Actual checklist of Tardigrada species (2009-2012, Ver. 21: 30-06 ...

<strong>Actual</strong> <strong>checklist</strong> <strong>of</strong> <strong>Tardigrada</strong> <strong>species</strong><br />

(<strong>2009</strong>-<strong>2012</strong>, <strong>Ver</strong>. <strong>21</strong>: <strong>30</strong>-<strong>06</strong>-<strong>2012</strong>)<br />

PETER DEGMA 1 ROBERTO BERTOLANI 2 ROBERTO GUIDETTI 2<br />

1 Department <strong>of</strong> Zoology, Comenius University in Bratislava, Mlynská dolina B-1, SK-84<strong>21</strong>5<br />

Bratislava, Slovakia; e-mail: degma@fns.uniba.sk<br />

2 Department <strong>of</strong> Biology, University <strong>of</strong> Modena and Reggio Emilia, via campi <strong>21</strong>3/D, 41100<br />

Modena, Italy; e-mail: roberto.bertolani@unimore.it, roberto.guidetti@unimore.it<br />

Introduction<br />

More than one thousand <strong>Tardigrada</strong> <strong>species</strong> were included in the recently published<br />

<strong>checklist</strong> (Guidetti, R. & Bertolani, R. 2005. Tardigrade taxonomy: an updated check list <strong>of</strong><br />

the taxa and a list <strong>of</strong> characters for their identification. Zootaxa, 845, 1–46.) plus the<br />

additions and corrections to this <strong>checklist</strong> (Degma, P. & Guidetti, R. 2007. Notes to the current<br />

<strong>checklist</strong> <strong>of</strong> <strong>Tardigrada</strong>. Zootaxa, 1579, 41–53.). For practical reasons, we have joined these<br />

two papers (without comments added to particular taxa as well as without references published in<br />

these papers) into an accurate combined version <strong>of</strong> the <strong>checklist</strong>s. This <strong>checklist</strong> is free for all<br />

users, but utilization <strong>of</strong> it requires the citation <strong>of</strong> the two original papers. This paper is the<br />

platform for occasional upgrades (date <strong>of</strong> latest upgrade is in the title). All changes, additions and<br />

corrections in comparison with both papers are written in red font. If you also use these changes,<br />

please also cite this <strong>checklist</strong> (Degma, P., Bertolani, R. & Guidetti, R. <strong>2009</strong>-<strong>2012</strong>. <strong>Actual</strong><br />

<strong>checklist</strong> <strong>of</strong> <strong>Tardigrada</strong> <strong>species</strong>. <strong>Ver</strong>. <strong>21</strong>: <strong>30</strong>-<strong>06</strong>-<strong>2012</strong>. http://www.tardigrada.modena.unimo.<br />

it/miscellanea/<strong>Actual</strong>%20<strong>checklist</strong>%20<strong>of</strong>%20<strong>Tardigrada</strong>.pdf, pp. 36). Please write us if you<br />

find any mistake or missing data in this <strong>checklist</strong>. Your help in its improvement will be<br />

acknowledged.<br />

Acknowledgements<br />

We are very grateful to Pr<strong>of</strong>essor Clark Beasley (McMurry University, Abilene, U.S.A.) for his<br />

suggestions as well as for help with improving the English in the last version <strong>of</strong> the manuscript.<br />

We also thank to Nigel Marley (University <strong>of</strong> Plymouth, U.K.), Gary Grothman (St. Mary’s<br />

University, Calgary, Canada) and Paulo Fontoura (University <strong>of</strong> Porto, Portugal) for their<br />

valuable comments to this <strong>checklist</strong>.<br />

This study was partly supported by the Slovak Scientific Grant Agency (VEGA) (grant No.<br />

1/3265/<strong>06</strong> and 1/0294/09 to PD), by University <strong>of</strong> Modena and Reggio Emilia (50% FAR) and by<br />

Fondazione Cassa di Risparmio di Modena (MoDNA project: Morphology and DNA) to RB and<br />

RG.<br />

Checklist <strong>of</strong> <strong>Tardigrada</strong><br />

(The assignment <strong>of</strong> synonyms are reported within square parenthesis ‘[ ]’. Amendments are<br />

reported within angle brackets ‘ < > ’. Comments are reported within brace parenthesis ‘ { } ’.<br />

1

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The <strong>species</strong> written in bold characters are the genus type <strong>species</strong>)<br />

2<br />

TARDIGRADA<br />

HETEROTARDIGRADA Marcus, 1927<br />

ARTHROTARDIGRADA Marcus, 1927<br />

Archechiniscidae Binda, 1978 <br />

[Archechiniscinae Grimaldi de Zio & D’Addabbo Gallo, 1987]<br />

Archechiniscus Schulz, 1953<br />

Archechiniscus marci Schulz, 1953<br />

Archechiniscus minutus Grimaldi de Zio & D'Addabbo Gallo, 1987<br />

Archechiniscus symbalanus Chang & Rho, 1998<br />

Batillipedidae Ramazzotti, 1962<br />

[Discopodidae Marcus, 1934]<br />

Batillipes Richters, 1909<br />

Batillipes acaudatus Pollock, 1971<br />

Batillipes adriaticus Grimaldi de Zio, Morone De Lucia, D'Addabbo Gallo & Grimaldi, 1979<br />

Batillipes africanus Morone De Lucia, D’Addabbo Gallo & Grimaldi de Zio, 1988<br />

Batillipes annulatus de Zio, 1962<br />

Batillipes bullacaudatus McGinty & Higgins, 1968<br />

Batillipes carnonensis Fize, 1957<br />

Batillipes crassipes Tchesunov & Mokievsky, 1995<br />

Batillipes dicrocercus Pollock, 1970<br />

Batillipes friaufi Riggin, 1962<br />

Batillipes gilmartini McGinty, 1969<br />

Batillipes lesteri Kristensen & Mackness, 2000<br />

Batillipes littoralis littoralis Renaud-Debyser, 1959<br />

Batillipes littoralis submersus d'Hondt, 1970<br />

Batillipes longispinosus Chang & Rho, 1997<br />

Batillipes marcelli Morone De Lucia, D'Addabbo Gallo & Grimaldi de Zio, 1988<br />

Batillipes mirus Richters, 1909 [Batillipes caudatus Hay, 1917]<br />

Batillipes noerrevangi Kristensen, 1978<br />

Batillipes orientalis Chang & Rho, 1997<br />

Batillipes pennaki Marcus, 1946<br />

Batillipes philippinensis Chang & Rho, 1997<br />

Batillipes phreaticus Renaud Debyser, 1959 [Batillipes littoralis submersus D'Hondt, 1970]<br />

Batillipes rosc<strong>of</strong>fensis Kristensen, 1978<br />

Batillipes rotundiculus Rho, Min & Chang, 1999<br />

Batillipes similis Schulz, 1955<br />

Batillipes spinicauda Gallo D’Addabbo, Sandulli & de Zio Grimaldi, 2005<br />

Batillipes tridentatus Pollock, 1989<br />

Batillipes tubernatis Pollock, 1971

---

Coronarctidae Renaud-Mornant, 1974<br />

Coronarctus Renaud-Mornant, 1974<br />

Coronarctus disparilis Renaud-Mornant, 1987<br />

Coronarctus fastigatus Renaud-Mornant, 1987<br />

Coronarctus laubieri Renaud-Mornant, 1987<br />

Coronarctus mexicus Romano III, Gallo, D'Addabbo, Accogli, Baguley & Montagna, 2011<br />

Coronarctus stylisetus Renaud-Mornant, 1987<br />

Coronarctus tenellus Renaud-Mornant, 1974<br />

Coronarctus verrucatus Hansen, 2007<br />

Trogloarctus Villora-Moreno, 1996<br />

Trogloarctus trionyches Villora-Moreno, 1996<br />

Halechiniscidae Thulin, 1928<br />

Dipodarctinae Pollock, 1995<br />

Dipodarctus Pollock, 1995<br />

Dipodarctus anaholiensis Pollock, 1995<br />

Dipodarctus borrori Pollock, 1995 [Hemitanarctus chimaera de Zio Grimaldi, D'Addabbo Gallo,<br />

Morone De Lucia & Troccoli, 1995/96]<br />

Dipodarctus subterraneus (Renaud-Debyser, 1959)<br />

Euclavarctinae Renaud-Mornant, 1983<br />

Clavarctus Renaud-Mornant, 1983<br />

Clavarctus falculus Renaud-Mornant, 1983<br />

Euclavarctus Renaud-Mornant, 1975<br />

Euclavarctus convergens Renaud-Mornant, 1983<br />

Euclavarctus thieli Renaud-Mornant, 1975<br />

Exoclavarctus Renaud-Mornant, 1983<br />

Exoclavarctus dineti Renaud-Mornant, 1983<br />

Moebjergarctus Bussau, 1992<br />

Moebjergarctus manganis Bussau, 1992<br />

Parmursa Renaud-Mornant, 1984<br />

Parmursa fimbriata Renaud-Mornant, 1984<br />

Parmursa torquata Hansen, 2007<br />

Proclavarctus Renaud-Mornant, 1983<br />

Proclavarctus fragilis Renaud-Mornant, 1983<br />

3

---

4<br />

Florarctinae Renaud-Mornant, 1982<br />

Florarctus Delamare Deboutteville & Renaud-Mornant, 1965<br />

Florarctus acer Renaud-Mornant, 1989<br />

Florarctus antillensis Van der Land, 1968<br />

Florarctus asper Renaud-Mornant, 1989<br />

Florarctus cervinus Renaud-Mornant, 1987<br />

Florarctus cinctus Renaud-Mornant, 1976<br />

Florarctus glareolus Noda, 1987<br />

Florarctus heimi Delamare Deboutteville & Renaud-Mornant, 1965<br />

Florarctus hulingsi Renaud-Mornant, 1976<br />

Florarctus kwoni Chang & Rho, 1997<br />

Florarctus pulcher Grimaldi de Zio, Lamarca, D’addabbo Gallo & Pietanza, 1999<br />

Florarctus salvati Delamare Deboutteville & Renaud-Mornant, 1965<br />

Florarctus stellatus Renaud-Mornant, 1989<br />

Florarctus vulcanius Renaud-Mornant, 1987<br />

Ligiarctus Renaud-Mornant, 1982<br />

Ligiarctus eastwardi Renaud-Mornant, 1982<br />

Wingstrandarctus Kristensen, 1984<br />

Wingstrandarctus corallinus Kristensen, 1984<br />

Wingstrandarctus crypticus Renaud-Mornant, 1989<br />

Wingstrandarctus intermedius (Renaud-Mornant, 1967)<br />

Halechiniscinae (Thulin, 1928)<br />

Chrysoarctus Renaud-Mornant, 1984<br />

Chrysoarctus briandi Renaud-Mornant, 1984<br />

Chrysoarctus flabellatus (Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella &<br />

Grimaldi, 1982)<br />

Halechiniscus Richters, 1908<br />

Halechiniscus chafarinensis Grimaldi de Zio & Villora Moreno, 1995<br />

Halechiniscus greveni Renaud-Mornant & Deroux, 1976<br />

Halechiniscus guiteli Richters, 1908<br />

Halechiniscus jejuensis Chang & Rho, 2002<br />

Halechiniscus macrocephalus Grimaldi de Zio, D’Addabbo Gallo & Morone De Lucia, 1988<br />

Halechiniscus paratuleari Grimaldi de Zio, D’Addabbo Gallo & Morone De Lucia, 1988<br />

Halechiniscus perfectus Schulz, 1955<br />

Halechiniscus remanei remanei Schulz, 1955<br />

Halechiniscus remanei antillensis Renaud-Mornant, 1984<br />

Halechiniscus tuleari Renaud-Mornant, 1979<br />

Paradoxipus Kristensen & Higgins, 1989<br />

Paradoxipus orzeliscoides Kristensen & Higgins, 1989

---

Orzeliscinae (Schulz, 1963)<br />

Orzeliscus du Bois-Reymond Marcus, 1952<br />

Orzeliscus belopus du Bois-Reymond Marcus, 1952<br />

Orzeliscus septentrionalis Schulz, 1953<br />

Opydorscus Renaud-Mornant, 1989<br />

Opydorscus fonsecae Renaud-Mornant, 1989<br />

Styraconyxinae Kristensen & Renaud-Mornant, 1983<br />

Angursa Pollock, 1979<br />

Angursa antarctica Villora-Moreno, 1998<br />

Angursa bicuspis bicuspis Pollock, 1979<br />

Angursa bicuspis clavifera Noda, 1985<br />

Angursa bicuspis abyssalis Renaud-Mornant, 1981<br />

Angursa capsula Bussau, 1992<br />

Angursa lanceolata Renaud-Mornant, 1981<br />

Angursa lingua Bussau, 1992<br />

Bathyechiniscus Steiner, 1926<br />

Bathyechiniscus tetronyx Steiner, 1926<br />

Lepoarctus Kristensen & Renaud-Mornant, 1983<br />

Lepoarctus coniferus (Renaud-Mornant, 1975)<br />

Paratanarctus D'Addabbo Gallo, Grimaldi de Zio, Morone De Lucia & Troccoli, 1992<br />

Paratanarctus kristenseni D'Addabbo Gallo, Grimaldi de Zio, Morone De Lucia & Troccoli,<br />

1992<br />

Pleocola Cantacuzène, 1951<br />

Pleocola limnoriae Cantacuzène, 1951<br />

Raiarctus Renaud-Mornant, 1981<br />

Raiarctus aureolatus Renaud-Mornant, 1981<br />

Raiarctus colurus Renaud-Mornant, 1981<br />

Raiarctus variabilis D’Addabbo Gallo, Grimaldi de Zio & Morone De Lucia, 1986<br />

Rhomboarctus Renaud-Mornant, 1984<br />

Rhomboarctus aslaki Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003<br />

Rhomboarctus duplicicaudatus Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003<br />

Rhomboarctus thomassini Renaud-Mornant, 1984<br />

Styraconyx Thulin, 1942<br />

Styraconyx craticuliformis Chang & Rho, 1998<br />

Styraconyx craticulus (Pollock, 1983)<br />

Styraconyx hallasi Kristensen, 1977<br />

5

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Styraconyx haploceros Thulin, 1942<br />

Styraconyx kristenseni kristenseni Renaud-Mornant, 1981<br />

Styraconyx kristenseni neocaledonensis Renaud-Mornant, 1981<br />

Styraconyx nanoqsunguak Kristensen & Higgins, 1984<br />

Styraconyx paulae Robotti, 1971<br />

Styraconyx qivitoq Kristensen & Higgins, 1984<br />

Styraconyx sardiniae D'Addabbo Gallo, Morone De Lucia & Grimaldi de Zio, 1989<br />

Styraconyx sargassi Thulin, 1942<br />

Styraconyx testudo D'Addabbo Gallo, Grimaldi de Zio & Morone De Lucia, 1984<br />

Styraconyx tyrrhenus D'Addabbo Gallo, Morone De Lucia & Grimaldi de Zio, 1989<br />

Tetrakentron Cuénot, 1892<br />

Tetrakentron synaptae Cuénot, 1892<br />

Tholoarctus Kristensen & Renaud-Mornant, 1983<br />

Tholoarctus natans natans Kristensen & Renaud-Mornant, 1983<br />

Tholoarctus natans pedunculatus D'Addabbo Gallo, de Zio Grimaldi, Morone De Lucia &<br />

Troccoli, 1992<br />

6<br />

Tanarctinae Renaud-Mornant, 1980<br />

Actinarctus Schulz, 1935<br />

Actinarctus doryphorus doryphorus Schulz, 1935<br />

Actinarctus doryphorus ocellatus Renaud-Mornant, 1971<br />

Actinarctus lyrophorus Renaud-Mornant, 1979<br />

Actinarctus neretinus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella &<br />

Grimaldi, 1982<br />

Actinarctus physophorus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella &<br />

Grimaldi, 1982<br />

Tanarctus Renaud-Debyser, 1959<br />

Tanarctus arborspinosus Lindgren, 1971<br />

Tanarctus bubulubus Jørgensen & Kristensen, 2001<br />

Tanarctus dendriticus Renaud-Mornant, 1980<br />

Tanarctus gracilis Renaud-Mornant, 1980<br />

Tanarctus helleouetae Renaud-Mornant, 1984<br />

Tanarctus heterodactylus Renaud-Mornant, 1980<br />

Tanarctus longisetosus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella &<br />

Grimaldi, 1982<br />

Tanarctus minotauricus Renaud-Mornant, 1984<br />

Tanarctus ramazzottii Renaud-Mornant, 1975<br />

Tanarctus tauricus Renaud-Debyser, 1959<br />

Tanarctus velatus McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Zioella Renaud-Mornant, 1987<br />

Zioella pavonina Renaud-Mornant, 1987

---

Neoarctidae (de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992)<br />

Neoarctus de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992<br />

Neoarctus primigenius de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992<br />

Neostygarctidae de Zio Grimaldi, D’Addabbo Gallo & De Lucia Morone, 1987<br />

Neostygarctus Grimaldi de Zio, D'Addabbo Gallo & Morone De Lucia, 1982<br />

Neostygarctus acanthophorus Grimaldi de Zio, D'Addabbo Gallo & Morone De Lucia, 1982<br />

Renaudarctidae Kristensen & Higgins, 1984<br />

Renaudarctus Kristensen & Higgins, 1984<br />

Renaudarctus psammocryptus Kristensen & Higgins, 1984<br />

Stygarctidae Schulz, 1951<br />

Megastygarctidinae Bello & de Zio Grimaldi, 1998<br />

Megastygarctides McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Megastygarctides christinae Hansen & Kristensen, 20<strong>06</strong><br />

Megastygarctides gerdae Hansen & Kristensen, 20<strong>06</strong><br />

Megastygarctides isounguis Renaud-Mornant, 1981<br />

Megastygarctides orbiculatus McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Megastygarctides setoloso Morgan & O'Reilly, 1988<br />

Stygarctinae Schulz, 1951<br />

Parastygarctus Renaud-Debyser, 1965<br />

Parastygarctus biungulatus Morone De Lucia, Grimaldi de Zio & D'Addabbo Gallo, 1984<br />

Parastygarctus higginsi Renaud-Debyser, 1965<br />

Parastygarctus mediterranicus Gallo D’Addabbo, Grimaldi de Zio & Sandulli, 2001<br />

Parastygarctus renaudae Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo,<br />

1987<br />

Parastygarctus sterreri Renaud-Mornant, 1970<br />

Pseudostygarctus McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Pseudostygarctus apuliae Gallo D’Addabbo, de Zio Grimaldi & D’Addabbo, 2000<br />

Pseudostygarctus intermedius (Renaud-Mornant, 1979)<br />

Pseudostygarctus mirabilis de Zio Grimaldi, D’Addabbo Gallo & Morone De Lucia, 1998<br />

Pseudostygarctus rugosus Gallo D’Addabbo, de Zio Grimaldi & Sandulli, 2001<br />

Pseudostygarctus triungulatus McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Stygarctus Schulz, 1951<br />

Stygarctus abornatus McKirdy, Schmidt & McGinty-Bayly, 1976<br />

Stygarctus bradypus Schulz, 1951<br />

Stygarctus gourbaultae Renaud-Mornant, 1981<br />

7

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Stygarctus granulatus Pollock, 1970<br />

Stygarctus lambertii Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo, 1987<br />

Stygarctus spinifer Hiruta, 1985<br />

8<br />

ECHINISCOIDEA Richters, 1926<br />

Echiniscoididae Kristensen & Hallas, 1980<br />

Anisonyches Pollock, 1975<br />

Anisonyches deliquus Chang & Rho, 1998<br />

Anisonyches diakidius Pollock, 1975<br />

Anisonyches mauritianus Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo,<br />

1987<br />

Echiniscoides Plate, 1888<br />

Echiniscoides andamanensis Chang & Rho, 1998<br />

Echiniscoides bruni D'Addabbo Gallo, Grimaldi de Zio, Morone De Lucia & Troccoli, 1992<br />

Echiniscoides higginsi Hallas & Kristensen, 1982<br />

Echiniscoides hoepneri Kristensen & Hallas, 1980<br />

Echiniscoides horningi Miller & Kristensen, 1999<br />

Echiniscoides pollocki Hallas & Kristensen, 1982<br />

Echiniscoides sigismundi sigismundi (M. Schultze, 1865)<br />

Echiniscoides sigismundi galliensis Kristensen & Hallas, 1980<br />

Echiniscoides sigismundi groenlandicus Kristensen & Hallas, 1980<br />

Echiniscoides sigismundi hispaniensis Kristensen & Hallas, 1980<br />

Echiniscoides sigismundi mediterranicus Kristensen & Hallas, 1980<br />

Echiniscoides sigismundi polynesiensis Renaud-Mornant, 1976<br />

Echiniscoides sigismundi porphyrae Grimaldi de Zio, Gallo D’Addabbo & Pietanza, 2000<br />

Echiniscoides sigismundi verrucariae Grimaldi de Zio, Gallo D’Addabbo & Pietanza, 2000<br />

Echiniscoides travei Bellido & Bertrand, 1981<br />

Carphania fluviatilis Binda, 1978<br />

Carphaniidae Binda & Kristensen, 1986<br />

Carphania Binda, 1978<br />

Oreellidae Puglia, 1959<br />

Oreella Murray, 1910<br />

Oreella chugachii Calloway, Miller, Johansson & Whiting, 2011<br />

Oreella mollis Murray, 1910 [Oreella minor Ramazzotti, 1964]<br />

Oreella vilucensis Rahm, 1931 <strong>species</strong> inquirenda [Oreella bonnensis Rahm, 1932]<br />

Echiniscidae Thulin, 1928<br />

Antechiniscus Kristensen, 1987<br />

Antechiniscus conversus (Horning & Schuster, 1983)

---

Antechiniscus jermani Rossi & Claps, 1989<br />

Antechiniscus lateromamillatus (Ramazzotti, 1964)<br />

Antechiniscus moscali Claxton, 2001<br />

Antechiniscus parvisentus (Horning & Schuster, 1983)<br />

Antechiniscus perplexus (Horning & Schuster, 1983)<br />

Bryochoerus Marcus, 1936<br />

Bryochoerus intermedius intermedius (Murray, 1910)<br />

Bryochoerus intermedius hawaiicus (Thulin, 1928)<br />

Bryochoerus intermedius laevis (Marcus, 1936)<br />

Bryodelphax Thulin, 1928<br />

Bryodelphax aaseae Kristensen, Michalczyk & Kaczmarek, 2010<br />

Bryodelphax alzirae (du Bois-Reymond Marcus, 1944)<br />

Bryodelphax amphoterus (Durante Pasa & Maucci, 1975)<br />

Bryodelphax asiaticus Kaczmarek & Michalczyk, 2004<br />

Bryodelphax atlantis Fontoura, Pilato & Lisi, 2008<br />

Bryodelphax brevidentatus Kaczmarek, Michalczyk & Degma, 2005<br />

Bryodelphax crossotus Grigarick, Schuster & Nelson, 1983<br />

Bryodelphax dominicanus (Schuster & T<strong>of</strong>tner, 1982)<br />

Bryodelphax iohannis Bertolani, Guidi & Rebecchi, 1996<br />

Bryodelphax lijiangensis Yang, 2002<br />

Bryodelphax mateusi (Fontoura, 1982)<br />

Bryodelphax meronensis Pilato, Lisi & Binda, 2010<br />

Bryodelphax ortholineatus (Bartoš, 1963)<br />

Bryodelphax parvulus Thulin, 1928<br />

Bryodelphax parvuspolaris Kaczmarek, Zawierucha, Smykla & Michalczyk, <strong>2012</strong><br />

Bryodelphax sinensis (Pilato, 1974)<br />

Bryodelphax tatrensis (Węglarska, 1959)<br />

Bryodelphax weglarskae (Pilato, 1972)<br />

Cornechiniscus Maucci & Ramazzotti, 1981<br />

Cornechiniscus brachycornutus Maucci, 1987<br />

Cornechiniscus ceratophorus (Maucci, 1973)<br />

Cornechiniscus cornutus (Richters, 19<strong>06</strong>) [Pseudechiniscus intermedius Mihelčič, 1970]<br />

Cornechiniscus holmeni (Petersen, 1951)<br />

Cornechiniscus lobatus (Ramazzotti, 1943) [Pseudechiniscus cornutus Mihelčič, 1966]<br />

Cornechiniscus madagascariensis Maucci, 1993<br />

Cornechiniscus schrammi (Dastych, 1979)<br />

Cornechiniscus subcornutus Maucci & Ramazzotti, 1981<br />

Cornechiniscus tibetanus (Maucci, 1979)<br />

Echiniscus C.A.S. Schultze, 1840<br />

Echiniscus africanus Murray, 1907<br />

Echiniscus aliquantillus Grigarick, Schuster & Nelson, 1983<br />

Echiniscus angolensis da Cunha & do Nascimento Ribeiro, 1964<br />

Echiniscus apuanus M. Bertolani, 1946<br />

9

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Echiniscus arcangelii Maucci, 1973-74<br />

Echiniscus arctomys Ehrenberg, 1853<br />

Echiniscus arthuri Pilato, Binda & Lisi, 2005<br />

Echiniscus azoricus Fontoura, Pilato & Lisi, 2008<br />

Echiniscus baius Marcus, 1928<br />

Echiniscus baloghi Iharos, 1973<br />

Echiniscus barbarae Kaczmarek & Michalczyk, 2002<br />

Echiniscus batramiae Iharos, 1936<br />

Echiniscus becki Schuster & Grigarick, 1966<br />

Echiniscus bigranulatus Richters, 1907<br />

Echiniscus bisculptus Maucci, 1983<br />

Echiniscus blumi blumi Richters, 1903 [Echiniscus ramazzottii Binda & Pilato, 1969]<br />

Echiniscus blumi schiz<strong>of</strong>ilus Bartoš, 1941<br />

Echiniscus calcaratus Richters, 1908<br />

Echiniscus calvus Marcus, 1931<br />

Echiniscus canadensis Murray, 1910 [Echiniscus punctulatus Mihelčič, 1955; Echiniscus bellus<br />

Mihelčič, 1967]<br />

Echiniscus canedoi da Cunha & do Nascimento Ribeiro, 1962<br />

Echiniscus capillatus Ramazzotti, 1956<br />

Echiniscus carsicus Mihelčič, 1966<br />

Echiniscus carusoi Pilato, 1972<br />

Echiniscus cavagnaroi Schuster & Grigarick, 1966<br />

Echiniscus cervicornis Murray, 19<strong>06</strong><br />

Echiniscus charrua Claps & Rossi, 1997<br />

Echiniscus cheonyoungi Moon & Kim, 1994<br />

Echiniscus cirinoi Binda & Pilato, 1993<br />

Echiniscus clavispinosus Fontoura, Pilato & Lisi, 2011<br />

Echiniscus clevelandi Beasley, 1999<br />

Echiniscus columinis Murray, 1911<br />

Echiniscus corrugicaudatus McInnes, <strong>2009</strong><br />

Echiniscus crassispinosus crassispinosus Murray, 1907<br />

Echiniscus crassispinosus fasciatus Marcus, 1928<br />

Echiniscus curiosus Claxton, 1996<br />

Echiniscus dariae Kaczmarek & Michalczyk, 2010<br />

Echiniscus darienae Miller, Horning & Dastych, 1995<br />

Echiniscus dearmatus Bartoš, 1935 [Echiniscus szaboi Iharos, 1973]<br />

Echiniscus dikenli Maucci, 1973<br />

Echiniscus diploglyptus Durante Pasa & Maucci, 1975<br />

Echiniscus divergens Marcus, 1936<br />

Echiniscus dreyfusi de Barros, 1942<br />

Echiniscus duboisi Richters, 1902<br />

Echiniscus egnatiae Durante Pasa & Maucci, 1979<br />

Echiniscus ehrenbergi Dastych & Kristensen, 1995<br />

Echiniscus elaeinae Pilato, Binda & Lisi, 2005<br />

Echiniscus elegans Richters, 19<strong>06</strong><br />

Echiniscus evelinae de Barros, 1942<br />

Echiniscus filamentosus Plate, 1888<br />

10

---

Echiniscus ganczareki Michalczyk & Kaczmarek, 2007<br />

Echiniscus glaber Bartoš, 1937<br />

Echiniscus granulatus (Doyère, 1840) [Echiniscus crassus Richters, 1904; Echiniscus fortis<br />

Bartoš, 1935; Echiniscus abanti Maucci, 1973]<br />

Echiniscus heterospinosus Maucci, 1954<br />

Echiniscus hexacanthus Maucci, 1973<br />

Echiniscus hoonsooi Moon & Kim, 1990<br />

Echiniscus horningi Schuster & Grigarick, 1971<br />

Echiniscus inocelatus Mihelčič, 1938 [Echiniscus inocellatus Mihelčič, 1964]<br />

Echiniscus insuetus Mihelčič, 1967<br />

Echiniscus jagodici Mihelčič, 1951<br />

Echiniscus jamesi Claxton, 1996<br />

Echiniscus japonicus Morikawa, 1951<br />

Echiniscus jenningsi Dastych, 1984<br />

Echiniscus kerguelensis Richters, 1904<br />

Echiniscus knowltoni Schuster & Grigarick, 1971<br />

Echiniscus k<strong>of</strong>ordi Schuster & Grigarick, 1966<br />

Echiniscus kosickii Kaczmarek & Michalczyk, 2010<br />

Echiniscus lapponicus Thulin, 1911<br />

Echiniscus laterosetosus Ito, 1993<br />

Echiniscus laterospinosus Rudescu, 1964<br />

Echiniscus latifasciatus Dudichev & Biserov, 2000<br />

Echiniscus lichenorum Maucci, 1983<br />

Echiniscus limai da Cunha & do Nascimento Ribeiro, 1964<br />

Echiniscus lineatus Pilato, Fontoura, Lisi & Beasley, 2008<br />

Echiniscus longispinosus Murray, 1907<br />

Echiniscus loxophthalmus Richters, 1911<br />

Echiniscus madonnae Michalczyk & Kaczmarek, 20<strong>06</strong><br />

Echiniscus maesi Séméria, 1985<br />

Echiniscus malpighii Biserov, 1994<br />

Echiniscus manuelae da Cunha & do Nascimento Ribeiro, 1962<br />

Echiniscus marcusi Pilato, Claxton & Binda, 1989<br />

Echiniscus marginatus Binda & Pilato, 1994<br />

Echiniscus marginoporus Grigarick, Schuster & Nelson, 1983<br />

Echiniscus markezi Mihelčič, 1971/72<br />

Echiniscus marleyi X. Li, 2007<br />

Echiniscus mauccii Ramazzotti, 1956<br />

Echiniscus mediantus Marcus, 19<strong>30</strong><br />

Echiniscus merokensis merokensis Richters, 1904 [Echiniscus iharosi Rudescu, 1964]<br />

Echiniscus merokensis suecicus Thulin, 1911<br />

Echiniscus migiurtinus Franceschi, 1957<br />

Echiniscus mihelcici Iharos, 1973<br />

Echiniscus militaris Murray, 1911<br />

Echiniscus molluscorum Fox & Garcia-Moll, 1962<br />

Echiniscus moniliatus Iharos, 1967<br />

Echiniscus montanus Iharos, 1982<br />

Echiniscus mosaicus Grigarick, Schuster & Nelson, 1983<br />

11

---

Echiniscus multispinosus da Cunha, 1944<br />

Echiniscus murrayi Iharos, 1969<br />

Echiniscus nelsonae X. Li, L. Wang & Yu, 2007<br />

Echiniscus nepalensis Dastych, 1975<br />

Echiniscus nigripustulus Horning, Schuster & Grigarick, 1978<br />

Echiniscus nobilis Mihelčič, 1967<br />

Echiniscus oihonnae Richters, 1903<br />

Echiniscus ollantaytamboensis Nickel, Miller & Marley, 2001<br />

Echiniscus osellai Maucci, 1975<br />

Echiniscus pajstunensis Bartoš,1941<br />

Echiniscus palmai Dastych, 1997<br />

Echiniscus perarmatus Murray, 1907<br />

Echiniscus peruvianus Binda & Pilato, 1994<br />

Echiniscus perviridis Ramazzotti, 1959<br />

Echiniscus phocae du Bois-Reymond Marcus, 1944<br />

Echiniscus polygonalis Ito, 1993<br />

Echiniscus pooensis Rodriguez-Roda, 1948<br />

Echiniscus porabrus Horning, Schuster & Grigarick, 1978<br />

Echiniscus postojnensis Mihelčič, 1967 [Echiniscus loxophtalmus Mihelčič, 1939]<br />

Echiniscus pseudelegans Séméria, 1994<br />

Echiniscus pseudowendti Dastych, 1984<br />

Echiniscus punctus McInnes, 1995<br />

Echiniscus pusae Marcus, 1928<br />

Echiniscus quadrispinosus quadrispinosus Richters, 1902 [Echiniscus scr<strong>of</strong>a Richters, 1902]<br />

Echiniscus quadrispinosus brachyspinosus Bartoš, 1934<br />

Echiniscus quadrispinosus cribrosus Murray, 1907<br />

Echiniscus quadrispinosus fissispinosus Murray, 1907<br />

Echiniscus quitensis Pilato, 2007<br />

Echiniscus rackae Dastych, 1986<br />

Echiniscus ranzii Ramazzotti, 1964<br />

Echiniscus reticulatus Murray, 1905<br />

Echiniscus reymondi Marcus, 1928<br />

Echiniscus robertsi Schuster & Grigarick, 1965<br />

Echiniscus rodnae Claxton, 1996<br />

Echiniscus rufoviridis du Bois-Reymond Marcus, 1944<br />

Echiniscus rugospinosus Marcus, 1928<br />

Echiniscus scabrospinosus Fontoura, 1982 <br />

Echiniscus semifoveolatus Ito, 1993<br />

Echiniscus shaanxiensis X. Li, L. Wang & Yu, 2007<br />

Echiniscus siegristi Heinis, 1911<br />

Echiniscus simba Marcus, 1928<br />

Echiniscus speciosus Mihelčič, 1967 [Echiniscus roseus Mihelčič, 1967]<br />

Echiniscus spiculifer Schaudinn, 1901<br />

Echiniscus spiniger Richters, 1904<br />

Echiniscus spinulosus (Doyère, 1840)<br />

Echiniscus storkani Bartoš, 1940<br />

Echiniscus sylvanus Murray, 1910<br />

12

---

Echiniscus taibaiensis L. Wang & X. Li, 2005<br />

Echiniscus tamus Mehlen, 1969 <br />

Echiniscus tardus Mihelčič, 1951<br />

Echiniscus tenuis Marcus, 1928<br />

Echiniscus tessellatus Murray, 1910<br />

Echiniscus testudo (Doyère, 1840) [Echiniscus bellermanni C.A.S. Schultze, 1840; Echiniscus<br />

inermis Richters, 1902; Echiniscus trifilis Rahm, 19<strong>21</strong>; Echiniscus filamentosus<br />

mongoliensis Iharos, 1973]<br />

Echiniscus trisetosus Cuénot, 1932<br />

Echiniscus trojanus Maucci, 1973<br />

Echiniscus tropicalis Binda & Pilato, 1995<br />

Echiniscus tympanista Murray, 1911<br />

Echiniscus velaminis Murray, 1910<br />

Echiniscus vinculus Horning, Schuster & Grigarick, 1978<br />

Echiniscus virginicus Riggin, 1962 <br />

Echiniscus viridianus Pilato, Fontoura & Lisi, 2007<br />

Echiniscus viridis Murray, 1910 <br />

Echiniscus viridissimus Péterfi, 1956<br />

Echiniscus walteri Pilato & Lisi, 2003<br />

Echiniscus weisseri Maucci, 1978<br />

Echiniscus wendti Richters, 1903<br />

Echiniscus zetotrymus Horning, Schuster & Grigarick, 1978<br />

Hypechiniscus Thulin, 1928<br />

Hypechiniscus exarmatus (Murray, 1907) [Oreella breviclava Grigarick, Schuster & Nelson,<br />

1983]<br />

Hypechiniscus gladiator gladiator (Murray, 1905) [Parechiniscus unispinosus da Cunha, 1947]<br />

Hypechiniscus gladiator bigladii (Iharos, 1973)<br />

Hypechiniscus gladiator fissigladii (Iharos, 1973)<br />

Hypechiniscus gladiator spinulosa (Iharos, 1973)<br />

Hypechiniscus papillifer Robotti, 1972<br />

Mopsechiniscus du Bois-Reymond Marcus, 1944<br />

Mopsechiniscus frenoti Dastych, 1999<br />

Mopsechiniscus granulosus Mihelčič, 1967<br />

Mopsechiniscus imberbis (Richters, 1907)<br />

Mopsechiniscus schusteri Dastych, 1999<br />

Mopsechiniscus tasmanicus Dastych & Moscal, 1992<br />

Novechiniscus Kristensen, 1987<br />

Novechiniscus armadilloides (R.O. Schuster, 1975)<br />

Parechiniscus Cuénot, 1926<br />

Parechiniscus chitonides Cuénot, 1926<br />

Parechiniscus unispinosus da Cunha, 1947<br />

Proechiniscus Kristensen, 1987<br />

13

---

Proechiniscus hanneae (Petersen, 1951)<br />

Pseudechiniscus Thulin, 1911<br />

Pseudechiniscus alberti Dastych, 1987<br />

Pseudechiniscus asper Abe, Utsugi & Takeda, 1998<br />

Pseudechiniscus bartkei Węglarska, 1962<br />

Pseudechiniscus beasleyi X. Li, L. Wang & Yu, 2007<br />

Pseudechiniscus bidenticulatus Bartoš, 1963<br />

Pseudechiniscus bispinosus (Murray, 1907)<br />

Pseudechiniscus brevimontanus Kendall-Fite & Nelson, 1996<br />

Pseudechiniscus clavatus Mihelčič, 1955<br />

Pseudechiniscus conifer (Richters, 1904)<br />

Pseudechiniscus dicrani Mihelčič, 1938<br />

Pseudechiniscus distinctus Mihelčič, 1951<br />

Pseudechiniscus facettalis Petersen, 1951 [Pseudechiniscus pseudoconifer facettalis Maucci,<br />

1954]<br />

Pseudechiniscus goedeni Grigarick, Mihelčič & Schuster, 1964<br />

Pseudechiniscus gullii Pilato & Lisi, 20<strong>06</strong><br />

Pseudechiniscus insolitus Maucci, 1991<br />

Pseudechiniscus islandicus (Richters, 1904)<br />

Pseudechiniscus jiroveci Bartoš, 1963<br />

Pseudechiniscus juanitae de Barros, 1939 [Pseudechiniscus suillus franciscae de Barros, 1942]<br />

Pseudechiniscus jubatus Biserov, 1990<br />

Pseudechiniscus megacephalus Mihelčič, 1951<br />

Pseudechiniscus nataliae Biserov & Maucci, 1986<br />

Pseudechiniscus novaezeelandiae novaezeelandiae (Richters, 1908)<br />

Pseudechiniscus novaezeelandiae aspinosa Iharos, 1963<br />

Pseudechiniscus novaezeelandiae laterospinosa Iharos, 1963<br />

Pseudechiniscus novaezeelandiae marinae Bartoš, 1934<br />

Pseudechiniscus occultus Dastych, 1980<br />

Pseudechiniscus papillosus X. Li, L. Wang, Liu & Su, 2005<br />

Pseudechiniscus pilatoi X. Li, 2007<br />

Pseudechiniscus pseudoconifer Ramazzotti, 1943<br />

Pseudechiniscus pulcher (Murray, 1910)<br />

Pseudechiniscus quadrilobatus Iharos, 1969<br />

Pseudechiniscus ramazzottii ramazzottii Maucci, 1952<br />

Pseudechiniscus ramazzottii facettalis Iharos, 1964 [Pseudechiniscus ramazzottii lineatus<br />

Maucci, 1973-74]<br />

Pseudechiniscus raneyi Grigarick, Mihelčič & Schuster, 1964<br />

Pseudechiniscus santomensis Fontoura, Pilato & Lisi, 2010<br />

Pseudechiniscus scorteccii Franceschi, 1952<br />

Pseudechiniscus shilinensis Yang, 2002<br />

Pseudechiniscus sinensis Rahm, 1937<br />

Pseudechiniscus spinerectus Pilato, Binda, Napolitano & Moncada, 2001<br />

Pseudechiniscus suillus (Ehrenberg, 1853) [Echiniscus mutabilis Murray, 1905]<br />

Pseudechiniscus transsylvanicus Iharos, 1936<br />

Pseudechiniscus victor (Ehrenberg, 1853) [Pseudechiniscus tridentifer Bartoš, 1935 ]<br />

14

---

Pseudechiniscus yunnanensis L. Wang, <strong>2009</strong><br />

Testechiniscus Kristensen, 1987<br />

Testechiniscus laterculus (Schuster, Grigarick & T<strong>of</strong>tner, 1980)<br />

Testechiniscus macronyx (Richters, 1907)<br />

Testechiniscus meridionalis (Murray, 19<strong>06</strong>)<br />

Testechiniscus spitsbergensis (Scourfield, 1897) [Echiniscus clavisetosus Mihelčič, 1958;<br />

Echiniscus marinellae Bartoš, 1935; Echiniscus melanophthalmus Bartoš, 1936; Echiniscus<br />

menzeli Heinis, 1917; Echiniscus rosaliae Mihelčič, 1951; Echiniscus spinuloides Murray,<br />

1907]<br />

MESOTARDIGRADA Rahm, 1937<br />

THERMOZODIA Ramazzotti & Maucci, 1983<br />

Thermozodiidae Rahm, 1937<br />

Thermozodium Rahm, 1937<br />

Thermozodium esakii Rahm, 1937<br />

EUTARDIGRADA Richters 1926<br />

APOCHELA Schuster, Nelson, Grigarick & Christenberry, 1980<br />

Milnesiidae Ramazzotti, 1962<br />

Bergtrollus Dastych, 2011<br />

Bergtrollus dzimbowski Dastych, 2011<br />

Limmenius Horning, Schuster & Grigarick, 1978<br />

Limmenius porcellus Horning, Schuster & Grigarick, 1978<br />

Milnesioides Claxton, 1999<br />

Milnesioides exsertum Claxton, 1999<br />

Milnesium Doyère, 1840<br />

Milnesium alabamae Wallendorf & Miller, <strong>2009</strong><br />

Milnesium almatyense Tumanov, 20<strong>06</strong><br />

Milnesium antarcticum Tumanov, 20<strong>06</strong><br />

Milnesium asiaticum Tumanov, 20<strong>06</strong><br />

Milnesium brachyungue Binda & Pilato, 1990<br />

Milnesium dujiangensis Yang, 2003<br />

Milnesium eurystomum Maucci, 1991 <br />

Milnesium granulatum Ramazzotti, 1962 <br />

Milnesium jacobi Meyer & Hinton, 2010<br />

Milnesium katarzynae Kaczmarek, Michalczyk & Beasley, 2004<br />

15

---

Milnesium krzyszt<strong>of</strong>i Kaczmarek & Michalczyk, 2007<br />

Milnesium longiungue Tumanov, 20<strong>06</strong><br />

Milnesium reductum Tumanov, 20<strong>06</strong><br />

Milnesium reticulatum Pilato, Binda & Lisi, 2002<br />

Milnesium swolenskyi Bertolani & Grimaldi, 2000 {fossil <strong>species</strong>}<br />

Milnesium tardigradum tardigradum Doyère, 1840 <br />

Milnesium tardigradum trispinosa Rahm, 1931<br />

Milnesium tetralamellatum Pilato & Binda, 1991<br />

Milnesium zsalakoae Meyer & Hinton, 2010<br />

16<br />

PARACHELA Schuster, Nelson Grigarick & Christenberry, 1980<br />

Eohypsibioidea Bertolani & Kristensen, 1987 in Marley et al. 2011<br />

Eohypsibiidae Bertolani & Kristensen, 1987<br />

<br />

[Amphibolidae R. Bertolani, 1981]<br />

Austeruseus Trygvadóttir & Kristensen 2011<br />

Austeruseus balduri Trygvadóttir & Kristensen 2011<br />

Austeruseus faeroensis Trygvadóttir & Kristensen 2011<br />

Austeruseus rokuri Trygvadóttir & Kristensen 2011<br />

Bertolanius Özdikmen, 2008<br />

<br />

[Amphibolus R. Bertolani, 1981 in Özdikmen 2008]<br />

Bertolanius mahunkai (Iharos, 1971)<br />

Bertolanius markevichi (Biserov, 1992)<br />

Bertolanius nebulosus (Dastych, 1983)<br />

Bertolanius portucalensis Fontoura, Pilato, Lisi & Morais, <strong>2009</strong><br />

Bertolanius smreczynskii (Węglarska, 1970)<br />

Bertolanius volubilis (Durante Pasa & Maucci, 1975)<br />

Bertolanius weglarskae (Dastych, 1972)<br />

Eohypsibius Kristensen, 1982<br />

<br />

Eohypsibius nadjae Kristensen, 1982<br />

Eohypsibius terrestris Ito, 1988<br />

Hypsibioidea Pilato, 1969 in Marley et al. 2011<br />

Calohypsibiidae Pilato, 1969<br />

Calohypsibius Thulin, 1928<br />

Calohypsibius ornatus (Richters, 1900) [Hypsibius (Calohypsibius) armatus Bartoš, 1938;<br />

Hypsibius (Calohypsibius) intermedius Mihelčič, 1939]<br />

Calohypsibius maliki Michalczyk & Kaczmarek, 2005

---

Calohypsibius placophorus (da Cunha, 1943)<br />

Calohypsibius schusteri Nelson & McGlothlin, 1996<br />

Calohypsibius verrucosus (Richters, 1900) [Calohypsibius scabrosus Thulin, 1928]<br />

Haplohexapodibius Pilato & Beasley, 1987<br />

Haplohexapodibius seductor Pilato & Beasley, 1987 [Hexapodibius beasleyi Maucci, 1988]<br />

Haplomacrobiotus May, 1948<br />

Haplomacrobiotus hermosillensis May, 1948<br />

Haplomacrobiotus utahensis Pilato & Beasley, 2005<br />

Hexapodibius Pilato, 1969<br />

Hexapodibius bindae Pilato, 1982<br />

Hexapodibius boothi Dastych & McInnes, 1994<br />

Hexapodibius christenberryae Pilato & Binda, 2003<br />

Hexapodibius micronyx Pilato, 1969<br />

Hexapodibius pseudomicronyx Robotti, 1972<br />

Hexapodibius reginae Vargha, 1995<br />

Parhexapodibius Pilato, 1969<br />

Parhexapodibius bactrianus Biserov, 1999<br />

Parhexapodibius castrii (Ramazzotti, 1964)<br />

Parhexapodibius lagrecai (Binda & Pilato, 1969)<br />

Parhexapodibius pilatoi (Bernard, 1977)<br />

Parhexapodibius ramazzottii Manicardi & Bertolani, 1987<br />

Hypsibiidae Pilato, 1969<br />

Diphasconinae Dastych, 1992<br />

Diphascon (Diphascon) Plate, 1888<br />

Diphascon (Diphascon) aculeatum (Maucci, 1951-1952)<br />

Diphascon (Diphascon) alpinum Murray, 19<strong>06</strong><br />

Diphascon (Diphascon) australianum Pilato & Binda, 1998<br />

Diphascon (Diphascon) bidropion Ito, 1995<br />

Diphascon (Diphascon) birkleh<strong>of</strong>i Rolf Schuster, 1999<br />

Diphascon (Diphascon) bisbullatum (Iharos, 1964)<br />

Diphascon (Diphascon) boreale Biserov, 1996<br />

Diphascon (Diphascon) brevipes (Marcus, 1936)<br />

Diphascon (Diphascon) bullatum Murray, 1905<br />

Diphascon (Diphascon) burti Nelson, 1991<br />

Diphascon (Diphascon) chilenense Plate, 1888<br />

Diphascon (Diphascon) claxtonae Pilato & Binda, 1998<br />

Diphascon (Diphascon) dastychi Pilato & Binda, 1999<br />

Diphascon (Diphascon) dolomiticum Pilato & Bertolani, 2005<br />

Diphascon (Diphascon) elongatum (Mihelčič, 1959)<br />

Diphascon (Diphascon) faialense Fontoura & Pilato, 2007<br />

17

---

Diphascon (Diphascon) gerdae (Mihelčič, 1951)<br />

Diphascon (Diphascon) granifer Greven, 1972<br />

Diphascon (Diphascon) halapiense (Iharos, 1964)<br />

Diphascon (Diphascon) higginsi Binda, 1971<br />

Diphascon (Diphascon) humicus Bertolani, Guidetti & Rebecchi, 1994<br />

Diphascon (Diphascon) hydrophilum Pilato, Binda, Bertolani & Lisi, 2005<br />

Diphascon (Diphascon) iharosi Vargha, 1995<br />

Diphascon (Diphascon) iltisi (Schuster & Grigarick, 1965)<br />

Diphascon (Diphascon) langhovdense (Sudzuki, 1964)<br />

Diphascon (Diphascon) latipes (Mihelčič, 1955)<br />

Diphascon (Diphascon) mirabilis Dastych, 1984<br />

Diphascon (Diphascon) mitrense Pilato, Binda & Qualtieri, 1999<br />

Diphascon (Diphascon) nelsonae Pilato, Binda, Bertolani & Lisi, 2005<br />

Diphascon (Diphascon) nobilei (Binda, 1969)<br />

Diphascon (Diphascon) nodulosum (Ramazzotti, 1957)<br />

Diphascon (Diphascon) nonbullatum (Mihelčič, 1951)<br />

Diphascon (Diphascon) oculatum oculatum Murray, 19<strong>06</strong><br />

Diphascon (Diphascon) oculatum alpinum (Mihelčič, 1964)<br />

Diphascon (Diphascon) oculatum canadense Murray, 1910 [Hypsibius vancouverensis Thulin,<br />

1911]<br />

Diphascon (Diphascon) ongulense (Morikawa, 1962)<br />

Diphascon (Diphascon) opisthoglyptum Maucci, 1987<br />

Diphascon (Diphascon) patanei Binda & Pilato, 1971<br />

Diphascon (Diphascon) pingue pingue (Marcus, 1936)<br />

Diphascon (Diphascon) pingue brunsvicense Argue, 1972<br />

Diphascon (Diphascon) pinguiforme Pilato & Binda, 1997/98<br />

Diphascon (Diphascon) platyungue Pilato, Binda, Bertolani & Lisi, 2005<br />

Diphascon (Diphascon) polare Pilato & Binda, 1999<br />

Diphascon (Diphascon) puniceum (Jennings, 1976)<br />

Diphascon (Diphascon) ramazzottii (Robotti, 1970)<br />

Diphascon (Diphascon) recamieri Richters, 1911<br />

Diphascon (Diphascon) rugocaudatum (Rodriguez Roda, 1952)<br />

Diphascon (Diphascon) rugosum (Bartoš, 1935)<br />

Diphascon (Diphascon) sanae Dastych, Ryan & Watkins, 1990<br />

Diphascon (Diphascon) secchii Bertolani & Rebecchi, 1996<br />

Diphascon (Diphascon) serratum Pilato, Binda, Bertolani & Lisi, 2005<br />

Diphascon (Diphascon) sexbullatum Ito, 1995<br />

Diphascon (Diphascon) stappersi Richters, 1911<br />

Diphascon (Diphascon) tenue Thulin, 1928<br />

Diphascon (Diphascon) trachydorsatum (Bartoš, 1937)<br />

Diphascon (Diphascon) victoriae Pilato & Binda, 1999<br />

Diphascon (Diphascon) zaniewi Kaczmarek & Michalczyk, 2004<br />

Diphascon (Adropion) Pilato, 1987<br />

Diphascon (Adropion) arduifrons Thulin, 1928<br />

Diphascon (Adropion) behanae Dastych, 1987<br />

Diphascon (Adropion) belgicae Richters, 1911 [Fujiscon diphasconiellum Ito, 1991]<br />

18

---

Diphascon (Adropion) carolae Binda & Pilato, 1969<br />

Diphascon (Adropion) clavatum (Bartoš, 1935)<br />

Diphascon (Adropion) gordonense Pilato, Claxton & Horning, 1991<br />

Diphascon (Adropion) greveni Dastych, 1984<br />

Diphascon (Adropion) linzhiensis X. Li, 2007<br />

Diphascon (Adropion) mauccii Dastych & McInnes, 1996<br />

Diphascon (Adropion) modestum Binda, Pilato & Dastych, 1984<br />

Diphascon (Adropion) montigenum Pilato & Dastych, 1974<br />

Diphascon (Adropion) onorei Pilato, Binda, Napolitano & Moncada, 2002<br />

Diphascon (Adropion) prorsirostre Thulin, 1928<br />

Diphascon (Adropion) scoticum scoticum Murray, 1905 [Diphascon crozetense Richters, 1907]<br />

Diphascon (Adropion) scoticum ommatophorum (Thulin, 1911)<br />

Diphascon (Adropion) scoticum qinlingensis X. Li & Liu, 2005<br />

Diphascon (Adropion) tricuspidatum Binda & Pilato, 2000<br />

Diphascon (Adropion) triodon (Maucci, 1996)<br />

Diphascon bicorne (Mihelčič, 1971)<br />

Diphascon coniferens (Bartoš, 1960)<br />

Diphascon marcuzzii (Mihelčič, 1971)<br />

Diphascon mariae (Mihelčič, 1951)<br />

Diphascon punctatum (Iharos, 1962)<br />

Diphascon rivulare (Mihelčič, 1967)<br />

Diphascon speciosum (Mihelčič, 1971)<br />

Bindius triquetrus Pilato, <strong>2009</strong><br />

Bindius Pilato, <strong>2009</strong><br />

Paradiphascon Dastych, 1992<br />

Paradiphascon manningi Dastych, 1992<br />

Hypsibiinae Pilato, 1969<br />

Acutuncus Pilato & Binda, 1997<br />

Acutuncus antarcticus (Richters, 1904) [Hypsibius simoizumii Sudzuki, 1964]<br />

Borealibius Pilato, Guidetti, Rebecchi, Lisi, Hansen & Bertolani, 20<strong>06</strong><br />

Borealibius zetlandicus (Murray, 1907)<br />

Hypsibius Ehrenberg, 1848<br />

Hypsibius allisoni Horning, Schuster & Grigarick, 1978<br />

Hypsibius antonovae (Biserov, 1990)<br />

Hypsibius arcticus (Murray, 1907) [Macrobiotus heinisi Richters, 1907]<br />

Hypsibius biscuitiformis Bartoš, 1960<br />

Hypsibius calcaratus Bartoš, 1935<br />

Hypsibius camelopardalis Ramazzotti & Maucci, 1983<br />

Hypsibius choucoutiensis Rahm, 1937<br />

Hypsibius conifer Mihelčič, 1938<br />

19

---

Hypsibius convergens (Urbanowicz, 1925)<br />

Hypsibius dujardini (Doyère, 1840) [Macrobiotus palustris Dujardin, 1851; Macrobiotus<br />

lacustris Dujardin, 1851; Macrobiotus tetradactylus Lance, 1896; Macrobiotus murrayi<br />

Richters, 1907; Macrobiotus samoanus Richters, 1908; Macrobiotus breckneri Richters,<br />

1910; Macrobiotus ursellus Della Valle, 1915; Hypsibius murrayi Marcus, 1929]<br />

Hypsibius fuhrmanni (Heinis, 1914)<br />

Hypsibius giusepperamazzotti Sudzuki, 1975<br />

Hypsibius heardensis Miller, McInnes & Bergstrom, 2005<br />

Hypsibius hypostomus Bartoš, 1935<br />

Hypsibius iskandarovi Tumanov, 1997<br />

Hypsibius janetscheki Ramazzotti, 1968<br />

Hypsibius klebelsbergi Mihelčič, 1959<br />

Hypsibius kunmingensis Yang, 2002<br />

Hypsibius macrocalcaratus Beasley, 1988<br />

Hypsibius maculatus Iharos, 1969<br />

Hypsibius marcelli Pilato, 1990<br />

Hypsibius microps Thulin, 1928 <br />

Hypsibius montanus Iharos, 1940 [Hypsibius iharosi Bartoš, 1941]<br />

Hypsibius morikawai Ito, 1995<br />

Hypsibius multituberculatus Pilato, Binda & Lisi, 2003<br />

Hypsibius novaezeelandiae Pilato & Binda, 1997<br />

Hypsibius pachyunguis Maucci, 1996<br />

Hypsibius pallidus Thulin, 1911 <br />

Hypsibius pallidoides Pilato, Kiosya, Lisi, Inshina & Biserov, 2011<br />

Hypsibius pedrottii Bertolani, Manicardi & Gibertoni, 1987<br />

Hypsibius pradellii Bertolani & Rebecchi, 1996<br />

Hypsibius ragonesei Binda & Pilato, 1985<br />

Hypsibius roanensis Nelson & McGlothlin, 1993<br />

Hypsibius runae Bartoš, 1941<br />

Hypsibius scaber Maucci, 1987<br />

Hypsibius scabropygus Cuénot, 1929<br />

Hypsibius septulatus Pilato, Binda, Napolitano & Moncada, 2004<br />

Hypsibius seychellensis Pilato, Binda & Lisi, 20<strong>06</strong><br />

Hypsibius shaanxiensis H. Li & X. Li, 2008<br />

Hypsibius stiliferus Abe, 2004<br />

Hypsibius thaleri Dastych, 2004<br />

Hypsibius valentinae Pilato, Kiosya, Lisi & Sabella, <strong>2012</strong><br />

20<br />

Itaquasconinae Rudescu, 1964<br />

Astatumen Pilato, 1997<br />

Astatumen bartosi (Węglarska, 1959)<br />

Astatumen tamaensis (Sudzuki, 1975)<br />

Astatumen tamurai (Ito, 1990)<br />

Astatumen trinacriae (Arcidiacono, 1962) [Itaquascon ramazzottii Iharos, 1966]<br />

Itaquascon de Barros, 1939

---

Itaquascon biserovi Pilato, Binda & Moncada, 1999<br />

Itaquascon cambewarrense Pilato, Binda & Claxton, 2002<br />

Itaquascon enckelli (Mihelčič, 1971/72)<br />

Itaquascon globuliferum Abe & Ito, 1994<br />

Itaquascon mongolicus Kaczmarek, Michalczyk & Węglarska, 2002<br />

Itaquascon pawlowskii Węglarska, 1973<br />

Itaquascon pisoniae Pilato & Lisi, <strong>2009</strong><br />

Itaquascon placophorum Maucci, 1973<br />

Itaquascon simplex (Mihelčič, 1971)<br />

Itaquascon umbellinae de Barros, 1939<br />

Itaquascon unguiculum Pilato, Binda & Claxton, 2002<br />

Mesocrista Pilato, 1987<br />

Mesocrista marcusi (Rudescu, 1964)<br />

Mesocrista spitzbergensis (Richters, 1903)<br />

Parascon Pilato & Binda, 1987<br />

Parascon nichollsae Pilato & Lisi, 2004<br />

Parascon schusteri Pilato & Binda, 1987<br />

Platicrista Pilato, 1987<br />

Platicrista affine (Mihelčič, 1951)<br />

Platicrista angustata (Murray, 1905)<br />

Platicrista cheleusis Kathman, 1990 [Diphascon craigi Beasley, 1990]<br />

Platicrista horribilis Kaczmarek & Michalczyk, 2003<br />

Platicrista itaquasconoide (Durante Pasa & Maucci, 1975)<br />

Platicrista ramsayi Marley, 20<strong>06</strong><br />

Microhypsibiidae Pilato, 1998<br />

Fractonotus Pilato, 1998<br />

Fractonotus caelatus (Marcus, 1928)<br />

Microhypsibius Thulin, 1928<br />

Microhypsibius bertolanii Kristensen, 1982<br />

Microhypsibius japonicus Ito, 1991<br />

Microhypsibius minimus Kristensen, 1982<br />

Microhypsibius truncatus Thulin, 1928<br />

Ramazzottidae Marley, McInnes & Sands, 2011<br />

Hebesuncus Pilato, 1987<br />

Hebesuncus conjungens (Thulin, 1911)<br />

Hebesuncus ryani Dastych & Harris, 1994<br />

Hebesuncus schusteri (Dastych, 1984)<br />

Ramazzottius Binda & Pilato, 1986<br />

<strong>21</strong>

---

Ramazzottius affinis Bertolani, Guidetti & Rebecchi, 1994<br />

Ramazzottius agannae Dastych, 2011<br />

Ramazzottius andreevi Biserov, 1997/98<br />

Ramazzottius anomalus (Ramazzotti, 1962)<br />

Ramazzottius baumanni (Ramazzotti, 1962)<br />

Ramazzottius belubellus Bartels, Nelson, Kaczmarek & Michalczyk, 2011<br />

Ramazzottius bunikowskae Kaczmarek, Michalczyk & Diduszko, 20<strong>06</strong><br />

Ramazzottius cataphractus (Maucci, 1974)<br />

Ramazzottius caucasicus Biserov, 1997/98<br />

Ramazzottius edmondabouti Séméria, 1993<br />

Ramazzottius horningi Binda & Pilato, 1994<br />

Ramazzottius ljudmilae Biserov, 1997/98<br />

Ramazzottius montivagus (Dastych, 1983)<br />

Ramazzottius nivalis Dastych, 20<strong>06</strong><br />

Ramazzottius novemcinctus (Marcus, 1936)<br />

Ramazzottius oberhaeuseri (Doyère, 1840)<br />

Ramazzottius rupeus Biserov, 1999<br />

Ramazzottius saltensis (Claps & Rossi, 1984)<br />

Ramazzottius semisculptus Pilato & Rebecchi, 1992<br />

Ramazzottius subanomalus (Biserov, 1985)<br />

Ramazzottius szeptycki (Dastych, 1980) <br />

Ramazzottius theroni Dastych, 1993<br />

Ramazzottius thulini (Pilato, 1970)<br />

Ramazzottius tribulosus Bertolani & Rebecchi, 1988<br />

Ramazzottius valaamis Biserov & Tumanov, 1993<br />

Ramazzottius varieornatus Bertolani & Kinchin, 1993<br />

22<br />

Isohypsibioidea Marley, McInnes & Sands, 2011<br />

Isohypsibiidae Marley, McInnes & Sands, 2011<br />

Doryphoribius Pilato, 1969<br />

Doryphoribius amazzonicus Lisi, 2011<br />

Doryphoribius barbarae Beasley & Miller, <strong>2012</strong><br />

Doryphoribius bertolanii Beasley & Pilato, 1987<br />

Doryphoribius bindae Lisi, 2011<br />

Doryphoribius dawkinsi Michalczyk & Kaczmarek, 2010<br />

Doryphoribius doryphorus (Binda & Pilato, 1969)<br />

Doryphoribius dupliglobulatus Ito, 1995<br />

Doryphoribius evelinae (Marcus, 1928)<br />

Doryphoribius flavus (Iharos, 1966) [Doryphoribius citrinus (Maucci, 1973)] <br />

Doryphoribius gibber Beasley & Pilato, 1987<br />

Doryphoribius huangguoshuensis H. Wang, L. Wang & X. Li, 2007<br />

Doryphoribius koreanus Moon, Kim & Bertolani, 1994<br />

Doryphoribius korganovae Biserov, 1994<br />

Doryphoribius longistipes Bartels, Nelson, Kaczmarek & Michalczyk, 2008

---

Doryphoribius macrodon Binda, Pilato & Dastych, 1980<br />

Doryphoribius maranguensis Binda & Pilato, 1995<br />

Doryphoribius mariae Pilato & Binda, 1990<br />

Doryphoribius mexicanus Beasley, Kaczmarek & Michalczyk, 2008<br />

Doryphoribius minimus Bartels, Nelson, Kaczmarek & Michalczyk, 2008<br />

Doryphoribius neglectus Pilato & Lisi, 2004<br />

Doryphoribius picoensis Fontoura, Pilato & Lisi, 2008<br />

Doryphoribius pilatoi R. Bertolani, 1984<br />

Doryphoribius polynettae Biserov, 1988<br />

Doryphoribius qinlingense X. Li, Su & Yu, 2004<br />

Doryphoribius quadrituberculatus Kaczmarek & Michalczyk, 2004<br />

Doryphoribius smokiensis Bartels, Nelson, Kaczmarek & Michalczyk, 2007<br />

Doryphoribius solidunguis Lisi, 2011<br />

Doryphoribius taiwanus X. Li & H. Li, 2008<br />

Doryphoribius tergumrudis Bartels, Nelson, Kaczmarek & Michalczyk, 2008<br />

Doryphoribius tessellatus Meyer, 2011<br />

Doryphoribius turkmenicus Biserov, 1999<br />

Doryphoribius vietnamensis (Iharos, 1969)<br />

Doryphoribius zappalai Pilato, 1971<br />

Doryphoribius zyxiglobus (Horning, Schuster & Grigarick, 1978) <br />

Eremobiotus Biserov, 1992<br />

Eremobiotus alicatai (Binda, 1969)<br />

Eremobiotus ovezovae Biserov, 1992<br />

Halobiotus Kristensen, 1982<br />

Halobiotus arcturulius Crisp & Kristensen, 1983<br />

Halobiotus crispae Kristensen, 1982<br />

Halobiotus stenostomus (Richters, 1908) [Macrobiotus appelloefi Richters, 1908; Hypsibius<br />

(Isohypsibius) geddesi Hallas, 1971]<br />

Isohypsibius Thulin, 1928<br />

Isohypsibius altai Kaczmarek & Michalczyk, 20<strong>06</strong><br />

Isohypsibius annulatus annulatus (Murray, 1905)<br />

Isohypsibius annulatus minor (Ramazzotti, 1945)<br />

Isohypsibius arbiter Binda, 1980<br />

Isohypsibius archangajensis Kaczmarek & Michalczyk, 2004<br />

Isohypsibius arcuatus (Bartoš, 1934)<br />

Isohypsibius asper (Murray, 19<strong>06</strong>)<br />

Isohypsibius austriacus (Iharos, 1966)<br />

Isohypsibius baicalensis (Ramazzotti, 1966)<br />

Isohypsibius baldii (Ramazzotti, 1945)<br />

Isohypsibius baldiioides Tumanov, 2003<br />

Isohypsibius barbarae Pilato & Binda, 2002<br />

Isohypsibius bartosi (Iharos, 1966)<br />

Isohypsibius basalovoi (Durante & Maucci, 1973)<br />

23

---

Isohypsibius belliformis (Mihelčič, 1971)<br />

Isohypsibius bellus (Mihelčič, 1971)<br />

Isohypsibius borkini Tumanov, 2003<br />

Isohypsibius brevispinosus (Iharos, 1966)<br />

Isohypsibius brevitubulatus Rho, Chang & Kim, 1997<br />

Isohypsibius brulloi Pilato & Pennisi, 1976<br />

Isohypsibius bulbifer (Mihelčič, 1957)<br />

Isohypsibius cameruni (Iharos, 1969)<br />

Isohypsibius campbellensis Pilato, 1996<br />

Isohypsibius canadensis (Murray, 1910)<br />

Isohypsibius ceciliae Pilato & Binda, 1987<br />

Isohypsibius changbaiensis Yang, 1999<br />

Isohypsibius chiarae Maucci, 1987 [Isohypsibius bertolanii Manicardi, 1989]<br />

Isohypsibius costatus (Mihelčič, 1971)<br />

Isohypsibius coulsoni Kaczmarek, Zawierucha, Smykla & Michalczyk, <strong>2012</strong><br />

Isohypsibius cyrilli (Mihelčič, 1951)<br />

Isohypsibius damxungensis Yang, 2007<br />

Isohypsibius dastychi Pilato, Bertolani & Binda, 1982<br />

Isohypsibius deconincki Pilato, 1971<br />

Isohypsibius deflexus (Mihelčič, 1960)<br />

Isohypsibius dudichi (Iharos, 1964)<br />

Isohypsibius duranteae (Maucci, 1978)<br />

Isohypsibius effusus (Mihelčič, 1971)<br />

Isohypsibius elegans Binda & Pilato, 1971<br />

Isohypsibius eplenyiensis (Iharos, 1970)<br />

Isohypsibius franzi (Mihelčič, 1951)<br />

Isohypsibius fuscus (Mihelčič, 1971/72)<br />

Isohypsibius gilvus Biserov, 1986<br />

Isohypsibius glaber (Durante Pasa & Maucci, 1979)<br />

Isohypsibius glazovi Biserov, 1999<br />

Isohypsibius gracilis (Iharos, 1966)<br />

Isohypsibius granditintinus Chang & Rho, 1996<br />

Isohypsibius granulifer granulifer Thulin, 1928<br />

Isohypsibius granulifer koreanensis (Iharos, 1971)<br />

Isohypsibius gyulai (Mihelčič, 1971)<br />

Isohypsibius hadzii (Mihelčič, 1938)<br />

Isohypsibius helenae (Iharos, 1964)<br />

Isohypsibius hydrogogianus Ito & Tagami, 1993<br />

Isohypsibius hypostomoides (Mihelčič, 1971)<br />

Isohypsibius improvisus Dastych, 1984<br />

Isohypsibius indicus (Murray, 1907)<br />

Isohypsibius irregibilis Biserov, 1992<br />

Isohypsibius jakieli Dastych, 1984<br />

Isohypsibius jingshanensis Yang, 2003<br />

Isohypsibius jinhouensis Yang, 2007<br />

Isohypsibius josephi (Iharos, 1964)<br />

Isohypsibius kenodontis Kendall-Fite & Nelson, 1996<br />

24

---

Isohypsibius kotovae Tumanov, 2003<br />

Isohypsibius kristenseni Pilato, Catanzaro & Binda, 1989<br />

Isohypsibius ladogensis Tumanov, 2003<br />

Isohypsibius laevis McInnes, 1995<br />

Isohypsibius latiunguis (Iharos, 1964)<br />

Isohypsibius leithaicus (Iharos, 1966)<br />

Isohypsibius liae X. Li & L. Wang, 20<strong>06</strong><br />

Isohypsibius lineatus (Mihelčič, 1969)<br />

Isohypsibius longiunguis Pilato, 1974<br />

Isohypsibius lunulatus (Iharos, 1966)<br />

Isohypsibius macrodactylus (Maucci, 1978) [Isohypsibius zierh<strong>of</strong>feri Dastych, 1980 ]<br />

Isohypsibius malawiensis Jørgensen, 2001<br />

Isohypsibius mammillosus (Iharos, 1964)<br />

Isohypsibius marcellinoi Binda & Pilato, 1971<br />

Isohypsibius marii R. Bertolani, 1982<br />

Isohypsibius mihelcici (Iharos, 1964)<br />

Isohypsibius monoicus R. Bertolani, 1982<br />

Isohypsibius monstruosus Maucci, 1991<br />

Isohypsibius montanus Mihelčič, 1938<br />

Isohypsibius myrops (du Bois-Reymond Marcus, 1944)<br />

Isohypsibius neoundulatus (Durante Pasa & Maucci, 1975)<br />

Isohypsibius nipponicus Sudzuki, 1975<br />

Isohypsibius nodosus (Murray, 1907)<br />

Isohypsibius novaeguineae (Iharos, 1967)<br />

Isohypsibius palmai Pilato, 1996<br />

Isohypsibius panovi Tumanov, 2005<br />

Isohypsibius papillifer papillifer (Murray, 1905)<br />

Isohypsibius papillifer bulbosus (Marcus, 1928)<br />

Isohypsibius papillifer indicus (Iharos, 1969)<br />

Isohypsibius pappi (Iharos, 1966)<br />

Isohypsibius pauper (Mihelčič, 1971)<br />

Isohypsibius pilatoi (Durante Pasa & Maucci, 1979)<br />

Isohypsibius pratensis (Iharos, 1964)<br />

Isohypsibius prosostomus prosostomus Thulin, 1928<br />

Isohypsibius prosostomus cambrensis (Morgan, 1976)<br />

Isohypsibius pseudoundulatus (da Cunha & do Nascimento Ribeiro, 1964)<br />

Isohypsibius pulcher (Mihelčič, 1971/72)<br />

Isohypsibius pushkini Tumanov, 2003<br />

Isohypsibius qinlingensis X. Li, L. Wang & Yu, 2005<br />

Isohypsibius rahmi X. Li & L. Wang, 20<strong>06</strong><br />

Isohypsibius reticulatus Pilato, 1973<br />

Isohypsibius roberti Biserov, 1996<br />

Isohypsibius ronsisvallei Binda & Pilato, 1969<br />

Isohypsibius rudescui (Iharos, 1966)<br />

Isohypsibius rugosus Guidi & Grabowski, 1996<br />

Isohypsibius sabellai Pilato, Binda, Napolitano & Moncada, 2004<br />

25

---

Isohypsibius sattleri (Richters, 1902) [Hypsibius bakonyiensis Iharos, 1964] {the correct<br />

reference <strong>of</strong> Dastych 1990 cited in the previous check lists is: Dastych, H. (1991)<br />

Isohypsibius sattleri (Richters 1902), a valid <strong>species</strong> (<strong>Tardigrada</strong>). Senckenbergiana<br />

biologica, 71, 181–189.}<br />

Isohypsibius schaudinni (Richters, 1909)<br />

Isohypsibius sculptus (Ramazzotti, 1962)<br />

Isohypsibius sellnicki (Mihelčič, 1962)<br />

Isohypsibius septentrionalis Thulin, 1928<br />

Isohypsibius silvicola (Iharos, 1966)<br />

Isohypsibius sismicus (Maucci, 1978)<br />

Isohypsibius solidus (Mihelčič, 1971)<br />

Isohypsibius taibaiensis X. Li & L. Wang, 2005<br />

Isohypsibius tetradactyloides (Richters, 1907)<br />

Isohypsibius theresiae (Iharos, 1964)<br />

Isohypsibius torulosus (Mihelčič, 1959)<br />

Isohypsibius truncorum (Iharos, 1964)<br />

Isohypsibius tuberculatus (Plate, 1888)<br />

Isohypsibius tuberculoides (Mihelčič, 1951)<br />

Isohypsibius tubereticulatus Pilato & Catanzaro, 1989<br />

Isohypsibius tucumanensis Claps & Rossi, 1984<br />

Isohypsibius undulatus Thulin, 1928<br />

Isohypsibius vejdovskyi (Bartoš, 1939)<br />

Isohypsibius verae Pilato & Catanzaro, 1989<br />

Isohypsibius verrucosus (Della Valle, 1915) [Isohypsibius gibbus (Marcus, 1928)]<br />

Isohypsibius wilsoni (Horning, Schuster & Grigarick, 1978)<br />

Isohypsibius woodsae Kathman, 1990<br />

Isohypsibius yunnanensis Yang, 2002<br />

Mixibius Pilato, 1992<br />

{to be confirmed in the family Isohypsibiidae, see Marley et al. 2011}<br />

Mixibius fueginus Pilato & Binda, 1996<br />

Mixibius ninguidus Biserov, 1999<br />

Mixibius ornatus Pilato, Binda, Napolitano & Moncada, 2002<br />

Mixibius pilatoi L. Wang, <strong>2009</strong><br />

Mixibius saracenus (Pilato, 1973)<br />

Mixibius schnurae Pilato, Lisi & Binda, 2010<br />

Mixibius sutirae Pilato, Binda & Lisi 2004<br />

Mixibius tibetanus H. Li & X. Li, 2008<br />

Pseudobiotus Nelson, 1980<br />

Pseudobiotus hirsutellus Pilato, Lisi & Binda, 2010<br />

Pseudobiotus kathmanae Nelson, Marley & Bertolani, 1999<br />

Pseudobiotus longiunguis (Iharos, 1968)<br />

Pseudobiotus matici (Pilato, 1971)<br />

Pseudobiotus megalonyx (Thulin, 1928)<br />

Pseudobiotus spinifer Chang, Kaczmarek, Lee & Michalczyk, 2007<br />

Pseudobiotus vladimiri Biserov, Dudichev & Biserova, 2001 <br />

26

---

Ramajendas Pilato & Binda, 1990<br />

Ramajendas frigidus Pilato & Binda, 1990<br />

Ramajendas heatwolei Miller, Horning & Dastych, 1995<br />

Ramajendas renaudi (Ramazzotti, 1972)<br />

Thalerius Dastych, <strong>2009</strong><br />

{to be confirmed in the family Isohypsibiidae, see Marley et al. 2011}<br />

Thalerius konradi Dastych, <strong>2009</strong><br />

Thulinius R. Bertolani, 2003<br />

[Thulinia R. Bertolani, 1982]<br />

Thulinius augusti (Murray, 1907) [Macrobiotus lacustris Wenck, 1914]<br />

Thulinius itoi (Tsurusaki, 1980)<br />

Thulinius ruffoi (R. Bertolani, 1982)<br />

Thulinius saltursus (Schuster, T<strong>of</strong>tner & Grigarick, 1978) <br />

Thulinius stephaniae (Pilato, 1974)<br />

Macrobiotoidea Thulin, 1928 in Marley et al. 2011<br />

Macrobiotidae Thulin, 1928<br />

Adorybiotus Maucci & Ramazzotti, 1981<br />

Adorybiotus granulatus (Richters, 1903)<br />

Biserovus Guidetti & Pilato, 2003<br />

Biserovus bindae (Christenberry & Higgins, 1979)<br />

Calcarobiotus (Calcarobiotus) Dastych, 1993<br />

Calcarobiotus (Calcarobiotus) digeronimoi Pilato, Binda & Lisi, 2004<br />

Calcarobiotus (Calcarobiotus) filmeri Dastych, 1993<br />

Calcarobiotus (Calcarobiotus) gildae (Maucci & Durante Pasa, 1980)<br />

Calcarobiotus (Calcarobiotus) hainanensis X. Li, D. Wang & L. Wang, 2008<br />

Calcarobiotus (Calcarobiotus) imperialis Abe & Takeda, 2000<br />

Calcarobiotus (Calcarobiotus) longinoi Kaczmarek, Michalczyk & Guidetti, 20<strong>06</strong><br />

Calcarobiotus (Calcarobiotus) occultus Dastych, 1993<br />

Calcarobiotus (Calcarobiotus) parvicalcar Pilato & Lisi, <strong>2009</strong><br />

Calcarobiotus (Discrepunguis) Guidetti & Bertolani, 2001<br />

Calcarobiotus (Discrepunguis) polygonatus (Binda & Guglielmino, 1991) [Macrobiotus<br />

eugranulatus Maucci, 1993]<br />

Calcarobiotus (Discrepunguis) tetrannulatus Pilato, Binda & Lisi, 2004<br />

Famelobiotus Pilato, Binda & Lisi, 2004<br />

Famelobiotus scalicii Pilato, Binda & Lisi, 2004<br />

27

---

Insuetifurca Guidetti & Pilato, 2003<br />

Insuetifurca arrowsmithi (Kathman & Nelson, 1989)<br />

Insuetifurca austronipponica Abe, 2005<br />

Insuetifurca fujiense (Ito, 1997)<br />

Insuetifurca xiae (X. Li, Su & Yu, 2004) <br />

Macrobiotus C.A.S. Schultze, 1834<br />

Macrobiotus almadai Fontoura, Pilato & Lisi, 2008<br />

Macrobiotus altitudinalis Biserov, 1997/98<br />

Macrobiotus alvaroi Pilato & Kaczmarek, 2007<br />

Macrobiotus anderssoni Richters, 1907<br />

Macrobiotus andinus Maucci, 1988<br />

Macrobiotus annae Richters, 1908<br />

Macrobiotus aradasi Binda, Pilato & Lisi, 2005<br />

Macrobiotus arguei Pilato & Sperlinga, 1975<br />

Macrobiotus ariekammensis Węglarska, 1965 [Macrobiotus adelges Dastych, 1977]<br />

Macrobiotus armatus Pilato & Binda, 1996<br />

Macrobiotus artipharyngis Iharos, 1940<br />

Macrobiotus ascensionis Richters, 1908<br />

Macrobiotus australis Pilato & D'Urso, 1976<br />

Macrobiotus baltatus McInnes, 1991<br />

Macrobiotus barabanovi Tumanov, 2005<br />

Macrobiotus barbarae Kaczmarek, Michalczyk & Degma, 2007<br />

Macrobiotus binieki Kaczmarek, Gołdyn, Prokop & Michalczyk, 2011<br />

Macrobiotus biserovi Bertolani, Guidi & Rebecchi, 1996<br />

Macrobiotus blocki Dastych, 1984<br />

Macrobiotus brevipes Mihelčič, 1971/72<br />

Macrobiotus caelicola Kathman, 1990<br />

Macrobiotus carsicus Maucci, 1954<br />

Macrobiotus caymanensis Meyer, 2011<br />

Macrobiotus contii Pilato & Lisi, 20<strong>06</strong><br />

Macrobiotus coronatus de Barros, 1942<br />

Macrobiotus creber Pilato & Lisi, <strong>2009</strong><br />

Macrobiotus crenulatus Richters, 1904 [Macrobiotus dentatus Binda, 1974]<br />

Macrobiotus danielisae Pilato, Binda & Lisi, 20<strong>06</strong><br />

Macrobiotus dariae Pilato & Bertolani, 2004<br />

Macrobiotus denticulus Dastych, 2002<br />

Macrobiotus diffusus Binda & Pilato, 1987<br />

Macrobiotus diguensis Pilato & Lisi, <strong>2009</strong><br />

Macrobiotus dimentmani Pilato, Lisi & Binda, 2010<br />

Macrobiotus divergens Binda, Pilato & Lisi, 2005<br />

Macrobiotus diversus Biserov, 1990<br />

Macrobiotus drakensbergi Dastych, 1993<br />

Macrobiotus echinogenitus Richters, 1904<br />

Macrobiotus erminiae Binda & Pilato, 1999<br />

28

---

Macrobiotus evelinae de Barros, 1938<br />

Macrobiotus furciger Murray, 1907 [Macrobiotus ehrenbergi Heinis, 19<strong>21</strong>; Macrobiotus furcatus<br />

Murray, 19<strong>06</strong>}<br />

Macrobiotus gemmatus Bartoš, 1963<br />

Macrobiotus glebkai Biserov, 1990<br />

Macrobiotus grandis Richters, 1911<br />

Macrobiotus hapukuensis Pilato, Binda & Lisi, 20<strong>06</strong><br />

Macrobiotus harmsworthi harmsworthi Murray, 1907<br />

Macrobiotus harmsworthi obscurus Dastych, 1985<br />

Macrobiotus hibiscus de Barros, 1942<br />

Macrobiotus hieronimi Pilato & Claxton, 1988<br />

Macrobiotus hufelandi hufelandi C.A.S. Schultze, 1833<br />

Macrobiotus hufelandi maculatus Iharos, 1973<br />

Macrobiotus humilis Binda & Pilato, 2001<br />

Macrobiotus hyperboreus Biserov, 1990<br />

Macrobiotus iharosi Pilato, Binda & Catanzaro, 1991<br />

Macrobiotus insignis Bartoš, 1963<br />

Macrobiotus insularis Pilato, 20<strong>06</strong><br />

Macrobiotus islandicus islandicus Richters, 1904 [Macrobiotus ruffoi Maucci, 1973]<br />

Macrobiotus islandicus nicaraguensis Séméria, 1985<br />

Macrobiotus joannae Pilato & Binda, 1983<br />

Macrobiotus kazmierskii Kaczmarek & Michalczyk, <strong>2009</strong><br />

Macrobiotus kirghizicus Tumanov, 2005<br />

Macrobiotus kolleri Mihelčič, 1951<br />

Macrobiotus komareki Bartoš, 1939<br />

Macrobiotus kovalevi Tumanov, 2004<br />

Macrobiotus krynauwi Dastych & Harris, 1995<br />

Macrobiotus kurasi Dastych, 1981<br />

Macrobiotus lazzaroi Maucci, 1986 [Macrobiotus spallanzanii Maucci, 1973]<br />

Macrobiotus lissostomus Durante Pasa & Maucci, 1979<br />

Macrobiotus liviae Ramazzotti, 1962<br />

Macrobiotus longipes Mihelčič, 1971/72<br />

Macrobiotus lusitanicus Maucci & Durante Pasa, 1984<br />

Macrobiotus macrocalix Bertolani & Rebecchi, 1993<br />

Macrobiotus madegassus Maucci, 1993<br />

Macrobiotus mandalaae Pilato, 1974<br />

Macrobiotus marlenae Kaczmarek & Michalczyk, 2004<br />

Macrobiotus martini Bartels, Pilato, Lisi & Nelson, <strong>2009</strong><br />

Macrobiotus mauccii Pilato, 1974<br />

Macrobiotus meridionalis Richters, 1909<br />

Macrobiotus modestus Pilato & Lisi, <strong>2009</strong><br />

Macrobiotus montanus Murray, 1910 [Macrobiotus morulatus Bartoš, 1936]<br />

Macrobiotus mottai Binda & Pilato, 1994<br />

Macrobiotus nelsonae Guidetti, 1998<br />

Macrobiotus neuquensis Rossi, Claps & Ardohain, <strong>2009</strong><br />

Macrobiotus norvegicus Mihelčič, 1971/72<br />

Macrobiotus nuragicus Pilato & Sperlinga, 1975<br />

29

---

Macrobiotus occidentalis occidentalis Murray, 1910<br />

Macrobiotus occidentalis striatus Dastych, 1974<br />

Macrobiotus ocotensis Pilato, 20<strong>06</strong><br />

Macrobiotus orcadensis Murray, 1907<br />

Macrobiotus ovidii Bartoš, 1937<br />

Macrobiotus ovostriatus Pilato & Patanè, 1998<br />

Macrobiotus ovovillosus Baumann, 1960<br />

Macrobiotus pallarii Maucci, 1954 [Macrobiotus aviglianae Robotti, 1970]<br />

Macrobiotus papillosus Iharos, 1963<br />

Macrobiotus patagonicus Maucci, 1988<br />

Macrobiotus patiens Pilato, Binda, Napolitano & Moncada, 2000<br />

Macrobiotus perfidus Pilato & Lisi, <strong>2009</strong><br />

Macrobiotus persimilis Binda & Pilato, 1972<br />

Macrobiotus personatus Biserov, 1990<br />

Macrobiotus peterseni Maucci, 1991<br />

Macrobiotus pilatoi Binda & Rebecchi, 1992<br />

Macrobiotus polaris Murray, 1910<br />

Macrobiotus polonicus Pilato, Kaczmarek, Michalczyk & Lisi, 2003<br />

Macrobiotus polyopus Marcus, 1928<br />

Macrobiotus porteri Rahm, 1931<br />

Macrobiotus potockii Węglarska, 1968<br />

Macrobiotus primitivae de Barros, 1942<br />

Macrobiotus priviterae Binda, Pilato, Moncada & Napolitano, 2001<br />

Macrobiotus psephus du Bois-Reymond Marcus, 1944<br />

Macrobiotus pseudocoronatus Pilato, Binda & Lisi, 20<strong>06</strong><br />

Macrobiotus pseud<strong>of</strong>urcatus Pilato, 1972<br />

Macrobiotus pseudoliviae Pilato & Binda, 1996<br />

Macrobiotus pseudonuragicus Pilato, Binda & Lisi, 2004<br />

Macrobiotus punctillus Pilato, Binda & Azzaro, 1990<br />

Macrobiotus radiatus Pilato, Binda & Catanzaro, 1991<br />

Macrobiotus ragonesei Binda, Pilato, Moncada & Napolitano, 2001<br />

Macrobiotus ramoli Dastych, 2005<br />

Macrobiotus rawsoni Horning, Schuster & Grigarick, 1978<br />

Macrobiotus recens Cuénot, 1932<br />

Macrobiotus reinhardti Michalczyk & Kaczmarek, 2003<br />

Macrobiotus rigidus Pilato & Lisi, 20<strong>06</strong><br />

Macrobiotus rollei Heinis, 19<strong>21</strong><br />

Macrobiotus rubens Murray, 1907<br />

Macrobiotus sandrae Bertolani & Rebecchi, 1993<br />

Macrobiotus santoroi Pilato & D'Urso, 1976<br />

Macrobiotus sapiens Binda & Pilato, 1984<br />

Macrobiotus semmelweisi Pilato, Binda & Lisi, 20<strong>06</strong><br />

Macrobiotus serratus Bertolani, Guidi & Rebecchi, 1996<br />

Macrobiotus seychellensis Biserov, 1994<br />

Macrobiotus shennongensis Yang, 1999<br />

Macrobiotus siamensis Tumanov, 20<strong>06</strong><br />

Macrobiotus sicheli Binda, Pilato & Lisi, 2005<br />

<strong>30</strong>

---

Macrobiotus simulans Pilato, Binda, Napolitano & Moncada, 2000<br />

Macrobiotus sklodowskae Michalczyk, Kaczmarek & Węglarska, 20<strong>06</strong><br />

Macrobiotus snaresensis Horning, Schuster & Grigarick, 1978<br />

Macrobiotus sottilei Pilato, Kiosya, Lisi & Sabella, <strong>2012</strong><br />

Macrobiotus spectabilis Thulin, 1928<br />

Macrobiotus spertii Ramazzotti, 1957<br />

Macrobiotus stellaris du Bois-Reymond Marcus, 1944<br />

Macrobiotus striatus Mihelčič, 1949<br />

Macrobiotus submorulatus Iharos, 1966<br />

Macrobiotus szeptyckii Kaczmarek & Michalczyk, <strong>2009</strong><br />

Macrobiotus tehuelchensis Rossi, Claps & Ardohain, <strong>2009</strong><br />

Macrobiotus terminalis Bertolani & Rebecchi, 1993<br />

Macrobiotus terricola Mihelčič, 1951<br />

Macrobiotus tetraplacoides Fontoura, 1981<br />

Macrobiotus topali Iharos, 1969<br />

Macrobiotus trunovae Biserov, Pilato & Lisi, 2011<br />

Macrobiotus virgatus Murray, 1910<br />

Macrobiotus vladimiri Bertolani, Biserov, Rebecchi & Cesari, 2011<br />

Macrobiotus wauensis Iharos, 1973<br />

Macrobiotus wuzhishanensis Yin, L. Wang & X. Li, 2011<br />

Macrobiotus yunshanensis Yang, 2002<br />

Macrobiotus zhejiangensis Yin, L. Wang & X. Li, 2011<br />

Minibiotus R.O. Schuster, 1980<br />

Minibiotus acadianus Meyer & Domingue, 2011<br />

Minibiotus acontistus (de Barros, 1942)<br />

Minibiotus aculeatus (Murray, 1910) [Macrobiotus intermedius subjulietae Horning, Schuster &<br />

Grigarick, 1978]<br />

Minibiotus africanus Binda & Pilato, 1995<br />

Minibiotus allani (Murray, 1913)<br />

Minibiotus aquatilis Claxton, 1998<br />

Minibiotus asteris Claxton, 1998<br />

Minibiotus bisoctus (Horning, Schuster & Grigarick, 1978)<br />

Minibiotus claxtonae Rossi, Claps & Ardohain, <strong>2009</strong><br />

Minibiotus constellatus Michalczyk & Kaczmarek, 2003<br />

Minibiotus continuus Pilato & Lisi, 20<strong>06</strong><br />

Minibiotus crassidens (Murray, 1907)<br />

Minibiotus decrescens Binda & Pilato, 1995<br />

Minibiotus diphasconides (Iharos, 1969)<br />

Minibiotus eichhorni Michalczyk & Kaczmarek, 2004<br />

Minibiotus ethelae Claxton, 1998<br />

Minibiotus fallax Pilato, Claxton & Binda, 1989<br />

Minibiotus floriparus Claxton, 1998<br />

Minibiotus furcatus (Ehrenberg, 1859) [nec Macrobiotus furcatus Murray, 19<strong>06</strong>]<br />

Minibiotus granatai (Pardi, 1941)<br />

Minibiotus gumersindoi Guil & Guidetti, 2005<br />

Minibiotus harrylewisi Meyer & Hinton, <strong>2009</strong><br />

31

---

Minibiotus hispidus Claxton, 1998<br />

Minibiotus hufelandioides (Murray, 1910)<br />

Minibiotus intermedius (Plate, 1888)<br />

Minibiotus jonesorum Meyer, Lyons, Nelson & Hinton, 2011<br />

Minibiotus julianae Meyer, <strong>2012</strong><br />

Minibiotus julietae (de Barros, 1942)<br />

Minibiotus keppelensis Claxton, 1998<br />

Minibiotus maculartus Pilato & Claxton, 1988<br />

Minibiotus marcusi (de Barros, 1942)<br />

Minibiotus milleri Claxton, 1998<br />

Minibiotus orth<strong>of</strong>asciatus Fontoura, Pilato, Lisi & Morais, <strong>2009</strong><br />

Minibiotus pilatus Claxton, 1998<br />

Minibiotus poricinctus Claxton, 1998<br />

Minibiotus pustulatus (Ramazzotti, 1959)<br />

Minibiotus ramazzottii Binda & Pilato, 1992<br />

Minibiotus scopulus Claxton, 1998<br />

Minibiotus sidereus Pilato, Binda & Lisi, 2003<br />

Minibiotus stuckenbergi (Dastych, Ryan & Watkins, 1990)<br />

Minibiotus subintermedius (Ramazzotti, 1962)<br />

Minibiotus taiti Claxton, 1998<br />

Minibiotus vinciguerrae Binda & Pilato, 1992<br />

Minibiotus weglarskae Michalczyk, Kaczmarek & Claxton, 2005<br />

Minibiotus weinerorum (Dastych, 1984)<br />

Minibiotus wuzhishanensis X. Li, D. Wang & L. Wang, 2008<br />

Minibiotus xavieri Fontoura, Pilato, Morais & Lisi, <strong>2009</strong><br />

Minilentus Guidetti & Pilato, 2003<br />

Minilentus dubius (Schuster & T<strong>of</strong>tner, 1982)<br />

Paramacrobiotus Guidetti, Schill, Bertolani, Dandekar & Wolf, <strong>2009</strong><br />

Paramacrobiotus alekseevi (Tumanov, 2005) <br />

Paramacrobiotus areolatus (Murray, 1907) <br />

Paramacrobiotus beotiae (Durante Pasa & Maucci, 1979) {<strong>species</strong> dubia; Guidetti et al. <strong>2009</strong>}<br />

Paramacrobiotus centesimus (Pilato, 2000) <br />

Paramacrobiotus chieregoi (Maucci & Durante Pasa, 1980) <br />

Paramacrobiotus corgatensis (Pilato, Binda & Lisi, 2002) <br />

Paramacrobiotus crenatus (Maucci, 1991) <br />

Paramacrobiotus csotiensis (Iharos, 1966) <br />

Paramacrobiotus danielae (Pilato, Binda, Napolitano & Moncada, 2001) <br />

32

---

Paramacrobiotus derkai (Degma, Michalczyk & Kaczmarek, 2008) <br />

Paramacrobiotus fairbanksi Schill, Förster, Dandekar & Wolf, 2010 {The <strong>species</strong> have been<br />

diagnosed based on genetic data only and no morphological difference from<br />

Paramacrobiotus richtersi has been found}<br />

Paramacrobiotus garynahi (Kaczmarek, Michalczyk & Diduszko, 2005) <br />

Paramacrobiotus gerlachae (Pilato, Binda & Lisi, 2004) <br />

Paramacrobiotus halei (Bartels, Pilato, Lisi & Nelson, <strong>2009</strong>) <br />

Paramacrobiotus huziori (Mihalczyk & Kaczmarek, 20<strong>06</strong>) <br />

Paramacrobiotus kenianus Schill, Förster, Dandekar & Wolf, 2010 {The <strong>species</strong> have been<br />

diagnosed based on genetic data only and no morphological difference from<br />

Paramacrobiotus richtersi has been found}<br />

Paramacrobiotus klymenki Pilato, Kiosya, Lisi & Sabella, <strong>2012</strong><br />

Paramacrobiotus lorenae (Biserov, 1996) <br />

Paramacrobiotus magdalenae (Mihalczyk & Kaczmarek, 20<strong>06</strong>) <br />

Paramacrobiotus palaui Schill, Förster, Dandekar & Wolf, 2010 {The <strong>species</strong> have been<br />

diagnosed based on genetic data only and no morphological difference from<br />

Paramacrobiotus richtersi has been found}<br />

Paramacrobiotus peteri (Pilato, Claxton & Binda, 1989) <br />

Paramacrobiotus richtersi (Murray, 1911) [Macrobiotus schultzei Greeff, 1866] <br />

Paramacrobiotus rioplatensis (Claps & Rossi, 1997) {doubly attributed to the genus; Guidetti et<br />

al. <strong>2009</strong>}<br />

Paramacrobiotus savai (Binda & Pilato, 2001) <br />

Paramacrobiotus tonollii (Ramazzotti, 1956) <br />

Paramacrobiotus vanescens (Pilato, Binda & Catanzaro, 1991) <br />

Paramacrobiotus walteri (Biserov, 1997/98) <br />

Pseudodiphascon Ramazzotti, 1965<br />

Pseudodiphascon inflexum (Arcidiacono, 1964)<br />

Pseudohexapodibius Bertolani & Biserov, 1996<br />

Pseudohexapodibius degenerans (Biserov, 1990)<br />

Richtersius Pilato & Binda, 1989<br />

[Richtersia Pilato & Binda, 1987]<br />

33

---

Richtersius coronifer (Richters, 1903)<br />

Schusterius Kaczmarek & Michalczyk, 20<strong>06</strong><br />

Schusterius tridigitus (R.O. Schuster, 1983)<br />

Tenuibiotus Pilato & Lisi, 2011<br />

Tenuibiotus bondavallii (Manicardi, 1989) <br />

Tenuibiotus bozhkae Pilato, Kiosya, Lisi, Inshina & Biserov, 2011<br />

Tenuibiotus ciprianoi (Guil, Guidetti & Machordom, 2007) <br />

Tenuibiotus danilovi (Tumanov, 2007) <br />

Tenuibiotus higginsi (Maucci, 1987) <br />

Tenuibiotus hyperonyx (Maucci, 1983) <br />

Tenuibiotus hystricogenitus (Maucci, 1978) <br />

Tenuibiotus kozharai (Biserov, 1999) <br />

Tenuibiotus mongolicus (Maucci, 1988) <br />

Tenuibiotus tenuiformis (Tumanov, 2007) <br />

Tenuibiotus tenuis (Binda & Pilato, 1972) <br />

Tenuibiotus voronkovi (Tumanov, 2007) <br />

Tenuibiotus willardi (Pilato, 1977) <br />

Xerobiotus Bertolani & Biserov, 1996<br />

Xerobiotus euxinus Pilato, Kiosya, Lisi, Inshina & Biserov, 2011<br />

Xerobiotus pseudohufelandi (Iharos, 1966) [Macrobiotus inermis Binda & Pilato, 1971]<br />

Xerobiotus xerophilus (Dastych, 1978)<br />

34<br />

Murrayidae Guidetti, Gandolfi, Rossi & Bertolani, 2005<br />

Dactylobiotus R.O. Schuster, 1980<br />

Dactylobiotus ambiguus (Murray, 1907)<br />

Dactylobiotus ampullaceus (Thulin, 1911)<br />

Dactylobiotus aquatilis Yang, 1999<br />

Dactylobiotus caldarellai Pilato & Binda, 1994<br />

Dactylobiotus dervizi Biserov, 1998<br />

Dactylobiotus dispar (Murray, 1907)<br />

Dactylobiotus grandipes (Schuster, T<strong>of</strong>tner & Grigarick, 1978)

---

Michalczyk 2010><br />

Dactylobiotus haplonyx Maucci, 1981<br />

Dactylobiotus henanensis Yang, 2002<br />

Dactylobiotus kansae Beasley, Miller & Shively, <strong>2009</strong><br />

Dactylobiotus lombardoi Binda & Pilato, 1999<br />

Dactylobiotus luci Kaczmarek, Michalczyk & Eggermont, 2008<br />

Dactylobiotus macronyx (Dujardin, 1851)<br />

Dactylobiotus octavi Guidetti, Altiero & Hansen, 20<strong>06</strong><br />

Dactylobiotus parthenogeneticus R. Bertolani, 1982<br />

Dactylobiotus selenicus R. Bertolani, 1982<br />

Macroversum Pilato & Catanzaro, 1988<br />

Macroversum mirum Pilato & Catanzaro, 1988<br />

Murrayon Bertolani & Pilato, 1988<br />

Murrayon dianeae (Kristensen, 1982)<br />

Murrayon hastatus (Murray, 1907)<br />

Murrayon hibernicus (Murray, 1911)<br />

Murrayon nocentiniae (Ramazzotti, 1961)<br />

Murrayon ovoglabellus (Biserov, 1988)<br />

Murrayon pullari (Murray, 1907)<br />

Murrayon stellatus Guidetti, 1998<br />

{the placement <strong>of</strong> following taxa into any superfamily remains still unclear, see Marley et al.<br />

2011}<br />

Beornidae Cooper, 1964<br />

Beorn leggi Cooper, 1964 {fossil <strong>species</strong>}<br />

Necopinatidae Ramazzotti & Maucci, 1983<br />

Necopinatum Pilato, 1971<br />

Necopinatum mirabile Pilato, 1971<br />

incertae sedis<br />

Apodibius Dastych, 1983<br />

Apodibius confusus Dastych, 1983<br />

Apodibius nuntius Binda, 1984 [Apodibius serventyi Morgan & Nicholls, 1986]<br />

Apodibius richardi Vargha, 1995<br />

References<br />

Bartels, P.J. & Nelson, D.R. (<strong>2012</strong>) An online key and field guide to the tardigrades <strong>of</strong> the Great<br />

Smoky Mountains National Park (North Carolina & Tennessee, USA, North America) with<br />

taxonomic revisions <strong>of</strong> two <strong>species</strong>. Zootaxa, 3249, 67–68.<br />

Chang, C.Y., Kaczmarek, Ł., Lee, J.M. & Michalczyk, Ł. (2007) Pseudobiotus spinifer, a new<br />

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tardigrade <strong>species</strong> (Eutardigrada: Hypsibiidae) from Nakdong River, South Korea, with a<br />

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Claxton, S.K., Beasley, C.W., Kaczmarek, L. & Michalczyk, L. (2010) Redescription <strong>of</strong><br />

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Guidetti, R., Schill, R.O., Bertolani, R., Dandekar, T. & Wolf, M. (<strong>2009</strong>) New molecular data for<br />

tardigrade phylogeny, with the erection <strong>of</strong> Paramacrobiotus gen. n. Journal <strong>of</strong> Zoological<br />

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Jørgensen A, Faurby, S., Hansen, J.G., Møbjerg, N. & Kristensen R.M. (<strong>2009</strong>) Molecular<br />

phylogeny <strong>of</strong> Arthrotardigrada (<strong>Tardigrada</strong>). Molecular Phylogenetics and Evolution, 54<br />

(2010), 10<strong>06</strong>–1015.<br />

Kaczmarek, Ł. & Michalczyk, Ł. (<strong>2009</strong>) Redescription <strong>of</strong> Hypsibius microps Thulin, 1928 and H.<br />

pallidus Thulin, 1911 (Eutardigrada: Hypsibiidae) based on the type material from the<br />

Thulin collection. Zootaxa, 2275: 60-68.<br />

Kaczmarek, Ł. & Michalczyk, Ł. (2010) Redescription <strong>of</strong> Dactylobiotus grandipes (Schuster,<br />

T<strong>of</strong>tner & Grigarick, 1978), the type <strong>species</strong> for the genus Dactylobiotus (<strong>Tardigrada</strong>:<br />

Macrobiotidae). Genus, <strong>21</strong>: 155-162.<br />

Kaczmarek, Ł., Bertolani, R., Nędzyńska-Stygar, M. (2010) Thulinius saltursus comb. nov.: a<br />

new systematic position for Isohypsibius saltursus Schuster, T<strong>of</strong>tner & Grigarick, 1978<br />

(Eutardigrada: Hypsibiidae) and a key for the genus Thulinius. Zootaxa, 2483: 58-64.<br />

Li, X. (<strong>2009</strong>) Revision <strong>of</strong> Biserovus xiae Li, Su and Yu, 2004 (<strong>Tardigrada</strong>: Macrobiotidae).<br />

Zootaxa, 2076, 63–68.<br />

Lisi, O. (2011) Remarks on Doryphoribius flavus (Iharos, 1966), and description <strong>of</strong> three new<br />

<strong>species</strong> (<strong>Tardigrada</strong>, Hypsibiidae). Zootaxa, 2834, 17–32.<br />

Marley, N.J., McInnes, S.J. & Sands, C.J. (2011) Phylum <strong>Tardigrada</strong>: A re-evaluation <strong>of</strong> the<br />

Parachela. Zootaxa, 2819, 51–64.<br />

Michalczyk, Ł., Wełnicz, W., Frohme M. & Kaczmarek, Ł. (<strong>2012</strong>) Redescriptions <strong>of</strong> three<br />

Milnesium Doyère, 1840 taxa (<strong>Tardigrada</strong>: Eutardigrada: Milnesiidae), including the<br />

nominal <strong>species</strong> for the genus. Zootaxa 3154, 1–20.<br />

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