Annotated list of diatom names by Horst Lange-Bertalot and co ...

Annotated list of diatom names by Horst Lange-Bertalot and co-workers 

Wolf-Henning Kusber & Regine Jahn 

Botanic Garden and Botanical Museum Berlin-Dahlem, Free University Berlin, Königin-Luise Str. 6-8, 14191 Berlin 

e-mail: w.h.kusber@bgbm.org, r.jahn@bgbm.org 

Abstract 

Kusber, W.-H. & Jahn, R. 2002 (2. August 2002): Annotated list of diatom names by Horst Lange-Bertalot and co-workers. – 

Version 2.0. [http://www.algaterra.org/Names_Version2_0.pdf]. 

The aim of this paper is to facilitate access to new diatom names of taxa described, combined or renamed by H. Lange- 

Bertalot and co-workers. These genera, subgenera, species, subspecies, varieties, and formae are listed in alphabetical order. 

For each name the page of the protologue and the figure is cited. New combinations are followed by the basionym, substitute 

names by the replaced name. Comments are given in square brackets. 

Introduction 

Since 1976 H. Lange-Bertalot is publishing papers with new diatom names. Lange-Bertalot and his co-workers (colleagues 

and students) have built up a network of related diatom names and taxonomic concepts by cross-referencing comments, 

corrections, and emendations. This seems to be hermetical for those not familiar with the complete output of the "Lange- 

Bertalot-School". The aim of this paper is to facilitate access to these names. 

The basis for this compilation are the two papers by Metzeltin & Kusber (2001) and Kusber & Metzeltin (2001), improved 

by Prof. H. Lange-Bertalot (Frankfurt, proof reading and contribution to validations) and by Prof. Dr. W. Greuter and Dr. N. 

Kilian (both BGBM Berlin, discussions on nomenclatural problems). 

Nomenclatural comments follow the St. Louis Code (Greuter et al. 2000). Author names were standardized according to 

Brummitt & Powell (1992). Those authors not included in this standard list will be reported to the IPNI-database 

[http://www.ipni.org/index.html]. All data is also made available to the algae names projects at Berkeley University (Index 

Nominum Algarum, University Herbarium, University of California, Berkeley. Compiled by Paul Silva. Available online at 

[http://ucjeps.herb.berkeley.edu/INA.html]) and the California Academy of Science (Diatom genus project, Fourtanier & 

Kociolek 1999, available online at [http://www.calacademy.org/research/diatoms/genproject/index.html]. 

Since this list is work in progress it will be maintained and successively updated at the BGBM Berlin 

[http://www.bgbm.org]. Major corrections and supplementary information will lead to new versions of this list. For special 

interest, older versions will be available in an electronic archive. A paper print out of each version will be deposited at the 

BGBM Library. 

This list is part of the data services provided by AlgaTerra [http://www.algaterra.org/default.htm], a BMBF funded project 

within the BIOLOG-Program. These data are evaluated in order to provide full protologue information of selected data 

subsets, which will be available in 2004 within the AlgaTerra Information System. 

The version 2.0 includes about half of the data of Kusber & Metzeltin (2001), ten publications are newly evaluated 

(Krammer 1980, 1981, Maidana et al. 2002, Reichardt 1984, 1997, Witkowski 1991, 1993 a-c, Wydrzycka & Lange-Bertalot 

2001). 

In the version 2.0 several changes are made to faciliate access. Standardized phrases: 

"≡" indicates that a name is a nomenclatural synonym (i.e. homotypic synonym, based on the same nomenclatural type). 

"Replaced synonym" indicates that the following name is a homotypic synomym (≡), which was either published 

illegitimately or would have resulted in an illegitimate name in a new combination. 

"Basionym" is the name bearing homotypic synonym (≡). 

"[Nomenclatural synonym of ...]" refers to a later homotypic synonym of this name (≡). 

"=" indicates that a name is a taxonomic synonym (i.e. heterotypic synonym, based on a different type). This subjective 

synonym is referenced. 

"[Taxonomic synonym of ...]" refers to a later heterotypic synonym of this name (=). This subjective synonym is referenced. 

"–" indicates a corresponding name, invalidly published (incl. so called "manuscript names").

Kusber & Jahn 2002: Annotated list of diatom names (V. 2.0) 

2 

To ensure data consistency, simultaneously with the version 2.0 [http://www.algaterra.org/Version2_0.pdf], a draft of version 

2.0 is published on the web [http://www.algaterra.org/Version2_0_Draft.pdf]. This "draft" differs from the original version 

by indicating all changes in this version in different colours. Red colour: new datasets, green colour: revised datasets from 

Kusber & Metzeltin (2001); blue colour: changes due to editing including standardization and correction of typographical 

errors. 

Diatom names in alphabetical order 

ACHNANTHEIOPSIS Lange-Bert., Arch. Protistenk 148, p. 200-201 (1997). [Typus generis: Achnantheiopsis lanceolata 

(Bréb.) Lange-Bert., Arch. Protistenk 148, p. 207 (1997). Comment: see 

http://www.calacademy.org/research/diatoms/hanna_db/DtmBrwsRecs.aspObj=vGenRef&Atr=GenusID&Val=999]. 

Achnantheiopsis apiculata (R.M.Patrick) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes lanceolata var. apiculata R.M. Patrick, Farlowia 2, p. 167, pl. 1: figs 4-5 (1945). 

[Nomenclatural synonym of Planothidium apiculatum (R.M.Patrick) Lange-Bert.] 

Achnantheiopsis aueri (Krasske) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes aueri Krasske, Ann. Acad. Scient. Fenn. Ser. A. IV. Biol. 14, p. 77, pl. 86: figs 23-32 (1949). 

[Nomenclatural synonym of Planothidium aueri (Krasske) Lange-Bert.] 

Achnantheiopsis biporoma (M.H.Hohn et Hellerman) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes biporoma M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82, p. 273, pl. 2: figs 5-6 (1963). 

[Nomenclatural synonym of Planothidium biporomum (M.H.Hohn et Hellerman) Lange-Bert.] 

[Comment: Erroneously Trans. Amer. Microscop. Soc. "80" is cited in Arch. Protistenk. 148, p. 206 (1997).] 

Achnantheiopsis borneolanceolata Lange-Bert., Arch. Protistenk 148, p. 208, (1997). [Achnantheiopsis borneolanceolata 

Lange-Bert. was already documented and described in Lange-Bertalot & Krammer (1989) Biblioth. Diatomol. 18, p. 30-31 

pl. 85: figs 11-14 and pl. 91: fig. 2 as "Achnanthes borneolanceolata (Manuskriptname) Lange-Bert.".] 

[Basionym of Planothidium borneolanceolatum (Lange-Bert.) Lange-Bert.] 

Achnantheiopsis chilensis (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes chilensis Hust., Arch. Hydrobiol. 18, p. 238, pl. 7: figs 3-4 (1927). 

[Nomenclatural synonym of Planothidium chilense (Hust.) Lange-Bert.] 

Achnantheiopsis daui (Foged) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes daui Foged, Danmark Geol. Undersoeg. 84, p. 14, pl. 1: fig. 10 (1962). 

[Nomenclatural synonym of Planothidium daui (Foged) Lange-Bert.] 

Achnantheiopsis delicatula (Kütz.) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnantheidium delicatulum Kütz., Kieselschalige Diatomeen, p. 75, pl. 5: fig. 4 (1844). 

Achnantheiopsis distincta (Messik.) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes distincta Messik., Hydrobiologia 6, p. 32, pl. 1: fig. 2 (1954). 

[Nomenclatural synonym of Planothidium distinctum (Messik.) Lange-Bert.] 

Achnantheiopsis dubia (Grunow) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes lanceolata var. dubia Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 32, (1880). 

[≡ Achnanthes lanceolata ssp. dubia (Grunow) Lange-Bert.] 

Achnantheiopsis engelbrechtii (Cholnoky) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes engelbrechtii Cholnoky, Ber. Deutsch. Bot. Ges. 68, p. 16, pl. 16: figs 1-8 (1955). 

[≡ Achnanthes delicatula ssp. engelbrechtii (Cholnoky) Lange-Bert.] 

Achnantheiopsis fonticolanceolata (Lange-Bert. et Schim.) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes fonticolanceolata Lange-Bert. et Schim., Biblioth. Diatomol. 18, p. 57, pl. 88: figs 1-7 (1989). 

[Nomenclatural synonym of Planothidium fonticolanceolatum (Lange-Bert. et Schim.) Lange-Bert.] 

Achnantheiopsis fragilarioides (J.B.Petersen) Lange-Bert., Arch. Protistenk. 148, p. 206 (1997). 

Basionym: Achnanthes fragilarioides J.B.Petersen, Dansk. Bot. Ark. 4, p. 18, fig. 6 (1924). 

Achnantheiopsis frequentissima (Lange-Bert.) Lange-Bert. var. frequentissima, Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes lanceolata ssp. frequentissima Lange-Bert., Biblioth. Diatomol. 27, p. 4 (1993). 

Achnantheiopsis frequentissima var. magna (F.Straub) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes rostrata var. magna F.Straub, Bull. Soc. Neuchâteloise Soc. Nat. 108, p. 139, pl. 10: fig. 142, pl. 11: 

[http://www.algaterra.org/Names_Version2_0.pdf] (2. August 2002)


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fig. 142b (1985). 

[Nomenclatural synonym of Planothidium frequentissimum var. magnum (F.Straub) Lange-Bert.] 

Achnantheiopsis frequentissima var. minor (Schulz-Danzig) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes lanceolata [var. elliptica] f. minor Schulz-Danzig, Bot. Arch. 13, p. 191, fig. 42 (1926). 

[≡ Achnanthes lanceolata var. minor (Schulz-Danzig) Lange-Bert.] 

[Nomenclatural synonym of Planothidium frequentissimum var. minus (Schulz-Danzig) Lange-Bert.] 

Achnantheiopsis grana (M.H.Hohn et Hellerman) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes grana M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82, p. 274, pl. 2: figs 9-10 (1963). 

[Nomenclatural synonym of Planothidium granum (M.H.Hohn et Hellerman) Lange-Bert.] 

Achnantheiopsis hauckiana (Grunow) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes hauckiana Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 21 (1880). 

[≡ Achnanthes delicatula ssp. hauckiana (Grunow) Lange-Bert. et Ruppel.] 

Achnantheiopsis haynaldii (Schaarschm.) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes haynaldii Schaarschm., Magyar. Növ. Lapok 5, p. 20 (1881). 

[Nomenclatural synonym of Planothidium haynaldii (Schaarschm.) Lange-Bert.] 

Achnantheiopsis heteromorpha (Grunow) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes heteromorpha Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 23 (1880). 

[Nomenclatural synonym of Planothidium heteromorphum (Grunow) Lange-Bert.] 

Achnantheiopsis holstii (Cleve) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes holstii Cleve, Kongl. Svenska Vetenskapsakad. Handl. 18, p. 13, pl. 16: figs 6-7 (1881). 

[Nomenclatural synonym of Planothidium holstii (Cleve) Lange-Bert.] 

Achnantheiopsis joursacense (Héribaud) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes joursacense Héribaud, Les Diatomées d’Auvergne, p. 89, pl. 12: fig. 20 (1903). 

[Nomenclatural synonym of Planothidium joursacense (Héribaud) Lange-Bert.] 

Achnantheiopsis lanceolata (Brébisson ex Kützing) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnantheidium lanceolatum Brébisson ex Kützing, Species Algarum, Arch. Protistenk. 148, p. 54 (1849). 

Achnantheiopsis lanceolatoides (Sovereign) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes lanceolatoides Sovereign, Trans. Amer. Microscop. Soc. 77, p. 115, pl. 2: figs 18-19 (1958). 

[≡ Achnanthes lanceolata ssp. lanceolatoides (Sovereign) Lange-Bert.] 

[Nomenclatural synonym of Planothidium lanceolatoides (Sovereign) Lange-Bert.] 

Achnantheiopsis linkei (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes linkei Hust., Abh. Naturwiss. Vereine Bremen 31, p. 607, figs 28-32 (1939). 

[Nomenclatural synonym of Planothidium linkei (Hust.) Lange-Bert.] 

Achnantheiopsis magnifica (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes magnifica Hust., Bot. Not. 4, p. 385, figs 62-65 (1952). 

[≡ Planothidium magnificum (Hust.) Lange-Bert.] 

Achnantheiopsis marina (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes marina Hust., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 415: figs 1-2 (1937). 

[Nomenclatural synonym of Planothidium marinum (Hust.) Lange-Bert.] 

Achnantheiopsis minutissima (Krasske) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes lanceolata var. minutissima Krasske, Arch. Hydrobiol. 33, p. 6, figs 18-22 (1938). 

[Nomenclatural synonym of Planothidium minutissimum (Krasske) Lange-Bert.] 

Achnantheiopsis miota (J.R.Carter) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes miota J.R.Carter, Nova Hedwigia Beih. 73, p. 286, pl. 1: fig. 9 (1982). 

[Nomenclatural synonym of Planothidium miotum (J.R.Carter) Lange-Bert.] 

Achnantheiopsis oestrupii (Cleve-Euler) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes lanceolata var. oestrupii Cleve-Euler, Acta Forest. Fenn. 22, p. 53, pl. 1: fig. 1 (1922). 

[Nomenclatural synonym of Planothidium oestrupii (Cleve-Euler) Round et Bukhtiyarova ex U.Rumrich, Lange-Bert. et 

M.Rumrich.] 

Achnantheiopsis peragalli (Brun et Hérib.) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes peragalli Brun et Héribaud in Héribaud, Les Diatomées d’Auvergne, p. 50, pl. 1: fig. 4 (1893). 



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Achnantheiopsis pericava (J.R.Carter) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes pericava J.R.Carter, Nova Hedwigia 11, p. 457, pl. 1: figs 5-8 (1966). 

[Nomenclatural synonym of Planothidium pericavum (J.R.Carter) Lange-Bert.] 

Achnantheiopsis pseudotanensis (Cleve-Euler) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes pseudotanensis Cleve-Euler, Kongl. Svenska Vetenskapsakad. Handl. 4, p. 25, fig. 524 (1953). 

[Nomenclatural synonym of Planothidium pseudotanense (Cleve-Euler) Lange-Bert.] 

Achnantheiopsis pungens (Cleve-Euler) Lange-Bert., Arch. Protistenk. 148, p. 207 (1997). 

Basionym: Achnanthes elliptica var. pungens Cleve-Euler, Soc. Scient. Fenn. Comm. Biol. 4, p. 38, fig. 40 (1934). 

[Nomenclatural synonym of Planothidium pungens (Cleve-Euler) Lange-Bert.] 

Achnantheiopsis robusta (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 208 (1997). 

Basionym: Achnanthes delicatula var. robusta Hust., Verh. Vortr. Bremer Wiss. Ges. 8-9, p. 378, figs 1-2 (1934). 

[Nomenclatural synonym of Planothidium robustum (Hust.) Lange-Bert.] 

Achnantheiopsis robustior Lange-Bert., Arch. Protistenk. 148, p. 208 (1997). 

Replaced synonym: Achnanthes lanceolata var. robusta Hust., Arch. Hydrobiol. Suppl. 15, p. 202, pl. 13: figs 47-49 (1934). 

[≡ Achnanthes lanceolata ssp. robusta (Hust.) Lange-Bert.] 

[Nomenclatural synonym of Planothidium robustius Lange-Bert.] 

Achnantheiopsis rostrata (Østrup) Lange-Bert., Arch. Protistenk. 148, p. 208 (1997). 

Basionym: Achnanthes rostrata Østrup, Flora of Koh Chang, Bot. Tidskr. 26, p. 253 ("1902", 1904). 

[≡ Achnanthes lanceolata ssp. rostrata (Østrup) Lange-Bert.] 

[Nomenclatural synonym of Planothidium rostratum (Østrup) Lange-Bert.] 

Achnantheiopsis salvadoriana (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 208 (1997). 

Basionym: Achnanthes salvadoriana Hust., Comun. Inst. Trop. Invest. Ci. 5-6, p. 132, fig. 1 (1953). 

[Nomenclatural synonym of Planothidium salvadorianum (Hust.) Lange-Bert.] 

Achnantheiopsis septentrionalis (Østrup) Lange-Bert., Arch. Protistenk. 148, p. 208 (1997). 

Basionym: Achnanthes septentrionalis Østrup, Meddel. Grønland, p. 215, pl. 13: fig. 21 (1910). 

[Nomenclatural synonym of Planothidium septentrionale (Østrup) Round et Bukhtiyarova ex U.Rumrich, Lange-Bert. et 

M.Rumrich.] 

ACHNANTHES Bory 

[http://www.calacademy.org/research/diatoms/hanna_db/DtmBrwsRecs.aspObj=vGenRef&Atr=GenusID&Val=843]. 

Achnanthes acidoclinata Lange-Bert., Iconogr. Diatomol. 2, p. 22, pl. 21: figs 22-24c, pl. 113: figs 1-7 (1996). 

[Basionym of Psammothidium acidoclinatum (Lange-Bert.) Lange-Bert.] 

Achnanthes alpestris (Brun) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 24 (1996). 

Basionym: Achnanthes flexella var. alpestris Brun, Diatomées des Alpes et du Jura et de la Region Suisse et Française des 

Environs de Genève, p. 30, fig. 26 (1880). 

[Nomenclatural synonym of Eucocconeis alpestris (Brun) Lange-Bert.] 

Achnanthes alteragracillima Lange-Bert., Biblioth. Diatomol. 27, p. 2 (1993). 

Replaced synonym: Microneis gracillima F.Meister, Kieselalgen der Schweiz, p. 97, pl. 12: figs 21-22 (1912). 

Achnanthes archibaldiana Lange-Bert., Biblioth. Diatomol. 18, p. 23, pl. 64: figs 7-16, pl. 66: figs 8-11 (1989). 

[Basionym of Achnanthidium archibaldianum (Lange-Bert.) Lange-Bert.] 

Achnanthes biasolettiana var. subatomus Lange-Bert., Biblioth. Diatomol. 18, p. 28, pl. 57: fig. 8, pl. 59: figs 1-11' (1989). 

Achnanthes borneolanceolata Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 30-31 pl. 85: figs 11-14 and pl. 91: fig. 2 

(1989). [Comment: Achnanthes borneolanceolata is lacking a Latin diagnosis and a type, hence it is invalidly published. 

Validation within the genus Achnantheiopsis Lange-Bert. as Achnantheiopsis borneolanceolata Lange-Bert., a synonym of 

Planothidium borneolanceolatum (Lange-Bert.) Lange-Bert.] 

Achnanthes boudoui Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 18, pl. 69: figs 9-14, pl. 70: figs 1, 4 (1998). 

Achnanthes bremeyeri Lange-Bert., Biblioth. Diatomol. 18, p. 31-32, pl. 76: figs 11-12 (1989). 

[Basionym of Astartiella bremeyeri (Lange-Bert.) Witkowski et Lange-Bert. 

Achnanthes butterfassiana Lange-Bert., Biblioth. Diatomol. 18, p. 34-35, pl. 71: figs 15-29, pl.72: figs 2-5 (1989). 

[Basionym of Nupela butterfassiana (Lange-Bert.) Lange-Bert.] 



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Achnanthes carissima Lange-Bert., Biblioth. Diatomol. 18, p. 36, pl. 28: fig. 1, pl. 36: figs 32-34, pl. 38: figs 1-4 (1989). 

[Comment: Achnanthes carissima is only the valve on the right hand side on pl. 28: fig. 1.] 

Achnanthes contrarea Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 34-35, pl. 5: figs 1-19 (1995). 

[Basionym of Achnanthidium contrarea (Lange-Bert. et Steindorf) Lange-Bert.] 

Achnanthes daonensis Lange-Bert., Biblioth. Diatomol. 18, p. 43, pl. 19: figs 27-32´, pl. 25: figs 1-6, pl. 95: figs 1-13 

(1989). 

[Basionym of Psammothidium daonense (Lange-Bert.) Lange-Bert.] 

Achnanthes delicatula ssp. engelbrechtii (Cholnoky) Lange-Bert., Biblioth. Diatomol. 27, p. 3 (1993). 

Basionym: Achnanthes engelbrechtii Cholnoky, Ber. Deutsch. Bot. Ges. 68, p. 16, figs 1-8 (1955). 

[Nomenclatural synonym of Achnantheiopsis engelbrechtii (Cholnoky) Lange-Bert.] 

Achnanthes delicatula ssp. hauckiana (Grunow) Lange-Bert. et Ruppel, Algol. Stud. 26, p. 6 (1980). 

Basionym: Achnanthes hauckiana Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 21 (1880). 

[Nomenclatural synonym of Achnantheiopsis hauckiana (Grunow) Lange-Bert.] 

Achnanthes dornii Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 49, pl. 38: figs 5-7, pl. 74: figs 10-12' (1989). 

[Comment: lacking a type and a Latin diagnosis, this name was invalidly published.] 

Achnanthes eutrophila Lange-Bert., Iconogr. Diatomol. 2, p. 25, pl. 78: figs 29-38 (1996). 

[Basionym of Achnanthidium eutrophilum (Lange-Bert.) Lange-Bert.] 

Achnanthes fonticolanceolata Lange-Bert. et Schim., Biblioth. Diatomol. 18, p. 57, pl. 88: figs 1-7 (1989). 

[Basionym of Achnantheiopsis fonticolanceolata (Lange-Bert. et Schim.) Lange-Bert.] 

[Basionym of Planothidium fonticolanceolatum (Lange-Bert. et Schim.) Lange-Bert.] 

Achnanthes frequentissima Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 58, pl. 86: figs 1-15', pl. 91: figs 6-7 (1989). 

[Comment: lacking a type and a Latin diagnosis, this name was invalidly published. Validly published as "Achnanthes 

lanceolata ssp. frequentissima" Lange-Bert.] 

Achnanthes helvetica (Hust.) Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 632 (2001). 

Basionym: Achnanthes austriaca var. helvetica Hust., Rabenhorsts Kryptogamenflora 7 (2) 3. Lieferung, 

p. 385, pl. 11: figs 831 g-k (1933). 

[Comment:validation of Achnanthes helvetica (Hust.) Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 63 (1989).] 

Achnanthes hyperboreoides Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 12, pl. 50: figs 13-19, pl. 75: figs 

2, 6, 8 (1996). 

Achnanthes impexa Lange-Bert., Biblioth. Diatomol. 18, p. 67-68, pl. 74: figs 1-6, pl. 75: fig. 1 (1989). [Comment: 

Achnanthes impexa on p. 316 (= caption of pl. 75: fig. 1) is cited as "A. impexiformis".] 

Achnanthes impexiformis Lange-Bert., Biblioth. Diatomol. 18, p. 68-69, pl. 74: figs 7-9, pl. 75: figs 2-4, 1 (1989). 

[Basionym of Nupela impexiformis (Lange-Bert.) Lange-Bert.] 

Achnanthes indicatrix Lange-Bert. et Steindorf, Biblioth. Diatomol. 38, p. 82-83, pl. 18: figs 10-14 (1998). 

[Basionym of Achnanthidium indicatrix (Lange-Bert. et Steindorf) Lange-Bert.] 

Achnanthes ingratiformis Lange-Bert., Biblioth. Diatomol. 18, p. 70-71, pl. 73: fig. 6, pl. 74: figs 13-18´ (1989). 

Achnanthes jan-marcinii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 12-13, pl. 50: figs 5-9, 22-23, pl. 

75: fig. 4, 9 (1996). 

[Basionym of Planothidium jan-marcinii (Witkowski, Metzeltin et Lange-Bert.) Witkowski, Lange-Bert. et Metzeltin.] 

Achnanthes koghisensis f. angustifasciata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 84, pl. 19: figs 

7-11 (1998). 

[Basionym of Achnanthidium koghisense f. angustifasciatum (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert.] 

Achnanthes koghisensis Gerd Moser, Lange-Bert. et Metzeltin f. koghisensis, Biblioth. Diatomol. 38, p. 84, pl. 19: figs 1-6 

(1998). 

[Basionym of Achnanthidium koghisense (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert.] 

Achnanthes kranzii Lange-Bert., Biblioth. Diatomol. 18, p. 74-75, pl. 55: figs 13-21, pl. 62: figs 6-8 (1989). 

Achnanthes kuelbsii Lange-Bert., Biblioth. Diatomol. 18, p. 79-80, pl. 36: figs 27-31, pl. 37: figs 1-4 (1989). 



6 

Achnanthes lanceolata ssp. apiculata (R.M.Patrick) Lange-Bert., Biblioth. Diatomol. 27, p. 3 (1993). 

Basionym: Achnanthes lanceolata var. apiculata R.M.Patrick, Farlowia 2 (2), p. 167, pl. 1: figs 4-5 (1945). 

[Nomenclatural synonym of Planothidium apiculatum (R.M.Patrick) Lange-Bert.] 

Achnanthes lanceolata ssp. dubia (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 3 (1993). 

Basionym: Achnanthes lanceolata var. dubia Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 23 (1880). 

[≡ Achnantheiopsis dubia (Grunow) Lange-Bert.] 

Achnanthes lanceolata ssp. frequentissima Lange-Bert., Biblioth. Diatomol. 27, p. 4 (1993). [Validation of Achnanthes 

lanceolata ssp. frequentissima Lange-Bert., nom. inval., Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), p. 78, pl. 44: 1-3, 15-23, pl. 45: fig. 18 (1991). Comment: in Lange-Bertalot (1993) and in the 

compilation by Kusber & Metzeltin (2001) Achnanthes lanceolata ssp. frequentissima was treated as a new name and status 

for Achnanthes lanceolata var. dubia f. minuta Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Type de 

Synopsis, no. 236 (1880-1887).] 

[Basionym of Planothidium frequentissimum (Lange-Bert.) Lange-Bert.] 

[≡ Achnantheiopsis frequentissima (Lange-Bert.) Lange-Bert.] 

Achnanthes lanceolata var. frequentissima Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 338, 348, pl. 86: figs 1-15', 

pl. 91: figs 6-7 (1989). 

[Comment: lacking a type and a Latin diagnosis, this name was invalidly published. Validly published as "Achnanthes 

lanceolata ssp. frequentissima" Lange-Bert.] 

Achnanthes lanceolata ssp. lanceolatoides (Sovereign) Lange-Bert., Biblioth. Diatomol. 27, p. 6 (1993). 


[≡ Achnantheiopsis lanceolatoides (Sovereign) Lange-Bert.] 

[Nomenclatural synonym of Planothidium lanceolatoides (Sovereign) Lange-Bert.] 

Achnanthes lanceolata ssp. robusta (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 6 (1993). 

Basionym: Achnanthes lanceolata var. robusta Hust., Arch. Hydrobiol. Suppl. 15, p. 202, pl. 13: figs 47-49 (1937). 

[≡ Achnantheiopsis robustior Lange-Bert.] 

[Nomenclatural synonym of Planothidium robustius Lange-Bert.] 

Achnanthes lanceolata ssp. rostrata (Østrup) Lange-Bert., Biblioth. Diatomol. 27, p. 6 (1993). 

Basionym: Achnanthes rostrata Østrup, Flora of Koh Chang, Bot. Tidskr. 26, p. 253 ("1902" 1904). 

[≡ Achnantheiopsis rostrata (Østrup) Lange-Bert.] 

[Nomenclatural synonym of Planothidium rostratum (Østrup) Lange-Bert.] 

Achnanthes lanceolata var. magna (F.Straub) Lange-Bert., Biblioth. Diatomol. 27, p. 5 (1993). 

Basionym: Achnanthes rostrata var. magna F.Straub, Bull. Soc. Neuchâteloise Soc. Nat. 108, p. 139, pl. 10: fig. 142; pl. 11: 

fig. 142 b (1985). 

[Nomenclatural synonym of Planothidium frequentissimum var. magnum (F.Straub) Lange-Bert.] 

Achnanthes lanceolata var. minor (Schulz-Danzig) Lange-Bert., Biblioth. Diatomol. 27, p. 4 (1993). 


[≡ Achnantheiopsis frequentissima var. minor (Schulz-Danzig) Lange-Bert.] 

[Nomenclatural synonym of Planothidium frequentissimum var. minus (Schulz-Danzig) Lange-Bert.] 

Achnanthes lanceolata var. rostratiformis Lange-Bert., Biblioth. Diatomol. 27, p. 5 (1993). 

Replaced synonym: Achnanthes lanceolata var. rostrata Hust., Arch. Naturwiss. Vereine Bremen 20, p. 279, pl. 3: fig. 34 

(1911). 

Achnanthes leonardii Witkowski et Lange-Bert., Iconogr. Diatomol. 7, p. 92, 420, pl. 50: figs 1-12 (2000). 

Achnanthes lineariformis Lange-Bert., Biblioth. Diatomol. 27, p. 7 (1993). [As a part of the protologue is cited: Krammer & 

Lange-Bertalot (1991), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 37: figs 19-23.] [Comment: in 

Krammer & Lange-Bertalot (1991b) Achnanthes lineariformis is documented as "Achnanthes linearis sensu auct. nonnull. 

(excl. Lectotypus)".] 

Achnanthes mercurii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 13, pl. 50: figs 30-32, pl. 75: figs 1, 5, 7 

(1996). 

Achnanthes metakryophila Lange-Bert. et Rol.Schmidt, J. Paleolimnol. 3, p. 64, figs 6 d-i (1990). 

Achnanthes minutissima var. jackii (Rabenh.) Lange-Bert. et Ruppel, Algol. Stud. 26, p. 18 (1980). 

Basionym: Achnanthidium jackii Rabenh., Algen Europas No. 1033 (1861); Flora Europaea Algarum, p. 107 (1864). 

Achnanthes nitidiformis Lange-Bert., Biblioth. Diatomol. 18, p. 109-110, pl. 74: figs 19-24 (1989). 



7 

Achnanthes obscura (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 130 (1996). 

Basionym: Navicula brevicostata var. obscura Krasske, Arch. Hydrobiol. 33, p. 513 (1938). 

Achnanthes parexigua Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 19, pl. 67: figs 1-11, pl. 68: figs 1-3 (1998). 

Achnanthes peridotitica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 85-86, pl. 18: figs 1-9 (1998). 

[Basionym of Achnanthidium peridotiticum (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert.] 

Achnanthes perpusilla (Østrup) Lange-Bert., Iconogr. Diatomol. 2, p. 26 (1996). 

Basionym: Pinnularia perpusilla Østrup, Danske Diatomeer, p. 96, pl. 3: fig. 62 (1910). 

[Nomenclatural synonym of Psammothidium perpusillum (Østrup) Lange-Bert.] 

Achnanthes pirogueana f. capitata Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 40, pl. 2: figs 18-26 (1995). 

Achnanthes placentuloides (Guslakov) Witkowski et Lange-Bert. Iconogr. Diatomol. 7, p. 93 (2000). 

Basionym: Cocconeis placentuloides Guslakov in Guslakov, Zakordoniec et Gerasimuk, Atlas diatomovykh vodorosley 

bentosa severozapadnoy tschasti Tschernogo Morja. Nukovaya Dumka Kiev, p. 59, pl. 80: figs 1-3 (1992). 

Achnanthes praecipua E.Reichardt in Round, Proc. Ninth Int. Diatom Symp., p. 391,figs 1-24 (1988). 

[Basionym of Nupela praecipua (E.Reichardt) E.Reichardt.] 

Achnanthes pusilla var. crassa (Hust.) Lange-Bert. et Ruppel, Algol. Stud. 26, p. 22 (1980). 

Basionym: Achnanthes crassa Hust., Arch. Hydrobiol. Suppl. 15, p. 194, pl. 13, figs 37-40 (1937). 

Achnanthes pusilla var. petersenii (Hust.) Lange-Bert. et Ruppel, Algol. Stud. 26, p. 22 (1980). 

Basionym: Achnanthes petersenii Hust., Bot. Arch. 38, p. 179, figs 10-14 (1936). 

Achnanthes pusilla var. procera (Hust.) Lange-Bert. et Ruppel, Algol. Stud. 26, p. 24 (1980). 

Basionym: Achnanthes procera Hust., Arch. Hydrobiol. 40, p. 908, pl. 39: figs 26-31 (1945). 

Achnanthes renei Lange-Bert. et Rol.Schmidt, J. Paleolimnol. 3, p. 64, figs 6 o-t (1990). 

Achnanthes rosenstockii Lange-Bert., Biblioth. Diatomol. 18, p. 131-132, pl. 61: figs 1-17, pl 63: figs 1-2, 3-5 (1989). 

[Comment: the citation of "pl. 63" missing on p. 131.] 

[Basionym of Psammothidium rosenstockii (Lange-Bert.) Lange-Bert.] 

Achnanthes scotica (J.R.Carter) Lange-Bert., Biblioth. Diatomol. 27, p. 8 (1993). 

Basionym: Achnanthes microcephala f. scotica J.R.Carter, Nova Hedwigia 33, p. 534, pl. 1: fig. 31 (1981). 

Achnanthes silvahercynia Lange-Bert., Biblioth. Diatomol. 18, p. 139, pl. 71: figs 30-37, pl. 73: figs 2-4 (1989). 

[Basionym of Nupela silvahercynia (Lange-Bert.) Lange-Bert.] 

Achnanthes straubiana Lange-Bert., Iconogr. Diatomol. 2, p. 26-27, pl. 78: fig. 21 a-b (1996). [Comment: pl. 78: figs 20, 

21c is not mentioned on p. 26.] [As a part of the protologue, Krammer & Lange-Bertalot (1991), Süßwasserflora von 

Mitteleuropa, Bacillariophyceae 2 (4), pl. 32: figs 53-56 is cited. Comment: in Krammer & Lange-Bertalot (1991b) 

Achnanthes straubiana is cited as "Achnanthes minutissima var. ".] 

[Basionym of Achnanthidium straubianum (Lange-Bert.) Lange-Bert.] 

Achnanthes subatomoides (Hust.) Lange-Bert. et R.E.M. Archibald, Biblioth. Diatomol. 9, p. 9, 97 (1985). 

Basionym: Navicula subatomoides Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 404: figs 33-35 (1936). 

Achnanthes thermalis var. rumrichiorum Lange-Bert., Biblioth. Diatomol. 18, p. 152-153, pl. 77: figs 30-38, pl. 78: figs 2- 

4 (1989). [Comment: Achnanthes thermalis var. rumrichiorum on p. 152, p. 320 and p. 322 is cited as "Achnanthes thermalis 

var. rumrichorum".] 

Achnanthes vistulana Witkowski, Biblioth. Diatomol. 28, p. 90, pl. 17: figs 1-14 (1994). 

Achnanthes ziegleri Lange-Bert., Biblioth. Diatomol. 27, p. 8, pl. 35: figs 4-7 (1993). [Comment: validation of Achnanthes 

ziegleri Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), 

p. 39, pl. 23: figs 33-38 (1991).] 

ACHNANTHIDIUM Kütz. 


Achnanthidium archibaldianum (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 270 (1999). 

Basionym: Achnanthes archibaldiana Lange-Bert., Biblioth. Diatomol. 18, p. 23, pl. 64: figs 7-16, pl. 66: figs 8-11 (1989). 

Achnanthidium arcus (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 270 (1999). 

Basionym: Achnanthes arcus Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 42, p. 38, figs 44-48 (1989). 



8 

Achnanthidium biasolettianum (Grunow) Lange-Bert., Iconogr. Diatomol. 6, p. 270 (1999). 

Basionym: Achnanthes biasolettiana Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 22 (1880). 

[Comment: In Kusber & Metzeltin (2001), p. 621, the basionym erroneously was cited as Achnanthes arcus Grunow.] 

Achnanthidium contrarea (Lange-Bert. et Steindorf) Lange-Bert., Iconogr. Diatomol. 6, p. 271 (1999). 

Basionym: Achnanthes contrarea Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 34, pl. 5: figs 1-19 (1995). 

Achnanthidium eutrophilum (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 271 (1999). 

Basionym: Achnanthes eutrophila Lange-Bert., Iconogr. Diatomol. 2, p. 25, pl. 78: figs 29-38 (1996). 

Achnanthidium indicatrix (Lange-Bert. et Steindorf) Lange-Bert., Iconogr. Diatomol. 6, p. 272 (1999). 

Basionym: Achnanthes indicatrix Lange-Bert. et Steindorf, Biblioth. Diatomol. 38, p. 82, pl. 18: figs 10-14 (1998). 

Achnanthidium koghisense (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert. f. koghisense, Iconogr. Diatomol. 6, p. 273 

(1999). 

Basionym: Achnanthes koghisensis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 84, pl. 19: figs 1-6 

(1998). 

[In Kusber & Metzeltin (2001), p. 622 the basionym authors erroneously are given as "Lange-Bert. et Steindorf".] 

Achnanthidium koghisense f. angustifasciatum (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert., Iconogr. Diatomol. 6, 

p. 273 (1999). 

Basionym: Achnanthes koghisensis f. angustifasciata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 38, 

pl. 19: figs 7-11 (1998). 

[In Kusber & Metzeltin (2001), p. 622 the basionym authors erroneously are given as "Lange-Bert. et Steindorf".] 

Achnanthidium peridotiticum (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert., Iconogr. Diatomol. 6, p. 273 (1999). 

Basionym: Achnanthes peridotitica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 85, pl. 18: figs 1-9 

(1998). 

Achnanthidium straubianum (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 273 (1999). 

Basionym: Achnanthes straubiana Lange-Bert., Iconogr. Diatomol. 2, p. 26, pl. 78: figs 21a, b (1996). 

Achnanthidium subatomus (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 273 (1999). 

Basionym: Achnanthes subatomus Hust., Ber. Deutsch. Bot. Ges. 56, p. 554, pl. 25: figs 1-2 (1939). 

Achnanthidium woltereckii (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes woltereckii Hust., Int. Rev. Gesamten. Hydrobiol. Hydrogr. 42, p. 36, figs 34-37 (1942). 

ACTINELLA F.W.Lewis 


Actinella modesta Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 86-87, pl. 10: figs 5-7 (1998). 

Actinella pararobusta Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 41, pl. 8: figs 3-4, pl. 9: figs 8-10 (1998). 

Actinella pseudohantzschia Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 42, pl. 57: figs 7-10 (1998). 

ADLAFIA Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 87-89 (1998). [Typus generis: Adlafia 

muscora (Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 87 (1998). 

http://www.calacademy.org/research/diatoms/hanna_db/DtmBrwsRecs.aspObj=vGenRef&Atr=GenusID&Val=997]. 

Adlafia aquaeductae (Krasske) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 89 (1998). 

Basionym: Navicula pseudopupula var. aquaeductae Krasske, Bot. Arch. 3, p. 197, fig. 8 (1923). 

[≡ Navicula bryophila var. aquaeductae (Krasske) Lange-Bert.] 

[Comment: in Lange-Bertalot, Diatoms of Europe 2, p. 142 (2001) the authors of Adlafia aquaeductae are cited as 

"(Krasske) Lange-Bertalot in Moser et al.".] 

Adlafia bryophila (J.B.Petersen) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 89 (1998). 

Basionym: Navicula bryophila J.B.Petersen 1928, Dansk. Bot. Ark. 5, p. 388, fig. 13. 

[– Naviculdicta bryophila (J.B. Petersen) Lange-Bert. et Metzeltin, nom. inval., Iconogr. Diatomol. 2, p. 172 (1996).] 

[Comment: in Lange-Bertalot, Diatoms of Europe 2, p. 142 (2001) the authors of Adlafia bryophila are cited as "(Petersen) 

Lange-Bertalot in Moser et al.".] 

Adlafia coringii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 20, pl. 86: figs 12-13 (1998). 

Adlafia drouetiana (R.M.Patrick) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 21 (1998). 

Basionym: Navicula drouetiana R.M.Patrick, Bol. Mus. Nac. Rio de Janeiro 2, p. 4, fig. 4 (1944). 



9 

Adlafia minuscula (Grunow) Lange-Bert. var. minuscula, Iconogr. Diatomol. 6, p. 32 (1999). 

Basionym: Navicula minuscula Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 14: fig. 27 (1880). 

Adlafia minuscula var. muralis (Grunow) Lange-Bert., Iconogr. Diatomol. 6, p. 32 (1999). 

Basionym: Navicula muralis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 14: fig. 27 (1880). 

[≡ Navicula minuscula var. muralis (Grunow) Lange-Bert.] 

[= Navicula cloacina Lange-Bert et Bonik, according to Lange-Bertalot, Diatoms of Europe 2, p. 144 (2001).] 

Adlafia parabryophila (Lange-Bert.) Lange-Bert., Biblioth. Diatomol. 38, p. 89 (1998). 

Basionym: Naviculadicta parabryophila Lange-Bert., Iconogr. Diatomol. 2, p. 86, pl. 27: figs 11-15; pl. 117: figs 5-7 (1996). 

Adlafia suchlandtii (Hust.) Lange-Bert., Biblioth. Diatomol. 38, p. 89 (1998). 

Basionym: Navicula suchlandtii Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 399: figs 24-28 (1934). 

ALINEA Lange-Bert. (section within the genus NAVICULA s. str.), Diatoms of Europe 2, p. 223, (2001). [Typus 

sectionis: Navicula alineae Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of continental Israel, 

p. 268, pl. 22: figs 175-178 (2000).] 

ALTERASYNEDRA Lange-Bert. (subgenus within FRAGILARIA), nom. illeg., Biblioth. Diatomol. 27, p. 52 (1993). 

[Typus subgeneris: Fragilaria ungeriana Grunow, p. 142, pl. 13: fig. 18 (1863). Comment: according to Lange-Bertalot & 

Compère (2001) ALTERASYNEDRA is a taxonomic synonym of FRAGILARIA subgen. ULNARIA Lange-Bert. et Compère, 

Diatom Res. 16, p. 103 (2001) .] 

ALVEOVALLUM Lange-Bert. et Krammer, Diatoms of Europe 1, p. 208, 241, pl. 217: figs 1-15 (2000). [Typus generis: 

Alveovallum beyensii Lange-Bert. et Krammer, Diatoms of Europe 1, p. 208, 241 (2000). Alveovallum was marked as new 

also in Diatoms of Europe 1 (2001 "2000") p. 207, 240-241.] 

Alveovallum beyensii Lange-Bert. et Krammer, Diatoms of Europe 1, p. 208, 241, pl. 217: figs 1-15 (2000). [Alveovallum 

beyensii is the type of the name of the genus ALVEOVALLUM Lange-Bert. & Krammer.] 

AMICULA Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 127, 420 (2000). [Typus generis: Amicula 

specululum (Witkowski) Witkowski, Iconogr. Diatomol. 7, p. 127 (2000).] 

Amicula specululum (Witkowski) Witkowski, Iconogr. Diatomol. 7, p. 127 (2000). 

Basionym: Navicula specululum Witkowski, Biblioth. Diatomol. 28, p. 159, pl. 36: figs 5-12 (1994). 

AMPHIPLEURA Kütz. 


Amphipleura chiapasensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 21, pl. 109: figs 1-5 (1998). 

AMPHORA Ehrenb. 


Amphora aequalis Krammer, Bacillaria 3, p. 212-213, figs 25-35 (1980). 

Amphora dissimilis Metzeltin et Krammer, Biblioth. Diatomol. 38, p. 90, pl. 43: figs 1-8 (1998). 

Amphora fogediana Krammer, Biblioth. Diatomol. 9, p. 9, pl. 10: figs 1-3 (1985). [As a part of the protologue, Krammer & 

Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 150: 14-17 is cited.] 

Amphora hassatica Krammer et S.Strecker, Biblioth. Diatomol. 37, p. 225-226, pl. 206: figs 1-8 (1997). 

Amphora inariensis Krammer, Bacillaria 3, p. 211, figs 21-24, 36-37, 43-45 (1980). 

Amphora jostesorum Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 141, 420, pl. 171: figs 1-9 (2000). 

[Comment: on p. 141 and 784 (caption of pl. 171) the authors are cited as Witkowski, Metzeltin et Lange-Bert.] 

Amphora oligotraphenta Lange-Bert., Iconogr. Diatomol. 2, p. 28 (1996). 

Replaced synonym: Amphora veneta var. capitata Haworth, Brit. Phycol. J. 9, p. 48, fig. 6 (1974). 

Amphora streckerae Krammer, Biblioth. Diatomol. 37, p. 226-227, pl. 206: figs 9-11 (1997). 

Amphora subholsatica Krammer, Biblioth. Diatomol. 37, p. 223-225, pl. 205: figs 1-10 (1997). 

[– Amphora hosatica sensu Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), 

pl. 152: figs 4-5.] 

Amphora tomiakae Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 7, p. 153, 421, pl. 169: figs 12-14 (2000). 



10 

ANAULUS Ehrenb. 


Anaulus simonsenii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 7, p. 23, 421, pl.10: figs 7-12 (2000). 

[Comment: on p. 23 and 462 (caption of pl. 10) the authors are cited as Witkowski et Metzeltin.] 

ANEUMASTUS D.G.Mann et A.J.Stickle 

[http://www.calacademy.org/research/diatoms/hanna_db/DtmBrwsRecs.aspObj=vGenRef&Atr=GenusID&Val=357] 

[http://128.32.109.44/cgi-bin/porp_cgi.plANEUMASTUSDGMANNAJSTICKLE1990]. 

Aneumastus albanicus Lange-Bert. et Miho, Diatoms of Europe 2, p. 216, pl. 123: figs 1-10 (2001). 

Aneumastus apiculatus (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 32 (1999). 

Basionym: Navicula lacustris var. apiculata Østrup, Danske Diatomeer, p. 88, pl. 3: fig. 59 (1910). 

Aneumastus aratus (Grunow) Lange-Bert., Diatoms of Europe 2, p. 153, (2001). 

Basionym: Navicula arata Grunow in Mojsisovics & Neumayr, Beitr. Kenntnis der fossilen Diatomeen Österreich-Ungarns 

und des Orients. 2 (4), p. 145, pl. 30: fig. 58 (1882). 

Aneumastus balticus Lange-Bert., Diatoms of Europe 2, p. 216, pl. 117: figs 11-15, pl. 118: figs 7-8 (2001). 

Aneumastus carolinianus (R.M.Patrick) Lange-Bert., Diatoms of Europe 2, p. 154, (2001). 

Basionym: Navicula caroliniana R.M.Patrick, Proc. Acad. Nat. Sci. Philadelphia 111, p. 107, pl. 7: fig. 11 (1959). 

Aneumastus humboldtianus Lange-Bert. et Miho, Diatoms of Europe 2, p. 217, pl. 120: figs 3-7 (2001). 

Aneumastus laetus (Ant.Mayer) Lange-Bert., Iconogr. Diatomol. 2, p. 29 (1996). 

Basionym: Navicula laeta Ant.Mayer, Kryptog. Forsch. 1, p. 204, pl. 7: fig. 59 (1919). 

Aneumastus maculosus (Donkin) Lange-Bert., Diatoms of Europe 2, p. 155, (2001). 

Basionym: Navicula maculosa Donkin, Natural Hist. Brit. Diatomaceae 2, p. 25, pl. 5: fig. 1 (1871). 

Aneumastus minor (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 9 (1993). 

Basionym: Navicula tuscula f. minor Hust., Süsswasserflora Mitteleuropas 10, p. 308, fig. 553 (1930). 

Aneumastus rosettae Lange-Bert. et Miho, Diatoms of Europe 2, p. 217, pl. 115: figs 1-6, pl. 116: figs 1-5 (2001). 

Aneumastus rostratus (Hust.) Lange-Bert., Diatoms of Europe 2, p. 156, (2001). 

Basionym: Navicula tuscula var. rostrata Hust., Abh. Naturwiss. Vereine Bremen 20, p. 287, pl. 3: fig. 22 (1911). 

Aneumastus tusculus f. angulatus (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 33 (1999). 

Basionym: Navicula tuscula f. angulata Hust., Arch. Hydrobiol. 39, p. 120, fig. 37 (1942). 

ANOMOEONEIS Pfitzer 


Anomoeoneis garrensis Lange-Bert. et Krammer, Biblioth. Diatomol. 9, p. 12, pl. 1: figs 1-5 (1985). [As a part of the 

protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 103a: 10-13 is 

cited.] 

Anomoeoneis starmuehlneri R.Maillard in Gerd Moser, Lange-Bertalot et Metzeltin, Biblioth. Diatomol. 38, p. 100 (1998). 

[Comment: The Holotypus is mentioned as "Neotypus".] 

[– Anomoeoneis starmuehlneri R.Maillard, nom. inval., Cah. O.R.S.T.O.M. Sér. Hydrobiol. 12 (2), p. 154, pl. 2: fig. 7 

(1978).] 

[Nomenclatural synonym of Brachysira starmuehlneri (R.Maillard) Gerd Moser, Lange-Bert. et Metzeltin.] 

ASTARTIELLA Witkowski, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 91, 357-358 (1998). [Typus generis: 

Astartiella bahusiensis (Grunow) Witkowski, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 357 (1998).] 

Astartiella bahusiensis (Grunow) Witkowski, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 359 (1998). 

Basionym: Navicula minuscula var. bahusiensis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 14: 

fig. 2 (1880). 

Astartiella bahusiensoides (Foged) Witkowski, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 359 (1998). 

Basionym: Navicula bahusiensoides Foged 1975, Biblioth. Pycol. 16, p. 36, pl. 19: figs 8-9. 

Astartiella bremeyeri (Lange-Bert.) Witkowski et Lange-Bert. var. bremeyeri, Biblioth. Diatomol. 38, p. 359 (1998). 

Basionym: Achnanthes bremeyeri Lange-Bert., Biblioth. Diatomol. 18, p. 31, pl. 76: figs 11-12 (1989). 



11 

Astartiella bremeyeri var. rostrata (Hust.) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 359 (1998). 

Basionym: Navicula bremeyeri var. rostrata Hust., Abh. Naturwiss. Vereine Bremen 31, p. 624, figs 73-74 (1939). 

Astartiella glacialis Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 100, 421-422, pl. 52: figs 5-10 (2000). 

Astartiella ignota (Hust.) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 359 (1998). 

Basionym: Achnanthes ignota Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 420: figs 20-23 (1958). 

Astartiella iks Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 100, 422, pl. 52: figs 35-39 (2000). 

Astartiella modica (Hust.) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 360 (1998). 

Basionym: Achnanthes modica Hust., Bot. Not. 4, p. 388, figs 72-75 (1952). 

Astartiella producta Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 101, 422, pl. 52: figs 1-4 (2000). 

Astartiella punctifera (Hust.) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 360 (1998). 

Basionym: Achnanthes punctifera Hust., Bull. Mar. Sta. Duke Univ. 6, p. 18, pl. 5: figs 26-28 (1955). 

Astartiella tenuis (Hust.) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 360 (1998). 

Basionym: Achnanthes tenuis Hust., Bull. Mar. Sta. Duke Univ. 6, p. 18, pl. 5: figs 29-31 (1955). 

Astartiella wellsiae (Reimer) Witkowski et Lange-Bert., Biblioth. Diatomol. 38, p. 360 (1998). 

Basionym: Achnanthes wellsiae Reimer in Patrick & Reimer, Monogr. Acad. Nat. Sci. Philadelphia 1, p. 255, pl. 16: figs 15- 

17 (1966). 

AUSTARIELLA Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 154, 422 (2000). [Typus generis: 

Austariella admissa (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 154 (2000).] 

Austariella admissa (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 154 (2000). 

Basionym: Navicula admissa Hust., Bull. Duke Univ. Mar. Sta. 6, p. 30, pl. 8: fig. 25 (1955). 

Austariella jamalinensis (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 155 (2000). 

Basionym: Navicula jamalinensis Cleve, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 13, pl. 2: fig. 40 (1880). 

Austariella jamalinensis var. subcircularis (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 155 

(2000). 

Basionym: Navicula jamalinensis var. subcircularis Østrup, Danske Diatomeer, p. 206, pl. 13: fig. 2 (1910). 

[Comment: In Iconogr. Diatomol. 7 (2000), p. 155 the basionym name is misspelled as Navicula jamalinensis var. 

"suborcircularis".] 

BACILLARIA J.F. Gmelin 


Bacillaria paxillifer var. tumidula (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 357 (2000). 

Basionym: Nitzschia paradoxa var. tumidula Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 86 (1880). 

BIREMIS D.G.Mann et E.J.Cox 


Biremis circumtexta (F.Meister ex Hust.) Witkowski et Lange-Bert., Iconogr. Diatomol. 7, p. 158 (2000). 

Basionym: Navicula circumtexta F.Meister ex Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 394: figs 33-35 

(1934). 

Biremis kriegeriana (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 166-167 (1996). 

Basionym: Pinnularia kriegeriana Krasske, Ber. Deutsch. Bot. Ges. 61, p. 86, fig. 17 (1943). 

[Nomenclatural synonym of Pulchella kriegeriana (Krasske) Krammer.] 

Biremis solitaria (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 159 (2000). 

Basionym: Navicula solitaria Cleve, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 22, p. 20, pl. 1: fig. 6 (1896). 

BRACHYSIRA Kütz. 


Brachysira acutirhomboides Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 10, pl. 25: figs 1-4 (1994). 

Brachysira acutissima Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 91-92, pl. 39: figs 20-23 (1998). 

Brachysira amoena Lange-Bert., Biblioth. Diatomol. 29, p. 11, pl. 37: figs 10-14 (1994). 

Brachysira archibaldii var. crassistriata Lange-Bert., Biblioth. Diatomol. 29, p. 16, pl. 32: figs 15-20 (1994). 



12 

Brachysira australofollis Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 17, pl. 20: fig. 4-15, pl. 21: fig. 4 (1994). 

Brachysira australofollis var. angustior Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 92-93, pl. 36: figs 

1-4 (1998). 

Brachysira babuschka Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 93-94, pl. 39: figs 1-8 (1998). 

Brachysira bouletiana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 94-95, pl. 35: figs 4-6 (1998). 

Brachysira brebissonii ssp. superba Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 22, pl. 24: figs 5-7 (1994). 

Brachysira calcicola Lange-Bert., Biblioth. Diatomol. 29, p. 23, pl. 41: figs 19-33 (1994). 

Brachysira calligraphica Lange-Bert., Biblioth. Diatomol. 29, p. 24, pl. 1: figs 1-2, pl. 37: figs 1-9 (1994). [Comment: the 

diagnosis was emended by Metzeltin et Lange-Bert. in Metzeltin & Lange-Bertalot, Iconogr. Diatomol. 5, p. 22-23 (1998) by 

excluding pl. 37: figs 6-9. According to Metzeltin & Lange-Bertalot (1998) these figures show B. steinitziae.] 

Brachysira coraliana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 23, pl. 103: figs 9-14 (1998). 

Brachysira estoniarum Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 160, 423-424, pl. 134: figs 1-4 

(2000). 

Brachysira guttiformis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 95-96, pl. 36: figs 5-7 (1998). 

Brachysira hofmanniae Lange-Bert., Biblioth. Diatomol. 29, p. 31, pl. 8: figs 1-18, pl. 9: figs 1-3 (1994). 

Brachysira inamoena Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 24, pl. 104: figs 6-9 (1998). 

Brachysira incognita Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 96-97, pl. 39: figs 16-19 (1998). 

[Comment:validation of Brachysira incognita Gerd Moser et Lange-Bert., nom. inval., Biblioth. Diatomol. 29, p. 32, pl. 25: 

fig. 9 (1994).] 

Brachysira inflata Lange-Bert., Biblioth. Diatomol. 29, p. 32-33, pl. 25: figs 5-8 (1994). 

Brachysira insolita Lange-Bert. et Gerd Moser, nom. inval., Biblioth. Diatomol. 29, p. 33-34, pl. 29: figs 19-20 (1994). 

[Comment: lacking a type and a Latin diagnosis this name is invalidly published.] 

Brachysira insulsa Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 24-25, pl. 107: figs 1-4 (1998). 

Brachysira irawanoides Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 36, pl. 30: figs 12-16 (1994). 

Brachysira julio Lange-Bert., Biblioth. Diatomol. 29, p. 37, pl. 40: figs 1-9 (1994). 

Brachysira krammeri Lange-Bert., Biblioth. Diatomol. 29, p. 37-38, pl. 16: figs 8-17, pl. 22: figs 4-6 (1994). 

Brachysira kuntzei (Reichelt) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 25 (1998). 

Basionym: Navicula kuntzei Reichelt in Kuntze, Revisio Generum plantarum, 3 (1), p. 391, fig. 3a (1898). 

Brachysira lange-bertalotii Metzeltin, Iconogr. Diatomol. 5, p. 25-26, pl. 103: figs 5-8, pl. 106: figs 6-7 (1998). 

Brachysira lecohui Lange-Bert., Biblioth. Diatomol. 29, p. 39, pl. 15: figs 8-13, pl. 22: figs 2-3, pl. 40: fig. 14 [valve on the 

right hand side] (1994). 

Brachysira lehmanniae Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 40, pl. 39: figs 13-17 (1994). 

Brachysira liliana Lange-Bert., Biblioth. Diatomol. 29, p. 41, pl. 10: figs 1-12 (1994). 

Brachysira macroserians Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 26, pl. 103: figs 1-4 (1998). 

Brachysira maillardii Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 43, pl. 29: figs 11-17 (1994). 

Brachysira manfredii Lange-Bert.. Biblioth. Diatomol. 29, p. 44, pl. 32: figs 6-14, pl. 48: figs 8-9 (1994). 

Brachysira metzeltinii Lange-Bert., Biblioth. Diatomol. 29, p. 45, pl. 17: figs 1-6, pl. 18: figs 5-6, pl. 40: fig. 14 [valve on 

the left hand side] (1994). 

Brachysira microclava Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 46, pl. 30: figs 1-9 (1994). 

Brachysira microserians Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 27, pl. 102: figs 1-5, 10-11 (1998). 

Brachysira nanoclava Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 97-98, pl. 37: figs 16-22 (1998). 

Brachysira neoacuta Lange-Bert., Biblioth. Diatomol. 29, p. 48, pl. 14: figs 9-10, pl. 15: figs 1-7, pl. 18: figs 3-4 (1994). 



13 

Brachysira neoexilis Lange-Bert., Biblioth. Diatomol. 29, p. 51-52, pl. 5: figs 1-35, pl. 6: figs 1-6, pl. 17: figs 7-11, pl. 32: 

figs 27-30, pl. 46: figs 19-27 (1994). [Comment: Brachysira neoexilis on p. 194 (= caption of pl. 46: figs 19-27) is cited as 

"Brachysira neoexilis Lange-Bert. or cf. neoexilis".] 

[Taxonomic (i.e. heterotypic) synonym of Brachysira microcephala (Grunow) Compère, according to Wolfe & Kling, 

Lange-Bertalot-Festschrift, p. 250 (2001).] 

Brachysira nubigena Lange-Bert., Biblioth. Diatomol. 29, p. 54, pl. 38: figs 1-4, pl. 39: figs 7-12 (1994). 

Brachysira procera Lange-Bert. et Gerd Moser, p. 55-56, pl. 7: figs 8-26, pl. 9: figs 4-6, pl. 32: figs 21-26 (1994). 

[Comment: Brachysira procera on p. 166 (= caption of pl. 32: figs 21-26) is cited as "Brachysira species cf. procera".] 

Brachysira pseudoacuta Lange-Bert., Biblioth. Diatomol. 29, p. 57, pl. 16: figs 1-7, pl. 18: figs 1-2 (1994). 

Brachysira pseudoexilis Lange-Bert. et Gerd Moser, nom. inval., Biblioth. Diatomol. 29, p. 58, pl. 17: figs 12-26, pl. 19: 

figs 1-6 (1994). [Comment: lacking a type and a Latin diagnosis this name is invalidly published.] 

Brachysira pulchra Lange-Bert., Biblioth. Diatomol. 29, p. 58-59, pl. 39: figs 1-6 (1994). 

Brachysira pumila Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 28, pl. 102: figs 13-17 (1998). 

Brachysira rostrata (Krasske) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 28 (1998). 

Basionym: Anomoeoneis serians var. rostrata Krasske, Svensk Bot. Tidskr. 42, p. 427, pl. 1: figs 28-29 (1948). 

Brachysira rumrichiae Lange-Bert., Biblioth. Diatomol. 29, p. 59-60, pl. 26: figs 1-7 (1994). 

Brachysira ruppeliana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 98-99, pl. 38: figs 1-3, pl. 39: figs 

12-15 (1998). 

Brachysira seippii Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 60. pl. 29: figs 6-10 (1994). 

Brachysira silvicola Lange-Bert., Biblioth. Diatomol. 29, p. 63, pl. 27: figs 16-23, pl. 28: figs 1-5 (1994). 

Brachysira simplex Lange-Bert., Biblioth. Diatomol. 29, p. 64, pl. 40: figs 10-13 (1994). 

Brachysira speluncola Lange-Bert., Biblioth. Diatomol. 29, p. 66, pl. 21: figs 5-6, pl. 22: fig. 1, pl. 23: figs 8-15 (1994). 

Brachysira spicula Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 99-100, pl. 37: figs 10-15 (1998). 

Brachysira starmuehlneri (R.Maillard) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 100 (1998). 

Basionym: Anomoeoneis starmuehlneri R.Maillard in Gerd Moser, Lange-Bertalot et Metzeltin, Bibioth. Diatomol. 38, p. 

100. 

[– Brachysira starmuehlneri (R. Maillard) Gerd Moser, Steindorf et Lange-Bert., nom. inval., Biblioth. Diatomol. 32, p. 55 

(1995).] 

Brachysira steindorfiana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 101, pl. 37: figs 1-4 (1998). 

Brachysira steinitziae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 29, pl. 102: figs 18-22, pl. 108: fig. 7 (1998). 

[Comment: see Brachysira calligraphica.] 

Brachysira sublinearis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 29, pl. 107: fig. 15, pl. 108: figs 5-6 (1998). 

Brachysira subrostrata Lange-Bert., Biblioth. Diatomol. 29, p. 68, pl. 35: figs 7-11, pl. 36: fig. 9 (1994). 

Brachysira supriniana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 103-104, pl. 39: figs 9-11, pl. 40: 

figs 1-5 (1998). 

Brachysira tenuiserians Lange-Bert., Biblioth. Diatomol. 29, p. 69, pl. 14: figs 5-8 (1994). 

Brachysira trinodis Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 29, p. 70, pl. 24: figs 1-4 (1994). 

Brachysira wygaschii Lange-Bert., Biblioth. Diatomol. 29, p. 72, pl. 2: figs 3-6, pl. 13: figs 1-11 (1994). 

BREVISIRA Krammer, Jahn et al., Lange-Bert.-Festschrift, p. 19 (2001) [Typus generis: Brevisira arentii (Kolbe) 

Krammer in Jahn et al., Lange-Bertalot-Festschrift, p. 19 (2001).] 

Brevisira arentii (Kolbe) Krammer in Jahn et al., Lange-Bertalot-Festschrift, p. 19 (2001) 

Basionym: Cyclotella arentii Kolbe, Svensk Bot. Tidskr. 42, p. 464, figs 9-10 (1948). [Brevisira arentii is the type of the 

name of the genus BREVISIRA.] 

CALONEIS Cleve 




14 

Caloneis arctica (Krasske) Lange-Bert., Iconogr. Diatomol. 6, p. 34 (1999). 

Basionym: Caloneis silicula [var. alpina] f. arctica Krasske, Arch. Hydrobiol. 33, p. 526 (1938). 

Caloneis constans E.Reichardt, Ber. Bayer. Bot. Ges. 64, p. 124, pl. 2: figs 1-7, pl. 4: fig. 6 (1994). 

Caloneis fontinalis (Grunow) Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 29 (1996). 

Basionym: Navicula fontinalis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, p. 103, pl. 12: fig. 33 (1880- 

1885). 

Caloneis hendeyi Lange-Bert., Iconogr. Diatomol. 6, p. 34 (1999). 

Replaced synonym: Stauroneis amphioxys var. obtusa Hendey, Fishery Investigations Ser. IV, Part V, Bacillariophyceae, p. 

220 (partim), pl. 43: fig. 9 (7, 8) (1964). 

Caloneis intermedia (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 26 

(1998). 

Basionym: Caloneis clevei var. intermedia Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 207 (1996). 

Caloneis kristinae Gerd Moser, Biblioth. Diatomol. 38, p. 105, pl. 54: figs 1-6 (1998). 

Caloneis lancettula (Schulz-Danzig) Lange-Bert. et Witkowski, Iconogr. Diatomol. 2, p. 29 (1996). 

Basionym: Caloneis aemula var. lancettula Schulz-Danzig, Bot. Arch. 13, p. 202, fig. 74e (1926). 

Caloneis odiosa (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 26 

(1998). 

Basionym: Pinnularia odiosa Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 206 (1996). 

Caloneis subclevei Krammer, Biblioth. Diatomol. 9, p. 21, pl. 15: figs 5-11 (1985). 

Caloneis undulatiformis Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 9, p. 100 (2000). 

Replaced synonym: Caloneis clevei var. undulata Krasske, Arch. Hydrobiol. 35, p. 379, pl. 11: figs 15-17 (1939). 

CAMPYLODISCUS Ehrenb. ex Kütz. 


Campylodiscus indianorum Lange-Bert. et Metzeltin, Nova Hedwigia Beih. 112, p. 325, fig. 13-20 (1996). 

CAPARTOGRAMMA Kuff. 


Capartogramma pumila Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 30-31, pl. 98: figs 6-9 (1998). 

CATENULA Mereschk. 


Catenula robusta Witkowski, Lange-Bert. et Metzeltin Iconogr. Diatomol. 7, p. 168, 424, pl. 170: figs 13-14 (2000). 

CAVINULA D.G. Mann & A.J. Stickle 


Cavinula cocconeiformis f. elliptica (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 30 (1996). 

Basionym: Navicula cocconeiformis f. elliptica Hust., Arch. Hydrobiol. 48, p. 470 (1954). 

Cavinula intractata (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 30 (1996). 

Basionym: Navicula intractata Hust., Rabenhorsts Kryptogamenflora 7 (3) 1. Lieferung, p. 136, fig. 1269 (1961). 

Cavinula lapidosa (Krasske) Lange-Bert., Iconogr. Diatomol. 2, p. 30 (1996). 

Basionym: Navicula lapidosa Krasske, Bot. Arch. 27, p. 354, fig. 7 (1929). 

Cavinula mollicula (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 31 (1996). 

Basionym: Navicula mollicula Hust., Arch. Hydrobiol. 40, p. 917, pl. 41: figs 34-36 (1945). 

Cavinula scutelloides (W.Sm.) Lange-Bert., Iconogr. Diatomol. 2, p. 31 (1996). 

Basionym: Navicula scutelloides W.Sm., Quart. J. Microscop. Sci. 4, p. 4, pl. 1: fig. 15 (1856). 

Cavinula thoroddsenii (Foged) Lange-Bert., Iconogr. Diatomol. 2, p. 31 (1996). 

Basionym: Navicula thoroddsenii Foged, Biblioth. Phycol. 15, p. 83, pl. 16: figs 7-10 (1974). 

CHAMAEPINNULARIA Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 32 (1996). [Typus generis: Chamaepinnularia 

vyvermanii Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 37 (1996).] 



15 

Chamaepinnularia alexandrowiczii Witkowski, Lange-Bert. et Metzeltin Iconogr. Diatomol. 7, p. 168, 423, pl. 69: figs 1-6 

(2000). 

[Comment: In Metzeltin & Kusber (2001), p. 579, the genus name was misspelled as "Chameapinnularia".] 

Chamaepinnularia begeri (Krasske) Lange-Bert., Iconogr. Diatomol. 2, p. 33 (1996). 

Basionym: Navicula begeri Krasske, Hedwigia 72, p. 113, pl. 3: fig. 16 (1932). 

Chamaepinnularia brasiliana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 31-32, pl. 86: figs 25-27 (1998). 

Chamaepinnularia brasilianopsis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 32, pl. 86: figs 21-24 (1998). 

Chamaepinnularia bremensis (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula bremensis Hust., Abh. Naturwiss. Vereine Bremen 34, p. 284, figs 34-35 (1957). 

Chamaepinnularia calida (Hendey) Lange-Bert., Iconogr. Diatomol. 6, p. 35 (1999). 

Basionym: Navicula calida Hendey, Fishery Investigations Ser. IV, Part V, Bacillariophyceae, p. 198, pl. 43: figs 10-11 

(1964). 

Chamaepinnularia circumborealis Lange-Bert., Iconogr. Diatomol. 6, p. 35, pl. 45: figs 15-18, pl. 48: figs 1-5, pl. 54: fig. 9 

(1999). 

Chamaepinnularia clamans (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 169 (2000). 

Basionym: Navicula clamans Hust., Abh. Naturwiss. Vereine Bremen 31, p. 624, figs 75-77 (1939). 

Chamaepinnularia distantepunctata (Simonsen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 169 (2000). 

Basionym: Navicula distantepunctata Simonsen 1974, The Diatom Plankton of the Indian Ocean. "Meteor" 

Forschungsergebnisse, D 19, p. 41, pl. 26: fig. 4. 

Chamaepinnularia evanida (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula evanida Hust., Ber. Deutsch. Bot. Ges. 60, p. 66, figs 20-21 (1942). 

Chamaepinnularia ferrarioana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 33, pl. 86: figs 28-31 (1998). 

Chamaepinnularia gandrupii (J.B.Petersen) Lange-Bert. et Krammer var. gandrupii, Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Pinnularia gandrupii J.B.Petersen, Kongel. Danske Vidensk.-Selsk. Skr. Afd. 12, Raekke 7, p. 16, fig. 4 (1924). 

[≡ Navicula gandrupii (J.B.Petersen) Krasske, Arch. Hydrobiol. 31, p. 528 (1938).] 

Chamaepinnularia gandrupii var. simplex (Krasske) Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula gandrupii var. simplex Krasske, Arch. Hydrobiol. 31, p. 528, pl. 11: figs 9-10 (1938). 

Chamaepinnularia justa (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 170 (2000). 

Basionym: Navicula justa Hust., Bull. Duke Univ. Mar. Sta. 6, p. 31, pl. 7: fig. 30 (1955). 

Chamaepinnularia krasskei Lange-Bert., Iconogr. Diatomol. 6, p. 36, pl. 46: figs 1-10 (1999). 

Chamaepinnularia krookiformis (Krammer) Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 37 (1999). 

Basionym: Pinnularia krookiformis Krammer, Biblioth. Diatomol. 26, p. 79, pl. 18: figs 14-21 (1992). 

Chamaepinnularia krookii (Grunow) Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 37 (1999). 

Basionym: Navicula krookii Grunow in E. Mojsisovics et M. Neumayer, Beiträge z. Paläontologie Österreich-Ungarns u. des 

Orients 2 (4), p. 155, pl. 30: fig. 40 (1882). 

Chamaepinnularia margaritacea (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula margaritacea Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 402: figs 61-62 (1936). 

Chamaepinnularia margaritiana (Witkowski) Witkowski, Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula margaritiana Witkowski, Biblioth. Diatomol. 28, p. 151, pl. 37: figs 1-9 (1994). 

Chamaepinnularia mediocriformis (Coste et Ricard) Lange-Bert., Iconogr. Diatomol. 2, p. 34 (1996). 

Basionym: Navicula mediocriformis Coste et Ricard, Cryptog. Algol. 3, p. 291, pl. 3: figs 21-22, pl. 6: fig. 14 (1982). 

Chamaepinnularia mediocris (Krasske) Lange-Bert. var. mediocris, Iconogr. Diatomol. 2, p. 35 (1996). 

Basionym: Navicula mediocris Krasske, Hedwigia 72, p. 113, pl. 3: fig. 15 (1932). 

Chamaepinnularia mediocris var. atomus (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 35 (1996). 

Basionym: Pinnularia atomus Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 390: figs 20-23 (1934). 

Chamaepinnularia reinventa Lange-Bert. et Wydrzycka ssp. reinventa, Brenesia 55-56, p. 5-6, pl. 4: figs 6-7 (2001). [As a 

part of the protologue is cited: Lange-Bertalot & Metzeltin Iconogr. Diatomol. 2, pl. 28: figs 60-63, pl. 118: figs 6-7 (1996) 



16 

in the Latin diagnosis. Comment: in Lange-Bertalot & Metzeltin (1996) Chamaepinnularia reinventa ssp. reinventa is cited 

as "Chamaepinnularia spec. Nr. 3 Julma Ölkky aff. begeri".] 

Chamaepinnularia reinventa ssp. urtroquarea Wydrzycka et Lange-Bert., Brenesia 55-56, p. 6, pl. 4: figs 8-11 (2001). 

Chamaepinnularia resalutata Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 6-7, pl. 4: figs 1-5 (2001). [As a part of the 

protologue is cited: Lange-Bertalot & Metzeltin (1996) Iconogr. Diatomol. 2, "fig. 37-39c (nec. d et e)" i.e. pl. 28: figs 37-38, 

39a-c. Comment: in Lange-Bertalot & Metzeltin (1996) Chamaepinnularia resalutata is cited as "Chamaepinnularia (nov.) 

spec. Nr. 2 Julma Ölkky".] 

Chamaepinnularia rhombelliptica Lange-Bert., Iconogr. Diatomol. 2, p. 35, pl. 28: figs 27-30 (1996). 

Chamaepinnularia rupicola (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 35 (1996). 

Basionym: Navicula rupicola Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 401: figs 103-105 (1936). 

Chamaepinnularia schauppiana Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 36, pl. 28: figs 43-47c (1996). 

Chamaepinnularia soehrensis (Krasske) Lange-Bert. et Krammer var. soehrensis, Iconogr. Diatomol. 2, p. 36 (1996). 

Basionym: Navicula soehrensis Krasske, Bot. Arch. 3, p. 198, fig. 2 (1923). 

Chamaepinnularia soehrensis var. capitata (Krasske) Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 37 (1996). 

Basionym: Navicula soehrensis var. capitata Krasske, Abh. Ber. Vereins Naturk. Kassel, 84-89, p. 47, pl. 2: fig. 37 (1925). 

Chamaepinnularia soehrensis var. hassica (Krasske) Lange-Bert., Iconogr. Diatomol. 2, p. 37 (1996). 

Basionym: Navicula hassica Krasske, Abh. Ber. Vereins Naturk. Kassel, 84-89, p. 47, pl. 2: fig. 26 (1925). 

[≡ Navicula soehrensis var. hassiaca (Krasske) Lange-Bert.] 

Chamaepinnularia soehrensis var. muscicola (J.B.Petersen) Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 37 (1996). 

Basionym: Pinnularia muscicola J.B.Petersen, p. 407, Botany of Iceland 2, fig. 27 (1928). 

[≡ Navicula soehrensis var. muscicola (J.B.Petersen) Krasske, Bot. Arch. 27, p. 373 (1929).] 

Chamaepinnularia submusciola (Krasske) Lange-Bert., Biblioth. Diatomol. 38, p. 27 (1998). 

Basionym: Navicula submusciola Krasske, Arch. Hydrobiol. 35, p. 385, pl. 11: fig. 32 (1939). 

Chamaepinnularia tongatensis (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 37 (1996). 

Basionym: Navicula tongatensis Hust., Rabenhorsts Kryptogamenflora 7 (3) 2. Lieferung, p. 227, fig. 1345 (1962). 

Chamaepinnularia truncata (Dieter König) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 171 (2000). 

Basionym: Caloneis truncata Dieter König, Z. Deutsch. Geol. Ges. 111, p. 54, pl. 2: figs 11a-c, pl. 3: figs 9a-d (1959). 

Chamaepinnularia vyvermanii Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 37 (1996). [As a part of the protologue is 

cited: Krammer & Lange-Bertalot (1985) Biblioth. Diatomol. 9, pl. 26: figs 1-6, 22-24 is cited. Comment: in Krammer & 

Lange-Bertalot (1985) Chamaepinnularia vyvermanii is cited as "Navicula spec. aus Irland und Finnland". In Iconogr. 

Diatomol. 2 on p. 37 "Krammer" is missing.] [Chamaepinnularia vyvermanii is the type of the name of the genus 

CHAMAEPINNULARIA Lange-Bert. et Krammer.] 

Chamaepinnularia wiktoriae (Witkowski et Lange-Bert.) Witkowski et Lange-Bert., Iconogr. Diatomol. 2, p. 38 (1996). 

Basionym: Navicula wiktoriae Witkowski et Lange-Bert., Biblioth. Diatomol. 28, p. 162, pl. 34: figs 1-13 (1994). 

[Comment: This new combination was again published as: Chamaepinnularia wiktoriae (Witkowski et Lange-Bert.) 

Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 171 (2000).] 

COCCONEIOPSIS Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 172, 424 (2000). [Typus generis: 

Cocconeiopsis orthoneoides (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 174 (2000).] 

Cocconeiopsis breviata (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 172 (2000). 

Basionym: Navicula breviata Hust., Rabenhorsts Kryptogamenflora 7 (3) 4. Lieferung, p. 637, fig. 1635 (1966). 

Cocconeiopsis fraudulenta (A.W.F.Schmidt) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 173 (2000). 

Basionym: Navicula fraudulenta A.W.F.Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 70: fig. 60 (1881). 

Cocconeiopsis girondica (Dieter König) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 173 (2000). 

Basionym: Navicula girondica Dieter König, Z. Deutsch. Geol. Ges. 111, p. 51, pl. 2: fig. 5, pl. 3: fig. 7 (1959). 

Cocconeiopsis kantsinensis (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 173 (2000). 

Basionym: Navicula kantsinensis Giffen, Nova Hedwigia 13, p. 269, figs 65-67 (1967). 

Cocconeiopsis orthoneoides (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 174 (2000). 

Basionym: Navicula orthoneoides Hust., Duke Univ. Mar. Sta. Bull. 6, p. 31, pl. 7: fig. 14 (1955). 



17 

Cocconeiopsis patrickae (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 174 (2000). 

Basionym: Navicula patrickae Hust., Duke Univ. Mar. Sta. Bull. 6, p. 26, pl. 8: figs 15-16 (1955). 

Cocconeiopsis pseudoorthoneoides (Foged) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 175 (2000). 

Basionym: Navicula pseudoorthoneoides Foged, Biblioth. Phycol. 16, p. 42, pl. 20: fig. 12 (1975). 

Cocconeiopsis pullus (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 175 (2000). 

Basionym: Navicula pullus Hust., Duke Univ. Mar. Sta. Bull. 6, p. 20, pl. 7: fig. 18 (1955). 

Cocconeiopsis regularis (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 176 (2000). 

Basionym: Navicula regularis Hust., Duke Univ. Mar. Sta. Bull. 6, p. 30, pl. 7: fig. 17 (1955). 

Cocconeiopsis wrightii (O´Meara) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 176 (2000). 

Basionym: Cocconeis wrightii O´Meara, Quart. J. Microscop. Soc. 7, p. 246, pl. 7: fig. 6. (1876). 

COCCONEIS Ehrenb. 


Cocconeis capensis (Cholnoky) Witkowski, Iconogr. Diatomol. 7, p. 103 (2000). 

Basionym: Cocconeis clandestina var. capensis Cholnoky, Bot. Mar. 5, p. 42, figs 6-7. (1963). 

Cocconeis egregia Gerd Moser, Steindorf et Lange-Bert., Biblioth. Diatomol. 32, p. 60, pl. 13: figs 1-9 (1995). 

Cocconeis hauniensis Witkowski emend. Witkowski, Iconogr. Diatomol. 7, p. 108 (2000). 

Cocconeis hauniensis Witkowski, Nord. J. Bot. 13, p. 468, pl. 1: figs A-N, pl. 2: figs A-C, pl. 3: figs A-D (1993). 

Cocconeis irregularis (Schulz-Danzig) Witkowski, Iconogr. Diatomol. 7, p. 109 (2000). 

Basionym: Cocconeis scutellum var. irregularis Schulz-Danzig, Bot. Arch. 13, p. 193, fig. 48. (1926). 

Cocconeis krammeri Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 39, pl. 19: figs 11-19 (1996). [Comment: Cocconeis 

krammeri is not documented on pl. 19: figs 20-22´ as cited on p. 39.] 

Cocconeis pseudothumensis E.Reichardt, Ber. Bayer. Bot. Ges. 53, p. 99, pl. 1: figs 1-5 (1982). 

COSMIONEIS D.G.Mann et A.J.Stickle in F.E. Round, R.M. Crawford & D.G. Mann 


Cosmioneis eta (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 176 (2000). 

Basionym: Navicula eta Cleve, Diatomiste 2, p. 13, pl. 1: fig. 5 (1893). 

COSTERICARDIA Lange-Bert. (subgenus within GOMPHOSPHENIA), Biblioth. Diatomol. 38, p. 192-193 (1998). 

[Typus subgeneris: Gomphosphenia oahuensis (Hust.) Lange-Bert., Biblioth. Diatomol. 38, p. 193 (1998).] 

COXIA Lange-Bert. et Gerd Moser, nom. illeg., Biblioth. Diatomol. 38, p. 107-110 (1998). [Comment: intended type of 

the name of the genus COXIA Lange-Bert. et Gerd Moser: Coxia guillauminii (Manguin [ex Kociolek et Reviers]) Lange- 

Bert., Biblioth. Diatomol. 38, p. 110 (1998). COXIA Lange-Bert. et Gerd Moser is a later homonym of COXIA Endlicher, 

hence illegitimate. For COXIA Endlicher see: http://www.ipni.org/ipni/IpniServletid=32773-1&query_type=by_id]. 

Coxia guillauminii (Manguin ex Kociolek et Reviers) Lange-Bert., Biblioth. Diatomol. 38, p. 110 (1998). 

Basionym: Amphipleura guillauminii Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 200 (1996). 

CRATICULA Grunow 


Craticula accomodiformis Lange-Bert., Biblioth. Diatomol. 27, p. 13, pl. 69: figs 6-8, 10, pl. 71: fig. 6 (1993). 

Craticula acidoclinata Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 41, pl. 26: figs 1-3 (1996). 

Craticula buderi (Hust.) Lange-Bert., Iconogr. Diatomol. 9, p. 101 (2000). 

Basionym: Navicula buderi Hust., Ber. Deutsch. Bot. Ges. 67, p. 279, figs 11-15 (1954). 

[Comment: Erroneously this combination again was marked as new by Lange-Bert. in E. Nevo & S.P. Wasser: Biodiversity 

of cyanoprocaryotes and algae of continental Israel, p. 236 (2000).] 

Craticula citrus (Krasske) E.Reichardt, Diatom Res. 12, p. 305 (1997). 

Basionym: Navicula citrus Krasske, Bot. Arch. 3, p. 199, fig. 14 (1923). 

Craticula dissociata (E.Reichardt) E.Reichardt, Diatom Res. 12, p. 305 (1997). 

Basionym: Navicula dissociata E.Reichardt, Biblioth. Diatomol. 6, p. 44, pl. 11: fig. 20 (1984). 

Craticula halopannonica Lange-Bert., Diatoms of Europe 2, p. 218, pl. 85: figs 1-6 (2001). 



18 

Craticula halophilioides (Hust.) Lange-Bert., Diatoms of Europe 2, p. 114, (2001). 

Basionym: Navicula halophilioides Hust., Oesterr. Bot. Z. 106, p. 402, figs 4-7 (1959). 

Craticula minusculoides (Hust.) Lange-Bert., Diatoms of Europe 2, p. 115, (2001). 

Basionym: Navicula minusculoides Hust., Ber. Deutsch. Bot. Ges. 60, p. 68, fig. 5 (1942). 

Craticula molesta (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 130 (1996). 

Basionym: Navicula molesta Krasske, Arch. Hydrobiol. 33, p. 528, pl. 11: fig. 20 (1938). 

Craticula molestiformis (Hust.) Lange-Bert., Iconogr. Diatomol. 9, p. 101 (2000). 

Basionym: Navicula molestiformis Hust., Expl. Parc. Natl. Albert. Mission Damas, p. 86, figs 5-9 (1949). 

Craticula riparia (Hust.) Lange-Bert. var. riparia, Biblioth. Diatomol. 27, p. 14 (1993). 

Basionym: Navicula riparia Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 42, p. 52, figs 77- 78 (1942). 

Craticula riparia var. mollenhaueri Lange-Bert., Biblioth. Diatomol. 27, p. 14-15, pl. 70: figs 10-13 (1993). 

Craticula subhalophila (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 16 (1993). 

Basionym: Navicula subhalophila Hust., Arch. Hydrobiol. Suppl. 15, p. 229, pl. 17: fig. 1 (1937). 

Craticula submolesta (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 42 (1996). 

Basionym: Navicula submolesta Hust., Mission H. Damas (1935-1936), Explor. Parc Natl. Albert. 8, p. 86, pl. 5: figs 16-18 

(1949). 

Craticula sverirschopkae Lange-Bert., Diatoms of Europe 2, p. 218, pl. 92: figs 17-19 (2001). 

Craticula vixnegligenda Lange-Bert., Biblioth. Diatomol. 27, p. 16, pl. 70: figs 15-18, pl. 72: figs 13-15 (1993). 

Craticula vixvisibilis (Hust.) Lange-Bert. var. vixvisibilis, Biblioth. Diatomol. 27, p. 17 (1993). 

Basionym: Navicula vixvisibilis Hust., Arch. Hydrobiol. Suppl. 15, p. 230, pl. 17: fig. 2 (1937). 

Craticula vixvisibilis var. distinctior Lange-Bert., Biblioth. Diatomol. 27, p. 18, pl. 72: figs 1-4 (1993). 

CULTRIA Metzeltin et Lange-Bert. (subgenus within EUNOTIA) Iconogr. Diatomol. 5, p. 46 (1998). [Typus subgeneris: 

Eunotia falcifera Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 60 (1998).] 

CYMATONEIS Cleve 


Cymatoneis margarita Witkowski, Iconogr. Diatomol. 7, p. 179, 425, pl. 109: figs 9-17 (2000). 

CYMBELLA C.Agardh 


Cymbella affiniformis Krammer, Diatoms of Europe 3, p. 162, pl. 25: figs 1-8 (2002). 

Cymbella affinis var. procera Krammer, Diatoms of Europe 3, p. 161, pl. 22: figs 8-13 (2002). 

Cymbella alpestris Krammer, Diatoms of Europe 3, p. 163, pl. 33: figs 1-13 (2002). 

Cymbella arctissima Metzeltin in Krammer, Diatoms of Europe 3, p. 170, pl. 106: figs 1-3 (2002). 

Cymbella bengaliformis Krammer, Diatoms of Europe 3, p. 172, pl. 142: figs 8-9 (2002). [Comment: Cymbella 

bengaliformis is not documented on pl. 133: figs 1-4 as cited on p. 172.] 

Cymbella buechleri Krammer, Diatoms of Europe 3, p. 171, pl. 116: fig. 1 (2002). 

Cymbella cantonatii Lange-Bert., Diatoms of Europe 3, p. 174, pl. 172: figs 1-7 (2002). 

Cymbella cistuliformis Krammer, Diatoms of Europe 3, p. 170, pl. 101: figs 1-4, pl. 102: figs 1-5 (2002). 

Cymbella descripta (Hust.) Krammer et Lange-Bert., Biblioth. Diatomol. 9, p. 24 (1985). 

Basionym: Navicula descripta Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 43, p. 170, fig. 25 (1943). 

Cymbella diversa Krammer, Diatoms of Europe 3, p. 168, pl. 68: figs 1-9 (2002). 

Cymbella diversiformis Krammer, Diatoms of Europe 3, p. 167, pl. 67: figs 1-2 (2002). 

Cymbella diversistigmata Krammer, Diatoms of Europe 3, p. 165, pl. 40: figs 1-11 (2002). 

Cymbella dorsirostrata Krammer, Diatoms of Europe 3, p. 159, pl. 10: figs 1-9 (2002). 

Cymbella duplopunctata Krammer, Diatoms of Europe 3, p. 171, pl. 116: figs 2-6 (2002). 



19 

Cymbella elginensis Krammer, Bacillaria 4, p. 136 (1981). 

Replaced synonym: Cymbella turgida W.Greg., Quart. J. Microsc. Sci. 4, p. 5, pl. 1: fig 18 (1856). 

Cymbella elizabethana Krammer, Diatoms of Europe 3, p. 83, 168, pl. 26: figs 9-10 (2002). [Comment: on p. 242 (= caption 

of pl. 26: figs 9-10) the authors are cited as "Krammer et Buechler".] 

Cymbella excisa var. angusta Krammer, Diatoms of Europe 3, p. 159, pl. 9: figs 8-13 (2002). 

Cymbella excisa var. procera Krammer, Diatoms of Europe 3, p. 159, pl. 9: figs 1-7 (2002). 

Cymbella excisa var. subcapitata Krammer, Diatoms of Europe 3, p. 159, pl. 10: figs 14-18 (2002). 

Cymbella excisiformis Krammer var. excisiformis, Diatoms of Europe 3, p. 160, pl. 11: figs 9-16 (2002). 

Cymbella excisiformis var. nonprotracta Krammer, Diatoms of Europe 3, p. 160, pl. 37: figs 8-15 (2002). 

Cymbella exigua Krammer, Diatoms of Europe 3, p. 159, pl. 10: figs 19-24 (2002). 

Cymbella falaisensis (Grunow) Krammer et Lange-Bert., Biblioth. Diatomol. 9, p. 26 (1985). 

Basionym: Navicula falaisensis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, p. 108, pl. 14: fig. 5 (1880). 

Cymbella geddiana Krammer et Lange-Bert., Diatoms of Europe 3, p. 62, 165, pl. 44: figs 14-18 (2002). [Comment: on p. 

165 "Lange-Bert." is missing.] 

Cymbella gemeinhardtii Metzeltin et Krammer, Diatoms of Europe 3, p. 172, pl. 152: figs 1-7 (2002). [Comment: Cymbella 

gemeinhardtii is not documented on pl. 146: figs 1-7 as cited on p. 172.] 

Cymbella gondwana Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 101-102, pl. 119: figs 6-11, pl. 121: figs 

5-6 (2000). 

Cymbella graciliformis Krammer, Diatoms of Europe 3, p. 167, pl. 54: figs 12-15 (2002). 

Cymbella guettingeri Krammer, Diatoms of Europe 3, p. 169, pl. 75: figs 1-9 (2002). 

Cymbella halophila Krammer, Diatoms of Europe 3, p. 172, pl. 135: figs 1-7 (2002). [Comment: Cymbella halophila is not 

documented on pl. 129: figs 1-7 as cited on p. 172.] 

Cymbella hantzschiana var. borealis Krammer, Diatoms of Europe 3, p. 162, pl. 29: figs 7-12 (2002). 

Cymbella hantzschiana var. strontiana Krammer, Diatoms of Europe 3, p. 162, pl. 30: figs 9-14 (2002). 

Cymbella hedinii Krammer, Diatoms of Europe 3, p. 159, pl. 29: figs 13-16 (2002). 

Cymbella hofmanniae Lange-Bert. et Krammer, Diatoms of Europe 3, p. 164, pl. 43: figs 1-11 (2002). 

Cymbella hustedtii var. compacta Krammer, Diatoms of Europe 3, p. 173, pl. 160: figs 19-20 (2002). [Comment: Cymbella 

hustedtii var. compacta is not documented on pl. 151a: figs 19-20 as cited on p. 173.] 

Cymbella hustedtii var. crassipunctata Krammer, Diatoms of Europe 3, p. 173, pl. 160: figs 21-27 (2002). [Comment: 

Cymbella hustedtii var. crassipunctata is not documented on pl. 151a: figs 21-27 as cited on p. 173.] 

Cymbella hustedtii var. rhombica Krammer, Diatoms of Europe 3, p. 173, pl. 160: figs 14-18 (2002). [Comment: Cymbella 

hustedtii var. rhombica is not documented on pl. 151a: figs 14-18 as cited on p. 173.] 

Cymbella hybrida var. lanceolata Krammer, Biblioth. Diatomol. 9, p. 27, pl. 8: fig. 12 (1985). [As a part of the protologue, 

Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 145: 4-5 is cited.] 

Cymbella inca Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 103, pl. 118: figs 9-10 (2000). 

Cymbella incerta var. crassipunctata Krammer, Biblioth. Diatomol. 9, p. 28, pl. 8: figs 6-7 (1985). [As a part of the 

protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 136: 1-3 is 

cited.] 

Cymbella incurvata Krammer, Diatoms of Europe 3, p. 163, pl. 35: figs 1-9 (2002). 

Cymbella italica Krammer, Diatoms of Europe 3, p. 163, pl. 32: figs 1-6 (2002). 

Cymbella kemiana Krammer, Diatoms of Europe 3, p. 169, pl. 115: figs 1-7 (2002). 

Cymbella kolbei var. angusta Krammer, Diatoms of Europe 3, p. 160, pl. 14: figs 24-28 (2002). 

Cymbella krasskei Krammer, Biblioth. Diatomol. 9, p. 28-29, pl. 9: figs 3-4 (1985). 



20 

Cymbella laevis var. lata Krammer, Diatoms of Europe 3, p. 174, pl. 192: figs 1-5 (2002). [Comment: on p. 174 the name is 

erroneously marked as "nov. spec.".] 

Cymbella lanceolata var. bottnica Krammer, Diatoms of Europe 3, p. 172, pl. 148: figs 1-6 (2002). [Comment: Cymbella 

lanceolata var. bottnica is not documented on pl. 142: figs 1-6 as cited on p. 172.] 

Cymbella lancettuliformis Krammer, Diatoms of Europe 3, p. 167, pl. 54: figs 7-11 (2002). 

Cymbella lange-bertalotii Krammer, Diatoms of Europe 3, p. 174, pl. 179: figs 1-4 (2002). 

Cymbella leptoceroides var. sublanceolata Krammer, Diatoms of Europe 3, p. 174, pl. 161: fig. 11 (2002). 

Cymbella maggiana Krammer, Diatoms of Europe 3, p. 165, pl. 45: figs 6-13 (2002). 

Cymbella metzeltinii Krammer, Diatoms of Europe 3, p. 166, pl. 49: figs 6-11 (2002). 

Cymbella mexicana var. lata Krammer, Diatoms of Europe 3, p. 174, pl. 169: figs 10-13 (2002). [Comment: on p. 174 the 

name is erroneously marked as "nov. comb.".] 

Cymbella modicepunctata Krammer, Diatoms of Europe 3, p. 171, pl. 117: figs 1-4 (2002). 

Cymbella neocistula Krammer var. neocistula, Diatoms of Europe 3, p. 169, pl. 86: figs 1, 6-7, pl. 87: figs 1-9 (2002). 

[Comment: on p. 95 additional figs are given: pl. 85: figs 1-4, pl. 86: figs 2-5, 8, pl. 88: figs 1-8, pl. 90: figs 1-8, pl. 91: figs 

1-6 and pl. 92: figs 1-3 (pl. 86: fig. 8 which is referred to on p. 95 shows Bacillaria cistula Ehrenb.).] 

Cymbella neocistula var. islandica Krammer, Diatoms of Europe 3, p. 170, pl. 93: figs 1-5 (2002). [Comment: Cymbella 

neocistula var. islandica is not documented on pl. 89: figs 1-5 as cited on p. 170.] 

Cymbella neocistula var. lunata Krammer, Diatoms of Europe 3, p. 169, pl. 89: figs 1-7 (2002). 

Cymbella neoleptoceros Krammer var. neoleptoceros, Diatoms of Europe 3, p. 173, pl. 156: figs 1-8, pl. 157: 1-19 (2002). 

[Comment: Cymbella neoleptoceros is not documented on pl. 150: figs 1-12 as cited on p. 173.] 

Cymbella neoleptoceros var. tenuistriata Krammer, Diatoms of Europe 3, p. 173, pl. 160: figs 1-6 (2002). [Comment: 

Cymbella neoleptoceros var. tenuistriata is not documented on pl. 151a: figs 1-6 as cited on p. 173.] 

Cymbella neuquina var. fastigata (Krasske) Krammer, Maidana et Díaz Villanueva, Nova Hedwigia 74, p. 341 (2002). 

Basionym: Cymbella cistula var. fastigata Krasske, Ann. Acad. Sci. Fenn., Ser. A, IV, Biol., p. 86, figs 27-28 (1949). 

Cymbella novazeelandiana Krammer, Diatoms of Europe 3, p. 166, pl. 47: figs 6-9 (2002). 

Cymbella obtusiformis Krammer, Diatoms of Europe 3, p. 167, pl. 66: figs 1-7 (2002). 

Cymbella oestrupii Lange-Bert. et Krammer, Diatoms of Europe 3, p. 163, pl. 34: figs 9-11 (2002). 

Cymbella omaniana Krammer, Diatoms of Europe 3, p. 166-167, pl. 55: figs 1-6 (2002). 

Cymbella pantocsekii Krammer, Diatoms of Europe 3, p. 162-163, pl. 55: figs 8-9 (2002). 

Cymbella parviformis Krammer, Diatoms of Europe 3, p. 160, pl. 15: figs 8-12 (2002). 

Cymbella parviformis Krammer, Diatoms of Europe 3, p. 168, pl. 69: figs 1-9 (2002). 

Cymbella peraffinis R.Tynni in Krammer, Diatoms of Europe 3, p. 160, pl. 14: figs 1-7 (2002). [Comment: validation of 

Cymbella peraffinis R.Tynni, nom. inval., Geol. Surv. Finland 296, p. 37, pl. 13: figs 111a-b (1978). A contribution of 

R.Tynni is deduced from p. 4.] 

Cymbella peraspera Krammer var. peraspera, Diatoms of Europe 3, p. 171, pl. 132: figs 1-6 (2002). 

Cymbella peraspera var. colombiana Krammer et Lange-Bert., Diatoms of Europe 3, p. 172, pl. 134: figs 1-6 (2002). 

[Comment: Cymbella peraspera var. colombiana is not documented on pl. 128: figs 1-7 as cited on p. 172.] 

Cymbella percapitata Krammer, Diatoms of Europe 3, p. 161, pl. 20: figs 1-8 (2002). 

Cymbella percymbiformis Krammer, Diatoms of Europe 3, p. 167, pl. 56: figs 1-7 (2002). 

Cymbella perfossilis Krammer, Diatoms of Europe 3, p. 170, pl. 100: figs 1-4 (2002). 

Cymbella perjaponica Krammer et Lange-Bert., Diatoms of Europe 3, p. 64, 165, pl. 46: figs 7-12 (2002). [Comment: 

erroneously pl. 46: fig. 6 is cited also in the protologue.] 

Cymbella perparva Krammer, Diatoms of Europe 3, p. 160, pl. 18: figs 1-15 (2002). 



21 

Cymbella pervarians Krammer, Diatoms of Europe 3, p. 164, pl. 41: figs 7-12 (2002). 

Cymbella proxima var. borealis Krammer, Diatoms of Europe 3, p. 170, pl. 111: figs 1-4 (2002). 

Cymbella proxima var. bottnica Krammer, Diatoms of Europe 3, p. 171, pl. 114: figs 1-7 (2002). 

Cymbella reichardtii Krammer, Biblioth. Diatomol. 9, p. 32, pl. 5: figs 16-25 (1985). [As a part of the protologue, Krammer 

& Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 119: 32-36 is cited.] 

Cymbella reviersiana Lange-Bert. et Gerd Moser, nom. inval., Biblioth. Diatomol. 32, p. 81, pl. 20: figs 9-12. (1995). 

[Comment: lacking a type and a Latin diagnosis, this name was invalidly published. Validly published as Encyonopsis 

reviersiana Lange-Bert., Gerd Moser et Krammer, Biblioth. Diatomol. 37, p. 126, 201, pl. 158: figs 13-15 (1997).] 

Cymbella robertii Krammer, Diatoms of Europe 3, p. 83, 168, pl. 25: fig. 14 (2002). [Comment: on p. 240 (= caption of pl. 

25: fig. 14) the authors are cited as "Krammer et Buechler". The type is indicated on p. 83.] 

Cymbella rossii Krammer et Lange-Bert., Nova Hedwigia Beih. 106, p. 94, figs 1-18 (1993). 

Cymbella rumrichae Krammer, Diatoms of Europe 3, p. 166, pl. 50: figs 1-13 (2002). 

Cymbella schilleri Krammer, Diatoms of Europe 3, p. 168, pl. 26: figs 7-8 (2002). 

Cymbella scotlandica Krammer, Diatoms of Europe 3, p. 162, pl. 31: figs 1-7 (2002). 

Cymbella scutariana Krammer, Diatoms of Europe 3, p. 165, pl. 45: figs 1-5 (2002). 

Cymbella simonsenii Krammer, Biblioth. Diatomol. 9, p. 33, pl. 7: figs 1-9 (1985). [As a part of the protologue, Krammer & 

Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 130: 1-3, pl. 10: fig. 6 is cited.] 

Cymbella sinensis Metzeltin et Krammer, Diatoms of Europe 3, p. 171, pl. 121: figs 1-8 (2002). 

Cymbella strontiana Krammer, Diatoms of Europe 3, p. 162, pl. 30: figs 1-8 (2002). 

Cymbella subantarctica Krammer, Diatoms of Europe 3, p. 168, pl. 73: figs 1-6 (2002). 

Cymbella subarctica Krammer, Diatoms of Europe 3, p. 170, pl. 105: figs 2, 6-7 (2002). 

Cymbella subaspera Krammer var. subaspera, Diatoms of Europe 3, p. 172, pl. 136: figs 1-5 (2002). [Comment: Cymbella 

subaspera var. subaspera is not documented on pl. 130: figs 1-5 as cited on p. 172.] 

Cymbella subaspera var. salina Krammer, Diatoms of Europe 3, p. 172, pl. 137: figs 1-8 (2002). [Comment: Cymbella 

subaspera var. salina is not documented on pl. 131: figs 1-7 as cited on p. 172.] 

Cymbella subcistula Krammer, Diatoms of Europe 3, p. 169, pl. 84: figs 1-9 (2002). 

Cymbella subhelvetica Krammer, Diatoms of Europe 3, p. 174, pl. 176: figs 1-7 (2002). [Comment: Cymbella subhelvetica 

is not documented on pl. 177: figs 1-7 as cited on p. 174.] 

Cymbella subkolbei Krammer, Diatoms of Europe 3, p. 160, pl. 15: figs 1-7 (2002). 

Cymbella subleptoceros Krammer, Diatoms of Europe 3, p. 173, pl. 154: figs 2-17 (2002). [Comment: Cymbella 

subleptoceros is not documented on pl. 147: figs 10-17 as cited on p. 173. On p. 498 and p. 500 the authors are cited as 

"(Ehrenb.) Kütz.".] 

Cymbella subtruncata Krammer var. subtruncata, Diatoms of Europe 3, p. 161, pl. 18: figs 16-21 (2002). 

Cymbella subtruncata var. similis Krammer, Diatoms of Europe 3, p. 161, pl. 19: figs 14-19 (2002). 

Cymbella subturgidula Krammer, Diatoms of Europe 3, p. 166, pl. 44: figs 19-21 (2002). 

Cymbella superparva Krammer, Diatoms of Europe 3, p. 161, pl. 15: figs 13-15 (2002). 

Cymbella terrafuegiana Krammer, Diatoms of Europe 3, p. 169, pl. 76: figs 1-3 (2002). 

Cymbella ticinensis Krammer, Diatoms of Europe 3, p. 161, pl. 24: figs 17-19 (2002). 

Cymbella towoetensis Krammer, Diatoms of Europe 3, p. 173-174, pl. 161: figs 13-14 (2002). [Comment: Cymbella 

towoetensis is not documented on pl. 152: figs 13-14 as cited on p. 173.] 

Cymbella transsilvanica Krammer, Diatoms of Europe 3, p. 173, pl. 158: figs 6-11 (2002). [Comment: Cymbella 

transsilvanica is not documented on pl. 151: figs 6-11 as cited on p. 173.] 



22 

Cymbella tropica Krammer var. tropica, Diatoms of Europe 3, p. 164, pl. 44: figs 7-10 (2002). [Comment: on p. 278 (= 

caption of pl. 44) also mentioned as Cymbella tropica Krammer et Metzeltin.] 

Cymbella tropica var. tenuipunctata Krammer, Diatoms of Europe 3, p. 164, pl. 49: figs 12-13 (2002). 

Cymbella tsoneka Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 104, pl. 119: figs 1-4 (2000). 

Cymbella tumidula var. lancettula Krammer, Biblioth. Diatomol. 9, p. 35, pl. 6: figs 5-8 (1985). [As a part of the 


cited.] 

Cymbella turgidula var. bengalensis Krammer, Diatoms of Europe 3, p. 166, pl. 48: figs 8-11 (2002). 

Cymbella turgidula var. venezolana Krammer, Diatoms of Europe 3, p. 166, pl. 48: figs 12-17 (2002). 

Cymbella turgiduliformis Krammer, Diatoms of Europe 3, p. 165, pl. 47: figs 1-5 (2002). 

Cymbella volkii U.Rumrich, Krammer et Lange-Bert., Biblioth. Diatomol. 27, p. 19, pl. 134: figs 1-10 (1993). 

Cymbella vulgata Krammer var. vulgata, Diatoms of Europe 3, p. 163-164, pl. 36: figs 1-11 (2002). 

Cymbella vulgata var. plivicensis Krammer, Diatoms of Europe 3, p. 164, pl. 32: figs 7-13 (2002). 

Cymbella weslawskii Krammer, Diatoms of Europe 3, p. 162, pl. 27: figs 1-7 (2002). 

Cymbella zambesiana Krammer, Diatoms of Europe 3, p. 164, pl. 44: figs 11-13 (2002). 

CYMBELLONITZSCHIA Hust. 


Cymbellonitzschia szulczewskii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 358, 425, pl. 213: figs 1-7 

(2000). 

CYMBELLOPSIS Krammer, Biblioth. Diatomol. 36, p. 157, (1997). [Typus generis: Cymbellopsis apiculata Krammer, 

Biblioth. Diatomol. 36, p. 157, pl. 2: figs 2-3 (1997).] 

Cymbellopsis apiculata Krammer, Biblioth. Diatomol. 36, p. 157, pl. 2: figs 2-3 (1997). 

Cymbellopsis guyana Krammer, Biblioth. Diatomol. 37, p. 208-209, pl. 196: figs 1-10 (1997). 

Cymbellopsis krammeri Lange-Bert. et Wydrzycka, Iconogr. Diatomol. 9, p. 105, pl. 111: figs 26-27 (2000). 

Cymbellopsis lanceolata Krammer, Biblioth. Diatomol. 37, p. 208, pl. 197: figs 7-13, pl. 198: figs 1-3 (1997). 

Cymbellopsis metzeltinii Krammer, Biblioth. Diatomol. 37, p. 209, pl. 197: figs 1-6, pl. 201: figs 4-8 (1997). 

Cymbellopsis mirabilis Krammer, Biblioth. Diatomol. 37, p. 209-210, pl. 194: figs 6-7 (1997). 

Cymbellopsis neocaledonica Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 111-112, pl. 43: figs 9-13 (1998). 

Cymbellopsis pumila Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 7, pl. 17: figs 10-14 (2001). 

Cymbellopsis terminis Krammer, Biblioth. Diatomol. 37, p. 209, pl. 194: figs 8-9, pl. 195: figs 1-9 (1997). 

CYMBOPLEURA (Krammer) Krammer, Iconogr. Diatomol 6, p. 284 (1999). 

Basionym: CYMBELLA subgen. CYMBOPLEURA Krammer in Helmcke & Krammer: Micromorphology of diatom valves 

11, p. 20 (1982). [Validation of CYMBOPLEURA Krammer, nom. inval., Biblioth. Diatomol. 36, p. 11 (1997). Typus generis 

Cymbella subaequalis Grunow in Van Heurck, Synopsis des diatomées de Belgique, pl. 3: fig. 2.] 

Cymbopleura aequistriata Krammer et Lange-Bert., nom. inval., Biblioth. Diatomol. 38, p. 112-113, pl. 44: figs 5-8 (1998). 

[Comment: since the genus Cymbopleura was not validated before 1999, this name is invalidly published.] 

Cymbopleura crassinervia Lange-Bert., Metzeltin et Krammer, nom. inval., Biblioth. Diatomol. 38, p. 114-115, pl. 45: figs 

1-4, pl. 46: figs 1-3 (1998). [Comment: since the genus Cymbopleura was not validated before 1999, this name is invalidly 

published.] 

Cymbopleura declivis Metzeltin et Krammer, nom. inval. Iconogr. Diatomol. 5, p. 34, pl. 131: figs 1-6, pl. 141: fig. 2 (1998). 


Cymbopleura delicatula ssp. judaica Lange-Bert. et Krammer in Nevo & Wasser, Biodiversity of cyanoprocaryotes and 

algae of continental Israel, p. 242, pl. 18: figs 130-133 (2000). 



23 

Cymbopleura fluminea (R.M.Patrick et Freese) Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 38 (1999). 

Basionym: Cymbella fluminea R.M.Patrick et Freese, Proc. Acad. Nat. Sci. Philadelphia 112 (6), p. 262, pl. 4: fig. 4 (1961). 

Cymbopleura heinii Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 38-39, pl. 57: fig. 1 (1999). 

Cymbopleura kanakarum Lange-Bert. et Krammer, nom. inval., Biblioth. Diatomol. 38, p. 116, pl. 44: figs 1-4 (1998). 


Cymbopleura michelcostei Metzeltin et Krammer, nom. inval., Iconogr. Diatomol. 5, p. 35, pl. 137: figs 1-8 (1998). 

[Comment: since the genus Cymbopleura was not validated before 1999, this name is invalidly published. On page 35 only 

pl. 137: figs 1-6 is referred to.] 

Cymbopleura neocaledonica Lange-Bert. et Krammer, nom. inval., Biblioth. Diatomol. 38, p. 117-118, pl. 44: figs 10-12 

(1998). [Comment: since the genus Cymbopleura was not validated before 1999, this name is invalidly published.] 

Cymbopleura semigibbosa (R.M.Patrick et Freese) Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 39 (1999). 

Basionym: Cymbella semigibbosa R.M.Patrick et Freese, Proc. Acad. Nat. Sci. Philadelphia 112 (6), p. 265, pl. 4: fig. 6 

(1961). 

DECUSSATA (R.M.Patrick) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 9, p. 670, (2000). 

Basionym: DECUSSATA R.M.Patrick [subgenus within NAVICULA Bory], Notul. Nat. Acad. Nat. Sci. Philadelphia 324, p. 

4 (1959). 

Decussata hexagona (Torka) Lange-Bert., Iconogr. Diatomol. 9, p. 671 (2000). 

Basionym: Navicula hexagona Torka, Hedwigia 73, p. 27, fig. 2 (1933). 

Decussata placenta (Ehrenb.) Lange-Bert. et Metzeltin var. placenta, Iconogr. Diatomol. 9, p. 671 (2000). 

Basionym: Navicula placenta Ehrenb., Mikrogeologie, pl. 33: fig. 12/23 (1854). 

Decussata placenta var. obtusa (F. Meister) Lange-Bert., Iconogr. Diatomol. 9, p. 671 (2000). 

Basionym: Navicula placenta var. obtusa F. Meister, Kieselalgen aus Asien, p. 37: fig. 99 (1932). 

DENTICULA Kütz. 


Denticula creticola (Østrup) Lange-Bert., Nova Hedwigia Beih. 106, p. 127 (1993). 

Basionym: Nitzschia creticola Østrup, Danske Diatomeer, p. 146, pl. 4: figs 98 a, b (1910). 

Denticula erifuga Lange-Bert., Iconogr. Diatomol. 3, p. 66 (1996). 

Replaced synonym: Denticula elegans f. constricta Krasske, Ann. Acad. Sient. Fenn. Ser. A, IV. Biol. 14, p. 87 (1949). 

Denticula eximia Krammer et Lange-Bert., Biblioth. Diatomol. 15, p. 64, pl. 41: figs 2-5 (1987). [As a part of the 

protologue, Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 98: figs 8-18 

is cited.] 

Denticula kuetzingii var. rumichiae Krammer, Biblioth. Diatomol. 15, p. 66, pl. 43: figs 11-14 (1987). [As a part of the 

protologue, Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 100: figs 18- 

22 is cited. Comment: Denticula kuetzingii var. rumichiae on p. 66 and p. 232 is misspelled as "Denticula kuetzingii var. 

rumrichae".] 

Denticula watanabei Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 132-133, pl. 60: figs 1-10 (1998). 

DENTICULOIDEA Lange-Bert. (subgenus within NITZSCHIA), Biblioth. Diatomol. 27, p. 153 (1993). [Typus 

subgeneris: Nitzschia semirobusta Lange-Bert., Biblioth. Diatomol. 27, p. 149, pl. 120: figs 7-10 (1993).] 

DIADESMIS Kütz. 


DIADESMIS subgen. PARADIADESMIS [see PARADIADESMIS]. 

Diadesmis andegavensis (H.Germ.) Lange-Bert., Biblioth. Diatomol. 38, p. 136 (1998). 

Basionym: Navicula brekkaensis var. andegavensis H.Germ., Cryptog. Algol. 1, p. 21, figs 20-34 (1980). 

Diadesmis arctica Lange-Bert. et Genkal, Iconogr. Diatomol. 6, p. 40, pl. 21: figs 8-14 (1999). [Comment: Diadesmis 

arctica is not documented on pl. 21: fig. 15 as cited in the 1 st edition on p. 40, reference corrected in the 2 nd edition.] 

Diadesmis arcuata Lange-Bert., Biblioth. Diatomol. 38, p. 136 (1998). 

[Comment: According to Art. 58.1 (St. Louis Code) this binomen is not a new combination but should be treated as a new 

name.] 



24 

Replaced synonym: Navicula arcuata Heiden in: Drygalski, Deutsche Südpolar-Expedition 1901-1903, nom. illeg., Bd. 8, H. 

5, p. 628, pl. 3: fig. 80 (1928); non Navicula arcuata Pantocsek, pl. 6: fig. 97 (1892). 

Diadesmis bigibba (Hust.) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 138 (1998). 

Basionym: Navicula brekkaensis var. bigibba Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 402: fig. 28 (1936). 

Diadesmis biscutella Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 138-139, pl. 28: figs 1-4, 10 (1998). 

[Comment: the authors are cited as "Lange-Bert. in Moser et al. 1998" in Lange-Bertalot & Werum (2001).] 

Diadesmis brekkaensoides (W.Bock) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 139 (1998). 

Basionym: Navicula brekkaensoides W.Bock, Nova Hedwigia 5, p. 224-225, fig. 1A, B (1963). 

[≡ Navicula brekkaensis var. brekkaensoides (W.Bock) Lange-Bert.] 

Diadesmis confervaceoides Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 106, pl. 81: figs 1-10 (2000). 

Diadesmis corrugata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 140-141, pl. 27: figs 1-3, 9 (1998). 

[Diadesmis corrugata is the type of the name of the subgenus PARADIADESMIS Lange-Bert. et Le Cohu. Comment: the 

authors are cited as "Lange-Bert. in Moser et al. 1998" in Lange-Bertalot & Werum (2001).] 

Diadesmis discordabilis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 142, pl. 27: figs 4-5, 10, 12-13 

(1998). 

Diadesmis dissimilis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 143, pl. 29: fig. 6 (1998). 

Diadesmis elegans Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 144-145, pl. 29: figs 7-8 (1998). 

Diadesmis fukushimae Lange-Bert., M.Werum et Broszinski, Diatom 17, p. 6, figs 33-38, 47-57 (2001). 

Diadesmis gallica var. perpusilla (Grunow) Lange-Bert., Iconogr. Diatomol. 2, p. 45 (1996). 

Basionym: Navicula perpusilla Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 10, p. 552, pl. 2: fig. 7 (1860). 

Diadesmis implicata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 145-146, pl. 27: figs 6-8, 11 (1998). 

Diadesmis lacunosa Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 147, pl. 29: figs 1-3 (1998). 

[Comment: the authors are cited as "Lange-Bert. in Moser et al. 1998" in Lange-Bertalot & Werum (2001).] 

Diadesmis pantropica Lange-Bert., Iconogr. Diatomol. 9, p. 109, pl. 82: figs 1-5 (2000). 

Diadesmis paracontenta Lange-Bert. et M.Werum, Iconogr. Diatomol. 6, p. 41, pl. 21: figs 2-5 (1999). [Comment: 

Diadesmis paracontenta is not documented on pl. 21: fig. 6 as cited in the 1 st edition on p. 41, reference corrected in the 2 nd 

edition. Comment: the Latin diagnosis erroneously refers to fig. 6, it should refer to fig. 5 as corrected in the 2 nd edition.] 

Diadesmis paracontenta ssp. magisconcava Lange-Bert., Diatom 17, p. 9, figs 11-14, 60-64 (2001). 

Diadesmis tabellariaeformis (Krasske) Lange-Bert. et Wojtal, Iconogr. Diatomol. 9, p. 110 (2000). 

Basionym: Navicula tabellariaeformis Krasske, Arch Hydrobiol. 35, p. 384, pl. 11: figs 48-49 (1939). 

Diadesmis tahitiensis Lange-Bert. et M.Werum, Diatom 17, p. 13, figs 22-29, 78-84 (2001). 

Diadesmis templiniana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 111, pl. 82: figs 6-18 (2000). 

Diadesmis virginiana Lange-Bert., Diatom 17, p. 17, figs 18-21, 100-105 (2001). 

DIATOMA Bory 


Diatoma ehrenbergii f. capitulata (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 22 (1993). 

Basionym: Diatoma vulgare var. capitulatum Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 12, p. 364 (1862). 

Diatoma moniliformis ssp. ovalis (Fricke) Lange-Bert., U.Rumrich et G.Hofmann, Acta Biol. Benrodis 3, p. 116 (1991). 

Basionym: Diatoma ovalis Fricke in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 268: figs 68-73 (1906). 

Diatoma ochridana Lange-Bert. et U.Rumrich, Acta Biol. Benrodis 3, p. 118, 120, figs 19-25, 45-48 (1991). 

Diatoma problematica Lange-Bert., Biblioth. Diatomol. 27, p. 23, pl. 6: figs 11-15, 21-27, pl. 7: figs 1-6, pl. 8: figs 1-5 

(1993). 

DIATOMELLA Grev. 


Diatomella lecohui Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 148, pl. 55: figs 6-16 (1998). 



25 

DICKIEIA Berk. ex Kütz. 


Dickieia resistans Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 181, 425-426, pl. 150: figs 16-20, 21 

(2000). 

Dickieia soodensis (Krasske) E.Reichardt, Diatom Res. 12, p. 314 (1997). 

Basionym: Navicula soodensis Krasske, Arch. Hydrobiol. 18, p. 272, figs 20-22 (1927). 

Dickieia wajdae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 181, 426, pl. 108: figs 19-23 (2000). 

DIDYMOSPHENIA M.Schmidt 


Didymosphenia clavaherculis (Ehrenb.) Metzeltin et Lange-Bert., Nova Hedwigia 60, p. 388 (1995). 

Basionym: Diomphala clavaherculis Ehrenb., Mikrogeologie, pl. A15: fig. 93 (1854). 

[= Diomphala clavaherculis Ehrenb., Ber. Bekanntm. Verh. Königl. Preuss. Akad. Wiss. Berlin 1842, p. 338 (1842), 

according to Metzeltin & Lange-Bertalot, Nova Hedwigia 60, p. 388 (1995).] 

Didymosphenia curvata (Skvortzov et D.I.Meyer 1928) Metzeltin et Lange-Bert., Nova Hedwigia 60, p. 390 (1995). 

Basionym: Diomphala geminata var. curvata Skvortzov et D.I.Meyer., Proc. Sungare River. Biol. Station 1 (5), p. 31: fig. 

137 (1928). 

Didymosphenia pumila Metzeltin et Lange-Bert., Nova Hedwigia 60, p. 390-391, pl. 8: figs 5-6, pl. 10: figs 1-6 (1995). 

DIPLONEIS (Ehrenb.) Cleve 


Diploneis caffra (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 183 (2000). 

Basionym: Diploneis interrupta var. caffra Giffen, Nova Hedwigia Beih. 31, p. 273, figs 29-30 (1970). 

Diploneis chilensis (Hust.) Lange-Bert., Iconogr. Diatomol. 9, p. 112 (2000). 

Basionym: Diploneis smithii var. chilensis Hust., Arch. Hydrobiol. 18, p. 242, pl. 8: fig. 9 (1927). 

Diploneis kahlii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 112-113, pl. 103: 1-9, pl. 104: figs 1-3 (2000). 

Diploneis kawabatae Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 149-150, pl. 42: figs 1-7 (1998). 

Diploneis krammeri Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 9, p. 652-653, pl. 4: figs 1-10, 12, pl. 5: figs 1-5, pl. 6: 

figs 1-5 (2000). 

Diploneis ovalis ssp. arctica Lange-Bert., Iconogr. Diatomol. 6, p. 42-43, pl. 43: figs 1-3, pl. 44: figs 1-5 (1999). [Comment: 

name and diagnosis on p. 43 only in the 2 nd edition.] 

EHRENBERGIA Witkowski, Lange-Bert. et Metzeltin, nom. illeg., Iconogr. Diatomol. 7, p. 30, 426 (2000). [Comment: 

intended type: Ehrenbergia granulosa (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 31 (2000). 

EHRENBERGIA Witkowski, Lange-Bert. et Metzeltin is a later homonym of EHRENBERGIA K.P.J.Sprengel, hence 

illegitimate. For EHRENBERGIA see also: http://www.ipni.org/ipni/IpniServletid=34541-1&query_type=by_id 

http://www.ipni.org/ipni/IpniServletid=41709-1&query_type=by_id]. 

Ehrenbergia granulosa (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 31 (2000). 

Basionym: Odontodiscus granulosus Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 113, pl. 7: fig. 130 (1880). 

Ehrenbergia hauckii (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 31 (2000). 

Basionym: Coscinodiscus hauckii Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 94: fig. 29 (1881). 

ENCYONEMA Kütz. 


Encyonema accedens Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 118-119, pl. 47: figs 1-4 (1998). 

Encyonema aequilongum Lange-Bert. et Krammer, Biblioth. Diatomol. 36, p. 61, 160, pl. 98: figs 17-21 (1997). 

Encyonema affine Metzeltin et Krammer, Biblioth. Diatomol. 37, p. 189, pl. 125: figs 6-8 (1997). [Comment: on p. 298 (= 

caption of pl. 125: figs 6-8 the authors are given as "Krammer et Metzeltin".] 

Encyonema africanum Krammer, Biblioth. Diatomol. 37, p. 191, pl. 129: figs 4-18 (1997). 

Encyonema alpiniforme Krammer, Biblioth. Diatomol. 37, p. 183-184, pl. 101: figs 6-14 (1997). 

Encyonema altiplanense Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 114, pl. 118: figs 11-14 (2000). 



26 

Encyonema amanianum Krammer, Biblioth. Diatomol. 36, p. 168, pl. 46: figs 4-8 (1997). 

Encyonema ampliatum Krammer et Lange-Bert., Biblioth. Diatomol. 36, p. 139, 177, pl. 54: figs 1-5 (1997). 

Encyonema angustecapitatum Krammer, Biblioth. Diatomol. 37, p. 192, pl. 130: figs 8-15 (1997). 

Encyonema apiculatum Krammer, Biblioth. Diatomol. 36, p. 176, pl. 76: figs 1-2 (1997). 

Encyonema blancheanum var. neocaledonicum (Manguin ex Kociolek et Reviers) Lange-Bert. in Jahn et al., Lange- 

Bertalot-Festschrift, p. 632 (2001). 

Basionym: Cymbella subturgida var. neocaledonica Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 210 (1996). 

[– Cymbella subturgida var. neocaledonica Manguin, nom. inval., Mém. Mus. Natl. Hist. Nat., Nouv. Sér. B, Bot. 12 (1), p. 

28, pl. 5: figs 7a, b (1962).] 

[– Encyonema blancheanum var. neocaledonicum (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, 

nom. inval., Biblioth. Diatomol. 38, p. 33 (1998).] 

Encyonema brehmiforme Krammer, Biblioth. Diatomol. 37, p. 182, pl. 104: figs 4-19 (1997). 

Encyonema brevicapitatum Krammer, Biblioth. Diatomol. 36, p. 170, pl. 34: figs 1-7 (1997). 

Encyonema canadense Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 41, 187-188, pl. 120: figs 6-7 (1997). 

[Comment: the type of the name of the species is referred to as Präp. "Can 50LF" (p. 188) respectivly "Can 49LF" (p. 41).] 

Encyonema carina Lange-Bert. et Krammer, Biblioth. Diatomol. 36, p. 152, 179-180, pl. 86: figs 1-8 (1997). 

Encyonema caronianum Krammer, Biblioth. Diatomol. 37, p. 189, pl. 131: figs 11-16 (1997). 

Encyonema cascadense Krammer, Biblioth. Diatomol. 37, p. 190, pl. 192: figs 6-12 (1997). 

Encyonema cespitosum var. comensis Krammer, Biblioth. Diatomol. 36, p. 173, pl. 67: figs 1-8 (1997). 

Encyonema cespitosum var. maxima Krammer, Biblioth. Diatomol. 36, p. 174, pl. 69: figs 1-8 (1997). 

Encyonema chilense Krammer, Biblioth. Diatomol. 36, p. 169, pl. 46: figs 1-3 (1997). 

Encyonema directiforme Krammer et Lange-Bert., Biblioth. Diatomol. 36, p. 149, 179, pl. 95: figs 6-11 (1997). [Encyonema 

directiforme Krammer et Lange-Bert. on p. 370 (caption of p. 95) is cited as "Encyonema directiforme Lange-Bert. et 

Krammer".] 

Encyonema distinctepunctatum Krammer, Biblioth. Diatomol. 36, p. 162-163, pl. 20: figs 1-5, pl. 98: figs 1-6 (1997). 

Encyonema distinctum Lange-Bert. et Krammer, Biblioth. Diatomol. 36, p. 56, 158-159, pl. 98: figs 13-16 (1997). 

[- "Cymbella nov. spec. Nr. 2 Mittersee" in Lange-Bertalot & Metzeltin (1996), Iconogr. Diatomol. 2, pl. 88: figs 21-24.] 

Encyonema donatoi Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 52, 189, pl. 125: figs 9-10 (1997). 

Encyonema dubium Krammer, Biblioth. Diatomol. 37, p. 186, pl. 109: figs 8-16 (1997). 

Encyonema elginense var. stigmoideum Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 36, pl. 133: figs 21, 24-25 (1998). 

Encyonema evergladianum Krammer, Biblioth. Diatomol. 37, p. 193, pl. 142: figs 1-7 (1997). 

Encyonema fogedii Krammer, Biblioth. Diatomol. 36, p. 171, pl. 26: figs 1-12, 18-28 (1997). 

Encyonema friedelhinzae Krammer, Biblioth. Diatomol. 36, p. 180, pl. 87: figs 1-15 (1997). 

Encyonema geisslerae Krammer, Nova Hedwigia 65, p. 141, figs 3, 6-7, 13-18 (1997). 

Encyonema graciloides Krammer, Biblioth. Diatomol. 37, p. 194, pl. 195: figs 11-13, pl. 199: figs 5-6 (1997). 

Encyonema guatemalense Krammer, Biblioth. Diatomol. 37, p. 193-194, pl. 142: figs 16-21 (1997). [Comment: on p. 332 

(= caption of pl. 142: figs 16-21) Encyonema guatemalense is cited as "Encyonema guatemalum".] 

Encyonema guettingeri Krammer, Biblioth. Diatomol. 37, p. 188, pl. 132: figs 1-9, 11-12, 10 (1997). 

Encyonema hebridiforme Krammer, Biblioth. Diatomol. 36, p. 165, pl. 11: figs 15-17 (1997). 

Encyonema hintzii Krammer, Biblioth. Diatomol. 36, p. 168, pl. 39: figs 6-9 (1997). 

Encyonema hophense Krammer, Biblioth. Diatomol. 36, p. 162, pl. 33: figs 6-13 (1997). 

Encyonema hungaricum Krammer, Biblioth. Diatomol. 36, p. 167, pl. 49: figs 1-4 (1997). 



27 

Encyonema hustedtii Krammer, Biblioth. Diatomol. 36, p. 166-167, pl. 42: figs 6-8 (1997). 

Encyonema hustedtii var. neocaledonicum Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 121, pl. 47: figs 12-14 

(1998). 

Encyonema inarense Krammer, Biblioth. Diatomol. 36, p. 173, pl. 60: figs 5-8 (1997). 

Encyonema incertum Krammer, Biblioth. Diatomol. 37, p. 190, pl. 137: figs 10-13, pl. 140: figs 5-6 (1997). 

Encyonema incurvatum Krammer, Biblioth. Diatomol. 36, p. 161, pl. 21: figs 21-23 (1997). 

Encyonema indistinctum Krammer, Biblioth. Diatomol. 37, p. 187, pl. 192: figs 1-5, pl. 194: fig. 5 (1997). 

Encyonema insulsum Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 121-122, pl. 47: figs 8-11 (1998). 

Encyonema jemtlandicum Krammer var. jemtlandicum, Biblioth. Diatomol. 36, p. 166, pl. 35: figs 1-9 (1997). 

Encyonema jemtlandicum var. stigmoidea Krammer, Biblioth. Diatomol. 37, p. 195, pl. 194: figs 2-4 (1997). 

Encyonema jemtlandicum var. venezolana Krammer, Biblioth. Diatomol. 36, p. 166, pl. 14: figs 1-5, pl. 50: fig. 8 (1997). 

Encyonema jordaniforme Krammer, Biblioth. Diatomol. 37, p. 191, pl. 201: figs 9-15 (1997). 

Encyonema kamtschaticum Krammer var. kamtschaticum, Biblioth. Diatomol. 36, p. 175, pl. 63: figs 4-11 (1997). 

Encyonema kamtschaticum var. parva Krammer, Biblioth. Diatomol. 36, p. 175, pl. 62: figs 9-13 (1997). 

Encyonema kuelbsii Lange-Bert. et Krammer, Biblioth. Diatomol. 36, p. 55, 158, pl. 98: figs 7-12 (1997). 

[– "Cymbella nov. spec. Nr. 1 Mittersee" in Lange-Bertalot & Metzeltin (1996), Iconogr. Diatomol. 2, pl. 88: figs 15-20.] 

Encyonema kukenanum Krammer, Biblioth. Diatomol. 37, p. 184-185, pl. 106: figs 7-9 (1997). 

Encyonema lancettulum Krammer, Biblioth. Diatomol. 36, p. 160, pl. 21: figs 9-15 (1997). 

Encyonema lange-bertalotii Krammer var. lange-bertalotii, Biblioth. Diatomol. 36, p. 169-170, pl. 5: figs 1-16 (1997). 

Encyonema lange-bertalotii var. inarensis Krammer, Biblioth. Diatomol. 36, p. 170, pl. 29: fig. 14 (1997). 

Encyonema lange-bertalotii var. obscuriformis Krammer, Biblioth. Diatomol. 36, p. 170, pl. 27: figs 13-16 (1997). 

Encyonema latareolatum Krammer, Biblioth. Diatomol. 36, p. 180, pl. 96: figs 1-13 (1997). 

Encyonema latecapitatum Krammer, Biblioth. Diatomol. 36, p. 170-171, pl. 34: figs 8-12 (1997). 

Encyonema latissimum Krammer, Biblioth. Diatomol. 36, p. 176, pl. 80: figs 1-5, pl. 81: figs 1-3 (1997). 

Encyonema latum Krammer, Biblioth. Diatomol. 36, p. 174, pl. 71: figs 1-6 (1997). 

Encyonema lineolatum Krammer var. lineolatum, Biblioth. Diatomol. 36, p. 161, pl. 21: figs 1-8 (1997). 

Encyonema lineolatum var. major Krammer, Biblioth. Diatomol. 36, p. 161-162, pl. 13: figs 1-8 (1997). 

Encyonema lunatum var. borealis Krammer, Biblioth. Diatomol. 36, p. 179, pl. 93: figs 3-5 (1997). 

Encyonema magnum Krammer, Biblioth. Diatomol. 36, p. 177, pl. 99: figs 1-2 (1997). 

Encyonema malaysianum Krammer, Biblioth. Diatomol. 37, p. 186, pl. 109: figs 17-24 (1997). 

Encyonema marginestriatum Krammer et Metzeltin, Biblioth. Diatomol. 37, p. 194, pl. 136: figs 15, 17; pl. 202: figs 16-18 

(1997). 

Encyonema melidense Krammer, Biblioth. Diatomol. 36, p. 168, pl. 37: figs 1-9 (1997). 

Encyonema menisculus Lange-Bert. et Krammer, Biblioth. Diatomol. 38, p. 123, pl. 48: figs 17-20 (1998). 

Encyonema metzeltinii Krammer, Biblioth. Diatomol. 37, p. 190, pl. 134: figs 1-6 (1997). 

Encyonema mexicanum Krammer, Biblioth. Diatomol. 36, p. 172, pl. 61: figs 5-8 (1997). 

Encyonema minutiforme Krammer, Biblioth. Diatomol. 36, p. 159-160, pl. 18: figs 11-15 (1997). 

Encyonema minutiforme var. apicodeclinatum Krammer, Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 115, pl. 111: 

figs 5-6, (7) (2000). 

Encyonema neocaledonicum var. leve Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 124, pl. 48: figs 21-25 (1998). 



28 

Encyonema neogracile Krammer var. neogracile, Biblioth. Diatomol. 36, p. 177-178, pl. 82: figs 1-7, 12-13 (1997). 

Encyonema neogracile var. tenuipunctata Krammer, Biblioth. Diatomol. 36, p. 178, pl. 85: figs 1-6 (1997). 

Encyonema neomuelleri Krammer, Nova Hedwigia 65, p. 142, figs 23-27 (1997). 

Encyonema norvegicum var. alpinum Krammer, Biblioth. Diatomol. 37, p. 185, pl. 107: figs 12-20 (1997). 

Encyonema norvegicum var. capitatum Krammer, Biblioth. Diatomol. 37, p. 185, pl. 107: figs 1-2 (1997). [Comment: on p. 

185 erroneously pl. 106: figs 1-2 is given. On pl. 106: figs. 1-2 E. distinctepunctatum Krammer is documented. On p. 262 (= 

caption of pl. 107: figs 1-2) var. capitatum is cited as "var. capitata".] 

Encyonema norvegicum var. crassipunctatum Krammer, Biblioth. Diatomol. 37, p. 185, pl. 108: figs 5-6 (1997). 

Encyonema obscuriforme Krammer, Biblioth. Diatomol. 36, p. 172, pl. 30: figs 11-15 (1997). 

Encyonema obscurum var. alpina Krammer, Biblioth. Diatomol. 36, p. 171, pl. 29: figs 1-11 (1997). 

Encyonema obtusum Krammer var. obtusum, Biblioth. Diatomol. 37, p. 20, 184, pl. 105: figs 1-7, 11-13 (1997). [Comment: 

Encyonema obtusum var. obtusum is not documented on pl. 105: figs 8-10 as mentioned on p. 184, on the other hand pl. 105: 

fig. 13 is missing on p. 184. On pl. 105: figs 8-10 Encyonema obtusum var. truncatum Krammer, nom. inval. is documented.] 

Encyonema obtusum var. truncatum Krammer, nom. inval., Biblioth. Diatomol. 37, p. 184, pl. 105: figs 8-10 (1997). 

[Comment: no type is given, hence this name is invalidly published.] 

Encyonema ochridanum Krammer, Biblioth. Diatomol. 36, p. 159, pl. 31: figs 3-10 (1997). 

Encyonema oestrupii Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 68, 191, pl. 192: figs 17-18 (1997). 

Encyonema pankowii Lange-Bert. et Krammer, Biblioth. Diatomol. 37, p. 69, 192, pl. 142: figs 9-11 (1997). 

Encyonema parvum Krammer, Biblioth. Diatomol. 37, p. 183, pl. 191: figs 10-15 (1997). 

Encyonema perelginense Krammer, Biblioth. Diatomol. 36, p. 173, pl. 61: figs 1-4 (1997). 

Encyonema pergracile Krammer, Biblioth. Diatomol. 36, p. 178, pl. 88: figs 1-8 (1997). 

Encyonema perminutum Krammer, Biblioth. Diatomol. 36, p. 161, pl. 25: figs 20-27 (1997). 

Encyonema perpusillum var. chilense Krammer, Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 115-116, pl. 111: figs 

20-23 (2000). 

Encyonema persilesiacum Krammer var. persilesiacum, Biblioth. Diatomol. 36, p. 165, pl. 10: figs 1-8 (1997). 

Encyonema persilesiacum var. ata-tau Krammer, Biblioth. Diatomol. 36, p. 165-166, pl. 12: figs 1-5 (1997). 

Encyonema polystigmoideum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 36, pl. 131: figs 8-11, pl. 144: figs 1-2 (1998). 

Encyonema ponteanum Krammer, Biblioth. Diatomol. 37, p. 192, pl. 142: figs 13-15 (1997). 

Encyonema procerum Krammer, Biblioth. Diatomol. 36, p. 169, pl. 32: figs 9-19 (1997). 

Encyonema pseudomuelleri Krammer, Biblioth. Diatomol. 36, p. 174, pl. 43: figs 1-6, pl. 44: figs 1-4, pl. 45: figs 1-4, 

(1997). [Comment: Encyonema pseudomuelleri was the intended new name and status for Cymbella muelleri var. javanica 

and Cymbella grossestriata var. javanica. Krammer published a new species with a Latin description, based on the lectotype, 

selected by Simonsen, Atlas and catalogue of the diatom types of Friedrich Hustedt 1, p. 130 (1987) for Cymbella 

grossestriata var. javanica (erroneously cited for Cymbella muelleri var. javanica).] [Homotypic synonym: Cymbella 

muelleri var. javanica Hust., Arch. Hydrobiol., Suppl., p. 425-426, pl. 26: figs. 1-4. Comment: Cymbella muelleri var. 

javanica was originally published as a new combination for Cymbella grossestriata var. javanica. Since Cymbella 

grossestriata var. javanica was invalidly published, Cymbella muelleri var. javanica is treated as a new described variety.] 

[- Cymbella grossestriata var. javanica Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 373: figs 8-9 

(1931). Comment: Cymbella grossestriata var. javanica is lacking a diagnosis, hence it is invalidly published, the term 

"Nähert sich Cymbella turgida Greg." can not be treated as a diagnosis.] 

Encyonema rifelanum Krammer, Biblioth. Diatomol. 37, p. 193, pl. 134: figs 7-14 (1997). 

Encyonema riotecense Krammer, Biblioth. Diatomol. 36, p. 169, pl. 41: figs 1-6 (1997). 

Encyonema rostratum Krammer, Biblioth. Diatomol. 37, p. 190-191, pl. 126: figs 1-14 (1997). 

Encyonema rugosiforme Krammer, Biblioth. Diatomol. 36, p. 175-176, pl. 63: figs 1-3 (1997). 



29 

Encyonema rumrichae Krammer, Biblioth. Diatomol. 37, p. 184, pl. 102: figs 9-17 (1997). 

Encyonema schimanskii Krammer, Biblioth. Diatomol. 36, p. 54, 158, pl. 14: figs 10-17 (1997). [Comment: Encyonema 

schimanskii on p. 208 (= caption of pl. 14: figs 10-17) is cited as "Encyonema minutum (Hilse) D.G.Mann".] 

Encyonema schneideri Krammer, Biblioth. Diatomol. 36, p. 166, pl. 40: figs 1-4 (1997). 

Encyonema schoemanii Krammer, Biblioth. Diatomol. 37, p. 183, pl. 136: figs 18-19 (1997). 

Encyonema semilanceolatum Krammer, Biblioth. Diatomol. 36, p. 172, pl. 33: figs 14-20 (1997). 

Encyonema sibericum Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 44, pl. 59: figs 4-6, pl. 60: figs 5-7 (1999). 

[Comment: Diploneis ovalis ssp. arctica is not documented on pl. 59: figs 4-5, pl. 60: figs 4-6 as cited in the 1 st edition, 

reference corrected in the 2 nd edition.] 

Encyonema silesiacum var. altensis Krammer, Biblioth. Diatomol. 36, p. 163, pl. 9: figs 9-19 (1997). [Comment: on p. 163 

pl. 9: figs 7-8 is erroneously cited. On pl. 9: figs 7-8 Encyonema silesiacum var. silesiacum is documented.] 

Encyonema silesiacum var. distigmata Krammer, Biblioth. Diatomol. 36, p. 164, pl. 8: fig. 8 (1997). 

Encyonema silesiacum var. distinctepunctata Krammer, Biblioth. Diatomol. 36, p. 164, pl. 11: figs 1-11 (1997). 

Encyonema silesiacum var. elegans Krammer, Biblioth. Diatomol. 36, p. 164, pl. 14: figs 6-9 (1997). 

Encyonema silesiacum var. excisa Krammer, Biblioth. Diatomol. 36, p. 164, pl. 11: figs 13-14 (1997). 

Encyonema silesiacum var. lata Krammer, Biblioth. Diatomol. 36, p. 164, pl. 12: figs 6-9 (1997). 

Encyonema silesiacum var. latarea Krammer, Biblioth. Diatomol. 36, p. 165, pl. 13: figs 9-12 (1997). 

Encyonema silesiacum var. latistriata Krammer, Biblioth. Diatomol. 36, p. 165, pl. 11: fig. 12 (1997). 

Encyonema silesiacum var. ventriformis Krammer, Biblioth. Diatomol. 36, p. 164, pl. 8: figs 9-15 (1997). 

Encyonema simile Krammer, Biblioth. Diatomol. 36, p. 160, pl. 18: figs 5-10 (1997). 

Encyonema sinicum Krammer, Biblioth. Diatomol. 36, p. 175, pl. 72: figs 1-9 (1997). 

Encyonema sparsipunctatum Krammer, Biblioth. Diatomol. 36, p. 179, pl. 95: figs 1-5 (1997). 

Encyonema spitsbergense Krammer, Biblioth. Diatomol. 36, p. 163, pl. 15: figs 1-4 (1997). 

Encyonema stigmoideum Krammer, Biblioth. Diatomol. 36, p. 163, pl. 15: figs 5-14 (1997). 

Encyonema subbrevistriatum Krammer, Biblioth. Diatomol. 37, p. 188-189, pl. 132: figs 17-23 (1997). 

Encyonema subelginense Krammer, Biblioth. Diatomol. 36, p. 173, pl. 62: figs 2-8 (1997). 

Encyonema subgracile Krammer, Biblioth. Diatomol. 36, p. 178-179, pl. 95: figs 12-19 (1997). [Comment: on p. 370 (= 

caption of pl. 95 the authors are cited as "Krammer et Lange-Bert.".] 

Encyonema subjavanicum Krammer, Biblioth. Diatomol. 37, p. 194, pl. 133: figs 13-20 (1997). 

Encyonema subkukenanum Krammer, Biblioth. Diatomol. 37, p. 185, pl. 106: figs 10-15 (1997). 

Encyonema sublanceolatum Krammer, Metzeltin et Lange-Bert., Biblioth. Diatomol. 37, p. 14, 183, pl. 136: fig. 16, pl. 

202: figs 13-15 (1997). 

Encyonema subminutum Krammer et Lange-Bert., Biblioth. Diatomol. 36, p. 58, 159, pl. 19: figs 17-24 (1997). [Comment: 

in the Latin diagnosis the author name "Lange-Bert." is missing. On p. 58 (German description) pl. 19: fig. 25, pl. 25: figs 

28-34 are also mentioned.] 

Encyonema subnorvegicum Krammer, Biblioth. Diatomol. 37, p. 186, pl. 112: figs 1-6 (1997). 

Encyonema subobscurum Krammer, Biblioth. Diatomol. 36, p. 171-172, pl. 27: figs 18-20 (1997). 

Encyonema subperpusillum Krammer, Biblioth. Diatomol. 37, p. 186-187, pl. 112: figs 12-14 (1997). 

Encyonema subrostratum Krammer, Biblioth. Diatomol. 37, p. 192-193, pl. 131: figs 1-10 (1997). 

Encyonema subtenerum Lange-Bert. et Krammer, Biblioth. Diatomol. 38, p. 126, pl. 48: figs 10-16 (1998). 

Encyonema subtenuissimum Krammer, Biblioth. Diatomol. 37, p. 182, pl. 128: figs 5-5a (1997). 



30 

Encyonema subtriste Krammer, Biblioth. Diatomol. 37, p. 188, pl. 131: figs 17-18 (1997). 

Encyonema supergracile Krammer et Lange-Bert., Biblioth. Diatomol. 36, p. 147, 178, pl. 89: figs 4-9 (1997). 

Encyonema tapajoz Krammer, Biblioth. Diatomol. 37, p. 188, pl. 130: figs 1-7 (1997). 

Encyonema tasmaniense Krammer, Biblioth. Diatomol. 37, p. 187, pl. 120: figs 1-5 (1997). 

Encyonema tenerum Krammer, Biblioth. Diatomol. 37, p. 182, pl. 151: fig. 10, pl. 192: figs 20-22 (1997). 

Encyonema tenuicephalum Lange-Bert. et Krammer, Biblioth. Diatomol. 38, p. 128, pl. 49: figs 6-8 (1998). 

Encyonema thermale Krammer, Biblioth. Diatomol. 37, p. 185-186, pl. 108: figs 7-10 (1997). 

Encyonema thioense Lange-Bert. et Krammer, Biblioth. Diatomol. 38, p. 129-130, pl. 49: figs 9-14 (1998). 

Encyonema trianguliforme Krammer, Biblioth. Diatomol. 36, p. 176, pl. 73: figs 1-5, pl. 74: figs 1-8 (1997). 

Encyonema tropicum Metzeltin et Krammer, Biblioth. Diatomol. 37, p. 195, pl. 203: figs 1-3 (1997). 

Encyonema truncatum Krammer, Biblioth. Diatomol. 37, p. 193, pl. 134: figs 15-18 (1997). 

Encyonema vanoyei (Cholnoky) Krammer var. vanoyei, Nova Hedwigia 65, p. 139 (1997). 

Basionym: Cymbella vanoyei Cholnoky, Oesterr. Bot. Z. 101, p. 412, figs 17-18 (1954). 

Encyonema vanoyei var. grande Krammer, Biblioth. Diatomol. 36, p. 177, pl. 51: figs 1-2 (1997). [Comment: on p. 138 and 

282 var. grande is cited as "var. grandis".] 

Encyonema venezolanum Krammer var. venezolanum, Biblioth. Diatomol. 36, p. 162, pl. 19: figs 1-8 (1997). 

Encyonema venezolanum var. similis Krammer, Biblioth. Diatomol. 36, p. 162, pl. 20: figs 6-10 (1997). 

Encyonema ventricosum var. angusta Krammer, Biblioth. Diatomol. 36, p. 170, pl. 6: figs 5-7 (1997). 

Encyonema vulgare Krammer var. vulgare, Biblioth. Diatomol. 36, p. 167, pl. 36: figs 4-10 (1997). 

Encyonema vulgare var. chilensis Krammer, Biblioth. Diatomol. 36, p. 168, pl. 38: figs 4-5 (1997). 

Encyonema vulgare var. tasmanensis Krammer, Biblioth. Diatomol. 36, p. 167, pl. 50: figs 1-2 (1997). [Comment: on p. 

280 var. tasmanensis is cited as "var. tasmaniensis ".] 

Encyonema wallaceanum var. neocaledonicum Krammer et Lange-Bert., Biblioth. Diatomol. 38, p. 130-131, pl. 48: fig. 26 

(1998). 

Encyonema wojtalae Metzeltin, Iconogr. Diatomol. 5, p. 37, pl. 136: figs 14-19, pl. 143: figs 3-4 (1998). 

Encyonema yellowstonianum Krammer, Biblioth. Diatomol. 36, p. 174-175, pl. 64: figs 4-6 (1997). 

Encyonema yucatanense Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 38, pl. 139: figs 15-17, 23-25 (1998). 

ENCYONOPSIS Krammer, Biblioth. Diatomol. 36, p. 156-157, (1997). [Typus generis: Navicula cesatii Rabenh., Süßwasserdiatomeen, 

p. 39, pl. 6: fig. 89 (1853).]. 

Encyonopsis alpina Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 97-98, 196, pl. 150: figs 4, 7-10, pl. 193: figs 1 

[only valve on the left hand side], 5, 7-9 (1997). 

Encyonopsis angusta Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 98-99, 196, pl. 148: figs 5-6, pl. 150: figs 2-3, 5-6, 

pl. 193: figs 2-3 (1997). 

Encyonopsis blanchensis Krammer, Biblioth. Diatomol. 37, p. 205, pl. 138: figs 1-3 (1997). 

Encyonopsis buedeliformis Krammer, Biblioth. Diatomol. 37, p. 205, pl. 181: figs 1-7 (1997). 

Encyonopsis buedelii Krammer, Biblioth. Diatomol. 37, p. 205, pl. 170: figs 4-11 (1997). 

Encyonopsis cesatiformis Krammer, Biblioth. Diatomol. 37, p. 206, pl. 178: figs 7-12 (1997). 

Encyonopsis cesatii var. geitleri Krammer, Biblioth. Diatomol. 37, p. 206, pl. 184: figs 4-13 (1997). 

Encyonopsis costaricana Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 7-8, pl. 17: figs 19-21 (2001). 

Encyonopsis degenerata Gerd Moser, Biblioth. Diatomol. 43, p. 125-126, pl. 1: figs 1-11 (1999). 

Encyonopsis descripta var. asymmetrica Krammer, Biblioth. Diatomol. 37, p. 200, pl. 156: figs 3-8 (1997). 



31 

Encyonopsis difficiliformis Krammer, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 37, p. 122-123, 200, pl. 162: figs 25-26, 

pl. 202: figs 7-12a (1997). 

Encyonopsis dubitabilis Lange-Bert., Gerd Moser et Krammer, Biblioth. Diatomol. 37, p. 143, 204, pl. 140: figs 14-16 

(1997). Comment: Valitated name for Encyonopsis dubitabilis Lange-Bert. et Gerd Moser, nom. inval., Moser et al., 

Biblioth. Diatomol. 32, p. 71, pl. 24: figs 9-14 (1995). In Metzeltin & Kusber (2001) the authors are erroneously cited as 

"Lange-Bert., Krammer et Gerd Moser".] 

Encyonopsis eifelana Krammer, Biblioth. Diatomol. 37, p. 197-198, pl. 150: figs 31-36, pl. 203: figs 7-8 (1997). 

Encyonopsis falaisensiformis Krammer, Biblioth. Diatomol. 37, p. 199-200, pl. 181: figs 8-13 (1997). 

Encyonopsis floridana Krammer, Biblioth. Diatomol. 37, p. 203, pl. 167: figs 1-6 (1997). 

Encyonopsis fragilis Krammer, Biblioth. Diatomol. 37, p. 204-205, pl. 169: figs 10-12 (1997). 

Encyonopsis frequentiformis Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 39, pl. 137: figs 13-14, 16 (1998). [Comment: 

on page 39 erroneously figs 11-12 are referred to.] 

Encyonopsis frequentis Krammer var. frequentis, Biblioth. Diatomol. 37, p. 201-202, pl. 157: figs 10-11, pl. 158: figs 1-3, 

pl. 202: figs 1-6 (1997). 

Encyonopsis frequentis var. brasiliana Krammer, Biblioth. Diatomol. 37, p. 202, pl. 186: figs 2-9 (1997). 

Encyonopsis frequentis var. fossilis Krammer, Biblioth. Diatomol. 37, p. 202, pl. 166: fig. 3 (1997). 

Encyonopsis glacialis Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 116, pl. 116: figs 11-12 (2000). 

[Comment: on p. 462 (= caption of pl. 111) the authors are cited as "Krammer, Rumrich et Lange-Bert.".] 

Encyonopsis gomphonemiformis Krammer, Biblioth. Diatomol. 37, p. 203, pl. 183: figs 4-6 (1997). 

Encyonopsis grunowii Krammer, Biblioth. Diatomol. 37, p. 207-208, pl. 178: figs 2-6 (1997). 

Encyonopsis kergueliformis Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 151, 205-206, pl. 189: figs 14-20 (1997). 

Encyonopsis krammerioides Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 117, pl. 116: figs 3-5 (2000). 

[Comment: Encyonopsis krammerioides in Metzeltin & Kusber (2001) was erroneously cited as "Encyonopsis 

krammeroides".] 

Encyonopsis krasskei Krammer, Biblioth. Diatomol. 37, p. 207, pl. 174: figs 5-8 (1997). 

Encyonopsis kriegeri var. fossilis Krammer, Biblioth. Diatomol. 37, p. 203, pl. 190: figs 8-10 (1997). 

Encyonopsis lanceo-elliptica Krammer, Biblioth. Diatomol. 37, p. 207, pl. 189: figs 1-8 (1997). 

Encyonopsis linearis Krammer, Biblioth. Diatomol. 37, p. 201, pl. 158: figs 4-9 (1997). 

Encyonopsis minuta Krammer et E.Reichardt, Biblioth. Diatomol. 37, p. 195, pl. 143a: figs 1-27 (1997). 

Encyonopsis moseri Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 104, 198, pl. 152: figs 14-18, pl. 153: figs 1-11 

(1997). 

Encyonopsis namibiana Krammer, Biblioth. Diatomol. 37, p. 206, pl. 181: figs 14-20 (1997). 

Encyonopsis neoamphioxis Krammer, Biblioth. Diatomol. 37, p. 141-142, 203, pl. 168: figs 1-9, 11-13 (1997). 

Encyonopsis opima (Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 35 (1998). 

Replaced synonym: Gomphonema opimum Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 211 (1996). 

Encyonopsis orientalis Krammer, Biblioth. Diatomol. 37, p. 199, pl. 153: figs 13-24 (1997). 

Encyonopsis patagonica Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 118, pl. 117: figs 21-22 (2000). 

Encyonopsis perborealis Krammer, Biblioth. Diatomol. 37, p. 198-199, pl. 154: figs 1-12 (1997). 

Encyonopsis radialis Krammer, Biblioth. Diatomol. 37, p. 200, pl. 164: figs 14-21, pl. 175: figs 6-7 (1997). 

Encyonopsis recta Krammer, Biblioth. Diatomol. 37, p. 202-203, pl. 165: figs 12-14 (1997). 

Encyonopsis reichardtii Krammer, Biblioth. Diatomol. 37, p. 202, pl. 166: figs 1-2 (1997). 

Encyonopsis reviersiana Lange-Bert., Gerd Moser et Krammer, Biblioth. Diatomol. 37, p. 126, 201, pl. 158: figs 13-15 

(1997). 



32 

[Comment: validated name for Cymbella reviersiana Lange-Bert. et Gerd Moser, nom. inval., Biblioth. Diatomol. 32, p. 81, 

pl. 20: figs 9-12. (1995). On p. 366 (= caption of pl. 158) Encyonopsis reviersiana is cited as "Encyonopsis reviersianum".] 

Encyonopsis rostrata Krammer, Biblioth. Diatomol. 37, p. 197, pl. 150: figs 24-30 (1997). 

Encyonopsis rumrichae Krammer, Biblioth. Diatomol. 37, p. 197, pl. 148: figs 10-24 (1997). 

Encyonopsis schneideri Krammer, Biblioth. Diatomol. 37, p. 207, pl. 176: figs 1-10, pl. 201: figs 1-3 (1997). 

Encyonopsis speciosa Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 119, pl. 117: figs 5-7 (2000). 

Encyonopsis spiculiformis Krammer, Biblioth. Diatomol. 37, p. 204, pl. 171: figs 6-9 (1997). 

Encyonopsis subcapitata Krammer, Biblioth. Diatomol. 37, p. 201, pl. 180: figs 1-4 (1997). 

Encyonopsis subfonticola Krammer, Biblioth. Diatomol. 37, p. 199, pl. 152: figs 7-13 (1997). 

Encyonopsis subminuta Krammer et E.Reichardt, Biblioth. Diatomol. 37, p. 195-196, pl. 144: figs 6-11 (1997). 

Encyonopsis subruttneri Krammer, Biblioth. Diatomol. 37, p. 198, pl. 151: figs 12-17 (1997). 

Encyonopsis subspicula Krammer, Biblioth. Diatomol. 37, p. 204, pl. 168: figs 8-9 (1997). 

Encyonopsis substodderi Krammer, Biblioth. Diatomol. 37, p. 206-207, pl. 172: figs 5-8 (1997). 

[Comment: the holotype erroneously is cited as "Lectotypus".] 

Encyonopsis symmetrica Krammer, Biblioth. Diatomol. 37, p. 208, pl. 179: figs 1-4 (1997). 

Encyonopsis tavirana Krammer, Biblioth. Diatomol. 37, p. 196, pl. 148: figs 1-3 (1997). 

Encyonopsis thumensis Krammer, Biblioth. Diatomol. 37, p. 198, pl. 154: figs 13-15, pl. 158: figs 16-22 (1997). 

Encyonopsis tiroliana Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 102, 197, pl. 203,: figs 10-12 (1997). 

[Comment: Encyonopsis tiroliana was already documented in Lange-Bertalot & Metzeltin, Iconogr. Diatomol. 2 (1996) pl. 

91: figs 18-20 as "Cymbella nov. spec. Nr. 10 Mittersee".] 

Encyonopsis vandamii Krammer et Lange-Bert., Biblioth. Diatomol. 37, p. 109-110, 199, pl. 151: figs 21-27, pl. 187: figs 9- 

11 (1997). 

ENTOMONEIS (Ehrenb.) Ehrenb. 


Entomoneis corrugata (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 198 (2000). 

Basionym: Amphiprora corrugata Giffen, Nova Hedwigia 13, p. 213, figs 6-7 (1967). 

Entomoneis densistriata (Krasske) Lange-Bert., Willmann et Metzeltin, Iconogr. Diatomol. 3, p. 38 (1996). 

Basionym: Amphiprora paludosa var. densestriata Krasske, Beih. Großbot. Zentralbl. 60, p. 342, pl. 13: fig. 1 (1940). 

Entomoneis kjellmanii var. striolata (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 198 (2000). 

Basionym: Amphiprora striolata Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 62, pl. 4: fig. 81 (1880). 

EOLIMNA Lange-Bert. et W.Schiller, Paläontol. Z. 71, p. 166, 168 (1997). [Typus generis: Eolimna martinii W.Schiller et 

Lange-Bert., Paläontol. Z. 71, p. 166, 168 (1997). Comment: in Moser et al. (1998), Biblioth. Diatomol. 38, p. 151 an 

additional Latin diagnosis is given.] 

Eolimna adnata (Hust.) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 152 (1998). 

Basionym: Navicula adnata Hust., Arch. Hydrobiol. Suppl. 15, p. 264 (1937). 

[– Navicula adnata Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde pl. 401: figs 37-40 (1936).] 

Eolimna inconspicua Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 152-153, pl. 23: figs 1-5 (1998). 

Eolimna martinii W.Schiller et Lange-Bert., Paläontol. Z. 71, p. 168, figs 2, 6-12 (1997). [The fossil Eolimna martinii is the 

type of the name of the genus EOLIMNA Lange-Bert. et W.Schiller. Comment: in Moser et al. (1998), Biblioth. Diatomol. 

38, p. 151 an additional Latin diagnosis is given.] 

Eolimna minima (Grunow) Lange-Bert., Biblioth. Diatomol. 38, p. 153 (1998). 

Basionym: Navicula minima Grunow in Van Heurck, Synopsis des Diatomées de Belgique, p. 107, pl. 14: fig. 15 (1880- 

1885). 

Eolimna neocaledonica (Manguin ex Kociolek et Reviers) Lange-Bert., Biblioth. Diatomol. 38, p. 35 (1998). 

Basionym: Navicula neocaledonica Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 203 (1996). 



33 

Eolimna rhombelliptica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 155, pl. 24: figs 1-9 (1998). 

Eolimna rhombica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 156, pl. 23: figs 11-20 (1998). 

Eolimna subadnata (Hust.) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 154 (1998). 

Basionym: Navicula subadnata Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 42, p. 69, figs 6-8 (1942). 

Eolimna subminuscula (Manguin) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 154 (1998). 

Basionym: Navicula subminuscula Manguin, Rev. Algol. 12, p. 139, pl. 2: fig. 39 (1941). 

Eolimna thioense Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 157, pl. 23: figs 6-10 (1998). 

Eolimna verecundaeformis (Manguin ex Kociolek et Reviers) Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 632 

(2001). 

Basionym: Navicula verecundaeformis Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 204, (1996). 

[– Navicula verecundaeformis Manguin, nom. inval., Mém. Mus. Natl. Hist. Nat., Nouv. Sér. B, Bot. 12 (1), p. 22, pl. 2: figs 

21a, b (1962).] 

[– Eolimna verecundaeformis (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, nom. inval., Biblioth. 

Diatomol. 38, p. 35, 52 (1998).] 

Eolimna zalokariae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 39-40, pl. 167: figs 1-2 (1998). 

EPHEMERA Paddock 


[http://128.32.109.44/cgi-bin/porp_cgi.plEPHEMERATBBPADDOCK1988]. 

Ephemera bjoernoeyaensis Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 14, pl. 55: figs 10-16, pl. 78: figs 

1-4 (1996). [Comment: Ephemera bjoernoeyaensis on p. 142 (= caption of pl. 55: fig. 16) is cited as"Proshkinia cf. 

bulnheimii (Grunow) Karayeva".] 

EPITHEMIA Kütz. 


Epithemia gracilis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 158, pl. 65: figs 1-9 (1998). 

Epithemia illuminata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 159, pl. 61: figs 1-4 (1998). 

Epithemia krammeri Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 160-161, pl. 66: figs 1-7 (1998). 

Epithemia miranda Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 163, pl. 62: figs 1-7, pl. 63: figs 1-6, 

pl. 64: figs 1-6 (1998). 

EUCOCCONEIS Cleve ex F.Meister 


Eucocconeis alpestris (Brun) Lange-Bert., Iconogr. Diatomol. 6, p. 45 (1999). 

Basionym: Achnanthes flexella var. alpestris Brun, Diatomées des Alpes et du Jura de la region Suisse et Française des 

environs de Genève, p. 30, fig. 26 (1880). 

[≡ Achnanthes alpestris (Brun) Lange-Bert. et Metzeltin.] 

Eucocconeis aretasii (Manguin) Lange-Bert., Iconogr. Diatomol. 6, p. 45 (1999). 

Basionym: Achnanthes aretasii Manguin, Mém. Inst. Sci. Madagaskar (Ser. B) 5, p. 19, pl. 2: fig. 16 a-c (1954). 

Eucocconeis austriaca (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 46 (1999). 

Basionym: Achnanthes austriaca Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 10, p. 75, figs 6-7 (1922). 

Eucocconeis depressa (Cleve) Lange-Bert., Iconogr. Diatomol. 6, p. 46 (1999). 

Basionym: Navicula depressa Cleve, Acta Soc. Fauna Fl. Fenn. 8 (2), p. 35, pl. 2: fig. 4 (1891). 

Eucocconeis diluviana (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 46 (1999). 

Basionym: Achnanthes diluviana Hust., Abh. Naturwiss. Vereine Bremen 34, p. 451, figs 11-12 (1954). 

Eucocconeis laevis (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 46 (1999). 

Basionym: Achnanthes laevis Østrup, Danske Diatomeer, p. 130, pl. 3: fig. 80 (1910). 

Eucocconeis leptostriata Lange-Bert., Iconogr. Diatomol. 6, p. 46, pl. 7: figs 4-5, pl. 8: figs 1-6 (1999). 

[Comment: the genus on p. 45 is erroneously not cited as Eucocconeis Cleve ex F.Meister, see Fourtanier & Kociolek 

(1999), p. 62.] 



34 

Eucocconeis ninckei (Guermeur et Manguin) Lange-Bert., Iconogr. Diatomol. 6, p. 48 (1999). 

Basionym: Achnanthes ninckei Guermeur et Manguin, Oesterr. Bot. Z. 100, p. 514, pl. 1: figs 10-11 (1953). 

Eucocconeis quadratarea (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 48 (1999). 

Basionym: Cocconeis quadratarea Østrup, Danske Diatomeer, p. 124, pl. 3: fig. 77 (1910). 

EUNOTIA Ehrenb. 


EUNOTIA subgen. CULTRIA [see CULTRIA]. 

Eunotia acicularis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 47-48, pl. 61: figs 8-9 (1998). [Comment: Eunotia 

acicularis on page 356 (= caption of pl. 61: figs 8-9) is cited as "Eunotia spec. Nr. 61/8, 9".] 

Eunotia actuariola Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 48-49, pl. 44: figs 2-8 (1998). [Comment: Eunotia 

actuariola on page 322 (= caption of pl. 44: figs 2-8) is misspelled as "E. acutuariola".] 

Eunotia acus Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 49, pl. 13: figs 8, 9A-C (1998). [Comment: Eunotia acus is 

not documented on pl. 12 as cited on p. 49.] 

Eunotia acutinasuta Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 50, pl. 59: figs 14-17 (1998). 

Eunotia anamargaritae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 51 (1998). 

Replaced synonym: Eunotia zygodon var. elongata Hust. ex Simonsen, Atlas and Catalogue of the Diatom Types of F. 

Hustedt 1, p. 37 (1987). 

Eunotia andinofrequens Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 120, pl. 18: figs 18-24 (2000). 

Eunotia arcubus Nörpel et Lange-Bert., Biblioth. Diatomol. 27, p. 24-25, pl. 16: figs 1-6, pl. 19: figs 1-10, pl. 20: figs 1-6, 

pl. 21: figs 1-11 (1993). 

Eunotia arculus Lange-Bert. et Nörpel, Biblioth. Diatomol. 27, p. 26 (1993). [Validation of Eunotia arculus (Grunow) 

Lange-Bert. et Nörpel, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 

(3), p. 213, pl. 157: figs 4-12 (1991).] [- Eunotia paludosa var. arculus Grunow, nom. inval., in Van Heurck, Synopsis des 

Diatomées de Belgique, Type de Synopsis, no. 274 (1880-1887).] 

Eunotia australominor Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 164, pl. 10: figs 8-13 (1998). 

Eunotia bicornigera Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 51, pl. 28: figs 17-24, 28 (1998). 

Eunotia bilunaris var. linearis (Okuno) Lange-Bert. et Nörpel-Schempp in Jahn et al., Lange-Bertalot-Festschrift, p. 633 

(2001). 

Basionym: Eunotia flexuosa var. linearis Okuno, Atlas of fossil diatoms from Japanese Diatomite deposits. Kawakita 

Printing Co., Kyoto, p. 31, pl. 23: fig. 18 (1952). 

[– Eunotia bilunaris var. linearis (Okuno) Lange-Bert. et Nörpel, nom. inval., Süßwasserflora von Mitteleuropa 2 (3), p. 180, 

pl. 137: figs 13-16 (1991); Biblioth. Diatomol. 27, p. 28 (1993).] 

Eunotia bilunaris var. mucophila Lange-Bert., Nörpel et Alles, Nova Hedwigia 53, p. 196, pl. 5: figs 12-21 (1991). 

Eunotia boreoalpina Lange-Bert. et Nörpel-Schempp, Iconogr. Diatomol. 5, p. 52 (1998). [As a part of the protologue 

Krammer & Lange-Bertalot (1991): Süßwasserflora von Mitteleuropa 2 (3), pl. 163: figs 1-7 is cited. Comment: in Krammer 

& Lange-Bertalot (1991a), p. 222 and p. 163 (= caption of pl. 163: figs 1-7) Eunotia boreoalpina is cited as "Eunotia incisa 

-Sippe".] 

Eunotia boreotenuis Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 46, pl. 9: figs 21-23 (1996). 

Eunotia botellus Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 165-166, pl. 15: figs 1-4 (1998). 

Eunotia botuliformis F. Wild, Nörpel et Lange-Bert., Biblioth. Diatomol. 27, p. 29, pl. 33: figs 2-15 (1993). 

Eunotia brinkmannii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 52-53, pl. 39: figs 4-6 (1998). [Comment: Eunotia 

brinkmannii on page 312 (caption of pl. 39: figs 4-6) is misspelled as "E. brinckmannii".] 

Eunotia cameliopsis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 53-54, pl. 29: figs 13 (14) (1998). [Comment: 

Eunotia cameliopsis is not documented on "pl. 21" as cited on p. 53.] 

Eunotia chelonia Nörpel-Schempp, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 47, pl. 13: figs 14-17 (1996). 

[Comment: on p. 144 (caption of pl. 13) the authors are cited as "Nörpel-Schempp, Metzeltin et Lange-Bert.".] 



35 

Eunotia circumborealis Lange-Bert. et Nörpel, Biblioth. Diatomol. 27, p. 30 (1993). [Validation of Eunotia circumborealis 

Lange-Bert. et Nörpel, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 

(3), p. 197, pl. 143: figs 16-23 (1991).] 

Eunotia coccinea Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 54, pl. 12: figs 12-15 (1998). 

Eunotia coringii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 55 (1998). 

Replaced synonym: Eunotia didyma var. media Hust. ex Simonsen, Atlas and Catalogue of the Diatom Types of F. Hustedt 

1, p. 30 (1987). 

Eunotia costaricana Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 8, pl. 2: figs 1-4 (2001). 

Eunotia crassispinata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 166-167, pl. 17: figs 1-4 (1998). 

Eunotia crassula Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 55-56, pl. 59: figs 11-13 (1998). 

Eunotia curtagrunowii Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 48 (1996). 

Replaced synonym: Eunotia praerupta var. curta Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 34: fig. 

24 (1881). 

Eunotia dacostae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 56, pl. 50: figs 5-6 (1998). 

Eunotia damasii ssp. melanesica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 167-168, pl. 16: figs 9- 

11 (1998). 

Eunotia deformis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 57, pl. 16: figs 9-11 (1998). 

Eunotia disproportionata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 58, pl. 11: figs 7-8, pl. 13: figs 2-5 (1998). 

Eunotia distinguenda Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 58-59, pl. 15: figs 10-15, pl. 46: fig. 1 (1998). 

Eunotia elongata (Grunow) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 59 (1998). 

Basionym: Eunotia formica var. elongata Grunow in Rabenhorst, Beiträge (Leipzig), 1, p. 3, pl. 1: figs 2a, b (1865). 

Eunotia eurycephala (Grunow) Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 48 (1996). 

Basionym: Eunotia flexuosa var. eurycephala Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 35: 

fig. 8 (1881). 

Eunotia eurycephaloides Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 48-49, pl. 9: figs 18-20, pl. 112: figs 2-3 

(1996). 

Eunotia exsecta (Cleve-Euler) Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 50 (1996). 

Basionym: Eunotia veneris var. exsecta Cleve-Euler, Kongl. Svenska Vetenskapsakad. Handl. 2 (2), p. 113, figs 450 c-e 

(1953). 

Eunotia falcifera Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 60-61, pl. 12: figs 19-26 (1998). [Eunotia falcifera is 

the type of the name of the subgenus CULTRIA Metzeltin et Lange-Bert.] 

Eunotia filum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 168-169, pl. 15: figs 5-8 (1998). 

Eunotia fukushimae Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 169-170, pl. 12: figs 1-2 (1998). 

Eunotia genufalcata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 170-171, pl. 15: figs 9-10 (1998). 

Eunotia genuflexa Nörpel-Schempp, Iconogr. Diatomol. 2, p. 50, pl. 9: figs 14-17 (1996). 

Eunotia georgii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 61, pl. 41: figs 1-7, pl. 42: figs 7-8 (1998). 

Eunotia glacialifalsa Lange-Bert. in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 3, 2, 

2. edition, p. 594-595, pl. 151: fig. 6, pl. 151A: figs 3-6 (2000). [Comment: Eunotia glacialifalsa on p. 532 (= caption of pl. 

151) is cited as "Eunotia glacialis Meister sensu stricto = Eunotia gracilis Ehrenberg sensu auct. nonnull.".] 

Eunotia gracillima (Krasske) Nörpel-Schempp, Iconogr. Diatomol. 3, p. 70 (1996). 

Basionym: Eunotia fallax var. gracillima Krasske, Bot. Arch. 27, p. 349, figs 2 a-c (1929). 

Eunotia hepaticola Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 8-9, pl. 1: figs 1-9 (2001). 

Eunotia incisatula Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 62, pl. 59: figs 25-30 (1998). 



36 

Eunotia indigenarum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 63 (1998). 

Replaced synonym: Eunotia zygodon var. lata Hust. ex Simonsen, Atlas and Catalogue of the Diatom Types of F. Hustedt 1, 

p. 36 (1987). 

Eunotia inflata (Grunow) Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 52 (1996). 

Basionym: Eunotia praerupta var. inflata Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 34: fig. 17 

(1881). 

Eunotia inspectabilis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 63-64, pl. 55: figs 8-11 (1998). 

Eunotia kanakarum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 171-172, pl. 13: figs 1-4 (1998). 

Eunotia krammeri Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 64, pl. 53: figs 1-7, pl. 63: fig. 1 (1998). 

Eunotia lenis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 65, pl. 60: figs 10-16, pl. 62: figs 3, 4, 5 (1998). [Comment: 

Eunotia lenis is not documented on pl. 62: fig. 6 as cited on p 65 and 354, p. 358 (caption of pl. 62) is correct.] 

Eunotia limax Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 172-173, pl. 16: fig. 8 (1998). 

Eunotia longicollis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 66, pl. 15: figs 6-8 (1998). 

Eunotia maxima (Hust. ex Simonsen) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 66 (1998). 

Basionym: Eunotia didyma var. maxima Hust. ex Simonsen Atlas and Catalogue of the Diatom Types of F. Hustedt 1, p. 30 

(1987). 

Eunotia melanogaster Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 173-174, pl. 12: figs 3-8 (1998). 

Eunotia meridiana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 67, pl. 59: figs 7-10 (1998). 

Eunotia michaelis Metzeltin, Witkowski et Lange-Bert., Iconogr. Diatomol. 4, p. 14-15, pl. 34: figs 2-8, pl. 90: figs 5-6 

(1996). [Comment: Eunotia michaelis is not documented on pl. 33: figs 2-8 and pl. 90: fig. 4 as cited on p. 14]. 

Eunotia monodontiforma Lange-Bert. et Nörpel-Schempp, Iconogr. Diatomol. 9, p. 121-122, pl. 21: figs 1-7 (2000). 

[Comment: validation of Eunotia monodontiforma Lange-Bert. et Nörpel, nom. inval., in Biblioth. Diatomol. 27, p. 37, pl. 

28: figs 1-6, pl. 29: figs 4-6. (1993).] 

Eunotia muscicola var. perminuta (Grunow) Nörpel et Lange-Bert., Biblioth. Diatomol. 27, p. 38 (1993). 

Basionym: Eunotia tridentula var. perminuta Grunow in Cleve et Möller, Diatoms, collection of 324 slides, no. 186 (1879). 

Eunotia muscicola var. tridentula Nörpel et Lange-Bert., Biblioth. Diatomol. 27, p. 39 (1993). [Validation of Eunotia 

muscicola var. tridentula Nörpel et Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von 

Mitteleuropa, Bacillariophyceae 2 (3), p. 217, pl. 156: figs 12-22 (1991).] 

Eunotia napoleonica Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 68, pl. 49: figs 4-8 (1998). 

Eunotia neofallax Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 3, p. 73-74, pl. 58: figs 7-29 (1996). 

Eunotia neomundana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 69, pl. 19: figs 1-6 (1998). [Comment: on p. 272 (= 

caption of pl. 19) the newly described species is erroneously cited as "nov. stat.".] 

Eunotia noerpeliana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 70, pl. 59: figs 1-6 (1998). 

Eunotia novaecaledonica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 39 (1998). 

Replaced synonym: Eunotia monodon var. neocaledonica Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 197- 

198 (1996). 

Eunotia odebrechtiana Metzeltin et Lange-Bert. var. odebrechtiana, Iconogr. Diatomol. 5, p. 71, pl. 56: figs 1-6, 13-14 

(1998). 

Eunotia odebrechtiana var. essequiboensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 72, pl. 57: figs 1-2 (1998). 

Eunotia parapectinalis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 123, pl. 19: figs 4-8 (2000). 

Eunotia parasiolii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 72-73, pl. 60: figs 17-30 (1998). 

Eunotia pileus var. guianense (Ehrenb.) E.Reichardt, Iconogr. Diatomol. 1, p. 15 (1995). 

Basionym: Himantidium guianense Ehrenb., Abh. Königl. Akad. Wissensch. Berlin. 1841, p. 417, pl. 2/1: fig. 4a-c (1843). 

[Again marked as a new combination in Iconogr. Diatomol. 1 (second edition), p. 31 (1995).] 

Eunotia pirarucu Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 73, pl. 49: figs 1-3 (1998). 



37 

Eunotia ponspendens Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 174-175, pl. 14: figs 1-6 (1998). 

Eunotia praerupta var. papilio (Grunow) Nörpel, Biblioth. Diatomol. 27, p. 39 (1993). 

Basionym: Eunotia robusta var. papilio Grunow in Van Heurck, Synopsis, Atlas pl. 33: fig. 8 (1881). 

[Nomenclatural synonym of Eunotia pseudopapilio Lange-Bert. et Nörpel-Schempp.] 

[Comment: In Iconogr. Diatomol. 5 (1998), p. 74 Eunotia praerupta var. papilio (Grunow) Nörpel is cited as Eunotia 

praerupta var. papilio (Grunow) "Nörpel ex Lange-Bertalot", Biblioth. Diatomol. 27, p. 39 (1993).] 

Eunotia praestans Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 175-176, pl. 11: fig. 6, pl. 12: figs 9-11 

(1998). 

Eunotia pseudopapilio Lange-Bert. et Nörpel-Schempp, Iconogr. Diatomol. 5, p. 74 (1998). 

Replaced name: Eunotia robusta var. papilio Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 33: fig. 

8 (1881). 

[≡ Eunotia praerupta var. papilio (Grunow) Nörpel.] 

Eunotia rabenhorstiana var. elongata (R.M.Patrick) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 76 (1998). 

Basionym: Desmogonium rabenhorstianum var. elongatum R.M.Patrick, Notul. Nat. Acad. Nat. Sci. Philadelphia 59, p. 3, 

figs 1-3 (1940). 

Eunotia reichardtii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 76 (1998). 

Replaced synonym: Eunotia didyma var. recta Hust. ex Simonsen, Atlas and Catalogue of the Diatom Types of F. Hustedt 1, 

p. 30 (1987). 

Eunotia rhynchocephala var. satelles Nörpel et Lange-Bert., Biblioth. Diatomol. 27, p. 39-40 (1993). 

[Validation of Eunotia rhynchocephala var. satelles Nörpel et Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (3), p. 203, pl. 154: figs 18-22 (1991).] 

[Nomenclatural synonym of Eunotia satelles (Nörpel-Schempp et Lange-Bert.) Nörpel-Schempp et Lange-Bert.] 

Eunotia satelles (Nörpel-Schempp et Lange-Bert.) Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 53 (1996). 

Basionym: Eunotia rhynchocephala var. satelles Nörpel-Schempp et Lange-Bert., Biblioth. Diatomol. 27, p. 39 (1993). 

Eunotia schneideri Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 77, pl. 15: figs 1-2 (1998). 

Eunotia seminulum Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 2, p. 53, pl. 17: figs 43-47 (1996). 

Eunotia serra-australis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 78, pl. 42: figs 1-5 (1998). 

Eunotia silvae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 79, pl. 39: figs 7-10 (1998). 

Eunotia silvahercynia Nörpel, Van Sull et Lange-Bert., Nova Hedwigia 53, p. 208, pl. 6: figs 22-44 (1991). 

Eunotia sioliopsis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 177, pl. 10: figs 1-4 (1998). 

Eunotia soleri Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 80 (1998). 

Replaced synonym: Eunotia frickei var. elongata Hust. ex Simonsen, Atlas and Catalogue of the Diatom Types of F. Hustedt 

1, p. 38 (1987). 

Eunotia sparsiornata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 178, pl. 4: figs 7-11, pl. 17: figs 5-13 

(1998). 

Eunotia subarcuatoides Alles, Nörpel et Lange-Bert., Nova Hedwigia 53, p. 188, pl. 4: figs 1-36 (1991). 

Eunotia tambaquina Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 81 (1998). 

Replaced synonym: Eunotia rabenhorstii var. maxima Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 285: fig. 28 

(1913). 

Eunotia tapacumae var. sphagnophila (Krasske) Nörpel-Schempp, Iconogr. Diatomol. 3, p. 79 (1996). 

Basionym: Eunotia sphagnophila Krasske, Svensk Bot. Tidskr. 42, p. 425, pl. 1: figs 16-17 (1948). 

Eunotia tapacuminor Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 81, pl. 54: fig. 2 (1998). 

Eunotia tapacumopsis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 83, pl. 54: figs 6-8 (1998). 

Eunotia torganiae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 84, pl. 28: figs 25-27 (1998). 

Eunotia transfuga Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 84-85, pl. 8: fig. 5, pl. 9: figs 1-3 (1998). 

Eunotia ursamaioris Lange-Bert. et Nörpel-Schempp, Iconogr. Diatomol. 6, p. 48-49, pl. 4: figs 18-21 (1999). 



38 

Eunotia varioundulata Nörpel-Schempp et Lange-Bert., Iconogr. Diatomol. 3, p. 68 (1996). 

Replaced synonym: Eunotia exigua var. tridentula Østrup, Danske Diatomeer, p. 172, pl. 5: fig. 106 (1910). 

Eunotia ventriosa var. brevis (R.M.Patrick) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 86 (1998). 

Basionym: Eunotia brevis R.M.Patrick, Notul. Nat. Acad. Nat. Sci. Philadelphia 59, p. 4, fig. 8 (1940). 

Eunotia vitiosa Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 179, pl. 11: figs 1-5 (1998). 

Eunotia weisingii Lange-Bert., Biblioth. Diatomol. 27, p. 40, pl. 15: figs 1-11 (1993). 

Eunotia yanomami Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 86-87, pl. 34: figs 1-6, pl. 35: figs 1-6, pl. 36: figs 1- 

13, pl. 37: figs 7-8, pl. 45: fig. 6, pl. 63: fig. 2 (1998). 

Eunotia yoshitakeae Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 180, pl. 16: figs 3-7, (1-2) (1998). 

FALLACIA A.J.Stickle et D.G.Mann 


[http://128.32.109.44/cgi-bin/porp_cgi.plFALLACIAAJSTICKLEDGMANN1990]. 

Fallacia balnearis (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 201 (2000). 

Basionym: Navicula pygmaea var. balnearis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 10: fig. 8 

(1880). 

Fallacia cassubiae Witkowski, Diatom Res. 6, p. 403, figs 2-28 (1991). 

Fallacia clepsidroides Witkowski, Biblioth. Diatomol. 28, p. 120, pl. 26: figs 1-12 (1994). 

Fallacia costericardii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 181, pl. 22: figs 4-6 (1998). 

Fallacia cunoniae (Cholnoky) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 203 (2000). 

Basionym: Navicula cunoniae Cholnoky, Bot. Mar. 5, p. 56, figs 46-47 (1963). 

Fallacia ecuadoriana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 125-126, pl. 69: figs 1-8 (2000). 

Fallacia florinae (M.Møller) Witkowski, Diatom Res. 8, p. 215 (1993). 

Basionym: Navicula florinae M.Møller, Folia Geogr. Danica 3, p. 204, fig. 9 (1950). 

Fallacia inscriptura (Hendey) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 206 (2000). 

Basionym: Navicula inscriptura Hendey in Simonsen, Proceedings of the Fourth Symposium on Recent and Fossil Marine 

Diatoms, p. 374, fig. 9 (1977). 

Fallacia lange-bertalotii (E.Reichardt) E.Reichardt, Iconogr. Diatomol. 9, p. 126 (2000). 

Basionym: Navicula lange-bertalotii E.Reichardt, Ber. Bayer. Bot. Ges. 56, p. 176, pl. 1: figs 18-28, pl. 5: figs 1-3 (1985). 

Fallacia marnieri (Manguin) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 207 (2000). 

Basionym: Navicula marnieri Manguin, Rev. Algol. N.S. 3, p. 127, fig. 35 (1957). 

Fallacia melanocephala (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 208 (2000). 

Basionym: Navicula melanocephala Giffen, Bot. Mar. 18, p. 84, figs 78-80 (1975). 

Fallacia minima (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 208 (2000). 

Basionym: Navicula forcipata var. minima Østrup, Meddel. Grønland 18, p. 436, pl. 5: fig. 57 (1895). 

Fallacia nicobarica (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 208 (2000). 

Basionym: Navicula nicobarica Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 13, p. 150, pl. 5: fig. 8 (1863). 

Fallacia reichardtii (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 212 (2000). 

Basionym: Navicula reichardtii Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 10: fig. 9 (1880). 

Fallacia schoemaniana (Foged) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 213 (2000). 

Basionym: Navicula schoemaniana Foged, Biblioth. Phycol. 16, p. 43, pl. 19: figs 19-20 (1975). 

Fallacia spatiata (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 213 (2000). 

Basionym: Navicula forcipata var. spatiata Østrup, Meddel. Grønland 18, p. 436, pl. 5: fig. 60 (1895). 

Fallacia suspirii (Cholnoky) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 214 (2000). 

Basionym: Navicula suspirii Cholnoky, Hydrobiologia 17, p. 312, figs 60-62 (1961). 

FISTULIFERA Lange-Bert., Arch. Protistenk 148, p. 73 (1997). [Typus generis: Fistulifera saprophila (Lange-Bert. et 

Bonik) Lange-Bert., Arch. Protistenk 148, p. 73 (1997).] 



39 

Fistulifera iranensis (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 73 (1997). 

Basionym: Navicula iranensis Hust., Rabenhorsts Kryptogamenflora 7 (3) 2. Lieferung, p. 262, fig. 1390A (1962). 

Fistulifera pelliculosa (Bréb. ex Kütz.) Lange-Bert., Arch. Protistenk. 148, p. 73 (1997). 

Basionym: Frustulia pelliculosa Bréb. ex Kütz., Species Algarum, p. 40 (1849). 

Fistulifera saprophila (Lange-Bert. et Bonik) Lange-Bert., Arch. Protistenk. 148, p. 73 (1997). 

Basionym: Navicula saprophila Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p.312, figs 8-9, 29 

(1976). [Fistulifera saprophila is the type of the name of the genus FISTULIFERA.] 

FOGEDIA Witkowski, Lange-Bert., Metzeltin et Bafana, Nova Hedwigia 65, p. 81 (1997). [Typus generis: Fogedia 

giffeniana (Foged) Witkowski, Lange-Bert., Metzeltin et Bafana, Nova Hedwigia 65, p. 81 (1997).] 

Fogedia acuta (Salah) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 216 (2000). 

Basionym: Navicula finmarchica var. acuta Salah, Hydrobiologia 7, p. 78, pl. 2: fig. 8 (1955). 

Fogedia christensenii Witkowski, Metzeltin et Lange-Bert., Nova Hedwigia 65, p. 85, figs 30-45 (1997). 

Fogedia finmarchia (Cleve et Grunow) Witkowski, Metzeltin et Lange-Bert., Nova Hedwigia 65, p. 87 (1997). 

Basionym: Stauroneis finmarchia Cleve et Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 47, pl. 3: fig. 63 (1880). 

Fogedia geisslerae Witkowski, Metzeltin et Lange-Bert., Nova Hedwigia 65, p. 90, figs 64-92 (1997). 

Fogedia giffeniana (Foged) Witkowski, Lange-Bert., Metzeltin et Bafana var. giffeniana, Nova Hedwigia 65, p. 81 (1997). 

Basionym: Navicula giffeniana Foged, Biblioth. Phycol. 16, p. 38, pl. 22: figs 16-17 (1975). 

Fogedia giffeniana var. yemenitica Bafana, Nova Hedwigia 65, p. 82, figs 23-29 (1997). 

Fogedia heterovalvata (Simonsen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 219 (2000). 

Basionym: Navicula heterovalvata Simonsen, Kieler Meeresforsch., p. 81, pl. 2: figs 7-12 (1959). 

FRAGILARIA Lyngb. 


FRAGILARIA subgen. ULNARIA [see ULNARIA]. 

FRAGILARIA subgen. ALTERASYNEDRA, nom. illeg. [see ALTERASYNEDRA and ULNARIA]. 

Fragilaria acidoclinata Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 41-42, pl. 14: figs 8-13 (1993). [Validation of 

Fragilaria acidoclinata Lange-Bert. et G.Hofmann, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von 


Fragilaria acus (Kütz.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 582 (2000). 

Basionym: Synedra acus Kütz., Kieselschalige Bacillarien, p. 68, pl. 15: fig. 7 (1844). 

[≡ Fragilaria ulna var. acus (Kütz.) Lange-Bert.] 

Fragilaria acutirostrata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 89, pl. 1: figs 18-19, 17, 32, pl. 2: fig. 4 

(1998). 

Fragilaria amicorum Witkowski et Lange-Bert., Limnologica 23, p. 62, fig. 3: a-d, j-k (1993). 

Fragilaria amphicephala (Kütz.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3),p. 581 (2000). 

Basionym: Synedra amphicephala Kütz., Kieselschalige Bacillarien, p. 64, pl. 3: fig. 12 (1844). 

[≡ Fragilaria capucina ssp. amphicephala (Kütz.) Lange-Bert.] 

Fragilaria austriaca (Grunow) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 581 (2000). 

Basionym: Synedra amphicephala var. austriaca Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Atlas, pl. 39: 

figs 16a, b (1881). 

[≡ Fragilaria capucina ssp. austriaca (Grunow) Lange-Bert.] 

[≡ Fragilaria capucina var. austriaca (Grunow) Lange-Bert.] 

Fragilaria barbararum Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 15, pl. 48: figs 32-35, pl. 73: figs 6, 

8-9 (1996). 

Fragilaria barbatula (Kütz.) Lange-Bert., Biblioth. Diatomol. 27, p. 54 (1993). 

Basionym: Synedra barbatula Kütz., Kieselschalige Bacillarien, Biblioth. Diatomol. 27, p. 68, pl. 15: fig. 4 (1844). 

Fragilaria berolinensis (Lemmerm.) Lange-Bert., Biblioth. Diatomol. 27, p. 43 (1993). 

Basionym: Synedra berolinensis Lemmerm., Ber. Deutsch. Bot. Ges. 18, p. 31 (1900). 

[Nomenclatural synonym of Staurosira berolinensis (Lemmerm.) Lange-Bert.] 



40 

Fragilaria biceps (Kütz.) Lange-Bert., Biblioth. Diatomol. 27, p. 43 (1993). 

Basionym: Synedra biceps Kütz., Kieselschalige Bacillarien, p. 66, pl. 14: fig. 18; pl. 14: fig. 21 (1844). 

[Nomenclatural synonym of Ulnaria biceps (Kütz.) Compère in Jahn et al., p. 100 (2001).] 

Fragilaria bronkei Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 48, 427, pl. 12: figs 1-12 (2000). 

Fragilaria canariensis Lange-Bert., Biblioth. Diatomol. 27, p. 43, pl. 14: figs 1-6 (1993). 

Fragilaria capitata (Ehrenb.) Lange-Bert., nom. illeg., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra capitata Ehrenb., Ann. Phys. Chem. II 38, p. 221, pl. 3: fig. 3 (1) (1836). 

[Nomenclatural synonym of Ulnaria capitata (Ehrenb.) Compère in Jahn et al., p. 100 (2001). According to Compère not 

identical with Fragilaria capitata Ehrenb.] 

Fragilaria capucina ssp. amphicephala (Kütz.) Lange-Bert., Biblioth. Diatomol. 27, p. 44 (1993). 

Basionym: Synedra amphicephala Kütz., Kieselschalige Bacillarien, p. 64, pl. 3: fig. 12 (1844). 

[Nomenclatural synonym of Fragilaria amphicephala (Kütz.) Lange-Bert.] 

Fragilaria capucina ssp. austriaca (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 45 (1993). 

Basionym: Synedra (amphicephala var. ) austriaca Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 39: 

figs 16a, b (1881). 

[≡ Fragilaria capucina var. austriaca (Grunow) Lange-Bert.] 

[Nomenclatural synonym of Fragilaria austriaca (Grunow) Lange-Bert.] 

Fragilaria capucina ssp. rumpens (Kütz.) Lange-Bert., Biblioth. Diatomol. 27, p. 45 (1993). 

Basionym: Synedra rumpens Kütz., Kieselschalige Bacillarien, p. 69, pl. 6: fig. 45 (1844). 

Fragilaria capucina var. austriaca (Grunow) Lange-Bert., Nova Hedwigia 33, p. 748 (1980). 

Basionym: Synedra (amphicephala var. ) austriaca Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 39: 

fig. 16 (1881). 

[≡ Fragilaria capucina ssp. austriaca (Grunow) Lange-Bert.] 

[Nomenclatural synonym of Fragilaria austriaca (Grunow) Lange-Bert.] 

Fragilaria capucina var. vaucheriae (Kütz.) Lange-Bert., Nova Hedwigia 33, p. 747 (1980). 

Basionym: Exilaria vaucheriae Kütz., Linnaea 8, p. 560, pl. 15: fig. 38 (1833). 

Fragilaria cassubica Witkowski et Lange-Bert., Limnologica 23, p. 65, fig. 4: a-f, n (1993). 

Fragilaria costata (Rehakova) Lange-Bert., Acta Biol. Benrodis 3, p. 120 (1991). 

Basionym: Diatoma costata Rehakova, Paleontologie 23, p. 112, pl. 13: figs 4, 9, 10 (1980). 

[Taxonomic synonym: Diatoma costata var. curta Rehakova, Paleontologie 23, p. 113, pl. 13: fig. 8 (1980), according to 

Lange-Bertalot in Acta Biol. Benrodis 3, p. 120 (1991).] 

Fragilaria crassa Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 89-90, pl. 1: figs 20-23, pl. 2: fig. 1 (1998). 

Fragilaria cyclopum (Brutschy) Lange-Bert., Nova Hedwigia 33, p. 752 (1980). 

Basionym: Synedra cyclopum Brutschy, Int. Rev. Gesamten Hydrobiol. Hydrogr. 10, p. 284, fig. 11 (1922). 

Fragilaria danica (Kütz.) Lange-Bert., Iconogr. Diatomol. 2, p. 54 (1996). 

Basionym: Synedra danica Kütz., Kieselschalige Bacillarien, p. 66, pl. 14: fig. 13 (1844). 

Fragilaria delicatissima (W.Sm.) Lange-Bert. var. delicatissima, Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra delicatissima W.Sm., Synopsis British Diatomaceae 1, p. 72, pl. 12: fig. 94 (1853). 

Fragilaria delicatissima var. angustissima (Grunow) Lange-Bert., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra delicatissima var. angustissima Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 39: fig. 

10 (1881). 

Fragilaria demerarae (Grunow) Lange-Bert., Nova Hedwigia 33, p. 752 (1980). 

Basionym: Synedra demararae Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 41, fig. 29 (1881). 

Fragilaria dilatata (Bréb.) Lange-Bert., Biblioth. Diatomol. 27, p. 45 (1993). 

Basionym: Frustulia dilatata Bréb., Considérations sur les Diatomées, p. 17 (1838). 

Fragilaria dorsiventralis (O.Müll.) Lange-Bert., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra dorsiventralis O.Müll., Bot. Jahrb. Syst. 45, p. 114, fig. 3 (1910). 

Fragilaria eichhornii Witkowski et Lange-Bert., Fragm. Florist. Geobot. 40, p. 731, fig. 1-9 (1995). 



41 

Fragilaria exiguiformis Lange-Bert., Biblioth. Diatomol. 27, p. 45 (1993). 

Replaced synonym: Fragilaria (virescens var.) exigua Grunow in Cleve et Möller, Diatoms, collection of 324 slides, no. 144. 

Fragilaria famelica (Kütz.) Lange-Bert. ssp. famelica, Nova Hedwigia 33, p. 749 (1980). 

Basionym: Synedra famelica Kütz., Kieselschalige Bacillarien, p. 64, pl. 14: fig. 8 (1) (1844). 

Fragilaria famelica ssp. littoralis (H.Germ.) Lange-Bert., Biblioth. Diatomol. 27, p. 46 (1993). 

Basionym: Fragilaria intermedia var. littoralis H.Germ., Flore des Diatomées, p. 68, pl. 20: figs 11-21 (1981). 

[Nomenclatural synonym of Fragilaria henryi Lange-Bert.] 

Fragilaria fasciculata (C.Agardh) Lange-Bert., Nova Hedwigia 33, p. 750 (1980). 

Basionym: Diatoma fasciculatum C. Agardh, Disp. Alg. Sveciae, p. 35 (1812). 

Fragilaria fossilis (Pant.) Lange-Bert., Acta Biol. Benrodis 3, p. 121 (1991). 

Basionym: Diatoma fossilis Pant., Beschreibung neuer Bacillarien Ungarns 3, Pozsony, p. 46 (1905). 

Fragilaria gaillonii (Bory) Lange-Bert., Nova Hedwigia 33, p. 751 (1980). 

Basionym: Navicula gaillonii Bory, Encyclop. Method. Hist. Nat. Zoophytes. 2, p. 564 (1824). 

[Comment: The basionym author is given as "Ehrenberg" in Nova Hedwigia 33, p. 750 (1980).] 

Fragilaria gedanensis Witkowski, Nova Hedwigia 56, p. 498, figs 1-18 (1993). 

Fragilaria geocollegarum Witkowski et Lange-Bert., Fragm. Florist. Geobot. 40, p. 731, 734, figs 16-22 (1995). [Validation 

of Fragilaria geocollegarum Witkowski et Lange-Bert., nom. inval., Bilioth. Diatomol. 28, p. 127, pl. 6: figs 1-5, pl. 9: fig. 4 

(1994).] 

Fragilaria germainii E.Reichardt et Lange-Bert. in Ricard, Ouvrage dédié à la Mémoire du Prof. H. Germain, p. 204, pl. 1: 

figs 1-13 (1990). 

Fragilaria germainii f. acostata Lange-Bert., Iconogr. Diatomol. 9, p. 127, pl. 5: figs 15-20, pl. 6: figs 4-6 (2000). 

Fragilaria guenter-grassii Witkowski et Lange-Bert., Limnologica 23, p. 65, fig. 5: a-h (1993). 

Fragilaria henryi Lange-Bert., Iconogr. Diatomol. 6, p. 50 (1999). 

Replaced synonym: Fragilaria intermedia var. littoralis H.Germ., Flore des Diatomées, p. 68, pl. 20: figs 11-21 (1981). 

[≡ Fragilaria famelica ssp. littoralis (H.Germ.) Lange-Bert.] 

Fragilaria improbula Witkowski et Lange-Bert., Fragm. Florist. Geobot. 40, p. 734, figs 28-38 (1995). 

Fragilaria incisa (C.S.Boyer) Lange-Bert., Nova Hedwigia 33, p. 748 (1980). 

Basionym: Synedra incisa C.S.Boyer, Bull. Torrey Bot. Club 47, p. 68, pl. 2: fig. 8 (1920). 

Fragilaria incognita E.Reichardt, Diatom Res. 3, p. 237-238, figs 1-10 (1988). 

Fragilaria labei Witkowski et Metzeltin, Iconogr. Diatomol. 4, p. 16, pl. 49: figs 31-40, pl. 71: figs 1-2, pl. 74: figs 4-6 

(1996). 

Fragilaria laevigata (Grunow) Lange-Bert., Nova Hedwigia 33, p. 752 (1980). 

Basionym: Synedra laevigata Grunow, Monthly Microscop. J. 18, p. 166, pl. 193: fig. 3 (1877). 

Fragilaria lanceolata (Kütz.) E.Reichardt, Nova Hedwigia 47, p. 85 (1988). 

Basionym: Synedra lanceolata Kütz., Kieselschalige Bacillarien, p. 66, pl. 30: fig. 31 (1844). 

[Nomenclatural synonym of Ulnaria lanceolata (Kütz.) Compère in Jahn et al., p. 100 (2001).] 

Fragilaria lenzii (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 184 (1996). 

Basionym: Synedra lenzii Krasske, Arch. Hydrobiol. 35, p. 554, figs 3-7 (1939). 

Fragilaria leptostauron var. trigona (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 83 (1996). 

Basionym: Fragilaria harrissonii var. trigona Krasske, Arch. Hydrobiol. 24, p. 437, fig. 7 (1932). 

[Nomenclatural synonym of Staurosira pinnata var. trigona (Krasske) Lange-Bert.] 

Fragilaria loetschertii Lange-Bert., Ann. Limnol. 21, p. 214-215, figs 13-17, 30-34 (1985). 

Fragilaria longiceps (Ehrenb.) Lange-Bert., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra longiceps Ehrenb., Ber. Bekanntm. Verh. Königl. Preuss. Akad. Wiss. Berlin 1845, p. 81 (1845). 

Fragilaria martyi (Hérib.) Lange-Bert., Biblioth. Diatomol. 27, p. 46 (1993). 

Basionym: Opephora martyi Hérib., Les Diatomées d’Auvergne, p. 43, pl. 8: fig. 20 (1902). 

[Nomenclatural synonym of Staurosira martyi (Hérib.) Lange-Bert.] 



42 

Fragilaria mazamaensis (Sovereign) Lange-Bert., Nova Hedwigia 33, p. 748 (1980). 

Basionym: Synedra mazamaensis Sovereign, Trans. Amer. Microscop. Soc. 77 (2), p. 111, pl. 2: figs 10-15 (1958). 

Fragilaria montana (Krasske) Lange-Bert., Nova Hedwigia 33, p. 747 (1980). 

Basionym: Synedra montana Krasske in Hustedt, Rabenhorsts Kryptogamenflora 7 (2) 2. Lieferung, p. 204, fig. 694 (1932); 

Krasske, Hedwigia 72, p. 99, pl. 2: fig. 2 (1932). 

Fragilaria nana (F.Meister) Lange-Bert., nom. illeg., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra nana F. Meister, Kryptogamenflora der Schweiz 4, p. 76, pl. 8: fig. 9 (1912). 

[Nomenclatural synonym of Fragilaria nanana Lange-Bert.] 

Fragilaria nanana Lange-Bert., Biblioth. Diatomol. 27, p. 48 (1993). 

Replaced synonym: Synedra nana F.Meister, Kieselalgen der Schweiz, p. 76, pl. 8: fig. 9 (1912). 

[≡ Fragilaria nana (F.Meister) Lange-Bert., nom. illeg., Nova Hedwigia 33, p. 746 (1980).] 

Fragilaria nanoides Lange-Bert., Iconogr. Diatomol. 2, p. 55, pl. 109: figs 2-6 (1996). 

Fragilaria neoelliptica Witkowski, Biblioth. Diatomol. 28, p. 128-129, pl. 10: figs 1-13 (1994). 

Fragilaria neoproducta Lange-Bert., Biblioth. Diatomol. 27, p. 48 (1993). 

[– Fragilaria neoproducta Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (3), p. 136, pl. 125: fig. 3, pl. 127: figs 1-5a (1991).] 

Fragilaria oldenburgioides Lange-Bert., Iconogr. Diatomol. 2, p. 55-56, pl. 7: figs 42-47, pl. 111: figs 8-9 (1996). 

Fragilaria opacolineata Lange-Bert., Iconogr. Diatomol. 2, p. 56, pl. 7: figs 36-41b, pl. 111: figs 2-3 (1996). 

Fragilaria perminuta (Grunow) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 581 (2000). 

Basionym: Synedra perminuta Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 40: fig. 23 (1881). 

Fragilaria polonica Witak et Lange-Bert., Fragm. Florist. Geobot. 40, p. 736, figs 39-45 (1995). 

Fragilaria pulchella (Ralfs ex Kütz.) Lange-Bert. var. pulchella, Nova Hedwigia 33, p. 749 (1980). 

Basionym: Exilaria pulchella Ralfs ex Kütz., Kieselschalige Bacillarien, p. 68 (1844). 

Fragilaria pulchella var. constricta (B.Liebetanz) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 53 (2000). 

Basionym: () Synedra pulchella [var. lanceolata] f. constricta B.Liebetanz, Bull. Int. Acad. Polon. Sci. Letters, Classe 

Math.-Nat. ser. B. 1925, p. 110 (1926). 

Fragilaria pulchella var. lanceolata (O`Meara) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 52 (2000). 

Basionym: Synedra pulchella var. lanceolata O`Meara, Proc. Roy. Irish Acad. (2. Ser.) 2, p. 304, pl. 28: fig. 20 (1876). 

Fragilaria quadrata (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 57 (1996). 

Basionym: Fragilaria undata var. quadrata Hust., Süßwasserflora Mitteleuropas 10, p. 144, fig. 149B (1930). 

Fragilaria quebecensis (Poulin, Hudon et Cardinal) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 53 

(2000). 

Basionym: Fragilaria virescens var. quebecensis Poulin, Hudon et Cardinal, Naturaliste Can. (Rev. Ecol. Syst.) 109, p. 280, 

pl. 1a: figs 3-6 (1982). 

Fragilaria radians (Kütz.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 581 (2000). 

Basionym: Synedra radians Kütz., Kieselschalige Bacillarien, p. 64, pl. 14: figs 7: 1-4 (1844). 

Fragilaria reicheltii (Max Voigt) Lange-Bert., p. 48. 

Basionym: Centronella reicheltii Max Voigt, Zool. Anzeiger 24 (Nr. 640), pl. 2: fig. 10 (1902). 

Fragilaria rolandschmidtii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 90, pl. 1: figs 14-16 (1998). 

Fragilaria santaremensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 91, pl. 1: figs 24-28, 33, pl. 2: fig. 2 (1998). 

Fragilaria socia (Wallace) Lange-Bert., Nova Hedwigia 33, p. 749 (1980). 

Basionym: Synedra socia Wallace, Notul. Nat. Acad. Nat. Sci. Philadelphia 331, p. 1, pl. 1: fig. 1 A-E (1960). 

Fragilaria sopotensis Witkowski et Lange-Bert., Limnologica 23, p. 67, fig. 6: a-p (1993). 

Fragilaria spinarum Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 57, pl. 7: figs 33-35 (1996). [Comment: pl. l7: figs 

31-32 is not mentioned on p. 57.] 



43 

Fragilaria subsalina (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 49 (1993). 

Basionym: Fragilaria virescens var. subsalina Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 44: fig. 5 

(1881). 

Fragilaria synegrotesca Lange-Bert., Biblioth. Diatomol. 27, p. 49-50, pl. 12: figs 1-11, pl. 13: figs 1-6 (1993). 

Fragilaria tabulata (C.Agardh) Lange-Bert. var. tabulata, Nova Hedwigia 33, p. 750 (1980). 

Basionym: Diatoma tabulatum C.Agardh, Conspectus Criticus Diatomacearum, p. 50 (1832). 

Fragilaria tabulata var. australis (Manguin) Lange-Bert., Nova Hedwigia 33, p. 750 (1980). 

Basionym: Synedra tabulata var. australis Manguin, Ann. Sci. Nat. Bot. 12, p. 301, pl. 13: fig. 142 (1960). 

Fragilaria tabulata var. obtusa (Pantocsek) Lange-Bert., Nova Hedwigia 33, p. 751 (1980). 

Basionym: Synedra tabulata var. obtusa Pantocsek, Beitr. zur Kenntnis der fossilen Bacillarien Ungarns Teil 2, p. 64, pl. 26: 

figs 377, 380 (1889). 

Fragilaria tabulata var. tenuistriata (F.Meister) Lange-Bert., Nova Hedwigia 33, p. 751 (1980). 

Basionym: Synedra tenuistriata F.Meister, Ber. Schweiz. Bot. Ges. 44, p. 104, fig. 95 (1935). 

Fragilaria tabulata var. truncata (Grev.) Lange-Bert., Nova Hedwigia 33, p. 751 (1980). 

Basionym: Echinella fasciculata var. truncata Grev., Scottish Cryptogam. Flora 1, p. 37, pl. 16: fig. 4 (1823). 

Fragilaria tenera (W.Sm.) Lange-Bert., Nova Hedwigia 33, p. 746 (1980). 

Basionym: Synedra tenera W.Sm., Synopsis British Diatomaceae 2, p. 98 (1856). 

Fragilaria ulna (Nitzsch) Lange-Bert. var. ulna, Nova Hedwigia 33, p. 745 (1980). 

Basionym: Bacillaria ulna Nitzsch, Neue Schriften Naturf. Ges. Halle 3, p. 99, pl. 5 (1817). 

[Nomenclatural synonym of Ulnaria ulna (Nitzsch) Compère in Jahn et al., p. 100 (2001).] 

Fragilaria ulna var. acus (Kütz.) Lange-Bert., Nova Hedwigia 33, p. 745 (1980). 

Basionym: Synedra acus Kütz., Kieselschalige Bacillarien, p. 68, pl. 15: fig. 7 (1844). 

[Nomenclatural synonym of Fragilaria acus (Kütz.) Lange-Bert.] 

Fragilaria ulna var. goulardi (Brébisson ex Cleve et Grunow) Lange-Bert., Nova Hedwigia 33, p. 745 (1980). 

Basionym: Synedra goulardi Brébisson ex Cleve et Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 107, pl. 6: 

fig. 119 (1880). 

Fragilaria ulna var. ungeriana (Grunow) Lange-Bert., Nova Hedwigia 33, p. 745 (1980). 

Basionym: Fragilaria ungeriana Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 13, p. 142, pl. 13: fig. 18 (1863). 

[Nomenclatural synonym of Ulnaria ungeriana (Grunow) Compère in Jahn et al., p. 100 (2001).] 

Fragilaria utermoehlii (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 52 (1993). 

Basionym: Synedra utermoehlii Hust., Rabenhorsts Kryptogamenflora 7 (2) 2. Lieferung, p. 185, fig. 687 (1932). 

Fragilaria viereckiana Gerd Moser, Biblioth. Diatomol. 43, p. 127-128, pl. 2: figs 1-17, pl. 3: fig. 2 (1999). 

FRAGILARIOPSIS Hust. 


Fragilariopsis bjoernoeyaensis Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 16, pl. 67: figs 20-21, pl. 87: 

fig. 8 (1996). 

Fragilariopsis humboldtianorum Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 17, pl. 67: figs 25-27, pl. 

87: figs 5, 9-10 (1996). 

Fragilariopsis reginae-jahniae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 361, 427, pl. 213: figs 31-36 

(2000). 

FRANKOPHILA Lange-Bert., Arch. Protistenk 148, p. 66 (1997). [Typus generis: Frankophila similioides Lange-Bert. et 

U.Rumrich, Arch. Protistenk 148, p. 66 (1997). ] 

Frankophila horstii U.Rumrich et M.Rumrich, Iconogr. Diatomol. 9, p. 128, pl. 9: figs 1-10 (2000). 

Frankophila loetschertii (Lange-Bert.) Lange-Bert., Arch. Protistenk. 148, p. 71 (1997). 

Basionym: Fragilaria loetschertii Lange-Bert., Ann. Limnol. 21, p. 214, figs 13-17, 30-34 (1985). 

Frankophila maillardii (Le Cohu) Lange-Bert., Arch. Protistenk. 148, p. 71 (1997). 

Basionym: Fragilaria maillardii Le Cohu, Ann. Limnol. 21, p. 213, figs 1-3, 18-24 (1985). 



44 

Frankophila similioides Lange-Bert. et U.Rumrich Arch. Protistenk 148, p. 66, fig. 1-15 (1997). [Frankophila similioides is 

the type of the name of the genus FRANKOPHILA Lange-Bert.]. 

FRUSTULIA Rabenh. 


Frustulia acidophilissima Wydrzycka et Lange-Bert., Brenesia 55-56, p. 9, pl. 10: figs 1-9 (2001). 

Frustulia alfinitosilviae Lange-Bert., Diatoms of Europe 2, p. 219, pl. 139: figs 1-6 (2001). 

Frustulia altimontana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 93, pl. 113: figs 1-5 (1998). 

Frustulia amosseana Lange-Bert., Iconogr. Diatomol. 9, p. 129 (2000). 

Replaced synonym: Berkella linearis R.Ross et Sims, Bacillaria 1, p. 156, figs 20-27 (1978). 

Frustulia angustata (Pant. et Greguss) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. angustata Pant. et Greguss, Bot. Közlem. 12, p. 211, pl. 7: fig. 80 (1913). 

Frustulia australis Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 182-183, pl. 35: figs 1-3 (1998). 

Frustulia baicalensis (Skvortzov et D.I.Mey.) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. baicalensis Skvortzov et D.I.Mey., Proc. Sungaree River Biol. Sta. 1 (5), p. 15, 

pl. 1: fig. 38 (1928). 

Frustulia chilensis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 131, pl. 91: figs 1-3 (2000). 

Frustulia corneliae Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 131-132, pl. 100: figs 1-9 (2000). 

Frustulia crassinervia (Bréb.) Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 57 (1996). 

Basionym: Navicula crassinervia Bréb. in W. Smith, Synopsis British Diatomaceae 1, p. 47, pl. 31: fig. 271 (1853). 

Frustulia crassipunctata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 94, pl. 112: figs 1-6 (1998). 

Frustulia disjuncta Lange-Bert., Diatoms of Europe 2, p. 219, pl. 135: figs 3-7 (2001). 

Frustulia ellipticoides Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Replaced synonym: Frustulia rhomboides var. elliptica Hust., Arch. Hydrobiol. Suppl. 15, p. 214, pl. 16: fig. 2 (1937). 

Frustulia elongata (Willi Krieg.) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. elongata Willi Krieg., Ber. Deutsch. Bot. Ges. 54, p. 82, pl. 10: fig. 16 (1936). 

Frustulia elongatissima (Manguin) Lange-Bert. et Metzeltin, Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. elongatissima Manguin, Algues d’eau douce de la Guadeloupe, SEDES, Paris, p. 54, 

fig. 47 (1952). 

Frustulia erifuga Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 58 (1996). 

Replaced synonym: Frustulia rhomboides var. viridula (Bréb. ex Kütz.) Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26, 

p. 94 (1894). 

[Comment: For the taxonomic concept of Frustulia erifuga Lange-Bert. et Krammer in Iconogr. Diatomol. 2, p. 58, pl. 39: 

figs 1-2 (1996) see Frustulia hustedtii (H.Germ.) Lange-Bert.] 

Frustulia gaertnerae Lange-Bert., Iconogr. Diatomol. 2, p. 59, pl. 39: figs 7-8, pl. 119: figs 3-4 (1996). [Comment: 

Frustulia gaertnerae on p. 59, 196, and 356 is misspelled as "Frustulia gaertneriae".] 

Frustulia guayanensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 95, pl. 115: figs 10-11 (1998). 

Frustulia guayanensis ssp. ecuadoriana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 133, pl. 99: figs 9-14 (2000). 

Frustulia huberi (F.Meister) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. huberi F.Meister, Z. Bot. 23, p. 458, fig. 6 (1930). 

Frustulia hustedtii (H.Germ.) Lange-Bert., Brenesia 55-56, p. 10 (2001). 

Basionym: Frustulia rhomboides [var. viridula] f. hustedtii H.Germ., Flore des Diatomées eaux douces et saumâtres du 

massif Armoricain, p. 138-140, pl. 53: figs 1-2 (1981). 

[– Frustulia erifuga sensu Lange-Bertalot et Krammer in Lange-Bertalot & Metzeltin, Iconogr. Diatomol. 2, p. 58, pl. 39: 

figs 1-2, according to Lange-Bertalot in Wydrzycka & Lange-Bertalot (2001), p. 10.] 

Frustulia kosmolliana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 133, pl. 84: fig. 5, pl. 95: figs 1-9 (2000). 



45 

Frustulia krammeri Lange-Bert. et Metzeltin, Iconogr. Diatomol. 5, p. 96 (1998). [Comment: Frustulia krammeri was 

documented already in Lange-Bertalot & Metzeltin (1996), Iconogr. Diatomol. 2, pl. 37: figs 1-3 as "Frustulia (nov.) spec. 

Nr. 1 Julma Ölkky" and on pl. 120: fig. 4 as "Frustulia nov. spec. Nr. 1 Julma Ölkky".] 

Frustulia lakatosii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 96-97, pl. 115: figs 7-9 (1998). 

Frustulia lange-bertalotii Metzeltin, Iconogr. Diatomol. 6, p. 51, pl. 37: fig. 8, pl. 38: figs 1-8 (1999). 

Frustulia leptocephala (Østrup) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. leptocephala Østrup, Meddel. Grønland 15, p. 257, pl. 1: fig 1 (1897). 

Frustulia longinqua Lange-Bert., Diatoms of Europe 2, p. 220, pl. 135: figs 8-10 (2001). 

Frustulia magaliesmontana ssp. americana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 97, pl. 116: figs 4-6 (1998). 

Frustulia magna Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 98, pl. 110: figs 1-4, pl. 113: fig. 6 (1998). 

Frustulia mamillaris Gerd Moser, Biblioth. Diatomol. 43, p. 129-130, pl. 4: figs 1-8, pl. 5: fig. 9 (1999). 

Frustulia mediocris Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 99, pl. 117: figs 16-17 (1998). 

Frustulia modesta Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 100, pl. 115: figs 1-6 (1998). 

Frustulia nana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 183-184, pl. 32: fig. 7, pl. 33: figs 6-10 

(1998). 

Frustulia neocaledonica var. minima Lange-Bert. et Gerd Moser, nom. inval., Biblioth. Diatomol. 32, p. 102, pl. 37: figs 5- 

6, pl. 40: figs 5-8, pl. 42: figs 10-11, pl. 44: fig. 7 (1995). [Comment: lacking a type and a Latin diagnosis, this name is 

invalidly published.] 

Frustulia neofrenguellii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 230, pl. 94: figs 1-7 (2000). 

Frustulia neomundana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 135, pl. 97: figs 1-12 (2000). 

Frustulia pangaea Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 101, pl. 111: figs 1-4 (1998). 

Frustulia pangaeopsis Lange-Bert., Diatoms of Europe 2, p. 220, pl. 130: figs 1-6 (2001). 

Frustulia pararhomboides Metzeltin et Lange-Bert. var. pararhomboides, Iconogr. Diatomol. 5, p. 102, pl. 114: figs 1-9, pl. 

116: figs 1-3 (1998). [Comment: Frustulia pararhomboides on p. 466 (caption of pl. 116: figs 1-3) is cited as "Frustulia aff. 

pararhomboides".] 

Frustulia pararhomboides var. subundosa Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 103, pl. 114: figs 10-12 

(1998). [Comment: on page 462 (caption of pl. 114: figs 10-12) the var. subundosa is misspelled as "var. subnodosa".] 

Frustulia peridotitica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 184-185, pl. 33: figs 1-5 (1998). 

Frustulia pumilio Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 136, pl. 99: figs 1-8 (2000). 

Frustulia pusilla (F.Meister) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. pusilla F.Meister, Kieselalgen aus Asien. Bornträger, Berlin, p. 35, pl. 12: fig. 91 

(1932). 

Frustulia quadrisinuata Lange-Bert., Iconogr. Diatomol. 2, 59-60, pl. 38: figs 10-12, pl. 119: figs 1-2 (1996). 

Frustulia quadrisinuata ssp. meridiana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 104, pl. 111: figs 5-6 (1998). 

Frustulia rhodesica (Cholnoky) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. rhodesica Cholnoky, Portugal. Acta. Biol. Ser. B. 4, p. 213, pl. 3: fig. 60 (1954). 

Frustulia rostratiformis Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Replaced synonym: Frustulia rhomboides var. rostrata Manguin, Travaux Algol. Sér. 1 (Rev. Algol. 13), p. 133, pl. 2: fig. 

26; non Frustulia rostrata Hust. 1937 (1942). 

Frustulia saxonica f. nipponica (Skvortzov) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. saxonica f. nipponica Skvortzov, Philipp. J. Sci. 61, p. 265, pl. 4: fig. 14 (1936). 

Frustulia septentrionalis Lange-Bert., Iconogr. Diatomol. 2, p. 61, pl. 37: figs 4-5, pl. 119: figs 5-6 (1996). 

Frustulia spicula ssp. judaica Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of continental 

Israel, p. 249, pl. 18: figs 122-129 (2000). 



46 

Frustulia stagnalis Gerd Moser, Biblioth. Diatomol. 43, p. 131, pl. 5: figs 1-6 (1999). 

Frustulia tenuirostris (Cleve-Euler) Lange-Bert., Syst. Geogr. Pl. 70, p. 258 (2000). 

Basionym: Frustulia rhomboides var. tenuirostris Cleve-Euler, Kongl. Svenska Vetenskapsakad. Handl. Fj. Ser. 3, p. 8, 

fig. 1326 b (1952). 

Frustulia undosa Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 105, pl. 116: figs 14-18, pl. 117: figs 1-7 (1998). 

Frustulia zizkae Lange-Bert., Iconogr. Diatomol. 5, p. 106-107, pl. 116: figs 7-13 (1998). 

GEISSLERIA Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 63 [Typus generis: Geissleria moseri Metzeltin, 

Witkowski et Lange-Bert., Iconogr. Diatomol. 2, p. 66 (1996).] 

Geissleria acceptata (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 64 (1996). 

Basionym: Navicula acceptata Hust., Arch. Hydrobiol. 43, p. 398, pl. 38: figs 66-67 (1950). 

Geissleria annulata (Grunow) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 64 (1996). 

Basionym: Navicula annulata Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 37 (1880). 

Geissleria arkonensis Lange-Bert., Diatoms of Europe 2, p. 220-221, pl. 96: figs 1-4 (2001). 

Geissleria boreosiberica Lange-Bert., Genkal et Vekhov, Diatoms of Europe 2, p. 121, 221, pl. 96: figs 12-15 (2001). 

[– Geissleria boreosiberica Lange-Bert., Genkal et Vekhov, nom. inval., Iconogr. Diatomol. 6, p. 28, pl. 20: figs 9-13, 14 

(1999).] 

[Comment: On p. 221 in Diatoms of Europe 2 the authors are missing.] 

Geissleria cummerowi (L.Kalbe) Lange-Bert., Diatoms of Europe 2, p. 122, (2001). 

Basionym: Navicula cummerowii L.Kalbe, Neue Brehm Bücherei, p. 116, pl. 19: figs 268-269 (1980). 

Geissleria declivis (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 65 (1996). 

Basionym: Navicula declivis Hust., Abh. Vortr. Bremer Wiss. Ges. 8-9, p. 384, fig. 12 (1934). 

Geissleria decussis (Østrup) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 65 (1996). 

Basionym: Navicula decussis Østrup, Danske Diatomeer, p. 77, pl. 2: fig. 50 (1910). 

Geissleria dolomitica (W.Bock) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 65 (1996). 

Basionym: Navicula dolomitica W.Bock, Nova Hedwigia Beih. 31, p. 420, pl. 1: figs 8-10, pl. 3: figs 56-57 (1970). 

Geissleria ignota (Krasske) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 65 (1996). 

Basionym: Navicula ignota Krasske, Hedwigia 72, p. 116, pl. 3: fig. 19 (1932). 

Geissleria kriegeri (Krasske) Lange-Bert., Iconogr. Diatomol. 2, p. 66 (1996). 

Basionym: Navicula kriegeri Krasske, Ber. Deutsch. Bot. Ges. 61, p. 86, fig. 16 (1943). 

[Comment: This combination was again marked as new in Iconogr. Diatomol. 5, p. 108 (1998). This superfluous data set 

appears in Kusber & Metzeltin (2001), p. 614 erroneously as "Geissleria kriegeri (Krasske) Metzeltin et Lange-Bert."] 

Geissleria lange-bertalotii Metzeltin, Iconogr. Diatomol. 4, p. 236, pl. 1: figs 1-16, pl. 2: figs 19-23, pl. 3: figs 23-27 (1996). 

Geissleria mirabilis (Krasske) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 108 (1998). 

Basionym: Stauroneis mirabilis Krasske, Arch. Hydrobiol. 44, p. 649, figs 2-3 (1951). 

Geissleria moseri Metzeltin, Witkowski et Lange-Bert., Iconogr. Diatomol. 2, p. 66, pl. 31: figs 1-2, 7-8, pl. 123: fig. 4 

(1996). [Geissleria moseri is the type of the name of the genus GEISSLERIA Lange-Bert. et Metzeltin.] 

Geissleria neotropica Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 108, pl. 75: figs 25-34 (1998). 

Geissleria paludosa (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 67 (1996). 

Basionym: Navicula lagerstedtii var. palustris Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 400: figs 27-29 

(1934). 

Geissleria perelegans (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 109 (1998). 

Basionym: Navicula perelegans Hust., Bot. Not. 4, p. 403, fig. 127 (1952). 

Geissleria schmidiae Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 138, pl. 66: figs 1-15 (2000). 

Geissleria schoenfeldii (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 67 (1996). 

Basionym: Navicula schoenfeldii Hust., Süßwasserflora Mitteleuropas 10, p. 301, fig. 520 (1930). 

Geissleria similis (Krasske) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 68 (1996). 

Basionym: Navicula similis Krasske, Bot. Arch. 27, p. 354, fig. 15 (1929). 



47 

Geissleria subdecussis (Hust.) Lange-Bert., Diatoms of Europe 2, p. 128, (2001). 

Basionym: Navicula subdecussis Hust., Int. Rev. Gesamten Hydrobiol. 42, p. 74, figs 136-138 (1942). 

Geissleria thingvallae (Østrup) Metzeltin et Lange-Bert., Iconogr. Diatomol. 2, p. 68 (1996). 

Basionym: Navicula thingvallae Østrup, Botany of Iceland 2 (1) no. 5, p. 29, pl. 3: fig. 43 (1918). 

GOMPHONEIS (Cleve) Lange-Bert. et E.Reichardt (subgenus within Gomphonema Ehrenb.), Biblioth. Diatomol. 27, p. 73 

(1993). 

Basionym: GOMPHONEIS Cleve, Kongl. Svenska Vetenskapsakad. Handl. Ser. 2, 26, p. 73 (1894) [as genus]. 

GOMPHONEMA Ehrenb. 


GOMPHONEMA subgen. GOMPHONEIS [see GOMPHONEIS]. 

Gomphonema aequirostrum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 110, pl. 151: figs 7-11 (1998). 

Gomphonema affine var. rhombicum E.Reichardt, Iconogr. Diatomol. 8, p. 15, pl. 10: figs 1-15 (1999). 

Gomphonema alineae Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 56, pl. 84: figs 1-7 (1993). [As a part of the 

protologue is cited: Krammer & Lange-Bertalot (1985) Biblioth. Diatomol. 9, pl. 40: figs 9-11 and Lange-Bertalot (1980) 

Bot. Not. 133, p. 4: figs A-D.] [Comment: In Biblioth. Diatomol. 9, Gomphonema alineae was documented as 

"Gomphonema spec." in Bot. Not. 133 as "Gomphonema aff. herculeanum aus dem See von Tiberias, Israel".] 

Gomphonema americobtusatum E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 33, pl. 34: figs 1-18, pl. 35: figs 1-7 

(1999). 

Gomphonema amoenum Lange-Bert., Biblioth. Diatomol. 9, p. 42, pl. 35: figs 1-6, pl. 42: figs 7-8 (1985). [As a part of the 


cited.] 

Gomphonema angusticephalum E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 49-50, pl. 55: figs 5-8, pl. 60: figs 1- 

26 (1999). 

Gomphonema angustivalva E.Reichardt, Nova Hedwigia 65, p. 112, pl. 6, figs 1-29 (1997). 

Gomphonema archaevibrio f. cuneatum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 111, pl. 160: figs 4-7 (1998). 

Gomphonema archaevibrio Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 1, p. 15-16, pl. 26: figs 8, 9, 10-11, pl. 27: figs 

1-7, pl. 28: figs 1-6 (1995). [Comment: Gomphonema archaevibrio on p. 46 (= caption of pl. 26: fig. 9 as "Gomphonema 

vibrio Ehrenberg". Gomphonema archaevibrio Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 1 (second edition), p. 32, pl. 

26: figs 8, 9, 10-11, pl. 27: figs 1-7, pl. 28: figs 1-6 (1995). Gomphonema archaevibrio on p. 92 (= caption of pl. 26: fig. 9 

as "Gomphonema vibrio Ehrenberg". In the second edition Gomphonema archaevibrio Lange-Bert. et E.Reichardt was again 

marked as new.] 

Gomphonema astridae E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 29 (1999). 

Replaced synonym: Gomphonema parvulum var. undulatum A. Cleve, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 21, Afd. 

3 (2), p. 21, pl. 1: fig. 18 (1895). 

Gomphonema augur var. sphaerophorum (Ehrenb.) Lange-Bert., Biblioth. Diatomol. 9, p. 43 (1985). 

Basionym: Gomphonema sphaerophorum Ehrenb., Ber. Bekanntm. Verh. Königl. Preuss. Akad. Wiss. Berlin 1845, p. 78 

(1845). 

Gomphonema augur var. turris (Ehrenb.) Lange-Bert., Biblioth. Diatomol. 9, p. 44 (1985). 

Basionym: Gomphonema turris Ehrenb., Abh. Königl. Akad. Wiss. Berlin 1841, p. 416 (1843). 

Gomphonema bavaricum E.Reichardt et Lange-Bert., Nova Hedwigia 53, p. 534, pl. 7: figs 1-12 (1991). 

Gomphonema bohemicum ssp. angustiminus E.Reichardt, Iconogr. Diatomol. 8, p. 54, pl. 62: figs 1-15, 18-19, 21-25 

(1999). 

Gomphonema bourbonense E.Reichardt, Nova Hedwigia 65, p. 118, pl. 9, figs 1-15 (1997). 

Gomphonema bozenae Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 68-69, pl. 65: figs 11-15, pl. 121: figs 4-5 

(1996). 

Gomphonema brasiliense ssp. pacificum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 185-186, pl. 50: 

figs 1-6 (1998). 



48 

Gomphonema calcifugum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 6, p. 53 (1999). 

Replaced synonym: Gomphonema olivaceum var. minutissimum Hust., Süßwasserflora von Mitteleuropa 10, p. 378, fig. 720 

(1930). 

Gomphonema camburnii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 113, pl. 159: figs 16-17 (1998). 

Gomphonema capitatum var. macilentum (Kociolek et Stoermer) E.Reichardt in Jahn et al., Lange-Bertalot-Festschrift, p. 

194 (2001) 

Basionym: Gomphonema truncatum var. macilentum Kociolek et Stoermer, Canad. J. Bot. 69, p. 1561, figs 40-41 (1991). 

Gomphonema cathedralis Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 69, pl. 65: figs 1-4 (1996). [Comment: on the 

"Errata" the name was corrected to "Gomphonema cathedrale".] 

Gomphonema clava E.Reichardt in Jahn et al., Lange-Bertalot-Festschrift, p. 214, pl. 12: figs 1-14, pl. 13: figs 1-8 (2001). 

Gomphonema clavatulum E.Reichardt, Iconogr. Diatomol. 8, p. 25, pl. 25: figs 1-23, 27-28 (1999). 

Gomphonema contraturris Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 57-58, pl. 78: figs 2-9, pl. 79: figs 1-5 

(1993). 

Gomphonema costei Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 115, pl. 154: figs 7-12 (1998). 

Gomphonema crassicingulatum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 186-187, pl. 50: figs 7-10 

(1998). 

Gomphonema cuneolus E.Reichardt, Nova Hedwigia 65, p. 123, pl. 11, figs 1-18 (1997). 

Gomphonema cymbelliclinum E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 36, pl. 39: figs 11-14, 24-26, pl. 40: figs 

1-33, pl. 41: figs 10-17 (1999). 

Gomphonema daphnoides E.Reichardt, Iconogr. Diatomol. 8, p. 15-16, pl. 12: figs 1-12 (1999). 

Gomphonema designatum E.Reichardt, Nova Hedwigia 65, p. 120, pl. 10, figs 1-13 (1997). 

Gomphonema distans (Cleve-Euler) Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 6, p. 54 (1999). 

Basionym: Gomphonema lagerheimii var. distans Cleve-Euler, Soc. Scient. Fenn., Com. Biol. 4, p. 72, pl. 5: fig. 120 (1934). 

Gomphonema drutelingense E.Reichardt, Iconogr. Diatomol. 8, p. 38, pl. 39: figs 21-23, pl. 42: figs 1-18 (1999). 

Gomphonema duplipunctatum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 3, p. 88 (1996). 

Replaced synonym: Gomphonema bipunctatum Krasske, nom. illeg., Ber. Deutsch. Bot. Ges. 61, p. 87, fig. 33 (1943); non 

Gomphonema bipunctatum Zimmermann (1915). 

Gomphonema ellipsis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 116, pl. 154: figs 6, 6A (1998). 

Gomphonema equadorianum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 116 (1998). 

Replaced synonym: Gomphonema puiggarianum var. aequatorialis Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26 (2), 

p. 189 (1894). Cleve, Diatomiste 2, p. 101, pl. 7: fig. 1 (1894). 

Gomphonema exiguum var. platypus (Østrup) Lange-Bert., Biblioth. Diatomol. 9, p. 47 (1985). 

Basionym: Gomphonema platypus Østrup, Danske Diatomeer, p. 65, fig. 49 (1910). 

Gomphonema exilissimum (Grunow) Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 70 (1996). 

Basionym: Gomphonema parvulum var. exilissimum Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 25: 

fig. 12 (1880). 

Gomphonema extentum E.Reichardt et Lange-Bert., Ber. Bayer. Bot. Ges. 64, p. 128, pl. 1: figs 16-21, pl. 5: figs 5-8 

(1994). 

Gomphonema frigidum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 6, p. 54, pl. 64: figs 1-8 (1999). 

Gomphonema geisslerae E.Reichardt et Lange-Bert., Nova Hedwigia 65, p. 114, pl. 8, figs 1-10 (1997). 

Gomphonema genkalii Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 6, p. 55-56, pl. 65: figs 1-7 (1999). 

Gomphonema grasmueckii Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 59, pl. 75: figs 1-5, pl. 77: figs 1-14 

(1993). 

Gomphonema grovei var. lingulatum (Hust.) Lange-Bert., Biblioth. Diatomol. 9, p. 47 (1985). 

Basionym: Gomphonema lingulatum Hust., Arch. Hydrobiol. 18, p. 166, pl. 5: fig. 5 (1927). 



49 

Gomphonema halbachii Lange-Bert. et E.Reichardt in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 253-254, pl. 20: figs 157-160 (2000). 

Gomphonema hastatum (Wislouch) Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 60 (1993). 

Basionym: Gomphonema quadripunctatum var. hastata Wislouch, Ber. Deutsch. Bot. Ges. 42, p. 166, figs 5a-c (1924). 

Gomphonema inkadoriae Lange-Bert. et E.Reichardt in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 255, pl. 21: figs 164-172 (2000). 

Gomphonema innocens E.Reichardt, Iconogr. Diatomol. 8, p. 32, pl. 33: figs 1-21, 24-29, 32-34 (1999). 

Gomphonema insigniforme E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 10, pl. 3: figs 1-22, pl. 4: figs 1-7 (1999). 

Gomphonema interpositum E.Reichardt, Iconogr. Diatomol. 8, p. 47-48, pl. 58: figs 1-8, 10-14 (1999). 

Gomphonema inversum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 187-188, pl. 51: figs 1-3 (1998). 

Gomphonema jadwigiae Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 70, pl. 98: figs 11-14 (1996). 

Gomphonema kaweckana Metzeltin, E.Reichardt et Lange-Bert., Iconogr. Diatomol. 4, p. 17-18, pl. 22: figs 6-10, 12-13 

(1996). 

Gomphonema kobayashiae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 117-118, pl. 154: figs 13-16 (1998). 

Gomphonema lacus-vulcani E.Reichardt et Lange-Bert., Nova Hedwigia 65, p. 116, pl. 8, figs 11-33 (1997). 

Gomphonema lateripunctatum E.Reichardt et Lange-Bert., Nova Hedwigia 53, p. 530, pl. 5: figs 1-19; pl. 6: figs 1-3 

(1991). 

Gomphonema laticollum E.Reichardt in Jahn et al., Lange-Bertalot-Festschrift, p. 199, pl. 5: figs 1-14 (2001). 

Gomphonema latipes E.Reichardt in Jahn et al., Lange-Bertalot-Festschrift, p. 196, pl. 3: figs 1-13 (2001). 

Gomphonema leptoproductum Lange-Bert. et Genkal, Iconogr. Diatomol. 6, p. 57, pl. 64: figs 9-15 (1999). [Comment: 

Gomphonema leptoproductum is also documented on pl. 64: fig. 15 although not cited on p. 57 in the 1 st edition, reference 

corrected in the 2 nd edition.] 

Gomphonema lingulatiforme Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 60-61, pl. 83: figs 1-11 (1993). 

[Validation of Gomphonema lingulatiforme Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 422, pl. 87: fig. 11, (1991). As a part of the protologue is cited: 

Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 166: figs 6-11 and pl. 167: 

figs 3-4.] [Comment: In Krammer & Lange-Bertalot (1986) Gomphonema lingulatiforme was documented as "Gomphonema 

grovei var. lingulatum (Hust.) Lange-Bert.".] 

[Basionym of Gomphosphenia lingulatiformis (Lange-Bert. et E.Reichardt) Lange-Bert.] 

Gomphonema lippertii E.Reichardt et Lange-Bert., Iconogr. Diatomol. 8, p. 21, pl. 21: figs 1-15 (1999). 

Gomphonema longilineare E.Reichardt, Iconogr. Diatomol. 8, p. 39, pl. 43: figs 1-15, pl. 44: figs 11-13 (1999). 

Gomphonema maclaughlinii E.Reichardt, Iconogr. Diatomol. 8, p. 22, pl. 22: figs 1-14 (1999). 

Gomphonema makarovae Lange-Bert., Iconogr. Diatomol. 6, p. 58, pl. 63: figs 1-6 (1999). 

Gomphonema malayense var. melanesicum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 188-189, pl. 

52: figs 1-4 (1998). 

Gomphonema micropumilum E.Reichardt, Nova Hedwigia 65, p. 107, pl. 4, figs 1-19 (1997). 

Gomphonema micropus var. aequale (W.Greg.) E.Reichardt, Iconogr. Diatomol. 8, p. 36 (1999). 

Basionym: Gomphonema aequale W.Greg., Quart. J. Microscop. Sci. 4, p. 12, pl. 1: fig. 41 (1856). 

Gomphonema minutiforme Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 61-62, pl. 83: figs 16-20 (1993). 

[Comment: validation of Gomphonema minutiforme Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 


Gomphonema minutum f. curtum (Hust.) Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 62 (1993). 

Basionym: Gomphonema curtum Hust., Arch. Hydrobiol. 40, p. 939, pl. 42: figs 37-43 (1945). 

Gomphonema minutum f. lemanense Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 63 (1993). [Comment: 

validation of Gomphonema minutum f. lemanense Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 410, pl. 81: figs 15-15A (1991).] 



50 

Gomphonema minutum f. pachypus Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 64 (1993). [Validation of 

Gomphonema minutum f. pachypus Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora 

von Mitteleuropa, Bacillariophyceae 2 (4), p. 410, pl. 81: figs 16-19 (1991).] 

Gomphonema minutum f. syriacum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 64 (1993). [Validation of 

Gomphonema minutum f. syriacum Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora 


Gomphonema neoapiculatum Lange-Bert., E.Reichardt et Metzeltin, Iconogr. Diatomol. 5, p. 120, pl. 157: figs 6-9 (1998). 

[Comment: in Metzeltin & Kusber (2001) the authors are erroneously cited as "Metzeltin, E.Reichardt et Lange-Bert.".] 

Gomphonema neobourrellyi Gerd Moser et Lange-Bert., Biblioth. Diatomol. 32, p. 110-111, pl. 45: figs 1-9 (1995). 

Gomphonema neobourrellyi ssp. prolongatum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 189-190, 

pl. 51: figs 8-10 (1998). 

Gomphonema neonasutum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 5, p. 121, pl. 156: figs 1-4 (1998). [Comment: in 

Metzeltin & Kusber (2001) the authors erroneously are cited as "Metzeltin et Lange-Bert".] 

Gomphonema netriviale Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 190-191, pl. 52: figs 5-10 (1998). 

Gomphonema occultum E.Reichardt et Lange-Bert., Nova Hedwigia 53, p. 538, pl. 10: figs 1-13 (1991). 

Gomphonema okavangoanum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 65, pl. 78: fig. 1, pl. 79A: figs 1-5 

(1993). 

Gomphonema olivaceum var. olivaceolacuum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 67, pl. 80: figs 1-8, pl. 

81: figs 3-4, pl. 82: figs 1-4 (1993). [Comment: validation of Gomphonema olivaceum var. olivaceolacuum Lange-Bert. et 

E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 424, 

pl. 88: figs 14-15 (1991).] 

Gomphonema ortalii Lange-Bert. et E.Reichardt in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 257-258, pl. 20: figs 149-156 (2000). 

Gomphonema pala E.Reichardt in Jahn et al., Lange-Bertalot-Festschrift, p. 212 pl. 10: figs 1-22, pl. 11: figs 4-7 (2001). 

Gomphonema paludosum E.Reichardt, Iconogr. Diatomol. 8, p. 26, pl. 26: figs 1-22, pl. 27: figs 1-9 (1999). 

Gomphonema parallelistriatum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 68, fig. 76: 11-16 (1993). [Comment: 

validation of Gomphonema parallelistriatum Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 


Gomphonema parvulius (Lange-Bert. et E.Reichardt) Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 71 (1996). 

Basionym: Gomphonema parvulum var. parvulius Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 70 (1993). 

Gomphonema parvulum f. saprophilum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 69 (1993). [Comment: 

validation of Gomphonema parvulum f. saprophilum Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 400, pl. 76: figs 8-13 (1991). The proptologue includes also 

Krammer & Lange-Bertalot (1991b) pl. 77: figs 5-9, originally on p. 402 documented as "() Gomphonema parvulum var. 

parvulum f. saprophilum Lange-Bert. & E.Reichardt, Umrißvariante ()".] 

Gomphonema parvulum var. parvulius Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 70 (1993). [Validation of 

Gomphonema parvulum var. parvulius Lange-Bert. et E.Reichardt, nom. inval., in Krammer & Lange-Bertalot, 


[Basionym of Gomphonema parvulius (Lange-Bert. et E.Reichardt) Lange-Bert. et E.Reichardt.] 

Gomphonema potaroense Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 122, pl. 146: figs 4-8, pl. 161: figs 8-10 (1998). 

Gomphonema pratense Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 8, p. 18-19, pl. 15: figs 4-7, pl. 16: figs 1-9 (1999). 

Gomphonema procerum E.Reichardt et Lange-Bert., Nova Hedwigia 53, p. 526, pl. 4: figs 1-12 (1991). 

Gomphonema productum (Grunow) Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 71 (1993). 

Basionym: Gomphonema angustatum var. productum Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 24: 

figs 52-55 (1880). 

Gomphonema pseudoaugur Lange-Bert., Arch. Hydrobiol. Suppl. 56,2 (Algol. Stud. 23), p. 213-214, figs 11-16, 79-80 

(1979). 



51 

Gomphonema pseudobohemicum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 71-72, pl. 66: figs 9-10 (1996). [As a 

part of the protologue, Krammer & Lange-Bertalot (1991), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 77: 

figs 16-21 is cited. Comment: in Krammer & Lange-Bertalot (1991b), p. 402, Gomphonema pseudobohemicum is cited as 

"() Gomphonema bohemicum Reichelt et Fricke".] 

Gomphonema pseudoclavatum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 3, p. 92, pl. 40: figs 1-15 (1996). 

Gomphonema pseudopusillum E.Reichardt, Iconogr. Diatomol. 8, p. 48-49, pl. 59: figs 1-15 (1999). 

Gomphonema pseudotenellum Lange-Bert., Biblioth. Diatomol. 9, p. 51, pl. 35: figs 7-12 (1985). [As a part of the 


cited.] [Comment: the diagnosis was emended by E.Reichardt in Nova Hedwigia 65, p. 114 (1997).] 

Gomphonema pumilum var. elegans E.Reichardt et Lange-Bert., Nova Hedwigia 65, p. 103, pl. 1, figs 1-6, pl. 2, figs 1-29, 

pl. 4, figs 20-23 (1997). 

Gomphonema pumilum var. rigidum E.Reichardt et Lange-Bert., Nova Hedwigia 65, p. 105, pl. 1, fig. 7, pl. 3, figs 1-41, 

pl. 4, figs 24-25 (1997). 

Gomphonema punae Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 140, pl. 129: figs 1-14 (2000). 

Gomphonema reichardtii Lange-Bert., Iconogr. Diatomol. 2, p. 72-73, pl. 65: figs 16-19, pl. 121: figs 1-3 (1996). [As a part 

of the protologue, Krammer & Lange-Bertalot (1991), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 78: figs 

20-23 is cited. Comment: in Krammer & Lange-Bertalot (1991b), p. 404, Gomphonema reichardtii is cited as "() 

Gomphonema lagerheimii A.Cleve-Euler (pro parte); () Gomphonema angustatum var. linearis Hust.; () Gomphonema 

angustatum var. cymbelloides Fusey".] 

Gomphonema rhombiciforme Lange-Bert., Iconogr. Diatomol. 5, p. 123 (1998). 

Replaced synonym: Gomphonema occidentale var. abbreviata M. Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 

216: figs 22-23 (1899). 

Gomphonema ricardii var. pachypus Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 32, p. 114, pl. 46: figs 11-12, 10 

(1995). 

Gomphonema riotecense Lange-Bert., E.Reichardt et Metzeltin, Iconogr. Diatomol. 5, p. 123-124, pl. 149: figs 1-6, pl. 150: 

figs 1-5 (1998). 

Gomphonema roehringeri E.Reichardt, Iconogr. Diatomol. 8, p. 55-56, pl. 66: figs 1-12 (1999). 

Gomphonema rosenstockianum Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 71, pl. 76: figs 1-10 (1993). 

Gomphonema salae Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 5, p. 124-125, pl. 157: figs 3-5 (1998). 

Gomphonema scharfii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 141, pl. 130: figs 1-14 (2000). 

Gomphonema scoticollegarum Lange-Bert., Iconogr. Diatomol. 8, p. 12, pl. 6: figs 1-9 (1999). 

Gomphonema spectabilissimum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 127, pl. 146: figs 1-3, pl. 162: figs 3-5 

(1998). 

Gomphonema sphenovertex Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 73-74, pl. 65: figs 22-23 (1996). 

Gomphonema stonei E.Reichardt, Iconogr. Diatomol. 8, p. 16, pl. 13: figs 1-17 (1999). 

Gomphonema subarcticum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 6, p. 59-60, pl. 65: figs 8-20, pl. 66: figs 1-5 

(1999). 

Gomphonema supersedens E.Reichardt, Iconogr. Diatomol. 8, p. 54-55, pl. 63: figs 6-7, pl. 64: figs 1-8, pl. 65: figs 1-9 

(1999). 

Gomphonema tristigmatum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 128, pl. 161: figs 1-7 (1998). 

Gomphonema tumidum (Skvortzov) Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 73 (1993). 

Basionym: Gomphonema quadripunctatum [var. genuina] f. tumida Skvortzov, Proc. Sungaree River Biol. Station 1, p. 27, 

fig. 99 (1928). 

Gomphonema utae Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 8, p. 27, pl. 28: figs 1-37, pl. 29: figs 1-7 (1999). 



52 

Gomphonema variscohercynicum Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 8, p. 37 (1999). 

Replaced synonym: Gomphonema angustatum var. cymbelloides Fusey, Bull. Microscop. Appl. 2. Ser., 1 (2), p. 46, pl. 2: 

fig. 44 (1951). 

Gomphonema vibrioides E.Reichardt et Lange-Bert., Nova Hedwigia 53, p. 524, pl. 3: figs 1-9 (1991). 

Gomphonema wiltschkorum Lange-Bert., Iconogr. Diatomol. 6, p. 61, pl. 67: figs 12-22 (1999). [Comment: Gomphonema 

wiltschkorum is also documented on 67: figs 12-16 although not cited in the 1 st edition, reference corrected in the 2 nd 

edition.] 

Gomphonema yucatanense Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 131-132, pl. 146: figs 9-10 (1998). 

Gomphonema zellense E.Reichardt, Iconogr. Diatomol. 8, p. 11, pl. 5: figs 1-11 (1999). 

GOMPHONEMOPSIS Medlin 


Gomphonemopsis lindae Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 18, pl. 54: figs 36, 39-41, pl. 77: 

figs 9-10, pl. 79: fig. 3 (1996). 

Gomphonemopsis obscura (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 91 (1996). 

Basionym: Gomphonema obscurum Krasske, Arch. Hydrobiol. 35, p. 406, pl. 12: figs 11-13 (1939). 

GOMPHOSEPTATUM Medlin 


Gomphoseptatum pseudoseptatum (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 222 (2000). 

Basionym: Gomphonema pseudoseptatum Giffen, Bot. Mar. 13, p. 92, figs 29-32 (1970). 

GOMPHOSPHENIA Lange-Bert., Nova Hedwigia 60, p. 242-243 (1995). [Typus generis: Gomphosphenia lingulatiformis 

(Lange-Bert. et E.Reichardt) Lange-Bert., Nova Hedwigia 60, p. 242 (1995).] 

GOMPHOSPHENIA subgen. COSTERICARDIA [see COSTERICARDIA]. 

GOMPHOSPHENIA subgen. RHOICOSPHENULA [see RHOICOSPHENULA]. 

Gomphosphenia grovei (M.Schmidt) Lange-Bert. var. grovei, Nova Hedwigia 60, p. 243 (1995). 

Basionym: Gomphonema grovei M.Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 214: figs 13-18 (1899). 

Gomphosphenia grovei var. lingulata (Hust.) Lange-Bert., Nova Hedwigia 60, p. 243 (1995). 

Basionym: Gomphonema ligulatum Hust., Arch. Hydrobiol. 18, p. 166, pl. 5: fig. 5 (1927). 

Gomphosphenia holmquistii (Foged) Lange-Bert., Nova Hedwigia 60, p. 244 (1995). 

Basionym: Gomphonema holmquistii Foged, Nova Hedwigia 16, p. 8, pl. 3: fig. 3 (1968). 

Gomphosphenia holmquistii var. alaskaensis (Foged) Lange-Bert., Nova Hedwigia 60, p. 244 (1995). 

Basionym: Gomphonema holmquistii var. alaskaensis Foged, Nova Hedwigia 16, p. 9, figs 3-4 (1968). 

Gomphosphenia lingulatiformis (Lange-Bert. et E.Reichardt) Lange-Bert., Nova Hedwigia 60, p. 242 (1995). 

Basionym: Gomphonema lingulatiforme Lange-Bert. et E.Reichardt, Biblioth. Diatomol. 27, p. 60, pl. 83: figs 1-11 (1993). 

[Gomphosphenia lingulatiformis is the type of the name of the genus GOMPHOSPHENIA.] 

Gomphosphenia paradoxa Lange-Bert., Nova Hedwigia 60, p. 246, 248, pl. 1: figs 1-18, pl. 2: figs 1-6, pl. 3: figs 1-7 

(1995). [Gomphosphenia paradoxa is the ype of the name of the subgenus RHOICOSPHENULA Lange-Bert.] 

Gomphosphenia pfannkuchae (Cholnoky) Lange-Bert., Nova Hedwigia 60, p. 244 (1995). 

Basionym: Gomphonema pfannkuchae Cholnoky, Nova Hedwigia Beih. 21, p. 24, figs 48-53 (1966). 

Gomphosphenia reicheltii (M.Schmidt) Lange-Bert., Nova Hedwigia 60, p. 244 (1995). 

Basionym: Gomphonema reicheltii M.Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 214: figs 19-22 (1899). 

Gomphosphenia tackei (Hust.) Lange-Bert., Nova Hedwigia 60, p. 244 (1995). 

Basionym: Gomphonema tackei Hust., Abh. Naturwiss. Vereine Bremen 42, p. 205, figs 16-18 (1942). 

Gomphosphenia tenerrima (Hust.) E.Reichardt, Iconogr. Diatomol. 8, p. 57 (1999). 

Basionym: Gomphonema tenerrimum Hust., Arch. Hydrobiol. Suppl. 15, p. 444, pl. 27: figs 23-24 (1938). 

[Taxonomic synonym of Gomphosphenia tenerrima (Hust.) E.Reichardt: Gomphonema wulasiense Foged, according to 

Reichardt, Iconogr. Diatomol. 8, p. 57 (1999).] 



53 

GRUNOWINITZSCHIA Lange-Bert. et U.Rumrich (subgenus within NITZSCHIA), Iconogr. Diatomol. 9, p. 191 (2000). 

[Typus subgeneris: Nitzschia andicola Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 193 (2000).] 

GYROSIGMA Hassall 


Gyrosigma gondwana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 194-195, pl. 41: figs 1-2 (1998). 

Gyrosigma moresbyana E.Reichardt, Nova Hedwigia 47, p. 92, pl. 5: figs 1-4 (1988). 

HANTZSCHIA Grunow 


Hantzschia abundans Lange-Bert., Biblioth. Diatomol. 27, p. 75, pl. 85: figs 12-18, pl. 89: figs 1-6, pl. 90: figs 1-6, pl. 92: 

fig. 1 (1993). 

Hantzschia amphilepta (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 76 (1993). 

Basionym: Hantzschia amphioxys var. amphilepta Grunow, J. Microscop. 3, p. 397, fig. 8 (1880). 

Hantzschia barckhausenii Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 75, pl. 66: figs 16-18, pl. 67: figs 1-2 (1996). 

Hantzschia brunneri Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 132, pl. 206: fig. 1 (1998). 

Hantzschia compacta (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 62 (1999). 

Basionym: Hantzschia amphioxys var. compacta Hust. in S. Hedin, Southern Tibet 6 (3) Bot. Stockholm, p. 145, fig. 42 

(1922). 

Hantzschia elegantula (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 362 (2000). 

Basionym: Hantzschia amphioxys var. elegantula Østrup, Danske Diatoméer, p. 143, pl. 4: fig. 94 (1910). 

Hantzschia eximia Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 195-196, pl. 59: figs 3-5 (1998). 

Hantzschia giessiana Lange-Bert. et U.Rumrich, Biblioth. Diatomol. 27, p. 80, pl. 94: figs 1-6, pl. 95: figs 1-6 (1993). 

Hantzschia graciosa Lange-Bert., Biblioth. Diatomol. 27, p. 81, pl. 101: figs 1-5, pl. 102: figs 1-6 (1993). 

Hantzschia graciosa ssp. neotropica Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 133, pl. 206: figs 2-5 (1998). 

Hantzschia hyperborea (Grunow) Lange-Bert., Biblioth. Diatomol. 27, p. 83 (1993). 

Basionym: Hantzschia amphioxys var. hyperborea Grunow, Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 48, p. 99, 

pl. 1: fig. 59 (1884). 

Hantzschia kaiseri Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 363, 427-428, pl. 175: figs 11-15; pl. 219: 

figs 1-5 (2000). 

Hantzschia pannonica Lange-Bert., Biblioth. Diatomol. 27, p. 83-84, pl. 97: fig. 6, pl. 98: figs 1-6, pl. 99: figs 1-6, pl. 100: 

fig. 1 (1993). 

Hantzschia subrupestris Lange-Bert., Biblioth. Diatomol. 27, p. 85 (1993). 

Replaced synonym: Hantzschia amphioxys [var. rupestris] f. capitellata Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, 

pl. 329: fig. 9 (1921). 

Hantzschia vivacior Lange-Bert., Biblioth. Diatomol. 27, p. 86, pl. 104: figs 1-6, pl. 105: figs 1-7, pl. 106: figs 1-6 (1993). 

HASLEA Simonsen 


Haslea britannica (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 223 (2000). 

Basionym: Navicula britannica Hust. in Hustedt & Aleem, J. Mar. Biol. Assoc. UK. 30, p. 184, fig. 1C (1951). 

Haslea spicula (Hickie) Lange-Bert., Arch. Protistenk. 148, p. 75 (1997). 

Basionym: Stauroneis spicula Hickie, Monthly Microscop. J. 12, p. 290 (1874). 

HIPPODONTA Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 249 (1996). [Typus generis: Hippodonta 

lueneburgensis (Grunow) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 262 (1996).] 

Hippodonta arctica (R.M.Patrick et Freese) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 252 (1996). 

Basionym: Navicula hungarica var. arctica R.M.Patrick et Freese, Proc. Acad. Nat. Sci. Philadelphia 112, p. 207, pl. 2: fig. 

16 (1961). 

Hippodonta arkonensis Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 252, pl. 4: figs 11-18 (1996). 



54 

Hippodonta avittata (Cholnoky) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 253 (1996). 

Basionym: Navicula subcostulata var. avittata Cholnoky, Oesterr. Bot. Z. 106, p. 51, figs 265-269 (1959). 

Hippodonta caotica Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 225, 428, pl. 131: figs 25-29 (2000). 

Hippodonta capitata (Ehrenb.) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 254 (1996). 

Basionym: Navicula capitata Ehrenb., Infusionsthierchen, p. 185, pl. 13: fig. 20 (1838). 

Hippodonta costulata (Grunow) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 254 (1996). 

Basionym: Navicula costulata Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 27 (1880). 

Hippodonta costulatiformis Lange-Bert., Metzeltin et Witkowski, p. 254-255, pl. 1: figs 27-28, pl. 4: figs 21-22 (1996). 

Hippodonta coxiae Lange-Bert., Diatoms of Europe 2, p. 221-222, pl. 76: figs 29-33, pl. 77: fig. 15 (2001). 

Hippodonta geocollegarum Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 255-256, pl. 4: figs 1-5 (1996). 

Hippodonta germainii Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 256-257, pl. 2: figs 6-11, pl. 3: fig. 6 

(1996). 

Hippodonta grunowii Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 257-258, pl. 1: fig. 41(1996). 

Hippodonta hungarica (Grunow) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 259 (1996). 

Basionym: Navicula hungarica Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 10, p. 539, pl. 1: fig. 30 (1860). 

Hippodonta kaiseri Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 259, pl. 2: figs 13-19, pl. 4: fig. 25 

(1996). 

Hippodonta kissii Lange-Bert., Diatoms of Europe 2, p. 222, pl. 78: figs 1-10 (2001). 

Hippodonta kociolekii Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 260, pl. 3: figs 7-7b (1996). 

Hippodonta lesmonensis (Hust.) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 261 (1996). 

Basionym: Navicula lesmonensis Hust., Abh. Naturwiss. Vereine Bremen 34, p. 293, figs 36-37 (1957). 

Hippodonta linearis (Østrup) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 261 (1996). 

Basionym: Navicula linearis Østrup, Danske Diatomeer, p. 79, pl. 2: fig. 53 (1910). 

[Comment: In Kusber & Metzeltin (2001), p. 604 the basionym was erroneously cited as "Navicula hungarica var. linearis 

Østrup", Danske Diatomeer, p. 79, pl. 2: fig. 53 (1910).] 

Hippodonta lueneburgensis (Grunow) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 249 (1996). 

Basionym: Navicula hungarica var. lueneburgensis Grunow in Mojsisovics et Neumayr, Beiträge zur Paläontologie 

Österreich-Ungarns und des Orients Bd. 2 (4), p. 156, pl. 30: figs 43-44 (1882). [Hippodonta lueneburgensis is the type of 

the name of the genus HIPPODONTA.] 

Hippodonta neglecta Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 262, pl. 1: figs 8-15, pl. 3: fig. 2, pl. 4: 

fig. 10 (1996). [Comment: Hippodonta neglecta is not documented on pl. 2: fig. 2 as cited on p. 262] 

Hippodonta pseudacceptata (H.Kobayasi) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 263 (1996). 

Basionym: Navicula pseudacceptata H.Kobayasi, Diatom 2, p. 96, figs 1-4, 7-12 (1986). 

Hippodonta pseudopinnularia Lange-Bert., Diatoms of Europe 2, p. 222, pl. 77: figs 1-8 (2001). 

Hippodonta rostrata (Hust.) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 263 (1996). 

Basionym: Navicula subcostulata var. rostrata Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 402: figs 65 (1936). 

Hippodonta ruthnielseniae Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 264, pl. 3: figs 8-15 (1996). 

[Comment: Hippodonta ruthnielseniae on p. 272 (caption of pl. 3: figs 8-15) is misprinted as "Hippodonta subtilissima".] 

Hippodonta subcostulata (Hust.) Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 264 (1996). 

Basionym: Navicula subcostulata Hust., Abh. Vortr. Bremer Wiss. Ges. 8-9, p. 386, fig. 13 (1934). 

Hippodonta subtilissima Lange-Bert., Metzeltin et Witkowski, Iconogr. Diatomol. 4, p. 265, pl. 3: figs 16-23 (1996). 

[Comment: Hippodonta subtilissima on p. 272 (caption of pl. 3: figs 16-23) is misprinted as "Hippodonta ruthnielseniae".] 

HYALINELLA Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 60, 428 (2000). [Typus generis: Hyalinella 

lateripunctata Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 60, 429 (2000).] 

Hyalinella lateripunctata Witkowski, Lange-Bert. et Metzeltin Iconogr. Diatomol. 7, p. 60, 429, pl. 32: figs 6-18 (2000). 

[Hyalinella lateripunctata is the type of the name of the genus HYALINELLA Witkowski, Lange-Bert. et Metzeltin.] 



55 

HYALOSIRA Kütz. 


Hyalosira delicatula var. gibbosa (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 61 (2000). 

Basionym: Striatella delicatula var. gibbosa Østrup, Bot. Tidsskr. 26, p. 276, pl. 1: fig. 13 (1904). 

HYGROPETRA Krammer et Lange-Bert., Diatoms of Europe 1, p. 206, 240 (2000). [Typus generis: Hygropetra 

balfouriana (Grunow ex Cleve) Krammer et Lange-Bert., Diatoms of Europe 1, p. 207 (2000).] 

Hygropetra balfouriana (Grunow ex Cleve) Krammer et Lange-Bert., Diatoms of Europe 1, p. 207 (2000). 

Basionym: Pinnularia balfouriana Grunow ex Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26, p. 80, pl. 1: fig. 18 

(1895). [Hygropetra balfouriana is the type of the name of the genus HYGROPETRA.] 

Hyalosira parietina Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 62, 429, pl. 21: figs 18-20 (2000). 

Hygropetra elongata Krammer et Lange-Bert., Diatoms of Europe 1, p. 207, 240, pl. 216: figs 10-13, 18 (2000). [Comment: 

on p. 692 (= caption of pl. 216: figs 10-13) the authors are cited as "Lange-Bert. et Krammer", Hygropetra elongata on p. 

692 (= caption of pl. 216: fig. 18) as "Hygropetra balfouriana (Grunow ex Cleve) Krammer et Lange-Bert.".] 

KOBAYASIA Lange-Bert., nom. illeg., Iconogr. Diatomol. 4, p. 279. [Intended type of the name of the genus: Kobayasia 

bicuneus Lange-Bert. Iconogr. Diatomol. 4, p. 279 (1996). [Comment: KOBAYASIA Lange-Bert. is a later homonym of 

KOBAYASIA Imai et A. Kkawamura, hence illegitimate. See: KOBAYASIELLA Lange-Bert. 1999.] 

Kobayasia bicuneus Lange-Bert., Iconogr. Diatomol. 4, p. 280-281, pl. 1: fig. 1-10 (1996) [Comment: Kobayasia bicuneus 

was the intended type of the name of the genus KOBAYASIA Lange-Bert., nom. illeg., see Kobayasiella bicuneus.] 

Kobayasia grandis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 134, pl. 94: figs 1-5 (1998). 

[Basionym of Kobayasiella grandis (Metzeltin et Lange-Bert.) Lange-Bert.] 

Kobayasia jaagii (F.Meister) Lange-Bert., Iconogr. Diatomol. 4, p. 281 (1996). 

Basionym: Navicula jaagii F.Meister, Ber. Schweiz. Bot. Ges. 44, p. 89, fig. 9 (1935). 

[Nomenclatural synonym of Kobayasiella jaagii (F.Meister) Lange-Bert.] 

Kobayasia krasskei Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 135, pl. 94: figs 13-15 (1998). 

[Basionym of Kobayasiella krasskei (Metzeltin et Lange-Bert.) Lange-Bert.] 

Kobayasia madumensis (E.G.Jørgensen) Lange-Bert., Iconogr. Diatomol. 4, p. 282 (1996). 

Basionym: Navicula madumensis E.G.Jørgensen, Kongel. Danske Vidensk.-Selsk. Skr. 5, p. 60, pl. 2: fig. 26 (1948). 

[Nomenclatural synonym of Kobayasiella madumensis (E.G.Jørgensen) Lange-Bert.] 

Kobayasia micropunctata (H.Germ.) Lange-Bert., Iconogr. Diatomol. 4, p. 282 (1996). 

Basionym: Navicula subtilissima var. micropunctata H.Germ., Flore Diatomées eaux douces et saumâtres, Boubée, Paris, p. 

234, pl. 169: figs 4-5 (1981). 

[≡ Navicula micropunctata (H.Germ.) H.Kobayasi et Nagumo, Bot. Mag. Tokyo 101: p 247 (1988).] 

[Nomenclatural synonym of Kobayasiella micropunctata (H.Germ.) Lange-Bert.] 

Kobayasia neocaledonica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 196-197, pl. 29: figs 9-14 

(1998). 

[Basionym of Kobayasiella neocaledonica (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert.] 

Kobayasia okadae (Skvortzov) Lange-Bert., Iconogr. Diatomol. 4, p. 282 (1996). 

Basionym: Anomoeoneis okadae Skvortzov, J. Jap. Bot. 14, p. 59, pl. 1: figs 37-38 (1938). 

[Nomenclatural synonym of Kobayasiella okadae (Skvortzov) Lange-Bert.] 

Kobayasia parasubtilissima (H.Kobayasi et Nagumo) Lange-Bert., Iconogr. Diatomol. 4, p. 282 (1996). 

Basionym: Navicula parasubtilissima H.Kobayasi et Nagumo, Bot. Mag. Tokyo 101, p. 245, figs 19-37 (1988). 

[Nomenclatural synonym of Kobayasiella parasubtilissima (H.Kobayasi et Nagumo) Lange-Bert.] 

Kobayasia pseudostauron (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 4, p. 282 (1996). 

Basionym: Naviculadicta pseudostauron Lange-Bert., Iconogr. Diatomol. 2, p. 86, pl. 35: figs 11-18; pl. 117: figs 1-2 

(1996). 

[Nomenclatural synonym of Kobayasiella pseudostauron (Lange-Bert.) Lange-Bert.] 

Kobayasia saxicola (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 42 

(1998). 

Basionym: Navicula saxicola Manguin ex Kociolek et Reviers, Cryptog. Algol. 17, p. 204 (1996). 

[Nomenclatural synonym of Kobayasiella saxicola (Manguin ex Kociolek et Reviers) Lange-Bert.] 



56 

Kobayasia subtilissima (Cleve) Lange-Bert., Iconogr. Diatomol. 4, p. 283 (1996). 

Basionym: Navicula subtilissima Cleve, Acta Soc. Fauna Fl. Fenn. 8, p. 37, pl. 2: fig. 35 (1891). 

[Nomenclatural synonym of Kobayasiella subtilissima (Cleve) Lange-Bert.] 

KOBAYASIELLA Lange-Bert., Iconogr. Diatomol. 6, p. 266 (1999). [Typus generis: Kobayasiella bicuneus (Lange-Bert.) 

Lange-Bert. Iconogr. Diatomol. 6, p. 266 (1999). Comment: new name for KOBAYASIA Lange-Bert. 1996, nom. illeg. In 

Iconogr. Diatomol. 6 (2 nd edition) on p. 272 (1999) again marked as new.] 

Kobayasiella bicuneus (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 266 (1999). 

Basionym: Kobayasia bicuneus Lange-Bert., Iconogr. Diatomol. 4, p. 280, figs 1-10 (1996). [Kobayasiella bicuneus is the 

type of the name of the genus KOBAYASIELLA.] 

Kobayasiella grandis (Metzeltin et Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 266 (1999). 

Basionym: Kobayasia grandis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 134, pl. 94: figs 1-5 (1998). 

Kobayasiella jaagii (F.Meister) Lange-Bert., Iconogr. Diatomol. 6, p. 266-267 (1999). 

Basionym: Navicula jaagii F.Meister, Ber. Schweiz. Bot. Ges. 44, p. 89, fig. 9 (1935). 

Kobayasiella krasskei (Metzeltin et Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 267 (1999). 

Basionym: Kobayasia krasskei Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 135, pl. 94: figs 13-15 (1998). 

Kobayasiella madumensis (E.G.Jørgensen) Lange-Bert., Iconogr. Diatomol. 6, p. 267 (1999). 

Basionym: Navicula madumensis E.G.Jørgensen, Kongel. Danske Vidensk.-Selsk. Skr. 5, p. 60, pl. 2: fig. 26 (1948). 

Kobayasiella micropunctata (H.Germ.) Lange-Bert., Iconogr. Diatomol. 6, p. 267 (1999). 

Basionym: Navicula subtilissima var. micropunctata H.Germ., Flore Diatomées eaux douces et saumâtres, Boubée, Paris, p. 

234, pl. 169: figs 4-5 (1981). 

[≡ Navicula micropunctata (H.Germ.) H.Kobayasi et Nagumo, Bot. Mag. Tokyo 101: p 247 (1988).] 

[Comment: In Iconogr. Diatomol. 6 (1999), p. 267 the basionym is given as "Navicula micropunctata H.Germ.".] 

Kobayasiella neocaledonica (Gerd Moser, Lange-Bert. et Metzeltin) Lange-Bert., Iconogr. Diatomol. 6, p. 267 (1999). 

Basionym: Kobayasia neocaledonica Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 196, pl. 29: figs 9-14 

(1998). 

Kobayasiella okadae (Skvortzov) Lange-Bert., Iconogr. Diatomol. 6, p. 267 (1999). 

Basionym: Anomoeoneis okadae Skvortzov, J. Jap. Bot. 14, p. 59, pl. 1: figs 37-38 (1938). 

Kobayasiella parasubtilissima (H.Kobayasi et Nagumo) Lange-Bert., Iconogr. Diatomol. 6, p. 268 (1999). 

Basionym: Navicula parasubtilissima H.Kobayasi et Nagumo, Bot. Mag. Tokyo 101, p. 245, figs 19-37 (1998). 

Kobayasiella pseudostauron (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 268 (1999). 

Basionym: Naviculadicta pseudostauron Lange-Bert., Iconogr. Diatomol. 2, p. 86, pl. 35: figs 11-18 (1996). 

Kobayasiella pseudosubtilissima (Manguin) Lange-Bert., Iconogr. Diatomol. 6, p. 268 (1999). 

Basionym: Navicula pseudosubtilissima Manguin in Bourrelly et Manguin, Algues de la Guadeloupe. Centre Nat. Rech. Sci. 

Paris, p. 64, pl. 3: fig. 71 (1952). 

Kobayasiella saxicola (Manguin ex Kociolek et Reviers) Lange-Bert., Iconogr. Diatomol. 6, p. 268 (1999). 

Basionym: Navicula saxicola Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 204 (1996). 

[≡ Kobayasia saxicola (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin.] 

Kobayasiella subtilissima (Cleve) Lange-Bert., Iconogr. Diatomol. 6, p. 268 (1999). 

Basionym: Navicula subtilissima Cleve, Acta Soc. Fauna Fl. Fenn. 8, p. 37, pl. 2: fig. 35 (1891). 

KRASSKELLA R.Ross et P.A.Sims 


Krasskella manguinii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 144, pl. 90: figs 4-5, 6 (2000). 

KURPISZIA Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 227, 429 (2000). [Typus generis: Kurpiszia 

semiaperta (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 228 (2000).] 

Kurpiszia kryokonites (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 227 (2000). 

Basionym: Navicula kryokonites Cleve, Vega Expedit. Vetensk. Iakttag. 3, p. 473, pl. 37: fig. 44 (1883). 

Kurpiszia semiaperta (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 228 (2000). 

Basionym: Navicula semiaperta Hust., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 401: figs 14-16 (1936). 



57 

Kurpiszia subprotracta (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 228 (2000). 

Basionym: Navicula kryokonites var. subprotracta Cleve, Vega Expedit. Vetensk. Iakttag. 3, p. 473, pl. 37: fig. 46 (1883). 

LECOHUIA Lange-Bert., Iconogr. Diatomol. 9, p. 146 (2000). [Typus generis: Lecohuia riverae Lange-Bert. et U.Rumrich, 

Iconogr. Diatomol. 9, p. 147 (2000).] 

Lecohuia corrugata (Manguin) Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 148 (2000). 

Basionym: Navicula corrugata Manguin, Mém. Inst. Sci. Madagaskar, Ser. B 4, p. 27, pl. 3: figs 32a, b (1954). 

Lecohuia geniculata (H.Germ.) Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 148 (2000). 

Basionym: Navicula geniculata H.Germ., Bull. Soc. Française Microscopie 6, p. 13, pl. 1: fig. 9 (1937). 

Lecohuia martinii (Krasske) Lange-Bert. et U. Rumrich, Iconogr. Diatomol. 9, p. 148 (2000). 

Basionym: Navicula martinii Krasske, Arch. Hydrobiol. 35, p. 391, pl. 11: fig. 30 (1939). 

Lecohuia riverae Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 147, pl. 64: figs 3-10 (2000). [Lecohuia riverae is the 

type of the name of the genus LECOHUIA Lange-Bert.] 

LUNELLA P.Snoeijs 


[http://128.32.109.44/cgi-bin/porp_cgi.plLUNELLAPSNOEIJS1996] 

Lunella ghalebii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 229, 430, pl. 163: figs 5-10 (2000). 

LUTICOLA D.G.Mann 


[http://128.32.109.44/cgi-bin/porp_cgi.plLUTICOLADGMANN1990]. 

Luticola acidoclinata Lange-Bert., Iconogr. Diatomol. 2, p. 76, pl. 24: figs 24-26, pl. 104: figs 10-16 (1996). 

Luticola clagesii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 197-198, pl. 30: figs 5-9 (1998). 

Luticola costei Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 136, pl. 87: fig. 13 (1998). [Comment: Luticola costei is 

not documented on pl. 87: fig. 12 as cited on p. 136; caption of pl. 87 on p. 408 is correct.] 

Luticola dapaloides (Frenguelli) Lange-Bert., Iconogr. Diatomol. 5, p. 137 (1998). 

Basionym: Navicula dapaloides Frenguelli, Rev. Mus. La Plata N.S. 8, p. 78, pl. 1: figs 20-21 (1953). 

Luticola frenguellii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 137, pl. 88: figs 1-6 (1998). 

Luticola incoactoides Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 149, pl. 61: figs 1-4 (2000). 

Luticola kraeuselii (Cholnoky) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 139 (1998). 

Basionym: Navicula kraeuselii Cholnoky, Bot. Not., p. 284, figs 79-80 (1954). 

Luticola mollis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 149-150, pl. 61: figs 5-6 (2000). 

Luticola tomesii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p 198-199, pl. 30: figs 1-4 (1998). 

LYRELLA N.I.Karayeva 


Lyrella david-mannii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 232 (2000). 

Replaced synonym: Navicula lyra var. elliptica A.W.F. Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, 

pl. 2: figs 29, 34 (January 1874); Jahresber. Komm. Unters. der Meere in Kiel 2, p. 89, pl. 1: figs 35, 38 (1874). 

Lyrella fogedii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 233, 430, pl. 99: figs 4-7, pl. 100: figs 1-3 

(2000). 

Lyrella opuntioides (Simonsen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 235 (2000). 

Basionym: Navicula opuntioides Simonsen, Kieler Meeresforsch. 15, p. 79, pl. 12: fig. 1 (1959). 

Lyrella sulcifera (Hust.) Witkowski, Biblioth. Diatomol. 28, p. 139 (1994). 

Basionym: Navicula sulcifera Hust., Bull. Duke Univ. Mar. Sta. 6, p. 24, pl. 8: fig. 1 (1955). 

MASTOGLOIA Thwaites 


Mastogloia rectirobusta Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 43 (1998). 

Replaced name: Mastogloia malayensis var. robusta Manguin ex Kociolek et Reviers, Cryptog. Algol. 17, p. 199 (1996). 



58 

Mastogloia urveae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 263, 431, pl. 84: figs 22-23 (2000). 

MAYAMAEA Lange-Bert., Arch. Protistenk 148, p. 71 (1997). [Typus generis: Mayamaea atomus (Kütz.) Lange-Bert., 

Arch. Protistenk 148, p. 71 (1997). ] 

Mayamaea agrestis (Hust.) Lange-Bert., Diatoms of Europe 2, p. 134, (2001). 

Basionym: Navicula agrestis Hust., Arch. Hydrobiol. Suppl. 15, p. 246, pl. 20: figs 21-22 (1937). 

[- Navicula agrestis Hust., nom. inval., in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 401: figs 27-29 (1936).] 

Mayamaea aliena (Krasske) Lange-Bert., Diatoms of Europe 2, p. 134, (2001). 

Basionym: Navicula aliena Krasske, Arch. Hydrobiol. 33, p. 529, figs 19-20 (1938). 

Mayamaea arida (W.Bock) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula arida W.Bock, Nova Hedwigia 5, p. 223, fig. 1D (1963). 

Mayamaea asellus (Weinhold) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula asellus Weinhold ex Hust., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 400: figs 1-5 (1934). 

Mayamaea atomus (Kütz.) Lange-Bert. var. atomus, Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Amphora atomus Kützing, Kieselschalige Bacillarien, p. 108, pl. 30: fig. 70 (1844). [Mayamaea atomus is the 

type of the name of the genus MAYAMAEA..] 

Mayamaea atomus var. alcimonica (E.Reichardt) E.Reichardt, Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula atomus var. alcimonica E.Reichardt, Biblioth. Diatomol. 6, p. 39-40, pl. 12: figs 10-14, pl. 30: figs 1-2 

(1984). 

Mayamaea atomus var. permitis (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula permitis Hust., Arch. Hydrobiol. 40, p. 919, pl. 41: figs 8-9 (1945). 

[≡ Navicula atomus var. permitis (Hust.) Lange-Bert.] 

Mayamaea destrica (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula destrica Hust., Ber. Deutsch. Bot. Ges. 60, p. 64, fig. 22 (1942). 

Mayamaea excelsa (Krasske) Lange-Bert., Arch. Protistenk. 148, p. 72. 

Basionym: Navicula excelsa Krasske, Bot. Arch. 3, p. 51, pl. 2: fig. 33 (1925). 

[≡ Navicula atomus var. excelsa (Krasske) Lange-Bert.] 

Mayamaea fossalis (Krasske) Lange-Bert. var. fossalis, Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula fossalis Krasske, Bot. Arch. 27, p. 354, fig. 10 (1929). 

Mayamaea fossalis var. obsidialis (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula obsidialis Hust., Ber. Deutsch. Bot. Ges. 60, p. 68, figs 1-3 (1942). 

[– Navicula fossalis var. obsidialis (Hust.) Lange-Bert., nom. inval., Biblioth. Diatomol. 9, p. 69 (1985).] 

Mayamaea fossaloides (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula fossaloides Hust., Abh. Naturwiss. Vereine Bremen 34, p. 275, fig. 17 (1957). 

Mayamaea lacunolaciniata (Lange-Bert. et Bonik) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula lacunolaciniata Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p. 313, figs 12-13, 

34 (1976). 

Mayamaea muraliformis (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula muraliformis Hust., Arch. Mikrobiol. 14, p. 440, fig. 5 (1949). 

Mayamaea nolensoides (W.Bock) Lange-Bert., Diatoms of Europe 2, p. 140, (2001). 

Basionym: Navicula nolensoides W.Bock, Nova Hedwigia Beih. 31, p. 424, pl. 1: figs 16-17, (1970) 

Mayamaea recondita (Hust.) Lange-Bert., Arch. Protistenk. 148, p. 72 (1997). 

Basionym: Navicula recondita Hust., A. Schmidt’s Atlas der Diatomaceenkunde, Atlas, pl. 400: figs 6-9 (1934). 

[≡ Navicula atomus var. recondita (Hust.) Lange-Bert.] 

MICROCOSTATUS J.R.Johans. et J.C.Sray 


[http://128.32.109.44/cgi-bin/porp_cgi.plMICROCOSTATUSJRJOHANSENJCSRAY1998]. 

Microcostatus andinus Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 151, pl. 76: figs 1-10 (2000). 

Microcostatus egregius (Hust.) Lange-Bert., Iconogr. Diatomol. 6 (2nd edition), p. 290 (1999). 

Basionym: Navicula egregia Hust., Ber. Deutsch. Bot. Ges. 60, p. 64, fig. 26 (1942). 



59 

Microcostatus kuelbsii (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6 (2nd edition), p. 290 (1999). 

Basionym: Navicula kuelbsii Lange-Bert., Biblioth. Diatomol. 9, p. 77, pl. 25: figs 4-7 (1985). 

Microcostatus maceria (Schim.) Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Basionym: Navicula maceria Schim., Nova Hedwigia 30, p. 586, pl. 11: figs 2, 9 (1978). 

[– Microcostatus maceria (Schim.) Lange-Bert., nom. inval., Iconogr. Diatomol. 6 (2nd edition), p. 290 (1999).] 

Microcostatus naumannii (Hust.) Lange-Bert., Iconogr. Diatomol. 6 (2nd edition), p. 291 (1999). 

Basionym: Navicula naumannii Hust., Arch. Hydrobiol. 39, p. 115, figs 22-24 (1942). 

[Comment: Erroneously the epitheton was misspelled as "naumanii".] 

Microcostatus ornicollegarum Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 10, pl. 16: figs 8-14 (2001). 

Microcostatus polonarum Lange-Bert., Brenesia 55-56, p. 11, pl. 15: figs 1-8 (2001). 

Microcostatus pseudomuralis (Hust.) Lange-Bert., Iconogr. Diatomol. 6 (2nd edition), p. 291 (1999). 

Basionym: Navicula pseudomuralis Hust., Arch. Hydrobiol. 15, p. 245, pl. 19: figs 25-27 (1937). 

[– Navicula pseudomuralis Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 401: figs 64-67 (1936).] 

Microcostatus stapputana Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 12, pl. 16: figs 1-7 (2001). 

MIOSIRA Krammer, Lange-Bert. et W.Schiller, Paläontol. Z. 71, p. 10 (1997). [Typus generis: Miosira rhoenana 

Krammer, Lange-Bert. et W.Schiller, Paläontol. Z. 71, p. 10, 17 (1997).] 

Miosira rhoenana Krammer, Lange-Bert. et W.Schiller, Paläontol. Z. 71, p. 10, 17, figs 1-27, 29 (1997). [Miosira rhoenana 

is the type of the name of the genus MIOSIRA Krammer, Lange-Bert. et W.Schiller.] 

MUELLERIA Van Heurck, nom. rej. 


Muelleria islandica (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 63 (1999). 

Basionym: Navicula islandica Østrup, Botany of Iceland 2, part 1, no. 5, p. 26, pl. 3: fig. 36 (1918), non Navicula islandica 

Grunow in Cleve et Möller, Diatoms, collection of 324 slides, no. 262 (1881). 

NAVICELLA Krammer, Biblioth. Diatomol. 36, p. 157, (1997). [Typus generis: Cymbella pusilla Grunow, A. Schmidt’s 

Atlas der Diatomaceenkunde, pl. 9: figs 36-37 (1875).] 

NAVICULA Bory 


NAVICULA sect. ALINEA [see ALINEA]. 

NAVICULA subsect. NAVISANTIQUA [see NAVISANTIQUA]. 

Navicula acidobionta Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 160-161, pl. 98: figs 28-44, pl. 99: figs 1-10 

(1989). 

[Comment: lacking a type and a Latin diagnosis, Navicula acidobionta Lange-Bert. was not validly published. Validly 

published as Stauroneis acidobionta Lange-Bert. et Wydrzycka.] 

Navicula agatkae Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 7, p. 265, 431, pl. 146: figs 1-8 (2000). 

Navicula aggerica E.Reichardt, Ber. Bayer. Bot. Ges. 53, p. 101, pl. 1: figs 25-33, pl. 4: figs 9-11 (1982). 

Navicula alineae Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of continental Israel, p. 268, pl. 

22: figs 175-178 (2000). [Navicula alineae is the type of the name of the section ALINEA Lange-Bert.] 

Navicula altiplanensis U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 152, pl. 55: figs 21-27, cf. pl. 53: figs 8-11, 13- 

15 (2000). 

Navicula amphiceropsis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 153, pl. 42: figs 1-12 (2000). 

Navicula antonii Lange-Bert., Iconogr. Diatomol. 9, p. 155 (2000). 

Replaced synonym: Navicula menisculus var. grunowii Lange-Bert., Biblioth. Diatomol. 27, p. 120, pl. 64: figs 1-11 (1993). 

Navicula aquaedurae Lange-Bert., Biblioth. Diatomol. 27, p. 95, pl. 46: figs 14-19, pl. 47: figs 1-2 (1993). [Validation of 

Navicula aquaedurae Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), p. 386, pl. 69: figs 1-10 (1991).] 

Navicula arctotenelloides Lange-Bert. et Metzeltin, Iconogr. Diatomol. 3, p. 97-98, pl. 9: figs 20-23 (1996). [Comment: 

Navicula arctotenelloides on p. 220 (= caption of pl. 9: figs 20-23) cited as "Navicula tenelloides Hustedt sensu Krasske".] 



60 

Navicula arenaria var. rostellata Lange-Bert., Biblioth. Diatomol. 9, p. 56, pl. 22: fig. 1 (1985). [As a part of the 

protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 39: fig. 2 is 

cited.] 

Navicula arkona Lange-Bert. et Witkowski, Diatoms of Europe 2, p. 17-18, 223, pl. 8: figs 8-11, pl. 73: fig. 5 (2001). 

Navicula arkona Lange-Bert. et Witkowski, Diatoms of Europe 2, p. 17-18, 223-224, pl. 8: figs 8-11, pl. 73: fig. 5 (2001). 

Navicula arvensis var. major Lange-Bert., Biblioth. Diatomol. 9, p. 56, pl. 25: figs 1-3 (1985). [As a part of the protologue, 

Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 80: figs 21, 22, 32 is cited. 

Comment: "fig. 32" is erroneously cited for fig. 22'.] 

[Nomenclatural synonym of Naviculadicta cosmopolitana Lange-Bert.] 

Navicula associata Lange-Bert., Diatoms of Europe 2, p. 18, (2001). 

Replaced synonym: Navicula reichardtiana var. crassa Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 133, pl. 47: 

figs 4-10 (1993). 

Navicula athenae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 267, 431-432, pl. 129: figs 15-16 (2000). 

Navicula atomus var. alcimonica E.Reichardt, Biblioth. Diatomol. 6, p. 39-40, pl. 12: figs 10-14, pl. 30: figs 1-2 (1984). 

[Basionym of Mayamaea atomus var. alcimonica (E.Reichardt) E.Reichardt. 

Navicula atomus var. excelsa (Krasske) Lange-Bert., Biblioth. Diatomol. 9, p. 57 (1985). 

Basionym: Navicula excelsa Krasske, Abh. Ber. Vereins Naturk. Kassel 84-89, p. 51, pl. 2: fig. 33 (1925). 

[Nomenclatural synonym of Mayamaea excelsa (Krasske) Lange-Bert.] 

Navicula atomus var. permitis (Hust.) Lange-Bert., Biblioth. Diatomol. 9, p. 57 (1985). 

Basionym: Navicula permitis Hust., Arch. Hydrobiol. 40, p. 919, pl. 41: figs 8-9 (1945). 

[Nomenclatural synonym of Mayamaea atomus var. permitis (Hust.) Lange-Bert.] 

Navicula atomus var. recondita (Hust.) Lange-Bert., Biblioth. Diatomol. 9, p. 58 (1985). 

Basionym: Navicula recondita Hust., Ber. Deutsch Bot. Ges. 60, p. 68 (1942) 

[– Navicula recondita Hust., nom. inval., in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 400: figs 6-9 (1934).] 

[Comment: The term "Differentialdiagnose von Hustedt (1962, p. 164 [Rabenhorsts Kryptogamenflora 7 (3), 2. Lieferung, p. 

168])" is treated as citation error for the first valid publication of Navicula recondita.] 

[Basionym of Mayamaea recondita (Hust.) Lange-Bert.] 

Navicula australomediocris Lange-Bert. et Rol.Schmidt, J. Paleolimnol. 3, p. 65, figs 7 i-k (1990). 

Navicula austrocollegarum Lange-Bert. et R.Voigt, Diatoms of Europe 2, p. 224, pl. 9: figs 14-19, pl. 67: fig. 5 (2001). 

Navicula brekkaensis var. brekkaensoides (W.Bock) Lange-Bert., Biblioth. Diatomol. 9, p. 59 (1985). 

Basionym: Navicula brekkaensoides W.Bock, Nova Hedwigia 5, p. 224, pl. 2: figs 128 u. 1 A, B (1963). 

[Nomenclatural synonym of Diadesmis brekkaensoides (W.Bock) Gerd Moser, Lange-Bert. et Metzeltin.] 

Navicula bicuneolus Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 139, pl. 76: figs 12-13, pl. 77: figs 5-6 (1998). 

Navicula bjoernoeyaensis Metzeltin, Witkowski et Lange-Bert., Iconogr. Diatomol. 4, p. 19, pl. 2: figs 14-18 (1996). 

Navicula borowkae Witkowski, Lange-Bert. et Metzeltin Iconogr. Diatomol. 7, p. 269, 432, pl. 145: figs 1-5 (2000). 

Navicula bourrellyivera Lange-Bert., Witkowski et Stachura, Cryptog. Algol. 19, p. 87, fig. 1-6 (1998). 

Navicula breitenbuchii Lange-Bert., Diatoms of Europe 2, p. 224-225, pl. 37: figs 8-15 (2001). 

Navicula broetzii Lange-Bert. et E.Reichardt, Iconogr. Diatomol. 2, p. 77, pl. 81: figs 1-7 (1996). 

Navicula canariana Lange-Bert., Biblioth. Diatomol. 27, p. 96-97, pl. 54: figs 10-11, pl. 55: figs 1-13 (1993). [As a part of 

the protologue is cited: Krammer & Lange-Bertalot (1991b), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 

67: figs 12-18.] [Comment: in Krammer & Lange-Bertalot (1991b) Navicula canariana was documented as "Navicula 

species von Tenerife aff. radiosafallax".] 

Navicula cantonatii Lange-Bert., Diatoms of Europe 2, p. 225, pl. 19: figs 1-8 (2001). 

Navicula cari var. cincta (Ehrenb.) Lange-Bert., Cryptog. Algol. 1, p. 37 (1980). 

Basionym: Pinnularia cincta Ehrenb., Mikrogeologie, pl. 10 (2): fig. 6 a-e (1854). 

Navicula cari var. recens Lange-Bert., Cryptog. Algol. 1, p. 37, pl. 6: figs 8-14 (1980). 



61 

Navicula cariocincta Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of continental Israel, p. 

271, pl. 23: figs 203-207 (2000). [Comment: Validation of Navicula cariocincta Lange-Bert., nom. inval., in Biblioth. 

Diatomol. 27, p. 97, pl. 44: figs 1-5. (1993).] 

Navicula catalanogermanica Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 98, pl. 64: figs 16-20 (1993). 

[– Navicula catalanogermanica Lange-Bert. et G.Hofmann, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von 


Navicula cataracta-rheni Lange-Bert., Biblioth. Diatomol. 27, p. 99-100, pl. 59: figs 13-15 (1993). 

[– Navicula cataractarheni Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 


Navicula celinei Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 20, pl. 59: figs 17-22, pl. 83: fig. 1, pl. 84: 

figs 6, 8 (1996). 

Navicula chiarae Lange-Bert. et Genkal, Iconogr. Diatomol. 6, p. 63-64, pl. 11: figs 12-15, pl. 18: figs 1-2 (1999). 

Navicula chilena Lange-Bert., nom. inval., Diatom Res. 11, p. 120, figs 69-70, 160 (1996). [Comment: lacking a type and a 

Latin description, this name was published invalidly. Erroneously fig. 159 was cited on p. 120 and p. 139 (= caption of figs 

159 and 160).] 

Navicula clagesii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 200, pl. 19: figs 17-20 (1998). 

Navicula cloacina Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p. 313, 315, figs 1d, 14-15 (1976). 

[Taxonomic synonym of Adlafia minuscula var. muralis (Grunow) Lange-Bert., according to Lange-Bertalot, Diatoms of 

Europe 2, p. 144 (2001).] 

Navicula cohnii (Hilse) Lange-Bert., Biblioth. Diatomol. 9, p. 61 (1985). 

Basionym: Stauroneis cohnii Hilse, Abh. Schles. Ges. Vaterl. Cult., Abth. Naturwiss. 40, p. 83 (1860). 

Navicula coraliana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 140, pl. 76: figs 8-11 (1998). 

Navicula crassulexigua E.Reichardt, Ber. Bayer. Bot. Ges. 64, p. 130, pl. 3: figs 4-11, figs 28-29 (1994). 

Navicula cryptocefalsa Lange-Bert., Biblioth. Diatomol. 27, p. 101, pl. 60: figs 1-2, pl. 61: figs 13-14 (1993). [As a part of 

the protologue is cited: Krammer & Lange-Bertalot (1991b), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 

64: figs 1-8.] [Comment: in Krammer & Lange-Bertalot (1991b) Navicula cryptocefalsa is documented as "Navicula 

cryptocephala Kützing".] 

Navicula cryptofallax Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 103, pl. 47: fig. 11, pl. 48: figs 1-4 (1993). 

[– Navicula cryptofallax Lange-Bert. et G.Hofmann, nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von 

Mitteleuropa, Bacillariophyceae 2 (4), p. 384 pl. 68: figs 5-9 (1991).] 

Navicula cryptotenella Lange-Bert., Biblioth. Diatomol. 9, p. 62-63, pl. 18: figs 22-23, pl. 19: figs 1-10, pl. 27: fig. 1 (1985). 

[As a part of the protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), 

pl. 33: figs 9-11, 13-17 is cited.] 

Navicula cryptotenelloides Lange-Bert., Biblioth. Diatomol. 27, p. 105, pl. 50: figs 9-12, pl. 51: figs 1-2 (1993). 

Navicula curtisterna Lange-Bert., Diatoms of Europe 2, p. 225, pl. 18: figs 1-8 (2001). 

Navicula dealpina Lange-Bert., Biblioth. Diatomol. 27, p. 106, pl. 44: figs 16-19 (1993). [Comment: validation of Navicula 

dealpina Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), 

p. 366, pl. 59: figs 1-3 (1991). As a part of the protologue is cited: Krammer & Lange-Bertalot (1986), Süßwasserflora von 

Mitteleuropa, Bacillariophyceae 2 (1), pl. 27: fig. 15.] [Comment: in Krammer & Lange-Bertalot (1986) Navicula dealpina is 

documented as "Navicula cari Ehrenberg als Navicula graciloides in Coll. Hustedt".] 

Navicula densilineolata (Lange-Bert.) Lange-Bert., Biblioth. Diatomol. 27, p. 107 (1993). 

Basionym: Navicula pseudolanceolata var. densilineolata Lange-Bert., Biblioth. Diatomol. 9, p. 89, figs 18-20 (1985). 

Navicula digitoconvergens Lange-Bert., Iconogr. Diatomol. 6, p 64-65, pl. 15: figs 1-9, pl. 16: figs 4-5 (1999). [Comment: 

name and diagnosis on p. 65 only in the 2 nd edition.] 

Navicula dissociata E.Reichardt, Biblioth. Diatomol. 6, p. 44-45, pl. 11: figs 20-21, pl. 12: figs 17-18 (1984). 

[Basionym of Craticula dissociata E.Reichardt, Diatom Res. 12, p. 305 (1997).] 

Navicula doehleri Lange-Bert., Diatoms of Europe 2, p. 226, pl. 16: figs 35-39 (2001). 



62 

Navicula ehrlichiae Lange-Bert. et E.Reichardt in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 275, pl. 23: figs 191-197 (2000). 

Navicula electrolytifuga Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 155-156, pl. 57: figs 1-5 (2000). 

Navicula elginensis var. cuneata (M.Møller ex Foged) Lange-Bert., Biblioth. Diatomol. 9, p. 68 (1985). 

Basionym: Navicula dicephala f. cuneata M.Møller ex Foged, Biblioth. Phycol. 34, p. 78, pl. 29: fig. 6 (1977). 

Navicula exiliformis E.Reichardt, Diatom Res. 3, p. 238, figs 11-21 (1988). 

Navicula germanopolonica Witkowski et Lange-Bert., Biblioth. Diatomol. 27, p. 110, pl. 45: figs 1-8 (1993). 

Navicula globuliferiformis Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Replaced synonym: Navicula globulifera var. robusta Hust., Arch. Hydrobiol. Suppl. 15, p. 266, pl. 19: figs 3-4 (1937). 

[– Navicula globulifera var. robusta Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 405: figs 20-22 

(1936).] 

[– Navicula globuliferiformis Lange-Bert., nom. inval., Biblioth. Diatomol. 27, p. 93 (1993).] 

Navicula gondwana Lange-Bert., Biblioth. Diatomol. 27, p. 111, pl. 57: figs 1-6 (1993). 

Navicula grasmueckii Lange-Bert., Biblioth. Diatomol. 27, p. 112, pl. 54: figs 1-9 (1993). 

Navicula halinae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 281, 432-433, pl. 130: figs 1-4 (2000). 

Navicula hamiltonii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 282, 433, pl. 130: figs 5-11 (2000). 

[Comment: Navicula hamiltonii is not documented on pl. 130: fig. 12 as cited on p. 282.] 

Navicula hanseatica Lange-Bert. et Stachura, Cryptog. Algol. 19, p. 87-88, fig. 23-27 (1998). 

Navicula hanseatica ssp. circumarctica Lange-Bert., Iconogr. Diatomol. 7, p. 282, 433, pl. 120: figs 7-10 (2000). 

[Comment: Navicula hanseatica ssp. circumarctica is not documented on pl. 120: figs 1-6 as cited on p. 433.] 

Navicula hastatula Lange-Bert. et Miho, Diatoms of Europe 2, p. 226, pl. 56: figs 1-8 (2001). 

Navicula heimansioides Lange-Bert., Biblioth. Diatomol. 27, p. 113, pl. 62: figs 7-10 (1993). [Comment: validation of 

Navicula heimansioides Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), p. 388, pl. 70: figs 1-8 (1991). As a part of the protologue is cited: Krammer & Lange-Bertalot 

(1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 29: figs 9-11.] [Comment: in Krammer & Lange- 

Bertalot (1986) Navicula heimansioides is documented as "Navicula cari Ehrenberg als Navicula leptostriata Jørgensen = 

Navicula heimansii Van Dam et Kooyman".] 

Navicula herbstiae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 141, pl. 76: figs 4-7, pl. 77: fig. 8 (1998). 

Navicula hintzii Lange-Bert., Biblioth. Diatomol. 27, p. 114-115, pl. 61: figs 15-18 (1993). 

[– Navicula hintzii Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), p. 374, pl. 63: figs 9-12 (1991). 

Navicula hofmanniae Lange-Bert., Biblioth. Diatomol. 27, p. 116, pl. 48: figs 12-18, pl. 49: figs 1-2 (1993). 

Navicula humboldtiana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 156, pl. 54: figs 1-6 (2000). 

Navicula incarum U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 158, pl. 49: figs 7-16 (2000). 

[Comment: in Metzeltin & Kusber (2001) the authors are erroneously cited as "Lange-Bert. et U.Rumrich".] 

Navicula ingapirca Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 159, pl. 52: figs 3-8 (2000). 

Navicula insulsa Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 142, pl. 75: figs 4-6 (1998). 

Navicula irmengardis Lange-Bert., Iconogr. Diatomol. 2, p. 78, pl. 80: figs 18-23 (1996). 

Navicula iserentantii Lange-Bert. et Witkowski, Iconogr. Diatomol. 7, p. 284, 433-434, pl. 115: figs 1-5 (2000). 

Navicula kohlenbachii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 160, pl. 53: figs 1-7 (2000). 

Navicula kohlmaieri Lange-Bert., Iconogr. Diatomol. 6, p. 66-67, pl. 17: figs 1-6 (1999). 

Navicula korzeniewskii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 287, 434, pl. 141: figs 30-33 (2000). 

Navicula krammerae Lange-Bert., Iconogr. Diatomol. 2, p. 79, pl. 80: figs 3-8 (1996). [Comment: Navicula krammerae on 

p. 79, and 278 is misspelled as "Navicula krammeriae".] 



63 

Navicula kuelbsii Lange-Bert., Biblioth. Diatomol. 9, p. 77, pl. 25: figs 4-7 (1985). [As a part of the protologue, Krammer & 

Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 78: fig. 29 and pl. 80: figs 33-35 are 

cited. Comment: "pl. 80: figs 33-35" is erroneously cited for pl. 80: figs 32-34.] 

[Basionym of Microcostatus kuelbsii (Lange-Bert.) Lange-Bert.] 

Navicula kuseliana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 161, pl. 43: figs 1-8 (2000). 

Navicula lacunolaciniata Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p. 313, figs 1e, 12-13 (1976). 

[Basionym of Mayamaea lacunolaciniata (Lange-Bert. et Bonik) Lange-Bert.] 

Navicula lange-bertalotii E.Reichardt, Ber. Bayer. Bot. Ges. 56, p. 176, pl. 1: figs 18-28, pl. 5: figs 1-3 (1985). 

[Basionym of Fallacia lange-bertalotii (E.Reichardt) E.Reichardt.] 

Navicula lauca U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 162, pl. 51: figs 1-17, 20-23, pl. 52: figs 1-2 (2000). 

Navicula leistikowii Lange-Bert., Biblioth. Diatomol. 27, p. 118, pl. 50: figs 1-8 (1993). [Comment: validation of Navicula 

leistikowii Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 

(4), p. 374, pl. 63: figs 14-20 (1991).] 

Navicula lohmannii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 163, pl. 36: figs 1-8, pl. 37: figs 1-4 (2000). 

Navicula loibl-sittlerii E.Reichardt, Nova Hedwigia 47, p. 97, pl. 4: figs 21-26 (1988). 

Navicula longicephala var. vilaplanii Lange-Bert. et Sabater, Nova Hedwigia 51, p. 180, pl. 3: figs 24-28; pl. 6: figs 7-10 

(1990). 

[Basionym of Navicula vilaplanii (Lange-Bert. et Sabater) Lange-Bert. et Sabater.] 

Navicula luciae Witkowski et Lange-Bert., Iconogr. Diatomol. 6, p. 68, pl. 16: fig. 20, pl. 17: figs 7-10 (1999). [Comment: 

Navicula luciae is not documented on pl. 16: fig. 19 as cited in the 1 st edition, reference corrected in the 2 nd edition.] 

Navicula lundii E.Reichardt, Ber. Bayer. Bot. Ges. 56, p. 180 (1985). 

Replaced synonym: Navicula cryptocephala f. terrestris J.W.G.Lund, New Phytologist 45, p. 86, 87, figs 9 H-W (1946). 

Navicula madeirensis Lange-Bert., Biblioth. Diatomol. 27, p. 119, pl. 52: figs 15-19 (1993). [As a part of the protologue is 

cited: Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), pl. 67: figs 5-10 (1991). 

[Comment: in Krammer & Lange-Bertalot (1991b) Navicula madeirensis is cited as "Navicula species von Madeira aff. 

radiosafallax".] 

Navicula mahoodii Witkowski et E.Witon, Iconogr. Diatomol. 7, p. 289, 434-435, pl. 145: figs 23-25 (2000). [Comment: in 

the protologue (p. 289 and p. 434) pl. 145: figs 24-26 is erroneously referred to.] 

Navicula maidanae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 142-143, pl. 76: figs 14-15 (1998). 

Navicula margalithii Lange-Bert., Biblioth. Diatomol. 9, p. 79-80, pl. 17: figs 1-3, pl. 28: figs 1-4 (1985). [As a part of the 

protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 26: figs 3-4, 

pl. 27: 4-6 is cited.] 

Navicula margaritiana Witkowski, Biblioth. Diatomol. 28, p. 151, pl. 37: figs 1-9 (1994). 

[Basionym of Chamaepinnularia margaritiana (Witkowski) Witkowski.] 

Navicula mediocostata E.Reichardt, Diatom Res. 3, p. 241, figs 22-29 (1988). 

Navicula melanesica Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 119-120, pl. 61: figs 1-7, 11-13 (1995). 

Navicula menisculus var. grunowii Lange-Bert., nom. illeg., Biblioth. Diatomol. 27, p. 120, pl. 64: figs 1-11 (1993). 

[Validation of Navicula menisculus var. grunowii Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora 


[Nomenclatural synonym of Navicula antonii Lange-Bert.] 

Navicula microcari Lange-Bert., Biblioth. Diatomol. 27, p. 121-122, pl. 58: figs 1-5 (1993). [Validation of Navicula 

microcari Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 

(4), p. 366, pl. 59: figs 4-7 (1991).] 

Navicula microdigitoradiata Lange-Bert., Biblioth. Diatomol. 27, p. 122 (1993). 

Replaced synonym: Navicula cincta f. minuta Van Heurck, Type de Synopsis, Texte, p. 82 (1885). 

Navicula minuscula var. muralis (Grunow) Lange-Bert. in Ross, Proceedings of the Sixth Symposium of Recent and Fossil 

Diatoms, p. 143 (1981) 



64 

Basionym: Navicula muralis Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 14: fig. 27 (1880). 

[Nomenclatural synonym of Adlafia minuscula var. muralis (Grunow) Lange-Bert.] 

Navicula moenofranconica Lange-Bert., Biblioth. Diatomol. 27, p. 123-124, pl. 59: figs 9-12 (1993). 

[– Navicula moenofranconica Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 


Navicula moskalii Metzeltin, Witkowski et Lange-Bert., Iconogr. Diatomol. 4, p. 20-21, pl. 5: figs 7-11 (1996). 

Navicula namibica Lange-Bert. et U.Rumrich, Biblioth. Diatomol. 27, p. 124-125, pl. 58: figs 16-24 (1993). [Comment: the 

authors on p. 280 (= caption of pl. 58) are cited as "Rumrich et Lange-Bert."]. 

Navicula nipponica (Skvortzov) Lange-Bert., Biblioth. Diatomol. 27, p. 126 (1993). 

Basionym: Navicula radiosa f. nipponica Skvortzov, Philipp. J. Sci. 61, p. 273, pl. 2: fig. 2; pl. 3: fig. 20 (1936). 

Navicula novaesiberica Lange-Bert., Biblioth. Diatomol. 27, p. 126-127, pl. 61: figs 1-8 (1993). 

[– Navicula novasiberica Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 


Navicula oligotraphenta Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 128, pl. 48: figs 6-11, pl. 49: figs 3-4 (1993). 

[As a part of the protologue is cited: Krammer & Lange-Bertalot (1991b), Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), pl. 60: figs 16-21.] [Comment: in Krammer & Lange-Bertalot (1991b) Navicula oligotraphenta was 

documented as "Navicula trivialis var. oligotraphenta Lange-Bert. et G.Hofmann".] 

Navicula paracari Lange-Bert., nom. inval., Biblioth. Diatomol. 27, p. 129, pl. 43: figs 11-24 (1993). [Comment: lacking a 

type and a Latin diagnosis this name is invalidly published.] 

Navicula parahasta Lange-Bert. et Miho, Diatoms of Europe 2, p. 226-227, pl. 55: figs 1-9 (2001). 

Navicula parinacota U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 164, pl. 50: figs 1-11 (2000). 

Navicula paul-schulzii Witkowski et Lange-Bert., Biblioth. Diatomol. 28, p. 154, pl. 33: figs 14-17, pl 36: figs 1, 3-4 

(1994). 

Navicula paupercula Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 165, pl. 55: figs 1-9 (2000). 

Navicula peregrinopsis Lange-Bert. et Witkowski, Iconogr. Diatomol. 7, p. 297, 435, pl. 112: figs 5-7 (2000). 

Navicula perfidissima Lange-Bert., Biblioth. Diatomol. 18, p. 162-163, pl. 70: figs 22-25', 26-27 (1989). 

Navicula phylleptosoma Lange-Bert., Iconogr. Diatomol. 6, p. 69-70, pl. 13: figs 1-5 (1999). 

Navicula podzorskii Lange-Bert., Biblioth. Diatomol. 27, p. 129-130, pl. 59: figs 1-7, pl. 60: figs 3-6 (1993). 

Navicula pseudanglica Lange-Bert. var. pseudanglica, Biblioth. Diatomol. 9, p. 86-87, pl. 23: figs 8-12 (1985). [As a part of 

the protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 46: figs 

13-15 is cited. Comment: "fig. 15" is cited erroneously on p. 86.] 

Navicula pseudokotschyi Lange-Bert., Biblioth. Diatomol. 9, p. 88, pl. 23: figs 13-14 (1985). [As a part of the protologue, 

Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), figs 16-19 is cited. Comment: 

This is an incomplete citation for pl. 60: figs 16-19.] 

Navicula pseudolanceolata Lange-Bert., Cryptog. Algol. 1, p. 32, pl. 2: figs 1-8 (1980). 

Navicula pseudolanceolata var. denselineolata Lange-Bert., Biblioth. Diatomol. 9, p. 89, pl. 18: figs 18-20 (1985). [As a 

part of the protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 36: 

fig. 8' is cited.] 

[Basionym of Navicula densilineolata (Lange-Bert.) Lange-Bert.] 

Navicula pseudoppugnata Lange-Bert. et Miho, Diatoms of Europe 2, p. 227, pl. 30: figs 1-7 (2001). 

Navicula quasidisjuncta Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 166, pl. 45: figs 9-13, pl. 49: figs 1-6 (2000). 

Navicula quechua Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 168, pl. 46: figs 30-33 (2000). 

Navicula radiosafallax Lange-Bert., Biblioth. Diatomol. 27, p. 131 (1993). 

Replaced synonym: Navicula radiosa var. parva J.H.Wallace, Notul. Nat. Acad. Nat. Sci. Philadelphia 331, p. 3, pl. 1: fig. 5 

(1960). 

Navicula radiosiola Lange-Bert., Biblioth. Diatomol. 27, p. 132, pl. 53: figs 4-8 (1993). 



65 

Navicula rajmundii Witkowski, Lange-Bert. et Metzeltin, p. 300, 435-436, pl. 138: figs 16-20, pl. 139: figs 1-4 (2000). 

Navicula rakowskae Lange-Bert., Diatoms of Europe 2, p. 227-228, pl. 57: figs 1-6 (2001). 

Navicula recurva (F.Meister) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 302 (2000). 

Basionym: Navicula gracilis var. recurva F.Meister, Ber. Schweiz. Bot. Ges. 47, p. 262, pl. 5: fig. 6 (1937). 

Navicula reichardtiana Lange-Bert. var reichardtiana., Biblioth. Diatomol. 18, p. 163-164, pl. 98: figs 19-27 (1989). 

Navicula reichardtiana var. crassa Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 133, pl. 47: figs 4-10 (1993). 

[– Navicula reichardtiana var. crassa Lange-Bert. et G.Hofmann, nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 384 pl. 68: figs 16-19 (1991).] 

[Basionym of Navicula associata Lange-Bert.] 

Navicula rhynchotella Lange-Bert., Biblioth. Diatomol. 27, p. 135, pl. 62: figs 4-6, pl. 63: figs 1-4 (1993). [Comment: 

validation of Navicula rhynchotella Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von 

Mitteleuropa, Bacillariophyceae 2 (4), p. 378, 380 pl. 65: figs 3-4, pl. 66: figs 3-4 (1991).] 

Navicula ricardae Lange-Bert., Diatoms of Europe 2, p. 228, pl. 37: figs 1-7 (2001). 

Navicula riediana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 169, pl. 43: figs 9-14 (2000). 

Navicula riotecensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 143, pl. 76: figs 1-3, pl. 77: fig. 7 (1998). 

Navicula rivulorum Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 170, pl. 55: figs 15-20 (2000). 

Navicula rolandii Wunsam, Witkowski et Lange-Bert., Cryptog. Algol. 19, p. 88, fig. 88-93. 

Navicula saprophila Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p. 312, figs 1c, 8-9, 29 (1976). 

[Basionym of Fistulifera saprophila (Lange-Bert. et Bonik) Lange-Bert.] 

Navicula seibigiana Lange-Bert., Biblioth. Diatomol. 27, p. 137, pl. 44: figs 6-15 (1993). [As a part of the protologue is 

cited: Krammer & Lange-Bertalot (1991b), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 374, pl. 63: figs 1- 

6.] [Comment: in Krammer & Lange-Bertalot (1991b) Navicula seibigiana was documented as "Navicula seibigii Lange- 

Bert.".] 

Navicula seibigii Lange-Bert. nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, 

Bacillariophyceae 2 (4), p. 374, pl. 63: figs 1-6. [Comment: lacking a type and a Latin diagnosis this name was invalidly 

published. Later published as Navicula seibigiana Lange-Bert.] 

Navicula siamexilis Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 138, pl. 53: figs 39-42 (1989). [Comment: lacking a 

type and a Latin diagnosis, this name is not validly published.] 

Navicula siamlinearis Lange-Bert., nom. inval., Biblioth. Diatomol. 18, p. 138, pl. 68: figs 39-46 (1989). [Comment: 

lacking a type and a Latin diagnosis, this name is not validly published.] 

Navicula sieminskiae Lange-Bert. et Witkowski, Diatoms of Europe 2, p. 228, pl. 18: figs 8-13 (2001). 

Navicula silvahercynia Lange-Bert., Biblioth. Diatomol. 18, p. 164-165, pl. 56: figs 11-12', pl. 98: figs 45-50 (1989). 

[Comment: erroneously pl. 56: fig. 13 is cited on p. 164.] 

Navicula specululum Witkowski, Biblioth. Diatomol. 28, p. 159, pl. 36: figs 5-12 (1994). 

[Basionym of Amicula specululum (Witkowski) Witkowski. Type of the name of the genus Amicula Witkowski, Lange-Bert. 

et Metzeltin.] 

Navicula stachurae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 306, 436, pl. 142: figs 6-9 (2000). 

Navicula starmachii Witkowski et Lange-Bert., Biblioth. Diatomol. 28, p. 160, pl. 33: figs 18-24 (1994). 

Navicula streckerae Lange-Bert. et Witkowski, Iconogr. Diatomol. 7, p. 307, 436-437, pl. 118: figs 8-15 (2000). 

Navicula structa Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 126-127, pl. 60: figs 1-5, 7-10, pl. 62: figs 11, 7 

(1995). [Comment: Navicula structa on p. 276 (= caption of pl. 62: fig. 7) is cited as "Navicula spec. 10".] 

Navicula subalpina E.Reichardt, Diatom Res. 3, p. 241, figs 30-41 (1988). 

Navicula subconcentrica Lange-Bert., Diatoms of Europe 2, p. 229, pl. 30: figs 8-13 (2001). 

Navicula supergregaria Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 171, pl. 45: figs 4-8 (2000). 

Navicula suprinii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 201, pl. 19: figs 12-16 (1998). 



66 

Navicula syvertsenii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 21, pl. 61: figs 33-36, pl. 82: figs 5, 7 

(1996). [Comment (not included in Metzeltin & Kusber (2001): pl. 61: fig. 37 is erroneously cited in the protologue. 

Navicula syvertsenii is not documented on pl. 82: fig. 6 as cited on p. 21 (see Navicula wasmundii).] 

Navicula tectissima Lange-Bert., Iconogr. Diatomol. 3, p. 150 (1996). 

Replaced synonym: Navicula tecta Krasske, Arch. Hydrobiol. 31, p. 41, pl. 8: figs 12-13 (1937). 

Navicula tridentula var. tenuis (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 151 (1996). 

Basionym: Navicula tenuis Krasske, Arch. Hydrobiol. 35, p. 385, pl. 11: fig. 51 (1939). 

Navicula trivialis Lange-Bert., var. trivialis, Cryptog. Algol. 1, p. 31, pl. 1: figs 5-9, pl. 9: figs 1-2 (1980). 

Navicula trivialis var. oligotraphenta Lange-Bert. et G.Hofmann, nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 368, pl. 60: figs 16-21 (1991). [Comment: lacking a type and a 

Latin diagnosis this name was invalidly published. New name and rank as Navicula oligotraphenta Lange-Bert. et 

G.Hofmann.] 

Navicula trophicatrix Lange-Bert., Iconogr. Diatomol. 2, p. 80-81, pl. 103: figs 28-31 (1996). 

Navicula tropicoidea Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 22, pl. 59: figs 23-27, pl. 60: figs 6-16, 

pl. 83: figs 2, 4, pl. 84: figs 1-3, 5, 7 (1996). [Comment: Navicula tropicoidea is not documented on pl. 83: figs 1, 3, 5, 7 as 

cited on p. 22.] 

Navicula ultratenelloides Lange-Bert., Iconogr. Diatomol. 2, p. 81-82, pl. 109: figs 12-13´ (1996). [Comment: Navicula 

ultratenelloides on p. 336 (caption of pl. 109) is misspelled as "N. subtenelloides".] 

Navicula vandamii var. mertensiae Lange-Bert., Iconogr. Diatomol. 7, p. 313, 437, pl. 124: figs 1-7 (2000). 

Navicula vaneei Lange-Bert., Cryptog. Algol. 19, p. 89, fig. 28-32 (1998). 

Navicula vekhovii Lange-Bert. et Genkal, p. 71, pl. 14: figs 1-7 (1999). 

Navicula ventraloconfusa Lange-Bert., Biblioth. Diatomol. 18, p. 165, pl. 79: figs 36-39 (1989). 

[Basionym of Naviculadicta ventraloconfusa (Lange-Bert.) Lange-Bert.] 

[Basionym of Sellaphora ventraloconfusa (Lange-Bert.) Metzeltin et Lange-Bert.] 

Navicula vilaplanii (Lange-Bert. et Sabater) Lange-Bert. et Sabater, Iconogr. Diatomol. 9, p. 173 (2000). 

Basionym: Navicula longicephala var. vilaplanii Lange-Bert. et Sabater, Nova Hedwigia 51, p. 180, pl. 3: figs 24-28, pl. 6: 

figs 7-10 (1990). 

Navicula viridula var. denselineolata E.Reichardt, Nova Hedwigia 47, p. 100, pl. 4: figs 16-18 (1988). 

Navicula viridula var. germainii (J.H.Wallace) Lange-Bert., Biblioth. Diatomol. 27, p. 139 (1993). 

Basionym: Navicula germainii J.H.Wallace, Notul. Nat. Acad. Nat. Sci. Philadelphia 331, p. 3, pl. 2: fig. 1 A-C (1960). 

Navicula viridula var. vulpina (Kütz.) Lange-Bert., Cryptog. Algol. 1, p. 35 (1980). 

Basionym: Navicula vulpina Kütz., Kieselschalige Bacillarien, p. 92, pl. 3: fig. 43 (1844). 

Navicula viridulacalcis Lange-Bert. ssp. viridulacalcis, Iconogr. Diatomol. 9, p. 174-175, pl. 38: fig. 5 (2000). [Comment: 

pl. 37: figs 5-8, pl. 38: figs 1-5 are cited for Navicula viridulacalcis Lange-Bert. on p. 174.] 

Navicula viridulacalcis ssp. neomundana Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 175, pl. 37: figs 5-8, pl. 38: 

figs 1-4 (2000). [Comment: Navicula viridulacalcis ssp. neomundana is erroneously cited on p. 314 (= caption of pl. 37: figs 

5-8) as Navicula viridulacalcis Lange-Bert. ssp. viridulacalcis but the type locality for Navicula viridulacalcis ssp. 

neomundana Lange-Bert. et U.Rumrich is given. The author on p. 316 (= caption of pl. 38: figs 1-4) is cited as "Lange-Bert." 

only.] 

Navicula wasmundii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 22-23, pl. 61: figs 27-28, pl. 82: figs 4, 6 

(1996). [Comment (not included in Metzeltin & Kusber (2001): Navicula wasmundii is not documented on pl. 82: fig. 5 as 

cited on p. 22 whereas pl. 82: fig. 6 is missing on p.22 (see Navicula syvertsenii).] 

Navicula wellneri Lange-Bert., Biblioth. Diatomol. 15, p. 123-124, pl. 40: figs 28-31 (1987). 

[Basionym of Nupela wellneri (Lange-Bert.) Lange-Bert.] 

Navicula wiesneri Lange-Bert., Biblioth. Diatomol. 27, p. 140, pl. 41: figs 23-37, pl. 42: figs 3-6 (1993). 

Navicula wiktoriae Witkowski et Lange-Bert., Biblioth. Diatomol. 28, p. 162-163, pl. 34: figs 1-13 (1994). 

[Basionym of Chamaepinnularia wiktoriae (Witkowski et Lange-Bert.) Witkowski, Lange-Bert. et Metzeltin.] 



67 

Navicula wildii Lange-Bert., Biblioth. Diatomol. 27, p. 141-142, pl. 46: figs 9-13 (1993). [Validation of Navicula wildii 

Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (4), p. 372 

pl. 62: figs 1-10 (1991). 

Navicula witkowskii Lange-Bert., Iserentant et Metzeltin, Cryptog. Algol. 19, p. 89-90, fig. 7-11 (1998). 

Navicula wunsamiae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 317, 437, pl. 138: figs 9-12 (2000). 

Navicula wygaschii Lange-Bert., Diatoms of Europe 2, p. 229, pl. 27: figs 1-9, pl. 69: fig. 6 (2001). 

NAVICULADICTA Lange-Bert., Biblioth. Diatomol. 29, p. 83-84 (1994). [Typus generis: Naviculadicta vaucheriae 

(J.B.Petersen) Lange-Bert., Biblioth. Diatomol. 29, p. 83 (1994).] 

Naviculadicta ambiguissima Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 202. pl. 20: figs 1-7 (1998). 

Naviculadicta angustata (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 

52 (1998). 

Basionym: Navicula perlucida f. angustata Manguin ex Kociolek et Reviers, Cryptog. Algol. 17, p. 203 (1996). 

Naviculadicta bremensiformis Lange-Bert., Iconogr. Diatomol. 2, p. 82, pl. 28: figs 21-23, 24-26 (1996). [Comment: 

Naviculadicta bremensiformis on p. 174 (caption of pl. 28: figs 21-23) is cited as "Naviculadicta (nov.) spec. Nr. 3 Julma 

Ölkky".] 

Naviculadicta chilensis (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 140 (1996). 

Basionym: Navicula ruttneri var. chilensis Krasske, Arch. Hydrobiol. 35, p. 382, pl. 11: fig. 25 (1939). 

Naviculadicta circumborealis Lange-Bert., Biblioth. Diatomol. 29, p. 84-85, pl. 52: figs 29-36 (1994). 

Naviculadicta cosmopolitana Lange-Bert., Iconogr. Diatomol. 9, p. 177 (2000). 

Replaced synonym: Navicula arvensis var. major Lange-Bert., Biblioth. Diatomol. 9, p. 56, pl. 25: figs 1-3. 

Naviculadicta digituloides Lange-Bert., Iconogr. Diatomol. 2, p. 83, pl. 109: figs 8-11, pl. 115: fig. 4 (1996). 

Naviculadicta digitulus (Hust.) Lange-Bert. et Metzeltin, nom. inval., Iconogr. Diatomol. 2, p. 84 (1996). [Comment: 

lacking a full and direct reference this name is invalidly published.] 

Naviculadicta elorantana Lange-Bert., Iconogr. Diatomol. 2, p. 84, pl. 28: figs 15-20, pl. 116: figs 3-4 (1996). 

Naviculadicta kaiseri Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 85, pl. 26: figs 14-18 (1996). 

Naviculadicta lepida Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 203, pl. 21: figs 8-11 (1998). 

Naviculadicta moratoria Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 204, pl. 22: figs 1-3 (1998). 

Naviculadicta multiconfusa Lange-Bert., Biblioth. Diatomol. 29, p. 86-87, pl. 52: figs 39-43 (1994). [Naviculadicta 

multiconfusa was documented already in Lange-Bertalot & Krammer (1989), Bibl. Diatomol. 18, pl. 79: figs 36, 39 as 

"Navicula ventraloconfusa Lange-Bert.".] 

Naviculadicta nagumoi Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 205, pl. 21: figs 12-15 (1998). 

Naviculadicta nana (Hust.) Lange-Bert., Iconogr. Diatomol. 2, p. 85 (1996). 

Basionym: Stauroneis nana Hust., Abh. Naturwiss. Vereine Bremen 34, p. 259, fig. 3 (1957). 

Naviculadicta nanogomphonema Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 178, pl. 67: figs 1-13 (2000). 

Naviculadicta obdurata (M.H.Hohn et Hellerman) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 86 (1996). 

Basionym: Navicula obdurata M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82, p. 307, pl. 4: figs 27-28 (1963). 

Naviculadicta parabryophila Lange-Bert., Iconogr. Diatomol. 2, p. 86, pl. 27: figs 11-15, pl. 117: figs 5-7 (1996). 

[Basionym of Adlafia parabryophila (Lange-Bert.) Lange-Bert.] 

Naviculadicta parasemen Lange-Bert., Iconogr. Diatomol. 6, p. 72, pl. 25: figs 1-5, pl. 26: figs 1-4 (1999). 

Naviculadicta pseudofallacia Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 23, pl. 50: figs 20-21, pl. 76: 

figs 9-12 (1996). [Comment: Naviculadicta pseudofallacia is not documented on pl. 76: fig. 8 as cited on p. 23. 

Naviculadicta pseudofallacia on p. 132 (caption of pl. 50: 20-21) is misprinted as "Achnanthes cf. pygmaea (Schum.) 

Cleve"]. 

Naviculadicta pseudofusticulus Lange-Bert. et Steindorf, Biblioth. Diatomol. 38, p. 205-206, pl. 21: figs 5-7 (1998). 

[– Naviculadicta pseudofusticulus Lange-Bert. et Steindorf, nom. inval., Biblioth. Diatomol. 32, p. 132, pl. 54: figs 14-15 

(1995).] 



68 

Naviculadicta pseudostauron Lange-Bert., Iconogr. Diatomol. 2, p. 87, pl. 35: figs 11-18, pl. 117: figs 1-2 (1996). 

[Basionym of Kobayasia pseudostauron (Lange-Bert.) Lange-Bert.] 

[Basionym of Kobayasiella pseudostauron (Lange-Bert.) Lange-Bert.] 

Naviculadicta raederae Lange-Bert., Iconogr. Diatomol. 2, p. 88, pl. 83: figs 37-40, pl. 116: figs 5-6 (1996). [Comment: 

Naviculadicta raederae on p. 88, 284, and 350 is misspelled as "Naviculadicta raederiae".] 

Naviculadicta rionautensis U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 179-180, pl. 70: 14-18; pl. 79: fig. 8 (2000). 

Naviculadicta sassiana Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 144-145, pl. 75: figs 9-12 (1998). 

Naviculadicta schneckenburgeri Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 207, pl. 22: figs 7-11 

(1998). 

Naviculadicta stauroneioides Lange-Bert., Iconogr. Diatomol. 2, p. 89, pl. 109: figs 22-27 (1996). 

Naviculadicta tomesii Gerd Moser, Lange-Bert. et Metzeltin var. tomesii, Biblioth. Diatomol. 38, p. 208, pl. 21: figs 1-2 

(1998). 

Naviculadicta tomesii var. clagesii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 209, pl. 21: figs 3-4 

(1998). 

Naviculadicta vaucheriae (J.B.Petersen) Lange-Bert., Biblioth. Diatomol. 29, p. 83 (1994). 

Basionym: Navicula vaucheriae J.B.Petersen, Kongel. Danske Vidensk.-Selsk. Skr. (Afd. 12) 7, p. 291, fig. 13 (1915). 

[Naviculadicta vaucheriae is the type of the name of the genus NAVICULADICTA.] 

Naviculadicta vekhovii Lange-Bert. et Genkal, Iconogr. Diatomol. 6, p. 74, pl. 20: figs 20-21 (1999). 

Naviculadicta ventraloconfusa (Lange-Bert.) Lange-Bert. var. ventraloconfusa, Biblioth. Diatomol. 29, p. 87 (1994). 

Basionym: Navicula ventraloconfusa Lange-Bert., partim, Biblioth. Diatomol. 18, p. 165, pl. 79: figs 37, 38, nec 36 (1989). 

[Nomenclatural synonym of Sellaphora ventraloconfusa (Lange-Bert.) Metzeltin et Lange-Bert.] 

Naviculadicta ventraloconfusa var. chilensis (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 155 (1996). 

Basionym: Navicula ventralis var. chilensis Krasske, Arch. Hydrobiol. 35, p. 383, pl. 11: fig. 45 (1939). 

Naviculadicta witkowskii Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 89-90, pl. 28: figs 31-35, pl. 115: fig. 2 (1996). 

NAVISANTIQUA Lange-Bert. (subsection within the genus NAVICULA s. str.), Diatoms of Europe 2, p. 223, (2001). 

[Typus subsectionis: Navicula perturbata Jurilj, Prirodosl. Istraz. 26, p. 141, fig. 43e (1954).] 

NEIDIOPSIS Lange-Bert. et Metzeltin, Diatoms of Europe 2, p. 130 (2001). [Typus generis: Neidiopsis vekhovii Lange- 

Bert. et Genkal, Diatoms of Europe 2, p. 230 (2001). Comment: the genus NEIDIOPSIS Lange-Bert. et Metzeltin was 

invalidly pulished in Iconogr. Diatomol. 6, p. 76. (1999), NEIDIOPSIS was based on Neidium vekhovii and Neidiopsis 

vekhovii, both names invalid at the time of publication. The genus NEIDIOPSIS in Iconogr. Diatomol. 6 was originally 

described with a Latin diagnosis. Since the genus was not intended to be monotypic, this diagnosis was not available for 

validating Neidiopsis vekhovii in 1999.] 

Neidiopsis levanderi (Hust.) Lange-Bert. et Metzeltin in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Basionym: Navicula levanderi Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 43, p. 174 (1943). 

[≡ Neidium levanderi (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 91 (1996).] 

[– Navicula levanderi Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 397: figs 30-32 (1934).] 

[– Navicula levanderi Hust., nom. inval., Bot. Arch. 38, p. 195 (1937).] 

[– Neidiopsis levanderi (Hust.) Lange-Bert. et Metzeltin, nom. inval., Iconogr. Diatomol. 6, p. 77 (1999).] 

[– Neidiopsis levanderi (Hust.) Lange-Bert. et Metzeltin, nom. inval. (Art. 33.3, St. Louis Code), Diatoms of Europe 2, p. 

131 (2001).] 

Neidiopsis vekhovii Lange-Bert. et Genkal, Diatoms of Europe 2, p. 230, pl. 101: fig. 22, pl. 102: figs 2-8 (2001) [Comment: 

validation of Neidiopsis vekhovii Lange-Bert. et Genkal, nom. inval., Iconogr. Diatomol. 6, p. 78 (1999). Neidiopsis vekhovii 

is the type of the genus NEIDIOPSIS Lange-Bert. et Metzeltin. Since Neidiopsis vekhovii was not validly published before 

2001, the genus NEIDIOPSIS also was not validly published before 2001.] 

Neidiopsis wulffii (J.B.Petersen) Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Basionym: Navicula wulffii J.B. Petersen, Meddel. Grønland 64, p. 312, fig. 1 (1924). 

[– Neidiopsis wulffii (J.B.Petersen) Lange-Bert., nom. inval., Iconogr. Diatomol. 6, p. 78 (1999).] 

[– Neidiopsis wulffii (J.B.Petersen) Lange-Bert., nom. inval. (Art. 33.3, St. Louis Code), Diatoms of Europe 2, p. 133 

(2001).] 



69 

NEIDIUM Pfitzer 


Neidium andinum Lange-Bert., Iconogr. Diatomol. 9, p. 180-181, pl. 102: figs 5-7 (2000). 

Neidium binodeforme Krammer, Biblioth. Diatomol. 9, p. 102-103, pl. 5: figs 14-15, pl. 43: figs 1-5 (1985). [As a part of 

the protologue, Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 100: figs 

6-8 is cited. Comment: in Krammer & Lange-Bertalot (1986), p. 640 (= caption of pl. 100: figs 6-8) Neidium binodeforme is 

cited as "Neidium binodeformis".] 

Neidium carteri Krammer, Biblioth. Diatomol. 9, p. 104, pl. 4: figs 1-8 (1985). [Comment: Neidium carteri on p. 104 and p. 

138 misspelled as "Neidium carterii".] 

Neidium catarinense (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 101 (1996). 

Basionym: Navicula catarinensis Krasske, Svensk Bot. Tidskr. 42, p. 431, pl. 2: fig. 4 (1948). 

Neidium coralieae Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 145, pl. 121: figs 11-12 (1998). 

Neidium cuneatum Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 146, pl. 124: fig. 6, pl. 127: fig. 2 (1998). 

Neidium demerariense Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 147, pl. 120: figs 2-7 (1998). 

Neidium dubiforme Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 148, pl. 124: figs 10-13 (1998). 

Neidium elongatum (Hust.) Lange-Bert., Metzeltin et Krammer, Iconogr. Diatomol. 2, p. 91 (1996). 

Basionym: Neidium dubium var. elongata Hust., Arch. Hydrobiol. 43, p. 448, Text-fig. 10 (1950). 

Neidium essequiboanum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 149, pl. 120: figs 13-16 (1998). 

Neidium excisum Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 149-150, pl. 123: figs 5-6, 8, pl. 126: figs 5-6 (1998). 

Neidium gracile f. simplex Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 32, p. 137, pl. 79: fig. 2 (1995). 

Neidium hamatum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 151, pl. 121: figs 1-2 (1998). 

Neidium infirmum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 152, pl. 118: figs 1-8 (1998). 

Neidium latum Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 153, pl. 119: figs 1-3, pl. 126: figs 1-4 (1998). 

Neidium levanderi (Hust.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 91 (1996). 

Basionym: Navicula levanderi Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 43, p. 174. 

[– Navicula levanderi Hust., nom inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 397: figs 30-32 (1934).] 

[Comment: The term "descr. Hustedt 1961 p. 98, fig. 1245" is treated as citation error for the first valid publication of 

Navicula levanderi.] 

[Nomenclatural synonym of Neidiopsis levanderi (Hust.) Lange-Bert. et Metzeltin.] 

Neidium obliquum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 154-155, pl. 124: figs 4-5, pl. 127: figs 3-4 (1998). 

Neidium subamphigomphus Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 155, pl. 120: figs 10-12 (1998). 

Neidium subdubium Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 156, pl. 119: figs 6-9 (1998). 

NITZSCHIA Hassall 


NITZSCHIA subgen. DENTICULOIDEA [see DENTICULOIDEA]. 

NITZSCHIA subgen. GRUNOWINITZSCHIA [see GRUNOWINITZSCHIA]. 

Nitzschia acidoclinata Lange-Bert., Nova Hedwigia 28, p. 277-278, pl. 7: figs 19-21, pl. 10: figs 1-2 (1977). 

Nitzschia alexandrina (Cholnoky) Lange-Bert. et Simonsen, Bacillaria 1, p. 16 (1978). 

Basionym: Nitzschia elliptica var. alexandrina Cholnoky, Hydrobiologia 11, p. 258, figs 29-30 (1958). 

Nitzschia alpinobacillum Lange-Bert., Biblioth. Diatomol. 27, p. 143, pl. 114: figs 2-6, pl. 115: fig. 5 (1993). [As a part of 

the protologue is cited: Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 78: 

figs 10-12.] [Comment: in Krammer & Lange-Bertalot (1988) Nitzschia alpinobacillum was documented as "Nitzschia 

bacillum Hustedt".] 

Nitzschia andicola Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 193, pl. 158: figs 3-9, pl. 159: figs 1-6 (2000). 

[Nitzschia andicola is the type of the name of the subgenus GRUNOWINITZSCHIA Lange-Bert. et U.Rumrich.] 



70 

Nitzschia angustatula Lange-Bert., Biblioth. Diatomol. 15, p. 6-7, pl. 18: figs 1-4, 5 (1987). [As a part of the protologue, 

Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 3: fig. 6, pl. 36: figs 6-10 

is cited.] 

Nitzschia angusteforaminata Lange-Bert., Bacillaria 3, p. 43, pl. 2: figs 44-51, pl. 9: figs 127-132 (1980). 

Nitzschia aniae Lange-Bert., Biblioth. Diatomol. 27, p. 144, pl. 116: figs 1-10 (1993). 

Nitzschia archibaldii Lange-Bert., Bacillaria 3, p. 44, pl. 1: figs 14-18, pl. 7: figs 115-121 (1980). 

Nitzschia blankaartensis Denys et Lange-Bert., Nova Hedwigia 67, p. 248, 250, fig. 1-18 (1998). 

Nitzschia brevior (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 53 

(1998). 

Basionym: Nitzschia luzonensis f. brevior Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 212 (1996). 

Nitzschia brunoi Lange-Bert., Iconogr. Diatomol. 2, p. 92, pl. 101: figs 11-15, pl. 102: fig. 1 (1996). 

Nitzschia chungara U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 182, pl. 166: figs 1-10 (2000). 

Nitzschia commutatoides Lange-Bert., Biblioth. Diatomol. 15, p. 11, pl. 21: figs 5-7 (1987). [As a part of the protologue, 

Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 34: figs 1-4 is cited. 

Comment: "pl. 34: figs 1-4" is erroneously cited for pl. 43: figs 1-4.] 

Nitzschia compressa var. pararostrata Lange-Bert., Iconogr. Diatomol. 3, p. 164, pl. 48: figs 6, 6' (1996). 

[Basionym of Nitzschia pararostrata (Lange-Bert.) Lange-Bert.] 

Nitzschia dealpina Lange-Bert. et G.Hofmann, Biblioth. Diatomol. 27, p. 146, pl. 114: figs 8-13, pl. 115: figs1-4 (1993). 

Nitzschia dissipata ssp. oligotraphenta Lange-Bert., Iconogr. Diatomol. 2, p. 93, pl. 99: figs 13-16 (1996). 

Nitzschia eglei Lange-Bert., Biblioth. Diatomol. 15, p. 15, pl. 28: figs 1-3 (1987). [As a part of the protologue, Krammer & 

Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 10: figs 6-9 is cited.] 

Nitzschia epithemiformis Lange-Bert., Bacillaria 3, p. 45, pl. 5: figs 105-109 (1980). 

Nitzschia fibula-fissa Lange-Bert., Bacillaria 3, p. 46-47, pl. 3: figs 63-66, pl. 12: figs 146-148 (1980). 

Nitzschia gaertnerae Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 94, pl. 68: figs 6-11 (1996). [Comment: Nitzschia 

gaertnerae on p. 94-95 and 254 is misspelled as "Nitzschia gaertneriae".] 

Nitzschia gisela Lange-Bert., Biblioth. Diatomol. 15, p. 21, pl. 20: figs 1-6 (1987). [As a part of the protologue, Krammer & 

Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 2: figs 2-3, pl. 42: figs 7-8 is cited.] 

Nitzschia graciliformis Lange-Bert. et Simonsen, Bacillaria 1, p. 33 (1978). 

Replaced synonym: Nitzschia graciloides Hust., nom. illeg., Veröff. Inst. Meeresforsch. Bremerhaven 6, p. 95, figs 4-5 

(1959); nec Nitzschia graciloides Hust., Arch. Hydrobiol. 47, p. 633, figs 10-11 (1953). 

Nitzschia homburgiensis Lange-Bert., Nova Hedwigia 30, p. 650-651, pl. 3: figs 27-28, pl. 4: figs 40-41 (1978). 

Nitzschia intermedia f. actinastroides (Lemmerm.) Lange-Bert., Bacillaria 3, p. 48 (1980). 

Basionym: Synedra actinastroides Lemmerm., Ber. Deutsch. Bot. Ges. 18, p. 30 (1900). 

Nitzschia kahlii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 183, pl. 165: figs 3-10 (2000). 

Nitzschia kanakarum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 209 (1998). 

Replaced synonym: Nitzschia sigma var. fonticola Hust., Arch. Hydrobiol. Suppl. 16, p. 486, pl. 40: figs 16-17 (1938). 

Nitzschia labella Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 210, pl. 59: figs 6-11 (1998). 

Nitzschia lacuum Lange-Bert., Bacillaria 3, p. 49, pl. 5: figs 91-97, pl. 11: figs 138-141 (1980). 

Nitzschia leistikowii Lange-Bert., Bacillaria 3, p. 50, pl. 1: figs 19-27, pl. 8: figs 122-126 (1980). 

Nitzschia lorenziana var. muscicola Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 211, pl. 59: figs 1-2 

(1998). 

Nitzschia macaronesica Lange-Bert., Biblioth. Diatomol. 27, p. 148, pl. 112: figs 6-10, pl. 113: fig. 6 (1993). 

Nitzschia machardyae Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 184, pl. 155: figs 1-3 (2000). 

Nitzschia metzeltinii Lange-Bert., Iconogr. Diatomol. 6, p. 78-79, pl. 72: figs 1-7 (1999). 



71 

Nitzschia monachorum Lange-Bert., Biblioth. Diatomol. 15, p. 35-36, pl. 6: figs 1-6 (1987). [As a part of the protologue, 

Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 1: figs 6-7, pl. 57: figs 5-8 

is cited.] 

Nitzschia neotropica Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 185, pl. 167: figs 4-8 (2000). 

Nitzschia novae-guineaensis E.Reichardt, Nova Hedwigia 47, p. 110, pl. 7: figs 16-22 (1988). 

Nitzschia oberheimiana U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 186, pl. 168: figs 1-17 (2000). 

Nitzschia pararostrata (Lange-Bert.) Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 320 (2000). 

Basionym: Nitzschia compressa var. pararostrata Lange-Bert., Iconogr. Diatomol. 3, p. 164, pl. 48: fig. 6 (1996). 

Nitzschia plioveterana Lange-Bert., Iconogr. Diatomol. 7, p. 401 (2000). 

Replaced synonym: Nitzschia angustata var. minuta Krasske, Arch. Hydrobiol. 24, p. 445, fig. 20 (1932). 

Nitzschia quadrangula (Kütz.) Lange-Bert., Bacillaria 3, p. 53 (1980). 

Basionym: Synedra quadrangula Kütz., Kieselschalige Bacillarien, p. 63, pl. 3: fig. 23 (1844). 

Nitzschia rectirobusta Lange-Bert., Iconogr. Diatomol. 2, p. 95 (1996). 

Replaced synonym: Nitzschia recta var. robusta Hust., Arch. Hydrobiol. 43, p. 403, pl. 41: figs 2-3 (1950). 

Nitzschia retusa Lange-Bert. et Bonik, Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16), p. 322, figs 1n, 22-23, 30-31 (1976). 

Nitzschia rosenstockii Lange-Bert., Bacillaria 3, p. 52-53, pl. 2: figs 30-33, pl. 10: figs 133-136 (1980). 

Nitzschia schwabei Krasske ex Lange-Bert., K. Külbs, T. Lauser, Nörpel-Schempp et Willmann, Iconogr. Diatomol. 3, p. 

164 (1996). 

Replaced synonym: Nitzschia incognita Krasske, nom. illeg., Arch. Hydrobiol. 38, p. 276, pl. 5: figs 2-3 (1941); non Legler 

& Krasske, Beih. Bot. Zentralbl. 60, p. 343, figs 3a, b (1940). 

Nitzschia semirobusta Lange-Bert., Biblioth. Diatomol. 27, p. 149, pl. 120: figs 3-21, pl. 122: fig. 7, pl. 123: figs 1-7 (1993). 

[Nitzschia semirobusta is the type of the name of the subgenus DENTICULOIDEA.] 

Nitzschia serpentiraphe Lange-Bert., Biblioth. Diatomol. 27, p. 151, pl. 125: figs 2-6, pl. 126: figs 1-9, pl. 127: figs 1-6, pl. 

128: figs 1-2 (1993). [Comment: Nitzschia serpentiraphe is not documented on pl. 125: fig. 1 and pl. 128: figs 3-6 as 

suggested on p. 151.] 

Nitzschia spinarum Lange-Bert., Iconogr. Diatomol. 2, p. 96, pl. 70: figs 34-35 (1996). 

Nitzschia strelnikovae Lange-Bert., Genkal et Vekhov, nom. inval., Iconogr. Diatomol. 6, p. 79, pl. 70: figs 6-11 (1999). 

[Comment: lacking a type and a Latin diagnosis, this name is invalidly published.] 

Nitzschia supralitorea Lange-Bert., Arch. Hydrobiol. Suppl. 56,2 (Algol. Stud. 23), p. 215, figs 25-27, 76-78 (1979). 

Nitzschia tenerifa Lange-Bert., Biblioth. Diatomol. 27, p. 153, pl. 112: figs 1-5, pl. 113: figs 1-5 (1993). 

[– Nitzschia tenerifa Lange-Bert., nom. inval., Biblioth. Diatomol. 15, p. 55, pl. 36: figs 11-19 (1987).] 

Nitzschia tsarenkoi Lange-Bert. in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of continental Israel, p. 325, 

pl. 25: figs 215-217 (2000). 

Nitzschia umbonata (Ehrenb.) Lange-Bert., Nova Hedwigia 30, p. 648 (1978). 

Basionym: Navicula umbonata Ehrenb., Infusionsthierchen, p. 177 (1838). 

Nitzschia valdecostata Lange-Bert. et Simonsen, Bacillaria 1, p. 58, pl. 16: figs 260-263; pl. 18: figs 269-270 (1978). 

[Comment: "Simonsen" is missing in Metzeltin & Kusber (2001).] 

Nitzschia vermicularoides Lange-Bert., Iconogr. Diatomol. 9, p. 187-188, pl. 157: figs 1-4 (2000). 

Nitzschia vildaryana U.Rumrich et Lange-Bert., Iconogr. Diatomol. 9, p. 189, pl. 169: figs 1-9 (2000). 

Nitzschia vixnegligenda Lange-Bert. et G.Hofmann, p. 190, pl. 173: figs 1-13 (2000). 

Nitzschia witkowskii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 157, pl. 206: fig. 7 (1998). 

Nitzschia wuellerstorffii Lange-Bert., Biblioth. Diatomol. 15, p. 61, pl. 1: figs 1-4 (1987). [As a part of the protologue, 

Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 4: fig. 3, pl. 6: figs 1-6 is 

cited.] 



72 

NUPELA Vyverman et Compère 


[http://128.32.109.44/cgi-bin/porp_cgi.plNUPELAWVYVERMANPCOMPERE1991]. 

Nupela astartiella Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 157-158, pl. 72: figs 46-55 (1998). 

Nupela brachysirioides Lange-Bert., Biblioth. Diatomol. 27, p. 154-155, pl. 39: figs 1-5 (1993). 

Nupela butterfassiana (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes butterfassiana Lange-Bert., Biblioth. Diatomol. 18, p. 34, pl. 71: figs 15-29, pl. 72: figs 2-5 (1989). 

Nupela comperei Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 211-212, pl. 31: figs 1-4 (1998). 

Nupela cymbelloidea Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 158, pl. 72: figs 38-41, pl. 73: fig. 5 (1998). 

[Comment: Nupela cymbelloidea is not documented on pl. 72: figs 39-42 as cited on p. 158. Nupela cymbelloidea on page 

380 (= caption of pl. 73: fig. 5) is cited as "Nupela (nov.) spec. Nr. 72/1".] 

Nupela deformis Lange-Bert., Biblioth. Diatomol. 29, p. 76, pl. 49: figs 23-24, pl. 51: fig. 6 (1994). 

Nupela encyonopsis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 159, pl. 72: figs 35-37, pl. 74: figs 3-4 (1998). 

Nupela gracillima (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 158 (1993). 

Basionym: Stauroneis gracillima Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 43, p. 154, figs 21-23 (1943). 

Nupela imperfecta (Schim.) Lange-Bert., Iconogr. Diatomol. 6, p. 80 (1999). 

Basionym: Achnanthes imperfecta Schim., Nova Hedwigia 30, p. 564, pl. 1: figs 1-5 (1978). 

Nupela impexiformis (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes impexiformis Lange-Bert., Biblioth. Diatomol. 18, p. 68, pl. 74: figs 7-9, pl. 75: figs 2-4 (1989). 

Nupela jahniae-reginae Lange-Bert. et Metzeltin, Iconogr. Diatomol. 9, p. 194-195, pl. 34: figs 1-8 (2000). [Comment: the 

author on p. 308 (= caption of pl. 34: figs 1-8) is cited as "Lange-Bert." only.] 

Nupela lapidosa (Krasske) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes lapidosa Krasske, Bot. Arch. 27, p. 350, fig. 9 (1929). 

[Comment: Erroneously the basionym author was given as "Lange-Bertalot".] 

Nupela lesothensis (Schoeman) Lange-Bert., Biblioth. Diatomol. 38, p. 213 (1998). 

Basionym: Rhoicosphenia lesothensis Schoeman, A systematic and ecological study of the diatom flora of Lesotho. Nat. Inst. 

Water Res., Pretoria, p. 221, figs 85-91 (1973). 

[Comment: This combination was again marked as new in Iconogr. Diatomol. 9, p. 195 (2000).] 

Nupela neotropica Lange-Bert., Biblioth. Diatomol. 29, p. 77, pl. 49: figs 7-9, pl. 51: figs 1-3 (1994). 

Nupela obliqua Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 160, pl. 72: figs 42-45, pl. 73: figs 2-4 (1998). 

Nupela paludigena (Scherer) Lange-Bert., Biblioth. Diatomol. 27, p. 158 (1993). 

Basionym: Anomoeoneis paludigena Scherer, Diatom Res. 3, p. 149, figs 42-45, 72 (1988). 

Nupela praecipua (E.Reichardt) E.Reichardt, Iconogr. Diatomol. 9, p. 196 (2000). 

Basionym: Achnanthes praecipua E.Reichardt in Round, Proc. Ninth Int. Diatom Symp. 1986, p. 391, figs 1-24 (1988). 

Nupela rhetica (Wuthrich) Lange-Bert., Biblioth. Diatomol. 27, p. 155 (1993). 

Basionym: Anomoeoneis rhetica Wuthrich, Ergebn. Wiss. Unters. Schweiz. Nationalpark 14, p. 310, pl. 15: figs 1-5 (1975). 

Nupela rumrichorum Lange-Bert., Biblioth. Diatomol. 29, p. 79, pl. 49: figs 14-22, pl. 50: figs 1-6 (1994). 

Nupela schoemaniana Lange-Bert., Biblioth. Diatomol. 27, p. 156, pl. 36: figs 1-9, pl. 37: figs 1-6 (1993). 

Nupela subpallavicinii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 161, pl. 72: figs 23-24, pl. 73: fig. 1, pl. 74: fig. 5 

(1998). 

Nupela silvahercynia (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 2, p. 97 (1996). 

Basionym: Achnanthes silvahercynia Lange-Bert., Biblioth. Diatomol. 18, p. 139, pl. 71: figs 30-37, pl. 72: figs 2-4 (1989). 

Nupela tenuicephala (Hust.) Lange-Bert., Biblioth. Diatomol. 27, p. 157 (1993). 

Basionym: Navicula tenuicephala Hust., Arch. Hydrobiol. 39, p. 113, figs 16-20 (1942). 

Nupela tenuistriata (Hust.) Metzeltin et Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Basionym: Achnanthes tenuistriata Hust., Bot. Not. 1952, p. 384, figs 57-59 (1952). 



73 

[– Achnanthes tenuistriata Hust., nom. inval., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 410: figs 8-9 (1937).] 

[– Nupela tenuistriata (Hust.) Metzeltin et Lange-Bert., nom. inval., Iconogr. Diatomol. 5, p. 161 (1998).] 

Nupela tropica (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes tropica Hust., Arch. Hydrobiol. Suppl. 15, p. 200, pl. 13: figs 28-32 (1937). 

Nupela vyvermanii Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 213, pl. 31: figs 7-10 (1998). 

Nupela wellneri (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 9, p. 196 (2000). 

Basionym: Navicula wellneri Lange-Bert., Biblioth. Diatomol. 15, p. 123, pl. 40: figs 28-31 (1987). 

Nupela zizkae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 162, pl. 71: figs 11-19 (1998). 

OESTRUPIA Heiden ex Hust. 


Oestrupia ergadensis (W.Greg.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 318 (2000). 

Basionym: Pinnularia ergadensis W.Greg., On the post-tertiary diatomaceous sand of Glenshire I and II. Micr. Soc. London 

4, p. 48, pl. 5: fig. 22 (1856). 

OPEPHORA P.Petit 


Opephora burchardtiae Witkowski, Metzeltin et Lange-Bert., Biblioth. Diatomol. 38, p. 214, 355-356, pl. 71: fig. 3, pl. 78: 

figs 1-13 (1998). [Comment: Opephora burchardtiae is not documented on pl. 71: fig. 2 as cited on p. 214 and 355.] 

Opephora horstiana Witkowski, Biblioth. Diatomol. 28, p. 173, pl. 12: figs 1-8 (1994). 

Opephora krumbeinii Witkowski, Witak et Stachura, Iconogr. Diatomol. 6, p. 80, pl. 3: figs 1-17, pl. 4: figs 1-3 (1999). 

Opephora minuta (Cleve-Euler) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 71 (2000). 

Basionym: Opephora marina var. minuta Cleve-Euler, Kongl. Svenska Vetenskapsakad. Handl. Ser. 4, p. 16, fig. 314c 

(1951). 

ORTHOSEIRA Thwaites 


Orthoseira roeseana var. tropica (Krasske) Lange-Bert. et Willmann, Iconogr. Diatomol. 3, p. 94 (1996). 

Basionym: Melosira roeseana var. tropica Krasske, Svensk Bot. Tidskr. 42, p. 422, pl. 1: figs 1-2 (1948). 

PARADIADESMIS Lange-Bert. et Le Cohu (subgenus within DIADESMIS), Iconogr. Diatomol. 9, p. 108 (2000). [Typus 

subgeneris: Diadesmis corrugata Lange-Bert. in Biblioth. Diatomol. 38, p. 140 (1998).] 

PARLIBELLUS E.J.Cox 


Parlibellus adnatus (Heiden) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 319 (2000). 

Basionym: Libellus adnatus Heiden in Heiden & Kolbe, Die marinen Diatomeen der Deutschen Süd-polarexpedition 1901- 

1903. Deutsch. Südpolar-Exped. 8, Bot (5): 450-715, p. 630, pl. 4: fig. 84 (1928). 

Parlibellus bennikei Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 319, 437-438, pl. 104: figs 11-15 (2000). 

Parlibellus calvus Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 24, pl. 54: figs 10-14 (1996). 

Parlibellus coxiae Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 4, p. 24, pl. 54: figs 27-31, pl. 79: figs 1-2 

(1996). 

Parlibellus crucicula (W.Sm.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 321 (2000). 

Basionym: Stauroneis crucicula W.Sm., Synopsis British Diatomaceae 1, p. 60, pl. 19: fig. 192 (1853). 

Parlibellus cruciculoides (C.Brockmann) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 321 (2000). 

Basionym: Navicula cruciculoides C.Brockmann, Abh. Senckenb. Naturf. Ges. 478, p. 15, pl. 4: figs 5-7 (1950). 

Parlibellus ellipticus (Gallik) Lange-Bert. et Witkowski in Nevo & Wasser, Biodiversity of cyanoprocaryotes and algae of 

continental Israel, p. 292 (2000). 

Basionym: Navicula protracta var. elliptica Gallik, Pannonhalmi Szemle 10, p. 65, fig. 7 (1935). 

Parlibellus hendeyi Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 323, 438, pl. 104: figs 21-23 (2000). 

Parlibellus perytii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 323, 438-439, pl. 105: figs 14-16 (2000). 



74 

Parlibellus phoebeae Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 7, p. 324, 439, pl. 105: figs 1-9 (2000). 

Parlibellus protracta (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 324 (2000). 

Basionym: Navicula protracta Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17 (2), p. 140 (1880). 

Parlibellus protractoides (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 325 (2000). 

Basionym: Navicula protractoides Hust., Abh. Naturwiss. Vereine Bremen 34, p. 283, figs 32-33 (1957). 

[Comment: In Iconogr. Diatomol. 9 (2000), p. 197 this combination is again marked as new. The Authors are given as 

"(Hust.) Witkowski et Lange-Bert."] 

Parlibellus rhombicula (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 325 (2000). 

Basionym: Navicula rhombicula Hust., Rabenhorsts Kryptogamenflora 7 (3) 2. Lieferung, p. 327, fig. 1442 (1962). 

PETRONEIS A.J.Stickle et D.G.Mann in F.E. Round, R.M. Crawford & D.G. Mann 


Petroneis glacialis (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 326 (2000). 

Basionym: Cocconeis glacialis Cleve, Bih. Kongl. Svenska Vetenk.-Akad. Handl. 1 (13), p. 14, pl. 3: fig. 12 (1873). 

Petroneis xi (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 330 (2000). 

Basionym: Navicula xi Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26, p. 41 (1895). 

PINNULARIA Ehrenb. 


Pinnularia acidophila G.Hofmann et Krammer, Diatoms of Europe 1, p. 213, pl. 14: figs 1-22 (2000). 

Pinnularia acorica Krammer et Metzeltin, Diatoms of Europe 1, p. 236-237, pl. 183: figs 1-2 (2000). 

Pinnularia acoricola var. elongatirhombica Wydrzycka, Lange-Bert. et Metzeltin, Brenesia 55-56, p. 13, pl. 6: figs 1-8 

(2001). 

Pinnularia acoricola var. linearis Wydrzycka, Lange-Bert. et Metzeltin, Brenesia 55-56, p. 13, pl. 5: figs 13-19, pl. 6: fig. 3 

(2001). 

Pinnularia acoricola var. pumilio Wydrzycka, Lange-Bert. et Metzeltin, Brenesia 55-56, p. 13-14, pl. 5: figs 29-41, pl. 6: 

figs 2, 4-7 (2001). 

Pinnularia acrosphaeria var. parva Krammer, Diatoms of Europe 1, p. 213, pl. 22: figs 3-6 (2000). 

Pinnularia acrosphaeria var. tumidula Krammer, Diatoms of Europe 1, p. 214, pl. 21: figs 8-9 (2000). [Comment: on p. 

300 (= caption of pl.21: figs 8-9) Pinnularia acrosphaeria var. tumidula erroneously is cited as "Pinnularia acrosphaeria 

var. turgidula Grunow ex Cleve".] 

Pinnularia acuminata var. carteri Krammer, Diatoms of Europe 1, p. 232, pl. 145: fig. 1 (2000). 

Pinnularia acuminata var. novazealandiae Krammer, Diatoms of Europe 1, p. 232, pl. 148: figs 1-5 (2000). 

Pinnularia alabamae Krammer, Diatoms of Europe 1, p. 239, pl. 212: figs 1-3 (2000). [Comment: on p. 239 erroneously pl. 

212: figs 4-5 is cited (pl. 212: figs 4-5 shows Pinnularia pulchella var. canadensis).] 

Pinnularia alta Krammer, Diatoms of Europe 1, p. 218, pl. 57: fig. 7 (2000). 

Pinnularia altiplanensis Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 197, pl. 145: figs 20-26 (2000). 

Pinnularia amabilis Krammer, Diatoms of Europe 1, p. 224 pl. 86: figs 1-9 (2000). 

[Taxonomic comment: Pinnularia amabilis is discussed by Idei & Mayama (2001) with respect to Pinnularia acidojaponica 

M.Idei et H.Kobayasi in Jahn et al.: Lange-Bertalot-Festschrift, p. 266, figs 1-29 (2001).] 

Pinnularia amazonica Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 163, pl. 176: figs 4-7 (1998). 

Pinnularia anglica Krammer, Biblioth. Diatomol. 26, p. 171, pl. 40: figs 21-23 (1992). 

Pinnularia angusta var. rostrata Krammer, Diatoms of Europe 1, p. 222, pl. 83: figs 21-23 (2000). 

Pinnularia angustiborealis Krammer et Lange-Bert. emend. Krammer et Lange-Bert., Diatoms of Europe 3, p. 174-175 

(2002). [Comment: intended validation of Pinnularia angustiborealis Krammer et Lange-Bert., Diatoms of Europe 1, p. 24, 

209, pl. 9: figs 5-6 (2000). The Latin diagnosis is rewritten, the name is marked again as new.] 

Pinnularia angustiborealis Krammer et Lange-Bert., Diatoms of Europe 1, p. 24, 209, pl. 9: figs 5-6 (2000). [Comment: 

emended diagnosis in: Diatoms of Europe 3, p. 174-175 (2002).] 



75 

Pinnularia angustistriata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 164-165, pl. 190: figs 7-9 (1998) [Comment: on p. 

164 "pl. 190: figs 6-8" is erroneously cited.] 

Pinnularia angustivalva Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 164, pl. 190: figs 10-11 (1998). 

Pinnularia appendiculata var. amaniana Krammer, Diatoms of Europe 1, p. 225, pl. 91: figs 10-11 (2000). 

Pinnularia appendiculata var. brevistriata Krammer, Diatoms of Europe 1, p. 225, pl. 91: fig. 33 (2000). 

Pinnularia araucanensis Lange-Bert., U.Rumrich et Krammer, Iconogr. Diatomol. 9, p. 198, pl. 150: figs 1-18 (2000). 

Pinnularia bacilliformis Krammer, Biblioth. Diatomol. 26, p. 171, pl. 50: figs 7-12 (1992). 

Pinnularia bertrandii Krammer var. bertrandii, Diatoms of Europe 1, p. 226, pl. 91: figs 22-28 (2000). 

Pinnularia bertrandii var. angustefasciata Krammer, Diatoms of Europe 1, p. 226, pl. 91: figs 29-30 (2000). 

Pinnularia bertrandii var. orleansiana Krammer, Diatoms of Europe 1, p. 226, pl. 91: figs 31-32, 34-35 (2000). 

Pinnularia bleischii Krammer, Diatoms of Europe 1, p. 235, pl. 174: figs 1-5 (2000). 

Pinnularia borealis var. islandica Krammer, Diatoms of Europe 1, p. 209, pl. 6: fig. 5, pl. 7: figs 1-5 (2000). 

Pinnularia borealis var. subislandica Krammer, Diatoms of Europe 1, p. 209, pl. 8: figs 1-5, 6 (2000). 

Pinnularia borealis var. sublinearis Krammer, Diatoms of Europe 1, p. 210, pl. 7: figs 14-19 (2000). 

Pinnularia borealis var. tenuistriata Krammer, Diatoms of Europe 1, p. 210, pl. 6: figs 7-10 (2000). 

Pinnularia boyeriformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 165, pl. 184: figs 4-6 (1998). 

Pinnularia brandeliformis Krammer, Biblioth. Diatomol. 26, p. 171, pl. 21: figs 1-4 (1992). 

Pinnularia brandeliformis var. linearis Krammer, Diatoms of Europe 1, p. 227, pl. 98: fig. 4 (2000). 

Pinnularia brebissonii var. lancarea Krammer, Diatoms of Europe 1, p. 216, pl. 47: fig. 6 (2000). 

Pinnularia brebissonii var. minuta Krammer, Diatoms of Europe 1, p. 216-217, pl. 47: figs 7-9 (2000). 

Pinnularia brebissoniiformis Krammer, Diatoms of Europe 1, p. 217, pl. 46: figs 1-5 (2000). 

Pinnularia bullacostae Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 82, pl. 50: figs 1-5 (1999). [Comment: Pinnularia 

bullacostae is also documented on pl. 50: fig. 5 although not cited in the 1 st edition, reference corrected in the 2 nd edition]. 

Pinnularia canadensis Krammer, Diatoms of Europe 1, p. 214, pl. 27: figs 3-7 (2000). 

Pinnularia carteri Krammer, Diatoms of Europe 1, p. 213, pl. 15: figs 3-4 (2000). 

Pinnularia castraregina Krammer, Diatoms of Europe 1, p. 211, pl. 10: figs 9-13 (2000). 

Pinnularia certa Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 166, pl. 178: figs 12-17 (1998). 

Pinnularia cherryfieldiana Krammer, Diatoms of Europe 1, p. 237, pl. 199: figs 1-5, pl. 200: fig. 1 (2000). 

Pinnularia claviculiformis Metzeltin et Krammer, Diatoms of Europe 1, p. 227, pl. 99: figs 1-3 (2000). 

Pinnularia clevei var. angusta Krammer, Diatoms of Europe 1, p. 231, pl. 131: fig. 5 (2000). 

Pinnularia cleveiformis Krammer, Biblioth. Diatomol. 26, p. 171, pl. 72: figs 1-2 (1992). 

Pinnularia complexa Krammer var. complexa, Diatoms of Europe 1, p. 230, pl. 129: figs 1-8 (2000). 

Pinnularia complexa var. minor Krammer, Diatoms of Europe 1, p. 230-231, pl. 130: figs 1-3 (2000). 

Pinnularia confirma Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 166-167, pl. 180: figs 11-13 (1998). 

Pinnularia copulata Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 83, pl. 54: figs 1-2 (1999). 

Pinnularia cruxarea Krammer, Diatoms of Europe 1, p. 230, pl. 123: figs 1-7 (2000). 

Pinnularia cuneatiformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 167, pl. 191: figs 3-4, pl. 195: fig. 3 (1998). 

Pinnularia cuneola E.Reichardt, Nova Hedwigia 34, p. 501, pl. 2: figs 15-20, pl. 6: fig. 23 (1981). 



76 

Pinnularia curticostata Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 84 (1999). 

Replaced synonym: Navicula curtestriata Østrup, Botany of Iceland 2 (1) no. 5, p. 25, pl. 3: fig. 32 (1920); non Pinnularia 

curtistriata Hust. nec Navicula curtistriata Pant. 

Pinnularia curtispinulosa Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 199, pl. 146: figs 12-21, pl. 148: 

figs 5-7 (2000). 

Pinnularia decrescens var. rhombarea Krammer, Diatoms of Europe 1, p. 216, pl. 39: figs 1-3 (2000). 

Pinnularia denotata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 168, pl. 190: figs 1-3, 4, 5 (1998). 

Pinnularia discrepantia Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 169, pl. 191: figs 1-2 (1998). 

Pinnularia dispar Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 170, pl. 183: figs 9-10 (1998). 

Pinnularia distinguenda var. obtusa Krammer, Diatoms of Europe 1, p. 237, pl. 193: figs 1-5 (2000). 

Pinnularia divergens var. ignorata Krammer, Biblioth. Diatomol. 26, p. 172, pl. 28: figs 7-8 (1992). 

Pinnularia divergens var. media Krammer, Diatoms of Europe 1, p. 215, pl. 34: figs 1-5 (2000). 

Pinnularia divergens var. mesoleptiformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 170, pl. 173: figs 1-4 (1998). 

Pinnularia divergens var. perundulata Krammer, Diatoms of Europe 1, p. 215, pl. 32: figs 2-3 (2000). 

Pinnularia divergens var. protracta Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 171, pl. 172: figs 1, 2-3 (1998). 

Pinnularia divergens var. rhombica Krammer, nom. inval., Diatoms of Europe 1, p. 215, pl. 35: figs 5-8 (2000). [Comment: 

the var. rhomica is combined with Pinnularia divergens on p. 215 and Pinnularia ovata on p. 64, hence both combinations 

are invalidly published.] 

Pinnularia divergens var. rhombundulata Krammer, Diatoms of Europe 1, p. 215, pl. 39: figs 4-5 (2000). 

Pinnularia divergens var. rostrata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 172, pl. 171: fig. 5 (1998). 

Pinnularia divergens var. subbacillaris Krammer, Diatoms of Europe 1, p. 215, pl. 34: figs 7, 11-12 (2000). 

Pinnularia divergens var. subcuneata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 172, pl. 171: figs 2-4 (1998). 

Pinnularia divergens var. sublineariformis Krammer, Diatoms of Europe 1, p. 215, pl. 34: fig. 9 (2000). 

Pinnularia divergentissima var. ardnamurchan Krammer, Diatoms of Europe 1 (corrected issue), p. 212 pl. 11: fig. 31 

(2001, "2000"). [Comment: Pinnularia divergentissima var. ardnamurchan is not documented on pl. 11: fig. 31, as cited on 

p. 212 in the original issue of Diatoms of Europe 1. The given figure (pl. 11: fig. 31) is published in Diatoms of Europe 1, 

corrected issue (2001 "2000"), as cited on p. 212 in both issues. Pinnularia divergentissima var. ardnamurchan on p. 280 (= 

caption of pl. 11) erroneously is cited as "Pinnularia divergentissima var.".] 

Pinnularia divergentissima var. minor Krammer, Biblioth. Diatomol. 26, p. 172, pl. 12: figs 11-13 (1992). 

Pinnularia divergentissima var. triundulata Krammer, Diatoms of Europe 1, p. 212, pl. 11: figs 26-30 (2000). [Comment: 

the given figures were published in Diatoms of Europe 1, corrected issue (2001 "2000").] 

Pinnularia diversa var. subcapitata Krammer et Lange-Bert., Diatoms of Europe 1, p. 46, 212, pl. 14: fig. 27 (2000). 

Pinnularia diversarea Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 173, pl. 174: figs 2-5 (1998). 

Pinnularia doloma var. fasciata Krammer, Diatoms of Europe 1, p. 211, pl. 58: figs 13-15 (2000). [Comment: Pinnularia 

doloma var. fasciata is not documented on pl. 58: figs 1-3 as cited on p. 211.] 

Pinnularia dornii Metzeltin, Iconogr. Diatomol. 2, p. 97, pl. 45: figs 19-21 (1996). 

Pinnularia dubitabilis var. minor Krammer, Diatoms of Europe 1, p. 210, pl. 7: fig. 20 (2000). 

Pinnularia elegans (W.Sm.) Krammer, Biblioth. Diatomol. 26, p. 73 (1992). 

Basionym: Navicula elegans W.Sm., Synopsis Brit. Diatomaceae 1, p. 49, pl. 16: fig. 137 (1853). 

[Nomenclatural synonym of Pinnunavis elegans (W.Sm.) Okuno.] 

[For Pinnunavis elegans (W.Sm.) Okuno see INA: http://128.32.109.44/images/JOB0877/004/0007/00070004.TIF] 

Pinnularia egregia Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 173, pl. 179: figs 1-8 (1998). 

Pinnularia episcopalis var. perelliptica Krammer, Diatoms of Europe 1, p. 216, pl. 41: fig. 6 (2000). 



77 

Pinnularia erratica var. fossilis Krammer, Diatoms of Europe 1, p. 221, pl. 73: fig. 8 (2000). 

Pinnularia esoxiformis var. angusta Krammer, Diatoms of Europe 1, p. 232, pl. 141: fig. 5 (2000). 

Pinnularia esoxiformis var. eifeliana Krammer, Biblioth. Diatomol. 26, p. 172, pl. 58: figs 5-9 (1992). 

Pinnularia falaiseana Krammer, Biblioth. Diatomol. 26, p. 172, pl. 41: fig. 6 (1992). 

Pinnularia ferrophila Krammer, Diatoms of Europe 1, p. 222-223, pl. 81: figs 1-9 (2000). 

Pinnularia fossilis Krammer, Diatoms of Europe 1, p. 231, pl. 133: fig. 9 (2000). 

Pinnularia frauenbergiana E.Reichardt, Ber. Bayer. Bot. Ges. 56, p. 182, pl. 2: figs 5-10 (1985). 

Pinnularia frequentis Krammer, Diatoms of Europe 1, p. 227, pl. 104: figs 1-8 (2000). 

Pinnularia genkalii Krammer et Lange-Bert., Diatoms of Europe 1, p. 182, 238, pl. 203: figs 1-4 (2000). 

Pinnularia gibbiformis Krammer, Biblioth. Diatomol. 26, p. 172, pl. 48: figs 1-7 (1992). 

Pinnularia gigaformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 175, pl. 201: figs 1-3 (1998). 

Pinnularia globiceps var. angustefasciata Krammer, Diatoms of Europe 1, p. 221, pl. 74: figs 14-15 (2000). 

Pinnularia globiceps var. linearis Krammer, Diatoms of Europe 1, p. 221, pl. 72: fig. 1, pl. 74: figs 10-13 (2000). 

[Comment: on p. 402 (= caption of p. 72: fig. 1) Pinnularia globiceps var. linearis is erroneously cited as "Pinnularia lundii 

var. linearis".] 

Pinnularia graciloides var. latecapitata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 176, pl. 180: figs 5-8 (1998). 

Pinnularia graciloides var. rumrichae Krammer, Diatoms of Europe 1, p. 226-227, pl. 99: fig. 4 (2000). 

Pinnularia grunowii Krammer, Diatoms of Europe 1, p. 222, pl. 77: figs 7-14, pl. 81: figs 10-17, pl. 82: figs 7-8 (2000). 

Pinnularia halophila Krammer, Biblioth. Diatomol. 26, p. 172-173, pl. 57: figs 1-2 (1992). 

Pinnularia hemiptera var. brevistriata Krammer, Iconogr. Diatomol. 5, p. 176, pl. 190: figs 12-14 (1998). 

Pinnularia hemipteriformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 177, pl. 174: figs 6-7 (1998). 

Pinnularia humilis Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 84-85, pl. 52b: figs 8-14 (1999). 

Pinnularia ilkaschoenfelderae Krammer, Diatoms of Europe 1, p. 238, pl. 204: figs 2-6 (2000). [Comment: on p. 238 pl. 

204: fig. 1 is cited (pl. 204: fig. 1 shows P. genkalii).] 

Pinnularia infirma Krammer, Biblioth. Diatomol. 9, p. 109-110, pl. 10: figs 16-17 (1985). [As a part of the protologue, 

Krammer & Lange-Bertalot (1986), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (1), pl. 183: figs 14-17 is cited.] 

Pinnularia instabiliformis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 177, pl. 174: figs 8-12 (1998). 

Pinnularia instabilis (A.W.F.Schmidt) Metzeltin, Iconogr. Diatomol. 5, p. 178 (1998). 

Basionym: Navicula instabilis A.W.F.Schmidt in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 43: figs 38-40, non figs 35- 

37 (1875). 

Pinnularia ivaloensis Krammer, Diatoms of Europe 1, p. 229-230, pl. 121: figs 2-6 (2000). 

Pinnularia julma Krammer et Metzeltin, Iconogr. Diatomol. 2, p. 98, pl. 45: figs 10-13 (1996). 

Pinnularia jungii Krammer, Diatoms of Europe 1, p. 212, pl. 58: figs 18-21 (2000). 

Pinnularia kanakarum Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 214-215, pl. 55: figs 1-5 (1998). 

Pinnularia kittelii Krammer, Diatoms of Europe 1, p. 221, pl. 71: figs 1-6 (2000). 

Pinnularia krammeri Metzeltin, Iconogr. Diatomol. 2, p. 98-99, pl. 57: figs 3-8 (1996). 

Pinnularia krookiformis Krammer, Biblioth. Diatomol. 26, p. 173, pl. 18: figs 18-21 (1992). 

[Basionym of Chamaepinnularia krookiformis (Krammer) Lange-Bert. et Krammer.] 

Pinnularia kuetzingii Krammer, Biblioth. Diatomol. 26, p. 173, pl. 36: figs 26-35 (1992). 

Pinnularia latarea Krammer var. latarea, Diatoms of Europe 1, p. 224, pl. 84: fig. 13-15 (2000). [Comment: erroneously pl. 

80: figs 1-6 is cited on p. 224. These figures show Pinnularia latarea var. thermophila.] 



78 

Pinnularia latarea var. thermophila Krammer, Diatoms of Europe 1, p. 224, pl. 80: figs 1-6 (2000). 

Pinnularia laucensis Lange-Bert., U.Rumrich et Krammer, Iconogr. Diatomol. 9, p. 200-201, pl. 145: figs 14-19 (2000). 

Pinnularia legumiformis Krammer, Biblioth. Diatomol. 26, p. 173, pl. 29: figs 1-4 (1992). 

Pinnularia linearis Lange-Bert., Wydrzycka et Metzeltin, Brenesia 55-56, p. 14, pl. 7: figs 12-14 (2001). 

Pinnularia lokana Krammer, Diatoms of Europe 1, p. 221, pl. 70: figs 1-6a (2000). 

Pinnularia lunata Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 85, pl. 52a: figs 9-13 (1999). 

Pinnularia lundii var. baltica Krammer, Biblioth. Diatomol. 26, p. 173, pl. 16: figs 5-8 (1992). 

Pinnularia lundii var. linearis Krammer, Diatoms of Europe 1, p. 222, pl. 75: figs 1-6 (2000). 

Pinnularia maculata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 179, pl. 177: figs 2, 3-5 (1998). 

Pinnularia magnacuneata Krammer, Diatoms of Europe 1, p. 236, pl. 181: figs 3-5 (2000). 

Pinnularia marchica Ilka Schönfelder var. marchica, Diatoms of Europe 1, p. 218-219, pl. 12: figs 11-17 (2000). 

[Comment: Pinnularia marchica var. marchica is not documented on pl. 90: figs 4-5 as cited on p. 218.] 

Pinnularia marchica var. subrostrata Krammer, Diatoms of Europe 1, p. 219, pl. 90: figs 4-5 (2000). [Comment: 

authorship according to p. 700.] 

Pinnularia mayeri var. similis Krammer, Biblioth. Diatomol. 26, p. 173, pl. 42: fig. 1 (1992). 

Pinnularia meridiana Metzeltin et Krammer var. meridiana, Iconogr. Diatomol. 5, p. 180, pl. 181: figs 1-2, 4-5 (1998). 

Pinnularia meridiana var. concava Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 180, pl. 181: figs 6-7 (1998). 

[Comment: Pinnularia meridiana var. concava is not documented on pl. 18 as referred to on p. 180.] 

Pinnularia meridiana var. parallela Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 181, pl. 181: fig. 3 (1998). 

Pinnularia metzeltinii Krammer, Diatoms of Europe 1, p. 219, pl. 59: figs 8-16 (2000). 

Pinnularia microstauron var. angusta Krammer, Diatoms of Europe 1, p. 217, pl. 51: figs 4-7 (2000). 

Pinnularia microstauron var. brasiliensis Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 181, pl. 179: figs 9-15 (1998). 

Pinnularia microstauron var. nonfasciata Krammer, Diatoms of Europe 1, p. 217, pl. 52: figs 1-13, pl. 55: figs 1-2 (2000). 

Pinnularia microstauron var. rostrata Krammer, Diatoms of Europe 1, p. 217, pl. 51: figs 8-18 (2000). 

Pinnularia minutiformis Krammer, Diatoms of Europe 1, p. 210, pl. 9: figs 18-19 (2000) [non Pinnularia minutiformis 

Krammer, nom. illeg., Diatoms of Europe 1, p. 29 which was renamed as Pinnularia minoricapitata Krammer, Diatoms of 

Europe 3, p. 175 (2002).] 

Pinnularia montgomeryana Krammer, Diatoms of Europe 1, p. 228, pl. 110: figs 6-8 (2000). 

Pinnularia neglectiformis Krammer, Diatoms of Europe 1, p. 233, pl. 158: figs 1-8 (2000). 

Pinnularia neomajor Krammer, Biblioth. Diatomol. 26, p. 174, pl. 62: fig. 2 (1992). 

Pinnularia neomajor var. cuneata Krammer, Diatoms of Europe 1, p. 234, pl. 152: figs 1-2, 6 (2000). 

Pinnularia neomajor var. frequentis Krammer, Diatoms of Europe 1, p. 234, pl. 137: figs 2-5 (2000). 

Pinnularia neomajor var. inflata Krammer, Diatoms of Europe 1, p. 233-234, pl. 171: figs 1-5 (2000). 

Pinnularia nitzschiophila U.Rumrich, Metzeltin et Lange-Bert., Iconogr. Diatomol. 9, p. 201, pl. 144: figs 1-6 (2000). 

Pinnularia nobilefasciata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 182, pl. 176: figs 8-9, 11-16 (1998). 

Pinnularia nobilis var. inarensis Krammer, Diatoms of Europe 1, p. 239, pl. 170: fig. 4 (2000). 

Pinnularia nobilis var. regularis Krammer, Diatoms of Europe 1, p. 239, pl. 207: figs 1-8, pl. 208: figs 1-5, pl. 209: figs 1-6 

(2000). 

Pinnularia nodosa var. percapitata Krammer, Diatoms of Europe 1, p. 214, pl. 26: figs 9-12, pl. 27: figs 9-10 (2000). 

Pinnularia nodosiformis Krammer, Diatoms of Europe 1, p. 214, pl. 27: figs 1-2 (2000). 

Pinnularia normandiana Krammer, Diatoms of Europe 1, p. 235, pl. 176: figs 1-5 (2000). 



79 

Pinnularia notabilis Krammer, Biblioth. Diatomol. 9, p. 110-111, pl. 11: figs 1-5 (1985). 

Pinnularia novazealandiae Krammer, Diatoms of Europe 1, p. 216, pl. 43: figs 3-7 (2000). 

Pinnularia obscuriformis var. subrostrata Krammer, Diatoms of Europe 1, p. 213, pl. 13: fig. 37 (2000). 

Pinnularia obtusicuneata Lange-Bert., Krammer et U.Rumrich, Iconogr. Diatomol. 9, p. 202, pl. 145: figs 9-12 (2000). 

Pinnularia oriunda Krammer, Biblioth. Diatomol. 26, p. 174, pl. 60: figs 5-6, pl. 61: figs 1-6 (1992). 

Pinnularia oriundiformis Krammer, Diatoms of Europe 1, p. 228, pl. 113: figs 1-4 (2000). 

Pinnularia ovata var. rhombica Krammer, nom. inval., Diatoms of Europe 1, p. 64, pl. 35: figs 5-8 (2000). [Comment: the 

var. rhomica is combined with Pinnularia ovata on p. 64 and Pinnularia divergens on p. 215, hence both combinations are 

invalidly published.] 

Pinnularia parvulissima Krammer, Diatoms of Europe 1, p. 220, pl. 65: figs 9-10, pl. 69: figs 7-11 (2000). 

Pinnularia patagonensis Lange-Bert. et Metzeltin, Iconogr. Diatomol. 9, p. 203, pl. 138: figs 4-5 (2000). 

Pinnularia percuneata Krammer var. percuneata, Diatoms of Europe 1, p. 232, pl. 139: figs 1-7 (2000). 

Pinnularia percuneata var. constricta Krammer, Diatoms of Europe 1, p. 232, pl. 137: fig. 1 (2000). 

Pinnularia percuneata var. minor Krammer, Diatoms of Europe 1, p. 232, pl. 140: figs 1-8 (2000). 

Pinnularia perinarensis Krammer, Diatoms of Europe 1, p. 230, pl. 126: figs 4-5 (2000). 

Pinnularia perinstabilis Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 183, pl. 193: figs 1-3 (1998). 

Pinnularia perinterrupta Krammer, Diatoms of Europe 1, p. 219, pl. 60: figs 7-14, pl. 61: fig. 7 (2000). 

Pinnularia perirrorata Krammer, Diatoms of Europe 1, p. 211, pl. 17: figs 11-22 (2000). 

Pinnularia permontana Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 183, pl. 198: fig. 1 (1998). 

Pinnularia perspicua Krammer, Diatoms of Europe 1, p. 229, pl. 120: figs 1-5 (2000). 

Pinnularia persudetica Krammer var. persudetica, Diatoms of Europe 1, p. 227-228, pl. 108: figs 1-6, pl. 109: fig. 3 (2000). 

Pinnularia persudetica var. silvatica Krammer, Diatoms of Europe 1, p. 228, pl. 109: figs 1-2, (2000). [Comment: on p. 228 

pl. 109: fig. 3 is erroneously cited (pl. 109: fig. 3 shows Pinnularia persudetica var. persudetica).] 

Pinnularia petersenii Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 86, pl. 52b: figs 4-7 (1999). 

Pinnularia pisciculus var. angusta Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 184, pl. 175: figs 6-12, pl. 194: figs 5-6 

(1998). 

Pinnularia pluvianiformis Krammer, Diatoms of Europe 1, p. 222, pl. 83: figs 10-12 (2000). 

Pinnularia poasana Lange-Bert., Wydrzycka et Metzeltin, Brenesia 55-56, p. 14, pl. 8: figs 16-18 (2001). 

Pinnularia podzorskii Krammer, Diatoms of Europe 1, p. 219-220, pl. 61: figs 1-6 (2000). 

Pinnularia polyonca var. stidolphii Krammer, Diatoms of Europe 1, p. 220, pl. 95: figs 8-11 (2000). 

Pinnularia polyonca var. sumatrana Krammer, Diatoms of Europe 1, p. 220, pl. 96: figs 4-7 (2000). 

Pinnularia problematica Ant.Mayer var. problematica, Denkschriften Bayerischen Bot. Ges. Regensburg 22, p. 152, pl. 10: 

figs 4-6 (1946). [Comment: Krammer (1992) Biblioth. Diatomol. 26, p. 170, 174 intended to validate Pinnularia 

problematica Ant.Mayer. Since Pinnularia problematica was already validly published, Krammer typified Pinnularia 

problematica by designating a lectotype, as "holotype", documented on pl. 75: figs 1-3. A Latin diagnosis by Krammer 

(1992), p. 73 is treated as additional information which does not emend Mayer's German diagnosis.] 

Pinnularia problematica var. subundulata Ant.Mayer, Denkschriften Bayerischen Bot. Ges. Regensburg 22, p. 152, pl. 11: 

fig. 1 (1946). 

[Taxonomic synonym of Pinnularia problematica var. problematica, according to Krammer, Biblioth. Diatomol. 26, p. 170 

(1992).] 

Pinnularia procera Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 184-185, pl. 178: figs 7-11 (1998). 

Pinnularia pseudacuminata Metzeltin et Krammer, Iconogr. Diatomol. 2, p. 99-100, pl. 53: figs 1-4 (1996). 



80 

Pinnularia pseudocardinalis E.Reichardt, Iconogr. Diatomol. 1, p. 16-17, pl. 15: figs 1-2 (1995). [Pinnularia 

pseudocardinalis E.Reichardt, Iconogr. Diatomol. 1, (second edition), p. 33, pl. 15: figs 1-2 (1995). In the second edition 

Pinnularia pseudocardinalis E.Reichardt was again marked as new.] 

Pinnularia pseudogibba Krammer var. pseudogibba, Biblioth. Diatomol. 26, p. 174, pl. 48: figs 8-14 (1992). 

Pinnularia pseudogibba var. rostrata Krammer, Biblioth. Diatomol. 26, p. 174, pl. 52: fig. 1 (1992). 

Pinnularia pseudoparva Krammer et Lange-Bert., Iconogr. Diatomol. 6, p. 87, pl. 54: fig. 7 (1999). 

Pinnularia pseudosimilis Krammer, Biblioth. Diatomol. 26, p. 174-175, pl. 13: figs 3-5 (1992). 

Pinnularia pulchella var. canadensis Krammer, Diatoms of Europe 1, p. 239, pl. 212: figs 4-5 (2000). 

Pinnularia pulchra var. subtilis Schim., Biblioth. Diatomol. 9, p. 112, pl. 10: fig. 14 (1985). [As a part of the protologue, 


Pinnularia rabenhorstii var. cuneata Krammer et Lange-Bert., Diatoms of Europe 1, p. 22, 209, pl. 5: figs 10-11 (2000). 

Pinnularia rabenhorstii var. franconia Krammer, Diatoms of Europe 1, p. 209, pl. 5: figs 5-9, pl. 6: figs 1-4 (2000). 

Pinnularia rathsbergiana Krammer, Diatoms of Europe 1, p. 235-236, pl. 1: figs 5-8, pl. 178: figs 1-6 (2000). 

Pinnularia reichardtii Krammer, Diatoms of Europe 1, p. 231, pl. 130: figs 4-7 (2000). 

Pinnularia renata Krammer, Biblioth. Diatomol. 26, p. 175, pl. 52: figs 2-8 (1992). 

Pinnularia renatiformis Krammer, Diatoms of Europe 1, p. 218, pl. 59: fig. 25 (2000). 

Pinnularia rhenohassiaca Krammer, Diatoms of Europe 1, p. 172, 236, pl. 180: figs 1-3. (2000). [Comment: Pinnularia 

rhenohassiaca is cited in the corrected issue as "Pinnularia rhenohassiaca Krammer et Lange-Bert.".] 

Pinnularia rhenohassiaca Krammer, Diatoms of Europe 1, p. 236, pl. 180: figs 1-3 (2000). 

Pinnularia rhombarea Krammer var. rhombarea, Iconogr. Diatomol. 5, p. 185, pl. 175: figs 1-5 (1998). 

Pinnularia rhombarea var. brevicapitata Krammer, Diatoms of Europe 1, p. 218, pl. 56: figs 7-11 (2000). 

Pinnularia rhombarea var. halophila Krammer, Diatoms of Europe 1, p. 217, pl. 54: figs 6-11 (2000). 

Pinnularia rhombarea var. undulata Krammer, Diatoms of Europe 1, p. 217, pl. 56: figs 1-3 (2000). 

Pinnularia rhombarea var. variarea Krammer, Diatoms of Europe 1, p. 218, pl. 56: figs 4-6 (2000). 

Pinnularia rhomboelliptica Krammer var. rhomboelliptica, Diatoms of Europe 1, p. 233, pl. 149: figs 4-9, pl. 159: figs 1-5, 

pl. 194: fig. 2 (2000). [Comment: on p. 233 erroneously pl. 151: figs 1-3 is cited (pl. 151: figs 1-3 shows Pinnularia 

rhomboelliptica var. rhombica).] 

Pinnularia rhomboelliptica var. inflata Krammer, Diatoms of Europe 1, p. 233, pl. 152: figs 3-5, pl. 153: figs 1-4 (2000). 

[Comment: on p. 233 erroneously pl. 159: fig. 1 is cited (pl. 159: fig. 1 shows Pinnularia rhomboelliptica var. 

rhomboelliptica).] 

Pinnularia rhomboelliptica var. novazealandiae Krammer, Diatoms of Europe 1, p. 233, pl. 149: figs 1-3, pl. 150: figs 1-3 

(2000). [Comment: on p. 233 the name is misprinted as "var. novazealandia".] 

Pinnularia rhomboelliptica var. rhombica Krammer, nom. inval., Diatoms of Europe 1, p. 161, pl. 151: 1-3 (2000). 

[Comment: lacking a Latin diagnosis, the name is invalidly published.] 

Pinnularia rhomboelliptica var. zuricana Krammer, Diatoms of Europe 1, p. 233, pl. 150: fig. 4 (2000). 

Pinnularia rhombofasciata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 174, pl. 168: figs 8-11 (1998). 

Pinnularia rostratissima var. parva Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 186, pl. 169: figs 6-9, pl. 203: figs 3, 6 

(1998). 

Pinnularia rostratissima var. ventricosa Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 186-187, pl. 169: figs 1-5, pl. 203: 

figs 1, 4 (1998). 

Pinnularia rumrichae Krammer, Diatoms of Europe 1, p. 223, pl. 84: figs 9-12 (2000). 

Pinnularia rupestris var. cuneata Krammer, Biblioth. Diatomol. 26, p. 175, pl. 56: figs 1-3 (1992). 

Pinnularia santafiorana Krammer, Diatoms of Europe 1, p. 234, pl. 169: figs 5-6 (2000). 



81 

Pinnularia saprophila Lange-Bert. et Krammer, Diatoms of Europe 1, p. 109, 223, pl. 85: figs 10-18 (2000). [Comment: on 

p. 428 (caption of pl. 85 in the original issue) the authors are cited as "Kobayashi, Lange-Bert. et Krammer", on p. 109 

(corrected issue) and p. 428 (caption of pl. 85 in the corrected issue) the authors are cited as "Lange-Bert., Kobayasi et 

Krammer".] 

[Taxonomic comment: Pinnularia saprophila is discussed by Idei & Mayama (2001) with respect to Pinnularia 

valdetolerans Mayama et H.Kobayasi in Jahn et al.: Lange-Bertalot-Festschrift, p. 270, figs 30-53 (2001).] 

Pinnularia savona Krammer, Diatoms of Europe 1, p. 236, pl. 182: figs 1-4 (2000). 

Pinnularia schimanskii Krammer, Diatoms of Europe 1, p. 210, pl. 9: figs 10-13 (2000). 

Pinnularia schoenfelderi Krammer, Biblioth. Diatomol. 26, p. 175, pl. 15: figs 1-2, 4-9 (1992). 

Pinnularia schroeterae Krammer var. schroeterae, Diatoms of Europe 1, p. 225, pl. 88: figs 1-15, 27-29; pl. 89: fig. 1 

(2000). [Comment: on p. 225 erroneously all figures for Pinnularia schroeterae var. schroeterae and Pinnularia schroeterae 

var. elliptica are cited (pl. 88: figs 1-30, pl. 89: figs 1-6).] 

Pinnularia schroeterae var. elliptica Krammer, Diatoms of Europe 1, p. 225, pl. 88: figs 16-26, 30; pl.89: figs 2-6 (2000). 

[Comment: on p. 225 erroneously pl. 88: figs 27-29 is cited (pl. 88: figs 27-29 shows Pinnularia schroeterae var. 

schroeterae).] 

Pinnularia scotica Krammer, Diatoms of Europe 1, p. 214-215, pl. 22: figs 7-14 (2000). 

Pinnularia septentrionalis var. americana Krammer, Diatoms of Europe 1, p. 222, pl. 83: fig. 7 (2000). 

Pinnularia siebenkittlii Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 142-143, pl. 69: figs 1-2 (1995). 

Pinnularia similiformis Krammer, Biblioth. Diatomol. 26, p. 175, pl. 13: figs 9-13 (1992). 

Pinnularia similiformis var. koreana Metzeltin et Krammer, Diatoms of Europe 1, p. 212-213, pl. 16: figs 3-9 (2000). 

Pinnularia sinistra Krammer, Biblioth. Diatomol. 26, p. 175-176, pl. 37: figs 1-16 (1992). 

Pinnularia spinosissima Lange-Bert., U.Rumrich et Krammer, Iconogr. Diatomol. 9, p. 204-205, pl. 146: figs 1-8, pl. 147: 

figs 1-6, pl. 148: figs 1-4 (2000). 

Pinnularia staurocuneata Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 58 (1998). 

Replaced synonym: Pinnularia subcapitata [var. stauroneiformis] f. subcuneata Manguin ex Kociolek et Reviers, Cryptog. 

Algol. 17 (3), p. 206 (1996). 

Pinnularia stidolphii Krammer, Diatoms of Europe 1, p. 231, pl. 134: figs 1-7, pl. 183: fig. 3 (2000). 

Pinnularia stomatophora var. irregularis Krammer, Diatoms of Europe 1, p. 226, pl. 101: figs 4-10 (2000). 

Pinnularia stomatophora var. salina Krammer, Diatoms of Europe 1, p. 226, pl. 99: figs 7-9 (2000). 

Pinnularia streckerae Krammer, Diatoms of Europe 1, p. 225, pl. 88: figs 31-41, pl. 89: figs 7-9 (2000). 

Pinnularia streptoraphe var. parva Krammer, Biblioth. Diatomol. 26, p. 176, pl. 74: figs 4-5 (1992). 

Pinnularia streptoraphe var. savona Krammer, Diatoms of Europe 1, p. 238, pl. 205: fig. 1 (2000). 

Pinnularia subacuminata Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 187, pl. 180: figs 9-10(1998). 

Pinnularia subacuta var. cuneata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 188, pl. 192: figs 1-2 (1998). 

Pinnularia subanglica Krammer, Diatoms of Europe 1, p. 223, pl. 78: fig. 7, pl. 84: figs 1-7 (2000). 

Pinnularia subbacillum Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 188, pl. 180: figs 1-4 (1998). 

Pinnularia subboyeri Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 189, pl. 183: figs 11-12, pl. 199: fig. 4 (1998). 

[Comment: on p. 632 (= caption of pl. 199) the authors are cited as "Krammer et Metzeltin".] 

Pinnularia subcapitata var. elongata Krammer, Biblioth. Diatomol. 26, p. 108, 176, pl. 38: figs 1-11, pl. 39: figs 1-15 

(1992). 

Pinnularia subcapitata var. semicruciata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 190, pl. 173: figs 5-13 (1998). 

Pinnularia subcapitata var. subrostrata Krammer, nom. inval., Biblioth. Diatomol. 26, p. 177, pl. 38: figs 12-18 (1992). 

[Comment: the Latin description can not be treated as a diagnosis.] 

Pinnularia subcommutata var. nonfasciata Krammer, Diatoms of Europe 1, p. 229, pl. 120: figs 6-11 (2000). 



82 

Pinnularia subdivergens Krammer, Biblioth. Diatomol. 26, p. 176, pl. 29: figs 5-7 (1992). 

Pinnularia subfalaiseana Krammer, Diatoms of Europe 1, p. 224, pl. 78: figs 8-12 (2000). 

Pinnularia subflamma Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 190, pl. 189: figs 4-7 (1998). 

Pinnularia subgibba Krammer var. subgibba, Biblioth. Diatomol. 26, p. 176, pl. 46: figs 1-4 (1992). 

Pinnularia subgibba var. angustarea Krammer et Metzeltin, Iconogr. Diatomol. 5, p. 191, pl. 184: figs 1-3 (1998). 

Pinnularia subgibba var. capitata Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 191-192, pl. 168: fig. 7, pl. 176: figs 1-2, 

3 (1998). 

Pinnularia subgibba var. hustedtii Krammer, Biblioth. Diatomol. 26, p. 177, pl. 47: fig. 1 (1992). 

Pinnularia subgibba var. sublinearis Krammer, Diatoms of Europe 1, p. 220, pl. 65: fig. 14 (2000). 

Pinnularia subgibba var. tirolensis Metzeltin et Krammer, Iconogr. Diatomol. 2, p. 100, pl. 86: figs 7-8 (1996). 

Pinnularia subgibba var. undulata Krammer, Biblioth. Diatomol. 26, p. 176-177, pl. 46: fig. 5 (1992). 

Pinnularia subgigas Krammer, Diatoms of Europe 1, p. 237-238, pl. 200: figs 2-5 (2000). [Comment: on p. 237 erroneously 

pl. 200: fig. 1 is cited (pl. 200: fig. 1 shows Pinnularia cherryfieldiana).] 

Pinnularia submicrostauron S.Schroet. in Krammer, Biblioth. Diatomol. 26, p. 177, pl. 39: figs 16-25 (1992). 

Pinnularia subrhombica Krammer var. subrhombica, Diatoms of Europe 1, p. 229, pl. 119: figs 8-11, pl. 154: figs 5-7 

(2000). 

Pinnularia subrhombica var. angusta Krammer, Diatoms of Europe 1, p. 229, pl. 119: fig. 7 (2000). 

Pinnularia subrupestris Krammer var. subrupestris, Biblioth. Diatomol. 26, p. 177, pl. 53: figs 8-13 (1992). 

Pinnularia subrupestris var. cruciata Krammer, Diatoms of Europe 1, p. 228, pl. 115: figs 3-5 (2000). 

Pinnularia subrupestris var. cuneata Krammer, Diatoms of Europe 1, p. 228, pl. 102: figs 15-17, pl. 147: figs 6-10 (2000). 

Pinnularia subrupestris var. parva Krammer, Biblioth. Diatomol. 26, p. 177, pl. 59: figs 8-9 (1992). 

Pinnularia subsilvatica Krammer et Lange-Bert., Diatoms of Europe 1, p. 43, 212, pl. 17: figs 27-29 (2000). 

Pinnularia substreptoraphe var. bornholmiana Krammer, Diatoms of Europe 1, p. 238, pl. 201: figs 1-3 (2000). 

[Comment: on p. 238 erroneously pl. 202: figs 1-6 is cited (pl. 202: figs 1-6 shows Pinnularia substreptoraphe var. 

substreptoraphe).] 

Pinnularia subumbrosa Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 192, pl. 182: figs 1-3, 4, 5-6 (1998). 

Pinnularia tirolensis var. julma Krammer, Diatoms of Europe 1, p. 220, pl. 65: figs 6, 11-13 (2000). [Comment: on p. 388 

(= caption of pl. 65: figs 6, 11-13 the authors are cited as "Krammer & Metzeltin".] 

Pinnularia toscana Krammer, Diatoms of Europe 1, p. 235, pl. 175: figs 1-4 (2000). 

Pinnularia transversiformis Krammer, Diatoms of Europe 1, p. 237, pl. 188: figs 1-7, pl. 189: figs 1-5 (2000). 

Pinnularia tsoneka Lange-Bert. et Metzeltin, Iconogr. Diatomol. 9, p. 205, fig. 137: 3-4, pl. 138: figs 1-3 (2000). 

Pinnularia tumescens Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 193, pl. 186: figs 1-5 (1998). 

Pinnularia tynnii Krammer, Diatoms of Europe 1, p. 216, pl. 40: figs 4-6 (2000). 

Pinnularia umbrosa var. tropica Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 194, pl. 182: figs 7-9 (1998). 

Pinnularia undula var. mesoleptiformis Krammer, Diatoms of Europe 1, p. 226, pl. 93: fig. 1 (2000). 

Pinnularia vaetterana Krammer, Diatoms of Europe 1, p. 211, pl. 10: figs 6-7 (2000). 

Pinnularia vangariana Krammer, Diatoms of Europe 1, p. 234, pl. 170: fig. 2 (2000). [On p. 600 (= caption of pl. 170: fig. 

2 as "Pinnularia spec.", slide Zt2/64 HB is cited wherasslide Zt1/64 HB is indicated as the type.] 

Pinnularia variarea Metzeltin et Krammer, Iconogr. Diatomol. 5, p. 194, pl. 181: figs 8-10, pl. 193: fig. 5 (1998). 

Pinnularia viridiformis var. minor Krammer, Diatoms of Europe 1, p. 234, pl. 138: figs 1-5 (2000). 



83 

PINNUNAVIS Okuno 


Pinnunavis armoricana (Amossé) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 331 (2000). 

Basionym: Navicula armoricana Amossé, Bull. Soc. Sci. Nat. Quest. 5 (2), p. 18, pl. 1: fig. 2 (1932). 

[Comment: Misspelled as Pinnuavis in Iconogr. Diatomol. 7.] 

Pinnunavis genustriata (Hust.) Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 81 (1999). 

Basionym: Navicula genustriata Hust., Int. Rev. Gesamten Hydrobiol. Hydrogr. 42, p. 77, figs 143-144 (1942). 

[Comment: Misspelled as Pinnuavis in Iconogr. Diatomol. 6.] 

PLACONEIS Mereschk. 


Placoneis centropunctata (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 195 (1998). 

Basionym: Navicula centropunctata Hust., Rabenhorsts Kryptogamenflora 7 (3) 4. Lieferung, 677, fig. 1678 (1966). 

Placoneis chilensis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 206, pl. 59: figs 8-9 (2000). 

Placoneis conveniens (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 195 (1998). 

Basionym: Navicula conveniens Hust., Bot. Not. 105, p. 402, fig. 123 (1952). 

Placoneis dacostae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 196, pl. 77: fig. 1, pl. 90: figs 4-8 (1998). [Comment: 

Placoneis dacostae on p. 388 is misspelled as "P. dacostei ".] 

Placoneis demeraroides (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 196 (1998). 

Basionym: Navicula demeraroides Hust., Rabenhorsts Kryptogamenflora 7 (3) 4. Lieferung, 676, fig. 1677 (1966). 

Placoneis disparilis (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 197 (1998). 

Basionym: Navicula diparilis Hust., Rabenhorsts Kryptogamenflora 7 (3) 4. Lieferung, p. 674, fig. 1674 (1966). 

Placoneis elginensis var. undulata (Østrup) Lange-Bert., Iconogr. Diatomol. 3, p. 133 (1996). 

Basionym: Navicula dicephala var. undulata Østrup, Botany of Iceland 2, p. 25, pl. 3: fig. 33 (1918). 

[Nomenclatural synonym of Placoneis undulata (Østrup) Lange-Bert.] 

Placoneis explanata (Hust.) Lange-Bert., Iconogr. Diatomol. 9, p. 207 (2000). 

Basionym: Navicula explanata Hust., Schweiz. Z. Hydrol. 11, p. 202, figs 7-8 (1948). 

Placoneis ignorata (Schim.) Lange-Bert., Iconogr. Diatomol. 9, p. 207 (2000). 

Basionym: Navicula ignorata Schim., Nova Hedwigia 30, p. 585, pl. 6: figs 1-9 (1978). 

Placoneis jatobensis (Krasske) Metzeltin et Lange-Bert. var. jatobensis, Iconogr. Diatomol. 5, p. 197 (1998). 

Basionym: Navicula jatobensis Krasske, Arch. Hydrobiol. 44, p. 651, fig. 12 (1951). 

Placoneis jatobensis var. capitata (Krasske) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 197 (1998). 

Basionym: Navicula jatobensis var. capitata Krasske, Arch. Hydrobiol. 44, p. 651, fig. 13 (1951). 

Placoneis molesta Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 197, pl. 89: figs 6-9 (1998). 

Placoneis neotropica Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 198-199, pl. 89: figs 1-5 (1998). 

Placoneis paraelginensis Lange-Bert., Iconogr. Diatomol. 9, p. 208, pl. 60: figs 17-20 (2000). 

Placoneis pellaifa Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 210, pl. 60: figs 9-10 (2000). 

Placoneis pseudodemerarae (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 199 (1998). 

Basionym: Navicula pseudodemerarae Hust. in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 370: fig. 9 (1930). 

Placoneis santaremensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 199, pl. 75: figs 18-24 (1998). 

Placoneis scharfii Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 211, pl. 59: figs 5-7 (2000). 

Placoneis surinamensis (Cleve) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 200 (1998). 

Basionym: Navicula surinamensis Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26, p. 9, pl. 2: fig. 1 (1895). 

Placoneis tersa (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 200 (1998). 

Basionym: Navicula tersa Hust. in Gessner & Vareschi, Venezuela Expedition 1, p. 116, figs 38-39 (1956). 

Placoneis undulata (Østrup) Lange-Bert., Iconogr. Diatomol. 9, p. 212 (2000). 

Basionym: Navicula dicephala var. undulata Østrup, Botany of Island 2, p. 25, pl. 3: fig. 33 (1918). 

[≡ Placoneis elginensis var. undulata (Østrup) Lange-Bert.] 



84 

Placoneis vicina (Hust.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 201 (1998). 

Basionym: Navicula vicina Hust., Bot. Not. 105, p. 407, fig. 128 (1952). 

Placoneis zimmermannii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 201, pl. 89: figs 10-13 (1998). 

PLAGIOGRAMMOPSIS Hasle, Stosch et Syvertsem 


Plagiogrammopsis crawfordii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 39, 439, pl. 10: figs 37-39, pl. 

12: figs 15-18 (2000). 

PLANOTHIDIUM Round et Bukhtiyarova 


[http://128.32.109.44/cgi-bin/porp_cgi.plPLANOTHIDIUMFEROUNDLBUKHTIYAROVA1996]. 

Planothidium apiculatum (R.M.Patrick) Lange-Bert., Iconogr. Diatomol. 6, p. 274 (1999). 

Basionym: Achnanthes lanceolata var. apiculata R.M.Patrick, Farlowia 2, p. 167, pl. 1: figs 4-5 (1945). 

[≡ Achnantheiopsis apiculata (R.M.Patrick) Lange-Bert.] 

[≡ Achnanthes lanceolata ssp. apiculata (R.M.Patrick) Lange-Bert.] 

Planothidium aueri (Krasske) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes aueri Krasske, Ann. Acad. Scient. Fenn. Ser. A. IV. Biol. 14, p. 77, pl. 86: figs 23-32 (1949). 

[≡ Achnantheiopsis aueri (Krasske) Lange-Bert.] 

Planothidium biporomum (M.H.Hohn et Hellerman) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes biporoma M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 80, p. 273, pl. 2: figs 5-6 (1963). 

[≡ Achnantheiopsis biporoma (M.H.Hohn et Hellerman) Lange-Bert.] 

Planothidium borneolanceolatum (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnantheiopsis borneolanceolata Lange-Bert., Arch. Protistenk. 148, p. 208, (1997). 

[Comment: The date of publication is given in the 2nd edition of Iconogr. Diatomol. 6, p. 281 (1999).] 

[– Achnanthes borneolanceolata Lange-Bert., nom. inval.] 

Planothidium chilense (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes chilensis Hust., Arch. Hydrobiol. 18, p. 238, pl. 72: figs 3-4 (1927). 

[≡ Achnantheiopsis chilensis (Hust.) Lange-Bert.] 

Planothidium daui (Foged) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes daui Foged, Danmark Geol. Undersoeg. 84, p. 14, pl. 1: fig. 10 (1962). 

[≡ Achnantheiopsis daui (Foged) Lange-Bert.] 

Planothidium deperditum (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 119 (2000). 

Basionym: Cocconeis deperdita Giffen. Bot. Mar. 18, p. 78, figs 26-28 (1975). 

Planothidium diplopunctatum (Simonsen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 119 (2000). 

Basionym: Achnanthes diplopunctata Simonsen, Kieler Meeresforsch. 15, p. 76, pl. 10: figs 4-7 (1959). 

Planothidium dispar (Cleve) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 120 (2000). 

Basionym: Achnanthes dispar Cleve, Acta Soc. Fauna Fl. Fenn. 8 (2), p. 52, pl. 3: fig. 2 (1891). 

Planothidium distinctum (Messik.) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes distincta Messik., Hydrobiologia 6, p. 32, pl. 1: fig. 2 (1954). 

[≡ Achnantheiopsis distincta (Messik.) Lange-Bert.] 

Planothidium fonticolanceolatum (Lange-Bert. et Schim.) Lange-Bert., Iconogr. Diatomol. 6, p. 275 (1999). 

Basionym: Achnanthes fonticolanceolata Lange-Bert. et Schim., Biblioth. Diatomol. 18, p. 57, pl. 88: figs 1-7 (1989). 

[≡ Achnantheiopsis fonticolanceolata (Lange-Bert. et Schim.) Lange-Bert.] 

Planothidium frequentissimum (Lange-Bert.) Lange-Bert. var. frequentissimum, Iconogr. Diatomol. 6, p. 276 (1999). 

Basionym: Achnanthes lanceolata ssp. frequentissima Lange-Bert., Biblioth. Diatomol. 27, p. 4 (1993). 

[≡ Achnantheiopsis frequentissima (Lange-Bert.) Lange-Bert.] 

Planothidium frequentissimum var. magnum (F.Straub) Lange-Bert., Iconogr. Diatomol. 6, p. 276 (1999). 

Basionym: Achnanthes rostrata var. magna F.Straub, Bull. Soc. Neuchâteloise Soc. Nat. 108, p. 139, pl. 10: fig. 142, pl. 11: 

fig. 142b (1985). 

[≡ Achnantheiopsis frequentissima var. magna (F.Straub) Lange-Bert.] 

[≡ Achnanthes lanceolata var. magna (F.Straub) Lange-Bert.] 



85 

Planothidium frequentissimum var. minus (Schulz-Danzig) Lange-Bert., Iconogr. Diatomol. 6, p. 276 (1999). 


[≡ Achnanthes lanceolata var. minor (Schulz-Danzig) Lange-Bert.] 

[≡ Achnantheiopsis frequentissima var. minor (Schulz-Danzig) Lange-Bert.] 

Planothidium granum (M.H.Hohn et Hellerman) Lange-Bert., Iconogr. Diatomol. 6, p. 276 (1999). 

Basionym: Achnanthes grana M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82, p. 274, pl. 2: figs 9-10 (1963). 

[≡ Achnantheiopsis grana (M.H.Hohn et Hellerman) Lange-Bert.] 

Planothidium haynaldii (Schaarschm.) Lange-Bert., Iconogr. Diatomol. 6, p. 276 (1999). 

Basionym: Achnanthes haynaldii Schaarschm., Magyar. Növ. Lapok 5, p. 20 (1881). 

[≡ Achnantheiopsis haynaldii (Schaarschm.) Lange-Bert.] 

Planothidium heidenii (Schulz-Danzig) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 120 (2000). 

Basionym: Achnanthes heidenii Schulz-Danzig, Bot. Arch. 13, p. 190, fig. 36 (1926). 

Planothidium heteromorphum (Grunow) Lange-Bert., Iconogr. Diatomol. 6, p. 276 (1999). 

Basionym: Achnanthes heteromorpha Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 23 (1880). 

[≡ Achnantheiopsis heteromorpha (Grunow) Lange-Bert.] 

Planothidium holstii (Cleve) Lange-Bert., Iconogr. Diatomol. 6, p. 277 (1999). 

Basionym: Achnanthes holstii Cleve, Kongl. Svenska Vetenskapsakad. Handl. 18, p. 13, pl. 16: figs 6-7 (1881). 

[≡ Achnantheiopsis holstii (Cleve) Lange-Bert.] 

Planothidium infrequens Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 212-213, pl. 27: figs 3-9 (2000). 

Planothidium jan-marcinii (Witkowski, Metzeltin et Lange-Bert.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 

7, p. 121 (2000). 

Basionym: Achnanthes jan-marcinii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 2, p. 12, pl. 50: figs 5-9 

(1996). 

Planothidium joursacense (Héribaud) Lange-Bert., Iconogr. Diatomol. 6, p. 277 (1999). 

Basionym: Achnanthes joursacense Héribaud, Les Diatomées d’Auvergne, p. 89, pl. 12: fig. 20 (1903). 

[≡ Achnantheiopsis joursacense (Héribaud) Lange-Bert.] 

Planothidium lanceolatoides (Sovereign) Lange-Bert., Iconogr. Diatomol. 6, p. 277. 


[≡ Achnantheiopsis lanceolatoides (Sovereign) Lange-Bert.] 

[≡ Achnanthes lanceolata ssp. lanceolatoides (Sovereign) Lange-Bert.] 

Planothidium lanceolatum (Bréb. ex Kütz.) Lange-Bert., Iconogr. Diatomol. 6, p. 281 (1999). 

Basionym: Achnanthes lanceolatum Bréb. ex Kütz., Bot. Z. 4, p. 247 (1846). 

Planothidium lilljeborgei (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 121 (2000). 

Basionym: Achnanthes lilljeborgei Grunow, Bot. Zentralbl. 181, p. 68 (1881). 

Planothidium linkei (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 277 (1999). 

Basionym: Achnanthes linkei Hust., Abh. Naturwiss. Vereine Bremen 31, p. 607, figs 28-32 (1939). 

[≡ Achnantheiopsis linkei (Hust.) Lange-Bert.] 

Planothidium neglectum Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 213, pl. 30: figs 7-10 (2000). 

Planothidium magnificum (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 277 (1999). 

Basionym: Achnanthes magnifica Hust., Bot. Not. 4, p. 385, figs 62-65 (1952). 

[≡ Achnantheiopsis magnifica (Hust.) Lange-Bert.] 

Planothidium marinum (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 277 (1999). 

Basionym: Achnanthes marina Hust., A. Schmidt’s Atlas der Diatomaceenkunde, pl. 415: figs 1-2 (1937). 

[≡ Achnantheiopsis marina (Hust.) Lange-Bert.] 

Planothidium minutissimum (Krasske) Lange-Bert., Iconogr. Diatomol. 6, p. 278 (1999). 

Basionym: Achnanthes lanceolata var. minutissima Krasske, Arch. Hydrobiol. 33, p. 6, figs 18-22 (1938). 

[≡ Achnantheiopsis minutissima (Krasske) Lange-Bert.] 

Planothidium miotum (J.R.Carter) Lange-Bert., Iconogr. Diatomol. 6, p. 278 (1999). 

Basionym: Achnanthes miota J.R.Carter, Nova Hedwigia Beih. 73, p. 286, pl. 1: fig. 9 (1982). 

[≡ Achnantheiopsis miota (J.R.Carter) Lange-Bert.] 



86 

Planothidium oculatum (Hust.) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 122 (2000). 

Basionym: Achnanthes oculata Hust., Bot. Not. 1952, p. 390, figs 91-94 (1952). 

Planothidium oestrupii (Cleve-Euler) Round et Bukhtiyarova ex U.Rumrich, Lange-Bert. et M.Rumrich, Iconogr. Diatomol. 

9, p. 214 (2000). 

Basionym: Achnanthes lanceolata var. oestrupii Cleve-Euler, Acta Forest. Fenn. 22, p. 53, pl. 1: fig. 1 (1922). 

[≡ Achnantheiopsis oestrupii (Cleve-Euler) Lange-Bert.] 

Planothidium pericavum (J.R.Carter) Lange-Bert., Iconogr. Diatomol. 6, p. 278 (1999). 

Basionym: Achnanthes pericava J.R.Carter, Nova Hedwigia 11, p. 457, pl. 1: figs 5-8 (1966). 

[≡ Achnantheiopsis pericava (J.R.Carter) Lange-Bert.] 

Planothidium polaris (Østrup) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 123 (2000). 

Basionym: Achnanthes polaris Østrup, Meddel. Grønland 18, p. 85, pl. 7: fig. 86 (1895). 

Planothidium pseudotanense (Cleve-Euler) Lange-Bert., Iconogr. Diatomol. 6, p. 207 (1999). 

Basionym: Achnanthes pseudotanensis Cleve-Euler, Kongl. Svenska Vetenskapsakad. Handl. 4, p. 25, fig. 524 (1953). 

[≡ Achnantheiopsis pseudotanensis (Cleve-Euler) Lange-Bert.] 

Planothidium pungens (Cleve-Euler) Lange-Bert., Iconogr. Diatomol. 6, p. 278 (1999). 

Basionym: Achnanthes elliptica var. pungens Cleve-Euler, Soc. Scient. Fenn. Comm. Biol. 4, p. 38, fig. 40 (1934). 

[≡ Achnantheiopsis pungens (Cleve-Euler) Lange-Bert.] 

Planothidium quarnerensis (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 123 (2000). 

Basionym: Rhaphoneis quarnerensis Grunow, Verh. K. K. Zool.-Bot. Ges. Wien 12, p. 381, pl. 7: fig. 24 (1862). 

Planothidium rhombicuneatum Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 214-215, pl. 29: figs 1-12 (2000). 

Planothidium robustius Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Replaced synonym: Achnanthes lanceolata var. robusta Hust., Arch. Hydrobiol. Suppl. 15, p. 202, pl. 13: figs 47-49 (1934). 

[≡ Achnantheiopsis robustior Lange-Bert.] 

[≡ Achnanthes lanceolata ssp. robusta (Hust.) Lange-Bert.] 

Planothidium robustum (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Basionym: Achnanthes delicatula var. robusta Hust., Verh. Vortr. Bremer Wiss. Ges. 8-9, p. 378, figs 1-2 (1934). 

[≡ Achnantheiopsis robusta (Hust.) Lange-Bert.] 

Planothidium rostratum (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Basionym: Achnanthes rostrata Østrup, Flora of Koh Chang, Bot. Tidskr. 26, p. 253 ("1902", 1904). 

[≡ Achnantheiopsis rostrata (Østrup) Lange-Bert.] 

[≡ Achnanthes lanceolata ssp. rostrata (Østrup) Lange-Bert.] 

Planothidium salvadorianum (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Basionym: Achnanthes salvadoriana Hust., Com. Inst. Tripical Investig. Cient. 5-6, p. 132, fig. 1 (1953). 

[≡ Achnantheiopsis salvadoriana (Hust.) Lange-Bert.] 

Planothidium schwabei (Krasske) Lange-Bert., Iconogr. Diatomol. 9, p. 216 (2000). 

Basionym: Achnanthes schwabei Krasske, Arch. Hydrobiol. 35, p. 371, pl. 11: figs 12-14 (1939). 

Planothidium septentrionale (Østrup) Round et Bukhtiyarova ex U.Rumrich, Lange-Bert. et M.Rumrich, Iconogr. Diatomol. 

9, p. 214 (2000). 

Basionym: Achnanthes septentrionalis Østrup, Danske Diatomeer, p. 215, figs 13, 27 (1910). 

[≡ Achnantheiopsis septentrionalis (Østrup) Lange-Bert.] 

PRAVIFUSUS Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 73, 439-440 (2000). [Typus generis: 

Pravifusus hyalinus Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 73, 440 (2000).] 

Pravifusus hyalinus Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 73, 440, pl. 12: figs 13-14, pl. 27: figs 

18-23 (2000). [Pravifusus hyalinus is the type of the name of the genus PRAVIFUSUS Witkowski, Lange-Bert. et Metzeltin.] 

PROSCHKINIA Karaeva 

[http://www.bgbm.org/scripts/asp/IAPT/ncugentry.aspname=Proschkinia] 


Proschkinia bulnheimii var. belgica (Grunow) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 340 (2000). 

Basionym: Navicula bulnheimii var. belgica Grunow in Van Heurck, Type de Synopsis, no. 113, pl. 56: fig. 10 (1885). 



87 

PSAMMOTHIDIUM Bukhtiyarova et Round 


Psammothidium acidoclinatum (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Basionym: Achnanthes acidoclinata Lange-Bert., Iconogr. Diatomol. 2, p. 22, pl. 21: figs 22-24, pl. 113: figs 1-7 (1996). 

Psammothidium chlidanos (M.H.Hohn et Hellerman) Lange-Bert., Iconogr. Diatomol. 6, p. 279 (1999). 

Basionym: Achnanthes chlidanos M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82, p. 273, pl. 2: figs 1-2 (1963). 

Psammothidium daonense (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 280 (1999). 

Basionym: Achnanthes daonensis Lange-Bert., Biblioth. Diatomol. 18, p. 43, pl. 19: figs 27-32, pl. 25: figs 1-6 (1989). 

Psammothidium perpusillum (Østrup) Lange-Bert., Iconogr. Diatomol. 6, p. 280 (1999). 

Basionym: Pinnularia perpusilla Østrup, Danske Diatomeer, p. 96, pl. 3: fig. 62 (1910). 

[≡ Achnanthes perpusilla (Østrup) Lange-Bert.] 

Psammothidium rechtensis (Leclercq) Lange-Bert., Iconogr. Diatomol. 6, p. 280 (1999). 

Basionym: Achnanthes rechtensis Leclercq, Bull. Jard. Bot. Nat. Belg. 53, p. 501, pl. 53: figs 1-17 (1983). 

Psammothidium rosenstockii (Lange-Bert.) Lange-Bert., Iconogr. Diatomol. 6, p. 280 (1999). 

Basionym: Achnanthes rosenstockii Lange-Bert., Biblioth. Diatomol. 18, p. 131, pl. 61: figs 1-17 (1989). 

PSEUDENCYONEMA Krammer, Biblioth. Diatomol. 36, p. 156, (1997). [Typus generis: Cymbella rossii Krammer et 

Lange-Bert., Nova Hedwigia Beih. 106, p. 93-99 (1993).] 

PSEUDOGOMPHONEMA Medlin 


Pseudogomphonema monicae Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 25, pl. 58: figs 11-22, pl. 80: 

figs 1-4, pl. 81: fig. 2 (1996). 

Pseudogomphonema plinskii Witkowski, Metzeltin et Lange-Bert., Iconogr. Diatomol. 4, p. 26, pl. 58: figs 7-10, pl. 80: fig. 

5, pl. 81: figs 3-6 (1996). [Comment: Pseudogomphonema plinskii on p. 192 (= caption of pl. 80) is cited as "() 

Pseudogomphonema spec.", P. plinskii on p. 194 (= caption of pl. 81: figs 5-6) is cited as "Pseudogomphonema spec." ]. 

PULCHELLA Krammer, Diatoms of Europe 1, p. 240 (2000). [Typus generis: Pinnularia kriegeriana Krasske, Ber. 

Deutsch. Bot. Ges. 61, p. 86, fig. 17 (1943).] 

Pulchella kriegeriana (Krasske) Krammer, Diatoms of Europe 1, p. 205 (2000). 

Basionym: Pinnularia kriegeriana Krasske, Ber. Deutsch. Bot. Ges. 61, p. 86, fig. 17 (1943). 

[≡ Biremis kriegeriana (Krasske) Lange-Bert.] 

RHOICOSPHENIA Grunow 


Rhoicosphenia abbreviata (C.Agardh) Lange-Bert., Bot. Not. 133, p. 586 (1980). 

Basionym: Gomphonema abbreviatum C.Agardh, Conspectus Criticus Diatomacearum 2, p. 34 (1831). 

RHOICOSPHENULA Lange-Bert. (subgenus within GOMPHOSPHENIA), Nova Hedwigia 60, p. 244, 246, pl. 1-3. 

Typus subgeneris: Gomphosphenia paradoxa Lange-Bert., Nova Hedwigia 60, p. 246, 248 (1995). 

RHOPALODIA O.Müll. 


Rhopalodia brebissonii Krammer, Biblioth. Diatomol. 15, p. 76, pl. 48: figs 7-10 (1987). [As a part of the protologue, 

Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 113: figs 7-13, pl. 113A: 

figs 7-12 is cited.] 

Rhopalodia gibba var. minuta Krammer, Biblioth. Diatomol. 15, p. 79-80, pl. 45: figs 1-6 (1987). [As a part of the 

protologue, Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 111A: figs 2-7 

is cited.] 

Rhopalodia globosa (Hust.) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 215 (1998). 

Basionym: Rhopalodia gibberula var. globosa Hust., Arch. Hydrobiol. Suppl. 15, p. 458, pl. 14: fig. 15 (1938). 

Rhopalodia pacifica Krammer, Biblioth. Diatomol. 15, p. 82-83, pl. 61: figs 9-13 (1987). 

Rhopalodia rumrichiae Krammer, Biblioth. Diatomol. 15, p. 83-84, pl. 62: figs 5-12 (1987). [Comment: on p. 83 pl. 62: fig. 

5 is not referred to. Rhopalodia rumrichiae on p. 83 and 270 misspelled as "Rhopalodia rumrichae".] 



88 

Rhopalodia sculpta Krammer, Biblioth. Diatomol. 15, p. 84, pl. 62: figs 1-4 (1987). 

Rhopalodia sterrenburgii Krammer, Biblioth. Diatomol. 15, p. 85, pl. 61: figs 5-8 (1987). 

SELLAPHORA Mereschk. 


Sellaphora alastos (M.H.Hohn et Hellerman) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 101(1996). 

Basionym: Navicula americana var. alastos M.H.Hohn et Hellerman, Trans. Amer. Microscop. Soc. 82 (3), p. 290, pl. 2: fig. 

36 (1963). 

Sellaphora amoena Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 202, pl. 84: fig. 4, pl. 85: figs 10-12 (1998). 

Sellaphora costei Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 203, pl. 85: figs 1-3 (1998). 

Sellaphora fusticulus (Østrup) Lange-Bert., Iconogr. Diatomol. 9, p. 216 (2000). 

Basionym: Navicula wittrockii f. fusticulus Østrup, Danske Diatoméer, p. 36, pl. 1: fig. 19 (1910). 

Sellaphora gibbula Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 204, pl. 85: figs 6-7 (1998). 

Sellaphora gregoryana (Cleve et Grunow) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 204 (1998). 

Basionym: Navicula bacillum var. gregoryana Cleve et Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, p. 44 (1880). 

Sellaphora guyanensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 205, pl. 100: figs 9-14, pl. 164: fig. 1 (1998). 

Sellaphora illustris Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 205-206, pl. 82: figs 1-2, pl. 84: figs 5-6 (1998). 

[Comment: Sellaphora illustris is not documented on pl. 82: figs 8-9 as cited on p. 205.] 

Sellaphora lambda (Cleve) Metzeltin et Lange-Bert. var. lambda, Iconogr. Diatomol. 5, p. 206 (1998). 

Basionym: Navicula lambda Cleve, Kongl. Svenska Vetenskapsakad. Handl. 26, pl. 5: fig. 19 (1894). 

Sellaphora lambda var. densistriata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 207, pl. 82: figs 3-4, pl. 84: fig. 7 

(1998). 

Sellaphora laterostellata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 208, pl. 82: figs 5-6 (1998). 

Sellaphora laterostrata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 208-209, pl. 81: figs 21-23 (1998). 

Sellaphora mutata (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 131 (1996). 

Basionym: Navicula mutata Krasske, Bot. Arch. 27, p. 354, fig. 16 (1929). 

[– Sellaphora mutata (Krasske) Lange-Bert., nom. inval., Iconogr. Diatomol. 2, p. 101 (1996).] 

Sellaphora parapupula Lange-Bert., Iconogr. Diatomol. 2, p. 101 (1996). 

Replaced synonym: Navicula pupula var. capitata Hust., Süßwasserflora Mitteleuropas 10, p. 281, fig. 467c (1930). 

Sellaphora pseudopupula (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 138 (1996). 

Basionym: Navicula pseudopupula Krasske, Bot. Arch. 3, p. 197, fig. 4 (1923). 

Sellaphora pupula var. densistriata Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 102, pl. 25: fig. 9 (1996). 

Sellaphora rectangularis (W.Greg.) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 102 (1996). 

Basionym: Stauroneis rectangularis W.Greg., Quart. J. Microscop. Sci. 2, p. 99, pl. 4: fig. 17 (1854). 

Sellaphora reviersii Lange-Bert. et Gerd Moser, Biblioth. Diatomol. 32, p. 135-136, pl. 63: figs 5-8 (1995). 

Sellaphora riojae (Cleve) E.Reichardt, Diatom Res. 12, p. 309 (1997). 

Basionym: Navicula riojae Cleve, Öfvers. Förh. Kongl. Svenska Vetensk.-Akad. 38 (10), p. 12, pl. 16: fig. 2 (1881). 

Sellaphora subdeliberata E.Reichardt, Diatom Res. 12, p. 311 (1997). 

[Comment: Intended validation of Navicula subdeliberata Manguin, nom. inval., and new combination with Sellaphora. 

Latin diagnosis by E.Reichard, pl. 25: fig. 8 in Manguin, Mém. Mus. Natl. Hist. Nat., B, Bot. 12 (1964) is designated as the 

holotype "iconotype".] 

[– Navicula subdeliberata Manguin, nom. inval., Mém. Mus. Natl. Hist. Nat., B, Bot. 12, p. 75, pl. 7: fig. 1, pl. 25: fig. 8 

(1964).] 

Sellaphora sublinearis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 209, pl. 81: figs 7-8 (1998). 

Sellaphora tapajosensis Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 210, pl. 81: figs 5-6 (1998). 

Sellaphora tau (Cleve) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 211 (1998). 

Basionym: Navicula tau Cleve, Le Diatomiste 2, p. 14, pl. 1: fig. 11 (1893). 



89 

Sellaphora tenerrima Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 211, pl. 81: figs 13-15 (1998). 

Sellaphora triundulata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 212, pl. 81: figs 16-19 (1998). 

Sellaphora ventraloconfusa (Lange-Bert.) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 212 (1998). 

Basionym: Navicula ventraloconfusa Lange-Bert., Biblioth. Diatomol. 18, p. 165, pl. 79: figs 37-38 nec 36 (1989). 

[≡ Naviculadicta ventraloconfusa (Lange-Bert.) Lange-Bert., Biblioth. Diatomol 29, p. 87 (1994).] 

Sellaphora verecundiae Lange-Bert., Biblioth. Diatomol. 29, p. 93, pl. 52: figs 19-28 (1994). 

SEMINAVIS D.G.Mann 


[http://128.32.109.44/cgi-bin/porp_cgi.plSEMINAVISDGMANN1990]. 

Seminavis barbara Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 348, 440, pl. 166: figs 1-4 (2000). 

SIEMINSKIA Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 213-214 (1998). [Typus generis: Sieminskia wohlenbergii 

(C.Brockmann) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 215 (1998).] 

Sieminskia wohlenbergii (C.Brockmann) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 215 (1998). 

Basionym: Navicula wohlenbergii C.Brockmann, Abh. Senckenb. Naturf. Ges. 478, p. 16, pl. 4: figs 4-6 (1950). 

Sieminskia zeta (Cleve) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 215 (1998). 

Basionym: Navicula zeta Cleve, Kongl. Svenska Vetenskapsakad. Handl. 27, p. 42 (1895). 

SIMONSENIA Lange-Bert., Bacillaria 2, p. 131. [Typus generis: Simonsenia delognei (Grunow) Lange-Bert., Bacillaria 2, 

p. 132 (1979).] 

Simonsenia delognei (Grunow) Lange-Bert. ssp. delognei, Bacillaria 2, p. 132 (1979). 

Basionym: Nitzschia delognei Grunow in Van Heurck, Synopsis des Diatomées de Belgique, Suppl. C, fig. 38 (1883). 

Simonsenia delognei ssp. rossii Lange-Bert. et Krammer, Nova Hedwigia Beih. 106, p. 122, figs 1-10 (1993). 

STAURONEIS Ehrenb. 


Stauroneis acidobionta Lange-Bert. et Wydrzycka, Brenesia 55-56, p. 15, pl. 9: figs 1-10 (2001). 

[– Navicula acidobionta Lange-Bert., nom. inval.] 

Stauroneis angustilancea Metzeltin et Lange-Bert., nom. inval., Iconogr. Diatomol. 5, p. 216, pl. 98: fig. 4 (1998). 

[Comment: Stauroneis angustilancea is lacking a type and a Latin diagnosis, hence the name is invalidly published. 

Stauroneis angustilancea was documented already in Lange-Bertalot & Metzeltin (1996), Iconogr. Diatomol. 2, p. 105 pl. 

34: figs 1-3 as "Stauroneis (nov.) spec. Nr. 3 Julma Ölkky".] 

Stauroneis angustevittata E.Reichardt, Nova Hedwigia 47, p. 93 (1988). 

Replaced synonym: Stauroneis anceps var. javanica Hust., Arch. Hydrobiol. Suppl.14, p. 154, pl. 1: fig. 10 (1935). 

Stauroneis aueri (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 98 (1996). 

Basionym: Navicula aueri Krasske, Ann. Acad. Sient. Fenn. Ser. A, IV. Biol. 14, p. 83, figs 8-9 (1949). 

Stauroneis brevis (Dippel) Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 216 (1998). 

Basionym: Stauroneis phoenicenteron var. brevis Dippel, Diatomeen der Rhein-Main-Ebene, p. 82, fig. 176 (1904). 

Stauroneis cataractae Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 215-216, pl. 31: figs 11-14 (1998). 

Stauroneis circumborealis Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 90, pl. 34: figs 1-4 (1999). 

Stauroneis conspicua Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 216, pl. 98: fig. 5 (1998). 

Stauroneis dubitabilior Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 217, pl. 98: figs 16-19 (1998). 

Stauroneis gracilior E.Reichardt, Iconogr. Diatomol. 1, p. 17 (1995). 

Replaced synonym: Stauroneis anceps f. gracilis Rabenh., Flora europaea algarum aquae dulcis et submarinae 1, p. 247 

(1864). 

[Comment: in Iconogr. Diatomol. 1 (second edition) (1995), p. 34 this combination was again marked as a new.] 

Stauroneis heinii Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 91, pl. 27: figs 1-4 (1999). 

Stauroneis hyperborea Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 92 (1999). 

Replaced synonym: Stauroneis obtusa var. lapponica Hust., Naturwiss. Unters. d. Sarekgebirges in Schwedisch Lappland 3 

(Botanik), p. 563 (1924). 



90 

Stauroneis koeltzii Metzeltin, Witkowski et Lange-Bert., Iconogr. Diatomol. 4, p. 26-27, pl. 10: figs 5-7 (1996). 

Stauroneis koniamboensis (Manguin ex Kociolek et Reviers) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, 

p. 59 (1998). 

Basionym: Navicula koniamboensis Manguin ex Kociolek et Reviers, Cryptog. Algol. 17 (3), p. 202 (1996). 

Stauroneis leguminiformis Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 93, pl. 33: figs 1-3 (1999). 

Stauroneis leguminopsis Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 94 (1999). 

Replaced synonym: Stauroneis legumen f. parva Grunow in Van Heurck, Synopsis des Diatomées de Belgique, pl. 4: fig. 11 

(1880). 

Stauroneis nebulosa (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 132 (1996). 

Basionym: Navicula nebulosa Krasske, Ann. Acad. Sient. Fenn. Ser. A, IV. Biol. 14, p. 84, figs 21a, b (1949). 

Stauroneis neofossilis Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 103, pl. 34: figs 4-5 (1996). 

Stauroneis neohyalina Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 104 (1996). 

Replaced name: Stauroneis anceps var. hyalina M.Perag. et Brun in Héribaud, Les Diatomees d’Auvergne, p. 78, pl. 3: fig. 

19 (1893). 

Stauroneis punensis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 217, pl. 88: figs 1-4, (5) (2000). 

Stauroneis resoluta Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 217, pl. 31: fig. 15, pl. 32: figs 1-6 

(1998). 

Stauroneis schimanskii Krammer, Biblioth. Diatomol. 9, p. 115-116, pl. 5: figs 1-6 (1985). [As a part of the protologue, 


Stauroneis siberica (Grunow) Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 104 (1996). 

Basionym: Stauroneis anceps var. siberica Grunow, Kongl. Svenska Vetenskapsakad. Handl. 17, 48, pl. 3: fig. 65 (1880). 

Stauroneis subgracilis Lange-Bert. et Krammer, Iconogr. Diatomol. 6, p. 96, pl. 29: figs 1-7 (1999). 

Stauroneis tackei (Hust.) Krammer et Lange-Bert. in Jahn et al., Lange-Bertalot-Festschrift, p. 633 (2001). 

Basionym: Navicula tackei Hust., Abh. Naturwiss. Vereine Bremen 32, p. 194, figs 6-7 (1942). 

[– Stauroneis tackei (Hust.) Krammer et Lange-Bert., nom. inval., Biblioth. Diatomol. 9, p. 116 (1985).] 

[Comment: according to Simonsen, 1987, Atlas and Catalogue of the Diatom Types of Friedrich Hustedt 1, p. 305 the 

holotype is N10/84.] 

Stauroneis valderostrata Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 218-219, pl. 95: fig. 1, pl. 98: figs 11-15 (1998). 

Stauroneis ventriosus Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 219, pl. 98: figs 1-2 (1998). 

STAURONELLA Mereschk. 


Stauronella amphoroides (Grunow) Lange-Bert., Iconogr. Diatomol. 6, p. 99 (1999). 

Basionym: Stauroneis amphoroides Grunow in A. Schmidt’s Atlas der Diatomaceenkunde, pl. 26: figs 37-39 (1875). 

Stauronella arctica (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 99 (1999). 

Basionym: Stauroneis arctica Hust. in Rabenhorsts Kryptogamenflora 7 (2) 6. Lieferung, p. 829, fig. 1171 (1959). 

Stauronella decipiens (Hust.) Lange-Bert., Iconogr. Diatomol. 6, p. 99 (1999). 

Basionym: Stauroneis decipiens Hust. in Rabenhorsts Kryptogamenflora 7 (2) 6. Lieferung, p. 827, fig. 1170 (1959). 

Stauronella indubitabilis Lange-Bert. et Genkal, Iconogr. Diatomol. 6, p. 100, pl. 36: figs 1-8 (1999). 

STAUROPHORA Mereschk. 


Staurophora anuschkae Witkowski, Iconogr. Diatomol. 7, p. 440-441, pl. 150: figs 1-15 (2000). 

STAUROSIRA Ehrenb. 


Staurosira altiplanensis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 220, pl. 14: figs 1-8 (2000). 

Staurosira aventralis Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 221, pl. 11: figs 1-8 (2000). 



91 

Staurosira berolinensis (Lemmerm.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 587 (2000). 

Basionym: Synedra berolinensis Lemmerm., Ber. Deutsch. Bot. Ges. 18, p. 31 (1900). 

[≡ Fragilaria berolinensis (Lemmerm.) Lange-Bert.] 

Staurosira construens var. exigua (W.Sm.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 585 (2000). 

Basionym: Triceratium exiguum W.Sm., Synopsis British Diatomaceae, p. 87 (1856). 

Staurosira lapponica (Grunow) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 586-587 (2000). 

Basionym: Fragilaria lapponica Grunow in Van Heurck, Synopsis des Diatomées des Belgique, Atlas, pl. 45: fig. 35 (1881). 

Staurosira laucensis Lange-Bert. et U.Rumrich var. laucensis, Iconogr. Diatomol. 9, p. 222-223, pl. 13: figs 10-20, 22-23 

(2000). 

Staurosira laucensis var. vulpina Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 223, pl. 10: figs 1-11 (2000). 

Staurosira martyi (Hérib.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 586. 

Basionym: Opephora martyi Hérib., Diatomées d’Auvergne, p. 43, pl. 8: fig. 20 (1903). 

[≡ Fragilaria martyi (Hérib.) Lange-Bert.] 

Staurosira mercedes Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 224, pl. 10: figs 12-14 (2000). [Comment: valid 

name on species rank for Fragilaria brevistriata var. trigona Lange-Bert., nom. inval., in Krammer & Lange-Bertalot, 

Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (3), p. 464, pl. 117: fig. 7b (1991).] 

Staurosira microstriata (Marciniak) Lange-Bert., Iconogr. Diatomol. 9, p. 225 (2000). 

Basionym: Fragilaria microstriata Marciniak, Acta Geol. Acad. Sci. Hung. 25, p. 165, pl. 2: figs 5-6 (1982). 

Staurosira oldenburgiana (Hust.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 587 (2000). 

Basionym: Fragilaria oldenburgiana Hust., Veröff. Inst. Meeresforsch. Bremerhaven 6, p. 29, pl. 1: figs 20-21 (1959). 

Staurosira pinnata var. trigona (Krasske) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 586 (2000). 

Basionym: Fragilaria harrisonii var. trigona Krasske, Arch. Hydrobiol. 24, p. 437, fig. 7 (1932). 

[≡ Fragilaria leptostauron var. trigona (Krasske) Lange-Bert. et Willmann.] 

Staurosira proboscoidea Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 226, pl. 10: figs 15-19 (2000). 

Staurosira pseudoconstruens (Marciniak) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 587 (2000). 

Basionym: Fragilaria pseudoconstruens Marciniak, Acta Geol. Acad. Sci. Hung. 25, p. 163, pl. 1: figs 1-2 (1982). 

Staurosira punctiformis Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 78, 441, pl. 24: figs 51-58, pl. 26: 

figs 1-2 (2000). [Comment: on p. 78 and 490 (caption of pl. 24) and 494 (caption of pl. 26) the authors are cited as 

"Witkowski, Metzeltin et Lange-Bert.".] 

Staurosira robusta (Fusey) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 587 (2000). 

Basionym: Fragilaria construens [var. binodis] f. robusta Fusey, Bull. Microscop. Appl. 2. Sér., 1 (2), p. 34, pl. 1: figs 2 

(1951). 

Staurosira subsalina (Hust.) Lange-Bert., Süßwasserflora von Mitteleuropa. 2 (3), p. 585 (2000). 

Basionym: Fragilaria construens var. subsalina Hust., Mitt. Geogr. Ges. Naturhist. Mus. Lübeck 30, p. 106, figs 3-8 (1925). 

STENONEIS Cleve 


Stenoneis wojtek-kowalskii Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 353, 441, pl. 156: figs 20-21 

(2000). 

STENOPTEROBIA Bréb. ex VanHeurck 


Stenopterobia capitata (Fontell) Lange-Bert. et Metzeltin, Iconogr. Diatomol. 2, p. 106 (1996). 

Basionym: Stenopterobia intermedia var. capitata Fontell, Ark. Bot. 14, p. 46, pl. 2: fig. 46 (1917). 

Stenopterobia crassior (N.Carter) Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 218 (1998). 

Basionym: Stenopterobia intermedia var. crassior N.Carter, J. Linnean Society 46, p. 60, fig. 1 J (1922). 

Stenopterobia eximia Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 218-219, pl. 68: figs 2-4, (5) 

(1998). 

Stenopterobia krammeri Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 220, pl. 217: figs 1-6 (1998). 

Stenopterobia minuscula Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 219-220, pl. 67: figs 3-5 (1998). 



92 

Stenopterobia muscicola (Krasske) Lange-Bert., Iconogr. Diatomol. 3, p. 180 (1996). 

Basionym: Surirella arctissima var. muscicola Krasske, Svensk Bot. Tidskr. 42, p. 439 (1948). 

Stenopterobia planctonica Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 221, pl. 210: fig. 4, pl. 219: fig. 3 (1998). 

Stenopterobia pumila Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 227, pl. 196: figs 1-7, pl. 197: figs 8-9 (2000). 

Stenopterobia ruppeliana Gerd Moser, Lange-Bert. et Steindorf, Biblioth. Diatomol. 32, p. 147-148, pl. 74: figs 1-8 (1995). 

Stenopterobia scolopendra Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 220-221, pl. 69: figs 3-4 

(1998). 

SURIRELLA Turpin 


Surirella brebissonii Krammer et Lange-Bert. var. brebissonii, Diatom Res. 2, p. 82, 85, figs 1, 4-5, 9, 21-33 (1987). 

Surirella brebissonii var. kuetzingii Krammer et Lange-Bert., Diatom Res. 2, p. 85-86, figs 52-68 (1987). 

Surirella brebissonii var. punctata Krammer, Biblioth. Diatomol. 15, p. 95, pl. 55: figs 1-10 (1987). [As a part of the 

protologue, Krammer & Lange-Bertalot (1988), Süßwasserflora von Mitteleuropa, Bacillariophyceae 2 (2), pl. 127: figs 9-13 

is cited.] 

Surirella bryophila Lange-Bert., Wydrzycka et Metzeltin, Brenesia 55-56, p. 16, pl. 20: figs 1-10 (2001). 

Surirella citrus Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 222, pl. 213: fig. 1 (1998). [Comment: Surirella citrus on 

p. 660 is cited as "S. citreus" (caption of pl. 213: fig. 1).] 

Surirella desikacharyi Lange-Bert. et Metzeltin, Nova Hedwigia Beih. 112, p. 322, figs 1-12 (1996). 

Surirella gotohiana Gerd Moser, Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 221-222, pl. 70: figs 1-3 (1998). 

Surirella lacus-vulcani Metzeltin, Lange-Bert. et Krammer, Iconogr. Diatomol. 2, p. 107, pl. 106: figs 17-18 (1996). 

Surirella marina Krammer, Biblioth. Diatomol. 15, p. 98-99, pl. 54: figs 1-9 (1987). 

Surirella muscicola (Krasske) Lange-Bert. et Metzeltin, Biblioth. Diatomol. 38, p. 222 (1998). 

Basionym: Surirella arctissima var. muscicola Krasske, Svensk Bot. Tidskr. 42, p. 439 (1948). 

Surirella patrimonii Sterrenburg, Biblioth. Diatomol. 15, p. 101-102, pl. 57: figs 1, 4-5 (1987). 

Surirella rottii Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 223, pl. 208: figs 1-3, pl. 211: fig. 2 (1998). 

Surirella rumrichorum Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 224, pl. 219: fig. 7, pl. 216: figs 2-3 (1998). 

Surirella schuwerkii E.Reichardt, Mem. Calif. Acad. Sci. 17, p. 130: figs 15-26 (1994). 

Surirella susanae Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 226, pl. 220: figs 1-3 (1998). 

Surirella tenerisilex Metzeltin et Lange-Bert., Iconogr. Diatomol. 5, p. 225, pl. 209: figs 4-6 (1998). 

Surirella terricola Lange-Bert. et Alles, Iconogr. Diatomol. 2, p. 107-108, pl. 106: fig. 16 (1996). [Comment: validation of 

Surirella terricola Lange-Bert. et Alles, nom. inval., Biblioth. Diatomol. 27, p. 159, pl. 53: figs 1-2'' (1993).] 

Surirella turkanensis E.Reichardt, Mem. Calif. Acad. Sci. 17, p. 129-130: figs 1-14 (1994). 

SYNEDRA Ehrenb. 


Synedra lata (Giffen) Witkowski, Iconogr. Diatomol. 7, p. 81 (2000). 

Basionym: Synedra tabulata var. lata Giffen, Bacillaria 3, p. 153, figs 40-41 (1980). 

TABELLARIA Ehrenb. ex Kütz. 


Tabellaria flocculosa var. andina Lange-Bert., Biblioth. Diatomol. 27, p. 160, pl. 11: figs 1-9 (1993). 

ULNARIA (Kütz.) Lange-Bert. et Compère (subgenus within FRAGILARIA), Diatom Res. 16, p. 103 (2001). 

Basionym: SYNEDRA [unranked] ULNARIA Kütz., Die kieselschaligen Bacillarien oder Diatomeen, p.65 (1844). [Synedra 

ulna (Nitzsch) Ehrenberg is the type of the name of the subgenus.] 



93 

ULNARIA (Kütz.) Compère in Jahn et al., Lange-Bertalot-Festschrift, p. 100 (2001). 

Basionym: SYNEDRA [unranked] ULNARIA Kütz., Die kieselschaligen Bacillarien oder Diatomeen, p.65 (1844). [Synedra 

ulna (Nitzsch) Ehrenberg is the type of the name of the genus.] 

VEIGALUDWIGIA Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 228-229 (2000). 

[Typus generis: Veigaludwigia urbana (Krasske) Lange-Bert. et U.Rumrich, Iconogr. Diatomol. 9, p. 228 (2000).] 

Veigaludwigia urbana (Krasske) Lange-Bert. et U. Rumrich, Iconogr. Diatomol. 9, p. 228 (2000). 

Basionym: Navicula urbana Krasske, Arch. Hydrobiol. 35, p. 384, pl. 12: fig. 14 (1939). 

Veigaludwigia willeri (Krasske) Lange-Bert. et U. Rumrich, Iconogr. Diatomol. 9, p. 229 (2000). 

Basionym: Navicula willeri Krasske, Arch. Hydrobiol. 35, p. 389, pl. 12: figs 3-4 (1939). 

VIKINGEA Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 125, 442 (2000). [Typus generis: Vikingea 

gibbocalyx (Brun) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 125 (2000).] 

Vikingea gibbocalyx (Brun) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 125 (2000). 

Basionym: Cocconeis gibbocalyx Brun, Mem. Soc. Phys. Hist. Nat. Genève 31 (1), p. 17, pl. 18: fig. 2 (1891). 

Vikingea promunturii (Giffen) Witkowski, Lange-Bert. et Metzeltin, Iconogr. Diatomol. 7, p. 126 (2000). 

Basionym: Achnanthes promunturii ["promonturii "] Giffen, Bot. Mar. 13, p. 88, figs 1-4 (1970). 

References 

Alles, E., Nörpel-Schempp, M. & Lange-Bertalot, H. (1991): Zur Systematik und Ökologie charakteristischer Eunotia-Arten. 

(Bacillariophyceae) in elektrolytarmen Bachoberläufen. – Nova Hedwigia 53: 171-213. 

Denys, L. & Lange-Bertalot, H. (1998): Observations on two taxa of the section Nitzschiae Lanceolatae (Bacillariophyceae): 

Nitzschia blankaartensis sp. nov. and N. bulnheimiana. – Nova Hedwigia 67: 247-258. 

Fourtanier, E. & Kociolek, P. (1999): Catalogue of the diatom genera. – Diatom. Res. 14: 1-190. 

Greuter, W., McNeill, J., Barrie, F. R., Burdet, H. M., Demoulin, V., Filgueiras, T. S., Nicolson, D.H., Silva, P.C., Skog, 

J.E., Trehane, P., Turland, N.J., Hawksworth D.L. (eds & compilers, 2000): International Code of Botanical 

Nomenclature (Saint Louis Code) adopted by the Sixteenth International Botanical Congress St. Louis, Missouri, July - 

August 1999. – Regnum Veg. 138: XVIII, 1-474. [http://www.bgbm.fuberlin.de/iapt/nomenclature/code/SaintLouis/0000St.Luistitle.htm] 

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[http://ucjeps.herb.berkeley.edu/INA.html] 

Jahn, R., Kociolek, J.P., Witkowski, A. & Compère, P. 2001 (eds): Lange-Bertalot-Festschrift: Studies on Diatoms. Gantner, 

Ruggell. 

Krammer, K. (1980): Morphologic and taxonomic investigations of some freshwater species of the diatom genus Amphora 

Ehr. – Bacillaria 3: 197-225. 

Krammer, K. (1981): Morphologic investigations of the valve and girdle of the diatom genus Cymbella Agardh. – Bacillaria 

4: 125-146. 

Krammer, K. (1992): Pinnularia Eine Monographie der europäischen Taxa. – Biblioth. Diatomol. 26: 1-353. 

Krammer, K. (1997): Taxonomische Probleme bei ähnlichen syntopischen Sippen am Beispiel einer Encyonema-Assoziation 

(Bacillariophyceae) aus dem Kivu-See, Zaire. – Nova Hedwigia 65: 131-146. 

Krammer, K. (1997a): Die cymbelloiden Diatomeen. Eine Monographie der weltweit bekannten Taxa. Teil 1. Allgemeines 

und Encyonema Part. – Biblioth. Diatomol. 36: 1-382. 

Krammer, K. (1997b): Die cymbelloiden Diatomeen. Eine Monographie der weltweit bekannten Taxa. Teil 2. Encyonema 

Part., Encyonopsis and Cymbellopsis. – Biblioth. Diatomol. 37: 1-469. 

Krammer, K. (1999a): Validierung von Cymbopleura nov. gen.– Iconogr. Diatomol. 6: 284. 

[Comment: This original article was published with different page numbers in: Iconogr. Diatomol. 6 (2 nd edition; “2 nd 

corrected printing”): 292.] 



94 

Krammer, K. (1999b): Validierung von Cymbopleura nov. gen.– Iconogr. Diatomol. 6 (2 nd corrected printing): 292. 

[Comment: This article is a reprint of the original article with different page number.] 

Krammer, K. (2000): The genus Pinnularia. – Diatoms of Europe 1: 1-703. Gantner Verlag, Ruggell. 

[Comment: Prior to the officially distributed corrected issue (January 2001) an original issue was distributed inofficially 

in a few copies (October 2000); nevertheless it is effectively published. Original issue and corrected issue look 

superficially the same and have the same ISBN number (ISBN 3-904144-24-3), but they differ in the protologues in a 

few cases. Plate 11 is missing in the original issue.] 

Krammer, K. (2001 "2000"): The genus Pinnularia. – Diatoms of Europe 1: 1-703. Gantner Verlag, Ruggell. 

[Comment: Prior to the officially distributed corrected issue (January 2001) an original issue was distributed inofficially 

in a few copies (October 2000); nevertheless it is effectively published. Original issue and corrected issue look 

superficially the same and have the same ISBN number (ISBN 3-904144-24-3), but they differ in the protologues in a 

few cases.] 

Krammer, K. (2001): Taxonomy and morphology of Brevisira arentii (Kolbe) Krammer gen. nov., comb. nov. – In: Jahn, R., 

Kociolek, J.P., Witkowski, A. & Compère, P. (eds): Lange-Bertalot-Festschrift: 9-20. Gantner, Ruggell. 

Krammer, K. (2002): Cymbella. – Diatoms of Europe 3: 1-584. Gantner Verlag, Ruggell. 

Krammer, K. & Lange-Bertalot, H. (1985): Naviculaceae. Neue und wenig bekannte Taxa, neue Kombinationen und 

Synonyme sowie Bemerkungen zu einigen Gattungen. – Biblioth. Diatomol. 9: 1-230. 

Krammer, K. & Lange-Bertalot, H. (1986): Süßwasserflora von Mitteleuropa. Bacillariophyceae 1. Teil, Naviculaceae. – In: 

Ettl, H., Gerloff, J., Heynig, H. & Mollenhauer, D. (eds): Süßwasserflora von Mitteleuropa. 2 (1). G. Fischer, Stuttgart & 

New York. 

[This volume documents 21 taxa, published as new in Krammer & Lange-Bertalot (1985) in Biblioth. Diatomol. 9. Some 

of these figures are referred to in Biblioth. Diatomol. 9 and thus be treated as part of the protologue of new diatom 

names.] 

Krammer, K. & Lange-Bertalot, H. (1987): Morphology and taxonomy of Surirella ovalis and related taxa. – Diatom Res. 2: 

77-95. 

Krammer, K. & Lange-Bertalot, H. (1988): Süßwasserflora von Mitteleuropa. Bacillariophyceae 1. Teil, Naviculaceae. – In: 

Ettl, H., Gerloff, J., Heynig, H. & Mollenhauer, D. (eds): Süßwasserflora von Mitteleuropa. 2 (2). G. Fischer, Stuttgart & 

New York. 

[This volume documents 9 taxa, published as new in Lange-Bertalot & Krammer (1985) in Biblioth. Diatomol. 15. Some 

of these figures are referred to in Biblioth. Diatomol. 15 and thus be treated as part of the protologue of new diatom 

names.] 

Krammer, K. & Lange-Bertalot, H. (1991a): Bacillariophyceae. 3. Teil: Centrales, Fragilariaceae, Eunotiaceae. – In: Ettl, H., 

Gerloff, J., Heynig, H. & Mollenhauer, D. (eds): Süßwasserflora von Mitteleuropa. 2 (3). G. Fischer, Stuttgart & Jena. 

[Comment: Some names first used in this volume were validated in Lange-Bertalot (1993).] 

Krammer, K. & Lange-Bertalot, H. (1991b): Bacillariophyceae. 4. Teil: Achnanthaceae. Kritische Ergänzungen zu Navicula 

(Lineolatae) und Gomphonema. Gesamtliteraturverzeichnis. Teil 1-4. – In: Ettl, H., Gerloff, J., Heynig, H. & 

Mollenhauer, D. (eds): Süßwasserflora von Mitteleuropa. 2 (4). G. Fischer, Stuttgart & Jena. 

[Comment: Some names first used in this volume were validated in Lange-Bertalot (1993).] 

Krammer, K. & Lange-Bertalot, H. (1993): Cymbella rossii spec. nov., a new "nonconformist" in the genus Cymbella. – 

Nova Hedwigia Beih. 106: 93-99. 

Krammer, K. & Lange-Bertalot, H. (2000): Süßwasserflora von Mitteleuropa. Bacillariophyceae 3. Teil, Centrales, 

Fragilariaceae, Eunotiaceae. Ergänzte und korrigierte 2. Auflage. – In: Ettl, H., Gerloff, J., Heynig, H. & Mollenhauer, D. 

(eds): Süßwasserflora von Mitteleuropa. 2 (3). Spektrum, Heidelberg & Berlin. 

Krammer, K., Lange-Bertalot, H. & Schiller, W. (1997): Miosira rhoenana n. gen. n. sp. (Bacillariophyceae) aus miozänen 

limnischen Sedimenten der Rhön im Vergleich mit anderen zentrischen Diatomeen. – Paläontol. Z. 71: 5-18. 

Kusber, W.-H. & Jahn, R. 2002: Annotated list of diatom names by Horst Lange-Bertalot and co-workers. - Version 1.0. 

[http://www.algaterra.org/Names_Version1_0.pdf] (pdf-file, 549 KB, Version 1.0; 17. May 2002). 

Kusber, W.-H. & Metzeltin, D. (2001): Checklist of new diatom combinations, replaced names and validations published by 

Horst Lange-Bertalot until the year 2000 and additional validations. – In: Jahn, R., Kociolek, J.P., Witkowski, A. & 

Compère, P. (eds): Lange-Bertalot-Festschrift: 585-633. Gantner, Ruggell. 



95 

Lange-Bertalot, H. (1977): Eine Revision zur Taxonomie der Nitzschiae lanceolatae Grunow. Die "klassischen" bis 1930 

beschriebenen Süßwasserarten Europas. – Nova Hedwigia 28: 253-307. 

Lange-Bertalot, H. (1978): Zur Systematik, Taxonomie und Ökologie des abwasserspezifisch wichtigen Formenkreises um 

"Nitzschia thermalis". – Nova Hedwigia 30: 635-652. 

Lange-Bertalot, H. (1979): Simonsenia a new genus with morphology intermediate between Nitzschia and Surirella. – 

Bacillaria 2: 127-136. 

Lange-Bertalot, H. (1979): Toleranzgrenzen und Populationsdynamik benthischer Diatomeen bei unterschiedlich starker 

Abwasserbelastung. – Arch. Hydrobiol. Suppl. 56,2 (Algol. Stud. 23): 184-219. 

Lange-Bertalot, H. (1980): New species, combinations and synonyms in the genus Nitzschia. – Bacillaria 3: 41-77. 

Lange-Bertalot, H. (1980): Zur taxonomischen Revision einiger ökologisch wichtiger "Naviculae lineolatae" Cleve. Die 

Formenkreise um Navicula lanceolata, N. viridula, N. cari – Cryptog. Algol. 1: 29-50. 

Lange-Bertalot, H. (1993): 85 Neue Taxa und über 100 weitere neu definierte Taxa ergänzend zur Süßwasserflora von 

Mitteleuropa Vol. 2 / 1-4. – Biblioth. Diatomol. 27: 1-428. 

Lange-Bertalot, H. (1995): Gomphosphenia paradoxa nov. spec. et nov. gen. und Vorschlag zur Lösung taxonomischer 

Probleme infolge eines veränderten Gattungskonzepts von Gomphonema (Bacillariophyceae). – Nova Hedwigia 60: 241- 

252. 

Lange-Bertalot, H. (1996): Kobayasia bicuneus gen. et spec. nov. – Iconogr. Diatomol. 4: 277-287. 

Lange-Bertalot, H. (1997): Frankophila, Mayamaea and Fistulifera: drei neue Gattungen der Klasse Bacillariophyceae. – Arch. 

Protistenk. 148: 65-76. 

Lange-Bertalot, H. (1997): Zur Revision der Gattung Achnanthes sensu lato (Bacillariophyceae): Achnantheiopsis, eine neue 

Gattung mit dem Typus generis A. lanceolata. – Arch. Protistenk. 148: 199-208. 

Lange-Bertalot, H. (1999a): Kobayasiella Lange-Bertalot nov. nom. ein neuer Gattungsname für Kobayasia Lange-Bertalot 

1996. – Iconogr. Diatomol. 6: 266-269. 

[Comment: This original article was published with different page numbers in: Iconogr. Diatomol. 6 (2 nd edition; “2 nd 

corrected printing”): 272-275.] 

Lange-Bertalot, H. (1999b): Kobayasiella Lange-Bertalot nov. nom. ein neuer Gattungsname für Kobayasia Lange-Bertalot 

1996. – Iconogr. Diatomol. 6 (2 nd corrected printing): 272-275. 

[Comment: This article is a reprint of the original article with different page numbers. Diatom names are again marked as 

new.] 

Lange-Bertalot, H. (2000): Transfer to the generic rank of Decussata Patrick as a subgenus of Navicula Bory sensu lato – 

Iconogr. Diatomol. 9: 670-673. 

Lange-Bertalot, H. (2001): Navicula sensu stricto – 10 genera separated from Navicula sensu lato - Frustulia. – Diatoms of 

Europe 2: 1-526. Gantner Verlag, Ruggell. 

Lange-Bertalot, H. & Bonik, K. (1976): Massenentwicklung bisher seltener und unbekannter Diatomeen als Indikator starker 

Abwasserbelastung in Flüssen. – Arch. Hydrobiol. Suppl. 49 (Algol. Stud. 16): 303-332. 

Lange-Bertalot, H. & Compère, P. (2001): Fragilaria subgen. Ulnaria, comb. nov., the correct name of the subgenus 

including Synedra ulna, when treated in Fragilaria – Diatom Res. 16: 103-104. 

Lange-Bertalot, H. & Jahn, R. (2000): On the identity of Navicula (Frustulia) rhomboides and Frustulia saxonica 

(Bacillariophyceae). – Syst. Geogr. Pl. 70: 255-261. 

Lange-Bertalot, H. & Krammer, K. (1987): Bacillariaceae, Epithemiaceae, Surirellaceae. Neue und wenig bekannte Taxa, 

neue Kombinationen und Synonyme sowie Bemerkungen und Ergänzungen zu den Naviculaceae. – Biblioth. Diatomol. 

15: 1-289. 

Lange-Bertalot, H. & Krammer, K. (1989): Achnanthes Eine Monographie der Gattung. – Biblioth. Diatomol. 18: 1-393. 

Lange-Bertalot, H. & Krammer, K. (1993): Observations on Simonsenia and some small species of Denticula and Nitzschia. 

– Nova Hedwigia Beih. 106: 121-131. 

Lange-Bertalot, H. & Le Cohu, R. (1985): Raphe like vestiges in the pennate diatom suborder Araphidinae – Ann. Limnol. 

21: 213-220. 



96 

Lange-Bertalot, H. & Metzeltin, D. (1996): Oligotrophie-Indikatoren. 800 Taxa repräsentativ für drei diverse Seen-Typen, 

kalkreich - oligodystroph - schwach gepuffertes Weichwasser. – Iconogr. Diatomol. 2: 1-390. 

Lange-Bertalot, H. & Metzeltin, D. (1996): Ultrastructure of Surirella desikacharyi sp. nov. and Campylodiscus indianorum 

sp. nov. (Bacillariophyta) in comparison with some other taxa of the “robustoid” type of construction. – Nova Hedwigia 

Beih. 112: 321-328. 

Lange-Bertalot, H. & Moser, G. (1994): Brachysira, Monographie der Gattung. – Biblioth. Diatomol. 29. 1-212. 

Lange-Bertalot, H. & Reichardt, E. (2000): Diploneis ovalis sensu stricto und Diploneis krammeri nov. spec. – Revision des 

aktuellen Konzepts von Diploneois ovalis (Hilse) Cleve. – Iconogr. Diatomol. 9: 650-669. 

Lange-Bertalot, H. & S.I. Genkal (1999a): Diatoms from Siberia I. Islands in the Artic Ocean (Yugorsky-Shar-Strait). – 


[Comment: Since the 1 st edition (ISBN 3-904144-09-X) and the 2 nd edition (“2 nd corrected printing”, ISBN 3-904144-12- 

X) differ in their protologues in some cases, both versions are cited.] 

Lange-Bertalot, H. & S.I. Genkal (1999b): Diatoms from Siberia I. Islands in the Artic Ocean (Yugorsky-Shar-Strait). – 

Iconogr. Diatomol. 6 (2 nd corrected printing): 1-271. 

[Comment: Since the 1 st edition (ISBN 3-904144-09-X) and the 2 nd edition (“2 nd corrected printing”, ISBN 3-904144-12- 

X) differ in their protologues in some cases, both versions are cited.] 

Lange-Bertalot, H. & Simonsen, R. (1978): A taxonomic revision of the Nitzschiae Lanceolatae Grunow. 2. European and 

related extra-European fresh water and brackish water taxa – Bacillaria 1: 11-111. 

Lange-Bertalot, H. & Werum, M. (2001): Diadesmis fukushimae sp. nov. and some other new or rarely observed taxa ot the 

subgenus Paradiadesmis Lange-Bertalot & Le Cohu – Diatom 17: 3-19. 

Lange-Bertalot, H., Külbs, K., Lauser, T., Nörpel-Schempp, M. & Willmann, M. (1996): Dokumentation und Revision der 

von Georg Krasske beschriebenen Diatomeen-Taxa. – Iconogr. Diatomol. 3: 1-358. 

Lange-Bertalot, H., Metzeltin, D. & Witkowski, A. (1996): Hippodonta gen. nov. - Umschreibung und Begründung einer 

neuen Gattung der Naviculaceae. – Iconogr. Diatomol. 4: 247-275. 

Lange-Bertalot, H., Rumrich, U. & Hofmann, G. (1991): Zur Revision der Gattung Diatoma Bory (Subgenus Diatoma, 

Bacillariophyceae). Identifikation ökologisch wichtiger, aber taxonomisch problematischer Arten. – Acta Biol. Benrodis 

3: 115-130. 

Maidana, N.I., Díaz Villanueva, V. & Krammer, K. (2002): Taxonomy and valv structure of Cymbella neuquina Frenguelli . 

(Bacillariophyceae), including a new combination, C. neuquina var. fastigata (Krasske) nov. comb. – Nova Hedwigia 74: 

339-348. 

Metzeltin, D. & Kusber, W.-H. (2001): Annotated list of new diatom taxa described by Horst Lange-Bertalot until the year 

2000. – In: Jahn, R., J.P. Kociolek, A. Witkowski & P. Compère (eds): Lange-Bertalot-Festschrift: 557-584. Gantner, 

Ruggell. 

Metzeltin, D. & Lange-Bertalot, H. (1995): Kritische Wertung der Taxa in Didymosphenia (Bacillariophyceae). – Nova 

Hedwigia 60: 381-405. 

Metzeltin, D. & Lange-Bertalot, H. (1998): Tropische Diatomeen in Südamerika I. 700 überwiegend wenig bekannte oder 

neue Taxa repräsentativ als Elemente der neotropischen Flora. – Iconogr. Diatomol. 5: 1-695. 

Metzeltin, D. & Witkowski, A. (1996): Diatomeen der Bären-Insel. Süßwasser- und marine Arten. – Iconogr. Diatomol. 4: 3- 

232. 

Metzeltin, D. (1996): Geissleria lange-bertalotii sp. nov.. – Iconogr. Diatomol. 4: 234-245. 

Moser, G. (1999): Die Diatomeenflora von Neukaledonien. Systematik, Geobotanik, Ökologie: Ein Fazit. – Biblioth. 

Diatomol. 43: 1-205. 

Moser, G., Lange-Bertalot, H. & Metzeltin, D. (1998): Insel der Endemiten. Geobotanisches Phänomen Neukaledonien. – 

Biblioth. Diatomol. 38: 1-464. 

Moser, G., Steindorf, A. & Lange-Bertalot, H. (1995): Neukaledonien. Diatomeenflora einer Tropeninsel. –Biblioth. 

Diatomol. 32: 1-340. 

Reichardt, E. (1981): Die Diatomeenflora quelliger und sumpfiger Standorte in der Umgebung von Treuchtlingen. – Nova 

Hedwigia 34: 487-519. 



97 

Reichardt, E. (1982): Die Diatomeenflora der "Steinernen Rinnen" in Mittelfranken. – Ber. Bayer. Bot. Ges. 53: 97-112. 

Reichardt, E. (1984): Die Diatomeen der Altmühl. – Biblioth. Diatomol. 6: 1-169. 

Reichardt, E. (1985): Diatomeen an feuchten Felsen des Südlichen Frankenjuras. – Ber. Bayer. Bot. Ges. 56: 167-187. 

Reichardt, E. (1988): Achnanthes praecipua n. sp., a new freshwater diatom from Mexico. – In: Round, F. E. (ed.): 

Proceedings of the Ninth International Diatom Symposium, Bristol 1986: 391-395. 

Reichardt, E. (1988): Neue Diatomeen aus Bayerischen und Nordtiroler Alpenseen. – Diatom Res. 3. 237-244. 

Reichardt, E. (1988): Süßwasser-Diatomeen von Papua-Neuguinea. – Nova Hedwigia 47: 81-127. 

Reichardt, E. (1994): Two new species of the genus Surirella from Lake Turkana, East Africa. – Mem. Calif. Acad. Sci. 17: 

129-134. 

Reichardt, E. (1994): Zur Diatomeenflora (Bacillariophyceae) tuffabscheidender Quellen und Bäche im Südlichen 

Frankenjura. – Ber. Bayer. Bot. Ges. 64. 119-133. 

Reichardt, E. (1995): Die Diatomeen (Bacillariophyceae) in Ehrenbergs Material von Cayenne, Guyana Gallica (1843). – 


[Comment: The first edition (ISBN 1-878762-69-9 USA, 3-87429-371-8 Germany) was followed by a second edition 

with corrected plate size and different page numbers (ISBN 3-87429-381-5 Germany, 3-878762-76-1 (“3-978762-76-1”) 

USA).] 

Reichardt, E. (1995): Die Diatomeen (Bacillariophyceae) in Ehrenbergs Material von Cayenne, Guyana Gallica (1843). – 

Iconogr. Diatomol. 1 (second edition): 1-99. 

[Comment: The second edition is not marked as such, but differs from the first edition by format, page numbers and 

ISBN (ISBN 3-87429-381-5 Germany, 3-878762-76-1 (“3-978762-76-1”) USA) for the second edition.] 

Reichardt, E. (1997): Taxonomische Revision des Artenkomplexes um Gomphonema pumilum (Bacillariophyceae). – Nova 

Hedwigia 65: 99-129. 

Reichardt, E. (1997): Morphologie und Taxonomie wenig bekannter Arten aus der Sammelgattung Navicula (excl. Navicula 

sensu stricto). – Diatom Res. 12: 299-320. 

Reichardt, E. (1999): Zur Revision der Gattung Gomphonema. Die Arten um G. affine/insigne, G. angustatum/micropus, G. 

acuminatum sowie gomphonemoide Diatomeen aus dem Oberoligozän in Böhmen. – Iconogr. Diatomol. 8: 1-203. 

Reichardt, E. (2001): Revision of the species around Gomphonema truncatum and G. capitatum. – In: Jahn, R., Kociolek, 

J.P., Witkowski, A. & Compère, P. (eds): Lange-Bertalot-Festschrift: 187-224. Gantner, Ruggell. 

Reichardt, E. & Lange-Bertalot, H. (1990): Fragilaria germainii, eine zweite Fragilaria-Art mit diatomoiden 

Rippenstrukturen. – In: Ricard, M. (ed.): Ouvrage dédié à la Mémoire du Prof. H. Germain (1902-1989): 203-209. 

Reichardt, E. & Lange-Bertalot, H. (1991): Taxonomische Revision des Artenkomplexes um Gomphonema angustum, G. 

dichotomum, G. intricatum, G. vibrio und ähnliche Taxa (Bacillariophyceae). – Nova Hedwigia 53: 519-544. 

Rumrich, U., Lange-Bertalot, H. & Rumrich, M. (2000): Diatomeen der Anden. Von Venezuela bis Patagonien/Feuerland. – 


Sabater, S., Tomàs, X., Cambra, J. & Lange-Bertalot, H. (1990): Diatom flora of the Cape of Creus peninsula, Catalonia, 

N.E. of Spain. – Nova Hedwigia 51: 165-195. 

Schiller, W. & Lange-Bertalot, H. (1997): Eolimna martinii n. gen., n. sp. (Bacillariophyceae) aus dem Unter-Oligozän von 

Sieblos/ Rhön im Vergleich mit ähnlichen rezenten Taxa. – Paläontol. Z. 71: (3-4): 163-172. 

Schmidt, R., Mäusbacher, R. & Müller, J. (1990): Holocene diatom flora and stratigraphy from sediment cores of two 

Antarctic lakes (King George Island). – J. Paleolimnol. 3: 55-74. 

Silva-Benavides, D. (1996): The epilithic diatom flora of a pristine and a polluted river in Costa Rica, Central America – 

Diatom Res. 11: 105-142. 

Tsarenko, P.M., Lange-Bertalot, H., Stupina, V.V. & Wasser, S.P. (2000): Bacillariophyta. – In: Nevo, E. & Wasser, S.P. 

(eds): Biodiversity of cyanoprocaryotes and algae of continental Israel: 195-335, pl. 14-26. Gantner, Ruggell. 

Witkowski, A. (1991): Fallacia cassubiae sp. nov.,a new brackish-water diatom from the Puck Bay (Southern Baltic Sea), 

Poland. – Diatom Res. 6: 401-409. 



98 

Witkowski, A. (1993): Cocconeis hauniensis sp. nov.,a new epipsammic diatom from Puck Bay (Southern Baltic Sea), 

Poland. – Nord. J. Bot. 13: 467-471. 

Witkowski, A. (1993): Fallacia florinae (Moeller) comb. nov., a marine, epipsammic diatom. – Diatom Res. 8: 215-219. 

Witkowski, A. (1993): Fragilaria gedanensis sp. nov. (Bacillariophyceae), a new epipsammic diatom species from the Baltic 

Sea. – Nova Hedwigia 56: 497-503. 

Witkowski, A. (1994): Recent and fossil diatom flora of the Gulf of Gdańsk, Southern Baltic Sea. – Biblioth. Diatomol. 28: 

1-313 

Witkowski, A. & Lange-Bertalot, H. (1993): Established and new diatom taxa related to Fragilaria schulzii Brockmann. – 

Limnologica 23: 59-70. 

Witkowski, A., Lange-Bertalot, H. & Metzeltin, D. (2000): Diatom flora of marine coasts I. – Iconogr. Diatomol. 7: 1-925. 

Witkowski, A., Lange-Bertalot, H. & Stachura, K. (1998): New and confused species in the genus Navicula 

(Bacillariophyceae) and the consequences of restrictive generic circumscription. – Cryptog. Algol. 19: 83-108. 

Witkowski, A., Lange-Bertalot, H. & Witak, M. (1995): Diatom taxa of unusual frustule structure belonging to the genus 

Fragilaria. – Fragm. Florist. Geobot. 40: 729-741. 

Witkowski, A., Metzeltin, D., Lange-Bertalot, H. & Bafana, G. (1997): Fogedia gen. nov. (Bacillariophyceae), a new 

naviculoid genus from the marine littoral. – Nova Hedwigia 65: 79-98. 

Wydrzycka, Ú. & Lange-Bertalot, H. (2001): Las diatomeas (Bacillariophyceae) acidófilas del río Agrio y sitios vinculados 

con su cuenca, volcán Poás, Costa Rica. – Brenesia 55-56: 1-68.

Annotated list of diatom names by Horst Lange-Bertalot and co ...

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