG] 

Trachylejeunea pandurantha (Spruce) Steph. [GU SU FG] 

Verdoornianthus griffinii Gradst. [FG] 

LEJEUNEACEAE - SYNONYMS AND 


Acrolejeunea torulosa (Lehm. & Lindenb.) Schiffn. var. obtusa 

Gradst. =Acrolejeunea torulosa 

Archilejeunea fuscescens (Hampe) Fulf. var. paroica Gradstein 

=Archilejeunea badia 

Aureolejeunea paramicola (Herz.) Schust. =Omphalanthus 

paramicola 

Brachiolejeunea phyllorhiza (Nees) Kruijt & Gradst. 

[specimens identified as B. conduplicata] 

Bryopteris fruticulosa Tayl. =Bryopteris filicina s.l. 

Bryopteris trinitensis (Lehm. & Lindenb.) Lehm. & Lindenb. 

=Bryopteris filicina s.l. 

Cyclolejeunea angulistipa (Steph.) Evans =Prionolejeunea 

denticulata 

Frullanoides densifolia Raddi [not in Guianas] 

Lopholejeunea muelleriana (Gott.) Schiffn. =Lopholejeunea 

nigricans 

Lopholejeunea subfusca (Nees) Schiffn. var. minor Gott. et al. 

=Lopholejeunea subfusca? 

Thysananthus comosus Lindenb. [not in Guianas] 

LEPIDOZIACEAE 

Arachniopsis diacantha (Mont.) Howe [GU SU FG] 

Arachniopsis pecten Spruce [GU FG] 

Bazzania acuminata (Lindenb. & Gott.) Trevis. [SU] 

Bazzania bidens (Nees) Trevis. [GU FG] 

Bazzania breuteliana (Lindenb. & Gott.) Trevis. [GU SU FG] 

Bazzania cubensis (Gott.) Pagán [GU] 

Bazzania cuneistipula (Gott. & Lindenb.) Trevis. [GU] 

Bazzania glaziovii (Gott.) Fulford [GU] 

Bazzania gracilis (Hampe & Gott.) Steph. [GU] 

Bazzania hookeri (Lindenb.) Trevis. [GU FG] 

Bazzania longa Trevis. [SU] 

Bazzania longistipula (Lindenb.) Trevis. [GU] 

Bazzania robusta Spruce [GU] 

Bazzania roraimensis (Steph.) Fulford [GU SU] 

Bazzania stolonifera (Swartz) Trevis. [GU FG] 

Bazzania teretiuscula (Lindenb. & Gott.) Trevis. [SU] 

Kurzia capillaris (Swartz) Grolle [GU] 

Lepidozia coilophylla Tayl. [GU SU] 

Lepidozia macrocolea Spruce [GU] 

Micropterygium campanense Spruce ex Reim. [GU] 

Micropterygium carinatum (Grev.) Reim. [GU] [Mt. Roraima] 

Micropterygium grandistipulum Steph. [GU] 

Micropterygium pterygophyllum (Nees) Trevis [GU] 

Micropterygium tenax (Steph.) Grolle [GU] 

Micropterygium trachyphyllum Reim. [GU SU FG] 

Micropterygium tumidulum Fulford [GU] 

Mytilopsis albifrons Spruce [GU] 

Paracromastigum bifidum (Steph.) Grolle [GU] 

Pteropsiella serrulata Spruce ex Steph. [GU SU] 

Telaranea nematodes (Gott. ex Aust.) Howe [GU SU FG] 

Zoopsidella monodactyla (Spruce) Schust. [GU] 

Zoopsidella serra (Spruce) Schust. [SU] 

Zoopsis antillana Steph. [GU] 

Zoopsis cynosuranda Spruce ex Steph. [GU SU] 

Zoopsis integrifolia (Spruce) Steph. [GU] 

Zoopsis macella (Spruce) Fulford [GU SU] 

MARCHANTIACEAE 

Dumortiera hirsuta (Swartz) Nees [FG] 

Marchantia chenopoda L. [SU FG] 

METZGERIACEAE 

Metzgeria adscendens Steph. ex Goebel [GU] 

Metzgeria australis Steph. [FG] 

Metzgeria convoluta Steph. [FG] 

Metzgeria decipiens (Mass.) Schiffn. [GU] 

Metzgeria effusa Steph. [GU] 

Metzgeria goebeliana Steph. [GU] 

Metzgeria innovans Steph. [GU] 

Metzgeria leptoneura Spruce [SU FG] 

Metzgeria procera Mitt. [GU] 

METZGERIACEAE - SYNONYMS AND 


Metzgeria hamata Lindenb. =Metzgeria leptoneura 

Metzgeria radicans Steph. =Metzgeria innovans 

Metzgeria thomeensis Steph. [specimens identified as M. 

australis] 

HEPATOPHYTA 

13

14 

HEPATOPHYTA 


MONOCLEACEAE 

Monoclea gottschei Lindb. subsp. elongata Gradst. & Mues 

[GU FG] 

MONOCLEACEAE - SYNONYMS AND 


Monoclea forsteri Hook. [specimens identified as M. 

gottschei subsp. elongata] 

PALLAVICINIACEAE 

Pallavicinia lyellii (Hook.) Carruth. [GU] 

Symphyogyna aspera Steph. [GU] 

Symphyogyna brasiliensis Nees [GU] 

Symphyogyna brongniartii Mont. [FG] 

Symphyogyna marginata Steph. [GU] 

PALLAVICINIACEAE - SYNONYMS AND 


Symphyogyna sinuata (Swartz) Nees & Mont. [not in 

Guianas] 

PLAGIOCHILACEAE 

Plagiochila abrupta Lehm. & Lindenb. [GU FG] 

Plagiochila aerea Tayl. [GU] 

Plagiochila amazonica Spruce [GU SU FG] 

Plagiochila arcuata Lindenb. [GU] 

Plagiochila bidens Gott. [GU] 

Plagiochila bursata (Desv.) Lindenb. [GU FG] 

Plagiochila contigua Gott. [GU FG] 

Plagiochila cristata Lindenb. [GU] 

Plagiochila depressa Spruce [GU SU] 

Plagiochila dichotoma (Nees) Grolle [SU] 

Plagiochila disticha Lehm. & Lindenb. [GU SU FG] 

Plagiochila divaricata Lindenb. [GU SU FG] 

Plagiochila dussiana Steph. [GU] 

Plagiochila echinella Gott. [GU] 

Plagiochila esmeraldana Steph. [GU] 

Plagiochila falcata Steph. [FG] 

Plagiochila guatemalensis Steph. [FG] 

Plagiochila guilleminiana Mont. [GU] 

Plagiochila gymnocalyx Inoue [GU] 

Plagiochila hansmeyeri Steph. [GU] 

Plagiochila irmscheri Steph. [GU] 

Plagiochila jamaicensis Lindenb. & Hampe [SU FG] 

Plagiochila kegeliana Steph. [GU SU FG] 

Plagiochila laetevirens Lindenb. [GU SU FG] 

Plagiochila lansbergeri Gott. [GU] 

Plagiochila macrifolia Tayl. [GU SU] 

Plagiochila micropterys Gott. [GU FG] 

Plagiochila montagnei Nees [FG] 

Plagiochila orbigniana Nees & Mont. [GU SU] 

Plagiochila patentissima Lindenb. [GU SU] 

Plagiochila patula (Swartz) Nees & Mont. [FG] 

Plagiochila pearceana Steph. [GU] 

Plagiochila perrotettiana Mont. [FG] 

Plagiochila pittieri Steph. [GU FG] 

Plagiochila quelchii Steph. [GU] 

Plagiochila rubescens (Lehm. & Lindenb.) Lindenb. [GU] 

Plagiochila rutilans Lindenb. [GU SU FG] 

Plagiochila semi-amplexicaulis Steph. [SU] 

Plagiochila serrata (Roth) Lindenb. [GU SU FG] 

Plagiochila subplana Lindenb. [GU] 

Plagiochila subsimplex (Swartz) Dum. [GU] 

Plagiochila surinamensis Molk. [SU] 

Plagiochila tarapotensis Steph. [FG] 

Plagiochila tenuis Lindenb. [GU FG] 

Plagiochila vincentina Lindenb. [GU] 

PLAGIOCHILACEAE - SYNONYMS AND 


Plagiochila hyalocoetis Spruce [specimens identified as P. 

vincentina] 

Plagiochila hypnoides Lindenb. =Plagiochila disticha 

PLEUROZIACEAE 

Pleurozia heterophylla Steph. ex Fulford [GU] 

RADULACEAE 

Radula decora Gott. [FG] 

Radula fendleri Gott. ex Steph. [SU] 

Radula flaccida Lindenb. & Gott. [GU SU FG] 

Radula gottscheana Tayl. [SU FG] 

Radula gradsteinii Yamada [GU] 

Radula guyanensis Yamada [GU] 

Radula husnotii Castle [GU FG] 

Radula involvens Spruce [GU] 

Radula kegelii Gott. ex Steph. [SU FG] 

Radula laxiramae Steph. [FG] 

Radula macrostachya Lindenb. & Gott. [GU SU FG] 

Radula mazarunensis Yamada [GU] 

Radula portoricensis Steph. [FG] 

Radula recubans Tayl. [GU] 

Radula stenocalyx Mont. [FG] 

Radula taylori Steph. [GU] 

Radula tenera Mitt. ex Steph. [GU] 

RADULACEAE - SYNONYMS AND 


Radula boryana (Weber) Nees [specimens identified as R. 

gottscheana] 

Radula caldana Ångstrom =Radula macrostachya


Radula surinamensis Steph. =Radula macrostachya 

RICCIACEAE 

Riccia subplana Steph. [GU FG] 

SCAPANIACEAE 

Scapania portoricensis Hampe & Gott. [GU] 

TRICHOCOLEACEAE 

Trichocolea filicaulis Steph. [FG] 

Trichocolea flaccida (Spruce) Jack & Steph. [FG] 

Trichocolea sprucei Steph. [FG] 

Trichocolea tomentosa (Swartz) Gott. [GU FG] 

ANTHOCEROPHYTA 

contributed by S.R. Gradstein (1992); additions 

and corrections by S.R. Gradstein (1996) 

ANTOCEROPHYTA REFERENCES: 

Gradstein, S.R. 1995. A Guide to the 

Bryophytes of Tropical America. 1. 

Liverworts and Hornworts. Draft version, 

Brussels, 255 pp. 

Gradstein, S.R. & W.H.A. Hekking. 1989. A 

catalogue of the bryophytes of the Guianas. I. 

Hepaticae and Anthocerotae. Journ. Hattori 

Bot. Lab. 66: 197-230. 





DENDROCEROTACEAE 

Dendroceros breutelii Nees var. surinamensis Lindenb. & 

Gott. [SU] 

Dendroceros crispus (Swartz) Nees [GU SU FG] 

NOTOTHYLADACEAE 

Notothylas breutelii Gott. [FG] 

BRYOPHYTA 

contributed by J. Florschütz-de Waard (1992); 

additions and corrections by A.E. Newton, 

W.R. Buck, & J. Florschütz-de Waard (1996) 

BRYOPHYTA REFERENCES: 

Allen, B.H. 1987. A revision of the genus 

Leucomium. Mem. New York Bot. Gard. 45: 

661-677. 

Allen, B.H. 1990. A revision of the genus 

Crossomitrium (Musci: Hookeriaceae). 

Tropical Bryologist 2: 3-34. 

Buck, W.R. 1987. Taxonomic and 

nomenclatural rearrangement in the 

Hookeriales with notes on West Indian taxa. 

Brittonia 39(2): 210-224. 

Buck, W.R. 1990. Contributions to the moss 

flora of Guyana. Mem. New York Bot. Gard. 

64: 184-196. 

Buck, W.R. 1991. Notes on neotropical 

Pterobryaceae. Brittonia 43(2): 96-101. 

Buck, W.R. & H. Crum. 1990. An evaluation of 

familial limits among the genera traditionally 

aligned with the Thuidiaceae and Leskeaceae. 

Contr. Univ. Mich. Herb. 17: 55-69. 

Buck, W.R. & H. Crum. 1993. Notes on 

Guayana mosses with new information on 

Sphagnum ornatum. Brittonia 45(1): 17-20. 

Cornelissen, J.H.C. & S.R. Gradstein. 1990. On 

the occurrence of bryophytes and 

macrolichens in different lowland rain forest 

types at Mabura Hill, Guyana. Tropical 


Delgadillo M., C., B. Bello, & A. Cárdenas S. 

1995. LATMOSS, a catalogue of Neotropical 

mosses. Monogr. Syst. Bot. Missouri Bot. 

Gard. 56: 1-191. 

Florschütz, P.A. 1964. Musci. In J. Lanjouw, 

Flora of Suriname 6(1): 1-271. 

Florschütz-de Waard, J. 1986. Musci (part II). 

In A.L. Stoffers & J.C. Lindeman, Flora of 

Suriname 6(1): 273-361. 

Florschütz-de Waard, J. 1990. A catalogue of 

the bryophytes of the Guianas. II. Musci. 

Tropical Bryology 3: 89-104. 

Florschütz-de Waard, J. 1992. A revision of the 

genus Potamium (Musci: Sematophyllaceae). 

Tropical Bryology 5: 109-121. 

Florschütz-des Waard, J. (in press). Musci (part 

III). In A.R.A. Görts-van Rijn, Flora of the 

Guianas. 

Frahm, J.-P. 1991. Dicranaceae p.p. Flora 

Neotropica Monogr. 54. 

Frahm, J.-P. & S.R. Gradstein. 1987. The genera 

Bryohumbertia and Campylopus in the 

Guianas. Crypt. Bryol. Lichén. 8: 311-319. 





HEPATOPHYTA, ANTHOCEROPHYTA, BRYOPHYTA 

15

16 

BRYOPHYTA 


Ireland, R.R. 1991. A preliminary study of the 

moss genus Isopterygium (Hypnaceae) in 

Latin America. Caldasia 16: 265-276. 

Ireland, R.R. 1992. The moss genus 

Isopterygium (Hypnaceae) in Latin America. 

Tropical Bryologist 6: 111-132. 

Ireland, R.R. & W.R. Buck. 1994. 

Stereophyllaceae. Flora Neotropica Monogr. 

65: 1-49. 

Pursell, R.A. 1994. Taxonomic notes on 

Neotropical Fissidens. Bryologist 97(3): 253- 

271. 

Reese, W.D. 1993. Calymperaceae. Flora 

Neotropica Monogr. 58. 

Salazar Allen, N. 1993. Leucophanaceae. Flora 


ANDREAEACEAE 

Andreaea nitida J.D. Hook. & Wilson [FG?] 

ARCHIDIACEAE 

Archidium ohioense C. Müll. [SU FG] 

BARTRAMIACEAE 

Breutelia tomentosa (Brid.) Jaeg. & Sauerb. [GU] 

Leiomela bartramioides (Hook.) Par. [GU] 

Philonotis uncinata (Schwaegr.) Brid. var. glaucescens 

(Hornsch.) Florsch. [GU SU FG] [=Philonotis 

glaucescens] 

Philonotis uncinata (Schwaegr.) Brid. var. gracillima (Ångstr.) 

Florsch. [GU SU] [=Philonotis gracillima] 

Philonotis uncinata (Schwaegr.) Brid. var. uncinata [GU SU 

FG?] 

BARTRAMIACEAE - SYNONYMS AND 


Philonotis glaucescens (Hornsch.) Broth. =Philonotis uncinata 

var. glaucescens 

Philonotis gracillima Ångstr. =Philonotis uncinata var. 

gracillima 

BRACHYTHECIACEAE 

Lepyrodontopsis trichophylla (Hedw.) Broth. [GU FG] 

BRYACEAE 

Brachymenium sipapoense E.B. Bartr. [SU FG] 

Brachymenium speciosum (Hook.f. & Wils.) Steere [GU] 

Brachymenium wrightii (Sull.) Broth. [SU FG] 

Bryum apiculatum Schwaegr. [SU FG] 

Bryum argenteum Hedw. [GU SU] 

Bryum beyrichianum (Hornsch.) C. Müll. [GU SU FG] 

[=Rhodobryum beyrichianum] 

Bryum billardieri Schwaegr. [GU SU FG] 

Bryum capillare Hedw. [SU FG] 

Bryum coronatum Schwaegr. [GU SU FG] 

BRYACEAE - SYNONYMS AND 


Bryum ellipsifolium C. Müll. [not in Guianas] 

Rhodobryum beyrichianum (Hornsch.) Par. =Bryum 

beyrichianum 

CALYMPERACEAE 

Calymperes afzelii Swartz [GU SU FG] 

Calymperes erosum C. Müll. [GU SU FG] 

Calymperes guildingii Hook. & Grev. [GU SU FG] 

Calymperes levyanum Besch. [SU] 

Calymperes lonchophyllum Schwaegr. [GU SU FG] 

Calymperes mitrafugax Florsch. [SU FG] 

Calymperes nicaraguense Ren. & Card. [GU SU? FG] 

Calymperes othmeri Herz. [GU SU FG] 

Calymperes palisotii Schwaegr. [GU SU FG] 

Calymperes pallidum Mitt. [SU FG] [to be expected in GU] 

Calymperes platyloma Mitt. [GU SU FG] 

Calymperes rubiginosum (Mitt.) Reese [GU SU FG] 

Calymperes smithii E.B. Bartram [GU SU FG] 

Calymperes venezuelanum (Mitt.) Broth. ex Pitt. [GU SU FG] 

Syrrhopodon circinatus (Brid.) Mitt. [FG?] 

Syrrhopodon cryptocarpos Dozy & Molk. [GU SU FG] 

Syrrhopodon cymbifolius C. Müll. [GU FG] 

Syrrhopodon elatus Mont. [SU FG] 

Syrrhopodon elongatus Sull. var. glaziovii (Hampe) Reese 

[GU] 

Syrrhopodon flexifolius Mitt. [GU SU FG] 

Syrrhopodon gaudichaudii Mont. [GU SU FG] 

Syrrhopodon graminicola Williams [GU SU FG] 

Syrrhopodon helicophyllus Mitt. [GU] 

Syrrhopodon hornschuchii Mart. [GU SU FG] 

Syrrhopodon incompletus Schwaegr. var. incompletus [GU 

SU FG] 

Syrrhopodon incompletus Schwaegr. var. lanceolatus (Hampe) 

Reese [GU SU FG] 

Syrrhopodon incompletus Schwaegr. var. luridus (Par. & 

Broth.) Florsch. [GU SU FG] 

Syrrhopodon incompletus Schwaegr. var. berteroanus (Brid.) 

Reese [FG] 

Syrrhopodon leprieurii Mont. [GU SU FG] 

Syrrhopodon ligulatus Mont. [GU SU FG] 

Syrrhopodon lycopodioides (Brid.) C. Müll. [GU] 

Syrrhopodon parasiticus (Brid.) Besch. [GU SU FG]


Syrrhopodon prolifer Schwaegr. var. acanthoneuros (C. Müll.) 

C. Müll. [GU SU FG] 

Syrrhopodon prolifer Schwaegr. var. cincinnatus (Hampe) 

Reese [GU SU FG] 

Syrrhopodon prolifer Schwaegr. var. prolifer [SU FG] 

Syrrhopodon prolifer Schwaegr. var. scaber (Mitt.) Reese 

[GU? SU FG] 

Syrrhopodon rigidus Hook. & Grev. [GU SU FG] 

Syrrhopodon simmondsii Steere [GU SU FG] 

Syrrhopodon tortilis Hampe [SU?] 

CALYMPERACEAE - SYNONYMS AND 


Calymperes donnellii Aust. =Calymperes afzelii 

Calymperes maguirei E.B. Bartram =Calymperes 

venezuelanum 

Calymperes reyi Broth. =Calymperes afzelii 

Calymperes richardii C. Müll. =Calymperes palisotii 

Calymperes uleanum Broth. =Calymperes pallidum 

Calymperopsis disciformis (C. Müll.) Broth. =Syrrhopodon 

graminicola 

Calymperopsis parasitica (Brid.) Broth. =Syrrhopodon 

parasiticus 

Syrrhopodon parasiticus (Brid.) Besch. var. disciformis (C. 

Müll.) Florsch. =Syrrhopodon graminicola 

Syrrhopodon parasiticus (Brid.) Besch. var. flexifolius (Mitt.) 

Reese =Syrrhopodon flexifolius 

Syrrhopodon prolifer Schwaegr. var. papillosus (C. Müll.) 

Reese =Syrrhopodon prolifer var. scaber 

Syrrhopodon surinamensis Dozy & Molk. =Syrrhopodon 

hornschuchii 

DICRANACEAE 

Atractylocarpus longisetus (Hook.) Bartr. [GU?] [Mt. 

Roraima] 

Bryohumbertia filifolia (Hornsch.) Frahm var. filifolia [GU 

SU? FG?] 

Campylopus arctocarpus (Hornsch.) Mitt. var. arctocarpus 

[GU SU FG] 

Campylopus cubensis Sull. [GU SU FG] 

Campylopus cuspidatus (Hornsch.) Mitt. var. cuspidatus [Mt. 

Roraima] 

Campylopus lamellinervis (C. Müll.) Mitt. var. lamellinervis 

[GU] 

Campylopus lamellinervis (C. Müll.) Mitt. var. exaltatus (C. 

Müll.) Frahm [FG] 

Campylopus luteus (C. Müll.) Par. [GU SU FG] 

Campylopus pilifer Brid. subsp. pilifer [GU SU FG] 

Campylopus richardii Brid. [GU] 

Campylopus savannarum (C. Müll.) Mitt. [GU SU FG] 

Campylopus subcuspidatus (Hampe) Jaeg. [GU SU FG] 

Campylopus surinamensis C. Müll. [GU SU FG] 

Campylopus trachyblepharon (C. Müll.) Mitt. [GU FG] 

Dicranella hilariana (Mont.) Mitt. [GU SU FG] 

Dicranodontium denudatum (Brid.) Britt. [Mt. Roraima] 

Dicranodontium pulchroalare Broth. [Mt. Roraima] 

Eucamptodontopsis brittonae (E.B. Bartram) Allen [GU] 

Eucamptodontopsis pilifera (Mitt.) Broth. [GU SU] 

Eucamptodontopsis tortuosa H. Robinson [GU] 

Holomitrium arboreum Mitt. [GU SU FG] 

Leucoloma cruegerianum (C. Müll.) Jaeg. & Sauerb. [GU SU 

FG] 

Leucoloma mariei Besch. [FG] 

Leucoloma serrulatum Brid. [GU SU FG] 

Leucoloma tortellum (Mitt.) Jaeg. [GU SU FG] 

Microdus muralis (Hampe) Par. [FG] 

DICRANACEAE - SYNONYMS AND 


Campylopus filifolius (Hornsch.) Mitt. =Bryohumbertia 

filifolia 

Dicranoloma brittonae E.B. Bartram =Eucamptodontopsis 

brittonae 

Diphyscium ulei C. Müll. [GU] 

DIPHYSCIACEAE 

EPHEMERACEAE 

Micromitrium telephorothecum (Florsch.) Crosby [SU] 

FISSIDENTACEAE 

Fissidens allionii Broth. [GU? FG] 

Fissidens angustifolius Sull. [FG] 

Fissidens diplodus Mitt. var. richardsii (Williams) Pursell [GU 

SU FG] 

Fissidens diplodus Mitt. var. diplodus [FG] 

Fissidens diplodus Mitt. var. wainionis [GU] 

Fissidens donnellii Aust. [GU] 

Fissidens elegans Brid. [SU FG] 

Fissidens flexinervis Mitt. [SU] 

Fissidens geijskesii Florsch. [SU] 

Fissidens guianensis Mont. [GU SU FG] 

Fissidens hornschuchii Mont. [FG] 

Fissidens inaequalis Mitt. [GU SU FG] 

Fissidens intramarginatus (Hampe) Jaeg. [SU] 

Fissidens leptophyllus Mont. [GU SU FG] 

Fissidens minutus Thwaites & Mitt. var. minutus [SU FG] 

Fissidens mollis Mitt. [SU FG] 

Fissidens neglectus Crum [SU] 

Fissidens oblongifolius Hook.f. & Wils. [GU SU FG] 

Fissidens obtusissimus (Florsch.) Pursell [SU FG] 

Fissidens ornatus Herz. [FG] 

Fissidens papillosus Lac. [GU?] 

Fissidens pellucidus Hornsch. [FG] 

Fissidens pennula Broth. [SU] 

Fissidens prionodes Mont. [GU SU FG] 

Fissidens radicans Mont. [GU SU FG] 

Fissidens ramicola Broth. [SU] 

Fissidens reticulosus (C. Müll.) Mitt. [SU FG] 

Fissidens scariosus Mitt. [GU SU FG] 

Fissidens splitgerberianus Dozy & Molk. [SU?] 

Fissidens submarginatus Bruch. [GU SU FG] 

BRYOPHYTA 

17

18 

Fissidens subramicola Broth. [SU FG] 

Fissidens subulatus Mitt. [GU?] 

Fissidens weirii Mitt. var. weirii [SU FG] 

Fissidens zollingeri Mont. [SU FG] 

BRYOPHYTA 


FISSIDENTACEAE - SYNONYMS AND 


Fissidens asplenioides Hedw. [not in Guianas] 

Fissidens austro-americanus Pursell & Reese =Fissidens 

subramicola 

Fissidens flavinervis Mitt. =Fissidens elegans 

Fissidens intermedius C. Müll. =Fissidens submarginatus 

Fissidens kegelianus C. Müll. =Fissidens zollingeri 

Fissidens microcladus Thwaites & Mitt. =Fissidens minutus 

var. minutus 

Fissidens pauperculus Howe [not in Guianas] 

Fissidens pauperculus Howe var. surinamensis Florsch. 

=Fissidens inaequalis 

Fissidens ravenellii Sull. =Fissidens elegans 

Fissidens similiretis Sull. =Fissidens oblongifolius 

FUNARIACEAE 

Funaria bonplandii (Hook.) Broth. [GU] [=Entosthodon 

bonplandii] 

Funaria hygrometrica Hedw. var. calvescens (Schwaegr.) 

Mont. [GU SU] 

FUNARIACEAE - SYNONYMS AND 


Entosthodon bonplandii (Hook.) Mitt. =Funaria bonplandii 

HEDWIGIACEAE 

Rhacocarpus purpurascens (Brid.) Par. [GU SU?] 

HELICOPHYLLACEAE 

Helicophyllum torquatum (Hook.) Brid. [GU] 

HOOKERIACEAE 

Brymela parkeriana (Hook. & Grev.) Buck [GU SU FG?] 

[=Hookeriopsis parkeriana] 

Callicosta bipinnata (Schwaegr.) C. Müll. [GU SU FG] 

[=Pilotrichum bipinnatum] 

Callicosta evanescens C. Müll. [GU SU FG] [=Pilotrichum 

evanescens] 

Callicosta fendleri (C. Müll.) Crosby [GU SU FG] 

[=Pilotrichum fendleri] 

Callicostella grossiretis E.B. Bartram [FG] [=Callicostella 

colombica Williams?] 

Callicostella guatemalensis (E.B. Bartram) J. Florsch. [SU] 

Callicostella merkelii (Hornsch.) Jaeg. [SU FG] 

Callicostella microcarpa Ångstr. [GU?] 

Callicostella mosenii (Broth.) Broth. [GU?] 

Callicostella pallida (Hornsch.) Ångstr. [GU SU FG] 

Callicostella rufescens (Mitt.) Jaeg. [GU SU FG] 

Callicostella sp. aff. rivularis (Mitt.) Jaeg. [GU FG] 

Callicostella subdepressa (Besch.) Buck [GU?] 

Crossomitrium epiphyllum (Mitt.) C. Müll. [GU? FG] 

Crossomitrium patrisiae (Brid.) C. Müll. [GU SU FG] 

Crossomitrium sintenisii C. Müll. [GU SU FG] 

Cyclodictyon cuspidatum Kuntze [GU?] 

Cyclodictyon limbatum (Hampe) Kuntze [GU?] 

Cyclodictyon varians (Sull.) O. Kuntze [GU SU FG?] 

Daltonia longifolia Tayl. [GU] 

Hemiragis aurea (Brid.) Kindb. [GU] 

Hypnella diversifolia (Mitt.) Jaeg. [GU] 

Hypnella guayanensis Allen & Buck [GU] 

Hypnella pallescens (Hook.) Jaeg. [GU SU FG] 

Lepidopilidium divaricatum (Dozy & Molk.) Broth. [SU?] 

Lepidopilidium portoricense (C. Müll.) Crum & Steere [GU 

SU FG] 

Lepidopilum affine C. Müll. [SU FG] 

Lepidopilum biductulosum (P. Beauv.) Wijk & Marg. [GU?] 

Lepidopilum brevipes Mitt. [FG] 

Lepidopilum cubense (Sull.) Mitt. [SU FG] 

Lepidopilum integrifolium Broth. [FG] 

Lepidopilum intermedium (C. Müll.) Mitt. [GU?] 

Lepidopilum polytrichoides (Hedw.) Brid. [SU FG] 

Lepidopilum purpurascens Schimp. ex Besch. [GU] 

Lepidopilum radicale Mitt. [GU? SU FG] 

Lepidopilum scabrisetum (Schwaegr.) Steere [GU SU FG] 

Lepidopilum stolonaceum C. Müll. [GU SU FG] 

Lepidopilum surinamense C. Müll. [GU SU FG] 

Lepidopilum tortifolium Mitt. [FG] 

Leskeodon andicola (Mitt.) Broth. [GU] 

Leskeodon cubensis (Mitt.) Thér. [GU] 

Thamniopsis diffusa (Wils.) Buck [GU] [=Hookeriopsis 

diffusa] 

Thamniopsis killipii (Williams) E.B. Bartram [GU SU] 

Thamniopsis undata (Hedw.) Buck [GU] [=Hookeriopsis 

undata] 

Trachyxiphium aduncum (Mitt.) Buck [GU] [=Hookeriopsis 

adunca] 

Trachyxiphium guadalupense (Brid.) Buck [GU SU FG] 

[=Hookeriopsis guadalupensis] 

Trachyxiphium variabile (Mitt.) Buck [GU?] [=Hookeriopsis 

variabilis] 

HOOKERIACEAE - SYNONYMS AND 


Callicostella aspera (Mitt.) Jaeg. =Callicostella pallida 

Callicostella depressa (Hedw.) Jaeg. =Callicostella pallida 

Hookeriopsis adunca (Mitt.) Jaeg. =Trachyxiphium aduncum 

Hookeriopsis crispa (C. Müll.) Jaeg. =Thamniopsis undata 

Hookeriopsis diffusa (Wils.) Jaeg. =Thamniopsis diffusa 

Hookeriopsis falcata (Hook.) Jaeg. =Trachyxiphium 

guadalupense 

Hookeriopsis guadalupensis (Brid.) Jaeg. =Trachyxiphium 

guadalupense


Hookeriopsis guatemalensis E.B. Bartram =Callicostella 

guatemalensis 

Hookeriopsis parkeriana (Hook. & Grev.) Jaeg. =Brymela 

parkeriana 

Hookeriopsis undata (Hedw.) Jaeg. =Thamniopsis undata 

Hookeriopsis variabilis (Mitt.) Jaeg. =Trachyxiphium variabile 

Pilotrichum bipinnatum (Schwaegr.) Brid. =Callicosta 

bipinnatum 

Pilotrichum evanescens (C. Müll.) Crosby =Callicosta 

evanescens 

Pilotrichum fendleri C. Müll. =Callicosta fendleri 

Schizomitrium grossiretis (E.B. Bartram) Florsch. 

=Callicostella grossiretis 

Schizomitrium guatemalensis (E.B. Bartram) J. Florsch. 

=Callicostella guatemalensis 

Schizomitrium merkelii (Hornsch.) Florsch. =Callicostella 

merkelii 

Schizomitrium microcarpum (Ångstr.) Yano =Callicostella 

microcarpa 

Schizomitrium pallidum (Hornsch.) Crum & Anderson 

=Callicostella pallida 

Schizomitrium rufescens (Mitt.) Florsch. =Callicostella 

rufescens 

HYDROPOGONACEAE 

Hydropogon fontinaloides (Hook.) Brid. [SU FG] 

Hydropogonella gymnostoma (Schimp.) Card. [GU SU] 

HYPNACEAE 

Chrysohypnum diminutivum (Hampe) Buck [GU SU FG] 

Ectropothecium leptochaeton (Schwaegr.) Buck [GU SU FG] 

Isopterygium acutifolium Ireland [GU] 

Isopterygium fruticola (C. Müll.) Par. [SU] 

Isopterygium subbrevisetum (Hampe) Broth. [SU FG] 

Isopterygium tenerifolium Mitt. [GU SU] 

Isopterygium tenerum (Swartz) Mitt. [GU SU FG] 

Mittenothamnium reptans (Hedw.) Card. [GU SU FG] 

Rhacopilopsis trinitensis (C. Müll.) Britt. & Dix. [GU SU FG] 

Vesicularia vesicularis (Schwaegr.) Broth. [GU SU FG] 

HYPNACEAE - SYNONYMS AND 


Ectropothecium globitheca (C. Müll.) Mitt. =Ectropothecium 

leptochaeton 

Ectropothecium isopterygioides Card. & Thér. =Isopterygium 

tenerum 

Hypnum lonchophyllum Mont. =Sematophyllum 

lonchophyllum [Sematophyllaceae] 

Hypnum richardii Schwaegr. =Sematophyllum lonchophyllum 

[Sematophyllaceae] 

Isopterygium radicisetum (C. Müll.) Broth. =Isopterygium 

tenerum 

Mittenothamnium diminutivum (Hampe) Britt. 

=Chrysohypnum diminutivum 

Rhaphidorrhynchium leptochaeton (Schwaegr.) Broth. 

=Ectropothecium leptochaeton 

Vesicularia surinamense (Dozy & Molk.) Broth. =Vesicularia 

vesicularis 

LEUCOBRYACEAE 

Holomitriopsis laevifolium (Broth.) Robins. [GU] 

Leucobryum albicans (Schwaegr.) Lindb. [GU] 

[=Paraleucobryum albicans] 

Leucobryum albidum (Brid. ex P. Beauv.) Lindb. [GU FG] 

Leucobryum antillarum Schimp. ex Besch. [GU FG] 

Leucobryum crispum C. Müll. [GU SU FG] 

Leucobryum giganteum C. Müll. [GU FG?] [see also 

Leucobryum albicans] 

Leucobryum martianum (Hornsch.) C. Müll. [GU SU FG] 

Leucophanes molleri C. Müll. [GU SU FG] [Leucophanaceae] 

Octoblepharum albidum Hedw. [GU SU FG] 

Octoblepharum cocuiense Mitt. [GU SU FG] 

Octoblepharum cylindricum Mont. [GU SU FG] 

Octoblepharum erectifolium Mitt. ex Williams [GU SU FG] 

Octoblepharum pellucidum C. Müll. [GU SU FG] 

Octoblepharum pulvinatum (Dozy & Molk.) Mitt. [GU SU 

FG] 

Octoblepharum stramineum Mitt. [GU SU FG] 

LEUCOBRYACEAE - SYNONYMS AND 


Leucobryum laevifolium Broth. =Holomitriopsis laevifolium 

Leucophanes mittenii Card. ex Par. =Leucophanes molleri 

Paraleucobryum albicans (Schwaegr.) Loeske =Leucobryum 

albicans 

LEUCODONTACEAE 

Henicodium geniculatum (Mitt.) Buck [GU SU FG] [placed in 

Pterobryaceae by Buck] 

Pseudocryphaea flagellifera (Brid.) Britt. [GU SU FG] 

[=Pseudocryphaea domingensis?] 

LEUCODONTACEAE - SYNONYMS AND 


Leucodontopsis geniculata (Mitt.) Crum & Steere 

=Henicodium geniculatum 

Pseudocryphaea domingensis (Spreng.) Buck [see 

Pseudocryphaea flagellifera] 

LEUCOMIACEAE 

Leucomium steerei Allen & Veling [GU] 

Leucomium strumosum (Hornsch.) Mitt. [GU SU FG] 

LEUCOMIACEAE - SYNONYMS AND 


BRYOPHYTA 

19

20 

BRYOPHYTA 


Leucomium cuspidatum (C. Müll.) Jaeg. =Leucomium 

strumosum 

Leucomium guianense C. Müll. =Leucomium strumosum 

METEORIACEAE 

Floribundaria cardotii (Broth.) Broth. [SU] 

Floribundaria floribunda (Dozy & Molk.) Fleisch. [SU FG] 

Meteoridium remotifolium (C. Müll.) Manuel [GU SU FG] 

Meteoriopsis piligera (Müll.) Broth. [FG?] 

Papillaria deppei (Hornsch. ex C. Müll.) Jaeg. & Sauerb. [SU] 

Papillaria nigrescens (Hedw.) Jaeg. & Sauerb. [GU SU FG] 

Pilotrichella flexilis (Hedw.) Jaeg. [FG?] 

Pilotrichella pentasticha (Brid.) Wijk & Marg. [GU SU FG] 

Squamidium leucotrichum (Tayl.) Broth. [GU SU FG] 

Zelometeorium patulum (Hedw.) Manuel [GU SU FG] 

MNIACEAE 

Plagiomnium rhynchophorum (Hook.) T. Kop. [GU?] 

NECKERACEAE 

Isodrepanium lentulum (Wils.) Britt. [GU SU FG] 

Neckeropsis disticha (Hedw.) Kindb. [GU SU FG] 

Neckeropsis undulata (Hedw.) Reichdt. [GU SU FG] 

Porotrichum korthalsianum (Dozy & Molk.) Mitt. [SU FG] 

Porotrichum lindigii (Hampe) Mitt. [SU] 

Porotrichum molliculum Broth. [FG] 

Porotrichum piniforme (Brid.) Mitt. [SU FG] 

Porotrichum substriatum (Hampe) Mitt. [SU FG] 

NECKERACEAE - SYNONYMS AND 


Porotrichum cavifolium (Card. & Herz.) Magill & Allen 

=Porotrichum lindigii 

ORTHOTRICHACEAE 

Groutiella apiculata (Hook.) Crum & Steere [GU SU FG] 

Groutiella obtusa (Mitt.) Florsch. [GU SU FG] 

Groutiella tomentosa (Hornsch.) Wijk & Marg. [GU SU FG] 

Leiomitrium scaberrimum (Broth.) Wijk & Marg. [GU] 

Macromitrium cirrosum (Hedw.) Brid. [GU SU FG] 

Macromitrium dubium Schimp. ex C. Müll. [GU] 

Macromitrium fusco-aureum Bartr. [GU] 

Macromitrium harrisii Par. [GU] 

Macromitrium leprieurii Mont. [GU SU FG] 

Macromitrium pellucidum Mitt. [GU SU FG] 

Macromitrium perichaetiale (Hook. & Grev.) C. Müll. [GU 

SU] 

Macromitrium podocarpi C. Müll. [SU FG] 

Macromitrium portoricense Williams [GU SU FG] 

[=Macromitrium podocarpi?] 

Macromitrium punctatum (Hook. & Grev.) Brid. [GU SU FG] 

Macromitrium schwaneckeanum Hampe [GU] 

[=Macromitrium cirrosum?] 

Macromitrium scoparium Mitt. [FG] 

Macromitrium standleyi E.B. Bartram [GU] 

Macromitrium stellulatum (Hornsch.) Brid. [GU SU FG] 

Macromitrium trachypodium Mitt. [GU] 

Macromitrium trinitense Williams [GU SU] 

Macromitrium ulophyllum Mitt. [GU] 

Schlotheimia jamesonii (Arnott) Brid. [SU FG] 

Schlotheimia kegeliana (C. Müll.) C. Müll. [SU?] 

Schlotheimia rugifolia (Hook.) Schwaegr. [GU SU FG] 

Schlotheimia torquata (Hedw.) Brid. [GU SU FG] 

ORTHOTRICHACEAE - SYNONYMS AND 


Macromitrium apiculatum (Hook.) Brid. =Groutiella apiculata 

PHYLLODREPANIACEAE 

Mniomalia viridis (Mitt.) C. Müll. [GU SU FG] 

Phyllodrepanium falcifolium (Schwaegr.) Crosby [GU SU FG] 

PHYLLOGONIACEAE 

Phyllogonium fulgens (Hedw.) Brid. [GU SU FG] 

Phyllogonium viride Brid. [GU SU FG] 

POLYTRICHACEAE 

Polytrichum commune Hedw. [GU] 

Polytrichum juniperinum Hedw. [GU] 

POTTIACEAE 

Barbula agraria Hedw. [GU SU FG] 

Barbula arcuata Griff. [GU] 

Barbula indica (Hook.) Spreng. [GU SU FG] 

Hyophila involuta (Hook.) Jaeg. & Sauerb. [GU SU FG] 

Trichostomum brachydontium Bruch [GU] 

Trichostomum duidense Bartr. [GU?] [=Trichostomum 

brachydontium?] 

Trichostomum tenuirostre (Hook. & Tayl.) Lindb. [GU]


PTEROBRYACEAE 

Hildebrandtiella guyanensis (Mont.) Buck [GU FG] 

[=Orthostichidium guyanense] 

Jaegerina scariosa (Lor.) Artz. [GU SU FG] 

Orthostichopsis praetermissa Buck [GU SU? FG] 

Orthostichopsis tortipilis (C. Müll.) Broth. [GU? SU? FG?] 

[Guianas specimens probably =Orthostichopsis 

praetermissa] 

Orthostichopsis tetragona (Hedw.) Broth. [GU SU FG] 

Pireella cymbifolia (Sull.) Card. [GU? SU FG] 

Pireella pohlii (Schwaegr.) Card. [GU SU FG] 

Renauldia paradoxica Allen [GU FG] 

Spiridentopsis longissima (Raddi) Broth. [GU] 

PTEROBRYACEAE - SYNONYMS AND 


Orthostichidium guyanense (Mont.) Broth. =Hildebrandtiella 

guyanensis 

Orthostichopsis crinita (Sull.) Broth. =Orthostichopsis 

tortipilis [Guianas specimens referred to Orthostichopsis 

praetermissa by Buck] 

Pireella cavifolia (Card. & Herz.) Card. =Porotrichum lindigii 

(Neckeraceae) 

RACOPILACEAE 

Racopilum tomentosum (Hedw.) Brid. [GU SU FG] 

RHIZOGONIACEAE 

Pyrrhobryum spiniforme (Hedw.) Mitt. [GU SU FG] 

Rhizogonium lindigii (Hampe) Mitt. [GU?] 

SEMATOPHYLLACEAE 

Acroporium pungens (Hedw.) Broth. [GU SU FG] 

Donnellia commutata (C. Müll.) Buck [FG] 

Maguireella vulpina (Mont.) Buck [GU SU FG?] 

[synonymized under Potamium deceptivum by 

Florschütz-de Waard] 

Meiothecium boryanum (C. Müll.) Mitt. [GU SU FG] 

Meiothecium commutatum (C. Müll.) Broth. [FG] 

Phyllodon truncatulus (C. Müll.) Buck [FG] [placed in 

Hypnaceae by Florschütz-de Waard] 

Pterogonidium pulchellum (Hook.) C. Müll. ex Broth. [GU SU 

FG] 

Sematophyllum cochleatum (Broth.) Broth. [SU FG] 

Sematophyllum galipense (C. Müll.) Mitt. [GU SU] 

Sematophyllum lonchophyllum (Mont.) Florsch. [SU FG] 

Sematophyllum maguireorum Buck [GU] [synonymized under 

Potamium vulpinum by Florschütz-de Waard] 

Sematophyllum pacimoniense (Mitt.) J. Florsch. [GU SU] 

Sematophyllum subpinnatum (Brid.) Britt. [GU SU FG] 

Sematophyllum subsimplex (Hedw.) Mitt. [GU SU FG] 

Taxithelium concavum (Hook.) Spruce [GU SU FG] 

[=Taxithelium planum?] 

Taxithelium planum (Brid.) Mitt. [GU SU FG] 

Taxithelium pluripunctatum (Ren. & Card.) Buck [GU SU FG] 

Trichosteleum bolivarense Robins. [GU SU] [=Trichosteleum 

cyparissoides?] 

Trichosteleum hornschuchii (Hampe) Jaeg. [GU SU FG] 

Trichosteleum intricatum (Thér.) J. Florsch. [GU SU FG] 

Trichosteleum martii (C. Müll.) Kindb. [FG] [=Trichosteleum 

hornschuchii?] 

Trichosteleum papillosum (Hornsch.) Jaeg. [GU SU FG] 

Wijkia costaricensis (E.B. Bartram & Dix.) Crum [SU FG] 

SEMATOPHYLLACEAE - SYNONYMS AND 


Acroporium guianense (Mitt.) Broth. =Acroporium pungens 

Acroporium intricatum Thér. =Trichosteleum intricatum 

Glossadelphus truncatulus (C. Müll.) Fleisch. =Phyllodon 

truncatulus 

Meiothecium urceolatum (Schwaeg.) Mitt. [not in Guianas] 

Potamium deceptivum Mitt. =Maguireella vulpina 

Potamium lonchophyllum (Mont.) Mitt. =Sematophyllum 

lonchophyllum 

Potamium recurvifolium Thér. =Sematophyllum cochleatum 

Potamium uleanum Broth. =Sematophyllum lonchophyllum 

Potamium vulpinum (Mont.) Mitt. =Maguireella vulpina [see 

also Sematophyllum maguireorum] 

Rhaphidorrhynchium subsimplex (Hedw.) Broth. 

=Sematophyllum subsimplex 

Rhaphidostichum guianense E.B. Bartram =Sematophyllum 

galipense 

Sematophyllum guianense Mitt. =Acroporium pungens 

Sematophyllum kegelianum (C. Müll.) Mitt. =Sematophyllum 

subpinnatum 

Sematophyllum octodiceroides (C. Müll.) Broth. 

=Sematophyllum lonchophyllum 

Taxithelium impellucidum (C. Müll.) Par. =Taxithelium 

concavum? 

Taxithelium patulifolium Thér. =Taxithelium pluripunctatum 

Taxithelium quelchii (C. Müll.) Par. =Taxithelium concavum 

Taxithelium suboctodiceras Broth. & Par. [not in Guianas] 

Trichosteleum cyparissoides (Hornsch.) Robins. [see 

Trichosteleum bolivarense] 

Trichosteleum fluviale (Mitt.) Jaeg. =Trichosteleum 

hornschuchii 

Trichosteleum guianae (C. Müll.) Broth. =Trichosteleum 

papillosum 

Trichosteleum micropyxis (C. Müll.) Broth. =Trichosteleum 

hornschuchii 

Trichosteleum subsimplex (Hedw.) Mitt. =Sematophyllum 

subsimplex 

BRYOPHYTA 

21

22 


SPHAGNACEAE 

Sphagnum boomii Crum [GU] 

Sphagnum chi-chiense Crum [GU] 

Sphagnum cuspidatum Ehrh. ex Hoffm. var. serrulatum 

(Schlieph.) Schlieph. [GU SU] 

Sphagnum denticulatum Brid. [GU] 

Sphagnum magellanicum Brid. [GU] 

Sphagnum negrense Mitt. [GU] 

Sphagnum ornatum Crum [GU] [Mt. Roraima] 

Sphagnum oxyphyllum Warnst. [GU?] 

Sphagnum palustre L. [GU?] 

Sphagnum perichaetiale Hampe [GU SU] 

Sphagnum portoricense Hampe [GU] 

Sphagnum sanguinale Warnst. [GU] 

Sphagnum tenerum Sull. & Lesq. [GU] 

SPLACHNACEAE 

Splachnobryum obtusum (Brid.) C. Müll. [SU] 

STEREOPHYLLACEAE 

Entodontopsis leucostega (Brid.) Buck & Ireland [GU] 

Entodontopsis nitens (Mitt.) Buck & Ireland [GU SU] 

Eulacophyllum cultelliforme (Sulliv.) Buck & Ireland [FG] 

Pilosium chlorophyllum (Hornsch.) C. Müll. [GU SU FG] 

Stereophyllum radiculosum (Hook.) Mitt. [GU] 

STEREOPHYLLACEAE - SYNONYMS AND 


Entodontopsis rhabdodonta Card. =Entodontopsis contorteoperculata 

(C. Müll.) Broth. [not in Guianas] 

Pilosium flaccisetum C. Müll. =Pilosium chlorophyllum 

Pilosium pseudoradiculosum Müll. [nomen nudum, =Pilosium 

chlorophyllum] 

Plagiothecium unilaterale C. Müll. =Pilosium chlorophyllum 

Stereophyllum leucostegum (Brid.) Mitt. =Entodontopsis 

leucostega 

Stereophyllum nitens Mitt. =Entodontopsis nitens 

THUIDIACEAE 

Cyrtohypnum involvens (Hedw.) Buck & Crum [GU? SU 

FG?] 

Cyrtohypnum scabrosulum (Mitt.) Buck & Crum [GU SU FG] 

Cyrtohypnum schistocalyx (C. Müll.) Buck [SU FG] 

Thuidium peruvianum Mitt. [GU] 

Thuidium tomentosum Schimp. [GU SU FG] 

BRYOPHYTA, PSILOTOPHYTA, LYCOPHYTA 

THUIDIACEAE - SYNONYMS AND 


Thuidium acuminatum Mitt. =Thuidium urceolatum Lor. [not 

in Guianas] 

Thuidium antillarum Besch. =Thuidium tomentosum 

Thuidium campanulatum Mitt. [not in Guianas] 

Thuidium involvens (Hedw.) Mitt. =Cyrtohypnum involvens 

Thuidium pseudodelicatulum Jaeg. [not in Guianas] 

Thuidium scabrosulum Mitt. =Cyrtohypnum scabrosulum 

Thuidium schistocalyx C. Müll. =Cyrtohypnum schistocalyx 

REFERENCES: 

PSILOTOPHYTA 

PSILOTACEAE 

Kramer, K.U. 1990. Psilotaceae. In K.U. 

Kramer & P.S. Green, Families and Genera of 

Vascular Plants. I. Pteridophytes and 

Gymnosperms. New York: Springer-Verlag. 

Smith, A.R. 1995. Psilotaceae. In J.A. 

Steyermark, P.E. Berry, & B.K. Holst, Flora 

of the Venezuelan Guayana 2: 250-251. 

Psilotum nudum (L.) P. Beauv. [GU SU FG] 

LYCOPHYTA 

LYCOPHYTA REFERENCES: 

Berry, P.E., B.K. Holst, & K. Yatskievych. 1995. 

Flora of the Venezuelan Guayana, vol. 2. 

Pteridophytes, Spermatophytes Acanthaceae - 

Araceae. Missouri Botanical Garden, St. 

Louis, & Timber Press, Portland, Oregon. 



Phanérogames et de Ptéridophytes de la 

Guyane française d'après l'Herbier du Centre 

ORSTOM de Cayenne. ORSTOM, Cayenne. 

Cremers, G. & M. Hoff. 1990. Inventaire 

taxonomique des plantes de la Guyane 

française. I. Les Ptéridophytes. Mus. Nat. 

Hist. Nat., Inventaires de Faune et de Flore 

54: 1-133. 

Kramer, K.U. 1978. The Pteridophytes of 

Suriname. An enumeration with keys of the 

ferns and fern-allies. Uitgaven 

"Natuurwetenschappelijke Studiekring voor 

Suriname en de Nederlandse Antillen" Utrecht 

no. 93. 

Kramer, K.U. & P.S. Green [ed.]. 1990. The 

Families and Genera of Vascular Plants. I.


Pteridophytes and Gymnosperms. New York: 

Springer-Verlag. 

Posthumous, O. 1928. The Ferns of Surinam and 

of French and British Guiana. In A. Pulle, 

Flora of Suriname supplement: 1-196. 

Tryon, R.M. & A.F. 1981. Ferns and Allied 

Plants. New York: Springer-Verlag. 

ISOETACEAE 

reviewed by D. Lellinger & G. Cremers (1992); 

reviewed by D. Lellinger (1996) 

REFERENCES: 

Fuchs-Eckert, H.P. von. 1982. Zur heutigen 

Kenntnis von Vorkommen und Verbreitung 

der Südamerikanischen Isoëtes-Arten. Proc. 

Kon. Ned. Akad. Wetensch. (ser. C) 85: 205- 

260. 

Hickey, R.J. & A.R. Smith. 1995. Isoëtaceae. In 

J.A. Steyermark, P.E. Berry, & B.K. Holst, 

Flora of the Venezuelan Guayana 2: 189-190. 

Isoetes killipii C. Morton [GU] [Mt. Roraima] 

Isoetes ovata Pfeiffer [GU FG] 

Isoetes schinzii H.P. Fuchs [GU FG] 

Isoetes triangula U. Weber [GU] 

LYCOPODIACEAE 



REFERENCES: 

Oellgaard, B. 1987. A revised classification of 

the Lycopodiaceae s.l. Opera Bot. 92: 153- 

178. 

Oellgaard, B. 1995. Lycopodiaceae. In J.A. 



Oellgaard, B. & P.G. Windisch. 1987. Sinopse 

das Licopodiáceas do Brasil. Bradea 5(1): 1- 

43. 

Huperzia beitelii B. Oellg. [Mt. Roraima] 

Huperzia dichotoma (Jacq.) Trevis. [GU SU FG] 

Huperzia funiformis (Spring) Trevis. [GU] 

Huperzia intermedia Trevis. [Mt. Roraima] 

Huperzia killipii (Herter) Oellg. [FG] 

Huperzia linifolia (L.) Trevis. var. jenmanii (Underw. & 

Lloyd) B. Oellg. & Windisch [GU SU FG] 

Huperzia linifolia (L.) Trevis. var. linifolia [FG] 

Huperzia myrsinites (Lam.) Trevis. [GU] 

Huperzia recurvifolia Rolleri [GU] 

Huperzia reflexa (Lam.) Trevis. [Mt. Roraima] 

Huperzia taxifolia (Swartz) Trevis. [GU SU FG] 

Huperzia tenuicaulis (Underw. & Lloyd) B. Oellg. [GU] 

Huperzia wilsonii (Underw. & Lloyd) B. Oellg. [SU] 

Lycopodiella alopecuroides (L.) Cranfill var. integerrima 

(Spring) B. Oellg. & Windisch [GU SU] 

Lycopodiella camporum B. Oellg. & Windisch [GU SU FG] 

Lycopodiella caroliniana (L.) Pic.-Ser. var. meridionalis 

(Underw. & Lloyd) B. Oellg. & Windisch [GU SU FG] 

Lycopodiella cernua (L.) Pic.-Ser. var. cernua [GU SU FG] 

Lycopodiella cernua (L.) Pic.-Ser. var. curvata (Sw.) Kartesz 

& Gandhi [GU?] 

Lycopodiella iuliformis (Underw. & Lloyd) B. Oellg. [GU 

SU] 

Lycopodium clavatum L. subsp. contiguum (Klotzsch) B. 

Oellg. [GU] 

LYCOPODIACEAE - SYNONYMS AND 


Huperzia jenmanii (Underw. & Lloyd) Holub =Huperzia 

linifolia var. jenmanii 

Lycopodium alopecuroides L. =Lycopodiella alopecuroides 

Lycopodium anomalum Hook. & Grev. =Selaginella 

porelloides (Selaginellaceae) 

Lycopodium carolinianum L. =Lycopodiella caroliniana 

Lycopodium cernuum L. =Lycopodiella cernua 

Lycopodium contiguum Klotzsch =Lycopodium clavatum 

subsp. contiguum 

Lycopodium dichotomum Jacq. =Huperzia dichotoma 

Lycopodium duidae A.C. Smith =Lycopodiella iuliformis 

Lycopodium funiforme Cham. ex Spring =Huperzia 

funiformis 

Lycopodium intermedium Spring =Huperzia intermedia 

Lycopodium iuliforme Underw. & Lloyd =Lycopodiella 

iuliformis 

Lycopodium jenmanii Underw. & Lloyd =Huperzia linifolia 

var. jenmanii 

Lycopodium linifolium L. =Huperzia linifolia 

Lycopodium meridionale Underw. & Lloyd =Lycopodiella 

caroliniana var. meridionalis 

Lycopodium myrsinites Lam. =Huperzia myrsinites 

Lycopodium parkeri Hook. & Grev. =Selaginella parkeri 

(Selaginellaceae) 

Lycopodium penniforme Lam. =Selaginella radiata 


Lycopodium plumosum sensu Aublet, non L. =Selaginella 

epirrhizos (Selaginellaceae) 

Lycopodium radiatum Aubl. =Selaginella radiata 


Lycopodium reflexum Lam. =Huperzia reflexa 

Lycopodium roraimense Underw. & Lloyd =Huperzia 

myrsinites 

Lycopodium taxifolium Swartz =Huperzia taxifolia 

Lycopodium wilsonii Underw. & Lloyd =Huperzia wilsonii 

SELAGINELLACEAE 



LYCOPHYTA 

23

24 

REFERENCES: 

LYCOPHYTA, PTEROPHYTA 


Alston, A.H.G., A.C. Jermy, & J.M. Rankin. 

1981. The genus Selaginella in tropical South 

America. Bull. Brit. Mus. (Nat. Hist.) Bot. 9: 

233-330. 

Smith, A.R. 1995. Selaginellaceae. In J.A. 



Selaginella amazonica Spring [GU] 

Selaginella breynii Spring [GU FG] 

Selaginella cladorrhizans A. Braun [FG] 

Selaginella coarctata Spring [GU] 

Selaginella conduplicata Spring [GU SU FG] 

Selaginella convoluta (Arnott) Spring [GU FG?] 

Selaginella dendricola Jenm. [GU SU FG] 

Selaginella densifolia Spruce ex Hook. [SU? FG?] 

Selaginella diffusa (K. Presl) Spring [SU FG] 

Selaginella epirrhizos Spring [GU SU FG] 

Selaginella falcata (P. Beauv.) Spring [GU? FG] 

Selaginella flagellata Spring [GU SU FG] 

Selaginella hirsuta Alston ex Crabbe & Jermy [GU] 

Selaginella kochii Hieron. [SU FG] 

Selaginella mazaruniensis Jenm. [GU] 

Selaginella minima Spring [FG] 

Selaginella muscosa Spring [GU] 

Selaginella pallescens (K. Presl) Spring [SU] 

Selaginella parkeri (Hook. & Grev.) Spring [GU SU FG] 

Selaginella plana (Desv. ex Poir.) Hieron. [GU] [introduced] 

Selaginella porelloides (Lam.) Spring [GU FG] 

Selaginella potaroensis Jenm. [GU SU FG] 

Selaginella producta Baker [GU SU FG] 

Selaginella pruskiana Valdespino [GU] 

Selaginella radiata (Aubl.) Spring [GU SU FG] 

Selaginella revoluta Baker [GU SU FG] 

Selaginella rhodostachya Baker [GU] 

Selaginella roraimensis Baker [GU] 

Selaginella sandwithii Alston [GU SU FG] 

Selaginella scalariformis A.C. Smith [GU] 

Selaginella seemannii Baker [GU SU FG] 

Selaginella sulcata (Desv. ex Poir.) Spring ex Mart. [GU SU 

FG] 

Selaginella terezoana Bautista [GU] 

Selaginella tuberculata Spruce ex Baker [GU SU FG] 

Selaginella valdepilosa Baker [GU] 

Selaginella vernicosa Baker [GU] 

SELAGINELLACEAE - SYNONYMS AND 


Selaginella affinis A. Braun =Selaginella epirrhizos 

Selaginella brachyclada Baker =Selaginella rhodostachya 

Selaginella caudorrhiza Baker =Selaginella sulcata 

Selaginella cayennensis Baker =Selaginella producta 

Selaginella diminutifolia Jenm. =Selaginella rhodostachya 

Selaginella guianensis Spring =Selaginella falcata 

Selaginella jenmani Baker =Selaginella porelloides 

Selaginella lucidinervia Spring =Selaginella parkeri 

Selaginella macroclada Baker =Selaginella parkeri 

Selaginella parkeri (Hook. & Grev.) Spring var. pedata 

(Klotzsch) Baker =Selaginella parkeri 

Selaginella pedata Klotzsch =Selaginella parkeri 

Selaginella penniformis (Lam.) Hieron. =Selaginella radiata 

Selaginella platyphylla Baker =Selaginella porelloides 

Selaginella platyphylla Baker var. laxa Jenm. =Selaginella 

porelloides 

Selaginella poeppigiana (Spring) Spring ex Splitg. var. 

guyanensis Spring =Selaginella epirrhizos 

Selaginella stellata Spring =Selaginella conduplicata 

Selaginella valdepilosa Baker var. tricholoma Jermy & Rankin 

=Selaginella rhodostachya 

PTEROPHYTA 

PTEROPHYTA REFERENCES: 

Berry, P.E., B.K. Holst, & K. Yatskievych. 1995. 

Flora of the Venezuelan Guayana, vol. 2. 

Pteridophytes, Spermatophytes Acanthaceae - 

Araceae. Missouri Botanical Garden, St. 

Louis, & Timber Press, Portland, Oregon. 



Phanérogames et de Ptéridophytes de la 

Guyane française d'après l'Herbier du Centre 




française. I. Les Ptéridophytes. Mus. Nat. 

Hist. Nat., Inventaires de Faune et de Flore 

54: 1-133. 

Jenman, G.S. 1909. The ferns and fern allies of 

the British West Indies and Guiana. Trinidad: 

Bull. Misc. Int. Bot. Dept. 

Kramer, K.U. 1978. The Pteridophytes of 

Suriname. An enumeration with keys of the 

ferns and fern-allies. Uitgaven 

"Natuurwetenschappelijke Studiekring voor 

Suriname en de Nederlandse Antillen" Utrecht 

no. 93. 

Kramer, K.U. & P.S. Green [ed.]. 1990. The 

Families and Genera of Vascular Plants. I. 

Pteridophytes and Gymnosperms. New York: 

Springer-Verlag. 

Posthumous, O. 1928. The Ferns of Surinam and 

of French and British Guiana. In A. Pulle, 

Flora of Suriname supplement: 1-196. 

Tryon, R.M. & A.F. 1981. Ferns and Allied 

Plants. New York: Springer-Verlag. 

Vareschi, V. 1968. Helechos. In T. Lasser, 

Flora de Venezuela 1: 1-1033. 

ADIANTACEAE 


reviewed by D. Lellinger (1996)

REFERENCES: 


Lellinger, D.B. 1967. Pterozonium. In B. 

Maguire & collaborators, Botany of the 

Guayana Highland part 7. Mem. New York 

Bot. Gard. 17(1): 1-439. 

Lellinger, D.B. 1972. Polypodiaceae subfam. 

Adiantoideae. In B. Maguire & collaborators, 

Botany of the Guayana Highland part 9. 

Mem. New York Bot. Gard. 23: 1-832. 

Smith, A.R. & D.B. Lellinger. 1995. 

Pteridaceae. In J.A. Steyermark, P.E. Berry, 

& B.K. Holst, Flora of the Venezuelan 

Guayana 2: 250-286. 

Tryon, A.F. 1970. A monograph of the fern 

genus Eriosorus. Contrib. Gray Herb. 200: 

54-174. 

Tryon, R.M. 1942. A revision of the genus 

Doryopteris. Contrib. Gray Herb. 143: 1-80. 

Adiantopsis monticola (Gardner) T. Moore [GU] 

Adiantopsis radiata (L.) Fée [GU SU FG] 

Adiantum adiantoides (J.E. Smith) C. Chr. [GU SU FG] 

Adiantum cajennense Willd. ex Klotzsch [GU SU FG] 

Adiantum cordatum Maxon [FG?] [specimens identified as sp. 

nov. by Cremers] 

Adiantum deflectens Mart. [FG] 

Adiantum dolosum Kunze [GU SU FG] 

Adiantum fructuosum Poepp. ex Spreng. [GU SU FG] 

Adiantum fuliginosum Fée [GU SU FG] 

Adiantum glaucescens Klotzsch [GU SU FG] 

Adiantum glaziovii Baker [GU FG] 

Adiantum humile Kunze [GU SU FG] [records probably due 

to misidentifications] 

Adiantum latifolium Lam. [GU SU FG] 

Adiantum leprieurii Hook. [GU SU FG] 

Adiantum lucidum (Cav.) Swartz [GU SU FG] 

Adiantum macrophyllum Swartz [GU SU FG] 

Adiantum obliquum Willd. [GU SU FG] 

Adiantum olivaceum Baker [GU] 

Adiantum oyapokense Jenm. [FG] 

Adiantum petiolatum Desv. [GU SU FG] 

Adiantum phyllitidis J. Smith [GU SU FG] 

Adiantum pulverulentum L. [GU SU FG] 

Adiantum serratodentatum Willd. [GU SU FG] 

Adiantum terminatum Kunze ex Miq. [GU SU FG] 

Adiantum tetraphyllum Humb. & Bonpl. ex Willd. [GU SU 

FG] 

Adiantum tomentosum Klotzsch [GU SU FG] 

Adiantum tripteris K.U. Kramer [SU] 

Adiantum villosum L. [FG?] 

Doryopteris collina (Raddi) J. Smith [GU SU] 

Doryopteris conformis K.U. Kramer & R. Tryon [GU SU] 

Doryopteris lomariacea Kunze ex Klotzsch [GU] 

Doryopteris pedata (L.) Fée var. multipartita (Fée) Tryon [GU 

SU] 

Doryopteris sagittifolia (Raddi) J. Smith [GU SU FG] 

Doryopteris varians (Raddi) J. Smith [GU] 

Eriosorus flexuosus (Kunth) Copeland var. flexuosus [GU] 

Eriosorus hispidulus (Kunze) Vareschi var. hispidulus [GU] 

Eriosorus paucifolius (A.C. Smith) Vareschi var. neblinae A. 

Tryon [GU] 

Eriosorus paucifolius (A.C. Smith) Vareschi var. paucifolius 

[GU?] [Mt. Roraima] 

Hemionitis palmata L. [GU SU FG] 

Hemionitis rufa (L.) Swartz [GU SU FG] 

Notholaena solitaria Tryon [SU] 

Pityrogramma calomelanos (L.) Link var. aureoflava (Hook.) 

Weatherby ex Bailey [GU] 

Pityrogramma calomelanos (L.) Link var. calomelanos [GU 

SU FG] 

Pityrogramma ebenea (L.) Proctor var. ebenea [Mt. Roraima] 

Pterozonium cyclophyllum (Baker) Diels [GU] 

Pterozonium elaphoglossoides (Baker) Lellinger [GU] 

Pterozonium paraphysatum (A.C. Smith) Lellinger [GU SU] 

Pterozonium reniforme (Mart.) Fée [GU] 

Pterozonium scopulinum Lellinger [GU] 

ADIANTACEAE - SYNONYMS AND 


Adiantum guianense Aubl. =Lindsaea guianensis 

(Dennstaedtiaceae) 

Adiantum hirtum Klotzsch =Adiantum terminatum 

Adiantum propinquum Fée =Adiantum pulverulentum 

Adiantum sagittatum Aubl. =Lindsaea sagittata 


Adiantum strictum Swartz =Lindsaea stricta 


Gymnopteris rufa (L.) Bernhouillii ex Underw. =Hemionitis 

rufa 

Hemionitis cajenensis Desv. =Antrophyum cajenense 

(Vittariaceae) 

ASPLENIACEAE 



REFERENCES: 

Morton, C.V. & D.B. Lellinger. 1966. The 

Polypodiaceae subfam. Asplenioideae in 

Venezuela. Mem. New York Bot. Gard. 15: 

1-49. 

Smith, A.R. 1995. Aspleniaceae. In J.A. 



Asplenium abscissum Willd. [FG] 

Asplenium angustum Swartz [GU SU FG] 

Asplenium auritum Swartz var. auritum [GU SU FG] 

Asplenium auritum Swartz var. bipinnatifidum Kunze [GU] 

Asplenium claussenii Hieron. [GU?] 

Asplenium cristatum Lam. [GU SU FG] 

Asplenium cuneatum Lam. [GU SU FG] 

Asplenium delitescens (Maxon) L.D. Gómez [FG] 

Asplenium feei Kunze ex Fée [SU FG] 

Asplenium formosum Willd. [GU FG] 

Asplenium harpeodes Kunze [GU] 

Asplenium hostmannii Hieron. [SU FG] 

Asplenium juglandifolium Lam. [GU SU FG] 

Asplenium laetum Swartz [GU SU FG] 

PTEROPHYTA 

25

26 

PTEROPHYTA 


Asplenium macilentum Kunze ex Klotzsch [SU] 

Asplenium pearcei Baker [GU SU FG] 

Asplenium pedicularifolium St. Hil. [SU FG] 

Asplenium praemorsum Swartz [GU] 

Asplenium pteropus Kaulf. [GU SU] 

Asplenium radicans L. var. cirrhatum (L.C. Rich. ex Willd.) 

Rosenstock [GU FG] 

Asplenium radicans L. var. radicans [GU] 

Asplenium repandulum Kunze [FG] 

Asplenium riparium Liebm. [FG] 

Asplenium rutaceum (Willd.) Mett. [GU SU FG] 

Asplenium salicifolium L. [GU SU FG] 

Asplenium serra Langsd. & Fischer [GU SU] 

Asplenium serratum L. [GU SU FG] 

Asplenium sessilifolium Desv. [FG?] 

Asplenium zamiifolium Willd. [GU SU FG] 

ASPLENIACEAE - SYNONYMS AND 


Asplenium anisophyllum Auct. non Kunze =Asplenium feei 

Asplenium auritum Swartz var. obtusum Kunze ex Mett. 

=Asplenium macilentum 

Asplenium cirrhatum Rich. ex Willd. =Asplenium radicans 

var. cirrhatum 

Asplenium dejectum (Baker) Pittier =Hymenophyllopsis 

dejecta (Hymenophyllopsidaceae) 

Asplenium dimidiatum Auct. non Swartz =Asplenium 

zamiifolium 

Asplenium haplophyllum Domin =Asplenium pearcei 

Asplenium integerrimum Spreng. =Asplenium juglandifolium 

Asplenium kapplerianum Kunze =Asplenium juglandifolium 

Asplenium perkinsii Jenm. =Asplenium rutaceum 

Asplenium semicordatum Raddi =Asplenium salicifolium 

Asplenium sulcatum Auct. non Lam. =Asplenium auritum 

BLECHNACEAE 



REFERENCES: 

Smith, A.R. 1995. Blechnaceae. In J.A. 



Blechnum asplenioides Swartz [GU SU FG] 

Blechnum brasiliense Desv. [GU] 

Blechnum gracile Kaulf. [GU SU FG] 

Blechnum lanceola Swartz [SU] 

Blechnum lehmannii Hieron. [GU] 

Blechnum lineatum (Swartz) C. Chr. [GU] 

Blechnum occidentale L. [GU SU FG] 

Blechnum polypodioides Raddi [GU SU FG] 

Blechnum proliferum Rosenst. [GU] 

Blechnum schomburgkii (Klotzsch) C. Chr. [GU] 

Blechnum serrulatum L.C. Rich. [GU SU FG] 

Blechnum stipitellatum (Sodiro) C. Chr. [GU] 

Blechnum violaceum (Fée) C. Chr. [GU] 

Salpichlaena hookeriana (Kuntze) Alston [GU SU FG] 

Salpichlaena volubilis (Kaulf.) J. Smith [GU SU FG] 

BLECHNACEAE - SYNONYMS AND 


Blechnum indicum Auct. non Burm. f. =Blechnum serrulatum 

Blechnum volubile Kaulf. =Salpichlaena volubilis 

Stenochlaena marginata Auct. non (Schrad.) C. Chr. 

=Lomariopsis japurensis (Lomariopsidaceae) 

CYATHEACEAE 



REFERENCES: 

Conant, D.C. 1983. A revision of the genus 

Alsophila in the Americas. Journ. Arn. Arb. 

64: 333-382. 

Smith, A.R. 1995. Cyatheaceae. In J.A. 



Stolze, R.G. 1974. A taxonomic revision of the 

genus Cnemidaria. Fieldiana Bot. 37: 1-98. 

Tryon, R.M. 1970. The classification of the 

Cyatheaceae. Contrib. Gray Herb. 200: 3-53. 

Tryon, R.M. 1976. A revision of the genus 

Cyathea. Contrib. Gray Herb. 206: 19-101. 

Windisch, P.G. 1977. Synopsis of the genus 

Sphaeropteris, with a revision of the 

neotropical exindusiate species. Bot. Jahrb. 

Syst. 98(2): 176-198. 

Alsophila cuspidata (Kunze) Conant [FG] 

Alsophila imrayana (Hook.) Conant var. basilaris (Christ) 

Lellinger [SU FG] 

Cnemidaria cruciata (Desv.) Stolze [GU FG] 

Cnemidaria roraimensis (Domin) R. Tryon [GU] 

Cnemidaria spectabilis (Kunze) R. Tryon var. spectabilis [GU 

SU FG] 

Cyathea akawaiorum P.J. Edwards [GU] 

Cyathea andina (H. Karst.) Domin [GU SU FG] 

Cyathea bipinnatifida (Baker) Domin [GU] 

Cyathea caracasana (Klotzsch) Domin [GU?] [Mt. Roraima] 

Cyathea cyatheoides (Desv.) K.U. Kramer [GU SU FG] 

Cyathea delgadii Sternb. [GU] 

Cyathea dissimilis (C. Morton) Stolze [GU] 

Cyathea divergens Kunze var. divergens [GU SU] 

Cyathea gibbosa (Klotzsch) Domin [GU] 

Cyathea infesta (Kunze) Domin [FG] 

Cyathea lasiosora (Kuhn) Domin [FG] 

Cyathea macrocarpa (K. Presl) Domin [GU SU FG] 

Cyathea macrosora (Baker) Domin var. reginae (Windisch) 

A.R. Smith [GU] [=Cyathea traillii?] 

Cyathea marginalis (Klotzsch) Domin [GU SU FG] 

Cyathea microdonta (Desv.) Domin [GU SU FG] 

Cyathea multiflora J.E. Smith [GU] 

Cyathea nanna (Barr.) Lellinger [GU] 

Cyathea platylepis (Hook.) Domin [GU] 

Cyathea procera (Willd.) Domin [GU] [=Cyathea pungens?]


Cyathea pubens Domin [GU] 

Cyathea pungens (Willd.) Domin [GU SU FG] 

Cyathea surinamensis (Miq.) Domin [GU SU FG] 

Cyathea traillii (Baker) Domin [GU] 

Cyathea villosa Willd. [GU] 

CYATHEACEAE - SYNONYMS AND 


Alsophila blechnoides (Klotzsch) Hook. =Metaxya rostrata 

(Metaxyaceae) 

Alsophila marginalis Klotzsch =Cyathea marginalis 

Alsophila microdonta (Desv.) Desv. =Cyathea microdonta 

Alsophila oblonga Klotzsch =Cyathea procera 

Alsophila phalerata Mart. var. infesta Kunze =Cyathea infesta 

Alsophila pungens (Willd.) Kaulf. =Cyathea pungens 

Alsophila vestita K. Presl =Cyathea sp. 

Cyathea arborea Auct non (L.) J.E. Smith =Alsophila 

imrayana 

Cyathea cuspidata Kunze =Alsophila cuspidata 

Cyathea imrayana Hook. =Alsophila imrayana 

Cyathea oblonga (Klotzsch) Domin =Cyathea procera 

Cyathea oyapoka Jenm. =Cyathea cuspidata 

Cyathea roraimensis (Domin) Domin =Cnemidaria roraimensis 

Cyathea spectabilis (Kunze) Domin =Cnemidaria spectabilis 

Hemitelia boryana Mett. ex Kuhn =Cyathea andina 

Hemitelia hirsuta (Desv.) Weath. =Cyathea surinamensis 

Hemitelia hostmannii Hook. =Cyathea cyatheoides 

Hemitelia macrocarpa K. Presl =Cyathea macrocarpa 

Hemitelia multiflora (J.E. Smith) Spreng. =Cyathea 

cyatheoides 

Hemitelia parkeri Hook. =Cyathea surinamensis 

Hemitelia spectabilis Kunze =Cnemidaria spectabilis 

Sphaeropteris cyatheoides (Desv.) Windisch =Cyathea 

cyatheoides 

Sphaeropteris dissimilis (C. Morton) Windisch =Cyathea 

dissimilis 

Sphaeropteris hirsuta (Desv.) R. Tryon =Cyathea surinamensis 

Sphaeropteris macrocarpa (K. Presl) R. Tryon =Cyathea 

macrocarpa 

Sphaeropteris macrosora (Baker) Windisch =Cyathea 

macrosora 

DAVALLIACEAE 



REFERENCES: 

Cremers, G., K.U. Kramer, R.C. Moran, & A.R. 

Smith. 1994. Dryopteridaceae. In A.R.A. 

Görts-van Rijn, Flora of the Guianas (series 

B) 6: 3-65. 

Rumohra adiantiformis (G. Forster) Ching [GU] 

DAVALLIACEAE - SYNONYMS AND 


Davallia parkeri Hook. =Lindsaea parkeri (Dennstaedtiaceae) 

DENNSTAEDTIACEAE 



REFERENCES: 

Cremers, G. & K.U. Kramer. 1991. 

Dennstaedtiaceae. In A.R.A. Görts-van Rijn, 

Flora of the Guianas (series B) 4: 20-81. 

Kramer, K.U. 1957. A revision of the genus 

Lindsaea in the New World. Acta Bot. Neerl. 

6: 97-290. 

Smith, A.R. & K.U. Kramer. 1995. 

Dennstaedtiaceae. In J.A. Steyermark, P.E. 

Berry, & B.K. Holst, Flora of the Venezuelan 

Guayana 2: 46-71. 

Dennstaedtia obtusifolia (Willd.) T. Moore [GU SU FG] 

[=Dennstaedtia dissecta (Sw.) T. Moore?] 

Histiopteris incisa (Thunb.) J. Smith [GU] 

Hypolepis "sp. 1" [GU] [Mt. Roraima] 

Hypolepis "sp. 2" [GU] [Mt. Roraima] 

Hypolepis "sp. 3" [GU?] [Mt. Roraima] 

Hypolepis guianensis Klotzsch [GU] [Mt. Roraima] 

Hypolepis hostilis (Kunze) K. Presl [GU SU FG] 

Hypolepis nigrescens Hook. [GU] [Mt. Roraima] 

Hypolepis parallelogramma (Kunze) K. Presl [GU FG] 

Hypolepis repens (L.) K. Presl [GU SU FG] 

Lindsaea coarctata K.U. Kramer [GU] 

Lindsaea cultriformis K.U. Kramer [GU] 

Lindsaea cyclophylla K.U. Kramer [GU] 

Lindsaea divaricata Klotzsch [GU SU FG] 

Lindsaea dubia Spreng. [GU SU FG] 

Lindsaea falciformis Hook. [GU] [Mt. Roraima] [taxon of 

uncertain status] 

Lindsaea guianensis (Aubl.) Dryander subsp. guianensis [GU 

SU FG] 

Lindsaea javitensis Humb. & Bonpl. ex Willd. [GU] 

Lindsaea lancea (L.) Bedd. var. elatior (Kunze) K.U. Kramer 

[GU SU FG] 

Lindsaea lancea (L.) Bedd. var. falcata (Dryander) Rosenstock 

[GU SU FG] 

Lindsaea lancea (L.) Bedd. var. lancea [GU SU FG] 

Lindsaea lancea (L.) Bedd. var. leprieurii (Hook.) K.U. 

Kramer [GU SU FG] 

Lindsaea lancea (L.) Bedd. var. longifolia K.U. Kramer [GU] 

Lindsaea lancea (L.) Bedd. var. remota (Kunze) K.U. Kramer 

[GU SU FG] 

Lindsaea macrophylla Kaulf. [FG] 

Lindsaea mazarunensis Jenm. [GU] [species of uncertain 

status, probably a hybrid] 

Lindsaea pallida Klotzsch [GU SU FG] 

Lindsaea parkeri (Hook.) Kuhn subsp. parkeri [GU SU FG] 

Lindsaea pendula Klotzsch [GU SU] 

Lindsaea pleioptera K.U. Kramer [GU] 

Lindsaea portoricensis Desv. [GU SU FG] 

Lindsaea pumila Klotzsch [GU] [dubious species] 

Lindsaea quadrangularis Raddi subsp. antillensis K.U. Kramer 

[GU SU FG] 

PTEROPHYTA 

27

28 

PTEROPHYTA 


Lindsaea reniformis Dryander [GU SU FG] 

Lindsaea sagittata (Aubl.) Dryander [GU SU FG] 

Lindsaea schomburgkii Klotzsch [GU] 

Lindsaea semilunata (C. Chr.) C. Chr. [GU] 

Lindsaea sphenomeridopsis K.U. Kramer [GU] 

Lindsaea stricta (Swartz) Dryander var. jamesoniiformis K.U. 


Lindsaea stricta (Swartz) Dryander var. parvula (Fée) K.U. 


Lindsaea stricta (Swartz) Dryander var. stricta [GU SU FG] 

Lindsaea surinamensis Posth. [GU SU FG] 

Lindsaea tenuis Klotzsch [GU] 

Lindsaea tetraptera K.U. Kramer [GU] 

Lindsaea ulei Hieron. [GU] 

Lonchitis hirsuta L. [SU FG] 

Microlepia speluncae (L.) T. Moore [FG] [introduced] 

Odontosoria fumarioides (Swartz) J. Smith [FG?] 

Ormoloma imrayanum (Hook.) Maxon [GU] [cited from FG in 

error] 

Pteridium aquilinum (L.) Kuhn subsp. caudatum (L.) 

Bonaparte [GU SU FG] 

Saccoloma elegans Kaulf. subsp. chartaceum Nair ex Cremers 

& K.U. Kramer [GU FG] 

Saccoloma inaequale (Kunze) Mett. [GU SU FG] 

DENNSTAEDTIACEAE - SYNONYMS AND 


Dennstaedtia adiantiodes (Willd.) T. Moore =Dennstaedtia 

obtusifolia 

Ithycaulon inaequale (Kunze) Copeland =Saccoloma 

inaequale 

Lindsaea aquatica Jenm. =Lindsaea stricta var. parvula 

Lindsaea coriifolia Lindm. =Lindsaea schomburgkii 

Lindsaea crenata Klotzsch =Lindsaea portoricensis 

Lindsaea falcata Dryander =Lindsaea lancea var. falcata 

Lindsaea falcata Dryander var. elatior Kunze =Lindsea lancea 

var. elatior 

Lindsaea falcata Dryander var. lancea (L.) Jenm. =Lindsaea 

lancea var. lancea 

Lindsaea falcata Dryander var. leprieurii (Hook.) Jenm. 

=Lindsaea lancea var. leprieurii 

Lindsaea falcata Dryander var. subrotundifolia Jenm. [name 

of uncertain application] 

Lindsaea filiformis Hook. =Lindsaea tenuis 

Lindsaea gracilis Klotzsch =Lindsaea stricta var. parvula 

Lindsaea guianensis (Aubl.) Dryander var. venosa Jenm. 

[name of uncertain application] 

Lindsaea lancea (L.) Beddome var. semilunata C. Chr. 

=Lindsaea semilunata 

Lindsaea leprieurii Hook. =Lindsaea lancea var. leprieurii 

Lindsaea nitidissima Rich. ex Willd. =Lindsaea lancea 

Lindsaea parvula Fée =Lindsaea stricta var. parvula 

Lindsaea pumila Klotzsch var. remota Kunze =Lindsaea 

lancea var. remota 

Lindsaea pusilla Splitg. =Lindsaea lancea var. remota 

Lindsaea raddiana Klotzsch =Lindsaea stricta 

Lindsaea rufescens Kunze =Lindsaea portoricensis 

Lindsaea trapeziformis Dryander var. erythromera Kunze 

=Lindsaea divaricata 

Odontoloma parkeri (Hook.) K. Presl =Lindsaea parkeri 

Pteridium aquilinum (L.) Kuhn var. arachnoideum (Kaulf.) 

Brade =Pteridium aquilinum subsp. caudatum [treated as 

P. arachnoideum by Lellinger] 

Pteridium arachnoideum (Kaulf.) Maxon =Pteridium 

aquilinum subsp. caudatum 

Pteridium caudatum (L.) Maxon =Pteridium aquilinum subsp. 

caudatum 

Saccoloma imrayanum Hook. =Ormoloma imrayanum 

Schizoloma macrophyllum (Kaulf.) K. Presl =Lindsaea 

macrophylla 

Schizoloma reniforme (Dryander) Diels =Lindsaea reniformis 

Schizoloma sagittatum (Aubl.) Diels =Lindsaea sagittata 

DICKSONIACEAE 



REFERENCES: 

Maxon, W.R. 1922. The genus Culcita. J. 

Wash. Acad. Sci. 12: 454-460. 

Smith, A.R. 1995. Dicksoniaceae. In J.A. 



Culcita coniifolia (Hook.) Maxon [GU] 

DRYOPTERIDACEAE 



REFERENCES: 




B) 6: 3-65. 

Moran, R.C. 1986. The Neotropical fern genus 

Olfersia. Amer. Fern J. 76: 161-178. 

Moran, R.C. 1987. Monograph of the 

Neotropical fern genus Polybotrya 

(Dryopteridaceae). Ill. Nat. Hist. Surv. Bull. 

34(1): 1-138. 

Moran, R.C. 1991. Monograph of the 

Neotropical fern genus Stigmatopteris 

(Dryopteridaceae). Ann. Missouri Bot. Gard. 

78(4):: 857-914. 

Smith, A.R. 1986. Revision of the Neotropical 

fern genus Cyclodium. Am. Fern J. 76(2): 56- 

98. 

Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. 

Dryopteridaceae. In J.A. Steyermark, P.E. 


Guayana 2: 73-128. 

Arachniodes denticulata (Swartz) Ching [GU] [Mt. Roraima] 

Arachniodes leucostegioides (C. Chr.) Ching [GU] [Mt. 

Roraima]


Arachniodes macrostegia (Hook.) R. Tryon & Conant [GU?] 

[Mt. Roraima] 

Arachniodes ochropteroides (Baker) Lellinger [GU SU] 

Cyclodium akawaiorum A.R. Smith [GU] [Mt. Roraima] 

Cyclodium guianense (Klotzsch) van der Werff ex L.D. 

Gómez [GU SU FG] 

Cyclodium inerme (Fée) A.R. Smith [GU SU FG] 

Cyclodium meniscioides (Willd.) K. Presl var. meniscioides 

[GU SU FG] 

Cyclodium meniscioides (Willd.) K. Presl var. rigidissimum 

(C. Chr.) A.R. Smith [GU] 

Cyclodium rheophilum A.R. Smith [FG] 

Cyclodium varians (Fée) A.R. Smith [GU] 

Didymochlaena truncatula (Swartz) J. Smith [GU SU FG] 

Olfersia cervina (L.) Kunze [GU SU FG] 

Polybotrya caudata Kunze [GU SU FG] 

Polybotrya cyathifolia Fée [FG] 

Polybotrya fractiserialis (Baker) J.E. Smith [GU SU FG] 

Polybotrya lechleriana Mett. [GU] [Mt. Roraima] 

Polybotrya osmundacea Humb. & Bonpl. ex Willd. [GU SU 

FG] 

Polybotrya sessilisora R.C. Moran [GU?] [Brazil/Guyana 

border area] 

Polybotrya sorbifolia Kuhn [to be expected in the Guianas] 

Stigmatopteris longicaudata (Liebm.) C. Chr. [FG] 

Stigmatopteris rotundata (Humb. & Bonpl. ex Willd.) C. Chr. 

[GU SU FG] 

DRYOPTERIDACEAE - SYNONYMS AND 


Aspidium confertum Kaulf. =Cyclodium meniscioides var. 

meniscioides 

Aspidium guianense Klotzsch =Cyclodium guianense 

Cyclodium confertum (Kaulf.) K. Presl =Cyclodium 

meniscioides var. meniscioides 

Cyclodium rigidissimum C. Chr. =Cyclodium meniscioides 

var. rigidissimum 

Dryopteris amplissima (K. Presl) Kunze var. subeffusa C. Chr. 

=Lastreopsis amplissima (Tectariaceae) 

Dryopteris arborea Brause =Thelypteris arborea 

(Thelypteridaceae) 

Dryopteris brachypoda (Baker) C. Chr. =Thelypteris 

brachypoda (Thelypteridaceae) 

Dryopteris carrii Auct. non (Baker) C. Chr. =Stigmatopteris 

rotundata 

Dryopteris decussata (L.) Urb. =Thelypteris decussata 


Dryopteris dejecta (Jenm.) C. Chr. =Thelypteris glandulosa 


Dryopteris demerarana (Baker) C. Chr. =Thelypteris 

demerarana (Thelypteridaceae) 

Dryopteris dentata Auct. non (Forsskal) C. Chr. =Thelypteris 

hispidula (Thelypteridaceae) 

Dryopteris effusa (Swartz) Urb. =Lastreopsis effusa 

(Tectariaceae) 

Dryopteris extensa (Blume) Kuntze =Thelypteris opulenta 


Dryopteris funesta (Kunze) C. Chr. =Triplophyllum funestum 


Dryopteris gongylodes (Schkuhr) Kuntze =Thelypteris 

interrupta (Thelypteridaceae) 

Dryopteris gonophora Weath. =Thelypteris gonophora 


Dryopteris guianensis (Klotzsch) Posth. =Cyclodium 

guianense 

Dryopteris hostmannii (Klotzsch) Maxon & C. Morton 

=Thelypteris arborescens (Thelypteridaceae) 

Dryopteris kanukuana Weath. =Thelypteris pennata 


Dryopteris leprieurii (Hook.) Kuntze =Thelypteris leprieurii 


Dryopteris macrophylla (Kunze) C. Chr. =Thelypteris 

macrophylla (Thelypteridaceae) 

Dryopteris martinii C. Chr. =Thelypteris connexa 

(Thelypteridaceae)[type erroneously cited from French 

Guiana] 

Dryopteris megalodus (Schkuhr) Urb. =Thelypteris pennata 


Dryopteris meniscioides (Willd.) Kuntze =Cyclodium 

meniscioides 

Dryopteris meniscioides (Willd.) Kuntze var. conferta (Kaulf.) 

C. Morton =Cyclodium meniscioides var. meniscioides 

Dryopteris nesiotica Maxon & C. Morton =Thelypteris 

nesiotica (Thelypteridaceae) 

Dryopteris ochropteroides (Baker) C. Chr. =Arachniodes 

ochropteroides 

Dryopteris poiteana (Bory) Urb. =Thelypteris poiteana 


Dryopteris protensa (Afzel. ex Swartz) C. Chr. var. funesta 

(Kunze) C. Chr. =Triplophyllum funestum (Tectariaceae) 

Dryopteris pyramidata (Fée) Maxon =Thelypteris abrupta 


Dryopteris quadrangularis (Fée) Alston =Thelypteris hispidula 


Dryopteris refulgens (Klotzsch) C. Chr. =Ctenitis refulgens 


Dryopteris refulgens (Klotzsch) C. Chr. =Ctenitis refulgens 


Dryopteris roraimensis Brause =Thelypteris arborea 


Dryopteris sancti-gabrieli (Hook.) Kuntze =Cyclodium 

guianense 

Dryopteris serrata (Cav.) C. Chr. =Thelypteris serrata 


Dryopteris sorbifolia Auct. non (Jacq.) Hieron. =Thelypteris 

hostmannii (Thelypteridaceae) 

Dryopteris subcuneata (Baker) Kuntze =Thelypteris abrupta 


Dryopteris subobliquata (Hook.) Kuntze =Cyclodium inerme 

Dryopteris tetragona (Swartz) Urb. =Thelypteris tetragona 


Polybotrya cervina (L.) Kaulf. =Olfersia cervina 

Polybotrya pubens Auct. non Mart. =Polybotrya caudata 

Polystichopsis ochropteroides (Baker) C. Morton 

=Arachniodes ochropteroides 

Polystichum inerme Fée =Cyclodium inerme 

Stigmatopteris guianensis (Klotzsch) C. Chr. =Cyclodium 

guianense 

Stigmatopteris meniscioides (Willd.) K.U. Kramer 

=Cyclodium meniscioides 

Stigmatopteris sancti-gabrieli (Hook.) C. Chr. =Cyclodium 

guianense 

Stigmatopteris varians (Fée) Alston =Cyclodium varians 

GLEICHENIACEAE 

PTEROPHYTA 

29

30 

PTEROPHYTA 




REFERENCES: 

Smith, A.R. 1995. Gleicheniaceae. In J.A. 



Dicranopteris flexuosa (Schrad.) Underw. [GU SU FG] 

Dicranopteris nervosa (Kaulf.) Maxon [GU] 

Dicranopteris pectinata (Willd.) Underw. [GU SU FG] 

Dicranopteris schomburgkiana (J.W. Sturm) C. Morton [Mt. 

Roraima] 

Gleichenia bifida (Willd.) Spreng. [GU SU FG] 

Gleichenia longipinnata Hook. [GU SU] 

Gleichenia nitidula Rosenst. [GU] 

Gleichenia pennigera (Mart.) T. Moore [GU] 

Gleichenia remota (Kaulf.) Spreng. [GU SU FG] 

Gleichenia revoluta Kunth [GU] [Mt. Roraima] 

GLEICHENIACEAE - SYNONYMS AND 


Dicranopteris nitidula (Rosenst.) Suesseng ex Losch 

=Gleichenia nitidula 

Dicranopteris remota (Kaulf.) Maxon =Gleichenia remota 

Gleichenia linearis Auct. non (Burm.) Clarke =Dicranopteris 

flexuosa 

Gleichenia pectinata (Willd.) K. Presl =Dicranopteris pectinata 

Sticherus bifidus (Willd.) Ching =Gleichenia bifida 

Sticherus pennigera (Mart.) Copel. =Gleichenia pennigera 

Sticherus remotus (Kaulf.) Chrysler =Gleichenia remota 

Sticherus revolutus (Kunth) Ching =Gleichenia revoluta 

GRAMMITIDACEAE 



REFERENCES: 

Bishop, L.E. 1977. The American species of 

Grammitis sect. Grammitis. Amer. Fern J. 67: 

101-106. 

Bishop, L.E. 1978. Revision of the genus 

Cochlidium. Amer. Fern J. 68: 76-94. 

Bishop, L.E. 1988. Ceradenia, a new genus of 

Grammitidaceae. Amer. Fern J. 78: 1-5. 

Bishop, L.E. & A.R. Smith. 1992. Revision of 

the fern genus Enterosora (Grammitidaceae) 

in the New World. Syst. Bot. 17: 345-362. 

Bishop, L.E. & A.R. Smith. 1995. 

Grammitidaceae. In J.A. Steyermark, P.E. 


Guayana 2: 135-158. 

Copeland, E.B. 1952. The American species of 

Xiphopteris. Amer. Fern J. 42: 41-52, 93- 

110. 

Smith, A.R. 1992. A review of the fern genus 

Micropolypodium (Grammitidaceae). Novon 

2(4): 419-425. 

Smith, A.R. 1993. Terpsichore, a new genus of 

Grammitidaceae (Pteridophyta). Novon 3(4): 

478-489. 

Smith, A.R. & R.C. Moran. 1992. Melpomene, a 

new genus of Grammitidaceae (Pteridophyta). 

Novon 2(4): 426-432. 

Smith, A.R., R.C. Moran, & L.E. Bishop. 1991. 

Lellingeria, a new genus of Grammitidaceae. 

Amer. Fern J. 81: 76-88. 

Ceradenia kookenamae (Jenm.) L.E. Bishop [GU] [Mt. 

Roraima] 

Cochlidium attenuatum A.C. Smith [GU] 

Cochlidium connellii (Baker ex C.H. Wright) A.C. Smith 

[GU] 

Cochlidium furcatum (Hook. & Grev.) C. Chr. [GU SU FG] 

Cochlidium linearifolium (Desv.) Maxon ex C. Chr. [GU SU 

FG] 

Cochlidium seminudum (Willd.) Maxon [GU? FG?] 

Cochlidium serrulatum (Swartz) L.E. Bishop [GU SU FG] 

Cochlidium tepuiense (A.C. Smith) L.E. Bishop [GU] 

Enterosora campbellii Baker subsp. campbellii [GU? FG?] 

Enterosora trifurcata (L.) L.E. Bishop [GU SU] 

Grammitis anfractuosa (Kunze ex Klotzsch) Proctor [GU] 

[=Melpomene anfractuosa] 

Grammitis asplenifolia (L.) Proctor [GU] [=Terpsichore 

asplenifolia] 

Grammitis blanchetii (C. Chr.) A.R. Smith [GU SU FG] 

[=Micropolypodium nanum] 

Grammitis bryophila (Maxon) Seymour [GU] 

Grammitis capillaris (Desv.) Proctor [Mt. Roraima] 

[=Ceradenia capillaris] 

Grammitis cultrata (Bory ex Willd.) Proctor [GU] 

[=Terpsichore cultrata] 

Grammitis discolor (Hook.) C. Morton [GU] [=Ceradenia 

discolor] 

Grammitis firma (J. Smith) C. Morton [GU] [=Melpomene 

firma] 

Grammitis flabelliformis (Poir.) C. Morton [GU?] [Mt. 

Roraima] [=Melpomene flabelliformis] 

Grammitis fragillima (Copel.) C. Morton [Mt. Roraima] 

[=Ceradenia fragillima] 

Grammitis kaieteura (Jenm.) C. Morton [GU] [=Grammitis 

tenuicula (Fée) Proctor?] 

Grammitis kalbreyeri (Baker) C. Morton [GU SU FG] 

[=Ceradenia kalbreyeri] 

Grammitis kegeliana (Kunze) Lellinger [SU FG] 

[=Terpsichore kegeliana] 

Grammitis leptopoda (C.H. Wright) Copeland [GU?] [Mt. 

Roraima] 

Grammitis melanosticta (Kunze) F. Seym. [GU SU] 

[=Melpomene melanosticta] 

Grammitis melanotrichia (Baker) Lellinger [Mt. Roraima] 

[=Lellingeria melanotrichia] 

Grammitis mollissima (Fée) Proctor [GU SU FG] 

[=Terpsichore mollissima] 

Grammitis moniliformis (Lag. ex Swartz) Proctor [GU] 

[=Melpomene moniliformis] 

Grammitis myosuroides (Swartz) Swartz [Mt. Roraima] 

[=Lellingeria myosuroides]


Grammitis paramicola L.E. Bishop [Mt. Roraima] 

Grammitis peritimundi L.E. Bishop & A.R. Smith [Mt. 

Roraima] 

Grammitis pilosissima (Martens & Galeotti) C. Morton [SU] 

[=Melpomene pilosissima] 

Grammitis randallii (Maxon) Proctor [GU] 

Grammitis shaferi (Maxon) Lellinger [SU] [=Lellingeria 

shaferi] 

Grammitis sinuosa A.R. Smith [GU SU] [=Lellingeria 

sinuosa] 

Grammitis staheliana (Posth.) Lellinger [SU FG] 

[=Terpsichore staheliana] 

Grammitis subsessilis (Baker) C. Morton [GU] [=Lellingeria 

subsessilis] 

Grammitis suspensa (L.) Proctor [GU SU FG] [=Lellingeria 

suspensa] 

Grammitis taenifolia (Jenm.) Proctor [GU SU FG] 

[=Micropolypodium taenifolium] 

Grammitis tatei Copeland [GU] 

Grammitis taxifolia (L.) Proctor [GU SU FG] [=Terpsichore 

taxifolia] 

Grammitis trichomanoides (Swartz) Ching [GU FG] 

[=Micropolypodium trichomanoides] 

Grammitis xanthotrichia (Klotzsch) Duek & Lellinger [Mt. 

Roraima] [=Terpsichore xanthotrichia] 

GRAMMITIDACEAE - SYNONYMS AND 


Ceradenia capillaris (Desv.) L.E. Bishop =Grammitis capillaris 

Ceradenia discolor (Hook.) L.E. Bishop =Grammitis discolor 

Ceradenia fragillima (Copel.) L.E. Bishop =Grammitis 

fragillima 

Ceradenia kalbreyeri (Baker) L.E. Bishop =Grammitis 

kalbreyeri 

Ctenopteris jubiformis (Kaulf.) J.E. Smith =Grammitis 

suspensa 

Ctenopteris kaieteura (Jenm.) Copeland =Grammitis kaieteura 

Ctenopteris kalbreyeri (Baker) Copeland =Grammitis 

kalbreyeri 

Ctenopteris kegeliana (Kunze) K.U. Kramer =Grammitis 

kegeliana 

Ctenopteris melanosticta (Kunze) Copeland =Grammitis 

melanosticta 

Ctenopteris mollissima (Fée) Copeland =Grammitis 

mollissima 

Ctenopteris pilosissima (Martens & Galeotti) Copeland 

=Grammitis pilosissima 

Ctenopteris staheliana (Posth.) K.U. Kramer =Grammitis 

staheliana 

Ctenopteris subsessilis (Baker) Copeland =Grammitis 

subsessilis 

Ctenopteris suspensa (L.) Copeland =Grammitis suspensa 

Ctenopteris taxifolia (L.) Copeland =Grammitis taxifolia 

Ctenopteris trifurcata (L.) Pic.-Ser. =Enterosora trifurcata 

Grammitis furcata Hook. & Grev. =Cochlidium furcatum 

Grammitis nana Fée =Grammitis blanchetii 

Grammitis serrulata (Swartz) Swartz =Cochlidium serrulatum 

Grammitis tenuicula (Fée) Proctor [see Grammitis kaieteura] 

Grammitis trifurcata (L.) Copeland =Enterosora trifurcata 

Lellingeria melanotrichia (Baker) A.R. Smith & R.C. Moran 

=Grammitis melanotrichia 

Lellingeria myosuroides (Swartz) A.R. Smith & R.C. Moran 

=Grammitis myosuroides 

Lellingeria shaferi (Maxon) A.R. Smith & R.C. Moran 

=Grammitis shaferi 

Lellingeria sinuosa (A.R. Smith) A.R. Smith & R.C. Moran 

=Grammitis sinuosa 

Lellingeria subsessilis (Baker) A.R. Smith & R.C. Moran 

=Grammitis subsessilis 

Lellingeria suspensa (L.) A.R. Smith & R.C. Moran 

=Grammitis suspensa 

Melpomene anfractuosa (Kunze ex Klotzsch) A.R. Smith & 

R.C. Moran =Grammitis anfractuosa 

Melpomene firma (J. Smith) A.R. Smith & R.C. Moran 

=Grammitis firma 

Melpomene flabelliformis (Poir.) A.R. Smith & R.C. Moran 

=Grammitis flabelliformis 

Melpomene melanosticta (Kunze) A.R. Smith & R.C. Moran 

=Grammitis melanosticta 

Melpomene moniliformis (Lagasca ex Sw.) A.R. Smith & R.C. 

Moran =Grammitis moniliformis 

Melpomene pilosissima (M. Martens & Galeotti) A.R. Smith 

& R.C. Moran =Grammitis pilosissima 

Micropolypodium nanum (Fée) A.R. Smith =Grammitis 

blanchetii 

Micropolypodium taenifolium (Jenm.) A.R. Smith =Grammitis 

taenifolia 

Micropolypodium trichomanoides (Sw.) A.R. Smith 

=Grammitis trichomanoides 

Monogramma linearifolia Desv. =Cochlidium linearifolium 

Terpsichore asplenifolia (L.) A.R. Smith =Grammitis 

asplenifolia 

Terpsichore cultrata (Bory ex Willd.) A.R. Smith =Grammitis 

cultrata 

Terpsichore kegeliana (Kunze) A.R. Smith =Grammitis 

kegeliana 

Terpsichore mollissima (Fée) A.R. Smith =Grammitis 

mollissima 

Terpsichore staheliana (Posth.) A.R. Smith =Grammitis 

staheliana 

Terpsichore taxifolia (L.) A.R. Smith =Grammitis taxifolia 

Terpsichore xanthotrichia (Klotzsch) A.R. Smith =Grammitis 

xanthotrichia 

Xiphopteris nana (Fée) Copeland =Grammitis blanchetii 

Xiphopteris serrulata (Swartz) Kaulf. =Cochlidium serrulatum 

Xiphopteris shaferi (Maxon) K.U. Kramer =Grammitis shaferi 

Xiphopteris taenifolia (Jenm.) Copeland =Grammitis taenifolia 

Xiphopteris trichomanoides (Swartz) Copeland =Grammitis 

trichomanoides 

HYMENOPHYLLACEAE 

contributed by D. Lellinger (1992); reviewed by 

D. Lellinger (1996) 

REFERENCES: 

Lellinger, D.B. 1984. Hymenophyllaceae. In B. 



Bot. Gard. 38: 1-84. 

Lellinger, D.B. 1991. Notes on neotropical 

Hymenophyllaceae. Amer. Fern J. 81: 42-46. 

Lellinger, D.B. 1994. Trichomanes 

polypodioides and its allies. Amer. Fern J. 

84(1): 1-4. 

PTEROPHYTA 

31

32 

PTEROPHYTA 


Lellinger, D.B. 1994. Hymenophyllaceae. In 

A.R.A. Görts-van Rijn, Flora of the Guianas 

(series B) 3: 1-66. 

Smith, A.R. 1995. Hymenophyllaceae. In J.A. 



Hymenophyllum antillense (Jenm.) Jenm. [Mt. Roraima] 

Hymenophyllum apiculatum Mett. ex Kuhn [GU SU] 

Hymenophyllum asplenioides (Swartz) Swartz [GU] 

Hymenophyllum brevifrons Kunze [FG] 

Hymenophyllum decurrens (Jacq.) Swartz [GU SU FG] 

Hymenophyllum dependens C. Morton [GU] 

Hymenophyllum elegans Spreng. [GU] 

Hymenophyllum fendlerianum J.W. Sturm [GU SU] 

Hymenophyllum fucoides (Swartz) Swartz [GU SU] 

Hymenophyllum funckii Bosch [FG?] 

Hymenophyllum hirsutum (L.) Swartz [GU SU FG] 

Hymenophyllum kaieteurum Jenm. [GU] 

Hymenophyllum lanatum Fée [SU] 

Hymenophyllum lehmannii Hieron. [GU] 

Hymenophyllum microcarpum Desv. [GU] [Mt. Roraima] 

Hymenophyllum polyanthos (Swartz) Swartz [GU SU FG] 

Hymenophyllum roraimense C. Morton [Mt. Roraima] 

Hymenophyllum trichomanoides Bosch [GU] 

Hymenophyllum trichophyllum Kunth [GU] 

Hymenophyllum valvatum Hook. & Grev. [GU] [Mt. 

Roraima] 

Trichomanes accedens K. Presl [GU SU FG] 

Trichomanes angustifrons (Fée) W. Boer [GU SU FG] 

Trichomanes ankersii Parker ex Hook. & Grev. [GU SU FG] 

Trichomanes anomalum Maxon & C. Morton [GU SU?] 

Trichomanes arbuscula Desv. [GU SU FG] 

Trichomanes bancroftii Hook. & Grev. [GU] 

Trichomanes bicorne Hook. [GU] 

Trichomanes botryoides Kaulf. [GU SU FG] 

Trichomanes caliginum Lellinger [GU SU] 

Trichomanes cellulosum Klotzsch [GU SU?] 

Trichomanes crinitum Swartz [GU] 

Trichomanes crispum L. [GU SU FG] 

Trichomanes cristatum Kaulf. [GU SU? FG] 

Trichomanes curranii Weath. [GU SU FG?] 

Trichomanes dactylites Sodiro [Mt. Roraima] 

Trichomanes diaphanum Kunth [GU SU FG] 

Trichomanes diversifrons (Bory) Mett. ex Sadebeck [GU SU 

FG] 

Trichomanes egleri Windisch [GU SU] 

Trichomanes elegans L.C. Rich. [GU SU FG] 

Trichomanes hostmannianum (Klotzsch) Kunze [GU SU FG] 

Trichomanes humboldtii (Bosch) Lellinger [GU] 

Trichomanes hymenophylloides van den Bosch [GU?] [Mt. 

Roraima] 

Trichomanes jenmanii Lellinger [GU] 

Trichomanes kapplerianum J.W. Sturm [GU SU FG] 

Trichomanes krausii Hook. & Grev. [GU SU FG] 

Trichomanes lucens Swartz [GU?] [Mt. Roraima] 

Trichomanes macilentum Bosch [GU] 

Trichomanes martiusii K. Presl [GU SU] 

Trichomanes membranaceum L. [GU SU FG] 

Trichomanes mougeotii Bosch [SU FG] 

Trichomanes osmundoides DC. ex Poir. [GU FG] 

Trichomanes ovale (Fourn.) W. Boer [SU] 

Trichomanes pedicellatum Desv. [GU SU FG] 

Trichomanes pellucens Kunze [GU SU FG] 

Trichomanes pinnatinervium Jenm. [GU SU FG] 

Trichomanes pinnatum Hedw. [GU SU FG] 

Trichomanes plumosum Kunze [GU] 

Trichomanes polypodioides L. [GU SU FG] 

Trichomanes punctatum Poir. subsp. labiatum (Jenm.) W. Boer 

[GU SU FG] 

Trichomanes pyxidiferum L. [FG] 

Trichomanes radicans Swartz var. radicans [GU SU FG] 

Trichomanes rigidum Swartz [GU SU FG] 

Trichomanes roraimense Jenm. [GU SU?] 

Trichomanes rupestre (Raddi) Bosch [FG] 

Trichomanes spruceanum Hook. [GU] 

Trichomanes sprucei Baker [GU?] [Mt. Roraima] 

Trichomanes steyermarkii Windisch & A.R. Smith [GU] [Mt. 

Roraima] 

Trichomanes tanaicum Hook. ex J.W. Sturm [GU] 

Trichomanes trollii Bergd. [GU SU FG] 

Trichomanes tuerckheimii Christ [GU SU FG] 

Trichomanes vandenboschii Windisch [GU SU] 

Trichomanes vittaria DC. ex Poir. [GU SU FG] 

Trichomanes windischianum Lellinger [GU SU] 

HYMENOPHYLLACEAE - SYNONYMS AND 


Davalliopsis elegans (Rich.) Copel. =Trichomanes elegans 

Feea boryi Bosch, nom. superfl. =Trichomanes diversifrons 

Feea botryoides (Kaulf.) Bosch =Trichomanes botryoides 

Feea humboldtii Bosch =Trichomanes humboldtii 

Feea nana Bory =Trichomanes botryoides 

Hymenophyllum amoenum (Klotzsch) J.W. Sturm 

=Hymenophyllum fendlerianum 

Hymenophyllum ciliatum (Swartz) Swartz =Hymenophyllum 

hirsutum 

Hymenophyllum ciliatum (Swartz) Swartz var. ornifolium 

Kunze =Hymenophyllum hirsutum 

Hymenophyllum dejectum Baker =Hymenophyllopsis dejecta 

(Hymenophyllopsidaceae) 

Hymenophyllum myriocarpum Hook. [records due to 

misidentifications] 

Hymenophyllum protrusum Hook. [records due to 


Hymenophyllum schomburgkii K. Presl ex J.W. Sturm 

=Hymenophyllum decurrens 

Hymenophyllum surinamense Bosch, nom. superfl. 

=Hymenophyllum hirsutum 

Hymenophyllum trapezoidale Liebm. [records due to 


Hymenophyllum undulatum (Swartz) Swartz var. 

fendlerianum (J.W. Sturm) Stolzez =Hymenophyllum 

fendlerianum 

Hymenostachys diversifrons Bory =Trichomanes diversifrons 

Hymenostachys elegans (Rudge) K. Presl =Trichomanes 

diversifrons 

Hymenostachys osmundoides (DC. ex Poir.) K. Presl 

=Trichomanes osmundoides 

Lacostea ankersii (Parker ex Hook. & Grev.) Prantl 

=Trichomanes ankersii 

Lacostea tanaica (Hook. ex J.W. Sturm) Prantl =Trichomanes 

tanaicum 

Microgonium kapplerianum (J.W. Sturm) Pic.-Ser. 

=Trichomanes kapplerianum


Neuromanes abruptum Bosch, nom. superfl. =Trichomanes 

hostmannianum 

Neuromanes kaulfussii Bosch, nom. superfl. =Trichomanes 

pinnatum 

Neuromanes schomburgkianum (J.W. Sturm) Bosch 

=Trichomanes pinnatum 

Neuromanes vittaria (DC. ex Poir.) Bosch =Trichomanes 

vittaria 

Neurophyllum hostmannianum Klotzsch =Trichomanes 

hostmannianum 

Neurophyllum pennatum (Kaulf.) K. Presl =Trichomanes 

pinnatum 

Neurophyllum rhizophyllum (Cav.) Fée =Trichomanes 

pinnatum 

Neurophyllum vittaria (DC. ex Poir.) K. Presl =Trichomanes 

vittaria 

Odontomanes hostmannianum (Klotzsch) K. Presl 

=Trichomanes hostmannianum 

Ptilophyllum accedens (K. Presl) Prantl =Trichomanes 

accedens 

Ptilophyllum bancroftii (Hook. & Grev.) Prantl =Trichomanes 

bancroftii 

Ptilophyllum bicorne (Hook.) Prantl =Trichomanes bicorne 

Ptilophyllum botryoides (Kaulf.) Prantl =Trichomanes 

botryoides 

Ptilophyllum elegans (Rudge) Prantl =Trichomanes 

diversifrons 

Ptilophyllum hostmannianum (Klotzsch) Prantl =Trichomanes 

hostmannianum 

Ptilophyllum pinnatum (Kaulf.) Prantl =Trichomanes 

pinnatum 

Ptilophyllum schomburgkianum (J.W. Sturm) Prantl 


Ptilophyllum vittaria (DC. ex Poir.) Prantl =Trichomanes 

vittaria 

Ragatelus eriophorus K. Presl =Trichomanes crispum 

Sphaerocionium crispum (Kunth) Klotzsch var. amoenum 

Klotzsch =Hymenophyllum fendlerianum 

Trichomanes abruptum (Bosch) Christ =Trichomanes 

hostmannianum 

Trichomanes ankersii Parker ex Hook. & Grev. var. tanaicum 

(Hook. ex J.W. Sturm) Sadeb. =Trichomanes tanaicum 

Trichomanes commutatum J.W. Sturm =Trichomanes 

pedicellatum 

Trichomanes ekmanii W. Boer [records due to 


Trichomanes elegans Rudge, non Rich. =Trichomanes 

diversifrons 

Trichomanes fimbriatum Backhouse ex T. Moore [records due 

to misidentification] 

Trichomanes floribundum Humb. & Bonpl. ex Willd. var. 

vittaria (DC. ex Poir.) Splitg. =Trichomanes vittaria 

Trichomanes fruticulosum Jenm. =Trichomanes punctatum 

subsp. labiatum 

Trichomanes furcatum Bosch, non Kaulf. ex Spreng. 

=Trichomanes pedicellatum 

Trichomanes gemmatum J. Smith ex Hook. & Baker, nom. 

superfl. =Trichomanes cellulosum 

Trichomanes guianense J.W. Sturm =Trichomanes ankersii 

Trichomanes hookeri Auct. non K. Presl =Trichomanes 

kapplerianum 

Trichomanes huberi H. Christ =Trichomanes hostmannianum 

Trichomanes labiatum Jenm. =Trichomanes punctatum subsp. 

labiatum 

Trichomanes leprieurii Hook., nom. superfl. =Trichomanes 

elegans 

Trichomanes myrioneuron Lindm. =Trichomanes punctatum 

subsp. labiatum 

Trichomanes nanum (Bory) Hook. =Trichomanes botryoides 

Trichomanes pennatum Kaulf. =Trichomanes pinnatum 

Trichomanes pennatum Kaulf. =Trichomanes pinnatum 

Trichomanes pilosum Raddi =Trichomanes martiusii 

Trichomanes pinnatum Hedw. var. kaulfussii Domin 


Trichomanes pinnatum Hedw. var. vittaria (DC. ex Poir.) 

Baker =Trichomanes vittaria 

Trichomanes prieurii Kunze =Trichomanes elegans 

Trichomanes rhizophyllum Cav. =Trichomanes pinnatum 

Trichomanes schomburgkianum J.W. Sturm =Trichomanes 

pinnatum 

Trichomanes schomburgkii Bosch =Trichomanes crispum 

Trichomanes sphenoides Auct. non Kunze =Trichomanes 

punctatum 

Trichomanes spicisorum Desv. =Trichomanes osmundoides 

Trichomanes subsessile Splitg. =Trichomanes pedicellatum 

Trichomanes trigonum Auct. non Desv. =Trichomanes 

fimbriatum 

HYMENOPHYLLOPSIDACEAE 

contributed by D. Lellinger (1992); reviewed by 

D. Lellinger (1996) 

REFERENCES: 

Lellinger, D.B. 1984. Hymenophyllopsidaceae. 

In B. Maguire & collaborators, Botany of the 


Bot. Gard. 38: 1-84. 


In A.R.A. Görts-van Rijn, Flora of the 

Guianas (series B) 4: 82-86. 


In J.A. Steyermark, P.E. Berry, & B.K. Holst, 

Flora of the Venezuelan Guayana 2: 186-188. 

Hymenophyllopsis asplenioides A.C. Smith [GU] [Mt. 

Roraima] 

Hymenophyllopsis ctenitoides Lellinger [GU?] [Mt. Roraima] 

Hymenophyllopsis dejecta (Baker) Goebel [GU] [Mt. 

Roraima] 

LOMARIOPSIDACEAE 



REFERENCES: 

Hennipman, E. 1977. A monograph of the fern 

genus Bolbitis. Leiden Bot. Ser. 2: 1-331. 



PTEROPHYTA 

33

34 

PTEROPHYTA 



Guayana 2: 73-128. 

Bolbitis nicotianifolia (Swartz) Alston [GU SU FG] 

Bolbitis semipinnatifida (Fée) Alston [GU SU FG] 

Bolbitis serratifolia (Mertens ex Kaulf.) Schott [GU SU FG] 

Elaphoglossum antioquianum Hieron. [GU] 

Elaphoglossum apodum (Kaulf.) Schott ex J. Smith [GU SU 

FG] 

Elaphoglossum brachyneuron (Fée) J. Smith [GU] 

Elaphoglossum crinitum (L.) Christ [GU] 

Elaphoglossum cuspidatum (Willd.) T. Moore [GU] 

Elaphoglossum decoratum (Kunze) T. Moore [GU] 

Elaphoglossum engelii (H. Karst.) Christ [GU] 

Elaphoglossum eximium (Mett.) Christ [GU] 

Elaphoglossum flaccidum (Fée) T. Moore [GU SU FG] 

Elaphoglossum glabellum J. Smith [GU SU FG] 

Elaphoglossum herminieri (Bory & Fée ex Fée) T. Moore [GU 

SU FG] 

Elaphoglossum laminarioides (Bory ex Fée) T. Moore [GU SU 

FG] 

Elaphoglossum latifolium (Swartz) J. Smith [GU FG] [see 

also Elaphoglossum sporadolepis] 

Elaphoglossum lingua (Raddi) Brackenridge [GU FG] 

Elaphoglossum lloense (Hook.) T. Moore [SU FG?] 

Elaphoglossum longicaudatum Mickel [FG] 

Elaphoglossum longifolium (K. Presl) J. Smith [SU FG] 

Elaphoglossum luridum (Fée) Christ [GU SU FG] 

Elaphoglossum macrophyllum (Mett. ex Kuhn) Christ [GU 

FG] 

Elaphoglossum minutum (Pohl ex Fée) T. Moore [Mt. 

Roraima] 

Elaphoglossum moritzianum (Klotzsch) T. Moore [GU] 

Elaphoglossum nigrescens (Hook.) T. Moore [GU] 

Elaphoglossum peltatum (Swartz) Urb. [GU] 

Elaphoglossum piloselloides (K. Presl) T. Moore [SU FG] 

Elaphoglossum plumosum (Fée) T. Moore [GU SU FG] 

Elaphoglossum praetermissum Mickel [GU] 

Elaphoglossum pteropus C. Chr. [GU SU FG] 

Elaphoglossum raywaense (Jenm.) Alston [GU SU FG] 

Elaphoglossum revolutum (Liebm.) T. Moore [FG] 

Elaphoglossum scandens (Bory ex Fée) T. Moore [SU? FG?] 

Elaphoglossum sporadolepis (Kunze ex Kuhn) T. Moore ex C. 

Chr. [GU SU FG] [=Elaphoglossum latifolium?] 

Elaphoglossum strictum (Raddi) T. Moore [GU SU FG] 

Elaphoglossum styriacum Mickel [FG?] 

Elaphoglossum tenuifolium (Liebm.) T. Moore [FG] 

Elaphoglossum tovarense (Mett. ex Kuhn) T. Moore ex C. 

Chr. [Mt. Roraima] 

Elaphoglossum wurdackii Vareschi [Mt. Roraima] 

Lomagramma guianensis (Aubl.) Ching [GU SU FG] 

Lomariopsis erythrodes (Kunze) Fée [FG?] 

Lomariopsis japurensis (Mart.) J. Smith [GU SU FG] 

LOMARIOPSIDACEAE - SYNONYMS AND 


Bolbitis aliena (Swartz) Auct. non Alston =Bolbitis 

semipinnatifida 

Bolbitis maguirei Maxon & C. Morton =Bolbitis serratifolia 

Bolbitis scopulina Maxon & C. Morton =Bolbitis 


Elaphoglossum rigidum (Aubl.) Urb. =Elaphoglossum 

flaccidum 

Elaphoglossum schomburgkii (Fée) T. Moore 

=Elaphoglossum luridum 

Elaphoglossum spathulatum Auct. non (Bory) T. Moore 

=Elaphoglossum piloselloides 

Leptochilus alienus Auct. non (Swartz) C. Chr. =Bolbitis 


Leptochilus guianensis (Aubl.) C. Chr. =Lomagramma 

guianensis 

Leptochilus nicotianifolius (Swartz) C. Chr. =Bolbitis 

nicotianifolia 

Leptochilus serratifolius (Mertens ex Kaulf.) C. Chr. =Bolbitis 

serratifolia 

LYGODIACEAE 



REFERENCES: 

Duek, J.J. 1978. A taxonomic revision of 

Lygodium in America. Feddes Repert. 89: 

411-423. 

Lellinger, D.B. 1969. Schizaeaceae. In B. 


Guayana Highland - Part VIII. Mem. New 

York Bot. Gard. 18(2): 2-11. 

Smith, A.R. 1995. Schizaeaceae. In J.A. 



Lygodium flexuosum (L.) Swartz [GU] 

Lygodium micans J.W. Sturm [GU FG] [=Lygodium 

volubile?] 

Lygodium venustum Swartz [GU SU FG] 

Lygodium volubile Swartz [GU SU FG] 

LYGODIACEAE - SYNONYMS AND 


Lygodium polymorphum Auct. non (Cav.) Kunth =Lygodium 

venustum 

MARATTIACEAE 



REFERENCES: 

Smith, A.R. 1995. Marattiaceae. In J.A. 



Danaea elliptica J.E. Smith [GU? FG] 

Danaea leprieurii Kunze [GU SU FG] 

Danaea moritziana K. Presl [FG]


Danaea nodosa (L.) J.E. Smith [SU FG] 

Danaea simplicifolia Rudge [GU SU FG] 

Danaea trifoliata Rchb. ex Kunze [GU SU FG] 

MARSILEACEAE 



REFERENCES: 

Johnson, D.M. 1986. Systematics of the New 

World species of Marsilea. Syst. Bot. 

Monogr. 11: 1-87. 

Reed, C.F. 1965. Index Marsileata et Salviniata. 

Suppl. Bol. Soc. Brot. II, 39: 259-302. 

Smith, A.R. 1995. Marsileaceae. In J.A. 



Marsilea deflexa A. Braun [GU] 

Marsilea polycarpa Hook. & Grev. [GU SU FG] 

METAXYACEAE 



REFERENCES: 

Smith, A.R. 1995. Metaxyaceae. In J.A. 



Metaxya rostrata (Kunth) K. Presl [GU SU FG] 

METAXYACEAE - SYNONYMS AND 


Amphidesmium rostratum (Kunth) J. Smith =Metaxya rostrata 

OLEANDRACEAE 



REFERENCES: 


Nephrolepidaceae. In A.R.A. Görts-van Rijn, 



Oleandraceae. In A.R.A. Görts-van Rijn, 


Smith, A.R. 1995. Davalliaceae. In J.A. 






Guayana 2: 73-128. 

Nephrolepis biserrata (Swartz) Schott [GU SU FG] 

Nephrolepis cordifolia (L.) K. Presl [GU SU FG] 

Nephrolepis pectinata (Willd.) Schott [GU] 

Nephrolepis rivularis (Vahl) Mett. ex Krug [GU SU FG] 

Oleandra articulata (Swartz) K. Presl [GU SU FG] 

Oleandra pilosa Hook. [GU SU FG] 

OLEANDRACEAE - SYNONYMS AND 


Aspidium pendulum Splitg. =Oleandra pilosa 

Nephrolepis pendula (Raddi) J. Smith =Nephrolepis cordifolia 

Oleandra nodosa (Willd.) K. Presl =Oleandra articulata 

OPHIOGLOSSACEAE 

contributed by W. H. Wagner (1992); reviewed 

by D. Lellinger (1996) 

REFERENCES: 

Clausen, R.T. 1938. A monograph of the 

Ophioglossaceae. Mem. Torrey Bot. Club 

19(2): 1-177. 

Smith, A.R. 1995. Ophioglossaceae. In J.A. 



Cheiroglossa palmata (L.) K. Presl [GU SU] 

Ophioglossum nudicaule L.f. [GU SU FG] 

Ophioglossum reticulatum L. [GU SU FG?] 

OPHIOGLOSSACEAE - SYNONYMS AND 


Ophioglossum ellipticum Hook. & Grev. =Ophioglossum 

nudicaule 

Ophioglossum macrorhizum Kunze =Ophioglossum nudicaule 

Ophioglossum palmatum L. =Cheiroglossa palmata 

PARKERIACEAE 



REFERENCES: 

Lloyd, R.M. 1974. Systematics of the genus 

Ceratopteris Brongn. II. Taxonomy. Brittonia 

26: 139-160. 

Smith, A.R. 1995. Parkeriaceae. In J.A. 



PTEROPHYTA 

35

36 

PTEROPHYTA 


Ceratopteris deltoidea R.C. Benedict [GU SU] [=Ceratopteris 

pteridoides?] 

Ceratopteris pteridoides (Hook.) Hieron. [GU SU FG] 

Ceratopteris richardii Brongn. [GU SU FG] 

Ceratopteris thalictroides (L.) Brongn. [GU SU FG] 

PARKERIACEAE - SYNONYMS AND 


Ceratopteris calomelanos (L.) Link =Pityrogramma 

calomelanos (Pteridaceae) 

Ceratopteris lockhartii (Hook. & Grev.) Kunze =C. pteridoides 

or C. thalictroides 

POLYPODIACEAE 



REFERENCES: 

Lellinger, D.B. 1972. A revision of the fern 

genus Niphidium. Amer. Fern J. 62: 101-120. 

Lellinger, D.B. 1988. Some new species of 

Campyloneurum and a provisional key to the 

genus. Amer. Fern J. 78(1): 14-35. 

Price, M.G. 1983. Pecluma, a new tropical 

American fern genus. Amer. Fern J. 73(3): 

109-116. 

Smith, A.R. 1995. Polypodiaceae. In J.A. 



Campyloneurum amphostenon (Kunze ex Klotzsch) Fée [Mt. 

Roraima] 

Campyloneurum angustifolium (Swartz) Fée [GU SU FG] 

Campyloneurum costatum (Kunze) K. Presl [FG] 

Campyloneurum fuscosquamatum Lellinger [GU SU FG] 

Campyloneurum occultum (Christ) L.D. Gómez [FG] 

Campyloneurum phyllitidis (L.) K. Presl [GU SU FG] 

Campyloneurum repens (Aubl.) K. Presl [GU SU FG] 

Dicranoglossum desvauxii (Klotzsch) Proctor [GU SU FG] 

Dicranoglossum furcatum (L.) J. Smith [SU FG] 

Microgramma fuscopunctata (Hook.) Vareschi [GU SU FG] 

Microgramma geminata (Schrad.) R. & A. Tryon [GU] 

Microgramma lycopodioides (L.) Copeland [GU SU FG] 

Microgramma megalophylla (Desv.) de la Sota [GU SU?] 

Microgramma persicariifolia (Schrad.) K. Presl [GU SU FG] 

Microgramma piloselloides (L.) Copel. [GU FG] 

Microgramma reptans (Cav.) A.R. Smith [GU SU FG] 

Microgramma tecta (Kaulf.) Alston [GU SU FG] 

Microgramma thurnii (Baker) R. Tryon & Stolze [SU FG] 

Neurodium lanceolatum (L.) Fée [FG?] 

Niphidium crassifolium (L.) Lellinger [GU SU FG] 

Pecluma consimilis (Mett.) Price var. consimilis [GU] 

Pecluma eurybasis (C. Chr.) Price [Mt. Roraima] 

Pecluma hygrometrica (Splitg.) Price [GU SU] 

Pecluma pectinata (L.) Price [GU SU FG] 

Pecluma plumula (Humb. & Bonpl. ex Willd.) Price [GU SU 

FG] 

Pecluma ptilodon (Kunze) Price var. caespitosa (Jenm.) 

Mickel & Beitel [SU FG] 

Pecluma ptilodon (Kunze) Price var. pilosa (Evans) Stolze 

[GU SU FG] 

Pecluma ptilodon (Kunze) Price var. ptilodon [GU] 

Phlebodium aureum (L.) J. Smith [GU SU FG] 

Phlebodium decumanum (Willd.) J. Smith [GU SU FG] 

Phlebodium pseudoaureum (Cav.) Lellinger [GU SU FG] 

Pleopeltis astrolepis (Liebm.) Fourn. [GU SU FG] 

Pleopeltis percussa (Cav.) Hook. & Grev. [GU SU FG] 

[=Microgramma percussa] 

Polypodium adnatum Kunze ex Klotzsch [GU SU FG] 

Polypodium attenuatum Humb. & Bonpl. ex Willd. [GU SU 

FG] 

Polypodium bombycinum Maxon [GU] 

Polypodium caceresii Sodiro [GU SU FG] 

Polypodium dissimile L. [GU SU FG] 

Polypodium dulce Poir. [GU SU FG] 

Polypodium flagellare Christ [SU FG] 

Polypodium fraxinifolium Jacq. [GU] 

Polypodium laevigatum Cav. [GU SU FG] 

Polypodium lasiopus Klotzsch [SU FG] 

Polypodium loriceum L. [GU] 

Polypodium panorense C. Chr. [GU] 

Polypodium polypodioides (L.) Watt var. burchellii (Baker) 

Weath. [GU SU FG] 

Polypodium polypodioides (L.) Watt var. polypodioides [GU 

FG?] 

Polypodium pseudofraternum A.C. Smith [GU] 

Polypodium sessilifolium Desv. [GU] 

Polypodium triseriale Swartz [GU SU FG] 

POLYPODIACEAE - SYNONYMS AND 


Campyloneurum costatum (Kunze) K. Presl =Campyloneurum 

phyllitidis 

Campyloneurum sphenodes (Kunze) Fée [records due to 


Eschatogramme desvauxii (Klotzsch) C. Chr. 

=Dicranoglossum desvauxii 

Eschatogramme furcata (L.) C. Chr. =Dicranoglossum 

furcatum 

Eschatogramme furcata Auct. non (L.) C. Chr. 

=Dicranoglossum desvauxii 

Microgramma acuminata (Fée) Lellinger =Microgramma 

thurnii 

Microgramma ciliata (Willd.) Alston =Microgramma reptans 

Microgramma percussa (Cav.) de la Sota =Pleopeltis percussa 

Paltonium lanceolatum (L.) K. Presl =Neurodium lanceolatum 

Pleopeltis fuscopunctata (Hook.) R. & A. Tryon 

=Microgramma fuscopunctata 

Polypodium adiantoides Aubl., non Burm. =Polybotrya 

caudata (Dryopteridaceae) 

Polypodium angustifolium Swartz =Campyloneurum 

angustifolium 

Polypodium astrolepis Liebm. =Pleopeltis astrolepis 

Polypodium aureum L. =Phlebodium aureum 

Polypodium brasiliense Poir. =Polypodium triseriale 

Polypodium ciliatum Willd. =Microgramma reptans 

Polypodium consimile Mett. =Pecluma consimilis 

Polypodium crassifolium L. =Niphidium crassifolium


Polypodium crispatum (J. Smith) C. Morton =Enterosora 

ecostata (Grammitidaceae)[not in Guianas] 

Polypodium cultratum Auct. non Willd. =Grammitis 

mollissima (Grammitidaceae) 

Polypodium decumanum Willd. =Phlebodium decumanum 

Polypodium demeraranum Baker =Thelypteris demerarana 


Polypodium dicranophyllum C. Chr. =Grammitis furcata 

(Grammitidaceae) 

Polypodium duale Maxon =Cochlidium serrulatum 


Polypodium fuscopunctatum Hook. =Microgramma 

fuscopunctata 

Polypodium glaucophyllum Kunze =Polypodium laevigatum 

Polypodium hostmannii (Klotzsch) C. Morton =Thelypteris 

arborescens (Thelypteridaceae) 

Polypodium hygrometricum Splitg. =Pecluma hygrometrica 

Polypodium jubaeforme Kaulf. =Grammitis suspensa 


Polypodium kegelianum Kunze =Grammitis kegeliana 


Polypodium l'herminieri Fée =Grammitis taxifolia 


Polypodium lanceolatum Auct. non L. =Polypodium astrolepis 

Polypodium lycopodioides L. =Microgramma lycopodioides 

Polypodium megalophyllum Desv. =Microgramma 

megalophylla 

Polypodium mollissimum Fée =Grammitis mollissima 


Polypodium nanum Fée =Grammitis blanchetii 


Polypodium nervosum Klotzsch =Thelypteris rudis 


Polypodium pectinatum L. =Pecluma pectinata 

Polypodium percussum Cav. =Pleopeltis percussa 

Polypodium persicariifolium Schrad. =Microgramma 

persicariifolia 

Polypodium phyllitidis L. =Campyloneurum phyllitidis 

Polypodium plumula Humb. & Bonpl. ex Willd. =Pecluma 

plumula 

Polypodium plumulum Humb. & Bonpl. ex Willd. =Pecluma 

plumula 

Polypodium pseudoaureum Cav. =Phlebodium pseudoaureum 

Polypodium ptilodon Kunze =Pecluma ptilodon 

Polypodium repens Aubl. =Campyloneurum repens 

Polypodium roraimense Baker =Thelypteris roraimensis 


Polypodium roraimense Brause =Grammitis kalbreyeri 


Polypodium serrulatum (Swartz) Mett. =Cochlidium 

serrulatum (Grammitidaceae) 

Polypodium sororium Humb. & Bonpl. ex Willd. 

=Polypodium dulce 

Polypodium sphenodes Kunze =Campyloneurum sphenodes 

(Kunze) Fée [not in Guianas] 

Polypodium subobliquatum Hook. =Cyclodium inerme 

(Dryopteridaceae) 

Polypodium surinamensis Jacq. =Microgramma lycopodioides 

Polypodium taenifolium Jenm. =Grammitis taenifolia 


Polypodium taxifolium L. =Grammitis taxifolia 


Polypodium tectum Kaulf. =Microgramma tecta 

Polypodium thurnii Baker =Microgramma thurnii 

Polypodium trichomanoides Swartz =Grammitis 

trichomanoides (Grammitidaceae) 

PTERIDACEAE 



REFERENCES: 

Smith, A.R. & D.B. Lellinger. 1995. 

Pteridaceae. In J.A. Steyermark, P.E. Berry, 


Guayana 2: 250-286. 

Acrostichum aureum L. [GU SU FG] 

Acrostichum danaeifolium Langsd. & Fischer [GU SU FG] 

Pteris altissima Poir. [GU SU FG] 

Pteris biaurita L. [GU SU FG] 

Pteris deflexa Link [GU SU FG] 

Pteris denticulata Swartz [GU] 

Pteris plumula Desv. [FG] 

Pteris polita Link [SU FG] [see also Pteris propinqua] 

Pteris propinqua Agardh [SU FG] [=Pteris polita?] 

Pteris pungens Willd. [GU SU FG] 

Pteris quadriaurita Retz. [FG] [neotropical material =P. 

plumula] 

Pteris splendens Kaulf. [GU] 

Pteris tripartita Swartz [SU FG] [naturalized] 

Pteris vittata L. [SU] [naturalized] 

PTERIDACEAE - SYNONYMS AND 


Pteris kunzeana Agardh =Pteris altissima 

SALVINIACEAE 



REFERENCES: 

Reed, C.F. 1965a. Distribution of Salvinia and 

Azolla in South America and Africa in 

connection with studies for control by insects. 

Phytologia 12(3): 121-130. 

Reed, C.F. 1965b. Index Marsileata et 

Salviniata. Suppl. Bol. Soc. Brot. II, 39: 259- 

302. 

Smith, A.R. 1995. Azollaceae. In J.A. 



Smith, A.R. 1995. Salviniaceae. In J.A. 



Svenson, H.K. 1944. The New World species of 

Azolla. Amer. Fern J. 34: 69-84. 

PTEROPHYTA 

37

38 

Azolla caroliniana Willd. [GU SU FG] 

Azolla mexicana K. Presl [GU] 

Azolla microphylla Kaulf. [GU] 

Salvinia auriculata Aubl. [GU SU FG] 

PTEROPHYTA 


SCHIZAEACEAE 


reviewed by D. Lellinger & C.L. Kelloff 

(1996) 

REFERENCES: 

Lellinger, D.B. 1969. Schizaeaceae. In B. 



Bot. Gard. 18(2): 1-290. 

Smith, A.R. 1995. Schizaeaceae. In J.A. 



Actinostachys pennula (Swartz) Hook. [GU SU FG] 

Actinostachys subtrijuga (Mart.) K. Presl [GU SU FG] 

Anemia buniifolia (Gardner) T. Moore [SU] 

Anemia collina Raddi [GU] 

Anemia ferruginea Kunth var. ahenobarba (Christ) Mickel 

[SU] 

Anemia ferruginea Kunth var. ferruginea [GU SU?] 

Anemia hirta (L.) Swartz [GU SU] 

Anemia oblongifolia (Cav.) Swartz [GU SU] 

Anemia pastinacaria Moritz ex Prantl [SU FG] 

Anemia phyllitidis (L.) Swartz [GU SU] 

Anemia villosa Humb. & Bonpl. ex Willd. [GU SU FG] 

Schizaea elegans (Vahl) Swartz [GU SU FG] 

Schizaea fluminensis Miers ex J.W. Sturm [GU SU FG] 

Schizaea incurvata Schkuhr [GU SU FG] 

Schizaea poeppigiana J.W. Sturm [GU] 

Schizaea stricta Lellinger [GU] 

SCHIZAEACEAE - SYNONYMS AND 


Actinostachys penicillata (Humb. & Bonpl. ex Willd.) Maxon 

=Actinostachys pennula 

Schizaea flabellum Mart. =Schizaea elegans 

Schizaea pennula Swartz =Actinostachys pennula 

Schizaea subtrijuga Mart. =Actinostachys subtrijuga 

TECTARIACEAE 



REFERENCES: 




B) 6: 3-65. 

Holttum, R.E. 1986. Studies in the fern genera 

allied to Tectaria Cav. V. Triplophyllum, a 

new genus of Africa and America. Kew Bull. 

41: 237-260. 




Guayana 2: 73-128. 

Tindale, M.D. 1965. A monograph of the genus 

Lastreopsis Ching. Contr. New South Wales 

Nat. Herb. 3: 249-339. 

Ctenitis falciculata (Raddi) Ching [GU] 

Ctenitis paranaensis (C. Chr.) Lellinger [GU] 

Ctenitis refulgens (Klotzsch ex Mett.) C. Chr. ex Vareschi 

[GU SU FG] 

Cyclopeltis semicordata (Swartz) J.E. Smith [GU SU FG] 

Lastreopsis amplissima (K. Presl) Tindale [GU] 

Lastreopsis effusa (Swartz) Tindale subsp. divergens (Willd.) 

Tindale [GU SU FG] 

Lastreopsis killipii (C. Chr. & Maxon) Tindale [GU?] 

Tectaria heracleifolia (Willd.) Underw. [GU SU] 

Tectaria incisa Cav. [GU SU FG] 

Tectaria plantaginea (Jacq.) Maxon var. macrocarpa (Fée) C. 

Morton [GU SU FG] 

Tectaria trifoliata (L.) Cav. [GU SU FG] 

Tectaria trinitensis Maxon [GU SU] 

Triplophyllum acutilobum Holttum [GU?] 

Triplophyllum angustifolium Holttum [GU SU FG] 

Triplophyllum dicksonioides (Fée) Holttum [GU SU FG] 

Triplophyllum funestum (Kunze) Holttum [GU SU FG] 

TECTARIACEAE - SYNONYMS AND 


Aspidium martinicense Spreng. =Tectaria incisa 

Aspidium plantagineum (Jacq.) Griseb. =Tectaria plantaginea 

Aspidium plumieri K. Presl =Tectaria trifoliata 

Aspidium schomburgkii Klotzsch =Ctenitis falciculata 

Bathmium macrocarpum Fée =Tectaria plantaginea var. 

macrocarpa 

Bathmium sinuatum Fée =Tectaria plantaginea var. 

macrocarpa 

Ctenitis effusa (Swartz) Copeland =Lastreopsis effusa 

Ctenitis protensa (Afzel. ex Swartz) Ching var. funesta 

(Kunze) Proctor =Triplophyllum funestum 

Lastreopsis effusa (Swartz) Tindale subsp. effusa [not in 


Phegopteris refulgens Klotzsch ex Mett. =Ctenitis refulgens 

Tectaria plantaginea (Jacq.) Maxon var. plantaginea [not in 


Triplophyllum crassifolium Holttum =Triplophyllum funestum 

Triplophyllum crassifolium Holttum var. caudatum Holttum 

=Triplophyllum funestum 

Triplophyllum funestum (Kunze) Holttum var. hirstutum 

Holttum =Triplophyllum funestum 

THELYPTERIDACEAE




REFERENCES: 

Smith, A.R. 1994. Thelypteridaceae. In A.R.A. 


B) 6: 77-115. 

Smith, A.R. 1995. Thelypteridaceae. In J.A. 



Thelypteris abrupta (Desv.) Proctor [GU SU FG] 

Thelypteris arborea (Brause) A.R. Smith [GU] [Mt. Roraima] 

Thelypteris arborescens (Humb. & Bonpl. ex Willd.) C. 

Morton [GU SU FG] 

Thelypteris balbisii (Spreng.) Ching [GU] 

Thelypteris brachypoda (Baker) C. Morton [GU] [Mt. 

Roraima] 

Thelypteris chrysodioides (Fée) C. Morton [GU] 

Thelypteris decussata (L.) Proctor var. decussata [GU SU? 

FG] 

Thelypteris demerarana (Baker) C. Reed [GU?] [Mt. Roraima] 

Thelypteris funckii (Mett.) Alston [GU] [Mt. Roraima] 

Thelypteris gemmulifera (Hieron.) A.R. Smith [FG] 

Thelypteris germaniana (Fée) Proctor [FG] 

Thelypteris glandulosa (Desv.) Proctor var. glandulosa [GU 

SU FG] 

Thelypteris gonophora (Weath.) A.R. Smith [GU] 

Thelypteris hispidula (Decne.) C. Reed var. hispidula [GU SU 

FG] 

Thelypteris holodictya K.U. Kramer [SU FG] 

Thelypteris interrupta (Willd.) Iwatsuki [GU SU FG] 

Thelypteris juruensis (C. Chr.) R. Tryon & Conant [FG] 

Thelypteris leprieurii (Hook.) R. Tryon var. leprieurii [GU SU 

FG] 

Thelypteris linkiana (K. Presl) R. Tryon [GU] [Mt. Roraima] 

Thelypteris macrophylla (Kunze) C. Morton [GU SU FG] 

Thelypteris nephrodioides (Klotzsch) Proctor [GU FG] 

Thelypteris nesiotica (Maxon & C. Morton) C. Morton [GU] 

Thelypteris opulenta (Kaulf.) Fosberg [GU SU FG?] 

[naturalized] 

Thelypteris pennata (Poir.) C. Morton [GU SU FG] 

Thelypteris peradenia A.R. Smith [Mt. Roraima] 

Thelypteris pilossisima C. Morton [Mt. Roraima] 

Thelypteris poiteana (Bory) Proctor [GU SU FG] 

Thelypteris pteroidea (Klotzsch) R. Tryon [Mt. Roraima] 

Thelypteris roraimensis (Baker) C. Reed [GU] [Mt. Roraima] 

Thelypteris rudis (Kunze) Proctor [GU] 

Thelypteris schomburgkii A.R. Smith [GU] 

Thelypteris serrata (Cav.) Alston [GU SU FG] 

Thelypteris tetragona (Swartz) Small [GU SU FG] 

Thelypteris tristis (Kunze) R. Tryon [GU SU FG] 

THELYPTERIDACEAE - SYNONYMS AND 


Aspidium gongylodes Schkuhr =Thelypteris interrupta 

Cyclosorus gongylodes (Schkuhr) Link =Thelypteris 

interrupta 

Goniopteris juruensis (C. Chr.) Brade =Thelypteris juruensis 

Goniopteris poiteana (Bory) Ching =Thelypteris poiteana 

Lastrea brachypoda (Baker) Vareschi =Thelypteris brachypoda 

Lastrea demerarana (Baker) Vareschi =Thelypteris demerarana 

Lastrea poiteana Bory =Thelypteris poiteana 

Lastrea roraimensis (Baker) Brade =Thelypteris roraimensis 

Meniscium guyanense Fée =Thelypteris macrophylla 

Nephrodium aureoviridum Jenm. [dubious species; 

=Thelypteris tetragona?] 

Nephrodium brachypodum Baker =Thelypteris brachypoda 

Nephrodium connexum Kuhn ex Baker =Thelypteris connexa 

[type erroneously cited from French Guiana] 

Nephrodium dejectum Jenm. =Thelypteris glandulosa 

Nephrodium leprieurii Hook. =Thelypteris leprieurii 

Nephrodium quandrangulare Fée =Thelypteris hispidula 

Nephrodium subcuneatum Baker =Thelypteris abrupta 

Nephrodium subfuscum Baker =Thelypteris germaniana 

Steiropteris glandulosa (Desv.) Pic.-Ser. =Thelypteris 

glandulosa 

Steiropteris leprieurii (Hook.) Pic.-Ser. =Thelypteris leprieurii 

Thelypteris connexa (Kuhn ex Baker) C. Reed [type 

erroneously cited from French Guiana] 

Thelypteris gongylodes (Schkuhr) Small =Thelypteris 

interrupta 

Thelypteris guyanensis (Fée) C. Morton =Thelypteris 

macrophylla 

Thelypteris hostmannii (Klotzsch) C. Morton =Thelypteris 

arborescens 

Thelypteris nervosa (Klotzsch) R. Tryon =Thelypteris rudis 

Thelypteris quadrangularis (Fée) Schelpe =Thelypteris 

hispidula 

VITTARIACEAE 



PTEROPHYTA 

39

40 

REFERENCES: 


Lellinger, D.B. 1972. Polypodiaceae subfam. 

Vittarioideae. In B. Maguire & colaborators, 



Smith, A.R. 1995. Vittariaceae. In J.A. 



Anetium citrifolium (L.) Splitg. [GU SU FG] 

Antrophyum cajenense (Desv.) Spreng. [GU SU FG] 

Antrophyum guayanense Hieron. [GU SU FG] 

Antrophyum jenmanii Benedict [GU SU FG] 

Antrophyum lanceolatum (L.) Kaulf. [GU SU FG] 

Hecistopteris pumila (Spreng.) J.E. Smith [GU SU FG] 

Vittaria costata Kunze [GU SU FG] 

Vittaria gardneriana Fée [GU SU] 

Vittaria graminifolia Kaulf. [GU SU] 

Vittaria lineata (L.) J.E. Smith [GU SU FG] 

Vittaria moritziana Mett. [GU] 

Vittaria remota Fée [GU SU FG] 

Vittaria stipitata Kunze [FG] 

VITTARIACEAE - SYNONYMS AND 


Ananthacorus angustifolius (Swartz) Underw. & Maxon 

=Vittaria costata 

Polytaenium brasilianum (Desv.) Benedict [records due to 


Polytaenium cajenense (Desv.) Benedict =Antrophyum 

cajenense 

Polytaenium guayanense (Hieron.) Alston =Antrophyum 

guayanense 

Vittaria angustifolia (Swartz) Baker =Vittaria costata 

Vittaria filifolia Fée =Vittaria graminifolia 

WOODSIACEAE 



REFERENCES: 




B) 6: 3-65. 




Guayana 2: 73-128. 

Diplazium celtidifolium Kunze [GU SU FG] 

Diplazium centripetale (Baker) Maxon [GU] [Mt. Roraima] 

Diplazium cristatum (Desr.) Alston [GU SU FG] 

Diplazium expansum Willd. [GU FG] 

PTEROPHYTA, SPERMATOPHYTA 

Diplazium gracilescens (Mett.) C. Chr. [SU FG] 

Diplazium grandifolium (Swartz) Swartz [GU FG] 

Diplazium lechleri (Mett.) T. Moore [GU SU] 

Diplazium radicans Desv. [SU FG] 

Diplazium roraimense Cremers & K.U. Kramer [GU] [Mt. 

Roraima] 

Diplazium striatum (L.) K. Presl [GU FG] 

Hemidictyum marginatum (L.) K. Presl [SU FG] 

WOODSIACEAE - SYNONYMS AND 


Diplazium arboreum (Willd.) K. Presl =Diplazium cristatum 

Diplazium shepherdii (Spreng.) Link =Diplazium cristatum 

SPERMATOPHYTA 

SPERMATOPHYTA REFERENCES: 

Cremers, G. 1985. Espèces nouvelles de Guyane 

française: richesse de l'herbier de Cayenne 

(CAY). Proc. Kon. Ned. Akad. Wet. ser. C, 

88(1): 15-37. 



Phanérogames et de Ptéridophytes de Guyane 

française d'après l'Herbier du Centre 


Cremers, G., J.J. de Granville, & M. Hoff. 1994. 

Les plantes "endemiques" de Guyane 

française. Mus. Nat. Hist. Nat., Coll. Patr. 

Nat. (Sér. Patr. Génétique) 18: 1-50. 



française. IV - Les Monocotyledones. Mus. 

Nat. Hist. Nat., Coll. Patr. Nat. (Sér. Patr. 

Génétique) 17: 1-252. 

Fanshawe, D.B. 1952. The vegetation of British 

Guiana: A preliminary review. Imperial 

Forestry Institute paper no. 29: 1-96. 

Funk, V.A. & C.L. Kelloff. 1991. A Preliminary 

Checklist of the Flowering Plants and Ferns of 

Kaieteur Falls National Park Guyana. 

Unpublished manuscript. Biological 

Diversity of the Guianas program, 

Smithsonian Institution, Washington, DC. 

Gleason, H.A. 1929. Studies on the flora of 

northern South America - XIII. The Tate 

collection from Mount Roraima and vicinity. 

Bull. Torr. Bot. Club 56(8): 391-408. 

Graham, E.H. 1934. Flora of the Kartabo 

Region, British Guiana. Reprinted from Ann. 

Carn. Mus. 22(1) (1933). 

Howard, R.A. 1983. The plates of Aublet's 

Histoire des Plantes de la Guiane Françoise. 

J. Arn. Arb. 64: 255-292. 

Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1989. 

Botanical explorations in Guyana, 1985. 

Annex: identifications. Institute of Systematic 

Botany, Utrecht.



Botanical explorations in the northern part of 

the western Kanuku Mountains (Guyana). 

Annex: identifications. Institute of Systematic 

Botany, Utrecht. 


Botanical exploration in the Wai-Wai area of 

southern Guyana. Annex: identifications. 

Herbarium Division, Department of Plant 

Ecology and Evolutionary Biology, 

University of Utrecht. 

Lemée, A. 1953-1956. Flore de la Guyane 

Française. Lechevalier, Paris. 

Lindeman, J.C. 1953. The Vegetation of the 

Coastal Region of Suriname. Drukkerij en 

Uitg.-Mij. v/h Kemink, Utrecht. 

Maas, P.J.M. & L.Y.Th. Westra. 1990. 

Botanical exploration in the northern part of 

the Rupununi Savanna and in the Mabura Hill 

area (Guyana). Appendix: identifications. 

Institute of Systematic Botany, Utrecht. 

Maguire, B. & collaborators. 1948. Plant 

explorations in Guiana in 1944, chiefly to the 

Tafelberg and the Kaieteur Plateau. Bull. 

Torrey Bot. Club 75: 56-115, 189-230, 286- 

323, 374-438, 523-580, 633-671. 

Mennega, E.A., W.C.M. Tammens-de Rooij, & 

M.J. Jansen-Jacobs. 1988. Check-List of 

Woody Plants of Guyana. The Tropenbos 

Foundation, Ede, The Netherlands. 

Mori, S.A. & collaborators. 1987. The 

Lecythidaceae of a lowland neotropical forest: 

La Fumée Mountain, French Guiana. Mem. 

New York Bot. Gard. 44: 1-190. 

Omawale. 1973. Guyana's Edible Plants. 

University of Guyana. 

Plotkin, M.J., B.M. Boom, & M. Allison. 1991. 

The ethnobotany of Aublet's Histoire des 

Plantes de la Guiane Françoise (1775). 

Monogr. Syst. Bot. 35: 1-108. 

Polak, A.M. 1992. Major Timber Trees of 

Guyana, a Field Guide. The Tropenbos 

Foundation, Wageningen. 

Pulle, A. 1906. An Enumeration of the Vascular 

Plants Known from Surinam, Together with 

their Distribution and Synonymy. E.J. Brill, 

Ltd., London. 

Roosmalen, M.G.M. van. 1985. Fruits of the 

Guianan Flora. Institute of Systematic 

Botany, Utrecht University, Utrecht, and 

Silvicultural Department of Wageningen 

Agricultural University, Wageningen. 

Steege, H. ter, M.J. Jansen-Jacobs, & M. Polak. 

1993. Checklist of plants collected around 

Mabura Hill, Guyana. The Tropenbos 

Foundation, Wageningen and Department of 

Plant Ecology and Evolutionary Biology, 

Utrecht University. 

Steyermark, J.A., P.E. Berry, & B.K. Holst. 

1995. Flora of the Venezuelan Guayana. 

Missouri Botanical Garden, St. Louis, & 

Timber Press, Portland, Oregon. 

ACANTHACEAE 

reviewed by D. Wasshausen (1992); reviewed by 

D. Wasshausen (1996) 

REFERENCES: 

Baum, V.M. & J.L. Reveal. 1983. Pulchranthus, 

a new genus from northern South America. 

Syst. Bot. 8(2): 221-220. 

Bremekamp, C.E.B. 1938. Acanthaceae. In A. 

Pulle, Flora of Suriname 4(2): 166-252. 

Bremekamp, C.E.B. 1938. Notes on the 

Acanthaceae of Surinam. Rec. Trav. Bot. 

Neerl. 35: 130-171. 

Graham, V.A.W. 1988. Delimitation and infrageneric 

classification of Justicia. Kew Bull. 

43(4): 551-624. 

Leonard, E.C. 1961. Acanthaceae. In B. 

Maguire, J.J. Wurdack, & collaborators, 

Botany of the Guayana Highland part IV. 

Mem. New York Bot. Gard. 10(4): 47-58. 

Wasshausen, D.C. 1975. The genus Aphelandra. 

Smiths. Contrib. Bot. 18: 1-157. 

Wasshausen, D.C. 1990. Justicia maguirei, a 

new species of Acanthaceae from Guyana. 


Wasshausen, D.C. 1991. New species of 

Dicliptera from the Guianas. Brittonia 43(1): 

1-6. 

Wasshausen, D.C. 1992a. New species and new 

combinations of Justicia (Acanthaceae) from 

the Venezuelan Guayana. Novon 2: 62-80. 

Wasshausen, D.C. 1992b. Nomenclatural 

changes in the Acanthaceae and two new 

species of Odontonema from Venezuela and 

Guyana. Novon 2: 149-152. 

Wasshausen, D.C. 1995. Acanthaceae. In J.A. 



Wasshausen, D.C. 1995. A new species, new 

combinations, and a new name in 

Acanthaceae from northern South America. 

Brittonia 47(4): 422-425. 

Anisacanthus secundus Leonard [GU SU FG] 

Aphelandra aurantiaca (Schweidw.) Lindl. var. aurantiaca 

[GU SU FG] 

Aphelandra pulcherrima (Jacq.) Kunth [GU SU] 

Aphelandra scabra (Vahl) Sm. [GU SU] 

Asystasia gangetica (L.) T. Anderson [GU] [cultivated, 

persists as an escape] 

Barleria lupulina Lindl. [FG] [introduced, possibly 

naturalized] 

Blechum pyramidatum (Lam.) Urb. [GU SU FG] 

Dicliptera ciliaris Juss. [GU? SU FG] 

Dicliptera granvillei Wasshausen [FG] 

Dicliptera knappii Wasshausen [GU] 

Dicliptera sexangularis (L.) Juss. [SU FG] [to be expected in 

GU] 

Eranthemum pulchellum Andr. [SU] [cultivated, naturalized] 

Fittonia sp. [GU] 

SPERMATOPHYTA: Acanthaceae 

41

42 


Gynocraterium guianense Bremek. [GU SU] 

Hemigraphis alternata (Burm.f.) T. Anderson [cultivated, 

persists as an escape] 

Herpetacanthus rotundatus (Lindau) Bremek. [SU FG] 

Hygrophila costata Nees [GU SU FG] 

Hygrophila erecta (Burm.f.) Hochr. [SU] [introduced?] 

Hygrophila sandwithii Bremek. [GU] 

Hygrophila surinamensis Bremek. [SU] [to be expected in 

GU] 

Justicia betonica L. [GU] [introduced?] 

Justicia calycina (Nees) V.A.W. Graham [GU SU FG] 

Justicia carthaginensis Jacq. [GU SU FG] 

Justicia cayennensis (Nees) Lindau var. cayennensis [GU SU 

FG] 

Justicia cayennensis (Nees) Lindau var. parvifolia Bremek. 

[SU] 

Justicia comata (L.) Lam. [GU SU FG] 

Justicia coppenamensis Wasshausen [SU] [nom. nov. for 

Beloperone modesta Bremek.] 

Justicia guianensis (N.E. Brown) Wasshausen [GU] 

Justicia laevilinguis (Nees) Lindau [SU FG] [to be expected in 

GU] 

Justicia maguirei Wasshausen [GU] 

Justicia martiana (Nees) Lindau [GU SU] 

Justicia monachinoi Wasshausen [GU] 

Justicia pectoralis Jacq. [GU SU FG] 

Justicia polystachya Lam. [SU FG] [to be expected in GU] 

Justicia potarensis (Bremek.) Wasshausen [GU SU] 

Justicia schomburgkiana (Nees) V.A.W. Graham [GU] 

Justicia secunda Vahl [GU SU FG] 

Justicia sprucei V.A.W. Graham [FG] 

Lepidagathis alopecuroidea (Vahl) R. Brown ex Griseb. [GU 

SU FG] 

Lepidagathis gracilis (Bremek.) Wasshausen [SU] [to be 

expected in GU] 

Lepidagathis medicaginea (Bremek.) Wasshausen [SU] [to be 


Lepidagathis nickeriensis Wasshausen [SU] [to be expected in 

GU] 

Lepidagathis surinamensis (Bremek.) Wasshausen [GU SU 

FG] 

Megaskepasma erythrochlamys Lindau [SU] 

Nelsonia campestris R. Brown [FG?] 

Odontonema mazarunensis Wasshausen [GU] 

Odontonema schomburgkianum (Nees) Kuntze [GU] 

Pachystachys coccinea (Aubl.) Nees [FG] 

Pachystachys spicata (Ruiz & Pav.) Wasshausen [GU FG] 

Polylychnis fulgens Bremek. [SU FG] 

Polylychnis radicans (Nees) Wasshausen, comb. nov. ined. 

[GU FG] 

Pulchranthus surinamensis (Bremek.) Baum, Reveal & 

Nowicke [SU] 

Pulchranthus variegatus (Aubl.) Baum, Reveal & Nowicke 

[SU FG] 

Ruellia geminiflora Kunth var. angustifolia (Nees) Griseb. 

[GU SU FG] 

Ruellia geminiflora Kunth var. geminiflora [GU SU FG] 

Ruellia inflata L.C. Rich. [SU FG] 

Ruellia longifolia L.C. Rich. [SU FG] 

Ruellia microcalyx (Nees) Lindau [GU SU] 

Ruellia prostrata Poir. [GU] 

Ruellia rubra Aubl. [GU SU FG] [includes several varieties 

without combinations in Ruellia] 

Ruellia saülensis Wasshausen [FG] 

SPERMATOPHYTA: Acanthaceae 

Ruellia tuberosa L. [GU SU FG] [naturalized] 

Ruellia violacea Aubl. [FG] 

Staurogyne fockeana Bremek. [SU] 

Staurogyne miqueliana Kuntze [SU] 

Staurogyne repens (Nees) Kuntze [GU] 

Staurogyne spraguei Wasshausen [GU] 

Staurogyne stahelii Bremek. [SU] 

Staurogyne stolonifera (Nees) Kuntze [SU] 

Staurogyne trinitensis Leonard [GU SU FG] 

Staurogyne wullschlaegeliana Bremek. [SU] 

Teliostachya cataractae Nees [GU FG] [=Lepidagathis sp.] 

Teliostachya coppenamensis Bremek. [SU] [=Lepidagathis 

sp.] 

Teliostachya lanceolata Nees [SU] [=Lepidagathis sp.] 

Teliostachya mazarunensis Bremek. [GU] [=Lepidagathis sp.] 

Thunbergia alata Bojer ex Sims [GU SU] [cultivated, 

naturalized] 

Thunbergia erecta (Benth.) T. Anderson [SU] [cultivated, 

naturalized] 

Thunbergia fragrans Roxb. [SU] [cultivated, naturalized] 

Thunbergia grandiflora (Roxb. ex Rottl.) Roxb. [SU] 

[cultivated, naturalized] 

Trichanthera gigantea (Humb. & Bonpl.) Nees var. guianensis 

Gleason [GU SU] 

ACANTHACEAE - SYNONYMS AND 


Anisacanthus boliviensis (Nees) Wasshausen [not in Guianas; 

records referable to A. secundus] 

Aphelandra deppeana Schlechtend. & Cham. =Aphelandra 

scabra 

Aphelandra paraensis Lindau =Aphelandra aurantiaca 

Aphelandra pectinata Willd. ex Nees =Aphelandra scabra 

Aphelandra tetragona (Vahl) Nees [not in Guianas] 

Arrhostoxylum coccineum Nees [not in Guianas] 

Arrhostoxylum rubrum (Aubl.) Nees =Ruellia rubra 

Asystasia nemorum Nees =Asystasia gangetica 

Beloperone modesta Bremek. =Justicia coppenamensis 

Beloperone schomburgkiana Nees =Justicia schomburgkiana 

Blechum brownei Juss. =Blechum pyramidatum 

Chaetochlamys ciliata Lindau =Justicia sprucei 

Dactylostegium assurgens (L.) Bremek. =Dicliptera 

sexangularis 

Dianthera spp. =Justicia spp. 

Dicliptera assurgens (L.) Juss. =Dicliptera sexangularis 

Dicliptera vahliana Nees =Dicliptera sexangularis 

Dipteracanthus angustifolius Bremek. =Ruellia geminiflora 

var. angustifolia 

Dipteracanthus paniculatus Nees =Ruellia sp.?[not in Guianas] 

Dipteracanthus surinamensis Miq. =Ruellia microcalyx 

Dipteracanthus violaceus (Nees) Lemée =Ruellia violacea 

Drejera boliviensis Nees =Anisacanthus boliviensis [not in 


Duvernoia potarensis Bremek. =Justicia potarensis 

Duvernoia potarensis Bremek. var. tetrapora Bremek. 

=Justicia potarensis 

Dyspemptemorion cayennense (Nees) Bremek. =Justicia 

cayennensis 

Ebermaiera repens Nees =Staurogyne repens 

Ebermaiera spp. =Staurogyne spp. 

Eranthemum nervosum (Vahl) R. Brown =Eranthemum 

pulchellum


Hygrophila guianensis Nees ex Benth. =Hygrophila costata 

Juruasia rotundata Lindau =Herpetacanthus rotundatus 

Justicia "divergens (Nees) Lindau" [invalid name, records 

based on misidentifications] 

Justicia acuminatissima (Miq.) Bremek. =Justicia calycina 

Justicia coccinea Aubl. =Pachystachys coccinea 

Justicia glabribracteata (Lindau) V.A.W. Graham [records 

apparently in error] 

Justicia moricandiana (Nees) Lemée =Justicia secunda 

Justicia obtusifolia (Nees) Lindau =Justicia laevilinguis 

Justicia pectoralis Jacq. var. latifolia Bremek. =Justicia 

pectoralis 

Justicia variegata Aubl. =Pulchranthus variegatus 

Nelsonia canescens (Lam.) Spreng. =Nelsonia campestris 

Odontonema surinamense Bremek. =Pulchranthus 

surinamensis 

Odontonema variegatum (Aubl.) Kuntze =Pulchranthus 

variegatus 

Pachystachys riedeliana Nees =Pachystachys spicata 

Polylychnis essequibensis Bremek. =Polylychnis radicans 

Psadacocalymma spp. =Justicia spp. 

Rhacodiscus calycinus (Nees) Bremek. =Justicia calycina 

Rhacodiscus secundus (Vahl) Bremek. =Justicia secunda 

Ruellia cordifolia Lindau =Ruellia inflata 

Ruellia pipericaulis Rizzini =Ruellia inflata 

Ruellia radicans (Nees) Lindau =Polylychnis radicans 

Ruellia tubiflora Kunth [not in Guianas, based on 


Staurogyne linearifolia Bremek. =Staurogyne stolonifera 

Staurogyne versteegii Bremek. =Staurogyne trinitensis 

Teliostachya alopecuroidea (Vahl) Nees =Lepidagathis 

alopecuroidea 

Teliostachya glandulosa Bremek. =Lepidagathis nickeriensis 

Teliostachya gracilis Bremek. =Lepidagathis gracilis 

Teliostachya medicaginea Bremek. =Lepidagathis 

medicaginea 

Teliostachya surinamensis Bremek. =Lepidagathis 

surinamensis 

AGAVACEAE 

Furcraea foetida (L.) Haw. [GU SU FG] 

AGAVACEAE - SYNONYMS AND 


Furcraea gigantea Vent. =Furcraea foetida 

AIZOACEAE 

reviewed by R.A. DeFilipps (1996) 

REFERENCES: 

Burger, W. 1983. Aizoaceae. In W. Burger, 

Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 

199-212. 

Eyma, P.J. 1934. Aizoaceae. In A. Pulle, Flora 

of Suriname 1(1): 158-161. 

Nevling, L Jr. 1961. Aizoaceae. In R.E. 

Woodson Jr., R.W. Schery, & collaborators, 

Flora of Panama. Ann. Missouri Bot. Gard. 

48(1): 80-85. 

Sesuvium portulacastrum (L.) L. [GU SU FG] 

AIZOACEAE - SYNONYMS AND 


Sesuvium acutifolium Miq. =Sesuvium portulacastrum 

ALISMATACEAE 

contributed by R. Haynes & L.B. Holm-Nielsen 

(1992) 

REFERENCES: 

Bogin, C. 1955. Revision of the genus 

Sagittaria. Mem. New York Bot. Gard. 9: 

179-233. 

Fassett, N.C. 1955. Echinodorus in the 

American tropics. Rhodora 57: 133-212. 

Haynes, R.R. & L.B. Holm-Nielsen. 1986. 

Notes on Echinodorus. Brittonia 38: 325-332. 


Alismataceae. Flora Neotropica Monograph 

64: 1-112. 

Jonker, F.P. 1943. Alismataceae. In A. Pulle, 


Jonker, F.P. 1968. Alismataceae (additions and 

corrections). In A. Pulle & J. Lanjouw, Flora 


Rataj, K. 1972. Revision of the genus Sagittaria. 

Part II. Annot. Zool. Bot. Bratislava 78: 1-61. 

Rataj, K. 1975. Revizion [sic] of the genus 

Echinodorus Rich. Studie CSAV (Praha) 2: 

1-156. 

Echinodorus berteroi (Spreng.) Fassett [GU] 

Echinodorus bolivianus (Rusby) Holm-Nielsen [GU] 

Echinodorus grisebachii Small [SU FG] 

Echinodorus macrophyllus (Kunth) Mich. subsp. macrophyllus 

[GU] 

Echinodorus macrophyllus (Kunth) Mich. subsp. scaber 

(Rataj) Haynes & Holms-Nielsen [GU SU FG?] 

Echinodorus paniculatus Mich. [GU SU?] 

Echinodorus reticulatus Haynes & Holm-Nielsen [SU] 

Echinodorus subalatus (Mart.) Griseb. subsp. andrieuxii 

(Hook. & Arnott) Haynes & Holm-Nielsen [GU] 

Echinodorus subalatus (Mart.) Griseb. subsp. subalatus [GU 

SU?] 

Echinodorus tenellus (Mart.) Buch. [GU SU?] 

Sagittaria guayanensis Kunth subsp. guayanensis [GU SU 

FG] 

Sagittaria lancifolia L. subsp. lancifolia [GU SU FG] 

Sagittaria rhombifolia Cham. [GU SU FG?] 

SPERMATOPHYTA: Acanthaceae - Alismataceae 

43

44 


ALISMATACEAE - SYNONYMS AND 


Alisma tenellum Mart. =Echinodorus tenellus 

Alisma tenellum Mart. f. latifolia Seub. =Echinodorus 

bolivianus 

Echinodorus andrieuxii (Hook. & Arnott) Small =Echinodorus 

subalatus subsp. andrieuxii 

Echinodorus gracilis Rataj =Echinodorus grisebachii 

Echinodorus grandiflorus (Cham. & Schlechtend.) Mich. [not 


Echinodorus guianensis (Kunth) Griseb. =Sagittaria 

guayanensis 

Echinodorus intermedius Griseb. [not in Guianas] 

Echinodorus latifolius (Seub.) Rataj =Echinodorus bolivianus 

Echinodorus scaber Rataj =Echinodorus macrophyllus subsp. 

scaber 

Lophotocarpus guayanensis (Kunth) J.G. Smith =Sagittaria 

guayanensis 

Sagittaria pugioniformis L. =Sagittaria lancifolia 

AMARANTHACEAE 


REFERENCES: 

Burger, W. 1983. Amaranthaceae. In W. 

Burger, Flora Costaricensis. Fieldiana Bot. 

(n.s.) 13: 142-180. 

Duke, J.A. 1961. Amaranthaceae. In R.E. 

Woodson, R.W. Schery, & collaborators, 

Flora of Panama 4(4): 348-392. 

Kellogg, E.A. 1988. Amaranthaceae. In R.A. 

Howard, Flora of the Lesser Antilles. 

Mears, J.A. 1977. The nomenclature and type 

collections of the widespread taxa of 

Alternanthera. Proc. Acad. Nat. Sci. Phila. 

129: 1-21. 

Morros, M.E., B. Trujillo, & M. Ponce. 1990. 

Descripcion del genero Amaranthus L. con 3 

nuevos registros para Venezuela y 

consiguiente clave para las especies. Ernstia 

58-59-60: 45-51. 

Nee, M.H. 1995. Amaranthaceae. In J.A. 



Pedersen, T.M. 1967. Studies in South 

American Amaranthaceae. Darwiniana 14(2- 

3): 430-462. 

Pedersen, T.M. 1976. Estudios sobre 

Amaranthaceae Sudamericanas, II. 

Darwiniana 20(1-2): 269-303. 

Scheygrond, A. 1932. Amarantaceae. In A. 


Sohmer, S.H. 1977. Revision of Chamissoa. 

Bull. Torr. Bot. Club 104: 111-126. 

Achyranthes aspera L. [GU SU FG] [naturalized] 

Alternanthera brasiliana (L.) Kuntze [GU SU FG] 

SPERMATOPHYTA: Alismataceae, Amaranthaceae 

Alternanthera halimifolia (Lam.) Standl. [GU SU FG] 

[=Alternanthera tenella?] 

Alternanthera littoralis P. Beauv. var. maritima (Mart.) 

Pedersen [FG?] 

Alternanthera paronychioides A. St. Hil. [GU SU FG] 

Alternanthera philoxeroides (Mart.) Griseb. [GU SU FG] 

Alternanthera sessilis (L.) R. Br. ex DC. [GU SU FG] 

Alternanthera tenella Colla [GU SU FG] 

Amaranthus australis (A. Gray) J.D. Sauer [SU] [introduced?] 

Amaranthus blitum L. [GU SU FG] [introduced?] 

Amaranthus caudatus L. [SU FG] [cultivated or escaped from 

cultivation] 

Amaranthus dubius Mart. ex Thellung [GU SU FG] 

Amaranthus hybridus L. [cultivated only?] 

Amaranthus spinosus L. [GU SU FG] [introduced?] 

Amaranthus tricolor L. [SU FG] [cultivated only?] 

Amaranthus viridis L. [GU SU FG] [introduced?] 

Blutaparon vermiculare (L.) Mears [GU SU FG] 

Celosia argentea L. [SU FG] [cultivated, naturalized] 

Chamissoa altissima (Jacq.) Kunth var. altissima [GU SU] 

Cyathula achyranthoides (Kunth) Moq. [GU SU FG] 

[introduced?] 

Cyathula prostrata (L.) Blume [GU SU FG] [introduced?] 

Gomphrena globosa L. [GU SU FG] [widely cultivated and 

escaping] 

Iresine diffusa Humb. & Bonpl. ex Willd. [GU SU] 

Pfaffia glauca (Mart.) Spreng. [GU SU FG] 

Pfaffia grandiflora (Hook.) R.E. Fr. [GU] 

AMARANTHACEAE - SYNONYMS AND 


Achyranthes indica (L.) Mill. =Achyranthes aspera 

Acnida cuspidata Bert. ex Spreng. =Amaranthus australis 

Alternanthera dentata (Moench) Scheygrond =Alternanthera 

brasiliana 

Alternanthera ficoidea (L.) R. Brown ex Roem. & Schult. 

=Alternanthera paronychoides [name often applied to A. 

tenella] 

Alternanthera ficoidea (L.) Smith =Alternanthera 

paronychoides 

Alternanthera flavogrisea (Urb.) Urb. =Alternanthera 

halimifolia 

Alternanthera flavogrisea (Urb.) Urb. subsp. diffusa (Mart.) 

Mears ex Fourn. =Alternanthera tenella 

Alternanthera maritima (Mart.) St. Hil. =Alternanthera 

littoralis var. maritima 

Alternanthera ramosissima R.E. Fries =Alternanthera 

brasiliana 

Amaranthus gracilis Desf. =Amaranthus viridis 

Amaranthus lividus L. =Amaranthus blitum 

Amaranthus paniculatus L. =Amaranthus hybridus 

Gomphrena brasiliana L. =Alternanthera brasiliana 

Iresine polymorpha Mart. =Iresine diffusa 

Iresine vermicularis (L.) Moq. =Blutaparon vermiculare 

Philoxerus vermicularis (L.) R. Brown =Blutaparon 

vermiculare 

Telanthera maritima Moq. =Alternanthera littoralis var. 

maritima


ANACARDIACEAE 

contributed by J.D. Mitchell (1992); additions 

and corrections by J.D. Mitchell (1996) 

REFERENCES: 

Barkley, F.A. 1968. Anacardiaceae: Rhoideae: 

Astronium. Phytologia 16: 107-152. 

Jansen-Jacobs, M.J. 1976. Anacardiaceae 

(additions and corrections). In J. Lanjouw & 

A.L. Stoffers, Flora of Suriname 2(2): 441- 

444. 

Mitchell, J.D. 1992. Additions to Anacardium. 

Anacardium amapaënse, a new species from 

French Guiana and eastern Amazonian Brazil. 

Brittonia 44(3): 331-338. 

Mitchell, J.D. 1993. Tapirira obtusa comb. nov. 

(Anacardiaceae). Novon 3(1): 66. 

Mitchell, J.D. 1995. Anacardiaceae. In J.A. 



Mitchell, J.D. (in press). Anacardiaceae. In 

A.R.A. Görts-van Rijn, Flora of the Guianas. 

Mitchell, J.D. & D.C. Daly. 1991. Cyrtocarpa 

Kunth (Anacardiaceae) in South America. 

Ann. Missouri Bot. Gard. 78: 184-189. 

Mitchell, J.D. & D.C. Daly. 1993. A revision of 

Thyrsodium. Brittonia 45(2): 115-129. 

Mitchell, J.D. & S. Mori. 1987. The cashew and 

its relatives (Anacardium: Anacardiaceae). 


Nannenga, E.T. 1934. Anacardiaceae. In A. 


Anacardium amapaënse J.D. Mitchell [FG] 

Anacardium fruticosum J.D. Mitchell & Mori [GU] 

Anacardium giganteum Hancock ex Engl. [GU SU] 

Anacardium occidentale L. [GU SU FG] 

Anacardium spruceanum Benth. ex Engl. [GU SU FG] 

Astronium fraxinifolium Schott [SU] 

Astronium lecointei Ducke [SU] 

Astronium ulei Mattick [GU SU FG] 

Cyrtocarpa velutinifolia (Cowan) J.D. Mitchell & Daly [GU] 

Loxopterygium sagotii J.D. Hook. [GU SU FG] 

Spondias mombin L. [GU SU FG] 

Tapirira bethanniana J.D. Mitchell [FG] 

Tapirira guianensis Aubl. [GU SU FG] 

Tapirira obtusa (Benth.) J.D. Mitchell [GU SU FG] 

Thyrsodium guianense Sagot ex March. [GU? SU FG] 

Thyrsodium puberulum J.D. Mitchell & Daly [SU FG] 

Thyrsodium spruceanum Benth. [GU SU FG] 

ANACARDIACEAE - SYNONYMS AND 


Garuga guianensis (Sagot) Engl. =Thyrsodium guianense 

Garuga schomburgkiana (Benth.) Engl. =Thyrsodium 

spruceanum 

Garuga spruceana (Benth.) Engl. =Thyrsodium spruceanum 

Mauria obtusa Benth. =Tapirira obtusa 

Tapirira marchandii Engl., nom. illeg. =Tapirira obtusa 

Tapirira peckoltiana Engl. =Tapirira obtusa 

Tapirira velutinifolia (Cowan) Marcano-Berti =Cyrtocarpa 

velutinifolia 

Thyrsodium dasytrichum Sandw. =Thyrsodium spruceanum 

Thyrsodium paraense Huber =Thyrsodium spruceanum 

Thyrsodium schomburgkianum Benth. =Thyrsodium 

spruceanum 

REFERENCES: 

ANISOPHYLLEACEAE 

Juncosa, A.M. & P.B. Tomlinson. 1988. A 

historical and taxonomic synopsis of 

Rhizophoraceae and Anisophylleaceae. Ann. 

Missouri Bot. Gard. 75: 1278-1295. 

Sandwith, N.Y. 1952. Contributions to the flora 

of tropical America LV. Discovery of 

Anisophyllea in America. Kew Bull. 1952: 

303-306. 

Anisophyllea guianensis Sandw. [GU] 

ANNONACEAE 

contributed by P.J.M. Maas (1992); additions and 

corrections by P.J. Maas & H. Rainer (1996) 

REFERENCES: 

Fries, R.E. 1930-1939. Revision der Arten 

einiger Annonaceen-Gattungen. Acta Horti 

Berg. 10(1): 1-128 (1930), 10(2): 129-341 

(1931), 12(1): 1-220 (1934), 12(2): 221-288 

(1937), 12(3): 289-577 (1939). 

Fries, R.E. 1940. Annonaceae. In A. Pulle, 


Jansen-Jacobs, M.J. 1976. Annonaceae 



687. 

Johnson, D.M. & N.A. Murray. 1995. Synopsis 

of the tribe Bocageeae (Annonaceae), with 

revisions of Cardiopetalum, Froesiodendron, 

Trigynaea, Bocagea, and Hornschuchia. 

Brittonia 47(3): 248-319. 

Maas, P.J.M. 1992. Rollinia. Flora Neotropica 

Monogr. 57: 1-192. 

Maas, P.J.M., J. Koek-Noorman, & L.Y.Th. 

Westra. 1993. Studies in Annonaceae XVIII. 

New species from the Neotropics and 

miscellaneous notes. Bot. Jahrb. 115(1): 77- 

95. 

Maas, P.J.M., E.A. Mennega, & L.Y.Th. Westra. 

1994. Studies in Annonaceae XXI. Index to 

species and infraspecific taxa of neotropical 

Annonaceae. Candollea 49(2): 389-481. 

SPERMATOPHYTA: Anacardiaceae - Annonaceae 

45

46 


Maas, P.J.M. & L.Y.Th. Westra. 1984-1985. 

Studies in Annonaceae II. A monograph of 

the genus Anaxagorea A. St. Hil. Part 1. Bot. 

Jahrb. Syst. 105: 73-134. Part 2. Bot. Jahrb. 

Syst. 105: 145-204. 

Steyermark, J.A., P.J.M. Maas, P.E. Berry, D.M. 

Johnson, N.A. Murray, & H. Rainer. 1995. 

Annonaceae. In J.A. Steyermark, P.E. Berry, 


Guayana 2: 413-469. 

Zuilen, C.M. van & P.J.M. Maas. 1994. Studies 

in Annonaceae XXII. A taxonomic revision 

of Duguetia A. St.-Hil., sections Alcmene and 

Xylopipetalum. Bot. Jahrb. 116(2): 221-242. 

Anaxagorea acuminata (Dunal) A. DC. [GU SU FG] 

Anaxagorea brevipedicellata Timmerman [FG] 

Anaxagorea brevipes Benth. [GU SU] 

Anaxagorea dolichocarpa Sprague & Sandw. [GU SU FG] 

Anaxagorea petiolata R.E. Fries [GU] 

Anaxagorea phaeocarpa Mart. [FG] 

Anaxagorea prinoides (Dunal) A. DC. [SU FG] 

Annona ambotay Aubl. [GU SU FG] 

Annona densicoma Mart. [GU SU FG] 

Annona echinata Dunal [SU FG] 

Annona foetida Mart. [SU FG] 

Annona glabra L. [GU SU FG] 

Annona haematantha Miq. [GU SU FG] 

Annona hypoglauca Mart. [GU SU FG] 

Annona montana Macfad. [GU SU FG] 

Annona muricata L. [GU SU FG] 

Annona paludosa Aubl. [GU SU FG] 

Annona sericea Dunal [GU SU FG] 

Annona symphyocarpa Sandw. [GU FG] 

Annona tenuiflora Mart. [FG] 

Annona tenuipes R.E. Fries [GU] 

Annona trunciflora R.E. Fries [FG] 

Annona ulei R.E. Fries [GU] 

Bocageopsis multiflora (Mart.) R.E. Fries [GU SU FG] 

Cardiopetalum surinamense R.E. Fries [SU FG] 

Cremastosperma brevipes (DC. ex Dunal) R.E. Fries [FG] 

Cymbopetalum brasiliense (Vell.) Benth. ex Baill. [GU SU 

FG] 

Duguetia amplexifolia R.E. Fries [SU FG] 

Duguetia cadaverica Huber [GU SU FG] 

Duguetia calycina Benoist [GU SU FG] 

Duguetia cauliflora R.E. Fries [GU SU FG] 

Duguetia decurrens R.E. Fries [GU] 

Duguetia eximia Diels [SU FG] 

Duguetia guianensis R.E. Fries [GU SU] 

Duguetia inconspicua Sagot [GU SU FG] 

Duguetia lepidota (Miq.) Pulle [SU] 

Duguetia lucida Urb. [GU] 

Duguetia macrocalyx R.E. Fries [GU] 

Duguetia megalophylla R.E. Fries [GU] 

Duguetia neglecta Sandw. [GU SU] 

Duguetia obovata R.E. Fries [GU SU FG] 

Duguetia paraensis R.E. Fries [GU FG] 

Duguetia pauciflora Rusby [GU FG] 

Duguetia pycnastera Sandw. [GU SU FG] 

Duguetia quitarensis Benth. [GU] 

Duguetia rigida R.E. Fries [GU] 

Duguetia riparia Huber [GU SU FG] 

SPERMATOPHYTA: Annonaceae 

Duguetia schulzii Jansen-Jacobs [SU] 

Duguetia surinamensis R.E. Fries [GU SU FG] 

Duguetia tenuis R.E. Fries [GU] 

Ephedranthus guianensis R.E. Fries [GU SU FG] 

Fusaea longifolia (Aubl.) Safford [GU SU FG] 

Guatteria atra Sandw. [GU] 

Guatteria brachypoda R.E. Fries [GU] 

Guatteria citriodora Ducke [GU SU FG] 

Guatteria clusiifolia D.M. Johnson & N.A. Murray [GU] 

Guatteria conspicua R.E. Fries [GU SU FG?] 

Guatteria discolor R.E. Fries [FG] 

Guatteria elongata Benth. [GU] 

Guatteria ferruginea A. St. Hil. [SU] 

Guatteria flexilis R.E. Fries [GU] 

Guatteria foliosa Benth. [GU SU? FG?] 

Guatteria gracilipes R.E. Fries [SU] 

Guatteria guianensis (Aubl.) R.E. Fries [FG] 

Guatteria insignis R.E. Fries [SU] 

Guatteria maguirei R.E. Fries [GU?] 

Guatteria maypurensis Kunth [GU] 

Guatteria megalophylla Diels [GU] 

Guatteria microsperma R.E. Fries [SU FG] 

Guatteria monticola R.E. Fries [GU] 

Guatteria oblonga R.E. Fries [FG] 

Guatteria obovata R.E. Fries [GU] 

Guatteria ouregou (Aubl.) Dunal [FG] 

Guatteria ovalifolia R.E. Fries [GU] 

Guatteria paludosa R.E. Fries [GU] 

Guatteria procera R.E. Fries [GU SU] 

Guatteria pteropus Benth. [SU] 

Guatteria punctata (Aubl.) R.A. Howard [GU SU FG] 

Guatteria recurvisepala R.E. Fries [GU] 

Guatteria richardii R.E. Fries [FG] 

Guatteria rubrinervis R.E. Fries [GU] 

Guatteria sagotiana R.E. Fries [FG] 

Guatteria scandens Ducke [GU SU FG] 

Guatteria schomburgkiana Mart. [GU SU FG] 

Guatteria stenopetala R.E. Fries [GU] 

Guatteria trichostemon R.E. Fries [SU FG] 

Guatteria umbonata R.E. Fries [GU] 

Guatteria wachenheimii Benoist [SU FG] 

Guatteria wessels-boerii Jansen-Jacobs [SU] 

Guatteriopsis blepharophylla (Mart.) R.E. Fries [SU FG] 

Malmea discolor R.E. Fries [GU SU] 

Malmea guianensis R.E. Fries [GU] 

Malmea sp. nov. 1 [SU] 

Malmea sp. nov. 2 [SU] 

Oxandra asbeckii (Pulle) R.E. Fries [GU SU FG] 

Oxandra guianensis R.E. Fries [GU] 

Oxandra krukoffii R.E. Fries [SU] 

Oxandra surinamensis Jansen-Jacobs [SU] 

Oxandra xylopioides Diels [GU SU FG] 

Pseudoxandra cuspidata Maas [FG] 

Pseudoxandra polyphleba (Diels) R.E. Fries [GU FG] 

Rollinia cuspidata Mart. [FG] 

Rollinia elliptica R.E. Fries [SU FG] 

Rollinia exsucca (DC. ex Dunal) A. DC. [GU SU FG] 

Rollinia mucosa (Jacq.) Baill. [GU SU FG] 

Trigynaea caudata (R.E. Fries) R.E. Fries [GU SU] 

Trigynaea duckei (R.E. Fries) R.E. Fries [GU] 

Unonopsis glaucopetala R.E. Fries [GU SU FG] 

Unonopsis guatterioides (A. DC.) R.E. Fries [GU SU FG] 

Unonopsis perrottetii (A. DC.) R.E. Fries [GU FG] 

Unonopsis rufescens (Baill.) R.E. Fries [GU SU FG] 

Unonopsis stipitata Diels [GU SU FG]


Xylopia amazonica R.E. Fries [SU] 

Xylopia aromatica (Lam.) Mart. [GU SU FG] 

Xylopia benthamii R.E. Fries [GU SU FG] 

Xylopia cayennensis Maas [GU SU FG] [replacement name 

for Xylopia longifolia] 

Xylopia crinita R.E. Fries [FG] 

Xylopia discreta (L.f.) Sprague & Hutchinson [GU SU FG] 

Xylopia excellens R.E. Fries [FG] 

Xylopia frutescens Aubl. [GU SU FG] 

Xylopia nervosa (R.E. Fries) Maas [FG] 

Xylopia nitida Dunal [GU SU FG] 

Xylopia parviflora Spruce [FG] 

Xylopia pulcherrima Sandw. [GU SU FG] 

Xylopia sericea A. St. Hil. [GU SU FG] 

Xylopia surinamensis R.E. Fries [GU SU FG] 

ANNONACEAE - SYNONYMS AND 


Aberemoa guianensis Aubl. =Guatteria guianensis 

Anaxagorea mutica R.E. Fries =Anaxagorea dolichocarpa 

Annona jenmanii Safford =Annona sericea 

Annona longifolia Aubl. =Fusaea longifolia 

Annona punctata Aubl. =Guatteria punctata 

Bocagea asbeckii Pulle =Oxandra asbeckii 

Cananga ouregou Aubl. =Guatteria ouregou 

Cremastosperma guianense R.E. Fries =Pseudoxandra 

polyphleba? 

Cremastosperma poiteaui (Diels) R.E. Fries =Cremastosperma 

brevipes 

Duguetia adiscandra Jansen-Jacobs =Duguetia cadaverica 

Duguetia cuspidata R.E. Fries =Duguetia calycina 

Duguetia elegans R.E. Fries =Duguetia calycina 

Duguetia friesii Jansen-Jacobs =Duguetia cadaverica 

Duguetia insculpta R.E. Fries =Duguetia riparia 

Duguetia latifolia R.E. Fries [not in Guianas] 

Duguetia sandwithii R.E. Fries =Duguetia calycina 

Duguetia tessmannii R.E. Fries [not in Guianas] 

Duguetia yeshidan Sandw. =Duguetia calycina 

Froesiodendron surinamense (R.E. Fries) R.E. Fries 

=Cardiopetalum surinamense 

Guatteria blepharophylla Mart. =Guatteriopsis blepharophylla 

Guatteria chrysopetala (Steud.) Miq. =Guatteria punctata 

Guatteria crassipetala (Dunal) DC ex Lemée =Guatteria 

ouregou 

Guatteria ouregou Griseb., non Dunal =Guatteria inuncta [not 


Guatteria ouregou Mart., non Dunal =Guatteria pteropus 

Guatteria podocarpa DC. ex Dunal =Guatteria ouregou 

Guatteria sandwithii R.E. Fries =Guatteria schomburgkiana 

Hornschuchia caudata R.E. Fries =Trigynaea caudata 

Hornschuchia duckei R.E. Fries =Trigynaea duckei 

Hornschuchia guianensis R.E. Fries =Trigynaea duckei 

Malmea obovata R.E. Fries [not in Guianas] 

Oxandra nervosa R.E. Fries =Xylopia nervosa 

Pseudoxandra guianensis (R.E. Fries) R.E. Fries 

=Pseudoxandra polyphleba? 

Raimondia tenuiflora (Mart.) R.E. Fries =Annona tenuiflora 

Rollinia cardiantha Diels =Rollinia cuspidata 

Rollinia puberula A. DC. =Rollinia exsucca 

Rollinia pulchrinervia A. DC. =Rollinia mucosa 

Rollinia sieberi A. DC. =Rollinia mucosa 

Rollinia surinamensis R.E. Fries =Rollinia exsucca 

Trigynaea guianensis (R.E. Fries) R.E. Fries =Trigynaea 

duckei 

Unona crassipetala Dunal =Guatteria ouregou 

Waria zeylanica Aubl. =Xylopia nitida?[non Uvaria zeylanica 

L.] 

Xylopia longifolia (Sagot) R.E. Fries =Xylopia cayennensis 

Xylopia longifolia A. DC. =Xylopia aromatica 

Xylopia nitida Dunal var. longifolia Sagot =Xylopia 

cayennensis 

APIACEAE 

reviewed by J.C. Lindeman (1992); reviewed by 

J.C. Lindeman (1996) 

REFERENCES: 

Berry, P.E. 1995. Apiaceae. In J.A. Steyermark, 

P.E. Berry, & B.K. Holst, Flora of the 

Venezuelan Guayana 2: 470-471. 

Mathias, M.E. 1936. The genus Hydrocotyle in 

northern South America. Brittonia 2: 201- 

237. 

Stafleu, F.A. 1951. Umbelliferae. In A. Pulle, 


Eryngium foetidum L. [GU SU FG] 

Hydrocotyle rotundifolia Roxb. [GU?] [introduced, no recent 

records] 

Hydrocotyle umbellata L. [GU SU FG] 

APOCYNACEAE 

reviewed by L. Allorge (1992); reviewed by 

A.J.M. Leeuwenberg & B.F. Hansen (1996) 

REFERENCES: 

Allorge, L. 1985. Monographie des 

Apocynacées - Tabernaemontanoidées 

Américaines. Mem. Mus. Nat. Hist. Naturelle 

(n.s.) Serie B, tome 30. 

Allorge, L. 1985. Distribution, relations et 

entérêt biogéographique des 

Tabernaemontanoideae (Apocynaceae) dans 

la région néotropicale. C.R. Soc. Biogéogr. 

61(4): 137-155. 

Allorge, L. & C. Poupat. 1991. Position 

systématique et révision du genre 

Aspidosperma (Apocynaceae) pour les trois 

Guyanes. Le point sur leur étude chimique. 

Bull. Soc. bot. Fr., Lettres bot. (4/5): 267-301. 

Allorge, L. & C. Sastre. 1990. Distribution 

géographique dans les Guyanes, 

d'Apocynaceae et d'Ochnaceae. Essai de mise 

en evidence de centres de speciation. C.R. 

Soc. Biogéogr. 67(2): 77-94. 

Leeuwenberg, A.J.M. 1988. Series of revisions 

of Apocynaceae. XXI. Notes on 

SPERMATOPHYTA: Annonaceae - Apocynaceae 

47

48 


Tabernaemontaneae. Agric. Univ. Wageningen 

Papers 87-5: 1-32. 

Leeuwenberg, A.J.M. 1994. A Revision of 

Tabernaemontana. Two. The New World 

Species and Stemmadenia. Royal Botanic 

Gardens, Kew, Richmond, Surrey, England. 

Markgraf, F. 1932. Apocynaceae. In A. Pulle, 


Markgraf, F. 1937. Apocynaceae (additions and 

corrections). In A. Pulle, Flora of Suriname 

4(1): 443-467. 

Monachino, J.V. 1943 [1944]. A revision of 

Couma and Parahancornia. Lloydia 6(4): 

229-247. 

Monachino, J. 1961. Apocynaceae. In B. 




Plumel, M.-M. 1991. Le genre Himatanthus, 

révision taxonomique. Bradea 5 suppl.: 1- 

118. 

Woodson, R.E., Jr. 1933-1936. Studies in the 

Apocynaceae. IV. The American genera of 

Echitoideae. Ann. Missouri Bot. Gard. 20: 

605-790, 22: 153-306, 23: 169-439. 

Zarucchi, J.L., G.N. Morillo, M.E. Endress, B.F. 

Hansen & A.J.M. Leeuwenberg. 1995. 

Apocynaceae. In J.A. Steyermark, P.E. Berry, 


Guayana 2: 471-571. 

Allamanda cathartica L. [GU SU FG] 

Allamanda schottii Pohl [FG] 

Allamanda setulosa Miq. [SU] 

Ambelania acida Aubl. [GU SU FG] 

Aspidosperma album (Vahl) Benoist ex Pichon [GU SU FG] 

Aspidosperma carapanauba Pichon [FG] 

Aspidosperma cruentum Woodson [GU SU FG] 

Aspidosperma excelsum Benth. [GU SU FG] 

Aspidosperma helstonei van Donselaar [SU FG] 

Aspidosperma macrophyllum Muell. Arg. subsp. 

macrophyllum [GU] 

Aspidosperma macrophyllum Muell. Arg. subsp. morii Allorge 

[FG] 

Aspidosperma marcgravianum Woodson [GU SU FG] 

Aspidosperma oblongum A. DC. [GU SU FG] 

Aspidosperma sandwithianum Markgr. [GU SU FG] 

Aspidosperma spruceanum Benth. ex Muell. Arg. [GU SU 

FG] 

Aspidosperma ulei Markgr. [GU SU] 

Aspidosperma vargasii A. DC. [GU SU] 

Catharanthus roseus (L.) G. Don [GU SU FG] [widely 

cultivated, escaping] 

Condylocarpon amazonicum (Markgr.) Ducke [SU FG?] 

Condylocarpon guyanense Desf. [GU FG] 

Condylocarpon intermedium Muell. Arg. var. intermedium 

[GU SU FG] 

Condylocarpon myrtifolium (Miq.) Muell. Arg. [GU SU FG] 

Couma catingae Ducke [GU] 

Couma guianensis Aubl. [GU SU FG] 

Couma macrocarpa Barb. Rodr. [GU] 

Couma rigida Muell. Arg. [GU] 

Forsteronia acouci (Aubl.) A. DC. [GU SU FG] 

SPERMATOPHYTA: Apocynaceae 

Forsteronia adenobasis Muell. Arg. [GU FG] 

Forsteronia diospyrifolia Muell. Arg. [GU] 

Forsteronia duckei Markgr. [GU] 

Forsteronia gracilis (Benth.) Muell. Arg. [GU SU] 

Forsteronia guyanensis Muell. Arg. [GU SU FG] 

Forsteronia obtusiloba Muell. Arg. [SU] [to be expected in 

GU] 

Forsteronia schomburgkii A. DC. [GU] 

Forsteronia umbellata (Aubl.) Woodson [FG] 

Galactophora schomburgkiana Woodson var. schomburgkiana 

[GU] 

Geissospermum argenteum Woodson [GU SU FG] 

Geissospermum laevis (Vell.) Miers [GU SU FG] 

Geissospermum sericeum (Sagot ex Prog.) Benth. & J.D. 

Hook. [GU SU FG] 

Himatanthus articulatus (Vahl) Woodson [GU SU FG] 

Himatanthus bracteatus (A. DC.) Woodson var. bracteatus 

[GU SU FG] 

Himatanthus bracteatus (A. DC.) Woodson var. revolutus 

(Huber) Plumel [GU] 

Himatanthus drasticus (Mart.) Plumel [GU SU FG] 

Himatanthus fallax (Muell. Arg.) Plumel [GU SU FG] 

Himatanthus obovatus (Muell. Arg.) Woodson [GU] 

Himatanthus speciosus (Muell. Arg.) Plumel [FG] 

Himatanthus stenophyllus Plumel [SU] 

Himatanthus sucuuba (Spruce ex Muell. Arg.) Woodson [GU 

SU FG] 

Kopsia arborea Blume [FG] 

Lacmellea aculeata (Ducke) Monachino [GU SU FG] 

Lacmellea arborescens (Muell. Arg.) Markgr. [GU?] 

Lacmellea floribunda (Poepp.) Benth. & J.D. Hook. [SU FG] 

Lacmellea guyanensis (Muell. Arg.) Monachino [FG] 

Lacmellea utilis (Arnott) Markgr. [GU FG?] 

Laxoplumeria baehniana Monachino [FG] 

Macoubea guianensis Aubl. [GU SU FG] 

Macropharynx spectabilis (Stadelm.) Woodson [GU SU] 

Macropharynx strigillosa Woodson [GU] 

Malouetia flavescens (Willd. ex Roem. & Schult.) Muell. Arg. 

[GU SU FG] 

Malouetia gracilis (Benth.) A. DC. [GU FG?] 

Malouetia pubescens Markgr. var. glabra M.E. Endress [GU] 

Malouetia tamaquarina (Aubl.) A. DC. [GU SU FG] 

Mandevilla benthamii (A. DC.) K. Schum. [GU] 

Mandevilla hirsuta (A. Rich.) K. Schum. [GU SU FG] 

Mandevilla leptophylla (A. DC.) K. Schum. [GU] 

Mandevilla rugellosa (L.C. Rich.) L. Allorge [GU SU FG] 

Mandevilla rugosa (Benth.) Woodson [GU] 

Mandevilla scaberula N.E. Brown [GU FG] 

Mandevilla scabra (Hoffmanns. ex Roem. & Schult.) K. 

Schum. [GU SU FG] 

Mandevilla spruceana (Muell. Arg.) K. Schum. [GU] 

Mandevilla subcarnosa (Benth.) Woodson var. angustata 

Steyerm. [GU] 

Mandevilla subcarnosa (Benth.) Woodson var. subcarnosa 

[GU] 

Mandevilla subspicata (Vahl) Markgr. [GU SU FG] 

Mandevilla surinamensis (Pulle) Woodson [GU SU FG] 

Mandevilla symphitocarpa (G. Mey.) Woodson [GU SU FG] 

Mandevilla tenuifolia (Miq.) Woodson [SU] 

Mandevilla vanheurckii (Muell. Arg.) Markgr. [GU?] 

Mesechites trifida (Jacq.) Muell. Arg. [GU SU FG] 

Nerium oleander L. [GU SU FG] [cultivated, perhaps 

escaping] 

Odontadenia geminata (Roem. & Schult.) Muell. Arg. [GU SU 

FG]


Odontadenia glauca Woodson [GU?] 

Odontadenia macrantha (Roem. & Schult.) Markgr. [GU SU 

FG] 

Odontadenia nitida (Vahl) Muell. Arg. [GU SU FG] 

Odontadenia perrottetii (A. DC.) Woodson [GU FG] 

Odontadenia puncticulosa (A. Rich.) Pulle [GU SU FG] 

Odontadenia sandwithiana Woodson [GU] 

Odontadenia verrucosa (Roem. & Schult.) K. Schum. ex 

Markgr. [GU SU] 

Pacouria guianensis Aubl. [GU SU FG] 

Parahancornia fasciculata (Poir.) Benoist ex Pichon [GU SU 

FG] 

Plumeria inodora Jacq. [GU] 

Plumeria rubra L. [GU SU FG] [cultivated & escaped] 

Prestonia acutifolia (Benth. ex Muell. Arg.) K. Schum. [GU 

SU] 

Prestonia annularis (L.f.) G. Don [GU SU FG] 

Prestonia cayennensis (A. DC.) Pichon [GU FG] 

Prestonia coalita (Vell.) Woodson [GU FG] 

Prestonia guianensis Gleason [GU] 

Prestonia ipomaeifolia A. DC. [FG] 

Prestonia lindleyana Woodson [GU?] 

Prestonia megagros (Vell.) Woodson [GU?] 

Prestonia perplexa Woodson [GU?] 

Prestonia quinquangularis Spreng. [GU?] 

Prestonia surinamensis Muell. Arg. [GU SU FG] 

Prestonia tomentosa R. Brown [GU SU?] 

Rauvolfia ligustrina Willd. ex Roem. & Schult. [GU SU] 

Rauvolfia pachyphylla Markgr. [GU] 

Rauvolfia paraensis Ducke [SU FG] 

Rauvolfia tetraphylla L. [FG] 

Rhabdadenia biflora (Jacq.) Muell. Arg. [GU SU FG] 

Rhabdadenia macrostoma (Benth.) Muell. Arg. [GU SU FG] 

Rhabdadenia pohlii Muell. Arg. [GU SU FG] 

Secondatia densiflora A. DC. [GU SU FG] 

Stemmadenia grandiflora (Jacq.) Miers [GU SU FG] 

Tabernaemontana albiflora (Miq.) Pulle [GU SU FG] 

[=Bonafousia albiflora] 

Tabernaemontana angulata Mart. ex Muell. Arg. [GU FG] 

[=Bonafousia angulata] 

Tabernaemontana attenuata (Miers) Urb. [GU SU FG] 

[=Anartia meyeri] 

Tabernaemontana cerea (Woodson) Leeuwenberg [GU SU] 

[=Anartia cerea] 

Tabernaemontana cymosa Jacq. [GU] [=Peschiera cymosa] 

Tabernaemontana disticha A. DC. [GU SU FG] [=Bonafousia 

disticha] 

Tabernaemontana divaricata (L.) R. Br. ex Roem. & Schult. 

[GU] [cultivated or escaped from cultivation] 

Tabernaemontana flavicans Willd. ex Roem. & Schult. [GU? 

SU? FG?] [=Anartia flavicans] 

Tabernaemontana heterophylla Vahl [GU SU FG] 

[=Stenosolen heterophyllus] 

Tabernaemontana lagenaria Leeuwenberg [SU? FG?] 

Tabernaemontana lorifera (Miers) Leeuwenberg [GU SU] 

[=Bonafousia lorifera] 

Tabernaemontana macrocalyx Muell. Arg. [GU SU FG] 

[=Bonafousia macrocalyx] 

Tabernaemontana pandacaqui Lam. [FG] [Asian species, 

cultivated only?] 

Tabernaemontana rupicola Benth. [GU SU FG] [=Bonafousia 

rupicola] 

Tabernaemontana sananho Ruiz & Pav. [GU SU FG] 

[=Bonafousia sananho] 

Tabernaemontana siphilitica (L.f.) Leeuwenberg [GU SU FG] 

[=Bonafousia siphilitica] 

Tabernaemontana undulata Vahl [GU SU FG] [=Bonafousia 

undulata] 

Thevetia peruviana (Pers.) K. Schum. [GU SU FG] 

APOCYNACEAE - SYNONYMS AND 


Ambelania sagotii Muell. Arg. =Ambelania acida 

Anacampta albescens (Rusby) Markgr. =Tabernaemontana 

undulata 

Anacampta disticha (A. DC.) Markgr. =Tabernaemontana 

disticha 

Anacampta echinata (Aubl.) Markgr. [see Tabernaemontana 

echinata, Peschiera echinata] 

Anacampta macrocalyx (Muell. Arg.) Miers 

=Tabernaemontana macrocalyx 

Anacampta rigida (Miers) Markgr. =Tabernaemontana 

muricata Link ex Roem. & Schult.[not in Guianas] 

Anacampta rupicola (Benth.) Markgr. =Tabernaemontana 

rupicola 

Anartia attenuata (Miers) Markgr. =Tabernaemontana 

attenuata 

Anartia cerea (Woodson) Allorge =Tabernaemontana cerea 

Anartia meyeri (G. Mey. ex G. Don) Miers =Tabernaemontana 

attenuata 

Anartia oblongifolia (A. DC.) Markgr. =Tabernaemontana 

flavicans 

Anartia olivacea (Muell. Arg.) Markgr. =Tabernaemontana 

flavicans 

Anechites nerium Urb. [not in Guianas] 

Anisolobus puncticulosus (A. Rich.) Miq. =Odontadenia 

puncticulosa 

Apocynum acouci Aubl. =Forsteronia acouci 

Apocynum umbellatum Aubl. =Forsteronia umbellata 

Aspidosperma desmanthum Benth. ex Muell. Arg. 

=Aspidosperma album 

Aspidosperma megalocarpon Muell. Arg. [not in Guianas] 

Aspidosperma nitidum Benth. ex Muell. Arg. [not in Guianas] 

Aspidosperma woodsonianum Markgr. =Aspidosperma 

spruceanum 

Bonafousia albiflora (Miq.) Boit. & Allorge 

=Tabernaemontana albiflora 

Bonafousia angulata (Mart. ex Muell. Arg.) Boit. & Allorge 

=Tabernaemontana angulata 

Bonafousia attenuata Miers =Tabernaemontana attenuata 

Bonafousia disticha (A. DC.) Boit. & Allorge 

=Tabernaemontana disticha 

Bonafousia lorifera (Miers) Boit. & Allorge 

=Tabernaemontana lorifera 

Bonafousia macrocalyx (Muell. Arg.) Boit. & Allorge 

=Tabernaemontana macrocalyx 

Bonafousia morettii Allorge =Tabernaemontana macrocalyx 

Bonafousia obliqua Miers =Tabernaemontana undulata 

Bonafousia rupicola (Benth.) Miers =Tabernaemontana 

rupicola 

Bonafousia rupicola (Benth.) Miers var. versicolor (Woods.) 

Allorge =Tabernaemontana rupicola 

Bonafousia sananho (Ruiz & Pav.) Markgr. 

=Tabernaemontana sananho 

Bonafousia siphilitica (L.f.) Allorge =Tabernaemontana 

siphilitica 

SPERMATOPHYTA: Apocynaceae 

49

50 


Bonafousia tetrastachya (Kunth) Markgr. =Tabernaemontana 

siphilitica 

Bonafousia undulata (Vahl) A. DC. =Tabernaemontana 

undulata 

Cameraria tamaquarina Aubl. =Malouetia tamaquarina 

Condylocarpon rauwolfiae Muell. Arg. =Condylocarpon 

isthmicum (Vell.) A. DC.[not in Guianas] 

Dipladenia surinamensis Pulle =Mandevilla surinamensis 

Echites grandiflora G. Mey. =Odontadenia macrantha 

Echites puncticulosa A. Rich. =Odontadenia puncticulosa 

Echites spectabilis Stadelm. =Macropharynx spectabilis 

Echites trifida Jacq. =Mesechites trifida 

Echites tubulosa Benth. =Mesechites trifida 

Ervatamia coronaria (Jacq.) Stapf =Tabernaemontana 

divaricata 

Ervatamia cumingiana (A. DC.) Markgr. =Tabernaemontana 

pandacaqui 

Forsteronia corymbifera (Miers) Sandw. =Forsteronia 

schomburgkii 

Forsteronia laurifolia (Benth.) A. DC. [not in Guianas] 

Forsteronia schomburgkii (Benth.) Muell. Arg. =Forsteronia 

acouci 

Haemadictyon cayennense A. DC. =Prestonia cayennensis 

Hortsmania myrtifolia Miq. =Condylocarpon myrtifolium 

Landolphia guyanensis (Aubl.) Pulle =Pacouria guianensis 

Lochnera rosea (L.) Rchb. =Catharanthus roseus 

Macoubea fasciculata (Poir.) Lemée =Parahancornia 

fasciculata 

Macoubea guianensis Aubl. var. reticulata (A. DC.) Allorge 

=Macoubea guianensis 

Macoubea reticulata (A. DC.) Markgr. =Macoubea guianensis 

Malouetia furfuracea Spruce ex Muell. Arg. =Malouetia 

tamaquarina 

Malouetia guianensis (Aubl.) Miers =Malouetia tamaquarina 

Malouetia obtusiloba A. DC. =Malouetia tamaquarina 

Malouetia odorata DC. =Malouetia tamaquarina 

Malouetia retroflexa Muell. Arg. =Malouetia cubana A. 

DC.[not in Guianas] 

Malouetia schomburgkii Muell. Arg. =Malouetia flavescens 

Mandevilla glabra N.E. Br. =Mandevilla vanheurckii 

Mandevilla lutea (Vell.) Markgr. =Mandevilla scaberula 

Mesechites syphilitica (L.f.) Lemée =Tabernaemontana 

siphilitica 

Odontadenia "schomburgkiana (Nees) Kr." [invalid name, 

error for Odontonema schomburgkiana (Acanthaceae)?] 

Odontadenia amazonica (Stadelm.) Malme =Odontadenia 

verrucosa 

Odontadenia cognata (Stadelm.) Woodson =Odontadenia 

verrucosa 

Odontadenia cururu (Mart.) K. Schum. =Odontadenia 

puncticulosa 

Odontadenia grandiflora (G. Mey.) Kuntze =Odontadenia 

macrantha 

Odontadenia hoffmannseggiana (Steud.) Woodson 

=Odontadenia macrantha 

Odontadenia silvestris (A. DC.) Muell. Arg. =Odontadenia 

macrantha 

Odontadenia surinamensis (Pulle) Woodson =Odontadenia 

verrucosa 

Orelia grandiflora Aubl. =Allamanda cathartica 

Parahancornia amapa (Huber) Ducke =Parahancornia 

fasciculata 

Parahancornia tabernaemontana Woodson =Macoubea 

guianensis 

SPERMATOPHYTA: Apocynaceae, Aquifoliaceae 

Peschiera cymosa (Jacq.) Dugand =Tabernaemontana cymosa 

Peschiera echinata (Aubl.) A. DC. [SU FG] [name misapplied 

to Tabernaemontana lagenaria] 

Plumeria articulata Vahl =Himatanthus articulatus 

Plumeria bracteata A. DC. =Himatanthus bracteatus 

Plumeria sucuuba R. Spruce ex Muell. Arg. =Himatanthus 

sucuuba 

Prestonia marginata (Benth.) Woodson =Prestonia cayennensis 

Rauvolfia hirsuta Jacq. =Rauvolfia tetraphylla 

Rauvolfia lamarckii DC. =Rauvolfia tetraphylla 

Rauvolfia ternifolia Kunth =Rauvolfia ligustrina 

Stemmadenia cerea Woodson =Tabernaemontana cerea 

Stenosolen heterophyllus (Vahl) Markgr. =Tabernaemontana 

heterophylla 

Taberna albiflora (Miq.) Markgr. =Tabernaemontana albiflora 

Tabernaemontana albescens Rusby =Tabernaemontana 

undulata 

Tabernaemontana aubletii Pulle, nom. illeg. =Macoubea 

guianensis 

Tabernaemontana citrifolia Muell. Arg. [not in Guianas] 

Tabernaemontana coronaria (Jacq.) Willd. =Tabernaemontana 

divaricata 

Tabernaemontana echinata Vell., non Aubl. 

=Tabernaemontana hystrix Steud.[not in Guianas] 

Tabernaemontana fasciculata Poir. =Parahancornia fasciculata 

Tabernaemontana grandiflora Jacq. =Stemmadenia grandiflora 

Tabernaemontana laurifolia L. [not in Guianas] 

Tabernaemontana longifolia Benth. =Tabernaemontana 

siphilitica 

Tabernaemontana macrophylla Muell. Arg. 

=Tabernaemontana muricata Link ex Roem. & Schult.[not 


Tabernaemontana meyeri G. Don =Tabernaemontana attenuata 

Tabernaemontana oblongifolia A. DC. =Tabernaemontana 

flavicans 

Tabernaemontana repanda E. Meyer =Tabernaemontana 

siphilitica 

Tabernaemontana speciosa Poir. =Tabernaemontana siphilitica 

Tabernaemontana submollis Mart. ex Muell. Arg. 

=Tabernaemontana coriacea Link ex Roem. & Schult.[not 


Tabernaemontana tetrastachya Kunth =Tabernaemontana 

siphilitica 

Tabernaemontana utilis Arnott =Lacmellea utilis 

Tabernaemontana versicolor Woodson =Tabernaemontana 

rupicola 

Thevetia ahouai A. DC. [not in Guianas] 

Willughbeja guianensis (Aubl.) J.F. Gmel. =Pacouria 

guianensis 

Zschokkea aculeata Ducke =Lacmellea aculeata 

AQUIFOLIACEAE 

reviewed by P.-A. Loizeau (1992)

REFERENCES: 


Amshoff, G.J.H. 1951. Aquifoliaceae. In A. 


Edwin, G. 1965. Aquifoliaceae. In B. Maguire 

& collaborators, Botany of the Guayana 

Highland Part VI. Mem. New York Bot. 

Gard. 12(3): 1-285. 

Steyermark, J.A. & P.E. Berry. 1995. 

Aquifoliaceae. In J.A. Steyermark, P.E. 


Guayana 2: 571-599. 

Wurdack, J.J. 1961. Aquifoliaceae. In B. 




Ilex apicidens N.E. Brown [GU] 

Ilex casiquiarensis Loes. [FG] 

Ilex costata Edwin [GU] 

Ilex daphnogenea Reissek [GU FG] 

Ilex fanshawei Edwin [GU] 

Ilex guianensis (Aubl.) Kuntze [GU SU FG] 

Ilex inundata Poepp. ex Reissek [SU FG?] 

Ilex jenmanii Loes. [GU SU] 

Ilex laureola Triana [GU] 

Ilex martiniana D. Don [GU SU FG] 

Ilex oliveriana Loes. [GU] 

Ilex ovalifolia G. Mey. [GU SU FG] 

Ilex retusa Klotzsch ex Reissek [GU] 

Ilex soderstromii Edwin var. ovata Edwin [GU] 

Ilex soderstromii Edwin var. soderstromii [GU] 

Ilex tepuiana Steyerm. ex Edwin [GU] 

Ilex thyrsiflora Klotzsch ex Reissek var. schomburgkii 

(Klotzsch ex Reissek) Loes. [GU] 

Ilex umbellata Klotzsch ex Reissek [GU] 

Ilex vacciniifolia Klotzsch ex Reissek [GU] 

Ilex vismiifolia Reissek [GU] 

AQUIFOLIACEAE - SYNONYMS AND 


Ilex acuminata Willd. =Ilex guianensis 

Ilex celastroides Klotzsch [nomen nudum] =Ilex vismiifolia 

Ilex congesta Reissek =Cybianthus lineatus (Myrsinaceae) 

Ilex humirioides Reissek =Ilex umbellata 

Ilex humirioides Reissek =Ilex umbellata 

Ilex lanceolata Klotzsch [nomen nudum] =Ilex martiniana 

Ilex laurina Klotzsch [nomen nudum] =Ilex daphnogenea 

Ilex macoucoua Pers. =Ilex guianensis 

Ilex riparia Reissek =Ilex inundata 

Ilex schomburgkii Klotzsch ex Reissek =Ilex thyrsiflora var. 

schomburgkii 

Macoucoua guianensis Aubl. =Ilex guianensis 

ARACEAE 

contributed by T. Croat, reviewed by D. Nicolson 

(1992); additions and corrections by T. Croat, 

reviewed by D. Nicolson (1996) 

REFERENCES: 

Bunting, G.S. 1960. A revision of 

Spathiphyllum. Mem. New York Bot. Gard. 

10(3): 1-54. 

Bunting, G.S. 1995. Araceae. In J.A. 



Croat, T.B. 1981. A revision of Syngonium. 


Croat, T.B. 1991. A revision of Anthurium 

section Pachyneurium. Ann. Missouri Bot. 

Gard. 78(3): 539-855. 

Croat, T.B. 1995 ["1994"]. Taxonomic status of 

Neotropical Araceae. Aroideana 17: 33-60. 

Croat, T.B. 1995. Two new species of Araceae 

for the Guianas. Novon 5(1): 24-29. 

Croat, T.B. & M.H. Grayum. 1994. 

Philodendron subgenus Pteromischum from 

Saül, French Guiana. Novon 4(3): 211-219. 

Jonker-Verhoef, A.M.E. & F.P. Jonker. 1953. 

Araceae. In A. Pulle, Flora of Suriname 1(2): 

1-80. 


Araceae (additions and corrections). In A. 

Pulle & J. Lanjouw, Flora of Suriname 1(2): 

380-412. 

Madison, M. 1977. A revision of Monstera. 

Contrib. Gray Herb. 207: 3-100. 

Madison, M. 1981. Notes on and its allies. 

Selbyana 5(3-4): 342-377. 

Alocasia macrorrhiza (L.) G. Don [SU] 

Anaphyllopsis americanum (Engl.) Hay [SU FG] 

Anthurium bakeri J.D. Hook. [GU] 

Anthurium bonplandii Bunting subsp. bonplandii [FG] 

Anthurium bonplandii Bunting subsp. guayanum (Bunting) 

Croat [GU SU FG?] 

Anthurium clavigerum Poepp. & Endl. [GU SU FG] 

Anthurium cowanii Croat [GU SU] 

Anthurium cremersii Croat, sp. nov. ined. [FG] 

Anthurium eminens Schott [GU SU FG] 

Anthurium expansum Gleason [GU SU FG] 

Anthurium gracile (Rudge) Lindl. [GU SU FG] 

Anthurium hookeri Kunth [GU SU FG] 

Anthurium jenmanii Engl. [GU SU FG] 

Anthurium kunthii Poepp. [GU?] 

Anthurium lanjouwii Jonker & Jonker [SU FG] 

Anthurium maguirei A.D. Hawkes [GU SU] 

Anthurium martianum C. Koch & Kolb [GU SU] 

Anthurium pentaphyllum (Aubl.) G. Don [GU SU FG] 

Anthurium ptarianum Steyerm. [GU] 

Anthurium roraimense N.E. Brown [GU] 

Anthurium rubrinervium (Link) G. Don [GU SU FG] 

Anthurium scandens (Aubl.) Engl. [GU SU FG] 

Anthurium sinuatum Benth. ex Schott [GU SU FG] 

Anthurium thrinax Madison [GU FG] 

Anthurium trinerve Miq. [GU SU FG] 

Caladium bicolor (Aiton) Ventenat [GU SU FG] 

Caladium picturatum K. Koch & Bouché [GU FG] 

[=Caladium bicolor?] 

Caladium schomburgkii Schott [GU SU FG] 

SPERMATOPHYTA: Aquifoliaceae, Araceae 

51

52 

SPERMATOPHYTA: Araceae 


Colocasia esculenta (L.) Schott [GU SU FG] [cultivated, 

naturalized] 

Cyrtosperma spruceanum (Schott) Engl. [SU] [=Dracontium 

spruceanum] 

Dieffenbachia duidae (Steyerm.) Bunting [GU] 

Dieffenbachia elegans Jonker & Jonker [GU SU FG] 

Dieffenbachia humilis Poepp. [SU FG] 

Dieffenbachia maculata (Lodd.) G. Don [FG] [specimens not 

seen; cultivated?] 

Dieffenbachia paludicola N.E. Brown ex Gleason [GU SU 

FG] 

Dieffenbachia seguine (Jacq.) Schott var. lingulata (Mart. ex 

Schott) Engl. [SU] 

Dieffenbachia seguine (Jacq.) Schott var. seguine [GU SU 

FG] 

Dieffenbachia seguine (Jacq.) Schott var. venteniana (Schott) 

Engl. [SU] 

Dieffenbachia seguine (Jacq.) Schott var. viridis Engl. [FG] 

Dieffenbachia sp. nov. #1 [SU FG] 

Dieffenbachia sp. nov. #2 [GU] 

Dracontium asperum K. Koch [SU FG] [to be expected in GU] 

Dracontium foecundum J.D. Hook. [GU SU] 

Dracontium polyphyllum L. [SU FG] 

Heteropsis flexuosa (Kunth) Bunting [GU SU FG] 

Heteropsis longispathacea Engl. [GU?] 

Heteropsis melinonii (Engl.) Jonker & Jonker [GU SU FG] 

Heteropsis oblongata (Kunth) Croat, comb. nov. ined. [GU] 

Heteropsis spruceana Schott [GU FG] 

Heteropsis tenuispadix Bunting [GU SU FG] 

Homalomena picturata (Linden & André) Regel [GU SU FG] 

[cultivated only?] 

Jasarum steyermarkii Bunting [GU] 

Monstera adansonii Schott var. klotzschiana (Schott) Madison 

[GU SU FG] 

Monstera adansonii Schott var. laniata (Schott) Madison [GU 

SU FG] 

Monstera dubia (Kunth) Engl. & Krause [SU? FG?] 

Monstera obliqua Miq. [GU SU FG] 

Monstera spruceana (Schott) Engl. [GU SU FG] 

Montrichardia arborescens (L.) Schott [GU SU FG] 

Montrichardia linifera (Arruda) Schott [GU SU FG] 

Philodendron acutatum Schott [GU SU FG] 

Philodendron auyantepuiense Bunting [GU] 

Philodendron billietiae Croat [FG] 

Philodendron bipennifolium Schott [SU] [to be expected in 

GU] 

Philodendron brevispathum Schott [FG] 

Philodendron callosum Krause subsp. callosum [GU FG?] 

Philodendron callosum Krause subsp. ptarianum (Steyerm.) 

Bunting [GU] 

Philodendron camposportoanum Barroso [GU SU FG] 

Philodendron cremersii Croat & Grayum [FG] 

Philodendron deflexum Poepp. ex Schott [GU SU FG] 

Philodendron dioscoreoides Gleason [GU SU] 

Philodendron duckei Croat & Grayum [SU FG] 

Philodendron ecordatum Schott [GU FG] 

Philodendron englerianum Steyerm. [GU] 

Philodendron exile Bunting [SU FG] 

Philodendron fragrantissimum (Hook.) Kunth [GU SU FG] 

Philodendron goeldii Barroso [SU FG] 

Philodendron grandifolium (Jacq.) Schott [GU SU FG] 

Philodendron guianense Croat & Grayum [GU SU FG] 

Philodendron hederaceum (Jacq.) Schott [SU FG?] [to be 


Philodendron houlletianum Engl. [FG] 

Philodendron hylaeae Bunting [GU SU FG] 

Philodendron insigne Schott [GU SU FG] 

Philodendron krauseanum Steyerm. [GU SU FG] [possibly not 

distinct from P. platypodum Schott] 

Philodendron linnaei Kunth [GU SU FG] 

Philodendron martinii Schott [FG] 

Philodendron mawarinumae Bunting [SU FG] 

Philodendron melinonii Brongn. ex Regel [GU SU FG] 

Philodendron muricatum Willd. ex Schott [GU SU] 

Philodendron ornatum Schott [GU SU FG] 

Philodendron pedatum (Hook.) Kunth [GU SU FG] 

Philodendron placidum Schott [FG] [to be expected in SU] 

Philodendron platypodum Schott [GU SU FG] 

Philodendron polypodioides Jonker & Jonker [SU FG] 

Philodendron riedelianum Schott [FG] 

Philodendron roraimae Krause [GU FG?] 

Philodendron rudgeanum Schott [GU SU FG] 

Philodendron scandens K. Koch & Sello [GU SU FG] 

Philodendron simsii Kunth [GU FG] 

Philodendron solimoesense A.C. Smith [GU SU FG] 

Philodendron sphalerum Schott [GU SU FG] 

Philodendron splitgerberi Schott [SU FG] 

Philodendron squamiferum Poepp. [SU FG] 

Philodendron surinamense (Schott) Engler [GU SU FG] 

Philodendron tatei Krause [GU] 

Philodendron wittianum Engl. [FG] 

Pistia stratiotes L. [GU SU FG] 

Rhodospatha brachypoda Bunting [GU FG?] 

Rhodospatha latifolia Poepp. [GU FG] 

Rhodospatha oblongata Poepp. [GU SU FG] 

Rhodospatha venosa Gleason [GU SU FG] 

Schismatoglottis americana Jonker & Jonker [SU FG] 

Spathiphyllum cannifolium (Dryand. ex Sims) Schott [GU] 

Spathiphyllum cuspidatum Schott [GU SU FG?] 

Spathiphyllum humboldtii Schott [GU SU FG] 

Spathiphyllum maguirei Bunting [GU] [=Spathiphyllum 

cuspidatum?] 

Spathiphyllum schomburgkii Schott [GU] 

Stenospermation ammiticum Bunting subsp. ammiticum [GU] 

Stenospermation maguirei Jonker & Jonker [GU SU] 

Stenospermation multiovulatum (Engl.) N.E. Brown [GU SU 

FG] 

Stenospermation spruceanum Schott [GU SU] 

Syngonium hastifolium Engl. [SU FG] 

Syngonium podophyllum Schott var. vellozianum (Schott) 

Croat [GU SU FG] 

Typhonium blumei Nicolson & Sivadasan [SU] [naturalized] 

Urospatha sagittifolia (Rudge) Schott var. sagittifolia [GU SU 

FG] 

Urospatha sagittifolia (Rudge) Schott var. spruceana (Schott) 

Engl. [GU SU] 

Xanthosoma belophyllum (Willd.) Schott [GU SU FG] 

[cultivated?] 

Xanthosoma caracu K. Koch & Bouché [SU?] [cultivated?] 

Xanthosoma conspurcatum Schott [SU FG] 

Xanthosoma cordatum N.E. Brown [GU FG] 

Xanthosoma granvillei Croat & Thompson [FG] 

Xanthosoma helleborifolium (Jacq.) Schott [GU SU] 

Xanthosoma striatipes (Kunth) Madison [GU FG] 

Xanthosoma striolatum Mart. ex Schott [FG] 

Xanthosoma undipes (K. Koch) K. Koch [GU SU FG] 

Xanthosoma violaceum Schott [FG] [widely cultivated]

ARACEAE - SYNONYMS AND 



Anthurium amoenum Kunth [not in Guianas] 

Anthurium andersonii Schott [not in Guianas] 

Anthurium atropurpureum R.E. Schultes & Maguire [not in 


Anthurium cordatum G. Don [not in Guianas] 

Anthurium crassinervium (Jacq.) Schott [not in Guianas] 

Anthurium digitatum (Jacq.) G. Don [not in Guianas] 

Anthurium galeottii (Hort.) C. Koch [not in Guianas] 

Anthurium huegelii Schott =Anthurium hookeri 

Anthurium martini Schott =Anthurium sinuatum 

Anthurium nigrescens Engl. [not in Guianas] 

Anthurium nymphaeifolium K. Koch & Bouché [not in 


Anthurium pentaphyllum (Aubl.) G. Don var. digitatum 

(Jacq.) Madison =Anthurium digitatum [not in Guianas] 

Anthurium polyrrhizum C. Koch & Augustin =Anthurium 

rubrinervium 

Anthurium salviniae Hemsley [not in Guianas] 

Anthurium scolopendrinum (Ham.) Kunth =Anthurium gracile 

Anthurium solitarium Schott [not in Guianas] 

Anthurium stahelii A.D. Hawkes =Philodendron deflexum 

Caladium striatipes (Kunth) Schott =Xanthosoma striatipes 

Cyrtosperma americanum Engl. =Anaphyllopsis americanum 

Dieffenbachia bolivarana Bunting =Dieffenbachia duidae 

Dieffenbachia picta Schott =Dieffenbachia seguine 

Dieffenbachia spruceana Schott =Dieffenbachia humilis 

Dracontium pentaphyllum Aubl. =Anthurium pentaphyllum 

Heteropsis jenmanii Oliver =Heteropsis flexuosa 

Heteropsis salicifolia Kunth [not in Guianas] 

Maguirea spathicarpoides A.D. Hawkes =Dieffenbachia 

paludicola 

Monstera epipremnoides Engl. [not in Guianas] 

Monstera expilata Schott =Monstera obliqua 

Monstera falcifolia Engl. =Monstera obliqua 

Monstera lechleriana Schott [not in Guianas] 

Monstera milleriana Schott =Monstera adansonii 

Monstera pertusa (L.) de Vriese =Monstera adansonii 

Monstera sagotiana Engl. =Monstera obliqua 

Monstera xanthospatha Madison [not in Guianas] 

Philodendron broadwayi N.E. Brown =Philodendron venosum 

(Willd.) Croat [not in Guianas] 

Philodendron cyclops A.D. Hawkes =Philodendron acutatum 

Philodendron demerarae Gleason =Philodendron 

fragrantissimum 

Philodendron grandipes K. Krause [not in Guianas] 

Philodendron guttiferum Kunth [not in Guianas] 

Philodendron guttiferum Kunth var. rudgeanum (Schott) 

Jonker & Jonker =Philodendron rudgeanum 

Philodendron jacquinii Schott =Philodendron hederaceum 

Philodendron jenmanii Krause =Philodendron muricatum 

Philodendron laciniatum (Vell.) Engl. =Philodendron pedatum 

Philodendron longipetiolatum Engl. =Philodendron sphalerum 

Philodendron megalophyllum Schott =Philodendron deflexum 

Philodendron myrmecophilum Engl. =Philodendron deflexum 

Philodendron nobile Bull. =Philodendron linnaei 

Philodendron scabrum K. Krause =Philodendron muricatum 

Philodendron tessmannii K. Krause =Philodendron guttiferum 

[not in Guianas] 

Philodendron tripartitum (Jacq.) Schott [not in Guianas] 

Philodendron wayombense Jonker & Jonker =Philodendron 

bipennifolium 

Rhodospatha spruceana (Schott) Macbr. =Stenospermation 

spruceanum 

Schismatoglottis spruceana (Schott) Bunting [not in Guianas] 

Spathiphyllum candolleanum Schott =Spathiphyllum 

humboldtii 

Spathiphyllum floribundum (Linden & André) N.E. Brown 


Spathiphyllum huberi Engl. =Spathiphyllum humboldtii 

Spathiphyllum lanceaefolium Schott [not in Guianas] 

Syngonium ternatum Gleason =Syngonium podophyllum 

Syngonium vellozianum Schott =Syngonium podophyllum 

var. vellozianum 

Typhonium trilobatum (L.) Schott [misidentification of T. 

blumei] 

Xanthosoma hoffmannii (Schott) Schott var. wendlandii 

(Schott) Engl. [not in Guianas] 

Xanthosoma jacquinii Schott =Xanthosoma undipes 

ARALIACEAE 



REFERENCES: 

Frodin, D.G. 1993. Studies in Schefflera, VI. 

New species and subordinate taxa in the 

Venezuelan Guayana and immediately 

adjacent areas. Novon 3: 367-403. 

Maguire, B., J.A. Steyermark, & D.G. Frodin. 

1984. Araliaceae. In B. Maguire & 

collaborators, Botany of the Guayana 

Highland Part XII. Mem. New York Bot. 

Gard. 38: 1-84. 

Muller, F.M. 1934. Araliaceae. In A. Pulle, 


Dendropanax arboreus (L.) Decne. & Planch. [GU] 

Oreopanax capitatus (Jacq.) Decne. & Planch. var. capitatus 

[GU FG] 

Schefflera ayangannensis Maguire, Steyerm. & Frodin [GU] 

Schefflera chimantensis (Steyerm. & Maguire) Maguire, 

Steyerm. & Frodin subsp. rugosifolia (Maguire, Steyerm. 

& Frodin) Frodin [GU] [Mt. Roraima] 

Schefflera coriacea (Él. Marchal) Harms [GU] 

Schefflera decaphylla (Seem.) Harms [GU SU FG] 

Schefflera dissidens Frodin [GU] [Mt. Roraima] 

Schefflera duidae Steyerm. subsp. duidae [to be expected in 

GU] 

Schefflera monosperma Maguire, Steyerm. & Frodin [GU] 

[Mt. Roraima] 

Schefflera morototoni (Aubl.) Maguire, Steyerm. & Frodin 

[GU SU FG] 

Schefflera psilophylla (Harms) Maguire, Steyerm. & Frodin 

[Mt. Roraima] 

Schefflera quinquecarinata Steyerm. [GU] 

SPERMATOPHYTA: Araceae, Araliaceae 

53

54 


Schefflera sessiliflora Splithof-Heerschop ex Frodin [GU] 

Schefflera splendens (Kunth) Frodin ex Lindeman [SU FG] 

Schefflera umbellata (N.E. Brown) Viguier [GU] [Mt. 

Roraima] 

ARALIACEAE - SYNONYMS AND 


Crepinella gracilis Él. Marchal =Schefflera monosperma 

Didymopanax morototoni (Aubl.) Decne. & Planch. 

=Schefflera morototoni 

Didymopanax rugosum N.E. Brown =Schefflera chimantensis 

subsp. rugosifolia 

Oreopanax capitatus (Jacq.) Decne. & Planch. var. multiflorus 

(DC.) Él. Marchal =Oreopanax capitatus var. capitatus 

Panax morototoni Aubl. =Schefflera morototoni 

Schefflera paraensis Huber ex Ducke =Schefflera decaphylla 

Schefflera reticulata (Gleason) Mag., Steyerm. & Frodin, non 

Philipson =Schefflera sessiliflora 

Schefflera rugosifolia Maguire, Steyerm. & Frodin 

=Schefflera chimantensis subsp. rugosifolia 

Sciadophyllum coriaceum Él. Marchal =Schefflera coriacea 

Sciadophyllum umbellatum N.E. Brown =Schefflera umbellata 

ARECACEAE 

Jessenia & Oenocarpus by M. Balick; reviewed 

by S.F. Glassman & J.-J. de Granville (1992); 

reviewed by A. Henderson & J.-J. de 

Granville (1996) 

REFERENCES: 

Balick, M.J. 1986. Systematics and economic 

botany of the Oenocarpus-Jessenia (Palmae) 

complex. Advances in Economic Botany 3: 

1-140. The New York Botanical Garden, 

New York. 

Dahlgren, B.E. 1936. Index of American Palms. 

Field Museum of Natural History, Botanical 

Series, 14: 1-436. 

Dransfield, J., D. Johnson, & H. Synge. 1988. 

The Palms of the New World, A Conservation 

Census. IUCN: Kew, Richmond, Surrey. 

Glassman, S.F. 1972. A revision of B.E. 

Dahlgren's Index of American Palms. 

Phanerogamarum Monographiae Tomus VI: 

1-294. 

Glassman, S.F. 1977. Preliminary taxonomic 

studies in the palm genus Attalea H.B.K. 

Fieldiana Bot. 38: 31-61. 

Glassman, S.F. 1977. Preliminary studies in the 

palm genus Orbignya Mart. Phytologia 36: 

89-115. 

Glassman, S.F. 1978. Preliminary taxonomic 

studies in the palm genus Maximiliana Mart. 

Phytologia 38: 161-172. 

Glassman, S.F. 1987. Revisions of the palm 

genus Syagrus Mart. and other selected genera 

in the Cocos alliance. Ill. Biol. Monogr. 56: 

1-230. 

SPERMATOPHYTA: Araliaceae, Arecaceae 

Granville, J.-J. de. 1975. Un nouveau palmier en 

Guyane Française: Geonoma oldemanii. 

Adansonia, sér. 2 14(4): 553-559. 

Granville, J.-J. de. 1986. The Palms of the 

Guianas. Flora of the Guianas Newsletter 3: 

36-39. 

Granville, J.-J. de. 1989. La distribucion de las 

palmas en Guyana Francesa. Acta Amaz. 19: 

115-138. 

Granville, J.-J. de. 1990. Les Palmiers de la 

Guyane Française. Bois et Forêts des 

Tropiques 220: 43-54. 

Granville, J.-J. de. 1992. The genus Bactris: 

taxonomy and distribution. Flora of the 

Guianas Newsletter 9: 6-11. 

Granville, J.-J. de. 1996 (in press). Arecaceae. 

In S.A. Mori et al., Vascular Plant Flora of 

Central French Guiana. The New York 

Botanical Garden, New York. 

Granville, J.-J. de & A. Henderson. 1988. A new 

species of Asterogyne (Palmae) from French 

Guiana. Studies on the Flora of the Guianas 

28. Brittonia 40(1): 76-80. 

Granville, J.-J. de & A. Henderson. 1994. A new 

species of Bactris (Palmae) from the Amazon 

region. Studies on the Flora of the Guianas 

78. Brittonia 46(2): 147-150. 

Henderson, A. 1990. Arecaceae part I. 

Introduction and Iriarteinae. Flora Neotropica 

Monogr. 53: 1-100. 

Henderson, A. 1994. The genus Attalea. Palm 

Journal 114: 49-53. 

Henderson, A. 1995. The Palms of the Amazon. 

Oxford University Press. 

Kahn, F. & J.-J. de Granville. 1992. Palms in 

Forest Ecosystems of Amazonia. Ecological 

Studies 95: 1-226. Springer-Verlag. 

Kahn, F. & B. Millan. 1992. Astrocaryum 

(Palmae) in Amazonia. A preliminary 

treatment. Bull. Inst. fr. études andines 21(2): 

459-531. 

Lindeman, J.C. 1984. Palmae (additions and 

corrections). In A.L. Stoffers & J.C. 

Lindeman, Flora of Suriname 5(1): 651-652. 

Wessels Boer, J.G. 1965. Palmae. In J. 

Lanjouw, Flora of Suriname 5(1): 1-172. 

Wessels Boer, J.G. 1968. The Geonomoid 

Palms. N.V. Noord-Hollandsche Uitgevers 

Maatschappij, Amsterdam. 

Wessels Boer, J.G. 1972. Palmae. In B. 



Bot. Gard. 23: 1-832. 

[status of names according to Henderson (1995) in brackets] 

Acrocomia lasiospatha Mart. [GU SU FG] [=Acrocomia 

aculeata] 

Asterogyne guianensis Granville & Henderson [FG] 

Astrocaryum acaule Mart. [GU] 

Astrocaryum aculeatum G. Mey. [GU SU]


Astrocaryum farinosum Barb. Rodr. [GU] [=Astrocaryum 

sciophilum] 

Astrocaryum gynacanthum Mart. [GU SU FG] 

Astrocaryum jauari Mart. [GU SU FG] 

Astrocaryum minus Trail [FG] 

Astrocaryum murumuru Mart. [FG] 

Astrocaryum paramaca Mart. [GU SU FG] 

Astrocaryum rodriguesii Trail [SU FG] [=Astrocaryum 

paramaca] 

Astrocaryum sciophilum (Miq.) Pulle [SU FG] 

Astrocaryum vulgare Mart. [GU SU FG] 

Bactris acanthocarpa Mart. var. intermedia A. Henderson [GU] 

Bactris acanthocarpoides Barb. Rodr. [GU? SU FG] 

Bactris aubletiana Trail [SU FG] 

Bactris balanophora Spruce [GU?] [not in Guianas, specimens 

= B. ptariana according to Granville] 

Bactris brongniartii Mart. [GU SU FG] [frequently confused 

with Bactris maraja] 

Bactris campestris Poepp. ex Mart. [GU SU FG] 

Bactris capinensis Huber [FG] [=Bactris tomentosa var. 

tomentosa] 

Bactris constanciae Barb. Rodr. [GU SU FG] 

Bactris cruegeriana Griseb. & H. Wendl. ex Griseb. [GU SU] 

[not in Guianas according to Henderson] 

Bactris elegans Barb. Rodr. & Trail [GU SU FG] 

Bactris floccosa Spruce [SU FG] [=Bactris cuspidata Mart.] 

Bactris gastoniana Barb. Rodr. [SU FG] 

Bactris gaviona (Trail) Trail ex Drude [SU] [=Bactris major 

var. infesta (Mart.) Henderson] 

Bactris hirta Mart. var. hirta [GU SU FG] 

Bactris humilis (Wallace) Burret [GU SU FG] [=Bactris 

acanthocarpa var. acanthocarpa] 

Bactris major N.J. Jacq. [GU SU FG] [var. major] 

Bactris maraja Mart. [GU SU FG] [var. maraja] [see also 

Bactris brongniartii] 

Bactris oligocarpa Barb. Rodr. & Trail ex Barb. Rodr. [SU 

FG] 

Bactris oligoclada Burret [GU] 

Bactris pectinata Mart. [FG] [=Bactris hirta var. hirta] 

Bactris penicillata Barb. Rodr. [FG] [=Bactris maraja var. 

juruensis (Trail) Henderson] 

Bactris pliniana Granville & Henderson [GU SU FG] 

Bactris ptariana Steyerm. [GU] 

Bactris rhaphidacantha Wessels Boer [GU SU FG] 

Bactris simplicifrons Mart. [GU SU FG] 

Bactris sp. nov. [FG] 

Bactris sp. nov. [FG] 

Bactris sp. nov. [GU] [=Bactris vesiculifera Trail, nomen 

nudum] 

Bactris trailiana Barb. Rodr. [SU FG] 

Bactris turbinocarpa Barb. Rodr. [SU] 

Cocos nucifera L. [GU SU FG] [cultivated, naturalized] 

Desmoncus macroacanthos Mart. [GU SU FG] [=Desmoncus 

polyacanthos var. polyacanthos] 

Desmoncus orthacanthos Mart. [GU SU FG] 

Desmoncus polyacanthos Mart. [GU SU FG] [var. 

polyacanthos] 

Desmoncus sp. cf. mitis Mart. [FG] 

Desmoncus sp. cf. pycnacanthos Mart. [FG] 

Desmoncus sp. cf. riparius Spruce [SU] [Desmoncus riparius = 

Desmoncus polyacanthos var. polyacanthos] 

Dictyocaryum ptariense (Steyerm.) H.E. Moore & Steyerm. 

[GU] 

Elaeis guineensis N.J. Jacq. [GU SU FG] [cultivated, 

naturalized] 

Elaeis oleifera (Kunth) Cortés [GU SU FG] [naturalized?] 

Euterpe aurantiaca H.E. Moore [GU] [not in Guianas 

according to Henderson] 

Euterpe longebracteata Barb. Rodr. [GU] 

Euterpe oleracea Mart. [GU SU FG] 

Euterpe precatoria Mart. [GU SU FG] [var. precatoria] 

Euterpe roraimae Dammer [GU?] [Mt. Roraima] 

Geonoma appuniana Spruce [GU] 

Geonoma arundinacea Mart. [FG] [not in Guianas according 

to Henderson] 

Geonoma baculifera (Poit.) Kunth [GU SU FG] 

Geonoma deversa (Poit.) Kunth [GU SU FG] 

Geonoma euspatha Burret [GU SU FG] [=Geonoma interrupta 

var. euspatha] 

Geonoma fusca Wessels Boer [GU FG] [=Geonoma 

aspidiifolia] 

Geonoma leptospadix Trail [GU SU FG] 

Geonoma maxima (Poit.) Kunth [GU SU FG] [var. maxima] 

Geonoma oldemanii Granville [FG] 

Geonoma pinnatifrons Willd. [SU] [not in Guianas according 

to Henderson] 

Geonoma piscicauda Dammer [SU FG] [=Geonoma stricta 

var. piscicauda] [not in Guianas (Henderson)] 

Geonoma poiteauana Kunth [GU SU FG] [=Geonoma 

macrostachys var. poiteauana] 

Geonoma pycnostachys Mart. [GU? FG] [=Geonoma stricta 

var. stricta] 

Geonoma sp. nov. #1 [FG] 

Geonoma sp. nov. #2 [FG] 

Geonoma stricta (Poit.) Kunth [GU SU FG] [var. stricta] 

Geonoma umbraculiformis Wessels Boer [GU SU FG] 

Hyospathe elegans Mart. [GU SU FG] 

Iriartella setigera (Mart.) H. Wendl. [GU] 

Jessenia bataua (Mart.) Burret subsp. bataua [GU? SU? FG] 

[=Oenocarpus bataua var. bataua] [not in Guianas 

according to Henderson)] 

Jessenia bataua (Mart.) Burret subsp. oligocarpa (Griseb. & H. 

Wendl.) Balick [GU SU FG] [=Oenocarpus bataua var. 

oligocarpa] 

Lepidocaryum guianense Becc. [GU] [=Lepidocaryum tenue 

var. gracile] 

Manicaria saccifera Gaertn. [GU SU FG] 

Mauritia flexuosa L.f. [GU SU FG] 

Mauritiella martiana (Spruce) Burret [GU SU FG] 

[=Mauritiella armata] 

xMaximbignya dahlgreniana (Bondar) Glassman, comb. & 

stat. nov. ined. [SU] [=Attalea dahlgreniana] 

Maximiliana maripa (Corrêa) Drude [GU SU FG] [=Attalea 

maripa] 

Oenocarpus bacaba Mart. [GU SU FG] 

Orbignya phalerata Mart. [GU SU] [=Attalea speciosa] 

Orbignya polysticha Burret [FG] [=Attalea microcarpa] 

Orbignya sagotii Trail ex Im Thurn [GU SU FG] [=Attalea 

microcarpa] 

Phoenix reclinata Jacq. [GU SU FG] [cultivated, naturalized] 

Prestoea tenuiramosa (Dammer) H.E. Moore [GU] 

Roystonea oleracea (N.J. Jacq.) O.F. Cook [GU SU FG] 


Scheelea camopiensis Glassman, sp. nov. ined. [FG] [=Attalea 

attaleoides] 

Scheelea degranvillei Glassman, sp. nov. ined. [FG] [=Attalea 

attaleoides] 

Scheelea guianensis Glassman, sp. nov. ined. [FG] [=Attalea 

attaleoides] 

SPERMATOPHYTA: Arecaceae 

55

56 

SPERMATOPHYTA: Arecaceae 


Scheelea maripensis Glassman, sp. nov. ined. [FG] [=Attalea 

attaleoides] 

Scheelea sp. cf. macrolepis Burret [FG] [=Attalea attaleoides] 

Socratea exorrhiza (Mart.) H. Wendl. [GU SU FG] 

Syagrus cocoides Mart. [GU] 

Syagrus inajai (Spruce) Beccari [SU FG] 

Syagrus stratincola Wessels Boer [GU SU FG] 

ARECACEAE - SYNONYMS AND 


Acrocomia aculeata (Jacq.) Lodd. ex Mart. [see Acrocomia 

lasiospatha] 

Acrocomia sclerocarpa Mart. =Acrocomia lasiospatha 

[=Acrocomia aculeata s.l.] 

Acrocomia wallaceana (Drude) Becc. =Acrocomia lasiospatha 

[=Acrocomia aculeata s.l.] 

Asterogyne spicata (H.E. Moore) Wessels Boer [not in 


Astrocaryum caudescens Barb. Rodr. =Astrocaryum 

aculeatum 

Astrocaryum guianense Splitg. ex Mart. =Astrocaryum vulgare 

Astrocaryum munbaca Mart. =Astrocaryum gynacanthum 

Astrocaryum plicatum Drude =Astrocaryum sciophilum 

Astrocaryum segregatum Drude =Astrocaryum vulgare 

Astrocaryum tucuma Mart. =Astrocaryum aculeatum 

Astrocaryum tucumoides Drude =Astrocaryum vulgare 

Attalea attaleoides (Barb. Rodr.) Wessels Boer =Scheelea 

sp.[non Scheelea attaleoides Karst.] 

Attalea dahlgreniana (Bondar) Wessels Boer =xMaximbignya 

dahlgreniana 

Attalea excelsa Mart. ex Spreng. =Scheelea martiana Burret 

[dubious species] 

Attalea funifera Mart. [not in Guianas] 

Attalea macropetala (Burret) Wessels Boer =Maximiliana 

maripa 

Attalea maripa (Corrêa) Mart. =Maximiliana maripa 

Attalea microcarpa Mart. [see Orbignya polysticha, Orbignya 

sagotii] 

Attalea monosperma Barb. Rodr. =Orbignya spectabilis 

(Mart.) Burret [not in Guianas] 

Attalea regia (Mart.) Wessels Boer =Maximiliana maripa 

Attalea sagotii (Trail ex Im Thurn) Wessels Boer =Orbignya 

sagotii 

Attalea speciosa Mart. [see Orbignya phalerata] 

Attalea spectabilis Mart. =Orbignya spectabilis (Mart.) Burret 


Bactris armata Barb. Rodr. [not in Guianas] 

Bactris concinna Mart. [not in Guianas] 

Bactris corossilla Karsten [not in Guianas] 

Bactris cuspidata Mart. var. tenuis (Wallace) Drude =Bactris 

simplicifrons 

Bactris exscapa Barb. Rodr. =Bactris humilis 

Bactris geonomoides Drude =Bactris hirta var. hirta 

Bactris integrifolia Wallace =Bactris hirta var. hirta 

Bactris leptocarpa Trail ex Im Thurn =Bactris campestris 

Bactris maraja Mart. var. juruensis (Trail) Henderson [see 

Bactris penicillata] 

Bactris megalocarpa Trail ex Im Thurn =Bactris cruegeriana 

Bactris mitis Mart. [not in Guianas] 

Bactris monticola Barb. Rodr. =Bactris maraja 

Bactris pallidispina Mart. =Bactris brongniartii 

Bactris rivularis Barb. Rodr. =Bactris brongniartii 

Bactris sphaerocarpa Trail [not in Guianas] 

Bactris tenuis Wallace =Bactris simplicifrons 

Bactris trichospatha Trail =Bactris maraja 

Bactris ulei Burret =Bactris simplicifrons 

Chamaedorea gracilis Willd. [name of uncertain application] 

Copernicia cerifera (Arruda da Camara) Mart. =Copernicia 

prunifera (Miller) Moore [cultivated only] 

Desmoncus guianensis Trail [apparently a nomen nudum] 

Desmoncus horridus Splitg. ex Mart. =Desmoncus 

orthacanthos 

Desmoncus kaieteurensis L.H. Bailey =Desmoncus 

macroacanthos 

Desmoncus maguirei L.H. Bailey =Desmoncus polyacanthos 

Desmoncus oxyacanthos Mart. =Desmoncus polyacanthos 

Desmoncus palustris Trail =Desmoncus orthacanthos 

Desmoncus parvulus L.H. Bailey =Desmoncus macroacanthos 

Dictyocaryum fuscum (Kersten) H. Wendl. [not in Guianas] 

Euterpe stenophylla Trail ex Burret =Euterpe precatoria 

Euterpe tenuiramosa Dammer =Prestoea tenuiramosa 

Geonoma acaulis (Poit.) Burret =Geonoma poiteauana 

Geonoma acaulis Mart. [not in Guianas] 

Geonoma ambigua Spruce =Geonoma maxima 

Geonoma aspidiifolia Spruce [see Geonoma fusca] 

Geonoma barthia Engel =Geonoma densa 

Geonoma bartlettii Dammer ex Burret =Geonoma deversa 

Geonoma densa Linden & H. Wendl. ex H. Wendl. [not in 


Geonoma elegans Mart. [not in Guianas] 

Geonoma interrupta (Ruiz & Pav.) Mart. var. euspatha 

(Burret) Henderson =Geonoma euspatha 

Geonoma macrostachys Mart. var. poiteauana (Kunth) 

Henderson =Geonoma poiteauana 

Geonoma maguirei L.H. Bailey =Geonoma stricta 

Geonoma multiflora Mart. [not in Guianas] 

Geonoma paniculigera Mart. =Geonoma deversa 

Geonoma platycaula Drude & Trail ex Drude =Geonoma 

rubescens H. Wendl. ex Drude [not in Guianas] 

Geonoma robusta Burret =Geonoma maxima 

Geonoma saramaccana L.H. Bailey =Geonoma leptospadix 

Geonoma schomburgkiana Spruce =Geonoma maxima 

Geonoma spruceana Trail =Geonoma maxima 

Geonoma stricta (Poit.) Kunth var. piscicauda (Dammer) 

Henderson =Geonoma piscicauda 

Geonoma tamandua Trail [not in Guianas] [=Geonoma 

macrostachys var. macrostachys] 

Geonoma triglochin Burret [not in Guianas] 

Guilielma speciosa Mart. =Bactris gasipaes Kunth [cultivated 

only] 

Iriartea exorrhiza Mart. =Socratea exorrhiza 

Iriartea fusca (Karst.) Drude =Dictyocaryum fuscum [not in 


Jessenia oligocarpa Griseb. & H. Wendl. =Jessenia bataua 

subsp. oligocarpa 

Lepidocaryum tenue Mart. var. gracile (Mart.) Henderson [see 

Lepidocaryum guianense] 

Markleya dahlgreniana Bondar =xMaximbignya dahlgreniana 

Mauritia martiana Spruce =Mauritiella martiana 

Mauritiella aculeata (Kunth) Burret [not in Guianas] 

Mauritiella armata (Mart.) Burret [see Mauritiella martiana] 

Maximiliana macropetala Burret =Maximiliana maripa 

Maximiliana martiana Karsten =Maximiliana maripa 

Maximiliana regia Mart. =Maximiliana maripa


Oenocarpus bataua Mart. =Jessenia bataua subsp. bataua 

Oenocarpus bataua Mart. var. oligocarpa (Griseb. & H. 

Wendl.) Henderson =Jessenia bataua subsp. oligocarpa 

Oenocarpus oligocarpa (Griseb. & H. Wendl.) Wessels Boer 

=Jessenia bataua subsp. oligocarpa 

Orbignya barbosiana Burret =Orbignya speciosa (Mart.) Barb. 

Rodr.[not in Guianas] 

Orbignya speciosa (Mart.) Barb. Rodr. [not in Guianas] 

[=Attalea speciosa] 

Orbignya spectabilis (Mart.) Burret [not in Guianas] [=Attalea 

spectabilis Mart.] 

Raphia taedigera (Mart.) Mart. [not in Guianas] 

Roystonea jenmanii (Wright) Burret =Roystonea regia (Kunth) 

Cook [cultivated] 

Syagrus sancona Karsten [not in Guianas] 

Yuyba dakamana L.H. Bailey =Bactris simplicifrons 

Yuyba essequiboensis L.H. Bailey =Bactris simplicifrons 

Yuyba gleasonii L.H. Bailey =Bactris simplicifrons 

Yuyba maguirei L.H. Bailey =Bactris simplicifrons 

Yuyba stahelii L.H. Bailey =Bactris simplicifrons 

ARISTOLOCHIACEAE 

contributed by C. Feuillet & O. Poncy (1992); 

additions and corrections by C. Feuillet 

(1996) 

REFERENCES: 

Hoehne, F.C. 1942. Aristolochiaceas. In F.C. 

Hoehne, Flora Brasílica 14(2): 1-141. 

Poncy, O. 1988. Quatre espèces nouvelles 

d'Aristolochia de Guyane françcaise, et 

remarques nomenclaturales concernant une 

espèce d'Aublet. Bull. Mus. Hist. Nat. Paris, 

sér. 4, 10 (sect. B), Adansonia 4: 337-344. 

Aristolochia amara (Aubl.) O. Poncy [FG] [based on sterile 

material, of uncertain application] 

Aristolochia bukuti O. Poncy [SU FG] 

Aristolochia consimilis Mast. [GU] 

Aristolochia cremersii O. Poncy [FG] 

Aristolochia daemoninoxia Mast. [GU] 

Aristolochia didyma S. Moore [FG] 

Aristolochia flava O. Poncy [FG] 

Aristolochia guianensis O. Poncy [SU FG] 

Aristolochia hians Willd. [GU] 

Aristolochia iquitensis O.C. Schmidt [GU SU FG] 

Aristolochia leprieurii Duch. [FG] 

Aristolochia mossii S. Moore [FG] 

Aristolochia paramaribensis Duch. [GU SU FG] 

Aristolochia peltatodeltoidea Hoehne [GU] 

Aristolochia rugosa Lam. [GU FG] 

Aristolochia stahelii O.C. Schmidt [SU FG] 

Aristolochia surinamensis Willd. [GU SU] 

Aristolochia trilobata L. [GU SU FG] 

Aristolochia weddellii Duch. var. rondoniana Hoehne [GU SU 

FG] 

ARISTOLOCHIACEAE - SYNONYMS AND 


Aristolochia barbata Jacq. =Aristolochia rugosa 

Aristolochia cornuta Masters [not in Guianas] 

Aristolochia dictyantha Duch. var. schomburgkii (Kl.) Duch. 

=Aristolochia rugosa 

Aristolochia dubia Hoehne =Aristolochia mossii 

Aristolochia gabrielis Briq. =Aristolochia leprieurii 

Aristolochia glaucescens Kunth [not in Guianas] 

Aristolochia longecaudata Mast. =Aristolochia weddellii var. 

rondoniana 

Aristolochia macrophylla Duch. =Aristolochia iquitensis 

Aristolochia macrota Duch. =Aristolochia surinamensis 

Aristolochia maxima L. [not in Guianas] 

Aristolochia platyloba Garcke =Aristolochia surinamensis 

ASCLEPIADACEAE 

contributed by G. Morillo (1992) 

REFERENCES: 

Fontella Pereira, J. 1977. Revisao taxonômica 

do gênero Tassadia Decaisne. Arq. Jard. Bot. 

Rio de Janeiro 21: 235-392. 

Jonker, F.P. 1940. Asclepiadaceae. In A. Pulle, 


Morillo, G. 1986. Asclepiadaceae II. Especies 

nuevas o interesantes en Venezuela y Guyana. 

Ernstia 37: 1-13. 

Morillo, G. 1988. Asclepiadaceae III, one new 

species and four new records. Proc. Kon. 

Ned. Akad. Wet. ser. C, 91(2): 161-164. 

Morillo, G. 1989. Contribución al conocimiento 

de las Asclepiadaceae suramericanas, 

principalmente de las Guayanas. Ernstia 51: 

2-15. 

Morillo, G. 1989. Veinte Asclepiadaceae 

sudamericanas nuevas para la ciencia y una 

nueva combinación. An. Jard. Bot. Madrid 

47(2): 349-359. 

Morillo, G. 1991. Once Asclepiadaceae 

suramericanas nuevas para la ciencia. Ernstia 

1(3): 109-120. 

Morillo, G. 1993. Asclepiadaceae nuevas o 

interesantes de Venezuela y Guyana. Ernstia 

3(2): 61-70. 

Morillo, G. 1994. Seis Asclepiadaceae 

Sudamericanas nuevas para la ciencia. 

Ernstia 4(1-2): 3-10. 

Asclepias curassavica L. [GU SU FG] 

Asclepias fruticosa L. [SU] 

Barjonia erecta (Vell.) K. Schum. [SU] 

Blepharodon glaucescens (Decne.) Fontella [GU] 

Blepharodon grandiflorum Benth. subsp. crassifolius 

(Schlechter) Morillo [GU] 

Blepharodon julianii Morillo [to be expected in GU] 

Blepharodon nitidus (Vell.) Macbr. [GU SU FG] 

Blepharodon tillettii Morillo [GU] 

SPERMATOPHYTA: Arecaceae - Asclepiadaceae 

57

58 


Blepharodon ulei Schlechter [GU] 

Calotropis gigantea (L.) Aiton [GU SU] [cultivated, 

naturalized] 

Cynanchum blandum (Decne.) Sundell [GU SU FG] 

Cynanchum görtsianum Morillo [FG] 

Cynanchum hirtellum (Oliver) Badillo [GU] 

Cynanchum jenmanii Morillo [GU] 

Cynanchum montevidense Spreng. [GU] 

Cynanchum parviflorum Swartz [GU] 

Cynanchum prevostiae Morillo [FG] 

Ditassa angustifolia Decne. [GU] 

Ditassa ayangannaensis Morillo [GU] 

Ditassa bolivarensis (R. Holm) Morillo [to be expected in GU] 

Ditassa consanguinea Decne. [GU SU FG] 

Ditassa gillespieae Morillo [GU] 

Ditassa lindemanii Morillo [SU] 

Ditassa obovata (R. Holm) Morillo [to be expected in GU] 

Ditassa pauciflora Decne. [GU] 

Ditassa roraimensis Schlechter [GU] 

Ditassa surinamensis Morillo [SU] 

Ditassa tatei Gleason & Moldenke [GU] 

Ditassa taxifolia Decne. [GU] 

Ditassa venamensis Morillo [GU] 

Fischeria stellata (Vell.) Fourn. [GU] 

Gonolobus germanianus Morillo [to be expected in GU] 

Gonolobus grandiflorus R. Brown [GU?] [specimens not seen, 

status uncertain] 

Gonolobus tobagensis Urb. [GU] 

Macroditassa carolina Morillo [GU] [Mt. Roraima] 

Macroditassa laurifolia (Decne.) Fontella [GU] 

Macroscepis hirsuta (Vahl) Schlecht. [GU] 

Marsdenia altissima (Jacq.) Dugand [GU SU FG] 

Marsdenia gillespieae Morillo [GU] 

Marsdenia macrophylla (Kunth) Fourn. [GU SU] 

Marsdenia rubrofusca Fourn. [GU SU FG] 

Marsdenia weddellii (Fourn.) Malme [SU] 

Matelea badilloi Morillo [GU FG] 

Matelea bolivarensis Morillo [GU] 

Matelea cayennensis Morillo [FG] 

Matelea coriacea Morillo [to be expected in GU] 

Matelea cremersii Morillo [FG] 

Matelea delascioi Morillo [GU] 

Matelea denticulata (Vahl) Fontella & Schwarz [GU SU FG] 

Matelea funkiana Morillo [GU] 

Matelea glaziovii (Fourn.) Morillo [SU] 

Matelea grenandii Morillo [FG] 

Matelea hoffmanii Morillo [GU] 

Matelea jansen-jacobsiae Morillo [SU] 

Matelea lourteigiae Morillo [GU SU FG] 

Matelea maritima (Jacq.) Woodson [GU] 

Matelea oldemanii Morillo [FG] 

Matelea palustris Aubl. [GU SU FG] 

Matelea planiflora (Jacq.) Dugand [GU] 

Matelea riparia Morillo [GU] 

Matelea sastrei Morillo [SU FG] 

Matelea squiresii (Rusby) Morillo [GU SU] 

Matelea stenopetala Sandw. [GU FG] 

Matelea surinamensis Morillo [SU] 

Metalepis albiflora Urb. [FG] 

Metalepis prevostiae Morillo [FG] 

Nephradenia linearis Benth. ex Fourn. [GU] 

Oxypetalum capitatum Mart. & Zucc. [GU] 

Sarcostemma clausum (Jacq.) Roem. & Schult. [GU SU FG] 

Stenomeria decalepis Turcz. [GU] 

SPERMATOPHYTA: Asclepiadaceae, Asteraceae 

Tassadia aristata (Benth. ex Fourn.) Fontella [to be expected in 

the Guianas] 

Tassadia berteriana (Spreng.) Stevens [GU SU] 

Tassadia decaisneana Miq. [GU SU] 

Tassadia guianensis Decne. [GU SU FG] 

Tassadia leptobotrys Decne. [SU] 

Tassadia martiana Decne. [to be expected in the Guianas] 

Tassadia obovata Decne. [GU SU FG] 

Tassadia ovalifolia (Fourn.) Fontella [to be expected in the 


Tassadia propinqua Decne. [GU SU] [to be expected in FG] 

Tassadia trailiana (Benth.) Fontella [GU] 

ASCLEPIADACEAE - SYNONYMS AND 


Funastrum clausum (Jacq.) Schlechter =Sarcostemma clausum 

Gonolobus ligustrinus Decne. [not in Guianas] 

Gonolobus riparius Kunth [not in Guianas] 

Gonolobus surinamensis Jonk. =Matelea lourteigiae 

Gonolobus viridiflorus Roem. & Schult. =Matelea denticulata 

Marsdenia maculata Hook. =Marsdenia macrophylla 

Marsdenia mollissima Fourn. =Marsdenia altissima 

Matelea hirsuta (Vahl) Woodson =Macroscepis hirsuta 

Matelea latifolia Aubl. =Matelea palustris 

Metastelma brasiliense K. Schum. [not in Guianas] 

Metastelma decaisneana Benth. & Hook. =Tassadia 

decaisneana 

Metastelma stenolobum Decne. =Tassadia berteriana 

Roulinia blanda Decne. =Cynanchum blandum 

Rouliniella guianensis (Decne.) Jonker =Cynanchum blandum 

Telminostelma corymbosum (Decne.) Fontella & Schwarz 

=Cynanchum blandum 

ASTERACEAE 

contributed by V.A. Funk, H. Robinson, & J. 

Pruski (1992); additions and corrections by 

V.A. Funk, H. Robinson, & J. Pruski (1996) 

REFERENCES: 

Aristeguieta, L. 1964. Compositae. In T. Lasser, 


Funk, V.A. 1991. The Compositae of the 

Guianas, I: Heliantheae (Heliantheae, 

Tageteae, Coreopsideae). Rhodora 93(875): 

256-267. 

Koster, J.T. 1938. Compositae. In A. Pulle, 


Pruski, J.F. 1991. Compositae of the Guayana 

Highland - V. The Mutisieae of the lost world 

of Brazil, Colombia, and Guyana. Bol. Mus. 

Paraense Emílio Goeldi (Bot.) 7: 335-392. 

Robinson, H. 1992. The Asteraceae of the 

Guianas, III: Vernonieae and restoration of 

the genus Xiphochaeta. Rhodora 94(880): 

348-361.


Acanthospermum australe (Loefl.) Kuntze [GU SU FG] 

Acanthospermum hispidum DC. [GU SU FG] 

Achyrocline flaccida DC. [GU] 

Achyrocline satureioides (Lam.) DC. [GU] 

Achyrocline vargasiana DC. [GU] 

Acmella brachyglossa Cass. [SU FG] 

Acmella ciliata (Kunth) Cass. [GU FG?] 

Acmella radicans (Jacq.) R.K. Jansen var. debilis (Kunth) R.K. 

Jansen [to be expected in the Guianas] 

Acmella repens (Walt.) L.C. Rich. ex Pers. [FG] 

Acmella uliginosa (Swartz) Cass. [GU SU FG] 

Ageratina ibaguensis (Schultz Bip. ex Hieron.) R. King & H. 

Robinson [to be expected in the Guianas] 

Ageratina roraimensis (N.E. Brown) R. King & H. Robinson 

[Mt. Roraima] 

Ageratum conyzoides L. [GU SU FG] [introduced] 

Ageratum houstonianum Mill. [SU] [widely cultivated, 

possibly naturalized] 

Ageratum scorpioideum Baker [GU] 

Ambrosia crithmifolia DC. [GU] 

Ambrosia microcephala DC. [SU FG] 

Ambrosia peruviana Willd. [to be expected in the Guianas] 

Austroeupatorium inulaefolium (Kunth) R. King & H. 

Robinson [GU] 

Ayapana amygdalina (Lam.) R. King & H. Robinson [GU SU 

FG] 

Ayapana trinitensis (Kuntze) R. King & H. Robinson [to be 

expected in the Guianas] 

Ayapana triplinervis (Vahl) R. King & H. Robinson [FG] 

Baccharis brachylaenoides DC. var. brachylaenoides [GU] 

Baccharis brachylaenoides DC. var. ligustrina (DC.) Maguire 

& Wurdack [GU] 

Baccharis densa (N.E. Brown) Badillo [GU] 

Baccharis rufescens Spreng. [GU] 

Baccharis schomburgkii Baker [GU] 

Baccharis trinervis (Lam.) Pers. [to be expected in the 


Baccharis varians Gardner [GU SU] 

Baccharis vitis-idaea Oliver [GU] 

Baltimora geminata (Brandegee) Stuessy [GU] 

Bidens alba (L.) DC. var. radiata (Schultz Bip.) Ballard [GU 

FG] 

Bidens cynapiifolia Kunth [GU SU FG] 

Bidens pilosa L. [GU SU FG] 

Bidens riparia Kunth var. refracta (Brandegee) O.E. Schultz 

[GU] 

Calea berteriana DC. [to be expected in the Guianas] 

Calea caleoides (DC.) H. Robinson [GU SU] 

Calea divaricata Benth. [GU] 

Calea lucidivenia Gleason & S.F. Blake [GU] 

Calea oliveri B.L. Robinson & Greenm. [GU] 

Calea solidaginea Kunth subsp. deltophylla (Cowan) Pruski 

[GU SU FG] 

Centratherum punctatum Cass. [GU FG] 

Chaptalia integerrima (Vell.) Burkart [GU] 

Chaptalia nutans (L.) Polakowsky [GU SU] 

Chromolaena corymbosa (Aubl.) R. King & H. Robinson [FG] 

Chromolaena dichotoma Schultz Bip. [SU] 

Chromolaena ivaefolia (L.) R. King & H. Robinson [GU] 

Chromolaena laevigata (Lam.) R. King & H. Robinson [GU 

SU FG] 

Chromolaena maximilianii (Schrad. ex DC.) R. King & H. 

Robinson [FG] 

Chromolaena odorata (L.) R. King & H. Robinson [GU SU 

FG] 

Chromolaena pharcidodes (B. Robinson) R. King & H. 

Robinson [GU] 

Chromolaena squalida (DC.) R. King & H. Robinson [GU SU] 

Chromolaena thurnii (B. Robinson) R. King & H. Robinson 

[GU] 

Clibadium armani (Balbis) Schultz Bip. ex O.E. Schulz [GU 

SU FG] 

Clibadium asperum DC. [GU SU FG] 

Clibadium surinamense L. [GU SU FG] 

Clibadium sylvestre (Aubl.) Baill. [GU SU FG] 

Condylidium iresinoides (Kunth) R. King & H. Robinson 

[GU] 

Conyza bonariensis (L.) Cronquist [GU SU FG] 

Conyza canadensis (L.) Cronquist [SU FG] [naturalized] 

Conyza primulaefolia (Lam.) Cuatrec. & Lourteig [GU SU] 

Cosmos caudatus Kunth [GU SU FG] [widely cultivated, 

probably escaping] 

Cyanthillium cinereum (L.) H. Robinson [GU SU FG] 

[naturalized] 

Cyrtocymura scorpioides (Lam.) H. Robinson [GU SU FG] 

Delilia biflora (L.) Kuntze [GU] 

Eclipta prostrata (L.) L. [GU SU FG] 

Egletes commixta Shinners [GU] 

Egletes florida Shinners [to be expected in the Guianas] 

Egletes prostrata (Swartz) Kuntze [GU] 

Elephantopus mollis Kunth [GU SU FG] 

Eleutheranthera ruderalis (Swartz) Schultz Bip. [GU SU FG] 

Emilia coccinea (Sims) G. Don [cultivated, perhaps escaping] 

Emilia fosbergii Nicolson [GU SU FG] 

Emilia sonchifolia (L.) DC. ex Wight [GU SU FG] 

Erechtites hieracifolia (L.) Raf. ex DC. [GU SU FG] 

Erechtites valerianaefolia (Wolf) Link ex Spreng. [GU FG] 

Erigeron laevigatus Rich. [FG] 

Fleischmannia microstemon (Cass.) R. King & H. Robinson 

[SU FG] 

Gamochaeta simplicicaulis Willd. ex Spreng. [GU] 

Gnaphalium elegans Kunth [to be expected in GU] 

Gnaphalium polycaulon Pers. [GU] 

Gongylolepis benthamiana Rob. Schomb. [GU] 

Guayania roupalifolia (B. Robinson) R. King & H. Robinson 

[GU] 

Hebeclinium macrophyllum (L.) DC. [GU SU FG] 

Heterocondylus vitalbae (DC.) R. King & H. Robinson [SU] 

Ichthyothere davidsei H. Robinson [FG] 

Ichthyothere granvillei H. Robinson [SU? FG] 

Ichthyothere terminalis (Spreng.) S.F. Blake [GU SU FG] 

Koanophyllon fuscum (N.E. Brown) R. King & H. Robinson 

[GU] 

Lagascea mollis Cav. [to be expected in the Guianas] 

Lepidaploa bolivarensis (Badillo) H. Robinson [GU SU] 

Lepidaploa cardonae (Aristeguieta) Pruski, comb. nov. ined. 

[GU] 

Lepidaploa chrysotricha (Alexander) H. Robinson [GU] 

Lepidaploa cotoneaster (Less.) H. Robinson [GU] 

Lepidaploa ehretiifolia (Benth.) H. Robinson [GU] 

Lepidaploa gracilis (Kunth) H. Robinson [GU] 

Lepidaploa pari (Badillo) H. Robinson [to be expected in the 


Lepidaploa remotiflora (L.C. Rich.) H. Robinson [GU SU FG] 

Lepidaploa salzmanii (DC.) H. Robinson [Mt. Roraima] 

Melanthera nivea (L.) Small [GU] 

Mikania banisteriae DC. [GU SU FG] 

Mikania boomii Pruski [GU] 

Mikania chlorolepis Baker [to be expected in GU] 

Mikania congesta DC. [GU SU FG] 

SPERMATOPHYTA: Asteraceae 

59

60 



Mikania cordifolia (L.f.) Willd. [GU SU FG] 

Mikania duidensis B.L. Robinson [GU] 

Mikania gleasonii B.L. Robinson [GU SU FG] 

Mikania guaco Humb. & Bonpl. [GU SU FG] 

Mikania hookeriana DC. [GU SU FG] 

Mikania lindleyana DC. [GU SU] 

Mikania lucida S.F. Blake [Mt. Roraima] 

Mikania micrantha Kunth [GU SU FG] 

Mikania microptera DC. [GU SU FG] 

Mikania nigropunctulata Hieron. [GU SU] 

Mikania pannosa Baker [GU] 

Mikania parviflora (Aubl.) Karsten [GU SU FG] 

Mikania psilostachya DC. [GU SU FG] 

Mikania rondonensis Badillo [Mt. Roraima] 

Mikania sprucei Baker [GU] 

Mikania trinitaria DC. [GU SU FG] 

Mikania vitifolia DC. [GU SU] 

Neurolaena lobata (L.) Cass. [SU FG] 

Orthopappus angustifolius (Swartz) Gleason [GU SU FG] 

Oyedaea scaberrima (Benth.) S.F. Blake [GU] 

Oyedaea tepuiana (Badillo) Pruski, comb. nov. ined. [to be 


Pacourina edulis Aubl. [GU SU FG] 

Parthenium hysterophorus L. [GU SU FG] 

Pectis elongata Kunth var. elongata [GU SU FG] 

Pectis elongata Kunth var. floribunda (A. Rich.) Keil [FG] 

Pectis humifusa Swartz [GU] 

Pectis linifolia L. var. linifolia [to be expected in the Guianas] 

Pectis venezuelensis Steyerm. [to be expected in the Guianas] 

Pentacalia freemanii (Britt. & Greenm.) Cuatrec. [GU] 

Pentacalia ptariana (Cuatrec.) Cuatrec. [to be expected in the 


Piptocarpha leprosa (Less.) Baker [GU] 

Piptocarpha opaca (Benth.) Baker [GU] 

Piptocarpha poeppigiana (DC.) Baker [GU] 

Piptocarpha polycephala Baker [GU] 

Piptocarpha triflora (Aubl.) Bennett ex Baker [GU SU FG] 

Piptocoma roraimensis (Steyerm.) Pruski, comb. nov. ined. 

[GU] 

Piptocoma schomburgkii (Schultz Bip.) Pruski, comb. nov. 

ined. [GU SU FG] 

Pluchea carolinensis (Jacq.) G. Don [GU] [probably cultivated 

only] 

Pluchea odorata (L.) Cass. [GU SU] 

Pluchea sagittalis (Lam.) Cabrera [to be expected in the 


Porophyllum ruderale (Jacq.) Cass. [GU SU FG] 

Praxelis asperulacea (Baker) R. King & H. Robinson [GU] 

Praxelis karuaiensis (V. Badillo) R. King & H. Robinson [GU] 

Praxelis pauciflora (Kunth) R. King & H. Robinson [GU SU 

FG] 

Pseudelephantopus spicatus (Juss.) Rohr [GU SU FG] 

Pseudogynoxys chenopodioides (Kunth) Cabrera [to be 


Pterocaulon alopecuroides (Lam.) DC. [GU SU] 

Quelchia conferta N.E. Brown [GU] 

Riencourtia glomerata Cass. [SU FG] 

Riencourtia latifolia Gardn. [GU] 

Riencourtia oblongifolia Gardn. [SU] 

Riencourtia pedunculosa (Rich.) Pruski, comb. nov. ined. [FG] 

Rolandra fruticosa (L.) Kuntze [GU SU FG] 

Salmea scandens DC. [to be expected in the Guianas] 

Sciadocephala pakaraimae (Maguire & Wurdack) R. King & 

H. Robinson [GU] 

Sigesbeckia orientalis L. [GU] [naturalized] 

Sonchus asper (L.) Hill. [GU] 

Sphagneticola brachycarpa (Baker) Pruski, comb. nov. ined. 

[GU] 

Sphagneticola trilobata (L.) Pruski, comb. nov. ined. [GU SU 

FG] 

Spilanthes urens Jacq. [to be expected in the Guianas] 

Stenopadus connellii (N.E. Brown) S.F. Blake [GU] 

Stenopadus megacephalus Pruski [GU] 

Stenopadus sericeus Maguire & Arist. [GU] 

Stenopadus talaumifolius S.F. Blake [GU] 

Stifftia cayennensis H. Robinson & Kahn [FG] 

Stomatochaeta condensata (Baker) Maguire & Wurdack [GU] 

Struchium sparganophorum (L.) Kuntze [GU SU FG] 

Symphyotrichum laeve (L.) Löve & Löve [GU FG] 


Synedrella nodiflora (L.) Gaertn. [GU SU FG] 

Tagetes erecta L. [GU SU] [cultivated, naturalized] 

Trichogonia arguta (Kunth) Benth. & Hook.f. ex Klatt [Mt. 

Roraima] 

Trichospira verticillata (L.) S.F. Blake [GU FG] 

Tridax procumbens L. [GU FG] 

Unxia camphorata L.f. [GU SU FG] 

Verbesina columbiana B.L. Robinson [to be expected in the 


Verbesina ligulata (Maguire & Wurdack) Pruski, comb. nov. 

ined. [to be expected in the Guianas] 

Verbesina schomburgkii Schultz Bip. ex Klatt [GU] 

Vernonanthura brasiliana (L.) H. Robinson [GU SU FG] 

Wedelia ambigens S.F. Blake [GU] 

Wedelia calycina L.C. Rich [GU] 

Wedelia fruticosa Jacq. [GU FG] 

Wulffia baccata (L.) Kuntze [GU SU FG] 

Wulffia rubens Alexander [GU] 

Xiphochaeta aquatica Poepp. [GU SU FG] 

Youngia japonica (L.) DC. [GU] 

ASTERACEAE - SYNONYMS AND 


Acanthospermum brasilium Shrank =Acanthospermum 

australe 

Acmella oppositifolia (Lam.) R.K. Jansen var. oppositifolia 

=Acmella repens 

Acmella radicans (Jacq.) R.K. Jansen var. radicans [not in 


Adenostemma pakaraimae Maguire & Wurdack 

=Sciadocephala pakaraimae 

Ambrosia cumanensis Kunth =Ambrosia peruviana 

Aster laevis L. =Symphyotrichum laeve 

Baillieria aspera Aubl. =Clibadium surinamense 

Calea cardonae Maguire & Wurdack var. orientalis Maguire & 

Wurdack =Calea lucidivenia 

Calea prunifolia Kunth [not in Guianas] 

Calea ternifolia Oliver =Calea oliveri 

Clibadium pedunculosum (Rich.) DC. =Riencourtia 

pedunculosa 

Clibadium rotundifolium DC. =Clibadium armani 

Conyza cayennensis Spreng. [name of uncertain application] 

Conyza chilensis Spreng. =Conyza primulaefolia 

Eclipta alba (L.) Hassk. =Eclipta prostrata 

Elephantopus angustifolius Swartz =Orthopappus 

angustifolius 

Elephantopus carolinianus Willd. [not in Guianas] 

Elephantopus pilosus Philipson =Elephantopus mollis


Elephantopus tomentosus L. =Elephantopus mollis 

Elephantopus virgatus Hamilton [name of uncertain 

application] 

Emilia javanica (Burm.) C.B. Robinson [not in Guianas] 

Emilia sagittata (Vahl) DC. [not in Guianas] 

Erechtites "surinamensis L." [invalid name, error for 

Clibadium surinamense L.] 

Erigeron bonariensis L. =Conyza bonariensis 

Erigeron canadensis L. =Conyza canadensis 

Eupatorium amygdalinum Lam. =Ayapana amygdalina 

Eupatorium corymbosum Aubl. =Chromolaena corymbosa 

Eupatorium denticulatum Vahl =Mikania cordifolia or 

Mikania micrantha 

Eupatorium laevigatum Lam. =Chromolaena laevigata 

Eupatorium macrophyllum L. =Hebeclinium macrophyllum 

Eupatorium martiusii DC. =Chromolaena squalida 

Eupatorium microstemon Cass. =Fleischmannia microstemon 

Eupatorium odoratum L. =Chromolaena odorata 

Eupatorium parviflorum Aubl. =Mikania parviflora 

Eupatorium pauciflorum Kunth =Praxelis pauciflora 

Eupatorium roupalifolium R. Robinson =Guayania 

roupalifolia 

Eupatorium triflorum Aubl. =Piptocarpha triflora 

Eupatorium triplinerve Vahl =Ayapana triplinervis 

Eupatorium vitalbae DC. =Heterocondylus vitalbae 

Gamochaeta americana (Mill.) Wedd. [not in Guianas] 

Gamochaeta purpurea (L.) Cabrera =Gamochaeta 

simplicicaulis 

Geissopappus caleoides (DC.) Benth. ex Baker =Calea 

caleoides 

Gnaphalium purpureum L. =Gamochaeta simplicicaulis 

Gnaphalium schomburgkii Schultz. Bip. [nomen nudum] 

=Gnaphalium polycaulon 

Gnaphalium spicatum Lam. =Gamochaeta americana (Mill.) 

Wedd. [not in Guianas] 

Gongylolepis erioclada S.F. Blake [not in Guianas] 

Gongylolepis glaberrima S.F. Blake [not in Guianas] 

Heterothalamus densus N.E. Brown =Baccharis densa 

Melampodium camphoratum (L.f.) Baker =Unxia camphorata 

Melanthera aspera (Jacq.) Small =Melanthera nivea 

Mikania amara (Vahl) Willd. =Mikania parviflora 

Mikania aschersonii sensu Pruski, non Hieron. =Mikania 

chlorolepis 

Mikania gracilipes B.L. Robinson [not in Guianas] 

Mikania hirsutissima DC. =Mikania banisteriae 

Mikania lanata Schultz Bip. [not in Guianas] 

Mikania mazaruniensis Holmes & McDaniel =Mikania 

banisteriae 

Mikania racemulosa Benth. =Mikania psilostachya 

Mikania scabra DC. =Mikania psilostachya 

Mikania tepuiensis Badillo =Mikania nigropunctulata 

Mikania tillettii R. King & H. Robinson =Mikania gleasonii 

Oliganthes schomburgkii Schultz Bip. =Piptocoma 

schomburgkii 

Oyedaea rusbyi S.F. Blake [not in Guianas] 

Pectis ciliaris L. [not in Guianas] 

Pluchea purpurascens (Swartz) DC. =Pluchea odorata 

Pollalesta milleri (Johnston) Aristeguieta [not in Guianas] 

Pollalesta roraimensis (Steyerm.) Aristegueta =Piptocoma 

roraimensis 

Pollalesta schomburgkii (Schultz Bip.) Aristeguieta 

=Piptocoma schomburgkii 

Porophyllum ellipticum (L.) Cass. =Porophyllum ruderale 

Pseudelephantopus spiralis (Less.) Cronq. [not in Guianas] 

Pterocaulon virgatum (L.) DC. [not in Guianas] 

Riencourtia pittieri S.F. Blake [not in Guianas] 

Rolandra argentea Rottb. =Rolandra fruticosa 

Sonchus oleraceus L. [not in Guianas] 

Sparganophorus sparganophora (L.) C. Jeffrey =Struchium 

sparganophorum 

Spilanthes acmella (L.) Murr. =Acmella sp. 

Spilanthes oppositifolia (Lam.) D'Arcy =Acmella repens 

Spilanthes paniculata Wall =Acmella sp.[not in Guianas] 

Spilanthes uliginosa Swartz =Acmella uliginosa 

Spiracantha cornifolia Kunth [not in Guianas] 

Stenopadus cardonae Maguire & Lasser =Stenopadus 

talaumifolius 

Stenopadus condensatus (Baker) S.F. Blake =Stomatochaeta 

condensata 

Stenopadus crassifolius S.F. Blake [not in Guianas] 

Stenopadus eurylepis S.F. Blake [not in Guianas] 

Stenopadus variabilis S.F. Blake =Stomatochaeta condensata 

Stifftia benthamiana (Rob. Schomb.) Baker =Gongylolepis 

benthamiana 

Stifftia condensata Baker =Stomatochaeta condensata 

Stifftia connellii N.E. Brown =Stenopadus connellii 

Stilpnopappus aquaticus (Poepp.) Dillon =Xiphochaeta 

aquatica 

Stilpnopappus viridis Benth. ex Baker =Xiphochaeta aquatica 

Stomatochaeta schomburgkii (Schultz Bip.) Maguire & 

Wurdack [nomen nudum] 

Trichospira menthoides Kunth =Trichospira verticillata 

Trixis pedunculosa Rich. =Riencourtia pedunculosa 

Verbesina alata L. [not in Guianas] 

Verbesina schomburgkii Schultz Bip. ex Klatt var. ligulata 

Maguire & Wurdack =Verbesina ligulata 

Vernonia bolivarensis Badillo =Lepidaploa bolivarensis 

Vernonia brasiliana (L.) Druce =Vernonanthura brasiliana 

Vernonia cardonae Aristeguieta =Lepidaploa cardonae 

Vernonia chrysotricha Alexander =Lepidaploa chrysotricha 

Vernonia cinerea (L.) Lessing =Cyanthillium cinereum 

Vernonia cotoneaster Less. =Lepidaploa cotoneaster 

Vernonia glandulosa Badillo =Lepidaploa bolivarensis 

Vernonia gracilis Kunth =Lepidaploa gracilis 

Vernonia moritziana Schultz Bip. =Lepidaploa gracilis 

Vernonia remotiflora L.C. Rich. =Lepidaploa remotiflora 

Vernonia schomburgkiana Schultz Bip. =Lepidaploa 

ehretiifolia 

Vernonia scorpioides (Lam.) Pers. =Cyrtocymura scorpioides 

Vernonia sericea L.C. Rich. =Lepidaploa sericea (L.C. Rich.) 

H. Robinson[not in Guianas] 

Vernonia sessiliflora Willd. =Lepidaploa gracilis 

Vernonia tricholepis DC. =Lepidaploa remotiflora 

Wedelia brachycarpa Baker =Sphagneticola brachycarpa 

Wedelia trilobata (L.) Hitchc. =Sphagneticola trilobata 

Zexmenia tepuiana Badillo =Oyedaea tepuiana 


61

62 


BALANOPHORACEAE 

reviewed by B. Hansen (1992); reviewed by B. 

Hansen (1996) 

REFERENCES: 

Hansen, B. 1980. Balanophoraceae. Flora 


Hansen, B. 1993. Balanophoraceae. In A.R.A. 


A) 14: 40-43. 

Lanjouw, J. 1932. Balanophoraceae. In A. 


Helosis cayennensis (Swartz) Spreng. var. cayennensis [GU 

SU FG] 

Langsdorffia hypogaea Mart. [GU] 

BALANOPHORACEAE - SYNONYMS AND 


Cynomorium cayennense Swartz =Helosis cayennensis 

Helosis guyanensis L.C. Rich., nom. illeg. superfl. =Helosis 

cayennensis 

BALSAMINACEAE 

Impatiens balsamina L. [GU] [probably escaped from 

cultivation] 

BASELLACEAE 


REFERENCES: 

Sperling, C.R. 1987. Systematics of the 

Basellaceae. Ph.D. Dissertation, Harvard 

University. U.M Dissertation Information 

Service, Ann Arbor, Michigan. 

Anredera vesicaria (Lam.) Gaertn. [FG] [probably escaped 

from cultivation] 

Basella alba L. [GU SU FG] [cultivated & naturalized] 

BASELLACEAE - SYNONYMS AND 


Anredera leptostachys (Moquin) van Steenis =Anredera 

vesicaria 

Basella rubra L. =Basella alba 

Boussingaltia leptostachya Moq. =Anredera vesicaria 

SPERMATOPHYTA: Balanophoraceae - Begoniaceae 

REFERENCES: 

BATACEAE 

Jonker, F.P. 1953. Batidaceae. In A. Pulle & J. 

Lanjouw, Flora of Suriname 1(2): 91-93. 

Batis maritima L. [GU SU FG?] 

BEGONIACEAE 


D. Wasshausen, J.C. Lindeman (1996) 

REFERENCES: 

Lindeman, J.C. 1984. Begoniaceae. In A.L. 

Stoffers & J.C. Lindeman, Flora of Suriname 

5(1): 576-584. 

Smith, L.B., D.C. Wasshausen, J. Golding, & 

C.E. Karegeannes. 1986. Begoniaceae 

(illustrated key and annotated species list). 

Smithsonian Contrib. Bot. 60: 1-584. 

Begonia fischeri Schrank [GU] 

Begonia glabra Aubl. [SU FG] 

Begonia heloisana Brade [GU] 

Begonia hirsuta Aubl. [FG] 

Begonia humilis Dryander [GU SU FG] 

Begonia jenmanii Tutin [GU] 

Begonia prieurii A. DC. [GU SU FG] 

Begonia semiovata Liebm. [GU SU FG] 

Begonia steyermarkii L.B. Smith & B.G. Schub. [GU] 

BEGONIACEAE - SYNONYMS AND 


Begonia diffusa L.B. Smith & B.G. Schub. [not in Guianas] 

Begonia filipes Benth. [not in Guianas] 

Begonia guaduensis Kunth [not in Guianas] 

Begonia guyanensis A. DC. =Begonia semiovata 

Begonia hirtella Link [not in Guianas] 

Begonia kaietukensis Tutin =Begonia fischeri 

Begonia kvistii L.B. Smith [nomen nudum] =Begonia 

jenmanii 

Begonia roraimensis Tutin =Begonia fischeri 

Begonia scandens Swartz =Begonia glabra 

Begonia tovarensis Klotzsch =Begonia fischeri


BIGNONIACEAE 

contributed by A.H. Gentry (1992) 

REFERENCES: 

Gentry, A.H. 1978. Bignoniaceae. In B. 

Maguire & collaborators, 1978. Botany of the 


Bot. Gard. 29: 245-283. 

Gentry, A.H. 1980. Bignoniaceae - Part I 

(Crescentieae and Tourretieae). Flora 

Neotropica Monogr. 25(I): 1-130. 

Gentry, A.H. 1982. Bignoniaceae. In Z. Luces 

de Febres, Flora de Venezuela 8: 1-462. 

Gentry, A.H. 1992. Bignoniaceae - Part II (Tribe 

Tecomeae). Flora Neotropica Monogr. 25(II): 

1-335. 

Sandwith, N.Y. 1938. Bignoniaceae. In A. 


Adenocalymna impressum (Rusby) Sandw. [SU FG] 

Adenocalymna inundatum Mart. ex DC. var. surinamense 

Bureau & K. Schum. [GU SU FG] 

Adenocalymna prancei A. Gentry [FG] 

Adenocalymna purpurascens Rusby [GU] 

Adenocalymna saulense A. Gentry [FG] 

Adenocalymna subincanum Huber [FG] 

Amphilophium paniculatum (L.) Kunth var. imatacense A. 

Gentry [GU SU FG] 

Anemopaegma chrysoleucum (Kunth) Sandw. [GU SU FG] 

Anemopaegma foetidum Bureau & K. Schum. [GU SU] 

Anemopaegma granvillei A. Gentry [FG] 

Anemopaegma ionanthum A. Gentry [GU FG] 

Anemopaegma jucundum Bureau & K. Schum. [GU] 

Anemopaegma karstenii Bureau & K. Schum. [GU] 

Anemopaegma longidens DC. [FG] 

Anemopaegma maguirei Sandw. [GU? SU] 

Anemopaegma oligoneuron (Sprague & Sandw.) A. Gentry 

[GU SU] 

Anemopaegma paraense Bureau & K. Schum. [GU? SU FG] 

Anemopaegma parkeri Sprague [GU SU FG] 

Anemopaegma robustum Bureau & K. Schum. [GU FG] 

Arrabidaea arthrerion (DC.) Bureau ex K. Schum. [SU?] 

Arrabidaea bilabiata (Sprague) Sandw. [GU SU FG] 

Arrabidaea brachypoda (DC.) Bureau [GU?] 

Arrabidaea candicans (L.C. Rich.) DC. [GU SU FG] 

Arrabidaea chica (Humb. & Bonpl.) Verl. [SU? FG] 

Arrabidaea cinerea Bureau ex K. Schum. var. cinerea [GU 

FG] 

Arrabidaea cinerea Bureau ex K. Schum. var. longipila Sandw. 

[GU] 

Arrabidaea cinnamomea (DC.) Sandw. [FG] 

Arrabidaea corallina (Jacq.) Sandw. [GU SU? FG] 

Arrabidaea egensis Bureau & K. Schum. [GU] 

Arrabidaea fanshawei Sandw. [GU SU FG] 

Arrabidaea florida DC. [GU SU FG] 

Arrabidaea grosourdyana (Baill.) Sandw. [GU FG] 

Arrabidaea inaequalis (DC. ex Splitg.) K. Schum. [GU SU 

FG] 

Arrabidaea japurensis (DC.) Bureau & K. Schum. [GU FG] 

Arrabidaea mollis (Vahl.) Bureau ex K. Schum. [GU SU FG] 

Arrabidaea nigrescens Sandw. [GU SU FG] 

Arrabidaea oligantha Bureau & K. Schum. [GU SU FG] 

Arrabidaea patellifera (Schlecht.) Sandw. [GU SU FG] 

Arrabidaea prancei A. Gentry [GU] 

Arrabidaea pubescens (L.) A. Gentry [GU] 

Arrabidaea revillae A. Gentry [GU] 

Arrabidaea spicata Bureau & K. Schum. [FG] 

Arrabidaea trailii Sprague [SU FG] 

Arrabidaea tuberculata DC. [GU SU? FG] 

Callichlamys latifolia (L.C. Rich.) K. Schum. [GU SU FG] 

Ceratophytum tetragonolobum (Jacq.) Sprague & Sandw. [GU 

SU] 

Clytostoma binatum (Thunb.) Sandw. [GU SU FG] 

Crescentia cujete L. [GU SU FG] [cultivated only?] 

Cuspidaria subincana A. Gentry [GU SU] 

Cybistax antisyphilitica (Mart.) Mart. ex DC. [SU] 

Cydista aequinoctialis (L.) Miers [GU SU FG] 

Cydista lilacina A. Gentry [SU FG] 

Digomphia densicoma (Mart. ex DC.) Pilger [GU] 

Digomphia laurifolia Benth. [GU] 

Distictella cremersii A. Gentry [FG] 

Distictella cuneifolia (DC.) Sandw. [FG] 

Distictella elongata (Vahl) Urb. [GU SU FG] 

Distictella magnoliifolia (Kunth) Sandw. [GU SU FG] 

Distictella obovata Sandw. [GU] 

Distictella parkeri (DC.) Sprague & Sandw. [GU] 

Distictella porphyrotricha Sandw. [GU] 

Distictella sp. cf. obovata Sandw. [FG] 

Distictis granulosa Bureau & K. Schum. [GU SU? FG] 

Distictis pulverulenta (Sandw.) A. Gentry [GU FG] 

Godmania aesculifolia (Kunth) Standl. [GU] 

Jacaranda copaia (Aubl.) D. Don subsp. copaia [GU SU FG] 

Jacaranda copaia (Aubl.) D. Don subsp. spectabilis (Mart. ex 

DC.) A. Gentry [GU SU FG] 

Jacaranda duckei Vattimo [SU FG] 

Jacaranda obtusifolia Humb. & Bonpl. subsp. rhombifolia (G. 

Mey.) A. Gentry [GU SU FG] 

Lundia corymbifera (Vahl) Sandw. [FG] 

Lundia densiflora DC. [GU SU] 

Lundia erionema DC. [GU SU FG] 

Macfadyena uncata (Andr.) Sprague & Sandw. [GU SU FG] 

Macfadyena unguis-cati (L.) A. Gentry [GU SU FG] 

Mansoa alliacea (Lam.) A. Gentry [GU SU FG] 

Mansoa kerere (Aubl.) A. Gentry [GU SU FG] 

Mansoa sp. cf. onohualcoides A. Gentry [SU] 

Mansoa sp. nov. aff. standleyi [FG] 

Mansoa standleyi (Steyerm.) A. Gentry [GU SU FG] 

Mansoa verrucifera (Schlecht.) A. Gentry [GU] 

Martinella obovata (Kunth) Bureau & K. Schum. [GU SU FG] 

Melloa quadrivalvis (Jacq.) A. Gentry [GU] 

Memora alba (Aubl.) Miers [FG] 

Memora bracteosa (DC.) Bureau ex K. Schum. [GU SU? FG] 

Memora flavida (DC.) Bureau & K. Schum. [GU SU FG] 

Memora flaviflora (Miq.) Pulle [GU SU FG] 

Memora heterophylla (Kraenzl.) Sandw. [GU] 

Memora moringifolia (DC.) Sandw. [SU FG] 

Memora racemosa A. Gentry [SU FG] 

Memora schomburgkii (DC.) Miers [GU SU FG] 

Memora tanaeciicarpa A. Gentry [GU FG] 

Mussatia hyacinthina (Standley) Sandw. [GU] 

Mussatia prieurei (DC.) Bureau ex K. Schum. [GU SU FG] 

Parabignonia steyermarkii Sandw. [GU SU FG] 

Paragonia pyramidata (L.C. Rich.) Bureau [GU SU FG] 

Phryganocydia corymbosa (Venten.) Bureau ex K. Schum. 

[GU FG] 

Pithecoctenium crucigerum (L.) A. Gentry [GU SU FG] 

SPERMATOPHYTA: Bignoniaceae 

63

64 


Pleonotoma albiflora (Salzm. ex DC.) A. Gentry [GU SU FG] 

Pleonotoma clematis (Kunth) Miers [GU SU FG] 

Pleonotoma echitidea Sprague & Sandw. [GU SU] 

Pleonotoma moringaefolia (DC.) Miers [FG] [unverified 

name] 

Pleonotoma sp. cf. dendrotricha Sandw. [FG] 

Potamoganos microcalyx (G. Mey.) Sandw. [GU SU] 

Pseudocalymma sagotii (Bureau & K. Schum.) Sandw. [FG] 

[unverified name] 

Pyrostegia dichotoma Miers ex K. Schum. [GU SU] 

Roentgenia sordida (Bureau & K. Schum.) Sprague & Sandw. 

[GU SU FG] 

Schlegelia fuscata A. Gentry [FG] 

Schlegelia macrophylla Ducke [FG] 

Schlegelia paraensis Ducke [SU FG] 

Schlegelia roseiflora Ducke [FG] 

Schlegelia spruceana Bureau & K. Schum. [GU] 

Schlegelia violacea (Aubl.) Griseb. [GU SU FG] 

Stizophyllum inaequilaterum Bureau & K. Schum. [GU SU 

FG] 

Stizophyllum riparium (Kunth) Sandw. [SU FG] 

Tabebuia aurea (S. Manso) Benth. & J.D. Hook. ex S. Moore 

[SU] 

Tabebuia capitata (Bureau & K. Schum.) Sandw. [GU SU FG] 

Tabebuia fluviatilis (Aubl.) DC. [GU SU FG] 

Tabebuia impetiginosa (Mart. ex DC.) Standl. [SU FG] 

Tabebuia insignis (Miq.) Sandw. var. insignis [GU SU FG] 

Tabebuia insignis (Miq.) Sandw. var. monophylla Sandw. [GU 

SU] 

Tabebuia obscura (Bureau & K. Schum.) Sandw. [FG?] 

Tabebuia ochracea (Cham.) Standl. subsp. heterotricha (DC.) 

A. Gentry [GU] 

Tabebuia serratifolia (Vahl) Nichols. [GU SU FG] 

Tabebuia stenocalyx Sprague & Stapf [GU FG] 

Tabebuia subtilis Sprague & Sandw. [GU] 

Tabebuia uleana (Kraenzl.) A. Gentry [GU] 

Tanaecium jaroba Swartz [GU] 

Tanaecium nocturnum (Barb. Rodr.) Bureau & K. Schum. [SU 

FG] 

Tynanthus pubescens A. Gentry [GU FG] 

Tynanthus sastrei A. Gentry [FG] 

Xylophragma seemannianum (Kuntze) Sandw. [GU SU] 

BIGNONIACEAE - SYNONYMS AND 


Adenocalymna bilabiatum (Sprague) Sandw. =Arrabidaea 

bilabiata 

Anemopaegma carrerense Armitage =Anemopaegma karstenii 

Anomoctenium granulosum (Bureau & K. Schum.) Sandw. 

=Distictis granulosa 

Arrabidaea obliqua (Kunth) Bureau =Arrabidaea corallina 

Bignonia alba Aubl. =Memora alba 

Bignonia aquatilis E. Mey. =Tabebuia fluviatilis 

Bignonia copaia Aubl. =Jacaranda copaia 

Bignonia digitata E. Mey. =Tabebuia fluviatilis 

Bignonia echinata Aubl. =Pithecoctenium crucigerum 

Bignonia filicifolia Anderson =Jacaranda obtusifolia subsp. 

rhombifolia 

Bignonia fluviatilis Aubl. =Tabebuia fluviatilis 

Bignonia incarnata Aubl. =Mansoa kerere 

Bignonia kerere Aubl. =Mansoa kerere 

Bignonia tinctoria Arruda =Arrabidaea chica? 

Bignonia unguis-cati L. =Macfadyena unguis-cati 

SPERMATOPHYTA: Bignoniaceae 

Ceratophytum tobagense (Urb.) Sprague & Sandw. 

=Ceratophytum tetragonolobum 

Distictella pulverulenta Sandw. =Distictis pulverulenta 

Distictella racemosa (Bureau & K. Schum.) Urb. =Distictella 

magnoliifolia 

Handroanthus durus (Bureau ex K. Schum.) Mattos =Tabebuia 

insignis var. insignis 

Jacaranda filicifolia D. Don =Jacaranda obtusifolia subsp. 

rhombifolia 

Jacaranda filicifolia D. Don var. puberula K. Schum. 

=Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda obtusifolia Humb. & Bonpl. var. rhombifolia (G. 

Mey.) Sandw. =Jacaranda obtusifolia subsp. rhombifolia 

Jacaranda paraensis (Huber) Vattimo =Jacaranda copaia subsp. 

spectabilis 

Jacaranda rhombifolia G. Mey. =Jacaranda obtusifolia subsp. 

rhombifolia 

Jacaranda spectabilis Mart. ex A.P. DC. =Jacaranda copaia 

subsp. spectabilis 

Martinella insculpta Sprague & Sandw. =Martinella obovata 

Memora ovata (Bureau & K. Schum.) Sprague & Sandw. 

=Memora schomburgkii 

Pachyptera alliacea (Lam.) A. Gentry =Mansoa alliacea 

Pachyptera kerere (Aubl.) Sandw. =Mansoa kerere 

Petastoma patelliferum (Schlechtend.) Miers =Arrabidaea 

patellifera 

Pithecoctenium echinatum (Jacq.) Baill. =Pithecoctenium 

crucigerum 

Pithecoctenium granulosum (Bureau & K. Schum.) Sprague & 

Sandw. =Distictis granulosa 

Pseudocalymma alliaceum (Lam.) Sandw. =Mansoa alliacea 

Pseudopaegma oligoneuron Sprague & Sandw. 

=Anemopaegma oligoneuron 

Roentgenia tuberculata Sprague & Sandw. [name of uncertain 

application, apparently a nomen nudum] 

Schlegelia lilacina Miq. =Schlegelia violacea 

Schlegelia parasitica (Swartz) Miers ex Griseb. [not in 


Tabebuia aquatilis (E. Mey.) Sprague & Sandw. =Tabebuia 

fluviatilis 

Tabebuia caraiba (Mart.) Bureau =Tabebuia aurea 

Tabebuia chrysotricha (Mart. ex DC.) Standl. [not in Guianas] 

Tabebuia dura (Bureau ex K. Schum.) Sprague & Sandw. 

=Tabebuia insignis var. insignis 

Tabebuia floccosa (Bureau & K. Schum.) Sprague & Sandw. 

=Xylophragma seemannianum 

Tabebuia hypolepra Sprague & Sandw. =Tabebuia capitata 

Tabebuia ipe (Mart.) Standl. var. integra (Sprague) Sandw. 

=Tabebuia impetiginosa 

Tabebuia latifolia A.P. DC. =Martinella obovata 

Tabebuia longipes Baker =Tabebuia insignis var. monophylla 

Tabebuia neurophylla Miq. =Arrabidaea patellifera 

Tabebuia roraimae Oliver =Tabebuia insignis var. insignis 

Tecoma conspicua A.P. DC. =Tabebuia serratifolia 

Tecoma dura Bureau ex K. Schum. =Tabebuia insignis var. 

insignis 

Tecoma fluviatilis (Aubl.) Miq. =Tabebuia fluviatilis 

Tecoma insignis Miq. =Tabebuia insignis 

Tecoma meyeriana A.P. DC. =Tabebuia fluviatilis


Tecoma patrisiana A.P. DC. =Tabebuia serratifolia 

Tecoma roraimae (Oliver) K. Schum. =Tabebuia insignis var. 

insignis 

Tecoma uleana Kraenzl. =Tabebuia uleana 

Tetrastictella inaequalis (DC. ex Splitg.) M. Pichon 

=Arrabidea inaequalis 

Zeyheria surinamensis Miq. =Paragonia pyramidata 

REFERENCES: 

BIXACEAE 

Poppendieck, H.H. 1981. Cochlospermaceae. 

Flora Neotropica Monogr. 27: 1-33. 

Bixa orellana L. [GU SU FG] 

Bixa urucurana Willd. [GU SU] [=Bixa orellana?] 

Cochlospermum orinocense (Kunth) Steudel [GU SU FG] 

Cochlospermum vitifolium (Willd.) Spreng. [GU SU FG?] 

BIXACEAE - SYNONYMS AND 


Cochlospermum insigne St. Hil. =Cochlospermum regium 

(Schr.) Pilger [not in Guianas] 

Cochlospermum parkeri Planchon =Cochlospermum 

orinocense 

Cochlospermum parviifolium Planch. =Cochlospermum 

orinocense 

Cochlospermum wentii Pulle =Cochlospermum orinocense 

BOMBACACEAE 

reviewed by L.J. Dorr (1992) 

REFERENCES: 

Alverson, W.S. 1994. New species and 

combinations of Catostemma and Pachira 

(Bombacaceae) from the Venezuelan 

Guayana. Novon 4(1): 3-8. 

Robyns, A. 1963. Essai de monographie du 

genre Bombax s.l. Bull. Jard. Bot. État 33: 

1-311. 

Robyns, A. 1967. Bombacaceae. In B. Maguire 


Highland part 7. Mem. New York Bot. Gard. 

17(1): 1-439. 

Steyermark, J.A. 1987. Bombacaceae. In Flora 

of the Venezuelan Guayana - III. Ann. 

Missouri Bot. Gard. 74(3): 636-646. 

Steyermark, J.A. & W.D. Stevens. 1988. Notes 

on Rhodognaphalopsis and Bombacopsis in 

the Guayanas. Ann. Missouri Bot. Gard. 75: 

396-398. 

Uittien, H. 1932. Bombacaceae. In A. Pulle, 


Bombacopsis macrocalyx (Ducke) A. Robyns [FG?] 

Bombacopsis nervosa (Uittien) A. Robyns [GU? SU FG] 

Bombacopsis quinata (Jacq.) Dugand [GU] 

Catostemma altsonii Sandw. [GU] 

Catostemma commune Sandw. [GU] 

Catostemma fragrans Benth. [GU SU FG] 

Ceiba pentandra (L.) Gaertn. [GU SU FG] 

Eriotheca crassa (Uittien) A. Robyns [SU FG] 

Eriotheca globosa (Aubl.) A. Robyns [GU SU FG] 

Eriotheca surinamensis (Uittien) A. Robyns [GU SU FG] 

Huberodendron swietenioides (Gleason) Ducke [FG] 

Matisia ochrocalyx K. Schum. [FG] 

Ochroma pyramidale (Cav. ex Lam.) Urb. [FG] 

Pachira aquatica Aubl. [GU SU FG] 

Pachira insignis (Swartz) Sav. [GU SU FG] 

Quararibea amazonica Ulbrich [FG] 

Quararibea duckei Huber [SU FG] 

Quararibea guianensis Aubl. [GU SU FG] 

Quararibea lasiocalyx (K. Schum.) Vischer [FG] 

Rhodognaphalopsis flaviflora (Pulle) A. Robyns var. flaviflora 

[GU SU FG] 

Rhodognaphalopsis flaviflora (Pulle) A. Robyns var. glabra A. 

Robyns [SU] 

Rhodognaphalopsis minor (Sims) A. Robyns var. minor [GU] 

Scleronema guianense Sandw. [GU] 

BOMBACACEAE - SYNONYMS AND 


Bombax aquaticum (Aubl.) K. Schum. =Pachira aquatica 

Bombax crassum Uittien =Eriotheca crassa 

Bombax flaviflorum Pulle =Rhodognaphalopsis flaviflora 

Bombax globosum Aubl. =Eriotheca globosa 

Bombax jenmanii Oliver =Rhodognaphalopsis minor 

Bombax nervosum Uittien =Bombacopsis nervosa 

Bombax spectabile Ulbr. =Pachira insignis 

Bombax surinamense Uittien =Eriotheca surinamensis 

Catostemma macrosperma (Sagot) Lemée =Catostemma 

fragrans 

Ceiba occidentalis (Spreng.) Burkhill =Ceiba pentandra 

Pochota flaviflora (Pulle) Steyerm. & W.D. Stevens 

=Rhodognaphalopsis flaviflora 

Pochota minor (Sims) Steyerm. & W.D. Stevens 

=Rhodognaphalopsis minor 

Quararibea turbinata Poir. [specimens apparently referable to 

Quararibea duckei] 

BONNETIACEAE 

reviewed by A. Weitzman (1992) 

REFERENCES: 

Kobuski, C.E. 1948. Studies in the Theaceae, 

XVII. A review of the genus Bonnetia. J. 

Arn. Arb. 29: 393-413. 

Maguire, B. 1972. Bonnetiaceae. In B. Maguire 



23: 1-832. 

SPERMATOPHYTA: Bignoniaceae - Bonnetiaceae 

65

66 


Archytaea triflora Mart. [GU] 

Bonnetia multinervia (Maguire) Steyerm. [to be expected in 

GU?] 

Bonnetia paniculata Spruce ex Benth. [GU] 

Bonnetia roraimae Oliver [GU] 

Bonnetia rubicunda (Sastre) A. Weitzman & Stevens, comb. 

nov. ined. [GU] 

Bonnetia sessilis Benth. [GU] 

Bonnetia tepuiensis Kobuski & Steyerm. [GU] 

BONNETIACEAE - SYNONYMS AND 


Archytaea multiflora Benth. =Archytaea triflora, syn. nov. 

ined. 

Neblinaria rubicunda Sastre =Bonnetia rubicunda 

Neogleasonia multinervia Maguire =Bonnetia multinervia 

BORAGINACEAE 

reviewed by C. Feuillet (1992); reviewed by C. 

Feuillet (1996) 

REFERENCES: 

Johnston, I.M. 1928. Studies in the 

Boraginaceae VII. 1. The South American 

species of Heliotropium. Contrib. Gray Herb. 

81: 3-72. 

Johnston, I.M. 1935. Studies in Boraginaceae X. 

The Boraginaceae of northeastern South 

America. J. Arn. Arb. 16: 1-64. 

Johnston, I.M. 1936. Boraginaceae. In A. Pulle, 


Uittien, H. 1937. Boraginaceae (additions and 


4(1): 496-497. 

Cordia alliodora (Ruiz & Pav.) Cham. ex DC. [GU SU FG] 

Cordia bicolor A. DC. [GU SU FG] 

Cordia candidula (Miers) I.M. Johnston [GU] 

Cordia curassavica (Jacq.) Roem. & Schult. [GU SU FG] 

Cordia exaltata Lam. var. exaltata [FG] 

Cordia exaltata Lam. var. melanoneura (Klotzsch) I.M. 

Johnston [GU] 

Cordia fallax I.M. Johnston [GU] 

Cordia fulva I.M. Johnston [SU FG] 

Cordia goeldiana Huber [FG] 

Cordia grandiflora (Desv.) Kunth [GU] 

Cordia hirta I.M. Johnston [GU SU FG] 

Cordia laevifrons I.M. Johnston [SU FG] 

Cordia lomatoloba I.M. Johnston [GU SU FG] 

Cordia multispicata Cham. [GU] 

Cordia nervosa Lam. [GU SU FG] 

Cordia nodosa Lam. [GU SU FG] 

Cordia panicularis Rudge [GU SU FG] 

Cordia polycephala (Lam.) I.M. Johnston [GU SU] 

Cordia sagotii I.M. Johnston [GU SU FG] 

Cordia scabrifolia A. DC. [GU] 

Cordia schomburgkii A. DC. [GU SU FG] 

Cordia sericicalyx A. DC. [GU SU] 

Cordia sprucei Mez [FG] 

SPERMATOPHYTA: Bonnetiaceae - Brassicaceae 

Cordia tetrandra Aubl. [GU SU FG] 

Cordia tomentosa Lam. ex Roem. & Schult. [SU FG] 

Cordia toqueve Aubl. [GU FG] 

Cordia wurdackii C. Feuillet, sp. nov. ined. [GU] 

Heliotropium curassavicum L. [GU SU FG?] 

Heliotropium filiforme Lehmann [GU SU FG] 

Heliotropium indicum L. [GU SU FG] [cultivated, persists as 

an escape] 

Heliotropium lagoense (Warm.) Guerke [SU] 

Heliotropium procumbens Mill. [GU] 

Heliotropium ternatum Vahl [GU] 

Lepidocordia punctata Ducke [GU] 

Tournefortia bicolor Swartz var. bicolor [GU SU FG] 

Tournefortia bicolor Swartz var. calycosa J.D. Smith [GU SU] 

Tournefortia cuspidata Kunth [GU SU] 

Tournefortia maculata Lam. [GU SU FG] 

Tournefortia melanochaeta DC. [FG] 

Tournefortia paniculata Cham. var. spigeliiflora (A. DC.) I.M. 

Johnston [GU SU FG] 

Tournefortia ulei Vaupel [GU SU FG] 

Tournefortia volubilis L. [GU FG] 

BORAGINACEAE - SYNONYMS AND 


Cordia collococa Aubl. non L. =Cordia nodosa 

Cordia flavescens Aubl. [name of uncertain application, 

referable to Lauraceae] 

Cordia heterophylla Willd. =Cordia toqueve 

Cordia lockhartii nom. inval. =Cordia bicolor A. DC. 

Cordia macrostachya (Jacq.) Roem. & Schult. =Cordia 

curassavica 

Cordia scandens Poir. =Dichapetalum rugosum 

(Dichapetalaceae) 

Cordia tetraphylla Aubl. =Buchenavia tetraphylla 

(Combretaceae) 

Tournefortia aubletii Macbr. =Tournefortia bicolor 

Tournefortia syringaefolia Vahl =Tournefortia maculata 

BRASSICACEAE 



REFERENCES: 

Al-Shehbaz, I. 1988. Cruciferae. In R.A. 

Howard, Flora of the Lesser Antilles 4: 276- 

292. 

Brassica juncea (L.) Czern. & Coss. [FG?] [introduced] 

Coronopus didymus (L.) J.E. Sm. [FG] [probably introduced] 

Lepidium virginicum L. [SU FG] [introduced] 

Rorippa dubia (Pers.) H. Hara [GU SU] [introduced] 

[=Rorippa indica (L.) Hiern] 

Rorippa nasturtium-aquaticum (L.) Hayek [FG] [introduced] 

[=Nasturtium officinale R. Br.] 

Rorippa palustris (L.) Besser [SU] [introduced] [=Rorippa 

islandica (Leder) Borbás]

Sinapis arvensis L. [GU] [introduced] 


BRASSICACEAE - SYNONYMS AND 


Lepidium micropterum Miq. =Lepidium virginicum 

Nasturtium palustre (L.) DC., non Crantz =Rorippa palustris 

Rorippa heterophylla (Blume) R.O. Williams =Rorippa dubia 

BROMELIACEAE 

reviewed by E.J. Gouda, J. Grant, & M. Spencer 

(1992); reviewed by B.K. Holst, J. Grant, E.J. 

Gouda, & H. Luther (1996) 

REFERENCES: 

Grant, J.R. 1995. The resurrection of Alcantarea 

and Werauhia, a new genus. Tropische und 

subtropische Pflanzenwelt 91: 1-59. 

Gouda, E.J. 1987. Bromeliaceae, subfamily 

Tillandsioideae. In A.R.A. Görts-van Rijn, 

Flora of the Guianas (series A) 3: 1-112. 

Holst, B.K. 1994. Checklist of Venezuelan 

Bromeliaceae with notes on species 

distribution by state and levels of endemism. 

Selbyana 15(1): 132-149. 

Kramer, K.U. & A.R.A. Görts-van Rijn. 1968. 

Bromeliaceae (additions and corrections). In 

A. Pulle & J. Lanjouw, Flora of Suriname 

1(2): 413-414. 

Smith, L.B. 1957. Bromeliaceae. In A. Pulle & 

J. Lanjouw, Flora of Suriname 1(2): 94-148. 

Smith, L.B. 1967. Bromeliaceae of the Guayana 

Highland. Mem. New York Bot. Gard. 14(3): 

15-68. 

Smith, L.B. 1986. Revision of the Guayana 

Highland Bromeliaceae. Ann. Missouri Bot. 

Gard. 73(4): 689-721. 

Smith, L.B. & R.J. Downs. 1974. 

Pitcairnioideae. Flora Neotropica Monogr. 

14, part 1: 1-658. 


Tillandsioideae. Flora Neotropica Monogr. 

14, part 2: 663-1492. 


Bromelioideae. Flora Neotropica Monogr. 

14, part 3: 1493-2142. 

Aechmea angustifolia Poepp. & Endl. [GU] 

Aechmea aquilega (Salisb.) Griseb. [GU SU FG] 

Aechmea brassicoides Baker [GU] 

Aechmea bromeliifolia (Rudge) Baker [GU SU FG] 

Aechmea campanulata L.B. Smith [GU] 

Aechmea castelnavii Baker [to be expected in FG] 

Aechmea contracta (Mart. ex Schult.f.) Baker [GU] 

Aechmea egleriana L.B. Smith var. egleriana [FG] 

Aechmea egleriana L.B. Smith var. major L.B. Smith [to be 

expected in FG] 

Aechmea fernandae (E. Morren) Baker [to be expected in FG] 

Aechmea lanjouwii (L.B. Smith) L.B. Smith [SU] 

Aechmea lingulata (L.) Baker [GU SU FG] 

Aechmea megalantha Harms [FG] [probably misident. of A. 

melinonii according to Gouda] 

Aechmea melinonii Hook. [GU SU FG] 

Aechmea mertensii (Mey.) Schult.f. [GU SU FG] 

Aechmea nudicaulis (L.) Griseb. var. cuspidata Baker [GU] 

Aechmea nudicaulis (L.) Griseb. var. nudicaulis [GU SU FG] 

Aechmea pallida L.B. Smith [GU] 

Aechmea penduliflora André [GU] 

Aechmea politii L.B. Smith [GU FG] 

Aechmea prancei L.B. Smith [to be expected in GU] 

Aechmea setigera Mart. ex Schult.f. [SU FG] 

Aechmea tillandsioides (Mart. ex Schult.f.) Baker [GU SU] 

Aechmea tocantina Baker [GU SU FG] 

Ananas ananassoides (Baker) L.B. Smith [GU SU FG] 

Ananas comosus (L.) Merrill [GU SU FG] [cultivated, 

naturalized] 

Ananas lucidus Mill. [FG] 

Ananas nanus (L.B. Smith) L.B. Smith [SU? FG?] 

Ananas parguazensis Camargo & L.B. Smith [GU SU] 

Araeococcus flagellifolius Harms [SU FG] 

Araeococcus goeldianus L.B. Smith [FG] 

Araeococcus micranthus Brongn. [GU SU FG] 

Billbergia brachysiphon L.B. Smith [to be expected in FG] 

Billbergia macrolepis L.B. Smith [GU] 

Billbergia pyramidalis (Sims) Lind. [FG] 

Billbergia rosea Hort. ex Beer [SU?] 

Billbergia violacea Beer [GU SU FG] 

Brocchinia acuminata L.B. Smith [to be expected in GU] 

Brocchinia hechtioides Mez [Mt. Roraima] 

Brocchinia melanacra L.B. Smith [to be expected in GU] 

Brocchinia micrantha (Baker) Mez [GU] 

Brocchinia reducta Baker [GU] 

Brocchinia steyermarkii L.B. Smith [GU] 

Brocchinia tatei L.B. Smith [GU] 

Bromelia agavifolia Brongn. ex Houllett [FG] 

Bromelia alta L.B. Smith [GU SU] 

Bromelia chrysantha Jacq. [to be expected in GU] 

Bromelia fosteriana L.B. Smith [SU] 

Bromelia goeldiana L.B. Smith [to be expected in GU] 

Bromelia humilis Jacq. [SU?] 

Bromelia morreniana (Regel) Mez [to be expected in FG] 

Bromelia oleveiriae L.B. Smith [FG?] 

Bromelia pinguin L. [GU SU] [cultivated only?] 

Bromelia plumieri (E. Morren) L.B. Smith [GU SU FG] 

Bromelia rondoniana L.B. Smith [to be expected in GU] 

Bromelia serra Griseb. [FG] 

Bromelia tubulosa L.B. Smith [FG?] 

Catopsis berteroniana (Schult.f.) Mez [GU SU FG] 

Catopsis sessiliflora (Ruiz & Pav.) Mez [SU FG] 

Connelia nutans L.B. Smith [to be expected in GU] 

Connellia augustae (Rich. Schomb.) N.E. Brown [GU] [Mt. 

Roraima] 

Connellia caricifolia L.B. Smith [Mt. Roraima] 

Connellia quelchii N.E. Brown [Mt. Roraima] 

Disteganthus basilateralis Lem. [FG] 

Disteganthus calatheoides (L.B. Smith) L.B. Smith & R.W. 

Read [FG] 

Disteganthus lateralis (L.B. Smith) Gouda [SU FG] 

Guzmania altsonii L.B. Smith [GU SU FG] 

Guzmania angustifolia (Baker) Wittmack [GU?] 

Guzmania brasiliensis Ule [possibly to be expected in GU] 

Guzmania calothyrsus Mez [GU] 

Guzmania erythrolepis Brongn. ex Planch. [FG?] [probably 

misident. of G. melinonis according to Gouda] 

SPERMATOPHYTA: Brassicaceae - Bromeliaceae 

67

68 


Guzmania lingulata (L.) Mez [GU SU FG] 

Guzmania melinonis Regel [GU SU FG] 

Guzmania plumieri (Griseb.) Mez [GU? SU? FG] 

Guzmania retusa L.B. Smith [GU] 

Guzmania roezlii (E. Morren) Mez [GU] 

Guzmania sphaeroidea (André) André ex Mez [GU] 

Guzmania squarrosa (Mez & Sodiro) L.B. Smith & Pittendrigh 

[GU] 

Guzmania steyermarkii L.B. Smith [to be expected in GU] 

Hohenbergia stellata Schult.f. [to be expected in GU] 

Lindmania geniculata L.B. Smith [GU] 

Lindmania gracillima (L.B. Smith) L.B. Smith [to be expected 

in GU] 

Lindmania guianensis (Beer) Mez [GU] 

Lindmania paludosa L.B. Smith [possibly to be expected in 

GU] 

Navia angustifolia (Baker) Mez [GU] 

Navia arida L.B. Smith & Steyerm. [GU] 

Navia barbellata L.B. Smith [GU] 

Navia brachyphylla L.B. Smith [GU?] 

Navia cataractarum Sandw. [GU] 

Navia duidae L.B. Smith [GU?] 

Navia gleasonii L.B. Smith [GU] 

Navia maguirei L.B. Smith [SU] 

Navia rupestris (Gleason) Sandw. [GU] 

Navia sandwithii L.B. Smith [GU] 

Navia splendens L.B. Smith [GU] 

Pitcairnia brittoniana Mez [GU] 

Pitcairnia caricifolia Mart. ex Schult.f. [GU SU FG] [=Pepinia 

caricifolia] 

Pitcairnia ctenophylla L.B. Smith [=Pepinia ctenophylla] [to 

be expected in GU] 

Pitcairnia geyskesii L.B. Smith [GU SU FG] [=Pepinia 

geyskesii] 

Pitcairnia heterophylla (Lindl.) Beer [to be expected in GU] 

Pitcairnia incarnata Baker [FG] [=Pepinia incarnata] 

Pitcairnia leprieurii Baker [GU FG] 

Pitcairnia maidifolia (C. Morren) Decne. [GU SU] 

Pitcairnia nuda Baker [GU SU] [=Pepinia nuda] 

Pitcairnia patentiflora L.B. Smith [FG] [=Pepinia patentiflora] 

Pitcairnia pusilla Mez [FG] 

Pitcairnia rubiginosa (Brongn.) Baker [FG] [=Pepinia 

rubiginosa] 

Pitcairnia sastrei L.B. Smith & R.W. Read [SU? FG] 

Pitcairnia sprucei Baker [FG] [=Pepinia sprucei] 

Streptocalyx longifolius (Rudge) Baker [SU FG] [=Aechmea 

longifolia] 

Streptocalyx poeppigii Beer [SU FG] [=Aechmea beeriana] 

Streptocalyx poitaei Baker [FG] [=Aechmea poitaei] 

Tillandsia adpressiflora Mez [GU? SU FG] 

Tillandsia anceps Lodd. [GU SU FG] 

Tillandsia balbisiana Schult.f. [possibly to be expected in GU] 

Tillandsia bulbosa Hook. [GU SU FG] 

Tillandsia compacta Griseb. [possibly to be expected in GU] 

Tillandsia complanata Benth. [GU] 

Tillandsia elongata Kunth var. subimbricata (Baker) L.B. 

Smith [GU?] 

Tillandsia fasciculata Swartz [GU? SU FG] 

Tillandsia fendleri Griseb. var. fendleri [GU] 

Tillandsia fendleri Griseb. var. reducta (L.B. Smith) L.B. 

Smith [possibly to be expected in GU] 

Tillandsia flexuosa Swartz [GU SU FG] 

Tillandsia geminiflora Brongn. [SU] 

Tillandsia jenmanii Baker [GU] [=Racinaea jenmanii] 

Tillandsia kegeliana Mez [GU SU FG] 

Tillandsia monadelpha (E. Morren) Baker [GU SU FG] 

SPERMATOPHYTA: Bromeliaceae 

Tillandsia paraensis Mez [GU SU FG] 

Tillandsia pruinosa Swartz [GU? SU] 

Tillandsia spiculosa Griseb. var. micrantha (Baker) L.B. Smith 

[GU] [=Racinaea spiculosa var. micrantha] 

Tillandsia spiculosa Griseb. var. spiculosa [GU SU FG] 

[=Racinaea spiculosa var. spiculosa] 

Tillandsia spiculosa Griseb. var. stenoglossa (L.B. Smith) 

Gouda [GU] [=Racinaea stenoglossa var. stenoglossa] 

Tillandsia stricta Solander ex Ker-Gawl. [GU SU] 

Tillandsia tenuifolia L. [GU SU FG] 

Tillandsia tetrantha Ruiz & Pav. var. caribaea (L.B. Smith) 

Gouda [GU?] [Mt. Roraima] [=Racinaea tetrantha var. 

caribaea] 

Tillandsia turneri Baker var. orientalis L.B. Smith [GU] 

Tillandsia usneoides (L.) L. [GU SU FG] 

Vriesea amazonica (Baker) Mez [SU FG] [=Werauhia 

gigantea] 

Vriesea capituligera (Griseb.) L.B. Smith & Pitt. 

[=Mezobromelia capituligera] [possibly to be expected in 

GU] 

Vriesea duidae (L.B. Smith) Gouda [GU] 

Vriesea gladioliflora (Wendl.) Antoine [GU SU FG] 

[=Werauhia gladioliflora] 

Vriesea heliconioides (Kunth) Hook. ex Walp. [GU? SU FG] 

Vriesea heterandra (André) L.B. Smith [SU] 

Vriesea hygrometrica (André) L.B. Smith & Pittendrigh 

[=Werauhia hygrometrica] [to be expected in GU] 

Vriesea incurva (Griseb.) R.W. Read [GU] [=Tillandsia 

incurva] 

Vriesea johnstonii (Mez) L.B. Smith & Pittendrigh [GU] 

[=Vriesea hygrometrica?] 

Vriesea jonghei (C. Koch) E. Morren [FG] 

Vriesea platynema Gaudichaud [GU] 

Vriesea pleiosticha (Griseb.) Gouda [GU SU FG] 

[=Mezobromelia pleiosticha] 

Vriesea procera (Mart. ex Schult.f.) Wittmack [GU SU FG] 

Vriesea ringens (Griseb.) Mez [GU] [=Werauhia ringens] 

Vriesea rubra (Ruiz & Pav.) Beer [GU] 

Vriesea soderstromii L.B. Smith [GU] 

Vriesea splendens (Brongn.) Lem. var. formosa Suringar ex 

Witte [GU] 

Vriesea splendens (Brongn.) Lem. var. splendens [GU SU 

FG] 

Vriesea viridiflora (Regel) Wittmack ex Mez [GU] 

[=Werauhia viridiflora] 

BROMELIACEAE - SYNONYMS AND 


Aechmea beeriana L.B. Smith & Spencer =Streptocalyx 

poeppigii 

Aechmea lateralis L.B. Smith =Disteganthus lateralis 

Aechmea longifolia (Rudge) L.B. Smith & Spencer 

=Streptocalyx longifolius 

Aechmea mucroniflora Hook. =Aechmea mertensii 

Aechmea poitaei (Baker) L.B. Smith & Spencer =Streptocalyx 

poitaei 

Aechmea spicata Mart. ex Schult.f. =Aechmea mertensii 

Aechmea tillandsioides (Mart. ex Schult.f.) Baker var. 

kienastii (E. Morren ex Mez) L.B. Smith =Aechmea 

tillandsioides


Ananas ananassoides (Baker) L.B. Smith var. nanus L.B. 

Smith =Ananas nanus 

Brocchinia bernardii L.B. Smith =Brocchinia melanacra 

Brocchinia paniculata Schult.f. [not in Guianas] 

Brocchinia secunda L.B. Smith =Brocchinia tatei 

Bromelia granvillei L.B. Smith [nomen nudum] 

Bromelia karatas L. =Bromelia plumieri 

Catopsis inconspicua Bak. =Catopsis nitida [not in Guianas] 

Chevaliera fernandae (E. Morren) L.B. Smith & W.J. Kress 

=Aechmea fernandae 

Chevaliera pallida (L.B. Smith) L.B. Smith & W.J. Kress 

=Aechmea pallida 

Cottendorfia geniculata (L.B. Smith) L.B. Smith =Lindmania 

geniculata 

Cottendorfia gracillima L.B. Smith =Lindmania gracillima 

Cottendorfia guianensis (Beer) Klotzsch ex Baker =Lindmania 

guianensis 

Cottendorfia paludosa (L.B. Smith) L.B. Smith =Lindmania 

paludosa 

Cottendorfia serrulata (L.B. Smith) L.B. Smith =Lindmania 

serrulata 

Disteganthus bromeliifolius L.B. Smith [nomen nudum] 

=Disteganthus lateralis 

Gravisia aquilega (Salisb.) Mez =Aechmea aquilega 

Gravisia lanjouwii (L.B. Smith) L.B. Smith =Aechmea 

lanjouwii 

Guzmania minor Mez =Guzmania lingulata 

Guzmania pleiosticha (Griseb.) Mez =Vriesea pleiosticha 

Guzmania splitgergeri Mez =Vriesea pleiosticha 

Guzmania venamensis L.B. Smith =Guzmania sphaeroidea 

Lamprococcus campanulatus (L.B. Smith) L.B. Smith & W.J. 

Kress =Aechmea campanulata 

Mezobromelia capituligera (Griseb.) J.R. Grant =Vriesea 

capituligera 

Mezobromelia pleiosticha (Griseb.) Utley & Luther =Vriesea 

pleiosticha 

Navia fluviatilis L.B. Smith =Navia rupestris 

Pepinia caricifolia (Mart. ex Schult.f.) Varadarajan & 

Gilmartin =Pitcairnia caricifolia 

Pepinia ctenophylla (L.B. Smith) Varadarajan & Gilmartin 

=Pitcairnia ctenophylla 

Pepinia geyskesii (L.B. Smith) Varadarajan & Gilmartin 

=Pitcairnia geyskesii 

Pepinia incarnata (Baker) Varadarajan & Gilmartin =Pitcairnia 

incarnata 

Pepinia nuda (Baker) Varadarajan & Gilmartin =Pitcairnia 

nuda 

Pepinia patentiflora (L.B. Smith) Varadarajan & Gilmartin 

=Pitcairnia patentiflora 

Pepinia rubiginosa (Brongn.) Varadarajan & Gilmartin 

=Pitcairnia rubiginosa 

Pepinia sprucei (Baker) Varadarajan & Gilmartin =Pitcairnia 

sprucei 

Pitcairnia kegeliana Schlecht. =Pitcairnia caricifolia 

Platyaechmea contracta (Mart. ex Schult.f.) L.B. Smith & W.J. 

Kress =Aechmea contracta 

Platyaechmea tillandsioides (Mart. ex Schult.f.) L.B. Smith & 

W.J. Kress =Aechmea tillandioides 

Pothuava nudicaulis (L.) L.B. Smith & W.J. Kress =Aechmea 

nudicaulis 

Quesnelia lamarckii Baker =Musa coccinea 

(Musaceae)[cultivated only] 

Racinaea jenmanii (Baker) Spencer & L.B. Smith =Tillandsia 

jenmanii 

Racinaea spiculosa (Griseb.) Spencer & L.B. Smith var. 

micrantha (Baker) Spencer & L.B. Smith =Tillandsia 

spiculosa var. micrantha 


spiculosa =Tillandsia spiculosa var. spiculosa 


stenoglossa (L.B. Smith) Spencer & L.B. Smith 

=Tillandsia spiculosa var. stenoglossa 

Racinaea tetrantha (Ruiz & Pav.) Spencer & L.B. Smith var. 

caribaea (L.B. Smith) Spencer & L.B. Smith =Tillandsia 

tetrantha var. caribaea 

Thecophyllum splitgerberi (Mez) Pittendrigh =Vriesea 

pleiosticha 

Tillandsia amazonica Baker =Vriesea amazonica 

Tillandsia appuniana Baker =Vriesea splendens var. splendens 

Tillandsia archeri L.B. Smith =Vriesea turneri 

Tillandsia compressa Bert. [not in Guianas] 

Tillandsia conspersa Miq. =Tillandsia stricta 

Tillandsia deppeana Steud. var. reducta (L.B. Smith) L.B. 

Smith =Tillandsia fendleri var. reducta 

Tillandsia duidae L.B. Smith =Vriesea duidae 

Tillandsia graminifolia Baker =Tillandsia monadelpha 

Tillandsia incurva Griseb. =Vriesea incurva 

Tillandsia multifolia Mez =Tillandsia turneri var. orientalis 

Tillandsia pachychlamys Mez =Vriesea gladioliflora 

Tillandsia parkeri Baker =Tillandsia spiculosa var. spiculosa 

Tillandsia pulchella Hook. =Tillandsia tenuifolia 

Tillandsia pulchella Hook. var. surinamensis Mez =Tillandsia 

tenuifolia 

Tillandsia punctulata Cham. & Schlechtend. [not in Guianas] 

Tillandsia rhodocincta Baker =Tillandsia turneri var. orientalis 

Tillandsia rhododactyla Mez =Vriesea rubra 

Tillandsia rigidula Baker =Guzmania roezlii 

Tillandsia rubra Ruiz & Pavon var. reducta L.B. Smith 

=Tillandsia fendleri var. reducta 

Tillandsia splendens Brongn. =Vriesea splendens 

Tillandsia stenoglossa L.B. Smith =Tillandsia spiculosa var. 

stenoglossa 

Tillandsia subimbricata Baker =Tillandsia elongata var. 

subimbricata 

Tillandsia surinamensis Miq. ex Mez =Tillandsia tenuifolia 

Tillandsia tenuifolia L. var. surinamensis (Mez) L.B. Smith 

=Tillandsia tenuifolia 

Tillandsia usneoides (L.) L. var. filiformis (André) Mez 

=Tillandsia usneoides 

Vriesea gigantea (Mart. ex Schult.f.) Mez =Vriesea amazonica 

Vriesea pachychlamys Mez =Vriesea gladioliflora 

Vriesea splitgerberi (Mez) L.B. Smith & Pittendrigh =Vriesea 

pleiosticha 

Werauhia gigantea (Mart. ex Schult.f.) J.R. Grant =Vriesea 

amazonica 

Werauhia gladioliflora (Wendl.) J.R. Grant =Vriesea 

gladioliflora 

Werauhia hygrometrica (André) J.R. Grant =Vriesea 

hygrometrica 

Werauhia ringens (Griseb.) J.R. Grant =Vriesea ringens 

Wittmackia odora Mez =Aechmea lingulata 

SPERMATOPHYTA: Bromeliaceae 

69

70 

REFERENCES: 


BRUNELLIACEAE 

Cuatrecasas, J. 1970. Brunelliaceae. Flora 


Brunellia comocladifolia Humb. & Bonpl. subsp. ptariana 

(Steyerm.) Cuatrecasas [Mt. Roraima] 

BURMANNIACEAE 

reviewed by P.J.M. Maas (1992); reviewed by 

P.J.M. Maas (1996) 

REFERENCES: 

Jonker, F.P. 1938. Burmanniaceae. In A. Pulle, 


Jonker, F.P. 1968. Burmanniaceae (additions 

and corrections). In A. Pulle & J. Lanjouw, 


Maas, P.J.M., H. Maas-van de Kamer, J. van 

Benthem, H.C.M. Snelders, & T. Rübsamen. 

1986. Burmanniaceae. Flora Neotropica 

Monogr. 42: 1-189. 

Maas, P.J.M. & H. Maas-van de Kamer. 1989. 

Burmanniaceae. In A.R.A. Görts-van Rijn, 


Maas, P.J.M. & H. Maas-van de Kamer. 1987. A 

new Thismia (Burmanniaceae) from French 

Guiana. Brittonia 39(3): 376-378. 

Apteria aphylla (Nutt.) Barnhart ex Small [GU SU FG] 

Burmannia bicolor Mart. [GU SU FG] 

Burmannia capitata (Walter ex J.F. Gmel.) Mart. [GU SU FG] 

Burmannia flava Mart. [GU SU FG] 

Burmannia tenella Benth. [GU] 

Campylosiphon purpurascens Benth. [GU SU FG] 

Cymbocarpa saccata Sandw. [GU] 

Dictyostega orobanchoides (Hook.) Miers subsp. 

orobanchoides [GU SU] 

Dictyostega orobanchoides (Hook.) Miers subsp. parviflora 

(Benth.) Snelders & Maas [GU SU FG] 

Gymnosiphon brachycephalus Snelders & Maas [GU SU] 

Gymnosiphon breviflorus Gleason [GU SU FG] 

Gymnosiphon capitatus (Benth.) Urb. [GU] 

Gymnosiphon cymosus (Benth.) Benth. & J.D. Hook. [SU] 

Gymnosiphon divaricatus (Benth.) Benth. & J.D. Hook. [GU 

SU FG] 

Gymnosiphon fimbriatus (Benth.) Urb. [GU] 

Gymnosiphon guianensis Gleason [GU SU] 

Gymnosiphon minutus Snelders & Maas [GU] 

Gymnosiphon recurvatus Snelders & Maas [GU FG?] 

Hexapterella gentianoides Urb. [GU SU] 

Miersiella umbellata (Miers) Urb. [GU FG] 

Thismia saulensis Maas & Maas [FG] 

BURMANNIACEAE - SYNONYMS AND 


SPERMATOPHYTA: Brunelliaceae - Burseraceae 

Apteria aphylla (Nuttall) Barnhart ex Small var. hymenanthera 

(Miq.) Jonk. =Apteria aphylla 

Apteria gracillima Sagot [nomen nudum] =Apteria aphylla 

Apteria hymenanthera Miq. =Apteria aphylla 

Burmannia brachystachya Miq. =Burmannia bicolor 

Dictyostega schomburgkii Miers =Dictyostega orobanchoides 

subsp. orobanchoides 

Dictyostega schomburgkii Miers var. parviflora Benth. 

=Dictyostega orobanchoides subsp. parviflora 

Gymnosiphon altsoni Gleason =Hexapterella gentianoides 

Gymnosiphon arcuatus Urb. =Gymnosiphon divaricatus 

Gymnosiphon cornutus (Benth.) Benth. & J.D. Hook. 

=Gymnosiphon divaricatus 

Gymnosiphon tenellus (Benth.) Urb. [not in Guianas] 

Miersiella aristata Sandw. =Miersiella umbellata 

BURSERACEAE 

reviewed by D. Daly (1992) 

REFERENCES: 

Daly, D. 1987. Studies in neotropical 

Burseraceae I. A synopsis of the genus 

Crepidospermum. Brittonia 39: 51-58. 


Burseraceae II. Generic limits in New World 

Protieae and Canarieae. Brittonia 41: 17-27. 


Burseraceae VI. New taxa and combinations 

in Protium Burm.f. Brittonia 44: 280-299. 

Swart, J.J. 1942. A monograph of the genus 

Protium and some allied genera. Rec. Trav. 

Bot. Neerl. 34: 211-446. 

Swart, J.J. 1951. Burseraceae. In A. Pulle, Flora 


Bursera simaruba (L.) Sarg. [GU FG] 

Crepidospermum goudotianum (Tul.) Triana & Planch. [GU 

FG] 

Crepidospermum rhoifolium (Benth.) Triana & Planch. [GU 

SU FG] 

Dacryodes belemensis Cuatrec. [SU?] 

Dacryodes cuspidata (Cuatrec.) Daly [FG] 

Dacryodes nitens Cuatrec. [GU SU FG] 

Dacryodes roraimensis Steyerm. & Maguire [GU] 

Dacryodes steyermarkii Sandw. [GU] 

Protium altsonii Sandw. [GU SU FG] 

Protium apiculatum Swart [SU FG] 

Protium aracouchini (Aubl.) March. [GU SU FG] 

Protium crassipetalum Cuatrec. [GU SU?] 

Protium crenatum Sandw. [GU] 

Protium cuneatum Swart [GU SU? FG] 

Protium decandrum (Aubl.) March. [GU SU FG] 

Protium demerarense Swart [GU FG] 

Protium divaricatum Engl. subsp. fumarium Daly [FG] 

Protium elegans Engl. [FG?]


Protium fimbriatum Swart [FG] 

Protium giganteum Engl. var. crassifolium (Engl.) Daly [SU 

FG] 

Protium giganteum Engl. var. giganteum [GU SU FG] 

Protium glabrescens Swart [SU?] 

Protium grandifolium Engl. [FG] 

Protium guianense (Aubl.) March. var. guianense [GU SU 

FG] 

Protium heptaphyllum (Aubl.) March. subsp. heptaphyllum 

[GU SU FG] 

Protium melinonis Engl. [FG] 

Protium morii Daly [FG] 

Protium nodulosum Swart [FG] 

Protium occultum Daly [FG] 

Protium opacum Swart subsp. rabelianum Daly [GU FG] 

Protium pallidum Cuatrec. [FG] 

Protium pilosum (Cuatrec.) Daly [FG] 

Protium plagiocarpum Benoist [GU? SU? FG] 

Protium poeppigianum Swart [FG] 

Protium polybotryum (Turcz.) Engl. subsp. polybotryum [GU 

SU FG] 

Protium pullei Swart [SU] [unverified name] 

Protium robustum (Swart) Porter [SU FG] 

Protium sagotianum March. [GU SU FG] 

Protium strumosum Daly [GU FG] 

Protium subserratum (Engl.) Engl. [GU FG] 

Protium tenuifolium (Engl.) Engl. [GU SU FG] 

Protium trifoliolatum Engl. [GU SU FG] 

Protium unifoliolatum Engl. [GU] 

Tetragastris altissima (Aubl.) Swart [GU SU FG] 

Tetragastris hostmannii (Engl.) Kuntze [GU SU FG] 

Tetragastris panamensis (Engl.) Kuntze [GU SU FG] 

Trattinnickia burserifolia Mart. [GU SU FG] 

Trattinnickia demerarae Sandw. [GU SU FG] 

Trattinnickia glaziovii Swart [FG] 

Trattinnickia rhoifolia Willd. [GU SU FG] 

BURSERACEAE - SYNONYMS AND 


Bursera velutinifolia Cowan =Cyrtocarpa velutinifolia 

(Anacardiaceae) 

Hemicrepidospermum rhoifolium (Benth.) Swart 

=Crepidospermum rhoifolium 

Icica altissima Aubl. =Tetragastris altissima 

Icica aracouchini Aubl. =Protium aracouchini 

Icica decandra Aubl. =Protium decandrum 

Icica enneandra Aubl. =Protium decandrum 

Icica guianensis Aubl. =Protium guianense 

Icica heptaphylla Aubl. =Protium heptaphyllum 

Protium araguense Cuatrec. [not in Guianas] 

Protium crassifolium Engl. =Protium giganteum var. 

crassifolium 

Protium firmum Swart =Protium apiculatum 

Protium hostmannii (Miq.) Engl. =Protium guianense var. 

guianense 

Protium insigne (Planch. & Triana) Engl. =Protium 

sagotianum 

Protium neglectum Swart =Protium tenuifolium 

Protium octandrum Swart =Protium heptaphyllum subsp. 

heptaphyllum 

Protium schomburgkianum Engl. =Protium decandrum 

Tetragastris pilosa Cuatrec. =Protium pilosum 

Trattinnickia cuspidata Cuatrec. =Dacryodes cuspidata 

CABOMBACEAE 

additions and corrections by J.H. Wiersema 

(1996) 

REFERENCES: 

Cramer, J. 1979. Cabombaceae. In A.L. Stoffers 

& J.C. Lindeman, Flora of Suriname 5(1): 

385-389. 

Fassett, N.C. 1953. A monograph of Cabomba. 

Castanea 18: 116-128. 

Oergaard, M., H.W.E. van Bruggen, & P.J. van 

der Vlugt. 1992. Die Familie Cabombaceae 

(Cabomba und Brasenia). Aqua-Planta 

sonderheft nr. 3. 

Wiersema, J.H. 1989. A new species of 

Cabomba from tropical America. Ann. 


Brasenia schreberi J.F. Gmelin [GU] 

Cabomba aquatica Aubl. [GU SU FG] 

Cabomba furcata Schult. & Schult.f. [GU SU?] 

Cabomba haynesii Wiersema [to be expected in GU] 

CABOMBACEAE - SYNONYMS AND 


Cabomba piauhyensis Gardn. var. albida Fassett =Cabomba 

haynesii 

Cabomba piauhyensis Gardn. var. piauhyensis =Cabomba 

furcata 

Cabomba warmingii Caspary =Cabomba furcata 

CACTACEAE 

contributed by B.E. Leuenberger (1992); 

additions and corrections by B.E. Leuenberger 

(1996) 

REFERENCES: 

Hunt, D. 1992. CITES Cactaceae Checklist. 

Royal Botanic Gardens, Kew. 

Leuenberger, B.E. 1987. A preliminary list of 

Cactaceae from the Guianas and 

recommendations for future collecting and 

preparation of specimens. Willdenowia 16: 

497-510. 

Leuenberger, B.E. 1989. Lectotypification of 

Cereus hexagonus (L.) Miller and taxonomic 

notes on related taxa from South America. 

Bot. Jahrb. Syst. 111(2): 145-164. 

Leuenberger, B.E. 1995 (in press). Cactaceae. 


Guianas (series A). 

Taylor, N.P. 1991. The genus Melocactus in 

Central and South America. Bradleya 9: 1-80. 

SPERMATOPHYTA: Burseraceae - Cactaceae 

71

72 


Cereus hexagonus (L.) Mill. [GU SU FG] 

Epiphyllum phyllanthus (L.) Haw. var. phyllanthus [GU SU 

FG] 

Hylocereus lemairei (Hook.) Britton & Rose [GU? SU? FG?] 

Hylocereus scandens (Salm-Dyck) Backeberg [GU SU] 

Melocactus neryi Schum. [GU SU] 

Melocactus smithii (Alexander) Buining [GU] 

Opuntia cochenillifera (L.) Mill. [SU FG] [cultivated only?] 

Pereskia aculeata Mill. [GU SU FG] [widely cultivated but 

apparently native in Guianas] 

Pilosocereus oligolepis (Vaupel) Byles & Rowley [GU] 

Rhipsalis baccifera (J. Mill.) W.T. Stearn [GU SU FG] 

Rhipsalis floccosa Pfeiffer [SU] 

Rhipsalis oblonga Löfgren [SU] 

Rhipsalis pachyptera Pfeiff. [GU? FG?] 

Selenicereus sp. sect. Salmdyckia [GU? SU? FG] 

CACTACEAE - SYNONYMS AND 


Cephalocereus kanukuensis Alexander =Pilosocereus 

oligolepis 

Cereus curtisii Otto [Guianan records dubious] 

Cereus grandiflorus Mill. [Guianan records dubious] 

Cereus jamacaru DC. [Guianan records dubious] 

Cereus longiflorus Alexander =Cereus hexagonus 

Cereus peruvianus Auct. =Cereus uruguayanus [Guianan 

records dubious] 

Cereus triangularis (L.) Haw. =Hylocereus triangularis (L.) 

Britton & Rose [Guianan records dubious] 

Epiphyllum hookeri Haw. =Epiphyllum phyllanthus var. 

hookeri [Guianan records dubious] 

Epiphyllum phyllanthus (L.) Haw. var. hookeri (Haw.) 

Kimnach [Guianan records dubious] 

Hylocereus triangularis (L.) Britton & Rose [Guianan records 

dubious] 

Melocactus schulzianus Buining =Melocactus neryi 

Nopalea cochenillifera (L.) Salm-Dyck =Opuntia 

cochenillifera 

Phyllocactus phyllanthus (DC.) Link =Epiphyllum phyllanthus 

Pilosocereus kanukuensis (Alexander) Leuenberger 

=Pilosocereus oligolepis 

Rhipsalis cassutha Gaertn. =Rhipsalis baccifera 

Rhipsalis minutiflora K. Schum. =Rhipsalis baccifera 

Rhipsalis occidentalis Barthlott & Rauh [not in Guianas] 

CAESALPINIACEAE 

Cassia, Chamaecrista, Dialium, Dicorynia, 

Dinizia, Martiodendron, & Senna contributed 

by R.C. Barneby (1992) 

REFERENCES: 

Amshoff, G.J.H. 1939. Papilionaceae. In A. 


Cowan, R.S. 1961. Leguminosae. In B. 




Cowan, R.S. 1968. Swartzia. Flora Neotropica 

Monogr. 1: 1-228. 

SPERMATOPHYTA: Cactaceae - Caesalpiniaceae 

Cowan, R.S. & J.C. Lindeman. 1989. 

Caesalpiniaceae p.p. In A.R.A. Görts-van 

Rijn, Flora of the Guianas (series A) 7: 1-167. 

Dwyer, J.D. 1951. The Central American, West 

Indian, and South American species of 

Copaifera (Caesalpiniaceae). Brittonia 7(3): 

143-172. 

Dwyer, J.D. 1954. The tropical American genus 

Tachigalia Aubl. (Caesalpiniaceae). Ann. 


Dwyer, J.D. 1957. The tropical American genus 

Sclerolobium Vogel (Caesalpiniaceae). 

Lloydia 20: 67-118, suppl.266-267. 

Dwyer, J.D. 1958. The New World species of 

Cynometra. Ann. Missouri Bot. Gard. 45: 

313-345. 

Fonseca Vaz, A.M.S. da. 1979. Consideraçoes 

sobre a taxonomia do gênero Bauhinia L. sect. 

Tylotaea Vogel (Leguminosae - 

Caesalpinioideae) do Brasil. Rodriguésia 

31(51): 127-234. 

Görts-van Rijn, A.R.A., K.U. Kramer, & J.C. 

Lindeman. 1976. Papilionaceae (additions 

and corrections). In J. Lanjouw & A.L. 

Stoffers, Flora of Suriname 2(2): 556-610. 

Irwin, H.S. & R.C. Barneby. 1982. The 

American Cassiinae. A synoptical revision of 

Leguminosae tribe Cassieae subtribe 

Cassiinae in the New World. Mem. New 

York Bot. Gard. 35: 1-918. 

Koeppen, R.C. 1967. Revision of Dicorynia. 

Brittonia 19(1): 42-61. 

Lee, Y.T. & J.H. Langenheim. 1975. 

Systematics of the genus Hymenaea L. Univ. 

Calif. Publ. Bot. 69: 1-109. 

Polhill, R.M. & P.H. Raven [ed.]. 1981. 

Advances in Legume Systematics. Royal 

Botanical Garden, Kew, England. 

Silva, M.F. da. 1976. Revisao taxonômica do 

gênero Peltogyne Vog. Acta Amaz. 6(1) 

suppl.: 1-61. 

Silva, M.F. da. 1986. Dimorphandra. Flora 


Aldina insignis (Benth.) Endl. var. insignis [GU] 

Aldina insignis (Benth.) Endl. var. retusa Cowan [GU] 

Aldina latifolia Spruce ex Benth. var. latifolia [to be expected 

in GU] 

Aldina latifolia Spruce ex Benth. var. pubescens Cowan [to 


Batesia floribunda Spruce ex Benth. [FG] 

Bauhinia aculeata L. [GU] 

Bauhinia bicuspidata Benth. [SU] 

Bauhinia breviloba Benth. [SU] 

Bauhinia cinnamomea DC. [GU SU FG] 

Bauhinia confertiflora Benth. [GU] 

Bauhinia coronata Benth. [FG] 

Bauhinia cupreonitens Ducke [GU SU FG] 

Bauhinia eilertsii Pulle [SU] 

Bauhinia glabra Jacq. [GU SU] 

Bauhinia guianensis Aubl. [GU SU FG]


Bauhinia kunthiana Vogel [GU SU FG] 

Bauhinia longicuspis Spruce ex Benth. [FG] 

Bauhinia longipetala (Benth.) Walp. [GU] 

Bauhinia macrostachya Benth. [GU] [=Bauhinia ungulata?] 

Bauhinia outimouta Aubl. [FG] 

Bauhinia poiteauana Vogel [FG] [=Bauhinia smilacina?] 

Bauhinia richardiana DC. [FG] 

Bauhinia rubiginosa Bong. [GU SU FG] [=Bauhinia 

guianensis?] 

Bauhinia scala-simiae Sandw. [GU] 

Bauhinia siqueiraei Ducke [GU FG] 

Bauhinia smilacina (Schott) Steud. [SU FG] 

Bauhinia sp. cf. rutilans Spruce ex Benth. [GU] 

Bauhinia splendens Kunth [SU FG] 

Bauhinia surinamensis Amsh. [GU SU FG] [=Bauhinia 

guianensis?] 

Bauhinia ungulata L. [GU] 

Bocoa alterna (Benth.) Cowan [GU] 

Bocoa prouacensis Aubl. [SU FG] 

Bocoa viridiflora (Ducke) Cowan [GU SU FG] 

Brownea guianensis Klotzsch [GU] 

Brownea latifolia Jacq. [GU] 

Caesalpinia bonduc (L.) Roxb. [GU SU FG] 

Caesalpinia pulcherrima (L.) Swartz [GU SU FG] [cultivated; 

naturalized?] 

Campsiandra comosa Benth. var. comosa [GU SU] 

Campsiandra comosa Benth. var. laurifolia (Benth.) Cowan 

[GU] 

Candolleodendron brachystachyum (DC.) Cowan [SU FG] 

Cassia cowanii Irwin & Barneby var. cowanii [FG] 

Cassia cowanii Irwin & Barneby var. guianensis (Sandw.) 

Irwin & Barneby [GU SU] 

Cassia fastuosa Willd. ex Benth. var. fastuosa [SU FG] 

Cassia fistula L. [GU FG] [widely cultivated, occasionally 

escaping] 

Cassia grandis L.f. [GU SU FG] 

Cassia moschata Kunth [GU] 

Cassia spruceana Benth. [FG] 

Chamaecrista adiantifolia (Benth.) Irwin & Barneby var. 

pteridophylla (Sandw.) Irwin & Barneby [GU] [to be 

expected in SU, FG] 

Chamaecrista apoucouita (Aubl.) Irwin & Barneby [GU SU 

FG] 

Chamaecrista calycioides (DC. ex Collad.) Greene [GU? FG] 

Chamaecrista desvauxii (Collad.) Killip var. brevipes (Benth.) 

Irwin & Barneby [GU SU?] 

Chamaecrista desvauxii (Collad.) Killip var. mollissima 

(Benth.) Irwin & Barneby [GU SU? FG?] 

Chamaecrista desvauxii (Collad.) Killip var. saxatilis (Amsh.) 

Irwin & Barneby [SU FG] 

Chamaecrista diphylla (L.) Greene [GU SU FG] 

Chamaecrista fagonioides (Vogel) Irwin & Barneby [GU SU 

FG] 

Chamaecrista flexuosa (L.) Greene [GU SU FG] 

Chamaecrista hispidula (Vahl) Irwin & Barneby [GU SU FG] 

Chamaecrista nictitans (L.) Moench var. disadena (Steud.) 

Irwin & Barneby [GU SU FG] 

Chamaecrista nictitans (L.) Moench var. glabrata (Vogel) 

Irwin & Barneby [GU] [widely naturalized] 

Chamaecrista nictitans (L.) Moench var. paraguariensis (Chod. 

& Hassl.) Irwin & Barneby [GU] 

Chamaecrista nictitans (L.) Moench var. patellaria (Collad.) 

Barneby, comb. nov. ined. [GU SU FG] 

Chamaecrista nictitans (L.) Moench var. praetexta (Vogel) 

Irwin & Barneby [GU SU FG] 

Chamaecrista polystachya (Benth.) Irwin & Barneby [GU] 

Chamaecrista ramosa (Vogel) Irwin & Barneby [GU SU FG] 

Chamaecrista roraimae (Benth.) Gleas. [GU] 

Chamaecrista rotundifolia (Pers.) Greene [GU?] [common 

weed to be expected in the Guianas] 

Chamaecrista serpens (L.) Greene var. serpens [GU] 

Chamaecrista viscosa (Kunth) Irwin & Barneby var. major 

(Benth.) Irwin & Barneby [GU SU] 

Chamaecrista viscosa (Kunth) Irwin & Barneby var. viscosa 

[to be expected in GU?] 

Copaifera epunctata Amsh. [SU] 

Copaifera guianensis Desf. [GU SU FG] 

Copaifera langsdorffii Desf. [GU] 

Copaifera officinalis L. [GU?] 

Copaifera pubiflora Benth. [GU] 

Copaifera reticulata Ducke [SU] 

Crudia amazonica Spruce ex Benth. [FG] 

Crudia aromatica (Aubl.) Willd. [SU FG] 

Crudia bracteata Benth. [FG] 

Crudia glaberrima (Steud.) Macbride [GU SU FG] 

Crudia oblonga Benth. [GU FG] 

Crudia pubescens Spruce ex Benth. [FG] 

Crudia spicata (Aubl.) Willd. [SU FG] 

Crudia tomentosa (Aubl.) Macbr. [GU FG] 

Cynometra bauhiniifolia Benth. var. bauhiniifolia [GU SU] 

Cynometra hostmanniana Tul. [GU SU FG] 

Cynometra marginata Benth. var. guianenis Dwyer [GU SU] 

Cynometra marginata Benth. var. marginata [GU SU FG] 

Cynometra parvifolia Tul. [SU FG] 

Cynometra spruceana Benth. var. spruceana [GU] 

Delonix regia (Bojer ex Hook.) Raf. [GU SU FG] [cultivated 

only?] 

Dialium guianense (Aubl.) Sandw. [GU SU FG] 

Dicorynia guianensis Amsh. [GU SU FG] 

Dicorynia paraensis Benth. [FG?] 

Dicymbe altsonii Sandw. [GU] 

Dicymbe bernardii Cowan [GU] 

Dicymbe corymbosa Spruce ex Benth. [GU] 

Dicymbe fraterna Cowan [GU] 

Dicymbe jenmanii Sandw. [GU] 

Dicymbe pharangophila Cowan [GU] 

Dicymbe uaiparuensis Cowan [to be expected in GU] 

Dimorphandra conjugata (Splitg.) Sandw. [GU SU] 

Dimorphandra cuprea Sprague & Sandw. subsp. cuprea [GU] 

Dimorphandra davisii Sprague & Sandw. [GU] 

Dimorphandra gonggrijpii Kleinh. [GU] 

Dimorphandra macrostachya Benth. subsp. congestiflora 

(Sprague & Sandw.) M.F. da Silva [GU] 

Dimorphandra macrostachya Benth. subsp. glabrifolia (Ducke) 

M.F. da Silva [GU FG] 

Dimorphandra macrostachya Benth. subsp. macrostachya [GU 

FG?] 

Dimorphandra polyandra Benoist [GU SU FG] 

Dimorphandra pullei Amsh. [SU FG] 

Dimorphandra williamii M.F. da Silva [GU] 

Dinizia excelsa Ducke [GU] [to be expected in FG] 

Elizabetha bicolor Ducke [FG?] 

Elizabetha coccinea Rich. Schomb. ex Benth. var. coccinea 

[GU] 

Elizabetha coccinea Rich. Schomb. ex Benth. var. oxyphylla 

(Harms) Cowan [GU SU] 

Elizabetha fanshawei Cowan [GU] 

Elizabetha grahamii Cowan [GU] 

Elizabetha macrostachya Benth. [GU SU] 

Elizabetha paraensis Ducke [SU? FG?] 

Elizabetha princeps Rich. Schomb. ex Benth. [GU SU FG] 

Eperua bijuga Mart. ex Benth. [FG] 

SPERMATOPHYTA: Caesalpiniaceae 

73

74 


Eperua falcata Aubl. [GU SU FG] 

Eperua glabra Cowan [GU FG?] 

Eperua grandiflora (Aubl.) Benth. subsp. grandiflora [SU FG] 

Eperua grandiflora (Aubl.) Benth. subsp. guyanensis Cowan 

[GU FG] 

Eperua jenmanii Oliver subsp. jenmanii [GU SU? FG] 

Eperua jenmanii Oliver subsp. sandwithii Cowan [GU] 

Eperua rubiginosa Miq. var. grandiflora Pulle [SU] 

Eperua rubiginosa Miq. var. rubiginosa [GU SU FG] 

Eperua schomburgkiana Benth. [GU SU FG] 

Eperua venosa Cowan [to be expected in GU] 

Exostyles sp. [SU] 

Haematoxylum campechianum L. [GU SU FG] [cultivated 

only?] 

Heterostemon ingifolius Sandw. [GU] 

Heterostemon mazarunensis Sandw. [GU] 

Heterostemon mimosoides Desf. var. complanatus Cowan 

[SU] 

Heterostemon mimosoides Desf. var. mimosoides [GU SU 

FG] 

Heterostemon otophorus Sandw. [GU] 

Hymenaea courbaril L. var. courbaril [GU SU FG] 

Hymenaea oblongifolia Huber var. davisii (Sandw.) Lee & 

Langenh. [GU] 

Hymenaea oblongifolia Huber var. oblongifolia [GU FG?] 

Hymenaea oblongifolia Huber var. palustris (Ducke) Lee & 

Langenh. [GU] [to be expected in SU, FG] 

Jacqueshuberia brevipes Barneby [GU] 

Lecointea amazonica Ducke [GU SU FG] 

Macrolobium acaciifolium (Benth.) Benth. [GU SU FG] 

Macrolobium acrothamnos Cowan [to be expected in GU] 

Macrolobium amplexans (Amsh.) Cowan [SU] 

Macrolobium angustifolium (Benth.) Cowan [GU SU FG] 

Macrolobium bifolium (Aubl.) Pers. [GU SU FG] 

Macrolobium campestre Huber var. arboreum Cowan [GU SU 

FG] 

Macrolobium furcatum Ducke [to be expected in GU] 

Macrolobium guianense (Aubl.) Pulle [SU FG] 

Macrolobium huberianum Ducke var. huberianum [GU] 

Macrolobium huberianum Ducke var. pubirachis Amsh. [GU] 

Macrolobium jenmanii (Gleason) Sandw. [GU] 

Macrolobium limbatum Spruce ex Benth. var. fanshawei 

(Cowan) Cowan [GU] 

Macrolobium longeracemosum Amsh. [GU SU] 

Macrolobium montanum Ducke var. potaroanum Cowan [GU] 

Macrolobium multijugum (DC.) Benth. var. multijugum [GU 

SU FG] 

Macrolobium pendulum Willd. ex Vogel [GU SU FG] 

Macrolobium stenopetalum Amsh. [SU] 

Macrolobium suaveolens Spruce ex Benth. var. pakarimense 

Cowan [GU] 

Macrolobium suaveolens Spruce ex Benth. var. suaveolens 

[GU] 

Macrolobium venulosum Benth. [possibly to be expected in 

GU] 

Martiodendron excelsum (Benth.) Gleason [GU] 

Martiodendron parviflorum (Amsh.) Koeppen [SU FG] 

Mora excelsa Benth. [GU SU] 

Mora gonggrijpii (Kleinh.) Sandw. [GU SU] 

Paloue brasiliensis Ducke [FG] 

Paloue guianensis Aubl. [GU SU FG] 

Paloue induta Sandw. [GU] 

Paloue riparia Pulle [GU SU FG] 

Paloveopsis emarginata Cowan [GU] 

Parkinsonia aculeata L. [SU] [cultivated only?] 

Peltogyne floribunda (Kunth) Benth. [GU] 


Peltogyne lecointei Ducke [FG?] 

Peltogyne paniculata Benth. subsp. paniculata [GU SU FG] 

Peltogyne paniculata Benth. subsp. pubescens (Benth.) M. F. 

da Silva [GU SU FG] 

Peltogyne venosa (Vahl) Benth. subsp. densiflora (Spruce ex 

Benth.) M. F. da Silva [GU SU FG] 

Peltogyne venosa (Vahl) Benth. subsp. venosa [GU SU FG] 

Peltophorum dubium (Spreng.) Taub. [GU] 

Peltophorum inerme (Rosb.) Naves [GU] 

Peltophorum pterocarpum (DC.) Backer ex K. Heyne [GU SU 

FG] [cultivated only?] 

Recordoxylon speciosum (Benoist) Gazel ex Barneby [GU 

FG] 

Sclerolobium albiflorum Benoist [SU FG] 

Sclerolobium amplifolium Ducke [FG] 

Sclerolobium bracteosum Harms [SU?] 

Sclerolobium guianense Benth. var. guianense [GU SU FG] 

Sclerolobium melinonii Harms [SU FG] 

Sclerolobium micropetalum Ducke [GU SU] 

Sclerolobium paraense Huber [SU FG] 

Sclerolobium roraimae Schomb. [GU?] [Mt. Roraima] 

Sclerolobium tinctorium Benth. [FG] 

Senna alata (L.) Roxb. [GU SU FG] 

Senna bacillaris (L.f.) Irwin & Barneby var. bacillaris [GU 

SU FG] 

Senna bicapsularis (L.) Roxb. [GU FG] 

Senna chrysocarpa (Desv.) Irwin & Barneby [GU SU FG] 

Senna georgica Irwin & Barneby [FG] 

Senna hirsuta (L.) Irwin & Barneby var. hirsuta [GU SU FG] 

Senna latifolia (G. Mey.) Irwin & Barneby [GU SU FG] 

Senna ligustrina (L.) Irwin & Barneby [GU?] 

Senna lourteigiana Irwin & Barneby [SU? FG] 

Senna macranthera (DC. ex Collad.) Irwin & Barneby 

[cultivated, perhaps escaping] 

Senna multijuga (L.C. Rich.) Irwin & Barneby var. multijuga 

[GU SU FG] 

Senna obtusifolia (L.) Irwin & Barneby [GU SU FG] 

Senna occidentalis (L.) Link [GU SU FG] 

Senna pallida (Vahl) Irwin & Barneby var. pallida [to be 


Senna pilifera (Vogel) Irwin & Barneby var. subglabra (S. 

Moore) Irwin & Barneby [to be expected in the Guianas] 

Senna polyphylla (Jacq.) Irwin & Barneby [cultivated, perhaps 

escaping] 

Senna quinquangulata (L.C. Rich.) Irwin & Barneby [GU SU 

FG] 

Senna reticulata (Willd.) Irwin & Barneby [GU SU FG] 

Senna rupununiensis Irwin & Barneby [GU] 

Senna sandwithiana Irwin & Barneby [GU] 

Senna siamea (Lam.) Irwin & Barneby [cultivated, 

occasionally escaping] 

Senna silvestris (Vell.) Irwin & Barneby var. silvestris [GU 

SU] [to be expected in FG] 

Senna sophera (L.) Roxb. [GU] 

Senna undulata (Benth.) Irwin & Barneby [GU] 

Senna velutina (Vogel) Irwin & Barneby [GU] 

Swartzia amshoffiana Cowan [SU FG] 

Swartzia anomala Cowan [GU] 

Swartzia apiculata Cowan [GU] 

Swartzia aptera DC. [FG] 

Swartzia arborescens (Aubl.) Pittier [GU SU FG] 

Swartzia bannia Sandw. [GU SU] 

Swartzia benthamiana Miq. var. benthamiana [GU SU FG] 

Swartzia brachyrachis Harms var. glabrata Cowan [GU SU] 

Swartzia coarctata Jackson [FG?] 

Swartzia conferta Spruce ex Benth. var. conferta [GU]


Swartzia davisii Sandw. [GU] 

Swartzia dipetala Willd. ex Vogel [GU] 

Swartzia eriocarpa Benth. [GU] 

Swartzia fanshawei Cowan [GU] 

Swartzia grandifolia Bong. ex Benth. [GU SU FG] 

Swartzia guianensis (Aubl.) Urb. [GU SU FG] 

Swartzia hostmannii Benth. [SU FG] 

Swartzia jenmanii Sandw. [GU] 

Swartzia katawa Cowan [GU] 

Swartzia laevicarpa Amsh. [GU SU FG?] 

Swartzia lamellata Ducke var. kaieteurensis Cowan [GU] 

Swartzia latifolia Benth. var. latifolia [GU] 

Swartzia latifolia Benth. var. sylvestris Cowan [GU SU] 

Swartzia laurifolia Benth. [to be expected in GU] 

Swartzia laxiflora Bong. ex Benth. [to be expected in the 


Swartzia leblondii Cowan [FG] 

Swartzia leiocalycina Benth. [GU] 

Swartzia leiogyne (Sandw.) Cowan [GU] 

Swartzia longicarpa Amsh. [SU FG] 

Swartzia longipedicellata Sandw. [GU] 

Swartzia oblanceolata Sandw. [GU FG] 

Swartzia oblonga Benth. [to be expected in GU] 

Swartzia pachyphylla Harms [Mt. Roraima] 

Swartzia panacoco (Aubl.) Cowan var. altsonii (Sandw.) 

Cowan [GU] 

Swartzia panacoco (Aubl.) Cowan var. cardonae (Cowan) 

Cowan [GU] 

Swartzia panacoco (Aubl.) Cowan var. kamarangensis Cowan 

[GU] 

Swartzia panacoco (Aubl.) Cowan var. panacoco [FG] 

Swartzia panacoco (Aubl.) Cowan var. polyanthera (Steud.) 

Cowan [GU SU FG] 

Swartzia panacoco (Aubl.) Cowan var. sagotii (Sandw.) 

Cowan [FG] 

Swartzia panacoco (Aubl.) Cowan var. sandwithiana (Cowan) 

Cowan [GU] 

Swartzia panacoco (Aubl.) Cowan var. tepuiensis (Schery) 

Cowan [to be expected in GU] 

Swartzia parvifolia Schery [to be expected in GU] 

Swartzia piarensis Cowan [to be expected in GU] 

Swartzia picta Spruce ex Benth. var. bolivarensis Cowan [to 

be expected in GU, possibly in SU] 

Swartzia polyphylla DC. [GU SU FG] 

Swartzia recurva Poepp. & Endl. [to be expected in GU?] 

Swartzia rediviva Cowan [SU] 

Swartzia remiger Amsh. [GU? SU FG?] 

Swartzia reticulata Ducke [SU] [to be expected in GU] 

Swartzia roraimae Sandw. [Mt. Roraima] 

Swartzia schomburgkii Benth. var. guayanensis Cowan [to be 


Swartzia schomburgkii Benth. var. schomburgkii [GU SU] 

Swartzia sericea Vogel var. sericea [GU? FG] 

Swartzia sprucei Benth. var. tessellata Cowan [GU] 

Swartzia steyermarkii Cowan [GU SU? FG] 

Swartzia stipellata Cowan [to be expected in GU] 

Swartzia tessmannii Harms [to be expected in GU] 

Swartzia tillettii Cowan [GU] 

Swartzia wurdackii Cowan [to be expected in GU] 

Swartzia xanthopetala Sandw. [GU] 

Tachigali bracteolata Dwyer [FG] 

Tachigali cavipes (Spruce ex Benth.) Macbr. [FG?] 

Tachigali myrmecophila (Ducke) Ducke [SU FG] 

Tachigali paniculata Aubl. var. angustifolia (Miq.) Dwyer [SU 

FG] 

Tachigali paniculata Aubl. var. paniculata [GU SU FG] 

Tachigali paniculata Aubl. var. sulcata (Benoist) Dwyer [FG] 

Tachigali pubiflora Benth. [GU] 

Tachigali rusbyi Harms [GU] 

Vouacapoua americana Aubl. [GU? SU FG] 

Vouacapoua macropetala Sandw. [GU] 

Zollernia paraënsis Huber [GU SU FG] 

CAESALPINIACEAE - SYNONYMS AND 


Allania insignis Benth. =Aldina insignis 

Apalatoa spicata Aubl. =Crudia spicata 

Arouna guianensis Aubl. =Dialium guianense 

Bauhinia brachystachya Benth. =Bauhinia glabra 

Bauhinia cumanensis Kunth =Bauhinia glabra 

Bauhinia guianensis Aubl. var. splendens (Kunth) Amsh. 

=Bauhinia splendens 

Bauhinia speciosa Vog., non Roxb. =Bauhinia rubiginosa 

Campsiandra laurifolia Benth. =Campsiandra comosa var. 

laurifolia 

Cassia alata L. =Senna alata 

Cassia apoucouita Aubl. =Chamaecrista apoucouita 

Cassia biflora L. [ambiguous name, probably applied to Senna 

pallida] 

Cassia calycioides DC. ex Collad. =Chamaecrista calycioides 

Cassia chrysocarpa Desv. =Senna chrysocarpa 

Cassia cultrifolia Kunth =Chamaecrista diphylla 

Cassia desvauxii Collad. =Chamaecrista desvauxii 

Cassia diphylla L. =Chamaecrista diphylla 

Cassia fagonoides Vogel =Chamaecrista fagonoides 

Cassia flexuosa L. =Chamaecrista flexuosa 

Cassia fruticosa Mill. =Senna fruticosa (Mill.) Irwin & 

Barneby [not in Guianas] 

Cassia glandulosa L. var. swartzii (Wikstr.) Macbr. 

=Chamaecrista glandulosa var. swartzii [not in Guianas] 

Cassia hirsuta L. =Senna hirsuta 

Cassia hispidula Vahl =Chamaecrista hispidula 

Cassia hoffmannseggii Mart. ex Benth. =Senna latifolia 

Cassia latifolia =Senna latifolia 

Cassia leiandra Benth. [Guianan records erroneous, but 

possibly to be expected] 

Cassia leiandra Benth. var. guianensis Sandw. =Cassia 

cowanii var. guianensis 

Cassia ligustrina L. =Senna ligustrina 

Cassia lucens Vogel =Senna silvestris var. silvestris 

Cassia macranthera DC. ex Collad. =Senna macranthera 

Cassia mimosoides L. =Chamaecrista mimosoides [not in 


Cassia multijuga L.C. Rich. =Senna multijuga 

Cassia occidentalis L. =Senna occidentalis 

Cassia paramariboensis Miq. =Aeschynomene sensitiva 

(Fabaceae) 

Cassia patellaria DC. =Chamaecrista nictitans var. patellaria 

Cassia pennelliana Amsh. =Chamaecrista nictitans var. 

paraguariensis [not in Guianas] 

Cassia polyphylla Jacq. =Senna polyphylla 

Cassia praetexta Vogel =Chamaecrista nictitans var. praetexta 

Cassia pteridophylla Sandw. =Chamaecrista adiantiflora var. 

pteridophylla 

Cassia quinquangulata Rich. =Senna quinquangulata 

Cassia racemosa Mill. =Senna racemosa (Mill.) Irwin & 

Barneby [not in Guianas] 

Cassia ramosa Vogel =Chamaecrista ramosa 

Cassia reticulata Willd. =Senna reticulata 

Cassia sagotiana Benth. =Cassia spruceana 


75

76 


Cassia saxatilis (Amsh.) Irwin =Chamaecrista desvauxii var. 

saxatilis 

Cassia siamea Lam. =Senna siamea 

Cassia stenocarpa Vogel =Chamaecrista nictitans var. 

disadena 

Cassia tetraphylla Desv. =Chamaecrista desvauxii var. 

desvauxii 

Cassia tora L. [not in Guianas] 

Cassia uniflora Spreng. [dubious name, not in Guianas] 

Cassia viminea L. =Senna viminea (L.) Irwin & Barneby [not 


Cassia viscosa Kunth =Chamaecrista viscosa 

Crudia obliqua Griseb. =Crudia glaberrima 

Crudia parivoa DC. =Crudia tomentosa 

Dimorphandra congestiflora Sprague & Sandw. 

=Dimorphandra macrostachya subsp. congestiflora 

Dimorphandra hohenkerkii Sprague & Sandw. 

=Dimorphandra polyandra 

Dimorphandra latifolia Tul. =Dimorphandra conjugata 

Elisabetha oxyphylla Harms =Elizabetha coccinea var. 

oxyphylla 

Eperua kourouensis Benoist =Eperua grandiflora 

Eperua stipulata Kleinh. =Eperua jenmanii 

Hymenaea davisii Sandw. =Hymenaea oblongifolia var. 

davisii 

Hymenaea palustris Ducke =Hymenaea oblongifolia var. 

palustris 

Macrolobium acaciifolium (Benth.) Benth. var. vestitum 

Sandw. =Macrolobium acaciifolium 

Macrolobium bifolium (Aubl.) Pers. var. amplexans Amsh. 

=Macrolobium amplexans 

Macrolobium chrysostachyum (Miq.) Benth. =Macrolobium 

angustifolium 

Macrolobium elegans Miq. =Macrolobium bifolium 

Macrolobium hymenaeoides Willd. =Macrolobium bifolium 

Macrolobium simira (Aubl.) J.F. Gmelin =Vouapa simira 

[incertae sedis] 

Macrolobium stamineum G. Mey. =Macrolobium bifolium 

Macrolobium unijugum (Poepp. & Endl.) Cowan 

=Macrolobium limbatum 

Macrolobium unijugum (Poepp. & Endl.) Cowan var. 

fanshawei Cowan =Macrolobium limbatum var. 

fanshawei 

Martiusia parviflora Amsh. =Martiodendron parviflorum 

Melanoxylon speciosum Benoist =Recordoxylon speciosum 

Outea acaciifolia Benth. =Macrolobium acaciifolium 

Outea guianensis Aubl. =Macrolobium guianense 

Outea multijuga DC. =Macrolobium multijugum 

Parivoa grandiflora Aubl. =Eperua grandiflora 

Parivoa tomentosa Aubl. =Crudia tomentosa 

Peltogyne densiflora Spruce ex Benth. =Peltogyne venosa 

subsp. densiflora 

Peltogyne porphyrocardia Griseb. ex Benth. =Peltogyne 

floribunda 

Peltogyne pubescens Benth. =Peltogyne paniculata subsp. 

pubescens 

Poinciana pulcherrima L. =Caesalpinia pulcherrima 

Possira arborescens Aubl. =Swartzia arborescens 

Recordoxylon amazonicum (Ducke) Ducke =Recordoxylon 

speciosum 

Rittera dodecandra Vahl =Swartzia arborescens 

Robinia panacoco Aubl. =Swartzia panacoco 

Senna spruceana Benth. =Cassia spruceana 

Swartzia alata Willd. =Swartzia guianensis 

Swartzia alterna Benth. =Bocoa alterna 

Swartzia apetala Raddi [not in Guianas] 


Swartzia apetala Raddi var. acuminata Amsh. =Swartzia 

amshoffiana 

Swartzia bifida Steud. =Swartzia arborescens 

Swartzia brachystachya DC. =Candolleodendron 

brachystachyum 

Swartzia corrugata Benth. [not in Guianas] 

Swartzia cuspidata Spruce ex Benth. [not in Guianas] 

Swartzia dodecandra (Vahl) Willd. =Swartzia arborescens 

Swartzia grandifolia Bong. ex Benth. var. leiogyne Sandw. 

=Swartzia leiogyne 

Swartzia longifolia DC. =Spirotropis longifolia (Fabaceae) 

Swartzia microstylis Benth. =Swartzia dipetala 

Swartzia minutiflora Kleinh. =Bocoa prouacensis 

Swartzia parviflora DC. =Swartzia arborescens 

Swartzia polyanthera Steud. =Swartzia panacoco var. 

polyanthera 

Swartzia prouacensis (Aubl.) Amsh. =Bocoa prouacensis 

Swartzia similis Benoist =Swartzia panacoco 

Swartzia simplex Spreng. [not in Guianas] 

Swartzia tepuiensis Schery =Swartzia panacoco var. tepuiensis 

Swartzia tomentosa (Willd.) DC. =Swartzia panacoco 

Swartzia tomentosa (Willd.) DC. var. altsoni Sandw. 

=Swartzia panacoco var. altsonii 

Swartzia tomentosa (Willd.) DC. var. cardonae Cowan 

=Swartzia panacoco var. cardonae 

Swartzia tomentosa (Willd.) DC. var. polyanthera (Steud.) 

Sandw. =Swartzia panacoco var. polyanthera 

Swartzia tomentosa (Willd.) DC. var. sagotii Sandw. 

=Swartzia panacoco var. sagotii 

Swartzia tomentosa (Willd.) DC. var. sandwithiana Cowan 

=Swartzia panacoco var. sandwithiana 

Swartzia viridiflora Ducke =Bocoa viridiflora 

Tachigali glauca Tul. =Tachigali cavipes 

Tachigali paniculata Aubl. var. cavipes Benth. =Tachigali 

cavipes 

Tachigali richardiana Tul. ex Lemée [apparently a nomen 

nudum] 

Tachigali sulcata Benoist =Tachigali paniculata var. sulcata 

Tachigali trigona Aubl. =Tachigali paniculata 

Touantea latifolia (Benth.) Taub. =Swartzia latifolia 

Touantea sericea (Vogel) Kuntze =Swartzia sericea 

Touchiroa aromatica Aubl. =Crudia aromatica 

Tounatea aptera (DC.) Taub. =Swartzia aptera 

Tounatea arborescens (Aubl.) Britton =Swartzia arborescens 

Tounatea benthamiana (Miq.) Taub. =Swartzia benthamiana 

Tounatea dodecandra (Vahl) Taub. =Swartzia arborescens 

Tounatea guianensis Aubl. =Swartzia guianensis 

Tounatea hostmanni (Benth.) Taub. =Swartzia hostmannii 

Tounatea schomburgkii (Benth.) Taub. =Swartzia 

schomburgkii 

Vouapa acaciifolia (Benth.) Baill. =Macrolobium acaciifolium 

Vouapa angustifolia Benth. =Macrolobium angustifolium 

Vouapa bifolia Aubl. =Macrolobium bifolium 

Vouapa chrysostachya Miq. =Macrolobium angustifolium


Vouapa guyanensis (Aubl.) Taub. =Macrolobium guianense 

Vouapa jenmani Gleason =Macrolobium jenmanii 

Vouapa multijuga (DC.) Taub. =Macrolobium multijugum 

Vouapa staminea (G. Mey.) DC. =Macrolobium bifolium 

Zollernia kanukuensis Cowan =Zollernia paraënsis 

CAMPANULACEAE 

reviewed by J.L. Luteyn & B. Stein (1992) 

REFERENCES: 

Lanjouw, J. 1934. Campanulaceae. In A. Pulle, 


Lanjouw, J. 1937. Campanulaceae (additions 

and corrections). In A. Pulle, Flora of 

Suriname 4(1): 494-495. 

Centropogon cornutus (L.) Druce [GU SU FG] 

Centropogon roraimanus Wimmer [GU] 

Hippobroma longiflora (L.) G. Don [GU SU] [to be expected 

in FG] 

Lobelia aquatica Cham. [GU SU FG] 

CAMPANULACEAE - SYNONYMS AND 


Centropogon solanifolius Benth. [not in Guianas] 

Centropogon surinamensis (L.) K. Presl =Centropogon 

cornutus 

Isotoma longiflora (L.) K. Presl =Hippobroma longiflora 

REFERENCES: 

CANELLACEAE 

Uittien, H. 1935. Canellaceae. In A. Pulle, Flora 


Wilson, T.K. 1960. The comparative 

morphology of the Canellaceae. I. Synopsis 

of genera and wood anatomy. Trop. Woods 

112: 1-27. 

Cinnamodendron tenuifolium Uittien [SU FG] 

CANNACEAE 


P.J.M. Maas (1996) 

REFERENCES: 

Maas, P.J.M. 1979. Cannaceae. In A.L. Stoffers 

& J.C. Lindeman, Flora of Suriname 5(1): 

412-415. 

Maas, P.J.M. 1985. Cannaceae. In A.R.A. 


A) 1: 69-73. 

Canna glauca L. [GU SU FG] 

Canna indica L. [GU SU FG] [naturalized?] 

CANNACEAE - SYNONYMS AND 


Canna coccinea Mill. =Canna indica 

Canna lambertii Lindl. =Canna indica 

CAPPARACEAE 



REFERENCES: 

Jansen-Jacobs, M.J. 1976. Capparaceae 



517. 

Went, J.C. 1937. Capparidaceae. In A. Pulle, 


Capparis amplissima Lam. [GU? SU] 

Capparis cynophallophora L. var. isthmensis (Eichl.) Iltis ex 

Jansen-Jacobs [SU] 

Capparis frondosa Jacq. [GU SU FG] [=Capparis baducca L.?] 

Capparis guaguaënsis Steyerm. [FG] [to be expected in GU] 

Capparis leprieurii Briq. [GU SU FG] 

Capparis maroniensis Benoist [GU SU FG] 

Capparis pulcherrima Jacq. [FG?] 

Capparis sola Macbr. [SU FG?] 

Capparis verrucosa Jacq. [GU?] 

Capparis yco (Mart.) Mart. & Eichl. var. guianensis Sagot 

[FG] 

Cleome aculeata L. [GU SU FG] 

Cleome frutescens Aubl. [FG] [=Cleome spinosa?] 

Cleome guianensis Aubl. [GU SU FG] 

Cleome gynandra L. [SU FG] [naturalized] 

Cleome latifolia Vahl ex DC. [GU SU FG] 

Cleome micrantha W. Hamilton [GU] [name of uncertain 

application] 

SPERMATOPHYTA: Caesalpiniaceae - Capparaceae 

77

78 

Cleome parviflora Kunth [GU SU FG] 

Cleome rutidosperma DC. [FG] [naturalized] 

Cleome serrata Jacq. [GU SU] 

Cleome speciosa Raf. [GU SU FG] 

Cleome spinosa Jacq. [GU SU FG] 

Cleome stenophylla Klotzsch ex Urb. [GU] 

Cleome viridiflora Schreb. [FG] 

Crateva tapia L. [GU SU FG] 

Morisonia americana L. [GU] 


CAPPARACEAE - SYNONYMS AND 


Capparidastrum baducca (L.) Hutchinson =Capparis baducca 

Capparis baducca L. [Capparis frondosa Jacq. synonymized 

here by some authors] 

Capparis isthmensis Eichl. =Capparis cynophallophora var. 

isthmensis 

Capparis montana (Aubl.) Lemée =Pradosia ptychandra 

(Sapotaceae) 

Capparis perseifolia Iltis ex Jansen-Jacobs [nomen nudum] 

=Capparis leprieurii 

Capparis surinamensis J.C. Went =Capparis maroniensis 

Cleome gigantea L. =Cleome viridiflora 

Cleome polygama L. =Cleome serrata 

Cleome psoraleaefolia DC. =Cleome parviflora 

Cleome surinamensis Miq. =Cleome aculeata 

Crateva acuminata DC. =Crateva tapia 

Crateva benthamii Eichl. =Crateva tapia 

Gynandropsis gynandra (L.) Briq. =Cleome gynandra 

Gynandropsis speciosa (Kunth) DC. =Cleome speciosa 

Physostemon guianense (Aubl.) Malme =Cleome guianensis 

Physostemon lanceolatum Mart. & Zucc. unnamed var. p.p. in 

Fl. Bras. 13(1): 244 (1865) =Cleome stenophylla 

Quadrella cynophallophora (L.) Hutchinson =Capparis 

cynophallophora 

Voyara montana Aubl. =Pradosia ptychandra (Sapotaceae) 

REFERENCES: 

CAPRIFOLIACEAE 

Killip, E.P. & A.C. Smith. 1929. The genus 

Viburnum in northwestern South America. 

Bull. Torrey Bot. Club 56: 265-274. 

Steyermark, J.A. 1953. Caprifoliaceae. In J.A. 

Steyermark & collaborators, Contributions to 

the Flora of Venezuela. Fieldiana Bot. 28(3): 

618-619. 

Viburnum tinoides L.f. var. roraimense (Killip & A.C. Smith) 

Steyerm. [GU] 

CAPRIFOLIACEAE - SYNONYMS AND 


Viburnum glabratum Kunth [not in Guianas] 

Viburnum roraimense Killip & A.C. Smith =Viburnum 

tinoides var. roraimense 

CARICACEAE 

SPERMATOPHYTA: Capparaceae - Caryocaraceae 

REFERENCES: 

Badillo, V.M. 1971. Monografia de la familia 

Caricaceae. Publ. Asoc. Prof., Univ. Central 

de Venezuela. Maracay. 

Lindeman, J.C. 1984. Caricaceae. In A.L. 


5(1): 585-591. 

Carica microcarpa Jacq. subsp. heterophylla (Poepp. & Endl.) 

Badillo [FG] 

Carica papaya L. [GU SU FG] [cultivated, naturalized] 

Jacaratia spinosa (Aubl.) A. DC. [GU SU FG] 

CARICACEAE - SYNONYMS AND 


Carica spinosa Aubl. =Jacaratia spinosa 

Papaya spinosa A. DC. =Jacaratia spinosa 

CARYOCARACEAE 

contributed by G.T. Prance (1992) 

REFERENCES: 

Lanjouw, J. & P.F. van Heerdt. 1941. 

Caryocaraceae. In A. Pulle, Flora of 

Suriname 3(1): 366-372. 

Prance, G.T. 1972. Caryocaraceae. In B. 



Bot. Gard. 23: 1-832. 

Prance, G.T. & M.F. da Silva. 1973. 

Caryocaraceae. Flora Neotropica Monogr. 

12: 1-75. 

Anthodiscus mazarunensis Gilly [GU SU] 

Anthodiscus trifoliatus G. Mey. [GU] 

Caryocar glabrum (Aubl.) Pers. subsp. album Prance & Freitas 

da Silva [GU] 

Caryocar glabrum (Aubl.) Pers. subsp. glabrum [GU SU FG] 

Caryocar microcarpum Ducke [GU SU FG] 

Caryocar montanum Prance [GU] 

Caryocar nuciferum L. [GU SU FG] 

Caryocar pallidum A.C. Smith [FG?] 

Caryocar villosum (Aubl.) Pers. [FG] 

CARYOCARACEAE - SYNONYMS AND 


Caryocar butyrosum (Aubl.) Willd. =Caryocar villosum 

Caryocar tomentosum (Aubl.) Willd. =Caryocar nuciferum 

Pekea butirosa Aubl. =Caryocar villosum 

Pekea tuberculosa Aubl. =Caryocar nuciferum

Saouari glabra Aubl. =Caryocar glabrum 

Saouari villosa Aubl. =Caryocar villosum 


CARYOPHYLLACEAE 


REFERENCES: 

Ooststroom, S.J. van. 1934. Caryophyllaceae. 

In A. Pulle, Flora of Suriname 1(1): 150-153. 

Drymaria cordata (L.) Willd. ex Roem. & Schult. [GU SU FG] 

Polycarpaea corymbosa (L.) Lam. [GU SU] 

CARYOPHYLLACEAE - SYNONYMS AND 


Polycarpaea atherophora Steud. =Polycarpaea corymbosa 

Polycarpaea corymbosa (L.) Lam. var. brasiliensis (Camb.) 

Chod. & Hassler =Polycarpaea corymbosa 

CASUARINACEAE 

[Old World family, one species widely cultivated] 

REFERENCES: 

Berg, C.C. 1992. Casuarinaceae. In A.R.A. 


A) 11: 140. 

Casuarina equisetifolia J.R. & G. Forster [cultivated] 

CECROPIACEAE 

contributed by C.C. Berg (1992); additions and 

corrections by C.C. Berg (1996) 

REFERENCES: 

Berg, C.C. 1992. Cecropiaceae. In A.R.A. 


A) 11: 93-124. 

Berg, C.C., R.W.A.P. Akkermans, & E.C.H. van 

Heusden. 1990. Cecropiaceae: Coussapoa 

and Pourouma. Flora Neotropica Monogr. 51: 

1-194. 

Berg, C.C. & G.P. DeWolf. 1975. Moraceae. In 

J. Lanjouw & A.L. Stoffers, Flora of 

Suriname 5(1): 265-294. 

Cecropia angulata I.W. Bailey [GU] 

Cecropia distachya Huber [FG] 

Cecropia granvilleana C.C. Berg [SU? FG] 

Cecropia kavanayensis Cuatrec. [GU?] 

Cecropia latiloba Miq. [GU SU FG] 

Cecropia obtusa Trécul [GU SU FG] 

Cecropia palmata Willd. [SU FG] 

Cecropia peltata L. [GU SU FG] 

Cecropia sciadophylla Mart. [GU SU FG] 

Cecropia silvae C.C. Berg [SU FG?] 

Coussapoa angustifolia Aubl. [GU SU FG] 

Coussapoa argentea Akkermans & C.C. Berg [GU?] 

Coussapoa asperifolia Trécul subsp. asperifolia [GU SU FG] 

Coussapoa asperifolia Trécul subsp. magnifolia (Trécul) 

Akkermans & C.C. Berg [GU] 

Coussapoa crassivenosa Mildbr. [GU?] 

Coussapoa ferruginea Trécul [SU FG] 

Coussapoa latifolia Aubl. [GU SU FG] 

Coussapoa leprieurii Benoist [FG] 

Coussapoa microcephala Trécul [GU SU FG] 

Coussapoa parvifolia Standl. [GU] 

Coussapoa trinervia Mildbr. [GU] 

Coussapoa viridifolia Cuatrec. [GU] 

Pourouma bicolor Mart. subsp. bicolor [GU SU FG] 

Pourouma bicolor Mart. subsp. digitata (Trécul) C.C. Berg & 

van Heusden [GU SU FG] 

Pourouma bolivarensis C.C. Berg [GU?] 

Pourouma guianensis Aubl. subsp. guianensis [GU SU FG] 

Pourouma melinonii Benoist subsp. melinonii [GU SU FG] 

Pourouma minor Benoist [GU SU FG] 

Pourouma mollis Trécul subsp. mollis [GU SU FG] 

Pourouma saulensis C.C. Berg & F. Kooy [FG] 

Pourouma stipulacea C.C. Berg [GU] 

Pourouma tomentosa Miq. subsp. essequiboensis (Standl.) 

C.C. Berg & Van Heusden [GU] 

Pourouma tomentosa Miq. subsp. maroniensis (Benoist) C.C. 

Berg & Van Heusden [SU FG] 

Pourouma velutina Miq. [SU FG] 

Pourouma villosa Trécul [GU SU FG] 

CECROPIACEAE - SYNONYMS AND 


Cecropia auyantepuiana Cuatrec. =Cecropia kavanayensis 

Cecropia bureauiana Al. Richter =Cecropia palmata 

Cecropia dielsiana Snethlage =Cecropia peltata 

Cecropia juranyiana Al. Richter =Cecropia sciadophylla 

Cecropia paraensis Huber =Cecropia latiloba 

Cecropia sciadophylla Mart. var. decurrens Snethlage 

=Cecropia sciadophylla 

Cecropia steyermarkii Cuatrec. =Cecropia kavanayensis 

Cecropia surinamensis Miq. =Cecropia peltata 

Coussapoa cayennensis Hawkes =Coussapoa asperifolia 

subsp. asperifolia 

Coussapoa cuneata Miq. =Coussapoa microcephala 

Coussapoa fagifolia Klotzsch =Coussapoa microcephala 

Coussapoa hypochlora Standl. =Coussapoa asperifolia subsp. 

magnifolia 

Coussapoa latifolia Aubl. var. obovata (Miq.) Miq. 

=Coussapoa latifolia 

Coussapoa magnifolia Trécul =Coussapoa asperifolia subsp. 

magnifolia 

Coussapoa microcephala Trécul subsp. cornifolia (Standl.) 

Akkermans & C.C. Berg =Coussapoa parvifolia 

SPERMATOPHYTA: Caryocaraceae - Cecropiaceae 

79

80 


Coussapoa obovata Miq. =Coussapoa latifolia 

Coussapoa steyermarkii Cuatrec. =Coussapoa crassivenosa 

Pourouma aspera Trécul =Pourouma bicolor subsp. bicolor 

Pourouma digitata Trécul =Pourouma bicolor subsp. digitata 

Pourouma essequiboensis Standl. =Pourouma tomentosa 

subsp. essequiboensis 

Pourouma laevis Benoist =Pourouma villosa 

Pourouma maroniensis Benoist =Pourouma tomentosa subsp. 

maroniensis 

CELASTRACEAE 

reviewed by D. Kearns (1996) 

REFERENCES: 

Loesener, T. 1942. Celastraceae. In A. Engler & 

K. Prantl, Die Natürlichen Pflanzenfamilien 

20b: 87-197. 

Elaeodendron xylocarpum (Vent.) DC. [possibly to be 


Goupia glabra Aubl. [GU SU FG] 

Maytenus alnifolia Reissek [FG?] 

Maytenus ficiformis Reissek [GU] 

Maytenus floribunda Reissek [FG] 

Maytenus guyanensis Klotzsch ex Reissek [GU] 

Maytenus kanukuensis A.C. Smith [GU] 

Maytenus myrsinoides Reissek [GU SU FG] 

Maytenus oblongata Reissek [GU SU FG] 

Maytenus planifolia A.C. Smith [GU] 

Maytenus pruinosa Reissek [GU SU FG] 

Maytenus tetragona Griseb. [FG] 

CELASTRACEAE - SYNONYMS AND 


Goupia paraensis Huber =Goupia glabra? 

Goupia tomentosa Aubl. =Goupia glabra 

Maytenus reissekii Urb. & Graebn. =Maytenus myrsinoides 

REFERENCES: 

CERATOPHYLLACEAE 

Les, D.H. 1988. The evolution of achene 

morphology in Ceratophyllum, II. Fruit 

variation and systematics of the "spinymargined" 

group. Syst. Bot. 13(1): 73-86. 

Les, D.H. 1989. The evolution of achene 

morphology in Ceratophyllum, IV. Summary 

of proposed relationships and evolutionary 

trends. Syst. Bot. 14(2): 254-262. 

Lowden, R.M. 1978. Studies on the submerged 

genus Ceratophyllum in the Neotropics. 

Aquatic Botany 4: 127-142. 

Wilmot-Dear, 1985. Ceratophyllum revised, a 

study in leaf and fruit variation. Kew Bull. 

40: 243-271. 

Ceratophyllum muricatum Cham. [GU SU] 

SPERMATOPHYTA: Cecropiaceae - Chrysobalanaceae 

CERATOPHYLLACEAE - SYNONYMS AND 


Ceratophyllum demersum L. var. cristatum (Spruce) K. 

Schum. =Ceratophyllum muricatum 

Ceratophyllum lleranae Fassett =Ceratophyllum muricatum 

CHENOPODIACEAE 


REFERENCES: 

Lanjouw, J. 1957. Chenopodiaceae. In A. Pulle 

& J. Lanjouw, Flora of Suriname 1(2): 291- 

292. 

Chenopodium ambrosioides L. [GU SU FG] [naturalized] 

CHLORANTHACEAE 

contributed by C.A. Todzia (1992); additions and 

corrections by C.A. Todzia (1996) 

REFERENCES: 

Todzia, C.A. 1988. Chloranthaceae: 

Hedyosmum. Flora Neotropica Monogr. 48: 

1-138. 

Todzia, C.A. 1993. New species of Hedyosmum 

from northern South America. Novon 3: 81- 

85. 

Hedyosmum tepuiense Todzia [GU] 

CHLORANTHACEAE - SYNONYMS AND 


Hedyosmum brasiliense Mart. [not in Guianas] 

Hedyosmum racemosum Auctt., non (Ruiz & Pav.) G. Don 

=Hedyosmum tepuiense 

CHRYSOBALANACEAE 


REFERENCES: 

Kleinhoonte, A. 1939. Rosaceae. In A. Pulle, 

Flora of Suriname 2(1): 426-456.


Prance, G.T. 1972. Chrysobalanaceae. Flora 


Prance, G.T. 1981. Notes on Couepia and 

Hirtella. Brittonia 33: 347-356. 

Prance, G.T. 1982. Chrysobalanaceae. Flora de 

Venezuela 14(2): 325-487. 

Prance, G.T. 1986a. Chrysobalanaceae. In 


(seriesA) 2: 1-146. 

Prance, G.T. 1986b. Studies on the flora of the 

Guianas. 19. New taxa of Chrysobalanaceae 

for the Flora of the Guianas. Proc. Kon. Ned. 

Akad. Wetensch., ser. C. 89: 111-116. 

Prance, G.T. 1989. Chrysobalanaceae 

(supplement). Flora Neotropica Monogr. 9S: 

1-267. 

Prance, G.T. 1992. New species and new 

records of neotropical Chrysobalanaceae. 

Kew Bull. 47: 633-646. 

Prance, G.T. & A.R.A. Görts-van Rijn. 1976. 

Chrysobalanaceae (additions and corrections). 

In J. Lanjouw & A.L. Stoffers, Flora of 

Suriname 2(2): 524-555. 

Acioa guianensis Aubl. [FG] 

Acioa somnolens Maguire [FG] 

Chrysobalanus icaco L. [GU SU FG] 

Couepia bracteosa Benth. [GU SU FG] 

Couepia canomensis (Mart.) Benth. ex J.D. Hook. [GU FG] 

Couepia caryophylloides Benoist [SU FG] 

Couepia cognata (Steud.) Fritsch var. cognata [GU SU] 

Couepia cognata (Steud.) Fritsch var. major Prance [GU] 

Couepia cognata (Steud.) Fritsch var. membranacea Prance 

[GU] 

Couepia comosa Benth. [GU] 

Couepia elata Ducke [GU] 

Couepia exflexa Fanshawe & Maguire [GU] 

Couepia foveolata Prance [GU] 

Couepia guianensis Aubl. subsp. divaricata (Hub.) Prance [GU 

SU FG] 

Couepia guianensis Aubl. subsp. glandulosa (Miq.) Prance 

[GU SU FG] 

Couepia guianensis Aubl. subsp. guianensis [GU SU FG] 

Couepia habrantha Standl. [GU FG] 

Couepia martinii Prance [FG] 

Couepia multiflora Benth. [GU] 

Couepia obovata Ducke [SU FG] 

Couepia paraensis (Mart. & Zucc.) Benth. subsp. glaucescens 

(Spr. ex Hook.f.) Prance [GU] 

Couepia parillo A.P. DC. [GU SU FG] 

Couepia sandwithii Prance [GU] 

Exellodendron barbatum (Ducke) Prance [GU SU FG] 

Exellodendron coriaceum (Benth.) Prance [GU] 

Hirtella angustissima Sandw. [GU FG] 

Hirtella araguariensis Prance [GU SU FG] 

Hirtella bicornis Mart. & Zucc. var. bicornis [SU FG] 

Hirtella bicornis Mart. & Zucc. var. pubescens Ducke [GU SU 

FG] 

Hirtella bullata Benth. [GU] 

Hirtella caduca Fanshawe & Maguire [GU] 

Hirtella ciliata Mart. & Zucc. [GU FG] 

Hirtella couepiiflora Prance [FG] 

Hirtella davisii Sandw. [GU FG] 

Hirtella deflexa Maguire [to be expected in GU] 

Hirtella duckei Huber [GU] 

Hirtella eriandra Benth. [to be expected in FG] 

Hirtella glandistipula Ducke [FG] 

Hirtella glandulosa Spreng. [GU SU FG] 

Hirtella guyanensis (Fritsch) Sandw. [GU] 

Hirtella hispidula Miq. [GU SU FG] 

Hirtella macrosepala Sandw. [GU SU FG] 

Hirtella margae Prance [GU SU] 

Hirtella mucronata Prance [GU SU] 

Hirtella obidensis Ducke [GU SU FG] 

Hirtella paniculata Swartz [GU SU FG] 

Hirtella physophora Mart. & Zucc. [GU SU FG] 

Hirtella piresii Prance [GU SU] 

Hirtella punctillata Ducke [GU SU FG] 

Hirtella racemosa Lam. var. hexandra (Willd. ex Roem.& 

Schult.) Prance [GU SU FG] 

Hirtella racemosa Lam. var. hispida Prance [GU FG] 

Hirtella racemosa Lam. var. racemosa [GU SU FG] 

Hirtella scabra Benth. [GU] 

Hirtella silicea Griseb. [GU SU FG] 

Hirtella suffulta Prance [FG] 

Hirtella tentaculata Poepp. & Endl. [GU] 

Hirtella tenuifolia Prance [FG] 

Hirtella triandra Swartz subsp. triandra [GU SU] 

Licania affinis Fritsch [GU SU FG] 

Licania alba (Bernoulli) Cuatrec. [GU SU FG] 

Licania albiflora Fanshawe & Maguire [GU SU FG] 

Licania amapaensis Prance [FG] 

Licania apetala (E. Mey.) Fritsch var. aperta (Benth.) Prance 

[GU SU FG] 

Licania apetala (E. Mey.) Fritsch var. apetala [GU SU FG] 

Licania arachnoidea Fanshawe & Maguire [GU] 

Licania boyanii Tutin [GU] 

Licania buxifolia Sandw. [GU] 

Licania canescens Benoist [GU SU FG] 

Licania caudata Prance [GU FG] 

Licania compacta Fritsch [GU] [Mt. Roraima] 

Licania coriacea Benth. [GU FG] 

Licania couepifolia Prance [GU SU] 

Licania cuprea Sandw. [GU] 

Licania cyathodes Benoist [FG] 

Licania davillifolia Benoist [SU FG] 

Licania densiflora Kleinh. [GU SU FG] 

Licania discolor Pilger [GU SU FG] 

Licania divaricata Benth. [GU SU] 

Licania elliptica Standl. [SU FG] 

Licania fanshawei Prance [GU SU FG] 

Licania foveolata Prance [GU] 

Licania glabriflora Prance [SU FG] 

Licania granvillei Prance [GU FG] 

Licania guianensis (Aubl.) Griseb. [GU SU FG] 

Licania heteromorpha Benth. var. glabra (Mart. ex J.D. Hook.) 

Prance [GU FG] 

Licania heteromorpha Benth. var. heteromorpha [GU SU FG] 

Licania heteromorpha Benth. var. perplexans Sandw. [GU] 

Licania hypoleuca Benth. var. hypoleuca [GU SU FG] 

Licania imbaimadaiensis Prance [GU] 

Licania incana Aubl. [GU SU FG] 

Licania intrapetiolaris Spruce ex J.D. Hook. [GU] 

Licania irwinii Prance [GU SU FG] 

Licania jimenezii Prance [GU SU] 

Licania kunthiana J.D. Hook. [GU SU FG] 

Licania laevigata Prance [GU SU FG] 

Licania lasseri Maguire [GU] 

Licania lata Macbr. [FG] 

Licania latifolia Benth. ex J.D. Hook. [GU FG] 

SPERMATOPHYTA: Chrysobalanaceae 

81

82 


Licania latistipula Prance [FG] 

Licania laxiflora Fritsch [GU SU FG] 

Licania leptostachya Benth. [GU SU FG] 

Licania licaniiflora (Sagot) Fritsch [GU SU FG] 

Licania longistyla (J.D. Hook.) Fritsch [GU SU FG] 

Licania macrophylla Benth. [SU FG] 

Licania majuscula Sagot [GU SU FG] 

Licania maxima Prance [SU? FG] 

Licania membranacea Sagot ex Laness. [GU FG] 

Licania micrantha Miq. [GU SU FG] 

Licania microphylla Fanshawe & Maguire [GU] 

Licania minutiflora (Sagot) Fritsch [GU SU FG] 

Licania obtusifolia Fritsch [FG] [dubious species] 

Licania octandra (Hoffmanns. ex Roem. & Schult.) Kuntze 

subsp. octandra [GU SU FG] 

Licania octandra (Hoffmanns. ex Roem. & Schult.) Kuntze 

subsp. pallida (J.D. Hook.) Prance [GU] 

Licania ovalifolia Kleinh. [SU FG] 

Licania parviflora Benth. [GU SU? FG] 

Licania parvifolia Hub. [GU] 

Licania parvifructa Fanshawe & Maguire [GU] 

Licania persaudii Fanshawe & Maguire [GU FG] 

Licania piresii Prance [GU FG] 

Licania polita Spruce ex J.D. Hook. [GU SU] 

Licania pruinosa Benoist [FG] 

Licania reticulata Prance [GU FG] 

Licania robusta Sagot [SU FG] 

Licania roraimensis Standl. [GU] [Mt. Roraima] 

Licania rufescens Klotzsch ex Fritsch [GU SU] 

Licania sandwithii Prance [GU] 

Licania silvae Prance [GU FG] 

Licania sprucei (J.D. Hook.) Fritsch [GU FG] 

Licania stricta Kleinh. [GU? SU] [dubious species] 

Parinari campestris Aubl. [GU SU FG] 

Parinari excelsa Sabine [GU SU FG] 

Parinari maguirei Prance [GU FG?] 

Parinari montana Aubl. emend. Ducke [SU FG] 

Parinari parvifolia Sandw. [GU FG] 

Parinari rodolphii Hub. [GU SU FG] 

CHRYSOBALANACEAE - SYNONYMS AND 


Chrysobalanus icaco L. var. pellocarpus (G. Mey.) Hook.f. 

=Chrysobalanus icaco 

Chrysobalanus incanus (Aubl.) Gómez de la Maza =Licania 

incana 

Chrysobalanus montanus (Aubl.) Gómez de la Maza =Parinari 

montana 

Chrysobalanus pellocarpus G. Mey. =Chrysobalanus icaco 

Chrysobalanus savannarum Britton =Chrysobalanus icaco 

Couepia glandulosa Miq. =Couepia guianensis subsp. 

glandulosa 

Couepia myrtifolia Benth. ex J.D. Hook. =Couepia guianensis 

subsp. glandulosa 

Couepia pauciflora Huber =Couepia parillo 

Couepia steudeliana Walp. =Couepia cognata 

Couepia surinamensis Kleinh. =Couepia guianensis subsp. 

guianensis 

Couepia versicolor Benoist =Couepia guianensis subsp. 

guianensis 

Couepia villosa Fanshawe & Maguire =Couepia parillo 

Ferolia coriacea (Benth.) Kuntze =Exellodendron coriaceum 

Ferolia montana (Aubl.) Kuntze =Parinari montana 

SPERMATOPHYTA: Chrysobalanaceae 

Hirtella americana Aubl., non. L. =Hirtella racemosa var. 

racemosa 

Hirtella apetala E. Mey. =Licania apetala 

Hirtella bracteosa Steud. =Hirtella paniculata 

Hirtella caudata Kleinh. =Hirtella bicornis var. pubescens 

Hirtella cauliflora Huber =Hirtella physophora 

Hirtella cognata Steud. =Couepia cognata 

Hirtella coppenamensis Kleinh. =Hirtella obidensis 

Hirtella cotticaensis Kleinh. =Hirtella physophora 

Hirtella lanceolata Kleinh. =Hirtella obidensis 

Hirtella manigera Kleinh. =Hirtella silicea 

Hirtella melinonii Benoist =Hirtella silicea 

Hirtella praealta Sagot =Hirtella bicornis var. bicornis 

Hirtella rubra Benth. =Hirtella ciliata 

Hirtella strigulosa Steud. =Hirtella racemosa var. racemosa 

Hirtella subsetosa Fanshawe & Maguire =Hirtella hispidula 

Hirtella velutina Pilger =Hirtella glandulosa 

Hirtella violacea Steud. =Hirtella racemosa var. racemosa 

Hirtella wachenheimii Benoist =Hirtella glandulosa 

Licania aperta Benth. =Licania apetala var. aperta 

Licania apetala (E. Mey.) Fritsch var. pendula (Benth.) Fritsch 

=Licania apetala var. apetala 

Licania arachnites Standl. =Licania parvifolia 

Licania aubletiana (Blume) Lemée =Acioa guianensis 

Licania axilliflora (Sagot) Hochr. =Licania leptostachya 

Licania crassifolia Benth. =Licania incana 

Licania cymosa Fritsch [not in Guianas] 

Licania exiguifolia Standl. =Couepia cognata var. cognata 

Licania floribunda Benth. =Licania apetala var. apetala 

Licania galibica Benoist =Licania membranacea 

Licania gracilis Kleinh. =Licania laxiflora 

Licania grisea Kleinh. =Licania hypoleuca 

Licania heteromorpha Benth. var. divaricata (Benth.) Fritsch 

=Licania divaricata 

Licania hostmannii Fritsch =Licania majuscula 

Licania incana Aubl. var. axilliflora Sagot =Licania 

leptostachya 

Licania kanukuensis Standl. =Licania densiflora 

Licania laxa Fanshawe & Maguire =Licania polita 

Licania leptostachya Benth. var. axilliflora (Sagot) Fritsch 

=Licania leptostachya 

Licania leptostachya Benth. var. crassifolia (Benth.) Benoist 

=Licania incana 

Licania longifolia Benoist =Licania alba 

Licania mollis Benth. [not in Guianas] 

Licania pachystachya Kleinh. =Licania robusta 

Licania paniculata Fanshawe & Maguire =Licania elliptica 

Licania parviflora (Blume) Lemée, nom. illeg. =Licania 

obtusifolia 

Licania pendula Benth. =Licania apetala var. apetala 

Licania poeppigii Fritsch =Licania polita 

Licania pubiflora Benth. =Licania apetala var. aperta 

Licania sclerophylla (Mart. ex J.D. Hook.) Fritsch [not in 


Licania stahelii Kleinh. =Licania ovalifolia 

Licania stipata Fanshawe & Maguire [nomen nudum] 

Licania takutuensis Standl. =Licania octandra subsp. octandra 

Licania turiuva Cham. & Schlechtend., nom. illeg. =Licania 

octandra subsp. octandra 

Licania utilis (J.D. Hook.) Fritsch =Licania octandra subsp. 

octandra 

Licania wilson-brownei Maguire =Licania licaniiflora


Moquilea aubletiana Blume, nom. illeg. =Acioa guianensis 

Moquilea comosa (Benth.) Walp. =Couepia comosa 

Moquilea floribunda (Benth.) Hook.f. =Licania apetala var. 

apetala 

Moquilea glandulosa (Miq.) Walp. =Couepia guianensis 

subsp. glandulosa 

Moquilea guianensis Aubl. =Licania guianensis 

Moquilea licaniiflora Sagot =Licania licaniiflora 

Moquilea minutiflora Sagot =Licania minutiflora 

Moquilea multiflora (Benth.) Walp. =Couepia multiflora 

Moquilea pendula (Benth.) Hook.f. =Licania apetala var. 

apetala 

Moquilea turiuva (Cham. & Schlechtend.) Hook.f. =Licania 

octandra 

Parinari guyanense (Aubl.) Lemée =Brosimum rubescens 

(Moraceae) 

Parinari hostmannii Fritsch =Couepia cognata 

Parinari lucidissima Standl. =Parinari rodolphii 

Parinarium brachystachum Benth. =Parinari excelsa 

Parinarium coriaceum Benth. =Exellodendron coriaceum 

Parinarium guyanense Fritsch =Hirtella guyanensis 

Petrocarya campestris (Aubl.) Willd. =Parinari campestris 

Petrocarya montana (Aubl.) Willd. =Parinari montana 

Tachibota guianensis Aubl. =Hirtella racemosa var. racemosa 

REFERENCES: 

CLETHRACEAE 

Sleumer, H. 1967. Monographia Clethracearum. 

Bot. Jahrb. Syst. 87: 36-175. 

Sleumer, H. 1971. Clethraceae. In T. Lasser, 

Flora de Venezuela 3(1): 171-183. 

Clethra guyanensis Klotzsch ex Meissn. [Mt. Roraima] 

CLUSIACEAE 

Hypericum, Vismia contributed by N.K.B. 

Robson (1992); reviewed by N.K.B. Robson 

& J. Pipoly (1996) 

REFERENCES: 

Berg, M.E. van der. 1979. Revisao das espécies 

brasileiras do gênero Rheedia L. Acta Amaz. 

9(1): 43-74. 

Eyma, P.J. 1934. Guttiferae. In A. Pulle, Flora 


Ewan, J. 1962. Synopsis of the South American 

species of Vismia. Contrib. U.S. Nat. Herb. 

35(5): 293-377. 

Kubitzki, K. 1978. Caraipa and Mahurea 

(Bonnetiaceae). In B. Maguire & 


Highland. Mem. New York Bot. Gard. 29: 

82-138. 

Maguire, B. 1958. Guttiferae. In B. Maguire, J. 

Wurdack & collaborators, Botany of the 

Guayana Highland Part III. Mem. New York 

Bot. Gard. 10(1): 49-61. 

Maguire, B. 1961. Guttiferae. In B. Maguire, 

J.J. Wurdack & collaborators, Botany of the 

Guayana Highland part IV. Mem. New York 

Bot. Gard. 10(4): 21-32. 

Maguire, B. 1964. Guttiferae subfam. 

Moronoboideae. In B. Maguire, J.J. 

Wurdack, & collaborators, Botany of the 

Guayana Highland Part V. Mem. New York 

Bot. Gard. 10(5): 123-135. 

Maguire, B. 1972. Clusiaceae - Quapoya and 

Renggeria. In B. Maguire & collaborators, 

Botany of the Guayana Highland Part IX. 


Robson, N.K.B. 1987. Studies in the genus 

Hypericum L. (Guttiferae). 7. Section 29. 

Brathys (part 1). Bull. Br. Mus. Nat. Hist. 

(Bot.) 16: 1-106. 

Robson, N.K.B. 1990. Studies in the genus 

Hypericum L. (Guttiferae). 8. Sections 29. 

Brathys (part 2) and 30. Trigynobrathys. 

Bull. Br. Mus. Nat. Hist. (Bot.) 20: 1-151. 


of tropical America: XXVI. The genus 

Tovomita in British Guiana. Kew Bull. 1936: 

210-221. 

Calophyllum brasiliense Camb. [GU SU FG] 

Calophyllum longifolium Willd. [GU SU] 

Calophyllum lucidum Benth. [GU FG?] 

Caraipa ampla Ducke [FG] 

Caraipa densifolia Mart. subsp. densifolia [GU SU FG] 

Caraipa llanorum Cuatrec. [GU] 

Caraipa parvifolia Aubl. [FG] 

Caraipa psilocarpa Kubitzki [GU?] 

Caraipa punctulata Ducke [GU SU FG] 

Caraipa racemosa Cambess. [GU FG] 

Caraipa richardiana Cambess. [GU SU FG] 

Caraipa savannarum Kubitzki [GU] 

Caraipa tereticaulis Tul. [GU] 

Clusia aishaltonensis Pipoly, sp. nov. ined. [GU] 

Clusia amazonica Planch. & Triana [GU] 

Clusia brachystyla Maguire var. brachystyla [GU] 

Clusia brongniartiana Planch. & Triana [FG] [dubious species] 

Clusia capituliflora Pipoly, sp. nov. ined. [GU] 

Clusia cardonae Maguire [GU] 

Clusia columnaris Engl. [GU SU] 

Clusia crassifolia Planch. & Triana [GU] 

Clusia cuneata Benth. [GU FG] 

Clusia duidae Gleason [GU] 

Clusia flaviflora Engl. [SU] 

Clusia fockeana Miq. [GU SU FG] 

Clusia gaudichaudii Choisy [SU] 

Clusia grandiflora Splitg. [GU SU FG] 

Clusia guayanae Pipoly, sp. nov. ined. [GU] 

Clusia insignis Mart. [GU SU FG] [=Clusia grandiflora?] 

Clusia kanukuana Maguire [GU FG?] 

Clusia leprantha Mart. [GU SU FG] [=Clusia purpurea?] 

Clusia lunanthera Maguire [SU] 

Clusia macropoda Engl. [GU SU] 

Clusia maguireana Pipoly, sp. nov. ined. [GU] 

Clusia martinii Sagot [GU] [=Clusia palmicida?] 

Clusia melchiorii Gleason [GU SU FG] 

SPERMATOPHYTA: Chrysobalanaceae - Clusiaceae 

83

84 

SPERMATOPHYTA: Clusiaceae 


Clusia microstemon Planch. & Triana [GU SU] 

Clusia minor L. [GU SU FG] 

Clusia mutica Maguire [GU] 

Clusia myriandra (Benth.) Planch. & Triana [GU] 

Clusia nemorosa G. Mey. [GU SU FG] 

Clusia palmicida L.C. Rich. [GU SU FG] 

Clusia panapanari (Aubl.) Choisy [GU SU FG] 

Clusia parvifolia Maguire [GU] 

Clusia phelpsiana Maguire [to be expected in GU] 

Clusia picta Pipoly, sp. nov. ined. [GU] 

Clusia platystigma Eyma [GU? SU FG] 

Clusia purpurea (Splitg.) Engl. [GU SU FG] 

Clusia pusilla Steyerm. subsp. pakaraimae Maguire [GU] 

Clusia pusilla Steyerm. subsp. pusilla [GU] 

Clusia renggerioides Triana & Planch. [GU] 

Clusia robusta Eyma [SU] 

Clusia rosea Jacq. [GU SU FG] [=Clusia major L.?] 

Clusia savannarum Maguire [GU] 

Clusia schomburgkiana (Planch. & Triana) Benth. ex Engl. 

[GU] 

Clusia schomburgkii Vesque [GU] 

Clusia scrobiculata Benoist [GU SU FG] 

Clusia sessilis Klotzsch ex Engl. [GU SU] 

Clusia stylosa Maguire [SU] 

Clusia tabulamontana Maguire [GU SU] 

Clusiella axillaris (Engl.) Cuatrec. [GU] 

Clusiella elegans Planch. & Triana [GU] 

Dystovomita sp. [GU] 

Haploclathra leiantha (Benth.) Benth. [GU] 

Havetiopsis flavida (Benth.) Planch. & Triana [GU FG] 

Hypericum marahuacanum N. Robson subsp. chimantaicum N. 

Robson [to be expected in GU] 

Hypericum roraimense Gleason [GU] [Mt. Roraima] 

Hypericum thesiifolium Kunth [to be expected in GU] 

Lorostemon bombaciflorum Ducke [GU] 

Mahurea exstipulata Benth. subsp. exstipulata [GU] 

Mahurea palustris Aubl. [GU? FG] 

Marila saramaccana Pulle [SU] 

Moronobea coccinea Aubl. [GU SU FG] 

Moronobea intermedia Engl. [GU] 

Moronobea jenmanii Engl. var. fanshawei Maguire [GU] 

Moronobea jenmanii Engl. var. jenmanii [GU FG?] 

Oedematopus obovatus (Spruce) Planch. & Triana [GU SU 

FG] 

Oedematopus octandrus (Poepp. & Endl.) Planch. & Triana 

[GU FG] 

Oedematopus quadratus Maguire [SU] 

Platonia insignis Mart. [GU SU FG] 

Quapoya bracteolata Sandw. [GU] 

Quapoya peruviana (Poepp. & Endl.) Kuntze [GU] 

Quapoya scandens Aubl. [FG] 

Quapoya sipapoana Maguire [GU] 

Renggeria comans Meisn. [FG] 

Rheedia acuminata (Ruiz & Pav.) Planch. & Triana [GU SU 

FG] [=Garcinia acuminata (Ruiz & Pav.) Hammel] 

Rheedia benthamiana Planch. & Triana [GU SU FG] 

[=Garcinia benthamiana (Planch. & Triana) Pipoly, comb. 

nov. ined.] 

Rheedia brasiliensis (Mart.) Planch. & Triana [FG] [=Garcinia 

brasiliensis Mart.] 

Rheedia lateriflora L. [GU] [=Garcinia humilis (Vahl) C. 

Adams] 

Rheedia macrophylla (Mart.) Planch. & Triana [GU SU FG] 

[=Garcinia macrophylla Mart.] 

Rheedia madruno (Kunth) Planch. & Triana [SU] [=Garcinia 

madruno (Kunth) Hammel] 

Rheedia martinii Maguire [SU] [=Garcinia sp.] 

Rheedia virens Planch. & Triana [GU] [=Garcinia sp.] 

Symphonia globulifera L.f. [GU SU FG] 

Thysanostemon fanshawei Maguire [GU] 

Thysanostemon pakaraimae Maguire [GU] 

Tovomita albiflora A.C. Smith [GU] 

Tovomita atropurpurea Steyerm. [GU] 

Tovomita brasiliensis (Mart.) Walpers [GU SU FG] 

Tovomita brevistaminea Engl. [GU SU FG] 

Tovomita calodictyos Sandw. [GU SU] 

Tovomita caloneura A.C. Smith [GU] 

Tovomita carinata Eyma [SU FG] 

Tovomita cephalostigma Vesque [GU SU FG] 

Tovomita choisyana Planch. & Triana [GU SU FG] 

Tovomita clusiifolia G. Don [FG] 

Tovomita divaricata Maguire [GU] 

Tovomita eggersii Vesque [GU] 

Tovomita fanshawei Maguire [GU] 

Tovomita grata Sandw. [GU SU FG] 

Tovomita guianensis Aubl. [GU FG] 

Tovomita longifolia (L.C. Rich.) Hochr. [GU SU FG] 

Tovomita macrophylla Walp. [GU] 

Tovomita melinonis Vesque [FG] 

Tovomita obovata Engl. [GU] 

Tovomita obscura Sandw. [GU] 

Tovomita pyrifolia Planch. & Triana [SU?] 

Tovomita schomburgkii Planch. & Triana [GU SU] 

Tovomita secunda Poepp. ex Planch. & Triana [SU FG] 

Tovomita tenuiflora Benth. [GU] 

Tovomita umbellata Benth. [GU FG] 

Tovomita uniflora G. Don [FG] 

Tovomita weddelliana Planch. & Triana [GU] 

Tovomitidium speciosa (Ducke) Ducke [to be expected in FG] 

Tovomitopsis membranacea (Planch. & Triana) D'Arcy [GU 

FG] [=Chrysochlamys membranacea Planch. & Triana] 

Tovomitopsis nicaraguensis (Oerst.) Planch. & Triana [GU] 

Tovomitopsis standleyana (L.O. Williams) D'Arcy [GU] 

Vismia baccifera (L.) Triana & Planch. subsp. dealbata 

(Kunth) Ewan [SU? FG?] 

Vismia cavalcantei v.d. Berg [SU] 

Vismia cayennensis (Jacq.) Pers. [GU SU FG] 

Vismia confertiflora Spruce ex Reichardt [GU? SU FG] 

Vismia glaziovii Ruhl. [GU SU FG] 

Vismia guianensis (Aubl.) Choisy var. acuminata (Lam.) v.d. 

Berg [GU SU FG] 

Vismia guianensis (Aubl.) Choisy var. guianensis [GU SU 

FG] 

Vismia japurensis Reichardt [GU SU] 

Vismia latifolia (Aubl.) Choisy [GU? SU FG] 

Vismia lauriformis (Lam.) Choisy [FG?] [to be expected in 

GU] 

Vismia laxiflora Reichardt [GU] 

Vismia macrophylla Kunth [GU SU FG] 

Vismia ramuliflora Miq. [GU SU FG] 

Vismia sandwithii Ewan [GU FG] 

Vismia sessilifolia (Aubl.) Choisy [GU SU FG] 

CLUSIACEAE - SYNONYMS AND 

MISIDENTIFICATIONS:


Caopia acuminata (Lam.) Kuntze =Vismia guianensis var. 

acuminata 

Caopia cayennensis (Jacq.) Kuntze =Vismia cayennensis 

Caopia guianensis (Aubl.) Lyons =Vismia guianensis 

Caopia latifolia (Aubl.) Kuntze =Vismia latifolia 

Caopia laxiflora (Reich.) Kuntze =Vismia laxiflora 

Caopia sessilifolia (Aubl.) Kuntze =Vismia sessilifolia 

Caraipa angustifolia Aubl. =Couepia sp. (Chrysobalanaceae) 

Caraipa glabrata Mart. =Caraipa grandifolia [not in Guianas] 

Caraipa lacerdaei Barb. Rodr. =Caraipa grandifolia [not in 


Caraipa latifolia Aubl. =Licania membranacea 

(Chrysobalanaceae) 

Caraipa laxiflora Benth. =Caraipa densifolia 

Caraipa longifolia Aubl. =Licania alba (Chrysobalanaceae) 

Caraipa paraensis J. Huber =Caraipa grandifolia [not in 


Caraipa psidiifolia Ducke =Caraipa richardiana 

Caraipa simplicior Sandw. =Caraipa tereticaulis? 

Caraipa variabilis Camb. =Caraipa densifolia? 

Chrysochlamys membranacea Planch. & Triana 

=Tovomitopsis membranacea 

Chrysochlamys weberbaueri Engl. =Tovomitopsis standleyana 

Clusia cerroana Steyerm. =Clusia sessilis 

Clusia colorans Klotzsch ex Engl. =Clusia panapanari 

Clusia cucullata Klotzsch [nomen nudum] 

Clusia jenmanii Engl. =Clusia myriandra 

Clusia parvicapsula Vesque =Clusia minor 

Clusia planchoniana Engl. var. oblongifolia Vesque =Clusia 

schomburgkiana 

Clusia roraimana Steyerm. =Clusia schomburgkii 

Clusia stahelii Maguire =Clusia fockeana 

Clusia venosa Jacq. =Clusia minor 

Garcinia acuminata (Ruiz & Pav.) Hammel =Rheedia 

acuminata 

Garcinia brasiliensis Mart. =Rheedia brasiliensis 

Garcinia macrophylla Mart. =Rheedia macrophylla 

Garcinia madruno (Kunth) Hammel =Rheedia madruno 

Havetiopsis flexilis (Benth.) Planch. & Triana =Havetiopsis 

flavida 

Hypericum acuminatum Lam. =Vismia guianensis var. 

acuminata 

Hypericum cayennense Jacq. =Vismia cayennensis 

Hypericum guianense Aubl. =Vismia guianensis 

Hypericum latifolium Aubl. =Vismia latifolia 

Hypericum reticulatum Poir. =Vismia macrophylla 

Hypericum rufescens Lam. =Vismia sessilifolia 

Hypericum sessilifolium Aubl. =Vismia sessilifolia 

Oedematopus duidae Gleason [recorded from Guianas in 

error] 

Quapoya acuminata (Planch. & Triana) Kuntze =Quapoya 

bracteolata 

Quapoya panapanari Aubl. =Clusia panapanari 

Renggeria montana Klotzsch [nomen nudum] =Quapoya 

bracteolata 

Rengifa acuminata Planch. & Triana =Quapoya bracteolata 

Rengifa scandens (Aubl.) Planch. & Triana =Quapoya 

scandens 

Rheedia kappleri Eyma =Rheedia acuminata 

Rheedia sagotiana Engl. =Rheedia benthamiana 

Vismia acuminata (Lam.) Pers. =Vismia guianensis var. 

acuminata 

Vismia amazonica Ewan =Vismia glaziovii 

Vismia angusta Miq. =Vismia macrophylla 

Vismia baccifera (L.) Triana & Planch. subsp. ferruginea 

(Kunth) Ewan =Vismia guianensis 

Vismia dealbata Kunth =Vismia baccifera subsp. dealbata 

Vismia falcata Rusby =Vismia laxiflora 

Vismia ferruginea Kunth =Vismia guianensis 

Vismia lindeniana Decne. [not in Guianas] 

Vismia pentagyna (Spreng.) Ewan [not in Guianas] 

Vismia reticulata (Poir.) Choisy =Vismia macrophylla 

Vismia rufescens (Lam.) Pers. =Vismia sessilifolia 

Vismia schomburgkiana Klotzsch ex Schomb. [nomen nudum] 

COMBRETACEAE 

contributed by C.A. Stace (1992); additions and 

corrections by C.A. Stace (1996) 

REFERENCES: 

Alwan Al-Mayah, A.-R. A. & C.A. Stace. 1985. 

Resurrection of two Eighteenth Century 

names in American Combretaceae. Nord. J. 

Bot. 5(5): 447-449. 

Exell, A.W. 1935. Combretaceae. In A. Pulle, 


Exell, A.W. 1953. The Combretum species of 

the New World. J. Linn. Soc. Lond. (Bot.) 

55: 103-141. 

Exell, A.W. & C.A. Stace. 1963. A revision of 

the genera Buchenavia and Ramatuella. Bull. 

Brit. Mus. (Nat. Hist.) 3: 1-46. 

Exell, A.W. & C.A. Stace. 1966. Revision of the 

Combretaceae. Bol. Soc. Brot. (ser. 2) 40: 5- 

26. 

Buchenavia amazonia Alwan & Stace [FG?] 

Buchenavia fanshawei Exell & Maguire [GU] 

Buchenavia grandis Ducke [GU SU FG] 

Buchenavia guianensis (Aubl.) Alwan & Stace [FG] 

Buchenavia macrophylla Eichl. [SU? FG] 

Buchenavia megalophylla Van Heurck & Muell. [GU] 

Buchenavia nitidissima (L.C. Rich.) Alwan & Stace [FG] 

Buchenavia parvifolia Ducke [GU FG] 

Buchenavia pulcherrima Exell & Stace [GU] 

Buchenavia tetraphylla (Aubl.) R. Howard [SU FG] 

Buchenavia viridiflora Ducke [FG] 

Combretum cacoucia (Baill.) Exell ex Sandw. [GU SU FG] 

Combretum fruticosum (Loefl.) Stuntz [GU] 

Combretum glabrum DC. [GU FG] 

Combretum laurifolium Mart. [GU] 

Combretum laxum Jacq. [GU SU FG] 

Combretum llewelynii Macbr. [GU?] 

Combretum pyramidatum Desv. ex Hamilton [GU SU FG] 

Combretum rohrii Exell [FG] 

Combretum rotundifolium L.C. Rich. [GU SU FG] 

Conocarpus erecta L. [GU SU FG] 

Laguncularia racemosa (L.) C.F. Gaertn. [GU SU FG] 

Terminalia amazonia (J.F. Gmelin) Exell [GU SU FG] 

Terminalia catappa L. [SU FG] [cultivated, naturalized] 

Terminalia dichotoma G. Mey. [GU SU FG] 

Terminalia guyanensis Eichl. [GU SU FG] 

SPERMATOPHYTA: Clusiaceae - Combretaceae 

85

86 


Terminalia lucida Hoffmanns. ex Mart. [GU SU FG] 

Terminalia oblonga (Ruiz & Pav.) Steud. [FG] 

Terminalia quintalata Maguire [GU] 

Thiloa paraguariensis Eichl. [FG] 

COMBRETACEAE - SYNONYMS AND 


Buceras bucida Crantz =Bucida buceras L.[not in Guianas] 

Buchenavia capitata (Vahl) Eichl. =Buchenavia tetraphylla 

Bucida buceras L. [not in Guianas] 

Cacoucia coccinea Aubl. =Combretum cacoucia 

Combretum brunnescens Gleason =Combretum laxum 

Combretum fulgens Gleason =Combretum laxum 

Combretum fusiforme Gleason =Combretum glabrum 

Combretum jacquinii Griseb. =Combretum laxum 

Combretum laxum Aubl. =Combretum rotundifolium 

Pamea guianensis Aubl. =Buchenavia guianensis 

Tanibouca guianensis Aubl. =Terminalia dichotoma 

Terminalia nitidissima L.C. Rich. =Buchenavia nitidissima 

Terminalia pamea DC. =Buchenavia guianensis 

COMMELINACEAE 

contributed by R. Faden (1992) 

REFERENCES: 

Aristeguieta, L. 1965. Notas sobre la familia 

Commelinaceae en Venezuela. Bol. Acad. 

Cienc. Físicas, Matemáticas y Naturales 

25(69): 94-142. 

Uittien, H. 1938. Commelinaceae. In A. Pulle, 


Aneilema umbrosum (Vahl) Kunth subsp. ovato-oblongum (P. 

Beauv.) J.K. Morton [GU SU FG] 

Buforrestia candolleana C.B. Clarke [SU FG] 

Callisia repens L. [FG] 

Commelina benghalensis L. [FG] [naturalized] 

Commelina caroliniana Walter [GU] 

Commelina diffusa Burm.f. [GU SU FG] 

Commelina erecta L. [GU SU FG] 

Commelina obliqua Vahl [FG] 

Commelina platyphylla Klotzsch ex C.B. Clarke [GU] 

Commelina rufipes Seub. var. glabrata (D.R. Hunt) Faden & 

D.R. Hunt [GU SU FG] 

Commelina rufipes Seub. var. rufipes [GU SU FG] 

Commelina schomburgkiana Klotzsch [GU] 

Dichorisandra diederichsanae Steyerm. [GU] 

Dichorisandra hexandra (Aubl.) Standl. [GU SU FG] 

Floscopa peruviana C.B. Clarke [SU FG] 

Gibasis geniculata (Jacq.) Rohweder [GU SU FG] 

Murdannia nudiflora (L.) Brenan [SU FG?] 

Murdannia schomburgkiana (Kunth) Brückner [GU] 

Tinantia umbellata (Vahl) Urb. [GU] 

Tradescantia zanonia (L.) Swartz [GU] 

Tradescantia zebrina Hort. ex Bosse [GU] 

Tripogandra glandulosa (Seub.) Rohw. [FG] 

Tripogandra serrulata (Vahl) Handlos [GU SU FG] 

SPERMATOPHYTA: Combretaceae - Connaraceae 

COMMELINACEAE - SYNONYMS AND 


Athyrocarpus persicariifolius (DC.) Hemsley =Commelina 

rufipes var. glabrata 

Athyrocarpus rufipes (Seub.) Standl. =Commelina rufipes var. 

rufipes 

Commelina hexandra Aubl. =Dichorisandra hexandra 

Commelina nudiflora L. =Murdannia nudiflora 

Commelina serrulata Vahl =Tripogandra serrulata 

Commelina virginica L. [not in Guianas] 

Commelinopsis persicariifolia (DC.) M. Pichon =Commelina 

rufipes var. glabrata 

Dichorisandra incurva Roem. & Schult. [not in Guianas] 

Dichorisandra leucophthalmos Hook. [not in Guianas] 

Rhoeo discolor (L'Hér.) Hance =Rhoeo spathacea (Swartz) 

Stearn[cultivated only] 

Tinantia erecta (Jacq.) Schlecht. [not in Guianas] 

Tradescantia elongata G. Mey. [not in Guianas] 

Tradescantia geniculata Jacq. =Gibasis geniculata 

Tradescantia glandulosa C.B. Clarke [not in Guianas] 

CONNARACEAE 

reviewed by E. Forero (1992); reviewed by E. 

Forero (1996) 

REFERENCES: 

Forero, E. 1983. Connaraceae. Flora Neotropica 

Monogr. 36: 1-207. 

Forero, E. & A.R.A. Görts-van Rijn. 1976. 

Connaraceae (additions and corrections). In J. 

Lanjouw & A.L. Stoffers, Flora of Suriname 

2(2): 654-657. 

Lanjouw, J. 1940. Connaraceae. In A. Pulle, 


Cnestidium guianense (Schellenb.) Schellenb. [GU SU FG] 

Connarus coriaceus Schellenb. [GU SU FG] 

Connarus erianthus Benth. ex Baker var. stipitatus Forero [SU 

FG?] 

Connarus fasciculatus (DC.) Planch. subsp. fasciculatus [SU 

FG] 

Connarus incomptus Planch. [GU] 

Connarus lambertii (DC.) Sagot [GU] 

Connarus megacarpus S.F. Blake [GU] 

Connarus patrisii (DC.) Planch. [GU SU FG] 

Connarus perrottetii (DC.) Planch. var. perrottetii [GU SU 

FG] 

Connarus perrottetii (DC.) Planch. var. rufus Forero [GU] 

Connarus punctatus Planch. [GU SU FG] 

Pseudoconnarus macrophyllus (Poepp. & Endl.) Radlk. [GU] 

Pseudoconnarus subtriplinervis (Radlk.) Schellenb. [GU SU] 

Rourea frutescens Aubl. [GU SU FG] 

Rourea grosourdyana Baill. var. grosourdyana [GU] 

Rourea kappleri Lanj. [SU] 

Rourea ligulata Baker [GU?] 

Rourea neglecta Schellenb. var. neglecta [FG] 

Rourea pubescens (DC.) Radlk. var. pubescens [FG]


Rourea pubescens (DC.) Radlk. var. spadicea (Radlk.) Forero 

[SU FG] 

Rourea revoluta Planch. var. glabra Baker [GU] 

Rourea revoluta Planch. var. revoluta [GU] 

Rourea surinamensis Miq. [GU SU FG] 

CONNARACEAE - SYNONYMS AND 


Bernardinia guianensis Schellenb. =Cnestidium guianense 

Connarus confertiflorus Baker =Connarus patrisii 

Connarus floribundus Planch. =Connarus perrottetii 

Connarus guianensis Lambert ex DC. =Connarus lambertii 

Connarus incomptus Planch. var. subcordata Baker 

=Connarus incomptus 

Connarus macrophyllus Poepp. & Endl. =Pseudoconnarus 

macrophyllus 

Connarus neglectus Schellenb. =Connarus coriaceus 

Connarus opacus Schellenb. =Connarus coriaceus 

Connarus pubescens DC. =Rourea pubescens 

Connarus schomburgkii Planch. =Connarus lambertii 

Omphalobium fasciculatum DC. =Connarus fasciculatus 

Omphalobium patrisii DC. =Connarus patrisii 

Omphalobium perrottetii DC. =Connarus perrottetii 

Robergia frutescens Willd. =Rourea frutescens 

Rourea glabra Kunth var. coriacea Baker =Rourea 

surinamensis 

Rourea induta Planch. var. concinna Baker =Rourea 

frutescens 

Rourea spadicea Radlk. =Rourea pubescens var. spadicea 

Rourea subtriplinervis Radlk. =Pseudoconnarus subtriplinervis 

Santalodes frutescens (Aubl.) Kuntze =Rourea frutescens 

Santalodes revolutum (Planch.) Kuntze =Rourea revoluta 

CONVOLVULACEAE 

contributed by D.F. Austin (1992); additions and 

corrections by D.F. Austin (1996) 

REFERENCES: 

Austin, D.F. 1982. Convolvulaceae. In T. 

Lasser, Flora de Venezuela. 

Austin, D.F. & P.B. Cavalcante. 1982. 

Convolvuláceas da Amazônia. Publ. Avulsas 

Mus. Paraense E. Goeldi 36: 1-134. 

Austin, D.F. & G.W. Staples III. 1981. 

Convolvulaceae. In B. Maguire & 


Highland Part XI. Mem. N.Y. Bot. Gard. 32: 

1-391. 

Ooststroom, S.J. van. 1932. Convolvulaceae. In 

A. Pulle, Flora of Suriname 4(1): 66-102. 

Ooststroom, S.J. van. 1937. Convolvulaceae 

(additions and corrections). In A. Pulle, Flora 


Aniseia cernua Moricand [GU SU FG] [=Aniseia minor 

(Choisy) McDonald?] 

Aniseia martinicensis (Jacq.) Choisy [GU SU FG] 

Bonamia maripoides H. Hallier [GU SU FG] 

Calycobolus glaber (Kunth) House [GU] 

Dicranostyles ampla Ducke var. ampla [GU SU?] 

Dicranostyles ampla Ducke var. castanea D. Austin [GU SU?] 

Dicranostyles guianensis Mennega [GU SU FG] 

Dicranostyles scandens Benth. [GU?] 

Dicranostyles villosus Mennega [SU FG] 

Evolvulus alsinoides (L.) L. [GU SU FG] 

Evolvulus convolvuloides (Willd.) Stearn [GU SU? FG?] 

Evolvulus filipes Mart. [GU SU FG] 

Evolvulus glomeratus Nees & Mart. [GU] 

Evolvulus nummularius (L.) L. [GU SU FG] 

Evolvulus sericeus Swartz [GU SU] 

Ipomoea alba L. [GU SU FG] 

Ipomoea angulata Mart. ex Choisy [FG?] 

Ipomoea anisomeres Robins. & Bart. [GU SU FG?] 

Ipomoea aquatica Forsk. [GU SU FG] 

Ipomoea argentea Meisn. [GU SU? FG?] 

Ipomoea asarifolia (Desr.) Roem. & Schult. [GU?] 

Ipomoea batatas (L.) Poir. [GU SU FG] [cultivated and 

frequently escaping] 

Ipomoea batatoides Choisy [GU SU FG] 

Ipomoea bignonioides Lindl. [FG] 

Ipomoea cairica (L.) Sweet [GU] 

Ipomoea carnea Jacq. subsp. fistulosa (Choisy) D.F. Austin 

[GU SU?] 

Ipomoea distans Choisy [FG] 

Ipomoea fimbrisepala Choisy [FG?] 

Ipomoea gabrieli Choisy [FG] 

Ipomoea hederifolia L. [GU SU] 

Ipomoea horsfalliae Hook. [SU] 

Ipomoea imperati (Vahl) Griseb. [GU SU FG] 

Ipomoea indica (Burm.f.) Merrill [GU?] 

Ipomoea killipiana O'Donell [FG] 

Ipomoea leprieurii D. Austin [FG] 

Ipomoea mauritiana Jacq. [GU SU FG] 

Ipomoea nil (L.) Roth [GU SU FG] 

Ipomoea pes-caprae (L.) R. Brown [GU SU FG] 

Ipomoea phillomega (Vell.) House [GU SU FG] 

Ipomoea piurensis O'Donell [GU] 

Ipomoea quamoclit L. [GU SU FG] 

Ipomoea rubens Choisy [GU SU FG] 

Ipomoea schomburgkii Choisy [GU SU] 

Ipomoea setifera Poir. [GU SU FG] 

Ipomoea squamosa Choisy [GU SU?] 

Ipomoea subrevoluta Choisy [GU SU FG] 

Ipomoea tiliacea (Willd.) Choisy [GU SU FG] 

Ipomoea trifida (Kunth) G. Don [GU?] 

Ipomoea violacea L. [GU SU FG] 

Iseia luxurians (Moric.) O'Donell [GU?] 

Jacquemontia agrestis (Choisy) Meissn. [GU] 

Jacquemontia ciliata Sandw. [GU FG] 

Jacquemontia guyanensis (Aubl.) Meissn. [GU SU FG] 

Jacquemontia pentantha (Jacq.) G. Don [FG] 

Jacquemontia sphaerostigma (Cav.) Rusby [GU] 

Jacquemontia tamnifolia (L.) Griseb. [GU SU FG] 

Jacquemontia velutina Choisy [SU?] 

Lysiostyles scandens Benth. [GU SU] 

Maripa glabra Choisy [GU SU FG] 

Maripa longifolia Sagot ex H. Hallier [FG] 

Maripa paniculata Barb. Rodr. [SU? FG] 

Maripa reticulata Ducke var. reticulata [SU FG] 

Maripa scandens Aubl. [GU SU FG] 

Maripa violacea (Aubl.) v. Ooststr. [GU SU FG] 

Merremia aegyptia (L.) Urb. [GU SU FG] 

SPERMATOPHYTA: Connaraceae - Convolvulaceae 

87

88 


Merremia aturensis (Kunth) H. Hallier [GU SU FG] 

Merremia cissoides (Lam.) H. Hallier [GU SU FG] 

Merremia digitata (Spreng.) H. Hallier [GU] 

Merremia dissecta (Jacq.) H. Hallier [GU SU FG] 

Merremia macrocalyx (Ruiz & Pav.) O'Donell [GU SU FG] 

Merremia quinquefolia (L.) H. Hallier [GU] 

Merremia ternifolia Pitt. [GU] 

Merremia umbellata (L.) H. Hallier [GU SU FG] 

Merremia wurdackii D. Austin & Staples [FG] 

Operculina hamiltonii (G. Don) Austin & Staples [GU SU FG] 

Operculina sericantha (Miq.) v. Ooststr. [SU FG] 

Operculina turpethum (L.) S. Manso [FG] [var. turpethum?] 

[probably introduced] 

Operculina turpethum (L.) S. Manso var. ventricosa (Bertero) 

Staples & Austin [SU] [probably introduced] 

Tetralocularia pennellii O'Donell [FG] 

CONVOLVULACEAE - SYNONYMS AND 


Convolvulus glaber Aubl. =Merremia macrocalyx 

Convolvulus guianensis Aubl. =Jacquemontia guyanensis 

Ipomoea digitata L. [not in Guianas] 

Ipomoea fragrans (Boj.) Boj. ex Choisy =Ipomoea rubens 

Ipomoea parkeri Choisy =Ipomoea rubens 

Ipomoea quinquefolia (L.) Griseb. =Merremia quinquefolia 

Ipomoea reptans (L.) Poir. =Ipomoea aquatica 

Ipomoea riedelii Meissn. =Ipomoea batatoides 

Ipomoea solanifolia L. [misapplied name, not in Guianas] 

Ipomoea squamisepala O'Donell =Ipomoea angulata 

Ipomoea stolonifera (Cyrillo) J.F. Gmelin =Ipomoea imperati 

Ipomoea tuba (Schlecht.) G. Don =Ipomoea violacea 

Ipomoea tuberosa L. =Merremia tuberosa (L.) Rendle [not in 


Jacquemontia violacea (Vahl) Choisy =Jacquemontia 

pentantha 

Maripa cayennensis Meisn. =Maripa scandens 

Maripa passifloroides Benth. =Maripa violacea 

Merremia glabra (Aubl.) H. Hallier =Merremia macrocalyx 

Mouroucoa violacea Aubl. =Maripa violacea 

Operculina alata (Ham.) Urb. =Operculina hamiltonii 

Operculina passifloroides (Benth.) Ducke =Maripa violacea 

Operculina triquetra (Vahl) H. Hallier =Operculina turpethum 

Operculina ventricosa Peter =Operculina turpethum var. 

ventricosa 

COSTACEAE 


P.J.M. Maas (1996) 

REFERENCES: 

Maas, P.J.M. 1972. Costoideae (Zingiberaceae). 


Maas, P.J.M. 1977. Costoideae (additions). 


Maas, P.J.M. 1979. Zingiberaceae. In A.L. 


5(1): 416-441. 

Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. 


A) 1: 29-67. 

SPERMATOPHYTA: Convolvulaceae - Crassulaceae 

Costus arabicus L. [GU SU FG] 

Costus claviger Benoist [GU SU FG] 

Costus congestiflorus L.C. Rich. ex Gagnep. [GU SU FG] 

Costus curcumoides Maas [FG] 

Costus erythrothyrsus Loes. [GU SU FG] 

Costus guanaiensis Rusby var. macrostrobilus (K. Schum.) 

Maas [GU] 

Costus lanceolatus Petersen subsp. pulchriflorus (Ducke) 

Maas [SU FG] 

Costus lasius Loes. [FG] 

Costus scaber Ruiz & Pav. [GU SU FG] 

Costus spiralis (Jacq.) Roscoe var. spiralis [GU SU FG] 

Costus spiralis (Jacq.) Roscoe var. villosus Maas [GU SU FG] 

Costus villosissimus Jacq. [GU FG?] 

Dimerocostus strobilaceus Kuntze subsp. gutierrezii (Kuntze) 

Maas [GU SU] 

COSTACEAE - SYNONYMS AND 


Costus bracteatus Gleason =Costus claviger 

Costus ciliatus Miq. =Costus scaber 

Costus congestiflorus L.C. Rich. ex Gagnep. var. glabrior K. 

Schum. =Costus congestiflorus 

Costus cylindricus Auct. non Jacq. =Costus scaber 

Costus cylindricus Jacq. var. ciliatus (Miq.) O.G. Petersen 

=Costus scaber 

Costus discolor Roscoe =Costus arabicus 

Costus guianicus Loes. =Costus claviger 

Costus niveus G. Mey. =Costus arabicus 

Costus pulchriflorus Ducke =Costus lanceolatus subsp. 

pulchriflorus 

Costus ramosus Woodson =Costus arabicus 

Costus scaberulus L.C. Rich. ex Gagnep. =Costus scaber 

Costus spicatus Auct. non Sessé & Moç. =Costus scaber 

REFERENCES: 

CRASSULACEAE 

Stoffers, A.L. 1984. Crassulaceae. In A.L. 


V(1): 573-575. 

Bryophyllum pinnatum (Lam.) Kurz [GU SU FG] [cultivated, 

naturalized] 

Kalanchoe crenata (Andr.) Haw. [GU] [cultivated or escaped 


CRASSULACEAE - SYNONYMS AND 


Bryophyllum calycinum Salisb. =Bryophyllum pinnatum 

Kalanchoe brasiliensis Cambess. =Kalanchoe crenata


Kalanchoe pinnata (Lam.) Pers. =Bryophyllum pinnatum 

CUCURBITACEAE 

reviewed by C. Jeffrey & M. Nee (1992); 

reviewed by M. Nee (1996) 

REFERENCES: 

Jeffrey, C. 1978. Further notes on 

Cucurbitaceae: IV. Some New World taxa. 

Kew Bull. 33: 347-380. 

Jeffrey, C. 1984. Cucurbitaceae. In A.L. 


5(1): 457-518. 

Jeffrey, C. & B. Trujillo. 1992. Cucurbitaceae. 

In G. Morillo, Flora de Venezuela 5(1): 11- 

201. 

Cayaponia angustiloba (Cogn.) Cogn. [GU SU] 

Cayaponia coriacea Cogn. [GU SU FG] 

Cayaponia cruegeri (Naud.) Cogn. [GU SU FG] 

Cayaponia guianensis C. Jeffrey [GU] 

Cayaponia jenmanii C. Jeffrey [GU SU FG] 

Cayaponia ophthalmica R.E. Schultes [GU SU FG] 

Cayaponia racemosa (Mill.) Cogn. [GU FG] 

Cayaponia rigida (Cogn.) Cogn. [GU SU FG] 

Cayaponia selysioides C. Jeffrey [GU SU FG] 

Cayaponia tessmannii Harms [FG?] 

Cayaponia triangularis (Cogn.) Cogn. [GU SU FG] 

Ceratosanthes palmata (L.) Urb. [GU SU FG] 

Citrullus lanatus (Thunb.) Matsumura & Nakai [GU SU FG] 

[cultivated, occasionally escaping] 

Coccinia grandis (L.) Voigt [GU] [cultivated, occasionally 

escaping] 

Cucumis anguria L. [GU? SU FG] [cultivated, naturalized] 

Cucumis melo L. [GU? SU FG?] [cultivated, occasionally 

escaping] 

Cucurbita moschata (Lam.) Poir. [GU SU FG] [cultivated, 


Fevillea cordifolia L. [FG] 

Fevillea trilobata L. [FG?] 

Gurania acuminata Cogn. [GU SU FG] 

Gurania bignoniacea (Poepp. & Endl.) C. Jeffrey [GU SU FG] 

Gurania brevipedunculata Cogn. [GU FG] 

Gurania huberi Cogn. [GU SU FG] 

Gurania oxyphylla C. Jeffrey [GU SU FG] 

Gurania reticulata Cogn. [FG] 

Gurania robusta Suessang. [SU FG] 

Gurania simplicifolia (Steyerm.) C. Jeffrey [GU] 

Gurania spinulosa (Poepp. & Endl.) Cogn. [GU SU FG] 

Gurania subumbellata (Miq.) Cogn. [GU SU FG] 

Helmontia cardiophylla Harms [FG] 

Helmontia leptantha (Schlechtend.) Cogn. [GU SU FG] 

Kedrostis foetidissima (Jacq.) Cogn. [SU] [naturalized?] 

Lagenaria siceraria (Molina) Standl. [GU SU FG] [cultivated, 


Luffa cylindrica (L.) M.J. Roem. [GU SU FG] [cultivated, 


Luffa sepium (G. Mey.) C. Jeffrey [GU SU FG] 

Melothria pendula L. [GU SU FG] 

Melothria trilobata Cogn. [GU SU] 

Momordica charantia L. [GU SU FG] [cultivated, naturalized] 

Posadaea sphaerocarpa Cogn. [GU SU FG] 

Psiguria triphylla (Miq.) C. Jeffrey [GU SU FG] 

Rytidostylis amazonica (Cogn.) Kuntze [FG] 

Selysia prunifera (Poepp. & Endl.) Cogn. [GU? SU FG] 

Siolmatra pentaphylla Harms [GU] 

Wilbrandia hibiscoides Manso [GU?] 

CUCURBITACEAE - SYNONYMS AND 


Anguria bignoniacea Poepp. & Endl. =Gurania bignoniacea 

Anguria guianensis Klotzsch ex Schlechtend. =Gurania 

bignoniacea 

Anguria kegeliana Schlechtend. =Gurania spinulosa 

Anguria leptantha Schlechtend. var. brevipes Schlechtend. 

=Helmontia leptantha 

Anguria longispica Sagot [name of uncertain application] 

Anguria multiflora Miq. =Gurania spinulosa 

Anguria subumbellata Miq. =Gurania subumbellata 

Anguria treslingiana Pulle =Psiguria triphylla 

Anguria trilobata sensu Pulle non (L.) Jacq. =Gurania 

spinulosa 

Anguria triphylla Miq. =Psiguria triphylla 

Anguria umbrosa Auct. non Kunth =Psiguria triphylla 

Apodanthera spp. [no records verified for Guianas] 

Cayaponia calycina Cogn. =Cayaponia cruegeri 

Cayaponia sclerocarpa Sagot [name of uncertain application] 

Cayaponia simplicifolia (Naud.) Cogn. [not in Guianas] 

Citrullus vulgaris Schrad. =Citrullus lanatus 

Elaterium amazonicum Mart. ex Cogn. =Rytidostylis 

amazonica 

Elaterium carthaginense Jacq. =Rytidostylis carthaginensis 

(Jacq.) Kuntze [not in Guianas] 

Gurania breviflora Cogn. =Gurania bignoniacea 

Gurania cissoides (Benth.) Cogn. =Gurania bignoniacea 

Gurania diffusa Cogn. =Gurania huberi 

Gurania diversifolia Cogn. =Gurania acuminata 

Gurania durandii Cogn. =Gurania spinulosa 

Gurania guianensis (Klotzsch ex Schlechtend.) Cogn. 

=Gurania bignoniacea 

Gurania kegeliana (Schlechtend.) Cogn. =Gurania spinulosa 

Gurania klotzschiana Cogn. =Gurania subumbellata 

Gurania levyana Cogn. var. glabra Cogn. =Gurania robusta 

Gurania multiflora (Miq.) Cogn. =Gurania spinulosa 

Gurania sagotiana Cogn. =Gurania reticulata 

Helmontia simplicifolia Cogn. =Helmontia leptantha 

Lagenaria vulgaris Ser. =Lagenaria siceraria 

Luffa operculata (L.) Cogn. [not in Guianas] 

Melothria fluminensis Gardn. =Melothria pendula 

Melothria guadalupensis (Spreng.) Cogn. =Melothria pendula 

Momordica balsamina L. [Guianan records dubious] 

Rytidostylis carthaginensis (Jacq.) Kuntze [not in Guianas] 

Trianosperm rigida Cogn. =Cayaponia rigida 

Trianosperma triangularis Cogn. =Cayaponia triangularis 

CUNONIACEAE 

reviewed by P. Berry (1996) 

SPERMATOPHYTA: Crassulaceae - Cunoniaceae 

89

90 

REFERENCES: 


Killip, E.P. & A.C. Smith. 1929. The genus 

Weinmannia in northern South America. 


Weinmannia brachystachya Willd. [to be expected in GU (Mt. 

Roraima)] 

Weinmannia fagaroides Kunth [GU?] [Mt. Roraima] 

Weinmannia guyanensis Klotzsch ex Engl. [GU] [Mt. 

Roraima] 

Weinmannia laxiramea Killip & A.C. Smith [GU] [Mt. 

Roraima] 

Weinmannia pinnata L. [GU?] [Mt. Roraima] 

Weinmannia velutina Schmidt [GU?] [Mt. Roraima] 

CUNONIACEAE - SYNONYMS AND 


Weinmannia pinnata L. var. caripensis (H.B.K.) Cuatr. 

=Weinmannia pinnata 

Weinmannia roraimensis Cuatrec. =Weinmannia fagaroides 

Weinmannia steyermarkii Cuatrec. =Weinmannia velutina 

REFERENCES: 

CUSCUTACEAE 

Austin, D.F. 1982. Convolvulaceae. In Flora de 

Venezuela. 

Yuncker, T.G. 1923. Revision of the South 

American species of Cuscuta. I. Am. J. Bot. 

9: 557-575; II. Am. J. Bot. 10: 1-17. 

Cuscuta americana L. [SU] 

Cuscuta umbellata Kunth [GU SU] 

REFERENCES: 

CYCADACEAE 

Stevenson, D.W. 1991. Cycadaceae. In A.R.A. 


A) 9: 3-7. 

Cycas spp. [species of this Old World family are widely 

cultivated] 

SPERMATOPHYTA: Cunoniaceae - Cyclanthaceae 

REFERENCES: 

CYCLANTHACEAE 

Harling, G. 1958. Monograph of the 

Cyclanthaceae. Acta Horti Berg. 18: 1-428. 

Harling, G. 1972. Cyclanthaceae. In B. Maguire 



23: 1-832. 

Asplundia brachyphylla Harling [GU SU FG] 

Asplundia fanshawei (Maguire) Harling [GU SU] 

Asplundia glandulosa (Gleason) Harling [GU SU FG] 

Asplundia gleasonii Harling [GU] 

Asplundia guianensis Harling [GU] 

Asplundia heteranthera Harling [SU FG] 

Asplundia maguirei Harling [GU SU] 

Carludovica palmata Ruiz & Pav. [FG?] 

Cyclanthus bipartitus Poit. [GU FG] 

Dicranopygium angustissimum (Sandw.) Harling [GU] 

Dicranopygium nanum (Gleason) Harling [GU] 

Dicranopygium pygmaeum (Gleason) Harling subsp. 

fimbriatum (Maguire) Harling [SU FG] 

Dicranopygium pygmaeum (Gleason) Harling subsp. 

pygmaeum [GU SU FG] 

Evodianthus funifer (Poit.) Lindm. subsp. funifer [GU SU 

FG] 

Evodianthus funifer (Poit.) Lindm. subsp. trailianus (Drude) 

Harling [GU SU FG] 

Ludovia lancifolia Brongn. [SU FG] [to be expected in GU] 

Stelestylis stylaris (Gleason) Harling [GU] 

Stelestylis surinamensis Harling [SU FG?] 

Thoracocarpus bissectus (Vell.) Harling [GU SU FG] 

CYCLANTHACEAE - SYNONYMS AND 


Carludovica acuminata (Ruiz & Pav.) Harling [not in Guianas] 

Carludovica angustissima Sandw. =Dicranopygium 

angustissimum 

Carludovica coronata Gleason =Evodianthus funifer subsp. 

funifer 

Carludovica disticha Neum. =Ludovia lancifolia 

Carludovica fanshawei Maguire =Asplundia fanshawei 

Carludovica fimbriata Maguire =Dicranopygium pygmaeum 

subsp. fimbriatum 

Carludovica glandulosa Gleason =Asplundia glandulosa 

Carludovica kegeliana Lemaire =Thoracocarpus bissectus 

Carludovica latifrons Drude =Asplundia latifrons (Drude) 

Harling [not in Guianas] 

Carludovica nana Gleason =Dicranopygium nanum 

Carludovica pygmaea Gleason =Dicranopygium pygmaeum 

Carludovica sarmentosa Sagot ex Drude =Thoracocarpus 

bissectus 

Carludovica stylaris Gleason =Stelestylis stylaris


CYPERACEAE 

contributed by P. Goetghebeur & M.T. Strong 

(1992); additions and corrections by M.T. 

Strong, 1996 

REFERENCES: 

Core, E.L. 1965. Cyperaceae tribe Sclerieae. In 

B. Maguire & collaborators, Botany of the 

Guayana Highland Part VI. Mem. New York 

Bot. Gard. 12(3): 1-285. 



française III - les Cyperaceae et les Poaceae. 

Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Sér. 

Patr. Génétique) 11: 1-213. 

Koyama, T. and B. Maguire. 1965. Cyperaceae 

tribe Lagenocarpeae. In B. Maguire & 



12(3): 1-285. 

Koyama, T. 1967. Cyperaceae - Mapanioideae. 



Bot. Gard. 17(1): 1-439. 

Koyama, T. 1972. Cyperaceae - 

Rhynchosporeae and Cladieae. In B. Maguire 



23: 1-832. 

Kral, R. 1971. A treatment of Abildgaardia, 

Bulbostylis, and Fimbristylis (Cyperaceae) for 

North America. Sida 4: 57-227. 

Lindeman, J.C. & A.R.A. Görts-van Rijn. 1968. 

Cyperaceae (additions and corrections). In A. 


310-314. 

Simpson, D.A. 1992. A revision of the genus 

Mapania. Royal Botanic Gardens, Kew. 

Thomas, W.W. 1984. The systematics of 

Rhynchospora section Dichromena. Mem. 


Uittien, H. 1934. Cyperaceae. In A. Pulle, Flora 


Uittien, H. 1943. Cyperaceae (additions and 

corrections). In A. Pulle [ed.], Flora of 

Suriname 1(1): 492-505. 

Abildgaardia ovata (Burm.f.) Kral [GU] 

Ascolepis brasiliensis (Kunth) Benth. ex C.B. Clarke [GU SU] 

Becquerelia cymosa Brongn. subsp. cymosa [GU SU FG] 

Becquerelia cymosa Brongn. subsp. merkeliana (Nees) T. 

Koyama [GU SU FG] 

Becquerelia tuberculata (Boeck.) H. Pfeiff. [GU SU] 

Bisboeckelera irrigua (Nees) Kuntze [GU] 

Bisboeckelera longifolia (Rudge) Kuntze [GU SU FG] 

Bisboeckelera microcephala (Boeck.) T. Koyama [GU SU FG] 

Bulbostylis capillaris (L.) C.B. Clarke var. capillaris [GU SU 

FG] 

Bulbostylis capillaris (L.) C.B. Clarke var. major Uittien [SU] 

Bulbostylis circinata (Nees) C.B. Clarke [GU SU FG] 

Bulbostylis conifera (Kunth) C.B. Clarke [GU SU FG] 

Bulbostylis fasciculata Uittien [GU SU FG] 

Bulbostylis hirta (Thunb.) Svenson [GU] 

Bulbostylis junciformis (Kunth) C.B.Clarke var. glabrescens 

Boeck. [SU] 

Bulbostylis junciformis (Kunth) C.B.Clarke var. junciformis 

[GU SU FG] 

Bulbostylis juncoides (Vahl) Kükenth. [GU FG] 

Bulbostylis lanata (Kunth) C.B. Clarke [GU SU FG] 

Bulbostylis paradoxa (Spreng.) Lindm. [GU SU FG] 

Bulbostylis stenocarpa Kükenth. [GU SU FG] 

Bulbostylis surinamensis Uittien [GU SU FG?] 

Bulbostylis tenuifolia (Rudge) Macbr. [GU SU FG] 

Bulbostylis truncata (Nees) M.T. Strong [GU SU] 

Bulbostylis vestita (Kunth) C.B. Clarke var. glabrescens 

Boeck. [SU] 

Bulbostylis vestita (Kunth) C.B. Clarke var. vestita [SU FG] 

Calyptrocarya bicolor (H. Pfeiff.) T. Koyama [GU SU FG] 

Calyptrocarya glomerulata (Brongn.) Urb. var. glomerulata 

[GU SU FG] 

Calyptrocarya luzuliformis T. Koyama [GU SU] 

Calyptrocarya monocephala Hochst. ex Steud. [GU SU FG] 

Calyptrocarya poeppigiana Kunth [GU SU FG] [records may 

be referable to another species] 

Carex polystachya Sw. ex Wahlenb. [GU] 

Cephalocarpus rigidus Gilly ex Gleason & Killip [GU SU] 

Cladium costatum Steyerm. [GU?] [Mt. Roraima] 

Cladium mariscus Pohl [SU?] 

Cyperus aggregatus (Willd.) Endl. [GU SU FG] 

Cyperus altsonii Kükenth. [GU] 

Cyperus amabilis Vahl [GU SU] 

Cyperus articulatus L. [GU SU FG] [var. articulatus?] 

Cyperus articulatus L. var. nodosus (Willd.) Kükenth. [FG] 

Cyperus capillifolius A. Rich. [GU?] 

Cyperus chlorocephalus (C.B. Clarke) Kükenth. [GU FG] 

Cyperus comosus Poir. [GU SU FG] 

Cyperus compressus L. [GU SU FG] 

Cyperus corymbosus Rottb. [GU SU] 

Cyperus croceus Vahl [GU SU FG] 

Cyperus cuspidatus Kunth [GU SU FG?] 

Cyperus difformis L. [GU] 

Cyperus digitatus Roxb. [GU SU] 

Cyperus distans L.f. [GU SU FG] 

Cyperus eragrostis Lam. [SU] 

Cyperus filifolius Willd. ex Kunth [GU] 

Cyperus flavescens L. [GU SU?] 

Cyperus gayi (C.B. Clarke) Kükenth. [FG] 

Cyperus giganteus Vahl [GU SU FG] 

Cyperus guianensis (C.B. Clarke) Kükenth. [GU SU] 

Cyperus haspan L. [GU SU FG] [subsp. haspan?] 

Cyperus haspan L. subsp. juncoides (Lam.) Kükenth. [GU SU 

FG] 

Cyperus haspan L. var. riparius (Nees) Kükenth. [GU FG] 

Cyperus hermaphroditus (Jacq.) Standl. [GU FG?] 

Cyperus hoppiifolius Uittien [SU FG] 

Cyperus imbricatus Retz. [GU] [var. imbricatus?] 

Cyperus imbricatus Retz. var. elongatus (Boeck.) Kükenth. 

[GU] 

Cyperus involucratus Rottb. [SU] [escaped from cultivation] 

Cyperus iria L. [GU] [introduced] 

Cyperus kappleri Hochst. ex Steud. [GU SU FG] 

Cyperus laxus Lam. s.l. [GU SU FG] 

Cyperus ligularis L. [GU SU FG] 

Cyperus luzulae (L.) Rottb. ex Retz. var. luzulae [GU SU FG] 

SPERMATOPHYTA: Cyperaceae 

91

92 


Cyperus luzulae (L.) Rottb. ex Retz. var. microcephalus (Nees) 

Nees [GU] 

Cyperus megapotamicus Kunth [SU?] 

Cyperus miliifolius Poepp. & Kunth [GU SU FG] 

Cyperus mutisii (Kunth) Griseb. [GU FG] 

Cyperus odoratus L. [GU SU FG] 

Cyperus oxylepis Nees ex Steud. [GU] 

Cyperus polystachyos Rottb. [GU SU FG] 

Cyperus prolixus Kunth [GU SU FG?] 

Cyperus pseudodistans Uittien [SU] 

Cyperus pumilus L. [FG] 

Cyperus rotundus L. [GU SU FG] 

Cyperus schomburgkianus Nees [GU] 

Cyperus simplex Kunth [GU SU FG] 

Cyperus sphacelatus Rottb. [GU SU FG] 

Cyperus subumbellatus Kükenth. [SU] 

Cyperus surinamensis Rottb. [GU SU FG] [var. 

surinamensis?] 

Cyperus surinamensis Rottb. var. lutescens Boeck. [SU FG] 

Cyperus trailii C.B. Clarke [GU SU] 

Cyperus unioloides R. Brown [SU] 

Cyperus virens Michx. [SU] 

Didymiandrum stellatum (Boeck.) Gilly [GU] 

Diplacrum africanum (Benth.) C.B. Clarke [SU] 

Diplacrum capitatum (Willd.) Boeck. [GU SU FG] 

Diplacrum guianense (Nees) T. Koyama [GU SU FG] 

Diplasia karataefolia L.C. Rich. [GU SU FG] 

Eleocharis acutangula (Roxb.) Schult. [GU SU] 

Eleocharis debilis Kunth [GU] 

Eleocharis elegans (Kunth) Roem. & Schult. [GU SU FG] 

Eleocharis filiculmis Kunth [GU SU FG] 

Eleocharis flaccida (Rchb. ex Spreng.f.) Urb. [GU SU FG] 

Eleocharis flavescens (Poir.) Urb. [GU SU FG] 

Eleocharis geniculata (L.) Roem. & Schult. [GU SU FG] 

Eleocharis interstincta (Vahl) Roem. & Schult. [GU SU FG] 

Eleocharis jelskiana Boeck. [GU SU FG] 

Eleocharis maculosa (Vahl) Roem. & Schult. [GU] 

Eleocharis minima Kunth [GU FG?] [var. minima?] 

Eleocharis minima Kunth var. bicolor (Chapm.) Svenson [GU] 

Eleocharis mitrata (Griseb.) C.B. Clarke [GU SU FG] 

Eleocharis mutata (L.) Roem. & Schult. [GU SU FG] 

Eleocharis nana Kunth [GU SU] 

Eleocharis nigrescens (Nees) Steud. [GU] 

Eleocharis nodulosa (Roth) Schult. [SU] 

Eleocharis pachystyla (C. Wright) C.B. Clarke [GU SU FG] 

Eleocharis plicarhachis (Griseb.) Svenson [GU SU FG] 

Eleocharis retroflexa (Poir.) Urb. [GU SU FG] 

Eleocharis sellowiana Kunth [GU FG] [var. sellowiana?] 

Eleocharis sellowiana Kunth var. homonyma (Steud.) H. 

Pfeiff. [FG] 

Eleocharis sp. aff. glauca Boeck. [GU] 

Eleocharis subfoliata C.B. Clarke [GU SU] 

Everardia angusta N.E. Brown [GU] 

Everardia debilis T. Koyama & Maguire [GU] 

Everardia disticha T. Koyama & Maguire [GU] 

Everardia montana Ridley ex im Thurn subsp. montana [GU] 

Everardia surinamensis Gilly [GU SU] 

Exochogyne amazonica C.B. Clarke [GU SU FG] 

Fimbristylis annua (Allioni) Roem. & Schult. [GU SU FG] 

Fimbristylis aspera (Schrad.) Boeck. [GU] 

Fimbristylis autumnalis (L.) Roem. & Schult. [GU? SU FG] 

Fimbristylis complanata (Retz.) Link [SU FG] 

Fimbristylis cymosa R. Brown [GU SU FG] 

Fimbristylis dichotoma (L.) Vahl [GU SU FG] 

Fimbristylis ferruginea (L.) Vahl [GU SU FG] 

Fimbristylis limosa Poepp. & Kunth [GU] 


Fimbristylis miliacea (L.) Vahl [GU SU FG] [naturalized] 

Fimbristylis quinquangularis (Vahl) Knuth [GU] 

Fimbristylis spadicea Vahl [GU SU FG] 

Fimbristylis vahlii Link [GU SU FG] 

Fuirena robusta Kunth [GU SU FG] 

Fuirena simplex Vahl [FG] 

Fuirena umbellata Rottb. [GU SU FG] 

Hypolytrum amplum Poepp. & Kunth ex Kunth [GU SU FG] 

Hypolytrum jenmanii C.B. Clarke subsp. jenmanii [GU SU 

FG] 

Hypolytrum jenmanii C.B. Clarke subsp. serratifolium 

(Uittien) T. Koyama [GU SU FG] 

Hypolytrum longifolium (L.C. Rich.) Nees subsp. longifolium 

[GU SU FG] 

Hypolytrum longifolium (L.C. Rich.) Nees subsp. rubescens 

(Huber ex C.B. Clarke) T. Koyama [GU] 

Hypolytrum longifolium (L.C. Rich.) Nees subsp. sylvaticum 

(Poepp. & Kunth) T. Koyama [GU SU FG] 

Hypolytrum nudum C.B. Clarke [GU] 

Hypolytrum pallidiceps S. Hooper & T. Koyama [GU] 

Hypolytrum pulchrum (Rudge) H. Pfeiff. [GU SU FG] 

Hypolytrum schraderianum Nees [GU FG] 

Hypolytrum spongiosum T. Koyama [GU] 

Hypolytrum stemonifolium T. Koyama [GU] 

Kyllinga brevifolia Rottb. [GU SU FG] 

Kyllinga nemoralis (J.R. & G. Forster) Dandy ex Hutchinson 

[SU FG?] 

Kyllinga odorata Vahl [GU SU FG] 

Kyllinga pumila Michx. [GU SU FG] 

Kyllinga pungens Link [GU SU FG] 

Kyllinga vaginata Lam. [SU FG] 

Lagenocarpus glomerulatus Gilly [GU] 

Lagenocarpus guianensis Lindl. & Nees ex Nees [GU SU FG] 

Lagenocarpus rigidus (Kunth) Nees subsp. tremulus (Nees) T. 

Koyama & Maguire [GU SU FG] 

Lagenocarpus sabanensis Gilly [SU FG] 

Lagenocarpus verticillatus (Spreng.) T. Koyama & Maguire 

[GU SU] 

Lipocarpha humboldtiana Nees [GU SU] 

Lipocarpha micrantha (Vahl) G. Tucker [GU SU FG] 

Lipocarpha salzmanniana Steud. [GU] 

Lipocarpha schomburgkii (Friedland) G. Tucker [GU] 

Mapania assimilis T. Koyama subsp. guianensis D.A. Simpson 

[FG] 

Mapania cuatrecasasii T. Koyama [GU] 

Mapania imeriensis (R. Gross) T. Koyama [GU SU] 

Mapania insignis Sandw. [GU] 

Mapania macrophylla (Boeck.) H. Pfeiff. [GU SU FG] 

Mapania maguireana T. Koyama & Steyerm. [GU] 

Mapania paradoxa J. Raynal [FG] 

Mapania pycnocephala (Benth.) Benth. subsp. fluviatilis 

(Sandw.) T. Koyama [GU FG] 

Mapania steyermarkii T. Koyama [GU] 

Mapania surinamensis Uittien [SU] 

Mapania sylvatica Aubl. subsp. sylvatica [GU SU FG] 

Mapania tepuiana (Steyerm.) T. Koyama [GU FG?] 

Mapaniopsis effusa C.B. Clarke [FG] 

Mapaniopsis micrococca T. Koyama [GU] 

Oxycaryum cubense (Poepp. & Kunth) Palla [GU SU FG] 

Pleurostachys guianensis Uittien [SU] 

Pleurostachys puberula Boeck. [GU FG] 

Remirea maritima Aubl. [SU FG] 

Rhynchocladium steyermarkii (T. Koyama) T. Koyama [GU] 

Rhynchospora albescens (Miq.) Kükenth. [GU SU] 

Rhynchospora albomarginata Kükenth. [GU]


Rhynchospora amazonica Poepp. & Kunth subsp. amazonica 

[GU] 

Rhynchospora amazonica Poepp. & Kunth subsp. guianensis 

(Kükenth.) T. Koyama [GU FG?] 

Rhynchospora arenicola Uittien [GU SU FG] 

Rhynchospora aristata Boeck. [GU] 

Rhynchospora armerioides Presl [GU SU] 

Rhynchospora barbata (Vahl) Kunth [GU SU FG] 

Rhynchospora bolivarana Steyerm. [GU] 

Rhynchospora brachychaeta C.Wright [GU] 

Rhynchospora brasiliensis Boeck. [GU] 

Rhynchospora brevirostris Griseb. [SU FG] 

Rhynchospora cajennensis Boeck. [GU FG] 

Rhynchospora canaliculata Boeck. [FG] 

Rhynchospora candida (Nees) Boeck. [GU SU FG?] 

Rhynchospora caracasana (Kunth) Boeck. [GU SU] 

Rhynchospora cephalotes (L.) Vahl [GU SU FG] [var. 

cephalotes?] 

Rhynchospora cephalotes (L.) Vahl var. angustifolia Uittien 

[SU] 

Rhynchospora comata (Link) Roem. & Schult. [GU SU FG] 

Rhynchospora contracta (Nees) J. Raynal [FG] 

Rhynchospora corymbosa (L.) Britton [GU SU FG] 

Rhynchospora curvula Griseb. [GU SU FG] 

Rhynchospora emaciata (Nees) Boeck. [GU FG] 

Rhynchospora exaltata Kunth [GU] [var. exaltata?] 

Rhynchospora exaltata Kunth var. microcephala Kükenth. 

[GU] 

Rhynchospora fallax Uittien [GU SU FG] 

Rhynchospora fascicularis (Michx.) Vahl var. distans (Michx.) 

Chapman [SU] 

Rhynchospora filiformis Vahl [GU SU FG] 

Rhynchospora flexuosa C.B. Clarke subsp. tepuiana 

(Steyerm.) T. Koyama [GU] 

Rhynchospora gigantea Link [GU SU FG] 

Rhynchospora globosa (Kunth) Roem. & Schult. [GU SU FG] 

Rhynchospora graminea Uittien [GU SU FG?] 

Rhynchospora hassleri C.B. Clarke [GU SU FG] 

Rhynchospora heterocaulis C.B. Clarke [SU] 

Rhynchospora hieronymii Boeck. [GU] 

Rhynchospora hirsuta (Vahl) Vahl [GU SU FG] 

Rhynchospora holoschoenoides (L.C. Rich.) Herter [GU SU 

FG] 

Rhynchospora junciformis (Kunth) Boeck. [GU SU FG] 

Rhynchospora longibracteata Boeck. [GU] 

Rhynchospora macrochaeta Steud. [GU] 

Rhynchospora maguireana T. Koyama [GU] 

Rhynchospora marisculus Nees [GU SU] 

Rhynchospora montana (Uittien) H. Pfeiff. [SU] 

Rhynchospora nervosa (Vahl) Boeck. subsp. ciliata T. 

Koyama [GU SU FG] 

Rhynchospora nervosa (Vahl) Boeck. subsp. nervosa [GU SU 

FG] 

Rhynchospora nitens (Vahl) A. Gray [GU] 

Rhynchospora polyphylla Vahl [GU SU FG] 

Rhynchospora pubera (Vahl) Boeck. subsp. parvula W.W. 

Thomas [GU] 

Rhynchospora pubera (Vahl) Boeck. subsp. pubera [GU SU 

FG] 

Rhynchospora radicans (Schlechtend. & Cham.) H. Pfeiff. 

subsp. microcephala (Bertero ex Spreng.) W. Thomas 

[GU SU FG] 

Rhynchospora reptans (L.C. Rich.) Boeck. [GU SU FG] 

Rhynchospora riparia (Nees) Boeck. [GU SU FG] 

Rhynchospora robusta (Kunth) Boeck. [GU FG?] 

Rhynchospora roraimae Kükenth. [GU] [Mt. Roraima] 

Rhynchospora rugosa (Vahl) Gale [GU SU FG] 

Rhynchospora schomburgkiana (Boeck.) T. Koyama [GU] 

Rhynchospora steyermarkii T. Koyama [GU] 

Rhynchospora stricta Boeck. var. demerarensis (C.B. Clarke) 

Kükenth. [GU] 

Rhynchospora subdicephala T. Koyama [FG] 

Rhynchospora sublanata Lindeman & Donsel. [GU SU] 

Rhynchospora subplumosa C.B. Clarke [GU SU] 

Rhynchospora tenella (Nees) Boeck. var. tenella [GU SU FG] 

Rhynchospora tenerrima Nees ex Spreng. [GU SU FG] 

Rhynchospora tenuis Willd. ex Link subsp. austrobrasiliensis 

T. Koyama [GU] 

Rhynchospora tenuis Willd. ex Link subsp. tenuis [GU SU 

FG] 

Rhynchospora trichochaeta C.B. Clarke [GU SU?] 

Rhynchospora trichodes C.B. Clarke [GU SU FG] 

Rhynchospora triflora (Poir.) Vahl [GU SU FG] 

Rhynchospora trispicata (Nees) Steud. [GU SU FG] 

Rhynchospora velutina (Kunth) Boeck. [GU SU FG] 

Rhynchospora watsonii (Britton) Davidse [GU SU FG] 

Schoenoplectus maritimus (L.) K. Lye [SU FG?] 

Schoenoplectus robustus (Pursh) M.T. Strong [GU SU FG?] 

Scleria acanthocarpa Boeck. [FG] 

Scleria bracteata Cav. [GU SU FG] 

Scleria cerradicola T. Koyama [GU] 

Scleria cyperina Willd. ex Kunth [GU SU FG] 

Scleria distans Poir. [FG] 

Scleria eggersiana Boeck. [GU SU] 

Scleria flagellum-nigrorum Berg. [GU SU] 

Scleria grandis Core [GU?] 

Scleria hirtella Swartz [GU SU FG] 

Scleria interrupta L.C. Rich. [GU FG] 

Scleria lacustris Wright [GU SU] 

Scleria latifolia Swartz [GU SU FG] 

Scleria lindleyana C.B. Clarke [FG?] 

Scleria lithosperma (L.) Swartz [FG] 

Scleria longigluma Kükenth. [GU] 

Scleria macrogyne C.B. Clarke [GU SU FG] 

Scleria macrophylla Presl [GU SU FG] 

Scleria martii (Nees) Steud. [SU FG] 

Scleria melaleuca Rchb. ex Cham. & Schltdl. [GU SU FG] 

Scleria microcarpa Nees [GU SU FG] 

Scleria micrococca (Liebm.) Steud. [GU SU FG] 

Scleria mitis Berg. [GU SU FG] 

Scleria nutans Willd. ex Kunth [GU?] 

Scleria pinetorum Britton [SU] 

Scleria pterota Presl var. melanocarpa Nees [SU] [=Scleria 

melaleuca?] 

Scleria scabra Willd. [GU SU] 

Scleria secans (L.) Urb. [GU SU FG] 

Scleria setacea Poir. [GU SU FG] 

Scleria splitgerberiana Henrard ex Uittien [SU] 

Scleria staheliana Uittien [SU FG] 

Scleria stipitata Uittien [GU SU] 

Scleria stipularis Nees [GU SU FG] 

Scleria tepuiensis Core [GU FG] 

Scleria triquetra M.T. Strong [FG] 

Scleria vaginata or cf. vaginata Steud. [SU] 

Scleria verticillata Muhl. ex Willd. [GU SU FG?] 

Scleria violacea Pilger [FG] 

Trilepis kanukuensis Gilly [GU SU FG] 

Websteria confervoides (Poir.) S. Hooper [GU SU? FG] 

CYPERACEAE - SYNONYMS AND 



93

94 


Becquerelia divaricata (Nees ex Boeck.) Palla =Becquerelia 

cymosa subsp. merkeliana 

Bisboeckelera angustifolia (Boeck.) Kuntze =Bisboeckelera 

longifolia 

Bolboschoenus maritimus (L.) Palla =Schoenoplectus 

maritimus 

Bolboschoenus robustus (Pursh) Soják =Schoenoplectus 

robustus 

Bulbostylis capillaris (L.) C.B. Clarke var. glabra Uittien 

=Bulbostylis truncata 

Bulbostylis capillaris (L.) C.B. Clarke var. tenuifolia (Rudge) 

C.B. Clarke =Bulbostylis tenuifolia 

Bulbostylis schomburgkiana (Steud.) M.T. Strong [type 

erroneously reported from Guyana] 

Bulbostylis spadicea (Kunth) Kükenth. =Bulbostylis paradoxa 

Calyptrocarya fragifera Nees ex Kunth [nomen nudum] 

=Calyptrocarya glomerulata 

Calyptrocarya intermedia C.B. Clarke =Calyptrocarya 

glomerulata 

Cyperus brevifolius (Rottb.) Hassk. =Kyllinga brevifolia 

Cyperus cayennensis Lam. =Cyperus aggregatus 

Cyperus chalaranthus Presl =Cyperus laxus s.l. 

Cyperus densicaespitosus Mattf. & Kükenth. =Kyllinga 

pumila 

Cyperus diffusus Vahl =Cyperus laxus s.l. 

Cyperus elegans Auct. non L. =Cyperus laxus s.l. 

Cyperus flavus (Vahl) Nees =Cyperus aggregatus 

Cyperus globulosus Aubl. =Cyperus luzulae 

Cyperus globulosus Auctt., non Aubl. =Cyperus croceus 

Cyperus huarmensis (Kunth) M.C. Johnston =Cyperus 

aggregatus 

Cyperus kyllingia Endl. =Kyllinga nemoralis 

Cyperus obtusatus (Presl) Mattf. & Kükenth. =Kyllinga 

pungens 

Cyperus pedunculatus (R. Br.) Kern =Remirea maritima 

Cyperus ruderalis Uittien =Cyperus trailii 

Cyperus sesquiflorus (Torrey) Mattf. & Kükenth. =Kyllinga 

odorata 

Cyperus tolucensis Kunth =Cyperus laxus s.l. 

Cyperus uncinatus Poir. [not in Guianas] 

Dichromena ciliata Vahl =Rhynchospora nervosa var. ciliata 

Dichromena monostachya (Boeck.) C.B. Clarke 

=Rhynchospora albescens 

Dichromena montana Uittien =Rhynchospora montana 

Dichromena pubera Vahl =Rhynchospora pubera 

Dichromena radicans Schlechtend. & Cham. =Rhynchospora 

radicans 

Dichromena repens Vahl =Rhynchospora reptans 

Diplacrum laevissimum Kükenth. =Diplacrum guianense 

Diplacrum longifolium (Griseb.) C.B. Clarke ex Durand & 

Schinz =Diplacrum capitatum 

Diplacrum notopterum (Nees ex Steud.) C.B. Clarke 

=Diplacrum guianense 

Eleocharis atropurpurea (Retz.) Presl [not in Guianas] 

Eleocharis caribaea (Rottb.) S.F. Blake =Eleocharis geniculata 

Eleocharis fistulosa (Poir.) Link =Eleocharis acutangula 

Eleocharis sagotii C.B. Clarke =Eleocharis jelskiana 

Eleocharis sagotii C.B. Clarke var. glochidiata C.B. Clarke 

=Eleocharis plicarhachis 

Eleocharis sulcata Nees =Eleocharis filiculmis 

Fimbristylis annua (Allioni) Roem. & Schult. var. diphylla 

(Retz.) Kükenth. =Fimbristylis dichotoma 

Fimbristylis cymosa R. Brown subsp. spathacea (Roth) T. 

Koyama =Fimbristylis cymosa 


Fimbristylis littoralis Guadichaud [= miliacea by some] 

Fimbristylis spathacea Roth =Fimbristylis cymosa 

Hemicarpha micrantha (Vahl) Pax =Lipocarpha micrantha 

Kyllinga monocephala Rottb., nom. illeg. =Kyllinga nemoralis 

Kyllinga peruviana Lam. =Kyllinga vaginata 

Kyllinga rigida Baldwin =Kyllinga vaginata 

Kyllinga sesquiflora Torrey =Kyllinga odorata 

Lagenocarpus amazonicus (C.B. Clarke) Pfeiff. =Exochogyne 

amazonica 

Lagenocarpus kunthii (Miq.) Uittien =Lagenocarpus 

verticillatus 

Lagenocarpus tremulus Nees =Lagenocarpus rigidus subsp. 

tremulus 

Lagenocarpus weigelti (Spreng.) Uittien =Lagenocarpus 

verticillatus 

Lipocarpha sellowiana Kunth =Lipocarpha humboldtiana 

Mariscus cayennensis (Lam.) Urb. =Cyperus aggregatus 

Mariscus chlorocephalus C.B. Clarke =Cyperus 

chlorocephalus 

Mariscus elatus Kunth =Cyperus aggregatus 

Mariscus flavus Vahl =Cyperus aggregatus 

Mariscus gayi C.B. Clarke =Cyperus gayi 

Mariscus guianensis C.B. Clarke =Cyperus guianensis 

Mariscus ligularis (L.) Urb. =Cyperus ligularis 

Mariscus mutisii Kunth =Cyperus mutisii 

Mariscus pedunculatus (R. Br.) T. Koyama =Remirea maritima 

Mariscus tovarii Kunth =Cyperus hermaphroditus 

Mariscus umbellatus (Rottb.) Vahl =Cyperus subumbellatus 

Pycreus capillifolius (A. Rich.) C.B. Clarke =Cyperus 

capillifolius 

Pycreus flavescens (L.) Rchb. =Cyperus flavescens 

Pycreus megapotamicus (Kunth) Nees =Cyperus 

megapotamicus 

Pycreus polystachyos (Rottb.) Beauv. =Cyperus polystachyos 

Pycreus pumilus (L.) Nees =Cyperus pumilus 

Pycreus unioloides (R. Brown) Urb. =Cyperus unioloides 

Rhynchospora aurea Vahl =Rhynchospora corymbosa 

Rhynchospora cyperoides (Swartz) Mart. =Rhynchospora 

holoschoenoides 

Rhynchospora denticulata Huber =Rhynchospora subplumosa 

Rhynchospora distans Vahl =Rhynchospora fascicularis var. 

distans 

Rhynchospora glauca Vahl =Rhynchospora rugosa 

Rhynchospora hispidula (Vahl) Boeck., nom. illeg. 

=Rhynchospora trichodes 

Rhynchospora longispiculata Boeck. =Rhynchospora 

filiformis 

Rhynchospora monostachya Boeck. =Rhynchospora albescens


Rhynchospora nervosa (Vahl) Boeck. var. ciliata (Vahl) 

Kükenth. =Rhynchospora nervosa subsp. ciliata 

Rhynchospora pilosa (Kunth) Boeck. subsp. arenicola 

(Uittien) T. Koyama =Rhynchospora arenicola 

Rhynchospora podosperma C. Wright =Rhynchospora 

filiformis 

Rhynchospora repens (Vahl) Boeckeler =Rhynchospora 

reptans 

Rhynchospora rufa (Nees) Boeck. =Rhynchospora velutina 

Rhynchospora setacea (Berg.) Boeck. =Rhynchospora 

tenerrima 

Rhynchospora stellata (Lam.) Griseb. =Rhynchospora colorata 


Rhynchospora tenerrima Nees ex Spreng. subsp. fallax 

(Uittien) T. Koyama =Rhynchospora fallax 

Rhynchospora tenuis Willd. ex Link subsp. riparia (Nees) T. 

Koyama =Rhynchospora riparia 

Rhynchospora viridilutea C.B. Clarke =Rhynchospora hassleri 

Scirpus cubensis Poepp. & Kunth =Oxycaryum cubense 

Scirpus maritimus L. =Schoenoplectus maritimus 

Scirpus micranthus Vahl =Lipocarpha micrantha 

Scirpus robustus Pursh =Schoenoplectus robustus 

Scirpus submersus C. Wright =Websteria confervoides 

Scleria arundinacea Kunth =Scleria latifolia 

Scleria foliosa C. Wright, nom. illeg. =Scleria microcarpa 

Scleria latifolia Nees =Scleria arundinacea [=Scleria latifolia 

Swartz] 

Scleria liebmannii Steud. =Scleria micrococca 

Scleria paludosa Kunth =Scleria macrophylla 

Scleria pterota Presl [nomen nudum] =Scleria melaleuca 

Scleria pyramidalis Hochst. ex Steud. =Scleria martii [=Scleria 

stipularis?] 

Scleria reflexa Kunth =Scleria secans 

Scleria tenella Kunth =Scleria verticillata 

Scleria weigeltiana Schrad. ex Boeck. =Scleria secans 

Torulinium ferax (L.C. Rich.) Urb. =Cyperus odoratus 

Torulinium odoratum (L.) S. Hooper =Cyperus odoratus 

REFERENCES: 

CYRILLACEAE 

Thomas, J.L. 1960. A monographic study of the 

Cyrillaceae. Contr. Gray Herb. 186: 1-114. 

Cyrilla antillana Michx. [GU] [=Cyrilla racemiflora s.l.] 

Cyrilla brevifolia N.E. Brown [GU] [=Cyrilla racemiflora s.l.] 

Cyrilla perrottetii Briq. [GU] [=Cyrilla racemiflora s.l.] 

Cyrilla racemiflora L. [GU SU FG] 

DICHAPETALACEAE 


REFERENCES: 

Prance, G.T. 1972. Dichapetalaceae. Flora 


Prance, G.T. 1979. A new species of 

Dichapetalum from Suriname. Bull. Torrey 

Bot. Club 106(4): 309-312. 

Prance, G.T. 1983. Additions to Neotropical 

Dichapetalaceae. Brittonia 35(1): 49-54. 

Stafleu, F.A. 1951. Dichapetalaceae. In A. 


Dichapetalum pedunculatum (DC.) Baill. [GU SU FG] 

Dichapetalum rugosum (Vahl) Prance [GU SU FG] 

Dichapetalum schulzii Prance [SU] 

Tapura amazonica Poepp. & Endl. [SU FG] 

Tapura capitulifera Baill. [GU SU FG] 

Tapura guianensis Aubl. [GU SU FG] 

Tapura singularis Ducke [FG] 

DICHAPETALACEAE - SYNONYMS AND 


Chailletia flavicans Klotzsch =Dichapetalum rugosum 

Chailletia pedunculata DC. =Dichapetalum pedunculatum 

Chailletia sessiliflora DC. =Tapura guianensis 

Chailletia vestita Benth. =Dichapetalum rugosum 

Dichapetalum flavicans (Klotzsch) Engl. =Dichapetalum 

rugosum 

Dichapetalum glabrum (Vahl) Prance =Dichapetalum 

pedunculatum 

Dichapetalum scandens (Poir.) Johnston =Dichapetalum 

rugosum 

Dichapetalum vestitum (Benth.) Baill. =Dichapetalum 

rugosum 

Rohria petioliflora Willd. =Tapura guianensis 

Rohria schreberi J.F. Gmel. =Tapura guianensis 

Rohria tapura J.F. Gmel. =Tapura guianensis 

Symphyllanthus glaber Vahl =Dichapetalum pedunculatum 

Symphyllanthus rugosus Vahl =Dichapetalum rugosum 

DILLENIACEAE 

contributed by G. Aymard & C.L. Kelloff (1992); 

additions and corrections by G. Aymard & 

reviewed by C.L. Kelloff (1996) 

REFERENCES: 

Aymard, G. & J. Miller. 1994. Dilleniaceae 

novae Neotropicae III. Sinopsis y adiciones a 

las Dilleniaceae del Perú. Candollea 49(1): 

169-182. 

Kubitzki, K. 1970. Die gattung Tetracera. Mitt. 

Bot. Münch. 8: 1-98. 

Kubitzki, K. 1971. Doliocarpus, Davilla und 

verwandte gattungen. Mitt. Bot. Münch. 9: 1- 

105. 


Dilleniaceae. In A. Pulle, Flora of Suriname 

3(1): 386-408. 

SPERMATOPHYTA: Cyperaceae - Dilleniaceae 

95

96 


Curatella americana L. [GU SU FG] 

Davilla alata (Venten.) Briq. [GU SU FG] 

Davilla kunthii A. St. Hil. [GU SU FG] 

Davilla lacunosa Mart. [FG] 

Davilla nitida (Vahl) Kubitzki [GU SU FG] 

Davilla rugosa Poir. var. rugosa [GU SU FG] 

Davilla steyermarkii Kubitzki [to be expected in GU] 

Doliocarpus brevipedicellatus Garcke subsp. brevipedicellatus 

[GU SU FG] 

Doliocarpus dentatus (Aubl.) Standl. subsp. dentatus [GU SU 

FG] 

Doliocarpus dentatus (Aubl.) Standl. subsp. esmeraldae 

(Steyerm.) Kubitzki [GU SU FG] 

Doliocarpus gracilis Kubitzki [SU FG] 

Doliocarpus guianensis (Aubl.) Gilg. [GU SU FG] 

Doliocarpus macrocarpus Mart. ex Eichl. [GU SU FG] 

Doliocarpus major J.F. Gmelin subsp. major [GU SU FG] 

Doliocarpus paraensis Sleumer [GU SU FG] 

Doliocarpus sagotianus Kubitzki [FG] 

Doliocarpus savannarum Sandw. [GU SU FG] 

Doliocarpus spraguei Cheesem. [GU SU FG] 

Pinzona coriacea Mart. & Zucc. [GU SU FG] 

Tetracera asperula Miq. [GU SU FG] 

Tetracera costata Mart. ex Eichl. subsp. costata [GU SU FG] 

Tetracera costata Mart. ex Eichl. subsp. rotundifolia (J.E. 

Smith) Kubitzki [GU FG] 

Tetracera surinamensis Miq. [GU SU FG] 

Tetracera tigarea DC. [GU SU FG] 

Tetracera volubilis L. subsp. volubilis [GU SU FG] 

Tetracera willdenowiana Steud. subsp. emarginata Kubitzki 

[FG] 

Tetracera willdenowiana Steud. subsp. willdenowiana [SU] 

DILLENIACEAE - SYNONYMS AND 


Calinea scandens Aubl. =Doliocarpus spraguei 

Davilla aspera (Aubl.) Benoist =Davilla kunthii 

Davilla multiflora (DC.) St. Hil. =Davilla nitida 

Doliocarpus calinea J.F. Gmelin =Doliocarpus spraguei 

Doliocarpus coriaceus (Mart. & Zucc.) Gilg =Pinzona coriacea 

Doliocarpus rolandri J.F. Gmelin =Doliocarpus major 

Doliocarpus scandens Poir. =Doliocarpus major 

Doliocarpus surinamensis Lanj. =Doliocarpus paraensis 

Doliocarpus virgatus Sagot =Doliocarpus macrocarpus 

Soramia guianensis Aubl. =Doliocarpus guianensis 

Tetracera ovalifolia DC. =Tetracera asperula 

Tetracera rotundifolia Smith =Tetracera asperula 

Tigarea aspera Aubl. =Tetracera tigarea 

Tigarea dentata Aubl. =Doliocarpus dentatus 

REFERENCES: 

DIOSCOREACEAE 

Gleason, H.A. 1925. Studies on the flora of 

northern South America. V. Miscellaneous 

new or noteworthy species. Bull. Torrey Bot. 

Club 52(5): 181-196. 

Knuth, R. 1924. Dioscoreaceae. In A. Engler, 

Das Pflanzenreich IV(43): 1-387. 

Dioscorea alata L. [GU SU FG] [introduced] 

SPERMATOPHYTA: Dilleniaceae - Dipterocarpaceae 

Dioscorea amazonum Mart. ex Griseb. [GU SU FG] 

Dioscorea bulbifera L. [SU FG] [introduced] 

Dioscorea cayennensis Lam. [GU SU FG] [introduced?] 

Dioscorea cordifolia Laness. 

Dioscorea crotalariifolia Uline [GU] 

Dioscorea dodecaneura Vell. [GU SU FG] 

Dioscorea esculenta (Lour.) Prain [FG] [cultivated only?] 

Dioscorea georgensis Knuth [GU] [=Dioscorea amazonum?] 

Dioscorea guianensis Knuth [GU] 

Dioscorea hastata Vell. [FG] [=Dioscorea hasslerana?] 

Dioscorea hitchcockii Knuth [GU] 

Dioscorea lanosa Gleason [GU SU] 

Dioscorea macrostachya Benth. [GU?] 

Dioscorea manicorensis Clayton, sp. nov. ined. [SU FG] 

Dioscorea megacarpa Gleason [GU SU FG] 

Dioscorea melastomatifolia Uline ex Prain & Burkill [FG] 

[=Dioscorea truncata?] 

Dioscorea microura Knuth [GU] 

Dioscorea oblonga Gleason [GU SU] 

Dioscorea pilosiuscula Bert. ex Spreng. [GU SU FG] 

Dioscorea piperifolia Humb. & Bonpl. ex Willd. [GU] 

Dioscorea polygonoides Humb. & Bonpl. ex Willd. [GU SU 

FG] 

Dioscorea pomeroonensis Knuth [GU] 

Dioscorea potarensis Knuth [GU] 

Dioscorea pubescens Poir. [FG] [insufficiently known species] 

Dioscorea quinquefoliolata Knuth [GU] 

Dioscorea riparia Kunth & Schomb. [GU SU] [=Dioscorea 

samydea?] 

Dioscorea sagittata Poir. [FG] [insufficiently known species] 

Dioscorea samydea Mart. ex Griseb. [GU SU] 

Dioscorea syringifolia Kunth & Schomb. [GU FG] 

Dioscorea trachyandra Griseb. [GU] 

Dioscorea trachycarpa Kunth [GU] [insufficiently known 

species] 

Dioscorea trichanthera Gleason [GU] 

Dioscorea trifida L.f. [GU SU FG] 

Dioscorea truncata Miq. [SU] [=Dioscorea trifida?] 

DIOSCOREACEAE - SYNONYMS AND 


Dioscorea affinis Kunth =Dioscorea trifida 

Dioscorea bulbifera Griseb., non L. =Dioscorea pilosiuscula 

Dioscorea cuspidata Klotzsch ex Kunth =Dioscorea 

amazonum 

Dioscorea elegantula Kunth =Dioscorea amazonum 

Dioscorea kegeliana Griseb. =Dioscorea polygonoides 

Dioscorea megalobotrya Kunth & Schomb. ex Kunth 

=Dioscorea amazonum 

Dioscorea schomburgkiana Kunth ex Schomb. =Dioscorea 

pilosiuscula 

Dioscorea surinamensis Mey. =Dioscorea amazonum 

Dioscorea truncata Schomb., non Miq. =Dioscorea megacarpa 

Dioscorea truncata Prain, non Miq. =Dioscorea megacarpa 

REFERENCES: 

DIPTEROCARPACEAE 

Maguire, B. & P.S. Ashton. 1977. 

Pakaraimoideae, Dipterocarpaceae of the 

Western Hemisphere. II. Systematic,


geographic, and phyletic considerations. 

Taxon 26(4): 343-368. 

Maguire, B. & P.S. Ashton. 1980. Pakaraimaea 

dipterocarpacea II. Taxon 29(2/3): 225-231. 

Maguire, B. & J.A. Steyermark. 1981. 

Pakaraimaea dipterocarpacea - III. In B. 



Bot. Gard. 32: 1-391. 

Pakaraimaea dipterocarpacea Maguire & Ashton subsp. 

dipterocarpacea [GU] 

DROSERACEAE 

contributed C.L. Kelloff (1992); reviewed by 

C.L. Kelloff (1996) 

REFERENCES: 

Brummer-Dinger, C.A. 1955. Notes on Guiana 

Droseraceae. Acta Bot. Neerl. 4(1): 136-138. 

Maguire, B. & J.J. Wurdack. 1957. Droseraceae. 

In B. Maguire, J.J. Wurdack, & collaborators, 



Drosera capillaris Poir. [GU SU] 

Drosera cayennensis Sagot ex Diels [GU FG] 

Drosera esmeraldae (Steyerm.) Maguire & Wurdack [GU] 

Drosera intermedia Hayne [GU SU] 

Drosera kaieteurensis Brummer-Dinger [GU] 

Drosera pusilla Kunth [GU SU] 

Drosera roraimae (Klotzsch ex Diels) Maguire & Laundon 

[GU] 

Drosera sessilifolia St. Hil. [GU] 

Drosera tenella Willd. ex Roem. & Schult. [FG] [=Drosera 

capillaris?] 

REFERENCES: 

EBENACEAE 

Cavalcante, P.B. 1963. Contribuçao ao 

conhecimento do gênero Diospyros Dalech. 

na Amazônia. Bol. Mus. Goeldi n.s. 20: 1-53. 

White, F. 1981. Ebenaceae. In B. Maguire & 


Highland part 11. Mem. New York Bot. 

Gard. 32: 1-391. 

Diospyros artantifolia Mart. ex Miq. [FG] 

Diospyros carbonaria Benoist [FG] 

Diospyros cauliflora Mart. [FG] 

Diospyros cayennensis A. DC. [FG] 

Diospyros dichroa Sandw. [GU SU FG] 

Diospyros discolor Willd. [FG] [cultivated; naturalized?] 

Diospyros duckei Sandw. [FG?] 

Diospyros glomerata Spruce [FG] 

Diospyros guianensis (Aubl.) Guerke subsp. akaraiensis (A.C. 

Smith) F. White [GU?] 

Diospyros guianensis (Aubl.) Guerke subsp. guianensis [GU 

SU FG] 

Diospyros ierensis Britton [GU SU? FG] 

Diospyros lissocarpoides Sandw. [GU SU FG] 

Diospyros martinii Benoist ex Amshoff [SU FG] 

Diospyros matheriana A.C. Smith [GU SU FG] 

Diospyros melinonii (Hiern) A.C. Smith [GU SU FG] 

Diospyros praetermissa Sandw. [perhaps to be expected in FG] 

Diospyros santaremnensis Sandw. [perhaps to be expected in 

FG] 

Diospyros sericea A. DC. [GU] 

Diospyros tetrandra Hiern [SU FG] 

Diospyros vestita Benoist [FG] 

EBENACEAE - SYNONYMS AND 


Diospyros akaraiensis A.C. Smith =Diospyros guianensis 

subsp. akaraiensis 

Diospyros paralea Steud. =Diospyros guianensis 

Diospyros subrotata Hiern =Diospyros artanthifolia 

Paralea guianensis Aubl. =Diospyros guianensis 

Ropourea guianensis Aubl. =Diospyros martinii 

ELAEOCARPACEAE 

reviewed by L.J. Dorr (1992) 

REFERENCES: 

Jansen-Jacobs, M.J. 1986. Elaeocarpaceae 

(additions and corrections). In A.L. Stoffers 

& J.C. Lindeman, Flora of Suriname 3(1-2): 

301-305. 

Smith, C.E., Jr. 1954. The New World species 

of Sloanea. Contr. Gray Herb. 175: 1-114. 

Uittien, H. 1932. Elaeocarpaceae. In A. Pulle, 


Sloanea acutiflora Uittien [SU] [=Sloanea laxiflora?] 

Sloanea brevipes Benth. [GU SU? FG] 

Sloanea durissima Spruce ex Benth. [GU] 

Sloanea echinocarpa Uittien [GU SU FG] 

Sloanea eichleri K. Schum. [GU SU FG] 

Sloanea garckeana K. Schum. [GU SU FG] 

Sloanea gracilis Uittien [SU] 

Sloanea grandiflora J.E. Smith [GU SU FG] 

SPERMATOPHYTA: Dipterocarpaceae - Elaeocarpaceae 

97

98 


Sloanea guianensis (Aubl.) Benth. [GU SU FG] 

Sloanea latifolia (Rich.) K. Schum. [GU FG] 

Sloanea laurifolia (Benth.) Benth. [GU FG?] 

Sloanea macrophylla Benth. ex Turcz. [FG?] 

Sloanea nitida G. Don [GU FG] 

Sloanea obtusifolia (Moric.) K. Schum. [FG?] 

Sloanea parviflora Planch. ex Benth. [GU SU FG] 

Sloanea picapica Standl. var. hirsuta C.E. Smith [GU] 

Sloanea pubescens (Poepp. & Endl.) Benth. [SU? FG] 

Sloanea robusta Uittien [GU SU] 

Sloanea rufa Planch. ex Benth. [SU FG] 

Sloanea schomburgkii Benth. [GU FG] 

Sloanea sinemariensis Aubl. [SU? FG] 

Sloanea synandra Spruce ex Benth. [GU FG] 

Sloanea terniflora (Moçino & Sessé ex DC.) Standl. [SU FG] 

Sloanea trichosticha R.O. Williams & Sandw. [GU] 

Sloanea tuerckheimii J.D. Smith [GU SU FG] 

ELAEOCARPACEAE - SYNONYMS AND 


Ablania guianensis Aubl. =Sloanea guianensis 

Courimari guianensis Aubl. =Sloanea sp.?[name of uncertain 

application] 

Sloanea amplifrons Johnston =Sloanea grandiflora 

Sloanea brachytepala Ducke =Sloanea laurifolia 

Sloanea bracteosa Ducke =Sloanea nitida 

Sloanea corymbiflora DC. =Sloanea latifolia 

Sloanea dentata L. [not in Guianas] 

Sloanea obtusa (Splitg.) K. Schum. =Sloanea parviflora 

Sloanea usurpatrix Sprague & Riley =Sloanea tuerckheimii 

EREMOLEPIDACEAE 

contributed by J. Kuijt (1992) 

REFERENCES: 

Kuijt, J. 1988. Monograph of the 

Eremolepidaceae. Syst. Bot. Monogr. 18: 1- 

60. 

Antidaphne viscoidea Poepp. & Endl. [GU] 

ERICACEAE 

contributed by J.L. Luteyn (1992) 

REFERENCES: 

Lanjouw, J. 1934. Ericaceae. In A. Pulle, Flora 


Lanjouw, J. 1937. Ericaceae (additions and 


4(1): 492-493. 

Luteyn, J.L. 1983. Ericaceae Part I. 

Cavendishia. Flora Neotrop. Monogr. 35: 1- 

290. 

SPERMATOPHYTA: Elaeocarpaceae - Ericaceae 

Luteyn, J.L. 1987. Orthaea: new species and 

redefinition of the genus. Nord. J. Bot. 7(1): 

31-37. 

Luteyn, J.L. (ed.). 1995. Ericaceae - Part II. The 

Superior-Ovaried Genera (Monotropoideae, 

Pyroloideae, Rhododendroideae, and 

Vaccinioideae p.p.). Flora Neotropica 

Monogr. 66: 1-560. 

Maguire, B., J.A. Steyermark, & J.L. Luteyn. 

1978. Ericaceae. In B. Maguire & 



139-203. 

Sleumer, H.O. 1936. Die Arten der Gattung 

Vaccinium L. in Zentral- und Südamerika. 

Notizbl. Bot. Gart. Berlin-Dahlem 13: 111- 

140. 

Smith, A.C. 1932. The American species of 

Thibaudieae. Contr. U.S. Nat. Herb. 28: 311- 

547. 

Smith, A.C. 1933. The genera Sphyrospermum 

and Disterigma. Brittonia 1: 203-232. 

Agarista duckei (Huber) Judd [to be expected in GU] 

Befaria [Bejaria] imthurnii N.E. Brown [GU?] [Mt. Roraima] 

Befaria [Bejaria] sprucei Meissn. [GU] 

Cavendishia callista J.D. Smith [GU SU FG] 

Ceratostema glandulifera Maguire, Steyerm. & Luteyn [GU] 

Disterigma humboldtii (Klotzsch) Nied. [GU] 

Gaultheria erecta Ventenat [GU] [Mt. Roraima] 

Gaultheria setulosa N.E. Brown [GU?] [Mt. Roraima] 

Ledothamnus guyanensis Meissn. [GU?] [Mt. Roraima] 

Ledothamnus sessiliflorus N.E. Brown [GU?] [Mt. Roraima] 

Notopora schomburgkii J.D. Hook. [GU] 

Orthaea apophysata (Griseb.) A.C. Smith [GU] 

Orthaea hispida A.C. Smith [GU] 

Orthaea merumensis Maguire, Steyerm. & Luteyn [GU] 

Orthaea venamensis Maguire, Steyerm. & Luteyn [GU] 

Pernettya marginata N.E. Brown [Mt. Roraima] 

Psammisia guianensis Klotzsch [GU] 

Psammisia urichiana (Britton) A.C. Smith [GU] 

Satyria carnosiflora Lanj. [GU SU] 

Satyria cerander (Dunal) A.C. Smith [SU? FG] 

Satyria panurensis (Benth.) Benth. & J.D. Hook. [GU FG] 

Sphyrospermum buxifolium Poepp. & Endl. [GU FG] 

Sphyrospermum cordifolium Benth. [GU FG] 

Thibaudia formosa (Klotzsch) Hoerold [GU] 

Thibaudia nutans Klotzsch ex Mansf. [GU SU] 

Vaccinium euryanthum A.C. Smith [GU] 

Vaccinium puberulum Klotzsch ex Meissn. s.l. [GU] 

Vaccinium roraimense N.E. Brown [Mt. Roraima] 

ERICACEAE - SYNONYMS AND 


Befaria aestuans Mutis ex L.f. [not in Guianas] 

Befaria glauca Bonpl. =Befaria aestuans [not in Guianas] 

Befaria guianensis Klotzsch ex N.E. Brown =Befaria sprucei 

Befaria guianensis Klotzsch ex Schomb. [nomen nudum] 

=Befaria sprucei 

Befaria puberula Klotzsch ex Mansfeld & Sleumer =Befaria 

sprucei


Befaria schomburgkiana Klotzsch ex Mansfeld & Sleumer 


Befaria schomburgkiana Klotzsch ex Schomb. [nomen nudum] 


Befaria tatei Gleason =Befaria imthurnii 

Brossea roraimae (Klotzsch ex Meissn.) O. Kuntze 

=Gaultheria erecta 

Cavendishia duidae A.C. Smith =Cavendishia callista 

Cavendishia salicifolia Maguire, Steyerm. & Luteyn 

=Cavendishia callista 

Epigaea cordifolia Swartz =Gaultheria domingensis [not in 


Gaultheria odorata Bredemeyer ex Willd. =Gaultheria erecta 

Gaultheria roraimae Klotzsch ex Meissn. =Gaultheria erecta 

Orthaea pubifolia Maguire, Steyerm. & Luteyn =Orthaea 

hispida 

Psammisia coriacea N.E. Brown =Thibaudia nutans 

Satyria maguirei Camp =Satyria carnosiflora 

Sphyrospermum majus Griseb. =Sphyrospermum cordifolium 

Thibaudia roraimae Mansf. =Thibaudia formosa 

Thibaudia ulei (Mansf.) A.C. Smith =Thibaudia nutans 

Vaccinium subcrenulatum Klotzsch ex Meissn. =Vaccinium 

puberulum s.l. 

ERIOCAULACEAE 

contributed by M. Hakki (1992); additions and 

corrections by M. Hakki (1996) 

REFERENCES: 

Giulietti, A.M., M.C.E. Amaral, & V. Bittrich. 

1995. Phylogenetic analysis of inter- and 

infrageneric relationships of Leiothrix 

Ruhland (Eriocaulaceae). Kew Bull. 50(1): 

55-71. 

Giulietti, A.M. & N. Hensold. 1991. 

Nomenclatural changes and range extension 

in Leiothrix flavescens (Bong.) Ruhl. 

(Eriocaulaceae). Novon 1(1): 45-49. 

Giulietti, A.M. & N. Hensold. 1991. 

Synonymization of the genera Comanthera 

and Carptotepala with Syngonanthus 

(Eriocaulaceae). Ann. Missouri Bot. Gard. 78 

(2): 460-464. 

Hensold, N. 1991. Revisionary studies in the 

Eriocaulaceae of Venezuela. Ann. Missouri 

Bot. Gard. 78(2): 424-440. 

Hensold, N. & A.M. Giulietti. 1991. Revision 

and redefinition of the genus Rondonanthus 

Herzog (Eriocaulaceae). Ann. Missouri Bot. 

Gard. 78(2): 441-459. 


Eriocaulaceae (additions and corrections). In 


1(2): 330-339. 

Moldenke, H.N. 1971. A Fifth Summary of the 

Verbenaceae, Avicenniaceae, Stilbaceae, 

Dicrostylidaceae, Symphoremaceae, 

Nyctanthaceae, and Eriocaulaceae of the 

World. H.N. Moldenke, Wayne, NJ. 

Uittien, H. & A.N.J. Heyn. 1938. Eriocaulaceae. 

In A. Pulle, Flora of Suriname 1(1): 213-224. 

Eriocaulon guyanense Körn. [GU FG] 

Eriocaulon heterodoxum Moldenke [GU] 

Eriocaulon humboldtii Kunth [GU SU] 

Eriocaulon melanocephalum Kunth [GU SU FG] 

Eriocaulon steyermarkii Moldenke [GU SU] 

Eriocaulon stramineum Körn. [FG?] 

Eriocaulon tenuifolium Klotzsch ex Körn. [GU] 

Leiothrix flavescens (Bong.) Ruhland [GU] 

Paepalanthus bifidus (Schrad.) Kunth [GU SU FG] 

Paepalanthus convexus Gleason [GU] 

Paepalanthus dichotomus Klotzsch ex Körn. [GU] 

Paepalanthus fasciculatus (Roettb.) Körn. [GU SU FG] 

Paepalanthus filipes Moldenke [GU] 

Paepalanthus fraternus N.E. Brown [GU] 

Paepalanthus gleasonii Moldenke [GU] 

Paepalanthus lamarckii Kunth [GU SU FG] 

Paepalanthus oyapockensis Herzog [GU SU FG] 

Paepalanthus pauper Moldenke [GU] 

Paepalanthus perplexans Moldenke [GU] 

Paepalanthus polytrichoides Kunth [GU SU] 

Paepalanthus roraimensis Moldenke [GU] 

Paepalanthus schomburgkii Klotzsch ex Körn. [GU] 

Paepalanthus sessiliflorus Mart. var. venezuelensis Moldenke 

[GU] 

Paepalanthus subcaulescens N.E. Brown [GU] 

Paepalanthus subtilis Miq. [GU SU] 

Paepalanthus tortilis (Bong.) Körn. [GU SU] 

Philodice hoffmannseggii Mart. [GU SU] 

Rondonanthus capillaceus (Klotzsch ex Körn.) Hensold & 

Giul. [GU] 

Rondonanthus roraimae (Oliver) Herzog [GU] 

Syngonanthus anomalus (Körn.) Ruhland [GU SU FG] 

Syngonanthus bisumbellatus (Steud.) Ruhland [GU] 

Syngonanthus caulescens (Poir.) Ruhland [GU SU FG] 

Syngonanthus fenestratus Hensold [GU] 

Syngonanthus gracilis (Bong.) Ruhland [GU SU FG] 

Syngonanthus heteropeploides Herzog [GU] 

Syngonanthus humboldtii (Kunth) Ruhland [GU SU] 

Syngonanthus jenmanii (Gleason) Giul. & Hensold [GU] 

Syngonanthus kegelianus (Körn.) Ruhland [GU SU] 

Syngonanthus leprieurii (Körn.) Ruhland [FG] 

Syngonanthus longipes Gleason [GU SU] 

Syngonanthus pakaraimensis Moldenke [GU] 

Syngonanthus simplex (Miq.) Ruhland [GU SU FG] 

Syngonanthus surinamensis Moldenke [SU] 

Syngonanthus umbellatus (Lam.) Ruhland [GU SU FG] 

Syngonanthus xeranthemoides (Bong.) Ruhland [GU] 

Tonina fluviatilis Aubl. [GU SU FG] 

ERIOCAULACEAE - SYNONYMS AND 


Carptotepala jenmanii (Gleason) Moldenke =Syngonanthus 

jenmanii 

Paepalanthus brunneus Moldenke =Paepalanthus 

oyapockensis 

Paepalanthus capillaceus Klotzsch ex Körn. =Rondonanthus 

capillaceus 

Paepalanthus congestus (Kunth) Kunth =Paepalanthus 

fasciculatus 

Paepalanthus griseus Moldenke =Paepalanthus oyapockensis 

SPERMATOPHYTA: Ericaceae - Eriocaulaceae 

99

100 


Paepalanthus leucocyaneus Tutin =Paepalanthus oyapockensis 

Paepalanthus maguirei Moldenke =Paepalanthus tortilis 

Paepalanthus tafelbergensis Moldenke =Paepalanthus 

oyapockensis 

Paepalanthus tatei Moldenke =Paepalanthus tortilis 

Paepalanthus viscosus Moldenke =Paepalanthus 

polytrichoides 

Syngonanthus biformis (N.E. Brown) Gleason =Syngonanthus 

simplex 

Syngonanthus glandulosus Gleason =Syngonanthus 

caulescens 

Syngonanthus guianensis Moldenke =Paepalanthus 

polytrichoides 

Syngonanthus heteropeplus (Körn.) Ruhland =Syngonanthus 

simplex 

Syngonanthus macrocaulon Ruhland =Syngonanthus 

anomalus 

Syngonanthus savannarum Moldenke =Paepalanthus 

dichotomus 

Syngonanthus tricostatus Gleason =Syngonanthus 

xeranthemoides 

ERYTHROXYLACEAE 


REFERENCES: 

Schulz, O.E. 1907. Erythroxylaceae. In A. 

Engler, Das Pflanzenreich IV(134): 1-176. 

Westhoff, V. 1942. Erythroxylaceae. In A. 


Erythroxylum amazonicum Peyr. [GU SU FG] 

Erythroxylum amplum Benth. [GU SU FG] 

Erythroxylum brevipes DC. [GU?] 

Erythroxylum campestre St. Hil. [GU?] 

Erythroxylum citrifolium St. Hil. [GU SU FG] 

Erythroxylum coelophlebium Mart. [SU] 

Erythroxylum cumanense Kunth [GU SU FG] 

Erythroxylum divaricatum Peyr. [GU] 

Erythroxylum fimbriatum Peyr. [FG] 

Erythroxylum gracilipes Peyr. [GU SU FG] 

Erythroxylum grisebachii Peyr. [SU] 

Erythroxylum impressum O.E. Schulz [GU] 

Erythroxylum kapplerianum Peyr. [GU SU FG] 

Erythroxylum leptoneurum O.E. Schulz [FG] 

Erythroxylum ligustrinum DC. [SU FG] 

Erythroxylum lineolatum DC. [GU SU FG] 

Erythroxylum macrophyllum Cav. [GU SU FG] 

Erythroxylum mucronatum Benth. [GU SU FG] 

Erythroxylum roraimae Klotzsch ex O.E. Schulz [GU SU FG] 

Erythroxylum rufum Cav. [GU FG] 

Erythroxylum schomburgkii Peyr. [GU] 

Erythroxylum squamatum Swartz [GU SU FG] 

Erythroxylum suberosum St. Hil. [GU FG] 

Erythroxylum vernicosum O.E. Schulz [GU FG?] 

ERYTHROXYLACEAE - SYNONYMS AND 


Erythroxylum coelophlebium Mart. var. grisebachii (Peyr.) 

O.E. Schulz =Erythroxylum grisebachii 

SPERMATOPHYTA: Eriocaulaceae - Euphorbiaceae 

Erythroxylum ectinocalyx Klotzsch =Erythroxylum 

divaricatum 

Erythroxylum orinocense Klotzsch =Erythroxylum 

schomburgkii 

Erythroxylum recurrens Huber =Erythroxylum gracilipes 

Erythroxylum squarrosum Klotzsch [nomen nudum] 

=Erythroxylum rufum 

Erythroxylum testaceum Peyr. =Erythroxylum suberosum 

Erythroxylum venezuelense Steyerm. =Erythroxylum 

mucronatum 

EUPHORBIACEAE 

contributed by L.J. Gillespie (1992) 

REFERENCES: 

Burger, W. & M. Huft. Rubiaceae. In W. 


(n.s.) 36: 1-169. 

Den Hollander, G. & C.C. Berg. 1986. Studies 

on the flora of the Guianas 21. Mabea species 

of the Guianas - a precursor. Proc. Kon. Ned. 

Akad. Wet. ser. C, 89(2): 147-157. 

Franco, P. 1990. The genus Hyeronima in South 

America. Bot. Jahrb. Syst. 111: 297-346. 

Gillespie, L.J. 1993. Euphorbiaceae of the 

Guianas: Annotated species checklist and key 

to the genera. Brittonia 45(1): 56-94. 

Gillespie, L.J. 1993. A synopsis of the 

Neotropical Plukenetia including two new 

species. Syst. Bot. 18(4): 575-592. 

Gillespie, L.J. 1994. A new section and two new 

species of Tragia from Venezuelan Guayana 

and French Guiana. Novon 4(4): 330-338. 

Gillespie, L.J. & W.S. Armbruster (in press). A 

contribution to the Guianan Flora: 

Dalechampia, Haematostemon, Omphalea, 

Pera, Plukenetia, and Tragia, with notes on 

subfamily Acalyphoideae. Smithsonian 

Contributions to Botany 86. 

Görts-van Rijn, A.R.A. 1976. Euphorbiaceae 



424. 

Hayden, W.J. 1990. Notes on Neotropical 

Amanoa. Brittonia 42(4): 260-270. 

Jablonski, E. 1965. Euphorbiaceae. In B. 


Guayana Highland Part 6. Mem. New York 

Bot. Gard. 12(3): 1-285. 

Jablonski, E. 1967. Euphorbiaceae. In B. 


Guayana Highland Part 7. Mem. New York 

Bot. Gard. 17(1): 1-439. 

Lanjouw, J. 1932. Euphorbiaceae. In A. Pulle, 



Lanjouw, J. 1939. Euphorbiaceae (additions and 


2(1): 457-470. 

Rogers, D.J. & S.G. Appan. 1973. Manihot and 

Manihotoides. Flora Neotropica Monogr. 13: 

1-272. 

Webster, G.L. & W.S. Armbruster. 1991. A 

synopsis of the neotropical species of 

Dalechampia. J. Linn. Soc. Bot. 105: 137- 

177. 

Acalypha arvensis Poepp. [SU FG] 

Acalypha diversifolia Jacq. [GU SU FG] 

Acalypha indica L. [GU FG] [naturalized] 

Acalypha lanceolata Willd. [GU] [naturalized] 

Acalypha macrostachya Jacq. [GU SU] 

Acalypha poirettii Spreng. [FG] 

Acalypha scandens Benth. [GU SU] 

Acalypha villosa Jacq. [GU] 

Actinostemon schomburgkii (Klotzsch) Hochr. [GU SU? FG] 

Adenophaedra grandifolia (Klotzsch) Muell. Arg. [GU FG] 

Alchornea schomburgkii Klotzsch [GU SU FG] 

Alchornea triplinervia (Spreng.) Muell. Arg. [GU SU FG] 

Alchorneopsis floribunda (Benth.) Muell. Arg. [GU SU FG] 

Amanoa congesta W.J. Hayden [FG] 

Amanoa guianensis Aubl. [GU SU FG] 

Amanoa neglecta W.J. Hayden [SU FG] 

Antidesma ghaesembilla Gaertn. [GU] [cultivated, naturalized] 

Aparisthmium cordatum (Adr. Juss.) Baill. [GU SU FG] 

Bernardia amazonica Croizat [FG] [specimens not seen] 

Bernardia sidoides (Klotzsch) Muell. Arg. [GU] 

Caperonia corchoroides Muell. Arg. [SU] [=Caperonia 

palustris?] 

Caperonia paludosa Klotzsch [GU FG] [=Caperonia 

castaneifolia (L.) A. St. Hil.?] 

Caperonia palustris (L.) A. St. Hil. [GU SU FG] 

Caperonia stenophylla Muell. Arg. [GU SU FG?] 

Chaetocarpus schomburgkianus (Kuntze) Pax & K. Hoffm. 

[GU SU FG] 

Chamaesyce adenoptera (Bertol.) Small [GU? FG] 

Chamaesyce hirta (L.) Millsp. [GU SU FG] 

Chamaesyce hypericifolia (L.) Millsp. [GU SU FG] 

Chamaesyce hyssopifolia (L.) Small [GU SU FG] 

Chamaesyce mesembryanthifolia (Jacq.) Dugand [FG?] 

Chamaesyce prostrata (Aiton) Small [GU SU FG] 

Chamaesyce serpens (Kunth) Small [GU FG] 

Chamaesyce thymifolia (L.) Millsp. [GU SU FG] 

Cnidoscolus urens (L.) J.C. Arthur [GU SU FG] 

Conceveiba guianensis Aubl. [GU SU FG] 

Conceveiba hostmannii Benth. [SU] 

Conceveiba martiana Baill. [FG] 

Croton argyrophylloides Muell. Arg. [FG?] 

Croton conduplicata Kunth [GU] [specimens not seen] 

Croton cuneatus Klotzsch [GU SU FG] 

Croton draconoides Muell. Arg. [FG] 

Croton essequiboensis Klotzsch [GU] [=Croton populifolius 

P. Mill.?] 

Croton galeopsifolius Lanj. [GU] 

Croton guianensis Aubl. [GU SU? FG] 

Croton hirtus L'Hér. [GU SU FG] 

Croton hostmannii Miq. [GU SU FG] 

Croton kaieteuri Jabl. [GU] [probably conspecific with Croton 

cuneatus] 

Croton kavayanensis Steyerm. [FG?] 

SPERMATOPHYTA: Euphorbiaceae 

101 

Croton lobatus L. [GU SU FG] 

Croton longiradiatus Lanj. [SU FG] 

Croton macradenis Görts & Punt [GU SU FG] 

Croton matourensis Aubl. [GU SU FG] 

Croton nervosus Klotzsch [GU] 

Croton nuntians Croizat [GU FG] 

Croton orinocensis Muell. Arg. [GU] 

Croton pakaraimae Jabl. [GU] 

Croton palanostigma Klotzsch [FG] 

Croton polypleurus Croizat [GU] 

Croton potaroensis Lanj. [GU] 

Croton pullei Lanj. var. glabrior Lanj. [SU FG] 

Croton pullei Lanj. var. pullei [GU? SU] 

Croton roraimensis Croizat var. roraimensis [GU] 

Croton rottlerifolius Baill. [FG?] 

Croton schiedeanus Schlechtend. [GU SU FG] 

Croton sipaliwiniensis Lanj. [SU] 

Croton stahelianus Lanj. [SU] 

Croton subincanus Muell. Arg. [GU SU FG] 

Croton tafelbergicus Croizat [SU FG] 

Croton trinitatis Millsp. [GU SU FG] 

Dalechampia affinis Muell. Arg. [GU SU FG] 

Dalechampia attenuistylus Armbruster [FG] 

Dalechampia brownsbergensis Webster & Armbruster [SU] 

Dalechampia dioscoreifolia Poepp. [FG] 

Dalechampia fragrans Armbruster [SU FG] 

Dalechampia magnoliifolia Muell. Arg. [GU FG] 

Dalechampia micrantha Poepp. [SU FG] 

Dalechampia parvibracteata Lanj. [GU SU FG] 

Dalechampia scandens L. [GU SU FG] 

Dalechampia tiliifolia Lam. [GU FG] 

Discocarpus essequiboensis Klotzsch [GU SU] 

Dodecastigma integrifolium (Lanj.) Lanj. & Sandw. [GU FG] 

Drypetes fanshawei Sandw. [GU] 

Drypetes variabilis Uittien [GU SU FG] 

Euphorbia comosa Vell. [SU?] 

Euphorbia cotinifolia L. [GU SU FG] 

Euphorbia cyathophora Murray [SU FG] [cultivated, 

naturalized] 

Euphorbia graminea Jacq. [FG] 

Euphorbia heterophylla L. [GU SU] [cultivated, naturalized] 

Euphorbia surinamensis Lanj. [GU SU] 

Gavarretia terminalis Baill. [GU] 

Glycydendron amazonicum Ducke [GU SU FG] 

Haematostemon guianensis Sandw. [GU] 

Hevea brasiliensis (Adr. Juss.) Muell. Arg. [FG?] [widely 

cultivated, possibly indigenous in FG] 

Hevea guianensis Aubl. [GU? FG] 

Hevea pauciflora (Spruce ex Benth.) Muell. Arg. [GU SU] 

Hura crepitans L. [GU SU FG] 

Hyeronima alchorneoides Allemao var. alchorneoides [GU 

SU FG] 

Hyeronima alchorneoides Allemao var. stipulosa Franco [FG] 

Hyeronima oblonga (Tul.) Muell. Arg. [GU FG] 

Jablonskia congesta (Benth. ex Muell. Arg.) Webster [GU SU 

FG?] 

Jatropha curcas L. [GU SU FG] [cultivated, naturalized] 

Jatropha gossypiifolia L. [GU SU FG] 

Mabea angularis den Hollander [GU] 

Mabea biglandulosa Muell. Arg. [GU] 

Mabea montana Muell. Arg. [GU] 

Mabea pakaraimaea Esser, sp. nov. ined. [GU] 

Mabea piriri Aubl. [GU SU FG] 

Mabea pulcherrima Muell. Arg. [GU SU FG] 

Mabea rubicunda Jabl. [GU] 

Mabea salicoides Esser [FG]

102 


Mabea speciosa Muell. Arg. subsp. A [GU SU FG] 

Mabea speciosa Muell. Arg. subsp. B [FG] 

Mabea speciosa Muell. Arg. subsp. speciosa [GU SU FG] 

Mabea subsessilis Pax & K. Hoffm. [GU SU FG] 

Mabea taquari Aubl. [GU SU FG] 

Manihot brachyloba Muell. Arg. [SU FG] 

Manihot esculenta Crantz [GU SU FG] [cultivated, 

naturalized] 

Manihot quinquepartita Huber ex Rogers & Appan [FG] 

Manihot surinamensis Rogers & Appan [GU SU FG] 

Manihot tristis Muell. Arg. subsp. saxicola (Lanjouw) Rogers 

& Appan [GU SU FG] 

Maprounea guianensis Aubl. [GU SU FG] 

Margaritaria nobilis L.f. [GU SU FG] 

Micrandra brownsbergensis Lanj. [SU FG] [probably 

conspecific with Micrandra elata] 

Micrandra glabra (R.E. Schultes) R.E. Schultes [GU SU] 

Micrandra gleasoniana (Croizat) R.E. Schultes [GU] 

Micrandra rossiana R.E. Schultes [GU] 

Micrandra siphonoides Benth. [FG] 

Micrandra spruceana (Baill.) R.E. Schultes [GU?] [record 

probably misident.] 

Omphalea diandra L. [GU SU FG] 

Pausandra fordii R. Secco [FG] 

Pausandra martinii Baill. [GU SU FG] 

Pausandra trianae (Muell. Arg.) Baill. [FG] [specimens not 

seen] 

Pedilanthus tithymaloides (L.) Poit. [GU SU FG] [native GU, 

SU; cultivated in all] 

Pera bicolor (Klotzsch) Muell. Arg. [GU SU FG] 

Pera decipiens (Muell. Arg.) Muell. Arg. [GU SU] 

Pera glabrata (Schott) Baill. [GU SU FG] 

Phyllanthus acuminatus Vahl [GU SU] 

Phyllanthus adiantoides Klotzsch [GU SU] 

Phyllanthus amarus Schumacher & Thonning [GU SU FG] 

Phyllanthus attenuatus Miq. [GU SU FG] 

Phyllanthus brasiliensis (Aubl.) Poir. [GU SU FG] 

Phyllanthus caroliniensis Walter subsp. caroliniensis [FG] 

Phyllanthus caroliniensis Walter subsp. guianensis (Klotzsch) 

Webster [GU SU FG] 

Phyllanthus caroliniensis Walter subsp. stenopterus (Muell. 

Arg.) Webster [FG] 

Phyllanthus debilis Klein ex Willd. [GU SU FG] [naturalized] 

Phyllanthus elsiae Urb. [GU SU] 

Phyllanthus gallinetae Jabl. [GU] 

Phyllanthus hyssopifolioides Kunth [SU? FG] 

Phyllanthus juglandifolius Willd. [GU FG] 

Phyllanthus majus Steyerm. [GU] 

Phyllanthus minutulus Muell. Arg. [GU] 

Phyllanthus niruri L. [SU FG] 

Phyllanthus orbiculatus L.C. Rich. [SU FG] 

Phyllanthus piscatorum Kunth [GU?] [specimens not seen] 

Phyllanthus pseudo-conami Muell. Arg. [GU] 

Phyllanthus pycnophyllus Muell. Arg. [GU] 

Phyllanthus stipulatus (Raf.) Webster [GU SU FG] 

Phyllanthus subglomeratus Poir. [GU FG] [cultivated only?] 

Phyllanthus tenellus Roxb. [GU SU] [naturalized] 

Phyllanthus urinaria L. [GU SU FG] [naturalized] 

Phyllanthus vacciniifolius (Muell. Arg.) Muell. Arg. subsp. 

vacciniifolius [GU] 

Piranhea trifoliata Baill. [GU?] 

Plukenetia loretensis Ule [GU] 

Plukenetia polyadenia Muell. Arg. [GU SU FG] 

Plukenetia supraglandulosa L.J. Gillespie [SU FG] 

Plukenetia verrucosa J.E. Smith [GU SU FG] 

Plukenetia volubilis L. [SU] 


Pogonophora schomburgkiana Miers ex Benth. [GU SU FG] 

Richeria grandis Vahl [GU FG] 

Ricinus communis L. [GU SU FG] [cultivated, naturalized] 

Sagotia brachysepala (Muell. Arg.) R. Secco [GU SU FG] 

Sagotia racemosa Baill. [GU SU FG] 

Sandwithia guyanensis Lanj. [GU FG] 

Sapium ciliatum Hemsley [SU FG] 

Sapium glandulosum (L.) Morong [GU SU FG] 

Sapium jenmanii Hemsley [GU SU] 

Sapium montanum Lanj. [SU FG?] [=Sapium argutum?] 

Sapium paucinervium Hemsley [GU SU FG] 

Sebastiania bidentata (Mart.) Pax [SU] 

Sebastiania corniculata (Vahl) Muell. Arg. [GU SU FG] 

Sebastiania guyanensis (Muell. Arg.) Muell. Arg. [GU] 

[=Gymnanthes guyanensis] 

Senefeldera macrophylla Ducke [GU SU FG] 

Senefelderopsis croizatii Steyerm. [GU] 

Tragia fendleri Muell. Arg. [GU] 

Tragia lessertiana (Baill.) Muell. Arg. [GU SU FG] 

Tragia tabulaemontana L.J. Gillespie [FG] 

Tragia volubilis L. [SU FG] 

EUPHORBIACEAE - SYNONYMS AND 


Acalypha persimilis Muell. Arg. var. corchorifolia (A. Rich.) 

Muell. Arg. [not in Guianas] 

Acalypha samydifolia Poepp. =Acalypha diversifolia 

Actinostemon depauperatus Pax & K. Hoffm. =Actinostemon 

schomburgkii 

Actinostemon guyanensis Pax & K. Hoffm. =Sandwithia 

guyanensis? 

Alchornea brachygyne Pax & K. Hoffm. =Alchornea 

schomburgkii 

Alchornea cordata (Juss.) Muell. Arg. =Aparisthium cordatum 

Alchorneopsis trimera Lanj. =Alchorneopsis floribunda 

Amanoa grandiflora (Muell. Arg.) Muell. Arg. =Amanoa 

guianensis 

Amanoa guianensis Aubl. var. grandiflora Muell. Arg. 

=Amanoa guianensis 

Angostyles tabulamontana Croizat [dubious species of 

uncertain placement] 

Apodandra loretensis (Ule) Pax & Hoffmann =Plukenetia 

loretensis 

Caperonia castaneifolia (L.) A. St. Hil. [see Caperonia 

paludosa] 

Chaetocarpus stipularis Gleason =Chaetocarpus 

schomburgkianus 

Chamaesyce glomerifera Millsp. =Chamaesyce hypericifolia 

Cleidion sp. [not in Guianas] 

Conami brasiliensis Aubl. =Phyllanthus brasiliensis 

Conceveibum cordatum Adr. Juss. =Aparisthium cordatum 

Croton argyrophyllus Muell. Arg. =Croton nervosus 

Croton bartlettii Lanj. =Croton potaroensis 

Croton glabellus L. [not in Guianas] 

Croton glandulosus L. [not in Guianas] 

Croton glandulosus L. subsp. hirtus (L'Hér.) Croizat =Croton 

hirtus 

Croton micans Swartz =Croton nervosus 

Croton miquelianus Ferg. =Croton trinitatis 

Croton populifolius P. Mill. [see Croton essequiboensis] 

Croton populifolius P. Mill. var. essequiboensis (Klotzsch) 

Muell. Arg. =Croton essequiboensis 

Croton surinamensis Muell. Arg. =Croton cuneatus 

Cunuria glabra R.E. Schult. =Micrandra glabra


Cunuria gleasoniana Croizat =Micrandra gleasoniana 

Cunuria spruceana Baill. =Micrandra spruceana 

Dactylostemon schomburgkii Klotzsch =Actinostemon 

schomburgkii 

Dalechampia spathulata (Scheidw.) Baill. [not in Guianas] 

Discocarpus mazarunensis Croizat =Chaetocarpus 


Dodecastigma mazarunense Croizat =Dodecastigma 

integrifolium 

Drypetes maguireana Monachino =Chaetocarpus 


Euphorbia adenoptera Bertol. =Chamaesyce adenoptera 

Euphorbia brasiliensis Lam. =Chamaesyce hyssopifolia 

Euphorbia buxifolia Lam. =Chamaescyce 

mesembryanthemifolia (Jacq.) Dugand 

Euphorbia cotinoides Miq. =Euphorbia cotinifolia 

Euphorbia heterophylla L. var. cyathophora (Murr.) Griseb. 

=Euphorbia cyathophora 

Euphorbia hirta L. =Chamaesyce hirta 

Euphorbia hypericifolia L. =Chamaesyce hypericifolia 

Euphorbia hyssopifolia L. =Chamaesyce hyssopifolia 

Euphorbia morisoniana Klotzsch =Euphorbia heterophylla 

Euphorbia multinodis Urban =Chamaesyce multinodis (Urban) 

Millsp.[not in Guianas] 

Euphorbia myrtillifolia L. [not in Guianas] 

Euphorbia prostrata Aiton =Chamaesyce prostrata 

Euphorbia serpens Kunth =Chamaesyce serpens 

Euphorbia thymifolia L. =Chamaesyce thymifolia 

Gaedawakka schomburgkiana Kuntze =Chaetocarpus 


Glochidion vacciniifolius Muell. Arg. =Phyllanthus 

vacciniifolius 

Gymnanthes guyanensis Muell. Arg. =Sebastiana guyanensis 

Hevea confusa Hemsley =Hevea pauciflora 

Hyeronima laxiflora (Tul.) Muell. Arg. =Hyeronima 

alchorneoides var. alchorneoides 

Jatropha urens L. =Cnidoscolus urens 

Mabea argutissima Croizat =Mabea subsessilis 

Mabea caudata Pax & K. Hoffm. =Mabea speciosa subsp. 

speciosa 

Mabea caudata Pax & K. Hoffm. var. concolor Lanj. =Mabea 

speciosa subsp. A 

Mabea ciliata Gleason =Mabea montana 

Mabea costata Pax & K. Hoffm. =Mabea montana 

Mabea eximia Ducke =Mabea pulcherrima 

Mabea lucida Pax & K. Hoffm. =Mabea montana 

Mabea montana Muell. Arg. subsp. biglandulosa (Muell. Arg.) 

den Hollander =Mabea biglandulosa 

Mabea montana Muell. Arg. subsp. lucida (Pax & K. Hoffm.) 

den Hollander =Mabea montana 

Mabea occidentalis Benth. [not in Guianas] 

Mabea occidentalis Benth. var. laevigata (Muell. Arg.) Muell. 

Arg. =Mabea biglandulosa 

Mabea occidentalis Benth. var. obovata (Muell. Arg.) Muell. 

Arg. =Mabea speciosa subsp. B 

Mabea occidentalis Benth. var. purpurascens (Muell. Arg.) 

Muell. Arg. =Mabea speciosa subsp. B 

Mabea piriri Aubl. var. laevigata Muell. Arg. =Mabea 

biglandulosa 

Mabea piriri Aubl. var. obovata Muell. Arg. =Mabea speciosa 

subsp. B 

Mabea piriri Aubl. var. purpurascens Muell. Arg. =Mabea 

speciosa subsp. B 

Mabea saramaccensis Croizat =Mabea speciosa subsp. A 

Mabea schomburgkii Benth. =Mabea taquari 


103 

Mabea speciosa Muell. Arg. subsp. concolor (Muell. Arg.) den 

Hollander =Mabea piriri 

Mabea taquari Aubl. var. angustifolia Muell. Arg. =Mabea 

taquari 

Manihot dulcis Pax. =Manihot esculenta 

Manihot glaziovii Muell. Arg. [not in Guianas] 

Manihot melanobasis Muell. Arg. [not in Guianas] 

Manihot saxicola Lanj. =Manihot tristis subsp. saxicola 

Manihot sprucei Pax =Manihot esculenta 

Manihot tristis Muell. Arg. subsp. tristis [not in Guianas] 

Meborea guianensis Aubl. =Phyllanthus sp.?[name of 

uncertain application] 

Micrandra elata (Didrichs.) Muell. Arg. [see Micrandra 

brownsbergensis] 

Microstachys guianensis Klotzsch =Sebastiana corniculata 

Pausandra flagellorhachis Lanj. =Pausandra martinii 

Pausandra integrifolia Lanj. =Dodecastigma integrifolium 

Pera echinocarpa Baill. [not in Guianas] 

Pera ferruginea (Schott) Muell. Arg. =Pera glabrata 

Pera nitida (Benth.) Jabl. =Pera decipiens 

Pera obovata Baill. [not in Guianas] 

Pera schomburgkiana Benth. ex Muell. Arg. =Pera bicolor 

Peridium bicolor Klotzsch =Pera bicolor 

Peridium bicolor Klotzsch var. nitidum Benth. =Pera 

decipiens 

Peridium decipiens Muell. Arg. =Pera decipiens 

Phyllanthus conami Swartz =Phyllanthus brasiliensis 

Phyllanthus diffusus Klotzsch =Phyllanthus stipulatus 

Phyllanthus epiphyllanthus L. [not in Guianas] 

Phyllanthus grandifolius L. [not in Guianas] 

Phyllanthus guianensis Klotzsch =Phyllanthus caroliniensis 

subsp. guianensis 

Phyllanthus guyanensis Muell. Arg. =Phyllanthus attenuatus 

Phyllanthus lathyroides Kunth [not in Guianas] 

Phyllanthus nobilis (L.f.) Muell. Arg. =Margaritaria nobilis 

Phyllanthus pentrandrus Schum. & Th. [not in Guianas] 

Plukenetia abutifolia (Ducke) Pax & Hoffmann =Plukenetia 

polyadenia 

Richeria laurifolia (Baill.) Baill. =Richeria grandis 

Sagotia racemosa Baill. var. brachysepala Muell. Arg. 

=Sagotia brachysepala 

Sagotia tafelbergii Croizat =Sagotia racemosa 

Sapium argutum Muell. Arg. [see Sapium montanum] 

Sapium aubletianum (Muell. Arg.) Hub. =Sapium 

glandulosum 

Sapium aucuparium Jacq. [see Sapium glandulosum] 

Sapium biglandulosum (L.) Muell. Arg. [see Sapium 

glandulosum] 

Sapium biglandulosum (L.) Muell. Arg. var. aubletianum 

Muell. Arg. =Sapium glandulosum 

Sapium biglandulosum (L.) Muell. Arg. var. klotzschianum 


Sapium biglandulosum (L.) Muell. Arg. var. lanceolatum 


Sapium cladogyne Hutch. =Sapium jenmanii

104 


Sapium ellipticum Hemsley [see Sapium glandulosum] 

Sapium hemsleyanum Huber [see Sapium glandulosum] 

Sapium hippomane G. Mey. =Sapium glandulosum 

Sapium klotzschianum (Muell. Arg.) Huber =Sapium 

glandulosum 

Sapium lanceolatum (Muell. Arg.) Huber =Sapium 

glandulosum 

Sapium microdentatum Lanj. =Sapium paucinervium 

Sapium obtusilobium Muell. Arg. =Sapium glandulosum 

Sapium prunifolium Klotzsch =Sapium glandulosum 

Sebastiania corniculata (Vahl) Muell. Arg. var. guyanensis 

(Klotzsch) Pax & K. Hoffm. =Sebastiana corniculata 

Sebastiania linearifolia Lanj. =Sebastiana bidentata 

Sebastiania micrantha (Benth.) Lanj. =Sebastiana corniculata 

Securinega congesta (Benth. ex Muell. Arg.) Muell. Arg 

=Jablonskia congesta 

Senefelderopsis sipapoensis Jabl. =Senefelderopsis croizatii 

Stilaginella laxiflora Tul. =Hyeronima alchorneoides var. 

alchorneoides 

Stilaginella oblonga Tul. =Hyeronima oblonga 

Traganthus sidoides Klotzsch =Bernardia sidoides 

Tragia corniculata Vahl =Sebastiana corniculata 

Tragia grandifolia Klotzsch =Adenophaedra grandifolia 

FABACEAE 

Aeschynomene, Chaetocalyx, & Machaerium by 

V.E. Rudd; Canavalia by G. Aymard; Clitoria 

by P.R. Fantz; Dioclea by R.H. Maxwell; 

Lonchocarpus by M. Sousa; Macroptilium, 

Phaseolus, & Vigna by A. Delgado Salinas; 

Crotalaria reviewed by D.R. Windler (1992); 

reviewed by P.R. Fantz, V.E. Rudd, & G. 

Aymard (1996) 

REFERENCES: 

Amshoff, G.J.H. 1939. Papilionaceae. In A. 


Aymard C., G. & N. Cuello A. 1991. Dos 

nuevas especies Neotropicales del genero 

Canavalia (Papilionoideae - Phaseoleae - 

Diocleinae). Biollania 8: 87-92. 

Burkart, A. 1971. El género Galactia (Legum.- 

Phaseoleae) en Sudamérica. Darwiniana 16: 

663-796. 





Ducke, A. 1942. Lonchocarpus, subgenus 

Phacelanthus Pittier, in Brazilian Amazonia. 

Trop. Woods 69: 2-7. 

Dwyer, J.D. 1980. Leguminosae subfamily 

Papilionoideae (conclusion). In R.E. 

Woodson, Jr., R.W. Schery, & collaborators, 


67: 523-818. 

Görts-van Rijn, A.R.A., K.U. Kramer, & J.C. 

Lindeman. 1976. Papilionaceae (additions 



SPERMATOPHYTA: Euphorbiaceae - Fabaceae 

Grear, J.W. 1971. A revision of the American 

species of Eriosema. Mem. New York Bot. 

Gard. 20(3): 1-98. 

Grear, J.W. 1978. A revision of the New World 

species of Rhynchosia. Mem. New York Bot. 

Gard. 31(1): 1-168. 

Krukoff, B.A. 1969. Supplementary notes on the 

American species of Erythrina. III. 

Phytologia 19: 113-175. 

Lima, H.C. de. 1982. Revisao taxonômica do 

género Vatairea Aublet (Leguminosae - 

Faboideae). Arquiv. Jard. Bot. Rio de Janeiro 

26: 173-213. 

Lima, H.C. de. 1985. Diplotropis Bentham. 

Estudo dos Táxons infragenéricos. Acta 

Amaz. 15: 61-75. 

Maxwell, R.H. 1970. The genus Cymbosema 

(Leguminosae): notes and distribution. Ann. 


Mohlenbrock, R.H. 1958. A revision of the 

genus Stylosanthes. Ann. Missouri Bot. Gard. 

44: 299-355. 

Mohlenbrock, R.H. 1961. A monograph of the 

leguminous genus Zornia. Webbia 16(1): 1- 

141. 

Mohlenbrock, R.H. 1963. Further considerations 

in Stylosanthes. Rhodora 65: 245-258. 




Ramírez, N. 1995. Revisión taxonómica del 

género Alexa Moq. (Fabaceae, Sophoreae). 

Ann. Missouri Bot. Gard. 82(4): 549-569. 

Rojo, J.P. 1972. Pterocarpus. Revised for the 

world. Phanerog. Monogr. 5: 1-119. 

Rudd, V.E. 1955. The American species of 

Aeschynomene. Contr. U.S. Nat. Herb. 32: 

1-172. 

Rudd, V.E. 1955. A revision of the genus 

Chaetocalyx. Contr. U.S. Nat. Herb. 32: 247- 

277. 

Rudd, V.E. 1963. The genus Dussia 

(Leguminosae). Contr. U.S. Nat. Herb. 32: 

247-277. 

Rudd, V.E. 1965. The American species of 

Ormosia. Contr. U.S. Nat. Herb. 32: 279-384. 

Rudd, V.E. 1981. Two new species of 

Paramachaerium and a brief resumé of the 

genus. Brittonia 33(3): 435-440. 

Rudd, V.E. 1987. Studies in Machaerium 

(Leguminosae) VI. Phytologia 62: 282-302. 

Rudd, V.E. 1989. Studies in Machaerium VII. 

Section II, Lineata. Part I. Species with 

wingless fruit. Phytologia 64: 1-12. 

Sauer, J. 1964. Revision of Canavalia. Brittonia 

16: 106-181. 

Verdcourt, B. 1970. Studies in the Leguminosae 

- Papilionoideae for the 'Flora of Tropical East 

Africa': IV. Kew Bull. 24: 507-569. 

Schultze-Kraft, R.J. & L. Coradin. 1990. 

Biogeography of Centrosema. In R. Schultze- 

Kraft & R.J. Clements (ed.), Centrosema: 

Biology, Agronomy, and Utilization. 

Williams, R.J. & R.R. Clements. 1990. 

Taxonomy of Centrosema. In R. Schultze-


Kraft & R.J. Clements (ed.), Centrosema: 

Biology, Agronomy, and Utilization. 

Yakovlev, G.P. 1969. A review of Sweetia and 

Acosmium. Notes Royal Bot. Gard. Edinb. 

29: 347-355. 

Abrus fruticulosus Wallich ex Wight & Walker-Arnott [SU] 

[introduced] 

Abrus precatorius L. [GU SU FG] [cultivated, naturalized] 

Acosmium nitens (Vogel) Yakovlev [GU SU FG] 

Acosmium praeclarum (Sandw.) Yakovlev [GU] 

Aeschynomene americana L. [GU FG] 

Aeschynomene brasiliana (Poir.) DC. [GU SU FG] 

Aeschynomene evenia Wright [GU?] 

Aeschynomene filosa Mart. ex Benth. [GU] 

Aeschynomene fluminensis Vell. [GU SU FG] 

Aeschynomene histrix Poir. var. densiflora (Benth.) Rudd 

[GU] 

Aeschynomene histrix Poir. var. histrix [GU SU FG] 

Aeschynomene interrupta Benth. [GU] 

Aeschynomene paniculata Willd. ex Vogel [GU SU] 

Aeschynomene pratensis Small var. caribaea Rudd [GU FG] 

Aeschynomene rudis Benth. [GU] 

Aeschynomene sensitiva Swartz [GU SU FG] 

Alexa cowanii Yakovlev [GU] 

Alexa grandiflora Ducke [FG?] 

Alexa imperatricis (Rob. Schomb.) Baill. [GU SU?] 

Alexa surinamensis Yakovlev [GU SU] 

Alexa wachenheimii Benoist [GU SU FG] 

Alysicarpus vaginalis (L.) DC. [GU SU FG] [introduced] 

Andira coriacea Pulle [SU FG] 

Andira grandistipula Amsh. [GU] 

Andira inermis (Wright) DC. [GU SU FG] 

Andira surinamensis (Bondt) Splitg. ex Amshoff [GU SU FG] 

Andira villosa Kleinh. [SU] 

Bowdichia virgilioides Kunth [GU SU] 

Cajanus cajan (L.) Millsp. [GU SU FG] [cultivated, 

naturalized] 

Calopogonium coeruleum (Benth.) Sauv. [GU SU FG] 

Calopogonium mucunoides Desv. [GU SU FG] 

Calopogonium velutinum (Benth.) Amsh. [GU SU FG] 

Canavalia brasiliensis Mart. ex Benth. [GU SU FG] 

Canavalia dictyota Piper var. dictyota [GU SU FG] 

Canavalia dictyota Piper var. ferruginea Aymard & Cuello 

[SU] 

Canavalia ensiformis (L.) DC. [GU? SU FG] [escaped from 

cultivation] 

Canavalia gladiata (Jacq.) DC. var. gladiata [SU] [escaped 


Canavalia grandiflora Benth. [GU] 

Canavalia plagiosperma Piper [GU SU] [probably escaped 


Canavalia rosea (Swartz) DC. [GU SU FG] 

Centrolobium paraense Tul. [GU FG] 

Centrosema angustifolium (Kunth) Benth. [GU SU] 

Centrosema brasilianum (L.) Benth. [GU SU FG] 

Centrosema capitatum (Rich.) Amsh. [GU SU FG] 

Centrosema macrocarpum Benth. [GU] 

Centrosema pascuorum Mart. ex Benth. [GU] 

Centrosema plumieri (Turp. ex Pers.) Benth. [GU SU? FG?] 

Centrosema sp. aff. molle Mart. ex Benth. [GU SU FG] 

[specimens previously identified as Centrosema 

pubescens] 

Centrosema triquetrum (Hoffm. ex Benth.) Benth. & Hook. 

[GU] 

Centrosema vexillatum Benth. [GU SU] 

Centrosema virginianum Benth. [GU? SU FG?] 

Chaetocalyx brasiliensis (Vogel) Benth. [SU FG?] 

Clathrotropis brachypetala (Tul.) Kleinh. var. brachypetala 

[GU SU FG] 

Clathrotropis brachypetala (Tul.) Kleinh. var. ferruginea 

Yakovlev [SU] 

Clathrotropis macrocarpa Ducke [GU] 

Clathrotropis paradoxa Sandw. [GU] 

Clitoria arborescens R. Brown [GU SU FG] 

Clitoria brachycalyx Harms [GU] 

Clitoria falcata Lam. var. falcata [GU SU FG] 

Clitoria guianensis (Aubl.) Benth. var. guianensis [GU SU 

FG] 

Clitoria kaieteurensis Fantz [GU] 

Clitoria laurifolia Poir. [GU SU FG] 

Clitoria leptostachya Benth. var. leptostachya [GU] 

Clitoria pendens Fantz [SU] 

Clitoria sagotii Fantz var. canaliculata Fantz [GU SU] 

Clitoria sagotii Fantz var. sagotii [FG] 

Clitoria ternatea L. var. ternatea [SU FG] [cultivated or 

escaped from cultivation] 

Coursetia ferruginea (Kunth) Lavin [GU] 

Crotalaria goreensis Guill. & Perr. [GU FG] [introduced] 

Crotalaria incana L. [GU] 

Crotalaria juncea Willd. [GU SU] [introduced] 

Crotalaria maypurensis Kunth [GU] 

Crotalaria micans Link [GU SU FG] 

Crotalaria nitens Kunth [GU] 

Crotalaria pallida Aiton [GU] [introduced] 

Crotalaria pilosa P. Mill. [GU SU FG] 

Crotalaria quinquefolia L. [GU] [introduced] 

Crotalaria retusa L. [GU SU FG] 

Crotalaria stipularia Desv. [GU SU FG] 

Crotalaria verrucosa L. [GU SU] [introduced] 

Crotalaria zanzibarica Benth. [GU] [introduced] 

Cymbosema roseum Benth. [to be expected in the Guianas] 

Dalbergia brownei (Jacq.) Urb. [GU SU FG] 

Dalbergia ecastaphyllum (L.) Taubert [GU SU FG] 

Dalbergia glauca (Desv.) Amsh. [GU SU FG?] 

Dalbergia inundata Spruce ex Benth. [GU] 

Dalbergia monetaria L.f. [GU SU FG] 

Dalbergia riedelii (Radlk.) Sandw. [GU SU FG] 

Dalbergia subcymosa Ducke [SU FG] 

Desmodium adscendens (Swartz) DC. [GU SU FG] 

Desmodium asperum (Poir.) Desv. [GU SU FG] 

Desmodium axillare (Swartz) DC. [GU SU FG] 

Desmodium barbatum (L.) Benth. & Oerst. [GU SU FG] 

Desmodium cajanifolium (Kunth) DC. [GU SU] 

Desmodium distortum (Aubl.) Macbr. var. perrottetii (DC.) 

Schind. [GU FG] 

Desmodium guianense DC. [FG] 

Desmodium incanum (Swartz) DC. [GU SU FG] 

Desmodium molliculum (Kunth) DC. [GU] 

Desmodium procumbens (Mill.) Hitchc. [SU] 

Desmodium sclerophyllum Benth. [GU SU] 

Desmodium scorpiurus (Swartz) Desv. [GU FG] 

Desmodium tortuosum (Swartz) DC. [SU] [introduced] 

Desmodium triflorum (L.) DC. [GU SU FG] [introduced] 

Desmodium velutinum DC. [FG] 

Desmodium wydlerianum Urb. [GU SU] 

Dioclea apurensis Kunth [SU] [unverified name] 

Dioclea coriacea Benth. [SU] 

Dioclea elliptica Maxwell, sp. nov. ined. [GU SU FG] 

SPERMATOPHYTA: Fabaceae 

105

106 



Dioclea guianensis Benth. [GU SU FG] 

Dioclea huberi Ducke [FG?] 

Dioclea macrantha Huber var. bracteolata (Merrill) Maxwell, 

comb. nov. ined. [GU SU?] 

Dioclea macrantha Huber var. macrantha [GU SU] 

Dioclea macrocarpa Huber [GU SU FG] 

Dioclea malacocarpa Ducke [GU SU FG] 

Dioclea megacarpa Rolfe [FG?] 

Dioclea reflexa J.D. Hook. [GU SU FG] 

Dioclea scabra (Rich.) Maxwell var. scabra [GU SU FG] 

Dioclea scabra (Rich.) Maxwell var. schulzii Maxwell [GU] 

Dioclea violacea Mart. ex Benth. [GU? SU? FG?] 

Dioclea virgata (Rich.) Amsh. var. crenata Maxwell [GU SU 

FG] 

Dioclea virgata (Rich.) Amsh. var. virgata [GU SU FG] 

Dioclea wilsonii Standl. [GU SU FG] 

Diplotropis purpurea (Rich.) Amsh. var. leptophylla (Kleinh.) 

Amsh. [SU FG] 

Diplotropis purpurea (Rich.) Amsh. var. purpurea [GU SU 

FG] 

Diplotropis racemosa (Hoehne) Amsh. var. racemosa [GU] 

Diplotropis rigidocarpa Lima [GU] 

Dipteryx odorata (Aubl.) Willd. [GU SU FG] 

Dipteryx punctata (Blake) Amsh. [GU SU FG] 

Dussia discolor (Benth.) Amsh. [GU SU FG] 

Eriosema crinitum (Kunth) G. Don var. crinitum [GU SU FG] 

Eriosema crinitum (Kunth) G. Don var. stipulare (Benth.) 

Fortunato [GU] 

Eriosema rufum (Kunth) G. Don. var. rufum [GU SU] 

Eriosema simplicifolium (Kunth) G. Don var. simplicifolium 

[GU SU FG] 

Eriosema violaceum (Aubl.) G. Don [GU SU FG] 

Erythrina amazonica Krukoff [GU SU FG] 

Erythrina fusca Loureiro [GU SU FG] 

Etaballia dubia (Kunth) Rudd [GU] 

Fissicalyx fendleri Benth. [GU] 

Flemingia strobilifera (L.) R. Brown [SU] [introduced] 

Galactia dubia DC. [FG?] 

Galactia gracillima Benth. [SU?] 

Galactia jussiaeana Kunth var. jussiaeana [GU SU] 

Galactia latisiliqua Desv. [GU] 

Galactia longifolia (Jacq.) Benth. [FG] 

Galactia striata (Jacq.) Urb. [GU] 

Gliricidia sepium (Jacq.) Steud. [GU SU FG] [widely 

cultivated & naturalized] 

Hymenolobium excelsum Ducke [FG?] 

Hymenolobium flavum Kleinh. [GU SU FG] 

Hymenolobium nitidum Benth. [GU] 

Hymenolobium petraeum Ducke [GU SU FG] 

Hymenolobium pulcherrimum Ducke [FG] 

Indigofera endecaphylla Jacq. [GU FG?] 

Indigofera hirsuta L. [GU FG] [cultivated only?] 

Indigofera lespedezioides Kunth [GU] 

Indigofera microcarpa Desv. [GU SU FG] 

Indigofera spicata Forsskal [FG] 

Indigofera suffruticosa Mill. [GU SU FG] [cultivated only?] 

Indigofera tinctoria L. [GU FG] [cultivated, perhaps escaped] 

Lablab purpureus (L.) Sweet [GU? SU FG] [widely cultivated] 

Lonchocarpus chrysophyllus Kleinh. [GU SU FG] 

Lonchocarpus densiflorus Benth. [GU] 

Lonchocarpus floribundus Benth. [GU SU FG] 

Lonchocarpus hedyosmus Miq. [SU FG] 

Lonchocarpus heptaphyllus (Poir.) DC. [GU SU FG] 

Lonchocarpus longifolius (Benth.) Ducke [SU?] 

Lonchocarpus margaritensis Pittier [GU] 

Lonchocarpus martynii A.C. Smith [GU] 

Lonchocarpus negrensis Benth. [SU FG] 

Lonchocarpus nicou (Aubl.) DC. [GU SU? FG] 

Lonchocarpus rariflorus Mart. ex Benth. [GU] 

Lonchocarpus rufescens Benth. [GU] 

Lonchocarpus scandens (Aubl.) Ducke [GU SU FG] 

Lonchocarpus sericeus (Poir.) DC. [to be expected in the 


Lonchocarpus spruceanus Benth. [FG] 

Luetzelburgia sp. [GU] 

Machaerium affine Benth. [GU] 

Machaerium altiscandens Ducke [SU FG] 

Machaerium aureiflorum Ducke [GU FG] 

Machaerium ferox (Mart. ex Benth.) Ducke [GU SU] 

Machaerium floribundum Benth. [GU FG] 

Machaerium inundatum (Mart. ex Benth.) Ducke [GU SU FG] 

Machaerium isadelphum (E. Mey.) Amsh. [SU] 

Machaerium kegelii Meisn. [GU SU FG] 

Machaerium leiophyllum (DC.) Benth. var. latifolium (Benth.) 

Rudd [GU FG] 

Machaerium leiophyllum (DC.) Benth. var. leiophyllum [GU 

SU FG] 

Machaerium lunatum (L.f.) Ducke [GU SU FG] 

Machaerium macrophyllum Benth. [GU SU FG] 

Machaerium madeirense Pittier [GU SU] 

Machaerium mutisii Killip ex Rudd [SU] 

Machaerium myrianthum Spruce ex Benth. [GU SU] 

Machaerium paraense Ducke [SU FG] 

Machaerium polyphyllum (Poir.) Benth. [SU FG] 

Machaerium quinata (Aubl.) Sandw. var. parviflorum (Benth.) 

Rudd [GU SU FG] 

Machaerium quinata (Aubl.) Sandw. var. quinata [GU SU 

FG] 

Machaerium robiniifolium (DC.) Vogel [GU] 

Machaerium trifoliolatum Ducke [GU SU FG] 

Macroptilium atropurpureum (Sessé & Moçino ex DC.) Urb. 

[FG] 

Macroptilium gracile (Poepp. ex Benth.) Urb. [GU FG] 

Macroptilium lathyroides (L.) Urban [GU SU FG] 

Macroptilium longepedunculatum (Mart. ex Benth.) Urb. [GU 

SU FG] 

Monopteryx inpae Rodrigues [FG] 

Mucuna pruriens (L.) DC. [SU] [cultivated, naturalized] 

Mucuna sloanei Fawcett & Rendle [GU SU FG] 

Mucuna urens (L.) DC. [GU SU FG] 

Muellera frutescens (Aubl.) Standl. [GU SU FG] 

Myroxylon peruiferum L.f. [GU] 

Ormosia cinerea Benoist [SU FG] 

Ormosia coarctata Jackson [GU SU FG] 

Ormosia coccinea (Aubl.) Jackson [GU SU FG] 

Ormosia costulata (Miq.) Kleinh. [GU SU] 

Ormosia coutinhoi Ducke [GU SU FG] 

Ormosia flava (Ducke) Rudd [SU FG] 

Ormosia guyanensis C.H. Stirton, sp. nov. ined. [GU] 

Ormosia lignivalvis Rudd [GU FG] 

Ormosia melanocarpa Kleinh. [SU FG] 

Ormosia nobilis Tul. [GU FG] 

Ormosia paraensis Ducke [GU SU FG] 

Ormosia stipularis Ducke [GU SU FG] 

Ormosia wayana Stirton [FG] 

Paramachaerium ormosioides (Ducke) Ducke [FG] 

Paramachaerium schomburgkii (Benth.) Ducke [GU FG] 

Phaseolus lunatus L. [GU SU FG] [cultivated, naturalized] 

Phaseolus reptans Ducke [SU?] 

Phaseolus vulgaris L. [GU SU FG] [probably cultivated only] 

Platymiscium filipes Benth. [FG] 

Platymiscium pinnatum (Jacq.) Dugand [GU FG]


Platymiscium trinitatis Benth. [GU SU FG] 

Platymiscium ulei Harms [SU FG] 

Poecilanthe effusa (Huber) Ducke [SU FG] 

Poecilanthe hostmannii (Benth.) Amsh. [GU SU FG] 

Poecilanthe ovalifolia Kleinh. [SU] 

Pterocarpus amazonum (Benth.) Amsh. [GU] 

Pterocarpus officinalis Jacq. subsp. officinalis [GU SU FG] 

Pterocarpus rohrii Vahl [GU SU FG] 

Pterocarpus santalinoides L'Hér. ex DC. [GU SU FG] 

Pueraria phaseoloides (Roxb.) Benth. [GU SU FG] 

[naturalized] 

Rhynchosia melanocarpa Grear [perhaps to be expected in the 


Rhynchosia minima (L.) DC. [GU SU FG] 

Rhynchosia phaseoloides (Swartz) DC. [GU SU FG] 

Rhynchosia schomburgkii Benth. [GU] 

Sesbania exasperata Kunth [SU FG] 

Sesbania grandiflora (L.) Pers. [cultivated, occasionally 

escaping] 

Sesbania sericea (Willd.) DC. [GU SU FG] [introduced] 

Sesbania sesban (L.) Merrill [SU] [introduced] 

Soemmeringia semperflorens Mart. [GU FG?] 

Spirotropis longifolia (DC.) Baill. [SU FG] 

Stylosanthes angustifolia Vogel [GU SU FG] 

Stylosanthes cayennensis Mohlenbrock [FG] [=Stylosanthes 

guianensis?] 

Stylosanthes guianensis (Aubl.) Swartz var. gracilis (Kunth) 

Vogel [GU SU] 

Stylosanthes guianensis (Aubl.) Swartz var. guianensis [GU 

SU FG] 

Stylosanthes hispida L.C. Rich. [GU SU FG] [=Stylosanthes 

guianensis?] 

Stylosanthes viscosa Swartz [GU SU FG] 

Taralea crassifolia (Spruce ex Benth.) Ducke [GU] 

Taralea nudipes (Tul.) Ducke [FG] 

Taralea oppositifolia Aubl. [GU SU FG] 

Taralea reticulata (Benth.) Ducke [GU] 

Tephrosia adunca Benth. [GU SU] 

Tephrosia candida DC. [SU] [introduced] 

Tephrosia cinerea (L.) Pers. [GU SU FG] 

Tephrosia purpurea (L.) Pers. [SU] 

Tephrosia sessiliflora (Poir.) Hassler [GU SU] 

Tephrosia sinapou (Bucholz) A. Chevalier [GU SU FG] 

Teramnus labialis Spreng. [GU] [introduced?] 

Teramnus uncinatus (L.) Swartz [FG?] 

Vatairea erythrocarpa Ducke [FG] 

Vatairea guianensis Aubl. [GU SU FG] 

Vatairea paraensis Ducke [SU FG] 

Vataireopsis speciosa Ducke [GU SU FG] [records may 

represent misidentifications] 

Vataireopsis surinamensis Lima [GU SU FG] 

Vigna adenantha (G. Mey.) Maréchal, Mascherpa & Stain 

[GU? SU FG] 

Vigna hosei (Craig) Baker [SU FG] [cultivated, naturalized] 

Vigna juruana (Harms) Verdcourt [GU SU FG] 

Vigna lasiocarpa (Benth.) Verdcourt [GU SU FG] 

Vigna linearis (Kunth) Maréchal, Mascherpa, & Stainier [GU 

SU] 

Vigna longifolia (Benth.) Verdcourt [FG] 

Vigna luteola (Jacq.) Benth. [GU SU FG] 

Vigna peduncularis (Kunth) Fawcett & Rendle [SU FG] 

Vigna trichocarpa (C. Wright) A. Delgado, comb. nov. ined. 

[GU SU FG] 

Vigna unguiculata (L.) Walpers [GU SU FG] [cultivated, 

naturalized] 

Vigna vexillata (L.) A. Rich. [GU SU FG] 

Zornia curvata Mohlenbrock [GU] 

Zornia diphylla (L.) Pers. [GU] 

Zornia glabra Desv. [GU SU FG] 

Zornia gracilis DC. [FG] 

Zornia guanipensis Pittier [SU] 

Zornia latifolia J.E. Smith var. latifolia [GU SU FG] 

Zornia pardina Mohlenbrock var. vichadana Killip ex 

Mohlenbrock [GU] 

Zornia reticulata Sm. [GU] 

FABACEAE - SYNONYMS AND 



107 

Acouroa violacea Aubl. =Dalbergia sp.? 

Aeschynomene biflora (Mill.) Fawc. & Rend. =Aeschynomene 

brasiliana 

Aeschynomene densiflora Benth. =Aeschynomene histrix var. 

densiflora 

Aeschynomene gracilis Miq. =Aeschynomene paniculata 

Aeschynomene hispida Willd. =Aeschynomene virginica [not 


Aeschynomene viscidula Michx. [not in Guianas] 

Agati grandiflora (L.) Desv. =Sesbania grandiflora 

Alexa leiopetala Sandw. =Alexa imperatricis 

Alexandra imperatricis (Rob. Schomb.) Baill. =Alexa 

imperatricis 

Amerimnum latifolium Willd. =Lonchocarpus heptaphyllus 

Amphiodon effusus Huber =Poecilanthe effusa 

Andira retusa (Lam.) Kunth =Andira surinamensis 

Bowdichia guianensis Ducke =Diplotropis purpurea 

Cajanus indicus Spreng. =Cajanus cajan 

Canavalia bracteolata Merrill =Dioclea macrantha var. 

bracteolata 

Canavalia maritima (Aubl.) Thouars =Canavalia rosea 

Canavalia obtusifolia DC. =Canavalia rosea 

Centrosema latissimum Ducke =Centrosema triquetrum 

Centrosema platycarpum Benth. [not in Guianas] 

Centrosema pubescens Auctt, non Benth. =Centrosema sp. aff. 

molle Mart. ex Benth. 

Clathrotropis flava Ducke =Ormosia flava 

Clathrotropis surinamensis Kleinh. =Ormosia flava 

Clitoria amoena Miq. =Clitoria arborescens 

Clitoria angustifolia Kunth =Centrosema angustifolium 

Clitoria arborea Mart. ex Benth. [not in Guianas] 

Clitoria brasiliana L. =Centrosema brasilianum 

Clitoria capitata Rich. =Centrosema capitatum 

Clitoria glycinoides DC. =Clitoria falcata 

Clitoria guyanensis Aubl. ex Desv. =Clitoria laurifolia 

Clitoria javitensis (Kunth) Benth. var. glabra Sagot =Clitoria 

sagotii 

Clitoria plumieri Turp. ex Pers. =Centrosema plumieri 

Clitoria poiteai Benth. =Clitoria arborescens 

Clitoria poiteai DC. =Clitoria arborescens 

Clitoria rubiginosa Juss. ex Pers. =Clitoria falcata var. falcata 

Coronilla sericea Willd. =Sesbania sericea 

Coublandia frutescens Aubl. =Muellera frutescens 

Coumarouna odorata Aubl. =Dipteryx odorata 

Coumarouna oppositifolia (Aubl.) Taub. =Taralea 

oppositifolia 

Coumarouna punctata Blake =Dipteryx punctata 

Crotalaria anagyroides Kunth =Crotalaria micans 

Crotalaria gajanensis Desv. =Clitoria laurifolia 

Crotalaria guianensis Aubl. =Clitoria guianensis 

Crotalaria paramariboensis Miq. =Crotalaria verrucosa 

Crotalaria pterocaula Desv. =Crotalaria pilosa

108 



Crotalaria saltiana Andr. =Crotalaria pallida 

Cyclolobium hostmannii Benth. =Poecilanthe hostmannii 

Cymbosema apurense (Kunth) Pittier =Dioclea apurensis 

Cytisus sessiliflorus Poir. =Tephrosia sessiliflora 

Cytisus violaceus Aubl. =Eriosema violaceum 

Dalbergia atropurpurea Ducke =Dalbergia glauca 

Deguelia scandens Aubl. =Lonchocarpus scandens 

Derris amazonica Killip =Lonchocarpus negrensis 

Derris guianensis Benth. =Lonchocarpus scandens 

Derris longifolia Benth. =Lonchocarpus longifolius 

Derris pterocarpa (DC.) Killip =Lonchocarpus scandens 

Derris scandens (Aubl.) Pitt. =Lonchocarpus scandens 

Desmodium canum (J.F. Gmelin) Schinz & Thellung 

=Desmodium incanum 

Desmodium frutescens (Jacq.) Schindler =Desmodium 

incanum 

Desmodium molle (Vahl) DC. =Desmodium glabrum (Mill.) 


Desmodium spectabilis Miq. =Desmodium asperum 

Desmodium supinum DC. =Desmodium incanum 

Dibrachion guianense Tul. =Diplotropis purpurea 

Dioclea glabra Benth. =Dioclea elliptica 

Dioclea lasiocarpa Mart. =Dioclea virgata 

Diplotropis brachypetala Tul. =Clathrotropis brachypetala var. 

brachypetala 

Diplotropis guianensis Benth. =Diplotropis purpurea 

Diplotropis leptophylla Kleinh. =Diplotropis purpurea var. 

leptophylla 

Diplotropis racemosa (Hoehne) Amsh. var. kaieteurensis 

Amsh. =Diplotropis racemosa var. racemosa 

Dipteryx crassifolia Spruce ex Benth. =Taralea crassifolia 

Dipteryx nudipes Tul. =Taralea nudipes 

Dipteryx oppositifolia (Aubl.) Willd. =Taralea oppositifolia 

Dolichos ensiformis L. =Canavalia ensiformis 

Dolichos gladiatus Jacq. =Canavalia gladiata 

Dolichos lablab L. =Lablab purpureus 

Dolichos luteolus Jacq. =Vigna luteola 

Dolichos maritimus Aubl. =Canavalia rosea 

Dolichos minimus L. =Rhynchosia minima 

Dolichos roseus Swartz =Canavalia rosea 

Dolichos unguiculatus L. =Vigna unguiculata 

Dolichos urens L. =Mucuna urens 

Dolichos virgatus Rich. =Dioclea virgata 

Drepanocarpus ferox Mart. ex Benth. =Machaerium ferox 

Drepanocarpus inundatus Mart. ex Benth. =Machaerium 

inundatum 

Drepanocarpus isadelphus E. Mey. =Machaerium isadelphum 

Drepanocarpus lunatus (L.f.) G. Mey. =Machaerium lunatum 

Drepanocarpus macrophyllus Ducke =Machaerium 

macrophyllum 

Dussia cayennensis Harms =Dussia discolor 

Ecastophyllum benthamianum Miq. =Dalbergia monetaria 

Ecastophyllum glaucum Desv. =Dalbergia glauca 

Ecastophyllum monetaria (L.f.) DC. =Dalbergia monetaria 

Ecastophyllum pubescens DC. =Dalbergia subcymosa 

Ecastophyllum riedelii Radlk. =Dalbergia riedelii 

Eriosema flaviflorum Miq. =Eriosema violaceum 

Eriosema fusiforme Rusby =Eriosema crinitum var. fusiforme 

(Rusby) Grear [not in Guianas] 

Eriosema stipulare Benth. =Eriosema crinitum var. stipulare 

Erythrina corallodendrum L. [not in Guianas] 

Erythrina glauca Willd. =Erythrina fusca 

Etaballia guianensis Benth. =Etaballia dubia 

Galega cinerea L. =Tephrosia cinerea 

Galega toxicaria Swartz =Tephrosia sinapou 

Geoffroea discolor Benth. =Dussia discolor 

Geoffroea inermis Wright =Andira inermis 

Geoffroea pubescens Rich. =Andira surinamensis 

Geoffroea retusa (Lam.) Poir. =Andira surinamensis 

Geoffroea surinamensis Bondt =Andira surinamensis 

Geoffroea violacea (Aubl.) Pers. =Acouroa violacea 

Aubl.[=Dalbergia sp.?] 

Glycine crinita Kunth =Eriosema crinitum 

Glycine phaseoloides Swartz =Rhynchosia phaseoloides 

Glycine rufa Kunth =Eriosema rufum 

Glycine simplicifolia Kunth =Eriosema simplicifolium 

Hedysarum adscendens Swartz =Desmodium adscendens 

Hedysarum asperum Poir. =Desmodium asperum 

Hedysarum axillare Swartz =Desmodium axillare 

Hedysarum barbatum L. =Desmodium barbatum 

Hedysarum brasilianum Poir. =Aeschynomene brasiliana 

Hedysarum cajanifolium Kunth =Desmodium cajanifolium 

Hedysarum diphyllum L. =Zornia diphylla 

Hedysarum ecastaphyllum L. =Dalbergia ecastophyllum 

Hedysarum procumbens Mill. =Desmodium procumbens 

Indigofera anil L. =Indigofera suffruticosa 

Indigofera pascuorum Benth. =Indigofera lespedezioides 

Leptolobium costulatum Miq. =Ormosia costulata 

Leptolobium nitens Vogel =Acosmium nitens 

Leptolobium nitidum Miq. =Acosmium nitens 

Lonchocarpus ernestii Harms =Lonchocarpus margaritensis 

Lonchocarpus latifolius (Willd.) Kunth =Lonchocarpus 

heptaphyllus 

Lonchocarpus nitidus Spruce ex Benth. =Lonchocarpus 

floribundus 

Lonchocarpus pentaphyllus (Poir.) DC. =Lonchocarpus 

heptaphyllus 

Lonchocarpus pterocarpus DC. =Lonchocarpus scandens 

Machaerium aculeatum Raddi [not in Guianas] 

Machaerium ferrugineum Pers. =Machaerium quinata 

Machaerium multifoliolatum Ducke [not in Guianas] 

Machaerium oblongifolium Vogel [not in Guianas] 

Machaerium schomburgkii Benth. =Paramachaerium 

schomburgkii 

Machaerium surinamense Miq. =Machaerium leiophyllum 

Moutouchi suberosa Aubl. =Pterocarpus officinalis 

Mucuna altissima (Jacq.) DC. =Mucuna urens 

Muellera moniliformis L.f. =Muellera frutescens 

Neurocarpum guianense (Aubl.) Desv. =Clitoria guianensis 

Nissolia leiophylla DC. =Machaerium leiophyllum var. 

leiophyllum 

Nissolia polyphylla Poir. =Machaerium polyphyllum 

Nissolia quinata Aubl. =Machaerium quinata 

Ormosia costulata (Miq.) Kleinh. var. trifoliata (Huber) Amsh. 

=Ormosia costulata 

Ormosia costulata (Miq.) Kleinh. var. trifoliata (Huber) Amsh. 

=Ormosia costulata 

Ormosia fastigiata Tul. [not in Guianas] 

Ormosia trifoliata Huber =Ormosia costulata 

Ormosiopsis flava Ducke =Ormosia flava 

Phaseolus adenanthus G. Mey. =Vigna adenantha


Phaseolus campestris Benth. =Macroptilium 

longepedunculatum 

Phaseolus juruanus Harms =Vigna juruana 

Phaseolus lasiocarpus Mart. ex Benth. =Vigna lasiocarpa 

Phaseolus lathyroides L. =Macroptilium lathyroides 

Phaseolus linearis Kunth =Vigna linearis 

Phaseolus longipedunculatus Mart. ex Benth. =Macroptilium 

longepedunculatum 

Phaseolus peduncularis Kunth =Vigna peduncularis 

Phaseolus pilosus Kunth =Vigna lasiocarpa 

Phaseolus schottii Benth. =Vigna longifolia 

Phaseolus surinamensis Miq. =Vigna adenantha 

Phaseolus trichocarpus Wright =Vigna trichocarpa 

Phaseolus truxillensis Kunth =Vigna adenantha 

Phaseolus vexillatus L. =Vigna vexillata 

Pterocarpus amazonicus Huber =Pterocarpus santalinoides 

Pterocarpus draco L. =Pterocarpus officinalis 

Pterocarpus esculentus Schum. & Thonn. =Pterocarpus 

santalinoides 

Pterocarpus lunatus L.f. =Machaerium lunatum 

Pterocarpus magnicarpus Schery =Pterocarpus rohrii 

Pterocarpus rufescens Benth. =Pterocarpus rohrii 

Pterocarpus suberosa (Aubl.) Pers. =Pterocarpus officinalis 

Rhadinocarpus brasiliensis Vogel =Chaetocalyx brasiliensis 

Robinia nicou Aubl. =Lonchocarpus nicou 

Robinia panacoco Aubl. =Swartzia panacoco 

(Caesalpiniaceae) 

Robinia sepium Jacq. =Gliricidia sepium 

Sesbania aegyptica Poir. =Sesbania sesban 

Singana guianensis Aubl. [name of uncertain application] 

Sphinctolobium glaucescens Miq. =Muellera frutescens 

Spirotropis candollei Tul. =Spirotropis longifolia 

Stenolobium brachycarpum (Desv.) Benth. =Calopogonium 

mucunoides 

Stenolobium coeruleum Benth. =Calopogonium coeruleum 

Stenolobium velutinum Benth. =Calopogonium velutinum 

Sweetia nitens (Vogel) Benth. =Acosmium nitens 

Sweetia praeclara Sandw. =Acosmium praeclarum 

Tachigali purpurea Rich. =Diplotropis purpurea var. purpurea 

Tephrosia brevipes Benth. =Tephrosia sessiliflora 

Tephrosia toxicaria (Swartz) Pers. =Tephrosia sinapou 

Trifolium guianense Aubl. =Stylosanthes guianensis 

Vatairea surinamensis Kleinh. =Vatairea guianensis 

Vigna repens (L.) Kuntze =Vigna luteola 

Zornia marajoara Huber =Zornia guanipensis 

Zornia surinamensis Miq. =Zornia diphylla 

FLACOURTIACEAE 

contributed by R.W. Kiger (1992) 

REFERENCES: 

Sleumer, H.O. 1980. Flacourtiaceae. Flora 


Sleumer, H.O. & H. Uittien. 1935. 

Flacourtiaceae. In A. Pulle, Flora of 

Suriname 3(1): 282-303. 

Banara guianensis Aubl. [GU SU FG] 

Carpotroche crispidentata Ducke [FG] 

Carpotroche longifolia (Poepp.) Benth. [GU SU FG] 

SPERMATOPHYTA: Fabaceae - Flacourtiaceae 

109 

Carpotroche surinamensis Uittien [GU SU FG] 

Casearia aculeata Jacq. [GU SU] 

Casearia acuminata DC. [GU? SU FG] 

Casearia arborea (L.C. Rich.) Urb. [GU SU FG] 

Casearia combaymensis Tul. [GU SU FG] 

Casearia commersoniana Camb. [GU SU FG] 

Casearia decandra Jacq. [GU SU FG] 

Casearia grandiflora Camb. [GU SU FG] 

Casearia guianensis (Aubl.) Urb. [GU SU FG] 

Casearia javitensis Kunth [GU SU FG] 

Casearia mariquitensis Kunth [GU SU FG] 

Casearia negrensis Eichl. [SU FG] 

Casearia pitumba Sleumer [GU SU FG] 

Casearia resinifera Spruce ex Eichl. [FG] 

Casearia rusbyana Briquet [GU SU FG] 

Casearia spinescens (Swartz) Griseb. [GU] 

Casearia sylvestris Swartz [GU SU FG] 

Casearia ulmifolia Vahl ex Ventenat [GU SU FG] 

Casearia zizyphoides Kunth [GU FG] 

Euceraea nitida Mart. [GU SU] 

Hasseltia floribunda Kunth [FG?] 

Hecatostemon completus (Jacq.) Sleumer [GU] 

Homalium guianense (Aubl.) Oken [GU SU FG] 

Homalium racemosum Jacq. [GU SU FG] 

Laetia cupulata Spruce ex Benth. [GU] 

Laetia procera (Poepp.) Eichl. [GU SU FG] 

Laetia suaveolens (Poepp.) Benth. [GU] 

Lindackeria paludosa (Benth.) Gilg [GU FG?] 

Mayna odorata Aubl. [GU SU FG] 

Neoptychocarpus apodanthus (Kuhlmann) Buchheim [SU FG] 

Neoptychocarpus killipii (Monachino) Buchheim [FG] 

Neosprucea sp. [FG?] 

Prockia crucis P. Browne ex L. [to be expected in the Guianas] 

Ryania pyrifera (L.C. Rich.) Sleumer & Uittien [GU SU FG] 

Ryania speciosa Vahl [GU SU FG] 

Xylosma benthamii (Tul.) Triana & Planch. [GU SU FG] 

Xylosma ciliatifolium (Clos) Eichl. [GU FG?] 

Xylosma tessmannii Sleumer [GU FG] 

FLACOURTIACEAE - SYNONYMS AND 


Casearia arguta Kunth [not in Guianas] 

Casearia bracteifera Sagot =Casearia combaymensis 

Casearia cambessedesii Eichl. =Casearia ulmifolia 

Casearia cotticensis Uittien =Casearia ulmifolia 

Casearia densiflora Benth. =Casearia commersoniana 

Casearia dentata (Aubl.) Sessé & Moçino =Casearia 

commersoniana 

Casearia laurifolia Benth. =Casearia commersoniana 

Casearia macrophylla Vahl =Casearia pitumba 

Casearia nitida Jacq. [not in Guianas] 

Casearia petraea Benth. =Casearia ulmifolia 

Homalium densiflorum Spruce ex Benth. =Homalium 

guianense 

Homalium napimoga Spreng. =Homalium guianense 

Iroucana guianensis Aubl. =Casearia guianensis 

Laetia calophylla Eichl. =Laetia suaveolens 

Laetia casearioides Sagot ex Benth. =Laetia procera 

Laetia hirtella Kunth =Hecatostemon completus

110 


Laetia obtusifolia Eichl. ex Benoist [nomen nudum] =Laetia 

procera 

Mayna denticulata Benth. =Mayna odorata 

Napimoga guianensis Aubl. =Homalium guianense 

Patrisia parviflora DC. =Ryania speciosa 

Patrisia pyrifera L.C. Rich. =Ryania pyrifera 

Patrisia speciosa (Vahl.) Monachino =Ryania speciosa 

Patrisia spruceana (Monachino) Lemée =Ryania spruceana 


Piparea dentata Aubl. =Casearia commersoniana 

Pitumba guianensis Aubl. =Casearia pitumba 

Ptychocarpus apodanthus Kuhlm. =Neoptychocarpus 

apodanthus 

Racoubea guianensis Aubl. =Homalium guianense 

Ryania pyrifera (L.C. Rich.) Sleumer & Uittien var. tomentosa 

(Miq.) Sleumer =Ryania speciosa 

Ryania sagotiana Eichl. =Ryania pyrifera 

Xylosma nitidum (Hellenius) A. Gray ex Griseb. [not in 


GENTIANACEAE 

contributed by M.J. Jansen-Jacobs & P.J.M. Maas 

(1992) 

REFERENCES: 

Jonker, F.P. 1936. Gentianaceae. In A. Pulle, 


Maas, P.J.M. 1985. Nomenclatural notes on 

neotropical Lisyantheae. Proc. Kon. Ned. 

Akad. Wetensch. (ser. c) 88(4): 405-412. 

Maas, P.J.M., S. Nilsson, A.M.C. Hollants, B.J.H. 

ter Welle, H. Persoon, & E.C.H. van Heusden. 

1984. Systematic studies in neotropical 

Gentianaceae - the Lisianthus complex. Acta 

Bot. Neerl. 32: 371-374. 

Maguire, B. 1981. Gentianaceae. In B. Maguire 


Highland part 11. Mem. N.Y. Bot. Gard. 32: 

1-391. 

Maguire, B. & B.M. Boom. 1989. Gentianaceae 

(part 3). In B. Maguire & collaborators, 


Memoirs of the New York Bot. Garden 51: 1- 

127. 

Maas, P.J.M. & P. Ruyters. 1986. Voyria and 

Voyriella (Saprophytic Gentianaceae). Flora 


Celiantha imthurniana (Oliver) Maguire [GU] 

Chorisepalum psychotrioides Ewan var. psychotrioides [GU] 

Coutoubea humilis Sandw. [GU] 

Coutoubea ramosa Aubl. [GU SU FG] 

Coutoubea reflexa Benth. [GU] 

Coutoubea spicata Aubl. [GU SU FG] 

Curtia tenuifolia (Aubl.) Knobl. var. tenella (Mart.) Grothe & 

Maas [FG] 

Curtia tenuifolia (Aubl.) Knobl. var. tenuifolia [GU SU? FG] 

Irlbachia alata (Aubl.) Maas subsp. alata [GU SU FG] 

Irlbachia alata (Aubl.) Maas subsp. angustifolia (Kunth) 

Persoon & Maas [GU] 

SPERMATOPHYTA: Flacourtiaceae - Gentianaceae 

Irlbachia alata (Aubl.) Maas subsp. longistyla Persoon & Maas 

[GU FG] 

Irlbachia alata (Aubl.) Maas subsp. viridiflora (Mart.) Persoon 

& Maas [GU] 

Irlbachia caerulescens (Aubl.) Griseb. [GU SU FG] 

Irlbachia nemorosa (Willd. ex Roem. & Schult.) Merrill [GU] 

Irlbachia purpurascens (Aubl.) Maas [GU SU FG] 

Irlbachia quelchii (N.E. Brown) Maas [GU] 

Irlbachia tatei (Gleason) Maguire [GU] 

Macrocarpaea sp. [GU] [Mt. Ayanganna] 

Neurotheca loeselioides (Spruce ex Progel) Baill. subsp. 

loeseloides [GU SU] 

Schultesia benthamiana Klotzsch ex Griseb. [GU SU?] 

Schultesia brachyptera Cham. [GU SU FG] 

Schultesia guianensis (Aubl.) Malme [GU SU FG] 

Schultesia pohliana Progel [GU SU] 

Schultesia subcrenata Klotzsch ex Griseb. [GU] 

Symbolanthus calygonus (Ruiz & Pav.) Griseb. s.l. [GU] 

Tachia gracilis Benth. [GU] 

Tachia grandiflora Maguire & Weaver [FG] 

Tachia guianensis Aubl. [GU SU FG] 

Tachia schomburgkiana Benth. [GU] 

Tachia smithii Maguire & Weaver [GU?] 

Tachia sp. nov. [SU] 

Tapeinostemon spenneroides Benth. [GU SU] 

Voyria aphylla (Jacq.) Pers. [GU SU FG] 

Voyria aurantiaca Splitg. [GU SU FG] 

Voyria caerulea Aubl. [GU SU FG] 

Voyria clavata Splitg. [GU SU FG] 

Voyria corymbosa Splitg. subsp. alba (Standl.) Ruyters & 

Maas [GU SU FG] 

Voyria corymbosa Splitg. subsp. corymbosa [SU FG] 

Voyria flavescens Griseb. [SU FG] 

Voyria pittieri (Standl.) L.O. Williams [GU] 

Voyria rosea Aubl. [GU SU FG] 

Voyria spruceana Benth. [GU SU FG] 

Voyria tenella Hook. [GU SU FG] 

Voyria tenuiflora Griseb. [GU SU FG] 

Voyriella parviflora (Miq.) Miq. [GU SU FG] 

GENTIANACEAE - SYNONYMS AND 


Calolisianthus imthurnianus (Oliver) Gleason =Celiantha 

imthurniana 

Calolisianthus tatei Gleason =Irlbachia tatei 

Chelonanthus alatus (Aubl.) Pulle =Irlbachia alata subsp. alata 

Chelonanthus angustifolius (Kunth) Gilg =Irlbachia alata 

subsp. angustifolia 

Chelonanthus chelonoides (L.f.) Gilg =Irlbachia alata subsp. 

alata 

Chelonanthus schomburgkii (Griseb.) Gilg =Irlbachia alata 

subsp. alata 

Chelonanthus uliginosus Griseb. =Irlbachia purpurascens 

Coutoubea ramosa Aubl. var. longifolia Benth. =Coutoubea 

ramosa 

Coutoubea ramosa Aubl. var. racemosa (G. Mey.) Benth. 

=Coutoubea ramosa 

Exacum guianense Aubl. =Schultesia guianensis 

Exacum tenuifolium Aubl. =Curtia tenuifolia subsp. tenuifolia


Leiothamnus elisabethae Schomb. =Symbolanthus calygonus 

Leiphamos calycina (Splitg. ex A. DC.) Miq. =Voyria caerulea 

Leiphamos leucantha (Miq.) Gilg. =Voyria tenuiflora 

Leiphamos montana Jonker =Voyria corymbosa subsp. alba 

Leiphamos obconica Auct., non (Progel) Gilg. =Voyria tenella 

Lisianthus alatus Aubl. =Irlbachia alata subsp. alata 

Lisianthus caerulescens Aubl. =Irlbachia caerulescens 

Lisianthus chelonoides L.f. =Irlbachia alata subsp. alata 

Lisianthus elisabethae (Schomb.) Griseb. =Symbolanthus 

calygonus 

Lisianthus fistulosus Poir. =Irlbachia alata subsp. alata 

Lisianthus grandiflorus Aubl. =Irlbachia alata subsp. alata 

Lisianthus imthurnianus Oliver =Celiantha imthurniana 

Lisianthus purpurascens Aubl. =Irlbachia purpurascens 

Lisianthus quelchii N.E. Brown =Irlbachia quelchii 

Lisianthus uliginosus Griseb. =Irlbachia purpurascens 

Macrocarpaea quelchii (N.E. Brown) Ewan =Irlbachia quelchii 

Rogersanthus quelchii (N.E. Brown) Maguire & Boom 

=Irlbachia quelchii 

Schultesia schomburgkiana Progel =Schultesia benthamiana 

Stahelia spenneroides (Benth.) Jonker =Tapeinostemon 

spenneroides 

Stahelia surinamensis Jonker =Tapeinostemon spenneroides 

Symbolanthus elisabethae (Schomb.) Gilg =Symbolanthus 

calygonus 

Symbolanthus quelchii (N.E. Brown) Gleason =Irlbachia 

quelchii 

Tapeinostemon surinamensis (Jonker) Steyerm. 

=Tapeinostemon spenneroides 

Tetrapollinia caerulescens (Aubl.) Maguire & Boom 

=Irlbachia caerulescens 

GESNERIACEAE 

reviewed by C. Feuillet & L.E. Skog (1992); 

reviewed by L.E. Skog & C. Feuillet (1996) 

REFERENCES: 

Leeuwenberg, A.J.M. 1958. The Gesneriaceae 

of Guiana. Acta Bot. Neerl. 7: 291-444. 

Leeuwenberg, A.J.M. 1984. Gesneriaceae. In 

A.L. Stoffers & J.C. Lindeman, Flora of 

Suriname 5(1): 592-650. 

Morton, C.V. 1939. A revision of Besleria. 

Contr. U.S. Nat. Herb. 26: 395-474. 

Wiehler, H.J. 1978. The genera Episcia, 

Alsobia, Nautilocalyx, and Paradrymonia. 

Selbyana 5: 11-60. 

Alloplectus cristatus (L.) Mart. var. epirotes Leeuwenberg 

[GU] 

Alloplectus savannarum C. Morton [GU SU] 

Besleria flavo-virens Nees & Mart. [GU SU FG] 

Besleria insolita C. Morton [GU FG] 

Besleria laxiflora Benth. [GU SU FG] 

Besleria maasii Wiehler [FG] 

Besleria patrisii DC. [FG] 

Besleria penduliflora Fritsch [GU] 

Besleria saxicola C. Morton [GU] 

Besleria verecunda C. Morton [GU SU FG] 

Chrysothemis pulchella (Donn ex Sims) Decne. [GU SU FG] 

Chrysothemis rupestris (Benth.) Leeuwenberg [GU SU] 

SPERMATOPHYTA: Gentianaceae - Gesneriaceae 

111 

Chrysothemis villosa (Benth.) Leeuwenberg [GU] 

Codonanthe calcarata (Miq.) Hanst. [GU SU FG] 

Codonanthe crassifolia (Focke) C. Morton [GU SU FG] 

Codonanthopsis dissimulata (H.E. Moore) Wiehler [GU FG] 

Columnea calotricha J.D. Smith [SU FG] 

Columnea guianensis C. Morton [GU] 

Columnea oerstediana Klotzsch ex Oersted [FG] 

Columnea sanguinea (Pers.) Hanst. [GU? SU FG] 

Corytoplectus deltoideus (C. Morton) Wiehler [GU] 

Drymonia antherocycla Leeuwenberg [SU FG] 

Drymonia coccinea (Aubl.) Wiehler [GU SU FG] 

Drymonia psilocalyx Leeuwenberg [SU FG] 

Drymonia serrulata (Jacq.) Mart. [GU SU FG] 

Episcia reptans Mart. [GU SU FG] 

Episcia sphalera Leeuwenberg [SU FG] 

Episcia xantha Leeuwenberg [FG] 

Gloxinia perennis (L.) Fritsch [GU SU FG] [cultivated or 


Gloxinia purpurascens (Rusby) Wiehler [GU FG] 

Kohleria hirsuta (Kunth) Regel [GU SU FG] 

Lembocarpus amoenus Leeuwenberg [SU FG] 

Napeanthus jelskii Fritsch [FG] 

Napeanthus macrostoma Leeuwenberg [SU FG] 

Nautilocalyx bryogeton (Leeuwenberg) Wiehler [GU] 

Nautilocalyx cordatus (Gleason) L.E. Skog [GU] 

Nautilocalyx kohlerioides (Leeuwenberg) Wiehler [FG] 

Nautilocalyx mimuloides (Benth.) C. Morton [GU FG] 

Nautilocalyx pallidus (Sprague) Sprague [FG] 

Nautilocalyx pictus (Hook.) Sprague [GU SU FG] 

Nautilocalyx porphyrotrichus (Leeuwenberg) Wiehler [GU] 

Nautilocalyx villosus (Kunth & Bouché) Sprague [GU SU? 

FG?] 

Paradrymonia campostyla (Leeuwenberg) Wiehler [SU FG] 

Paradrymonia ciliosa (Mart.) Wiehler [GU SU FG] 

Paradrymonia densa (C.H. Wright) Wiehler [GU SU FG] 

Paradrymonia maculata (J.D. Hook.) Wiehler [GU FG] 

Rhoogeton cyclophyllus Leeuwenberg [GU] 

Rhoogeton viviparus Leeuwenberg [GU] 

Sinningia incarnata (Aubl.) D. Denham [GU SU FG] 

Sinningia schomburgkiana (Kunth & Bouché) Chautems [GU] 

Tylopsacas cuneatum (Gleason) Leeuwenberg [GU] 

GESNERIACEAE - SYNONYMS AND 


Alloplectus calotrichus (J.D. Smith) Stearn =Columnea 

calotricha 

Alloplectus coccineus (Aubl.) Mart. =Drymonia coccinea 

Alloplectus meridensis Klotzsch ex Hanst. [not in Guianas] 

Alloplectus patrisii DC. =Drymonia coccinea 

Besleria bivalvis L.f. =Mendoncia bivalvis (L.f.) Merr. 

(Mendonciaceae) 

Besleria coccinea Aubl. =Drymonia coccinea 

Besleria incarnata Aubl. =Sinningia incarnata 

Besleria lutea L. [not in Guianas] 

Besleria violacea Aubl. =Schlegelia violacea (Bignoniaceae) 

Centrosolenia densa (C.H. Wright) Sprague =Paradrymonia 

densa

112 


Centrosolenia hirsuta Benth. =Nautilocalyx cordatus 

Codonanthe bipartita L.B. Smith =Codonanthe calcarata 

Codonanthe carnosa Hanst. [not in Guianas] 

Codonanthe confusa Sandw. =Codonanthe crassifolia 

Codonanthopsis ulei Fritsch [not in Guianas] 

Columnea aureonitens Hook. =Columnea sanguinea 

Columnea scandens L. [not in Guianas] 

Crantzia patrisii (DC.) Fritsch =Drymonia coccinea 

Drymonia calcarata Mart. =Drymonia serrulata 

Drymonia campostyla Leeuwenberg =Paradrymonia 

campostyla 

Drymonia cristata Miq. =Drymonia serrulata 

Drymonia sarmentosula Lem. =Drymonia serrulata 

Drymonia turrialvae Hanst. [not in Guianas] 

Episcia bryogeton Leeuwenberg =Nautilocalyx bryogeton 

Episcia ciliosa (Mart.) Hanst. =Paradrymonia ciliosa 

Episcia cordata Gleason =Nautilocalyx cordatus 

Episcia densa C.H. Wright =Paradrymonia densa 

Episcia hirsuta (Benth.) Hanst. =Nautilocalyx cordatus 

Episcia maculata J.D. Hook. =Paradrymonia maculata 

Episcia mimuloides Benth. =Nautilocalyx mimuloides 

Episcia porphyrotricha Leeuwenberg =Nautilocalyx 

porphyrotrichus 

Gesneria schomburgkiana Kunth & Bouché =Sinningia 


Gloxinia trichantha Miq. =Gloxinia perennis 

Hypocyrta crassifolia Focke =Codonanthe crassifolia 

Nautilocalyx adenosiphon (Leeuwenberg) Wiehler [not in 


Nautilocalyx lacteus Sandw. =Nautilocalyx pictus 

Nematanthus calcaratus Miq. =Codonanthe calcarata 

Rechsteineria incarnata (Aubl.) Leeuwenberg =Sinningia 

incarnata 

Rechsteineria schomburgkiana (Kunth & Bouché) 

Leeuwenberg =Sinningia schomburgkiana 

Tussacia rupestris Benth. =Chrysothemis rupestris 

Tylosperma cuneatum (Gleason) Leeuwenberg =Tylopsacas 

cuneatum 

GNETACEAE 

contributed by D. Stevenson (1992) 

REFERENCES: 

Markgraf, F. 1965. New discoveries of Gnetum 

in tropical America. Ann. Missouri Bot. 

Gard. 52: 379-386. 

Stevenson, D. & T. Zanoni. 1991. Gnetaceae. In 


(series A) 9: 12-18. 

Gnetum camporum (Markgr.) D. Stevenson & T. Zanoni [to be 


Gnetum leyboldii Tul. [to be expected in the Guianas] 

Gnetum nodiflorum Brongn. [GU SU FG] 

Gnetum paniculatum Spruce ex Benth. [GU FG] 

Gnetum urens (Aubl.) Blume [GU SU FG] 

GNETACEAE - SYNONYMS AND 


Gnetum cruzianum Gleason =Gnetum nodiflorum 

SPERMATOPHYTA: Gesneriaceae - Haloragaceae 

Gnetum melinonii Benoist =Gnetum urens 

Gnetum thoa Brongn. =Gnetum urens 

Gnetum urens (Aubl.) Blume var. camporum Markgr. 

=Gnetum camporum 

Thoa urens Aubl. =Gnetum urens 

HAEMODORACEAE 

contributed by P.J.M. Maas (1992); additions and 

corrections by P.J.M. Maas (1996) 

REFERENCES: 

Lanjouw, J. 1938. Haemodoraceae. In A. Pulle, 



Haemodoraceae. Flora Neotropica Monogr. 

61: 1-44. 


Haemodoraceae. In A.R.A. Görts-van Rijn, 


Villarroel, F. 1971. Haemodoraceae. In T. 

Lasser, Flora de Venezuela 3(1): 207-222. 

Schiekia orinocensis (Kunth) Meissn. subsp. orinocensis [GU 

SU] 

Schiekia orinocensis (Kunth) Meissn. subsp. silvestris Maas & 

Stoel [GU SU FG] 

Xiphidium caeruleum Aubl. [GU SU FG] 

HAEMODORACEAE - SYNONYMS AND 


Schiekia orinocensis (Kunth) Meissn. subsp. savannarum 

Maguire & Wurdack =Schiekea orinocensis subsp. 

orinocensis 

Troschelia orinocensis (Kunth) Klotzsch & M.R. Schomb. 

[nomen nudum] =Schiekia orinocensis 

Xiphidium fockeanum Miq. =Xiphidium caeruleum 

REFERENCES: 

HALORAGACEAE 

Orchard, A.E. 1981. A revision of South 

American Myriophyllum, and its 

repercussions on some Australian and North 

American species. Brunonia 4: 27-65. 

Laurembergia tetrandra (Schott) Kanitz subsp. tetrandra [to be 


Myriophyllum aquaticum (Vell.) Verdcourt [GU? FG?] 

HALORAGACEAE - SYNONYMS AND 

MISIDENTIFICATIONS:


Myriophyllum proserpinacoides Gillies ex Hook. & Arnott 

=Myriophyllum aquaticum 

HELICONIACEAE 


P.J.M. Maas & W.J. Kress (1996) 

REFERENCES: 

Berry, F. & W.J. Kress. 1991. Heliconia: An 

Identification Guide. Washington, DC: 

Smithsonian Institution Press. 

Maas, P.J.M. 1985. Musaceae. In A.R.A. Görtsvan 

Rijn, Flora of the Guianas (series A) 1: 1- 

28. 

Maas, P.J.M. & M.J.M. de Rooij. 1979. 

Musaceae. In A.L. Stoffers & J.C. Lindeman, 


Heliconia acuminata L.C. Rich. subsp. acuminata [GU SU 

FG] 

Heliconia aemygdiana Burle-Marx [FG] 

Heliconia bihai (L.) L. [GU SU FG] 

Heliconia chartacea Lane ex Barreiros [GU SU FG] 

Heliconia dasyantha Koch & Bouché [SU FG] 

Heliconia densiflora B. Verl. subsp. angustifolia L. Andersson 

[GU] 

Heliconia densiflora B. Verl. subsp. densiflora [SU FG] 

Heliconia episcopalis Vell. [SU] 

Heliconia hirsuta L.f. [GU SU FG] 

Heliconia lourteigiae Mello & E. Santos [GU SU FG] 

Heliconia marginata (Griggs) Pittier [GU SU FG?] 

Heliconia pendula Wawra [GU FG] 

Heliconia psittacorum L.f. [GU SU FG] 

Heliconia richardiana Miq. [GU SU FG] 

Heliconia spathocircinata Aristeguieta [GU SU FG] 

Heliconia stricta Huber [GU SU] 

Heliconia xnickeriensis Maas & de Rooij [GU SU] 

HELICONIACEAE - SYNONYMS AND 


Bihai silvestris Gleason =Heliconia psittacorum 

Heliconia ballia L.C. Rich. =Heliconia psittacorum 

Heliconia brasiliensis Hook. =Heliconia farinosa Raddi [not in 


Heliconia brasiliensis Hook. var. concolor O.G. Petersen 

=Heliconia psittacorum 

Heliconia cannoidea L.C. Rich. =Heliconia psittacorum 

Heliconia glauca Poit. ex B. Verl. =Heliconia richardiana 

Heliconia hirsuta L.f. var. cannoidea (L.C. Rich.) Baker 

=Heliconia psittacorum 

Heliconia humilis (Aubl.) Jacq. =Heliconia psittacorum 

Heliconia psittacorum L.f. var. schomburgkiana (Klotzsch) 

Baker =Heliconia psittacorum 

Heliconia psittacorum L.f. var. spathacea Eichler ex O.G. 

Petersen =Heliconia psittacorum 

Heliconia schomburgkiana Klotzsch =Heliconia psittacorum 

Heliconia silvestris (Gleason) L.B. Smith =Heliconia 

psittacorum 

REFERENCES: 

HERNANDIACEAE 

SPERMATOPHYTA: Haloragaceae - Hippocrateaceae 

113 

Kostermans, A.J.G.H. 1937. Hernandiaceae. In 


Kubitzki, K.U. 1969. Monographie der 

Hernandiaceen. Bot. Jahrb. Syst. 89: 78-205. 

Kubitzki, K.U. 1976. Hernandiaceae (additions 



Kubitzki, K.U. 1982. Hernandiaceae. In Z. 

Luces de Febres, Flore de Venezuela 4(2): 

317-324. 

Hernandia guianensis Aubl. [GU SU FG] 

Sparattanthelium aruakorum Tutin [GU SU] 

Sparattanthelium guianense Sandw. [GU FG?] 

Sparattanthelium uncigerum (Meissn.) Kubitzki [GU SU FG] 

Sparattanthelium wonotoboense Kosterm. [GU SU] 

HERNANDIACEAE - SYNONYMS AND 


Hernandia sonora L. [not in Guianas] 

Sparattanthelium botocudorum Mart. [not in Guianas] 

Sparattanthelium botocudorum Mart. var. uncigerum Meissn. 

=Sparattanthelium uncigerum 

Sparattanthelium macusiorum A.C. Smith =Sparattanthelium 

wonotoboense 

Sparattanthelium melinonis Baill. ex Lemée [nomen nudum] 

=Sparattanthelium uncigerum 

HIPPOCRATEACEAE 

contributed by A.R.A. Görts-van Rijn (1992); 

additions and corrections by A.R.A. Görtsvan 

Rijn (1996) 

REFERENCES: 

Görts-van Rijn, A.R.A. & A.M.W. Mennega. 

1994. Hippocrateaceae. In A.R.A. Görts-van 


157. 

Loesener, T. 1942. Hippocrateaceae. In A. 

Engler & K. Prantl, Die Natürlichen 

Pflanzenfamilien 20b: 198-231. 


Hippocrateaceae. Brittonia 3: 341-355. 

Anthodon decussatum Ruiz & Pav. [FG] 

Cheiloclinium anomalum Miers [GU SU] 

Cheiloclinium belizense (Standl.) A.C. Smith [GU FG] 

Cheiloclinium cognatum (Miers) A.C. Smith [GU SU FG] 

Cheiloclinium diffusiflorum (Miers) A.C. Smith [GU SU FG]

114 


Cheiloclinium hippocrateoides (Peyr.) A.C. Smith [GU SU 

FG] 

Cheiloclinium minutiflorum (A.C. Smith) A.C. Smith [FG] 

Cuervea kappleriana (Miq.) A.C. Smith [GU SU FG] 

Elachyptera floribunda (Benth.) A.C. Smith [GU SU FG] 

Hippocratea volubilis L. [GU SU FG] 

Hylenaea comosa (Swartz) Miers [GU SU FG] 

Hylenaea praecelsa (Miers) A.C. Smith [to be expected in the 


Hylenaea unguiculata A.M. Mennega [SU] 

Peritassa compta Miers [GU] 

Peritassa huanucana (Loes.) A.C. Smith [GU SU FG] 

Peritassa laevigata (Hoffmanns. ex Link) A.C. Smith [GU SU 

FG?] 

Prionostemma aspera (Lam.) Miers [GU SU FG] 

Pristimera caudata A.M. Mennega [SU] 

Pristimera holdeniana (A.C. Smith) A.C. Smith [GU] 

Pristimera nervosa (Miers) A.C. Smith [GU SU FG] 

Pristimera tenuiflora (Mart. ex Peyr.) A.C. Smith [SU FG] 

Pristimera verrucosa (Kunth) Miers [GU] 

Salacia amplectens A.C. Smith [GU SU FG] 

Salacia cordata (Miers) A.M. Mennega [SU?] 

Salacia duckei A.C. Smith [SU? FG] 

Salacia elliptica (Mart.) G. Don [GU SU FG] 

Salacia impressifolia (Miers) A.C. Smith [GU SU FG] 

Salacia insignis A.C. Smith [SU FG] 

Salacia juruana Loes. [GU SU FG] 

Salacia kanukuensis A.C. Smith [GU SU] 

Salacia maburensis A.M. Mennega [GU] 

Salacia miqueliana Loes. [SU FG] 

Salacia multiflora (Lam.) DC. subsp. mucronata (Rusby) A.M. 

Mennega [SU FG] 

Salacia multiflora (Lam.) DC. subsp. multiflora [GU SU FG] 

Salacia pachyphylla (Miers) Peyr. [GU] 

Tontelea attenuata Miers [GU FG] 

Tontelea coriacea A.C. Smith [GU SU FG] 

Tontelea cylindrocarpa (A.C. Smith) A.C. Smith [SU FG] 

Tontelea glabra A.C. Smith [GU SU FG] 

Tontelea laxiflora (Benth.) A.C. Smith [GU SU FG] 

Tontelea nectandrifolia (A.C. Smith) A.C. Smith [GU SU 

FG?] 

Tontelea ovalifolia (Miers) A.C. Smith subsp. ovalifolia [GU 

FG] 

Tontelea ovalifolia (Miers) A.C. Smith subsp. richardii (Peyr.) 

Görts & A.M. Mennega [FG] 

Tontelea sandwithii A.C. Smith [GU FG] 

HIPPOCRATEACEAE - SYNONYMS AND 


Amphizoma laxiflora (Benth.) Miers =Tontelea laxiflora 

Anthodon laxiflorum Benth. =Tontelea laxiflora 

Cheiloclinium gleasonianum (A.C. Smith) A.C. Smith 

=Cheiloclinium hippocrateoides 

Cheiloclinium jenmanii A.C. Smith =Cheiloclinium belizense 

Cheiloclinium lineolatum (A.C. Smith) A.C. Smith 

=Cheiloclinium cognatum 

Cheiloclinium parviflorum (Miers) A.C. Smith 

=Cheiloclinium diffusiflorum 

Cheiloclinium podostemmum (Sandw.) A.C. Smith 

=Cheiloclinium belizense 

Hippocratea aggregata Peyr. =Cuervea kappleriana 

Hippocratea ambigua Peyr. [insufficiently known taxon] 

Hippocratea aspera Lam. =Prionostemma aspera 

Hippocratea comosa Swartz =Hylenaea comosa 

SPERMATOPHYTA: Hippocrateaceae 

Hippocratea emarginata Rudge =Salacia multiflora subsp. 

multiflora 

Hippocratea flava Gleason =Cuervea kappleriana 

Hippocratea holdeniana A.C. Smith =Pristimera holdeniana 

Hippocratea kappleriana Miq. =Cuervea kappleriana 

Hippocratea lutea Gleason =Cuervea kappleriana 

Hippocratea malpighiifolia Rudge =Prionostemma aspera 

Hippocratea multiflora Lam. =Salacia multiflora 

Hippocratea obovata L.C. Rich. =Salacia multiflora subsp. 

multiflora 

Hippocratea schomburgkii Klotzsch [nomen nudum] 

=Prionostemma aspera 

Hippocratea tenuiflora Mart. ex Peyr. =Pristimera tenuiflora 

Hylenaea capilliflora Miers =Hylenaea comosa 

Peritassa cuneata A.M. Mennega =Peritassa huanucana 

Peritassa granulata (Urb.) A.C. Smith =Peritassa laevigata 

Prionostemma malpighiifolia (Rudge) Miers =Prionostemma 

aspera 

Pristimera apiculata Miers =Cuervea kappleriana 

Pristimera caribaea (Urb.) A.C. Smith [not in Guianas] 

Raddia lacunosa Miers =Salacia elliptica 

Raddia pachyphylla Miers =Salacia pachyphylla 

Salacia anomala (Miers) Peyr. =Cheiloclinium anomalum 

Salacia gleasoniana A.C. Smith =Cheiloclinium 

hippocrateoides 

Salacia guianensis Klotzsch [nomen nudum] =Salacia elliptica 

Salacia guianensis (Aubl.) Lemée =Macahanea guianensis 

Aubl.[incertae sedis; excluded from Hippocrateaceae by 

A.C. Smith] 

Salacia hippocrateoides Peyr. =Cheiloclinium hippocrateoides 

Salacia lacunosa (Miers) Peyr. =Salacia elliptica 

Salacia laevigata (Hoffmanns. ex Link) DC. =Peritassa 

laevigata 

Salacia laevigata (Hoffmanns. ex Link) DC. subsp. obtusa 

Peyr. =Peritassa compta 

Salacia laxiflora (Benth.) Peyr. =Tontelea laxiflora 

Salacia laxiflora (Benth.) Peyr. var. amygdalifolia Peyr. 

=Tontelea laxiflora 

Salacia macrantha A.C. Smith [not in Guianas] 

Salacia macrophylla Miq. =Salacia miqueliana 

Salacia mucronata Rusby =Salacia multiflora subsp. 

mucronata 

Salacia obovata (L.C. Rich.) Peyr. =Salacia multiflora subsp. 

multiflora 

Salacia obovata (L.C. Rich.) Peyr. var. guianensis Peyr. 

=Salacia multiflora subsp. multiflora 

Salacia parviflora Sagot ex Peyr. =Cheiloclinium 

diffusiflorum 

Salacia podostemma Sandw. =Cheiloclinium belizense 

Salacia richardii Peyr. =Tontelea ovalifolia subsp. richardii 

Salacia rugulosa Sagot [nomen nudum]=Salacia multiflora 

subsp. multiflora 

Salacia scabra DC. =Prionostemma aspera 

Salacia scandens (Aubl.) Griseb. =Elachyptera floribunda? 

Thermophila emarginata (Rudge) Miers =Salacia multiflora 


Thermophila macrophylla (Miq.) Miers =Salacia miqueliana 

Thermophila obovata (L.C. Rich.) Miers =Salacia multiflora 


Thermophila rugulosa Miers =Salacia multiflora subsp. 

multiflora


Tonsella malpighiifolia (Rudge) G. Mey. =Prionostemma 

aspera 

Tontelea aubletiana Miers =Tontelea sp. 

Tontelea fluminensis (Peyr.) A.C. Smith =Tontelea ovalifolia 

subsp. richardii 

Tontelea parviflora Miers =Cheiloclinium diffusiflorum 

Tontelea richardii (Peyr.) A.C. Smith =Tontelea ovalifolia 

subsp. richardii 

Tontelea scandens Aubl. =Elachyptera floribunda? 

HUGONIACEAE 

reviewed by D. Sabatier (1992) 

REFERENCES: 

Bakhuizen van den Brink f., R.C. 1941. 

Linaceae. In A. Pulle, Flora of Suriname 

3(1): 409-411. 

Secco, R. de S. & S.M.B. Silva. 1990. Materiaes 

para a flora amazônica - VIII. Contribuçao à 

sistemática das Linaceae da Amazônia 

Brasileira. Bol. Mus. Paraense Emílio Goeldi 

(Bot.) 6(1): 113-135. 

Hebepetalum humiriifolium (Planch.) Benth. [GU SU FG] 

Roucheria calophylla Planch. [GU SU? FG?] 

Roucheria laxiflora Winkl. [SU FG] 

Roucheria schomburgkii Planch. [GU] 

HUGONIACEAE - SYNONYMS AND 


Roucheria humeriifolia Planch. =Hebepetalum humiriifolium 

HUMIRIACEAE 


REFERENCES: 

Bakhuizen van den Brink f., R.C. 1941. 

Humiriaceae. In A. Pulle, Flora of Suriname 

3(1): 412-421. 

Cuatrecasas, J. 1961. A taxonomic revision of 

the Humiriaceae. Contrib. U.S. Nat. Herb. 35: 

25-214. 

Sabatier, D. 1987. Studies on the Flora of the 

Guianas 12. Quelques nouveautés chez les 

Humiriaceae. Proc. Kon. Ned. Akad. Wet. 

ser. C, 90(2): 203-210. 

Endopleura uchii (Huber) Cuatrec. [GU?] 

Humiria balsamifera (Aubl.) St. Hil. var. balsamifera [GU SU 

FG] 

Humiria balsamifera (Aubl.) St. Hil. var. coriacea Cuatrec. 

[GU SU] 

Humiria balsamifera (Aubl.) St. Hil. var. floribunda (Mart.) 

Cuatrec. [GU SU FG] 

Humiria balsamifera (Aubl.) St. Hil. var. guianensis (Benth.) 

Cuatrec. [GU SU] 

Humiria balsamifera (Aubl.) St. Hil. var. imbaimadaiensis 

Cuatrec. [GU] 

Humiria balsamifera (Aubl.) St. Hil. var. laurina (Urb.) 

Cuatrec. [GU] 

Humiria balsamifera (Aubl.) St. Hil. var. savannarum 

(Gleason) Cuatrec. [GU] 

Humiria crassifolia Mart. ex Urb. var. acutipetala Cuatrec. 

[GU] 

Humiria crassifolia Mart. ex Urb. var. crassifolia [GU] 

Humiriastrum excelsum (Ducke) Cuatrec. [FG] 

Humiriastrum obovatum (Benth.) Cuatrec. [GU] 

Humiriastrum subcrenatum (Benth.) Cuatrec. [FG] 

Sacoglottis amazonica Mart. [GU FG?] 

Sacoglottis cydonioides Cuatrec. [GU SU FG] 

Sacoglottis guianensis Benth. var. guianensis [GU SU FG] 

Sacoglottis guianensis Benth. var. hispidula Cuatrec. [GU? 

SU] 

Schistostemon densiflorum (Benth.) Cuatrec. [GU SU] 

Schistostemon dichotomum (Urb.) Cuatrec. [SU FG] 

Schistostemon sylvaticum Sabatier [FG] 

Vantanea guianensis Aubl. [GU SU FG] 

Vantanea minor Benth. [GU] 

Vantanea parviflora Lam. var. parviflora [SU FG] 

HUMIRIACEAE - SYNONYMS AND 


Houmiri balsamifera Aubl. =Humiria balsamifera 

Humiria floribunda Mart. =Humiria balsamifera var. 

floribunda 

Humiria subcrenata Urb. =Humiriastrum subcrenatum 

Humirium guianense Benth. =Humiria balsamifera var. 

guianensis 

Sacoglottis densiflora (Benth.) Urb. =Schistostemon 

densiflorum 

Sacoglottis dichotoma Urb. =Schistostemon dichotomum 

Sacoglottis kaboeriensis Bakh. v.d. Brink f. =Schistostemon 

densiflorum 

Sacoglottis mattogrossensis Malme [not in Guianas; 

specimens represent a new species (fide Sabatier)] 

Sacoglottis obovata (Benth.) Urb. =Humiriastrum obovatum 

REFERENCES: 

HYDROCHARITACEAE 

115 

Cook, C.D. 1985. A revision of the genus 

Apalanthe. Aquatic Bot. 21: 157-164. 

Cook, C.D. & K. Urmi-König, 1983. A revision 

of the genus Limnobium including 

Hydromystria. Aquatic Bot. 17: 1-27. 

Jonker, F.P. 1943. Hydrocharitaceae. In A. 


Lowden, R.M. 1992. Floral variation and 

taxonomy of Limnobium L.C. Richard. 

Rhodora 94: 111-134. 

Phillips, R.C. & E.G. Meñez. 1988. Seagrasses. 

Smiths. Contrib. Marine Sci. 34: 1-104. 

SPERMATOPHYTA: Hippocrateaceae - Hydrocharitaceae

116 


Apalanthe granatensis (Humb. & Bonpl.) Planch. [GU SU FG] 

Limnobium spongia (Bosc) Steud. subsp. laevigatum (Humb. 

& Bonpl. ex Willd.) Lowden [GU SU FG] 

HYDROCHARITACEAE - SYNONYMS AND 


Elodea granatensis Humb. & Bonpl. =Apalanthe granatensis 

Elodea guyanensis L.C. Rich. =Elodea granatensis 

Hydromystria laevigata (Willd.) Hanzeker =Limnobium 

spongia subsp. laevigatum 

Limnobium laevigatum (Humb. & Bonpl. ex Willd.) Heine 

=Limnobium spongia subsp. laevigatum 

Limnobium stoloniferum (G. Mey.) Griseb. =Limnobium 

spongia subsp. laevigatum 

Philotria granatensis (Humb. & Bonpl.) Victorin =Apalanthe 

granatensis 

REFERENCES: 

HYDROPHYLLACEAE 

Constance, L. 1982. Hydrophyllaceae. In Z. 

Luces de Febres, Flora de Venezuela 8(3): 7- 

14. 

Davenport, L.J. 1988. A monograph of 

Hydrolea. Rhodora 90 (862): 169-208. 

Hydrolea elatior Schott [GU] 

Hydrolea palustris (Aubl.) Raeusch. [FG] [naturalized?] 

Hydrolea spinosa L. var. spinosa [GU SU FG] 

HYDROPHYLLACEAE - SYNONYMS AND 


Hydrolea minima Brand =Hydrolea elatior 

Hydrolea multiflora Mart. ex Choisy =Hydrolea elatior 

Sagonea palustris Aubl. =Hydrolea palustris 

ICACINACEAE 

reviewed by A.C. de Roon (1992); reviewed by 

A.C. de Roon (1996) 

REFERENCES: 

Howard, R.A. 1942. Studies in the Icacinaceae, 

III. A revision of Emmotum. J. Arn. Arb. 23: 

479-494. 

Howard, R.A. 1942. Studies of the Icacinaceae 

IV. Considerations of the New World genera. 

Contrib. Gray Herb. 142: 3-60. 

Howard, R.A. 1978. Icacinaceae. In B. Maguire 


Highland part 10. Mem. New York Bot. 

Gard. 29: 62-68. 

Howard, R.A. 1992. A revision of Casimirella, 

including Humirianthera. Brittonia 44(2): 

166-172. 

SPERMATOPHYTA: Hydrocharitaceae - Iridaceae 

Jansen-Jacobs, M.J. 1979. Icacinaceae. In A.L. 


5(1): 344-355. 

Roon, A.C. de. 1994. Icacinaceae. In A.R.A. 


A) 16: 1-157. 

Sleumer, H. 1942. Icacinaceae. In A. Engler & 


20b: 322-396. 

Casimirella ampla (Miers) R. Howard [FG] 

Dendrobangia boliviana Rusby [GU SU FG] 

Discophora guianensis Miers [GU SU FG] 

Emmotum acuminatum (Benth.) Miers [Guianan records 

erroneous, but possibly to be expected] 

Emmotum conjunctum R.A. Howard [GU] 

Emmotum fagifolium Desv. ex Hamilton [GU SU FG] 

Emmotum fulvum R.A. Howard [GU?] [Mt. Roraima] 

Emmotum glabrum Benth. ex Miers [GU] 

Emmotum orbiculatum (Benth.) Miers [Guianan records 

erroneous, but possibly to be expected] 

Leretia cordata Vell. [GU SU FG] 

Pleurisanthes artocarpi Baill. [GU FG] 

Pleurisanthes emarginata Tiegh. [SU FG] 

Pleurisanthes flava Sandw. [GU] 

Pleurisanthes parviflora (Ducke) Howard [SU FG] 

Poraqueiba guianensis Aubl. [GU SU FG] 

ICACINACEAE - SYNONYMS AND 


Clavapetalum surinamense Pulle =Dendrobangia boliviana 

Emmotum argenteum Gleason =Emmotum glabrum 

Emmotum ptarianum Steyerm. =Emmotum glabrum 

Humirianthera ampla (Miers) Baehni =Casimirella ampla 

Humirianthera duckei Huber =Casimirella ampla 

Leretia parviflora Ducke =Pleurisanthes parviflora 

Mappia ampla Miers =Casimirella ampla 

Mappia parviflora (Ducke) Baehni =Pleurisanthes parviflora 

Pleurisanthes brasiliensis Tiegh. [not in Guianas] 

Pleurisanthes simpliciflora Sleumer [not in Guianas] 

Pogopetalum acuminatum Benth. =Emmotum acuminatum 

Pogopetalum acutum Benth. =Emmotum fagifolium 

Pogopetalum orbiculatum Benth. =Emmotum orbiculatum 

Poraqueiba surinamensis Miers =Poraqueiba guianensis 

IRIDACEAE 

reviewed by P. Goldblatt (1992) 

REFERENCES: 

Boterenbrood, M.J.A. 1943. Iridaceae. In A. 


Goldblatt, P. & T.M. Howard. 1992. Notes on 

Alophia and a new species, A. veracruzana, 

from Vera Cruz, Mexico. Ann. Missouri Bot. 

Gard. 79(4): 901-905.


Goldblatt, P. & N. Snow. 1991. Systematics and 

chromosome cytology of Eleutherine Herbert. 


Alophia drummondii (Graham) R. Foster [GU] 

Cipura paludosa Aubl. [GU SU FG] 

Eleutherine bulbosa (P. Mill.) Urb. [GU SU FG] 

Neomarica caerulea (Ker-Gawl.) Sprague [escaped from 

cultivation] 

Sisyrinchium vaginatum Spreng. [GU SU] [to be expected in 

FG] 

Trimezia fosteriana Steyerm. [GU] 

Trimezia guianensis Ravenna [GU] 

Trimezia martinicensis (L.) Herbert [GU? FG] 

IRIDACEAE - SYNONYMS AND 


Ixia americana Aubl. =Eleutherine bulbosa 

Sisyrinchium alatum Hook. =Sisyrinchium vaginatum 

Sisyrinchium americanum (Aubl.) Lemée =Eleutherine 

bulbosa 

Sisyrinchium marchio (Vell.) Steud. =Sisyrinchium vaginatum 

IXONANTHACEAE 


REFERENCES: 

Forman, L.L. 1965. A new genus of 

Ixonanthaceae with notes on the family. Kew 

Bull. 19: 517-526. 

Kuhlmann, J.G. 1925. Cyrillaceae, in 

Contribuiçao para o conhecimento de algumas 

plantas novas. Arq. Jard. Bot. Rio de Janeiro 

4: 356-358. 

Steyermark, J.A. & J.L. Luteyn. 1980. Revision 

of the genus Ochthocosmus. Brittonia 32: 

128-143. 

Secco, R. de S. & S.M.B. Silva. 1990. Materiaes 

para a flora amazônica - VIII. Contribuçao à 

sistemática das Linaceae da Amazônia 

Brasileira. Bol. Mus. Paraense Emílio Goeldi 

(Bot.) 6(1): 113-135. 

Cyrillopsis paraensis Kuhlmann [FG] 

Ochthocosmus attenuatus Steyerm. & Luteyn [GU] 

Ochthocosmus barrae Hallier f. [GU?] 

Ochthocosmus longipedicellatus Steyerm. & Luteyn [GU] 

Ochthocosmus parvifolius Hallier f. [GU?] [incertae sedis] 

Ochthocosmus roraimae Benth. var. roraimae [GU] 

IXONANTHACEAE - SYNONYMS AND 


Ochthocosmus grandifolius Steyerm. =Ochthocosmus 

roraimae var. grandifolius 

Ochthocosmus roraimae Benth. var. parvifolius (Hallier f.) 

Steyerm. & Luteyn =Ochthocosmus parvifolius 

REFERENCES: 

JUGLANDACEAE 

SPERMATOPHYTA: Iridaceae - Lacistemataceae 

117 

Manning, W.E. 1960. The genus Juglans in 

South America and the West Indies. Brittonia 

12: 1-26. 

Juglans sp. [FG?] [none known from Guianas, probably based 

on misidentifications or cultivated material] 

JUGLANDACEAE - SYNONYMS AND 


Juglans brasiliensis Dode. =Juglans australis Griseb.[not in 


KRAMERIACEAE 

contributed by B.B. Simpson (1992); additions 

and corrections by B.B. Simpson (1996) 

REFERENCES: 

Simpson, B.B. 1989. Krameriaceae. Flora 


Krameria ixine Loefling [GU] 

Krameria spartioides Klotsch ex O. Berg [GU] 

Krameria tomentosa A. St. Hil. [GU?] 

REFERENCES: 

LACISTEMATACEAE 

Sleumer, H.O. 1980. Flacourtiaceae. Flora 


Vaandrager, G. 1938. Lacistemaceae. In A. 


Lacistema aggregatum (Bergius) Rusby [GU SU FG] 

Lacistema grandifolium Schnizlein [SU FG] 

Lacistema polystachyum Schnizlein [GU SU FG]

118 


LAMIACEAE 

reviewed by R.M. Harley (1992) 

REFERENCES: 

Epling, C. 1935-1937. Synopsis of the South 

American Labiatae. Repert. Sp. Nov. Beih. 

85: 1-341. 

Epling, C. 1949. Revisión del género Hyptis 

(Labiatae). Rev. Mus. La Plata (Sect. Bot.) 7: 

153-497. 

Harley, R.M. 1988. Revision of generic limits in 

Hyptis Jacq. (Labiatae) and its allies. Bot. J. 

Linn. Soc. 98: 87-95. 

Kostermans, A.J.G.H. 1936. Labiatae. In A. 


Kostermans, A.J.G.H. 1937. Labiatae (additions 


Suriname 4(1): 498. 

Eriope crassipes Benth. [FG] 

Hypenia salzmannii (Benth.) Harley [GU] 

Hyptidendron arboreum (Benth.) Harley [GU] 

Hyptis atrorubens Poit. [GU SU FG] 

Hyptis brevipes Poit. [SU FG] [frequently confused with H. 

lanceolata] 

Hyptis capitata Jacq. [FG?] 

Hyptis dilatata Benth. [GU] 

Hyptis hirsuta Kunth [GU SU] 

Hyptis lacustris St. Hil. ex Benth. [GU] 

Hyptis lanceolata Poir. [GU SU FG] 

Hyptis lantanifolia Poit. [GU SU FG] 

Hyptis microphylla Pohl ex Benth. [GU SU] 

Hyptis mutabilis (L.C. Rich.) Briquet [SU FG] 

Hyptis pachycephala Epling [FG] 

Hyptis parkeri Benth. [GU SU? FG?] 

Hyptis pectinata (L.) Poit. [GU SU? FG] 

Hyptis pulegioides Pohl ex Benth. [GU] 

Hyptis recurvata Poit. [GU SU FG] 

Hyptis spicigera Lam. [GU FG] 

Hyptis suaveolens (L.) Poit. [GU FG] 

Leonotis nepetifolia (L.) R. Brown [GU SU FG] 

Leonurus japonicus Houtt. [GU SU FG] [naturalized] 

Marsypianthes chamaedrys (Vahl) Kuntze [GU SU FG] 

Ocimum americanum L. [SU] 

Ocimum campechianum P. Mill. [GU SU FG] 

Ocimum gratissimum L. [SU] [naturalized] 

Scutellaria leucantha Loesen [FG] 

Scutellaria uliginosa St. Hil. [FG] 

LAMIACEAE - SYNONYMS AND 


Hyptis americana (Aubl.) Urb. =Hyptis spicigera 

Hyptis arborea Benth. =Hyptidendron arboreum 

Hyptis radiata Willd. =Hyptis alata (Raf.) Shinners [not in 


Hyptis salzmannii Benth. =Hypenia salzmannii 

Leonurus sibiricus Auct. non L. =Leonurus japonicus 

Ocimum micranthum Willd. =Ocimum campechianum 

SPERMATOPHYTA: Lamiaceae - Lauraceae 

LAURACEAE 

reviewed by H. van der Werff (1992); reviewed 

by H. van der Werff (1996) 

REFERENCES: 

Allen, C.K. 1964. Lauraceae. In B. Maguire, 

J.J. Wurdack, & collaborators, Botany of the 


Bot. Gard. 10(5): 1-278. 

Allen, C.K. 1965. Lauraceae. In B. Maguire & 



12(3): 1-285. 

Jansen-Jacobs, M.J. 1976. Lauraceae (additions 



Kostermans, A.J.G.H. 1936-1938. Revision of 

the Lauraceae. Rec. Trav. Bot. Neerl. 33: 

719-757 (I), 34: 500-609 (II), 35: 56-129 (III), 

35: 834-931 (V). 

Kostermans, A.J.G.H. 1936. Lauraceae. In A. 


Kostermans, A.J.G.H. 1939. Lauraceae 



Kubitzki, K. & S. Renner. 1982. Lauraceae I 

(Aniba and Aiouea). Flora Neotropica 

Monogr. 31: 1-124. 

Rohwer, J.G. 1986. Prodromus einer 

Monographie der Gattung Ocotea Aubl., 

sensu lato. Mitt. Inst. Allg. Bot. Hamburg 20: 

3-278. 

Rohwer, J.G. 1993. Lauraceae: Nectandra. 

Flora Neotrop. Monogr. 60: 1-332. 

Rohwer, J.G., H.G. Richter, & H. van der Werff. 

1991. Two new genera of neotropical 

Lauraceae and critical remarks on the generic 

delimitation. Ann. Missouri Bot. Gard. 78: 

388-400. 

Werff, H. van der. 1986. Kubitzkia van der 

Werff, a new name for a genus of neotropical 

Lauraceae. Taxon 35(1): 164-166. 

Werff, H. van der. 1987. A revision of 

Mezilaurus. Ann. Missouri Bot. Gard. 74: 

153-182. 

Werff, H. van der. 1991. A key to the genera of 

Lauraceae in the New World. Ann. Missouri 

Bot. Gard. 78: 377-387. 

Werff, H. van der. 1995. Two new species of 

Lauraceae from central French Guiana. 

Brittonia 47(4): 372-375. 

Aiouea guianensis Aubl. [GU SU FG] 

Aiouea impressa (Meissn.) Kosterm. [FG] 

Aiouea laevis (Mart.) Kosterm. [GU SU] 

Aiouea longipetiolata van der Werff [FG] 

Aiouea opaca van der Werff [FG] 

Aniba canelilla (Kunth) Mez [GU SU?]


Aniba citrifolia (Nees) Mez [GU FG] 

Aniba excelsa Kosterm. [GU] 

Aniba guianensis Aubl. [GU SU FG] 

Aniba hostmanniana (Nees) Mez [GU SU FG] 

Aniba hypoglauca Sandw. [GU SU] 

Aniba jenmanii Mez [GU SU FG] 

Aniba kappleri Mez [GU SU FG] 

Aniba megaphylla Mez [GU SU FG] 

Aniba panurensis (Meissn.) Mez [GU SU FG] 

Aniba parviflora (Meissn.) Mez [SU FG] 

Aniba percoriacea C.K. Allen [SU] 

Aniba permollis (Nees) Mez [GU] 

Aniba riparia (Nees) Mez [GU SU] 

Aniba rosaeodora Ducke [GU SU FG] 

Aniba taubertiana Mez [SU FG] 

Aniba terminalis Ducke [FG] 

Aniba williamsii O.C. Schmidt [GU? SU FG] 

Beilschmiedia curviramea (Meissn.) Kosterm. [GU] 

Beilschmiedia hexanthera van der Werff [FG] 

Cassytha filiformis L. [GU SU FG] 

Chlorocardium rodiei (Rob. Schomb.) Rohwer, Richter, & van 

der Werff [GU SU FG] 

Cinnamomum cinnamomifolia (Kunth) Kosterm. [recorded 

from FG in error, but perhaps to be expected in GU] 

Cryptocarya guianensis Meissn. [FG] 

Cryptocarya maroniensis Benoist [FG] 

Dicypellium caryophyllaceum (Mart.) Nees [FG?] 

Endlicheria anomala (Nees) Mez [GU SU] 

Endlicheria bracteolata (Meissn.) C.K. Allen [GU SU FG] 

Endlicheria endlicheriopsis (Mez) Kosterm. [GU SU FG] 

Endlicheria gracilis Kosterm. [GU SU FG] 

Endlicheria multiflora (Miq.) Mez [GU SU FG] 

Endlicheria paradoxa Mez [SU?] 

Endlicheria punctulata (Mez) C.K. Allen [GU SU FG] 

Endlicheria pyriformis (Nees) Mez [SU FG] 

Endlicheria reflectens (Nees) Mez [GU] 

Endlicheria sericea Nees [SU FG] 

Kubitzkia mezii (Kosterm.) van der Werff [GU SU FG] 

Licaria armeniaca (Nees) Kosterm. [GU] 

Licaria aurea (Huber) Kosterm. [GU SU] 

Licaria brasiliensis (Nees) Kosterm. [SU?] 

Licaria camara (Rob. Schomb.) Kosterm. [GU FG] 

Licaria cannella (Meissn.) Kosterm. [GU SU FG] 

Licaria chrysophylla (Meissn.) Kosterm. [GU SU FG] 

Licaria debilis (Mez) Kosterm. [GU SU FG] 

Licaria guianensis Aubl. [GU SU FG] 

Licaria martiniana (Mez) Kosterm. [GU SU FG] 

Licaria multiflora (Kosterm.) Kosterm. [GU] 

Licaria oppositifolia (Nees) Kosterm. [GU] 

Licaria pachycarpa (Meissn.) Kosterm. [GU FG] 

Licaria polyphylla (Nees) Kosterm. [GU SU FG] 

Licaria subbullata Kosterm. [GU SU FG] 

Licaria vernicosa (Mez) Kosterm. [GU SU FG] 

Mezilaurus itauba (Meissn.) Taub. ex Mez [SU FG] 

Mezilaurus lindaviana Schwacke & Mez [GU] 

Mezilaurus thoroflora van der Werff [GU] 

Nectandra amazonum Nees [GU SU] 

Nectandra cissiflora Nees [GU SU FG] 

Nectandra cuspidata Nees [GU SU] 

Nectandra globosa (Aubl.) Mez [GU SU FG] 

Nectandra hihua (Ruiz & Pav.) Rohwer [GU SU FG] 

Nectandra paucinervia Coe-Teixeira [GU] 

Nectandra pearcei Mez [GU SU] 

Nectandra pulverulenta Nees [FG] 

Nectandra reticulata (Ruiz & Pav.) Mez [SU FG] 

SPERMATOPHYTA: Lauraceae 

119 

Nectandra sanguinea Roland. ex Rottb. [GU SU] 

Ocotea aciphylla (Nees) Mez [GU SU FG] 

Ocotea amazonica (Meissn.) Mez [FG] 

Ocotea argyrophylla Ducke [SU FG] 

Ocotea bijuga (Rottb.) Bernardi [SU] [=Ocotea neesiana?] 

Ocotea canaliculata (L.C. Rich.) Mez [GU SU FG] 

Ocotea caracasana (Nees) Mez [GU] 

Ocotea ceanothifolia (Nees) Mez [FG] 

Ocotea cernua (Nees) Mez, s.l. [GU SU FG] 

Ocotea commutata (Nees) Mez [FG] 

Ocotea crassifolia (Nees) Mez [GU?] [Mt. Roraima] 

Ocotea cujumari Mart. [FG] 

Ocotea cymbarum Kunth [GU SU FG] 

Ocotea discrepens C.K. Allen [GU SU FG] 

Ocotea dissimilis C.K. Allen [FG] [=Ocotea splendens?] 

Ocotea divaricata (Nees) Mez [FG] 

Ocotea fasciculata (Nees) Mez [GU SU FG] 

Ocotea fendleri (Meissn.) Rohwer [FG] 

Ocotea floribunda (Swartz) Mez [GU SU FG] 

Ocotea glomerata (Nees) Mez [GU SU FG] 

Ocotea gracilis (Meissn.) Mez [GU] 

Ocotea guianensis Aubl. var. guianensis [GU SU FG] 

Ocotea guianensis Aubl. var. subsericea Kosterm. [GU SU] 

Ocotea hemsleyana Mez [GU] 

Ocotea indirectinervia C.K. Allen [FG] 

Ocotea leucoxylon (Swartz) de Lanessan [GU SU FG] 

Ocotea mucronata (Poir.) Kosterm. [FG] 

Ocotea neesiana (Miq.) Kosterm. [GU SU FG] [see also 

Ocotea bijuga] 

Ocotea nigra Benoist [GU SU FG] 

Ocotea oblonga (Meissn.) Mez [GU SU FG] 

Ocotea pauciflora (Nees) Mez [GU] 

Ocotea petalanthera (Meissn.) Mez [SU FG] 

Ocotea puberula (Rich.) Nees [GU SU FG] 

Ocotea rubra Mez [GU SU FG] 

Ocotea rubrinervis Mez [GU? FG] 

Ocotea rufovestita Ducke [FG] 

Ocotea sandwithii Kosterm. [GU] 

Ocotea scandens Kosterm. [FG] 

Ocotea schomburgkiana (Nees) Mez [GU SU FG] 

Ocotea splendens (Meissn.) Mez [GU SU FG] 

Ocotea tillettsiana C.K. Allen [GU] 

Ocotea tomentella Sandw. [GU FG] 

Persea americana P. Mill. [GU SU FG] [cultivated, 


Persea jenmanii Mez [GU] 

Persea julianae van der Werff [SU] 

Persea nivea Mez [GU SU FG] 

Pleurothyrium amapaense C.K. Allen [SU] 

Rhodostemonodaphne grandis (Mez) Rohwer [GU SU FG] 

Rhodostemonodaphne kunthiana (Nees) Rohwer [GU SU FG] 

Rhodostemonodaphne miranda (Sandw.) Rohwer [GU] 

Rhodostemonodaphne mirecolorata (C.K. Allen) Rohwer 

[GU] 

LAURACEAE - SYNONYMS AND 


Acrodiclidium aubletii Kosterm. =Licaria guianensis 

Acrodiclidium canella (Meissn.) Mez =Licaria cannella 

Acrodiclidium cayennense (Meissn.) Mez =Licaria cannella 

Acrodiclidium debile Mez =Licaria debilis 

Acrodiclidium guianense Nees =Licaria polyphylla 

Acrodiclidium martinianum Mez =Licaria martiniana

120 


Acrodiclidium rigidum Kosterm. =Licaria chrysophylla 

Aiouea demararensis Kosterm. =Aiouea guianensis 

Aiouea densiflora Nees =Aiouea laevis 

Aiouea schomburgkii Meissn. =Aiouea guianensis 

Aniba castanea C.K. Allen =Aniba jenmani 

Aniba duckei Kosterm. =Aniba rosaeodora 

Aniba firmula (Nees & Mart.) Mez [not in Guianas] 

Aniba gonggrijpii Kosterm. =Aniba panurensis 

Aniba koumaroucapa Kosterm. =Aniba megaphylla 

Aniba mas Kosterm. =Aniba panurensis 

Aniba muca (Ruiz & Pav.) Mez [not in Guianas] 

Aniba muelleriana Mez =Aniba citrifolia 

Aniba murcana C.K. Allen =Aniba williamsii 

Aniba ovalifolia Mez =Aniba hypoglauca 

Aniba salicifolia (Nees) Mez =Aniba guianensis 

Aniba sericea (Nees) Brooks =Endlicheria sericea 

Aniba trinitatis (Meissn.) Mez =Aniba citrifolia 

Cassytha americana Nees =Cassytha filiformis 

Clinostemon maguireanum (Allen) Kurz =Mezilaurus 

thoroflora 

Cryptocarya moschata Mart. [not in Guianas] 

Cryptocarya mucronata (Poir.) Spreng. =Ocotea mucronata 

Endlicheria impressa Mez =Aiouea impressa 

Endlicheria melinonii Benoist =Endlicheria multiflora 

Licaria appelii (Mez) Kosterm. =Licaria guianensis 

Licaria cayennensis (Meissn.) Kosterm. =Licaria cannella 

Licaria foveolata (Mez) Lemée =Licaria martiniana 

Licaria maguireana C.K. Allen =Mezilaurus thoroflora 

Licaria rigida (Kosterm.) Kosterm. =Licaria chrysophylla 

Licaria simulans C.K. Allen =Licaria vernicosa 

Licaria wilhelminensis C.K. Allen =Licaria subbullata 

Mezilaurus wurdackiana C.K. Allen =Mezilaurus lindaviana 

Misanteca maguireana (Allen) Lundell =Mezilaurus thoroflora 

Nectandra ambigua Meissn. =Nectandra amazonum 

Nectandra angustifolia (Schrad.) Nees [not in Guianas] 

Nectandra canescens Nees [not in Guianas] 

Nectandra grandis (Mez) Kosterm. =Rhodostemonodaphne 

grandis 

Nectandra guianensis Meissn. =Nectandra sanguinea 

Nectandra kaburiensis Kosterm. =Nectandra pearcei 

Nectandra kunthiana (Nees) Kosterm. =Rhodostemonodaphne 

kunthiana 

Nectandra laurel Nees [not in Guianas] 

Nectandra lucida Nees (1833) =Endlicheria paniculata 

(Spreng.) Macbr.[not in Guianas] 

Nectandra lucida Nees (1836) =Nectandra hihua 

Nectandra magnoliifolia Meissn. =Nectandra canescens Nees 


Nectandra maguireana C.K. Allen [not in Guianas] 

Nectandra miranda Sandw. =Rhodostemonodaphne miranda 

Nectandra mollis (Kunth) Nees =Nectandra reticulata 

Nectandra pallida Nees =Nectandra amazonum 

Nectandra pichurim (Kunth) Mez [not in Guianas] 

Nectandra pisi Miq. =Nectandra globosa 

Nectandra praeclara Sandw. =Rhodostemonodaphne grandis 

Nectandra rubra (Mez) C.K. Allen =Ocotea rubra 

Nectandra schomburgkii Meissn. =Nectandra hihua 

Nectandra surinamensis Mez =Nectandra cissiflora 

Nectandra viburnoides Meissn. [not in Guianas] 

Ocotea abbreviata Schwacke & Mez [not in Guianas] 

Ocotea acutangula (Miq.) Mez =Ocotea leucoxylon 

Ocotea barcellensis (Meissn.) Mez =Ocotea cymbarum 

Ocotea caudata (Nees) Mez =Ocotea cernua s.l. 

Ocotea costulata (Nees) Mez =Ocotea aciphylla 

Ocotea duotincta C.K. Allen =Ocotea leucoxylon 

SPERMATOPHYTA: Lauraceae - Lecythidaceae 

Ocotea endlicheriopsis Mez =Endlicheria endlicheriopsis 

Ocotea fallax (Miq.) Mez =Ocotea neesiana 

Ocotea finium C.K. Allen =Ocotea aciphylla 

Ocotea firmula Mart. [nomen nudum] =Aniba firmula (Nees) 

Mez [not in Guianas] 

Ocotea glaucina (Meissn.) Mez =Ocotea notata (Nees) Mez 


Ocotea globifera Mez =Ocotea splendens 

Ocotea indecora (Schott) Mez [not in Guianas] 

Ocotea jenmanii Mez [nomen nudum] 

Ocotea lasseriana C.K. Allen =Ocotea leucoxylon 

Ocotea laxiflora (Meissn.) Mez =Ocotea pauciflora 

Ocotea marowynensis (Miq.) Mez =Ocotea cernua 

Ocotea martiniana (Nees) Mez =Ocotea puberula 

Ocotea mirecolorata C.K. Allen =Rhodostemonodaphne 

mirecolorata 

Ocotea opifera Mart. =Ocotea longifolia 

Ocotea pendula (Poepp.) Engl. [nomen nudum] 

Ocotea persulcata C.K. Allen =Ocotea leucoxylon 

Ocotea pubescens (Nees) Mez [not in Guianas] 

Ocotea punctulata Mez =Endlicheria punctulata 

Ocotea rhynchophylla (Meissn.) Mez =Ocotea aciphylla 

Ocotea rodiei (Rob. Schomb.) Mez =Chlorocardium rodiei 

Ocotea roraimae Mez =Ocotea aciphylla 

Ocotea subalveolata C.K. Allen =Ocotea schomburgkiana 

Ocotea subglabra Benoist =Ocotea puberula 

Ocotea urophylla (Meissn.) Mez =Ocotea cernua 

Ocotea vernicosa Mez =Licaria vernicosa 

Ocotea wachenheimii Benoist =Ocotea floribunda 

Persea coriacea Kosterm. [type based on mixed collection] 

Phoebe cinnamomifolia (Kunth) Nees =Cinnamomum 

cinnamomifolia 

Pleurothyrium cowaniana C.K. Allen =Rhodostemonodaphne 

kunthiana 

Systemonodaphne geminiflora Mez =Kubitzkia mezii 

Systemonodaphne mezii Kosterm. =Kubitzkia mezii 

LECYTHIDACEAE 

contributed by S.A. Mori & G.T. Prance (1992); 

additions and corrections by S.A. Mori (1996) 

REFERENCES: 

[for a more extensive bibliography, see Mori & 

Prance 1993] 

Eyma, P.J. 1934. Lecythidaceae. In A. Pulle, 


Mori, S.A. & collaborators. 1987. The 

Lecythidaceae of a lowland neotropical forest: 

La Fumée Mountain, French Guiana. Mem. 


Mori, S.A. & G.T. Prance. 1990. Lecythidaceae 

- part II. Flora Neotropica Monogr. 21(II): 1- 

373. 

Mori, S.A. & G.T. Prance. 1993. Lecythidaceae. 


Guianas (series A) 12: 1-144.


Prance, G.T. & S.A. Mori. 1979. Lecythidaceae 

- part I. Flora Neotropica Monogr. 21(I): 1- 

270. 

Bertholletia excelsa Humb. & Bonpl. [GU SU FG] 

Corythophora amapaensis Pires ex Mori & Prance [FG] 

Corythophora labriculata (Eyma) Mori & Prance [SU] 

Corythophora rimosa W. Rodrigues subsp. rubra Mori [FG] 

Couratari calycina Sandw. [GU FG] 

Couratari gloriosa Sandw. [GU SU FG] 

Couratari guianensis Aubl. emend. Prance [GU SU FG] 

Couratari multiflora (J.E. Smith) Eyma [GU SU FG] 

Couratari oblongifolia Ducke & Knuth [GU SU FG] 

Couratari oligantha A.C. Smith [GU] 

Couratari riparia Sandw. [GU] 

Couratari sandwithii Prance [SU] 

Couratari stellata A.C. Smith [GU SU FG] 

Couroupita guianensis Aubl. [GU SU FG] 

Eschweilera alata A.C. Smith [GU FG] 

Eschweilera apiculata (Miers) A.C. Smith [SU FG] 

Eschweilera boltenii Mori [SU] 

Eschweilera chartaceifolia Mori [FG] 

Eschweilera collina Eyma [GU SU FG] 

Eschweilera congestiflora (Benoist) Eyma [GU? SU FG] 

Eschweilera coriacea (A.P. DC.) Mori [GU SU FG] 

Eschweilera decolorans Sandw. [GU SU FG] 

Eschweilera eperuetorum Sandw. [GU SU] 

Eschweilera fanshawei Sandw. [GU] 

Eschweilera grandiflora (Aubl.) Sandw. [FG] 

Eschweilera laevicarpa Mori [FG] 

Eschweilera micrantha (Berg) Miers [GU SU FG] 

Eschweilera parviflora (Aubl.) Miers [GU FG] 

Eschweilera parvifolia Mart. ex A.P. DC. [GU] 

Eschweilera pedicellata (Rich.) Mori [GU SU FG] 

Eschweilera piresii Mori subsp. viridipetala Mori [FG] 

Eschweilera potaroensis Sandw. [GU] 

Eschweilera praeclara Sandw. [GU] 

Eschweilera sagotiana Miers [GU SU FG] 

Eschweilera simiorum (Benoist) Eyma [GU SU FG] 

Eschweilera squamata Mori [SU FG] 

Eschweilera subglandulosa (Steud. ex Berg) Miers [GU SU 

FG] 

Eschweilera wachenheimii (Benoist) Sandw. [GU SU FG] 

Gustavia augusta L. [GU SU FG] 

Gustavia gigantophylla Sandw. [GU] 

Gustavia hexapetala (Aubl.) J.E. Smith [GU SU FG] 

Gustavia poeppigiana Berg [GU] 

Lecythis alutacea (A.C. Smith) Mori [GU SU FG] 

Lecythis brancoensis (R. Knuth) Mori [GU] 

Lecythis chartacea Berg [GU SU FG] 

Lecythis confertiflora (A.C. Smith) Mori [GU SU FG] 

Lecythis corrugata Poit. subsp. corrugata [GU SU FG] 

Lecythis corrugata Poit. subsp. rosea (Spruce ex Berg) Mori 

[GU] 

Lecythis holcogyne (Sandw.) Mori [GU SU FG] 

Lecythis idatimon Aubl. [SU FG] 

Lecythis persistens Sagot subsp. aurantiaca Mori [FG] 

Lecythis persistens Sagot subsp. persistens [GU SU? FG] 

Lecythis pneumatophora Mori [FG] 

Lecythis poiteaui Berg [SU FG] 

Lecythis schomburgkii Berg [GU] 

Lecythis zabucajo Aubl. [GU SU FG] 

LECYTHIDACEAE - SYNONYMS AND 


SPERMATOPHYTA: Lecythidaceae 

121 

Chytroma amara (Aubl.) Miers =Lecythis idatimon 

Chytroma apiculata Miers =Eschweilera apiculata 

Chytroma brancoensis Knuth =Lecythis brancoensis 

Chytroma chartacea (Berg) Miers =Lecythis chartacea 

Chytroma collina (Eyma) Knuth =Eschweilera collina 

Chytroma congestiflora (Benoist) Knuth =Eschweilera 

congestiflora 

Chytroma corrugata (Poit.) Nied. =Lecythis corrugata 

Chytroma foetida Knuth =Lecythis poiteaui 

Chytroma holcogyne (Sandw.) Knuth =Lecythis holcogyne 

Chytroma idatimon (Aubl.) Nied. =Lecythis idatimon 

Chytroma labriculata (Eyma) Knuth =Corythophora 

labriculata 

Chytroma laevicula (Miers) Knuth =Lecythis chartacea 

Chytroma marawynensis (Berg) Miers =Lecythis chartacea 

Chytroma perspicua Miers =Eschweilera pedicellata 

Chytroma praeclara (Sandw.) Kunth =Eschweilera praeclara 

Chytroma rubiflora Miers =Lecythis corrugata subsp. 

corrugata 

Chytroma salebrosa (Berg) Miers =Lecythis corrugata subsp. 

corrugata 

Chytroma schomburgkiana (Berg) Miers =Lecythis 

schomburgkii 

Chytroma simiorum (Benoist) Knuth =Eschweilera simiorum 

Chytroma simiorum (Benoist) Knuth var. latifolia (Eyma) 

Knuth =Eschweilera simiorum 

Couratari coriacea Mart. ex O. Berg =Couratari multiflora 

Couratari fagifolia (Miq.) Eyma =Couratari multiflora 

Couratari pulchra Sandw. =Couratari guianensis 

Couratari reticulata A.C. Smith =Couratari stellata 

Couroupita guianensis Aubl. var. surinamensis (Mart. ex Berg) 

Eyma =Couroupita guianensis 

Couroupita surinamensis Mart. ex Berg =Couroupita 

guianensis 

Eschweilera alutacea A.C. Smith =Lecythis alutacea 

Eschweilera amara (Aubl.) Nied. =Lecythis idatimon 

Eschweilera chartacea (O. Berg) Eyma =Lecythis chartacea 

Eschweilera conduplicata A.C. Smith =Lecythis corrugata 

subsp. rosea 

Eschweilera confertiflora A.C. Smith =Lecythis confertiflora 

Eschweilera confusa (Benoist) Knuth [nomen nudum] 

Eschweilera corrugata (Poit.) Miers =Lecythis corrugata 

Eschweilera flaccida Miers =Eschweilera pedicellata 

Eschweilera floribunda Eyma =Eschweilera micrantha 

Eschweilera gracilipes (Sagot) Knuth =Eschweilera micrantha 

Eschweilera grata Sandw. =Eschweilera parviflora 

Eschweilera holcogyne Sandw. =Lecythis holcogyne 

Eschweilera idatimon (Aubl.) Nied. =Lecythis idatimon 

Eschweilera idatimonoides (Berg) Miers =Eschweilera 

pedicellata 

Eschweilera jenmanii Knuth =Lecythis corrugata subsp. 

corrugata 

Eschweilera jucunda (Benoist) Knuth =Eschweilera sagotiana 

Eschweilera labriculata Eyma =Corythophora labriculata 

Eschweilera longipes (Poit.) Miers =Eschweilera pedicellata 

Eschweilera lutea (Aubl.) Miers =Lecythis idatimon 

Eschweilera macrophylla (Berg) Miers =Eschweilera 

pedicellata 

Eschweilera marawynensis (Berg) Nied. =Lecythis chartacea

122 


Eschweilera melinonis (Sagot) Knuth =Eschweilera sagotiana 

Eschweilera montana A.C. Smith =Eschweilera parviflora 

Eschweilera odora (Poepp.) Miers =Eschweilera coriacea 

Eschweilera ovalis Knuth =Lecythis persistens subsp. 

persistens 

Eschweilera pachycarpa Mart. [nomen nudum] 

Eschweilera pallida Miers =Eschweilera coriacea 

Eschweilera patrisii Knuth =Lecythis corrugata subsp. 

corrugata 

Eschweilera persaudii Knuth =Lecythis alutacea 

Eschweilera platycarpa (Poit.) Miers =Eschweilera pedicellata 

Eschweilera poiteaui (Berg) Nied. =Lecythis poiteaui 

Eschweilera retroflexa (Benoist) Knuth =Eschweilera coriacea 

Eschweilera salebrosa (Berg) Nied. =Lecythis corrugata subsp. 

corrugata 

Eschweilera schomburgkii (O. Berg) Nied. =Lecythis 

schomburgkii 

Eschweilera simiorum (Benoist) Eyma var. latifolia Eyma 

=Eschweilera simiorum 

Eschweilera tapuya (Benoist) Knuth =Eschweilera sagotiana 

Grias aubletiana Miers =Gustavia augusta 

Grias tetrapetala (Aubl.) Nied. =Gustavia augusta 

Gustavia augusta L. var. guianensis Berg =Gustavia augusta 

Gustavia augusta L. var. verrucosa Mart. ex Berg =Gustavia 

augusta 

Gustavia fastuosa Willd. =Gustavia hexapetala 

Gustavia fastuosa Willd. var. angustisepala Berg =Gustavia 

hexapetala 

Gustavia fastuosa Willd. var. latisepala Berg =Gustavia 

hexapetala 

Gustavia tetrapetala (Aubl.) A.P. DC. =Gustavia augusta 

Gustavia urceolata Poit. =Gustavia augusta 

Japarandiba augusta (L.) Kuntze =Gustavia augusta 

Japarandiba fastuosa (Willd.) Kuntze =Gustavia hexapetala 

Japarandiba hexapetala (Aubl.) Kuntze =Gustavia hexapetala 

Jugastrum poiteaui (Berg) Miers =Lecythis poiteaui 

Lecythis alba Mori [nomen nudum] =Lecythis persistens 

subsp. persistens 

Lecythis amara Aubl. =Lecythis idatimon 

Lecythis chartacea Berg var. calyce Sagot =Lecythis 

chartacea 

Lecythis cognata Miers =Lecythis corrugata subsp. corrugata 

Lecythis congestiflora Benoist =Eschweilera congestiflora 

Lecythis crassinoda Miers =Lecythis zabucajo 

Lecythis davisii Sandw. =Lecythis zabucajo 

Lecythis davisii Sandw. var. gracilipes Eyma =Lecythis 

zabucajo 

Lecythis gracilipes Sagot =Eschweilera micrantha 

Lecythis grandiflora Aubl. =Eschweilera grandiflora 

Lecythis jucunda Benoist =Eschweilera sagotiana 

Lecythis laevicula Miers =Lecythis chartacea 

Lecythis lutea Aubl. =Lecythis idatimon 

Lecythis macrophylla Berg =Eschweilera pedicellata 

Lecythis marawynensis Berg =Lecythis chartacea 

Lecythis martinii Berg =Lecythis corrugata subsp. corrugata 

Lecythis melinonis Sagot =Eschweilera sagotiana 

Lecythis micrantha Berg =Eschweilera micrantha 

Lecythis multiflora J.E. Smith =Couratari multiflora 

Lecythis parviflora Aubl. =Eschweilera parviflora 

Lecythis parviflora Sagot ex Miers =Eschweilera sagotiana 

Lecythis platycarpa Poiteau =Eschweilera pedicellata 

Lecythis racemiflora Sagot =Lecythis poiteaui 

Lecythis retroflexa (Benoist) Knuth =Eschweilera coriacea 

Lecythis rosea Spruce ex Berg =Lecythis corrugata subsp. 

rosea 

SPERMATOPHYTA: Lecythidaceae - Lemnaceae 

Lecythis rubicunda Miers =Lecythis corrugata subsp. 

corrugata 

Lecythis sagotiana (Miers) Sagot =Eschweilera sagotiana 

Lecythis salebrosa Berg =Lecythis corrugata subsp. corrugata 

Lecythis simiorum Benoist =Eschweilera simiorum 

Lecythis subglandulosa Steud. ex Berg =Eschweilera 

subglandulosa 

Lecythis tapuya Benoist =Eschweilera sagotiana 

Lecythis tumefacta Miers =Lecythis zabucajo 

Lecythis venusta Miers =Lecythis corrugata subsp. corrugata 

Lecythis wachenheimii Benoist =Eschweilera wachenheimii 

Lecythis wullschlaegeliana Berg =Eschweilera pedicellata 

Pirigara hexapetala Aubl. =Gustavia hexapetala 

Pirigara tetrapetala Aubl. =Gustavia augusta 

LEMNACEAE 


REFERENCES: 

Daubs, E.H. 1965. A monograph of Lemnaceae. 

Ill. Biol. Monogr. 34: 1-118. 

Hartog, C. den. & F. van der Plas. 1970. A 

synopsis of the Lemnaceae. Blumea 18: 355- 

368. 

Landolt, E. 1986-1987. The family of 

Lemnaceae - a monographic study. Veröff. 

Geobot. Inst. E.T.H. Stift. Rübel, Zürich 71: 

1-566, 95: 1-638. 

Lemna aequinoctialis Welwitsch [GU SU FG] 

Lemna valdiviana Phil. [SU] 

Spirodela intermedia W. Koch [SU] 

Spirodela punctata (G. Mey.) Thompson [GU] 

Wolffia brasiliensis Weddell [SU] 

Wolffiella lingulata (Hegelm.) Hegelm. [SU FG] 

Wolffiella neotropica Landolt [SU] 

Wolffiella welwitschii (Hegelm.) Monod [GU SU FG] 

LEMNACEAE - SYNONYMS AND 


Lemna gibba L. [not in Guianas] 

Lemna minor L. [not in Guianas] 

Lemna paucicostata Hegelm. =Lemna aequinoctialis 

Lemna perpusilla Torrey [not in Guianas] 

Lemna trinervis sensu Small, non (Austin) Small =Lemna 

aequinoctialis 

Spirodela biperforata W. Koch =Spirodela intermedia 

Spirodela polyrrhiza (L.) Schleid. [not in Guianas] 

Wolffiopsis welwitschii (Hegelm.) den Hartog & van der Plas 

=Wolffiella welwitschii


LENTIBULARIACEAE 

contributed by P. Taylor (1992); additions and 

corrections by P. Taylor (1996) 

REFERENCES: 

Taylor, P. 1967. Lentibulariaceae. In B. 



Bot. Gard. 17(1): 1-439. 

Taylor, P. 1989. The genus Utricularia - a 

taxonomic monograph. Kew Bull. Add. Ser. 

XIV: 1-724. 

Genlisea filiformis St. Hil. [GU SU FG] 

Genlisea guianensis N.E. Brown [GU] 

Genlisea pygmaea St. Hil. [GU SU FG] 

Genlisea repens Benj. [GU SU] 

Genlisea roraimensis N.E. Brown [GU] 

Utricularia adpressa Salzm. ex St. Hil. & Girard [GU SU FG] 

Utricularia alpina Jacq. [GU SU] 

Utricularia amethystina Salzm. ex St. Hil. & Girard [GU SU 

FG] 

Utricularia benjaminiana Oliver [GU SU FG] 

Utricularia breviscapa Wright ex Griseb. [GU] 

Utricularia calycifida Benj. [GU SU] 

Utricularia campbelliana Oliver [GU] 

Utricularia choristotheca P. Taylor [SU FG] 

Utricularia cucculata A. St. Hil. & Girard [GU SU FG] 

Utricularia determannii P. Taylor [SU] 

Utricularia erectiflora St. Hil. & Girard [GU SU] 

Utricularia foliosa L. [GU SU FG] 

Utricularia gibba L. [GU SU FG] 

Utricularia guyanensis A. DC. [GU SU FG] 

Utricularia hispida Lam. [GU SU FG] 

Utricularia humboldtii Rob. Schomb. [GU] 

Utricularia hydrocarpa Vahl [GU SU FG] 

Utricularia jamesoniana Oliver [GU SU FG] 

Utricularia juncea Vahl [GU SU FG] 

Utricularia lloydii Merl [SU FG] 

Utricularia longeciliata A. DC. [GU SU FG] 

Utricularia myriocista St. Hil. & Girard [GU SU FG] 

Utricularia nana St. Hil. & Girard [GU SU FG] 

Utricularia olivacea Wright ex Griseb. [GU SU] 

Utricularia pubescens J.E. Smith [GU FG] 

Utricularia pusilla Vahl [GU SU FG] 

Utricularia quelchii N.E. Brown [GU] 

Utricularia sandwithii P. Taylor [GU SU] 

Utricularia simulans Pilger [GU SU FG] 

Utricularia subulata L. [GU SU FG] 

Utricularia tenuissima Tutin [GU SU FG] 

Utricularia trichophylla Spruce ex Oliver [GU SU FG] 

Utricularia triloba Benj. [GU SU FG] 

Utricularia viscosa Spruce ex Oliver [GU SU FG] 

LENTIBULARIACEAE - SYNONYMS AND 


SPERMATOPHYTA: Lentibulariaceae - Liliaceae 

123 

Akentra inflata Benj. =Utricularia benjaminiana 

Biovularia olivacea (Wright) Kam. =Utricularia olivacea 

Genlisea anfractuosa Tutin =Genlisea filiformis 

Genlisea esmeraldae Steyerm. =Genlisea pygmaea 

Genlisea nigrocaulis Steyerm. =Genlisea pygmaea + Genlisea 

repens [based on mixed collection] 

Genlisea pulchella Tutin =Genlisea repens 

Polypompholyx laciniata Benj. =Utricularia simulans 

Utricularia alutacea Tutin =Utricularia amethystina 

Utricularia angulosa Poir. =Utricularia juncea 

Utricularia angustifolia Benj. =Utricularia hispida 

Utricularia arenicola Tutin =Utricularia nana 

Utricularia concinna N.E. Brown =Utricularia jamesoniana 

Utricularia connellii N.E. Brown =Utricularia pubescens 

Utricularia fimbriata Kunth [not in Guianas] 

Utricularia flaccida A. DC. [not in Guianas] 

Utricularia fockeana Miq. =Utricularia hydrocarpa 

Utricularia guianensis Splitg. ex De Vries =Utricularia foliosa 

Utricularia hirtella St. Hil. & Girard =Utricularia amethystina 

Utricularia inflata Walter [sensu stricto, not in Guianas] 

Utricularia kaieteurensis Steyerm. =Utricularia amethystina 

Utricularia leptantha Benj. =Utricularia pusilla 

Utricularia magnifica L. =Utricularia myriocista 

Utricularia maguirei Steyerm. =Utricularia calycifida 

Utricularia muscosa Benj. =Utricularia calycifida 

Utricularia obovata Miq. =Utricularia amethystina 

Utricularia obtusa Swartz =Utricularia gibba 

Utricularia oligosperma St. Hil. =Utricularia foliosa 

Utricularia parkeriana A. DC. =Utricularia gibba 

Utricularia peckii S.F. Blake =Utricularia guyanensis 

Utricularia pectinata Split. ex De Vries =Utricularia 

longeciliata 

Utricularia peltata Spruce ex Oliver =Utricularia pubescens 

Utricularia praelonga St. Hil. & Girard [not in Guianas] 

Utricularia purpurea Walt. [not in Guianas] 

Utricularia roraimensis N.E. Brown =Utricularia amethystina 

Utricularia rubricaulis Tutin =Utricularia guyanensis 

Utricularia sciaphila Tutin =Utricularia pubescens 

Utricularia spatulata Splitg. ex De Vries =Utricularia 

amethystina 

Utricularia spatulifolia Pilger =Utricularia amethystina 

Utricularia stricta G. Mey. =Utricularia juncea 

Utricularia tenuifolia Benj. =Utricularia gibba 

Utricularia tenuiscapa Pilger =Utricularia subulata 

LILIACEAE 

revised by J.K. Boggan (1996) 

REFERENCES: 

Baker, J.G. 1888. Handbook of the 

Amaryllideae. London: George Bell & Sons. 

Boterenbrood, M.J.A. 1943. Amaryllidaceae. In 


Cruden, R.W. 1991. A revision of Isidrogalvia: 

recognition for Ruíz & Pavón's genus. Syst. 

Bot. 16: 270-282.

124 


Killip, E.P. 1936. Bomarea, a genus of showy 

Andean plants. Nat. Hort. Mag. 15(2): 115- 

128. 

Maas, P.J.M. 1968. Amaryllidaceae (additions 


Flora of Suriname 1(2): 376. 

Martyn, E.B. 1936. Amaryllidaceae of British 

Guiana. Herbertia 3: 71. 

Sealy, J.R. 1954. Review of the genus 

Hymenocallis. Kew Bull. 1954: 201-240. 

Traub, H.P. & H.N. Moldenke. 1949. 

Amaryllidaceae: Tribe Amarylleae. American 

Plant Life Society. 

Uphof, J.C. Th. 1942. A review of the species of 

Crinum. Herbertia 9: 63-84. 

Alstroemeria sp. [FG] 

Bomarea edulis (Tussac) Herbert [GU SU] 

Crinum erubescens L.f. ex Solander [GU SU FG] 

Curculigo scorzonerifolia (Lam.) Baker [GU SU FG] 

Hippeastrum barbatum Herbert [SU] 

Hippeastrum elegans (Spreng.) H.E. Moore [GU] 

Hippeastrum puniceum (Lam.) Kuntze [GU SU FG] 

Hippeastrum stylosum Herbert [distribution described as 

"Guiana"] 

Hippeastrum vanleestenii (Traub) H.E. Moore [SU] 

[=Hippeastrum puniceum?] 

Hymenocallis caribaea (L. emend. Ker-Gawl.) Herbert [GU 

SU FG] [cultivated or escaped from cultivation] 

Hymenocallis littoralis (Jacq.) Salisb. [SU FG] [cultivated or 


Hymenocallis tubiflora Salisb. [GU SU FG] 

Hypoxis decumbens L. [GU? SU? FG?] [most, if not all, 

records represent Curculigo scorzonerifolia] 

Isidrogalvia schomburgkiana (Oliver) Cruden [GU] 

Nietneria corymbosa Klotzsch & Rich. Schomb. [GU?] [Mt. 

Roraima] 

Nietneria paniculata Steyerm. [GU] 

Zephyranthes citrina Baker [GU?] [type attributed to Guyana, 

possibly erroneously] 

Zephyranthes tubispatha (Ker-Gawl.) Herbert [SU] 

LILIACEAE - SYNONYMS AND 


Amaryllis belladonna L. [name incorrectly applied to 

Hippeastrum puniceum] 

Amaryllis belladonna L. var. barbata (Herbert) Traub & 

Moldenke =Hippeastrum barbatum 

Amaryllis stylosa (Herbert) Sweet =Hippeastrum stylosum 

Amaryllis vanleestenii Traub =Hippeastrum vanleestenii 

Bomarea fuscata Klotzsch ex Schomb. [nomen nudum] 

Bomarea guianensis Kränzl. =Bomarea edulis 

Crinum commelyni Jacq. =Crinum erubescens 

Crinum guianense M. Roem. =Crinum erubescens? 

Crinum lancei Herb. ex Sweet =Crinum erubescens? 

Crinum lindleyanum Schult.f. ex Seub. =Crinum erubescens 

Echeandia sp. [recorded from Guianas in error, but possibly to 

be expected] 

Hippeastrum equestre (Ait.) Herbert =Hippeastrum puniceum 

Hippeastrum solandriflorum Herbert =Hippeastrum elegans 

Hymenocallis guianensis (Ker-Gawl.) Herbert =Hymenocallis 

tubiflora 

SPERMATOPHYTA: Liliaceae - Loganiaceae 

Hymenocallis obtusata (Griseb.) Walp. =Hymenocallis 

caribaea 

Isidrogalvia guianensis Klotzsch [nomen nudum] 

=Isidrogalvia schomburgkiana 

Pancratium guianense Ker-Gawl. =Hymenocallis tubiflora 

Tofieldia guianensis (Klotzsch) Schulze [nomen nudum] 

=Isidrogalvia schomburgkiana 

Tofieldia rupestris Gleason =Navia rupestris (Gleason) Sandw. 

(Bromeliaceae) 

Tofieldia schomburgkiana Oliver =Isidrogalvia 


LIMNOCHARITACEAE 


(1992) 

REFERENCES: 


Limnocharitaceae. Flora Neotropica Monogr. 

56: 1-34. 

Jonker, F.P. 1943. Butomaceae. In A. Pulle, 


Hydrocleys nymphoides (Humb. & Bonpl. ex Willd.) Buch. 

[GU SU FG?] 

Limnocharis flava (L.) Buchenau [FG?] [record not verified, 

but to be expected in the Guianas] 

LISSOCARPACEAE 

Lissocarpa benthamii Baill. [GU] 

Lissocarpa guianensis Gleason [GU] 

LOASACEAE 

Mentzelia aspera L. [FG?] 

LOGANIACEAE 

reviewed by A.J.M. Leeuwenberg (1996) 

REFERENCES: 

Hurley, H.H. 1967. A taxonomic revision of the 

genus Spigelia. George Washington 

University, Washington, DC. [Unpublished 

Ph.D. thesis] 

Krukoff, B.A. & J. Monachino. 1942. The 

American species of Strychnos. Brittonia 4: 

248-322. [See also the subsequent series of 

supplementary notes in various journals.]


Leeuwenberg, A.J.M. 1961. The Loganiaceae of 

Africa II. A revision of Mostuea Didr. 

Meded. Landb. Wag. 61(4): 1-31. 

Leeuwenberg, A.J.M. 1969. Notes on American 

Loganiaceae. III. Revision of Bonyunia 

Rich. Schomb. Acta Bot. Neerl. 18(1): 152- 

158. 

Leeuwenberg, A.J.M. 1971. Notes on American 

Loganiaceae. V. Key to the genera. Acta 

Bot. Neerl. 20(5): 539-542. 

Leeuwenberg, A.J.M. 1975. The Loganiaceae of 

Africa XII. A revision of Mitreola L. Meded. 

Landb. Wag. 74(23): 1-28. 

Leeuwenberg, A.J.M. [ed.]. 1980. Loganiaceae. 

In Engler & Prantl, Die Natürlichen 

Pflanzenfamilian 28bI: 1-255. 

Raalte, M.H. van. 1932. Loganiaceae. In A. 


Smith, L.B. 1960. Notes on South American 

phanerogams - III. Wrightia 2(2): 90-102. 

Uittien, H. 1937. Loganiaceae (additions and 


4(1): 472-474. 

Antonia ovata Pohl [GU SU FG] 

Bonyunia minor N.E. Brown [GU] 

Bonyunia superba Rich. Schomb. [GU] [Mt. Roraima] 

Mitreola petiolata (J.F. Gmelin) Torrey & A. Gray [SU FG?] 

Mostuea surinamensis Benth. [SU] [to be expected in FG] 

Polypremnum procumbens L. [SU?] 

Potalia amara Aubl. [SU FG] 

Spigelia anthelmia L. [GU SU FG] 

Spigelia flemmingiana Cham. & Schlechtend. [GU] 

Spigelia humilis Benth. [GU] 

Spigelia multispica Steud. [GU SU FG] 

Spigelia polystachya Klotzsch ex Prog. [GU] 

Strychnos bredemeyeri (Schult.) Sprague & Sandw. [GU] 

Strychnos castelnaeana Wedd. [FG?] 

Strychnos cayennensis Krukoff & Barneby [FG] 

Strychnos cogens Benth. [GU SU FG] 

Strychnos darienensis Seem. [GU] 

Strychnos diaboli Sandw. [GU] 

Strychnos divaricans Ducke [SU FG] 

Strychnos erichsonii Rich. Schomb. ex Progel [GU SU FG] 

Strychnos eugeniifolia Monachino [SU FG] 

Strychnos glabra Sagot ex Progel [GU SU FG] 

Strychnos guianensis (Aubl.) Mart. [GU SU FG] 

Strychnos hirsuta Spruce ex Benth. [GU FG] 

Strychnos jobertiana Baill. [FG] 

Strychnos medeola Sagot ex Progel [SU FG] 

Strychnos melinoniana Baill. [GU SU FG] 

Strychnos mitscherlichii Rich. Schomb. var. mitscherlichii 

[GU SU FG] 

Strychnos oiapocensis Froes [SU FG] 

Strychnos panurensis Sprague & Sandw. [FG] 

Strychnos peckii B.L. Robinson [GU SU FG] 

Strychnos ramentifera Ducke [SU FG?] 

Strychnos rondeletioides Spruce ex Benth. [FG] 

Strychnos subcordata Spruce ex Benth. [to be expected in FG] 

Strychnos tomentosa Benth. [GU SU FG] 

Strychnos toxifera Rob. Schomb. ex Benth. [GU SU FG] 

LOGANIACEAE - SYNONYMS AND 


SPERMATOPHYTA: Loganiaceae - Loranthaceae 

125 

Mostuea brasiliensis Huber =Mostuea surinamensis 

Rouhamon guianensis Aubl. =Strychnos guianensis 

Spigelia guianensis (Aubl.) Lemée =Montira guianensis 

Aubl.[incertae sedis; excluded from Spigelia by Hurley] 

Spigelia humboldtiana Cham. & Schlechtend. =Spigelia scabra 

Cham. & Schlechtend.[not in Guianas] 

Spigelia schomburgkiana Benth. =Spigelia flemmingiana 

Strychnos pedunculata (A. DC.) Benth. =Strychnos 

bredemeyeri 

Strychnos rouhamon Benth. =Strychnos guianensis 

Strychnos solerederi Gilg =Strychnos mitscherlichii var. 

mitscherlichii 

LORANTHACEAE 


REFERENCES: 

Krause, K. 1932. Loranthaceae. In A. Pulle, 


Kuijt, J. 1976. Revision of the genus 

Oryctanthus. Bot. Jahrb. Syst. 95: 478-534. 

Kuijt, J. 1985. The genus Cladocolea. J. Arn. 

Arb. 56: 265-335. 


Loranthaceae (additions and corrections). In 


1(2): 295-300. 

Rizzini, C.T. 1978. Loranthaceae. In B. 



Bot. Gard. 29: 23-36. 

Rizzini, C.T. 1982. Loranthaceae. In Z. Luces 

de Febres, Flora de Venezuela 4(2): 7-316. 

Cladocolea elliptica Kuijt [SU] 

Cladocolea micrantha (Eichl.) Kuijt [GU] 

Cladocolea sandwithii (Maguire) Kuijt [GU] 

Gaiadendron punctatum (Ruiz & Pav.) G. Don [GU] 

Ixocactus clandestinus (Mart.) Kuijt [GU] 

Oryctanthus alveolatus (Kunth) Kuijt [GU SU FG] 

Oryctanthus florulentus (L.C. Rich.) Urb. [GU SU FG] 

Oryctanthus occidentalis (L.) Eichl. subsp. continentalis Kuijt 

[GU SU] 

Oryctanthus spicatus (Jacq.) Eichl. [GU] 

Oryctina myrsinites (Eichl.) Kuijt [GU SU FG] 

Phthirusa coarctata A.C. Smith [GU] 

Phthirusa disjectifolia (Rizzini) Kuijt [GU SU] 

Phthirusa guyanensis Eichl. [GU SU FG] 

Phthirusa monetaria Sandw. [GU] 

Phthirusa nitens (Mart.) Eichl. [GU] 

Phthirusa pycnostachya Eichl. [GU FG] 

Phthirusa pyrifolia (Kunth) Eichl. [GU SU FG] 

Phthirusa retroflexa (Ruiz & Pav.) Kuijt [FG] 

Phthirusa rufa (Mart.) Eichl. [GU SU] 

Phthirusa stelis (L.) Kuijt, comb. nov. ined. [GU SU FG] 

Psittacanthus clusiifolius (Willd.) Eichl. [GU SU?]

126 


Psittacanthus collum-cygni Eichl. [SU] 

Psittacanthus cordatus (Hoffmanns.) Blume [GU FG] 

Psittacanthus cucullaris (Lam.) Blume [GU? SU FG] 

Psittacanthus lamprophyllus Eichl. [FG] 

Psittacanthus lasianthus Sandw. [GU SU] 

Psittacanthus melinonii (Tiegh.) Engl. [FG] 

Psittacanthus peronopetalus Eichl. [FG] 

Psittacanthus plagiophyllus Eichl. [GU SU FG] 

Psittacanthus redactus Rizzini [FG] 

Struthanthus concinnus Mart. [GU SU] 

Struthanthus confertus Mart. [SU?] 

Struthanthus dichotrianthus Eichl. [GU SU FG] 

Struthanthus gracilis (Gleason) Steyerm. & Maguire [GU] 

Struthanthus marginatus (Desr.) Blume [GU] 

Struthanthus syringifolius Mart. [GU SU FG] 

Tripodanthus acutifolius (Ruiz & Pav.) Tiegh [to be expected 

in the Guianas] 

LORANTHACEAE - SYNONYMS AND 


Cladocolea clandestina (Mart.) Kuijt =Ixocactus clandestinus 

Furarium disjectifolium Rizzini =Phthirusa disjectifolia 

Loranthus patrisii DC. =Combretum sp. (Combretaceae) 

Oryctanthus botryostachys Eichl. =Oryctanthus alveolatus 

Oryctanthus ruficaulis (Poepp. & Endl.) Eichl. =Oryctanthus 

florulentus 

Phthirusa adunca (G. Mey.) Maguire =Phthirusa stelis 

Phthirusa angulata Krause =Phthirusa stelis 

Phthirusa myrsinites Eichl. =Oryctina myrsinites 

Phthirusa pauciflora Eichl. =Dendropemon pauciflorus 

(Swartz) Tiegh.[not in Guianas] 

Phthirusa phaeocladus Eichl. =Phthirusa guyanensis 

Phthirusa sandwithii Maguire =Cladocolea sandwithii 

Phthirusa savannarum Maguire =Oryctina myrsinites 

Phthirusa seitzii Krug & Urb. =Phthirusa stelis 

Phthirusa squamulosa Klotzsch ex Eichl. =Phthirusa rufa 

Phthirusa theobromae (Willd.) Eichl. =Phthirusa stelis 

Psittacanthus cupulifer (Kunth) Eichler =Psittacanthus 

cucullaris 

Struthanthus retroflexa (Ruiz & Pav.) Kuijt =Phthirusa stelis 

Struthanthus squamulosus Klotzsch ex Eichl. =Phthirusa rufa 

Struthanthus umbellatus Kuijt =Phthirusa pycnostachya 

LYTHRACEAE 

contributed by A. Lourteig (1992); additions and 

corrections by A. Lourteig (1996) 

REFERENCES: 

Cook, C.D.K. 1979. A revision of the genus 

Rotala. Boissiera 29: 1-156. 

Graham, S.A. 1985. A revision of Ammannia in 

the western hemisphere. J. Arnold Arb. 56: 

265-335. 

Jonker, F.P. 1941. Lythraceae. In A. Pulle, 


Lourteig, A. 1964. Lythraceae 

Austroamericanae, addenda et corrigenda. 

Sellowia 16: 119-162. 

SPERMATOPHYTA: Loranthaceae - Magnoliaceae 

Lourteig, A. 1986. Revision del genero 

Lafoensia Vandelli (Litraceas). Mem. Soc. 

Cienc. Nat. La Salle 45(123): 115-157. 

Lourteig, A. 1986. Revision de dos secciones 

del genero Cuphea P. Browne. Phytologia 

60(1): 17-55. 

Lourteig, A. 1987. Lythraceae 

Austroamericanae, addenda et corrigenda II. 

Sellowia 39: 15-48. 

Ammannia auriculata Willd. [to be expected in the Guianas] 

Ammannia coccinea Rottb. [to be expected in the Guianas] 

Ammannia latifolia L. [FG] 

Crenea maritima Aubl. [GU SU FG] 

Cuphea anagalloidea St. Hil. [GU] 

Cuphea antisyphilitica Kunth var. acutifolia Benth. [GU] 

Cuphea antisyphilitica Kunth var. antisyphilitica [GU] 

Cuphea blackii Lourteig [FG] 

Cuphea carthagenensis (Jacq.) Macbr. [SU FG] 

Cuphea dactylophora Koehne [GU] 

Cuphea melvilla Lindl. [GU FG] 

Cuphea micrantha Kunth [GU] 

Cuphea repens Koehne [to be expected in the Guianas] 

Cuphea rigidula Benth. [GU] 

Lafoensia vandelliana Cham. & Schlechtend. [to be expected 

in the Guianas] 

Lagerstroemia speciosa (L.) Pers. [GU] [cultivated or escaped 


Lawsonia inermis L. [GU SU] 

Rotala mexicana Cham. & Schlechtend. [SU FG] [to be 


Rotala ramosior (L.) Koehne [to be expected in the Guianas] 

LYTHRACEAE - SYNONYMS AND 


Crenea surinamensis (L.f.) Koehne =Crenea maritima 

Cuphea balsamona Cham. & Schlechtend. =Cuphea 

carthagenensis 

Cuphea gracilis Kunth [not in Guianas] 

Cuphea mimuloides Cham. & Schlechtend. var. guianensis 

Koehne =Cuphea anagalloidea 

Cuphea speciosa L. =Cuphea melvilla 

Lafoensia pacari St. Hil. [not in Guianas] 

REFERENCES: 

MAGNOLIACEAE 

Lozano Contreras, G. 1994. Dugandiodendron y 

Talauma (Magnoliaceae) en el Neotrópica. 

Academia de Ciencias Exactas, Fisicas y 

Naturales Coleccion Jorge Alvarez Lleras No. 

3. 

Steyermark, J.A. 1951. Magnoliaceae. In 

Botanical Exploration in Venezuela - I. 

Fieldiana Bot. 28(1): 233-234. 

Dugandiodendron ptaritepuianum (Steyerm.) G. Lozano-C. 

[Mt. Roraima]


MAGNOLIACEAE - SYNONYMS AND 


Magnolia ptaritepuiana Steyerm. =Dugandiodendron 

ptaritepuianum 

Magnolia roraimae Steyerm. =Dugandiodendron 

ptaritepuianum 

MALPIGHIACEAE 

reviewed by W.R. Anderson; Stigmaphyllon by 

C. Anderson (1992); additions and corrections 

by W.R. Anderson & C. Anderson (1996) 

REFERENCES: 

Anderson, C. 1993. The identities of the 

sericeous-leaved species of Stigmaphyllon in 

the Amazon region. Contr. Univ. Mich. Herb. 

19: 393-413. 

Anderson, W.R. 1981. Malpighiaceae. In B. 


Guayana Highland part 11. Mem. N.Y. Bot. 

Gard. 32: 1-391. 

Anderson, W.R. 1990. The taxonomy of 

Jubelina. Contr. Univ. Mich. Herb. 17: 21-37. 

Anderson, W.R. 1993. Notes on Neotropical 

Malpighiaceae - IV. Contr. Univ. Mich. 

Herb. 19: 355-392. 

Anderson, W.R. 1994. New species of Hiraea 

from the Guianas and adjacent Brazil. 

Brittonia 46(2): 126-133. 

Gates, B. 1982. Banisteriopsis, Diplopterys. 


Görts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 

1976. Malpighiaceae (additions and 

corrections). In J. Lanjouw & A.L. Stoffers, 


Jonker, F.P. 1939. Malpighiaceae (additions and 


2(1): 478-480. 

Kostermans, A.J.G.H. 1936. Malpighiaceae. In 


Niedenzu, F. 1928. Malpighiaceae. In A. 

Engler, Das Pflanzenreich IV. 141: 1-870. 

Banisteriopsis cinerascens (Benth.) B. Gates [GU] 

Banisteriopsis cristata (Griseb.) Cuatrec. [GU SU] 

Banisteriopsis lucida (L.C. Rich.) Small [GU SU FG] 

Banisteriopsis martiniana (Adr. Juss.) Cuatrec. var. martiniana 

[GU SU FG] 

Banisteriopsis mathiasiae (W.R. Anderson) W.R. Anderson 

[GU] 

Banisteriopsis muricata (Cav.) Cuatrec. [GU FG] 

Banisteriopsis nummifera (Adr. Juss.) B. Gates [FG] 

Banisteriopsis pulcherrima (Sandw.) B. Gates [GU] 

Banisteriopsis schwannioides (Griseb.) B. Gates [FG] 

Banisteriopsis wurdackii B. Gates [FG] 

Blepharandra fimbriata B. MacBryde [GU] 

Blepharandra hypoleuca (Benth.) Griseb. [GU] 

Bunchosia apiculata Huber [FG] 

Bunchosia argentea (Jacq.) DC. [GU SU? FG] 

Bunchosia decussiflora W.R. Anderson [FG] 

Bunchosia mollis Benth. [GU SU?] 

Bunchosia polystachia (Andr.) DC. [FG] 

Burdachia sphaerocarpa Adr. Juss. [GU] 

Byrsonima aerugo Sagot [GU SU FG] 

Byrsonima altissima DC. [FG] 

Byrsonima arthropoda Adr. Juss. [GU] 

Byrsonima chalcophylla Nied. var. carraoana (Steyerm.) W. 

Anderson [GU] 

Byrsonima chalcophylla Nied. var. chalcophylla [GU] 

Byrsonima christianeae W.R. Anderson [GU] 

Byrsonima coccolobifolia Kunth [GU SU?] 

Byrsonima concinna Benth. [GU] 

Byrsonima crassifolia (L.) Kunth [GU SU FG] 

Byrsonima crispa Adr. Juss. [FG] 

Byrsonima densa (Poir.) DC. [GU SU FG] 

Byrsonima dubia W.R. Anderson [GU] 

Byrsonima eugeniifolia Sandw. [GU SU] 

Byrsonima euryphylla Pilger [SU] 

Byrsonima fanshawei W.R. Anderson [GU] 

Byrsonima gymnocalycina Adr. Juss. [GU SU? FG?] 

Byrsonima incarnata Sandw. [GU] 

Byrsonima laevigata (Poir.) DC. [SU FG] 

Byrsonima pachypoda W.R. Anderson [GU] 

Byrsonima rubrobracteata W.R. Anderson [GU] 

Byrsonima schomburgkiana Benth. [GU] 

Byrsonima sericea DC. [FG] 

Byrsonima spicata (Cav.) DC. [GU SU FG] 

Byrsonima stipulacea Adr. Juss. [GU SU FG] 

Byrsonima surinamensis W.R. Anderson [SU] 

Byrsonima tillettii W.R. Anderson [GU] 

Byrsonima verbascifolia (L.) DC. [GU SU FG] 

Camarea affinis St. Hil. [GU SU] 

Coleostachys genipifolia Adr. Juss. [FG] 

Diplopterys pauciflora (G. Mey.) Nied. [GU FG] 

Heteropterys biglandulosa Adr. Juss. [FG] 

Heteropterys cristata Benth. [GU] 

Heteropterys lasseri W.R. Anderson [GU] 

Heteropterys macradena (DC.) W. Anderson [GU SU? FG] 

Heteropterys macrostachya Adr. Juss. [GU SU FG] 

Heteropterys maguirei W.R. Anderson [GU] 

Heteropterys mathewsana Adr. Juss. [SU] 

Heteropterys multiflora (DC.) Hochr. [GU SU FG] 

Heteropterys nervosa Adr. Juss. [GU SU FG] 

Heteropterys oligantha W.R. Anderson [GU] 

Heteropterys siderosa Cuatrec. [GU FG] 

Heteropterys subhelicina Nied. [GU] 

Hiraea adenophora Sandw. [GU] 

Hiraea affinis Miq. [GU SU FG] 

Hiraea fagifolia (DC.) Adr. Juss. [GU SU FG] 

Hiraea faginea (Swartz) Nied. [GU SU FG] 

Hiraea gracieana W.R. Anderson [SU FG] 

Hiraea longipes W.R. Anderson [FG] 

Hiraea longipilifera W.R. Anderson [GU] 

Hiraea morii W.R. Anderson [FG] 

Hiraea propinqua W.R. Anderson [SU FG] 

Hiraea quapara (Aubl.) Sprague [FG] 

Jubelina riparia Adr. Juss. [FG] 

Jubelina rosea (Miq.) Nied. [SU FG] 

Lophopterys euryptera Sandw. [GU] 

Lophopterys splendens Adr. Juss. [FG] 

Lophopterys surinamensis (Kosterm.) Sandw. [SU] 

Mascagnia divaricata (Kunth) Nied. [SU? FG] 

127 

SPERMATOPHYTA: Magnoliaceae - Malpighiaceae

128 


Mascagnia guianensis W.R. Anderson [GU SU FG] 

Mascagnia macrodisca (Triana & Planch.) Nied. [GU SU] 

Mascagnia microcarpa (Sandw.) W. Anderson [GU] 

Mascagnia sepium (Adr. Juss.) Griseb. [GU SU FG] 

Mascagnia sinemariensis (Aubl.) Griseb. [GU FG] 

Mascagnia surinamensis (Kosterm.) W. Anderson [GU SU] 

Mezia includens (Benth.) Cuatrec. [GU FG] 

Pterandra guianensis W.R. Anderson [GU] 

Pterandra sericea W.R. Anderson [GU] 

Spachea elegans (G. Mey.) Adr. Juss. [GU SU FG] 

Stigmaphyllon bannisterioides (L.) C. Anderson [GU SU FG] 

Stigmaphyllon ciliatum (Lam.) Adr. Juss. [GU?] 

Stigmaphyllon convolvulifolium Adr. Juss. [GU SU FG] 

Stigmaphyllon palmatum (Cav.) Adr. Juss. [FG] 

Stigmaphyllon puberum (L.C. Rich.) Adr. Juss. [GU SU FG] 

Stigmaphyllon sinuatum (DC.) Adr. Juss. [GU SU FG] 

Tetrapterys acutifolia Cav. [GU SU FG] 

Tetrapterys calophylla Adr. Juss. [SU FG] 

Tetrapterys crispa Adr. Juss. [GU SU FG] 

Tetrapterys discolor (G. Mey.) DC. [GU SU FG] 

Tetrapterys fimbripetala Adr. Juss. [GU SU FG?] 

Tetrapterys maranhamensis Adr. Juss. [GU? SU?] 

Tetrapterys megalantha W.R. Anderson [GU] 

Tetrapterys mucronata Cav. [GU SU? FG] 

Tetrapterys pusilla Steyerm. [GU] 

Tetrapterys rhodoptera Oliver [GU] 

Tetrapterys styloptera Adr. Juss. [GU SU FG] 

MALPIGHIACEAE - SYNONYMS AND 


Banisteria leptocarpa Benth. =Banisteriopsis martiniana 

Banisteria lucida Rich. =Banisteriopsis lucida 

Banisteria martiniana Adr. Juss. =Banisteriopsis martiniana 

Banisteria quapara Aubl. =Hiraea quapara 

Banisteria reticulata (Poir.) C. Robinson =Heteropterys 

multiflora 

Banisteria sinemariensis Aubl. =Mascagnia sinemariensis 

Banisteriopsis elegans (Triana & Planch.) Sandw. subsp. 

cordata (Nied.) Sandw. [not in Guianas] 

Banisteriopsis gardneriana (Adr. Juss.) W. Anderson & B. 

Gates [reported from FG in error] 

Brachypterys ovata (Cav.) Small =Stigmaphyllon 

bannisterioides 

Bunchosia maritima (Vell.) Macbride [not in Guianas] 

Bunchosia nitida (Jacq.) L.C. Rich. [not in Guianas] 

Bunchosia tuberculata DC. [dubious name of uncertain status] 

Byrsonima aubletii Kosterm. =Byrsonima altissima 

Byrsonima bracteolaris Benth. =Byrsonima concinna 

Byrsonima coriacea (Swartz) Kunth [not in Guianas] 

Byrsonima coriacea (Swartz) Kunth var. spicata (Cav.) Nied. 

=Byrsonima spicata 

Byrsonima cydoniifolia Adr. Juss. [not in Guianas] 

Byrsonima discolor Pilger =Byrsonima altissima 

Byrsonima obversa Miq. =Byrsonima laevigata 

Byrsonima poeppigiana Adr. Juss. [not in Guianas] 

Byrsonima rugosa Benth. =Byrsonima stipulacea 

Diplopterys includens (Benth.) Nied. =Mezia includens 

Diplopterys microcarpa Sandw. =Mascagnia microcarpa 

Diplopterys riparia (Adr. Juss.) Nied. =Jubelina riparia 

Diplopterys rosea (Miq.) Nied. =Jubelina rosea 

Diplopterys spruceana (Nied.) Nied. =Mascagnia bracteosa 

Griseb.[not in Guianas] 

SPERMATOPHYTA: Malpighiaceae - Malvaceae 

Dolichopterys surinamensis Kosterm. =Lophopterys 

surinamensis 

Heteropterys acutifolia Adr. Juss. =Heteropyterys orinocensis 

(H.B.K.) Adr. Juss.[not in Guianas] 

Heteropterys catoptera W.R. Anderson =Heteropterys 

subhelicina 

Heteropterys reticulata (Poir.) Nied. =Heteropterys multiflora 

Heteropterys suberosa Griseb. =Heteropterys nervosa 

Hiraea chrysophylla Adr. Juss. =Hiraea faginea 

Hiraea gaudichaudiana Adr. Juss. [not in Guianas] 

Hiraea multiradiata Adr. Juss. =Hiraea quapara 

Lophopterys splendens Adr. Juss. var. obovata Nied. 

=Lophopterys euryptera 

Malpighia altissima Aubl. =Byrsonima altissima 

Malpighia crassifolia Aubl. =Byrsonima crassifolia 

Malpighia moureila Aubl. =Byrsonima crassifolia 

Malpighia verbascifolia Aubl. =Byrsonima verbascifolia 

Mascagnia anisopetala (Adr. Juss.) Griseb. [not in Guianas] 

Mascagnia anisopetala (Adr. Juss.) Griseb. var. macrodisca 

(Triana & Planch.) Kosterm. =Mascagnia macrodisca 

Mascagnia leucanthele Griseb. [not in Guianas] 

Mascagnia multiglandulosa Nied. [not in Guianas] 

Mascagnia multiglandulosa Nied. var. surinamensis Kosterm. 

=Mascagnia surinamensis 

Mascagnia nitida (Kunth) Nied. [not in Guianas] 

Mascagnia ovatifolia (Kunth) Griseb. [not in Guianas] 

Mascagnia volubilis Nied. =Mascagnia sinemariensis 

Stigmaphyllon angustilobum Adr. Juss. [not in Guianas] 

Stigmaphyllon fulgens Adr. Juss. (nom. superfl.) 

=Stigmaphyllon sinuatum 

Stigmaphyllon hypoleucum Miq. =Stigmaphyllon sinuatum 

Stigmaphyllon latifolium Benth. =Stigmaphyllon 

convolvulifolium 

Stigmaphyllon martianum Adr. Juss. =Stigmaphyllon 

sinuatum 

Stigmaphyllon mathiasiae W.R. Anderson =Banisteriopsis 

mathiasiae 

Stigmaphyllon ovatum (Cav.) Cuatrec. =Stigmaphyllon 

bannisterioides 

Stigmaphyllon periplocifolium Adr. Juss. [not in Guianas] 

Stigmaphyllon purpureum Benth. =Stigmaphyllon sinuatum 

Stigmaphyllon sagittatum Adr. Juss. =Stigmaphyllon 

palmatum 

Stigmaphyllon tiliifolium Nied. [not in Guianas] 

Tetrapodenia glandifera Gleason =Burdachia sphaerocarpa 

Tetrapterys includens Benth. =Mezia includens 

Tetrapterys oleifolia (Benth.) Nied. [not in Guianas; type 

erroneously reported from GU] 

Tetrapterys squarrosa (Griseb.) Griseb. =Tetrapterys styloptera 

MALVACEAE 

contributed by M.J. Jansen-Jacobs (1992); 

additions and corrections by M.J. Jansen- 

Jacobs & L.J. Dorr (1996) 

REFERENCES: 

Clement, I.D. 1957. Studies in Sida. I. Contrib. 

Gray Herb. 180: 1-91. 

Jansen-Jacobs, M.J. 1986. Malvaceae (additions 

and corrections). In A.L. Stoffers & J.C.


Lindeman, Flora of Suriname 3(1-2): 259-276. 

Kearney, T.H. 1957. A tentative key to the 

South American species of Hibiscus L. Leafl. 

W. Bot. 8(7): 161-168. 


South American species of Pavonia Cav. 

Leafl. W. Bot. 8(10): 225-248. 


South American species of Sida L. Leafl. W. 

Bot. 8(11): 249-270. 

Krapovickas, A. & C.L. Cristobal. 1965. 

Revisión del genero Peltaea. Kurtziana 2: 

135-216. 

Uittien, H. 1932. Malvaceae. In A. Pulle, Flora 


Uittien, H. 1941. Malvaceae (additions and 


3(1): 433-435. 

Abelmoschus spp. [GU SU FG] [cultivated] 

Abutilon indicum (L.) Sweet [GU SU] [cultivated] 

Allosidastrum pyramidatum (Desp. ex Cav.) Krapov., Fryxell 

& D.M. Bates [SU] 

Bastardia viscosa (L.) Kunth [FG?] [introduced] 

Briquetia spicata (Kunth) Fryxell [GU SU FG] 

Cienfuegosia affinis (Kunth) Hochr. [GU] 

Cienfuegosia phlomidifolia (St. Hil.) Garcke [GU] 

Gaya occidentalis (L.) Sweet [GU] 

Gossypium barbadense L. [GU SU FG] [cultivated only?] 

Gossypium hirsutum L. [GU SU FG] [cultivated only?] 

Herissantia crispa (L.) Brizicky [GU] 

Hibiscus bifurcatus Cav. [GU SU FG] 

Hibiscus dimidiatus Schrank [GU] 

Hibiscus furcellatus Desr. [GU SU FG] 

Hibiscus guianensis Aubl. [FG] [name of uncertain 

application] 

Hibiscus ingratus Miq. [SU] 

Hibiscus mutabilis L. [SU] [cultivated?] 

Hibiscus pernambucensis Arruda [GU SU? FG] [=Hibiscus 

tiliaceus?] 

Hibiscus phoeniceus Jacq. [GU FG] [cultivated?] 

Hibiscus prunifolius F.G. Dietr. [GU?] 

Hibiscus radiatus Cav. [GU SU] [cultivated] 

Hibiscus rosa-sinensis L. [GU SU FG] [cultivated] 

Hibiscus sabdariffa L. [GU SU FG] [cultivated] 

Hibiscus schizopetalus (Dyer) Hook.f. [GU SU FG] 

[cultivated] 

Hibiscus sororius L.f. [GU SU FG] 

Hibiscus striatus Cav. var. lambertianus (Kunth) O. Blanch. 

[GU SU] 

Hibiscus tiliaceus L. [GU SU FG] [see also Hibiscus 

pernambucensis] 

Hibiscus trilobus Aubl. [FG] [name of uncertain application] 

Hibiscus varians Splitg. [GU?] 

Hibiscus verbasciformis Klotzsch ex Hochr. [GU SU FG] 

Malachra alceifolia Jacq. var. alceifolia [GU FG] 

Malachra alceifolia Jacq. var. rotundifolia (Schrank) Guerke 

[GU SU] 

Malachra fasciata Jacq. [GU SU FG] 

Malachra radiata (L.) L. [GU SU] 

Malvastrum coromandelianum (L.) Garcke [GU FG] 

Pavonia angustifolia Benth. [GU] 

Pavonia cancellata (L.) Cav. [GU SU FG] 

129 

Pavonia castanaefolia St. Hil. & Naud. [GU SU FG] 

Pavonia flavispina Miq. [FG?] 

Pavonia fruticosa (Mill.) Fawcett & Rendle [GU SU FG] 

Pavonia geminiflora Moric. [GU] 

Pavonia paludicola Nicolson [GU SU FG] 

Pavonia paniculata Cav. [GU] 

Pavonia piptocalyx Krapov. [FG] 

Pavonia schiedeana Steud. [GU SU FG] 

Peltaea riedelii (Guerke) Standl. [GU SU] 

Peltaea speciosa (Kunth) Standl. [GU SU] 

Peltaea surumuensis (Ulbr.) Krapov. & Cristóbal [possibly to 

be expected in the Guianas] 

Peltaea trinervis (K. Presl) Krapov. & Cristóbal [GU SU] 

Sida acuta Burm.f. [GU SU FG] 

Sida aggregata K. Presl [GU] 

Sida ciliaris L. [GU] 

Sida cordata (Burm.f.) Borssum Waalkes [SU] 

Sida cordifolia L. [GU SU FG] 

Sida glomerata Cav. [GU SU FG] 

Sida linifolia Cav. [GU SU FG] 

Sida rhombifolia L. [GU SU FG] 

Sida serrata Willd. [GU SU FG] 

Sida setosa Mart. ex Colla [GU SU FG] 

Sida spinosa L. [GU FG] 

Sida surumuensis Ulbr. [GU] 

Sida urens L. [GU SU FG] 

Sida viarum St. Hil. [GU SU] 

Sidastrum micranthum (St. Hil.) Fryxell [GU] 

Sidastrum quinquenervium (Duch. ex Triana & Planch.) E.G. 

Baker [GU SU FG] 

Thespesia lampas (Cav.) Dalz. & Gibs. [SU] [cultivated] 

Thespesia populnea (L.) Solander ex Correa [GU SU] 

[cultivated or escaped from cultivation] 

Urena lobata L. [GU SU FG] 

Wissadula contracta (Link) R.E. Fries [GU] 

Wissadula hirsuta Presl [GU] 

Wissadula patens (St. Hil.) Garcke [GU SU] 

Wissadula periplocifolia (L.) K. Presl ex Thwaites [GU SU] 

MALVACEAE - SYNONYMS AND 


Abutilon crispum (L.) Medic. =Herissantia crispa 

Hibiscus abelmoschus L. =Abelmoschus moschatus 

Medic.[cultivated] 

Hibiscus brasiliensis L. [not in Guianas] 

Hibiscus elatus Swartz =Hibiscus tiliaceus 

Hibiscus esculentus L. =Abelmoschus esculentus (L.) Moench 

[cultivated] 

Hibiscus lambertianus Kunth =Hibiscus striatus var. 

lambertianus 

Hibiscus lampas Cav. =Thespesia lampas 

Hibiscus spathulatus Garcke =Hibiscus verbasciformis 

Malachra alceifolia Jacq. var. fasciata (Jacq.) A. Robyns 

=Malachra fasciata 

Malachra capitata L. =Malachra alceifolia 

Pavonia julianae Uittien =Peltaea riedelii 

Pavonia racemosa Swartz [not in Guianas] 

Pavonia rosea Schlechtend. =Pavonia schiedeana 

Pavonia sessiliflora Kunth =Peltaea sessiliflora (Kunth) 

Standley [not in Guianas] 

Pavonia speciosa Kunth =Peltaea speciosa 

Pavonia spicata Cav. =Pavonia paludicola 

Pavonia spinifex Cav. [not in Guianas] 

SPERMATOPHYTA: Malvaceae

130 


Pavonia trinervis (K. Presl) A. Robyns =Peltaea trinervis 

Pavonia typhalaea Cav. =Pavonia fruticosa 

Pseudabutilon spicatum (Kunth) R.E. Fries =Briquetia spicata 

Sida guianensis K. Schum. =Sidastrum quinquenervium 

Sida micrantha St. Hil. =Sidastrum micranthum 

Sida quinquenervia (Duch. ex Triana & Planch.) Baker 

=Sidastrum quinquenervium 

Sida veronicifolia Lam. =Sida cordata 

Urena reticulata Cav. =Urena lobata 

Wissadula spicata (Kunth) K. Presl =Briquetia spicata 

MARANTACEAE 

contributed by H. Kennedy (1992) 

REFERENCES: 

Andersson, L. 1977. The genus Ischnosiphon. 

Opera Bot. 43: 1-113. 

Andersson, L. 1986. Revision of Maranta 

subgen. Maranta. Nord. J. Bot. 6(6): 729-756. 

Andersson, L. & H. Kennedy. 1986. Four new 

species of Calathea from French Guiana. 

Nordic J. Bot. 6(4): 447-455. 

Hagberg, M. 1990. Two new species of 

Monotagma from the Venezuelan Guayana. 



Marantaceae. In A. Pulle & J. Lanjouw, Flora 


Kennedy, H. 1995. Calathea maasiorum 

(Marantaceae), a new species from French 

Guiana and Surinam. Brittonia 47(2): 156- 

159. 

Calathea altissima (Poepp. & Endl.) Körn. [GU SU FG] 

Calathea casupito (Jacq.) Schult. [GU] 

Calathea cataractum K. Schum. [GU] 

Calathea comosa (L.f.) Lindl. [SU FG?] 

Calathea cyclophora Baker [GU] 

Calathea cylindrica (Roscoe) K. Schum. [FG] 

Calathea densa (Koch) Regel [SU?] 

Calathea dilabens L. Andersson & H. Kennedy [FG] 

Calathea elliptica (Roscoe) K. Schum. [GU SU FG] 

Calathea erecta L. Andersson & H. Kennedy [FG] 

Calathea fragilis Gleason [GU SU] 

Calathea grandis Petersen [GU SU FG] 

Calathea granvillei L. Andersson & H. Kennedy [GU SU FG] 

Calathea legrelleana (Linden) Regel [GU SU FG] 

Calathea lutea (Aubl.) Schult. [GU SU FG] 

Calathea maasiorum H. Kennedy [SU FG] 

Calathea mansonis Körn. [SU FG] 

Calathea micans (Mathieu) Körn. [GU SU FG] 

Calathea microcephala (Poepp. & Endl.) Körn. [FG] 

Calathea ovata (Nees & Mart.) Lindl. [SU? FG?] 

Calathea polytricha Baker [GU] 

Calathea propinqua (Poepp. & Endl.) Körn. [GU SU FG] 

Calathea splendida (Verschaff. ex Lem.) Regel [FG] 

Calathea squarrosa L. Andersson & H. Kennedy [SU FG] 

Calathea variegata Linden ex Körn. [GU SU] 

Calathea villosa Lindl. [GU SU FG] 

SPERMATOPHYTA: Malvaceae - Marantaceae 

Calathea zingiberina Körn. [GU SU FG] 

Hylaeanthe hexantha (Poepp. & Endl.) Jonker & Jonker [SU 

FG] 

Hylaeanthe polystachya (Pulle) Jonker-Verhoef & Jonker [SU 

FG?] 

Hylaeanthe unilateralis (Poepp. & Endl.) Jonker & Jonker [SU 

FG] 

Ischnosiphon arouma (Aubl.) Körn. [GU SU FG] 

Ischnosiphon bambusaceus (Poepp. & Endl.) Körn. [SU? FG?] 

Ischnosiphon centricifolius L. Andersson [GU FG] 

Ischnosiphon enigmaticus L. Andersson [GU FG] 

Ischnosiphon foliosus Gleason [GU] 

Ischnosiphon gracilis (Rudge) Körn. subsp. gracilis [SU FG] 

Ischnosiphon gracilis (Rudge) Körn. subsp. sandwithii L. 

Andersson [GU] 

Ischnosiphon leucophaeus (Poepp. & Endl.) Körn. subsp. 

leucophaeus [GU? SU FG] 

Ischnosiphon longiflorus K. Schum. subsp. angustifolius L. 

Andersson [GU SU] 

Ischnosiphon martianus Eichl. ex Petersen [SU FG] 

Ischnosiphon obliquus (Rudge) Körn. [GU SU FG] 

Ischnosiphon petiolatus (Rudge) L. Andersson [GU SU FG] 

Ischnosiphon puberulus Loes. var. scaber (Petersen) L. 

Andersson [GU SU FG] 

Ischnosiphon puberulus Loes. var. verruculosus (Macbride) L. 

Andersson [FG] 

Ischnosiphon ursinus L. Andersson [FG] 

Koernickanthe orbiculata (Körn.) L. Andersson [SU] 

Maranta amplifolia K. Schum. [GU SU FG] 

Maranta arundinacea Plumier ex L. [GU SU FG] 

Maranta friedrichsthaliana Körn. [FG] 

Maranta gibba J.E. Smith [GU SU] 

Maranta humilis Aubl. [GU? SU FG] 

Maranta protracta Miq. [GU SU] 

Maranta ruiziana Körn. [GU FG] 

Maranta rupicola L. Andersson [GU SU FG] 

Monophyllanthe oligophylla K. Schum. [SU FG] 

Monotagma contractum Huber [SU FG] 

Monotagma exile Hagberg, sp. nov. ined. [FG] 

Monotagma juruanum Loes. [FG] 

Monotagma laxum (Poepp. & Endl.) K. Schum. [GU SU FG] 

Monotagma ovatum Hagberg [GU] 

Monotagma plurispicatum (Körn.) K. Schum. var. 

plurispicatum [GU? SU FG] 

Monotagma secundum (Peters.) K. Schum. [GU SU] 

Monotagma spicatum (Aubl.) Macbr. [GU SU FG] 

Monotagma ulei Schum. ex Loes. [FG] 

Monotagma vaginatum Hagberg [FG] 

Myrosma cannifolia L.f. [GU SU FG?] 

Stromanthe tonckat (Aubl.) Eichl. [SU FG] 

Thalia geniculata L. [GU SU FG] 

MARANTACEAE - SYNONYMS AND 


Calathea albicans Brongn. ex K. Schum. =Calathea micans 

Calathea allouia (Aubl.) Lindl. =Calathea legrelleana 

Calathea strobilifera (Miq.) Körn. =Renealmia cernua 

(Zingiberaceae) 

Ischnosiphon surinamensis (Miq.) Körn. =Ischnosiphon 

leucophaeus 

Ischnosiphon violaceus Pulle =Ischnosiphon petiolatus 

Maranta allouia Aubl. =Calathea legrelleana


Maranta divaricata Roscoe [not in Guianas] 

Maranta orbiculata (Körn.) K. Schum. =Koernikanthe 

orbiculata 

Maranta strobilifera Miq. =Renealmia cernua (Zingiberaceae) 

Monotagma exannulatum K. Schum. =Monotagma laxum 

Monotagma guianense (Körn.) K. Schum. =Monotagma 

plurispicatum 

Monotagma parkeri (Roscoe) K. Schum. =Monotagma 

spicatum 

Monotagma parkeri (Roscoe) Schum. =Monotagma spicatum 

Myrosma polystachya Pulle =Hylaeanthe polystachya 

Phrynium strobiliferum (Miq.) K. Koch =Renealmia cernua 

(Zingiberaceae) 

Saranthe urceolata Peters. =Maranta humilis 

Thalia trichocalyx Gagnep. =Thalia geniculata 

MARCGRAVIACEAE 

reviewed by A.C. de Roon (1992); additions and 

corrections by A.C. de Roon & S. Dressler 

(1996) 

REFERENCES: 

Bailey, I.W. 1922. The pollination of 

Marcgravia: a classical case of ornithophily? 

Amer. J. Bot. 9: 370-384. 


Marcgraviaceae. In A. Pulle, Flora of 

Suriname 3(1): 373-385. 

Marcgravia coriacea Vahl [GU SU FG] 

Marcgravia magnibracteata Lanj. & van Heerdt [SU FG] 

Marcgravia parviflora L.C. Rich. ex Wittm. [GU SU FG] 

Marcgravia purpurea I.W. Bailey [GU] 

Norantea guianensis Aubl. [GU SU FG] 

Norantea pendula Lanj. & van Heerdt [GU SU FG] 

Norantea tepuiensis de Roon [GU] 

Souroubea guianensis Aubl. subsp. guianensis [GU SU FG] 

MARCGRAVIACEAE - SYNONYMS AND 


Marcgravia cuyuniensis I.W. Bailey =Marcgravia coriacea 

Marcgravia dasystachya Gilg ex Ule [nomen nudum] 

Marcgravia gracilis Sagot =Marcgravia parviflora 

Marcgravia picta Willd. [dubious species] 

Marcgravia roraimae Gilg ex Werderm. =Marcgravia coriacea 

Marcgravia roraimensis Gilg ex Ule [nomen nudum] 

Marcgravia umbellata L. [Guianan records misident.] 

MAYACACEAE 



REFERENCES: 

131 

Lanjouw, J. 1938. Mayacaceae. In A. Pulle, 


Lourteig, A. 1952. Mayacaceae. Notul. Syst. 14: 

234-248. 

Lourteig, A. 1971. Mayacaceae. In T. Lasser, 


Mayaca fluviatilis Aubl. [GU SU FG] 

Mayaca longipes Mart. ex Seub. [GU SU FG] 

Mayaca sellowiana Kunth [GU SU FG] 

MAYACACEAE - SYNONYMS AND 


Mayaca aubletii Michx. =Mayaca fluviatilis 

MELASTOMATACEAE 

contributed by J.J. Wurdack; Mouriri, Votomita 

by T. Morley (1992); reviewed by T. Morley 

(1996) 

REFERENCES: 

[for a more extensive bibliography, see Wurdack 

et al. 1993] 

Gleason, H.A. 1932. A synopsis of the 

Melastomataceae of British Guiana. Brittonia 

1: 127-184. 

Gleason, H.A. 1935. Melastomaceae. In A. 


Görts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 

1986. Melastomataceae (additions and 



Morley, T. 1976. Memecyleae 

(Melastomataceae). Flora Neotropica 

Monogr. 15: 1-295. 

Renner, S.S. 1989. Systematic studies in the 

Melastomataceae: Bellucia, Loreya, and 

Macairea. Mem. New York Bot. Gard. 50: 1- 

112. 

Wurdack, J.J. 1961. Melastomataceae. In B. 




Wurdack, J.J. 1973. Melastomataceae. In T. 

Lasser, Flora de Venezuela 8: 1-819. 

[Mouriri & Votomita contributed by T. 

Morley] 

Wurdack, J.J., T. Morley, & S. Renner. 1993. 

Melastomataceae. In A.R.A. Görts-van Rijn, 


Aciotis aequatorialis Cogn. [GU SU FG] 

Aciotis annua (DC.) Triana [GU SU FG] 

Aciotis circaeifolia (Bonpl.) Triana [GU] 

Aciotis dichotoma (Benth.) Cogn. [GU SU FG] 

SPERMATOPHYTA: Marantaceae - Melastomataceae

132 


Aciotis fragilis (DC.) Cogn. [GU SU FG] 

Aciotis laxa (DC.) Cogn. var. A [GU] 

Aciotis laxa (DC.) Cogn. var. B [GU SU FG] 

Aciotis laxa (DC.) Cogn. var. kappleriana (Naud.) Cogn. [GU 

SU FG] 

Aciotis laxa (DC.) Cogn. var. laxa [GU SU FG] 

Aciotis longifolia (DC.) Triana [FG] 

Aciotis ornata (Miq.) Gleason [GU SU FG] 

Aciotis purpurascens (Aubl.) Triana [GU SU FG] 

Aciotis rubricaulis (DC.) Triana [SU FG] 

Acisanthera bivalvis (Aubl.) Cogn. [GU SU FG] 

Acisanthera crassipes (Naud.) Wurdack [GU FG] 

Acisanthera hedyotidea (K. Presl) Triana [GU SU] 

Acisanthera limnobios (DC.) Triana [GU SU] 

Acisanthera nana Ule [GU] 

Acisanthera rosulans Huber [SU FG] 

Acisanthera tetraptera (Cogn.) Gleason [SU FG?] 

Acisanthera uniflora (Vahl) Gleason [GU SU FG] 

Adelobotrys adscendens (Swartz) Triana [GU SU FG] 

Adelobotrys ayangannensis Wurdack [GU] 

Adelobotrys ciliata (Naud.) Triana [GU? SU FG] 

Adelobotrys monticola Gleason [SU] 

Adelobotrys permixta Wurdack [GU] 

Adelobotrys scandens (Aubl.) DC. [SU FG] 

Adelobotrys spruceana Cogn. [FG] 

Appendicularia thymifolia (Bonpl.) DC. [GU SU FG] 

Bellucia grossularioides (L.) Triana [GU SU FG] 

Bellucia pentamera Naud. [GU] 

Boyania ayangannae Wurdack [GU] 

Chaetolepis anisandra Naud. [GU] 

Clidemia anisophylla DC. [GU SU FG] 

Clidemia aphanantha (Naud.) Sagot [FG] 

Clidemia attenuata (Naud.) Cogn. [GU FG] 

Clidemia ayangannensis Wurdack [GU] 

Clidemia bullosa DC. [GU SU FG] 

Clidemia buntingii Wurdack [GU] 

Clidemia capitata Benth. [GU] 

Clidemia capitellata (Bonpl.) D. Don var. capitellata [GU SU 

FG] 

Clidemia capitellata (Bonpl.) D. Don var. dependens (D. Don) 

Macbride [GU SU FG] 

Clidemia charadrophylla Tutin [GU] 

Clidemia conglomerata DC. [GU SU FG] 

Clidemia debilis Crueger [reported and perhaps to be 

expected, but no material seen] 

Clidemia dentata D. Don [GU SU FG] 

Clidemia epibaterium DC. [GU] 

Clidemia epiphytica (Triana) Cogn. [FG] 

Clidemia granvillei Wurdack [FG] 

Clidemia heptamera Wurdack [GU] 

Clidemia hirta (L.) D. Don var. elegans (Aubl.) Griseb. [GU 

SU FG] 

Clidemia hirta (L.) D. Don var. hirta [GU SU FG] 

Clidemia involucrata DC. [GU SU FG] 

Clidemia japurensis DC. var. heterobasis (DC.) Wurdack [GU 

SU FG] 

Clidemia japurensis DC. var. japurensis [GU SU FG] 

Clidemia kappleri Cogn. [GU SU] 

Clidemia laevifolia Gleason [GU SU FG] 

Clidemia micrantha Sagot [GU FG] 

Clidemia microthyrsa R.O. Williams [GU SU FG] 

Clidemia minutiflora (Triana) Cogn. [GU SU FG] 

Clidemia novemnervia (DC.) Triana [GU SU] 

Clidemia octona (Bonpl.) L.O. Williams subsp. guayanensis 

Wurdack [GU SU FG] 

SPERMATOPHYTA: Melastomataceae 

Clidemia ostentata Wurdack [GU] 

Clidemia pakaraimae Wurdack [GU] 

Clidemia petiolata DC. [SU FG] 

Clidemia pustulata DC. [GU SU FG] 

Clidemia pycnaster Tutin subsp. pycnaster [GU SU] 

Clidemia pycnaster Tutin subsp. robusta Wurdack [GU SU] 

Clidemia rubra (Aubl.) Mart. [GU SU FG] 

Clidemia sandwithii Wurdack [GU] 

Clidemia saulensis Wurdack [FG] 

Clidemia septuplinervia Cogn. [SU FG] 

Clidemia sericea D. Don [GU SU FG] 

Clidemia silvicola Gleason [GU SU FG] 

Clidemia stellipilis (Gleason) Wurdack [GU] 

Clidemia strigillosa (Swartz) DC. [GU SU] 

Clidemia tepuiensis Wurdack [GU] 

Clidemia urceolata DC. [GU FG] 

Clidemia venosa (Gleason) Wurdack [GU SU FG] 

Comolia ayangannae Wurdack [GU] 

Comolia latifolia (Aubl.) Cogn. [FG] [dubious species, no 

material found] 

Comolia microphylla Benth. [GU SU FG] 

Comolia smithii Wurdack [GU] 

Comolia vernicosa (Benth.) Triana [GU SU] 

Comolia villosa (Aubl.) Triana var. A [GU SU FG] 

Comolia villosa (Aubl.) Triana var. B [GU SU FG] 

Comolia villosa (Aubl.) Triana var. villosa [GU SU FG] 

Desmoscelis villosa (Aubl.) Naud. [GU SU FG] 

Ernestia blackii Brade & Markgr. [SU FG] 

Ernestia cataractae Tutin [GU] 

Ernestia confertiflora Wurdack [SU FG] 

Ernestia glandulosa Gleason [GU SU? FG] 

Ernestia granvillei Wurdack [SU FG] 

Ernestia minor Gleason [SU] 

Ernestia pullei Gleason [GU SU FG] 

Ernestia rubra Pulle [SU FG] 

Ernestia subglabra Wurdack [FG] 

Graffenrieda caryophyllea Triana [GU] 

Graffenrieda caudata Wurdack [GU] 

Graffenrieda intermedia Triana [GU] 

Graffenrieda irwinii Wurdack [GU] 

Graffenrieda obliqua Triana [GU] 

Graffenrieda sessilifolia Triana [GU] 

Graffenrieda weddellii Naud. [GU] 

Henriettea granulata Berg ex Triana [GU] 

Henriettea maroniensis Sagot [GU SU FG] 

Henriettea multiflora Naud. [GU SU FG] 

Henriettea patrisiana DC. [GU SU FG] 

Henriettea ramiflora (Swartz) DC. [GU SU FG] 

Henriettea stellaris O. Berg ex Triana [GU SU FG] 

Henriettea succosa (Aubl.) DC. [GU SU FG] 

Henriettella caudata Gleason [GU SU FG] 

Henriettella duckeana Hoehne [FG] 

Henriettella flavescens (Aubl.) Triana [SU FG] 

Henriettella ininiensis Wurdack [FG] 

Henriettella ovata Cogn. [GU] 

Leandra agrestis (Aubl.) Raddi [FG] 

Leandra chaetodon (DC.) Cogn. [GU] 

Leandra clidemioides (Naud.) Wurdack [GU FG] 

Leandra cremersii Wurdack [FG] 

Leandra cuneata (Mart.) Cogn. [GU] 

Leandra divaricata (Naud.) Cogn. [GU SU FG] 

Leandra edentula Gleason [GU] 

Leandra lindeniana (Naud.) Cogn. [GU] 

Leandra micrantha Wurdack [FG] 

Leandra micropetala (Naud.) Cogn. [SU FG]


Leandra paleacea Wurdack [SU FG] 

Leandra polyadena Ule [GU] 

Leandra procumbens Ule [GU] 

Leandra purpurea Gleason [GU FG] 

Leandra rufescens (DC.) Cogn. [GU SU FG] 

Leandra sanguinea Gleason subsp. sanguinea [GU] 

Leandra sanguinea Gleason subsp. tepuiensis Wurdack [GU 

SU] 

Leandra solenifera Cogn. [GU SU FG] 

Leandra verticillata (DC.) Cogn. [FG] 

Loreya arborescens (Aubl.) DC. [GU SU? FG] 

Loreya mespiloides Miq. [GU SU FG] 

Loreya spruceana Benth. ex Triana [GU] 

Loreya subrotundifolia (Wurdack) Renner [FG] 

Macairea lasiophylla (Benth.) Wurdack [GU] 

Macairea multinervia Benth. [GU] 

Macairea pachyphylla Benth. [GU SU] 

Macairea parvifolia Benth. [GU] 

Macairea thyrsiflora DC. [GU] 

Macrocentrum anfractum Wurdack [GU] 

Macrocentrum brevipedicellatum Wurdack [GU] 

Macrocentrum cristatum (DC.) Triana var. cristatum [GU SU 

FG] 

Macrocentrum cristatum (DC.) Triana var. microphyllum 

Cogn. [GU SU FG] 

Macrocentrum cristatum (DC.) Triana var. parviflorum (DC.) 

Cogn. [GU SU FG] 

Macrocentrum droseroides Triana [GU] 

Macrocentrum fasciculatum (DC.) Triana [GU SU FG] 

Macrocentrum fruticosum Gleason [GU SU FG] 

Macrocentrum gesneriaceum Sandw. [GU] 

Macrocentrum gracile Wurdack [GU] 

Macrocentrum latifolium Wurdack [FG] 

Macrocentrum minus Gleason [GU] 

Macrocentrum neblinae Wurdack [GU] 

Macrocentrum parvulum Gleason [SU] 

Macrocentrum repens (Gleason) Wurdack [GU] 

Macrocentrum stipulaceum Wurdack [GU] 

Macrocentrum vestitum Sandw. [GU FG] 

Maguireanthus ayangannae Wurdack [GU] 

Maieta guianensis Aubl. [GU SU FG] 

Maieta poeppigii Mart. ex Cogn. [GU SU FG] 

Marcetia taxifolia (St. Hil.) DC. [GU] 

Meriania crassiramis (Naud.) Wurdack [GU] 

Meriania sclerophylla (Naud.) Triana [GU] 

Meriania urceolata Triana [GU] 

Miconia acinodendron (L.) Sweet [GU SU FG] 

Miconia acuminata (Steud.) Naud. [SU FG] 

Miconia acutifolia Ule [GU] 

Miconia affinis DC. [GU SU FG] 

Miconia alata (Aubl.) DC. [GU SU FG] 

Miconia albicans (Swartz) Triana [GU SU FG] 

Miconia alborufescens Naud. [GU] 

Miconia aliquantula Wurdack [SU FG] 

Miconia alternans Naud. [GU SU FG] 

Miconia amacurensis Wurdack [GU FG] 

Miconia ampla Triana [GU SU FG] 

Miconia aplostachya (Bonpl.) DC. [GU SU FG] 

Miconia argyrophylla DC. subsp. argyrophylla [GU SU FG] 

Miconia argyrophylla DC. subsp. gracilis Wurdack [GU SU 

FG] 

Miconia bracteata (DC.) Triana [GU SU FG] 

Miconia brevipes Benth. [GU SU] 

Miconia bubalina (D. Don) Naud. [GU] 

Miconia cacatin (Aubl.) Renner [SU FG] 


133 

Miconia cacumina Wurdack [GU] 

Miconia campestris (Benth.) Triana [GU SU] 

Miconia centrodesma Naud. [GU SU? FG] 

Miconia ceramicarpa (DC.) Cogn. var. candolleana Cogn. 

[FG] 

Miconia ceramicarpa (DC.) Cogn. var. ceramicarpa [GU SU 

FG] 

Miconia chrysophylla (Rich.) Urb. [GU SU FG] 

Miconia ciliata (Rich.) DC. [GU SU FG] 

Miconia cowanii Wurdack [perhaps to be expected in FG] 

Miconia diaphanea Gleason [SU FG] 

Miconia dispar Benth. [FG] 

Miconia dodecandra (Desr.) Cogn. [GU SU] 

Miconia egensis Cogn. [GU SU FG] 

Miconia elata (Swartz) DC. [FG] 

Miconia eriodonta DC. [FG] 

Miconia fallax DC. [GU SU] 

Miconia fanshawei Wurdack [GU] 

Miconia fragilis Naud. [GU FG] 

Miconia fragrans Cogn. [GU] 

Miconia francavillana Cogn. [SU? FG] 

Miconia gratissima Benth. ex Triana [GU SU FG] 

Miconia holosericea (L.) DC. [GU SU FG] 

Miconia hypoleuca (Benth.) Triana [GU SU FG] 

Miconia ibaguensis (Bonpl.) Triana [GU SU] 

Miconia kappleri Naud. [SU FG] 

Miconia lappacea (DC.) Triana [SU FG] 

Miconia lasseri Gleason [GU SU] 

Miconia lateriflora Cogn. subsp. lateriflora [GU SU FG] 

Miconia lateriflora Cogn. subsp. monticellensis Wurdack [GU 

SU] 

Miconia lepidota DC. [GU SU FG] 

Miconia longifolia (Aubl.) DC. [GU SU FG] 

Miconia longispicata Triana [FG] 

Miconia macrothyrsa Benth. [GU SU FG] 

Miconia maguirei Gleason [GU] 

Miconia marginata Triana [GU] 

Miconia mariae Wurdack [GU] 

Miconia matthaei Naud. [GU SU FG] 

Miconia melinonis Naud. [FG] 

Miconia minutiflora (Bonpl.) DC. [SU FG] 

Miconia mirabilis (Aubl.) L.O. Williams [GU SU FG] 

Miconia mutabilis (DC.) Triana [FG] 

Miconia myriantha Benth. [GU SU FG] 

Miconia navioensis Wurdack [perhaps to be expected in FG] 

Miconia nervosa (Sm.) Triana [GU SU FG] 

Miconia oldemanii Wurdack [FG] 

Miconia phaeophylla Triana [GU SU FG] 

Miconia pileata DC. [FG] 

Miconia platypoda Gleason [SU] 

Miconia plukenetii Naud. [GU SU FG] 

Miconia poeppigii Triana [GU? SU FG] 

Miconia polita Gleason [GU] 

Miconia prasina (Swartz) DC. [GU SU FG] 

Miconia pseudoaplostachya Cogn. [FG] 

Miconia pubipetala Miq. [GU SU FG] 

Miconia punctata (Desr.) D. Don [GU SU FG] 

Miconia pyrifolia Naud. [GU SU FG?] 

Miconia racemosa (Aubl.) DC. [GU SU FG] 

Miconia radulaefolia (Benth.) Naud. [GU] 

Miconia regelii Cogn. [GU FG?] 

Miconia rhytidophylla Naud. [to be expected in GU, SU] 

Miconia roraimensis Ule [GU] 

Miconia rubiginosa (Bonpl.) DC. [GU SU] 

Miconia rufescens (Aubl.) DC. [GU SU FG]

134 


Miconia ruficalyx Gleason [GU SU FG] 

Miconia rugosa Triana [GU SU] 

Miconia rupestris Ule [GU] 

Miconia sagotiana Cogn. [GU SU FG] 

Miconia sarmentosa Cogn. [SU FG] 

Miconia sastrei Wurdack [FG] 

Miconia serialis DC. [GU SU FG] 

Miconia serrulata (DC.) Naud. [GU SU FG] 

Miconia silicicola Gleason [GU] 

Miconia splendens (Swartz) Griseb. [GU SU FG] 

Miconia stenostachya DC. [GU SU] 

Miconia stephananthera Ule [GU SU FG?] 

Miconia superba Ule [GU] 

Miconia tetraspermoides Wurdack [GU SU FG] 

Miconia theaezans (Bonpl.) Cogn. [to be expected in GU] 

Miconia tillettii Wurdack [GU SU FG] 

Miconia tinifolia Naud. var. roraimensis Wurdack [GU] 

Miconia tomentosa (Rich.) D. Don ex DC. [GU SU FG] 

Miconia traillii Cogn. [GU SU FG] 

Miconia trimera Wurdack [SU FG] 

Miconia trinervia (Swartz) D. Don ex Loud. [FG] 

Miconia tschudyoides Cogn. [SU FG] 

Microlicia benthamiana Triana ex Cogn. [GU] 

Monochaeum bonplandii (Kunth) Naud. [GU] 

Mouriri acutiflora Naud. [GU SU FG] 

Mouriri angulicosta Morley [SU FG] 

Mouriri brevipes Hook. [GU SU] 

Mouriri collocarpa Ducke [SU FG] 

Mouriri crassifolia Sagot [GU? SU FG] 

Mouriri duckeana Morley [GU SU FG] 

Mouriri dumetosa Cogn. [FG] 

Mouriri francavillana Cogn. [FG] 

Mouriri grandiflora DC. [GU SU FG] 

Mouriri guianensis Aubl. [GU SU FG] 

Mouriri huberi Cogn. [GU SU FG] 

Mouriri myrtifolia Spruce ex Triana [to be expected in GU] 

Mouriri nervosa Pilger [SU FG] 

Mouriri nigra (DC.) Morley [GU SU FG] 

Mouriri oligantha Pilger [FG] 

Mouriri sagotiana Triana [GU SU FG] 

Mouriri sideroxylon Sagot ex Triana [GU SU FG] 

Mouriri subumbellata Triana [FG] 

Mouriri vernicosa Naud. [GU SU FG] 

Mouriri viridicosta Morley [FG] 

Myriaspora egensis DC. [GU SU FG] 

Myrmidone macrosperma (Mart.) Mart. [GU] 

Nepsera aquatica (Aubl.) Naud. [GU SU FG] 

Ochthephilus repentinus Wurdack [GU] 

Ossaea coarctiflora Wurdack [FG] 

Ossaea quadrisulca (Naud.) Wurdack [FG] 

Pachyloma coriaceum (DC.) [GU] 

Phainantha laxiflora (Triana) Gleason [GU] 

Phainantha maguirei Wurdack [GU] 

Phainantha myrteoloides Wurdack [to be expected in GU] 

Pterogastra divaricata (Bonpl.) Naud. [GU] 

Pterolepis glomerata (Rottb.) Miq. [GU SU FG] 

Pterolepis pumila (Bonpl.) Cogn. [GU FG] 

Pterolepis trichotoma (Rottb.) Cogn. [GU SU FG] 

Rhynchanthera dichotoma (Desr.) DC. [GU] 

Rhynchanthera grandiflora (Aubl.) DC. [GU SU FG] 

Rhynchanthera hispida Naud. [GU? SU] 

Rhynchanthera serrulata (L.C. Rich.) DC. [GU SU FG] 

Salpinga glandulosa (Gleason) Wurdack [to be expected in 

GU] 

Salpinga paleacea Wurdack [GU] 


Siphanthera cordifolia (Benth.) Gleason [GU] 

Siphanthera foliosa (Naud.) Wurdack [GU?] 

Siphanthera hostmannii Cogn. [GU SU] 

Tibouchina aspera Aubl. var. aspera [GU SU FG] 

Tibouchina aspera Aubl. var. asperrima Cogn. [GU SU FG] 

Tibouchina fraterna N.E. Brown [GU] 

Tibouchina geitneriana (Schlechtend.) Cogn. [GU] 

Tibouchina gracilis (Bonpl.) Cogn. [GU] 

Tibouchina multiflora (Gardn.) Cogn. [FG] 

Tococa aristata Benth. [GU SU FG?] 

Tococa desiliens Gleason [GU] 

Tococa erythrophylla (Ule) Wurdack [GU] 

Tococa guianensis Aubl. [GU SU FG] 

Tococa nitens (Benth.) Triana [GU] 

Tococa subciliata (DC.) Triana [GU] 

Topobea ferruginea Gleason [perhaps to be expected in GU] 

Topobea parasitica Aubl. [SU FG] 

Tryssophyton merumense Wurdack [GU] 

Votomita guianensis Aubl. [GU SU FG] 

Votomita sp. aff. monadelpha (Ducke) Morley [SU] 

MELASTOMATACEAE - SYNONYMS AND 


Aciotis acutiflora (Mart.) Triana [not in Guianas] 

Aciotis cordata Macb. [not in Guianas] 

Aciotis dichotoma (Benth.) Cogn. var. anomala (Miq.) Cogn. 

=Aciotis dichotoma 

Aciotis discolor D. Don =Aciotis purpurascens 

Aciotis dysophylla (Benth.) Triana =Aciotis annua 

Aciotis fragilis (DC.) Cogn. var. lancifolia Cogn. =Aciotis 

fragilis 

Aciotis herbacea Cogn. =Aciotis ornata 

Aciotis indecora (Bonpl.) Triana var. glabrescens Cogn. 

=Aciotis laxa var. A 

Aciotis indecora (Bonpl.) Triana var. sagotiana Cogn. 

=Aciotis laxa var. B 

Aciotis purpurascens (Aubl.) Triana var. pellucida (DC.) 

Cogn. =Aciotis purpurascens 

Acisanthera beccabunga (Mart. ex Naud.) Triana =Acisanthera 

bivalvis 

Acisanthera boissieriana Cogn. =Acisanthera hedyotidea 

Acisanthera brevifolia (DC.) Griseb. =Acisanthera bivalvis 

Acisanthera gracilis Ule =Acisanthera crassipes 

Acisanthera inundata (DC.) Triana [not in Guianas] 

Acisanthera inundata (DC.) Triana var. pusilla Cogn. 

=Acisanthera limnobios 

Acisanthera lasiophylla (Benth.) Gleason =Macairea 

lasiophylla 

Acisanthera recurva (L.C. Rich.) Griseb. =Acisanthera 

uniflora 

Acisanthera trivalvis (Aubl.) Cogn. =Acisanthera bivalvis 

Adelobotrys guianensis (DC.) Gleason =Adelobotrys 

adscendens 

Adelobotrys laxiflora Triana =Phainantha laxiflora 

Adelobotrys monticola Gleason subsp. occidentalis Wurdack 

=Adelobotrys monticola 

Apatitia blakeoides Desv. =Bellucia grossularioides 

Arthrostemma glomeratum (Rottb.) Cham. =Pterolepis 

glomerata 

Arthrostemma ladanoides (Rich.) DC. =Pterolepis trichotoma 

Arthrostemma latifolium Don [dubious species, type not 

found] 

Arthrostemma villosum (Aubl.) DC. =Comolia villosa


Bellucia arborescens (Aubl.) Baill. =Loreya arborescens 

Bellucia aubletii Naud. =Bellucia grossularioides 

Bellucia cacatin (Aubl.) Sagot =Miconia cacatin 

Bellucia hostmannii Naud. =Bellucia grossularioides 

Bellucia mespiloides (Miq.) Macbr. =Loreya mespiloides 

Bellucia nervosa Raf. =Bellucia grossularioides 

Bellucia quinquenervia (Aubl.) Karst. =Bellucia 

grossularioides 

Bellucia subrotundifolia Wurdack =Loreya subrotundifolia 

Blakea cacatin (Aubl.) D. Don ex DC. =Miconia cacatin 

Blakea quinquenervia Aubl. =Bellucia grossularioides 

Blakea triplinervia L.f. =Bellucia grossularioides 

Brachypremna petiolata Gleason =Ernestia pullei 

Capitellaria benthamii Naud. =Clidemia capitata 

Centronia crassiramis (Naud.) Triana =Meriania crassiramis 

Chaenopleura fulva (Rich.) Griseb. =Miconia chrysophylla 

Chaenopleura hypoleuca Benth. =Miconia hypoleuca 

Chaenopleura longifolia (Aubl.) Griseb. =Miconia longifolia 

Chaetogastra callichaeta Benth. =Pterolepis glomerata 

Chaetogastra glomerata (Rottb.) Benth. =Pterolepis glomerata 

Chaetogastra hypericoides DC. =Desmoscelis villosa 

Chaetogastra lasiophylla Benth. =Macairea lasiophylla 

Charianthus spp. [Caribbean, none in Guianas] 

Chitonia fothergilla (Desr.) D. Don 

Clidemia affinis (Naud.) Cogn. =Clidemia pustulata 

Clidemia agrestis (Aubl.) D. Don =Leandra agrestis 

Clidemia benthamiana Miq. =Clidemia hirta var. hirta 

Clidemia berbiceana DC. =Miconia acinodendron 

Clidemia biformis Gleason =Clidemia anisophylla 

Clidemia bonplandii Cogn. =Clidemia debilis 

Clidemia bracteata DC. =Miconia bracteata 

Clidemia campestris Benth. =Miconia campestris 

Clidemia candolleana Cogn. =Clidemia bullosa 

Clidemia coccinea DC. =Miconia ceramicarpa var. 

ceramicarpa 

Clidemia coriacea (Naud.) Cogn. =Clidemia tepuiensis 

Clidemia depauperata DC. =Leandra agrestis 

Clidemia dependens D. Don =Clidemia capitellata var. 

dependens 

Clidemia divaricata Naud. =Leandra divaricata 

Clidemia drosera Sagot =Clidemia micrantha? 

Clidemia elegans (Aubl.) D. Don =Clidemia hirta var. elegans 

Clidemia glabrata Steud. =Miconia ibaguensis 

Clidemia heteroclita Naud. =Clidemia japurensis var. 

japurensis 

Clidemia lappacea DC. =Miconia lappacea 

Clidemia maculata Benth. =Miconia ibaguensis 

Clidemia miconioides Benth. =Miconia ibaguensis 

Clidemia mutabilis DC. =Miconia mutabilis 

Clidemia novemnervia (DC.) Triana var. affinis (Naud.) 

Wurdack =Clidemia pustulata 

Clidemia petiolata DC. var. angustifolia Cogn. =Clidemia 

petiolata 

Clidemia petiolata DC. var. colorata Cogn. =Clidemia 

petiolata 

Clidemia radulaefolia Benth. =Miconia radulaefolia 

Clidemia rariflora Benth. =Clidemia bullosa 

Clidemia rhamnifolia Naud. var. macrodon Naud. =Leandra 

divaricata 

Clidemia rubra (Aubl.) Mart. var. biacuta (Naud.) Cogn. 

=Clidemia sericea 

Clidemia smithii Gleason =Clidemia laevifolia x Clidemia 

hirta? 

Clidemia spicata (Aubl.) DC. =Clidemia capitellata var. 

dependens 


135 

Clidemia surinamensis Miq. =Clidemia novemnervia 

Clidemia tiliifolia DC. =Clidemia hirta var. tiliifolia (DC.) 

Macbr.[not in Guianas] 

Clidemia trichodes DC. =Miconia bracteata 

Clidemia tristis (Naud.) Sagot =Miconia bracteata 

Clidemia umbonata DC. =Clidemia strigillosa 

Clidemia violacea DC. =Miconia ceramicarpa var. 

ceramicarpa 

Comolia angustifolia Gleason =Comolia villosa var. A 

Comolia hirtella Naud. =Comolia villosa var. villosa 

Comolia kuhlmannii Hoehne =Comolia villosa var. A 

Comolia lythrarioides (Steud.) Naud. =Comolia villosa var. B 

Comolia neglecta Cogn. =Comolia villosa var. A 

Comolia purpurea Miq. =Comolia villosa var. villosa 

Comolia surinamensis Miq. =Comolia villosa var. B 

Comolia tetraptera Cogn. =Acisanthera tetraptera 

Comolia veronicaefolia Benth. =Comolia villosa var. A 

Conostegia acuminata Steud. =Miconia acuminata 

Conostegia icosandra (Sw.) Urb. [not in Guianas] 

Conostegia petiolata Gleason =Conostegia montana (Sw.) 


Copedesma nitens Gleason =Miconia ruficalyx 

Coryphadenis abnormis (Naud.) Morley =Votomita guianensis 

Davya ciliata Naud. =Adelobotrys ciliata 

Davya crassiramis Naud. =Meriania crassiramis 

Davya guianensis DC. =Adelobotrys adscendens 

Davya scandens (Aubl.) Naud. =Adelobotrys scandens 

Davya sclerophylla Naud. =Meriania sclerophylla 

Decaraphe floribunda Miq. =Miconia acinodendron 

Decaraphe fockeana Miq. =Miconia acinodendron 

Decaraphe hostmannii Steud. =Miconia serrulata 

Diolena repens Gleason =Macrocentrum repens 

Diplochita florida DC. =Miconia mirabilis 

Diplochita fothergilla (Desr.) DC. =Miconia mirabilis 

Diplochita leucocephala DC. =Miconia serrulata 

Diplochita mucronata (Desr.) DC. =Miconia holosericea 

Diplochita parviflora Benth. =Miconia pubipetala 

Diplochita serrulata DC. var. latifolia DC. =Miconia serrulata 

Diplochita tomentosa (Rich.) Griseb. =Miconia tomentosa 

Fothergilla guianensis Aubl. =Miconia mirabilis 

Fothergilla mirabilis Aubl. =Miconia mirabilis 

Graffenrieda ovalifolia Naud. =Graffenrieda weddellii 

Graffenrieda stenopetala Ule =Graffenrieda intermedia 

Hartigia barbigera (DC.) Miq. =Miconia ciliata 

Hartigia oblongifolia Miq. =Miconia racemosa 

Hartigia spectabilis Miq. =Miconia racemosa 

Henriettea succosa (Aubl.) DC. var. guianensis Gleason 

=Henriettea ramiflora 

Henriettea succosa (Aubl.) DC. var. kappleriana Steud. ex 

Naud. =Henriettea succosa 

Henriettea surinamensis Miq. =Henriettea ramiflora 

Henriettea trinervia Naud. =Henriettea ramiflora 

Henriettella longistyla Ule =Henriettella ovata 

Henriettella patrisiana (DC.) Naud. =Henriettea patrisiana 

Henriettella venosa Gleason =Clidemia venosa 

Jucunda tomentosa (Rich.) Benth. =Miconia tomentosa 

Lasiandra tibouchina (Desr.) Naud. =Tibouchina aspera var. 

aspera 

Leandra anisophylla (DC.) Cogn. =Clidemia anisophylla 

Leandra longicoma Cogn. [not in Guianas] 

Leandra montana Gleason =Aciotis laxa var. kappleriana 

Leandra pulverulenta (DC.) Cogn. =Miconia aliquantula 

Leandra rhamnifolia (Naud.) Cogn. var. macrodon (Naud.) 

Cogn. =Leandra divaricata

136 


Leandra rufescens (DC.) Cogn. var. asperiuscula (DC.) Cogn. 

=Leandra rufescens 

Leandra scabra DC. =Leandra melastomoides Raddi [not in 


Leiostegia vernicosa Benth. =Comolia vernicosa 

Loreya acutifolia O. Berg ex Triana =Loreya arborescens 

Loreya flavescens (Aubl.) Naud. =Henriettella flavescens 

Macairea aspera N.E. Brown =Macairea thyrsiflora 

Macairea calvescens Naud. =Macairea pachyphylla 

Macrocentrum montanum Gleason =Macrocentrum cristatum 

var. microphyllum 

Maieta dispar Miq. =Maieta guianensis 

Meisneria cordifolia Benth. =Siphanthera cordifolia 

Meisneria microlicioides Naud. =Siphanthera cordifolia 

Melastoma agrestis Aubl. =Leandra agrestis 

Melastoma alata Aubl. =Miconia alata 

Melastoma aquatica Aubl. =Nepsera aquatica 

Melastoma arborescens Aubl. =Loreya arborescens 

Melastoma aromatica Vahl =Tibouchina aspera var. aspera 

Melastoma bivalvis Aubl. =Acisanthera bivalvis 

Melastoma brunnea Vahl =Henriettea succosa 

Melastoma cacatin Aubl. =Miconia cacatin (Aubl.) Renner 

Melastoma chrysophylla Rich. =Miconia chrysophylla 

Melastoma ciliata Rich. =Miconia ciliata 

Melastoma diffusa Desr. =Acisanthera uniflora 

Melastoma elegans Aubl. =Clidemia hirta var. elegans 

Melastoma flavescens Aubl. =Henrietella flavescens 

Melastoma fothergilla Desr. =Miconia mirabilis 

Melastoma fulva Rich. =Miconia chrysophylla 

Melastoma grandiflora Aubl. =Rhynchanthera grandiflora 

Melastoma holosericea L. =Miconia holosericea 

Melastoma laevigata Aubl. =Miconia prasina 

Melastoma longifolia Aubl. =Miconia longifolia 

Melastoma maieta Desr. =Maieta guianensis 

Melastoma mucronata Desr. =Miconia holosericea 

Melastoma parviflora Aubl. =Miconia prasina 

Melastoma physophora Vahl =Tococa guianensis 

Melastoma pleurocarpum E. Mey. =Miconia acinodendron 

Melastoma purpurascens Aubl. =Aciotis purpurascens 

Melastoma purpurea Willd. =Aciotis purpurascens 

Melastoma racemosa Aubl. =Miconia racemosa 

Melastoma rubra Aubl. =Clidemia rubra 

Melastoma rufescens Aubl. =Miconia rufescens 

Melastoma scandens Aubl. =Adelobotrys scandens 

Melastoma spicata Aubl. =Clidemia capitellata var. dependens 

Melastoma succosa Aubl. =Henriettea succosa 

Melastoma tamonea Sw. =Miconia mirabilis 

Melastoma tibouchina Desr. =Tibouchina aspera var. aspera 

Melastoma tococo Desr. =Tococa guianensis 

Melastoma tomentosa Rich. =Miconia tomentosa 

Melastoma trivalvis Aubl. =Acisanthera bivalvis 

Melastoma villosa Aubl. =Desmoscelis villosa 

Meliandra monadelpha (Ducke) Morley =Votomita 

monadelpha 

Miconia agrestis (Aubl.) Baill. =Leandra agrestis 

Miconia argyrophylla DC. var. attenuata Cogn. =Miconia 

argyrophylla subsp. argyrophylla 

Miconia attenuata DC. =Miconia prasina 

Miconia attenuata DC. var. subquintuplinervia DC. =Miconia 

prasina 

Miconia axilliflora Naud. =Miconia acuminata 

Miconia barbigera DC. =Miconia ciliata 

Miconia berbiceana (DC.) Naud. =Miconia acinodendron 

Miconia bracteolaris Naud. =Miconia eriodonta 


Miconia bracteolaris Naud. var. cardiophora Naud. =Miconia 

eriodonta 

Miconia ceramicarpa (DC.) Cogn. var. navioensis Wurdack 

=Miconia ceramicarpa var. candolleana 

Miconia ceramicarpa (DC.) Cogn. var. violacea (DC.) Cogn. 

=Miconia ceramicarpa var. ceramicarpa 

Miconia chamissonis Naud. [not in Guianas] 

Miconia clidemioides Steud. =Miconia acinodendron 

Miconia cristulata Naud. =Miconia prasina 

Miconia demerarensis Gleason =Miconia bracteata 

Miconia disparilis (Standl.) R.O. Williams =Miconia 

lateriflora 

Miconia eriodonta DC. var. oblongifolia DC. =Miconia 

eriodonta 

Miconia eriophora Miq. =Miconia ibaguensis 

Miconia eriopoda Benth. =Miconia acinodendron 

Miconia erythrophylla Ule =Tococa erythrophylla 

Miconia erythropila Steud. =Miconia ceramicarpa var. 

ceramicarpa 

Miconia florida (DC.) Naud. =Miconia mirabilis 

Miconia fockeana Miq. =Miconia nervosa 

Miconia fothergilla (Desr.) Naud. =Miconia mirabilis 

Miconia fulva (Rich.) DC. =Miconia chrysophylla 

Miconia fulva (Rich.) DC. var. angustifolia Cogn. =Miconia 

chrysophylla 

Miconia fulva (Rich.) DC. var. aubletiana (Naud.) Cogn. 

=Miconia chrysophylla 

Miconia globuliflora (Rich.) Cham. [not in Guianas] 

Miconia guianensis (Aubl.) Cogn. =Miconia mirabilis 

Miconia heterochroa Miq. =Miconia albicans 

Miconia holosericea (L.) DC. var. mucronata (Desr.) Cogn. 

=Miconia holosericea 

Miconia holosericea (L.) DC. var. subquintuplinervia Cogn. 

=Miconia holosericea 

Miconia hypargyrea Miq. =Miconia stenostachya 

Miconia ibaguensis (Bonpl.) Triana var. glabrata (Steud.) 

Cogn. =Miconia ibaguensis 

Miconia iodopila Steud. =Miconia ceramicarpa var. 

ceramicarpa 

Miconia laevigata (L.) DC. [not in Guianas] 

Miconia leschenaultiana (Naud.) Sagot =Miconia ceramicarpa 

var. ceramicarpa 

Miconia leucocephala (DC.) Naud. =Miconia serrulata 

Miconia longifolia (Aubl.) DC. var. aubletiana Naud. 

=Miconia chrysophylla 

Miconia longistyla Steud. =Miconia argyrophylla subsp. 

argyrophylla 

Miconia macrophylla Steud. =Miconia prasina 

Miconia macrophylla (D. Don) Triana var. hostmannii (Steud.) 

Cogn. =Miconia serrulata 

Miconia macrophylla (D. Don) Triana var. latifolia (DC.) 


Miconia macrophylla (D. Don) Triana var. leucocephala (DC.) 


Miconia miqueliana Naud. =Miconia acinodendron 

Miconia mucronata (Desr.) Naud. =Miconia holosericea 

Miconia mucronulata Ule =Miconia prasina 

Miconia naudiniana Miq. =Miconia lepidota 

Miconia nemoralis Naud. =Miconia prasina 

Miconia nitens Benth. =Tococa nitens 

Miconia panicularis Gleason =Miconia egensis 

Miconia parviflora (Benth.) Cogn. =Miconia pubipetala 

Miconia plebeia Naud. =Miconia brevipes 

Miconia plumosa Gleason =Miconia rugosa


Miconia prasina (Sw.) DC. var. attenuata (DC.) Cogn. 

=Miconia prasina 

Miconia pterophora Miq. =Miconia alata 

Miconia pteropoda Benth. =Miconia prasina 

Miconia purpurascens (Aubl.) DC. =Aciotis purpurascens 

Miconia racemosa (Aubl.) DC. var. barbigera (DC.) Cogn. 

=Miconia ciliata 

Miconia racemosa (Aubl.) DC. var. ciliata (Rich.) Griseb. 

=Miconia ciliata 

Miconia revoluta Benth. =Miconia prasina 

Miconia robusta (Sagot) Cogn. =Miconia tschudyoides 

Miconia rufescens (Aubl.) DC. var. grandifolia Cogn. 

=Miconia rufescens 

Miconia spondylantha DC. =Miconia nervosa 

Miconia spondylantha DC. var. leschenaultiana Naud. 

=Miconia ceramicarpa var. ceramicarpa 

Miconia surinamensis Gleason =Miconia poeppigii 

Miconia tomentella Cogn. =Miconia serialis 

Miconia trichodes (DC.) Triana =Miconia bracteata 

Miconia verticilliflora Steud. ex Naud. =Miconia pubipetala 

Miconia virgulata Gleason =Miconia ciliata 

Microlicia bivalvis (Aubl.) DC. =Acisanthera bivalvis 

Microlicia brevifolia DC. =Acisanthera bivalvis 

Microlicia bryanthoides Oliver =Microlicia benthamiana 

Microlicia recurva (Rich.) DC. =Acisanthera uniflora 

Mouriri abnormis Naud. =Votomita guianensis 

Mouriri acutiflora Naud. var. amblyodon Naud. =Mouriri 

acutiflora 

Mouriri acutiflora Naud. var. oligantha Gleason =Mouriri 

collocarpa 

Mouriri anomala Pulle =Mouriri crassifolia 

Mouriri pendulifolia Cogn. =Mouriri acutiflora 

Mouriri plasschaertii Pulle =Mouriri nigra 

Mouriri polyantha Miq. =Mouriri guianensis 

Mouriri princeps Naud. =Mouriri grandiflora 

Myriaspora decipiens Naud. =Myriaspora egensis 

Myriaspora surinamensis Steud. =Myriaspora egensis 

Noterophila beccabunga Mart. ex Naud. =Acisanthera bivalvis 

Noterophila brevifolia (DC.) Naud. =Acisanthera bivalvis 

Noterophila pusilla Naud. =Acisanthera limnobios 

Onoctonia calcarata Naud. =Acisanthera crassipes 

Onoctonia crassipes Naud. =Acisanthera crassipes 

Onoctonia pauciflora Naud. =Acisanthera crassipes 

Osbeckia glomerata (Rottb.) DC. =Pterolepis glomerata 

Ossaea duckeana Hoehne =Clidemia epibaterium 

Ossaea epibaterium (DC.) Triana =Clidemia epibaterium 

Ossaea flavescens (Aubl.) DC. =Henriettella flavescens 

Oxymeris agrestis (Aubl.) Triana =Leandra agrestis 

Oxymeris anisophylla (DC.) Triana =Clidemia anisophylla 

Oxymeris asperiuscula (DC.) Triana =Leandra rufescens 

Oxymeris pulverulenta (DC.) Triana =Miconia aliquantula 

Oxymeris rufescens (DC.) Triana =Leandra rufescens 

Petaloma mouriri Sw. =Mouriri guianensis 

Platycentrum clidemioides Naud. =Leandra clidemioides 

Pleroma tibouchinum (Desr.) Triana =Tibouchina aspera var. 

aspera 

Poteranthera calcarata (Naud.) Triana =Acisanthera crassipes 

Poteranthera crassipes (Naud.) Triana =Acisanthera crassipes 

Poteranthera pauciflora (Naud.) Triana =Acisanthera crassipes 

Pterolepis hispida (Rich.) Gleason =Pterolepis trichotoma 

Pterolepis hostmannii Steud. =Pterolepis glomerata 

Pterolepis ladanoides (Rich.) Triana =Pterolepis trichotoma 

Pterolepis lasiophylla (Benth.) Triana =Macairea lasiophylla 

Pterolepis pumila (Bonpl.) Cogn. var. procera Cogn. 

=Pterolepis pumila 


137 

Pterolepis pusilla Miq. =Pterolepis trichotoma 

Rhexia aquatica (Aubl.) Sw. =Nepsera aquatica 

Rhexia aspera (Aubl.) Willd. =Tibouchina aspera var. aspera 

Rhexia bicornis Rottb. =Acisanthera uniflora 

Rhexia bivalvis (Aubl.) Vahl =Acisanthera bivalvis 

Rhexia glomerata Rottb. =Pterolepis glomerata 

Rhexia hispida Rich. =Pterolepis trichotoma 

Rhexia hypericoides Willd. =Desmoscelis villosa 

Rhexia ladanoides Rich. =Pterolepis trichotoma 

Rhexia latifolia Aubl. =Comolia latifolia 

Rhexia recurva Rich. =Acisanthera uniflora 

Rhexia serrulata Rich. =Rhynchanthera serrulata 

Rhexia strigosa Willd. ex Spreng. =Pterolepis trichotoma 

Rhexia thymifolia Bonpl. =Appendicularia thymifolia 

Rhexia trichotoma Rottb. =Pterolepis trichotoma 

Rhexia trivalvis (Aubl.) Vahl =Acisanthera bivalvis 

Rhexia villosa Aubl. =Comolia villosa 

Rhexia villosissima Rich. =Desmoscelis villosa 

Rhynchanthera acuminata Benth. =Rhynchanthera grandiflora 

Rhynchanthera acuminata Benth. var. sublaevis Cogn. 

=Rhynchanthera grandiflora 

Rhynchanthera adenophora Miq. =Acisanthera uniflora 

Rhynchanthera ambigua Naud. =Rhynchanthera grandiflora 

Rhynchanthera betulifolia Cogn. =Rhynchanthera grandiflora 

Rhynchanthera grandiflora (Aubl.) DC. var. microphylla 

Naud. =Rhynchanthera grandiflora 

Rhynchanthera intermedia Ule =Rhynchanthera grandiflora 

Rhynchanthera latifolia Cogn. [not in Guianas] 

Rhynchanthera microphylla Gleason =Rhynchanthera 

grandiflora 

Rhynchanthera modesta Naud. =Rhynchanthera serrulata 

Rhynchanthera parviflora Naud. =Rhynchanthera serrulata 

Rhynchanthera virgata Wurdack =Rhynchanthera hispida 

Sagraea aphanantha (Naud.) Triana =Clidemia aphanantha 

Sagraea capitata (Benth.) Triana =Clidemia capitata 

Sagraea cognata Steud. =Clidemia rubra 

Sagraea coriacea (Naud.) Triana =Clidemia tepuiensis 

Sagraea petiolata (DC.) Triana =Clidemia petiolata 

Sagraea rubra (Aubl.) Triana =Clidemia rubra 

Salpinga cristata DC. =Macrocentrum cristatum 

Salpinga fasciculata DC. =Macrocentrum fasciculatum 

Salpinga parviflora DC. =Macrocentrum cristatum var. 

parviflorum 

Sarmentaria decora Naud. =Adelobotrys adscendens? 

Siphanthera capitata Gleason =Siphanthera hostmannii 

Siphanthera jenmanii Gleason =Siphanthera cordifolia 

Siphanthera microlicioides (Naud.) Cogn. =Siphanthera 

cordifolia 

Siphanthera paraensis Huber =Siphanthera hostmannii 

Siphantheropsis williamii Brade =Macairea lasiophylla 

Spennera anomala Miq. =Aciotis dichotoma 

Spennera aquatica (Aubl.) Mart. ex DC. =Nepsera aquatica 

Spennera dichotoma Benth. =Aciotis dichotoma 

Spennera fragilis DC. =Aciotis fragilis 

Spennera grandifolia Miq. =Aciotis purpurascens 

Spennera hydrophila Miq. =Nepsera aquatica 

Spennera kappleriana Naud. =Aciotis laxa var. kappleriana 

Spennera laxa DC. =Aciotis laxa 

Spennera longifolia DC. =Aciotis longifolia 

Spennera ornata Miq. =Aciotis ornata 

Spennera pellucida DC. =Aciotis purpurascens

138 


Spennera quintuplinervia Miq. =Clidemia japurensis var. 

japurensis 

Spennera tetraptera Miq. =Aciotis annua 

Staphidiastrum aphananthum Naud. =Clidemia aphanantha 

Staphidiastrum attenuatum Naud. =Clidemia attenuata 

Staphidiastrum cognatum (Steud.) Naud. =Clidemia rubra 

Staphidiastrum coriaceum Naud. =Clidemia tepuiensis 

Staphidiastrum pulverulentum (DC.) Naud. =Miconia 

aliquantula 

Staphidiastrum rubrum (Aubl.) Naud. =Clidemia rubra 

Staphidiastrum rubrum (Aubl.) Naud. var. biacutum Naud. 

=Clidemia sericea 

Staphidium affine Naud. =Clidemia pustulata 

Staphidium agreste (Aubl.) Naud. =Leandra agrestis 

Staphidium benthamianum (Miq.) Naud. =Clidemia hirta var. 

hirta 

Staphidium bracteosum Naud. =Clidemia capitellata var. 

dependens 

Staphidium conglomeratum (DC.) Naud. =Clidemia 

conglomerata 

Staphidium elegans (Aubl.) Naud. =Clidemia hirta var. 

elegans 

Staphidium glabratum (Steud.) Naud. =Miconia ibaguensis 

Staphidium hostmannii Naud. =Clidemia dentata 

Staphidium involucratum (DC.) Naud. =Clidemia involucrata 

Staphidium lappaceum (DC.) Naud. =Miconia lappacea 

Staphidium mutabile (DC.) Naud. =Miconia mutabilis 

Staphidium petiolatum (DC.) Naud. =Clidemia petiolata 

Staphidium spicatum (Aubl.) Naud. =Clidemia capitellata var. 

dependens 

Staphidium triste Naud. =Miconia bracteata 

Tamonea guianensis Aubl. =Miconia mirabilis 

Tetrameris trivalvis Naud. =Macairea multinervia 

Tetrameris villosa (Aubl.) Naud. =Comolia villosa 

Tetrazygia discolor (L.) DC. [not in Guianas] 

Tibouchina lasiophylla (Benth.) Cogn. =Macairea lasiophylla 

Tibouchina multinervia (Benth.) Baill. =Macairea multinervia 

Tibouchina pseudomollis Gleason =Tibouchina geitneriana 

Tococa castrata Naud. =Tococa subciliata 

Tococa maieta D. Don =Maieta guianensis 

Tococa roreimi Benth. =Tococa guianensis 

Tococa subnuda Benth. =Tococa subciliata 

Topobea "guianensis Aubl." [invalid name from herbarium 

records, = Topobea parasitica] 

Topobea cuspidata Gleason =Topobea parasitica 

Tschudya asperiuscula DC. =Leandra rufescens 

Tschudya berbiceana (DC.) Griseb. =Miconia acinodendron 

Tschudya pulverulenta DC. =Miconia aliquantula 

Tschudya robusta Sagot =Miconia tschudyoides 

Tschudya rufescens DC. =Leandra rufescens 

Tschudya spondylantha (DC.) Griseb. =Miconia nervosa 

Tschudya verticillata DC. =Leandra verticillata 

Uranthera hostmannii Naud. =Acisanthera uniflora 

Uranthera recurva (Rich.) Naud. =Acisanthera uniflora 

Webera quinquenervia (Aubl.) G.F. Gmel. =Bellucia 

grossularioides 

REFERENCES: 

MELIACEAE 

SPERMATOPHYTA: Melastomataceae - Meliaceae 

Pennington, T.D., B.T. Styles, & D.A.H. Taylor. 

1981. Meliaceae. Flora Neotropica Monogr. 

28: 1-470. 

Pennington, T.D. & A.R.A. Görts-van Rijn. 

1984. Meliaceae. In A.L. Stoffers & J.C. 


Azadirachta indica Adr. Juss. [cultivated, naturalized] 

Cabralea canjerana (Vell.) Mart. subsp. canjerana [GU FG?] 

Carapa guianensis Aubl. [GU SU FG] 

Carapa procera A. DC. [GU? SU FG] 

Cedrela odorata L. [GU SU FG] 

Guarea carinata Ducke [SU FG] 

Guarea cinnamomea Harms [FG?] 

Guarea costata Adr. Juss. [SU FG] 

Guarea glabra Vahl [GU SU FG?] 

Guarea gomma Pulle [GU? SU FG] 

Guarea grandifolia A. DC. [GU SU FG] 

Guarea guidonia (L.) Sleumer [GU SU FG] 

Guarea kunthiana Adr. Juss. [GU SU FG] 

Guarea macrophylla Vahl subsp. pachycarpa (C. DC.) 

Pennington [SU? FG] 

Guarea macrophylla Vahl subsp. pendulispica (C. DC.) 

Pennington [GU] 

Guarea michel-moddei Pennington & Mori [FG] 

Guarea pubescens (L.C. Rich.) Adr. Juss. subsp. pubescens 

[GU SU FG] 

Guarea pubescens (L.C. Rich.) Adr. Juss. subsp. pubiflora 

(Adr. Juss.) Pennington [SU FG?] 

Guarea scabra Adr. Juss. [GU SU FG] 

Guarea silvatica C. DC. [GU? SU FG] 

Guarea truncifolia C. DC. [GU] 

Melia azedarach L. [cultivated, naturalized] 

Swietenia macrophylla King [FG] 

Trichilia cipo (Adr. Juss.) C. DC. [GU SU? FG] 

Trichilia elegans Adr. Juss. [GU] 

Trichilia euneura C. DC. [GU? SU FG] 

Trichilia lecointei Ducke [GU] 

Trichilia lepidota Mart. subsp. leucastera (Sandw.) Pennington 

[GU SU] 

Trichilia martiana C. DC. [GU? SU FG] 

Trichilia mazanensis Macbr. [to be expected in the Guianas] 

Trichilia micrantha Benth. [GU SU FG] 

Trichilia micropetala Pennington [FG] 

Trichilia moschata Swartz [GU] 

Trichilia pallida Swartz [GU SU FG] 

Trichilia pleeana (Adr. Juss.) C. DC. [GU] 

Trichilia quadrijuga Kunth subsp. quadrijuga [GU SU FG] 

Trichilia rubra C. DC. [GU] 

Trichilia schomburgkii C. DC. subsp. schomburgkii [GU SU 

FG] 

Trichilia septentrionalis C. DC. [GU SU FG] 

Trichilia surinamensis (Miq.) C. DC. [GU SU FG] 

Trichilia surumuensis C. DC. [GU] 

MELIACEAE - SYNONYMS AND 


Carapa surinamensis Miq. =Carapa procera 

Cedrela guianensis Adr. Juss. =Cedrela odorata 

Granatum surinamensis (Miq.) O. Kuntze =Carapa procera 

Guarea affinis Adr. Juss. =Guarea pubescens subsp. pubescens


Guarea aubletii Adr. Juss. =Guarea guidonea 

Guarea concinna Sandw. =Guarea pubescens subsp. pubescens 

Guarea davisii Sandw. =Guarea pubescens subsp. pubescens 

Guarea guara (Jacq.) P.G. Wilson =Guarea guidonia 

Guarea pallida C. DC. =Guarea pubescens 

Guarea pedicellata C. DC. =Guarea silvatica 

Guarea pubiflora Adr. Juss. =Guarea pubescens subsp. 

pubiflora [this subspecies doubtfully in Guianas] 

Guarea richardiana Adr. Juss. =Guarea pubescens subsp. 

pubescens 

Guarea schomburgkii C. DC. =Guarea glabra 

Guarea surinamensis Miq. ex C. DC. [nomen nudum] =Guarea 

guidonia 

Guarea trichilioides L. =Guarea guidonia 

Moschoxylum propinquum Miq. =Trichilia quadrijuga 

Portesia echinocarpa de Vriese =Trichilia pallida 

Trichilia acariaeantha Harms =Trichilia micrantha 

Trichilia alternans C. DC. =Trichilia surinamensis 

Trichilia brachystachya Klotzsch ex C. DC. =Trichilia pallida 

Trichilia compacta A.C. Smith =Trichilia quadrijuga subsp. 

quadrijuga 

Trichilia cuneifolia Pulle =Trichilia schomburgkii 

Trichilia davisii Sandw. =Trichilia pallida 

Trichilia echinocarpa (de Vriese) Walp. =Trichilia pallida 

Trichilia fuscescens Radlk. =Trichilia martiana 

Trichilia guianensis Klotzsch ex C. DC. =Trichilia rubra 

Trichilia leucastera Sandw. =Trichilia lepidota subsp. 

leucastera 

Trichilia maynasiana C. DC. =Trichilia surinamensis 

Trichilia poeppigii C. DC. =Trichilia schomburgkii 

Trichilia propinqua (Miq.) C. DC. =Trichilia quadrijuga subsp. 

quadrijuga 

Trichilia roraimana C. DC. =Trichilia micrantha 

Trichilia smithii C. DC. =Trichilia pleeana 

Trichilia stelligera Radlk. =Trichilia euneura 

Trichilia subsessilifolia C. DC. =Trichilia schomburgkii 

Trichilia trinitensis Adr. Juss. =Trichilia pallida 

MENDONCIACEAE 

reviewed by D. Wasshausen (1992); reviewed by 

D. Wasshausen (1996) 

REFERENCES: 

Bremekamp, C.E.B. 1938. Acanthaceae. In A. 


Wasshausen, D.C. 1989. New species of 

Mendoncia from the Venezuelan Guayana. 


Mendoncia aspera (Ruiz & Pav.) Nees [GU SU FG] 

Mendoncia bivalvis (L.f.) Merr. [GU SU FG] 

Mendoncia crenata Lindau [FG] 

Mendoncia glabra (Poepp. & Endl.) Nees [GU FG] 

Mendoncia hoffmannseggiana Nees [GU SU FG] 

Mendoncia squamuligera Nees [GU SU FG] 

MENDONCIACEAE - SYNONYMS AND 


Mendoncia perrottetiana Nees =Mendoncia bivalvis 

Mendoncia schomburgkiana Nees =Mendoncia 

hoffmannseggiana 

MENISPERMACEAE 

contributed by P. Hiepko (1992); additions and 

corrections by P. Hiepko (1996) 

REFERENCES: 

SPERMATOPHYTA: Meliaceae - Menispermaceae 

139 

Diels, L. 1934. Menispermaceae. In A. Pulle, 


Barneby, R.C. 1970. Revision of Neotropical 

Menispermaceae tribe Tinosporeae. Mem. 

New York Bot. Gard. 20(2): 81-158. 

Barneby, R.C. & B.A. Krukoff. 1971. 

Supplementary notes in American 

Menispermaceae. VIII. A generic survey of 

the American Triclisieae and 

Anomospermeae. Mem. New York Bot. 

Gard. 22(2): 1-89. 

Jansen-Jacobs, M.J. 1976. Menispermaceae 



440. 

Jonker, F.P. 1939. Menispermaceae (additions 


Suriname 2(1): 476-477. 

Krukoff, B.A. & R.C. Barneby. 1938. Studies of 

American Menispermaceae, with special 

reference to species used in preparation of 

arrow-poisons. Brittonia 3: 1-74. [See also 

numerous articles by R.C. Barneby, B.A. 

Krukoff, and H.N. Moldenke in a series of 

papers published from 1941 through 1971 in 

Bull. Torrey Bot. Club and Mem. New York 

Bot. Gard.] 

Abuta barbata Miers [GU SU FG] 

Abuta bullata Moldenke [GU FG] 

Abuta candollei Triana & Planch. [GU SU FG] 

Abuta grandifolia (Mart.) Sandw. [GU SU FG] 

Abuta grisebachii Triana & Planch. [FG?] 

Abuta imene (Mart.) Eichl. [GU SU FG] 

Abuta obovata Diels [GU SU FG] 

Abuta rufescens Aubl. [GU SU FG] 

Abuta sandwithiana Krukoff & Barneby [SU FG] 

Abuta solimoesensis Krukoff & Barneby [FG] 

Anomospermum chloranthum Diels subsp. chloranthum [FG] 

Anomospermum chloranthum Diels subsp. confusum Krukoff 

& Barneby [FG] 

Anomospermum grandifolium Eichl. [GU] 

Anomospermum steyermarkii Krukoff & Barneby [FG] 

Borismene japurensis (Mart.) Barneby [SU?] 

Caryomene foveolata Barneby & Krukoff [SU] 

Caryomene glaucescens (Moldenke) Barneby & Krukoff [FG] 

Caryomene olivascens Barneby & Krukoff [FG] 

Cissampelos andromorpha DC. [GU SU FG]

140 


Cissampelos fasciculata Benth. [GU SU FG?] 

Cissampelos ovalifolia DC. [GU SU] 

Cissampelos pareira L. [GU SU] 

Curarea candicans (L.C. Rich.) Barneby & Krukoff [GU SU 

FG] 

Disciphania moriorum Barneby, sp. nov. ined. [FG] 

Disciphania unilateralis Barneby [FG] 

Elephantomene eburnea Barneby & Krukoff [SU FG] 

Hyperbaena domingensis (DC.) Benth. [GU SU FG] 

Odontocarya tamoides (DC.) Miers var. canescens (Miers) 

Barneby [GU SU] 

Odontocarya tamoides (DC.) Miers var. tamoides [GU FG?] 

Odontocarya wullschlaegelii (Eichl.) Barneby [GU SU FG] 

Orthomene prancei Barneby & Krukoff [FG] 

Orthomene schomburgkii (Miers) Barneby & Krukoff [GU SU 

FG] 

Orthomene verruculosa (Krukoff & Barneby) Barneby & 

Krukoff [FG] 

Sciadotenia cayennensis Benth. [GU SU FG] 

Sciadotenia duckei Moldenke [FG] 

Sciadotenia eichleriana Moldenke [FG] 

Sciadotenia sagotiana (Eichl.) Diels [GU SU FG] 

Telitoxicum duckei (Diels) Moldenke [FG] 

Telitoxicum inopinatum Moldenke [GU SU FG] 

Telitoxicum krukovii Moldenke [SU] 

Telitoxicum minutiflorum (Diels) Moldenke [FG?] 

MENISPERMACEAE - SYNONYMS AND 


Abuta amara Aubl. =Aristolochia amara (Aristolochiaceae) 

Abuta concolor Poepp. & Endl. =Abuta grandifolia 

Abuta klugii Moldenke =Abuta imene 

Abuta macrocarpa Moldenke =Anomospermum grandifolium 

Abuta pullei Diels =Curarea candicans 

Abuta splendida Krukoff & Moldenke =Abuta rufescens 

Abuta tomentosa Sagot =Sciadotaenia sagotiana 

Abuta umbellata Sagot =Sciadotenia cayennensis 

Anomospermum schomburgkii Miers =Orthomene 

schomburgkii 

Chondrodendron candicans (L.C. Rich.) Sandw. =Curarea 

candicans 

Odontocarya paupera (Griseb.) Diels =Odontocarya tamoides 

var. canescens 

Sciadotenia candicans Diels =Curarea candicans 

Somphoxylon wullschlaegelii Eichl. =Odontocarya 

wullschlaegelii 

REFERENCES: 

MENYANTHACEAE 

Jonker, F.P. 1936. Menyanthaceae. In A. Pulle, 


Nymphoides humboldtianum (Kunth) Kuntze [GU SU FG] 

[=Nymphoides indica (L.) Kuntze] 

MENYANTHACEAE - SYNONYMS AND 


SPERMATOPHYTA: Menispermaceae - Mimosaceae 

Nymphoides indica (L.) Kuntze subsp. occidentalis A. Raynal 

=Nymphoides humboldtianum 

MIMOSACEAE 

contributed by R.C. Barneby & J. Grimes; Inga 

by O. Poncy (1992); additions and corrections 

by R.C. Barneby & J. Grimes; Inga by O. 

Poncy (1996) 

REFERENCES: 

Barneby, R.C. 1991. Sensitivae Censitae. A 

description of the genus Mimosa 

(Mimosaceae) in the New World. Mem. New 

York Bot. Gard. 65: 1-835. 

Barneby, R.C. & J. Grimes (in press). 

Mimosaceae. In A.R.A. Görts-van Rijn, Flora 

of the Guianas (series A). 





Hopkins, H.C.F. 1986. Parkia. Flora Neotropica 

Monogr. 43: 1-123. 

Jansen-Jacobs, M.J. 1976. Mimosaceae 



653. 

Kleinhoonte, A. 1940. Mimosaceae. In A. Pulle, 





Poncy, O. 1985. Le genre Inga 

(Léguminoseuses, Mimosoideae) en Guyane 

française. Systématique, morphologie des 

formes juveniles, ecologie. Mém. Mus. Nat. 

Hist. Nat. (n.s.), Sér. B 31: 1-153. 

Poncy, O. 1991. Le genre Inga dans les 

Guyanes: notes phytogeographiques. C.R. 

Soc. Biogéogr. 67(2): 95-111. 

Poncy, O. 1991. Deux nouvelles espèces de Inga 

(Mimosaceae) et notes nomenclaturales et 

taxonomiques sur trois autres espèces. Bull. 

Mus. natl. Hist. nat., Paris, sér. 4, 13, sect. B, 

Adansonia, 3-4: 147-154. 

Abarema barbouriana (Standl.) Barneby & Grimes, comb. nov. 

ined. var. arenaria (Ducke) Barneby & Grimes, comb. & 

stat. nov. ined [GU] 

Abarema barbouriana (Standl.) Barneby & Grimes, comb. nov. 

ined. var. barbouriana [GU FG] 

Abarema commutata Barneby & Grimes, sp. nov. ined. [GU 

SU] 

Abarema curvicarpa (Irwin) Barneby & Grimes var. 

curvicarpa [GU FG] 

Abarema ferruginea (Benth.) Pittier [GU?] 

Abarema floribunda (Spruce ex Benth.) Barneby & Grimes, 

comb. ined. [GU?] 

Abarema gallorum Barneby & Grimes, sp. nov. ined. [FG]


Abarema jupunba (Willd.) Britton & Killip var. jupunba [FG] 

Abarema jupunba (Willd.) Britton & Killip var. trapezifolia 

(Vahl) Barneby & Grimes, comb. nov. ined. [GU SU FG] 

Abarema laeta (Poepp. & Endl.) Barneby & Grimes, comb. 

ined. [SU FG] 

Abarema longipedunculata (Irwin) Barneby & Grimes [to be 


Abarema mataybifolia (Sandw.) Barneby & Grimes, comb. 

nov. ined. [GU FG] 

Acacia alemquerensis Huber [to be expected in the Guianas] 

Acacia articulata Ducke [SU] 

Acacia farnesiana (L.) Willd. [GU SU FG] 

Acacia floribunda (Spruce ex Benth.) Barneby & Grimes [GU] 

Acacia macracantha Willd. [GU FG] [introduced] 

Acacia polyphylla DC. [GU] 

Acacia tenuifolia (L.) Willd. var. producta Grimes [SU FG] 

Acacia tenuifolia (L.) Willd. var. tenuifolia [GU SU FG] 

Adenanthera pavonina L. [SU] [cultivated, naturalized] 

Albizia barinensis Cardenas [GU] 

Albizia glabripetala (Irwin) G.P. Lewis & P.E. Owen [GU] 

Albizia lebbek (L.) Benth. [GU SU FG] [cultivated, 

naturalized] 

Albizia niopoides (Spruce ex Benth.) Burkart [GU] 

Albizia subdimidiata (Splitg.) Barneby & Grimes, comb. nov. 

ined. var. minor Barneby & Grimes, var. nov. ined. [GU] 

Albizia subdimidiata (Splitg.) Barneby & Grimes, comb. nov. 

ined. var. subdimidiata [GU SU] 

Anadenanthera peregrina (L.) Speg. [GU SU FG] 

[naturalized?] 

Balizia pedicellaris (DC.) Barneby & Grimes, gen. & comb. 

nov. ined. [GU SU FG] 

Calliandra calothyrsus Meisn. [cultivated, perhaps escaping] 

Calliandra coriacea (Willd.) Benth. [GU SU] [to be expected 

in FG] 

Calliandra glomerulata Karsten [GU] 

Calliandra hymenaeodes (Pers.) Benth. [GU SU FG] 

Calliandra rigida Benth. [GU] 

Calliandra riparia Pittier [GU] 

Calliandra stipulacea Benth. [GU] 

Calliandra surinamensis Benth. [GU SU FG] 

Calliandra tenuiflora Benth. [GU SU FG] 

Calliandra trinervia Benth. [FG] 

Cedrelinga cateniformis (Ducke) Ducke [SU FG] 

Chloroleucon acacioides (Ducke) Barneby & Grimes, comb. 

nov. ined. [SU FG] 

Desmanthus pernambucanus (L.) Thell. [GU SU] 

Desmanthus virgatus (L.) Willd. [GU SU] [to be expected in 

FG] 

Dichrostachys glomerata (Forsk.) Chiov. [FG] [naturalized] 

Entada polyphylla Benth. [GU SU FG] 

Entada polystachya (L.) DC. [GU SU FG] 

Enterolobium cyclocarpum (Jacq.) Griseb. [GU SU?] 

Enterolobium oldemanii Barneby & Grimes, sp. nov. ined. 

[FG] 

Enterolobium schomburgkii (Benth.) Benth. [GU SU FG] 

Hydrochorea corymbosa (A. Rich.) Barneby & Grimes, gen. & 

comb. ined. [GU SU FG] 

Hydrochorea gonggrijpii (Kleinh.) Barneby & Grimes, gen. & 

comb. ined. [GU SU FG?] 

Inga acreana Harms [SU FG] 

Inga acrocephala Steud. [GU SU FG] 

Inga alata Benoist [GU FG] 

Inga alba (Swartz) Willd. [GU SU FG] 

Inga auristellae Harms [GU SU FG] 

Inga bourgoni (Aubl.) DC. [GU SU FG] 

SPERMATOPHYTA: Mimosaceae 

141 

Inga brachystachys Ducke [SU FG] 

Inga bracteosa Benth. [GU SU FG] 

Inga brevipes Benth. [GU] 

Inga calanthoides Amsh. [SU] 

Inga capitata Desv. [GU SU FG] 

Inga cayennensis Sagot ex Benth. [GU SU FG] 

Inga cinnamomea Spruce ex Benth. [SU] 

Inga crassiflora Ducke [FG] 

Inga cylindrica Mart. [FG] 

Inga disticha Benth. [GU SU FG] 

Inga edulis (Vell.) Mart. [GU SU FG] 

Inga fanchoniana O. Poncy [GU? FG] 

Inga fastuosa (N.J. Jacq.) Willd. [GU FG] 

Inga flagelliformis Mart. [FG] 

Inga graciliflora Benth. [GU SU FG] 

Inga gracilifolia Ducke [GU FG] 

Inga heterophylla Willd. [GU SU FG] 

Inga huberi Ducke [SU FG] 

Inga ingoides (L.C. Rich.) Willd. [GU SU FG] 

Inga java Pittier [GU FG] 

Inga jenmanii Sandw. [GU FG] 

Inga lateriflora Miq. [GU SU FG] 

Inga laurina (Swartz) Willd. [GU] 

Inga leiocalycina Benth. [GU SU FG] 

Inga leptingoides Amsh. [SU] 

Inga lomatophylla (Benth.) Pitt. [GU FG] 

Inga longipedunculata Ducke [FG] 

Inga meissneriana Miq. [GU SU FG] 

Inga melinonis Sagot [GU FG] 

Inga nobilis Willd. [GU SU FG] 

Inga nubium Poncy [FG] 

Inga nuda Salzm. ex Benth. [GU SU FG] 

Inga paraensis Ducke [FG] 

Inga pedunculata Ducke [FG] 

Inga pezizifera Benth. [GU SU FG] 

Inga pilosula (L.C. Rich.) Macbride [GU SU FG] 

Inga poeppigiana Benth. [FG] 

Inga retinocarpa Poncy [FG] 

Inga rhynchocalyx Sandw. [GU FG] 

Inga rubiginosa (L.C. Rich.) DC. [GU SU FG] 

Inga sarmentosa Glaz. ex Harms [GU SU FG] 

Inga semialata (Vell.) Mart. [GU SU FG] 

Inga sertulifera DC. [GU SU FG] 

Inga splendens Willd. [GU SU FG] 

Inga stenocalyx Spruce ex Benth. [GU FG] 

Inga stenoptera Benth. [GU] 

Inga stipularis DC. [GU SU FG] 

Inga thibaudiana DC. [GU SU FG] 

Inga tubaeformis Benoist [FG] 

Inga umbellifera (Vahl) Steud. ex DC. [GU SU FG] 

Inga virgultosa (Vahl) Desv. [SU FG] 

Leucaena leucocephala (Lam.) de Wit [GU SU FG] 


Lysiloma bahamensis Benth. [FG] [introduced, dubiously 

persisting] 

Macrosamanea kegelii (Meisn.) Kleinh. [SU FG] 

Macrosamanea pubiramea (Steud.) Barneby & Grimes, comb. 

nov. ined. [GU SU FG] 

Mimosa annularis Benth. var. odora Barneby [GU SU FG] 

Mimosa annularis Benth. var. xinguensis (Ducke) Barneby 

[FG] 

Mimosa bimucronata (DC.) Kuntze [cultivated or escaped 


Mimosa camporum Benth. [GU SU FG?] 

Mimosa casta L. [FG] [possibly introduced]

142 


Mimosa debilis Humb. & Bonpl. ex Willd. [GU? SU FG] 

Mimosa diplotricha C. Wright ex Sauv. [GU SU FG] 

Mimosa dormiens Humb. & Bonpl. ex Willd. [GU] [record 

unconfirmed, but to be expected] 

Mimosa guilandinae (DC.) Barneby var. duckei (Huber) 

Barneby [SU FG] 

Mimosa guilandinae (DC.) Barneby var. guilandinae [FG] 

Mimosa invisa Mart. ex Colla var. spiciflora (Karst.) Barneby 

[GU SU?] 

Mimosa microcephala Humb. & Bonpl. ex Willd. var. 

cataractae (Ducke) Barneby [SU FG] 

Mimosa microcephala Humb. & Bonpl. ex Willd. var. lumaria 

Barneby [GU FG?] 

Mimosa myriadenia (Benth.) Benth. var. dispersa Barneby [SU 

FG] 

Mimosa myriadenia (Benth.) Benth. var. myriadenia [GU SU 

FG] 

Mimosa pigra L. [GU SU FG] 

Mimosa polydactyla Humb. & Bonpl. ex Willd. [GU SU FG] 

Mimosa pudica L. var. hispida Brenan [GU SU] 

Mimosa pudica L. var. pastoris Barneby [GU] 

Mimosa pudica L. var. tetrandra (Humb. & Bonpl. ex Willd.) 

DC. [GU SU FG] 

Mimosa pudica L. var. unijuga (Walp. & Duchaiss.) Griseb. 

[GU SU FG] 

Mimosa quadrivalvis L. var. leptocarpa (DC.) Barneby [GU 

FG] 

Mimosa rufescens Benth. [GU] [to be expected in SU, FG] 

Mimosa schomburgkii Benth. [GU FG] 

Mimosa schrankioides Benth. var. sagotiana (Benth.) Barneby 

[FG] 

Mimosa schrankioides Benth. var. schrankioides [GU] 

Mimosa somnians Humb. & Bonpl. ex Willd. var. somnians 

[GU FG] 

Mimosa somnians Humb. & Bonpl. ex Willd. var. viscida 

(Willd.) Barneby [FG] 

Mimosa surumuensis Harms [GU] 

Neptunia natans (L.f.) Druce [GU SU FG] [to be expected in 

FG] 

Neptunia plena (L.) Benth. [GU SU FG] 

Parkia decussata Ducke [FG] 

Parkia gigantocarpa Ducke [GU] 

Parkia igneiflora Ducke [GU] 

Parkia nitida Miq. [GU SU FG] 

Parkia pendula (Willd.) Benth. ex Walpers [GU SU FG] 

Parkia reticulata Ducke [FG] 

Parkia ulei (Harms) Kuhlmann var. surinamensis Kleinh. [GU 

SU FG] 

Parkia velutina Benoist [SU FG] [to be expected in GU] 

Pentaclethra macroloba (Willd.) Kuntze [GU SU FG] 

Piptadenia floribunda Kleinh. [SU FG] 

Piptadenia imatacae Barneby [to be expected in GU] 

Piptadenia leucoxylon Barneby & Grimes [GU] 

Piptadenia moniliformis Benth. [to be expected in the 


Piptadenia uaupensis Spruce ex Benth. [FG?] 

Pithecellobium dulce (Roxb.) Benth. [GU SU? FG] 


Pithecellobium roseum (Vahl) Barneby & Grimes, comb. nov. 

ined. [GU] 

Pithecellobium unguis-cati (L.) Benth. [GU FG] [cultivated, 

naturalized] 

Plathymenia reticulata Benth. [SU] 

Pseudopiptadenia psilostachya (DC.) Lewis & M.P. Lima [GU 

SU FG] 


Pseudopiptadenia suaveolens (Miq.) Grimes [GU SU FG] 

Samanea inopinata (Harms) Barneby & Grimes [GU] 

Samanea saman (Jacq.) Merrill [GU] [cultivated, perhaps 

escaping] 

Stryphnodendron guianense (Aubl.) Benth. [GU SU FG] 

Stryphnodendron moricolor Barneby & Grimes [GU FG] 

Stryphnodendron polystachyum (Miq.) Kleinh. [GU SU FG] 

Stryphnodendron pulcherrimum (Willd.) Hochr. [GU SU FG] 

Zapoteca portoricensis (Jacq.) H. Hern. subsp. flavida (Urb.) 

H. Hern. [GU] [specimens not seen, but to be expected] 

Zygia ampla (Benth.) Pittier [GU] 

Zygia cataractae (Kunth) L. Rico [GU SU FG] 

Zygia collina (Sandw.) Barneby & Grimes, comb. nov. ined. 

[GU] 

Zygia eperuetorum (Sandw.) Barneby & Grimes, comb. nov. 

ined. [GU] 

Zygia inaequalis (Humb. & Bonpl. ex Willd.) Pittier [GU SU 

FG] 

Zygia inundata (Ducke) H. de Lima [FG] 

Zygia juruana (Harms) L. Rico [GU] 

Zygia latifolia (L.) Fawcett & Rendle var. lasiopus (Benth.) 

Barneby & Grimes, stat. nov. ined. [GU SU FG] 

Zygia latifolia (L.) Fawcett & Rendle var. latifolia [GU SU? 

FG?] 

Zygia palustris Barneby & Grimes, sp. nov. ined. [GU] 

Zygia potaroensis Barneby & Grimes, sp. nov. ined. [GU] 

Zygia racemosa (Ducke) Barneby & Grimes, comb. nov. ined. 

[GU SU FG] 

Zygia sabatieri Barneby & Grimes, sp. nov. ined. [FG] 

Zygia tetragona Barneby & Grimes, sp. nov. ined. [FG] 

Zygia unifoliolata (Benth.) Pittier [to be expected in GU] 

MIMOSACEAE - SYNONYMS AND 


Abarema claviflora (Spruce) Kleinh. [not in Guianas] 

Abarema racemosa (Ducke) Kleinh. =Zygia racemosa 

Acacia chrysantha DC. [name of uncertain application, type 

not located] 

Acacia glandulosa (Vahl) DC. [nomen nudum] 

Acacia ouyrarema DC. [nomen nudum] 

Acacia paniculata Willd. =Acacia tenuifolia 

Acuan virgatum (L.) Medic. =Desmanthus virgatus 

Albizia caribea (Urban) Britton & Rose =Albizia niopoides 

Albizia corymbosa (A. Rich.) G.P. Lewis & P.E. Owen 

=Hydrochorea corymbosa 

Albizia inopinata (Harms) G.P. Lewis =Samanea inopinata 

Anneslia calothyrsus (Meissn.) Kleinh. =Calliandra 

calothyrsus 

Anneslia fasciculata (Willd.) Kleinh. =Calliandra surinamensis 

Anneslia tergemina (L.) Britton & Rose =Calliandra tergemina 


Arthrosamanea corymbosa (A. Rich.) Kleinh. =Hydrochorea 

corymbosa 

Arthrosamanea gonggrijpii (Kleinh.) Kleinh. =Hydrochorea 

gonggrijpii 

Arthrosamanea multiflora (Kunth) Kleinh. [sensu Kleinh. 

=Albizia subdimidiata] 

Arthrosamanea pullei (Kleinh.) Kleinh. =Hydrochorea 

gonggrijpii 

Calliandra anthoniae Grimes =Calliandra coriacea 

Calliandra pakaraimensis Cowan =Calliandra rigida 

Calliandra patrisii Sagot =Calliandra hymenaeodes? 

Calliandra tergemina (L.) Benth. [not in Guianas]


Entada gigas Fawc. & Rendle [not in Guianas] 

Enterolobium racemosum Ducke [apparently a nomen nudum] 

Inga acuminata Benth. [material identified as I. rhynchocalyx] 

Inga aggregata G. Don =Inga alba 

Inga angustifolia Willd. [not in Guianas] 

Inga commewijnensis Miq. =Inga nuda 

Inga conglomerata Benoist =Inga nobilis 

Inga coriacea (Pers.) Desv. =Inga sertulifera 

Inga crevauxii Sagot =Inga disticha 

Inga dysantha Benth. =Inga cayennensis 

Inga fagifolia (L.) Willd. ex Benth. =Inga laurina 

Inga longiflora Spruce ex Benth. [not in Guianas] 

Inga marginata Willd. =Inga semialata 

Inga myriantha Poepp. & Endl. =Inga umbellifera 

Inga nitida Willd. =Inga pilosula 

Inga panurensis Spruce ex Benth. [not in Guianas] 

Inga perrottetii Sagot =Inga nuda 

Inga pilosiuscula Desv. =Inga pilosula 

Inga prieurei Sagot =Inga nuda 

Inga ricardorum Bernardi & Spichiger [material identified as 

Inga java] 

Inga scabriuscula Benth. =Inga edulis 

Inga sciadion Steud. =Inga umbellifera 

Inga setifera DC. =Inga pilosula 

Inga speciosa Spruce ex Benth. =Inga lomatophylla 

Inga spuria Humb. & Bonpl. [not in Guianas] 

Inga strigillosa Spruce ex Benth. =Inga leiocalycina 

Inga urnifera Kleinh. =Inga pezizifera 

Inga velutina (Poir.) Willd. [not in Guianas] 

Klugiodendron laetum (Benth.) Britton & Killip =Abarema 

laeta 

Leucaena glauca (L.) Benth. [not in Guianas] 

Macrosamanea discolor (Humb. & Bonpl.) Britt. & Rose ex 

Britt. & Killip [not in Guianas] 

Macrosamanea pedicellaris (DC.) Kleinh. =Balizia pedicellaris 

Marmaroxylon collinum (Sandw.) L. Rico =Zygia collina 

Marmaroxylon eperuetorum (Sandw.) L. Rico =Zygia 

eperuetorum 

Marmaroxylon racemosum (Ducke) Killip ex Record =Zygia 

racemosa 

Mimosa acacioides Benth. =Anadenanthera peregrina (L.) 

Speg. [cultivated] 

Mimosa asperata L. [not in Guianas] 

Mimosa bourgoni Aubl. =Inga bourgoni 

Mimosa ceratonia L. [not in Guianas] 

Mimosa farnesiana L. =Acacia farnesiana 

Mimosa guianensis Aubl. =Stryphnodendron guianense 

Mimosa micracantha Benth. =Mimosa rufescens 

Mimosa obovata Benth. =Mimosa laticifera Rizzini & Mattos 

Filho [not in Guianas] 

Mimosa paniculata Benth. =Mimosa annularis var. odora 

Mimosa pellita Humb. & Bonpl. ex Willd. =Mimosa pigra 

Mimosa plumaeifolia Kleinh. =Mimosa microcephala var. 

cataractae 

Mimosa sagotiana Benth. =Mimosa schrankioides var. 

sagotiana 

Mimosa viva L. [not in Guianas] 

Mimosa xinguensis Ducke =Mimosa annularis var. xinguensis 

Neptunia oleracea Lour. =Neptunia natans 

Neptunia prostrata (Lam.) Baill. =Neptunia natans 

Newtonia psilostachya (DC.) Brenan =Pseudopiptadenia 

psilostachya 

Newtonia suaveolens (Miq.) Brenan =Pseudopiptadenia 

suaveolens 

Parkia oppositifolia Spruce ex Benth. =Parkia nitida 


143 

Pentaclethra filamentosa Benth. =Pentaclethra macroloba 

Piptadenia polystachya Miq. =Stryphnodendron polystachyum 

Piptadenia psilostachya (DC.) Benth. =Pseudopiptadenia 

psilostachya 

Piptadenia suaveolens Miq. =Pseudopiptadenia suaveolens 

Pithecellobium acacioides Ducke =Chloroleucon acacioides 

Pithecellobium adiantifolium (Kunth) Benth. =Macrosamanea 

discolor [not in Guianas] 

Pithecellobium arenarium Ducke =Abarema barbouriana var. 

arenaria 

Pithecellobium barbourianum Standl. =Abarema barbouriana 

var. barbouriana 

Pithecellobium basijugum Ducke =Zygia basijuga (Ducke) 

Barneby & Grimes [not in Guianas] 

Pithecellobium cauliflorum (Willd.) Mart. =Zygia latifolia 

Pithecellobium collinum Sandw. =Zygia collina 

Pithecellobium corymbosum (A. Rich.) Benth. =Hydrochorea 

corymbosa 

Pithecellobium curvicarpum Irwin =Abarema curvicarpa 

Pithecellobium elegans Ducke =Balizia elegans (Ducke) 


Pithecellobium eperuetorum Sandw. =Zygia eperuetorum 

Pithecellobium fanshawei Sandw. =Abarema barbouriana 

Pithecellobium forfex (Kunth) Benth. =Pithecellobium roseum 

Pithecellobium glomeratum (DC.) Benth. =Zygia cataractae 

Pithecellobium gonggrijpii Kleinh. =Hydrochorea gonggrijpii 

Pithecellobium huberi Ducke =Zygia latifolia 

Pithecellobium inaequale (Humb. & Bonpl. ex Willd.) Benth. 

=Zygia inaequalis 

Pithecellobium jupunba (Willd.) Urb. =Abarema jupunba 

Pithecellobium kegelii Meissn. =Macrosamanea kegelii 

Pithecellobium laetum Benth. =Abarema laeta 

Pithecellobium lanceolatum Benth. [not in Guianas] 

Pithecellobium lasiopus Benth. =Zygia latifolia var. lasiopus 

Pithecellobium latifolium (L.) Benth. =Zygia latifolia var. 

latifolia 

Pithecellobium mangense (Jacq.) Macbride =Chloroleucon 

mangense (Jacq.) Britton & Rose [not in Guianas] 

Pithecellobium mataybifolium Sandw. =Abarema mataybifolia 

Pithecellobium multiflorum (Kunth) Benth. =Arthrosamanea 

multiflora?[sensu Bentham, =Albizia subdimidiata] 

Pithecellobium niopoides (Benth.) Spruce ex Benth. =Albizia 

niopoides 

Pithecellobium pedicellare (DC.) Benth. =Balizia pedicellaris 

Pithecellobium polycephalum Benth. =Albizia polycephala 


Pithecellobium pullei Kleinh. =Hydrochorea gonggrijpii 

Pithecellobium racemosum Ducke =Zygia racemosa 

Pithecellobium trapezifolium (Vahl) Benth. =Abarema 

jupunba var. trapezifolia 

Pithecellobium villiferum Ducke =Abarema villifera (Ducke) 


Samanea polycephala (Benth.) Pittier =Albizia polycephala 

[not in Guianas]

144 


Schrankia leptocarpa DC. =Mimosa quadrivalvis var. 

leptocarpa 

Stryphnodendron angustum Benth. =Stryphnodendron 

pulcherrimum 

Stryphnodendron flammatum Kleinh. =Dimorphandra sp.? 

Stryphnodendron melinonis Sagot =Stryphnodendron 

pulcherrimum 

Stryphnodendron purpureum Ducke =Stryphnodendron 

guianense 

Zygia cauliflora (Mart.) Killip ex Record =Zygia latifolia 

Zygia glomerata (DC.) Pittier =Zygia cataractae 

Zygia trunciflorum (Ducke) L. Rico =Zygia juruana 

MOLLUGINACEAE 


REFERENCES: 

Burger, W. 1983. Aizoaceae. In W. Burger, 

Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 

199-212. 

Eyma, P.J. 1934. Aizoaceae. In A. Pulle, Flora 


Nevling, L Jr. 1961. Aizoaceae. In R.E. 



48(1): 80-85. 

Glinus radiatus (Ruiz & Pav.) Rohrb. [GU FG] 

Mollugo verticillata L. [GU SU FG] 

Trianthema portulacastrum L. [pantropical weed, to be 

expected] 

MONIMIACEAE 

reviewed by S. Renner (1996) 

REFERENCES: 

Jansen-Jacobs, M.J. 1976. Monimiaceae 



426. 

Lanjouw, J. 1939. Monimiaceae (additions and 


2(1): 471-472. 

Peixoto, A.L. 1987. Nota sobre as espécies de 

Mollinedia (Monimiaceae, Monimioideae) das 

Guianas. Bradea 4(41): 331-336. 

Perkins, J. & E. Gilg. 1901. Monimiaceae. In A. 

Engler, Das Pflanzenreich IV(101): 1-122. 

Petter, H.F.M. 1932. Monimiaceae. In A. Pulle, 


Steyermark, J.A. 1951. Monimiaceae. In 

Botanical Exploration in Venezuela - I. 

Fieldiana Bot. 28(1): 235-236. 

SPERMATOPHYTA: Mimosaceae - Moraceae 

Mollinedia grazielae A.L. Peixoto [GU SU] 

Mollinedia laurina Tul. [GU SU FG] 

Mollinedia ptariensis Steyerm. [GU] 

Mollinedia roraimensis Steyerm. [GU] 

Siparuna cristata (Poepp. & Endl.) A. DC. [FG] 

Siparuna cuspidata (Tul.) A. DC. [GU SU FG] 

Siparuna decipiens (Tul.) A. DC. [GU SU FG] 

Siparuna guianensis Aubl. [GU SU FG] 

Siparuna pachyantha A.C. Smith [FG] 

Siparuna sprucei A. DC. [GU SU FG] 

Siparuna surinamensis Lanj. [GU SU] 

MONIMIACEAE - SYNONYMS AND 


Citriosma amazonica (Mart.) A. DC. =Siparuna sprucei 

Citriosma cristata Poepp. & Endl. =Siparuna cristata 

Citriosma cuspidata Tul. =Siparuna cuspidata 

Citriosma decipiens Tul. =Siparuna decipiens 

Citriosma guianensis (Aubl.) Tul. =Siparuna guianensis 

Conuleum guianense A. Rich. =Siparuna decipiens 

Siparuna amazonica (Mart.) A. DC. =Siparuna sprucei 

Siparuna argyrochrysea Perk. =Siparuna cuspidata 

Siparuna emarginata Cowan =Siparuna pachyantha 

MORACEAE 



REFERENCES: 

Berg, C.C. 1972. Olmedieae and Brosimeae 

(Moraceae). Flora Neotropica Monogr. 7: 1- 

229. 

Berg, C.C. 1992. Moraceae. In A.R.A. Görtsvan 

Rijn, Flora of the Guianas (series A) 11: 

10-92. 

Berg, C.C. & R.W.A.P. Akkermans. 1985. 

Studies on the flora of the Guianas 14. New 

taxa and combinations in Sorocea and a key to 

its species. Proc. Kon. Ned. Akad. Wet. ser. 

C, 88(4): 381-394. 

Berg, C.C. & G.P. DeWolf. 1975. Moraceae. In 

J. Lanjouw & A.L. Stoffers, Flora of 

Suriname 5(1): 173-265. 

Berg, C.C., M. Vásquez Avila, & F. Kooy. 1984. 

Ficus species of Brazilian Amazonia and the 

Guianas. Acta Amaz. 14(1/2) suppl.: 159- 

194. 

Lindeman, J.C. 1984. Moraceae (additions and 



Bagassa guianensis Aubl. [GU SU FG] 

Batocarpus amazonicus (Ducke) Fosb. [FG] 

Brosimum acutifolium Huber subsp. acutifolium [SU FG] 

Brosimum acutifolium Huber subsp. obovatum (Ducke) C.C. 

Berg [GU FG] 

Brosimum alicastrum Swartz subsp. bolivarense (Pitt.) C.C. 

Berg [GU]


Brosimum guianense (Aubl.) Huber [GU SU FG] 

Brosimum lactescens (S. Moore) C.C. Berg [GU SU FG] 

Brosimum parinarioides Ducke subsp. parinarioides [GU SU 

FG] 

Brosimum rubescens Taub. [GU SU FG] 

Brosimum utile (Kunth) Pitt. subsp. ovatifolium (Ducke) C.C. 

Berg [GU? SU? FG] 

Clarisia ilicifolia (Spreng.) Lanj. & Rossberg [GU SU FG] 

Clarisia racemosa Ruiz & Pav. [SU] 

Dorstenia brasiliensis Lam. [FG] 

Dorstenia cayapia Vell. subsp. asaroides (Hook.) C.C. Berg 

[to be expected in SU] 

Dorstenia contrajerva L. [to be expected in GU] 

Ficus albert-smithii Standl. [GU SU FG] 

Ficus amazonica (Miq.) Miq. [GU SU FG] 

Ficus americana Aubl. [GU] 

Ficus blepharophylla Vazquez Avila [Mt. Roraima] 

Ficus broadwayi Urb. [GU SU FG] 

Ficus caballina Standl. [GU SU FG] 

Ficus catappifolia Kunth & Bouché [GU SU FG] 

Ficus cremersii C.C. Berg [FG] 

Ficus donnell-smithii Standl. [FG] 

Ficus eximia Schott [GU] 

Ficus gomelleira Kunth & Bouché [GU SU FG] 

Ficus greiffiana Dugand [GU FG] 

Ficus guianensis Desv. ex Hamilton [GU SU FG] 

Ficus hebetifolia Dugand [FG] 

Ficus insipida Willd. subsp. scabra C.C. Berg [GU SU FG] 

Ficus krukovii Standl. [SU FG] 

Ficus leiophylla C.C. Berg [SU FG] 

Ficus malacocarpa Standl. [GU SU FG] 

Ficus maroniensis Benoist [GU SU FG] 

Ficus mathewsii (Miq.) Miq. [GU SU] 

Ficus matiziana Dugand [FG] [to be expected in GU] 

Ficus maxima Mill. [GU SU FG] 

Ficus nymphaeifolia Mill. [GU SU FG] 

Ficus padifolia Kunth [GU] 

Ficus padifolia Kunth [GU?] 

Ficus pakkensis Standl. [GU SU FG] 

Ficus paludica Standl. [GU SU FG] 

Ficus panurensis Standl. [GU SU FG] 

Ficus paraensis (Miq.) Miq. [GU SU FG] 

Ficus pertusa L.f. [GU SU FG] 

Ficus piresiana Vazquez Avila & C.C. Berg [FG] 

Ficus pulchella Schott [FG?] 

Ficus roraimensis C.C. Berg [FG?] 

Ficus schumacheri (Liebm.) Griseb. [GU SU FG] 

Ficus subapiculata (Miq.) Miq. [GU? SU? FG] 

Ficus tepuiensis C.C. Berg & Simonis [to be expected in GU] 

Ficus trigona L.f. [GU SU FG] 

Ficus trigonata L. [GU?] 

Helicostylis pedunculata Benoist [SU FG] 

Helicostylis tomentosa (Poepp. & Endl.) Rusby [GU SU FG] 

Maclura tinctoria (L.) G. Don ex Steud. subsp. tinctoria [GU] 

Maquira calophylla (Poepp. & Endl.) C.C. Berg [FG] 

Maquira guianensis Aubl. [GU SU FG] 

Maquira sclerophylla (Ducke) C.C. Berg [GU SU FG] 

Naucleopsis glabra Spruce ex Pitt. [SU FG] 

Naucleopsis guianensis (Mildbr.) C.C. Berg [GU SU FG] 

Naucleopsis ulei (Warb.) Ducke subsp. ulei [to be expected in 

FG] 

Perebea guianensis Aubl. subsp. guianensis [SU FG] 

Perebea mollis (Poepp. & Endl.) Huber subsp. mollis [SU] 

Perebea mollis (Poepp. & Endl.) Huber subsp. rubra (Trécul) 

C.C. Berg [SU FG] 

SPERMATOPHYTA: Moraceae 

145 

Poulsenia sp. [FG] 

Pseudolmedia laevigata Trécul [GU FG] 

Pseudolmedia laevis (Ruiz & Pav.) Macbr. [GU SU FG] 

Sorocea hirtella Mildbr. subsp. oligotricha Akkermans & C.C. 

Berg [GU SU] 

Sorocea muriculata Miq. subsp. muriculata [GU? SU] 

Sorocea muriculata Miq. subsp. uaupensis (Baill.) C.C. Berg 

[SU FG] 

Sorocea sprucei (Baill.) Macbr. subsp. sprucei [GU] 

Trymatococcus oligandrus (Benoist) Lanj. [GU SU FG] 

Trymatococcus paraensis Ducke [GU SU FG] 

MORACEAE - SYNONYMS AND 


Acanthosphaera amara (Ducke) Ducke =Naucleopsis ulei 

Alicastrum guianense (Aubl.) Kuntze =Brosimum guianense 

Aliteria sagotii Benoist =Clarisia ilicifolia 

Anonocarpus amazonicus Ducke =Batocarpus amazonicus 

Bagassa sagotiana Bureau ex Benth. & J.D. Hook. =Bagassa 

guianensis 

Bagassa tiliifolia (W. Hamilton) Benoist =Bagassa guianensis 

Brosimopsis acutifolia (Huber) Ducke =Brosimum acutifolium 

Brosimopsis obovata Ducke =Brosimum acutifolium subsp. 

obovatum 

Brosimum angustifolium Ducke =Brosimum rubescens 

Brosimum aubletii Poepp. & Endl. =Brosimum guianense 

Brosimum bolivarense (Pitt.) Romero =Brosimum alicastrum 

subsp. bolivarense 

Brosimum discolor Schott =Brosimum guianense 

Brosimum lanciferum Ducke =Brosimum rubescens 

Brosimum lemeei (Benoist) Lemée =Brosimum guianense 

Brosimum ovatifolium Ducke =Brosimum utile subsp. 

ovatifolium 

Brosimum paraense Huber =Brosimum rubescens 

Brosimum rigidum Ducke =Brosimum utile subsp. ovatifolium 

Brosimum rotundatum Standl. =Brosimum guianense 

Brosimum velutinum (S.F. Blake) Ducke =Brosimum 

guianense 

Chlorophora tinctoria (L.) Gaudichaud =Maclura tinctoria 

Clarisia ilicifolia (Spreng.) Lanj. & Rossberg var. cuyunensis 

(Lanj.) Lanj. & Rossman =Clarisia ilicifolia 

Clarisia ilicifolia (Spreng.) Lanj. & Rossberg var. guianensis 

(Lanj.) Lanj. & Rossberg =Clarisia ilicifolia 

Clarisia strepitans (Allemao) Lanj. var. cuyunensis Lanj. 

=Clarisia ilicifolia 

Clarisia strepitans (Allemao) Lanj. var. guianensis Lanj. 

=Clarisia ilicifolia 

Ferolia guianensis Aubl. =Brosimum rubescens 

Ficus acarouaniensis Benoist =Ficus gomelleira 

Ficus anguina Benoist =Ficus nymphaeifolia 

Ficus angustifolia (Miq.) Miq. =Ficus amazonica 

Ficus arctocarpa Standl. =Ficus panurensis 

Ficus arukensis Standl. =Ficus paraensis 

Ficus citrifolia Mill. [sensu stricto, not in Guianas] 

Ficus clusiifolia Schott =Ficus guianensis 

Ficus ernestiana Pittier [sensu stricto, not in Guianas] 

Ficus erratica Standl. =Ficus guianensis 

Ficus erythrosticta (Miq.) Miq. =Ficus pertusa 

Ficus fagifolia (Miq.) Miq. =Ficus trigona 

Ficus fanshawei Standl. =Ficus trigona

146 


Ficus gardneriana (Miq.) Miq. =Ficus obtusifolia Kunth [not 


Ficus gigantea Kunth =Ficus citrifolia [not in Guianas] 

Ficus glaucescens (Liebm.) Miq. =Ficus maxima 

Ficus gleasonii Standl. =Ficus mathewsii 

Ficus hartii Warb. =Ficus trigonata 

Ficus kanukuensis Standl. =Ficus pertusa 

Ficus lanjouwii DeWolf =Ficus maroniensis 

Ficus leucosticta (Miq.) Miq. =Ficus paludica 

Ficus longifolia Schott [sensu stricto, not in Guianas] 

Ficus maguirei Standl. =Ficus panurensis 

Ficus manicariarum Standl. =Ficus paludica 

Ficus martinii Miq. =Ficus guianensis 

Ficus mensalis Standl. =Ficus guianensis 

Ficus obtusifolia Kunth [not in Guianas] 

Ficus pallida Vahl [not in Guianas] 

Ficus parkeri Miq. =Ficus maxima 

Ficus parkeriana (Miq.) Sandw. =Ficus guianensis 

Ficus perforata L. =Ficus americana 

Ficus prinoides Humb. & Bonpl. ex Willd. =Ficus pallida Vahl 


Ficus radula Humb. & Bonpl. ex Willd. =Ficus maxima 

Ficus ramiflora Standl. =Ficus caballina 

Ficus rolanderi (Liebm.) Miq. =Ficus pertusa 

Ficus savannarum Standl. =Ficus broadwayi 

Ficus surinamensis Miq. =Ficus amazonica 

Ficus thelephora Benoist =Ficus paraensis 

Ficus velutina Willd. [not in Guianas] 

Ficus vulpina Benoist =Ficus trigona 

Greeneina affinis (Miq.) Kuntze =Helicostylis tomentosa 

Helicostylis affinis Steud. ex Miq. =Helicostylis tomentosa 

Helicostylis bolivarensis Pitt. =Brosimum alicastrum subsp. 

bolivarense 

Helicostylis poeppigiana (Mart.) Trécul =Helicostylis 

tomentosa 

Lanessania oligandra Benoist =Trymatococcus oligandrus 

Laurea tiliifolia Gaudichaud =Bagassa guianensis 

Naucleopsis amara Ducke =Naucleopsis ulei subsp. ulei 

Noyera mollis (Poepp. & Endl.) Ducke =Perebea mollis 

Noyera rubra Trécul =Perebea mollis subsp. rubra 

Ogcodeia amara (Ducke) Ducke =Naucleopsis ulei subsp. ulei 

Ogcodeia glabra (Spruce ex Pitt.) Mildbr. =Naucleopsis glabra 

Ogcodeia guianensis Mildbr. =Naucleopsis guianensis 

Olmedia asperula Standl. =Helicostylis tomentosa 

Olmedia guianensis (Aubl.) Trécul =Maquira guianensis 

Olmedia maquira Steud. =Maquira guianensis 

Olmedia mollis Poepp. & Endl. =Perebea mollis 

Olmedioperebea sclerophylla Ducke =Maquira sclerophylla 

Perebea laurifolia Trécul =Maquira guianensis 

Perebea lecointei Huber =Perebea mollis subsp. mollis 

Perebea paraensis Huber =Perebea mollis subsp. mollis 

Perebea xinguana Standl. =Maquira sclerophylla 

Pharmacosycea guyanensis Miq. =Ficus maxima 

Pharmacosycea parkeriana Miq. =Ficus guianensis 

Piratinera acutifolia (Huber) Pitt. =Brosimum acutifolium 

Piratinera guianensis Aubl. =Brosimum guianense 

Piratinera lancifera (Ducke) Benoist =Brosimum rubescens 

Piratinera lemeei Benoist =Brosimum guianense 

Piratinera paraensis (Huber) Benoist =Brosimum rubescens 

Piratinera scabridula S.F. Blake =Brosimum guianense 

Piratinera velutina S.F. Blake =Brosimum guianense 

Pseudolmedia sagotii Benoist =Pseudolmedia laevis 

Sorocea dentata Huber =Sorocea muriculata subsp. muriculata 

Sorocea guayanensis W. Burger =Sorocea muriculata subsp. 

uaupensis 

SPERMATOPHYTA: Moraceae - Myristicaceae 

Trymatococcus amazonicus Poepp. & Endl. [not in Guianas] 

Urostigma angustifolium Miq. =Ficus amazonica 

Urostigma erythrostictum Miq. =Ficus pertusa 

Urostigma fagifolium Miq. =Ficus trigona 

Urostigma leucostictum Miq. =Ficus paludica 

Urostigma paraense Miq. =Ficus paraensis 

Urostigma rolanderi Liebm. =Ficus pertusa 

Urostigma schumacheri Liebm. =Ficus schumacheri 

MORINGACEAE 

[Old World family, one species widely cultivated] 

REFERENCES: 

Stoffers, A.L. 1984. Moringaceae. In A.L. 


5(1): 570-572. 

Moringa oleifera Lam. [cultivated] 

MUSACEAE 

[s.s., Old World family, cultivated in Guianas] 

Musa spp. [cultivated] 

MUSACEAE - SYNONYMS AND 


Musa humilis Aubl. =Heliconia psittacorum 

REFERENCES: 

MYOPORACEAE 

Lanjouw, J. 1938. Myoporaceae. In A. Pulle, 


Bontia daphnoides L. [GU SU FG] [cultivated; dubiously 

native or naturalized] 

MYRISTICACEAE 

contributed by W.A. Rodrigues (1992) 

REFERENCES:


Jansen-Jacobs, M.J. 1976. Myristicaceae 



429. 

Ooststroom, S.J. van. 1934. Myristicaceae. In 


Ooststroom, S.J. van. 1939. Myristicaceae 



Rodrigues, W.A. 1980. Revisao taxonômica das 

espécies de Virola Aublet do Brasil. Acta 

Amaz. 10(1) suppl. 1-127. 

Smith, A.C. & R.P. Wodehouse. 1937. The 

American species of Myristicaceae. Brittonia 

2(5): 393-510. 

Compsoneura ulei Warb. [GU FG] 

Iryanthera hostmannii (Benth.) Warb. [GU SU FG] 

Iryanthera juruensis Warb. [GU SU FG] 

Iryanthera lancifolia Ducke [GU SU] 

Iryanthera macrophylla (Benth.) Warb. [GU] 

Iryanthera paraensis Huber [GU SU FG] 

Iryanthera sagotiana (Benth.) Warb. [GU SU FG] 

Iryanthera tessmannii Markgr. [SU FG?] 

Iryanthera ulei Warb. [GU] 

Osteophloeum platyspermum (Spruce ex A. DC.) Warb. [GU 

FG] 

Virola calophylla Warb. [GU SU] 

Virola elongata (Benth.) Warb. [GU] 

Virola michelii Heckel [GU SU FG] 

Virola multicostata Ducke [FG] 

Virola sebifera Aubl. [GU SU FG] 

Virola surinamensis (Rolander) Warb. [GU SU FG] 

Virola venosa (Benth.) Warb. [SU] 

MYRISTICACEAE - SYNONYMS AND 


Myristica platysperma Spruce ex A. DC. =Osteophloeum 

platyspermum 

Virola melinonii (Benoist) A.C. Smith =Virola michelii 

Virola mycetis Pulle =Virola sebifera p.p., Virola surinamensis 

p.p. 

MYRSINACEAE 

reviewed by J.J. Pipoly (1992); reviewed by J.J. 

Pipoly (1996) 

REFERENCES: 

Agostini, G. 1980. Una nueva clasificacion del 

genero Cybianthus (Myrsinaceae). Acta Biol. 

Venez. 10: 129-185. 

Bottelier, H.P. 1936. Myrsinaceae. In A. Pulle, 


Pipoly, J.J. 1983. Contributions toward a 

monograph of Cybianthus, III: a revision of 

subgenus Laxiflorus. Brittonia 35(1): 61-80. 

Pipoly, J.J. 1992. The genus Cybianthus 

subgenus Conomorpha in Guayana. Ann. 


SPERMATOPHYTA: Myristicaceae - Myrsinaceae 

147 

Ardisia elliptica Thunb. [GU] 

Ardisia guianensis (Aubl.) Mez [GU SU FG] 

Ardisia semicrenata Mart. [FG?] 

Cybianthus amplus (Mez) Agostini [GU FG] 

Cybianthus apiculatus (Steyerm.) Agostini [GU] 

Cybianthus duidae (Gleason & Moldenke) Agostini [GU] 

Cybianthus fabiolae Pipoly [GU] 

Cybianthus fulvopulverulentus (Mez) Agostini subsp. 

fulvopulverulentus [GU SU] 

Cybianthus fulvopulverulentus (Mez) Agostini subsp. 

magnoliifolius (Mez) Pipoly [GU SU FG] 

Cybianthus fuscus Mart. [SU] 

Cybianthus gardneri (A. DC.) Agostini [FG] 

Cybianthus grandifolius (Mez) Agostini [GU FG] 

Cybianthus guianensis (A. DC.) Miq. subsp. guianensis [GU 

FG] 

Cybianthus guianensis (A. DC.) Miq. subsp. pseudoicacoreus 

(A. DC.) Agostini [GU SU FG] 

Cybianthus lepidotus (Gleason) Agostini [GU?] 

Cybianthus leprieurii Agostini [FG] 

Cybianthus lineatus (Benth.) Pipoly [GU] 

Cybianthus microbotrys A. DC. [SU FG] 

Cybianthus multidenticulatus Pipoly, sp. nov. ined. [GU] 

Cybianthus pakaraimae Pipoly, sp. nov. ined. [GU] 

Cybianthus potiaei Mez [GU FG] [=Cybianthus 

microbotrys?] 

Cybianthus prieurii A. DC. [GU FG] 

Cybianthus ptariensis (Steyerm.) Pipoly [GU] 

Cybianthus punctatus (Mez) Agostini [GU] 

Cybianthus quelchii (N.E. Brown) Agostini [GU] 

Cybianthus roraimae (Mez) Agost. [GU] 

Cybianthus spicatus (Kunth) Agostini [GU] 

Cybianthus surinamensis (Spreng.f.) Agostini [GU SU] 

Cybianthus venezuelanus Mez [GU SU FG] 

Myrsine coriacea (Sw.) Roem. & Schult. [GU] 

Myrsine guianensis (Aubl.) Kuntze [GU SU FG] 

Myrsine maguireana Pipoly [GU] 

Myrsine minima (Steyerm.) Pipoly [Mt. Roraima] 

Myrsine nitida (Mez) Pipoly [GU] 

Myrsine roraimensis (A.C. Smith) Pipoly [GU] 

Stylogyne depauperata Mez [FG] 

Stylogyne kappleri Mez [SU FG] 

Stylogyne lasseri (Lundell) Pipoly [GU] 

Stylogyne latifolia A.C. Smith [GU] 

Stylogyne longifolia (Mart. ex Miq.) Mez [GU SU FG] 

Stylogyne micans Mez [GU FG] 

Stylogyne micrantha (Kunth) Mez [FG] 

Stylogyne schomburgkiana (DC.) Mez [GU FG] 

Stylogyne spruceana Mez [SU] 

Stylogyne surinamensis (Miq.) Mez [GU SU FG] 

MYRSINACEAE - SYNONYMS AND 


Ardisia acuminata Willd. [not in Guianas] 

Ardisia quelchii N.E. Brown =Cybianthus quelchii 

Ardisia surinamensis Miq. =Stylogyne surinamensis 

Conomorpha apiculata Steyerm. =Cybianthus apiculatus

148 


Conomorpha fulvopulverulenta Mez =Cybianthus 

fulvopulverulentus 

Conomorpha magnoliifolia Mez =Cybianthus 

fulvopulverulentus subsp. magnoliifolius 

Conomorpha multipunctata (A. DC.) Miq. =Cybianthus 

guianensis subsp. pseudoicacoreus 

Conomorpha peruviana DC. var. guianensis (A. DC.) Mez 

=Cybianthus guianensis subsp. guianensis 

Conomorpha ptariensis Steyerm. =Cybianthus ptariensis 

Conomorpha rigida Mez =Cybianthus fulvopulverulentus 

Conomorpha sessilis A.C. Smith =Cybianthus punctatus 

Cybianthus brownii Gleason =Cybianthus venezuelanus 

Cybianthus comatus Mez =Cybianthus prieurii 

Cybianthus gracillimus (Warm.) Mez [not in Guianas] 

Cybianthus multipunctatus A. DC. =Cybianthus guianensis 

subsp. pseudoicacoreus 

Cybianthus nitidus Miq. [not in Guianas] 

Cybianthus resinosus Mez [not in Guianas] 

Cybianthus viridiflorus A.C. Smith =Cybianthus prieurii 

Grammadenia lineata Benth. =Cybianthus lineatus 

Icacorea acuminata (Willd.) Lundell =Ardisia acuminata 

Willd.[not in Guianas] 

Icacorea guianensis Aubl. =Ardisia guianensis 

Myrsine resinosa (A.C. Smith) Pipoly [not in Guianas] 

Myrsine schomburgkiana Miq. =Elaeoluma schomburgkiana 

(Sapotaceae) 

Rapanea guianensis Aubl. =Myrsine guianensis 

Rapanea nitida Mez =Myrsine nitida 

Rapanea roraimensis A.C. Smith =Myrsine roraimensis 

Stylogyne turbacensis Mez [not in Guianas] 

Weigeltia grandifolia Mez =Cybianthus grandifolius 

Weigeltia microbotrys (DC.) Mez =Cybianthus microbotrys 

Weigeltia parviflora Mez =Cybianthus leprieurii 

Weigeltia potiaei Mez =Cybianthus potiaei 

Weigeltia surinamensis (Spreng.f.) Mez =Cybianthus 

surinamensis 

MYRTACEAE 

contributed by B. Holst & P. Hiepko (1992) 

REFERENCES: 

Amshoff, G.J.H. 1951. Myrtaceae. In A. Pulle, 


Kawasaki, M.L. & B.K. Holst. 1994. New 

species and a new combination in Myrtaceae 

from northeastern South America. Brittonia 

46(2): 137-143. 

Landrum, L.R. 1986. Campomanesia, Pimenta, 

Blepharocalyx, Legrandia, Acca, Myrrhinium, 

and Luma. Flora Neotropica Monogr. 45: 1- 

178. 

McVaugh, R. 1958. Myrtaceae (Calyptranthes 

and Marlierea). In B. Maguire, J.J. Wurdack, 



10(1): 1-156. 

McVaugh, R. 1969. Myrtaceae. In B. Maguire 



18(2): 1-290. 

SPERMATOPHYTA: Myrsinaceae - Myrtaceae 

Sobral, M. 1993. Sinopse de Myrciaria. Napaea 

9: 13-41. 

Blepharocalyx eggersii (Kiaerskov) Landrum [GU] 

Calycolpus cochleatus McVaugh [GU] 

Calycolpus goetheanus (Mart. ex DC.) O. Berg [GU SU FG] 

Calycolpus revolutus (Schauer) O. Berg [GU SU FG] 

Calycolpus roraimensis Steyerm. [GU] 

Calycolpus surinamensis McVaugh [GU SU] 

Calycorectes batavorum McVaugh [SU] 

Calycorectes bergii Sandw. [GU SU FG] 

Calycorectes grandifolius O. Berg [SU FG] 

Calycorectes maximus McVaugh [SU] 

Calyptranthes amshoffae McVaugh [SU FG] 

Calyptranthes bracteata M.L. Kawasaki & B. Holst [FG] 

Calyptranthes fasciculata O. Berg [GU SU FG] 

Calyptranthes lepida McVaugh [FG] 

Calyptranthes lucida Mart. ex DC. [SU FG] 

Calyptranthes ovoidea McVaugh [GU] 

Calyptranthes pulchella DC. [GU SU] 

Calyptranthes pullei Burret ex Amsh. [GU SU] 

Calyptranthes smithii McVaugh [GU] 

Calyptranthes speciosa Sagot var. speciosa [SU FG] 

Calyptranthes spicata Amsh. [SU] 

Calyptranthes vexata McVaugh [GU] 

Campomanesia aromatica (Aubl.) Griseb. [GU SU FG] 

Campomanesia grandiflora (Aubl.) Sagot [GU SU FG] 

Eugenia albicans (O. Berg) Urb. [SU FG] 

Eugenia amshoffae McVaugh [GU] 

Eugenia anastomosans DC. [GU SU FG] 

Eugenia arawakorum Sandw. [GU] 

Eugenia armeniaca Sagot [GU FG] 

Eugenia biflora (L.) DC. [GU SU FG] 

Eugenia brownsbergii Amsh. [SU FG] 

Eugenia chrysophyllum Poir. [SU FG] 

Eugenia citrifolia Poir. [SU FG] 

Eugenia coffeifolia DC. [GU SU FG] 

Eugenia conjuncta Amsh. [GU] 

Eugenia cowanii McVaugh [GU SU FG] 

Eugenia cucullata Amsh. [GU SU FG] 

Eugenia cupulata Amsh. var. cupulata [SU FG] 

Eugenia denigrata McVaugh [FG] 

Eugenia egensis DC. [GU SU FG?] 

Eugenia essequiboensis Sandw. [GU] 

Eugenia eurycheila O. Berg [GU] 

Eugenia exaltata Rich. ex O. Berg [FG] 

Eugenia excelsa O. Berg [GU SU FG] 

Eugenia feijoi O. Berg [GU SU FG] 

Eugenia ferreireana O. Berg [GU SU FG] 

Eugenia flavescens DC. [GU SU FG] 

Eugenia florida DC. [GU SU FG] 

Eugenia foetida Pers. [FG] [unverified name] 

Eugenia gomesiana O. Berg [GU SU] 

Eugenia gongylocarpa M.L. Kawasaki & B. Holst [FG] 

Eugenia griseiflora McVaugh [SU] 

Eugenia incanescens Benth. [GU] 

Eugenia kaieteurensis Amsh. [GU] 

Eugenia lambertiana DC. var. hispidula McVaugh [SU] 

Eugenia lambertiana DC. var. lambertiana [GU SU FG] 

Eugenia latifolia Aubl. [GU SU FG] 

Eugenia ligustrina (Swartz) Willd. var. hebecarpa Amsh. [GU 

SU] 

Eugenia ligustrina (Swartz) Willd. var. ligustrina [GU]


Eugenia luciae Amsh. [GU SU FG] 

Eugenia macrocalyx (Rusby) McVaugh [GU SU FG] 

Eugenia marowynensis Miq. [SU FG] 

Eugenia mimus McVaugh [GU FG] 

Eugenia minuscula McVaugh [SU] 

Eugenia monticola (Swartz) DC. var. latifolia Krug & Urb. 

[GU] 

Eugenia monticola (Swartz) DC. var. racemosa Amsh. [GU? 

SU FG?] 

Eugenia muricata DC. [SU FG] 

Eugenia omissa McVaugh [GU SU FG] 

Eugenia patens Poir. [FG] 

Eugenia patrisii Vahl [GU SU FG] 

Eugenia polystachya Rich. [GU FG] 

Eugenia polystachyoides Amsh. [GU SU] 

Eugenia producta DC. [FG] 

Eugenia pseudopsidium Jacq. [GU SU FG] 

Eugenia punicifolia (Kunth) DC. [GU SU FG] 

Eugenia quadriovulata Amsh. [GU SU] 

Eugenia ramiflora Desv. var. montana Amsh. [SU] 

Eugenia ramiflora Desv. var. ramiflora [SU FG] 

Eugenia sigillata McVaugh [GU] 

Eugenia tafelbergica Amsh. [GU SU] 

Eugenia tapacumensis O. Berg [GU SU FG] 

Eugenia tetramera (McVaugh) M.L. Kawasaki & B. Holst 

[GU SU FG] 

Eugenia trinervia Vahl [GU? FG] 

Eugenia undulata Aubl. [SU FG] 

Eugenia uniflora L. [SU] [cultivated; naturalized?] 

Eugenia wullschlaegeliana Amsh. [SU FG] 

Marlierea biptera Amsh. [SU] 

Marlierea buxifolia Amsh. [GU] 

Marlierea cuprea Amsh. [GU] 

Marlierea ferruginea (Poir.) McVaugh [GU SU FG] 

Marlierea guildingiana (Griseb.) Krug & Urb. [GU] 

Marlierea karuaiensis (Steyerm.) McVaugh [GU] 

Marlierea ligustrina McVaugh [GU?] 

Marlierea lituatinervia (O. Berg) McVaugh [GU] 

Marlierea macrophylla Amsh. [GU] 

Marlierea montana (Aubl.) Amsh. [GU SU FG] 

Marlierea multiglomerata Amsh. [SU FG] 

Marlierea salticola Amsh. [GU] 

Marlierea schomburgkiana O. Berg [GU SU] 

Marlierea summa McVaugh [GU] 

Marlierea umbraticola (Kunth) O. Berg [FG] 

Marlierea vicina McVaugh [GU] 

Melaleuca leucadendron L. [SU FG?] [cultivated; 

naturalized?] 

Melaleuca quinquenervia (Cav.) S.T. Blake [FG] [cultivated? 

naturalized?] 

Myrcia albidotomentosa (Amsh.) McVaugh [GU SU] 

Myrcia amazonica DC. [GU SU FG] 

Myrcia bolivarensis (Steyerm.) McVaugh [GU] 

Myrcia bracteata (L.C. Rich.) DC. [GU SU FG] 

Myrcia calycampa Amsh. [GU SU FG?] 

Myrcia citrifolia (Aubl.) Urb. [GU SU FG] 

Myrcia clausa McVaugh [to be expected in GU] 

Myrcia coumete (Aubl.) DC. [SU FG] 

Myrcia cuprea (O. Berg) Kiaerskou [SU FG] 

Myrcia decorticans DC. [GU SU FG] 

Myrcia deflexa (Poir.) DC. [GU SU FG] 

Myrcia ehrenbergiana (O. Berg) McVaugh [GU] 

Myrcia exploratoris McVaugh [GU] 

Myrcia extranea McVaugh [GU] 

Myrcia fallax (L.C. Rich.) DC. [GU SU FG] 

SPERMATOPHYTA: Myrtaceae 

149 

Myrcia fenestrata DC. [FG] 

Myrcia fenzliana O. Berg [GU] 

Myrcia gigas McVaugh [FG] 

Myrcia gonini McVaugh [SU] 

Myrcia graciliflora Sagot [GU SU FG] 

Myrcia grandis McVaugh [GU] 

Myrcia guianensis (Aubl.) DC. var. guianensis [GU SU FG] 

Myrcia imperfecta McVaugh [GU] 

Myrcia inaequiloba (DC.) Legrand [GU SU FG] 

Myrcia magnoliifolia DC. [GU SU FG] 

Myrcia minutiflora Sagot [GU SU FG] 

Myrcia multiflora (Lam.) DC. [GU SU FG] 

Myrcia myriantha McVaugh [GU] 

Myrcia nigrescens DC. [GU] 

Myrcia nubicola McVaugh [GU] 

Myrcia paivae O. Berg [GU SU] 

Myrcia pistrinalis McVaugh [SU] 

Myrcia platyclada DC. [GU SU FG] 

Myrcia porphyrea McVaugh [GU] 

Myrcia pyrifolia (Hamilton) Nied. [GU SU FG] 

Myrcia quitarensis (Benth.) Sagot [GU] 

Myrcia rotundata (Amsh.) McVaugh var. rotundata [GU] 

Myrcia rufipila McVaugh [GU SU FG] 

Myrcia rupta M.L. Kawasaki & B. Holst [FG] 

Myrcia saxatilis (Amsh.) McVaugh [SU FG] 

Myrcia servata McVaugh [GU SU FG] 

Myrcia sororopanensis Steyerm. [GU] 

Myrcia speciosa (Amsh.) McVaugh [GU] 

Myrcia splendens (Swartz) DC. [GU SU FG] 

Myrcia subobliqua (Benth.) Nied. [GU SU] 

Myrcia subsessilis O. Berg [FG] 

Myrcia sylvatica (G. Mey.) DC. [GU SU FG] 

Myrcia tafelbergica Amsh. [GU SU] 

Myrcia tomentosa (Aubl.) DC. [GU SU FG] 

Myrcianthes prodigiosa McVaugh [GU SU] 

Myrciaria amazonica O. Berg [GU? SU FG] 

Myrciaria cordata O. Berg [GU] 

Myrciaria dubia (Kunth) McVaugh [GU] 

Myrciaria floribunda (West ex Willd.) O. Berg [GU SU FG] 

Myrciaria vismiifolia (Benth.) O. Berg [GU SU FG] 

Plinia costata Amsh. [GU SU] 

Plinia pinnata L. [GU? SU?] 

Pseudanamomis umbellulifera (Kunth) Kausel [GU] 

Psidium acutangulum Mart. ex DC. [GU SU] 

Psidium densicomum DC. [GU] 

Psidium grandifolium Mart. ex DC. [SU] 

Psidium guajava L. [GU SU FG] [cultivated, naturalized] 

Psidium guineense Swartz [GU SU FG] 

Psidium laruotteanum Camb. [GU] 

Psidium persoonii McVaugh [GU SU FG] 

Psidium salutare (Kunth) O. Berg [GU SU] 

Psidium sartorianum (O. Berg) Nied. [GU SU FG] 

Psidium striatulum DC. [GU SU] 

Psidium submetrale McVaugh [GU] 

Siphoneugena dussii (Krug & Urb.) Proença var. salicifolia 

(McVaugh) Proença [to be expected in GU] 

Siphoneugena dussii (Krug & Urb.) Proença var. dussii [GU 

SU] 

Syzygium cumini (L.) Skeels [GU SU] [cultivated; 

naturalized?] 

Syzygium jambos (L.) Alston [GU SU FG] [cultivated, 


Ugni myricoides (Kunth) O. Berg [GU]

150 


MYRTACEAE - SYNONYMS AND 


Aulomyrcia divaricata O. Berg =Myrcia rufipila 

Aulomyrcia grandiflora O. Berg [not in Guianas] 

Aulomyrcia hostmanniana O. Berg =Myrcia amazonica 

Aulomyrcia obtusa (Schauer) O. Berg =Myrcia guianensis 

Aulomyrcia schaueriana (Miq.) Amsh. =Myrcia graciliflora 

Aulomyrcia spp. =Myrcia spp. 

Aulomyrcia tetramera Amsh. =Myrcia decorticans? 

Aulomyrcia triflora O. Berg =Myrcia citrifolia 

Calycolpus chnoiophyllus Riley =Calycolpus revolutus 

Calycolpus glaber (Benth.) O. Berg =Calycolpus goetheanus 

Calycolpus pyrifer Riley =Calycolpus revolutus 

Calyptranthes apoda McVaugh, nom. illeg. =Calyptranthes 

vexata 

Campomanesia tetramera McVaugh =Eugenia tetramera 

Catinga aromatica Aubl. =Eugenia feijoi 

Catinga moschata Aubl. =Calycorectes bergii 

Eugenia adenocalyx DC. =Eugenia citrifolia 

Eugenia atropunctata Steud. =Eugenia florida 

Eugenia benthamii O. Berg =Eugenia punicifolia 

Eugenia brachybotrya DC. =Mouriri guianensis 

(Melastomataceae) 

Eugenia comete Aubl. =Myrcia coumete 

Eugenia compta O. Berg =Eugenia pseudopsidium 

Eugenia coriacea O. Berg =Eugenia polystachyoides 

Eugenia cryptadena Amsh. var. gracilis Amsh. =Eugenia 

cowanii 

Eugenia cumini L. =Syzygium cumini 

Eugenia fasciculiflora O. Berg =Eugenia feijoi? 

Eugenia guianensis Aubl. =Myrcia guianensis 

Eugenia jambos L. =Syzygium jambos 

Eugenia mini Aubl. =Eugenia biflora 

Eugenia montana Aubl. =Marlierea montana 

Eugenia prosoneura O. Berg =Eugenia gomesiana 

Eugenia protracta Steud. =Myrciaria floribunda 

Eugenia psidioides DC. =Eugenia pseudopsidium 

Eugenia punicifolia (Kunth) DC. var. brachypoda (DC.) Kr. & 

Urb. =Eugenia ramiflora 

Eugenia pyrifolia Hamilton =Myrcia pyrifolia 

Eugenia racemifera Sagot =Eugenia florida 

Eugenia roraimana O. Berg =Eugenia tapacumensis 

Eugenia schomburgkii Benth. =Eugenia lambertiana 

Eugenia sinemariensis Aubl. =Eugenia sp.[name of uncertain 

application] 

Eugenia tomentosa Aubl. =Myrcia tomentosa 

Eugenia wentii Amsh. =Eugenia macrocalyx 

Krugia ferruginea (Poir.) Urb. =Marlierea ferruginea 

Marlierea dussii Krug & Urb. ex Urb. =Siphoneugena dussii 

Marlierea gleasonii McVaugh =Myrcia decorticans 

Myrcia berberis DC. =Myrcia fallax 

Myrcia divaricata Berg =Myrcia rufipila 

Myrcia grandiflora (O. Berg) Lemée =Aulomyrcia grandiflora 


Myrcia hostmanniana Kiaerk =Myrcia amazonica 

Myrcia kegeliana O. Berg =Myrcia fallax 

Myrcia obtusa Schauer =Myrcia guianensis 

Myrcia paniculata (Jacq.) Krug & Urb. =Myrcia citrifolia 

Myrcia schomburgkiana O. Berg =Myrcia servata 

Myrcia tetramera Amsh. =Myrcia decorticans? 

Myrciaria nitida (Benth.) O. Berg =Myrcia inaequiloba 

Myrciaria verticillata O. Berg =Myrciaria floribunda 

SPERMATOPHYTA: Myrtaceae - Nyctaginaceae 

Myrtus umbellata Desv. ex Hamilton =Mouriri guianensis 

(Melastomataceae) 

Plinia dussii (Krug & Urb.) Urb. =Siphoneugena dussii 

Psidium aquaticum Benth. =Psidium striatulum 

Psidium araca Raddi =Psidium guineense 

Psidium aromaticum Aubl. =Campomanesia aromatica 

Psidium benthamianum O. Berg =Psidium guineense 

Psidium grandiflorum Aubl. =Campomanesia grandiflora 

Psidium guianense sensu Amshoff, not Persoon =Psidium 

persoonii 

Psidium guyanense Persoon =Psidium guineense 

Psidium ovatifolium O. Berg var. glabrum Amsh. =Psidium 

densicomum 

Psidium parviflorum Benth. =Psidium striatulum 

Psidium quinquedentatum Amsh. =Psidium sartorianum? 

Siphoneugena densiflora O. Berg [not in Guianas] 

NAJADACEAE 

reviewed by R. Haynes & L.B. Holm-Nielsen 

(1992) 

REFERENCES: 

Lowden, R.M. 1986. Taxonomy of the genus 

Najas in the Neotropics. Aquatic Bot. 24: 

147-184. 

Najas guadalupensis (Spreng.) Magnus [FG?] 

Najas microcarpa K. Schum. [FG] [to be expected in GU, SU] 

NAJADACEAE - SYNONYMS AND 


Najas conferta (A. Brown) A. Brown [not in Guianas] 

Najas marina L. [widespread, perhaps to be expected] 

Najas microdon A. Brown =Najas guadalupensis 

Najas wrightiana A. Brown [as circumscribed by Lowden, not 


REFERENCES: 

NELUMBONACEAE 

Cramer, J. 1979. Nelumbonaceae. In A.L. 


5(1): 390-392. 

Nelumbo nucifera Gaertn. [GU SU FG?] [cultivated, 

naturalized] 

NYCTAGINACEAE 

revised by R.A. DeFilipps (1996) 

REFERENCES:


DeFilipps, R.A. & S.L. Maina (in press). 

Nyctaginaceae. In A.R.A. Görts-van Rijn, 

Flora of the Guianas (series A). 

Steyermark, J.A. 1987. Nyctaginaceae. In Flora 

of the Venezuelan Guayana - III. Ann. 


Standley, P.C. 1931. The Nyctaginaceae of 

northwestern South America. Field Mus. Bot. 

11: 73-114. 

Boerhavia diffusa L. [GU SU FG] 

Bouganvillea spp. [several species cultivated] 

Guapira cuspidata (Heimerl) Lundell [GU SU] 

Guapira eggersiana (Heimerl) Lundell [GU SU FG] 

Guapira kanukuensis (Standl.) Lundell [GU] 

Guapira salicifolia (Heimerl) Lundell [GU SU FG] 

Mirabilis jalapa L. [GU SU FG] [cultivated, escaping and 

naturalized] 

Neea constricta Spruce ex Schmidt [GU FG] 

Neea floribunda Poepp. & Endl. [GU SU FG] 

Neea mollis Spruce ex Schmidt [GU FG] 

Neea ovalifolia Spruce ex Schmidt [GU SU FG] 

Neea spruceana Heimerl [GU FG] 

Pisonia macranthocarpa (Donn. Sm.) Donn. Sm. [GU] 

NYCTAGINACEAE - SYNONYMS AND 


Boerhavia caribaea Jacq. =Boerhavia diffusa 

Boerhavia coccinea Mill. =Boerhavia diffusa 

Boerhavia paniculata L. Rich. =Boerhavia diffusa 

Boerhavia surinamensis Miq. =Boerhavia diffusa 

Guapira fragrans (Dum.-Cours.) Little [not in Guianas] 

Guapira glabra (Heimerl) Steyerm. =Guapira eggersiana 

Guapira graciliflora sensu Mennega et al. (1988), non (Mart.) 

Lundell =Guapira salicifolia 

Guapira guianensis Aubl. =Guapira eggersiana? 

Guapira heimerliana (Standl.) Lundell =Guapira salicifolia 

Guapira olfersiana (Link, Klotzsch & Otto) Lundell [not in 


Guapira pacurero (Kunth) Little [not in Guianas] 

Guapira riedeliana (Fisch. ex Heimerl) Lundell [not in 


Guapira schomburgkiana (Heimerl) Lundell =Guapira 

cuspidata 

Neea cauliflora Heimerl =Neea floribunda 

Neea nigricans (Sw.) Heimerl [not in Guianas] 

Neea tristis Heimerl [not in Guianas] 

Pisonia aculeata L. var. macranthocarpa Donn. Sm. =Pisonia 

macranthocarpa 

Pisonia albiflora Heimerl =Guapira salicifolia 

Pisonia cuspidata Heimerl =Guapira cuspidata 

Pisonia eggersiana Heimerl =Guapira eggersiana 

Pisonia glabra Heimerl =Guapira eggersiana 

Pisonia guianensis (Aubl.) Howard =Guapira eggersiana? 

Pisonia inermis Jacq. =Guapira fragrans [not in Guianas] 

Pisonia nitida Mart. ex Schmidt =Guapira eggersiana 

Pisonia olfersiana sensu Maguire (1948), non Link, Klotzsch 

& Otto =Guapira eggersiana 

Pisonia pacurero sensu Pulle (1906), non Kunth =Guapira 

eggersiana 

Pisonia salicifolia Heimerl =Guapira salicifolia 

Pisonia schomburgkiana Heimerl =Guapira cuspidata 

Torrubia cuspidata (Heimerl) Standl. =Guapira cuspidata 

Torrubia heimerliana Standl. =Guapira salicifolia 

Torrubia kanukuensis Standl. =Guapira kanukuensis 

Torrubia laurina Standl. =Guapira eggersiana 

Torrubia olfersiana (Link, Klotzsch, & Otto) Standl. [not in 


Torrubia salicifolia (Heimerl) Standl. =Guapira salicifolia 

Torrubia schomburgkiana (Heimerl) Standl. =Guapira 

cuspidata 

NYMPHAEACEAE 

additions and corrections by J.H. Wiersema 

(1996) 

REFERENCES: 

SPERMATOPHYTA: Nyctaginaceae - Ochnaceae 

151 

Conard, H.S. 1905. The Waterlilies: A 

Monograph of the Genus Nymphaea. 

Carnegie Institution of Washington. 

Cramer, J. 1979. Nymphaeaceae. In A.L. 


5(1): 370-384. 

Wiersema, J.H. 1987. A monograph of 

Nymphaea subgenus Hydrocallis. Syst. Bot. 

Monogr. 16: 1-112. 

Nymphaea amazonum Mart. & Zucc. subsp. amazonum [GU 

SU FG] 

Nymphaea ampla (Salisb.) DC. [GU SU] 

Nymphaea conardii Wiersema [GU] 

Nymphaea gardneriana Planch. [GU] 

Nymphaea glandulifera Rodschied [GU SU FG] 

Nymphaea jamesoniana Planch. [GU] 

Nymphaea lotus L. [GU] [cultivated, naturalized] 

Nymphaea odorata Solander [GU SU] 

Nymphaea rubra Roxb. ex Salisb. [GU SU] [cultivated, 

naturalized] 

Nymphaea rudgeana G. Mey. [GU SU FG] 

Nymphaea tenerinervia Caspary [to be expected in GU] 

Victoria amazonica (Poepp.) Sowerby [GU] 

NYMPHAEACEAE - SYNONYMS AND 


Castalia spp. =Nymphaea spp. 

Nymphaea blanda G. Mey. =Nymphaea glandulifera 

Nymphaea odorata Solander var. gigantea Tricker 

=Nymphaea odorata 

Nymphaea parkeriana Lehm. =Nymphaea odorata 

Victoria regia Lindl. =Victoria amazonica 

OCHNACEAE 

contributed by C. Sastre (1992) 

REFERENCES:

152 

SPERMATOPHYTA: Ochnaceae 


Allorge, L. & C. Sastre. 1990. Distribution 

géographique dans les Guyanes, 

d'Apocynaceae et d'Ochnaceae. Essai de mise 

en evidence de centres de speciation. C.R. 

Soc. Biogeogr. 67(2): 77-94. 

Dwyer, J.D. 1943. The taxa of the monogeneric 

tribe Elvasieae (Ochnaceae). Bull. Torrey 

Bot. Club 70: 42-49. 

Dwyer, J.D. 1945. The taxonomy of the genus 

Sauvagesia (Ochnaceae). Bull. Torrey Bot. 

Club 72(6): 521-540. 

Maguire, B., & J.J. Wurdack. 1961. Ochnaceae. 

In Botany of the Guayana Highland part IV. 


Maguire, B. & J.A. Steyermark. 1989. Ouratea 

in Guayana and adjacent Amazonian hylaea. 


Guayana Highland part XIII. Mem. New 

York Bot. Gard. 51: 56-102. 

Sastre, C. 1971. Distribution géographique des 

espèces de Sauvagesia L. C.R. Soc. 

Biogéogr. 421: 47-59. 

Sastre, C. 1984. Etude sur le genre Sauvagesia 

L. en Guyane française. Saussurea 15: 111- 

118. 

Sastre, C. 1986. Studies on the Flora of the 

Guianas 23. Cinq espèces nouvelles 

d'Ouratea Aublet (Ochnaceae) des Guyanes. 

Bull. Mus. Nat. Hist. ser. 4, 8, sect. B, 

Adansonia 4: 413-422. 

Sastre, C. 1987. Studies on the Flora of the 

Guianas 30. Considérations 

phytogéographiques sur les Ochnacées 

guyanaises. C.R. Soc. Biog. 63(3): 89-97. 

Sastre, C. 1988. Synopsis generis Ouratea Aubl. 

Bull. Mus. natn. Hist. nat. Paris, sér. 4, 10, 

sect. B, Adansonia 1: 47-67. 

Sastre, C. 1991. Duae novae species Ourateae 

(Ochnaceae) in Guianis collectae. Studies on 

the Flora of the Guianas. 59. Candollea 

46(2): 395-398. 

Sastre, C. 1994. Three new taxa of Ouratea 

(Ochnaceae) from South America. Brittonia 

46(4): 309-313. 

Sastre, C. 1995. Novelties in the Neotropical 

genus Ouratea Aublet (Ochnaceae). Novon 

5(2): 193-200. 

Wehlburg, C. 1937. Ochnaceae. In A. Pulle, 


Adenanthe bicarpellata Maguire, Steyerm. & Wurdack [GU] 

Cespedesia spathulata (Ruiz & Pav.) Planch. [GU] 

Elvasia brevipedicellata Ule [GU] 

Elvasia elvasioides (Planch.) Gilg [GU SU FG] 

Elvasia essequibensis Engl. [GU SU] 

Elvasia macrostipularis Sastre & Lescure [FG] 

Elvasia quinqueloba Spruce ex Engl. [GU] 

Elvasia sphaerocarpa Cowan [GU] 

Ouratea acuminata (DC.) Engl. [GU?] 

Ouratea angulata Tiegh. [GU? SU FG] 

Ouratea attenuata Tiegh. [GU] 

Ouratea bipartita Sastre [GU] 

Ouratea candollei (Planch.) Tiegh. [SU FG] 

Ouratea cardiosperma (Lam.) Engl. [GU? SU FG] 

Ouratea castaneifolia (DC.) Engl. [GU SU] 

Ouratea cataractarum Sandw. [GU] 

Ouratea cerebroidea Sastre [GU FG] 

Ouratea cernuiflora Sandw. [GU] 

Ouratea crenata (Spruce) Tiegh. [GU] 

Ouratea decagyna Maguire [GU? SU FG] 

Ouratea engleri Tiegh. [FG] 

Ouratea erecta Sastre [SU FG] 

Ouratea fasciculata Maguire & Steyerm. [GU] 

Ouratea ferruginea Engl. [GU FG] 

Ouratea francinae Sastre [FG] 

Ouratea frontium Sleumer [GU] 

Ouratea gigantophylla (Ehrhard) Engl. [SU] 

Ouratea gillyana (Dwyer) Sandw. & Maguire [GU] 

Ouratea guianensis Aubl. [GU SU FG] 

Ouratea guildingii (Planch.) Urb. [GU] 

Ouratea impressa (Tiegh.) Lemée [FG] 

Ouratea leblondii (Tiegh.) Lemée [GU SU FG] 

Ouratea leprieurii Tiegh. [FG] 

Ouratea longifolia (Lam.) Engl. [GU FG] 

Ouratea maasorum Sastre [GU] 

Ouratea macrocarpa Sastre [GU SU FG] 

Ouratea mazaruniensis A.C. Smith & Dwyer [GU] 

Ouratea melinonii (Tiegh.) Lemée [SU FG] 

Ouratea occultinervis Sastre [SU FG] 

Ouratea olivaeformis (St. Hil.) Engl. [GU] [unverified name] 

Ouratea pendula (Poepp.) Engl. [SU FG] 

Ouratea pseudotatei Maguire & Steyerm. [GU] 

Ouratea rigida Engl. [GU] 

Ouratea riparia Sleumer [SU FG] 

Ouratea roraimae Engl. [GU] 

Ouratea rupununiensis Engl. [GU] 

Ouratea sagotii (Tiegh.) Lemée [GU SU FG] 

Ouratea saülensis Sastre [FG] 

Ouratea schomburgkii (Planch.) Engl. [GU SU] 

Ouratea scottii Sastre subsp. scottii [GU SU FG] 

Ouratea sculpta (Tiegh.) Sastre [GU SU FG] 

Ouratea soderstromii Sastre [GU] 

Ouratea spruceana Engl. [GU SU?] 

Ouratea superba Engl. [GU SU] 

Ouratea tatei Gleason [GU] 

Ouratea timehriensis Sastre [GU] 

Ouratea vaccinioides (St. Hil. & Tul.) Engl. [GU?] 

Ouratea verruculosa Engl. [FG] 

Poecilandra pumila Steyerm. [GU] 

Poecilandra retusa Tul. [GU] 

Sauvagesia aliciae Sastre subsp. aratayensis Sastre [FG] 

Sauvagesia amoena Ule [GU?] 

Sauvagesia angustifolia Ule [GU SU] 

Sauvagesia deficiens A.C. Smith [GU] [unverified name] 

Sauvagesia elata Benth. [GU SU? FG] 

Sauvagesia erecta L. subsp. erecta [GU SU FG] 

Sauvagesia guianensis (Eichl.) Sastre [GU] 

Sauvagesia imthurniana (Oliver) Dwyer [GU] 

Sauvagesia longifolia Eichl. [GU] 

Sauvagesia longipes Steyerm. [GU] 

Sauvagesia pulchella Planch. [GU SU] 

Sauvagesia ramosissima Spruce ex Eichl. [SU FG] 

Sauvagesia roraimensis Ule [GU SU] 

Sauvagesia rubiginosa St. Hil. [GU SU FG] 

Sauvagesia sprengelii St. Hil. [GU SU FG] 

Sauvagesia tafelbergensis Sastre [SU FG] 

Sauvagesia tenella Lam. [GU SU FG]


OCHNACEAE - SYNONYMS AND 


Cespedesia amazonica Huber =Cespedesia spathulata 

Elvasia calophyllea DC. [not in Guianas] 

Elvasia hostmannia Planch. =Elvasia elvasioides 

Elvasia sagoti (Tiegh.) Lemée =Elvasia elvasioides 

Elvasia schomburgkii Tiegh. =Elvasia calophyllea DC.[not in 


Gomphia candollei Planch. =Ouratea candollei 

Gomphia surinamensis Planch. =Ouratea cardiosperma 

Kaieteurea gillyana Dwyer =Ouratea gillyana 

Leitgebia imthurniana Oliver =Sauvagesia imthurniana 

Ouratea discophora Ducke [not in Guianas] 

Ouratea integrifolia (DC.) Sagot [nomen nudum] 

Ouratea polygyna Engl. [not in Guianas] 

Ouratea racemiformis Ule =Ouratea schomburgkii 

Ouratea surinamensis (Planch.) Wehlburg =Ouratea 

cardiosperma 

Roraimanthus imthurnianus (Oliver) Gleason =Sauvagesia 

imthurniana 

Sauvagesia adima Aubl. =Sauvagesia erecta 

Sauvagesia erecta Aubl., non L. =Sauvagesia sprengelii 

Sauvagesia kappleri Miq. =Sauvagesia sprengelii 

Sauvagesia salicifolia Ging. =Sauvagesia erecta 

Sauvagesia smithiana Dwyer =Sauvagesia roraimensis 

Sauvagesia surinamensis Miq. =Sauvagesia rubiginosa 

OLACACEAE 

contributed by P. Hiepko (1992) 

REFERENCES: 

Amshoff, G.J.H. 1938. Olacaceae. In A. Pulle, 


Hiepko, P. 1993. Olacaceae. In A.R.A. Görtsvan 


3-35. 

Sleumer, H.O. 1984. Olacaceae. Flora 


Aptandra liriosmoides Spruce ex Miers [GU] 

Cathedra acuminata (Benth.) Miers [GU FG] 

Chaunochiton kappleri (Sagot ex Engl.) Ducke [GU SU FG] 

Dulacia crassa (Monachino) Sleumer [GU] 

Dulacia guianensis (Engl.) Kuntze [GU SU FG] 

Dulacia redmondii Steyerm. [GU] 

Heisteria barbata Cuatrec. [FG] 

Heisteria cauliflora J.E. Smith [GU SU FG] 

Heisteria densifrons Engl. [GU SU FG] 

Heisteria maguirei Sleumer [GU] 

Heisteria ovata Benth. [GU SU FG] 

Heisteria scandens Ducke [GU SU FG] 

Maburea trinervis P. Maas [GU] 

Minquartia guianensis Aubl. [GU SU FG] 

Ptychopetalum olacoides Benth. [GU SU FG] 

Schoepfia lucida Pulle [SU] 

Ximenia americana L. var. americana [GU SU FG] 

OLACACEAE - SYNONYMS AND 


SPERMATOPHYTA: Ochnaceae - Onagraceae 

153 

Cathedra guianensis Sandw. =Cathedra acuminata 

Chaunochiton tubicinum (Poepp.) Lemée =Aptandra tubicina 


Dulacia inopiflora (Miers) Kuntze [not in Guianas] 

Heisteria coccinea Jacq. [not in Guianas] 

Heisteria kappleri Sagot ex Engler =Chaunochiton kappleri 

Heisteria microcalyx Sagot =Heisteria densifrons 

Heisteria surinamensis Amsh. =Heisteria ovata 

Heymassoli inermis Aubl. =Ximenia americana 

Heymassoli spinosa Aubl. =Ximenia americana 

Liriosma crassa Monachino =Dulacia crassa 

Liriosma guianensis Engl. =Dulacia guianensis 

Liriosma inopiflora Miers =Dulacia inopiflora [not in 


Olax schomburgkii Klotzsch [nomen nudum] =Dulacia 

guianensis 

Secretania loranthacea Muell. Arg. =Minquartia guianensis 

REFERENCES: 

OLEACEAE 

Green, P.S. 1994. A revision of Chionanthus in 

S. America and the description of 

Priogymnanthus, gen. nov. Kew Bull. 49(2): 

261-286. 

Stearn, W.T. 1976. Union of Chionanthus and 

Linociera. Ann. Missouri Bot. Gard. 63: 355- 

357. 

Chionanthus guianensis (Aubl.) Pers. [FG] 

Jasminum spp. [cultivated, perhaps escaping] 

OLEACEAE - SYNONYMS AND 


Chionanthus guianensis (Aubl.) Stearn, comb. superfl. 

Chionanthus mayepea Vahl =Chionanthus guianensis 

Chionanthus tetrandra Vahl =Chionanthus guianensis 

Linociera tetrandra (Vahl) Roem. & Schult. =Chionanthus 

guianensis 

Mayepea guianensis Aubl. =Chionanthus guianensis 

ONAGRACEAE 

reviewed by E.M. Zardini & P.H. Raven (1992); 

reviewed by E.M. Zardini (1996) 

REFERENCES: 

Jonker, F.P. 1942. Oenotheraceae. In A. Pulle, 


154 


Zardini, E.M. & P.H. Raven. 1991. Onagraceae. 


Guianas (series A) 10: 1-44. 

Ludwigia affinis (DC.) Hara [GU SU FG] 

Ludwigia decurrens Walter [GU SU FG] 

Ludwigia dodecandra (DC.) Zardini & Raven [GU FG] 

Ludwigia elegans (Camb.) Hara [SU] 

Ludwigia erecta (L.) Hara [GU SU FG] 

Ludwigia foliobracteolata (Munz) Hara [GU FG] 

Ludwigia helminthorriza (Mart.) Hara [GU SU] 

Ludwigia hyssopifolia (G. Don) Exell [GU SU FG] 

Ludwigia inclinata (L.f.) Gómez [GU SU] 

Ludwigia latifolia (Benth.) Hara [GU SU FG] 

Ludwigia leptocarpa (Nutt.) Hara [GU SU FG] 

Ludwigia mexiae (Munz) Hara [SU] 

Ludwigia nervosa (Poir.) Hara [GU SU FG] 

Ludwigia octovalvis (Jacq.) Raven [GU SU FG] 

Ludwigia rigida (Miq.) Sandw. [GU SU] 

Ludwigia sedoides (Humb. & Bonpl.) Hara [GU] 

Ludwigia torulosa (Arnott) Hara [GU SU FG] 

ONAGRACEAE - SYNONYMS AND 


Jussiaea affinis DC. =Ludwigia affinis 

Jussiaea affinis DC. var. dodecandra (DC.) Munz =Ludwigia 

dodecandra 

Jussiaea affinis DC. var. multiflora Jonker =Ludwigia affinis 

Jussiaea aluligera Miq. =Ludwigia leptocarpa 

Jussiaea decurrens (Walt.) DC. =Ludwigia decurrens 

Jussiaea dodecandra DC. =Ludwigia dodecandra 

Jussiaea elegans Camb. =Ludwigia elegans 

Jussiaea erecta L. =Ludwigia erecta 

Jussiaea erecta L. var. sebana DC. =Ludwigia erecta 

Jussiaea foliobracteolata Munz =Ludwigia foliobracteolata 

Jussiaea helminthorriza Mart. =Ludwigia helminthorrhiza 

Jussiaea hexamera Miq. =Ludwigia affinis 

Jussiaea inclinata L.f. =Ludwigia inclinata 

Jussiaea latifolia Benth. =Ludwigia latifolia 

Jussiaea leptocarpa Nutt. =Ludwigia leptocarpa 

Jussiaea leptocarpa Nutt. var. aluligera (Miq.) Jonker 

=Ludwigia leptocarpa 

Jussiaea leptocarpa Nutt. var. meyeriana (Kuntze) Munz 

=Ludwigia leptocarpa 

Jussiaea linifolia Vahl =Ludwigia hyssopifolia 

Jussiaea mexiae Munz =Ludwigia mexiae 

Jussiaea micrantha Kunze =Ludwigia hyssopifolia 

Jussiaea nervosa Poir. =Ludwigia nervosa 

Jussiaea octonervia Lam. =Ludwigia octovalvis 

Jussiaea palustris G. Mey. =Ludwigia decurrens 

Jussiaea pterophora Miq. =Ludwigia decurrens 

Jussiaea rigida Miq. =Ludwigia rigida 

Jussiaea suffruticosa L. =Ludwigia octovalvis 

Jussiaea surinamensis Miq. =Ludwigia leptocarpa 

Jussiaea torulosa Arnott =Ludwigia torulosa 

Jussiaea variabilis G. Mey. var. affinis (DC.) O. Kuntze 

=Ludwigia affinis 

Jussiaea variabilis G. Mey. var. meyeriana Kuntze =Ludwigia 

leptocarpa 

Ludwigia affinis (DC.) Hara var. dodecandra (DC.) Munz 

=Ludwigia dodecandra 

Ludwigia micrantha (Kunze) Hara =Ludwigia hyssopifolia 

SPERMATOPHYTA: Onagraceae - Orchidaceae 

Oocarpon torulosum (Arnott) Urb. =Ludwigia torulosa 

OPILIACEAE 

contributed by P. Hiepko (1992) 

REFERENCES: 

Hiepko, P. 1993. Opiliaceae. In A.R.A. Görtsvan 


36-39. 

Agonandra brasiliensis Miers ex Benth. [GU] 

Agonandra silvatica Ducke [GU SU FG] 

ORCHIDACEAE 

contributed by E.A. Christenson & J.K. Boggan; 

Acineta, Braemia, Catasetum, Coryanthes, 

Gongora, Houlletia, Kegeliella, Mormodes, 

Paphinia, Peristeria, Polycycnis, Sievekingia, 

& Stanhopea by R. Jenny; Dichaea by J. 

Folsom; Encyclia & Epidendrum by E. 

Hagsater; Habenaria by J. Renz (1992); 

additions and corrections by E.A. Christenson 

& J.K. Boggan, reviewed by M.C.M. 

Werkhoven & M.E. Medley (1996) 

REFERENCES: 

Cremers, G., M. Hoff, D. Barthelemy, E.A. 

Christenson, Ly. & L. Leneveu, & Ch. & Cl. 

Pawilowski. 1992. Inventaire taxonomique 

des plantes de la Guyane française II - les 

Orchidacées. Mus. Nat. Hist. Nat., Coll. Patr. 

Nat. (Sér. Patr. Génétique) 7: 1-144. 

Dunsterville, G.C.K. & L.A. Garay. 1959-1976. 

Venezuelan Orchids Illustrated. 6 vol. 

Foldats, E. 1970. Orchidaceae. In T. Lasser, 

Flora de Venezuela. 

Renz, J. 1992. The genus Habenaria in the 

Guianas. Candollea 47: 483-512. 

Schweinfurth, C. 1967. Orchidaceae of the 

Guayana Highland. Mem. New York Bot. 

Gard. 14(3): 69-214. 

Szlachetko, D.L. & Y. Veyret. 1994. Two new 

species of Spiranthinae (Orchidaceae) from 

French Guyana. Fragm. Flor. Geobot. 39(2): 

471-474. 

Werkhoven, M.C.M. 1986. Orchids of 

Suriname. Uitgeversmaatschappij: 

Paramaribo, Suriname. 

Acacallis fimbriata (Rchb.f.) Schlechter [GU SU] 

Acineta alticola C. Schweinf. [GU?] 

Acineta superba (Kunth) Rchb.f. [SU] 

Ada keiliana (Rchb.f. ex Lindl.) N.H. Wms. [GU] 

Aganisia pulchella Lindl. [GU SU FG]


Aspasia variegata Lindl. [GU SU FG] 

Aspidogyne foliosa (Poepp. & Endl.) Garay [GU SU FG] 

Aspidogyne longicornu (Cogn.) Garay [GU SU] 

Aspidogyne multifoliata (Schweinf.) Garay [GU] 

Aspidogyne pumila (Cogn.) Garay [GU SU] 

Baskervilla venezuelana Garay & Dunsterv. [GU] 

Batemannia armillata Rchb.f. [SU] 

Batemannia colleyi Lindl. [GU SU FG] 

Batemannia lepida Rchb.f. [SU] 

Beloglottis costaricensis (Rchb.f.) Schlechter [SU] 

Bifrenaria longicornis Lindl. [GU SU FG] 

Bletia altilamellata Garay [GU] 

Bletia purpurea (Lam.) DC. [SU? FG] 

Bollea hemixantha Rchb.f. [GU FG?] 

Bollea violacea (Lindl.) Rchb.f. [GU SU FG] 

Brachionidium brevicaudatum Rolfe [GU] 

Brachystele guyanensis (Lindl.) Schlechter [GU SU FG] 

Braemia vittata (Lindl.) Jenny [GU SU? FG] 

Brassavola cucullata R. Brown [GU SU FG] 

Brassavola gardneri Cogn. [SU? FG] 

Brassavola martiana Lindl. [GU SU FG] 

Brassavola nodosa (L.) Lindl. [GU FG] 

Brassia angusta Lindl. [GU] 

Brassia bidens Lindl. [GU] 

Brassia caudata (L.) Lindl. [GU SU FG] 

Brassia chloroleuca Barb. Rodr. [GU FG] 

Brassia huebneri Schlechter [FG] 

Brassia lanceana Lindl. [GU SU FG] 

Brassia lawrenceana Lindl. [GU SU FG] 

Brassia macrostachya Lindl. [GU SU] 

Brassia neglecta Rchb.f. [GU] 

Brassia pumila Lindl. [GU FG] 

Brassia villosa Lindl. [GU] 

Brassia wageneri Rchb.f. [GU] 

Bulbophyllum bracteolatum Lindl. [GU SU FG] 

Bulbophyllum exaltatum Lindl. [GU] 

Bulbophyllum geraense Rchb.f. [GU] 

Bulbophyllum oerstedii (Rchb.f.) Hemsl. [SU FG] 

Bulbophyllum pachyrrachis (A. Rich.) Griseb. [FG] 

Bulbophyllum quadrisetum Lindl. [SU FG] 

Bulbophyllum roraimense Rolfe [GU FG] 

Bulbophyllum setigerum Lindl. [GU] 

Caluera surinamensis Dodson & R.O. Determann [SU] 

Caluera vulpina Dodson & R.O. Determann [FG] 

Campylocentrum fasciola (Lindl.) Cogn. [GU SU FG] 

Campylocentrum micranthum (Lindl.) Rolfe [GU SU FG] 

Campylocentrum pachyrrhizum (Rchb.f.) Rolfe [SU? FG] 

Campylocentrum poeppigii (Rchb.f.) Rolfe [GU] 

Campylocentrum steyermarkii Foldats [GU] 

Catasetum barbatum (Lindl.) Lindl. var. barbatum [GU SU 

FG] 

Catasetum barbatum (Lindl.) Lindl. var. spinosum Rolfe [FG?] 

Catasetum bicallosum Cogn. [GU?] 

Catasetum bicolor Klotzsch [GU?] 

Catasetum callosum Lindl. [GU] [var. callosum?] 

Catasetum callosum Lindl. var. carunculatum Mansf. [GU SU? 

FG] 

Catasetum comosum Cogn. [GU?] 

Catasetum cristatum Lindl. [GU SU] 

Catasetum deltoideum (Lindl.) Mutel [GU SU FG?] 

Catasetum discolor (Lindl.) Lindl. [GU SU FG] 

Catasetum ferox Kraenzl. [GU?] 

Catasetum gnomus Linden & Rchb.f. [GU?] 

Catasetum longifolium Lindl. [GU SU FG] 

Catasetum luridum (Link) Lindl. [GU] 

SPERMATOPHYTA: Orchidaceae 

155 

Catasetum macrocarpum L.C. Rich. ex Kunth [GU SU FG] 

Catasetum ochraceum Lindl. [GU?] 

Catasetum pileatum Rchb.f. [GU?] 

Catasetum planiceps Lindl. [GU SU FG?] 

Catasetum poriferum Schomb. ex Lindl. [GU] 

Catasetum saccatum Lindl. [GU] 

Catasetum thompsonii Dodson [GU] 

Catasetum xdunstervillei Romero & Carnevali [GU?] 

Catasetum xguianense Jenny & Romero [GU SU? FG] 

Catasetum xroseo-album (Hook.) Lindl. [GU SU FG?] 

Catasetum xtapiriceps Rchb.f. [GU? FG?] 

Catasetum xwendlingeri Foldats [GU] 

Cattleya jenmannii Rolfe [GU] 

Cattleya lawrenceana Rchb.f. [GU] 

Cattleya violacea (Kunth) Rolfe [GU FG] 

Caularthron bicornutum (Hook.) Raf. [GU SU] 

Chamelophyton kegelii (Rchb.f.) Garay [SU] 

Chaubardia surinamensis Rchb.f. [GU SU FG] 

Chaubardiella tigrina (Garay & Dunsterv.) Garay [GU SU FG] 

Cheiradenia cuspidata Lindl. [GU SU FG] 

Cleistes grandiflora (Aubl.) Schlechter [GU FG] 

Cleistes moritzii (Rchb.f.) Garay & Dunsterv. [GU] 

Cleistes paludosa Rchb.f. [SU FG] 

Cleistes parviflora Lindl. [GU SU?] 

Cleistes rosea Lindl. [GU SU FG] 

Cleistes tenuis (Rchb.f. ex Griseb.) Schlechter [GU SU FG] 

Cleistes unifoliata (Schweinf.) Carnevali & Ramirez [SU] 

Clowesia warscewiczii (Lindl. & Paxt.) Dodson [GU? SU?] 

Cochleanthes guianensis Laf., Gerl. & Sengh. [GU SU FG] 

Comparettia falcata Poepp. & Endl. [FG] 

Coryanthes macrantha (Hook.) Hook. [GU SU FG] 

Coryanthes maculata Hook. [GU FG] 

Coryanthes punctata Beer [GU] [=Coryanthes speciosa var. 

puncata (Beer) Gerlach] 

Coryanthes speciosa (Hook.) Hook. [GU SU FG] 

Corymborkis sp. [GU?] 

Cranichis diphylla Swartz [SU] 

Cranichis sp. [SU FG] 

Cryptarrhena guatemalensis Schlechter [GU SU FG] 

Cryptarrhena kegelii Rchb.f. [GU SU FG] 

Cryptocentrum sp. nov. [GU] [Cryptocentrum peruvianum 

aucct., Cryptocentrum gracillimum auctt.] 

Cyclopogon bifidus (Ridl.) Schlechter [GU] 

Cyclopogon elatus (Swartz) Schlechter [SU] 

Cyclopogon olivaceus (Rolfe) Schlechter [GU SU FG] 

Cycnoches chlorochilon Klotzsch [GU? SU? FG] 

Cycnoches haagii Barb. Rodr. [SU FG] 

Cycnoches loddigesii Lindl. [GU SU FG] 

Cyrtopodium andersonii (Lamb.) R. Brown [GU SU FG] 

Cyrtopodium cristatum Lindl. [GU SU FG] 

Cyrtopodium intermedium Brade [SU] 

Cyrtopodium paniculatum (Ruiz & Pav.) Garay [GU SU FG] 

Cyrtopodium parviflorum Lindl. [GU SU FG] 

Degranvillea dermaptera R.O. Determann [FG] 

Dichaea angustifolia H.G. Jones [GU] 

Dichaea brevicaulis Cogn. [FG?] 

Dichaea bryophila Rchb.f. [FG?] 

Dichaea camaridioides Schlechter [GU?] [Mt. Roraima] 

Dichaea cornuta S. Moore [SU] 

Dichaea histrio Rchb.f. [GU? SU?] 

Dichaea hookeri Garay & Sweet [GU SU FG] 

Dichaea kegelii Rchb.f. [GU SU FG?] 

Dichaea morrisii Fawc. & Rendle [GU?] 

Dichaea ochracea Lindl. [GU] 

Dichaea pendula (Aubl.) Cogn. [GU FG]

156 


Dichaea picta Rchb.f. [GU SU FG] 

Dichaea rendlei Gleason [GU SU FG] 

Dichaea sp. cf. trulla Rchb.f. [GU SU FG] 

Dichaea splitgerberi Rchb.f. [GU SU FG] 

Dichaea trulla Rchb.f. [GU SU FG?] 

Dichaea weigeltii Rchb.f. [GU SU] 

Dimerandra elegans (Focke) Siegerist [GU SU FG] 

Dimerandra emarginata (Mey.) Hoehne [GU SU FG] 

Duckeela alticola C. Schweinf. [to be expected in GU] 

Elleanthus brasiliensis (Lindl.) Rchb.f. [FG] 

Elleanthus caravata (Aubl.) Rchb.f. [GU SU FG] 

Elleanthus cephalotus Garay & Sweet [GU SU FG] 

Elleanthus crinipes Rchb.f. [FG] 

Elleanthus furfuraceus (Lindl.) Rchb.f. [GU SU?] 

Elleanthus graminifolius (Barb. Rodr.) Loijtnant [GU SU FG] 

Elleanthus trilobatus Ames & Schweinf. [GU] 

Eloyella panamensis (Dressler) Dodson [GU] 

Eltroplectris calcarata (Swartz) Garay & Sweet [SU] 

Encyclia aemula (Lindl.) Carnevali & Ramirez [GU SU FG] 

Encyclia calamaria (Lindl.) Pabst [GU SU FG] 

Encyclia chloroleuca (W.J. Hook.) Neum. [GU SU] 

Encyclia cochleata (L.) Lemée [GU SU? FG] 

Encyclia conchichila (Barb. Rodr.) Porto & Brade [GU] 

Encyclia cordigera (Kunth) Dressler [GU FG] 

Encyclia dichroma (Lindl.) Schlechter [SU] 

Encyclia diurna (Jacq.) Schlechter [GU SU FG] 

Encyclia granitica (Lindl.) Schlechter [GU SU FG] 

Encyclia guianensis Carnevali & G. Romero [GU] 

Encyclia ionosma (Lindl.) Schlechter [GU SU? FG] 

Encyclia latipetala (C. Schweinf.) Pabst [GU SU?] 

Encyclia longifolia Barb. Rodr. [SU?] 

Encyclia odoratissima (Lindl.) Schlechter [GU FG] 

Encyclia pachyantha (Lindl.) Hoehne [GU FG] 

Encyclia picta (Lindl.) Hoehne [GU] 

Encyclia pygmaea (W.J. Hook.) Dressler [GU SU FG] 

Encyclia tarumana Schlechter [FG] 

Encyclia tigrina (Lind. ex Lindl.) Carnevali & Ramírez [GU] 

Encyclia vespa (Vell.) Dressler [GU SU FG] 

Epidendrum anceps Jacq. [GU SU FG] 

Epidendrum apaganum Mansf. [SU] 

Epidendrum bakrense Hagsater & Cremers, sp. nov. ined. [FG] 

Epidendrum calanthum Rchb.f. [GU] 

Epidendrum carpophorum Barb. Rodr. [GU SU FG] 

Epidendrum churunense Garay & Dunsterv. [GU] 

Epidendrum ciliare L. [GU SU FG] 

Epidendrum compressum Griseb. [GU SU FG] 

Epidendrum cooperianum Batem. [GU] 

Epidendrum coronatum Ruiz & Pav. [SU] 

Epidendrum cristatum Ruiz & Pav. [GU SU] 

Epidendrum dendrobioides Thunb. [GU] 

Epidendrum densiflorum Hook. [GU SU FG] 

Epidendrum difforme Jacq. [GU SU FG] 

Epidendrum diffusum Swartz var. depauperatum Cogn. [SU] 

[=Epidendrum compressum?] 

Epidendrum dipus Lindl. [FG] 

Epidendrum ecostatum Pabst [GU FG] 

Epidendrum elongatum Jacq. [GU] 

Epidendrum flexuosum Mey. [GU SU FG] 

Epidendrum geminiflorum Kunth [SU] 

Epidendrum gracillimum Lindl. [SU?] 

Epidendrum gramineum Lindl. [SU?] 

Epidendrum gratissimum (Rchb.f.) Hagsater [GU] 

Epidendrum ibaguense Kunth [GU SU? FG] 

Epidendrum imitans Schlechter [GU FG] 

Epidendrum imthurnii Ridl. [GU FG] 


Epidendrum isomerum Schlechter [GU] 

Epidendrum latilabre Lindl. [GU SU? FG] 

Epidendrum longicolle Lindl. [GU] 

Epidendrum macrocarpum (L.C. Rich.) Garay, comb. nov. 

ined. [GU SU FG] 

Epidendrum microphyllum Lindl. [GU SU FG] 

Epidendrum miserrimum Rchb.f. [GU SU] 

Epidendrum nocturnum Jacq. [GU SU FG] 

Epidendrum nutans Swartz [SU] 

Epidendrum oldemanii Christenson [FG] 

Epidendrum orchidiflorum Salzm. ex Lindl. [GU] 

Epidendrum paniculatum Ruiz & Pav. [GU SU FG] 

[=Epidendrum densiflorum?] 

Epidendrum patens Swartz [GU?] 

Epidendrum prostratum (Lindl.) Cogn. [GU] 

Epidendrum purpurascens Focke [GU SU FG] 

Epidendrum ramosum Jacq. [GU SU FG] 

Epidendrum revertianum (Stehle) Hagsater [GU] 

Epidendrum rigidum Jacq. [GU SU FG] 

Epidendrum rostratum Garay & Dunsterv. [FG] 

Epidendrum saxatile Lindl. [GU] 

Epidendrum schlechterianum Ames [GU SU] 

Epidendrum sculptum Rchb.f. var. linearifolium Rchb.f. [SU] 

Epidendrum sculptum Rchb.f. var. sculptum [GU SU FG] 

Epidendrum smaragdinum Lindl. [GU SU FG] 

Epidendrum stiliferum Dressler [SU?] 

Epidendrum strobiliferum Rchb.f. [GU SU FG] 

Epidendrum strobiloides Garay & Dunsterv. [GU SU FG] 

Epidendrum tenuifolium Lindl. [GU] 

Epidendrum tridens Poepp. & Endl. [GU SU] 

Epidendrum unguiculatum (Schweinf.) Garay & Dunsterv. 

[SU FG] 

Epidendrum violascens Ridl. [GU] 

Epidendrum viviparum Lindl. [GU FG?] 

Epistephium amplexicaule Poepp. & Endl. [GU] 

Epistephium duckei Huber [GU] 

Epistephium ellipticum L.O. Williams & Summerh. [GU] 

Epistephium lucidum Cogn. [GU] 

Epistephium parviflorum Lindl. [GU SU FG] 

Epistephium sclerophyllum Lindl. [SU?] 

Epistephium subrepens Hoehne [GU] 

Eriopsis biloba Lindl. [GU SU FG] 

Eriopsis sceptrum Rchb.f. & Warsc. [SU?] 

Erythrodes hirtella (Swartz) Fawc. & Rendle [GU SU? FG] 

Erythrodes major (Presl) Ames [GU SU] 

Erythrodes pumila (Cogn.) Pabst [SU] 

Eulophia alta (L.) Fawc. & Rendle [GU SU FG] 

Galeandra baueri Lindl. var. baueri [FG] 

Galeandra baueri Lindl. var. lutea Wats. [FG] 

Galeandra beyrichii Rchb.f. [to be expected in the Guianas] 

Galeandra devoniana Schomb. ex Lindl. [GU] 

Galeandra dives Rchb.f. [GU SU FG] 

Galeandra lagoensis Rchb.f. & Warm. [SU] 

Galeandra stylomisantha (Vell.) Hoehne [GU SU FG] 

Galeottia burkei (Rchb.f.) Dressler & Christenson [GU SU?] 

Galeottia jorisiana (Rolfe) Dressler & Christenson [GU] 

Gomphichis adnata (Ridl.) Schlechter [GU] [=Gomphichis 

viscosa?] 

Gomphichis viscosa (Rchb.f.) Schlechter [GU] 

Gongora atropurpurea W.J. Hook. [GU SU FG] 

Gongora histrionica Rchb.f. [GU? SU FG?] 

Gongora maculata Lindl. [GU FG?] 

Gongora nigrita Lindl. [GU SU?] 

Gongora pleiochroma Rchb.f. [GU FG] 

Govenia utriculata (Swartz) Lindl. [SU?]


Habenaria abortiens Lindl. [FG?] 

Habenaria alata Hook. [SU?] 

Habenaria alterosula Snuverink & Westra [SU FG] 

Habenaria amalfitana Lehm. & Kraenzl. [GU] [Mt. Roraima] 

Habenaria amambayensis Schlechter [GU FG] 

Habenaria ayangannensis Renz [GU] 

Habenaria caldensis Kraenzl. [GU SU FG?] 

Habenaria dusenii Schlechter [SU? FG] 

Habenaria entomantha (Llave & Lex.) Lindl. [GU SU FG?] 

Habenaria ernestii Schlechter [GU] 

Habenaria heptadactyla Rchb.f. [GU SU] 

Habenaria lehmanniana Kraenzl. [GU FG] 

Habenaria leprieurii Rchb.f. [GU SU FG] 

Habenaria longicauda Hook. subsp. ecalcarata Snuverink & 

Westra [GU SU FG] 

Habenaria longicauda Hook. subsp. longicauda [SU] 

Habenaria macilenta (Lindl.) Rchb.f. [GU SU FG] 

Habenaria monorrhiza (Swartz) Rchb.f. [to be expected in the 


Habenaria montis-wilhelminae Renz [SU] 

Habenaria obtusa Lindl. [GU SU FG?] 

Habenaria odontopetala Rchb.f. [GU] 

Habenaria parvidens Lindl. [SU?] 

Habenaria parviflora Lindl. [GU SU? FG?] 

Habenaria platydactyla Kraenzl. [GU SU FG] 

Habenaria pratensis (Lindl.) Rchb.f. var. parviflora Cogn. 

[GU? SU FG] 

Habenaria quinqueseta (Michx.) A. Eaton [GU SU? FG?] 

Habenaria repens Nuttall [GU SU FG] 

Habenaria rodeiensis Barb. Rodr. [GU FG] 

Habenaria rodriquezii Cogn. [FG] 

Habenaria roraimensis Rolfe [GU] [Mt. Roraima] 

Habenaria santensis Barb. Rodr. [SU?] 

Habenaria schomburgkii Lindl. ex Benth. [GU SU] 

Habenaria setacea Lindl. [GU FG] 

Habenaria seticauda Lindl. ex Benth. [GU FG] 

Habenaria sprucei Cogn. [GU SU FG] 

Habenaria trifida Kunth [GU SU FG] 

Hexisea bidentata Lindl. [GU SU? FG] 

Hexisea imbricata (Lindl.) Rchb.f. [GU SU] 

Houlletia odoratissima Linden ex Lindl. & Paxt. [GU SU?] 

Houlletia roraimensis Rolfe [GU] 

Huntleya lucida (Rolfe) Rolfe [GU] 

Huntleya meleagris Lindl. [GU] 

Huntleya sessiliflora Batem. ex Lindl. [GU] 

Ionopsis satyrioides (Swartz) Rchb.f. [GU SU FG] 

Ionopsis utricularioides (Swartz) Lindl. [GU SU FG] 

Isochilus linearis (Jacq.) R. Brown [GU SU FG] 

Jacquiniella globosa (Jacq.) Schlechter [GU SU FG] 

Jacquiniella leucomelana (Rchb.f.) Schlechter [GU] 

Jacquiniella steyermarkii Carnevali & Dressler [GU] 

Jacquiniella teretifolia (Swartz) Britt. & Wils. [GU SU FG] 

Kefersteinia lafontainei Senghas & Gerlach [SU FG] 

Kegeliella houtteana (Rchb.f.) L.O. Williams [SU FG] 

Koellensteinia carraoënsis Garay & Dunsterv. [SU FG] 

Koellensteinia eburnea (Barb. Rodr.) Schlechter [GU SU] 

Koellensteinia graminea (Lindl.) Rchb.f. [GU SU FG] 

Koellensteinia hyacinthoides Schlechter [SU? FG?] 

Koellensteinia kellneriana Rchb.f. [GU SU FG] 

Koellensteinia tricolor (Lindl.) Rchb.f. [GU] 

Lepanthes helicocephala Rchb.f. [GU SU FG] 

Lepanthes ruscifolia Rchb.f. [SU FG] 

Lepanthopsis floripecten (Rchb.f.) Ames [GU FG] 

Leucohyle mutica (Rchb.f. & Wullschl.) Schlechter [SU] 

Leucohyle subulata (Sw.) Schlechter [GU] 

Ligeophila clavigera (Rchb.f.) Garay [GU SU FG] 

Ligeophila juruensis (Hoehne) Garay [SU] 

Ligeophila petersiana (Cogn.) Garay [SU FG] 

Ligeophila rosea (Lindl.) Garay [SU?] 

Ligeophila stigmatoptera (Rchb.f.) Garay [SU] 

Liparis jamaicensis Griseb. [GU] 

Liparis nervosa (Thunb.) Lindl. [GU? SU FG] 

Lockhartia biserra (L.C. Rich.) Christenson & Garay, comb. 


Lockhartia imbricata (Lam.) Hoehne [GU SU FG] 

Lockhartia latilabris C. Schweinf. [GU] 

Lockhartia parthenocomos Rchb.f. [GU] 

Lycaste macrophylla (Poepp. & Endl.) Lindl. [SU FG] 

Lyroglossa grisebachii (Cogn.) Schlechter [GU SU FG] 

Macradenia loxoglottis H. Focke & Rchb.f. ex Rchb.f. [SU] 

Macradenia lutescens R. Brown [GU SU FG] 

Macroclinium mirabile (Schweinf.) Dodson [GU SU] 

Macroclinium wullschlaegelianum (Focke) Dodson [GU SU 

FG] 

Malaxis excavata (Lindl.) O. Ktze. [SU FG] 

Malaxis maguirei C. Schweinf. [GU SU] 

Malaxis umbellifera Swartz [GU] 

Masdevallia cuprea Lindl. [SU FG] 

Masdevallia guianensis Lindl. [GU SU FG] 

Masdevallia infracta Lindl. [SU] 

Masdevallia minuta Lindl. [GU SU FG] 

Masdevallia norae Luer [SU FG] 

Masdevallia picturata Rchb.f. [GU] 

Maxillaria acutiflora Lindl. [GU FG] 

Maxillaria alba (Hook.) Lindl. [GU SU FG] 

Maxillaria attenuata Ames & Schweinf. [SU] 

Maxillaria auyantepuiensis Foldats [GU SU] 

Maxillaria bolivarensis Schweinf. [GU] 

Maxillaria brachybulbon Schlechter [GU FG] 

Maxillaria brunnea Linden & Rchb.f. [GU SU FG] 

Maxillaria caespitifica Rchb.f. [GU SU FG] 

Maxillaria camaridii Rchb.f. [GU SU FG] 

Maxillaria candida Lodd. ex Lindl. [GU] 

Maxillaria chlorantha Lindl. [GU] 

Maxillaria christobalensis Rchb.f. [GU SU FG] 

Maxillaria connellii Rolfe [GU] 

Maxillaria crassifolia (Lindl.) Rchb.f. [GU FG] 

Maxillaria desvauxiana Rchb.f. [GU SU FG] 

Maxillaria discolor (Lodd. ex Lindl.) Rchb.f. [GU SU FG] 

Maxillaria grobyoides Garay & Dunsterv. [GU] 

Maxillaria imbricata Barb. Rodr. [GU] 

Maxillaria jenischiana (Rchb.f.) Schweinf. [GU SU] 

Maxillaria kegelii Rchb.f. [SU FG] 

Maxillaria lasallei Foldats [GU] 

Maxillaria leucaimata Barb. Rodr. [FG] 

Maxillaria liparophylla Summerh. [GU] 

Maxillaria mapiriensis (Kraenzl.) L.O. Williams [GU SU] 

Maxillaria melina Lindl. [GU] 

Maxillaria nasuta Rchb.f. [GU] 

Maxillaria pannieri Foldats [GU] 

Maxillaria parkeri Hook. [GU SU FG] 

Maxillaria parviflora (Poepp. & Endl.) Garay [GU SU FG] 

Maxillaria pauciflora Barb. Rodr. [GU] 

Maxillaria pendens Pabst [GU] 

Maxillaria ponerantha Rchb.f. [SU FG] 

Maxillaria porrecta Lindl. [GU FG] 

Maxillaria prolifera Ruiz & Pav. [SU] 

Maxillaria pterocarpa Barb. Rodr. [GU] 

Maxillaria pumila Hook. [GU] 

Maxillaria quelchii Rolfe [GU SU] 


157

158 


Maxillaria ramosa Ruiz & Pav. [SU FG] 

Maxillaria reichenheimiana Endres & Rchb.f. [GU SU FG] 

Maxillaria rufescens Lindl. [GU SU FG] 

Maxillaria setigera Lindl. [GU SU?] 

Maxillaria splendens Poepp. & Endl. [SU FG] 

Maxillaria stenophylla Rchb.f. [GU SU FG] 

Maxillaria superflua Rchb.f. [GU SU FG] 

Maxillaria tarumaensis Hoehne [GU SU?] 

Maxillaria trinitatis Ames [GU FG] 

Maxillaria uncata Lindl. [GU SU FG] 

Maxillaria villosa (Barb. Rodr.) Cogn. [GU SU FG] 

Maxillaria violaceopunctata Rchb.f. [GU SU FG] 

Maxillaria xylobiiflora Schlechter [GU SU] 

Mesadenella cuspidata (Lindl.) Garay [GU SU] 

Mormodes atropurpurea Lindl. [GU?] 

Mormodes buccinator Lindl. [SU FG] 

Mormodes igneum Lindl. & Paxton [FG] 

Mormodes vernixioidea Pabst [GU? SU] 

Mormodes vernixium Rchb.f. [GU?] 

Myoxanthus aspasicensis (Rchb.f.) Luer [GU] 

Myoxanthus octomeriae (Schlechter) Luer [GU] 

Myoxanthus parahybunensis (Barb. Rodr.) Luer [GU SU] 

Myoxanthus parvilabius (Schweinf.) Luer [GU SU] 

Myoxanthus simplicicaulis (Schweinf.) Luer [GU] 

Myoxanthus trachychlamys (Schlechter) Luer [GU SU] 

Nidema ottonis (Rchb.f.) Britt. & Millsp. [GU] 

Notylia angustifolia Cogn. [FG] 

Notylia aromatica Barker ex Lindl. [GU SU FG] 

Notylia bungerothii Rchb.f. [GU] 

Notylia durandiana Cogn. [SU] 

Notylia fragrans Wullschl. ex H. Focke [GU SU FG?] 

Notylia laxa Rchb.f. [SU] 

Notylia micrantha Lindl. [GU] 

Notylia microchila Cogn. [GU] 

Notylia peruviana (Schlechter) C. Schweinf. [FG] 

Notylia rhombilabia Schweinf. [GU] 

Notylia sagittifera (Kunth) Link, Klotzsch, & Otto [GU SU 

FG] 

Notylia yauaperiensis Barb. Rodr. [FG] 

Octomeria amazonica Pabst [FG] 

Octomeria brevifolia Cogn. [GU SU FG] 

Octomeria complanata C. Schweinf. [GU] 

Octomeria connellii Rolfe [GU] 

Octomeria deltoglossa Garay [GU SU FG] 

Octomeria erosilabia Schweinf. [GU] 

Octomeria exigua C. Schweinf. [GU] [var. exigua?] 

Octomeria exigua C. Schweinf. var. elata C. Schweinf. [GU] 

Octomeria filifolia Schweinf. [GU] 

Octomeria flaviflora C. Schweinf. [GU] 

Octomeria graminifolia R. Brown [FG] 

Octomeria integrilabia Schweinf. [GU] 

Octomeria minor Schweinf. [GU SU] 

Octomeria monticola Schweinf. [GU] 

Octomeria oxychela Barb. Rodr. [GU] 

Octomeria parvifolia Rolfe [GU] 

Octomeria parvula Schweinf. [GU] 

Octomeria petulans Rchb.f. [FG] 

Octomeria sarthouae Luer [FG] 

Octomeria surinamensis Focke [GU SU FG] 

Odontoglossum naevium Lindl. & Paxt. [GU] 

Oeceoclades maculata (Lindl.) Lindl. [GU SU FG] 

Oliveriana egregia Rchb.f. [GU] 

Oncidium baueri Lindl. [GU SU FG] 

Oncidium cebolleta (Jacq.) Swartz [GU SU FG] 

Oncidium heteranthum Poepp. & Endl. [SU] 


Oncidium lanceanum Lindl. [GU SU FG] 

Oncidium luridum Lindl. [GU FG] 

Oncidium nanum Lindl. [GU FG] 

Oncidium nigratum Lindl. & Paxt. [GU] 

Oncidium orthostates Ridl. [GU] 

Oncidium pirarense Rchb.f. [GU] 

Oncidium tenellum Fr. Gerard [FG] 

Orleansia amazonica Barb. Rodr. [GU SU FG] 

Ornithocephalus bicornis Lindl. [GU SU FG] 

Ornithocephalus bonplandii Rchb.f. [SU] 

Ornithocephalus ciliatus Lindl. [GU SU FG] 

Ornithocephalus dasyrhizus Kraenzl. [FG?] 

Ornithocephalus gladiatus Hook. [GU SU FG] 

Ornithocephalus kruegeri Rchb.f. [SU FG] 

Otostylis lepida (Linden & Rchb.f.) Schlechter [GU SU] 

Palmorchis guianensis (Schlechter) Schweinf. & Correll [GU 

SU] 

Palmorchis kuhlmannii (Schlechter) L.O. Williams [SU] 

Palmorchis lobulata (Mansf.) Schweinf. & Corr. [FG] 

Palmorchis pabstii Veyret [FG] 

Palmorchis prospectorum Veyret [SU FG] 

Palmorchis pubescens Barb. Rodr. [GU FG] 

Palmorchis trilobulata L.O. Williams [SU] 

Paphinia cristata (Lindl.) Lindl. [GU SU FG] 

Paphinia lindeniana Rchb.f. [GU?] 

Pelexia callifera (C. Schweinf.) Garay [SU FG] 

Pelexia delicatula Szlachetko & Veyret [FG] 

Pelexia goninensis (Pulle) Schlechter [GU SU FG] 

Pelexia laxa (Poepp. & Endl.) Lindl. [SU?] 

Peristeria cerina Lindl. [GU] 

Peristeria guttata Knowles & Westcott [GU SU FG] 

Peristeria pendula W.J. Hook. [GU SU FG] 

Peristeria rossiana Rchb.f. [GU? SU] 

Peristeria selligera Rchb.f. [GU] 

Phragmipedium klotzscheanum (Rchb.f.) Rolfe [GU] 

Phragmipedium lindleyanum (Schomb.) Rolfe [GU SU FG] 

Platystele ovalifolia (H. Focke) Garay & Dunsterv. [GU SU 

FG] 

Platystele stenostachya (Rchb.f.) Garay [SU] 

Platythelys maculata (Hook.) Garay [SU FG] 

Plectrophora cultrifolia (Barb. Rodr.) Cogn. [FG] 

Plectrophora iridifolia H. Focke [GU SU FG] 

Pleurothallis arachnopsis H.G. Jones [GU] 

Pleurothallis archidiaconi Ames [GU SU FG] 

Pleurothallis aristata W.J. Hook. [GU SU FG] 

Pleurothallis barboselloides Schlechter [SU] 

Pleurothallis barbulata Lindl. [GU SU FG] 

Pleurothallis barthelemyi Luer [FG] 

Pleurothallis biglandulosa Schlechter [FG] 

Pleurothallis brevipes H. Focke [GU SU FG] 

Pleurothallis breviscapa Schweinf. [GU FG] 

Pleurothallis cabellensis Rchb.f. [FG] 

Pleurothallis ciliolata Schlechter [GU SU FG] 

Pleurothallis consimilis Ames [GU SU FG] 

Pleurothallis corniculata (Swartz) Lindl. [GU] 

Pleurothallis curvifructa H.G. Jones [GU] 

Pleurothallis determannii Luer [GU SU] 

Pleurothallis discoidea Lindl. [SU FG] 

Pleurothallis ephemera Lindl. [SU FG] 

Pleurothallis erinacea Rchb.f. [GU SU?] 

Pleurothallis exigua Cogn. [GU] 

Pleurothallis fockei Lindl. [GU SU] 

Pleurothallis glandulosa Ames [GU SU FG] 

Pleurothallis grobyi J. Bateman ex Lindl. [GU SU FG] 

Pleurothallis hitchcockii Ames [GU]


Pleurothallis hymenantha Lindl. [GU] 

Pleurothallis imraei Lindl. [GU SU] 

Pleurothallis kegelii Rchb.f. [SU] 

Pleurothallis lanceana Lodd. [GU SU FG] 

Pleurothallis lancipetala (Karst.) Schlechter [GU] 

Pleurothallis loranthophylla Rchb.f. [GU] 

Pleurothallis minima Schweinf. [GU] 

Pleurothallis miqueliana (H. Focke) Lindl. [GU SU] 

Pleurothallis monophylla (W.J. Hook.) Fawc. & Rendle [GU] 

Pleurothallis moritzii Rchb.f. [GU] [Mt. Roraima] 

Pleurothallis nanifolia Foldats [FG?] 

Pleurothallis obovata (Lindl.) Lindl. [GU] 

Pleurothallis omoglossa Luer [FG] 

Pleurothallis picta Lindl. [GU SU FG] 

Pleurothallis polygonoides Griseb. [GU SU FG] 

Pleurothallis pruinosa Lindl. [GU SU FG] 

Pleurothallis pubescens Lindl. [SU FG] 

Pleurothallis revoluta (Ruiz & Pav.) Garay [SU] 

Pleurothallis rhombipetala Rolfe [GU] 

Pleurothallis ruscifolia (Jacq.) R. Brown [GU SU FG] 

Pleurothallis sclerophylla Lindl. [GU SU? FG] 

Pleurothallis secunda Poepp. & Endl. [FG] 

Pleurothallis semperflorens Lindl. [GU SU FG] 

Pleurothallis seriata Lindl. [GU SU FG] 

Pleurothallis spiculifera Lindl. [GU SU FG] 

Pleurothallis stenocardium Schlechter [GU] 

Pleurothallis stenopetala Lodd. ex Lindl. [GU] [=Pleurothallis 

sclerophylla?] 

Pleurothallis suspensa Luer [GU SU FG] 

Pleurothallis testaefolia (Swartz) Lindl. [SU?] 

Pleurothallis tridentata Klotzsch [GU] 

Pleurothallis uniflora Lindl. [GU SU FG] 

Pleurothallis yauaperiensis Barb. Rodr. [SU] 

Polycycnis surinamensis C. Schweinf. [GU SU] 

Polystachya concreta (Jacq.) Garay & Sweet [GU SU FG] 

Polystachya d'agremondiana Kraenzl. [SU] [dubious species] 

Polystachya foliosa (Hook.) Rchb.f. [GU SU FG] 

Polystachya stenophylla Schlechter [GU SU FG] 

Ponthieva diptera Lind. & Rchb.f. [GU] 

Ponthieva ovatilabia Schweinf. [GU] 

Prescottia carnosa C. Schweinf. [GU] [Mt. Roraima] 

Prescottia oligantha (Sw.) Lindl. [GU] 

Prescottia stachyodes (Swartz) Lindl. [GU SU FG] 

Psilochilus macrophyllus (Lindl.) Ames [GU] 

Psilochilus physurifolius (Rchb.f.) Lojtnant [GU] 

Psychopsis kramerianum (Rchb.f.) H.G. Jones [SU] 

Psychopsis papilio (Lindl.) H.G. Jones [SU FG] 

Psychopsis versteegianum (Pulle) Luckel & Braem [SU] 

Psygmorchis glossomystax (Rchb.f.) Dodson & Dressler [GU 

SU FG] 

Psygmorchis pumilio (Rchb.f.) Dodson & Dressler [SU FG] 

Psygmorchis pusilla (L.) Dodson & Dressler [GU SU FG] 

Quekettia jenmanii Rchb.f. [GU] 

Quekettia microscopica Lindl. [GU SU FG] 

Quekettia papillosa Garay [SU] 

Quekettia vermeuleniana R.O. Determann [GU SU] 

Reichenbachanthus reflexus (Lindl.) Brade [GU SU FG] 

Rodriguezia candida (Lindl.) Rchb.f. [SU FG] 

Rodriguezia cinnabarina H.G. Jones [GU] 

Rodriguezia flavida Garay & Dunsterv. [GU SU] 

Rodriguezia lanceolata Ruiz & Pav. [GU SU FG] 

Rodriguezia luteola N.E. Brown [GU] 

Rudolfiella aurantiaca (Lindl.) Hoehne [GU SU FG] 

Rudolfiella bicornaria (Rchb.f.) Hoehne [SU] 

Sacoila hassleri (Cogn.) Garay [SU] 


159 

Sacoila lanceolata (Aubl.) Garay [GU SU FG] 

Sarcoglottis acaulis (J.E. Smith) Schlechter [GU SU FG] 

Sarcoglottis amazonica Pabst [SU FG] 

Sarcoglottis aphylla (Ridl.) Schlechter [GU] 

Sarcoglottis degranvillei Szlachetko & Veyret [FG] 

Sarcoglottis grandiflora (Hook.) Klotzsch [GU SU FG?] 

Sarcoglottis metallica (Rolfe) Schlechter [GU] 

Sarcoglottis rupestris (Lindl.) Barb. Rodr. [SU] 

Sarcoglottis sagittata (Rchb.f. & Warm.) Schlechter [FG] 

Sarcoglottis simplex (Griseb.) Schlechter [GU] 

Sarcoglottis stergiosii Carnevali & I. Ramírez [GU] 

Scaphosepalum breve (Rchb.f.) Rolfe [GU] 

Scaphyglottis bifida (Rchb.f.) Schweinf. [SU FG] 

Scaphyglottis dunstervillei (Garay) Foldats [SU] 

Scaphyglottis felskyi (Rchb.f.) Schlechter [SU] 

Scaphyglottis fusiformis (Griseb.) Schultes [GU SU FG] 

Scaphyglottis graminifolia (Ruiz & Pav.) Poepp. & Endl. [GU 

FG] 

Scaphyglottis grandiflora Ames & Schweinf. [GU] 

Scaphyglottis lindeniana (A. Rich. & Gal.) L.O. Williams [GU 

SU] 

Scaphyglottis modesta (Rchb.f.) Schlechter [GU SU FG] 

Scaphyglottis ochroleuca Schlechter [SU FG] 

Scaphyglottis prolifera Cogn. [GU SU FG] 

Scaphyglottis propinqua Schweinf. [SU] 

Scaphyglottis sickii Pabst [GU SU FG] 

Scaphyglottis stellata Lodd. ex Lindl. [GU SU FG] 

Scaphyglottis violacea Lindl. [GU SU FG] 

Scelochilus ecalcaratus Determann [FG] 

Schlimia alpina Rchb.f. & Warsc. [GU] 

Schomburgkia crispa Lindl. [GU? SU? FG?] [most (all?) 

records in Guianas =S. marginata] 

Schomburgkia gloriosa Rchb.f. [GU SU?] 

Schomburgkia marginata Lindl. [GU SU FG] 

Schomburgkia rosea Linden ex Lindl. [GU] 

Scuticaria hadwenii Hort. ex Planch. var. dodgsoni (B.S. 

Wms.) Stein [GU] 

Scuticaria steelei (Hook.) Lindl. [GU SU FG] 

Selenipedium palmifolium (Lindl.) Rchb.f. [GU SU FG] 

Selenipedium steyermarkii Foldats [GU] 

Sievekingia jenmanii Rchb.f. [GU SU? FG] 

Sigmatostalix amazonica Schlechter [SU FG] 

Sobralia crocea (Poepp. & Endl.) Rchb.f. [SU?] 

Sobralia fimbriata Poepp. & Endl. [SU FG] 

Sobralia fragrans Lindl. [GU SU FG] 

Sobralia infundibuligera Garay & Dunsterv. [GU] 

Sobralia liliastrum Lindl. [GU SU? FG] 

Sobralia macrophylla Rchb.f. [GU SU FG] 

Sobralia oliva-estevae Carnevali & Ramirez [GU] 

Sobralia sclerophylla Lindl. [FG] 

Sobralia sessilis Lindl. [GU SU FG] 

Sobralia stenophylla Lindl. [GU SU? FG?] 

Sobralia suaveolens Rchb.f. [GU SU FG] 

Sobralia valida Rolfe [GU] 

Sobralia violacea Linden ex Lindl. [SU?] 

Sobralia yauaperyensis Barb. Rodr. [FG] 

Solenidium lunatum (Lindl.) Kraenzl. [GU] 

Spiranthes tenuis Lindl. [GU SU FG] 

Spiranthes torta (Thunb.) Garay & Sweet [FG?] 

Stanhopea candida Barb. Rodr. [GU?] 

Stanhopea grandiflora (Lodd.) Lindl. [GU SU FG] 

Stelis aprica Lindl. [GU SU] 

Stelis argentata Lindl. [GU SU FG] 

Stelis cuprea Lindl. [FG] [dubious record] 

Stelis grandiflora Lindl. [GU]

160 


Stelis guianensis Rolfe [GU FG] 

Stelis intermedia Poepp. & Endl. [GU SU] 

Stelis muscifera Lindl. [GU] 

Stelis ophioglossoides Swartz [SU? FG] 

Stelis papaquerensis Rchb.f. [SU? FG] 

Stelis perparva Schweinf. [GU] 

Stelis pygmaea Cogn. [FG] 

Stelis santiagoensis Mansf. [GU FG] 

Stelis schomburgkii Fawc. & Rendle [GU] 

Stelis tristyla Lindl. [GU SU] 

Stelis zonata Rchb.f. [GU] [=Stelis muscifera?] 

Stenia pallida Lindl. [GU] 

Teuscheria dodsonii Dressl. [SU?] 

Trichocentrum fuscum Lindl. [SU FG] 

Trichocentrum purpureum Lindl. ex Rchb.f. [GU] 

Trichocentrum recurvum Lindl. [GU SU] 

Trichopilia wageneri Rchb.f. [GU SU] 

Trichosalpinx ciliaris (Lindl.) Luer [GU SU FG] 

Trichosalpinx egleri (Pabst) Luer [GU SU FG] 

Trichosalpinx foliata (Griseb.) Luer [SU] 

Trichosalpinx memor (Rchb.f.) Luer [GU SU] 

Trichosalpinx orbicularis (Lindl.) Luer [GU SU FG] 

Trichosalpinx roraimensis (Rolfe) Luer [GU] 

Trigonidium acuminatum Batem. [GU SU FG] 

Trigonidium egertonianum Batem. ex Lindl. [GU] 

Trigonidium macranthum Barb. Rodr. [GU] 

Trigonidium obtusum Lindl. [GU SU? FG] 

Triphora amazonica Schlechter [GU SU? FG] 

Triphora surinamensis (Lindl. ex Benth.) Britton [GU SU FG] 

Trisetella triglochin (Rchb.f.) Luer [SU FG] 

Trizeuxis falcata Lindl. [GU] 

Uleiorchis ulei (Cogn.) Handro [GU FG] 

Vanilla acuta Rolfe [GU SU FG] 

Vanilla appendiculata Rolfe [GU] 

Vanilla barrereana Veyret & Szlachetko [FG] 

Vanilla bicolor Lindl. [GU SU FG] 

Vanilla chamissonis Klotzsch [FG] 

Vanilla cristato-callosa Hoehne [GU] 

Vanilla fimbriata Rolfe [GU] 

Vanilla grandiflora Lindl. [GU SU FG?] 

Vanilla guianensis Splitg. [SU] [name of uncertain 

application; =V. mexicana?] 

Vanilla hostmanni Rolfe [SU] 

Vanilla latisegmenta Ames & Schweinf. [GU] 

Vanilla leprieurii R. Portères [FG] 

Vanilla marowynensis Pulle [SU] 

Vanilla mexicana P. Mill. [GU SU FG] 

Vanilla odorata Presl [FG] 

Vanilla ovata Rolfe [GU? FG] 

Vanilla palmarum Lindl. [GU SU FG] 

Vanilla planifolia G. Jackson [GU SU] 

Vanilla porteresiana Veyret & Szlachetko [FG] 

Vanilla surinamensis Rchb.f. [SU] 

Wullschlaegelia calcarata Benth. [GU SU FG] 

Xerorchis amazonica Schlechter [SU] 

Xerorchis trichorhiza (Kraenzl.) Garay [GU FG] 

Xylobium chapadense (Barb. Rodr.) Cogn. [GU] 

Xylobium colleyi (Batem.) Rolfe [GU SU?] 

Xylobium foveatum (Lindl.) Nichols. [GU SU FG] 

Xylobium pallidiflorum (Hook.) Nichols. [SU] 

Xylobium palmifolium (Swartz) Benth. ex Fawc. [GU] 

Xylobium steyermarkii Foldats [GU] 

Xylobium variegatum (Ruiz & Pav.) Garay & Dunsterv. [SU 

FG] 

Zygosepalum kegelii (Rchb.f.) Rchb.f. [SU] 


Zygosepalum labiosum (L.C. Rich.) Schweinf. [GU SU FG] 

Zygosepalum lindeniae (Rolfe) Garay & Dunsterv. [FG] 

Zygosepalum tatei (Ames & Schweinf.) Garay & Dunsterv. 

[GU] 

Zygostates sp. nov. [SU] 

ORCHIDACEAE - SYNONYMS AND 


Aganisia fimbriata Rchb.f. =Acacallis fimbriata 

Angraecum fasciola Lindl. =Campylocentrum fasciola 

Aulizia ciliaris (L.) Salisb. =Epidendrum ciliare 

Bifrenaria aurantiaca Lindl. =Rudolfiella aurantiaca 

Bifrenaria bicornaria Rchb.f. =Rudolfiella bicornaria 

Bifrenaria sabulosa Barb. Rodr. =Bifrenaria longicornis 

Bletia verecunda R. Brown =Bletia purpurea 

Bollea guyanensis Klotzsch [nomen nudum] 

Bonatea macilenta (Lindl.) Rchb.f. =Habenaria macilenta 

Brassavola angustata Lindl. =Brassavola martiana 

Brassavola surinamensis Focke =Brassavola martiana 

Brassia attenuata H.G. Jones =Brassia chloroleuca 

Brassia verrucosa Auctt., non Lindl. =Brassia spp. 

Burlingtonia candida Lindl. =Rodriguezia candida 

Campylocentrum burchellii Cogn. [not in Guianas] 

Catasetum discolor (Lindl.) Lindl. var. roseo-album (Hook.) 

Mansf. =Catasetum xroseo-album 

Catasetum fuliginosum Lindl. =Catasetum callosum var. 

carunculatum? 

Cattleya pumila Schomb. =Cattleya lawrenceana 

Cattleya schomburgkii Lodd. ex Lindl. =Cattleya violacea 

Cattleya superba Schomb. ex Lindl. =Cattleya violacea 

Chaubardia tigrina Garay & Dunsterv. =Chaubardiella tigrina 

Cheiradenia imthurnii Cogn. =Cheiradenia cuspidata 

Cladobium violaceum Lindl. =Scaphyglottis violacea 

Cochleanthes amazonica (Rchb.f.) Schultes & Garay [not in 


Cochleanthes flabelliformis (Swartz) Schultes & Garay [not in 


Cryptarrhena acrensis Schlechter =Cryptarrhena guatemalensis 

Cryptarrhena ghillanyi Pabst =Cryptarrhena guatemalensis 

Cryptarrhena lunata Auctt., non R. Brown =Cryptarrhena 

kegelii 

Cyclopogon stictophyllus Schlechter =Cyclopogon olivaceus 

Cypripedium klotzscheanum Rchb.f. ex Schomb. 

=Phragmipedium klotzscheanum 

Cypripedium lindleyanum Schomb. =Phragmipedium 

lindleyanum 

Cypripedium palmifolium Lindl. =Selenipedium palmifolium 

Cypripedium schomburgkianum Klotzsch ex Schomb. 

=Phragmipedium klotzscheanum 

Cyrtopera longifolia Rchb.f. =Eulophia alta 

Cyrtopodium glutiniferum Raddi [dubious in Guianas] 

Cyrtopodium punctatum Lindl. [dubious in Guianas] 

Diacrium bicornutum Benth. =Caularthron bicornutum 

Dichaea anchoraelabia Schweinf. [Peruvian taxon, unlikely in 

Guianan flora] 

Dichaea brachypoda Rchb.f. [Costa Rican taxon, name widely 

misapplied] 

Dichaea graminoides (Swartz) Lindl. [name (mis?)applied to 

D. rendlei in Guianas] 

Dichaea muricata (Swartz) Lindl. [rejected name] 

Dichaea trinitensis Gleason =Dichaea picta 

Dicrypta discolor Lodd. ex Lindl. =Maxillaria discolor 

Dimerandra isthmii Schlechter =Dimerandra elegans


Dimerandra stenopetala Auctt., non (Hook.) Schlechter 

=Dimerandra spp.[name widely misapplied] 

Diothonea gratissima Rchb.f. =Epidendrum gratissimum 

Elleanthus linifolius Auctt., non K. Presl =Elleanthus 

graminifolius 

Elleanthus strobilifer (Poepp. & Endl.) Rchb.f. [not in 


Encyclia abbreviata Auctt., non (Schlechter) Dressl. =Encyclia 

calamaria 

Encyclia atropurpurea Schlechter =Encyclia cordigera 

Encyclia ciliaris (L.) Lemée =Epidendrum ciliare 

Encyclia crassilabia (Poepp. & Endl.) Lemée =Encyclia vespa 

Encyclia fragrans (Swartz) Lemée var. aemula (Lindl.) 

Dressler =Encyclia aemula 

Encyclia oncidioides (Lindl.) Schlechter [not in Guianas] 

Encyclia palacea (Rchb.f.) Lemée =Nidema boothii [not in 


Encyclia selligera (Batem. ex Lindl.) Schlechter [not in 


Epidendropsis violascens (Ridl.) Garay & Dunsterv. 

=Epidendrum violascens 

Epidendrum aemulum Lindl. =Encyclia aemula 

Epidendrum alsum Ridl. =Epidendrum dendrobioides 

Epidendrum bicornutum Hook. =Caularthron bicornutum 

Epidendrum bifidum Aubl. [not in Guianas] 

Epidendrum boothii L. Will. =Nidema boothii [not in Guianas] 

Epidendrum caespitosum Barb. Rodr. =Epidendrum 

orchidiflorum 

Epidendrum chloroleucum W.J. Hook. =Encyclia chloroleuca 

Epidendrum clavatum Lindl. =Epidendrum purpurascens 

Epidendrum cochleatum L. =Encyclia cochleata 

Epidendrum coriifolium Lindl. [all material probably referable 

to E. imitans in Guianas] 

Epidendrum decipiens Auctt., non Lindl. =Epidendrum 

ibaguense 

Epidendrum diurnum (Jacq.) Cogn. =Encyclia diurna 

Epidendrum durum Lindl. =Epidendrum dendrobioides 

Epidendrum floribundum Kunth =Epidendrum densiflorum 

Epidendrum florijugum Barb. Rodr. =Epidendrum sculptum 

Epidendrum fragrans Swartz =Encyclia fragrans [in Guianas, 

=Encyclia aemula] 

Epidendrum fulgens Brongn. =Epidendrum ibaguense? 

Epidendrum fuscatum Sm. =Epidendrum anceps 

Epidendrum globosum Jacq. =Jacquiniella globosa 

Epidendrum graniticum Lindl. =Encyclia granitica 

Epidendrum huebneri Schlechter =Epidendrum orchidiflorum 

Epidendrum ibaguense Kunth var. schomburgkii (Lindl.) 

Schweinf. =Epidendrum macrocarpum 

Epidendrum imatophyllum Lindl. =Epidendrum flexuosum 

Epidendrum imbricatum Lam. =Lockhartia imbricata 

Epidendrum incisum Vell. =Epidendrum macrocarpum 

Epidendrum ionosmum Lindl. =Encyclia ionosma 

Epidendrum labiosum L.C. Rich. =Zygosepalum labiosum 

Epidendrum latifolium (Lindl.) Garay & Sweet =Epidendrum 

carpophorum 

Epidendrum latipetalum C. Schweinf. =Encyclia latipetala 

Epidendrum maculatum Aubl. =Oncidium luridum 

Epidendrum montigenum Ridl. =Epidendrum dendrobioides 

Epidendrum nocturnum Jacq. var. latifolium Lindl. 

=Epidendrum carpophorum 

Epidendrum nocturnum Jacq. var. minus Cogn. =Epidendrum 

longicolle 

Epidendrum nocturnum Jacq. var. tumuc-humaciense Veyret 

=Epidendrum tridens 

Epidendrum odoratissimum Lindl. =Encyclia odoratissima 


161 

Epidendrum oncidioides Lindl. =Encyclia oncidioides (Lindl.) 

Schlechter [not in Guianas] 

Epidendrum pachyanthum Lindl. =Encyclia pachyantha 

Epidendrum pictum Lindl. =Encyclia picta 

Epidendrum pusillum L. =Psygmorchis pusilla 

Epidendrum raniferum Auctt., non Lindl. =Epidendrum 

cristatum 

Epidendrum rectopedunculatum C. Schweinf. =Epidendrum 

orchidiflorum 

Epidendrum schomburgkii Lindl. =Epidendrum macrocarpum 

Epidendrum secundum Auctt., non Jacq. =Epidendrum anceps 

Epidendrum spruceanum Lindl. =Epidendrum nocturnum 

Epidendrum stenopetalum Hook. =Dimerandra stenopetala 

[name misapplied] 

Epidendrum ulei Schlechter =Epidendrum imthurnii 

Epidendrum variegatum Hook. =Encyclia vespa 

Eriopsis schomburgkii (Rchb.f.) Rchb.f. =Eriopsis biloba 

Erythrodes petersiana (Cogn.) Ames =Ligeophila petersiana 

Erythrodes plantaginea (L.) Fawc. & Rendle [dubiously in 


Erythrodes santensis (Kraenzl.) C. Schweinf. =Ligeophila 

juruensis 

Fractiunguis reflexa Schlechter =Reichenbachanthus reflexus 

Galeandra juncea Lindl. =Galeandra stylomisantha 

Galeandra montana Barb. Rodr. =Galeandra lagoensis 

Reichb.f. & Warm.[dubious in Guianas] 

Gongora macrantha Hook. =Coryanthes macrantha 

Gongora quinquenervis Ruiz & Pav. [not in Guianas] 

Goodyera guyanensis Lindl. =Brachystele guyanensis 

Habenaria amazonica Schlechter =Habenaria platydactyla 

Habenaria culmiformis Schlechter =Habenaria leprieurii 

Habenaria demerarensis Rchb.f. ex Schomb. =Habenaria 

parviflora 

Habenaria fastor Auctt., non Warming =Habenaria longicauda 

Habenaria goyazensis Cogn. [as circumscribed by Renz, not in 


Habenaria leaoana Schlechter =Habenaria sprucei 

Habenaria macroceratilis Willd. =Habenaria quinqueseta 

Habenaria moritzii Ridl. =Habenaria entomantha 

Habenaria muelleriana Cogn. =Habenaria macilenta 

Habenaria pauciflora (Lindl.) Rchb.f. =Habenaria trifida 

Habenaria savannensis Renz =Habenaria amalfitana 

Habenaria staminodiata Schlechter =Habenaria macilenta 

Hexadesmia fusiformis Griseb. =Scaphyglottis fusiformis 

Hexisea reflexa (Lindl.) Rchb.f. ex Griseb. 

=Reichenbachanthus reflexus 

Houlletia vittata Lindl. =Braemia vittata 

Humboldtia ciliata (Knowl. & Westc.) Kuntze =Pleurothallis 

lanceana 

Humboldtia fockei (Lindl.) Kuntze =Pleurothallis fockei 

Humboldtia grobyi (Lindl.) Kuntze =Pleurothallis grobyi 

Humboldtia miqueliana (Focke) Kuntze =Pleurothallis 

miqueliana 

Humboldtia pruinosa (Lindl.) Kuntze =Pleurothallis pruinosa 

Huntleya violacea Lindl. =Bollea violacea 

Ionopsis paniculata Lindl. =Ionopsis utricularioides 

Ionopsis teres Lindl. =Ionopsis satyrioides 

Isochilus elegans Focke =Dimerandra elegans 

Kefersteinia pusilla Auct., non (Schweinf.) Schweinf. 

=Kefersteinia lafontanei 

Kegelia houtteana Rchb.f. =Kegeliella houtteana 

Laelia crispa (Lindl.) Lemée =Schomburgkia crispa 

Laelia marginata (Lindl.) L.O. Williams =Schomburgkia 

marginata

162 


Lanium microphyllum (Lindl.) Benth. =Epidendrum 

microphyllum 

Lanium subulatum Rolfe =Epidendrum stiliferum 

Limodorum grandiflorum Aubl. =Cleistes grandiflora 

Limodorum lanceolatum Aubl. =Sacoila lanceolata 

Limodorum pendulum Aubl. =Dichaea pendula 

Lindleyella aurantiaca (Lindl.) Schlechter =Rudolfiella 

aurantiaca 

Liparis elliptica (Rchb.f.) Rchb.f., non Wight =Liparis 

jamaicensis 

Liparis kappleri (Rchb.f.) Rchb.f. =Liparis nervosa 

Liparis vexillifera (Llave & Lex.) Cogn. [not in Guianas] 

Lockhartia elegans Hook. =Lockhartia imbricata 

Lockhartia micrantha Rchb.f. [not in Guianas] 

Lockhartia weigeltii Rchb.f. =Lockhartia imbricata 

Lycomormium minus Kraenzl. =Peristeria cerina 

Macradenia surinamensis Rchb.f. & Wullschl. =Macradenia 

lutescens 

Macradenia triandra Lindl. =Macradenia lutescens 

Masdevallia brevis Rchb.f. =Scaphosepalum breve 

Masdevallia cayennensis Rchb.f. =Masdevallia cuprea 

Masdevallia guttulata Auctt., non Rchb.f. =Masdevallia norae 

Masdevallia huebneri Schlechter =Trisetella triglochin 

Masdevallia surinamensis Focke =Masdevallia minuta 

Maxillaria conferta (Griseb.) Schweinf. ex Leon =Maxillaria 

parviflora 

Maxillaria foveata Lindl. =Xylobium foveatum 

Maxillaria graminea Lindl. =Koellensteinia graminea 

Maxillaria nana Hook. =Maxillaria uncata 

Maxillaria ringens Rchb.f. =Maxillaria brunnea 

Maxillaria steelei Hook. =Scuticaria steelei 

Maxillaria variabilis Auctt., non J. Bateman ex Lindl. 

=Maxillaria caespitifica 

Maxillaria vestita (Swartz) Lemée =Maxillaria parviflora 

Menadenium kegelii (Rchb.f.) Cogn. =Zygosepalum kegelii 

Menadenium labiosum (L.C. Rich.) Cogn. =Zygosepalum 

labiosum 

Mendoncella burkei (Rchb.f.) Garay =Galeottia burkei 

Mesadenella esmeraldae (Lindl. & Rchb.f.) Pabst & Garay 

=Mesadenella cuspidata 

Monachanthus discolor Lindl. =Catasetum discolor 

Myanthus barbatus Lindl. =Catasetum barbatum 

Myanthus deltoideus Lindl. =Catasetum deltoideum 

Nanodes discolor Lindl. =Epidendrum schlechterianum 

Neobartlettia guianensis Schlechter =Palmorchis guianensis 

Neolehmannia apagana (Mansf.) Garay & Dunsterv. 

=Epidendrum apaganum 

Nidema boothii (Lindl.) Schlechter [not in Guianas] 

Notylia amesiana Smith & Harris =Notylia microchila 

Notylia bicolor Lindl. =Macroclinium bicolor (Lindl.) 

Dodson[not in Guianas] 

Notylia mirabilis Schweinf. =Macroclinium mirabile 

Notylia wullschlaegeliana Focke =Macroclinium 

wullschlaegelianum 

Octomeria grandiflora Lindl. [not in Guianas] 

Octomeria pusilla Lindl. [not in Guianas] 

Oncidium altissimum Auctt., non (Jacq.) Swartz =Oncidium 

baueri 

Oncidium emarginatum Mey. =Dimerandra emarginata 

Oncidium glossomystax Rchb.f. =Psygmorchis glossomystax 

Oncidium guianense (Aubl.) Garay [not in Guianas] 

Oncidium kappleri Rchb.f. =Oncidium baueri 

Oncidium lunatum Lindl. =Solenidium lunatum 

Oncidium maculatum Urb. =Oncidium luridum 

Oncidium papilio Lindl. =Psychopsis papilio 


Oncidium pusillum (L.) Rchb.f. =Psygmorchis pusilla 

Oncidium ultrajectinum Pulle =Oncidium cebolleta 

Oncidium versteegianum Pulle =Psychopsis versteegianum 

Ophrys guianensis Aubl. =Oncidium guianense (Aubl.) Garay 


Ophrys peruviana Aubl. =Spiranthes torta 

Ornithocephalus avicula Rchb.f. =Ornithocephalus kruegeri 

Ornithocephalus falcatus Focke =Ornithocephalus gladiatus 

Otostylis brachystalix (Rchb.f.) Schlechter =Otostylis lepida 

Paphiopedilum klotzscheanum Pfitz. =Phragmipedium 

klotzscheanum 

Paphiopedilum lindleyanum Pfitz. =Phragmipedium 

lindleyanum 

Pelexia aphylla Ridl. =Sarcoglottis aphylla 

Peristeria aspersa Rolfe =Peristeria rossiana 

Pilumna wageneri Rchb.f. =Trichopilia wageneri 

Pleurothallis acuminata Focke =Pleurothallis semperflorens 

Pleurothallis acutissima Lindl. =Pleurothallis spiculifera 

Pleurothallis blaisdellii S. Wats. =Trichosalpinx blaisdellii 

(Wats.) Luer [not in Guianas] 

Pleurothallis ciliaris (Lindl.) L.O. Wiliams =Trichosalpinx 

ciliaris 

Pleurothallis ciliata Knowl. & Westc. =Pleurothallis lanceana 

Pleurothallis diffusiflora Schweinf. =Pleurothallis seriata 

Pleurothallis leptopetala Cogn. =Pleurothallis discoidea 

Pleurothallis longirostris Focke =Pleurothallis ephemera 

Pleurothallis marginalis Rchb.f. =Pleurothallis grobyi 

Pleurothallis memor Rchb.f. =Trichosalpinx memor 

Pleurothallis monocardia Auctt., non Rchb.f. =Pleurothallis 

archidiaconi 

Pleurothallis orbicularis (Lindl.) Lindl. =Trichosalpinx 

orbicularis 

Pleurothallis ovalifolia Rchb.f. =Platystele ovalifolia 

Pleurothallis parvilabia Schweinf. =Myoxanthus parvilabius 

Pleurothallis peduncularis Lindl. =Myoxanthus 

parahybunensis 

Pleurothallis penduliflora Auctt., non Kraenzl. =Pleurothallis 

suspensa 

Pleurothallis pertenuis Schweinf. =Pleurothallis glandulosa 

Pleurothallis roraimensis Rolfe =Trichosalpinx roraimensis 

Pleurothallis striata Focke =Pleurothallis uniflora 

Pleurothallis tricarinata Focke =Pleurothallis fockei 

Pleurothallis villosa Knowl. & Westc. [in Guianas, 

=Trichosalpinx egleri] 

Pleurothallis vittariifolia Schlechter =Pleurothallis glandulosa 

Pogonia grandiflora Rchb.f. =Cleistes grandiflora 

Pogonia macrophylla Lindl. =Psilochilus macrophyllus 

Pogonia paludosa (Rchb.f.) Rchb.f. =Cleistes paludosa 

Pogonia parviflora (Lindl.) Rchb.f. =Cleistes parviflora 

Pogonia physurifolia Rchb.f. =Psilochilus physurifolius 

Pogonia rosea (Lindl.) Rchb.f. =Cleistes rosea 

Pogonia surinamensis Lindl. =Triphora surinamensis 

Pogonia tenuis Rchb.f. =Cleistes tenuis 

Pogonia triflora Steyerm. =Cleistes moritzii 

Pogonia unifoliata Schweinf. =Cleistes unifoliata 

Polystachya estrellensis Rchb.f. 

Polystachya geraensis Barb. Rodr. 

Polystachya luteola (Swartz) Hook. =Polystachya concreta 

Polystachya minuta Britt.


Polystachya nana Poepp. & Endl. 

Ponera felskii Rchb.f. =Scaphyglottis felskyi 

Pseuderiopsis schomburgkii Rchb.f. =Eriopsis biloba 

Restrepia kegelii Rchb.f. =Chamelophyton kegelii 

Rodriguezia secunda Kunth =Rodriguezia lanceolata 

Sarcoglottis maasorum Pabst =Sarcoglottis metallica 

Scaphosepalum ochthodes Pfitz. [not in Guianas] 

Scaphyglottis amethystina (Rchb.f.) Schlechter =Scaphyglottis 

stellata 

Scaphyglottis cuneata Schlechter =Scaphyglottis prolifera 

Scaphyglottis reflexa Lindl. =Reichenbachanthus reflexus 

Schlechterella aurantiaca (Lindl.) Hoehne =Rudolfiella 

aurantiaca 

Selenipedium lindleyanum Rchb.f. =Phragmipedium 

lindleyanum 

Selenipedium schomburgkianum Rchb.f. =Phragmipedium 

klotzscheanum 

Serapias caravata Aubl. =Elleanthus caravata 

Specklinia miqueliana Focke =Pleurothallis miqueliana 

Specklinia orbicularis Lindl. =Trichosalpinx orbicularis 

Spiranthes acaulis (J.E. Smith) Cogn. =Sarcoglottis acaulis 

Spiranthes aphylla (Ridl.) Cogn. =Sarcoglottis aphylla 

Spiranthes bifida Ridl. =Cyclopogon bifidus 

Spiranthes callifera Schweinf. =Pelexia callifera 

Spiranthes elata (Swartz) L.C. Rich. =Cyclopogon elatus 

Spiranthes goninensis (Pulle) Schweinf. =Pelexia goninensis 

Spiranthes grisebachii Cogn. =Lyroglossa grisebachii 

Spiranthes guianensis (Lindl.) Cogn. =Brachystele guyanensis 

Spiranthes lanceolata (Aubl.) Leon =Sacoila lanceolata 

Spiranthes metallica Rolfe =Sarcoglottis metallica 

Spiranthes quinqueloba (Poir.) Urb. =Spiranthes torta 

Stanhopea eburnea Lindl. =Stanhopea grandiflora 

Stelis drosophila Barb. Rodr. =Stelis intermedia 

Stelis flavida Focke =Pleurothallis pruinosa 

Stelis foliosa W.J. Hook. =Polystachya foliosa 

Stenoptera adnata Ridl. =Gomphichis adnata 

Stenorrhynchos goninensis Pulle =Pelexia goninensis 

Stenorrhynchos lanceolatum (Aubl.) L.C. Rich. ex Spreng. 

=Sacoila lanceolata 

Sturmia kappleri Rchb.f. =Liparis nervosa 

Trichocentrum cornucopiae Linden & Rchb.f. =Trichocentrum 

fuscum 

Trichopilia fragrans Auctt., non (Lindl.) Rchb.f. =Trichopilia 

wageneri 

Trichopilia mutica Rchb.f. & Wullschl. =Leucohyle mutica 

Trichosalpinx blaisdellii Auctt., non (Wats.) Luer 

=Trichosalpinx egleri 

Trigonidium tenue Lodd. ex Lindl. =Trigonidium acuminatum 

Trisetella huebneri (Schlechter) Luer =Trisetella triglochin 

Vanilla aromatica Swartz =Vanilla mexicana 

Vanilla ensifolia Rolfe =Vanilla odorata 

Vanilla inodora Schiede =Vanilla mexicana 

Vanilla pompona Auctt., non Schiede =Vanilla grandiflora 

Vanilla wrightii Rchb.f. =Vanilla bicolor 

Wullschlaegelia aphylla (Swartz) Rchb.f. [not in Guianas] 

Wullschlaegelia ulei Cogn. =Uleiorchis ulei 

Zygopetalum brachystalix Rchb.f. =Otostylis lepida 

Zygopetalum burkei Rchb.f. =Galeottia burkei 

Zygopetalum kegelii Rchb.f. =Zygosepalum kegelii 

Zygopetalum rostratum Hook. =Zygosepalum labiosum 

Zygopetalum tricolor Lindl. =Koellensteinia tricolor 

OXALIDACEAE 



REFERENCES: 

SPERMATOPHYTA: Orchidaceae - Papaveraceae 

163 

Hammel, B.E. & N.A. Zamora. 1993. 

Ruptiliocarpon (Lepidobotryaceae): a new 

arborescent genus and tropical American link 

to Africa, with a reconsideration of the family. 

Novon 3(4): 408-417. 

Jonker, F.P. 1942. Oxalidaceae. In A. Pulle, 


Lourteig, A. 1980. Oxalidaceae. In R.E. 



67: 823-850. 

Lourteig, A. 1994. Oxalis L. subgénero 

Thamnoxys (Endl.) Reiche emend. Lourt. 

Bradea 7(1): 1-199. 

Oxalis barrelieri L. [GU SU FG] 

Oxalis corniculata L. [GU SU FG] [naturalized] 

Oxalis debilis Kunth var. corymbosa (DC.) Lourteig [GU] 

Oxalis frutescens L. [GU SU FG] 

Oxalis juruensis Diels [GU SU FG] 

Oxalis mucronulata Norlind [GU] 

Ruptiliocarpon sp. aff. caracolita Hammel & Zamora [SU] 

[referred to segregate family Lepidobotryaceae by 

Hammel & Zamora] 

OXALIDACEAE - SYNONYMS AND 


Acetosella schomburgkiana (Prog.) Kuntze =Oxalis frutescens 

Oxalis amazonica Prog. =Oxalis barrelieri 

Oxalis schomburgkiana Prog. =Oxalis frutescens 

Oxalis schomburgkiana Prog. var. lasiocarpa Prog. =Oxalis 

frutescens 

Oxalis schomburgkiana Prog. var. leiocarpa Prog. =Oxalis 

frutescens 

Oxalis sepium St. Hil. [not in Guianas] 

PAPAVERACEAE 

contributed by R.W. Kiger (1992) 

REFERENCES: 

Stoffers, A.L. 1979. Papaveraceae. In A.L. 


5(1): 331-334. 

Argemone mexicana L. [SU FG] [cultivated, naturalized]

164 


PASSIFLORACEAE 

reviewed by C. Feuillet (1992); reviewed by C. 

Feuillet (1996) 

REFERENCES: 

Escobar, L.K. 1994. Two new species and a key 

to Passiflora subg. Astrophea. Syst. Bot. 

19(2): 203-210. 

Feuillet, C. 1986. Deux Passifloraceae nouvelles 

et quelques espèces rares en Guyane 

française. Candollea 41: 173-178. 

Feuillet, C. 1989. Diversity and distribution on 

Guianan Passifloraceae. In L.B. Holm- 

Nielsen, I.C. Nielsen & H. Balslev, Tropical 

Forests, Botanical Dynamics, Speciation and 

Diversity. Academic Press, London. 

Feuillet, C. 1994. Two new species of Passiflora 

(Passifloraceae) from French Guiana. Novon 

4(3): 236-241. 

Feuillet, C. & G. Cremers. 1984. Passifloraceae 

nouvelles ou méconnues de Guyane française. 

Proc. Kon. Ned. Akad. Wet. ser. C, 87: 377- 

386. 

Killip, E.P. 1937. Passifloraceae. In A. Pulle, 


Killip, E.P. 1938. The American species of 

Passifloraceae. Publ. Field Mus. Nat. Hist., 

Bot. Ser., 19: 1-613. 

Ancistrothyrsus sp. cf. tessmannii Harms [GU FG] 

Dilkea johannesii Barb. Rodr. [FG] 

Dilkea wallisii Masters [FG] 

Mitostemma glaziovii Masters [GU] 

Mitostemma jenmanii Masters [GU] 

Passiflora acuminata DC. [GU FG] 

Passiflora amoena Escobar [FG] 

Passiflora auriculata Kunth [GU SU FG] 

Passiflora candida (Poepp. & Endl.) Masters [SU FG] 

Passiflora capparidifolia Killip [GU] 

Passiflora cardonae Killip [GU] 

Passiflora ceratocarpa F. Silveira [GU] 

Passiflora cirrhiflora Juss. [GU SU FG] 

Passiflora citrifolia (Juss.) Masters [FG] 

Passiflora coccinea Aubl. [GU SU FG] 

Passiflora coriacea Juss. [GU] 

Passiflora costata Masters [GU SU FG] 

Passiflora crenata Feuillet & Cremers [FG] 

Passiflora deficiens Mast. [GU] 

Passiflora exura Feuillet [FG] 

Passiflora fanchonae Feuillet [FG] 

Passiflora foetida L. var. foetida [GU SU FG] 

Passiflora foetida L. var. hispida (DC.) Killip ex Gleason [GU 

SU FG] 

Passiflora foetida L. var. moritziana (Planch.) Killip [SU FG] 

Passiflora fuchsiiflora Hemsley [GU SU] 

Passiflora garckei Masters [GU SU FG] 

Passiflora glandulosa Cav. [GU SU FG] 

Passiflora kawensis Feuillet [FG] 

Passiflora laurifolia L. [GU SU FG] 

Passiflora leptopoda Harms [GU SU] 

Passiflora longiracemosa Ducke [GU] 

SPERMATOPHYTA: Passifloraceae - Pedaliaceae 

Passiflora maguirei Killip [GU] 

Passiflora misera Kunth [GU SU FG] 

Passiflora nitida Kunth [GU SU FG] 

Passiflora ovata Martin ex DC. [FG] 

Passiflora pachyantha Killip [GU] 

Passiflora pedata L. [GU SU] 

Passiflora picturata Ker-Gawler [SU] 

Passiflora plumosa Feuillet & Cremers [FG] 

Passiflora quadrangularis L. [FG] [cultivated or escaped from 

cultivation] 

Passiflora quadriglandulosa Rodsch. [GU] 

Passiflora quelchii N.E. Brown [GU] 

Passiflora retipetala Mast. [GU SU] 

Passiflora riparia Mast. [GU] 

Passiflora rubra L. [FG] 

Passiflora rufostipulata Feuillet [FG] 

Passiflora sclerophylla Harms [GU] 

Passiflora securiclata Masters [GU] 

Passiflora serrato-digitata L. [GU SU FG] 

Passiflora spicata Mast. [GU] 

Passiflora spinosa (Poepp. & Endl.) Mast. [GU] 

Passiflora stipulata Aubl. [FG] 

Passiflora trialata Feuillet & MacDougal, sp. nov. ined. [FG] 

Passiflora tuberosa Jacq. [GU] 

Passiflora variolata Poepp. & Endl. [FG?] [records probably 

represent a new species] 

Passiflora vespertilio L. [GU SU FG] 

PASSIFLORACEAE - SYNONYMS AND 


Passiflora capsularis L. [not in Guianas] 

Passiflora lonchophora Harms =Passiflora retipetala 

Passiflora maliformis L. [not in Guianas] 

Passiflora murucuia L. [not in Guianas] 

Passiflora rotundifolia L. [not in Guianas] 

Passiflora serratifolia L. [not in Guianas] 

Passiflora suberosa L. [not in Guianas] 

REFERENCES: 

PEDALIACEAE 

Eseltine, G.P. van. 1929. A preliminary study of 

the unicorn plants (Martyniaceae). N.Y. State 

Ag. Expt. Sta. (Geneva) Tech. Bull. 149: 1- 

41. 

Pulle, A. 1932. Pedaliaceae. In A. Pulle, Flora 


Seegeler, C.J.P. 1989. Sesamum orientale L. 

(Pedaliaceae): sesame's correct name. Taxon 

38(4): 656-659. 

Craniolaria annua L. [GU] [to be expected in SU] 

Sesamum orientale L. [GU SU FG] [cultivated & escaping] 

PEDALIACEAE - SYNONYMS AND 

MISIDENTIFICATIONS:

Sesamum indicum L. =Sesamum orientale 

REFERENCES: 


PERIDISCACEAE 


of tropical America: LXIX. A new genus of 

Peridiscaceae. Kew Bull. 15: 467-471. 

Whittonia guianensis Sandw. [GU] 

PHYTOLACCACEAE 


REFERENCES: 

Burger, W. 1983. Phytolaccaceae. In W. 


(n.s.) 13: 199-212. 

Raeder, K. 1961. Phytolaccaceae. In R.E. 



48(1): 66-79. 

Rohwer, J. 1982. A taxonomic revision of the 

genera Seguieria Loefl. and Gallesia Casar. 

Mitt. Bot. Staats. München 18: 231-288. 

Stoffers, A.L. 1957. Phytolaccaceae. In A. Pulle 

& J. Lanjouw, Flora of Suriname 1(2): 209- 

217. 

Walter, H. 1909. Phytolaccaceae. In A. Engler, 

Das Pflanzenreich 4(83): 1-154. 

Microtea debilis Swartz [GU SU FG] 

Microtea maypurensis (Kunth) G. Don [GU SU] 

Petiveria alliacea L. [GU SU FG] 

Phytolacca rivinoides Kunth & Bouché [GU SU FG] 

Phytolacca thyrsiflora Fenzl ex Schmidt [GU FG] 

Rivina humilis L. [GU SU FG] 

Seguieria americana L. [GU SU FG] 

Seguieria macrophylla Benth. [GU FG?] 

Trichostigma octandra (L.) H. Walter [GU FG?] 

PHYTOLACCACEAE - SYNONYMS AND 


Phytolacca americana L. [not in Guianas] 

Seguieria foliosa Benth. =Seguieria americana 

PINACEAE 


REFERENCES: 

Stevenson, D. & T. Zanoni. 1991. Pinaceae. In 


(series A) 9: 19-22. 

Pinus caribaea Morelet [GU SU FG] [cultivated, naturalized] 

PIPERACEAE 

reviewed by A.R.A. Görts-van Rijn (1996) 

REFERENCES: 

SPERMATOPHYTA: Pedaliaceae - Piperaceae 

165 

Kramer, K.U. & A.R.A. Görts-van Rijn. 1968. 

Piperaceae (additions and corrections). In A. 


415-421. 

Steyermark, J.A. 1984. Piperaceae. Flora de 

Venezuela II(2): 1-619. 

Tebbs, M.C. 1989. Revision of Piper in the New 

World. 1. Review of characters and 

taxonomy of Piper section Macrostachys. 

Bull. Brit. Mus. Nat. Hist. (Bot.) 19: 117-158. 


World. 2. The taxonomy of Piper section 

Churumayu. Bull. Brit. Mus. Nat. Hist. (Bot.) 

20(2): 193-236. 


World. 3. The taxonomy of Piper sections 

Lepianthes and Radula. Bull. Nat. Hist. Mus. 

23(1): 1-50. 

Trelease, W. & T.G. Yuncker. 1950. The 

Piperaceae of Northern South America. 

Urbana, Illinois: University of Illinois Press. 

Yuncker, T.G. 1957. Piperaceae. In A. Pulle & 

J. Lanjouw, Flora of Suriname 1(2): 218-290. 

Yuncker, T.G. 1972. The Piperaceae of Brazil. 

Hoehnea 2: 19-366. 

Peperomia acuminata Ruiz & Pav. [GU?] [Mt. Roraima] 

Peperomia alata Ruiz & Pav. [GU SU FG?] 

Peperomia angustata Kunth [GU FG] 

Peperomia blanda (Jacq.) Kunth [GU? SU] 

Peperomia circinnata Link [SU?] 

Peperomia delascioi Steyerm. [GU?] [Mt. Roraima] 

Peperomia distachya (L.) A. Dietr. [SU FG] [to be expected in 

GU] 

Peperomia duidana Trel. ex Gleason [GU] 

Peperomia elongata Kunth var. elongata [GU SU FG] 

Peperomia elongata Kunth var. guianensis Yunck. [GU FG] 

Peperomia elongata Kunth var. piliramea Trel. & Yunck. [GU 

SU] 

Peperomia emarginella (Swartz) C. DC. [GU FG] 

Peperomia galioides Kunth [Mt. Roraima] 

Peperomia glabella (Swartz) A. Dietr. var. glabella [GU SU 

FG] 

Peperomia glabella (Swartz) A. Dietr. var. nervulosa (C. DC.) 

Yunck. [GU SU FG] 

Peperomia lancifolia Hook. [GU] 

Peperomia longemucronata C.DC. [Mt. Roraima]

166 

SPERMATOPHYTA: Piperaceae 


Peperomia macrostachya (Vahl) A. Dietr. var. macrostachya 

[GU SU FG] 

Peperomia macrostachya (Vahl) A. Dietr. var. nematostachya 

(Link) Trel. & Yunck. [GU FG] 

Peperomia magnoliifolia (Jacq.) A. Dietr. [GU SU] 

Peperomia maguirei Yunck. [SU FG] 

Peperomia obtusifolia (L.) A. Dietr. var. obtusifolia [GU SU 

FG] 

Peperomia ouabianae C. DC. var. ouabianae [GU SU FG] 

Peperomia pellucida (L.) Kunth [GU SU FG] 

Peperomia pernambucensis Miq. [GU FG] 

Peperomia purpurinervis C.DC. [GU?] [Mt. Roraima] 

Peperomia quadrangularis (Thompson) A. Dietr. [GU SU FG] 

Peperomia rotundata Kunth var. trinervula (C.DC.) Steyerm. 

[Mt. Roraima] 

Peperomia rotundifolia (L.) Kunth var. rotundifolia [GU SU 

FG] 

Peperomia schomburgkii C. DC. [GU] 

Peperomia serpens (Swartz) J.C. Loudon [GU SU FG] 

Peperomia tenella (Swartz) A. Dietr. var. tenella [GU?] [Mt. 

Roraima] 

Peperomia tetraphylla (G. Forst.) Hook. & Arn. [GU?] 

Peperomia transparens Miq. [SU FG] 

Piper adenandrum (Miq.) C. DC. [GU SU FG] 

Piper aduncum L. [GU SU FG] 

Piper aequale Vahl [GU SU FG] 

Piper alatabaccum Trel. & Yunck. [GU SU FG] 

Piper amalago L. var. medium (Jacq.) Yunck. [GU SU] 

Piper amapense Yunck. [GU? FG] 

Piper amplectenticaule Trel. & Yunck. [FG] 

Piper amplum Kunth [FG] 

Piper angustifolium Lam. [FG] 

Piper anonifolium (Kunth) C. DC. var. anonifolium [GU SU 

FG] 

Piper anonifolium (Kunth) C. DC. var. parkerianum (C. DC.) 

Steyerm. [GU] 

Piper arboreum Aubl. var. arboreum [GU SU FG] [subsp. 

arboreum] 

Piper arboreum Aubl. var. hirtellum Yunck. [GU SU FG] 

[subsp. arboreum] 

Piper arboreum Aubl. subsp. tuberculatum (Jacq.) Tebbs [GU 

SU FG] 

Piper augustum Rudge var. augustum [GU SU FG] 

Piper augustum Rudge var. pubinerve Trel. & Yunck. [GU 

FG] 

Piper auritum Kunth [SU FG] 

Piper avellanum (Miq.) C. DC. var. angustifolium Trel. & 

Yunck. [SU] 

Piper avellanum (Miq.) C. DC. var. avellanum [GU SU FG] 

Piper bartlingianum (Miq.) C. DC. [GU SU FG] 

Piper bolivaranum Yunck. [GU] 

Piper brownsbergense Yunck. [SU FG] 

Piper cernuum Vell. var. cernuum [FG] 

Piper consanguineum Kunth [GU SU FG] 

Piper coruscans Kunth [GU FG] 

Piper crassinervium Kunth [FG] 

Piper cyrtopodon (Miq.) C. DC. [GU SU FG] 

Piper demeraranum (Miq.) C. DC. [GU SU FG] 

Piper dilatatum L.C. Rich. [GU SU FG] 

Piper divaricatum G. Mey. [GU SU FG] 

Piper dumosum Rudge [GU SU FG] 

Piper eucalyptifolium Rudge [GU FG] 

Piper fanshawei Yunck. [GU] 

Piper flexicaule (Miq.) C. DC. [GU] 

Piper foveolatum C. DC. [SU FG] [to be expected in GU] 

Piper fuligineum Kunth [SU FG] 

Piper gabrielianum (Miq.) C. DC. [FG] 

Piper glabrescens (Miq.) C. DC. var. caparonum (C. DC.) 

Yunck. [GU] 

Piper glabrescens (Miq.) C. DC. var. glabrescens [GU SU] 

Piper guianense (Klotzsch) C.DC. [GU FG?] 

Piper hirtilimbum Trel. & Yunck. [GU] 

Piper hispidum Swartz [GU SU FG] 

Piper hostmannianum (Miq.) C. DC. [GU SU FG] 

Piper humistratum Görts & K.U. Kramer [SU FG] 

Piper inaequale C. DC. [FG] 

Piper insigne (Kunth) C. DC. [GU SU FG] 

Piper kappleri C. DC. [GU SU FG] 

Piper kegelianum (Miq.) C. DC. [GU SU FG] 

Piper kegelii C. DC. [SU] 

Piper laevigatum Kunth [FG?] 

Piper marginatum Jacq. [GU SU FG] 

Piper microstachyum Vahl [FG] [=Piper angustifolium?] 

Piper modestum (Miq.) C. DC. [FG] [insufficiently known 

taxon] 

Piper nematanthera (Miq.) C. DC. var. nematanthera [GU SU 

FG] 

Piper nematanthera (Miq.) C. DC. var. sagotii (C. DC.) Trel. & 

Yunck. [FG] 

Piper nigrispicum C. DC. [FG] 

Piper nigrum L. [cultivated, perhaps escaping] 

Piper obliquum Ruiz & Pav. [GU SU FG] 

Piper oblongifolium (Klotzsch) C. DC. var. glabrum C. DC. 

[GU] 

Piper oblongifolium (Klotzsch) C. DC. var. oblongifolium 

[GU SU FG] 

Piper paramaribense C. DC. [SU FG] 

Piper pedunculare (Miq.) C. DC., comb. illeg. [GU] [taxon 

without valid combination in Piper] 

Piper perstipulare Steyerm. [GU] 

Piper piscatorum Trel. & Yunck. [GU FG] 

Piper pseudoglabrescens Trel. & Yunck. [GU] 

Piper pulleanum Yunck. [GU SU FG] 

Piper reticulatum L. [GU SU FG] 

Piper rudgeanum (Miq.) C. DC. [SU FG?] 

Piper rupununianum Trel. & Yunck. [GU] 

Piper salicifolium Vahl [SU] 

Piper schiedeanum Steud. [GU? SU?] 

Piper subciliatum C. DC. [FG] 

Piper tectoniifolium Kunth [FG] 

Piper trichoneuron (Miq.) C. DC. [GU SU FG] 

Piper wachenheimii Trel. [GU SU FG] 

Pothomorphe peltata (L.) Miq. [GU SU FG] [=Piper peltatum] 

[=Lepianthes peltata] 

PIPERACEAE - SYNONYMS AND 


Artanthe peduncularis Miq. =Piper pedunculare 

Artanthe schomburgkii Klotzsch =Piper pedunculare 

Lepianthes peltata (L.) Raf. =Pothomorphe peltata 

Peperomia angulata Kunth =Peperomia quadrangularis 

Peperomia caulibarbis Miq. =Peperomia glabella var. glabella 

Peperomia glabella (Swartz) A. Dietr. var. melanostigma 

(Miq.) Dahlst. =Peperomia glabella var. nervulosa 

Peperomia lanjouwii Yunck. =Peperomia rotundifolia 

Peperomia longifolia C.DC. =Peperomia pernambucensis 

Peperomia myosuroides A. Dietr. =Peperomia macrostachya 

var. macrostachya


Peperomia myriocarpa Miq. =Peperomia elongata var.elongata 

Peperomia nematostachya Link =Peperomia macrostachya var. 

nematostachya 

Peperomia nummularifolia Kunth =Peperomia rotundifolia 

var. rotundifolia 

Peperomia paniculata Regel =Peperomia pernambucensis 

Peperomia parkeriana Miq. =Peperomia macrostachya var. 

macrostachya 

Peperomia reflexa (L.f.) A. Dietr., nom illeg., non Kunth 

=Peperomia tetraphylla 

Peperomia surinamensis C. DC. =Peperomia elongata var. 

elongata 

Peperomia tafelbergensis Yunck. =Peperomia ouabianae var. 

ouabianae 

Peperomia trifolia (L.) A. Dietr. [not in Guianas] 

Piper acarouanyanum C. DC. =Piper avellanum 

Piper adenophorum C. DC. =Piper dumosum 

Piper affine C. DC. =Piper paramaribense 

Piper amplectens (Miq.) C. DC. =Piper fuligineum 

Piper angremondii DC. =Piper hispidum 

Piper asperifolium Rich. =Piper hispidum 

Piper benoistii Trel. ex Lemée [apparently a nomen nudum] 

Piper brachystachyum C. DC. =Piper eucalyptifolium 

Piper caracasanum Bredem. =Piper obliquum 

Piper carniconnectivum C. DC. [not in Guianas] 

Piper citrifolium Lam. =Piper anonifolium var. annonifolium 

Piper citrifolium Lam. var. parkerianum (C. DC.) Trel. & 

Yunck. =Piper anonifolium var. parkerianum 

Piper colubrinum Link =Piper divaricatum 

Piper corylifolium Kunth =Piper dilatatum 

Piper crassum Vell. =Piper divaricatum 

Piper demeraranum (Miq.) C. DC. var. rubescens (C. DC.) 

Trel. & Yunck. =Piper demeraranum 

Piper deminutum Yunck. =Piper bolivaranum 

Piper diospyrifolium Kunth [not in Guianas] 

Piper flexuosum Rudge =Piper flexicaule 

Piper fulgidum Yunck. [not in Guianas] 

Piper geayi Trel. ex Lemée [apparently a nomen nudum] 

Piper geniculatum Sw. =Piper arboreum 

Piper glabellum (Miq.) C. DC. =Piper divaricatum 

Piper gleasonii Yunck. =Piper trichoneuron 

Piper godebertianum Trel. ex Lemée [apparently a nomen 

nudum] 

Piper hirsutum Swartz =Piper hispidum 

Piper hohenackeri C. DC. =Piper anonifolium 

Piper insipiens Trel. & Yunck. =Piper divaricatum 

Piper lapathifolium Steud. [not in Guianas] 

Piper lenormandianum C. DC. =Piper kappleri 

Piper leprieurii Pulle =Piper consanguineum 

Piper maguirei Yunck. =Piper paramaribense, syn. nov. ined. 

Piper marowinianum C. DC. =Piper kappleri 

Piper marowynense C. DC. =Piper avellanum 

Piper medium Jacq. =Piper amalago var. medium 

Piper miquelii C. DC. =Piper eucalyptifolium, syn. nov. ined. 

Piper mollipilosum C. DC. [not in Guianas] 

Piper moraballianum Trel. =Piper avellanum 

Piper nitidum Vahl =Piper divaricatum 

Piper orenocanum C. DC. =Piper coruscans [=Piper coruscans 

var. membranaceum (C.DC.) Steyerm.?] 

Piper orenocanum C. DC. var. koriaboense Trel. & Yunck. 

=Piper coruscans 

Piper parakegelianum Sagot ex Lemée [apparently a nomen 

nudum] 

Piper parkerianum (Miq.) C. DC. =Piper anonifolium var. 

parkerianum 

SPERMATOPHYTA: Piperaceae - Poaceae 

167 

Piper peltatum L. =Pothomorphe peltata 

Piper piresii Yunck. [not in Guianas] 

Piper poiteanum Kunth =Piper foveolatum 

Piper pseudomarowynense Trel. ex Lemée [apparently a 

nomen nudum] 

Piper purpurascens Desf. 

Piper rhododendrifolium Kunth =Piper anonifolium 

Piper rhynchostachyum C. DC. =Piper eucalyptifolium 

Piper romboutsii Yunck. =Piper divaricatum, syn. nov. ined. 

Piper rubescens C.DC. =Piper demeraranum var. rubescens 

Piper rugosum Vahl =Piper anonifolium 

Piper sagoti C. DC. =Piper nematanthera var. sagotii 

Piper saramaccanum Yunck. =Piper obliquum 

Piper scabrum Swartz =Piper hispidum 

Piper submelanostictum C. DC. var. amelanostictum Yunck. 

=Piper obliquum 

Piper submelanostictum C. DC. var. submelanostictum =Piper 

obliquum 

Piper tiliifolium W. Hamilton =Bagassa guianensis 

(Moraceae) 

Piper trigonum C. DC. [not in Guianas] 

Piper tuberculatum Jacq. =Piper arboreum subsp. tuberculatum 

Piper warakabacoura (Miq.) C. DC. =Piper bartlingianum 

Quebitea guianensis Aubl. =Piper humistratum 

REFERENCES: 

PLANTAGINACEAE 

D'Arcy, W.G. 1971. Plantaginaceae. In R.E. 



58(3): 363-369. 

Lanjouw, J. 1938. Plantaginaceae. In A. Pulle, 


Plantago lanceolata L. [FG] [introduced] 

Plantago major L. [GU SU FG] [introduced] 

PLUMBAGINACEAE 

contributed by J.L. Luteyn (1992) 

Plumbago auriculata Lam. [widely cultivated, perhaps 

escaping] 

Plumbago scandens L. [GU? SU FG] 

POACEAE 

reviewed by E.J. Judziewicz (1992); additions 

and corrections by E.J. Judziewicz (1996)

168 

REFERENCES: 

SPERMATOPHYTA: Poaceae 


[for a more extensive bibliography, see 

Judziewicz 1990] 

Amshoff, G.J.H. & J.T. Henrard. 1943. 

Gramineae. In A. Pulle, Flora of Suriname 

1(1): 273-442. 



française III - les Cyperaceae et les Poaceae. 

Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Sér. 

Patr. Génétique) 11: 1-213. 

Hitchcock, A.S. 1922. Grasses of British 

Guiana. Contr. U.S. Natl. Herb. 22: 443-514. 

Judziewicz, E. 1990. Poaceae. In A.R.A. Görtsvan 


725. 


Gramineae (additions and corrections). In A. 


343-375. 

Swallen, J. 1948. Gramineae. In B. Maguire et 

al., Plant explorations in Guiana in 1944, 

chiefly to the Tafelberg and the Kaieteur 

Plateau - I. Bull. Torrey Bot. Club 75: 81-91. 

Acroceras macrum Stapf [GU SU] [naturalized] 

Acroceras zizanioides (Kunth) Dandy [GU SU FG] 

[naturalized] 

Aegopodon cenchroides Humb. & Bonpl. ex Willd. [Mt. 

Roraima] 

Andropogon angustatus (J. Presl) Steud. [GU SU] 

Andropogon bicornis L. [GU SU FG] 

Andropogon fastigiatus Swartz [GU] 

Andropogon gayanus Kunth [SU] [cultivated, naturalized] 

Andropogon glomeratus (Well.) BSP [GU?] 

Andropogon leucostachyus Kunth [GU SU FG] 

Andropogon selloanus (Hack.) Hack. [GU SU FG] 

Andropogon virgatus Desv. [GU SU FG] 

Anthephora hermaphrodita (L.) Kuntze [FG] 

Arberella flaccida (Doell) Soderstrom & Calderón [SU FG] 

Aristida capillacea Lam. [GU SU FG] 

Aristida longifolia Trin. [GU] 

Aristida recurvata Kunth [GU SU] 

Aristida riparia Trin. [SU] 

Aristida setifolia Kunth [GU SU] 

Aristida torta (Nees) Kunth [GU SU FG] 

Arthrostylidium scandens McClure [GU SU] 

Arthrostylidium venezuelae (Steud.) McClure [GU] 

Arundinella hispida (Willd.) Kuntze [GU SU FG] 

Arundo donax L. [SU FG?] [cultivated and naturalized] 

Aulonemia deflexa (N.E. Brown) McClure [GU] 

Aulonemia nitida Judziewicz, sp. nov. ined. [GU] 

Axonopus anceps (Mez) Hitchc. [GU SU] 

Axonopus aureus P. Beauv. [GU SU FG] 

Axonopus canescens (Nees) Pilg. [GU SU FG] 

Axonopus capillaris (Lam.) Chase [GU SU FG] 

Axonopus caulescens (Mez) Henr. [GU] 

Axonopus chrysoblepharis (Lag.) Chase [FG] 

Axonopus chrysostachyus (Schrad.) Pilg. [SU] 

Axonopus compressus (Swartz) P. Beauv. [GU SU FG] 

Axonopus eminens (Nees) G.A. Black [SU] 

Axonopus equitans Hitchc. & Chase [GU SU FG] 

Axonopus fissifolius (Raddi) Kuhlm. [GU] 

Axonopus flabelliformis Swallen [GU SU FG] 

Axonopus leptostachyus (Flüggé) Hitchc. [GU SU FG] 

Axonopus longispicus (Doell) Kuhlm. [GU SU FG] 

Axonopus oiapocensis G.A. Black [FG] 

Axonopus passourae G.A. Black [FG] 

Axonopus pubivaginatus Henr. [GU SU FG] 

Axonopus purpusii (Mez) Chase [GU SU FG] 

Axonopus ramosus Swallen [SU FG] [to be expected in GU] 

Axonopus senescens (Doell) Henr. [FG] 

Axonopus surinamensis (Steud.) Henr. [GU SU FG] 

Bothriochloa bladhii (Retz.) S.T. Blake [GU SU] [naturalized] 

Bouteloua americana (L.) Lams.-Scribn. [GU?] [introduced? 

no recent collections] 

Brachiaria fasciculata (Swartz) Parodi [GU SU FG] 

Brachiaria mollis (Swartz) Parodi [GU FG] 

Brachiaria mutica (Forssk.) Stapf [GU SU FG] [naturalized] 

Brachiaria plantaginea (Link) Hitchc. [FG] 

Cenchrus brownii Roem. & Schult. [GU SU FG] 

Cenchrus echinatus L. [GU SU FG] 

Chloris barbata Swartz [GU SU FG] 

Chloris dandyana C.D. Adams [GU] 

Chloris radiata (L.) Swartz [GU SU FG] 

Chusquea linearis N.E. Brown [GU] 

Coeloachis aurita (Steud.) A. Camus [GU SU FG] 

Coix lacryma-jobi L. [GU SU FG] [cultivated, naturalized] 

Cortaderia roraimensis (N.E. Brown) Pilg. [GU] 

Cryptochloa capillata (Trin.) Soderstrom [FG] 

Ctenium cirrosum (Nees) Kunth [SU] 

Cymbopogon citratus (DC.) Stapf [GU SU FG] [cultivated, 

naturalized] 

Cynodon dactylon (L.) Pers. [GU SU FG] 

Dactyloctenium aegyptium (L.) Willd. [GU SU FG] 

Dichanthium annulatum (Forssk.) Stapf [GU] [cultivated, 

naturalized] 

Dichanthium aristatum (Poir.) Hubb. [GU] [cultivated, 

naturalized] 

Digitaria ciliaris (Retz.) Koeler [GU SU FG] 

Digitaria decumbens Stent [GU SU] [cultivated, naturalized] 

Digitaria fragilis (Steud.) Luces [GU] 

Digitaria fuscescens (J. Presl) Henr. [GU SU FG] [naturalized] 

Digitaria horizontalis Willd. [GU SU FG] 

Digitaria insularis (L.) Mez ex Ekman [GU SU FG] 

Digitaria lanuginosa (Nees) Henr. [GU FG] 

Digitaria laxa (A. Spreng.) Parodi [GU SU FG] 

Digitaria longiflora (Retz) Pers. [GU SU] [naturalized?] 

Digitaria nuda Schumach. [GU FG] [naturalized?] 

Digitaria setigera Roth ex Roem. & Schult. [GU SU FG] 

[naturalized] 

Digitaria swazilandensis Stent [SU] [cultivated, naturalized] 

Digitaria tenuis (Nees) Henr. [GU] 

Digitaria venezuelae Henr. [SU FG] 

Digitaria violascens Link [GU SU FG] [cultivated, 

naturalized] 

Echinochloa colona (L.) Link [GU SU FG] 

Echinochloa crus-pavonis (Kunth) Schult. [GU SU FG] 

Echinochloa polystachya (Kunth) Hitchc. [GU SU FG] 

Echinochloa pyramidalis (Lam.) Hitchc. & Chase [FG] 


Echinolaena inflexa (Poir.) Chase [GU SU FG] 

Eleusine indica (L.) Gaertn. [GU SU FG] [naturalized] 

Elyonurus muticus (Spreng.) Kuntze [GU SU FG] 

Eragrostis acutiflora (Kunth) Nees [GU FG] 

Eragrostis airoides Nees [to be expected in GU]


Eragrostis amabilis (L.) Wight & Arnott [GU SU FG] 

[naturalized] 

Eragrostis ciliaris (L.) R. Brown [GU SU FG] [naturalized] 

Eragrostis guianensis Hitchc. [GU] 

Eragrostis hypnoides (Lam.) Britton, Sterns, & Poggenb. [GU 

SU FG] 

Eragrostis japonica (Thunb.) Trin. [GU] [naturalized] 

Eragrostis maypurensis (Kunth) Steud. [GU SU FG] 

Eragrostis pilosa (L.) P. Beauv. [GU FG] [naturalized] 

Eragrostis plurinodis Swallen ex Luces [GU] 

Eragrostis polytricha Nees [GU] 

Eragrostis refracta (Elliott) Lams.-Scribn. [SU] 

Eragrostis riobrancensis Judziewicz & P. Peterson [GU] 

Eragrostis scaligera Salzm. ex Steud. [FG] 

Eragrostis secundiflora J. Presl [GU] [to be expected in SU] 

Eragrostis tephrosanthos Schult. [GU SU FG] 

Eragrostis unioloides (Retz.) Nees [GU SU FG] [naturalized] 

Eriochloa polystachya Kunth [GU SU FG] 

Eriochloa procera (Retz.) Hubb. [GU SU] [naturalized] 

Eriochloa punctata (L.) W. Hamilton [GU SU FG] 

Eriochrysis cayennensis P. Beauv. [GU SU FG] 

Froesiochloa boutelouoides G.A. Black [FG] 

Guadua angustifolia Kunth [GU] 

Guadua glomerata Munro [GU FG] 

Guadua latifolia (Humb. & Bonpl.) Kunth [GU SU FG] 

Guadua macrostachya Rupr. [FG] [to be expected in SU] 

Guadua paniculata Munro [to be expected in GU] 

Guadua superba Huber [GU SU] 

Gymnopogon foliosus (Willd.) Nees [GU SU FG] 

Gymnopogon spicatus (Spreng.) Kuntze [GU] 

Gynerium sagittatum (Aubl.) P. Beauv. [GU SU FG] 

Hackelochloa granularis (L.) Kuntze [GU SU FG] 

[naturalized] 

Hemarthria altissima (Poir.) Stapf & Hubb. [GU FG] 

[naturalized] 

Heteropogon contortus (L.) P. Beauv. ex Roem. & Schult. 

[GU] 

Heteropogon melanocarpus (Elliott) Elliott ex Benth. [FG] [to 


Homolepis aturensis (Kunth) Chase [FG] [naturalized?] 

Homolepis glutinosa (Swartz) Zuloaga & Soderstrom [GU] 

Homolepis isocalycia (G. Mey.) Chase [GU SU FG] 

Hymenachne amplexicaulis (Rudge) Nees [GU SU FG] 

Hymenachne donacifolia (Raddi) Chase [GU SU FG] 

Hyparrhenia bracteata (Willd.) Stapf [GU SU] 

Hyparrhenia rufa (Nees) Stapf [GU SU FG] [cultivated, 

naturalized] 

Ichnanthus breviscrobs Doell [GU SU FG] 

Ichnanthus calvescens (Nees) Doell [GU SU] 

Ichnanthus dasycoleus Tutin [GU SU] 

Ichnanthus hoffmannseggii (Roem. & Schult.) Doell [FG] 

Ichnanthus leiocarpus (Spreng.) Kunth [GU SU FG] 

Ichnanthus nemoralis (Schrad.) Hitchc. & Chase [GU SU FG] 

Ichnanthus pallens (Swartz) Munro ex Benth. [GU SU FG] 

Ichnanthus panicoides P. Beauv. [GU SU FG] 

Ichnanthus ruprechtii Doell [GU SU FG] 

Ichnanthus tenuis (J. Presl) Hitchc. & Chase [GU SU FG] 

Imperata brasiliensis Trin. [GU SU FG] 

Imperata contracta (Kunth) Hitchc. [GU SU FG] 

Isachne ligulata Swallen [GU] 

Isachne polygonoides (Lam.) Doell [GU SU FG] 

Ischaemum guianense Kunth ex Hack. [GU SU FG] 

Ischaemum indicum (Houtt.) Merrill [GU SU] [cultivated, 

naturalized] 

Ischaemum rugosum Salisb. [GU SU FG] [naturalized] 


169 

Ischaemum timorense Kunth [GU SU] [cultivated, naturalized] 

Ixophorus unisetus (J. Presl) Schlechtend. [cultivated but to be 


Lasiacis anomala Hitchc. [GU SU] 

Lasiacis ligulata Hitchc. & Chase [GU SU FG] 

Lasiacis nigra Davidse [GU] 

Lasiacis procerrima (Hack.) Hitchc. [GU] 

Lasiacis sorghoidea (Desv.) Hitchc. & Chase [GU SU FG] 

Leersia hexandra Swartz [GU SU FG] 

Leersia ligularis Trin. [to be expected in GU] 

Leptochloa filiformis (Lam.) P. Beauv. [GU] 

Leptochloa scabra Nees [GU SU FG] 

Leptochloa virgata (L.) P. Beauv. [GU SU FG] 

Leptocoryphium lanatum (Kunth) Nees [GU SU FG] 

Lithachne pauciflora (Swartz) P. Beauv. [FG] [to be expected 

in GU] 

Luziola bahiensis (Steud.) Hitchc. [GU SU FG] 

Luziola subintegra Swallen [GU SU FG] 

Melinis minutiflora P. Beauv. [GU SU] [cultivated, 

naturalized] 

Merostachys sp. aff. retrorsa McClure [GU] 

Mesosetum cayennense Steud. [SU FG] 

Mesosetum chaseae Luces [GU] 

Mesosetum loliiforme (Steud.) Chase [GU SU FG] 

Mesosetum rottboellioides (Kunth) Hitchc. [GU SU] 

Microchloa indica (L.f.) P. Beauv. [introduced species to be 


Myriocladus distantiflorus Swallen [GU] [other species of this 

genus to be expected as well] 

Neurolepis angusta Swallen [to be expected in GU] 

Olyra caudata Trin. [GU SU FG] 

Olyra ciliatifolia Raddi [GU FG] 

Olyra ecaudata Doell [GU SU FG] 

Olyra latifolia L. [GU SU FG] 

Olyra longifolia Kunth [GU SU FG] 

Olyra micrantha Kunth [GU SU FG] 

Olyra obliquifolia Steud. [SU FG] 

Oplismenus hirtellus (L.) P. Beauv. [GU SU FG] 

Orthoclada laxa (Rich.) P. Beauv. [GU SU FG] 

Oryza grandiglumis (Doell) Prodoehl [FG] 

Oryza latifolia Desv. [GU SU FG] 

Oryza rufipogon Griff. [GU SU FG] [naturalized] 

Oryza sativa L. [cultivated, occasionally escaping] 

Otachyrium succisum (Swallen) Sendulsky & Soderstrom [GU 

SU FG] 

Otachyrium versicolor (Doell) Henr. [GU] 

Panicum aequivaginatum Swallen [GU] 

Panicum altum Hitchc. & Chase [GU] 

Panicum arctum Swallen [SU FG] 

Panicum bergii Arechav. [GU] 

Panicum caricoides Nees [GU SU FG] 

Panicum cayennense Lam. [SU FG] 

Panicum chnoodes Trin. [GU] 

Panicum cordovense E. Fourn. [FG] [to be expected in GU] 

Panicum cyanescens Nees [GU SU FG] 

Panicum dichotomiflorum Michx. [GU SU FG] 

Panicum discrepans Doell [GU FG] 

Panicum elephantipes Nees [GU SU FG] 

Panicum eligulatum N.E. Brown [GU] 

Panicum grande Hitchc. & Chase [GU SU FG] 

Panicum granuliferum Kunth [GU] 

Panicum hirsutum Swartz [GU] 

Panicum hirtum Lam. [GU SU FG] 

Panicum hylaeicum Mez [GU SU FG] 

Panicum laxum Swartz [GU SU FG]

170 



Panicum maximum Jacq. [GU SU FG] [naturalized] 

Panicum mertensii Roth [GU SU FG] 

Panicum micranthum Kunth [GU SU FG] 

Panicum miliaceum L. [GU FG] [cultivated; naturalized?] 

Panicum millegrana Poir. [GU SU FG] 

Panicum nervosum Lam. [GU SU FG] 

Panicum olyroides Kunth [GU SU FG] 

Panicum pandum Swallen [GU] 

Panicum pantrichum Hack. [FG] [to be expected in GU] 

Panicum parvifolium Lam. [GU SU FG] 

Panicum pilosum Swartz [GU SU FG] 

Panicum polycomum Trin. [GU SU FG] 

Panicum polygonatum Schrad. [GU FG] 

Panicum pycnoclados Tutin [GU SU] 

Panicum pyrularium Hitchc. & Chase [SU FG] 

Panicum rivale Swallen [GU SU FG] 

Panicum rudgei Roem. & Schult. [GU SU FG] 

Panicum scabridum Doell [GU SU FG] 

Panicum schwackeanum Mez [GU SU FG] 

Panicum sciurotoides Zuloaga & Morrone [GU FG] 

Panicum sellowii Nees [GU] 

Panicum stenodes Griseb. [GU SU FG] 

Panicum stoloniferum Poir. [GU SU FG] 

Panicum telmatum Swallen [to be expected in GU] 

Panicum trichanthum Nees [GU SU FG] 

Panicum trichocladum Hack. ex K. Schum. [SU] [naturalized] 

Panicum trichoides Swartz [GU SU FG] 

Panicum vinaceum Swallen [to be expected in GU] 

Pappophorum pappiferum (Lam.) Kuntze [GU] [naturalized] 

Paratheria prostrata Griseb. [GU FG] 

Pariana campestris Aubl. [GU? SU FG] 

Pariana radiciflora Sagot ex Doell [GU SU FG] 

Parodiolyra lateralis (J. Presl) Soderstrom & Zuloaga [GU SU] 

Parodiolyra luetzelburgii (Pilg.) Soderstrom & Zuloaga [GU 

SU FG] 

Paspalidium geminatum (Forssk.) Stapf [GU SU FG] 

Paspalum albidulum Henr. [SU] 

Paspalum altsonii Chase [GU SU] 

Paspalum ammodes Trin. [GU] 

Paspalum arenarium Schrad. [GU SU FG] 

Paspalum arundinaceum Poir. [FG] 

Paspalum bifidifolium Soderstrom [GU] 

Paspalum boscianum Flüggé [GU SU FG] 

Paspalum carinatum Humb. & Bonpl. ex Flüggé [GU SU] 

Paspalum cinerascens (Doell) Burman & Bastos [SU FG] 

Paspalum clavuliferum C. Wright [GU] 

Paspalum conjugatum Berg. [GU SU FG] 

Paspalum conspersum Schrad. [GU SU] 

Paspalum convexum Humb. & Bonpl. ex Flüggé [GU SU FG] 

Paspalum corcovadense Raddi [GU] 

Paspalum coryphaeum Trin. [GU SU FG] 

Paspalum decumbens Swartz [GU SU FG] 

Paspalum delicatum Swallen [FG] 

Paspalum densum Poir. [GU SU FG] 

Paspalum dilatatum Poir. [GU SU] [cultivated, naturalized] 

Paspalum distichum L. [GU SU FG?] 

Paspalum fasciculatum Willd. [SU FG] [cultivated; 

naturalized] 

Paspalum fimbriatum Kunth [GU?] [introduced, no recent 

collections] 

Paspalum gardnerianum Nees [GU SU FG] 

Paspalum goeldii Swallen [GU SU FG] 

Paspalum hyalinum Nees [GU SU] 

Paspalum lacustre Swallen [records erroneous, but perhaps to 

be expected] 

Paspalum lanciflorum Trin. [GU SU] 

Paspalum laxum Lam. [FG] 

Paspalum lineare Trin. [GU] 

Paspalum loefgrenii Ekman [GU] 

Paspalum maculosum Trin. [GU SU] 

Paspalum maritimum Trin. [GU SU FG] 

Paspalum melanospermum Desv. ex Poir. [GU SU FG] 

Paspalum millegrana Schrad. [GU SU FG] 

Paspalum morichalense Davidse, Zuloaga & Filgueiras [SU] 

[specimens previously identified as Paspalum lacustre] 

Paspalum multicaule Poir. [GU SU FG] 

Paspalum multinervium A.G. Burman [FG] 

Paspalum notatum Flüggé [GU SU FG] 

Paspalum nudatum Luces [GU SU] 

Paspalum nutans Lam. [GU SU FG] 

Paspalum oligostachyum Salzm. ex Steud. [SU FG] 

Paspalum orbiculatum Poir. [GU SU FG] 

Paspalum paniculatum L. [GU SU FG] 

Paspalum parviflorum Rhode ex Flüggé [SU FG] 

Paspalum pectinatum Nees [GU SU] 

Paspalum petilum Chase [GU] 

Paspalum pictum Ekman [GU] 

Paspalum pilosum Lam. [GU SU FG] 

Paspalum plicatulum Michx. [GU SU FG] 

Paspalum pulchellum Kunth [GU SU FG] 

Paspalum pumilum Nees [GU SU FG] 

Paspalum repens Berg. [GU SU FG] 

Paspalum riparium Nees [GU SU FG] 

Paspalum scrobiculatum L. [SU FG] [cultivated; naturalized?] 

Paspalum serpentinum Hochst. [GU SU FG] 

Paspalum setiglume Chase [GU] 

Paspalum stellatum Humb. & Bonpl. [to be expected in GU] 

Paspalum subciliatum Chase [GU FG] 

Paspalum trinii Swallen [GU] 

Paspalum vaginatum Swartz [GU SU FG] 

Paspalum virgatum L. [GU SU FG] 

Paspalum wrightii Hitchc. & Chase [GU] 

Pennisetum ciliare (L.) Link [GU] [cultivated; naturalized?] 

Pennisetum polystachion (L.) Schult. [GU FG] 

Pennisetum purpureum Schumach. [GU FG] [cultivated, 

naturalized] 

Pharus lappulaceus Aubl. [GU SU FG] 

Pharus latifolius L. [GU SU FG] 

Pharus parvifolius Nash subsp. elongatus Judziewicz [GU] 

Pharus parvifolius Nash subsp. parvifolius [GU SU FG] 

Pharus virescens Doell [GU SU FG] 

Phragmites australis (Cav.) Trin. ex Steud. [GU SU FG] 

Piresia goeldii Swallen [SU FG] 

Piresia macrophylla Soderstrom [FG] 

Piresia sympodica (Doell) Swallen [GU SU FG] 

Pseudechinolaena polystachya (Kunth) Stapf [GU] 

Raddia guianensis (Brongn.) Hitchc. [FG] [to be expected in 

SU] 

Raddiella esenbeckii (Steud.) Calderón & Soderstrom [GU SU 

FG] 

Raddiella kaieteurana Soderstrom [GU SU] 

Raddiella potaroensis Soderstrom [GU] 

Rehia nervata (Swallen) Fijten [GU SU FG] 

Reimarochloa aberrans (Doell) Chase [FG] 

Reimarochloa acuta (Flüggé) Hitchc. [GU SU FG] 

Rhipidocladum racemiflorum (Steud.) McClure [GU? SU] 

Rhipidocladum sibilans Davidse, Judziewicz, & L.G. Clark 

[Mt. Roraima] 

Rhipidocladum sp. aff. maxonii (Hitchc.) McClure [GU] 

Rhynchelytrum repens (Willd.) Hubb. [GU] [naturalized]


Rhytachne gonzalezii Davidse [SU] 

Rhytachne guianensis (Hitchc.) W.D. Clayton [GU SU] 

Rottboellia cochinchinensis (Lour.) W.D. Clayton [SU FG] 

[naturalized] 

Saccharum asperum (Nees) Steud. [to be expected in GU] 

Saccharum trinii (Hack.) Renvoize [GU SU] 

Sacciolepis angustissima (Steud.) Kuhlm. [GU SU FG] 

Sacciolepis indica (L.) Chase [GU] [naturalized?] 

Sacciolepis myuros (Lam.) Chase [GU SU FG] 

Sacciolepis otachyrioides Judziewicz [GU] 

Sacciolepis striata (L.) Nash [GU SU FG] 

Sacciolepis vilvoides (Trin.) Chase [GU SU FG] 

Schizachyrium brevifolium (Swartz) Nees ex Büse [GU SU? 

FG] 

Schizachyrium condensatum (Kunth) Nees [GU SU FG] 

Schizachyrium maclaudii (Jacques-Félix) S.T. Blake [GU SU 

FG] 

Schizachyrium sanguineum (Retz.) Alston [GU SU FG] 

Schizachyrium tenerum Nees [GU] 

Setaria barbata (Lam.) Kunth [FG] [naturalized] 

Setaria italica (L.) P. Beauv. [FG] [naturalized?] 

Setaria magna Griseb. [SU FG] 

Setaria parviflora (Poir.) Kerguélen [GU SU FG] 

Setaria poiretiana (Schult.) Kunth [GU SU FG] 

Setaria setosa (Swartz) P. Beauv. [FG] [naturalized?] 

Setaria tenax (Rich.) Desv. [GU SU FG] 

Setaria vulpiseta (Lam.) Roem. & Schult. [GU SU FG] 

Sorghastrum setosum (Griseb.) Hitchc. [GU FG] 

Sorghastrum stipoides (Kunth) Nash [GU SU FG] 

Sorghum bicolor (L.) Moench [GU SU FG] [cultivated, 

sparingly naturalized] 

Sorghum halepense (L.) Pers. [GU FG] [cultivated, 

naturalized] 

Spartina alterniflora Loisel. [GU SU FG] 

Spheneria kegelii (K. Muell. berol.) Pilg. [GU SU] 

Sporobolus ciliatus J. Presl [GU SU FG] 

Sporobolus cubensis Hitchc. [GU SU FG] 

Sporobolus jacquemontii Kunth [GU SU FG] 

Sporobolus pyramidatus (Lam.) Hitchc. [SU FG] 

Sporobolus temomairemensis Judziewicz & P. Peterson [SU] 

Sporobolus tenuissimus (Schrank) Kuntze [GU SU FG] 

Sporobolus virginicus (L.) Kunth [GU SU FG] 

Steirachne barbata (Trin.) Renvoize [GU] 

Stenotaphrum secundatum (Walt.) Kuntze [GU SU FG] 

Streptochaeta spicata Schrad. ex Nees [to be expected in GU] 

Streptogyna americana Hubb. [GU SU FG] 

Streptostachys asperifolia Desv. [GU SU FG] 

Thrasya achlysophila Soderstrom [GU] 

Thrasya axillaris (Swallen) A.G. Burman [SU] [to be expected 

in GU] 

Thrasya granitica A.G. Burman [SU] 

Thrasya petrosa (Trin.) Chase [GU SU] 

Thrasya robusta Hitchc. & Chase [SU FG] 

Thrasya scandens (Tutin) Soderstrom ex A.G. Burm. [GU SU 

FG] 

Thrasya trinitensis Mez [GU] 

Trachypogon spicatus (L.f.) Kuntze [GU SU FG] 

Tridens flaccidus (Doell) Parodi [GU] 

Tripogon spicatus (Nees) Ekman [to be expected in GU] 

Tripsacum andersonii J.R. Gray [GU SU FG] [introduced] 

Tripsacum australe Cutler & E.S. Anderson [GU SU FG] 

Tripsacum dactyloides (L.) L. [GU SU FG] 

Urochloa reptans (L.) Stapf [GU FG] [naturalized] 

Vetiveria zizanioides (L.) Nash [GU SU FG] [cultivated; 

naturalized?] 


171 

Zea luxurians (E.A. Durand & Aschers.) Bird [FG] [cultivated 

or escaped from cultivation] 

POACEAE - SYNONYMS AND 


Agrostis littoralis Lam. =Sporobolus virginicus 

Agrostis villosa Rchb. ex A. Spreng. =Sporobolus ciliatus 

Anatherum zizanioides (L.) Hitchc. & Chase =Vetiveria 

zizanioides 

Andropogon brevifolius Sw. =Schizachyrium brevifolium 

Andropogon dactyloides Steud. =Trachypogon spicatus 

Andropogon nodosus (Willemet) Nash =Dichanthium 

aristatum 

Andropogon sorghum (L.) Brot. =Sorghum bicolor 

Andropogon squarrosus non L.f., sensu Pulle =Vetiveria 

zizanioides 

Apluda zeugites non L., sensu Aublet =Ichnanthus pallens 

Aristida elegans Rudge =Aristida capillacea 

Aristida tincta Trin. & Rupr. =Aristida torta 

Arthrostylidium capillifolium Griseb. =Arthrostylidium 

farctum [West Indian species, not in Guianas] 

Arthrostylidium farctum (Aubl.) Soderstrom & Lourteig [West 

Indian species, not in Guianas] 

Arthrostylidium schomburgkii (Benn.) Munro [not in Guianas] 

Arunda farcta Aubl. =Arthrostylidium farctum [West Indian 

species, not in Guianas] 

Arundinaria deflexa N.E. Brown =Aulonemia deflexa 

Arundinaria schomburgkii Benn. =Arthrostylidium 

schomburgkii [not in Guianas] 

Arundo roraimensis N.E. Brown =Cortaderia roraimensis 

Axonopus "perpusillus Swallen" [invalid name, error for 

Axonopus purpurellus Swallen] 

Axonopus affinis non Chase, sensu Lindeman & Görts-van 

Rijn =Axonopus purpusii 

Axonopus attenuatus non (J. Presl) Hitchc., sensu Hitchcock 

=Axonopus flabelliformis 

Axonopus caracarahyensis Black & Fróes =Axonopus anceps 

Axonopus caulescens non (Mez) Henr., sensu Swallen 

=Axonopus ramosus 

Axonopus chrysites (Steud.) Kuhlm. =Axonopus aureus 

Axonopus erectus Swallen =Axonopus anceps 

Axonopus fissifolius non (Raddi) Kuhlm. sensu Amsh. & 

Henr. =Axonopus purpusii 

Axonopus flabelliformis Swallen var. camporum G.A. Black 


Axonopus flabelliformis Swallen var. decipiens G.A. Black 


Axonopus fockei (Mez) Henr. =Axonopus surinamensis 

Axonopus fockei non (Mez) Henr., sensu Amsh. & Henr. 

=Axonopus equitans 

Axonopus gentilis Henr. =Axonopus eminens 

Axonopus hitchcockii G.A. Black =Axonopus longispicus 

Axonopus kaietukensis Swallen =Axonopus flabelliformis 

Axonopus leptostachyus non (Flüggé) Hitchc. sensu 

Hitchcock =Axonopus longispicus 

Axonopus maguirei G.A. Black [not in Guianas] 

Axonopus pruinosus Henr. =Axonopus anceps 

Axonopus pulcher (Nees) Kuhlm. =Axonopus canescens 

Axonopus purpurellus Swallen =Axonopus flabelliformis 

Axonopus scoparius (Flüggé) Hitchc. sensu Hitchcock 

=Axonopus pubivaginatus 

Axonopus stragulus Chase =Axonopus fissifolius

172 



Axonopus stragulus non Chase, sensu Amsh. & Henr. 

=Axonopus ramosus 

Bambusa macrostachya (Rupr.) McClure =Guadua 

macrostachya 

Bambusa surinamensis Rupr. =Bambusa vulgaris Schrad. ex 

J.C. Wendl.[cultivated] 

Bothriochloa ischaemum non (L.) Keng, sensu Hitchcock 

=Bothriochloa bladhii 

Bothriochloa pertusa non (L.) Willd., sensu Hitchcock 

=Bothriochloa bladhii 

Brachiaria purpurascens (Raddi) Henr. =Brachiaria mollis 

Bulbulus nervatus Swallen =Rehia nervata 

Capriola dactylon (L.) Kuntze =Cynodon dactylon 

Cenchrus dactylolepis Steud. =Cenchrus brownii 

Cenchrus inflexus Poir. =Echinolaena inflexa 

Cenchrus marginalis Rudge =Echinolaena inflexa 

Cenchrus viridis Spreng. =Cenchrus brownii 

Chaetochloa geniculata (Lam.) Millsp. & Chase =Setaria 

parviflora 

Chaetochloa poiretiana (Schult.) Hitchc. =Setaria poiretiana 

Chaetochloa tenax (Rich.) Hitchc. =Setaria tenax 

Chloris inflata Link =Chloris barbata 

Chloris polydactyla (L.) Sw. =Chloris dandyana 

Cryptochloa concinna non (J.D. Hook.) Swallen, sensu 

Swallen =Piresia goeldii 

Cymbopogon bracteatus (Willd.) Hitchc. =Hyparrhenia 

bracteata 

Digitaria adscendens (Kunth) Henrard =Digitaria ciliaris 

Digitaria argillacea non (Hitchc. & Chase) Fern., sensu 

Swallen =Digitaria fragilis 

Digitaria malacophylla (Hitchc.) Henr. =Digitaria lanuginosa 

Digitaria microbachne (J. Presl) Henr. =Digitaria setigera 

Digitaria sanguinalis non (L.) Scop., sensu Lemée =Digitaria 

ciliaris 

Elyonurus adustus (Trin.) Ekman =Elyonurus muticus 

Eragrostis bahiensis non Schrad. ex Schult., sensu Lemée 

=Eragrostiss scaligera 

Eragrostis capillaris non (L.) Nees, sensu Lemée =Eragrostis 

pilosa 

Eragrostis fragilis Swallen =Eragrostis polytricha 

Eragrostis glomerata (Walt.) L.H. Dewey =Eragrostis japonica 

Eragrostis mexicana Auctt., non (Lag.) Link. =Eragrostis 

pilosa 

Eragrostis poaeoides non P. Beauv., sensu Pulle =Eragrostis 

tephrosanthos 

Eragrostis reptans sensu Auctt., non (Michx.) Nees 

=Eragrostis hypnoides 

Eragrostis rufescens Schrad. ex Schult. var. laurentiana 

Benoist =Eragrostis scaligera 

Erianthus saccharoides Michx. var. trinii Hack. =Saccharum 

trinii 

Erianthus trinii (Hack.) Hack. =Saccharum trinii 

Eriochloa kunthii G. Mey. =Eriochloa punctata 

Eriochloa ramosa (Retz.) Kuntze =Eriochloa procera 

Eriochloa subglabra (Nash) Hitchc. =Eriochloa polystachya 

Euchlaena luxurians E.A. Durand & Aschers. =Zea luxurians 

Guadua dioica Steud. =Guadua macrostachya 

Guadua tagoara non (Nees) Kunth, sensu Pulle =Guadua 

latifolia 

Helopus cognatus Steud. =Eriochloa punctata 

Helopus kunthii (G. Mey.) Trin. ex Steud. =Eriochloa punctata 

Holcus sorghum L. =Sorghum bicolor 

Holcus sorghum L. var. sudanensis (Piper) Hitchc. =Sorghum 

halapense 

Homalocenchrus hexandrus (Sw.) Kuntze =Leersia hexandra 

Hymenachne auriculata (Spreng.) Chase =Hymenachne 

donacifolia 

Hypogynium virgatum (Desv.) Dandy =Andropogon virgatus 

Ichnanthus acuminatus Swallen =Ichnanthus calvescens 

Ichnanthus amplus Swallen =Ichnanthus calvescens 

Ichnanthus areolatus K.E. Rogers =Ichnanthus pallens 

Ichnanthus axillaris (Nees) Hitchc. & Chase =Ichnanthus 

pallens 

Ichnanthus candicans non (Nees) Doell, sensu Lemée 

=Ichnanthus tenuis 

Ichnanthus candicans (Nees) Doell var. virescens Doell 

=Ichnanthus tenuis 

Ichnanthus confertus K.E. Rogers =Ichnanthus pallens 

Ichnanthus hitchcockii K.E. Rogers =Ichnanthus pallens 

Ichnanthus ichnodes (Griseb.) Hitchc. & Chase =Ichnanthus 

calvescens 

Ichnanthus nemorosus (Sw.) Doell [not in Guianas] 

Ichnanthus pallidus Swallen =Ichnanthus calvescens 

Ichnanthus petiolatus (Nees) Doell var. lanceolatus Doell 

=Ichnanthus nemoralis 

Ichnanthus riedelii non (Trin.) Doell., sensu Hitchcock 

=Ichnanthus nemoralis 

Imperata arundinacea Cirillo var. americana Anderss. 

=Imperata brasiliensis 

Imperata caudata (G. Mey.) Trin. =Imperata contracta 

Imperata flexuosa Swallen =Imperata contracta 

Ischaemum ciliare Retz. =Ischaemum indicum 

Ischaemum guianense Kunth ex Hack. var. schomburgkii 

Hack. =Ischaemum guianense 

Lasiacis divaricata non (L.) Hitchc., sensu Amshoff & Henrard 

=Lasiacis ligulata 

Lasiacis latifolium non L., sensu Pulle =Lasiacis ligulata 

Leptochloa domingensis (Jacq.) Trin. =Leptochloa virgata 

Leptochloa mutica Steud. =Leptochloa virgata 

Leptochloa virgata (L.) P. Beauv var. mutica (Steud.) Doell 

=Leptochloa virgata 

Leptostachys domingensis G. Mey. =Leptochloa virgata 

Leptostachys filiformis G. Mey. =Leptochloa filiformis 

Leptostachys virgata G. Mey. =Leptochloa virgata 

Luziola spruceana Auctt., non Benth. ex Doell =Luziola 

subintegra 

Manisuris granularis (L.) Naezén =Hackelochloa granularis 

Manisuris guianensis Hitchc. =Rhytachne guianensis 

Mesosetum elongatum Mez =Mesosetum loliiforme 

Mesosetum tenuifolium Swallen =Mesosetum loliiforme 

Olyra cordifolia Kunth =Olyra latifolia 

Olyra dimidiata Hochst. ex Steud. =Olyra caudata 

Olyra floribunda Raddi var. microphylla Doell =Raddia 

guianensis 

Olyra kegelii Mez =Olyra longifolia 

Olyra lateralis (J. Presl) Chase =Parodiolyra lateralis 

Olyra lateralis non (J. Presl) Chase, sensu Amshoff & Henrard 

=Parodiolyra luetzelburgii 

Olyra latifolia L. var. pubescens (Raddi) Doell =Olyra 

latifolia 

Olyra latifolia L. var. vestita Henr. =Olyra latifolia 

Olyra leutzelburgii Pilg. =Parodiolyra luetzelburgii 

Olyra nana Doell =Raddiella esenbeckii 

Olyra ovata Desv. =Ichnanthus panicoides 

Olyra pubescens Raddi =Olyra latifolia 

Olyra scabra Nees =Olyra latifolia 

Olyra speciosa Mez =Olyra caudata 

Olyra surinamensis Hochst. ex Steud. =Olyra latifolia 

Olyra surinamensis Auctt. non Hochst. ex Steud. =Olyra 

longifolia


Olyra sympodica Doell =Piresia macrophylla 

Oplismenus pseudo-undulatifolius (Roem. & Schult.) Kunth 

=Panicum stoloniferum 

Oplismenus velutinus (G. Mey.) Schult. =Oplismenus hirtellus 

Orthoclada rariflora (Lam.) P. Beauv. =Orthoclada laxa 

Oryza glumaepatula Hochst. ex Steud. =Oryza rufipogon 

Panicum albociliatum Swallen =Panicum pycnoclados 

Panicum altissimum G. Mey. =Panicum mertensii 

Panicum amplexicaule Rudge =Hymenachne amplexicaulis 

Panicum amplifolium Steud. =Setaria vulpiseta 

Panicum angulosum Swallen =Panicum rivale 

Panicum angustissimum Hochst. ex Steud. =Sacciolepis 

angustissima 

Panicum appressifolium Swallen =Panicum aequivaginatum 

Panicum appressum (Lam.) Doell =Paspalidium geminatum 

Panicum asperifolium (Desv.) Hitchc. =Streptostachys 

asperifolia 

Panicum axipilium Steud. =Panicum hirtum 

Panicum blackii Swallen =Panicum micranthum 

Panicum breviscrobs (Doell) Pilg. =Ichnanthus breviscrobs 

Panicum calvescens Nees =Ichnanthus calvescens 

Panicum caricoides non Nees, sensu Lemée =Panicum 

stenodes 

Panicum caricoides Nees var. pilosum Doell =Panicum 

caricoides 

Panicum caudatum Lam. =Setaria setosa 

Panicum cayennense Lam. var. curtatum Doell =Panicum 

cayennense 

Panicum cayennense Lam. var. divaricatum Doell =Panicum 

rudgei 

Panicum cayennense Lam. var. patulum Doell =Panicum 

cayennense 

Panicum chloroticum Nees =Panicum dichotomiflorum 

Panicum chrysites Steud. =Axonopus aureus 

Panicum coenosum Doell =Panicum pilosum 

Panicum colonum L. =Echinochloa colona 

Panicum commelinaefolium Rudge =Panicum nervosum 

Panicum compositum Auctt., non L. =Oplismenus hirtellus 

Panicum cordovense non E. Fourn., sensu Amshoff & Henrard 

=Panicum pycnoclados 

Panicum crus-galli L. var. aristatum G. Mey. =Echinochloa 

crus-pavonis 

Panicum cyanescens Nees var. lamarckianum Doell =Panicum 

cyanescens 

Panicum distichum Lam. var. luxurians G. Mey. =Panicum 

pilosum 

Panicum echinolaena Nees =Echinolaena inflexa 

Panicum elatior Kunth =Panicum mertensii 

Panicum errabundum Hitchc. =Panicum schwackeanum 

Panicum esenbeckii Nees ex Steud. =Raddiella esenbeckii 

Panicum fasciculatum Swartz =Brachiaria fasciculata 

Panicum fistulosum Hochst. ex Steud. =Panicum elephantipes 

Panicum frondescens G. Mey. =Panicum stoloniferum 

Panicum fusciflorum Steud. =Leptocoryphium lanatum 

Panicum glutinosum Sw. =Homolepis glutinosa 

Panicum graniticum Swallen =Panicum pandum 

Panicum guianense Hitchc. =Panicum hylaeicum 

Panicum hoffmannseggii Roem. & Schult. =Ichnanthus 

hoffmannseggii 

Panicum horizontale (Willd.) G. Mey. =Digitaria horizontalis 

Panicum imberbe Poir. =Setaria parviflora 

Panicum insulare (L.) G. Mey. =Digitaria insularis 

Panicum isocalycium G. Mey. =Homolepis isocalycia 

Panicum kaietukense Tutin =Panicum parvifolium 

Panicum kappleri Steud. =Panicum micranthum 


173 

Panicum leprieurii Steud. =Panicum stoloniferum 

Panicum leptachyrium Doell =Paratheria prostrata 

Panicum loliaceum Auctt., non Lam. =Oplismenus hirtellus 

Panicum loliiforme Hochst. ex Steud. =Mesosetum loliiforme 

Panicum longepaniculatum Benoist =Setaria poiretiana 

Panicum luticola Hitchc. =Panicum laxum 

Panicum macrostachyum (Kunth) Doell =Setaria vulpiseta 

Panicum magnum Hitchc. =Ichnanthus breviscrobs 

Panicum megiston Schult. =Panicum mertensii 

Panicum milleflorum Hitchc. & Chase =Panicum pilosum 

Panicum molle Swartz =Brachiaria mollis 

Panicum multiflorum non Poir., sensu Pulle =Panicum grande 

Panicum musei Steud. =Echinochloa colona 

Panicum myosurus Rich. =Sacciolepis myuros 

Panicum myrianthum Mez =Panicum grande 

Panicum myuros Lam. =Sacciolepis myuros 

Panicum numidianum non Lam., sensu Pulle =Brachiaria 

mutica 

Panicum pedunculare Willd. ex Steud. =Panicum cayennense 

Panicum perpusillum Benoist =Panicum polycomum 

Panicum pilcomayense Hack. =Panicum bergii 

Panicum pilisparsum G. Mey. =Panicum pilosum 

Panicum pilosissimum Roth =Panicum rudgei 

Panicum polygonoides Lam. =Isachne polygonoides 

Panicum prieurii Mez =Panicum scabridum 

Panicum proximum Steud. =Panicum mertensii 

Panicum pseudo-undulatifolium Roem. & Schult. =Panicum 

stoloniferum 

Panicum purpurascens Raddi =Brachiaria mutica 

Panicum racemiferum Steud. =Setaria poiretiana? 

Panicum ramuliflorum Hochst. ex Steud. =Panicum laxum 

Panicum recalvum non (Nees) Kunth, sensu Pulle =Digitaria 

laxa 

Panicum reptans L. =Urochloa reptans 

Panicum retroflexum Steud. =Setaria sp.? 

Panicum rugulosum Trin. var. subvelutinum Doell =Panicum 

millegrana 

Panicum sanguinale non L., sensu Pulle =Digitaria ciliaris 

Panicum savannarum Soderstrom =Panicum cyanescens 

Panicum sciurotis Auctt., non Trin. =Panicum sciurotoides 

Panicum scoparium Rudge =Panicum rudgei 

Panicum sellowii Nees var. subvelutinum (Doell) Henr. 

=Panicum millegrana 

Panicum setarioides Steud. =Echinochloa pyramidalis 

Panicum spectabile Nees =Echinochloa polystachya 

Panicum spissifolium Swallen =Panicum granuliferum 

Panicum stenodoides Hubb. =Panicum caricoides 

Panicum sulcatum Aubl. =Setaria palmifolia 

Panicum surinamense Hochst. ex Steud. =Axonopus 

surinamensis 

Panicum tatei Swallen =Panicum chnoodes 

Panicum tenax Rich. =Setaria tenax 

Panicum tenuiculmum G. Mey. =Panicum laxum 

Panicum tropidoblephare Tutin =Panicum chnoodes 

Panicum velutinum G. Mey. =Oplismenus hirtellus 

Panicum zizanioides Kunth =Acroceras zizanioides 

Pariana debilis Tutin =Pariana radiciflora 

Pariana gleasonii Hitchc. =Pariana radiciflora 

Pariana inaequalis Miq. =Pariana campestris 

Pariana longiflora Tutin =Pariana radiciflora 

Pariana lunata Nees =Pariana campestris 

Pariana maynensis Huber =Pariana radiciflora 

Pariana mollis Nees =Pariana campestris 

Pariana pulcherrima Tutin =Pariana radiciflora 

Pariana vulgaris Tutin =Pariana radiciflora

174 



Pariana zingiberina Doell =Pariana radiciflora 

Paspalum abrahamii Chase =Paspalum maritimum 

Paspalum arenicolum K. Muell. =Paspalum pulchellum 

Paspalum argyrocondylon Steud. =Paspalum serpentinum 

Paspalum axillare Swallen =Thrasya axillaris 

Paspalum bistipulatum Hochst. ex Steud. =Paspalum repens 

Paspalum caespitosum non Flüggé, sensu Pulle =Paspalum 

maritimum 

Paspalum caespitosum Steud. =Paspalum melanospermum 

Paspalum canum Sohns =Paspalum ammodes 

Paspalum capillare Lam. =Axonopus capillaris 

Paspalum carinatum Humb. & Bonpl. ex Flüggé var. kappleri 

(Steud.) Doell =Paspalum carinatum 

Paspalum caulescens Mez =Axonopus caulescens 

Paspalum ciliatum Lam. =Paspalum conjugatum 

Paspalum conjugatum Berg. var. pubescens Doell =Paspalum 

conjugatum 

Paspalum contractum Pilger =Paspalum lanciflorum 

Paspalum densiflorum non Doell, sensu Pulle =Paspalum 

maritimum 

Paspalum densum Poir. var. ciliatum Doell =Paspalum 

densum 

Paspalum echinotrichum Mez =Paspalum lanciflorum 

Paspalum elatum Rich. ex Doell =Paspalum arundinaceum 

Paspalum fockei Mez =Axonopus surinamensis 

Paspalum foveolatum Steud. =Paspalum melanospermum 

Paspalum froesianum Swallen =Paspalum loefgrenii 

Paspalum furcatum non Flüggé, sensu Pulle =Axonopus 

surinamensis 

Paspalum geniculatum Steud. =Paspalum orbiculatum 

Paspalum gnaphaloideum K. Müll. =Axonopus 

chrysostachyus 

Paspalum gracile Rudge =Paspalum repens 

Paspalum hirtigluma Steud. =Paspalum maritimum 

Paspalum humile Steud. =Paspalum melanospermum 

Paspalum kappleri Hochst. ex Steud. =Paspalum carinatum 

Paspalum kegelii K. Muell. =Spheneria kegelii 

Paspalum lacustre sensu Judziewicz, non Swallen =Paspalum 

morichalense 

Paspalum longebrachiatum Steud. =Paspalum arundinaceum 

Paspalum longissimum Hochst. ex Steud. =Paspalum 

conjugatum 

Paspalum lucidulum Swallen =Paspalum altsonii 

Paspalum maculosum Trin. var. rotundiflorum Doell 

=Paspalum serpentinum 

Paspalum montanum Henr. =Paspalum oligostachyum 

Paspalum orbiculatum Poir. var. lanuginosum Henr. 

=Paspalum orbiculatum 

Paspalum orbiculatum Poir. var. potaroense Chase =Paspalum 

orbiculatum 

Paspalum pectinatum non Nees, sensu Hitchcock =Paspalum 

lanciflorum 

Paspalum pedunculatum Poir. =Paspalum decumbens 

Paspalum platycaulon Poir. var. parviflorum Doell 

=Axonopus compressus 

Paspalum plicatulum Michx. var. microspermum Doell 

=Paspalum melanospermum 

Paspalum polychaetum Mez =Paspalum hyalinum 

Paspalum pulchrum Nees =Axonopus canescens 

Paspalum pusillum Venten. ex Flüggé =Paspalum orbiculatum 

Paspalum scandens Tutin =Thrasya scandens 

Paspalum senescens Doell =Axonopus senescens 

Paspalum setifolium Doell =Spheneria kegelii 

Paspalum vaginiflorum Steud. =Paspalum decumbens 

Paspalum wullschlaegelii Mez =Paspalum maritimum 

Pennisetum dasistachyum Desv. =Pennisetum polystachion 

Pennisetum richardii Kunth =Pennisetum polystachion 

Pennisetum setosum (Swartz) L. Rich. =Pennisetum 

polystachion 

Pharus glaber Kunth =Pharus lappulaceus 

Pharus glaber non Kunth, sensu Lemée =Pharus latifolius 

Pharus scaber Kunth var. pictus Doell =Pharus latifolius 

Phragmites communis Trin. =Phragmites australis 

Raddia capillata (Trin) Hitchc. =Cryptochloa capillata 

Raddia malmeana non (Ekman) Hitchc., sensu Hitchcock 

=Raddiella kaieteurana 

Raddia nana (Doell) Chase =Raddiella esenbeckii 

Raddia sympodica (Doell) Hitchc. =Piresia sympodica 

Raddiella maipuriensis Soderstrom =Raddiella potaroensis 

Raddiella malmeana non (Ekman) Swallen, sensu Hitchcock 

=Raddiella kaieteurana 

Raddiella nana (Doell) Swallen =Raddiella esenbeckii 

Raddiella truncata Swallen =Parodiolyra lateralis 

Reimarachloa brasiliensis (Spreng.) Hitchc. =Reimarachloa 

acuta 

Reimaria aberrans Doell =Reimarachloa aberrans 

Reimaria laxa Rchb. ex A. Spreng. =Digitaria laxa 

Rottboellia compressa L.f. =Hemiarthria altissima 

Rottboellia exaltata L.f. =Rottboellia cochinchinensis 

Saccharum caudatum G. Mey. =Imperata contracta 

Saccharum cayennense (P. Beauv.) Benth. =Eriochrysis 

cayennensis 

Saccharum sagittatum Aubl. =Gynerium sagittatum 

Schizachyrium riedelii (Trin.) A. Camus =Schizachyrium 

sanguineum 

Schizachyrium riedelii (Trin.) A. Camus var. multirameum 

(Hack.) Henr. =Schizachyrium sanguineum 

Schizachyrium semiberbe Nees =Schizachyrium sanguineum 

Setaria geniculata Beauv. =Setaria parviflora 

Setaria sulcata (Aubl.) A. Camus =Setaria palmifolia 

Sorghum vulgare Pers. =Sorghum bicolor 

Spartina brasiliensis Raddi =Spartina alterniflora 

Spheneria setifolia (Doell) Kuhlm. =Spheneria kegelii 

Sporobolus aeneus non (Trin.) Kunth, sensu Hitchcock 

=Sporobolus cubensis 

Sporobolus indicus Auctt., non (L.) R. Brown =Sporobolus 

jacquemontii 

Sporobolus littoralis (Lam.) Kunth =Sporobolus virginicus 

Sporobolus poiretii Auct., non (Roem. & Schult.) Hitchc. 

=Sporobolus jacquemontii 

Sporobolus villosus (A. Spreng.) Kunth =Sporobolus ciliatus 

Steirachne diandra non Ekman, sensu Swallen =Steirachne 

barbata 

Stenotaphrum glabrum Trin. var. americanum (Schrank) Doell 

=Stenotaphrum secundatum 

Strephium guianense Brongn. =Raddia guianensis 

Streptogyna crinita Auctt., non Beauv. =Streptogyna 

americana 

Streptostachys hirsuta P. Beauv. =Streptostachys asperifolia


Streptostachys vaginaeflorum Steud. =Streptostachys 

asperifolia 

Syntherisma chinensis (Nees) Hitchc. =Digitaria violascens 

Syntherisma cuyabensis (Trin.) Hitchc. =Digitaria lanuginosa 

Syntherisma digitata non (Sw.) Hitchc., sensu Hitchcock 

=Digitaria horizontalis 

Syntherisma longiflora (Retz.) Skeels =Digitaria longiflora 

Syntherisma malacophylla Hitchc. =Digitaria lanuginosa 

Syntherisma sanguinalis non (L.) Dulac, sensu Hitchcock 

=Digitaria ciliaris 

Thrasya campylostachya non (Hack.) Chase, sensu Amshoff & 

Henrard =Thrasya granitica 

Thrasya cinerascens (Doell) Judziewicz =Paspalum 

cinerascens 

Thrasya guianensis Swallen =Thrasya scandens 

Thrasya reticulata Swallen =Paspalum multinervium 

Trachypogon dactyloides (Steud.) E. Fourn. =Trachypogon 

spicatus 

Trachypogon plumosus (Willd.) Nees =Trachypogon spicatus 

Trachypogon polymorphus Hack. var. plumosus (Willd.) 

Hack. =Trachypogon spicatus 

Trachypogon ramosus Swallen =Trachypogon spicatus 

Trichachne insularis (L.) Nees =Digitaria insularis 

Triodia flaccida (Doell) Hitchc. =Tridens flaccidus 

Tripsacum latifolium non Hitchc. (1906), sensu Hitchcock 

(1922) =Tripsacum australe 

Tripsacum laxum non Nash, sensu Ostendorf =Tripsacum 

andersonii 

Valota insularis (L.) Chase =Digitaria insularis 

Valota laxa (A. Spreng.) Hitchc. & Chase =Digitaria laxa 

Vilfa villifera Steud. =Sporobolus ciliatus 

PODOCARPACEAE 


REFERENCES: 

Laubenfels, D.J. de. 1982. Podocarpaceae. In 


Laubenfels, D.J. de. 1985. A taxonomic revision 

of the genus Podocarpus. Blumea 30: 251- 

278. 

Stevenson, D. 1991. Podocarpaceae. In A.R.A. 


A) 9: 23-30. 

Podocarpus brasiliensis de Laubenfels [to be expected in GU] 

Podocarpus buchholzii de Laubenfels [to be expected in GU] 

Podocarpus magnifolius Buchholz & N. Gray [to be expected 

in GU] 

Podocarpus roraimae Pilger [GU?] [Mt. Roraima] 

Podocarpus steyermarkii Buchholz & N. Gray [to be expected 

in GU] 

Podocarpus tepuiensis Buchholz & N. Gray [to be expected in 

GU] 

REFERENCES: 

PODOSTEMACEAE 

SPERMATOPHYTA: Poaceae - Podostemaceae 

175 

Den Hollander, G. & C.C. Berg. 1983. Studies 

on the flora of the Guianas 4. A new species 

of Apinagia and a key to the Apinagia species 

in Suriname. Acta Bot. Neerl. 32(3): 223- 

226. 

Royen, P. van. 1951-1954. The Podostemaceae 

of the New World. Med. Bot. Mus. Herb. 

Rijks Univ. Utrecht 107: 1-151 (1951); 115: 

1-21 (1953); 119: 215-263. 

Schnell, R. 1969. Contribution à l'étude des 

Podostémacées de Guyane. Adansonia ser. 2 

9(2): 249-271. 

Apinagia arminensis van Royen [SU FG] 

Apinagia corymbosa (Tul.) Engl. var. corymbosa [GU SU] 

Apinagia crispa van Royen [SU] 

Apinagia digitata van Royen [FG] 

Apinagia divertens Went [SU FG] 

Apinagia exilis (Tul.) van Royen [GU SU] 

Apinagia flexuosa (Tul.) van Royen [GU SU FG] 

Apinagia goejei Went [SU] 

Apinagia guyanensis (Pulle) van Royen [SU FG] 

Apinagia hulkiana (Went) van Royen [GU SU] 

Apinagia imthurnii (Goebel) van Royen [GU SU FG?] 

Apinagia itanensis R. Schnell [FG] 

Apinagia latifolia (Goebel) van Royen [GU] 

Apinagia leptophylla (Goebel) van Royen [GU] 

Apinagia longifolia (Tul.) van Royen [GU SU FG] 

Apinagia marowynensis (Went) van Royen [SU FG?] 

Apinagia nana Went [SU] [=Apinagia pilgeri?] 

Apinagia penicillata (van Royen) van Royen [SU] 

Apinagia petiolata den Hollander [SU] 

Apinagia pilgeri Mildbraed [SU] 

Apinagia pusilla Tul. [GU] 

Apinagia richardiana (Tul.) van Royen [GU SU FG] 

Apinagia secundiflora (Tul.) Pulle [GU SU FG] 

Apinagia staheliana (Went) van Royen [GU SU] 

Apinagia surumuensis (Engl.) van Royen [GU] 

Apinagia treslingiana (Went) van Royen [SU] 

Apinagia versteegiana (Went) van Royen [SU] 

Jenmaniella ceratophylla Engl. var. ceratophylla [GU] 

Jenmaniella ceratophylla Engl. var. hexandra Engl. [GU] 

Jenmaniella guianensis Engl. [GU] 

Jenmaniella isoetifolia van Royen [GU] 

Jenmaniella jenmanii (Engl.) van Royen [GU] 

Jenmaniella tridactylitifolia Engl. [GU] 

Jenmaniella varians Engl. [GU] 

Marathrum capillaceum (Pulle) van Royen [SU FG] 

Marathrum pauciflorum Tul. var. heterophyllum van Royen 

[GU] 

Marathrum pauciflorum Tul. var. pauciflorum [GU] 

Marathrum squamosum Weddell [FG] 

Mourera fluviatilis Aubl. [GU SU FG] 

Oserya flabellifera Tul. & Wedd. [SU] [not in Guianas?] 

Oserya minima van Royen [SU] 

Oserya perpusilla (Went) van Royen [SU FG] 

Oserya sphaerocarpa Tul. & Wedd. [GU] 

Podostemum dichotomum Klotzsch [GU] [dubious species] 

Rhyncholacis apiculata van Royen [GU] 

Rhyncholacis applanata Goebel var. applanata [GU] 

Rhyncholacis applanata Goebel var. laxipinnata van Royen 

[GU]

176 


Rhyncholacis brevistamina van Royen [GU] 

Rhyncholacis coronata van Royen [GU] 

Rhyncholacis cristata van Royen [SU] 

Rhyncholacis dentata van Royen [SU] 

Rhyncholacis guyanensis van Royen [GU] 

Rhyncholacis hydrocichorium Tul. [GU] 

Rhyncholacis jenmanii Engl. [GU] 

Rhyncholacis macrocarpa Tul. [GU SU?] [dubious species] 

Rhyncholacis palmettifolia van Royen var. palmettifolia [GU] 

Rhyncholacis palmettifolia van Royen var. rosea van Royen 

[GU] 

Tristicha trifaria (Bory ex Willd.) Spreng. [GU FG] 

Weddellina squamulosa Tul. [GU SU FG] 

PODOSTEMACEAE - SYNONYMS AND 


Apinagia flabellifera (Tul. & Wedd.) Went =Oserya 

flabellifera 

Apinagia perpusilla Went =Oserya perpusilla 

Ligea flexuosa Tul. =Apinagia flexuosa 

Ligea longifolia (Tul.) Tul. =Apinagia longifolia 

Ligea richardiana Tul. =Apinagia richardiana 

Ligea secundiflora Tul. =Apinagia secundiflora 

Lophogyne capillacea Pulle =Marathrum capillaceum 

Neolacis secundiflora (Tul.) Wedd. =Apinagia secundiflora 

Oenone flexuosa (Tul.) Wedd. =Apinagia flexuosa 

Oenone guyanensis Pulle =Apinagia guyanensis 

Oenone hulkiana Went =Apinagia hulkiana 

Oenone imthurnii Goebel =Apinagia imthurnii 

Oenone longifolia Tul. =Apinagia longifolia 

Oenone marowynensis Went =Apinagia marowynensis 

Oenone penicillata van Royen =Apinagia penicellata 

Oenone richardiana (Tul.) Warm. =Apinagia richardiana 

Oenone secundiflora (Tul.) Engl. =Apinagia secundiflora 

Oenone treslingiana Went =Apinagia treslingiana 

Oenone versteegiana Went =Apinagia versteegiana 

Tristicha hypnoides (St. Hil.) Spreng. =Tristicha trifaria 

REFERENCES: 

POLYGALACEAE 

Chodat, R. 1893. Monographia Polygalacearum. 

Mém. Soc. phys. et d'hist. nat. Genève 31 part 

2(2): 1-500. 

Görts-van Rijn, A.R.A. 1974. Notes on 

Polygalaceae from Suriname. Acta Bot. 

Neerl. 23(2): 189-191. 

Görts-van Rijn, A.R.A. 1976. Polygalaceae 



523. 

Marques, M.d.C.M. 1979. Revisao das espécies 

do gênero Polygala L. (Polygalaceae) do 

estado do Rio de Janeiro. Rodriguésia 31(48): 

69-339. 

Marques, M.d.C.M. 1980. Revisao das espécies 

do gênero Bredemeyera Willd. (Polygalaceae) 

do Brasil. Rodriguésia 32(54): 269-321. 

Oort, A.J.P. 1939. Polygalaceae. In A. Pulle, 


SPERMATOPHYTA: Podostemaceae - Polygalaceae 

Barnhartia floribunda Gleason [GU SU FG] 

Bredemeyera densiflora A.W. Benn. var. densiflora [GU SU?] 

Bredemeyera densiflora A.W. Benn. var. glabra A.W. Benn. 

[GU SU] 

Bredemeyera floribunda Willd. [GU] 

Bredemeyera lucida (Benth.) A.W. Benn. [GU] 

Monnina cacumina N.E. Brown [GU] 

Moutabea guianensis Aubl. [GU SU FG] 

Moutabea longifolia Poepp. & Endl. [GU FG] 

Polygala adenophora DC. [GU SU FG] 

Polygala appressa Benth. var. appressa [GU SU FG] 

Polygala appressa Benth. var. gracillima A.W. Benn. [GU] 

Polygala blakeana Steyerm. [GU SU FG] 

Polygala coelosioides Mart. [GU?] 

Polygala echinosperma Görts [SU FG] 

Polygala galioides Poir. [GU SU FG] 

Polygala glochidiata Kunth [GU] 

Polygala hebeclada DC. [GU?] 

Polygala hygrophila Kunth [GU SU] [to be expected in FG] 

Polygala leptocaulis Torrey & Gray [GU SU FG] 

Polygala longicaulis Kunth [GU SU FG] 

Polygala maguirei Wurdack [GU] 

Polygala martiana A.W. Benn. [FG] 

Polygala membranacea (Miq.) Görts [SU FG] 

Polygala microspora S.F. Blake [GU] 

Polygala monticola Kunth [SU FG] 

Polygala paniculata L. [GU] 

Polygala planellasi Molinet & Gómez de la Maza [FG?] 

Polygala pseudocoelosioides Chodat [GU] 

Polygala savannarum Chodat [GU] 

Polygala spectabilis DC. [GU SU FG] 

Polygala spruceana A.W. Benn. [GU] 

Polygala subtilis Kunth [GU SU] 

Polygala timoutou Aubl. [GU SU FG] 

Polygala trichosperma L. [SU FG] 

Polygala variabilis Kunth [GU SU FG] 

Polygala violacea Aubl. emend. Marques [GU SU FG] 

Securidaca cayennensis S.F. Blake [FG] 

Securidaca coriacea Bonpl. [GU SU] 

Securidaca diversifolia (L.) S.F. Blake [GU SU FG] 

Securidaca lateralis A.W. Benn. [GU] 

Securidaca longifolia Poepp. & Endl. [GU?] 

Securidaca macrophylla (Benth.) Wurdack [FG] 

Securidaca marginata Benth. [GU] 

Securidaca orinocensis Rusby [GU?] [=Securidaca bialata?] 

Securidaca paniculata L.C. Rich var. lasiocarpa Oort [GU SU 

FG] 

Securidaca paniculata L.C. Rich var. paniculata [GU SU FG] 

Securidaca pubescens DC. [GU SU FG] 

Securidaca pubiflora Benth. [GU] 

Securidaca pyramidalis Sprague [GU] 

Securidaca retusa Benth. [GU FG] 

Securidaca rivinaefolia St. Hil. & Moq. var. parvifolia A.W. 

Benn. [GU SU] 

Securidaca rivinaefolia St. Hil. & Moq. var. rivinaefolia [GU 

SU] 

Securidaca spinifex Sandw. [GU SU FG] 

Securidaca tomentosa St. Hil. & Moq. [FG?] 

Securidaca uniflora Oort [GU SU FG]

Securidaca volubilis L. [GU SU FG] 


POLYGALACEAE - SYNONYMS AND 


Catocoma lucida Benth. =Bredemeyera lucida 

Monnina duidae S.F. Blake =Monnina cacumina 

Monnina membranacea Miq. =Polygala membranacea 

Polygala angustifolia Kunth =Polygala violacea 

Polygala angustifolia Kunth var. latifolia St. Hil. ex Chodat 

=Polygala violacea 

Polygala brizoides St. Hil. =Polygala violacea 

Polygala camporum Benth. =Polygala violacea 

Polygala mollis Kunth =Polygala monticola 

Polygala monticola Kunth var. bryzoides (St. Hil. & Moq.) 

Steyerm. =Polygala violacea 

Polygala paludosa St. Hil. =Polygala tenuis DC.[not in 


Polygala paludosa sensu Chodat, non St. Hil. =Polygala 

leptocaulis 

Polygala peduncularis A. Rich. =Polygala planellasi 

Polygala sororia Miq. =Polygala variabilis 

Polygala stipulata Chodat =Polygala membranacea 

Polygala violacea Vahl, non Aubl. =Polygala martiana 

Securidaca corytholobium A.W. Benn. =Securidaca 

macrophylla 

Securidaca densiflora Oort =Securidaca pubescens 

Securidaca hostmannii Miq. =Securidaca paniculata 

Securidaca latifolia Benth. =Securidaca pubiflora 

Securidaca major Sagot =Securidaca pubescens 

POLYGONACEAE 

reviewed by J. Brandbyge (1992) 

REFERENCES: 

Brandbyge, J. 1986. A revision of the genus 

Triplaris (Polygonaceae). Nord. J. Bot. 6(5): 

545-570. 

Cocucci, A.E. 1961. Revision del genero 

Ruprechtia (Polygonaceae). Kurtziana 1: 

217-269. 

Eyma, P.J. 1934. Polygonaceae. In A. Pulle, 


Howard, R.A. 1960. Studies in the genus 

Coccoloba, IX. A conspectus of the South 

American species. J. Arn. Arb. 41: 213-258, 

357-390. 

Howard, R.A. 1961. Studies in the genus 

Coccoloba, X. New species and a summary 

of distribution in South America. J. Arn. Arb. 

42: 87-95. 


Polygonaceae (additions and corrections). In 


1(2): 303-309. 

Wilson, K.L. 1990. Some widespread species of 

Persicaria and their allies. Kew Bull. 45: 621- 

636. 

SPERMATOPHYTA: Polygalaceae - Polygonaceae 

177 

Antigonon leptopus Hook. & Arnott [GU SU FG] [cultivated, 

naturalized] 

Coccoloba acuminata Kunth [GU] 

Coccoloba ascendens Duss ex Lindau [GU SU FG] 

Coccoloba charitostachya Standl. [GU] 

Coccoloba conduplicata Maguire [SU] 

Coccoloba cruegeri Lindau [GU SU] 

Coccoloba declinata (Vell.) Mart. [SU FG] 

Coccoloba densifrons Mart. ex Meisn. [GU] 

Coccoloba excelsa Benth. [GU SU FG] 

Coccoloba gymnorrhachis Sandw. [GU SU FG] 

Coccoloba latifolia Lam. [GU SU FG] 

Coccoloba lucidula Benth. [GU SU FG] 

Coccoloba marginata Benth. [GU SU FG] 

Coccoloba mollis Casaretto [SU FG] 

Coccoloba ovata Benth. [GU SU? FG?] 

Coccoloba parimensis Benth. [GU SU FG] 

Coccoloba savannarum Standl. [GU] 

Coccoloba schomburgkii Meisn. [GU] 

Coccoloba striata Benth. [GU] 

Coccoloba uvifera (L.) L. [GU SU FG] 

Polygonum acuminatum Kunth [GU SU FG] 

Polygonum ferrugineum Weddell [FG?] 

Polygonum glabrum Willd. [FG?] 

Polygonum hydropiperoides Michx. [SU] [to be expected in 

GU] 

Polygonum meisnerianum Cham. & Schlechtend. var. 

beyrichianum (Cham. & Schlechtend.) Meissn. [FG] 

Polygonum punctatum Elliott [GU SU FG] 

Ruprechtia brachysepala Meissn. [GU SU FG] 

Ruprechtia brachystachya Meisn. [GU] 

Ruprechtia tenuiflora Benth. [GU] 

Symmeria paniculata Benth. [GU SU] 

Triplaris americana L. [to be expected in GU] 

Triplaris weigeltiana (Rchb.) Kuntze [GU SU FG] 

POLYGONACEAE - SYNONYMS AND 


Coccoloba excoriata L. =Coccoloba tenuifolia L.[not in 


Coccoloba grandifolia Jacq. =Coccoloba pubescens L.[not in 


Coccoloba guianensis Meissn. =Coccoloba marginata 

Coccoloba micropunctata Eyma =Coccoloba excelsa 

Coccoloba sagotii Lindau =Coccoloba lucidula 

Coccoloba sparsifolia Lindau =Coccoloba ascendens 

Persicaria acuminata (Kunth) Gomez de la Maza =Polygonum 

acuminatum 

Persicaria densiflora (Meissn.) Moldenke =Polygonum 

glabrum s.l. 

Persicaria glabra (Willd.) Gomez de la Maza =Polygonum 

glabrum 

Persicaria hydropiperoides (Michx.) Small =Polygonum 

hydropiperoides 

Persicaria meisneriana (Cham. & Schlechtend.) Gomez de la 

Maza =Polygonum meisnerianum 

Persicaria punctata (Elliott) Small =Polygonum punctatum 

Ruprechtia marowynensis Eyma =Ruprechtia brachysepala 

Triplaris americana Aubl., non L. =Triplaris weigeltiana 

Triplaris schomburgkiana Benth. =Triplaris americana 

Triplaris surinamensis Cham. =Triplaris weigeltiana 

Triplaris surinamensis Cham. var. benthamiana Meisn. 

=Triplaris weigeltiana

178 


Triplaris surinamensis Cham. var. chamissoana Meisn. 

=Triplaris weigeltiana 

Triplaris surinamensis Cham. var. crassifolia Benth. 

=Triplaris weigeltiana 

Triplaris vahliana Fisch. & Mey. ex C.A. Meyer =Triplaris 

weigeltiana 

PONTEDERIACEAE 

contributed by C.N. Horn (1992); additions and 

corrections by C.N. Horn (1996) 

REFERENCES: 

Castellanos, A. 1958. Las Pontederiaceae de 

Brasil. Arq. Jard. Bot. Rio de Janeiro 16: 

149-216. 

Horn, C.N. 1994. Pontederiaceae. In A.R.A. 


A) 15: 91-102. 

Jonker-Verhoef, A.M.E. 1953. Pontederiaceae. 

In A. Pulle & J. Lanjouw, Flora of Suriname 

1(2): 81-90. 

Lowden, R.M. 1973. A revision of the genus 

Pontederia. Rhodora 75: 426-487. 

Eichhornia azurea (Swartz) Kunth [GU SU FG] 

Eichhornia crassipes (Mart.) Solms-Laubach [GU SU FG] 

Eichhornia diversifolia (Vahl) Urb. [GU SU FG] 

Eichhornia heterosperma Alexander [GU SU FG] 

Eichhornia paniculata (Spreng.) Solms-Laubach [GU] 

[probably naturalized] 

Pontederia rotundifolia L. [GU SU FG] 

Pontederia subovata (Seub.) Lowden [GU] 

Pontederia triflora (Seub.) Agostini, D. Velázquez & J. 

Velásquez [GU] 

PONTEDERIACEAE - SYNONYMS AND 


Eichhornia polysperma [dubious name of uncertain origin] 

Heteranthera diversifolia Vahl =Eichhornia diversifolia 

Heteranthera formosa Miq. =Eichhornia crassipes 

Heteranthera grandiflora Klotzsch [nomen nudum] 

=Eichhornia diversifolia 

Heteranthera zosterifolia Mart. [not in Guianas, misident. for 

Eichhornia diversifolia] 

Pontederia cordata L. [not in Guianas] 

Pontederia eriantha Miq. =Pontederia rotundifolia 

Pontederia schomburgkiana Klotzsch [nomen nudum] 

=Pontederia subovata 

SPERMATOPHYTA: Polygonaceae - Proteaceae 

PORTULACACEAE 

revised by R.A. DeFilipps (1996) 

REFERENCES: 

Legrand, C.D. 1962. Las especies americanas de 

Portulaca. An. Mus. Hist. Nat. Montevideo 

(ser. 2) 7: 1-147. 

Ooststroom, S.J. van. 1943. Portulacaceae. In 


Portulaca oleracea L. [GU SU FG] [introduced] 

Portulaca pilosa L. [GU SU? FG] 

Portulaca quadrifida L. [GU] [introduced?] 

Portulaca rubricaulis Kunth [FG?] 

Portulaca sedifolia N.E. Brown [GU SU FG] 

Portulaca teretifolia Kunth [FG?] 

Talinum fruticosum (L.) Juss. [SU?] 

Talinum paniculatum (Jacq.) Gaertn. [GU FG] 

PORTULACACEAE - SYNONYMS AND 


Portulaca cayennensis Legrand =Portulaca sedifolia 

Portulaca fruticosa L. =Talinum fruticosum 

Portulaca lanata Rich. =Portulaca pilosa 

Portulaca mucronata Link [not in Guianas] 

Portulaca paniculata Jacq. =Talinum paniculatum 

Portulaca triangulare Jacq. =Talinum fruticosum 

Portulaca umbraticola Kunth =Portulaca oleracea 

Talinum triangulare (Jacq.) Willd. =Talinum fruticosum 

PROTEACEAE 

reviewed by G.T. Prance (1992) 

REFERENCES: 

Mennega, A.M.W. 1934. Proteaceae. In A. 


Mennega, A.M.W. 1968. Proteaceae (additions 



Sleumer, H.O. 1954. Proteaceae americanae. 

Bot. Jahrb. 76: 139-211. 

Euplassa glaziovii (Mez) Steyerm. [Mt. Roraima] 

Euplassa pinnata (Lam.) Johnston [GU SU FG] 

Panopsis rubescens (Pohl) Pittier var. rubescens [GU SU] 

Panopsis rubescens (Pohl) Pittier var. simulans Macbr. [GU 

SU FG] 

Panopsis sessilifolia (L.C. Rich) Sandw. [GU SU FG] 

Roupala montana Aubl. var. dentata (R. Brown) Sleumer [GU 

FG?] 

Roupala montana Aubl. var. montana [GU SU FG] 

Roupala nitida Rudge [FG]

Roupala obtusata Klotzsch [GU FG] 

Roupala schomburgkii Klotzsch [GU SU] 

Roupala schulzii Mennega [SU] 

Roupala suaveolens Klotzsch [GU] 


PROTEACEAE - SYNONYMS AND 


Adenostephanus guianensis Meisn. =Euplassa pinnata 

Andriapetalum cayennensis Klotzsch ex Meissn. =Panopsis 

sessilifolia 

Andriapetalum rubescens Pohl =Panopsis rubescens 

Andriapetalum sessilifolium (L.C. Rich.) Klotzsch =Panopsis 

sessilifolia 

Euplassa meridionalis Salisb. =Euplassa pinnata 

Panopsis hameliifolia (Rudge) Salisb. =Panopsis sessilifolia 

Panopsis schomburgkii (Klotzsch) Pittier =Roupala 

schomburgkii 

Roupala dentata R. Brown =Roupala montana var. dentata 

Roupala hameliifolia Rudge =Panopsis sessilifolia 

Roupala laurifolia Salisb. & Knight =Roupala nitida 

Roupala media R. Brown =Roupala montana var. montana 

Roupala pinnata Lam. =Euplassa pinnata 

Roupala pullei Mennega =Roupala schomburgkii 

Roupala pyrifolia Salisb. & Knight =Roupala montana var. 

montana 

Roupala sessilifolia L.C. Rich. =Panopsis sessilifolia 

REFERENCES: 

QUIINACEAE 


Quiinaceae. In A. Pulle, Flora of Suriname 

3(1): 355-365. 

Lacunaria crenata (Tul.) A.C. Smith [GU SU FG] 

Lacunaria jenmanii (Oliver) Ducke [GU SU FG] 

Lacunaria umbonata Pires [FG] 

Quiina albiflora A.C. Smith [GU] 

Quiina amazonica A.C. Smith [GU?] 

Quiina brevensis Pires [GU] 

Quiina congesta Cowan [SU] 

Quiina cruegeriana Griseb. [GU FG] 

Quiina guianensis Aubl. [GU SU FG] 

Quiina indigofera Sandw. [GU] 

Quiina integrifolia Pulle [SU FG] 

Quiina leptoclada Tul. [SU FG] 

Quiina macrophylla Tul. [FG] 

Quiina obovata Tul. [GU SU FG] 

Quiina oiapocensis Pires [GU? FG] 

Quiina parvifolia Lanj. & v. Heerdt [SU] 

Quiina pteridophylla (Radlk.) Pires [GU SU FG] 

Quiina rhytidopus Tul. [GU] 

Quiina sessilis Choisy [FG] 

Touroulia guianensis Aubl. [GU SU FG] 

QUIINACEAE - SYNONYMS AND 


Lacunaria silvatica (Pulle) A.C. Smith =Lacunaria jenmanii 

Quiina crenata Tul. =Lacunaria crenata 

Quiina decaisneana Planch. & Triana =Lacunaria sp. 

(jenmanii?) 

Quiina oblanceolata Sandw. =Quiina obovata 

Quiina silvatica Pulle =Lacunaria jenmanii 

Touroulia jenmanii Oliver =Lacunaria jenmanii 

Touroulia pteridophylla Radlk. =Quiina pteridophylla 

REFERENCES: 

RAFFLESIACEAE 

Gentry, A.H. 1973. Rafflesiaceae. In R.E. 

Woodson et al., Flora of Panama. Ann. 


Vattimo, I. de. 1956. Notes on Apodanthes 

caseariae Poit. and Pilostyles calliandrae 

(Gardn.) R. Br. Not. Syst. 15(2): 225-229. 

Vattimo, I. de. 1971. Contribuiçao ao 

conhecimento de tribo Apodanthea [sic] R. 

Br. Parte I - Conspecto das espécies. 

Rodriguesia 26(38): 37-62. 

Apodanthes caseariae Poit. [GU SU FG] 

Apodanthes surinamensis Pulle [SU] 

REFERENCES: 

RANUNCULACEAE 

SPERMATOPHYTA: Proteaceae - Rapateaceae 

179 

Lourteig, A. 1952. Ranunculáceas de 

Sudamérica templada. Darwiniana 9: 397- 

609. 

Lourteig, A. 1971. Ranunculaceae. In T. Lasser, 


Clematis dioica L. var. brasiliana (DC.) Eichl. [FG?] 

RANUNCULACEAE - SYNONYMS AND 


Clematis caracasana DC. =Clematis dioica var. brasiliana 

RAPATEACEAE 

reviewed by P. Berry with additions by J.J. de 

Granville (1992); reviewed by P. Berry (1996) 

REFERENCES: 

Boom, B.M. 1994. A new species of Rapatea 

section Paludosa from French Guiana. 

Brittonia 46(4): 314-316. 


Rapateaceae (additions and corrections). In

180 


A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 

326-328. 

Maguire, B. 1958. Rapateaceae. In B. Maguire, 



Bot. Gard. 10(1): 1-156. 

Maguire, B. 1965. Rapateaceae. In B. Maguire 



12(3): 1-285. 

Maguire, B. 1982. Rapateaceae. In Z. Luces de 

Febres, Flora de Venezuela 6(2): 85-203. 

Uittien, H. 1938. Rapateaceae. In A. Pulle, 


Cephalostemon affinis Körn. [SU] 

Cephalostemon gracilis (Poepp. & Endl.) Rob. Schomb. [SU?] 

Epidryos guayanensis Maguire [GU] 

Monotrema aemulans Körn. [GU] 

Potarophytum riparium Sandw. [GU] 

Rapatea fanshawei Maguire var. fanshawei [GU] 

Rapatea fanshawei Maguire var. minor Maguire [GU] 

Rapatea linearis Gleason [GU] 

Rapatea membranacea Maguire [GU] 

Rapatea paludosa Aubl. var. paludosa [GU SU FG] 

Rapatea paludosa Aubl. var. sessiliflora Maguire [SU] 

Rapatea saülensis Mori [FG] 

Rapatea steyermarkii Maguire [GU] 

Rapatea ulei Pilger [GU FG] 

Rapatea xiphoides Sandw. [GU] 

Saxofridericia aculeata Körn. [SU FG] 

Saxofridericia regalis Rob. Schomb. [GU SU] 

Spathanthus unilateralis (Rudge) Desv. var. abruptus Maguire 

[GU? FG] 

Spathanthus unilateralis (Rudge) Desv. var. jenmanii (N.E. 

Brown) Maguire [GU FG] 

Spathanthus unilateralis (Rudge) Desv. var. unilateralis [GU 

SU FG] 

Stegolepis angustata Gleason [GU] 

Stegolepis ferruginea Baker [GU] 

Stegolepis guianensis Klotzsch ex Körn. [GU] 

Stegolepis ptaritepuiensis Steyerm. [GU] 

Stegolepis sp. nov. [GU] [Mt. Ayanganna] 

Stegolepis steyermarkii Maguire [GU] 

Windsorina guianensis Gleason [GU] 

RAPATEACEAE - SYNONYMS AND 


Mnasium unilateralis Rudge =Spathanthus unilateralis var. 

unilateralis 

Rapatea unilateralis (Rudge) Schult. =Spathanthus unilateralis 

var. unilateralis 

Rapatea viscosa Gleason =Saxofridericia regalis 

Spathanthus jenmanii N.E. Brown =Spathanthus unilateralis 

var. jenmani 

RHABDODENDRACEAE 

REFERENCES: 

SPERMATOPHYTA: Rapateaceae - Rhamnaceae 

Prance, G.T. 1972. Rhabdodendraceae. Flora 


Rhabdodendron amazonicum (Spruce ex Benth.) Huber [GU 

SU FG] 

RHABDODENDRACEAE - SYNONYMS AND 


Lecostemon amazonicum Spruce ex Benth. =Rhabdodendron 

amazonicum 

Lecostemon sylvestre Gleason =Rhabdodendron amazonicum 

Rhabdodendron crassipes (Spruce ex Benth.) Huber 

=Rhabdodendron amazonicum 

Rhabdodendron crassipes (Spruce ex Benth.) Huber var. 

cayenense Benth. =Rhabdodendron amazonicum 

Rhabdodendron sylvestre (Gleason) Maguire 

=Rhabdodendron amazonicum 

REFERENCES: 

RHAMNACEAE 

Johnston, M.C. 1971. Revision of Colubrina. 

Brittonia 23: 2-53. 

Johnston, M.C. & L.V.A. Johnston. 1978. 

Rhamnus. Flora Neotropica Monogr. 20: 

1-96. 

Lanjouw, J. 1932. Rhamnaceae. In A. Pulle, 


Suessenguth, K. 1953. Rhamnaceae. In A. 

Engler & K. Prantl, Die Natürlichen 

Pflanzenfamilien 20d: 7-173. 

Ampelozizyphus amazonicus Ducke [GU SU FG] 

Colubrina glandulosa Perkins var. glandulosa [GU] 

Gouania blanchetiana Miq. [GU SU FG] 

Gouania canescens L.C. Rich. [FG] 

Gouania colurnifolia Reissek [GU] 

Gouania frangulifolia (Schult.) Radlk. [FG] 

Gouania lupuloides (L.) Urb. s.l. [FG] 

Gouania polygama (Jacq.) Urb. [GU SU] 

Gouania pubescens Poir. [FG?] 

Gouania striata L.C. Rich. [FG] 

Gouania velutina Reissek [GU SU] 

Gouania virgata Reissek ["Guiana"] 

Rhamnus ulei Pilger [GU?] [Mt. Roraima] 

Ziziphus cinnamomum Triana & Planch. [GU FG] 

Ziziphus jujuba Mill. [GU SU FG] [cultivated; records may be 

referable to Ziziphus mauritiana] 

Ziziphus mauritiana Lam. [GU SU FG] [cultivated, 

naturalized] 

RHAMNACEAE - SYNONYMS AND 


Ziziphus jujuba (L.) Lam., non Mill. =Ziziphus mauritiana

REFERENCES: 


RHIZOPHORACEAE 

Alston, A.H.G. 1925. Revision of Cassipourea. 

Kew Bull. 1925: 241-276. 

Breteler, F.J. 1969. The Atlantic species of 

Rhizophora. Acta Bot. Neerl. 18: 434-444. 

Ding Hou. 1960. A review of the genus 

Rhizophora. Blumea 10: 625-634. 

Jonker, F.P. 1942. Rhizophoraceae. In A. Pulle, 


Juncosa, A.M. & P.B. Tomlinson. 1988. A 

historical and taxonomic synopsis of 

Rhizophoraceae and Anisophylleaceae. Ann. 


Prance, G.T. et al. 1975. Revisao taxonômica 

das espécies amazonicas de Rhizóphoraceae. 

Acta Amazonica 5: 5-22. 

Steyermark, J.A. & R. Liesner. 1983. Revision of 

the genus Sterigmapetalum. Ann. Missouri 

Bot. Gard. 70: 179-193. 

Cassipourea elliptica (Swartz) Poir. [SU FG] [=Cassipourea 

guianensis?] 

Cassipourea guianensis Aubl. [GU SU FG] 

Cassipourea lasiocalyx Alston [GU] 

Cassipourea serrata Benth. [GU] 

Rhizophora harrisonii Leechm. [GU SU] [putative hybrid 

species] 

Rhizophora mangle L. [GU SU FG] 

Rhizophora racemosa G. Mey. [GU SU FG] 

Sterigmapetalum exappendiculatum Steyerm. & Liesner 

[perhaps to be expected in GU] 

Sterigmapetalum guianense Steyerm. subsp. guianense [SU] 

[to be expected in GU] 

Sterigmapetalum resinosum Steyerm. & Liesner [perhaps to be 


REFERENCES: 

ROSACEAE 

Kleinhoonte, A. 1939. Rosaceae. In A. Pulle, 


Prunus myrtifolia (L.) Urb. var. accumulans Koehne [GU SU 

FG] 

Rubus guyanensis Focke [GU] 

RUBIACEAE 

reviewed by C.M. Taylor (1992); reviewed by 

C.M. Taylor & J.H. Kirkbride (1996) 

REFERENCES: 

181 

Andersson, L. 1992. A Provisional Checklist of 

Neotropical Rubiaceae. National Botanic 

Garden of Belgium, Meise. 

Andersson, L. 1995. Tribes and genera of the 

Cinchoneae complex (Rubiaceae). Ann. 


Boom, B.M. 1984. A revision of Isertia 

(Isertieae: Rubiaceae). Brittonia 36(4): 425- 

454. 

Bremekamp, C.E.B. 1934. Rubiaceae. In A. 


Bremekamp, C.E.B. 1937. Rubiaceae (additions 


Suriname 4(1): 475-491. 

Burger, W. & C.M. Taylor. 1993. Rubiaceae. In 

W. Burger, Flora Costaricensis. Fieldiana 

Bot. (n.s.) 33: 1-333. 

Howard, R.A. 1989. Rubiaceae. In R.A. 

Howard, Flora of the Lesser Antilles 6: 389- 

471. 

Kirkbride, J.H. Jr. 1977. Index to the Rubiaceae 

by Julian A. Steyermark in the Botany of the 

Guayana Highland by B. Maguire and 

collaborators. Phytologia 36(4): 324-366. 

Steyermark, J.A. 1964. Rubiaceae. In B. 



Bot. Gard. 10(5): 1-278. 




Bot. Gard. 12(3): 1-285. 




Bot. Gard. 17(1): 1-439. 




Bot. Gard. 23: 1-832. 

Steyermark, J.A. 1974. Rubiaceae. In T. Lasser, 


Steyermark, J.A. 1981. New species of 

Rubiaceae from French Guiana, Brazil, and 

Colombia. Brittonia 33(3): 385-400. 

Steyermark, J.A. 1984. New Rubiaceae from 

French Guiana. Brittonia 36(2): 154-160. 

Taylor, C.M. 1992. Revision of Cosmibuena 

(Rubiaceae). Ann. Missouri Bot. Gard. 79: 

886-900. 

Taylor, C.M. 1994. Revision of Hillia 

(Rubiaceae). Ann. Missouri Bot. Gard. 81(4): 

571-609. 

Alibertia acuminata (Benth.) Sandw. var. acuminata [GU FG] 

Alibertia edulis (L.C. Rich.) A. Rich ex DC. [GU SU FG] 

Alibertia latifolia (Benth.) K. Schum. var. latifolia [GU SU 

FG] 

Alibertia myrciifolia K. Schum. var. myrciifolia [GU SU FG] 

Alibertia surinamensis (Bremek.) Steyerm. [SU FG] 

Alibertia triflora (A. Rich.) K. Schum. [GU SU FG] 

SPERMATOPHYTA: Rhizophoraceae - Rubiaceae

182 

SPERMATOPHYTA: Rubiaceae 


Alibertia triloba Steyerm. [GU] 

Alseis floribunda Schott [FG] 

Alseis longifolia Ducke var. pentamera Bremek. [GU SU FG] 

Alseis smithii Standl. [GU] 

Amaioua brevidentata Steyerm. [FG?] 

Amaioua corymbosa Kunth [GU SU FG] 

Amaioua guianensis Aubl. var. guianensis [GU SU FG] 

Antirhea triflora J.H. Kirkbride, sp. nov. ined. [FG] 

Bertiera diversiramea Steyerm. [SU FG] [to be expected in 

GU] 

Bertiera guianensis Aubl. subsp. guianensis var. guianensis 

[GU SU FG] 

Bertiera palustris A. Rich. ex DC. [FG] [excluded from 

Bertiera by Steyermark] 

Borreria alata (Aubl.) DC. [GU FG] 

Borreria capitata (Ruiz & Pav.) DC. var. capitata [GU SU FG] 

Borreria capitata (Ruiz & Pav.) DC. var. suaveolens (G. Mey.) 

Steyerm. [GU SU] 

Borreria capitata (Ruiz & Pav.) DC. var. tenella (Kunth) 

Steyerm. [GU SU FG] 

Borreria cataractarum Steyerm. [GU] 

Borreria confertifolia Steyerm. [GU] 

Borreria densiflora DC. [GU FG] 

Borreria hispida Spruce ex K. Schum. var. hispida [GU SU 

FG] 

Borreria latifolia (Aubl.) K. Schum. var. latifolia [GU SU FG] 

Borreria oligodonta Steyerm. [GU SU] 

Borreria prostrata (Aubl.) Miq. [GU SU FG] 

Borreria verticillata (L.) G. Mey. [GU SU FG] 

Borreria xanthophylla Bremek. [SU] 

Bothriospora corymbosa Hook.f. [GU] 

Capirona decorticans Spruce [SU FG] [to be expected in GU] 

Chalepophyllum guyanense J.D. Hook. var. guyanense [GU] 

Chalepophyllum longilobum Steyerm. [GU] 

Chimarrhis cymosa Jacq. [GU] 

Chimarrhis microcarpa Standl. var. microcarpa [GU SU FG] 

Chimarrhis turbinata DC. [GU SU FG] 

Chiococca alba (L.) Hitchc. subsp. alba [GU FG] 

Chiococca multipedunculata Steyerm. [GU] 

Chiococca nitida Benth. var. amazonica Muell. Arg. [GU] 

Chiococca nitida Benth. var. nitida [GU FG] 

Chiococca pubescens Humb. & Bonpl. ex Roem. & Schult. 

[FG] [to be expected in GU] 

Chomelia angustifolia Benth. [GU] 

Chomelia glabriuscula Steyerm. [FG] 

Chomelia malaneoides Muell. Arg. [FG] 

Chomelia obtusa Cham. & Schlechtend. [GU] 

Chomelia schomburgkii Steyerm. [GU] 

Chomelia splitgerberi Bremek. [SU] 

Chomelia tenuiflora Benth. [GU SU FG] 

Chomelia volubilis (Standl.) Steyerm. [GU?] 

Coccocypselum aureum (Spreng.) Cham. & Schlechtend. var. 

capitatum (Benth.) Steyerm. [GU FG] 

Coccocypselum brevipetiolatum Steyerm. [GU?] 

Coccocypselum guianense (Aubl.) K. Schum. var. guianense 

[GU SU FG] 

Coccocypselum guianense (Aubl.) K. Schum. var. patens 

Steyerm. [GU FG] 

Coccocypselum hirsutum Bartling ex DC. [GU FG] 

Coccocypselum lanceolatum (Ruiz & Pav.) Pers. [GU?] 

Coccocypselum tontanea Kunth [GU FG] 

Coffea arabica L. [GU SU FG] [cultivated or escaped from 

cultivation] 

Cosmibuena grandiflora (Ruiz & Pav.) Rusby [GU SU? FG?] 

Coussarea amapaënsis Steyerm. [GU FG] 

Coussarea fanshawei Steyerm. [GU] 

Coussarea grandis Muell. Arg. [FG?] 

Coussarea hallei Steyerm. [FG] 

Coussarea hirticalyx Standl. var. glabrior Steyerm. [FG?] 

Coussarea leptophragma Muell. Arg. [SU] 

Coussarea mapourioides Bremek. [SU FG] 

Coussarea micrococca Bremek. [SU FG] 

Coussarea paniculata (Vahl) Standl. [GU SU FG] 

Coussarea racemosa A. Rich. [GU SU FG] 

Coussarea surinamensis Bremek. [SU] 

Coussarea violacea Aubl. [GU SU FG] 

Coutarea hexandra (Jacq.) K. Schum. var. hexandra [GU SU 

FG] 

Declieuxia fruticosa (Willd. ex Roem. & Schult.) Kuntze [GU 

SU FG] 

Declieuxia tenuiflora (Willd.) Steyerm. & Kirkbride [GU?] 

Didymochlamys connellii N.E. Brown [GU] 

Diodia apiculata (Willd. ex Roem. & Schult.) K. Schum. [GU] 

Diodia hyssopifolia (Willd. ex Roem. & Schult.) Cham. & 

Schlechtend. [GU SU FG] 

Diodia ocimifolia (Willd. ex Roem. & Schult.) Bremek. [GU 

SU FG] 

Diodia pulchristipula Bremek. [GU? SU] 

Diodia sarmentosa Swartz [GU SU FG] 

Diodia spicata Miq. [GU SU FG] 

Diodia teres Walter [GU] 

Duroia amapana Steyerm. [SU FG] 

Duroia aquatica (Aubl.) Bremek. [SU FG] 

Duroia eriopila L.f. var. eriopila [GU SU FG] 

Duroia eriopila L.f. var. tafelbergensis Steyerm. [SU] 

Duroia fusifera Hook.f. ex K. Schum. [GU] 

Duroia genipoides J.D. Hook. ex K. Schum. [GU SU FG] 

Duroia longiflora Ducke [GU SU FG] 

Duroia martiniana (Miers) Bremek. [FG] 

Duroia merumensis Steyerm. [GU] 

Duroia micrantha (Ladbr.) Zarucchi & Kirkbride [GU SU FG] 

Duroia plicata Benoist [FG] 

Elaeagia maguirei Standl. var. maguirei [GU SU] 

Emmeorrhiza umbellata (Spreng.) K. Schum. [GU SU FG] 

Faramea brevipes Steyerm. [GU] 

Faramea calyciflora A. Rich. ex DC. [FG] 

Faramea capillipes Muell. Arg. [GU SU FG] [specimens in 

Guianas may represent misidentifications] 

Faramea corymbosa Aubl. [GU SU FG] 

Faramea costata Steyerm. [SU FG] 

Faramea crassifolia Benth. [GU] 

Faramea cyanea Muell. Arg. [GU?] 

Faramea egregia Sandw. [GU] 

Faramea fanshawei Steyerm. [GU FG] 

Faramea guianensis (Aubl.) Bremek. [GU SU FG] 

Faramea irwinii Steyerm. [GU SU FG] 

Faramea lourteigiana Steyerm. [SU? FG] 

Faramea maguirei Steyerm. [GU] 

Faramea multiflora A. Rich. var. epedunculata Steyerm. [GU 

SU FG] 

Faramea multiflora A. Rich. var. multiflora [GU FG] 

Faramea multiflora A. Rich. var. salicifolia (K. Presl) Steyerm. 

[FG?] 

Faramea occidentalis (L.) A. Rich. subsp. occidentalis [GU 

SU FG] 

Faramea paniculata (Aubl.) Benth. [FG] [=Coussarea sp.?] 

Faramea quadricostata Bremek., emend Steyerm. [GU SU FG] 

Faramea sessiliflora Aubl. var. pedunculata Bremek. [SU FG] 

Faramea sessiliflora Aubl. var. sessiliflora [FG] 

Faramea sessilifolia (Kunth) A. DC. [GU FG]


Faramea spathacea Muell. Arg. ex Standl. [GU] 

Faramea tinguana Muell. Arg. [FG?] 

Faramea torquata Muell. Arg. [GU?] 

Faramea umbelliflora Bremek. [SU FG] 

Ferdinandusa goudotiana K. Schum. var. eciliata Steyerm. 

[GU] 

Ferdinandusa goudotiana K. Schum. var. goudotiana [Mt. 

Roraima] 

Ferdinandusa goudotiana K. Schum. var. psilocarpa Steyerm. 

[GU] 

Ferdinandusa guainiae Spruce ex K. Schum. [GU] 

Ferdinandusa paraensis Ducke [GU SU FG] 

Ferdinandusa rudgeoides (Benth.) Wedd. [GU SU FG] 

Galium hypocarpium (L.) Endl. ex Griseb. subsp. 

hypocarpium [GU] 

Galium surinamense Dempster [SU] 

Genipa americana L. [GU SU FG] 

Genipa spruceana Steyerm. [GU SU FG] [=Genipa americana 

L.?] 

Geophila cordifolia Miq. var. cordifolia [GU SU FG] 

Geophila gracilis (Ruiz & Pav.) DC. [FG?] 

Geophila granvillei Steyerm. [FG] [nomen nudum?] 

Geophila repens (L.) I.M. Johnston [GU SU FG] 

Geophila tenuis (Muell. Arg.) Standl. [GU FG] 

Gonzalagunia bunchosioides Standl. [GU] 

Gonzalagunia cornifolia (Kunth) Standl. [GU FG] 

Gonzalagunia dicocca Cham. & Schlechtend. subsp. dicocca 

var. guianensis Steyerm. [GU SU FG] 

Gonzalagunia dicocca Cham. & Schlechtend. subsp. 

venezuelensis Steyerm. [SU] 

Gonzalagunia spicata (Lamb.) Gómez [GU SU FG] 

Gonzalagunia surinamensis Bremek. [GU SU] 

Guettarda acreana Krause [GU SU FG] 

Guettarda argentea Lam. [FG] 

Guettarda aromatica Poepp. & Endl. [GU SU FG] 

Guettarda divaricata (Roem. & Schult.) Standl. [GU] 

Guettarda fanshawei Steyerm. [GU FG] 

Guettarda macrantha Benth. [GU SU FG] 

Guettarda pohliana Muell. Arg. [SU] 

Guettarda roupalaefolia Rusby [FG] 

Guettarda spruceana Muell. Arg. [GU SU FG] 

Guettarda viburnoides Cham. & Schlechtend. [GU] 

Hamelia axillaris Swartz [FG] 

Hamelia patens N.J. Jacq. [GU] 

Henriquezia jenmanii K. Schum. [GU] 

Hillia illustris (Vell.) K. Schum. [GU SU FG] 

Hillia parasitica Jacq. [GU FG] 

Hillia ulei K. Krause [SU] 

Hoffmannia megistophylla Standl. [GU FG] 

Isertia coccinea (Aubl.) J.F. Gmelin [GU SU FG] 

Isertia hypoleuca Benth. [GU SU FG] 

Isertia laevis (Triana) B. Boom [GU] 

Isertia parviflora Vahl [GU SU FG] 

Isertia rosea Spruce ex K. Schum. [FG] 

Isertia spiciformis DC. [GU SU FG] 

Isertia wilhelminensis Steyerm. [SU] 

Ixora acuminatissima Muell. Arg. [GU SU FG] 

Ixora aluminicola Steyerm. [GU SU FG] 

Ixora coccinea L. [GU SU FG] [cultivated or escaped from 

cultivation] 

Ixora davisii Sandw. [GU FG] 

Ixora ferrea (Jacq.) Benth. [GU?] 

Ixora graciliflora Benth. [GU SU FG] 

Ixora intensa K. Krause [FG] 

Ixora intropilosa Steyerm. [GU] 

183 

Ixora panurensis Muell. Arg. [GU] 

Ixora parviflora Vahl [GU] 

Ixora piresii Steyerm. [SU FG] 

Ixora potaroensis Steyerm. [GU FG] 

Ixora pubescens Willd. ex Roem. & Schult. [FG] 

Ixora schomburgkiana Benth. [GU FG] 

Ixora ulei Krause [GU SU] 

Ixora versteegii Bremek. [SU FG] 

Kotchubaea insignis Fisch. ex DC. [GU FG] 

Ladenbergia lambertiana (A. Braun ex Mart.) Klotzsch [GU] 

Limnosipanea schomburgkii Hook.f. [GU] 

Limnosipanea spruceana Hook.f. [GU?] 

Machaonia brasiliensis (Humb.) Cham. & Schlechtend. [GU] 

Maguireothamnus speciosus (N.E. Brown) Steyerm. [GU] 

Malanea ciliolata Steyerm. [GU] 

Malanea cruzii Steyerm. [GU] 

Malanea hypoleuca Steyerm. [GU FG] 

Malanea macrophylla Bartling ex Griseb. var. macrophylla 

[GU SU FG] 

Malanea obovata Hochr. [GU] 

Malanea sarmentosa Aubl. [GU SU FG] 

Malanea schomburgkii Steyerm. [GU] 

Malanea tafelbergensis Steyerm. [GU SU] 

Malanea ueiensis Steyerm. [GU] 

Manettia alba (Aubl.) Wernham [GU SU FG] 

Manettia coccinea (Aubl.) Willd. var. coccinea [GU SU FG] 

Manettia coccinea (Aubl.) Willd. var. spraguei (Wernham) 

Steyerm. [SU] [to be expected in GU] 

Manettia reclinata L. [GU FG] 

Merumea plicata Steyerm. [GU] 

Mitracarpus frigidus (Willd. ex Roem. & Schult.) K. Schum. 

var. fruticosus (Standl.) Steyerm. [GU] 

Mitracarpus hirtus (L.) DC. [GU SU FG] 

Mitracarpus humboldtianus Cham. & Schlechtend. [GU? SU? 

FG?] 

Mitracarpus microspermus K. Schum. [GU SU] 

Mitracarpus scabrellus Benth. [GU] 

Morinda aurantiaca (K. Krause) Steyerm. [GU] 

Morinda brachycalyx (Bremek.) Steyerm. [GU SU FG] 

Morinda calycina (Benth.) Steyerm. [GU] 

Morinda citrifolia L. [GU] [cultivated, naturalized] 

Morinda debilis (Sandw.) Steyerm. [GU] 

Morinda longiloba (Steyerm.) Steyerm. [GU] 

Morinda peduncularis Kunth [GU] 

Morinda sparsiflora (Steyerm.) Steyerm. [GU] 

Morinda surinamensis (Bremek.) Steyerm. [SU] 

Morinda tenuiflora (Benth.) Steyerm. var. leiophylla 

(Steyerm.) Steyerm. [GU FG] 

Morinda tenuiflora (Benth.) Steyerm. var. tenuiflora [GU 

FG] 

Neobertiera gracilis Wernham [GU SU FG] 

Oldenlandia corymbosa L. [GU SU FG] [=Hedyotis 

corymbosa (L.) Lam.] 

Oldenlandia lancifolia (Schumacher) DC. [GU SU FG] 

[naturalized] [=Hedyotis lancifolia Schumacher] 

Oldenlandia tenuis K. Schum. [GU] 

Oldenlandiopsis callitrichoides (Griseb.) Terrell & W.H. 

Lewis [GU] [introduced] 

Pagamea capitata Benth. subsp. capitata [GU SU] 

Pagamea capitata Benth. subsp. thibaudiifolia (Wernh.) 


Pagamea coriacea Spruce ex Benth. var. acuta Steyerm. [GU] 

Pagamea coriacea Spruce ex Benth. var. coriacea [GU] 

Pagamea guianensis Aubl. var. guianensis [GU SU FG] 

Pagamea thyrsiflora Spruce ex Benth. [GU] 

SPERMATOPHYTA: Rubiaceae

184 



Palicourea amapaënsis Steyerm. [FG] 

Palicourea calophylla DC. [GU SU FG] 

Palicourea corymbifera (Muell. Arg.) Standl. [GU FG] 

Palicourea crocea (Swartz) Roem. & Schult. [GU FG] 

Palicourea croceoides W. Hamilton [GU SU FG] [=Palicourea 

riparia] 

Palicourea expetens Standl. [GU FG] 

Palicourea formosa (A. Rich.) Lemée [FG] 

Palicourea grandiflora (Kunth) Standl. [FG] 

Palicourea grandifolia (Willd.) Standl. [FG?] 

Palicourea guianensis Aubl. subsp. guianensis [GU SU FG] 

Palicourea guianensis Aubl. subsp. occidentalis Steyerm. [GU] 

Palicourea irwinii Steyerm. [FG] 

Palicourea longiflora (Aubl.) A. Rich. [GU SU FG] 

Palicourea longistipulata (Muell. Arg.) Standl. subsp. 

chrysorrachis (Bremek.) Steyerm. [GU SU FG] 

Palicourea lutea (Aubl.) Bremek. ex Lemée [FG] [=Palicourea 

longiflora?] 

Palicourea marcgravii St. Hil. [FG] 

Palicourea obtusata Krause [GU] 

Palicourea pedunculosa (A. Rich.) DC. ex Lemée [FG] 

[dubious name] 

Palicourea perquadrangularis Wernh. [GU] 

Palicourea quadrifolia (Rudge) DC. [GU SU FG] 

Palicourea rigida Kunth [GU SU FG] 

Palicourea triphylla DC. [GU FG] 

Patabea tenuiflora DC. [FG] [=Psychotria sp.] 

Patima guianensis Aubl. [GU FG] 

Perama dichotoma Poepp. & Endl. var. dichotoma [GU] 

Perama dichotoma Poepp. & Endl. var. scaposa (Gleason & 

Standl.) Steyerm. [GU SU] 

Perama galioides (Kunth) Poir. var. galioides [GU] 

Perama galioides (Kunth) Poir. var. intermedia Steyerm. [GU] 

Perama hirsuta Aubl. var. hirsuta [GU SU FG] 

Perama hirsuta Aubl. var. stricta (Benth.) Bremek. [GU SU?] 

Platycarpum eglandulosum Steyerm. [GU] 

Posoqueria coriacea Mart. & Gal. [GU] 

Posoqueria latifolia (Rudge) Roem. & Schult. subsp. gracilis 

(Rudge) Steyerm. [SU FG] 

Posoqueria latifolia (Rudge) Roem. & Schult. subsp. latifolia 

[GU SU FG] 

Posoqueria longiflora Aubl. [GU SU FG] 

Posoqueria panamensis (Walp. & Duchass.) Walp. [GU FG] 

Posoqueria trinitatis DC. [GU SU FG] 

Psychotria acuminata Benth. [GU FG] 

Psychotria aetantha (Sandw.) Steyerm. [GU] 

Psychotria alba Ruiz & Pav. [GU SU FG] 

Psychotria aligera Steyerm. [GU] 

Psychotria alloantha Steyerm. [FG] 

Psychotria altsonii (Sandw.) Steyerm. [GU] 

Psychotria amplectens Benth. [SU] 

Psychotria anceps Kunth var. anceps [GU SU FG] 

Psychotria antennaeformis Steyerm. [FG] 

Psychotria apoda Steyerm. [GU SU FG] 

Psychotria appuniana Steyerm. [GU] 

Psychotria astrellantha Wernham [GU SU FG] 

Psychotria aubletiana Steyerm. [GU FG] 

Psychotria ayangannensis Steyerm. [GU] 

Psychotria bahiensis DC. var. bahiensis [GU SU FG] 

Psychotria bahiensis DC. var. cornigera (Benth.) Steyerm. 

[GU SU FG] 

Psychotria barbiflora DC. [GU SU FG] 

Psychotria berteriana DC. subsp. venezuelica Steyerm. [GU] 

Psychotria blepharophylla (Standl.) Steyerm. [FG] 

Psychotria borjensis Kunth [SU FG] 

Psychotria bostrychothyrsus Sandw. [GU SU? FG] 

Psychotria brachybotrya Muell. Arg. [GU SU FG] 

Psychotria bracteocardia (DC.) Muell. Arg. [GU SU FG] 

Psychotria bremekampiana Steyerm. [SU FG] 

Psychotria brevipes DC. [FG] 

Psychotria callithrix (Miq.) Steyerm. var. callithrix [SU FG] 

Psychotria callithrix (Miq.) Steyerm. var. tontaneoides (Britt. 

& Standl.) Steyerm. [GU FG] 

Psychotria campylopoda Steyerm. [GU] 

Psychotria capitata Ruiz & Pav. subsp. amplifolia (Raeusch.) 


Psychotria capitata Ruiz & Pav. subsp. capitata [GU] 

Psychotria capitata Ruiz & Pav. subsp. inundata (Benth.) 

Steyerm. [GU] 

Psychotria carthagenensis Jacq. [GU SU FG] 

Psychotria cincta Standl. [GU] 

Psychotria colorata (Willd. ex Roem. & Schult.) Muell. Arg. 

subsp. colorata [GU FG] 

Psychotria concinna Oliver [GU?] [Mt. Roraima] 

Psychotria cordifolia Kunth subsp. perpusilla Steyerm. [GU 

SU FG] 

Psychotria coussareoides Standl. var. ciliata Steyerm. [FG] 

Psychotria crassa Benth. [GU?] [Mt. Roraima] 

Psychotria crocochlamys Sandw. [GU] 

Psychotria ctenophora Steyerm. [SU FG] 

Psychotria cupularis (Muell. Arg.) Standl. [GU SU FG] 

Psychotria cypellantha Steyerm. [FG] 

Psychotria deflexa DC. subsp. campyloneura (Muell. Arg.) 


Psychotria deflexa DC. subsp. deflexa [GU SU FG] 

Psychotria deflexa DC. subsp. venulosa (Muell. Arg.) 

Steyerm. [GU FG] 

Psychotria deinocalyx Sandw. [GU SU FG] 

Psychotria erecta (Aubl.) Standl. & Steyerm. [GU SU FG] 

Psychotria ernestii Krause [FG?] 

Psychotria everardii Wernh. subsp. everardii [Mt. Roraima] 

Psychotria fanshawei (Standl.) Steyerm. [GU] 

Psychotria farameoides Bremek. [SU] 

Psychotria galbaoensis Steyerm. [GU? FG] 

Psychotria granvillei Steyerm. [FG] 

Psychotria guadalupensis (DC.) Howard [GU FG] [subsp. 

venezuelica Steyerm.?] 

Psychotria hemicephaelis Wernham [GU] 

Psychotria hoffmannseggiana (Willd. ex Roem. & Schult.) 

Muell. Arg. var. erythrophylla (Muell. Arg.) Steyerm. 

[SU? FG] 

Psychotria hoffmannseggiana (Willd. ex Roem. & Schult.) 

Muell. Arg. var. hoffmannseggiana [GU SU FG] 

Psychotria horizontalis Swartz var. glaucescens (Kunth) 

Steyerm. [GU] 

Psychotria horizontalis Swartz var. horizontalis [GU] 

Psychotria hyptoides Benth. [GU] 

Psychotria imthurniana Oliver [GU] 

Psychotria iodotricha Muell. Arg. subsp. atricapilla (Bremek.) 

Steyerm. [SU FG] 

Psychotria iodotricha Muell. Arg. subsp. iodotricha [GU SU 

FG] 

Psychotria irwinii Steyerm. [GU SU FG?] 

Psychotria kaieteurensis Sandw. [GU] 

Psychotria kappleri (Miq.) Muell. Arg. ex Benoist [SU FG] 

Psychotria lateralis Steyerm. [FG] 

Psychotria ligularis (Rudge) Steyerm. var. carapichea (Poir.) 

Steyerm. [SU FG] 

Psychotria ligularis (Rudge) Steyerm. var. ligularis [GU SU 

FG]


Psychotria lupulina Benth. subsp. lupulina [GU SU FG] 

Psychotria macrophylla Ruiz & Pav. [GU FG] [subsp. 

macrophylla?] 

Psychotria macrophylla Ruiz & Pav. subsp. anomothyrsa (K. 

Schum. & J.D. Smith) Steyerm. [FG] 

Psychotria maguireorum Steyerm. [GU] 

Psychotria mapourioides DC. var. chionantha (DC.) Steyerm. 

[GU SU FG] 

Psychotria mapourioides DC. var. mapourioides [GU SU FG] 

Psychotria mapourioides DC. var. opaca (Bremek.) Steyerm. 

[GU SU FG] 

Psychotria mazaruniensis Standl. [GU FG] 

Psychotria medusula Muell. Arg. [GU FG] 

Psychotria merumensis Steyerm. [GU] 

Psychotria microbotrys Ruiz ex Standl. [FG] 

Psychotria microbracteata Steyerm. [FG] 

Psychotria microdon (DC.) Urb. [GU] 

Psychotria morindoides (A. Rich.) Lemée [FG] 

Psychotria moroidea Steyerm. [SU FG] 

Psychotria multiramosa Steyerm. [GU] 

Psychotria muscosa (Jacq.) Steyerm. subsp. breviloba 


Psychotria nivea (Sandw.) Steyerm. [GU] 

Psychotria oblita Wernham [GU?] [Mt. Roraima] 

Psychotria oblonga (DC.) Steyerm. [SU FG] 

Psychotria officinalis (Aubl.) Sandw. subsp. officinalis [GU 

SU FG] 

Psychotria officinalis (Aubl.) Sandw. subsp. wilhelminensis 

Steyerm. [SU] 

Psychotria oiapoquensis Steyerm. [FG] 

Psychotria pacimonica Muell. Arg. [FG] 

Psychotria pakaraimensis Steyerm. [GU] 

Psychotria paniculata (Aubl.) Raeusch. [GU SU FG] 

Psychotria perferruginea Steyerm. [FG] 

Psychotria phaneroloma Standl. & Steyerm. var. phaneroloma 

[GU] 

Psychotria platypoda DC. [GU SU FG] 

Psychotria plocamipes Wernham [GU] 

Psychotria podocephala (Muell. Arg.) Standl. [FG] 

Psychotria poeppigiana Muell. Arg. subsp. barcellana (Muell. 

Arg.) Steyerm. [GU SU FG] 

Psychotria poeppigiana Muell. Arg. subsp. poeppigiana [GU 

SU FG] 

Psychotria polycephala Benth. [GU] 

Psychotria potaroensis (Sandw.) Steyerm. [GU] 

Psychotria prancei Steyerm. [FG] 

Psychotria pseudinundata Wernham [SU FG] 

Psychotria psittacina Steyerm. [GU] 

Psychotria puberulenta Steyerm. [GU] 

Psychotria pullei Bremek. [SU FG] 

Psychotria pungens Steyerm. [FG] 

Psychotria racemosa (Aubl.) Raeusch. [GU SU FG] 

Psychotria ramiflora Rusby [GU] 

Psychotria remota Benth. [GU FG] 

Psychotria rosea (Benth.) Muell. Arg. var. rosea [GU FG] 

Psychotria saulensis Steyerm. [FG] 

Psychotria schlechtendaliana Muell. Arg. [FG?] 

Psychotria schomburgkii Benth. [GU] 

Psychotria sessilis (Vell.) Muell. Arg. var. angustifolia (Muell. 

Arg.) Steyerm. [GU] 

Psychotria spectabilis Steyerm. [FG] 

Psychotria speluncae Standl. & Steyerm. subsp. speluncae 

[Mt. Roraima] 

Psychotria squamelligera Steyerm. [FG] 

Psychotria subundulata Benth. [GU SU FG] 

185 

Psychotria surinamensis Bremek. [SU FG] 

Psychotria tapajozensis Standl. [GU SU] 

Psychotria tepuiensis (Steyerm.) Steyerm. [GU] 

Psychotria tetramera Steyerm. [GU] 

Psychotria tillettii Steyerm. [GU] 

Psychotria transiens Wernham [GU] 

Psychotria tricholoba Muell. Arg. [GU?] 

Psychotria trichophoroides Muell. Arg. [SU FG] 

Psychotria uliginosa Swartz [GU SU FG] 

Psychotria ulviformis Steyerm. [GU SU FG] 

Psychotria urceolata Steyerm. [FG] 

Psychotria urniformis Steyerm. [GU] 

Psychotria variegata Steyerm. [GU SU FG] 

Psychotria viridibractea Steyerm. [FG] 

Psychotria wessels-boeri Steyerm. [SU] 

Psychotria wilhelminensis Steyerm. [SU] 

Psychotria wonotobensis (Bremek.) Steyerm. [SU] 

Randia armata (Swartz) DC. [GU SU FG] 

Randia asperifolia (Sandw.) Sandw. [GU] 

Randia formosa (Jacq.) K. Schum. var. densiflora Bartling ex 

K. Schum. [GU SU FG] 

Randia formosa (Jacq.) K. Schum. var. formosa [GU] 

Randia hebecarpa Benth. [GU FG] 

Randia ruizana DC. [GU SU FG] 

Remijia roraimae (Benth.) K. Schum. var. guianensis Steyerm. 

[GU] 

Remijia roraimae (Benth.) K. Schum. var. roraimae [GU] 

Retiniphyllum concolor (Spruce ex Benth.) Muell. Arg. [GU] 

Retiniphyllum guianense Steyerm. [GU] 

Retiniphyllum laxiflorum (Benth.) N.E. Brown var. laxiflorum 

[GU SU] 

Retiniphyllum maguirei Standl. [SU] 

Retiniphyllum scabrum Benth. var. ayangannense Steyerm. 

[GU] 

Retiniphyllum scabrum Benth. var. scabrum [GU] 

Retiniphyllum schomburgkii (Benth.) Muell. Arg. subsp. 

schomburgkii [GU SU] 

Retiniphyllum truncatum Muell. Arg. [GU] 

Richardia brasiliensis Gomes [GU] 

Richardia scabra L. [FG] 

Rudgea ayangannensis Steyerm. [GU] 

Rudgea corniculata Benth. [FG] 

Rudgea cornifolia (Humb. & Bonpl. ex Roem. & Schult.) 

Stanley [GU SU FG] 

Rudgea cornigera Bremek. [GU SU FG] 

Rudgea crassiloba (Benth.) B.L. Robinson [GU SU? FG] 

Rudgea duidae (Standl.) Steyerm. [FG?] 

Rudgea enervia Steyerm. [GU] 

Rudgea graciliflora Standl. [GU?] 

Rudgea guyanensis (A. Rich.) Sandw. [FG] 

Rudgea hostmanniana Benth. subsp. hostmanniana [GU SU 

FG] 

Rudgea lanceifolia Salisb. [GU FG] 

Rudgea merumensis Steyerm. [GU] 

Rudgea oldemanii Steyerm. [FG] 

Rudgea sandemanii Sandw. [FG] 

Rudgea standleyana Steyerm. [GU SU FG] 

Rudgea stipulacea (DC.) Steyerm. [SU FG] [to be expected in 

GU] 

Rudgea tanaosepala Sandw. [GU FG] 

Sabicea aspera Aubl. [GU SU FG] 

Sabicea cinerea Aubl. [SU FG] [to be expected in GU] 

Sabicea colombiana Wernham [FG?] 

Sabicea glabrescens Benth. [GU SU FG] 

Sabicea oblongifolia (Miq.) Steyerm. [GU SU FG] 


186 



Sabicea paraensis (Schum.) Wernh. [FG] 

Sabicea romboutsii Bremek. [SU] 

Sabicea surinamensis Bremek. [SU] 

Sabicea velutina Benth. subsp. velutina [GU SU FG] 

Sabicea villosa Willd. ex Roem. & Schult. var. adpressa 

(Wernham) Standl. [FG?] 

Sabicea villosa Willd. ex Roem. & Schult. var. villosa [GU 

SU FG] 

Schradera polycephala DC. [GU SU FG] 

Schradera surinamensis Standl. [GU SU] 

Schradera ternata Steyerm. [GU] 

Semaphyllanthe intonsa (Steyerm.) L. Andersson [GU] 

Semaphyllanthe merumensis (Steyerm.) L. Andersson [GU] 

Semaphyllanthe spectabilis (Steyerm.) L. Andersson [GU] 

Semaphyllanthe venezuelensis (Steyerm.) L. Andersson [GU] 

Simira rubescens (Benth.) Bremek. ex Steyerm. [FG?] 

Simira tinctoria Aubl. [SU FG] 

Sipanea ayangannensis Steyerm. [GU] 

Sipanea biflora (L.f.) Cham. & Schlechtend. [GU SU FG] 

Sipanea cowanii Steyerm. [GU] 

Sipanea galioides Wernh. [GU] 

Sipanea glaberrima (Bremek.) Steyerm. [SU] 

Sipanea gleasonii Steyerm. [GU] 

Sipanea hispida Benth. ex Wernham [GU FG?] 

Sipanea micrantha Sandw. [GU] 

Sipanea ovalifolia Bremek. var. ovalifolia [SU FG] 

Sipanea ovalifolia Bremek. var. villosissima Steyerm. [FG] 

Sipanea pratensis Aubl. var. dichotoma (Kunth) Steyerm. [GU 

SU FG] 

Sipanea pratensis Aubl. var. pratensis [GU SU FG] 

Sipanea stahelii Bremek. [SU FG?] 

Sipanea wilson-brownei Cowan [GU FG] 

Spermacoce assurgens Ruiz & Pav. [GU SU FG] 

Spermacoce hexangularis Aubl. [FG] [name of uncertain 

application] 

Spermacoce radicans Aubl. [FG] [=Diodia sp.?] 

Spermacoce scandens Aubl. ex J.F. Gmel. [FG] [name of 


Sphinctanthus maculatus Spruce ex K. Schum. [FG] 

Sphinctanthus striiflorus (DC.) Hook.f. [FG?] 

Thieleodoxa nitidula Bremek. [SU] [=Alibertia sp.] 

Tocoyena formosa (Cham. & Schlechtend.) K. Schum. [GU?] 

Tocoyena guianensis K. Schum. var. communis Steyerm. [GU 

SU FG] 

Tocoyena guianensis K. Schum. var. guianensis [FG] 

Tocoyena longiflora Aubl. [FG] 

Tocoyena neglecta N.E. Brown [GU] 

Tocoyena surinamensis Bremek. [SU] 

Uncaria guianensis (Aubl.) J.F. Gmelin [GU SU FG] 

Uncaria tomentosa (Willd. ex Roem. & Schult.) DC. [GU SU 

FG] 

Warszewiczia coccinea (Vahl) Klotzsch [to be expected in the 


RUBIACEAE - SYNONYMS AND 


Alibertia lanceolata (Cham.) Mennega [nomen nudum] 

Anisomeris obtusa (Cham. & Schlechtend.) K. Schum. 

=Chomelia obtusa 

Anisomeris tenuiflora (Benth.) Pulle =Chomelia tenuiflora 

Anisomeris volubilis Standl. =Chomelia volubilis 

Appunia aurantiaca (K. Krause) Sandw. =Morinda aurantiaca 

Appunia brachycalyx (Bremek.) Steyerm. =Morinda 

brachycalyx 

Appunia calycina (Benth.) Sandw. =Morinda calycina 

Appunia debilis Sandw. =Morinda debilis 

Appunia longiloba Steyerm. =Morinda longiloba 

Appunia sparsiflora Steyerm. =Morinda sparsiflora 

Appunia surinamensis (Bremek.) Steyerm. =Morinda 

surinamensis 

Appunia tenuiflora (Benth.) Jackson & Hook.f. =Morinda 

tenuiflora 

Appunia tenuiflora (Benth.) Jackson & Hook.f. var. leiophylla 

Steyerm. =Morinda tenuiflora var. leiophylla 

Basanacantha asperifolia Sandw. =Randia asperifolia 

Basanacantha spinosa (Jacq.) K. Schum. =Randia armata 

Bellynkxia brachycalyx Bremek. =Morinda brachycalyx 

Bellynkxia surinamensis Bremek. =Morinda surinamensis 

Borreria fockeana Miq. =Borreria latifolia var. latifolia 

Borreria kappleriana Miq. =Borreria capitata var. capitata 

Borreria laevis Auctt., non (Lamb.) Griseb. =Spermacoce 

assurgens 

Borreria laevis (Lamb.) Griseb. =Spermacoce tenuoir L.[not in 


Borreria latifolia (Aubl.) K. Schum. var. fockeana (Miq.) 

Bremek. =Borreria latifolia var. latifolia 

Borreria ocimoides Auctt., non (Burm.f.) DC. =Borreria 

prostrata 

Borreria splitgerberi Bremek. =Borreria latifolia var. latifolia 

Borreria suaveolens G. Mey. =Borreria capitata var. 

suaveolens 

Brignolia pubigera Benth. =Isertia parviflora 

Bruinsmania isertioides Miq. =Isertia parviflora 

Calycophyllum intonsum Steyerm. =Semaphyllanthe intonsa 

Calycophyllum merumense Steyerm. =Semaphyllanthe 

merumensis 

Calycophyllum spectabile Steyerm. =Semaphyllanthe 

spectabilis 

Calycophyllum venezuelense Steyerm. =Semaphyllanthe 

venezuelensis 

Capirona leiophloea Benoist =Capirona decorticans 

Capirona surinamensis Bremek. =Capirona decorticans 

Carapichea guianensis Aubl. =Psychotria ligularis var. 

carapichea 

Carapichea kappleri Miq. =Psychotria kappleri 

Cephaelis aetantha Sandw. =Psychotria aetantha 

Cephaelis alba (Aubl.) Wernham =Psychotria ulviformis 

Cephaelis altsonii Sandw. =Psychotria altsonii 

Cephaelis amoena Bremek. =Psychotria colorata subsp. 

colorata 

Cephaelis barcellana (Muell. Arg.) Standl. =Psychotria 

poeppigiana subsp. barcellana 

Cephaelis blepharophylla Standl. =Psychotria blepharophylla 

Cephaelis bracteocardia DC. =Psychotria bracteocardia 

Cephaelis callithrix Miq. =Psychotria callithrix 

Cephaelis colorata Willd. ex Roem. & Schult. =Psychotria 

colorata 

Cephaelis dichotoma Rudge =Psychotria platypoda 

Cephaelis duckei Standl. =Psychotria spectabilis 

Cephaelis ernestii Krause =Psychotria transiens 

Cephaelis fanshawei Standl. =Psychotria cephaelis 

Cephaelis glabrescens (Muell. Arg.) Standl. =Psychotria 

glabrescens Muell. Arg.[not in Guianas] 

Cephaelis guianensis (Aubl.) Standl. =Psychotria ligularis var. 

carapichea 

Cephaelis hemicephaelis (Wernh.) Steyerm. =Psychotria 

hemicephaelis


Cephaelis hirta Miq. =Psychotria bremekampiana 

Cephaelis hoffmannseggiana Willd. ex Roem. & Schult. 

=Psychotria hoffmannseggiana 

Cephaelis jenmanii Wernham =Psychotria mazaruniensis 

Cephaelis kaieteurensis Wernham =Psychotria appuniana 

Cephaelis kappleri (Miq.) Standl. =Psychotria kappleri 

Cephaelis ligularis (Rudge) A. Rich. =Psychotria ligularis 

Cephaelis microcephala Miq. =Psychotria hoffmannseggiana 

Cephaelis muscosa (Jacq.) Swartz =Psychotria muscosa 

Cephaelis nivea Sandw. =Pschotria nivea 

Cephaelis patrisii Dietr. =Psychotria sp. 

Cephaelis potaroensis Sandw. =Psychotria potaroensis 

Cephaelis pubescens Hoffmanns. ex Roem. & Schult. 

=Psychotria bracteocardia 

Cephaelis purpurea (Aubl.) Willd. =Psychotria variegata 

Cephaelis rosea Benth. =Psychotria rosea 

Cephaelis rubra Hoffmanns. ex Roem. & Schult. =Psychotria 


Cephaelis surinamensis Standl. =Psychotria variegata 

Cephaelis sylvatica Bremek. =Psychotria ctenophora 

Cephaelis tatei Standl. =Psychotria crocochlamys 

Cephaelis tepuiensis Steyerm. =Psychotria tepuiensis 

Cephaelis tomentosa (Aubl.) Vahl =Psychotria poeppigiana 

subsp. poeppigiana 

Cephaelis tontaneoides (Britt. & Standl.) Williams & Cheesm. 

=Psychotria callithrix var. tontaneoides 

Cephaelis trichophoroides (Muell. Arg.) Standl. =Psychotria 

trichophoroides 

Cephaelis violacea (Aubl.) Swartz =Psychotria apoda 

Chalepophyllum speciosum N.E. Brown =Maguireothamnus 

speciosus 

Chimarrhis longistipulata Bremek. =Chimarrhis microcarpa 

var. microcarpa 

Chiococca cordata Cowan =Chiococca nitida var. amazonica 

Chiococca erubescens Wernh. =Chiococca nitida var. nitida 

Chytropsia astrellantha (Wernham) Bremek. =Psychotria 

astrellantha 

Chytropsia guianensis Bremek. =Psychotria moroidea 

Coffea guianensis Aubl. =Faramea sp.?[=Ixora guianensis 

(Aubl.) Spreng.] 

Coffea paniculata Aubl. =Faramea paniculata 

Cosmibuena triflora (Benth.) Klotzsch =Cosmibuena 

grandiflora var. grandiflora 

Coupoui aquatica Aubl. =Duroia aquatica 

Coupoui martiniana (Miers) Wernh. =Duroia martiniana 

Coupoui micrantha Ladbr. =Duroia micrantha 

Coussarea multiflora (A. Rich.) Lemée =Faramea multiflora 

Coussarea paniculata (Aubl.) Lemée, nom. illeg. =Faramea 

paniculata 

Coussarea schomburgkiana (Benth.) Benth. & J.D. Hook. 

=Coussarea violacea 

Coutarea speciosa Aubl. =Coutarea hexandra 

Cupirana martiniana Miers =Duroia martiniana 

Declieuxia roraimensis Wernh. =Psychotria capitata subsp. 

inundata 


Schlechtend. var. articulata (Pohl ex DC.) Steyerm. 

=Diodia hyssopifolia 


Schlechtend. var. linearis (Willd. ex Roem. & Schult.) 

Steyerm. =Diodia hyssopifolia 

Diodia surinamensis Miq. =Diodia hyssopifolia 

Duroia sprucei Rusby =Duroia micrantha 

Evea guianensis Aubl. =Faramea guianensis 

187 

Evea tontaneoides Britt. & Standl. =Psychotria callithrix var. 

tantaneoides 

Faramea longifolia Benth. =Faramea sessilifolia 

Faramea salicifolia K. Presl =Faramea multiflora var. 

salicifolia 

Faramea truncata DC. =Faramea corymbosa 

Gamotopea alba (Aubl.) Bremek. =Psychotria ulviformis 

Gamotopea callithrix (Miq.) Bremek. =Psychotria callithrix 

Gamotopea oblonga (DC.) Lemée =Psychotria oblonga 

Gamotopea purpurea (Aubl.) Bremek. =Psychotria variegata 

Gamotopea surinamensis (Standl.) Bremek. =Psychotria 

variegata 

Genipa americana L. var. caruto (Kunth) K. Schum. =Genipa 

americana 

Geophila herbacea (Jacq.) K. Schum. =Geophila repens 

Geophila violacea (Aubl.) DC. =Coccocypselum sp.? 

Guettarda coccinea Aubl. =Isertia coccinea 

Guettarda grandiflora Steud. =Guettarda aromatica 

Hamelia patens N.J. Jacq. var. glabra Oersted =Hamelia 

patens 

Hemidiodia ocimifolia (Willd.) K. Schum. =Diodia ocimifolia 

Hillia tubiflora Cham. =Hillia illustris 

Ibetralia surinamensis Bremek. =Alibertia surinamensis 

Isertia commutata Miq. =Isertia spiciformis 

Isertia parviflora Vahl var. hirta Steyerm. =Isertia parviflora 

Isertia pterantha Bremek. =Isertia spiciformis 

Ixora floribunda (Rich.) Griseb. [not in Guianas] 

Ixora guianensis (Aubl.) Spreng. [see Coffea guianensis] 

Ixora mazarunensis Standl. =Ixora davisii 

Ixora orinocensis Spruce ex Standl. [nomen nudum] 

Ixora surinamensis Bremek. =Ixora graciliflora 

Ixora tenuiflora (DC.) Lemée =Patabea tenuiflora 

Ixora xantholoba Standl. =Ixora graciliflora 

Malanea angustifolia Bartling =Malanea obovata 

Mapouria alba (Ruiz & Pav.) Wernh. =Pschotria alba 

Mapouria brownei (Spreng.) Lemée =Psychotria brownei 

Spreng.[not in Guianas] 

Mapouria chionantha (DC.) Muell. Arg. =Psychotria 

mapourioides var. chionantha 

Mapouria cupularis Muell. Arg. =Psychotria cupularis 

Mapouria ficigemma (DC.) Lemée =Psychotria carthagenensis 

Mapouria fockeana (Miq.) Bremek. =Psychotria 

carthagenensis 

Mapouria guianensis Aubl. =Psychotria mapourioides 

Mapouria mahuriensis Bremek. =Psychotria cupularis 

Mapouria microdon (DC.) Bremek. =Pschotria microdon 

Mapouria opaca Bremek. =Psychotria mapouroides var. opaca 

Mapouria ornithophila Bremek. =Psychotria anceps 

Mapouria patrisii (DC.) Lemée =Psychotria anceps 

Mapouria sororia (DC.) Lemée =Psychotria sp.[dubious name] 

Mapouria umbrosa Muell. Arg. =Psychotria borjensis 

Mapouria wonotobensis Bremek. =Psychotria wonotobensis 

Mitracarpus discolor Miq. =Mitracarpus humboldtianus 

Mitracarpus villosus (Swartz) Cham. & Schlechtend. ex DC. 

=Mitracarpus hirtus 

Mitrocarpum puberulum Benth. =Borreria prostrata 

Nacibea alba Aubl. =Manettia alba 

Nacibea coccinea Aubl. =Manettia coccinea 

Naletonia violacea (Aubl.) Bremek. =Psychotria capitata 

subsp. amplifolia 

Nonatelia longiflora Aubl. =Palicourea longiflora 

Nonatelia lutea Aubl. ex Steud. =Palicourea longiflora 

Nonatelia lutea Aubl. =Palicourea lutea [=Palicourea 

longiflora?] 

Nonatelia officinalis Aubl. =Psychotria officinalis 


188 



Nonatelia paniculata Aubl. =Psychotria paniculata 

Nonatelia racemosa Aubl. =Psychotria racemosa 

Nonatelia violacea Aubl. =Psychotria capitata 

Notopleura tapajozensis (Standl.) Bremek. =Psychotria 

tapajozensis 

Notopleura uliginosa (Swartz) Bremek. =Psychotria uliginosa 

Oldenlandia herbacea (L.) DC. [not in Guianas] 

Ourouparia guianensis Aubl. =Uncaria guianensis 

Palicourea "latifolia (Rudge) Roem. & Schult." [invalid name, 

error for Posoqueria latifolia] 

Palicourea cayennensis Desv. =Palicourea pedunculosa 

Palicourea crocea (Swartz) Roem. & Schult. var. riparia 

(Benth.) Griseb. =Palicourea croceoides 

Palicourea divaricata Schlechtend. =Psychotria 

schlechtendaliana 

Palicourea guianensis Aubl. var. tetramera Bremek. 

=Palicourea guianensis subsp. guianensis 

Palicourea guianensis Aubl. var. trimera Bremek. =Palicourea 

guianensis subsp. guianensis 

Palicourea nicotianifolia Auctt., not Cham. & Schlechtd. 

=Palicourea calophylla 

Palicourea riparia Benth. =Palicourea croceoides 

Palicourea roraimae Standl. =Psychotria oblita 

Patabea coccinea Aubl. =Ixora davisii 

Petagomoa atricapilla Bremek. =Psychotria iodotricha subsp. 

atricapilla 

Petagomoa glabra Bremek. =Psychotria sp. 

Petagomoa hirta (Miq.) Bremek. =Psychotria bremekampiana 

Petagomoa nigricans Bremek. =Psychotria trichophoroides 

Petagomoa tricholoba (Muell. Arg.) Bremek. =Psychotria 

tricholoba 

Posoqueria gracilis (Rudge) Roem. & Schult. =Posoqueria 

latifolia subsp. gracilis 

Posoqueria intermedia Miq. =Posoqueria trinitatis 

Pseudochimarrhis difformis Benoist =Chimarrhis sp. 

Pseudochimarrhis turbinata Benoist =Chimarrhis turbinata 

Psychotria amplifolia Raeusch. =Psychotria capitata subsp. 

amplifolia 

Psychotria amplinoda Steyerm. =Psychotria tapajozensis 

Psychotria amplinoda Steyerm. var. ayangannae Steyerm. 

=Psychotria tapajozensis 

Psychotria arcuata Benth. =Psychotria capitata var. inundata 

Psychotria barcellana Muell. Arg. =Psychotria poeppigiana 

subsp. barcellana 

Psychotria bracteata A. DC. =Psychotria pseudinundata 

Psychotria campyloneura Muell. Arg. =Psychotria deflexa 

subsp. campyloneura 

Psychotria capitata Ruiz & Pav. var. roraimensis (Wernh.) 

Steyerm. =Psychotria capitata subsp. inundata 

Psychotria chionantha (DC.) Britton =Psychotria mapourioides 

var. chionantha 

Psychotria chlorantha Benth. =Psychotria anceps 

Psychotria cornigera Benth. =Psychotria bahiensis var. 

cornigera 

Psychotria cuspidata Bredem. ex Roem. & Schult. [not in 


Psychotria dichotoma (Rudge) Bremek. =Psychotria platypoda 

Psychotria didymocarpos (A. Rich.) Lemée =Psychotria 

bahiensis var. bahiensis 

Psychotria flexuosa Willd. =Psychotria paniculata 

Psychotria fockeana Miq. =Psychotria carthaginensis 

Psychotria glabrescens Muell. Arg. [not in Guianas] 

Psychotria guianensis (Aubl.) Rusby =Palicourea guianensis 

Psychotria guianensis Raeusch. [nomen nudum] 

Psychotria inundata Benth. =Psychotria capitata subsp. 

inundata 

Psychotria involucrata Swartz =Psychotria officinalis 

Psychotria maguirei Standl. =Rudgea stipulacea 

Psychotria mapouria Roem. & Schult. =Psychotria 

mapourioides 

Psychotria mapuria Roem. & Schult. =Psychotria 

mapourioides 

Psychotria microcephala (Miq.) Miq. =Psychotria 


Psychotria nervosa Benth. =Psychotria lupulina subsp. 

lupulina 

Psychotria patens Swartz [not in Guianas] 

Psychotria pedunculosa A. Rich. =Palicourea pedunculosa 

Psychotria racemifera Standl. =Psychotria polycephala 

Psychotria romboutsii Bremek. =Psychotria amplectens 

Psychotria roraimensis Wernh. =Psychotria capitata subsp. 

inundata 

Psychotria rubra (Willd.) Muell. Arg. =Psychotria 


Psychotria rudgei Bremek. =Psychotria lupulina subsp. 

lupulina 

Psychotria setifera Benth. =Psychotria capitata subsp. inundata 

Psychotria sororopanensis Standl. & Steyerm. =Psychotria 

sessilis var. angustifolia 

Psychotria venulosa Muell. Arg. =Psychotria deflexa subsp. 

venulosa 

Psychotria violacea Aubl. =Coccocypselum sp.?[name of 


Relbunium guadalupense (Spreng.) Urban [not in Guianas] 

Relbunium hypocarpium (L.) Hemsl. subsp. hypocarpium 

=Galium hypocarpium subsp. hypocarpium 

Retiniphyllum maguirei Standl. var. reticulatum Steyerm. 

=Retiniphyllum maguirei 

Ronabea erecta Aubl. =Psychotria erecta 

Ronabea latifolia Aubl. =Psychotria erecta 

Ronabea morindoides A. Rich. =Psychotria morindoides 

Rondeletia glomerulata Spreng. [dubious name] 

Rosenbergiodendron densiflorum Fagerl. =Randia formosa 

var. densiflora 

Rosenbergiodendron formosum (Jacq.) Fagerl. =Randia 

formosa var. formosa 

Rosenbergiodendron longiflorum Fagerl. =Randia ruizana 

Rudgea obscura Sandw. =Psychotria astrellantha 

Rudgea ovalifolia Salisb. =Rudgea lanceifolia 

Rudgea tillettii Steyerm. =Psychotria coussareoides 

Sabicea glabrescens Benth. var. oblongifolia (Miq.) Sandw. 

=Sabicea oblongifolia 

Sabicea guianensis (Aubl.) Baill. =Patima guianensis 

Schradera ligularis Rudge =Psychotria ligularis var. lupulina 

Sickingia tinctoria (Aubl.) Lemée =Simira tinctoria


Sipanea coppenamensis Bremek. =Sipanea pratensis var. 

pratensis 

Solena latifolia Rudge =Posoqueria latifolia 

Spermacoce alata Aubl. =Borreria alata 

Spermacoce aspera Aubl. =Mitracarpus hirtus 

Spermacoce caerulescens Aubl. =Borreria latifolia var. 

latifolia 

Spermacoce densiflora (DC.) Liogier =Borreria densiflora 

Spermacoce guianensis Bremek. =Spermacoce assurgens 

Spermacoce hyssopifolia Willd. ex Roem. & Schult. =Diodia 

hyssopifolia 

Spermacoce latifolia Aubl. =Borreria latifolia var. latifolia 

Spermacoce longifolia Aubl. ex Miq. =Diodia ocimifolia 

Spermacoce longifolia Aubl. =Mitracarpus hirtus 

Spermacoce prostrata Aubl. =Borreria prostrata 

Spermacoce verticillata L. =Borreria verticillata 

Strempelia fimbriata (Benth.) Bremek. =Rudgea cornifolia 

Strempelia guyanensis A. Rich. =Rudgea guyanensis 

Tapogomea alba Aubl. =Psychotria ulviformis 

Tapogomea carapichea Poir. =Psychotria ligularis var. 

carapichea 

Tapogomea glabra Aubl. =Psychotria blepharophylla 

Tapogomea purpurea Aubl. =Psychotria variegata 

Tapogomea tomentosa Aubl. =Psychotria poeppigiana 

Tapogomea violacea Aubl. =Psychotria apoda 

Tontanea guianensis Aubl. =Coccocypselum guianense 

Uragoga alba (Aubl.) Kuntze =Psychotria ulviformis 

Uragoga glabra (Aubl.) Kuntze =Psychotria blepharophylla 

Uragoga hirta (Miq.) Pulle =Psychotria bremekampiana 

Uragoga kappleri (Miq.) Pulle =Psychotria kappleri 

Uragoga purpurea (Aubl.) Kuntze =Psychotria variegata 

Uragoga tomentosa (Aubl.) Kuntze =Psychotria poeppigiana 

subsp. poeppigiana 

Uragoga violacea (Aubl.) Pulle =Psychotria apoda 

RUPPIACEAE 


(1992) 

Ruppia maritima L. [GU SU FG] 

RUTACEAE 

reviewed by J. Kallunki (1996) 

REFERENCES: 

Albuquerque, B.W.P. de. 1977 ("1976"). 

Revisao taxonômica das Rutaceae do Estado 

do Amazonas. Acta Amazonica suppl. 6(3): 

1-67. 

Albuquerque, B.W.P. de. 1981. Novas 

combinaçoes em Angosture Roemer & 

Schultes (Rutaceae). Acta Amazonica 11(4): 

849-852. 

Cowan, R.S. 1960. Rutaceae. In B. Maguire & 

J.J. Wurdack, Botany of the Guayana 

Highland part IV. Mem. New York Bot. 

Gard. 10(2): 24-37. 

SPERMATOPHYTA: Rubiaceae - Rutaceae 

189 

Cowan, R.S. 1967. Rutaceae of the Guayana 

Highland. Mem. New York Bot. Gard. 14(3): 

1-14. 

Elias, T.S. 1970. The correct name for the genus 

Cusparia (Rutaceae). Taxon 19(4): 573-575. 

Emmerich, M. 1978. Contribuçao ao 

conhecimento da tribo Cusparineae [sic] 

(Rutaceae). Rodriguesia 45: 223-307. 

Kaastra, R.C. 1982. Pilocarpinae (Rutaceae). 


Kallunki, J.A. 1990. An emended description of 

and new combinations in Raputia Aublet. 

Brittonia 42: 175-177. 

Kallunki, J.A. 1992. A revision of Erythrochiton 

s.l. Brittonia 44(2): 107-139. 

Kallunki, J.A. 1994. Revision of Raputia Aubl. 

(Cuspariinae, Rutaceae). Brittonia 46(4): 

279-295. 

Porter, D.M. & T.S. Elias. 1979. Rutaceae. In 

R.E. Woodson Jr., R.W. Schery, & 

collaborators, Flora of Panama. Ann. 


Waterman, P.G. 1975. New combinations in 

Zanthoxylum L. (1753). Taxon 24: 361-366. 

Amyris guianensis Aubl. [FG] 

Citrus spp. [species and cultivars cultivated and perhaps 

escaping] 

Conchocarpus acuminatus (Pilger) Kallunk & Pirani, comb. 

nov. ined. [FG] 

Conchocarpus fanshawei (Sandw.) Kallunki & Pirani, comb. 


Conchocarpus guianensis (Pulle) Kallunki & Pirani, comb. 


Conchocarpus pentagynus (St. Hil.) Kallunki & Pirani, comb. 


Conchocarpus toxicarius (Engl.) Kallunki & Pirani, comb. 

nov. ined. [to be expected in GU] 

Conchocarpus ucayalinus (Huber) Kallunki & Pirani, comb. 


Ertela trifolia (L.) Kuntze [GU SU FG] [=Monnieria trifolia 

L.] 

Erythrochiton brasiliensis Nees & Mart. [GU SU FG] 

Esenbeckia cowanii Kaastra [FG] 

Esenbeckia grandiflora Mart. subsp. grandiflora [GU? SU 

FG] 

Esenbeckia pilocarpoides Kunth subsp. pilocarpoides [GU SU 

FG] 

Galipea davisii Sandw. [GU FG] 

Galipea trifoliata Aubl. [SU FG] 

Hortia regia Sandw. [GU] 

Metrodorea flavida Krause [SU] 

Murraya paniculata (L.) Jack [GU SU FG] [cultivated; 

naturalized?] 

Neoraputia paraensis (Ducke) Emmerich [FG] 

Pilocarpus demerarae Sandw. [GU] 

Pilocarpus microphyllus Stapf ex Wardleworth [SU] 

Pilocarpus racemosus Vahl subsp. racemosus [FG] 

Raputia aromatica Aubl. [FG] 

Raputia brevipedunculata Kallunki [FG] 

Rauia resinosa Nees & Mart. [GU SU] 

Rauia spicata Haye [SU] 

Raveniopsis ruellioides (Oliver) Cowan [GU]

190 


Spathelia ulei (Engl. ex Harms) Cowan & Brizicky [GU] 

Spiranthera guianensis Sandw. [GU] 

Spiranthera parviflora Sandw. [GU] 

Ticorea foetida Aubl. [SU FG] 

Ticorea longiflora DC. [SU FG] 

Ticorea pedicellata DC. [SU FG] 

Triphasia trifolia (Burm.) P. Wils. [GU FG] [cultivated and 

apparently escaped] 

Zanthoxylum apiculatum (Sandw.) Waterm. [GU] 

Zanthoxylum caribaeum Lam. [GU] 

Zanthoxylum ekmanii (Urban) B. Alain [FG] 

Zanthoxylum flavum Vahl [FG?] 

Zanthoxylum hermaphroditum Willd. [type "Guiana"] 

Zanthoxylum nigrum Mart. [SU?] 

Zanthoxylum pentandrum (Aubl.) R. Howard [GU SU FG] 

Zanthoxylum pterotum (Kunth) Waterm. [FG] 

Zanthoxylum rhoifolium Lam. [GU SU FG] 

RUTACEAE - SYNONYMS AND 


Almeidea guyanensis Pulle =Conchocarpus guianensis 

Angostura acuminata (Pilger) Albuquerque =Conchocarpus 

acuminatus 

Angostura fanshawei (Sandw.) Albuquerque =Conchocarpus 

fanshawei 

Angostura fontanesiana (St. Hil.) Albuquerque [not in 


Angostura pentagyna (St. Hil.) Albuquerque =Conchocarpus 

pentagynus 

Angostura toxicaria (Spruce ex Engl.) Albuquerque 

=Conchocarpus toxicarius 

Angostura ucayalina (Huber) Albuquerque =Conchocarpus 

ucayalinus 

Cusparia fanshawei Sandw. =Conchocarpus fanshawei 

Cusparia toxicaria Spruce ex Engl. =Conchocarpus toxicarius 

Diomma ulei Engl. ex Harms =Spathelia ulei 

Fagara apiculata Sandw. =Zanthoxylum apiculatum 

Fagara caribaea (Lam.) Krug & Urban =Zanthoxylum 

caribaeum 

Fagara flava Krug & Urban =Zanthoxylum flavum 

Fagara guianensis Lam. =Zanthoxylum hermaphroditum 

Fagara pentandra Aubl. =Zanthoxylum pentandrum 

Fagara perrottetii DC. =Zanthoxylum rhoifolium 

Fagara rhoifolia (Lam.) Engl. =Zanthoxylum rhoifolium 

Galipea aromatica (Aubl.) Spreng. =Raputia aromatica 

Galipea fissa Miq. =Galipea trifoliata 

Monnieria trifolia L. =Ertela trifolia 

Murraya exotica L. =Murraya paniculata 

Pilocarpus latifolius St. Hil. =Pilocarpus racemosus subsp. 

racemosus 

Raputia paraensis Ducke =Neoraputia paraensis 

Rauia prancei Rodrigues & Silva [not in Guianas] 

Rauia racemosa Nees & Mart. [not in Guianas] 

Rauia ulei Krause =Conchocarpus ucayalinus 

Ravenia ruellioides Oliver =Raveniopsis ruellioides 

Sciuris aromatica (Aubl.) Spreng. =Raputia aromatica 

Zanthoxylum belizense Lund. [specimens re-determined as 

Zanthoxylum ekmanii] 

Zanthoxylum lucidum St. Hil. [specimens re-determined as 

Zanthoxylum pentandrum] 

Zanthoxylum microcarpum Griseb. [specimens re-determined 

as Zanthoxylum rhoifolium] 

Zanthoxylum perrottetii DC. =Zanthoxylum rhoifolium 

SPERMATOPHYTA: Rutaceae - Sapindaceae 

REFERENCES: 

SABIACEAE 

Barneby, R.C. 1972. Meliosmaceae - 

Ophiocaryon. In B. Maguire & collaborators, 



Steyermark, J.A. & A.H. Gentry. 1992. 

Sabiaceae. In G. Morillo, Flora de Venezuela 

5(1): 203-220. 

Ophiocaryon chironectes Barneby [FG?] 

Ophiocaryon heterophyllum (Benth.) Urb. [GU?] [records 

from Guyana probably represent Ophiocaryon maguirei] 

Ophiocaryon maguirei Barneby [GU] 

Ophiocaryon paradoxum Rob. Schomb. [GU FG?] 

SAPINDACEAE 

contributed by P. Acevedo R. (1992) 

REFERENCES: 

Kramer, K.U. 1973 ("1972"). Studies in the New 

World Sapindaceae. I. Notes on the genus 

Talisia Aublet. Acta Bot. Neerl. 21: 671-678. 

Kramer, K.U. 1976. Sapindaceae (additions and 

corrections). In J. Lanjouw & A.L. Stoffers, 


Radlkofer, L. 1931-1934. Sapindaceae. In A. 

Engler, Das Pflanzenreich IV,165 Bd. I: 1- 

1018, Bd. II: 1019-1539. 

Uittien, H. 1937. Sapindaceae. In A. Pulle, 


Allophylus edulis (St. Hil.) Radlk. [GU] 

Allophylus punctatus (Poepp. & Endl.) Radlk. [SU] 

Allophylus racemosus Swartz [GU] 

Allophylus robustus Radlk. [GU FG] 

Allophylus sp. cf. acutatus Radlk. [FG] 

Cardiospermum corindum L. [GU?] 

Cardiospermum grandiflorum Swartz [GU] 

Cardiospermum halicacabum L. [GU SU FG] 

Cupania americana L. [SU FG?] 

Cupania diphylla Vahl [SU FG] 

Cupania hirsuta Radlk. [GU SU FG] 

Cupania hispida Radlk. [FG] 

Cupania lachnocarpa Radlk. [FG] 

Cupania lanuginosa Sagot ex Radlk. [FG] 

Cupania scrobiculata L.C. Rich. [GU SU FG] [var. 

scrobiculata?] 

Cupania scrobiculata L.C. Rich. var. reticulata (Camb.) Radlk. 

[GU] 

Dodonaea viscosa Jacq. [GU SU FG] 

Matayba adenanthera Radlk. [FG] 

Matayba arborescens (Aubl.) Radlk. [GU SU FG] 

Matayba camptoneura Radlk. [GU SU]


Matayba elegans Radlk. [GU? FG] 

Matayba guianensis Aubl. [GU? SU FG] 

Matayba inelegans Spruce ex Radlk. [GU FG] 

Matayba laevigata (Miq.) Radlk. [SU FG] 

Matayba macrolepis Radlk. [GU] 

Matayba macrostylis Radlk. [GU FG] 

Matayba oligandra Sandw. [GU FG] 

Matayba opaca Radlk. var. fallax (Radlk.) Uitt. [GU SU] 

Matayba opaca Radlk. var. opaca [GU SU FG?] 

Matayba ptariana Steyerm. [GU] 

Matayba purgans Radlk. [GU SU? FG] 

Matayba scrobiculata (Kunth) Radlk. [FG] 

Melicoccus bijugatus Jacq. [GU SU FG?] [cultivated or 


Paullinia acuminata Uittien [SU FG?] 

Paullinia alata (Ruiz & Pav.) G. Don [SU FG?] 

Paullinia anisoptera Turcz. [GU] 

Paullinia anodonta Radlk. [SU FG] 

Paullinia arenicola Gleason [GU] 

Paullinia bernhardi Uittien [SU] 

Paullinia caloptera Radlk. [GU? SU FG] 

Paullinia cambessedesii Triana & Planch. [FG] 

Paullinia capreolata (Aubl.) Radlk. [GU SU FG] 

Paullinia conduplicata Radlk. [GU FG] 

Paullinia cururu L. [GU SU] 

Paullinia curvicuspis Radlk. [FG] 

Paullinia dasygonia Radlk. [GU SU FG] 

Paullinia fasciculata Radlk. [FG] 

Paullinia fibulata Rich. ex Juss. [FG] 

Paullinia fuscescens Kunth [GU SU] 

Paullinia grandifolia Benth. ex Radlk. [SU] 

Paullinia hitchcockii Gleason [GU] 

Paullinia imberbis Radlk. [FG] 

Paullinia ingaefolia Rich. ex Juss. [GU FG] 

Paullinia latifolia Benth. ex Radlk. [GU? SU FG?] 

Paullinia leiocarpa Griseb. [GU FG] 

Paullinia novemalata Uittien [SU] 

Paullinia pinnata L. [GU SU FG] 

Paullinia plagioptera Radlk. [SU FG?] 

Paullinia rubiginosa Camb. [FG] 

Paullinia rufescens Rich ex Juss. [GU SU FG] 

Paullinia rugosa Benth. ex Radlk. [GU] 

Paullinia sphaerocarpa Rich. ex Juss. [GU? SU FG] 

Paullinia spicata Benth. [SU FG] 

Paullinia stellata Radlk. [FG] 

Paullinia stenopetala Sagot [FG] 

Paullinia subnuda Radlk. [GU] 

Paullinia tetragona Aubl. [FG] 

Paullinia tricornis Radlk. [SU FG] 

Paullinia venosa Radlk. [SU FG] 

Paullinia verrucosa Radlk. [GU FG] 

Paullinia xestophylla Radlk. [GU] 

Pentascyphus thyrsiflorus Radlk. [FG] 

Porocystis toulicioides Radlk. [GU FG] 

Pseudima frutescens (Aubl.) Radlk. [GU SU FG] 

Pseudima pallidum Radlk. [SU FG] 

Sapindus saponaria L. var. inaequalis (DC.) Radlk. [GU? SU 

FG?] 

Scyphonichium multiflorum (Mart.) Radlk. [FG] 

Serjania adusta Radlk. [GU] 

Serjania caracasana (Jacq.) Willd. [GU FG] 

Serjania chartacea Radlk. [GU FG] 

Serjania exarata Radlk. [GU] 

Serjania grandifolia Sagot [SU FG] 

Serjania membranacea Splitg. [GU SU FG] 

Serjania oblongifolia Radlk. [GU SU FG] 

Serjania paucidentata DC. [GU SU FG] 

Serjania pedicellaris Radlk. [SU FG] 

Serjania prancei Acevedo R., sp. nov. ined. [GU] 

Serjania pyramidata Radlk. [FG] 

Talisia acuminata Radlk. [FG] 

Talisia acutifolia Radlk. [SU] 

Talisia carinata Radlk. [SU FG] 

Talisia clathrata Radlk. [FG] 

Talisia elephantipes Sandw. [GU SU FG] 

Talisia esculenta (St. Hil.) Radlk. [GU] 

Talisia eximia K.U. Kramer [SU FG] 

Talisia furfuracea Sandw. [GU SU FG] 

Talisia guianensis Aubl. [GU SU FG] 

Talisia hemidasya Radlk. [GU SU FG] 

Talisia hexaphylla Vahl [GU FG] 

Talisia longifolia (Benth.) Radlk. [SU FG] 

Talisia megaphylla Sagot ex Radlk. [GU SU FG] 

Talisia micrantha Radlk. [SU FG] 

Talisia microphylla Uittien [SU FG] 

Talisia mollis Kunth ex Camb. [SU FG] 

Talisia olivaeformis (Kunth) Radlk. [GU FG] 

Talisia pachycarpa Radlk. [FG] 

Talisia pedicellaris Sagot ex Radlk. [SU FG] 

Talisia pilosula Sagot ex Radlk. [FG] 

Talisia praealta (Sagot) Radlk. [SU FG] 

Talisia pulverulenta Radlk. [FG] 

Talisia reticulata Radlk. [SU FG?] 

Talisia retusa Cowan [GU SU FG?] 

Talisia simaboides K.U. Kramer [SU FG] 

Talisia sp. cf. cupularis Radlk. [FG] 

Talisia sp. cf. nervosa Radlk. [FG] 

Talisia squarrosa Radlk. [GU SU FG] 

Talisia sylvatica (Aubl.) Radlk. [GU SU FG] 

Thinouia myriantha Triana & Planch. [GU SU? FG] 

Toulicia elliptica Radlk. [GU? SU FG?] 

Toulicia guianensis Aubl. [GU SU FG] 

Toulicia patentinervis Radlk. [GU SU] 

Toulicia pulvinata Radlk. [GU SU FG?] 

Urvillea ulmacea Kunth [GU SU? FG] 

Vouarana guianensis Aubl. [GU? SU FG] 

SAPINDACEAE - SYNONYMS AND 


Allophylus occidentalis (Swartz) Radlk. =Allophylus 

racemosus 

Enourea capreolata Aubl. =Paullinia capreolata 

Matayba fallax Radlk. =Matayba opaca var. fallax 

Matayba miquelii Uittien =Matayba laevigata 

Melicocca bijugata L. =Melicoccus bijugatus 

Racaria sylvatica Aubl. =Talisia sylvatica 

Sapindus arborescens Aubl. =Matayba arborescens 

Sapindus frutescens Aubl. =Pseudima frutescens 

Talisia glandulifera Steyerm. =Talisia hemidasya 

Talisia macrophylla (Mart.) Radlk. [dubious name] 

REFERENCES: 

SAPOTACEAE 

SPERMATOPHYTA: Sapindaceae - Sapotaceae 

191

192 

SPERMATOPHYTA: Sapotaceae 


[for a more extensive bibliography, see 

Pennington 1990] 

Aubréville, A. 1972. Sapotaceae. In B. Maguire 


Highland part IX. Mem. New York Bot. 

Gard. 23: 1-832. 

Eyma, P.J. 1936. Sapotaceae. In A. Pulle, Flora 


Pennington, T.D. 1990. Sapotaceae. Flora 

Neotropica Monograph 52: 1-770. 

Chromolucuma baehniana Monachino [GU] 

Chrysophyllum amazonicum Pennington [GU] 

Chrysophyllum argenteum Jacq. subsp. auratum (Miq.) 

Pennington [GU SU FG] 

Chrysophyllum argenteum Jacq. subsp. nitidum (G. Mey.) 

Pennington [GU SU FG] 

Chrysophyllum cainito L. [GU SU FG] [cultivated, 

naturalized] 

Chrysophyllum cuneifolium (Rudge) A. DC. [SU FG] 

Chrysophyllum eximium Ducke [SU] [to be expected in FG] 

Chrysophyllum lucentifolium Cronq. subsp. pachycarpum 

Pires & Pennington [FG] 

Chrysophyllum manaosense (Aubr.) Pennington [GU SU] 

Chrysophyllum pomiferum (Eyma) Pennington [GU SU FG] 

Chrysophyllum prieurii A. DC. [SU FG] 

Chrysophyllum sanguinolentum (Pierre) Baehni subsp. balata 

(Ducke) Pennington [GU] 

Chrysophyllum sanguinolentum (Pierre) Baehni subsp. 

sanguinolentum [GU SU FG] 

Chrysophyllum sparsiflorum Klotzsch ex Miq. [GU SU? FG] 

Chrysophyllum venezuelanense (Pierre) Pennington [FG] 

Diploön cuspidatum (Hoehne) Cronquist [GU FG] 

Ecclinusa dumetorum (Baehni) Pennington [SU FG?] 

Ecclinusa guianensis Eyma [GU SU FG] 

Ecclinusa lanceolata (Mart. & Eichl.) Pierre [GU FG] 

Ecclinusa psilophylla Sandw. [GU SU] 

Ecclinusa ramiflora Mart. [SU FG] [to be expected in GU] 

Ecclinusa ulei (Krause) Gilly ex Cronquist [Mt. Roraima] 

Elaeoluma glabrescens (Mart. & Eichl.) Aubr. [GU] 

Elaeoluma nuda (Baehni) Aubr. [SU] [to be expected in GU] 

Elaeoluma schomburgkiana (Miq.) Baill. [GU] 

Manilkara bidentata (A. DC.) Chevalier subsp. bidentata [GU 

SU FG] 

Manilkara bidentata (A. DC.) Chevalier subsp. surinamensis 

(Miq.) Pennington [GU SU FG] 

Manilkara excelsa (Ducke) Standl. [GU?] 

Manilkara huberi (Ducke) Chevalier [GU SU FG] 

Manilkara paraensis (Huber) Standl. [FG] 

Manilkara pubicarpa Monachino [GU] 

Manilkara zapota (L.) van Royen [widely cultivated and 

escaping] 

Micropholis acutangula (Ducke) Eyma [FG] 

Micropholis cayennensis Pennington [FG] 

Micropholis egensis (A. DC.) Pierre [GU? SU FG] 

Micropholis gardneriana (A. DC.) Pierre [GU?] 

Micropholis guyanensis (A. DC.) Pierre subsp. duckeana 

(Baehni) Pennington [FG] 

Micropholis guyanensis (A. DC.) Pierre subsp. guyanensis 

[GU SU FG] 

Micropholis longipedicellata Aubr. [GU SU FG] 

Micropholis melinoniana Pierre [SU FG] 

Micropholis mensalis (Baehni) Aubr. [GU SU FG] 

Micropholis obscura Pennington [GU FG] 

Micropholis porphyrocarpa (Baehni) Monachino [GU FG] 

Micropholis sanctae-rosae (Baehni) Pennington [FG] 

Micropholis splendens Gilly ex Aubr. [FG?] 

Micropholis venulosa (Mart. & Eichl.) Pierre [GU SU FG] 

Mimusops spp. [cultivated or escaped from cultivation] 

Pouteria ambelaniifolia (Sandw.) Pennington [GU FG] 

Pouteria aubrevillei Bernardi [FG] 

Pouteria bangii (Rusby) Pennington [SU FG] [to be expected 

in GU] 

Pouteria benai (Aubr. & Pellegr.) Pennington [FG] 

Pouteria bilocularis (Winkler) Baehni [GU FG] 

Pouteria brachyandra (Aubr. & Pellegr.) Pennington [GU SU 

FG] 

Pouteria caimito (Ruiz & Pav.) Radlk. [GU SU FG] 

Pouteria campanulata Baehni [possibly to be expected in GU] 

Pouteria cayennensis (A. DC.) Eyma [GU FG] 

Pouteria cicatricata Pennington [FG] 

Pouteria cladantha Sandw. [GU SU FG] 

Pouteria congestifolia Pilz [FG] 

Pouteria coriacea (Pierre) Pierre [GU SU FG] 

Pouteria crassiflora Pires & Pennington [possibly to be 

expected in FG] 

Pouteria cuspidata (A. DC.) Baehni subsp. cuspidata [GU SU] 

[to be expected in FG] 

Pouteria cuspidata (A. DC.) Baehni subsp. dura (Eyma) 

Pennington [GU SU FG?] 

Pouteria cuspidata (A. DC.) Baehni subsp. robusta (Mart. & 

Eichl.) Pennington [GU? SU FG] 

Pouteria decorticans Pennington [FG] 

Pouteria deliciosa Pennington [FG] 

Pouteria durlandii (Standl.) Baehni subsp. durlandii [GU] 

Pouteria egregia Sandw. [GU SU FG] 

Pouteria elegans (A. DC.) Baehni [GU] 

Pouteria engleri Eyma [GU SU FG] 

Pouteria eugeniifolia (Pierre) Baehni [GU SU FG] 

Pouteria filipes Eyma [GU SU FG] 

Pouteria fimbriata Baehni [GU FG] 

Pouteria franciscana Baehni [FG] 

Pouteria glomerata (Miq.) Radlk. subsp. glomerata [GU? SU? 

FG] 

Pouteria glomerata (Miq.) Radlk. subsp. stylosa (Pierre) 

Pennington [SU] 

Pouteria gonggrijpii Eyma [SU FG] 

Pouteria gongylocarpa Pennington, sp. nov. ined. [FG] 

Pouteria grandis Eyma [GU SU FG] 

Pouteria guianensis Aubl. [GU SU FG] 

Pouteria hispida Eyma [GU SU FG] 

Pouteria jariensis Pires & Pennington [SU? FG] 

Pouteria kaieteurensis Pennington [GU] 

Pouteria laevigata (Mart.) Radlk. [FG] 

Pouteria latianthera Pennington [to be expected in FG] 

Pouteria macrocarpa (Mart.) Dietr. [FG?] 

Pouteria macrophylla (Lam.) Eyma [GU? SU FG] 

Pouteria manaosensis (Aubr. & Pellegr.) Pennington [SU] 

Pouteria melanopoda Eyma [GU SU FG] 

Pouteria oblanceolata Pires [FG] 

Pouteria penicillata (Baehni) Aubr. [GU] 

Pouteria pullei Eyma [SU] [imperfectly known species] 

Pouteria reticulata (Engl.) Eyma subsp. reticulata [SU FG] [to 


Pouteria reticulata (Engl.) Eyma subsp. surinamensis 

Pennington [GU SU] 

Pouteria retinervis Pennington [FG]


Pouteria rigida (Mart. & Eichl.) Radlk. subsp. rigida [GU] 

Pouteria rigida (Mart. & Eichl.) Radlk. subsp. tomentosa 

(Aubr.) Pennington [GU] 

Pouteria rodriguesiana Pires & Pennington [SU FG] 

Pouteria sagotiana (Baill.) Eyma [GU SU FG] 

Pouteria singularis Pennington [FG] 

Pouteria speciosa (Ducke) Baehni [GU SU FG] 

Pouteria surumuensis Baehni [GU] 

Pouteria tenuisepala Pires & Pennington [FG] 

Pouteria torta (Mart.) Radlk. subsp. glabra Pennington [GU 

FG] 

Pouteria torta (Mart.) Radlk. subsp. tuberculata (Sleumer) 

Pennington [FG] 

Pouteria trigonosperma Eyma [GU SU] 

Pouteria venosa (Mart.) Baehni subsp. amazonica Pennington 

[GU SU FG] 

Pouteria virescens Baehni [GU FG] 

Pouteria williamii (Aubr. & Pellegr.) Pennington [FG] 

Pradosia beardii (Monachino) Pennington [GU] 

Pradosia cochlearia (Lecomte) Pennington subsp. cochlearia 

[FG] 

Pradosia huberi (Ducke) Ducke [FG] 

Pradosia ptychandra (Eyma) Pennington [SU FG] 

Pradosia schomburgkiana (A. DC.) Cronq. subsp. 

schomburgkiana [GU SU] 

Pradosia surinamensis (Eyma) Pennington [GU SU FG] 

Pradosia verticillata Ducke [FG] 

Sarcaulus brasiliensis (A. DC.) Eyma subsp. brasiliensis [GU 

SU FG] 

SAPOTACEAE - SYNONYMS AND 


Achras balata Aubl. =Mimusops sp.[described from cultivated 

material] 

Achras zapota L. =Manilkara zapota 

Achrouteria pomifera Eyma =Chrysophyllum pomiferum 

Amorphospermum schomburgkianum (Miq.) Baehni 

=Elaeoluma schomburgkiana 

Bumelia amazonica Krause =Pouteria cuspidata subsp. 

cuspidata 

Bumelia cuneifolia Rudge =Chrysophyllum cuneifolium 

Bumelia nervosa Vahl =Pouteria macrophylla 

Calocarpum cooperi (Cronquist) Lundell =Pouteria torta 

subsp. tuberculata 

Calocarpum mammosum (L.) Pierre =Pouteria sapota 

Chrysophyllum alnifolium Engl., non Baker =Pouteria engleri 

Chrysophyllum ambelaniifolium Sandw. =Pouteria 

ambelaniifolia 

Chrysophyllum auratum Miq. =Chrysophyllum argenteum 

subsp. auratum 

Chrysophyllum auratum Miq. var. majus Miq. 

=Chrysophyllum argenteum subsp. auratum 

Chrysophyllum beardii Monachino =Pradosia beardii 

Chrysophyllum brasiliense A. DC. =Sarcaulus brasiliensis 

Chrysophyllum brasiliense A. DC. var. minus Miq. 

=Sarcaulus brasiliensis subsp. brasiliensis 

Chrysophyllum cayennense A. DC. =Pouteria cayennensis 

Chrysophyllum cochlearium Lecomte =Pradosia cochlearia 

Chrysophyllum dumetorum Baehni =Ecclinusa dumetorum 

Chrysophyllum emarginatum Klotzsch ex Rich. Schomb. 

[name of uncertain application] 

Chrysophyllum guianense (Eyma) Baehni =Ecclinusa 

guianensis 

193 

Chrysophyllum guyanense Klotzsch ex Miq. =Chrysophyllum 

argenteum subsp. auratum 

Chrysophyllum macoucou Aubl. =Aubletella macoucou 

(Aubl.) Pierre [Incertae sedis; excluded from Sapotaceae 

by Pennington] 

Chrysophyllum macrophyllum Mart., non Lam. =Sarcaulus 

brasiliensis subsp. brasiliensis 

Chrysophyllum macrophyllum Lam. =Pouteria macrophylla 

Chrysophyllum melinonii Engl. =Micropholis guyanensis 

subsp. guyanensis 

Chrysophyllum monachinoanum Steyerm. =Pradosia beardii 

Chrysophyllum nitidum G. Mey. =Chrysophyllum argenteum 

subsp. nitidum 

Chrysophyllum psilophyllum A. DC. =Chrysophyllum 

argenteum subsp. nitidum 

Chrysophyllum psilophyllum (Sandw.) Baehni =Ecclinusa 

psilophylla 

Chrysophyllum pyriforme Willd. =Aubletella macoucou 

(Aubl.) Pierre [Incertae sedis; excluded from Sapotaceae 

by Pennington] 

Chrysophyllum rufocupreum Ducke =Chrysophyllum 

eximium 

Chrysophyllum schomburgkianum A. DC. =Pradosia 


Chrysophyllum sericeum A. DC. =Chrysophyllum argenteum 

subsp. auratum 

Chrysophyllum sericeum A. DC. var. acutifolium Miq. 


Chrysophyllum sericeum A. DC. var. obtusifolium Miq. 


Chrysophyllum sessiliflorum Poir. =Pouteria cayennensis 

Chrysophyllum ulei Krause =Ecclinusa ulei 

Cynodendron nitidum (G. Mey.) Aubr. =Chrysophyllum 

argenteum subsp. nitidum 

Ecclinusa balata Ducke =Chrysophyllum sanguinolentum 

subsp. balata 

Ecclinusa cochlearia (Lecomte) Aubr. =Pradosia cochlearia 

Ecclinusa cuneifolia (Rudge) Aubr. =Chrysophyllum 

cuneifolium 

Ecclinusa prieurii (A. DC.) Aubr. =Chrysophyllum prieurii 

Ecclinusa sanguinolenta (Pierre) Engl. =Chrysophyllum 

sanguinolentum 

Englerella macrocarpa Pierre =Pouteria speciosa 

Englerella speciosa Ducke =Pouteria speciosa 

Eremoluma sagotiana Baill. =Pouteria sagotiana 

Eremoluma williamii Aubr. & Pellegr. =Pouteria williamii 

Franchetella gonggrijpii (Eyma) Aubr. =Pouteria gonggrijpii 

Ichthyophora elegans (Miq.) Baehni =Pouteria elegans 

Ichthyophora leptocarpa (Miq.) Baehni =Pouteria elegans 

Labatia laevigata Mart. =Pouteria laevigata 

Labatia pedunculata Willd. =Pouteria guianensis 

Lucuma duckei Huber =Pouteria venosa subsp. amazonica 

Lucuma glabrescens Mart. & Eichl. =Elaeoluma glabrescens 

Lucuma jenmanii Pittier =Pouteria torta subsp. glabra 

Lucuma pulverulenta Mart. & Eichl. =Pouteria cayennensis 

Lucuma rigida Mart. & Eichl. =Pouteria rigida 

Lucuma sagotiana (Baill.) Engl. =Pouteria sagotiana 

Lucuma sericea Krause =Pouteria surumuensis 

Lucuma speciosa Ducke =Pouteria speciosa 

Manilkara amazonica (Hub.) Chev. =Manilkara bidentata 

subsp. surinamensis 

Manilkara balata (Pierre) Dubard =Manilkara bidentata subsp. 

bidentata 

Manilkara surinamensis (Miq.) Dubard =Manilkara bidentata 

subsp. surinamensis 

SPERMATOPHYTA: Sapotaceae

194 

SPERMATOPHYTA: Sapotaceae 


Meioluma guianensis Baill. =Micropholis venulosa 

Micropholis burchelliana Pierre =Micropholis gardneriana 

Micropholis calophylloides Pierre =Micropholis venulosa 

Micropholis casiquiarensis Aubr. [not in Guianas] 

Micropholis cowanii Aubr. =Micropholis guyanensis subsp. 

guyanensis 

Micropholis eugeniifolia Pierre =Pouteria eugeniifolia 

Micropholis melinonii (Engl.) Mennega et al. =Micropholis 

guyanensis subsp. guyanensis 

Micropholis mucronata Pierre =Micropholis venulosa 

Micropholis paraensis Huber ex Eyma =Micropholis 

acutangula 

Micropholis robusta (Mart. & Eichl.) Mennega et al. =Pouteria 

cuspidata subsp. robusta 

Mimusops balata (Aubl.) C.F. Gaertn. [see Achras balata] 

Mimusops balata Pierre var. gutta Pierre =Manilkara bidentata 

subsp. bidentata 

Mimusops balata Pierre var. melinonis Pierre =Manilkara 

bidentata subsp. bidentata 

Mimusops bidentata A. DC. =Manilkara bidentata 

Mimusops huberi Ducke =Manilkara huberi 

Mimusops paraensis Huber =Manilkara paraensis 

Mimusops surinamensis Miq. =Manilkara bidentata subsp. 

surinamensis 

Myrtiluma eugeniifolia (Pierre) Aubr. =Pouteria eugeniifolia 

Nemaluma engleri (Eyma) Aubr. & Pellegr. =Pouteria engleri 

Neopometia ptychandra (Eyma) Aubr. =Pradosia ptychandra 

Neoxythece cladantha (Sandw.) Aubr. =Pouteria cladantha 

Neoxythece dura (Eyma) Aubr. & Pellegr. =Pouteria cuspidata 

subsp. dura 

Neoxythece robusta (Mart. & Eichl.) Aubr. & Pellegr. 

=Pouteria cuspidata subsp. robusta 

Neoxythece roraimaensis Aubr. =Pouteria rigida subsp. rigida 

Neoxythece schulzii Aubr. =Pouteria cuspidata subsp. dura 

Neoxythece tomentosa Aubr. =Pouteria rigida subsp. 

tomentosa 

Oxythece ambelaniifolia (Sandw.) Cronq. =Pouteria 


Oxythece dura (Eyma) Gilly =Pouteria cuspidata subsp. dura 

Oxythece elegans (A. DC.) Urb. =Pouteria elegans 

Oxythece guianensis Lec. =Pouteria guianensis? 

Oxythece leptocarpa Miq. =Pouteria elegans 

Oxythece rigida (Mart. & Eichl.) Monach. =Pouteria rigida 

Oxythece schomburgkiana (Miq.) Cronq. =Elaeoluma 


Paralabatia acutangula (Ducke) Baehni =Micropholis 

acutangula 

Paramicropholis acutangula (Ducke) Aubr. & Pellegr. 

=Micropholis acutangula 

Planchonella elegans (A. DC.) Baehni =Pouteria elegans 

Planchonella guianensis van Royen =Chrysophyllum 

pomiferum 

Planchonella melinonii (Engl.) Baehni =Micropholis 


Platyluma calophylloides (Pierre) Baill. =Micropholis 

venulosa 

Podoluma benai Aubr. & Pellegr. =Pouteria benai 

Pouteria acutangula (Ducke) Baehni =Micropholis acutangula 

Pouteria benoistii Aubr. =Pouteria hispida 

Pouteria burchelliana (Pierre) Baehni =Micropholis 

gardneriana 

Pouteria demerarae Sandw. =Pouteria guianensis 

Pouteria duckeana Baehni =Micropholis guyanensis subsp. 

duckeana 

Pouteria dura Eyma =Pouteria cuspidata subsp. dura 

Pouteria echinocarpa W. Rodrigues =Pouteria torta subsp. 

tuberculata 

Pouteria elegans var. micrantha (Miq.) Baehni =Pouteria 

cuspidata subsp. cuspidata 

Pouteria jenmanii (Pittier) Sandw. =Pouteria torta subsp. 

glabra 

Pouteria lucentifolia (Standl.) Baehni =Chrysophyllum 

venezuelanense 

Pouteria melinoniana (Pierre) Baehni =Micropholis 

melinoniana 

Pouteria melinonii (Engl.) Baehni =Micropholis guyanensis 

subsp. guyanensis 

Pouteria mensalis Baehni =Micropholis mensalis 

Pouteria minutiflora (Britton) Sandw. =Pouteria coriacea 

Pouteria multiflora (A. DC.) Eyma [not in Guianas] 

Pouteria nuda Baehni =Elaeoluma nuda 

Pouteria obidensis Huber =Pouteria guianensis 

Pouteria pedunculata (Willd.) Poir. =Pouteria guianensis 

Pouteria pomifera (Eyma) Baehni =Chrysophyllum pomiferum 

Pouteria porphyrocarpa Baehni =Micropholis porphyrocarpa 

Pouteria ptychandra Eyma =Pradosia ptychandra 

Pouteria robusta (Mart. & Eichl.) Eyma =Pouteria cuspidata 

subsp. robusta 

Pouteria robusta var. longifolia Eyma =Pouteria cuspidata 


Pouteria rufa (Mart. & Eichl. ex Mart.) Lemée =Micropholis 

guyanensis 

Pouteria scytalophora Eyma =Pouteria bangii 

Pouteria sericea (Aiton) Baehni [Australian species, recorded 

from GU in error] 

Pouteria suffulta Baehni =Pouteria durlandii subsp. durlandii 

Pouteria surinamensis Eyma =Pradosia surinamensis 

Pouteria venulosa (Mart. & Eichl.) Baehni =Micropholis 

venulosa 

Prieurella cuneifolia (Rudge) Pierre ex Aubr. =Chrysophyllum 

cuneifolium 

Prieurella prieurii (A. DC.) Aubr. =Chrysophyllum prieurii 

Pseudocladia minutiflora (Britton) Aubr. =Pouteria coriacea 

Pseudocladia scytalophora (Eyma) Aubr. =Pouteria bangii 

Pseudolabatia filipes (Eyma) Aubr. =Pouteria filipes 

Pseudolabatia penicillata (Baehni) Aubr. =Pouteria penicillata 

Pseudoxythece ambelaniifolia (Sandw.) Aubr. =Pouteria 


Radlkoferella brachyandra Aubr. & Pellegr. =Pouteria 

brachyandra 

Radlkoferella grandis (Eyma) Aubr. =Pouteria grandis 

Radlkoferella trigonosperma (Eyma) Aubr. =Pouteria 

trigonosperma 

Radlkoferella venosa (Mart.) Aubr. =Pouteria venosa 

Ragala sanguinolenta (Pierre) Aubr. =Chrysophyllum 

sanguinolentum 

Ragala ulei (Krause) Aubr. =Ecclinusa ulei 

Richardella cayennensis (A. DC.) Aubr. =Pouteria 

cayennensis 

Richardella cladantha (Sandw.) Baehni =Pouteria cladantha 

Richardella macrophylla (Lam.) Aubr. =Pouteria macrophylla 

Richardella surumuensis (Baehni) Aubr. =Pouteria 

surumuensis 

Richardella trigonosperma (Eyma) Baehni =Pouteria 

trigonosperma


Sandwithiodoxa egregia (Sandw.) Aubr. =Pouteria egregia 

Sideroxylon acutangulum Ducke =Micropholis acutangula 

Sideroxylon calophylloides (Pierre) Engl. =Micropholis 

venulosa 

Sideroxylon cuspidatum A. DC. =Pouteria cuspidata subsp. 

cuspidata 

Sideroxylon cuspidatum var. crassifolium Miq. =Pouteria 

cuspidata subsp. dura 

Sideroxylon cuspidatum A. DC. var. ellipticum Miq. 

=Pouteria cuspidata subsp. cuspidata 

Sideroxylon egense A. DC. =Micropholis egensis 

Sideroxylon elegans A. DC. =Pouteria elegans 

Sideroxylon elegans var. micranthum Miq. =Pouteria 

cuspidata subsp. cuspidata 

Sideroxylon eugeniifolium (Pierre) Engl. =Pouteria 

eugeniifolia 

Sideroxylon guianense (Baill.) Engl. =Micropholis venulosa 

Sideroxylon guyanense A. DC. =Micropholis guyanensis 

Sideroxylon robustum Mart. & Eichl. =Pouteria cuspidata 


Sideroxylon rufum Mart. & Eichl. ex Mart. =Micropholis 


Sideroxylon rugosum Roem. & Schult. var. parviflorum Miq. 

=Micropholis melinoniana 

Sideroxylon venulosum Mart. & Eichl. =Micropholis venulosa 

Vitellaria pulverulenta (Mart. & Eichl.) Radlk. =Pouteria 

cayennensis 

SARRACENIACEAE 

contributed by K.M. Cameron (1992) 

REFERENCES: 

Maguire, B. 1978. Sarraceniaceae. In B. 



Bot. Gard. 29: 36-62. 

Heliamphora nutans Benth. [GU] 

REFERENCES: 

SCROPHULARIACEAE 

D'Arcy, W.G. 1979. Scrophulariaceae. In R.E. 



66: 173-272. 

Diaz-Miranda, D. 1991. Scrophulariaceae en la 

flora de Venezuela I. Introducción. Ernstia 

1(3): 95-108. 

Pennell, F.W. 1940. Some new species of 

Scrophulariaceae from Guiana. Not. Nat. 

Acad. Nat. Sci. Phila. 46: 1-6. 

Philcox, D. 1965. Revision of the New World 

species of Buchnera L. Kew Bull. 18: 

275-315. 

Turner, B.L. & C.C. Cowan. 1993. Taxonomic 

overview of Stemodia for South America. 

Phytologia 75(4): 281-324. 

195 

Achetaria guianensis Pennell [GU SU FG] 

Achetaria ocimoides (Cham. & Schlechtend.) Wettst. [GU SU 

FG] 

Achetaria scutellarioides (Benth.) Kuntze [GU FG] 

Agalinis hispidula (Mart.) D'Arcy [GU SU FG] 

Alectra aspera (Cham. & Schlechtend.) L.O. Williams [GU 

FG] 

Angelonia angustifolia Benth. [SU FG] [probably cultivated 

only] 

Angelonia biflora Benth. [GU] 

Angelonia salicariifolia Humb. & Bonpl. [GU SU] [probably 

cultivated only] 

Bacopa aquatica Aubl. [GU SU FG] 

Bacopa bacopoides (Benth.) Pulle [SU] 

Bacopa chamaedroides Wettst. [FG] 

Bacopa connata (Pennell) Pennell [GU] 

Bacopa gratioloides (Benth.) Chodat [GU] 

Bacopa laxiflora (Benth.) Edwall [GU SU] 

Bacopa monnieri (L.) Wettst. [FG] 

Bacopa repens (Sw.) Wettst. [GU] 

Bacopa salzmannii (Benth.) Edwall [GU] 

Bacopa sessiliflora (Benth.) Edwall [GU SU FG] 

Bacopa stricta (Schrad.) Edwall [GU] 

Benjaminia reflexa (Benth.) D'Arcy [GU SU FG] 

Buchnera longifolia Kunth [GU FG] 

Buchnera palustris (Aubl.) Spreng. [GU SU FG] 

Buchnera pusilla Kunth [GU] 

Buchnera rosea Kunth [GU SU] 

Buchnera weberbaueri Diels [GU] 

Capraria biflora L. [GU SU FG] 

Conobea aquatica Aubl. [GU SU FG] 

Lindernia brachyphylla Pennell [GU] 

Lindernia crustacea (L.) F. Mueller [GU SU FG] 

Lindernia diffusa (L.) Wettst. [GU SU FG] 

Lindernia dubia (L.) Pennell [GU SU FG] 

Lindernia microcalyx Pennell & Stehlé [FG] 

Lindernia rotundifolia (L.) Standl. & L.O. Williams [FG] 

[naturalized] 

Mecardonia procumbens (P. Mill.) Small [FG] 

Micranthemum umbrosum (Walt. ex Gmel.) Blake [GU SU] 

Russelia sarmentosa Jacq. [FG] 

Scoparia dulcis L. [GU SU FG] 

Stemodia pratensis (Aubl.) C. Cowan [GU FG] [=Stemodia 

foliosa Benth.] 

Stemodia verticillata (Mill.) Hassl. [GU SU] 

Torenia fournieri Linden ex Fourn. [GU] [probably cultivated 

or escaped from cultivation 

Torenia thouarsii (Cham. & Schlechtend.) Kuntze [GU SU] 

[naturalized] 

Vellosiella spathacea (Oliver) Melchior [GU?] [Mt. Roraima] 

SCROPHULARIACEAE - SYNONYMS AND 


Alectra brasiliensis Benth. =Alectra aspera 

Anisantherina hispidula (Mart.) Pennell ex N.L. Britton 

=Agalinis hispidula 

Bacopa ciliata (Pennell) Pennell =Bacopa salzmannii 

Bacopa reflexa (Benth.) Edwall =Benjaminia reflexa 

SPERMATOPHYTA: Sapotaceae - Scrophulariaceae

196 


Beyrichia ocimoides Cham. & Schlechtend. =Achetaria 

ocimoides 

Bramia monnieri (L.) Drake =Bacopa monnieri 

Buchnera capillaris Desv. =Lindernia crustacea 

Buchnera elongata Swartz =Buchnera longifolia [records may 

represent Buchnera pusilla] 

Buchnera virgata Kunth [name dubiously applied to Guianan 

specimens] 

Capraria integerrima Miq. =Sesamum orientale L. 

(Pedaliaceae) 

Curanga bivalvis (L.f.) Druce =Mendoncia bivalvis (L.f.) 

Merr. (Mendonciaceae) 

Drupina cristata L. =Mendoncia bivalvis (L.f.) Merr. 


Gerardia hispidula Mart. =Agalinis hispidula 

Herpestis ciliata Pennell =Bacopa salzmannii 

Herpestis connata Pennell =Bacopa connata 

Herpestis reflexa Benth. =Benjaminia reflexa 

Ilysanthes riparia Raf. =Lindernia dubia 

Ilysanthes rotundifolia Benth. =Lindernia rotundifolia 

Matourea pratensis Aubl. =Stemodia pratensis 

Melasma brasiliensis Chodat & Hassl. =Alectra aspera 

Melasma spathaceum Oliver =Vellosiella spathacea 

Mella laxiflora (Benth.) Pennell =Bacopa laxiflora 

Micranthemum orbiculatum Michx. =Micranthemum 

umbrosum 

Picria surinamensis Spreng. =Mendoncia bivalvis (L.f.) Merr. 


Piripea palustris Aubl. =Buchnera palustris 

Stemodia foliosa Benth. =Stemodia pratensis 

Stemodia parviflora Ait. =Stemodia verticillata 

Stemodia pusilla Benth. [name apparently misapplied to S. 

pratensis] 

Stemodia surinamensis Miq. [name of uncertain application] 

Vandellia crustacea (L.) Benth. =Lindernia crustacea 

Vandellia diffusa L. =Lindernia diffusa 

REFERENCES: 

SIMAROUBACEAE 

Cavalcante, P.B. 1983. Revisao taxonômica do 

gênero Simaba Aublet na América do Sul. 

Publ. Avulsas Mus. Goeldi 37: 1-85. 

Cronquist, A. 1944. Studies in the 

Simaroubaceae. II. The genus Simarouba. 


Cronquist, A. 1944. Studies in Simaroubaceae. 

III. The genus Simaba. Lloydia 7: 81-92. 

Cronquist, A. 1944. Studies in Simaroubaceae. 

IV. Resume of the American genera. 

Brittonia 5: 128-147. 

Cronquist, A. 1945. Additional notes on the 

Simaroubaceae. Brittonia 5: 469-470. 

Feuillet, C. 1983. Etudes sur les Simaroubaceae. 

II. Un Simaba nouveau de Guyane française 

dans la section Floribundae Engl.: S. morettii. 

Candollea 38(2): 745-750. 

Jansen-Jacobs, M.J. 1979. Simaroubaceae. In 

A.L. Stoffers & J.C. Lindeman, Flora of 

Suriname 5(1): 319-330. 

Nooteboom, H.P. 1962. Generic delimitation in 

Simaroubaceae tribus Simaroubeae and a 

SPERMATOPHYTA: Scrophulariaceae - Simaroubaceae 

conspectus of the genus Quassia L. Blumea 

11: 509-528. 

Thomas, W.W. 1984. A new species of Simaba 

(Simaroubaceae) from Pará, Brazil, with a key 

to the species north of the Amazon River. 

Brittonia 36(3): 244-247. 

Thomas, W.W. 1985. The Simaba guianensis 

complex in northern South America. Acta 

Amazonica 15(1-2) suppl.: 71-79. 

Picramnia guianensis (Aubl.) Jansen-Jacobs [GU SU FG] 

Picramnia longissima Tul. [GU] [=Picramnia latifolia Tul.?] 

Picramnia macrostachys Klotzsch ex Engl. [GU SU FG] 

[=Picramnia latifolia Tul.?] 

Picramnia sellowii Planch. [GU FG] 

Picramnia spruceana Engl. [GU SU FG] 

Picrasma excelsa (Swartz) Planch. [FG?] 

Picrolemma pseudocoffea Ducke [FG] 

Quassia amara L. [GU SU FG] [cultivated, naturalized] 

Simaba cedron Planch. [GU SU FG] [cultivated, naturalized] 

Simaba guianensis Aubl. subsp. ecaudata (Cronquist) 

Cavalcante [GU SU FG] 

Simaba guianensis Aubl. subsp. guianensis [GU SU FG] 

Simaba monophylla (Oliv.) Cronquist [GU] 

Simaba morettii Feuillet [FG] 

Simaba multiflora Adr. Juss. [GU SU FG] 

Simaba orinocensis Kunth [SU FG] [to be expected in GU] 

Simaba polyphylla (Cavalcante) W. Thomas [FG] 

Simarouba amara Aubl. [GU SU FG] 

SIMAROUBACEAE - SYNONYMS AND 


Aeschrion excelsa (Swartz) Kuntze =Picrasma excelsa 

Aruba guianensis Aubl. =Simaba guianensis 

Picramnia latifolia Tul. [see Picramnia macrostachys, 

Picramnia longissima] 

Picramnia tariri DC. =Picramnia guianensis 

Quassia cedron (Planch.) Baill. =Simaba cedron 

Quassia cuspidata (Spruce ex Engl.) Nooteboom =Simaba 

guianensis 

Quassia guianensis (Aubl.) Dietr. =Simaba guianensis 

Quassia monophylla (Oliv.) Nooteboom =Simaba monophylla 

Quassia multiflora (Adr. Juss.) Nooteboom =Simaba 

multiflora 

Quassia simarouba L.f. =Simarouba amara 

Simaba alata Maguire =Simaba multiflora 

Simaba cuspidata Spruce ex Engl. =Simaba guianensis 

Simaba guianensis Aubl. subsp. polyphylla Cavalcante 

=Simaba polyphylla 

Simaba trichilioides A. St. Hil. [not in Guianas] 

Tariri guianensis Aubl. =Picramnia guianensis


SMILACACEAE 


REFERENCES: 

Mitchell, J.D. 1994. Smilax saülensis, a new 

species from French Guiana. Brittonia 46(3): 

225-227. 

Sipman, H. 1979. Liliaceae. In A.L. Stoffers & 

J.C. Lindeman, Flora of Suriname 5(1): 442- 

456. 

Smilax cinnamommea Desf. ex DC. [FG] 

Smilax cordato-ovata L.C. Rich. [FG] 

Smilax cumanensis Willd. [GU SU FG] 

Smilax cuspidata Poir. [FG] 

Smilax immersa A.C. Smith [GU] 

Smilax krukovii A.C. Smith [FG?] 

Smilax kunthii Killip & C. Morton [GU SU FG] 

Smilax lasseriana Steyerm. [GU SU FG] [=Smilax kunthii?] 

Smilax latipes Gleason [GU SU FG] 

Smilax longifolia L.C. Rich. [SU? FG] 

Smilax megalophylla Duhamel [GU SU FG?] 

Smilax pseudosyphilitica Kunth [FG] 

Smilax quinquenervia Vell. [FG] 

Smilax riedeliana A. DC. [FG] 

Smilax santaremensis DC. [GU SU FG] 

Smilax saülensis J.D. Mitchell [FG] 

Smilax schomburgkiana Kunth [GU SU FG] 

Smilax syphilitica Willd. [GU SU FG] 

Smilax talbotiana A. DC. [GU] 

SMILACACEAE - SYNONYMS AND 


Smilax flavescens Hamilton =Smilax longifolia 

Smilax floribunda Hamilton =Smilax cuspidata? 

Smilax globifera G. Mey. =Smilax cumanensis 

Smilax hostmanniana Kunth =Smilax cumanensis 

Smilax papyracea Duhamel =Smilax longifolia 

Smilax pirarensis Kunth & Rich. Schomb. =Smilax 

cumanensis 

Smilax procera Griseb. =Smilax quinquenervia 

Smilax surinamensis Miq. =Smilax cumanensis 

SOLANACEAE 

reviewed by M. Nee (1992); reviewed by M. Nee 

(1996) 

REFERENCES: 

Bernardello, L.M. & A.T. Hunziker. 1987. A 

synoptical revision of Solandra (Solanaceae). 

Nord. J. Bot. 7: 639-652. 

Bitter, G. 1919. Die Gattung Lycianthes. Ahb. 

Naturwiss. Vereine Bremen 24(2): 292-520. 

Bohs, L. 1994. Cyphomandra (Solanaceae). 

Flora Neotropica Monograph 63: 1-175. 

SPERMATOPHYTA: Smilacaceae - Solanaceae 

197 

Carvalho, L. d'A Freire de. 1978. O gênero 

Schwenckia D. van Rooyen ex Linnaeus no 

Brasil - Solanaceae. Rodriguésia 44: 307- 

524. 

Cuatrecasas, J. 1958. Notes on American 

Solanaceae. Feddes Repert. 61(1): 74-86. 

D'Arcy, W.G. 1973. Solanaceae. In R.E. 



60: 573-780. 

Edmonds, J.M. 1972. A synopsis of the 

taxonomy of Solanum sect. Solanum 

(Maurella) in South America. Kew Bull. 27: 

95-114. 

Francey, P. 1935-1936. Monographie du genre 

Cestrum L. Candollea 6: 46-398, 7: 1-132. 

Hawkes, J.G., R.N. Lester, M. Nee, & N. Estrada- 

R. 1991. Solanaceae III. Taxonomy, 

Chemistry, Evolution. Royal Botanic 

Gardens, Kew. 

Plowman, T. 1979. The genus Brunfelsia: a 

conspectus of the taxonomy and 

biogeography. Pp. 475-491 in J.G. Hawkes, 

R.N. Lester, & A.D. Skelding, The Biology 

and Taxonomy of the Solanaceae. Linnaean 

Society of London, Academic Press. 

Whalen, M.D. 1984. Conspectus of species 

groups in Solanum subgenus Leptostemonum. 

Gentes Herb. 12(4): 179-282. 

Whalen, M.D., D.E. Costich, & C.B. Heiser. 

1981. Taxonomy of Solanum section 

Lasiocarpa. Gentes Herb. 12(2): 41-129. 

Brunfelsia guianensis Benth. [GU SU FG] 

Brunfelsia martiana Plowman [GU FG] 

Brunfelsia uniflora D. Don [GU] 

Capsicum annuum L. var. annuum [GU FG] [cultivated, 


Capsicum annuum L. var. glabriusculum (Dunal) Heiser & 

Pick. [GU SU] 

Capsicum frutescens L. [GU SU FG] 

Cestrum acuminatissimum Dunal [FG] [=Cestrum 

megalophyllum?] 

Cestrum latifolium Lam. [GU SU FG] 

Cestrum megalophyllum Dunal [GU] 

Cestrum multiflorum Roem. & Schult. [FG] 

Cestrum racemosum Ruiz & Pav. [FG?] 

Cyphomandra endopogon Bitter subsp. guianensis Bohs [FG] 

Cyphomandra hartwegii (Miers) Walp. subsp. hartwegii [GU 

SU FG] 

Cyphomandra oblongifolia Bohs [SU] 

Cyphomandra tegore (Aubl.) Walp. [SU FG] 

Cyphomandra tobagensis Sandw. [GU] 

Datura metel L. [SU FG] [cultivated or escaped from 

cultivation] 

Datura stramonium L. [FG?] [naturalized?] 

Lycianthes geminata Bitter [FG] [to be expected in GU, SU] 

Lycianthes pauciflora (Vahl) Bitter [GU SU FG] 

Markea camponoti Ducke [GU FG] 

Markea coccinea L.C. Rich. [GU SU FG] 

Markea formicarum Dammer [SU FG] [to be expected in GU]

198 

SPERMATOPHYTA: Solanaceae 


Markea longiflora Miers [GU FG] [doubtful species, of 

uncertain status] 

Markea sessiliflora Ducke [GU SU FG] 

Nicotiana tabacum L. [GU SU FG] [cultivated, perhaps 

escaping] 

Physalis angulata L. [GU SU FG] 

Physalis pubescens L. [GU SU FG] 

Schwenckia americana L. var. americana [GU SU FG] 

Schwenckia americana L. var. hirta (Klotzsch) Carvalho [GU 

SU FG] 

Schwenckia elegans Carvalho [GU] 

Schwenckia grandiflora Benth. [GU SU FG?] 

Schwenckia guianensis Benth. [SU? FG] ["Guianas"] 

Schwenckia mollissima Nees & Mart. [SU] 

Solandra longiflora Tussac [SU FG?] [to be expected in GU] 

Solandra paraensis Ducke [FG] 

Solanum accrescens Standl. & C. Morton [FG?] 

Solanum adhaerens Roem. & Schult. [GU FG] 

Solanum agrarium Sendtn. [GU] [to be expected in SU, FG] 

Solanum americanum Mill. [GU SU FG] 

Solanum anceps Ruiz & Pav. [GU SU FG] 

Solanum asperum L.C. Rich. [GU SU FG] 

Solanum aturense Dunal [FG?] 

Solanum bicolor Roem. & Schult. [GU] 

Solanum campaniforme Roem. & Schult. [GU FG] 

Solanum capsicoides Allioni [GU SU] [introduced] 

Solanum cordovense Sessé & Moc. [GU] 

Solanum coriaceum Dunal [GU SU FG] 

Solanum costatum M. Nee, sp. nov. ined. [GU FG] 

Solanum crinitum Lam. [GU SU FG] 

Solanum fulvidum Bitter [GU FG] 

Solanum jamaicense Mill. [GU SU FG] 

Solanum lepidotum Dunal [GU FG?] 

Solanum leucocarpon Dunal [GU SU FG] 

Solanum mammosum L. [GU SU FG] 

Solanum maroniense Poit. [FG] 

Solanum monachophyllum Dunal [GU FG] 

Solanum morii S. Knapp [GU FG] 

Solanum oocarpum Sendtn. [GU SU?] 

Solanum oppositifolium Ruiz & Pav. [GU FG] 

Solanum paludosum Moricand [GU SU FG] 

Solanum pensile Sendtn. [GU SU FG] 

Solanum rubiginosum Vahl [GU SU FG] 

Solanum rugosum Dunal [GU SU FG] 

Solanum schlechtendalianum M. Walpers [GU SU FG] 

Solanum schomburgkii Sendtn. [GU SU FG] 

Solanum semotum M. Nee [GU FG] 

Solanum sendtnerianum Heurck & Muell. Arg. [FG] 

Solanum sessiliflorum Dunal [SU FG] [probably cultivated or 


Solanum stramoniifolium Jacq. [GU SU FG] 

Solanum subinerme Jacq. [GU SU FG] 

Solanum torvum Swartz [GU FG] 

Solanum tricuspidatum L.C. Rich. ex Dunal [FG] 

Solanum vanheurckii Muell. Arg. [FG] 

Solanum velutinum Dunal [GU SU FG] 

Witheringia solanacea L'Hér. [FG?] 

SOLANACEAE - SYNONYMS AND 


Bassovia sylvatica Aubl. =Solanum anceps 

Brunfelsia maritima Benth. =Brunfelsia latifolia (Pohl) 

Benth.[not in Guianas] 

Brunfelsia spruceana Benth. =Brunfelsia martiana 

Capsicum conoide Mill. =Capsicum annuum? 

Capsicum curvipes Dun. =Capsicum annuum? 

Cestrum baenitzii Lingelsh. =Cestrum megalophyllum 

Cestrum oliganthum Dunal =Cestrum latifolium 

Cestrum prieurei Dunal =Cestrum latifolium 

Cestrum silvaticum Francey =Cestrum microcalyx Francey 


Cyphomandra bolivarensis Steyerm. =Cyphomandra 

tobagensis 

Datura fastuosa L. =Datura metel 

Lycianthes guianensis (Dunal) Bitter =Lycianthes pauciflora 

Markea porphyrobaphes Sandw. =Markea sessiliflora 

Physalis capsicifolia Dunal =Physalis sp. [name of uncertain 

application] 

Physalis hirsuta Dunal =Physalis sp. [name of uncertain 

application] 

Physalis surinamensis Miq. =Physalis pubescens 

Schwenckia hirta Klotzsch =Schwenckia americana var. hirta 

Solanum americanum Mill. var. nodiflorum (Jacq.) Edmonds 

=Solanum americanum 

Solanum antillarum O.E. Schulz =Solanum nudum 

Solanum arboreum Dunal [not in Guianas] 

Solanum aubletii Pulle =Solanum anceps 

Solanum boissieri Dunal =Solanum schomburghii 

Solanum brachybotryon Dunal =Solanum velutinum 

Solanum cuneifolium Dunal =Solanum jamaicense 

Solanum demerarense Dunal =Solanum stramoniifolium 

Solanum diphyllum L. [not in Guianas] 

Solanum enoplocalyx Dunal =Solanum adhaerens 

Solanum foetidum Ruiz & Pav. [not in Guianas] 

Solanum guianense Dunal =Lycianthes guianensis 

Solanum guianense Klotzsch [nomen nudum] 

Solanum hostmannii Dunal =Solanum schomburghii 

Solanum intermedium Sendtner [not in Guianas] 

Solanum japurense Dunal =Lycianthes sp. 

Solanum lanceifolium Jacq. [records based on 


Solanum longiflorum Vahl =Solanum subinerme 

Solanum macranthum Dunal =Solanum crinitum 

Solanum manaense Sagot ex Lemée [nomen nudum] 

Solanum miquelii C. Morton =Solanum pensile 

Solanum nigrum L. [not in Guianas, misident. for Solanum 

americanum] 

Solanum nodiflorum Jacq. =Solanum americanum 

Solanum nudum Dunal [not in Guianas] 

Solanum ochraceo-ferrugineum (Dunal) Fernald [not in 


Solanum oleraceum Dunal =Solanum americanum 

Solanum richardii (Dunal) Lemée =Solanum anceps? 

Solanum sacupanense Rusby =Solanum monachophyllum 

Solanum salviifolium Lam. =Solanum asperum 

Solanum salzmannii Dunal =Solanum paludosum 

Solanum scandens L. =Solanum pensile 

Solanum sempervirens Dunal =Solanum pensile 

Solanum siparunoides Ewan =Solanum aturense 

Solanum surinamense Steud. =Solanum leucocarpon 

Solanum sylvaticum (Aubl.) Bitter =Solanum anceps 

Solanum tegore Aubl. =Cyphomandra tegore 

Solanum topiro Dunal =Solanum sessiliflorum 

Solanum toxicarium L. Rich. =Solanum stramoniifolium 

Solanum trichocarpum Miq. =Solanum stramoniifolium 

Solanum verbascifolium L. [not in Guianas]

Strelitziaceae 


SPHENOCLEACEAE 

Sphenoclea zeylanica Gaertn. [GU SU FG] [naturalized?] 

STERCULIACEAE 

contributed by L.J. Dorr (1992) 

REFERENCES: 

Barnett, L.C. & L.J. Dorr. 1990. A new, 

arborescent species of Byttneria from French 

Guiana. Brittonia 42(4): 271-275. 

Cristóbal, C.L. 1976. Estudio taxonómico del 

género Byttneria Loefl. Bonplandia 4: 1-428. 

Cuatrecasas, J. 1964. Cacao and its allies. A 

taxonomic revision of the genus Theobroma. 

Contr. U.S. Nat. Herb. 35: 379-614. 

Goldberg, A. 1967. The genus Melochia L. 

Contrib. United States National Herbarium 

34(5): 191-363. 

Uittien, H. 1932. Sterculiaceae. In A. Pulle, 


Uittien, H. 1941. Sterculiaceae (additions and 


3(1): 437. 

Ayenia praeclara Sandw. [GU SU] 

Ayenia tomentosa L. [GU SU] 

Byttneria aurantiaca Mildbr. [FG] 

Byttneria cordifolia Sagot [SU FG] 

Byttneria divaricata Benth. var. divaricata [GU SU] 

Byttneria genistella Triana & Planch. [GU] 

Byttneria morii L. Barnett & Dorr [FG] 

Byttneria scabra L. [GU SU FG] 

Byttneria uaupensis Spruce ex K. Schum. [GU] 

Guazuma ulmifolia Lam. [GU SU FG] 

Helicteres baruensis Jacq. [GU SU] 

Helicteres guazumaefolia Kunth [GU] 

Helicteres pentandra L. [GU? SU FG] 

Herrania kanukuensis R.E. Schultes [GU SU FG] 

Herrania lemniscata (Rich. Schomb.) R.E. Schultes [GU] 

Melochia arenosa Benth. [GU SU] 

Melochia graminifolia St. Hil. [GU] 

Melochia lanceolata Benth. [GU SU FG?] 

Melochia melissifolia Benth. [GU SU FG] 

Melochia parvifolia Kunth var. fasciculata (Benth.) Hassler 

[GU SU?] 

Melochia parvifolia Kunth var. parvifolia [GU] 

Melochia pyramidata L. var. pyramidata [FG?] 

Melochia simplex St. Hil. [GU SU] 

Melochia spicata (L.) Fryxell [GU SU FG] 

Melochia tomentosa L. [SU FG?] 

Melochia ulmifolia Benth. [GU SU FG] 

Sterculia frondosa A. Rich. [FG] 

Sterculia guianensis Sandw. [GU] 

Sterculia lisae E. Taylor, sp. nov. ined. [FG] 

Sterculia multiovula E. Taylor, sp. nov. ined. [GU FG] 

Sterculia parviflora (Ducke) E. Taylor, comb. nov. ined. [SU 

FG] 

SPERMATOPHYTA: Sphenocleaceae - 

199 

Sterculia pruriens (Aubl.) K. Schum. var. glabrescens E. 

Taylor, var. nov. ined. [GU] 

Sterculia pruriens (Aubl.) K. Schum. var. pruriens [GU SU 

FG] 

Sterculia rugosa R. Brown [GU] 

Sterculia speciosa K. Schum. [GU FG] 

Sterculia villifera Steud. [GU? SU FG] 

Theobroma cacao L. subsp. spherocarpum (Chev.) Cuatrec. 

[GU SU FG] 

Theobroma grandiflorum (Willd. ex Spreng.) K. Schum. [GU] 

Theobroma obovatum Klotzsch ex Bernoulli [GU] 

Theobroma subincanum Mart. [GU SU FG] 

Theobroma velutinum Benoist [SU FG] 

Waltheria indica L. [GU SU FG] 

Waltheria involucrata Benth. [GU] 

Waltheria viscosissima St. Hil. [GU] 

STERCULIACEAE - SYNONYMS AND 


Buttneria scabra Aubl. =Byttneria scabra 

Byttneria sparganiocarpa Uittien =Byttneria divaricata 

Cacao guianensis Aubl. =Theobroma cacao 

Cacao sylvestris Aubl. =Theobroma subincanum 

Herrania mariae (Mart.) Decne. ex Goudot [not in Guianas] 

Ivira pruriens Aubl. =Sterculia pruriens 

Melochia cinerea St. Hil. & Naud. =Melochia arenosa 

Melochia fasciculata Benth. =Melochia parvifolia var. 

fasciculata 

Melochia hirsuta Cav. =Melochia spicata 

Melochia ramuliflora Fries =Melochia ulmifolia 

Melochia villosa (Mill.) Fawc. & Rendle =Melochia spicata 

Sterculia excelsa Mart. [not in Guianas] 

Theobroma alba Bernoulli =Licania alba (Chrysobalanaceae) 

Theobroma guianense (Aubl.) Gmelin =Theobroma cacao 

Theobroma mariae Schum. =Herrania mariae (Mart.) Decne. 

ex Goud.[not in Guianas] 

Theobroma speciosum Mart. [not in Guianas] 

Theobroma sylvestre Mart., non Cacao sylvestris Aubl. [not in 


Waltheria americana L. =Waltheria indica 

Xylosterculia rugosa (R. Brown) Kosterm. =Sterculia rugosa 

STRELITZIACEAE 

reviewed by P.J.M. Maas (1992) 

REFERENCES: 

Maas, P.J.M. 1985. Musaceae. In A.R.A. Görtsvan 


28. 

Maas, P.J.M. & M.J.M. de Rooij. 1979. 

Musaceae. In A.L. Stoffers & J.C. Lindeman, 


Phenakospermum guyannense (L.C. Rich.) Endl. ex Miq. [GU 

SU FG]

200 


STRELITZIACEAE - SYNONYMS AND 


Ravenala guyannensis (L.C. Rich.) O.G. Peters. 

=Phenakospermum guyannense 

Urania guyannensis L.C. Rich. =Phenakospermum 

guyannense 

STYRACACEAE 


REFERENCES: 

Maguire, B. & Yung-Chau Huang. 1978. 

Styracaceae. In B. Maguire & collaborators, 



Perkins, J. 1907. Styracaceae. In A. Engler, Das 

Pflanzenreich IV(241): 1-111. 

Styrax fanshawei Sandw. [GU] 

Styrax glaber Swartz [GU] 

Styrax glabratus Schott ex Spreng. [SU FG] 

Styrax guianensis A. DC. [GU SU FG] 

Styrax hypochryseus Perkins [GU] 

Styrax longifolius Standl. [GU FG] 

Styrax oblongus (Ruiz & Pav.) A. DC. [GU] 

Styrax pallidus A. DC. [FG] 

Styrax roraimae Perkins [GU?] [Mt. Roraima] 

Styrax tafelbergensis Maguire [GU? SU] 

Styrax tessmannii Perkins [GU] 

REFERENCES: 

SURIANACEAE 

Howard, R.A. 1988. Surianaceae. In R.A. 

Howard, Flora of the Lesser Antilles 4(1): 

574-575. 

Suriana maritima L. [FG?] 

SYMPLOCACEAE 


REFERENCES: 

Brand, A. 1901. Symplocaceae. In A. Engler, 

Das Pflanzenreich IV(242): 1-100. 

Maguire, B. & Yung-Chau Huang. 1978. 

Symplocaceae. In B. Maguire & 



223-230. 

SPERMATOPHYTA: Strelitziaceae - Theaceae 

Symplocos guianensis (Aubl.) Guerke [GU SU FG] 

Symplocos martinicensis Jacq. [GU FG] 

Symplocos pilosiuscula Brand [GU FG] 

Symplocos pycnophylla Sleumer [Mt. Roraima] 

Symplocos schomburgkii (Klotzsch ex Schomb.) Brand [GU?] 

[Mt. Roraima] 

Symplocos ulei Brand [GU] 

SYMPLOCACEAE - SYNONYMS AND 


Ciponima guianensis Aubl. =Symplocos guianensis 

Symplocos roraimensis Steyerm. =Symplocos pycnophylla 

REFERENCES: 

TACCACEAE 

Drenth, E. 1972. A revision of the family 

Taccaceae. Blumea 20: 367-406. 

Tacca parkeri Seem. [GU SU?] 

THEACEAE 

reviewed by A. Weitzman (1992) 

REFERENCES: 

Jansen-Jacobs, M.J. & J.C. Lindeman. 1979. 

Theaceae. In A.L. Stoffers & J.C. Lindeman, 


Kobuski, C.E. 1942. Studies on the Theaceae, 

XII. Notes on the South American species of 

Ternstroemia. J. Arn. Arb. 23: 298-343. 

Freziera carinata A. Weitzman [to be expected in GU] 

Freziera roraimensis Tul. [GU?] [Mt. Roraima] 

Gordonia fruticosa (Schrad.) H. Keng [GU SU FG] 

Ternstroemia browniana Kobuski [GU SU FG?] 

Ternstroemia candolleana Wawra [SU] 

Ternstroemia crassifolia Benth. [GU] 

Ternstroemia dehiscens Huber [FG?] 

Ternstroemia delicatula Choisy [GU SU FG] 

Ternstroemia dentata (Aubl.) Swartz [GU SU FG] 

Ternstroemia gleasoniana Kobuski [GU] 

Ternstroemia grandiosa Kobuski [GU] 

Ternstroemia laevigata Wawra [GU] 

Ternstroemia longipes Klotzsch ex Wawra [GU] 

Ternstroemia punctata (Aubl.) Swartz [GU SU FG] 

Ternstroemia schomburgkiana Benth. [GU SU] 

Ternstroemia verticillata Klotzsch ex Wawra [GU SU]

THEACEAE - SYNONYMS AND 



Laplacea fruticosa (Schrad.) Kobuski =Gordonia fruticosa 

Laplacea semiserrata (Nees) Camb. =Gordonia fruticosa 

Taonabo dentata Aubl. =Ternstroemia dentata 

Taonabo punctata Aubl. =Ternstroemia punctata 

Ternstroemia cayennensis Planch. [nomen nudum] 

Ternstroemia circumscissilis Kobuski [not in Guianas] 

Ternstroemia corymbosa Smith [name of uncertain 

application, not Ternstroemia] 

THEOPHRASTACEAE 

reviewed by J.C. Lindeman (1992) 

REFERENCES: 

Lindeman, J.C. 1979. Theophrastaceae. In A.L. 


5(1): 367-369. 

Ståhl, B. 1991. A revision of Clavija. Opera 

Bot. 107: 1-77. 

Clavija imatacae Ståhl [GU] 

Clavija lancifolia Desf. subsp. chermontiana (Standl.) Ståhl 

[GU SU FG] 

Clavija lancifolia Desf. subsp. lancifolia [SU FG] 

Clavija macrophylla (Link ex Roem. & Schult.) Radlk. [GU] 

THURNIACEAE 

reviewed by M.T. Strong (1992) 

REFERENCES: 

Schnee, L. 1943. El genero Thurnia (con 

descripción de una nueva especie). Bol. Soc. 

Venez. Cienc. Nat. 8(55): 241-243. 

Uittien, H. 1938. Thurniaceae. In A. Pulle, 


Thurnia jenmanii J.D. Hook. [GU] 

Thurnia sphaerocephala (Rudge) J.D. Hook. [GU SU FG] 

REFERENCES: 

THYMELAEACEAE 

Domke, W. 1934. Untersuchungen über die 

systematische und geographische Gliederung 

der Thymelaeaceen. Bibliotheca Botanica 

111: 1-151. 

Nevling, L 1959. A revision of the genus 

Daphnopsis. Ann. Missouri Bot. Gard. 46: 

257-358. 

Daphnopsis longipedunculata Gilg ex Domke [Mt. Roraima] 

[=Funifera sp.?] 

Daphnopsis steyermarkii Nevling [GU] 

Goodallia guianensis Benth. [GU] 

Lasiadenia rupestris Benth. [GU] 

Schoenobiblus daphnoides Mart. [FG?] 

TILIACEAE 

contributed by M.J. Jansen-Jacobs & W. Meijer 

(1992); additions and corrections by M.J. 

Jansen-Jacobs (1996) 

REFERENCES: 

SPERMATOPHYTA: Theaceae - Tiliaceae 

201 

Jansen-Jacobs, M.J. 1986. Tiliaceae (additions 

and corrections). In A.L. Stoffers & J.C. 

Lindeman, Flora of Suriname 3(1-2): 293- 

300. 

Jansen-Jacobs, M.J. & W. Meijer. 1995. 

Tiliaceae. In A.R.A. Görts-van Rijn, Flora of 

the Guianas (series A) 17: 5-49. 

Jansen-Jacobs, M.J. & L.Y.Th. Westra. 1995. A 

new species of Apeiba (Tiliaceae) from the 

Venezuelan-Brazilian border. Brittonia 47(3): 

335-339. 

Ko Ko Lay 1950. The American species of 

Triumfetta L. Ann. Missouri Bot. Gard. 

37(3): 315-395. 

Kubitzki, K. 1995. Asterophorum and Tahitia 

congeneric with Christiana (Tiliaceae). Bot. 

Jahrb. Syst. 116: 537-542. 

Meijer, W. 1978. Mollia (Tiliaceae). A revision 

of the South American species. In B. Maguire 


Highland - Part X. Mem. New York Bot. 

Gard. 29: 68-82. 

Uittien, H. 1932. Tiliaceae. In A. Pulle, Flora of 

Suriname 3(1): 49-57. 

Uittien, H. 1941. Tiliaceae (additions and 


3(1): 438-441. 

Apeiba glabra Aubl. [SU FG] 

Apeiba intermedia Uittien [SU] 

Apeiba macropetala Ducke [GU] 

Apeiba membranacea Spruce ex Benth. [GU] 

Apeiba petoumo Aubl. [GU SU FG] 

Apeiba schomburgkii Szyszyl. [GU SU] 

Apeiba tibourbou Aubl. [GU SU FG] 

Christiana africana DC. [GU] 

Christiana mennegae (Jansen-Jacobs & Westra) Kubitzki [SU] 

Corchorus aestuans L. [GU SU FG] 

Corchorus hirtus L. [GU SU] 

Corchorus orinocensis Kunth [GU] 

Corchorus siliquosus L. [GU] 

Luehea paniculata Mart. & Zucc. [SU] 

Luehea speciosa Willd. [GU SU FG]

202 


Lueheopsis rosea (Ducke) Burret [GU SU FG] 

Lueheopsis rugosa (Pulle) Burret [GU SU FG] 

Mollia glabrescens Benth. [GU] 

Mollia globularis Meijer [GU] 

Mollia lepidota Spruce ex Benth. subsp. sphaerocarpa 

(Gleason ex Record) Meijer [GU] 

Mollia ulei Burret [GU] 

Triumfetta althaeoides Lam. [GU SU FG] 

Triumfetta lappula L. [SU FG] 

Triumfetta rhomboidea Jacq. [GU FG] 

Triumfetta semitriloba Jacq. [GU SU?] 

Vasivaea alchorneoides Baill. [GU] 

TILIACEAE - SYNONYMS AND 


Apeiba albiflora Ducke =Apeiba tibourbou 

Apeiba aspera Aubl. =Apeiba petoumo 

Apeiba burchellii Sprague =Apeiba glabra 

Apeiba echinata Gaertn. =Apeiba petoumo 

Apeiba hypoleuca Steud. =Apeiba petoumo 

Apeiba surinamensis Uittien =Apeiba schomburgkii 

Asterophorum mennegae Jansen-Jacobs & Westra =Christiana 

mennegae 

Corchorus acutangulus Lam. =Corchorus aestuans 

Corchorus argutus Kunth =Corchorus orinocensis 

Corchorus argutus Kunth var. benthamii K. Schum. 

=Corchorus orinocensis 

Guenetia macrosperma Sagot =Catostemma fragrans 

(Bombacaceae) 

Luehea maroniensis Benoist =Lueheopsis rugosa 

Lueheopsis flavescens (Uittien) Burret =Lueheopsis rosea 

Mollia scabra Meijer =Mollia ulei 

Mollia speciosa Mart. & Zucc. [not in Guianas] 

Mollia sphaerocarpa Gleason ex Record =Mollia lepidota 

subsp. sphaerocarpa 

Mollia tomentosa Spruce ex Benth. [not in Guianas] 

Triumfetta bartramia sensu Lay, non L. =Triumfetta 

rhomboidea 

Triumfetta hostmannii Miq. =Triumfetta lappula 

Triumfetta macrophylla Vahl =Triumfetta althaeoides 

Triumfetta sinuosa Miq. =Triumfetta lappula 

Triumfetta surinamensis Steud. =Triumfetta althaeoides 

TRIGONIACEAE 

reviewed by E. Lleras-Pérez (1992) 

REFERENCES: 

Lleras, E. 1978. Trigoniaceae. Flora Neotropica 

Monogr. 19: 1-73. 

Stafleu, F.A. 1951. Trigoniaceae. In A. Pulle, 


Trigonia candelabra Lleras [SU] 

Trigonia coppenamensis Stafl. [SU] 

Trigonia hypoleuca Griseb. [GU SU FG] 

Trigonia laevis Aubl. var. laevis [GU FG] 

Trigonia laevis Aubl. var. microcarpa (Sagot ex Warm.) Sagot 

[GU SU FG] 

SPERMATOPHYTA: Tiliaceae - Triuridaceae 

Trigonia nivea Cambess. var. nivea [GU] 

Trigonia reticulata Lleras [GU] 

Trigonia subcymosa Benth. [GU] 

Trigonia villosa Aubl. var. macrocarpa (Benth.) Lleras [GU 

FG?] 

Trigonia villosa Aubl. var. villosa [GU SU? FG] 

TRIGONIACEAE - SYNONYMS AND 


Trigonia hypoleuca Griseb. var. pubescens Warm. =Trigonia 

hypoleuca 

Trigonia kaieteurensis Maguire =Trigonia laevis var. laevis 

Trigonia macrocarpa Benth. =Trigonia villosa var. macrocarpa 

Trigonia microcarpa Sagot ex Warm. =Trigonia laevis var. 

microcarpa 

Trigonia villosa Aubl. var. cuneata DC. =Trigonia villosa var. 

villosa 

Trigonia villosa Aubl. var. oblonga DC. =Trigonia villosa var. 

villosa 

Trigonia villosa Aubl. var. obtusata DC. =Trigonia villosa 

var. villosa 

Trigonia xanthopila Garke =Trigonia hypoleuca 

TRIURIDACEAE 

reviewed by P.J.M. Maas (1992); additions and 

corrections by P.J.M. Maas (1996) 

REFERENCES: 

Jonker, F.P. 1943. Triuridaceae. In A. Pulle, 


Maas, P.J.M. & T. Rübsamen. 1986. 

Triuridaceae. Flora Neotropica Monogr. 40: 

1-55. 


Triuridaceae. In A.R.A. Görts-van Rijn, Flora 

of the Guianas (series A) 5: 1-16. 

Peltophyllum luteum Gardner [GU] 

Sciaphila albescens Benth. [GU SU FG] 

Sciaphila picta Miers [GU?] 

Sciaphila polygyna Maas [FG] 

Sciaphila purpurea Benth. [GU] 

Soridium spruceanum Miers [SU FG] 

Triuris hexophthalma Maas [GU] 

Triuris hyalina Miers [GU SU] 

TRIURIDACEAE - SYNONYMS AND 


Sciaphila guianensis Sandw. =Sciaphila albescens 

Sciaphila spruceana (Miers) Benth. & J.D. Hook. =Soridium 

spruceanum 

Soridium guianense (Sandw.) Giesen =Sciaphila albescens


TURNERACEAE 

contributed by M. Hoff (1992); revised by M. 

Hoff & M.-M. Arbo (1996) 

REFERENCES: 

Arbo, M.M. 1990. Turneraceae: novedades para 

la Guayana Venezolana. Ann. Missouri Bot. 

Gard. 77: 340-352. 

Arbo, M.M. 1995. Tuneraceae - Parte I. 

Piriqueta. Flora Neotropica Monogr. 67: 1- 

156. 

Bremekamp, E. 1941. Turneraceae. In A. Pulle, 


Lemée, A. 1953. Flore de la Guyane française 3: 

82. Paul Lechevalier, Paris. 

Pulle, A. 1906. An enumeration of the Vascular 

Plants known from Surinam, together with 

their distribution and Synonymy. Leiden. 

Urban, I. 1883a. Monographie des Familie der 

Turneraceen. Jahrbuch des Königlichen 

Botanischen Gartens und des botanischen 

Museums zu Berlin, 2: 1-152. 

Urban, I. 1883b. Turneraceae. In C.F.P. 

Martius, Flora Brasiliensis 13(3): 88-170, tab. 

31-48. 

Piriqueta cistoides (L.) Griseb. subsp. cistoides [GU SU FG] 

Piriqueta guianensis N.E. Brown subsp. guianensis [GU] 

Piriqueta viscosa Griseb. subsp. viscosa [GU FG] 

Turnera aurantiaca Benth. [GU SU FG] 

Turnera brasiliensis Willd. ex Roem. & Schultes [GU SU FG] 

Turnera caerulea Moçino & Sessé ex DC. var. surinamensis 

(Urban) Arbo, comb. nov. ined. [GU SU] 

Turnera cicatricosa Arbo [GU] 

Turnera glaziovii Urb. [GU? SU FG] 

Turnera guianensis Aubl. [GU SU FG] 

Turnera lineata Urb. [GU] 

Turnera odorata L.C. Rich. [GU SU FG] 

Turnera pumilea L. [FG] 

Turnera rupestris Aubl. var. frutescens (Aubl.) Urb. [GU SU 

FG] 

Turnera rupestris Aubl. var. rupestris [GU SU FG] 

Turnera scabra Millsp. [GU FG?] 

Turnera schomburgkiana Urb. [GU] 

Turnera subulata J.E. Smith [GU SU FG] 

Turnera velutina Benth. [GU] 

Turnera waltherioides Urb. [GU] 

TURNERACEAE - SYNONYMS AND 


Piriqueta cistoides (L.) Griseb. var. bracteolata Urb. 

=Piriqueta cistoides subsp. cistoides 

Piriqueta cistoides (L.) Griseb. var. foliosa (Garcke) Urb. 

=Piriqueta cistoides subsp. cistoides 

Piriqueta cistoides (L.) Griseb. var. genuina Urban =Piriqueta 

cistoides subsp. cistoides 

Piriqueta cistoides (L.) Griseb. var. latifolia Urb. =Piriqueta 

cistoides subsp. cistoides 

Piriqueta foliosa Garcke =Piriqueta cistoides subsp. cistoides 

SPERMATOPHYTA: Turneraceae - Ulmaceae 

203 

Piriqueta lanceolata Benth. =Piriqueta cistoides subsp. 

cistoides 

Piriqueta stenophylla Klotzsch =Piriqueta cistoides subsp. 

cistoides 

Piriqueta villosa Aubl. =Piriqueta cistoides subsp. cistoides 

Turnera aspera Poir. =Piriqueta cistoides subsp. cistoides 

Turnera benthamiana Schomb. =Turnera velutina 

Turnera chamaedrys Klotzsch [excluded taxon] 

Turnera cistoides L. =Piriqueta cistoides 

Turnera corchorifolia Willd. =Turnera odorata 

Turnera corchoroides Klotzsch [excluded taxon] 

Turnera desvauxii DC. =Piriqueta cistoides subsp. cistoides 

Turnera frutescens Aubl. =Turnera rupestris var. frutescens 

Turnera grandifolia Bremek. =Turnera glaziovii 

Turnera hirta Desv., non Willd. =Piriqueta cistoides subsp. 

cistoides 

Turnera intermedia (Urb.) Auct. =Turnera scabra 

Turnera opifera Mart. =Turnera caerulea var. surinamensis 

Turnera parviflora Benth. =Turnera odorata 

Turnera refracta Klotzsch =Turnera caerulea var. surinamensis 

Turnera subglabra Klotzsch =Turnera aurantiaca 

Turnera ulmifolia L. var. elegans (Otto) Urb. =Turnera 

subulata 

Turnera ulmifolia L. var. intermedia Urb. =Turnera scabra 

Turnera ulmifolia L. var. surinamensis Urb. =Turnera 

caerulea var. surinamensis 

TYPHACEAE 

reviewed by M.T. Strong (1992) 

REFERENCES: 

Lanjouw, J. 1938. Typhaceae. In A. Pulle, Flora 


Typha domingensis Pers. [GU SU FG] 

TYPHACEAE - SYNONYMS AND 


Typha angustifolia L. [not in Guianas] 

ULMACEAE 



REFERENCES: 

Berg, C.C. 1992. Ulmaceae. In A.R.A. Görtsvan 


3-9. 

Ooststroom, S.J. van. 1932. Ulmaceae. In A. 

Pulle, Flora of Suriname 1(1): 47-48.

204 

Ampelocera edentula Kuhlm. [GU SU FG] 

Celtis iguanaea (Jacq.) Sarg. [GU SU FG] 

Trema micrantha (L.) Blume [GU SU FG] 


URTICACEAE 



REFERENCES: 

Berg, C.C. 1992. Urticaceae. In A.R.A. Görtsvan 


125-139. 

Rooij, M.J.M. de. 1975. Urticaceae. In J. 

Lanjouw & A.L. Stoffers, Flora of Suriname 

5(1): 300-318. 

Boehmeria cylindrica (L.) Swartz [SU FG] 

Boehmeria ramiflora Jacq. [SU FG] 

Laportea aestuans (L.) Chew [GU SU FG] 

Phenax sonneratii (Poir.) Wedd. [FG] [naturalized?] 

Pilea imparifolia Wedd. [GU SU FG] 

Pilea microphylla (L.) Liebm. [GU SU FG] 

Pilea pubescens Liebm. [GU SU FG] 

Pilea tabularis C.C. Berg [FG] 

Urera baccifera (L.) Gaudichaud ex Wedd. [GU SU FG] 

Urera caracasana (Jacq.) Griseb. [GU SU FG] 

Urera laciniata Wedd. [SU FG] 

URTICACEAE - SYNONYMS AND 


Fleurya aestuans (L.) Gaudichaud ex Miq. =Laportea aestuans 

Pilea dendrophila Miq. [not in Guianas] 

Pilea guyanensis Wedd. =Pilea pubescens 

Pilea pubescens Liebm. var. guyanensis (Wedd.) Wedd. 

=Pilea pubescens 

Urtica aestuans L. =Laportea aestuans 

Urtica alceaefolia Poir. =Urera caracasana 

REFERENCES: 

VALERIANACEAE 

Xena de Enrech, N. 1992. Valerianaceae. In G. 

Morillo, Flora de Venezuela 5(1): 221-267. 

Valeriana scandens L. var. candolleana (Gardn.) Muell. [Mt. 

Roraima] 

REFERENCES: 

VELLOZIACEAE 

SPERMATOPHYTA: Ulmaceae - Verbenaceae 

Maguire, B. 1969. Velloziaceae. In B. Maguire 


Highland part VIII. Mem. New York Bot. 

Gard. 18(2): 1-290. 

Smith, L.B. & E.S. Ayensu. 1976. A revision of 

American Velloziaceae. Smiths. Contrib. Bot. 

30: 1-172. 

Vellozia tubiflora (A. Rich.) Kunth [GU] 

VELLOZIACEAE - SYNONYMS AND 


Barbacenia alexandrinae Schomb. ex Hook. =Vellozia 

tubiflora 

Vellozia alexandrinae (Schomb. ex Hook.) Goeth. & Henr. 

=Vellozia tubiflora 

VERBENACEAE 

contributed by M.J. Jansen-Jacobs (1992); 

additions and corrections by M.J. Jansen- 

Jacobs (1996) 

REFERENCES: 

Jansen-Jacobs, M.J. 1988. Verbenaceae. In 


(series A) 4: 1-114. 

Lopez-Palacios, S. 1977. Verbenaceae. In Flora 

de Venezuela. 

Moldenke, H.N. 1940a. Avicenniaceae. In A. 


Moldenke, H.N. 1940b. Verbenaceae. In A. 


Moldenke, H.N. 1971. A Fifth Summary of the 

Verbenaceae, Avicenniaceae, Stilbaceae, 

Dicrostylidaceae, Symphoremaceae, 

Nyctanthaceae, and Eriocaulaceae of the 

World. H.N. Moldenke, Wayne, N.J. 

Rueda, R.M. 1994. Systematics and evolution of 

the genus Petrea. Ann. Missouri Bot. Gard. 

81(4): 610-652. 

Aegiphila bracteolosa Moldenke [GU] 

Aegiphila deppeana Steud. [FG] 

Aegiphila gleasonii Moldenke [GU] [insufficiently known 

taxon] 

Aegiphila integrifolia (Jacq.) B.D. Jacks. [GU SU FG] 

Aegiphila laevis (Aubl.) J.F. Gmelin [GU SU FG] 

Aegiphila laxiflora Benth. [GU] 

Aegiphila lhotzkiana Cham. [SU FG] 

Aegiphila longifolia Turcz. [GU] 

Aegiphila macrantha Ducke [GU SU FG] 

Aegiphila membranacea Turcz. [GU SU FG] 

Aegiphila racemosa Vell. [GU SU FG] 

Aegiphila roraimensis Moldenke [GU] 

Aegiphila villosa (Aubl.) J.F. Gmelin [GU FG]


Aegiphila vitelliniflora Klotzsch ex Walp. var. egleri 

Moldenke [FG?] [insufficiently known taxon] 

Amasonia campestris (Aubl.) Moldenke [GU SU FG] 

Avicennia germinans (L.) Stearn [GU SU FG] 

Avicennia schaueriana Stapf & Leechman ex Moldenke [GU 

SU] 

Citharexylum macrophyllum Poir. [GU SU FG] 

Citharexylum spinosum L. [GU SU FG] 

Clerodendrum aculeatum (L.) Schlechtend. [GU SU FG] 

Clerodendrum fragrans (Venten.) Willd. var. pleniflorum 

Schauer [GU SU FG] [cultivated, naturalized] 

Clerodendrum indicum (L.) Kuntze [GU SU FG] [cultivated, 

naturalized] 

Clerodendrum japonicum (Thunb.) Sweet [GU SU] 


Clerodendrum thomsonae Balf. [GU SU FG] [cultivated, 

naturalized] 

Cornutia pubescens C.F. Gaertn. [FG] 

Cornutia pyramidata L. [SU] 

Lantana camara L. [GU SU FG] 

Lantana canescens Kunth [GU] 

Lantana radula Swartz [GU] 

Lantana trifolia L. [GU SU FG] 

Lippia alba (Mill.) N.E. Brown [GU SU FG] 

Lippia aristata Schauer [GU] 

Lippia betulifolia Kunth [GU SU] 

Lippia micromera Schauer [GU] 

Lippia origanoides Kunth [GU] 

Petrea bracteata Steud. [GU SU FG] 

Petrea macrostachya Benth. [GU SU FG] 

Petrea sulphurea M.J. Jansen-Jacobs [FG] 

Petrea volubilis L. [GU SU FG] 

Priva lappulacea (L.) Pers. [GU SU FG] 

Stachytarpheta angustifolia (Mill.) Vahl [GU SU FG] 

Stachytarpheta cayennensis (L.C. Rich.) Vahl [GU SU FG] 

Stachytarpheta jamaicensis (L.) Vahl [GU SU FG] 

Stachytarpheta mutabilis (Jacq.) Vahl [GU SU? FG] 

Stachytarpheta sprucei Moldenke [GU] 

Tamonea spicata Aubl. [GU SU FG] 

Vitex agnus-castus L. [GU SU FG] [cultivated, naturalized] 

Vitex capitata Vahl [GU SU] 

Vitex compressa Turcz. [GU SU FG] 

Vitex guianensis Moldenke [GU FG] 

Vitex orinocensis Kunth var. multiflora (Miq.) Huber [GU SU 

FG] 

Vitex panshiniana Moldenke [SU FG?] 

Vitex schomburgkiana Schauer [GU] 

Vitex stahelii Moldenke [GU SU FG] 

Vitex triflora Vahl [GU SU FG] 

VERBENACEAE - SYNONYMS AND 


Aegiphila cowanii Moldenke =Aegiphila integrifolia 

Aegiphila elata Swartz [not in Guianas] 

Aegiphila guianensis Moldenke =Aegiphila integrifolia 

Aegiphila integrifolia (Jacq.) Jacks. var. guianensis 

(Moldenke) López-Palacios =Aegiphila integrifolia 

Aegiphila lutea Lam. =Aegiphila laevis 

Aegiphila martinicensis Jacq. [not in Guianas] 

Amasonia arborea Kunth =Amasonia campestris 

Amasonia campestris (Aubl.) Moldenke var. surinamensis 

Moldenke =Amasonia campestris 

Amasonia erecta L.f. =Amasonia campestris 

SPERMATOPHYTA: Verbenaceae - Violaceae 

205 

Amasonia erecta L.f. var. punicea (Vahl) Pulle =Amasonia 

campestris 

Amasonia punicea Vahl =Amasonia campestris 

Avicennia marina (Forskal) Vierhapper =Avicennia germinans 

Avicennia nitida Jacq. =Avicennia germinans 

Citharexylum fruticosum L. =Citharexylum spinosum 

Clerodendrum aculeatum (L.) Schlechtend. var. guianense 

Moldenke =Clerodendrum aculeatum 

Cornutia cayennensis Schauer =Cornutia pubescens 

Cryptocalyx nepetifolia Benth. =Lippia betulifolia 

Ghinia spicata (Aubl.) Moldenke =Tamonea spicata 

Lantana armata Schauer var. guianensis Moldenke =Lantana 

camara 

Lippia schomburgkiana Schauer =Lippia origanoides 

Lippia stoechadifolia (L.) Kunth =Phyla stoechadifolia (L.) 

Small [not in Guianas] 

Manabea arborescens Aubl. =Aegiphila integrifolia 

Manabea laevis Aubl. =Aegiphila laevis 

Manabea villosa Aubl. =Aegiphila villosa 

Petrea blanchetiana Schauer [not in Guianas] 

Petrea kohautiana K. Presl =Petrea volubilis 

Petrea martiana Schauer =Petrea bracteata 

Petrea rivularis Moldenke =Petrea volubilis 

Petrea rugosa Kunth [not in Guianas] 

Petrea schomburgkiana Schauer =Petrea bracteata 

Petrea subserrata Cham. =Petrea racemosa Nees [not in 


Phyla betulaefolia (Kunth) Greene =Lippia betulifolia 

Phyla nodiflora (L.) Greene [not in Guianas] 

Stachytarpheta dichotoma (Ruiz & Pav.) Vahl =Stachytarpheta 

cayennensis 

Stachytarpheta elatior Schrad. ex Schult. & Roem. 

=Stachytarpheta angustifolia 

Stachytarpheta elatior Schrad. ex Schult. & Roem. var. 

jenmanii Moldenke =Stachytarpheta angustifolia 

Stachytarpheta orubica (L.) Vahl [not in Guianas] 

Stachytarpheta roraimensis Moldenke =Stachytarpheta sprucei 

Stachytarpheta surinamensis Miq. [nomen nudum] 

=Stachytarpheta angustifolia 

Taligalea campestris Aubl. =Amasonia campestris 

Verbena cayennensis L.C. Rich. =Stachytarpheta cayennensis 

Verbena suffruticosa Aubl. =Phyla stoechadifolia (L.) Small 


Vitex appuni Moldenke =Vitex schomburgkiana 

Vitex brittoniana Moldenke =Vitex compressa 

Vitex multiflora Miq. =Vitex orinocensis var. multiflora 

Vitex wittrockiana Moldenke =Vitex capitata 

VIOLACEAE 

reviewed by W.H.A. Hekking (1992); reviewed 

by W.H.A. Hekking (1996) 

REFERENCES: 

Hekking, W.H.A. 1988. Violaceae part I - 

Rinorea and Rinoreocarpus. Flora Neotropica 

Monogr. 46: 1-208.

206 


Amphirrhox latifolia Mart. ex Eichl. [GU SU FG] 

[=Amphirrhox longifolia s.l.] 

Amphirrhox longiflora (Oudemans) S.F. Blake [GU SU FG] 

[=Amphirrhox longifolia s.l.] 

Amphirrhox longifolia (St. Hil.) Spreng. [GU SU FG] 

Amphirrhox surinamensis Eichl. [GU SU FG] [=Amphirrhox 

longifolia s.l.] 

Corynostylis arborea (L.) S.F. Blake [GU SU FG] 

Gloeospermum sphaerocarpum Triana & Planch. [SU FG] 

Hybanthus calceolaria (L.) G.K. Schulze [GU SU FG] 

Hybanthus oppositifolius (L.) Taub. [GU SU FG] 

Leonia glycycarpa Ruiz & Pav. [GU SU FG] 

Noisettia orchidiflora (Rudge) Gingins [SU FG] 

Paypayrola confertiflora Tul. [GU SU FG] 

Paypayrola grandiflora Tul. [GU FG] 

Paypayrola guianensis Aubl. [GU SU FG] 

Paypayrola hulkiana Pulle [SU FG] 

Paypayrola longifolia Tul. [GU SU FG] 

Rinorea amapensis Hekking [SU FG] 

Rinorea bahiensis (Moric.) Kuntze [FG] 

Rinorea brevipes (Benth.) S.F. Blake [GU SU FG] 

Rinorea endotricha Sandw. [GU] 

Rinorea falcata (Mart. ex Eichl.) Kuntze [SU FG] 

Rinorea flavescens (Aubl.) Kuntze [GU SU FG] 

Rinorea guianensis Aubl. [FG] 

Rinorea lindeniana (Tul.) Kuntze var. lindeniana [GU SU] 

Rinorea macrocarpa (Mart. ex Eichl.) Kuntze [GU SU FG] 

Rinorea melanodonta S.F. Blake [GU] 

Rinorea neglecta Sandw. [GU SU FG] 

Rinorea paniculata (Mart.) Kuntze [FG] 

Rinorea pectino-squamata Hekking [FG] 

Rinorea pubiflora (Benth.) Sprague & Sandw. var. grandifolia 

(Eichl.) Hekking [GU SU FG] 

Rinorea pubiflora (Benth.) Sprague & Sandw. var. pubiflora 

[GU SU FG] 

Rinorea riana Kuntze [GU SU FG] 

Rinoreocarpus ulei (Melchior) Ducke [SU FG] 

VIOLACEAE - SYNONYMS AND 


Alsodeia brevipes Benth. =Rinorea brevipes 

Alsodeia flavescens (Aubl.) Spreng. =Rinorea flavescens 

Alsodeia guianensis (Aubl.) Eichl. var. brevipes (Benth.) 

Eichl. =Rinorea brevipes 

Alsodeia guianensis (Aubl.) Eichl. var. laxiflora (Benth.) 

Eichl. =Rinorea brevipes 

Alsodeia guianensis (Aubl.) Eichl. var. lindeniana (Tul.) Eichl 

=Rinorea lindeniana 

Alsodeia guianensis (Aubl.) Eichl. var. parviflora Eichl. 

=Rinorea falcata 

Alsodeia laxiflora Benth. =Rinorea brevipes 

Alsodeia longiflora Oudemans =Amphirrhox longiflora 

Alsodeia martini Turcz. =Rinorea pubiflora var. pubiflora 

Alsodeia pallida Klotzsch ex Schomb. [nomen nudum] 

Alsodeia piparea Spreng. =Casearia commersoniana 

(Flacourtiaceae) 

Alsodeia prunifolia Spreng. =Rinorea riana 

Alsodeia pubiflora Benth. =Rinorea pubiflora 

Alsodeia riana (DC. ex Gingins) Turcz. =Rinorea riana 

Amphirrhox surinamensis Eichl. =Amphirrhox longifolia 

Conohoria aubletii D. Dietr. =Casearia commersoniana 

(Flacourtiaceae) 

Conohoria brevipes (Benth.) Miq. =Rinorea brevipes 

SPERMATOPHYTA: Violaceae - Viscaceae 

Conohoria flavescens Aubl. =Rinorea flavescens 

Conohoria laxiflora (Benth.) Miq. =Rinorea brevipes 

Conohoria passoura DC. ex Gingins =Rinorea pubiflora var. 

pubiflora 

Conohoria riana DC. ex Gingins =Rinorea riana 

Hybanthus ipecacuanha (L. non Vent.) Baill. ex Laness 

=Hybanthus calceolaria 

Hybanthus poaya (St. Hil.) Baill. [not in Guianas] 

Ionidium ipecacuanha (L.) Vent. =Hybanthus calceolaria 

Ionidium oppositifolium (L.) Roem. & Schult. =Hybanthus 

oppositifolius 

Leonia melinoniana (Baill.) Baill. ex Sagot =Leonia 

glycycarpa 

Noisettia longifolia Kunth =Noisettia orchidiflora 

Passoura guianensis Aubl. =Rinorea pubiflora var. pubiflora 

Riana guianensis Aubl. =Rinorea riana 

Rinorea laxiflora (Benth.) Melchior =Rinorea brevipes 

Rinorea martini (Turcz.) Blake =Rinorea pubiflora var. 

pubiflora 

Rinorea passoura (DC. ex Gingins) Kuntze, nom. illeg. 

=Rinorea pubiflora var. pubiflora 

Rinorea passoura Kuntze =Rinorea pubiflora var. pubiflora 

Rinorea passoura (DC. ex Gingins) Kuntze var. andersonii 

Sandw. ex Hekking =Rinorea pubiflora var. grandifolia 

Rinorea pulleana Melchior =Rinorea pubiflora var. grandifolia 

Rinorea riana (DC. ex Gingins) Kuntze, nom. illeg. =Rinorea 

riana 

Rinorea surinamensis Melchior =Rinorea falcata 

Viola hybanthus Aubl. =Corynostylis arborea 

Viola itoubou Aubl. =Hybanthus calceolaria 

VISCACEAE 


REFERENCES: 

Krause, K. 1932. Loranthaceae. In A. Pulle, 


Kuijt, J. 1961. A revision of Dendrophthora. 

Wentia 6: 1-145. 


Loranthaceae (additions and corrections). In 


1(2): 295-300. 

Rizzini, C.T. 1978. Loranthaceae. In B. 


Guayana Highland, part 10. Mem. New York 

Bot. Gard. 29: 23-36. 

Rizzini, C.T. 1982. Loranthaceae. In Z. Luces 

de Febres, Flora de Venezuela 4(2): 7-316. 

Dendrophthora decipiens Kuijt [to be expected in the Guianas] 

Dendrophthora densifrons (Ule) Kuijt, comb. nov. ined. [to be 


Dendrophthora elliptica (Gardn.) Krug & Urb. [GU] 

Dendrophthora roraimae (Oliver) Ule [GU] 

Dendrophthora tepuiana (Steyerm.) Kuijt [GU?] 

Phoradendron acinacifolium Mart. ex Eichl. [GU SU]


Phoradendron aphyllum Steyerm. [GU] 

Phoradendron bathyoryctum Eichl. [GU SU] 

Phoradendron chrysocladon A. Gray [GU] 

Phoradendron crassifolium (Pohl ex DC.) Eichl. [GU SU FG] 

Phoradendron dimidiatum (Miq.) Eichl. [GU SU FG] 

Phoradendron dipterum (Eichl.) [GU SU] 

Phoradendron fanshawei Maguire [GU SU?] 

Phoradendron harmsianum Ule [GU SU] 

Phoradendron inaequidentatum Rusby [GU SU FG] 

Phoradendron morsicatum Rizzini [GU] 

Phoradendron mucronatum (DC.) Krug & Urb. [GU] 

Phoradendron nervosum Oliver [GU?] 

Phoradendron obtusissimum (Miq.) Eichl. [GU SU] 

Phoradendron perfurcatum Rizzini [GU] 

Phoradendron perrottetii (DC.) Eichl. [GU SU? FG] 

Phoradendron piperoides (Kunth) Trel. [GU SU FG] 

Phoradendron pteroneuron Eichl. [GU SU] 

Phoradendron pulleanum Krause [SU] 

Phoradendron quadrangulare (Kunth) Krug & Urb. [GU] 

Phoradendron racemosum (Aubl.) Krug & Urb. [GU SU FG] 

Phoradendron strongyloclados Eichl. [GU SU] 

Phoradendron trinervium (Lam.) Griseb. [GU SU] 

VISCACEAE - SYNONYMS AND 


Phoradendron essequibense Trel. =Phoradendron 

strongyloclados 

Phoradendron flavens (Swartz) Griseb. =Phoradendron 

chrysocladon 

Phoradendron jenmanii Trel. =Phoradendron inaequidentatum 

Phoradendron pennivenium (DC.) Eichl. =Oryctanthus 

florulentus (Loranthaceae) 

Phoradendron platycaulon Eichl. [not in Guianas] 

Phoradendron schottii A. Gray =Phoradendron piperoides 

Phoradendron surinamense Pulle =Phoradendron 

strongyloclados 

Phoradendron tepuianum Steyerm. =Dendrophthora tepuiana 

Phoradendron zuloagae Trel. =Phoradendron trinervium 

REFERENCES: 

VITACEAE 

Descoings, B. 1991. Contribution à l'étude des 

Vitacées d'Amerique tropicale: deux Cissus 

nouveaux des Guyanes. Bull. Soc. Bot. 

France, Lett. Bot. 138(3): 249-256. 

Görts-van Rijn, A.R.A. 1979. Vitaceae. In A.L. 


5(1): 335-343. 

Lombardi, J.A. 1995. Typification of names of 

South American Cissus (Vitaceae). Taxon 

44(2): 193-206. 

Suessenguth, K. 1953. Vitaceae. In A. Engler & 


20d: 174-333. 

Cissus erosa L.C. Rich. [GU SU FG] 

Cissus guyanensis Descoings [FG] 

Cissus haematantha Miq. [SU FG] 

Cissus spinosa Cambess. [GU SU FG] 

Cissus subrhomboidea (Baker) Planch. [GU SU] 

Cissus surinamensis Descoings [SU FG] 

Cissus ulmifolia (Baker) Planch. [SU] 

Cissus venezuelensis Steyerm. [GU] 

Cissus verticillata (L.) Nicolson & Jarvis [GU SU FG] 

Vitis tiliifolia Willd. ex Roem. & Schult. [FG?] 

VITACEAE - SYNONYMS AND 


Cissus cordifolia L. =Cissus verticillata 

Cissus elongata Miq. =Cissus erosa 

Cissus microcarpa Vahl =Cissus erosa 

Cissus parkeri (Baker) Planch. =Cissus spinosa 

Cissus salutaris Kunth =Cissus erosa 

Cissus sicyoides L. =Cissus verticillata 

Cissus trifoliata (L.) L. =Cissus erosa? 

Vitis erosa Baker =Cissus erosa 

Vitis miqueliana Baker =Cissus erosa 

Vitis parkeri Baker =Cissus spinosa 

Vitis sicyoides (L.) Baker =Cissus verticillata 

Vitis spinosa (Cambess.) Baker =Cissus spinosa 

VOCHYSIACEAE 

contributed by L. Marcano-Berti (1992) 

REFERENCES: 

SPERMATOPHYTA: Viscaceae - Vochysiaceae 

207 

Lleras, E. 1976. Revision and taxonomic 

position of the genus Euphronia Martius ex 

Martius & Zuccarini (Vochysiaceae). Acta 

Amaz. 6: 43-47. 

Marcano-Berti, L. 1969. Un nuevo genero de las 

Vochysiaceae. Pittieria 2: 3-27. 

Marcano-Berti, L. 1989. Vochysiaceae: 

novedades y correciones. Pittieria 18: 5-14. 

Marcano-Berti, L. 1989. Euphroniaceae: una 

nueva familia. Pittieria 18: 15-17. 

Stafleu, F.A. 1948-1954. A monograph of the 

Vochysiaceae. I. Salvertia & Vochysia. Rec. 

Trav. Bot. Neerl. 41: 397-540(1948). III. 

Qualea. Acta Bot. Neerl. 2: 144-217 (1953). 

IV. Erisma. Acta Bot. Neerl. 3: 459-480 

(1954). 

Stafleu, F.A. 1951. Vochysiaceae. In A. Pulle, 


Steyermark, J.A. 1987. Vochysiaceae: a 

reevaluation of the genus Euphronia. In J.A. 

Steyermark, Flora of the Venezuelan Guayana 

- II. Ann. Missouri Bot. Gard. 74(1): 85-116. 

Erisma calcaratum (Link) Warm. [GU] 

Erisma floribundum Rudge var. floribundum [FG] 

Erisma nitidum DC. [GU FG] 

Erisma uncinatum Warm. [GU SU FG] 

Euphronia guianensis (Rob. Schomb.) H. Hallier [GU] 

[=Euphronia hirtelloides s.l.] 

Qualea acuminata Spruce ex Warm. [FG]

208 


Qualea amapaënsis Balslev & Mori [FG?] 

Qualea caerulea Aubl. [SU FG] 

Qualea dinizii Ducke [GU SU FG] 

Qualea grandiflora Mart. [SU] 

Qualea mori-boomii Marcano-Berti [FG] 

Qualea paraensis Ducke [GU] 

Qualea polychroma Stafl. [GU] 

Qualea psidiifolia Spruce ex Warm. [FG] 

Qualea rosea Aubl. [SU FG] 

Qualea schomburgkiana Warm. [GU] 

Qualea tricolor Benoist [FG] 

Ruizterania albiflora (Warm.) Marcano-Berti [GU SU FG] 

Ruizterania ferruginea (Steyerm.) Marcano-Berti [GU] 

Ruizterania rigida (Stafl.) Marcano-Berti [GU] 

Salvertia convallariodora A. St. Hil. [SU] 

Vochysia cayennensis Warm. [FG] 

Vochysia costata Warm. [GU] 

Vochysia crassifolia Warm. [GU] 

Vochysia densiflora Spruce ex Warm. [GU SU FG] 

Vochysia ferruginea Mart. [GU] 

Vochysia glaberrima Warm. [GU SU] 

Vochysia guianensis Aubl. [GU SU FG] 

Vochysia maguirei Marcano-Berti [GU] 

Vochysia neyratii Normand [FG] 

Vochysia sabatieri Marcano-Berti [FG] 

Vochysia schomburgkii Warm. [GU] 

Vochysia speciosa Warm. [FG] 

Vochysia surinamensis Stafl. var. surinamensis [GU SU FG] 

Vochysia tetraphylla (G. Mey.) DC. [GU SU FG] 

Vochysia tomentosa (G. Mey.) DC. [GU SU FG] 

Vochysia tucanorum Mart. var. tucanorum [SU] 

VOCHYSIACEAE - SYNONYMS AND 


Euphronia hirtelloides Mart. ex Mart. & Zucc. [not in 


Lightia guianensis Rob. Schomb. =Euphronia guianensis 

Qualea albiflora Warm. =Ruizterania albiflora 

Qualea ferruginea Steyerm. =Ruizterania ferruginea 

Qualea rigida Stafleu =Ruizterania rigida 

Qualea rubiginosa Stafleu =Ruizterania ferruginea 

Ruizterania rubiginosa (Stafl.) Marcano-Berti =Ruizterania 

ferruginea 

Vochysia pinkusii A.C. Smith [not in Guianas] 

Vochysia racemosa Poir. [name of uncertain application, not 

Vochysia] 

REFERENCES: 

WINTERACEAE 


Drimys. J. Arn. Arb. 24: 1-33. 

Drimys brasiliensis Miers var. roraimensis A.C. Smith [GU?] 

[Mt. Roraima] 

SPERMATOPHYTA: Vochysiaceae - Xyridaceae 

XYRIDACEAE 

reviewed by R. Kral (1992); reviewed by R. Kral 

(1996) 

REFERENCES: 

Kral, R. 1988. The genus Xyris in Venezuela 

and contiguous northern South America. 


Kral, R. 1992. A treatment of American 

Xyridaceae exclusive of Xyris. Ann. 


Kral, R. 1994. Xyridaceae. In A.R.A. Görts-van 

Rijn, Flora of the Guianas (series A) 15: 3-90. 

Lanjouw, J. 1938. Xyridaceae. In A. Pulle, 


Maguire, B. 1958. Xyridaceae. In B. Maguire, 


Guayana Highland part III. Mem. New York 

Bot. Gard. 10(1): 1-156. 

Maguire, B. & L.B. Smith. 1964. Xyridaceae. 

In B. Maguire, J.J. Wurdack, & collaborators, 

Botany of the Guayana Highland part V. 


Mennega, A.M.W., J.C. Lindeman, & A.R.A. 

Görts-van Rijn. 1968. Xyridaceae (additions 



Abolboda acaulis Maguire var. acaulis [GU] 

Abolboda acaulis Maguire var. scaposa Kral [GU] 

Abolboda americana (Aubl.) Lanj. [GU SU FG] 

Abolboda grandis Griseb. var. grandis [GU SU] 

Abolboda grandis Griseb. var. rigida Malme [GU SU] 

Abolboda killipii Lasser [GU SU] 

Abolboda macrostachya Spruce ex Malme var. macrostachya 

[GU] 

Abolboda macrostachya Spruce ex Malme var. robustior 

Steyerm. [GU] 

Abolboda poarchon Seub. [SU?] 

Abolboda pulchella Humb. & Bonpl. [GU SU FG] 

Orectanthe ptaritepuiana (Steyerm.) Maguire [GU] 

Orectanthe sceptrum (F. Oliver) Maguire subsp. sceptrum 

[GU] 

Xyris albescens Steyerm. [GU] [Mt. Roraima] 

Xyris anceps Lam. [GU? SU? FG] 

Xyris bicephala Gleason [GU] 

Xyris concinna N.E. Brown [GU] [Mt. Roraima] 

Xyris connosepala Lanj. & Lindeman [SU] 

Xyris cyperoides Gleason [GU SU] 

Xyris decussata Gleason [GU] [Mt. Roraima] 

Xyris fallax Malme [GU SU FG] 

Xyris guianensis Steud. [GU SU] 

Xyris hymenachne Mart. [GU] [Mt. Roraima] 

Xyris involucrata Nees [GU SU?] 

Xyris jupicai L.C. Rich. [GU SU FG] 

Xyris kukenaniana Kral [to be expected in GU] 

Xyris lacerata Pohl ex Seub. [to be expected in GU] 

Xyris lanulobractea Steyerm. [GU] 

Xyris laxifolia Mart. var. laxifolia [GU SU FG]


Xyris malmeana L.B. Smith [GU SU FG] 

Xyris paraensis Poepp. ex Kunth var. longiceps (Malme) L.B. 

Smith & Downs [GU SU FG] 

Xyris paraensis Poepp. ex Kunth var. paraensis [GU SU FG] 

Xyris roraimae Malme [GU] [Mt. Roraima] 

Xyris savanensis Miq. [GU SU FG] 

Xyris setigera F. Oliver [GU] 

Xyris seubertii Nilsson [GU] [Mt. Roraima] 

Xyris spathacea Lanj. [GU? SU FG] 

Xyris subuniflora Malme [GU SU] 

Xyris surinamensis Spreng. [GU SU FG?] 

Xyris tenella Kunth var. leprieurii Malme [FG] 

Xyris tenella Kunth var. tenella [GU?] 

Xyris thysanolepis Maguire & L.B. Smith var. sipapoa 

(Maguire & L.B. Smith) Kral & L.B. Smith [to be 


Xyris thysanolepis Maguire & L.B. Smith var. thysanolepis [to 


Xyris uleana Malme var. angustifolia Lanj. [GU SU] 

Xyris uleana Malme var. uleana [GU SU] 

Xyris witsenioides F. Oliver [GU] [Mt. Roraima] 

XYRIDACEAE - SYNONYMS AND 


Abolboda grandis Griseb. var. guayanensis Maguire 

=Abolboda grandis var. rigida 

Abolboda grandis Griseb. var. minor Spruce ex Malme 

=Abolboda grandis var. grandis 

Abolboda imberbis Kunth =Abolboda americana 

Abolboda psammophila Maguire =Abolboda killipii 

Abolboda ptaritepuiana Steyerm. =Orectanthe ptaritepuiana 

Abolboda rigida Gleason =Abolboda macrostachya var. 

robustior 

Abolboda sceptrum F. Oliver =Orectanthe sceptrum 

Xyris acuminata Miq. ex Steud. =Xyris jupicai 

Xyris americana Aubl. =Abolboda americana 

Xyris asterocephala Seub. =Xyris involucrata 

Xyris caroliniana Walter var. major (Mart.) Idrobo & L.B. 

Smith =Xyris laxifolia 

Xyris communis Kunth =Xyris jupicai 

Xyris connectens Malme =Xyris uleana var. angustifolia 

Xyris dolichosperma Lanj. =Xyris fallax 

Xyris eriophylla Rchb. ex Kunth =Xyris surinamensis 

Xyris erythema Maguire & L.B. Smith =Xyris fallax 

Xyris glabrata (Seub.) Griseb. sensu Griseb. =Xyris malmeana 

Xyris guianensis Klotzsch [nomen nudum] =Xyris jupicai 

Xyris jupicai L.C. Rich. var. major (Mart.) L.B. Smith & 

Downs =Xyris laxifolia 

Xyris leptostachya Malme =Xyris paraensis var. longiceps 

Xyris longiceps Malme =Xyris paraensis var. longiceps 

Xyris macrocephala Auct. non Vahl =Xyris laxifolia 

Xyris macrocephala Vahl var. major (Mart.) A. Nilsson 

=Xyris laxifolia 

Xyris submontana Gleason =Xyris hymenachne 

Xyris surinamensis Miq. =Xyris jupicai 

ZAMIACEAE 


REFERENCES: 

Jones, D.L. 1993. Cycads of the World. 

Washington, DC: Smithsonian Institution 

Press. 

Sabato, S. 1990. West Indian and South 

American Cycads. Mem. New York Bot. 

Gard. 57: 173-185. 

Stevenson, D. 1991. Zamiaceae. In A.R.A. 


A) 9: 7-11. 

Zamia lecointei Ducke [to be expected in the Guianas] 

Zamia muricata Willd. [to be expected in the Guianas] 

Zamia ulei U. Dammer [to be expected in the Guianas] 

ZINGIBERACEAE 


P.J.M. Maas (1996) 

REFERENCES: 

SPERMATOPHYTA: Xyridaceae - Zingiberaceae 

209 

Maas, P.J.M. 1977. Renealmia. Flora 


Maas, P.J.M. 1979. Zingiberaceae. In A.L. 


5(1): 416-441. 

Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. 


A). 

Hedychium coronarium J. Koenig [GU SU FG] [cultivated, 

naturalized] 

Renealmia alpinia (Rottb.) Maas [GU SU FG] 

Renealmia aromatica (Aubl.) Griseb. [GU] 

Renealmia cernua (Swartz ex Roem. & Schult.) Macbr. [SU] 

Renealmia floribunda K. Schum. [GU SU FG] 

Renealmia guianensis Maas [GU SU FG] 

Renealmia monosperma Miq. [GU SU FG] 

Renealmia orinocensis Rusby [GU SU FG] 

Renealmia thyrsoidea (Ruiz & Pav.) Poepp. & Endl. subsp. 

chrysantha Maas [GU SU] 

Zingiber spp. [several species cultivated, possibly escaping] 

ZINGIBERACEAE - SYNONYMS AND 


Alpinia aromatica Aubl. =Renealmia aromatica 

Alpinia tubulata Ker-Gawl. =Renealmia alpinia 

Renealmia exaltata L.f. =Renealmia alpinia 

Renealmia gracilis O.G. Petersen =Renealmia guianensis 

Renealmia humilis O.G. Petersen =Renealmia monosperma 

Renealmia occidentalis (Swartz) Sweet =Renealmia aromatica 

Renealmia occidentalis (Swartz) Sweet var. cylindrica O.G. 

Petersen =Renealmia aromatica 

Renealmia occidentalis (Swartz) Sweet var. macrocarpa Loes. 

=Renealmia aromatica

210 


Renealmia pedicellaris Gleason =Renealmia floribunda 

REFERENCES: 

ZOSTERACEAE 

Phillips, R.C. & E.G. Meñez. 1988. Seagrasses. 

Smiths. Contrib. Marine Sci. 34: 1-104. 

Zostera marina L. [FG?] [the record by Lemée must be 

considered extremely dubious] 

ZYGOPHYLLACEAE 

reviewed by B.E. Leuenberger (1992); reviewed 

by B.E. Leuenberger (1996) 

REFERENCES: 

Howard, R.A. 1988. Zygophyllaceae. In R.A. 

Howard, Flora of the Lesser Antilles 4(1): 

551-555. 

Lasser, T. 1971. Zygophyllaceae. In T. Lasser, 


Stafleu, F.A. 1951. Zygophyllaceae. In A. 


Guaiacum officinale L. [GU SU] 

Kallstroemia maxima (L.) Hook. & Arnott [FG] 

Tribulus cistoides L. [GU SU] 

SPECIES INCERTAE SEDIS 

Aubletella macoucou (Aubl.) Pierre [FG] [Chrysophyllum 

macoucou Aubl.] 

Macahanea guianensis Aubl. [FG] [Salacia guianensis (Aubl.) 

Lemée] 

Managa guianensis Aubl. [FG] 

Montira guianensis Aubl. [FG] [Spigelia guianensis (Aubl.) 

Lemée] 

Sarcaulus melinonianum Pierre [FG] 

Senapea guianensis Aubl. [FG] 

Tampoa guianensis Aubl. [FG] 

Tapirocarpus talisia Sagot [FG] 

Vouapa simira Aubl. [FG] [Macrolobium simira (Aubl.) J.F. 

Gmelin] 

SPERMATOPHYTA: Zingerberaceae - Zygophyllaceae

A 

Abarema 140, 142 

Abelmoschus 129 

Aberemoa 47 

Abildgaardia 91 

Ablania 98 

Abolboda 208, 209 

Abrus 105 

Abuta 139, 140 

Abutilon 129 

Acacallis 154 

Acacia 141, 142 

Acalypha 101, 102 

ACANTHACEAE 41-43 

Acanthocoleus 11 

Acanthospermum 59, 60 

Acanthosphaera 145 

Acetosella 163 

Achetaria 195 

Achras 193 

Achrouteria 193 

Achyranthes 44 

Achyrocline 59 

Acineta 154 

Acioa 81 

Aciotis 131, 134 

Acisanthera 132, 134 

Acmella 59, 60 

Acnida 44 

Acosmium 105 

Acouroa 107 

ACROBOLBACEAE 9 

Acroceras 168 

Acrocomia 54, 56 

Acrodiclidium 119 

Acrolejeunea 11, 13 

Acroporium 21 

Acrostichum 37 

Actinostachys 38 

Actinostemon 101, 102 

Acuan 142 

Ada 154 

ADELANTHACEAE 9 

Adelanthus 9 

Adelobotrys 132, 134 

Adenanthe 152 

Adenanthera 141 

Adenocalymna 63, 64 

Adenophaedra 101 

Adenostemma 60 

Adenostephanus 179 

ADIANTACEAE 24-25 

Adiantopsis 25 

Adiantum 25 

Aechmea 67, 68 

Aegiphila 204, 205 


INDEX 

Aegopodon 168 

Aeschrion 196 

Aeschynomene 105, 107 

Agalinis 195 

Aganisia 154, 160 

Agarista 98 

Agati 107 

AGAVACEAE 43 

Ageratina 59 

Ageratum 59 

Agonandra 154 

Agrostis 171 

Aiouea 118, 120 

AIZOACEAE 43 

Akentra 123 

Albizia 141, 142 

Alchornea 101, 102 

Alchorneopsis 101, 102 

Aldina 72 

Alectra 195 

Alexa 105, 107 

Alexandra 107 

Alibertia 181, 186 

Alicastrum 145 

Alisma 44 

ALISMATACEAE 43-44 

Aliteria 145 

Allamanda 47 

Allania 75 

Allophylus 190, 191 

Alloplectus 111 

Allosidastrum 129 

Almeidea 190 

Alobiella 10 

Alocasia 51 

Alophia 117 

Alpinia 209 

Alseis 182 

Alsodeia 206 

Alsophila 26 

Alstroemeria 124 

Alternanthera 44 

Alysicarpus 105 

Amaioua 182 

Amanoa 101, 102 

AMARANTHACEAE 44 

Amaranthus 44 

Amaryllis 124 

Amasonia 205 

Ambelania 48, 49 

Ambrosia 59, 60 

Amerimnum 107 

Ammannia 126 

Amorphospermum 193 

Ampelocera 204 

Ampelozizyphus 180 

Amphidesmium 35 

Amphilophium 63 

Amphiodon 107 

Amphirrhox 206 

Amphizoma 114 

Amyris 189 

Anacampta 49 

ANACARDIACEAE 45 

Anacardium 45 

Anadenanthera 141 

Ananas 67, 69 

Ananthacorus 40 

Anaphyllopsis 51 

Anartia 49 

Anastrophyllum 11 

Anatherum 171 

Anaxagorea 46, 47 

Ancistrothyrsus 164 

Andira 105, 107 

Andreaea 16 

ANDREAEACEAE 16 

Andriapetalum 179 

Andropogon 168, 171 

Anechites 49 

Aneilema 86 

Anemia 38 

Anemopaegma 63, 64 

Anetium 40 

Aneura 10 

ANEURACEAE 10 

Angelonia 195 

Angostura 190 

Angostyles 102 

Angraecum 160 

Anguria 89 

Aniba 118, 120 

Anisacanthus 41 

Anisantherina 195 

Aniseia 87 

Anisolobus 49 

Anisomeris 186 

Anisophyllea 45 

ANISOPHYLLEACEAE 45 

Anneslia 142 

Annona 46, 47 

ANNONACEAE 45-47 

Anomoclada 10 

Anomoctenium 64 

Anomospermum 139, 140 

Anonocarpus 145 

Anoplolejeunea 

Anredera 62 

Anthephora 168 

ANTHOCEROPHYTA 15 

Anthodiscus 78 

Anthodon 113, 114 

Anthurium 51, 53 

Antidaphne 98 

211 

INDEX

212 

Antidesma 101 

Antigonon 177 

Antirhea 182 

Antonia 125 

Antrophyum 40 

Apalanthe 116 

Apalatoa 75 

Aparisthmium 101 

Apatitia 134 

Apeiba 201, 202 

Aphanolejeunea 11 

Aphelandra 41 

APIACEAE 47 

Apinagia 175, 176 

Apluda 171 

APOCYNACEAE 47-50 

Apocynum 49 

Apodandra 102 

Apodanthera 89 

Apodanthes 179 

Appendicularia 132 

Appunia 186 

Aptandra 153 

Apteria 70 

AQUIFOLIACEAE 50-51 

ARACEAE 51-53 

Arachniodes 29 

Arachniopsis 13 

Araeococcus 67 

ARALIACEAE 53-54 

Arberella 168 

ARCHIDIACEAE 16 

Archidium 16 

Archilejeunea 11, 13 

Archytaea 66 

Ardisia 147 

ARECACEAE 54-57 

Argemone 163 

Aristida 168, 171 

Aristolochia 57 

ARISTOLOCHIACEAE 57 

Arouna 75 

Arrabidaea 63, 64 

Arrhostoxylum 42 

Artanthe 166 

Arthrosamanea 142 

Arthrostemma 134 

Arthrostylidium 168, 171 

Aruba 196 

Arunda 171 

Arundinaria 171 

Arundinella 168 

Arundo 168, 171 

ASCLEPIADACEAE 57-58 

Asclepias 57 

Ascolepis 91 

Aspasia 155 

Aspidium 29, 35, 38, 39 

Aspidogyne 155 

Aspidosperma 48, 49 

INDEX 


ASPLENIACEAE 25-26 

Asplenium 25-26 

Asplundia 90 

Aster 60 

ASTERACEAE 58-61 

Asterella 10 

Asterogyne 54, 56 

Asterophorum 202 

Astrocaryum 54, 56 

Astronium 45 

Asystasia 41 

Athyrocarpus 86 

Atractylocarpus 17 

Attalea 56 

Aubletella 210 

Aulizia 160 

Aulomyrcia 150 

Aulonemia 168 

Aureolejeunea 13 

Austroeupatorium 59 

Avicennia 205 

Axonopus 168, 171 

Ayapana 59 

Ayenia 199 

AYTONIACEAE 10 

Azadirachta 138 

Azolla 38 

B 

Baccharis 59 

Bacopa 195 

Bactris 55, 56 

Bagassa 144, 145 

Baillieria 60 

BALANOPHORACEAE 62 

BALANTIOPSACEAE 10 

Balizia 141 

BALSAMINACEAE 62 

Baltimora 59 

Bambusa 172 

Banara 109 

Banisteria 128 

Banisteriopsis 127, 128 

Barbacenia 204 

Barbula 20 

Barjonia 57 

Barleria 41 

Barnhartia 176 

BARTRAMIACEAE 16 

Basanacantha 186 

Basella 62 

BASELLACEAE 62 

Baskervilla 155 

Bassovia 198 

Bastardia 129 

BATACEAE 62 

Batemannia 155 

Batesia 72 

Bathmium 39 

Batis 62 

Batocarpus 144 

Bauhinia 72, 75 

Bazzania 13 

Becquerelia 91, 94 

Befaria 98 

Begonia 62 

BEGONIACEAE 62 

Beilschmiedia 119 

Bejaria 98 

Bellucia 132, 135 

Bellynkxia 186 

Beloglottis 155 

Beloperone 42 

Benjaminia 195 

Bernardia 101 

Bernardinia 87 

Bertholletia 121 

Bertiera 182 

Besleria 111 

Beyrichia 196 

Bidens 59 

Bifrenaria 155, 160 

Bignonia 64 

BIGNONIACEAE 63-65 

Bihai 113 

Billbergia 67 

Biovularia 123 

Bisboeckelera 91, 94 

Bixa 65 

BIXACEAE 65 

Blakea 135 

BLECHNACEAE 26 

Blechnum 26 

Blechum 41 

Blepharandra 127 

Blepharocalyx 148 

Blepharodon 57 

Bletia 155, 160 

Blutaparon 44 

Bocagea 47 

Bocageopsis 46 

Bocoa 73 

Boehmeria 204 

Boerhavia 151 

Bolbitis 34 

Bolboschoenus 94 

Bollea 155, 160 

Bomarea 124 

BOMBACACEAE 65 

Bombacopsis 65 

Bombax 65 

Bonafousia 49 

Bonamia 87 

Bonatea 160 

Bonnetia 66 

BONNETIACEAE 65-66 

Bontia 146 

Bonyunia 125 

BORAGINACEAE 66

Borismene 139 

Borreria 182, 186 

Bothriochloa 168, 172 

Bothriospora 182 

Bouganvillea 151 

Boussingaltia 62 

Bouteloua 168 

Bowdichia 105, 107 

Boyania 132 

Brachiaria 168, 172 

Brachiolejeunea 13 

Brachiolejeunia 11 

Brachionidium 155 

Brachymenium 16 

Brachypremna 135 

Brachypterys 128 

Brachystele 155 

BRACHYTHECIACEAE 16 

Braemia 155 

Bramia 196 

Brasenia 71 

Brassavola 155, 160 

Brassia 155, 160 

Brassica 66 

BRASSICACEAE 66-67 

Bredemeyera 176 

Breutelia 16 

Brignolia 186 

Briquetia 129 

Brocchinia 67, 69 

Bromelia 67, 69 

BROMELIACEAE 67-69 

Brosimopsis 145 

Brosimum 144, 145 

Brossea 99 

Brownea 73 

Bruinsmania 186 

Brunellia 70 

BRUNELLIACEAE 70 

Brunfelsia 197, 198 

BRYACEAE 16 

Brymela 18 

Bryohumbertia 17 

Bryophyllum 88 

BRYOPHYTA 15-22 

Bryopteris 11, 13 

Bryum 16 

Buceras 86 

Buchenavia 85, 86 

Buchnera 195, 196 

Bucida 86 

Buforrestia 86 

Bulbophyllum 155 

Bulbostylis 91, 94 

Bulbulus 172 

Bumelia 193 

Bunchosia 127, 128 

Burdachia 127 

Burlingtonia 160 

Burmannia 70 


BURMANNIACEAE 70 

Bursera 70, 71 

BURSERACEAE 70-71 

Buttneria 199 

Byrsonima 127, 128 

Byttneria 199 

C 

Cabomba 71 

CABOMBACEAE 71 

Cabralea 138 

Cacao 199 

Cacoucia 86 

CACTACEAE 71-72 

Caesalpinia 73 

CAESALPINIACEAE 72-77 

Cajanus 105, 107 

Caladium 51, 53 

Calathea 130 

Calea 59, 60 

Calinea 96 

Calliandra 141, 142 

Callichlamys 63 

Callicosta 18 

Callicostella 18 

Callisia 86 

Calocarpum 193 

Calolisianthus 110 

Calophyllum 83 

Calopogonium 105 

Calotropis 58 

Caluera 155 

Calycolobus 87 

Calycolpus 148, 150 

Calycophyllum 186 

Calycorectes 148 

CALYMPERACEAE 16-17 

Calymperes 16 

Calymperopsis 17 

Calypogeia 10 

CALYPOGEIACEAE 10 

Calyptranthes 148, 150 

Calyptrocarya 91, 94 

Camarea 127 

Cameraria 50 

CAMPANULACEAE 77 

Campomanesia 148, 150 

Campsiandra 73, 75 

Campylocentrum 155, 160 

Campyloneurum 36 

Campylopus 17 

Campylosiphon 70 

Cananga 47 

Canavalia 105, 107 

Candolleodendron 73 

CANELLACEAE 77 

Canna 77 

CANNACEAE 77 

Caopia 85 

Caperonia 101, 102 

Capirona 182, 186 

Capitellaria 135 

CAPPARACEAE 77-78 

Capparidastrum 78 

Capparis 77, 78 

Capraria 195, 196 

CAPRIFOLIACEAE 78 

Capriola 172 

Capsicum 197, 198 

Caraipa 83, 85 

Carapa 138 

Carapichea 186 

Cardiopetalum 46 

Cardiospermum 190 

Carex 91 

Carica 78 

CARICACEAE 78 

Carludovica 90 

Carpotroche 109 

Carptotepala 99 

Caryocar 78 

CARYOCARACEAE 78-79 

Caryomene 139 

Caryophyllaceae 

Casearia 109 

Casimirella 116 

Cassia 73, 75 

Cassipourea 181 

Cassytha 119, 120 

Castalia 151 

Casuarina 79 

CASUARINACEAE 79 

Catasetum 155, 160 

Catharanthus 48 

Cathedra 153 

Catinga 150 

Catocoma 177 

Catopsis 67, 69 

Catostemma 65 

Cattleya 155, 160 

Caudalejeunea 11 

Caularthron 155 

Cavendishia 98, 99 

Cayaponia 89 

Cecropia 79 

CECROPIACEAE 79-80 

Cedrela 138 

Cedrelinga 141 

Ceiba 65 

CELASTRACEAE 80 

Celiantha 110 

Celosia 44 

Celtis 204 

Cenchrus 168, 172 

Centratherum 59 

Centrolobium 105 

Centronia 135 

Centropogon 77 

Centrosema 105, 107 

213 

INDEX

214 

Centrosolenia 111 

Cephaelis 186 

Cephalocarpus 91 

Cephalocereus 72 

Cephalostemon 180 

Cephalozia 10 

CEPHALOZIACEAE 10 

CEPHALOZIELLACEAE 10 

Ceradenia 30 

Ceratolejeunea 11 

CERATOPHYLLACEAE 80 

Ceratophyllum 80 

Ceratophytum 63, 64 

Ceratopteris 36 

Ceratosanthes 89 

Ceratostema 98 

Cereus 72 

Cespedesia 152, 153 

Cestrum 197, 198 

Chaenopleura 135 

Chaetocalyx 105 

Chaetocarpus 101, 102 

Chaetochlamys 42 

Chaetochloa 172 

Chaetogastra 135 

Chaetolepis 132 

Chailletia 95 

Chalepophyllum 182, 187 

Chamaecrista 73 

Chamaedorea 56 

Chamaesyce 101, 102 

Chamelophyton 155 

Chamissoa 44 

Chaptalia 59 

Chara 9 

CHARACEAE 9 

Charianthus 135 

Chaubardia 155, 160 

Chaubardiella 155 

Chaunochiton 153 

Cheiloclinium 113, 114 

Cheilolejeunea 11 

Cheiradenia 155, 160 

Cheiroglossa 35 

Chelonanthus 110 

CHENOPODIACEAE 80 

Chenopodium 80 

Chevaliera 69 

Chimarrhis 182, 187 

Chiococca 182, 187 

Chionanthus 153 

Chitonia 135 

CHLORANTHACEAE 80 

Chloris 168, 172 

Chlorocardium 119 

Chloroleucon 141 

Chlorophora 145 

CHLOROPHYTA 9 

Chomelia 182 

Chondrodendron 140 

INDEX 


Chorisepalum 110 

Christiana 201 

Chromolaena 59 

Chromolucuma 192 

CHRYSOBALANACEAE 80-83 

Chrysobalanus 81, 82 

Chrysochlamys 85 

Chrysohypnum 19 

Chrysophyllum 192, 193 

Chrysothemis 111 

Chusquea 168 

Chytroma 121 

Chytropsia 187 

Cienfuegosia 129 

Cinnamodendron 77 

Cinnamomum 119 

Ciponima 200 

Cipura 117 

Cissampelos 139 

Cissus 207 

Citharexylum 205 

Citriosma 144 

Citrullus 89 

Citrus 189 

Cladium 91 

Cladobium 160 

Cladocolea 125, 126 

Clarisia 145 

Clathrotropis 105, 107 

Clavapetalum 116 

Clavija 201 

Cleidion 102 

Cleistes 155 

Clematis 179 

Cleome 77, 78 

Clerodendrum 205 

Clethra 83 

CLETHRACEAE 83 

Clibadium 59, 60 

Clidemia 132, 135 

Clinostemon 120 

Clitoria 105, 107 

Clowesia 155 

Clusia 83, 85 

CLUSIACEAE 83-85 

Clusiella 84 

Clytostoma 63 

Cnemidaria 26 

Cnestidium 86 

Cnidoscolus 101 

Coccinia 89 

Coccocypselum 182 

Coccoloba 177 

Cochleanthes 155, 160 

Cochlidium 30 

Cochlospermum 65 

Cocos 55 

Codonanthe 111, 112 

Codonanthopsis 111, 112 

Coeloachis 168 

Coffea 182, 187 

Coix 168 

Coleostachys 127 

Colocasia 52 

Cololejeunea 11 

Colubrina 180 

Columnea 111, 112 

Colura 11 

COMBRETACEAE 85-86 

Combretum 85, 86 

Commelina 86 

COMMELINACEAE 86 

Commelinopsis 86 

Comolia 132, 135 

Comparettia 155 

Compsoneura 147 

Conami 102 

Conceveiba 101 

Conceveibum 102 

Conchocarpus 189 

Condylidium 59 

Condylocarpon 48, 50 

CONNARACEAE 86-87 

Connarus 86, 87 

Connelia 67 

Connellia 67 

Conobea 195 

Conocarpus 85 

Conohoria 206 

Conomorpha 147-148 

Conostegia 135 

Conuleum 144 

CONVOLVULACEAE 87-88 

Convolvulus 88 

Conyza 59, 61 

Copaifera 73 

Copedesma 135 

Copernicia 56 

Corchorus 201, 202 

Cordia 66 

Cornutia 205 

Coronilla 107 

Coronopus 66 

Cortaderia 168 

Coryanthes 155 

Corymborkis 155 

Corynostylis 206 

Coryphadenis 135 

Corythophora 121 

Corytoplectus 111 

Cosmibuena 182, 187 

Cosmos 59 

COSTACEAE 88 

Costus 88 

Cottendorfia 69 

Coublandia 107 

Couepia 81, 82 

Couma 48 

Coumarouna 107 

Coupoui 187

Couratari 121 

Courimari 98 

Couroupita 121 

Coursetia 105 

Coussapoa 79 

Coussarea 182, 187 

Coutarea 187 

Coutoubea 110 

Cranichis 155 

Craniolaria 164 

Crantzia 112 

CRASSULACEAE 88-89 

Crateva 78 

Cremastosperma 46, 47 

Crenea 126 

Crepidospermum 70 

Crepinella 54 

Crescentia 63 

Crinum 124 

Crossomitrium 18 

Crossotolejeunea 12 

Crotalaria 105, 108 

Croton 101, 102 

Crudia 73, 76 

Cryptarrhena 155, 160 

Cryptocalyx 205 

Cryptocarya 119, 120 

Cryptocentrum 155 

Cryptochloa 168, 172 

Ctenitis 38 

Ctenium 168 

Ctenopteris 31 

Cucumis 89 

Cucurbita 89 

CUCURBITACEAE 89 

Cuervea 114 

Culcita 28 

CUNONIACEAE 89-90 

Cunuria 103 

Cupania 190 

Cuphea 126 

Cupirana 187 

Curanga 196 

Curarea 140 

Curatella 96 

Curculigo 124 

Curtia 110 

Cuscuta 90 

CUSCUTACEAE 90 

Cusparia 190 

Cuspidaria 63 

Cyanthillium 59 

Cyathea 26-27 

CYATHEACEAE 26-27 

Cyathula 44 

Cybianthus 147, 148 

Cybistax 63 

CYCADACEAE 90 

Cycas 90 

CYCLANTHACEAE 90 


Cyclanthus 90 

Cyclodictyon 18 

Cyclodium 29 

Cyclolejeunea 12, 13 

Cyclolobium 108 

Cyclopeltis 38 

Cyclopogon 155, 160 

Cyclosorus 39 

Cycnoches 155 

Cydista 63 

Cylindrocolea 10 

Cymbocarpa 70 

Cymbopetalum 46 

Cymbopogon 168, 172 

Cymbosema 105, 108 

Cynanchum 58 

Cynodendron 193 

Cynodon 168 

Cynometra 73 

Cynomorium 62 

CYPERACEAE 91-95 

Cyperus 91, 94 

Cyphomandra 197, 198 

Cypripedium 160 

Cyrilla 95 

CYRILLACEAE 95 

Cyrillopsis 117 

Cyrtocarpa 45 

Cyrtocymura 59 

Cyrtohypnum 22 

Cyrtolejeunea 12 

Cyrtopera 160 

Cyrtopodium 155, 160 

Cyrtosperma 52, 53 

Cystolejeunea 12 

Cytisus 108 

D 

Dacryodes 70 

Dactyloctenium 168 

Dactylostegium 42 

Dactylostemon 103 

Dalbergia 105, 108 

Dalechampia 101, 103 

Daltonia 18 

Danaea 35 

Daphnopsis 201 

Datura 197, 198 

Davallia 27 

DAVALLIACEAE 27 

Davalliopsis 32 

Davilla 96 

Davya 135 

Decaraphe 135 

Declieuxia 182, 187 

Degranvillea 155 

Deguelia 108 

Delilia 59 

Delonix 73 

Dendrobangia 116 

Dendroceros 15 

DENDROCEROTACEAE 15 

Dendropanax 53 

Dendrophthora 206 

Dennstaedtia 27 

DENNSTAEDITIACEAE 27 

Derris 108 

Desmanthus 141 

Desmodium 105, 108 

Desmoncus 55, 56 

Desmoscelis 132 

Diacrium 160 

Dialium 73 

Dianthera 42 

Dibrachion 108 

Dichaea 155, 160 

Dichanthium 168 

DICHAPETALACEAE 95 

Dichapetalum 95 

Dichorisandra 86 

Dichromena 94 

Dichrostachys 141 

DICKSONIACEAE 28 

Dicliptera 41 

Dicorynia 73 

DICRANACEAE 17 

Dicranella 17 

Dicranodontium 17 

Dicranoglossum 36 

Dicranolejeunea 12 

Dicranoloma 17 

Dicranopteris 30 

Dicranopygium 90 

Dicranostyles 87 

Dicrypta 161 

Dictyocaryum 55, 56 

Dictyostega 70 

Dicymbe 73 

Dicypellium 119 

Didymiandrum 92 

Didymochlaena 29 

Didymochlamys 182 

Didymopanax 54 

Dieffenbachia 52, 53 

Digitaria 168, 172 

Digomphia 63 

Dilkea 164 

DILLENIACEAE 95-96 

Dimerandra 156, 161 

Dimerocostus 88 

Dimorphandra 73, 76 

Dinizia 73 

Dioclea 105, 108 

Diodia 182, 187 

Diolena 135 

Diomma 190 

Dioscorea 96 

DIOSCOREACEAE 96 

Diospyros 97 

215 

INDEX

216 

Diothonea 161 

DIPHYSCIACEAE 17 

Diphyscium 17 

Diplacrum 92, 94 

Dipladenia 50 

Diplasia 92 

Diplasiolejeunea 12 

Diplazium 40 

Diplochita 135 

Diplopterys 127, 128 

Diplotropis 106, 108 

Diploön 192 

Dipteracanthus 42 

DIPTEROCARPACEAE 96-97 

Dipteryx 106, 108 

Disciphania 140 

Discocarpus 101, 103 

Discophora 116 

Disteganthus 67, 69 

Disterigma 98 

Distictella 63, 64 

Distictis 63 

Ditassa 58 

Dodecastigma 101, 103 

Dodonaea 190 

Dolichopterys 128 

Dolichos 108 

Doliocarpus 96 

Donnellia 21 

Dorstenia 145 

Doryopteris 25 

Dracontium 52, 53 

Drejera 42 

Drepanocarpus 108 

Drepanolejeunea 12 

Drimys 208 

Drosera 97 

DROSERACEAE 97 

Drupina 196 

Drymaria 79 

Drymonia 111, 112 

DRYOPTERIDACEAE 28-30 

Dryopteris 29 

Drypetes 101, 103 

Duckeela 156 

Dugandiodendron 126 

Duguetia 46, 47 

Dulacia 153 

Dumortiera 13 

Duroia 182 

Dussia 106, 108 

Duvernoia 42 

Dyspemptemorion 42 

Dystovomita 84 

E 

EBENACEAE 97 

INDEX 


Ebermaiera 42 

Ecastophyllum 108 

Ecclinusa 192, 193 

Echeandia 124 

Echinochloa 

Echinocolea 12 

Echinodorus 43 

Echinolaena 168 

Echites 50 

Eclipta 59, 61 

Ectropothecium 19 

Egletes 59 

Eichhornia 178 

Elachyptera 114 

Elaeagia 182 

Elaeis 55 

ELAEOCARPACEAE 97-98 

Elaeodendron 80 

Elaeoluma 192 

Elaphoglossum 34 

Elaterium 89 

Eleocharis 92, 94 

Elephantomene 140 

Elephantopus 59, 61 

Eleusine 168 

Eleutheranthera 59 

Eleutherine 117 

Elisabetha 76 

Elizabetha 73 

Elleanthus 156, 161 

Elodea 116 

Eloyella 156 

Eltroplectris 156 

Elvasia 152, 153 

Elyonurus 168, 172 

Emilia 59, 61 

Emmeorrhiza 182 

Emmotum 116 

Encyclia 156, 161 

Endlicheria 119, 120 

Endopleura 115 

Englerella 193 

Enourea 191 

Entada 141, 143 

Enterolobium 141, 143 

Enterosora 30 

Entodontopsis 22 

Entosthodon 18 

Eperua 74, 76 

Ephedranthus 46 

EPHEMERACEAE 17 

Epidendropsis 161 

Epidendrum 156, 161 

Epidryos 180 

Epigaea 99 

Epiphyllum 72 

Episcia 111, 112 

Epistephium 156 

Eragrostis 169, 172 

Eranthemum 41 

Erechtites 59, 61 

EREMOLEPIDACEAE 98 

Eremoluma 193 

Erianthus 172 

ERICACEAE 98-99 

Erigeron 59, 61 

ERIOCAULACEAE 99-100 

Eriocaulon 99 

Eriochloa 169, 172 

Eriochrysis 169 

Eriope 118 

Eriopsis 156, 161 

Eriosema 106, 108 

Eriosorus 25 

Eriotheca 65 

Erisma 207 

Ernestia 132 

Ertela 189 

Ervatamia 50 

Eryngium 47 

Erythrina 106, 108 

Erythrochiton 189 

Erythrodes 156, 161 

ERYTHROXYLACEAE 100 

Erythroxylum 100 

Eschatogramme 36 

Eschweilera 121 

Esenbeckia 189 

Etaballia 106, 108 

Eucamptodontopsis 17 

Euceraea 109 

Euchlaena 172 

Eugenia 148, 150 

Eulacophyllum 22 

Eulophia 156 

Eupatorium 61 

Euphorbia 101, 103 

EUPHORBIACEAE 100-104 

Euphronia 207, 208 

Euphroniaceae 

Euplassa 178, 179 

Euterpe 55, 56 

Evea 187 

Everardia 92 

Evodianthus 90 

Evolvulus 87 

Exacum 110 

Exellodendron 81 

Exochogyne 92 

Exostyles 74 

F 

FABACEAE 104-109 

Fagara 190 

Faramea 182, 187 

Feea 32 

Ferdinandusa 183 

Ferolia 82, 145 

Fevillea 89

Ficus 145 

Fimbristylis 92, 94 

Fischeria 58 

Fissicalyx 106 

Fissidens 17-18 

FISSIDENTACEAE 17-18 

Fittonia 41 

FLACOURTIACEAE 109-110 

Fleischmannia 59 

Flemingia 106 

Fleurya 204 

Floribundaria 20 

Floscopa 86 

Forsteronia 48 50 

Fossombronia 10 

FOSSOMBRONIACEAE 10 

Fothergilla 135 

Fractiunguis 161 

Franchetella 193 

Freziera 200 

Froesiochloa 169 

Froesiodendron 47 

Frullania 10 

FRULLANIACEAE 10 

Frullanoides 12, 13 

Fuirena 92 

Funaria 18 

FUNARIACEAE 18 

Funastrum 58 

Furarium 126 

Furcraea 43 

Fusaea 46 

G 

Gaedawakka 103 

Gaiadendron 125 

Galactia 106 

Galactophora 48 

Galeandra 156, 161 

Galega 108 

Galeottia 156 

Galipea 189, 190 

Galium 183 

Gamochaeta 59, 61 

Gamotopea 187 

Garcinia 85 

Garuga 45 

Gaultheria 98, 99 

Gavarretia 101 

Gaya 129 

Geissopappus 61 

Geissospermum 48 

Genipa 183, 187 

Genlisea 123 

GENTIANACEAE 110-111 

GEOCALYCACEAE 10-11 

Geoffroea 108 

Geonoma 55, 56 

Geophila 183, 187 


Gerardia 196 

Gesneria 111, 112 

GESNERIACEAE 111-112 

Ghinia 205 

Gibasis 86 

Gleichenia 30 

GLEICHENIACEAE 30 

Glinus 144 

Gliricidia 106 

Glochidion 103 

Gloeospermum 206 

Glossadelphus 21 

Gloxinia 111, 112 

Glycine 108 

Glycydendron 101 

Gnaphalium 59, 61 

GNETACEAE 112 

Gnetum 112 

Godmania 63 

Gomphia 153 

Gomphichis 156 

Gomphrena 44 

Gongora 156, 161 

Gongylolepis 59, 61 

Goniopteris 39 

Gonolobus 58 

Gonzalagunia 183 

Goodallia 201 

Goodyera 161 

Gordonia 200 

Gossypium 129 

Gouania 180 

Goupia 80 

Govenia 157 

Graffenrieda 132, 135 

Grammadenia 148 

GRAMMITIDACEAE 30-31 

Grammitis 30 

Granatum 138 

Gravisia 69 

Greeneina 146 

Grias 122 

Groutiella 20 

Guadua 169, 172 

Guaiacum 210 

Guapira 151 

Guarea 138 

Guatteria 46, 47 

Guatteriopsis 46 

Guayania 59 

Guazuma 199 

Guenetia 202 

Guettarda 183, 187 

Guilielma 56 

Gurania 89 

Gustavia 121, 122 

Guzmania 67, 69 

Gymnanthes 103 

Gymnopogon 169 

Gymnopteris 25 

Gymnosiphon 70 

Gynandropsis 78 

Gynerium 169 

Gynocraterium 42 

H 

Habenaria 157, 161 

Hackelochloa 169 

Haemadictyon 50 

Haematostemon 101 

Haematoxylum 74 

HAEMODORACEAE 112 

Haesselia 10 

HALORAGACEAE 112-113 

Hamelia 183, 187 

Handroanthus 64 

Haploclathra 84 

Haplolejeunea 12 

Harpalejeunea 12 

Hartigia 135 

Hasseltia 109 

Havetiopsis 84, 85 

Hebeclinium 59 

Hebepetalum 115 

Hecatostemon 109 

Hecistopteris 40 

HEDWIGIACEAE 18 

Hedychium 209 

Hedyosmum 80 

Hedysarum 108 

Heisteria 153 

Heliamphora 195 

Heliconia 113 

HELICONIACEAE 113 

HELICOPHYLLACEAE 18 

Helicophyllum 18 

Helicostylis 145, 146 

Helicteres 199 

Heliotropium 66 

Helmontia 89 

Helopus 172 

Helosis 62 

Hemarthria 169 

Hemicarpha 94 

Hemicrepidospermum 71 

Hemidictyum 40 

Hemidiodia 187 

Hemigraphis 42 

Hemionitis 25 

Hemiragis 18 

Hemitelia 27 

Henicodium 19 

Henriettea 132, 135 

Henriettella 132, 135 

Henriquezia 183 

HEPATOPHYTA 9-15 

HERBERTACEAE 11 

Herbertus 11 

Herissantia 129 

217 

INDEX

218 

Hernandia 113 

HERNANDIACEAE 113 

Herpestis 196 

Herpetacanthus 42 

Herrania 199 

Heteranthera 178 

Heterocondylus 59 

Heteropogon 169 

Heteropsis 52, 53 

Heteropterys 127, 128 

Heteroscyphus 11 

Heterostemon 74 

Heterothalamus 61 

Hevea 101, 103 

Hexadesmia 161 

Hexapterella 70 

Hexisea 157, 161 

Heymassoli 153 

Hibiscus 129 

Hildebrandtiella 21 

Hillia 183, 187 

Himatanthus 48 

Hippeastrum 124 

Hippobroma 77 

Hippocratea 114 

HIPPOCRATEACEAE 113-115 

Hiraea 127, 128 

Hirtella 81, 82 

Histiopteris 27 

Hoffmannia 183 

Hohenbergia 68 

Holcus 172 

Holomitriopsis 19 

Holomitrium 17 

Homalium 109 

Homalocenchrus 172 

Homalomena 52 

Homolepis 169 

HOOKERIACEAE 18-19 

Hookeriopsis 18 

Hornschuchia 47 

Hortia 189 

Hortsmania 50 

Houlletia 157, 161 

Houmiri 115 

Huberodendron 65 

HUGONIACEAE 115 

Humboldtia 161 

Humiria 115 

HUMIRIACEAE 115 

Humirianthera 116 

Humiriastrum 115 

Humirium 115 

Huntleya 157, 161 

Huperzia 23 

Hura 101 

Hybanthus 206 

HYDROCHARITACEAE 115-116 

Hydrochorea 141 

Hydrocleys 124 

INDEX 


Hydrocotyle 47 

Hydrolea 116 

Hydromystria 116 

HYDROPHYLLACEAE 116 

Hydropogon 19 

HYDROPOGONACEAE 19 

Hydropogonella 19 

Hyeronima 101, 103 

Hygrophila 42 

Hylaeanthe 130 

Hylenaea 114 

Hylocereus 72 

Hymenachne 169, 172 

Hymenaea 74, 76 

Hymenocallis 124 

Hymenolobium 106 

HYMENOPHYLLACEAE 32-33 

HYMENOPHYLLOPSIDACEAE 

33-34 

Hymenophyllopsis 33-34 

Hymenophyllum 32 

Hymenostachys 33 

Hyophila 20 

Hyospathe 55 

Hyparrhenia 169 

Hypenia 118 

Hyperbaena 140 

Hypericum 84, 85 

HYPNACEAE 19 

Hypnella 18 

Hypnum 19 

Hypocyrta 112 

Hypogynium 172 

Hypolepis 27 

Hypolytrum 92 

Hypoxis 124 

Hyptidendron 118 

Hyptis 118 

I 

Ibetralia 187 

ICACINACEAE 116 

Icacorea 148 

Ichnanthus 169, 172 

Ichthyophora 193 

Ichthyothere 59 

Icica 71 

Ilex 51 

Ilysanthes 196 

Impatiens 62 

Imperata 169, 172 

Indigofera 106, 108 

Inga 141, 143 

Ionidium 206 

Ionopsis 157, 161 

Ipomoea 87, 88 

Iresine 44 

Iriartea 

Iriartella 55, 56 

IRIDACEAE 116-117 

Irlbachia 110 

Iroucana 109 

Iryanthera 147 

Isachne 169 

Ischaemum 169, 172 

Ischnosiphon 130 

Iseia 87 

Isertia 183, 187 

Isidrogalvia 124 

Isochilus 157, 161 

Isodrepanium 20 

ISOETACEAE 23 

Isoetes 23 

Isopterygium 19 

Isotachis 10 

Isotoma 77 

Ithycaulon 28 

Ivira 199 

Ixia 117 

Ixocactus 125 

IXONANTHACEAE 117 

Ixophorus 169 

Ixora 183, 187 

J 

Jablonskia 101 

Jacaranda 63, 64 

Jacaratia 78 

Jacquemontia 87, 88 

Jacqueshuberia 74 

Jacquiniella 157 

Jaegerina 21 

Jamesoniella 11 

Japarandiba 122 

Jasarum 52 

Jasminum 153 

Jatropha 101, 103 

Jenmaniella 175 

Jessenia 55, 56 

Jubelina 127 

Jucunda 135 

Jugastrum 122 

JUGLANDACEAE 117 

Juglans 117 

JUNGERMANNIACEAE 11 

Juruasia 43 

Jussiaea 154 

Justicia 42 

K 

Kaieteurea 153 

Kalanchoe 88 

Kallstroemia 210 

Kedrostis 89 

Kefersteinia 157, 162 

Kegelia 162 

Kegeliella 157

Klugiodendron 143 

Koanophyllon 59 

Koellensteinia 157 

Koernickanthe 130 

Kohleria 111 

Kopsia 48 

Kotchubaea 183 

Krameria 117 

KRAMERIACEAE 117 

Krugia 150 

Kubitzkia 119 

Kurzia 13 

Kyllinga 92, 94 

L 

Labatia 193 

Lablab 106 

Lacistema 117 

LACISTEMATACEAE 117 

Lacmellea 48 

Lacostea 33 

Lacunaria 179 

Ladenbergia 183 

Laelia 162 

Laetia 109 

Lafoensia 126 

Lagascea 59 

Lagenaria 89 

Lagenocarpus 92, 94 

Lagerstroemia 126 

Laguncularia 85 

LAMIACEAE 118 

Lamprococcus 69 

Landolphia 50 

Lanessania 146 

Langsdorffia 62 

Lanium 162 

Lantana 205 

Laplacea 201 

Laportea 204 

Lasiacis 169, 172 

Lasiadenia 201 

Lasiandra 135 

Lastrea 39 

Lastreopsis 38 

LAURACEAE 118-120 

Laurea 146 

Laurembergia 112 

Lawsonia 126 

Laxoplumeria 48 

Leandra 132, 135 

Lecointea 74 

Lecostemon 180 

LECYTHIDACEAE 120-122 

Lecythis 121, 122 

Ledothamnus 98 

Leersia 169 

Leiomela 16 

Leiomitrium 20 


Leiostegia 136 

Leiothamnus 111 

Leiothrix 99 

Leiphamos 111 

Leitgebia 153 

Lejeunea 12 

LEJEUNEACEAE 11-13 

Lellingeria 31 

Lembocarpus 111 

Lemna 122 

LEMNACEAE 122 

LENTIBULARIACEAE 123 

Leonia 206 

Leonotis 118 

Leonurus 118 

Lepanthes 157 

Lepanthopsis 157 

Lepianthes 166 

Lepidagathis 42 

Lepidaploa 59 

Lepidium 66, 67 

Lepidobotryaceae 

Lepidocaryum 55, 56 

Lepidocordia 66 

Lepidolejeunea 12 

Lepidopilidium 18 

Lepidopilum 18 

Lepidozia 13 

LEPIDOZIACEAE 13 

Leptochilus 34 

Leptochloa 169, 172 

Leptocoryphium 169 

Leptolejeunea 12 

Leptolobium 108 

Leptoscyphus 11 

Leptostachys 172 

Lepyrodontopsis 16 

Leretia 116 

Leskeodon 18 

Leucaena 141, 143 

LEUCOBRYACEAE 19 

Leucobryum 19 

LEUCODONTACEAE 19 

Leucodontopsis 19 

Leucohyle 157 

Leucolejeunea 12 

Leucoloma 17 

LEUCOMIACEAE 19-20 

Leucomium 19-20 

Leucophanes 19 

Licania 81, 82 

Licaria 119, 120 

Ligea 176 

Ligeophila 157 

Lightia 208 

LILIACEAE 123-124 

Limnobium 116 

Limnocharis 124 

LIMNOCHARITACEAE 124 

Limnosipanea 183 

Limodorum 162 

Lindackeria 109 

Lindernia 195 

Lindleyella 162 

Lindmania 69 

Lindsaea 27-28 

Linociera 153 

Liparis 157, 162 

Lipocarpha 92, 94 

Lippia 205 

Liriosma 153 

Lisianthus 111 

Lissocarpa 124 

LISSOCARPACEAE 124 

Lithachne 169 

LOASACEAE 124 

Lobelia 77 

Lochnera 50 

Lockhartia 157, 162 

LOGANIACEAE 124-125 

Lomagramma 34 

LOMARIOPSIDACEAE 34 

Lomariopsis 34 

Lonchitis 28 

Lonchocarpus 106, 108 

Lophocolea 11 

Lophogyne 176 

Lopholejeunea 12, 13 

Lophopterys 127, 128 

Lophotocarpus 44 

LORANTHACEAE 125-126 

Loranthus 126 

Loreya 133, 136 

Lorostemon 84 

Loxopterygium 45 

Lucuma 193 

Ludovia 90 

Ludwigia 154 

Luehea 201, 202 

Lueheopsis 202 

Luetzelburgia 106 

Luffa 89 

Lundia 63 

Luziola 169, 172 

Lycaste 157 

Lycianthes 197, 198 

Lycomormium 162 

LYCOPHYTA 22-24 

LYCOPODIACEAE 23 

Lycopodiella 23 

Lycopodium 23 

LYCOPODIACEAE 23 

LYGODIACEAE 34 

Lygodium 34 

Lyroglossa 157 

Lysiloma 141 

Lysiostyles 87 

LYTHRACEAE 126 

M 

219 

INDEX

220 

Mabea 101, 103 

Maburea 153 

Macahanea 210 

Macairea 133, 136 

Macfadyena 63 

Machaerium 106, 108 

Machaonia 183 

Maclura 145 

Macoubea 48, 50 

Macoucoua 51 

Macradenia 157, 162 

Macrocarpaea 110, 111 

Macrocentrum 133, 136 

Macroclinium 157 

Macroditassa 58 

Macrolobium 74, 76 

Macromitrium 20 

Macropharynx 48 

Macroptilium 106 

Macrosamanea 141, 143 

Macroscepis 58 

Magnolia 127 

MAGNOLIACEAE 126-127 

Maguirea 53 

Maguireanthus 133 

Maguireella 21 

Maguireothamnus 183 

Mahurea 84 

Maieta 133, 136 

Malachra 129 

Malanea 183, 187 

Malaxis 157 

Malmea 46, 47 

Malouetia 48, 50 

Malpighia 128 

MALPIGHIACEAE 127-128 

MALVACEAE 128-130 

Malvastrum 129 

Manabea 205 

Managa 210 

Mandevilla 48, 50 

Manettia 183 

Manicaria 55 

Manihot 102, 103 

Manilkara 192, 193 

Manisuris 172 

Mansoa 63 

Mapania 92 

Mapaniopsis 92 

Mapouria 187 

Mappia 116 

Maprounea 102 

Maquira 145 

Maranta 130 

MARANTACEAE 130-131 

Marathrum 175 

MARATTIACEAE 35 

Marcetia 133 

Marcgravia 131 

INDEX 


MARCGRAVIACEAE 131 

Marchantia 13 

Marchantiaceae 13 

Marchesinia 12 

Margaritaria 102 

Marila 84 

Maripa 87, 88 

Mariscus 94 

Markea 197, 198 

Markleya 56 

Marlierea 149, 150 

Marmaroxylon 143 

Marsdenia 58 

Marsilea 35 

MARSILEACEAE 35 

Marsupidium 9 

Marsypianthes 118 

Martinella 63, 64 

Martiodendron 74 

Martiusia 76 

Mascagnia 127 

Masdevallia 157, 162 

Mastigolejeunea 12 

Matayba 190, 191 

Matelea 58 

Matisia 65 

Matourea 196 

Mauria 45 

Mauritia 55, 56 

Mauritiella 55, 56 

Maxillaria 157, 162 

Maximbignya 55 

Maximiliana 55, 56 

Mayaca 131 

MAYACACEAE 131 

Mayepea 153 

Mayna 109, 110 

Maytenus 80 

Meborea 103 

Mecardonia 195 

Megaskepasma 42 

Meioluma 194 

Meiothecium 21 

Meisneria 136 

Melaleuca 149 

Melampodium 61 

Melanoxylon 76 

Melanthera 59, 61 

Melasma 196 

Melastoma 136 

MELASTOMATACEAE 131-138 

Melia 138 

MELIACEAE 138-139 

Meliandra 136 

Melicocca 

Melicoccus 191 

Melinis 169 

Mella 196 

Melloa 63 

Melocactus 72 

Melochia 199 

Melothria 89 

Melpomene 31 

Memora 63, 64 

Menadenium 162 

Mendoncella 162 

Mendoncia 139 

MENDONCIACEAE 139 

Meniscium 39 

MENISPERMACEAE 139-140 

Mentzelia 124 

MENYANTHACEAE 140 

Meriania 133 

Merostachys 169 

Merremia 87, 88 

Merumea 183 

Mesadenella 158, 162 

Mesechites 48, 50 

Mesosetum 169, 172 

Metalepis 58 

Metastelma 58 

Metaxya 35 

METAXYACEAE 35 

METEORIACEAE 20 

Meteoridium 20 

Meteoriopsis 20 

Metrodorea 189 

Metzgeria 13 

METZGERIACEAE 13-14 

Mezia 128 

Mezilaurus 119, 120 

Mezobromelia 69 

Miconia 133, 136 

Micrandra 102, 103 

Micranthemum 195, 196 

Microchloa 169 

Microdus 17 

Microgonium 33 

Microgramma 36 

Microlejeunea 12 

Microlepia 28 

Microlicia 134, 137 

Micromitrium 17 

Micropholis 192, 194 

Micropolypodium 31 

Micropterygium 13 

Microstachys 103 

Microtea 165 

Miersiella 70 

Mikania 60, 61 

Mimosa 141, 143 

MIMOSACEAE 140-144 

Mimusops 192, 194 

Minquartia 153 

Mirabilis 151 

Misanteca 120 

Mitostemma 164 

Mitracarpus 183, 188 

Mitreola 125 

Mitrocarpum 188

Mittenothamnium 19 

Mnasium 180 

MNIACEAE 20 

Mniomalia 20 

Mollia 202 

Mollinedia 144 

MOLLUGINACEAE 144 

Mollugo 144 

Momordica 89 

Monachanthus 162 

MONIMIACEAE 144 

Monnieria 190 

Monnina 176, 177 

Monochaeum 134 

Monoclea 14 

MONOCLEACEAE 14 

Monogramma 31 

Monophyllanthe 130 

Monopteryx 106 

Monotagma 130, 131 

Monotrema 180 

Monstera 52, 53 

Montira 210 

Montrichardia 52 

Moquilea 83 

Mora 74 

MORACEAE 144-146 

Morinda 183 

Moringa 146 

MORINGACEAE 146 

Morisonia 78 

Mormodes 158 

Moronobea 84 

Moschoxylum 139 

Mostuea 125 

Mourera 175 

Mouriri 134, 137 

Mouroucoa 88 

Moutabea 176 

Moutouchi 108 

Mucuna 106, 108 

Muellera 106, 108 

Murdannia 86 

Murraya 189, 190 

Musa 146 

MUSACEAE 146 

Mussatia 63 

Myanthus 162 

MYOPORACEAE 146 

Myoxanthus 158 

Myrcia 149, 150 

Myrcianthes 149 

Myrciaria 149, 150 

Myriaspora 134, 137 

Myriocladus 169 

Myriophyllum 112, 113 

Myristica 147 

MYRITICACEAE 146-147 

Myrmidone 134 

Myrosma 130, 131 


Myroxylon 106 

MYRSINACEAE 147-148 

Myrsine 147, 148 

MYRTACEAE 148-150 

Myrtiluma 194 

Myrtus 150 

Mytilopsis 13 

N 

Nacibea 188 

NAJADACEAE 150 

Najas 150 

Naletonia 188 

Nanodes 162 

Napeanthus 111 

Napimoga 110 

Nasturtium 67 

Naucleopsis 145, 146 

Nautilocalyx 111, 112 

Navia 68, 69 

Neblinaria 66 

NECKERACEAE 20 

Neckeropsis 20 

Nectandra 119, 120 

Neea 151 

Neesioscyphus 10 

Nelsonia 42 

Nelumbo 150 

NELUMBONACEAE 150 

Nemaluma 194 

Nematanthus 112 

Neobartlettia 162 

Neobertiera 183 

Neogleasonia 66 

Neolacis 176 

Neolehmannia 162 

Neomarica 117 

Neopometia 194 

Neoptychocarpus 109 

Neoraputia 189 

Neosprucea 109 

Neoxythece 194 

Nephradenia 58 

Nephrodium 39 

Nephrolepis 35 

Nepsera 134 

Neptunia 142, 143 

Nerium 48 

Neurocarpum 108 

Neurodium 36 

Neurolaena 60 

Neurolejeunea 12 

Neurolepis 169 

Neuromanes 33 

Neurophyllum 33 

Neurotheca 110 

Newtonia 143 

Nicotiana 198 

Nidema 158, 162 

Nietneria 124 

Niphidium 36 

Nissolia 108 

Nitella 9 

Noisettia 206 

Nonatelia 188 

Nopalea 72 

Norantea 131 

Noterophila 137 

Notholaena 25 

Notopleura 188 

Notopora 98 

NOTOTHYLADACEAE 15 

Notothylas 15 

Notylia 158, 162 

Nowellia 10 

Noyera 146 

NYCTAGINACEAE 150-151 

Nymphaea 151 

NYMPHAEACEAE 151 

Nymphoides 140 

O 

OCHNACEAE 151-153 

Ochroma 65 

Ochthephilus 134 

Ochthocosmus 117 

Ocimum 118 

Ocotea 119, 120 

Octoblepharum 19 

Octomeria 158, 162 

Odontadenia 48, 50 

Odontocarya 140 

Odontoglossum 158 

Odontolejeunea 12 

Odontoloma 28 

Odontomanes 33 

Odontonema 42 

Odontoschisma 10 

Odontosoria 28 

Oeceoclades 158 

Oedematopus 84, 85 

Oenocarpus 55, 56 

Oenone 176 

Ogcodeia 146 

OLACACEAE 153 

Olax 153 

Oldenlandia 183, 188 

Oldenlandiopsis 183 

OLEACEAE 153 

Oleandra 35 

OLEANDRACEAE 35 

Olfersia 29 

Oliganthes 61 

Oliveriana 158 

Olmedia 146 

Olmedioperebea 146 

Olyra 169, 172 

Omphalanthus 12 

221 

INDEX

222 

Omphalea 102 

Omphalobium 87 

ONAGRACEAE 153-154 

Oncidium 158, 162 

Onoctonia 137 

Oocarpon 154 

Operculina 88 

Ophiocaryon 190 

OPHIOGLOSSACEAE 35-36 

Ophioglossum 35 

Ophrys 162 

OPILIACEAE 154 

Oplismenus 169, 173 

Opuntia 72 

Orbignya 55, 56 

ORCHIDACEAE 154-163 

Orectanthe 208 

Orelia 50 

Oreopanax 53, 54 

Orleansia 158 

Ormoloma 28 

Ormosia 106, 108 

Ormosiopsis 108 

Ornithocephalus 158, 162 

Orthaea 98, 99 

Orthoclada 169, 173 

Orthomene 140 

Orthopappus 60 

Orthostichidium 21 

Orthostichopsis 21 

ORTHOTRICHACEAE 20 

Oryctanthus 125, 126 

Oryctina 125 

Oryza 169, 173 

Osbeckia 137 

Oserya 175 

Ossaea 134, 137 

Osteophloeum 147 

Otachyrium 169 

Otostylis 158, 162 

Ouratea 152, 153 

Ourouparia 188 

Outea 76 

OXALIDACEAE 163 

Oxalis 163 

Oxandra 46, 47 

Oxycaryum 92 

Oxymeris 137 

Oxypetalum 58 

Oxythece 194 

Oyedaea 60, 61 

P 

Pachira 65 

Pachyloma 134 

Pachyptera 64 

Pachystachys 42 

Pacouria 49 

Pacourina 60 

INDEX 


Paepalanthus 99 

Pagamea 183 

Pakaraimaea 97 

Palicourea 184, 188 

Pallavicinia 14 

PALLAVICINIACEAE 14 

Palmorchis 158 

Paloue 74 

Paloveopsis 74 

Paltonium 37 

Pamea 86 

Panax 54 

Pancratium 124 

Panicum 169, 173 

Panopsis 178, 179 

PAPAVERACEAE 163 

Papaya 78 

Paphinia 158 

Paphiopedilum 162 

Papillaria 20 

Pappophorum 170 

Parabignonia 63 

Paracromastigum 13 

Paradrymonia 111 

Paragonia 63 

Parahancornia 49, 50 

Paralabatia 194 

Paralea 97 

Paraleucobryum 19 

Paramachaerium 106 

Paramicropholis 194 

Paratheria 170 

Pariana 170, 174 

Parinari 82, 83 

Parinarium 83 

Parivoa 76 

PARKERIACEAE 36 

Parkia 142, 143 

Parkinsonia 74 

Parodiolyra 170 

Parthenium 60 

Paspalidium 170 

Paspalum 170, 174 

Passiflora 164 

PASSIFLORACEAE 164 

Passoura 206 

Patabea 184, 188 

Patima 184 

Patrisia 110 

Paullinia 191 

Pausandra 102, 103 

Pavonia 129 

Paypayrola 206 

Pecluma 36 

Pectis 60, 61 

PEDALIACEAE 164-165 

Pedilanthus 102 

Pekea 78 

Pelexia 158, 162 

Peltaea 129 

Peltogyne 74, 76 

Peltophorum 74 

Peltophyllum 202 

Pennisetum 170, 174 

Pentacalia 60 

Pentaclethra 142, 143 

Pentascyphus 191 

Peperomia 165, 166 

Pepinia 69 

Pera 102, 103 

Perama 184 

Perebea 145, 146 

Pereskia 72 

PERIDISCACEAE 165 

Peridium 103 

Peristeria 158, 162 

Peritassa 114 

Pernettya 98 

Persea 119, 120 

Persicaria 177 

Peschiera 50 

Petagomoa 188 

Petaloma 137 

Petastoma 64 

Petiveria 165 

Petrea 205 

Petrocarya 83 

Pfaffia 44 

Phainantha 134 

Pharmacosycea 146 

Pharus 170, 174 

Phaseolus 107, 108 

Phegopteris 39 

Phenakospermum 199 

Phenax 204 

Philodendron 52, 53 

Philodice 99 

Philonotis 16 

Philotria 116 

Philoxerus 44 

Phlebodium 36 

Phoebe 120 

Phoenix 55 

Phoradendron 206-207 

Phragmipedium 158 

Phragmites 170, 174 

Phryganocydia 64 

Phrynium 131 

Phthirusa 125, 126 

Phyla 205 

Phyllanthus 102, 103 

Phyllocactus 72 

Phyllodon 21 

PHYLLODREPANIACEAE 20 

Phyllodrepanium 20 

PHYLLOGONIACEAE 20 

Phyllogonium 20 

Physalis 198 

Physostemon 78 

Phytolacca 165

PHYTOLACCACEAE 165 

Picramnia 196 

Picrasma 196 

Picria 196 

Picrolemma 196 

Pictolejeunea 12 

Pilea 204 

Pilocarpus 189 190 

Pilosium 22 

Pilosocereus 72 

Pilotrichella 20 

Pilotrichum 19 

Pilumna 162 

PINACEAE 165 

Pinus 165 

Pinzona 96 

Piparea 110 

Piper 166, 167 

PIPERACEAE 165-167 

Piptadenia 142, 143 

Piptocarpha 60 

Piptocoma 60 

Piranhea 102 

Piratinera 146 

Pireella 21 

Piresia 170 

Pirigara 122 

Piripea 196 

Piriqueta 203 

Pisonia 151 

Pistia 52 

Pitcairnia 68, 69 

Pithecellobium 142, 143 

Pithecoctenium 64 

Pitumba 110 

Pityrogramma 25 

Plagiochila 14 

PLAGIOCHILACEAE 14 

Plagiomnium 20 

Plagiothecium 22 

Planchonella 194 

PLANTAGINACEAE 167 

Plantago 167 

Plathymenia 142 

Platonia 84 

Platyaechmea 69 

Platycarpum 184 

Platycentrum 137 

Platyluma 194 

Platymiscium 107 

Platystele 158 

Platythelys 158 

Plectrophora 158 

Pleonotoma 64 

Pleopeltis 36 

Pleroma 137 

Pleurisanthes 116 

Pleurostachys 93 

Pleurothallis 158, 162 

Pleurothyrium 119, 120 


Pleurozia 14 

PLEUROZIACEAE 14 

Plinia 149, 150 

Pluchea 60, 61 

Plukenetia 102, 103 

PLUMBAGINACEAE 167 

Plumbago 167 

Plumeria 49, 50 

POACEAE 167-175 

Pochota 65 

PODOCARPACEAE 175 

Podocarpus 175 

Podoluma 194 

PODOSTEMACEAE 175-176 

Podostemum 175 

Poecilandra 152 

Poecilanthe 107 

Pogonia 162 

Pogonophora 102 

Pogopetalum 116 

Poinciana 76 

Pollalesta 61 

Polybotrya 29 

Polycarpaea 79 

Polycycnis 159 

Polygala 176, 177 

POLYGALACEAE 176-177 

POLYGONACEAE 177-178 

Polygonum 177 

Polylychnis 42 

POLYPODIACEAE 36-37 

Polypodium 36 

Polypompholyx 123 

Polypremnum 125 

Polystachya 159, 162 

Polystichopsis 29 

Polystichum 29 

Polytaenium 40 

POLYTRICHACEAE 20 

Polytrichum 20 

Ponera 163 

Pontederia 178 

PONTEDERIACEAE 178 

Ponthieva 159 

Poraqueiba 116 

Porocystis 191 

Porophyllum 60, 61 

Porotrichum 20 

Portesia 139 

Portulaca 178 

PORTULACACEAE 178 

Posadaea 89 

Posoqueria 184, 188 

Possira 76 

Potalia 125 

Potamium 21 

Potamoganos 64 

Potarophytum 180 

Poteranthera 137 

Pothomorphe 166 

Pothuava 69 

POTTIACEAE 20-21 

Poulsenia 145 

Pourouma 79, 80 

Pouteria 192-193, 194 

Pradosia 193 

Praxelis 60 

Prescottia 159 

Prestoea 55 

Prestonia 49, 50 

Prieurella 194 

Prionolejeunea 12 

Prionostemma 114 

Pristimera 114 

Priva 205 

Prockia 109 

PROTEACEAE 178-179 

Protium 70, 71 

Prunus 181 

Psadacocalymma 43 

Psammisia 98, 99 

Pseudabutilon 130 

Pseudanamomis 149 

Pseudechinolaena 170 

Pseudelephantopus 60, 61 

Pseuderiopsis 163 

Pseudima 191 

Pseudocalymma 64 

Pseudochimarrhis 188 

Pseudocladia 194 

Pseudoconnarus 86 

Pseudocryphaea 19 

Pseudogynoxys 60 

Pseudolabatia 194 

Pseudolmedia 145, 146 

Pseudopaegma 

Pseudopiptadenia 142 

Pseudoxandra 46, 47 

Pseudoxythece 194 

Psidium 149, 150 

Psiguria 89 

Psilochilus 159 

PSILOTOPHYTA 22 

PSILOTACEAE 22 

Psilotum 22 

Psittacanthus 125 

Psychopsis 159 

Psychotria 184, 188 

Psygmorchis 159 

Pterandra 128 

PTERIDACEAE 37 

Pteridium 28 

Pteris 37 

PTEROBRYACEAE 21 

Pterocarpus 107, 109 

Pterocaulon 60, 61 

Pterogastra 134 

Pterogonidium 21 

Pterolepis 134, 137 

PTEROPHYTA 24-40 

223 

INDEX

224 

Pteropsiella 13 

Pterozonium 25 

Ptilophyllum 33 

Ptychocarpus 110 

Ptychopetalum 153 

Pueraria 107 

Pulchranthus 42 

Pycnolejeunea 13 

Pycreus 94 

Pyrostegia 64 

Pyrrhobryum 21 

Q 

Quadrella 78 

Qualea 207 

Quapoya 84, 85 

Quararibea 65 

Quassia 196 

Quebitea 167 

Quekettia 159 

Quelchia 60 

Quesnelia 69 

Quiina 179 

QUIINACEAE 179 

R 

Racaria 191 

Racinaea 69 

RACOPILACEAE 21 

Racopilum 21 

Racoubea 110 

Raddia 114, 170, 174 

Raddiella 170, 174 

Radlkoferella 194 

Radula 14 

RADULACEAE 14-15 

RAFFLESIACEAE 179 

Ragala 194 

Ragatelus 33 

Raimondia 47 

Randia 185 

RANUNCULACEAE 179 

Rapanea 148 

Rapatea 180 

RAPATEACEAE 179-180 

Raphia 57 

Raputia 189, 190 

Rauia 189, 190 

Rauvolfia 49, 50 

Ravenala 200 

Ravenia 190 

Raveniopsis 189 

Rechsteineria 112 

Recordoxylon 74, 76 

Rectolejeunea 13 

Rehia 170 

Reichenbachanthus 159 

Reimarachloa 174 

INDEX 


Reimaria 174 

Reimarochloa 170 

Relbunium 188 

Remijia 185 

Remirea 93 

Renauldia 21 

Renealmia 209 

Renggeria 84, 85 

Rengifa 85 

Restrepia 163 

Retiniphyllum 185, 188 

Rhabdadenia 49 

RHABDODENDRACEAE 180 

Rhabdodendron 180 

Rhacocarpus 18 

Rhacodiscus 43 

Rhacopilopsis 19 

Rhadinocarpus 109 

RHAMNACEAE 180 

Rhamnus 180 

Rhaphidolejeunea 13 

Rhaphidorrhynchium 19, 21 

Rhaphidostichum 21 

Rheedia 84, 85 

Rhexia 137 

Rhipidocladum 171 

Rhipsalis 72 

RHIZOGONIACEAE 21 

Rhizogonium 21 

Rhizophora 181 

RHIZOPHORACEAE 181 

Rhodobryum 16 

Rhodognaphalopsis 65 

Rhodospatha 52, 53 

Rhodostemonodaphne 119 

Rhoeo 86 

Rhoogeton 111 

Rhynchanthera 134, 137 

Rhynchelytrum 171 

Rhynchocladium 93 

Rhyncholacis 175 

Rhynchosia 107 

Rhynchospora 93, 94 

Rhytachne 171 

Riana 206 

Riccardia 10 

Riccia 15 

RICCIACEAE 15 

Richardella 194 

Richardia 185 

Richeria 102, 103 

Ricinus 102 

Riencourtia 60, 61 

Rinorea 206 

Rinoreocarpus 206 

Rittera 76 

Rivina 165 

Robergia 87 

Robinia 76, 109 

Rodriguezia 159, 163 

Roentgenia 64 

Rogersanthus 111 

Rohria 95 

Rolandra 60, 61 

Rollinia 46, 47 

Ronabea 188 

Rondeletia 188 

Rondonanthus 99 

Ropourea 97 

Roraimanthus 153 

Rorippa 66, 67 

ROSACEAE 181 

Rosenbergiodendron 188 

Rotala 126 

Rottboellia 171, 174 

Roucheria 115 

Rouhamon 125 

Roulinia 58 

Rouliniella 58 

Roupala 178, 179 

Rourea 86, 87 

Roystonea 55, 57 

RUBIACEAE 181-189 

Rubus 181 

Rudgea 185, 188 

Rudolfiella 159 

Ruellia 42 

Ruizterania 208 

Rumohra 27 

Ruppia 189 

RUPPIACEAE 189 

Ruprechtia 177 

Ruptiliocarpon 163 

Russelia 195 

RUTACEAE 189-190 

Ryania 109, 110 

Rytidostylis 89 

S 

SABIACEAE 190 

Sabicea 186, 188 

Saccharum 171, 174 

Sacciolepis 171 

Saccoloma 28 

Sacoglottis 115 

Sacoila 159 

Sagittaria 43 

Sagonea 116 

Sagotia 102, 103 

Sagraea 137 

Salacia 114 

Salmea 60 

Salpichlaena 26 

Salpinga 134, 137 

Salvertia 208 

Salvinia 38 

SALVINIACEAE 38 

Samanea 142, 143 

Sandwithia 102

Sandwithiodoxa 195 

Santalodes 87 

Saouari 79 

SAPINDACEAE 190-191 

Sapindus 191 

Sapium 102, 103 

SAPOTACEAE 191-195 

Saranthe 131 

Sarcaulus 193, 210 

Sarcoglottis 159, 163 

Sarcostemma 58 

Sarmentaria 137 

SARRACENIACEAE 195 

Satyria 98, 99 

Sauvagesia 152, 153 

Saxofridericia 180 

Scapania 15 

SCAPANIACEAE 15 

Scaphosepalum 159, 163 

Scaphyglottis 159, 163 

Scelochilus 159 

Scheelea 55 

Schefflera 53, 54 

Schiekia 112 

Schiffneriolejeunea 13 

Schismatoglottis 52, 53 

Schistostemon 115 

Schizachyrium 171, 174 

Schizaea 38 

SCHIZAEACEAE 38 

Schizoloma 28 

Schizomitrium 19 

Schlechterella 163 

Schlegelia 64 

Schlimia 159 

Schlotheimia 20 

Schoenobiblus 201 

Schoenoplectus 93 

Schoepfia 153 

Schomburgkia 159 

Schradera 186, 188 

Schrankia 144 

Schultesia 110, 111 

Schusterolejeunea 13 

Schwenckia 198 

Sciadocephala 60 

Sciadophyllum 54 

Sciadotenia 140 

Sciaphila 202 

Scirpus 95 

Sciuris 190 

Scleria 93, 95 

Sclerolobium 74 

Scleronema 65 

Scoparia 195 

SCROPHULARIACEAE 195-196 

Scutellaria 118 

Scuticaria 159 

Scyphonichium 191 

Sebastiania 102, 104 


Secondatia 49 

Secretania 153 

Securidaca 176, 177 

Securinega 104 

Seguieria 165 

Selaginella 24 

SELAGINELLACEAE 23-24 

Selenicereus 72 

Selenipedium 159, 163 

Selysia 89 

Semaphyllanthe 186 

SEMATOPHYLLACEAE 21 

Sematophyllum 21 

Senapea 210 

Senefeldera 102, 104 

Senefelderopsis 102 

Senna 74, 76 

Serapias 163 

Serjania 191 

Sesamum 164, 165 

Sesbania 107, 109 

Sesuvium 43 

Setaria 171, 174 

Sickingia 188 

Sida 129, 130 

Sidastrum 129 

Sideroxylon 195 

Sievekingia 159 

Sigesbeckia 60 

Sigmatostalix 159 

Simaba 196 

Simarouba 196 

SIMAROUBACEAE 196 

Simira 186 

Sinapis 67 

Singana 109 

Sinningia 111 

Siolmatra 89 

Sipanea 186, 189 

Siparuna 144 

Siphanthera 134, 137 

Siphantheropsis 137 

Siphoneugena 149, 150 

Sisyrinchium 117 

Sloanea 97 

SMILACACEAE 197 

Smilax 197 

Sobralia 159 

Socratea 56 

Soemmeringia 107 

SOLANACEAE 197-198 

Solandra 198 

Solanum 198 

Solena 189 

Solenidium 159 

Somphoxylon 140 

Sonchus 60, 61 

Soramia 96 

Sorghastrum 171 

Sorghum 171, 174 

225 

Soridium 202 

Sorocea 145, 146 

Souroubea 131 

Spachea 128 

Sparattanthelium 113 

Sparganophorus 61 

Spartina 171, 174 

Spathanthus 180 

Spathelia 190 

Spathiphyllum 52, 53 

SPECIES INCERTAE SEDIS 210 

Specklinia 163 

Spennera 137 

Spermacoce 186, 189 

SPERMATOPHYTA 40-210 

Sphaerocionium 33 

Sphaeropteris 27 

SPHAGNACEAE 22 

Sphagneticola 60 

Sphagnum 22 

Spheneria 171, 174 

Sphenoclea 199 

SPHENOCLEACEAE 199 

Sphinctanthus 186 

Sphinctolobium 109 

Sphyrospermum 98, 99 

Spigelia 125 

Spilanthes 60, 61 

Spiracantha 61 

Spiranthera 190 

Spiranthes 159, 163 

Spiridentopsis 21 

Spirodela 122 

Spirotropis 107, 109 

SPLACHNACEAE 22 

Splachnobryum 22 

Spondias 45 

Sporobolus 171, 174 

Squamidium 20 

Stachytarpheta 205 

Stahelia 111 

Stanhopea 159, 163 

Staphidiastrum 138 

Staphidium 138 

Staurogyne 42 

Stegolepis 180 

Steirachne 171, 174 

Steiropteris 39 

Stelestylis 90 

Stelis 159, 163 

Stemmadenia 49, 50 

Stemodia 195, 196 

Stenia 160 

Stenochlaena 26 

Stenolobium 109 

Stenomeria 58 

Stenopadus 60, 61 

Stenoptera 163 

Stenorrhipis 11 

Stenorrhynchos 163 

INDEX

226 

Stenosolen 50 

Stenospermation 52 

Stenotaphrum 171, 174 

Sterculia 199 

STERCULIACEAE 199 

STEREOPHYLLACEAE 22 

Stereophyllum 22 

Sterigmapetalum 181 

Sticherus 30 

Stictolejeunea 13 

Stifftia 60, 61 

Stigmaphyllon 128 

Stigmatopteris 29 

Stilaginella 104 

Stilpnopappus 61 

Stizophyllum 64 

Stomatochaeta 60, 61 

STRELITZIACEAE 199-200 

Strempelia 189 

Strephium 174 

Streptocalyx 68 

Streptochaeta 171 

Streptogyna 171, 174 

Streptostachys 171, 174 

Stromanthe 130 

Struchium 60 

Struthanthus 126 

Strychnos 125 

Stryphnodendron 142, 144 

Sturmia 163 

Stylogyne 147, 148 

Stylosanthes 107 

STYRACACEAE 200 

Styrax 200 

Suriana 200 

SURIANACEAE 200 

Swartzia 74-75, 76 

Sweetia 109 

Swietenia 138 

Syagrus 56, 57 

Symbiezidium 13 

Symbolanthus 110 111 

Symmeria 177 

Symphonia 84 

Symphyllanthus 95 

Symphyogyna 14 

Symphyotrichum 60 

SYMPLOCACEAE 200 

Symplocos 200 

Synedrella 60 

Syngonanthus 99, 100 

Syngonium 52, 53 

Syntherisma 175 

Syrrhopodon 16 

Systemonodaphne 120 

Syzygiella 11 

Syzygium 149 

T 

INDEX 


Tabebuia 64 

Taberna 50 

Tabernaemontana 49, 50 

Tacca 200 

TACCACEAE 200 

Tachia 110 

Tachibota 83 

Tachigali 75, 76, 109 

Tagetes 60 

Taligalea 205 

Talinum 178 

Talisia 191 

Tamonea 138, 205 

Tampoa 210 

Tanaecium 64 

Tanibouca 86 

Taonabo 201 

Tapeinostemon 110, 111 

Tapirira 45 

Tapirocarpus 210 

Tapogomea 189 

Tapura 95 

Taralea 107 

Tariri 196 

Tassadia 58 

Taxilejeunea 13 

Taxithelium 21 

Tecoma 64 

Tectaria 38 

TECTARIACEAE 38-39 

Telanthera 44 

Telaranea 13 

Teliostachya 42 

Telitoxicum 140 

Telminostelma 58 

Tephrosia 107, 109 

Teramnus 107 

Terminalia 85, 86 

Ternstroemia 200, 201 

Terpsichore 31 

Tetracera 96 

Tetragastris 71 

Tetralocularia 88 

Tetrameris 138 

Tetrapodenia 128 

Tetrapollinia 111 

Tetrapterys 128 

Tetrastictella 65 

Tetrazygia 138 

Teuscheria 160 

Thalia 130, 131 

Thamniopsis 18 

THEACEAE 200-201 

Thecophyllum 69 

THELYPTERIDACEAE 39 

Thelypteris 39 

Theobroma 199 

THEOPHRASTACEAE 201 

Thermophila 114 

Thespesia 129 

Thevetia 49, 50 

Thibaudia 98, 99 

Thieleodoxa 186 

Thiloa 86 

Thinouia 191 

Thismia 70 

Thoa 112 

Thoracocarpus 90 

Thrasya 171, 175 

THUIDIACEAE 22 

Thuidium 22 

Thunbergia 42 

Thurnia 201 

THURNIACEAE 201 

THYMELAEACEAE 201 

Thyrsodium 45 

Thysananthus 13 

Thysanostemon 84 

Tibouchina 134, 138 

Ticorea 190 

Tigarea 96 

TILIACEAE 201-202 

Tillandsia 68, 69 

Tinantia 86 

Tococa 134, 138 

Tocoyena 186 

Tofieldia 124 

Tonina 99 

Tonsella 115 

Tontanea 189 

Tontelea 114, 115 

Topobea 134, 138 

Torenia 195 

Torrubia 151 

Torulinium 95 

Touantea 76 

Touchiroa 76 

Toulicia 191 

Tounatea 76 

Tournefortia 66 

Touroulia 179 

Tovomita 84 

Tovomitidium 84 

Tovomitopsis 84 

Trabacellula 10 

Trachylejeunea 13 

Trachypogon 171, 175 

Trachyxiphium 18 

Tradescantia 86 

Traganthus 104 

Tragia 102, 104 

Trattinnickia 71 

Trema 204 

Trianosperm 89 

Trianosperma 89 

Trianthema 144 

Tribulus 210 

Trichachne 175 

Trichanthera 42 

Trichilia 138, 139

Trichocentrum 160, 163 

Trichocolea 15 

TRICHOCOLEACEAE 15 

Trichogonia 60 

Trichomanes 32 

Trichopilia 160, 163 

Trichosalpinx 160, 163 

Trichospira 60, 61 

Trichosteleum 21 

Trichostigma 165 

Trichostomum 20-21 

Tridax 60 

Tridens 171 

Trifolium 109 

Trigonia 202 

TRIGONIACEAE 202 

Trigonidium 160, 163 

Trigynaea 46, 47 

Trilepis 94 

Trimezia 117 

Triodia 175 

Triphasia 190 

Triphora 160 

Triplaris 177 

Triplophyllum 38 

Tripodanthus 126 

Tripogandra 86 

Tripogon 171 

Tripsacum 171, 175 

Trisetella 160, 163 

Tristicha 176 

Triumfetta 202 

TRIURIDACEAE 202 

Triuris 202 

Trixis 61 

Trizeuxis 160 

Troschelia 112 

Trymatococcus 145, 146 

Tryssophyton 134 

Tschudya 138 

Turnera 203 

TURNERACEAE 203 

Tussacia 112 

Tylimanthus 9 

Tylopsacas 111 

Tylosperma 112 

Tynanthus 64 

Typha 203 

TYPHACEAE 203 

Typhonium 52, 53 

U 

Ugni 149 

Uleiorchis 160 

ULMACEAE 203-204 

Uncaria 186 

Unona 47 

Unonopsis 46 

Unxia 60 


Uragoga 189 

Urania 200 

Uranthera 138 

Urena 129, 130 

Urera 204 

Urochloa 171 

Urospatha 52 

Urostigma 146 

Urtica 204 

URTICACEAE 204 

Urvillea 191 

Utricularia 123 

V 

Vaccinium 98, 99 

Valeriana 204 

VALERIANACEAE 204 

Valota 175 

Vanaea 11 

Vandellia 196 

Vanilla 160, 163 

Vantanea 115 

Vasivaea 202 

Vatairea 107, 109 

Vataireopsis 107 

Vellosiella 195 

Vellozia 204 

VELLOZIACEAE 204 

Verbena 205 

VERBENACEAE 204-205 

Verbesina 60, 61 

Verdoornianthus 13 

Vernonanthura 60 

Vernonia 61 

Vesicularia 19 

Vetiveria 171 

Viburnum 78 

Victoria 151 

Vigna 107, 109 

Vilfa 175 

Viola 206 

VIOLACEAE 205-206 

Virola 147 

VISCACEAE 206-207 

Vismia 84, 85 

VITACEAE 207 

Vitellaria 195 

Vitex 205 

Vitis 207 

Vittaria 40 

VITTARIACEAE 39-40 

Vochysia 208 

VOCHYSIACEAE 207-208 

Votomita 134 

Vouacapoua 75 

Vouapa 76, 210 

Vouarana 191 

Voyara 78 

Voyria 110 

Voyriella 110 

Vriesea 68, 69 

W 

Waltheria 199 

Waria 47 

Warszewiczia 186 

Webera 138 

Websteria 94 

Weddellina 176 

Wedelia 60, 61 

Weigeltia 148 

Weinmannia 90 

Werauhia 69 

Whittonia 165 

Wijkia 21 

Wilbrandia 89 

Willughbeja 50 

Windsorina 180 

WINTERACEAE 208 

Wissadula 129, 130 

Witheringia 198 

Wittmackia 69 

Wolffia 122 

Wolffiella 122 

Wolffiopsis 122 

WOODSIACEAE 40 

Wulffia 60 

Wullschlaegelia 160, 163 

X 

Xanthosoma 52, 53 

Xerorchis 160 

Ximenia 153 

Xiphidium 112 

Xiphochaeta 60 

Xiphopteris 31 

Xylobium 160 

Xylophragma 64 

Xylopia 47 

Xylosma 109, 110 

Xylosterculia 199 

XYRIDACEAE 208-209 

Xyris 208, 209 

Y 

Youngia 60 

Yuyba 57 

Z 

Zamia 209 

ZAMIACEAE 209 

Zanthoxylum 190 

Zapoteca 142 

Zea 171 

227 

INDEX

228 

Zelometeorium 20 

Zephyranthes 124 

Zexmenia 61 

Zeyheria 65 

Zingiber 209 

ZINGIBERACEAE 209-210 

Ziziphus 180 

Zollernia 75, 76 

Zoopsidella 13 

Zoopsis 13 

Zornia 107, 109 

Zostera 210 

ZOSTERACEAE 210 

Zschokkea 50 

Zygia 142, 144 

Zygopetalum 163 

ZYGOPHYLLACEAE 210 

Zygosepalum 160, 163 

Zygostates 160 

INDEX 

CHECKLIST OF THE PLANTS OF THE GUIANAS

Pedro Acevedo R. 

Department of Botany, NHB 166 


Washington, DC 20560 

USA 


Lucile Allorge 

Museum National d'Histoire Naturelle 

Laboratoire de Phanerogamie 

16, rue Buffon 

F-75005 Paris 

FRANCE 

Christiane Anderson 

University of Michigan Herbarium 

North University Building 

University of Michigan 

Ann Arbor, MI 48109-1057 

USA 

William R. Anderson 

University of Michigan Herbarium 

North University Building 


Ann Arbor, MI 48109-1057 

USA 

Daniel F. Austin 

Department of Biological Sciences 

Florida Atlantic University 

Boca Raton, FL 33431-0991 

USA 

Gerardo A. Aymard C. 

UNELLEZ-Guanare, Programa R.N.R. 

Herbario Universitario 

Mesa de Cavacas 

Edo. Portuguesa 3323 

VENEZUELA 

APPENDIX A. 

Contributors and reviewers. 

Michael J. Balick 

New York Botanical Garden 

Bronx, NY 10458-5126 

USA 

Rupert C. Barneby 


Bronx, NY 10458-5126 

USA 

APPENDIX A 

229 

Cornelis C. Berg 

The Norwegian Arboretum and Botanical Garden 

University of Bergen 

N-5067 Store Milde 

NORWAY 

Paul E. Berry 

Missouri Botanical Garden 

P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

John K.Boggan 




USA 

John Brandbyge 

Skejbygardsvej 80 

DK-8240 Risskov 

DENMARK

230 

Kenneth M. Cameron 

Department of Biology 

University of North Carolina 

Chapel Hill, NC 27599-3280 

USA 

Eric A. Christenson 

1646 Oak Street 

Sarasota, FL 34236 

USA 

APPENDIX A 


Georges Cremers 

Herbier, Centre ORSTOM-Cayenne 

B.P. 165 

F-97323 Cayenne cedex (French Guiana) 

FRANCE 

Thomas B. Croat 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Douglas Daly 


Bronx, NY 10458-5126 

USA 

Alfonso Delgado-Salinas 

Herbario Nacional, Departamento de Botanica 

Instituto de Biologia, UNAM 

Apartado Postal 70-233 

04510 Mexico, D.F. 

MEXICO 

Pierre Detienne 

Centre Technique Forestier Tropical 

45 Bis Avenue de la Belle Gabrielle 

94736 Nogent sur Marne-Cedex 

FRANCE 

Laurence J. Dorr 




USA 

Robert B. Faden 




USA 

Paul R. Fantz 

Department of Horticultural Science 

North Carolina State University 

Box 7609, Raleigh, NC 27695-7609 

USA 

Christian Feuillet 

c/o Department of Botany, NHB 166 



USA 

Jeanne Florschütz-de Waard 

Institute of Systematic Botany 

P.O. Box 80.102 

Heidelberglaan 2 

3508 TC Utrecht 

THE NETHERLANDS 

James P. Folsom 

Huntington Botanical Gardens 

1151 Oxford Road 

San Marino, CA 91108 

USA 

Enrique Forero 


Bronx, NY 10458-5126 

USA

Vicki A. Funk 




USA 

Alwyn H. Gentry 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Lynn J. Gillespie 

Research Division 

Canadian Museum of Nature 

P.O. Box 3443, Station D 

Ottawa, Ontario K1P 6P4 

CANADA 


Sidney F. Glassman 

Herbarium, Botany Department 

Field Museum of Natural History 

Roosevelt Road at Lake Shore Drive 

Chicago, IL 60605 

USA 

A.R.A. Görts-van Rijn 


P.O. Box 80.102 




Paul Goetghebeur 

Laboratory of Plant Morphology, Systematics, 

and Ecology 

University Gent 

Ledeganckstraat 35 

9000 Gent 

BELGIUM 

Peter Goldblatt 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Eric J. Gouda 


P.O. Box 80.102 




S.R. Gradstein 


P.O. Box 80.102 




Jason R. Grant 

Department of Plant Biology 

University of Maryland 

College Park, MD 20742 

USA 

Jean-Jacques de Granville 


B.P. 165 


FRANCE 

James Grimes 


Bronx, NY 10458-5126 

USA 

Micheline Guerlesquin 

Laboratoire de Biologie végétale et de 

Phytogeographie 

Institut de Recherche 

3, Place Andre Leroy, B.P. 808 

49005 Angers Cedex 01 

FRANCE 

Eric Hagsater 

Asociacion Mexicana de Orquideologia, A.C. 



MEXICO 

APPENDIX A 

231

232 

APPENDIX A 


Madjit Hakki 

Botanischer Garten und Botanisches Museum 

Berlin-Dahlem 

Konigin-Luise-Strasse 6-8 

D-1000 Berlin 33 

GERMANY 

Bertel Hansen 

Botanical Museum and Herbarium 

Gothersgade 130 

DK-1123 Copenhagen 

DENMARK 

Ray M. Harley 

Herbarium, Royal Botanic Gardens 

Kew, Richmond, Surrey 

England TW9 3AB 

UNITED KINGDOM 

Robert R. Haynes 

Department of Biological Sciences 

Box 870344 

University of Alabama 

Tuscaloosa, AL 35487-0344 

USA 

Willem H.A. Hekking 


P.O. Box 80.102 




Paul Hiepko 


Berlin-Dahlem 



GERMANY 

Michel Hoff 

ORSTOM 

Institut de Botanique 

28, rue Goethe 

F-67083 Strasbourg cedex 

FRANCE 

Lauritz B. Holm-Nielsen 

Herbarium Jutlandicum, Botanical Institute 

University of Aarhus 

68 Nordlandsvej 

DK-8240 Risskov 

DENMARK 

Bruce K. Holst 

Marie Selby Botanical Gardens 

811 South Palm Avenue 


USA 

Charles N. Horn 

Biology Department 

Newberry College 

Newberry, SC 29108 

USA 

Marion J. Jansen-Jacobs 


P.O. Box 80.102 




C. Jeffrey 



TW9 3AB England 


Rudolf Jenny 

Moosweg 9 

3112 Allmendingen 

SWITZERLAND 

Emmet J. Judziewicz 

Department of Botany, Birge Hall 

University of Wisconsin 

430 Lincoln Drive 

Madison, WI 53706 

USA

Carol L. Kelloff 




USA 

Helen Kennedy 

Herbarium, Botany Department 

University of British Columbia 

3529-6270 University Boulevard 

Vancouver, BC V6T 2B1 

CANADA 


Robert W. Kiger 

Hunt Institute for Botanical Documentation 

Carnegie Mellon University 

Pittsburgh, PA 15213-3890 

USA 

Robert Kral 

Vanderbilt University 

Box 1705, Sta. B 

Nashville, TN 37235 

USA 

Job Kuijt 


University of Victoria 

P.O. Box 1700 

Victoria, BC V8W 2Y2 

CANADA 

David B. Lellinger 




USA 

Beat Ernst Leuenberger 


Berlin-Dahlem 



GERMANY 

Jan C. Lindeman 


P.O. Box 80.102 




Eduardo Lleras-Perez 

Centro Nacional de Recursos Geneticos 

Empresa Brasileira de Pesquisa Agropec 

Caixa Postal 10-2372 

70770 Brasilia, D.F. 

BRAZIL 

Pierre-André Loizeau 

Conservatoire et Jardin botaniques 

Case postale 60 

Ch-1292 Chambesy/GE 

SWITZERLAND 

Alicia Lourteig 


57 rue Cuvier 

F-75005 Paris 

FRANCE 

James L. Luteyn 


Bronx, NY 10458-5126 

USA 

Paul J.M. Maas 


P.O. Box 80.102 




H. Maas-van de Kamer 


P.O. Box 80.102 




APPENDIX A 

233

234 

Luis Marcano-Berti 

Kometenlaan 413 

3721 JM Bilthoven 


APPENDIX A 


Richard H. Maxwell 

Herbarium, Indiana University Southeast 

4201 Grant Line Rd. 

New Albany, IN 47150 

USA 

Willem Meijer 

Herbarium, T.H. Morgan School of Biological 

Sciences 

University of Kentucky 

Lexington, KY 40506-0225 

USA 

John D. Mitchell 


Bronx, NY 10458-5126 

USA 

Scott Mori 


Bronx, NY 10458-5126 

USA 

Gilberto Morillo 

Herbario L. Ruiz Terán, MERF 

Facultad de Farmacia, U.L.A. 

5101 Mérida, Edo. Mérida 

VENEZUELA 

Thomas Morley 

c/o Plant Biology Department 

University of Minnesota 

220 Biological Sciences Center 

1445 Gortner Ave. 

St. Paul, MN 55108-1095 USA 

Michael Nee 


Bronx, NY 10458-5126 

USA 

Dan H. Nicolson 




USA 

John J. Pipoly 

Botanical Research Insitute of Texas 

509 Pecan Street 

Ft. Worth, TX 76102 

USA 

Odile Poncy 




F-75005 Paris 

FRANCE 

Ghillean T. Prance 



England TW9 3AB 


John Pruski 




USA 

Peter H. Raven 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA

Harold Robinson 




USA 

Norman K.B. Robson 

Botany Department 

British Museum (Natural History) 

Cromwell Road 

London SW7 5BD, England 



William A. Rodrigues 

Departamento de Botânica-Herbário 

Universidade Federal do Paraná 

Centro Politécnico-Jardim das Américas 

C.P. 19031, 81531-970 Curitiba, Paraná 

BRASIL 

Adriaan C. de Roon 


P.O. Box 80.102 




Velva E. Rudd 

P.O. Box 19 

Reseda, CA 91337 

USA 

Daniel Sabatier 


B.P. 165 


FRANCE 

Claude Sastre 




F-75005 Paris 

FRANCE 

Beryl B. Simpson 

Department of Botany, BIO 308 

University of Texas 

Austin, TX 78713-7640 

USA 

Laurence E. Skog 




USA 

Flemming Skov 

National Environmental Research Institute 

Division of Wildlife Ecology, Kalo 

Grenavej 12, DK-8410 Ronde 

DENMARK 

Mario Sousa Sanchez 

Herbario Nacional, Departamento de Botanica 

Instituto de Biologia, UNAM 



MEXICO 

Clive A. Stace 

Herbarium, Department of Botany 

University of Leicester 

Leicester LE1 7RH, England 


Bruce Stein 

The Nature Conservancy 

1815 N. Lynn St. 

Arlington, VA 22209 

USA 

Dennis W. Stevenson 


Bronx, NY 10458-5126 

USA 

APPENDIX A 

235

236 

Mark T. Strong 




USA 

Charlotte M. Taylor 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Peter Taylor 

4 Laneland Cottages 

Plaistow Rd 

Kirdford, Billingshurst 

West Sussex RH14 0JY 


APPENDIX A 


Carol A. Todzia 

Lundell Herbarium, Department of Botany 

University of Texas 

Austin, TX 78712 

USA 

Warren H. Wagner, Jr. 



Ann Arbor, MI 48109 

USA 

Dieter Wasshausen 




USA 

Anna Weitzman 




USA 

Henk van der Werff 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Donald R. Windler 

Herbarium, Biology Department 

Towson State University 

8000 York Road 

Baltimore, MD 21204 

USA 

John J. Wurdack 




USA 

Elsa M. Zardini 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA


Maria Mercedes Arbo 

Facultad de Ciencias Agrarias 

Instituto de Botanica del Nordeste IBONE 

C.C. 209 

3400 Corrientes 

ARGENTINA 

William R. Buck 


Bronx, NY 10458-5126 

USA 

Robert A. DeFilipps 




USA 

Stefan Dressler 

Rijksherbarium/Hortus Botanicus 

P.O. Box 9514 

NL-2300 RA Leiden 


Bruce F. Hansen 


University of South Florida 

4202 East Fowler Avenue, LIF 169 

Tampa, FL 33620 

USA 

Andrew Henderson 


Bronx, NY 10458-5126 

USA 

APPENDIX B. 

Contributors and reviewers 

for the revised edition. 

Jacquelyn A. Kallunki 


Bronx, NY 10458-5126 

USA 

Denis M. Kearns 


P.O. Box 299 

St. Louis, MO 63166-0299 

USA 

Joseph H. Kirkbride, Jr. 

Systematic Botany and Mycology Laboratory 

USDA/ARS 

Building 011A, Room 304, BARC-West 

Beltsville, MD 20705-2350 

W. John Kress 




USA 

Antony J.M. Leeuwenberg 

Department of Plant Taxonomy 

Wageningen Agricultural University 

P.O.B. 8010, 6700 ED Wageningen 


Harry E. Luther 

Marie Selby Botanical Gardens 

811 South Palm Avenue 


USA 

APPENDIX B 

237

238 

Max E. Medley 


College of Arts and Sciences 

University of Louisville 

Louisville, KY 40292 

USA 

Heimo Rainer 

Institute of Botany 

University of Vienna 

Rennweg 14 

A-1030 Vienna 

AUSTRIA 

APPENDIX B 


Susanne S. Renner 

Institut für Spezielle Botanik und Botanischer 

Garten 

Johannes Gutenburg-Universität Mainz 

Anselm-F.-v.-Bentzel-Weg 2 

D-6500 Mainz 

GERMANY 

Marga C.M. Werkhoven 

National Herbarium of Suriname 

Anton de Kom Universiteit van Suriname 

Universiteitscomplex Leysweg P.O.B. 9212 

Paramaribo 

SURINAME 

John H. Wiersema 

Systematic Botany and Mycology Laboratory 

USDA/ARS 

Building 011A, Room 304, BARC-West 

Beltsville, MD 20705-2350 

USA

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