Checklist of the mosses of sub-Saharan Africa

Tropical Bryology 10: 91-198, 1995 

Checklist of the mosses of sub-Saharan Africa 

Brian O’Shea 

141 Fawnbrake Avenue, London SE24 OBG, U.K. 

Abstract. 2939 moss taxa are listed for sub-Saharan Africa and adjacent islands, with distribution by 

country. Each distribution record is supported by a literature reference. The following new 

combinations are made: Calyptrochaeta cristata (Hedw.) O’Shea, Groutiella elimbata (Thér) O’Shea, 

Meiothecium undulatum (Ren. & Card.) O’Shea, Orthodontium ruwenzorensis (Thér. & Nav.) O’Shea, 

Pohlia lacouturei (Thér.) O’Shea, Sematophyllum corticolum (Aongstr.) O’Shea, Sematophyllum 

dixonii (Thér.) O’Shea, Sematophyllum nanopyxis (Geh.) O’Shea, Sematophyllum rigescens (Card.) 

O’Shea, and Thamnobryum malgachum (Card.) O’Shea. 

Introduction 

Identifying mosses collected on the BBS Tropical 

Bryology Group expedition to Mulanje Mountain, 

Malawi, in 1991 proved difficult because the 

available literature was so diverse and it was 

difficult to know which taxa were worth pursuing 

in the literature as possible identifications for our 

material. Although there was a great deal of 

expertise available, there seemed no published 

view available of the recorded bryoflora. This 

checklist is intended to give an overall view of the 

distribution of tropical African mosses, and a list 

of sub-Saharan hepatics is also being produced, 

which should be published shortly (84). [It should 

be noted that all references in this paper are given 

as numbers, but these are translated into 

‘author+year’ references in the bibliography.] 

The need for checklists containing “current 

names” (rather than just a list of all names ever 

created) has been identified as one of the most 

vital tools needed to progress conservation and 

ecological issues in botany (1639). Such checklists 

are seen as tactical steps towards a global 

plant species information system. There has also 

been debate about the most appropriate approach 

to putting together the taxonomic backbone of 

such an information system and what information 

should be stored, and the data in this checklist 

covers the basic requirement, common across all 

plant groups: species name, important synonyms, 

generic and family placement, geographical 

distribution and bibliographic citation. 

91

The scope of the checklist is all of sub-Saharan 

Africa, together with the Atlantic and Indian 

Ocean islands in the tropics. This does not quite 

conform to the Index Muscorum divisions, as part 

of Africa-2 is excluded to the north (the Sahelian 

area within the tropics), and part of Africa-4 is 

excluded to the south (the islands in the southern 

Atlantic and Indian Oceans). For practical reasons 

it was necessary to draw a definite boundary 

round the region, even when this cut across 

existing affinities, such as the north Atlantic islands 

(Cape Verde is included, the Canaries, Madeira 

and Azores are excluded). This differs slightly 

from the combined Bryologia Africana and Flora 

of Southern Africa areas, by excluding Mauritania, 

and including the islands of Cape Verde, Ascension 

and St. Helena . 

All African countries apart from Sudan have been 

covered by checklists although some of these are 

now rather old (see map). All of these have been 

entered into a database, and updated with later 

publications. The publications on which the list 

is based are shown in the references. I would be 

grateful for pointers to any omitted publications, 

or monographs that include African taxa, that are 

not already included in the references. Where 

data has been taken from an existing checklist, I 

have not included all the individual references and 

have instead just given the name of the list - e.g. 

east African references will typically show (46) 

(=Kis, 1985) as the reference. It will be necessary 

to go to the source document quoted to get the 

detailed references. The resulting list contains 

2939 taxa, with over 9000 taxon/country occurrences 

- an average of 3 countries (or part countries) per 

taxon. All taxa in the source lists should appear 

in the checklist as either accepted taxa, or as 

synonyms or excluded taxa. 

This document is not intended as a taxonomic 

work. The quality of the data in this list is thus 

variable in its taxonomic quality, although it 

attempts to present the latest position for all taxa. 

Some difficult and complex families where revision 

is almost complete (e.g. Calymperaceae) are in a 

good state, and the list represents a useful synthesis 

of the current position. Where the process of 

revision is still underway (e.g. Fissidentaceae) 

there are now large numbers of synonyms, but 

clearly a large number of taxa still to tackle. In other 

areas work has hardly begun, such as 

Sematophyllaceae and parts of Hypnaceae, and 

this will continue to cause problems for those 

attempting to identify plants in these groups. The 

list will at least indicate the scale of the problem. 

Producing and maintaining the list would have 

been unmanageable without computer assistance, 

and all data used in the list is held on the TAXA 

system (115), and the main taxon distribution list 

and list of references was generated directly from 

TAXA. This system is available to anyone 

requiring it, but it is being continually enhanced, 

and so potential users should contact the author 

for the latest version. 

Although I have checked this list back against the 

source documents, this project has taken some 

time, and has led to considerable manipulation of 

data to take account of synonyms and new publications, 

and what was once correct may no longer 

be so. Please let me know of any mistakes. 

This checklist is expected to be a source and 

foundation for future work, rather than a final 

statement. Future updates are planned. 

Assumptions, Comments & Compromises 

A list this size cannot be produced without taking 

a pragmatic view, and some will disagree with the 

approach I have taken in some areas. Listed below 

are some of the problems faced and assumptions 

taken. 

- The basic list has been built from the latest 

‘authoritative’ list for the country concerned 

- i.e. on lists that validate the occurrence in a 

country via earlier lists or collections, as 

shown on the map. Taxa mentioned in Index 

Muscorum as occurring in Africa-2, 3 or 4 are 

not included unless they have appeared in a 

country list. Records not localised to a 

country have been ignored, and taxa that are 

not identified at least to subspecies or variety 

level (i.e. forms) are marked as excluded. The 

basic list has then been enhanced by data 

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94 

from references published since the original 

country lists. 

- Where the source information does not 

provide sufficient detail, group names have 

been used (e.g. Senegambia, for Senegal + 

Gambia). Where there is sufficient detail, and 

the units are sufficiently large, intra-country 

divisions have been used (e.g. South Africa 

is divided into four regions), and also where 

the parts are particularly isolated (e.g. the two 

major islands belonging to Equatorial Guinea, 

Fernando Po (Bioko) and Annobon are 

recorded separately from mainland Rio Muni). 

When it is not clear where all the records are 

from, the group name may be used in 

conjunction with an island or ‘part-country’ 

name (e.g. MAS(40), REU(105)). 

- Where there is more than one reference for 

the occurrence of a taxon in a country, I have 

not in all cases checked whether the specimens 

referred to are different. Where I have checked, 

I have chosen between duplicates by retaining 

those with: 1) more widely published 

references, 2) more authoritative/detailed 

references, 3) references with wider 

information. 

- All references seen up to the end of February 

1995 have been included. 

- It has been assumed that if one plant is 

synonymised with another, then its distribution 

data can also be transferred. A situation 

where this approach could cause problems is 

exemplified by Rhodobryum spathulatum, 

where the type is actually Bryum aubertii, but 

the collections called R. spathulatum are in 

fact R. ontariense. I hope I have managed to 

identify most of such cases. 

- Where names from the source lists have 

been synonymised, the original names under 

which taxa were collected have not been 

included in the distribution data; this data 

should be available in the source reference. 

Synonyms are only listed in alphabetic sequence 

with the rest of the names, and not 

under the valid taxa to which they apply. This 

means that it may not be possible to work back 

from a reference to the original names(s) under 

which it was collected, other than by working 

from the source document. 

- No distinction has been made between 

nomenclatural and taxonomic synonyms. 

- I have found a number of taxa that do not 

appear in Index Muscorum (1521), or the 

Index of Mosses (80, 1644). The status of such 

taxa is not always clear (although usually 

nomina nuda), and they tend to get missed 

from taxonomic revisions. 

- I have made no judgement on the age of 

records - all apparently valid records have 

been accepted. 

New combinations 

During the compilation of this Checklist, a number 

of taxa were noticed that appeared to be in need of 

names in different genera. This situation has 

occurred for two reasons: 

- nomenclatural changes have been made 

that leave ‘orphaned’ varieties or other subtaxa, 

e.g. Drepanocladus fluitans has been 

transferred to the genus Warnstorfia, but its 

variety monodii was not included in the 

transfer, and nor was it synonymised; 

- where taxa have been transferred en bloc 

from one genus to another (for instance 

because of the discovery of an earlier valid 

name for the genus) sometimes taxa have 

been left behind, e.g. Eriopus cristata was 

not transferred to Calyptrochaeta when Eriopus 

was made a synonym of Calyptrochaeta. 

In the first case I have left the taxon in its original 

genus, with a note pointing to the correct location 

if the variety proves to be distinct. In the second 

case I have made new combinations. 

There are a number of nomina nuda on the checklist 

that also need a change of name, but these have 

been commented on in the checklist text, and are 

not covered here. 

My intention is to leave the taxonomic problems 

(if any) clearer. I appreciate that these transfers 

may detract from the taxonomic purity of the 

receiving genus, but I feel it is better than leaving 

taxa in an ‘invalid’ genus. If they are not valid taxa 

they should have been synonymised or excluded 

as part of the original transfer. Keeping them 

visible in this way may help resolve their status.

Calyptrochaeta cristata (Hedw.) O’Shea, comb. 

nov. 

Basionym: Leskea cristata Hedw., Spec. 

Musc. 2:211. 49. 1801. 

[Subsequently recombined as Eriopus cristatus 

(Hedw.) Brid., Bryol. Univ. 2:788. 1827.] 

Eriopus C.Müll. nom. illeg. has been replaced by 

Calyptrochaeta Desvaux (1521). 

Groutiella elimbata (Thér.) O’Shea, comb. nov. 

Basionym: Micromitrium elimbatum Thér., 

Recueil Publ. Soc. Havraise Étud. Div. 

1929:113. ic. 1930. 

Micromitrium Schimp. in Besch. is a synonym of 

Groutiella Steere (1521). 

Meiothecium undulatum (Ren. & Card.) O’Shea, 

comb. nov. 

Basionym: Pterogoniella undulata Ren. & 

Card., Prodr. Fl. Bryol. Madag. Suppl. 67. 

1909. 

Pterogoniella Schimp. in Broth. has been replaced 

by Meiothecium Mitt. (1521) 

Orthodontium ruwenzorensis (Thér. & Nav.) 

O’Shea, comb. nov. 

Basionym: Stableria ruwenzorense Thér. & 

Nav., Bull. Soc. R. Bot. Belg. 60:34. 15. 1927. 

Stableria Lindb. ex Braithw. is a synonym of 

Orthodontium Schwaegr. sect. Stableria 

(Braithw.)Meijer (1521). 

Pohlia lacouturei (Thér.) O’Shea, comb. nov. 

Basionym: Webera lacouturei Thér., Recueil 

Publ. Soc. Havraise Étud. Div. 89:124. ic. 

1922. 

Webera Hedw. hom. illeg. is replaced by Pohlia 

Hedw. (1521). 

Sematophyllum corticolum (Aongstr.) O’Shea, 

comb. nov. 

Basionym: Plagiothecium corticola 

Aongstr., Defr. K. Vet. Ak. Foerh. 30(5):143. 

1873. [“um”]. 

This synonymisation was first suggested by Brothe- 

rus in 1908 (Nat. Pfl. 1(3):1087), and referred to 

subsequently in Wijk et al. (1521) and Crosby et 

al.(40). 

Sematophyllum dixonii (Thér.) O’Shea, comb. 

nov. 

Basionym: Rhaphidostegium dixonii Thér., 

Recueil Publ. Soc. Havraise Étud. Div. 

1924(2):96. 1924. [“-i”] (R. densirete Thér., 

1923 hom. illeg.). 

Rhaphidostegium (BS&G)De. Not. nom. illeg. incl. 

gen. prior. is replaced by Sematophyllum (1521). 

Sematophyllum nanopyxis (Geh. )O’Shea, comb. 

nov. 

Basionym: Hypnum nanopyxis C.Müll. ex 

Geh., Abh. Naturw. Ver. Bremen 7:212. 1881. 

[Subsequently recombined as Rhaphidostegium 

nanopyxis (Geh.)Par., Ind. 

Bryol.:1100. 1897.] 

See note under Sematophyllum dixonii. 

Sematophyllum rigescens (Card.) O’Shea, comb. 

nov. 

Basionym: Rhapidostegium rigescens Card. 

in Grand., Hist. Madag. 39:499. 1915. 

See note under Sematophyllum dixonii. 

Thamnobryum malgachum (Card.) O’Shea, comb. 

nov. 

Basionym: Thamnium malgachum Card. in 

Grand., Hist. Madag. 39:382. 1915. 

De Sloover (15) (p. 148) lists this taxon as doubtful 

but gives no reason, and presumably had not seen a 

collection. Cardot gives a clear description of what 

he says is a very distinctive plant. Until the type 

is examined, I have transferred the plant to Tham- 

nobryum. 

Checklist 

The list is in alphabetic order of the current version 

of taxon names, and any changes to earlier names 

(taxonomic and nomenclatural synonyms) are 

shown inset, in a smaller font, giving the new 

name, as are comments and excluded species. 

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96 

Only synonyms that postdate the source list have 

been given: earlier synonyms can be found in the 

source list and its references. The assumed family 

of each taxon is shown in Appendix B. 

The countries in which a taxon is recorded are 

shown by a three character abbreviation. These 

abbreviations are interpreted in Appendix A. 

Country abbreviations are listed for each taxon in 

alphabetic order of country name, not abbreviation, 

so, for example, CBS (Cape Province, South 

Africa) comes before CAR (Central African 

Republic). The numbers in brackets after each 

country (and after synonyms) are the literature 

references, which are translated to ‘author+year’ 

in the References section. 

Abietinella abietina (Hedw.) Fleisch. = Thuidium 

abietinum (71). 

Acanthorrhynchium decolor (Besch.) Fleisch. 

SEY(40). 

Acanthorrhynchium loucoubense (Besch.) 

Fleisch. COM(40), MAD(40). 

Acanthorrhynchium serratum (Ren. & Card.) 

Fleisch. COM(40), MAD(40). 

Acaulon leucochaete Stone CBS(1531). 

Acaulon recurvatum Magill CBS(1531). 

Acaulon rufochaete Magill = Microbryum rufochaete 

(1420). 

Acrocryphaea emarginata Thér. & Nav. in P.Varde, 

nom. nud. See notes under Schoenobryum. 

Acrocryphaea subrobusta Broth. & P.Varde = Schoe- 

nobryum subrobustum (1291). 

Acroporium depressum Thér. MAD(40). 

Acroporium eburensis Bizot IVO(4). 

Acroporium laevifolium (Ren.) Broth. MAD(40). 

Acroporium le-testui P.Varde GAB(47). 

Acroporium megasporum (Duby) Fleisch. = Macro- 

hymenium megasporum (46). 

Acroporium megasporum var. densum (Ren. & 

Card.) Wijk & Marg. MAD(40). 

Acroporium megasporum var. heterodictyon 

P.Varde GAB(47). 

Acroporium megasporum vars. A. megasporum has 

been transferred to Macrohymenium (46). These 

varieties also are in need of revision. 

Acroporium pocsii Biz. CAM(47). 

Acroporium prionophylax Buck ZAI(65). 

Acroporium pungens (Hedw.) Broth. RWA(65), 

ZAI(65). 

Acroporium stellatum (Ren. & Card.) Broth. 

MAD(40). 

Actinodontium dusenii Broth. CAM(47). 

Actinodontium streptopogoneum Broth. 

CAM(47). 

Adelothecium bogotense (Hampe) Mitt. 

MAD(40), TAN(1347). 

Aerobryidium subpiligerum (Hampe) Card. 

COM(40), KEN(46), MAD(40), MAL(67), 

MAU(105), REU(105), TAN(46), ZAI(65). 

Aerobryidium subpiligerum var. majus (Ren. & 

Card.) Wijk & Marg. REU(105). 

Aerobryopsis capensis (C.Müll.) Fleisch. 

CBS(1531), MAL(67), NAT(1531), TAN(46), 

TRV(1531), ZAM(46), ZIM(46). 

Aerobryopsis cirrifolia (Schwaegr.) Paris 

MAU(105). 

Aerobryopsis cymatocheilos (Kindb.) Broth. 

TAN(46). 

Aerobryopsis longipendula (Dus.) Broth. 

CAM(47). 

Aerobryopsis longissima (Dozy & Molk.) Fleisch. 

SEY(40). 

Aerobryopsis pseudocapensis (C.Müll.) Fleisch. 

ETH(50). 

Aerobryopsis trachyptera (Dus.) Broth. CAM(47), 

GAB(47). 

Aerolindigia capillacea (Hornsch.) Menzel 

MAD(40), RWA(65), UGA(46), ZAI(65). 

Aloina bifrons (De Not.) Delg. CBS(1531). 

Aloina brevirostris (Hook. & Grev.) Kindb. 

ZIM(61). 

Aloina pilifera (De Not.) Crum & Steere = A. bifrons 

(81). 

Aloina rigida (Hedw.) Limpr. DJI(1569), KEN(411). 

Amblystegium camisassae Tosco & Piovano 

KEN(46). 

Amblystegium chalaropelma Ren. MAD(40). 

Amblystegium riparium (Hedw.) BS&G 

BOT(1531), KEN(46), MAD(40), MAL(67), 

NAT(1531), OFS(1531), TRV(1531), ZAI(65), 

ZAM(46), ZIM(46). 

Amblystegium riparium fo. elongatum (BS&G) 

Moenk. Excluded. Any records for this form 

have been associated with the owning species. 

Amblystegium serpens var. serpens (Hedw.) BS&G 

UGA(46). 

Amphidium aloysii-sabaudiae Negri UGA(46). 

Amphidium cyathicarpum (Mont.) Broth. = A.

tortuosum (65). 

Amphidium lapponicum (Hedw.) Schimp. 

LES(1494,1531), NAT(1531,1494). 

Amphidium le-testui Thér. & P.Varde GAB(47), 

ZAI(65). 

Amphidium sp. ETH(50) 

Amphoridium semense Schimp. in Par., nom. nud. is 

the base name for this taxon. Amphoridium is 

a synonym for Amphidium, but this taxon cannot 

be dealt with satisfactorily until a type is nominated 

and described, or it is synonymised with another 

taxon. There is a ‘type’ sheet in BM for this 

taxon, containing specimens from Herb. Hampe, 

leg. Schimper. 

Amphidium tortuosum (Hornsch.) Robins. 

CAM(47), CBS(1531), ETH(50), KEN(46), 

LES(1494,1531), NAT(1531), TAN(46), 

UGA(46), ZAI(65). 

Amphoridium semense Schimp. in Par., nom. nud. 

See note under Amphidum sp. 

Anacamptodon africanus P.Varde CAR(47), 

TAN(46). 

Anacolia abyssinica (C.Müll.) Schimp. = Flowersia 

abyssinica (182). 

Anacolia breutelii var. breutelii (C.Müll.) Magill 

CBS(1531), LES(1531), NAT(1531), OFS(1531). 

Anacolia breutelii var. squarrifolia (Sim) Magill 

CBS(1531), LES(1531), NAT(1531). 

Anacolia cameruniae Dix. CAM(47), TAN(46). 

Anacolia laevisphaera (Tayl.) Flowers CAM(47), 

KEN(46), REU(105), RWA(65), SUD(1419), 

TAN(46), UGA(46), ZAI(65). 

Anacolia scioana (Brizi) Cuf. ETH(50). 

Andreaea bistratosa Magill CBS(1531). 

Andreaea borbonica Besch. REU(105). 

Andreaea camerunensis Richards CAM(47). 

Andreaea cucullata Dix. KEN(46), TAN(46). 

Andreaea firma C.Müll. KEN(46), RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Andreaea mildbraedii Broth. KEN(46), 

RWA(65,46), TAN(46), UGA(46), ZAI(65), 

ZIM(46). 

Andreaea nitida Hook.f. & Wils. CBS(1531). 

Andreaea obovata var. obovata Thed. KEN(46), 

RWA(65), TAN(46), ZAI(65). 

Andreaea rupestris var. rupestris Hedw. 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TAN(1410), ZIM(61). 

Andreaea subulata Harv CBS(1531), MAD(40), 

UGA(46). 

Andreaea tsaratananae Thér. MAD(40), 

REU(105). 

Anisothecium horridum P.Varde CAR(47). 

Anisothecium madagassum Thér. MAD(40). 

Anisothecium ugandae P.Varde RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Anoectangium aestivum (Hedw.) Mitt. CAM(47), 

CAR(47), IVO(1458), SUD(1419), TAN(46), 

TOG(1878). 

Anoectangium afrocompactum C.Müll. ex Dus. 

CAM(47). 

Anoectangium borbonense Besch. COM(40), 

MAU(105), NIG(334), REU(105). 

Anoectangium brotherusii Kis KEN(46), MAL(67), 

TAN(46). 

Anoectangium euchloron (Schwaegr.) Mitt. = A. 

aestivum (46). 

Anoectangium euchloron var. laxum P.Varde 

CAR(47). 

Anoectangium euchlorum var. laxum P.Varde. The 

owning species of this variety has been transferred 

to A. aestivum (46). This taxon is also in need 

of revision. 

Anoectangium eukilimandscharicum Dix. 

TAN(46). 

Anoectangium flexuosum Negri UGA(46). 

Anoectangium flexuosum var. gracile Negri 

UGA(46). 

Anoectangium fuscum Negri UGA(46). 

Anoectangium hanningtonii Mitt. TAN(46), 

ZAM(62). 

Anoectangium humblotii Ren. & Card. COM(40). 

Anoectangium impressum Hampe MAD(40), 

REU(105). 

Anoectangium keniae (P.Varde) Zander KEN(46). 

Anoectangium madagassum Ren. & Par. 

MAD(40). 

Anoectangium mafatense Ren. & Card. REU(105), 

UGA(46), ZAI(65). 

Anoectangium nigerianum Broth. & Par. MAI(47). 

Anoectangium rhaphidostegium Besch. 

COM(40), REU(105). 

Anoectangium rufoviride Besch. COM(40), 

ERI(50), MAD(40). 

Anoectangium rufoviride var. eucollum Besch. 

COM(40). 

Anoectangium schimperi Mitt. ETH(50). 

Anoectangium sellae Negri UGA(46). 

Anoectangium socotranum (Mitt.) Broth. 

ETH(50), SOC(50). 

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98 

Anoectangium spathulatum Mitt. CAM(47). 

Anoectangium torquatum Broth. = A. brotherusii 

(46). 

Anoectangium wilmsianum (C.Müll.) Par. 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TRV(1531). 

Anomobryum auratum (Mitt.) Jaeg. KEN(46), 

MAL(67), TAN(1404,46), UGA(46), ZAI(65), 

ZAM(62). 

Anomobryum compressulum (C.Müll.) Broth. 

KEN(46), TAN(46), UGA(46). 

Anomobryum cygnicollum (C.Müll.) Hampe ex 

Jaeg. ERI(50). 

Anomobryum drakensbergense Van Rooy 

LES(1531), NAT(1531), OFS(1531). 

Anomobryum filiforme (Dicks.) Husn = A julaceum 

(46). 

Anomobryum filiforme var. madagassum Ren. & 

Card. MAD(40). 

Anomobryum filiforme var. madagassum Ren. & 

Card. This variety presumably belongs to A. 

julaceum, following the synonymisation of A. 

filiforme (46). 

Anomobryum julaceum (Gaertn. et al.) Schimp. 

ANG(46), CAM(47), CBS(1531), COM(40), 

ETH(50), KEN(46), LES(1531), MAD(40), 

MAL(67), NAT(1531), OFS(1531), 

RWA(46,65), SOC(40,1626), SUD(1419), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZIM(46,61). 

Anomobryum julaceum var. abyssinicum Schimp., 

nom. nud. ERI(50), ETH(50). 

Anomobryum julaceum var. juliforme (Solms) 

Husn. CAM(47). 

Anomobryum laceratum (Besch.) Broth. REU(105). 

Anomobryum laceratum var. comorense (Besch.) 

Paris COM(40). 

Anomobryum maclennanii Broth. = A. sulcatum 

(46) (= A. auratum). 

Anomobryum minutirete (C.Müll.) Kis KEN(46), 

TAN(46). 

Anomobryum perimbricatum (Broth.) Broth. 

CAM(47). 

Anomobryum sulcatum Thér. & P.Varde = Bryum 

auratum (23) = Anomobryum auratum (1521). 

Antitrichia curtipendula (Hedw.) Brid. Specimens 

given this name in Africa should be referred to A. 

kilimandscharica (804). 

Antitrichia kilimandscharica Broth. ETH(50), 

KEN(46,136), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Aongstroemia filiformis (P.Beauv.) Wijk & Marg. 

LES(1074), MAD(40), MAL(67), NAT(1531), 

REU(105), TAN(46), TRV(1531), ZIM(61). 

Aongstroemia julacea (Hook.) Mitt. LES(1531), 

MAD(40), MAL(67), NAT(1531), OFS(1531), 

REU(105), TAN(46). 

Aongstroemiopsis julacea (Dozy & Molk.) Fleisch. 

LES(1074,1531). 

Aphanorrhegma magdalenae (De Sloover) Ochyra. 

See Physcomitrella patens ssp. magdalenae. 

Aphanorrhegma is only found in the neotropics 

(Fife, pers. comm.) 

Archidium acanthophyllum Snider NAT(1531), 

TRV(1531), ZAI(65,30), ZAM(62), ZIM(61). 

Archidium africanum Mitt. = A. ohioense (1471) (= 

A. globiferum). 

Archidium alternifolium (Hedw.) Schimp. All speci- 

mens given this name from Africa south of the 

Sahara should be referred to A. ohioense (1471) 

(= A. globiferum). 

Archidium amplexicaule C.Müll. CBS(1531). 

Archidium andersonianum Snider CBS(1531). 

Archidium brevinerve P.Varde GAB(47,30). 

Archidium capense Hornsch. CBS(1531), 

LES(1531), OFS(1531), ZAI(65). 

Archidium dinteri (Irmsch.) Snider NAM(30,1531). 

Archidium globiferum (Brid.) Frahm BUF(1423), 

CBS(1531), CAR(47,30), IVO(63,1458), 

KEN(46), LES(1531), MAL(46,67), MAI(47,30), 

MAU(30), NAM(30,1531), NAT(1531), 

NIG(334,1492), NIR(79), SOM(30), SWA(1531), 

TAN(46), TOG(58,1878), TRV(1531), ZAM(62), 

ZIM(61). 

Archidium johannis-negrii Tong. Excluded. Snider 

(30) states that this taxon (from Eritrea (50)) is 

not an Archidium, and probably a member of the 

Bryaceae. 

Archidium julicaule C.Müll. CBS(1531), 

LES(1531). 

Archidium laxirete P.Varde CAR(47,30). 

Archidium leptophyllum P.Varde = A. ohioense (1471) 


Archidium microthecium Dix. & P.Varde 

NAM(1531), NAT(1531), TAN(46,30), 

TRV(1531), ZAM(411). 

Archidium muellerianum Snider CBS(1531), 

ZIM(61). 

Archidium ohioense Schimp. ex C.Müll. = A. globi- 

ferum (1458).

Archidium petrophyllum P.Varde = A. ohioense (1471) 


Archidium rehmannii Mitt. CBS(1531). 

Archidium subulatum C.Müll. CBS(1531). 

Archidium tenellum P.Varde = A. ohioense (1471) (= 

A. globiferum). 

Aschisma aethiopicum (Welw. & Duby) Lindb. = 

Bryoceuthospora aethiopica (1420). 

Astomiopsis amblyocalyx C.Müll. 

LES(1074,1471,1531). 

Astomum borbonicum Biz. & Onr. REU(105). 

Astomum borbonicum Biz. & Onr. The genus 

Astomum is considered synonymous with Weissia 

(1420), but the above taxon may belong to 

Trichostomum. It is left here pending revision. 

Atractylocarpus alticaulis (Broth.)Williams 

KEN(46), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Atractylocarpus capillifolius Dix. = A. alticaulis (804). 

Atractylocarpus flexifolius Dix. = A. alticaulis (465). 

Atractylocarpus madagascariensis (Thér.) Padberg 

& Frahm MAD(40), REU(411). 

Atrichum androgynum (C.Müll.) Jaeg. 

CBS(1531,289), LES(1531), MAD(40,289), 

MAL(46,67), MAU(105,289), MOZ(357), 

NAT(1531,289), OFS(1531), REU(105,289), 

TRV(1531,289), ZIM(46). 

Atrichum borbonicum Besch. = A. androgynum (289). 

Atrichum brevilamellatum Dix. & Thér. = Pogonatum 

congolense (289) 

Aulacomnium abyssinicum Schimp. ex Par. 

ETH(50). 

Aulacomnium palustre fo. perpapillosum P.Varde. 

Excluded. Any records for this form have been 

associated with the owning species. 

Aulacomnium palustre var. palustre (Hedw.) 

Schwaegr. ETH(50), KEN(46), UGA(46). 

Aulacomnium turgidum (Wahlenb.) Schwaegr. 

KEN(46). 

Aulacomnium turgidum var. papillosum Dix. 

KEN(46). 

Aulacopilum trichophyllum Aongstr. CBS(1531), 

NAT(1531), TRV(1531), UGA(46), ZIM(46,61). 

Barbella capillicaulis var. angustifolia Dix. & 

Thér. MAU(105). 

Barbella capillicaulis var. capillicaulis (Ren. & 

Card.) Card. MAD(40), MAU(105). 

Barbella eusebii Dix. ex Tosco & Piovano 

KEN(46). 

Barbula acuta (Brid.) Brid. = Didymodon rigidulus 

var. gracilis (1420). 

Barbula acutata C.Müll. = Pseudocrossidium 

replicatum (1420). 

Barbula afrofontana (C.Müll.) Broth. = B. ehrenbergii 

(1158). 

Barbula afrofontana var. acutiuscula P.Varde 

ZIM(46). 

Barbula aristata Bruch. & Schimp. ex Par., nom. 

nud. ETH(50). 

Barbula bolleana (C.Müll.) Broth. CAP(1296). 

Barbula calycina Schwaegr. CBS(1531). 

Barbula congoana Thér. TAN(46), UGA(46), 

ZAI(65). 

Barbula convoluta var. convoluta Hedw. 

KEN(411), TAN(46), ZAM(411). 

Barbula crinita Schultz. = Pseudocrossidium crinitum 

(1420). 

Barbula cylindrica var. flaccidus B.S.G. = Didymodon 

vinealis (1420). 

Barbula dioritica C.Müll. ERI(50). 

Barbula dusenii C.Müll. ex Broth. CAM(47). 

Barbula ehrenbergii (Lor.) Fleisch. CBS(1531), 

MAD(40), NAM(1531), NAT(1531), 

SAF(1158), TAN(46), TRV(1531), ZAI(65), 

ZAM(62). 

Barbula elliotii Broth. UGA(46). 

Barbula eubryum C.Müll. ANG(46), BUR(65), 

CAR(47), KEN(46), NAT(1531), RWA(65), 

SWA(1531), TAN(46), TRV(1531), UGA(46), 

ZAI(65), ZAM(62,411), ZIM(46,61). 

Barbula hornschuchiana Schultz = Pseudocrossidium 

hornshuchianum (1420). 

Barbula inclinans Schimp. ex Besch. MAU(105). 

Barbula indica var. indica (Hook.) Spreng. 

CAM(1158), CBS(1531), GAB(1158), KEN(46), 

MAD(40), MAL(67,411), MOZ(357), 

NAT(1531), ROD(105), RWA(65), SEY(40), 

SOC(40), SWA(1531), TAN(46), TRV(1531), 

ZAI(65,259), ZAM(411), ZIM(46,61). 

Barbula indica var. scaberrima Dix. TAN(46). 

Barbula kiaerii Broth. COM(40), MAD(40). 

Barbula kivuensis Leroy & P.Varde ZAI(65). 

Barbula lambarenense P.Varde = B. indica (1420). 

Barbula longirostris Hampe = Hennediella limbata 

(1420). 

Barbula lorentzii Kis. = B. indica (1158) (B. lorentzii 

is based on B. papillinervis (Lor.) Broth., hom. 

illeg. = B. indica. The valid name (before 

synonymisation) was B. isoindica Zander, nom. 

nov. for B. papillinervis). 

99

100 

Barbula madagassa Ren. & Card. MAD(40). 

Barbula meidensis Cuf. SOM(50). 

Barbula microcalycina Magill LES(1531), 

NAT(1531), OFS(1531). 

Barbula mucronulata Ren. & Card., hom. illeg. = B. 

indica (1158). 

Barbula nana C.Müll. = Trichostomum brachydontium 

(1660). 

Barbula pertorquata C.Müll. TAN(46), ZAM(46). 

Barbula porphyroneura C.Müll. in De Vent. = Pseu- 

docrossidium porphyroneurum (1420). 

Barbula pseudogracilis C.Müll. in De Vent. = B. acuta 

(1420) (= Didymodon rigidulus var. gracilis). 

Barbula punctulata (Ren. & Card.) Broth., hom. 

illeg. MAD(40). 

Barbula pygmaea C.Müll. TAN(46). 

Barbula rehmannii C.Müll. = B. indica (1158). 

Barbula salisburiensis Dix. ZIM(46,61). 

Barbula schweinfurthiana C.Müll. SOC(349,1626). 

Barbula somaliae C.Müll. SOM(50). 

Barbula stuhlmanii (Broth.) Brid = B. indica (1158). 

Barbula subobtusa Thér. MAD(40). 

Barbula subscabrinervis Dix. & Nav. = Bellibarbula 

recurva (1420). 

Barbula sulcata Geh. CAP(1296). 

Barbula tisserantii (P.Varde) P.Varde CAR(47). 

Barbula tuberculosa (Ren. & Car.) Card. 

MAD(40). 

Barbula umtaliensis Magill ZIM(46,61). 

Barbula unguiculata Hedw. ANG(46), SUD(1707). 

Barbula vinealis Brid. = Didymodon vinealis (1420). 

Barbula vinealis ssp. cylindrica (Tayl.) Podp.= Didy- 

modon vinealis (1420). 

Barbula zambesiaca Magill ZIM(46,61). 

Bartramia afro-ithyphylla Broth. KEN(46), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Bartramia angustissima Dix. UGA(46). 

Bartramia aprica C.Müll. ERI(50). 

Bartramia aristaria C.Müll. CBS(1531), 

NAT(1531), TRV(1531), ZIM(61). 

Bartramia boulayi Ren. & Card. MAD(40), 

REU(105). 

Bartramia brachypus Bruch & Schimp. ex C.Müll. 

ETH(50). 

Bartramia capensis (R.Br.) Wijk & Marg. 

CBS(1531), NAT(1531), TAN(46). 

Bartramia commutata Mitt. CAM(47). 

Bartramia compacta var. compacta Hornsch. 

CBS(1531). 

Bartramia compacta var. macowaniana (C.Müll.) 

Magill CBS(1531). 

Bartramia elliottii Broth. = Anacolia laevisphaera 

(46). 

Bartramia fragilicuspis Broth. MAD(40). 

Bartramia gigantea Bory TAN(46). 

Bartramia gigantea Bory. This plant was described 

from Réunion, but does not appear (in any obvious 

guise) in Crosby & Schultze-Motel (40). It is 

possible that it may be a synonym of B. patens. 

Bartramia halleriana Hedw. CAM(47), TAN(46). 

Bartramia hampeana C.Müll. CBS(1531), 

LES(1531), NAT(1531), OFS(1531), TAN(46), 

TRV(1531), ZIM(61). 

Bartramia ithyphylla Brid. TAN(46), ZAI(65). 

Bartramia jungneri Par. CAM(47), KEN(46), 

TAN(46), ZAI(65). 

Bartramia microcarpa P.Varde TAN(46). 

Bartramia patens Brid. MAD(40), REU(105). 

Bartramia radiculosa Mitt. = B. radicosa (1521). 

Bartramia ruwenzorensis Broth. KEN(46), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Bartramia stricta Hedw. CAM(47). 

Bartramia strictula C.Müll. KEN(46), TAN(46). 

Bellibarbula recurva (Griff.) Zander UGA(46), 

ZAI(65). 

Blindia acuta (Hedw.) BS&G KEN(46), ZAI(65). 

Blindia magellanica Schimp. ex C.Müll. 

LES(1531), REU(105). 

Brachydontium flexisetum Hampe RWA(65). 

Brachymenium abyssinicum (C.Müll.) Jaeg. 

ETH(50), ZAI(65). 

Brachymenium acuminatum Harv. ANG(46), 

BOT(1531), BUF(47), CAM(47), CBS(1531), 

CAR(47), ERI(50), GUI(47,1458), IVO(1458), 

LES(1531), MAD(40), MAL(67), MAU(105), 

NAM(1531), NAT(1531), NIG(47,334), 

OFS(1531), SWA(1531), TAN(46), TOG(1878), 

TRV(1531), UGA(46), ZAI(65), ZAM(46,62), 

ZIM(46,61). 

Brachymenium altipes Par. & Broth. = B. acuminatum 

(354). 

Brachymenium angolense (Welw. & Duby) Jaeg. 

ANG(46), KEN(46), MAL(67,411), NAM(1531), 

NAT(1531), TAN(46), TRV(1531), UGA(46), 

ZAI(65), ZAM(46,62), ZIM(46,61). 

Brachymenium angusti-limbatum Broth. & Par. = B. 

leptophyllum (65). 

Brachymenium angustirete Ochi ETH(354). 

Brachymenium anthracodictyon P.Varde = B. 

speirocladum (354).

Brachymenium apiculatum (P.Beauv.) Wijk & 

Marg. MAD(40), MAU(105), ROD(105). 

Brachymenium aureo-nitens Broth. & Par. = B. 

acuminatum (354). 

Brachymenium bequaertii Dix. & Thér. = B. 

flexifolium (354). 

Brachymenium borgenianum Hampe = B. 

acuminatum Harv (46). 

Brachymenium capitulatum (Mitt.)Par. CAM(47), 

CAR(47), DJI(1569), GHA(47), GUI(47), 

KEN(46), MAD(40), MAL(67), NIG(334), 

RWA(65), TAN(46), UGA(46), ZAI(65), 

ZAM(46). 

Brachymenium commutatum var. attenuatum 

Thér. & Trab. MAI(47). 

Brachymenium commutatum var. commutatum 

(C.Müll.) Jaeg. ERI(50). 

Brachymenium curvitheca Dix. ETH(50). 

Brachymenium dicranoides (Hornsch.) Jaeg 

CBS(1531), ZIM(46). 

Brachymenium djolibae Broth. & Par. = B. 


Brachymenium dusenii Broth. = B. acuminatum 

(354). 

Brachymenium elgonense Dix. KEN(46), UGA(46), 

ZAI(65). 

Brachymenium erythraeae Broth. ex Chiov. 

ERI(50). 

Brachymenium eurychelium Besch. MAU(105), 

REU(105). 

Brachymenium exile (Dozy & Molk.) Bosch & 

Lac. CAR(47), DJI(1569), ERI(50), GUI(47,1458), 

IVO(1458), MAD(40), NIG(334), SEN(1458), 

TAN(46), TOG(1878), ZAM(62), ZIM(46). 

Brachymenium flexifolium Bruch & Schimp. ex 

Jaeg. CAM(47), CON(354), ETH(50), KEN(46), 

ZAI(65). 

Brachymenium fusiforme Dix. = B. flexifolium (354). 

Brachymenium geniculatum Dix. = B. leptophyllum 

(354). 

Brachymenium leptophyllum (C.Müll.) B&S ex 

Jaeg. CAM(47), CAR(47), ETH(50), GAB(47), 

GUI(47,1458), IVO(63), KEN(46), MAD(40), 

MAL(67), MAS(40), NAT(1531), NIG(334), 

RWA(65), SIL(47), TAN(46), TOG(58,1878), 

TRV(1531), UGA(46), ZAI(65), ZAM(62), 

ZIM(46,61). 

Brachymenium lonchopus P.Varde = B. acuminatum 

(354). 

Brachymenium longicolle Thér. MAL(67), 

MAU(105), TAN(46). 

Brachymenium maclaudii (Broth. & Par.) P.Varde = 

B. acuminatum (65). 

Brachymenium madagassum Hampe MAD(40). 

Brachymenium megalacrion (Schwaegr.) Jaeg. = B. 

apiculata (1521). 

Brachymenium meruense (C.Müll.) Lindau 

TAN(46). 

Brachymenium mielichhoferioides C.Müll. in De Vent. 

= B. acuminatum (354). 

Brachymenium mildbraedii Broth. in Mild. = B. 


Brachymenium mildbraedii var. subelimbatum Thér. 

& P.Varde = B. leptophyllum. Ochi (23) does not 

mention this variety when he synonymises the 

main species with B. leptophyllum. Considering 

the variable nature of the plant, particularly the 

soft and easily-decaying leaves, I have assumed 

that the variety should also be referred here. 

Brachymenium minutifolium Broth. & P.Varde = B. 


Brachymenium molleri (C.Müll.) Kindb. = B. 

subuliferum (354). 

Brachymenium myurella (C.Müll.) Broth. GUI(47), 

KEN(46). 

Brachymenium nepalense Hook. FER(47), 

MAD(40), MAL(67), MAU(105), NAT(1531), 

RWA(65), SOC(40,1626), TRV(1531), 

ZAI(65,354). 

Brachymenium nigrescens Besch. MAD(40). 

Brachymenium pachyloma (R.C.) Thér. ZAI(65). 

Brachymenium philonotula Broth. BUR(65), 

KEN(46), MAD(40), TAN(46), ZAI(65). 

Brachymenium pobeguinii Broth. & Par. = B. 


Brachymenium procerrimum Broth. CAM(47), 

KEN(46), TAN(46), UGA(46). 

Brachymenium pulchrum Hook. CBS(1531), 

KEN(46), MAD(40), MAL(67), NAT(1531), 

REU(105), ROD(105), SWA(1531), TAN(46), 

TRV(1531), ZIM(46,61). 

Brachymenium recurvatum Broth. & P.Varde = B. 


Brachymenium revolutum Broth. CAM(47). 

Brachymenium rigidum Broth. & Par. ANG(46), 

GUI(47), KEN(46), MAD(40), RWA(65), 

TAN(46), UGA(46), ZIM(46,61). 

Brachymenium rigidum var. leptocarpum Broth. & 

Par. = B. capitulatum (354). 

Brachymenium rosulatum P.Varde, nom. illeg.. = B. 

101

102 

speirocladum (23). 

Brachymenium ruwenzorense Thér. & Nav. = B. 

nepalense (354). 

Brachymenium spathidophyllum Besch. 

REU(105). 

Brachymenium speirocladum C.Müll. ex Besch. 

CAM(47), CAR(47), COM(40), MAL(67), 

MAS(40), TAN(46), TOG(1878), UGA(46), 

ZAI(65). 

Brachymenium stereoneurum C.Müll. = B. exile (354). 

Brachymenium suberectum (Mitt.) Jaeg. = B. 

nepalense (354). 

Brachymenium subflexifolium var. pachypoma Ren. 

& Card (“pachyloma” in Crosby & Schultze- 

Motel, (40)) = B. nepalense (23). 

Brachymenium subuliferum (Mitt.) Jaeg. SAO(47), 

ZAI(65). 

Brachymenium torquescens Schimp. ex Par., nom. 

nud. ETH(50). 

Brachymenium turgiditheca Dix. = B. leptophyllum 

(354). 

Brachymitrion pocsii (A.Kop.) A.Kop. TAN(46). 

Brachythecium abyssinicum Par., nom. nud. 

ETH(50). 

Brachythecium afro-glareosum (Broth.) Par. 

KEN(46,136), TAN(46), UGA(46), ZAI(65). 

Brachythecium athrocladum Mitt. STH(1296). 

Brachythecium borgenii (Hampe) Jaeg. COM(40), 

KEN(46), MAD(40), MAU(105), REU(105), 

TAN(46), UGA(46). 

Brachythecium caespitans Schimp. ex Par. 

ETH(50). 

Brachythecium chauvetii Ren. & Card. REU(105). 

Brachythecium decurrens Card. MAD(40). 

Brachythecium dummeri Dix. KEN(46), RWA(65), 

TAN(46), UGA(46). 

Brachythecium gloriosum (C.Müll.) Kindb. = B. 

dummeri (46). 

Brachythecium hedbergii P.Varde RWA(65), 

TAN(46), UGA(46). 

Brachythecium implicatum (Hornsch.) Jaeg. 

BUR(65), CBS(1531), KEN(46), LES(1531), 

NAT(1531), OFS(1531), RWA(46,65), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZIM(46). 

Brachythecium inconditum Besch. MAS(40). 

Brachythecium mildeanum (Schimp.) Milde 

TAN(46), UGA(46). 

Brachythecium negrii Tosco & Piovano KEN(46). 

Brachythecium nigroviride (C.Müll.) Kindb. 

TAN(46), UGA(46). 

Brachythecium pinnatum Dix. CBS(1531). 

Brachythecium plumosum (Hedw.) BS&G 

CBS(1531), KEN(46), LES(1531), NAT(1531), 

RWA(65), TAN(146), TRV(1531), UGA(46), 

ZAI(65). 

Brachythecium populeum (Hedw.) BS&G 

CBS(1531), TAN(46). 

Brachythecium preussii (Broth.) Par. CAM(47). 

Brachythecium pseudopopuleum (C.Müll.) 

Schimp. CBS(1531). 

Brachythecium pseudovelutinum (C.Müll.) Jaeg. 

CBS(1531). 

Brachythecium ramicola Broth. KEN(46), 

RWA(46), TAN(46), UGA(46), ZAI(65). 

Brachythecium rigens Buck RWA(65). 

Brachythecium rivulare BS&G KEN(46). 

Brachythecium roccatii Negri UGA(46). 

Brachythecium rutabulum BS&G TAN(146). 

Brachythecium salebrosum (Web. & Mohr) BS&G 

CBS(1531), MAL(67), NAT(1531), RWA(65), 

SWA(1531), TRV(1531), ZAI(65), ZIM(61). 

Brachythecium spectabile Broth. KEN(46), 

RWA(46), TAN(46), UGA(46), ZAI(65). 

Brachythecium starkei var. sublaeve Thér. 

ZAI(65). 

Brachythecium subfurcatum C.Müll. ex Par., nom. 

nud. ETH(50). 

Brachythecium sublaetum Broth. RWA(65), 

TAN(146). 

Brachythecium subrutabulum (C.Müll.) Jaeg. 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TRV(1531). 

Brachythecium ugandae Dix. KEN(46), TAN(46), 

UGA(46). 

Brachythecium valentinii Besch. REU(105). 

Brachythecium vellereum (Mitt.) Jaeg. CAM(47), 

ETH(50), KEN(46,136), RWA(46,65), TAN(46), 

UGA(46), ZAI(65). 

Brachythecium vesiculariopsis Dix. CAM(47). 

Braunia arbuscula (Welw. & Duby) Gepp 

ANG(46), KEN(46), TAN(46), UGA(46), 

ZAI(65). 

Braunia brachytheca (Dix.) Broth. = B. secunda 

(1521). 

Braunia camptoclada P.Varde & Thér. KEN(46), 

TAN(46), ZAI(65). 

Braunia entodonticarpa C.Müll. COM(40), 

KEN(46). 

Braunia laevicuspis Broth. ex Engl. ETH(50). 

Braunia rupestris (Mitt.) Jaeg. CAM(47), ETH(50).

Braunia schimperi BS&G = B. schimperianum (1521). 

Braunia schimperiana (Hampe) Jaeg. ERI(50), 

ETH(50), KEN(46), TAN(46). 

Braunia schimperiana var. latifolia Broth. ex 

Engl. ETH(50). 

Braunia secunda (Hook.) BS&G ANG(46), 

CBS(1531), KEN(46), MAL(67), MOZ(357), 

NAT(1531), RWA(65), SWA(1531), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZIM(46,61). 

Braunia secunda fo. longipila Dix. ex Tosco & 

Piovano. Excluded. Any records for this form 


Braunia secunda fo. xerophitica Dix. ex Tosco & 



Breutelia abyssinica Schimp. ex Par. Excluded. Fide 

De Sloover, (282) (nomen nudum). 

Breutelia angustifolia Rehm. ex Sim = B. microdonta 

(829). 

Breutelia borbonica De Sloover REU(105), 

TAN(46). 

Breutelia diffracta Mitt. CAM(47), CBS(1531), 

KEN(46), LES(1635), MAD(40), MAL(67,411), 

NAT(1531), RWA(65), TAN(46), TRV(1531), 

UGA(46), ZAI(65), ZIM(61). 

Breutelia elliptica Magill CBS(1531). 

Breutelia gnaphalea (P.Beauv.) Mitt. CAM(47), 

MAD(40), MAL(67), REU(105). 

Breutelia gracillima Broth. RWA(65), UGA(46), 

ZAI(65). 

Breutelia humbertii P.Varde & Thér. KEN(46), 

RWA(65), TAN(46). 

Breutelia kilimandscharica (C.Müll.) Par. Excluded. 

Fide De Sloover, (282) (lacking adequate descrip- 

tion and type). 

Breutelia madagassa var. kivuensis P.Varde 

ZAI(65). 

Breutelia madagassa var. madagassa Thér. 

MAD(40). 

Breutelia magdalenae De Sloover MAU(105), 

REU(105). 

Breutelia microdonta (Mitt.) Broth. 

CBS(1531,1882), NAT(1531), OFS(1531), 

TRV(1531), ZIM(61). 

Breutelia muhavurensis P.Varde & Thér. 

KEN(46,259), RWA(65), TAN(46,259). 

Breutelia perrieri Thér. MAD(40), REU(105), 

TAN(46). 

Breutelia sciuroides Thér. MAD(40). 

Breutelia sclerodictya Card. MAD(40). 

Breutelia stenodictyon (Ren. & Card.) Broth. 

MAD(40), REU(105). 

Breutelia stricticaulis Dix. KEN(46), RWA(65), 

UGA(46), ZAI(65). 

Breutelia stuhlmannii Broth. KEN(46), MAD(40), 

MAS(40), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Breutelia subgnaphalea (C.Müll.) Kindb. = B. diffracta 

(65). 

Breutelia substricta (C.Müll.) Magill 

CBS(1531,1882), LES(1531), NAT(1531), 

OFS(1531). 

Breutelia tabularis Dix. ex Sim. CBS(1531). 

Breutelia tomentosa (Brid.) Jaeg. Excluded. Fide De 

Sloover, (282) (doubtful for the area). 

Breutelia viguieri Card. MAD(40). 

Brothera leana (Sull.) C.Müll. MAL(134). 

Bruchia brevipes Harv. ex Hook. CBS(1531). 

Bruchia eckloniana C.Müll. CBS(1531). 

Bruchia queenslandica Stone NAM(1531). 

Bryobartramia novae-valesiae (Broth.) Stone & 

Scott CBS(1531). 

Bryoceuthospora aethiopica (Welw. & Duby) 

Zander ANG(46). 

Bryoerythrophyllum afrorubellum (Broth. & Wag.) 

De Sloover = B. campylocarpum (1420). 

Bryoerythrophyllum alpigenum (Vent.) Chen. See B. 

recurvirostrum var. alpigenum (Sollman. pers. 

comm.) 

Bryoerythrophyllum campylocarpum (C.Müll.) 

Crum ANG(259), BUR(65), CAM(47), 

CBS(1531), CAR(47), CHA(47), GAB(47), 

GHA(47), GUI(47), IVO(44), KEN(46), 

LES(1531), MAL(67), NAT(1531), OFS(1531), 

RWA(65), SIL(47), TAN(46), TOG(58), 

TRV(1531), UGA(1158,46), ZAI(47,65), 

ZAM(62), ZIM(61). 

Bryoerythrophyllum ferruginascens (Stirt.) Giac. 

ETH(1862), KEN(1862), TAN(1862). 

Bryoerythrophyllum inaequalifolium (Tayl.) 

Zander ETH(1158). 

Bryoerythrophyllum jamesonii (Tayl.) Crum auct. 

afr. = B. campylocarpum (1420). 

Bryoerythrophyllum recurvirostrum (Hedw.) 

Chen CBS(1531), ETH(50), KEN(46,136), 

LES(1531), MOZ(357), NAT(1531), RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Bryoerythrophyllum recurvirostrum var. alpigenum 

(Vent.) Giac. KEN(46), TAN(46), UGA(46). 

Bryoerythrophyllum recurvirostrum var. 

103

104 

angustifolium (P.Varde) Wijk & Marg. = B. 

afrorubellum (1420) (= B. campylocarpum). 

Bryoerythrophyllum rubrum (Jur. ex Geh.) Chen 

TAN(46). 

Bryoerythrophyllum rubrum (Jur. ex Geh.) Chen. 

Sollman (pers. comm.) believes that the occurrence 

of this taxon in Africa is doubtful, and that the 

material should be checked. 

Bryohumbertia filifolia (Hornsch.) Frahm 

MAD(105), MAU(105), REU(105). 

Bryohumbertia flavicoma (C.Müll.) Frahm 

ANG(146), BUR(65), CAM(47), RWA(65), 

TAN(46), UGA(46), ZAI(65), ZAM(62). 

Bryosedgwickia barlassinae Tosco & Piovano 

KEN(46). 

Bryosedgwickia densa (Hook.) Biz. & P.Varde 

CAR(47), GUI(47), NIG(334), ZAI(65), 

ZAM(46). 

Bryotestua brevicuspis Thér. & P.Varde CAR(47). 

Bryotestua longicuspis Thér. & P.Varde CAR(47). 

Bryum afro-plumosum Broth. & Par. = B. apiculatum 

(354). 

Bryum afrocalophyllum P.Varde KEN(46), 

TAN(46). 

Bryum albo-marginatum Card., hom. illeg. 

MAD(40). 

Bryum albo-pulvinatum C.Müll. ANG(46), 

CAM(47), CAR(47), GAB(47), GUI(47), 

KEN(46), NIG(47), SIL(47), ZAI(65). 

Bryum albolimbatum Jaeg. MAD(40), TAN(46). 

Bryum algovicum var. rutheanum (Warnst.) 

Crundw. KEN(46), TAN(46). 

Bryum alpinulum var. latinerve P.Varde & Thér. 

MAD(40). 

Bryum alpinulum var. levieri Ren. & Card. 

MAD(40). 

Bryum alpinum Huds. ex With. CAM(47), 

CBS(1531), KEN(46), LES(1531), MAD(40,23), 

MAL(67), NAT(1531), OFS(1531), TAN(46), 

TRV(1531), UGA(46), ZIM(61). 

Bryum alpinum var. perrevolutum P.Varde, nom. 

inval. KEN(46), TAN(46), UGA(46). 

Bryum alpinum var. viride Husn. KEN(46), 

TAN(46). 

Bryum andicola Hook. CBS(1531), CAR(21), 

KEN(46), LES(1531), MAL(67), NAT(1531), 

OFS(1531), SIL(47), SWA(1531), TAN(46,21), 

TRV(1531), ZAI(65,21), ZIM(61). 

Bryum anomodon Mont. CAP(1296). 

Bryum apiculatum Schwaegr. BUF(1423), 

CAM(47), CBS(1531), CAR(47), COM(40), 

FER(47), GAB(47), GUI(47), IVO(44,1880), 

KEN(46), MAD(40), MAU(105), MOZ(357), 

NAM(1531), NAT(1531), OFS(1531), REU(105), 

ROD(105), SAO(47), SEY(40), SIL(47), 

TAN(46), TOG(1878), TRV(1531), ZAI(65), 

ZAM(62). 

Bryum appressum Ren. & Card. MAD(21), 

SAF(21). 

Bryum arachnoideum C.Müll. ANG(46), BUF(47), 

BUR(65), IVO(1458), KEN(46), MAD(40), 

MAL(67), NIG(334), RWA(65), TAN(46), 

UGA(46), ZAM(62), ZIM(46). 

Bryum areoblastum C.Müll. = B. apiculatum (354). 

Bryum argenteum fo. julaceum Hampe. Excluded. 

Any records for this form have been associated 

with the owning species. 

Bryum argenteum fo. laxifolium Thér. Excluded. 



Bryum argenteum fo. majus (Schwaegr.) Podp. Ex- 

cluded. Any records for this form have been 


Bryum argenteum var. argenteum Hedw. ANG(46), 

ASC(23,1296), BOT(1531), BUR(65), CAM(47), 

CBS(1531), CAR(47), COM(40), ERI(50), 

ETH(50), GAB(47), GUI(47), KEN(46), 

LES(1531), MAD(40,23), MAL(67), MAU(105), 

MOZ(357), NAM(1531), NAT(1531), 

NIG(23,334), OFS(1531), REU(23,105), 

RWA(65), SWA(1531), TAN(46), 

TOG(58,1878), TRV(1531), UGA(46), ZAI(65), 

ZAM(46), ZIM(46,61). 

Bryum argenteum var. argyrotrichum (C.Müll.) Mitt. 

= B. argenteum var. lanatum (1521). 

Bryum argenteum var. australe Rehm. ex Dix. 

KEN(46), ZIM(46). 

Bryum argenteum var. lanatum (P.Beauv.) Hampe 

BUR(65), CAM(47), CAR(47), CHA(47), 

COM(40), ETH(50), GAB(47), KEN(46,136), 

MAD(40), MAL(67), MAU(105), REU(105), 

RWA(65), SAO(47), STH(1296), TAN(46), 

ZAI(65), ZIM(46). 

Bryum argenteum var. proliferum Sim ZIM(46). 

Bryum argenteum var. rotundifolium Sim ZIM(46). 

Bryum argenteum var. submuticum Besch. 

REU(105). 

Bryum argentisetum C.Müll. TAN(46). 

Bryum argyrotrichum C.Müll. = B. argenteum var. 

lanatum (1521).

Bryum aubertii (Schwaegr.) Brid. BUR(65), 

CBS(1531), IVO(1458), LES(1531), MAD(40), 

MAS(40), NAT(1531), OFS(1531), TAN(46), 

TRV(1531), ZAI(65). 

Bryum aulacomnioides C.Müll. MAD(40). 

Bryum bessonii Ren. & Card. ANG(46), COM(40), 

MAD(40). 

Bryum bicolor Dicks. BOT(1531), CBS(1531), 

MAD(40), NAM(1531), NAT(1531), OFS(1531), 

TAN(46), TRV(1531), ZIM(46,61). 

Bryum billardieri Schwaegr. MAD(40), 

MAU(105), REU(105). 

Bryum bulbillicaule C.Müll. KEN(46). 

Bryum bullosum C.Müll. ex Broth. CAM(47). 

Bryum cadetii Biz. & Onr. REU(105). 

Bryum caespiticium var. caespiticium Hedw. 

ANG(46), CBS(1531), LES(1531), NAT(1531), 

UGA(46). 

Bryum campylopodioides C.Müll., nom. nud. 

MAU(105). 

Bryum canariense Brid. CBS(1531), GUI(47,23), 

LES(1531), NAT(1531), OFS(1531), TAN(46), 

TRV(1531), ZAM(62), ZIM(46,61). 

Bryum candidum C.Müll. = B. argenteum (1521). 

Bryum capillare var. capillare Hedw. BOT(1531), 

CAM(47), CBS(1531), COM(40,23), ERI(50), 

ETH(50,23), GUI(47,23), KEN(46), LES(1531), 

MAD(40,23), MAL(67), MAI(67), MOZ(46), 

NAM(1531), NAT(1531), OFS(1531), SEY(40), 

SIL(47,23), SUD(23), SWA(1531), TAN(46), 

TOG(1878), TRV(1531), UGA(46), ZAI(65), 

ZIM(46). 

Bryum capillare var. torquescens (De Not.) Husn. = 

B. torquescens (1520). 

Bryum cellulare Hook. LES(1531), NAM(1531), 

NAT(1531), OFS(1531), REU(354,105), 

TRV(1531), ZIM(61). 

Bryum cellulare ssp. lipopae (Thér. & P.Varde) 

P.Varde = B. euryphyllum (354). 

Bryum chalarorhodon C.Müll. ex Broth. = B. huillense 

(354). 

Bryum coronatum Schwaegr. ANG(46), 

ASC(1296), BUR(65), CAM(47), CAR(47), 

CHG(40), FER(47), GAB(47), GHA(47), GUI(47), 

IVO(47,1458,1880), KEN(46), LIB(47), 

MAD(40), MAL(67), MAU(105), MOZ(357), 

NIR(79), NIG(47,334), REU(105), RWA(65), 

SAO(47), SEN(47), SIL(47), TAN(46), 

TOG(58,1878), UGA(46), ZAI(65), ZAM(46,62), 

ZIM(61). 

Bryum coronatum var. latifolium Broth. & P.Varde 

CAR(47). 

Bryum cyathiphyllum Dix. & Herz. UGA(46), 

ZAI(65). 

Bryum depressum C.Müll. ex Broth. CAM(47), 

GUI(1458), IVO(1458), SIL(47), ZAI(65). 

Bryum dichotomum Hedw. SOC(40). 

Bryum dongolense Brizi ERI(50). 

Bryum donianum Grev. CBS(1531). 

Bryum elatum Broth. & Par. = B. rhypariocaulon 

(354). 

Bryum ellipsifolium C.Müll. KEN(46), TAN(46), 

UGA(46). 

Bryum ellipticifolium var. ellipticifolium Brizi 

ETH(50). 

Bryum erythrocarpulum C.Müll., nom. nud. 

MAU(105). 

Bryum erythrocarpum var. madagassum Ren. & 

Card., nom. illeg. MAD(40). 

Bryum erythrocaulon (Schwaegr.) Brid. CAM(47), 

CBS(1531), MAD(21,40), MAU(21,105), 

NAT(1531), ROD(105), SIL(47), TAN(21), 

TRV(1531), ZAI(65). 

Bryum euryphyllum Dix. & P.Varde GAB(47), 

TAN(46), ZAI(65). 

Bryum fluminale C.Müll. ex Broth. = B. erythrocaulon 

(21). 

Bryum fouta-djallonii Par. = Brachymenium exile 

(354). 

Bryum fragiferum P.Varde = B. apiculatum (354). 

Bryum fragiferum var. angustifolium P.Varde = B. 

apiculatum (354). 

Bryum fusco-tomentosum Demar. & Leroy. Excluded. 

According to Ochi, the only collection of this 

taxon (from Zaire) is not a Bryum, but probably 

an Entosthodon (354). 

Bryum grammothecium C.Müll., nom. nud. 

COM(40). 

Bryum griseum Dus. ex Thér. = B. argenteum (354). 

Bryum haematoneurum C.Müll. ERI(50), TAN(46). 

Bryum hedbergii P.Varde KEN(46), TAN(46). 

Bryum homalobolax C.Müll. ex Ren. = B. huillense 

(46). 

Bryum huillense Welw. et Duby ANG(46), 

BUR(65), CAM(47,23), CAR(47), COM(40), 

ETH(23), GUI(47,23), IVO(44,47), KEN(46), 

MAD(40,23), MAL(67), MAU(23,105), 

NIG(334), SAO(47,23), TAN(46), TOG(58,1878), 

UGA(46), ZAI(65), ZAM(62). 

Bryum interruptinerve C.Müll., nom. nud. 

105

106 

COM(40). 

Bryum keniae C.Müll. BUR(65), CAM(47), 

CBS(1531), GUI(47), KEN(46), MAL(67), 

MOZ(46), NAT(1531), RWA(65), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZAM(46), ZIM(61). 

Bryum kilimandscharicum (C.Müll.) Zander. Exclu- 

ded. See note under Phascum kilimandscharicum. 

Bryum kindiae Par. & Broth. = B. canariense (354). 

Bryum klinggraeffii Schimp. SUD(1707). 

Bryum leptoneurum P.Varde KEN(46), TAN(46). 

Bryum leptospeiron C.Müll. COM(40), MAD(40), 

MAU(105), SEY(40). 

Bryum leptotorquescens C.Müll. ex Broth. 

ANG(46), CAM(47), CAR(47), SIL(47), 

UGA(46). 

Bryum lonchophyllum Broth. CAM(47). 

Bryum madagasso-ramosum Broth. = B. 

erythrocaulon (46). 

Bryum megalacrion Schwaegr. MAD(40), 

MAS(40). 

Bryum micro-erythrocarpum C.Müll. & Kindb. 

CBS(1531), NAT(1531), TRV(1531). 

Bryum microdontium Dix. hom. illeg. = B. andicola 

(21). 

Bryum microdontium var. compactum P.Varde = B. 

andicola (21). 

Bryum minutirosatum C.Müll. KEN(46), TAN(46). 

Bryum muehlenbeckii BS&G ZAI(65). 

Bryum mundtii C.Müll. ZIM(46). 

Bryum nanocapillare C.Müll. ERI(50). 

Bryum nanotorquescens Broth. & Par. = B. capillare 

(354). 

Bryum nigerianum var. germainii P.Varde & Demar. 

= B. capillare (354). 

Bryum nigerianum var. nigerianum Broth. & Par. = 

B. capillare (354). 

Bryum nitens Hook. = B. apiculatum (46). 

Bryum ocobiense Broth. & P.Varde = B. apiculatum 

(354). 

Bryum orthocladon Bruch & Schimp. in Par., nom. 

nud. ETH(50), KEN(23). 

Bryum pallens var. pallens Sw. RWA(65). 

Bryum pallescens Schleich. ex Schwaegr. 

CAM(47), ZAI(65). 

Bryum peralatum P.Varde. Excluded. According to 

Ochi, the only collection of this taxon (from 

Zaire) is not a Bryum, but probably an Entosthodon 

(354). 

Bryum perlimbatum Card. Excluded. Ochyra (1442) 

concludes that the specimen was collected in 

Puerto Natale, Chile, not in Natal, South Africa. 

Bryum perrieri Thér. MAD(40). 

Bryum perspinidens Broth. KEN(354), UGA(354). 

Bryum petrophilum P.Varde = B. billardieri Schwaegr 

(354) = B. andicola (21). 

Bryum piliferum Bruch & Schimp. in Par., nom. 

nud. ETH(50). 

Bryum plano-marginatum Dix. = B. pseudotriquetrum 

(46). 

Bryum plumosum Doz. & Molk. ROD(105). 

Bryum pobeguinii Broth. & P.Varde = B. capillare 

(354). 

Bryum pottiaefolium C.Müll. = B. capillare (354). 

Bryum preussii Broth. CAM(47), KEN(1408,411), 

ZAI(65). 

Bryum pseudotorquescens C.Müll. = B. capillare 

(354). 

Bryum pseudotriquetrum var. bimum (Brid.) Lilj. 

SUD(1419). 

Bryum pseudotriquetrum var. pseudotriquetrum 

(Hedw.) Schwaegr. CBS(1531), LES(1531), 

NAT(1531), OFS(1531), RWA(65), TAN(354), 

TRV(1531), ZAI(65), ZAM(62), ZIM(61). 

Bryum purpureo-nigrum Duby MAD(40). 

Bryum pycnophyllum (Dix.) Mohamed CBS(1531), 

LES(1531), NAT(1531), OFS(1531), 

SWA(1531), TAN(46), TRV(1531), ZAM(62), 

ZIM(46,61). 

Bryum radiculosum Brid. CBS(1531), NAM(1531). 

Bryum rhypariocaulon C.Müll. CAR(47), FER(23), 

GUI(47,23), TOG(1878), UGA(46). 

Bryum robustum Hampe MAD(40). 

Bryum roscheri Lor. TAN(46). 

Bryum rosulatulum C.Müll. = B. andicola (21). 

Bryum rubicundum C.Müll. Excluded - according to 

Ochi (23), this is probably not a Bryum, but the 

specimen is too fragmentary to determine. 

Bryum rubro-costatum C.Müll. Excluded - according 

to Ochi (23) this is probably a Mielichhoferia, but 

the specimen is too scanty to be more definite. 

Bryum semense Par., nom. nud. = B. orthocladon 

(354). 

Bryum spiralifolium Dix. = B.andicola (21). 

Bryum splendidifolium C.Müll. ERI(50). 

Bryum staudtii Broth. CAM(47), MAL(67), 

SIL(47). 

Bryum stenophyllum Dix. UGA(46). 

Bryum subapiculatum Hampe TAN(46). 

Bryum subappressum var. forsythii (Ren. & Card.)


Bryum subargenteum Hampe MAD(40), TAN(46), 

ZAI(65). 

Bryum subdepressum Broth. & Par. = B. 

rhypariocaulon (354). 

Bryum subhomalobolax Ren. & Card. = B. huillense 

(354). 

Bryum subturbinatum C.Müll. = B. turbinatum (23). 

Bryum taitae C.Müll. KEN(46). 

Bryum tenerum P.Varde. Excluded. According to 

Ochi, the type is not referable with certainty to 

a specific taxon (354). 

Bryum thomeanum P.Varde SAO(47). 

Bryum tisserantii P.Varde CAR(47), GHA(47). 

Bryum tisserantii var. sublimbatum P.Varde. Excluded. 

According to Ochi, the type of this taxon (from 

Zaire) is probably a member of the Mniaceae (23). 

Koponen (1036) disputes this and states that it 

belongs to the Bryaceae. The status of this taxon 

is thus uncertain. 

Bryum torquescens Bruch ex De Not. CBS(1531), 

ETH(50), KEN(46), LES(1531), NAT(1531), 

OFS(1531), SUD(1419), TAN(46), TRV(1531), 

ZIM(46). 

Bryum truncorum (Brid.) Brid. MAD(21,40), 

MAU(298,105), REU(21,105), ROD(105), 

TAN(46), ZIM(46). 

Bryum turbinatum (Hedw.) Turn. CBS(1531), 

ETH(50), LES(1531), NAT(1531), OFS(1531). 

Bryum turbinitiforme Schimp. & Par., nom. nud. 

ETH(50). 

Bryum umbraculum Bruch. ex Hook. CBS(1531), 

ETH(23), KEN(46), NAT(1531), OFS(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZIM(46). 

Bryum usambaricum Dix. = B. coronatum (46). 

Bryum viguieri Thér. ANG(46), MAD(40), 

MAL(67), TAN(46). 

Bryum viridescens Welw. & Duby ANG(46), 

CBS(1531), NAT(1531), SWA(1531), 

TRV(1531), ZIM(61). 

Bryum voeltzkowii Broth. MAD(21,40), TAN(46). 

Bryum zygodontoides C.Müll. = B. coronatum (1521). 

Caduciella mariei (Besch.) Enroth COM(40), 

TAN(1439). 

Callicostella (C.Müll.) Mitt. Menzel (1750) summa- 

rises the position over the use of this name. 

Crosby pointed out the priority of Schizomitrium 

Schimp. over Callicostella, but Koponen and 

Isoviita proposed the conservation of Callicostella, 

to avoid the nomenclatural changes. This 

proposal was accepted by the Committee for 

Bryophyta of IAPT, for submission to the 1993 

Tokyo Congress, and confimed there. 

Nevertheless, in the interim, a large number of 

nomenclatural changes were made. These have 

been reversed in this checklist. 

Callicostella africana Mitt. CAM(47), CAR(47), 

COM(40), CON(47), FER(47), GAB(47), 

GHA(47), IVO(1880,47), LIB(47), MAD(40), 

NIG(47), TAN(46), UGA(46), ZAI(65), 

ZAM(62). 

Callicostella ascensionis (C.Müll.)Broth. 

ASC(1296). 

Callicostella attenuata (C.Müll.) Kindb. 

CAM(47), CAR(47), GAB(47), LIB(47), ZAI(65). 

Callicostella brevipes (Broth.) Broth. CAM(47), 

CAR(47), CON(47), FER(47), GAB(47), 

GHA(47), IVO(1880), MAD(40), MAU(105), 

SAO(47), SEY(40), SIL(47), TAN(46), UGA(46), 

ZAI(65). 

Callicostella chevalieri Mitt. CAR(47), GAB(47), 

NIG(47), ZAI(65). 

Callicostella chionophylla (C.Müll.) Kindb. 

SAO(47). 

Callicostella constricta (C.Müll.) Kindb. 

CAM(47), CAR(47), COM(40), FER(47), 

GAB(47), IVO(1880), MAD(40), NIG(47), 

TAN(46), ZAI(65). 

Callicostella emarginatula Broth. in Corb. 

ANG(46), CAM(47), CAR(47), CON(47), 

GAB(47), ZAI(65). 

Callicostella eroso-truncata Card. CAM(47), 

CAR(47), CON(47), GAB(47), IVO(1880), 

MAU(105), NIG(47), TAN(46), ZAI(65). 

Callicostella fissidentella (Besch.) Kindb. 

IVO(1880), LIB(47), MAD(40), MAU(105), 

PRI(47), REU(105), SAO(47), TAN(46), 

ZAI(65). 

Callicostella gabonensis Broth. & P.Varde 

CAR(47), CON(47), GAB(47), IVO(1880), 

ZAI(65). 

Callicostella lacerans (C.Müll.) Jaeg. COM(40), 

LIB(47), MAD(40), MAU(105), TAN(46). 

Callicostella laeviuscula Mitt. CAM(47), 

GAB(47), ROD(105). 

Callicostella leptocladula (Broth.) Broth. 

CAM(47), MAD(40). 

Callicostella maclaudii (Par. & Broth.) Broth. 

GUI(47). 

107

108 

Callicostella papillata (Mont.) Mitt. COM(40), 

MAD(40), MAS(40). 

Callicostella papillata var. brevifolia Fleisch. 

MAD(40). 

Callicostella papillosula Broth. & P.Varde 

CAR(47), CON(47), GAB(47). 

Callicostella parvocellulata Mitt. MAD(40). 

Callicostella parvula Broth. in Mildbraed, nom. 

nud. ANN(47). 

Callicostella parvula Broth. in Mildbraed, nom. nud. 

Mildbraed (1619) gives a locality (Santa Mina 

Gipfel) and a collection number [6706] from a 

German expedition of 1911-12, so there is hope 

that the specimen can be found. 

Callicostella perpapillata Broth. & P.Varde 

CAM(47), CAR(47), CON(47), GAB(47), 

IVO(47), PRI(47), TAN(46), UGA(46), ZAI(65). 

Callicostella perrotii (Par.) Broth. IVO(1880), 

MAD(40). 

Callicostella pusilla Demar. & P.Varde CAM(47), 

CAR(47), GHA(47), IVO(1880), MAD(40), 

UGA(46), ZIM(61). 

Callicostella robillardei Fleisch., nom. nud. 

MAU(105). 

Callicostella salaziae (Besch.) Kindb. CAM(47), 

REU(105), SAO(47). 

Callicostella seychellensis (Besch.) Kindb. 

ANN(47), CAM(47), CAR(47), GAB(47), 

GUI(47), IVO(1880), LIB(47), MAD(40), 

MAU(105), SEY(40), ZAI(65). 

Callicostella subemarginatula Broth. & P.Varde 

ANG(46), CAM(47), CAR(47), CON(47), 

GAB(47), GUI(47), TOG(1878), ZAI(65). 

Callicostella tristis (C.Müll.) Broth. CBS(1531), 

CAR(47), MAI(47), NAT(1531), SWA(1531), 

TRV(1531). 

Callicostella usambarica (C.Müll.) Broth. 

CAR(47), GAB(47), IVO(1880), TAN(46). 

Calliergon keniae Dix. = Sarmentypnum sarmentosum 

(1884). 

Calliergon sarmentosum (Wahlenb.) Kindb. = Sar- 

mentypnum sarmentosum (453). 

Calliergon sarmentosum var. subflavum Dix. = Sar- 

mentypnum sarmentosum (1884). 

Calliergon stramineum (Brid.) Kindb. Excluded. 

Ochyra (pers. comm.) believes that the sub-fossil 

specimen cited by Spence & Mahaney, (1497) is 

likely to be a form of Sarmentypnum 

sarmentosum. The voucher specimen cannot 

now be located at UBC. 

Calymperes acanthoneuron P.Varde = C. decolorans 

(1473) (= C. pallidum). 

Calymperes afzelii Sw. ANG(46), BUR(65), 

CAM(57), CAR(57), COM(40), GAB(57), 

GHA(57), GUI(57,1458), IVO(44,1458,1880), 

KEN(46), MAD(40), NIG(57,334), SAO(57), 

SIL(57), TAN(46), TOG(1878), UGA(46), 

ZAI(57,65), ZAM(62), ZIM(46,61). 

Calymperes annobonense Broth. in Mildbr., nom. 

nud. ANN(47). 

Calymperes ascensionis C.Müll. = C. erosum (1473). 

Calymperes asperum C.Müll. ex Besch. = C. erosum 

(57). 

Calymperes asterigium C.Müll. ex Besch. = C. palisotii 

ssp. palisotii (57). 

Calymperes asteristylium C.Müll. ex Besch. = C. 

afzelii (57). 

Calymperes borgenii Kiaer ex Besch. = C. hispidum 

(1371). 

Calymperes brachypelma C.Müll. ex Besch. = C. 

palisotii ssp. palisotii (57). 

Calymperes brevifolium Card. = C. palisotii (1624). 

Calymperes cacazouense Besch. = C. tenerum (1373). 

Calymperes campylopodioides C.Müll. ex Besch. = C. 

palisitii var. palisotii (57). 

Calymperes casamancae Card. = C. erosum (1473). 

Calymperes caudatum C.Müll. = C. tenerum (1473). 

Calymperes cernense Besch. MAD(40). 

Calymperes chevalieri Thér. & Corb. = C. decolorans 

(1473) (= C. pallidum). 

Calymperes chevalieri var. angustifolium Thér. & 

P.Varde = C. decolorans (1473) (= C. pallidum). 

Calymperes chrysoblastum C.Müll. = C. erosum (57). 

Calymperes cochlearifolium Par. = C. palisotii (1473). 

Calymperes congolense (Ren. & Card.) Card. = C. 

decolorans (1473) (= C. pallidum). 

Calymperes conterminum Thér. & P.Varde = C. 

lonchophyllum ssp. microblastum (57). 

Calymperes corbieri Thér. = C. decolorans (1473) (= 

C. pallidum). 

Calymperes crassilimbatum Ren. & Card. = C. hispidum 

(1511). 

Calymperes decolorans C.Müll. = C. pallidum (1518). 

Calymperes decolorans var. latifolium Ren. & Card. 

= C. decolorans (1473) (= C. pallidum). 

Calymperes dilatatum Besch. = C. decolorans (1473) 

(= C. pallidum). 

Calymperes disjunctum Besch. = C. palisotii (1370). 

Calymperes dusenii C.Müll. ex Besch. = C. afzelii 

(57).

Calymperes erosulum Ren. & Par. = C. decolorans 

(1473) (= C. pallidum). 

Calymperes erosum C.Müll. ANG(57), ASC(1296), 

BEN(47), CAM(57), GAB(57), GHA(57), 

GUI(57,1458), IVO(1458,1880), LIB(57), 

MAD(40), NIG(57,334), SEN(57), SEY(40), 

SIL(57), TAN(46), TOG(1878), ZAI(57,65,1552). 

Calymperes gabonense Thér. & P.Varde = C. erosum 

(1473). 

Calymperes garciae Mitt. = C. graeffeanum (1371). 

Calymperes gilletii Ren. & Card. = C. erosum (1473). 

Calymperes graeffeanum C.Müll. CHG(40), 

MAU(105), SEY(40). 

Calymperes heudelotii Besch. = C. erosum (57). 

Calymperes hildebrandtii C.Müll. = C. afzelii (1473). 

Calymperes hispido-costatum Dix. = C. pallidum 

(1518). 

Calymperes hispidum Ren. & Card. MAD(40), 

MAU(105), REU(105), SEY(1371). 

Calymperes hyalino-limbatum Par. = C. afzelii (1473). 

Calymperes incertum Card. = C. palisotii (1370). 

Calymperes integrifolium C.Müll. = C. afzelii (1473). 

Calymperes intra-limbatum C.Müll. = C. erosum 

(57). 

Calymperes isleanum Besch. = C. motleyi (1370). 

Calymperes jardinii Besch. = C. erosum (57). 

Calymperes jollyi Broth. & Par. = C. rabenhorstii (57) 


Calymperes konkourae Par. = C. erosum (57). 

Calymperes laevifolium Mitt. = C. palisotii (1370). 

Calymperes lamellosulum Card. in Ren & Card. = C. 

loucoubense (765). 

Calymperes lecomptei Besch. (“lecomtei”) = C. afzelii 

(57). 

Calymperes leucocoleus C.Müll. ex Besch. = C. erosum 

(57). 

Calymperes leucomitrium C.Müll. = C. afzelii (1473). 

As existing collections under this name appear to 

be of either C. afzelii or C. erosum (1473), 

collections listed in Schultze-Motel (47) and 

Bizot et al. (1880) have been ignored. 

Calymperes levyanum Besch. CBS(1531). 

Calymperes ligulare Mitt. = C. tenerum (1473). 

Calymperes linealifolium C.Müll. ex Besch. = C. 

afzelii (1473). As there is some doubt about 

collections with this name, the collection from 

Cameroon listed in Schultze-Motel (47) has been 

ignored. 

Calymperes lonchophyllum ssp. microblastum 

(C.Müll.) S.Edwards CAM(47), GAB(47), 

GHA(47), IVO(1880), NIG(47), ZAI(65). 

Calymperes lonchophyllum ssp. saxatile (C.Müll.) 

S.Edwards LIB(47), SIL(47), TOG(47). 

Calymperes loucoubense Besch. MAD(40). 

Calymperes loucoubense var. minus Ren. & Par. = C. 

decolorans (1473) (= C. pallidum). 

Calymperes mahense P.Varde = C. decolorans (1473) 


Calymperes malimbae C.Müll. = C. erosum (1473). 

Calymperes mariei Besch. = C. palisotii (1370). 

Calymperes maroantsetrae Card. = C. afzelii (1473). 

Calymperes maroantsetrae var. angustifolium Card. 

= C. erosum (1473). 

Calymperes mathieui Rem. & Card. = C. erosum 

(1473). 

Calymperes mayottense Besch. = C. afzelii (767). 

Calymperes megamitrium C.Müll. ex Besch. = C. 

erosum (57). 

Calymperes microblastum C.Müll. ex Besch. = C. 

lonchophyllum ssp. microblastum (57). 

Calymperes minus Besch. = C. pallidum (1518). 

Calymperes motleyi Mitt. ex Dozy & Molk. 

SEY(1373). 

Calymperes norkettii L.T.Ellis SEY(1374). 

Calymperes nossi-beanum (Besch.) Broth. = C. 

taitense (1473). 

Calymperes nossi-combae Besch. = C. afzelii (1518). 

Calymperes occidentale Besch. = C. palisotii (1473). 

Calymperes orthophyllaceum C.Müll. = C. afzelii 

(57). 

Calymperes pachyloma Hampe = C. taitense (1518). 

Calymperes palisotii ssp. palisotii Schwaegr. 

CAM(57), GAB(57), GHA(57), GUI(57), 

IVO(44,1880), LIB(57), MAD(40), MAU(105), 

NIG(57), ROD(105), SEN(57), SEY(40), SIL(57), 

TAN(1370), TOG(1878), ZAI(57). 

Calymperes pallidum Mitt. ANG(57), BEN(57), 

BUR(65), COM(40), GHA(57), GUI(57), 

IVO(57,1880), MAD(40), NAT(1531), 

NIG(57,334), ROD(105), SEY(40), SIL(57), 

TAN(46), TOG(57), TRV(1531), UGA(46), 

ZAI(65,57), ZIM(61). 

Calymperes palmicola Besch = C. pallidum (1518). 

Calymperes patentifolium Par. = C. erosum (57). 

Calymperes perlimbatum Par. = C. afzelii (1473). 

Calymperes perrottetii Besch. = C. afzelii (1473). 

Calymperes perserratum Broth. & Par. = C. loncho- 

phyllum ssp. saxatile (57). 

Calymperes pintasii C.Müll. ex Besch. Excluded. 

There is doubt about the identity of this taxon, 

109

110 

recorded from GUI, LIB and SAO (47). See 

Bescherelle (1625). 

Calymperes polii Besch. = C. loucoubense (765). 

Calymperes principis Broth. = C. tenerum (1473). 

Calymperes proligerum C.Müll. ex Dus. LIB(47). 

Calymperes pygmaeum Par. = Syrrhopodon 

parasiticus var. parasiticus (1473). 

Calymperes quintasii Broth. = C. afzelii (1473). 

Calymperes rabenhorstii Hampe & C.Müll. = C. 

pallidum (1372). 

Calymperes rabenhorstii var. longemitrata Broth. & 

Thér. = C. decolorans (1473) (= C. pallidum). 

Calymperes reduncum Besch. = C. hispidum (40). 

Calymperes renauldii Card. = C. afzelii (1518). 

Calymperes rhypariophyllum C.Müll. = C. afzelii 

(57). 

Calymperes robillardii Besch. = C. graeffeanum 

(1518). 

Calymperes rotundatulum C.Müll. ex Dus., nom. 

nud. Excluded. Following Schultze-Motel (47), 

this taxon has been treated as an orthographic 

variant of C. rotundatum (= C. afzelii). 

Calymperes rotundatum C.Müll. ex Besch. = C. afzelii 

(1473). 

Calymperes rupestre C.Müll. ex Dus. = C. lonchophyl- 

lum ssp. microblastum (57). 

Calymperes sanctae-mariae Besch. = C. tenerum 

(1473). 

Calymperes sanctae-mariae var. mayottense 

Besch. COM(40). 

Calymperes sankaranae Par. = C. afzelii (57). 

Calymperes saxatile C.Müll. ex Besch. = C. loncho- 

phyllum ssp. saxatile (57). 

Calymperes secundulum C.Müll. ex Besch. = C. afzelii 

(57). 

Calymperes seychellarum Besch. = C. erosum (1373). 

Calymperes subasterigerum Par. = C. afzelii (57). 

Calymperes subcrassilimbatum P.Varde = C. erosum 

(1373). 

Calymperes subdecolorans Card. = C. rabenhorstii 

(57) (= C. pallidum). 

Calymperes subdecolorans var. remotifolium Thér. 

= C. decolorans (1473) (= C. pallidum). 

Calymperes subdiaphanum Leroy = C. tenerum 

(1473). 

Calymperes subtenellum Broth. in Mildbr., nom. 

nud. ANN(47). 

Calymperes taitense (Sull.) Mitt. COM(40), 

MAD(40), REU(105), SEY(105). 

Calymperes tenellum C.Müll. ex Besch. = C. palisotii 

var. palisotii (57). 

Calymperes tenerum var. edamense Fleisch. = C. 

tenerum (1485). 

Calymperes tenerum var. tenerum C.Müll. 

AMI(40), CHG(40), COM(40), GHA(57), 

IVO(44,1880), KEN(46), MAD(40), NAT(1531), 

NIG(57), TAN(46,57), ZAI(65). 

Calymperes theriotii Ren. & Card. = C. palisotii ssp. 

palisotii (57). 

Calymperes thomeanum C.Müll. Excluded. There 

is doubt about the identity of this taxon, recorded 

from SAO (47). See Bescherelle (1625). 

Calymperes thompsonii Besch = C. pallidum (1518). 

Calymperes usambaricum Broth. = C. afzelii (1473). 

Calymperes victoriae Dix. = C. afzelii (1473). 

Calymperes zanzibarense Besch. = C. tenerum (1473). 

Calymperopsis (C.Müll.) Fleisch. The view of Reese 

& Orbán (132) was that this genus should be 

treated as a subgenus of Syrrhopodon, and Reese 

has now revised several of the taxa (1708, 1711), 

provided a synopsis of the subgenus (1909) and 

annotated all BM specimens with a name in 

Syrrhopodon. The annotations suggest that 

previous understanding of the subgenus has been 

highly suspect. All records have now been 

transferred to Syrrhopodon, but see reservations 

expressed under that genus regarding distribution 

data. 

Calymperopsis crenulata Tixier = Syrrhopodon 

crenulatus (1909). 

Calymperopsis disciformis (C.Müll.) Tixier = Syrrho- 

podon africanus ssp. africanus (1909). 

Calymperopsis elimbatus Tixier = Syrrhopodon 

tixieri (1909).. 

Calymperopsis madagascariensis (Thér.) Broth. = 

Syrrhopodon parasiticus (1909). 

Calymperopsis mandrakensis Tixier = Syrrhopodon 

africanus ssp. mandrakensis (1909). 

Calymperopsis martinicensis (Broth.) Broth. = Syrr- 

hopodon disciformis (1909). 

Calymperopsis subdisciformis (Besch.) Fleisch. = 

Syrrhopodon subdisciformis (1909). 

Calymperopsis viguieri (Thér.) Broth. = Syrrhopodon 

parasiticus (1909). 

Calyptothecium acutifolium var. acutifolium 

(Brid.) Broth. ANG(46,2), CBS(1531,2), 

CAR(47), MAL(67,2), NAT(1531), REU(2,105), 

RWA(65), TAN(46), TRV(1531), UGA(46,2), 

ZAI(65,2), ZAM(46,62,2), ZIM(46). 

Calyptothecium acutifolium var. antitrichioides

(Besch.) Par. REU(2,105). 

Calyptothecium acutifolium var. breviusculum 

(C.Müll. ex Dus.) Argent CAM(47,2), 

TOG(1878). 

Calyptothecium acutifolium var. longiusculum 

(Dus.) Argent CAM(47,2), TAN(46). 

Calyptothecium bescherellei (Ren.) Broth. 

MAD(40). 

Calyptothecium hoehnelii (C.Müll.) Argent 

CBS(1531), KEN(46,2), MAL(67,2), 

NAT(1531,2), RWA(65,2), TAN(46,2), 

TRV(1531), UGA(46,2), ZAI(65,2), ZIM(61). 

Calyptothecium hoehnelii var. angustifolium 

(Dix.) Kis KEN(46), TAN(46), UGA(46). 

Calyptothecium hoehnelii var. angustifolium (Dix.) 

Kis. This variety is not mentioned by Argent (2), 

but he does discuss the great variability of the 

species in terms of leaf size, and this variety may 

prove to be within the scope of the main species. 

Calyptothecium planifrons (Ren. & Par.) Argent 

CAR(47,2), CHA(47,2), MAD(40,2), TAN(46), 

ZAI(2,411), ZAM(46,62,2). 

Calyptothecium pterobryoides Argent 

UGA(46,2). 

Calyptothecium subbreviusculum Thér. & P.Varde = 

C. acutifolium var. acutifolium (2). 

Calyptrochaeta asplenioides (Brid.) Crosby 

CBS(1531), COM(40), MAD(40), REU(105), 

TAN(46). 

Calyptrochaeta cristata (Hedw.) O’Shea 

MAU(40). 

Camptochaete fallax (Ren. & Card.) Ren. 

MAD(40). 

Camptochaete ramulosa (Mitt.) Jaeg. ZAI(65). 

Campyliadelphus polygamus (BS&G) Kanda 

TAN(1405). 

Campylium hispidulum var. sommerfeltii (Myr.) 

J.Lange Excluded. According to Crundwell et al. 

(489), both African collections given this name 

- CAR(47) and KEN(46) - refer to other taxa. 

Campylium stellatum var. stellatum (Hedw.) 

J.Lange & C.Jens KEN(66). 

Campylodontium dusenii (Broth.) Broth. = C. flaves- 

cens (1878) (= Mesonodon flavescens). 

Campylodontium flavescens (Hook.) Besch. & Lac. 

= Mesonodon flavescens (715). 

Campylodontium flavescens var. viride Card. 

MAD(40). 

Campylodontium flavescens var. viride Card. This 

variety presumably belongs with the owning spe- 

cies, in Mesonodon. Revision is required. 

Campylopodium euphorocladum var. laevigatum 

(Thér.) Thér. = Microcampylopus laevigatus 

(507). 

Campylopodium laevigatum Thér. = 

Microcampylopus laevigatus (507). 

Campylopodium proscriptum (Hornsch.) Broth. = 

Dicranella proscripta (506). 

Campylopus abyssinicus De Not. Excluded. No 

herbarium specimen available (54). 

Campylopus afro-concolor C.Müll. ex Broth. = C. 

savannarum (54). 

Campylopus alpicola (C.Müll.) Jaeg. = C. fragilis 

(54). 

Campylopus ambiguus Thér. Excluded. No herbarium 

material available (54). 

Campylopus amblyacron (Ren.) Ren. & Card. = C. 

cambouei (1751). 

Campylopus ampliretis (C.Müll.) Card. = C. flaccidus 

(54). 

Campylopus aongstroemii C.Müll. = C. aureonotens 

(54). 

Campylopus arbogastii Ren. & Card. = C. julaceus 

ssp. arbogastii (54). 

Campylopus arbogastii var. capitulifer Ren. & Card. 

= C. julaceus ssp. arbogastii (54). 

Campylopus arctocarpus ssp. arctocarpus (Hornsch.) 

Mitt. All records of this species for Africa belong 

to ssp. madecassus. Ssp. arctocarpus is neotro- 

pical. (54). 

Campylopus arctocarpus ssp. madecassus 

(Besch.) Frahm COM(54), MAD(54,40), 

MAL(67), MAU(54), REU(54), SEY(54), 

TAN(54). 

Campylopus arcuatus (Brid.) Jaeg. ASC(54,1296), 

MAD(54,40), MAU(54), REU(54), 

STH(54,1296). 

Campylopus argutidens Broth. & Par. = C. 


Campylopus argutidens var. angustifolius Thér. & 

P.Varde = C. savannarum (54). 

Campylopus aspericuspis Thér. & P.Varde = C. 


Campylopus assimilis Thér. & P.Varde = C. savanna- 

rum (54). 

Campylopus atro-sordidus C.Müll. ex Broth. = C. 

pilifer (54). 

Campylopus atroluteus (C.Müll.) Par. = C. bartrami- 

naceus & C. bicolor ssp. atroluteus (see discussion 

in Frahm, (54) about the distribution of these two 

111

112 

plants). 

Campylopus aureonitens (C.Müll.) Jaeg. ANG(54), 

ASC(54), CAM(40), CBS(1531,54), 

COM(54,40), KEN(54), MAD(54,40), MAL(67), 

MAU(54), NAT(1531,54), OFS(1531,54), 

REU(54), SWA(1531), TAN(54), TRV(1531,54), 

ZIM(54). 

Campylopus aureus Bosch. & Lac. = C. schmidii 

(1751). 

Campylopus bartramiaceus (C.Müll.) Thér. 

CBS(1531,54), LES(1531), NAM(1531), 

NAT(1531,54), OFS(1531), REU(105), 

SWA(1531), TRV(1531,54). 

Campylopus bequaertii Thér. & Nav. = C. savannarum 

(54). 

Campylopus bequaertii var. gabonensis Thér. & 

P.Varde = C. savannarum (1751). 

Campylopus bicolor (C.Müll.) Wils. is shown in 

Crosby et al. (40) for Madagascar, on the basis of 

C. pseudobicolor (shown as a synonym). This 

is not a synonym of C. bicolor, which only occurs 

in Africa as ssp. atroluteus (54). 

Campylopus bicolor ssp. atroluteus (C.Müll.) 

Frahm CBS(1531,54). 

Campylopus boivinianus Beschl. = C. flexuosus (54). 

Campylopus brevirameus Dix. = C. pilifer (54). 

Campylopus cailleae Ren. & Card. = C. perpusillus 

(54). 

Campylopus calvus Ren. & Card. = C. aureus (54). 

Campylopus cambouei Ren. & Card. CBS(54), 

GUI(54,47), KEN(411), MAD(54,40), MAL(67), 

MAU(54), MOZ(54), TAN(54), TRV(54). 

Campylopus capillaceus (Brid.) Jaeg., hom. illeg. 

Excluded. No herbarium material available (54). 

Campylopus capitiflorus Mont. = C. arcuatus (54). 

Campylopus capitiflorus var. pachycomus Besch. = 

C. arcuatus (54). 

Campylopus cardotii Thér. = C. controversus (1701). 

Campylopus carolinae Grout RWA(65). 

Campylopus catarractilis (C.Müll.) Par. 

CBS(1531,54), MAD(54), TRV(1531,54). 

Campylopus chevalieri Broth. & Thér. 

CAM(47,54), CON(54,47), GAB(54,47), 

IVO(54,47), NIG(54,47), SIL(54,47). 

Campylopus chlorotrichus (C.Müll.) Par. = C. steno- 

pelma (54). 

Campylopus chrismarii (C.Müll.) Mitt. = C. nivalis 

(54). 

Campylopus chryseolus Besch. = C. aureus (54). 

Campylopus clavatus (R.Br.) Wils. Excluded. Not 

found in Africa - see Frahm (54). 

Campylopus comatus Ren. & Card. = C. trachyble- 

pharon var. comatus (54). 

Campylopus comatus var. subcomatus (Ren. & Card.) 

Thér. = C. trachyblepharon var. comatus (54). 

Campylopus compactus Par. & Broth., hom. illeg. = 

C. savannarum (54). 

Campylopus consociatus Thér. = C. flaccidus (54). 

Campylopus controversus (Hampe) Jaeg. 

CBS(1531,54), LES(1531), MAD(54,40), 

MAL(67), MAU(56,40), MOZ(54), 

NAT(1531,54), REU(54), RWA(65), STH(54), 

SWA(1531), TRV(1531,54), ZAI(65). 

Campylopus crateris Besch. COM(54,40), 

KEN(54), MAD(54,40), REU(54,40). 

Campylopus crispatulus Thér. Excluded. No 

herbarium material available (54). 

Campylopus cummingii Duby = C. introflexus (54). 

Campylopus decaryi Thér. CBS(54), MAD(54,40), 

TRV(54), ZAI(65). 

Campylopus deciduus Ren. & Card. = C. aureus (54). 

Campylopus denticuspis Broth. in Mildbr. = C. nivalis 

(54). 

Campylopus denticuspis var. acutifolius Broth. = C. 

nivalis (1751). 

Campylopus dicranoides Thér. KEN(54), 

RWA(65), UGA(54), ZAI(54,65). 

Campylopus dissitus C.Müll. ex Broth. = C. savanna- 

rum (54). 

Campylopus dissitus var. parvifolius Thér. & P.Varde 

= C. savannarum (1751). 

Campylopus divaricatus (Mitt.) Jaeg. = C. savannarum 

(54). 

Campylopus dolosus Besch. = Bryohumbertia sp 

(54). 

Campylopus drouhardii Card. & Copp. = C. aureo- 

notens (54). 

Campylopus dusenii C.Müll. in Par., nom. nud. = C. 


Campylopus echernieri Besch. = C. jamesonii (54). 

Campylopus eckendorfii Thér. & P.Varde = C. pilifer 

(54). 

Campylopus erythrocaulon Broth. = C. savannarum 

(54). 

Campylopus eximius Reichdt. ASC(54). 

Campylopus filescens Ren. & Card. = Bryohumbertia 

sp (54). 

Campylopus filifolius (Hornsch.) Mitt. = Bryohum- 

bertia filifolia (481) 

Campylopus flaccidus Ren. & Card. BUR(54,65),

CBS(1531), KEN(411), MAD(54,40), MAL(67), 

MOZ(357), NAT(1531,54), OFS(54), RWA(65), 

SUD(54), TAN(54), TRV(1531,54), ZAI(65), 

ZIM(54). 

Campylopus flagei Ren. & Card. = C. filifolius (19) 

(=Bryohumbertia filifolia). 

Campylopus flavicoma C.Müll. ex Broth. = Bryo- 

humbertia flavicoma (54). 

Campylopus flexuosus (Hedw.) Brid. MAD(54,40), 

MAU(54), REU(54), RWA(65), SAO(54,47), 

STH(1296), TAN(54,146), UGA(54), ZAI(54,65), 

ZAM(62). 

Campylopus flexuosus var. incacorralis (Herz.) 

Frahm MAU(54), REU(298), TAN(54), 

UGA(54), ZAI(54,65). 

Campylopus fragilis (Brid.) BS&G CAM(54), 

ETH(54), KEN(411), MAL(67), REU(105), 

RWA(54), TAN(54), TRV(1531,54), UGA(54), 

ZAI(54,65). 

Campylopus fusco-lutescens Ren. & Card. = C. 

controversus (1701). 

Campylopus gallienii Par. = C. robillardei (54). 

Campylopus gallienii var. brevifolius Thér. = C. 

robillardei (1751). 

Campylopus guineensis P.Varde & Thér. = C. obrutus 

(54). 

Campylopus helenicus C.Müll. = C. arcuatus (54). 

Campylopus hensii Ren. & Card. ANG(54), 

CAM(54,47), CAR(54,47), CON(47), EGU(54), 

GAB(54,47), IVO(54), KEN(54), LIB(54), 

MAL(67), NAT(54), NIG(54,334,47), RWA(54), 

TAN(54), UGA(54), ZAI(54,65), ZAM(54). 

Campylopus heribaudii C.Müll. = C. robillardei (54). 

Campylopus hildebrandtii (C.Müll.) Jaeg. 

BUR(54,65), CAM(54), CBS(54), COM(54,40), 

KEN(54), MAD(40), MAL(67), NAT(54), 

REU(54,298), TAN(54), UGA(54), ZAI(65,54). 

Campylopus hispidus Ren. & Card. = C. pilifer (1751). 

Campylopus horridus Welw. & Duby = C. savannarum 

(54). 

Campylopus incacorralis Herz. = C. flexuosus var. 

incacorralis (1751). 

Campylopus inchangae (C.Müll.) Par. = C. robillardei 

(54). 

Campylopus inerangae (C.Müll.) Par. Orthographic 

variant for C. inchangae, but the specimen from 

Madagascar (40) given this name was C. arctocar- 

pus ssp. madecassus, not C. robillardei (54). 

Campylopus interrupulus Besch. = C. filifolius (1641) 

(= Bryohumbertia filifolia). 

Campylopus introflexus (Hedw.) Brid. BOT(1531), 

CBS(1531,54), LES(1531), MAU(105), 

NAM(1531), NAT(1531), OFS(1531), REU(54), 

STH(54,1296), TRV(1531,54). 

Campylopus introflexus (Hedw.) Brid. Some published 

collections with this name are misidentified (e.g. 

Sierra Leone and Cameroon in (47), which are C. 

pilifer) (54). 

Campylopus introflexus var. altecristatus (Ren. & 

Card.) Thér. = C. pilifer (54). 

Campylopus introflexus var. mauritianus (Ren. & 

Card.) Thér. = C. pilifer (54). 

Campylopus introflexus var. polytrichoides (De Not.) 

Giac. = C. pilifer (54). 

Campylopus jamesonii (Hook.) Jaeg. CAM(54,47), 

KEN(54), MAD(54,40), MAL(67), MOZ(54), 

REU(54,40), RWA(54,65), TAN(54), 

TRV(1531,54), UGA(54), ZAI(54,65), ZIM(54). 

Campylopus johannis-meyeri (C.Müll.) Par. 

CAM(54), KEN(54), TAN(54), ZAI(54,65). 

Campylopus julaceus ssp. arbogastii (Ren. & 

Card.) Frahm MAD(54,40), MAU(54), 

OFS(1531,54), REU(54), SEY(54). 

Campylopus kivuensis P.Varde & Thér. 

ZAI(54,65). 

Campylopus labeensis Broth. & Par. = C. cambouei 

(54). 

Campylopus laxobasis Ren. & Card. = C. cambouei 

(54). 

Campylopus lecomtei Besch. in P.Varde, nom. nud. 

Excluded. No herbarium material available (54). 

Campylopus leptodrepanium (C.Müll.) Par. = C. 

nivalis (54). 

Campylopus leptotrichaceus (C.Müll.) Par. = C. py- 

riformis (804). Some plants recorded under this 

name are C. praetermissus (e.g. Gabon (47)). 

Campylopus letestui Thér. & P.Varde = C. nanophyllus 

(54). 

Campylopus leucochlorus (C.Müll.) Par. = C. 

hildebrandtii (804). 

Campylopus lonchocladus C.Müll. = C. aureonotens 

(54). 

Campylopus longifolius Besch. = C. arcuatus (54). 

Campylopus maclaudii Par. & Broth. = C. nanophyllus 

(54). 

Campylopus macrotis C.Müll. ex Broth. = C. savan- 

narum (54). 

Campylopus madecassus Besch. = C. arctocarpus 

ssp. madecassus (54). 

Campylopus madegassus Dix. = C. arctocarpus ssp. 

113

114 

madecassus (54). 

Campylopus mangenotii Biz. & Kilb. = C. smaragdinus 

(1751). 

Campylopus marginii Thér. = C. nivalis (54). 

Campylopus matarensis Besch. = C. arctocarpus ssp. 

madecassus (54). 

Campylopus mathieui Thér. = C. savannarum (54). 

Campylopus montanus Welw. & Duby = C. savanna- 

rum (54). 

Campylopus mozambicus P.Varde = C. cambouei 

(54). 

Campylopus nanophyllus C.Müll. ex Broth. 

CAM(54,47), CAR(54,47), GAB(54,47), 

GUI(54,47), IVO(1458), MAD(40), MAL(67), 

REU(54), RWA(65), TRV(54), ZIM(54). 

Campylopus naumanni C.Müll. = C. arcuatus (54). 

Campylopus nivalis (Brid.) Brid. CAM(54,47), 

KEN(54), MAD(54), MAL(67), MAU(105), 

NAT(54), REU(54), RWA(54,65), TAN(54), 

UGA(54), ZAI(54). 

Campylopus obrutus Thér. & P.Varde BUR(65), 

CAM(54,47), CON(47,54), GAB(54,47), 

GUI(54,1458,47), NIG(54,1492), RWA(65), 

ZAI(54,65). 

Campylopus ochrodictyon Aongstr. = C. introflexus 

(54). 

Campylopus orzeszkoanus Ren. & Card. = C. crateri 

(54). 

Campylopus pallescens Besch. = C. hildebrandtii 

(54). 

Campylopus pallidisetus Thér. = Bryohumbertia sp 

(54). 

Campylopus pallido-nitens Thér. = C. robillardei 

(54). 

Campylopus paludicola Broth. = C. pyriformis (1562). 

Campylopus parisii Ren. = C. smaragdinus (54). 

Campylopus perichaetialis P.Varde & Thér. 

IVO(1880), KEN(54), RWA(54,65), ZAI(54,65). 

Campylopus perpusillus Mitt. BUR(65), 

CAM(54,47), CAR(54,47), CON(47), 

GAB(54,47), GUI(54,47), KEN(54), 

MAD(54,40), MOZ(54), NAT(1531), 

NIG(54,47,334), RWA(54,65), SIL(54,47), 

TAN(54,46), TRV(1531), UGA(54,46), 

ZAI(54,65), ZAM(54,46), ZIM(54,46). 

Campylopus perrieri Thér. = C. flaccidus (54). 

Campylopus pilifer Brid. ANG(54), ASC(54,1296), 

BUR(54,65), CAM(54,47), CBS(1531,54), 

CAP(56,1296), CAR(54,47), CHA(54), 

COM(54,40), ETH(54), KEN(54), LES(54,1531), 

LIB(54), MAD(54,40), MAL(67), MAU(54), 

NAM(1531), NAT(1531,54), OFS(1531,54), 

REU(54), RWA(54,65), SEY(40), SIL(54,47), 

SOC(54,1626,40), SWA(1531), TAN(54), 

TRV(1531,54), UGA(54), ZAI(54,65), ZIM(54). 

Campylopus pobeguinii Par. & Broth. = C. 


Campylopus pobeguinii var. minor Par. = C. savan- 

narum (1751). 

Campylopus polytrichoides De Not. = C. pilifer (54). 

Campylopus praetermissus Frahm GAB(54,47), 

KEN(54), MAU(54), REU(298), RWA(65), 

SAF(54), TAN(54), ZAI(65). 

Campylopus procerus (C.Müll.) Kindb. = C. jamesonii 

(54). 

Campylopus pseudo-nigrescens Thér. = C. flaccidus 

(54). 

Campylopus pseudobicolor C.Müll. ex Ren. & 

Card. CBS(54), MAD(54,40), MAU(105). 

Campylopus pseudovirescens Ren. & Card. Excluded. 

Probably C. arcuatus (1751). 

Campylopus purpurascens Lor. = C. pilifer (54). 

Campylopus pyriformis var. pyriformis (Schultz) 

Brid. CBS(1531,54), CAP(56), KEN(54,411), 

MAL(67), NAT(1531,54), OFS(1531), 

REU(298), RWA(54,65), TAN(54), 

TRV(1531,54), ZAI(65,54). 

Campylopus quintasii Broth. = C. flexuosus (54) 

Campylopus reticulatus Par. & Broth. Excluded. 

Type specimen is not Campylopus (54). 

Campylopus ripicola Besch. = C. aureus (54). 

Campylopus robillardei Besch. CBS(1531,54), 

MAD(54,40), MAL(67), MAU(54), MOZ(357), 

NAT(1531,54), REU(54), RWA(65), 

SWA(1531), TAN(54), TRV(1531,54), UGA(54), 

ZAI(65), ZIM(54). 

Campylopus salessianus Broth. = C. savannarum 

(54). 

Campylopus savannarum (C.Müll.) Mitt. ANG(54), 

BOT(1531), BUR(54,65), CAM(54,47), 

CBS(1531,54), CAR(54,47), CON(47,54), 

ETH(50), GAB(54,47), GHA(54), GUI(54,47), 

IVO(54,1458), KEN(54), LIB(54,47), MAL(67), 

NAT(1531), NIG(54,1492,47), PRI(54,47), 

RWA(65), SAO(54,47), SIL(54,47), SWA(1531), 

TAN(54), TOG(1878), TRV(1531,54), UGA(54), 

ZAI(54,65), ZAM(54), ZIM(54). 

Campylopus scabricuspis Mitt. = C. flexuosus (1751). 

Campylopus schmidii (C.Müll.) Jaeg. COM(54,40), 

MAD(54,40), REU(54,40), ZAI(65).

Campylopus serricuspis Thér. & P.Varde = C. 


Campylopus simii Schelpe = C. julaceus ssp. arbogastii 

(54). 

Campylopus smaragdinus (Brid.) Jaeg. 

ASC(54,1296), KEN(54), MAD(54,40), REU(54), 

TAN(54), ZAI(54,65). 

Campylopus stenopelma (C.Müll.) Par. = C. 

controversus (1701). 

Campylopus stramineus (Mitt.) Jaeg. = C. nivalis 

(54) 

Campylopus subambiguus Thér. = C. nanophyllus 

(54). 

Campylopus subchlorophyllosus C.Müll. ex Rabenh. 

= Sphaerothecium subchlorophyllosus (1475). 

Campylopus suberythrocaulon Broth. in Mildbr. = C. 

hensii (54) Some specimens given this name 

belong to C. savannarum (e.g Congo (47)) (54). 

Campylopus suberythrocaulon var. parvifolius P. 

Varde, nom. inval. = C. hensii (1751) 

Campylopus subflexuosus Par., nom. nud. = C. fragilis 

(54). 

Campylopus subleptodrepanium Broth. = C. nivalis 

(1751) 

Campylopus subobrutus Thér. & P.Varde = C. obrutus 

(54) 

Campylopus subvirescens Ren. & Card. = C. arctocar- 

pus ssp. madecassus (54). 

Campylopus torrentis Thér. & P.Varde 

GAB(54,47), NIG(54,1492). 

Campylopus trachyblepharon ssp. comatus (Ren. 

& Card.) Frahm MAD(54,40), MAU(54), 

REU(298). 

Campylopus tsaratananae Thér. = C. flexuosus (54). 

Campylopus valentinii Besch. = C. aureonotens (54). 

Campylopus verdolinii Par. & Ren. = C. flexuosus 

(54). 

Campylopus virescens Besch. = C. flexuosus (54). 

Campylopus viridatulus C.Müll. ex Broth. = C. 


Cardotiella appendiculata (Ren. & Card.) Vitt 

MAD(40), REU(105). 

Cardotiella renauldii (Broth.) Vitt. MAD(40). 

Cardotiella secunda (C.Müll.) Vitt CBS(1531), 

NAT(1531). 

Cardotiella subappendiculata (Broth.) Vitt 

MAD(40), MAS(40). 

Catagonium nitens (Brid.) Card. COM(40), 

MAD(40), MAU(105), REU(105), 

TAN(46,1881). 

Catagonium nitens ssp. maritimum (Hook.) Lin 

CBS(1531,1881), NAT(1531,1881), 

SWA(1531,1881), TRV(1531,1881). 

Ceratodon purpureus ssp. purpureus (Hedw.) 

Brid. CAM(47), CBS(1882), KEN(46), MAL(67), 

MAU(105), RWA(65), STH(1296), TAN(46), 

UGA(46), ZAI(65). 

Ceratodon purpureus ssp. stenocarpus (C.Müll.) 

Dix. CBS(1531), LES(1531), MAD(40), 

NAT(1531), REU(105), RWA(65), TRV(1531), 

ZAI(65). 

Ceratodon purpureus var. brevidens Besch. 

MAU(105), REU(105). 

Ceratodon purpureus var. congolensis Thér. 

RWA(65). 

Ceratodon purpureus var. robustus P.Varde, 

hom. illeg. UGA(46). 

Ceratodon purpureus var. xanthopus Sull. 

ZAI(65). 

Ceratodon stenocarpus C.Müll. = Ceratodon 

purpureus ssp. stenocarpus (1589). 

Ceratodon viridatus (C.Müll.) Broth. TAN(46). 

Chaetomitrium borbonicum Besch. MAU(105), 

REU(105). 

Chaetomitrium comorense C.Müll. COM(40). 

Chaetomitrium dusenii var. brevinerve (P.Varde) 

P.Varde CAR(47), GAB(47), TAN(46). 

Chaetomitrium dusenii var. dusenii C.Müll. ex 

Broth. CAM(47), GAB(47), IVO(44), ZAI(65). 

Chamaebryum pottioides Thér. & Dix. CBS(1531), 

NAM(1531). 

Chameleion cryptocarpus (Dozy & Molk.) Ellis & 

Eddy in Eddy GAB(24), GUI(363), ZAI(65,24). 

Cheilothela chilensis (Mont.) Broth. CBS(1531), 

LES(1074). 

Cheilothela jaegeri Bizot SIL(4). 

Chenia leptophylla (C.Müll.) Zander 

BOT(1531,68), CBS(1531), MAL(68,67), 

NAT(1531), TAN(68), TRV(1531), ZIM(68). 

Chorisodontium falcatum Magill CBS(1531). 

Chryso-hypnum cavifolium (Dix.) Ochyra & Sharp 

BUR(65), ETH(146), KEN(46), MAL(67), 

NAT(1531), TAN(46), TRV(1531), ZAM(146), 

ZIM(61). 

Chryso-hypnum demaretii (P.Varde) Higuchi 

UGA(46), ZAI(65). 

Chryso-hypnum frondosum (Mitt.) Buck 

CAM(47), FER(47), KEN(46), RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Cinclidotus fontinaloides (Hedw.) P.Beauv. 

115

116 

KEN(46), TAN(259). 

Cladophascum gymnomitrioides Dix. & Sim 

CBS(1531), NAM(1531), OFS(1531), 

TAN(1437), TRV(1531), ZIM(46,61). 

Clastobryella foliicola P.Varde = Gammiella merrillii 

(46) 

Clastobryella pusilla (Thér.) Wijk & Marg. 

MAD(40). 

Clastobryophilum rufo-viride (Besch.) Fleisch. 

MAD(40), REU(105), SEY(40). 

Conostomum crassinervium P.Varde KEN(46). 

Conostomum pentastichum (Brid.) Lindb. 

LES(1074,1531), NAT(1531). 

Cratoneuron curvicaule fo. densirete P.Varde, nom. 

inv. descr. gall. = C. filicinum (1533). 

Cratoneuron filicinum var. curvicaule (Jur.) Moenk. 

= C. filicinum (1533). 

Cratoneuron filicinum var. filicinum (Hedw.) 

Spruce KEN(46), NAT(1531,1533), TAN(46), 

UGA(1533), ZAI(65). 

Cratoneuron procerum (Dix.) Broth. = C. filicinum 

(1533). 

Cratoneuron subcurvicaule P.Varde = C. filicinum 

(1533). 

Crossidium apiculatum Magill CBS(1531). 

Crossidium chloronotos (Brid.) Limpr. = C. 

squamiferum var. squamiferum (1420). 

Crossidium crassinerve var. laevipilum (Thér. & 

Trab.) Delgadillo = C. laevipilum (1420). 

Crossidium laevipilum Thér. & Trab. KEN(411). 

Crossidium spiralifolium Magill CBS(1531), 

NAM(1531). 

Cryphaea bollei Broth. & Geh., nom. nud. in 

sched. CAP(1296). 

Cryphaea exigua (C.Müll.) Jaeg. CBS(1531), 

KEN(46), NAT(1531), RWA(65), SWA(1531), 

TAN(46), TRV(1531). 

Cryphaea filivaga C.Müll. in Par., nom. nud. 

ETH(50). 

Cryphaea lasioides C.Müll. in Par., nom. nud. 

ETH(50). 

Cryphaea longicuspis P.Varde & Thér. TAN(46), 

ZAI(65). 

Cryphaea protensa Bruch & Schimp. ex C.Müll. 

ETH(50), KEN(46), TAN(46). 

Cryphaea ragazzii (Brizi) Broth. ETH(50). 

Cryphaea robusta Broth. & Thér. KEN(46), 

TAN(46). 

Cryphaea rutenbergii C.Müll. MAD(40). 

Cryphaea scariosa C.Müll. = Antitrichia curtipendula 

(1521). 

Cryphaea tenuiretis Thér. MAD(40). 

Cryphaea welwitschii Duby = Schoenobryum welwit- 

schii (1291). 

Cryptoleptodon flexuosus (Harv.) Ren. & Card. = C. 

pluvinii (301). 

Cryptoleptodon longisetus (Mont.) Enroth 

CAP(301). 

Cryptoleptodon pluvinii (Brid.) Broth. 

ETH(50,301). 

Ctenidium floribundarioides Dix. ex Tosco & 

Piovano TAN(46). 

Ctenidium mauritianum Broth. MAS(40). 

Ctenidium squarrifolium (Broth.) Broth. 

CAM(47), CAR(47), CON(47), GAB(47), 

UGA(46). 

Cyathophorella africana (Dix.) Broth. KEN(46), 

RWA(65), TAN(46), UGA(46). 

Cyclodictyon albicans (Hedw.) Kuntze REU(105). 

Cyclodictyon annobonense Broth. in Mildbr., 

nom. nud. ANN(47). 

Cyclodictyon aubertii (P.Beauv.) O.Kuntze 

MAD(40), MAL(67), MAU(105), REU(105), 

TAN(46), ZAI(65), ZIM(61). 

Cyclodictyon bidentatum Demar. CAM(47). 

Cyclodictyon borbonicum (Besch.) Broth. 

REU(105), RWA(46), TAN(46), UGA(46), 

ZAI(65). 

Cyclodictyon brevifolium Broth. BUR(65), 

REU(105), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Cyclodictyon crassicaule Broth. in Mildbr. 

RWA(65). 

Cyclodictyon delicatulum P.Varde GAB(47), 

ZIM(46). 

Cyclodictyon dixonianum Demar. KEN(46). 

Cyclodictyon filicuspis P.Varde GAB(47), PRI(47), 

ZAI(65). 

Cyclodictyon hildebrandtii (C.Müll.) O.Kuntze 

COM(40), ZAI(65). 

Cyclodictyon immersum Broth. & P.Varde 

GAB(47). 

Cyclodictyon krebedjense Broth. in Corb. 

CAM(47), CAR(47), CON(47), GAB(47), 

ZAI(65). 

Cyclodictyon krebedjense var. argutidens P.Varde 

GAB(47), ZAI(65). 

Cyclodictyon laete-virens (Hook. & Tayl.) Mitt. 

FER(47), MAD(40), SAO(47), ZAI(65), ZIM(61). 

Cyclodictyon lebrunii Demar. & P.Varde TAN(46),

ZAI(65). 

Cyclodictyon perlimbatum Broth. TAN(46). 

Cyclodictyon perrottetii Demar. & P.Varde 

REU(105). 

Cyclodictyon preussii (Broth.) Broth. CAM(47). 

Cyclodictyon recognitum Demar. & P.Varde 

ANG(46). 

Cyclodictyon spectabile Broth. in Mildbr. 

ZAI(65). 

Cyclodictyon subbrevifolium Broth. TAN(46). 

Cyclodictyon subobtusifolium Dix. ex Dem. & 

P.Varde ZAI(65). 

Cyclodictyon vallis-gratiae (C.Müll.) O.Kuntze 

ANG(46), CAM(47), CBS(1531), GAB(47), 

KEN(46), MAD(40), MAU(105), NAT(1531), 

RWA(65), TAN(46), TRV(1531), UGA(46), 

ZAI(65), ZIM(61). 

Cyclodictyon vallis-gratiae fo. breutelianum (Hampe) 

Demar. & P.Varde. Excluded. Any records for 

this form have been associated with the owning 

species. 

Cyclodictyon vesiculosum (Brid.) O.Kuntze 

MAU(105), REU(105), TAN(46). 

Cygnicollum immersum Fife & Magill CBS(1531). 

Cynodontium tanganyikae P.Varde TAN(46). 

Daltonia angustifolia var. angustifolia Doz. & 

Molk. CAM(47), MOZ(46), TAN(46). 

Daltonia angustifolia var. strictifolia (Mitt.) 

Fleisch. KEN(46), MAD(40), MAL(67), 

REU(105). 

Daltonia cardotii Biz. & Onr. COM(40), MAD(40), 

TAN(46). 

Daltonia constricta P.Varde MOZ(46). 

Daltonia dusenii C.Müll. ex Broth. CAM(47). 

Daltonia euryloma P.Varde CAM(47). 

Daltonia forsythii Card. MAD(40). 

Daltonia latimarginata Besch. COM(40), 

MAD(40), REU(105). 

Daltonia latimarginata var. madagassa Ren. 

MAD(40). 

Daltonia longinervis Mitt. FER(47). 

Daltonia mildbraedii Broth. KEN(46), RWA(65), 

TAN(46), ZAI(65). 

Daltonia mildbraedii var. laevis Demar. & Leroy 

RWA(65), TAN(46). 

Daltonia minuta Thér. TAN(46), ZAI(65). 

Daltonia mittenii Kis FER(47), MAL(67), TAN(46). 

Daltonia onraedtii Biz. REU(105). 

Daltonia patula Mitt., hom. illeg. = D. mittenii (46). 

Daltonia plicata P.Varde UGA(46). 

Daltonia splachnoides (Sm.) Hook. & Tayl. 

FER(47). 

Daltonia tortifolia Demar. & P.Varde ZAI(65). 

Desmatodon africanus P.Varde = Scopelophila cata- 

ractae (1420). 

Desmatodon bogosicus C.Müll. = Tortula bogosica 

(1420). 

Desmatodon convolutus (Brid.) Grout = Tortula atro- 

virens (1420). 

Desmatodon corticola Schimp. in Par., nom. nud. 

See Tortula sp. 

Desmatodon longipedunculatus (C.Müll.) Magill = 

Hennediella longipedunculata (1420). 

Dicranella acroclada Card. MAD(40). 

Dicranella ascensionica Mitt. ASC(1296). 

Dicranella borbonica Besch. MAS(40). 

Dicranella borbonica var. comorensis Thér. 

COM(40). 

Dicranella borbonica var. madagascariensis 

Thér. MAD(40). 

Dicranella borgeniana (Hampe) Jaeg. ex Par. 

MAD(40), TAN(46). 

Dicranella cameruniae C.Müll. ex Dus. CAM(47), 

KEN(46). 

Dicranella cardotii (R.Br. ter.) Dix. SAF(1500). 

Dicranella condensata Aongstr. STH(1296). 

Dicranella cratericola Besch. REU(105). 

Dicranella cygnea Aongstr. = Campylopodium pro- 

scriptum (1521). 

Dicranella densifolia (Thér. & P.Varde) Thér. & 

P.Varde CON(47), GAB(47), IVO(44). 

Dicranella falcularia C.Müll. ex Dus. CAM(47), 

GAB(47), GHA(47), GUI(47), IVO(1880), PRI(47). 

Dicranella flavipes Besch. REU(105). 

Dicranella grandispora Dix. KEN(46). 

Dicranella heteromalla (Hedw.) Schimp. KEN(46). 

Dicranella kenyae Dix. KEN(46), TAN(259). 

Dicranella madagassa Ren. COM(40), MAD(40). 

Dicranella nodicoma C.Müll ex Dus. CAM(47), 

KEN(46). 

Dicranella pertenella C.Müll. ex Dus. CAM(47), 

IVO(1880), ZAI(65). 

Dicranella pervaginata Broth. TAN(46). 

Dicranella pervilleana Besch. COM(40), 

MAD(40). 

Dicranella polii Ren. & Card. MAD(40). 

Dicranella proscripta (Hornsch.) Mitt. in Mell. 

ASC(506), STH(1296). 

Dicranella rigida Dix. ex Sim CBS(1531). 

Dicranella rivalis C.Müll. ex Dus. CAM(47), 

117

118 

KEN(46), ZAI(65). 

Dicranella subfalcularia Thér. CAM(47), 

GAB(47), TOG(1878). 

Dicranella subsubulata (Hampe ex C.Müll.) Jaeg. 

BUR(65), CBS(1531), CAR(47), GAB(47), 

LES(1635), MAL(46,67), NAT(1531), NIG(47), 

RWA(65), TAN(46), TRV(1531), UGA(46), 

ZAI(65), ZIM(46). 

Dicranella symonsii Dix. CBS(1531), 

LES(1531,1074), NAT(1531), OFS(1531). 

Dicranella usambarica P.Varde KEN(46), 

TAN(46). 

Dicranodontium humile P.Varde RWA(65,46), 

UGA(46). 

Dicranodontium interruptum P.Varde MAD(40). 

Dicranodontium tanganyikae J.Tayl. ex P.Varde 

TAN(46). 

Dicranoloma billardieri (Brid.) Par. CBS(1531), 

KEN(46), MAD(40), MAL(67), MAU(105), 

REU(105), RWA(65), TAN(46), TRV(1531), 

UGA(46), ZAI(65). 

Dicranoloma billardieri var. drepanocladium 

(C.Müll.) P.Varde TAN(46), ZAI(65). 

Dicranoloma billardieri var. patentifolium (Ren. 

& Par.) Thér. MAD(40), TAN(46). 

Dicranoloma billardieri var. scopareolum 

(C.Müll.) Thér. COM(40), MAD(40), REU(105), 

TAN(46). 

Dicranoloma borbonicum Ren. & Card. REU(105). 

Dicranoloma cataractarum Demar. & P.Varde 

ZAI(65). 

Dicranoloma entabeniense Magill TRV(1531). 

Dicranoloma onraedtii Biz. REU(105). 

Dicranoweisia africana Dix. KEN(46), TAN(46). 

Dicranum acanthoneurum C.Müll. KEN(46), 

RWA(65), TAN(46), ZAI(65). 

Dicranum jashii var. latifolium Thér. & Nav. 

ZAI(65). 

Dicranum johnstonii Mitt. BUR(65), GAB(47), 

KEN(46), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Dicranum kilimandscharicum Broth. TAN(46). 

Dicranum obliquatum Mitt. FER(47). 

Dicranum petrophyllum Negri UGA(46). 

Dicranum stuhlmanii Broth. = D. johnstonii (46) 

Didymodon abyssinicus Schimp. ex Par., nom. 

nud. ETH(50). 

Didymodon acutus (Brid.) Saito = D. rigidulus var 

gracilis (1420). 

Didymodon afrorubellus Broth. & Wag. in Dix. = 

Bryoerythrophyllum afrorubellum (1420) (= B. 

campylocarpum). 

Didymodon asperifolius (Mitt.) Crum et al. 

KEN(46). 

Didymodon australasiae (Hook. & Grev.) Zander 

CBS(1531), ETH(1865), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), TRV(1531). 

Didymodon capitatus Dix. ETH(50). 

Didymodon ceratodonteus (C.Müll.) Dix. 

BOT(1531), CBS(1531), LES(1531), MOZ(46), 

NAM(1531), NAT(1531), OFS(1531), 

TRV(1531), ZIM(46). 

Didymodon distans (Hoppe) Jaeg. = Trichostomum 

distans (1420). 

Didymodon eckendorfii P.Varde CAR(47), 

TOG(1878). 

Didymodon guineensis Broth. & Par. GUI(47). 

Didymodon integrifolius var. integrifolius Broth. 

RWA(65). 

Didymodon integrifolius var. paucidentatus Thér. = 

Bryoerythrophyllum campylocarpum (1420). 

Didymodon jackvancei Zander LES(1531). 

Didymodon michiganensis (Steere) K.Saito 

ETH(758), REU(758). 

Didymodon papillinervis (Dix. ex Herz.) Demar. = 

Bryoerythrophyllum recurvirostrum (1420). 

Didymodon paucidentatus (C.Müll.) Broth. = Bryoe- 

rythrophyllum campylocarpum (1420). 

Didymodon recurvirostris (Hedw.) Jenn. = Bryoeryth- 

rophyllum recurvirostrum (1521). 

Didymodon recurvirostris var. angustifolius (P.Varde) 

Demar. = Bryoerythrophyllum recurvirostrum 

var. angustifolium (46) (= B. campylocarpum). 

Didymodon rigidifolius Dix. = Bellibarbula recurva 

(1420). 

Didymodon rigidulus var. acutus Biz. TAN(46). 

Didymodon rigidulus var. gracilis (Schleich. ex 

Hook. & Grev.) Zander KEN(411), SUD(1419). 

Didymodon subfontanus Dix. MAL(67). 

Didymodon sublingulatus Dix. = Bryoerythrophyllum 

campylocarpum (1420). 

Didymodon subrevolutus (Hampe) Broth. 

MAD(40). 

Didymodon subulirostris Schimp. ex Par., nom. 

nud. ETH(50). 

Didymodon tophaceopsis Zander CBS(1531), 

LES(1531), NAM(1531), NAT(1531), 

OFS(1531), ZAM(62), ZIM(46). 

Didymodon tophaceus (Brid.) Lisa Sollman (pers. 

comm.) has seen specimens of this taxon from

South Africa, and it is likely that some collections 

listed for D. ceratodonteus may belong here. 

Didymodon tophaceus fo. humilis (Schimp.) Marg. 

= Barbula tophaceus var. humilis (1420). 

Didymodon trivialis (C.Müll.) Guerra CBS(1531), 

LES(1531), NAT(1531), OFS(1531). 

Didymodon xanthocarpus (C.Müll.) Magill 

CBS(1531), LES(1531), NAT(1531), TRV(1531). 

Dimerodontium africanum C.Müll. CBS(1531), 

NAT(1531), TRV(1531). 

Dimerodontium kenyae Dix. ex Tosco & Piovano 

KEN(46). 

Diphyscium pocsii (Biz.) Zander TAN(46). 

Diplostichum longirostre (Brid.) Mont. = Eustichia 

longirostris (83). 

Distichium capillaceum (Hedw.) BS&G CAM(47), 

ETH(50), LES(1531), MAD(40), NAT(1531), 

TAN(46). 

Distichium kilimandscharicum C.Müll. KEN(46), 

TAN(46). 

Distichophyllidium africanum Demar. & P.Varde 

TAN(46), ZAI(65). 

Distichophyllum annobonense Broth. in Mildbr., 


Distichophyllum mascarenicum Besch. 

REU(105), SEY(40). 

Distichophyllum mniifolium (Hornsch.) Sim 

CBS(1531), TAN(46). 

Distichophyllum procumbens Mitt. CAM(47), 

FER(47), GHA(47). 

Distichophyllum rakotomariae Crosby 

MAD(40). 

Distichophyllum rigidicaule (Dus.) Broth. 

CAM(47), GAB(47), IVO(1880). 

Distichophyllum rigidicaule var. gabonense 

(P.Varde) P.Varde GAB(47), ZAI(65). 

Ditrichum brachypodum (C.Müll.) Broth. 

CBS(1531), NAT(1531), OFS(1531), 

SWA(1531), TRV(1531), ZIM(61). 

Ditrichum difficile (Duby) Fleisch. 

CBS(1531,1882), KEN(46), MAD(40), MAL(67), 

MAU(105), NAT(1531), REU(105), RWA(65), 

SWA(1531), TAN(46), TRV(1531), ZIM(61). 

Ditrichum luteum Dix. & Thér. MAD(40). 

Ditrichum madagassum (Ren. & Card.) Par. 

MAD(40). 

Ditrichum pallidum (Hedw.) Hampe KEN(46), 

MAD(40), TAN(46), UGA(46), ZAI(65). 

Ditrichum postscriptum Hornsch. = Campylopodium 

proscriptum (1521). 

Ditrichum punctulatum Mitt. CBS(1531), 

NAT(1531). 

Ditrichum strictum (Hook.f. & Wils.) Hampe 

LES(1531), NAT(1531). 

Ditrichum tisserantii P.Varde ANG(46). 

Donnellia laxiuscula (Card.) Buck ZAI(65). 

Donnellia matutina Buck RWA(65), ZAI(65). 

Drepanocladus aduncus (Hedw.) Warnst. 

KEN(46), RWA(65), ZAI(65), ZIM(61). 

Drepanocladus exannulatus (Schimp.) Warnst. = 

Warnstorfia exannulata (453). 

Drepanocladus fluitans (Hedw.) Warnst. = Warnstorfia 

fluitans (1124). 

Drepanocladus fluitans var. monodii P.Varde = Warn- 

storfia fluitans (According to Ochyra (pers. 

comm.) this is just an aquatic phenotype of 

otherwise typical W. fluitans.). 

Drepanocladus hallii Broth. & Dix. CBS(1531), 

NAT(1531). 

Drepanocladus hallii Broth. & Dix. According to 

Ochyra (pers. comm.) the type of this taxon is D. 

sendtneri, but this has not yet been published. 

Drepanocladus sparsus C.Müll. CBS(1531), 

LES(1531), NAT(1531), OFS(1531), TRV(1531), 

ZIM(46). 

Drepanocladus sparsus C.Müll. According to Ochyra 

(pers. comm.) the type of this taxon is 

Leptodictyum riparium (=Amblystegium 

riparium in this checklist), but this has not yet 

been published. 

Drepanocladus uncinatus (Hedw.) Warnst. = Sanionia 

uncinata (453). 

Eccremidium exiguum (Hook.f. & Wils.) Hampe 

NAT(1531), TRV(1531). 

Ectropothecium affine Broth. TAN(46). 

Ectropothecium afro-molluscum Broth. 

CAM(47), CAR(47), GAB(47), GHA(47), 

SIL(47), TAN(46), ZAI(65). 

Ectropothecium anisophyllum Broth. CAM(47), 

CAR(47), CON(47), GAB(47), GUI(47), 

IVO(1880), LIB(47), ZAI(65). 

Ectropothecium aureo-crispum Broth. CAM(47). 

Ectropothecium bingerianum Par. & Broth. 

GUI(47). 

Ectropothecium brachycarpum (Dix.) Magill 

CBS(1531), MOZ(46), NAT(1531), TRV(1531), 

ZIM(61). 

Ectropothecium brachycladulum (C.Müll.) Broth. 

IVO(1880), TAN(46). 

Ectropothecium brevifalcatum (C.Müll.) Kindb. 

119

120 

CAM(47), GUI(47), MAI(47), SAO(47), 

TOG(1878). 

Ectropothecium buluense Broth., nom. nud. 

TAN(46). 

Ectropothecium capillisetum (C.Müll.) Kindb. 

ANG(46). 

Ectropothecium chenagonii Ren. & Card. 

MAD(40). 

Ectropothecium chenagonii var. hamatulum Ren. 

& Card. MAD(40). 

Ectropothecium dafilae Broth. & Par. GUI(47). 

Ectropothecium diffusum (Mitt.) Jaeg. FER(47), 

PRI(47). 

Ectropothecium doliare Mitt. ROD(105). 

Ectropothecium dolosum Card. ZAI(65). 

Ectropothecium drepanophyllum Broth. 

SAO(47). 

Ectropothecium duemmeri Dix. TAN(46), 

UGA(46). 

Ectropothecium engleri Broth. TAN(46). 

Ectropothecium engleri var. planissimum Broth. 

LIB(47), TAN(46). 

Ectropothecium fiorinae Tosco & Piovano 

TAN(46). 

Ectropothecium guineense Par. & Broth. GUI(47), 

IVO(47). 

Ectropothecium intertextum Ren. & Card. 

MAD(40), MAU(105). 

Ectropothecium isopterygioides Broth., nom. 

nud. TAN(46). 

Ectropothecium latericola Broth. CAM(47), 

KEN(46), UGA(46). 

Ectropothecium leptoblastum Broth., nom. nud. 

TAN(46). 

Ectropothecium mauritianum Ren. & Card. 

MAD(40), MAU(105). 

Ectropothecium micro-pyxis Broth., nom. nud. 

TOG(58). 

Ectropothecium mollicellum (Mitt.) Jaeg. 

CAM(47). 

Ectropothecium occultum Ren. & Card. COM(40), 

MAD(40), REU(105). 

Ectropothecium paillotii Ren. & Card. MAD(40), 

MAU(105), ZIM(61). 

Ectropothecium perrevolutum P.Varde CAR(47), 

GUI(47), IVO(1880), TOG(1878). 

Ectropothecium perrieri Thér. MAD(40). 

Ectropothecium perrotii Ren. & Card. MAD(40), 

MAL(67), TRV(1531), ZAI(65), ZIM(46,61). 

Ectropothecium protractulum (C.Müll.) Broth. 

COM(40). 

Ectropothecium regulare (Brid.) Jaeg. BUR(65), 

CAM(47), CBS(1531), COM(40), CON(47), 

GAB(47), GUI(47), IVO(1880), MAD(40), 

MAL(67), MAU(105), NAT(1531), REU(105), 

SIL(47), TAN(46), TRV(1531), ZAM(62), 

ZIM(46,61). 

Ectropothecium regulare var. angustifolium Thér. 

MAU(105). 

Ectropothecium regulare var. laxitextum Thér. 

REU(105). 

Ectropothecium revolutum Broth. CAM(47), 

IVO(1880), TAN(46), ZAI(65). 

Ectropothecium sericeum P.Varde TAN(46). 

Ectropothecium seychellarum Besch. MAD(40), 

SEY(40). 

Ectropothecium subulosum Mitt. ROD(105). 

Ectropothecium tamatavense Broth. MAD(40). 

Ectropothecium taxiforme (Brid.) Broth. ANN(47), 

CAM(47), LIB(47). 

Ectropothecium triviale (C.Müll.) Kindb. FER(47). 

Ectropothecium valentinii Besch. MAU(105), 

REU(105). 

Ectropothecium valentinii var. minus Ren. 

MAU(105). 

Ectropothecium viridulum (Brid.) Jaeg. REU(105). 

Encalypta ciliata Hedw. CBS(1531), ETH(50), 

LES(1531), NAT(1531), OFS(1531), TAN(46). 

Encalypta cuspidata B&S ex C.Müll. ETH(50). 

Encalypta hedbergii P.Varde TAN(46). 

Encalypta vulgaris Hedw. CBS(1531), LES(1531). 

Encalypta vulgaris var. intermedia Dix. ERI(50). 

Entodon adyris Buck ZAI(65). 

Entodon breve (Schimp.) Par. ETH(50). 

Entodon brevirameus Dix. = E. macropodus (53). 

Entodon cameruniae C.Müll. in Par., nom. nud. = 

Entodontella cameruniae (715). 

Entodon corbieri Ren. & Card. MAD(40). 

Entodon cymbifolius Wager ex Dix. LES(1531), 

NAT(1531), TRV(1531), ZIM(46). 

Entodon denticuspis P.Varde UGA(46), ZAI(715). 

Entodon dregeanus (Hornsch.) C.Müll. = E. macro- 

podus (70). 

Entodon dregeanus var. borbonicus Ren. & Card. 

REU(105). 

Entodon dregeanus var. borbonicus Ren. & Card. E. 

dregeanus is now a synonym of E. macropodus 

(70). This variety is also in need of revision. 

Entodon felicis Ren. & Card. MAD(40). 

Entodon felicis var. julaceus Ren. & Card.

MAD(40). 

Entodon geminidens (Besch.) Broth. CAM(47), 

REU(715), TAN(46), TRV(1531). 

Entodon lacunosus Broth. TAN(46). 

Entodon macropodus (Hedw.) C.Müll. BUR(65), 

CBS(1531), KEN(46), MAL(67,70), NAT(1531), 

NIG(334), OFS(1531), RWA(65), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZAM(46), ZIM(46,70). 

Entodon madagassus Geh. MAD(40). 

Entodon motelayi (Besch.) Par. MAU(105,715). 

Entodon natalensis Rehm. ex C.Müll. LES(1531), 

NAT(1531), TRV(1531). 

Entodon piovanoi Biz. ETH(715), TAN(46). 

Entodon pseudo-seductrix C.Müll. CAP(1296). 

Entodon rutenbergii Geh. MAD(40). 

Entodon stereophylloides Broth. TAN(46), 

UGA(715). 

Entodon subcompressus (C.Müll.) Fleisch., nom. 

nud. ETH(50), TAN(46). 

Entodon subgeminidens P.Varde CAR(47), 

CON(715). 

Entodon usambaricus Broth. KEN(46), TAN(46). 

Entodon vulcanicus Demar. & Leroy KEN(46), 

RWA(65), TAN(46), ZAI(65). 

Entodontella cameruniae Broth. CAM(47), 

GUI(47). 

Entodontopsis leucostega (Brid.) Buck & Ireland 

ANG(46), TOG(1878). 

Entodontopsis nitens (Mitt.) Buck ANG(46), 

CAM(47), CAR(47), COM(40), FER(47), 

GAB(47), GHA(47), GUI(47,1458), 

IVO(1458,1880), KEN(411), LIB(47), MAD(40), 

MAL(67), NIG(47,334), RWA(65), TAN(46), 

TOG(1878), UGA(46), ZAI(65), ZAM(62). 

Entodontopsis rhynchostegioides (Broth. & Par.) 

Buck & Ireland GUI(47). 

Entodontopsis rufescens (Broth.) Buck & Ireland 

TAN(46). 

Entodontopsis tenuinervis (Broth.) Buck & Ireland 

CAR(47), GHA(47), GUI(1458,47), 

MAD(40), MAL(67), NIG(334), SIL(47), 

ZAI(65). 

Entosthodon amblyodon Schimp. in Jaeg. 

ETH(50). 

Entosthodon bergianus (Hornsch.) C.Müll. 

CBS(1531), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), TRV(1531). 

Entosthodon borbonicus Besch. MAD(40), 

MAU(105), REU(105), RWA(65). 

Entosthodon clavatus Mitt. CBS(1531). 

Entosthodon curvipes (C.Müll.) C.Müll. CAM(47), 

ETH(50). 

Entosthodon curvipes var. robustior C.Müll. 

ETH(50). 

Entosthodon dixonii Sim TRV(1531), ZIM(46,61). 

Entosthodon hildebrandtii C.Müll. KEN(46), 

TAN(46). 

Entosthodon holstii (Broth.) Par. TAN(46). 

Entosthodon lepervanchei Besch. REU(105). 

Entosthodon limbatus C.Müll. CBS(1531), 

NAT(1531), TAN(1410), TRV(1531), ZIM(61). 

Entosthodon marginatulus C.Müll. ex Geh. 

MAD(40). 

Entosthodon marginatus C.Müll. TAN(46), 

ZAM(46), ZIM(46). 

Entosthodon mauritianus Schimp. ex Besch. 

MAU(105). 

Entosthodon pertenellus (Broth.) Kis TAN(46). 

Entosthodon rottleri (Schwaegr.) C.Müll. 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TAN(46), TRV(1531). 

Entosthodon urceolatus Mitt. CBS(1531), 

NAT(1531), OFS(1531), ZIM(61). 

Entosthodon usambaricus (Broth.) Par. TAN(46). 

Entosthodon volkensii (Broth.) Par. KEN(46), 

TAN(46), UGA(46), ZAI(65). 

Entosthodon wichurae Fleisch. UGA(46). 

Ephemerum capense C.Müll. CBS(1531). 

Ephemerum diversifolium Mitt. in Harv. 

CBS(1531). 

Ephemerum namaquense Magill CBS(1531). 

Ephemerum pechuelii C.Müll. CAR(47), NIG(334), 

ZAI(65). 

Ephemerum perminutum C.C.Townsend 

TAN(46). 

Ephemerum rehmannii (C.Müll.) Broth. 

CBS(1531), NAT(1531), OFS(1531), ZIM(46). 

Epipterygium convalleum Dus. CAM(47). 

Epipterygium diversifolium Ren. & Card. 

MAD(40). 

Epipterygium tozeri (Grev.) Lindb. RWA(65). 

Eriopus cristatus (Hedw.) Brid. = Calyptrochaeta 

cristata (See “New Combinations”). 

Erpodium beccarii C.Müll. ex Vent. BOT(1531), 

ERI(50), KEN(46), MAD(40), NAM(1531), 

NAT(1531), OFS(1531), SOM(50), TAN(46), 

TRV(1531), ZAM(46), ZIM(46,61). 

Erpodium biseriatum (Aust.) Aust. TAN(46). 

Erpodium coronatum (Hook. & Wils.) Mitt. 

121

122 

BUF(47), BUR(65), CAM(47), CAR(47), 

GUI(47,1458), IVO(1458), MAI(47), NIG(334), 

SEN(47), ZAI(1410), ZAM(62). 

Erpodium distichum Wag. & Dix. NAT(273), 

TRV(1531). 

Erpodium grossirete C.Müll. MOZ(46), 

NAM(1531), ZIM(46,61). 

Erpodium hanningtonii Mitt. = E. beccarii (46). 

Erpodium holstii Broth. KEN(46), TAN(46). 

Erpodium madagassum Par. & Ren. MAD(40). 

Erpodium perrottetii (Mont.) Jaeg. & Sauerb. 

SEG(273). 

Erpodium pobeguinii Par. & Broth. = E. coronatum 

(273). 

Erpodium schimperi C.Müll. ex Jaeg., nom. nud. 

ETH(50). 

Erpodium theriotii Broth. in Corb. SEN(47). 

Erpodium transvaaliense Broth. & Wag. ex Dix. 

NAM(1531), NAT(1531), TRV(1531). 

Erythrodontium barteri (Mitt.) Broth. ANG(46), 

CAM(47), CAR(47), GAB(47), GUI(47), IVO(63), 

NIG(47,334), TOG(1878). 

Erythrodontium bicolor Gepp in Hiern, hom. 

illeg. ANG(46). 

Erythrodontium cameruniae C.Müll. CAM(47), 

GUI(47). 

Erythrodontium engleri (Broth.) Par. KEN(46), 

MAL(67), TAN(46), UGA(46). 

Erythrodontium julaceum Hook. STH(1296). 

Erythrodontium lacoutourei Ren. & Card. 

MAD(40). 

Erythrodontium lamoruense Thér. KEN(46). 

Erythrodontium perjulaceum Par., nom. nud. 

CAM(47), GUI(47). 

Erythrodontium rotundifolium (C.Müll.) Par. 

TAN(46), ZIM(46). 

Erythrodontium squarrosum (Hampe) Par. 

GUI(1458), IVO(1458), RWA(65). 

Erythrodontium subbiforme (Ren. & Card.) Broth. 

ZAI(65). 

Erythrodontium subjulaceum (C.Müll.) Par. 

ANG(46), CAM(47), CAR(47), GAB(47), 

GUI(47), KEN(46), MAL(67), NIG(334), 

RWA(65), SWA(1531), TAN(46), TRV(1531), 

UGA(46), ZAI(65), ZAM(62), ZIM(61). 

Erythrodontium subjulaceum var. chalcophyllum 

(Thér. & P.Varde) P.Varde CAR(47). 

Erythrodontium subjulaceum var. impolitum 

Gepp. ANG(46). 

Erythrodontium vanderystii Card. ZAI(65). 

Eucladium verticillatum (Brid.) BS&G RWA(65), 

TAN(46), ZAI(65). 

Eurhynchiella decurrens P.Varde KEN(46), 

RWA(46), TAN(46), UGA(46), ZAI(65). 

Eurhynchium acicladium Besch. REU(105). 

Eurhynchium hians (Hedw.) Sande Lac. IVO(44), 

KEN(46), TAN(46), UGA(46). 

Eurhynchium praelongum var. praelongum BS&G = 

Kindbergia praelonga var. praelonga (1640) 

Eurhynchium praelongum var. stokesii (Turn.) Dix. 

= Kindbergia praelonga var. stokesii (1640) 

Eurhynchium spiculosum (Mitt.) Jaeg. CAM(47). 

Eurhynchium spinulinerve Card. MAD(40). 

Eurhynchium stokesii Turn. = E. praelongum var. 

stokesii (1521) (= Kindbergia praelonga var. 

stokesii). 

Eurhynchium stokesii var. teneriffae Ren. & Card. = 

E. praelongum (1295) (= Kindbergia praelonga 

var. praelonga). 

Eurhynchium striatum (Hedw.) Schimp. MAD(40). 

Eurhynchium tenuivagum (C.Müll. ex Broth.) 

Buck CAM(47), CAR(47), KEN(46), UGA(46). 

Eustichia longirostris (Brid.) Brid. LES(1531), 

MAD(40), NAT(1531), OFS(1531), REU(105). 

Fabronia abyssinica C.Müll. CBS(1531), 

DJI(1569), ERI(50), ETH(50), KEN(46), 

NAT(1531), SUD(1419), TAN(46), TRV(1531), 

ZAI(65), ZAM(46), ZIM(46). 

Fabronia angolensis Welw. & Duby = F. pilifera (46) 

Fabronia bizotii Pócs CAM(47). 

Fabronia breutelii Hampe in C.Müll. CBS(1531). 

Fabronia campenonii Ren. & Card. MAD(40). 

Fabronia claviramea Broth. in Mildbr. RWA(65). 

Fabronia condensata Schimp. ex Jaeg. ETH(50). 

Fabronia congolensis Card. TAN(46), ZAI(65). 

Fabronia crassiretis Ren. & Card. CAM(47), 

MAD(40). 

Fabronia eckloniana Hampe in C.Müll. 

CBS(1531). 

Fabronia fastigiata Ren. & Card. MAD(40). 

Fabronia fastigiata var. asperula Ren. & Par. 

MAD(40). 

Fabronia garnieri (Par. & Ren.) Ren. & Par. 

MAD(40). 

Fabronia goetzei Broth. TAN(46). 

Fabronia gueinzii Hampe CBS(1531), NAT(1531), 

TAN(259), TRV(1531). 

Fabronia lachenaudii Ren. MAD(40). 

Fabronia lachenaudii var. nairobiensis Thér. & 

P.Varde CAR(47), KEN(46).

Fabronia leikipiae C.Müll. CBS(1531), KEN(46), 

TAN(46,259), TRV(1531), ZAM(62), ZIM(46). 

Fabronia longipila Broth. KEN(46), TAN(46), 

UGA(46), ZAM(46). 

Fabronia longipila var. gracilis Broth. TAN(46). 

Fabronia motelayi Ren. & Card. MAD(40). 

Fabronia perciliata C.Müll. CBS(1531), 

NAT(1531), TRV(1531). 

Fabronia pilifera Hornsch. ANG(46), BOT(1531), 

BUF(47), CBS(1531), CAR(47), ETH(50), 

KEN(46), LES(1531), MAL(67), MOZ(357), 

NAT(1531), OFS(1531), SWA(1531), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZAM(46,62), 

ZIM(46). 

Fabronia pobeguinii Broth. & Par. CAR(47), 

GUI(47). 

Fabronia pobeguinii var. subintegrifolia Par. 

GUI(47). 

Fabronia pocsii var. cameruniae Bizot = F. bizotii 

(108). 

Fabronia pocsii var. pocsii Bizot ETH(1886). 

Fabronia rehmannii C.Müll. CBS(1531), ERI(50), 

MAL(67), NAT(1531), SWA(1531), TRV(1531), 

ZIM(46). 

Fabronia schimperiana Hampe ex Jaeg., hom. 

illeg. ETH(50), TAN(46). 

Fabronia socotrana Mitt. SOC(40). 

Fabronia trichophora Card. ANG(46). 

Fabronia trichophylla C.Müll. ERI(50). 

Fabronia victoriae Dix. CBS(1531), TRV(1531), 

ZIM(46). 

Fabronia villaumii Ren. & Card. MAD(40). 

Fabronia wageri Dix. CBS(1531). 

Fissidens acanthophyllus P.Varde = F. smilioides 

(1749). 

Fissidens aciphyllus Dix. CBS(1531), NAT(1531), 

OFS(1531), TRV(1531). 

Fissidens aegrotus Biz. = F. asplenioides var. aegrotus 

(1748). 

Fissidens afropapillosus P.Varde KEN(46). 

Fissidens ake-assii Bizot & Dury TOG(4). 

Fissidens algarvicus Solms TAN(46). 

Fissidens alomoides C.Müll. ex Dus. CAM(47), 

CAR(47), GUI(47), IVO(47,1880), SIL(47). 

Fissidens ambiguus Biz. ex Pócs = F. intramarginatus 

(90). 

Fissidens amblyophyllus C.Müll. = F. asplenioides 

(90). 

Fissidens androgynous Bruch & Krauss = F. bryoides 

(82). 

Fissidens angolensis Welw. & Duby ANG(46), 

KEN(46), TAN(46), UGA(46). 

Fissidens angustinervis Thér. MAD(40). 

Fissidens aoristoloma P.Varde = F. borgenii (90). 

Fissidens arbogastii Ren. & Card. = F. zippelianus 

(76). 

Fissidens arboreus Broth. TOG(1878). 

Fissidens arenivagus P.Varde ZAI(65). 

Fissidens argyroloma P.Varde GAB(47). 

Fissidens asplenioides Hedw. CAM(47), 

CBS(1531), COM(105,40), GUI(47), KEN(46), 

MAD(40), MAL(67), MAS(40), NAT(1531), 

REU(105), RWA(65), TAN(46), TRV(1531), 

ZAI(65), ZAM(62), ZIM(46). 

Fissidens asplenioides var. aegrotus (Biz.)Brugg.- 

Nann., Pursell & Iwats TAN(46). 

Fissidens asselii Biz. CAR(47,1749). 

Fissidens atroviridis Besch. COM(40), MAD(40), 

MAU(105), TAN(46), ZAM(46). 

Fissidens badoglioi Tong. ETH(50). 

Fissidens bambariensis P.Varde = F. desertorum 

(1749). 

Fissidens basicarpus C.Müll. LIB(47). 

Fissidens bessouensis Corb. CAR(47), 

ZAI(65,1552). 

Fissidens bocarangensis P.Varde CAR(47). 

Fissidens bogosicus C.Müll. in De Vent. ERI(41). 

Fissidens boivinianus var. boivinianus Besch. = F. 

asplenioides (1748). 

Fissidens boivinianus var. longifolius Besch. 

MAS(40), TAN(46). 

Fissidens boivinianus var. major Par., nom. nud. 

MAS(40). 

Fissidens bonnotii Bizot CAR(1749), GAB(47), 

TAN(46). 

Fissidens borgenii Hampe BUF(47), CAM(47), 

CBS(1531), CAR(47,1749), CHA(90), CON(47), 

GAB(47), IVO(1880), LIB(47), MAD(90), 

MOZ(46), NAT(1531), NIG(47,334), OFS(1531), 

RWA(65), SAF(90), SIL(47), TAN(46), 

TRV(1531), ZAI(65), ZIM(46). 

Fissidens boryanus Besch. MAS(40). 

Fissidens brachycaulon Broth. TAN(46). 

Fissidens brenanii P.Varde CAM(47). 

Fissidens brevifrons Mitt. ROD(105). 

Fissidens brotheri Par. TOG(58). 

Fissidens brunnthaleri Broth. CAR(47), NIG(47), 

TAN(46). 

Fissidens brunnthaleri var. filipes Dix. & P.Varde 

CAR(47). 

123

124 

Fissidens bryoides Hedw. ANG(46), CAM(47), 

CBS(1531), CAR(47,1749), ETH(50), GAB(47), 

GHA(47), GUI(47), IVO(63,1880), KEN(46), 

LES(1531), MAL(67), MAS(40), NAT(1531), 

NIG(47,334), OFS(1531), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZIM(46). 

Fissidens bryoides ssp. viridulus (Sw.) Kindb. 

CAM(47). 

Fissidens bryoides var. crispulus (Brid.) Wijk & 

Marg. MAS(40). 

Fissidens bryoides var. longifolius (Brid.) Hampe 

MAS(40). 

Fissidens bryum C.Müll. ex Dus. CAM(47), 

CAR(90,1749), GAB(47), GUI(47), NIG(47,334), 

RWA(65), ZAI(65). 

Fissidens bryum var. terrestris P.Varde GAB(47). 

Fissidens buettneri Broth. CAR(47,1749), 

GHA(47), GUI(47), IVO(1880), LIB(47), TOG(58). 

Fissidens bukobensis Broth. in Mild. = F. intramar- 

ginatus (90) 

Fissidens cadetii Biz. REU(105). 

Fissidens calabariae C.Müll. ex Dusen. = F. laxus 

(90). 

Fissidens calcicolus C.Müll. SOM(50). 

Fissidens caloglottis C.Müll. = F. asplenioides (1748). 

Fissidens cameruniae C.Müll. = F. zippelianus (76). 

Fissidens capriviensis Magill NAM(1531). 

Fissidens casamancae Card. SEN(47). 

Fissidens cataractarum Demar. & P.Varde 

ZAI(65). 

Fissidens centrafricanus Biz. CAR(47,1749), 

GUI(47). 

Fissidens chevalieri Corb. CAR(47), NIG(334). 

Fissidens chioneuros C.Müll. = F. helenicus (1854, 

p. 18). 

Fissidens comorensis C.Müll. COM(40), 

MAD(40), MAS(40), NIG(334), TAN(46). 

Fissidens comorensis var. acuminatus Besch. 

COM(40), MAD(40). 

Fissidens comorensis var. sordidus Ren. & Card. 

MAD(40). 

Fissidens congolensis Ren. & Card. BEN(47), 

CAM(47), CAR(47,1749), GAB(47), GHA(47), 

GUI(47), MAL(67), NIG(47), SIL(47), TAN(46), 

ZAI(65). 

Fissidens coriaceifolius C.Müll. = F. zippelianus (76). 

Fissidens corrugatulus Dix. ZAM(62), 

ZIM(46,61). 

Fissidens crassipes ssp. warnstorfii (Fleisch.) 

Brugg.-Nann. TAN(46), ZAM(46), ZIM(61,46). 

Fissidens crassipes var. philibertii Besch. = F. crassipes 

ssp. warnstorfii (1898). 

Fissidens crateris Dix. = F. bryoides (90). 

Fissidens crateris var. sererikae Dix. = F. bryoides 

(90). 

Fissidens cremersii Biz. & Onr. = F. laxus (90). 

Fissidens crispatulus Ren. & Card. = F. zippelianus 

(76). 

Fissidens crispifolius Broth. & P.Varde CAR(47), 

GAB(47), NIG(47). 

Fissidens crispopachyloma Dix. = F. leucocinctus 

(90). 

Fissidens crispulus Brid. MAS(40). 

Fissidens cryptoneuron P.Varde CAR(47), 

GUI(47), MAD(40), UGA(46), ZAM(46). 

Fissidens curvatus Hornsch. CBS(1531), 

NAT(1531), OFS(1531), SWA(1531), 

TRV(1531), ZIM(46,61). 

Fissidens curvifolius Mitt. NIG(47,334), TAN(46). 

Fissidens cuynetii Biz. ex Pócs UGA(46). 

Fissidens danckelmannii C.Müll. EGU(47). 

Fissidens darntyi Schimp. CAM(47), IVO(1880), 

MAD(40), MAU(105), REU(105), TAN(46). 

Fissidens dasyphus Welw. & Duby ANG(46), 

ZAI(65). 

Fissidens decaryi Biz. MAD(40). 

Fissidens dendeliensis Par. & Broth. CAM(47), 

GAB(47), GUI(47), NIG(334). 

Fissidens desertorum (C.Müll.) Par. CAR(47,1749), 

NIR(79), NIG(47), TAN(46), TOG(1878), 

ZAI(65). 

Fissidens deslooveri Biz.. = F. intramarginatus (90). 

Fissidens diaphanodontus (P.Varde) Biz. 

CAR(47), TAN(46). 

Fissidens diaphanus Biz. CAM(47), 

CAR(47,1749), IVO(1880), NIG(47). 

Fissidens dispersus Card. = F. laxus (90). 

Fissidens dubiosus Dix. GAB(47). 

Fissidens dubiosus var. fluitans P.Varde GAB(47). 

Fissidens dubyanus Jaeg. ANG(46), GAB(47), 

GUI(47). 

Fissidens durus Par. & Broth. CAR(47), GUI(47). 

Fissidens eckendorfii P.Varde CAM(47), 

CAR(47,1749), NIG(334). 

Fissidens eenii Biz. = F. papillosus (580). 

Fissidens ellipticus Besch. REU(105). 

Fissidens enervis Sim NAT(1531), TAN(46), 

ZIM(61). 

Fissidens enervis ssp. hedbergii (P.Varde) Biz. 

TAN(46), ZAM(46).

Fissidens erikssonii Bartr. SEY(40). 

Fissidens erosulus (C.Müll.) Par. BOT(1531), 

CBS(1531), CAR(47), LES(1635), MOZ(46), 

NAT(1531), SWA(1531), TRV(1531), UGA(46), 

ZAM(62), ZIM(46). 

Fissidens eschowensis Broth. & Bryhn = F. submar- 

ginatus (82). 

Fissidens euryloma P.Varde GAB(47). 

Fissidens exasperatus Ren. & Card. = F. papillosus 

Sande Lac (76). 

Fissidens fasciculatus Hornsch. CBS(1531), 

NAT(1531), TRV(1531). 

Fissidens ferrugineus C.Müll. MAD(40), 

REU(105). 

Fissidens filiicola Par. & Broth. GUI(47). 

Fissidens flaccidus Mitt. NIG(47,334), ROD(105), 

ZIM(61). 

Fissidens flavo-limbatus Besch. = F. intramarginatus 

(90). 

Fissidens fluminalis Dus. CAM(47), CAR(47), 

GUI(47). 

Fissidens fontanus (Pyl.) Stend. = F. palmifolius (85). 

Fissidens fouta-djallonii Par. = F. gardneri (1470). 

Fissidens foveolatus Card. SEN(47), ZAI(65). 

Fissidens fremyi P.Varde = F. zippelianus (76). 

Fissidens fusco-lutescens Card. = F. zippelianus (76). 

Fissidens gabonensis Broth. & P.Varde GAB(47). 

Fissidens gardneri Mitt. CAR(47), GUI(47), 

NIG(47,334), SEN(47), TAN(46). 

Fissidens ghanae Biz. = F. laxus (90). 

Fissidens gibbonii Biz.ex Pócs = F. intramarginatus 

(90) 

Fissidens gladiolus Mitt. CAR(47,1749), GAB(47), 

NIG(47,334), TAN(46), UGA(46). 

Fissidens glaucescens Hornsch. CBS(1531), 

KEN(46), LES(1531), MAS(40), NAT(1531), 

OFS(1531), SWA(1531), TAN(46), TRV(1531), 

ZAI(65), ZAM(62), ZIM(46,61). 

Fissidens glaucissimus Welw. & Duby ANG(46), 

CAM(47), CAR(47,1749), GAB(47), GUI(47), 

IVO(1880), NIG(47,334), PRI(47), SAO(47), 

TAN(46), TOG(58), UGA(46), ZAI(65). 

Fissidens glaucissimus var. minor Welw. & Duby 

ANG(46). 

Fissidens glaucopteris C.Müll. ex Dus. IVO(1880), 

LIB(47), SIL(47), TOG(1878). 

Fissidens glauculus C.Müll. ex Dus. BUF(1423), 

CAM(47), CAR(47,1749), GAB(47), GHA(47), 

GUI(47), IVO(63,1880), LIB(47), NIG(47,334), 

TOG(1878), UGA(46), ZAI(65). 

Fissidens glauculus ssp. dupuisii (Ren. & Card.) 

P.Varde = F. glauculus (90). 

Fissidens glauculus var. aculeifrons P.Varde in 

Thér. GAB(47). 

Fissidens glauculus var. circinicaulis (Card.) 

P.Varde GAB(47), SEN(47). 

Fissidens glauculus var. gymnandrus P.Varde 

GAB(47). 

Fissidens gomae P.Varde & Leroy TAN(46), 

ZAI(65). 

Fissidens grainvillei Thér. MAD(40). 

Fissidens grandifolius Broth. & P.Varde CAR(47), 

GAB(47), GUI(47), LIB(47), NIR(79), NIG(47), 

TOG(1878). 

Fissidens grandifrons Brid. MAS(40). 

Fissidens grandifrons var. flaccidus Brid. 

MAS(40). 

Fissidens grandiretis Ren. & Card. = F. laxus (90). 

Fissidens grossiretis C.Müll. CAM(47), 

CAR(1749), GAB(47), IVO(1880). 

Fissidens guineensis Par. & Broth. = F. microcarpus 

(73). 

Fissidens gumangensis (C.Müll.) Par. = F. sciophyllus 

(90). 

Fissidens haakonii Broth. & Bryhn = F. borgenii 

(82). 

Fissidens helenicus Mitt. STH(1296). 

Fissidens helictocaulos C.Müll. ERI(50), 

TAN(46), ZAM(46). 

Fissidens herpetineuron Broth. & P.Varde = F. mo- 

nostictus (1749) (= F. zollingeri). 

Fissidens hians (Hampe) Par. CAR(47), GAB(47), 

GUI(47), NIG(47,334). 

Fissidens hildebrandtii C.Müll. SOM(50). 

Fissidens hillieri Thér. MAD(40). 

Fissidens hirsutus Biz. ex Pócs TAN(46). 

Fissidens hoeegii P.Varde CBS(1531), NAT(1531), 

OFS(1531), TAN(46), TRV(1531). 

Fissidens holstii Broth. = F. laxus (90). 

Fissidens holstii var. perintegrifolius P.Varde = F. 

laxus (90). 

Fissidens hookerioides Biz. & Onr. MAD(40). 

Fissidens horizonticarpus C.Müll. CAR(47,1749), 

CON(47), GUI(47), TAN(46). 

Fissidens hymenodon Besch. COM(40), 

MAD(40), TAN(46). 

Fissidens idanreensis P.Varde NIG(47), RWA(65), 

TAN(46), ZAM(46). 

Fissidens inaequalilimbatus Par. & Broth. 

CAR(47,1749), GUI(47). 

125

126 

Fissidens inclinatulus (C.Müll.) Par. 

CAR(47,1749), NIG(47). 

Fissidens inclusus Biz. ex Dury & Pócs TAN(46). 

Fissidens inflatus (C.Müll.) Par. CAR(1749), 

NIG(47), ZAI(65). 

Fissidens intralimbatulus Broth. & P.Varde = F. 

borgenii (90). 

Fissidens intralimbatulus var. brevifolius P.Varde 

BUF(47). 

Fissidens intralimbatulus var. brevifolius P.Varde. 

The owning species of this variety has been made 

a synonym of F. borgenii. This variety also needs 

revision. 

Fissidens intramarginatus (Hampe) Mitt. 

KEN(46), MAD(40), REU(105), RWA(65), 

SUD(90), TAN(46), UGA(46), ZAI(65). 

Fissidens itarensis Broth. = F. intramarginatus (90) 

Fissidens jeffreyi Biz. SEY(40). 

Fissidens jonesii Biz. ex Pócs TAN(46). 

Fissidens jonesii fo. terrestris Bix. & Pócs. Excluded. 



Fissidens jordanii J.Tayl. & P.Varde SIL(47). 

Fissidens jovet-astiae Biz. CAR(47,1749). 

Fissidens kegelianus C.Müll. = F.zollingerii (76). 

Fissidens konkourae Par. & Broth. = F. microcarpus 

(73). 

Fissidens lachmanii Biz. IVO(47,1880). 

Fissidens lacouturei Thér. MAD(40), TAN(46), 

UGA(46). 

Fissidens lanceolatus Bruch, nom. nud. = F. glauces- 

cens (1521). 

Fissidens latifolius Dix. CAM(47), KEN(46), 

TAN(46), ZAM(46), ZIM(46). 

Fissidens laurentiorum Card. = F. zippelianus (76). 

Fissidens laxus Sull. & Lesq. CAR(47,1749), 

CON(47), GAB(47), GHA(47), GUI(47), 

IVO(47,1880), LIB(47), MAD(40), NIG(47), 

REU(90), SEY(90), SUD(90), TAN(46), ZAI(65). 

Fissidens le-testui P.Varde GAB(47), IVO(1880), 

ZAI(65). 

Fissidens leptocheilos C.Müll. SOM(50). 

Fissidens leucocaulis C.Müll. ex Biz. CAM(47), 

CAR(47), GAB(47), MAL(67), NIG(47), 

SAO(47), TAN(46). 

Fissidens leucocinctus Hampe CAM(47), 

KEN(46), MAD(40), SAF(40), TAN(46), 

UGA(40), ZAI(65). 

Fissidens ligulinus C.Müll. MAD(40). 

Fissidens linderi P.Varde in Thér. = F. laxus (90). 

Fissidens linearilimbatus C.Müll. KEN(46), 

TAN(46). 

Fissidens loennbergii P.Varde KEN(46). 

Fissidens longelimbatus Broth.. = F. intramarginatus 

(90) 

Fissidens longevittatus P.Varde GAB(47), 

NIG(47), TAN(46). 

Fissidens longidens Biz. ex Pócs TAN(46). 

Fissidens longipes Welw. & Duby. = F. bryoides (90). 

Fissidens loyei Biz. CON(47). 

Fissidens luridus Ren. & Card. MAD(40). 

Fissidens maclaudii Par. & Broth. GUI(47). 

Fissidens macowanianus C.Müll. = F. rufescens (82). 

Fissidens macroglossus Broth. CAM(47), 

IVO(1458). 

Fissidens madecassus C.Müll. MAD(40). 

Fissidens mammillosus C.Müll. = F. zippelianus (76). 

Fissidens maniae Par. & Ren. = F. zippelianus (76). 

Fissidens marginatus Schimp. ex C.Müll. 

CBS(1531), TAN(46). 

Fissidens mariei (Besch.) Broth. MAD(40). 

Fissidens marthae Card. CAR(47,1749), GHA(47), 

IVO(1880), NIG(47), SEN(47), TAN(46). 

Fissidens mathieui Card. CAR(47,1749), SEN(47). 

Fissidens menyhartii C.Müll. ZAM(46,62), 

ZIM(46). 

Fissidens metzgeria (C.Müll.) Broth. TAN(46). 

Fissidens microandrogynus Dix. BOT(1531), 

LES(1531), NAT(1531), OFS(1531), TAN(46), 

TRV(1531), ZAM(46,62), ZIM(46,61). 

Fissidens microcarpus Mitt. CAM(47,73), 

CAR(47), CON(73), GAB(73), GUI(47,73), 

NIG(47), SIL(47), ZAI(65). 

Fissidens microcellatus Dix. ZAI(65). 

Fissidens minutifolius Broth. & P.Varde 

CAR(1749), GAB(47). 

Fissidens minutus Thwaites & Mitt. in Mitt. 

CAR(1470), TAN(1470), ZAM(1470). 

Fissidens mobukensis Negri UGA(46). 

Fissidens moenkemeyeri C.Müll.= F. zippelianus (76). 

Fissidens molliculus P.Varde CAR(47), ZAI(65). 

Fissidens monostictus Broth. & P.Varde = F. 

kegelianus (74) (= F. zollingeri). 

Fissidens motelayi Ren. & Card. MAD(40). 

Fissidens nigerianus Biz. NIG(47,334). 

Fissidens nitens Rehm. ex Salm. = F. porrectus (90). 

Fissidens norkettii P.Varde = F. borgenii (90). 

Fissidens nossianus Besch. MAD(40). 

Fissidens oblongifolius Hook.f. & Wils. CAR(47), 

GAB(47).

Fissidens obsoletidens C.Müll. COM(40), 

MAD(40), TAN(46). 

Fissidens obsoletidens var. schistophilla Broth. 

RWA(65). 

Fissidens ocellatus P.Varde GAB(47). 

Fissidens oligistoloma P.Varde CAR(1749). 

Fissidens onraedtii Biz. MAD(40). 

Fissidens opacifolius Mitt. NIG(47), TAN(46), 

ZAI(65). 

Fissidens oussouyensis Card. CAR(1749), 

SEN(47). 

Fissidens ovatus fo. cymatophyllus (C.Müll.) Bix. ex 

Kis. Excluded. Any records for this form have 

been associated with the owning species. 

Fissidens ovatus var. elatior Ren. & Card. 

MAS(40). 

Fissidens ovatus var. ochroloma Ren. et Card. 

MAD(40), MAL(67), NIG(334), TAN(46). 

Fissidens ovatus var. ovatus Brid. MAD(40), 

MAU(105), REU(105), TAN(46), ZAM(46). 

Fissidens ovatus var. planifolius Besch. 

MAD(40). 

Fissidens pachyloma C.Müll. ETH(50). 

Fissidens pachylomadelphus Demar. & P.Varde = F. 

bryoides (90). 

Fissidens pachylomoides P.Varde = F. porrectus 

(90). 

Fissidens pachylomoides var. subdenticulatus P.Varde 

= F. porrectus (90). 

Fissidens palmifolius (P.Beauv.) Broth. ANG(46), 

CBS(1531), MAL(67), MAU(105), NAT(1531), 

OFS(1531), REU(105), TRV(1531), ZAM(46,62), 

ZIM(46,61). 

Fissidens palustris (C.Müll.) Par. CAM(47), 

CAR(47,1749), LIB(47). 

Fissidens palvadeaui Biz. = F. laxus (90). 

Fissidens papillosus Sande Lac KEN(580,46), 

MAD(40), MAU(580,105), TAN(580,46). 

Fissidens parkii Mitt. CAR(47), CHA(47), GUI(47), 

IVO(1880), MAI(47), NIG(47), SEN(47), 

TAN(46). 

Fissidens parvilimbatus Sim CBS(1531), 

LES(1531), NAT(1531), TAN(46), TRV(1531). 

Fissidens paucilimbatus P.Varde = F. borgenii (90). 

Fissidens pechuelii (C.Müll.) Jaeg. CAR(47), 

TAN(46), ZAI(65). 

Fissidens perfoliatus (C.Müll.) Par. CAR(1749), 

ZAI(65). 

Fissidens perintegrifolius P.Varde CAR(1749), 

GAB(47), IVO(1880), NIG(47), SIL(47). 

Fissidens perlimbatus Broth. & P.Varde CAR(47), 

GAB(47), IVO(1880), NIG(47). 

Fissidens perlimbatus fo. albinervis P.Varde. Exclu- 

ded. Any records for this form have been 


Fissidens perparviretis Dix. CAR(47), KEN(46), 

ZAI(65). 

Fissidens pictus Biz. ex Pócs = F. laxus (90). 

Fissidens pierrotii Biz. CAR(47,1749), TAN(46). 

Fissidens planifrons Besch. COM(40), MAD(40), 

TAN(46). 

Fissidens planifrons var. corticeus Besch. 

COM(40), MAD(40). 

Fissidens platybryoides C.Müll. = F. bryoides (65) 

Fissidens platybryoides var. subimmarginatus Dix. = 

F. bryoides (90). 

Fissidens platyneuros Ren. & Card. MAD(40). 

Fissidens plumosus Hornsch. CBS(1531), 

MAL(67), MAU(105), NAT(1531), REU(105), 

TAN(46), TRV(1531), ZIM(46,61). 

Fissidens pobeguinii Par. & Broth. CAM(47), 

CAR(47), GUI(47), NIG(47), SIL(47), TOG(1878), 

ZAI(65). 

Fissidens pocsii Biz. & Dury ex Pócs TAN(46). 

Fissidens porrectus Mitt. CAM(47), GAB(47), 

GUI(47), IVO(1880), NAT(1531), NIG(47), 

RWA(65), TAN(46), UGA(46), ZAI(65), 

ZIM(61). 

Fissidens procumbens Mitt. ROD(105). 

Fissidens propinquus P.Varde = F. laxus (90). 

Fissidens pseudoeeni Biz. & Dury ex Pócs 

TAN(46). 

Fissidens pseudolimbatus C.Müll. ex De Vent. 

ETH(50). 

Fissidens pseudoplumosus Biz. & Onr. REU(105). 

Fissidens pseudorufescens C.Müll. KEN(46), 

TAN(46). 

Fissidens pseudoserratus (C.Müll.) Jaeg. 

CBS(1531), MOZ(357), NAT(1531), TRV(1531), 

ZIM(61). 

Fissidens pulcher C.Müll. ex Dusen. = F. zippelianus 

(76). 

Fissidens purpureocaulis C.Müll. CAM(47), 

MAD(90), REU(90), RWA(65), SAO(47), 

TAN(46), ZAI(65). 

Fissidens pygmaeus Hornsch. CBS(1531), 

ZIM(46,61). 

Fissidens ramulosus Mitt. CAR(47), NIG(47). 

Fissidens recurvatifolius C.Müll. ex Par., nom. 

nud. ETH(50). 

127

128 

Fissidens reflexus Hampe MAD(40), TAN(46), 

ZAM(62). 

Fissidens ripensis Mitt. = F. laxus (90). 

Fissidens rivicola Broth. & P.Varde = F. laxus (90). 

Fissidens rivicola var. gabonensis P.Varde = F. laxus 

(90). 

Fissidens robynsianus P.Varde ZAI(65). 

Fissidens rosulatus P.Varde MAD(40). 

Fissidens rotereaui P.Varde MAD(40). 

Fissidens rotundatus Dix. CAR(47,1749), GUI(47), 

IVO(63), TOG(1878). 

Fissidens rufescens Hornsch. BOT(1531), 

CBS(1531), DJI(1569), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), TAN(46), TRV(1531), 

ZIM(61). 

Fissidens rufolimbatus Card. = F. porrectus (90). 

Fissidens rufolimbatus var. latifolius P.Varde = F. 

porrectus (90). 

Fissidens rufolimbatus var. ramosus P.Varde = F. 


Fissidens rufolimbatus var. torrentium P.Varde = F. 


Fissidens ruwenzorensis Thér. & Nav. = F. 

asplenioides (1748). 

Fissidens sarcophyllus C.Müll. ex Dus. = F. borgenii 

(90). 

Fissidens savellii Par. & Ren. = F. zippelianus (76). 

Fissidens schelpei P.Varde KEN(46). 

Fissidens schmidii C.Müll. CAM(47), CAR(47), 

GAB(47), GUI(47), NIG(47), UGA(46). 

Fissidens schnellii P.Varde = F. porrectus (90). 

Fissidens schweinfurthii (C.Müll.) Par. 

CAR(47,1749), TAN(46). 

Fissidens sciophyllus Mitt. CHA(90), GAB(47), 

GHA(90), MAD(90), NIG(334), RWA(65), 

SUD(90), TAN(46), UGA(46), ZAI(65), 

ZAM(46,62). 

Fissidens scleromitrius (Besch.) Broth. CAR(47), 

MAD(40), NAT(1531), ZAI(65). 

Fissidens semi-obscurus (C.Müll.) Par. 

CAR(47,1749), CHA(47), IVO(1880), NIG(47), 

TAN(46), ZAM(46). 

Fissidens sericeus Broth. TAN(46). 

Fissidens seychellensis Dury & Onr. SEY(40). 

Fissidens sigmocarpoides P.Varde CAR(47). 

Fissidens sigmocarpus C.Müll. ex Dus. CAM(47), 

ZAI(65). 

Fissidens simensis Schimp. ex C.Müll. = F. bryoides 

(90). 

Fissidens simii Schelpe in Magill & Schelpe 

CBS(1531), NAT(1531), TAN(46), TRV(1531), 

ZIM(61). 

Fissidens smilioides P.Varde CAR(47,1749), 

TAN(46). 

Fissidens somaliae C.Müll. SOM(50). 

Fissidens spinosolimbatus Biz & Dury ex Pócs = F. 

intramarginatus (90). 

Fissidens splachnifolius Hornsch. CBS(1531). 

Fissidens splendens Brugg.-Nann. TAN(46). 

Fissidens stellenboschianus Dix. in Sim 

CBS(1531), TAN(46). 

Fissidens subambiguus Biz. & Dury ex Pócs 

TAN(46). 

Fissidens subarboreus Broth. & P.Varde = F. oblon- 

gifolius (1748). 

Fissidens subceylonensis Dury & Onr. SEY(40). 

Fissidens subcongolensis Biz. & Dury ex Pócs = F. 

intramarginatus (90). 

Fissidens subdurus Broth. & P.Varde CAR(47). 

Fissidens subelimbatus Broth. & P.Varde 

CAM(47), CAR(47,1749), ZAI(65). 

Fissidens subexasperatus Biz. & Dury ex Pócs = F. 

papillosus (580). 

Fissidens subglaucissimus Broth.= F. glaucissimus 

(255). 

Fissidens submadecassus Biz. ZAI(65). 

Fissidens submarginatus Bruch ex Krauss 

CBS(1531), NAT(1531), SWA(1531), TAN(46), 

TRV(1531), ZAM(62), ZIM(46,61). 

Fissidens subobtusatus C.Müll. CBS(1531), 

DJI(1569), NAM(1531), NAT(1531), OFS(1531), 

TAN(46), TRV(1531), ZAM(46). 

Fissidens subpictus Biz. ex Pócs = F. laxus (90). 

Fissidens subplanifrons Biz. & Onr. MAU(105). 

Fissidens subpycnophyllus P.Varde CAR(47). 

Fissidens subsarcophyllus Biz. & Onr. MAD(40). 

Fissidens subscleromitrius Biz. & Dury TAN(46). 

Fissidens subsphaerocarpus (P.Varde) Brugg.-Nann. 

= F. microcarpus (73). 

Fissidens subundatus Dury MAD(40). 

Fissidens sylvaticus Griff. Iwatsuki & Suzuki (1892, 

p.400) gave this as a synonym of F. taxifolius, but 

Bruggeman-Nannenga (pers. comm.) says that all 

African material named F. sylvaticus seen by her 

is F. zippelianus. All African records are thus 

shown for F. zippelianus. 

Fissidens sylvaticus fo. robusta Dix. Excluded. Any 

records for this form have been associated with 

the owning species. 

Fissidens sylvaticus var savelii (Par. & Ren.) Wijk &

Mar. = F. zippelianus (76). 

Fissidens taeniocladus P.Varde = F. laxus (90). 

Fissidens taxifolius ssp. taxifolius Hedw. 

MAS(40). 

Fissidens tenuisetus Card. = F. platybryoides (1749) 

(= F. bryoides). 

Fissidens termitidarum Par. & Broth. GUI(47), 

TOG(1878). 

Fissidens thallangae Dix. UGA(46). 

Fissidens tisserantii Broth. & P.Varde = F. glauculus 

(90). 

Fissidens tisserantii var. sublaevifolius P.Varde 

CAR(47). 

Fissidens tisserantii var. sublaevifolius P.Varde. The 

owning species of this variety has been transferred 

to F. glauculus (90). This variety should also be 

revised. 

Fissidens ugandae Dix. & P.Varde = F. intramarginatus 

(90) 

Fissidens ulna (C.Müll.) Par. CAR(47,1749), 

GAB(47), GUI(47), IVO(47), NIG(47), TAN(46), 

UGA(46), ZAI(65). 

Fissidens ulna var. fallax P.Varde CAR(47). 

Fissidens undatus (C.Müll.) Par. KEN(46). 

Fissidens undifolius C.Müll. CAR(47,1749), 

KEN(46), TAN(46), ZAI(65). 

Fissidens usambaricus Broth. CAM(90), 

CBS(1531), CAR(90,1749), NAT(1531), 

RWA(65), SAF(90), TAN(46), TRV(1531), 

ZAI(65). 

Fissidens usambaricus var. acutifolius P.Varde 

CAR(47). 

Fissidens vanderijstii Card. CAR(47), GAB(47), 

ZAI(65). 

Fissidens ventroalaris Card. MAD(40), MAL(67), 

TAN(46). 

Fissidens vesiculosus Demar. et Leroy MAL(67), 

TAN(46), ZAI(65). 

Fissidens vogelianus Mitt. CAM(47), CAR(47), 

NIG(47), ROD(105), TAN(46), ZAI(65). 

Fissidens vogelianus var. latifolius P.Varde, nom. 

nud. CAR(47), NIG(47). 

Fissidens wageri Dix. ex Wag. CBS(1531), 

NAT(1531). 

Fissidens waterlotii Thér. MAD(40). 

Fissidens welwitschii Duby ANG(46), TAN(46). 

Fissidens woodii Tayl. & P.Varde = F. porrectus (90). 

Fissidens zenkeri Broth. ex Biz. CAM(47). 

Fissidens zippelianus Dozy & Molk. CAM(47), 

CAR(47,1749), COM(40), EGU(76), ERI(50), 

ETH(50), FER(47), GAB(47), IVO(1458,63), 

KEN(46), MAD(40), MAL(67), MAU(105), 

NIG(47), RWA(46), SIL(47), SUD(76), TAN(46), 

TOG(1878), UGA(46), ZAI(65,1552), ZAM(46), 

ZIM(61). 

Fissidens zollingeri Mont. CAR(47,1749), 

KEN(46), NIG(334,47), TAN(46), ZAI(65). 

Fissidentella perpusilla Card. = Fissidens fouta-djal- 

lonii (76) (= F. gardneri). 

Floribundaria cameruniae (Dus.) C.Müll. = F. flori- 

bunda (65). 

Floribundaria floribunda (Dozy & Molk.) Fleisch. 

CAM(47), CAR(47), CON(47), GAB(47), 

GHA(47), IVO(47,1880), KEN(46), MAD(40), 

MAU(105,1882), NAT(1531), REU(105), 

RWA(65), TAN(46), TRV(1531), UGA(46), 

ZAI(65), ZAM(46). 

Floribundaria floribunda var. capilliramea 

(C.Müll.) Fleisch. TAN(46). 

Floribundaria floribunda var. madagassa (Ren. 

& Card.) Ren. & Card. MAD(40). 

Floribundaria floribundula (C.Müll.) Fleisch. 

COM(40), MAD(40). 

Floribundaria golae Tosco & Piovano TAN(46). 

Floribundaria jumboana (Dus.) C.Müll. 

CAM(47), CAR(47), GAB(47), GUI(47). 

Floribundaria octodiceras (C.Müll.) C.Müll. 

CAM(47). 

Floribundaria patentissima (C.Müll.) Fleisch. 

CAR(47), GAB(47), KEN(411), MAD(40), 

MAU(105), REU(105), SAO(47), TAN(46). 

Floribundaria patentissima var. squarrosa (Ren. 

& Card.) Ren & Card. MAD(40), TAN(46). 

Floribundaria vaginans (Welw. & Dub.) Broth. 

ANG(46), MAL(67). 

Flowersia abyssinica (C.Müll.) Griffin & Buck 

ERI(50), ETH(50). 

Fontinalis abyssinica Schimp. nom. nud. ETH(50). 

Fontinalis antipyretica var. erythraea Tong 

ERI(50). 

Fontinalis antipyretica var. gracilis (Hedw.) 

Schimp. CBS(1531). 

Fontinalis fiorii Tong ERI(50). 

Fontinalis squamosa var. squamosa Hedw. 

CBS(1531). 

Forsstroemia dixonii Tosco & Piovano = Levierella 

fabroniacea (408). 

Forsstroemia flagellacea (C.Müll.) Par. = F. producta 

(408). 

Forsstroemia incrassata (Broth.) Par. = F. producta 

129

130 

(408). 

Forsstroemia producta (Hornsch.) Par. 

CBS(1531,408), ETH(408), KEN(46,408), 

MAL(67), NAT(1531,408), RWA(65), 

TAN(46,408), TRV(1531,408), UGA(46,408). 

Forsstroemia producta var. brachycystis Thér. & 

P.Varde. = F. producta (408). 

Funaria abyssinica Schimp. in Par., nom. nud. 

ETH(50). 

Funaria acicularis C.Müll. SAO(47). 

Funaria bergiana (Hornsch.) Broth. See Entosthodon 

bergianus. 

Funaria bogosica C.Müll. ERI(50). 

Funaria borbonica (Besch.) Broth. See Entosthodon 

borbonicus. 

Funaria cameruniae Dix. CAM(47). 

Funaria clavata (Mitt.) Magill See Entosthodon 

clavatus. 

Funaria convexa Spruce TAN(46). 

Funaria curvipes (C.Müll.) Broth. See Entosthodon 

curvipes. 

Funaria decaryi Thér. MAD(40). 

Funaria delicatula Thér. MAD(40). 

Funaria hygrometrica fo. breviseta (Broth.) Cuf. 


associated with Funaria hygrometrica var. cal- 

vescens. 

Funaria hygrometrica var. calvescens (Schwaegr.) 

Mont. BUR(65), CAM(47), CAR(47), COM(40), 

ERI(50), ETH(50), GUI(47), IVO(1458), 

MAD(40), MAL(67), MAU(105), REU(105), 

RWA(65), TAN(46), UGA(46), ZAI(65), 

ZAM(46). 

Funaria hygrometrica var. hygrometrica Hedw. 

CAM(47), CBS(1531), ETH(50), KEN(46), 

LES(1531), MAL(67), NAM(1531), NAT(1531), 

OFS(1531), REU(105), SAO(47), SUD(1707), 

SWA(1531), TAN(46), TRV(1531), ZIM(46,61). 

Funaria hygrometrica var. mauritiana 

(Besch.)Par. MAU(105), REU(105). 

Funaria imerinensis Card. MAD(40). 

Funaria kilimandscharica C.Müll. KEN(46), 

TAN(46), ZAI(65). 

Funaria lepervanchei (Besch.) Broth. See 

Entosthodon lepervanchei. 

Funaria limbata (C.Müll.) Broth. See Entosthodon 

limbatus. 

Funaria longicollis Dix. = Entosthodon dixonii (46) 

Funaria marginata (C.Müll.) Broth., hom. illeg. = 

Entosthodon ampliretis (C.Müll.) Broth (1521) 

= Funaria limbata (C.Müll.) Broth (83) ( = 

Entosthodon limbatus). 

Funaria marginatula (Besch.) Broth. See 

Entosthodon marginatulus. 

Funaria mauritiana (Besch.) Broth. See Entosthodon 

mauritianus. 

Funaria mayottensis (Besch.) Broth. COM(40). 

Funaria nutans Mitt. SUD(1707). 

Funaria pallescens Schimp. in Jaeg., nom. nud. 

ETH(50). 

Funaria perlaxa Thér. KEN(46), TAN(46), 

ZAI(65). 

Funaria renauldii (Thér.) Card. MAD(40). 

Funaria rhomboidea Shaw CBS(1531), 

NAM(1531), OFS(1531), TRV(1531), ZIM(61). 

Funaria robustior (C.Müll.) Broth. See Entosthodon 

curvipes var. robustior. 

Funaria rottleri (Schwaegr. )Broth. See Entosthodon 

rottleri. 

Funaria spathulata Schimp. ex C.Müll. CBS(1531), 

LES(1531), NAT(1531), ZIM(61). 

Funaria subleptopoda Hampe MAD(40). 

Funaria succuleata (Wager & Wright) Magill 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TRV(1531), ZIM(61). 

Funaria urceolata (Mitt.) Magill. See Entosthodon 

urceolatus. 

Funaria volkensii Broth. See Entosthodon volkensii. 

Gammiella merrillii (Broth.) Tix. CON(47), 

GAB(47), TAN(46). 

Garckea comosa (Dozy & Molk.) Wijk & Marg. = G. 

phascoides (40). 

Garckea elongata Ren. & Card. ZAI(65). 

Garckea mathieui Thér. CAM(47), CAR(47), 

GAB(47), SIL(47). 

Garckea moenkemeyeri C.Müll. CAR(47), 

GAB(47), GUI(47,1458), IVO(1458), NIG(47), 

SIL(47). 

Garckea phascoides C.Müll. COM(40), MAD(40), 

REU(105), TAN(46), ZAI(65). 

Garckea phascoides var. comorensis Thér. 

COM(40). 

Gigaspermum repens (Hook.) Lindb. CBS(1531), 

LES(1531), MAD(40), NAM(1531), OFS(1531). 

Glossadelphus abruptus (Mitt.) Robins. 

IVO(1880). 

Glossadelphus congolensis Broth. & P.Varde 

CAR(47), GAB(47), TOG(1878), ZAI(65,1552). 

Glossadelphus congolensis var. falcatulus Broth. 

ZAI(65).

Glossadelphus eckendorffii P.Varde GAB(47). 

Glossadelphus euryphyllus P.Varde SIL(47). 

Glossadelphus ovalifolius (Besch.) Broth. 

MAD(40). 

Glossadelphus perplanicaulis (Broth.) Fleisch. 

CAM(47), CAR(47). 

Glossadelphus scutellifolius (Besch.) Fleisch. 

MAD(40). 

Glossadelphus serpyllifolius P.Varde UGA(46), 

ZAI(65). 

Glossadelphus truncatulus (C.Müll.) Fleisch. 

TOG(1878). 

Glossadelphus truncatus (Welw. & Duby) Fleisch. 

ANG(46), CAM(47), CAR(47), GAB(47), 

ZAI(65). 

Glyphomitrium helenicum Mitt. = Ptychomitrium 

helenicum (1521). 

Gollania bartramiophylla (C.Müll.) Broth. ex 

Par. TAN(46). 

Gollania demaretii P.Varde = Chryso-hypnum dema- 

retii (65). 

Gollania le-testui Dix. & P.Varde GAB(47). 

Gollania monodii P.Varde SAO(47). 

Goniomitrium africanum (C.Müll.) Broth. 

CBS(1531), LES(1531), NAM(1531), OFS(1531), 

TRV(1531). 

Goniomitrium speluncae P.Varde CAR(47). 

Grimmia abyssinica (C.Müll.) Mitt. = G. apiculata 

(46). 

Grimmia aethiopica C.Müll. ERI(50), ETH(50). 

Grimmia affinis Hornsch. KEN(46), TAN(46), 

UGA(46), ZIM(61). 

Grimmia afro-incurva Broth. TAN(46), UGA(46), 

ZAI(65). 

Grimmia apiculata Hornsch. CAM(47), ETH(50), 

KEN(46), TAN(46). 

Grimmia argyrotricha C.Müll. TAN(46). 

Grimmia campylotricha C.Müll. TAN(46). 

Grimmia conferta Funck = Schistidium apocarpum 

var. confertum (1521). 

Grimmia donniana var. donniana Sm. TAN(146). 

Grimmia erythraea (C.Müll.) Cuf. ERI(50). 

Grimmia immergens C.Müll. TAN(46). 

Grimmia laevigata (Brid.) Brid. CBS(1531), 

KEN(46), LES(1531), NAT(1531), OFS(1531), 

REU(105), SUD(1419), TAN(46), UGA(46). 

Grimmia longicaulis Dix. KEN(46). 

Grimmia madagassa Ren. & Card. MAD(40). 

Grimmia mammosa Gao & Cao MAL(66). 

Grimmia obtusolinearis C.Müll. TAN(46). 

Grimmia ovalis (Hedw.) Lindb. CBS(1531), 

LES(1531), NAT(1531), OFS(1531). 

Grimmia ovalis (auct. afr., non (Hedw.) Lindb.). 

According to Kis, African specimens with this 

name should be referred to G. affinis (46). The 

above distribution has been confirmed with Dr. R. 

Ochyra. 

Grimmia pulvinata var. africana (Hedw.) Hook.f. 

ETH(50). 

Grimmia pulvinata var. pulvinata (Hedw.) Sm. 

CBS(1531), ETH(50), LES(1531), NAM(1531), 

NAT(1531), OFS(1531). 

Grimmia schimperi B&S ex C.Müll., nom. nud. 

ETH(50). 

Grimmia subleucophaea C.Müll. ETH(50). 

Grimmia subleucophaea C.Müll. According to 

Ochyra (pers. comm.), the type of this taxon is 

indistinguishable from G. laevigata. 

Grimmia trichophylla var. brachycarpa De Not. 

KEN(46), TAN(46). 

Grimmia trichophylla var. trichophylla Grev. 

RWA(65). 

Grimmia vulcanica Besch. MAD(40), REU(105). 

Groutiella elimbata (Thér.) O’Shea MAD(40). 

Groutiella laxotorquata (Besch.) Wijk. & Marg. 

CAM(47), CAR(47), COM(40), GAB(47), 

GHA(47), GUI(47), IVO(44,1880), MAD(40), 

MAU(105), NIG(47), SIL(47), TAN(46), 

TOG(58,1878), ZAM(62,411). 

Groutiella sarcotricha (Broth.) Wijk & Marg. 

NIG(334), ZAI(65). 

Groutiella thraustophylla (Broth.) Wijk & Marg. 

CAM(47). 

Gymnostomiella tanganyikae De Sloover 

BUR(65). 

Gymnostomum aeruginosum Sm. CBS(1531), 

LES(1531), NAT(1531), OFS(1531), TAN(46), 

TRV(1531). 

Gymnostomum bescherellei Broth. & Geh. 

ASC(1296). 

Gymnostomum bewsii Dix. CBS(1531), LES(1531), 

NAT(1531), OFS(1531), TRV(1531). 

Gymnostomum calcareum Nees & Hornsch. 

MAL(67). 

Gymnostomum kenyae P.Varde = Anoectangium 

keniae (1420). 

Gymnostomum leptostomum Brid., hom. illeg. 

MAD(40). 

Gymnostomum lessonii Besch. Excluded. According 

to Zander (1420), this is a species of Racopilum. 

131

132 

Gymnostomum lingulatum Rehm. ex Sim 

TRV(1531). 

Gymnostomum splachnobryoides Biz. ZAI(65). 

Gyroweisia hildebrandtii (C.Müll.) Broth. = Hyme- 

nostylium hildebrandtii (1420). 

Gyroweisia latifolia var. tanneri Biz. ex Pócs = 

Husnotiella latifolia var. tanneri (46). 

Gyroweisia pocsii Biz. = Weisiopsis nigeriana (1420). 

Gyroweisia tophicola (C.Müll.) Broth. = Hymenosty- 

lium recurvirostrum (1420). 

Haplocladium angustifolium (Hampe & C.Müll.) 

Broth. CBS(1531), KEN(411), NAT(1531), 

TAN(46), TRV(1531), UGA(46), ZAM(46), 

ZIM(46,61). 

Haplocladium angustifolium var. afro-capillatum 

(Broth.) Thér. ZAI(65). 

Haplocladium angustifolium var. viride Broth. & 

Bryhn ZAI(65). 

Haplocladium jacquemontii (C.Müll.) Broth. 

ETH(50). 

Haplodontium clavatum (Bruch & Schimp.) Broth. 

= Mielichhoferia clavata (1532). 

Haplodontium nitidum P.Varde = Mielichhoferia 

vardei (1532). 

Haplodontium ovale (Mitt.) Broth. = Mielichhoferia 

ovalis (1532). 

Haplodontium pusillum Card. = Mielichhoferia 

cardotii (1532). 

Haplohymenium pseudotriste (C.Müll.) Broth. 

CBS(1531), MAU(105), NAT(1531), ROD(105), 

TAN(46), TRV(1531). 

Haplohymenium pseudotriste var. robustum Biz. 

MAU(105). 

Haplohymenium tenuissimum (Besch.) Broth. 

MAU(105). 

Haplohymenium triste (Ces.) Kindb. KEN(46). 

Hedwigia ciliata fo. leucophaea Jones. See H. ciliata 

var. leucophaea BS&G. 

Hedwigia ciliata var. ciliata (Hedw.) P.Beauv. 

CBS(1531), KEN(46), LES(1531), MAD(40), 

MAL(67), NAT(1531), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZAI(65). 

Hedwigia ciliata var. leucophaea BS&G KEN(46), 

TAN(46), UGA(46). 

Hedwigidium integrifolium (P.Beauv.) Dix. 

CAM(47), CBS(1531), KEN(46), LES(1531), 

NAT(1531), OFS(1531), REU(105), 

RWA(46,65), SWA(1531), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZIM(61). 

Helicodontium fabroniopsis C.Müll. MAD(40). 

Helicodontium guineense Broth. & Par. GUI(47). 

Helicodontium jacobi-felicis P.Varde CAM(47). 

Helicodontium lanceolatum (Hampe & C.Müll.) 

Jaeg. CBS(1531), NAT(1531), TRV(1531). 

Helicodontium usagarum Mitt. TAN(46). 

Heliconema cryptocarpus (Dozy & Molk.) Ellis & 

Eddy, nom. inval. = Chameleion cryptocarpus 

(Dozy & Molk.) Ellis & Eddy in Eddy (755). 

Henicodium geniculatum (Mitt.) Buck CAM(47), 

CAR(47), GAB(47), GHA(47), IVO(44), NIG(47), 

SUD(1339), ZAI(65). 

Hennediella limbata (Mitt.) Zander ETH(50), 

KEN(46). 

Hennediella longipedunculata (C.Müll.) Zander 

CBS(1531), KEN(46), NAM(1531). 

Herpetineuron toccoae (Sull. & Lesq.) Card. 

CAR(47), GUI(47), MAL(67), NAT(1531), 

TAN(46), TRV(1531). 

Herzogiella cylindrocarpa (Card.) Iwats. 

RWA(65). 

Heterophyllium albescens P.Varde GAB(47). 

Heterophyllium albo-alare (Broth.) Broth. 

CAR(47). 

Heterophyllium bulbilliferum Dix. CAM(47). 

Heterophyllium flexile (Ren. & Card.) Thér. & P.Varde 

= Rhacopilopsis transvaaliensis (717). 

Heterophyllium guineense (Broth. & Par.) Broth. = 

Rhacopilopsis guineensis (717). 

Heterophyllium humbertii P.Varde ZAI(65). 

Heterophyllium julicaule (Broh. & Par.) Broth. = 


Heterophyllium julicaule var. auriculatum P.Varde = 


Heterophyllium kilimandscharicum (Broth. & 

P.Varde) Thér. & P.Varde (= “kilimanjaricum”) 

= Rhacopilopsis kilimandscharica (717). 

Heterophyllium scabrifolium J.Tayl. & P.Varde 

RWA(65), SIL(47). 

Heterophyllium transvaaliense (Thér. & Dix.) Thér. 

& P.Varde. = Rhacopilopsis transvaaliensis (717). 

Heterophyllium variegatum (Welw. & Duby) 

Broth. ANG(46), GUI(47). 

Hildebrandtiella cuspidans Besch. COM(40). 

Hildebrandtiella endotrichelloides C.Müll. 

COM(40), MAD(40). 

Hildebrandtiella longiseta Ren. & Card. 

MAD(40). 

Hildebrandtiella nitens Bosw., nom. nud. 

MAU(105). 

Hildebrandtiella pachyclada Besch.

MAU(2,105), REU(2,105), ROD(105), 

TAN(46,2). 

Hildebrandtiella rotundifolia Ren. MAD(40), 

REU(105). 

Hildebrandtiella rotundifolia Ren. This taxon is 

treated by Argent (2) as a variety of H. 

endotrichelloides, but he does not make the name 

change. 

Himantocladium ascensionis (Besch.) Par. = H. im- 

planum (302). 

Himantocladium cyclophyllum (C.Müll.) Fleisch. 

SEY(40). 

Himantocladium implanum (Mitt.) Fleisch. 

ASC(1296). 

Himantocladium seychellarum Dix. = H. 

cyclophyllum (1300). 

Holomitrium angolense P.Varde ANG(46), 

UGA(46), ZAM(411). 

Holomitrium angolense fo. brevifolia P.Varde. Ex- 



Holomitrium borbonicum Besch. MAD(40), 

MAU(105), REU(105). 

Holomitrium borbonicum fo. longisetum Besch. Ex- 



Holomitrium borbonicum var. hamatum (C.Müll.) 

Thér. MAD(40), REU(105), TAN(46). 

Holomitrium calycinum (Hedw.) Mitt. REU(105). 

Holomitrium camptocarpum P.Varde MAD(40). 

Holomitrium comorense C.Müll. COM(40). 

Holomitrium cylindraceum var. cucullatum 

(Besch.) Wijk & Marg. CBS(1531), CAR(47), 

MAD(40), NAT(1531), REU(105), TAN(46), 

TRV(1531), ZIM(46). 

Holomitrium cylindraceum var. cylindraceum 

(P.Beauv.) Wijk & Marg. CAR(47), MAD(40), 

MAU(105), REU(105). 

Holomitrium cylindraceum var. obtusifolium 

(Besch.) Wijk & Marg. MAD(40), MAU(105), 

UGA(46). 

Holomitrium dubiosum Dix. ANG(46), ZAI(65). 

Holomitrium gracilisetum Thér. MAD(40). 

Holomitrium hillieri Thér. MAD(40). 

Holomitrium lepervanchei Thér. MAD(40), 

MAS(40). 

Holomitrium rhyparodictyon P.Varde MAD(40). 

Holomitrium subglobosum Thér. MAD(40). 

Homalia pygmaea Broth. = Symphyodon pygmaeus 

(924). 

Homalia subexigua C.Müll., nom. nud. MAU(105). 

Homaliodendron exiguum (Bosch. & Lac.) 

Fleisch. MAD(40), MAU(105), REU(105), 

TAN(46). 

Homaliodendron exiguum var. semimarginatum 

(Thér.) W.Sch.-Mot. Excluded. Pócs (108) 

states that the type is a mixture of two plants, 

neither new, and declares this name nomina 

reiicienda. 

Homaliodendron flabellatum (Sm.) Fleisch. 

MAU(1629). 

Homaliodendron piniforme (Brid.) Enroth 

CAM(785,47), COM(40), CON(785,47), 

GAB(47), KEN(785), LIB(785), MAD(785,40), 

MAS(785), MAU(105), NIG(785,47), 

TAN(785,46), UGA(785), ZAI(65). 

Homaliodendron scapellifolium (Mitt.) Fleisch. = H. 

flabellatum (1386). 

Homalothecium afrostriatum (C.Müll.) Ochyra = 

Palamocladium sericeum (717). 

Homalothecium mandoni Mitt. CAP(1296). 

Homalothecium nilgheriense (Mont.) Robins. = Pa- 

lamocladium nilgheriense (1608). 

Homalothecium sericeum BS&G. See Palamocladium 

sericeum. 

Hookeria contracta Gepp ANG(46). 

Hookeria splachnifolia (Brid.) Arnott REU(105). 

Hookeriopsis ambigua P.Varde BUR(65), 

GAB(47), RWA(65), ZAI(65). 

Hookeriopsis angolensis (Welw. & Duby) Par. 

ANG(46). 

Hookeriopsis balazsii Biz. CAM(47). 

Hookeriopsis cheiloneura (Broth.) Broth. 

CAM(47). 

Hookeriopsis darntyi (Besch.) Broth. MAU(105). 

Hookeriopsis diversifolia (Ren. & Card.) Broth. 

ex Card. COM(40), MAD(40). 

Hookeriopsis gabonensis Broth. & P.Varde 

GAB(47). 

Hookeriopsis mittenii P.Varde CAM(47), FER(47), 

GAB(47), KEN(46), TAN(46), UGA(46), 

ZIM(61). 

Hookeriopsis pallidifolia (Mitt.) Geh. & Herz. 

STH(1296). 

Hookeriopsis papillosula Broth. & P.Varde 

GAB(47). 

Hookeriopsis pappeana (Hampe) Jaeg. 

CBS(1531), KEN(411), NAT(1531), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65). 

Hookeriopsis pocsii Biz. TAN(46). 

133

134 

Hookeriopsis staudtii (Broth.) Broth. CAM(47). 

Husnotiella latifolia (Dix.) Zander & Magill = Didy- 

modon tophaceopsis (1420). 

Husnotiella latifolia var. tanneri (Biz. & Pócs) 

Kis TAN(46). 

Husnotiella latifolia var. tanneri (Biz. ex Pócs) Kis. 

Husnotiella latifolia has been made a synonym of 

Didymodon tophaceopsis by Zander (1420), but 

he did not deal with this variety. If it is distinct, 

it also should be transferred. 

Husnotiella plicata Magill = Didymodon jackvancei 

(1420). 

Hydrogonium afro-fontanum C.Müll. = Barbula eh- 

renbergii (1158). 

Hydrogonium brotheri Ren. & Par. = Barbula ehren- 

bergii (1158). 

Hydrogonium clavicostatum (Ren. & Card.) Hilp. = 

Barbula indica (1158). 

Hydrogonium dorrii (Ren. & Card.) Hilp. = Barbula 


Hydrogonium ehrenbergii (Lor.) Jaeg. = Barbula 

ehrenbergii (46). 

Hydrogonium fontanum (C.Müll.) Jaeg. = Barbula 

meidensis (1420). 

Hydrogonium tisserantii (P.Varde) Sch.-Mot. = Bar- 

bula tisserantii (1420). 

Hygroamblystegium caudicaule (C.Müll.) Broth. = 

Pseudoleskea chilensis (1533). 

Hygroamblystegium tenax (Hedw.) Jenn. Excluded. 

Tanzanian collections given this name by Potier 

de la Varde have been shown to be Cratoneuron 

filicinum (1533). 

Hygrohypnum caussequei (Ren. & Card.) Card. 

MAD(40). 

Hygrohypnum hedbergii P.Varde = Platyhypnidium 

hedbergii (65). 

Hylocomiopsis cylindricarpa Thér. BUR(65), 

CAM(32,47), KEN(46), RWA(65), TAN(46), 

ZAI(65). 

Hylocomium brevirostre BS&G REU(105). 

Hylocomium proliferum L. = H. splendens (1521). 

Hylocomium splendens (Hedw.) BS&G KEN(46), 

RWA(65), STH(1296), TAN(46), UGA(46), 

ZAI(65). 

Hylocomium splendens var. alaskanum (Lesq. & 

Jam.) Limpr. TAN(1410). 

Hylocomium splendens var. alpinum Schlieph. 

ZAI(65). 

Hymenostomum abyssinicum Schimp. ex Par., 

nom. nud. ETH(50). 

Hymenostomum abyssinicum Schimp. ex Par. nom. 

nud. Hymenostomum R.Brown is now a synonym 

of Weissia Hedw. (1420) but this taxon is not 

covered by Zander (1420) or Index Muscorum 

(1521). Cufodontis (40) states that a specimen 

exists in BM, but I could not find one. Revision 

is required. 

Hymenostomum ayresii (Schimp.) Broth. = Weissia 

ayresii (1420). 

Hymenostomum brachypelma (C.Müll.) Kindb. = 

Weissia brachypelma (1420). 

Hymenostomum chloropus (Besch.) Broth. = Weissia 

ayresii (1420). 

Hymenostomum eckendorfii P.Varde = Tortella ec- 

kendorfii (1420). 

Hymenostomum guineense Broth. & Par. GUI(47). 

Hymenostomum humicolum (C.Müll.) Par. = Weissia 

humicola (1420). 

Hymenostomum lineaefolium (C.Müll.) Par. = Weissia 

lineafolia (1420). 

Hymenostomum pellucidum Broth. in Mich., nom. 

nud. See note under H. abyssinicum regarding 

transfer to Weissia. 

Hymenostomum pulicare Besch. = Trichostomum 

pulicara (1420). 

Hymenostomum socotranum (Mitt.) Broth. = Weissia 

socotrana (1420). 

Hymenostylium congoanum Dix. & Nav. KEN(46). 

Hymenostylium crassinervium Broth. & Dix. 

KEN(46), TAN(46), ZAI(65), ZAM(46), 

ZIM(46). 

Hymenostylium crispulum Broth. & Par. CAR(47), 

GUI(47), TOG(1878). 

Hymenostylium hildebrandtii (C.Müll.) Zander 

SOM(50). 

Hymenostylium recurvirostrum (Hedw.) Dix. 

CBS(1531), KEN(46), LES(1531), NAT(1531), 

SOM(50), TAN(46), TRV(1531), UGA(46). 

Hymenostylium recurvirostrum var. latifolium (Zett.) 

Wijk & Marg. = H. recurvirostrum (1420). 

Hymenostylium rigescens (C.Müll.) Broth. 

DJI(1569), ERI(50), ETH(50). 

Hymenostylium scaturiginosum (C.Müll.) Broth. 

MAU(105), REU(105). 

Hymenostylium secundum C.Müll. = Molendoa sendt- 

nerianum (1420). 

Hymenostylium subcrispulum Thér. MAD(40). 

Hymenostylium xanthocarpum var. gracilescens 

(Hook. ex Mitt.) Cuf. ETH(50). 

Hyophila Brid. It is likely that many taxa in this genus

will be synonymised. Following examination of 

nearly all relevant types, Sollman (pers. comm.) 

believes that there are only two taxa (at species 

level) present in tropical Africa: H. involuta 

(Hook.) Jaeg. and H. nymeniana (Fleisch.) Menzel. 

Hyophila acuminata Broth. & P.Varde IVO(47), 

KEN(46), MAD(40), TAN(46), UGA(46), 

ZAI(65), ZAM(46). 

Hyophila acutiuscula Broth. = Trichostomum acu- 

tiusculum (1420). 

Hyophila angustifolia Par. & Ren. = Trichostomuum 

platyphyllum (1420). 

Hyophila anoectangioides C.Müll. ex Dus. 

LIB(47). 

Hyophila atrovirens (C.Müll.) Broth. = H. involuta 

(46). 

Hyophila atrovirens var. oubanguiensis Thér. & 

P.Varde CAR(47). 

Hyophila baginsensis C.Müll. CAR(47), KEN(46), 

MOZ(357), NAM(1531), NAT(1531), 

TRV(1531), ZAM(62), ZIM(46,61). 

Hyophila bingeri Broth. & Par. GUI(47), IVO(47), 

ZAI(65). 

Hyophila calymperoides Thér. & Nav. = Bryoeryth- 

rophyllum campylocarpum (1420). 

Hyophila compacta (Welw. & Duby) Jaeg. = Tricho- 

stomum brachydontium (1158) (as Pottia 

compacta Welw. & Duby). 

Hyophila congolensis Thér. & Nav. ZAI(65). 

Hyophila crenulata C.Müll. ex Dus. = H. involuta 

(1420). 

Hyophila cuspidatissima Par. & Broth. GUI(47). 

Hyophila excurrentinervis Par. & Broth. = Trichosto- 

mum brachydontium (1158). 

Hyophila fouta-djalloni Par. & Broth. GUI(47), 

IVO(47). 

Hyophila girodii Ren. & Card. = Barbula indica 

(1158). 

Hyophila glaucoviridis Par. & Broth. CAR(47), 

GUI(47), TOG(1878). 

Hyophila gymnostomioides (Welw. & Duby) Jaeg. = 

Trichostomum brachydontium (1158) (as Pottia 

gymnostomioides Welw. & Duby). 

Hyophila holstii Broth. TAN(46). 

Hyophila integrifolia Dix. & Thér. ZAI(65). 

Hyophila involuta (Hook.) Jaeg. CAR(47), 

GUI(1458), IVO(1458,1880), KEN(46), 

LES(1531), MAL(67), MOZ(46), NAT(1531), 

TAN(46), TOG(1878), TRV(1531), ZAI(1552,65), 

ZAM(62), ZIM(46,61). 

Hyophila kerensis Broth. in Mich., nom. nud. = 

Trichostomum brachydontium (1158). 

Hyophila lanceolata Ren. & Card. MAD(40), 

SEY(40). 

Hyophila latifolia Broth. CON(47). 

Hyophila leikipiae (C.Müll.) Broth. = Trichostomum 

brachydontium (1158) (as Trichostomum 

leikipiae C.Müll.). 

Hyophila leioneura Ren. & Par. = Barbula indica 

(1420). 

Hyophila ligulaefolia Broth. & Par. = Trichostomum 

ligulaefolium (1420). 

Hyophila niam-niamiae C.Müll. TAN(46). 

Hyophila nymaniana (Fleisch.) Menzel BUF(47), 

CAR(47), SIL(47), ZAI(1460). 

Hyophila parietalis Card. MAD(40). 

Hyophila perpendiculata Dix. ZAI(65). 

Hyophila potieri Besch. MAD(40), REU(105), 

TAN(46). 

Hyophila potieri var. atroviridis Ren. & Card. 

MAD(40). 

Hyophila potieri var. denticulata Broth. TAN(46). 

Hyophila procera Par. & Broth. GUI(47). 

Hyophila punctulata (Mitt.) Kindb. SOC(40). 

Hyophila sakalavensis Par. & Ren. COM(40), 

MAD(40). 

Hyophila somaliae C.Müll. ex Dus. = Tortella somaliae 

(1420). 

Hyophila subacutiuscula P.Varde & Thér. 

RWA(65). 

Hyophila tisserantii P.Varde = H. nymaniana (1460). 

Hyophila usambarica Broth. = Trichostomum bra- 

chydontium (1158). 

Hyophila victoriae C.Müll. ex Dus. CAM(47), 

CAR(47), SIL(47). 

Hypnella abrupta (Mitt.) Jaeg. CAM(47), FER(47). 

Hypnella guineensis Broth. & Par. GUI(47). 

Hypnella semi-scabra Ren. & Card. MAD(40). 

Hypnella viridis Ren. & Card. MAD(40). 

Hypnum aduncoides (Brid.) C.Müll. = H. macrogynum 

(1913). 

Hypnum africanum (P.Varde) Ochyra RWA(46), 

TAN(46), UGA(46), ZAI(65). 

Hypnum bicolor Bel. REU(105). 

Hypnum boryanum Schwaegr. REU(105). 

Hypnum cupressiforme fo. filiforme Krahmer. See H. 

cupressiforme var. filiforme Brid. 

Hypnum cupressiforme var. cupressiforme Hedw. 

ANG(169), CBS(1531,169,1882), ETH(169), 

KEN(46,169,136), LES(1531,169), 

135

136 

NAT(1531,169), OFS(1531), RWA(46,65), 

SWA(1531,169), TAN(46,169), TRV(1531,169), 

UGA(46,169), ZAI(65), ZIM(61). 

Hypnum cupressiforme var. ericetorum BS&G. = H. 

jutlandicum (162). 

Hypnum cupressiforme var. filiforme Brid. ZAI(65). 

Hypnum cupressiforme var. integrifolium Thér. = H. 

cupressiforme var. cupressiforme (169). 

Hypnum cupressiforme var. lacunosum Brid. 

CBS(169), ETH(169), KEN(169), LES(169), 

NAT(169), RWA(65), SWA(169), TRV(169). 

Hypnum cupressiforme var. mossmanianum 

(C.Müll.) Ando ZAI(65). 

Hypnum cupressiforme var. resupinatum (Tayl.) 

Schimp. RWA(65), ZAI(65). 

Hypnum cupressiforme var. townsendii Ando 

KEN(679), RWA(65), ZAI(65). 

Hypnum ericetorum BS&G = H. cupressiforme var. 

ericetorum (1521) (= H. jutlandicum). 

Hypnum gracilirameum C.Müll. ex Ren. 

MAU(105). 

Hypnum jutlandicum Holmen & Warncke 

CAR(47), MAD(40), REU(105), ZAI(65). 

Hypnum macrogynum Besch. CBS(1531), 

KEN(46), LES(1531), MAD(40), MAL(46), 

NAT(1531), REU(105), TAN(46), TRV(1531), 

ZIM(46). 

Hypnum macrogynum Besch. H. aduncoides was 

treated as a synonym of H. cupressiforme var. 

cupressiforme by Ando in 1989 (169), although 

he mentioned its similarity to the Asian H. 

macrogynum. He has now synonymised H. 

aduncoides with H. macrogynum. The records 

shown here represent all collections listed in the 

literature as H. aduncoides. 

Hypnum multiflorum Schwaegr., hom. illeg. 

MAS(40). 

Hypnum radiatum Schwaegr. REU(105). 

Hypnum subcompressum C.Müll. = Entodon sub- 

compressus (1521). 

Hypnum usagarum Mitt. TAN(46). 

Hypnum vaucheri Lesq. ERI(50). 

Hypodontium dregei (Hornsch.) C.Müll. 

CBS(1531), LES(1531), MAL(67), NAT(1531), 

OFS(1531), SWA(1531), TRV(1531), ZAM(46), 

ZIM(46,61). 

Hypodontium pomiforme (Hook.) C.Müll. 

CBS(1531), NAT(1531), TRV(1531). 

Hypopterygium brevifolium Broth. SAO(47). 

Hypopterygium congoanum Dix. & Thér. ZAI(65). 

Hypopterygium grandi-stipulaceum Ren. & Card. 

MAD(40). 

Hypopterygium laricinum (Hook.) Brid. 

CBS(1531), ETH(50), FER(47), MAD(40), 

MAU(105), NAT(1531), REU(105), RWA(65), 

SAO(47), TAN(46), TRV(1531), ZAI(65), 

ZIM(46,61). 

Hypopterygium laricinum ssp. incrassato-limbatum 

(C.Müll.) Kindb. MAS(40). 

Hypopterygium mauritianum Hampe ex Besch. = H. 

laricinum (40). 

Hypopterygium mildbraedii Broth. KEN(46), 

MAL(67), RWA(65), TAN(46). 

Hypopterygium pirottae Brizi ERI(50), ETH(50). 

Hypopterygium sphaerocarpum Ren. MAD(40), 

MAU(105). 

Hypopterygium subhumile Ren. & Card. 

MAD(40), TAN(46). 

Hypopterygium tamariscinum (Hedw.) Brid. 

MAU(105). 

Hypopterygium tenellum C.Müll. FER(47). 

Hypopterygium torulosum var. kameruniae Broth. 

CAM(47). 

Hypopterygium torulosum var. nossi-beanum 

Besch. MAD(40). 

Hypopterygium torulosum var. torulosum Schimp. 

MAD(40), MAU(105), REU(105). 

Hypopterygium usambaricum Broth. TAN(46). 

Hypopterygium viridissimum C.Müll. CAM(47), 

COM(40), GAB(47), KEN(46), MAL(67), 

TAN(46), UGA(46). 

Ischyrodon lepturus (Tayl.) Schelpe CBS(1531), 

NAM(1531). 

Isopterygium ambreanum Ren. & Card. MAD(40). 

Isopterygium antsihanakense Card. MAD(40). 

Isopterygium antunesii Broth. ANG(46). 

Isopterygium aptychopsis (C.Müll.) Broth. = I. argy- 

roleucum (1878). 

Isopterygium aquaticum Dix. ZIM(46). 

Isopterygium argentatum Broth., nom. nud. 

TOG(58). 

Isopterygium argillicola (Ren. & Card.) Broth. 

MAD(40). 

Isopterygium argyroleucum Besch. ANG(46), 

BEN(47), CAM(47), CAR(47), COM(40), 

GAB(47), GUI(47), IVO(1880), MAD(40), 

MAU(105), NIG(47,334), SEY(40), TOG(1878), 

ZAI(65). 

Isopterygium asymmetricum (Ren. & Card.) Card. 

ZAI(65).

Isopterygium austro-denticulatum (Ren. & Card.) 

Broth. MAD(40). 

Isopterygium baurii Broth. TAN(46). 

Isopterygium boivinii Besch. = I. argyroleucum 

(1878). 

Isopterygium boivinii var. terrestre Ren. & Card. = I. 

argyroleucum (1878). 

Isopterygium brevicuspis Broth. in Corb. CON(47). 

Isopterygium chryseolum Besch. COM(40), 

MAS(40). 

Isopterygium citrinellum Ren. & Card. REU(105). 

Isopterygium combae Besch. MAD(40). 

Isopterygium conangium Broth. CAM(47), 

CAR(47), GAB(47), GUI(47), IVO(1880), LIB(47), 

TOG(1878). 

Isopterygium congoanum Dix. ZAI(65). 

Isopterygium dewevrei Ren. & Card. IVO(1880). 

Isopterygium dubyi Gepp = I. argyroleucum (1878). 

Isopterygium elodes P.Varde UGA(46). 

Isopterygium gabonense Broth. & P.Varde 

CAR(47), GAB(47), IVO(1880). 

Isopterygium gibbosulum (C.Müll.) Jaeg. 

COM(40). 

Isopterygium gracile Ren. & Card. MAD(40). 

Isopterygium hygrophilum Broth. RWA(65), 

ZAI(65). 

Isopterygium intortum (P.Beauv.) Jaeg. MAD(40), 

MAU(105), REU(105). 

Isopterygium intortum var. chenagonii Ren. & 


Isopterygium ivoirense Broth. & Thér. IVO(47). 

Isopterygium kilimanjaricum Broth. KEN(46), 

TAN(46). 

Isopterygium leiotheca Ren. & Card. MAS(40). 

Isopterygium leptoblastum (C.Müll.) Jaeg. 

COM(40), MAD(40). 

Isopterygium leucophanes (C.Müll.) Jaeg. 

CBS(1531), COM(40), GAB(47), MAL(67), 

MOZ(357), NAT(1531), SWA(1531), 

TRV(1531), ZAM(46,62), ZIM(46). 

Isopterygium leucopsis (C.Müll.) Par. NAT(1531), 

TRV(1531). 

Isopterygium luteo-nitens (Par.) Par. MAD(40). 

Isopterygium m’bangae (C.Müll.) Jaeg. GAB(47), 

TAN(46), UGA(46), ZAI(65). 

Isopterygium meylanii Card. MAD(40). 

Isopterygium microthecium Broth. & Par. GUI(47). 

Isopterygium minutum Ren. & Card. COM(40). 

Isopterygium molle Thér. REU(105). 

Isopterygium nanoglobum (C.Müll.) Par. SAO(47). 

Isopterygium nematosum Broth. & Par. GUI(47). 

Isopterygium nitentivirens Card. ZAI(65). 

Isopterygium nivescens (Broth.) Broth. CAM(47), 

GUI(47). 

Isopterygium palmarum (Broth.) Broth. CAM(47), 

GAB(47), IVO(1880), TOG(1878). 

Isopterygium pedunculatum Broth. & Par. 

GAB(47), GUI(47). 

Isopterygium phlyctitheca P.Varde TAN(46), 

ZAI(65), ZAM(146). 

Isopterygium phlyctitheca var. brachycladum 

P.Varde ZAI(65). 

Isopterygium plumigerum Broth. CAM(47), 

GAB(47). 

Isopterygium podorrhizum (Ren. & Card.) Card. 

MAD(40). 

Isopterygium praenitens Card. GAB(47), ZAI(65). 

Isopterygium prasiellum Besch. CON(47). 

Isopterygium punctulatum Broth. & Wag. 

CBS(1531), NAT(1531), TRV(1531). 

Isopterygium radicans (Brid.) Jaeg. REU(105). 

Isopterygium saperense Besch. COM(40). 

Isopterygium seligeri (Brid.) Dix. TAN(46). 

Isopterygium sericifolium Dix. KEN(46), TAN(46), 

UGA(46). 

Isopterygium strangulatum (C.Müll.) Broth. 

CBS(1531). 

Isopterygium subaptychopsis Broth. in Corb. 

CON(47). 

Isopterygium subconangium Broth. & P.Varde 

CAR(47). 

Isopterygium subleptoblastum C.Müll. COM(40), 

MAD(40). 

Isopterygium submicrothecium Broth. & Par. 

ANN(47), GUI(47). 

Isopterygium subpinnatum Thér., hom. illeg. 

LIB(47). 

Isopterygium taxithellioides Broth. & Bryhn 

NAT(1531). 

Isopterygium taylorii Sim. CBS(1531). 

Isopterygium tisserantii Broth. & P.Varde 

CAR(47). 

Isopterygium tisserantii var. viride P.Varde 

CAR(47). 

Isopterygium verruculosum (C.Müll.) Jaeg. 

COM(40). 

Isopterygium welwitschii Gepp ANG(46). 

Isothecium kenyae Dix. ex Tosco & Piovano 

KEN(46). 

Jaegerina excurrens Dix. = J. robillardii (1). 

137

138 

Jaegerina formosa Besch. MAU(94). 

Jaegerina plicata Card. COM(40). 

Jaegerina retrosquarrosa Allen, Crosby & Magill 

MAU(94). 

Jaegerina robillardii C.Müll. MAU(105). 

Jaegerina scariosa (Lor.) Arz. CAM(47,1), 

CAR(47,1), GAB(47,1), GHA(47,1), GUI(47), 

NIG(47,1). 

Jaegerina solitaria (Brid.) Jaeg. COM(40), 

MAU(105), REU(105). 

Jaegerina solitaria var. nossi-beana Besch. 

MAD(40). 

Jaegerina solitaria var. ramosa Besch. REU(105). 

Jaegerina stolonifera ssp. laticuspis (J.Tayl.& 

P.Varde) Argent MAL(67,1), TAN(46,1), 

UGA(46,1). 

Jaegerina stolonifera ssp. stolonifera (C.Müll.) 

C.Müll. CBS(1531), COM(40,1), KEN(46,1), 

MAD(40,1), TAN(46), UGA(46,1). 

Jaegerina stolonifera var. clavata C.Müll. = J. stolo- 

nifera var. stolonifera (1). 

Jonesiobryum sphaerocarpum Biz. ex Allen & 

Pursell NIG(47,334). 

Juratzkaea incisa (Buck) Catcheside & Stone 

CBS(1554). 

Kindbergia africana (Herz.) Ochyra KEN(46), 

RWA(65), UGA(46), ZAI(65). 

Kindbergia africana var. latifolia (Demar. & 

Leroy) Ochyra RWA(717). 

Kindbergia praelonga var. congoana (Thér.) 

Ochyra RWA(65). 

Kindbergia praelonga var. praelonga (BS&G) 

Ochyra CAP(1296), TAN(46). 

Kindbergia praelonga var. stokesii (Turn.) 

Ochyra STH(1296). 

Kleioweisiopsis denticulata Dix. KEN(46). 

Kleioweisiopsis involuta Biz. = Weissia bizotii (1420). 

Leiomela africana Thér. & Nav. KEN(46), 

REU(105), RWA(65), TAN(46), ZAI(65). 

Leiomitrium plicatum (P.Beauv.) Mitt. REU(105), 

ROD(105). 

Lepidopilidium attenuatum Thér. MAD(40). 

Lepidopilidium brunneolum (C.Müll.) Par. 

COM(40). 

Lepidopilidium cespitosum (Besch.) Broth. 

BUR(65), REU(105). 

Lepidopilidium chenagonii (Ren. & Card.) Card. 

MAD(40). 

Lepidopilidium corbieri (Ren. & Card.) Card. 

BUR(65), MAD(40). 

Lepidopilidium cyrtostegium (Ren. & Card.) Card. 

ZAI(65). 

Lepidopilidium devexum (Mitt.) Broth. CAM(47), 

TAN(46). 

Lepidopilidium flexuosum (Besch.) Par. REU(105). 

Lepidopilidium hanningtonii (Mitt.) Broth. 

CBS(1531), NAT(1531), SWA(1531), TAN(46), 

TRV(1531), UGA(46). 

Lepidopilidium isleanum (Besch.) Broth. 

REU(105). 

Lepidopilidium lastii Mitt. RWA(65), ZAI(65). 

Lepidopilidium parvulum Card. MAD(40). 

Lepidopilidium subdevexum (Broth.) Broth. 

CAM(47), CAR(47). 

Lepidopilidium subdevexum var. intermedium 


Lepidopilidium subrevolutum (Ren. & Card.) 

Card. MAD(40), MAU(105). 

Lepidopilidium therioti Nav. in Dix. RWA(65), 

ZAI(65). 

Lepidopilum callochlorum C.Müll. ex Broth. 

CAM(47), FER(47), GUI(47), NIG(47). 

Lepidopilum carrougeauanum Thér. & P.Varde 

MAD(40). 

Lepidopilum dusenii C.Müll. ex Broth. CAM(47), 

IVO(1880), SAO(47), ZAI(65). 

Lepidopilum filiferum Broth., hom. illeg. 

RWA(65). 

Lepidopilum hirsutum (Besch.) Broth. REU(105). 

Lepidopilum hirsutum var. tuberculatum Thér. 

MAD(40). 

Lepidopilum humblotii Ren. & Card. COM(40). 

Lepidopilum lastii Mitt. KEN(46), MAL(67), 

TAN(46). 

Lepidopilum niveum (C.Müll.) Kindb. CAM(47), 

CAR(47), CON(47), GAB(47), GUI(47), SAO(47). 

Lepidopilum verrucipes Card. MAD(40). 

Leptobryum pyriforme (Hedw.) Wils. CBS(1531), 

LES(1531), MAD(40), MOZ(357), NAT(1531), 

ZAI(65). 

Leptodictyum riparium (Hedw.) BS&G = 

Amblystegium riparium (71). 

Leptodictyum riparium fo. elongatum (BS&G) 

Warnst. = Amblystegium riparium fo. elongatum 

(65). 

Leptodon boryanus Besch. = L. fuciformis (301). 

Leptodon fuciformis (Brid.) Enroth MAD(40), 

MAU(105), REU(301,105). 

Leptodon smithii (Hedw.) Web. & Mohr 

CBS(1531), DJI(1569), ERI(50), ETH(50),

KEN(46), NAT(1531), OFS(1531), RWA(65), 

TAN(46), TRV(1531), UGA(46), ZIM(46). 

Leptodon smithii fo. laxus Tong. Excluded. Any 

records for this form have been associated with 

the owning species. 

Leptodon smithii var. beccarii (C.Müll.) Tong. 

DJI(50), ERI(50), ETH(50), KEN(46), TAN(46), 

UGA(46). 

Leptodontiopsis elata Dix. KEN(46). 

Leptodontiopsis fragilifolia Broth. KEN(46,136), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Leptodontiopsis fragilifolia var. brotheri P.Varde 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Leptodontiopsis macrocarpa Dix. KEN(46). 

Leptodontium abyssinicum Broth. = L. capituligerum 

(46). 

Leptodontium allorgei Biz. ex Pócs. Excluded. De 

Sloover (290) states that the specimen is probably 

a species of Cynodontium. 

Leptodontium brachyphyllum Broth. & Thér. 

CBS(1531), LES(1531), NAT(1531), 

SWA(1531), TRV(1531). 

Leptodontium capituligerum C.Müll. ETH(50), 

KEN(46). 

Leptodontium filicaule Dix. = L. flexifolium (1420). 

Leptodontium flexifolium (Dicks.) Hampe 

CAM(47), FER(47), KEN(46), LES(1158), 

REU(290), RWA(46,65), TAN(46), UGA(46), 

ZAI(65). 

Leptodontium gambaragarae Negri KEN(46), 

UGA(46). 

Leptodontium insolutum var. insolutum Thér. & 

P.Varde = Barbula eubryum (1420). 

Leptodontium insolutum var. perundulatum Thér. & 

P.Varde = Barbula eubryum (1420). 

Leptodontium joannis-meyeri C.Müll. = L. pungens 

(1420). 

Leptodontium joannis-meyeri var. cameruniae Broth. 

= L. pungens (1420). 

Leptodontium latifolium Broth. RWA(65), 

UGA(46), ZAI(65). 

Leptodontium longicaule ssp. stellatum (Brid.) 

De Sloover BUR(65), REU(290), TAN(46). 

Leptodontium longicaule var. longicaule Mitt. 

BUR(65), KEN(46), MAL(67), MAS(40), 

NAT(1531), RWA(65), SWA(1531), TAN(46), 

TRV(1531), ZAI(65), ZIM(61). 

Leptodontium longicaule var. microuncinatum 

(Dus.) Zander KEN(46), REU(105). 

Leptodontium luteum (Tayl.) Mitt. KEN(46), 

TAN(46). 

Leptodontium pumilum (C.Müll.) Kindb. = L. pungens 

(1420). 

Leptodontium pungens (Mitt.) Kindb. CAM(47), 

ETH(1865), KEN(46), REU(290), RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Leptodontium repens (C.Müll.) Kindb. TAN(46). 

Leptodontium rhynchophorum Dix. Excluded. De 

Sloover (290) states that the specimen is probably 

a species of Zygodon. 

Leptodontium serrifolium (C.Müll.) Broth. = L. 

pungens (1420). 

Leptodontium squarrosum (Hook.) Par. = L. viticu- 

losoides (1521). 

Leptodontium stellatum (Brid.) Ren. = L. longicaule 

ssp. stellatum (65). 

Leptodontium tenerascens Broth. = L. flexifolium 

(290). 

Leptodontium tenerascens var. planifolium P.Varde. 

Excluded. De Sloover (290) states that the 

specimen from Rwanda called such by Potier de 

la Varde does not belong to Leptodontium. 

Leptodontium undatifolium C.Müll. ZAI(65). 

Leptodontium viticulosoides (P.Beauv.) Wijk & 

Marg. BUR(65), CAM(47), COM(40), ETH(50), 

KEN(46), MAD(40), MAL(67), MOZ(46), 

NAT(1531), REU(105), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZAM(46), 

ZIM(46,61). 

Leptodontium volkensii Broth. DJI(1569). 

Leptodontium wallisii (C.Müll.) Kindb. RWA(65), 

TAN(1404), UGA(46), ZAI(65). 

Leptohymenium stramineum (C.Müll.) Jaeg. 

ETH(50). 

Leptoischyrodon congoanus Dix. MAL(67), 

TRV(1531), ZAI(65). 

Leptopterigynandrum austro-alpinum C.Müll. 

LES(1074,1531), NAT(1531). 

Leptotheca gaudichaudii Schwaegr. CBS(1531). 

Leskea filivaga (C.Müll.) Broth. KEN(46). 

Leskea guineensis Par. & Broth. GUI(47). 

Leskea mauritiana Besch. MAU(105). 

Leskeella zuluensis Broth. & Bryhn NAT(1531). 

Leskeodon mniifolius (Hornsch.) Bizot = Disticho- 

phyllum mniifolium (1519). 

Leucobryum acutifolium (Mitt.) Card. ANG(46), 

CBS(1531), MAD(40), MAL(67), NAT(1531), 

SWA(1531), TAN(46), TRV(1531), ZAI(65), 

ZAM(46,62), ZIM(46,61). 

Leucobryum afro-glaucum C.Müll. CAM(47), 

139

140 

CAR(47), GAB(47), GUI(47), ZAI(65). 

Leucobryum babetii Thér. & P.Varde CON(47), 

GAB(47). 

Leucobryum bistratosum Broth. TAN(46). 

Leucobryum boivinianum Besch. MAD(40). 

Leucobryum boryanum Besch. MAD(40), 

REU(105). 

Leucobryum boryanum var. minus Card. MAD(40). 

Leucobryum cambouei Card. MAD(40). 

Leucobryum cameruniae C.Müll. ex Ren. & Card. 

CAM(47), CAR(47), CON(47), GAB(47), 

IVO(44). 

Leucobryum comorense C.Müll. COM(40), 

MAD(40), MAU(105). 

Leucobryum cucullatum Broth. ANG(259), 

GAB(47), KEN(46), MAD(40), MAL(67), 

TAN(46), ZAI(65), ZAM(46). 

Leucobryum cucullatum var. rutenbergii (Geh.) 

Card. ANG(259), MAD(40). 

Leucobryum fouta-djalloni Par. & Card. CAR(47), 

GUI(47), KEN(46), MAL(67), NIG(334), 

SAO(47). 

Leucobryum galinonii Card. & Par. MAD(40). 

Leucobryum heterodictyon Besch. MAD(40). 

Leucobryum heterodictyon var. microphyllum 

(Card.) Wijk & Marg. MAD(40). 

Leucobryum homalophyllum Broth. SAO(47). 

Leucobryum isleanum Besch. COM(40), 

MAD(40), MAU(105), REU(105). 

Leucobryum isleanum var. molle (C.Müll.) Card. 

COM(40), MAD(40), MAU(105), TAN(46), 

ZAM(46). 

Leucobryum javense (Brid.) Mitt. REU(105). 

Leucobryum juniperoideum (Brid.) C.Müll. 

MAU(105), REU(105). 

Leucobryum laeve Mitt., nom. nud. MAD(40). 

Leucobryum letestui P.Varde GAB(47). 

Leucobryum leucophanoides C.Müll. SAO(47). 

Leucobryum madagassum Besch. ANG(46), 

BUR(65), KEN(46), MAD(40), MAL(67), 

RWA(65), SWA(1531), TAN(46), TRV(1531), 

ZAM(62), ZIM(61). 

Leucobryum mauritianum C.Müll., nom. nud. 

MAU(105). 

Leucobryum mayottense Card. COM(40), KEN(46), 

MAD(40), MAU(105), REU(105), TAN(46). 

Leucobryum molliculum Broth. CAM(47), 

MOZ(46), TAN(46), UGA(46). 

Leucobryum parvulum Card. MAD(40). 

Leucobryum perrotii Ren. & Card. MAD(40), 

MAU(105), TAN(46). 

Leucobryum pseudo-madagassum Card. 

MAD(40). 

Leucobryum rehmannii C.Müll. CBS(1531). 

Leucobryum sanctae-mariae Card. MAD(40). 

Leucobryum sphagnoides (Welw. & Duby) Card. 

ANG(46). 

Leucodon assimilis (C.Müll.) Jaeg. = L. maritimus 

(46). 

Leucodon bescherellei Broth. & Geh. ASC(1296). 

Leucodon canariensis (Brid.) Schwaegr. 

MAD(40). 

Leucodon capensis Schimp. = L. maritimus (1519). 

Leucodon dracaenae Solms. ex Vent. DJI(1569), 

ERI(50), ETH(50), KEN(46,136), TAN(46). 

Leucodon dracaenae var. schweinfurthii (C.Müll.) 

Fleisch. = L. dracaenae (1521). 

Leucodon laxifolius C.Müll. & Fleisch. ERI(50), 

ETH(50), KEN(46), TAN(46). 

Leucodon maritimus (Hook.) Wijk et Marg. 

CBS(1531), KEN(46), MAD(40), MAL(67), 

NAT(1531), OFS(1531), REU(105,1294), 

RWA(65), SWA(1531), TAN(46), TRV(1531), 

UGA(46), ZAI(65), ZAM(46), ZIM(46). 

Leucodon rutenbergii C.Müll. ex Geh. MAD(40), 

TAN(46). 

Leucodon schweinfurthii C.Müll. = L. dracaenae 

(1521). 

Leucodon schweinfurthii var. gracilior Broth ex Mich. 

= L. dracaenae (1521). 

Leucodon sciuroides var. sciuroides (Hedw.) 

Schwaegr. KEN(46), TAN(46). 

Leucodontopsis cameruniae (Broth.) Broth. = Leu- 

codontopsis geniculata (28) (= Henicodium 

geniculatum). 

Leucodontopsis geniculata (Mitt.) Crum & Steere = 

Henicodium geniculatum (1339). 

Leucoloma albocinctum Ren. & Card. MAD(40), 

MAL(67). 

Leucoloma albocinctum var. subelimbatum Ren., 

nom. nud. MAD(40). 

Leucoloma allorgei Thér. MAD(40). 

Leucoloma amblyacron Besch. MAU(105). 

Leucoloma ambreanum Ren. & Card. MAD(40). 

Leucoloma aspericuspis P.Varde KEN(46), 

MAL(67), TAN(46). 

Leucoloma bifidum (Brid.) Brid. KEN(46), 

MAD(40), MAS(40), TAN(46). 

Leucoloma bifidum var. tanganikae P.Varde & 

Thér. TAN(46).

Leucoloma boivinianum var. angasizae (Ren.) 

Par. COM(40). 

Leucoloma boivinianum var. boivinianum Besch. 

COM(40), MAD(40), REU(105). 

Leucoloma boivinianum var. brevifolium Ren. 

MAD(40). 

Leucoloma brevioperculatum Dix. MAD(40). 

Leucoloma brotheri Ren. MAD(40), TAN(46). 

Leucoloma candidulum Besch. MAU(105), 

REU(105). 

Leucoloma candidum Broth. MAD(40). 

Leucoloma capillifolium Ren. MAD(40). 

Leucoloma cespitulans (C.Müll.) Jaeg. COM(40). 

Leucoloma charrieri Thér. & P.Varde MAD(40). 

Leucoloma chlorophyllum Broth. TAN(46). 

Leucoloma chrysobasilare (C.Müll.) Jaeg. 

COM(40), MAD(40), NAT(1531), TAN(46), 

TRV(1531), ZIM(46,61). 

Leucoloma chrysobasilare var. gracilicaulon 

(C.Müll.) Jaeg. COM(40), MOZ(46), TAN(46), 

ZIM(46). 

Leucoloma chrysobasilare var. procerum (Ren.) 

Broth. COM(40). 

Leucoloma cinclidotioides Besch. 

MAU(298,105), REU(105). 

Leucoloma cirrosulum Ren. MAD(40). 

Leucoloma comorae Ren. COM(40), MAD(40). 

Leucoloma convolutaceum Ren. MAD(40). 

Leucoloma crepinii Ren. & Card. MAD(40), 

MAU(105), REU(105). 

Leucoloma crepinii var. orthothecioides (Besch.) 

Ren. MAU(105). 

Leucoloma crosbyi LaFarge-England MAD(1617). 

Leucoloma cuneifolium (C.Müll. ex Geh.) Wright 

MAD(40), TAN(46). 

Leucoloma decaryi Thér. MAD(40). 

Leucoloma decaryi var. comorense Thér. COM(40). 

Leucoloma decolor Wright TAN(46). 

Leucoloma delicatulum Ren. MAD(40). 

Leucoloma delicatulum var. laxobasis Ren. & 


Leucoloma dichelymoides (C.Müll.) Jaeg. 

COM(40), MAD(40), MAU(105). 

Leucoloma dubyanum Besch. MAD(40), 

MAU(105). 

Leucoloma fontinaloides Dix. MAD(40). 

Leucoloma forsythii Thér. MAD(40). 

Leucoloma fuscifolium Besch. MAD(40), 

MAU(105), REU(105). 

Leucoloma fuscifolium var. crispatulum Ren. 

MAD(40), REU(105). 

Leucoloma garnieri Par. & Ren. MAD(40). 

Leucoloma garnieri fo. aquaticum Thér. Excluded. 



Leucoloma gracilescens Broth. SAO(47). 

Leucoloma grandidieri Ren. & Card. MAD(40). 

Leucoloma grimmioides P.Varde MAD(40), 

TAN(46). 

Leucoloma guineense Broth. & Par. GUI(47). 

Leucoloma holomitrioides C.Müll. MAD(40). 

Leucoloma holstii Broth. MAL(67), RWA(65), 

TAN(46). 

Leucoloma humbertii P.Varde MAD(40). 

Leucoloma incrassatum Thér. MAD(40). 

Leucoloma isleanum Par. COM(40), MAU(105), 

SEY(40). 

Leucoloma isleanum var. subtortile Ren. 

COM(40). 

Leucoloma latifolium Broth. & P.Varde GAB(47). 

Leucoloma lepervanchei Besch. REU(105). 

Leucoloma leptocladum Ren. MAD(40). 

Leucoloma levieri Ren. MAD(40). 

Leucoloma longifolium (Brid.) Wijk & Marg. 

MAD(40), MAU(105), REU(105). 

Leucoloma mafatense Ren. REU(105). 

Leucoloma meteorioides P.Varde MAD(40). 

Leucoloma normandii Par. & Broth. ex Ren. 

GUI(47), NIG(47), SIL(47). 

Leucoloma normandii var. alpinum Par. & Broth. 

GUI(47). 

Leucoloma ochrobasilare Ren. MAD(40). 

Leucoloma pallidulum Thér. MAU(105). 

Leucoloma pennequinii Par., nom. nud. MAD(40). 

Leucoloma perrieri Thér. MAD(40). 

Leucoloma persecundum Besch. MAD(40), 

MAU(105). 

Leucoloma persecundum var. perrotii Ren. 

MAD(40). 

Leucoloma pobeguinii Par. & Broth. GUI(47). 

Leucoloma pumilum Wright MAD(40). 

Leucoloma pusillum Card. MAD(40). 

Leucoloma pygmaeum Par. GUI(47). 

Leucoloma rehmannii (C.Müll.) Rehm. ex Par. 

CBS(1531), MAL(67), NAT(1531), TRV(1531), 

ZAM(46), ZIM(46,61). 

Leucoloma rutenbergii (Geh.) Wright MAD(40). 

Leucoloma rutenbergii var. abbreviatum Ren. 

MAD(40). 

Leucoloma rutenbergii var. elatum Ren. 

141

142 

MAD(40), REU(298). 

Leucoloma rutenbergii var. perrotii Ren. 

MAD(40). 

Leucoloma rutenbergii var. porosis Thér. 

MAU(105). 

Leucoloma sanctae-mariae Besch. MAD(40). 

Leucoloma sanctae-mariae var. leptolimbatum 

Ren. MAD(40). 

Leucoloma schelpei P.Varde MOZ(46). 

Leucoloma secundifolium Mitt. CAM(47), 

SAO(47). 

Leucoloma sericeum P.Varde SIL(47). 

Leucoloma serraticuspis P.Varde MOZ(46). 

Leucoloma seychellense Besch. SEY(40). 

Leucoloma silvaticum Ren. MAD(40). 

Leucoloma sinuosulum C.Müll. ex Besch. 

COM(40), MAD(40), MAU(105), REU(105), 

TAN(46). 

Leucoloma sprengelianum (C.Müll.) Jaeg. 

CBS(1531), TRV(1531). 

Leucoloma squarrosulum (Geh.) Wright 

MAD(40). 

Leucoloma squarrosulum ssp. subplicatum (Ren. 

& Card.) Broth. MAD(40). 

Leucoloma subbifidum Ren. MAD(40). 

Leucoloma subcespitulans Besch. REU(105). 

Leucoloma subchrysobasilare Ren. MAD(40). 

Leucoloma subsecundifolium Broth. TAN(46). 

Leucoloma subtuberculosum Ren. MAD(40), 

MAS(40). 

Leucoloma subtuberculosum var. longifolium 


Leucoloma syrrhopodontioides Broth. 

CBS(1531), TRV(1531), ZIM(61). 

Leucoloma talazaccii Ren. & Card. MAD(40). 

Leucoloma talazaccii var. fragilifolium Thér. & 

P.Varde MAD(40). 

Leucoloma tanganyikae P.Varde TAN(46). 

Leucoloma terricola Broth. TAN(46). 

Leucoloma thraustum Besch. MAD(40). 

Leucoloma thuretii Besch. MAD(40). 

Leucoloma tuberculosum Ren. MAD(40). 

Leucoloma villaumei Thér. MAD(40). 

Leucoloma volkensii Broth. KEN(46), TAN(46). 

Leucoloma zanzibarense Broth. TAN(46). 

Leucomium aneurodictyon (C.Müll.) Jaeg. 

CAR(47), COM(40), GHA(47), IVO(1880), 

LIB(47), MAD(40), SEY(40), TAN(46), ZAI(65). 

Leucomium commixtum (C.Müll.) Kindb. 

CAM(47). 

Leucomium golungense Gepp in Hiern. = L. strumosum 

(1878). 

Leucomium le-testui Broth. & P.Varde = L. strumosum 

(1878). 

Leucomium mahorense Besch. = L. aneurodictyon 

(40). 

Leucomium perglaucum C.Müll. ex Broth. 

CAM(47), CAR(47), GAB(47), TAN(46), 

ZAI(65). 

Leucomium strumosum (Hornsch.) Mitt. ANG(46), 

CAR(47), CON(47), FER(47), GAB(47), 

SAO(47), TAN(46), TOG(1878). 

Leucoperichaetium eremophilum Magill 

NAM(1531). 

Leucophanella revoluta (Dozy & Molk.) Fleisch. = 

Syrrhopodon revolutus (1521) (= S. involutus). 

Leucophanes angustifolium Ren. & Card. 

CAM(47,1896), COM(1149,1896), 

GHA(1149,1896), MAU(105), 

REU(1149,105,1896), SEY(1149,1896), 

TAN(46,1896), UGA(46), ZAI(65). 

Leucophanes annobonense Broth. in Mildbr., 


Leucophanes annobonense Broth. in Mildbr., nom. 

nud. This taxon is not covered in Salazar Allen’s 

world-wide revision of the genus (1896). 

Leucophanes cameruniae C.Müll ex Dus. = L. molleri 

(1149). 

Leucophanes denticuspis C.Müll ex Dus. = L. angu- 

stifolium (1149). 

Leucophanes glaucum (Schwaegr.) Mitt. 

SEY(1896). 

Leucophanes heterodictyon Onr. = L. renauldii 

(1896). 

Leucophanes hildebrandtii C.Müll. 

COM(40,1896), MAD(40,1896), 

MAU(105,1896), REU(105), TAN(46,1896). 

Leucophanes hildebrandtii fo. rigidum Besch. Ex- 



Leucophanes lecomptei Besch. = L. molleri (1149). 

Leucophanes mayottense Card. = L. angustifolium 

(1149). 

Leucophanes molleri C.Müll. CAM(47,1896), 

CAR(47), CON(47,1896), GAB(47), 

GHA(1149,1896), GUI(47), IVO(47,1896,1880), 

LIB(47,1896), MAU(1896), SAO(47,1896), 

TAN(46,1896), UGA(1896), ZAI(65,1896). 

Leucophanes octoblepharioides Brid. CAR(47), 

GAB(47), MAU(1896), SEY(1896).

Leucophanes renauldii Card. COM(1896), 

MAD(40,1896), MAU(105,1896), TAN(46). 

Leucophanes rodriguezii C.Müll. COM(40), 

MAD(1896), MAU(1896), REU(1896), 

TAN(46). 

Leucophanes scabridens P.Varde = L. 

octoblepharoides (1896). 

Leucophanes seychellarum Besch. SEY(40,1896), 

TAN(46). 

Leucophanes unguiculatum Mitt. NIG(1896), 

PRI(47,1896). 

Levierella fabroniacea C.Müll. KEN(408,46), 

NAT(1531), SWA(1531), TRV(1531). 

Levierella fabroniacea var. abyssinica Dix. = L. 

perserrata (46). 

Levierella perserrata P.Varde & Leroy ERI(50), 

ETH(50), KEN(46), TAN(46), UGA(46), ZAI(65), 

ZAM(46). 

Levierella subfabroniacea Broth. & Par. GUI(47). 

Lindbergia abbreviata (Schimp. ex Jaeg.) Broth. 

ETH(50). 

Lindbergia erythraea (C.Müll.) Fleisch. ERI(50). 

Lindbergia haplocladioides Dix. CBS(1531), 

KEN(46), NAT(1531), OFS(1531), TRV(1531), 

ZIM(61). 

Lindbergia patentifolia Dix. CBS(1531), DJI(1569), 

KEN(46), NAT(1531), OFS(1531), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZIM(46,61). 

Lindbergia patentifolia fo. perrobusta P.Varde. Ex- 



Lindbergia pseudoleskeoides Dix. CBS(1531), 

LES(1531), MOZ(46), NAT(1531), OFS(1531), 

TRV(1531), ZIM(46). 

Lindbergia viridis Dix. CBS(1531), NAT(1531), 

OFS(1531), SWA(1531), TRV(1531). 

Lindigia africana Broth. = Aerolindigia capillacea 

(65). 

Lindigia hildebrandtii Ren. MAD(40). 

Loiseaubryum ephemeroides Biz. CHA(4), 

NIG(1883). 

Lopidium campenonii (Ren. & Card.) Fleisch. = L. 

struthiopteris var. campenonii (108). 

Lopidium hemiloma (C.Müll.) Fleisch. COM(40), 

TAN(46), ZAI(65). 

Lopidium pennaeforme (Brid.) Fleisch. CBS(1531), 

NAT(1531), TAN(46), ZIM(61). 

Lopidium struthiopteris (Brid.) Fleisch. KEN(46), 

REU(105), TAN(46). 

Lopidium struthiopteris var. campenonii (Ren. & 

Card.) Biz. KEN(46), MAD(40), TAN(46). 

Lopidium subtrichocladon (Broth.) Fleisch. 

SAO(47), TAN(46). 

Macrocoma abyssinica (C.Müll.) Vitt. CAM(47), 

ETH(50), KEN(46), MAL(67), RWA(65), 

SOC(40,1626), TAN(46), ZAI(65). 

Macrocoma lycopodioides (Schwaegr.) Vitt 

CBS(1531), NAT(1531), OFS(1531), 

SWA(1531), TRV(1531). 

Macrocoma lycopodioides (Schwaegr.) Vitt. Van 

Rooy (1547), fide Vitt, says that this taxon is 

restricted to southern Africa, and the record for 

Zimbabwe (61) is erroneus. Similarly I have 

assumed that Gepp’s 1901 collection from Angola 

is also unreliable. 

Macrocoma pulchella (Hornsch.) Vitt CBS(1531). 

Macrocoma pulchella (Schwaegr.) Vitt. Van Rooy 

(1547), fide Vitt, says that this taxon is restricted 

to southern Africa, and the record for Zimbabwe 

(61) is erroneus. Similarly I have assumed that 

Potier de la Varde’s 1947 Kenya record is also 

unreliable. 

Macrocoma tenue ssp. tenue (Hook. & Grev.) Vitt 

ANG(46), CBS(1531), KEN(46), LES(1531), 

MAD(40), MAL(67), MOZ(357), NAT(1531), 

OFS(1531), REU(105), RWA(65), STH(1296), 

SWA(1531), TAN(46), TRV(1531), ZAI(65), 

ZIM(46,61). 

Macrohymenium acidodon (Mont.) Dozy & Molk. 

KEN(46), MAD(40), REU(105), SEY(40), 

TAN(46). 

Macrohymenium acidodon fo. minutissimum Besch. 



Macrohymenium incrassatum (Dix. & Thér.) Kis 

UGA(46), ZAI(65). 

Macrohymenium megasporum (Duby) Kis 

CAM(47), COM(40), GAB(47), MAD(40), 

MAU(105), REU(105), SEY(40), TAN(46). 

Macromitrium abessinianum var. angustifolium 

Broth., nom. dub. ex err. TAN(46). 

Macromitrium abyssinicum C.Müll. = Macrocoma 

abyssinica (65). 

Macromitrium adelphinum Card. MAD(40). 

Macromitrium amaniense P.Varde TAN(46). 

Macromitrium ancistrophyllum Card. MAD(40). 

Macromitrium anomodictyon Card. MAD(40). 

Macromitrium astroideum Mitt. ROD(105). 

Macromitrium belangeri C.Müll. REU(105). 

143

144 

Macromitrium bequaerti Thér. & Nav. ZAI(65). 

Macromitrium calocalyx Geh. MAD(40). 

Macromitrium calomicron Broth. MAU(105). 

Macromitrium chlororete C.Müll. ex Par., nom. 

nud. ETH(50). 

Macromitrium cucullatum C.Müll. ex Brizi ERI(50). 

Macromitrium dusenii C.Müll. ex Broth. CAM(47), 

GAB(47), IVO(1880), LIB(47), NIG(47), ZAI(65). 

Macromitrium eckendorfii Thér. & P.Varde = M. 

sulcatum (1878). 

Macromitrium ecrispatum Dix. TAN(46). 

Macromitrium fasciculare Mitt. MAD(40), 

REU(105), TAN(46). 

Macromitrium fasciculare var. angustifolium 

P.Varde MAD(40). 

Macromitrium fimbriatum (P.Beauv.) Schwaegr. 

MAD(40), MAU(105), REU(105). 

Macromitrium fimbriatum var. chloromitrium 

Besch. MAU(105). 

Macromitrium fragile Negri UGA(46). 

Macromitrium funicaule Besch. MAU(105). 

Macromitrium gimalacii Biz. & Onr. REU(105). 

Macromitrium hildebrandtii C.Müll. COM(40). 

Macromitrium lanceolatum Broth. MAU(105). 

Macromitrium lebomboense Van Rooy 

CBS(1531,1438), NAT(1531,1438). 

Macromitrium leucoblastum C.Müll. ex Par., nom. 

nud. ETH(50). 

Macromitrium levatum Mitt. = M. sulcatum (1878). 

Macromitrium liliputanum C.Müll. KEN(46). 

Macromitrium macropelma C.Müll. CBS(1531). 

Macromitrium mauritianum Schwaegr. DIE(40), 

MAD(40), MAU(105), REU(105), TAN(46). 

Macromitrium mauritianum var. viride Broth. 

MAU(105). 

Macromitrium megalosporum Thér. & Nav. 

KEN(46), ZAI(65). 

Macromitrium microphyllum Hook. & Grev. = M. 

tenue (1521) (= Macrocoma tenue). 

Macromitrium nanothecium Card. MAD(40). 

Macromitrium nanothecium var. sublaeve Thér. 

MAD(40). 

Macromitrium onraedtii Biz. MAD(40). 

Macromitrium pallidum (P.Beauv.) Wijk & Marg. 

MAU(105), REU(105). 

Macromitrium proximum Thér. MAD(40). 

Macromitrium recurvum Mitt., nom. nud. 

MAU(105). 

Macromitrium rhizomatosum C.Müll. MAD(40). 

Macromitrium richardii Schwaegr. 

CBS(1531,1438), NAT(1531,1438). 

Macromitrium rufescens Besch. COM(40), 

KEN(46), MAD(40), REU(105), TAN(46). 

Macromitrium sanctae-mariae Ren. & Card. 

MAD(40). 

Macromitrium sclerodictyon Card. MAD(40). 

Macromitrium scleropodium Besch. CAM(47), 

GAB(47), MAD(40), REU(105), ZAI(65). 

Macromitrium seemannii Mitt. STH(1296). 

Macromitrium semipapillosum Thér. & P.Varde 

MAD(40). 

Macromitrium seritum Par. & Broth. GUI(47). 

Macromitrium serpens (Hook. & Grev.) Brid. 

CBS(1531), MAD(40), MAL(67), MOZ(46), 

NAT(1531), REU(105). 

Macromitrium soulae Ren. & Card. MAD(40). 

Macromitrium subpungens Hampe ex C.Müll. 

COM(40), MAD(40), SEY(40), TAN(46). 

Macromitrium subpungens var. madagassum 


Macromitrium subtortum (Hook. & Grev.) 

Schwaegr. MAD(40). 

Macromitrium sulcatum (Hook.) Brid. CAM(47), 

CBS(1531), CAR(47), COM(1447), FER(47), 

GAB(47), GHA(47), IVO(47), KEN(46), 

MAD(40), MAL(67), MOZ(46), NAT(1531), 

NIG(47), PRI(47), RWA(65), SAO(47), SIL(47), 

TAN(46), TOG(1878), TRV(1531), ZAI(65), 

ZAM(46,62), ZIM(46,61). 

Macromitrium tristratosum Dix. MAD(40), 

TAN(46). 

Macromitrium trollii Dix. KEN(46). 

Macromitrium urceolatum C.Müll. ex Geh. 

DIE(40), MAD(40), STH(1296), TAN(46). 

Macromitrium virescens C.Müll. ex Vent. ERI(50). 

Macromitrium voeltzkowii Broth. MAU(105), 

REU(105). 

Meesia kenyae P.Varde KEN(46). 

Meiotheciopsis usambarica (Broth.) Buck. 

TAN(46). 

Meiothecium chlorocladum (Broth.) Broth. 

CAM(47). 

Meiothecium chlorocladum var. intermedium 

Thér. ZAI(65). 

Meiothecium crassirete (Ren. & Card.) Card. 

MAD(40). 

Meiothecium fuscescens (Jaeg.) Broth. CBS(1531). 

Meiothecium guineense (Broth. & Par.) Sch.- 

Mot. GUI(47). 

Meiothecium madagascariense (Brid.) Broth.

MAD(40), ROD(1631). 

Meiothecium pobeguinii (Broth. & Par.) Broth. 

GUI(47). 

Meiothecium undulatum (Ren. & Card.) O’Shea 

MAD(40). 

Mesonodon flavescens (Hook.) Buck CAM(47), 

GUI(47), MAD(40), MAL(67), MAU(105), 

TAN(46), TOG(1878). 

Metzlerella madagascariensis Thér. = Atractylocarpus 

madagascariensis (465). 

Microbryum davallianum var. conicum (Schleich. 

ex Schwaegr.) Zander CBS(1531). 

Microbryum rufochaete (Magill) Zander 

CBS(1531). 

Microbryum subplanomarginatum (Dix.) Zander 

CBS(1531), NAM(1531). 

Microcampylopus kouroussensis (Ren. & Par.) Rich. 

& Clear = Campylopus kouroussensis Ren. & Par 

(507) = Campylopus perpusillus (54). 

Microcampylopus laevigatus (Thér.) Giese & 

Frahm KEN(46), MAD(40), MAL(46,67), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Microcampylopus nanus (C.Müll.) Broth. ex Par. = 

Dicranella subsubulata (507). 

Microcampylopus perpusillus (Mitt.) Broth. ex Par. 

= Campylopus perpusillus (507). 

Microdus afro-exiguus (Dus.) Par. CAM(47). 

Microdus ampullaceus (Dus.) Par. MAL(67), 

NIG(47). 

Microdus congolense Ren. & Card. ZAI(65). 

Microdus glabrifolium (C.Müll.) Kindb. 

RWA(65), ZAI(65). 

Microdus ligulifolius (C.Müll.) Broth. NIG(47). 

Microdus lutarius Besch. REU(105). 

Microdus minutus (Hampe) Besch. CBS(1531), 

COM(40), LES(1531), MAD(40), MAL(67), 

MAU(105), NAT(1531), SWA(1531), TAN(46), 

TRV(1531). 

Microdus moenkemeyeri (C.Müll.) Par. NIG(47). 

Microdus nitidulus (Mitt.) Par. CAR(47), GAB(47), 

GHA(47), IVO(1880), NIG(47), TOG(58,1878), 

ZAI(65). 

Microdus pallidisetus (Brid.) Besch. MAD(40). 

Microdus subnitidulus Thér. & P.Varde CAR(47), 

GAB(47), ZAI(65). 

Micromitrium elimbatum Thér. = Groutiella elimbata 

(See “New Combinations”). 

Micropoma bukobense Broth. = Physcomitrium bu- 

kobense (1680). 

Micropoma niloticum (Del.) Lindb. = Physcomitrium 

niloticum (1680). 

Microthamnium afro-elegantulum C.Müll. = Mitte- 

nothamnium horridulum (1521) 

Mielichhoferia basilaris Bruch. & Schimp. ex C.Müll. 

= Schizymenium basilaris (1487). 

Mielichhoferia bequaertii Dix. & Nav. = Pohlia 

bequaertii (65). 

Mielichhoferia borbonica Thér. REU(105). 

Mielichhoferia bryoides (Harv.) Wijk et Marg. 

CBS(1531), LES(1531), MAL(67), NAT(1531), 

OFS(1531), TRV(1531), ZIM(46). 

Mielichhoferia cardotii Shaw & Crum MAD(40). 

Mielichhoferia clavata Bruch & Schimp. ETH(50), 

SUD(1419). 

Mielichhoferia commixta Thér. = Schizymenium com- 

mixtum (1487). 

Mielichhoferia cratericola Broth. = Schizymenium 

cratericola (65). 

Mielichhoferia elgonensis P.Varde TAN(46), 

UGA(46). 

Mielichhoferia elongata (Hoppe & Hornsch.) Nees & 

Hornsch. = M. mielichhoferi var. elongata (1521). 

Mielichhoferia hariotii Thér. = Schizymenium hariotii 

(1487). 

Mielichhoferia hymenostoma Bruch. & Schimp. ex 

C.Müll. = Schizymenium hymenostomum (1487). 

Mielichhoferia hymenostomoides P.Varde 

UGA(46). 

Mielichhoferia linderi Thér. RWA(65). 

Mielichhoferia madagassa Ren. & Card. ex Card. = 

Schizymenium madagassum (1487). 

Mielichhoferia madagassa var. gracilis Thér. 

MAD(40). 

Mielichhoferia madagassa var. gracilis Thér. Mie- 

lichhoferia madagassa has now been transferred 

to Schizymenium, but this variety was not 

considered at the time of the transfer. It should 

be synonymised or transferred. 

Mielichhoferia mildbraedii Broth. = Schizymenium 

mildbraedii (65). 

Mielichhoferia ovalis Mitt. CAM(47), IVO(1458). 

Mielichhoferia ruwenzorensis Thér. & Nav. = Schizy- 

menium ruwenzorensis (65). 

Mielichhoferia schimperi Besch. ex Paris ETH(50). 

Mielichhoferia speluncae P.Varde UGA(46). 

Mielichhoferia squarrulosa var. fragilis Thér. & 

Nav. ZAI(65). 

Mielichhoferia subbasilaris Broth. RWA(65), 

UGA(46). 

Mielichhoferia subnuda Sim CBS(1531), 

145

146 

LES(1531), NAT(1531), OFS(1531). 

Mielichhoferia vardei Shaw & Crum CAM(47). 

Mielichhoferia woodii P.Varde UGA(46). 

Mittenothamnium aureum (Besch.) Card. = M. reptans 

(717). 

Mittenothamnium bescherellei (Ren. & Card.) 

Card. REU(105). 

Mittenothamnium brevicuspis (P.Varde) Wijk & Marg. 

= Chryso-hypnum frondosum (717). 

Mittenothamnium caudiforme (C.Müll.) Card. 

MOZ(46). 

Mittenothamnium cavifolium (Dix.) Wijk & Marg. = 

Chryso-hypnum cavifolium (717). 

Mittenothamnium ctenidioides (Dix.) Schelpe 

CBS(1531), TAN(46). 

Mittenothamnium cygnicollum (Dix.) Wijk & Marg. 

= M. reptans (717). 

Mittenothamnium frondosum (Mitt.) Card. = Chryso- 

hypnum frondosum (717). 

Mittenothamnium fruticellum (Mitt.) Card. = M. 

reptans (717). 

Mittenothamnium horridulum (Broth.) Card. 

CAM(47), GUI(47), MAD(40), NAT(1531), 

NIG(334). 

Mittenothamnium isopterygioides (Ren. & Card.) 

Card. GAB(47). 

Mittenothamnium lacouturei (Ren. & Card.) Card. 

MAD(40). 

Mittenothamnium leptoreptans (Broth.) Card. 

SAO(47). 

Mittenothamnium limosum (Besch.) Card. 

MAD(40), REU(105). 

Mittenothamnium madagassum (Besch.) Card. 

COM(40), MAD(40), REU(105). 

Mittenothamnium microthamnioides (Geh.) Wijk 

& Marg. MAD(40), MOZ(46), ZAM(46). 

Mittenothamnium microthamnioides var. glauculum 

(Card.) Wijk & Marg. MAD(40). 

Mittenothamnium overlaetii Thér. & Nav. 

TAN(46), ZAI(65). 

Mittenothamnium overlaetii var. kilimandscharicum 

(Broth.) Kis TAN(46). 

Mittenothamnium pallescens Broth. in Mildbr. 

ANN(47). 

Mittenothamnium patens (Hampe) Card. 

CBS(1531), NAT(1531), TAN(46), TRV(1531). 

Mittenothamnium perrieri (Thér.) W.Sch.-Mot. 

CAR(47). 

Mittenothamnium perrieri (Thér.) Schultze-Motel. 

Crosby & Schultze-Motel (40) suggest that this is 

probably a synonym of M. microthamnioides 

var. glauculum. 

Mittenothamnium plano-squarrosum (Broth.) Card. 

= Rhachopilopsis trinitensis (717). 

Mittenothamnium pobeguinii (Broth. & Par.) Card. 

= Rhachopilopsis trinitensis (717). 

Mittenothamnium proreptans (C.Müll.) Card. 

BUR(65), ZAI(65). 

Mittenothamnium pseudoreptans (C.Müll.) Card. 

CBS(1531), KEN(46), NAT(1531), TAN(46), 

TRV(1531), UGA(46), ZIM(46). 

Mittenothamnium reptans (Hedw.) Mitt. 

CAM(47), CBS(1531), FER(47), KEN(46), 

MAD(40), MAL(67), NAT(1531), REU(105), 

RWA(65), TAN(46), TRV(1531), ZAI(65), 

ZIM(46). 

Mittenothamnium saproadelphus (Broth.) Card. 

CAM(47), MAU(105). 

Mittenothamnium serratum (Jaeg.) Card. = M. reptans 

(717). 

Mittenothamnium stuhlmannii (Broth.) Card. = Rha- 

chopilopsis kilimandscharica (717). 

Mittenothamnium subelegantulum (Broth.) Card. = 

Rhachopilopsis trinitensis (717). 

Mittenothamnium subpatens (Broth.) Kis 

TAN(46). 

Mitthyridium chenagonii (Ren. & Card.) Robins. = 

Serpotortella chenagonii (1474). 

Mitthyridium cyrtophyllum (Besch.) Robins. = Serpo- 

tortella cyrtophylla (1474). 

Mitthyridium fasciculatum (Hook. & Grev.) 

H.Robins. MAD(1375), MAU(105). 

Mitthyridium guineense (Broth. & Par.) Sch.- 

Mot. GUI(47). 

Mitthyridium letestui (Thér. & P.Varde) Robins. = M. 

undulatum (1474). 

Mitthyridium micro-undulatum (Dix.) H.Robins. 

SEY(40). 

Mitthyridium micro-undulatum ssp. comoroense 

Reese COM(1713). 

Mitthyridium subflavum (Ren. & Card.) Robins. = 

Syrrhopodon parasiticus var. disciformis (1474) 

(= Calymperopsis disciformis). 

Mitthyridium undulatum (Dozy & Molk.) 

H.Robins. GAB(47). 

Mniobryum linderi Thér. ZAI(65). 

Mniobryum pergracile Thér. & P.Varde CAR(47), 

KEN(46), TAN(259), UGA(46), ZAI(65). 

Mniobryum perlaxum Thér. & P.Varde MAD(40). 

Mniobryum ruwenzorense P.Varde UGA(46).

Mniobryum wahlenbergii (Web. & Mohr) Jenn. 

MAU(105). 

Mnium punctatum (Schreb.) Hedw. = Rhizomnium 

punctatum. Excluded. According to Koponen 

(1036), this taxon is not known from sub-Saharan 

Africa. 

Mnium rostratum (Schrad.) Schwaegr. = Plagiomnium 

rostratum (1036) (= P. rhynchophorum). 

Moenkemeyera diaphanodonta P.Varde = Fissidens 

diaphanodontus (46). 

Moenkemeyera etessei Broth. & Par. = Fissidens 

fouta-djalloni (76) (= F. gardneri). 

Moenkemeyera flexipes P.Varde = F. parkii (72). 

Moenkemeyera hians (Hampe) C.Müll. See Fissidens 

hians (Hampe)Par. It is likely that these two 

taxa, described in successive years are homotypic, 

based on a collection made by Rabenhorst near 

Lagos. (Bruggeman-Nannenga, pers. comm.) 

Moenkemeyera konkourae (Par. & Broth.) P.Varde = 

Fissidens microcarpus (73). 

Moenkemeyera macrocarpa Broth. & Par. = Fissidens 

parkii (72). 

Moenkemeyera macroglossa Broth. = Fissidens mac- 

roglossus (1458 (fide Bruggeman-Nannenga)). 

Moenkemeyera macroglossoides Broth. 

CAM(47). 

Moenkemeyera mirabilis C.Müll. = F. parkii (72). 

Moenkemeyera scleromitria (Besch.) P.Varde = F. 

scleromitrius (65). 

Moenkemeyera subsphaerocarpa P.Varde = Fissidens 

microcarpus (73). 

Molendoa sendtneriana (BS&G) Limpr. SOM(50). 

Molendoa sublaevis Demar. & P.Varde ZAI(65). 

Nanobryum dummeri Dix. CBS(1531), NAT(1531). 

Nanobryum gladiolum (Mitt.) Biz. = Fissidens gla- 

diolus (46). 

Neckera ascensionis Besch. = Himantocladium as- 

censionis (1521). 

Neckera complanata var. maxima Dix. KEN(46). 

Neckera ehrenbergii C.Müll. = N. remota (1521). 

Neckera extans Besch. = Neckeropsis disticha (286). 

Neckera hedbergii P.Varde UGA(46). 

Neckera hedbergii P.Varde. De Sloover (286) states 

that without sporophytes, it is impossible to 

assign this taxon to a species, but suggests that it 

might be a synonym of N. remota. 

Neckera hoehneliana C.Müll. ERI(50), KEN(46). 

Neckera homoclada C.Müll. in Par., nom. nud. 

ETH(50). 

Neckera intermedia Brid. CAP(1296). 

Neckera nelloi Broth. ex Broth. & Tong ERI(50). 

Neckera nelloi Broth. ex Broth. & Tong. De Sloover 

(286) suggests that this taxon is probably a 

synonym of N. remota. 

Neckera occidentalis Besch. = Neckeropsis disticha 

(286). 

Neckera pennata Hedw. FER(47), ZIM(61). 

Neckera pennata Hedw. De Sloover (286) states that 

is likely that the identifications of the above 

specimens are in error. 

Neckera pertruncata Card. = Neckeropsis lepineana 

(1848). 

Neckera platyantha (C.Müll.) Par. BUR(65), 

KEN(46), RWA(65,46), TAN(46), UGA(46), 

ZAI(65). 

Neckera remota Bruch & Schimp. ex C.Müll. 

BUR(65), CAM(47), DJI(50), ERI(50), ETH(50), 

KEN(46), MAD(40), RWA(65), TAN(46), 

UGA(46), ZAI(65). 

Neckera submacrocarpa Dix. KEN(46,136), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Neckera subremota C.Müll. ex Dus. = N. remota (46). 

Neckera valentiniana Besch. CBS(1531), 

MAD(40), NAT(1531), REU(105), TRV(1531), 

ZAI(65). 

Neckera valentiniana Besch. De Sloover (286) states 

that the records of this plant for ‘central Africa’ 

are in error. 

Neckeropsis afrovictoriae (Dus.) Fleisch. = N. foveolata 

(787). 

Neckeropsis angustiretis Dix. ex Tosco & Piovano, 

nom. nud. Excluded. Enroth (787) states that 

not only has this taxon no description, there are 

no records of Neckeropsis for Kenya (the reputed 

origin of the specimen), and it is thus a doubtful 

record. In addition, Tosco & Piovano (407) state 

that although Dixon must have seen a specimen, 

there is now only an empty folder, and the 

specimen is missing. 

Neckeropsis bequaerti Thér. & Nav. = N. madecassa 

(787). 

Neckeropsis boiviniana (Besch.) Card. MAD(40). 

Neckeropsis disticha (Hedw.) Kindb. BUR(65), 

CAM(47), CAR(47), COM(40), CON(47), 

FER(47), GAB(47), GHA(47), IVO(787,1880), 

LIB(787,108), NIG(787), PRI(787), 

TOG(58,1878), UGA(46), ZAI(65). 

Neckeropsis foveolata (Mitt.) Broth. ex Par. 

CAM(47), CAR(47), IVO(1880), LIB(47), 

NIG(47). 

147

148 

Neckeropsis hookeriacea (Dus.) Fleisch. = N. made- 

cassa (787). 

Neckeropsis lepineana (Mont.) Fleisch. 

CAM(787), COM(40), GAB(47), MAD(40), 

MAU(105), REU(105), ROD(105), SAF(787), 

TAN(46), UGA(46), ZAI(65). 

Neckeropsis lepineana var. comorae (C.Müll.) Card. 

= N. lepineana (787). 

Neckeropsis lepineana var. pervilleana (C.Müll.) Card. 

= N. lepineana (787). 

Neckeropsis liliana (Ren.) Broth. ex Par. CAM(47), 

CAR(47), GHA(787), IVO(787,1880), ZAI(787). 

Neckeropsis madecassa (Besch.) Fleisch. 

CAM(47), GAB(47), MAD(40), REU(785), 

ZAI(65). 

Neckeropsis pocsii Enroth & Magill COM(510). 

Neckeropsis spurio-truncata (Dus.) Fleisch. 

CAM(47), CON(47), GAB(47), GUI(47), IVO(47), 

ZAI(1552). 

Neckeropsis spuriotruncata var. latifolia Broth. & 

P.Varde = N. madecassa (787). 

Neckeropsis subtruncata Fleisch., nom. nud. = N. 

disticha (787). 

Neorutenbergia usagarae (Dix.) Bizot & Pócs 

TAN(46). 

Ochrobryum dendeliae Broth. & Par. GUI(47). 

Ochrobryum maclaudii Card. & Par. GUI(47). 

Ochrobryum normandii Card. & Par. CAR(47), 

GUI(47). 

Ochrobryum normandii var. fragilifolium Dix. & 


Ochrobryum normandii var. glaucescens Par. 

GUI(47). 

Ochrobryum obtusissimum Dix. MAL(67), 

TAN(46). 

Ochrobryum sakalavum Card. & Per. MAD(40). 

Octoblepharum africanum (Broth.) Card. 

MAD(40). 

Octoblepharum albidum Hedw. ANG(46), 

ANN(47), BEN(47), BUR(65), CAM(47), 

CBS(1531), CAR(47), COM(40), CON(47), 

GAB(47), GHA(47), GUI(47,1458), 

IVO(63,1458,1880), LIB(47), MAD(40), 

MAL(67), MAU(105,1882), MOZ(46), 

NAT(1531), NIG(47,334), PRI(47), REU(105), 

ROD(105), SAO(47), SEN(47), SEY(40), SIL(47), 

SWA(1531), TAN(46), TOG(58,1878), 

TRV(1531), UGA(46), ZAI(65), ZAM(46,62), 

ZIM(46,61). 

Octoblepharum albidum fo. longifolia Dix. Excluded. 



Octoblepharum brevisetum Townsend = O. albidum 

(762). 

Octoblepharum leptodontium Broth. in Lindau 

TAN(46). 

Octoblepharum leptoneuron Card. SEN(47). 

Octodiceras fontanus Pyl. = F. fontanus (Pyl.) Stend. 

= F. palmifolius (85). 

Oedipodiella australis (Wager & Dix.) Dix. 

CBS(1531), LES(1531), NAT(1531), TRV(1531). 

Oligotrichum afrolaevigatum (Dix.) G.L.Smith 

LES(1531,289), NAT(1531,289). 

Oligotrichum capense Schelpe & Fanshawe 

CBS(1531), NAT(1531). 

Oligotrichum cavallii (Negri) Smith KEN(289), 

RWA(46,65), TAN(46,289), UGA(46,289), 

ZAI(65). 

Oligotrichum tetragonum Schelpe & Fanshawe 

CBS(1531). 

Oligotrichum wageri (Broth.) G.L.Smith 

NAT(1531), OFS(1531,289). 

Oreoweisia erosa (C.Müll.) Kindb. CBS(1531), 

LES(225,1531), MAD(40), NAT(1531), 

OFS(1531), TRV(1531), ZAI(65). 

Orthodontium brevifolium Broth. = O. loreifolium 

(1521). 

Orthodontium gracile Schwaegr. ex BS&G 

CAM(47), ETH(50), KEN(411), RWA(65), 

TAN(46), ZAI(65). 

Orthodontium haplohymenium Dix. & Nav. = O. 

loreifolium (1521). 

Orthodontium lineare Schwaegr. CBS(1531), 

MAL(67), NAT(1531), TAN(146). 

Orthodontium loreifolium Besch. KEN(46), 

MAL(67), REU(105), RWA(65), TAN(46), 

UGA(46), ZAI(65). 

Orthodontium ruwenzorensis (Thér. & Nav.) 

O’Shea UGA(46), ZAI(65). 

Orthostichella pandurifolia (C.Müll.) Buck 

CBS(1531), KEN(46), MAL(67), NAT(1531), 

SWA(1531), TAN(46), TRV(1531), ZIM(61). 

Orthostichella pentasticha (Brid.) Buck CAM(47), 

FER(47), KEN(46), MAD(40), MAL(67), 

REU(105), SIL(47), TAN(46), ZAM(62). 

Orthostichella turgidellacea (C.Müll. ex Dusen) 

Buck CAM(47). 

Orthostichidium involutifolium ssp. involutifolium 

(Mitt.) Broth. ANG(46,2), CAM(47,2), 

CAR(47), FER(47,2), GAB(47), GHA(47,2),

GUI(47,2), IVO(44), LIB(47,2), MAD(40), 

NIG(47,334,2), ROD(105), SAO(47), SIL(47,2), 

ZAI(65). 

Orthostichidium involutifolium ssp. thomeanum 

(Broth.) Argent CAM(47,2), FER(47,2), PRI(47), 

REU(105), SAO(47,2). 

Orthostichopsis debilinervis (Ren. & Card.) Broth. 

REU(105). 

Orthostichopsis longinervis (Ren. & Card.) Broth. 

MAD(40). 

Orthostichopsis madagassa Dix. MAD(40). 

Orthostichopsis pinnatella (Broth.) Broth. 

MAL(67), NAT(1531), TAN(46). 

Orthostichopsis scaberula (Ren. & Card.) Fleisch. 

MAD(40). 

Orthostichopsis subimbricata (Hampe) Broth. 

CBS(1531), COM(40,2), KEN(46), MAD(40,2), 

REU(2,105), TAN(46), TRV(1531), UGA(46). 

Orthostichopsis subimbricata var. borbonica 

(Ren. & Card.) Card. REU(105). 

Orthostichopsis subimbricata var. flageyi (Ren. 

& Card.) Card. MAD(40). 

Orthostichopsis sublivens (Besch.) Broth. 

REU(105). 

Orthotrichum aequatoreum Mitt. KEN(411), 

RWA(65), TAN(46), UGA(46). 

Orthotrichum affine Brid. ETH(347,50), KEN(46), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Orthotrichum afro-fastigiatum C.Müll. Excluded. 

Considered by Lewinsky & van Rooy (1065) to 

be nomen dubium, and replaced by O. incurvo- 

marginatum. 

Orthotrichum arborescens Thér. & Nav. 

KEN(46,136), RWA(65), ZAI(65). 

Orthotrichum armatum Lewinsky & Van Rooy 

CBS(1531,1065). 

Orthotrichum denticulatum Lewinsky 

ETH(347,50), RWA(65), TAN(46). 

Orthotrichum diaphanum Schrad. ex Brid. 

CBS(1531), ETH(347), KEN(46), NAT(1531), 

OFS(1531), SOM(50), TRV(1531). 

Orthotrichum firmum Vent. CBS(1531,1065), 

ERI(50), ETH(50), KEN(46), NAT(1531,1065), 

RWA(65), TAN(46), TRV(1531,1065), UGA(46), 

ZAI(65). 

Orthotrichum incurvomarginatum Lewinsky & 

Van Rooy CBS(1531,1065). 

Orthotrichum latifolium Dix. ex Sappa = O. denticu- 

latum (347). 

Orthotrichum leikipiae C.Müll. = Excluded. See 

Lewinsky (347). 

Orthotrichum leptocarpum Bruch & Schimp. ex 

C.Müll. = O. affine (347). 

Orthotrichum mirum Lewinsky = Stoneobryum mirum 

(1531). 

Orthotrichum mollisimum C.Müll. = Excluded. See 

Lewinsky (347). 

Orthotrichum oreophilum Lewinsky & Van Rooy 

LES(1531,1065). 

Orthotrichum rupestre Schleich. ex Schwaegr. 

CBS(1531), KEN(46), LES(1065,1531), TAN(46), 

UGA(46). 

Orthotrichum simense Schimp. ex Par. = O. firmum 

(347). 

Orthotrichum subexsertum Schimp. ex C.Müll. 

CBS(1531), KEN(46), NAT(1531), RWA(65). 

Orthotrichum transvaaliense Rehm. ex Sim 

NAT(1531), OFS(1531), TRV(1531). 

Orthotrichum undulatifolium C.Müll. ETH(50), 

TAN(46), ZAI(65). 

Oxyrrhynchium confervoidum Sim CBS(1531), 

NAT(1531). 

Oxyrrhynchium kenyae Dix., nom. nud. ETH(50). 

Oxyrrhynchium petronelli Tosco & Piovano 

KEN(46). 

Oxyrrhynchium porothamnioides P.Varde & 

Levier ZAI(65). 

Oxyrrhynchium ruscifome Warnst. = Platyhypnidium 

riparioides (1521). 

Oxyrrhynchium subasperum Sim CBS(1531), 

NAT(1531), TRV(1531). 

Oxyrrhynchium swartzii (Turn.) Warnst. 

CAM(47). 

Oxystegus cylindricus (Brid.) Hilp. = Trichostomum 

tenuirostre (1420). 

Oxystegus rhodesiae (Brid.) Hilp. = Trichostomum 

rhodesiae (1420). 

Oxystegus soulae (C.Müll. in Ren. & Card.) Wijk & 

Marg. = Trichostomum soulae (1420). 

Oxystegus tenuirostris var. tenuirostris (Hook. & 

Tayl.) AJE Smith = Trichostomum tenuirostre 

(1420). 

Palamocladium nilgheriense (Mont.) C.Müll. 

COM(40), MAD(40), REU(105,1626), RWA(65), 

SOC(40,1626), SOM(1626). 

Palamocladium schoanum C.Müll., nom. nud. 

ETH(50). 

Palamocladium sericeum (Jaeg.) C.Müll. 

CBS(1531), CAP(1296), ETH(50), KEN(46), 

LES(1531), NAT(1531), RWA(47,65), 

149

150 

SWA(1531), TAN(46), TRV(1531), UGA(46), 

ZAI(47,65). 

Palamocladium subsericeus (Broth. & Par.) Broth. 

DJI(50), ETH(50). 

Papillaria acinacifolia Besch. MAU(105). 

Papillaria africana (C.Müll.) Jaeg. ANG(46), 

CBS(1531), COM(40), KEN(46), MAD(40), 

MAL(67), MAU(105), NAT(1531), REU(105), 

SWA(1531), TAN(46), TRV(1531), UGA(46), 

ZAM(411), ZIM(46,61). 

Papillaria africana fo. pinnata Dix. ex Tosco & 



Papillaria africana var. flagellacea Biz. TAN(46). 

Papillaria africana var. gracilis (Ren. & Card.) 

Wijk & Marg. MAD(40). 

Papillaria bazzaccoi Tosco & Piovano TAN(46). 

Papillaria borchgrevinkii Kiaer MAD(40), 

TAN(46). 

Papillaria breviculifolia C.Müll. TAN(46). 

Papillaria filifunalis C.Müll. KEN(46), TAN(46). 

Papillaria flaccidula Card. MAD(40). 

Papillaria perichaetialis (Hampe) Jaeg. 

MAD(40). 

Papillaria renauldii Besch. MAU(105). 

Papillaria serpentina C.Müll. TAN(46). 

Paraleucobryum longifolium ssp. brasiliense 

(Broth.) P.Müller & Frahm ZAI(65). 

Pelekium velatum Mitt. TAN(46). 

Phasconica tisserantii P.Varde = Trichostomum un- 

guiculatum (1420). 

Phascum kilimandscharicum (C.Müll.) Broth. = 

Bryum kilimandscharicum (C.Müll.)Zander 

(1420). However, there is already a Bryum 

kilimandscharicum Broth., which is a synonym 

of Bryum algovicum var. rutheanum (23). I have 

thus provisionally excluded this species, recorded 

for Tanzania by Kis (46). 

Phascum leptophyllum C.Müll. = Chenia leptophylla 

(1420). 

Phascum peraristatum C.Müll. CBS(1531), 

KEN(411). 

Philonotis africana (C.Müll.) Rehm. ex Par. 

CBS(1531), MAL(67), NAT(1531), SWA(1531), 

TRV(1531), ZAM(46,62), ZIM(61). 

Philonotis amblystegioides Demar. & P.Varde 

ZAI(65). 

Philonotis androgyna (Hampe) Jaeg. TAN(46), 

ZIM(46). 

Philonotis angustiretis Broth., nom. nud. 

UGA(46). 

Philonotis annobonensis Broth. in Mildbr. 

ANN(47). 

Philonotis bescherellei Thér., nom. nud. REU(105). 

Philonotis brevicuspis Broth. CAM(47), CAR(47), 

ZAI(65,1552). 

Philonotis brevicuspis var. perobtusata P.Varde 

CAR(47). 

Philonotis byssiformis C.Müll. MAD(40). 

Philonotis cernua (Wils.) Griffin & Buck CAM(47), 

FER(47). 

Philonotis comosa (Broth.) Griffin & Buck 

CBS(1531). 

Philonotis curvula (C.Müll.) Kindb. KEN(46). 

Philonotis dregeana (C.Müll.) Jaeg. BOT(1531), 

CBS(1531), CAR(47), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), SWA(1531), 

TRV(1531), ZAI(65), ZIM(61). 

Philonotis falcata (Hook.) Mitt. BOT(1531), 

CBS(1531), LES(1531), MAL(67), NAM(1531), 

NAT(1531), OFS(1531), RWA(65), TRV(1531), 

ZAI(65), ZAM(62), ZIM(61). 

Philonotis flaccidifolia (Mitt.) Par. ROD(105). 

Philonotis flavinervis (C.Müll.) Kindb. FER(47). 

Philonotis fontana (Hedw.) Brid. ETH(50), 

KEN(46), STH(1296), TAN(46), UGA(46), 

ZAI(65). 

Philonotis fugacissima Par. GUI(47). 

Philonotis gemmascens (C.Müll.) Kindb. TAN(46), 

ZAI(65). 

Philonotis globosa (C.Müll.) Griffin & Buck 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TAN(46), TRV(1531), UGA(46), ZAM(46), 

ZIM(46,61). 

Philonotis gracilescens Schimp. MAU(105), 

REU(105). 

Philonotis hastata (Duby) Wijk & Marg. ANG(46), 

BOT(1531), BUF(47), CAM(47), CBS(1531), 

CAR(47), COM(40), GAB(47), GHA(47), 

IVO(47), KEN(46), LES(1531), MAD(40), 

MAL(67), MOZ(357), NAT(1531), NIG(334), 

OFS(1531), REU(105), RWA(65), SIL(47), 

SWA(1531), TAN(46), TOG(1878), TRV(1531), 

UGA(46), ZAI(65,1552), ZIM(46,61). 

Philonotis heleniana (Mitt.) Herz. = P. fontana 

(1521). 

Philonotis helenica (Besch.) Par. STH(1296). 

Philonotis heterophylla Mitt. MAD(40). 

Philonotis incrassata (C.Müll.) Kindb. ANG(46). 

Philonotis jungneri Broth. CAM(47), ZAI(65).

Philonotis laeviuscula Dix. ZIM(46). 

Philonotis le-testui (P.Varde) Griffin & Buck 

CAR(47). 

Philonotis marangensis Broth. KEN(46), 

TAN(46), ZAI(65). 

Philonotis marchica (Hedw.) Brid. SUD(1419). 

Philonotis mauritiana Aongstr. MAD(40), 

MAU(105), REU(105), TAN(46), UGA(46). 

Philonotis mauritiana var. gemmiclada Broth. 

ZAI(65). 

Philonotis mauritiana var. stricta Ren. & Card. 

MAD(40). 

Philonotis microthamnia Broth. CAM(47), 

CAR(47), CON(47), GAB(47), IVO(1880), 

KEN(46), UGA(46), ZAI(65). 

Philonotis mnelabiensis P.Varde GAB(47), 

ZAI(65). 

Philonotis mniobryoides Broth. CAM(47), 

CAR(47), GAB(47), IVO(1458), TOG(1878), 

ZAI(65). 

Philonotis mniobryoides var. angustifolia Thér. 

& P.Varde GAB(47), GHA(47), ZAI(65). 

Philonotis nanothecia (C.Müll.) Kindb. BEN(47), 

CAR(47), SAO(47), TOG(1878), ZAI(65). 

Philonotis nanothecioidea Par. & Broth. CAR(47), 

GAB(47), GUI(47). 

Philonotis obtusata C.Müll. ex Ren. & Card. 

KEN(46), TAN(46). 

Philonotis pechuelii (C.Müll.) Kindb. CAR(47), 

ZAI(65). 

Philonotis penicillata Wright ASC(1296). 

Philonotis perconferta Broth. CAM(47). 

Philonotis perigonialis Besch. REU(105). 

Philonotis perigonialis fo. procerior Besch. Excluded. 



Philonotis perpusilla Card. MAD(40). 

Philonotis perrieri Thér. MAD(40). 

Philonotis plana Dix. & Herz. UGA(46). 

Philonotis platyneura P.Varde TAN(46), 

UGA(46). 

Philonotis pungens (Mitt.) Mitt. ROD(105), 

SOC(40). 

Philonotis rigida Brid. CAP(1296). 

Philonotis ruwenzorensis Thér. & Nav. TAN(46), 

UGA(46), ZAI(65). 

Philonotis scabrifolia (Hook.f. & Wils.) Braithw. 

CBS(1531), LES(1531), NAT(1531). 

Philonotis schroederi Broth. TAN(46). 

Philonotis soulii Thér. & P.Varde KEN(46). 

Philonotis sparsifolia (Hampe) Jaeg. KEN(46), 

MAD(40). 

Philonotis speirophylla Dix. KEN(46). 

Philonotis spongiosa (Welw. & Duby) Gepp 

ANG(46). 

Philonotis strictula Card. CAM(47), CAR(47), 

GAB(47), IVO(1458), SIL(47), UGA(46), 

ZAI(65). 

Philonotis submarchica Besch. REU(105). 

Philonotis submarchica var. plumosa Ren. & 

Card. REU(105). 

Philonotis subolescens (C.Müll.) Par. ASC(1296). 

Philonotis subsimplex Broth. & Par. MAI(47). 

Philonotis tenuis Tayl. = P. fontana var. capillaris 

(1521). 

Philonotis tomentella Mol. KEN(46), RWA(65), 

TAN(46), ZAI(65). 

Philonotis trichodonta (C.Müll.) Kindb. SAO(47). 

Philonotis tricolor (C.Müll.) Kindb. TAN(46). 

Philonotis usambarica Broth. TAN(46). 

Philonotis vagans (Hook.f. & Wils.) Mitt. 

CBS(1531). 

Philonotis vanderystii Card. CAM(47), CAR(47), 

ZAI(65). 

Phyllodrepanium falcifolium (Schwaegr.) Crosby 

REU(105). 

Phyllogonium chevalieri Corb. = Calyptothecium 

planifrons (1). 

Phyllogonium fulgens (Hedw.) Brid. 

REU(348,105). 

Phyllogonium viride Brid. REU(348). 

Phyllogonium viscosum (P.Beauv.) Mitt. 

MAD(40), MAU(105), REU(348,105,1881), 

SAF(348). 

Phyllogonium viscosum fo. viride Besch. = P. viride 

(348). 

Physcomitrella patens ssp. magdalenae (De 

Sloover) Tan RWA(65). 

Physcomitrellopsis africana Wag. & Broth. ex 

Dix. CBS(1531), NAT(1531), TAN(46). 

Physcomitrium aubertii Besch. MAU(105). 

Physcomitrium bukobense (Broth.) Fife TAN(46). 

Physcomitrium coarctatum Card. MAD(40). 

Physcomitrium dilatatum Ren. & Card. MAD(40). 

Physcomitrium eurystomum Sendtn. KEN(46), 

MAL(67), TAN(46), UGA(46). 

Physcomitrium flexifolium Mitt. STH(1296). 

Physcomitrium mirabile (Broth. & Par.) Zander 

GUI(47). 

Physcomitrium niloticum (Del.) C.Müll. MOZ(46), 

151

152 

NAT(1531), SUD(1707), ZIM(46,61). 

Physcomitrium spathulatum var. sessile (Shaw) 

Magill CBS(1531). 

Physcomitrium spathulatum var. spathulatum 

(Hornsch.) C.Müll. CBS(1531), KEN(46), 

MOZ(46), NAM(1531), NAT(1531), TRV(1531), 

ZIM(46). 

Physcomitrium subspathulatum Thér. & Nav. 

RWA(65), ZAI(65). 

Pilopogon africanus Broth. BUR(65), KEN(411), 

RWA(65), TAN(46), ZAI(65). 

Pilopogon africanus var. kivuensis P.Varde & 


Pilotrichella ampullacea (C.Müll.) Jaeg. 

COM(40), KEN(46), MAL(46,67), RWA(65), 

TAN(46), UGA(46), ZAI(65), ZAM(46). 

Pilotrichella attenuata Broth. TAN(46). 

Pilotrichella calomicra Broth. CAM(47), GUI(47), 

SAO(47). 

Pilotrichella capillicaulis Kindb. KEN(46). 

Pilotrichella communis C.Müll. ex Dus. CAM(47), 

CON(47), FER(47), GAB(47), GHA(47), GUI(47), 

IVO(47), SAO(47), SIL(47), TOG(1878). 

Pilotrichella cordata Broth. in P.Varde, nom. 

nud. KEN(46), TAN(46). 

Pilotrichella curvifrons Kindb. KEN(46). 

Pilotrichella cuspidata Broth. = P. panduraefolia 

(1519) (= Orthostichella pandurifolia). 

Pilotrichella cuspidata var. gracilescens Tosco 

& Piovano KEN(46). 

Pilotrichella cuspidata var. gracilescens Tosco & 

Piovano. The owning species of this variety has 

been synonymised with Orthostichella 

pandurifolia. This variety also needs revision. 

Pilotrichella flexilis (Hedw.) Jaeg. MAS(40). 

Pilotrichella gracilicaulis C.Müll. ex Dus. 

CAM(47). 

Pilotrichella grimaldii Ren. & Card. MAD(40), 

TAN(46). 

Pilotrichella guineensis Aongstr. in C.Müll. 

GUI(47). 

Pilotrichella holstii Broth. MOZ(46), TAN(46). 

Pilotrichella imbricatula C.Müll. ETH(50). 

Pilotrichella incurva Broth. ANG(46). 

Pilotrichella inflatifolia C.Müll. SAO(47). 

Pilotrichella isleana Besch. KEN(46), MAD(40), 

MAL(46,67), REU(105), TAN(46). 

Pilotrichella isleana var. nervosa (Ren. & Card.) 


Pilotrichella isleana var. virescens Besch. 

MAS(40). 

Pilotrichella latiramea C.Müll. ex Dus. CAM(47), 

CAR(47), GAB(47), IVO(1880), KEN(46), 

TAN(46). 

Pilotrichella leptoclada C.Müll. CON(47), 

SAO(47). 

Pilotrichella mascarenica (C.Müll.) Jaeg. 

MAD(40), REU(105). 

Pilotrichella moenkemeyeri (C.Müll.) Kindb. 

NIG(47). 

Pilotrichella muelleri Dus. CAM(47). 

Pilotrichella panduraefolia (C.Müll.) Jaeg. = Ortho- 

stichella pandurifolia (1714). 

Pilotrichella pentasticha (Brid.) Wijk & Marg. = 

Orthostichella pentasticha (1714). 

Pilotrichella percordata Broth. in P.Varde, nom. 

nud. KEN(46), TAN(46). 

Pilotrichella perrobusta P.Varde MAD(40), 

TAN(46). 

Pilotrichella phleoides (Brid.) Jaeg. REU(105). 

Pilotrichella profusicaulis Par. KEN(46), 

TAN(46), ZAI(65). 

Pilotrichella pseudo-imbricata (C.Müll.) Jaeg. 

COM(40), MAD(40). 

Pilotrichella pseudo-imbricata var. pallescens 

(C.Müll.) Par. COM(40). 

Pilotrichella pseudo-imbricata var. rufescens 

(C.Müll.) Par. COM(40). 

Pilotrichella pseudoimbricata Jaeg. MAL(67). 

Pilotrichella ragazzii Brizi ETH(50). 

Pilotrichella sericea Kindb. KEN(46), TAN(46). 

Pilotrichella sordido-viridis C.Müll. ex Dus. 

CAM(47), CAR(47), CON(47), GAB(47), 

GHA(47), IVO(1458,1880), LIB(47), NIG(47). 

Pilotrichella stuhlmannii Broth. KEN(46), 

TAN(46). 

Pilotrichella subpanduraefolia Kindb. CAM(47). 

Pilotrichella tenellula Par. TAN(46). 

Pilotrichella turgidellacea C.Müll. ex Dus. = Ortho- 

stichella turgidellacea (1714). 

Pilotrichella welwitschii (Duby) Gepp ANG(46). 

Pilotrichum denticulatum P.Beauv. REU(105). 

Pinnatella africana (Welw. & Duby). Broth. = P. 

minuta (1830). 

Pinnatella allamanoi Tosco & Piovano = Porotrichum 

molliculum (1300). 

Pinnatella braunii (Broth.). Fleisch. = 

Homaliodendron piniforme (1300). 

Pinnatella chalaropteris (Dus.). Broth. = Homalio- 

dendron piniforme (1300).

Pinnatella dupuisii (Ren. & Card.). Broth. = P. 


Pinnatella engleri (Broth.). Fleisch. = P. minuta 

(1830). 

Pinnatella flagellacea (Mitt.). Broth. ex Par. = .Ho- 

maliodendron piniforme (1300) 

Pinnatella fuciformis (Brid.) Touw (“fuciforme”) = 

Leptodon fuciformis (301). 

Pinnatella geheebii (C.Müll.). Fleisch. = P. minuta 

(1830). 

Pinnatella le-testui Thér. & P.Varde = P. minuta 

(1830). 

Pinnatella mariei (Besch.) Broth. = Caduciella mariei 

(780). 

Pinnatella mayumbensis (Besch.). Fleisch. = Homa- 

liodendron piniforme (1300). 

Pinnatella minuta (Mitt.) Broth. ANG(46), 

CAM(47), CAR(47), COM(40), CON(47), 

EGU(1830), GAB(47), GUI(47), IVO(44,1880), 

KEN(46), MAD(40), MAU(1830), NAT(1531), 

PRI(1830), REU(1830), SAO(47), TAN(46), 

UGA(46), ZAI(65). 

Pinnatella mucronata (Bosch & Sande Lac.) 

Fleisch. SEY(1830). 

Pinnatella oblongifrondea (Broth.). Broth. = P. 


Pinnatella pechuelii (C.Müll.). Broth. = P. minuta 

(1830). 

Pinnatella tamariscina (Hampe). Broth. = Homalio- 

dendron piniforme (1300). 

Pinnatella usagara (Mitt.). Broth. = Porotrichum 

usagarum (299). 

Pinnatella vignolo-lutatii Tosco & Piovano, nom. 

inval. = Porotrichum elongatum (1830). 

Pireella filicina (Hedw.) Card. Excluded. It is unlikely 

that this neotropical taxon occurs in Africa (A.E. 

Newton, pers. comm.), and this 1827 record from 

Réunion (105) is thus suspect. It appears in Bridel 

(77) amongst various neotropical records. 

Plagiobryum laxum Demar. & P.Varde ZAI(65). 

Plagiobryum piliferum P.Varde TAN(46). 

Plagiobryum zierii (Hedw.) Lindb. LES(1531), 

NAT(1531). 

Plagiomnium cuspidatum (Hedw.) Kop. 

KEN(411), UGA(46,1036). 

Plagiomnium rhynchophorum (Hook.) T.Kop. 

CAM(47), ETH(50,1036), FER(47), KEN(46), 

MAD(40), MAL(67), REU(105), RWA(65), 

SAO(47), TAN(46), UGA(46), ZAI(65), 

ZAM(62), ZIM(46,61). 

Plagiomnium rhynchophorum var. reidii (Dix.) 

T.Kop. CBS(1036,1531), KEN(46,1036), 

LES(1036,1531), NAT(1036,1531), OFS(1531), 

RWA(65,1036), TAN(46,1036), 

TRV(1036,1531), ZAI(65). 

Plagiomnium rostratum (Schrad.) Kop. auct. afr. = P. 

rhynchophorum (1036). 

Plagiomnium undulatum (Hedw.) Kop. 

ETH(1336). 

Plagiopus oederi (Brid.) Limpr. LES(1074,1531), 

NAT(1531). 

Plagiothecium capense Schimp. ex Jaeg. ETH(50). 

Plagiothecium corticola Aongstr. = Sematophyllum 

corticolum (See “New Combinations”). 

Plagiothecium denticulatum var. denticulatum 

BS&G MAS(40). 

Plagiothecium membranosulum C.Müll. 

CBS(1531), NAT(1531), TRV(1531). 

Plagiothecium mildbraedii Broth. RWA(46,65), 

TAN(46), UGA(46), ZAI(65). 

Plagiothecium nitens Dix. KEN(46), RWA(65), 

TAN(46), ZAI(65). 

Plagiothecium nitidifolium (Mitt.) Jaeg. FER(47), 

RWA(65), TAN(46). 

Plagiothecium rhynchostegioides C.Müll. 

CBS(1531), NAT(1531), TRV(1531), ZIM(46). 

Platyhypnidium afro-rusciforme (Broth.) Fleisch. 

CAM(47). 

Platyhypnidium hedbergii (P.Varde) Ochyra & 

Sharp KEN(46), TAN(46), ZAI(65). 

Platyhypnidium macowanianum (Par.) Fleisch. 

TRV(1531), ZIM(46). 

Platyhypnidium riparioides (Hedw.) Fleisch. 

KEN(46), RWA(65), TAN(46). 

Pleuridium acuminatum Lindb. MAL(67). 

Pleuridium ecklonii (C.Müll.) Snider CBS(1531), 

NAT(1531). 

Pleuridium nervosum (Hook.) Mitt. CBS(1531), 

LES(1531), MAD(40), NAT(1531), OFS(1531). 

Pleuridium papillosum Magill CBS(1531). 

Pleuridium pappeanum (C.Müll.) Jaeg. CBS(1531), 

LES(1531), ZIM(61). 

Pleurochaete beccarii Vent. ERI(50), SOM(50), 

ZAI(65). 

Pleurochaete beccarii Vent. Zander (1420) reports 

that African material identified as this taxon at 

BM and NY is P. squarrosa var. squarrosa, and 

that type material of P. beccarii at TR could not 

be obtained on loan, but is probably synonymous. 

Pleurochaete malacophylla (C.Müll.) Broth. 

153

154 

ERI(50). 

Pleurochaete malacophylla (C.Müll.) Broth. Zander 

(1420) states that this taxon is “doubtfully distinct 

from P. squarrosa var. squarrosa”. 

Pleurochaete squarrosa (Brid.) Lindb. CAP(1296), 

DJI(1569), ETH(50), KEN(46), SOC(349), 

TAN(46). 

Pleuropus bonplandii Hook. = Palamocladium les- 

keoides (1521). 

Pleuropus sericeus (C.Müll.) Broth. = Palamocladium 

sericeum (1521). 

Pocsiella hydrogonioides Biz. TAN(46). 

Pogonatum aloides (Hedw.) P.Beauv. Excluded. All 

African material examined by Hyvönen had been 

misidentified (56). 

Pogonatum belangeri (C.Müll.) Jaeg. BUR(56,65), 

CAM(56), KEN(56), MAD(56), MAL(67), 

MAU(56), REU(56), RWA(56,65), SAF(56), 

TAN(56), UGA(56), ZAI(56,65), ZIM(61). 

Pogonatum borgenii (Hampe) Jaeg. = P. capense 

(56). 

Pogonatum brachythecium Besch. = P. belangeri 

(56). 

Pogonatum brachythecium var. hildebrandtii Thér. 

= P. belangeri (56). 

Pogonatum brachythecium var. madagassum Card. 

= P. belangeri (56). 

Pogonatum capense (Hampe) Jaeg. ANG(56,289), 

CBS(1531), MAD(56,40,289), MAU(289), 

NAT(1531,289), SWA(56,1531), 

TRV(1531,289), ZIM(61,289). 

Pogonatum congolense Card. CAM(56,289,47), 

GAB(56), ZAI(56,65,289). 

Pogonatum convolutum (Hedw.) P.Beauv. 

MAD(56,40,289), MAU(56,289), REU(56,289). 

Pogonatum convolutum var. madagassum Card. = P. 

convolutum (56). 

Pogonatum flexuosum Mitt. Excluded. Hyvönen 

(56) states that the only specimen of the taxon 

is mislabelled, and is not from Africa, but from 

Borneo. It is now a synonym of P. cirratum var. 

macrophyllum (Dozy & Molk.) Hyvönen. 

Pogonatum gracilifolium Besch. ANG(289), 

BUR(65), CAM(56,289), COM(40), 

ETH(56,289), FER(289), KEN(56,289), LIB(289), 

MAD(40), MAL(56,67,289), MAU(289), 

REU(56), RWA(56,65,289), SAO(56,289), 

TAN(56,289), TRV(289), UGA(56,289), 

ZAI(56,65,289). 

Pogonatum macei Thér. = P. perichaetiale subsp. 

oligodus (56). 

Pogonatum marginatum Mitt. MAU(289). 

Pogonatum marginatum Mitt. This species is said 

by Hyvönen (56) to occur only in SE Asia, and is 

shown as such in his distribution maps, but he 

quotes De Sloover’s illustration (289), which is of 

a plant collected in Mauritius. A similar disjunction 

is shown by P. proliferum (Griff.) Mitt. 

Pogonatum molleri (C.Müll.) Par. = P. gracilifolium 

Besch (56). 

Pogonatum oligodus (C.Müll.) Mitt. = P. perichaetiale 

ssp. oligodus (56). 

Pogonatum perichaetiale ssp. oligodus (C.Müll.) 

Hyvönen BUR(56,65,289), CAM(56,289), 

COM(56,40), ETH(56,50,289), KEN(56,289), 

LES(56,1531,289), NAT(1531,289), OFS(1531), 

REU(56,289), RWA(56,65,289), TAN(56,289), 

TRV(1531,289), UGA(56,289). 

Pogonatum proliferum (Griff.) Mitt. REU(758,105), 

RWA(56,65,289), UGA(56,289). 

Pogonatum rubenti-viride (C.Müll.) Par. = P. graci- 

lifolium Besch (56). 

Pogonatum rutenbergii (Geh.) Par. = P. convolutum 

(56). 

Pogonatum semilamellatum (Leroy) = P. congolense 

Card (56). 

Pogonatum simense (Bruch & Schimp. ex C.Müll.) 

Jaeg. = P. perichaetiale ssp. oligodus (56). 

Pogonatum therioti Demar. & Leroy = P. belangeri 

(56). 

Pogonatum ugandae P.Varde = P. proliferum (56). 

Pogonatum urnigerum (Hedw.) P.Beauv. 

CAM(56,289,47), KEN(289,46), 

RWA(56,65,289,46), TAN(56,289,46), 

UGA(56,46), ZAI(56,65,289). 

Pogonatum usambaricum (Broth.) Par. KEN(56), 

REU(56), RWA(56), SAF(56), SAO(56), 

TAN(56), ZAI(56). 

Pogonatum waterlotii Thér. = P. belangeri (56). 

Pohlia afro-cruda (C.Müll.) Broth. KEN(46), 

TAN(46), UGA(46), ZAI(65). 

Pohlia aloysii-sabaudiae Negri UGA(46). 

Pohlia baronii Wijk & Marg. KEN(46), LES(1531), 

MAD(40), MAL(67), NAT(1531), TRV(1531), 

UGA(46), ZAI(65). 

Pohlia bequaertii (Dix. & Nav.) Shaw UGA(46), 

ZAI(65). 

Pohlia chrysoblasta (Thér. & Nav.) Demar. 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Pohlia cratericola Broth. CAM(47), TAN(46),

ZAI(65). 

Pohlia cruda (Hedw.) Lindb. LES(1531), 

NAT(1531). 

Pohlia elongata var. elongata Hedw. CBS(1531), 

KEN(46), LES(1531), MAD(40), MAL(67), 

NAT(1531), OFS(1531), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZIM(61). 

Pohlia grammophylla (Besch.) Broth. COM(40). 

Pohlia gromieri (Thér.) Kis KEN(46). 

Pohlia heribaudii var. madagassa (Ren. & Card.) 

Wijk & Marg. MAD(40). 

Pohlia kenyae (Dix.) Kis KEN(46). 

Pohlia lacouturei (Thér.) O’Shea MAD(40). 

Pohlia longicollis (Hedw.) Lindb. KEN(46), 

TAN(46). 

Pohlia longicollis var. alpina (Hoppe & Hornsch.) 

Will. TAN(46). 

Pohlia mielichhoferiacea (C.Müll.) Broth. 

KEN(146), RWA(65), TAN(46), ZAM(62). 

Pohlia nutans (Hedw.) Lindb. CBS(1531), 

TAN(46), ZIM(61). 

Pohlia philonotula (C.Müll.) Broth. KEN(46), 

TAN(46), ZAM(62). 

Pohlia sp. ETH(50) 

Webera cavifolia Bruch. & Schimp (in sched.) ex Par., 

nom. nud. This taxon cannot be usefully 

transferred to its correct genus (Pohlia) without 

its status being investigated. Paris mentions a 

specific collection of Schimper [459], which it 

was not possible to locate at BM. If it is likely 

that a specimen exists, this taxon should remain 

here until described or synonymised. 

Pohlia sp. ETH(50) 

Webera philibertii Par., nom. nud. This taxon is listed 

by Paris, but a specific collection is not mentioned. 

Until its status is confimed, this taxon cannot be 

transferred to Pohlia, or synonymised with 

another taxon. If a specimen is not found, it 

should be excluded. 

Polytrichastrum alpinum (Hedw.) Smith ZAI(65). 

Polytrichastrum formosum (Hedw.) G.L.Smith 

CBS(1531), LES(1635), MAD(40), MAU(289), 

TRV(289). 

Polytrichum appressum Brid. = P. subpilosum (289). 

Polytrichum commune var. commune Hedw. 

ANG(46,289), BUR(65,289), CAM(47,289), 

CBS(1531,289), COM(40), ETH(50), FER(47), 

KEN(46), LES(1531), MAD(40,289), MAL(67), 

MAU(289), MOZ(46,289), NAT(1531), 

NIG(47), OFS(1531,289), REU(289), 

RWA(65,289), SIL(47), SWA(1531), 

TAN(46,289), TRV(1531,289), UGA(46), 

ZAI(65,289), ZAM(289), ZIM(46). 

Polytrichum commune var. humile Sw. ZAI(65). 

Polytrichum commune var. perigoniale (Michx.) 

Hampe ANG(46). 

Polytrichum commune var. trichodes (C.Müll.) 

Dix. ZIM(46). 

Polytrichum cupreum Negri UGA(46). 

Polytrichum elegans Welw. & Duby ANG(46). 

Polytrichum glabrum Schrad. MAD(40), 

MAS(40). 

Polytrichum integrum C.Müll. = P. juniperinum 

(1521). 

Polytrichum juniperellum Geh. MAD(40). 

Polytrichum juniperinum Hedw. CAM(47), 

CBS(1531,289), LES(1531), OFS(1531), 

STH(1296). 

Polytrichum keniae Dix. = Pogonatum urnigerum 

(289). 

Polytrichum madagassum (Hampe) Jaeg. = P. 

commune (289). 

Polytrichum piliferum Hedw. CBS(1531,289), 

COM(289), KEN(46,289), LES(1531,289), 

MAL(67,289), RWA(46,65,289), TAN(46,289), 

UGA(46,289), ZAI(65,289), ZAM(46), ZIM(61). 

Polytrichum piliferum var. australe Ren. & Card. 

COM(40). 

Polytrichum preussii Broth. = P. subpilosum (289). 

Polytrichum pungens C.Müll. = P. subpilosum (289). 

Polytrichum robustum Card., hom. illeg. = P. subpi- 

losum (289). 

Polytrichum subappressum Besch.. = P. subpilosum 

(289). 

Polytrichum subattenuatum Thér. = Polytrichastrum 

formosum (289). 

Polytrichum subformosum Besch. = P. subpilosum 

(289). 

Polytrichum subformosum var. anomalum Thér. = 

P. commune (289). 

Polytrichum subformosum var. parvisetum 

(C.Müll.) Card. MAD(40). 

Polytrichum subformosum var. parvisetum (C.Müll.) 

Card. The owning species of this variety, and 

other varieties of this species, have all been 

transferred to other names (289). This variety 

also needs revision. 

Polytrichum subformosum var. pervillei (Besch.) 

Card. in Grand. = P. subpilosum (289). 

Polytrichum subpilosum P.Beauv. BUR(65), 

155

156 

CAM(47), CBS(1531), COM(40), KEN(46), 

MAD(40), MAL(67), MAU(105), REU(105), 

RWA(65), SEY(40), SWA(1531), TAN(46), 

TRV(1531), UGA(46), ZAI(65), ZAM(62), 

ZIM(61). 

Polytrichum trichobasis C.Müll., nom. nud. 

MAD(40). 

Polytrichum tristanii Duby = P. juniperinum (1521). 

Porothamnium comorense (C.Müll.) Sim. = Porotri- 

chum elongatum (15). 

Porothamnium comorense var planissimum (Dus.) 

Wijk & Marg. = P. stipitatum (15). 

Porothamnium comorense var. scioanum (Tong.) 

Cuf. = P. stipitatum (15). 

Porothamnium leptopteris (Dusen.) Fleisch. = P. 

stipitatum (15). 

Porothamnium minutum (Broth. & Par.) Broth. = 

Porotrichum caudatum (15). 

Porothamnium molliculum (Broth.) Fleisch. = Poro- 

trichum molliculum (15). 

Porothamnium pennaefrondeum (C.Müll.) Card. = 

Porotrichum madagassum (15). 

Porothamnium punctulatum C.Müll. ex Dus. = Poro- 

trichum caudatum (15). 

Porothamnium stipitatum (Mitt.) Touw ex De 

Sloover ANG(46,15), BUR(15,65), CAM(15), 

CBS(1531), COM(15), CON(15), ETH(15), 

FER(47,15), GAB(15), GHA(15), GUI(15), 

IVO(44), KEN(46,15), MAD(15), MAL(15,67), 

MAU(105), MOZ(357), NAT(1531), 

REU(15,105), RWA(15,65), SAO(15), 

SWA(1531), TAN(46,15), TRV(1531), 

UGA(46,15), ZAI(15,65), ZAM(46,62,15), 

ZIM(46,15). 

Porothamnium variifolioides De Sloover FER(15), 

KEN(411), MAD(15), NIG(1492), REU(15), 

RWA(65,15), TAN(46,15), ZAI(65,15). 

Porothamnium viguieri Thér. = P. molliculum (15). 

Porothamnium viguieri var. laxirete Demar. & Leroy 

= Porotrichum molliculum (15). 

Porotrichum annobonense Broth. in Mildbr., 


Porotrichum cameruniae Dix. = P. quintasii (15). 

Porotrichum caudatum Broth. CAM(47,15), 

CAR(15), ETH(15), GAB(15), GHA(15), 

GUI(15,1458), IVO(411,1880), NIG(15), 

SAO(47,15), SEN(15), TAN(46,15), 

TOG(58,15,1878), ZAI(65,15). 

Porotrichum dentatum Gepp MAL(1849). 

Porotrichum elongatum (Welw. & Duby) Gepp 

ANG(46,15), ANN(15), BUR(65,15), 

CAB(46,15), CAM(15), CAR(15), COM(15), 

CON(15), ETH(15), FER(15), GAB(15), GUI(15), 

IVO(15,1880), KEN(46,15), LIB(15), MAD(15), 

MAL(46,67), MAU(15), NAT(1531), NIG(15), 

REU(15), RWA(65), SAO(15), SEY(15), 

TAN(46,15), TOG(1878), TRV(1531), 

UGA(46,15), ZAI(65,15), ZIM(46,61). 

Porotrichum jollyi Broth. & Par. = P. elongatum 

(15). 

Porotrichum jollyi var. elongatum P.Varde = P. 

elongatum (15). 

Porotrichum laurentii Ren. & Card. = P. elongatum 

(15). 

Porotrichum longirostre (Hook.) Mitt. CAM(47). 

Porotrichum madagassum Kiaer ex Besch. 

CAM(15), CBS(1531), COM(15), ETH(15), 

KEN(46,15), LES(15), MAD(15), MAL(15,67), 

MAU(15), MOZ(46,15), NAT(1531), 

OFS(1531), REU(15), RWA(65,15), TAN(46,15), 

TRV(1531), UGA(46,15), ZAI(65,15), 

ZIM(46,15). 

Porotrichum molliculum Broth. ANG(46,15), 

BUR(15,65), CAB(46), CAM(15), CBS(1531), 

COM(15), GAB(15), KEN(46,15), MAD(15), 

MAL(46,67), MAU(15), NAT(1531), REU(15), 

RWA(15,65), TAN(46,15), TRV(1531), 

UGA(46,15), ZAI(15,65), ZIM(61). 

Porotrichum pennaefrondeum C.Müll. = P. mada- 

gassum (15). 

Porotrichum pobeguinii (Par. & Broth.) Par. = P. 

elongatum (15). 

Porotrichum punctulatum C.Müll. ex Dus. = P. cau- 

datum (15). 

Porotrichum quintasii Broth. CAM(15), CAR(15), 

CON(15), GAB(15), GUI(15), LIB(15), SAO(47). 

Porotrichum usagarum Mitt. TAN(785,46). 

Potamium sanctae-mariae (Besch.) Broth. 

MAD(40). 

Pottia elatior (C.Müll.) C.Müll. = P. pusilla var. 

elatior (1521) (= Tortula minor var. elatior). 

Pottia macowanianum C.Müll. = Microbryum daval- 

lianum var. conicum (1420). 

Pottia minor (C.Müll.) Cuf. = Tortula minor (1420). 

Pottia mirabilis Broth. & Par. = Physcomitrium 

mirabile (1420). 

Pottia namaquensis Magill CBS(1531). 

Pottia pusilla var. elatior (C.Müll.) Warnst. = Tortula 

minor var. elatior (1420). 

Pottia pusilla var. pusilla Warnst., hom. illeg. =

Tortula minor (1420). 

Pottia splachnoides (Hornsch.) Broth. = Chenia 

leptophylla (1420). 

Pottia stenocarpa P.Varde CAR(47), NIG(334). 

Pottia subplanomarginata Dix. = Microbryum sub- 

planomarginatum (1420). 

Prionodon balboi Tosco & Piovano KEN(46). 

Prionodon ciliatus Besch. COM(40), KEN(46), 

MAD(40), MAU(105), REU(105), RWA(65), 

TAN(46), UGA(46). 

Prionodon densus (Hedw.) C.Müll. CBS(1531), 

NAT(1531), TRV(1531). 

Prionodon rehmannii Mitt. MAU(105). 

Prionodon rehmannii Mitt. This taxon is given as 

a synonym of P. ciliatus by Kis (46), and of P. 

densus by Perold & van Rooy (1531). I have left 

the taxon as it is until its correct position is 

determined. 

Pseudephemerum laxifolium (Ren. & Card.) Thér. 

ZAI(65). 

Pseudephemerum nitidum (Hedw.) Reim. 

KEN(46), MAD(40), RWA(65), TAN(46), 

ZAI(65). 

Pseudocrossidium crinitum (Schultz) Zander 

CBS(1531), ETH(50), LES(1531), MOZ(46), 

NAM(1531), NAT(1531), OFS(1531), 

TRV(1531), ZIM(46,61). 

Pseudocrossidium hornschuchianum (Schultz) 

Zander CBS(1531), ETH(50). 

Pseudocrossidium porphyreoneurum (C.Müll.) 

Zander BOT(1531), CBS(1531), DJI(1569), 

ERI(50), KEN(411), MAL(67,411), MOZ(357), 

NAM(1531), TAN(46), TRV(1531), UGA(46), 

ZAM(62), ZIM(46,61). 

Pseudocrossidium replicatum (Tayl.) Zander 

BOT(1531), CBS(1531), LES(1531), NAM(1531), 

OFS(1531), SOM(50), SUD(1419), TRV(1531). 

Pseudoleskea carbonarii Broth. ex Mich. ERI(50). 

Pseudoleskea chilensis (Lor.) Ochyra CBS(1531). 

Pseudoleskea claviramea (C.Müll.) Jaeg. = Pseudo- 

leskeopsis claviramea (717). 

Pseudoleskea dispersa C.Müll. ex Broth = Hyloco- 

miopsis cylindricarpa (46). 

Pseudoleskea falcifolia Dix. & Thér. UGA(46), 

ZAI(65). 

Pseudoleskea leikipiae (C.Müll.) Par. KEN(46). 

Pseudoleskea leskeoides (Par.) C.Müll. 

CBS(1531), LES(1531), NAT(1531), TRV(1531), 

ZIM(46). 

Pseudoleskea leskeoides var. macowanianum 

Thér. TAN(46). 

Pseudoleskea obtusiuscula Ren. & Card. 

MAD(40). 

Pseudoleskea penzigii Brizi ERI(50). 

Pseudoleskea plagiostoma C.Müll. ex Vent. 

ERI(50). 

Pseudoleskea plagiostoma var. attenuata C.Müll. 

ex Vent. ERI(50). 

Pseudoleskea pseudoattenuata (C.Müll.) Broth. 

CBS(1531), ERI(50), ETH(50), KEN(46), 

LES(1531), NAT(1531), TAN(46), TRV(1531), 

UGA(46). 

Pseudoleskea semensis C.Müll. ex Vent. ETH(50). 

Pseudoleskeopsis claviramea (C.Müll.) Thér. 

CBS(1531), DJI(1569), LES(1531), MOZ(46), 

NAT(1531), OFS(1531), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZIM(46). 

Pseudoleskeopsis leikipiae (C.Müll.) Thér. 

ERI(50). 

Pseudoleskeopsis pseudoattenuata (C.Müll.) Thér. = 

Pseudoleskea pseudoattenuata (46). 

Pseudoleskeopsis unilateralis Dix. CBS(1531). 

Pseudopohlia microstoma (Harv.) U.Miz. 

MAD(40), REU(105). 

Pseudoscleropodium purum (Hedw.) Fleisch. 

REU(105), STH(1296). 

Pseudosymblepharis Broth. It is likely that there is 

only one pantropical species present in this genus 

(1158). 

Pseudosymblepharis angustata (Mitt.) Hilp. 

SOC(40,1626). 

Pseudosymblepharis bescherellei Thér. = Oxystegus 

cylindricus (Brid.) Hilp (46) (= Trichostomum 

tenuirostre). 

Pseudosymblepharis circinnatula (Broth. in 

Voeltzk.) Zander COM(40), MAU(105), 

REU(105). 

Pseudosymblepharis socotrana (Mitt.) Thér. = Weissia 

artocosana (1420). 

Pseudosymblepharis syrrhopodontoides (Dix.) 

Zander KEN(46). 

Pseudotimmiella pocsii Biz. = Diphyscium pocsii 

(1420). 

Psilopilum brevilamellatum Thér. = Pogonatum 

congolense (289). 

Pterobryon flagelliferum Mitt. TAN(46). 

Pterobryon julaceum Broth. TAN(46). 

Pterobryon stipitatum (Mitt.) Touw ex De Sloover. 

Orthographic error in (65) for Porothamnium 

stipitatum. 

157

158 

Pterobryopsis longissima Dix. MAD(40). 

Pterobryopsis rehmannii Magill NAT(1531). 

Pterogonium gracile (Hedw.) Sm. CBS(1531), 

DJI(50), ETH(50), KEN(46), MAD(40), 

NAT(1531), REU(105), RWA(65), TAN(46), 

TRV(1531), UGA(46), ZAI(65). 

Pterogonium gracile var. kilimandscharicum 

(C.Müll.) Fleisch. TAN(46). 

Pterogonium gracile var. madagassum (Geh.) 

Fleisch COM(40), MAD(40), REU(105). 

Pterygoneurum macleanum Warnst. CBS(1531). 

Ptychomitriopsis africana Dix. NAT(1531), 

TRV(1531). 

Ptychomitrium crassinervium (C.Müll.) Schimp. 

ex Par. CBS(1531), ZAM(62), ZIM(46). 

Ptychomitrium crispatum (Hedw.) Jaeg. 

CBS(1531), LES(1531), MAD(40), NAT(1531), 

OFS(1531), SWA(1531), TAN(1406), 

TRV(1531), ZIM(46,61). 

Ptychomitrium cucullatifolium (C.Müll.) Jaeg. 

CBS(1531), LES(1531), NAT(1531), OFS(1531), 

TRV(1531), ZIM(46). 

Ptychomitrium depressum (C.Müll.) Par. 

CBS(1531), NAT(1531), OFS(1531), 

SWA(1531), TRV(1531). 

Ptychomitrium diexaratum Magill ex Magill & 

Schelpe LES(1531), NAT(1531). 

Ptychomitrium drouhardii Thér. MAD(40). 

Ptychomitrium eurybasis Dix. CBS(1531), 

MAL(67), SWA(1531), TRV(1531), ZIM(46,61). 

Ptychomitrium exaratifolium Robinson 

NAT(1531), SWA(1531), TRV(1531). 

Ptychomitrium helenicum (Mitt.) Par. STH(1296). 

Ptychomitrium marginatum (Wager & Dix.) Dix. = 

P. subcrispatum (1519). 

Ptychomitrium subcrispatum Thér. & P.Varde 

CBS(1531), LES(1531), MAS(40), MOZ(357), 

NAM(1531), NAT(1531), OFS(1531), 

SWA(1531), TRV(1531), ZIM(46,61). 

Ptychomnion aciculare (Brid.) Mitt. MAU(105). 

Pylaisiella frahmii Buck RWA(65). 

Pylaisiobryum abyssinicum (C.Müll.) Wijk & 

Marg. CAM(47), ETH(50), GAB(47), KEN(46). 

Pyrrhobryum mauritianum (Besch.) Manuel 

MAU(105), REU(105). 

Pyrrhobryum spiniforme (Hedw.) Mitt. 

CBS(1531), COM(40), FER(47), GAB(47), 

KEN(46), MAD(40), MAL(67), MAU(105), 

MOZ(46), NAT(1531), REU(105), RWA(65), 

SAO(47), SEY(40), TAN(46), TRV(1531), 

ZAI(65), ZIM(46). 

Pyrrhobryum spiniforme var. acunhense (Besch.) 

Manuel MAU(105). 

Pyrrhobryum spiniforme var. brevifolium (Besch.) 

Manuel REU(105). 

Pyrrhobryum spiniforme var. comorense (C.Müll.) 

Manuel COM(40). 

Pyrrhobryum vallis-gratiae (Hampe) Manuel 

CBS(1531). 

Quathlamba debilicostata Magill LES(1531), 

NAT(1531). 

Racomitrium alare (Broth.) Par. = Racomitrium 

subsecundum (1124). 

Racomitrium crispipilum (Tayl.) Jaeg. RWA(65). 

Racomitrium crispulum (Hook.f. & Wils.) Hook.f. 

& Wils. CBS(1531), NAT(1531), ZIM(61). 

Racomitrium crispulum var. tasmanicum (Hampe) 

Lawton = R. membranaceum (1915). 

Racomitrium durum (Broth.) Par. = Racomitrium 

subsecundum (1124). 

Racomitrium hypnoides L. = R. lanuginosum (1521). 

Racomitrium lamprocarpum (C.Müll.) Jaeg. 

CBS(1531), KEN(46), RWA(65), TAN(46), 

UGA(46), ZAI(65). 

Racomitrium lanuginosum (Hedw.) Brid. 

CBS(1531,1882), MAU(105), REU(105), 

STH(1296), ZAI(65). 

Racomitrium lanuginosum var. squalidum (Brid.) 

Brid. = R. lanuginosum (1914). 

Racomitrium lepervanchei Besch. REU(287). 

Racomitrium nigroviride (Broth.) Par. = Racomitrium 

lamprocarpum (1124). 

Racomitrium membranaceum (Mitt.) Par. 

MAD(40), REU(105), SEY(40). 

Racomitrium seychellarum Besch. = R. 

membranaceum (1915). 

Racomitrium subsecundum (Hook. & Grev.) Mitt. 

& Wi. CAM(47), KEN(46), REU(105), RWA(65), 

TAN(46), UGA(46), ZAI(65). 

Racopilum africanum Mitt. ANG(46), CAM(47), 

CAR(47), COM(40), CON(47), DJI(50), ERI(50), 

ETH(50), GAB(47), GUI(47), IVO(1880), 

KEN(46), MAD(40), MAU(105), REU(105), 

ROD(105), SIL(47), TAN(46), TOG(58), 

UGA(46), ZAI(65). 

Racopilum africanum Mitt. According to van 

Zanten (pers. comm.), all described African species 

of Racopilum may belong to this species. 

Racopilum angusti-stipulaceum C.Müll. 

COM(40).

Racopilum ayresii Mitt. REU(105). 

Racopilum brevipes C.Müll ex Broth. CAM(47), 

GUI(47), IVO(63), TOG(58). 

Racopilum buettneri Broth. CAR(47), GAB(47), 

GUI(47,1458), IVO(1458), SIL(47), TOG(58), 

ZAI(65). 

Racopilum buettneri fo. brevirostre Thér. & P.Varde. 



Racopilum capense C.Müll. ex Broth. BUR(65), 

CAM(47), CBS(1531), CAR(47), GAB(47), 

GHA(47), IVO(44), KEN(46), MAL(67), 

NAT(1531), REU(105), RWA(65), SIL(47), 

SWA(1531), TAN(46), TOG(58,1878), 

TRV(1531), UGA(46), ZAI(65), ZAM(46,62), 

ZIM(46,61). 

Racopilum capense var. subintegrifolium Thér. 

& P.Varde CAR(47). 

Racopilum cardotii Ren. MAD(40). 

Racopilum chevalieri Thér. CAR(47), GUI(47), 

TOG(1878). 

Racopilum crassicuspidatum Thér. & Corb. 

CAR(47), GAB(47). 

Racopilum ellipticum Ren. MAD(40). 

Racopilum gracile Mitt. ASC(1296). 

Racopilum leptotapes C.Müll. ex Card. CAR(47), 

IVO(47), ZAI(65). 

Racopilum longe-aristatum C.Müll. ex Vent = Raco- 

pilum africanum (1521). 

Racopilum macrocarpum Broth. BUR(65), 

TAN(46), UGA(46), ZAI(65). 

Racopilum madagassum Ren. MAD(40), ZAI(65). 

Racopilum madagassum var. grandistipulaceum 

Thér. & Ren. MAD(40). 

Racopilum marginatum Dix. CAR(47), GAB(47), 

GUI(47), LIB(47), MAD(40), UGA(46), ZAI(65). 

Racopilum mauritianum C.Müll. ex Besch. 

ETH(50), MAD(40), MAU(105), REU(105). 

Racopilum microdictyon Besch. COM(40), 

MAD(40). 

Racopilum naumannii C.Müll. ASC(1296). 

Racopilum orthocarpioides Broth. CAM(47), 

CAR(47), CON(47), GAB(47), GHA(47), 

GUI(47), IVO(44,1880), LIB(47), NIG(334), 

SAO(47), SIL(47), TOG(1878), ZAI(65). 

Racopilum penzigii C.Müll. ERI(50). 

Racopilum perrieri Thér. MAD(40). 

Racopilum plicatum Ren. & Card. COM(40), 

KEN(46), MAD(40), TAN(46). 

Racopilum schmidii C.Müll. REU(105). 

Racopilum speluncae C.Müll. ERI(50), ETH(50), 

KEN(46), TAN(46), UGA(46), ZAI(65). 

Racopilum speluncae var. grandistipulaceum Dix. 

ex Tosc. & Piov., nom. inv. KEN(46). 

Racopilum thomeanum Broth. GUI(47), SAO(47). 

Racopilum tomentosum (Hedw.) Brid. ANG(46), 

DIE(40), FER(47), GUI(47), MAD(40), 

MAU(105), REU(105). 

Racopilum ugandae Dix. UGA(46), ZAI(65). 

Radulina borbonica (Bel.) Buck. ANN(47), 

CAM(47), CAR(47), COM(40), CON(47), 

GAB(47), IVO(1880), LIB(47), MAD(40), 

MAU(105), NIG(47), REU(105), RWA(65), 

SAO(47), SEY(40), TAN(46), UGA(46), ZAI(65). 

Rauiella subfilamentosa (Besch.) Wijk & Marg. 

KEN(46), MAD(32,40), MAL(67), NAT(32), 

TAN(46), TRV(1531), ZIM(46). 

Regmatodon Brid. An unpublished world-wide 

revision by Eakin (1908) states that only R. 

polycarpus (Griff.) Mitt. occurs in Africa, and all 

other African names are made synonyms. 

Regmatodon bevicuspis P.Varde & Leroy. = R. secun- 

dus (46). 

Regmatodon imbricata Broth. Excluded. Clerical 

error in Born et al. (65): taxon does not exist 

(Frahm, pers. comm.) 

Regmatodon newtonii C.Müll. ex Broth. CAM(47), 

FER(47), GUI(47). 

Regmatodon secundus Kiaer KEN(46), MAD(40), 

MAL(67), TAN(46), ZAI(65). 

Renauldia dusenii (Broth.) Broth. CAM(47,2), 

FER(47,2). 

Renauldia hildebrandtielloides C.Müll. 

MAD(40,2). 

Renauldia lycopodioides Biz. ex Pócs TAN(46). 

Renauldia patentissima (Hampe) Broth. 

MAD(40,2). 

Renauldia patentissima var. mutica Thér. 

MAD(40). 

Renauldia patentissima var. tenuis Thér., nom. 

nud. MAD(40). 

Rhabdoweisia africana Dix. & Nav. TAN(46), 

UGA(46), ZAI(65). 

Rhabdoweisia crispata (Dicks.) Lindb. LES(1448), 

NAT(1531). 

Rhabdoweisia fugax BS&G NAT(1531), 

TRV(1531), ZIM(61). 

Rhabdoweisia lineata Rich. & Arg. CAM(47), 

RWA(65), TAN(46), ZAI(65). 

Rhachitheciopsis tisserantii P.Varde = Rhachithecium 

159

160 

tisserantii (80). 

Rhachithecium perpusillum (Thwait. & Mitt.) 

Broth. BUR(65), CAR(47), GUI(47), MAD(40), 

MAL(67), NIG(334,333), TAN(46,333), 

TRV(1531), ZAI(65,333), ZAM(62,333), 

ZIM(46,61). 

Rhachithecium tisserantii (P.Varde) Robins. 

CAR(47). 

Rhachithecium welwitschii (Duby) Zander 

ANG(46). 

Rhacocarpus purpurascens (Brid.) Par. CBS(1531), 

KEN(46), LES(1531), MAD(40), MAL(67), 

REU(105), RWA(46,65), TAN(46), TRV(1531), 

UGA(46), ZAI(65), ZIM(61). 

Rhacocarpus purpurascens var. rufipilus (Ren. 

& Card.) Wijk & Marg. MAD(40), REU(105). 

Rhacocarpus rehmannianus (C.Müll.) Wijk & Marg. 

= R. purpurascens (1531). 

Rhacopilopsis flexilis (Ren. & Card.) Buck 

MAD(717). 

Rhacopilopsis guineensis (Broth. & Par.) Buck 

CAR(47), GAB(47), GUI(47), LIB(47), SIL(47), 

ZAI(65). 

Rhacopilopsis kilimandscharica (Broth. & 

P.Varde) Buck RWA(65), TAN(46), ZAI(65). 

Rhacopilopsis transvaaliensis (Thér. & Dix. ex 

Sim) Buck BUR(65), MAD(40), MAL(67), 

MOZ(46,357), NAT(1531), RWA(65), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZAM(46), ZIM(61). 

Rhacopilopsis trinitensis (C.Müll.) Britt. ex Dix. 

ANG(46), CAM(47), CAR(47), CON(47), 

GAB(47), GHA(47), GUI(47), IVO(47,1458,1880), 

NIG(47), SAO(47), TAN(46), TOG(1878), 

UGA(46), ZAI(65), ZAM(46). 

Rhacopilopsis trinitensis fo. densa Dix. ex Tosco & 



Rhacopilopsis trinitensis var. acuminata (Card.) 

P.Varde CAR(47), CON(47), GAB(47), GUI(47), 

ZAI(65). 

Rhacopilopsis trinitensis var. gracilis (Card.) 

P.Varde CAR(47), GAB(47). 

Rhacopilopsis trinitensis var. pendula P.Varde 

GAB(47). 

Rhacopilopsis trinitensis var. perserrata J.Tayl. 

& P.Varde UGA(46). 

Rhaphidorrhynchium amoenum (Hedw.) Fleisch. 

MAU(105), REU(105). 

Rhaphidorrhynchium angustissimum (Geh.) 


Rhaphidorrhynchium argyrophyllum (Besch.) 

Broth. CAR(47), CON(47), GAB(47). 

Rhaphidorrhynchium brevihorridum (Broth.) 

Broth. CAM(47), CAR(47), CON(47), GAB(47), 

IVO(1880), NIG(334). 

Rhaphidorrhynchium cambouei (Ren. & Card.) 


Rhaphidorrhynchium crispans (Besch.) Broth. 

REU(105). 

Rhaphidorrhynchium gabonense Broth. & 


Rhaphidorrhynchium lamprosericeum (Ren. & 

Card.) Broth. IVO(1880), ZAI(65). 

Rhaphidorrhynchium nigricaule (Brid.) Broth. 

CAM(47), GAB(47), GHA(47), IVO(1880), 

SIL(47). 

Rhaphidorrhynchium pervilleanum (Geh.) Broth. 

MAD(40). 

Rhaphidorrhynchium plumularium (Mitt.) Broth. 

STH(1296). 

Rhaphidorrhynchium pseudo-brachythecium 

(Broth.) Broth. = Sematophyllum subsimplex 

(717). 

Rhaphidorrhynchium rubricaule (Besch.) Broth. 

MAD(40), REU(105). 

Rhaphidostegium crassiusculum Brid. = Semato- 

phyllum crassiusculum (1521). 

Rhaphidostegium dixonii Thér. = Sematophyllum 

dixonii (See “New Combinations”). 

Rhaphidostegium nanopyxis Geh. = Sematophyllum 

nanopyxis (See “New Combinations”). 

Rhaphidostegium nanothecium Jaeg., nom. nud. See 

note under Sematophyllum sp. 

Rhaphidostegium plumularium Mitt. = Rhaphidorr- 

hynchium plumularium (1521). 

Rhaphidostegium rigescens Card. = Sematophyllum 

rigescens (See “New Combinations”). 

Rhaphidostegium rutenbergii C.Müll., nom. nud. 

Excluded. This taxon was recorded from Mada- 

gascar (40). Rhaphidostegium is a synonym of 

Sematophyllum, and thus the taxa should be 

transferred there, but this name is not in Index 

Muscorum (1521), and I could not find a specimen 

in BM. 

Rhaphidostichum gracile Dix. CAM(47). 

Rhaphidostichum mahense (Besch.) Broth. 

SEY(40). 

Rhaphidostichum subluxurians Dix. & Thér. 

GHA(47), LIB(47).

Rhizofabronia perpilosa (Broth.) Broth. CAM(47), 

KEN(46), TAN(46), UGA(46), ZAI(65). 

Rhizofabronia persoonii var. persoonii 

(Schwaegr.) Fleisch. MAU(105), REU(105), 

RWA(65), SAO(47), TAN(46). 

Rhizofabronia persoonii var. sphaerocarpa 

(Dus.) Bizot ex Ochyra CAM(47), KEN(46), 

MAD(40), MAL(67), TAN(46), ZAI(65), 

ZIM(61). 

Rhizofabronia sphaerocarpa (Dus.) Fleisch. = R. 

persoonii var. sphaerocarpa (46). 

Rhizogonium spiniforme var. acunhense Besch. = 

Pyrrhobryum spiniforme var. acunhense (80). 

Rhizomnium punctatum (Hedw.) Kop. Excluded. 

Although this appears on the list for Ethiopia 

(50), Koponen (1036) does not consider it as 

present in Africa south of the Sahara. 

Rhodobryum alto-roseum (Broth.) Par. = Bryum 

huillense (354). 

Rhodobryum aubertii (Schwaegr.) Thér. = Bryum 

aubertii (354). 

Rhodobryum commersonii (Schwaegr.) Brid. 

CBS(1531), COM(40), KEN(46), MAD(40), 

MAL(46,67), MAU(105), NAT(1531), 

REU(105), SWA(1531), TAN(46), TRV(1531), 

UGA(46), ZIM(46). 

Rhodobryum ellipticifolium (Brizi) Broth.= Bryum 

ellipticifolium (1521). 

Rhodobryum ellipticifolium fo. compacto-gregaria 

Tong. Excluded. The records for this form have 

been associated with Bryum ellipticifolium (1521). 

Rhodobryum ellipticifolium fo. laxo-verticillata Tong. 

Excluded. The records for this form have been 

associated with Bryum ellipticifolium (1521). 

Rhodobryum giganteum (Schwaegr.) Par. 

MAD(40), REU(105). 

Rhodobryum huillense (Welw. & Duby) Touw = 

Bryum huillense (74). 

Rhodobryum jungneri (Broth.) Par. = Bryum keniae 

(354). 

Rhodobryum keniae (C.Müll.) Broth. = Bryum keniae 

(74). 

Rhodobryum leptoroseum Par., nom. nud. = Bryum 

keniae (354). 

Rhodobryum ontariense (Kindb.) Kindb. 

CBS(1531), KEN(46), LES(1531), MAD(40), 

MAL(67), NAT(1531), TAN(46), TRV(1531), 

UGA(46). 

Rhodobryum perspinidens (Broth.) Pócs KEN(46), 

TAN(46), UGA(46). 

Rhodobryum pertomentosum = Bryum keniae (354). 

Rhodobryum preussii Broth. & Par. = Bryum preussii 

(74). 

Rhodobryum pseudo-homalobolax Par. & Broth. = 

Bryum huillense (354). 

Rhodobryum quintasii (Broth.) Par. = Bryum huillense 

(354). 

Rhodobryum roseum (Hedw.) Limpr. auct. afr. See 

note under R. spathulatum. 

Rhodobryum saprophilum Par. = Bryum saprophilum 

Broth., hom. illeg (1521) = Bryum capillare 

(354). 

Rhodobryum spathulatum (Hornsch.) Pócs. 

According to Touw (1502), Pócs incorrectly used 

this name as the earliest name for R. ontariense, 

but in fact the type of R. spathulatum is Bryum 

aubertii. I have thus assumed that all plants called 

either R. roseum or R. spathulatum should be 

called R. ontariense. 

Rhodobryum staudtii (Broth.) Par. = Bryum staudtii 

(74). 

Rhodobryum umbraculum (Hook.) Schimp. ex Par. 

= Bryum umbraculum (23). 

Rhynchostegiella bequaertii Dix. & Thér. 

UGA(46), ZAI(65). 

Rhynchostegiella brunelii Tixier TOG(1878). 

Rhynchostegiella holstii (Broth.) Broth. 

CBS(1531), KEN(46), MAL(67), TAN(46), 

ZAI(65), ZIM(61). 

Rhynchostegiella keniae Dix. KEN(46). 

Rhynchostegiella linderi Thér. ZAI(65). 

Rhynchostegiella microtheca (Ren. & Card.) 

Broth. COM(40), MAD(40). 

Rhynchostegiella pendula Thér. ZAI(65). 

Rhynchostegiella sublaevipes Broth. & Bryhn 

NAT(1531). 

Rhynchostegiella tanneri Biz. ex Pócs TAN(46). 

Rhynchostegiella tenella var. tenella (Dicks.) 

Limpr. KEN(46), ZIM(46). 

Rhynchostegiella tenelliformis (Ren. & Card.) 

Broth. REU(105). 

Rhynchostegiella zeyheri (C.Müll.) Broth. 

CBS(1531), NAT(1531), TRV(1531), ZAM(46), 

ZIM(46). 

Rhynchostegium angustifolium Ren. & Card. 

MAD(40). 

Rhynchostegium bello-intricatum (Broth.) Par. = 

Schimperella bello-intricatum (717). 

Rhynchostegium bequaerti Thér. & Nav. ZAI(65). 

Rhynchostegium brachypterum (Hornsch.) Jaeg. 

161

162 

CAM(47), CBS(1531), MAL(67), NAT(1531), 

RWA(65), TRV(1531), ZAI(65), ZIM(46). 

Rhynchostegium buluense (Broth.) Par. GAB(47), 

TAN(46). 

Rhynchostegium cataractarum Thér. & P.Varde 

CAR(47). 

Rhynchostegium comorae (C.Müll.) Jaeg. 

COM(40), MAD(40), TAN(46), ZAI(65). 

Rhynchostegium distans Besch. COM(40), 

MAD(40), MAU(105), REU(105), TAN(46), 

UGA(46). 

Rhynchostegium gracilipes Thér. BUR(65), 

UGA(46). 

Rhynchostegium hopfferi (Welw. & Duby) Gepp 

SAO(47). 

Rhynchostegium horridum Broth. KEN(46), 

TAN(46), UGA(46), ZAI(65). 

Rhynchostegium luteonitens (Welw. & Duby) 

Jaeg. ANG(46). 

Rhynchostegium macowanianum Par. = Platyhypni- 

dium macowanianum (1521). 

Rhynchostegium megapelma (Broth.) Par. = Rhyn- 

chostegium brachypterum (717). 

Rhynchostegium megapolitanum var. meridionale 

Schimp. CAP(1296). 

Rhynchostegium nervosum (Ren.) Card. MAD(40). 

Rhynchostegium omocrates Buck RWA(65). 

Rhynchostegium oxyodon (Welw. & Duby) Gepp 

ANG(46). 

Rhynchostegium pectinatum (Mitt.) Par. 

MAS(40). 

Rhynchostegium pseudo-distans Card. MAD(40), 

REU(105). 

Rhynchostegium rhaphidorrhynchum (C.Müll.) 

Jaeg. CBS(1531), NAT(1531), TRV(1531), 

ZIM(61). 

Rhynchostegium riparioides (Hedw.) Card. = Platy- 

hypnidium riparioides (Hedw.) Fleisch.. 

Rhynchostegium ruwenzorense Broth. UGA(46). 

Rhynchostegium schoanum C.Müll. ex Par., nom. 

nud. ETH(50). 

Rhynchostegium subbrachypterum Broth. & 

Bryhn CBS(1531), NAT(1531), TRV(1531). 

Rhynchostegium tenuivagum (C.Müll. ex Broth.) 

Par. = Eurhynchium tenuivagum (717). 

Rhynchostegium tenuivagum var. congolense 

Card. ZAI(65). 

Rhynchostegium tenuivagum var. congolense Card. 

This variety was not dealt with when the species 

was transferred to Eurhynchium (717). It should 

be transferred if the variety proves to be distinct. 

Rhynchostegium volkensii (Broth.) Par. KEN(46), 

TAN(46), UGA(46), ZAI(65). 

Rhynchostegium volkensii fo. latifolium Tosco & 



Rhynchostegium volkensii fo. majus Tosco & Piovano. 



Rhytidium rugosum (Hedw.) Kindb. KEN(46), 

UGA(46). 

Rigodium kilimandscharicum (Broth.) Par. = R. 

toxarion (1202). 

Rigodium niveum Thér. = R. toxarion (1202). 

Rigodium toxarion (Schwaegr.) Jaeg. 

MAD(40,1202), MAL(67), MOZ(1202), 

TAN(46), TRV(1531), ZIM(61). 

Rutenbergia borbonica Besch. MAD(40), 

REU(105). 

Rutenbergia cirrata Ren. & Card. MAD(40). 

Rutenbergia limbata (Hampe) Besch. MAD(40). 

Rutenbergia madagassa Geh. & Hampe MAD(40). 

Rutenbergia madagassa var. flagelliformis Ren. 

& Card. MAD(40). 

Rutenbergia prionodon (Besch.) Ren. MAU(105), 

REU(105). 

Saelania glaucescens (Hedw.) Broth. LES(1531), 

NAT(1531), OFS(1531). 

Sanionia uncinata (Hedw.) Loeske GAB(47), 

KEN(46), OFS(1531), RWA(65), TAN(46), 

UGA(46), ZAI(65). 

Sarmentypnum sarmentosum (Wahlenb.) Tuom. 

& T.Kop KEN(46), TAN(1403). 

Sauloma africana Dix. & Thér. ZAI(65). 

Sauloma tisserantii P.Varde CAR(47). 

Schimperella atrotheca (P.Varde) P.Varde = Schim- 

perella bello-intricata (717). 

Schimperella bello-intricata (C.Müll. ex Broth.) 

Buck BUR(65), CAM(47), KEN(46), RWA(65), 

TAN(46), ZAI(65). 

Schimperella katalensis P.Varde = Schimperella 

bello-intricata (717). 

Schimperella rhynchostegioides Thér. MAD(40). 

Schistidium agassizii auct. afr., non Sull. & Lesq. 

Records for this taxon should be referred to S. 

rivulare (1124). 

Schistidium angustissimum (P.Varde) Ochyra 

UGA(46). 

Schistidium apocarpum var. apocarpum (Hedw.) 

BS&G CBS(1531), KEN(46), LES(1531),

NAT(1531), OFS(1531), REU(105), RWA(65), 

SAF(1882), TAN(46), TRV(1531), UGA(46). 

Schistidium apocarpum var. confertum (Funck) 

Moell. ETH(50). 

Schistidium cribrodontium (Herz.) Ochyra 

UGA(46). 

Schistidium perichaetiale (P.Varde) Ochyra 

KEN(46), TAN(1407), UGA(46). 

Schistidium perichaetiale var. piliferum (Demar.) 

Ochyra KEN(46). 

Schistidium rivulare (Brid.) Podp. RWA(65), 

TAN(46). 

Schizomitrium africanum (Mitt.) Ochyra See Cal- 

licostella africana. 

Schizomitrium ascensione (C.Müll.) Ochyra See 

Callicostella ascensionis. 

Schizomitrium attenuatum (C.Müll.) Ochyra See 

Callicostella attenuata. 

Schizomitrium brevipes (Broth.) Ochyra See Calli- 

costella bevipes. 

Schizomitrium chevalieri (Broth. in Corb.) Ochyra 

See Callicostella chevalieri. 

Schizomitrium chionophyllum (C.Müll.) Ochyra See 

Callicostella chionophylla 

Schizomitrium constrictum (C.Müll.) Ochyra See 

Callicostella constricta 

Schizomitrium emarginatulum (Broth.) Ochyra See 

Callicostella emarginatula . 

Schizomitrium eroso-truncatum (Card.) Ochyra See 

Callicostella eroso-truncata . 

Schizomitrium fissidentellum (Besch.) Ochyra See 

Callicostella fissidentella . 

Schizomitrium gabonense (Broth. & P.Varde) Ochyra 

See Callicostella gabonensis . 

Schizomitrium lacerans (C.Müll.) Ochyra See Cal- 

licostella lacerans . 

Schizomitrium laeviusculum (Mitt.) Ochyra See 

Callicostella laeviuscula ). 

Schizomitrium leptocladulum (C.Müll. ex Broth.) 

Ochyra See Callicostella leptocladula 

Schizomitrium maclaudii (Par. & Broth.) Ochyra 

See Callicostella maclaudii . 

Schizomitrium papillatum (Mont.) Sull. See Callico- 

stella papillata . 

Schizomitrium papillatum var. brevifolium (Fleisch.) 

Miller et. al. See Callicostella papillata var. 

bevifolia. 

Schizomitrium papillosulum (Broth. & P.Varde) 

Ochyra See Callicostella papillosula . 

Schizomitrium parvocellulatum (Dem. & P.Varde) 

Ochyra See Callicostella parvocellulata . 

Schizomitrium perpapillatum (Broth. & P.Varde) 

Ochyra See Callicostella perpapillata . 

Schizomitrium perrotii (Ren. & Card. ex Par.) Ochyra 

See Callicostella perrotii 

Schizomitrium pusillum (Broth.) Ochyra See Calli- 

costella pusilla . 

Schizomitrium salaziae (Besch.) Ochyra See Calli- 

costella salaziae. 

Schizomitrium seychellense (Besch.) Ochyra See 

Callicostella seychellensis. 

Schizomitrium subemarginatulum (Broth. & P.Varde) 

Ochyra See Callicostella subemarginatula. 

Schizomitrium triste (C.Müll.) Ochyra See Callico- 

stella tristis. 

Schizomitrium usambaricum (Broth.) Ochyra See 

Callicostella usambarica. 

Schizymenium basilaris (Schimp. ex C.Müll.) Shaw 

CAM(47), ETH(50). 

Schizymenium commixtum (Thér.) Shaw 

MAD(40). 

Schizymenium cratericola (Broth. in Mildbr.) 

Ochyra & Sharp KEN(46), TAN(46), UGA(46), 

ZAI(65). 

Schizymenium hariotii (Thér.) Shaw UGA(46). 

Schizymenium hymenostomum (Bruch & Schimp. 

ex C.Müll.) Shaw ETH(50). 

Schizymenium madagassum (Ren. & Card. ex 

Card.) Shaw MAD(40). 

Schizymenium mildbraedii (Broth.) Shaw 

KEN(46), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Schizymenium ruwenzorense (Thér. & Nav.) 

Ochyra & Sharp RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Schlotheimia abbreviata Broth., nom. nud. in 

sched. TAN(46). 

Schlotheimia acutifolia Ren. & Par. MAD(40). 

Schlotheimia acutifolia var. angustifolia Thér. 

MAD(40). 

Schlotheimia angulosa P.Beauv. MAU(105), 

REU(105), TAN(46). 

Schlotheimia badiella Besch. MAD(40), 

MAU(105), REU(105). 

Schlotheimia badiella var. brevifolia Ren. 

MAU(105). 

Schlotheimia badiella var. helicophylla Besch. 

MAD(40), REU(105). 

Schlotheimia badiella var. longicaulis Besch. 

REU(105). 

163

164 

Schlotheimia balfourii Mitt. SOC(40,1626). 

Schlotheimia bequaerti Thér. & Nav. ZAI(65). 

Schlotheimia boiviniana Besch. COM(40), 

MAD(40). 

Schlotheimia brachyphylla Ren. & Card. 

REU(105). 

Schlotheimia brachypodia Thér. & Nav. ANG(46), 

MAL(67), RWA(65), ZAI(65). 

Schlotheimia congolensis Card. ZAI(65). 

Schlotheimia conica Ren. & Card. MAD(40). 

Schlotheimia cyrtophylla P.Varde = S. grevilleana 

(1878). 

Schlotheimia excorrugata C.Müll. ex Card. 

MAD(40), TAN(46). 

Schlotheimia ferruginea (Hook. & Grev.) Brid. 

CBS(1531), NAT(1531), SWA(1531), TAN(46), 

ZAM(46,62), ZIM(61). 

Schlotheimia fornicata Duby MAD(40), 

MAU(105), REU(105). 

Schlotheimia fornicata var. apiculata P.Varde 

MAD(40). 

Schlotheimia fornicata var. microtheca P.Varde 

MAD(40). 

Schlotheimia foveolata Ren. & Card. MAD(40). 

Schlotheimia foveolata var. torquescens Card. 

MAD(40). 

Schlotheimia fragilifolia Thér. & Nav. TAN(46), 

ZAI(65). 

Schlotheimia gauthieri Thér. COM(40), MAD(40). 

Schlotheimia grevilleana Mitt. CAR(47), KEN(46), 

TAN(46), TOG(1878), UGA(46), ZIM(61). 

Schlotheimia holstii Broth., nom. nud. in sched. 

TAN(46). 

Schlotheimia illecebra Schimp. ex Besch. 

REU(105), ZIM(61). 

Schlotheimia laetevirens Broth. MAL(67), 

TAN(46). 

Schlotheimia malacophylla Besch. MAD(40), 

REU(105), TAN(46). 

Schlotheimia microcarpa Geh. MAD(40), 

REU(105). 

Schlotheimia microphylla var. farlowii Ren. & 

Card. MAU(105). 

Schlotheimia nossi-beana C.Müll. MAD(40). 

Schlotheimia nossi-beana var. linealis (Geh.) 


Schlotheimia pauli Ren. & Card. MAD(40). 

Schlotheimia percuspidata C.Müll. CBS(1531), 

MAL(67), NAT(1531), TAN(46), TRV(1531), 

ZIM(46). 

Schlotheimia perrieri Thér. MAD(40). 

Schlotheimia perrotii Ren. & Card. MAD(40). 

Schlotheimia perrotii var. breviseta Thér. 

MAD(40). 

Schlotheimia pobeguinii Par. & Broth. GUI(47). 

Schlotheimia poecilodictyon P.Varde & Thér. 

TAN(46). 

Schlotheimia quadrifolia Brid. = S. angulosa (1521). 

Schlotheimia richardii Besch. MAU(105), 

REU(105). 

Schlotheimia rigescens Broth. KEN(46), TAN(46), 

UGA(46), ZAI(65). 

Schlotheimia robillardii Duby MAD(40), 

MAU(105), REU(105), TAN(46). 

Schlotheimia robusta Thér. MAD(40), TAN(46). 

Schlotheimia rufoaeruginosa C.Müll. ANG(46), 

ZIM(46). 

Schlotheimia rufopallens C.Müll. CBS(1531), 

NAT(1531), TRV(1531). 

Schlotheimia rugifolia (Hook.) Schwaegr. 

ANG(46). 

Schlotheimia schweinfurthii C.Müll. = S. grevilleana 

(1878). 

Schlotheimia semidiaphana (Ren. & Card.) Card. 

MAD(40). 

Schlotheimia squarrosa Brid. MAD(40), 

REU(105). 

Schlotheimia subfornicata Thér. MAD(40), 

REU(105). 

Schlotheimia tenuiseta Geh. MAD(40). 

Schlotheimia torquata (Hedw.) Brid. MAS(40). 

Schlotheimia trichophora Ren. & Card. MAD(40). 

Schlotheimia trichophora var. stenophylla (Ren. 

& Card.) Thér. MAD(40). 

Schlotheimia trypanoclada Geh. MAD(40). 

Schoenobryum latifolium (Broth. & P.Varde) 

Manuel CAR(47), TAN(46). 

Schoenobryum latifolium var. microsporum 

(P.Varde) Manuel CAR(47). 

Schoenobryum madagassum (Geh.) Manuel 

MAD(40). 

Schoenobryum plicatulum (Dix. & Thér.) Manuel 

ZAI(65). 

Schoenobryum robustum (Broth.) Manuel 

DJI(1569), KEN(46), MAL(67), RWA(65), 

TAN(46), ZAI(65). 

Schoenobryum sp. CAR(47). 

Acrocryphaea emarginata Thér. & Nav. in P.Varde, 

nom. nud. was not dealt with by Manuel (1291) 

in his transfer of Acrocryphaea to Schoenobryum,

presumably because it was nom. nud. The list 

entry in Potier de la Varde (1567) (quoted by 

Index Muscorum (1521) as the basionym for this 

taxon) does not claim it as a new species, but there 

is no indication of where if anywhere it was 

described, or where the specimen is, although a 

locality is given. 

Schoenobryum subintegrum var. idanreense (Rich. 

& Edw.) Manuel NIG(47). 

Schoenobryum subrobustum (Broth. & P.Varde) 

Manuel CAR(47), NIG(334). 

Schoenobryum tisserantii (Thér. & P.Varde) 

Manuel CAR(47). 

Schoenobryum welwitschii (Duby) Manuel 

ANG(46), MAD(40), TAN(46). 

Schwetschkea brevis (Schimp.) Fleisch. ETH(50). 

Schwetschkea brotheri Dus. CAM(47), CAR(47), 

GAB(47), GUI(47), TAN(46), ZAI(65). 

Schwetschkea capillifolia Dix. ex Herz. ERI(50). 

Schwetschkea fabronioides (Welw. & Duby) 

Broth. ANG(46), CAM(47), CAR(47), GAB(47), 

GHA(47), LIB(47), TAN(146), TOG(1878), 

UGA(46), ZAI(65). 

Schwetschkea grateloupii (Mont.) C.Müll. 

COM(40), REU(105). 

Schwetschkea peracuminata Demar. & Leroy 

ZAI(65). 

Schwetschkea schweinfurthii C.Müll. CAR(47), 

IVO(1880), TAN(46), UGA(46). 

Schwetschkea usambarica Broth. TAN(46), 

ZAI(65), ZAM(46). 

Scleropodium purum L. = Pseudoscleropodium 

purum (1521). 

Scopelophila cataractae (Mitt.) Broth. CAR(47), 

RWA(65). 

Scopelophila ligulata (Spruce) Spruce RWA(65). 

Scorpiurium circinatum (Brid.) Fleisch. & Loeske 

DJI(1569). 

Sematophyllum alticaule Dix. & Thér. KEN(46), 

UGA(46), ZAI(65). 

Sematophyllum amblystegiocarpum (C.Müll.) 

Broth. SAO(47). 

Sematophyllum aneuron (Duby) Jaeg. MAU(105). 

Sematophyllum angusticymbeum (C.Müll.) Broth. 

COM(40). 

Sematophyllum annobonense Broth. in Mildbr., 


Sematophyllum brachycarpum (Hampe) Broth. 

CBS(1531), KEN(46), MOZ(46), NAT(1531), 

OFS(1531), SWA(1531), TRV(1531), UGA(46), 

ZAM(62), ZIM(46). 

Sematophyllum brachytheciiforme (Broth.) Broth. 

CAM(47), RWA(65), ZAI(65). 

Sematophyllum caespitosum (Hedw.) Mitt.= S. sub- 

pinnatum (46). 

Sematophyllum caespitosum var. sphaerotheca 

(C.Müll.) P.Varde, nom. inv. UGA(46). 

Sematophyllum caespitosum var. sphaerotheca 

(C.Müll.) P.Varde, nom. inv. The species for this 

variety has now been transferred to S. 

subpinnatum (46). This variety should be 

synonymised, or validated and transferred. 

Sematophyllum cellulosum Buck. RWA(65). 

Sematophyllum congolensis (Card.) Broth. 

ZAI(65). 

Sematophyllum constrictum Mitt. MAS(40). 

Sematophyllum coppeyi (Card.) Broth. MAD(40). 

Sematophyllum corticolum (Aongstr.) O’Shea 

MAS(40). 

Sematophyllum crassirete (Card.) Broth. CAR(47), 

CON(47), GAB(47), ZAI(65). 

Sematophyllum crassiusculum (Brid.) Broth. 

BUR(65), MAU(105), REU(105). 

Sematophyllum danckelmannii (C.Müll.) Broth. 

CAM(47). 

Sematophyllum dixonii (Thér.) O’Shea MAD(40). 

Sematophyllum dregei (C.Müll.) Bartr. CBS(1531), 

MAL(67), NAT(1531), SWA(1531), TAN(46), 

TRV(1531), ZIM(46). 

Sematophyllum dubium (Ren.) Broth. MAD(40). 

Sematophyllum elgonense (Dix.) Broth. KEN(46), 

RWA(65), TAN(46), UGA(46), ZAI(65), 

ZAM(259). 

Sematophyllum erythrocaulon Aongstr. 

STH(1296). 

Sematophyllum euryphyllum Dix. & Thér. ZAI(65). 

Sematophyllum flavovesiculosum Buck RWA(65). 

Sematophyllum friesiorum P.Varde KEN(46), 

UGA(46). 

Sematophyllum fulvifolium Mitt. CAR(47), 

GAB(47), GUI(1458), IVO(1458,1880), 

ROD(105), TAN(46). 

Sematophyllum fulvifolium var. patulifolium 


Sematophyllum gueinzii (Hampe) Magill 

CBS(1531), NAT(1531), OFS(1531), 

SWA(1531), TRV(1531), ZIM(61). 

Sematophyllum helenicum Mitt. STH(1296). 

Sematophyllum humillimum Dix. & Thér. TAN(46), 

ZAI(65). 

165

166 

Sematophyllum incurvum Mitt. ROD(105). 

Sematophyllum laetevirens (Broth. & Par.) Broth. 

CAR(47), GUI(47), TAN(46), UGA(46), ZAI(65). 

Sematophyllum laetevirens var. homomallum 


Sematophyllum longinerve (Duby) Jaeg. 

MAU(105). 

Sematophyllum nanopyxis (Geh.) O’Shea 

MAD(40). 

Sematophyllum nebulosum Buck ZAI(65). 

Sematophyllum nematocaulon (C.Müll.) Fleisch. 

COM(40). 

Sematophyllum nigro-alare Dix. ZAI(65). 

Sematophyllum obtusifolium (Ren. & Card.) 

Broth. CAM(47), CAR(47), GAB(47), GUI(47), 

MAD(40), TAN(46), ZAM(46). 

Sematophyllum obtusifolium var. laxifolium 


Sematophyllum parisii Broth. CAR(47), GUI(47). 

Sematophyllum perrevolutum (Broth.) Broth. 

TAN(46), ZAI(65). 

Sematophyllum physaophyllos (Welw. & Duby) 

Broth. ANG(46). 

Sematophyllum plagiothecioides Dix. & Thér. 

TAN(259), UGA(46), ZAI(65). 

Sematophyllum pobeguinii (Broth. & Par.) Broth. 

GUI(47). 

Sematophyllum rigescens (Card.) O’Shea 

MAD(40). 

Sematophyllum rivuletorum (Broth.) Broth. 

CAM(47). 

Sematophyllum ruwenzorense Dix. & Thér. 

ZAI(65). 

Sematophyllum schimperi (Besch.) Broth. 

MAU(105), REU(105), TAN(46). 

Sematophyllum serandii (Thér.) Broth. GUI(47). 

Sematophyllum sinuosulum (Besch.) Broth. 

KEN(46), MAD(40), REU(105), TAN(46). 

Sematophyllum sinuosulum var. latifolium (Card.) 


Sematophyllum sp. MAU(105). 

Rhaphidostegium nanothecium Jaeg., nom. nud. A 

specimen in BM of the type of this taxon appears 

to be a Sematophyllum. The taxon, being nom. 

nud., is thus left here awaiting revision of Sema- 

tophyllum. 

Sematophyllum sphaeropyxis (C.Müll.) Broth. 

CBS(1531), NAT(1531), SWA(1531), 

TRV(1531), ZIM(61). 

Sematophyllum stylites Buck ZAI(65). 

Sematophyllum subbrachytheciiforme P.Varde 

KEN(46), TAN(46), ZAI(65), ZAM(46,62). 

Sematophyllum subcochleatum Dix. & Thér. 

ZAI(65). 

Sematophyllum subjulaceum (Broth. & Par.) Broth. 

GUI(47). 

Sematophyllum subpinnatum (Brid.) Britt. 

ANG(259), BUR(65), CAM(47), CBS(1531), 

CAR(47), CON(47), DIE(40), GAB(47), 

GUI(47,1458), IVO(47,1880), LIB(47), MAD(40), 

MAL(67), MAU(105), MOZ(357), NAT(1531), 

NIG(47), OFS(1531), REU(105), RWA(65), 

SIL(47), SWA(1531), TAN(46), TOG(1878), 

TRV(1531), UGA(46), ZAI(65), ZIM(46). 

Sematophyllum subrugifolium Dix. MAD(40). 

Sematophyllum subscabrellum Ren. & Card. 

MAD(40). 

Sematophyllum subsimplex (Hedw.) Mitt. 

CAM(47), CAR(47), CON(47), GAB(47), 

GUI(47), IVO(1880), ZAI(65). 

Sematophyllum trachelocarpum (Kindb.) Broth. 

NIG(47). 

Sematophyllum wageri Wright & Wag. 

CBS(1531), MOZ(46), NAT(1531), SWA(1531), 

TRV(1531), ZIM(61). 

Sematophyllum zuluense (Sim) Magill CBS(1531), 

NAT(1531), TRV(1531). 

Semibarbula lambarenensis (P.Varde) Hilp. = Barbula 


Semibarbula orientalis (Web.) Wijk & Marg.= 

Barbula indica (1620). 

Serpotortella chenagonii (Ren. & Card.) Reese & 

Zander MAD(40), REU(105). 

Serpotortella cyrtophylla (Besch.) Reese & 

Zander MAD(1474), REU(105). 

Serpotortella madegassa Dix. = S. chenagonii (1474). 

Serpotortella marginata Dix. = S. chenagonii (1474). 

Sphaerothecium subchlorophyllosum (C.Müll. 

ex Rabenh.) Frahm CBS(1531), LES(1475,1531), 

NAT(1531), TRV(1531). 

Sphagnum africanum Welw. & Duby ANG(59), 

MAL(67), ZAI(59), ZAM(59,62), ZIM(59). 

Sphagnum australe Mitt. = S. lacteolum Besch. = S. 

strictum ssp. pappeanum (59). 

Sphagnum bernieri Besch. = S. cuspidatum (59) 

Sphagnum bessonii Warnst. = S. planifolium var. 

planifolium (59). 

Sphagnum bordasii Besch. = S. truncatum var. bordasii 

(59). 

Sphagnum bourbonense Crum REU(1632).

Sphagnum capense Hornsch. CBS(1531), 

MAD(59), MAL(67), MOZ(357), NAT(1531), 

REU(59), TRV(1531), ZIM(59). 

Sphagnum ceylonicum Mitt. ex Warnst. KEN(59), 

MAD(59), REU(59), TAN(259), UGA(59), 

ZAI(65). 

Sphagnum condensatum Brid. REU(105). 

Sphagnum condensatum Brid. Not treated by Eddy 

(59), but included in Crosby et al (40), and shown 

by Index Muscorum (1521) as present in Africa- 

3. 

Sphagnum cordemoyi Warnst. = S. violascens (59). 

Sphagnum cuspidatulum C.Müll. MAD(59), 

RWA(59). 

Sphagnum cuspidatum Hoffm. CON(59), GAB(59), 

MAD(59), RWA(59,65), ZAI(59,65). 

Sphagnum davidii Warnst. CAM(59), IVO(59), 

KEN(59), MAL(67), NIG(59), REU(59), 

RWA(59,65), SIL(59), TAN(59), UGA(59), 

ZAI(59,65). 

Sphagnum davidii var flavo-fuscescens Warnst. = S. 

davidii Warnst (59). 

Sphagnum drouhardii Card. = S. perichaetiale (59). 

Sphagnum ericetorum Brid. MAD(59), REU(59). 

Sphagnum fimbriatum Wils. TRV(1531). 

Sphagnum helenicum Warnst. STH(1296). 

Sphagnum humbertii Card. = S. capense (59). 

Sphagnum kerstenii Hampe = S. ceylonicum (59). 

Sphagnum laxirete Thér. = S. pappeanum C.Müll 

(1521) = S. strictum var. pappeanum (59). 

Sphagnum madegassum C.Müll. = S. planifolium 

var. planifolium (59). 

Sphagnum magellanicum C. Müll. auct. afr. = S. 

magellanicum ssp. grandirete (59). 

Sphagnum magellanicum ssp. grandirete 

(Warnst.) Eddy MAD(59). 

Sphagnum mathieui var. subsquarrosum Warnst. 

Excluded. Not mentioned by Eddy (59). 

Sphagnum mathieui Warnst. = S. truncatum var. 

bordasii (59). 

Sphagnum meridense (Hampe) C.Müll. Excluded. 

This is an American taxon, not known in Africa 

(59). The synonym of this name given in Crosby 

et al (40) is S. cymbifolium var. patens (Brid.) 

Brid., nom. illeg., but this is treated by Eddy (59) 

as nomina ambigua, as there is no specimen and 

the description is “inadequate for positive iden- 

tification”. 

Sphagnum nemoreum Scop. REU(105). 

Sphagnum nemoreum Scop. Not treated by Eddy 

(59), but listed in Crosby et al.(40), and shown in 

Index Muscorum (1521) for Africa-3 and Africa- 

4. 

Sphagnum obovatum Warnst. = S. rutenbergii (59). 

Sphagnum obtusifolium Warnst. Excluded. This 

taxon is listed for Madagascar and the Mascarenes 

(40), but is not treated by Eddy (59). However, 

this is unlikely to be the correct name, as the only 

entry in Index Muscorum (1521) is S. obtusifolium 

Ehrh., which is a synonym of S. palustre. The 

two synonyms of S. obtusifolium Warnst. given 

in Crosby & Schultze-Motel (40) are both now 

synonyms of S. violascens (59) which belongs to 

a different section of the genus. 

Sphagnum pallidum Warnst. = S. rutenbergii (59). 

Sphagnum palustre L. Excluded. Omitted from Eddy 

(59), but included in Crosby et al (40) for Réunion, 

although not shown in Index Muscorum (1521) 

for Africa. 

Sphagnum perichaetiale Hampe ANG(59), 

CBS(1531), MAD(59), MAU(59), NAT(1531), 

REU(59). 

Sphagnum perrieri Thér. MAD(40). 

Sphagnum perrieri Thér. Not dealt with by Eddy 

(59), although shown as valid for Africa-3 in 

Index Muscorum (1521), and included in Crosby 

et al (40). 

Sphagnum planifolium var. angustilimbatum 

(Warnst.) Eddy MAD(59), TAN(59), UGA(59), 

ZAI(65), ZIM(59). 

Sphagnum planifolium var. planifolium C.Müll. 

CAM(59), GAB(59), IVO(59), KEN(59), 

MAD(59), MAL(67), RWA(65), SAF(59), 

TAN(59), UGA(59), ZAI(59,65), ZAM(59), 

ZIM(59). 

Sphagnum planifolium var. rugegense (Warnst.) 

Eddy KEN(59), RWA(65), TAN(59), UGA(59), 

ZAI(65), ZAM(59). 

Sphagnum purpureum Schimp. ex Warnst. = S. 

violascens (59) 

Sphagnum pycnocladulum C.Müll. CBS(1531), 

MAL(67), RWA(59). 

Sphagnum rutenbergii C.Müll. MAD(59), 

MAU(59), REU(59). 

Sphagnum salvanii Warnst. = S. rutenbergii (59) 

Sphagnum slooveri Eddy BUR(59). 

Sphagnum strictum Sull. ZAI(59,65). 

Sphagnum strictum ssp. pappeanum (C.Müll.) 

Eddy CBS(1531), MAD(59), NAT(1531), 

REU(59), RWA(59), TAN(59), TRV(1531), 

167

168 

UGA(59), ZAI(59,65). 

Sphagnum truncatum Hornsch. CBS(1531), 

MOZ(59), NAT(1531), SWA(1531), TRV(1531). 

Sphagnum truncatum var. bordasii (Besch.) Eddy 

BUR(65), MAD(59), MAU(59), NAT(59), 

REU(59), TRV(59). 

Sphagnum tumidulum var. confusum Eddy 

REU(59). 

Sphagnum tumidulum var. microphyllum Warnst.= 

S. tumidulum var. tumidulum (59) 

Sphagnum tumidulum var. tumidulum Besch. 

MAD(59), MAU(59), REU(59). 

Sphagnum violascens C.Müll. CBS(1531), 

MAD(59), MAL(67), MAU(59), MOZ(59,357), 

REU(59), SAF(59). 

Splachnobryum acutifolium Thér. LIB(47). 

Splachnobryum aquaticum C.Müll. SOM(50). 

Splachnobryum boivinii C.Müll. MAD(40). 

Splachnobryum brotheri Par. GAB(47), GUI(47), 

MAL(67). 

Splachnobryum erosulum C.Müll. ex Dus. 

CAM(47), SIL(47). 

Splachnobryum gracile Besch. COM(40), 

MAD(40). 

Splachnobryum inundatum C.Müll. MAD(40). 

Splachnobryum obtusum (Brid.) C.Müll. 

CAM(47), CAR(47), GAB(47), GUI(47), 

MAD(40), SIL(47). 

Splachnobryum rostratum Broth. & Par. CAR(47), 

GAB(47), GUI(47), NIG(334). 

Splachnobryum suborbifolium Par. & Broth. 

CAR(47), GUI(47), ZAI(65). 

Splachnobryum tenerum C.Müll. ex Dus. CAR(47), 

LIB(47), ZAI(65). 

Splachnum adolphi-friederici Broth. ZAI(65). 

Squamidium biforme (Hampe) Broth. = S. brasiliense 

(93). 

Squamidium biforme var. congesta (Ren. & Card.) 

Een = S. brasiliense (93). 

Squamidium biforme var. densirameum (Broth.) 

Biz. = S. brasiliense (93). 

Squamidium brasiliense (Hornsch.) Broth. 

CBS(1531), COM(40), KEN(46), MAD(40), 

NAT(1531), REU(105), RWA(65), SWA(1531), 

TAN(1406,46), TRV(1531), UGA(46). 

Stableria aethiopica (C.Müll.) Broth. = Orthodontium 

gracile (1521). 

Stableria ruwenzorense Thér. & Nav. = Orthodontium 

ruwenzorensis (See “New Combinations”). 

Stableria tenella (Mitt.) Broth.= Orthodontium gracile 

(1521). 

Stenocarpidiopsis perfiliaris (C.Müll.) Fleisch. 

ETH(50). 

Stereophyllum acutum var. acutum Broth. & Par. in 

Par. = Entodontopsis tenuinervis (719). 

Stereophyllum acutum var. viridis Broth. & Par. in 

Par. = Entodontopsis tenuinervis (719). 

Stereophyllum andungense (Welw. & Duby) 

Gepp ANG(46), CAR(47), GHA(47), NIG(334), 

TOG(1878), UGA(46). 

Stereophyllum auriculatum Gepp = Entodontopsis 

leucostega (1349). 

Stereophyllum brunthaleri Broth. Excluded. Probably 

a synonym of S. radiculosum (719). 

Stereophyllum conterminum Card. = Entodontopsis 

nitens (65). 

Stereophyllum favoroi Tosco & Piovano KEN(46). 

Stereophyllum flavobrunneum C.Müll. in Par., 

nom. nud. SEN(47). 

Stereophyllum guineense Broth. & Par. = S. radicu- 

losum (719). 

Stereophyllum indicum (Bel.) Mitt. = S. radiculosum 

(1349). 

Stereophyllum laetevirens Broth. = Entodontopsis 

nitens (1349). 

Stereophyllum laxirete Broth. & P.Varde CAR(47). 

Stereophyllum leucomioides Broth. & Par. = Ento- 

dontopsis tenuinervis (719). 

Stereophyllum limnobioides Ren. Excluded. Probably 


Stereophyllum lingulaefolium (Welw. & Duby) Gepp 

= Entodontopsis nitens (1349) 

Stereophyllum loseanum Broth. & Par. = 

Entodontopsis tenuinervis (719). 

Stereophyllum macrocarpum J.Tayl. & P.Varde 

ETH(719). 

Stereophyllum microcarpum Thér. = Entodontopsis 

tenuinervis (719). 

Stereophyllum natalense Sim = S. radiculosum (719). 

Stereophyllum nitens Mitt. = Entodontopsis nitens 

(717). 

Stereophyllum odontocalyx (C.Müll.) Jaeg.& Sauerb. 

= S. radiculosum (719). 

Stereophyllum omalosekos (Welw. & Duby) Jaeg. & 

Sauerb. Excluded. Probably a synonym of S. 

radiculosum (719). 

Stereophyllum pallidifolium Ren. & Card. = Ento- 

dontopsis nitens (1349). 

Stereophyllum pobeguinii Broth. & Par. in Par. = 

Entodontopsis nitens (1349).

Stereophyllum radiculosum (Hook.) Mitt. 

ANG(46), CAM(47), CBS(1531), CAR(47), 

ETH(50), GUI(47), IVO(44,1458,63), KEN(46), 

MAD(40), MAL(67), MAU(105), NAT(1531), 

NIG(47,334), REU(105), SWA(1531), TAN(46), 

TOG(47), TRV(1531), ZAI(65), ZAM(46,62), 

ZIM(61,46). 

Stereophyllum reclinatum Par. & Broth. in Par. 

Excluded. Probably a synonym of S. radiculosum 

(719). 

Stereophyllum rhynchostegioides Broth. & Par. = 

Entodontopsis rhynchostegioides (719). 

Stereophyllum rigescens Broth. Excluded. Probably 


Stereophyllum rivale P.Varde = S. radiculosum (1349). 

Stereophyllum rivulare Mitt. = S. radiculosum (1349). 

Stereophyllum rufescens Broth. & Par. = 

Entodontopsis rufescens (719). 

Stereophyllum tenuinerve Broth. & Par. = Entodon- 

topsis tenuinervis (719). 

Stereophyllum tisserantii Broth. & P.Varde in P.Varde 

= S. radiculosum (1349). 

Stereophyllum woodii (Sim) Magill in Magill & Schelpe 

Excluded. Probably a synonym of S. radiculosum 

(719). 

Stereophyllum zuluense Broth. & Bryhn in Bryhn = 

Entodontopsis nitens (1349). 

Stoneobryum mirum (Lewinsky) Norris & Robinson 

CBS(347), NAT(1531,347). 

Streblotrichum convolutum (Hedw.)P.Varde = 

Barbula convoluta (46). 

Streptopogon calymperes C.Müll. ex Geh. 

MAD(40), RWA(65). 

Streptopogon calymperoides Demaret & P.Varde 

ZAI(65). 

Streptopogon erythrodontus var. erythrodontus 

(Tayl.) Wils. RWA(65), ZAI(65). 

Streptopogon erythrodontus var. rutenbergii 

(C.Müll. ex Geh.) Salm. CAM(47), KEN(46), 

MAD(40), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Streptopogon subelimbatus Card. MAD(40). 

Symblepharis rhacomitrioides Dix. CAM(47). 

Symphyodon pygmaeus (Broth.) He & Snider 

MAD(40), MOZ(357), REU(357). 

Syntrichia alpestris (Dix. in Herz.) Zander KEN(46). 

Syntrichia ammonsiana (Crum & Anderson) 

Ochyra CBS(1531), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), TRV(1531). 

Syntrichia amphidacea (C.Müll.) Zander 

TAN(46), ZIM(61). 

Syntrichia antarctica (Hampe in C.Müll.) Zander 

CBS(1863), LES(1863), TRV(1863). 

Syntrichia austroafricana (Kramer) Zander 

CBS(1863), LES(1863). 

Syntrichia cavallii (Negri) Ochyra ETH(1158), 

KEN(46,136), RWA(65), TAN(46), UGA(46), 

ZAI(65). 

Syntrichia chisosa (Magill, Delg. & Stark) Zander 

CBS(1531), LES(1531), NAM(1531), TRV(1531). 

Syntrichia fragilis (Tayl.) Ochyra CBS(1531), 

CAR(47), DJI(1569), ERI(50), ETH(50), KEN(46), 

LES(1531), MOZ(46), NAM(1531), NAT(1531), 

OFS(1531), RWA(65), SOM(50), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZIM(46,61). 

Syntrichia laevipila var. laevipila Brid. KEN(411). 

Syntrichia leucostega var. leucostega (C.Müll.) 

Zander CBS(1863), LES(1863), OFS(1863), 

TRV(1863). 

Syntrichia leucostega var. trachyneura (Dix.) 

Zander CBS(1863), OFS(1863), TRV(1863). 

Syntrichia mollis (B&S ex C.Müll.) Zander 

ETH(50). 

Syntrichia norvegica var. norvegica Web. 

LES(1863), NAT(1863). 

Syntrichia pagorum (Milde) Amann CBS(1531), 

NAT(1531), SWA(1531), TRV(1531), ZIM(61). 

Syntrichia pagorum (Milde) Amann. The differences 

between this taxon and S. laeviplia may be too 

slight to justify retention of this taxon (Sollman, 

pers. comm.). 

Syntrichia papillosa (Wils. in Spruce) Jur. 

CBS(1531), KEN(411), NAT(1531), OFS(1531). 

Syntrichia princeps (De Not.) Mitt. CBS(1531), 

NAT(1531). 

Syntrichia princeps (De. Not.) Mitt. Material called 

by this name from the southern hemisphere may 

belong to S. antarctica (Sollman, pers. comm.), 

and it is possible that S. princeps does not occur 

in sub-Saharan Africa. 

Syntrichia rubella (Hook.f. & Wils.) Zander 

CBS(1863). 

Syntrichia ruralis ssp. ruralis (Hedw.) Web. & 

Mohr. CBS(1531), ETH(1865), LES(1531,1863), 

NAT(1531), OFS(1531), SUD(1419), TRV(1531). 

Syntrichia subaristata (Broth. & Schimp. ex 

C.Müll.)Zand. ERI(50), ETH(50). 

Syrrhopodon Schwaegr. All records previously 

recorded for the genus Calymperopsis have now 

169

170 

been transferred to this genus (see note under 

Calymperopsis). As Reese’s revisions do not 

discuss the African distribution in detail, the data 

shown here should for subgenus Pseudocalymperes 

should be treated with caution, considering 

previous misunderstanding of the subgenus, until 

the distribution has been revised, . The 

distributions of S. africanus var. africanus and S. 

subdisciformis are most likely to be suspect. 

Syrrhopodon aculeato-serratus Besch. = S. gardneri 

(132). 

Syrrhopodon africanus ssp. africanus (Mitt.) 

Par. BEN(400), CAM(47,1708), CON(47), 

GAB(47), GUI(47,1458), MAD(40), NIG(47,761), 

SIL(47), TAN(46), ZAI(1125,65). 

Syrrhopodon africanus ssp. africanus (Mitt.) Par. 

See comments above (under Syrrhopodon) about 

distribution data for this taxon. 

Syrrhopodon africanus ssp. mandrakensis (Tixier) 

Reese MAD(1346). 

Syrrhopodon afrociliatus C.Müll. ex Dus. = S. armatus 

(24). 

Syrrhopodon apertifolius Besch. = S. prolifer var. 

acanthoneuros (1344). 

Syrrhopodon armatissimus Reese MAD(1375). 

Syrrhopodon armatus Mitt. CAM(47), CON(47), 

GAB(47), GUI(363), IVO(44), NIG(47), ZAI(65). 

Syrrhopodon armatus fo. rubrotomentosus = S. 

armatus (24). 

Syrrhopodon armatus ssp. insularus (Biz. & 

Onraedt) Orban & Reese MAD(361), 

REU(361), SEY(361), TAN(46,361), ZAI(1125). 

Syrrhopodon armatus var. latifolius P.Varde = S. 

armatus (Reese, pers. comm.). 

Syrrhopodon asper Mitt. GAB(24), KEN(46), 

MAD(105), MAL(67), MOZ(46), REU(105), 

RWA(65), TAN(46), TRV(1531), ZAI(65), 

ZAM(62), ZIM(61). 

Syrrhopodon brevifolius Broth. & P.Varde = S. 

gardneri (24). 

Syrrhopodon brevifolius var. overlaetii (Thér. & 

Nav.) Dix. & Thér. = S. gardneri (24). 

Syrrhopodon brevivaginans Demar. & Leroy = S. 

incompletus var. incompletus (132). 

Syrrhopodon chevalieri P.Varde = S. planifolius (24). 

Syrrhopodon chlorothrix (C.Müll.) Broth. 

ZAI(65). 

Syrrhopodon congolensis Besch. Excluded. See 

Orbán (24). 

Syrrhopodon crenulatus (Tixier) Reese GAB(47). 

Syrrhopodon croceus Mitt. SEY(1889). 

Syrrhopodon cryptocarpus Dozy & Molk. = Helico- 

nema cryptocarpus (Dozy & Molk.) Ellis & 

Eddy, nom. inval (= Chameleion cryptocarpus) 

(755). 

Syrrhopodon cuneifolius Thér. MAD(24,40). 

Syrrhopodon decipiens P.Varde = S. planifolius (24). 

Syrrhopodon disciformis Dusen CAM(47,1708), 

CAR(47), GAB(47), IVO(1880), NIG(47), 

ZAI(400). 

Syrrhopodon gabonensis Broth. & P.Varde = S. 

gardneri (24). 

Syrrhopodon gabonensis var. aspericuspis P.Varde 

= S. asper (24). 

Syrrhopodon gardneri (Hook.) Schwaegr. 

ANG(46), BUR(65), CAM(24), COM(40), 

GAB(24,47), GUI(24,363), MAD(24,40), 

MAL(67), REU(24,105), RWA(65), SAO(24), 

SIL(24), TAN(46,24), ZAI(65). 

Syrrhopodon gaudichaudii Mont. CBS(1531), 

COM(24), GAB(24), GUI(24,363), MAD(24,40), 

MAL(67), MOZ(46), NAT(1531), OFS(1531), 

REU(24), SWA(1531), TAN(46,362), 

TRV(1531), UGA(46,363), ZAI(65), ZIM(24). 

Syrrhopodon graminifolius Ren. & Card. 

MAD(24,40), MAU(1375). 

Syrrhopodon hispido-costatus Ren. & Card. in Ren. 

= S. prolifer var. hispidocostatus (1344). 

Syrrhopodon incompletus var. incompletus 

Schwaegr. TAN(46,362), ZAI(65,24). 

Syrrhopodon insularus Biz. & Onraedt = S. armatus 

var. insularus (132). 

Syrrhopodon involutus Schwaegr. CAR(47), 

CHG(40), MAD(24,40), MAU(24), REU(24), 

SEY(24,40), ZAI(24,65). 

Syrrhopodon isleanus Besch. = S. prolifer var. prolifer 

(1344). 

Syrrhopodon jacobi Besch. = Serpotortella chenagonii 

(1474) 

Syrrhopodon lamprocarpus Mitt. CAM(47), 

CON(24), FER(47), GAB(47), RWA(65), 

SAO(47), ZAI(65). 

Syrrhopodon leptodontoides Besch. = Serpotortella 

cyrtophylla (1474) 

Syrrhopodon lineaefolius P.Varde = S. cryptocarpa 

(24) (= Chameleion cryptocarpus). 

Syrrhopodon linealis = S. gaudichaudii (132). 

Syrrhopodon lisowskii Orban GUI(363), ZAI(65). 

Syrrhopodon mahensis Besch. REU(105), 

SEY(24,40).

Syrrhopodon mauritianus C.Müll. ex Aongstr. 

COM(24), MAD(24,40,362), MAU(24,362), 

REU(24,362), TAN(46,362). 

Syrrhopodon microblepharis P.Varde = S. gardneri 

(132). 

Syrrhopodon mildbraedii Broth. in Mildbr. = S. asper 

(24). 

Syrrhopodon mildbraedii var. parvifolius P.Varde = 

S. asper (24). 

Syrrhopodon obuduensis Egunyomi & Olarinmoye = 

S. africanus ssp. africanus (1909) 

Syrrhopodon oxyphyllus P.Varde = S. gaudichaudii 

(24). 

Syrrhopodon parasiticus (Brid.) Besch. 

CAM(1708), MAD(40). 

Syrrhopodon patentifolius P.Varde, nom. nud. 

MAD(40). 

Syrrhopodon patulifolius P.Varde, hom. illeg. 

MAD(40). 

Syrrhopodon paucifimbriatus C.Müll. ex Dus. = S. 

armatus (24). 

Syrrhopodon perdusenii Reese CAM(1711). 

Syrrhopodon phragmidaceus C.Müll. Excluded. See 

Orbán (24). 

Syrrhopodon planifolius fo. subaciliatus P.Varde. 



Syrrhopodon planifolius var. planifolius P.Varde 

BUF(24), GAB(24,47), GUI(24), IVO(1458), 

NIG(24). 

Syrrhopodon pobeguinii Par. & Broth. = S. 

gaudichaudi (24). 

Syrrhopodon prolifer var. acanthoneuros 

(C.Müll.) C.Müll. COM(24,40), MAD(24,40), 

MAU(24), REU(24), TAN(46). 

Syrrhopodon prolifer var. hispidocostatus (Ren. 

& Card. in Ren.)Orban & Reese MAD(24,40), 

MAU(1375), REU(24). 

Syrrhopodon prolifer var. prolifer Schwaegr. 

COM(105), MAD(24,40), MAU(105), 

REU(105), SEY(24,40), TAN(46). 

Syrrhopodon pulvinaris Par. = S. gaudichaudii (24). 

Syrrhopodon quintasii Broth. = S. gardneri (24). 

Syrrhopodon revolutus Dozy & Molk. = S. involutus 

(1579). 

Syrrhopodon rodriguezii Ren. & Card. REU(24). 

Syrrhopodon spiralis Ren. & Card. = S. prolifer var. 

prolifer (1344). 

Syrrhopodon stuhlmannii Broth. ex Orban 

TAN(46,361). 

Syrrhopodon subdisciformis Dusen BEN(400), 

CAM(47,1708), GAB(47), GHA(47), LIB(47), 

NIG(47), TAN(46), ZAI(400). 

Syrrhopodon subdisciformis Dusen See comments 

above (under Syrrhopodon) about distribution 

data for this taxon. 

Syrrhopodon tenellifolius (Dus.) Besch. = S. armatus 

(24). 

Syrrhopodon tixieri Reese MAD(1346). 

Syrrhopodon uncinifolius C.Müll. = S. gaudichaudii 

(24). 

Syrrhopodon usambaricus Broth. ex Orban 

TAN(46,362). 

Syrrhopodon virgulicola Reese CAM(1711). 

Taxiphyllum gabonense Broth. & P.Varde 

CAR(47), CON(47), GAB(47), IVO(1880), 

SIL(47), TAN(46), ZAI(65). 

Taxiphyllum gabonense var. ectropothecioides 

P.Varde CAR(47), GAB(47). 

Taxiphyllum gabonense var. plagiothecioides 

P.Varde CAR(47), GAB(47), TAN(46), ZAI(65). 

Taxiphyllum laxalare Buck ZAI(65). 

Taxiphyllum maniae (Ren. & Card.) Fleisch. 

COM(40), MAD(40). 

Taxithelium afro-acuminulatum (C.Müll.) Par. 

CAM(47). 

Taxithelium andongense Gepp ANG(46), 

CAR(47). 

Taxithelium argyrophyllum Ren. & Card. 

MAD(40). 

Taxithelium chloropterum (C.Müll.) Ren. & Card. 

FER(47). 

Taxithelium compressicaule Broth. CAM(47), 

CAR(47), GAB(47), IVO(1880), LIB(47). 

Taxithelium confusum Card. MAD(40). 

Taxithelium glabriusculum Broth. CAM(47), 

CAR(47), GAB(47), LIB(47), SIL(47), 

TOG(58,47). 

Taxithelium glaucophyllum Besch. MAD(40). 

Taxithelium glaucovirens Card. ZAI(65). 

Taxithelium guineense Broth. & Par. GUI(47), 

ZAI(1552). 

Taxithelium hirtellum Par. & Ren. MAD(40). 

Taxithelium homalophyllum (Mitt.) Broth. 

NIG(47). 

Taxithelium kuilui (C.Müll.) Ren. & Card. 

CON(47). 

Taxithelium latitruncatum Card. MAD(40). 

Taxithelium lepto-punctatum Broth. LIB(47). 

Taxithelium nigerianum Broth. & Par. MAI(47). 

171

172 

Taxithelium nossianum Besch. MAD(40). 

Taxithelium perglabrum Broth. & Par. CAR(47), 

GAB(47), GUI(47). 

Taxithelium perminutum Broth. CAM(47). 

Taxithelium planulum Besch. COM(40), 

MAD(40). 

Taxithelium planum (Brid.) Mitt. ASC(1296), 

NIG(47). 

Taxithelium pobeguinii Broth. & Par. CAR(47), 

GAB(47), GUI(47), SIL(47). 

Taxithelium pseudo-amoenum (Bel.) Ren. 

MAU(105), REU(105). 

Taxithelium ramivagum Broth. CAM(47), 

CAR(47), CON(47), GAB(47), GUI(47), 

IVO(47,1880), SIL(47). 

Taxithelium ramivagum var. elongatum P.Varde 

GAB(47). 

Taxithelium rhaphidostegioides (Broth.) Broth. 

TAN(46). 

Taxithelium rhaphidostegioides var. pallidum 

Broth. TAN(46). 

Taxithelium rhaphidostegioides var. viride Broth. 

TAN(46). 

Taxithelium rotundatulum Broth. CAM(47), 

CAR(47), GAB(47), GUI(47), IVO(47,1880), 

LIB(47), SIL(47). 

Taxithelium rotundatulum var. minus Broth. ex 

Corb. CAR(47). 

Taxithelium stenosecos (Welw. & Duby) Broth. 

ANG(46). 

Taxithelium suboctodiceras Broth. & Par. 

CAR(47), CON(47), GAB(47), GUI(47), MAI(47). 

Taxithelium subrotundatulum Broth. & Par. 

CAM(47), CAR(47), CON(47), GAB(47), 

GUI(47), IVO(1880), TAN(46), ZAI(65). 

Taxithelium voeltzkowii Broth. MAU(105). 

Tayloria indica Mitt. MAD(40), MAS(40). 

Tayloria isleana (Besch.) Broth. MAD(40), 

NAT(1531), REU(123), TAN(46), ZIM(1547). 

Tayloria jamesonii (Tayl.) C.Müll. CAM(47). 

Tayloria kilimandscharica Broth. KEN(136), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Tayloria limbata Dix., hom. illeg. Excluded. An- 

notation on type (from Kenya) in BM: “not 

Splachnaceae. det. Aune Koponen 1981.” No 

alternative identification is offered, although 

Dixon (16) considers Bryaceae as a possible 

alternative family. 

Tayloria orthodonta (P.Beauv.) Demar. CAM(47), 

GUI(47), KEN(46), MAD(40), MAL(67), 

REU(123), RWA(65), SIL(47), TAN(46), 

TRV(1531), ZAI(65), ZIM(46,61). 

Tayloria serrata var. tenuis (With.) BS&G 

KEN(46), UGA(46). 

Tayloria solitaria (Hedw.) T.Kop & W.Web. 

ANG(46), CAM(47,123), GUI(47), MAD(40), 

SAO(47), TAN(46), UGA(46), ZAI(65). 

Tayloria thomeana (Broth.) Broth. = T. solitaria 

(107). 

Tetraplodon bryoides Lindb. = T. mnioides (46). 

Tetraplodon mnioides (Hedw.) BS&G KEN(46), 

TAN(46), UGA(46), ZAI(65). 

Tetrapterum tetragonum (Hook.) Andrews 

CBS(1531). 

Thamnium malgachum Card. = Thamnobryum mal- 

gachum (See “New Combinations”). 

Thamnium stipitatum (Mitt.) Broth. ex Par. = Poro- 

thamnium stipitatum (15). 

Thamnobryum malgachum (Card.) O’Shea 

MAD(40). 

Thuidium abietinum ssp. abietinum BS&G 

LES(1635), NAT(1531). 

Thuidium abietinum ssp. abietinum BS&G. Touw 

(pers. comm.) believes that the occurrence of this 

taxon in South Africa is very unlikely. 

Thuidium aculeoserratum Ren. & Card. 

MAD(32,40), REU(32). 

Thuidium annobonense Broth. in Mildbr., nom. nud. 

= T. gratum (32). (The only collection is cited 

in Touw (32), although the name is omitted from 

the paper’s list of synonyms.) 

Thuidium borbonicum Besch. = T. versicolor (32). 

Thuidium byssoideum Besch. COM(32,40), 

MAD(32,40). 

Thuidium chenagonii C.Müll. ex Ren. & Card. 

BUR(32,65), CAM(32), CAR(32), COM(32,40), 

GAB(32), GUI(32), IVO(32), KEN(46), 

MAD(32,40), MAL(67), MOZ(46), RWA(65), 

SIL(32), TAN(46), UGA(46), ZAI(32,65), 

ZAM(411), ZIM(46). 

Thuidium chloropsis (C.Müll.) Par. = T. gratum var. 

gratum (32). 

Thuidium gratum (P.Beauv.) Jaeg. ANG(32), 

ANN(32), BUF(32), BUR(32), CAM(32), 

CAR(32), COM(32,40), CON(32), FER(32), 

GAB(32), GHA(32), GUB(32), GUI(32,1458), 

IVO(32,1458,1880), LIB(32), MAD(32,40), 

MAL(67), MAI(32), NIG(32,334), SEN(32), 

SEY(40), SIL(32), SUD(32), TAN(32), 

TOG(46,1878), UGA(32), ZAI(32,65).

Thuidium gratum ssp. gratum (Besch.) Touw = T. 

gratum (1636). 

Thuidium gratum ssp. subscissum (Besch.) Touw = T. 

gratum (1636). 

Thuidium intricatum Jaeg. BUR(32), CAM(32), 

RWA(32), ZAI(32). 

Thuidium involvens ssp. involvens (Hedw.) Mitt.. 

Excluded. The collection from Angola given this 

name by Gepp is Thuidium involvens ssp. tho- 

meanum (32). 

Thuidium involvens ssp. thomeanum (Broth.) 

Touw ANG(46), CAM(32), CAR(32), CON(32), 

GAB(32), GHA(32), IVO(32), NIG(32), SAO(32), 

TOG(58,1878), ZAI(32). 

Thuidium laevipes Mitt. = T. varians (32). 

Thuidium loricalycinum (C.Müll.) Kindb. 

TAN(46). 

Thuidium matarumense Besch. CBS(1531,32), 

COM(32,40), KEN(46), LES(32,1531), 

MAD(32,40), NAT(1531,32), OFS(32,1531), 

REU(32,287), TAN(46), TRV(1531,32), 

UGA(46), ZIM(61). 

Thuidium minusculum (Mitt.) Jaeg. MAL(1636), 

TAN(1636). 

Thuidium nigerianum (Mitt.) Par. = T. involvens var. 

thomeanum (32). 

Thuidium perscissum (C.Müll.) Jaeg. Exclude. All 

specimens given this name have proved to be 

other taxa. It is probably a synonym of T. 

chenagonii, but the diagnosis gives insufficient 

information to confirm this (32). 

Thuidium pseudo-involvens (C.Müll.) Jaeg. 

COM(32,40), MAD(32,40), MAU(32), 

TAN(46). 

Thuidium pycnangiellum C.Müll. ex Broth. = T. 

chenagonii (32). 

Thuidium ramusculosum (Mitt.) Jaeg. ANG(46), 

CAM(32), CBS(32), FER(32), GUI(32), KEN(46), 

MAD(32,40), MAL(67), NAT(1531,32), 

OFS(1531), RWA(65), TAN(46), TRV(1531,32), 

UGA(46), ZAM(46,62), ZIM(46). 

Thuidium spurio-involvens Broth. and Par. = T. 

varians (32). 

Thuidium strangulatum P.Varde = T. tenuissimum 

(32). 

Thuidium subfalcatum (Mitt.) Par. = T. involvens var. 

thomeanum (32). 

Thuidium subperscissum C.Müll., nom. nud. = T. 

pseudo-involvens (32). 

Thuidium tamariscellum (C.Müll.) Bosch. & Lac. 

TAN(46), UGA(46). 

Thuidium tamariscinum (Hedw.) BS&G 

REU(32,287), TAN(46). 

Thuidium tenuisetum Welw. & Duby = T. varians 

(32). 

Thuidium tenuissimum Welw. & Duby ANG(46), 

CAM(32), CAR(32), FER(32), GAB(32), GUI(32), 

IVO(63), NIG(32), ZAI(32). 

Thuidium trachynotum Ren. & Par. = T. ramusculosum 

(32). 

Thuidium varians Welw. et Duby ANG(46), 

BUR(32), CAM(32), CAR(32), COM(32,40), 

GAB(32), GHA(32), GUI(32), IVO(32,1880), 

MAD(32,40), MAL(67), MOZ(46), 

NAT(1531,32), NIG(334), RWA(65), SIL(32), 

SUD(32), SWA(32,1531), TAN(46), 

TOG(58,1878), UGA(46), ZAI(32), 

ZAM(62,411). 

Thuidium venustulum Besch. Excluded. The record 

of this taxon (TAN(46)) is doubtful (Touw, pers. 

comm.) 

Thuidium versicolor (C.Müll.) Broth. CAM(32), 

CBS(1531,32), KEN(46), LES(32), MAD(32,40), 

MAL(67), NAT(1531,32), REU(32,287), 

RWA(32,65), TAN(46), TRV(1531,32), 

UGA(46), ZAI(32,65), ZAM(46), ZIM(46). 

Timmiella anomala (BS&G) Limpr. KEN(411), 

TAN(46). 

Timmiella barbula (De Not.) Schwaegr.= T. barbuloi- 

des (1420). 

Timmiella barbuloides (Brid.) Moenk. CAP(1296), 

DJI(1569), ERI(50). 

Timmiella brevidens Dix. UGA(46). 

Timmiella cameruniae Broth. CAM(47), TAN(46). 

Timmiella pelindaba Magill CBS(1531), 

LES(1531), OFS(1531), TRV(1531), ZIM(61). 

Tisserantiella pulchella (Thér. & Hilp.) Zander 

CAM(47). 

Tisserantiella spathulata P.Varde = T. pulchella (1420). 

Tortella aprica (C.Müll.) Broth. ERI(50). 

Tortella caespitosa Schwaegr. = T. humilis (1420). 

Tortella eckendorffii (P.Varde) Zander CAR(47). 

Tortella fragilis (Hook.f. & Wils.) Limpr. 

CBS(1531), LES(1531), NAT(1531), RWA(65), 

ZAI(65). 

Tortella fragillima P.Varde TAN(46). 

Tortella humilis (Hedw.) Jenn. CBS(1531), 

DJI(1569), KEN(46), LES(1531), MAD(40), 

MAU(1637), MOZ(46), NAT(1531), SOC(40), 

STH(1296), SWA(1531), TAN(46), TRV(1531), 

173

174 

UGA(46), ZAI(65), ZIM(46). 

Tortella smithii Townsend SOC(40,1626). 

Tortella somaliae (C.Müll.) Zander SOM(50). 

Tortella syrrhopodontoides Dix. = Pseudosymble- 

pharis syrrhopodontoides (1420). 

Tortella theriotii Broth. & P.Varde = T. humilis 

(1420). 

Tortella theriotii var. angustata Dix. & P.Varde = T. 

humilis (1420). 

Tortella torquescens (Schimp.) Broth. = Tortula pilifera 

Hook (1521) = Pseudocrossidium crinitum 

(1420). 

Tortella tortuosa (Hedw.) Limpr. MAL(67). 

Tortella tortuosa (Hedw.) Limpr. Sollman (pers. 

comm.) believes that this taxon is unlikely in 

Africa, but has yet to examine the collection. 

Tortella vernicosa (Ren. & Card.) Broth. REU(105). 

Tortella xanthocarpa (C.Müll.) Broth. CBS(1531), 

LES(1531), NAM(1531), NAT(1531), 

OFS(1531), SWA(1531), TRV(1531), ZIM(61). 

Tortula abyssinica De Not. ETH(50). 

Tortula alpestris Dix. = Syntrichia alpestris (1420). 

Tortula ammonsiana Crum & Anderson = Syntrichia 

ammonsiana (1420). 

Tortula atrovirens (Sm.) Lindb. CBS(1531), 

ERI(50), KEN(411), LES(1531), NAM(1531), 

NAT(1531), OFS(1531), TRV(1531). 

Tortula bogosica (C.Müll.) Zander ERI(50), 

ETH(1865), KEN(46), SUD(1419,1707), 

TRV(1531). 

Tortula brevi-mucronata (C.Müll.) Broth. = Syntrichia 

fragilis (1420). 

Tortula cavallii Negri = Syntrichia cavallii (1420). 

Tortula chisosa Magill, Delgadillo & Stark = Syntrichia 

chisosa (1420). 

Tortula cochlearifolia P.Varde TAN(46). 

Tortula cuspidatissima (C.Müll.) Broth. 

ASC(1296). 

Tortula erubescens (C.Müll.) Broth. = T. fragilis (65) 

(= Syntrichia fragilis). 

Tortula eubryum (C.Müll.) Broth. = Barbula eubryum 

(1420). 

Tortula fragilis Tayl. DJI(1569). 

Tortula fragilis Tayl. = Syntrichia fragilis (1420). 

Tortula fusco-mucronata (C.Müll.) Broth. = Pseu- 

docrossidium replicatum (1419). 

Tortula hyalinoblasta Broth. in Mich., nom. nud. 

ERI(50). 

Tortula incerta (Mitt.) Broth. = Trichostomum 

incertum (1420). 

Tortula inermis (Brid.) Mont. = Syntrichia inermis 

(1420). 

Tortula kerensis Broth. in Mich., nom. nud. ERI(50). 

Tortula laevipila var. laevipila (Brid.) Schwaegr. = 

Syntrichia laevipila var. laevipila (1420). 

Tortula laevipila var. wachteri Barkm. KEN(411). 

Tortula laevipila var. wachteri Barkm. This variety 

was not dealt with when the species was transferred 

to Syntrichia (1420). If the variety proves to be 

distinct, it should be transferred, otherwise syno- 

nymised. 

Tortula laxiretis Broth. in Mich., nom. nud. ERI(50). 

Tortula leiostomoides P.Varde UGA(46). 

Tortula leucochlora C.Müll. ASC(1296). 

Tortula longirostris (Hampe) Broth. = Barbula lon- 

girostris (1420) (= Hennediella limbata). 

Tortula madagassa Dix. MAD(40). 

Tortula microtricha Broth. in Mich., nom. nud. 

ERI(50). 

Tortula minor (C.Müll.) Zander ERI(50). 

Tortula minor var. elatior (C.Müll.) Zander 

ERI(50). 

Tortula mollis (Broth. & Schimp.) Broth. = Syntrichia 

mollis (1420). 

Tortula muralis var. madagascariensis Thér. 

MAD(40). 

Tortula muralis var. muralis Hedw. 

CBS(1531,1882), NAM(1531), NAT(1531). 

Tortula muralis var. obcordata (Schimp.) Moenk. 

ZAI(65). 

Tortula nana (C.Müll.) Broth. = Barbula nana (1521) 

(= Trichostomum brachydontium). 

Tortula omissa Thér. MAD(40). 

Tortula pagorum (Milde) De Not. = Syntrichia 

pagorum (1420). 

Tortula papillosa Wils. in Spruce = Syntrichia papillosa 

(1420). 

Tortula pertorquata Broth. in Mich., nom. nud. 

ERI(50). 

Tortula pierrotii Biz. TAN(46). 

Tortula porphyreoneura (C.Müll.) Townsend. = Pseu- 

docrossidium porphyreoneurum (1420). 

Tortula princeps De Not. = Syntrichia princeps (1420). 

Tortula rehmannii (C.Müll.) Sim = Barbula rehmannii 

(82) (= Barbula indica.) 

Tortula rhizophylla (Sak.) Iwats. & K.Saito = Phascum 

leptophyllum C.Müll (68) (= Chenia leptophylla). 

Tortula rufa (Besch.) Broth., hom. illeg. MAU(105). 

Tortula ruralis ssp. ruralis (Hedw.) Gaertn. = Syntrichia 

ruralis var. ruralis (1420).

Tortula sabinae Townsend SUD(1419). 

Tortula sp. ETH(50). 

Desmatodon corticola Schimp. in Par., nom. nud. 

Desmatodon Brid. is now a synonym of Tortula 

Hedw (1420). This taxon is not dealt with by 

Zander (1420) or Index Muscorum (1521), and 

requires investigation. 

Tortula subaristata (Broth. & Schimp.) Broth. = 

Syntrichia subaristata (1420). 

Tortula subaristata fo. robusta Dix. in Herz. Excluded. 


with Syntrichia subaristata, to which name the 

owning species has transferred. 

Tortula subrufa Card. MAD(40). 

Tortula subspathulata (C.Müll.) Broth. = Syntrichia 

fragilis (1621). 

Tortula subtorquatifolia Dix. = T. atrovirens (1420). 

Tortula tanganyikae Dix. = Syntrichia amphidaceus 

(1420). 

Trachycarpidium tisserantii Dix. & P.Varde 

CAR(47), NIG(334), UGA(46). 

Trachyphyllum borgenii (Ren. & Card.) Broth. 

MAD(12,40). 

Trachyphyllum curvifolium (Mitt.) Gepp. = T. inflexum 

(12). 

Trachyphyllum dilatatum (Ren.) Ren. = T. inflexum 

(12). 

Trachyphyllum dusenii (Broth.) Broth. CAM(47), 

CAR(47), GHA(12), GUI(12,1458), IVO(1880), 

MAI(12), MOZ(357), NIR(12), NIG(12,334), 

SEN(12), SIL(12), TOG(1878), ZAI(12,65), 

ZAM(46,62), ZIM(61). 

Trachyphyllum fabronioides (Besch.) Gepp. = T. 

inflexum (12). 

Trachyphyllum fabronioides fo. comorense Besch. = 

T. inflexum (12). 

Trachyphyllum fabronioides var. imbricatum (Ren. 

& Par.) Card. = T. inflexum. This is not included 

as a synonym by Buck (12), but in his discussion 

of T. inflexum (p.382), it is clear that this variety 

would in all likelihood be included here. 

Trachyphyllum gastrodes (Duby) Gepp ANG(46), 

MAL(67), NAT(1531), SEN(47), SWA(1531), 

TRV(1531), ZAI(12), ZAM(46,62), ZIM(46). 

Trachyphyllum inflexum (Harvey) Gepp ALD(12), 

COM(12,40), MAD(12,40), MAU(12), 

MOZ(357), REU(12), ROD(105), SUD(12), 

TAN(46), ZAM(62). 

Trachyphyllum pinnatum (Broth. & Par.) Broth. = T. 

dusenii (12) 

Trachypodopsis laxoalaris Broth. MAL(67), 

UGA(46), ZAI(65). 

Trachypodopsis normandii (Broth. & Par.) Fleisch. 

GUI(47), SIL(47). 

Trachypodopsis normandii fo. viridis Par., nom. nud. 



Trachypodopsis serrulata var. crispatula (Hook.) 

Zant. CAM(47), NIG(47), TAN(46). 

Trachypodopsis serrulata var. serrulata (P.Beauv.) 

Fleisch. CAM(47), COM(40), FER(47), 

IVO(44,1880), KEN(46), MAD(40), MAL(67), 

NIG(47), REU(105), RWA(65), SAO(47), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZIM(46). 

Trachypus appendiculatus (Ren. & Card.) Broth. 

MAD(40), TAN(46). 

Trachypus bicolor Reinw. & Hornsch. NAT(1531), 

TAN(46), TRV(1531). 

Trachypus bicolor var. hispidus (C.Müll.) Card. 

GUI(47), TAN(46). 

Trachypus bicolor var. viridulus (Mitt.) Zant. 

MAD(40), REU(105), SAO(47), TAN(46). 

Trachythecium le-testui Thér. & P.Varde 

CAM(47), CAR(47), GAB(47), IVO(1880). 

Trematodon angolensis Welw. ex Duby ANG(46). 

Trematodon borbonicus Besch. REU(105). 

Trematodon congolensis Leroy ZAI(65). 

Trematodon curvicollis Crad. MAD(40). 

Trematodon decaryi Thér. MAD(40). 

Trematodon dentatus Broth. & P.Varde CAR(47). 

Trematodon divaricatus Bruch CBS(1531), 

MAL(67), MOZ(46), NAT(1531), SAO(47), 

TAN(46), TRV(1531), ZAI(65). 

Trematodon hildebrandtii C.Müll. COM(40). 

Trematodon intermedius Welw. & Duby. = T. para- 

doxus (46). 

Trematodon lacunosus Ren. & Card. MAD(40). 

Trematodon le-testui P.Varde GAB(47). 

Trematodon longicollis Michx. KEN(411), 

MAL(67,411), NAT(1531), OFS(1531), 

RWA(65), TAN(411), TRV(1531), UGA(46), 

ZAI(65), ZIM(46,61). 

Trematodon mayottensis Besch. COM(40), 

MAL(67), NAT(1531), OFS(1531), TAN(46), 

ZAM(62,411), ZIM(46). 

Trematodon minutulus C.Müll. ex Dus. CAM(47), 

ZAI(65). 

Trematodon nudus C.Müll. ex Dus. CAM(47). 

Trematodon pallidens C.Müll. COM(40), 

175

176 

MAD(40). 

Trematodon paradoxus Hornsch. ANG(46), 

CBS(1531), MAL(67), NAM(1531), NAT(1531), 

OFS(1531), REU(105), RWA(65), SWA(1531), 

TRV(1531), UGA(46), ZAI(65), ZAM(62,411), 

ZIM(46,61). 

Trematodon paradoxus var. nanus (Welw. & 

Duby) Sim ANG(46), CAR(47), TAN(46), 

UGA(46), ZAI(65), ZIM(46). 

Trematodon pillansii Dix. ex Sim CBS(1531). 

Trematodon platybasis C.Müll. MAD(40). 

Trematodon reticulatus C.Müll. ex Geh. MAD(40). 

Trematodon schroederi Broth. MAL(67), 

TAN(46), ZAI(65). 

Trematodon subambiguus Besch. MAU(105), 

REU(105). 

Trematodon tisserantii Broth. & P.Varde CAR(47). 

Trematodon usambaricus Broth. TAN(46). 

Trematodon victoriae C.Müll. ex Dus. CAM(47), 

ZAI(65). 

Trematodon viguieri Card. COM(40), MAD(40). 

Trichosteleum asselii Tixier IVO(1880). 

Trichosteleum borbonicum (Bel.) Jaeg. = Radulina 

borbonica (717) 

Trichosteleum borbonicum var. brachycarpum 

Ren. & Card. MAD(40). 

Trichosteleum borbonicum var. brachycarpum Ren. 

& Card. Although Buck (717) does not specifically 

mention this variety when making several new 

synonyms for Radulina borbonica, he states that 

“Although several names have been used for 

species in this group in tropical Africa, all specimens 

seen from throughout the area represent a single 

taxon.”. I assume this taxon will at some point 

become formally recognised as a synonym. 

Trichosteleum chrysophyllum P.Varde CAR(47), 

GAB(47), IVO(1880). 

Trichosteleum constrictum (Brid.) Ren. REU(105). 

Trichosteleum debettei (Besch.) Kindb. REU(105). 

Trichosteleum debettei var. laevisetum Card. 

MAD(40). 

Trichosteleum dicranelloides Broth. LIB(47), 

SAO(47). 

Trichosteleum dimorphum Dix. IVO(1880), SIL(47). 

Trichosteleum fusco-virescens Par. & Broth. 

GUI(47). 

Trichosteleum gabonense Broth. & P.Varde 

GAB(47), GUI(47), IVO(1880). 

Trichosteleum grossepapillosum Par. & Broth. 

CAR(47), GAB(47), GUI(47,1458), IVO(1880), 

LIB(47), NIG(1492). 

Trichosteleum heterophyllum Biz. & Onr. 

MAD(40). 

Trichosteleum hornschuchii (Hampe) Jaeg. 

IVO(1458). 

Trichosteleum humbertii P.Varde BUR(65), 

KEN(46), TAN(46), ZAI(65). 

Trichosteleum jaegeri P.Varde SIL(47). 

Trichosteleum jonesii Biz. ex Pócs TAN(46). 

Trichosteleum lachmanii Biz. IVO(47,1880). 

Trichosteleum le-testui P.Varde CON(47), 

GAB(47). 

Trichosteleum leviusculum Ren. & Card. 

MAD(40). 

Trichosteleum lorifolium P.Varde GAB(47). 

Trichosteleum madagassum Jaeg., nom. nud. 

MAD(40). 

Trichosteleum mammillipes Broth. = Radulina bor- 

bonica (717). 

Trichosteleum microcalyx Ren. & Card. GAB(47), 

ZAI(65). 

Trichosteleum microdontum (Besch.) Ren. 

MAD(40). 

Trichosteleum microdontum var. megapterum 

Ren. & Card. MAD(40). 

Trichosteleum papillosum (Hornsch.) Jaeg. 

NIG(47). 

Trichosteleum perchlorosum Broth. & Bryhn 

CBS(1531), TRV(1531), ZIM(46). 

Trichosteleum perhamosum Broth. = Radulina bor- 

bonica (717). 

Trichosteleum permixtum Card. MAD(40). 

Trichosteleum perrotii (Ren. & Card.) Ren. & 

Card. MAD(40), MAU(105). 

Trichosteleum perrotii var. eurydictyon Ren. & 


Trichosteleum pervilleanum (C.Müll. ex Geh.) 

Buck ZAI(65). 

Trichosteleum pobeguinii Par. & Broth. CAR(47), 

GUI(47), IVO(1880). 

Trichosteleum staudtii Broth. CAM(47), GAB(47), 

IVO(1880). 

Trichosteleum staudtii var. densirameum P.Varde 

GAB(47). 

Trichosteleum stictum (Besch.) Kindb. SEY(40). 

Trichosteleum sublaeve P.Varde CAR(47), 

IVO(1880). 

Trichosteleum subpycnocylindricum Broth. = 

Radulina borbonica (717). 

Trichosteleum subscabrisetulum (C.Müll.) Jaeg.

COM(40). 

Trichosteleum subulatulum (C.Müll.) Jaeg. 

COM(40), RWA(65), ZAI(65). 

Trichosteleum tisserantii P.Varde CAR(47), 

IVO(1880). 

Trichosteleum trachypyxis (Geh.) Ren. & Card. 

MAD(40). 

Trichosteleum usambaricum Broth. TAN(46). 

Trichostomum acutiusculum (Broth.) Zander 

TAN(46). 

Trichostomum ayresianum Besch. MAU(105). 

Trichostomum borbonicum Een REU(105). 

Trichostomum brachydontium Bruch ANG(46), 

BOT(1531), CBS(1531), CAP(1296), CAR(47), 

ERI(50), GUI(47), IVO(1458), KEN(46,411), 

LES(1531), MAD(40), MAL(67,411), MOZ(46), 

NAM(1531), NAT(1531), NIG(334), OFS(1531), 

REU(105), SWA(1531), TAN(46), TRV(1531), 

ZAM(62), ZIM(46,61). 

Trichostomum brachydontium ssp. mutabile 

(Bruch) Giac. REU(105). 

Trichostomum calymperaceum Broth. & Par. 

GUI(47). 

Trichostomum cardotii Biz. = T. borbonicum (1420). 

Trichostomum circinatulum Broth. in Voeltzk. = 

Pseudosymblepharis circinatula (1420). 

Trichostomum circinatum (Besch.) Broth. = 

Oxystegus cylindricus (Brid.) Hilp (1521) (= 

Trichostomum tenuirostre). 

Trichostomum crispulum Bruch CAP(1296), 

DJI(1569), STH(1296), TAN(46). 

Trichostomum decurvifolium Par. & Broth. = T. 

brachydontium (1158). 

Trichostomum distans Hampe in C.Müll. ETH(50). 

Trichostomum flavescens Dix. ZAI(65). 

Trichostomum fragilifolium Dix. KEN(46), 

TAN(46). 

Trichostomum incertum (Mitt.) Zander ROD(105). 

Trichostomum ligulaefolium (Broth. & Par.) 

Zander CAR(47), GUI(47), TOG(1878). 

Trichostomum lorifolium Broth. & Par. CAR(47), 

GUI(47), NIG(334), TAN(46), ZAI(65). 

Trichostomum mutabile Bruch = T. brachydontium 

var. mutabile (1420). 

Trichostomum muticum Par. ROD(105). 

Trichostomum pennequinii Ren. & Par. MAD(40). 

Trichostomum perplexum P.Varde CAR(47). 

Trichostomum perrieri Thér. MAD(40). 

Trichostomum platyphyllum (Broth. ex Ihs.) Chen 

MAD(40). 

Trichostomum pulicare (Besch.) Zander COM(40). 

Trichostomum rhodesiae Broth. ZIM(46,61). 

Trichostomum ruvenzorense (Broth.) Broth. 

KEN(46), TAN(46), UGA(46), ZAI(65). 

Trichostomum soulae (C.Müll. in Ren. & Card.) 

Zander MAD(40). 

Trichostomum sparsifolium (Ren. & Card.) Card. = T. 

brachydontium (1158) (as Barbula sparsifolia 

Ren. & Card.) 

Trichostomum sporaphyllum (Ren. & Card.) Card. 

MAD(40). 

Trichostomum subintegrum (Broth.) Broth. 

UGA(46). 

Trichostomum tenuirostre (Hedw. & Tayl.) Lindb. 

ANG(46), CAM(47), CBS(1531), COM(40), 

LES(1531), MAD(40), MAL(67), MAU(105), 

NAT(1531), OFS(1531), REU(105), 

RWA(65,46), TAN(46), UGA(46), ZAI(65), 

ZAM(62). 

Trichostomum trirete Dix. ZAI(65). 

Trichostomum unguiculatum (Mitt.) Zander 

CBS(1531), ZIM(61). 

Trichostomum usambaricum (Broth.) Broth. 

CAM(47), TAN(46). 

Trichostomum villaumei Thér. MAD(40). 

Triquetrella tristicha (C.Müll.) C.Müll. CBS(1531), 

ETH(50), LES(1531), OFS(1531), TRV(1531). 

Trismegistia rigidicaulis (Broth.) Broth. CAM(47). 

Tristichium mirabile (C.Müll.) Herz. KEN(146), 

LES(146,1531), NAT(1531), TAN(146). 

Ulea welwitschii (Duby) Broth. = Rhachithecium 

welwitschii (1420). 

Ulota ecklonii (Hornsch.) Jaeg. CBS(1531). 

Ulota fulva Brid. MAD(40), MAU(105), REU(105). 

Ulota tanganyikae P.Varde TAN(46). 

Vesicularia albo-viridis (Ren.) Broth. MAU(105). 

Vesicularia albo-viridis var. rufula (Ren. & Card.) 


Vesicularia angustifolia Dix. ZAI(65). 

Vesicularia annobonensis Broth. in Mildbr., nom. 

nud. ANN(47). 

Vesicularia ayresii (Besch.) Broth. MAD(40), 

MAU(105). 

Vesicularia bescherellei (Ren.) Broth. MAU(105). 

Vesicularia codonopyxis (C.Müll.) Broth. 

FER(47). 

Vesicularia combae (Ren. & Card.) Broth. 

MAD(40). 

Vesicularia crassiramea (Ren. & Card.) Broth. 

COM(40), MAD(40). 

177

178 

Vesicularia crassiramea var. laxifolia (Ren. & 

Card.) Card. MAD(40). 

Vesicularia densiramea Broth., nom. nud. 

UGA(46). 

Vesicularia fonticola P.Varde GAB(47). 

Vesicularia galerulata (Duby.) Broth. ANG(46), 

CAM(47), CBS(1531), COM(40), DIE(40), 

GAB(47), IVO(63,1880), MAD(40), MAU(105), 

NAT(1531), NIG(334), REU(105), SWA(1531), 

TAN(46), TRV(1531), UGA(46), ZAI(65), 

ZAM(62), ZIM(61). 

Vesicularia galerulata var. subpiligera (Ren. & 

Par.) Wijk & Marg. MAD(40). 

Vesicularia glaucissima (C.Müll.) Broth. 

COM(40). 

Vesicularia glaucula (Broth.) Broth. CAM(47), 

CAR(47), CON(47), GAB(47), IVO(1880), 

SAO(47), TOG(1878), ZAI(65). 

Vesicularia golungensis (Welw. & Duby) Broth. 

ANG(46). 

Vesicularia hapalyptera (C.Müll.) Broth. 

GAB(47). 

Vesicularia hygrobium (Besch.) Broth. MAD(40), 

MAS(40). 

Vesicularia immunda (Ren.) Card. MAD(40). 

Vesicularia ischyropteris (Broth.) C.Müll. 

CAM(47), CAR(47), GAB(47), GHA(47), 

TOG(1878). 

Vesicularia latiramea Broth. CAR(47), GAB(47), 

IVO(1880), TAN(46). 

Vesicularia lepervanchei (Besch.) Broth. 

COM(40), REU(105). 

Vesicularia longo-fluitans (Broth.) Broth. 

CAM(47). 

Vesicularia longo-fluitans var. latifolia P.Varde 

CAR(47). 

Vesicularia mayumbensis (Besch.) Broth. 

CAM(47), CAR(47), CON(47), GAB(47), 

GUI(47). 

Vesicularia montagnei (Bel.) Broth. DIE(40). 

Vesicularia nigeriana Broth. & Par. CAR(47), 

MAI(47), TAN(46), UGA(46), ZAI(65). 

Vesicularia oreadelphus (Broth.) Broth. CAM(47), 

CAR(47), CON(47), GAB(47), GUI(47), IVO(47), 

RWA(65), TAN(46), ZAI(65). 

Vesicularia oreadelphus var. glauco-viridis 

(Broth.) Par. IVO(47). 

Vesicularia oreadelphus var. sulphurea-flava 

(Broth.) Broth. CAM(47). 

Vesicularia perpallida (Broth.) Broth. CAM(47), 

ZAI(65). 

Vesicularia rodriguezii (Ren. & Card.) Broth. 

REU(105). 

Vesicularia rootii (Par. & Broth.) Broth. MAI(47). 

Vesicularia sarcoblasta (Broth.) Broth. CAM(47), 

CAR(47). 

Vesicularia scaturigina (Brid.) Broth. FER(47), 

NIG(47), PRI(47), REU(105), ROD(105). 

Vesicularia sigmangia (Broth.) Broth. CAM(47), 

RWA(65). 

Vesicularia soyauxii (C.Müll.) Broth. CON(47), 

GAB(47). 

Vesicularia sphaerocarpa (Jaeg.) Broth. ZAI(65). 

Vesicularia strephomischos (Welw. & Duby) 

Broth. ANG(46). 

Vesicularia subenervis (Mitt.) Broth. ROD(105). 

Vesicularia subsarcoblasta (Broth.) Broth. 

CAM(47). 

Vesicularia subsphaerica (C.Müll.) Broth. 

REU(105). 

Vesicularia tenaci-inserta (C.Müll.) Broth. 

CAR(47), CON(47), GAB(47), IVO(1880). 

Vesicularia tenuatipes (C.Müll.) Broth. CAR(47), 

GAB(47), IVO(1880), NIG(47). 

Vesicularia terrestris (C.Müll.) Broth. FER(47). 

Vesicularia thollonii (Besch.) Broth. CON(47). 

Vittia pachyloma (Mont.) Ochyra CBS(1531), 

NAT(1531), TRV(1531). 

Warburgiella adhaerens (Besch.) Broth. 

REU(105). 

Warburgiella leptorrhyncha (Jaeg.) Broth. 

KEN(46), MAD(40), MAU(105), REU(105), 

TAN(46), ZAI(65). 

Wardia hygrometrica Harv. & Hook. CBS(1531). 

Warnstorfia exannulata var. exannulata (BS&G) 

Loeske KEN(46), RWA(65), ZAI(65). 

Warnstorfia fluitans var. fluitans (Hedw.) Loeske 

FER(47), RWA(65,46), UGA(46). 

Webera cavifolia Bruch. & Schimp. ex Par., nom. nud. 

in sched. See note under Pohlia sp. 

Webera lacouturei Thér. = Pohlia lacouturei (See 

“New Combinations”). 

Webera philibertii Par., nom. nud. See note under 

Pohlia sp. 

Weisiopsis involuta Magill BOT(1531), 

NAM(1531). 

Weisiopsis nigeriana (Egun. & Olar.) Zander 

TAN(46), ZAM(46). 

Weisiopsis plicata (Mitt.) Broth. MAD(40), 

TAN(46), TRV(1531), ZAM(62), ZIM(46,61).

Weisiopsis pulchriretis Dix. NAT(1531). 

Weissia artocosana Zander SOC(40,1626). 

Weissia ayresii Schimp. in Besch. MAD(40), 

MAU(105), REU(105), RWA(65). 

Weissia bizotii Zander TAN(46). 

Weissia brachypelma C.Müll. KEN(46). 

Weissia brachypoma Townsend TAN(46). 

Weissia controversa var. controversa Hedw. 

CBS(1531), KEN(46), LES(1531), NAT(1531), 

NIG(334), OFS(1531), REU(105), TAN(46), 

TRV(1531), UGA(46), ZIM(46). 

Weissia controversa var. gymnostoma (Dix.) 

Sainsb. TAN(46). 

Weissia controversa var. macrophylla (Thér. & 

P.Varde) Wijk & Marg. KEN(46). 

Weissia cucullata C.Müll. CBS(1531). 

Weissia dieterlenii Thér. CBS(1531), LES(1531), 

OFS(1531). 

Weissia humicola C.Müll. CBS(1531), TAN(46), 

ZIM(46). 

Weissia kilimandscharica Broth., nom. nud. 

TAN(46). 

Weissia latiuscula C.Müll. CBS(1531), MOZ(46), 

NAM(1531), NAT(1531), TRV(1531), ZAM(62), 

ZIM(46,61). 

Weissia lineaefolia C.Müll. CAR(47). 

Weissia opaca (Dix.) Magill in Magill & Schelpe 

TAN(46), ZAM(62), ZIM(46,61). 

Weissia papillosa Dix. & Nav. ANG(46), ZAI(65). 

Weissia ricciae Brid. REU(105). 

Weissia rutilans (Hedw.) Lindb. ETH(50). 

Weissia socotrana Mitt. ETH(50), KEN(46), 

SOC(40), ZIM(46). 

Weissia unguiculata (Mitt.) Crundw. & Nyh. 

CAR(47), MAL(67), TAN(46), ZAM(62). 

Wijkia albescens (Thér. & P.Varde) Crum CAR(47), 

GAB(47). 

Wijkia bessonii (Ren. & Card.) Crum MAD(40). 

Wijkia cuynetii (Biz.) Schultze-Motel CAM(47), 

IVO(44), KEN(46), TAN(46). 

Wijkia jungneri (Broth.) Crum BUR(65), CAM(47), 

GAB(47), IVO(1880), TAN(46), ZAI(65). 

Wijkia le-testui (Thér. & P.Varde) Crum GAB(47). 

Wijkia madagassa (Thér.) Crum MAD(40). 

Wijkia protensa (Ren. & Card.) Crum REU(105). 

Wijkia rutenbergii (C.Müll.) Crum MAD(40), 

TAN(46). 

Wijkia rutenbergii var. eflagellaris (Card.) Crum 

MAD(40). 

Wijkia trichocolea (C.Müll.) Crum GAB(47), 

KEN(46), LES(1531), MOZ(46), RWA(65), 

TAN(46), TOG(1878), TRV(1531), UGA(46), 

ZAI(65), ZAM(62), ZIM(46). 

Wijkia trichocolea ssp. subintegrifolia (Thér. & 

P.Varde) Crum GAB(47). 

Wijkia trichocoleoides (C.Müll.) Crum CAM(47), 

KEN(46), RWA(65), SAO(47), TAN(46), 

ZAI(65). 

Wijkiella kenyae Bizot & Lewinsky KEN(46). 

Wilsoniella crispidens C.Müll. ex Broth. CAM(47). 

Zelometeorium ambiguum (Hornsch. in Mart.) 

Manuel ZAI(1890). 

Zygodon barbuloides Broth. ex Malta TAN(46). 

Zygodon bolleanus C.Müll. Excluded. Wijk et al 

(1521) say this is probably an Anoectangium 

(presumably indeterminate). 

Zygodon conoideus var. latifolius Brid. MAU(105). 

Zygodon dioicus Broth. RWA(65), TAN(46), 

ZAI(65). 

Zygodon dixonii Sim NAT(1531). 

Zygodon erosus Mitt. KEN(46), NAT(1531), 

TAN(46), TRV(1531), ZAI(65). 

Zygodon erosus var. circinatifolius Thér. & Nav. 

ZAI(65). 

Zygodon gemmaceus Dix. UGA(46). 

Zygodon hirsutus Negri UGA(46). 

Zygodon intermedius BS&G CAM(47), 

KEN(46,136), REU(758), RWA(65), TAN(46), 

UGA(46), ZAI(65), ZAM(62). 

Zygodon intermedius var. dusenii Malta CAM(47). 

Zygodon intermedius var. strictifolius Thér. 

RWA(65), UGA(46), ZAI(65). 

Zygodon johnstonii Broth. ex Malta TAN(46). 

Zygodon laxifolius Thér. MAD(40). 

Zygodon leptobolax C.Müll. CBS(1531), 

NAT(1531), TRV(1531). 

Zygodon leptocarpus Thér. MAD(40). 

Zygodon microtheca Dix. ex Malta KEN(46), 

RWA(65), TAN(46), UGA(46), ZAI(65). 

Zygodon obtusifolius Hook. ZAI(65). 

Zygodon pungens C.Müll. TAN(46). 

Zygodon reinwardtii (Hornsch.) Braun KEN(46), 

MAD(40), REU(105), TAN(46), UGA(46), 

ZAI(65). 

Zygodon reinwardtii var. madacassus Card. 

MAD(40). 

Zygodon reinwardtii var. ruwenzorensis (Thér. 

& Nav.) Demar. ZAI(65). 

Zygodon robustus Broth. ex Malta TAN(46). 

Zygodon roccatii Negri RWA(65), UGA(46). 

179

180 

Zygodon roccatii var. diffusus Negri UGA(46). 

Zygodon runcinatus C.Müll. CBS(1531), TAN(46). 

Zygodon schimperi Hampe ex C.Müll. ETH(50). 

Zygodon semitortus Mitt. CAM(47), ZAI(65). 

Zygodon seriatus Thér. & Nav. ETH(50), KEN(46), 

RWA(46,65), TAN(46), UGA(46), ZAI(65). 

Zygodon trichomitrius Hook. & Wils. CBS(1531), 

NAT(1531), TAN(46), TRV(1531), ZIM(61). 

Zygodon trichomitrius var. mildbraedii Broth. 

RWA(65), TAN(46), UGA(46), ZAI(65) 

Acknowledgements 

I have received assistance on this project from a large 

number of people, but I would particularly like to thank 

Len Ellis for the time he has spent in searching out 

specimens and references, and Martin Wigginton for 

discussions especially in the early days of the checklist, and 

many pointers to literature references. Thanks also go 

to those who have commented on the final family 

treatments: Ida Bruggeman-Nannenga (Fissidentaceae), 

Len Ellis (Calymperacae), Johannes Enroth 

(Neckeraceae), Allan Fife (Funariaceae), Jan-Peter Frahm 

(Dicranaceae), Angela Newton (Pterobryaceae), Ryszard 

Ochyra (Amblystegiaceae, Brachytheciaceae, Grimmia- 

ceae), Philip Sollman (Pottiaceae), Dries Touw 

(Thuidiaceae s.l.), and to Tamás Pócs, who gave thorough 

comments on the whole paper in a very short space of 

time. As I did not always accept the advice given, any 

mistakes are mine. 

References 

Use this table to translate the numeric references given in 

the text of this paper to the ‘author + year’ equivalent. 

The alphabetic list of references follows the table. 

(1) Argent 1973a 

(2) Argent 1973b 

(4) Bizot 1976 

(12) Buck 1979 

(15) De Sloover 1983 

(16) Dixon 1938 

(19) Frahm 1982 

(21) Mohamed 1979 

(23) Ochi 1972 

(24) Orbán 1981 

(28) Richards et al. 1968 

(30) Snider 1975 

(32) Touw 1976 

(40) Crosby et al. 1983 

(44) Frahm 1984 

(46) Kis 1985 

(47) Schultze-Motel 1975 

(50) Cufodontis 1951 

(51) Jelenc 1955. 

(52) Jelenc 1967 

(53) Magill et al. 1979 

(54) Frahm 1985 

(56) Hyvönen 1989 

(57) Edwards 1980 

(58) Schultze-Motel 1979 

(59) Eddy 1985 

(61) Best 1990 

(62) Phiri et al. 1985 

(63) Prendergast 1984 

(64) El-Saadawi et al. 1977 

(65) Born et al. 1993 

(66) O’Shea 1994 

(67) O’Shea 1993a 

(68) Arts et al. 1990 [1993] 

(70) Townsend 1991 

(71) Corley et al. 1981 

(72) Bruggeman-Nannenga et al. 1990a 

(73) Bruggeman-Nannenga et al. 1990b 

(74) Bizot et al.1979 

(75) Pursell 1979 

(76) Pursell et al. 1992 

(77) Bridel-Brideri 1827 

(79) Tixier 1981 

(80) Crosby et al. 1992 

(81) Delgadillo M. 1973 

(82) Magill 1981 

(83) Magill 1987a 

(84) Wigginton (in press) 

(85) Pursell 1987 

(89) Bruggeman-Nannenga 1978 

(90) Bruggeman-Nannenga 1993 

(93) Allen et al. 1986a 

(94) Allen et al. 1986b 

(105) Een 1993b 

(107) Bizot et al. 1980 

(108) Bizot et al. 1982 

(115) O’Shea 1993b 

(123) Koponen et al. 1972 

(132) Orbán et al. 1986 

(134) Pócs 1993

(136) Pócs et al. 1983 

(146) Ochyra et al. 1988 

(162) Ando 1985 

(169) Ando 1989 

(175) Buck et al. 1986 

(182) Griffin III et al. 1989 

(225) Griffin III et al. 1989 

(255) Bizot 1968. 

(259) Bizot et al. 1986 

(273) Crum et al. 1972 [1973] 


(286) De Sloover 1977a 

(287) De Sloover 1977b 



(298) Een 1989 

(299) Enroth 1991a 

(301) Enroth 1992a 

(302) Enroth 1992b 

(333) Jones 1985a 

(334) Jones 1985b 

(347) Lewinsky 1978 

(348) Lin 1983 

(349) Long 1986 

(354) Ochi 1973 

(355) Ochyra et al. 1982 

(357) Ochyra et al. 1985 [1987] 

(361) Orbán 1977 

(362) Orbán 1978 

(363) Orbán 1987 

(400) Richards et al 1972 

(408) Stark 1987 

(411) Townsend 1987 

(453) Tuomikoski et al. 1979 

(465) Padberg et al. 1985. 

(489) Crundwell et al. 1962 

(506) Giese et al. 1985 

(507) Giese et al. 1985 

(510) Enroth et al. 1994 

(580) Dirkse et al. 1991 

(679) Ando 1993 

(715) Buck 1980 

(717) Buck 1993 

(719) Buck et al. 1985 

(755) Eddy 1990 

(758) Een 1993 

(761) Egunyomi et al. 1982 

(762) Egunyomi et al. 1977 

(765) Ellis 1991a 

(767) Ellis 1991b 

(780) Enroth 1991b 

(781) Enroth 1991c 

(785) Enroth 1993b 

(787) Enroth 1993a 

(804) Frahm 1993 

(829) Griffin III 1992 

(924) He et al. 1992 

(1036) Koponen 1981 

(1065) Lewinsky et al. 1990 

(1074) Magill 1987b 

(1124) Ochyra 1993 

(1125) Orbán 1993 

(1149) Salazar Allen 1993b 

(1157) Sollman 1990 

(1158) Sollman 1993 

(1202) Zomlefer 1993 

(1291) Manuel 1977b 

(1294) Een 1978a 

(1295) Hedenäs 1992 

(1296) Geheeb 1910 

(1300) Enroth 1990 

(1336) Koponen 1993 

(1339) Buck 1989 

(1341) Whittemore et al. 1989 

(1344) Orbán et al. 1990 

(1346) Tixier, P. 1992 

(1347) Ochyra et al. 1992e 

(1349) Ireland et al. 1994 

(1350) Whitehouse et al. 1991 

(1357) Corley et al. 1991 

(1370) Ellis 1987 

(1371) Ellis 1988a 

(1372) Ellis 1988b 

(1373) Ellis 1989a 

(1374) Ellis 1989b 

(1375) Reese 1991 

(1386) Enroth 1989 

(1393) Frey et al. 1988a 

(1403) Ochyra et al. 1992a 

(1404) Ochyra et al. 1992b 

(1405) Ochyra et al. 1992c 

(1406) Ochyra et al. 1992d 

(1407) Ochyra et al. 1993a 

(1408) Ochyra et al. 1993b 

(1409) Ochyra et al. 1993c 

(1410) Ochyra et al. 1993d 

(1419) Townsend 1984 [1985] 

(1420) Zander 1993 

(1421) Sollman 1983 

(1423) Prendergast 1983 

181

182 

(1424) Vitt 1980 

(1437) Stevenson 1992 

(1438) Van Rooy 1990 

(1439) Mattila et al. 1990 

(1442) Ochyra 1989 

(1447) Van Rooy et al. 1992 

(1448) Van Rooy 1991b 

(1458) Frahm et al. 1994. 

(1460) Sollman 1994 

(1470) Pursell et al. 1993 

(1471) Snider 1987 

(1473) Reese 1987 

(1474) Reese et al. 1987 

(1475) Frahm 1986 

(1477) Fife 1986. 

(1485) Reese et al. 1985 

(1487) Shaw 1985 

(1492) Egunyomi 1984. 

(1494) Van Rooy 1991a 

(1497) Spence et al. 1988 

(1500) Ochyra et al. 1985 

(1502) Touw 1983. 

(1511) Ellis 1991c 

(1517) Crosby et al. 1993 

(1518) Ellis (in press)a 

(1519) Magill (in press) 

(1520) Syed 1973 

(1521) Wijk et al. 1959-1969 

(1527) Beever et al. 1992 

(1531) Perold et al. 1993 

(1532) Shaw et al. 1984 

(1533) Ochyra 1994 

(1547) Van Rooy 1992 

(1552) Leonard 1993 

(1554) Catcheside et al. 1980 

(1562) Frahm 1994 

(1567) Potier de la Varde 1939 

(1569) Kürschner et al. 1993 

(1579) Mohamed et al. 985 

(1589) Burley et al. 1990 

(1593) Vitt 1984 

(1608) Frey et al. 1988b 

(1617) LaFarge-England 1992 

(1620) Saito 1975 

(1621) Ochyra 1992 

(1624) Reese et al. 1986 

(1625) Bescherelle 1895. 

(1626) Townsend 1969 

(1627) Mitten 1887 [1888] 

(1629) De Sloover 1982 

(1631) Townsend 1978 

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190 

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Appendix A - Country Abbreviations 

There are 68 countries, part-countries or islands on the 

list. 

Abb Full name Group/ Index Muscorum 

zone 

ALD Aldabra Africa-3 

AMI Amirante Islands Africa-3 

ANG Angola Africa-2 

ANN Annobon Is. Equatorial Guinea 

ASC Ascension Is. Africa-2 

BEN Benin Africa-2 

BOT Botswana Africa-4 

BUF Burkina Faso Africa-2 

BUR Burundi Africa-2 

CAB Cabinda Angola 

CAM Cameroon Africa-2 

CBS Cape (of Good Hope) South Africa 

CAP Cape Verde Africa-2 

CAR Central African Rep. Africa-2 

CHA Chad Africa-2 

CHG Chagos Archipelago Africa-3 

COM Comoros Africa-3 

CON CongoAfrica-2 

DIE Diego Garcia Africa-3 

DJI Djibouti Africa-2 

EGU Equatorial Guinea Africa-2 

ERI Eritrea Africa-2 

ETH Ethiopia Africa-2 

FER Fernando Po (Bioko) Equatorial Guinea 

GAB Gabon Africa-2 

GAM Gambia Senegambia 

GHA Ghana Africa-2 

GUB Guinea Bissau Africa-2 

GUI GuineaAfrica-2 

IVO Ivory Coast Africa-2 

KEN Kenya Africa-2

LES Lesotho Africa-4 

LIB LiberiaAfrica-2 

MAD Madagascar Africa-3 

MAL Malawi Africa-2 

MAI Mali Africa-2 

MAS Mascarene Islands Africa-3 

MAU Mauritius Mascarenes 

MOZ Mozambique Africa-2 

NAM Namibia Africa-4 

NAT Natal South Africa 

NIR Niger Africa-2 

NIG Nigeria Africa-2 

OFS Orange Free State South Africa 

PRI Principe Africa-2 

REU Reunion Mascarenes 

RIM Rio Muni Equatorial Guinea 

ROD Rodrigues Mascarenes 

RWA Rwanda Africa-2 

SAF South Africa Africa-4 

SAO Sao Tome Africa-2 

SEN Senegal Senegambia 

SEG Senegambia Africa-2 

SEY Seychelles Africa-3 

SIL Sierra Leone Africa-2 

SOC Socotra Africa-2 

SOM Somalia Africa-2 

PAU Sao Paulo Africa-2 

STH St. Helena Africa-2 

SUD Sudan Africa-2 

SWA Swaziland Africa-4 

TAN Tanzania Africa-2 

TOG Togo Africa-2 

TRV Transvaal South Africa 

UGA Uganda Africa-2 

ZAI Zaire Africa-2 

ZAM Zambia Africa-2 

ZIM Zimbabwe Africa-2 

Appendix B - Family Placement 

There are 75 families and 353 genera on the checklist. 

Family placement is based on Crosby & Magill (ref. 1752), 

and family sequence on Vitt (1593) with later amendments 

from Buck & Ireland (719), Buck & Vitt (175), Buck (715) 

and Whittemore and Allen (1341). 

Sphagnaceae 

Sphagnum 

Andreaeaceae 

Andreaea 

Polytrichaceae 

Atrichum 

Oligotrichum 

Pogonatum 

Polytrichastrum 

Polytrichum 

Archidiaceae 

Archidium 

Funariaceae 

Cygnicollum 

Entosthodon 

Funaria 

Goniomitrium 

Loiseaubryum 

Physcomitrella 

Physcomitrellopsis 

Physcomitrium 

Gigaspermaceae 

Chamaebryum 

Gigaspermum 

Oedipodiella 

Ephemeraceae 

Ephemerum 

Splachnaceae 

Brachymitrion 

Splachnum 

Tayloria 

Tetraplodon 

Splachnobryaceae 

Splachnobryum 

Orthotrichaceae 

Amphidium 

Cardotiella 

Groutiella 

Kleioweisiopsis 

Leiomitrium 

Leptodontiopsis 

Macrocoma 

Macromitrium 

191

192 

Orthotrichum 

Schlotheimia 

Stoneobryum 

Ulota 

Zygodon 

Erpodiaceae 

Aulacopilum 

Erpodium 

Rhachitheciaceae 

Jonesiobryum 

Rhachithecium 

Tisserantiella 

Bryaceae 

Anomobryum 

Brachymenium 

Bryum 

Epipterygium 

Leptobryum 

Mielichhoferia 

Mniobryum 

Orthodontium 

Plagiobryum 

Pohlia 

Pseudopohlia 

Rhodobryum 

Schizymenium 

Mniaceae 

Plagiomnium 

Bartramiaceae 

Anacolia 

Bartramia 

Breutelia 

Conostomum 

Flowersia 

Leiomela 

Philonotis 

Plagiopus 

Quathlamba 

Aulacomniaceae 

Aulacomnium 

Meesiaceae 

Meesia 

Rhizogoniaceae 

Leptotheca 

Pyrrhobryum 

Racopilaceae 

Racopilum 

Hypopterygiaceae 

Hypopterygium 

Lopidium 

Buxbaumiaceae 

Diphyscium 

Encalyptaceae 

Encalypta 

Pottiaceae 

Acaulon 

Aloina 

Anoectangium 

Astomum 

Barbula 

Bellibarbula 

Bryoceuthospora 

Bryoerythrophyllum 

Chenia 

Crossidium 

Didymodon 

Eucladium 

Gymnostomiella 

Gymnostomum 

Hennediella 

Husnotiella 

Hymenostomum 

Hymenostylium 

Hyophila 

Hypodontium 

Leptodontium 

Microbryum 

Molendoa 

Phascum 

Pleurochaete 

Pottia 

Pseudocrossidium 

Pseudosymblepharis 

Pterygoneurum 

Scopelophila 

Streptopogon 

Syntrichia

193 

Tetrapterum 

Timmiella 

Tortella 

Tortula 

Trachycarpidium 

Trichostomum 

Triquetrella 

Weisiopsis 

Weissia 

Cinclidotaceae 

Cinclidotus 

Serpotortellaceae 

Serpotortella 

Calymperaceae 

Calymperes 

Calymperopsis 

Chameleion 

Leucophanes 

Mitthyridium 

Octoblepharum 

Syrrhopodon 

Bryobartramiaceae 

Bryobartramia 

Dicranaceae 

Anisothecium 

Aongstroemia 

Aongstroemiopsis 

Atractylocarpus 

Brothera 

Bruchia 

Bryohumbertia 

Bryotestua 

Campylopus 

Chorisodontium 

Cladophascum 

Cynodontium 

Dicranella 

Dicranodontium 

Dicranoloma 

Dicranoweisia 

Dicranum 

Holomitrium 

Leucoloma 

Microcampylopus 

Microdus 

Ochrobryum 

Oreoweisia 

Paraleucobryum 

Pilopogon 

Pocsiella 

Pseudephemerum 

Rhabdoweisia 

Sphaerothecium 

Symblepharis 

Wilsoniella 

Leucobryaceae 

Leucobryum 

Ditrichaceae 

Astomiopsis 

Ceratodon 

Cheilothela 

Distichium 

Ditrichum 

Eccremidium 

Garckea 

Pleuridium 

Saelania 

Trematodon 

Tristichium 

Eustichiaceae 

Eustichia 

Phyllodrepaniaceae 

Phyllodrepanium 

Nanobryaceae 

Nanobryum 

Fissidentaceae 

Fissidens 

Moenkemeyera 

Seligeriaceae 

Blindia 

Brachydontium 

Ptychomitriaceae 

Ptychomitriopsis 

Ptychomitrium 

Grimmiaceae 

Grimmia

194 

Leucoperichaetium 

Racomitrium 

Schistidium 

Fontinalaceae 

Fontinalis 

Echinodiaceae 

Isothecium 

Thuidiaceae 

Haplocladium 

Pelekium 

Rauiella 

Thuidium 

Trachyphyllum 

Rigodiaceae 

Rigodium 

Leskeaceae 

Hylocomiopsis 

Leptopterigynandrum 

Leskea 

Leskeella 

Lindbergia 

Pseudoleskea 

Pseudoleskeopsis 

Rhytidiaceae 

Chryso-hypnum 

Gollania 

Rhytidium 

Amblystegiaceae 

Amblystegium 

Campyliadelphus 

Campylium 

Cratoneuron 

Drepanocladus 

Hygrohypnum 

Sanionia 

Sarmentypnum 

Warnstorfia 

Vittiaceae 

Vittia 

Lembophyllaceae 

Camptochaete 

Orthostichella 

Stereophyllaceae 

Entodontopsis 

Juratzkaea 

Stereophyllum 

Fabroniaceae 

Anacamptodon 

Campylodontium 

Dimerodontium 

Fabronia 

Ischyrodon 

Leptoischyrodon 

Levierella 

Rhizofabronia 

Schwetschkea 

Brachytheciaceae 

Aerolindigia 

Brachythecium 

Eurhynchiella 

Eurhynchium 

Homalothecium 

Kindbergia 

Oxyrrhynchium 

Palamocladium 

Platyhypnidium 

Pseudoscleropodium 

Rhynchostegiella 

Rhynchostegium 

Schimperella 

Scorpiurium 

Stenocarpidiopsis 

Myriniaceae 

Helicodontium 

Regmatodontaceae 

Regmatodon 

Entodontaceae 

Entodon 

Erythrodontium 

Mesonodon 

Pylaisiobryum 

Sematophyllaceae 

Acanthorrhynchium 

Acroporium 

Clastobryella 

Clastobryophilum

195 

Donnellia 

Gammiella 

Glossadelphus 

Heterophyllium 

Macrohymenium 

Meiotheciopsis 

Meiothecium 

Potamium 

Radulina 

Rhaphidorrhynchium 

Rhaphidostichum 

Sematophyllum 

Taxithelium 

Trichosteleum 

Trismegistia 

Warburgiella 

Wijkia 

Wijkiella 

Hylocomiaceae 

Hylocomium 

Leptohymenium 

Catagoniaceae 

Catagonium 

Plagiotheciaceae 

Plagiothecium 

Hypnaceae 

Bryosedgwickia 

Ctenidium 

Ectropothecium 

Entodontella 

Herzogiella 

Hypnum 

Isopterygium 

Mittenothamnium 

Pylaisiella 

Rhacopilopsis 

Taxiphyllum 

Trachythecium 

Vesicularia 

Neckeraceae 

Himantocladium 

Homalia 

Homaliodendron 

Neckera 

Neckeropsis 

Pilotrichum 

Pinnatella 

Porothamnium 

Porotrichum 

Thamnobryum 

Cryphaeaceae 

Cryphaea 

Schoenobryum 

Leptodontaceae 

Caduciella 

Cryptoleptodon 

Forsstroemia 

Leptodon 

Anomodontaceae 

Haplohymenium 

Herpetineuron 

Hedwigiaceae 

Braunia 

Hedwigia 

Hedwigidium 

Leucodontaceae 

Antitrichia 

Leucodon 

Pterogonium 

Ptychomniaceae 

Ptychomnion 

Wardiaceae 

Wardia 

Rhacocarpaceae 

Rhacocarpus 

Rutenbergiaceae 

Neorutenbergia 

Rutenbergia 

Prionodontaceae 

Prionodon 

Phyllogoniaceae 

Phyllogonium 

Pterobryaceae

196 

Calyptothecium 

Henicodium 

Hildebrandtiella 

Jaegerina 

Orthostichidium 

Orthostichopsis 

Pterobryon 

Pterobryopsis 

Renauldia 

Trachypodaceae 

Trachypodopsis 

Trachypus 

Meteoriaceae 

Aerobryidium 

Aerobryopsis 

Barbella 

Floribundaria 

Lindigia 

Papillaria 

Pilotrichella 

Squamidium 

Zelometeorium 

Symphyodontaceae 

Symphyodon 

Hookeriaceae 

Actinodontium 

Callicostella 

Chaetomitrium 

Cyclodictyon 

Hookeria 

Hookeriopsis 

Hypnella 

Lepidopilidium 

Lepidopilum 

Leucomium 

Sauloma 

Daltoniaceae 

Adelothecium 

Calyptrochaeta 

Cyathophorella 

Daltonia 

Distichophyllidium 

Distichophyllum

197

198

Checklist of the mosses of sub-Saharan Africa

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