Hepatics and Anthocerotes of the Mediterranean, an ... - NTNU

354 R.M. Ros, V. Mazimpaka et al.Fig. 1. Map of the study area indicating the abbreviation used the countries and islands for whicha separate list is included in the checklist (cf. also Table 1).The lengthy duration of the project is explained by its complexity and thefact that the members of the OPTIMA Bryophyte Commission had otherconcurrent activities and responsibilities. However the experience gained duringthe project will serve as a sound basis for the checklist of mosses still to beproduced.Criteria for the list of accepted namesTaxonomy and NomenclatureThe list includes published names and records to the end of May 2007.The checklist by Grolle & Long (2000) was the basic reference work for acceptednames in Europe and Macaronesia. We have made some exceptions to this list inthe light of more recently published taxonomic papers, and we have added thenames of taxa from African and Asian countries that were outside the geographicalscope of Grolle & Long’s list. We have also included some names that are notcited in Grolle & Long’s list in view of their frequent use in Mediterranean areasand the lack of consensus in recent literature on their taxonomic value. For example,Grolle & Long (2000) cited very few infra-specific names. We took the viewthat the varieties and subspecies of some polymorphic taxa that have been recognizedin relatively recent studies merited inclusion (e.g., those of Bazzania trilobata(L.) Gray, Blepharostoma trichophyllum (L.) Dumort., Jubula hutchinsiae(Hook.) Dumort., Marchantia polymorpha L. and several species of Riccia ). Incases where infraspecific taxa have not been distinguished in the literature for a

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 355Table 1. Countries and island groups treated in the checklist, with the names of the responsibleauthors. Country abbreviations are based on the two-letters code defined by ISO 3166-1 forcountry names. For the islands that are not independent countries a two-letter code as used inSöderström et al. (2002) has been used except for Madeira for which MD has been used.For the purpose of the checklist, Andorra has not been considered. Sinai has been included inEgypt. Israel includes some adjacent areas (Gaza, the West Bank, the former Cis-Jordan, andparts of both the Golan Heights and Mt. Hermon). The list of Malta includes the Maltese islands:Malta, Gozo and Comino. The Spanish North-African territories have been included in Morocco;Kosovo in Serbia; Gibraltar in Spain. Turkey includes the European and Asian parts, although nosegregation between them has been done.Countries and Islands Abbreviations AuthorsAlbania AL M. Sabovljevi †Algeria DZ R.M. Ros & M.J. CanoBaleares BL M. Brugués & R.M. CrosBosnia-Herzegovina BA M. Sabovljevi †Bulgaria BG A. GanevaCanary Islands CN A. Losada-Lima, G.M. Dirkse, S. Rodríguez-Nuñez& J.M. González-ManceboCorsica CO L. SöderströmCrete CT T.L. BlockeelCroatia HR M. Sabovljevi †Cyprus CY T. L. BlockeelEgypt EG W. El Saadawi, H. Shabbara, U. Abou-Salama & M.S. RefaiFrance FR L. SöderströmGreece GR T.L. BlockeelIsrael IL I. HerrnstadtItaly IT M. AleffiJordan JO H. KürschnerLebanon LB H. Kürschner & A. Erda ∑Libya LY R.M. Ros & M.J. CanoMacedonia MK M. Sabovljevi †Madeira MD M. Sim-SimMalta MT E. LanfrancoMontenegro ME M. Sabovljevi †Morocco MA R.M. Ros & M.J. CanoPortugal PT C. SérgioSardinia SA M. AleffiSerbia RS M. Sabovljevi †Sicilia SC M.G. DiaSlovenia SI M. Sabovljevi †Spain ES M. Brugués & R.M. CrosSyria SY W. El Saadawi, H. Shabbara, U. Abou-Salama, M.S. Refai& K. KhalilTunisia TN R.M. Ros & M.J. CanoTurkey TR H. Kürschner & A. Erda ∑

356 R.M. Ros, V. Mazimpaka et al.given territory, then the relevant occurrences have been indicated in the checklistat species level. For the most part, however, we have been restrictive in our inclusionof infraspecific taxa. Thus, if the majority of recent authors have not recognizeda taxon, then we have also ignored it here. We have not recognized thecategory of “form”. We have compared the contents of our list as they relate toEuropean territories with the data and subsequent amendments published bySöderström et al. (2002; 2007) (see http://www.bio.ntnu.no/ECCB/ for a completechecklist). Taxa erroneously reported from the territories covered by Söderströmet al. (2007) and noted as such by them are not repeated here. Taxa whose presencein a given area is doubtful have been excluded at the discretion of the authorresponsible for that area. We have ignored names previously rejected in publishedpapers, but have included an explanatory annotation for those rejected here.The names of the accepted taxa and their occurrence in individualterritories are presented in tabular format. Taxa are arranged alphabeticallywithin taxonomic unit, and territories are listed in alphabetic sequence of theirrespective abbreviations. The reported presence of a taxon in a territory is markedby one of the following symbols indicating the nature of the record. The absenceof a symbol means that the taxon has not been reported in the territory or hasbeen rejected in previous publications. Annotations concerning taxonomic criteriaand floristic data are indicated by a number following the name of the taxon, orin the relevant cell of the grid. Report based on collections published before 1962. Report based on at least one collection published during or after 1962.( ) Doubtful report based on collections published before 1962.( ) Doubtful report based on at least one collection published during or after 1962.+ Report without precise collection data or locality.Criteria for the list of synonymsOur objective was to compile all the names published for Mediterraneanliverworts, and we have therefore included all available synonyms. The authors ofthe national and island lists were asked to include all the names published fortheir designated territories. All these names were checked using standard workson liverworts and other important recent publications, including general andregional floras (Müller, 1954-1957; Schuster, 1966, 1969, 1980, 1988, 1992a, 1992b;Paton, 1999; Schumacker & Vá≈ a, 2000, 2 005; Damsholt, 2002), generic monographs(Grolle, 1972; Jovet-Ast, 1986; Bis chler-Causse, 1993; Sim-Sim, 1999; Engel &Merrill, 2004; Long, 2006, etc.) and earlier lists (e.g., Grolle, 1983, Grolle & Long,2000; Söderström et al ., 2002). Our intention was to include all the names used inthe literature, even if they had been misapplied or had not been validly published.However it became clear that the authorities given for names cited in floristic publicationswere sometimes erroneous, and we have therefore ignored such errors.The names and authorities of taxa were first checked against IndexHepaticarum (Bonner, 1962a, 1962b, 1963a, 1963b, 1965, 1966; Bischler, 1977;Geissler & Bischler, 1985, 1987, 1989, 1990) and Index of Hepatics (Crosby &Engel, 2006), and the nomenclature was then further verified using the publicationscited above. Author abbreviations follow Brummitt & Powell (1992), andannotations are included for problematic names. Invalid combinations have beenincluded, especially basionyms and their subsequent combinations, with the aim offacilitating their interpretation.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 357For ease of consultation, all the synonyms and disputed names used inthe Mediterranean countries have been arranged alphabetically in a singleconsolidated list, with a cross-reference where appropriate to the annotation inwhich the taxonomic problem is discussed. This list also includes names rejectedfor the whole of our area.Summarizing the results of three centuries work in a critical list is acomplex and difficult task, especially considering that advances in taxonomy andnomenclature have run in parallel with floristic exploration. While we haveattempted to check all names against the available literature, we do not assumethat the result is perfect. We would welcome any further observations regardingomissions, errors and inconsistencies, since they will serve to improve futureeditions of the checklist.

358 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 359List of accepted namesAnthocerotaeAL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRAnthoceros L.agrestis Paton [2] ● ● + ● ● + ● ● ● +caucasicus Steph. ● ● ● ● ●punctatus L. [2] ● ● ● ● ● ● ● ● [1] ● ● ● ● ●Notothylas Sull. ex A. Grayorbicularis (Schwein.) A. GrayPhaeoceros Prosk.carolinianus (Michx.) Prosk. ● [3] + ● ● ( ● ) ● + ● ●laevis (L.) Prosk. [4] [5] ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ( ● ) ● ● ● ● ●Phymatoceros Stotler, W. T. Doyle& Crand.-Stotl. [6]bulbiculosus (Brot.) Stotler,W. T. Doyle & Crand.-Stotl. [5]● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ●HepaticaeAcanthocoleus R. M. Schust.aberrans (Lindenb.& Gottsche) Kruijt●●Acrobolbus Neeswilsonii Nees ( ) ●Adelanthus Mitt.decipiens (Hook.) Mitt. ● ● ●Anastrepta (Lindb.) Schiffn.orcadensis (Hook.) Schiffn. ● ● ●Anastrophyllum (Spruce) Steph.assimile (Mitt.) Steph.●hellerianum (Nees ex Lindenb.)R. M. Schust.● ● ● ● michauxii (F. Weber) H. Buch ● ● ● ●minutum (Schreb.) R. M. Schust. ● ● ● ● ● ● saxicola (Schrad.) R. M. Schust.●Aneura Dumort.maxima (Schiffn.) Steph. ● ●pinguis (L.) Dumort. ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ●Anthelia (Dumort.) Dumort.julacea (L.) Dumort. [7] ● juratzkana (Limpr.) Trevis. [7] ● ● ● ● ●Aphanolejeunea A. Evansazorica (V. Allorge & Ast)Bernecker & Pócs●

360 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 361AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRmadeirensis (Schiffn.) Grolle●microscopica (Taylor) A. Evans ● ● ● ●sintenisii Steph. ● ●Apomarsupella R. M. Schust.revoluta (Nees) R. M. Schust. ● ●Apometzgeria Kuwah.pubescens (Schrank) Kuwah. ● ● ● ● ● ● ● ● ● Arnellia Lindb.fennica (Gottsche) Lindb. ● Asterella P. Beauv. [8]africana (Mont.) A. Evans ● ● ● ● ● ●gracilis (F. Weber) Underw. [9] ● ● ● ● ● ●lindenbergiana (Corda ex Nees)Arnell● ● ( ) saccata (Wahlenb.) A. Evans [10] ( ) ● ●Athalamia Falconerhyalina (Sommerf.) S. Hatt. [9] ● ● ● ● ● ● ● ● ( ) ● [1] ● ●spathysii (Lindenb.) S. Hatt. ● ● ● ● ● ( ● ) ● ● ● ● ● ● + ● ● ●Barbilophozia Loeskeatlantica (Kaal.) Müll. Frib. ● ●attenuata (Mart.) Loeske + ● ● ● barbata (Schmidel ex Schreb.)Loeske● ● ● ● ● ● ● ● [1] ● ● ● ●binsteadii (Kaal.) Loeske●floerkei (F. Weber & D. Mohr)Loeske● ● ● ● ● ● ● hatcheri (A. Evans) Loeske ● ● ● ● ● ● ● ● ● ● ● ● ● ●kunzeana (Huebener) Müll. Frib. ● ● ● ●lycopodioides (Wallr.) Loeske ● ● ● ● ● ● ● ● ● quadriloba (Lindb.) Loeske ● ● rubescens (R. M. Schust. &Damsh.) Kartt. & L. Söderstr.+Bazzania Grayflaccida (Dumort.) Grolle ● ● ● ( ● ) ● [1] ● ●tricrenata (Wahlenb.) Lindb. ● ● ● + ● ●trilobata (L.) Grayvar. depauperata (Müll. Frib.)Grolle● +var. trilobata ● ● ● ● ● ● ● ●Blasia L.pusilla L. ● ● ● ● ● ( ) ( ● ) ●Blepharostoma (Dumort.) Dumort.trichophyllum (L.) Dumort. [1]subsp. brevirete (Bryhn & Kaal.)R. M. Schust.+ ●subsp. trichophyllum ● ● ● ● ● ● ● ● ● ● ● ●

362 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 363AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRCalypogeia Raddiarguta Nees & Mont. ● ● ● ● ● ● ● ● ● ● ● ●azorica Bischl.●azurea Stotler & Crotz [11] [12] ● ● ( ● ) ● ● ● ( ) ( ● ) ( ) ● ● ( ) ● ( ) ●fissa (L.) Raddi ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●integristipula Steph. ● ● ● ( ● ) ●muelleriana (Schiffn.) Müll. Frib.[11]● [1] ● ● ● ● ● ● ● ( ● ) ●neesiana (C. Massal. & Carestia)Müll. Frib.● ● ● ● ● sphagnicola (Arnell & J. Perss.)Warnst. & Loeske● ● ● ● ● ● ● ●suecica (Arnell & J. Perss.) Müll.Frib.● ● ● ● ● ● ● ● Cephalozia (Dumort.) Dumort.affinis Lindb. ex Steph.●ambigua C. Massal. ● ● ● ●bicuspidata (L.) Dumort. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●catenulata (Huebener) Lindb. ● ( ● ) ● ● ● ● ● ● connivens (Dicks.) Lindb. ● [1] ● ● ● ● ( ) ● ●crassifolia (Lindenb. & Gottsche)Fulford●●lacinulata J. B. Jack ex Spruce + ● leucantha Spruce ● ● ●loitlesbergeri Schiffn. + ● ● ●lunulifolia (Dumort.) Dumort. ● ● ● ● ● ● ● ● ● ● ● macrostachya Kaal. [13] ● ●pleniceps (Austin) Lindb. ● ● ● ● ●Cephaloziella (Spruce) Schiffn.baumgartneri Schiffn. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●calyculata (Durieu & Mont.) Müll.Frib. [14]● ● + ● ● ● ● ● ● [1] ● ● ● ● ●dentata (Raddi) Steph. ● ● ● ● ● ● ●divaricata (Sm.) Schiffn. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●elachista (J. B. Jack ex Gottsche &Rabenh.) Schiffn.( ) ● ● elegans (Heeg) Schiffn. [15] ● ● granatensis (J. B. Jack ex Steph.)Fulford●grimsulana (J. B. Jack ex Gottsche& Rabenh.) Lacout.● ● ●hampeana (Nees) Schiffn. [14] + ● ● ● ● ● ● ● ●integerrima (Lindb.) Warnst. ● ● ●massalongi (Spruce) Müll. Frib. + ● ● ● ● ● ●phyllacantha (C. Massal. &Carestia) Müll. Frib.( ● ) ●rubella (Nees) Warnst. [15] ● ● ● ● ● ● ● ● ( ● ) ● ● +spinigera (Lindb.) Warnst. ● ●stellulifera (Taylor ex Spruce)Schiffn.● ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●turneri (Hook.) Müll. Frib. ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● +varians (Gottsche) Steph.

364 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 365AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRChiloscyphus Cordapallescens (Ehrh. ex Hoffm.)Dumort.[16] ● ● ● ● ● ● ● ● ● ● ● ● ●polyanthos (L.) Corda ● ● ● ● ● ● ● ● ● ● ● ● ●Cladopodiella H. Buchfluitans (Nees) H. Buch ● ● ● ● ●francisci (Hook.) Jörg. ● ● ●Cololejeunea (Spruce) Schiffn.calcarea (Lib.) Schiffn. ● ● ● ● ● ● ● ● ● ● ● ●minutissima (Sm.) Schiffn. ● ● ● ● ● ● [1] ● ● ● ● ● ●rossettiana (C. Massal.) Schiffn. ● ● ● ● ● ● ● ● ● ● ● ● ● ● +schaeferi Grolle ● ●Colura (Dumort.) Dumort.calyptrifolia (Hook.) Dumort. ● ● ●Conocephalum Hillconicum (L.) Dumort. ● ● ● ● ● ● ● ● ● ● ● ● ● ( ● ) ● ● ● ● ● ● ● ● ● ● ●salebrosum Szweyk., Buczkowska& Odrzykoski [17]●●Corsinia Raddicoriandrina (Spreng.) Lindb. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●Cryptothallus Malmb.mirabilis Malmb. ● ●Cyathodium Kunzefoetidissimum Schiffn.●Diplophyllum (Dumort.) Dumort.albicans (L.) Dumort. ● ● ● [1] ● ● ● ● ● ● ● ● ●obtusatum (R. M. Schust.) R. M.Schust.obtusifolium (Hook.) Dumort. ● ● ● ● ● ( ● ) ● taxifolium (Wahlenb.) Dumort. ● ● ● ● ● ● Douinia (C. E. O. Jensen) H. Buchovata (Dicks.) H. Buch ● ● ● ●Drepanolejeunea (Spruce) Schiffn.hamatifolia (Hook.) Schiffn. ● ● ● ● ●Dumortiera Neeshirsuta (Sw.) Nees ● [1] ● ● ( ) ● ● ●Eremonotus Lindb. & Kaal. exPearsonmyriocarpus (Carrington) Pearson ● ● ● ●Exormotheca Mitt.pustulosa Mitt. ● ● ● ● ● ●welwitschii Steph. ● ●

366 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 367AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRFossombronia Raddi [18] [19]angulosa (Dicks.) Raddi ● ● ● ● ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ●caespitiformis De Not. ex Rabenh.[20] [21]subsp. caespitiformis ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●subsp. multispira (Schiffn.) J. R.Bray & D. C. Cargill● ● ● ● ● ● ● ● ● [1] ● ● ● ● ●crispa Nees [22] ( ● )echinata Macvicar [20] ( ● ) ● ● ● + ● ● ● ● ● ● ● ● ● ●foveolata Lindb. ● ● ● ● +incurva Lindb.●leucoxantha Lehm.●mittenii Tind. ● ● ●pusilla (L.) Nees [21] [23] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●wondraczekii (Corda) Lindb. [20][23]● ● ● + (● ) ● ● ● ● ● ● ●Frullania Raddiazorica Sim-Sim et al. [24] ● ● dilatata (L.) Dumort. [25] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●ericoides (Nees) Mont. ● ●fragilifolia (Taylor) Gottsche et al. ● ● ● ● ● ● ● ● ● ● inflata Gottsche ● + ● jackii Gottsche [26] ● ● ● ●microphylla (Gottsche) Pearson ● ● ● ● ●oakesiana Austin ● ●parvistipula Steph. ● ●polysticta Lindenb. ● ●riparia Hampe ex Lehm. ● ● [24] ● ● ● sergiae Sim-Sim et al.●tamarisci (L.) Dumort. [25] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●teneriffae (F. Weber) Nees ● ● ● ● ●Geocalyx Neesgraveolens (Schrad.) Nees ● ● ( ) +Gongylanthus Neesericetorum (Raddi) Nees ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●Gymnocolea (Dumort.) Dumort.inflata (Huds.) Dumort. + ● ● ● ● ( ) ● ● ● ( ) ●Gymnomitrion Cordaapiculatum (Schiffn.) Müll. Frib. + concinnatum (Lightf.) Corda ● ● ● ● ●corallioides Nees ● ● ● ● crenulatum Gottsche ex Carrington ● ● ●obtusum Lindb. ● ● ●Haplomitrium Neeshookeri (Sm.) Nees ● ● ●Harpalejeunea (Spruce) Schiffn.molleri (Steph.) Grolle ● ● ● ● ( ) ● ● ●

368 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 369AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRHarpanthus Neesflotovianus (Nees) Nees ● ● scutatus (F. Weber & D. Mohr)Spruce + ● ● ( ) ●Heteroscyphus Schiffn.denticulatus (Mitt.) Schiffn. ● ●Hygrobiella Sprucelaxifolia (Hook.) Spruce ● ●Jamesoniella (Spruce) F. Leesautumnalis (DC.) Steph. ● ● ● ● ( ) ● ●undulifolia (Nees) Müll. Frib.●Jubula Dumort.hutchinsiae (Hook.) Dumort. [27]subsp. hutchinsiae ● [27] ● ● ● ●[27]subsp. javanica (Steph.) Verd. ● [27]Jungermannia L. [28] [29]atrovirens Dumort. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● borealis Damsh. & Vá≈ a ● ●caespiticia Lindenb.●callithrix Lindenb. & Gottsche●caucasica Vá≈ a●confertissima Nees ● ● ● exsertifolia Steph.subsp. cordifolia (Dumort.) Vá≈ a ● ● ● ● ● gracillima Sm. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●handelii (Schiffn.) Amakawa ● hyalina Lyell [30] ● ● ● ● ● ● ● ● ● ● ( ● ) ● ● ● ● ( ) ●leiantha Grolle ● ● ● ● ● ● ● [1] ● ● ● ●lignicola (Schiffn.) Grolleobovata Nees ● ● [1] ● ● ● ● ● ● paroica (Schiffn.) Grolle ● ●polaris Lindb. ● ● ● ● ● ●pumila With. ● ● [1] ● ● ● ● ( ) ● sphaerocarpa Hook. ● ● ● ● ● ● ● ( ) ● ●subelliptica (Lindb. ex Kaal.) Levier ( ) ● ● ●subulata A. Evans ● ●subtilissima (Schiffn.) GrolleKurzia G. Martenspauciflora (Dicks.) Grolle ● ● ● ● sylvatica (A. Evans) Grolle ● ● ●trichoclados (Müll. Frib.) Grolle ● ● ( )Leiocolea (Müll. Frib.) H. Buchbadensis (Gottsche) Jörg. ● ● ● ● ● ● +bantriensis (Hook.) Jörg. [31] ● ● ● ● ● ● ● ● ( ● ) ●collaris (Nees) Schljakov [31] ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ● ●gillmanii (Austin) A. Evans ● ●

370 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 371AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRheterocolpos (Thed. ex Hartm.) H. ● ● ● ● ● ● ● Buchturbinata (Raddi) H. Buch ● ● ● ● ● ● ● ● ● ● ● ● ● [32] ● ● ● ● ● ● ●Lejeunea Lib.canariensis (Steph.) Steph. ● ●cavifolia (Ehrh.) Lindb. [33] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●eckloniana Lindenb. ● ● ● ●flava (Sw.) Nees ● ●[34]hibernica Bischl. et al. ex Grolle●lamacerina (Steph.) Schiffn. ● ● ● ● ● ● ● ● ●mandonii (Steph.) Müll. Frib. ● ● ● ●patens Lindb. [33] ( ● ) ● ● ● ● Lepidozia (Dumort.) Dumort. [35]cupressina (Sw.) Lindenb. ● ● ● ●reptans (L.) Dumort. ● ● ● ● ● ● ● ● ● ● ● Leptoscyphus Mitt.cuneifolius (Hook.) Mitt.●Lophocolea (Dumort.) Dumort.bidentata (L.) Dumort. ● ● ● ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ●fragrans (Moris & De Not.)Gottsche et al.● ● ● ● ● ● ●heterophylla (Schrad.) Dumort. ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ● ● ● ● ●minor Nees ● ● ● ● [1] ● ● ● ● ● ● ● ●Lophozia (Dumort.) Dumort. [36]ascendens (Warnst.) R. M. Schust. ● ● ● ● ●bicrenata (Schmidel ex Hoffm.)Dumort.● ● ● ● ● ● ● ● ● ●capitata (Hook.) Macoun ( ) ●decolorans (Limpr.) Steph. ● ●elongata Steph.excisa (Dicks.) Dumort. + ● ● ● ● ● ● ● ● ( ) ● ●grandiretis (Lindb. ex Kaal.)Schiffn.( ● ) incisa (Schrad.) Dumort. ● ● ● ● ● ● ●laxa (Lindb.) Grolle●longidens (Lindb.) Macoun ● ● ● ● ● longiflora (Nees) Schiffn. ● ● ● ● ( ) ● ( ) ● ● ●obtusa (Lindb.) A. Evans ● ● ● ● ● ● ●opacifolia Culm. ex Meyl. ● ● ●perssonii H. Buch & S. W. Arnell ● ●polaris (R. M. Schust.) R. M.Schust. & Damsh.silvicola H. Buch [37] + ● ● ● +sudetica (Nees ex Huebener) Grolle ● ● ● ● ● ● ● ● ventricosa (Dicks.) Dumort. [38] ● ● ● ● ● ● ● ● ( ) ● ●wenzelii (Nees) Steph. ● ● ● ● ●Lunularia Adans.cruciata (L.) Lindb. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●

374 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 375AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRMylia Grayanomala (Hook.) Gray ● ● ● ● taylorii (Hook.) Gray ● ● ● ●Nardia Graybreidleri (Limpr.) Lindb. ● ● ●compressa (Hook.) Gray ● ● ● ● ( ) ● [1] ● ● geoscyphus (De Not.) Lindb. [41] ● ( ) ● ● ● ● ● ● ● insecta Lindb. [41]●scalaris Gray ● ● ● ● ● ● ● ● ● ● ●Nowellia Mitt.curvifolia (Dicks.) Mitt. ● ● ● ● ● ● ● ● ● Odontoschisma (Dumort.)Dumort. [12]denudatum (Mart.) Dumort. ● ● ● ● elongatum (Lindb.) A. Evans ● ● ●macounii (Austin) Underw.prostratum (Sw.) Trevis.●sphagni (Dicks.) Dumort. + ● ● ● ● Oxymitra Bisch. ex Lindenb.incrassata (Brot.) Sérgio & Sim-Sim ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ●Pallavicinia Graylyellii (Hook.) Carruth. ● ● ● ● ● ●Pedinophyllum (Lindb.) Lindb.interruptum (Nees) Kaal. ● ● ● ● ● ● ● ● ● ● ● ● ●Pellia Raddiendiviifolia (Dicks.) Dumort. ● ● ● ● ● ● ● ● [42] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●epiphylla (L.) Corda [43] ● ● ● ( ● ) ● ● ● ● ● [1] ● ● ● ● ● ● ● ●neesiana (Gottsche) Limpr. ● ● ● ● ● ( ) ● ● ●Peltolepis Lindb.quadrata (Saut.) Müll. Frib. ● Petalophyllum Nees & Gottsche exLehm.ralfsii (Wilson) Nees & Gottsche ● ● ● ● ● ● ● ● ● ● ● ●Plagiochasma Lehm. & Lindenb.appendiculatum Lehm. & Lindenb .microcephalum (Steph.) Steph.var. tunesicum Bischl.rupestre (J. R. Forst. & G. Forst.)Steph.Plagiochila (Dumort.) Dumort.[44]asplenioides (L. emend. Taylor)Dumort. [45]●●● ● ● ● ● + ● ● ● ● ● ● + ( ) ● ● ● ● ● ● ● ●● ( ● ) ● ● ( ) ● ● ● ( ● ) ● ● ( ● ) (● ) ● ● ● ● ●

376 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 377AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRbifaria (Sw.) Lindenb. ● ● ● ● ● ● [1]britannica Paton●exigua (Taylor) Taylor ● ● ● ● ●heterophylla Lindenb. ex Lehm.[46]+maderensis Gottsche ex Steph. [47] ● ●porelloides (Torrey ex Nees)Lindenb. [45] ● ● ● + ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ●punctata (Taylor) Taylor ● ● ● ● ●retrorsa Gottsche●spinulosa (Dicks.) Dumort. ● ● ● ●stricta Lindenb. ● ●virginica A. Evans ● ●Pleurocladula Grollealbescens (Hook.) Grolle ● ● ● ●Porella L.arboris-vitae (With.) Grolle ● ● ●( ● )[48]● ● ● ● ● ● [1] ● ● ● ● ● ●x baueri (Schiffn.) C. E. O. Jensen ● ● ● ● ● ● ( ) ● ● ● ●canariensis (F. Weber) Underw. ● ● [49] ● ●cordaeana (Huebener) Moore ● ● ● ● ● ● ● ● ● ● ● ● [50] ● ● ● ● ● ● ● ●inaequalis (Gottsche ex Steph.)Perss.●obtusata (Taylor) Trevis. [16] ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ●pinnata L. ● ● ● [50] ● ●platyphylla (L.) Pfeiff. ● ● ● ● ● [48] ● ● ● ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ●Prasanthus Lindb.suecicus (Gottsche) Lindb.●Preissia Cordaquadrata (Scop.) Nees [51] ● ● ● ● ● ● ● ● ● ● ● ● ● ●Ptilidium Neesciliare (L.) Hampe ● ● ● ● ● ● pulcherrimum (Weber) Vain. ● ● ● ● ● ● ● Radula Dumort. [52]aquilegia (Hook. f. & Taylor)Gottsche et al.● ● ●carringtonii J. B. Jack ● ●complanata (L.) Dumort. ● ● ● ● ● ● + ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ●holtii Spruce ● ● ● ●jonesii Bouman et al. ● ●lindenbergiana Gottsche ex C.Hartm.● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●nudicaulis Steph.●visianica C. Massal.wichurae Steph. ● ●Reboulia Raddihemisphaerica (L.) Raddi [51] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●

378 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 379AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRRiccardia Graychamedryfolia (With.) Grolle ● ● ● ● ● ● ● ● ● ( ) ● ● ● ( ) ●incurvata Lindb. ( ) ● ● ● ●latifrons (Lindb.) Lindb. ● [1] ● ● ● ● ● ● ● ● ●multifida (L.) Gray ● ● ● ● ● ● ● ● ● ● ( ● ) ● ● ● ●palmata (Hedw.) Carruth. ● ● ● ● ● ● ● ●Riccia L. [53] [54] [55]atlantica Sérgio & Perold●atromarginata Levier ● ● ● ● ● ● ● ● ● ● ● ● ● ( ) ● + ●beyrichiana Hampe ex Lehm. ● ● ● ● ● ● ● ● ● ●bicarinata Lindb. ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● + ● ●bifurca Hoffm. [56] ● ● ● ● ● ● ● ● [1] ● ● ● ● ●breidleri Jur. ex Steph. ● ●canaliculata Hoffm. ● + ● ● ● + ● ● ● + +cavernosa Hoffm. ● ● ● ● ● ( ) ● ● ● ● ● ● ●ciliata Hoffm. [57] ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ●ciliifera Link ex Lindenb. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + + ●congoana Steph.●crinita Taylor [57] ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ●crozalsii Levier ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● + ● ●crustata Trab. ● ● + ● ● ● ● + ● ● ●crystallina L. emend. Raddi ● ● ● ● ● ● ● ● ● ● ● [1] ● ● ● ● ● ● ● ● ● ●duplex Lorb. ● ● ● ●fluitans L. ● ● ● ● ● ● ● ( ) ● + ● ● ● ●frostii Austin ● ● ● ● ● + ● + [58] ●glauca L. ( ● ) ● ● var. ciliaris Warnst. ● ● ● ● ● ● ● ●var. glauca ● ● ● ● ● ● ● ● ● + ● ( ) ● ● ● ●gougetiana Durieu & Mont. + var. armatissima Levier ex Müll.Frib.● ● ● ● ● ● ● ● ● ●var. gougetiana ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ●huebeneriana Lindenb. ( ) ● ● ● [1] ●lamellosa Raddi ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● + ● ●ligula Steph. ● [1] ● + ● ( ) ●macrocarpa Levier ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●melitensis C. Massal.●michelii Raddi ● ● ● + ● ● ● ● ● ● ● ● [1] ● ● ● ● ● + ● ●nigrella DC. ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ●papillosa Moris ● ● ● ● ● ● ● ● ● ● ● ● ● +perennis Steph. ● ● ● ● ● ● ●rhenana Lorb. ex Müll. Frib. ● ●sommieri Levier ● ● ● ● ●sorocarpa Bisch. ●var. heegii Schiffn. ● ● + ● ● + ● ● ●var. sorocarpa ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● + ● ●subbifurca Warnst. ex Croz. ● ● ● ● ● ● ● ● ● ● ● ● +teneriffae S. W. Arnelltrabutiana Steph. ● ● ● ● + ● ● ● + ● ● ● ● ● ● ● ● ●warnstorfii Limpr. ex Warnst. ● [1]var. subinermis Warnst. ● var. warnstorfii ( ) ● ● ● ● ● ● ( ) ●

380 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 381AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRRicciocarpos Cordanatans (L.) Corda ● ● ● ● ● ● ( ● ) ● ●Riella Mont.affinis M. Howe & Underw. ● [59] ●bialata Trab.cossoniana Trab. ● ● ● ● ● [1] +cyrenaica Maire●helicophylla (Bory & Mont.) Mont. ● ● ● ● ● ● ● +notarisii (Mont.) Mont. ● ● ● ● ● ● ● ● ●numidica Trab. [60] ● parisii Gottsche ( ) ● [1] sersuensis Trab. [60]Saccogyna Dumort.viticulosa (L.) Dumort. ● ● ● ● ● ● ● ( )Sauteria Neesalpina (Nees) Nees ● ● Scapania (Dumort.) Dumort.aequiloba (Schwägr.) Dumort. ● ● ● ● ● ● [1] ● ● ● ● apiculata Spruce ● ● ●aspera Bernet & M. Bernet ● ● ● ● ● ● ● ● ● [1] ● ( ) ● ( ) ● ● brevicaulis Taylor +[61]calcicola (Arnell & J. Perss.)Ingham● ● [62] ● ● ● ● ● ● carinthiaca J. B. Jack ex Lindb.●compacta (A. Roth) Dumort. [63] ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●crassiretis Bryhn ● ●curta (Mart.) Dumort. ● ● ● ● ● ● ● ● ● ● ( ) cuspiduligera (Nees) Müll. Frib. ● ● ( ) ● gracilis Lindb. ● ● [1] ● ● ( ) ● ● ( )gymnostomophila Kaal. ● ● ●helvetica Gottsche ● ( ● ) ● ● irrigua (Nees) Nees ● ● ● ● ● ● ● ●lingulata H. Buch ● ● ●mucronata H. Buch ● ● ● ● ● + ●nemorea (L.) Grolle ● ● ● ● ● ● ● ● ● ● ● ● ● ●obscura (Arnell & C. E. O. Jensen)Schiffn.●paludicola Loeske & Müll. Frib. ● ● ● ●paludosa (Müll. Frib.) Müll. Frib. ● ● ● ( ● ) ● ●praetervisa Meyl. ● ● ●scandica (Arnell & H. Buch)Macvicar● ● ● ● ● ●scapanioides (C. Massal.) Grolle●subalpina (Nees ex Lindenb.)Dumort.● ● ( ) ● ● ● ● ● ● ●uliginosa (Sw. ex Lindenb.)Dumort.( ● ) ● ●umbrosa (Schrad.) Dumort. ● ● ● ● ● ● ● ● ● undulata (L.) Dumort. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●verrucosa Heeg ● ● ● ● ●

382 R.M. Ros et al.Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 383AL BA BG BL CN CO CT CY DZ EG ES FR GR HR IL IT JO LB LY MA MD ME MK MT PT RS SA SC SI SY TN TRSouthbya Sprucenigrella (De Not.) Henriq. ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●tophacea (Spruce) Spruce ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ( ) ● ● ●Sphaerocarpos Boehm.michelii Bellardi ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●stipitatus Bisch. ex Lindenb.●texanus Austin ● ● ● ●( )[64]● ● ● ● ● ( ) ● [1] ●Targionia L.hypophylla L. ● ● ● ● ● ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●lorbeeriana Müll. Frib. ● ● + ● ● ● ● ● ● ● ● ● ● ● ● ● ● ● ●Telaranea Spruce ex Schiffn.azorica ( H. Buch & Perss.) Pócs exSchumacker & Vá≈ a [65]europaea J. J. Engel & G. L. S.Merril [66] [67]●● ● ● ● ●Tetralophozia (R. M. Schust.)Schljakovfiliformis (Steph.) Urmi●setiformis (Ehrh.) Schljakov ( ● ) Trichocolea Dumort.tomentella (Ehrh.) Dumort. ● ● ● ● ( ) ● [68] ● ● Tritomaria Schiffn. ex Loeskeexsecta (Schmidel ex Schrad.)Loeske ● ● ● ● ● ● ( ) ●exsectiformis (Breidl.) Loeske ● ● ● ● ● ● ● ● ●polita (Nees) Jörg. ● ● quinquedentata (Huds.) H. Buch ● ● ● ● ● ● ● ● ● scitula (Taylor) Jörg. + ● ●Tylimanthus Mitt.madeirensis Grolle & Perss.●

384 R.M. Ros et al.List of synonyms and names used in the studied areaSymbols used in the list: Name used in the Mediterranean literature, but not found in reference works and there areuncertainties about the correct synonymy and authorship.☞ The author/s of the name is/are doubtful.? Name used in the Mediterranean literature, but not found in reference works and theauthorship is therefore uncertain, although there is no doubt about the taxon referred(the name was probably created in a floristic publication).# Name used in the Mediterranean literature and found in independent reference works,but the synonymy could not be confirmed.$ Synonym used in the Mediterranean literature, but not formally published (the namewas undoubtedly created in a floristic publication) therefore no authors are indicated.AnthocerotaeAnthoceros beltrani Casares-Gil [4]Anthoceros bulbiculosus Brot. = Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle &Crand.-Stotl.Anthoceros caespiticius De Not. = Anthoceros punctatus L.Anthoceros carolinianus Michx. = Phaeoceros carolinianus (Michx.) Prosk.Anthoceros constans Lindb. = Anthoceros punctatus L.Anthoceros crispulus (Mont.) Douin = Anthoceros punctatus L.Anthoceros crispulus auct. non (Mont.) Douin = Anthoceros agrestis PatonAnthoceros dichotomus Raddi = Phymatoceros bulbiculosus (Brot.) Stotler, W. T. Doyle &Crand.-Stotl.Anthoceros dichotomus f. stricta Casares-Gil = Phymatoceros bulbiculosus (Brot.) Stotler,W. T. Doyle & Crand.-Stotl.Anthoceros dichotomus var. gussonei Zodda = Phaeoceros laevis (L.) Prosk. [5]Anthoceros husnotii Steph. = Anthocerospunctatus L.Anthoceros laevis L. = Phaeoceros laevis (L.) Prosk.Anthoceros mandonii Steph. = Anthoceros caucasicus Steph.Anthoceros multifidus L . = Anthoceros punctatus L. [2]Anthoceros multifidus auct. = Anthoceros agrestis Paton [2]Anthoceros multilobulus Lindb. = Anthoceros punctatus L.Anthoceros polymorphus Raddi = Anthoceros punctatus L.Anthoceros punctatus var. cavernosus auct. non (Nees) Gottsche et al. = Anthoceros agrestisPatonAnthoceros punctatus var. crispatus Mont. = Anthoceros punctatus L.Anthoceros punctatus var. crispatus auct. non Mont. = Anthoceros agrestis PatonAnthoceros punctatus var. crispulus Mont. = Anthoceros punctatus L.Anthoceros punctatus var. crispulus auct. non Mont. = Anthoceros agrestis PatonAnthoceros punctatus var. husnotii (Steph.) Zodda = Anthoceros punctatus L.Anthoceros punctatus f. multifidus (L.) Nees = Anthoceros punctatus L. [2]Anthoceros punctatus f. multifidus auct. non (L.) Nees = Anthoceros agrestis Paton [2]Anthoceros punctatus var. multifidus (L.) Schmidel ex Swartz = Anthoceros punctatus L. [2]Anthoceros punctatus var. multifidus auct. non (L.) Schmidel ex Swartz = Anthocerosagrestis Paton [2]Anthoceros validus Steph. = Phaeoceros carolinianus (Michx.) Prosk.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 385# Anthoceros torreyi Austin = Anthoceros punctatus L.Aspiromitus madeirensis Steph. = Anthoceros caucasicus Steph.Carpobolus orbicularis (Schwein.) Schwein. = Notothylas orbicularis (Schwein.) A. GrayCarpolipum fertile Lehm. = Notothylas orbicularis (Schwein.) A. GrayCarpolipum valvatum (Sull.) Nees = Notothylas orbicularis (Schwein.) A. GrayChamaeceros fertilis (Lehm.) Milde = Notothylas orbicularis (Schwein.) A. GrayNotothylas fertilis (Milde) Lehm. = Notothylas orbicularis (Schwein.) A. GrayNotothylas valvata Sull. = Notothylas orbicularis (Schwein.) A. GrayPhaeoceros bulbiculosus (Brot.) Prosk. = Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle & Crand.-Stotl.Phaeoceros dichotomus (Raddi) Prosk. = Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle & Crand.-Stotl.Phaeoceros laevis subsp. carolinianus (Michx.) Prosk. = Phaeoceros carolinianus (Michx.)Prosk.HepaticaeAchiton quadratum (Scop.) Corda = Preissia quadrata (Scop.) Müll. Frib. [51]Acolea adusta (Nees emend. Limpr.) C. Massal. hom. illeg. non (Gray) Trevis. = Marsupellaadusta (Nees emend. Limpr.) SpruceAcolea alpina ( Gottsche ex Husn.) C. Massal. = Marsupella alpina (Gottsche ex Husn.)BernetAcolea brevissima Dumort. = Marsupella brevissima (Dumort.) GrolleAcolea concinnata (Lightf.) Dumort. = Gymnomitrion concinnatum (Lightf.) CordaAcolea conferta (Limpr.) C. Massal. & Carestia = Marsupella brevissima (Dumort.) GrolleAcolea corallioides (Nees) Dumort. = Gymnomitrion corallioides NeesAcolea revoluta (Nees) Steph. = Apomarsupella revoluta (Nees) R. M. Schust.Acolea varians (Lindb.) Steph. = Marsupella brevissima (Dumort.) GrolleAitonia italica (Sassi) Lindb. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Alicularia breidleri Limpr. = Nardia breidleri (Limpr.) Lindb.Alicularia compressa (Hook.) Nees = Nardia compressa (Hook.) GrayAlicularia geoscypha De Not. = Nardia geoscyphus (De Not.) Lindb.Alicularia geoscypha f. insecta (Lindb.) Müll. Frib. = Nardia insecta Lindb.Alicularia minor (Nees) Limpr. = Nardia geoscyphus (De Not.) Lindb.Alicularia pachyphylla De Not. = Nardia compressa (Hook.) GrayAlicularia rotaeana De Not. = Nardia scalaris GrayAlicularia scalaris (Gray) Corda = Nardia scalaris GrayAlicularia scalaris var. minor ( Nees) Nees = Nardia geoscyphus (De Not.) Lindb.Anastrophyllum minutum var. grande (Gottsche ex Lindb.) R. M. Schust. = Anastrophyllumminutum (Schreb.) R. M. Schust.Anastrophyllum minutum var. weberi (Mart.) Kartt. = Anastrophyllum minutum (Schreb.)R. M. Schust.Anastrophyllum myriocarpum (Carrington) R. M. Schust. ex Vá≈ a = Eremonotusmyriocarpus (Carrington) PearsonAnastrophyllum nardioides (Lindb.) Kaal. = Anastrophyllum assimile (Mitt.) Steph.Anastrophyllum reichardtii (Gottsche in Jur.) Steph. = Anastrophyllum assimile (Mitt.)Steph.Aneura incurvata (Lindb.) Steph. = Riccardia incurvata Lindb.Aneura latifrons Lindb. = Riccardia latifrons (Lindb.) Lindb.Aneura multifida (L.) Dumort. = Riccardia multifida (L.) Gray

386 R.M. Ros et al.Aneura multifida var. major (Dicks. ex Hook.) Nees = Riccardia chamedryfolia (With.)Grolle☞ Aneura multifida var. rivularis Rabenh. = Riccardia chamedryfolia (With.) GrolleAneura palmata (Hedw.) Dumort. = Riccardia palmata (Hedw.) Carruth.Aneura pinguis var. angustior (Hook.) Dumort. = Aneura pinguis (L.) Dumort.Aneura pinnatifida (Sw.) Dumort. hom. illeg. = Riccardia chamedryfolia (With.) GrolleAneura sinuata (Dicks. ex Hook.) Dumort. = Riccardia chamedryfolia (With.) GrolleAnomomarsupella cephalozielloides R. M. Schust. = Eremonotus myriocarpus (Carrington)PearsonAnthelia andina Herzog = Anthelia juratzkana (Limpr.) Trevis.Anthelia julacea subsp. juratzkana (Limpr.) Meyl. = Anthelia juratzkana (Limpr.) Trevis.Anthelia nivalis (Sw. ex F. Weber & D. Mohr) Lindb. [7]Anthelia phyllacantha C. Massal. & Carestia = Cephaloziella phyllacantha (C. Massal. &Carestia) Müll. Frib.Anthelia setiformis (Ehrh.) Dumort. = Tetralophozia setiformis (Ehrh.) SchljakovAnthelia turneri (Hook.) Dumort. = Cephaloziella turneri (Hook.) Müll. Frib.Antoiria vulgaris Raddi nom. illeg. = Porella platyphylla (L.) Pfeiff.Antrocephalus italicus Sassi=Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Aphanolejeunea teotonii V. Allorge & Ast = Aphanolejeunea sintenisii Steph.Aplozia amplexicaulis (Dumort.) Dumort. = Jungermannia sphaerocarpa Hook.Aplozia anomala (Hook.) Warnst. = Mylia anomala (Hook.) GrayAplozia atrovirens (Dumort.) Dumort. = Jungermannia atrovirens Dumort.Aplozia autumnalis (DC.) Heeg = Jamesoniella autumnalis (DC.) Steph.Aplozia confertissima (Nees) Dumort. = Jungermannia confertissima NeesAplozia cordifolia Dumort. = Jungermannia exsertifolia subsp. cordifolia (Dumort.) V á ≈ aAplozia crenulata (Mitt.) Lindb. = Jungermannia gracillima Sm.Aplozia crenulata var. inundata (Schiffn.) Macvicar = Jungermannia gracillima Sm.Aplozia crenulata var. gracillima (Sm.) Heeg = Jungermannia gracillima Sm.Aplozia gracillima (Sm.) Dumort. = Jungermannia gracillima Sm.Aplozia hyalina (Lyell) Dumort. = Jungermannia hyalina LyellAplozia lanceolata (L.) Dumort. = Jungermannia atrovirens Dumort.Aplozia lanceolata auct. non (L.) Dumort. = Jungermannia leiantha GrolleAplozia lurida (Dumort.) Dumort. = Jungermannia sphaerocarpa Hook.Aplozia lurida sensu Breidl. = Jungermannia confertissima NeesAplozia nana (Nees) Breidl. = Jungermannia sphaerocarpa Hook.Aplozia oblongifolia (Müll. Frib.) Jörg. = Jungermannia pumila With.Aplozia obovata (Nees) Loeske = Jungermannia obovata NeesAplozia polaris (Lindb.) Bryhn & Kaal. = Jungermannia polaris Lindb.Aplozia pumila (With.) Dumort. = Jungermannia pumila With.Aplozia riparia (Taylor) Dumort. = Jungermannia atrovirens Dumort.Aplozia riparia var. potamophila Müll. Arg. ex Bernet = Jungermannia atrovirens Dumort.Aplozia riparia var. rivularis Bernet = Jungermannia atrovirens Dumort.Aplozia scalariformis (Nees) Dumort. = Jungermannia confertissima NeesAplozia schiffneri Loitl. = Jungermannia polaris Lindb.Aplozia sphaerocarpa (Hook.) Dumort. = Jungermannia sphaerocarpa Hook.Aplozia sphaerocarpa var. nana (Nees ) Macvicar = Jungermannia sphaerocarpa Hook.Aplozia sphaerocarpoidea (De Not.) Dumort. = Jungermannia atrovirens Dumort.Aplozia tristis (Nees) Dumort. = Jungermannia atrovirens Dumort.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 387Asterella bonjeanii (De Not.) Trevis. = Asterella lindenbergiana (Corda ex Nees) ArnellAsterella elegans (Spreng.) Trevis. REJECTED FROM THE AREA [8]Asterella fragrans (Schleich.) Trevis. = Asterella saccata (Wahlenb.) A. Evans [10]Asterella hemisphaerica (L.) P. Beauv. = Reboulia hemisphaerica (L.) RaddiAsterella ludwigii (Schwägr.) Underw. ex A. Evans nom. inval. = Mannia triandra (Scop.)GrolleAsterella ludwigii auct. non (Schwägr.) Underw. ex A. Evans nom. inval. = Asterella gracilis(F. Weber) Underw.Asterella nana (Lindenb.) C. Massal. = Athalamia hyalina (Sommerf.) S. Hatt. [9]Asterella pilosa (Wahlenb.) Trevis. = Asterella gracilis (F. Weber) Underw.Aytonia rupestris J. R. Forst. & G. Forst. = Plagiochasma rupestre (J. R. Forst. & G. Forst.)Steph.Barbilophozia baueriana (Schiffn.) Loeske = Barbilophozia hatcheri (A. Evans) LoeskeBarbilophozia gracilis (Schleich. ex Steph.) Müll. Frib. = Barbilophozia attenuata (Mart.)LoeskeBarbilophozia hatcheri var. grandiretis H. Buch ex Lammes = Barbilophozia rubescens(R. M. Schust. & Damsh.) Kartt. & L. Söderstr.Barbilophozia lycopodioides var. hatcheri (Evans) Schljakov = Barbilophozia hatcheri(A. Evans) Loeske? Barbilophozia lycopodioides var. parvifolia Schiffn. = Barbilophozia hatcheri (A. Evans)LoeskeBarbilophozia obtusa (Lindb.) H. Buch = Lophozia obtusa (Lindb.) A. EvansBazzania deflexa (Mart.) Trevis. = Bazzania tricrenata (Wahlenb.) Lindb.Bazzania denudata auct. non (Torrey ex Gottsche et al .) Trevis. = Bazzania flaccida(Dumort.) GrolleBazzania implexa (Nees) Müll. Frib. = Bazzania flaccida (Dumort.) GrolleBazzania triangularis (Schleich.) Lindb. nom. illeg. = Bazzania tricrenata (Wahlenb.)Lindb.Bazzania triangularis var. implexa (Nees) Zahlbr. = Bazzania flaccida (Dumort.) Grolle$ Bazzania tricrenata var. implexa = Bazzania flaccida (Dumort.) GrolleBellincinia laevigata (Schrad.) Kuntze = Porella arboris-vitae (With.) GrolleBellincinia montana Raddi = Porella arboris-vitae (With.) GrolleBellincinia platyphylla (L.) Kuntze = Porella platyphylla (L.) Pfeiff.Bellincinia porella (Dicks.) Kuntze = Porella pinnata L.Bellincinia thuja (Hook.) Kuntze = Porella arboris-vitae (With.) GrolleBlasia pusilla var. gemmifera Nees = Blasia pusilla L.Blepharostoma arachnoideum var. brevirete (Bryhn & Kaal.) Frye & L. Clark = Blepharostomatrichophyllum subsp. brevirete (Bryhn & Kaal.) R. M. Schust.Blepharostoma trichophyllum var. brevirete Bryhn & Kaal. = Blepharostoma trichophyllumsubsp. brevirete (Bryhn & Kaal.) R. M. Schust.Blepharozia ciliaris (L.) Dumort. = Ptilidium ciliare (L.) Hampe? Blepharozia ciliaris var. wallrothiana Nees = Ptilidium pulcherrimum (Weber) Vain.Blepharozia pulcherrima (Weber) Lindb. = Ptilidium pulcherrimum (Weber) Vain.Blyttia lyellii (Hook.) Endl. ex Gottsche et al. = Pallavicinia lyellii (Hook.) Carruth.Blyttia lyellii var. hibernica (Hook.) Nees = Moerckia hibernica (Hook.) GottscheBlyttia moerckii Nees = Moerckia blyttii (Moerch ex Hornem.) Brockm.Brissocarpus riccioides Bisch. = Corsinia coriandrina (Spreng.) Lindb.Calycularia blyttii (Moerch ex Hornem.) Steph. = Moerckia blyttii (Moerch ex Hornem.)Brockm.Calycularia hibernica (Hook.) Steph. = Moerckia hibernica (Hook.) Gottsche

388 R.M. Ros et al.Calypogeia adscendens (Nees) Warnst. = Calypogeia muelleriana (Schiffn.) Müll. Frib.Calypogeia ericetorum Raddi = Gongylanthus ericetorum (Raddi) NeesCalypogeia flagellifera Raddi = Gongylanthus ericetorum (Raddi) NeesCalypogeia meylanii H. Buch = Calypogeia integristipula Steph.Calypogeia muelleriana f. sphagnicola (Arnell & J. Perss.) Schljakov = Calypogeiasphagnicola (Arnell & J. Perss.) Warnst. & LoeskeCalypogeia neesiana var. laxa Meyl. ex Müll. Frib. = Calypogeia muelleriana (Schiffn.)Müll. Frib.Calypogeia neesiana var. meylanii (H. Buch) R. M. Schust. = Calypogeia integristipulaSteph.Calypogeia paludosa Warnst. = Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. &LoeskeCalypogeia tenuis (Austin) A. Evans. = Calypogeia sphagnicola (Arnell & J. Perss.)Warnst. & LoeskeCalypogeia trichomanis (L.) Corda nom. rejic. = Calypogeia fissa (L.) RaddiCalypogeia trichomanis auct. non (L.) Corda = Calypogeia azurea Stotler & Crotz [11]# Calypogeia trichomanis var. communis Nees = Calypogeia fissa (L.) RaddiCalypogeia trichomanis var. erecta Müll. Frib. = Calypogeia muelleriana (Schiffn.) Müll.Frib.Calypogeia trichomanis var. fissa (L.) De Not. = Calypogeia fissa (L.) Raddi# Calypogeia trichomanis f. papillosa Warnst. = Calypogeia azurea Stotler & CrotzCalypogeia trichomanis var. sphagnicola (Arnell & J. Perss.) Meyl. = Calypogeiasphagnicola (Arnell & J. Perss.) Warnst. & LoeskeCalypogeia trichomanis var . sprengelii (Mart.) C. Massal. = Calypogeia fissa (L.) RaddiCalypogeia trichomanis var. suecica (Arnell & J. Perss.) Casares-Gil = Calypogeia suecica(Arnell & J. Perss.) Müll. Frib.Candollea carinata Raddi = Scapania compacta (A. Roth) Dumort.Cephalozia albescens (Hook.) Dumort. = Pleurocladula albescens (Hook.) Grolle# Cephalozia asperifolia f. belsensis Douin = Cephaloziella divaricata (Sm.) Schiffn.Cephalozia baueri (Mart.) C. Massal. = Nowellia curvifolia (Dicks.) Mitt.Cephalozia bicuspidata subsp. ambigua (C. Massal.) R. M. Schust. = Cephalozia ambiguaC. Massal.Cephalozia bicuspidata subsp. lammersiana (Huebener) R. M. Schust. = Cephaloziabicuspidata (L.) Dumort.Cephalozia bicuspidata var. alpicola (C. Massal.) C. Massal. & Carestia = Cephaloziapleniceps (Austin) Lindb.Cephalozia bicuspidata var. lammersiana (Huebener) Breidl. = Cephalozia bicuspidata (L.)Dumort.Cephalozia bicuspidata var. setulosa Spruce = Cephalozia bicuspidata (L.) Dumort.Cephalozia bifida Lindb. = Cephaloziella rubella (Nees) Warnst.Cephalozia borealis Lindb. = Cladopodiella francisci (Hook.) Jörg.Cephalozia bryhnii Kaal. = Cephaloziella rubella (Nees) Warnst.Cephalozia byssacea (A. Roth) Dumort. nom. rejic. = Cephaloziella hampeana (Nees)Schiffn.Cephalozia byssacea auct. non (A. Roth) Dumort. = Cephaloziella divaricata (Sm.) Schiffn.Cephalozia columbae Camus = Cephaloziella massalongi (Spruce) Müll. Frib.Cephalozia compacta Warnst. = Cephalozia connivens (Dicks.) Lindb.Cephalozia curvifolia (Dicks.) Dumort. = Nowellia curvifolia (Dicks.) Mitt.Cephalozia dentata (Raddi) Lindb. = Cephaloziella dentata (Raddi) Steph.Cephalozia divaricata (Sm.) Dumort. = Cephaloziella divaricata (Sm.) Schiffn.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 389Cephalozia divaricata Heeg nom. inval. non (Sm.) Dumort. = Cephaloziella rubella (Nees)Warnst.Cephalozia divaricata var. scabra M. Howe = Cephaloziella divaricata (Sm.) Schiffn.Cephalozia divaricata var. stellulifera Taylor ex Spruce = Cephaloziella stellulifera (Taylorex Spruce) Schiffn.Cephalozia divaricata var. starkei auct. non (Funck ex Nees) Spruce nom. illeg. =Cephaloziella divaricata (Sm.) Schiffn.Cephalozia elegans Heeg = Cephaloziella elegans (Heeg) Schiffn.Cephalozia fluitans (Nees) Spruce = Cladopodiella fluitans (Nees) H. BuchCephalozia francisci (Hook.) Dumort. = Cladopodiella francisci (Hook.) Jörg.Cephalozia granatensis J.B. Jack ex Steph. = Cephaloziella granatensis (J.B. Jack ex Steph.)FulfordCephalozia helleri (Nees ex Lindenb.) Lindb. nom. illeg. = Anastrophyllum hellerianum(Nees ex Lindenb.) R. M. Schust.Cephalozia hibernica Spruce ex Pearson = Cephalozia crassifolia (Lindenb. & Gottsche)FulfordCephalozia hypogyna Steph. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn.Cephalozia integerrima Lindb. = Cephaloziella integerrima (Lindb.) Warnst.Cephalozia lammersiana (Huebener) Carrington = Cephalozia bicuspidata (L.) Dumort.Cephalozia laxifolia (Hook.) Lindb. = Hygrobiella laxifolia (Hook.) Spruce☞ Cephalozia limprichtii (Warnst.) Steph. = Cephaloziella stellulifera (Taylor ex Spruce)Schiffn.Cephalozia lucens (A. Evans) Steph. = Metahygrobiella lucens (A. Evans) H. A. MillerREJECTED FROM THE AREA# Cephalozia lunulaefolia var. gasilieni Corb. = Cephalozia affinis Lindb. ex Steph.Cephalozia massalongi Spruce = Cephaloziella massalongi (Spruce) Müll. Frib.Cephalozia media Lindb. = Cephalozia lunulifolia (Dumort.) Dumort.# Cephalozia media var. gasilieni Corb. = Cephaloziaaffinis Lindb. ex Steph.Cephalozia multiflora (Huds.) Lindb. = Cephalozia connivens (Dicks.) Lindb.Cephalozia multiflora Spruce nom. illeg. = Cephalozia lunulifolia (Dumort.) Dumort.Cephalozia myriocarpa (Carrington) Lindb. = Eremonotus myriocarpus (Carrington)PearsonCephalozia nematodes sensu auct. eur. non Gottsche ex Austin = Teleranea europea J.J.Engel & G.L.S. Merrill [66]Cephalozia notarisiana C.Massal. = Hygrobiella laxifolia (Hook.) Spruce☞ Cephalozia nudis Lindb. = Cephalozia lunulifolia (Dumort.) Dumort.Cephalozia obtusiloba Lindb. = Cladopodiella fluitans (Nees) H. BuchCephalozia pallida (Spruce) Pearson = Cephalozia lunulifolia (Dumort.) Dumort.Cephalozia papillosa Douin = Cephaloziella divaricata (Sm.) Schiffn.Cephalozia patula Steph. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn.Cephalozia phyllacantha (C. Massal. & Carestia) C. Massal. & Carestia = Cephaloziellaphyllacantha (C. Massal. & Carestia) Müll. Frib.Cephalozia pleniceps var. macrantha (Kaal. & Nicholson) Müll. Frib. = Cephaloziapleniceps (Austin) Lindb.Cephalozia raddiana (C.Massal.) C. Massal. = Cephaloziella rubella (Nees) Warnst.Cephalozia reclusa (Taylor) Dumort. = Cephaloziacatenulata (Huebener) Lindb.Cephalozia serriflora Lindb. = Cephalozia catenulata (Huebener) Lindb.Cephalozia sphagni (Dicks.) Spruce = Odontoschisma sphagni (Dicks.) Dumort.Cephalozia starkei auct. non (Funck ex Nees) Dumort. ex Cogn. nom. illeg. = Cephalozielladivaricata (Sm.) Schiffn.

390 R.M. Ros et al.Cephalozia symbolica (Gottsche) Breidl. = Cephalozia lunulifolia (Dumort.) Dumort.Cephalozia symbolica var. sphagnorum C. Massal. = Cephalozia pleniceps (Austin) Lindb.Cephalozia turneri (Hook.) Lindb. = Cephaloziella turneri (Hook.) Müll. Frib.Cephaloziella algeriensis (Douin) Trab. = Cephaloziella baumgartneri Schiffn.Cephaloziella alpina Douin = Cephaloziella varians (Gottsche) Steph.Cephaloziella arctica Bryhn & Douin = Cephaloziella varians (Gottsche) Steph.Cephaloziella baumgartneri var. algeriensis Douin = Cephaloziella baumgartneri Schiffn.Cephaloziella bifidoides Douin = Cephaloziella rubella (Nees) Warnst.Cephaloziella byssacea (A. Roth) Warnst. nom. rejic. = Cephaloziella hampeana (Nees)Schiffn.Cephaloziella byssacea auct. non (A. Roth) Warnst. = Cephaloziella divaricata (Sm.)Schiffn.☞ Cephaloziella byssacea var. asperifolia (Taylor) R. M. Schust. = Cephaloziella divaricata(Sm.) Schiffn.# Cephaloziella carnutensis Douin = Cephaloziella divaricata (Sm.) Schiffn.Cephaloziella columbae (Camus) Müll. Frib. = Cephaloziella massalongi (Spruce) Müll.Frib.Cephaloziella compacta (Jörg.) Müll. Frib. = Cephaloziella massalongi (Spruce) Müll. Frib.Cephaloziella curnowii Douin = Cephaloziella hampeana (Nees) Schiffn.Cephaloziella divaricata var. asperifolia (Taylor) Damsh. = Cephaloziella divaricata (Sm.)Schiffn.Cephaloziella divaricata var. polystratosa (R. M. Schust. & Damsh.) Potemkin =Cephaloziella divaricata (Sm.) Schiffn.Cephaloziella divaricata var. rupestris (C. E. O. Jensen) Damsh. nom. inval. = Cephalozielladivaricata (Sm.) Schiffn.Cephaloziella divaricata var. scabra (M. Howe) S.W. Arnell = Cephaloziella divaricata(Sm.) Schiffn.Cephaloziella erosa Limpr. ex Warnst. = Cephaloziella hampeana (Nees) Schiffn.? Cephaloziella francisci (Hook.) Dumort. = Cladopodiella francisci (Hook.) Jörg.Cephaloziella gallica (Douin) Douin = Cephaloziella hampeana (Nees) Schiffn. [14]Cephaloziella gracillima (Douin) Douin = Cephaloziella stellulifera (Taylor ex Spruce)Schiffn.Cephaloziella hampeana var. subtilis (Velen.) Macvicar = Cephaloziella hampeana (Nees)Schiffn.Cephaloziella latzeliana Schiffn. = Cephaloziella divaricata (Sm.) Schiffn.☞ Cephaloziella limprichtii Warnst. = Cephaloziella stellulifera (Taylor ex Spruce) Schiffn.Cephaloziella limprichtii var. stellulifera (Taylor ex Carrington & Pearson) Müll. Frib. =Cephaloziella divaricata (Sm.) Schiffn.Cephaloziella massalongi var. algarvica Douin = Cephaloziella massalongi (Spruce) Müll.Frib.Cephaloziella massalongi var. compacta (Jörg.) Müll. Frib. = Cephaloziella massalongi(Spruce) Müll. Frib.Cephaloziella myriantha (Lindb.) Schiffn. = Cephaloziella rubella (Nees) Warnst. [15]Cephaloziella nigrimonasteriensis Douin = Cephaloziella divaricata (Sm.) Schiffn.Cephaloziella papillosa (Douin) Schiffn. = Cephaloziella divaricata (Sm.) Schiffn.Cephaloziella patula (Steph.) Schiffn. = Cephaloziella stellulifera (Taylor ex Spruce)Schiffn.Cephaloziella piriflora Douin = Cephaloziella integerrima (Lindb.) Warnst.Cephaloziella pulchella (C. E. O. Jensen) Douin = Cephaloziella rubella (Nees) Warnst.Cephaloziella raddiana (C. Massal.) Schiffn. = Cephaloziella rubella (Nees) Warnst.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 391Cephaloziella rubella var. elegans (Heeg) R. M. Schust. nom. illeg. = Cephaloziella elegans(Heeg) Schiffn.Cephaloziella rubella var . pulchella (C. E. O. Jensen) R. M. Schust. = Cephaloziella rubella(Nees) Warnst.Cephaloziella rubella var. sullivantii (Austin) Müll. Frib. = Cephaloziella rubella (Nees)Warnst.Cephaloziella starkei auct. non (Funck ex Nees) Schiffn. nom. illeg. = Cephalozielladivaricata (Sm.) Schiffn.# Cephaloziella starkei var. examphigastriata Douin ex Müll. Frib. = Cephalozielladivaricata (Sm.) Schiffn.☞ Cephaloziella starkei var. papillosa (Douin) Casares-Gil = Cephaloziella divaricata(Sm.) Schiffn.Cephaloziella starkei var. scabra (M. Howe) Frye & L. Clark = Cephaloziella divaricata(Sm.) Schiffn.# Cephaloziella starkei var. serriflora Douin = Cephaloziella divaricata (Sm.) Schiffn.☞ Cephaloziella stellulifera var. gracillima Douin = Cephaloziella stellulifera (Taylor exSpruce) Schiffn.# Cephaloziella stellulifera var. hericensis Douin = Cephaloziella stellulifera (Taylor exSpruce) Schiffn.Cephaloziella stellulifera var. gracillima Douin = Cephaloziella stellulifera (Taylor exSpruce) Schiffn.Cephaloziella stellulifera var. limprichtii (Warnst.) Macvicar = Cephaloziella stellulifera(Taylor ex Spruce) Schiffn.Cephaloziella striatula (C. E. O. Jensen) Douin = Cephaloziella spinigera (Lindb.) Warnst.Cephaloziella subdentata Warnst. = Cephaloziella spinigera (Lindb.) Warnst.Cephaloziella vaucherioides Cout. = Cephaloziella baumgartneri Schiffn.Cephaloziella verrucosa (C.E.O. Jensen) Bryhn & Kaal. hom. illeg. = Cephaloziella varians(Gottsche) Steph.Cesia adusta (Nees emend. Limpr.) Lindb. = Marsupella adusta (Nees emend. Limpr.)SpruceCesia concinnata (Lightf.) Lindb. = Gymnomitrion concinnatum (Lightf.) Corda Cesia conferta (Limpr.) Pearson = Marsupella brevissima (Dumort.) GrolleCesia corallioides (Nees) Carruth. = Gymnomitrion corallioides NeesCesia revoluta (Nees) Lindb. = Apomarsupella revoluta (Nees) R. M. Schust.Cesia varians (Lindb.) Lindb. = Marsupella brevissima (Dumort.) GrolleCesius concinnatus (Lightf.) Gray nom. rejic. = Gymnomitrion concinnatum (Lightf.) CordaChaetopsis trichophylla (L.) Mitt. = Blepharostoma trichophyllum (L.) Dumort.Chandonanthus setiformis (Ehrh.) Lindb. = Tetralophozia setiformis (Ehrh.) SchljakovCheilolejeunea boaventurae Steph. = Lejeunea eckloniana Lindenb.Chiloscyphus canariensis Bryhn = Lophocolea fragrans (Moris & De Not.) Gottsche et al .Chiloscyphus denticulatus Mitt. = Heteroscyphus denticulatus (Mitt.) Schiffn.Chiloscyphus fragilis (A. Roth) Schiffn. = Chiloscyphus pallescens (Ehrh. ex Hoffm.)Dumort.Chiloscyphus lophocoleoides Nees = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort.Chiloscyphus minor (Nees) J. J. Engel & R. M. Schust. = Lophocolea minor NeesChiloscyphus pallescens var. fontana Müll. Frib. = Chiloscyphus pallescens (Ehrh. exHoffm.) Dumort.Chiloscyphus pallescens var. fragilis (A. Roth) Müll. Frib. = Chiloscyphus pallescens (Ehrh.ex Hoffm.) Dumort.

392 R.M. Ros et al.Chiloscyphus polyanthos var. erectus Boulay = Chiloscyphus pallescens (Ehrh. ex Hoffm.)Dumort.# Chiloscyphus polyanthos var. gracilescens Boulay = Chiloscyphus polyanthos (L.) CordaChiloscyphus polyanthos var. pallescens (Ehrh. ex Hoffm.) C. Hartm. = Chiloscyphus pallescens(Ehrh. ex Hoffm.) Dumort.Chiloscyphus polyanthos var. rivularis (Schrad.) Nees = Chiloscyphus polyanthos (L.)CordaChiloscyphus rivularis (Schrad.) Hazsl. = Chiloscyphus polyanthos (L.) CordaChomiocarpon commutatus (Lindenb.) Lindb. = Preissia quadrata (Scop.) Müll. Frib.Chomiocarpon quadratus (Scop.) Lindb. = Preissia quadrata (Scop.) Nees [51]Cincinnulus argutus (Nees & Mont.) Dumort. = Calypogeia arguta Nees & Mont.Cincinnulus suecicus (Arnell & J. Perss.) Müll. Frib. = Calypogeia suecica (Arnell &J. Perss.) Müll. Frib.Cincinnulus trichomanis (L.) Dumort. = Calypogeia fissa (L.) RaddiCincinulus trichomanis auct. non (L.) Dumort. = Calypogeia azurea Stotler & Crotz [11]Cincinnulus trichomanis var. fissus (L.) Boulay = Calypogeia fissa (L.) RaddiCladopus francisci (Hook.) Meyl. = Cladopodiella francisci (Hook.) Jörg.Clevea crassa Trab. nom. inval. = Athalamia hyalina (Sommerf.) S. Hatt.Clevea hyalina (Sommerf.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt.Clevea rousselliana (Mont.) Leitg. ex Steph. = Athalamia spathysii (Lindenb.) S. Hatt.Clevea spathysii (Lindenb.) Müll. Frib. = Athalamia spathysii (Lindenb.) S. Hatt.Clevea suecica (Lindb.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt.Clevea trabutiana Steph. = Athalamia hyalina (Sommerf.) S. Hatt.Codonia pusilla (L.) Dumort.nom. inval. = Fossombronia pusilla (L.) NeesCodonia ralfsii (Wilson) Dumort. = Petalophyllum ralfsii (Wilson) Nees & GottscheCodonia wondraczekii (Corda) Dumort. = Fossombronia wondraczekii (Corda) Lindb.Coleochila anomala (Hook.) Dumort. = Mylia anomala (Hook.) GrayColeochila stillicidiorum (Raddi) Dumort. = Southbya tophacea (Spruce) SpruceCololejeunea azorica V. Allorge & Ast = Aphanolejeunea azorica (V. Allorge & Ast)Bernecker & PócsCololejeunea inconspicua (Raddi) Schiffn. = Cololejeunea minutissima (Sm.) Schiffn.Cololejeunea madeirensis Schiffn. = Aphanolejeunea madeirensis (Schiffn.) GrolleCololejeunea microscopica (Taylor) Schiffn. = Aphanolejeunea microscopica (Taylor) A.EvansCololejeunea teotonii (V. Allorge & Ast) Grolle = Aphanolejeunea sintenisii Steph.☞ Conocephalum hemisphaericum Dumort. = Preissia quadrata (Scop.) NeesConocephalum trioicum F. H. Wigg. nom. superfl. = Conocephalum conicum (L.) Dumort.Conocephalus officinalis Trevis. = Conocephalum conicum (L.) Dumort.Conocephalus vulgaris Bisch. = Conocephalum conicum (L.) Dumort.Corbierella algeriensis Douin & Trab. = Exormotheca welwitschii Steph.? Corbierella welwitschii (Steph.) Douin & Trab. ex Paris = Exormotheca welwitschiiSteph.Cordaea blyttii (Moerch ex Hornem.) Corda = Moerckia blyttii (Moerch ex Hornem.)Brockm.Cordaea flotoviana Nees = Moerckia hibernica (Hook.) GottscheCorsinia coriandrina var . latifrons Trab. nom. inval. = Corsinia coriandrina (Spreng.)Lindb.Corsinia marchantioides Raddi = Corsinia coriandrina (Spreng.) Lindb.Corsinia reticulata (J.F. Gmel.) Dumort. nom. illeg. = Corsinia coriandrina (Spreng.) Lindb.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 393Crossocalyx hellerianus (Nees ex Lindenb.) Meyl. = Anastrophyllum hellerianum (Nees exLindenb.) R. M. Schust.Crossogyna autumnalis (DC.) Schljakov = Jamesoniella autumnalis (DC.) Steph.Cyathophora commutata (Lindenb.) Trevis. = Preissia quadrata (Scop.) Müll. Frib.☞ Dichiton calyculatum (Durieu & Mont.) Trevis. = Cephaloziella calyculata (Durieu &Mont.) Müll. Frib.Dichiton gallicum Douin = Cephaloziella hampeana (Nees) Schiffn. [14]Dichiton integerrimum (Lindb.) H. Buch = Cephaloziella integerrima (Lindb.) Warnst.Dichiton perpusillum Mont. nom. illeg. = Cephaloziella calyculata (Durieu & Mont.) Müll.Frib.Dichominum cruciatum (L.) Trevis. = Lunularia cruciata (L.) Lindb.Dicranolejeunea johnsoniana (Mitt.) Vanden Berghen ex S. W. Arnell = Acanthocoleusaberrans (Lindenb. & Gottsche) KruijtDilaena blyttii (Moerch ex Hornem.) Dumort. = Moerckia blyttii (Moerch ex Hornem.)Brockm.Dilaena hibernica (Hook.) Dumort. = Moerckia hibernica (Hook.) GottscheDilaena lyellii (Hook.) Dumort. = Pallavicinia lyellii (Hook.) Carruth.Diplolaena blyttii (Moerch ex Hornem.) Nees = Moerckia blyttii (Moerch ex Hornem.)Brockm.Diplolaena lyellii (Hook.) Dumort. = Pallavicinia lyellii (Hook.) Carruth.Diplolaena lyellii var. hibernica (Hook.) Nees = Moerckia hibernica (Hook.) Gottsche☞ Diplolaena lyellii var. lamellata (Nees) Macvicar = Petalophyllum ralfsii (Wilson) Nees& GottscheDiplolaena lyellii f. lamellata Nees = Petalophyllum ralfsii (Wilson) Nees & GottscheDiplomitrium blyttii (Moerch ex Hornem.) Corda = Moerckia blyttii (Moerch ex Hornem.)Brockm.Diplomitrium hibernicum (Hook.) Corda = Moerckia hibernica (Hook.) GottscheDiplophylleia albicans (L.) Trevis. = Diplophyllum albicans (L.) Dumort.Diplophylleia minuta (Schreb.) Trevis. = Anastrophyllum minutum (Schreb.) R. M. Schust.Diplophylleia obtusifolia (Hook.) Trevis. = Diplophyllum obtusifolium (Hook.) Dumort.Diplophylleia taxifolia (Wahlenb.) Trevis. = Diplophyllum taxifolium (Wahlenb.) Dumort.Diplophyllum apiculatum var . obtusatum R. M. Schust. = Diplophyllum obtusatum (R. M.Schust.) R. M. Schust.Diplophyllum dicksoni (Hook.) Dumort. = Douinia ovata (Dicks.) H. BuchDiplophyllum gymnostomophilum (Kaal.) Kaal. = Scapania gymnostomophila Kaal.Diplophyllum ovatum (Dicks.) Lindb. ex Steph. = Douinia ovata (Dicks.) H. BuchDiplophyllum scapanioides (C. Massal.) Steph. = Scapania scapanioides (C. Massal.) GrolleDumortiera irrigua (Wilson ex Hook.) Nees = Dumortiera hirsuta (Sw.) NeesDumortiera spathysii (Lindenb.) Nees = Athalamia spathysii (Lindenb.) S. Hatt.Duriaea helicophylla Bory & Mont. = Riella helicophylla (Bory & Mont.) Mont.Duriaea notarisii (Mont.) Bory & Mont. nom. illeg. = Riella notarisii (Mont.) Mont.Duvalia pilosa (Hornem.) Lindb. = Mannia pilosa (Hornem.) Frye & L. ClarkDuvalia rupestris Nees = Mannia triandra (Scop.) GrolleEucalyx crenulatus (Mitt.) Loeske = Jungermannia gracillima Sm.Eucalyx gracillimus (Sm.) Loeske = Jungermannia gracillima Sm.☞ Eucalyx hyalinus (Lyell) Breidl. = Jungermannia hyalina LyellEucalyx muellerianus (Schiffn.) Müll. Frib. = Jungermannia hyalina LyellEucalyx obovatus (Nees) Breidl. = Jungermannia obovata Nees

394 R.M. Ros et al.Eucalyx subellipticus (Lindb. ex Kaal.) Breidl. = Jungermannia subelliptica (Lindb. exKaal.) LevierEucephalozia bicuspidata (L.) Schiffn. = Cephalozia bicuspidata (L.) Dumort.Eulejeunea canariensis Steph. = Lejeunea canariensis (Steph.) Steph.Eulejeunea cavifolia (Ehrh.) Lindb. = Lejeunea cavifolia (Ehrh.) Lindb.Eulejeunea flava (Sw.) Steph. = Lejeunea flava (Sw.) NeesEulejeunea lamacerina Steph. = Lejeunea lamacerina (Steph.) Schiffn.Eulejeunea pitardii Steph. = Lejeunea eckloniana Lindenb.Eulejeunea serpyllifolia (Dicks.) Schiffn. nom. illeg. = Lejeunea patens Lindb. [33]? Eulejeunea serpyllifolia var. cavifolia (Ehrh.) Lindb. = Lejeunea cavifolia (Ehrh.) Lindb.? Eulejeunea serpyllifolia var. patens (Lindb.) A. L. M. Guim. = Lejeunea patens Lindb.Eulejeunea ulicina (Taylor) Schiffn. = Microlejeunea ulicina (Taylor) A. EvansEvansia dentata (Raddi) Douin = Cephaloziella dentata (Raddi) Steph.Exormotheca africana Steph. = Exormotheca pustulosa Mitt.Exormotheca bullosa sensu Müll. Frib. = Exormotheca welwitschii Steph.Fegatella conica (L.) Corda = Conocephalum conicum (L.) Dumort.Fegatella officinalis Raddi nom. superfl. = Conocephalum conicum (L.) Dumort.Fimbriaria africana Mont. = Asterella africana (Mont.) A. EvansFimbriaria blumeana Nees = Asterella blumeana (Nees) Kachroo REJECTED FROMTHE AREAFimbriaria bonjeanii De Not. = Asterella lindenbergiana (Corda ex Nees) ArnellFimbriaria elegans auct. eur. non Spreng. = Asterella africana (Mont.) A. EvansFimbriaria fragrans (Balbis)Nees = Mannia fragrans (Balbis) Frye & L. Clark [10]Fimbriaria fragrans auct. = Asterella saccata (Wahlenb.) A. Evans. [10]Fimbriaria lindenbergiana Corda ex Nees = Asterella lindenbergiana (Corda ex Nees)ArnellFimbriaria ludwigii auct. non (Schwägr.) Limpr. = Asterella gracilis (F. Weber) Underw.Fimbriaria ludwigii (Schwägr.) Limpr. ex Müll. Frib. nom. inval. = Mannia triandra (Scop.)GrolleFimbriaria nana Lindenb. = Athalamia hyalina (Sommerf.) S. Hatt. [9]Fimbriaria pilosa (Wahlenb.) Nees = Asterella gracilis (F. Weber) Underw.Fimbriaria raddii Corda ex Nees = Asterella wallichiana (Lehm. & Lindenb.) Pandé et al .REJECTED FROM THE AREAFimbriaria saccata (Wahlenb.) Nees = Asterella saccata (Wahlenb.) A. EvansFimbriaria stahlii Steph. = Asterella pringlei Underw. REJECTED FROM THE AREAFimbriaria tenella auct. non L. = Asterella gracilis (F. Weber) Underw.Fossombronia angulosa var. caespitiformis Raddi = Fossombronia caespitiformis De Not.ex Rabenh.Fossombronia caespitiformis var. husnotii (Corb.) Corb. = Fossombronia caespitiformissubsp. multispira (Schiffn.) J.R. Bray and D.C. Cargill in Stotler et al .Fossombronia caespitiformis var. intermedia (Corb. ex Trab.) Trab. ex Bonner nom. inval. =Fossombronia echinata Macvicar [20]Fossombronia caespitiformis var. subcristata Schiffn. = Fossombronia caespitiformis DeNot. ex Rabenh. subsp. caespitiformisFossombronia corbulaeformis Trab. = Petalophyllum ralfsii (Wilson) Nees & Gottsche# Fossombronia crispula (Brot.) R. M. Schust. [21]Fossombronia cristata Lindb. = Fossombronia wondraczekii (Corda) Lindb.Fossombronia cristata var. wondraczekii (Corda) Lindb . = Fossombronia wondraczekii(Corda) Lindb.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 395Fossombronia crozalsii Corb. = Fossombronia mittenii Tind.Fossombronia dumortieri Huebener & Genth ex Lindb. nom. inval. = Fossombroniafoveolata Lindb.Fossombronia husnotii Corb. = Fossombronia caespitiformis subsp. multispira (Schiffn.)J.R. Bray and D.C. Cargill in Stotler et al .Fossombronia intermedia Corb. ex Trab. nom. inval. = Fossombronia echinata Macvicar[20]Fossombronia intestinalis Taylor REJECTED FROM THE AREA [19]Fossombronia loitlesbergeri Schiffn. = Fossombronia pusilla (L.) Nees [23]Fossombronia mauritanica Trab. nom. inval . = Fossombronia caespitiformis subsp.multispira (Schiffn.) J. R. Bray and D. C. Cargill in Stotler et al .Fossombronia maritima (Paton) Paton = Fossombronia pusilla (L.) NeesFossombronia pusilla var. corsa Lindb. ex Bonner nom. nud. = Fossombronia pusilla (L.)NeesFossombronia pusilla var. decipiens Corb. = Fossombronia pusilla (L.) NeesFossombronia pusilla var. maritima Paton = Fossombronia pusilla (L.) Nees Fossombronia pusilla var. major Jur. nom. nud. = Fossombronia pusilla (L.) NeesFossombronia verrucosa Lindb. = Fossombronia caespitiformis subsp. multispira (Schiffn.)J. R. Bray and D. C. Cargill in Stotler et al .Frullania aeolotis Mont. & Nees = Frullania riparia Hampe ex Lehm.Frullania bryhnii Müll. Frib. = Frullania polysticta Lindenb.Frullania calcarifera Steph. = Frullania tamarisci (L.) Dumort.Frullania caucasica Steph. = Frullania parvistipula Steph.Frullania cesatiana De Not. = Frullania riparia Hampe ex Lehm.Frullania cesatiana var. muscicola (Steph.) Bisang et al . nom. nud. REJECTED FROMTHE AREA [24]Frullania cesatiana var. muscicola auct. eur. non (Steph.) Bisang et al . nom. nud. = Frullaniaazorica Sim-Sim et al. [24]Frullania cleistostoma Schiffn. & Wollny = Frullania inflata GottscheFrullania davurica auct. eur. non Hampe = Frullania jackii Gottsche [26]Frullania davurica subsp. jackii (Gottsche) S. Hatt. = Frullania jackii GottscheFrullania dilatata var. anomala Corb. = Frullania dilatata (L.) Dumort.Frullania dilatata var. briziana C.Massal. = Frullania dilatata (L.) Dumort.Frullania dilatata var. elongata Losada Lima & Vanden Berghen [25]Frullania dilatata var. microphylla (Wallroth) Nees = Frullania dilatata (L.) Dumort.Frullania eboracencis subsp. caucasica (Steph.) R. M. Schust. = Frullania parvistipulaSteph.Frullania eboracencis subsp. parvistipula (Steph.) R. M. Schust. = Frullania parvistipulaSteph.Frullania fragilifolia var. parvistipula (Steph.) Schiffn. = Frullania parvistipula Steph.Frullania funchalensis Gottsche nom. nud. = Frullania dilatata (L.) Dumort.Frullania germana (Taylor) Taylor = Frullania teneriffae (F. Weber) NeesFrullania hispanica Nees = Frullania tamarisci (L.) Dumort.Frullania hispanica Hampe non Nees = Frullania polysticta Lindenb.Frullania hutchinsiae (Hook.) Nees = Jubula hutchinsiae (Hook.) Dumort.Frullania illyrica Grolle = Frullania inflata Gottsche? Frullania inflata subsp. illyrica (Grolle) R. M. Schust. = Frullania inflata GottscheFrullania inflata var. illyrica (Grolle) R. M. Schust. = Frullania inflata GottscheFrullania major Raddi nom. superfl. = Frullania tamarisci (L.) Dumort.

396 R.M. Ros et al.Frullania maritima Steph. p. p. = Frullania tamarisci (L.) Dumort.Frullania microphylla var. deciduifolia Grolle = Frullania microphylla (Gottsche) PearsonFrullania muscicola Steph. REJECTED FROM THE AREA [24]Frullania muscicola sensu Sérgio non Steph. = Frullania azorica Sim-Sim et al. [24]Frullania nervosa Mont. = Frullania tamarisci (L.) Dumort.Frullania nervosa var. microcellularis Müll. Frib. nom. nud. = Frullania tamarisci (L.)Dumort.Frullania nervosa var. polysticta Nees nom. herb. = Frullania polysticta Lindenb.Frullania obscurifolia Mitt. REJECTED FROM THE AREAFrullania saxicola Austin = Frullania inflata GottscheFrullania squarrosa Dumort. = Frullania ericoides (Nees) Mont.Frullaniatamarisci var. atrovirens Carrington = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. calcarifera Steph. = Frullania tamarisci (L.) Dumort.Frullaniatamarisci var. hispanica (Nees) Nees = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. mediterranea (De Not.) Nees = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. mediterranea f. robustior Schiffn. = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. heterophylla Corb. = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. heterophylla f. brunnea Schiffn. = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. microphylla Gottsche = Frullania microphylla (Gottsche) PearsonFrullania tamarisci var. robusta Lindb. = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. sardoa (De Not.) De Not. = Frullania tamarisci (L.) Dumort.Frullania tamarisci var. schiffneri Nicholl = Frullania tamarisci (L.) Dumort.Frullania tenera Lindb. ex Steph. non Spruce = Frullania parvistipula Steph.Frullania trabutiana Steph. = Frullania dilatata (L.) Dumort.Frullania welwitschii Steph. = Frullania ericoides (Nees) Mont.Frullania willkommii Steph. = Frullania tamarisci (L.) Dumort.Gongylanthus flagelliferus (Raddi) Nees = Gongylanthus ericetorum (Raddi) NeesGrimaldia androgyna (L.) Lindb. = Mannia androgyna (L.) A. EvansGrimaldia angustifolia (Neck .)Lindb. ex Schiffn. = Mannia androgyna (L.) A. EvansGrimaldia atlantica Trab. nom. inval. [39]Grimaldia barbifrons Bisch. = Mannia fragrans (Balbis) Frye & L. ClarkGrimaldia carnica C.Massal. = Mannia pilosa (Hornem.) Frye & L. ClarkGrimaldia dichotoma Raddi nom. illeg. = Mannia androgyna (L.) A. EvansGrimaldia fragrans (Balbis) Corda = Mannia fragrans (Balbis) Frye & L. Clark [10]Grimaldia hemisphaerica (L.) Lindenb. = Reboulia hemisphaerica (L.) RaddiGrimaldia pilosa (Hornem.) Lindb. = Mannia pilosa (Hornem.) Frye & L. ClarkGrimaldia punicea Wallr. = Athalamia hyalina (Sommerf.) S. Hatt.Grimaldia rupestris (Nees) Lindenb. = Mannia triandra (Scop.) GrolleGrimaldia triandra (Scop.) Lindb. = Mannia triandra (Scop.) GrolleGrimaldia ventricosa Wallr. = Reboulia hemisphaerica (L.) RaddiGuentheria graveolens Trevis. = Corsinia coriandrina (Spreng.) Lindb.Gymnocolea acutiloba (Schiffn.) Müll. Frib. = Gymnocolea inflata (Huds.) Dumort.Gymnocolea huebeneriana (Nees) Dumort. = Hygrobiella laxifolia (Hook.) SpruceGymnocolea inflata subsp. acutiloba (Schiffn.) R. M. Schust. & Damsh. ex L. Söderstr. &Vá≈ a = Gymnocolea inflata (Huds.) Dumort.Gymnocolea inflata var . acutiloba (Schiffn.) S. W. Arnell = Gymnocolea inflata (Huds.)Dumort.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 397Gymnocolea inflata var. heterostipa (Carrington & Spruce) Müll. Frib. = Gymnocoleainflata (Huds.) Dumort.Gymnocolea laxifolia (Hook.) Dumort. = Hygrobiella laxifolia (Hook.) SpruceGymnomitrion adustum Nees emend. Limpr. = Marsupella adusta (Nees emend. Limpr.)SpruceGymnomitrion alpinum (Gottsche ex Husn.) Schiffn. = Marsupella alpina (Gottsche exHusn.) BernetGymnomitrion cochleare (Lindb.) Müll. Frib. = Marsupella brevissima (Dumort.) GrolleGymnomitrion commutatum (Limpr.) Schiffn. = Marsupella commutata (Limpr.) BernetGymnomitrion concinnatum var. intermedium Limpr. = Gymnomitrion concinnatum(Lightf.) CordaGymnomitrion condensatum Ångstr. ex C. Hartm. = Marsupella condensata (Ångstr. ex C.Hartm.) Kaal.Gymnomitrion confertum (Limpr.) Limpr. = Marsupella brevissima (Dumort.) GrolleGymnomitrion crassifolium Carrington = Marsupella brevissima (Dumort.) GrolleGymnomitrion revolutum (Nees) Philib. = Apomarsupella revoluta (Nees) R. M. Schust.Gymnomitrion varians (Lindb.) Schiffn. = Marsupella brevissima (Dumort.) GrolleHaplozia amplexicaulis (Dumort.) Warnst. = Jungermannia sphaerocarpa Hook.Haplozia atrovirens (Dumort.) Müll. Frib. = Jungermannia atrovirens Dumort.Haplozia autumnalis (DC.) Warnst. = Jamesoniella autumnalis (DC.) Steph.Haplozia breidleri Müll. Frib. = Jungermannia confertissima NeesHaplozia cordifolia (Dumort.) Müll. Frib. = Jungermannia exsertifolia subsp. cordifolia(Dumort.) Vá≈ aHaplozia crenulata (Mitt.) Müll. Frib. = Jungermannia gracillima Sm.Haplozia crenulata var. gracillima (Sm.) C. E. O. Jensen = Jungermannia gracillima Sm.Haplozia crenulata f. gracillima (Sm.) Müll. Frib. = Jungermannia gracillima Sm.Haplozia hyalina (Lyell) Warnst. = Jungermannia hyalina LyellHaplozia levieri (Steph.) Müll. Frib. = Jungermannia confertissima NeesHaplozia lurida (Dumort.) Müll. Frib. = Jungermannia sphaerocarpa Hook.Haplozia muelleriana (Schiffn.) Casares-Gil = Jungermannia hyalina LyellHaplozia nigrella (De Not.) Dumort. = Southbya nigrella (De Not.) Henriq.Haplozia oblongifolia Müll. Frib. = Jungermannia pumila With.Haplozia obovata (Nees) Casares-Gil = Jungermannia obovata NeesHaplozia polaris (Lindb.) Müll. Frib. = Jungermannia polaris Lindb.Haplozia pumila (With.) Müll. Frib. = Jungermannia pumila With.Haplozia riparia (Taylor) Müll. Frib. = Jungermannia atrovirens Dumort.Haplozia riparia var. rivularis (Bernet) Müll. Frib. = Jungermannia atrovirens Dumort.Haplozia rivularis (Schiffn.) Casares-Gil = Jungermannia pumila With.Haplozia schiffneri (Loitl.) Müll. Frib. = Jungermannia polaris Lindb.Haplozia schraderi (Mart.) Dumort. = Jamesoniella autumnalis (DC.) Steph.Haplozia sphaerocarpa (Hook.) Müll. Frib. = Jungermannia sphaerocarpa Hook.Haplozia sphaerocarpa var. amplexicaulis (Dumort.) Müll. Frib. = Jungermanniasphaerocarpa Hook.Haplozia sphaerocarpa var. nana (Nees ex Flot.) Müll. Frib. = Jungermannia sphaerocarpaHook.Haplozia subelliptica (Lindb. ex Kaal.) Casares-Gil = Jungermannia subelliptica (Lindb. exKaal.) LevierHaplozia tristis (Nees) Müll. Frib.= Jungermannia atrovirens Dumort.

398 R.M. Ros et al.Harpalejeunea ovata (Hook.) Schiffn. non (Dicks.) Schiffn. = Harpalejeunea molleri(Steph.) GrolleHepatica conica (L.) Lindb. = Conocephalum conicum (L.) Dumort.Herpetium deflexum (Mart.) Nees = Bazzania tricrenata (Wahlenb.) Lindb.Herpetium reptans (L.) Nees = Lepidozia reptans (L.) Dumort.Herpetium trilobatum (L.) Nees = Bazzania trilobata (L.) GrayHomalolejeunea mackaii (Hook.) Gray ex Casares-Gil = Marchesinia mackaii (Hook.)GrayHygrobiella myriocarpa (Carrington) Spruce = Eremonotus myriocarpus (Carrington)PearsonHygrophila irrigua (Wilson ex Hook.) Taylor nom. superfl. = Dumortiera hirsuta (Sw.)NeesHypenantron africanum (Mont.) Kuntze = Asterella africana (Mont.) A. Evans? Hypenantron lindenbergianum (Corda ex Nees) Kuntze = Asterella lindenbergiana(Corda ex Nees) ArnellHypnenatron nanum (Lindenb.) Trevis. = Athalamia hyalina (Sommerf.) S. Hatt.Isopaches bicrenatus (Schmidel ex Hoffm.) H. Buch = Lophozia bicrenata (Schmidel exHoffm.) Dumort.Isopaches decolorans (Limpr.) H. Buch = Lophozia decolorans (Limpr.) Steph.Isopaches hellerianus (Nees ex Lindenb.) H. Buch = Anastrophyllum hellerianum (Nees exLindenb.) R. M. Schust.Jamesoniella autumnalis f. subapicalis (Nees) Müll. Frib. = Jamesoniella autumnalis (DC.)Steph.Jamesoniella subapicalis (Nees) Schiffn. = Jamesoniella autumnalis (DC.) Steph.Jubula hutchinsiae subsp. hutchinsiae var. integrifolia (Steph.) S. Hatt. [27]Jubula hutchinsiae var. sullivanti Spruce [27]Jubula hutchinsiae var. warburgii auct. eur. non Schiffn. = Jubula hutchinsiae (Hook.)Dumort. subsp. hutchinsiaeJubula javanica Steph. = Jubula hutchinsiae subsp. javanica (Steph.) Verd.Jubula pennsylvanica (Steph.) A. Evans [27]Jungermannia acuta Lindenb. nom. illeg. = Leiocolea collaris (Nees) SchljakovJungermannia acuta var. aeruginosa Lindenb. = Leiocolea turbinata (Raddi) H. BuchJungermannia acutiloba Kaal. non Hook. f. & Taylor = Gymnocolea inflata (Huds.)Dumort.Jungermannia aequiloba Schwägr. = Scapania aequiloba (Schwägr.) Dumort.Jungermannia affinis Wilson ex Hook. nom. inval. = Leiocolea turbinata (Raddi) H. BuchJungermannia albescens Hook. = Pleurocladula albescens (Hook.) GrolleJungermannia albicans L. = Diplophyllum albicans (L.) Dumort.# Jungermannia albicans var. rubescens Grognot = Diplophyllum albicans (L.) Dumort.Jungermannia albicans var. taxifolia (Wahlenb.) Wallr. = Diplophyllum taxifolium(Wahlenb.) Dumort.Jungermannia algeriensis Gottsche & Rabenh. = Leiocolea turbinata (Raddi) H. BuchJungermannia alicularia De Not. = Southbya tophacea (Spruce) SpruceJungermannia alpestris Schleich. ex F. Weber = Leiocolea collaris (Nees) SchljakovJungermannia alpestris auct. non Schleich. ex F. Weber = Lophozia sudetica (Nees exHuebener) GrolleJungermannia amplexicaulis Dumort. = Jungermannia sphaerocarpa Hook.Jungermannia angulosa Dicks. = Fossombronia angulosa (Dicks.) RaddiJungermannia anomala Hook. = Mylia anomala (Hook.) Gray

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 399Jungermannia aquilegia Hook. f. & Taylor = Radula aquilegia (Hook. f. & Taylor)Gottsche et al.Jungermannia arboris-vitae With. = Porella arboris-vitae (With.) GrolleJungermannia arenaria Nees = Lophozia excisa (Dicks.) Dumort.Jungermannia asplenioides L. emend. Taylor = Plagiochila asplenioides (L. emend. Taylor)Dumort.Jungermannia assimilis Mitt. = Anastrophyllum assimile (Mitt.) Steph.Jungermannia attenuata (Mart.) Lindenb. = Barbilophozia attenuata (Mart.) LoeskeJungermannia attenuata var. gracilis Schleich. ex Lindenb. = Barbilophozia attenuata(Mart.) Loeske# Jungermannia attenuata var. gradata Desm. = Barbilophozia attenuata (Mart.) LoeskeJungermannia autumnalis DC. = Jamesoniella autumnalis (DC.) Steph.Jungermannia badensis Gottsche = Leiocolea badensis (Gottsche) Schiffn.Jungermannia bantriensis Hook. = Leiocolea bantriensis (Hook.) Jörg.Jungermannia bantriensis var. acuta Lindb. = Leiocolea collaris (Nees) SchljakovJungermannia bantriensis var. muelleri (Nees ex Lindenb.) Lindb. = Leiocolea collaris(Nees) SchljakovJungermannia barbata Schmidel ex Schreb. = Barbilophozia barbata (Schmidel ex Schreb.)LoeskeJungermannia barbata var. attenuata (Mart.) Nees = Barbilophozia attenuata (Mart.)LoeskeJungermannia barbata var. floerkei (F. Weber & D. Mohr) Dumort. = Barbilophoziafloerkei (F. Weber & D. Mohr) LoeskeJungermannia barbata var. lycopodioides (Wallr.) Nees = Barbilophozia lycopodioides(Wallr.) LoeskeJungermannia barbata var. quinquedentata (Huds.) Nees = Tritomaria quinquedentata(Huds.) H. BuchJungermannia barbata var. schreberi Nees nom. illeg. = Barbilophozia barbata (Schmidel exSchreb.) LoeskeJungermannia bartlingii Hampe ex Nees = Scapania cuspiduligera (Nees) Müll. Frib.Jungermannia baueri Mart. = Nowellia curvifolia (Dicks.) Mitt.Jungermannia baueriana (Schiffn.) Arnell = Barbilophozia hatcheri (A. Evans) LoeskeJungermannia bicalyculata Raddi = Cephalozia bicuspidata (L.) Dumort.Jungermannia bicornis F. Weber & D. Mohr = Anastrophyllum minutum (Schreb.) R. M.Schust.Jungermannia bicrenata Schmidel ex Hoffm. = Lophozia bicrenata (Schmidel ex Hoffm.)Dumort.Jungermannia bicuspidata L. = Cephalozia bicuspidata (L.) Dumort.Jungermannia bidentata L. = Lophocolea bidentata (L.) Dumort.Jungermannia bifaria Sw. = Plagiochila bifaria (Sw.) Lindenb.Jungermannia blasia DC. = Blasia pusilla L.Jungermannia blyttii Moerch ex Hornem. = Moerckia blyttii (Moerch ex Hornem.) Brockm.Jungermannia breidleri (Müll. Frib.) Gams = Jungermannia confertissima NeesJungermannia byssacea A. Rothnom. rejic. = Cephaloziella hampeana (Nees) Schiffn.Jungermannia byssacea auct. non A.Roth = Cephaloziella divaricata (Sm.) Schiffn.Jungermannia calycina Taylor = Pellia endiviifolia (Dicks.) Dumort.Jungermannia calyculata Durieu & Mont. = Cephaloziella calyculata (Durieu & Mont.)Müll. Frib.Jungermannia calypogeia Raddi nom. illeg. = Calypogeia fissa (L.) RaddiJungermannia catenulata Huebener = Cephalozia catenulata (Huebener) Lindb.

400 R.M. Ros et al.Jungermannia cavifolia Ehrh. = Lejeunea cavifolia (Ehrh.) Lindb.Jungermannia chamedryfolia With. = Riccardia chamedryfolia (With.) GrolleJungermannia ciliaris L. = Ptilidium ciliare (L.) HampeJungermannia collaris Nees = Leiocolea collaris (Nees) SchljakovJungermannia compacta A. Roth = Scapania compacta (A. Roth) Dumort.Jungermannia compacta Mazziari nom. nud. non Roth [28]Jungermannia commutata Huebener = Lophozia bicrenata (Schmidel ex Hoffm.) Dumort.Jungermannia complanata L. = Radula complanata (L.) Dumort.Jungermannia compressa Hook. = Nardia compressa (Hook.) GrayJungermannia concinnata Lightf. = Gymnomitrion concinnatum (Lightf.) CordaJungermannia confervoides Raddi = Cephaloziella divaricata (Sm.) Schiffn.Jungermannia connivens Dicks. = Cephalozia connivens (Dicks.) Lindb.Jungermannia corcyraea Nees = Leiocolea turbinata (Raddi) H. BuchJungermannia cordaeana Huebener = Porella cordaeana (Huebener) Moore# Jungermannia cordifolia Brot.non Hook. [63]Jungermannia cordifolia Hook. non Brot. nec Ehrh. ex F. Weber nom. illeg. = Jungermanniaexsertifolia subsp. cordifolia (Dumort.) Vá≈ aJungermannia crassifolia Lindenb. & Gottsche = Cephalozia crassifolia (Lindenb. &Gottsche) FulfordJungermannia crenulata Sm. nom. illeg. non Schmidel = Jungermannia gracillima Sm.Jungermannia crenulata var. gracillima (Sm.) Hook. = Jungermannia gracillima Sm.# Jungermannia crispula Brot. [21]Jungermannia crocata De Not. = Lophocolea minor NeesJungermannia cuneifolia Hook. = Leptoscyphus cuneifolius (Hook.) Mitt.Jungermannia cupressina Sw. = Lepidozia cupressina (Sw.) Lindenb.Jungermannia curta Mart. = Scapania curta (Mart.) Dumort.Jungermannia curvifolia Dicks. = Nowellia curvifolia (Dicks.) Mitt.Jungermannia cuspiduligera Nees = Scapania cuspiduligera (Nees) Müll. Frib.Jungermannia cylindracea Dumort. = Lophozia excisa (Dicks.) Dumort.Jungermannia decipiens Hook. = Adelanthus decipiens (Hook.) Mitt.Jungermannia decolorans Limpr. = Lophozia decolorans (Limpr.) Steph.Jungermannia deflexa var. flaccida Schleich. ex Lindenb. nom. illeg. = Bazzania flaccida(Dumort.) GrolleJungermannia dentata Raddi = Cephaloziella dentata (Raddi) Steph.Jungermannia denudata (Mart.) Nees = Odontoschisma denudatum (Mart.) Dumort.Jungermannia dilatata L. = Frullania dilatata (L.) Dumort.Jungermannia distans Schwein. = Porella pinnata L.Jungermannia divaricata Sm. = Cephaloziella divaricata (Sm.) Schiffn.Jungermannia emarginata Ehrh. = Marsupella emarginata (Ehrh.) Dumort.Jungermannia endiviifolia Dicks. = Pellia endiviifolia (Dicks.) Dumort.Jungermannia epiphylla L. = Pellia epiphylla (L.) CordaJungermannia epiphylla var. furcigera Hook. = Pellia endiviifolia (Dicks.) Dumort.Jungermannia epiphylla var. longifolia Lam. & DC. = Pellia endiviifolia (Dicks.) Dumort.Jungermannia ericoides Nees = Frullania ericoides (Nees) Mont.Jungermannia excisa Dicks. = Lophozia excisa (Dicks.) Dumort.Jungermannia exigua Taylor = Plagiochila exigua (Taylor) TaylorJungermannia exsecta Schmidel ex Schrad. = Tritomaria exsecta (Schmidel ex Schrad.)Loeske

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 401Jungermannia exsectiformis Breidl. = Tritomaria exsectiformis (Breidl.) LoeskeJungermannia falcata Raddi nom. illeg. = Diplophyllum albicans (L.) Dumort.Jungermannia fennica Gottsche = Arnellia fennica (Gottsche) Lindb.Jungermannia fissa (L.) Scop. = Calypogeia fissa (L.) RaddiJungermannia fissidentoidea Huebener = Diplophyllum albicans (L.) Dumort.Jungermannia flaccida Huebener = Jungermannia obovata NeesJungermannia flava Sw. = Lejeunea flava (Sw.) NeesJungermannia floerkei F. Weber & D. Mohr = Barbilophozia floerkei (F. Weber &D. Mohr) LoeskeJungermannia flotoviana Nees = Harpanthus flotovianus (Nees) NeesJungermannia fluitans Nees = Cladopodiella fluitans (Nees) H. BuchJungermannia foscardiana Mazziari nom. nud. [28]Jungermannia fragilifolia Taylor = Frullania fragilifolia (Taylor) Gottsche et al .Jungermannia fragilis A. Roth = Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort.Jungermannia fragrans Moris & De Not. = Lophocolea fragrans (Moris & De Not.)Gottsche et al.Jungermannia francisci Hook. = Cladopodiella francisci (Hook.) Jörg.Jungermannia fruticulosa auct. eur. non (O. F. Müll.) Sm. = Metzgeria violacea (Ach.)Dumort. [40]Jungermannia fuciformis Bory [43]Jungermannia funckii F. Weber & D. Mohr = Marsupella funckii (F. Weber & D. Mohr)Dumort.Jungermannia furcata L. = Metzgeria furcata (L.) Dumort.Jungermannia germana Taylor = Frullania teneriffae (F. Weber) NeesJungermannia gillmanii Austin = Lophozia gillmanii (Austin) R. M. Schust.Jungermannia gracilis (Schleich. ex Lindenb.) Lindb. = Barbilophozia attenuata (Mart.)LoeskeJungermannia grandiretis Lindb. ex Kaal. = Lophozia grandiretis (Lindb. ex Kaal.) Schiffn.Jungermannia graveolens Schrad. = Geocalyx graveolens (Schrad.) NeesJungermannia grimsulana J. B. Jack ex Gottsche & Rabenh. = Cephaloziella grimsulana(J.B. Jack ex Gottsche & Rabenh.) Lacout.Jungermannia guilfortiana Mazziari nom. nud. [28]Jungermannia guttulata Lindb. & Arnell = Lophozia longiflora (Nees) Schiffn.Jungermannia hamatifolia Hook. = Drepanolejeunea hamatifolia (Hook.) Schiffn.Jungermannia hampeana Nees = Cephaloziella hampeana (Nees) Schiffn.Jungermannia hatcheri A. Evans = Barbilophozia hatcheri (A. Evans) LoeskeJungermannia helleriana Nees ex Lindenb. = Anastrophyllum hellerianum (Nees exLindenb.) R. M. Schust.Jungermannia heterocolpos Thed. ex Hartm. = Leiocolea heterocolpos (Thed. ex Hartm.)H. BuchJungermannia heterophylla Schrad. = Lophocolea heterophylla (Schrad.) Dumort.Jungermannia hibernica Hook. = Moerckia hibernica (Hook.) GottscheJungermannia hoffmannii Wallr. = Ptilidium ciliare (L.) HampeJungermannia hornschuchiana Nees = Leiocolea bantriensis (Hook.) Jörg.Jungermannia hutchinsiae Hook. = Jubula hutchinsiae (Hook.) Dumort. subsp. hutchinsiaeJungermannia incisa Schrad. = Lophozia incisa (Schrad.) Dumort.Jungermannia inconspicua Raddi = Cololejeunea minutissima (Sm.) Schiffn.Jungermannia inflata Huds. = Gymnocolea inflata (Huds.) Dumort.

402 R.M. Ros et al.Jungermannia inflata var. fluitans (Nees) Gottsche et al. = Cladopodiella fluitans (Nees)H. BuchJungermannia integrifolia Lehm. & Lindenb. = Chiloscyphus integrifolius (Lehm. &Lindenb.) Lehm. & Lindenb. REJECTED FROM THE AREAJungermannia jenseniana Grolle REJECTED FROM THE AREA [29]Jungermannia intermedia Lindenb. nom. illeg. = Lophozia excisa (Dicks.) Dumort.Jungermannia interrupta Nees = Pedinophyllum interruptum (Nees) Kaal.Jungermannia irrigua Nees = Scapania irrigua (Nees) NeesJungermannia julacea L. = Anthelia julacea (L.) Dumort.Jungermannia julacea var. clavuligera Nees = Anthelia juratzkana (Limpr.) Trevis.Jungermannia juratzkana Limpr. = Anthelia juratzkana (Limpr.) Trevis.Jungermannia karl-muelleri Grolle = Jungermannia pumila With.Jungermannia kaurinii Limpr. = Leiocolea gillmanii (Austin) R. M. Schust.Jungermannia kunzeana Huebener = Barbilophozia kunzeana (Huebener) Müll. Frib.Jungermannia lacera Mazziari nom. nud. [28]Jungermannia laevigata Schrad. = Porella arboris-vitae (With.) GrolleJungermannia lammersiana Huebener = Cephalozia bicuspidata (L.) Dumort.Jungermannia lanceolata L. nom. rejic. = Jungermannia atrovirens Dumort.Jungermannia lanceolata auct. non L. = Jungermannia leiantha GrolleJungermannia laurentiana De Not. = Leiocolea collaris (Nees) SchljakovJungermannia laxifolia Hook. = Hygrobiella laxifolia (Hook.) SpruceJungermannia leersii A. Roth = Ptilidium ciliare (L.) HampeJungermannia laevigata Schrad. = Porella arboris-vitae (With.) GrolleJungermannia levieri (Steph.) H. Buch, A. Evans & Verd. = Jungermannia confertissimaNeesJungermannia limosa Mazziari nom. nud. [28]Jungermannia limprichtii (Lindb. ex C. Massal. & Carestia) Warnst. = Lophozia excisa(Dicks.) Dumort.Jungermannia longidens Lindb. = Lophozia longidens (Lindb.) MacounJungermannia longiflora Nees = Lophozia longiflora (Nees) Schiffn.Jungermannia lunulifolia Dumort. = Cephalozia lunulifolia (Dumort.) Dumort.Jungermannia lurida Dumort. nom. illeg. = Jungermannia sphaerocarpa Hook.Jungermannia lycopodioides Wallr. = Barbilophozia lycopodioides (Wallr.) LoeskeJungermannia lycopodioides var. floerkei (F. Weber & D. Mohr) Lindb. = Barbilophoziafloerkei (F. Weber & D. Mohr) LoeskeJungermannia lyellii Hook. = Pallavicinia lyellii (Hook.) Carruth.Jungermannia lyonii Taylor = Tritomaria quinquedentata (Huds.) H. BuchJungermannia mackaii Hook. = Marchesinia mackaii (Hook.) GrayJungermannia macounii Austin = Cephalozia macounii (Austin) AustinJungermannia menzelii Corda = Cephalozia bicuspidata (L.) Dumort.Jungermannia michauxii F. Weber = Anastrophyllum michauxii (F. Weber) H. BuchJungermannia minuta Schreb. = Anastrophyllum minutum (Schreb.) R. M. Schust.Jungermannia minuta var. procera Nees = Anastrophyllum michauxii (F. Weber) H. BuchJungermannia minuta var. robusta C. Massal. & Carestia nom. inval. = Anastrophyllumassimile (Mitt.) Steph.Jungermannia minutissima Sm. = Cololejeunea minutissima (Sm.) Schiffn.Jungermannia muelleri Nees ex Lindenb. = Leiocolea collaris (Nees) SchljakovJungermannia multifida L. = Riccardia multifida (L.) Gray

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 403Jungermannia multifida var. sinuata Dicks. ex Hook. = Riccardia chamedryfolia (With.)GrolleJungermannia myriocarpa Carrington = Eremonotus myriocarpus (Carrington) PearsonJungermannia nana Nees ex Flot. = Jungermannia sphaerocarpa Hook.Jungermannia nardioides Lindb. = Anastrophyllum assimile (Mitt.) Steph.Jungermannia naumannii Nees ex Mart. = Barbilophozia floerkei (F. Weber & D. Mohr)LoeskeJungermannia nemorea L. = Scapania nemorea (L.) GrolleJungermannia nemorosa L. nom. illeg. = Scapania nemorea (L.) GrolleJungermannia nemorosa var. recurvifolia Hook. = Scapania gracilis Lindb.Jungermannia nevicensis Carrington = Marsupella boeckii (Austin) Lindb.Jungermannia nigrella De Not. = Southbya nigrella (De Not.) Henriq.Jungermannia nigricans Lam. nom. illeg. = Frullania tamarisci (L.) Dumort.Jungermannia obtusa Lindb. = Lophozia obtusa (Lindb.) A. EvansJungermannia obtusifolia Hook. = Diplophyllum obtusifolium (Hook.) Dumort.# Jungermannia oppositifolia Lacroix = Gongylanthus ericetorum (Raddi) NeesJungermannia orcadensis Hook. = Anastrepta orcadensis (Hook.) Schiffn.Jungermannia pallescens Ehrh. ex Hoffm. = Chiloscyphus pallescens (Ehrh. ex Hoffm.)Dumort.Jungermannia pallescens var. rivularis Schrad. = Chiloscyphus polyanthos (L.) CordaJungermannia palmata Hedw. = Riccardia palmata (Hedw.) Carruth.Jungermannia pauciflora Dicks. = Kurzia pauciflora (Dicks.) GrolleJungermannia pinguis L. = Aneura pinguis (L.) Dumort.Jungermannia platyphylla L. = Porella platyphylla (L.) Pfeiff.Jungermannia platyphylla var. thuja Hook. = Porella arboris-vitae (With.) GrolleJungermannia platyphylla var. canariensis F. Weber = Porella canariensis (F. Weber)Underw.Jungermannia platyphylloidea Schwein. = Porella platyphylla (L.) Pfeiff.Jungermannia pleniceps Austin = Cephalozia pleniceps (Austin) Lindb.Jungermannia plicata Hartm. = Barbilophozia kunzeana (Huebener) Müll. Frib.Jungermannia polita Nees = Tritomaria polita (Nees) Schiffn.Jungermannia polyanthos L. = Chiloscyphus polyanthos (L.) CordaJungermannia porella Dicks. nom. illeg. = Porella pinnata L.Jungermannia porelloides Torrey ex Nees = Plagiochila porelloides (Torrey ex Nees)Lindenb.Jungermannia porphyroleuca Nees. nom. illeg. = Lophozia longiflora (Nees) Schiffn.Jungermannia porphyroleuca var. guttulata (Lindb. & Arnell) Warnst. = Lophozia longiflora(Nees) Schiffn.Jungermannia prostrata Sw. = Odontoschisma prostratum (Sw.) Trevis .Jungermannia pubescens Schrank = Apometzgeria pubescens (Schrank) Kuwah.Jungermannia pulcherrima Weber = Ptilidium pulcherrimum (Weber) Vain.Jungermannia pulvinata Raddi = Marsupella emarginata (Ehrh.) Dumort.Jungermannia pumila subsp. polaris (Lindb.) R. M. Schust. = Jungermannia polaris Lindb.Jungermannia pumila var. rivularis ( Schiffn.) Frye & L. Clark = Jungermannia pumilaWith.Jungermannia punctata Taylor = Plagiochila punctata (Taylor) TaylorJungermannia pusilla L. = Fossombronia pusilla (L.) NeesJungermannia quinquedentata Huds. = Tritomaria quinquedentata (Huds.) H. Buch

404 R.M. Ros et al.Jungermannia quinquedentata var. attenuata (Mart.) Steud. = Barbilophozia attenuata(Mart.) LoeskeJungermannia raddiana C. Massal. = Cephaloziella rubella (Nees) Warnst.Jungermannia ralfsii Wilson = Petalophyllum ralfsii (Wilson) Nees & GottscheJungermannia reclusa Taylor = Cephalozia catenulata (Huebener) Lindb.Jungermannia reichardtii Gottsche ex Jur. = Anastrophyllum assimile (Mitt.) Steph.Jungermannia reptans L. = Lepidozia reptans (L.) Dumort.Jungermannia resupinata Hook. nom. inval. = Scapania compacta (A. Roth) Dumort.Jungermannia resupinata L. = Scapania undulata (L.) Dumort.Jungermannia rigida Wallr. nom. illeg. = Aneura pinguis (L.) Dumort.Jungermannia rigidula Huebener nom. illeg. = Cephalozia bicuspidata (L.) Dumort.Jungermannia riparia Taylor = Jungermannia atrovirens Dumort.$ Jungermannia riparia var. rotundifolia= Jungermannia atrovirens Dumort.Jungermannia rosacea Corda = Scapania curta (Mart.) Dumort.Jungermannia rostellata Huebener = Jungermannia pumila With.Jungermannia rubella Nees = Cephaloziella rubella (Nees) Warnst.Jungermannia saxicola Schrad. = Anastrophyllum saxicola (Schrad.) R. M. Schust.Jungermannia scalariformis Nees = Jungermannia confertissima NeesJungermannia scalaris Schmidel ex Schreb. nom. dub. [12]Jungermannia scalaris Hook. homon. illeg. = Nardia scalaris GrayJungermannia scalaris var. denudata Mart. = Odontoschisma denudatum (Mart.) Dumort.Jungermannia scalaris var. minor Nees = Nardia geoscyphus (De Not.) Lindb.Jungermannia scapanioides C. Massal. = Scapania scapanioides (C. Massal.) GrolleJungermannia schiffneri (Loitl.) A. Evans = Jungermannia polaris Lindb.Jungermannia schraderi Mart. = Jamesoniella autumnalis (DC.) Steph.Jungermannia schraderi auct. non Mart. = Jamesoniella undulifolia (Nees) Müll. Frib.Jungermannia schreberi S. Hatt. nom. inval. = Barbilophozia barbata (Schmidel) LoeskeJungermannia scitula Taylor = Tritomaria scitula (Taylor) Jörg.Jungermannia scutata F. Weber & D. Mohr = Harpanthus scutatus (F. Weber & D. Mohr)SpruceJungermannia serpyllifolia var. ovata auct. non Hook. = Harpalejeunea molleri (Steph.)GrolleJungermannia setiformis Ehrh. = Tetralophozia setiformis (Ehrh.) SchljakovJungermannia sinuata Dicks. nom. illeg. = Riccardia chamedryfolia (With.) GrolleJungermannia socia Nees = Lophozia excisa (Dicks.) Dumort.Jungermannia sphacelata Gieseke ex Lindenb. = Marsupella sphacelata (Gieseke exLindenb.) Dumort.Jungermannia sphaerocarpa var. amplexicaulis (Dumort.) Frye & L. Clark = Jungermanniasphaerocarpa Hook.Jungermannia sphaerocarpoidea De Not. = Jungermannia atrovirens Dumort.Jungermannia sphagni Dicks. = Odontoschisma sphagni (Dicks.) Dumort.Jungermannia spinulosa Dicks. = Plagiochila spinulosa (Dicks.) Dumort.Jungermannia sprengelii Mart. = Calypogeia fissa (L.) RaddiJungermannia starkei auct. non Funk ex Nees nom. illeg. = Cephaloziella divaricata (Sm.)Schiffn.Jungermannia stellulifera Taylor ex Carrington & Pearson = Cephaloziella divaricata (Sm.)Schiffn.Jungermannia stillicidiorum (Raddi) De Not. = Southbya tophacea (Spruce) SpruceJungermannia stipulacea Hook. = Harpanthus scutatus (F. Weber & D. Mohr) Spruce

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 405Jungermannia subalpina Nees ex Lindenb. = Scapania subalpina (Nees ex Lindenb.)DumortJungermannia subapicalis Nees = Jamesoniella autumnalis (DC.) Steph.Jungermannia sudetica Nees ex Huebener = Lophozia sudetica (Nees ex Huebener) GrolleJungermannia sullivantii Austin = Cephaloziella rubella (Nees) Warnst.Jungermannia tamarisci L. = Frullania tamarisci (L.) Dumort.Jungermannia tamarisci var. mediterranea De Not. = Frullania tamarisci (L.) Dumort.Jungermanniatamarisci var. sardoa De Not. = Frullania tamarisci (L.) Dumort.Jungermannia tamariscifolia L. nom. illeg. = Frullania tamarisci (L.) Dumort.Jungermannia taxifolia Wahlenb. = Diplophyllum taxifolium (Wahlenb.) Dumort.Jungermannia taylorii Hook. = Mylia taylorii (Hook.) GrayJungermannia taylorii var. anomala (Hook.) Nees = Mylia anomala (Hook.) GrayJungermannia teneriffae F. Weber = Frullania teneriffae (F. Weber) NeesJungermannia tenerrima Raddi nom. illeg. = Jungermannia gracillima Sm.Jungermannia tersa Nees = Jungermannia sphaerocarpa Hook.Jungermannia thuja Dicks.nom. illeg. = Porella arboris-vitae (With.) GrolleJungermannia thuja auct. non Dicks. nom. illeg. = Porella obtusata (Taylor) Trevis.Jungermannia tomentella Ehrh. = Trichocolea tomentella (Ehrh.) Dumort.Jungermannia tophacea Spruce = Southbya tophacea (Spruce) SpruceJungermannia trichomanis (L.) Scop. = Calypogeia fissa (L.) RaddiJungermannia trichomanis auct. non (L.) Scop. = Calypogeia azurea Stotler & Crotz [11]Jungermannia trichophylla L. = Blepharostoma trichophyllum (L.) Dumort.Jungermannia tricrenata Wahlenb. = Bazzania tricrenata (Wahlenb.) Lindb.Jungermannia trilobata L. = Bazzania trilobata (L.) GrayJungermannia tristis Nees = Jungermannia atrovirens Dumort.Jungermannia turbinata Raddi = Leiocolea turbinata (Raddi) H. BuchJungermannia turneri Hook. = Cephaloziella turneri (Hook.) Müll. Frib.Jungermannia ulicina Taylor = Microlejeunea ulicina (Taylor) A. EvansJungermannia uliginosa Sw. ex Lindenb. = Scapania uliginosa (Sw. ex Lindenb.) Dumort.Jungermannia umbrosa Schrad. = Scapania umbrosa (Schrad.) Dumort.Jungermannia undulata L. = Scapania undulata (L.) Dumort.Jungermannia varians Gottsche = Cephaloziella varians (Gottsche) Steph.Jungermannia ventricosa Dicks. = Lophozia ventricosa (Dicks.) Dumort.Jungermannia ventricosa var. guttulata (Lindb. & Arnell) C. Massal. = Lophozia longiflora(Nees) Schiffn.Jungermannia ventricosa var. longidens (Lindb.) C. Massal. & Carestia = Lophozia longidens(Lindb.) MacounJungermannia ventricosa var. porphyroleuca Hartm. = Lophozia longiflora (Nees) Schiffn.Jungermannia viticulosa L. = Saccogyna viticulosa (L.) Dumort.Jungermannia wenzelii Nees = Lophozia wenzelii (Nees) Steph.Jungermannia wilsonii (Nees)Hook. f. & Taylor = Acrobolbus wilsonii NeesJungermannia wilsoniana Nees = Leiocolea turbinata (Raddi) H. BuchJungermannia wondraczekii Corda = Fossombronia wondraczekii (Corda) Lindb.Jungermannia zeyheri Huebener = Jungermannia pumila With.Kantius(Kantia) argutus(arguta) (Nees & Mont.) Lindb. = Calypogeia arguta Nees & Mont.Kantius(Kantia) calypogeus(calypogea) (Raddi) Lindb. = Calypogeia fissa (L.) RaddiKantius(Kantia) fissus(fissa) (L.) Lindb. = Calypogeia fissa (L.) Raddi

406 R.M. Ros et al.Kantius(Kantia) muellerianus(muelleriana) Schiffn. = Calypogeia muelleriana (Schiffn.)Müll. Frib.Kantius (Kantia) neesianus (neesiana) (C. Massal. & Carestia) Arnell & C. E. O. Jensen =Calypogeia neesiana (C. Massal. & Carestia) Müll. Frib.Kantius(Kantia) sphagnicolus(sphagnicola) Arnell & J. Perss. = Calypogeia sphagnicola(Arnell & J. Perss.) Warnst. & LoeskeKantius(Kantia) sprengelii (Mart.) Trevis. = Calypogeia fissa (L.) RaddiKantius(Kantia) submersus(submersa) Arnell = Calypogeia sphagnicola (Arnell &J. Perss.) Warnst. & LoeskeKantius(Kantia) suecicus(suecica) Arnell & J. Perss. = Calypogeia suecica (Arnell. & J.Perss.) Müll. Frib.Kantius(Kantia) trichomanis (L.) Gray = Calypogeia fissa (L.) RaddiKantius(Kantia) trichomanis auct. non (L.) Gray = Calypogeia azurea Stotler & Crotz [11]Kantius(Kantia) trichomanis var. fissus(fissa) (L.) Lindb. = Calypogeia fissa (L.) RaddiKantius(Kantia) trichomanis var. neesianus(neesiana) C. Massal. & Carestia = Calypogeianeesiana (C. Massal. & Carestia) Müll. Frib.Leiocolea alpestris (Schleich. ex F. Weber) Isov. = Leiocolea collaris (Nees) SchljakovLeiocolea kaurinii (Limpr.) Jörg. = Leiocolea gillmanii (Austin) A. EvansLeiocolea muelleri (Nees ex Lindenb.) Jörg. = Leiocolea collaris (Nees) SchljakovLeiocolea obtusa (Lindb.) H. Buch = Lophozia obtusa (Lindb.) A. EvansLeiocolea turbinata f. aquatica Jelenc nom. nud. [32]Leioscyphus anomalus (Hook.) Steph. = Mylia anomala (Hook.) GrayLeioscyphus cuneifolius (Hook.) Steph. = Leptoscyphus cuneifolius (Hook.) Mitt.Lejeunea aberrans Lindenb. & Gottsche = Acanthocoleus aberrans (Lindenb. & Gottsche)KruijtLejeunea azorica Steph. = Lejeunea lamacerina (Steph.) Schiffn.Lejeunea calcarea Lib. = Cololejeunea calcarea (Lib.) Schiffn.Lejeunea calcarea var. rossettiana (C. Massal.) Mansion & Clerbois = Cololejeunea rossettiana(C. Massal.) Schiffn.Lejeunea cavifolia var. loitlesbergeri Schiffn. nom. inval. = Lejeunea cavifolia (Ehrh.)Lindb.Lejeunea diversiloba Spruce = Lejeunea hibernica Bischl. et al. ex GrolleLejeunea echinata (Hook.) Taylor ex Gottsche et al. nom. inval. = Cololejeunea calcarea(Lib.) Schiffn.Lejeunea flava subsp. moorei (Lindb.) R. M. Schust. [34]Lejeunea hamatifolia (Hook.) Spreng. = Drepanolejeunea hamatifolia (Hook.) Schiffn.Lejeunea hamatifolia Dumort. nom. inval. = Drepanolejeunea hamatifolia (Hook.) Schiffn.Lejeunea holtii Spruce = Lejeunea eckloniana Lindenb.Lejeunea inconspicua (Raddi) De Not. ex Rabenh. = Cololejeunea minutissima (Sm.)Schiffn.Lejeunea lamacerina var. azorica (Steph.) Greig-Smith = Lejeunea lamacerina (Steph.)Schiffn.Lejeunea mackaii (Hook.) Spreng. = Marchesinia mackaii (Hook.) GrayLejeunea macvicarii Pearson = Lejeunea mandonii (Steph.) Müll. Frib.Lejeunea microscopica (Taylor) Gottsche et al. = Aphanolejeunea microscopica (Taylor) A.EvansLejeunea minutissima (Sm.) Spreng. = Cololejeunea minutissima (Sm.) Schiffn.Lejeunea molleri Steph. = Harpalejeunea molleri (Steph.) GrolleLejeunea moorei Lindb. [34]Lejeunea ovata (Dicks.) Taylor ex Gottsche = Douinia ovata (Dicks.) H. Buch

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 407Lejeunea planiuscula (Lindb.) H. Buch = Lejeunea lamacerina (Steph.) Schiffn.Lejeunea rossettiana C. Massal. = Cololejeunea rossettiana (C. Massal.) Schiffn.Lejeunea serpyllifolia (Dicks.) Lib. nom. illeg. = Lejeunea patens Lindb. [33]Lejeunea serpyllifolia Scop. = Lejeunea cavifolia (Ehrh.) Lindb. [33]Lejeunea serpyllifolia var. cavifolia (Ehrh.) Lindb. nom. inval. = Lejeunea cavifolia (Ehrh.)Lindb.☞ Lejeunea serpyllifolia var. loitlesbergeri Schiffn. = Lejeunea cavifolia (Ehrh.) Lindb.Lejeunea serpyllifolia var. planiuscula C. Massal. nom. illeg. = Lejeunea lamacerina (Steph.)Schiffn.Lejeunea serpyllifolia f. planiuscula Lindb. nom. illeg. = Lejeunea lamacerina (Steph.)Schiffn.Lejeunea taylorii Spruce = Cololejeunea minutissima (Sm.) Schiffn.Lejeunea ulicina (Taylor) Gottsche et al. = Microlejeunea ulicina (Taylor) Steph.Lepidozia azorica H. Buch & Perss. = Telaranea azorica (H. Buch & Perss.) Pócs ex Schumacker& Vá≈ a [65]Lepidoziapearsonii Spruce REJECTED FROM THE AREA [35]Lepidozia pinnata (Hook.) Dumort. = Lepidozia cupressina (Sw.) Lindenb.Lepidozia reptans var. tenera (Huebener) Nees = Lepidozia reptans (L.) Dumort.Lepidozia setacea (Weber) Mitt. = Kurzia pauciflora (Dicks.) GrolleLepidozia sylvatica A. Evans = Kurzia sylvatica (A. Evans) GrolleLepidozia trichoclados Müll. Frib. = Kurzia trichoclados (Müll. Frib.) GrolleLeptoscyphus anomalus (Hook.) Lindb. = Mylia anomala (Hook.) GrayLeptoscyphus interruptus (Nees) Lindb. = Pedinophyllum interruptum (Nees) Kaal.Leptoscyphus taylorii (Hook.) Mitt. = Mylia taylorii (Hook.) GrayLiochlaena lanceolata Nees = Jungermannia leiantha GrolleLiochlaena subulata (A. Evans) Schljakov = Jungermannia subulata A. EvansLophocolea alata Mitt. ex Larter = Lophocolea bidentata (L.) Dumort.☞ Lophocolea bidentata var. alata (Mitt. ex Larter) Nees = Lophocolea bidentata (L.)Dumort.Lophocolea bidentata var. cuspidata (Nees) Nees = Lophocolea bidentata (L.) Dumort.Lophocolea bidentata var. fluitans F. A. Camus ex Husn. = Lophocolea bidentata (L.)Dumort.Lophocolea bidentata var. latifolia (Nees) Rabenh. nom. inval. = Lophocolea bidentata (L.)Dumort.Lophocolea bidentata var. rivularis (Raddi) Warnst. = Lophocolea bidentata (L.) Dumort.Lophocolea crocata (De Not.) Nees = Lophocolea minor NeesLophocolea cuspidata (Nees) Limpr. = Lophocolea bidentata (L.) Dumort. Lophocolea heterophylla f. gemmifera Andres = Lophocolea heterophylla (Schrad.)Dumort.Lophocolea heterophylla var. minor (Nees) Douin = Lophocolea minor NeesLophocolea heterophylla f. propagulifera Feld nom. inval. = Lophocolea heterophylla(Schrad.) Dumort.Lophocolea hirticalyx Corb. & Steph. = Lophocolea fragrans (Moris & De Not.) Gottscheet al .Lophocolea hookeriana Nees = Lophocolea bidentata (L.) Dumort.Lophocolea incisa Lindb. = Lophocolea heterophylla (Schrad.) Dumort.Lophocolea lateralis Dumort. = Lophocolea bidentata (L.) Dumort.Lophocolea latifolia Nees = Lophocolea bidentata (L.) Dumort.Lophocolea preauxiana Mont. = Saccogyna viticulosa (L.) Dumort.

408 R.M. Ros et al.☞ Lophocolea spicata Gottsche et al. = Lophocolea fragrans (Moris & De Not.) Gottscheet al .Lophocolea vogesiaca Huebener ex Nees = Harpanthus flotovianus (Nees) NeesLophozia acutiloba Schiffn. = Gymnocolea inflata (Huds.) Dumort.Lophozia alpestris (Schleich. ex F. Weber) A. Evans = Leiocolea collaris (Nees) SchljakovLophozia alpestris auct. non (Schleich. ex F. Weber) A. Evans = Lophozia sudetica (Nees exHuebener) GrolleLophozia attenuata (Mart.) Dumort. = Barbilophozia attenuata (Mart.) LoeskeLophozia atlantica (Kaal.) Schiffn. = Barbilophozia atlantica (Kaal.) Müll. Frib.Lophozia badensis (Gottsche) Schiffn. = Leiocolea badensis (Gottsche) Jörg.Lophozia bantriensis (Hook.) Steph. = Leiocolea bantriensis (Hook.) Jörg.Lophozia barbata (Schmidel ex Schreb.) Dumort. = Barbilophozia barbata (Schmidel exSchreb.) LoeskeLophozia baueriana (Schiffn.) Schiffn. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia canariensis Bryhn = Marsupella emarginata (Ehrh.) Dumort.Lophozia capitata subsp. laxa (Lindb.) Bisang = Lophozia laxa (Lindb.) GrolleLophozia collaris (Nees) Dumort. = Leiocolea collaris (Nees) SchljakovLophozia confertifolia Schiffn. = Lophozia ventricosa (Dicks.) Dumort. [38]Lophozia ehrhartiana (F. Weber) Inoue & Steere = Lophozia ventricosa (Dicks.) Dumort.Lophozia elongata Steph. REJECTED FROM THE AREALophozia excisa var. cylindracea (Dumort.) Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia excisa var. jurensis (Meyl. ex Müll. Frib.) Müll. Frib. = Lophozia excisa (Dicks.)Dumort.Lophozia exsecta (Schmidel ex Schrad.) Dumort. = Tritomaria exsecta (Schmidel exSchrad.) LoeskeLophozia exsectiformis (Breidl.) Jaap = Tritomaria exsectiformis (Breidl.) LoeskeLophozia floerkei (F. Weber & D. Mohr) Schiffn. = Barbilophozia floerkei (F. Weber & D.Mohr) LoeskeLophozia gillmanii (Austin) R. M. Schust. = Leiocolea gillmanii (Austin) A. EvansLophozia gracillima H. Buch = Lophozia ascendens (Warnst.) R. M. Schust.Lophozia gracilis Schleich. ex Steph. nom. illeg. = Barbilophozia attenuata (Mart.) LoeskeLophozia guttulata (Lindb. & Arnell) A. Evans = Lophozia longiflora (Nees) Schiffn.Lophozia hatcheri (A. Evans) Steph. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia heterocolpos (Thed. ex Hartm.) M. Howe = Leiocolea heterocolpos (Thed. exHartm.) H. BuchLophozia hornschuchiana (Nees) Macoun = Leiocolea bantriensis (Hook.) Jörg.Lophozia incisa subsp. opacifolia (Culm. ex Meyl.) R. M. Schust. & Damsh. = Lophoziaopacifolia Culm. ex Meyl.Lophozia inflata (Huds.) M. Howe = Gymnocolea inflata (Huds.) Dumort.Lophozia jurensis Meyl. ex Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia kaurinii (Limpr.) Steph. = Leiocolea gillmanii (Austin) A. EvansLophozia kiaeri Jörg. p. p. = Lophozia excisa (Dicks.) Dumort.Lophozia kunzeana (Huebener) A. Evans = Barbilophozia kunzeana (Huebener) Müll.Frib.Lophozia longiflora sensu auct. = Lophozia ventricosa (Dicks.) Dumort.Lophozia lycopodioides (Wallr.) Cogn. = Barbilophozia lycopodioides (Wallr.) LoeskeLophozia lycopodioides var. parvifolia Schiffn. = Barbilophozia hatcheri (A. Evans)LoeskeLophozia lyoni (Taylor) Steph. = Tritomaria quinquedentata (Huds.) H. Buch

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 409Lophozia minuta (Schreb.) Schiffn. = Anastrophyllum minutum (Schreb.) R. M. Schust.Lophozia muelleri (Nees ex Lindenb.) Dumort. = Leiocolea collaris (Nees) SchljakovLophozia porphyroleuca (Nees) Schiffn. nom. illeg. = Lophozia longiflora (Nees) Schiffn.Lophozia quadriloba (Lindb.) A. Evans = Barbilophozia quadriloba (Lindb.) LoeskeLophozia propagulifera (Gottsche) Steph. = Lophozia latifolia R. M. Schust. [36]Lophozia quinquedentata (Huds.) Cogn. = Tritomaria quinquedentata (Huds.) H. BuchLophozia rufescens Schljakov = Lophozia sudetica (Nees ex Huebener) GrolleLophozia schreberi Boulay nom. illeg. = Barbilophozia barbata (Schmidel ex Schreb.)LoeskeLophozia scutata (F. Weber & D. Mohr) Dumort. = Harpanthus scutatus (F. Weber & D.Mohr) SpruceLophozia turbinata (Raddi) Steph. = Leiocolea turbinata (Raddi) H. BuchLophozia turbinata var. wilsoniana (Nees) C. Massal. = Leiocolea turbinata (Raddi) H.BuchLophozia ventricosa var. longiflora (Nees)Macoun = Lophozia longiflora (Nees) Schiffn.Lophozia ventricosa var. longiflora auct. non (Nees) Macoun = Lophozia ventricosa(Dicks.) Dumort.Lophozia ventricosa var. porphyroleuca (Nees) Müll. Frib. = Lophozia longiflora (Nees)Schiffn.Lophozia ventricosa var. silvicola (H. Buch) E. W. Jones ex R. M. Schust. = Lophozia silvicolaH. BuchLophozia ventricosa var. uliginosa Breidl. ex Schiffn. = Lophozia ventricosa (Dicks.)Dumort.Lunularia alpina Nees = Sauteria alpina (Nees) NeesLunularia dillenii Le Jol. = Lunularia cruciata (L.) Lindb.Lunularia michelii Le Jol. = Lunularia cruciata (L.) Lindb.Lunularia vulgaris Raddi nom. illeg. = Lunularia cruciata (L.) Lindb.Madotheca baueri Schiffn. = Porella xbaueri (Schiffn.) C. E. O. JensenMadotheca canariensis (F. Weber) Nees = Porella canariensis (F. Weber) Underw.# Madotheca canariensis var. subsquarrosa Schiffn. = Porella canariensis (F. Weber)Underw.Madotheca cordaeana (Huebener) Dumort. = Porella cordaeana (Huebener) MooreMadotheca cordaeana var. distans (Müll. Frib.) Müll. Frib. = Porella cordaeana (Huebener)MooreMadotheca dentata (Hartm.) C.Massal. = Porella cordaeana (Huebener) MooreMadotheca inaequalis Gottsche ex Steph. = Porella inaequalis (Gottsche ex Steph.) Perss.Madotheca jackii Schiffn. = Porella platyphylla (L.) Pfeiff.Madotheca kabylica Trab.nom. inval. = Porella cordaeana (Huebener) MooreMadotheca laevigata (Schrad.) Dumort. = Porella arboris-vitae (With.) GrolleMadotheca laevigata var. obscura Nees = Porella arboris-vitae (With.) GrolleMadotheca laevigata var. killarniensis (Pearson) Macvicar = Porella arboris-vitae (With.)GrolleMadotheca laevigata var. subintegra (Kaal.) Schiffn. = Porella arboris-vitae (With.) Grolle☞ Madotheca laevigata var. thuja (Nees) Gottsche et al. = Porella arboris-vitae (With.)GrolleMadotheca levieri (J. B. Jack & Steph.) Copp. = Porella cordaeana (Huebener) MooreMadotheca obscura (Nees) Boulay = Porella arboris-vitae (With.) GrolleMadotheca obscura (Nees) Warnst. nom. inval. = Porella arboris-vitae (With.) GrolleMadotheca obtusata Taylor = Porella obtusata (Taylor) Trevis.

410 R.M. Ros et al.Madotheca platyphylla (L.) Dumort. = Porella platyphylla (L.) Pfeiff.Madotheca platyphylla var. subsquarrosa Schiffn. = Porella platyphylla (L.) Pfeiff.Madotheca platyphylloidea (Schwein.) Nees = Porella platyphylla (L.) Pfeiff.Madotheca porella Nees nom. illeg. = Porella pinnata L.Madotheca porelloides De Not. ex Lindb. nom. inval. = Porella platyphylla (L.) Pfeiff.Madotheca rivularis Nees = Porella cordaeana (Huebener) MooreMadotheca simplicior J. E. Zetterst. = Porella cordaeana (Huebener) MooreMadotheca thuja (Hook.) Dumort. nom. illeg. = Porella arboris-vitae (With.) GrolleMadotheca thuja auct. non (Hook.) Dumort.=Porella obtusata (Taylor) Trevis.Madotheca torva De Not. ex Bertol. nom. inval. = Porella obtusata (Taylor) Trevis.Mannia atlantica Jelenc nom. inval. [39]Mannia dichotoma Udar & V. Chandra nom. inval. = Mannia androgyna (L.) A. EvansMannia dichotoma S. W. Arnell = Mannia androgyna (L.) A. EvansMannia rupestris (Nees) Frye & L. Clark = Mannia triandra (Scop.) GrolleMarchantia alpestris (Nees) Burgeff = Marchantia polymorpha subsp. montivagans Bischl.& BoisselierMarchantia androgyna L. = Mannia androgyna (L.) A. EvansMarchantia angustifolia Neck. nom. illeg. = Mannia androgyna (L.) A. EvansMarchantia aquatica (Nees) Burgeff = Marchantia polymorpha L. subsp. polymorphaMarchantia circumscissa Biv. = Mannia androgyna (L.) A. EvansMarchantia commutata Lindenb. = Preissia quadrata (Scop.) Müll. Frib.Marchantia conica L. = Conocephalum conicum (L.) Dumort.Marchantia cruciata L. = Lunularia cruciata (L.) Lindb.Marchantia cruciata var. hyalina Sommerf. = Athalamia hyalina (Sommerf.) S. Hatt.$ Marchantia dillenii = Lunularia cruciata (L.) Lindb.Marchantia fragrans Balbis = Mannia fragrans (Balbis) Frye & L. Clark [10]Marchantia glauca Biv. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Marchantia gracilis F. Weber = Asterella gracilis (F. Weber) Underw.Marchantia hemisphaerica L. = Reboulia hemisphaerica (L.) RaddiMarchantia hirsuta Sw. = Dumortiera hirsuta (Sw.) NeesMarchantia hyalina Sommerf. = Athalamia hyalina (Sommerf.) S. Hatt.Marchantia irrigua Wilson ex Hook. = Dumortiera hirsuta (Sw.) NeesMarchantia pilosa Hornem. = Mannia pilosa (Hornem.) Frye & L. ClarkMarchantia polycephala Schleich. nom. inval. = Asterella gracilis (F. Weber) Underw.Marchantia polymorpha auct . = Marchantia polymorpha subsp. ruderalis Bischl. & BoisselierMarchantia polymorpha var. alpestris (Nees) Nees = Marchantia polymorpha subsp. montivagansBischl. & BoisselierMarchantia polymorpha var . aquatica (Nees) Gottsche et al. = Marchantia polymorpha L.subsp. polymorphaMarchantia polymorpha f. domestica Nees nom. illeg. = Marchantia polymorpha subsp.ruderalis Bischl. & BoisselierMarchantia polymorpha var. mamillata I. Hag . ex Schiffn. = Marchantia polymorphasubsp. montivagans Bischl. & BoisselierMarchantia quadrata Scop. = Preissia quadrata (Scop.) Müll. Frib. [51]Marchantia spathysii Lindenb. = Athalamia spathysii (Lindenb.) S. Hatt.Marchantia stellata Scop. = Marchantia polymorpha subsp. ruderalis Bischl. & BoisselierMarchantia triandra Scop. = Mannia triandra (Scop.) Grolle

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 411Marchantia umbellata Scop. = Marchantia polymorpha subsp. ruderalis Bischl. & BoisselierMarchantia vittata Raddi = Marchantia polymorpha L.subsp. polymorphaMarchesinia johnsoniana (Mitt.) Herz. = Acanthocoleus aberrans (Lindenb. & Gottsche)Kruijt☞ Marsilia epiphylla (L.) Lindb. nom. illeg. = Pellia epiphylla (L.) CordaMarsupella aemula (Limpr.) Lindb. ex Kaal. = Marsupella condensata (Ångstr. ex C.Hartm.) Kaal.Marsupella aquatica (Lindenb.) Schiffn. = Marsupella emarginata (Ehrh.) Dumort.Marsupella badensis Schiffn. = Marsupella funckii (F. Weber & D. Mohr) Dumort.Marsupella capillaris (Limpr.) H. Bern. = Marsupella boeckii (Austin) Lindb.Marsupella conferta (Limpr.) Spruce = Marsupella brevissima (Dumort.) GrolleMarsupella emarginata subsp. aquatica (Lindenb.) Meyl. = Marsupella emarginata (Ehrh.)Dumort.Marsupella emarginata var. aquatica (Lindenb.) Dumort. = Marsupella emarginata (Ehrh.)Dumort.Marsupella emarginata var. minor C. Massal. ex Schiffn. nom. illeg. = Marsupella emarginata(Ehrh.) Dumort.Marsupella erythrorhiza Schiffn. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort.☞ Marsupella filiformis (Lindb.) Lindb. ex Kaal. = Marsupella boeckii (Austin) Lindb.Marsupella funckii var. major (Nees) Boulay = Marsupella funckii (F. Weber & D. Mohr)Dumort.☞ Marsupella gracilis (C. Massal. & Carestia) Pearson = Marsupella sprucei (Limpr.) BernetMarsupella hungarica Boros & Vajda nom. inval. = Marsupella funckii (F. Weber & D.Mohr) Dumort.Marsupella joergensenii Schiffn. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort.Marsupella latifolia Lindb. nom. inval. = Marsupella boeckii (Austin) Lindb.☞ Marsupella neglecta (Limpr.) Lindb. = Marsupella sprucei (Limpr.) BernetMarsupella nevicensis (Carrington) Pearson = Marsupella boeckii (Austin) Lindb.Marsupella pearsonii Schiffn. ex Macvicar = Marsupella emarginata (Ehrh.) Dumort.Marsupella pygmaea (Limpr.) Steph. = Marsupella funckii (F. Weber & D. Mohr) Dumort.Marsupella revoluta (Nees) Dumort. = Apomarsupella revoluta (Nees) R. M. Schust.Marsupella robusta (De Not.) A. Evans = Marsupella emarginata (Ehrh.) Dumort.Marsupella styriaca (Limpr.) Kaal. = Marsupella sparsifolia (Lindb.) Dumort.Marsupella sullivantii (De Not.) A. Evans = Marsupella sphacelata (Gieseke ex Lindenb.)Dumort.Marsupella ustulata (Huebener) Spruce ex Pearson hom. illeg. = Marsupella emarginata(Ehrh.) Dumort.Marsupella ustulata Spruce non (Huebener) Spruce ex Pearson hom. illeg. = Marsupellasprucei (Limpr.) BernetMarsupella ustulata var. neglecta (Limpr.) Müll. Frib. = Marsupella sprucei (Limpr.) BernetMarsupella varians (Lindb.) Müll. Frib. = Marsupella brevissima (Dumort.) GrolleMartinellia aequiloba (Schwägr.) Lindb. nom. illeg. = Scapania aequiloba (Schwägr.)Dumort.☞ Martinellia apiculata (Spruce) Trevis. = Scapania apiculata Spruce☞ Martinellia aspera (Bernet & M. Bernet) Broth. = Scapania aspera Bernet & M. BernetMartinellia calcicola Arnell & J. Perss. = Scapania calcicola (Arnell & J. Perss.) InghamMartinellia convexa (Scop.) Lindb. = Scapania umbrosa (Schrad.) Dumort.Martinellia crassiretis (Bryhn) Arnell = Scapania crassiretis BryhnMartinellia curta (Mart.) Lindb. = Scapania curta (Mart.) Dumort.

412 R.M. Ros et al.Martinellia cuspiduligera (Nees) Arnell = Scapania cuspiduligera (Nees) Müll. Frib.Martinellia gracilis (Lindb.) Lindb. = Scapania gracilis Lindb.☞ Martinellia helvetica (Gottsche) Trevis. = Scapania helvetica GottscheMartinellia mucronata (H. Buch) Arnell & C. E. O. Jensen = Scapania mucronata H. BuchMartinellia nemorosa Lindb. nom. illeg. = Scapania nemorea (L.) GrolleMartinellia obliqua Arnell = Scapania uliginosa (Sw. ex Lindenb.) Dumort.Martinellia obscura Arnell & C. E. O. Jensen = Scapania obscura (Arnell & C. E. O.Jensen) Schiffn.Martinellia paludicola (Loeske & Müll. Frib.) C. E. O. Jensen = Scapania paludicolaLoeske & Müll. Frib.☞ Martinellia paludosa (Müll. Frib.) Arnell & C. E. O. Jensen = Scapania paludosa (Müll.Frib.) Müll. Frib.Martinellia resupinata (L.) Gray ex Lindb. = Scapania undulata (L.) Dumort.☞ Martinellia resupinata (Hook.) Lindb. nom. inval. = Scapania compacta (A. Roth)Dumort.Martinellia rosacea (Corda) Lindb. = Scapania curta (Mart.) Dumort.Martinellia scandica Arnell & H. Buch = Scapania scandica (Arnell & H. Buch) Macvicar☞ Martinellia subalpina (Nees ex Lindenb.) Trevis. = Scapania subalpina (Nees exLindenb.) Dumort.☞ Martinellia uliginosa (Sw. ex Lindenb.) Trevis. = Scapania uliginosa (Sw. ex Lindenb.)Dumort.☞ Martinellia umbrosa (Schrad.) Gray = Scapania umbrosa (Schrad.) Dumort.Martinellia undulata (L.) Gray = Scapania undulata (L.) Dumort.Massula capitata (Hook.) Schljakov = Lophozia capitata (Hook.) MacounMassula grandiretis (Lindb. ex Kaal.) Schljakov = Lophozia grandiretis (Lindb. ex Kaal.)Schiffn.Massula incisa (Schrad.) Schljakov nom. illeg.= Lophozia incisa (Schrad.) Dumort.Massula opacifolia (Culm. ex Meyl.) Schljakov nom. illeg. = Lophozia opacifolia Culm. exMeyl.Mastigobryum deflexum (Mart.) Nees = Bazzania tricrenata (Wahlenb.) Lindb.Mastigobryum triangulare Schleich. ex Steph. nom. illeg. = Bazzania tricrenata (Wahlenb.)Lindb.Mastigobryum trilobatum (L.) Nees = Bazzania trilobata (L.) GrayMesophylla crenulata (Mitt.) Corb. = Jungermannia gracillima Sm.Mesophylla crenulata var. gracillima (Sm.) Boulay nom. illeg. = Jungermannia gracillimaSm.Mesophylla hyalina (Lyell) Corb. = Jungermannia hyalina LyellMesophylla hyalina var. gracillima (Schiffn.) Boulay = Jungermannia hyalina LyellMesophylla nigrella (De Not.) Boulay = Southbya nigrella (De Not.) Henriq.Mesophylla obovata (Nees) Corb. = Jungermannia obovata NeesMesophylla rotaeana (De Not) Dumort. = Nardia scalaris GrayMesophylla scalaris (Gray) Dumort. = Nardia scalaris GrayMesophylla stillicidiorum (Raddi) Boulay = Southbya tophacea (Spruce) SpruceMetzgeria conjugata subsp. simplex (Lorb. ex Müll. Frib.) R. M. Schust. = Metzgeria simplexLorb. ex Müll. Frib.# Metzgeria conjugata f. subpinnata Jörg . = Metzgeria conjugata Lindb.Metzgeria densiseta Steph. = Metzgeria furcata (L.) Dumort.Metzgeria fruticulosa auct. eur. non (O. F. Müll.) Evans = Metzgeria violacea (Ach.)Dumort. [40]

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 413Metzgeria furcata var. fruticulosa auct. eur. non (O. F. Müll.) Lindb. = Metzgeria violacea(Ach.) Dumort. [40]☞ Metzgeria furcata var. ulvula (Nees) Gottsche et al. = Metzgeria furcata (L.) Dumort.Metzgeria glabra Raddi nom. illeg. = Metzgeria furcata (L.) Dumort.Metzgeria glabra f. ulvula (Nees) Zodda = Metzgeria furcata (L.) Dumort.Metzgeria hamata Lindb. nom. illeg. = Metzgeria leptoneura SpruceMetzgeria nitida Mitt. = Metzgeria leptoneura SpruceMetzgeria pubescens (Schrank) Raddi = Apometzgeria pubescens (Schrank) Kuwah.Microlejeunea diversiloba auct. non (Spruce) Müll. Frib. = Lejeunea hibernica Bischl. et al.ex GrolleMicrolejeunea mandonii Steph. = Lejeunea mandonii (Steph.) Müll. Frib.Microlepidozia setacea auct. non (G. Web.) Jørg. = Kurzia pauciflora (Dicks.) GrolleMicrolepidozia sylvatica (A. Evans) Jørg. = Kurzia sylvatica (A. Evans) GrolleMicrolepidozia trichoclados (Müll. Frib.) Jørg. = Kurzia trichoclados (Müll. Frib.) GrolleMnium fissum L. = Calypogeia fissa (L.) RaddiMnium trichomanis L. = Calypogeia fissa (L.) RaddiMnium trichomanis auct. non L. = Calypogeia azurea Stotler & Crotz [11]Moerckia flotoviana (Nees) Schiffn. = Moerckia hibernica (Hook.) GottscheMylia cuneifolia (Hook.) Gray = Leptoscyphus cuneifolius (Hook.) Mitt.Nardia alpina (Gottsche ex Husn.) Trevis. nom. inval. = Marsupella alpina (Gottsche exHusn.) BernetNardia boeckii (Austin) Lindb. = Marsupella boeckii (Austin) Lindb.Nardia brevissima (Dumort.) Lindb. = Marsupella brevissima (Dumort.) GrolleNardia cochlearis Lindb. = Marsupella brevissima (Dumort.) GrolleNardia commutata (Limpr.) C. Massal. & Carestia = Marsupella commutata (Limpr.)BernetNardia compressa var. rigida Lindb. = Nardia compressa (Hook.) GrayNardia crassula Lorb. nom. inval. = Nardia insecta Lindb.Nardia crenulata (Mitt.) Lindb. = Jungermannia gracillima Sm.Nardia crenulata var. cristulata (Dumort.) Schiffn. = Jungermannia gracillima Sm.Nardia crenulata var. gracillima (Sm.) Lindb. = Jungermannia gracillima Sm.Nardia crenulata var. inundata Schiffn. = Jungermannia gracillima Sm.☞ Nardia densifolia (Nees) Trevis. = Marsupella emarginata (Ehrh.) Dumort.Nardia emarginata (Ehrh.) Gray = Marsupella emarginata (Ehrh.) Dumort.Nardia emarginata var. aquatica (Lindenb.) Carrington = Marsupella emarginata (Ehrh.)Dumort.Nardia filiformis Lindb. = Marsupella boeckii (Austin) Lindb.☞ Nardia funckii (F. Weber & D. Mohr) Carrington ex Lindb. = Marsupella funckii(F. Weber & D. Mohr) Dumort.Nardia geoscyphus var. insecta (Lindb.) Schiffn. = Nardia insecta Lindb.Nardia gracilis C. Massal. & Carestia = Marsupella sprucei (Limpr.) BernetNardia gracillima (Sm.) Lindb. = Jungermannia gracillima Sm.Nardia haematosticta Nees ex Lindb. nom. inval. = Nardia geoscyphus (De Not.) Lindb.Nardia hyalina (Lyell) Carrington = Jungermannia hyalina LyellNardia hyalina var. colorata Samp. nom. inval. [30]Nardia latifolia Lindb. nom. inval. = Marsupella boeckii (Austin) Lindb.Nardia levieri Steph. = Jungermannia confertissima NeesNardia lignicola Schiffn. = Jungermannia lignicola (Schiffn.) Grolle

414 R.M. Ros et al.Nardia muelleriana Schiffn. = Jungermannia hyalina LyellNardia obovata (Nees) Lindb. = Jungermannia obovata NeesNardia obovata var. minor Carrington = Jungermannia subelliptica (Lindb. ex Kaal.)LevierNardia pachyphylla (De Not.) Trevis. = Nardia compressa (Hook.) GrayNardia revoluta (Nees) Lindb. = Apomarsupella revoluta (Nees) R. M. Schust.☞ Nardia robusta (De Not.) Lindb. ex Carrington = Marsupella emarginata (Ehrh.)Dumort.☞ Nardia silvrettae (Gottsche ex Dumort.) Trevis. [41]Nardia sparsifolia (Lindb.) Lindb. = Marsupella sparsifolia (Lindb.) Dumort.Nardia sparsifolia var. minor (Schleich.) Lindb. nom. inval. = Marsupella brevissima(Dumort.) GrolleNardia sphacelata (Gieseke ex Lindenb.) Carrington = Marsupella sphacelata (Gieseke exLindenb.) Dumort.Nardia sprucei (Limpr.) C. Massal. & Carestia = Marsupella sprucei (Limpr.) BernetNardia subelliptica Lindb. ex Kaal. = Jungermannia subelliptica (Lindb. ex Kaal.) LevierNardia subtilissima Schiffn. = Jungermannia subtilissima (Schiffn.) GrolleNardia varians Lindb. = Marsupella brevissima (Dumort.) GrolleNeesiella carnica (C. Massal.) Schiffn. = Mannia pilosa (Hornem.) Frye & L. ClarkNeesiella pilosa (Hornem.) Schiffn. = Mannia pilosa (Hornem.) Frye & L. ClarkNeesiella rupestris (Nees) Schiffn. = Mannia triandra (Scop.) GrolleObtusifolium obtusum (Lindb.) S. W. Arnell = Lophozia obtusa (Lindb.) A. EvansOdontoschisma cavifolium Steph. = Odontoschisma denudatum (Mart.) Dumort.Odontoschisma denudatum subsp. elongatum (Lindb.) Potemkin = Odontoschisma elongatum(Lindb.) A. EvansOdontoschisma denudatum var. elongatum (Lindb.) Lindb. = Odontoschisma elongatum(Lindb.) A. EvansOdontoschisma denudatum f. elongatum Lindb. = Odontoschisma elongatum (Lindb.) A.EvansOdontoschisma remotifolium Warnst. = Odontoschisma sphagni (Dicks.) Dumort.Odontoschisma sphagni var. denudatum (Mart.) C. Massal. & Carestia = Odontoschismadenudatum (Mart.) Dumort.Odontoschisma sphagni var. macrior Bernet = Odontoschisma denudatum (Mart.) Dumort.Odontoschisma tessellatum (Berggr.) C. E. O. Jensen = Odontoschisma macounii (Austin)Underw.Orthocaulis atlanticus (Kaal.) H. Buch = Barbilophozia atlantica (Kaal.) Müll. Frib.Orthocaulis attenuatus (Mart.) A. Evans = Barbilophozia attenuata (Mart.) LoeskeOrthocaulis floerkei (F. Weber & D. Mohr) H. Buch = Barbilophozia floerkei (F. Weber &D. Mohr) LoeskeOrthocaulis gracilis (Schleich. ex DC.) H. Buch = Barbilophozia attenuata (Mart.) LoeskeOrthocaulis kunzeanus (Huebener) H. Buch = Barbilophozia kunzeana (Huebener) Müll.Frib.Otiona crinita (Michx.) Corda = Reboulia hemisphaerica (L.) RaddiOtiona italica (Sassi) Dumort. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Otiona rupestris (J. R. Forst. & G. Forst.) Dumort. = Plagiochasma rupestre (J. R. Forst. &G. Forst.) Steph.Oxymitra paleacea Bisch. ex Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-SimOxymitra pyramidata (Raddi) Huebener non (Willd.) Huebener = Oxymitra incrassata(Brot.) Sérgio & Sim-Sim

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 415Oxymitra pyramidata var. linkii (Lindenb.) Lindenb. = Oxymitra incrassata (Brot.) Sérgio& Sim-SimOxymitra pyramidata var. paleacea (Bisch. ex Lindenb.) Gottsche et al. = Oxymitraincrassata (Brot.) Sérgio & Sim-SimPallavicinia blyttii (Moerch ex Hornem.) Lindb. = Moerckia blyttii (Moerch ex Hornem.)Brockm.Pallavicinia flotoviana (flotowii error pro flotoviana) (Nees) Lindb. = Moerckia hibernica(Hook.) GottschePallavicinia flotoviana var. hibernica (Hook.) Lindb. nom. illeg. = Moerckia hibernica(Hook.) GottschePallavicinia hibernica (Hook.) Gray = Moerckia hibernica (Hook.) GottschePedinophyllum interruptum var. pyrenaicum (Spruce) Kaal. = Pedinophyllum interruptum(Nees) Kaal.Pedinophyllum pyrenaicum (Spruce) Lindb. = Pedinophyllum interruptum (Nees) Kaal.Pedinophyllum pyrenaicum var. interruptum (Nees) Schiffn. nom. incorrect. = Pedinophylluminterruptum (Nees) Kaal.Pellia borealis Lorb. = Pellia epiphylla (L.) CordaPellia calycina (Taylor) Nees = Pellia endiviifolia (Dicks.) Dumort.Pellia endiviifolia f. furcigera (Hook.) Schiffn. = Pellia endiviifolia (Dicks.) Dumort.Pellia epiphylla subsp. borealis (Lorb.) Messe = Pellia epiphylla (L.) CordaPellia epiphylla f. neesiana Gottsche = Pellia neesiana (Gottsche) Limpr.Pellia fabbroniana auct. non Raddi = Pellia endiviifolia (Dicks.) Dumort.$ Pellia fabbroniana var. furcigera = Pellia endiviifolia (Dicks.) Dumort.Pellia fabbroniana f. furcigera (Hook.) C. Massal. = Pellia endiviifolia (Dicks.) Dumort.Pellia fabbroniana f. undulata Jelenc nom. inval. [42]Pellia fuciformis (Bory) Nees [43]Peltolepis grandis (Lindb.) Lindb. = Peltolepis quadrata (Saut.) Müll. Frib.Peltolepis sibirica Lindb. = Peltolepis quadrata (Saut.) Müll. Frib.Petalophyllum lamellatum (Nees) Lindb. = Petalophyllum ralfsii (Wilson) Nees & GottschePhragmicoma mackaii (Hook.) Dumort. = Marchesinia mackaii (Hook.) GrayPhysocolea rossettiana (C. Massal.) Steph. = Cololejeunea rossettiana (C. Massal.) Schiffn.Plagiochasma algericum Steph. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Plagiochasma aitonia Lindenb. & Nees = Plagiochasma rupestre (J. R. Forst. & G. Forst.)Steph.Plagiochasma beccarianum Steph. REJECTED FROM THE AREAPlagiochasma italicum (Sassi) De Not. = Plagiochasma rupestre (J. R. Forst. & G. Forst.)Steph.Plagiochasma roussellianum Mont. = Athalamia spathysii (Lindenb.) S. Hatt.Plagiochasma sauteria Gottsche ex Jelenc nom. inval. = Athalamia spathysii (Lindenb.)S. Hatt.Plagiochila aequiloba (Schwägr.) Mont. & Nees = Scapania aequiloba (Schwägr.) Dumort.Plagiochila africana Steph. REJECTED FROM THE AREA [44]Plagiochila asplenioides auct. = Plagiochila porelloides (Torrey ex Nees) Lindenb.Plagiochila asplenioides var. humilis (Nees) Lindenb. = Plagiochila porelloides (Torrey exNees) Lindenb.Plagiochila asplenioides var. major Nees = Plagiochila asplenioides (L. emend. Taylor)Dumort.Plagiochila asplenioides var. minor Lindenb. = Plagiochila porelloides (Torrey ex Nees)Lindenb.

416 R.M. Ros et al.Plagiochila asplenioides subsp . porelloides (Torrey ex Nees)Kaal. = Plagiochila porelloides(Torrey ex Nees) Lindenb.Plagiochila asplenioides var. riparia Breidl. = Plagiochila porelloides (Torrey ex Nees)Lindenb.Plagiochila atlantica F. Rose = Plagiochila heterophylla Lindenb. ex Lehm. [46]Plagiochila compacta (A. Roth) Mont. & Nees = Scapania compacta (A. Roth) Dumort.Plagiochila corniculata auct. non (Dumort.) Dumort. = Plagiochila exigua (Taylor) TaylorPlagiochila curta (Mart.) Mont. & Nees = Scapania curta (Mart.) Dumort.Plagiochila diacantha Taylor ex Steph. = Plagiochila bifaria (Sw.) Lindenb.Plagiochila dubia Lindenb. & Gottsche = Plagiochila patula (Sw.) Lindenb. REJECTEDFROM THE AREAPlagiochila interrupta (Nees) Mont. & Nees = Pedinophyllum interruptum (Nees) Kaal.Plagiochila javanica (Sw.) Nees & Mont. REJECTED FROM THE AREAPlagiochila killarniensis Pearson = Plagiochila bifaria (Sw.) Lindenb.Plagiochila major (Nees) S. W. Arnell nom. illeg. = Plagiochila asplenioides (L. emend.Taylor) Dumort.Plagiochila punctata var. owenii (Steph.) Macvicar = Plagiochila punctata (Taylor) TaylorPlagiochila pyrenaica Spruce = Pedinophyllum interruptum (Nees) Kaal.Plagiochila sharpii H. L. Blomq. = Plagiochila retrorsa GottschePlagiochila spinulosa var. inermis Carrington = Plagiochila bifaria (Sw.) Lindenb.Plagiochila spinulosa var. killarniensis (Pearson) Macvicar = Plagiochila bifaria (Sw.)Lindenb.Plagiochila spinulosa var. macaronesiana Allorge & V. Allorge = Plagiochila bifaria (Sw.)Lindenb.Plagiochila spinulosa var. punctata (Taylor) Carrington = Plagiochila punctata (Taylor)TaylorPlagiochila spinulosa var. tridenticulata auct. non (Michx.) Husn. nom. illeg. = Plagiochilaexigua (Taylor) TaylorPlagiochila stableri Pearson = Plagiochila exigua (Taylor) TaylorPlagiochila tridenticulata auct. non (Michx.) Dumort. = Plagiochila exigua (Taylor) TaylorPlagiochila undulata (L.) Mont. & Nees = Scapania undulata (L.) Dumort.Plectocolea crenulata (Mitt.) A. Evans = Jungermannia gracillima Sm.Plectocolea crenulata var. gracillima (Sm.) Frye & L. Clark = Jungermannia gracillima Sm.Plectocolea hyalina (Lyell) Mitt. = Jungermannia hyalina LyellPlectocolea obovata (Nees) Lindb. = Jungermannia obovata NeesPlectocolea subelliptica (Lindb. ex Kaal.) A. Evans = Jungermannia subelliptica (Lindb. exKaal.) LevierPleuroclada albescens (Hook.) Spruce nom. illeg. = Pleurocladula albescens (Hook.) GrollePleuroschisma flaccidum Dumort. = Bazzania flaccida (Dumort.) GrollePleuroschisma implexum (Nees) Meyl. = Bazzania flaccida (Dumort.) GrollePleuroschisma sphagni (Dicks.) Dumort. = Odontoschisma sphagni (Dicks.) Dumort.$ Pleuroschisma triangulare var. implexa = Bazzania flaccida (Dumort.) GrollePleuroschisma tricrenatum (Wahlenb.) Dumort. = Bazzania tricrenata (Wahlenb.) Lindb.Pleuroschisma tricrenatum var. implexum (Nees) Müll. Frib. = Bazzania flaccida (Dumort.)GrollePleuroschisma trilobatum (L.) Dumort. = Bazzania trilobata (L.) GrayPorella arboris-vitae var. killarniensis (Pearson) M. F. V. Corley = Porella arboris-vitae(With.) Grolle

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 417Porella arboris-vitae var. obscura (Nees) M.F.V. Corley = Porella arboris-vitae (With.)GrollePorella cordaeana var. distans (Müll. Frib.) Jelenc = Porella cordaeana (Huebener) MoorePorella cordaeana var. simplicior (J.E. Zetterst.) Arnell = Porella cordaeana (Huebener)MoorePorella dentata ( Hartm.) Lindb. = Porella cordaeana (Huebener) MoorePorella kabylica Jelenc nom. inval. = Porella cordaeana (Huebener) MoorePorella laevigata (Schrad.) Pfeiff. = Porella arboris-vitae (With.) GrollePorella laevigata var. obscura (Nees) Arnell = Porella arboris-vitae (With.) Grolle☞ Porella laevigata var. thuja (Nees)Arnell = Porella arboris-vitae (With.) GrollePorella levieri J. B. Jack & Steph. = Porella cordaeana (Huebener) MoorePorella notarisii Trevis. nom. inval. = Porella platyphylla (L.) Pfeiff.Porella platyphylla var. subsquarrosula (Schiffn.) Arnell = Porella platyphylla (L.) Pfeiff.Porella platyphylloidea (Schwein.) Lindb. = Porella platyphylla (L.) Pfeiff.? Porella porelloides De Not . = Porella platyphylla (L.) Pfeiff.Porella rivularis (Nees) Pfeiff. = Porella cordaeana (Huebener) MoorePorella thuja (Dicks.) Lindb. nom. illeg. = Porella arboris-vitae (With.) GrollePorella thuja auct. non (Dicks.) Lindb. = Porella obtusata (Taylor) Trevis.Preissia commutata (Lindenb.) Nees = Preissia quadrata (Scop.) Müll. Frib.Preissia cucullata Nees & Mont. = Lunularia cruciata (L.) Lindb.☞ Preissia hemisphaerica Cogn. = Preissia quadrata (Scop.) Müll. Frib.Preissia italica Corda = Preissia quadrata (Scop.) Müll. Frib.Prionolobus turneri (Hook.) Schiffn. = Cephaloziella turneri (Hook.) Müll. Frib.☞ Ptilidium ciliare var. pulcherrimum (Weber) Warnst. = Ptilidium pulcherrimum(Weber) Vain.Ptilidium trichophyllum (L.) Mitt. = Blepharostoma trichophyllum (L.) Dumort.Radula bornmuelleri Schiffn. = Radula lindenbergiana Gottsche ex C. Hartm.Radula commutata Gottsche ex J. B. Jack = Radula lindenbergiana Gottsche ex C. Hartm.Radula complanata subsp. lindbergiana (Gottsche ex C. Hartm.) R. M. Schust. = Radulalindenbergiana Gottsche ex C. Hartm.Radula complanata f. propagulifera Nees = Radula lindenbergiana Gottsche ex C. Hartm.Radula complanata var. propagulifera (Nees) Gottsche et al. = Radula lindenbergianaGottsche ex C. Hartm.Radula germana J. B. Jack = Radula lindenbergiana Gottsche ex C. Hartm.Radula limbata Schiffn. = Radula wichurae Steph.Radula lindbergiana Gottsche ex Jack nom. inval. = Radula lindenbergiana Gottsche ex C.Hartm.Radula lindbergii Gottsche ex Lindb. nom. inval. = Radula lindenbergiana Gottsche ex C.Hartm.Radula notarisii Steph. = Radula complanata (L.) Dumort.Radula ovata J. B. Jack ex Steph. = Radulalindenbergiana Gottsche ex C. Hartm.Radula physoloba Mont. REJECTED FROM THE AREA [52]Reboulia charrieri Douin & R.C.V. Douin = Reboulia hemisphaerica (L.) Raddi☞ Reboulia hemisphaerica var. macrocephala C. Massal. = Reboulia hemisphaerica (L.)RaddiReboulia hemisphaerica var. microspora Schiffn. = Reboulia hemisphaerica (L.) RaddiReboulia hemisphaerica var. pangiensis (Kashyap) Kashyap = Reboulia hemisphaerica (L.)Raddi

418 R.M. Ros et al.Reboulia hemisphaerica subsp. paradoxa R. M. Schust. nom. inval. = Reboulia hemisphaerica(L.) RaddiReboulia occidentalis Douin & R. C. V. Douin = Reboulia hemisphaerica (L.) RaddiReboulia quadrata (Scop.) Bertol. = Preissia quadrata (Scop.) Nees [51]Rhakiocarpon conspersum Corda = Reboulia hemisphaerica (L.) RaddiRiccardia latifrons var. sinuata (Dicks. ex Hook.) Lindb. = Riccardia chamedryfolia (With.)GrolleRiccardia major Lindb. = Riccardia chamedryfolia (With.) GrolleRiccardia pinguis (L.) Gray = Aneura pinguis (L.) Dumort.Riccardia pinguis f. angustior (Hook.) P. Allorge = Aneura pinguis (L.) Dumort. Riccardia pinguis f . submersa Loeske ex Jaapnom. inval. = Aneura pinguis (L.) Dumort.Riccardia sinuata (Dicks. ex Hook.) Trevis. = Riccardia chamedryfolia (With.) GrolleRiccia aculeata Trab.nom. nud. = Riccia crozalsii Levier [55]Riccia aegyptiaca S. W. Arnell = Riccia congoana Steph.Riccia affinis Milde = Riccia ciliata Hoffm.Riccia aggregata Underw. = Riccia nigrella DC.Riccia americana (M. Howe) M. Howe = Riccia lamellosa RaddiRiccia armatissima (Levier ex Müll. Frib.) Trab. = Riccia gougetiana var. armatissimaLevier ex Müll. Frib.Riccia arvensis Austin = Riccia bifurca Hoffm.Riccia atromarginata var. glabra Levier ex Müll. Frib. = Riccia trabutiana Steph.Riccia austinii Steph. = Riccia lamellosa RaddiRiccia battandieri Trab.nom. nud. [55]Riccia baumgartneri Schiffn. = Riccia subbifurca Warnst. ex Croz.Riccia bavarica Warnst. = Riccia warnstorfii Limpr. ex Warnst.Riccia beckeriana Steph. = Riccia frostii AustinRiccia bifurca f. subinermis (Heeg) Müll. Frib. = Riccia bifurca var. subinermis HeegRiccia bischoffii Huebener = Riccia ciliifera Link ex Lindenb.Riccia bischoffii f. armata Trab. nom. nud . = Riccia gougetiana var. armatissima Levier exMüll. Frib. [55]Riccia bischoffii var. ciliifera (Link. ex Lindenb.) Müll. Frib. = Riccia ciliifera Link exLindenb.Riccia bischoffii var. fimbriata Gottsche et al. = Riccia ciliifera Link ex Lindenb.Riccia bischoffii f. maxima Levier = Riccia gougetiana Durieu & Mont.Riccia bischoffii var. subtumida Milde = Riccia ciliata Hoffm.Riccia bullosa Link ex Lindenb. REJECTED FROM THE AREARiccia bullosa sensu Müll. Frib. = Exormotheca welwitschii Steph.Riccia cana Durieu ex Trab. nom. illeg. = Riccia crinita TaylorRiccia canescens Steph. = Riccia crinita TaylorRiccia capensis Steph. = Riccia limbata Bisch. ex Krauss REJECTED FROM THE AREARiccia capillata Schmidel nom. illeg. = Ricciocarpos natans (L.) CordaRiccia centrifuga Arnell = Riccia fluitans L.Riccia chinensis Herz. [54]Riccia chudoana Steph. [54]Riccia ciliata Raddi nom. illeg. = Riccia michelii RaddiRiccia ciliata var. intumescens Bisch. = Riccia ciliata Hoffm. [57]Riccia ciliata var. epilosa Warnst. = Riccia ciliata Hoffm.Riccia ciliifera f. pedemontana (Steph.) Müll. Frib. = Riccia ciliifera Link ex Lindenb.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 419Riccia ciliata var. violacea Kny = Riccia ciliata Hoffm.Riccia commutata J. B. Jack ex Levier = Riccia warnstorfii Limpr. ex Warnst.Riccia commutata var. acrotricha Levier ex Sommier = Riccia warnstorfii var. subinermisWarnst.Riccia concava Bisch. REJECTED FROM THE AREARiccia convexa Steph. [54]Riccia coriandrina Spreng. = Corsinia coriandrina (Spreng.) Lindb. Riccia crozalsii f. major C. Massal. nom. nud. = Riccia crozalsii LevierRiccia crystallina auct. = Riccia cavernosa Hoffm.Riccia crystallina var. angustior Lindenb. ex Nees = Riccia cavernosa Hoffm.Riccia crystallina subsp. austrigena Schiffn. = Riccia crystallina L. emend. RaddiRiccia crystallina var. major Lindb. = Riccia crystallina L. emend. RaddiRiccia crystallina var. vulgaris Lindenb. = Riccia crystallina L. emend. RaddiRiccia dufourii Nees = Riccia lamellosa RaddiRiccia echinulata Trab. nom. nud. = Riccia ciliifera Link ex Lindenb. [55]Riccia epicarpa Wallr. ex Gottsche et al. = Riccia sorocarpa Bisch.Riccia erinacea Schiffn. = Riccia gougetiana var. armatissima Levier ex Müll. Frib.Riccia eudichotoma Bisch. nom. illeg. = Riccia canaliculata Hoffm.Riccia eudichotoma var. fluitans (L.) Bisch. nom. incorrect. & nom. illeg. = Riccia fluitans L.Riccia fluitans var. canaliculata (Hoffm.) A. Roth = Riccia canaliculata Hoffm.Riccia fluitans var. purpurascens Klinggr . = Riccia huebeneriana Lindenb.Riccia frostii var . crystallinoides Schiffn. [58]Riccia fruticulosa auct. eur. non O. F. Müll. = Metzgeria violacea (Ach.) Dumort. [40]Riccia glauca var . subinermis (Lindb.) Warnst. = Riccia glauca var. ciliaris Warnst.Riccia glaucescens Carrington = Riccia beyrichiana Hampe ex Lehm.Riccia gougetiana var. erinacea Schiffn. nom. inval. = Riccia gougetiana var. armatissimaLevier ex Müll. Frib.Riccia henriquesii Levier = Riccia bicarinata Lindb.Riccia henriquesii var. mediterranea C. Massal. = Riccia bicarinata Lindb.Riccia heyeri (Huebener ex Genth) Müll. Frib. = Riccia canaliculata Hoffm.Riccia huebeneriana var. cavernosa (Hoffm.) Ast = Riccia cavernosa Hoffm.Riccia huebeneriana var. pseudo-frostii Schiffn. = Riccia huebeneriana Lindenb.Riccia hyi Levier ex Müll. Frib. nom. inval. = Riccia glauca var. ciliaris Warnst.Riccia incrassata Brot. = Oxymitra incrassata (Brot.) Sérgio & Sim-SimRiccia insularis Levier ex Steph. = Riccia sorocarpa Bisch.Riccia intumescens (Bisch.) Heeg = Riccia ciliata Hoffm. [57]Riccia intumescens Underw. nom. inval. = Ricciacrinita Taylor [57]Riccia klinggraeffii Gottsche = Riccia huebeneriana Lindenb.Riccia lamellosa var. americana M. Howe = Riccia lamellosa RaddiRiccia latzelii Schiffn. = Riccia ciliifera Link ex Lindenb.Riccia lescuriana Austin = Riccia beyrichiana Hampe ex Lehm.Riccia lescuriana var. glaucescens (Carrington) Müll. Frib. nom. incorrect. = Riccia beyrichianaHampe ex Lehm.Riccia lesquereuxii Lindb. = Riccia beyrichiana Hampe ex Lehm.Riccia levieri Schiffn. = Riccia beyrichiana Hampe ex Lehm.Riccia levieri var. algarvica Schiffn. ex Nicholson = Riccia beyrichiana Hampe ex Lehm.Riccia lindenbergiana Saut. = Riccia sorocarpa Bisch.Riccia linkii Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim

420 R.M. Ros et al.Riccia lusitanica Levier ex Steph. = Riccia bicarinata Lindb.Riccia major A. Roth = Riccia glauca L.Riccia mamillata Trab. ex Steph. [54]Riccia michelii var. ciliaris Levier nom. illeg. = Riccia michelii RaddiRiccia michelii var. subinermis Levier nom. illeg. = Riccia michelii RaddiRiccia microspora Steph. = Riccia frostii AustinRiccia minima L. p. p. = Riccia sorocarpa Bisch.Riccia minima L. p. p. = Riccia nigrella DC.Riccia minutissima Steph. = Riccia bifurca Hoffm. [56]Riccia montagnei Steph. = Riccia cavernosa Hoffm.Riccia munita Trab.nom. nud. = Riccia sorocarpa Bisch. [55]Riccia natans L . = Ricciocarpos natans (L.) CordaRiccia negrii Gola = Riccia crystallina L. emend. RaddiRiccia palaestina S. W. Arnell = Riccia frostii AustinRiccia palmata Lindenb.=Riccia michelii RaddiRiccia palmata var. minor Lindenb. = Riccia michelii RaddiRiccia panormitana Levier & Jack ex Lojacono-Pojero = Riccia sorocarpa Bisch.Riccia paradoxa De Not. = Riccia michelii RaddiRiccia pearsoni Steph. = Riccia nigrella DC.Riccia pedemontana Steph. = Riccia ciliifera Link ex Lindenb.Riccia plana Taylor = Riccia crystallina L. emend. RaddiRiccia polycarpa (Trab.) Jelenc [54]Riccia pseudo-frostii (Schiffn.) Müll. Frib. = Riccia huebeneriana Lindenb.Riccia pseudopapillosa Levier ex Steph. = Riccia papillosa MorisRiccia pusilla Warnst. = Riccia bifurca Hoffm.Riccia pyramidata Raddi non Willd.= Oxymitra incrassata (Brot.) Sérgio & Sim-SimRiccia raddiana J. B. Jack & Levier ex Steph. = Riccia sorocarpa Bisch.Riccia reticulata J. F. Gmel. nom. illeg. = Corsinia coriandrina (Spreng.) Lindb.Riccia riatensis Trab. nom. nud. = Riccia lamellosa Raddi [55]Riccia ruppinensis Steph. = Riccia glauca L. var. glaucaRiccia sanguinea Kashyap = Riccia frostii AustinRiccia sorocarpa var. raddiana (J. B. Jack & Levier ex Steph.) Croz. ex Casares-Gil = Ricciasorocarpa Bisch.Riccia sorocarpa f. raddiana (J. B. Jack & Levier ex Steph.) Chevassut & Quézel nom. inval.= Riccia sorocarpa Bisch.Riccia spinosissima Steph. = Riccia ciliata Hoffm.Riccia subciliata Trab. nom. inval. = Riccia glauca var. ciliaris Warnst.Riccia subcrispula Warnst. = Riccia bifurca Hoffm.Riccia subinermis Lindb. = Riccia glauca var. ciliaris Warnst.Riccia subtumida Milde = Riccia ciliata Hoffm.Riccia tellinii Gola = Riccia cavernosa Hoffm.Riccia terracianoi Gola = Riccia cavernosa Hoffm.Riccia trichocarpa M. Howe = Riccia crinita TaylorRiccia trichophylla Gasp. ex Müll. Frib. nom. inval. = Riccia papillosa MorisRiccia tumida Lindenb. = Riccia michelii RaddiRiccia velutina Wilson ex Hook. = Ricciocarpos natans (L.) CordaRiccia venosa A. Roth = Riccia glauca L. var. glauca

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 421Riccia warnstorfii f. subinermis (Warnst.) Warnst. ex Kavina = Riccia warnstorfii var.subinermis Warnst.Riccia watsonii Austin = Riccia frostii AustinRiccia zachariae Lorb. ex Müll. Frib. = Riccia gougetiana Durieu & Mont. var. gougetianaRicciella fluitans (L.) A. Braun = Riccia fluitans L.Ricciella fluitans var. canaliculata (Hoffm.) Hartm. = Riccia canaliculata Hoffm.Ricciella fluitans f. terrestris Jaapnom. inval. = Riccia fluitans L.Ricciella heyeri Huebener ex Genth = Riccia canaliculata Hoffm.Ricciella huebeneriana (Lindenb.) Dumort. = Riccia huebeneriana Lindenb.Ricciella huebeneriana var. cavernosa (Hoffm.) Casares-Gil = Riccia cavernosa Hoffm.Ricciella polycarpa Trab. = Riccia polycarpa (Trab.) JelencRicciella rautanenii Steph. = Riccia cavernosa Hoffm.Riccinia perennis (Steph.) Trab. = Riccia perennis Steph.Ricciocarpos velutina (Wilson ex Hook.) Steph. = Ricciocarpos natans (L.) CordaRiella battandieri Trab. = Riella notarisii (Mont.) Mont.Riella clausonis Letourn. = Riella parisii GottscheRiella cossoniana var. echinata Müll. Frib.nom. inval . = Riella helicophylla (Bory & Mont.)Mont.Riella gallica Balansa = Riella notarisii (Mont.) Mont.Riella reuteri Mont. = Riella notarisii (Mont.) Mont.Riella reuteri f. gallica (Balansa) Trab. = Riella notarisii (Mont.) Mont.Roemeria multifida (L.) Raddi nom. illeg. = Riccardia multifida (L.) GrayRoemeria palmata (Hedw.) Raddi nom. illeg. = Riccardia palmata (Hedw.) Carruth.Ruppinia italica (Sassi) Trevis. = Plagiochasma rupestre (J. R. Forst. & G. Forst.) Steph.Rupinia pyramidata (Raddi) Corda non (Willd.) Corda = Oxymitra incrassata (Brot.)Sérgio & Sim-SimRupinia rupestris (J. R. Forst. & G. Forst.) Sw. = Plagiochasma rupestre (J. R. Forst. &G. Forst.) Steph.Saccobasis polita (Nees) H. Buch = Tritomaria polita (Nees) Jörg.Saccogyna graveolens (Schrad.) Lindb. = Geocalyx graveolens (Schrad.) NeesSarcocyphos aemulus Limpr. = Marsupella condensata (Ångstr. ex C. Hartm.) Kaal.Sarcocyphos alpinus Gottsche ex Husn. = Marsupella alpina (Gottsche ex Husn.) BernetSarcocyphos boeckii Austin = Marsupella boeckii (Austin) Lindb.Sarcocyphos capillaris Limpr. = Marsupella boeckii (Austin) Lindb.Sarcocyphos commutatus Limpr. = Marsupella commutata (Limpr.) BernetSarcocyphos confertus Limpr. = Marsupella brevissima (Dumort.) GrolleSarcocyphos densifolius Nees = Marsupella emarginata (Ehrh.) Dumort.☞ Sarcocyphos emarginatus (Ehrh.) C. Hartm. = Marsupella emarginata (Ehrh.) Dumort.Sarcocyphos ehrhardtii Corda nom. illeg. = Marsupella emarginata (Ehrh.) Dumort.Sarcocyphos ehrhardtii var. aquaticus (Nees) Gottsche et al. = Marsupella emarginata(Ehrh.) Dumort.Sarcocyphos ehrhardtii var. robustus De Not. = Marsupella emarginata (Ehrh.) Dumort.Sarcocyphos funckii (F. Weber & D. Mohr) Nees = Marsupella funckii (F. Weber & D.Mohr) Dumort.Sarcocyphos muelleri Nees = Marsupella funckii (F. Weber & D. Mohr) Dumort.Sarcocyphos neglectus Limpr. = Marsupella sprucei (Limpr.) BernetSarcocyphos pygmaeus Limpr. = Marsupella funckii (F. Weber & D. Mohr) Dumort.Sarcocyphos revolutus Nees = Apomarsupella revoluta (Nees) R. M. Schust.

422 R.M. Ros et al.Sarcocyphos sparsifolius Lindb. = Marsupella sparsifolia (Lindb.) Dumort.Sarcocyphos sphacelatus (Gieseke ex Lindenb.) Nees = Marsupella sphacelata (Gieseke exLindenb.) Dumort.Sarcocyphos sprucei Limpr. = Marsupella sprucei (Limpr.) BernetSarcocyphos styriacus Limpr. = Marsupella sparsifolia (Lindb.) Dumort.Sarcocyphos sullivantii De Not. = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort.Sarcocyphos ustulatus auct . = Marsupella sprucei (Limpr.) BernetSauteria grandis Lindb. = Peltolepis quadrata (Saut.) Müll. Frib.Sauteria hyalina (Sommerf.) Lindb. = Athalamia hyalina (Sommerf.) S. Hatt.Sauteria quadrata Saut. = Peltolepis quadrata (Saut.) Müll. Frib.Sauteria seriata Lindb. nom. illeg. = Athalamia hyalina (Sommerf.) S. Hatt.Scapania aconiensis De Not. = Scapania nemorea (L.) GrolleScapania bartlingii (Hampe ex Nees) Nees nom. illeg. = Scapania cuspiduligera (Nees)Müll. Frib.Scapania biroliana C.Massal. & Carestia = Scapania compacta (A. Roth) Dumort.Scapania buchii Müll. Frib. nom. illeg. = Scapania lingulata H. BuchScapania carestiae De Not. = Scapania cuspiduligera (Nees) Müll. Frib.Scapania carinthiaca var. massalongi Müll. Frib. = Scapania carinthiaca J. B. Jack ex Lindb.Scapania casaresana Steph. = Scapaniagracilis Lindb.Scapania compacta var. biroliana (C. Massal. & Carestia) C. Massal. = Scapania compacta(A. Roth) Dumort.Scapania convexa (Scop.) Pearson = Scapania umbrosa (Schrad.) Dumort.Scapania curta var. geniculata (C. Massal.) Müll. Frib. = Scapania helvetica Gottsche☞ Scapania curta var. spinulosa (Nees) Gottsche et al. = Scapania carinthiaca J. B. Jack exLindb.Scapania dentata (Dumort.) Dumort. = Scapania undulata (L.) Dumort.Scapania franzoniana De Not. = Scapania subalpina (Nees ex Lindenb.) Dumort.Scapania geniculata C. Massal. = Scapania helvetica Gottsche☞ Scapania gracilis var. laxifolia (Dumort.) Dumort. = Scapania gracilis Lindb.Scapania gracilis var. minor (Pearson) Macvicar = Scapania gracilis Lindb.Scapania intermedia (Husn.) Pearson = Scapania undulata (L.) Dumort.Scapania joergensenii Schiffn. ex Müll. Frib. = Scapania nemorea (L.) GrolleScapania massalongii (Müll. Frib.) Müll. Frib. = Scapania carinthiaca J. B. Jack ex Lindb.Scapania microphylla Warnst. = Scapania lingulata H. BuchScapania microscopica Culm. = Diplophyllum obtusifolium (Hook.) Dumort.Scapania mucronata var. praetervisa (Meyl.) H. Buch = Scapania praetervisa Meyl.Scapania nemorea subsp. crassiretis (Bryhn) Potemkin = Scapania crassiretis BryhnScapania nemorosa Dumort. nom. illeg. = Scapania nemorea (L.) GrolleScapania nemorosa var. aconiensis (De Not.) C. Massal. = Scapania nemorea (L.) GrolleScapania nemorosa var. fallaciosa Schiffn. = Scapania nemorea (L.) GrolleScapania oakesii Austin = Scapania undulata (L.) Dumort.Scapania obliqua (Arnell) Schiffn. = Scapania uliginosa (Sw. ex Lindenb.) Dumort.Scapania recurva Steph. ex Müll. Frib. = Scapania gracilis Lindb.Scapania resupinata (L.) Dumort. = Scapania undulata (L.) Dumort.Scapania resupinata sensu A. L. M. Guim. = Scapania gracilis Lindb.Scapania rosacea (Corda) Nees nom. inval. = Scapania curta (Mart.) Dumort.Scapania rotundata Warnst. = Scapania paludicola Loeske & Müll. Frib.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 423Scapania subalpina var. undulifolia (Nees) Gottsche et al. = Scapania subalpina (Nees exLindenb.) Dumort.Scapania tyrolensis (Nees) Nees = Scapania aequiloba (Schwägr.) Dumort.Scapania undulata var. aequatiformis (De Not.) C. Massal. & Carestia = Scapania undulata(L.) Dumort.Scapania undulata var. dentata (Dumort.) Douin = Scapania undulata (L.) Dumort.Scapania undulata var. dentata (Dumort.) McArdle. nom. inval. = Scapania undulata (L.)Dumort.Scapania undulata var. integrifolia Lindb. = Scapania undulata (L.) Dumort.Scapania undulata var. paludosa Müll. Frib. = Scapania paludosa (Müll. Frib.) Müll. Frib.Scapania undulata var. purpurascens (Huebener) Cogn. = Scapania undulata (L.) Dumort.Scapania undulata var. purpurea Neesnom. inval. = Scapania undulata (L.) Dumort.Scapania vexata C. Massal. nom. illeg. = Scapania scapanioides (C. Massal.) GrolleScapaniella carinthiaca (J. B. Jack ex Lindb.) H. Buch = Scapania carinthiaca J. B. Jack exLindb.Scapaniella massalongi (Müll. Frib.) H. Buch = Scapania carinthiaca J. B. Jack ex Lindb.Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstantinova = Lophozia grandiretis (Lindb.ex Kaal.) Schiffn.Schistochilopsis incisa (Schrad.) Konstantinova = Lophozia incisa (Schrad.) Dumort.Schistochilopsis opacifolia (Culm. ex Meyl.) Konstantinova = Lophozia opacifolia Culm. exMeyl.Solenostoma amplexicaule (Dumort.) Steph. = Jungermannia sphaerocarpa Hook.Solenostoma atrovirens (Dumort.) Müll. Frib. nom. illeg. = Jungermannia atrovirensDumort.Solenostoma caespiticium (Lindenb.) Steph. = Jungermannia caespiticia Lindenb.Solenostoma confertissimum (Nees) Schljakov = Jungermannia confertissima NeesSolenostoma cordifolium (Dumort.) Steph. = Jungermannia exsertifolia subsp. cordifolia(Dumort.) Vá≈ aSolenostoma crenulatum (Sm.) R. M. Schust. = Jungermannia gracillima Sm.Solenostoma crenulatum (Sm.) Mitt. nom. illeg. = Jungermannia gracillima Sm.Solenostoma handelii (Schiffn.) Müll. Frib. = Jungermannia handelii (Schiffn.) AmakawaSolenostoma hyalinum (Lyell) Mitt. = Jungermannia hyalina LyellSolenostoma lanceolatum (L.) Steph. = Jungermannia atrovirens Dumort.Solenostoma lanceolatum auct. non (L.) Steph. = Jungermannia leiantha GrolleSolenostoma levieri (Steph.) Steph. = Jungermannia confertissima NeesSolenostoma oblongifolium (Müll. Frib.) Müll. Frib. = Jungermannia pumila With.Solenostoma obovatum (Nees) C. Massal. = Jungermannia obovata NeesSolenostoma ontariense R. M. Schust. = Jungermannia hyalina LyellSolenostoma polare (Lindb.) R. M. Schust. = Jungermannia polaris Lindb.Solenostoma pumilum (With.) Müll. Frib. = Jungermannia pumila With.Solenostoma pusillum (C. E. O. Jensen) Steph. = Jungermannia jenseniana GrolleSolenostoma pusillum var. vinaceum R. M. Schust. = Jungermannia confertissima NeesSolenostoma scalare (Gray) Mitt. = Nardia scalaris GraySolenostoma schiffneri (Loitl.) Müll. Frib. = Jungermannia polaris Lindb.Solenostoma sphaerocarpoideum (De Not.) Paton & E. F. Warb. = Jungermannia atrovirensDumort.Solenostoma sphaerocarpum (Hook.) Steph. = Jungermannia sphaerocarpa Hook.Solenostoma subellipticum (Lindb. ex Kaal.) R.M. Schust. = Jungermannia subelliptica(Lindb. ex Kaal.) Levier

424 R.M. Ros et al.Solenostoma triste (Nees) Müll. Frib. = Jungermannia atrovirens Dumort.Southbya alicularia C. Massal. = Southbya tophacea (Spruce) SpruceSouthbya crenulata (Mitt.) Bernet = Jungermannia gracillima Sm.Southbya fennica (Gottsche) C. Hartm. = Arnellia fennica (Gottsche) Lindb.Southbya hyalina (Lyell) Husn. = Jungermannia hyalina LyellSouthbya obovata (Nees) Lindb. = Jungermannia obovata NeesSouthbya stillicidiorum (Raddi) Lindb. = Southbya tophacea (Spruce) SpruceSphaerocarpos algeriensis Trab.nom. nud. = Sphaerocarpos texanus Austin [64]Sphaerocarpos californicus Austin = Sphaerocarpos texanus AustinSphaerocarpos europaeus Lorb. = Sphaerocarpos texanus AustinSphaerocarpos notarisii Mont. = Riella notarisii (Mont.) Mont.Sphaerocarpos sphaerocarpos (Dicks.) M. Howe nom. illeg. = Sphaerocarpos micheliiBellardiSphaerocarpos terrestris Sm. nom. illeg. = Sphaerocarpos michelii BellardiSphagnoecetis communis Nees nom. illeg. = Odontoschisma sphagni (Dicks.) Dumort.Sphagnoecetis macounii Austin = Odontoschisma macounii (Austin) Underw.Sphenolobus ascendens Warnst. = Lophozia ascendens (Warnst.) R.M. Schust.Sphenolobus exsectiformis (Breidl.) Steph. = Tritomaria exsectiformis (Breidl.) LoeskeSphenolobus exsectiformis var. aequilobus (Culm.) Müll. Frib. = Tritomaria scitula (Taylor)Jörg.Sphenolobus exsectus (Schmidel ex Schrad.) Steph. = Tritomaria exsecta (Schmidel exSchrad.) LoeskeSphenolobus hellerianus (Nees ex Lindenb.) Steph. = Anastrophyllum hellerianum (Nees exLindenb.) R. M. Schust.Sphenolobus michauxii (F. Weber) Steph. = Anastrophyllum michauxii (F. Weber)H. BuchSphenolobus minutus (Schreb.) Berggr. = Anastrophyllum minutum (Schreb.) R. M. Schust.Sphenolobus politus (Nees) Steph. = Tritomaria polita (Nees) Jörg.Sphenolobus saxicolus (Schrad.) Steph. = Anastrophyllum saxicola (Schrad.) R. M. Schust.Sphenolobus scitulus (Taylor) Steph. = Tritomaria scitula (Taylor) Jörg.Staurophora pulchella Willd. nom. illeg. = Lunularia cruciata (L.) Lindb.Stephanina complanata (L.) Kuntze = Radula complanata (L.) Dumort.Strozzia conica (L.) Müll. Frib. = Conocephalum conicum (L.) Dumort.Strozzia hemisphaerica (L.) Müll. Frib. = Reboulia hemisphaerica (L.) RaddiStrozzius conicus (L.) Gray = Conocephalum conicum (L.) Dumort.Strozzius hemisphaericus (L.) Gray = Reboulia hemisphaerica (L.) RaddiTargionia bifurca Nees & Mont. = Targionia hypophylla L.Targionia convoluta Lindenb. & Gottsche = Targionia hypophylla L.Targionia germanica Corda nom. inval. = Targionia hypophylla L.Targionia hypophylla var. fimbriata (Müll. Frib.) Müll. Frib. = Targionia hypophylla L.Targionia hypophylla var. lorbeeriana (Müll. Frib.) Aleffi & Schumacker nom. inval. =Targionia lorbeeriana Müll. Frib.Targionia michelii Corda = Targionia hypophylla L.Targionia sphaerocarpos Dicks. = Sphaerocarpos michelii BellardiTelaranea nematodes sensu auct. eur. non (Gottsche ex Austin) M. Howe = Telaraneaeuropaea J.J. Engel & G.L.S. Merril [66]Telaranea sejuncta auct. eur. non (Ångstr.) S. W. Arnell = Telaranea europaea J.J. Engel &G.L.S. Merril [67]Telaranea setacea auct. non (Weber) Müll. Frib. = Kurzia pauciflora (Dicks.) Grolle

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 425Telaranea sylvatica (A. Evans) Müll. Frib. = Kurzia sylvatica (A. Evans) GrolleTelaranea trichoclados (Müll. Frib.) Müll. Frib. = Kurzia trichoclados (Müll. Frib.) GrolleTemnoma setiforme (Ehrh.) M. Howe = Tetralophozia setiformis (Ehrh.) SchljakovTesselina coriandrina (Spreng.) Dumort. = Corsinia coriandrina (Spreng.) Lindb.☞ Tesselina paleacea auct. = Oxymitra incrassata (Brot.) Sérgio & Sim-SimTesselina pyramidata (Raddi) Dumort. non (Willd.) Dumort. = Oxymitra incrassata (Brot.)Sérgio & Sim-SimTesselina pyramidata var. paleacea Lindb. = Oxymitra incrassata (Brot.) Sérgio & Sim-SimTesselina pyramidata var. paleacea (Bisch. ex Lindenb.) Crozals nom. illeg. = Oxymitraincrassata (Brot.) Sérgio & Sim-SimTesselina pyramidata var. polycarpa (Bisch. ex Lindenb.) Lindb. = Oxymitra incrassata(Brot.) Sérgio & Sim-SimThedenia blyttii (Moerch ex Hornem.) Hartm. nom. inval. = Moerckia blyttii (Moerch exHornem.) Brockm.☞ Trigonanthus curvifolius (Dicks.) Spruce = Nowellia curvifolia (Dicks.) Mitt.Trigonanthus francisci (Hook.) Hartm. = Cladopodiella francisci (Hook.) Jörg.Tritomaria polymorpha (R. M. Schust.) Grolle = Tritomaria polita (Nees) Jörg.Tritomaria scitula var. savoica Müll. Frib. = Tritomaria scitula (Taylor) Jörg.Annotations1. Bischler (2004) reported several taxa from Mediterranean countries withoutdetails of locality. The following require verification as they are the only knownreports for their respective countries.–For Algeria: Calypogeia muelleriana (Schiffn.) Müll. Frib., Cephaloziaconnivens (Dicks.) Lindb., C. lunulifolia (Dumort.) Dumort., Diplophyllumalbicans (L.) Dumort., Dumortiera hirsuta (Sw.) Nees , Jungermannia obovataNees , J. pumila With., Leiocolea collaris (Nees) Schljakov , Lophocoleaheterophylla (Schrad.) Dumort., L. minor Nees , Marsupella emarginata (Ehrh.)Dumort., Riccardia latifrons (Lindb.) Lindb., Riccia ligula Steph. and Scapaniagracilis Lindb.–For Lebanon: Anthoceros punctatus L., Athalamia hyalina (Sommerf.)S. Hatt., Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Bazzania flaccida(Dumort.) Grolle, Blepharostoma trichophyllum (L.) Dumort., Cephaloziellacalyculata (Durieu & Mont.) Müll. Frib., Cololejeunea minutissima (Sm.) Schiffn.,Fossombronia angulosa (Dicks.) Raddi, F. caespitiformis subsp. multispira(Schiffn.) J. R. Bray & D. C. Cargill (as F. husnotii Corb.), Jungermannia leianthaGrolle, Lophocolea bidentata (L.) Dumort., Nardia compressa (Hook.) Gray ,Pellia epiphylla (L.) Corda , Plagiochila porelloides (Torrey ex Nees) Lindenb.,Porella arboris-vitae (With.) Grolle, P. platyphylla (L.) Pfeiff., Radula complanata(L.) Dumort., Riccia bifurca Hoffm., R. crystallina L. emend. Raddi, R. micheliiRaddi, Scapania aequiloba (Schwägr.) Dumort. and S. aspera Bernet & M. Bernet .–For Morocco: Riccia huebeneriana Lindenb., Riella cossoniana Trab.and R. parisii Gottsche.–For Tunisia: Riccia warnstorfii Lindenb. and Sphaerocarpos texanusAustin.–For Turkey: Plagiochila bifaria (Sw.) Lindenb.–For Europe, see Söderström et al. (2007).2. The nomenclatural problems of Anthoceros agrestis Paton and its relationshipwith A. punctatus L. have been discussed by Söderström et al . (2002). In the

426 R.M. Ros et al.Mediterranean area the two species have been regularly confused, both ontaxonomic and nomenclatural basis, but most records probably belong to A.punctatus. Further studies of Mediterranean material are needed (including A.multifidus L.) to clarify the distribution of the two taxa.3. Phaeoceros carolinianus (Michx.) Prosk. was reported from Crete by Jovet-Ast& Bischler (1968). This is a doubtful record that is rejected here. Jovet-Ast &Bischler include Crete in the distribution of “ P. laevis subsp. carolinianus”, citingthe paper by Düll (1966, p. 743). However in this paper Düll does not distinguishsubsp. carolinianus and subsp. laevis . He lists only P. laevis .4 . Anthoceros beltrani Casares-Gil was described by Casares-Gil (1915) fromSpain, and later synonymized by Proskauer (1958) with Phaeoceros laevis (L.)Prosk. Nevertheless this synonymy is questionable because of the paroicouscondition of A. beltrani. Further studies are needed.5 . Anthoceros dichotomus var. gussonei Zodda was synonymized with Phaeoceroslaevis (L.) Prosk. by Proskauer (1958), although it often has been erroneouslyinterpreted in the Mediterranean area as a synonym of Phymatoceros bulbiculosus(Brot.) Stotler, W. T. Doyle & Crand.-Stotl.6. For the taxonomy of the genus Phymathoceros, we have followed Stotler et al.(2005) and Crandall-Stotler et al. (2006).7. According to Schuster (1988), Anthelia nivalis (F. Weber & D. Mohr) Lindb. is anomen dubium and could correspond to Anthelia juratzkana (Limpr.) Trevis. p. p.and A. julacea (L.) Durmort. p. p. (type probably A. julacea ).8. The presence of the American Asterella elegans (Spreng.) Trevis. was rejectedfrom Corsica by Söderström et al . (2002) and the report from Turkey (Bornmüller,1931) must also be rejected.9 . There has been some confusion in the literature with the name Fimbriaria nanaLindenb., that has been considered by some authors to be a synonym of Asterellagracilis (F. Weber) Underw., but Grolle (1980) clearly established its identity withAthalamia hyalina (Sommerf.) S. Hatt.10. Fimbriaria fragrans (Balbis) Nees 1825 ( ≡ Marchantia fragrans Balbis) isMannia fragrans (Balbis) Frye & L. Clark while F. fragrans (Schleich.) Nees 1838( ≡ Marchantia fragrans Schleich.) is a synonym to Asterella saccata (Wahlenb.) A.Evans. Both variants of F. fragrans have been used in the study area. It is likelythat most of the Mediterranean reports correspond to Mannia fragrans rather thanAsterella saccata , but further revision is needed.11. The problems in applying the name Calypogeia trichomanis (L.) Corda arediscussed by Söderström et al . (2002), who note that this name has been variouslyinterpreted in Europe as both C. azurea Stotler & Crotz (Grolle & Long, 2000) andC. muelleriana (Schiffn.) Müll. Frib . The same is true of the Mediterranean area.12. Jungermannia scalaris Schmidel ex Schreb. is a nomen dubium and couldcorrespond either to Calypogeia trichomanis (L.) Corda nom. rejic. (currentlyequated by most authors with Calypogeia azurea Stotler & Crotz), or toOdontoschisma sp. (Vá≈ a, 1976).13. Cephalozia macrostachya Kaal. was erroneously included in the checklist ofSpain by Casas (1998). Therefore it should be excluded from Spain.

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 42714. Cephaloziella gallica (Douin) Douin was considered by Müller (1954-1957) tobe a synonym of Cephaloziella calyculata (Durieu & Mont.) Müll. Frib., but it hasbeen synonymized with C. hampeana (Nees) Schiffn. by recent authors(Schumacker & Vá≈ a, 2000, 2005; Söderström et al., 2002) and that is the criterionfollowed here.15. Cephaloziella myriantha (Lindb.) Schiffn. was considered by Schuster (1980) tobe a synonym of C. rubella var. elegans (Heeg) R. M. Schust. (= C. elegans (Heeg)Schiffn.), but Damsholt (2002) referred it to C. rubella (Nees) Warnst. var. rubella ,which is the interpretation followed here.16. Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort. and Porella obtusata(Taylor) Trevis., have been reported from localities formerly included in theAlbanian territory, but now belong to Montenegro.17. Conocephalum salebrosum Szweyk., Buczkowska & Odrzykoski has beenrecently described by Szweykowski et al. (2005) as a separated species from C.conicum (L.) Dumort. The new species is in some areas of Europe more commonthan C. conicum and it is impossible to know which species is meant by the lattername prior to 2005. Conocephalum salebrosum will certainly turn out to be morewidespread in the Mediterranean area.18. Recent studies on the genus Fossombronia Raddi by Stotler & Crandall-Stotler(2006) propose a new treatment for several taxa, and this treatment has beenfollowed here.19. Fossombronia intestinalis Taylor was erroneously reported from Algeria bySöderström et al. (2002). It is an Australian taxon that is also known from SouthAfrica and it should be rejected from the Mediterranean area.20. According to Scott & Pike (1988), Fossombronia intermedia Corb. ex Trab.nom. inval. (= Fossombronia caespitiformis var. intermedia Corb. ex Trab. nom.inval.) could be ascribed either to F. caespitiformis De Not. ex Rabenh. or to F.wondraczekii (Corda) Lindb. However Stotler & Crandall-Stotler (2006) considerit to be a synonym of F. echinata Macvicar. At present it is not possible to know theidentity of the Mediterranean specimens so named.21. Jungermannia crispula Brot. was described from Portuguese material and waslater transferred to Fossombronia as F. crispula (Brot.) R. M. Schust. by Schuster(1992a), but no other report of this species has been found in European works.Machado (1925) included J. crispula in Fossombronia pusilla (L.) Nees p. p. and F.caespitiformisp. p., but according to Sérgio & Carvalho (2003), no original materialof J. crispula has been found. Therefore it should be considered a nomen dubium.22. Fossombronia crispa is problematic and should be considered doubtful forEurope (cf. Söderström et al., 2007).23. The identity of Fossombronia loitlesbergeri Schiffn. is controversial. Paton(1999) and Scott & Pike (1988) considered it to be related to Fossombroniawondraczekii (Corda) Lindb., but Stotler & Crandall-Stotler (2006), whom wefollow here, treated it as a synonym of F. pusilla (L.) Nees.24. Frullania muscicola Steph. is an Asian species that was reported fromMacaronesia by Sérgio (1985). Later Bisang et al. (1989) considered that the Asianand Macaronesian specimens would be better considered as Frullania cesatianavar. muscicola Bisang et al ., and Sim-Sim et al. (1995) assigned the Macaronesianspecimens to the endemic F. azorica Sim-Sim et al.

428 R.M. Ros et al.The reports from the Canary islands of Frullania riparia Hampe ex Lehm. byArnell (1961) also correspond to this taxon (Casas et al ., 1989).25 . Several infraspecific taxa have been recognized in Frullania dilatata (L.)Dumort. and F. tamarisci (L.) Dumort. for Portugal and Madeira by Sim-Sim(1999). In addition Losada Lima & Vanden Berghen (1985) described Frullaniadilatata var. elongata from the Canary Islands, though it was subsequentlysynonymized with F. ericoides (Nees) Mont. by Bisang et al. (1989). However theseterritories form only a relatively small part of the Mediterranean Basin, and thecriteria used by Sim-Sim (1989) are difficult to apply to the whole region. We havetherefore adopted a conservative treatment in this checklist.26. Grolle & Long (2000) referred Frullania davurica Hampe to F. jackii Gottsche.The Mediterranean material identified as F. davurica probably corresponds to F.jackii, which is also the interpretation of Schumacker & Vá≈ a (2005). Furtherconfirmation is needed.27. Jubula pennsylvanica (Steph.) A. Evans was reported doubtfully from Turkeyby Schiffner (1909) sub Jubula hutchinsiae var. sullivanti Spruce. According toGrolle & Long (2000), it could be the same taxon as Jubula hutchinsiae subsp.javanica (Steph.) Verd. The relationship between the two taxa needs to beassessed. Two varieties have also been recognized in Macaronesia by Sim-Sim et al.(2002): Jubula hutchinsiae subsp . hutchinsiae var. hutchinsiae and Jubulahutchinsiae subsp. hutchinsiae var. integrifolia Lindb. However the correct authorcitation of the latter name is Jubula hutchinsiae subsp. hutchinsiae var. integrifolia(Steph.) S. Hatt., and it has previously been synonymized with subsp. javanica (seeSöderström et al., 2007).28. Jungermannia compacta Mazziari nom. nud. non Roth., J. foscardiana Mazziarinom. nud., J . guilfortiana Mazziari nom. nud., J. lacera Mazziari nom. nud., and J.limosa Mazziari nom. nud. were reported from Greece by Mazziari (1851) but noother reference to these names has been found in the literature.29. Jungermannia jenseniana Grolle was reported from Spain by Lloret et al.(1997), but after revision of the specimen by M. Infante, it was assigned to Nardiascalaris Gray. Therefore the species is at the moment rejected from the area.30. Nardia hyalina var. colorata Samp. is a nomen nudum and is based on a separatetype (from Portugal) than Nardia hyalina var. colorata (Mart.) Dalla Torre &Sarnth. (type from Germany). Vá≈ a (1975) treated the latter as a synonym ofJungermannia gracillima . Later Sérgio & Carvalho (2003) stated that “ Nardiahyalina var. colorata Samp. = ?? Jungermannia hyalina Lyell”. Consequently theidentity of the name remains uncertain and requires further studies.31. Leiocolea collaris (Nees) Schljakov is sometimes treated as a synonym of L.bantriensis (Hook.) Jørg. Thus, the literature reports of the latter must be treatedwith caution.32. Leiocolea turbinata f. aquatica Jelenc nom. nud. was reported by Jelenc (1955)from Morocco but no other record or reference to it has been found.33. According to Schuster (1980), Jungermannia serpyllifolia Dicks. nom. illeg. correspondsto Lejeunea patens Lindb. but the name Lejeunea serpyllifolia sensu Lib.nom. illeg. refers to another taxon, which led Bonner & Miller (1960) to proposethe new name Lejeunea libertiae Bonner & H. A. Miller. This was later synonymizedwith L. cavifolia (Ehrh.) Lindb. The literature reports of L. serpyllifoliaare difficult to place but do mostly correspond to L. cavifolia .

Hepatics and Anthocerotes of the Mediterranean, an annotated checklist 42934. Schuster (1980) confirmed the presence of L. flava subsp. moorei (Lindb.) R.M.Schust. in Ireland. In Macaronesia it has also been reported from Madeira.According to Grolle & Long (2000), the identity of the European andMacaronesian specimens referred to L. flava remains to be confirmed. In thispaper they have been included in Lejeunea flava (Sw.) Nees.35. Lepidozia pearsonii Spruce was reported by Reinoso (1985) from Spain andconsidered by Casas (1998) as a species whose occurrence in Spain neededconfirmation, which led to Söderström et al . (2002) to exclude it from Spain. Afterthat, the specimen has been revised by R. Schumacker as Lepidozia reptans (L.)Dumort. Therefore the species should be rejected from the Mediterranean area.36. The report of Lophozia latifolia R. M. Schust. (sub L. propagulifera (Gottsche)Steph.) from Serbia given in Sabovljevi † & Natcheva (2006) is almost certainlyer roneous. This is an arctic taxon that has been syn onymized with Lophozia jurensisMüll. Frib. (whose type is from the Alps), but this synonymy is very questionableand “true” L. propagulifera has never been reported from central Europe (seeSöderström et al., 2007).37. Recent molecular studies (Roo et al., 2007) show that Lophozia silvicola H.Buch is a species well separated from L. ventricosa (Dicks.) Dumort.38. Lophozia confertifolia Schiffn. is not conspecific with true L. wenzelii (Nees)Steph. as stated by Müller (1954-1957), and we have followed Schuster (1969) andSchumacker & Vá≈ a (2005) who consider Lophozia confertifolia to be a synonymof Lophozia ventricosa (Dicks.) Dumort (see Söderström et al., 2007).39. Grimaldia atlantica Trab. nom. inval. ( Mannia atlantica Jelenc nom. inval.) wasreported by Trabut (1941) from Algeria, but no other record or reference to ithave been found.40. According to Grolle & So (2003), Metzgeria violacea (Ach.) Dumort. is thecorrect name for M. fruticulosa auct. eur.41. The identity of the plants named Nardia silvrettae (Gottsche ex Dumort.)Trevis. is doubtful since it has been synonymized with both N. geoscyphus (DeNot.) Lindb. (Vá≈ a, 1976) and N. insecta Lindb. (Schumacker & Vá≈ a, 2005).According to Geissler & Bischler (1989), its basionym is Marsupella silvrettae Gott.ex Dumort., 1874 ( ≡ Jungermannia silvrettae Gottsche & Rabenh. nom. inval.).Schumacker & Vá≈ a (2000, 2005) refer it to Nardia insecta Lindb., but cite it with adifferent authority – Nardia silvrettae Douin.42. Pellia fabbroniana f. undulata Jelenc nom. inval. was reported by Jelenc (1955)from Algeria but no other record or reference to it has been found.43. Jungermannia fuciformis Bory (= Pellia fuciformis (Bory) Nees) is an uncertainEuropean name reported from Greece by Bory de Saint-Vicent (1832). Its identityis still to be assessed but Coppey (1908) stated that it could be a sterile form ofPellia epiphylla (L.) Corda.44. Plagiochila africana Steph. was erroneously reported from Libya bySöderström et al. (2002). It is a west African tropical species that should be rejectedfrom the Mediterranean area.45. The nomenclatural history of Plagiochila asplenioides (L. emend. Taylor)Dumort. is complex and it is not always possible to know whether a particularreport belongs to P. asplenioides s. str. or P. porelloides (Nees) Lindenb. Mostearlier reports probably belong to the latter (cf. Söderström et al., 2002).

430 R.M. Ros et al.46. Plagiochila atlantica Lindenb. ex Lehm. has been synonymized with P.heterophylla Lindenb. ex Lehm by Heinrichs (2002).47. Plagiochila maderensis Gottsche ex Steph. has recently been reinstated forMadeira and the Canary Islands by Rycroft et al. (2004).48. Fontinha (2005) excluded Porella arboris-vitae (With.) Grolle and Porellaplatyphylla (L.) Pfeiff. from Madeira. Bischler et al. (2006) stated that the firstspecies is restricted to southern continental areas while the second has a“European and Asiatic (North American?)” distribution. Further studies areneeded to establish conclusively their presence in the Canary Islands, in view ofthe ecological differences that characterize the individual islands.49. Madotheca canariensis (F. Weber) Nees has been reported from NorthernAfrica by Trabut (1941) and others without mention of locality or country andBischler (2004) placed it in Morocco. Trabut used this name for specimensintermediate between Madotheca laevigata (Schrad.) Dumort. and Madothecathuja (Dicks.) Dumort. The two latter names are synonyms of Porella arboris-vitae(With.) Grolle, but M. thuja was earlier frequently used for P. obtusata (Taylor)Trevis. The identity of the North African plants is therefore questionable andPorella canariensis (F. Weber) Underw. should be rejected until it is positivelyverified.50. Ros et al. (1999) erroneously reported Porella cordaeana (Huebener) Moorefrom Libya based on a supposed record of Rungby (1962). But this authorreported this species only for Morocco in Northern Africa. In addition, Porellapinnata L. was reported erroneously by Ros et al. (1999) from Morocco and shouldbe rejected from this country.51. In several works Achiton quadratum (Scop.) Corda has been erroneouslyassigned to Reboulia hemisphaerica (L.) Raddi. According to Bonner (1962b) thebasionym of this name is Marchantia quadrata Scop., which actually is thebasionym of Preissia quadrata (Scop.) Nees.52. Düll (1983) included the name Radula physoloba auct. as a synonym of R.aquilegia (Hook. f. & Taylor) Gottsche et al. Later Sérgio & Carvalho (2003)excluded this species from Portugal and considered it to be a synonym of Radulalindenbergiana Gottsche ex C. Hartm. In both cases the use of this name seems tobe erroneous, as Radula physoloba Mont. is a species of New Zealand. Thus, itshould be excluded from the Mediterranean lists.53. For the taxonomy of the genus Riccia L., the monographic treatment by Jovet-Ast (1986) has been followed here.54. Several species that have been reported from Northern Africa were notincluded in the monograph of the genus Riccia L. in the Mediterranean area byJovet-Ast (1986), but mentioned by Söderström et al . (2002), and their identityremains doubtful.– Riccia chinensis Herz. was proposed as new to Algeria by Hariot(1913) as Riccia chinensis Steph. ex Hariot from Tahount Arak (Mouydir), withthe observation that the specific name given by Stephani was provisional becauseit seemed to be unsuitable for an African species. Moreover it lacked a Latindiagnosis and was therefore an invalid name. Herzog subsequently validated thename but using material from China as type. The identity of the Algerian materialis therefore doubtful.

432 R.M. Ros et al.60. Despite the statements by Grolle & Long (2000) and Geissler & Bischler(1990), Riella numidica Trab. and Riella sersuensis Trab. were validly described,since the protologue of both species includes a Latin diagnosis (in agreement withthe article 36.1 of the Missouri ICBN).61. Scapania brevicaulis Taylor has been reported from Italy by Potemkin (2002),and this is the only report for the Mediterranean area. Because the author did notgive any indication of the locality, the report should be considered doubtful untilspecimens can be checked.62. Scapania calcicola (Arnell & J. Perss.) Ingham was erroneously reported fromthe Balearic Islands by Casas et al. (2004) without giving any locality, and shouldbe rejected.63. Jungermannia cordifolia Brot. non Hook. was described from Portugal. On thebasis of the original description, Sim-Sim (1989) stated that this taxoncorresponded to Scapania compacta (A. Roth) Dumort. and proposed thissynonymy without studying the type. Since the type cannot be located a formalsynonymyzation should be made following designation of a neotype.64. Sphaerocarpos algeriensis Trab. nom. nud. was reported by Trabut (1941) fromAlgeria. Jovet-Ast (1956) indicated that the morphological characters given for S.algeriensis fall within the range of variation observed in S. texanus Austin inMorocco and that it is probably only a synonym of that species.65. Schumacker & Vá≈ a (2005) proposed a new combination for Lepidozia azoricain the genus Telaranea as T. azorica (H. Buch & Perss.) Pócs ex Schumacker &Vá≈ a.66. Telaranea nematodes (Gottsche ex Austin) M. Howe was considered to be thecorrect name for the species present in Europe and Macaronesia until Engel &Merrill (2004) established a new name for it, Telaranea europaea J. J. Engel & G.L. S. Merrill.67. Telaranea sejuncta auct. non (Angst.) Arnell is a confusing taxon. It wasassigned to T. nematodes ( Gottsc he ex Austin) M. Howe by Arnell (1963), and thissynonymy has been widely used. According to Engel & Merrill (2004), “all thepublished statements of distribution, descriptions and illustrations of T. sejuncta(apart from the type) are suspect, since they may include some or all the taxaformerly included in T. nematodes , as well as T. diacantha , when, in fact, it is noneof these”. In our current state of knowledge, it can be assumed that all theMediterranean reports of T. nematodes and T. sejuncta correspond to T. europaea .68. Trichocolea tomentella (Ehrh.) Dumort. was recorded from Malta by Gulia(1858-59) and Sommier & Caruana Gatto (1915) as “Jungermannia tomentella(Ehr.)”. These reports are almost certainly erroneous and are here rejected.Acknowledgements. The authors wish to express their gratitude to David Longand Jesús Muñoz who kindly provided information concerning various taxa, and to SusanaRams and Olaf Werner for their help with the tables and figure. Ana Séneca helped tocoordinate the list with the European checklist. Two of the co-authors, Lars Söderströmand Tom Blockeel, made an extra contribution to improve the quality of the manuscript:LS revised the whole list and made important corrections and additions, while TB revisedthe English. This research has been partially funded by the Spanish Ministerio deEducación y Ciencia (Projects CGL2005-00028 and CGL2004-03513).

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