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Tropical Bryology Research Reports 

No. 5 http://www.tropicalbryologyresearch.co.uk/ 2004 

Checklist and distribution of the liverworts and 

hornworts of sub-Saharan Africa, including the 

East African Islands 

(edition 2, September 2004). 

Martin J. Wigginton 

36, Big Green, Warmington, Peterborough, PE8 6TU, U.K. 

September 2004

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Editors: Brian J. O'Shea and Martin J. Wigginton 

Copyright © 2004, Tropical Bryology Research 

Date of publication: September 2004 

Printed in United Kingdom 

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written permission from the publisher

Reports (2004) 5: 1-102 Research Bryology Tropical 

Checklist and distribution of the liverworts and hornworts of sub- 

Saharan Africa, including the East African Islands 

(edition 2, September 2004). 

Martin J. Wigginton 

36, Big Green, Warmington, Peterborough PE8 6TU, U.K. M.J.Wigginton@btinternet.com 

Abstract. The distribution by country or other geopolitical area is provided for each of the 1071 liverwort and 

hornwort taxa currently accepted in sub-Saharan Africa (i.e. Index Muscorum regions Afr 2, Afr 3 and Afr 4, 

excluding the part of Afr 4 to the south of South Africa). All country records are supported by at least one literature 

reference, and the number of taxa recorded for each country is given. The 1992 synonyms with their 

owning taxa are intercalated alphabetically in the list. 

Introduction 

Catalogues of the Hepaticae and Anthocerotae have been published for the East African Islands 

(Grolle, 1995 (1)) and for the rest of sub-Saharan Africa, including Atlantic Ocean islands and Socotra 

(Wigginton & Grolle 1996 (2)). A similar checklist of the bryophytes of North Africa was published 

by Ros et al. 1999 (3). These catalogues and checklist sought to bring together all published records of 

liverworts for these regions, and provided for each accepted taxon at least one literature reference for 

each of the countries in which it had been recorded. Together they summarised current knowledge of 

the distribution of liverworts throughout the continent. 

Since the publication of the East African Islands (1) and sub-Saharan African (2) catalogues, a considerable 

bulk of new data on the liverworts of these regions has been published, and this has provided 

the stimulus to produce a new checklist. Clearly, this checklist is not a substitute for (1) and (2): those 

publications contain much additional information and should be consulted for details of synonymy and 

literature sources for country records. Grolle 1995 (1) in addition provides essential notes on selected 

genera and on many excluded taxa. 

The area covered by this checklist includes all of the Index Muscorum regions Afr 2, Afr 3 and Afr 4, 

except for the following areas: 

1) the part of Afr 4 which lies to the south of the African mainland (south of latitude 35°S). This area 

encompasses cool-temperate and subantarctic islands from St. Paul and Amsterdam islands in the 

southern Indian Ocean to the Tristan da Cunha group in the southern Atlantic, all of whose floristic 

affinities are antipodal rather than African. A catalogue of the liverworts of this area, except the Tristan 

da Cunha group, was published in Grolle (2002). 

2) Western Sahara, whose floristic affinities are North African. There are, however, no bryophytes 

recorded from that geographical region (Ros et al. 1999). 

Catalogues (1) and (2) have provided the main basis for the present checklist, and most country records 

are referred to those publications, which should be consulted for details of primary literature. In 

the few years since the catalogues appeared, many new publications have provided additional country 

records, and new taxa and synonymy, the latter being listed in Appendix 1. Also incorporated in this 

checklist are: 

a) the correction of errors, including some authorities of names, which remained undetected prior to 

the publication of (2), and the inclusion of some country records from overlooked literature, 

b) the reassignment of taxa which were placed in (2) in the genera Aneura, Aspiromitus, Anthoceros, 

Fimbriaria, Hygrolejeunea, Madotheca, Schisma and Strepsilejeunea,

2 M. J. WIGGINTON 

c) the reassignment of some records to Eritrea or Ethiopia following the re-checking of the geographical 

locations of places cited in (121, 502). 

In sub-Saharan Africa, including the islands, 1071 taxa (1029 species and 47 additional taxa of lower 

rank) are currently accepted in 146 genera and 47 families of the Marchantiopsida and Anthocerotopsida. 

One family (Wiesnerellaceae), 8 genera (Allisoniella, Bryopteris, Geocalyx, Metalejeunea, 

Otolejeunea, Symbiezidium, Wiesnerella, Xylolejeunea) and 207 species or lower rank taxa have been 

recorded in Africa only from the East African Islands. 

The list of currently accepted taxa excludes those which have been placed in synonymy or have been 

published as 'Dubia', 'Excludenda' or 'Dubia. Indeterminate', but does include many taxa which have 

been insufficiently studied (or perhaps not re-examined since first described - which may be in the 

19th century), and a few of somewhat doubtful occurrence in Africa. It seems likely that at least some 

of the latter will eventually be placed in synonymy or rejected for other reasons. Insufficiently studied 

taxa are not differentiated in this checklist, and reference should be made to (1) and (2) for further information. 

Data for the checklist are held on the TAXA system (541), and the main taxon distribution list was 

generated directly from TAXA. The format of this checklist follows that established by O'Shea (171, 

540) for the Checklist of the mosses of sub-Saharan Africa, and subsequently adopted by Ros et al. (3) 

for the Bryophyte checklist of Northern Africa. 

The checklist is not a taxonomic document, and no new taxonomy is published here: it is merely a 

summation of published records. In its compilation, no judgements have been made by the author on 

the validity of taxa, or on the acceptability of regional records. Despite checking and re-checking, 

some errors doubtless remain undetected in this document, and literature overlooked, and the author 

would be very grateful to know of any of these. 

This checklist is published both as a printed version and an electronic version (via 

http://www.tropicalbryologyresearch.co.uk/) in order to make it as widely available as possible. It is 

intended that further editions will be published in the same way at intervals of a few years. 

Changes in edition 2 

1. The full genus name is given for all 'owners' of synonyms. 

2. In response to feedback on edition 1, a literature source is now provided for synonyms. The literature 

reference numbers that refer to source literature are placed in round brackets following the 

name of the owning taxon. 

For heterotypic synonyms, underlined literature reference numbers indicate the sources where the 

synonymy was first published, or where first publication seems a reasonable assumption. However, 

since 'syn. nov' is not stated in much of the earlier literature, there are probably some errors, particularly 

where the synonymy was first published in the nineteenth century. Where the place of first 

synonymy is unclear or I have not yet discovered it, the lit. ref. number is not underlined. Reference 

to early synonymy is sometimes supported by an additional reference (e.g. (973, 471) indicates 

that the synonym was first published in Mitten 1886, and the synonymy is accepted in 

Grolle's 1963 revision). 

For homotypic synonyms, no attempt has been made to find the earliest source, and the literature 

reference numbers for such synonyms are not underlined. 

3. Additional notes are added for some taxa. 

4. Changes since edition 1 are listed in Appendix 1. 

I should be most grateful to know of any errors in this list, and would welcome suggestions for things 

that could be useful in future editions. Comments please to M.J.Wigginton@btinternet.com 

Under consideration for the next edition is the inclusion of summary data on world distributions of 

sub-Saharan taxa that occur outside the continent.

CHECKLIST OF THE LIVERWORTS AND HORNWORTS OF SUB-SAHARAN AFRICA 3 

Acknowledgments 

I am indebted to Brian O'Shea for a careful reading of the text and detecting transcription and other errors.. 

However, any mistakes that may still remain in this document are entirely the responsibility of the author. 

Map1. The countries of Index Muscorum regions Afr 2, Afr 3 and Afr 4 covered in this checklist. 

AGA Agalega Islands 

ALD Aldabra 

AMI Amirante Islands 

ANG Angola 

ANN Annobón Island 

ASC Ascension Island 

BEN Benin 

BIO Bioko (Fernando Po). 

BKN Burkina Faso 

BOT Botswana 

BUR Burundi 

CAB Cabinda 

CAF Central African Republic 

CGS Chagos Archipelago 

CHA Chad 

CMN Cameroon 

COM Comoro Islands 

CON Congo-Brazzaville 

CVI Cape Verde Islands 

DIE Diego Garcia 

DJI Djibouti 

ERI Eritrea 

ETH Ethiopia 

GAB Gabon 

GAM Gambia 

GHA Ghana 

GNB Guinea Bissau 

GUI Guinea 

IVO Ivory Coast 

KEN Kenya 

LBR Liberia 

LES Lesotho 

MAU Mauritius 

MDG Madagascar 

MLI Mali 

MLW Malawi 

MOZ Mozambique 

MTN Mauritania 

NAM Namibia 

NGA Nigeria 

NGR Niger 

PRI Príncipe 

REU Réunion 

RIM Rio Muni 

ROD Rodrigues 

RWA Rwanda 

SAF South Africa 

SAO São Tomé 

SEN Senegal 

SEY Seychelles 

SIE Sierra Leone 

SOC Socotra 

SOM Somalia 

STH St. Helena 

SUD Sudan 

SWZ Swaziland 

TAN Tanzania 

TOG Togo 

UGA Uganda 

ZAI Democratic Republic 

of the Congo (former 

Zaire). 

ZAM Zambia 

ZIM Zimbabwe


Synopsis of families and genera 

Families are arranged alphabetically within each Order. The placement of genera closely follows 

Grolle 1983a (421), except for the Lejeuneaceae which follows Gradstein 1994 (122). The 1071 taxa 

are assigned to 146 genera in 47 families. The numbers in brackets against each genus indicate the 

number of taxa currently accepted within the genus (but this number includes some doubtful taxa 

which have not (yet) been synonymised or rejected, and taxa identified only to the genus). The numbers 

in a small font refer to the African Index Muscorum region(s) in which the genus has been recorded. 

Since the publication of Grolle 1995 (1) and Wigginton & Grolle 1996 (2), thirteen generic 

names have been added to the sub-Saharan list: Apomarsupella, Aureolejeunea, Kymatocalyx, 

Mnioloma, Paracromastigum, Sphenolobopsis and Xylolejeunea included in edition 1 of this checklist, 

and Amphicephalozia, Denotarisia, Pellia, Plicanthus, Sauteria and Southbya in the present edition. 

Capillolejeunea and Stenorrhipis have been placed in synonymy, Folioceros recognised at generic 

level, and Hyalolepidozia, which was removed in edition 1 of this checklist, is reinstated. 

Subclass Sphaerocarpiidae 

Order SPHAEROCARPALES 

Family Monocarpaceae 

Monocarpus (1) 4N 

Family Riellaceae 

Riella (5) 4N 

Family Sphaerocarpaceae 

Sphaerocarpos (1) 4N 

Subclass Marchantiidae 

Order MARCHANTIALES 

Family Aytoniaceae 

2 3 4N 

Asterella (17) 

Cryptomitrium (1) 4N 

2 4N 

Mannia (2) 

Plagiochasma (6) 

Reboulia (1) 2 

Family Cleveaceae 

2 4N 

Athalamia (2) 

Sauteria (1) 4N 

2 3 4N 

Subclass Metzgeriidae 

ORDER METZGERIALES 

Family Allisoniaceae 

Calycularia (1) 2 

Family Aneuraceae 

2 3 4N 

Aneura (4) 

Riccardia (23) 

2 3 4N 

Family Fossombroniaceae 

2 3 4N 

Fossombronia (35) 

Family Metzgeriaceae 

2 3 4N 

Metzgeria (8) 

Division Marchantiophyta (Hepaticae, Liverworts). 

CLASS MARCHANTIOPSIDA 

Family Exormothecaceae 

2 3 4N 

Exormotheca (3) 

Family Lunulariaceae 

2 4N 

Lunularia (1) 

Family Marchantiaceae 

2 3 4N 

Dumortiera (2) 

2 3 4N 

Marchantia (8) 

Family Oxymitraceae 

Oxymitra (2) 4N 

Family Ricciaceae 

2 3 4N 

Riccia (87) 

Ricciocarpos (1) 

2 4N 

Family Targioniaceae 

2 3 4N 

Cyathodium (3) 

2 3 4N 

Targionia (3) 

Family Wiesnerellaceae 

Wiesnerella (1) 3 

CLASS JUNGERMANNIOPSIDA 

Family Pallaviciniaceae 

2 3 4N 

Jensenia (1) 

2 3 4N 

Pallavicinia (2) 

2 3 4N 

Symphyogyna (3) 

2 3 

Symphyogynopsis (1) 

Family Pelliaceae 

Pellia (1) 2 

Order CALOBRYALES 

Family Haplomitriaceae 

2 4N 

Haplomitrium (2)

Subclass Jungermanniidae 

ORDER JUNGERMANNIALES 

Family Acrobolbaceae 

Acrobolbus (1) 4N 

2 3 4N 

Lethocolea (1) 

2 3 4N 

Marsupidium (1) 

2 3 4N 

Tylimanthus (3) 

Family Adelanthaceae 

2 3 4N 

Adelanthus (2) 

Family Antheliaceae 

Anthelia (1) 2 

Family Arnelliaceae 

Gongylanthus (4) 

Southbya (1) 2 


2 3 4N 

Family Balantiopsaceae 

2 3 4N 

Isotachis (1) 

Family Calypogeiaceae 

2 3 4N 

Calypogeia (9) 

2 3 4N 

Mnioloma (1) 

Family Cephaloziaceae 

2 3 4N 

Alobiellopsis (2) 

2 3 4N 

Cephalozia (3) 

2 3 

Iwatsukia (1) 

Odontoschisma (1) 

2 3 4N 

Family Cephaloziellaceae 

Allisoniella (1) 3 

Amphicephalozia (1) 3 

Cephalojonesia (1) 2 

2 3 4N 

Cephaloziella (18) 

2 3 4N 

Cylindrocolea (7) 

2 3 

Kymatocalyx (4) 

Family Chonecoleaceae 

Chonecolea (1) 2 

Family Geocalycaceae 

Clasmatocolea (2) 

2 3 

Conoscyphus (1) 

Geocalyx (1) 3 

Heteroscyphus (5) 

2 3 4N 

Leptoscyphus (4) 

2 3 4N 

Lophocolea (19) 

2 3 4N 

2 3 4N 

Family Gymnomitriaceae 

Apomarsupella (1) 2 

2 4N 

Gymnomitrion (2) 

2 3 4N 

Marsupella (3) 

Stephaniella (1) 4N 

Family Herbertaceae 

2 3 4N 

Herbertus (11) 

Family Jubulaceae 

2 3 4N 

Frullania (54) 

Family Jungermanniaceae 

2 3 4N 

Anastrophyllum (4) 

2 3 

Andrewsianthus (2) 

2 3 4N 

Cryptochila (1) 

Denotarisia (1) 3 

2 3 

Gottschelia (1) 

2 4N 

Gymnocoleopsis (2) 

2 3 4N 

Jamesoniella (3) 

2 3 4N 

Jungermannia (11) 

2 4N 

Lophozia (4) 

2 3 

Nardia (1) 

2 3 4N 

Notoscyphus (1) 

2 3 4N 

Plicanthus (4) 

Sphenolobopsis (1) 2 

2 3 

Syzygiella (3) 

Tetralophozia (1) 2 

2 4N 

Tritomaria (2) 

Family Lejeuneaceae 

Ptychanthoideae 

Ptychantheae 

2 3 

Acrolejeunea (4) 

Archilejeunea (11) 

Bryopteris (1) 3 

Caudalejeunea (11) 

2 3 4N 

Frullanoides (1) 

Lopholejeunea (22) 

2 3 

Marchesinia (5) 

2 3 

Mastigolejeunea (5) 

2 3 4N 

Ptychanthus (1) 

Schiffneriolejeunea (11) 

2 3 4N 

Thysananthus (2) 

Lejeuneoideae 

Brachiolejeuneae 

Acanthocoleus (3) 

Brachiolejeunea (1) 

Odontolejeunea (1) 

2 3 

Stictolejeunea (1) 

Symbiezidium (2) 3 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 

Lejeuneae 

Aureolejeunea (1) 2 

Ceratolejeunea (10) 

Cheilolejeunea (29) 

Cladolejeunea (1) 2 

Drepanolejeunea (21) 

Evansiolejeunea (1) 2 

2 3 

Haplolejeunea (2) 

2 3 

Harpalejeunea (2) 

2 3 4N 

Lejeunea (57) 

2 3 

Lepidolejeunea (3) 

2 3 

Leptolejeunea (5) 

Leucolejeunea (3) 

Metalejeunea (1) 3 

Microlejeunea (15) 

Otolejeunea (2) 3 

2 3 

Prionolejeunea (2) 

2 3 

Pycnolejeunea (1) 

2 3 4N 

Taxilejeunea (6) 

Xylolejeunea (2) 3 

2 3 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N


Cololejeuneae 

Aphanolejeunea (10) 

Cololejeunea (110) 

2 3 4N 

Colura (22) 

Diplasiolejeunea (29) 

Family Lepicoleaceae 

Lepicolea (1) 4N 

2 3 4N 

2 3 4N 

Family Lepidoziaceae 

Acromastigum (1) 4N 

2 3 4N 

Arachniopsis (2) 

2 3 4N 

Bazzania (18) 

Hyalolepidozia (1) 4N 

2 3 4N 

Kurzia (5) 

2 3 4N 

Lepidozia (14) 

Paracromastigum (1) 4N 

Psiloclada (1) 4N 

2 3 4N 

Telaranea (2) 

2 3 4N 

Family Mastigophoraceae 

2 3 

Mastigophora (1) 

ORDER ANTHOCEROTALES 

Family Anthocerotaceae 

2 3 4N 

Anthoceros (21) 

2 3 

Folioceros (2) 

2 3 4N 

Phaeoceros (15) 

Family Dendrocerotaceae 

Family Plagiochilaceae 

2 3 4N 

Plagiochila (70) 

Family Pleuroziaceae 

2 3 

Pleurozia (1) 

Family Porellaceae 

2 3 4N 

Porella (13) 

Family Pseudolepicoleaceae 

Blepharostoma (1) 2 

Family Radulaceae 

2 3 4N 

Radula (22) 

Family Scapaniaceae 

Diplophyllum (1) 2 

2 4N 

Scapania (4) 

Family Schistochilaceae 

2 3 4N 

Gottschea (3) 

2 3 4N 

Schistochila (3) 

Division Anthocerotophyta (Hornworts). 

CLASS ANTHOCEROTOPSIDA 

The Checklist 

Dendroceros (5) 

Megaceros (1) 2 

2, 3 

ORDER NOTOTHYLALES 

Family Notothyladaceae 

Notothylas (5) 2 

The checklist is in alphabetical order of taxon name, with current names shown in bold face italic font. 

Names placed in synonymy are indented and shown in a smaller font, together with the owning taxon. 

For synonymies, an = sign is used throughout, no distinction being made between nomenclatural and 

taxonomic synonyms. Taxa marked as 'Dubia' are those that have been reported from sub-Saharan Africa, 

but are now considered either to be of doubtful occurrence in the area, or are doubtfully distinct. 

Taxa marked as 'Excludenda' are those that have been recorded from the area, but are now either excluded 

from the African list, or the records referred to other taxa. These doubtful and excluded taxa are 

shown in a smaller font. 

Explanatory notes are appended to selected taxa, especially where there has been an amendment to the 

distribution given in (1) and (2). Except for Jovet-Ast, the authorities of names follow Brummitt & 

Powell 1992 (166), with post-1992 author names following the same rules, including the suppression 

of spaces between initials and surname. 

The countries in which each current taxon has been recorded are denoted by 3-character abbreviations, 

which follow Hollis & Brummitt 1992 (539), with few exceptions. The name of the country and the 

Index Muscorum region to which it is assigned is shown in Appendix 2. The countries are given in 

alphabetic order of the abbreviation and not of the country name, which means that some names are 

slightly out of sequence - for example, Mauritius (MAU) being listed before Madagascar (MDG). 

However, the alphabetic arrangement of abbreviations makes it easier to 'read' the checklist. Literature


references are given in brackets after each country abbreviation, and are translated in the References 

and Literature sections following this alphabetic list. 

Acanthocoleus aberrans Lindenb. et Gottsche) Kruijt ANG(2), CMN(2), KEN(2), MDG(1), NGA(2), 

SAF(2), TAN(2), UGA(2), ZAI(2), ZIM(2). Two varieties are recognised in the neotropics. African records may 

belong to var. laevis Gradst. (122), but this is yet to be confirmed. 

Acanthocoleus chrysophyllus (Lehm.) Kruijt CMN(2), IVO(53), KEN(2), LES(75), NGA(2), 

RWA(2), SAF(2,75), TAN(2), UGA(2), ZAM(2), ZIM(2). 

Acanthocoleus madagascariensis (Steph.) Kruijt BUR(2), COM(1), GUI(2), MAU(1), 

MDG(1,4,720), REU(1), RWA(2), SAF(2), SIE(2), TAN(2), ZIM(2). 

Acolea africana Steph. 1917 = Marsupella africana Steph. ex Bonner 1953 (416) = Apomarsupella africana 

(Steph. ex Bonner) R.M.Schust. 1996 (153). 

Acolea lacerata Steph. 1917 = Gymnomitrion laceratum (Steph.) Horik. 1943 (416). 

Acrobolbus bilobus (Mitt.) Steph. 1902 = Andrewsianthus bilobus (Mitt.) Grolle 1963 (315). 

Acrobolbus campylodontus (Hook.f. et Taylor) Steph. 1902 = Adelanthus decipiens (Hook.) Mitt. 1864 (827, 

465). 

Acrobolbus excisus (Mitt.) Schiffn. 1893 = Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust. 1961 

(451). 

Acrobolbus madagascariensis Steph. 1902 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa 1999 

(74). 

Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust. SAF(2). 

Acrolejeunea abnormis (Gottsche) Pearson 1892 = Phragmicoma abnormis Gottsche 1882 (1, 35) [Dubia(1)]. 

Acrolejeunea africana Steph. 1912, nom. inval. TAN(2). [Dubia]. This name seems only to have used by 

Stephani, and the identity of the collections is uncertain. The name is not mentioned in Gradstein 1975 (35). 

Acrolejeunea angustispica Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Acrolejeunea aulacophora (Mont.) Steph. COM(1), MAU(1), MDG(1), MOZ(2), REU(1), TAN(2). 

Listed in error from Uganda in (2). 

Acrolejeunea borgenii Steph. 1892 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (35). 

Acrolejeunea confertissima Steph. 1892 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975 

(35). 

Acrolejeunea convexa Steph. 1911 = Evansiolejeunea renistipula (Steph.) Vanden Berghen 1948 (851) = 

Evansiolejeunea roccatii (Gola) Vanden Berghen 1960 (850). 

Acrolejeunea emergens var. emergens (Mitt.) Steph. ALD(34,35), ANG(2), BEN(11), BUR(2), 

CAF(2), COM(1), CON(2), CVI(2,37), GHA(2), GUI(2), IVO(2,53), KEN(2), MAU(1,4), 

MDG(1,720), MLW(2,563), NGA(2), REU(1), RIM(10), ROD(1,723), RWA(2), SEN(2), SEY(1,4), 

SIE(2), TAN(2), UGA(2,42,50,96), ZAI(2), ZAM(2). 

Acrolejeunea emergens var. confertissima (Steph.) Gradst. CMN(2), NGA(2), RIM(10), SEN(2), 

SIE(2). 

Acrolejeunea ferruginea Steph. 1895 = Schiffneriolejeunea ferruginea (Steph.) Gradst. 1974 (832). 

Acrolejeunea fulva (Gottsche) Pearson 1892 = Phragmicoma fulva Gottsche 1882 (35) = Schiffneriolejeunea 

pappeana (Nees) Gradst. 1974 (831). 

Acrolejeunea fuscescens Gola 1907 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 (35). 

Acrolejeunea inflexa (Gottsche) Pearson 1892 = Phragmicoma inflexa Gottsche 1882 (1) [Dubia(1)]. 

Acrolejeunea molleri (Steph.) Paris 1906 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (832). 

Acrolejeunea occulta Steph. 1888 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974 (832). 

Acrolejeunea orchidae Pearson 1922 = Ptychocoleus molleri (Steph.) Steph. (834) = Schiffneriolejeunea 

polycarpa (Nees) Gradst. 1974 (35). 

Acrolejeunea pappeana (Nees) Schiffn. 1894 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (35). 

Acrolejeunea parviloba Steph. Nov 1890 = Schiffneriolejeunea parviloba (Steph.) Gradst. 1974 (832). 

Acrolejeunea pusilla Steph. 1908, nom. nud. [non (Steph.) Grolle et Gradst. 1974] = Acrolejeunea emergens 

(Mitt.) Steph. 1895 (35). 

Acrolejeunea pycnoclada (Taylor) Schiffn. GHA(2), MDG(1), NGA(2), REU(1), SEY(1), ZAI(2). 

Acrolejeunea renauldii Steph. 1888, nom. inval. = Acrolejeunea aulacophora (Mont.) Steph. 1895 (832, 35). 

Acrolejeunea renauldii (Steph.) Steph. 1912 = Acrolejeunea aulacophora (Mont.) Steph. 1895 (832, 35). 

Acrolejeunea roccatii Gola 1907 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960 (850). 

Acromastigum exile (Lindenb.) A.Evans SAF(2). 

Adelanthus decipiens (Hook.) Mitt. BIO(2,33), CMN(2), COM(4), ETH(2), MAU(1), MDG(720), 

MLW(2,109), REU(1), RWA(142), SAF(2,75), STH(2), TAN(2), UGA(42,50), ZAI(2). 

Adelanthus lindenbergianus (Lehm.) Mitt. KEN(2), LES(75), MAU(1), MDG(1), MLW(109),


REU(1), RWA(2), SAF(2,75), TAN(2), UGA(2), ZAI(2). 

Adelanthus sphalerus (Hook.f. et Taylor) Steph. 1908 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864 

(827, 465). 

Adelanthus unciformis (Hook.f. et Taylor) Spruce 1876 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864 

(1105, 465). 

Aitonia appendiculata (Lehm. et Lindenb.) Kuntze 1891 = Plagiochasma appendiculata Lehm. et Lindenb. 

1832 (254) 

Aitonia eximia Schiffn. in Steph. 1895 = Plagiochasma eximium (Schiffn. in Steph.) Steph. 1898 (254). 

Aitonia fischeriana Steph. 1895 = Plagiochasma appendiculatum Lehm. et Lindenb. 1832 (254). 

Aitonia microcephala Steph. 1895 = Plagiochasma microcephalum (Steph.) Steph. 1898 (254). 

Alicularia flexuosa (Lehm.) Nees 1844 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (883, 97). 

Alicularia vermicularis (Lehm.) Gottsche et al. 1844 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 

(97). 

Allisoniella nigra (Rodway) R.M.Schust. REU(1). 

Alobiella borbonica (Steph.) Steph. 1908 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (1). 

Alobiella chevalieri Steph. 1908 = Cylindrocolea chevalieri (Steph.) R.M.Schust. 1969 (615). 

Alobiella heteromorpha (Lehm.) Steph. 1908 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969 

(844). 

Alobiella pillansii (Sim) S.W.Arnell 1963 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969 (844). 

Alobiellopsis heteromorpha (Lehm.) R.M.Schust. LES(75), MAS(1), MAU(1), MLW(114), SAF(2). 

Alobiellopsis pillansii (Sim) R.M.Schust. SAF(2). 

Amphicephalozia geisslerae Pócs et Váňa MDG(712,720). 

Anastrophyllum auritum (Lehm.) Steph. BIO(36), CMN(2), ETH(2), KEN(2,71,665,759), LES(75), 

MLW(114), REU(45), RWA(2), SAF(2,75), TAN(2), UGA(2,42,96), ZAI(2). 

Anastrophyllum bessonii Steph. 1893 = Gottschelia schizopleura (Spruce) Grolle 1968 (901). 

Anastrophyllum calcaratum Steph. 1911 = Anastrophyllum gambaragarae Gola 1907 (7) = Anastrophyllum 

auritum (Lehm.) Steph. 1901 (918). 

Anastrophyllum gambaragarae Gola 1907 = Anastrophyllum auritum (Lehm.) Steph. 1901 (918). 

Anastrophyllum grossitextum Steph. 1911 = Anastrophyllum gambaragarae Gola 1907 (7) = Anastrophyllum 

auritum (Lehm.) Steph. 1901 (918). 

Anastrophyllum hintzeanum Steph. 1917 = Anastrophyllum auritum (Lehm.) Steph. 1901 (918). 

Anastrophyllum leucocephalum (Taylor) Steph. 1893 = Anastrophyllum auritum (Lehm.) Steph. 1901 (778). 

Anastrophyllum minutum (Schreb.) R.M.Schust. CMN(2), KEN(2), SAF(2,75), TAN(2), UGA(2), 

ZAI(667). 

Anastrophyllum obtusatum (Hook.f. et Taylor) Steph. 1901 = Anastrophyllum subcomplicatum (Lehm. et 

Lindenb.) Steph. 1893 (917, 163). 

Anastrophyllum piligerum (Nees) Steph. ASC(2), MAU(1), MDG(1,4,720), MLW(114), REU(1), 

RWA(2), SEY(1,4), TAN(2), ZIM(2). 

Anastrophyllum schizopleurum (Spruce) Steph. 1893 = Gottschelia schizopleura (Spruce) Grolle 1968 (901). 

Anastrophyllum subcomplicatum (Lehm. et Lindenb.) Steph. STH(2). 

Andrewsianthus aberrans (Nees et Mont.) Grolle MAU(1), MDG(4), REU(1,4). 

Andrewsianthus bilobus (Mitt.) Grolle BIO(2,33,36), CMN(2), ETH(2), KEN(759), RWA(2), 

TAN(2), UGA(2,776), ZAI(2). Listed in error from South Africa in (2); the specimen is Gymnocoleopsis capensis 

(163, 164). 

Andrewsianthus jamesonii (Mont.) Váňa 1983 = Lophozia jamesonii (Mont.) R.M.Schust. 2002 (779). 

Andrewsianthus kilimanjaricus (S.W.Arnell) Grolle et Váňa 1980 = Lophozia jamesonii (Mont.) 

R.M.Schust. 2002 (974). Placed in synonymy in Gradstein et al, 1983 (with Andrewsianthus jamesonii), but 

Schuster (2002) provisionally treated Lophozia (Andrewsianthus) kilimanjarica as a distinct species. 

Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust. 1966. [Excludenda(416)]. Reports from Africa 

are based on misidentifications. 

Aneura amazonica Spruce 1885 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979 (846, 760). 

Aneura angusticosta Steph. 1913 "angusticostata" = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 

(845) = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979 (351, 760). 

Aneura caespitans Steph. 1891 = Riccardia sp. (1). 

Aneura comosa Steph. 1890 = Riccardia sp. (1). 

Aneura compacta Steph. 1893 = Riccardia compacta (Steph.) S.W.Arnell 1952 (717). 

Aneura congoana Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893 (845). 

Aneura congoensis Steph. 1917 = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 (845) = Riccardia 

amazonica (Spruce) Schiffn. ex Gradst. 1979 (351, 760). 

Aneura corbieri Steph. 1917 = Riccardia sp. (1). 

Aneura dismieri Steph. 1917 = Riccardia sp. (1).


Aneura erosa Steph. 1891 = Riccardia erosa (Steph.) E.W.Jones 1956 (845). 

Aneura erronea Steph. TAN(2). 

Aneura exigua Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893 (847). 

Aneura fastigiata (Lehm.) Gottsche et al. 1846 = Riccardia fastigiata (Lehm.) Trevis. 1877 (351). 

Aneura grosselimbata Steph. 1917 = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 (845) = Riccardia 


Aneura holstii Steph. 1917 = Riccardia holstii (Steph.) E.W.Jones 1956 (845). 

Aneura inconspicua Steph. 1893 = Riccardia erosa (Steph.) E.W.Jones 1956 (845). 

Aneura keniae Gola 1914 = Riccardia sp. (845). 

Aneura latissima Spruce MAU(1), PRI(128), REU(1). 

Aneura laurentiana Steph. 1917 = Riccardia pinguis (L.) Gray (351, 652) = Aneura pinguis (L.) Dumort. 

1831 sens. lat. (652, 845). 

Aneura lepervanchei Steph. 1899 = Riccardia sp. (1). 

Aneura limbata Steph. 1891 = Riccardia limbata (Steph.) E.W.Jones 1956 (351). 

Aneura longispica Steph. 1890 = Riccardia longispica (Steph.) Pearson 1893 (845, 351). 

Aneura multifida (L.) Dumort. 1822 = Riccardia multifida (L.) Gray 1821 (653). 

Aneura nudiflora Steph. 1890 = Riccardia nudiflora (Steph.) Grolle 1995 (1). 

Aneura palmata (Hedw.) Dumort. 1831 = Riccardia palmata (Hedw.) Carruth. 1865 (1) [Excludenda(1)]. 

Aneura papulosa Steph. 1917, hom. illeg. [non Steph. 1893] = Riccardia longispica (Steph.) Pearson 1893 

(847). 

Aneura piliflora Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893 (845, 351). 

Aneura pinguis (L.) Dumort. MAS(1), SAF(652), SEY(1), TAN(2). 

Aneura pinguis (L.) Dumort. sens. lat. ANG(2), CMN(2), GHA(2), NGA(2), SAO(2), SIE(2), 

ZAI(2). 

Aneura pinnatifida (F.Weber) Nees 1838 = Riccardia pinnatifida (F.Weber) Carruth. 1865 (1) [Excludenda(1)]. 

Aneura pseudopinguis (Herzog) Pócs IVO(2), LES(75), MLW(114), RIM(10), SAF(2), SEY(4), 

TAN(2), UGA(42,50), ZAI(2), ZIM(2). 

Aneura ramosissima Steph. 1891 = Riccardia ramosissima (Steph.) Grolle 1995 (845). 

Aneura reticulata Steph. 1891 = Riccardia limbata (Steph.) E.W.Jones 1956 (845). 

Aneura saccatiflora Steph. 1890 = Riccardia saccatiflora (Steph.) S.W.Arnell 1952 (351). 

Aneura stephanii Besch. ex Steph. 1899 = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 (845) = 

Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979 (351, 760). 

Aneura submarginata S.W.Arnell 1952 = Riccardia pinguis (L.) Gray (845) = Aneura pinguis (L.) Dumort. 

1831 sens. lat. (652). 

Aneura tenera Steph. 1917 = Riccardia erosa (Steph.) E.W.Jones 1956 (845). 

Aneura travisiana Pearson 1921 = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 (845) = Riccardia 


Anomalolejeunea decemplicata Steph. 1913 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 1980 

(878). 

Anomalolejeunea pluriplicata (Pearson) Schiffn. 1893 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 

1980 (878). 

Anomalolejeunea pluriplicata var. tabularis S.W.Arnell 1953 = Anomalolejeunea pluriplicata (Pearson) 

Schiffn. 1893 (7) = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 1980 (878). 

Anthelia hirtella (F.Weber) Steph. 1891 = Plicanthus hirtellus (F.Weber) R.M.Schust. 2003 (780). 

Anthelia juratzkana (Limpr.) Trevis. TAN(2). 

Anthoceros aethyopicus Gola 1914 = Phaeoceros sp. 

Anthoceros brunnthaleri Steph. 1913 = Phaeoceros sp. 

Anthoceros buettneri Steph. GHA(2), TOG(2). In (2), listed in error from Sierra Leone. 

Anthoceros caucasicus Steph. SIE(2), ZAI(2). 

Anthoceros chevalieri Steph. ZAI(2). 

Anthoceros communis Steph. 1897 = Phaeoceros carolinianus (Michx.) Prosk. 1951 (990). 

Anthoceros cristatus Steph. ASC(2). 

Anthoceros decurvus Mitt. 1860 = Notothylas decurva (Mitt.) Steph. 1901 (988). 

Anthoceros dichotomus Raddi 1818 = Phaeoceros bulbiculosus (Brot.) Prosk. 1954 (992). 

Anthoceros dilatatus Steph. BIO(2,33). 

Anthoceros dimorphus Sim SAF(2), ZIM(2). 

Anthoceros ecklonii Steph. 1916 = Phaeoceros carolinianus (Michx.) Prosk. 1951 (170). 

Anthoceros flagellaris Mitt. 1873 = Megaceros flagellaris (Mitt.) Steph. 1916 (993). 

Anthoceros floribundus Steph. ASC(2). 

Anthoceros fuciformis Mont. 1843 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975 (996).


Anthoceros fulvisporus Steph. 1916 = Phaeoceros fulvisporus (Steph.) J.Haseg. 1993 (991). 

Anthoceros fusiformis Austin MAU(1). 

Anthoceros gasangorii Gola KEN(2). 

Anthoceros incurvus Steph.1893 = Folioceros incurvus (Steph.) D.C.Bharadwaj 1972 (1103). 

Anthoceros javanicus Nees 1830 = Dendroceros javanicus (Nees) Gottsche et al. 1846 (994). 

Anthoceros jollyanus Steph. 1916 = Phaeoceros jollyanus (Steph.) S.W.Arnell 1956 (813). 

Anthoceros laevis L. 1753 = Phaeoceros laevis (L.) Prosk. 1951 (1) [Excludenda(1)]. 

Anthoceros mandonii Steph. 1916 = Anthoceros caucasicus Steph. 1914 (947). 

Anthoceros mildbraedii Steph. 1923 = Phaeoceros sp. 

Anthoceros minutus Mitt. 1877 = Phaeoceros minutus (Mitt.) S.W.Arnell 1963 (170). 

Anthoceros myriandroecius Steph. KEN(2,665), RWA(2), TAN(2), UGA(96,115). 

Anthoceros natalensis Steph. LES(75), MLW(2), SAF(2). 

Anthoceros parvifrons Steph. 1912 = Phaeoceros sp. 

Anthoceros pinnatus Steph. CMN(2), SAO(2), TAN(2). 

Anthoceros pseudocostus Steph. 1916 = Phaeoceros sp. 

Anthoceros punctatus L. CVI(2,37), RWA(2), SOC(764), ZAI(2). 

Anthoceros aff. punctatus L. ZIM(9). This plant was published in (9) as Anthoceros aff. laevis, but in error (pers. 

comm. H.J. During, 2002). 

Anthoceros rossoi Gola 1914 = Phaeoceros sp. 

Anthoceros sambesianus Steph. LES(75), RHO(2), RWA(2), UGA(115). 

Anthoceros schroederi Steph. TAN(2). 

Anthoceros stuhlmannii Steph. 1916 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975 (131). 

Anthoceros tenuissimus Steph. 1893 = Phaeoceros sp. 

Anthoceros tigrinus Gola 1914 = Phaeoceros sp. 

Anthoceros usambarensis Steph. 1916 probably = Phaeoceros carolinianus (Michx.) Prosk. 1951 (170). 

Anthoceros volkensii Steph. TAN(2). 

Anthoceros sp. (Aspiromitus dismieri Steph.) MDG(1). 

Anthoceros sp. (Aspiromitus expansus Steph.) CMN(2). Probably a species of Anthoceros, as Stephani 1916 

(886) described it as having "frons cavernosa ... sporae fuscae". 

Anthoceros sp. (Aspiromitus hildebrandtii Steph.) MDG(1). 

Anthoceros sp. ROD(723). 

Aphanolejeunea capensis (S.W.Arnell) S.W.Arnell BUR(2), MLW(2,563), RWA(2), SAF(2), 

TAN(2), UGA(50), ZAI(2), ZIM(2). Pócs synonymised A. capensis (South Africa) with A. mamillata (383), but 

later (384) considered that the synonymy needed further proof. Accordingly, the records included under A. capensis for 

the time being, pending a resolution of this question. 

Aphanolejeunea corniculata Tixier in Pócs 1984, nom. inval. MAD(1). [Dubia(1)]. 

Aphanolejeunea diaphana (A.Evans) R.M.Schust. 1980. hom. illeg. = Aphanolejeunea subdiaphana (Jovet- 

Ast) Pócs 1984 (383) = Aphanolejeunea truncatifolia Horik. 1934 (117). 

Aphanolejeunea exigua A.Evans 1911 = Aphanolejeunea microscopica var. exigua (A.Evans) Pócs et 

Lücking 1996 (117). 

Aphanolejeunea exigua var. africana Pócs 1984 = Aphanolejeunea microscopica var. africana (Pócs) Pócs 

et Lücking 1996 (117). 

Aphanolejeunea fadenii (Pócs) Pócs KEN(2), RWA(2), TAN(2), UGA(115). 

Aphanolejeunea jovetastiana Pócs CMN(2), GHA(2), IVO(2), RIM(10). 

Aphanolejeunea lisowskii Pócs RIM(10), UGA(115), ZAI(2). 

Aphanolejeunea mamillata (Ångstr.) Steph. 1916. Perhaps synonymous with Aphanolejeunea capensis - see note 

under the latter species. 

Aphanolejeunea microscopica var. africana (Pócs) Pócs et Lücking KEN(2), MLW(563), RWA(2), 

TAN(2), UGA(42,96), ZAI(2). 

Aphanolejeunea microscopica var. exigua (A.Evans) Pócs et Lücking COM(1), IVO(2), KEN(2), 

MDG(1), MLW(563), RWA(2), TAN(2), UGA(42), ZAI(2). 

Aphanolejeunea microscopica var. microscopica (Taylor) A.Evans COM(4), MLW(563), TAN(2), 

UGA(42). 

Aphanolejeunea minima Tixier CMN(2). 

Aphanolejeunea moramangae (Tixier) Pócs COM(1), MDG(1), RIM(10), TAN(2), UGA(42,115), 

ZAI(2). 

Aphanolejeunea runssorensis Steph. 1916 = Cololejeunea runssorensis (Steph.) Pócs 1975 (378). 

Aphanolejeunea subdiaphana (Jovet-Ast) Pócs 1984 = Aphanolejeunea truncatifolia Horik. 1934 (117). 

Aphanolejeunea truncatifolia Horik. BIO(72), MLW(563), UGA(115). 

Aphanolejeunea zenkeri Steph. 1916 = Cololejeunea zenkeri (Steph.) E.W.Jones 1954 (858).


Apomarsupella africana (Steph. ex Bonner) R.M.Schust. KEN(2), RWA(2), TAN(2), UGA(2), 

ZAI(2). 

Arachniopsis coactilis Spruce 1882 = Arachniopsis diacantha (Mont.) M.Howe 1902 (382). 

Arachniopsis diacantha (Mont.) M.Howe ANN(2,33), BIO(36), COM(4), GHA(2), IVO(2), 

KEN(71,665), MAU(1), MDG(1,4,720), MLW(2,92), MOZ(2), REU(1), RIM(10), RWA(2), 

SAF(2), SEY(1,4,13), SIE(2), TAN(2), UGA(42,50), ZAI(2), ZAM(2), ZIM(2). 

Arachniopsis diplopoda Pócs COM(4), MDG(1,4), REU(4). The record from D.R. Congo (ZAI), formerly 

considered to be doubtful, is now discounted. This species is now regarded as endemic to the East African Islands (Pócs 

1995 (4)). 

Arachniopsis madagascariensis Steph. 1922 = Arachniopsis diacantha (Mont.) M.Howe 1902 (382). 

Arachniopsis succulenta (Sim) S.W.Arnell 1955 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Archilejeunea abbreviata (Mitt.) Vanden Berghen ANG(2), CAF(2), CMN(2), GAB(2), GHA(2), 

GUI(2), MLW(563), NGA(2), RIM(10), SIE(2), TAN(2), UGA(50). 

Archilejeunea africana Steph. BIO(2,33), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), IVO(53), 

NGA(2), PRI(48), RIM(10), SIE(2), TAN(2), UGA(42,50,96), ZAI(2). 

Archilejeunea alata Steph. COM(1), MAU(1), MDG(1). 

Archilejeunea apiculata Pearson 1931 = Marchesinia excavata (Mitt.) Schiffn. 1893 (891). 

Archilejeunea autoica Vanden Berghen CAF(2), CMN(2), GAB(2), GHA(2), IVO(2,53), NGA(2), 

RIM(10), SIE(2), UGA(42,115). 

Archilejeunea brevilobula Steph. MDG(1). 

Archilejeunea elobulata Steph. CAF(2), IVO(2,53), SAF(2), TAN(2), ZAI(2). 

Archi-lejeunea erronea Steph. 1888 = Leucolejeunea clypeata (Schwein.) A.Evans 1907 [Excludenda]. 

Grolle 1995 (1) placed the South African Archilejeunea erronea in synonymy with Leucolejeunea rotundistipula, but 

Gradstein & Geissler 1997 (55) re-examined the holotype, and found it inseparable from Leucolejeunea clypeata, a 

species endemic to eastern North America. Because of this extreme disjunction, they concluded that specimens had 

been confused by Stephani, and that there had been a labelling error. 

Archilejeunea jonesii Vanden Berghen CAF(2), IVO(53), NGA(2), RIM(10). 

Archilejeunea linguifolia Steph. CAF(2), CMN(2), GAB(2), IVO(2), NGA(2), RIM(10), SIE(2), 

TOG(2), ZAI(2). 

Archilejeunea mariana (Gottsche) Steph. 1897 = Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 

1989 (1) [Excludenda(1)]. 

Archilejeunea mauritiana Lindenb. ex Steph. MAS(1), MAU(1), TAN(2). 

Archilejeunea nobilis (Gottsche) Steph. MDG(1). 

Archilejeunea palustris Sim 1926 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122, 170). 

Archilejeunea pappeana (Nees) Sim 1926 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (833). 

Archilejeunea parvistipula Steph. 1924 = Archilejeunea africana Steph. 1911 (907). 

Archilejeunea planiuscula (Mitt.) Steph. SEY(1). 

Archilejeunea pobeguinii Steph. 1911 = Archilejeunea abbreviata (Mitt.) Vanden Berghen 1951 (907). 

Archilejeunea rotalis (Hook.f. et Taylor) Steph. 1911 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 

Geissler 1997 (55). 

Archilejeunea rotundistipula (Lindenb. ex Lehm.) Sim 1926 = Leucolejeunea rotundistipula (Lindenb. ex 

Lehm.) A.Evans 1907 (806). 

Archilejeunea staudtiana Steph. 1911 = Archilejeunea africana Steph. 1911 (907). 

Archilejeunea subinermis Steph. 1911, nom. nud. = Archilejeunea africana Steph. 1911 (907). 

Archilejeunea taxiantha Herzog, in sched. = Archilejeunea africana Steph. 1911 (907). 

Archilejeunea xanthocarpa (Lehm. et Lindenb.) Steph. 1895 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907 (430). 

Aspiromitus dilatatus (Steph.) Steph. 1916 = Anthoceros dilatatus Steph. 1887 (979). 

Aspiromitus dismieri Steph. 1916 = Anthoceros sp. (1). 

Aspiromitus expansus Steph. 1916 = Anthoceros sp. (2). 

Aspiromitus fuciformis (Mont.) Steph. 1916 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975 (1). 

Aspiromitus hildebrandtii Steph. 1916 = Anthoceros sp. (1). 

Aspiromitus incurvus (Steph.) Steph. 1916 = Anthoceros incurvus Steph. 1893 (1103). 

Aspiromitus maranguensis Steph. 1916 = Phaeoceros maranguensis (Steph.) Bapna 1967 (995). 

Aspiromitus pinnatus (Steph.) Steph. 1916 = Anthoceros pinnatus Steph. 1887 (979). 

Aspiromitus schroederi Steph. 1916 = Anthoceros schroederi Steph. 1912 (979). 

Asterella abyssinica (Gottsche) Grolle BUR(2), CMN(2), ETH(2), MLW(85), RWA(2), SAF(85), 

SIE(2), TAN(2), ZAI(2), ZIM(2). 

Asterella angolensis (Steph.) S.W.Arnell 1963 = Asterella wilmsii (Steph.) S.W.Arnell 1963 (997).


Asterella bachmannii (Steph.) S.W.Arnell BOT(2), LES(2,75), MLW(2,29), SAF(2,75), SWZ(2), 

ZAI(2), ZIM(2). 

Asterella boryana (Mont.) H.A.Mill. 1981, nom. illeg. = Asterella syngenesica (Bory) Grolle 1974 (849, 1). 

Asterella coronata (Steph.) H.A.Mill. MDG(1). 

Asterella dissoluta (Steph.) Grolle TAN(2). 

Asterella khasyana (Griff.) Pandé et al. BUR(93), UGA(93). 

Asterella linearis (Steph.) M.Howe KEN(2,71), TAN(2). 

Asterella marginata (Nees) S.W.Arnell SAF(2). 

Asterella muscicola (Steph.) S.W.Arnell BOT(2), LES(2,75), MLW(2), SAF(2), ZIM(2). 

Asterella pappii (Gola) Grolle ERI(121), ETH(2), SOC(2). 

Asterella syngenesica (Bory) Grolle MAU(1), REU(1). 

Asterella wilmsii (Steph.) S.W.Arnell ANG(2), LES(2,75), MDG(1), MLW(2,29), MOZ(2), 

SAF(2,75), SWZ(2), ZIM(2). 

Asterella sp. (Fimbriaria gigantea Steph.) TAN(2). Presumed Asterella sp. 

Asterella sp. (Fimbriaria incrassata Steph.) ETH(2). Provisionally placed in synonymy with Asterella abyssinica 

by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria kamerunensis Steph.) CMN(2). Provisionally placed in synonymy with Asterella 

abyssinica by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria pirottae Gola) ETH(2). Presumed Asterella sp. 

Asterella sp. (Fimbriaria preussii Schiffn. ex Steph.) CMN(2). Provisionally placed in synonymy with Asterella 

bachmannii by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria volkensii Steph.) KEN(2), TAN(2), UGA(2). Provisionally placed in synonymy 

with Asterella dissoluta by Long & Grolle, in litt. 

Athalamia pulcherrima (Steph.) S.Hatt. ETH(2). 

Athalamia spathysii (Lindenb.) S.Hatt. NAM(2), SOC(764), TAN(2). 

Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler STH(2,49,55). 

Aytonia abyssinica (Gola) Cufod. 1952 = Plagiochasma rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 

1898 (502). 

Aytonia appendiculata var. erythraeae (Gola) Cufod. 1952 = Plagiochasma appendiculatum Lehm. et Lindenb. 

1832 (254). 

Aytonia beccariana (Steph.) Cufod. 1952 = Plagiochasma beccarianum Steph. 1898 (254). 

Aytonia eximia Schiffn. ex Steph. 1895 = Plagiochasma eximium (Schiffn. ex Steph.) Steph. 1898 (254). 

Aytonia microcephala Steph. 1895 = Plagiochasma microcephalum (Steph.) Steph. 1898 (254). 

Aytonia rupestris J.R.Forst. et G.Forst. 1775 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 

(254). 

Aytonia rupestris var. albociliata (Lindenb. et Nees) Cufod. 1952 = Plagiochasma rupestre (J.R.Forst. et 

G.Forst.) Steph. 1898 (254). 

Aytonia schimperi (Steph.) Cufod. 1952 = Plagiochasma eximium (Schiffn.) Steph. 1898 (254). 

Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 REU(1). [Excludenda(1)]. 

Bazzania approximata Onr. SEY(1,4). 

Bazzania borbonica (Steph.) Steph. 1891 = Bazzania praerupta (Reinw. et al.) Trevis. 1877 (1). 

Bazzania boschiana (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 

1877 (823). 

Bazzania comorensis Steph. COM(1). 

Bazzania convexa (Lindenb.) Trevis. 1877 = Bazzania nitida (F.Weber) Grolle 1960 (888). 

Bazzania cucullata Onr. SEY(1,4). 

Bazzania curvidens Steph. MDG(1). 

Bazzania deciduifolia Onr. MDG(1). 

Bazzania decrescens (Lehm. et Lindenb.) Trevis. BIO(36), COM(1,4), CON(2), GAB(2), 

KEN(2,731), MAS(1), MAU(1), MDG(1,720), MLW(2,92), MOZ(2), REU(1), RWA(2), SAF(2), 

SEY(1,4), TAN(2), UGA(2), ZAI(2), ZAM(2), ZIM(2). 

Bazzania decrescens Mitt. in Melliss 1875, nom. inval. = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 

1877. 

Bazzania decrescens var. ambahatrae Pócs MDG(720). 

Bazzania decrescens subsp. molleri (Steph.) E.W.Jones ANN(2,33), BIO(72), CMN(2), CON(2), 

GAB(2), GHA(2), IVO(51), MLW(92), MOZ(2), RIM(10), SAO(2), SIE(2), ZAI(2), ZIM(2). 

Bazzania decrescens subsp. pumila (Mitt.) Pócs COM(4), KEN(731), MDG(4), MLW(92), RWA(2), 

TAN(2), UGA(50), ZAI(2). 

Bazzania fusca Steph. 1893 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (884, 1).


Bazzania integra (Nees et Mont.) Steph. 1894 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 

(884, 1). 

Bazzania lacouturei (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania sp. [B. decrescens complex] (1). 

Bazzania laxifolia (Steph.) Demaret 1942 [= Mastigobryum laxifolium Steph. in Mildbr. 1911] = Bazzania 

roccatii Gola 1907 (7). 

Bazzania longa (Nees) Trevis. 1877 MAU(1). [Excludenda(1)]. 

Bazzania madagassa (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania sp. [B. decrescens complex]. (1). 

Bazzania mascarena (Steph.) Herzog COM(1), MDG(1), REU(1). 

Bazzania molleri Steph. 1891 = Bazzania decrescens subsp. molleri (Steph.) E.W.Jones 1975 (870). 

Bazzania nitida (F.Weber) Grolle BIO(36), BUR(2), CMN(2), COM(1), MAU(1,4), MDG(1,4,720), 

MLW(2,92), REU(1), RWA(2), SAF(2), SEY(1,4), TAN(2), UGA(50), ZAI(2). 

Bazzania papillosa S.W.Arnell MAU(1). 

Bazzania pellucida Sim 1926 = Acromastigum exile (Lindenb.) A.Evans 1934 (170). 

Bazzania perrotana (Steph.) E.W.Jones MDG(1). 

Bazzania perrotana (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania mascarena (Steph.) Herzog 1947 

(823). 

Bazzania praerupta (Reinw. et al.) Trevis. BIO(36), MAS(1), MAU(1), MDG(1,4), MLW(2,92), 

REU(1), SAF(2), TAN(2). 

Bazzania pulvinata Steph. 1891 = Bazzania decrescens subsp. pumila (Mitt.) Pócs 1994 (889). 

Bazzania pumila Mitt. 1886 = Bazzania decrescens subsp. pumila (Mitt.) Pócs 1994 (889). 

Bazzania radicans Sim 1926 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (170). 

Bazzania reflexa (Gottsche) Steph. MDG(1). 

Bazzania robertii (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 

1877 (870, 1). 

Bazzania roccatii Gola CMN(2), MLW(92), RWA(2), TAN(2), UGA(2,42), ZAI(2). 

Bazzania schismoidea (Steph.) Steph. 1891 = Bazzania borbonica Steph. 1894 (870) = Bazzania praerupta 

(Reinw. et al.) Trevis. 1877 (1). 

Bazzania seychellarum Gottsche ex Steph. 1893 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 

(847). 

Bazzania sp. (Mastigobryum lacouturei Steph. = Bazzania lacouturei (Steph.) S.W.Arnell, nom. inval.) 

MDG(1). 

Bazzania sp. (Mastigobryum madagassum Steph. = Bazzania madagassa (Steph.) S.W.Arnell, nom. inval.) 

MDG(1). 

Bazzania stephanii (J.B.Jack) Steph. 1891 = Bazzania nitida (F.Weber) Grolle 1960 (870). 

Bazzania stolonifera (Sw.) Trevis. 1877 REU(1). [Excludenda(1)]. 

Bellincinia abyssinica (Nees) Kuntze 1891 = Porella abyssinica (Nees) Trevis. 1877 (502). 

Blasia javanica Sande Lac. 1856 = Notothylas javanica (Sande Lac.) Gottsche 1858 (1003). 

Blepharostoma cavallii Gola 1907 = Tetralophozia cavallii (Gola) Váňa 1993 (892). 

Blepharostoma sejunctum Ångstr. 1877 = Arachniopsis diacantha (Mont.) M.Howe 1902 (848). 

Blepharostoma trichophyllum (L.) Dumort. KEN(2), RWA(2), TAN(2), UGA(2,96). 

Blytia byssophora (Lehm. et Lindenb.) Nees 1846 = Pallavicinia lyellii (Hook.) Carruth. 1865 (146). 

Brachiolejeunea africana Steph. 1912 = Mastigolejeunea florea (Mitt.) Steph. 1912 (906). 

Brachiolejeunea assimilis Steph. 1911 = Mastigolejeunea nigra (Steph.) Steph. 1895 (795, 906). 

Brachiolejeunea bidens Steph. 1912 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 

auriculata (Wilson) Schiffn. 1893 (835). 

Brachiolejeunea camerunensis E.W.Jones et Vanden Berghen 1951 = Brachiolejeunea tristis Steph. 1912 

(850) = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Brachiolejeunea camerunensis var. angolensis S.W.Arnell 1962 = Frullanoides tristis (Steph.) van Slageren 

1985 (795). 

Brachiolejeunea confertifolia Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Brachiolejeunea crenata Sim 1926 = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Brachiolejeunea hildebrandtii Steph. 1912 = Mastigolejeunea auriculata (Wilson) Schiffn. 1893 (795). 

Brachiolejeunea hildebrandtii var. pluriplicata Gola 1914 = Frullanoides tristis (Steph.) van Slageren 1985 

(795). 

Brachiolejeunea laxifolia (Taylor) Schiffn. 1893 SAO(2). [Dubia(795)]. 

Brachiolejeunea natalaensis Sim 1926 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 (795). 

Brachiolejeunea nigra (Steph.) Steph. 1912 = Mastigolejeunea nigra Steph. 1895 (25). 

Brachiolejeunea parviflora Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. MDG(1), SAF(2), ZIM(2). The types of Dicranolejeunea 

lillieana, from Kenya, placed by Kruijt & Gradstein 1986 (133) under Brachiolejeunea phyllorhiza, are species of 

Acanthocoleus (Gradstein 1994 (122)).


Brachiolejeunea rhodesica Vanden Berghen 1951 = Mastigolejeunea rhodesica (Vanden Berghen) 

E.W.Jones 1957 (134). 

Brachiolejeunea thomeensis Steph. 1912 = Brachiolejeunea laxifolia (Taylor) Schiffn. 1893. (795) [Dubia(795)]. 

Brachiolejeunea tristis (Steph.) Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Brachiolejeunea usambarensis Steph. 1912 = Brachiolejeunea nigra (Steph.) Steph. 1912 (906) = Mastigolejeunea 

nigra Steph. 1895 (25). 

Bryopteris filicina (Sw.) Nees 1845 MAU(1). [Excludenda(1)]. 

Bryopteris filicina var. fruticosa Gottsche et al. 1845, nom. inval. = Dendrolejeunea fruticosa (Lindenb. et 

Gottsche) Lacout. 1908 (1) [Excludenda(1)]. 

Bryopteris fruticosa Lindenb. et Gottsche 1847 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) Lacout. 

1908 (1) [Excludenda(1)]. 

Bryopteris gaudichaudii Gottsche MDG(1,4), REU(1). 

Bryopteris madagascariensis Steph. 1912 = Bryopteris gaudichaudii Gottsche 1857 (890). 

Bryopteris madagassa Steph. 1902 = Bryopteris gaudichaudii Gottsche 1957 (890). 

Bryopteris trinitensis (Lehm. et Lindenb.) Gottsche et al. 1845 = Bryopteris filicina (Sw.) Nees 1845 (122) 

[Excludenda(795)]. 

Calobryum blumii Nees 1846 = Haplomitrium blumii (Nees) R.M.Schust. 1963 (1004). 

Calycularia crispula Mitt. ETH(2), KEN(71), MLW(2,29), TAN(2), ZAM(2). 

Calycularia golae Gerola 1947 = Calycularia crispula Mitt. 1861 (104). 

Calypogeia afrocaerulea E.W.Jones BIO(36), BUR(2), CMN(2), KEN(2), REU(4), RWA(2), 

SAO(2), SEY(4), TAN(2), UGA(42,50), ZAI(2). 

Calypogeia annabonensis Steph. ANN(2,33), GUI(2), KEN(2), MDG(720), SEY(1,4), TAN(2). 

Calypogeia arguta Nees et Mont. BUR(2), MAU(1,4), MDG(1), MLW(2), REU(1,4), SAF(2), 

SWZ(2), TAN(2), ZAI(2). 

Calypogeia bidentula (F.Weber) Nees LES(75), MDG(1), REU(1), RWA(2), SAF(2,75), ZIM(2). 

Calypogeia bidentula var. grandiretis S.W.Arnell 1963 = Calypogeia bidentula (F.Weber) Nees 1845 (253). 

Calypogeia capensis S.W.Arnell 1953 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (253, 118). 

Calypogeia ericetorum Raddi 1818 = Gongylanthus ericetorum (Raddi) Nees 1836 (862). 

Calypogeia fissa (L.) Raddi ANG(2), ANN(2,33), CMN(2), COM(4), KEN(71), REU(4), RWA(2), 

SAF(2), SAO(2), SIE(2), SWZ(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Calypogeia fusca (Lehm.) Steph. 1908 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Calypogeia latifolia Steph. 1908 = Calypogeia arguta Nees et Mont. 1838 (253). 

Calypogeia longifolia Steph. BUR(2), LES(2), MDG(1), SAF(2), SIE(2), SWZ(2), ZIM(2). 

Calypogeia mascarenensis Bischl. COM(4), MAU(1), MDG(1), REU(1,4). 

Calypogeia microstipula (Steph.) Steph. COM(1), MDG(1), REU(1,4), SAF(2). 

Calypogeia muelleriana (Schiffn.) Müll.Frib. ETH(2). 

Calypogeia repandifolia Steph. 1924 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (253, 118). 

Campylolejeunea peculiaris (Herzog) Amakawa 1960 = Cololejeunea inflectens (Mitt.) Benedix 1953 (878). 

Capillolejeunea mascarena S.W.Arnell 1965 = Drepanolejeunea mascarena (S.W.Arnell) R.L.Zhu et Grolle 

2003 (765) 

Cardiolejeunea dipterocarpa (E.W.Jones) R.M.Schust. 1980 = Lejeunea dipterocarpa E.W.Jones 1972 

(430). 

Cardiolejeunea papilionacea (Steph.) R.M.Schust. 1980 = Lejeunea papilionacea Steph. 1892 (430, 670). 

Carpolepidum dichotomum P.Beauv. 1805 = Plagiochila dichotoma (P.Beauv.) Nees et Mont. 1836 (988). 

Carrpos sphaerocarpus (D.J.Carr) Prosk. 1961 = Monocarpus sphaerocarpus D.J.Carr 1956 (1034). 

Caudalejeunea acutifolia Gerola ETH(2). 

Caudalejeunea africana (Steph.) Steph. BIO(2,33), ETH(2), GHA(2), IVO(2), KEN(2), NGA(2), 

SAF(2), TAN(2), ZAI(2). 

Caudalejeunea dusenii Steph. CMN(2), GHA(2), NGA(2), TAN(2), UGA(96). 

Caudalejeunea fruticosa (Lindenb. et Gottsche) Steph. 1912 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) 

Lacout. 1908 [Excludenda(1)]. 

Caudalejeunea grolleana Gradst. MDG(1). 

Caudalejeunea hanningtonii (Mitt.) Schiffn. BEN(22), CMN(2), GHA(2), GNB(2), GUI(2), 

IVO(2,53), KEN(2,731), MDG(1), MLW(563), NGA(2), PRI(48), RIM(10), SAF(2), SAO(48), 

SIE(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Caudalejeunea katangensis Vanden Berghen ZAI(2). 

Caudalejeunea lehmanniana (Gottsche) A.Evans BEN(22), CMN(2), GNB(2), NGA(2), SIE(2). 

Caudalejeunea lessonii Steph. 1895 = Caudalejeunea recurvistipula (Gottsche) Schiffn. 1893 (613). 

Caudalejeunea lewallei Vanden Berghen BUR(2), ETH(2), MDG(1), MLW(563), RWA(2), TAN(2),


UGA(50,96), ZAI(2), ZIM(2). 

Caudalejeunea madagassa Steph. 1912 = Caudalejeunea recurvistipula (Gottsche) Schiffn. 1893 (1). 

Caudalejeunea recurvistipula (Gottsche) Schiffn. COM(1), MDG(1). 

Caudalejeunea reniloba (Gottsche) Steph. 1912 MAU(1), REU(1). [Excludenda(1)]. 

Caudalejeunea stephanii Spruce ex Steph. 1912 = Caudalejeunea reniloba (Gottsche) Steph. 1912 (613) 


Caudalejeunea tricariniata E.W.Jones 1953 = Caudalejeunea lehmanniana (Gottsche) A.Evans 1907 (430). 

Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones CMN(2), CON(2), GHA(23), RWA(2), 

UGA(2,42), ZAI(2). 

Caudalejeunea zenkeri Steph. 1912 = Caudalejeunea hanningtonii (Mitt.) Schiffn. 1893 (891). 

Caudalejeunea sp. (Dicranolejeunea africana Steph.) MAU(1). 

Cephalojonesia incuba Grolle et Vanden Berghen KEN(2), NGA(2), ZAI(2). 

Cephalozia africana Váňa RWA(2), ZAI(2). 

Cephalozia "atro-virens" Sim 1926, nom. inval. - a typographical error for C. atroviridis Sim 1926. 

Cephalozia atro-viridis Sim 1926 = Cylindrocolea atroviridis (Sim) Váňa 1979 (156). 

Cephalozia bicuspidata (L.) Dumort. KEN(2), MLW(2,114), REU(1), RWA(2), SAF(2), TAN(2), 

UGA(2,42), ZAI(2). 

Cephalozia borbonica (Steph.) Besch. 1895 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (1). 

Cephalozia caffra J.B.Jack ex Bonner 1963, nom. nud. = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Cephalozia capensis Sim 1926 = Cephaloziella capensis (Sim) S.W.Arnell 1952 (494). 

Cephalozia cavifolia S.W.Arnell 1956 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Cephalozia connivens subsp. fissa Váňa BIO(2,33), CMN(2), CON(2), GAB(2), GHA(2), 

MDG(1,4,720), MLW(114), NGA(2), PRI(2), RIM(10), SAF(2), SAO(2), TAN(2), TOG(2), 

UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Cephalozia connivens var. flagellifera Pearson 1886 = Cephalozia connivens subsp. fissa Steph. 1891 (894, 

494). 

Cephalozia crassicaulis Steph. 1908 = Cephalozia connivens subsp. fissa Steph. 1891 (494). 

Cephalozia denudata var. africana Pearson 1887 = Odontoschisma africanum (Pearson) Sim 1926 (894, 

888). 

Cephalozia fissa Steph. = Cephalozia connivens ssp. fissa (Steph.) Váňa 1988 (494) 

Cephalozia grossitexta Steph. 1908 = Cephalozia connivens subsp. fissa Steph. 1891 (494). 

Cephalozia heteromorpha (Lehm.) Trevis. 1877 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969 

(494). 

Cephalozia jishibae Steph. 1924 = Iwatsukia jishibae (Steph.) N.Kitag. 1965 (472). 

Cephalozia kiaeri (Austin) Pearson 1887 = Cephaloziella kiaeri (Austin) Douin 1920 (494). 

Cephalozia ligulata (Steph.) Besch. 1895 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (937). 

Cephalozia lycopodioides Sim 1926 = Cephaloziella lycopodioides (Sim) S.W.Arnell 1952 (494). 

Cephalozia minutissima Kiaer et Pearson 1893 = Cephaloziella kiaeri (Austin) Douin 1920 (494). 

Cephalozia natalensis Sim 1926 = Cephaloziella natalensis (Sim) S.W.Arnell 1952 (494). 

Cephalozia nematodes Gottsche ex Austin 1879 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902 

(382). 

Cephalozia pillansii Sim 1926 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969 (494, 844). 

Cephalozia radicans Sim 1926 = Cephaloziella kiaeri (Austin) Douin 1920 (170, 494). 

Cephalozia robusta Steph. 1908 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Cephalozia tenuissima (Lehm. et Lindenb.) Trevis. 1877 = Cephaloziella tenuissima (Lehm. et Lindenb.) 

Steph. 1893 (494). 

Cephalozia vaginans (Steph.) Steph. 1924 = Cephaloziella vaginans Steph. 1911 (494). 

Cephalozia vallis-gratiae Gottsche ex Steph. 1908 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Cephalozia villaumei Steph. 1924 = Cephaloziella sp. (494). 

Cephalozia vulcanicola Steph. 1911 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Cephalozia welwitschii Steph. 1908 = Cephaloziella welwitschii (Steph.) Douin 1920 (494). 

Cephaloziella abyssinica Gola ETH(2). 

Cephaloziella anthelioides S.W.Arnell SAF(2). 

Cephaloziella "atro-virens" (Sim) S.W.Arnell 1955, nom. inval. = Cephaloziella atroviridis (Sim) 

E.W.Jones 1958 (158) = Cylindrocolea atroviridis (Sim) Váňa 1979 (156). 

Cephaloziella atroviridis (Sim) E.W.Jones 1958 = Cylindrocolea atroviridis (Sim) Váňa 1979 (156). 

Cephaloziella biokoensis Váňa et F.Müller BIO(775). 

Cephaloziella capensis (Sim) S.W.Arnell SAF(2). 

Cephaloziella chevalieri (Steph.) Grolle 1964 = Cylindrocolea chevalieri (Steph.) R.M.Schust. 1969 (615). 

Cephaloziella garsidei S.W.Arnell SAF(2), UGA(50).


Cephaloziella flagellaris S.Hatt. 1948 = Iwatsukia jishibae (Steph.) N.Kitag. 1965 (472). 

Cephaloziella gittinsii E.W.Jones 1960 = Cylindrocolea gittinsii (E.W.Jones) R.M.Schust. 1972 (615). 

Cephaloziella jishibae (Steph.) S.Hatt. 1952 = Iwatsukia jishibae (Steph.) N.Kitag. 1965 (472). 

Cephaloziella kiaeri (Austin) Douin CMN(2), KEN(759), LES(17), MDG(1,720), MLW(2,114), 

MOZ(2), REU(1), SAF(2,75), SEY(1), TAN(2), ZAI(2). 

Cephaloziella leonensis S.W.Arnell 1952 = Cephaloziella capensis (Sim) S.W.Arnell 1952 (170). 

Cephaloziella lycopodioides (Sim) S.W.Arnell SAF(2). 

Cephaloziella natalensis (Sim) S.W.Arnell SAF(2). 

Cephaloziella nigerica E.W.Jones 1960 = Cylindrocolea nigerica (E.W.Jones) R.M.Schust. 1972 (615). 

Cephaloziella pentagona Schiffn. ex Douin 1920 = Cephaloziella willisana (Steph. ) N.Kitag. 1969 (1035) = 

Cephaloziella kiaeri (Austin) Douin 1920 (156). 

Cephaloziella pillansii (Sim) S.W.Arnell 1955 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969 (844). 

Cephaloziella retusa E.W.Jones 1960 = Cylindrocolea chevalieri (E.W.Jones) R.M.Schust. 1969 (898, 615). 

Cephaloziella schelpei S.W.Arnell SAF(2). 

Cephaloziella tabularis S.W.Arnell SAF(2). 

Cephaloziella tenuissima (Lehm. et Lindenb.) Steph. LES(75), SAF(2,75). 

Cephaloziella transvaalensis S.W.Arnell MAU(719), MLW(114), SAF(2), TAN(659), UGA(50), 

ZIM(143). 

Cephaloziella triplicata S.W.Arnell SAF(2). 

Cephaloziella ugandica E.W.Jones 1960 = Cylindrocolea ugandica (E.W.Jones) R.M.Schust. 1972 (615). 

Cephaloziella umtaliensis S.W.Arnell MLW(114), MOZ(2), UGA(776), ZIM(2). 

Cephaloziella vaginans Steph. LES(75), MLW(114), REU(45), RWA(2), SIE(2), TAN(2), 

UGA(776), ZAI(2). Listed in error from Cameroon in (2). 

Cephaloziella vaginans var. camerunensis S.W.Arnell CMN(2). 

Cephaloziella welwitschii (Steph.) Douin ANG(2). 

Cephaloziella sp. (Cephalozia villaumei Steph.) MDG(1). 

Ceratolejeunea aberrans Steph. 1923 = Cladolejeunea aberrans (Steph.) Zwickel 1933 (1036). 

Ceratolejeunea belangeriana (Gottsche) Steph. COM(1), MAU(1), MDG(1,720), REU(1), SEY(1). 

Listed in error from Tanzania in (2). 

Ceratolejeunea beninensis E.W.Jones et Vanden Berghen BEN(2), CMN(2), GHA(2), IVO(53), 

NGA(2). 

Ceratolejeunea boschiana Steph. 1913 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 (1). 

Ceratolejeunea calabariensis Steph. ANN(2,33), BIO(2,33), CMN(2), COM(4), CON(2), GAB(2), 

GHA(2), IVO(2,53), MDG(4,720), MLW(563), NGA(2), PRI(2), RIM(10), SEY(4,108), SIE(2), 

TAN(2), UGA(42,96), ZAI(2). 

Ceratolejeunea cornuta (Lindenb.) Steph. 1895 MAD(1), REU(1). [Excludenda(1)]. 

Ceratolejeunea cornutissima Steph. 1892 perhaps = Ceratolejeunea diversicornua Steph. 1891 (733). 

Ceratolejeunea diversicornua Steph. CMN(2), GHA(2), ZAI(2). 

Ceratolejeunea floribunda Steph. SAO(2). 

Ceratolejeunea jungneri Steph. 1895 = Ceratolejeunea calabariensis Steph. 1895 (27). 

Ceratolejeunea mascarena (Steph.) Steph. 1913 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 

(847). 

Ceratolejeunea mauritiana (Steph.) Steph. 1892 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 (1). 

Ceratolejeunea papuliflora Steph. MDG(1). 

Ceratolejeunea renauldii Steph. 1913 = Lejeunea alata Gottsche 1845 (828). 

Ceratolejeunea saxbyi Pearson 1921 = Ceratolejeunea calabariensis Steph. 1895 (25). 

Ceratolejeunea stictophylla Herzog ex Vanden Berghen NGA(2). 

Ceratolejeunea umbonata Steph. BIO(2,33), CMN(2). 

Ceratolejeunea usambarensis Steph. 1913 = Ceratolejeunea jungneri Steph. 1895 (732) = Ceratolejeunea 

calabariensis Steph. 1895 (27). 

Ceratolejeunea variabilis (Lindenb.) Schiffn. MAU(1), MDG(1), REU(1). Dauphin (790) placed C. variabilis 

in synonymy with C. cornuta. Both species were reported in the 19th century from the East African Islands. Grolle 

(1) referred a collection from Madagascar to C. belangeriana, but whether all African records should be likewise referred, 

or some to C. cornuta remains to be seen. 

Ceratolejeunea zenkeri Steph. BIO(2,33), CMN(2), GAB(2), GHA(2), IVO(53), NGA(2), SIE(2), 

TAN(2), ZAI(2). 

Chandonanthus birmensis Steph. 1899 = Plicanthus birmensis (Steph.) R.M.Schust. 2002 (780). 

Chandonanthus cavallii (Gola) S.W.Arnell 1956 = Blepharostoma cavallii Gola 1907 (7) = Tetralophozia 

cavallii (Gola) Váňa 1993 (892). 

Chandonanthus giganteus Steph. 1911 = Plicanthus giganteus (Steph.) R.M.Schust. 2002 (780).


Chandonanthus hirtellus (F.Weber) Mitt. 1873 subsp. hirtellus = Plicanthus hirtellus (F.Weber) 

R.M.Schust. 2002 (780). 

Chandonanthus hirtellus subsp. giganteus (Steph.) Vanden Berghen 1965 = Plicanthus giganteus (Steph.) 

R.M.Schust. 2002 (780). 

Chandonanthus perloi Gola 1914 = presumed Plicanthus sp., probably P. hirtellus. 

Chandonanthus quadrifidus Steph. 1911 = Chandonanthus cavallii (Gola) S.W.Arnell 1956 (7) = Tetralophozia 

cavallii (Gola) Váňa 1993 (892). 

Chandonanthus setiformis sensu Vanden Berghen 1965 [non (Ehrh.) Lindb.] = Tetralophozia cavallii (Gola) 

Váňa 1993 (780). 

Cheilolejeunea africana (Steph.) E.W.Jones 1954 = Cheilolejeunea silvestris var. involuta (Steph.) 

E.W.Jones 1974 (867) = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (872). 

Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle ASC(2), STH(2). 

Cheilolejeunea brachytoma (Gottsche) R.M.Schust. 1980 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 

1964 (847). 

Cheilolejeunea brevifissa (Gottsche) R.M.Schust. 1963 = Cheilolejeunea krakakammae (Lindenb.) 

R.M.Schust. 1963 (878). 

Cheilolejeunea camerunensis S.W.Arnell CMN(2), NGA(2). 

Cheilolejeunea compressa (Herzog) Grolle COM(1). 

Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell SAF(2). 

Cheilolejeunea cordigera (Steph.) Grolle MAU(1), MDG(1,720). 

Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones BIO(2,33), CMN(2), ETH(2), 

KEN(2,731,759), MDG(1), MLW(563), REU(1), TAN(2), UGA(2). 

Cheilolejeunea crenulata Steph. in Paris 1902, nom. inval. = Cheilolejeunea principensis Steph. ex Paris 

1888, nom. inval (856) = Cheilolejeunea serpentina (Mitt.) Mizut. 1963 (574). 

Cheilolejeunea cucullata Steph. 1914 = Haplolejeunea cucullata (Steph.) Grolle 1979 (580). 

Cheilolejeunea decurrens (Steph.) X.-L.He 1995 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (2). 

Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. CAF(2), CMN(2), COM(1), GUI(2), 

MDG(1,720), MLW(2), REU(1), SIE(2), TAN(2), ZAI(2). 

Cheilolejeunea diversifolia Augier CMN(2). 

Cheilolejeunea ecarinata Vanden Berghen REU(1,4). 

Cheilolejeunea exigua Steph. 1914 = Lejeunea caespitosa Lindenb. 1845 (157). 

Cheilolejeunea exinnovata E.W.Jones GHA(2), SIE(2), ZIM(2). 

Cheilolejeunea inflata Steph. 1914 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897 (574). 

Cheilolejeunea intertexta (Lindenb.) Steph. BEN(22), CMN(2), GHA(2), IVO(2), MDG(1,720), 

NGA(2), REU(1), SAF(2), SIE(2), TAN(2), UGA(42,50). 

Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. BUR(2), COM(1), ETH(2), KEN(2,731,759), 

MDG(1,720), MLW(563), REU(1), RWA(2), SAF(2,75), TAN(2), UGA(2,42), ZAI(2), ZIM(2). 

Cheilolejeunea kurzii Steph. 1890 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897 (574). 

Cheilolejeunea latiflora Steph. CAF(2). 

Cheilolejeunea latistipula Steph. 1913 = Lejeunea wilmsii (Steph.) Steph. 1915 [= Eulejeunea wilmsii Steph. 

1892] (158) = Lejeunea eckloniana Lindenb. 1845 (869). 

Cheilolejeunea laurentii Steph. ZAI(2). 

Cheilolejeunea longiflora (Taylor) R.M.Schust. 1980 SAO(2). [Dubia(2)]. 

Cheilolejeunea madagassa Steph. 1914 = Cheilolejeunea principensis Steph. ex Paris 1888, nom. inval 

(856) = Cheilolejeunea serpentina (Mitt.) Mizut. 1963 (574). 

Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 1989 SEY(1). [Excludenda(1)]. 

Cheilolejeunea montagnei (Gottsche) R.M.Schust. COM(1), KEN(2), MAU(1), MDG(1,720), 

REU(1,4), RWA(2), SAO(2), TAN(2), ZAI(2). 

Cheilolejeunea multiflora Steph. 1895 = Lepidolejeunea multiflora (Steph.) R.M.Schust. 1980 (582). 

Cheilolejeunea newtonii Steph. ex Schiffn. CAF(2), CON(2), ETH(2), GHA(2), GUI(2), KEN(2), 

NGA(48), PRI(2), SIE(2). 

Cheilolejeunea obtusistipula Steph. 1914 = Lejeunea obtusistipula (Steph.) E.W.Jones 1989 (879). 

Cheilolejeunea omphalogastria Pócs RWA(2). 

Cheilolejeunea pluriplicata (Pearson) R.M.Schust. ANG(2), ETH(2), KEN(2,71), MDG(1), 

MLW(563), MOZ(2), SAF(2), TAN(2), UGA(2,96), ZAI(2). 

Cheilolejeunea pocsii E.W.Jones KEN(21,731,759), MDG(1), MLW(563), RWA(2), SAF(2), 

TAN(2), UGA(115). 

Cheilolejeunea principensis Steph. ex Paris 1888, nom. inval. = Cheilolejeunea serpentina (Mitt.) Mizut. 

1963 (574).


Cheilolejeunea principensis Steph. ex Paris 1906 = Cheilolejeunea serpentina (Mitt.) Mizut. 1963 (574). 

Cheilolejeunea robillardii (Steph.) R.M.Schust. 1980 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 

(849). 

Cheilolejeunea rufescens (Lindenb.) Grolle SAF(2). A little-studied species from Argentina. Mitten's specimen 

from South Africa should be re-examined. 

Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust. UGA(2), ZAI(2). 

Cheilolejeunea serpentina (Mitt.) Mizut. ANN(2,33), BIO(2,33), CMN(2), COM(1,4), GHA(2), 

GUI(2), IVO(2), KEN(48), MAU(1,108), MDG(1,4,720), NGA(2), PRI(2), REU(1), RIM(10), 

SEY(1,4,13,108), SIE(2), TAN(2), ZAI(2). 

Cheilolejeunea silvestris (Gottsche) E.W.Jones 1974 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 

(872). 

Cheilolejeunea silvestris var. involuta (Steph.) E.W.Jones 1974 = Cheilolejeunea surrepens (Mitt.) 

E.W.Jones 1976 (872). 

Cheilolejeunea surrepens (Mitt.) E.W.Jones ANN(2,33), CMN(2), COM(1,4), GHA(2), IVO(2,53), 

KEN(2,731), MAU(1), MDG(1,720), MLW(563), NGA(2), REU(1), ROD(1,723), SEY(1,4), 

SIE(2), TAN(2,736), UGA(2,42,96), ZAI(2). 

Cheilolejeunea tisserantii Jovet-Ast et Vanden Berghen 1951 = Cheilolejeunea decursiva (Sande Lac.) 

R.M.Schust. 1963 (828). 

Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust. ZAI(2). 

Cheilolejeunea trifaria (Reinw. et al.) Mizut. BIO(33), CAF(2), CMN(2), GAB(2), GHA(2), 

MAU(1), MDG(1,720), REU(1), ROD(723), RWA(2), SAO(2), SEY(1,4,108), SIE(2), TAN(2), 

TOG(48), UGA(2,96), ZAI(2), ZIM(2). 

Cheilolejeunea usambarana (Steph.) Grolle CMN(2), CON(2), GUI(2), KEN (731), MLW(563), 

REU(1), SIE(2), TAN(2). 

Cheilolejeunea ventricosa (Schiffn.) X.-L.He MAU(108). 

Cheilolejeunea sp. (Strepsilejeunea vatovae Gerola) ETH(2). 

Chiloscyphus ankefinensis Gottsche ex Steph. 1907 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Chiloscyphus angustistipulus (Steph.) J.J.Engel et R.M.Schust. 1984 = Lophocolea angustistipulus Steph. 

1922 (447) 

Chiloscyphus argutus (Reinw. et al.) Nees 1845 = Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910 (1) 


Chiloscyphus armatistipulus Steph. 1893 = Chiloscyphus dubius Gottsche 1882 (165) = Heteroscyphus dubius 

(Gottsche) Schiffn. 1910 (1). 

Chiloscyphus concretus (Mont.) J.J.Engel et R.M.Schust. 1985 = Lophocolea concreta Mont. 1845 (1). 

Chiloscyphus congoanus (Steph.) J.J.Engel et R.M.Schust. 1985 = Lophocolea martiana Nees 1845 (974). 

Chiloscyphus connatus (Sw.) J.J.Engel et R.M.Schust. 1984 = Lophocolea connata (Sw.) Mont. et Nees 

1839 (1) [Excludenda(1)]. 

Chiloscyphus cuspidatus (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea bidentata (L.) Dumort. 1835 

(975). 

Chiloscyphus decurrens Nees 1845 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984 (623). 

Chiloscyphus difformis (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea difformis Nees 1845 (1). 

Chiloscyphus dubius Gottsche 1882 = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (1). 

Chiloscyphus expansus (Lehm.) Nees 1845 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Chiloscyphus fasciculatus (Nees) Gottsche et al. 1845 = Clasmatocolea fasciculata (Nees) Grolle 1960 (314, 

97). 

Chiloscyphus fragrans (Moris et De Not.) J.J.Engel et R.M.Schust. 1985 = Lophocolea fragrans (Moris et 

De Not.) Gottsche et al. 1845 (1). 

Chiloscyphus grandistipus Steph. 1890 = Heteroscyphus grandistipus (Steph.) Schiffn. 1910 (1). 

Chiloscyphus hamatistipulus Steph. 1907 = Chiloscyphus dubius Gottsche 1882 (165) = Heteroscyphus dubius 

(Gottsche) Schiffn. 1910 (1). 

Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust. 1984 = Lophocolea lepervanchei Steph. 1907 

(1). 

Chiloscyphus lindenbergianus ("lindenbergii") Nees 1845 = Lophocolea semiteres (Lehm.) Mitt. 1877 

(313). 

Chiloscyphus loangensis Steph. 1893 probably = Chiloscyphus dubius Gottsche 1882 (165) = Heteroscyphus 

dubius (Gottsche) Schiffn. 1910 (1). 

Chiloscyphus lucidus (Spreng. ex Lehm.) J.J.Engel et R.M.Schust. 1985 [non Mitt. 1876] = Lophocolea 

lucida (Spreng. ex Lehm.) Mont. 1839 (313). 

Chiloscyphus madagascariensis (Gottsche) J.J.Engel et R.M.Schust. 1984 = Lophocolea madagascariensis 

Gottsche 1882 (1). 

Chiloscyphus martianus (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea martiana Nees 1845 (1).


Chiloscyphus mascarenensis S.W.Arnell 1965 = Chiloscyphus decurrens (Reinw. et al.) Nees 1845 (380) = 

Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984 (623). 

Chiloscyphus minor (Nees) J.J.Engel et R.M.Schust. 1984 = Lophocolea minor Nees 1836 (1). 

Chiloscyphus mollis (Nees) J.J.Engel et R.M.Schust. 1984 = Lophocolea mollis (Nees) Gottsche et al. 1845 

(1). 

Chiloscyphus muhavurensis S.W.Arnell 1956 = Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs 

1979 (950). 

Chiloscyphus muricatus (Lehm.) J.J.Engel et R.M.Schust. 1985 = Lophocolea muricata (Lehm.) Nees 1845 

(1). 

Chiloscyphus oblongifolius Mitt. 1860, hom. illeg. [non (Hook.f. et Taylor) Gottsche et al. 1847] = Chiloscyphus 

dubius Gottsche 1882 (884) = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (1). 

Chiloscyphus occidentalis (Steph.) J.J.Engel et R.M.Schust. 1985 = Chiloscyphus dubius Gottsche 1882 

(165) = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (1). 

Chiloscyphus rabenhorstii Steph. 1906 = Lophocolea fasciculata (Nees ) S.W.Arnell et Grolle in Grolle 

1959 (313) = Clasmatocolea fasciculata (Nees) Grolle 1960 (97). 

Chiloscyphus regularis Steph. 1893 [non Steph. 1906] = Lophocolea onraedtii Grolle 1984 (927). 

Chiloscyphus renauldii Steph. 1907 = Heteroscyphus cf. dubius (Gottsche) Schiffn. 1910 (1). 

Chiloscyphus semiteres (Lehm.) Lindenb. 1845 = Lophocolea semiteres (Lehm.) Mitt. 1877 (313). 

Chiloscyphus silvestris (Gottsche) J.J.Engel et R.M.Schust. 1984 = Lophocolea silvestris Gottsche 1882 

[Dubia (1)]. 

Chiloscyphus spectabilis Steph. 1891 = Heteroscyphus spectabilis (Steph.) Schiffn. 1910 (623). 

Chiloscyphus thomeensis Steph. 1893 perhaps = Heteroscyphus spectabilis (Steph.) Schiffn. 1910 (163). 

Chiloscyphus trapezioides Sande Lac. 1855 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898 (1). 

Chonecolea ruwenzorensis E.W.Jones UGA(2,50,96). 

Ciliolejeunea capensis S.W.Arnell 1953 = Lejeunea arnelliana R.M.Schust. 1965 (869) = Lejeunea villaumei 

(Steph.) Grolle 1977 (828). 

Cincinnulus bidentulus (F.Weber) Steph. 1895 = Calypogeia bidentula (F.Weber) Nees 1845 (253). 

Cladolejeunea aberrans (Steph.) Zwickel TAN(2). 

Clasmatocolea fasciculata (Nees) Grolle SAF(2,97). 

Clasmatocolea vermicularis (Lehm.) Grolle BUR(2,97), CMN(2), ETH(2), LES(2,75), MDG(97), 

MLW(2,29), NGA(2), REU(1,97), RWA(2), SAF(2,97), TAN(2,97), UGA(115), ZAI(2,97), 

ZIM(2,97). 

Clevea pulcherrima Steph. 1895 = Athalamia pulcherrima (Steph.) S.Hatt. 1954 (829). 

Clevea spathysii (Lindenb.) Müll.Frib. 1940 = Athalamia spathysii (Lindenb.) S.Hatt. 1954 (829). 

Cololejeunea adhaesiva (Mitt.) R.M.Schust. TAN(2). 

Cololejeunea adnata Tixier MDG(1). 

Cololejeunea africana (Steph.) R.M.Schust. CMN(2), KEN(2,731), SAO(2), SIE(2), TAN(2). 

Cololejeunea amaniensis Pócs TAN(2). 

Cololejeunea andapania Tixier MDG(1). 

Cololejeunea androphylla E.W.Jones CMN(2), CON(2), GHA(2), IVO(2), NGA(2), RIM(10), 

SEY(1), SIE(2). 

Cololejeunea androphylla var. madecassa Tixier MDG(1). 

Cololejeunea ankaiana Tixier MDG(1). 

Cololejeunea apiculata (E.W.Jones) R.M.Schust. CMN(2), TAN(2), UGA(50). 

Cololejeunea appressa (A.Evans) Benedix COM(1), IVO(2), KEN(2), MDG(1,720), MLW(563), 

REU(1), SEY(1), SIE(2), TAN(2), UGA(2), ZAI(2). 

Cololejeunea attilana Pócs TAN(736). 

Cololejeunea augieri Tixier RWA(2). 

Cololejeunea auriculata (E.W.Jones) R.M.Schust. CMN(2), KEN(2), MDG(1,720), NGA(2), 

SEY(1), UGA(42). 

Cololejeunea autoica (Steph.) Grolle COM(1). 

Cololejeunea bandamiae Tixier COM(1). 

Cololejeunea bebourensis Tixier COM(1), REU(1). 

Cololejeunea bichiana Tixier 1974 = C. platyneura (Spruce) A.Evans 1902 (914). 

Cololejeunea bidentula (Steph.) E.W.Jones KEN(2,731), MAU(1), MDG(1), REU(1), TAN(2), 

UGA(42). 

Cololejeunea bolombensis (Steph.) Vanden Berghen 1972 = Cololejeunea tonkinensis Steph. 1895 (135). 

Cololejeunea borbonica Tixier MDG(1). 

Cololejeunea borhidiana Pócs TAN(2).


Cololejeunea bosseriana Tixier REU(1). 

Cololejeunea calcarata E.W.Jones GHA(2). 

Cololejeunea camusii Tixier MDG(1). 

Cololejeunea capensis S.W.Arnell 1953 = Aphanolejeunea capensis (S.W.Arnell) S.W.Arnell 1957 (143). 

Cololejeunea capuronii Tixier MDG(1). 

Cololejeunea cardiocarpa (Mont.) A.Evans ASC(2), BUR(2), CMN(2), KEN(2,665,731), MDG(1), 

MLW(563), SAF(2), SIE(2), SWZ(2), TAN(2,736), UGA(2), ZAI(2), ZIM(2). 

Cololejeunea cardiocarpioides Tixier RWA(2). 

Cololejeunea ceatocarpa (Ångstr.) Steph. REU(1). 

Cololejeunea chuahiana Pócs KEN(738). 

Cololejeunea cornuta E.W.Jones GHA(2), NGA(2), RIM(10), SIE(2). 

Cololejeunea crenatiflora Steph. 1891, nom. inval. perhaps = Cololejeunea obliqua (Nees et Mont.) Schiffn. 

1897 (1). But synonymy uncertain. Jones (858) considered C. crenatiflora and C. obliqua to be closely related but 

probably not conspecific, and Tixier (914) noted that C. crenatiflora "diffère de C. obliqua par de petits détails". 

Grolle (1) accepted the synonymy with hesitation. 

Cololejeunea crenatiflora (Steph.) Vanden Berghen 1972 perhaps = Cololejeunea obliqua (Nees et Mont.) 

Schiffn. 1897 (1). See above. 

Cololejeunea crenatiflora var. orientalis Tixier 1985 perhaps = Cololejeunea obliqua (Nees et Mont.) 

Schiffn. 1897 (1). See above. 

Cololejeunea cristata (Steph.) R.M.Schust. KEN(2,731), MAU(1,108), MDG(1), ZAI(2). 

Cololejeunea cuneata (Lehm. et Lindenb.) Herzog COM(1), MAU(1), MDG(1,4), REU(1), ROD(1), 

TAN(2). 

Cololejeunea cuneifolia Steph. CMN(2), GHA(2), GUI(2), IVO(2), NGA(2), RIM(10), SIE(2), 

TAN(2), UGA(2,42,50), ZAI(2). 

Cololejeunea cuspidata Steph. ex Tixier SEY(4). 

Cololejeunea decemplicata (Steph.) Tixier MDG(1), REU(1). 

Cololejeunea dentata (E.W.Jones) R.M.Schust. CMN(2), GHA(2), KEN(2), SIE(2), TAN(2), 

UGA(50), ZAI(2). 

Cololejeunea deroinii Tixier MDG(1). 

Cololejeunea deslooveri Vanden Berghen ZAI(2). 

Cololejeunea diplasiolejeuneoides Tixier MDG(1). 

Cololejeunea disciflora Tixier MDG(1). 

Cololejeunea dissita E.W.Jones 1954 = Cololejeunea minutissima subsp. myriocarpa (Nees et Mont.) 

R.M.Schust. 1955 (25). 

Cololejeunea distalopapillata (E.W.Jones) R.M.Schust. COM(1), KEN(2,731), MDG(1,720), 

MLW(563), RWA(2), TAN(2), UGA(42,50), ZAI(2), ZIM(2). 

Cololejeunea dolichodonta Tixier MDG(1). 

Cololejeunea duvignaudii E.W.Jones COM(1), GAB(2), KEN(2,731), MDG(1), MLW(563), 

REU(1), RIM(10), RWA(2), TAN(2), UGA(115), ZAI(2), ZIM(2). 

Cololejeunea duvignaudii var. papillata Tixier MDG(1,720), REU(1). 

Cololejeunea elegans Steph. CMN(2), COM(1), GHA(2), KEN(2), MDG(1,720), MLW(563), 

RIM(10), TAN(2). 

Cololejeunea erectifolia Steph. 1895 = Leptocolea subciliata (Mitt.) E.W.Jones 1953 (858) = Cololejeunea 

cuneata (Lehm. et Lindenb.) Herzog 1947 (152). 

Cololejeunea fadenii Pócs 1975 = Aphanolejeunea fadenii (Pócs) Pócs 1984 (383). 

Cololejeunea falcata var. madecassa Tixier MDG(1). 

Cololejeunea filicaulis Steph. 1892 = Cololejeunea hildebrandii (Austin) Steph. 1897 (135). 

Cololejeunea fischeri Tixier RWA(2), ZAI(2). 

Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn. IVO(2), MDG(1), SIE(2). 

Cololejeunea floccosa var. trivittata Tixier MDG(1). 

Cololejeunea frahmii Tixier RWA(2). 

Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust. MDG(1), NGA(2), SEY(4), 

TAN(2,736). 

Cololejeunea grolleana Pócs TAN(2). 

Cololejeunea grossidens Steph. 1900, nom. nud. = Cololejeunea malanjae Steph. 1902 (842). 

Cololejeunea grossidens (Steph.) E.W.Jones 1953 = Cololejeunea malanjae Steph. 1902 (842). 

Cololejeunea guehoi Tixier MAU(1). 

Cololejeunea harrisii Pócs BUR(2), TAN(2).


Cololejeunea harrisii sensu Vanden Berghen 1977 = Cololejeunea magna (Tixier) Infante et Heras 1999 

(39). 

Cololejeunea harrisii var. magna Tixier 1995 = Cololejeunea magna (Tixier) Infante et Heras 1999 (39). 

Cololejeunea haskarliana (Lehm. et Lindenb.) Schiffn. REU(1), SEY(1). 

Cololejeunea heterolobula Tixier RWA(2), ZAI(2). 

Cololejeunea hildebrandii (Austin) Steph. CMN(2), COM(1), ETH(2), KEN(2,731), MDG(1), 

REU(1), RWA(2), TAN(2), ZAI(2). 

Cololejeunea himalayensis (Pandé et Misra) R.M.Schust. 1963 = Cololejeunea latilobula (Herzog) Tixier 

1985 (135). 

Cololejeunea hyalinomarginata (Nees ex Mont.) Grolle 1981 = Cololejeunea marginata (Lehm. et Lindenb.) 

Schiffn. 1898 (1). 

Cololejeunea inflectens (Mitt.) Benedix COM(1), SEY(1,4,108). 

Cololejeunea iradieri Infante et Heras RIM(39). 

Cololejeunea jonesii Pócs TAN(2). 

Cololejeunea kahuziensis Tixier ZAI(2). 

Cololejeunea lanceolata E.W.Jones CMN(2), GHA(2), NGA(2), SIE(2). 

Cololejeunea latilobula (Herzog) Tixier BUR(2), COM(4), MLW(2,563), NGA(2), RIM(10), 

SEY(1), TAN(2), UGA(2,42), ZAI(2), ZAM(2). 

Cololejeunea leloutrei (E.W.Jones) R.M.Schust. CMN(2), COM(1), GAB(2), GHA(25), 

KEN(2,731), MAU(1), MDG(1,720), RIM(10), SAO(2), SEY(1,4,108), SIE(25), TAN(2,108), 

ZAI(2). Current opinion favours the view that C. marginata is endemic to the East African islands. Pending a clarification 

of this difficult complex, records from Ghana and Sierra Leone, uncertainly assigned to C. marginata in (2), have been 

re-assigned in this checklist to C. leloutrei. The record from Tanzania is also transferred. 

Cololejeunea leloutrei var. microlobulata Tixier MDG(1). 

Cololejeunea leloutrei var. ulugurica Pócs in Tixier CMN(2), COM(1), KEN(2), MDG(1), TAN(2). 

Cololejeunea lemuriana Tixier MDG(1), TAN(108,129). 

Cololejeunea leonardii Vanden Berghen 1971 = Cololejeunea runssorensis (Steph.) Pócs 1975 (378). 

Cololejeunea lobulilineata Tixier ZAI(2). 

Cololejeunea magillii Pócs COM(1). 

Cololejeunea magna (Tixier) Infante et Heras RWA(2). 

Cololejeunea malanjae Steph. BUR(2), KEN(2,731), MLW(2,563), RWA(2), TAN(2), UGA(115), 

ZAI(2), ZIM(2). 

Cololejeunea marginata (Lehm. et Lindenb.) Schiffn. COM(1), MAU(1,4,108), MDG(1,720), 

REU(1), SEY(4,108). See note under C. leloutrei. 

Cololejeunea mariana Tixier 1979 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (135). 

Cololejeunea microscopica (Taylor) Schiffn. 1985 = Aphanolejeunea microscopica (Taylor) A.Evans 1911 

(384) 

Cololejeunea minutissima (Sm.) Schiffn. CVI(2,37), IVO(53), KEN(759), MDG(1), MLW(2,563), 

REU(1), ROD(1), RWA(2), SAF(2,75), TAN(2), UGA(42,50), ZAI(2), ZIM(2). 

Cololejeunea minutissima (Sm.) Schiffn. subsp. minutissima BEN(22), UGA(115). 

Cololejeunea minutissima subsp. myriocarpa (Nees et Mont.) R.M.Schust. GHA(2), LES(75), 

MLW(2,563), NGA(2), SAF(2), TAN(2), UGA(42,115). 

Cololejeunea minutissima subsp. utriculifera Vanden Berghen BUR(2), RWA(2), TAN(2). 

Cololejeunea minutissima subsp. myriantha (Herzog) R.M.Schust. 1955 = Cololejeunea minutissima subsp. 

myriocarpa (Nees et Mont.) R.M.Schust. 1955 (936). 

Cololejeunea mocambiquensis S.W.Arnell MDG(1), MLW(563), MOZ(2), SAF(2), TAN(2), 

UGA(2). 

Cololejeunea moramangae Tixier 1979 = Aphanolejeunea moramangae (Tixier) Pócs 1984 (384). 

Cololejeunea myriantha (Herzog) S.W.Arnell 1953 = Cololejeunea minutissima subsp. myriocarpa (Nees et 

Mont.) R.M.Schust. 1955 (936). 

Cololejeunea myriocarpa (Nees et Mont.) C.Massal. 1906 = Cololejeunea minutissima subsp. myriocarpa 

(Nees et Mont.) R.M.Schust. 1955 (936). 

Cololejeunea nigerica (E.W.Jones) R.M.Schust. CMN(2), GHA(2), IVO(2), NGA(2), SIE(2), 

TAN(736), TOG(2,101), UGA(42,115), ZAI(2). 

Cololejeunea nymanii (Steph.) Benedix 1953 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (135). 

Cololejeunea obliqua (Nees et Mont.) Schiffn. BUR(2), CMN(2), COM(1), GHA(2), GUI(2), 

IVO(2), MAU(1), MDG(1,720), NGA(2), REU(1), RIM(10), SAO(2), SEY(1,4,108), SIE(2), 

TAN(2), UGA(2,42,115), ZAI(2), ZAM(2). 

Cololejeunea obtusifolia (E.W.Jones) Tixier BEN(22), BUR(2), CMN(2), ETH(2), GHA(2),


GNB(2), GUI(2), IVO(2), KEN(2), MDG(1,720), MLW(563), NGA(2), RIM(10), RWA(2), SIE(2), 

TAN(2), UGA(42,50), ZAI(2). 

Cololejeunea occidentalis (E.W.Jones) Vanden Berghen KEN(2), MLW(563), NGA(2), SAF(2), 

TAN(2), ZIM(2). 

Cololejeunea oleana Sim SAF(2). 

Cololejeunea onraedtii Tixier COM(1), ROD(723), SEY(1). 

Cololejeunea parva Vanden Berghen RWA(2), UGA(115). 

Cololejeunea peculiaris (Herzog) Benedix 1953 = Cololejeunea inflectens (Mitt.) Benedix 1953 (878). 

Cololejeunea peponiformis Mizut. COM(1), REU(1), SEY(1). 

Cololejeunea peraffinis var. elegans Benedix GUI(2,106), NGA(2,104,106), SIE(2,25). The plants 

from Guinea, Nigeria and Sierra Leone were published by E.W. Jones in (25, 104, 106) as C. peraffinis var. elegans. However, 

he later acknowledged (in 1037) that they showed conflicting characters, approaching C. peraffinis var. elegans in 

some, and matching C. usambarica (= C. platyneura) in others. (see also Pócs, in Pócs & Streimann 1999 (103)). 

Cololejeunea peraffinis var. serrulata Benedix MDG(720). 

Cololejeunea plagiochiliana Tixier MDG(1). 

Cololejeunea planissima (Mitt.) Abeyw. TAN(702). 

Cololejeunea platyneura (Spruce) A.Evans MDG(1,720), MLW(563), RWA(2), TAN(2), ZAI(2). 

Cololejeunea praeruptorum Tixier MDG(1). 

Cololejeunea pseudo-obliqua Tixier RWA(2), ZAI(2). 

Cololejeunea pseudopusilla Tixier ZAI(2). 

Cololejeunea pteroporum Tixier MDG(1). 

Cololejeunea punctata (E.W.Jones) R.M.Schust. 1963, nom. illeg. = Cololejeunea vidaliana Tixier 1973 

(847). 

Cololejeunea punctata (Gottsche) Pearson 1892 MAD(1). [Dubia(1)]. 

Cololejeunea pusilla Steph. CMN(2), GAB(2), GHA(2), GUI(2), NGA(2), RIM(10), RWA(2), 

TAN(108), TOG(101). 

Cololejeunea pusilla var. obtusifolia E.W.Jones 1954 = Cololejeunea obtusifolia (E.W.Jones) Tixier 1995 

(130). 

Cololejeunea raduliloba Steph. COM(1), MDG(1), SEY(1). 

Cololejeunea runssorensis (Steph.) Pócs BUR(2), KEN(2), MLW(563), RWA(2), UGA(2,115), 

ZAI(2). 

Cololejeunea saltuum Tixier MDG(1). 

Cololejeunea sambiroana Tixier MDG(1). 

Cololejeunea scabriflora Gottsche ex Steph. 1895 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 

(135). 

Cololejeunea scabrifolia Schiffn. 1900, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 

(135). 

Cololejeunea selangorensis Tixier MDG(1). 

Cololejeunea sphaerocarpa Tixier RWA(2), ZAI(2). 

Cololejeunea stenophylla Herzog 1947 = Cololejeunea hildebrandii (Austin) Steph. 1897 (152, 913). 

Cololejeunea subciliata (Mitt.) R.M.Schust. 1963 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 

(152). 

Cololejeunea subdiaphana Jovet-Ast 1947 = Aphanolejeunea truncatifolia Horik. 1934 (117). 

Cololejeunea subinflata Tixier MDG(1). 

Cololejeunea succinea Tixier MDG(152). 

Cololejeunea takamakae Tixier REU(1). 

Cololejeunea tamatavensis Tixier MDG(1). 

Cololejeunea tanneri Pócs MLW(563), TAN(2). 

Cololejeunea tanzaniae Pócs COM(1), KEN(731), MAU(108), MDG(1,720), MLW(563), TAN(2). 

Cololejeunea tenella Benedix RWA(2). 

Cololejeunea tenuiparietata Tixier RWA(2), ZAI(2). 

Cololejeunea tonkinensis Steph. BUR(2), CMN(2), COM(1), GUI(2), IVO(2), KEN(2,731), 

MDG(1,720), MLW(563), NGA(2), RIM(10), SIE(2), TAN(2), UGA(2,42), ZAI(2), ZIM(2). 

C. tonkinensis and C. bolombensis are treated as synonyms of C. lanciloba Steph. in Zhu & So, 2000 (702). 

Cololejeunea tribracteata Tixier ZAI(2). 

Cololejeunea usambarica E.W.Jones 1954 = Cololejeunea platyneura (Spruce) A.Evans 1902 (914, 915). 

Cololejeunea vidaliana Tixier KEN(2), MAU(1), MDG(1), ROD(723), SEY(1,4,108), TAN(2). 

Cololejeunea xavieri (Lacout. ex Steph.) E.W.Jones COM(1), KEN(2). Tixier 1979 (152) made C. xavieri a 

synonym of C. elegans, but it is considered probably distinct by others, and is kept separate here.


Cololejeunea zenkeri (Steph.) E.W.Jones BUR(2), CMN(2), MDG(720), MLW(563), PRI(48), 

RIM(10), RWA(2), SEY(4,108), SIE(2), TAN(2), UGA(50), ZAI(2). 

Colura alata (Gottsche) Trevis. 1877 = Lejeunea alata Gottsche 1845 (828). 

Colura benoistii Jovet-Ast COM(1), MDG(1). 

Colura berghenii Jovet-Ast RWA(2), TAN(2). 

Colura bicornis Jovet-Ast MDG(1,4). 

Colura calyptrifolia (Hook.) Dumort. MLW(110), RWA(2), SAF(2,75), TAN(2), UGA(42,50). 

Colura calyptrifolia subsp. tenuicornis (A.Evans) Vanden Berghen 1972 = Colura tenuicornis (A.Evans) 

Steph. 1916 (25). 

Colura cylindrica Herzog TAN(2). 

Colura digitalis (Mitt.) Steph. ANN(2,33), BUR(2), CAF(2), CMN(2), COM(1,4), CON(2), GAB(2), 

GHA(2), GUI(2), IVO(2), KEN(731), MAU(1,108), MDG(4,720), MLW(110), NGA(2), PRI(2), 

REU(1), RIM(10), RWA(2), SAO(2), SEY(1,108), SIE(2), TAN(2), UGA(2,42), ZAI(2). 

Colura digitalis var. mucronata Pócs SEY(4). 

Colura dusenii (Steph.) Steph. CMN(2), GHA(2), KEN(2), NGA(2), SIE(2), TAN(2). 

Colura hattoriana Pócs COM(1), MLW(110). 

Colura hedbergiana Pócs TAN(2), UGA(42). 

Colura heimii Jovet-Ast COM(1), MAU(1,108), MDG(1), REU(1). 

Colura humbertii Jovet-Ast MAU(1), MDG(1), REU(1). 

Colura inornata Jovet-Ast MDG(1,4). 

Colura kilimanjarica Pócs et Jovet-Ast KEN(759), TAN(2), UGA(42). 

Colura mauritiana Pócs MAU(6,108). 

Colura obesa Jovet-Ast MDG(1), REU(1). 

Colura obtusa (Steph.) Steph. 1916 = Colura digitalis (Mitt.) Steph. 1916 (336, 836). 

Colura obvoluta Jovet-Ast MAU(108), MDG(1), SEY(4,108). 

Colura pluridentata Jovet-Ast SEY(1). 

Colura pungens Herzog 1947 = Colura tenuicornis (A.Evans) Steph. 1916 (336). 

Colura saroltae Pócs RWA(2), TAN(2). 

Colura superba (Mont.) Steph. 1916 REU(1). [Excludenda(1)]. 

Colura tenuicornis (A.Evans) Steph. ASC(2), BUR(2), CMN(2), COM(1), GAB(2), KEN(2), 

MAU(1), MDG(1), MLW(110), REU(1), RIM(10), RWA(2), SAF(2), SAO(2), SEY(1,4), SIE(2), 

TAN(2), UGA(42,50), ZAI(2), ZIM(733). 

Colura usambarica E.W.Jones KEN(731), MLW(2,110), TAN(2). 

Colura zoophaga Eb.Fisch. KEN(52). Chuah-Petiot & Pócs (759) considered that C. zoophaga "seems to us to be a 

synonym of C. kilimanjarica…", but they did not formally place it in synonymy. 

Colurolejeunea dusenii Steph. 1892 = Colura dusenii (Steph.) Steph. 1916 (336). 

Colurolejeunea obtusa Steph. 1891 = Colura obtusa (Steph.) Steph. 1916 (979) = Colura digitalis (Mitt.) 

Steph. 1916 (336). 

Colurolejeunea tenuicornis A.Evans 1900 = Colura tenuicornis (A.Evans) Steph. 1916 (336). 

Conoscyphus inflexifolius Mitt. 1873 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898 (884). 

Conoscyphus trapezioides (Sande Lac.) Schiffn. MDG(1,4,720), MLW(29), REU(1,4), SAO(2), 

SEY(1,4), SIE(2), TAN(2). Erroneously listed from Cameroon in (2). 

Crossotolejeunea kilimandjarica S.W.Arnell 1960 = Taxilejeunea pulchriflora Pearson 1924 (872). 

Cryptochila grandiflora (Lindenb. et Gottsche) Grolle MDG(1), MLW(114), REU(1,4), SAF(2). 

Cryptomitrium oreades Perold LES(2,75). 

Cuspidatula contracta (Reinw. et al.) Steph. 1901 = Jamesoniella contracta (Reinw. et al.) N.Kitag. 1981 

(527). 

Cyathodium africanum Mitt. ANN(2,33), BIO(2,33,36), CAF(2), CMN(2), COM(1), CON(2), 

CVI(2,20,37), GHA(2), GUI(2), IVO(2), MDG(720), MLW(2), NAM(2), NGA(2), SAO(2), SIE(2), 

TAN(2), TOG(2), UGA(42,50,96), ZAI(2), ZIM(2). In edition 1, Srivastava & Dixit (119) was followed in 

placing C. africanum in synonymy with C. cavernarum. However, since its status seems as yet unclear, it is reinstated in 

this edition. 

Cyathodium aureonitens (Griff.) Mitt., non Schiffn. CMN(2). 

Cyathodium foetidissimum Schiffn. CMN(2), UGA(96). The Uganda plants are sterile, and Lye & Pócs (96) 

considered their identity to be uncertain. 

Cyathodium griffithii Schiffn. 1938, nom. illeg. = Cyathodium aureonitens (Griff.) Mitt. 1886 (650). 

Cyathodium smaragdinum Schiffn. 1910 COM(1). [Excludenda(1) et Dubia(2)]. Srivastava & Dixit 1996 (119) 

cited a collection from Tanzania (Jones 679) under C. smaragdinum, but cited the same specimen also under C. cavernarum. 

Clarification is required.


Cyclolejeunea annobonensis S.W.Arnell 1956 = Prionolejeunea kindbergii Steph. 1892 (815) = Prionolejeunea 

grata (Gottsche) Schiffn. 1893 (847). 

Cylindrocolea atroviridis (Sim) Váňa CAF(2), CMN(2), GHA(2), IVO(51), LES(75), MDG(1), 

MOZ(2), NGA(2), RWA(2), SAF(2), SIE(2), UGA(50), ZAI(2), ZIM(2). 

Cylindrocolea chevalieri (Steph.) R.M.Schust. CON(2), GAB(2), GHA(2), NGA(2). 

Cylindrocolea gittinsii (E.W.Jones) R.M.Schust. CMN(2), KEN(21), RWA(2), UGA(2,776). 

Cylindrocolea madagascariensis (Steph.) R.M.Schust. MDG(1). 

Cylindrocolea nigerica (E.W.Jones) R.M.Schust. DJI(2), NGA(2). 

Cylindrocolea ugandica (E.W.Jones) R.M.Schust. KEN(21), UGA(2,42). 

Cylindrocolea cf. ugandica SIE(25). 

Cystolejeunea lineata (Lindenb. et Lehm.) A.Evans 1906 REU(1). [Excludenda(1)]. 

Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche et al. STH(2). 

Dendroceros africanus Steph. CMN(2), TAN(2). 

Dendroceros borbonicus Steph. MAU(1), REU(1), SEY(1,108). 

Dendroceros crispatus Nees SAO(95). 

Dendroceros javanicus (Nees) Gottsche et al. TAN(2). 

Dendroceros lacerus Nees 1846 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975 (1). 

Dendrolejeunea fruticosa (Lindenb. et Gottsche) Lacout. 1908 MAU(1). [Excludenda(1)]. 

Denotarisia linguifolia (De Not.) Grolle MDG(720). 

Dicranolejeunea africana Steph. 1912 = Caudalejeunea sp. [non C. africana (Steph.) Schiffn. 1893]. (1). 

Dicranolejeunea annobonensis S.W.Arnell 1956 = Marchesinia sp. (542). 

Dicranolejeunea atra (Mitt.) Vanden Berghen 1953 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 

1988 (542). 

Dicranolejeunea bovonei Gola 1920 ZAI(2). [Dubia(542)]. 

Dicranolejeunea chrysophylla (Lehm.) Grolle 1966 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 

(542). 

Dicranolejeunea lillieana Pearson 1924 = Acanthocoleus sp. (122). 

Dicranolejeunea madagascariensis Steph. 1912 = Acanthocoleus madagascariensis (Steph.) Kruijt 1988 

(542). 

Dicranolejeunea madagascariensis var. obtusifolia E.W.Jones 1970 = Acanthocoleus chrysophyllus (Lehm.) 

Kruijt 1988 (542). 

Dicranolejeunea natalensis (Sim) S.W.Arnell 1961 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 

(542). 

Dicranolejeunea phyllorhiza (Nees) Schiffn. 1893 = Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. 

1986 (133). 

Dicranolejeunea usambarensis Steph. 1912 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988 

(542). 

Diplasiolejeunea albifolia (Taylor) E.W.Jones KEN(2,731), NGA(2), TAN(2). 

Diplasiolejeunea albifolia subsp. orientalis Pócs, ined. TAN(129). 

Diplasiolejeunea aulae E.W.Jones GHA(2), MDG(1), RWA(2), UGA(50). 

Diplasiolejeunea auriculata Tixier MDG(1). 

Diplasiolejeunea bifidula Steph. 1916 = Diplasiolejeunea cornuta Steph. 1916 (1). 

Diplasiolejeunea brachyclada A.Evans 1912 = Diplasiolejeunea cavifolia Steph. 1895 (900, 940). 

Diplasiolejeunea cavifolia Steph. COM(1), GHA(2), KEN(2), MAU(1,108), MDG(1,720), NGA(2), 

RIM(10), RWA(2), SAF(2), SAO(2), TAN(2), UGA(42), ZAI(2). 

Diplasiolejeunea cobrensis Gottsche ex Steph. GHA(2), MDG(1,720), SIE(2). 

Diplasiolejeunea cobrensis subsp. antsirananae Pócs MDG(709,720). 

Diplasiolejeunea cogoensis Infante, Heras et Pócs RIM(38). 

Diplasiolejeunea comorensis Tixier COM(1), MAU(108), MDG(1), ZAI(2). 

Diplasiolejeunea cornuta Steph. COM(1), GHA(2), KEN(2), MAU(1), MDG(1,4), MLW(563), 

REU(1,4), RWA(2), SEY(4), TAN(2), UGA(42,115), ZAI(2), ZIM(2). 

Diplasiolejeunea cornuta var. laciniata Herzog 1947 = Diplasiolejeunea cornuta Steph. 1916 (1). 

Diplasiolejeunea cyangugensis Tixier RWA(2). 

Diplasiolejeunea deslooveri Vanden Berghen RWA(2), TAN(2). 

Diplasiolejeunea drepanolejeunoides Tixier 1977 = Diplasiolejeunea magnistipulata Tixier 1975 (135). 

Diplasiolejeunea ensifera Tixier MDG(1). 

Diplasiolejeunea gradsteinii Tixier ZAI(2). 

Diplasiolejeunea hamata Tixier MDG(1). 

Diplasiolejeunea harpaphylla Steph. 1916 = Diplasiolejeunea cobrensis Gottsche ex Steph. 1916 (1).


Diplasiolejeunea incurvata Jovet-Ast et Tixier 1962 = Diplasiolejeunea harpaphylla Steph. 1916 (25) = 

Diplasiolejeunea cobrensis Gottsche ex Steph. 1916 (1). 

Diplasiolejeunea insignis Tixier MDG(1). 

Diplasiolejeunea integerrima Tixier 1979 MAD(1). [Dubia(1)]. 

Diplasiolejeunea jonesii Tixier MDG(1). 

Diplasiolejeunea kraussiana (Lindenb.) Steph. BUR(2), MDG(1), MLW(563), RWA(2), SAF(2), 

TAN(2), UGA(50), ZAI(2), ZIM(2). 

Diplasiolejeunea lemuriana Tixier MDG(1). 

Diplasiolejeunea magnistipulata Tixier CMN(2), MDG(1). 

Diplasiolejeunea ocellata Steph. 1916 = Diplasiolejeunea cavifolia Steph. 1895 (1). 

Diplasiolejeunea palustrium Tixier MDG(1). 

Diplasiolejeunea pellucida (Meissn. ex Spreng.) Schiffn. 1893 MAS(1), MAU(1), REU(1). [Excludenda(1)]. 

This name has sometimes been applied erroneously to African specimens of D. albifolia (Taylor) 

E.W.Jones. 

Diplasiolejeunea phyllarthonii Tixier MDG(1). 

Diplasiolejeunea ramicola Tixier MDG(1). 

Diplasiolejeunea runssorensis Steph. BUR(2), KEN(2,731), MDG(1), RWA(2), TAN(2), UGA(50), 

ZAI(2). 

Diplasiolejeunea runssorensis var. australis E.W.Jones 1974 = Diplasiolejeunea villaumei Steph. 1916 

(941). 

Diplasiolejeunea subcornuta Tixier MDG(1), MLW(563), SEY(4). 

Diplasiolejeunea symoensii Vanden Berghen BUR(2), COM(1), KEN(2), MDG(1), MLW(563), 

RWA(2), TAN(2), UGA(42,50), ZAI(2). 

Diplasiolejeunea tridentata 1979 = Diplasiolejeunea aulae E.W.Jones 1974 (1107). 

Diplasiolejeunea tubulata Grolle, nom. herb. = Diplasiolejeunea zakiae Tixier 1977 (507). 

Diplasiolejeunea utriculata Steph. MDG(1), TAN(545). 

Diplasiolejeunea vandenberghenii Grolle 1960 = Diplasiolejeunea cavifolia Steph. 1895 (900, 940). 

Diplasiolejeunea villaumei Steph. MDG(1), MLW(563), TAN(2). 

Diplasiolejeunea zakiae Tixier KEN(2,731), MDG(26,720), TAN(2). 

Diplophylleia biapiculata (Hook.f. et Taylor) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893 

(916, 163). 

Diplophylleia imbricata (Wilson) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893 (883, 163). 

Diplophylleia incumbens (Lehm. et Lindenb.) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893 

(883, 163). 

Diplophylleia piligera (Nees) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893 (163). 

Diplophyllum africanum S.W.Arnell KEN(2,71), RWA(2), TAN(2). 

Drepanolejeunea africana Steph. 1913 possibly = Drepanolejeunea capulata (Taylor) Steph. 1913 (1037). 

Drepanolejeunea angustifolia (Mitt.) Grolle 1965 COM(1), REU(1). [Excludenda(1)]. 

Drepanolejeunea ankasica E.W.Jones GHA(2). 

Drepanolejeunea cambouena Steph. COM(1), MAU(1), MDG(1), SEY(1,108), TAN(2). 

Drepanolejeunea capensis Steph. 1913 = Drepanolejeunea hamatifolia (Hook.) Schiffn. 1893 (170). 

Drepanolejeunea capulata (Taylor) Steph. NGA(2). 

Drepanolejeunea clavicornis Steph. 1913 probably = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 

(1037). In (649), “clavicornis” was erroneously transcribed as “claviformis”. 

Drepanolejeunea cristata Steph. 1891 = Drepanolejeunea cultrella (Mitt.) Steph. 1913 (649, 1013). 

Drepanolejeunea cultrella (Mitt.) Steph. BIO(72), BUR(2), CMN(2), COM(1), GUI(2), IVO(2), 

KEN(2,731), MDG(720), MLW(563), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(115), 

ZAI(2). 

Drepanolejeunea deslooveri Vanden Berghen RWA(733). Treated as a synonym of D. hamatifolia (Hook.) 

Schiffn. by Tixier 1995 (130) and accepted uncritically as such in (2) and in edition 1 of this checklist. However, no published 

supporting evidence for the synonymy has been found, and D. deslooveri is therefore reinstated pending further evidence 

coming to light. 

Drepanolejeunea filicuspis Steph. 1913 = Harpalejeunea filicuspis (Steph.) Mizut. 1973 (345). 

Drepanolejeunea friesii Vanden Berghen 1951 probably = Drepanolejeunea physaefolia (Gottsche) Steph. 

1913 (1037). 

Drepanolejeunea geisslerae Pócs MDG(709,720). 

Drepanolejeunea gomphiae Steph. 1913 = Leptolejeunea truncatiloba Steph. 1892 (649) = Leptolejeunea 

maculata (Mitt.) Schiffn. 1898 (924). 

Drepanolejeunea hamatifolia (Hook.) Schiffn. MOZ(2), RWA(2), SAF(2,75), ZAI(2), ZIM(2). 

Drepanolejeunea helenae Pócs REU(6).


Drepanolejeunea intorta (Besch. et Spruce) Steph. 1913 = Drepanolejeunea physaefolia (Gottsche) Steph. 

1913 (1). 

Drepanolejeunea madagascariensis (Steph.) Grolle COM(1), KEN(2,731), MAU(1), MDG(1,720), 

REU(1), SEY(1,4,108), TAN(2). 

Drepanolejeunea mascarena S.W.Arnell COM(4,765), MAU(1,765), MDG(765). 

Drepanolejeunea mauritiana Tixier MAU(1). 

Drepanolejeunea micholitzii Steph. 1913 = Drepanolejeunea pentadactyla (Mont.) Steph. 1913 (745). 

Drepanolejeunea molleri Steph. 1913 = Drepanolejeunea cultrella (Mitt.) Steph. 1913 (649). 

Drepanolejeunea papillosa S.W.Arnell SAF(2). 

Drepanolejeunea pentadactyla (Mont.) Steph. MDG(744). 

Drepanolejeunea physaefolia (Gottsche) Steph. BIO(2,33), CMN(2), COM(1,4), GUI(2), IVO(2), 

KEN(2,71,731,759), MAU(1), MDG(1,4,720), MLW(2,563), MOZ(2), PRI(2), REU(1), RIM(10), 

RWA(2), SAF(2), SAO(2), SEY(1), TAN(2), UGA(2), ZAI(2), ZIM(2). D. physaefolia (Gottsche) Steph. 

was treated as a synonym of D. vesiculosa by Tixier (130), and for that reason records were placed under the latter name in 

(2). However, Grolle 1995 (1) considered that synonymy had yet to be proven and listed East African Island collections 

under D. physaefolia. In this checklist, all African records are placed under D. physaefolia. 

Drepanolejeunea pocsii Grolle COM(1), KEN(2,731), MDG(720), MLW(563), TAN(2), ZIM(2). 

Drepanolejeunea riddleana Steph. 1914 = Harpalejeunea filicuspis (Steph.) Mizut. 1973 (345). 

Drepanolejeunea ruandensis Vanden Berghen RWA(2), ZAI(2). 

Drepanolejeunea securifolia (Gottsche) Steph. 1913 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 

(1). 

Drepanolejeunea symoensii Vanden Berghen et Grolle BUR(2), RWA(2), UGA(115), ZAI(2). 

Drepanolejeunea symoensii var. minor Tixier RWA(2). 

Drepanolejeunea taxiophthalma Herzog 1947 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Drepanolejeunea tenuis (Nees) Schiffn. 1898 = Drepanolejeunea angustifolia (Mitt.) Grolle 1965 (1) [Excludenda(1)]. 

Drepanolejeunea ternatensis (Gottsche) Schiffn. KEN(759), SEY(1,4,108). 

Drepanolejeunea thwaitesiana (Mitt.) Steph. 1913 MAU(1). [Excludenda(1)]. 

Drepanolejeunea trematodes (Nees) Bischl. COM(1), KEN(2,731), MAU(1,108), MDG(1,720), 

REU(1), SEY(108), TAN(2). 

Drepanolejeunea vesiculosa (Mitt.) Steph. - see note under D. physaefolia (Gottsche) Steph. 

Drepanolejeunea vesiculosa subsp. affinis Schiffn. ex Herzog 1939, nom. inval. = Drepanolejeunea vesiculosa 

(Mitt.) Steph. 1913 (1) [Excludenda(1)]. 

Drepanolejeunea sp. (D. ?friesii) GHA(25). This taxon, published in Jones & Harrington (1983) as D. ?friesii, 

was included under D. vesiculosa in (2). Since its placement is uncertain, it is listed separately here. 

Dumortiera denudata Mitt. 1861 = Wiesnerella denudata (Mitt.) Steph. 1899 (1). 

Dumortiera hirsuta (Sw.) Nees ANG(2), BIO(2,33,36), BUR(2), CMN(2), COM(1), ETH(2), 

KEN(2,71,665,731), MAU(1), MLW(2), MOZ(2), REU(1), RWA(2), SAF(2), SAO(2), SIE(2), 

SWZ(2), TAN(2), UGA(42,50,115), ZAI(2), ZIM(2). 

Dumortiera hirsuta var. nepalensis (Taylor) R.M.Schust. ZAI(2). 

Dumortiera trichocephala (Hook.) Nees 1838 = Dumortiera hirsuta var. nepalensis (Taylor) R.M.Schust. 

1992 (141). 

Eulejeunea abyssinica Gola 1914 = Lejeunea abyssinica (Gola) Cufod. 1952 (502). 

Eulejeunea aloba (Sande Lac.) Schiffn. = Lejeunea aloba Sande Lac. 1856 (969). 

Eulejeunea breutelii Steph. 1896 [= Lejeunea breutelii Steph. 1896] = Lejeunea caespitosa Lindenb. 1845 

(806). 

Eulejeunea camerunensis Steph. 1923 = Lejeunea camerunensis (Steph.) E.W.Jones 1972 (25) = Lejeunea 

tuberculosa Steph. 1915 (720). 

Eulejeunea compressistipula Pearson 1922 [= Lejeunea compressistipula Pearson 1921] perhaps = Lejeunea 

confusa E.W.Jones 1972 (157). 

Eulejeunea corbieri Steph. 1912 perhaps = Lejeunea ibadana A.J.Harr. et E.W.Jones 1982 (25). 

Eulejeunea cucullata (Reinw. et al.) Schiffn. 1898 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995 (1). 

Eulejeunea cuculliloba Steph. 1891, nom. nud. = Lejeunea caespitosa Lindenb. 1845 (157). 

Eulejeunea ecarinata Steph. Nov 1890 = Lejeunea ecarinata (Steph.) Steph. Dec 1890 (1). 

Eulejeunea ecklonii Lindenb. ex Steph. 1890, nom. inval. = Lejeunea eckloniana Lindenb. 1845 (869). 

Eulejeunea elobulata Sim 1926 = Lejeunea elobulata (Sim) S.W.Arnell 1963 (170). 

Eulejeunea expansa Steph. 1912 = Lejeunea expansa (Steph.) Steph. 1915 (982) = Lejeunea flava subsp. 

tabularis (Spreng.) S.W.Arnell 1963 (864) = Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (969). 

Eulejeunea flava (Sw.) Steph. Mar-Apr 1888 = Lejeunea flava (Sw.) Nees 1838 (1). 

Eulejeunea flavovirens (Ångstr.) Steph. 1913 = Lejeunea flavovirens Ångstr. 1873 (1).


Eulejeunea hepaticola Steph. 1888 = Lejeunea hepaticola (Steph.) Steph. 1915 (982). 

Eulejeunea isomorpha (Gottsche) Steph. 1901 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (864, 1). 

Eulejeunea jungneri Steph. 1896 = Lejeunea jungneri Steph. 1896 (982) perhaps = Taxilejeunea conformis 

(Nees et Mont.) Steph. 1914 (863). 

Eulejeunea lamacerina Steph. 1896 = Lejeunea lamacerina (Steph.) Schiffn. 1902 (971). 

Eulejeunea microclada Pearson 1931 perhaps = Lejeunea camerunensis (Steph.) E.W.Jones 1972 (25) = 

Lejeunea tuberculosa Steph. 1915 (720). 

Eulejeunea obtusata (Gottsche) Pearson 1892, nom. inval. = Lejeunea obtusata Gottsche 1882 (1). 

Eulejeunea rodriguezii Steph. Mar-Apr 1888 = Lejeunea rodriguezii (Steph.) Steph. 1891 (1). 

Eulejeunea saccatiloba Steph. Sept-Oct 1891 = Lejeunea caespitosa Lindenb. 1845 (157). 

Eulejeunea setacea Steph. 1912 = Lejeunea setacea (Steph.) Steph. 1914 (1). 

Eulejeunea squarrosa Steph. 1895 = Lejeunea squarrosa (Steph.) Steph. 1915 (982). 

Eulejeunea vulgariformis (Gottsche) Pearson 1892, nom. inval. = Lejeunea vulgariformis Gottsche 1882 (1). 

Eulejeunea wilmsii Steph. 1892 = Lejeunea eckloniana Lindenb. 1845 (869). 

Euosmolejeunea africana (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea surrepens (Mitt.) 

E.W.Jones 1976 (867, 872). 

Euosmolejeunea brachytoma (Gottsche) Steph. 1914 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 

(847, 1040). 

Euosmolejeunea contigua (Nees) Steph. 1890 = Pycnolejeunea contigua (Nees) Grolle 1979 (14). 

Euosmolejeunea grandistipula (Steph.) Steph. 1914 = Euosmolejeunea brachytoma (Gottsche) Steph. 1914 

(867) = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 (847). 

Euosmolejeunea inflata (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea intertexta (Lindenb.) Steph. 

1897 (574). 

Euosmolejeunea longiflora (Taylor) Steph. 1914 = Cheilolejeunea longiflora (Taylor) R.M.Schust. 1980 

(1040). [Dubia(2)]. 

Euosmolejeunea montagnei (Gottsche) Steph. 1895 = Cheilolejeunea montagnei (Gottsche) R.M.Schust. 

1963 (912). 

Euosmolejeunea principensis (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea serpentina (Mitt.) 

Mizut. 1963 (574). 

Euosmolejeunea robillardii Steph. 1914 = Cheilolejeunea robillardii (Steph.) R.M.Schust. 1980 (1040) = 

Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 (849). 

Euosmolejeunea ruwenzorensis S.W.Arnell 1956 = Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust. 

1963 (912). 

Euosmolejeunea serpentina (Mitt.) Steph. 1914 = Cheilolejeunea serpentina (Mitt.) Mizut. 1963 (574). 

Euosmolejeunea thomeensis Steph. 1914 = Euosmolejeunea montagnei (Gottsche) Steph. (855) = Cheilolejeunea 

montagnei (Gottsche) R.M.Schust. 1963 (912). 

Euosmolejeunea trifaria (Reinw. et al.) Steph. 1888 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 

(1042). 

Evansiolejeunea renistipula (Steph.) Vanden Berghen 1948 = E. roccatii (Gola) Vanden Berghen 1960 

(850). 

Evansiolejeunea roccatii (Gola) Vanden Berghen RWA(2), UGA(2,96), ZAI(2). 

Exormotheca africana Steph. 1924 = Exormotheca pustulosa Mitt. 1870 (401). 

Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf NAM(19). 

Exormotheca holstii Steph. NAM(2,19), SAF(2,19), TAN(2), ZIM(2,9). 

Exormotheca megastomata Marquand 1930 = Exormotheca holstii Steph. 1899 (1043). 

Exormotheca pustulosa Mitt. ANG(2), BOT(2), CHA(2), COM(1), CVI(2,37), ETH(2), KEN(2), 

LES(2,75), NAM(2), REU(1), SAF(2), SOC(746), STH(2), TAN(2), ZIM(2,9). 

Exormotheca youngii S.W.Arnell 1954 = Exormotheca holstii Steph. 1899 (170, 1043). 

Fimbraria gondarensis Gola 1914 = Asterella pappii (Gola) Grolle 1976 (999). 

Fimbraria marginata Nees 1820 = Asterella marginata (Nees) S.W.Arnell 1963 (997). 

Fimbriaria abyssinica Gottsche 1846 = Asterella abyssinica (Gottsche ) Grolle 1972 (158). 

Fimbriaria angolensis Steph. 1899 = Asterella angolensis (Steph.) S.W.Arnell 1963 = Asterella wilmsii 

(Steph.) S.W.Arnell 1963 (997). 

Fimbriaria bachmannii Steph. 1894 = Asterella bachmannii (Steph.) S.W.Arnell 1963 (997). 

Fimbriaria boryana Mont. 1838, nom. illeg. = Asterella syngenesica (Bory) Grolle 1974 (849, 1). 

Fimbriaria coronata Steph. 1917 = Asterella coronata (Steph.) H.A.Mill. 1981 (1). 

Fimbriaria dissoluta Steph. 1895 = Asterella dissoluta (Steph.) Grolle 1989 (998). 

Fimbriaria gigantea Steph. 1899 = Asterella sp. 

Fimbriaria incrassata Steph. 1899 = Asterella sp. 

Fimbriaria kamerunensis Steph. 1917 = Asterella sp. 

Fimbriaria linearis Steph. 1895 = Asterella linearis (Steph.) M.Howe 1898 (1044). 

Fimbriaria muscicola Steph. 1892 = Asterella muscicola (Steph.) S.W.Arnell 1963 (997).


Fimbriaria pappii Gola 1914 = Asterella pappii (Gola) Grolle 1976 (992). 

Fimbriaria pirottae Gola 1914 = Asterella sp. 

Fimbriaria preussii Schiffn. in Steph. 1895 = Asterella sp. 

Fimbriaria pusilla Mitt. 1888 = Asterella pappii (Gola) Grolle 1976 (999). 

Fimbriaria volkensii Steph. 1899 = Asterella sp. 

Fimbriaria wilmsii Steph. 1892 = Asterella wilmsii (Steph.) S.W.Arnell 1963 (997). 

Folioceros fuciformis Mont. ANN(2,33), REU(1), ROD(1), TAN(2). 

Folioceros incurvus (Steph.) D.C.Bharadwaj CMN(2). 

Fossombronia angulifolia Perold LES(86,91). 

Fossombronia angulosa (Dicks.) Raddi CVI(2,37). 

Fossombronia caespitiformis De Not. ex Rabenh. ETH(2). 

Fossombronia capensis S.W.Arnell SAF(2,91). 

Fossombronia capensis var. spiralis Perold SAF(87,91). 

Fossombronia cederbergensis Perold SAF(84,91). 

Fossombronia crispa Nees SAF(2,83,91), SOC(692), ZIM(2). Al-Gifri et al, 1995 (692) reported F. crispa 

from Socotra, but Kürschner, 2001 (558) considered that the record should probably be referred to R. caespitiformis. See 

also Perold 1997f (83). 

Fossombronia cristata Lindb. 1873 = Fossombronia wondraczekii (Corda) Dumort. ex Lindb. 1873 (948, 1). 

Fossombronia densilamellata S.W.Arnell SAF(2,79,91). 

Fossombronia elsieae Perold SAF(88,91). 

Fossombronia foveolata Lindb. MAU(719), REU(45), SAF(2). 

Fossombronia gemmifera Perold LES(86), SAF(91). 

Fossombronia glenii Perold LES(86), SAF(91). 

Fossombronia grandis Steph. KEN(2), TAN(2), UGA(96). 

Fossombronia hamato-hirta Steph. 1894 = Fossombronia wondraczekii (Corda) Dumort. ex Lindb. 1873 

(1045). 

Fossombronia husnotii Corb. CMN(2), ETH(2), SIE(2), TAN(2), ZAI(2), ZAM(2). 

Fossombronia hyalorhiza Perold SAF(90,91). 

Fossombronia indica Steph. CMN(2), GHA(2), NGA(2), SIE(2), TOG(2). 

Fossombronia intestinalis Taylor SAF(2). 

Fossombronia leucoxantha (Lehm.) Gottsche et al. SAF(2,91). 

Fossombronia marindae Perold SAF(90,91). 

Fossombronia montaguensis S.W.Arnell LES(75), SAF(2,91). 

Fossombronia monticola Perold SAF(90,91). 

Fossombronia nyikaensis Perold MLW(664). 

Fossombronia occidento-africana S.W.Arnell 1952 = Fossombronia indica Steph. 1917 (1045). 

Fossombronia porphyrorhiza (Nees) Prosk. 1955 [Dubia(2)]. According to Jones (679), this species was once 

recorded, as Noteroclada porphyrorhiza, from Mt. Kilimanjaro, Tanzania. However, as no published details have been 

located, the record is treated as dubious pending further information coming to light. 

Fossombronia pulvinata Steph. RWA(2). 

Fossombronia pusilla (L.) Nees CVI(2,37), LES(75), MLW(2), REU(1), RWA(2), ZAI(2). Though 

reported from southern Africa, Perold 1999e (91) considered the presence there of F. pusilla is yet to be confirmed. The 

specimen reported from Lesotho in Hodgetts et al. 1999 (75) should be reassessed. 

Fossombronia renateae Perold SAF(89,91). 

Fossombronia rwandaensis Perold RWA(85). 

Fossombronia spinifolia Steph. SAF(2,91). 

Fossombronia spinosa Perold SAF(88,91). 

Fossombronia stephanii Schiffn. ex Steph. REU(1). 

Fossombronia straussiana Perold MLW(79,91), SAF(79,91). 

Fossombronia swaziensis Perold SAF(86,91), SWZ(86,91). 

Fossombronia tumida Mitt. SAF(2,91). 

Fossombronia wondraczekii (Corda) Dumort. ex Lindb. ASC(2), MAU(1). 

Fossombronia zeyheri Steph. 1900 = Fossombronia crispa Nees 1846 (82, 83). 

Fossombronia zuurbergensis Perold SAF(663). 

Fossombronia sp. MDG(720). 

Frullania abyssinica Steph. 1910 perhaps = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania acutiloba Gerola [non Mitt. 1861] ETH(2). 

Frullania affinis Nees et Mont. 1843 MAD(1), MAU(1), REU(1). [Dubia(1)]. 

Frullania africana Steph. 1891 = Frullania riojaneirensis (Raddi) Ångstr. 1876 (949).


Frullania albertii Steph. BUR(2), KEN(2). 

Frullania anderssonii Ångstr. COM(1), MAU(1,43), MDG(1), REU(1,4), SAF(2), SEY(1). 

Frullania angolensis Steph. 1910 = Frullania obscurifolia Mitt. 1879 (147). 

Frullania angulata Mitt. var. angulata ANG(2), ANN(2,33), BIO(2,33,36,72), BUR(2), CMN(2), 

COM(1), ETH(2), KEN(2,665,731), MAU(1), MDG(1,720), MLW(2,114), MOZ(2), NGA(2), 

REU(1,4), RWA(2), SAO(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Frullania angulata var. goetzeana (Steph.) Demaret et Vanden Berghen 1950 = Frullania angulata Mitt. 

1863 var. angulata (147). 

Frullania angulata var. laciniata Demaret et Vanden Berghen MDG(1), MLW(2). 

Frullania angulata fo. serratoides Vanden Berghen ZAI(2). 

Frullania angustifolia Steph. 1910 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania apertilobula Gerola ETH(2). 

Frullania apicalis Mitt. ANN(2), BIO(2,33,36), BUR(2), COM(1), ETH(2), GHA(2), GUI(2), 

IVO(2), KEN(2,71,731), LBR(2), MAU(1,4), MDG(1,4), MLW(114), MOZ(2), NGA(2), REU(1,4), 

RIM(10), ROD(1,723), RWA(2), SAO(2), SEY(4,108), SIE(2), TAN(2), UGA(2), ZAI(2), ZIM(2). 

Frullania apicalis var. camerunensis Vanden Berghen CMN(2), KEN(731). 

Frullania apiculata (Reinw. et al.) Nees CMN(2), COM(1), GAB(2), GHA(2), GUI(2), MAU(1), 

MDG(1,4,720), REU(1,4), RIM(10), RWA(2), SEY(1,4), ZAI(2). 

Frullania apiculata Dumort., nom. inval. = Frullania apiculata (Reinw. et al.) Nees in Gottsche et al. 1845 

(147). 

Frullania apiculata Pearson 1887 [non (Reinw, et al.)Nees 1835] = Frullania serrata Gottsche 1845 var. 

serrata (147). 

Frullania arecae (Spreng.) Gottsche BIO(2,33), BUR(2), CMN(2), ETH(2), GHA(2), 

KEN(2,71,665,731,759), MDG(1), MLW(2), MOZ(2), NGA(2), REU(1), RWA(2), SAF(2,75), 

SAO(2), SIE(2), TAN(2), UGA(2,42,96), ZAI(2), ZIM(2). 

Frullania armatistipula Steph. 1910 = Frullania ericoides (Nees) Mont. 1839 (147). 

Frullania bangiensis Steph. 1910 = Frullania squarrosa Dumort. 1835 (951, 147) = Frullania ericoides 

(Nees) Mont. 1839 (920, 147). 

Frullania bessonii Steph. 1893 probably = Frullania loricata Pearson 1891 (147). 

Frullania bilabiata Steph. 1911 = Frullania loricata Pearson 1891 (147). 

Frullania boivinii Steph. 1911 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania borbonica Lindenb. MAU(1), MDG(1), REU(1). 

Frullania borgenii Pearson 1891 = Frullania obscurifolia Mitt. 1879 (147). 

Frullania brasiliensis Raddi 1822 MAD(1). [Excludenda(1)]. 

Frullania brunnea (Spreng.) Gottsche et al. MLW(2), SAF(2). 

Frullania brunnthaleri Steph. 1911 = Frullania apicalis Mitt. 1879 var. apicalis (147). 

Frullania bullata Steph. MAU(1), MDG(1), MOZ(2), TAN(2). 

Frullania bursicula Steph. 1911 = Frullania socotrana Mitt. 1888 (147). 

Frullania bystroemii S.W.Arnell 1961 = Frullania spongiosa Steph. 1894 (147). 

Frullania caffraria Steph. ANG(2), BIO(2,33), CMN(2), DJI(2), ERI(2), ETH(2), 

KEN(2,21,71,665,731,759), MAU(1), MDG(1), MLW(2,114), NGA(2), REU(1), RWA(2), SAF(2), 

TAN(2), UGA(2,50,96), ZAI(2), ZAM(2), ZIM(2). 

Frullania caffraria fo. anomala Vanden Berghen ZAI(2). 

Frullania cambouena Steph. 1891 = Frullania loricata Pearson 1891 (147). 

Frullania camerunensis Steph. 1911 = Frullania apiculata (Reinw. et al.) Nees 1845 (147). 

Frullania capensis Gottsche MAU(1), MDG(1), MLW(114), MOZ(2), REU(1,4), SAF(2), TAN(2), 

ZIM(2). 

Frullania capensis sensu Sim 1926 [non Gottsche 1845] = Frullania obscurifolia Mitt. 1879 (147). 

Frullania cardotii Steph. 1910 = Frullania obscurifolia Mitt. 1879 (147). 

Frullania cavallii Gola UGA(2). 

Frullania chiovendae Steph. ETH(2). 

Frullania clavellifera Steph. 1911 = Frullania lindenbergii Lehm. 1844 (147). 

Frullania colonica Steph. 1910 = Frullania spongiosa Steph. 1894 (951, 147). 

Frullania conjugata Steph. 1911 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania cordata Mitt. 1863 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania cordifolia Steph. 1911 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania creberrima Steph. 1911 = Frullania loricata Pearson 1891 (147). 

Frullania crenuliflora Pearson 1924 perhaps = Frullania trinervis (Lehm.) Drège 1843 (147). Taxon erroneously 

transcribed as F. crenatiflora in (147).


Frullania crispistipula Steph. 1894 = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Frullania cuculliloba Steph. 1911 = Frullania obscurifolia Mitt. 1879 (147). 

Frullania cyparioides (Schwägr.) Nees 1838 MAU(1). [Excludenda(1)]. 

Frullania dalleana Gerola 1947 perhaps = Frullania socotrana Mitt. 1888 (147). 

Frullania densepinnata Steph. 1911 = Frullania apicalis Mitt. 1878 var apicalis (147). 

Frullania dentilobula Steph. 1894 = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Frullania depressa Mitt. BUR(2), CMN(2), COM(4), ETH(2), KEN(2,759), LES(75), MDG(1), 

MLW(114), NGA(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Frullania dilatata (L.) Dumort. CVI(37,124). 

Frullania diptera (Lehm.) Drège ANG(2), ANN(2,33), BEN(22), GHA(2), KEN(2,665), MAU(1), 

MDG(1,720), MLW(114), NGA(2), REU(1,4), SAF(2), SAO(2), TAN(2), TOG(101), UGA(42,50), 

ZAI(2). 

Frullania dupuisana Steph. ex Bonner 1965, nom. nud. = Frullania ericoides (Nees) Mont. 1839 (951, 147). 

Frullania dusenii Schiffn. in sched. [non Steph. 1905] = Frullania spongiosa Steph. 1894 (147). 

Frullania ecklonii (Spreng.) Gottsche et al. CMN(2), KEN(2,759), MLW(2), SAF(2), SAO(2), 

TAN(2), UGA(2,96), ZAI(2). 

Frullania elliotii Steph. 1894 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania emergens Mitt. 1863 = Frullania ecklonii (Spreng.) Gottsche et al. 1845 (147). 

Frullania epiphylla Vanden Berghen 1976, hom. illeg. [non S.Hatt. 1974] = Frullania vandenberghenii Pócs 

1979 (950). 

Frullania eplicata Steph. GHA(2), MLW(114), MOZ(2), SAF(2), TAN(2), ZAI(2), ZAM(2), 

ZIM(2). 

Frullania ericicola Steph. in Bonner 1965 perhaps = Frullania trinervis Steph. 1894 (147). 

Frullania ericoides (Nees) Mont. ALD(34), ANG(2), ANN(2,33), ASC(2), BEN(22), BIO(2,33), 

BUR(2), CAF(2), CMN(2), COM(1), CON(2), CVI(2,37), ERI(121), ETH(2), GAB(2), GHA(2), 

GUI(2), IVO(51,53), KEN(2,731), MAU(1,43), MDG(1), MLW(2,114), MOZ(2), NGA(2), PRI(2), 

REU(1), ROD(1,723), RWA(2), SAF(2,75), SEY(1,4), SIE(2), SOC(2), TAN(2), TOG(2,101), 

UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Frullania erosolimbata Steph. 1924 = Frullania caffraria Steph. 1894 (147). 

Frullania erronea Steph. 1911 = Frullania depressa Mitt. 1863 (147). 

Frullania exigua Steph. 1910 = Frullania obscurifolia Mitt. 1879 (951, 147). 

Frullania filiformis Steph. 1910 = Frullania loricata Pearson 1891 (147). 

Frullania fusca Steph. 1894 = Frullania lindenbergii Lehm. 1844 (147). 

Frullania gabonensis Vanden Berghen CMN(2), GAB(2), MDG(1,4). 

Frullania garsidei S.W.Arnell 1953 = Frullania variegata Steph. 1894 (147). 

Frullania goetzeana Steph. 1901 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania gracilis (Reinw. et al.) Gottsche et al. 1845 MAU(1). [Excludenda(1)]. 

Frullania grossiclava Steph. MDG(1,4,720), MLW(114), MOZ(2), TAN(2), ZAI(2), ZIM(2). 

Frullania hamatirostra Gerola 1947 perhaps = Frullania ericoides (Nees) Mont. 1839 (147). 

Frullania hedbergii Vanden Berghen 1953 = Frullania albertii Steph. 1916 (949). 

Frullania hians (Lehm. et Lindenb.) Mont. et Nees 1839 = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Frullania hildebrandtii Steph. 1910 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania hispida Steph. 1911 = Frullania angulata Mitt 1863 var. angulata (147). 

Frullania hoehneliana Steph. 1894 = Frullania caffraria Steph. 1894 (952, 147). 

Frullania holstii Steph. 1894 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania humbertii Vanden Berghen MDG(1), REU(1). 

Frullania imerinensis Steph. MDG(1,720), RWA(2), SEY(4). 

Frullania inflatiloba Steph. 1910 = Frullania diptera (Lehm. et Lindenb.) Drège 1843 (147). 

Frullania insularis Beauverd 1924 = Frullania apicalis Mitt. 1878 var. apicalis (147). 

Frullania johannensis Steph. 1911 = Frullania serrata var. pertenuis (Nees) Gottsche et al. 1845 (147). 

Frullania junghuhniana Gottsche 1845 MAS(1), MAU(1), REU(1). [Excludenda(1)]. 

Frullania jungneri Steph. 1911 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania kilimanjarica Vanden Berghen 1953 = Frullania apicalis Mitt. 1879 var. apicalis (147). 

Frullania kinangopii Gola 1916 perhaps = F. apicalis Mitt. 1879 (147). 

Frullania kizarensis Steph. 1911 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania lacerata Steph. 1911 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania laceriloba Steph. 1911 = Frullania apicalis Mitt. 1879 var. apicalis (147). 

Frullania laceriloba var. acutifolia S.W.Arnell 1956 = Frullania apicalis Mitt. 1879 var. apicalis (147). 

Frullania laxepinnata Steph. 1895, nom. nud. [non Steph 1911] = Frullania angulata Mitt. 1963 var. angulata 

(147).


Frullania lepervanchii Steph. 1911 = Frullania lindenbergii Lehm. 1844 (147). 

Frullania letestui Vanden Berghen CAF(2), CMN(2), GAB(2), ZAI(2). 

Frullania levicalyx Steph. 1912 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania lindenbergii Lehm. KEN(2), MAU(1), MDG(1,4,720), MLW(114), MOZ(2), REU(1,4), 

SAF(2), SEY(4,108), TAN(2), UGA(50), ZIM(2). 

Frullania lobangensis Steph. 1910 = Frullania caffraria Steph. 1894 (147). 

Frullania longirostris Steph. 1894 = Frullania ecklonii (Spreng.) Gottsche et al. 1845 (949). 

Frullania longistipula Steph. MDG(1). 

Frullania loricata Pearson MDG(1,720), MLW(2,114), TAN(2), UGA(2). 

Frullania loricata var. laxa Pearson MDG(1). 

Frullania macroloba Steph. 1910 = Frullania anderssonii Ångstr. 1873 (147). 

Frullania madagascariensis Gottsche 1882, nom. dub. MAD(1). [Dubia(1)]. 

Frullania mauritiana Austin 1870, nom. dub. MAU(1). [Dubia(1)]. 

Frullania mildbraedii Steph. 1911 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania molleri Steph. 1894 perhaps = Frullania caffraria Steph. 1894 (147). 

Frullania multiramosa Steph. 1894 = Frullania capensis Gottsche 1845 (147). 

Frullania mundiana Lindenb. et Gottsche 1847 = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Frullania natalensis Sim 1926 = Frullania depressa Mitt. 1863 (147). 

Frullania nervosa Mont. 1840 = Frullania tamarisci (L.) Dumort. 1835 (953, 147). 

Frullania newtonii Steph. 1911 perhaps = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania nigricans Steph. 1911 = Frullania apiculata (Reinw. et al.) Nees 1845 (147). 

Frullania nodulosa (Reinw. et al.) Gottsche et al. CMN(2), COM(1), GHA(2), IVO(2), MDG(1,4), 

NGA(2), SEY(1), SIE(2), TAN(2), ZAI(2). 

Frullania nutans Steph. 1901 perhaps = Frullania caffraria Steph. 1894 (147). 

Frullania obconica Steph. 1910 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania obscura Steph. 1910, hom. illeg. [non (Lehm. et Lindenb.) Nees 1845] = Frullania anderssonii 

Ångstr. 1873 (147). 

Frullania obscurifolia Mitt. ANG(2), ANN(2,33), BUR(2), CAF(2), CMN(2), COM(4), CON(2), 

DJI(2), ETH(2), GHA(2), KEN(2,21,71,665,759), LES(75), MAU(1), MDG(1,4), MLW(2,114), 

MOZ(2), NGA(2), ROD(1,723), RWA(2), SAF(2,75), SEY(1,4), SIE(2), SWZ(2), TAN(2), 

UGA(2,42), ZAI(2), ZAM(2), ZIM(2). 

Frullania onraedtii Vanden Berghen MAU(1). 

Frullania ovalifolia Kiaer et Pearson 1891 = Frullania anderssonii Ångstr. 1873 (147). 

Frullania ovifolia Kiaer et Pearson ex Steph. 1910 = Frullania anderssonii Ångstr. 1873 (147). 

Frullania parisii Steph. 1908 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania perrotana Steph. 1911 = Frullania apiculata (Reinw. et al.) Nees 1845 (147). 

Frullania picta Steph. 1894 MAU(1). [Excludenda(1)]. 

Frullania platyflora Pearson 1922 = Frullania eplicata Steph. 1911 (147). 

Frullania purpurea Steph. COM(4), MAU(1), MDG(1,4,720), NGA(2), RIM(10), SAO(2), SEY(4), 

SIE(2). 

Frullania purpureopicta Steph. 1910 = Frullania socotrana Mitt. 1888 (951). 

Frullania rehmannii Steph. 1911 = Frullania capensis Gottsche 1845 (147). 

Frullania renauldii Steph. 17 Jul 1911 [non 28 Feb 1911] = Frullania apicalis Mitt. 1878 var. apicalis 

(147). 

Frullania renauldii Steph. 28 Feb 1911 [non 17 Jul 1911] = Frullania variegata Steph. 1894 (147). 

Frullania renistipula Steph. 1910 = Frullania anderssonii Ångstr. 1873 (147). 

Frullania repandistipula Sande Lac. COM(1), MDG(1), REU(1,4), SEY(1,4,108), TAN(2). 

Frullania rigida Steph. MDG(1). 

Frullania riojaneirensis (Raddi) Ångstr. CMN(2), GAB(2), GHA(2), GUI(2), NGA(2), PRI(2), 

SAO(2), TAN(2), TOG(101), ZAI(2). 

Frullania riparia Hampe ex Lehm. 1838. [Excludenda(2)]. According to Arnell (160), Frullania riparia was recorded 

in error for F. ericoides (819) from the Cape Verde Islands. Grolle & Long (160) pointed out that F. riparia is 

confirmed only from Europe and North America. 

Frullania robillardii Steph. 1893 = Frullania anderssonii Ångstr. 1873 (147). 

Frullania robusta Kiaer 1891, nom. inval. [non (Spruce) Steph. 1911] = Frullania loricata Pearson 1891 

(147). 

Frullania rodriguezii Steph. 1894 = Frullania apicalis Mitt. var. apicalis 1878 (147). 

Frullania rupestris Steph. 1894 = Frullania schimperi Nees 1845 var. schimperi (147). 

Frullania rutenbergii Gottsche 1882 = Frullania apicalis Mitt. 1879 var. apicalis (147). 

Frullania sandei Kiaer ex Steph. 1894, with some doubt = Frullania caffraria Steph. 1894 (147).


Frullania schimperi Nees var. schimperi BUR(2), CMN(2), ETH(2), KEN(2,71,665), MDG(1,720), 

RWA(2), SOC(764), TAN(2), UGA(50). 

Frullania schimperi var. laciniata Vanden Berghen MDG(1). 

Frullania semienana Gola ETH(2). 

Frullania serrata Gottsche var. serrata CMN(2), COM(1), ETH(2), KEN(2,731), MAU(1), 

MDG(1,4,720), MLW(2,114), MOZ(2), REU(1,4), RWA(2), SAF(2), SAO(2), TAN(2), UGA(2,96), 

ZAI(2), ZIM(2). 

Frullania serrata var. pertenuis (Nees) Gottsche et al. COM(1), MDG(1,4), RWA(2), TAN(2), 

ZAI(2). 

Frullania silvestris Sim 1926, nom. illeg. [non Steph. 1908]. SAF(894). [Dubia(147)]. Placed in synonymy 

with F. natalensis by Arnell (812), but Vanden Berghen (147) was unable to come to any conclusion as to its identity. 

Frullania socotrana Mitt. CMN(2), CVI(2,37), KEN(2), LES(75), MDG(1), MOZ(2), SAF(2), 

SOC(2), TAN(2), UGA(2,50,96), ZAI(2). 

Frullania sphaeroflora Steph. 1911 = Frullania bullata Steph. 1910 (147). 

Frullania spongiosa Steph. BEN(22), CMN(2), CVI(2,37), GHA(2), IVO(2,53), MDG(4), NGA(2), 

RIM(10), SAF(2), SEY(4), TAN(2), TOG(101), UGA(50), ZAI(2), ZIM(2). 

Frullania squarrosa Gottsche et al. 1845 = Frullania ericoides (Nees) Mont. 1839 (920, 147). 

Frullania staudtiana Steph. 1911 probably = Frullania apiculata (Reinw. et al.) Nees 1845 (147). 

Frullania stephanii Schiffn. 1893 perhaps = Frullania caffraria Steph. 1894 (147). 

Frullania stipatiflora Steph. 1911 = Frullania apicalis Mitt. 1878 var. apicalis (147). 

Frullania stricta (F.Weber) Lindenb. 1845 = Frullania serrata Gottsche 1845 var. serrata (147). 

Frullania stylata Gerola 1947 perhaps = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania subatrata Steph. 1911 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania subplana Gottsche ex Steph. 1910 = Frullania bullata Steph. 1910 (147). 

Frullania substricta Steph. 1913 probably = Frullania angulata Mitt. 1863 (147). 

Frullania tamarisci (L.) Dumort. CVI(2,20,37). 

Frullania tatanarivensis Steph. 1894 = Frullania ericoides (Nees) Mont. 1839 (951, 147). 

Frullania thomeensis Steph. 1910 probably = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Frullania tixieriana Vanden Berghen 1976 = Frullania repandistipula Sande Lac. 1855 (847). 

Frullania traversiana Steph. 1910 = Frullania depressa Mitt. 1863 (147). 

Frullania trinervis (Lehm.) Drège ANG(2), BIO(2,33), CAF(2), CON(2), ETH(2), GHA(2), GUI(2), 

KEN(2,759), LES(2,75), MAU(4), MDG(1), MLW(2,114), MOZ(2), NGA(2), SAF(75), SEY(4), 

SIE(2), SOC(94), TAN(2), TOG(101), UGA(2,42), ZAI(2), ZAM(2), ZIM(2). 

Frullania truncatiloba sensu Vanden Berghen 1961 [non Steph. 1910] = Frullania depressa Mitt. 1863 

(147). 

Frullania truncatiloba Steph. 1910 [non sensu Vanden Berghen 1961] = Frullania africana Steph. 1891 

(147) = Frullania riojaneirensis (Raddi) Ångstr. 1876 (949). 

Frullania ubangiensis Steph. 1910 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Frullania usagara Mitt. 1886 = Frullania ericoides (Nees) Mont. 1839 (951). 

Frullania usambarana Schiffn. ex Steph. MDG(1,4), REU(4), TAN(2). 

Frullania usambarana var. reducta Vanden Berghen MDG(1), REU(1). 

Frullania usambarensis Steph. 1910 = Frullania spongiosa Steph. 1894 (147). 

Frullania vandenberghenii Pócs MDG(1,4), UGA(50). 

Frullania vanderystii Steph. 1910 = Frullania obscurifolia Mitt. 1879 (951). 

Frullania varia Gottsche 1882 = Frullania angulata Mitt. 1863 var. angulata (147). 

Frullania variegata Steph. BUR(2), CMN(2), MAU(1), MDG(1,4,720), MLW(114), REU(1), 

RWA(2), SAF(2), TAN(2). 

Frullania welwitschii Steph. 1901 = Frullania squarrosa Gottsche et al. 1845 (822) = Frullania ericoides 

(Nees) Mont. 1839 (920). 

Frullanoides riojaneirensis Raddi 1822 = Frullania riojaneirensis (Raddi) Ångstr. 1876 (949). 

Frullanoides tristis (Steph.) van Slageren ANG(2), BUR(2), CAF(2), CMN(2), ETH(2), IVO(2), 

KEN(2), MDG(1), NGA(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(50), ZAI(2). 

Gamochaetium concretum (Mont.) Trevis. 1879 = Lophocolea concreta Mont. 1845 (1). 

Geocalyx orientalis Besch. et Spruce REU(1). 

Gongylanthus ericetorum (Raddi) Nees BUR(2), CMN(2), ETH(2), KEN(2), LES(75), MDG(1), 

MLW(2,29), REU(1), SAF(2,75), SIE(2), TAN(2), UGA(776), ZAI(2), ZAM(2), ZIM(2). Listed in 

error from Rwanda in (2). 

Gongylanthus renifolius (Mitt.) Steph. RWA(2), SAF(2), TAN(2), UGA(776). Listed in error from Burundi 

in (2). 

Gongylanthus richardsii E.W.Jones CMN(2), SIE(2), TAN(2), UGA(42).


Gongylanthus scariosus (Lehm.) Steph. CMN(2), ETH(2), MLW(29), REU(1), SAF(2). 

Gottschea alata (Lehm.) Nees 1844 = Schistochila alata (Lehm.) Schiffn. 1893 (954). 

Gottschea borbonica (Steph.) Bonner 1966 = Gottschea neesii Mont. 1843 (871, 954). 

Gottschea engleriana (Steph.) Grolle et Zijlstra COM(1,4), MLW(2,109), RWA(2), TAN(2). 

Gottschea neesii Mont. COM(4), MAS(1), MAU(1,4), MDG(1,4,720), REU(1,4). 

Gottschea schwaegricheniana Nees et Mont. 1843, nom. inval. = Gottschea neesii Mont. 1843 (954). 

Gottschea sphagnoides (Schwägr.) Lindb. MAU(1), MLW(2,109), MOZ(2), REU(1,4), SAF(2), 

ZIM(2). 

Gottschea thouarsii (Hook.) Nees 1844 = Gottschea sphagnoides (Schwägr.) Lindb. 1876 (885, 954). 

Gottschelia schizopleura (Spruce) Grolle MDG(1,4,720), MLW(2,114), REU(1,4), TAN(2). 

Grimaldia abyssinica Gola 1914 = Plagiochasma eximium (Schiffn. in Steph.) Steph. 1898 (254). 

Grimaldia capensis Steph. 1898 = Mannia capensis (Steph.) S.W.Arnell 1957 (715). 

Grimaldia delessertii Mont. 1846, nom. inval. = Marchantia globosa Brid. ex F.Weber 1815 (132). 

Grimaldia dichotoma Raddi 1818, nom. illeg. = Mannia androgyna (L. emend. Lindb.) A.Evans 1938 (920). 

Gymnanthe anisodonta (Hook.f. et Taylor) Gottsche 1864 = Tylimanthus anisodontus (Hook.f. et Taylor) 

Mitt. 1875 (1046). 

Gymnanthe belangeriana (Lehm. et Lindenb.) Steph. 1894 = Notoscyphus lutescens (Lehm. et Lindenb.) 

Mitt. 1873 (916, 1). 

Gymnanthe biloba Mitt. 1863 = Andrewsianthus bilobus (Mitt.) Grolle 1963 (315). 

Gymnanthe decipiens (Hook.) Mitt. 1863 = Adelanthus decipiens (Hook.) Mitt. 1864 (465). 

Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust. SAF(2). 

Gymnocoleopsis multiflorus (Steph.) R.M.Schust. TAN(2), ZAI(2). 

Gymnomitrion africanum (Steph.) Horik. 1943 = Marsupella africana Steph. ex Bonner 1953 (416) = Apomarsupella 

africana (Steph. ex Bonner) R.M.Schust. 1996 (153). 

Gymnomitrion andinum (Herzog) Herzog SAF(2,111). 

Gymnomitrion belangerianum (Lehm. et Lindenb.) Gottsche 1844 = Notoscyphus lutescens (Lehm. et Lindenb.) 

Mitt. 1873 (916, 1). 

Gymnomitrion elgonense S.W.Arnell 1956 = Gymnomitrion laceratum (Steph.) Horik. 1943 (416). 

Gymnomitrion elgonense var. squarrosum S.W.Arnell 1956 = Gymnomitrion laceratum (Steph.) Horik. 

1943 (416). 

Gymnomitrion laceratum (Steph.) Horik. KEN(2), RWA(2), SAF(2,75), TAN(2), UGA(7), ZAI(2). 

Gymnomitrion scariosum (Lehm.) Nees 1844 = Gongylanthus scariosus (Lehm.) Steph. 1906 (170). 

Haplolejeunea cucullata (Steph.) Grolle CMN(2). 

Haplolejeunea sticta Grolle MDG(1,720), REU(4). 

Haplomitrium andinum (Spruce) R.M.Schust. 1963 = Haplomitrium blumii (Nees) R.M.Schust. 1963 

(1004). 

Haplomitrium blumii (Nees) R.M.Schust. RWA(2), ZAI(2). 

Haplomitrium gibbsiae (Steph.) R.M.Schust. LES(75), UGA(115). 

Harpalejeunea delessertii (Nees et Mont.) Steph. 1913 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 

1984 (582). 

Harpalejeunea filicuspis (Steph.) Mizut. COM(1,571), SEY(1,571). 

Harpalejeunea fischeri Tixier RWA(2), UGA(42). 

Harpalejeunea marginalis (Hook.f. et Taylor) Steph. 1913. SAF(2). [Excludenda(2)]. 

Harpalejeunea punctata (Herzog) S.W.Arnell 1965 MAU(1). [Excludenda(1)]. 

Harpalejeunea riddleana (Steph.) Mizut. 1973 = Harpalejeunea filicuspis (Steph.) Mizut. 1973 (345). 

Harpalejeunea solitaria (Gottsche) Steph. 1913 MAD(1). [Dubia(1)]. 

Herbertus capensis (Steph.) Sim COM(1), ETH(2), MDG(1), MLW(2), SAF(2,75), UGA(50,96). 

Herbertus capillaris Steph. 1892 MAD(1), REU(1). [Dubia(1)]. 

Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. ETH(2), KEN(2,71), MAU(1), MDG(1,4), 

REU(1), RWA(2), TAN(2), UGA(2,96), ZAI(2). Collections from Ethiopia, Tanzania, D.R. Congo and Madagascar 

were described under H. subdentatus (Steph.) Fulford (2), but Hodgetts (761) has shown that the type differs from 

the African material. 

Herbertus doggeltianus (Steph.) Demaret UGA(50), ZAI(2). 

Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle MAU(1), MDG(1,720), SEY(1,4). 

Herbertus juniperinus Trevis. 1877 = Herbertus juniperoideus (Sw.) Grolle 1961 (1) [Excludenda(1)]. 

Herbertus juniperoideus (Sw.) Grolle 1961 MAD(1), REU(1). [Excludenda(1)]. 

Herbertus leiocladus (Taylor) Kuntze 1891 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 (895). 

Herbertus lobatus (Steph.) Demaret UGA(50), ZAI(2). 

Herbertus mascarenicus (Steph.) S.W.Arnell ex Váňa, in Váňa et al. MAS(1), REU(1). Belongs to the 

H. capensis complex (847), but treated as a separate species by Váňa (156).


Herbertus mollis (Steph.) Dusén CMN(2), SAO(2). 

Herbertus ochroleucus (Spreng.) Schiffn. 1898 = Lepicolea ochroleuca (Spreng.) Spruce 1885 (107). 

Herbertus seriatus (Steph.) S.W.Arnell 1965, nom. inval. = Herbertus grossevittatus (Steph.) S.W.Arnell ex 

Grolle 1995 (847). 

Herbertus stuhlmannii (Steph.) Demaret UGA(2), ZAI(2). 

Herbertus subdentatus auct. [non (Steph.) Fulford] = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 

1877 (761). 

Herbertus sp. (Schisma balboi Gola) KEN(2). 

Herbertus sp. (Schisma kilimandjarense Steph.) TAN(2). 

Herbertus sp. (Schisma mascarenicum var. kikujense Gola) KEN(2). 

Herpetium integrum Nees et Mont. 1843 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (884, 1). 

Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910 MAU(1). [Excludenda(1)]. 

Heteroscyphus decurrens (Nees) Schiffn. 1910 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984 

(623). 

Heteroscyphus dubius (Gottsche) Schiffn. CAF(2), CMN(2), GHA(2,165), IVO(2), KEN(731), 

MAU(1), MDG(1), MLW(2,29), NGA(2), PRI(2), REU(1), RIM(10), ROD(1), SAF(2), SEY(1,13), 

SIE(2), TAN(2), TOG(101), UGA(50,115), ZAI(2). The record from British Togoland published in Jones 

1953b (165) was erroneously listed in (2) under Togo. The site is in Ghana. 

Heteroscyphus fasciculatus (Nees) Schiffn. 1910 = Clasmatocolea fasciculata (Nees) Grolle 1960 (314, 97). 

Heteroscyphus grandistipus (Steph.) Schiffn. REU(1). 

Heteroscyphus grandistipus subsp. henkelii (Sim) Grolle ZIM(2). 

Heteroscyphus hamatistipulus (Steph.) Schiffn. 1910 = Chiloscyphus hamatistipulus Steph. 1907 (982) = 

Chiloscyphus dubius Gottsche 1882 (165) = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (1). 

Heteroscyphus loangensis (Steph.) Schiffn. 1910 = Chiloscyphus loangensis Steph. 1893 (892) probably = 

Chiloscyphus dubius Gottsche 1882 (165) = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (1). 

Heteroscyphus lucidus (Spreng. ex Lehm.) Schiffn. 1910 = Lophocolea lucida (Spreng. ex Lehm.) Mont. 

1839 (313). 

Heteroscyphus rabenhorstii (Steph.) Schiffn. 1910 = Lophocolea fasciculata (Nees ) S.W.Arnell et Grolle in 

Grolle 1959 (313) = Clasmatocolea fasciculata (Nees) Grolle 1960 (97). 

Heteroscyphus spectabilis (Steph.) Schiffn. CMN(2), SAO(2). 

Heteroscyphus splendens (Lehm. et Lindenb.) Grolle MAU(1), MDG(4,720), REU(1), SEY(1), 

TAN(2). 

Homalolejeunea acutiloba (Hook.f. et Taylor) Steph. 1890 = Phragmicoma acutiloba (Hook.f. et Taylor) 

Gottsche et al. 1847 = Marchesinia brachiata (Sw.) Schiffn. 1893 (831) 

Homalolejeunea excavata (Mitt.) Steph. 1912 = Marchesinia excavata (Mitt.) Schiffn. 1893 (891). 

Homalolejeunea henriquesii Steph. 1888 = Marchesinia excavata (Mitt.) Schiffn. 1893 (886). 

Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle SAF(2,504). Following the redefinition of 

Hyalolepidozia, South African records of H. bicuspidata were provisionally reassigned by Schuster (504) to Paracromastigum 

tristanianum. However, since its taxonomic position is not settled, the taxon is retained under H. bicuspidata in this 

checklist. 

Hygrolejeunea acuta (Mitt.) Vanden Berghen 1961 = Lejeunea acuta Mitt. 1863 (1). 

Hygrolejeunea alata (Gottsche) Steph. 1914 = Lejeunea alata Gottsche 1845 (828). 

Hygrolejeunea breuteliana Steph. 1914 = Lejeunea eckloniana Lindenb. 1845 (580). 

Hygrolejeunea cordigera Steph. 1896 = Cheilolejeunea cordigera (Steph.) Grolle 1977 (828). 

Hygrolejeunea dismieri Steph. 1913 = Dicranolejeunea madagascariensis Steph. 1912 (828) = Acanthocoleus 

madagascariensis (Steph.) Kruijt 1988 (542). 

Hygrolejeunea grossecristata Steph. 1896 = Lejeunea grossecristata (Steph.) E.W.Jones 1968 (864). 

Hygrolejeunea leucosis (Besch. et Spruce) Steph. 1914 = Lejeunea leucosis Besch. et Spruce 1890 (1). 

Hygrolejeunea lyratiflora Steph., Hedwigia 31: 169. May-Jun 1892, nom. inval. [= Hygrolejeunea lyratiflora 

(Steph.) Steph. 1914] = Lejeunea lyratiflora Steph. 1892 (733). 

Hygrolejeunea papilionacea (Steph.) Steph. 1914 = Lejeunea papilionacea Steph. 1892 (157, 670). 

Hygrolejeunea parva Steph. 1914 = Lejeunea eckloniana Lindenb. 1845 (580). 

Hygrolejeunea patellirostris Steph. 1896 = Lejeunea sp. 

Hygrolejeunea pterota (Hook.f. et Taylor) Steph. 1914 = Cheilolejeunea ascensionis (Hook.f. et Taylor) 

Grolle 1988 (930). 

Hygrolejeunea pulcherrima Steph. 1914 = Lejeunea acuta Mitt. 1863 (140). 

Hygrolejeunea staudtiana Steph. 1914 = Lejeunea sp. 

Hypenantron abyssinicum (Gottsche) Kuntze 1891 = Asterella abyssinica (Gottsche) Grolle 1972 (85). 

Hypenantron gondarense (Gola) Cufod. 1952 = Fimbraria gondarensis Gola 1914 (502) = Asterella pappii 

(Gola) Grolle 1976 (999). 

Hypenantron incrassatum (Steph.) Cufod. 1952 = Fimbriaria incrassata Steph. 1899 (502).


Hypenantron pappii (Gola) Cufod. 1952 = Asterella pappii (Gola) Grolle 1976 (502, 992). 

Hypenantron pirottae (Gola) Cufod. 1952 = Fimbriaria pirottae Gola 1914 (502). 

Inflatolejeunea capensis S.W.Arnell 1953 = Lejeunea capensis Gottsche 1845 (170). 

Inflatolejeunea ecarinata (Steph.) S.W.Arnell 1961 = Lejeunea ecarinata (Steph.) Steph. 1890 (863). 

Inflatolejeunea eplicata (Steph.) S.W.Arnell 1961 = Lejeunea eplicata Steph. 1915 (863) = Lejeunea ramosissima 

Steph. 1886 (25). 

Isopaches decolorans (Limpr.) Buch 1932 = Lophozia decolorans (Limpr.) Steph. 1902 (163). 

Isotachis aspera Steph. 1911 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Isotachis aspera var. transvaalensis S.W.Arnell 1963, nom. inval. = Isotachis aubertii (Schwägr.) Mitt. 1886 

(163). 

Isotachis aubertii (Schwägr.) Mitt. ASC(2), BUR(2), CMN(2), KEN(2), MAU(1), MDG(1,720), 

MLW(114), REU(1,4), ROD(1), RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2), ZIM(2). 

Isotachis capensis Herzog 1952 = Lophocolea fasciculata (Nees ) S.W.Arnell et Grolle in Grolle 1959 (313) 

= Clasmatocolea fasciculata (Nees) Grolle 1960 (97). 

Isotachis conistipula Steph. 1911 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Isotachis gordonii Steph. 1895 = Isotachis aubertii (Schwägr.) Mitt. 1886 (496). 

Isotachis perfoliata Steph. 1886 = Heteroscyphus spectabilis (Steph.) Schiffn. 1910 (163). 

Isotachis renistipula Steph. 1911 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Isotachis rutenbergii Gottsche 1882 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Isotachis serrulata (Sw.) Gottsche 1864 MAU(1). [Excludenda(1)]. 

Isotachis uncinata (F.Weber) Gottsche 1864 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Iwatsukia jishibae (Steph.) N.Kitag. BIO(36), COM(4), MAU(1), MDG(4,720), MLW(114), 

REU(1,4), SEY(1,4). 

Jamesoniella aspera Steph. 1917 = Gongylanthus ericetorum (Raddi) Nees 1836 (318). 

Jamesoniella colorata (Lehm.) Schiffn. SAF(2,75). 

Jamesoniella contracta (Reinw. et al.) N.Kitag. COM(4), MAU(1), MDG(4), REU(4), SEY(1,4), 

TAN(2). 

Jamesoniella grandiflora (Lindenb. et Gottsche) Steph. 1901 = Cryptochila grandiflora (Lindenb. et Gottsche) 

Grolle 1971 (318). 

Jamesoniella oenops (Lindenb. et Gottsche) Steph. 1901 = Jamesoniella colorata (Lehm.) Schiffn. 1893 

(318). 

Jamesoniella purpurascens Steph. MAU(1), MDG(1,4,720), MLW(2,114), REU(1), SAF(2), 

TAN(2), ZAI(2), ZAM(2). 

Jamesoniella rehmannii (Steph.) Steph 1901 = Jamesoniella purpurascens Steph. 1891 (318). 

Jamesoniella seychellarum Steph. 1917 = Cuspidatula contracta (Reinw. et al.) Steph. (318) = Jamesoniella 

contracta (Reinw. et al.) N.Kitag. 1981 (527). 

Jensenia spinosa (Lindenb. et Gottsche) Grolle KEN(2,71,665,759), MAU(1), MLW(2,29), REU(1), 

RWA(2), SAF(2,955), STH(2), TAN(2), UGA(50), ZAI(2). 

Jubula cambouena Steph. 1924 = Jubula tonkinensis Steph. 1908 (1) [Excludenda(1)]. 

Jubula hutchinsiae subsp. javanica (Steph.) Verd. 1928 MAD(1). [Excludenda(1)]. 

Jubula javanica Steph. 1911 = Jubula hutchinsiae subsp. javanica (Steph.) Verd. 1928 (1) [Excludenda(1)]. 

Jubula tonkinensis Steph. 1908 MAD(1). [Excludenda(1)]. 

Jungermannia aberrans Nees et Mont. 1843 = Andrewsianthus aberrans (Nees et Mont.) Grolle 1963 (315). 

Jungermannia abyssinica Nees ETH(2), KEN(2), TAN(2), UGA(2). 

Jungermannia acutiloba Hook.f. et Taylor 1845 = Phragmicoma acutiloba (Hook.f. et Taylor) Gottsche et 

al. 1847 (988) = Marchesinia brachiata (Sw.) Schiffn. 1893 (831). 

Jungermannia adnexa Lehm. et Lindenb. 1832 = Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 (1) 


Jungermannia alata Lehm. 1829 = Schistochila alata (Lehm.) Schiffn.1893 (871). 

Jungermannia anisodonta Hook.f. et Taylor 1845 = Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. 1875 

(1046). 

Jungermannia apiculata Reinw. et al. 1824 = Frullania apiculata (Reinw. et al.) Nees 1845 (147). 

Jungermannia arboris-vitae With. 1776 = Porella arboris-vitae (With.) Grolle 1969 (921). 

Jungermannia arecae Spreng. 1821 = Frullania arecae (Spreng.) Gottsche 1863 (147). 

Jungermannia arguta Reinw. et al. 1824 = Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910 (1) [Excludenda(1)]. 

Jungermannia ascensionis Hook.f. et Taylor 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle 

1988 (930). 

Jungermannia atrata var. β stricta F.Weber 1815 = Frullania serrata Gottsche 1845 var. serrata (147). 

Jungermannia aubertii Schwägr. 1814 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Jungermannia auriculata Wilson 1841 = Mastigolejeunea auriculata (Wilson) Schiffn. 1893 (430).


Jungermannia aurita Lehm. 1829 = Anastrophyllum auritum (Lehm.) Steph. 1901 (163). 

Jungermannia austroafricana S.W.Arnell 1958 = Jungermannia mildbraedii Steph. 1911 (412). 

Jungermannia balfourii Váňa MAU(1). 

Jungermannia belangeriana Lehm. et Lindenb. 1833 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 

1873 (916, 1). 

Jungermannia bergiana Lehm. et Lindenb. 1832 = Clasmatocolea fasciculata (Nees) Grolle 1960 (917, 97). 

Jungermannia biapiculata Hook.f. et Taylor 1845 = Anastrophyllum piligerum (Nees) Steph. 1893 (916, 

163). 

Jungermannia bicuspidata L. 1753 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Jungermannia bidentata L. 1753 = Lophocolea bidentata (L.) Dumort. 1835 (253). 

Jungermannia bidentula F.Weber 1815 = Calypogeia bidentula (F.Weber) Nees 1845 (253). 

Jungermannia borgenii Gottsche ex Pearson ANG(2), BOT(2), CMN(2), CON(2), KEN(2), 

MAU(1), MDG(1), MLW(2,114), REU(1), RWA(2), SAF(2), SAO(2), TAN(2), UGA(2,96), 

ZAI(2), ZAM(2), ZIM(2). 

Jungermannia boryana F.Weber 1815 = Radula boryana (F.Weber) Mont. 1842 (661). 

Jungermannia brasiliensis Nees 1830, hom. illeg. [non Raddi 1822] = Symphyogyna brasiliensis Nees et 

Mont. 1836 (970). 

Jungermannia brunnea Spreng. 1827 = Frullania brunnea (Spreng. ) Gottsche et al. 1845 (147). 

Jungermannia bursata Desv. 1814, nom. illeg. = Plagiochila bursata Lindenb. 1840, nom. illeg. = Plagiochila 

aerea Taylor 1846 (959). 

Jungermannia byssophora Lehm. et Lindenb. 1832 = Pallavicinia lyellii (Hook.) Carruth. 1865 (146). 

Jungermannia calyptrifolia Hook. 1816 = Colura calyptrifolia (Hook.) Dumort. 1835 (336). 

Jungermannia campylodonta Hook.f. et Taylor 1845 = Adelanthus decipiens (Hook.) Mitt. 1864 (827, 465). 

Jungermannia capillaris Sw. 1788 = Kurzia capillaris (Sw.) Grolle 1964 (382). 

Jungermannia chrysophylla Lehm. 1835 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 (542). 

Jungermannia colorata Lehm. 1829 = Jamesoniella colorata (Lehm.) Schiffn. 1893 (318). 

Jungermannia concreta Gottsche 1863 = Syzygiella concreta (Gottsche) Spruce 1876 (416). 

Jungermannia congesta Lehm. 1829 = Lethocolea congesta (Lehm.) S.W.Arnell 1955 (538). 

Jungermannia connata Sw. 1788 = Lophocolea connata (Sw.) Mont. et Nees 1839 (1) [Excludenda(1)]. 

Jungermannia contigua Nees 1833 = Pycnolejeunea contigua (Nees) Grolle 1979 (14). 

Jungermannia contracta Reinw. et al. 1824 = Cuspidatula contracta (Reinw. et al.) Steph. (318) = Jamesoniella 

contracta (Reinw. et al.) N.Kitag. 1981 (527). 

Jungermannia convexa Thunb. 1800, hom. illeg. [non Scop. 1772] = Bazzania nitida (F.Weber) Grolle 1960 

(888). 

Jungermannia convoluta Hook.f. et Taylor 1845, hom. illeg. [non Huebener 1834] = Symphyogyna brasiliensis 

Nees et Mont. 1836 (970). 

Jungermannia cordistipula var. pertenuis Nees 1830 = Frullania serrata var. pertenuis (Nees) Gottsche 

1845 (147). 

Jungermannia cornuta Lindenb. 1829 = Ceratolejeunea cornuta (Lindenb.) Steph. 1895 (1) [Excludenda(1)]. 

Jungermannia crispa Spreng. in Gottsche et al. 1846 = Fossombronia crispa Nees 1846 (82). 

Jungermannia cucullata Reinw. et al. 1824 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995 (1). 

Jungermannia cucullata var. maior Nees 1830 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995 (1). 

Jungermannia cuneata Lehm. et Lindenb. 1832 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 

(152). 

Jungermannia cupressina Sw. 1788 = Lepidozia cupressina (Sw.) Lindenb. 1845 (382). 

Jungermannia cuspiduligera Nees 1833 = Scapania cuspiduligera (Nees) Müll.Frib. 1915 (992). 

Jungermannia decipiens Hook. 1813 = Adelanthus decipiens (Hook.) Mitt. 1864 (465). 

Jungermannia decolorans Limpr. 1880 = Lophozia decolorans (Limpr.) Steph. 1902 (163). 

Jungermannia decrescens Lehm. et Lindenb. 1832 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 

(870). 

Jungermannia decurrens Reinw. et al. 1824, hom. illeg. [non Desv. 1818] = Heteroscyphus splendens 

(Lehm. et Lindenb.) Grolle in Grolle & Piippo 1984 (623). 

Jungermannia diacantha Mont. 1856 = Arachniopsis diacantha (Mont.) M.Howe 1902 (1). 

Jungermannia dichotoma var. Nees 1831 = Plagiochila pectinata Willd. ex Lindenb. 1839 (917, 1). 

Gottsche 

Jungermannia dichotoma var. β 

et al. 1844 = Plagiochila pectinata Willd. ex Lindenb. 1839 (917, 

1). 

Jungermannia dichotoma var. γ Nees 1831 = Plagiochila paucidentata Mont. et Gottsche 1856 (917, 1). 

Jungermannia diclados Brid. ex F.Weber 1815 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 

(27). 

Jungermannia diptera Lehm. 1835 = Frullania diptera (Lehm.) Drège 1843 (147). 

Jungermannia dusenii (Steph.) Steph. CMN(2), SAO(2).


Jungermannia dominicensis Spruce 1895 = Kymatocalyx dominicensis (Spruce) Váňa 1970 (74). 

Jungermannia ecklonii Spreng. 1827 = Frullania ecklonii (Spreng.) Gottsche et al. 1845 (949). 

Jungermannia elliptica Lehm. et Lindenb. 1833 = Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. 1893 

(430). 

Jungermannia elongata Lehm. 1835 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Jungermannia emarginata Ehrh. 1784 = Marsupella emarginata (Ehrh.) Dumort. 1835 (992). 

Jungermannia ericoides Nees 1833 = Frullania ericoides (Nees) Mont. 1839 (147). 

Jungermannia exigua Taylor 1844 = Plagiochila exigua (Taylor) Taylor 1846 (1). 

Jungermannia expansa Lehm. 1829 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Jungermannia exsecta Schmidel 1797 = Tritomaria exsecta (Schmidel) Loeske 1909 (992). 

Jungermannia fasciculata Nees 1820 = Clasmatocolea fasciculata (Nees) Grolle 1960 (314, 97). 

Jungermannia fastigiata Lehm. 1829 = Riccardia fastigiata (Lehm.) Trevis. 1877 (351). 

Jungermannia filicina Sw. 1788 = Bryopteris filicina (Sw.) Nees 1845 (1) [Excludenda(1)]. 

Jungermannia filiformis Sw. 1788 = Omphalanthus filiformis (Sw.) Nees 1845 (1) [Excludenda(1)]. 

Jungermannia flava Sw. 1788 = Lejeunea flava (Sw.) Nees 1838 (864). 

Jungermannia flexuosa Lehm. 1829 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (883, 97). 

Jungermannia floccosa Lehm. et Lindenb. 1833 = Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn. 1898 

(106). 

Jungermannia fragrans Moris et De Not. 1839 = Lophocolea fragrans (Moris et De Not.) Gottsche et al. 

1845 (313). 

Jungermannia fulvifolia Hook.f. et Taylor 1845 = Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845 

(661). 

Jungermannia furcata L. 1753 = Metzgeria furcata (L.) Dumort. 1835 (767). 

Jungermannia fusca Lehm. 1829 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Jungermannia geminiflora Nees 1833 = Omphalanthus filiformis (Sw.) Nees 1845 (1) [Excludenda(1)]. 

Jungermannia geminifolia Mitt. 1863 = Syzygiella geminifolia (Mitt.) Steph. 1902 (416). 

Jungermannia gigantea F.Weber 1815 = Pleurozia gigantea (F.Weber) Lindb. 1874 (1012). 

Jungermannia gracilis Reinw. et al. 1824 = Frullania gracilis (Reinw. et al.) Gottsche et al. 1845 (1) [Excludenda(1)]. 

Jungermannia grandiflora Lindenb. et Gottsche 1847 = Cryptochila grandiflora (Lindenb. et Gottsche) 

Grolle 1971 (318). 

Jungermannia heteromorpha Lehm. 1829 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969 (844). 

Jungermannia hians Lehm. et Lindenb. in Gottsche et al. 1847 = Frullania arecae (Spreng.) Gottsche 1863 

(147). 

Jungermannia hirtella F.Weber 1815 = Plicanthus hirtellus (F.Weber) R.M.Schust. 2003 (780). 

Jungermannia humistrata Hook.f. et Taylor 1845 = Lophocolea humistrata (Hook.f. et Taylor) Gottsche et 

al. 1847 (2). 

Jungermannia incerta Gottsche 1882 MAD(1). [Dubia(1)]. 

Jungermannia jamesonii Mont. 1856 = Lophozia jamesonii (Mont.) R.M.Schust. 2002 (779). 

Jungermannia javanica Sw. 1781 = Plagiochila javanica (Sw.) Dumort. 1835 (1) [Excludenda(1, 2)]. 

Jungermannia juratzkana Limpr. 1876 = Anthelia juratzkana (Limpr.) Trevis. 1877 (992). 

Jungermannia kiaeri Austin 1875 = Cephaloziella kiaeri (Austin) Douin 1920 (1). 

Jungermannia laevifolia Hook.f. et Taylor 1844 [non Lindb. 1892] = Lepidozia laevifolia (Hook.f. et Taylor) 

Gottsche et al. 1845 (984). 

Jungermannia laevigata Schrad. 1797 = Porella arboris-vitae (With.) Grolle 1969 (921). 

Jungermannia leioclada Hook.f. et Taylor 1845 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 

(895). 

Jungermannia leucoxantha Lehm. 1829 = Fossombronia leucoxantha (Lehm.) Gottsche et al. 1845 (91). 

Jungermannia limbata Hook.f. et Taylor 1845 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. var. 

rupestre 1898 (1047, 254). 

Jungermannia lindenbergiana Lehm. 1829 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864 (465). 

Jungermannia lineata Lehm. et Lindenb. 1832 = Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans 1906 

(1)[Excludenda(1)]. 

Jungermannia longa Nees 1831 = Bazzania longa (Nees) Trevis. 1877 (1) [Excludenda(1)]. 

Jungermannia lucida Spreng. ex Lehm. 1833 = Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839 (313). 

Jungermannia lunulata F.Weber 1815 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (837). 

Jungermannia lutescens Lehm. et Lindenb. 1832 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 

(1). 

Jungermannia lyellii Hook. 1816 = Pallavicinia lyellii (Hook.) Carruth. 1865 (146). 

Jungermannia macrocalyx Mont. 1843 = Syzygiella macrocalyx (Mont.) Spruce 1876 (1) [Excludenda(1)]. 

Jungermannia marginalis Hook.f. et Taylor 1845 = Harpalejeunea marginalis (Hook.f. et Taylor) Steph. 

1913 (571). [Dubia et Excludenda(2)].


Jungermannia marginata Lehm. et Lindenb. 1833 [non Ehrh. ex Hartman 1820] = Cololejeunea marginata 

(Lehm. et Lindenb.) Schiffn. 1898 (1). 

Jungermannia mauritiana Nees 1831 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (1011, 843). 

Jungermannia microphylla var. β Hook. 1818 = Lepidozia africana Steph. 1922 (382). 

Jungermannia microscopica Taylor 1836 = Aphanolejeunea microscopica (Taylor) A.Evans 1911 (384). 

Jungermannia microscypha Hook.f. et Taylor 1845 = Phragmicoma microscypha (Hook.f. et Taylor) Mitt. 

1875 (988) = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 (831). 

Jungermannia mildbraedii Steph. RWA(2), SAF(2), TAN(2), UGA(2). 

Jungermannia minuta Schreb. ex Cranz 1770 = Anastrophyllum minutum (Schreb.) R.M.Schust. 1949 (943). 

Jungermannia mollis Nees 1830 = Lophocolea mollis (Nees) Gottsche et al. 1845 (1). 

Jungermannia multifida L. 1753 = Riccardia multifida (L.) Gray 1821 (653). 

Jungermannia multifida var. pinnatifida F.Weber 1815 = Riccardia pinnatifida (F.Weber) Carruth. 1865 (1) 


Jungermannia muricata Lehm. 1829 = Lophocolea muricata (Lehm.) Nees 1845 (313). 

Jungermannia nemoides Hook.f. et Taylor 1845 = Kurzia nemoides (Hook.f. et Taylor) Grolle 1964 (382). 

Jungermannia nitida F.Weber 1815 = Bazzania nitida (F.Weber) Grolle 1960 (888). 

Jungermannia nodulosa Reinw. et al. 1824 = Frullania nodulosa (Reinw. et al.) Nees 1845 (147). 

Jungermannia obtusata Hook.f. et Taylor 1845 = Anastrophyllum subcomplicatum (Lehm. et Lindenb.) 

Steph. 1893 (917, 163). 

Jungermannia ochroleuca Spreng. 1827 = Lepicolea ochroleuca (Spreng.) Spruce 1884 (170). 

Jungermannia oenops Lindenb. et Gottsche, in Gottsche et al. 1845 = Jamesoniella colorata (Lehm.) 

Schiffn. 1893 (318). 

Jungermannia onraedtii Váňa MDG(1), REU(1). 

Jungermannia pallens Sw. 1788 = Radula pallens (Sw.) Nees et Mont. 1839 (1) [Excludenda(1)]. 

Jungermannia palmata Hedw. 1784 = Riccardia palmata (Hedw.) Carruth. 1865 (1) [Excludenda(1)]. 

Jungermannia pearsonii Spruce 1881 = Sphenolobopsis pearsonii (Spruce) R.M.Schust. 1972 (615). 

Jungermannia pectinata Willd. ex F.Weber 1815, nom. inval. = Plagiochila pectinata Willd. ex Lindenb. 

1839 (722). 

Jungermannia pellucida Meissn. ex Spreng. 1827 = Diplasiolejeunea pellucida (Meissn. ex Spreng.) 

Schiffn. 1893 (1) [Excludenda(1)]. 

Jungermannia perloi (Gola) Váňa KEN(2), REU(1). 

Jungermannia phyllorhiza Nees 1833 = Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. 1986 (133). 

Jungermannia piligera Nees 1824 = Anastrophyllum piligerum (Nees) Steph. 1893 (163). 

Jungermannia pinguis L. 1753 = Aneura pinguis (L.) Dumort. 1831 (351, 652). 

Jungermannia pocsii Váňa KEN(2), SAF(2), TAN(2). 

Jungermannia podophylla Thunb. 1800 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846 (956). 

Jungermannia polycarpa Nees 1833 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (832). 

Jungermannia praerupta Reinw. et al. 1824 = Bazzania praerupta (Reinw. et al.) Trevis. 1877 (1). 

Jungermannia procumbens Hook.f. et Taylor 1845 = Symphyogyna brasiliensis Nees et Mont. 1836 (970). 

Jungermannia pterota Hook.f. et Taylor 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle 1988 

(930). 

Jungermannia pumila With. LES(75), SAF(75), TAN(2), UGA(776). 

Jungermannia purpurascens (Steph.) Besch. 1895 = Jamesoniella purpurascens Steph. 1891 (318). 

Jungermannia pusilla L. 1753 = Fossombronia pusilla (L.) Nees 1838 (992). 

Jungermannia pusilla Lehm. = Fossombronia crispa Nees 1846 (82). 

Jungermannia refracta Brid. ex Gottsche 1857, nom. inval. = Plagiochila paucidentata Mont. et Gottsche 

1856 (1010, 1). 

Jungermannia rehmannii Steph. 1892 = Jamesoniella purpurascens (Steph.) Steph. 1901 (318). 

Jungermannia renauldii Steph. MDG(1), REU(1), UGA(2,96). 

Jungermannia repanda Schwägr. 1814 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (843, 1). 

Jungermannia reptans L. 1753 = Lepidozia reptans (L.) Dumort. 1835 (992). 

Jungermannia retusa Reinw. et al. 1824 = Thysananthus retusus (Reinw. et al.) B.Thiers et Gradst. 1989 (1) 


Jungermannia rotalis Hook.f. et Taylor 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 

1997 (55). 

Jungermannia rotundistipula Lindenb. ex Lehm. 1829 = Leucolejeunea rotundistipula (Lindenb. ex Lehm.) 

A.Evans 1907 (806). 

Jungermannia sarmentosa Lehm. et Lindenb. 1835 = Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb. 

1844 (722). 

Jungermannia scariosa Lehm. 1829 = Gongylanthus scariosus (Lehm.) Steph. 1906 (170). 

Jungermannia schizopleura Spruce 1885 = Gottschelia schizopleura (Spruce) Grolle 1968 (901).


Jungermannia semiteres Lehm. 1829 = Lophocolea semiteres (Lehm.) Mitt. 1877 (313). 

Jungermannia serrulata Sw. 1788 = Isotachis serrulata (Sw.) Gottsche 1864 (1) [Excludenda(1)]. 

Jungermannia simplex Sw. 1788 = Plagiochila simplex (Sw.) Lindenb. 1840 (1) [Excludenda(1)]. 

Jungermannia spathulistipa Reinw. et al. 1824 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845 

(972). 

Jungermannia sphaerocarpa Hook. CMN(2), LES(75), MDG(1), MLW(2), REU(1), RWA(2), 

SAF(75), TAN(2), UGA(2,42), ZAI(2), ZIM(2). 

Jungermannia sphaerophora Lehm. et Lindenb. 1833 = Lopholejeunea sphaerophora (Lehm. et Lindenb.) 

Steph. 1912 (839). 

Jungermannia sphagnoides Schwägr. 1814 = Schistochila sphagnoides (Schwägr.) Steph. 1909 (871) = 

Gottschea sphagnoides (Schwägr.) Lindb. 1870 (954). 

Jungermannia sphagnoides sensu Richard ex Hook. 1818, hom. illeg. [non Schwägr. 1814] = Pleurozia gigantea 

(F.Weber) Lindb. 1874 (1012). 

Jungermannia splendens Lehm. et Lindenb. 1832 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 

1984 (623). 

Jungermannia squarrosa Reinw. et al. 1824, hom. illeg. [non Hook 1818] = Frullania ericoides (Nees) 

Mont. 1839 (920, 147). 

Jungermannia stolonifera Sieber ex Nees 1831, hom. illeg. [non Sw. 1788] = Bazzania longa (Nees) Trevis. 

1877 (1) [Excludenda(1)]. 

Jungermannia stolonifera (Steph.) Steph. 1901, hom. illeg. [non Sw. 1788] = Jungermannia borgenii Gottsche 

ex Pearson 1893 (412). 

Jungermannia stolonifera Sw. 1788 = Bazzania stolonifera (Sw.) Trevis. 1877 (1) [Excludenda(1)]. 

Jungermannia striata Lehm. et Lindenb. 1832 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (158, 

972). 

Jungermannia subaequifolia Nees et Mont. 1844 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 

(895). 

Jungermannia subcomplicata Lehm. et Lindenb. 1838 = Anastrophyllum subcomplicatum (Lehm. et Lindenb.) 

Steph. 1901 (163). 

Jungermannia subfusca Nees 1830 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (839). 

Jungermannia subintegerrima Reinw. et al. 1824 = Syzygiella subintegerrima (Reinw. et al.) Spruce 1895 

(1) [Excludenda(1)]. 

Jungermannia subintegra Hook.f. et Taylor 1845 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (314, 

97). 

Jungermannia subtilissima (Schiffn.) Grolle 1964. CVI(125). [Excludenda]. A record from the Cape Verde Islands 

(125) is based on a misidentification (37). 

Jungermannia superba Nees ex Spreng. 1827 = Plagiochila superba (Nees ex Spreng.) Mont. et Nees 1839 


Jungermannia tabularis Spreng. 1827 = Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell 1963 (864) = 

Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (969). 

Jungermannia tamarisci L. 1753 = Frullania tamarisci (L.) Dumort. 1835 (147). 

Jungermannia tenuis Reinw. et al. 1824, hom. illeg. [non Ehrh. 1789] = Drepanolejeunea angustifolia 

(Mitt.) Grolle 1965 (1) [Excludenda(1)]. 

Jungermannia tenuissima Lehm. et Lindenb. 1845 = Cephaloziella tenuissima (Lehm. et Lindenb.) Steph. 

1893 (803). 

Jungermannia thouarsii Hook. 1818 = Gottschea sphagnoides (Schwägr.) Lindb. 1870 (885, 954). 

Jungermannia tortuosa Lehm. et Lindenb. 1832 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (836). 

Jungermannia trichophylla L. 1753 = Blepharostoma trichophyllum (L.) Dumort. 1835 (942). 

Jungermannia trifaria Reinw. et al. 1824 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 (1042). 

Jungermannia trinervis Lehm. 1835 = Frullania trinervis (Lehm.) Drège 1843 (147). 

Jungermannia trinitensis Lehm. et Lindenb. 1833 = Bryopteris filicina (Sw.) Nees 1845 (122) [Excludenda(1)]. 

Jungermannia tuberculata Lehm. 1835 = Frullania ericoides (Nees) Mont. 1839 (147). 

Jungermannia ulicina Taylor 1841 = Microlejeunea ulicina (Taylor) A.Evans 1902 (984). 

Jungermannia ulothrix Schwägr. 1814 = Lepidozia ulothrix (Schwägr.) Lindenb. in Gottsche et al. 1845 

(382). 

Jungermannia unciformis Hook.f. et Taylor 1844 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864 (465). 

Jungermannia uncinata F.Weber 1815 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Jungermannia vallis-gratiae Gottsche ex Steph. 1908 = Cephalozia bicuspidata (L.) Dumort. 1835 (494). 

Jungermannia vermicularis Lehm. 1829 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (97). 

Jungermannia verrucosa (Steph.) Steph. 1901 = Jungermannia dusenii (Steph.) Steph. 1901 (412). 

Jungermannia vitrea Nees 1830 = Leptolejeunea vitrea (Nees) Schiffn. 1893 (984). 

Jungermannia volkensii Steph. 1917 = Jungermannia borgenii Gottsche ex Pearson 1893 (412).


Jungermannia wondraczekii Corda in Sturm 1830 = Fossombronia wondraczekii (Corda) Dumort. ex Lindb. 

1873 (992). 

Jungermannia xanthocarpa Lehm. et Lindenb. 1833 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907 (430). 

Kantius bidentulus (F.Weber) Trevis. 1877 [Kantia auct.] = Calypogeia bidentula (F.Weber) Nees 1845 

(253). 

Kantius heteromorphus (Lehm.) Steph. 1888 [Kantia auct.] = Jungermannia heteromorpha Lehm. 1829 

(986) = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969 (844). 

Kantius microstipulus Steph. 1895 [Kantia auct.] = Calypogeia microstipula (Steph.) Steph. 1908 (253). 

Kantius muelleriana Schiffn. 1900 [Kantia auct.] = Calypogeia muelleriana (Schiffn.) Müll.Frib. 1908 

(992). 

Kurzia capillaris (Sw.) Grolle subsp. capillaris MDG(4,720), MLW(2,92), RWA(2), SAF(2), 

SEY(4), TAN(2). 

Kurzia capillaris subsp. capillaris var. verrucosa (Steph.) Pócs TAN(2). 

Kurzia capillaris subsp. stephanii (Renauld ex Steph.) Pócs MAU(1), MDG(1,4,720), MLW(2), 

REU(1,4), TAN(2). 

Kurzia irregularis (Steph.) Grolle KEN(759), RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2). 

Kurzia nemoides (Hook.f. et Taylor) Grolle STH(2,49). 

Kurzia stephanii (Renauld ex Steph.) Grolle 1964 = Kurzia capillaris subsp. stephanii (Renauld ex Steph.) 

Pócs 1984 (382). 

Kurzia tabularis (Steph.) Grolle 1964 = Kurzia capillaris (Sw.) Grolle 1964 (382). 

Kymatocalyx africanus Váňa et Wigginton MLW(74), TAN(74). 

Kymatocalyx dominicensis (Spruce) Váňa MDG(74), UGA(776). 

Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa COM(4,74), MAU(1,4,74), 

MDG(1,4,74,720), REU(1,4,74), TAN(2). 

Kymatocalyx rhizomatica (Herzog) Gradst. et Váňa MDG(720). 

Leioscyphus borbonicus Steph. 1892 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (471). 

Leioscyphus gottscheanus (Lindenb. ex Lehm.) Steph. 1906 = Leptoscyphus expansus (Lehm.) Grolle 1963 

(471, 812). 

Leioscyphus infuscatus Mitt. 1886 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (471). 

Leioscyphus iversenii Pearson 1887 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Leioscyphus motleyi Mitt. 1886 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (471). 

Leioscyphus repens Mitt. 1863, hom. illeg. [non Mitt. in Hooker 1854] = Leptoscyphus infuscatus (Mitt.) 

E.W.Jones 1963 (471, 973). 

Lejeunea aberrans Lindenb. et Gottsche 1845 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988 

(542). 

Lejeunea abbreviata Mitt. 1863 = Archilejeunea abbreviata (Mitt.) Vanden Berghen 1951 (907). 

Lejeunea abnormis (Gottsche) Steph. 1891 [non (Herzog) R.M.Schust.1962] = Phragmicoma abnormis 

Gottsche 1882 (1) [Dubia(1)]. 

Lejeunea abortiva Mitt. 1879 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122). 

Lejeunea abyssinica (Gola) Cufod. ETH(2). 

Lejeunea acuta Mitt. ANN(2,33), BIO(2,33,72), BUR(2), CMN(2), COM(1), KEN(2,731), RIM(10), 

RWA(2), SAO(2), TAN(2), UGA(50), ZAI(667). 

Lejeunea acutiloba (Hook.f. et Taylor) Gottsche et al. 1845 = Phragmicoma acutiloba (Hook.f. et Taylor) 

Gottsche et al. 1847 = Marchesinia brachiata (Sw.) Schiffn. 1893 (831). 

Lejeunea adhaesiva Mitt. 1886 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963 (654, 912). 

Lejeunea aethiopica E.W.Jones ETH(2). 

Lejeunea africana (Steph.) Steph. 1891 = Microlejeunea africana Steph. 1888 (986). 

Lejeunea alata (Steph.) Steph. 1893, hom. illeg. [non Gottsche 1845] = Archilejeunea alata Steph. 1893 

(828). 

Lejeunea alata Gottsche var. alata COM(1), MAS(1), MAU(1), MDG(1,720), REU(1), SEY(1), 

TAN(2). 

Lejeunea albifolia Taylor 1845 = Diplasiolejeunea albifolia (Taylor) E.W.Jones 1954 (857). 

Lejeunea alata var. patriciae Pócs MDG(709,720). 

Lejeunea aloba Sande Lac. KEN(759), ZAI(2). 

Lejeunea amaniensis E.W.Jones CMN(2), MDG(1), RWA(2), TAN(2). 

Lejeunea amentulifera Steph. UGA/ZAI(2). 

Lejeunea amplectens (Steph.) Steph. 1890 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (35). 

Lejeunea angulifolia Mitt. ROD(1). 

Lejeunea angustifolia Mitt. 1861 = Drepanolejeunea angustifolia (Mitt.) Grolle 1965 (1) [Excludenda(1)].


Lejeunea arnelliana R.M.Schust. 1965 = Lejeunea villaumei (Steph.) Grolle 1977 (828). 

Lejeunea ascensionis (Hook.f. et Taylor) Gottsche et al. 1845 = Cheilolejeunea ascensionis (Hook.f. et 

Taylor) Grolle 1988 (930). 

Lejeunea asperifolia Steph. ZAI(2). 

Lejeunea astroidea Mitt. 1860 = Leptolejeunea astroidea (Mitt.) Steph. 1913 (99). 

Lejeunea atra Mitt. 1886 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988 (542). 

Lejeunea aulacophora (Mont.) Steph. 1890 = Acrolejeunea aulacophora (Mont.) Steph. 1895 (35). 

Lejeunea autoica R.M.Schust. BIO(2,33), CMN(2), GHA(2), TAN(2). 

Lejeunea balfourii Mitt. 1879 = Stictolejeunea balfourii (Mitt.) E.W.Jones 1976 (872). 

Lejeunea barbiflora Lindenb. et Gottsche 1851 = Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans 

1908 (1031). 

Lejeunea belangeri Gottsche ex Ångstr. 1874, nom. inval. = Ceratolejeunea belangeriana (Gottsche) Steph. 

1913 (1). 

Lejeunea belangeriana Gottsche 1845 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 (1). 

Lejeunea borbonica Hampe et Gottsche 1895, nom. inval. = Lopholejeunea borbonica Steph. 1896 (1). 

Lejeunea borgenii Steph. 1890 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (35, 833). 

Lejeunea brachytoma Gottsche 1882 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 (847). 

Lejeunea brenanii E.W.Jones CMN(2), GHA(2), RIM(10), SIE(2). 

Lejeunea breutelii (Steph.) Steph. 1901, hom. illeg. [non Gottsche 1845] = Lejeunea caespitosa Lindenb. 

1845 (806). 

Lejeunea brevifissa Gottsche 1882 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963 (878). 

Lejeunea brittoniae (A.Evans) Grolle 1995 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Lejeunea byssoides Gottsche 1882 = Microlejeunea byssoides (Gottsche) Pearson 1892 (1) [Dubia(1)]. 

Lejeunea caespitosa Lindenb. ANG(2), ANN(33), BIO(2,33), BUR(2), CAF(2), CMN(2), COM(1), 

CON(2), CVI(2,37), GHA(2), IVO(2,53), KEN(2,71,731), MAU(1), MDG(720), MLW(563), 

NGA(2), PRI(2), REU(1), RIM(10), ROD(723), RWA(2), SAF(2), SAO(2), SEY(1,4,108), SIE(2), 

TAN(2), UGA(2,42,96), ZAI(2), ZIM(2). 

Lejeunea calyptrifolia (Hook.) Spreng. 1827, hom. illeg. [non Dumort. 1822] = Colura calyptrifolia (Hook.) 

Dumort. 1835 (336). 

Lejeunea calyptrifolia Dumort. 1822, nom. nud. = Colura calyptrifolia (Hook.) Dumort. 1835 (170). 

Lejeunea camerunensis (Steph.) E.W.Jones 1972 = Lejeunea tuberculosa Steph. 1915 (720). 

Lejeunea cancellata Nees et Mont. 1842 MAS(1). [Excludenda(1)]. 

Lejeunea cantabrigiensis E.W.Jones CMN(2). 

Lejeunea capensis Gottsche KEN(2), LES(75), MDG(1), MLW(2), RWA(2), SAF(2), TAN(2), 

ZAI(2). 

Lejeunea capensis (S.W.Arnell) R.M.Schust. 1963, hom. illeg. [non Gottsche 1845] = Ciliolejeunea capensis 

S.W.Arnell 1953 (912) = Lejeunea villaumei (Steph.) Grolle 1977 (828). 

Lejeunea (Inflatolejeunea) capensis (S.W.Arnell) R.M.Schust. 1963 = Inflatolejeunea capensis S.W.Arnell 

1953 (912) = Lejeunea capensis Gottsche 1845 (170). 

Lejeunea capulata Taylor 1846 = Drepanolejeunea capulata (Taylor) Steph. 1913 (106). 

Lejeunea cardiantha Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (886). 

Lejeunea cardiocarpa Mont. 1842 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902 (495). 

Lejeunea cardiophylla Gottsche 1882 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (579). 

Lejeunea cavifolia (Ehrh.) Lindb. SOC(2,764). 

Lejeunea cavifolia Steph. 1886, hom. illeg. [non (Ehrh.) Lindb. 1875] = Diplasiolejeunea vandenberghenii 

Grolle 1960 (888) = Diplasiolejeunea cavifolia Steph. 1895 (900, 940). 

Lejeunea chrysophylla (Lehm.) Gottsche et al. 1845 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 

(542). 

Lejeunea compressistipula (Pearson) Demaret 1942 perhaps = Lejeunea confusa E.W.Jones 1972 (157). 

Lejeunea confertissima (Steph.) Steph. 1892 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 

1975 (35). 

Lejeunea conformis Nees et Mont. 1843 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Lejeunea confusa E.W.Jones CAF(2), CMN(2), COM(1), GAB(2), GHA(2), KEN(2), MDG(720), 

MLW(563), NGA(2), RIM(10), RWA(2), SEY(1), SIE(2), TAN(2), UGA(50,96), ZAI(2). 

Lejeunea contigua (Nees) Mont. 1840 = Pycnolejeunea contigua (Nees) Grolle 1979 (14). 

Lejeunea convexa S.W.Arnell 1953 = Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell 1955 (878). 

Lejeunea corbieri Steph. 1915 perhaps = Lejeunea ibadana A.J.Harr. et E.W.Jones 1982 (25). 

Lejeunea cornuta (Lindenb.) Nees 1838 = Ceratolejeunea cornuta (Lindenb.) Steph. 1895 (1) [Excludenda(1)]. 

Lejeunea cornutissima Steph. 1892 perhaps = Ceratolejeunea diversicornua (Steph.) Steph. 1913 (733).


Lejeunea crenatiflora (Steph.) Steph. 1891 ("crenatifolia") = Cololejeunea obliqua (Nees et Mont.) Schiffn. 

1897 (914). See note under Cololejeunea crenatiflora. 

Lejeunea cristata (Steph.) Steph. 1891 = Drepanolejeunea cristata Steph. 1891 (986) = Drepanolejeunea 

cultrella (Mitt.) Steph. 1913 (649) 

Lejeunea cucullata (Reinw. et al.) Nees 1838 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995 (1). 

Lejeunea cuculliloba (Steph.) Steph. 1891 = Lejeunea caespitosa Lindenb. 1845 (157). 

Lejeunea cultrella Mitt. 1863 = Drepanolejeunea cultrella (Mitt.) Steph. 1913 (649). 

Lejeunea cuneata (Lehm. et Lindenb.) Gottsche et al. 1845 = Cololejeunea cuneata (Lehm. et Lindenb.) 

Herzog 1947 (152). 

Lejeunea cuneifolia (Steph.) Steph. Nov-Dec 1892 [non Spruce 1884] = Cololejeunea cuneifolia Steph. Jul- 

Aug 1892 (986). 

Lejeunea cupulirostris Augier CMN(2). 

Lejeunea cyathearum E.W.Jones COM(1,4), MLW(2), RWA(2), TAN(2), UGA(776), ZAI(2). 

Lejeunea decursiva Sande Lac. 1855 = Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. 1963 (828). 

Lejeunea delessertii Nees et Mont. 1843 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 1984 (582). 

Lejeunea deplanata Mitt. 1886, hom. illeg. [non Nees 1845] = Prionolejeunea grata (Gottsche) Schiffn. 

1893 (847). 

Lejeunea digitalis Mitt. 1886 = Colura digitalis (Mitt.) Steph. 1916 (336). 

Lejeunea dipterocarpa E.W.Jones CMN(2), NGA(2). 

Lejeunea dispar (Jovet-Ast) R.M.Schust. 1962 = Microlejeunea dispar Jovet-Ast 1959 (1033). 

Lejeunea diversicornua (Steph.) Steph. 1891 = Ceratolejeunea diversicornua (Steph.) Steph. 1913 (986). 

Lejeunea duncaniae (Sim) S.W.Arnell "duncanii" SAF(2). 

Lejeunea dusenii (Steph.) Steph. 1891 [non (Steph.) Steph. 1892] = Taxilejeunea dusenii Steph. 1891 (989) 

= Lejeunea ramosissima Steph. 1886 (863). 

Lejeunea dusenii (Steph.) Steph. 1892, hom. illeg. [non (Steph.) Steph. 1981] = Colura dusenii (Steph.) 

Steph. 1916 (986). 

Lejeunea ecarinata (Steph.) Steph. MDG(1), REU(1). 

Lejeunea eckloniana Lindenb. ANG(2), CMN(2), CON(2), CVI(2,37), DJI(2), GHA(2), GUI(2), 

KEN(2,21,71), LES(75), MAU(1), MDG(1,720), MLW(2,563), NGA(2,127), REU(1), ROD(1,723), 

RWA(2), SAF(2,75), SIE(2), TAN(2), TOG(2,101), UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Lejeunea ecklonii Lindenb. ex Steph. 1890, nom. inval. = Lejeunea eckloniana Lindenb. 1845 (869). 

Lejeunea elegans (Steph.) Steph. Nov-Dec 1891, hom. illeg. [non Gottsche 1845] = Cololejeunea elegans 

Steph. 1891 (986). 

Lejeunea elliptica (Lehm. et Lindenb.) Mont. 1840 = Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. 

1893 (430). 

Lejeunea elobulata (Sim) S.W.Arnell, nom. inval. SAF(2). 

Lejeunea emergens (Mitt.) Steph. 1894 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Lejeunea epiphylla Mitt. 1861 = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (130). 

Lejeunea epiphyta (Steph.) Steph. 1891, hom. illeg. [non Gottsche 1845] = Taxilejeunea epiphyta Steph. 

1891 (989) = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Lejeunea eplicata Steph. 1915 = Lejeunea ramosissima Steph. 1887 (25). 

Lejeunea eulopha Taylor 1846 = Lopholejeunea eulopha (Taylor) Schiffn. 1893 (122). 

Lejeunea excavata Mitt. 1860 = Marchesinia excavata (Mitt.) Schiffn. 1893 (891). 

Lejeunea expansa (Steph.) Steph. 1915 = Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell 1963 (864) = 

Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (969). 

Lejeunea filicaulis (Steph.) Steph. Nov-Dec 1892 = Cololejeunea hildebrandii (Austin) Steph. 1897 (135). 

Lejeunea filiformis (Sw.) Nees ex Mont. 1839 = Omphalanthus filiformis (Sw.) Nees 1845 (1) [Excludenda(1)]. 

Lejeunea fimbriata Gottsche 1880 = Lopholejeunea eulopha (Taylor) Schiffn. 1893 (972, 578). 

Lejeunea flava (Sw.) Nees CAF(2), COM(1), CON(2), CVI(37), GHA(2), KEN(71,731), MDG(1), 

RWA(2), SEY(1,4,108), SIE(2), TAN(2), TOG(101), UGA(2,96), ZAI(2). 

Lejeunea flava agg. BIO(72), ETH(2), KEN(2), SAO(2), ZAI(2), ZAM(2). 

Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell 1963 = Lejeunea tabularis (Spreng.) Gottsche et al. 

1845 (969). 

Lejeunea flava var. convexiuscula Pearson 1886 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (806). 

Lejeunea flavovirens Ångstr. COM(1), MAU(1), MDG(1,720), REU(1), RWA(2), SIE(2), TAN(2), 

UGA(2,50), ZAI(2). 

Lejeunea fulva (Gottsche) Steph. 1890, hom. illeg. [non Spruce 1884] = Phragmicoma fulva Gottsche 1882 

(35) = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (831). 

Lejeunea furva Mitt. 1878 = Lejeunea eckloniana Lindenb. 1845 (869). 

Lejeunea fusca (Lehm.) Drège 1843 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118).


Lejeunea galliotii Steph. 1915 probably = Lejeunea eckloniana Lindenb. 1845 (869). 

Lejeunea gaudichaudii (Gottsche) Steph. 1894 [non Lindenb. 1845] = Bryopteris gaudichaudii Gottsche 

1857 (890). 

Lejeunea gracillima Mitt. 1861 = Microlejeunea ulicina (Taylor) A.Evans 1902 (986). 

Lejeunea grandistipula Steph. 1886 = Euosmolejeunea brachytoma (Gottsche) Steph. 1914 (867) = Cheilolejeunea 

trifaria (Reinw. et al.) Mizut. 1964 (847). 

Lejeunea grata Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Lejeunea grossecristata (Steph.) E.W.Jones ANN(2,33), BIO(2,33), CMN(2), GHA(2), IVO(2), 

KEN(2), NGA(2), RWA(2), SAO(2), SIE(2), TAN(2), UGA(2,96), ZAI(2). 

Lejeunea hamatifolia (Hook.) Spreng. 1893 = Drepanolejeunea hamatifolia (Hook.) Schiffn. 1895 (170). 

Lejeunea hanningtonii Mitt. 1886 = Caudalejeunea hanningtonii (Mitt.) Schiffn. 1893 (891). 

Lejeunea haskarliana (Gottsche) Spruce 1884 "hasskarliana", comb. illeg. = Schiffneriolejeunea tumida var 

haskarliana (Gottsche) Gradst. et Terken 1981 (1038). 

Lejeunea haskarliana Lehm. et Lindenb. in Lehmann 1844 = Cololejeunea haskarliana (Lehm. et Lindenb.) 

Schiffn. 1898 (28). 

Lejeunea helenae Pearson ANG(2), ETH(2), KEN(2), MLW(2), RWA(2), SAF(2,75), TAN(2), 

UGA(2,96), ZAI(2), ZIM(2). 

Lejeunea hepaticola (Steph.) Steph. KEN(2,731,759), MLW(563), TAN(2), UGA(115), ZAI(2). 

Lejeunea hildebrandii Austin 1876 = Cololejeunea hildebrandii (Austin) Steph. 1897 (135). 

Lejeunea holtii Spruce 1887 ANN(2), UGA(2). [Dubia(2)]. A European and Macaronesian species. Arnell's records 

from Annobón and Uganda are likely to represent other species. 

Lejeunea hyalinomarginata Nees ex Mont. 1842 = Cololejeunea marginata (Lehm. et Lindenb.) Schiffn. 

1898 (1). 

Lejeunea ibadana A.J.Harr. et E.W.Jones CAF(2), GHA(2), IVO(2), NGA(2), RIM(10), SIE(2), 

UGA(2). 

Lejeunea inflexa (Gottsche) Steph. 1891, hom. illeg. [non Lehm. et Lindenb. 1838] = Phragmicoma inflexa 

Gottsche 1882 (1) [Dubia(1)]. 

Lejeunea intermedia Lindenb. 1845 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (976). 

Lejeunea intertexta Lindenb. 1845 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897 (574). 

Lejeunea intorta Besch. et Spruce 1890 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 (1). 

Lejeunea isomorpha Gottsche 1882 = Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell 1963 (864) = 

Lejeunea tabularis (Spreng.) Gottsche et al. 1845 (969). However, Jones 1968 (106) pointed out that the names 

L. isomorpha and L. isophylla have been confused. Vanden Berghen 1972 (158) considered that plants from southern 

Africa named as L. isomorpha might often represent L. isophylla. 

Lejeunea isophylla E.W.Jones BUR(2), COM(1), IVO(2), KEN(2,731), MDG(1), MLW(2,563), 

MOZ(2), RWA(2), SAF(2), SEY(1), TAN(2), UGA(50,96), ZAI(2), ZIM(2). 

Lejeunea julacea Steph. MDG(1). 

Lejeunea jungneri (Steph.) Steph. 1901 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Lejeunea kamerunensis (Steph.) Vanden Berghen 1972 = Microlejeunea kamerunensis Steph. 1915 (25). 

Lejeunea kilimandjarensis Steph. TAN(2). 

Lejeunea kindbergii Steph. 1892 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Lejeunea krakakammae Lindenb. 1845 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963 (878). 

Lejeunea kraussiana Lindenb. 1845 = Diplasiolejeunea kraussiana (Lindenb.) Steph. 1895 (850). 

Lejeunea kurzii (Steph.) Steph. 1890 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897 (574). 

Lejeunea lamacerina (Steph.) Schiffn. CVI(2,37). 

Lejeunea lehmanniana Gottsche 1845 = Caudalejeunea lehmanniana (Gottsche) A.Evans 1907 (430). 

Lejeunea letabaensis S.W.Arnell 1954 = Lejeunea rhodesiae (Sim) R.M.Schust. 1962 (868). 

Lejeunea leucosis Besch. et Spruce MDG(1). 

Lejeunea lineata (Lehm. et Lindenb.) Nees 1838 = Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans 1906 

(1) [Excludenda(1)]. 

Lejeunea lomana E.W.Jones SIE(2), UGA(115). 

Lejeunea longifissa Steph. 1915 perhaps = Lejeunea caespitosa Lindb. 1845 (157). 

Lejeunea longirostris (Steph.) E.W.Jones 1968, hom. illeg. [non Spruce 1884] = Lejeunea grossecristata 

(Steph.) E.W.Jones 1968 (874). 

Lejeunea lunulata (F.Weber) Nees 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (837). 

Lejeunea lyratiflora Steph. BUR(2), CMN(2), RWA(2), TAN(2), UGA(50). 

Lejeunea maculata Mitt. 1861 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Lejeunea madagascariensis Nees et Mont. 1836 = Porella madagascariensis (Nees et Mont.) Trevis. 1877 

(1). 

Lejeunea madagascariensis Gottsche 1882, hom. illeg. [non Nees et Mont. 1836] = Acanthocoleus madagascariensis 

(Steph.) Kruijt 1988 (542).


Lejeunea marginata (Lehm. et Lindenb.) Gottsche et al. 1845 = Cololejeunea marginata (Lehm. et Lindenb.) 

Schiffn. 1898 (135). 

Lejeunea mariana Gottsche 1845 = Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 1989 (578) [Excludenda(1)]. 

Lejeunea mascarena Steph. 1890 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 (847). 

Lejeunea mauritiana Steph. 1890 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913 (1). 

Lejeunea microloba Gola 1920 perhaps = Lejeunea wilmsii (Steph.) Steph. 1915 [= Eulejeunea wilmsii 

Steph. 1892] (158) = Lejeunea eckloniana Lindenb. 1845 (869). 

Lejeunea microlobulata S.W.Arnell 1953 = (Eu)Lejeunea wilmsii (Steph.) Steph. 1915 (170) = Lejeunea 

eckloniana Lindenb. 1845 (869). 

Lejeunea microscopica (Taylor) Gottsche et al. 1845 = Aphanolejeunea microscopica (Taylor) A.Evans 

1911 (453). 

Lejeunea minutissima Dumort. 1822, nom. inval. = Cololejeunea minutissima (Sm.) Schiffn. 1893 (430). 

Lejeunea minutissima (Sm.) Spreng. 1827 = Cololejeunea minutissima (Sm.) Schiffn. 1893 (1). 

Lejeunea mitracalyx (Eifrig) Mizut. 1970 = Lejeunea alata Gottsche 1845 (828). 

Lejeunea montagnei Gottsche 1843 = Euosmolejeunea montagnei (Gottsche) Steph. (855) = Cheilolejeunea 

montagnei (Gottsche) R.M.Schust. 1963 (912). 

Lejeunea multilacera (Steph.) Steph. Dec 1890 = Lopholejeunea multilacera Steph. Nov 1890 (986). 

Lejeunea myriocarpa Nees et Mont. 1842 = Cololejeunea minutissima subsp. myriocarpa (Nees et Mont.) 

R.M.Schust. 1955 (936). 

Lejeunea nigra Steph. 1891 = Mastigolejeunea nigra (Steph.) Steph. 1895 (986). 

Lejeunea nigricans Lindenb. 1845 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122). 

Lejeunea nobilis Gottsche 1882 = Archilejeunea nobilis (Gottsche) Steph. 1911 (1). 

Lejeunea obliqua Nees et Mont. 1843 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (1). 

Lejeunea oblongistipula Gottsche 1882 = Microlejeunea oblongistipula (Gottsche) Steph. 1915 (1). 

Lejeunea obtusa (Steph.) Steph. 1891 = Colura obtusa (Steph.) Steph. 1916 (986) = Colura digitalis (Mitt.) 

Steph. 1916 (336). 

Lejeunea obtusata Gottsche GHA(2), MAU(1), MDG(1), SIE(2), TAN(2), UGA(2,96). 

Lejeunea obtusistipula (Steph.) E.W.Jones GUI(2), SIE(2). 

Lejeunea (subgen. Acrolejeunea) occulta Steph. 1888 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974 

(832). 

Lejeunea ocellifera (S.W.Arnell) R.M.Schust. 1955, nom. inval. = Microlejeunea ocellifera S.W.Arnell 

1953 (829) = Microlejeunea africana Steph. 1888 (865). 

Lejeunea okomuensis E.W.Jones CMN(2), IVO(2), NGA(2), UGA(42). 

Lejeunea pallidissima Gola ZAI(2). 

Lejeunea papilionacea Steph. CMN(2,670), COM(1,670), GHA(2,670), GUI(670), MDG(1,670), 

REU(670), RIM(10), SIE(2,670). 

Lejeunea pappeana (Nees) Mitt. 1863 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (833). 

Lejeunea papulosa Gottsche 1882 = Microlejeunea papulosa (Gottsche) Pearson 1892 (1). 

Lejeunea parisii Steph. 1915 = Lejeunea caespitosa Lindenb. 1845 (157). 

Lejeunea parviloba Steph. Dec 1890, hom. illeg. [non Ångstr. 1877] = Schiffneriolejeunea parviloba 

(Steph.) Gradst. 1974 (832). 

Lejeunea pellucida (Meissn. ex Spreng.) Gottsche et al. 1845 = Diplasiolejeunea pellucida (Meissn. ex 

Spreng.) Schiffn. 1893 (1) [Excludenda(1)]. 

Lejeunea perforata Schiffn. 1886 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Lejeunea phyllobola Nees et Mont. COM(1), KEN(2), MDG(4), SAF(2), SEY(4), SIE(2), TAN(2), 

ZAI(2). Lejeunea brittoniae was placed in synonymy with L. phyllobola by Reiner-Drehwald 2000 (168), and records of 

the former species in Africa have accordingly been transferred in this checklist. However, Africa is not included in the 

stated distribution of L. phyllobola in (168), and the African material should be reassessed. 

Lejeunea physaefolia Gottsche 1882 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 (1). 

Lejeunea planiuscula Mitt. 1861 = Archilejeunea planiuscula (Mitt.) Steph. 1911 (578). 

Lejeunea platyneura Spruce 1884 = Cololejeunea platyneura (Spruce) A.Evans 1902 (1106). 

Lejeunea pluriplicata Pearson 1887 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 1980 (878). 

Lejeunea polyandra Gola 1920 = Lejeunea wilmsii (Steph.) Steph. 1915 [= Eulejeunea wilmsii Steph. 1892] 

(158) = Lejeunea eckloniana Lindenb. 1845 (869). 

Lejeunea pterota (Hook.f. et Taylor) Gottsche et al. 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) 

Grolle 1988 (930). 

Lejeunea pulcherrima Steph. 1886 = Lejeunea acuta Mitt. 1863 (140). 

Lejeunea punctata Gottsche 1882 = Cololejeunea punctata (Gottsche) Pearson 1892 (1) [Dubia(1)]. 

Lejeunea pycnoclada (Taylor) Mitt. 1861 = Acrolejeunea pycnoclada (Taylor) Schiffn. 1893 (35). 

Lejeunea quintasii Steph. 1891 = Leptolejeunea quintasii (Steph.) Steph. 1913 (986) = Leptolejeunea epiphylla 

(Mitt.) Steph. 1913 (130). See also note under Leptolejeunea quintasii.


Lejeunea ramosissima Steph. ANN(2,33), BIO(2,33), CMN(2), CON(2), GUI(2), PRI(2), REU(1), 

RWA(2), SAO(2), SEY(1), SIE(2), TAN(2), TOG(2), UGA(96), ZAI(2). 

Lejeunea recurvistipula Gottsche 1845 = Caudalejeunea recurvistipula (Gottsche) Schiffn. 1893 (1). 

Lejeunea renauldii Renauld et Cardot 1891, nom. illeg. = Acrolejeunea aulacophora (Mont.) Steph. 1895 

(35). 

Lejeunea renauldii Steph. Dec 1890 = Lejeunea alata Gottsche 1845 (828). 

Lejeunea repanda Schiffn 1886 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Lejeunea rhodesiae (Sim) R.M.Schust. ANG(2), CAF(2), CON(2), KEN(21), LES(75), MLW(2), 

ROD(723), RWA(2), SAF(2), SOC(746), TAN(2), UGA(2,50), ZAI(2), ZAM(2), ZIM(2). 

Lejeunea rodriguezii (Steph.) Steph. MAU(1), MDG(1), REU(1). 

Lejeunea rotalis (Hook.f. et Taylor) Gottsche et al. 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. 

et Geissler 1997 (55). 

Lejeunea rotundistipula (Lindenb. ex Lehm.) Mont. 1842 [non (Steph.) S.Hatt. 1952] = Leucolejeunea rotundistipula 

(Lindenb. ex Lehm.) A.Evans 1907 (1). 

Lejeunea rufescens Lindenb. 1845 = Cheilolejeunea rufescens (Lindenb.) Grolle 1982 (1). 

Lejeunea rutenbergiana Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Lejeunea rutenbergii Gottsche ex Steph. 1891, nom. inval. = Prionolejeunea grata (Gottsche) Schiffn. 1893 

(1). 

Lejeunea saccatiloba (Steph.) Steph. Nov-Dec 1891 = Lejeunea caespitosa Lindenb. 1845 (157). Jones 1972 

stated that most plants Stephani named as L. saccatiloba are L. confusa, but that the type is L. caespitosa. 

Lejeunea sagraeana (Mont.) Gottsche et al. 1845 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (122). 

Lejeunea scabriflora Gottsche 1882, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 

(135). 

Lejeunea securifolia Gottsche 1882 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 (1). 

Lejeunea serpentina Mitt. 1861 = Cheilolejeunea serpentina (Mitt.) Mizut. 1963 (574). 

Lejeunea serrula Mitt. 1886 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Lejeunea setacea (Steph.) Steph. BEN(22), BUR(2), CAF(2), CMN(2), COM(1), CON(2), GHA(2), 

IVO(2), KEN(2), NGA(2), PRI(2), SAF(2), SAO(2), SIE(2), ZAI(2), ZAM(2). 

Lejeunea sieberiana Gottsche 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (837). 

Lejeunea sikorae (Steph.) Steph. 1894 = Taxilejeunea sikorae Steph. 1893 (1). 

Lejeunea silvestris Gottsche 1882 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (872). 

Lejeunea sinuata Mitt. 1886 = Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. 1913 (977). 

Lejeunea solitaria Gottsche 1882 = Harpalejeunea solitaria (Gottsche) Steph. 1913 (1) [Dubia(1)]. 

Lejeunea spathulistipa (Reinw. et al.) Mitt. 1861 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 

1845 (972). 

Lejeunea sphaerophora (Lehm. et Lindenb.) Gottsche et al. 1845 = Lopholejeunea sphaerophora (Lehm. et 

Lindenb.) Steph. 1912 (839). 

Lejeunea squarrosa (Steph.) Steph. TAN(2). 

Lejeunea stephanii Spruce ex Steph. 1892, nom. inval. = Caudalejeunea reniloba (Gottsche) Steph. 1912 (1) 


Lejeunea striata (Lehm. et Lindenb.) Steph. 1890 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (1). 

Lejeunea subciliata Mitt. 1878 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 (152). 

Lejeunea subfusca (Nees) Nees et Mont. 1836 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (1). 

Lejeunea subinermis (Steph.) Steph. 1901 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (839). 

Lejeunea subintegra (Hook.f. et Taylor) Gottsche et al. 1845 = Clasmatocolea vermicularis (Lehm.) Grolle 

1960 (314, 97). 

Lejeunea surrepens Mitt. 1878 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (872). 

Lejeunea tabularis (Spreng.) Gottsche et al. BIO(2,33), BUR(2), CAF(2), CMN(2), COM(1), 

CON(2), GUI(2), IVO(2), KEN(2,731), LES(75), MAU(108), MDG(1,720), MLW(2,563), MOZ(2), 

NGA(2), PRI(2), RWA(2), SAF(2,75), TAN(2), UGA(2,42,96), ZAI(2). 

Lejeunea tenuis (Nees) Gottsche et al. 1845 [non (Ehrh.) Gottsche et al. 1845] = Drepanolejeunea angustifolia 

(Mitt.) Grolle 1965 (1) [Excludenda(1)]. 

Lejeunea thomeensis Steph. 1886 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Lejeunea thwaitesiana Mitt. 1861 = Drepanolejeunea thwaitesiana (Mitt.) Steph. 1913 (1) [Excludenda(1)]. 

Lejeunea tortuosa Lehm. et Lindenb. 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (836, 837). 

Lejeunea trematodes Nees 1836 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Lejeunea trichomanoides Mont. et Nees 1845 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Lejeunea trifaria (Reinw. et al.) Gottsche et al. 1845 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964 

(1048). 

Lejeunea trinitensis Lindenb. COM(4). 

Lejeunea triquetra Mitt. 1863 = Thysananthus spathulistipus (Reinw. et al.) Gottsche et al. 1845 (25).


Lejeunea tristis Steph. 1890 = Frullanoides tristis (Steph.) van Slageren 1985 (795). 

Lejeunea truncatiloba Steph. 1892 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (161). 

Lejeunea tuberculosa Steph. ANG(2), ANN(2,33), CAF(2), CMN(2), GHA(2), GUI(2), IVO(2), 

KEN(2), MAU(1), MDG(1,720), MLW(2,563), MOZ(2), NGA(2), REU(1), SEY(1,4), SIE(2), 

TAN(2), UGA(96), ZIM(2). 

Lejeunea turgida Steph. 1892, hom. illeg. [non Mitt. 1861] = Mastigolejeunea turgida Steph. 1912 (986). 

Lejeunea ulicina (Taylor) Gottsche et al. 1945 = Microlejeunea ulicina (Taylor) A.Evans 1902 (986). 

Lejeunea ulicina subsp. africana (Steph.) Vanden Berghen 1972 = Microlejeunea africana Steph. 1888 (25). 

Lejeunea ulicina subsp. ocellifera (S.W.Arnell) R.M.Schust. 1962 = Microlejeunea africana Steph. 1888 

(865). 

Lejeunea unciloba Lindenb. 1845 = Leucolejeunea unciloba (Lindenb.) A.Evans 1907 (430). 

Lejeunea variabilis Lindenb. 1845 = Ceratolejeunea variabilis (Lindenb.) Schiffn. 1893 (1). 

Lejeunea villaumei (Steph.) Grolle KEN(2), MDG(1,720), MLW(563), REU(1), RWA(2), SAF(2), 

SEY(1), TAN(2), UGA(776). 

Lejeunea vogelii Taylor 1846 = Lejeunea caespitosa Lindenb. 1845 (157). 

Lejeunea vulgariformis Gottsche MDG(1). 

Lejeunea wilmsii (Steph.) Steph. 1915 [= Eulejeunea wilmsii Steph. 1892] = Lejeunea eckloniana Lindenb. 

1845 (869). 

Lejeunea xanthocarpa (Lehm. et Lindenb.) Gottsche et al. 1845 = Leucolejeunea xanthocarpa (Lehm. et 

Lindenb.) A.Evans 1907 (430). 

Lejeunea sp. (Hygrolejeunea patellirostris Steph. 1896). H. patellirostris was treated as synonymous with 

Lejeunea acuta Mitt. in Tixier 1995 (130). However, E.W.Jones (in litt.) previously commented that although a specimen 

under the name H. patellirostris in Hb. Stephani (G) is L. acuta, it is not the type, "nor does it agree well with the description 

of the type". 

Lejeunea sp. (Hygrolejeunea staudtiana Steph.) BIO(2), CMN(2), UGA(2). 

Lejeunea sp. (Prionolejeunea corbisieri Pearson) ZAI(2). 

Lembidium borbonicum Steph. 1893 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (1). 

Lembidium heteromorphum (Lehm.) Steph. 1891 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969 

(844). 

Leperoma ochroleuca (Spreng.) Mitt. 1877 = Lepicolea ochroleuca (Spreng.) Spruce 1885 (170). 

Lepicolea ochroleuca (Spreng.) Spruce SAF(2,107). 

Lepidolejeunea bidentula (Steph.) R.M.Schust. var. bidentula MDG(1,582), SEY(1,4,108,582). 

Lepidolejeunea delessertii (Nees et Mont.) Grolle MAU(1,582), REU(1,4,582). 

Lepidolejeunea serrulata (Steph.) Grolle CMN(2,582), GHA(2,582), RIM(10). 

Lepidozia abyssinica S.W.Arnell 1960 = Lepidozia stuhlmannii Steph. 1895 (92) 

Lepidozia africana Steph. MAU(1), MDG(1), REU(1,4), TAN(2). 

Lepidozia bicuspidata C.Massal. 1885 = Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle 1964 

(821). 

Lepidozia capillaris (Sw.) Lindenb. 1845 = Kurzia capillaris (Sw.) Grolle 1964 (382). 

Lepidozia carnosa Steph. 1911 = Lepidozia stuhlmannii var. carnosa (Steph.) Pócs (96). 

Lepidozia cupressina (Sw.) Lindenb. KEN(2), LES(75), RWA(2), SAF(2), TAN(2), UGA(2,96), 

ZAI(2). 

Lepidozia cupressina subsp. natalensis (Steph.) Pócs SAF(2), TAN(2). 

Lepidozia cupressina subsp. quinquefida (Steph.) Pócs TAN(2), ZAI(2). 

Lepidozia hexiloba Pearson GHA(2). 

Lepidozia hyalina Steph. 1911 = Lepidozia pearsonii Spruce 1891 (382). 

Lepidozia irregularis Steph. 1911 = Kurzia irregularis (Steph.) Grolle 1982 (382). 

Lepidozia lacerata Steph. 1909 = Lepidozia stuhlmannii Steph. 1895 (889). 

Lepidozia laevifolia (Hook.f. et Taylor) Gottsche et al. STH(2). 

Lepidozia laurentii Steph. 1909 = Sprucella succida (Mitt.) Steph. 1891 (902) = Lepidozia succida Mitt. 

1860 (250). 

Lepidozia longitexta Steph. 1909 = Sprucella succida (Mitt.) Steph. 1891 (902) = Lepidozia succida Mitt. 

1860 (250). 

Lepidozia microphylla var. ß (Hook.) Lindenb. in Gottsche et al. 1845 [quoad pl. Madagascar] = Lepidozia 

africana Steph. 1922 (382). 

Lepidozia natalensis Steph. 1909 = Lepidozia cupressina subsp. natalensis (Steph.) Pócs 1984 (382). 

Lepidozia nematodes (Gottsche ex Austin) Spruce 1885 = Telaranea nematodes (Gottsche ex Austin) 

M.Howe 1902 (382). 

Lepidozia nemoides (Hook.f. et Taylor) Lindenb. et Gottsche 1846 = Kurzia nemoides (Hook.f. et Taylor) 

Grolle 1964 (382).


Lepidozia pearsonii Spruce MLW(2), SAF(2), UGA(2), ZAI(2). 

Lepidozia pearsonii var. lacerata (Steph.) Pócs 1984 = Lepidozia lacerata Steph. 1909 (382) = Lepidozia 

stuhlmannii Steph. 1895 (889). 

Lepidozia pearsonii var. pulvinata (Steph.) Pócs 1984 = Lepidozia stuhlmannii subsp. pulvinata (Steph.) 

Pócs 1994 (889). 

Lepidozia procumbens Mitt. 1875 = Lepidozia laevifolia (Hook.f. et Taylor) Gottsche et al. 1845 (382). 

Lepidozia pulvinata Steph. 1911 = Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs 1994 (889). 

Lepidozia quinquefida Steph. 1911 = Lepidozia cupressina subsp. quinquefida (Steph.) Pócs 1984 (382). 

Lepidozia quintasii Steph. 1922 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902 (382). 

Lepidozia redacta Steph. 1911 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902 (382). 

Lepidozia reducta Steph. 1922, a typographical error for Lepidozia redacta Steph. 1911 [= Telaranea 

nematodes (Gottsche ex Austin) M.Howe 1902]. 

Lepidozia reptans (L.) Dumort. ZAI(2). 

Lepidozia spinosa S.W.Arnell 1954 = Psiloclada clandestina Mitt. 1855 (382). 

Lepidozia stephanii Renauld ex Steph. 1890 = Kurzia capillaris subsp. stephanii (Renauld) Pócs 1984 (382). 

Lepidozia stuhlmannii Steph. MLW(92), UGA(96). 

Lepidozia stuhlmannii var. abyssinica (S.W.Arnell) Pócs 1994 = Lepidozia stuhlmannii Steph. 1895 var. 

stuhlmannii (92). 

Lepidozia stuhlmannii var. carnosa (Steph.) Pócs KEN(7), UGA(7,96), ZAI(274). 

Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs KEN(2), RWA(2), UGA(2), ZAI(2). 

Lepidozia stuhlmannii Steph. subsp. stuhlmannii BIO(36), CMN(2), ETH(2), KEN(2), MLW(2), 

RWA(2), SAF(2), TAN(2), UGA(2,42,50), ZAI(2). 

Lepidozia succida Mitt. ANG(2), BIO(2,33,36), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), 

GUI(2), IVO(2), MDG(1,4), NGA(2), RIM(10), SAO(2), SIE(2), UGA(2,42,96), ZAI(2). 

Lepidozia ubangiensis Steph. CAF(2), CON(2), GAB(2), ZAI(2). Treated as a subspecies (Lepidozia succida 

subsp. ubangiensis) by Vanden Berghen 1983 (250), but accorded species status by Pócs 1994b (889). 

Lepidozia succulenta Sim 1926 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001 (556). 

Lepidozia tabularis Steph. 1909 = Kurzia capillaris (Sw.) Grolle 1964 (382). 

Lepidozia trifida Steph. 1911 = Telaranea trifida (Steph.) R.M.Schust. 1966 (382). 

Lepidozia truncatella Nees 1845 = Lepidozia cupressina (Sw.) Lindenb. 1845 (804, 382). 

Lepidozia ulothrix (Schwägr.) Lindenb. MAU(1). 

Lepidozia verrucosa Steph. 1885 = Kurzia capillaris var. verrucosa (Steph.) Pócs 1984 (382). 

Leptocolea adhaesiva (Mitt.) E.W.Jones 1953 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963 (912). 

Leptocolea africana (Steph.) E.W.Jones 1953 = Cololejeunea africana (Steph.) R.M.Schust. 1963 (912). 

Leptocolea apiculata E.W.Jones 1954 = Cololejeunea apiculata (E.W.Jones) R.M.Schust. 1963 (912). 

Leptocolea appressa A.Evans 1912 = Cololejeunea appressa (A.Evans) Benedix 1953 (106). 

Leptocolea auriculata E.W.Jones 1953 = Cololejeunea auriculata (E.W.Jones) R.M.Schust. 1963 (912). 

Leptocolea autoica (Steph.) E.W.Jones 1953, nom. inval. = Cololejeunea autoica (Steph.) Grolle 1995 (1). 

Leptocolea bolombensis (Steph.) E.W.Jones 1953 = Cololejeunea bolombensis (Steph.) Vanden Berghen 

1972 (495) = Cololejeunea tonkinensis Steph. 1895 (135). 

Leptocolea cardiocarpa (Mont.) A.Evans 1911 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902 (495). 

Leptocolea crenatiflora Steph. 1915 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (914). See note 

under Cololejeunea crenatiflora. 

Leptocolea crenatifolia Steph. ex S.W.Arnell 1965, nom. inval. = Cololejeunea obliqua (Nees et Mont.) 

Schiffn. 1897 (914). 

Leptocolea cristata (Steph.) E.W.Jones 1953 = Cololejeunea cristata (Steph.) R.M.Schust. 1963 (152). 

Leptocolea cristata var. lanciloba S.W.Arnell 1963 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902 

(1060, 135). 

Leptocolea cristata var. occidentalis E.W.Jones 1953 = Cololejeunea occidentalis (E.W.Jones) Vanden Berghen 

1978 (842). 

Leptocolea cuneifolia (Steph.) A.Evans 1911 = Cololejeunea cuneifolia Steph. 1892 (25). 

Leptocolea dentata E.W.Jones 1953 = Cololejeunea dentata (E.W.Jones) R.M.Schust. 1963 (912). 

Leptocolea distalopapillata E.W.Jones 1957 = Cololejeunea distalopapillata (E.W.Jones) R.M.Schust. 1963 

(912). 

Leptocolea furcilobulata Berrie et E.W.Jones 1954 = Cololejeunea furcilobulata (Berrie et E.W.Jones) 

R.M.Schust. 1963 (912). 

Leptocolea himalayensis Pandé et Misra 1943 = Cololejeunea latilobula (Herzog) Tixier 1985 (135). E.W. 

Jones (1037) preferred to maintain C. himalayensis as a distinct species. 

Leptocolea latilobula Herzog 1930 = Cololejeunea latilobula (Herzog) Tixier 1985 (135). 

Leptocolea leloutrei E.W.Jones 1953 = Cololejeunea leloutrei (E.W.Jones) R.M.Schust. 1963 (912).


Leptocolea mascarena Steph. ex S.W.Arnell 1965, nom. inval. = Leptolejeunea maculata (Mitt.) Schiffn. 

1898 (924). 

Leptocolea montagnei (Gottsche) Steph. 1916 = Cheilolejeunea montagnei (Gottsche) Schiffn. 1898 (1). 

Leptocolea nigerica E.W.Jones 1953 = Cololejeunea nigerica (E.W.Jones) R.M.Schust. 1963 (912). 

Leptocolea nymanii Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (135). 

Leptocolea perforata Schiffn. ex S.W.Arnell 1965, nom. inval. = Drepanolejeunea trematodes (Nees) 

Bischl. 1968 (924). 

Leptocolea perforata sensu S.W.Arnell 1965 [non Schiffn. ex S.W.Arnell 1965, nom. inval.] = Drepanolejeunea 

madagascariensis (Steph.) Grolle 1974 (924). 

Leptocolea punctata E.W.Jones 1957 = Cololejeunea vidaliana Tixier 1973 (847). 

Leptocolea quintasii var mascarenica S.W.Arnell 1965, nom. inval. = Leptolejeunea epiphylla (Mitt.) Steph. 

1913 (136). 

Leptocolea scabrifolia var. africana Herzog 1947, nom. inval. = Cololejeunea obliqua (Nees et Mont.) 

Schiffn. 1897 (1). 

Leptocolea subciliata (Mitt.) E.W.Jones 1953 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 

(152). 

Leptocolea tonkinensis (Steph.) Steph. 1916 = Cololejeunea tonkinensis Steph. 1895 (135). 

Leptolejeunea adhaesiva (Mitt.) Steph. 1895 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963 (912). 

Leptolejeunea astroidea (Mitt.) Steph. BUR(2), CMN(2), CON(2), GHA(2), GNB(2), GUI(2), 

IVO(2), NGA(2), RIM(10), SIE(2), UGA(42,50,115), ZAI(2), ZIM(2). 

Leptolejeunea convexistipa Bischl. 1969 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Leptolejeunea degelii S.W.Arnell 1962 = Diplasiolejeunea symoensii Vanden Berghen 1960 (495). 

Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. COM(1). 

Leptolejeunea epiphylla (Mitt.) Steph. CMN(2), COM(1), CON(2), GHA(2), GUI(2), IVO(2), 

KEN(2), MAU(1,4), MDG(720), NGA(2), RIM(10), SAO(2), SEY(4,108), SIE(2), TAN(2,736), 

UGA(42), ZAI(2). 

Leptolejeunea maculata (Mitt.) Schiffn. CMN(2), COM(1,4), GAB(2), GHA(2), GUI(2), IVO(2), 

KEN(2,731), MAU(1), NGA(2), RIM(10), ROD(723), SAO(2), SEY(1,4,108), SIE(2), TAN(2), 

TOG(2), UGA(42,50,96), ZAI(2). 

Leptolejeunea madagascariensis Steph. 1913 = Drepanolejeunea madagascariensis (Steph.) Grolle 1974 

(924). 

Leptolejeunea mascarena Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Leptolejeunea papuliflora Steph. 1913 = Leptolejeunea quintasii Steph. 1891 (99) = Leptolejeunea epiphylla 

(Mitt.) Steph. 1913 (130). 

Leptolejeunea perforata (Schiffn.) Schiffn. 1897 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Leptolejeunea quintasii (Steph.) Steph. 1913 = Leptolejeunea epiphylla (Mitt.) Steph. 1913 (130). Vanden 

Berghen (733) considered that L. epiphylla is perhaps synonymous with L. quintasii, and Tixier (130) treated them as 

synonymous without further comment. 

Leptolejeunea quintasii var. mascarenica S.W.Arnell 1965, nom. inval. = Leptolejeunea epiphylla (Mitt.) 

Steph. 1913 (136). 

Leptolejeunea repanda (Schiffn.) Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Leptolejeunea symoensii Vanden Berghen 1961 = Drepanolejeunea symoensii (Vanden Berghen) Grolle 

1976 (924). 

Leptolejeunea thomeensis (Steph.) Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898 (924). 

Leptolejeunea trematodes (Nees) Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Leptolejeunea triocellata Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968 (924). 

Leptolejeunea truncatiflora Steph. 1911 = Leptolejeunea astroidea (Mitt.) Steph. 1913 (99). 

Leptolejeunea truncatiloba (Steph.) Steph. 1896 = Leptolejeunea thomeensis (Steph.) Steph. (161) = Leptolejeunea 

maculata (Mitt.) Schiffn. 1898 (924). 

Leptolejeunea vitrea (Nees) Schiffn. MAS, with doubt(1). 

Leptoscyphus bewsii Sim 1926 = Lophocolea bewsii (Sim) Grolle 1959 (471). 

Leptoscyphus diversifolius (Gottsche) Grolle SAF(2). 

Leptoscyphus expansus (Lehm.) Grolle KEN(731), LES(75), MDG(1), MLW(29), RWA(2), SAF(2), 

ZAI(2), ZIM(2). 

Leptoscyphus gottscheanus (Lindenb. ex Lehm.) Gottsche 1858 = L. expansus (Lehm.) Grolle 1963 (812, 

471). 

Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. KEN(2,731), RWA(2), TAN(2), UGA(2,96), 

ZAI(2). 

Leptoscyphus henkelii Sim 1926 = Heteroscyphus grandistipus subsp. henkelii (Sim) Grolle 1963 (471). 

Leptoscyphus infuscatus (Mitt.) E.W.Jones BIO(2,33,36), CMN(2), ETH(2), MDG(720), 

MLW(2,29), REU(1,4), RWA(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2).


Leptoscyphus iversenii (Pearson) Sim 1926 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Leptoscyphus leightonii Sim 1926 = Lophocolea newtonii Steph. 1907 (812) = Lophocolea congoana Steph. 

in Renauld 1902/3 (313) = Lophocolea martiana Nees 1845 (974). 

Leptoscyphus stephensii Sim 1926 = Lophocolea cuspidata (Nees) Limpr. 1876 (812) = Lophocolea bidentata 

(L.) Dumort. 1835 (975). 

Lethocolea congesta (Lehm.) S.W.Arnell COM(1,4), KEN(2), LES(75), REU(1,4), RWA(2), 

SAF(2), TAN(2), UGA(2), ZAI(2). 

Leucolejeunea capensis S.W.Arnell 1953 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 

(806, 867). 

Leucolejeunea knysnana S.W.Arnell 1953 = Leucolejeunea unciloba (Lindenb.) A.Evans 1907 (867). 

Leucolejeunea rotalis (Hook.f. et Taylor) S.W.Arnell 1961 = Aureolejeunea rotalis (Hook.f. et Taylor) 

Gradst. et Geissler 1997 (55). 

Leucolejeunea rotundistipula (Lindenb. ex Lehm.) Steph. LES(75), MAU(1), MLW(563), REU(1), 

SAF(2,75), ZIM(2). 

Leucolejeunea unciloba (Lindenb.) A.Evans KEN(731), SAF(2), TAN(2), UGA(776). 

Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans BIO(2,33), COM(4), ETH(2), 

KEN(2,731), MDG(4), MLW(563), MOZ(2), REU(1,4), RWA(2), SAF(2), STH(2), TAN(2), 

UGA(2,42), ZAI(2), ZIM(2). 

Lindigina renifolia Mitt. 1877 = Gongylanthus renifolius (Mitt.) Steph. 1906 (170). 

Lindigina scariosa (Lehm.) Mitt. 1877 = Gongylanthus scariosus (Lehm.) Steph. 1906 (170). 

Lophocolea amanica Steph. 1922 = Lophocolea congoana Steph. in Renauld 1902/3 (313) = Lophocolea 

martiana Nees 1845 (974). 

Lophocolea angustistipula Steph. STH(887). Overlooked in (2) and in edition 1 of this checklist. 

Lophocolea armatistipula Steph. 1911 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (165). 

Lophocolea ascensionis Steph. ASC(2). 

Lophocolea bewsii (Sim) Grolle SAF(2). 

Lophocolea bidentata (L.) Dumort. BIO(2,33,36), BUR(2), CMN(2), CVI(2,37), ETH(2), 

KEN(2,21,731), MAU(1), MDG(1,720), MLW(2,29), REU(1), RWA(2), SAF(2), STH(49), 

TAN(2), UGA(2,42,96), ZAI(2). 

Lophocolea bidentata sensu Sim 1926 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis 

Nees 1845 (313). 

Lophocolea bidentata var. δ capensis Gottsche et al. 1845 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea 

difformis Nees 1845 (313). 

Lophocolea borbonica Steph. 1890 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 

1845 (313). 

Lophocolea cagnii Gola 1907 = Lophocolea difformis Nees 1845 (814). 

Lophocolea cambouena Steph. 1907 = Lophocolea subrotunda Mitt. 1879 (165) = Lophocolea concreta 

Mont. 1845 (313). 

Lophocolea concreta Mont. ANN(2,33), BUR(2), GHA(2), KEN(2), LES(75), MAU(1), 

MDG(1,720), MLW(2,29), NGA(2), REU(1), ROD(1,723), RWA(2), SAF(2,75), SIE(2), SWZ(2), 

TAN(2), TOG(2), UGA(42,50), ZAI(2), ZAM(2), ZIM(2). 

Lophocolea congoana Steph. 1905 = Lophocolea martiana Nees 1845 (974). 

Lophocolea connata (Sw.) Mont. et Nees 1839 MAU(1). [Excludenda(1)]. 

Lophocolea cuspidata (Nees) Limpr. 1876 = Lophocolea bidentata (L.) Dumort. 1835 (975). 

Lophocolea devexa Mitt. 1863 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898 (916). 

Lophocolea difformis Nees BIO(2,33,36), CAF(2), CMN(2), ETH(2), GHA(2), IVO(51), 

KEN(2,731), LES(75), MAU(4), MDG(720), MLW(2,29), NGA(2), REU(1), RWA(2), SAF(2,75), 

SAO(2), SIE(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZIM(2). 

Lophocolea diversifolia Gottsche 1845 = Leptoscyphus diversifolius (Gottsche) Grolle 1963 (471). 

Lophocolea diversifolia sensu Sim 1926 [non Gottsche 1845] = Lophocolea moelleri Steph. 1886 (165) = 

Lophocolea difformis Nees 1845 (313). 

Lophocolea dusenii Steph. 1907 = Lophocolea congoana Steph. 1905 [= Lophocolea martiana Nees 1845] 

or Lophocolea opposita Mitt. 1879 [= Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839] (165). 

Lophocolea esterhuysenii S.W.Arnell ex E.W.Jones 1953 = Lophocolea bewsii (Sim) Grolle 1959 (313). 

Lophocolea expansa (Lehm.) Drège 1843 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Lophocolea fasciculata (Nees) S.W.Arnell et Grolle 1959 = Clasmatocolea fasciculata (Nees) Grolle 1960 

(314, 97). 

Lophocolea fragrans (Moris et De Not.) Gottsche et al. COM(1), MLW(29), REU(1), SAF(2), 

TAN(2), ZAI(2). 

Lophocolea frappieri Steph. 1907 = Lophocolea fragrans (Moris et De Not.) Gottsche et al. 1845 (313).


Lophocolea hians Steph. 1901 perhaps = Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839 (165). 

Lophocolea humistrata (Hook.f. et Taylor) Gottsche et al. STH(2). 

Lophocolea inflata Steph. 1911 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (165). 

Lophocolea integrifolia Steph. 1892 = Leptoscyphus expansus (Lehm.) Grolle 1963 (1). 

Lophocolea lepervanchei Steph. REU(1). 

Lophocolea lillieana Steph. KEN(2). 

Lophocolea longifolia Steph. 1890 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 

1845 (313). 

Lophocolea longispica Steph. 1892 = Lophocolea cuspidata (Nees) Limpr. 1876 (165) = Lophocolea 

bidentata (L.) Dumort. 1835 (975). 

Lophocolea lucida (Spreng. ex Lehm.) Mont. CMN(2), COM(1), MDG(1,720), MLW(2,29), 

REU(1), ROD(1), SAF(2), SEY(1), TAN(2), UGA(96), ZAI(2), ZAM(2). 

Lophocolea macleana Steph. 1907 = Lophocolea moelleri Steph. 1886 (165) = Lophocolea difformis Nees 

1845 (313). 

Lophocolea madagascariensis Gottsche MDG(1). 

Lophocolea magnistipula S.W.Arnell 1954 = Lophocolea semiteres (Lehm.) Mitt. 1877 (313). 

Lophocolea martiana Nees ANG(2), BIO(36,72), BUR(2), CAF(2), CMN(2), CON(2), GAB(2), 

GHA(2), IVO(2,53), MAU(1), MDG(1,720), MLW(29), NGA(2), PRI(2), REU(1), RIM(10), 

RWA(2), SAF(2), SEY(1,4), SIE(2), SWZ(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZIM(2). 

Lophocolea minor Nees ?REU(1). The record needs verification (1). 

Lophocolea moelleri Steph. 1886 = Lophocolea difformis Nees 1845 (313). 

Lophocolea mollis (Nees) Gottsche et al. The distribution of this taxon is given only as "îles austro-africaines de 

l'Indien Océan". It is close to L. difformis and may be conspecific (Grolle 1995 (1)). 

Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs KEN(2), TAN(2), UGA(2). 

Lophocolea muricata (Lehm.) Nees BIO(33,36,72), BUR(2), CMN(2), COM(1), ETH(2), 

MDG(1,720), MLW(29), MOZ(2), REU(1), RWA(2), SAF(2), TAN(2), UGA(2,42,50,96), ZAI(2), 

ZIM(2). 

Lophocolea newtonii Steph. 1907 = Lophocolea congoana Steph. in Renauld 1902/3 (313) = Lophocolea 

martiana Nees 1845 (974). 

Lophocolea obscura Steph. 1907 = Lophocolea opposita Mitt. 1879 (165) = Lophocolea lucida (Lehm.) 

Mont. 1839 (812). 

Lophocolea occidentalis Steph. 1922 = Heteroscyphus dubius (Gottsche) Schiffn. 1910 (165). 

Lophocolea onraedtii Grolle MDG(1), REU(1). 

Lophocolea opposita Mitt. 1879 = Lophocolea lucida (Lehm.) Mont. 1839 (812). 

Lophocolea ovistipula Herzog 1952 = Lophocolea subintegra (Hook.f. et Taylor) Grolle 1959 (313) = Clasmatocolea 

vermicularis (Lehm.) Grolle 1960 (314). 

Lophocolea rehmannii Steph. 1892 = Lophocolea subrotunda Mitt. 1879 (165) = Lophocolea concreta 

Mont. 1845 (313). 

Lophocolea rubescens Steph. 1890 = Lophocolea opposita Mitt. 1879 (165) = Lophocolea lucida (Lehm.) 

Mont. 1839 (812). 

Lophocolea scott-elliottii Steph. 1907 = Lophocolea cuspidata (Nees) Limpr. 1876 (165) = Lophocolea 

bidentata (L.) Dumort. 1835 (975). 

Lophocolea semiteres (Lehm.) Mitt. SAF(2). 

Lophocolea setacea sensu Sim 1926 [non Steph. 1892] = Lophocolea moelleri Steph. 1886 (165) = Lophocolea 

difformis Nees 1845 (313). 

Lophocolea setacea Steph. 1892 = Lophocolea cuspidata (Nees) Limpr. 1876 (165) = Lophocolea bidentata 

(L.) Dumort. 1835 (975). 

Lophocolea silvestris Gottsche 1882 MAD(1). [Dubia(1)]. 

Lophocolea spiniflora Steph. 1907 = Lophocolea muricata (Lehm.) Nees 1845 (313). 

Lophocolea staudtiana Steph. 1907 = Lophocolea congoana Steph. 1905 (165) = Lophocolea martiana Nees 

1845 (974). 

Lophocolea subintegra (Hook.f. et Taylor) Grolle 1959 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 

(314, 97). 

Lophocolea subrotunda Mitt. 1879 = Lophocolea concreta Mont. 1845 (313). 

Lophocolea subulistipa Herzog 1952 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (314, 97). 

Lophocolea sylvestris Gottsche ex Steph. 1907, nom. inval. = Lophocolea silvestris Gottsche 1882 (1) [Dubia(1)]. 

Lophocolea vermicularis Herzog 1952 = Lophocolea fasciculata (Nees ) S.W.Arnell et Grolle in Grolle 

1959 (313) = Clasmatocolea fasciculata (Nees) Grolle 1960 (97). 

Lopholejeunea abbreviata (Mitt.) Steph. 1912 = Archilejeunea abbreviata (Mitt.) Vanden Berghen 1951 

(907).


Lopholejeunea abortiva (Mitt.) Steph. 1912 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122). 

Lopholejeunea abortiva var. alata Vanden Berghen 1984 = Lopholejeunea nigricans (Lindenb.) Schiffn. 

1898 (1). 

Lopholejeunea abortiva var. fragilis (Steph.) Vanden Berghen 1984 = Lopholejeunea nigricans (Lindenb.) 

Schiffn. 1898 (122). 

Lopholejeunea angustifolia Steph. 1912 = Lopholejeunea eulopha (Taylor) Schiffn. 1893 (841). 

Lopholejeunea atra (Mitt.) Steph. 1895 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988 (542). 

Lopholejeunea boivinii Steph. 1912 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (839). 

Lopholejeunea borbonica Steph. COM(1), MDG(1,720), REU(1), TAN(2). 

Lopholejeunea camerunensis Steph. 1911, nom. nud = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 

(839, 122). 

Lopholejeunea erugata B.Thiers MDG(1). 

Lopholejeunea eulopha (Taylor) Schiffn. COM(1), MAU(1), MDG(1,720), REU(1), RWA(2), 

SEY(1), TAN(2). 

Lopholejeunea fimbriata (Gottsche) Schiffn. 1893 = Lopholejeunea eulopha (Taylor) Schiffn. 1893 (972, 

578). 

Lopholejeunea fragilis Steph. 1912 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122). 

Lopholejeunea gabonensis Vanden Berghen GAB(2). 

Lopholejeunea grandicrista Steph. MDG(1,720), REU(1). 

Lopholejeunea intermedia (Lindenb.) Steph. 1912 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 

(976) 

Lopholejeunea jonesii Vanden Berghen BIO(2,33), CMN(2), IVO(2), NGA(2), SIE(2). 

Lopholejeunea kilimanjarica E.W.Jones 1957 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (839). 

Lopholejeunea laceriloba Steph. 1923 = Lopholejeunea eulopha (Taylor) Schiffn. 1893 (972, 839). 

Lopholejeunea laciniata E.W.Jones MLW(563), TAN(2), ZIM(2). 

Lopholejeunea leioptera Gyarmati MDG(708,720). 

Lopholejeunea lepidoscypha Kiaer et Pearson MDG(1). 

Lopholejeunea minima Vanden Berghen REU(1). 

Lopholejeunea moenkemeyeri Steph. 1912 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (839). 

Lopholejeunea multilacera Steph. MAU(1), MDG(1), REU(1). 

Lopholejeunea nigricans (Lindenb.) Schiffn. BIO(2,33), BUR(2), CAF(2), CMN(2), CON(2), 

ETH(2), GAB(2), GHA(2), GUI(2), IVO(2,53), MAU(1), MDG(1), MLW(2,563), MOZ(2), 

NGA(2), REU(1), RIM(10), ROD(1,723), RWA(2), SAF(2,75), SEY(1), SIE(2), TAN(2), TOG(2), 

ZAI(2), ZAM(2), ZIM(2). 

Lopholejeunea obtusilacera Herzog CMN(2), IVO(2), NGA(2). 

Lopholejeunea onraedtii Vanden Berghen MDG(1). 

Lopholejeunea palustris (Sim) S.W.Arnell 1953 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122, 

170). 

Lopholejeunea paramultilacera Vanden Berghen REU(1). 

Lopholejeunea quinquecarinata Vanden Berghen MDG(1), REU(1), TAN(2). 

Lopholejeunea revoluta E.W.Jones UGA(2,50). 

Lopholejeunea sagraeana (Mont.) Schiffn. 1893 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (838). 

Lopholejeunea sinuata (Mitt.) Steph. 1912 = Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. 1912 

(977). 

Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. MAU(1). 

Lopholejeunea subfusca (Nees) Schiffn. ANG(2), ANN(2,33), BIO(2,33), CAF(2), CMN(2), 

COM(1), GAB(2), GHA(2), GUI(2), IVO(2,53), KEN(2), MAU(1), MDG(1,720), NGA(2), 

REU(1,2), RIM(10), RWA(2), SAF(2), SEY(1), SIE(2), TAN(2), TOG(2,101), UGA(96,115), 

ZAI(2). 

Lopholejeunea subfusca var. elongata Vanden Berghen SEY(1). 

Lopholejeunea subinermis Steph. 1895 probably = Lopholejeunea abortiva (Mitt.) Steph. 1912 (839) = Lopholejeunea 

nigricans (Lindenb.) Schiffn. 1898 (122). 

Lopholejeunea tixieriana Vanden Berghen MDG(1). 

Lopholejeunea utriculata Steph. MDG(1). 

Lopholejeunea vermoesenii Pearson 1922 probably = Lopholejeunea abortiva (Mitt.) Steph. 1912 (839) = 

Lopholejeunea nigricans (Lindenb.) Schiffn. 1898 (122). 

Lopholejeunea zenkeri Steph. 1912 = Lopholejeunea abortiva (Mitt.) Steph. 1912 (839) = Lopholejeunea 

nigricans (Lindenb.) Schiffn. 1898 (122).


Lophozia aberrans (Nees et Mont.) Steph. 1901 = Andrewsianthus aberrans (Nees et Mont.) Grolle 1963 

(1). 

Lophozia argentina (Steph.) R.M.Schust. LES(75), SAF(2). 

Lophozia capensis S.W.Arnell 1953 = Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust. 1995.(164). 

Lophozia decolorans (Limpr.) Steph. CMN(2), TAN(2), UGA(776), ZAI(2). 

Lophozia hedbergii S.W.Arnell LES(75), UGA(2). 

Lophozia incisa (Schrad.) Dumort. 1835. The plant named as L. incisa in Váňa, Pócs & De Sloover 1979 (156) is 

Lophozia jamesonii (Mont.) R.M.Schust. 2002. 

Lophozia jamesonii (Mont.) R.M.Schust. KEN(2), RWA(2), TAN(2), UGA(2), ZAI(2). 

Lophozia kilimanjarica S.W.Arnell 1956 = Lophozia jamesonii (Mont.) R.M.Schust. 2002 (974). Placed in 

synonymy in Gradstein et al (1983), but Schuster (2002) treated L. kilimanjarica as a distinct species. 

Lophozia kinangopii Gola 1914 = Isotachis aubertii (Schwägr.) Mitt. 1886 (163). 

Lophozia madagascariensis Steph. 1917 = Cylindrocolea madagascariensis (Steph.) R.M.Schust. 1969 

(615). 

Lophozia montaguensis S.W.Arnell 1954 = Lophozia argentina (Steph.) R.M.Schust. 1966 (163). 

Lophozia multiflora Steph. 1917 = Gymnocoleopsis multiflorus (Steph.) R.M.Schust. 1978 (163). 

Lophozia ruwenzorensis S.W.Arnell 1956 = Andrewsianthus bilobus (Mitt.) Grolle 1963 (414, 163). 

Lunularia cruciata (L.) Dumort. ex Lindb. CMN(48), CVI(2), ERI(2), ETH(2,113), KEN(2,71,665), 

MLW(2,114), NAM(2), RWA(2), SAF(2), TAN(2), UGA(2,42,96), ZAI(2), ZIM(2). 

Lunularia vulgaris Raddi 1818, nom. illeg. = Lunularia cruciata (L.) Dumort. ex Lindb. 1822 (141). 

Madotheca abyssinica Nees 1845 = Porella abyssinica (Nees) Trevis. 1877 (861). 

Madotheca capensis Gottsche 1845 = Porella capensis (Gottsche) Mitt. 1886 (861). 

Madotheca cucullistipula (Steph.) Paris 1903 = Porella cucullistipula Steph. 1893 (861). 

Madotheca hoehneliana Steph. 1895 = Porella hoehnelii Steph. 1891 (861) = Porella abyssinica var. 

hoehnelii (Steph.) Pócs 1994 (1002). 

Madotheca katangae Gola 1920 = Porella abyssinica (Nees) Trevis. 1877 (158). 

Madotheca keniana Steph. 1924 = Porella hoehnelii Steph. 1891 (861) = Porella abyssinica var. hoehnelii 

(Steph.) Pócs 1994 (1002). 

Madotheca lobata Steph. 1910 = Porella montantii (Steph.) E.W.Jones 1963 (861). 

Madotheca madagascariensis (Nees et Mont.) Gottsche et al. 1845 = Porella madagascariensis (Nees et 

Mont.) Trevis. 1877 (1). 

Madotheca matongae Herzog 1952 = Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 1955 (472). 

Madotheca mattirolii Gola 1914 = Porella mattirolii (Gola) S.W.Arnell 1960 (861). 

Madotheca montantii Steph. 1910 = Porella montantii (Steph.) E.W.Jones 1963 (861). 

Madotheca nilgheriensis Mont. 1842 = Porella nilgheriensis (Mont.) Trevis. 1877 (1) [Excludenda(1)]. 

Madotheca ovifolia Steph. 1910 = Porella triquetra (Steph.) E.W.Jones 1963 (861). 

Madotheca prolixa Gottsche ex Steph. 1910 = Porella prolixa (Gottsche ex Steph.) E.W.Jones 1963 (861). 

Madotheca schroederi Steph. 1924 = Porella hoehnelii Steph. 1891 (861) = Porella abyssinica var. 

hoehnelii (Steph.) Pócs 1994 (1002). 

Madotheca steudneri Gottsche 1867, nom. inval. = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994 

(885, 1002). 

Madotheca subdentata Mitt. 1863 = Porella subdentata (Mitt.) E.W.Jones 1963 (861). 

Madotheca thomeensis Steph. 1910 = Porella subdentata (Mitt.) E.W.Jones 1963 (861). 

Madotheca triquetra Steph. 1895 = Porella triquetra (Steph.) E.W.Jones 1963 (861). 

Madotheca vallis-gratiae Gottsche ex Steph. 1910 = Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 

1955 (861). 

Madotheca vastifolia Steph. 1910 = Porella triquetra (Steph.) E.W.Jones 1963 (861). 

Mannia androgyna (L. emend. Lindb.) A.Evans CHA(2,3), CVI(2,37), ERI(2), ETH(2), SOC(746). 

Mannia capensis (Steph.) S.W.Arnell NAM(2), SAF(2), TAN(2), ZAM(2), ZIM(2). 

Mannia dichotoma S.W.Arnell 1961, nom. illeg. = Mannia androgyna (L.) A.Evans 1938 (920). 

Mannia capensis var. pallida S.W.Arnell 1963 = Mannia capensis (Steph.) S.W.Arnell 1957 (716) 

Marchantia androgyna L. 1753 = Mannia androgyna (L. emend. Lindb.) A.Evans 1938 (920). 

Marchantia balboi Gola 1914 KEN(120). [Dubia. Indeterminate(132)]. 

Marchantia balboi var. acutisquamata Gerola 1947 ETH(503). [Dubia. Indeterminate(132)]. 

Marchantia berteroana Lehm. et Lindenb. ASC(2), SAF(2), STH(2), ZIM(2). 

Marchantia berteroana var. anactis Gottsche 1846 = Marchantia berteroana Lehm. et Lindenb. 1834 (132). 

Marchantia cagnii Gola 1907 UGA(1149). [Dubia. Indeterminate(132)]. 

Marchantia cengiana Gerola 1947 ETH(503). [Dubia. Indeterminate(132)]. 

Marchantia chevalieri Steph. ex Bonner 1953 = Marchantia debilis Goebel 1915 (132). 

Marchantia contracta Bisch. ex C.Krauss 1846 = Marchantia berteroana Lehm. et Lindenb. 1834 (132). 

Marchantia crassa A.Richard 1834 = Marchantia berteroana Lehm. et Lindenb. 1834 (132).


Marchantia cruciata L. 1753 = Lunularia cruciata (L.) Dumort. ex Lindb. 1822 (421). 

Marchantia debilis Goebel ANG(2), BUR(2), CHA(2), CMN(2), COM(1), CON(2), GAB(2), 

GHA(2), GUI(2), IVO(2), LES(2,75), MDG(1), MLW(2), NGA(2), RWA(2), SAF(2,75), SIE(2), 

SWZ(2), TAN(2), TOG(2), UGA(2,42), ZAI(2), ZAM(2), ZIM(2). 

Marchantia flavescens Steph. ex Bonner 1953 = Marchantia pappeana Lehm. subsp. pappeana 1857 (132). 

Marchantia globosa Brid. ex F.Weber COM(1), MAU(1), REU(1). 

Marchantia hemisphaerica L. 1753 = Reboulia hemisphaerica (L.) Raddi 1818 (421). 

Marchantia hirsuta Sw. 1788 = Dumortiera hirsuta (Sw.) Nees 1824 (426). 

Marchantia keniae Gola 1914 KEN(120). [Dubia. Indeterminate(132)]. 

Marchantia marginata (Nees) Lehm. 1823 [non Steph. ex Bonner 1953] = Asterella marginata (Nees) 

S.W.Arnell 1963 (132, 997). 

Marchantia paleacea Bertol. CVI(2,37), ETH(2), REU(1), SAF(2). 

Marchantia pappeana Lehm. subsp. pappeana ANG(2), BIO(2,33,36), BUR(2), CMN(2), COM(1), 

CON(2), CVI(2,20,37), ETH(2), GHA(2), GUI(2), IVO(2), KEN(2,665,731), LES(2,75), MDG(1), 

MLW(2,114), RWA(2), SAF(2,75), SAO(2), SIE(2), SWZ(2), TAN(2), TOG(2), UGA(2,96), 

ZAI(2), ZAM(2), ZIM(2). 

Marchantia papyracea Gola 1907 UGA(1149). [Dubia. Indeterminate(132)]. 

Marchantia parviloba Steph. 1895 = Marchantia pappeana Lehm. subsp. pappeana 1857 (132). 

Marchantia planiloba Steph. 1886 = Marchantia pappeana Lehm. subsp. pappeana 1857 (132). 

Marchantia planiloba subsp. walteri Burgeff 1943 = Marchantia pappeana Lehm. subsp. pappeana 1857 

(132) 

Marchantia polymorpha L. BUR(2), ETH(2), KEN(2), REU(1), RWA(2), TAN(2), UGA(2), ZAI(2). 

Reported in Frahm, Lindlar et al. 1996 (37) as occurring in the Cape Verde Islands, but the specimen they cite (Chevalier 

H23) is identified in Bischler-Causse 1993 (132) as M. paleacea. 

Marchantia polymorpha subsp. montivagans Bischl. et Boisselier ETH(2), UGA(96). 

Marchantia polymorpha subsp. ruderalis Bischl. et Boisselier SAF(2). 

Marchantia sellae Gola 1907 UGA(1149). [Dubia. Indeterminate(132)]. 

Marchantia spathysii Lindenb. 1829 = Athalamia spathysii (Lindenb.) S.Hatt. 1954 (992). 

Marchantia stephanii Vanden Berghen 1954 = Marchantia pappeana Lehm. subsp. pappeana 1857 (132). 

Marchantia syngenesica Bory 1804 = Asterella syngenesica (Bory) Grolle 1974 (849, 1). 

Marchantia tabularis Nees 1838 = Marchantia berteroana Lehm. et Lindenb. 1834 (1005, 132). 

Marchantia tusii Gola 1914 KEN(120). [Dubia. Indeterminate(132)]. 

Marchantia umbellata Steph. 1895, hom. illeg. [non Scopoli 1772] = Marchantia pappeana Lehm. subsp. 

pappeana 1857 (132). 

Marchantia viridula Lehm. et Lindenb. 1857 = Marchantia globosa Brid. ex F.Weber 1815 (132). 

Marchantia wilmsii Steph. 1892 = Marchantia pappeana Lehm. subsp. pappeana 1857 (1006, 132). 

Marchantia winkleri Steph. ex Bonner 1953 = Marchantia pappeana Lehm. subsp. pappeana 1857 (1006, 

132). 

Marchesinia acutiloba (Hook.f. et Taylor) Trevis. 1877 = Phragmicoma acutiloba (Hook.f. et Taylor) Gottsche 

et al. 1847 = Marchantia brachiata (Sw.) Schiffn. 1893 (831). 

Marchesinia brachiata (Sw.) Schiffn. STH(2). 

Marchesinia chrysophylla (Lehm.) Steph. 1912 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988 (542). 

Marchesinia deslooveri Vanden Berghen RWA(2), TAN(2). Geissler 1997 (126) named this taxon Marchesinia 

cf. acutiloba. 

Marchesinia excavata (Mitt.) Schiffn. ANN(2,33), BIO(2,33,72), CMN(2), GAB(2), GHA(2), 

GUI(2), KEN(2,731), MDG(720), MLW(2), NGA(2), RIM(10), SAO(2), SIE(2), SOC(558), 

TAN(2), TOG(2), UGA(776), ZAI(2), ZAM(2). 

Marchesinia mackaii (Hook.) Gray CVI(124). The collection from Socotra reported in Kürschner 2000 (94) was 

later redetermined as M. excavata (Kürschner 2001 (558)). 

Marchesinia madagassa Steph. BUR(2), CMN(2), COM(4), KEN(2,665,731), MDG(1), 

MLW(2,563), REU(1), SIE(2), TAN(2), ZAI(2), ZIM(2). The (sterile) plant from Mt Kasigau, Kenya reported 

in Bizot & Pócs 1974 (136), is probably a Lopholejeunea (Geissler 1997 (126)). 

Marchesinia microscypha (Hook.f. et Taylor) Trevis. 1877 = Phragmicoma microscypha (Hook.f. et Taylor 

1845) Mitt. 1875 (987, 988) = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 (831). 

Marchesinia moelleriana Pearson 1924 = Marchesinia madagassa Steph. 1912 (969). 

Marchesinia obtusifolia Vanden Berghen 1984 = Marchesinia madagassa Steph. 1912 (1). 

Marchesinia pappeana (Nees) Kuntze 1891 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (833). 

Marsupella africana Steph. ex Bonner 1953 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 

1996 (153). 

Marsupella aurita (Lehm.) Sim 1926 = Anastrophyllum auritum (Lehm.) Steph. 1901 (163). 

Marsupella boeckii var. ruwenzorensis S.W.Arnell 1957 = Nardia arnelliana Grolle 1964 (317).


Marsupella capensis S.W.Arnell 1957 = Gymnomitrion andinum (Herzog) Herzog 1934 (111). 

Marsupella emarginata (Ehrh.) Dumort. RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Marsupella hedbergii S.W.Arnell 1956 = Marsupella africana Steph. ex Bonner 1953 (416) = Apomarsupella 

africana (Steph. ex Bonner) R.M.Schust. 1996 (153). 

Marsupella integra N.Kitag. 1967 = Marsupella subintegra S.W.Arnell 1956 (416). 

Marsupella sparsifolia (Lindb.) Dumort. SAF(2). 

Marsupella subintegra S.W.Arnell MLW(29), REU(1), RWA(2), UGA(2). 

Marsupella subquadrata Steph., in sched. et icones ined. = Marsupella africana Steph. ex Bonner 1953 

(416) = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 (153). 

Marsupidium brevifolium Steph. 1908 = Adelanthus decipiens (Hook.) Mitt. 1864 (446, 465). 

Marsupidium excisum Mitt. 1877 = Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust. 1961 (451). 

Marsupidium limbatum (Steph.) Grolle MAU(1), MLW(114), REU(1), SAF(2), TAN(2). 

Mastigobryum borbonicum Steph. 1885 = Bazzania borbonica Steph. 1894 (870) = Bazzania praerupta 


Mastigobryum boschianum Steph. 1924 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (823). 

Mastigobryum comorense (Steph.) Steph. 1908 = Bazzania comorense Steph. 1891. [B. decrescens complex]. 

(1). 

Mastigobryum convexum Lindenb. 1845 = Bazzania nitida (F.Weber) Grolle 1960 (888, 1). 

Mastigobryum curvidens (Steph.) Steph. 1908 = Bazzania curvidens Steph. 1891. [B. decrescens complex]. 

(1). 

Mastigobryum decrescens (Lehm. et Lindenb.) Gottsche et al. 1845 = Bazzania decrescens (Lehm. et Lindenb.) 

Trevis. 1877 (1). 

Mastigobryum diclados (in Stephani 1909 (885)) - a typographical error for Mastigophora diclados. 

Mastigobryum exile Lindenb. 1845 = Acromastigum exile (Lindenb.) A.Evans 1934 (170). 

Mastigobryum fuscum (Steph.) Besch. 1895 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (884, 

1). 

Mastigobryum integrum (Nees et Mont.) Gottsche et al. 1845 = Bazzania decrescens (Lehm. et Lindenb.) 

Trevis. 1877 (884, 1). 

Mastigobryum lacouturei Steph. 1924 = Bazzania sp. [B. decrescens complex]. (1). 

Mastigobryum laxifolium Steph. 1911 = Bazzania roccatii Gola 1907 (7). 

Mastigobryum madagassum Steph. 1924 = Bazzania sp. [B. decrescens complex]. (1). 

Mastigobryum mascarenum Steph. 1886 = Bazzania mascarenum (Steph.) Herzog 1947. [B. decrescens 

complex]. (1). 

Mastigobryum molleri Steph. 1886 = Bazzania decrescens subsp. molleri (Steph.) E.W.Jones 1975 (870). 

Mastigobryum novae-hollandiae Nees 1845 = Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 (1) [Excludenda(1)]. 

Mastigobryum pellucidum Steph. 1908 = Bazzania decrescens subsp. molleri (Steph.) E.W.Jones 1975 

(870). 

Mastigobryum perrotanum Steph. 1908 [non Steph. 1924] = Bazzania mascarena (Steph.) Herzog 1947 

(823, 1). [B. decrescens complex]. 

Mastigobryum perrotanum Steph. 1924, hom. illeg. [non Steph. 1908] = Bazzania perrotana E.W.Jones 

1975 (870). 

Mastigobryum pulvinatum Steph. 1891 = Bazzania decrescens subsp. pumila (Mitt.) Pócs 1994 (870, 889). 

Mastigobryum pumilum (Mitt.) Steph. 1908 = Bazzania decrescens subsp. pumila (Mitt.) Pócs 1994 (870, 

889). 

Mastigobryum reflexum Gottsche 1882 = Bazzania reflexa (Gottsche) Steph. 1891 (870). 

Mastigobryum robertii Steph. 1908 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877 (870, 1). 

Mastigobryum schismoideum Steph. 1888 = Bazzania borbonica Steph. 1894 (870) = Bazzania praerupta 


Mastigobryum seychellarum (Gottsche ex Steph.) Steph. 1908 = Bazzania decrescens (Lehm. et Lindenb.) 

Trevis. 1877 (847). 

Mastigobryum staudtianum Steph. 1908 = Bazzania decrescens subsp. molleri (Steph.) E.W.Jones 1975 

(870). 

Mastigobryum stephanii J.B.Jack in Steph. 1886 = Bazzania stephanii (J.B.Jack) Steph. 1891 (1) = Bazzania 

nitida (F.Weber) Grolle 1960 (870). 

Mastigobryum stoloniferum (Sw.) Lindenb. 1845 = Bazzania stolonifera (Sw.) Trevis. 1877 (1) [Excludenda(1)]. 

Mastigolejeunea africana Steph. ex Paris 1911, nom. inval. = Mastigolejeunea florea (Mitt.) Steph. 1912 

(903). 

Mastigolejeunea auriculata (Wilson) Schiffn. ANN(2,33), BEN(22), CAF(2), CMN(2), CON(2), 

GAB(2), GHA(2), GUI(2), IVO(2,53), MAU(4), MDG(1), MOZ(2), NGA(2,127), REU(1),


RIM(10), ROD(1,723), SAO(2), SEN(2), SIE(2), SOC(746), SUD(2), TAN(2), TOG(2,101), 

UGA(96), ZAI(2). 

Mastigolejeunea brevifissa (Gottsche) Steph. 1924. MDG(1150). [Dubia(903)]. 

Mastigolejeunea buettneri Steph. 1888 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea camerunensis Steph. 1924. CMN(887). [Dubia(903)]. 

Mastigolejeunea carinata (Mitt.) Steph. 1912 = Mastigolejeunea auriculata (Wilson) Schiffn. 1893 (835). 

Mastigolejeunea corbieri Steph. 1912 = Mastigolejeunea florea (Mitt.) Steph. 1912 (903). 

Mastigolejeunea crispula Steph. 1912 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea dusenii Steph. 1912 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea florea (Mitt.) Paris BEN(22), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), 

GUI(2), IVO(2), NGA(2), PRI(48), RIM(10), SIE(2), TAN(2), TOG(2,101), UGA(115), ZAI(2). 

Mastigolejeunea fusco-virens Steph. 1912 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea gilletana Steph. 1912 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea humilis (Gottsche) Schiffn. 1893 = Mastigolejeunea auriculata (Wilson) Schiffn. 1893 

(578). 

Mastigolejeunea jungneri Steph. 1912 CMN(2). [Dubia (903)]. Vanden Berghen (903) considered it unlikely to 

be a Mastigolejeunea. Only sterile specimens remain of the only collection. 

Mastigolejeunea lacomblezii Pearson 1922 = Brachiolejeunea nigra (Steph.) Steph. 1912 (906) = Mastigolejeunea 

nigra Steph. 1895 (25). 

Mastigolejeunea microscypha (Hook.f. et Taylor) Steph. 1912 = Phragmicoma microscypha (Hook.f. et 

Taylor 1845) Mitt. 1875 (987, 988) = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 

Mastigolejeunea nigra Steph. BEN(22), CMN(2), GAB(2), GHA(2), GUI(2), IVO(2), MLW(2), 

NGA(2), SAO(2), SIE(2), TAN(2), UGA(96,776), ZAI(2). 

Mastigolejeunea rhodesica (Vanden Berghen) E.W.Jones CAF(2), KEN(2), MDG(1), MLW(2), 

REU(1), TAN(2), ZAI(2), ZAM(2). 

Mastigolejeunea robusta Steph. 1912 = Ptychocoleus ferrugineus (Steph.) Steph. 1912 (903) = Schiffneriolejeunea 

ferruginea (Steph.) Gradst. 1974 (832). 

Mastigolejeunea sudanensis Herzog 1948 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea tarkwana Pearson 1931 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845 (25). 

Mastigolejeunea trigona Steph. 1895 = Mastigolejeunea carinata (Mitt.) Steph. 1912 (903) = Mastigolejeunea 


Mastigolejeunea turgida Steph. BIO(2,33), CMN(2), GHA(2), NGA(2), PRI(2), SAO(2). 

Mastigophora africana S.W.Arnell 1957 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 (847). 

Mastigophora africana var. annobonensis S.W.Arnell 1956, nom. inval. probably = Mastigophora diclados 

(Brid. ex F.Weber) Nees 1838. M. africana was synonymised with M. diclados by Grolle (847), and it is likely the 

records of var. annobonensis cited by Arnell (813) belong to M. diclados. 

Mastigophora diclados (Brid. ex F.Weber) Nees ANN(2,33), ASC(2), BIO(2,33), CMN(2), 

COM(1,4), ETH(2), GAB(2), MAS(1), MAU(1,4), MDG(1,720), MLW(114), REU(1), RIM(10), 

RWA(2), SAO(2), SEY(1,4,13), TAN(2), UGA(50). 

Mastigophora leioclada (Hook.f. et Taylor) Mitt. 1875 = Mastigophora diclados (Brid. ex F.Weber) Nees 

1838 (895). 

Mastigophora madagassa Steph. 1909 = Chandonanthus hirtellus (F.Weber) Mitt. 1887 (985) = Plicanthus 

hirtellus (F.Weber) R.M.Schust. 2003 (780). 

Mastigophora mascarenica (Mitt.) Steph. 1891 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 

(895). 

Megaceros flagellaris (Mitt.) Steph. TAN(2). 

Megaceros lacerus (Nees) Steph. 1916 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975 (1). 

Metacalypogeia fusca (Lehm.) N.Kitag. 1988 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Metalejeunea cucullata (Reinw. et al.) Grolle COM(1), MDG(1), SEY(1,4). 

Metzgeria africana Steph. 1917 = Metzgeria nudifrons Steph. 1892 (767). 

Metzgeria agnewiae Kuwah. 1973 "agnewii" = Metzgeria consanguinea Schiffn. 1893 (767). 

Metzgeria albinea Spruce The collection from the Seychelles reported as M. albinea belongs to M. madagassa Steph. 

(767). 

Metzgeria allanii Steph. 1917 = Metzgeria leptoneura Spruce 1885 (768). 

Metzgeria allionii Steph. 1917. The collections reported by Kuwahara (769) from Malawi probably belong to M.


nudifrons Steph. 1892 (767). 

Metzgeria attenuata Steph. 1916 = Metzgeria consanguinea Schiffn. 1893 (767). 

Metzgeria australis Steph. 1889 = Metzgeria leptoneura Spruce 1885 (729). The entry of M. australis into the 

African checklist seems to have arisen because Kuwahara (624) placed the African taxa M. thomeensis and M. madagassa 

in synonymy with it. 

Metzgeria borneensis Kuwah. 1965 = Metzgeria leptoneura Spruce 1885 (771). M. borneensis was erroneously 

recorded from Madagascar (1000). The collection should presumably be referred to M. leptoneura Spruce. 

Metzgeria brunnthaleri Steph. 1913 = Metzgeria nudifrons Steph. 1892 (167). 

Metzgeria bystroemii S.W.Arnell 1956 = Metzgeria quadrifaria Steph. 1899 (767). 

Metzgeria camerunensis Steph. 1899 = Metzgeria madagassa Steph. 1899 (767). 

Metzgeria capensis S.W.Arnell 1953 = Metzgeria furcata (L.) Dumort. 1835 (486, 550). 

Metzgeria conjugata Lindb. 1875 [Excludenda]. All records of this species from sub-Saharan African [MAU(1), 

MDG(1), MOZ(2), REU(1), SAF(2), ZAI (2)] are old, and are treated here as erroneous. The species is not accepted 

as African in So (767). 

Metzgeria consanguinea Schiffn. BUR(2,767), CMN(767), ETH(2,767), KEN(2,767), LES(75,767), 

MDG(767), MLW(767), REU(767), RWA(2,767), SAF(2,75,767), TAN(2,767), UGA(2,42,767), 

ZAI(2,767), ZAM(767), ZIM(767). 

Metzgeria convexa Vanden Berghen 1951 = Metzgeria consanguinea Schiffn. 1893 (767). 

Metzgeria corralensis Steph. 1899. The presence of this species in Africa is doubtful, and the collection from South 

Africa cited by Kuwahara (769) probably belongs to M. quadrifaria Steph. (767). 

Metzgeria crassipilis (Lindb.) A.Evans BUR(767), MLW(767), TAN(2), UGA(767), ZAI(767). Metzgeria 

propagulifera was reported from TAN in Bizot & Pócs 1974 (136), and although the collection was not mentioned in 

So 2004 (767), it should presumably be assigned to M. crassipilis. 

Metzgeria decipiens (C.Massal.) Schiffn. 1890 = Metzgeria furcata (L.) Dumort. 1835 (550). 

Metzgeria elliotii Steph. 1899 = Metzgeria consanguinea Schiffn. 1893 (767). 

Metzgeria epiphylla A.Evans 1923. The record of M. epiphylla from Uganda (769) belongs to M. consanguinea 

(767). 

Metzgeria furcata (L.) Dumort. BUR(767), CMN(767), KEN(767), LES(767), LBR(767), 

MAU(767), MDG(767), MLW(767), PRI(767), REU(767), SAF(533,767), SAO(2,767), SIE(2,767), 

TAN(767), UGA(767), ZAI(767). 'M. thomeensis' has been reported from ANN(2,33), BIO(36), COM(1), 

GHA(2), IVO(2), MOZ(2), NGA(2), RIM(10), RWA(2) and SEY(4), but collections from these countries were not 

examined by So (767). She pointed out that many collections deposited in various herbaria and annotated as M. thomeensis, 

belong either to M. furcata or M. madagassa. 

Metzgeria furcata subsp. crassipilis Lindb. 1878 = Metzgeria crassipilis (Lindb.) A.Evans 1909 (767). 

Metzgeria furcata var. decipiens C.Massal. 1885 = Metzgeria furcata (L.) Dumort. 1835 (550). 

Metzgeria hamata auct. [non Lindb. 1878] = Metzgeria leptoneura Spruce 1885 (1008). 

Metzgeria hamata Lindb. 1878 [non M. hamata auct.] = Metzgeria procera Mitt. 1855 (1) [Excludenda(1)]. 

Metzgeria hedbergii Vanden Berghen 1953 = Metzgeria quadrifaria Steph. 1899 (767). 

Metzgeria latifrons Steph. 1910 = Metzgeria nudifrons Steph. (767). 

Metzgeria leptoneura Spruce BIO(767), CMN(767), GUI(767), KEN(767), MAU(1,767), 

MDG(1,767), MLW(2,767), NGA(767), PRI(767), REU(1,767), RWA(2), SAO(2,767), SAF(767), 

SEY(4), TAN(2,767), UGA(2,96,767), ZAI(2), ZIM(2). 

Metzgeria limbato-setosa Steph. 1910 = Metzgeria madagassa Steph. 1899 (767). 

Metzgeria luzonensis Kuwah. = Metzgeria leptoneura Spruce 1885 (771). The collection of M. luzonensis from 

Madagascar (770) belongs to M. madagassa (767). 

Metzgeria madagassa Steph. CMN(767), ETH(767), LBR(767), MAU(767), MDG(1,767), 

MLW(767), NGA(767), REU(767), RWA(767), SAF(767), SEY(767), TAN(767), UGA(767), 

ZAI(767), ZIM(767). 

Metzgeria muscicola Steph. 1899 = Metzgeria quadrifaria Steph. 1899 (767). 

Metzgeria nicomariei Veltman et Potgieter 2000 = Metzgeria furcata (L.) Dumort. 1835 (767). 

Metzgeria nudifrons Steph. CMN(16), LES(75,767), MDG(767), MLW(767), SAF(75,767), 

TAN(767), UGA(767), ZAI(2), ZAM(767), ZIM(767). 

Metzgeria papulosa Steph. 1899 = Metzgeria leptoneura Spruce 1885 (424). The record of M. papulosa from 

Madagascar (1001) is treated as an error, and likely to refer to M. leptoneura (767). 

Metzgeria perrotana Steph. 1899 = Metzgeria furcata (L.) Dumort. 1835 (486, 550). 

Metzgeria procera Mitt. 1855 MAD(1), MAU(1), REU(1). [Excludenda(1)]. 

Metzgeria propagulifera Vanden Berghen 1948 = M. crassipilis (Lindb.) A.Evans 1909 (154, 767). 

Metzgeria quadrifaria Steph. BIO(2,767), CMN(767), KEN(2,767), LES(75,767), MDG(767), 

RWA(2), SOM(2,767), TAN(767), UGA(767), ZAI(767), ZAM(767). 

Metzgeria recurva Steph. 1886 = Metzgeria leptoneura Spruce 1885 (167). 

Metzgeria renauldii Steph. 1899 = Metzgeria leptoneura Spruce 1885 (424).


Metzgeria rufula Spruce 1885. The records from Cameroon and Uganda of this neotropical species (155) could refer 

to M. madagassa (767). 

Metzgeria ruwenzorensis Gola 1907 = Metzgeria furcata (L.) Dumort. 1835 (767). 

Metzgeria saxbyi Pearson BIO(2,33), CMN(2,767), GHA(767), UGA(96,767). Arnell (170) reported M. 

saxbyi from South Africa and Swaziland, but confirmation is required. 

Metzgeria tabularis Steph. 1913 = Metzgeria furcata (L.) Dumort. 1835 (767). 

Metzgeria thomeensis Steph. 1891 = Metzgeria furcata (L.) Dumort. 1835 (767). 

Metzgeria tusii Gola 1914 = Metzgeria leptoneura Spruce 1885 (767). 

Metzgeria vandenberghenii Kuwah. 1983 = Metzgeria consanguinea Schiffn. 1893 (767). 

Metzgeria violacea (Ach.) Dumort. 1835. This species has been reported from several African countries, but recent 

examination (So, 2004) has shown that many collections identified as this species are not true M. violacea. Its presence 

in Africa is therefore considered doubtful (767). 

Metzgeria violascens (Ach.) Dumort. ex S.W.Arnell 1965, nom. inval. = Metzgeria violacea (Ach.) Dumort. 

1835 (1). 

Metzgeria vivipara A.Evans 1910. The report of this species from Réunion should probably be referred to M. nudifrons 

(767). 

Metzgeria warnstorfii Steph. 1899 "warnstorffii" = Metzgeria madagassa Steph. 1899 (767). 

Microlejeunea africana Steph. ANN(2,33), BEN(2), BIO(2,33,72), BUR(2), CAF(2), CMN(2), 

COM(1), CON(2), ETH(2), GHA(2), GNB(2), GUI(2), IVO(2,53), KEN(2,665,731,759), LES(75), 

MAU(108), MDG(1), MLW(2,563), MOZ(2), NGA(2), PRI(2), RIM(10), ROD(723), RWA(2), 

SAF(2,75), SAO(2), SEY(1,108), SIE(2), SOC(94), TAN(2,736), TOG(101), UGA(2,42), ZAI(2), 

ZIM(2). 

Microlejeunea ankasica E.W.Jones GHA(2), SIE(2). 

Microlejeunea byssoides (Gottsche) Pearson 1892 MAD(1). [Dubia(1)]. 

Microlejeunea cochlearifolia Steph. 1888 perhaps = Microlejeunea kamerunensis Steph. 1915 (865). 

Microlejeunea cucullata (Reinw. et al.) Steph. 1915 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995 

(1). 

Microlejeunea dispar Jovet-Ast MAU(1), REU(1). 

Microlejeunea fissistipula Steph. MDG(1,720). 

Microlejeunea furva (Mitt.) Steph. 1915 = Lejeunea furva Mitt. 1879 (987) = Lejeunea eckloniana Lindenb. 

1845 (869). 

Microlejeunea globosa (Spruce) Steph. KEN(759). 

Microlejeunea gracillima sensu Sim 1926 [? non (Mitt.) Steph. 1895] presumed = Microlejeunea africana 

Steph. 1888 (fide illustration in Sim (894)). 

Microlejeunea gracillima (Mitt.) Steph. = Microlejeunea ulicina (Taylor) A.Evans 1902 (987, 984). 

Microlejeunea helenae (Pearson) Sim 1926 = Lejeunea helenae Pearson 1886 (894). 

Microlejeunea heterophylla Steph. 1915 = Lejeunea caespitosa Lindenb. 1845 (157). 

Microlejeunea inflata Steph. COM(1), MDG(1,720). 

Microlejeunea jungneri Steph. 1915 [non Lejeunea jungneri (Steph.) Steph. 1901] = Lejeunea confusa 

E.W.Jones 1972 (157). 

Microlejeunea kamerunensis Steph. BUR(2), CMN(2), IVO(2,53), KEN(2), MLW(563), MOZ(2), 

NGA(2), PRI(2), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Microlejeunea magnilobula Gola UGA(2). 

Microlejeunea minutistipula Steph. TAN(2). 

Microlejeunea nyandaruensis Pócs KEN(738,759), RWA(738). 

Microlejeunea oblongistipula (Gottsche) Pearson MDG(1), ROD(723), SEY(4,108). 

Microlejeunea ocellifera S.W.Arnell 1953 = Microlejeunea africana Steph. 1888 (865). 

Microlejeunea ovistipula Steph. ZAI(2). 

Microlejeunea papulosa (Gottsche) Pearson MDG(1). 

Microlejeunea saccula Steph. 1915 = Microlejeunea africana Steph. 1888 (1). 

Microlejeunea ulicina (Taylor) A.Evans CVI(37). 

Microlejeunea usambarensis Steph. TAN(2). 

Microlepidozia stephanii (Renauld ex Steph.) S.W.Arnell 1965, nom. inval. = Kurzia capillaris subsp. 

stephanii (Renauld) Pócs 1984 (382). 

Microlepidozia succulenta (Sim) S.W.Arnell 1962 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Microlepidozia tabularis (Steph.) S.W.Arnell 1962 = Kurzia capillaris (Sw.) Grolle 1964 (382). 

Mnioloma fuscum (Lehm.) R.M.Schust. BIO(36), CMN(2), ETH(2), LES(2), MDG(1,720), 

REU(1,4), RWA(2), SAF(2), SEY(1,4), STH(2), SWZ(2), TAN(2), UGA(2), ZAI(2). 

Monocarpus sphaerocarpus D.J.Carr. SAF(2,76).


Monoclea adglutinata Hook.f. et Taylor 1845 = Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche et 

al. 1846 (917, 982). 

Monoclea gottschei Lindb. 1886 - erroneously recorded from Ethiopia; the specimens are Calycularia crispula 

(928). 

Mylia diversifolia (Gottsche) E.W.Jones ex S.W.Arnell 1955 = Leptoscyphus diversifolius (Gottsche) Grolle 

1963 (471). 

Mylia expansa (Lehm.) S.W.Arnell 1955 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Mylia hedbergii S.W.Arnell 1956 = Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. 1959 (471). 

Mylia infuscata (Mitt.) S.W.Arnell 1956 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963 (471). 

Mylia iversenii (Pearson) S.W.Arnell 1955 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Nardia abyssinica (Nees) Cufod. 1952 = Jungermannia abyssinica Nees 1844 (412). 

Nardia arnelliana Grolle REU(1), UGA(2). 

Nardia aurita (Lehm.) Trevis. 1877 = Anastrophyllum auritum (Lehm.) Steph. 1901 (163). 

Nardia dusenii Steph. 1891 = Jungermannia dusenii (Steph.) Steph. 1901 (412). 

Nardia exserta Steph. ex Besch. 1895, hom. inval. [non A.Evans 1891] = Jungermannia perloi (Gola) Váňa 

1973 (1). 

Nardia jackii Steph. 1892 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1983 (937). 

Nardia stolonifera Steph. 1892 = Jungermannia borgenii Gottsche ex Pearson 1893 (412). 

Nardia verrucosa Steph. 1892 = Jungermannia dusenii (Steph.) Steph. 1901 (412). 

Nesolejeunea intercalaris Herzog 1947 = Lejeunea acuta Mitt. 1864 (968). 

Noteroclada porphyrorhiza (Nees) Mitt. 1855 = Fossombronia porphyrorhiza (Nees) Prosk. 1955 (2) [Dubia(2)]. 

Notoscyphus africanus (Pearson) S.W.Arnell 1955 = Odontoschisma africanum (Pearson) Sim 1926 (888). 

Notoscyphus belangerianus (Lehm. et Lindenb.) Mitt. 1877 = Notoscyphus lutescens (Lehm. et Lindenb.) 

Mitt. 1873 (916, 1). 

Notoscyphus congoanus Steph. 1917 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (937). 

Notoscyphus flexuosus (Lehm.) Sim 1926 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (883, 97). 

Notoscyphus jackii (Steph.) Steph. 1901 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (937). 

Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. BUR(2), CMN(2), COM(1,4), MAU(1), 

MDG(1,4,720), MLW(114), REU(1), RWA(2), SAF(2), ZAI(2). 

Notoscyphus natalensis Sim 1926 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (937). 

Notoscyphus variifolius Mitt. 1877 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (883, 97). 

Notoscyphus vermicularis (Lehm.) Steph. 1901 = Clasmatocolea vermicularis (Lehm.) Grolle 1960 (97). 

Notothylas angolensis Steph. 1917 = Notothylas decurva (Mitt.) Steph. 1901 (104). 

Notothylas decurva (Mitt.) Steph. ANG(2), CMN(2), GHA(2), NGA(2), SIE(2). Following Schuster 1992 

(141), N. decurva was treated in (2) as a synonym of N. orbicularis. However, Jones 1985 (104) described some observed 

differences between the two taxa, and concluded "on present evidence" that N. decurva should be maintained. It is unclear 

whether the two taxa have been compared in sufficient detail, and for that reason Jones (1985) is followed in this checklist, 

pending any further evidence presented to support synonymy with N. orbicularis. 

Notothylas flabellata Steph. ANG(2), CMN(2), TAN(2). 

Notothylas indica Kashyap GHA(2). 

Notothylas javanica (Sande Lac.) Gottsche IVO(2). 

Notothylas minuta (Mitt.) Steph. 1916 = Phaeoceros minutus (Mitt.) S.W.Arnell 1963 (170). 

Notothylas sp. SIE(25). Jones (in litt.) considered the Sierra Leone plant may represent a new taxon, but that its status 

must remain uncertain until more copious material is available. 

Odontolejeunea hanningtonii (Mitt.) Steph. 1895 = Caudalejeunea hanningtonii (Mitt.) Schiffn. 1909 (891). 

Odontolejeunea lunulata (F.Weber) Schiffn. BIO(2,33), BUR(2), CMN(2), COM(1), CON(2), 

ETH(2), GUI(2), IVO(2), KEN(2,731), MAU(1), MDG(1,4,720), MOZ(2), NGA(2), REU(2), 

RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(42,50), ZAI(2). 

Odontolejeunea mauritiana Steph. 1912 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (837). 

Odontolejeunea sieberiana (Gottsche) Schiffn. 1893 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 

(837). 

Odontolejeunea sieberiana var. africana Steph. 1892, nom. inval. = Odontolejeunea lunulata (F.Weber) 

Schiffn. 1893 (836, 837). 

Odontolejeunea thomeensis Steph. 1912 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 (836, 837). 

Odontolejeunea tortuosa (Lehm. et Lindenb.) Steph. 1912 = Odontolejeunea lunulata (F.Weber) Schiffn. 

1893 (836, 837). 

Odontoschisma africanum (Pearson) Sim MAU(1), MLW(114), REU(45), SAF(2), TAN(2). 

Odontoschisma denudatum var. africanum (Pearson) S.W.Arnell 1957, nom. inval. = Odontoschisma africanum 

(Pearson) Sim 1926 (888). 

Odontoschisma ligulatum Steph. 1890 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (937).


Odontoschisma variabile Sim 1926, hom. illeg. [non (Lindenb. et Gottsche) Trevis. 1877] = Clasmatocolea 

vermicularis (Lehm.) Grolle 1960 (816, 97). 

Omphalanthus filiformis (Sw.) Nees 1845 MAD(1). [Excludenda(1)]. Omphalanthus is a neotropical genus, absent 

from Africa (852). 

Omphalanthus geminiflorus (Nees) Gottsche et al. 1845 = Omphalanthus filiformis (Sw.) Nees 1845 (1) 


Omphalanthus renistipulus Steph. 1911, hom. illeg. [non Lindenb. 1845] = Evansiolejeunea renistipula 

(Steph.) Vanden Berghen 1948 (851) = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960 (850). 

Omphalanthus roccatii (Gola) R.M.Schust. 1963 = Acrolejeunea roccatii Gola (988) = Evansiolejeunea roccatii 

(Gola) Vanden Berghen 1960 (850). 

Omphalolejeunea filiformis (Sw.) Lacout. 1908 = Omphalanthus filiformis (Sw.) Nees 1845 (1) [Excludenda(1)]. 

Otigoniolejeunea villaumei Steph. 1914 = Lejeunea villaumei (Steph.) Grolle 1977 (828). 

Otiona aitonia (Lindenb. et Nees) Mitt. 1877 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 

(502). 

Otolejeunea moniliata Grolle MDG(1). 

Otolejeunea rabenorii Tixier MDG(1). 

Oxymitra cristata Garside ex Perold NAM(2), SAF(2). 

Oxymitra incrassata (Brot.) Sérgio et Sim-Sim SOC(764). 

Pallavicinia attenuata Steph. 1893 = Pallavicinia lyellii (Hook.) Carruth. 1865 (146). 

Pallavicinia byssophora (Lehm. et Lindenb.) Trevis. 1877 = Pallavicinia lyellii (Hook.) Carruth. 1865 

(146). 

Pallavicinia camisassai Gola KEN(120). 

Pallavicinia capensis S.W.Arnell 1954 = Symphyogyna brasiliensis Nees et Mont. 1836 (170). 

Pallavicinia lyellii (Hook.) Carruth. BUR(2), LES(75), MLW(2,29), REU(1), RWA(2), SAF(2,75), 

SAO(2), TAN(2), UGA(50,96), ZAI(2), ZIM(2). 

Pallavicinia pilifera Steph. 1891 = Pallavicinia lyellii (Hook.) Carruth. 1865 (158). 

Pallavicinia procumbens (Hook.f. et Taylor) Mitt. 1875 = Symphyogyna brasiliensis Nees et Mont. 1836 

(970). 

Pallavicinia rubescens Steph. 1917 = Symphyogyna brasiliensis Nees et Mont. 1836 (970). 

Pallavicinia serrata (Mitt.) Grolle 1984 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986 (956). 

Pallavicinia spinosa (Lindenb. et Gottsche) Grolle 1979 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 

1986 (955). 

Pallavicinia stephanii J.B.Jack 1892 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986 (956). 

Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. SAF(2,556). 

Paracromastigum tristanianum (R.M.Schust.) J.J.Engel et R.M.Schust. 1974. See note under Hyalolepidozia 

bicuspidata. 

Paraschistochila engleriana (Steph.) R.M.Schust. 1971 = Gottschea engleriana (Steph.) Grolle et Zijlstra 

1984 (954). 

Paraschistochila neesii (Mont.) R.M.Schust 1971 = Gottschea neesii Mont. 1843 (954). 

Pedinolejeunea himalayensis (Pandé et Misra) Chen et Wu 1964 = Cololejeunea latilobula (Herzog) Tixier 

1985 (135). 

Pellia epiphylla (L.) Corda SOC(746). 

Phaeoceros bolusii (Sim) S.W.Arnell SAF(2), ZIM(2). 

Phaeoceros bulbiculosus (Brot.) Prosk. CMN(2), ERI(2). 

Phaeoceros carolinianus (Michx.) Prosk. ANG(2), CMN(2), CVI(2,37), ETH(2), KEN(2), LES(75), 

MDG(1), MLW(2,114), NAM(2), NGA(2), REU(1), ROD(723), RWA(2), SAF(2,75), SIE(2), 

STH(2,49), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Phaeoceros fulvisporus (Steph.) J.Haseg. RWA(2), SAF(2), TAN(2). 

Phaeoceros jollyanus (Steph.) S.W.Arnell BIO(2,33). 

Phaeoceros laevis (L.) Prosk. 1951 MAD(1), ROD(1). [Excludenda(1)]. 

Phaeoceros laevis subsp. carolinianus (Michx.) Prosk. 1954 = Phaeoceros carolinianus (Michx.) Prosk. 

1951 (991, 683). 

Phaeoceros maranguensis (Steph.) Bapna TAN(2). 

Phaeoceros minutus (Mitt.) S.W.Arnell SAF(2), ZIM(2). 

Phaeoceros sp. (Anthoceros aethyopicus Gola) ETH(2). Probably a species of Phaeoceros as it is described as 

having "frons haud cavernosa" (121). 

Phaeoceros sp. (Anthoceros brunnthaleri Steph.) TAN(2). Probably a species of Phaeoceros as Stephani 1916 

(886) described it as having "frons solida ... sporae pallidae".


Phaeoceros sp. (Anthoceros mildbraedii Steph.) CAM(2). Probably a species of Phaeoceros, as Stephani 1923 

(887) described it as having "frons solida ... sporae pallidae". E.W. Jones (in litt. ca. 1990) considered it is probably synonymous 

with Phaeoceros carolinianus (Michx.) Prosk. 

Phaeoceros sp. (Anthoceros parvifrons Steph.) TAN(2). Probably a species of Phaeoceros as Stephani 1912 

(138) described it as having "frons solida ... sporae pallidae". 

Phaeoceros sp. (Anthoceros pseudocostus Steph.) MDG(1). The spores are described as "pallidae" (Stephani 

1916 (886)), hence probably a Phaeoceros species. 

Phaeoceros sp. (Anthoceros rossoi Gola) KEN(2). Probably a species of Phaeoceros as it is described as "frons 

haud cavernosa ... sporae flavescentes" (120). 

Phaeoceros sp. (Anthoceros tenuissimus Steph.) TAN(2). West Africa (Lobango)(2). Probably a species 

of Phaeoceros as the spores are described as "flavidae" (887). 

Phaeoceros sp. (Anthoceros tigrinus Gola) ETH(2). Probably a species of Phaeoceros as it is described as having 

"frons haud cavernosa ... sporae flavae" (121). 

Phragmicoma abnormis Gottsche 1882 MAD(1). [Dubia(1)]. 

Phragmicoma acutiloba (Hook.f. et Taylor) Gottsche et al. 1847 = Marchesinia brachiata (Sw.) Schiffn. 

1893 (831). 

Phragmicoma amplectens Steph. 1886 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (35, 831). 

Phragmicoma aulacophora Mont. 1843 = Acrolejeunea aulacophora (Mont.) Steph. 1895 (35). 

Phragmicoma carinata Mitt. 1879 = Mastigolejeunea auriculata (Wilson) Schiffn. 1893 (835). 

Phragmicoma emergens Mitt. 1879 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Phragmicoma excavata (Mitt.) Steph. 1886 = Marchesinia excavata (Mitt.) Schiffn. 1893 (891). 

Phragmicoma florea Mitt. 1886 = Mastigolejeunea florea (Mitt.) Paris 1906 (903). 

Phragmicoma fulva Gottsche 1882 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (831). 

Phragmicoma inflexa Gottsche 1882 MAD(1). [Dubia(1)]. 

Phragmicoma microscypha (Hook.f. et Taylor) Mitt. 1875 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907 (831). 

Phragmicoma molleri Steph. 1886 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (832). 

Phragmicoma palisoti Taylor ex Mitt. 1886, nom. nud. = Odontolejeunea lunulata (F.Weber) Schiffn. 1893 

(836). 

Phragmicoma pappeana Nees 1845 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (832). 

Phragmicoma reniloba Gottsche 1845 = Caudalejeunea reniloba (Gottsche) Steph. 1912 (1) [Excludenda(1)].. 

Phragmicoma rotalis (Hook.f. et Taylor) Mitt. 1875 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 


Phragmicoma sagraeana Mont. 1842 = Lopholejeunea subfusca (Nees) Schiffn. 1897 (838). 

Phragmicoma sphaerophora (Lehm. et Lindenb.) Schiffn. 1886 = Lopholejeunea sphaerophora (Lehm. et 

Lindenb.) Steph. 1912 (839). 

Phragmilejeunea molleri (Steph.) R.M.Schust. 1961 = Schiffneriolejeunea polycarpa (Steph.) Gradst. 1974 

(832). 

Phragmilejeunea occulta (Steph.) Bischl. 1965, nom. inval. = Schiffneriolejeunea occulta (Steph.) Gradst. 

1974 (832). 

Phragmilejeunea pappeana (Nees) R.M.Schust. 1954 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 

(832). 

Physiotium sphagnoides Richard ex Nees 1838, nom. illeg. = Pleurozia gigantea (F.Weber) Lindb. 1874 

(1012). 

Physocolea adhaesiva (Mitt.) Steph. 1916 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963 (912). 

Physocolea africana Steph. 1916 = Cololejeunea africana (Steph.) R.M.Schust. 1963 (912). 

Physocolea autoica Steph. 1916 = Cololejeunea autoica (Steph.) Grolle 1995 (1). 

Physocolea bidentula Steph. 1916 = Cololejeunea bidentula (Steph.) E.W.Jones 1954 (858). 

Physocolea bolombensis Steph. 1916 = Cololejeunea tonkinensis Steph. 1895 (135, 654). 

Physocolea cristata Steph. 1916 = Cololejeunea cristata (Steph.) R.M.Schust. 1963 (912). 

Physocolea cuneata (Lehm. et Lindenb.) Steph. 1916 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 

1947 (152). 

Physocolea decemplicata Steph. 1916 = Cololejeunea decemplicata (Steph.) Tixier 1979 (152). 

Physocolea elegans (Steph.) Steph. 1916 = Cololejeunea elegans Steph. 1891 (858). 

Physocolea erectifolia (Steph.) Steph. 1916 = Cololejeunea erectifolia Steph. 1895 (988) = Leptocolea subciliata 

(Mitt.) E.W.Jones 1953 (858) = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 (152). 

Physocolea filicaulis (Steph.) Steph. 1916 = Cololejeunea filicaulis Steph. 1892 (858) = Cololejeunea hildebrandii 

(Austin) Steph. 1897 (135). 

Physocolea flegelii Steph. 1916 = Leptocolea crenatiflora Steph. 1915 (858). See also note under the latter species. 

Physocolea grossidens Steph. 1916 = Cololejeunea malanjae Steph. 1902 (842).


Physocolea malanjae Steph. 1916 = Cololejeunea malanjae Steph. 1902 (853). 

Physocolea myriantha Herzog 1952 = Cololejeunea minutissima subsp. myriocarpa (Nees et Mont.) 

R.M.Schust. 1955 (936). 

Physocolea peculiaris Herzog 1931 = Cololejeunea inflectens (Mitt.) Benedix 1953 (878). 

Physocolea punctata (Gottsche) Steph. 1916 = Cololejeunea punctata (Gottsche) Pearson 1892 (1) [Dubia(1)]. 

Physocolea pusilla (Steph.) Steph. 1916 = Cololejeunea pusilla Steph. 1895 (858). 

Physocolea scabrifolia Schiffn. ex Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (135). 

Physocolea subciliata (Mitt.) Steph. 1916 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947 (152). 

Physocolea villaumei Steph. 1916 = Cololejeunea hildebrandii (Austin) Steph. 1897 (135). 

Physocolea vittata Steph. 1916 = Leptocolea crenatiflora Steph. 1915 [= Cololejeunea crenatiflora (Steph.) 

Vanden Berghen 1972] or Cololejeunea usambarica E.W.Jones 1953 [= Cololejeunea platyneura (Spruce) 

A.Evans 1902] (858). 

Physocolea xavieri Lacout. ex Steph. 1916 = Cololejeunea xavieri (Lacout. ex Steph.) E.W.Jones 1954 

(858). 

Plagiochasma abyssinicum Gola 1914 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 var. rupestre 

(254). 

Plagiochasma aitonia Lindenb. et Nees 1836 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 

var. rupestre (254). 

Plagiochasma appendiculatum Lehm. et Lindenb. ERI(2), KEN(2), SAF(2), SOC(2), TAN(2), 

ZIM(2). 

Plagiochasma appendiculatum var. erythraeum Gola 1914 = Plagiochasma appendiculatum Lehm. et Lindenb. 

1832 (254). 

Plagiochasma beccarianum Steph. ERI(2), MLW(29), NAM(2), SAF(2), SOC(2), TAN(2), 

UGA(115), ZAM(2). 

Plagiochasma capense Sim 1926 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 var. rupestre 

(170, 254). 

Plagiochasma dinteri Steph. 1901 = Plagiochasma microcephalum (Steph.) Steph. 1898 (254). 

Plagiochasma dschallanum Steph. 1898 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 var. 

rupestre (254). 

Plagiochasma eximium (Schiffn. in Steph.) Steph. CMN(2), CVI(2,37), DJI(2), ERI(2), ETH(2,113), 

GUI(2), KEN(2,731), REU(1), SAF(2,75), SIE(2), SOC(2), TAN(2), UGA(2,96), ZAI(2), 

ZIM(727). The collection from Malawi (Crundwell 369), has been re-determined by T. Pócs as Plagiochasma beccarianum. 

Plagiochasma fischerianum (Steph.) Steph. 1898 ("P. fischeri") = Plagiochasma appendiculatum Lehm. et 

Lindenb. 1832 (254). 

Plagiochasma limbatum (Hook.f. et Taylor) Nees 1846 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) 

Steph. 1898 (1047, 254). 

Plagiochasma microcephalum var. microcephalum (Steph.) Steph. ANG(2), ETH(2,113), MDG(1), 

NAM(2), SAF(2), TAN(2), UGA(2). 

Plagiochasma muricatum Steph. 1901 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 (254). 

Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. CHA(3), CVI(37), LES(75), MAU(1), REU(1), 

SAF(75), UGA(42). 

Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. var. rupestre ANG(2), ASC(2), CHA(2), 

CVI(2,254), DJI(2), ERI(2), ETH(2,113), KEN(2,731), LES(2), NAM(2), SAF(2), STH(2), SUD(2), 

TAN(2), UGA(2), ZAI(2), ZIM(2). 

Plagiochasma rupestre var. volkii Bischl. LES(2,75), NAM(2), SAF(2), ZIM(2). 

Plagiochasma schimperi Steph. 1898 = Plagiochasma eximium (Schiffn.) Steph. 1898 (254). 

Plagiochasma tenue Steph. 1898 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898 var. rupestre 

(254). 

Plagiochila abyssinica Mitt. 1886 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila aerea Taylor 1846 REU(1). [Excludenda(1)]. 

Plagiochila africana Steph. BIO(2,33), CAF(2), CMN(2), GHA(2), GUI(48), IVO(2,53), NGA(2), 

RIM(10), SIE(2). 

Plagiochila aloysii-sabaudiae Gola 1907 = Adelanthus unciformis (Taylor) Mitt. 1864 (829) = Adelanthus 

lindenbergianus (Lehm.) Mitt. 1864 (465). 

Plagiochila amphigastriata Herzog 1938 = Plagiochila pinniflora Steph. 1891 (722). 

Plagiochila amplifolia Steph. 1901 perhaps = Plagiochila fusifera Taylor 1846 (722). 

Plagiochila andongensis Steph. 1904 = Plagiochila divergens var. capensis (Steph.) E.W.Jones 1962 (722). 

P. divergens was synonymised with P. kiaeri by Grolle (1), but the status of P. divergens var. capensis is yet to be reassessed.


Plagiochila angolensis Steph. ex Paris 1908, nom. inval. ANG(2). [Dubia(2)]. 

Plagiochila angusta Lindenb. MDG(1,720), REU(1). 

Plagiochila angustifolia Steph. 1913 = Plagiochila strictifolia Steph. 1891 (722). 

Plagiochila angustissima Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila angustitexta Steph. MLW(114), TAN(2). 

Plagiochila anisodonta (Hook.f. et Taylor) Gottsche et al. 1847 = Tylimanthus anisodontus (Hook.f. et 

Taylor) Mitt. 1875 (1046). 

Plagiochila anjouana Gottsche ex Dugas 1928 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (1). 

Plagiochila ankefinensis Steph. 1904 = Plagiochila kiaeri Gottsche 1882 (843). 

Plagiochila armata Steph. 1891 = Plagiochila barteri Mitt. 1886 var. barteri (722). 

Plagiochila attenuata Steph. 1911 = Adelanthus unciformis (Taylor) Mitt. 1864 (829) = Adelanthus lindenbergianus 

(Lehm.) Mitt. 1864 (465). 

Plagiochila bamingensis Steph. CAF(2). 

Plagiochila barteri Mitt. BIO(2,33,36), CMN(2), COM(1), ETH(2), GAB(2), KEN(2,71), MAU(1), 

MDG(1,720), MLW(2,114), PRI(2), REU(1), RWA(2), SAO(2), SIE(2), TAN(2), TOG(2), ZAI(2), 

ZIM(2). 

Plagiochila barteri subsp. colorans (Steph.) Pócs ETH(2), KEN(2,71,731), RWA(2), TAN(2), 

UGA(2,96), ZAI(2). 

Plagiochila barteri var. valida (Steph.) Vanden Berghen ETH(2), MLW(2), REU(1). 

Plagiochila batangensis Steph. 1917 = Plagiochila praemorsa Steph. 1886 (722). 

Plagiochila berthieui Steph. 1893 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843). 

Plagiochila bidentula Steph. ANN(2,33), CMN(2). 

Plagiochila blanda Steph. 1911 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila boivinii Steph. MDG(1), REU(1). 

Plagiochila bomanensis Steph. 1891 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila borbonica Mont. et Gottsche 1856 = Plagiochila pectinata Willd. ex Lindenb. 1839 (843). 

Plagiochila borgenii Gottsche 1882, nom. inval. = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 

(1). 

Plagiochila boryana Gottsche ex Steph. MDG(1), REU(1,4), TAN(2), UGA(42). Heinrichs 2002 (961) 

considered the record from Mauritius to be dubious. 

Plagiochila bovonei Gola 1920 = Plagiochila fusifera Taylor 1846 (158). 

Plagiochila breviramea Steph. 1911 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila brevisedens Steph. 1918 = Plagiochila divergens Steph. 1891 (722) = Plagiochila kiaeri Gottsche 

1882 (1). 

Plagiochila brunneola Steph. ETH(2), SAO(2). 

Plagiochila brunnthaleri Steph. 1913 = Plagiochila effusa Steph. 1895 (722). 

Plagiochila bueensis Steph. 1904 = Plagiochila moenkemeyeri Steph. 1904 (722). 

Plagiochila buettneriana Steph. 1895 = Plagiochila strictifolia Steph. 1891 (722). 

Plagiochila bursata (Desv.) Lindenb. 1840 nom. illeg. = Plagiochila aerea Taylor 1846 (959). 

Plagiochila cacuminis Steph. 1918 = Plagiochila praemorsa Steph. 1887 (722). 

Plagiochila calva (Nees) Lindenb. 1839 = Plagiochila javanica (Sw.) Dumort. 1836 (1) [Excludenda(1, 2)]. 

Plagiochila cambouena Steph. 1904, nom. inval. = Plagiochila cambuena Steph. 1890 (1). 

Plagiochila cambuena Steph. "cambouena" IVO(53), MDG(1), MLW(114), ROD(723), TAN(2). 

Plagiochila camerunensis Steph. 1902 = Plagiochila africana Steph. 1902 (722). 

Plagiochila candelabra Steph. 1904 = Plagiochila pinniflora Steph. 1891 (722). 

Plagiochila capensis Steph. 1904 = Plagiochila divergens var. capensis (Steph.) E.W.Jones 1962 (722). P. 

divergens was synonymised with P. kaieri by Grolle (1), but the status of P. divergens var. capensis is yet to be reassessed. 

Plagiochila carduifolia Steph. 1904 = Plagiochila africana Steph. 1902 (722). 

Plagiochila chenagonii Steph. 1892 = Plagiochila angusta Lindenb. 1843 (1). 

Plagiochila chiovendae Gola ETH(2). 

Plagiochila circumvoluta Gerola ETH(2). 

Plagiochila clavaeflora Steph. 1891 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila collicalyx Steph. 1904 = Plagiochila flabellata Steph. 1886 (722). 

Plagiochila colorans Steph. 1911 = Plagiochila barteri subsp. colorans (Steph.) Pócs 1993 (969). 

Plagiochila comorensis Steph. 1888 = Plagiochila pectinata Willd. ex Lindenb. 1839 (1). 

Plagiochila comorensis fo. johannensis Herzog 1947, nom. inval. = Plagiochila terebrans Nees et Mont. ex 

Lindenb. 1840 (1). 

Plagiochila congoana Steph. 1918 = Plagiochila neckeroidea Mitt. 1860 (722). 

Plagiochila corniculata auct. [non Dumort. 1835] = Plagiochila exigua (Taylor) Taylor 1846 (684). 

Plagiochila corymbulosa Pearson 1886 = Plagiochila heterostipa Steph. 1892 (722).


Plagiochila crispi-crista Steph. 1904 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila crispulo-caudata Gottsche 1882 = Plagiochila squamulosa var. crispulo-caudata (Gottsche) 

Vanden Berghen 1981 (843). 

Plagiochila cristato-dentata Steph. 1904 = Adelanthus unciformis (Taylor) Mitt. 1864 (829) = Adelanthus 

lindenbergianus (Lehm.) Mitt. 1864 (465). 

Plagiochila crollii Steph. 1904 = Plagiochila incerta Gottsche 1857 (1). 

Plagiochila cumminsiana Steph. 1918 = Plagiochila praemorsa Steph. 1886 (722). 

Plagiochila curvatifolia Steph. 1886 = Plagiochila neckeroidea Mitt. 1860 (722). 

Plagiochila decurrens Steph. 1901 = Plagiochila moenkemeyeri Steph. 1904 (722). 

Plagiochila dichotoma (P.Beauv.) Nees et Mont. NGA(2). 

Plagiochila dichotoma var. elongata Lindenb. ex Gottsche et al. 1844 = Plagiochila paucidentata Mont. et 

Gottsche 1856 (1010, 1). 

Plagiochila dichotoma var. laxa Gottsche et al. 1844 = Plagiochila integerrima Steph. 1886 (1). 

Plagiochila dicrana Mitt. 1886 = Plagiochila drepanophylla Sande Lac. 1857 (883, 1). 

Plagiochila divergens Steph. 1891 = Plagiochila kiaeri Gottsche 1882 (1). 

Plagiochila divergens var. capensis (Steph.) E.W.Jones ANG(2), SAF(2,75), ZIM(2). Plagiochila divergens 

was synonymised with P. kiaeri by Grolle (1), but the status of P. divergens var. capensis is yet to be reassessed. 

Plagiochila. divergens var. myriocarpa (Pearson) E.W.Jones ETH(2), KEN(2,21), MLW(2,114), 

RWA(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2). Plagiochila divergens was synonymised with P. kaieri by 

Grolle (1), but the status of P. divergens var. myriocarpa is yet to be reassessed. 

Plagiochila divergens var. sparsa (Steph.) Vanden Berghen 1981 - treated by Grolle (1) under Plagiochila kiaeri Gottsche 

1882. 

Plagiochila drepanophylla Sande Lac. MDG(1), REU(1), TAN(2). 

Plagiochila dschaggana Steph. 1891 probably = Plagiochila sinuosa Mitt. 1886 (722) = Plagiochila squamulosa 

var. sinuosa (Mitt.) Vanden Berghen 1981 (843). 

Plagiochila effusa Steph. ANG(2), BIO(33,72), KEN(2,731), MLW(114), RIM(10), TAN(2), 

UGA(2). 

Plagiochila elgonensis Pearson 1924 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila elongata Gottsche ex Geissler et Bischl. 1989, nom. inval. = Plagiochila paucidentata Mont. et 

Gottsche 1856 (1010, 1). 

Plagiochila emarginata Mont. et Gottsche 1856 = Adelanthus decipiens (Hook.) Mitt. 1864 (1). 

Plagiochila engleriana Steph. 1895 = Plagiochila lastii Mitt. 1886 (722). 

Plagiochila ericicola Steph. KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Plagiochila erythraeae Herzog ERI(2). 

Plagiochila esterhuysenii S.W.Arnell 1954 probably = Plagiochila squamulosa var. crispulo-caudata (Gottsche) 


Plagiochila evansii Steph. 1904 = Plagiochila kiaeri Gottsche 1882 (1). 

Plagiochila exigua (Taylor) Taylor ETH(2), KEN(2,759), LES(75), MLW(114), REU(1), SAF(2,75), 

TAN(2), UGA(42), ZAI(2,667). 

Plagiochila expallescens Steph. 1904 = Plagiochila divergens Steph. 1891 (722) = Plagiochila kiaeri Gottsche 

1882 (1). 

Plagiochila filicicola Steph. UGA(2). 

Plagiochila filiformis Gottsche ex Dugas 1928 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (1). 

Plagiochila fissicalyx Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843). 

Plagiochila flabellata Steph. CMN(2), REU(1), SAO(2), SEY(1), TAN(2). 

Plagiochila flagellaris Steph. ex Renauld 1909, nom. inval. = Plagiochila integerrima Steph. 1886 (1009, 

722). 

Plagiochila fracta Pócs MDG(720). 

Plagiochila fragilifolia Pearson 1922 = Plagiochila praemorsa Steph. 1886 (722). 

Plagiochila frapieri Dugas 1928 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (1). 

Plagiochila frappieri Gottsche ex Besch. 1895, nom. inval. = Plagiochila terebrans Nees et Mont. ex Lindenb. 

1840 (1). 

Plagiochila frondescens (Nees) Lindenb. 1840 ETH(2). [Excludenda]. Records by Gola 1914 (121) of this East 

Asian taxon must represent other species. 

Plagiochila furcata Steph. 1892 = Plagiochila angusta Lindenb. 1843 (843). 

Plagiochila fuscovirens Steph. 1918 probably = Plagiochila squamulosa var. crispulo-caudata (Gottsche) 


Plagiochila fusifera Taylor CAF(2), CMN(2), ETH(2), GAB(2), GHA(2), GUI(2), IVO(2,53), 

KEN(2,731), MLW(2,114), NGA(2), RWA(2), SEG(722), SIE(2), SOC(1069), SOM(2), TAN(2), 

TOG(2,101), UGA(2), ZAI(2), ZAM(2). 

Plagiochila aff. fusifera Taylor SEY(4).


Plagiochila gaffatensis Gottsche ETH(2). 

Plagiochila gasangorii Gola 1914 = Plagiochila lastii Mitt. 1886 (722). 

Plagiochila gentiliana Steph. 1902 = Plagiochila neckeroidea Mitt. 1860 (722). 

Plagiochila gibbiflora Steph. SAO(2). 

Plagiochila gottscheana Lindenb. ex Lehm. 1844 = Leptoscyphus expansus (Lehm.) Grolle 1963 (812, 471). 

Plagiochila granditexta Steph. MDG(1). 

Plagiochila grossealata Steph. 1918 perhaps = Plagiochila praemorsa Steph. 1886 or Plagiochila moenkemeyeri 

Steph. 1904 (722). 

Plagiochila haumannii Herzog KEN(21), MOZ(2), SAF(2), UGA(2), ZAI(2). 

Plagiochila hedbergii S.W.Arnell 1956 = Plagiochila ericicola Steph. 1904 (722). 

Plagiochila henriquesii Steph. 1904 perhaps = Plagiochila divergens Steph. 1891 [Plagiochila kiaeri Gottsche 

1882] or Plagiochila heterostipa Steph. 1892 (722). 

Plagiochila heterostipa Steph. ANG(2), BUR(2), GAB(2), IVO(53), KEN(2), MLW(2,114), 

RWA(2), SAF(2), SWZ(2), UGA(50,96), ZAI(2), ZIM(2). 

Plagiochila heudelotiana Steph. 1904 = Plagiochila fusifera Taylor 1846 (722). 

Plagiochila heudelotiana Steph. ex Paris 1902, nom. inval. = Plagiochila fusifera Taylor 1846 (722). 

Plagiochila hildebrandtii Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843). 

Plagiochila hochstetteri Nees ex Lindenb. 1843 perhaps = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila imerinensis Steph. 1904 = Plagiochila kiaeri Gottsche 1882 (843). 

Plagiochila incerta Gottsche MAU(1), MDG(1,720), REU(1). 

Plagiochila injasutiensis S.W.Arnell SAF(2,75). 

Plagiochila insularia Mitt. STH(2). 

Plagiochila integerrima Steph. BIO(2,33,36,72), BUR(2), CAF(2), CMN(2), CON(2), GAB(2), 

GHA(2), IVO(2,53), KEN(2,731), MAU(1), MDG(1), MLW(114), NGA(2), REU(1), RIM(10), 

RWA(2), SAO(2), SEY(1), SIE(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Plagiochila javanica (Sw.) Nees et Mont. 1836 MAD(1), MAU(1), REU(1). [Excludenda(1, 2)]. 

Plagiochila johannensis Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843). 

Plagiochila jollyana Steph. 1902 = Plagiochila neckeroidea Mitt. 1860 (722). 

Plagiochila jungneri Steph. 1904 = Plagiochila barteri Mitt. 1886 subsp. barteri (722). 

Plagiochila kamerunensis Steph. 1918 = Plagiochila strictifolia Steph. 1891 (722). 

Plagiochila katangae Gola 1920 = Plagiochila crispulo-caudata Gottsche 1882 (158) = Plagiochila squamulosa 

var. crispulo-caudata (Gottsche) Vanden Berghen 1981 (843). 

Plagiochila kiaeri Gottsche ANG(2), BUR(2), KEN(2), MDG(1,720), MLW(2,114), RWA(2), 

SEY(4), TAN(2), UGA(2), ZAI(2). 

Plagiochila knysnana S.W.Arnell 1954 = Leptoscyphus expansus (Lehm.) Grolle 1963 (471). 

Plagiochila laevifolia Gola KEN(2). 

Plagiochila lastii Mitt. ANG(2), ETH(2), KEN(2,71,731), MLW(2,114), RWA(2), SAF(2,75), 

TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Plagiochila laxifolia Gottsche 1882 perhaps = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila lecontei Steph. Congo francais(2). 

Plagiochila ledieui Steph. Congo(2). 

Plagiochila lenormandii Gottsche 1882, nom. inval. = Plagiochila repanda (Schwägr.) Lindenb. 1840 

(1011, 1). 

Plagiochila ligulata Steph. 1904 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (1). 

Plagiochila liliputana Herzog 1947 = Plagiochila rodriguezii Steph. 1890 (1). 

Plagiochila lindblomii Pearson 1924 probably = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 

(722). 

Plagiochila lindenbergiana (Lehm.) Gottsche et al. 1844 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864 

(465). 

Plagiochila loloensis Steph. BIO(33), CMN(2), NGA(2), SAO(2). 

Plagiochila longicauda Steph. 1921 = Plagiochila pinniflora Steph. 1891 (722). 

Plagiochila lophophora R.M.Schust. 1959 = Plagiochila exigua (Taylor) Taylor 1846 (1088, 684). 

Plagiochila lunata S.W.Arnell LES(75), SAF(2). 

Plagiochila lurida Steph. 1911 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila macrostachya Lindenb. 1840 REU(1). [Excludenda(1)]. 

Plagiochila madagascariensis Gottsche 1882, nom. inval. [non Steph. 1904] = Plagiochila repanda 

(Schwägr.) Lindenb. 1840 (1011, 1). 

Plagiochila madagascariensis Steph. 1904 [non Gottsche 1882] = Plagiochila cambuena Steph. 1890 (843). 

Plagiochila maranguana Steph. 1895 = Plagiochila divergens Steph. 1891 (722) = Plagiochila kiaeri Gottsche 

1882 (1).


Plagiochila mascarena Gottsche 1857 = Plagiochila paucidentata Mont. et Gottsche 1856 (1). 

Plagiochila mauritiana (Nees) Lindenb. 1840 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (1011, 843). 

Plagiochila mediocris Dugas 1928 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (1). 

Plagiochila moenkemeyeri Steph. BIO(2,33), CMN(2), ETH(2), GHA(2), NGA(2), RIM(10), 

SAO(2), SIE(2), TAN(2), ZAI(2). 

Plagiochila molleri Steph. 1886 probably = Plagiochila flabellata Steph. 1886 (722). 

Plagiochila moschiensis Steph. 1904 perhaps = Plagiochila strictifolia Steph. 1891 or Plagiochila moenkemeyeri 

Steph. 1904 (722). 

Plagiochila multiflora Steph. TAN(2). 

Plagiochila multispica Steph. 1905 = Plagiochila moenkemeyeri Steph. 1904 (722). 

Plagiochila myriocarpa Pearson 1924 = Plagiochila divergens var. myriocarpa (Pearson) E.W.Jones (722). 

P. divergens was synonymised with P. kaieri by Grolle (1), but the status of P. divergens var. myriocarpa is yet to be 

reassessed. 

Plagiochila natalensis Pearson 1886 perhaps = Plagiochila heterostipa Steph. 1892 (722). 

Plagiochila naveauiana Pearson 1922 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila neckeroidea Mitt. BIO(36), CMN(2), CON(2), GAB(2), GHA(2), IVO(2,53), LBR(2), 

NGA(2), RIM(10), SIE(2), UGA(42), ZAI(2). 

Plagiochila negrii Gola ETH(2). 

Plagiochila nemophila Gottsche 1882 = Plagiochila incerta Gottsche 1857 (1). 

Plagiochila noditexta Steph. 1904 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (1). 

Plagiochila nudicaulis Steph. 1895 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila nudiuscula Gottsche 1882 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (1). 

Plagiochila oblongotrigona Steph. 1921 = Plagiochila effusa Steph. 1895 (722). 

Plagiochila ovatotrigona Steph. 1904 = Plagiochila moenkemeyeri Steph. 1904 (722). 

Plagiochila pallida Gottsche 1882 = Plagiochila angusta Lindenb. 1843 (1). 

Plagiochila pallidevirens Steph. 1921 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila paucidentata Mont. et Gottsche MAS(1), MAU(1), MDG(1,720), REU(1). 

Plagiochila pearsoni Steph. 1921 = Plagiochila praemorsa Steph. 1886 or Plagiochila heterostipa Steph. 

1892 or Plagiochila kiaeri Gottsche 1882 (722). 

Plagiochila pectinata Willd. ex Lindenb. BIO(72), CMN(2), COM(1,4), GHA(2), KEN(731), 

MAS(1), MAU(1), MDG(1,720), MLW(114), REU(1), RIM(10), SAO(2), SEY(1,4), SIE(2), 

UGA(50), ZAI(2). 

Plagiochila perflorens Gola 1920 = Plagiochila crispulo-caudata Gottsche 1882 (158) = Plagiochila squamulosa 

var. crispulo-caudata (Gottsche) Vanden Berghen 1981 (843). 

Plagiochila perloi Gola 1914 = Plagiochila sinuosa Mitt. 1886 (722) = Plagiochila squamulosa var. sinuosa 

(Mitt.) Vanden Berghen 1981 (843). 

Plagiochila perrotana Steph. 1904 = Plagiochila repanda var. perrotana (Steph.) Vanden Berghen 1981 

(843). 

Plagiochila pinniflora Steph. CMN(2), CON(2), GAB(2), GHA(2), IVO(2), NGA(2), RIM(10), 

SIE(2), TOG(2), UGA(2,42), ZAI(2). 

Plagiochila pomai Pearson 1922 = Plagiochila neckeroidea Mitt. 1860 (722). 

Plagiochila praemorsa Steph. ANN(2,33), BIO(2,33,36), CAF(2), CMN(2), CON(2), GAB(2), 

GHA(2), IVO(2,53), NGA(2), RIM(10), SAO(2), SIE(2), UGA(2), ZAI(2), ZAM(2). 

Plagiochila pseudoattenuata S.W.Arnell 1962. SAF(2,75). Arnell first published a description of this taxon in 

1962, but in Hep. South Africa 1963 (170) treated the taxon as a synonym of Plagiochila subalpina. However, in a discussion 

of P. subalpina, Vanden Berghen 1981 (843) stated that the description of P. subalpina by S. Arnell (Hep. South Africa: 

246, fig 178, 1963) appears to apply to another taxon. Accordingly, it seems safer to list only South African records 

here, and all other African records under P. subalpina pending further study of these and related species. 

Plagiochila punctata Taylor BIO(2,33). 

Plagiochila quintasii Steph. 1904 = Plagiochila barteri Mitt. 1886 subsp. barteri (722). 

Plagiochila radicans Steph. 1902 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila ragazzii Gola ERI(121), ETH(2). 

Plagiochila ramentacea Steph. 1921 = Plagiochila pinniflora Steph. 1891 (722). 

Plagiochila refracta Brid. ex Gottsche 1857, nom. inval. = Plagiochila paucidentata Mont. et Gottsche 1856 

(1010, 1). 

Plagiochila renauldii Steph. MAU(1), REU(1). 

Plagiochila repanda (Schwägr.) Lindenb. KEN(2), MAU(1), MDG(1), MLW(2), REU(1). 

Plagiochila repanda var. perrotana (Steph.) Vanden Berghen COM(1), MDG(1). 

Plagiochila replicatula Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843). 

Plagiochila richardiana Steph. 1902 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (843).


Plagiochila rivana Steph. 1921 perhaps = Plagiochila fusifera Taylor 1846 or Plagiochila moenkemeyeri 

Steph. 1904 (722). 

Plagiochila rivularis Steph. 1921 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila rodriguezii Steph. COM(1), MDG(1), REU(1). 

Plagiochila rotundifolia Steph. 1904, hom. illeg. [non Colenso 1888] = Plagiochila loloensis Steph. 1904 

(1). 

Plagiochila cf. rodriguezii Steph. ROD(723). 

Plagiochila rubricaulis Steph. ZAI(2,159). 

Plagiochila rudolfii Pócs TAN(2). 

Plagiochila runssorensis Steph. 1895 = Plagiochila divergens Steph. 1891 (722) = Plagiochila kiaeri Gottsche 

1882 (1). 

Plagiochila rupicola Steph. 1895 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila ruspoliana Steph. 1904 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila rutenbergiana Gottsche 1882 = Plagiochila repanda (Schwägr.) Lindenb. 1840 (883, 1). 

Plagiochila rutenbergii Gottsche 1882, nom. inval. = Plagiochila repanda (Schwägr.) Lindenb. 1840 (883, 

1). 

Plagiochila ruwenzorensis Steph. 1911 = Syzygiella geminifolia (Mitt.) Steph. 1902 (416). 

Plagiochila ruwenzoris Steph. 1921 = Plagiochila colorans Steph. ex Mildbr. 1911 (722) = Plagiochila 

barteri subsp. colorans (Steph.) Pócs 1993 (969). 

Plagiochila salvadorica Steph. ANG(2), CAF(2), GHA(2), GUI(48), IVO(53), SIE(2), ZAI(2). 

Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb. SAF(2). 

Plagiochila schimperiana Steph. 1895 = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila securifolia Nees in Lindenb. 1840 = Syzygiella securifolia (Nees in Lindenb.) Inoue 1979 

(1089). [Dubia(2)]. 

Plagiochila sikorae Steph 1893 = Plagiochila barteri Mitt. 1886 subsp. barteri (843). 

Plagiochila simplex (Sw.) Lindenb. 1840 REU(1). [Excludenda(1)]. 

Plagiochila sinuosa Mitt. 1886 = Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981 (843). 

Plagiochila socorana Gerola 1947 probably = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila sparsa Steph. 1903 = Plagiochila kiaeri Gottsche 1882 (843). 

Plagiochila sprengeri Steph. 1904 = Plagiochila lastii Mitt. 1886 (722). 

Plagiochila squamulosa Mitt. BIO(2,33), BUR(2), CMN(2), DJI(2), ERI(2), ETH(2), 

KEN(2,21,71,665,759), MLW(114), RWA(2), SOM(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2). 

Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen CMN(2), ETH(2), 

KEN(2,21,71,665), LES(75), MDG(1), MLW(2,114), MOZ(2), REU(1), RWA(2), SAF(2,75), 

TAN(2), UGA(50), ZAI(2), ZAM(2), ZIM(2). 

Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen ANG(2), BUR(2), ETH(2), 

KEN(2,21,71,665,731), MAU(1), MDG(1), MLW(2,114), REU(1), RWA(2), TAN(2), UGA(2,96), 

ZAI(2), ZAM(2), ZIM(2). 

Plagiochila squarrosifolia Steph. 1921 = Plagiochila integerrima Steph. 1886 (722). 

Plagiochila stahlii Steph. 1921 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila strictifolia Steph. ANG(2), ANN(2,33), BIO(2), CAF(2), CMN(2), GHA(2), IVO(2,53), 

KEN(731), MLW(2,114), NGA(2), PRI(2), SAO(2), SIE(2), TAN(2), TOG(2,101), UGA(2,96), 

ZAI(2), ZAM(2). 

Plagiochila stuhlmannii Steph. 1895 = Plagiochila divergens Steph. 1891 (722) = Plagiochila kiaeri Gottsche 

1882 (1). 

Plagiochila subalpina Steph. KEN(2,731), LES(75), MDG(1,720), ?MLW(2), RWA(2), TAN(2), 

UGA(2), ZAI(2). See note under P. pseudoattenuata. 

Plagiochila subcrispula Steph. 1921 = Plagiochila africana Steph. 1902 (722). 

Plagiochila subintegerrima (Reinw. et al.) Mont. et Nees 1839 = Syzygiella subintegerrima (Reinw. et al.) 

Spruce 1895 (1) [Excludenda(1)]. 

Plagiochila sublinearis Gottsche 1882 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (1). 

Plagiochila subrepanda Steph. 1921 = Plagiochila pinniflora Steph. 1891 (722). 

Plagiochila subtenuis Steph. 1902 REU(1). [Excludenda(1)]. 

Plagiochila superba (Nees ex Spreng.) Mont. et Nees 1839 MAU(1). [Excludenda(1)]. 

Plagiochila telekii Steph. 1891 probably = Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden 

Berghen 1981 or Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981 (722). 

Plagiochila tenax Steph. 1890 = Plagiochila pectinata Willd. ex Lindenb. 1839 (883, 722). 

Plagiochila terebrans Nees et Mont. ex Lindenb. ANG(2), ANN(2,33), BIO(36), CMN(2), 

COM(1,4), IVO(53), KEN(2), MAS(1), MDG(1,720), MLW(2,114), REU(1), RIM(10), RWA(2), 

SAF(2), SAO(2), TAN(2), UGA(2,96), ZAI(2).


Plagiochila thomeensis Steph. 1886 = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 (722). 

Plagiochila togoensis Steph. 1895 = Plagiochila strictifolia Steph. 1891 (722). 

Plagiochila triangularis Steph. 1886 = Plagiochila barteri Mitt. 1886 var. barteri (722). 

Plagiochila tricuspis Steph. ANN(2), BIO(33), SAO(2). 

Plagiochila tridenticulata auct. [non Dumort. 1835] = Plagiochila exigua (Taylor) Taylor 1846 (684). 

Plagiochila ubangiensis Steph. 1921 perhaps = Plagiochila terebrans Nees et Mont. ex Lindenb. 1840 

(722). 

Plagiochila uhligii Steph. 1921 = Plagiochila colorans Steph. ex Mildbr. 1911 (722) = Plagiochila barteri 

subsp. colorans (Steph.) Pócs 1993 (969). 

Plagiochila undulata Dugas 1928, hom. illeg. [non (L.) Mont. et Nees 1838] = Plagiochila cambuena Steph. 

1890 (1). 

Plagiochila uondana Gola 1914 probably = Plagiochila squamulosa Mitt. 1863 (722). 

Plagiochila valida Steph. 1904 = Plagiochila barteri var. valida (Steph.) Vanden Berghen 1981 (843). 

Plagiochila vignolii Gola 1914 = Plagiochila sinuosa Mitt. 1886 (722) = Plagiochila squamulosa var. sinuosa 

(Mitt.) Vanden Berghen 1981 (843). 

Plagiochila volkensii Steph. 1895 = Plagiochila lastii Mitt. 1886 (722). 

Plagiochila wilmsiana Steph. SAF(2). 

Plagiochila winteri Steph. 1904 = Plagiochila strictifolia Steph. 1891 or Plagiochila moenkemeyeri Steph. 

1904 (722). 

Plagiochila sp. (sect. Choachinae) CMN(722). Placed under P. wilmsiana in (2) and in edition 1, but the identities 

of the specimens are uncertain (722). 

Plagiochila sp. (sect. Zonatae) CMN(722). Overlooked in (2). According to Jones (1962), resembling P. yakushimensis 

(sic) [= P. semidecurrens (Lehm. et Lindenb.) Lindenb.], but could not be named (722). 

Pleuroschisma convexa Steph. 1895 = Jungermannia convexa Thunb. 1800, hom. illeg. [non Scop. 1772] 

(988) = Bazzania nitida (F.Weber) Grolle 1960 (888). 

Pleurozia gigantea (F.Weber) Lindb. MAU(1), MDG(720), REU(1,4), SAO(2), SEY(4), STH(2,49), 

TAN(2). 

Pleurozia gigantea var. major (J.B.Jack) A.Evans 1892 = Pleurozia gigantea (F.Weber) Lindb. 1874 (1012). 

Pleurozia sphagnoides Trevis. 1877 = Pleurozia gigantea (F.Weber) Lindb. 1874 (1012). 

Plicanthus birmensis (Steph.) R.M.Schust. MDG(1), MLW(114). 

Plicanthus giganteus (Steph.) R.M.Schust. ETH(2), KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Plicanthus hirtellus (F.Weber) R.M.Schust. BIO(36), CMN(2), COM(1), ETH(2), KEN(2,71), 

MAU(1), MDG(1,720), MLW(2,114), REU(1,4), RWA(2), SAF(2), SAO(2), SEY(1), SIE(2), 

TAN(2), UGA(2,42,50,96), ZAI(2), ZIM(2). 

Plicanthus sp. (Chandonanthus perloi Gola) KEN(2). 

Porella abyssinica (Nees) Trevis. ETH(2), KEN(2,71), RWA(2), TAN(2), ZAI(2). 

Porella abyssinica var. hoehnelii (Steph.) Pócs BUR(2), ETH(2), KEN(2,21,665), MDG(1), 

MLW(109), MOZ(2), SAF(2), TAN(2), UGA(2,96). 

Porella arboris-vitae (With.) Grolle CVI(2,37). 

Porella capensis (Gottsche) Mitt. ETH(2), KEN(2), MLW(109), SAF(2,75), TAN(2), UGA(96), 

ZIM(2). 

Porella cucullistipula Steph. COM(1), MAU(1,43), MDG(1). 

Porella hoehneliana Steph. ex S.W.Arnell 1957, nom. inval. = Porella hoehnelii Steph. 1891 (861) = 

Porella abyssinica var. hoehnelii (Steph.) Pócs 1994 (1002). 

Porella hoehnelii Steph. 1891 = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994 (1002). 

Porella keniana (Steph.) S.W.Arnell 1956, nom. inval. = Porella hoehnelii Steph. 1891 (861) = Porella 

abyssinica var. hoehnelii (Steph.) Pócs 1994 (1002). 

Porella laevigata (Schrad.) Pfeiffer 1855 = Porella arboris-vitae (With.) Grolle 1969 (921). 

Porella madagascariensis (Nees et Mont.) Trevis. MDG(1). 

Porella matongae (Herzog) S.W.Arnell 1963 = Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 1955 

(472). 

Porella mattirolii (Gola) S.W.Arnell ETH(2). 

Porella montantii (Steph.) E.W.Jones MDG(1). 

Porella nilgheriensis (Mont.) Trevis. 1877 MAD(1). [Excludenda(1)]. 

Porella prolixa (Gottsche ex Steph.) E.W.Jones REU(1). 

Porella subdentata var. camerunensis E.W.Jones BIO(2,33), CMN(2), GHA(2). 

Porella subdentata (Mitt.) E.W.Jones var. subdentata ANG(2), CAF(2), CMN(2), ETH(2), GAB(2), 

GHA(2), GUI(2), IVO(51), KEN(2,665), MLW(109), NGA(2), PRI(2), RWA(2), SAO(2), SIE(2), 

TAN(2), TOG(2), UGA(2,96), ZAI(2), ZAM(2). 

Porella triquetra (Steph.) E.W.Jones ETH(2), MDG(1), MLW(2), REU(1), TAN(2), ZIM(2).


Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell MOZ(2), SAF(2,75). 

Potamolejeunea holtii (Spruce) Greig-Smith 1948 = Lejeunea holtii Spruce 1887 (458). 

Prionolejeunea aberrans Steph. 1923 = Lejeunea papilionacea Steph. 1892 (25). 

Prionolejeunea corbisieri Pearson 1922 = Lejeunea sp (836). 

Prionolejeunea deplanata Schiffn. 1893 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea exarmata Steph. 1913 = Lejeunea papilionacea Steph. 1892 (123). 

Prionolejeunea fissistipula Steph. 1913 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913 (2). 

Prionolejeunea grata (Gottsche) Schiffn. ANN(2,33), BIO(72), CMN(2), COM(1,4), GAB(2), 

GHA(2), IVO(2), MAU(1), MDG(1,720), MLW(563), NGA(2), PRI(2), REU(1), RIM(10), 

RWA(2), SAO(2), SEY(1), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Prionolejeunea hamidens Herzog 1947 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea kindbergii Steph. 1892, nom. inval. = Prionolejeunea serrula (Mitt.) Steph. 1913 (27) = 

Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea kindbergii (Steph.) Schiffn. 1893 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea madagascariensis Steph. 1923 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea principensis Vanden Berghen PRI(2). 

Prionolejeunea pyriflora Steph. 1923 = Cheilolejeunea cordigera (Steph.) Grolle 1977 (828). 

Prionolejeunea rutenbergiana (Gottsche) Steph. 1913 = Prionolejeunea grata (Gottsche) Schiffn. 1893 

(847). 

Prionolejeunea rutenbergii Gottsche ex Pearson 1892, nom. inval. = Prionolejeunea grata (Gottsche) 

Schiffn. 1893 (847). 

Prionolejeunea serrula (Mitt.) Steph. 1913 = Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Prionolejeunea uncatifolia Pearson 1922 probably = Prionolejeunea serrula (Mitt.) Steph. 1913 (27) = 

Prionolejeunea grata (Gottsche) Schiffn. 1893 (847). 

Pseudomarsupidium decipiens (Hook.) Grolle 1963 = Adelanthus decipiens (Hook.) Mitt. 1864 (465). 

Psiloclada clandestina Mitt. SAF(2). The South African taxon was assigned to subsp. spinosa (S.W.Arnell) 

R.M.Schust. by Schuster (504). 

Ptychanthus africanus Steph. 1910 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (972, 158). 

Ptychanthus dioicus (Sim) S.W.Arnell 1953 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (972, 

136). 

Ptychanthus integrifolius Steph. 1912 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (972). 

Ptychanthus pallidus Steph. 1912 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (972). 

Ptychanthus pycnocladus Taylor 1846 = Acrolejeunea pycnoclada (Taylor) Schiffn. 1893 (35). 

Ptychanthus squarrosus Mont. ex Lehm. 1844 = Ptychanthus striatus (Lehm. ex Lindenb.) Nees 1838 (972) 

Ptychanthus striatus (Lehm. et Lindenb.) Nees BUR(2), CMN(2), ETH(2), GUI(48), IVO(51), 

KEN(2,731), MDG(1), MLW(2,563), MOZ(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(2,96), 

ZAI(2), ZAM(2), ZIM(2). 

Ptychanthus tumidus Nees 1838 = Schiffneriolejeunea tumida (Nees) Gradst. 1974 (832). 

Ptychocoleus abnormis (Gottsche) Steph. 1912 = Phragmicoma abnormis Gottsche 1882 (1) [Dubia(1)]. 

Ptychocoleus africanus Steph. 1912 = Ptychocoleus molleri (Steph.) Steph. (834) = Schiffneriolejeunea 


Ptychocoleus amplectens Steph. 1912 = Lejeunea amplectens (Steph.) Steph. 1890 (988, 986) = Schiffneriolejeunea 


Ptychocoleus borgenii Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (35, 833). 

Ptychocoleus confertissimus (Steph.) Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) 

Gradst. 1975 (35). 

Ptychocoleus convexus (Steph.) Steph. 1912 = Evansiolejeunea renistipula (Steph.) Vanden Berghen 1948 

(851) = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960 (850). 

Ptychocoleus emergens (Mitt.) Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus ferrugineus (Steph.) Steph. 1912 = Schiffneriolejeunea ferruginea (Steph.) Gradst. 1974 (832). 

Ptychocoleus flagelliferus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (830, 35). 

Ptychocoleus floribundus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus fulvus (Gottsche) Steph. 1912 = Phragmicoma fulva Gottsche 1882 (988) = Schiffneriolejeunea 

pappeana (Nees) Gradst. 1974 (831) 

Ptychocoleus fuscescens (Gola) Steph. 1912, hom. illeg. [non (Hampe ex Lehm.) Trevis. 1877] = Leucolejeunea 

xanthocarpa (Lehm. et Lindenb.) A.Evans 1907 (988, 35). 

Ptychocoleus grossispicus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus henriquesii Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus inermis Steph. 1912 = Lopholejeunea erugata B.Thiers 1984 [non Lopholejeunea inermis 

Steph. 1912] (893). 

Ptychocoleus inflexus (Gottsche) Steph. 1912 = Phragmicoma inflexus Gottsche 1882 [Dubia(1)].


Ptychocoleus madagascariensis Steph. 1912 = Schiffneriolejeunea madagascariensis (Steph.) Gradst. 1974 

(832). 

Ptychocoleus molleri (Steph.) Steph. 1912 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (35). 

Ptychocoleus occultus (Steph.) Steph. 1912 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974 (832). 

Ptychocoleus orchidae (Pearson) Demaret 1942 = Ptychocoleus molleri (Steph.) Steph. (834) = Schiffneriolejeunea 


Ptychocoleus pappeanus (Nees) Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 (833). 

Ptychocoleus parvilobus (Steph.) Steph. 1912 = Schiffneriolejeunea parviloba (Steph.) Gradst. 1974 (832). 

Ptychocoleus polycarpus (Nees) Trevis. 1877 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974 (832). 

Ptychocoleus pusillus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus quintasii Steph. 1912 = Ptychocoleus pappeanus (Nees) Steph. 1912 (830) = Schiffneriolejeunea 

pappeana (Nees) Gradst. 1974 (833). 

Ptychocoleus renauldii Steph. 1912 = Acrolejeunea aulacophora (Mont.) Steph. 1895 (35). 

Ptychocoleus renauldii var. victoriae E.W.Jones 1954 = Acrolejeunea emergens (Mitt.) Steph. 1895 (35). 

Ptychocoleus roccatii (Gola) Steph. 1912 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960 (850). 

Ptychocoleus rupestris Herzog 1948 = Acrolejeunea emergens (Mitt.) Steph. 1895 (829, 35). 

Ptychocoleus vanderystii Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975 

(830, 35). 

Ptychocoleus yangambiensis Vanden Berghen 1951 = Caudalejeunea yangambiensis (Vanden Berghen) 

E.W.Jones 1957 (841). 

Ptycholejeunea dioica Sim 1926 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838 (972). 

Ptycholejeunea striata (Lehm. et Lindenb.) Steph. 1895 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 

1838 (158). 

Pycnolejeunea africana Steph. 1923 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (867, 872). 

Pycnolejeunea angustiflora Steph. 1914 = Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones 1985 

(877). 

Pycnolejeunea bidentula Steph. in J.B.Jack et Steph. 1894 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 

1980 (582). 

Pycnolejeunea contigua (Nees) Grolle COM(4), GHA(2,14), MDG(1,14), NGA(2,14), RIM(10), 

SIE(2,14), TAN(2,14). 

Pycnolejeunea decurrens Steph. 1914 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (2). 

Pycnolejeunea delessertii (Nees et Mont.) Grolle 1974 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 

1984 (582). 

Pycnolejeunea involuta Steph. 1914 = Cheilolejeunea silvestris var. involuta (Steph.) E.W.Jones 1974 (867) 

= Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (872). 

Pycnolejeunea multiflora (Steph.) Steph. 1914 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 1980 (582). 

Pycnolejeunea nicobarica Steph. 1896 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 1980 (582). 

Pycnolejeunea silvestris (Gottsche) Steph. 1914 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976 (872). 

Pycnolejeunea ventricosa Schiffn. 1898 = Cheilolejeunea ventricosa (Schiffn.) X.-L.He 1999 (fide 14). 

Radula allamanoi Gola 1914 = Radula voluta Taylor ex Gottsche et al. 1845 (963). 

Radula angustata Steph. 1886 = Radula appressa Mitt. 1879 (873). 

Radula ankefinensis Gottsche ex Steph. BIO(36), BUR(2), CMN(2), ETH(2), KEN(661,665,731), 

MAU(1), MDG(1,661,720), MLW(2,114), REU(1,661), RWA(2), SAF(2,661), SWZ(2), 

TAN(2,661), UGA(2,96), ZAI(2,661), ZIM(2). The collections from Ghana and Sierra Leone, reported as 

Radula ?holstiana, could not be named with certainty (25). Grolle (661) stated that past reports from Ethiopia require verification. 

Radula aphylla Mitt. ex Steph., nom. inval. TAN(2). 

Radula appressa Mitt. ANN(2,33), BIO(2,33,72), CAF(2), CMN(2), COM(1), GHA(2), GUI(2), 

IVO(53), KEN(71), MAU(1), MDG(1,4,720), MLW(114), MOZ(2), NGA(2), PRI(2), REU(1,112), 

RIM(10), ROD(1,723), SAO(2), SEY(1), SIE(2), SOC(94), TAN(2), UGA(96), ZAI(2). 

Radula autoica Steph. 1913 = Radula holstiana Steph. 1895 (433) = Radula ankefinensis Gottsche ex Steph. 

1884 (661). 

Radula bipinnata Mitt. 1863 = Radula boryana (F.Weber) Mont. 1842 (660). 

Radula boryana (F.Weber) Mont. ANN(2,33), BIO(2,33,36), CMN(2), COM(1,661), GHA(2), 

KEN(2,661,665,731), MAU(1), MDG(1,661,720), MLW(2,114), PRI(2), REU(1,112,661), 

RIM(10), RWA(2,661), SAF(2,661), SAO(2,661), TAN(2,661), TOG(2), UGA(2), ZAI(2,661), 

ZIM(2,661). The collection from Ethiopia (669) has been redetermined as R. fulvifolia (661). 

Radula caespitosa Steph. 1888 = Radula madagascariensis Gottsche 1882 (873). 

Radula capensis Steph. 1884 = Radula lindenbergiana Gottsche ex C.Hartm. 1864 (873). 

Radula carringtonii J.B.Jack REU(112). 

Radula commutata Gottsche ex J.B.Jack 1881 = R. lindenbergiana Gottsche ex C.Hartm. 1864 (967).


Radula comorensis Steph. COM(1), LES(75), MAU(1), MDG(720), MLW(114), REU(1,112), 

RWA(2), UGA(50), SOC(764), ZAI(2). 

Radula complanata (L.) Dumort. 1831 SAF(2). [Excludenda]. R. complanata has a panboreal distribution and 

Yamada (1041), for example, considered its presence in tropical and subtropical region to be very doubtful. In accordance 

with this view, the sole African record (Mitten, 1877) is here discounted. 

Radula corniculata auct. [non Dumort. 1831] = Plagiochila exigua (Taylor) Taylor 1846 (684). 

Radula delessertii Steph. 1893 possibly = Radula lespangnolii Steph. 1910 (873, 1) [= Radula voluta Taylor 

ex Gottsche et al. 1845 (661)]. 

Radula epiphylla Mitt. ex Steph. 1884 = Radula flaccida Lindenb. et Gottsche 1847 (145). 

Radula evelynae Yamada KEN(2,665,731), REU(1), TAN(2), ZAI(2). 

Radula exigua Steph. 1910 = Radula madagascariensis Gottsche 1882 (661). 

Radula flaccida Lindenb. et Gottsche ANG(2), CMN(2), COM(1), GHA(2), GUI(2), IVO(2), 

NGA(2), PRI(145), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(2,42), ZAI(2). 

Radula foliicola Steph. 1910 = Radula flaccida Lindenb. et Gottsche 1847 (145). 

Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. CAF(661), CMN(661), COM(661), ETH(2,661), 

GHA(661), IVO(728), KEN(2,661,665,731), MAU(661), MDG(661), PRI(661), REU(661,728), 

RIM(661), SAF(2,661,728), STH(2,49,661,728), TAN(2,661), UGA(2,96,661), ZAI(2,661), 

ZIM(661). 

Radula fuscorufa Steph. 1924 = Radula boryana (F.Weber) Nees ex Mont. 1842 (660). 

Radula guineensis Steph. 1884 = Radula appressa Mitt. 1879 (873). 

Radula hallapensis Mont. ex Castle SAF(2). 

Radula hans-meyeri Steph. ex S.W.Arnell 1960, nom. inval. - an orthographic variant of R. meyeri Steph. (989). 

Radula heteroica Steph. 1910 = Radula holstiana Steph. 1895 (433) = Radula ankefinensis Gottsche ex 

Steph. 1884 (661). 

Radula holstiana Steph. 1895 = Radula ankefinensis Gottsche ex Steph. 1884 (661). 

Radula javanica Gottsche 1845 MDG(1), MAU(1), REU(1). [Excludenda(1)]. 

Radula lespagnolii Steph. 1910 = Radula macroloba Steph. 1892 (873) = Radula voluta Taylor ex Gottsche 

et al. 1845 (661). 

Radula lindbergiana Gottsche ex Lindb. 1881, nom. inval. - an orthographic variant of R. lindenbergiana Gottsche 

ex C.Hartm. 

Radula lindenbergiana Gottsche ex C.Hartm. CVI(2,37), LES(75), MLW(114), SAF(2,75), TAN(2), 

ZIM(2). 

Radula macroloba Steph. 1892 = Radula voluta Taylor ex Gottsche et al. 1845 (661). 

Radula macroloba fo. gigantea S.W.Arnell 1957, nom. inval. = Radula allamanoi Gola 1914 (873) = Radula 

voluta Taylor ex Gottsche et al. 1845 (963). 

Radula madagascariensis Gottsche COM(661), KEN(661,665,731), MAU(1,108,661), 

MDG(1,4,661), MLW(114), REU(1,661), TAN(2,661). 

Radula marojezica E.W.Jones MDG(1). 

Radula mascarena Steph. 1884 = Radula appressa Mitt. 1878 (873). 

Radula mauritiana Mitt. ex Steph. 1884 = Radula complanata (L.) Dumort. 1831 (873). 

Radula meyeri Steph. 1888 = Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845 (661). 

Radula molleri Steph. 1910 = Radula appressa Mitt. 1878 (873). 

Radula multiramea Gola 1920 = Radula voluta Taylor ex Gottsche et al. 1845 (2). 

Radula multiramosa Steph. 1924 = Radula boryana (F.Weber) Mont. 1842 (660). 

Radula newtonii Steph. 1910 = Radula stipatiflora Steph. 1910 (435) = Radula voluta Taylor ex Gottsche et 

al. 1845 (661). 

Radula oblongiloba Steph. 1884 = Radula appressa Mitt. 1878 (965). 

Radula pallens (Sw.) Nees et Mont. 1839 MAU(1). [Excludenda(1)]. 

Radula perrottetii Gottsche ex Steph. 1884 COM(1), MDG(1), MAU(1), REU(1). [Excludenda(1)]. 

Radula physoloba Mont. 1843 SAF(2). [Dubia(2)]. The South African record (Mitten, 1877) of this antipodean 

species seems likely to represent another species, probably Radula lindenbergiana (894). 

Radula pirottae Gola 1914 = Radula recurvifolia Steph. 1888 (873) = Radula quadrata Gottsche 1845 

(963). 

Radula pseudoflaccida E.W.Jones TAN(2). 

Radula quadrata Gottsche CMN(2), COM(1), ERI(121), ETH(2), KEN(2,71,665,731,759), MAU(1), 

MDG(1), MLW(114), REU(1), RWA(2), SAF(2), SOC(764), TAN(2), UGA(96), ZAI(2). 

Radula recurvifolia Steph. 1888 = Radula quadrata Gottsche 1845 (963). 

Radula retroflexa Taylor REU(112). 

Radula sarogliae Gola 1914 = Radula boryana (F.Weber) Mont. 1842 (966). 

Radula silvestris Gottsche MDG(1).


Radula socorana Gerola ETH(503). 

Radula spongiosa Steph. 1910 = Radula recurvifolia Steph. 1888 (873) = Radula quadrata Gottsche et al. 

1845 (963). 

Radula stenocalyx Mont. BUR(2), CMN(2), COM(1), GAB(2), GHA(2), GUI(2), KEN(2,731), 

MAU(1), MDG(1,4,720), MLW(114), REU(1), RIM(10), RWA(2), SAF(2), SAO(2), SEY(73), 

SIE(2), TAN(2), UGA(50), ZAI(2). 

Radula stipatiflora Steph. 1910 = Radula voluta Taylor ex Gottsche et al. 1845 (661). 

Radula tabularis Steph. SAF(2,661). 

Radula tamariscina Mitt. 1863 = Radula boryana (F.Weber) Mont. 1842 (661). 

Radula thomeensis Steph. 1910 = Radula boryana (F.Weber) Mont. 1842 (660). 

Radula tubiflora Steph. 1892 ("tubaeflora") = Radula stenocalyx Mont. 1855 (145). 

Radula vaginata Steph. 1910 = Radula stenocalyx Mont. 1855 (145). 

Radula voluta Taylor ex Gottsche et al. BIO(2,33,36), ETH(2,661), KEN(2,71,661,665,731), 

MAU(1), MDG(1,4,661,720), MLW(2,114), REU(1,112), RWA(2), SAF(2,75,661), SOC(764), 

TAN(2,661), UGA(2,96,661), ZAI(2,661). 

Radula wilmsiana Steph. ex Castle, nom. inval. SAF(2). 

Radula woodiana Steph. 1910 = Radula boryana (F.Weber) Mont. 1842 (660). 

Radula xalapensis Nees et Mont. 1836 MAU(1), REU(1). [Excludenda(1)]. 

Reboulia hemisphaerica (L.) Raddi CHA(2,3), CVI(2,37), ERI(2), ETH(2), MLW(2), SOC(2), 

TAN(2). 

Rectolejeunea aloba (Sande Lac.) Steph. 1914 = Lejeunea aloba Sande Lac. 1856 (969). 

Rectolejeunea arnellii E.W.Jones 1974 = Rectolejeunea brittoniae A.Evans 1911 (430) = Lejeunea phyllobola 

Nees et Mont. 1842 (168). 

Rectolejeunea brittoniae A.Evans 1911 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Rectolejeunea pililoba (Spruce) R.M.Schust. 1980 = Lejeunea pililoba Spruce 1895 (430) = Lejeunea 

trinitensis Lindenb.1845 (123). 

Rectolejeunea rhodesiae (Sim) S.W.Arnell 1953 [non Stylolejeunea rhodesiae Sim 1926] = Rectolejeunea 

arnellii E.W.Jones 1974 (868) = Rectolejeunea brittoniae A.Evans 1911 (430) = Lejeunea phyllobola 

Nees et Mont. 1842 (168). 

Rectolejeunea santae-mariae Steph. 1914 probably = Cheilolejeunea intertexta (Lindenb.) Steph. 1897 

(868). 

Rectolejeunea setacea (Steph.) R.M.Schust. 1980 = Lejeunea setacea (Steph.) Steph. 1914 (1). 

Riccardia amazonica (Spruce) Schiffn. ex Gradst. BIO(36), CMN(2), CON(2), GAB(2), GHA(2), 

IVO(51), KEN(2), LES(75), MDG(720), NGA(2), RIM(10), SAF(2,760), SIE(2), TAN(2), 

UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Riccardia angusticosta (Steph.) Grolle 1973 "angusticostata" = Riccardia amazonica (Spruce) Schiffn. ex 

Gradst. 1979 (760, 845). 

Riccardia caespitans (Steph.) E.W.Jones 1956, nom. inval. [= Aneura caespitans Steph. 1891] = Riccardia 

sp. (1). 

Riccardia campanuliflora S.W.Arnell SAF(760). Placed in synonymy with R. amazonica by Arnell (1963), but 

Perold (760) has shown it to be a different taxon. Accordingly, the species is reinstated, pending clarification of its taxonomic 

status. 

Riccardia capensis S.W.Arnell SAF(2). 

Riccardia comosa (Steph.) E.W.Jones 1956, nom. inval. [= Aneura comosa Steph. 1890] = Riccardia sp. (1) 

Riccardia compacta (Steph.) S.W.Arnell SAF(2,717), TAN(2,717), UGA(2). Meenks & Pócs questioned 

the identity of the specimen from Uganda published as R. compacta, but the record is retained under that name for the time 

being. 

Riccardia congoana (Steph.) Demaret 1942 = Riccardia longispica (Steph.) Pearson 1893 (845). 

Riccardia erosa (Steph.) E.W.Jones CMN(2), CON(2), ETH(2), MDG(720), SAO(2), TAN(2). 

Riccardia exigua (Steph.) S.W.Arnell 1957, nom. inval. = Riccardia longispica (Steph.) Pearson 1893 (847). 

Riccardia fastigiata (Lehm.) Trevis. CAF(2), CON(2), ETH(2), KEN(2,665), LES(75), SAF(2,75), 

SEY(4), TAN(2), UGA(96), ZAI(2), ZAM(2), ZIM(2). 

Riccardia holstii (Steph.) E.W.Jones ?CMN(2), TAN(2), UGA(115), ZAI(2), ZIM(2). 

Riccardia kilimandjarica S.W.Arnell TAN(717). R. kilimandjarica was placed in synonymy with R. compacta 

by Meenks & Pócs (1985), but Perold (717) subsequently examined the holotype from Tanzania and was reluctant to accept 

this synonymy. Accordingly, the species is reinstated, pending further clarification. 

Riccardia laurentiana (Steph.) Demaret 1942 = Riccardia pinguis (L.) Gray (845) = Aneura pinguis (L.) 

Dumort. 1831 sens. lat. (652). 

Riccardia lepervanchei (Steph.) S.W.Arnell 1965, nom. inval. [= Aneura lepervanchei Steph. 1899] = Riccardia 

sp. (1).


Riccardia limbata (Steph.) E.W.Jones ANN(2,33), BIO(36), CMN(2), CON(2), GHA(2), 

KEN(2,665), MAU(1), MLW(2), NGA(2), REU(1), RIM(10), RWA(48), SAF(2), SAO(2), SIE(2), 

SWZ(2), TAN(2), TOG(2), ZAI(2), ZAM(2). 

Riccardia longispica (Steph.) Pearson BIO(36), CMN(2), COM(1), MAU(1), MDG(1,720), REU(1), 

RIM(10), SEY(1), TAN(2), UGA(115), ZAI(2). 

Riccardia multifida (L.) Gray MDG(1), SAF(653). 

Riccardia nudiflora (Steph.) Grolle MAU(1), MDG(1), REU(1). 

Riccardia obtusa S.W.Arnell SAF(2,734). 

Riccardia palmata (Hedw.) Carruth. 1865 MDG(1). [Excludenda(1)]. 

Riccardia pinguis (L.) Gray 1821 = Aneura pinguis (L.) Dumort. 1831 (351, 652). 

Riccardia pinnatifida (F.Weber) Carruth. 1865 MDG(1), REU(1). [Excludenda(1)]. 

Riccardia pseudopinguis Herzog 1942 = Aneura pseudopinguis (Herzog) Pócs 1983 (351, 1). 

Riccardia ramosissima (Steph.) E.W.Jones 1956, nom. inval. = Riccardia ramosissima (Steph.) Grolle 1995 

(1). 

Riccardia ramosissima (Steph.) Grolle REU(1). 

Riccardia rhodesiae S.W.Arnell ZIM(2). 

Riccardia saccatiflora (Steph.) S.W.Arnell KEN(2), MAU(1), MDG(1), REU(1,653), RIM(10), 

TAN(2), UGA(96). Perold 2001c (653) showed that Arnell's records of R. saccatiflora from South Africa should be 

referred to R. multifida. 

Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956, hom. illeg. [non S.Hatt. 1944] = Riccardia amazonica 

(Spruce) Schiffn. ex Gradst. 1979 (351). 

Riccardia submarginata S.W.Arnell 1953 (845) = Riccardia pinguis (L.) Gray (845) = Aneura pinguis (L.) 

Dumort. 1831 sens. lat. (652). 

Riccardia sp. (Aneura caespitans Steph.) CMN(2), KEN(2), MAU(1), MDG(1), REU(1), UGA(2). 

Riccardia sp. (Aneura comosa Steph.) MAU(1), REU(1). 

Riccardia sp. (Aneura corbieri Steph.) MDG(1). 

Riccardia sp. (Aneura dismieri Steph.) MDG(1). 

Riccardia sp. (Aneura keniae Gola) KEN(2, 845). 

Riccardia sp. (Aneura lepervanchei Steph.) REU(1). 

Riccardia sp. GHA(25), ?MAU(25), ?RWA(25), ?ZAI(25). 

Riccia abnormis (Steph.) Steph. 1898 = Riccia moenkemeyeri Steph. 1887 (859). 

Riccia aegyptiaca S.W.Arnell 1963 = Riccia congoana Steph. 1898 (547). 

Riccia africana Sim 1926 = Riccia crozalsii Levier 1902 (170). 

Riccia alatospora O.H.Volk et Perold SAF(2). 

Riccia albolimbata S.W.Arnell KEN(2), NAM(2), SAF(2), TAN(2). 

Riccia albomarginata Bisch. ex C.Krauss SAF(2). 

Riccia alboporosa Perold SAF(2). 

Riccia albornata O.H.Volk et Perold SAF(2). 

Riccia albosquamata S.W.Arnell 1957 = Riccia albolimbata S.W.Arnell 1957 (1020). 

Riccia albovestita O.H.Volk NAM(2), SAF(2). 

Riccia ampullacea Perold LES(2,75), SAF(2). 

Riccia angolensis Steph. ANG(2), BOT(2), KEN(2), NAM(2), SAF(2). 

Riccia argenteolimbata O.H.Volk et Perold BOT(2), KEN(2), NAM(2), SAF(2), SOC(764), TAN(2). 

Riccia atromarginata var. jovet-astiae Rauh et Buchloh MDG(1), SOC(764). 

Riccia atropurpurea Sim ANG(2), BEN(11), BOT(2), GHA(2), MLW(2), NAM(2), NGA(2), 

RWA(2), SAF(2), SWZ(2), TAN(2), UGA(2), ZAM(2), ZIM(2,9). 

Riccia beccarii Steph. ex Chiov., nom. inval. ERI(2). 

Riccia berriei E.W.Jones 1957 = Riccia congoana Steph. 1898 (1014). 

Riccia bicolorata Perold SAF(2). 

Riccia biokoensis Perold BIO(98). 

Riccia bullosa Link ex Lindenb. LES(2,75), SAF(2). 

Riccia campbelliana M.Howe 1899 = Riccia macrocarpa Levier 1894 (1026). 

Riccia canescens Steph. 1898 = Riccia trichocarpa M.Howe 1898 (1025) = Riccia crinita Taylor 1846 

(555). 

Riccia capensis auct. [non Steph. 1913] = Riccia nigrella DC. 1815 (1016). 

Riccia capensis Steph. 1913 = Riccia limbata Bisch. ex C.Krauss 1846 (1016). 

Riccia cavernosa Hoffm. ANG(2), BOT(2), CHA(2), CVI(37), DJI(2), ERI(2), KEN(2), LES(75), 

MDG(1), MOZ(2), MTN(3,725), NAM(2), NGR(2,3), REU(719), SAF(2), SUD(2), TAN(2),


ZAI(2), ZAM(2), ZIM(2). 

Riccia chevalieri Steph. 1917 probably = Riccia moenkemeyeri Steph. 1887 (859). 

Riccia ciliata Hoffm. CVI(2,37). 

Riccia ciliata var. austroafricana S.W.Arnell 1953 = Riccia crozalsii Levier 1902 (1132). 

Riccia ciliata subsp. canescens (Steph.) R.M.Schust. 1992 = Riccia crinita Taylor 1846 (555). 

Riccia ciliata subsp. trichocarpa (M.Howe) R.M.Schust. 1992 = Riccia crinita Taylor 1846 (555). 

Riccia commutata J.B.Jack ex Levier 1894 = Riccia warnstorfii Limpr. ex Warnst. 1885 (1027). 

Riccia compacta Garside SAF(2). 

Riccia concava Bisch. ex C.Krauss SAF(2). 

Riccia congoana Steph. ANG(2), BEN(11), BOT(2), CHA(2), CON(2), GHA(2), KEN(2,665), 

MLW(2), NAM(2), NGA(2), RWA(2), SAF(547), SIE(2), SUD(2), SWZ(2), TAN(2), UGA(2,96), 

ZAI(2), ZAM(2), ZIM(2). 

Riccia convexa Steph. ex Har. 1913, nom. inval. [? non Steph. 1917] = Riccia frostii Austin 1875 (1024). 

Riccia coronata Sim 1926. This species is insufficiently known. The specimen cited by Sim cannot be traced, and 

another specimen, which had been identified as 'near R. coronata' is placed under R. alatospora (Perold 1999g 

(1132)). 

Riccia crassa Nees ex Lindenb. 1829 = Riccia bullosa Link ex Lindenb.1829 (1019). 

Riccia crinita Taylor BOT(2), KEN(2), LES(2), MDG(1), MLW(2), NAM(2), NGR(2), RWA(2), 

SAF(2), SOC(2), TAN(2), UGA(2), ZIM(2). 

Riccia crozalsii Levier SAF(2), TAN(2). 

Riccia crystallina L. emend. Raddi CHA(3), CVI(2,37), ETH(2), SAF(2), TAN(2), ZIM(2). It is uncertain 

whether some early records represent this species or R. cavernosa. 

Riccia crystallina var. angustior sensu Gillet et Jovet-Ast 1957 = Riccia cavernosa Hoffm. 1796 (1025). 

Riccia cupulifera A.V.Duthie SAF(2). 

Riccia curtisii (James ex Austin) Austin CHA(2), SAF(2). 

Riccia dinteri Steph. ex S.W.Arnell 1956, nom. inval. = Riccia stricta (Lindenb.) Perold 1990 (1018). 

Riccia discolor Lehm. et Lindenb. BEN(11), IVO(2,60), KEN(2), MLW(2), NGA(2), NGR(2), 

TAN(2), UGA(2), ZAI(2), ZAM(2). 

Riccia duthieae O.H.Volk et Perold 1985 = Riccia albovestita O.H.Volk 1981 (1017). 

Riccia elongata Perold SAF(2). 

Riccia erubescens Perold ANG(2), MLW(2), MOZ(2), ZAM(2). 

Riccia erythraeae Steph. ex Chiov. 1919, nom. inval. ERI(502), ETH(502). A taxon of uncertain status, 

considered by Gola, 1914 (121) as a possible subspecies of R. lanceolata. 

Riccia esulcata Steph. ex Har. 1913, nom. inval. [non Steph. 1917] = Riccia frostii Austin 1875 (1024). 

Riccia fluitans sens. lat. IVO(2), SAO(2), TAN(2), TOG(2), ZAM(2). 

Riccia fluitans var. δ stricta Lindenb. 1837 = Riccia stricta (Lindenb.) Perold 1990 (1018). 

Riccia frostii Austin CHA(2,3), CVI(2), MTN(2,3), SUD(2). 

Riccia furfuracea Perold SAF(2). 

Riccia garsidei Sim SAF(2). 

Riccia glauca L. ETH(2). 

Riccia gougetiana Durieu et Mont. CVI(2,37). 

Riccia hantamensis Perold SAF(2). 

Riccia hariotii Steph. ex Har., nom. inval. SUD(2). 

Riccia helenae Jovet-Ast REU(1). 

Riccia hirsuta O.H.Volk et Perold SAF(2). 

Riccia hortorum Bory ex Lindenb. MAU(1), REU(1). 

Riccia huebeneriana Lindenb. ETH(2), REU(45). 

Riccia intermedia E.W.Jones 1957 = Riccia discolor Lehm. et Lindenb. 1832 (1015). 

Riccia kassaica Steph. ex E.W.Jones 1957 probably = R. congoana Steph. 1898 (859). 

Riccia lamellosa Raddi CHA(2,3), ERI(2), TAN(2). 

Riccia lanceolata Steph. BEN(2,11), GHA(2), IVO(2,60), MLW(2,114), MOZ(2), NGA(2), NGR(2), 

RWA(2), TAN(2), UGA(2,96), ZAM(2). 

Riccia laxisquamata (Steph.) Steph. ETH(2). 

Riccia limbata Bisch. ex C.Krauss ETH(2), SAF(2). 

Riccia limbatoides O.H.Volk 1984, nom. prov. = Riccia congoana Steph. 1898 (547). 

Riccia macrocarpa Levier SAF(2), SOC(2). 

Riccia mammifera O.H.Volk et Perold SAF(2). 

Riccia membranacea Gottsche et Lindenb. ANG(2), CMN(2), GHA(2), KEN(2), NGA(2), SIE(2), 

SUD(2), TAN(2), ZAI(2).


Riccia microciliata O.H.Volk et Perold ANG(2), BOT(2), MLW(2), MOZ(2), NAM(2), SAF(2), 

SWZ(2), TAN(2), ZIM(9). 

Riccia moenkemeyeri Steph. ANG(2), BEN(11), BIO(33), CAF(2), CMN(2), GAM(2), GHA(2), 

IVO(2,60), MLW(2), NGA(2), RWA(2), SAF(2), SIE(2), TAN(2), ZAI(2), ZAM(2), ZIM(2). 

Riccia montaguensis S.W.Arnell 1952 = Riccia bullosa Link ex Lindenb. 1829 (1019). 

Riccia montana Perold LES(2,75), SAF(2). 

Riccia namaquensis Perold SAF(2). 

Riccia natalensis Sim SAF(2), ZIM(9). 

Riccia natans L. 1759 = Ricciocarpos natans (L.) Corda 1829 (655). 

Riccia negrii Gola 1914 = Riccia crystallina L. 1753 emend. Raddi 1818 (1023). 

Riccia nigerica E.W.Jones CMN(2), NGA(2). 

Riccia nigrella DC. CVI(37), LES(2,75), NAM(2), SAF(2). 

Riccia nigrosquamata E.W.Jones 1957 = Riccia congoana Steph. 1898 (547). 

Riccia okahandjana S.W.Arnell ANG(2), BOT(2), LES(75), MLW(2), MOZ(2), NAM(2), RWA(2), 

SAF(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Riccia papillispora Steph. ERI(2). 

Riccia parvo-areolata O.H.Volk et Perold SAF(2). 

Riccia perssonii Sultan Khan BOT(2), CHA(2), MDG(655), MLI(2), NAM(2). 

Riccia plana Taylor 1846 = Riccia crystallina L. 1753 emend. Raddi 1818 (1023). 

Riccia pottsiana Sim SAF(2). 

Riccia pseudolimbata S.W.Arnell 1957 = Riccia angolensis Steph. 1898 (1028). 

Riccia pulveracea Perold SAF(2). 

Riccia purpurascens Lehm. SAF(2). 

Riccia radicosa Pearson ZAI(2). 

Riccia rautanenii (Steph.) Steph. 1898 = Riccia cavernosa Hoffm. 1796 (1021). 

Riccia rhodesiae S.W.Arnell 1952 = Riccia congoana Steph. 1898 (547). 

Riccia rosea O.H.Volk et Perold ANG(2), BOT(2), KEN(2), NAM(2), SAF(2), TAN(2), UGA(2), 

ZAI(2), ZAM(2), ZIM(11). 

Riccia rubricollis Garside et A.V.Duthie ex Perold SAF(2). 

Riccia runssorensis Steph. ANG(2), BOT(2), NAM(2), SAF(2), TAN(2), UGA(2). 

Riccia saharensis Steph. ex Jovet-Ast NGR(2,3). Erroneously listed from Mauritania in (2). 

Riccia sarcosa O.H.Volk et Perold 1986 = Riccia albovestita O.H.Volk 1981 (1017). 

Riccia schelpei O.H.Volk et Perold SAF(2). 

Riccia schroederi Steph. TAN(2). 

Riccia schweinfurthii Steph. SUD(2). 

Riccia sibayenii Perold SAF(651). 

Riccia simii Perold LES(75), SAF(2). 

Riccia somaliensis Perold SOM(2). 

Riccia sorocarpa Bisch. CHA(2,3), CVI(37), KEN(2), LES(2), SAF(2), TAN(2). 

Riccia sorocarpa var. heegii Schiffn. CVI(2). 

Riccia spongosa S.W.Arnell 1952 = Riccia curtisii (James ex Austin) Austin 1879 (1132). 

Riccia stricta (Lindenb.) Perold ANG(2), BIO(36), BOT(2), BUR(2), CMN(2), ETH(2), GHA(2), 

IVO(2), KEN(2,21,665,731), LES(75), MLW(2), MOZ(2), NAM(2), NGA(2), NGR(2), RWA(2), 

SAF(2), SUD(2), TAN(2), TOG(2), UGA(2,96,776), ZAI(2), ZAM(2), ZIM(2,9). 

Riccia stricta A.V.Duthie ex S.W.Arnell 1963, nom. inval. = Riccia stricta (Lindenb.) Perold 1990 (1018). 

Riccia sudanensis Steph. ex Har., nom. inval. SUD(2). 

Riccia suntica Pettet 1967 = Riccia cavernosa Hoffm. 1796 (1022). 

Riccia symoensii Vanden Berghen ZAI(2). 

Riccia tellinii Gola 1914 = Riccia cavernosa Hoffm. 1796 (1023). 

Riccia terracianoi Gola 1914 = Riccia cavernosa Hoffm. 1796 (1023). 

Riccia tomentosa O.H.Volk et Perold SAF(2). 

Riccia trabutiana Steph. SOC(2,558). 

Riccia trachyglossum Perold LES(2). 

Riccia triangularis Steph. CAF(2). 

Riccia trichocarpa M.Howe 1898 = Riccia crinita Taylor 1846 (555). 

Riccia undulata S.W.Arnell 1952 = Riccia moenkemeyeri Steph. 1886 (25). 

Riccia villosa Steph. SAF(2). 

Riccia vitrea Perold SAF(2).


Riccia volkii S.W.Arnell NAM(2), SAF(2), ZIM(2,9). 

Riccia vulcanicola Eb.Fisch. RWA(2). 

Riccia warnstorfii Limpr. SAF(2). Twice collected from a garden in 1954, but not subsequently recorded from 

South Africa, and considered to have been a short-lived introduction (Perold 1999g (1132)). 

Riccia welwitschii Steph. 1898 = Riccia membranacea Gottsche et Lindenb. 1846 (859). 

Ricciella abnormis Steph. 1891 = Riccia moenkemeyeri Steph. 1887 (859). 

Ricciella laxisquamata Steph. 1895 = Riccia laxisquamata (Steph.) Steph. 1898 (502). 

Ricciella purpurascens (Lehm.) Lehm. et Lindenb. 1832 = Riccia purpurascens Lehm. 1829 (1132). 

Ricciella rautanenii Steph. 1895 = Riccia cavernosa Hoffm. 1796 (1021). 

Ricciella stricta (Lindenb.) Trevis. 1877 = Riccia stricta (Lindenb.) Perold 1990 (1132). 

Ricciella tellinii (Gola) Cufod. 1952 = Riccia tellini Gola 1914 (502) = Riccia cavernosa Hoffm. 1796 

(1023). 

Ricciella terracianoi (Gola) Cufod. 1952 = Riccia terracianoi Gola 1914 (502) = Riccia cavernosa Hoffm. 

1796 (1023). 

Ricciella undulata S.W.Arnell 1952 = Riccia moenkemeyeri Steph. 1886 (1058). 

Ricciella welwitschii Steph. 1901 = Riccia membranacea Gottsche et Lindenb. 1846 (859). 

Ricciocarpos natans (L.) Corda BOT(2), GHA(2), KEN(2,665), MOZ(2), NGA(2), RWA(2), 

SAF(2), TAN(2), UGA(96), ZAI(2), ZAM(2). 

Riella affinis M.Howe et Underw. SAF(2,78). 

Riella alatospora Wigglesworth SAF(2,78). 

Riella capensis Cavers SAF(2,78). 

Riella echinospora Wigglesworth NAM(2), SAF(2,78). 

Riella purpureospora Wigglesworth SAF(2,78). 

Sarcocyphos auritus (Lehm.) Nees 1844 = Anastrophyllum auritum (Lehm.) Steph. 1901 (163). 

Sarcocyphos sparsifolius Lindb. 1868 = Marsupella sparsifolia (Lindb.) Dumort. 1874 (944). 

Sauteria nyikaensis Perold MLW(766). 

Scapania calcicola (S.W.Arnell et J.Perss.) Ingham LES(75). 

Scapania cuspiduligera (Nees) Müll.Frib. LES(75), SAF(75), UGA(776), ZAI(2). 

Scapania esterhuyseniae S.W.Arnell TAN(2), UGA(2). 

Scapania hedbergii S.W.Arnell KEN(2). 

Schiffneriolejeunea altimontana Vanden Berghen RWA(2), TAN(2), UGA(50), ZAI(2). 

Schiffneriolejeunea ferruginea (Steph.) Gradst. CMN(2), NGA(2). 

Schiffneriolejeunea fragilis Gradst. et E.W.Jones CMN(2), CON(2), GAB(2), GHA(2), GUI(2), 

IVO(2), NGA(2), SIE(2), UGA(42), ZAI(2). 

Schiffneriolejeunea madagascariensis (Steph.) Gradst. MDG(1). 

Schiffneriolejeunea occulta (Steph.) Gradst. CON(2), GHA(2), NGA(2), RIM(10), SIE(2). 

Schiffneriolejeunea pappeana (Nees) Gradst. var. pappeana ANG(2), ANN(2,33), BIO(2,33), 

CAF(2), CMN(2), ETH(2), GUI(2), KEN(2,731), MDG(1,720), MLW(2,563), REU(1), RWA(2), 

SAF(2), SAO(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Schiffneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen MAU(1), MDG(1), 

REU(1). 

Schiffneriolejeunea pappeana var. integra Gradst. et Vanden Berghen MDG(1). 

Schiffneriolejeunea parviloba (Steph.) Gradst. MAU(1), MDG(720), REU(1). 

Schiffneriolejeunea polycarpa (Nees) Gradst. ANG(2), BIO(2,33), CAF(2), CON(2), GAB(2), 

GHA(2), GUI(2), MDG(1,720), MLW(563), MOZ(2), NGA(2), SAF(2), SAO(2), SIE(2), TAN(2), 

UGA(42,50,96), ZAI(2), ZAM(2), ZIM(2). 

Schiffneriolejeunea tumida (Nees) Gradst. SEY(1). 

Schisma balboi Gola 1914 = Herbertus sp (2). 

Schisma capense Steph. 1909 = Herbertus capensis (Steph.) Sim 1926 (1). 

Schisma capillare (Steph.) Steph. 1909 = Herbertus capillaris Steph. 1892 (1) [Dubia(1)]. 

Schisma dicranum (Taylor ex Gottsche et al.) Steph. 1895 = Herbertus dicranus (Taylor ex Gottsche et al.) 

Trevis. 1877 (1). 

Schisma doggeltianum Steph. 1909 = Herbertus doggeltianus (Steph.) Demaret 1942 (982). 

Schisma dusenii Steph. 1909 = Herbertus mollis (Steph.) Dusén 1893. 

Schisma grossevittatum Steph. 1909 = Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle 1995 (1). 

Schisma hildebrandtii Steph. 1909 = Herbertus capensis (Steph.) Sim 1926 (847). 

Schisma kilimandjarense Steph. 1909 = Herbertus sp. 

Schisma lobatum Steph. 1911 = Herbertus lobatus (Steph.) Demaret 1942 (982).


Schisma mascarenicum Steph. 1909 = Herbertus mascarenicus (Steph.) S.W.Arnell ex Váňa, in Váňa et al. 

1979 (156). 

Schisma mascarenicum var. kikujense Gola 1914 = Herbertus sp. 

Schisma molle (Steph.) Steph. 1909 = Herbertus mollis (Steph.) Dusén 1893. 

Schisma seriatum Steph. 1909 = Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle 1995 (847). 

Schisma stuhlmannii Steph. 1909 = Herbertus stuhlmannii (Steph.) Demaret 1942 (274). 

Schisma subdentatum Steph. 1901 = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 1877 (761). 

Schistochila alata (Lehm.) Schiffn. MDG(1,4), SAF(2). 

Schistochila borbonica Steph. 1890 = Schistochila neesii (Mont.) Trevis. 1877 (871) = Gottschea neesii 

Mont. 1843 (954). 

Schistochila congoana Steph. CON(2). 

Schistochila engleriana Steph. 1909 = Gottschea engleriana (Steph.) Grolle et Zijlstra 1984 (954). 

Schistochila limbata S.W.Arnell 1957 = Schistochila sphagnoides (Schwägr.) Steph. 1909 (871) = Gottschea 

sphagnoides (Schwägr.) Lindb. 1870 (954). 

Schistochila neesii (Mont.) Lindb. 1872 = Gottschea neesii Mont. 1843 (954). 

Schistochila pauciserrata Kiaer et Pearson 1892 = Schistochila alata (Lehm.) Schiffn. 1893 (720). 

Schistochila piligera Steph. MDG(1,4,720). 

Schistochila sphagnoides (Schwägr.) Steph. 1909 = Gottschea sphagnoides (Schwägr.) Lindb. 1870 (954). 

Schistochila thouarsii (Hook.) Dumort. ex Trevis. 1877 = Schistochila sphagnoides (Schwägr.) Steph. 1909 

(885, 871) = Gottschea sphagnoides (Schwägr.) Lindb. 1870 (954). 

Schistochilaster englerianus (Steph.) H.A.Mill. 1970 = Gottschea engleriana (Steph.) Grolle et Zijlstra 1984 

(954). 

Schistochilaster neesii (Mont.) H.A.Mill. 1970 = Gottschea neesii Mont. 1843 (954). 

Schistochilaster piliger (Steph.) H.A.Mill. 1970 "piligerum" = Schistochila piligera Steph. 1890 (1). 

Sendtnera capillaris (Steph.) Renauld 1898 = Herbertus capillaris Steph. 1892 (1) [Dubia(1)]. 

Sendtnera diclados (Brid. ex F.Weber) Endl. ex Gottsche et al. 1845 = Mastigophora diclados (Brid. ex 

F.Weber) Nees 1838 (27). 

Sendtnera dicrana Taylor ex Gottsche et al. 1845 = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 

1877 (1). 

Sendtnera juniperina Nees 1845 = Herbertus juniperoideus (Sw.) Grolle 1961 (1) [Excludenda(1)]. 

Sendtnera mascarenica Mitt. 1860 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838 (895). 

Sendtnera mollis Steph. 1886 = Herbertus mollis (Steph.) Dusén 1893. 

Simia atroviridis (Sim) S.W.Arnell 1962, nom. inval. = Cylindrocolea atroviridis (Sim) Váňa 1979 (615). 

Solenostoma abyssinicum (Nees) Steph. 1901 = Jungermannia abyssinica Nees 1844 (412). 

Solenostoma atrovirens Steph. 1901 = Jungermannia perloi (Gola) Váňa 1973 (412). 

Solenostoma austroafricanum (S.W.Arnell) S.W.Arnell 1963 = Jungermannia mildbraedii Steph. 1914 

(412). 

Solenostoma belangerianum Steph. 1917 = Notoscyphus belangerianus (Lehm. et Lindenb.) Mitt. 1877 

(416) = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873 (916, 1). 

Solenostoma borgenii (Gottsche ex Pearson) Steph. 1901 = Jungermannia borgenii Gottsche ex Pearson 

1893 (412). 

Solenostoma lacouturei Steph. 1917 = Jungermannia borgenii Gottsche ex Pearson 1893 (412). 

Solenostoma perloi Gola 1914 = Jungermannia perloi (Gola) Váňa 1973 (412). 

Solenostoma sphaerocarpum (Hook.) Steph. 1901 = Jungermannia sphaerocarpa Hook. 1815 (1030). 

Solenostoma stoloniferum (Steph.) S.W.Arnell 1963 = Jungermannia borgenii Gottsche ex Pearson 1893 

(412). 

Southbya nigrella (De Not.) Henriq. SOC(746). 

Sphaerocarpos stipitatus Bisch. ex Lindenb. SAF(2,77). 

Sphenolobopsis pearsonii (Spruce) R.M.Schust. KEN(102), MLW(102). 

Sphenolobus argentinus Steph. 1902 = Lophozia argentina (Steph.) R.M.Schust. 1966 (163). 

Sphenolobus crispifolius Steph. 1924 = Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden 

Berghen 1981 (163). 

Sphenolobus flagellaris (S.Hatt.) Grolle 1963 = Iwatsukia jishibae (Steph.) N.Kitag. 1965 (472). 

Sphenolobus laceratus Steph. 1902 [non Gymnomitrion laceratum (Steph.) Horik.] = Gymnomitrion andinum 

(Herzog) Herzog 1934 (429). 

Sphenolobus minutus (Schreb.) Berggr. 1898 = Anastrophyllum minutum (Schreb.) R.M.Schust. 1949 (943). 

Sphenolobus nigrus Rodway 1916 = Allisoniella nigra (Rodway) R.M.Schust. 1972 (416). 

Sphenolobus perigonialis (Hook f. et Taylor) Steph. 1902 = Andrewsianthus perigonialis (Hook.f. et Taylor) 

R.M.Schust. 1966 (416) [Excludenda(416)]. 

Sphenolobus savioi Gola 1914 = Anastrophyllum auritum (Lehm.) Steph. 1901 (918). 

Spruceanthus marianus (Gottsche) Mizut. 1966 = Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 

1989 [Excludenda(1)].


Sprucella moenkemeyeri Steph. 1886 = Sprucella succida (Mitt.) Steph. 1891 (144, 902) = Lepidozia succida 

Mitt. 1860 (250). 

Sprucella succida (Mitt.) Steph. 1891 = Lepidozia succida Mitt. 1860 (250). 

Sprucella ubangiensis (Steph.) Fulford et J.Taylor 1959 = Lepidozia ubangiensis Steph. (889). 

Stenorrhipis madagascariensis (Steph.) Grolle 1963 = Kymatocalyx madagascariensis (Steph.) Gradst. et 

Váňa 1999 (74). 

Stenorrhipis rhizomatica Herzog 1950 = Kymatocalyx rhizomatica (Herzog) Gradst. et Váňa (74). 

Stephaniella paraphyllina J.B.Jack LES(75), SAF(2,75,100). 

Stictolejeunea africana Vanden Berghen 1950 = S. balfourii (Vanden Berghen) E.W.Jones 1976 (872). 

Stictolejeunea balfourii (Mitt.) E.W.Jones CAF(2), CMN(2), GHA(2), IVO(2,53), MAU(1), 

NGA(2), REU(1), RIM(10), ROD(1), SIE(2). 

Strepsilejeunea acuta (Mitt.) Steph. 1895 = Lejeunea acuta Mitt. 1863 (1). 

Strepsilejeunea brevifissa (Gottsche) Steph. 1895 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 

1963 (878). 

Strepsilejeunea brevifissa sensu Vanden Berghen 1960, p.p. = Cheilolejeunea usambarana (Steph.) Grolle 

1979 (2). 

Strepsilejeunea cardiophylla (Gottsche) Steph. 1913 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 

(579). 

Strepsilejeunea compressa Herzog 1947 = Cheilolejeunea compressa (Herzog) Grolle 1995 (1). 

Strepsilejeunea cordistipula Steph. 1913 = Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones 1985 

(877). 

Strepsilejeunea decemplicata Steph. in Levier 1901, nom. inval. = Cheilolejeunea pluriplicata (Pearson) 

R.M.Schust. 1980 (878). 

Strepsilejeunea decemplicata Steph. in Renauld 1909, nom. inval. = Cheilolejeunea pluriplicata (Pearson) 

R.M.Schust. 1980 (878). 

Strepsilejeunea georgensis S.W.Arnell 1953 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963 

(878). 

Strepsilejeunea inflata Steph. 1913 [non Cheilolejeunea inflata Steph. 1914] = Cheilolejeunea krakakammae 

(Lindenb.) R.M.Schust. 1963 "krakagamae" (1). 

Strepsilejeunea knysnana S.W.Arnell 1953 = Strepsilejeunea krakakammae (Lindenb.) Steph. 1913 (812) = 

Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963 (878). 

Strepsilejeunea krakakammae (Lindenb.) Steph. 1913 = Cheilolejeunea krakakammae (Lindenb.) 

R.M.Schust. 1963 (878). 

Strepsilejeunea usambarana Steph. 1913 = Cheilolejeunea usambarana (Steph.) Grolle 1979 (878). 

Strepsilejeunea vatovae Gerola 1947 presumably = Cheilolejeunea sp. 

Stylolejeunea duncaniae Sim 1926 "duncanii" = Lejeunea duncaniae (Sim) S.W.Arnell 1961 (820). 

Stylolejeunea rhodesiae Sim 1926 = Lejeunea rhodesiae (Sim) R.M.Schust. 1962 (1033). 

Stylolejeunea setacea (Steph.) S.W.Arnell 1953 = Lejeunea setacea (Steph.) Steph. 1914 (865). 

Symbiezidium balfourii (Mitt.) Steph. 1912 = Stictolejeunea balfourii (Mitt.) E.W.Jones 1976 (872). 

Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans MDG(713). 

Symbiezidium madagascariense Steph. MDG(1,4), SEY(1). One of the two collections from Madagascar 

published in (4) subsequently proved to be of S. barbiflorum (713). 

Symbiezidium rotale (Hook.f. et Taylor) Trevis. 1877 = Aureolejeunea rotalis (Hook f. et Taylor) Gradst. et 


Symphyogyna aloysii-sabaudiae Gola 1907 probably = Symphyogyna volkensii Steph. 1900 (829). 

Symphyogyna brasiliensis Nees et Mont. ANG(2), ASC(2), BUR(2), CMN(2), GHA(2), MAU(1), 

MDG(1), MLW(2,29), NGA(2), REU(1), RWA(2), SAF(2), SIE(2), STH(2,49), SWZ(2), TAN(2), 

UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Symphyogyna convoluta Gottsche 1846 = Symphyogyna brasiliensis Nees et Mont. 1836 (970). 

Symphyogyna filicum Nadeaud 1873 = Symphyogynopsis gottscheana (Mont. et Nees) Grolle 1987 (929). 

Symphyogyna gottscheana Mont. et Nees 1846 = Symphyogynopsis gottscheana (Mont. et Nees) Grolle 1987 

(929). 

Symphyogyna harveyana Taylor 1846 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846 (894, 956). 

Symphyogyna lehmanniana Mont. et Nees 1846 = Symphyogyna brasiliensis Nees et Mont. 1836 (970). 

Symphyogyna podophylla (Thunb.) Mont. et Nees CMN(2), ETH(2), KEN(2,71), MAU(1), 

MLW(2,29), REU(1), RWA(2), SAF(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Symphyogyna rhizobola (Schwägr.) Nees 1846 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846 

(956). 

Symphyogyna rigida Steph. 1911 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846 (829). 

Symphyogyna sellae Gola 1907 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846 (7). 

Symphyogyna serrata Mitt. 1875 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986 (956).


Symphyogyna spinosa Lindenb. et Gottsche 1847 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986 

(146). 

Symphyogyna tenuicosta - an orthographic error for Symphyogyna tenuicostata Steph. (in Arnell 1956 (7)). 

Symphyogyna tenuicostata Steph. 1895 = Symphyogyna brasiliensis Nees et Mont. 1836 (970). 

Symphyogyna valida Steph. 1917 = Symphyogyna lehmanniana Nees et Mont. 1846 (829) = Symphyogyna 

brasiliensis Nees et Mont. 1836 (970). 

Symphyogyna volkensii Steph. KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Symphyogynopsis filicum (Nadeaud) Grolle 1986 = Symphyogynopsis gottscheana (Mont. et Nees) Grolle 

1987 (929). 

Symphyogynopsis gottscheana (Mont. et Nees) Grolle BIO(36), CMN(146), COM(1), MAU(1), 

MDG(1), MLW(29), REU(1). 

Symphyomitra africana Steph. 1917 = Lethocolea congesta (Lehm.) S.W.Arnell 1955 (812). 

Symphyomitra glossophylla sensu Renauld 1909 = Lethocolea congesta (Lehm.) S.W.Arnell 1955 (1). 

Symphyomitra glossophylla Spruce 1885 COM(1). [Excludenda(1)]. 

Symphyomitra tabularis S.W.Arnell 1953 = Lethocolea congesta (Lehm.) S.W.Arnell 1955 (812). 

Synhymenium aureonitens Griff. 1849 = Cyathodium aureonitens (Griff.) Mitt. 1886 (119). 

Syzygiella concreta (Gottsche) Spruce KEN(2), MDG(1,720), RWA(2), TAN(2), ZAI(2). 

Syzygiella geminifolia (Mitt.) Steph. BIO(36), ETH(2), MDG(1,4,720), REU(1,4), RWA(2), SAO(2), 

TAN(2), ZAI(2). 

Syzygiella macrocalyx (Mont.) Spruce 1876 MAU(1), REU(1). [Excludenda(1)]. 

Syzygiella ruwenzorensis Steph. 1917 = Syzygiella geminifolia (Mitt.) Steph. 1902 (156). 

Syzygiella securifolia (Nees in Lindenb.) Inoue 1979 PRI(2). [Dubia(2)]. 

Syzygiella subintegerrima (Reinw. et al.) Spruce 1895 REU(1). [Excludenda(1)]. 

Targionia capensis Huebener 1834 = Targionia hypophylla L. 1753 (1027, 169). 

Syzygiella sp. SAF(416). 

Targionia elongata Bisch. ex Gottsche ETH(2). 

Targionia fiorii Gola ERI(121). 

Targionia hypophylla L. CHA(2,3), CMN(2), COM(4), CVI(37), DJI(2), ERI(2), ETH(2,113), 

GHA(2), KEN(2,665), LES(2,75), MDG(1), MLW(2,114), NAM(2), NGA(2), REU(1), SAF(2,170), 

SOC(764), TAN(2), UGA(42,96), ZAI(2), ZAM(2), ZIM(2). 

Targionia hypophylla var. capensis (Huebener) C.Krauss 1846 = Targionia hypophylla L. 1753 (169). 

Targionia lorbeeriana Müll.Frib. MDG(1), KEN(2), REU(1), TAN(2). T. lorbeeriana was placed in synonymy 

with T. hypophylla by Garcia-Zamora et al. 1989 (557), and the conspecificity of the two taxa was supported by 

Boisselier-Dubayle & Bischler 1999 (625). However, Grolle 1995 (1) and Grolle & Long 2000 (160) considered the complex 

far from clarified. 

Targionia orbicularis Schwein. 1821 = Notothylas orbicularis (Schwein.) Sull. 1846 (141). 

Taxilejeunea acuta (Mitt.) Steph. 1914 = Lejeunea acuta Mitt. 1863 (140). 

Taxilejeunea borbonica S.W.Arnell 1965 = Lejeunea eckloniana Lindenb. 1845 (869). 

Taxilejeunea conformis (Mont. et Nees) Steph. ANG(2), BUR(2), CAF(2), CMN(2), COM(1), 

ETH(2), GHA(2), IVO(2), KEN(2,71,731), MAU(1), MDG(1,720), MLW(2,563), REU(1), 

RWA(2), SAF(2,75), SEY(1), SIE(2), TAN(2), TOG(2), UGA(2), ZAI(2). 

Taxilejeunea cuneistipula Steph. CMN(2). 

Taxilejeunea dusenii Steph. 1891 = Lejeunea ramosissima Steph. 1887 (863). 

Taxilejeunea epiphyta Steph. 1914 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Taxilejeunea furcicornuta Grolle MAU(1), MDG(1), REU(1,4). 

Taxilejeunea heterofolia Steph. 1914 = Lejeunea eckloniana Lindenb. 1845 (869). Tixier 1995 (130) treated T. 

heterofolia as a distinct species and recorded it from Rwanda. Chuah-Petiot 1997 (71) recorded it from Kenya. 

Taxilejeunea krakakammae (Lindenb.) Sim 1926 = Strepsilejeunea krakakammae (Lindenb.) Steph. 1913 = 

Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963 (878). 

Taxilejeunea lindblomii Pearson 1924 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (579). 

Taxilejeunea longirostris Steph. 1914 = Lejeunea grossecristata (Steph.) E.W.Jones 1968 (874). 

Taxilejeunea lyratiflora (Steph.) Tixier 1995 = Lejeunea lyratiflora Steph. 1892 (2). 

Taxilejeunea mitracalyx Eifrig 1936 = Lejeunea alata Gottsche 1845 (828). 

Taxilejeunea nicholsonii Pearson 1931 = Lejeunea grossecristata (Steph.) E.W.Jones 1968 (25). 

Taxilejeunea pulchriflora Pearson CAF(2), CMN(2), GHA(2), GUI(2), IVO(2,53), KEN(2), LBR(2), 

RIM(10), RWA(2), SIE(2), TAN(2), TOG(2), UGA(115), ZAI(2). 

Taxilejeunea pusilla Pearson 1924, hom. illeg. [non Steph. 1916] = Lejeunea eckloniana Lindenb. 1845 

(869). 

Taxilejeunea ramosissima (Steph.) Steph. 1914 = Lejeunea ramosissima Steph. 1886 (863). 

Taxilejeunea rotundifolia Steph. 1925 perhaps = Lejeunea caespitosa Lindenb. 1845 (157).


Taxilejeunea sikorae Steph. MDG(1). 

Taxilejeunea trichomanoides (Mont. et Nees) Steph. 1914 = Lejeunea trichomanoides Mont. et Nees 1844 

(989) = Taxilejeunea conformis (Nees et Mont.) Steph. 1914 (863). 

Taxilejeunea vallis-gratiae Steph. LES(75), SAF(2,75). 

Telaranea nematodes (Gottsche ex Austin) M.Howe BIO(72), BUR(2), COM(1), GAB(2), GHA(2), 

KEN(48), MDG(1), MLW(2,92), REU(4), RIM(10), RWA(2), SAF(2), SAO(2), SIE(2), TAN(2), 

TOG(2), UGA(42,50,96), ZAI(2), ZAM(2), ZIM(2). 

Telaranea sejuncta (Ångstr.) S.W.Arnell 1957 = Arachniopsis diacantha (Mont.) M.Howe 1902 (848). 

Telaranea succulenta (Sim) Grolle 1964 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 

2001 (556). 

Telaranea trifida (Steph.) R.M.Schust. CMN(2), RWA(2), UGA(42), ZAI(2). 

Tetralophozia cavallii (Gola) Váňa ETH(2), KEN(2), MLW(114), RWA(2), TAN(2), UGA(2,42,96), 

ZAI(2), ZIM(2). 

Thysananthus africanus (Sim) S.W.Arnell, hom. illeg. [non Steph. 1887] SAF(2). 

Thysananthus africanus Steph. 1887 = Caudalejeunea africana (Steph.) Steph. 1895 (891). 

Thysananthus fruticosus (Lindenb. et Gottsche) Schiffn. 1893 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) 

Lacout. 1908 [Excludenda(1)]. 

Thysananthus moenkemeyeri Steph. 1895, nom. nud. = Caudalejeunea africana (Steph.) Steph. 1895 (886, 

891). 

Thysananthus monoicus Steph. 1912 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845 (972). 

Thysananthus planus Sande Lac. 1855 = Thysananthus retusus (Reinw. et al.) B.Thiers et Gradst. 1989 (1) 


Thysananthus retusus (Reinw. et al.) B.Thiers et Gradst. 1989 MAS(1). [Excludenda(1)]. 

Thysananthus spathulistipus (Reinw. et al.) Lindenb. CMN(2), GAB(2), GHA(2), IVO(2,53), 

MAU(1), MDG(1,720), NGA(2), REU(1), RIM(10), SEY(1,4), SIE(2), ZAI(2). See also comment in 

(969). 

Thysananthus triquetrus (Mitt.) Steph. 1912 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845 

(25). 

Thysanolejeunea africana Sim 1926 = Thysananthus africanus (Sim) S.W.Arnell 1953, hom. illeg. [non 

Steph. 1887] (170). 

Thysanolejeunea spathulistipa (Reinw. et al.) J.B.Jack et Steph. 1894 = Thysananthus spathulistipus (Reinw. 

et al.) Lindenb. 1845 (905). 

Trachylejeunea grolleana Pócs 1999 = Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 2001 (149). 

Trachylejeunea serrulata Steph. 1913 = Lepidolejeunea serrulata (Steph.) Grolle 1984 (582). 

Tritomaria camerunensis S.W.Arnell CMN(2). 

Tritomaria exsecta (Schmidel) Loeske ETH(2), KEN(2), MLW(114), RWA(2), SAF(75), TAN(2), 

UGA(2,42). 

Tylimanthus aetiopicus S.W.Arnell 1961 "aetiopica" = Tylimanthus ruwenzorensis S.W.Arnell 1956 (875). 

Tylimanthus africanus Pearson SAF(2). 

Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. STH(2,49). 

Tylimanthus chenagonii Steph. 1905 = Plagiochila integerrima Steph. 1886 (931). 

Tylimanthus congoensis Steph. 1922 = Plagiochila integerrima Steph. 1886 (931). 

Tylimanthus lespagnolii Steph. 1905 = Plagiochila integerrima Steph. 1886 (931). 

Tylimanthus limbatus Steph. 1911 = Marsupidium limbatum (Steph.) Grolle 1965 (538). 

Tylimanthus madagascariensis Steph. 1901, nom. inval. = Kymatocalyx madagascariensis (Steph.) Gradst. 

et Váňa 1999 (74). 

Tylimanthus retusus Steph. 1922 = Plagiochila integerrima Steph. 1886 (931). 

Tylimanthus ruwenzorensis S.W.Arnell ETH(2), KEN(2), MLW(114), REU(1,4), RWA(2), TAN(2), 

UGA(2,96). A collection of Tylimanthus from Ghana could not be named with certainty (25). 

Tylimanthus schweinfurthii Steph. 1905 = Plagiochila integerrima Steph. 1886 (931). 

Tylimanthus wilmsii Steph. 1922 = Marsupidium limbatum (Steph.) Grolle 1965 (538). 

Wiesnerella denudata (Mitt.) Steph. REU(1). 

Xylolejeunea grolleana (Pócs) X.-L.He et Grolle MDG(5,149). 

Xylolejeunea muricella X.-L.He et Grolle SEY(149).


References 

The full literature reference can be found by translating the numerical reference shown in brackets after 

the country (or in additional notes on the taxon) to the author + year equivalent and crossreferencing 

with the literature list below. 

1 Grolle 1995 

2 Wigginton & Grolle 1996 

3 Ros et al. 1999 

4 Pócs 1995 

5 Pócs 1999b 

6 Pócs 1997 

7 Arnell 1956a 

9 During & Moyo 1999 

10 Infante & Heras 1998 

11 Frahm & Porembski 1998 

13 Een & Kristoferson 1999 

14 He 1999 

16 Augier 1985 

17 Anon 1994 

19 Bornefeld et al. 1996 

20 Cardoso 1915 

21 Chuah-Petiot 1996 

22 Dilg & Frahm 2000 

23 Wigginton 2000 

25 Jones & Harrington 1983 

26 Tixier 1977 

27 Vanden Berghen 1973 

28 Benedix 1953 

29 Wigginton & Porley 2001 

33 Heras Pérez & Infante 

Sánchez 1996 

34 Seaward et al. 1996 

35 Gradstein 1975 

36 Müller 1996 

37 Frahm et al. 1996 

38 Infante et al. 1999 

39 Infante & Heras 1999 

42 Pócs & Lye 1999 

43 Een 2000 


48 Müller et al. 2000 


50 Porley et al. 1999 

51 Müller & Schaefer- 

Verwimp 1999 

52 Fischer 1999 

53 Gyarmati 2001 

55 Gradstein & Geissler 1997 

60 Frahm et al. 1996 


72 Infante et al. 1997 

73 O'Shea et al. 1997 

74 Gradstein & Váňa 1999 

75 Hodgetts et al. 1999 

76 Perold 1999f 

77 Perold 2000a 

78 Perold 2000b 


80 Perold 1997c 

81 Perold 1997d 

82 Perold 1997e 

83 Perold 1997f 









92 Hodgetts & Pócs 1999 

93 Long 2001 

94 Kürschner 2000 

95 Mitten 1863 

96 Lye & Pócs 1997 

97 Engel 1980 


99 Vanden Berghen 1953b 

100 Schmitt & Winkler 1968 

101 Tixier 1983 

102 Váňa 1998 

103 Pócs & Streimann 1999 

104 Jones 1985b 

106 Jones 1968b 

107 Scott 1960 

108 Pócs & Tóthmérész 1997 

109 Hodgetts 1997 

110 Porley 1997 

111 Váňa 1999 

112 Arts & Yamada 1998 

113 Wigginton 2001a 

114 O'Shea et al. 2001 

115 Wigginton et al. 2001 

117 Pócs & Lücking 1996 

118 Schuster 1995a 

119 Srivastava & Dixit 1996 

120 Gola 1914a 

121 Gola 1914b 


123 Grolle 1981b 

124 Jovet-Ast 1946 

125 Muhle 1982 

126 Geissler 1997 

127 Oyesiku & Egunyomi 2001 

128 Stephani 1888 

129 Pócs 1999a 

130 Tixier 1995 

131 Hasegawa 1995 

132 Bischler-Causse 1993 

133 Kruijt & Gradstein 1986 

134 Jones 1957a 

135 Tixier 1985 

136 Bizot & Pócs 1974 




141 Schuster 1992 

142 Fischer 1993 



145 Castle 1939 

146 Grolle & Piippo 1986 


148 Long & Grolle 1992 

149 He & Grolle 2001 

152 Tixier 1979 


154 Kuwahara 1981 


156 Váňa et al. 1979 

157 Jones 1972 



160 Grolle & Long 2000 

161 Vanden Berghen 1963a 

163 Váňa 1982 

164 Schuster 1995b 

165 Jones 1953c 

166 Brummitt & Powell 1992 


168 Reiner-Drehwald 2000 


170 Arnell 1963 

171 O'Shea 1995 

172 Engel 1978 


253 Bischler 1970 

254 Bischler 1978 

274 Demaret 1942 

313 Grolle 1959 

314 Grolle 1960a 



318 Grolle 1971 


345 Grolle 1979c 

351 Meenks & Pócs 1985 

378 Pócs 1975 

380 Pócs 1976 

382 Pócs 1984a 

383 Pócs 1984b 

384 Pócs 1984c 

401 Schiffner 1942 

412 Váňa 1974a 

414 Váňa 1980 

416 Váňa 1985 

420 Engel & Braggins 1998 


424 Piippo 1991 




433 Castle 1964 

435 Castle 1965 

446 Engel & Grolle 1971 

447 Engel & Schuster 1985 

451 Grolle 1964 

453 Mizutani 1961 

465 Grolle 1972 



486 Kuwahara 1983b 

494 Váňa 1988 


496 Hatcher 1960-61 

502 Cufodontis 1952 

503 Gerola 1947 


507 Bizot & Pócs 1982 

527 Kitagawa 1981 

533 Veltman et al. 2000 


539 Hollis & Brummitt 1992 

540 O'Shea 1999


541 O'Shea 1993 

542 Kruijt 1988 

543 Kuwahara 1983a 

545 Pócs 1978 

547 Perold 1986 

550 Grolle 2002 


556 Engel & Smith Merrill 

2001 

557 Garcia-Zamora et al. 1989 


563 Wigginton 2001b 

571 Grolle & Reiner-Drehwald 

1999 

574 Grolle 1979d 

578 Thiers & Gradstein 1989 

579 Grolle 1991 


582 Piippo 1986 



623 Grolle & Piippo 1984 


625 Boisselier-Dubayle & 

Bischler 1999 


650 Jones 1952 




654 Jones 1953a 


656 Hyvönen & Piippo 1993 

657 Piippo 1993 


659 Pócs 1994a 

660 Castle 1937 

661 Grolle 2001 




667 Eggers et al. 2001 

669 Arnell 1960 

670 Zhu & Grolle 2001 

679 Jones 1990 

683 Proskauer 1954 


692 Al-Gifri et al. 1995 

696 Uribe & Gradstein 1998 

702 Zhu & So 2000 

708 Sass-Gyarmati 2001 

709 Pócs 2001 

712 Pócs & Váňa 2001 

713 Sass-Gyarmati & Pócs 

2002 

715 Arnell 1957c 

716 Perold 1994 


719 Müller 2002 

720 Pócs & Geissler 2002 

722 Jones 1962 

723 Müller & Pócs 2002 

725 Gonzáles-Mancebo & Ros 

2002 


728 Müller et al. 2001 

729 So 2002 

731 Bytebier & Chuah-Petiot 

2002 




736 Pócs 2002a 

738 Pócs 2002b 


745 Grolle 1980 


759 Chuah-Petiot & Pócs 2003 


761 Hodgetts 2003 

763 Grolle & So 2003 

764 Kürschner 2004a 

765 Zhu & Grolle 2003 


767 So 2004 


769 Kuwahara 1983c 

770 Kuwahara & Onraedt 1969 

771 So 2003 

775 Váňa & Müller 2003 

776 O'Shea et al. 2003 

778 Gradstein & da Costa 2003 



790 Dauphin 2000 

795 Slageren 1985 

803 Arnell 1952 


806 Arnell 1953b 

812 Arnell 1955 









823 Arnell 1965 

827 Mitten 1864 

828 Grolle 1977 


830 Jones 1954d 

831 Geissler & Gradstein 1994 


833 Gradstein & Vanden Berghen 

1985 


835 Gradstein & Inoue 1980 


837 Teeuwen 1989 



840 Verdoorn 1930 





845 Jones 1956 

846 Gradstein & Hekking 1979 

847 Grolle 1978 


849 Grolle & Onraedt 1974 


851 Vanden Berghen 1948c 

852 Gradstein et al. 2001 

853 Jones 1953b 

855 Jones 1954a 

856 Jones 1954b 

857 Jones 1954c 

858 Jones 1954e 

859 Jones 1957b 

861 Jones 1963 

862 Jones 1964 

863 Jones 1967 

864 Jones 1968a 

865 Jones 1969 

867 Jones 1974a 

868 Jones 1974b 

869 Jones 1974c 

870 Jones 1975 

871 Jones 1976a 

872 Jones 1976b 

873 Jones 1977 

874 Jones 1979 

875 Jones 1981 

877 Jones 1985a 

878 Jones 1988 

879 Jones 1989 

883 Stephani 1901-1906 

884 Stephani 1907-1909 

885 Stephani 1909-1912 

886 Stephani 1912-1916 

887 Stephani 1917-1925 


889 Pócs 1994b 

890 Stotler et al. 1974 

891 Vanden Berghen 1948d 

892 Váňa 1993 

893 Thiers 1984 

894 Sim 1926 

895 Inoue 1971 

898 Grolle 1964d 

900 Grolle 1966 






907 Vanden Berghen 1951c 


913 Tixier 1979b 

914 Tixier 1989 

915 Pócs 1993a 


917 Gottsche et al. 1845 

918 Váňa 1975 


921 Grolle 1969 


927 Grolle 1984 

928 Grolle 1985 

929 Grolle 1987 

930 Grolle 1988 


936 Reiner-Drehwald 1994 

937 Grolle 1964c 

940 Grolle 1973 

941 Tixier 1984 




947 Sérgio 1987 

948 Lindberg 1847 

949 Yuzawa 1991 

950 Bizot & Pócs 1979


951 Demaret & Vanden Berghen 

1947 


953 Grolle 1970 

954 Grolle & Zijlstra 1984 



959 Grolle & Heinrichs 1999 

961 Heinrichs 2002 

963 Yamada 1993 

965 Castle 1966 

966 Castle 1969 

967 Castle 1968 


969 Pócs 1993b 



972 Verdoorn 1934 

973 Mitten 1886 

974 Gradstein et al. 1983 

975 Smith 1990 


977 Thiers 1983 

979 Bonner 1962-63 

982 Bonner 1964 

984 Bonner 1976 

986 Bonner 1978 

987 Geissler & Bischler 1985 








995 Bapna 1965 

996 Bharadwaj 1975 



999 Frey & Kürschner 1988 



1002 Pócs 1994c 


1004 Bartholomew-Began 1991 



1008 Engel 1976 

1009 Dugas 1928 

1010 Gottsche 1857 

1011 Gottsche 1882 

1012 Thiers 1993 

1013 Jones 1958 

1014 Berrie 1975 

1015 Pandé & Udar 1957 

1016 Perold & Volk 1988 

1017 Perold 1990a 

1018 Perold 1990b 

1019 Perold 1991a 

1020 Perold 1989a 


1022 Jovet-Ast & Bischler 1971 




1026 Sérgio 1991 

Literature 

1027 Müller 1952 (1951-58). 

1028 Perold 1989b 

1029 Perold 1991b 

1030 Váňa 1974b 

1031 Gradstein & van Beek 1985 


1034 Proskauer 1961 

1035 Kitagawa 1969 

1036 Zwickel 1933 

1037 Wigginton 2004 

1038 Gradstein & Terken 1981 


1041 Yamada 1979 


1043 Perold 1994 

1044 Howe 1898 

1045 Scott & Pike 1988 

1046 Grolle & Persson 1966 

1047 Evans 1915 


1058 Perold 1992a 

1060 Perold 1992b 

1069 Kürschner 2004b 

1088 Inoue 1980 

1089 Inoue 1979 

1103 Bharadwaj 1972 


1106 Zhu & So 1998 

1107 Pócs 1994c 

1132 Perold 1999g 

1149 Gola 1907 


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17 Anon. 1994. List of PRE holdings for Lesotho (Computer diskette with files dated 25/03/1994). 

803 Arnell, S.W. 1952. Hepaticae collected in South and West Africa, 1951. New and little-known species. Botaniska 

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804 Arnell, S.W. 1953a. Notes on South African Hepaticae. Revue Bryologique et Lichénologique 22: 3-5. 

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815 Arnell, S.W. 1958a. New Hepaticae from Cameroon Mountain. Svensk Botanisk Tidskrift 52: 63-67. 

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Appendix 1. Changes to the checklist of sub-Saharan liverworts and hornworts 

Edition 2 (September 2004). 

Taxa added to the checklist since edition 1 (February 2002). 

Amphicephalozia geisslerae Pócs et Váňa 

Anthoceros sp. ROD 

Aphanolejeunea mamillata (Ångstr.) Steph. 

Bazzania decrescens var. ambahatrae Pócs 

Caudalejeunea sp. (Dicranolejeunea africana Steph.) 

Cephaloziella biokoensis Váňa et F.Müller 

Ceratolejeunea cornutissima Steph. 

Cololejeunea attilana Pócs 

Cololejeunea chuahiana Pócs 

Cololejeunea peraffinis var. serrulata Benedix 

Cololejeunea succinea Tixier 

Cyathodium africanum Mitt. 

Cylindrocolea ?ugandica 

Denotarisia linguifolia (De Not.) Grolle 

Drepanolejeunea deslooveri Vanden Berghen 

Drepanolejeunea mascarena (S.W.Arnell) R.L.Zhu et 

Grolle 

Drepanolejeunea pentadactyla (Mont.) Steph. 

Folioceros fuciformis Mont. 

Folioceros incurvus (Steph.) D.C.Bharadwaj 

Frullania ericicola Steph. in Bonner 

Frullania kinangopii Gola 1916 

Frullania substricta Steph. 1913 

Kymatocalyx rhizomatica Gradst. et Váňa 

Lejeunea tuberculosa Steph. 

Lophocolea angustistipula Steph. 

Lophozia jamesonii (Mont.) R.M.Schust. 

Metzgeria madagassa Steph. 

Microlejeunea globosa (Spruce) Steph. 

Microlejeunea nyandaruensis Pócs 

Oxymitra incrassata (Brot.) Sérgio et Sim-Sim 

Pellia epiphylla (L.) Corda 

Plagiochila divergens var. capensis (Steph.) E.W.Jones 

Plagiochila divergens var. myriocarpa (Pearson) E.W.Jones 

Plagiochila fracta Pócs ined. 

Plagiochila cf. rodriguezii Steph. 

Plagiochila subalpina Steph. 

Plagiochila sp. (sect. Choachinae) 

Plagiochila sp. (sect. Zonatae) 

Plicanthus birmensis (Steph.) R.M.Schust. 

Plicanthus giganteus (Steph.) R.M.Schust. 

Plicanthus hirtellus (F.Weber) R.M.Schust. 

Plicanthus sp. (Chandonanthus perloi Gola) 

Riccardia campanuliflora S.W.Arnell 

Riccardia kilimandjarica S.W.Arnell 

Riccia erythraeae Steph. ex Chiov. 

Sauteria nyikaensis Perold 

Southbya nigrella (De Not.) Henriq. 

Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans 

Targionia lorbeeriana Mull.Frib. 

Changes to synonymy since edition 1. The following list includes new synonyms, transfers of an owner to another 

genus, amendments, overlooked synonyms and some removals. 

Acolea lacerata Steph. 1917 = Gymnomitrion laceratum (Steph.) Horik. 1943. 

Acrolejeunea angustispica Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Acrolejeunea renauldii (Steph.) Steph. 1912 = Acrolejeunea aulacophora (Mont.) Steph. 1895. 

Aitonia appendiculata (Lehm. et Lindenb.) Kuntze 1891 = Plagiochasma appendiculata Lehm. et Lindenb. 1832. 

Alicularia flexuosa (Lehm.) Nees 1844 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Anastrophyllum crebrifolium (Hook.f. et Taylor) Steph. = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Anastrophyllum leucocephalum (Taylor) Steph. 1893 = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Andrewsianthus jamesonii(Mont.) Váňa 1983 = Lophozia jamesonii (Mont.) R.M.Schust. 2002. 

Andrewsianthus kilimanjaricus (S.W.Arnell) Grolle et Váňa 1980 = Lophozia jamesonii (Mont.) R.M.Schust. 2002. 

Aneura angusticosta Steph. 1913 "angusticostata" = Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956 (845) = Riccardia 

amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura congoana Steph. 1917 = Riccardia longispica (Steph.) E.W.Jones 1956. In (2) and edition 1 of this checklist, A. 

congoana was erroneously given as a synonym of Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura congoensis Steph. 1917 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. In (2) and edition 1 of this checklist, 

A. congoensis was erroneously given as a synonym of Riccardia longispica (Steph.) E.W.Jones 1956. 

Aneura laurentiana Steph. 1917 = Aneura pinguis (L.) Dumort. 1831 sens. lat. It is uncertain whether this taxon should be 

assigned to Aneura pinguis or Aneura pseudopinguis. 

Aneura multifida (L.) Dumort. 1822 = Riccardia multifida (L.) Gray 1821. 

Anthelia hirtella (F.Weber) Steph. 1891 = Plicanthus hirtellus (F.Weber) R.M.Schust. 2003. 

Anthoceros fuciformis Mont. 1843 = Folioceros fuciformis (Mont.) D.C.Bharadwaj 1975. 

Anthoceros incurvus Steph.1893 = Folioceros incurvus (Steph.) D.C.Bharadwaj 1972. 

Aphanolejeunea mamillata sens. auct. [non (Ångstr.) Steph. 1916] perhaps = Aphanolejeunea capensis (S.W.Arnell) 

S.W.Arnell 1957. 

Archilejeunea palustris Sim 1926 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Blepharostoma sejunctum Ångstr. 1877 = Arachniopsis diacantha (Mont.) M.Howe 1902. Omitted in error from edition 1.


Brachiolejeunea natalaensis Sim 1926 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Bryopteris madagascariensis Steph. 1912 = Bryopteris gaudichaudii Gottsche 1857. 

Bryopteris madagassa Steph. 1902 = Bryopteris gaudichaudii Gottsche 1957. 

Capillolejeunea mascarena S.W.Arnell 1965 = Drepanolejeunea mascarena (S.W.Arnell) R.L.Zhu et Grolle. 

Cephalozia fissa Steph. = Cephalozia connivens ssp. fissa ((Steph.) Váňa 1988. 

Cephaloziella flagellaris S.Hatt. 1948 = Iwatsukia jishibae (Steph.) N.Kitag. 1965. 

Cephaloziella jishibae (Steph.) S.Hatt. 1952 = Iwatsukia jishibae (Steph.) N.Kitag. 1965. 

Cephaloziella radicans Sim 1926 [= Cephaloziella kiaeri (Austin) Douin 1920]. Removed. A transcription error for Cephalozia 

radicans in (2) and in edition 1. 

Ceratolejeunea cornutissima Steph. 1892 perhaps = Ceratolejeunea diversicornua Steph. 1891. The synonymy only suggested 

in (733) - see note in the main list. 

Ceratolejeunea usambarensis Steph. 1913 = Ceratolejeunea calabariensis Steph. 1895. 

Chandonanthus birmensis Steph. 1899 = Plicanthus birmensis (Steph.) R.M.Schust. 2002. 

Chandonanthus giganteus Steph. 1911 = Plicanthus giganteus (Steph.) R.M.Schust. 2002. 

Chandonanthus hirtellus (F.Weber) Mitt. 1873 subsp. hirtellus = Plicanthus hirtellus (F.Weber) R.M.Schust. 2002. 

Chandonanthus hirtellus subsp. giganteus (Steph.) Vanden Berghen 1965 = Plicanthus giganteus (Steph.) R.M.Schust. 2002. 

Chandonanthus perloi Gola 1914 = presumed Plicanthus sp., probably P. hirtellus. 

Cheilolejeunea multiflora Steph. 1895 = Lepidolejeunea multiflora (Steph.) R.M.Schust. 1980. 

Chiloscyphus angustistipulus (Steph.) J.J.Engel et R.M.Schust. 1984 = Lophocolea angustistipulus Steph. 1922. 

Cololejeunea crenatiflora Steph. 1891, nom. inval. perhaps = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897 (1). See 

note in main list. 

Cololejeunea crenatiflora (Steph.) Vanden Berghen 1972 perhaps = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. See 


Cololejeunea crenatiflora var. orientalis Tixier 1985 perhaps = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. See 


Cololejeunea microscopica (Taylor) Schiffn. 1985 = Aphanolejeunea microscopica (Taylor) A.Evans 1911. 

Cyathodium africanum Mitt. 1886 = Cyathodium cavernarum Kunze 1834. Removed. See note under C. africanum in main 

list. 

Dicranolejeunea atra (Mitt.) Vanden Berghen 1953 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988. 

Dicranolejeunea usambarensis Steph. 1912 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988. 

Diplasiolejeunea cornuta var. laciniata Herzog 1947 = Diplasiolejeunea cornuta Steph. 1916. 

Diplasiolejeunea ocellata Steph. 1916 = Diplasiolejeunea cavifolia Steph. 1895. 

Diplasiolejeunea tridentata 1979 = Diplasiolejeunea aulae E.W.Jones 1974. 

Drepanolejeunea africana Steph. 1913 possibly = Drepanolejeunea capulata (Taylor) Steph. 1913. 

Drepanolejeunea claviformis Steph. 1896 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea deslooveri Vanden Berghen 1977 = Drepanolejeunea hamatifolia (Hook.) Schiffn. 1893. Removed. See 

note under Drepanolejeunea deslooveri. 

Drepanolejeunea micholitzii Steph. 1913 = Drepanolejeunea pentadactyla (Mont.) Steph. 1913. 

Dumortiera denudata Mitt. 1861 = Wiesnerella denudata (Mitt.) Steph. 1899. 

Eulejeunea aloba (Sande Lac.) Schiffn. = Lejeunea aloba Sande Lac. 1856. 

Eulejeunea camerunensis Steph. 1923 = Lejeunea tuberculosa Steph. 1915. 

Eulejeunea microclada Pearson 1931 perhaps = Lejeunea tuberculosa Steph. 1915. 

Fimbriaria gigantea Steph. 1899 = Asterella sp. 

Fimbriaria pirottae Gola 1914 = Asterella sp. 

Frullania crispistipula Steph. 1894. Erroneously as Frullania crististipula in edition 1 of this checklist. 

Frullania ericicola Steph. in Bonner 1965 perhaps = Frullania trinervis Steph. 1894. 

Frullania kinangopii Gola 1916 perhaps = Frullania apicalis Mitt. 1879. 

Frullania silvestris Sim 1926, nom. illeg. [non Steph. 1908]. Synonymy uncertain - see note under the species. 

Frullania substricta Steph. 1913 probably = Frullania angulata Mitt. 1863. 

Herbertus subdentatus (Steph.) Fulford should not be applied to African collections, which should be referred to H. subdentatus 

auct. 

Herbertus subdentatus auct. [non (Steph.) Fulford] = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 1877. 

Homalolejeunea acutiloba (Taylor) Steph. 1890 = Marchesinia brachiata (Sw.) Schiffn. 1893. Erroneously transcribed as 

'acutifolia' in (2) and in edition 1. 

Hygrolejeunea punctata Herzog 1951 = Harpalejeunea punctata (Herzog) S.W.Arnell 1965. 

Jungermannia arguta Reinw. et al. 1824 = Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910. 

Jungermannia bursata Desv. 1814, nom. illeg. = Plagiochila bursata Lindenb. 1840, nom. illeg. = Plagiochila aerea Taylor 

1846. 

Jungermannia connata Sw. 1788 = Lophocolea connata (Sw.) Mont. et Nees 1839. 

Jungermannia dichotoma var. Gottsche et al. 1844 = Plagiochila pectinata Willd. ex Lindenb. 1839. 

Jungermannia dominicensis Spruce 1895 = Kymatocalyx dominicensis (Spruce) Váňa 1970. 

Jungermannia flexuosa Lehm. 1829 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Jungermannia gracilis Reinw. et al. 1824 = Frullania gracilis (Reinw. et al.) Gottsche et al. 1845. 

Jungermannia hians Lehm. et Lindenb. in Gottsche et al. 1847 = Frullania arecae (Spreng.) Gottsche 1863. 

Jungermannia hirtella F.Weber 1815 = Plicanthus hirtellus (F.Weber) R.M.Schust. 2002. 

Jungermannia jamesonii Mont. 1856 = Lophozia jamesonii (Mont.) R.M.Schust. 2002. 

Jungermannia laevigata Schrad. 1797 = Porella arboris-vitae (With.) Grolle 1969. 

Jungermannia mollis Nees 1830 = Lophocolea mollis (Nees) Gottsche et al. 1845. 

Jungermannia multifida L. 1753 = Riccardia multifida (L.) Gray 1821. 

β


Jungermannia oenops Lindenb. et Gottsche, in Gottsche et al. 1845 = Jamesoniella colorata (Lehm. ) Schiffn. 1893. 

Jungermannia pallens Sw. 1788 = Radula pallens (Sw.) Nees et Mont. 1839. 

Jungermannia pearsonii Spruce 1881 = Sphenolobopsis pearsonii (Spruce) R.M.Schust. 1972. 

Jungermannia retusa Reinw. et al. 1824 = Thysananthus retusus (Reinw. et al.) B.Thiers et Gradst. 1989. 

Jungermannia subintegerrima Reinw. et al. 1824 = Syzygiella subintegerrima (Reinw. et al.) Spruce 1895. 

Jungermannia subintegra Hook.f. et Taylor 1845 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Jungermannia ulothrix Schwägr. 1814 = Lepidozia ulothrix (Schwägr.) Lindenb. in Gottsche et al. 1845. 

Jungermannia vallis-gratiae Gottsche ex Steph. 1908 = Cephalozia bicuspidata (L.) Dumort. 1835. 

Jungermannia wondraczekii Corda in Sturm 1830 = Fossombronia wondraczekii (Corda) Dumort. ex Lindb. 1873. 

Lejeunea aberrans Lindenb. et Gottsche 1845 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988. 

Lejeunea atra Mitt. 1886 = Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt 1988. 

Lejeunea aulacophora (Mont.) Steph. 1890 = Acrolejeunea aulacophora (Mont.) Steph. 1895. 

Lejeunea barbiflora Lindenb. et Gottsche 1851 = Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans 1908. 

Lejeunea camerunensis (Steph.) E.W.Jones 1972 = Lejeunea tuberculosa Steph. 1915. 

Lejeunea cuculliloba (Steph.) Steph. Nov-Dec 1891, nom. nud. = Lejeunea caespitosa Lindenb. 1845. 

Lejeunea haskarliana (Gottsche) Spruce 1884 "hasskarliana", comb. illeg. = Schiffneriolejeunea tumida var haskarliana 

(Gottsche) Gradst. et Terken 1981. 

Lejeunea longifissa Steph. 1915 perhaps = Lejeunea caespitosa Lindb. 1845. 

Lejeunea minutissima Dumort. 1822, nom. inval. = Cololejeunea minutissima (Sm.) Schiffn. subsp. minutissima 1893. 

Lejeunea recurvistipula Gottsche 1845 = Caudalejeunea recurvistipula (Gottsche) Schiffn. 1893. 

Lejeunea tuberculiflora E.W.Jones ex Pócs 1979, nom. illeg. = Lejeunea tuberculosa Steph. 1915. 

Lepidozia muenchiana Steph. 1909 = Lepidozia cupressina (Sw.) Lindenb. 1845. Removed. This synonymy was made by 

Pócs 1984 (382), but later appparently abandoned (fide 696: 45). 

Lepidozia wallisiana Steph. 1909 = Lepidozia cupressina (Sw.) Lindenb. 1845 (382). Removed. The synonym was given in 

Pócs 1984 (382), but the neotropical L. wallichiana has recently been accepted in later checklists (e.g. 696: 45). 

Lopholejeunea angustifolia Steph. 1912 = Lopholejeunea eulopha (Taylor) Schiffn. 1893. Erroneously transcribed as 'angustiflora' 

in edition 1 of this checklist. 

Lopholejeunea intermedia (Lindenb.) Steph. 1912 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Mannia capensis var. pallida S.W.Arnell 1963 = Mannia capensis (Steph.) S.W.Arnell 1957. 

Marchantia berteroana var. anactis Gottsche 1846 = Marchantia berteroana Lehm. et Lindenb. 1834. 

Marchantia planiloba subsp. walteri Burgeff 1943 = Marchantia pappeana Lehm. subsp. pappeana 1857. 

Mastigophora africana var. annobonensis S.W.Arnell 1956, nom. inval. probably = Mastigophora diclados (Brid. ex 

F.Weber) Nees 1838. Formal synonymy has not been made - see note in main list. 

Mastigophora madagassa Steph. 1909 = Plicanthus hirtellus (F.Weber) R.M.Schust. 2003. 

Metzgeria africana Steph. 1917 = Metzgeria nudifrons Steph. 1892. 

Metzgeria agnewiae Kuwah. 1973 "agnewii" = Metzgeria consanguinea Schiffn. 1893. 

Metzgeria allanii Steph. 1917 = Metzgeria leptoneura Spruce 1885. 

Metzgeria attenuata Steph. 1916 = Metzgeria consanguinea Schiffn. 1893. 

Metzgeria bystroemii S.W.Arnell 1956 = Metzgeria quadrifaria Steph. 1899. 

Metzgeria camerunensis Steph. 1899 = Metzgeria madagassa Steph. 1899. 

Metzgeria capensis S.W.Arnell 1953 = Metzgeria furcata (L.) Dumort 1835. 

Metzgeria convexa Vanden Berghen 1951 = Metzgeria consanguinea Schiffn. 1893. 

Metzgeria decipiens (C.Massal.) Schiffn. 1890 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria elliotii Steph. 1899 = Metzgeria consanguinea Schiffn. 1893. 

Metzgeria furcata var. decipiens C.Massal. 1885 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria hamata Lindb. 1878 [non Metzgeria hamata auct.] = Metzgeria procera Mitt. 1855. 

Metzgeria hedbergii Vanden Berghen 1953 = Metzgeria quadrifaria Steph. 1899. 

Metzgeria latifrons Steph. 1910 = Metzgeria nudifrons Steph. 1892. 

Metzgeria limbato-setosa Steph. 1910 = Metzgeria madagassa Steph. 1899. 

Metzgeria muscicola Steph. 1899 = Metzgeria quadrifaria Steph. 1899. 

Metzgeria nicomariei Veltman et Potgieter 2000 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria perrotana Steph. 1899 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria propagulifera Vanden Berghen 1948 = Metzgeria crassipilis (Lindb.) A.Evans 1909. 

Metzgeria ruwenzorensis Gola 1907 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria tabularis Steph. 1913 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria thomeensis Steph. 1891 = Metzgeria furcata (L.) Dumort. 1835. 

Metzgeria tusii Gola 1914 = Metzgeria leptoneura Spruce 1885. 

Metzgeria vandenberghenii Kuwah. 1983 = Metzgeria consanguinea Schiffn. 1893. 

Metzgeria warnstorfii Steph. 1899 "warnstorffii" = Metzgeria madagassa Steph. 1899. 

Notoscyphus flexuosus (Lehm.) Sim 1926 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Plagiochila amplifolia Steph. 1901 perhaps = Plagiochila fusifera Taylor 1846. 

Plagiochila andongensis Steph. 1904 = Plagiochila divergens var. capensis (Steph.) E.W.Jones 1962. Overlooked in edn. 1. 

Plagiochila anisodonta (Hook.f. et Taylor) Gottsche et al. 1847 = Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. 1875. 

Overlooked in edition 1. 

Plagiochila bursata (Desv.) Lindenb. 1840 nom. illeg. = Plagiochila aerea Taylor 1846. 

Plagiochila capensis Steph. 1904 = Plagiochila divergens var. capensis (Steph.) E.W.Jones 1962. 

Plagiochila grossealata Steph. 1918 = Plagiochila praemorsa Steph. 1886 or Plagiochila moenkemeyeri Steph. 1904. 

Plagiochila maranguana Steph. 1895 = Plagiochila divergens Steph. 1891 = Plagiochila kiaeri Gottsche 1882. 

Plagiochila myriocarpa Pearson 1924 = Plagiochila divergens var. myriocarpa (Pearson) E.W.Jones 1962.


Plagiochila noditexta Steph. 1904 = Plagiochila repanda (Schwägr.) Lindenb. 1840. Corrected. 

Plagiochila pearsoni Steph. 1921 = Plagiochila praemorsa Steph. 1886 or Plagiochila heterostipa Steph. 1892 or Plagiochila 

kiaeri Gottsche 1882. 

Plagiochila perrotana Steph. 1904 = Plagiochila repanda var. perrotana (Steph.) Vanden Berghen 1981. Corrected. 

Ptychanthus squarrosus Mont. ex Lehm. 1844 = Ptychanthus striatus (Lehm. ex Lindenb.) Nees 1838. Overlooked in edn. 1. 

Plagiochila subalpina Steph. 1891, hom. illeg. [non (Nees ex Lindenb.) Mont. et Nees 1838] = Plagiochila pseudo-attenuata 

S.W.Arnell 1962. Removed. 

Plagiochila telekii Steph. 1891 probably = Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen 1981 

or Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981. 

Ptychanthus squarrosus Mont. ex Lehm. 1844 = Ptychanthus striatus (Lehm. ex Lindenb.) Nees 1838. 

Ptychanthus tumidus Nees 1838 = Schiffneriolejeunea tumida (Nees) Gradst. 1974. 

Ptychocoleus fulvus (Gottsche) Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974 

Radula delessertii Steph. 1893 possibly = Radula voluta Taylor ex Gottsche et al. 1845. 

Radula fuscorufa Steph. 1924 = Radula boryana (F.Weber) Nees ex Mont. 1842. 

Rectolejeunea pililoba (Spruce) R.M.Schust. 1980 = Lejeunea trinitensis Lindenb.1845. 

Rectolejeunea rhodesiae (Sim) S.W.Arnell 1953 [non Stylolejeunea rhodesiae Sim 1926] = Lejeunea phyllobola Nees et 

Mont. 1842. 

Riccardia campanuliflora S.W.Arnell 1952 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. (760). Synonymy removed. 

Riccardia congoana (Steph.) Demaret 1942 = Riccardia longispica (Steph.) E.W.Jones 1956. [see also Aneura congoana 

above]. 

Riccardia kilimandjarica S.W.Arnell 1959 = Riccardia compacta (Steph.) S.W.Arnell (717). Synonymy removed. 

Riccia ciliata var. austroafricana S.W.Arnell 1953 = Riccia crozalsii Levier 1902. Overlooked in edition 1. 

Riccia crassa Nees ex Lindenb. 1829 = Riccia bullosa Link ex Lindenb.1829. Overlooked in edition 1. 

Riccia erythraea (Gola) Cufod. 1952 = Riccia lanceolata Steph. 1888. Removed. Erroneous assignment in (2). 

Riccia erythraea Steph. ex Choiv. 1919 = Riccia lanceolata Steph. 1888. Removed. Erroneous assignment in (2). 

Riccia kassaica Steph. ex E.W.Jones 1957 probably = Riccia congoana Steph. 1898. Overlooked in edition 1. 

Riccia lanceolata var erythraea Gola 1914 = Riccia lanceolata Steph. 1888. Removed. Erroneous assignment in (2). 

Ricciella undulata S.W.Arnell 1952 = Riccia moenkemeyeri Steph. 1886. 

Schisma mascarenicum Steph. 1909 = Herbertus mascarenicus (Steph.) S.W.Arnell ex Váňa, in Váňa et al. 1979. 

Schisma molle (Steph.) Steph. 1909 = Herbertus mollis (Steph.) Dusén 1893. Overlooked in edition 1. 

Schisma subdentatum Steph. 1901 = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 1877. 

Schistochila pauciserrata Kiaer et Pearson 1892 = Schistochila alata (Lehm.) Schiffn. 1893. 

Sphenolobus laceratus Steph. 1902 [non Gymnomitrion laceratum (Steph.) Horik.] = Gymnomitrion andinum (Herzog) Herzog 

1934. Wrongly assigned to Gymnomitrion laceratum in (2) and in edition 1. 

Symphyomitra glossophylla sensu Renauld 1909 = Lethocolea congesta (Lehm.) S.W.Arnell 1955. 

Stenorrhipis rhizomatica Herzog 1950 = Kymatocalyx rhizomatica (Herzog) Gradst. et Váňa. 

Symphyomitra glossophylla sensu Renauld 1909 = Lethocolea congesta (Lehm.) S.W.Arnell 1955. 

Targionia lorbeeriana Mull.Frib. 1940 = Targionia hypophylla L. 1753. Removed. 

Taxilejeunea rotundifolia Steph. 1925 perhaps = Lejeunea caespitosa Lindenb. 1845. 

Taxa removed since edition 1. 

Acanthocoleus aberrans var. laevis Gradst. Records are transferred from var. laevis to the species in this edition - see note 

under Acanthocoleus aberrans in main list. 

Acrolejeunea africana Steph., nom. inval. Removed. 

Metzgeria conjugata Lindb. Removed. 

Metzgeria corralensis Steph. Removed. 

Metzgeria epiphylla A.Evans Removed. 

Metzgeria rufula Spruce Removed. 

Metzgeria violacea (Ach.) Dumort. Removed. 

Metzgeria vivipara A.Evans Removed. 

Plagiochila frondescens (Nees) Lindenb. This East Asian species is removed from the African list. 

Plagiochila kiaeri var. capensis (Steph.). Removed. The variety has not been formally transferred to Plagiochila kiaeri, and 

is retained as a variety of Plagiochila divergens. 

Plagiochila kiaeri var. myriocarpa (Pearson). Removed. The variety has not been formally transferred to Plagiochila kiaeri, 

and is retained as a variety of Plagiochila divergens. 

Ptychanthus squarrosus Mont. ex Lehm. Placed in synonymy. 

Radula complanata (L.) Dumort. Removed. 

Country records removed since edition 1. 

Asterella sp. (Fimbriaria preussii Schiffn. ex Steph.) Removed ETH, wrongly inserted for CMN in edition 1. 

Cololejeunea magna (Tixier) Infante et Heras Removed RIM. Included in error in edition 1. 

Cololejeunea platyneura (Spruce) A.Evans Removed SIE. See notes under Cololejeunea peraffinis var. elegans in the main 

list. 

Colura calyptrifolia (Hook.) Dumort. Removed BUR, ZAI and ZIM. The records in (495) and (733) refer to Colura calyptrifolia 

subsp. tenuicornis (A.Evans) Vanden Berghen = Colura tenuicornis (A.Evans) Steph. 

Cylindrocolea ugandica Removed SIE, and reassigned record to Cylindrocolea ?ugandica. 

Drepanolejeunea physaefolia (Gottsche) Steph. Removed GHA. The collection from Ghana, published in Jones & Harrington 

(1983) as Drepanolejeunea ?friesii, was included under Drepanolejeunea vesiculosa in (2).


Mannia androgyna (L. emend. Lindenb.) A.Evans Removed NAM. Reported from Namibia by Arnell (715), but the specimen 

is sterile and unreliably separated from M. capensis. Perold (716) stated that M. capensis is the only southern African 

representative of Mannia. 

Plagiochila pseudoattenuata S.W.Arnell All country records, except SAF, transferred to Plagiochila subalpina. 

Plagiochila barteri Mitt. Removed UGA. Listed in error in edition 1 for Plagiochila barteri subsp. colorans. 

Plagiochila boryana Gottsche ex Steph. Removed MAU. Heinrichs 2002 (961) considered the record to be dubious. 

Plagiochila fusifera Taylor Removed SEN. Listed in error in (2) for SEG (Senegambia). 

Plagiochila wilmsiana Steph. Removed CMN, and assigned to Plagiochila sp. (aff. Plagiochila subalpina). 

Taxa added to the checklist in edition 1. 

Anastrophyllum crebrifolium (Hook.f. et Taylor) Steph. 

Aphanolejeunea microscopica var. africana (Pócs) Pócs et Lücking 

Aphanolejeunea microscopica var. exigua (A.Evans) Pócs et Lücking 

Aphanolejeunea truncatifolia Horik. 

Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 

Asterella khasyana (Griff.) Pandé, K.P.Srivast. et Sultan Khan 

Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 

Cheilolejeunea ventricosa (Schiffn.) X.-L.He 

Cololejeunea cuspidata Steph. ex Tixier 

Cololejeunea iradieri Infante et Heras 

Cololejeunea magna (Tixier) Infante et Heras 

Cololejeunea peraffinis var. elegans Benedix 

Cololejeunea planissima (Mitt.) Abeyw. 

Colura digitalis var. mucronata Pócs 

Colura mauritiana Pócs 

Colura zoophaga Eb.Fischer 

Cyathodium cavernarum Kunze 

Diplasiolejeunea albifolia subsp. orientalis Pócs ined. 

Diplasiolejeunea cobrensis subsp. antsirananae Pócs 

Diplasiolejeunea cogoensis Infante, Heras et Pócs 

Drepanolejeunea geisslerae Pócs 

Drepanolejeunea helenae Pócs 

Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf 

Fossombronia angulifolia Perold 

Fossombronia capensis var. spiralis Perold 

Fossombronia cederbergensis Perold 

Fossombronia elsieae Perold 

Fossombronia gemmifera Perold 

Fossombronia glenii Perold 

Fossombronia hyalorhiza Perold 

Fossombronia marindae Perold 

Edition 1 (February 2002) 

Fossombronia monticola Perold 

Fossombronia nyikaensis Perold 

Fossombronia renateae Perold 

Fossombronia rwandaensis Perold 

Fossombronia spinosa Perold 

Fossombronia straussiana Perold 

Fossombronia swaziensis Perold 

Fossombronia zuurbergensis Perold 

Gymnomitrion andinum (Herzog) Herzog 

Haplomitrium gibbsiae (Steph.) R.M.Schust. 

Kymatocalyx africanus Váňa et Wigginton 

Kymatocalyx dominicensis (Spruce) Váňa 

Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa 

Lejeunea alata var. patriciae Pócs 

Lejeunea phyllobola Nees et Mont. 

Lejeunea trinitensis Lindenb. 

Lepidozia stuhlmannii var. carnosa (Steph.) Pócs 

Lopholejeunea leioptera Gyarmati 

Marchesinia mackaii (Hook.) Gray 

Metzgeria africana Steph. 

Metzgeria nicomariei Veltman et Potgeiter 

Mnioloma fuscum (Lehm.) R.M.Schust. 

Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 

Radula ankefinensis Gottsche ex Steph. 

Radula carringtonii J.B.Jack 

Radula retroflexa Taylor 

Riccia biokoensis Perold 

Riccia sibayenii Perold 

Scapania calcicola (S.W.Arnell et J.Perss.) Ingham 

Sphenolobopsis pearsonii (Spruce) R.M.Schust. 

Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 

Xylolejeunea muricella X.-L.He et Grolle 

Changes to synonymy in edition 1. The following list includes new synonymy, synonymy overlooked in (2), and other corrections. 

Transfers to other genera are also listed, e.g. Anthoceros species to Phaeoceros. 

Acolea africana Steph. 1917 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996. 

Acrobolbus madagascariensis Steph. 1902 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa 1999. 

Archilejeunea rotalis (Hook.f. et Taylor) Steph. 1911 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Aneura keniae Gola 1914 = Riccardia sp. 

Aneura laurentiana Steph. 1917 = Aneura pinguis (L.) Dumort. 1831. 

Aneura submarginata S.W.Arnell 1952 = Aneura pinguis (L.) Dumort. 1831. 

Anthoceros aethyopicus Gola 1914 = Phaeoceros sp. 

Anthoceros brunnthaleri Steph. 1913 = Phaeoceros sp. 

Anthoceros mildbraedii Steph. 1923 = Phaeoceros sp. 

Anthoceros parvifrons Steph. 1912 = Phaeoceros sp. 

Anthoceros rossoi Gola 1914 = Phaeoceros sp. 

Anthoceros tenuissimus Steph. 1893 = Phaeoceros sp. 

Anthoceros tigrinus Gola 1914 = Phaeoceros sp. 

Aphanolejeunea diaphana (A.Evans) R.M.Schust. 1980 = Aphanolejeunea truncatifolia Horik. 1934. 

Aphanolejeunea exigua var. africana Pócs 1984 = Aphanolejeunea microscopica var. africana (Pócs) Pócs et Lücking 1996. 

Aphanolejeunea exigua A.Evans 1911 = Aphanolejeunea microscopica var. exigua (A.Evans) Pócs et Lücking 1996. 

Aphanolejeunea subdiaphana (Jovet-Ast) Pócs 1984 = Aphanolejeunea truncatifolia Horik. 1934. 

Arachniopsis succulenta (Sim) S.W.Arnell 1955 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001. 

Aspiromitus expansus Steph. 1916 = Anthoceros sp. 

Calypogeia capensis S.W.Arnell 1953 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Calypogeia fusca (Lehm.) Steph. 1908 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Calypogeia repandifolia Steph. 1924 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Ceratolejeunea jungneri Steph. 1895 = Ceratolejeunea calabariensis Steph. 1895. 

Cololejeunea harrisii var. magna Tixier 1995 = Cololejeunea magna (Tixier) Infante et Heras 1999. 

Cololejeunea harrisii sensu Vanden Berghen 1977 = Cololejeunea magna (Tixier) Infante et Heras 1999. 

Cololejeunea subdiaphana Jovet-Ast 1947 = Aphanolejeunea truncatifolia Horik. 1934.


Cyathodium africanum Mitt. 1886 = Cyathodium cavernarum Kunze 1834. 

Fimbriaria incrassata Steph. 1899 = Asterella sp. (provisionally treated as A. abyssinica). 

Fimbriaria kamerunensis Steph. 1917 = Asterella sp. (provisionally treated as A. abyssinica). 

Fimbriaria preussii Schiffn. ex Steph. 1895 = Asterella sp. (provisionally treated as A. bachmannii). 

Fimbriaria volkensii Steph. 1899 = Asterella sp. (provisionally treated as A. dissoluta). 

Fossombronia zeyheri Steph. 1900 = Fossombronia crispa Nees 1846. 

Gymnomitrion africanum (Steph.) Horik. 1943 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996. 

Hygrolejeunea patellirostris Steph. 1896 = Lejeunea sp. 

Hygrolejeunea staudtiana Steph. 1914 = Lejeunea sp. 

Jungermannia crispa Spreng. in Gottsche et al. 1846 = Fossombronia crispa Nees 1846. 

Jungermannia fusca Lehm. 1829 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Jungermannia rotalis Hook.f. et Taylor 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Lejeunea brittoniae (A.Evans) Grolle 1995 = Lejeunea phyllobola Nees et Mont. 1842. 

Lejeunea fusca (Lehm.) Drège 1843 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Lejeunea rotalis (Hook.f. et Taylor) Gottsche et al. 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Lepidozia abyssinica S.W.Arnell 1960 = Lepidozia stuhlmannii var. stuhlmannii Steph. 1895. 

Lepidozia carnosa Steph. 1911 = Lepidozia stuhlmannii var. carnosa (Steph.) Pócs 1997. 

Lepidozia stuhlmannii var. abyssinica (S.W.Arnell) Pócs 1994 = Lepidozia stuhlmannii var. stuhlmannii Steph. 1895. 

Lepidozia succulenta Sim 1926 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001. 

Leptocolea cristata var. occidentalis E.W.Jones 1953 = Cololejeunea occidentalis (E.W.Jones) Vanden Berghen 1978. 

Leucolejeunea rotalis (Hook.f. et Taylor) S.W.Arnell 1961 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Madotheca mattirolii Gola 1914 = Porella mattirolii (Gola) S.W.Arnell 1960. 

Marsupella africana Steph. ex Bonner 1953 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996. 

Marsupella capensis S.W.Arnell 1957 = Gymnomitrion andinum (Herzog) Herzog 1934. 

Marsupella hedbergii S.W.Arnell 1956 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996. 

Marsupella subquadrata Steph., in sched. et icones ined. = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996. 

Metacalypogeia fusca (Lehm.) N.Kitag. 1988 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995. 

Metzgeria camerunensis Steph. 1899 = Metzgeria australis Steph. 1889. 

Metzgeria papulosa Steph. 1899 = Metzgeria leptoneura Spruce 1885. 

Metzgeria thomeensis Steph. 1891 = Metzgeria australis Steph. 1889. 

Microlepidozia succulenta (Sim) S.W.Arnell 1962 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001. 

Phragmicoma rotalis (Hook.f. et Taylor) Mitt. 1875 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Pycnolejeunea ventricosa Schiffn. 1898 = Cheilolejeunea ventricosa (Schiffn.) X.-L.He 1999. 

Radula autoica Steph. 1913 = Radula ankefinensis Gottsche ex Steph. 1884. 

Radula delessertii Steph. 1893 = Radula voluta Taylor ex Gottsche et al. 1845. 

Radula exigua Steph. 1910 = Radula madagascariensis Gottsche 1882. 

Radula heteroica Steph. 1910 = Radula ankefinensis Gottsche ex Steph. 1884. 

Radula holstiana Steph. 1895 = Radula ankefinensis Gottsche ex Steph. 1884. 

Radula lespagnolii Steph. 1910 = Radula voluta Taylor ex Gottsche et al. 1845. 

Radula macroloba Steph. 1892 = Radula voluta Taylor ex Gottsche et al. 1845. 

Radula meyeri Steph. 1888 = Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845. 

Radula stipatiflora Steph. 1910 = Radula voluta Taylor ex Gottsche et al. 1845. 

Radula tamariscina Mitt. 1863 = Radula boryana (F.Weber) Mont. 1842. 

Radula woodiana Steph. 1910 = Radula boryana (F.Weber) Mont. 1842. 

Rectolejeunea brittoniae A.Evans 1911 = Lejeunea phyllobola Nees et Mont. 1842. 

Riccardia submarginata S.W.Arnell 1953 = Aneura pinguis (L.) Dumort. 1831. 

Riccia canescens Steph. 1898 = Riccia crinita Taylor 1846. 

Riccia ciliata subsp. canescens (Steph.) R.M.Schust. 1992 = Riccia crinita Taylor 1846. 

Riccia ciliata subsp. trichocarpa (M.Howe) R.M.Schust. 1992 = Riccia crinita Taylor 1846. 

Riccia trichocarpa M.Howe 1898 = Riccia crinita Taylor 1846. 

Schisma balboi Gola 1914 = Herbertus sp. 

Schisma kilimandjarense Steph. 1909 = Herbertus sp. 

Schisma mascarenicum var. kikujense Gola 1914 = Herbertus sp. 

Stenorrhipis madagascariensis (Steph.) Grolle 1963 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa 1999. 

Strepsilejeunea vatovae Gerola 1947 = Cheilolejeunea sp. 

Symbiezidium rotale (Hook.f. et Taylor) Trevis. 1877 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 1997. 

Targionia lorbeeriana Müll.Frib. 1940 = Targionia hypophylla L. 1753. 

Telaranea succulenta (Sim) Grolle 1964 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001. 

Trachylejeunea grolleana Pócs 1999 = Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 2001. 

Tylimanthus madagascariensis Steph. 1901, nom. inval. = Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa 1999. 

Appendix 2. Country abbreviations and numbers of taxa 

The list of geopolitical areas includes 65 countries, part-countries or islands. Country abbreviations follow Hollis 

& Brummitt 1992 (539), except for the four Gulf of Guinea islands and the Republic of South Africa, which are 

not provided with 3-letter abbreviations in the Hollis & Brummitt list. 

The abbreviations for Madagascar (MDG) and Cameroon (CMN) differ from those given in (1) and (2), and 

RIM (Rio Muni) is an additional abbreviation. RHO is adopted for former Rhodesia (Zambia + Zimbabwe) for 

one record where the location is unclear, and SEG for Senegambia (Senegal + Gambia).


The 'Group/Index Muscorum zone' indicates the Index Muscorum zone together with the group name to which it 

belongs in the 'Full name' column, if relevant. The sub-Saharan part of part of Africa-4 is designated Africa-4N 

(= North) to differentiate it from the subantarctic part (not included in this checklist). 

In the table below, the numbers of taxa excludes those marked in the main list as 'Dubia' or 'Excludenda', but 

includes taxa identified only to the genus. 

Abbr. Full name No. of taxa Group/ Index Muscorum zone 

AGA Agalega Islands 0 Africa-3 

ALD Aldabra 2 Seychelles [Africa-3] 

AMI Amirante Islands 0 Seychelles [Africa-3] 

ANG Angola 60 Africa-2 

ANN Annobón Island 37 Equatorial Guinea [Africa-2] 

ASC Ascension Island 14 Africa-2 

BEN Benin 20 Africa-2 

BIO Bioko (Fernando Po) 94 Equatorial Guinea [Africa-2] 

BKN Burkina Faso 0 Africa-2 

BOT Botswana 17 Africa-4N 

BUR Burundi 79 Africa-2 

CAB Cabinda 0 Angola [Africa-2] 

CAF Central African Republic 56 Africa-2 

CGS Chagos Archipelago 0 Africa-3 

CHA Chad 14 Africa-2 

CMN Cameroon 228 Africa-2 

COM Comoro Islands 143 Africa-3 

CON Congo-Brazzaville 49 Africa-2 

CVI Cape Verde 37 Africa-2 

DIE Diego Garcia 0 Chagos Archipelago [Africa-3] 

DJI Djibouti 9 Africa-2 

EQG Equatorial Guinea 0 Africa-2 

ERI Eritrea 22 Africa-2 

ETH Ethiopia 125 Africa-2 

GAB Gabon 50 Africa-2 

GAM Gambia 1 Senegambia [Africa-2] 

GHA Ghana 136 Africa-2 

GNB Guinea Bissau 5 Africa-2 

GUI Guinea 58 Africa-2 

IVO Ivory Coast 95 Africa-2 

KEN Kenya 240 Africa-2 

LBR Liberia 5 Africa-2 

LES Lesotho 82 Africa-4N 

MAS Mascarene Islands 13 Africa-3 

MAU Mauritius 170 Mascarenes [Africa-3] 

MDG Madagascar 376 Africa-3 

MLI Mali 1 Africa-2 

MLW Malawi 231 Africa-2 

MOZ Mozambique 52 Africa-2 

MTN Mauritania 2 Africa-2 

NAM Namibia 31 Africa-4N 

NGA Nigeria 135 Africa-2 

NGR Niger 6 Africa-2 

PRI Príncipe 33 Africa-2 

REU Réunion 237 Mascarenes [Africa-3] 

RHO former Rhodesia 1 Africa-2 

RIM Rio Muni 88 Equatorial Guinea [Africa-2] 

ROD Rodrigues 31 Mascarenes [Africa-3] 

RWA Rwanda 219 Africa-2


SAF South Africa 317 Africa-4N 

SAO São Tomé 85 Africa-2 

SEG Senegambia 1 Africa-2 

SEN Senegal 3 Senegambia [Africa-2] 

SEY Seychelles 110 Africa-3 

SIE Sierra Leone 142 Africa-2 

SOC Socotra 32 Africa-2 

SOM Somalia 4 Africa-2 

STH St. Helena 23 Africa-2 

SUD Sudan 10 Africa-2 

SWZ Swaziland 21 Africa-4N 

TAN Tanzania 395 Africa-2 

TOG Togo 41 Africa-2 

UGA Uganda 295 Africa-2 

ZAI Democratic Republic of Congo 296 Africa-2 

(former Zaire) 

ZAM Zambia 60 Africa-2 

ZIM Zimbabwe 143 Africa-2

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