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Catalogue of Tenebrionidae (Coleoptera) of North America
expand article infoYves Bousquet, Donald B. Thomas§, Patrice Bouchard, Aaron D. Smith|, Rolf L. Aalbu, M. Andrew Johnston#, Warren E. Steiner Jr.¤
‡ Agriculture and Agri-Food Canada, Ottawa, Canada
§ USDA ARS, Cattle Fever Tick Research Laboratory, Edinburg, United States of America
| Northern Arizona University, Flagstaff, United States of America
¶ California Academy of Sciences, San Francisco, United States of America
# Arizona State University, Tempe, United States of America
¤ mithsonian Institution, Washington, United States of America
† Deceased author
Open Access

Abstract

This catalogue includes all valid family-group (8 subfamilies, 52 tribes, 14 subtribes), genus-group (349 genera, 86 subgenera), and species-group names (2825 species, 215 subspecies) of darkling beetles (Coleoptera: Tenebrionidae) known to occur in North America1 and their available synonyms. Data on extant, subfossil and fossil taxa are given. For each name the author and year and page number of the description are provided, with additional information (e.g., type species for genus-group names, author of synonymies for invalid taxa) depending on the taxon rank.

Several new nomenclatural acts are included. One new genus, Lepidocnemeplatia Bousquet and Bouchard, is described. Spelaebiosis Bousquet and Bouchard [for Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [for Blapstinus kulzeri Marcuzzi, 1977], and Hymenorus campbelli Bouchard [for Hymenorus oculatus Doyen and Poinar, 1994] are proposed as new replacement names. Supporting evidence is provided for the conservation of usage of Tarpela micans (Fabricius, 1798) nomen protectum over Tarpela vittata (Olivier, 1793) nomen oblitum. The generic names Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 and Euconibius Casey, 1895 [= Conibius LeConte, 1851] are new synonyms (valid names in square brackets). The following 127 new synonymies of species-group names, listed in their original combination, are proposed (valid names, in their current combination, placed in square brackets): Bothrasida mucorea Wilke, 1922 [= Pelecyphorus guanajuatensis (Champion, 1884)]; Parasida zacualpanicola Wilke, 1922 [= Pelecyphorus asidoides Solier, 1836]; Stenosides kulzeri Pallister, 1954, Stenosides bisinuatus Pallister, 1954, and Parasida trisinuata Pallister, 1954 [= Pelecyphorus dispar (Champion, 1892)]; Asida favosa Champion, 1884 and Asida similata Champion, 1884 [= Pelecyphorus fallax (Champion, 1884)]; Ologlyptus bicarinatus Champion, 1884 [= Pelecyphorus indutus (Champion, 1884)]; Parasida laciniata Casey, 1912 and Parasida cristata Pallister, 1954 [= Pelecyphorus liratus (LeConte, 1854)]; Parasida esperanzae Wilke, 1922 and Parasida mixtecae Wilke, 1922 [= Pelecyphorus longipennis (Champion, 1884)]; Parasida tolucana Casey, 1912 [= Pelecyphorus scutellaris (Champion, 1884)]; Parasida purpusi Wilke, 1922 [= Pelecyphorus tristis (Champion, 1884)]; Astrotus nosodermoides Champion, 1892 [= Pelecyphorus erosus (Champion, 1892)]; Astrotus seticornis var. humeralis Champion, 1884 [= Pelecyphorus seticornis (Champion, 1884)]; Pactostoma breviuscula Casey, 1912, Pactostoma exoleta Casey, 1912, Pactostoma luteotecta Casey, 1912, Pactostoma monticola Casey, 1912, Pactostoma obtecta Casey, 1912, and Pactostoma sigillata Casey, 1912 [=Pelecyphorus anastomosis (Say, 1824)]; Ologlyptus canus Champion, 1884 and Ologlyptus sinuaticollis Champion, 1884 [= Pelecyphorus graciliformis (Solier, 1836)]; Gonasida elata reducta Casey, 1912, Gonasida elata prolixa Casey, 1912, and Gonasida aucta Casey, 1912 [= Philolithus elatus compar (Casey, 1912)]; Gonasida alaticollis Casey, 1912 [= Philolithus elatus difformis (LeConte, 1854)]; Gonasida gravida Casey, 1912 [= Philolithus elatus elatus (LeConte, 1853)]; Pelecyphorus aegrotus limbatus Casey, 1912 [= Philolithus aegrotus aegrotus (LeConte, 1861)]; Pelecyphorus corporalis Casey, 1912, Pelecyphorus reptans Casey, 1912, Pelecyphorus socer Casey, 1912, Pelecyphorus abscissus Casey, 1912, Pelecyphorus fumosus Casey, 1912, Pelecyphorus parvus Casey, 1912, Pelecyphorus morbillosus pacatus Casey, 1912, Pelecyphorus morbillosus sobrius Casey, 1912, Pelecyphorus piceus Casey, 1912, Pelecyphorus piceus crudelis Casey, 1912, Pelecyphorus snowi Casey, 1912, and Pelecyphorus subtenuis Casey, 1912 [= Philolithus morbillosus (LeConte, 1858)]; Bothrasida sanctae-agnae Wilke, 1922 [= Stenomorpha funesta (Champion, 1884)]; Asida flaccida Horn, 1896 [= Stenomorpha embaphionides (Horn, 1894)]; Asida angustula Casey, 1890, Stethasida stricta Casey, 1912, Stethasida muricatula languida Casey, 1912, Stethasida pertinax Casey, 1912, Stethasida socors Casey, 1912, Stethasida angustula inepta Casey, 1912, Stethasida tenax Casey, 1912, and Stethasida vegrandis Casey, 1912 [= Stenomorpha muricatula (LeConte, 1851)]; Stethasida obsoleta expansa Casey, 1912, Stethasida obsoleta opacella Casey, 1912, Stethasida brevipes Casey, 1912, Stethasida torpida Casey, 1912, Stethasida convergens Casey, 1912, Stethasida discreta Casey, 1912, Stethasida longula Casey, 1912, Stethasida adumbrata Casey, 1912, Stethasida occulta Casey, 1912, Stethasida tarsalis Casey, 1912, Stethasida unica Casey, 1912, and Pelecyphorus laevigatus Papp, 1961 [= Stenomorpha obsoleta (LeConte, 1851)]; Trichiasida eremica Wilke, 1922 [= Stenomorpha difficilis (Champion, 1884)]; Trichiasida lineatopilosa Casey, 1912 [= Stenomorpha hirsuta (LeConte, 1851)]; Trichiasida tenella Casey, 1912 [= Stenomorpha hispidula (LeConte, 1851)]; Trichiasida duplex Casey, 1912 [= Stenomorpha villosa (Champion, 1884)]; Alaudes squamosa Blaisdell, 1919, Alaudes testacea Blaisdell, 1919, and Alaudes fallax Fall, 1928 [= Alaudes singularis Horn, 1870]; Edrotes barrowsi Dajoz, 1999 [=Edrotes ventricosus LeConte, 1851]; Nyctoporis tetrica Casey, 1907 and Nyctoporis maura Casey, 1907 [= Nyctoporis aequicollis Eschscholtz, 1831]; Nyctoporis pullata Casey, 1907 [= Nyctoporis sponsa Casey, 1907]; Eleodes tibialis forma oblonga Blaisdell, 1909 [= Eleodes tibialis Blaisdell, 1909]; Eleodes (manni var.) variolosa Blaisdell, 1917 [= Eleodes constrictus LeConte, 1858]; Eleodes cordata forma sublaevis Blaisdell, 1909, Eleodes cordata forma intermedia Blaisdell, 1909, Eleodes cordata forma oblonga Blaisdell, 1909, Eleodes cordata forma elongata Blaisdell, 1909, and Eleodes (cordata var.) adulterina Blaisdell, 1917 [= Eleodes cordata Eschscholtz, 1829]; Eleodes hornii var. monticula Blaisdell, 1918 and Eleodes manni sierra Blaisdell, 1925 [= Eleodes fuchsii Blaisdell, 1909]; Eleodes parvicollis var. squalida Blaisdell, 1918 [= Eleodes parvicollis Eschscholtz, 1829]; Eleodes reflexicollis Mannerheim, 1843 and Eleodes parvicollis forma farallonicaBlaisdell, 1909 [= Eleodes planata Eschscholtz, 1829]; Eleodes indentata Blaisdell, 1935 [= Eleodes rotundipennis LeConte, 1857]; Eleodes intricata Mannerheim, 1843 [= Eleodes scabrosa Eschscholtz, 1829]; Eleodes horni fenyesi Blaisdell, 1925 [= Eleodes tenebrosa Horn, 1870]; Eleodes cordata var. horrida Blaisdell, 1918 [= Eleodes tuberculata Eschscholtz, 1829]; Eleodes oblonga Blaisdell, 1933 [= Eleodes versatilis Blaisdell, 1921]; Eleodes dentipes marinae Blaisdell, 1921 [= Eleodes dentipes Eschscholtz, 1829]; Eleodes carbonaria forma glabra Blaisdell, 1909 [= Eleodes carbonaria carbonaria (Say, 1824)]; Eleodes granosa forma fortis Blaisdell, 1909 [= Eleodes granosa LeConte, 1866]; Eleodes pilosa forma ordinata Blaisdell, 1909 [= Eleodes pilosa Horn, 1870]; Trogloderus costatus pappi Kulzer, 1960 [= Trogloderus tuberculatus Blaisdell, 1909]; Trogloderus costatus mayhewi Papp, 1961 [= Trogloderus vandykei La Rivers, 1946]; Bolitophagus cristatus Gosse, 1840 [= Bolitotherus cornutus (Fabricius, 1801)]; Eleates explanatus Casey, 1890 [= Eleates depressus (Randall, 1838)]; Blapstinus sonorae Casey, 1890 [= Blapstinus brevicollis LeConte, 1851]; Blapstinus falli Blaisdell, 1929 [= Blapstinus castaneus Casey, 1890]; Blapstinus brunneus Casey, 1890 and Blapstinus coronadensis Blaisdell, 1892 [=Blapstinus histricus Casey, 1890]; Blapstinus hesperius Casey, 1890 [=Blapstinus intermixtus Casey, 1890]; Blapstinus cinerascens Fall, 1929 [= Blapstinus lecontei Mulsant and Rey, 1859]; Blapstinus niger Casey, 1890 and Blapstinus cribricollis Casey, 1890 [= Blapstinus pimalis Casey, 1885]; Blapstinus arenarius Casey, 1890 [= Blapstinus pratensis LeConte, 1859]; Blapstinus gregalis Casey, 1890 [= Blapstinus substriatus Champion, 1885]; Blapstinus hydropicus Casey, 1890 [= Blapstinus sulcatus LeConte, 1851]; Blapstinus hospes Casey, 1890 [= Blapstinus vestitus LeConte, 1859]; Notibius reflexus Horn, 1894 [= Conibius opacus (LeConte, 1866)]; Notibius affinis Champion, 1885 [=Conibius rugipes (Champion, 1885)]; Conibius parallelus LeConte, 1851 [= Conibius seriatus LeConte, 1851]; Nocibiotes rubripes Casey, 1895 [=Nocibiotes caudatus Casey, 1895]; Nocibiotes gracilis Casey, 1895 and Nocibiotes acutus Casey, 1895 [=Nocibiotes granulatus (LeConte, 1851)]; Conibius alternatus Casey, 1890 [= Tonibius sulcatus (LeConte, 1851)]; Pedinus suturalis Say, 1824 [= Alaetrinus minimus (Palisot de Beauvois, 1817)]; Menedrio longipennis Motschulsky, 1872 [= Tenebrio obscurus Fabricius, 1792]; Hymenophorus megops Hatch, 1965 and Telesicles magnus Hatch, 1965 [= Hymenorus sinuatus Fall, 1931]; Andrimus concolor Casey, 1891 and Andrimus convergens Casey, 1891 [= Andrimus murrayi (LeConte, 1866)]; Mycetochara marshalli Campbell, 1978 [= Mycetochara perplexata Marshall, 1970]; Phaleria globosa LeConte, 1857 [= Phaleria picta Mannerheim, 1843]. The following subspecies of Trogloderus costatus LeConte, 1879 are given species rank: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909, and Trogloderus vandykei La Rivers, 1946. The following taxa, previously thought to be junior synonyms, are considered valid: Amphidora Eschscholtz, 1829; Xysta Eschscholtz, 1829; Helops confluens (Casey, 1924). Two new combinations are proposed: Stenomorpha spinimana (Champion, 1892) and Stenomorpha tenebrosa (Champion, 1892) [from the genus Parasida Casey, 1912]. The type species [placed in square brackets] of the following 12 genus-group taxa are designated for the first time: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant and Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Two species previously known from South America [Nilio lebasi J. Thomson and Platydema erotyloides Chevrolat] are reported for the first time from North America.

Resumen

Este catálogo incluye todos los grupos válidos de nivel famili (8 subfamilias, 52 tribus, 14 subtribus), de nivel género (349 géneros, 86 subgéneros) y de nivel especie (2825 especies, 215 subespecies) de tenebriónidos (Coleoptera: Tenebrionidae) presentes en América del Norte y sus sinónimos disponibles. Se dan datos sobre los táxones existentes, subfósiles y fósiles. Para cada nombre, el autor, el año y el número de página de la descripción se complementan con información adicional dependiendo de la categoría del taxon (por ejemplo, especies tipo para nombres de nivel género, autor de sinonimias para táxones no válidos).

Se incluyen varios nuevos actos nomenclaturales. Un nuevo género, Lepidocnemeplatia Bousquet y Bouchard, se describe. Spelaebiosis Bousquet y Bouchard [para Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [para Blapstinuskulzeri Marcuzzi, 1977] e Hymenorus campbelli Bouchard [para Hymenorus oculatus Doyen y Poinar, 1994] se proponen como nuevos nombres de reemplazo. Se proporciona evidencia de uso para la conservación de Tarpela micans (Fabricius, 1798) nomen protectum sobre Tarpela vittata (Olivier, 1793) nomen oblitum. Los nombres genéricos Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 y Euconibius Casey, 1895 [= Conibius LeConte, 1851] sonnuevos sinónimos (nombres válidos en corchetes). Se proponen las siguientes 127 nuevas sinonimias de nombres de nivel especie (ver lista en el «Resumen»), enumerados en su combinación original (nombres válidos, listados en su combinación actual, colocados entre corchetes). Las siguientes subespecies de Trogloderus costatus LeConte, 1879 reciben categoría de especie: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909 y Trogloderus vandykei La Rivers, 1946. Los siguientes táxones, que anteriormente se consideraban sinónimos más modernos, se consideran válidos: Amphidora Eschscholtz, 1829, Xysta Eschscholtz, 1829, Helops confluens (Casey, 1924). Se proponen dos nuevas combinaciones: Stenomorpha spinimana (Champion, 1892) y Stenomorpha tenebrosa (Champion, 1892) [del género Parasida Casey, 1912]. Las especies tipo [colocadas entre corchetes] de los siguientes 12 táxones del nivel génerosse designan por primera vez: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant y Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Dos especies previamente conocidas de América del Sur [Nilio lebasi J. Thomson y Platydema erotyloides Chevrolat] se mencionan por primera vez de América del Norte.

Résumé

Ce catalogue liste tous les noms de niveau famille (8 sous-familles, 52 tribus, 14 sous-tribus), genre (349 genres, 86 sous-genres), et espèce (2825 espèces, 215 sous-espèces) de Tenebrionidae (Coleoptera:) recensés en Amérique du Nord ainsi que leurs synonymes disponibles. Le catalogue inclut les taxons actuels, subfossiles et fossiles. L’auteur ainsi que l’année et la pagination de la description sont inclus pour chaque nom scientifique et, pour certains de ces noms, des renseignements additionnels (par exemple, l’espèce-type pour les noms de niveau genre, l’auteur de la synonymie des noms invalides des niveaux genre et espèce).

Plusieurs nouveaux actes nomenclaturaux sont proposés. On décrit un nouveau genre, Lepidocnemeplatia Bousquet et Bouchard. Spelaebiosis Bousquet et Bouchard [pour Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [pour Blapstinus kulzeri Marcuzzi, 1977] et Hymenorus campbelli Bouchard [pour Hymenorus oculatus Doyen and Poinar, 1994] sont des nouveaux noms de remplacement. On fournit les évidences nécessaires pour conserver l’usage actuel du nom Tarpela micans (Fabricius, 1798) nomen protectum sur celui de Tarpela vittata (Olivier, 1793) nomen oblitum. Les noms génériques Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 et Euconibius Casey, 1895 [= Conibius LeConte, 1851] sont des nouveaux synonymes (noms valides placés entre crochets). On propose 127 nouveaux synonymes de noms de niveau espèce (voir la liste dans « Abstract »), listés par leur combinaison originelle (noms valides, listés par leur combinaison actuelle, placés entre crochets). On élève au rang d’espèce les sous-espèces suivantes de Trogloderus costatus LeConte, 1879: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909, et Trogloderus vandykei La Rivers, 1946. On redonne le statut de taxons valides aux entités suivantes considérées à tort comme invalides: Amphidora Eschscholtz, 1829, Xysta Eschscholtz, 1829, et Helops confluens (Casey, 1924). On propose les combinaisons nouvelles suivantes: Stenomorpha spinimana (Champion, 1892) et Stenomorpha tenebrosa (Champion, 1892) [auparavant placés dans le genre Parasida Casey, 1912]. On désigne pour la première fois l’espèce type [placée entre crochets] des 12 noms suivants de niveau genre: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz, 1829]; Discogenia LeConte, 1866 [Eleodes scabricula LeConte, 1858]; Metablapylis Blaisdell, 1909 [Eleodes nigrina LeConte, 1858]; Steneleodes Blaisdell, 1909 [Eleodes longicollis LeConte, 1851]; Scaptes Champion, 1886 [Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866)]; Aspidius Mulsant and Rey, 1859 [Blaps punctata Fabricius, 1792]; Cryptozoon Schaufuss, 1882 [Cryptozoon civile Schaufuss, 1882]; Halophalerus Crotch, 1874 [Phaleria rotundata LeConte, 1851]; Dignamptus LeConte, 1878 [Dignamptus stenochinus LeConte, 1878]. Deux espèces, auparavant documentées seulement de l’Amérique du Sud [Nilio lebasi J. Thomson and Platydema erotyloides Chevrolat], sont rapportées pour la première fois d’Amérique du Nord.

Keywords

North America, nomenclature, Lepidocnemeplatia , new genus, new replacement names

Introduction

Darkling beetles (Coleoptera: Tenebrionidae) form a species-rich and morphologically diverse family with approximately 2300 genera and 20000 species worldwide (Matthews et al. 2010), and many more taxa to be described. The first thorough classification of the Tenebrionidae was provided by Lacordaire (1859) and was based entirely on the external morphology of adults. With relatively few exceptions, his classification schema was followed by subsequent workers for approximately 100 years (Watt 1967). The family classification was eventually reviewed using morphological characters of immature stages (Watt 1975) although significant changes did not appear until the first comprehensive investigation of adult internal structures including defense glands, female ovipositor, and female genital tube (Tschinkel and Doyen 1980, Doyen and Tschinkel 1982). The first higher-level phylogeny of the family based on molecular data was published only recently (Kergoat et al. 2014). As a result of these comparative and phylogenetic studies, several taxa previously treated as separate families (e.g., Lagriidae, Alleculidae, Nilionidae) are now included within Tenebrionidae. Additionally, many taxa previously included in Tenebrionidae are now classified in other families (see Table 1 for North American genera, Aalbu (2006) for worldwide taxa).

Table 1.

List of North American genera previously included in Tenebrionidae but currently classified in another family.

Genus Current placement
Boros Herbst, 1797 Boridae
Dacoderus LeConte, 1858 Salpingidae
Megazopherus Casey, 1907 Zopheridae
Meralius Casey, 1907 Zopheridae
Noserodes Casey, 1907 Zopheridae
Noserus LeConte, 1862 Zopheridae
Nosoderma Solier, 1841 Zopheridae
Phellopsis LeConte, 1862 Zopheridae
Phloeodes LeConte, 1862 Zopheridae
Sesaspis Casey, 1907 Zopheridae
Usechus Motschulsky, 1845 Zopheridae
Verodes Casey, 1907 Zopheridae
Zopherinus Casey, 1907 Zopheridae
Zopherodes Casey, 1907 Zopheridae
Zopherus Laporte, 1840 Zopheridae

The aim of this work is to synthesize available taxonomic, nomenclatural, and distributional information for all darkling beetles known from North America.

Methods

Nomenclatural data

All nomenclaturally available family-, genus- and species-group names are included. Extant taxa and subfossils from the Pleistocene (see Doyen and Miller 1980, Doyen and Poinar 1994) are given in the main catalogue. Impression fossils from major North American deposits are listed in Appendix 1, although the taxonomic assignment of these often-fragmentary fossils needs to be confirmed. Fossil species described from amber are listed in Appendix 2. Taxa incorrectly recorded from North America are given in Appendix 3. Subfamilies are listed in a phylogenetic framework but valid tribal, generic, and specific names are given in alphabetic order; listings of all invalid names are chronological.

The author and year and page number of the original description are provided for each scientific name. The type genus for each family-group name and the type species and type fixation for each genus-group name are included. The reference in which a given generic or specific name is first placed in synonymy with the current valid name is listed. Type-species designations in Lucas (1920) were accepted when a single species was listed under a particular genus-group name (see Alonso-Zarazaga and Lyal 2009, Bousquet et al. 2015). For genera with valid subgenera, synonyms are given under the nominotypical subgenus when relevant. Every species-group name are listed in its original combination, as given in the publication even if the agreement in gender with the generic name is incorrect. In cases where a species-group name older than the one currently recognized as valid is available but of doubtful application (e.g., Latridius pubescens Say, 1826), we have retained usage of the younger, accepted name as valid and treated the older name as nomen dubium. The synonymy list for adventive species focuses primarily on names used in the North American literature, other sources (e.g., Löbl et al. 2008a) can be consulted for data on all invalid names.

The classification used follows Bouchard et al. (2011) but also includes corrections and subsequent additions.

The gender of all valid genera listed in the catalogue has been determined following the provision of Article 30 (ICZN 1999) and indicated after the name using the initials M [Masculine], F [Feminine], and N [Neuter]. Therefore, the gender of the generic names Liodema, Platydema, and Scaphidema is herein treated as feminine following Article 30.1.3 (ICZN 1999; see Kerzhner (2003) and Löbl and Smetana (2010: 34) for further comments); the ending is derived from the Greek “demas (body silhouette).” The gender of Alaudes is treated as masculine following Article 30.2.4 (ICZN 1999). The gender of Eleodes, originally treated as feminine by Eschscholtz (1829), was changed to masculine by Somerby and Doyen (1976) and has been followed subsequently in the literature. However, Eleodes is feminine following Article 30.1.4.4 (ICZN 1999) which says that “a compound genus-group name ending in the suffix ... -odes is to be treated as masculine unless its author, when establishing the name, stated that it had another gender or treated it as such by combining it with an adjectival species-group name in another gender form.”

If necessary, the ending of all valid species-group names has been modified according to the gender of the generic name with which the species is currently combined. All specific names that are nouns in apposition need not agree in gender with the generic name and retain their original endings.

The author(s) of every new nomenclatural act proposed in the catalogue is given in square brackets (e.g., “[ADS]”) except for the first typification of genus-group names. One of these new acts needs further development. The genus-group taxon Lepidocnemeplatia was proposed by Kaszab (1938: 80) as a subgenus of Cnemeplatia Costa, 1847 to include two species, Cnemeplatia laticollis Champion, 1885 and C. sericea Horn, 1870. Unfortunately, Kaszab did not designate a type species for his new genus and therefore the name is unavailable from that date (ICZN 1999: Article 13.3). Löbl and Merkl (2003: 245) designated C. sericea Horn, 1870 as type species of Lepidocnemeplatia, the first typification for the taxon, and subsequently Löbl et al. (2008a: 140) credited authorship of the name to Löbl and Merkl (2003). However, since Löbl and Merkl (2003: 245) failed to indicate that they were establishing a new nominal taxon, a mandatory requirement for all new names published after 1999 (ICZN 1999: Article 16.1), the name cannot be attributed to them and is still a nomen nudum. In order to make the name available we here proposed the generic name Lepidocnemeplatia2 Bousquet and Bouchard, new genus; type species (here designated): Cnemeplatia sericea Horn, 1870. The reader is referred to Kaszab (1938: 79–80, couplet 1" of his key) for a description of the character states that differentiate the taxon (ICZN 1999: Article 13.1.2). A summary of all new nomenclatural acts is available in the Abstract.

Distributional data

This catalogue documents all species of Tenebrionidae from Greenland, Alaska, and Canada south to Panama, and also includes islands of the West Indies. Records from the Netherlands Antilles (consisting of several islands in the West Indies), the Venezuelan island Margarita, and Trinidad and Tobago off the northeast coast of Venezuela are not included since the fauna of these islands is more closely affiliated with the South American fauna. When known, the states (for Mexico and continental United States) and the provinces and territories (for Canada) are listed in parentheses for each species. For West Indian records, we give the political unit names for species found in the Lucayan Archipelago and Greater Antilles, occasionally with specific islands in parenthesis; political unit names are not provided for species occurring in the Lesser Antilles (LAN) and Virgin Islands (VIS), though sometimes we include specific islands in parentheses, especially when the species is known from only one or two islands. Further details regarding the West Indies geographical units can be found in Ivie and Hart (2016: Table 1).

Many Mexican state records in this catalogue came from localities listed by Champion in volume IV, parts 1 and 2, of the Biologia Centrali-Americana [1884–1893] and the gazetteer of Selander and Vaurie (1962) was used consistently to associate localities with states (including the federal district).

Amongst the geographical units used (see list below), HIS (for Hispaniola) is used when only the island record is known and LC (for Lower California) when only the peninsula record is known. South America (SA) is placed at the end of a species record list to indicate that the species extends into South America. Distributional records listed in square brackets (e.g., “[NM]”) are considered doubtful. South American species introduced accidentally into North America are indicated with a subscript “i” beside the country record (i.e., “USAi”).

Bibliographic data

References are provided for all scientific names included if a page number (related to the description of the taxon) is provided after the year of publication. Based on evidence previously published (see Bousquet 2016a: 211, 265), the dates of publication of Germar’s Coleopterorum species novae dated 1824 on its title page, and of Hope’s The Coleopterist’s manual, part the third, dated 1840 on its title page, are given as 1823 and 1841 respectively.

As discussed by Bousquet (2016b), Mäklin's “Monographie der Gattung Strongylium” was published in 1867, not 1864 as given by several authors.

List of acronyms used for geographic units3

BAH Bahamas

BEL Belize

BER Bermuda

CAN Canada [AB: Alberta; BC: British Columbia; MB: Manitoba; NB: New Brunswick; NF: Newfoundland and Labrador; NS: Nova Scotia; NT: Northwest Territories; NU: Nunavut; ON: Ontario; PE: Prince Edward Island; QC: Quebec; SK: Saskatchewan; YT: Yukon Territory]

CAY Cayman Islands

CRI Costa Rica

CUB Cuba

DOM Dominican Republic

GRE Greenland

GUA Guatemala

HAI Haiti

HIS Hispaniola

HON Honduras

JAM Jamaica

LAN Lesser Antilles (including among others Anguilla, Antigua and Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, Saint Lucia, Grenada, Barbados)

LC Lower California

MEX Mexico [AG: Aguascalientes; BC: Baja California; BS: Baja California Sur; CA: Campeche; CH: Chihuahua; CI: Chiapas; CL: Colima; CO: Coahuila; DU: Durango; FD: Federal District; GE: Guerrero; GU: Guanajuato; HI: Hidalgo; JA: Jalisco; ME: México; MI: Michoacán; MO: Morelos; NA: Nayarit; NL: Nuevo León; OA: Oaxaca; PU: Puebla; QR: Quintana Roo; QU: Querétaro; SI: Sinaloa; SL: San Luis Potosí; SO: Sonora; TA: Tamaulipas; TB: Tabasco; TL: Tlaxcala; VE: Veracruz; YU: Yucatán; ZA: Zacatecas]

NIC Nicaragua

PAN Panama

PRI Puerto Rico [includes Vieques]

SAL El Salvador

TUR Turks and Caicos Islands

USA United States of America [AK: Alaska; AL: Alabama; AR: Arkansas; AZ: Arizona; CA: California; CO: Colorado; CT: Connecticut; DC: District of Columbia; DE: Delaware; FL: Florida; GA: Georgia; IA: Iowa; ID: Idaho; IL: Illinois; IN: Indiana; KS: Kansas; KY: Kentucky; LA: Louisiana; MA: Massachusetts; MD: Maryland; ME: Maine; MI: Michigan; MN: Minnesota; MO: Missouri; MS: Mississippi; MT: Montana; NC: North Carolina; ND: North Dakota; NE: Nebraska; NH: New Hampshire; NJ: New Jersey; NM: New Mexico; NV: Nevada; NY: New York; OH: Ohio; OK: Oklahoma; OR: Oregon; PA: Pennsylvania; RI: Rhode Island; SC: South Carolina; SD: South Dakota; TN: Tennessee; TX: Texas; UT: Utah; VA: Virginia; VT: Vermont; WA: Washington; WI: Wisconsin; WV: West Virginia; WY: Wyoming]

VIS Virgin Islands [includes US Virgin Islands and British Virgin Islands: Saint Thomas, Saint Croix]

SA South America

Results

Overall diversity

A total of 128 valid and invalid family-group, 612 genus-group, and 4065 species-group taxa (excluding fossils listed in Appendices 1, 2) are listed in this catalogue. The subfamily Pimeliinae is the most diverse with 908 valid species-group taxa, followed by the Tenebrioninae (808), Alleculinae (418), Stenochiinae (361), Diaperinae (260), Lagriinae (256), Phrenapatinae (22), and Nilioninae (6). Thirty-seven species in three subfamilies are adventive (Table 2), several of which are pests of stored grain products. Mexico and the continental United States of America are by far the most diverse political regions with 1224 and 1230 valid species-group taxa respectively, while at the other extreme Greenland and Bermuda only have two each (Table 3).

Table 2.

List of adventive species documented in North America. Data on the origin, date of first detection in North America, and microhabitat associations in nature are given for each species as far as known. * = pest of stored grain products.

Species Origin Date of detection Microhabitat Placement
Alphitobius diaperinus (Panzer, 1797) Africa <1866 *Animal nests, caves, guano Tenebrioninae: Alphitobiini
Alphitobius laevigatus (Fabricius, 1781) Africa <1866 *Animal nests, caves, guano Tenebrioninae: Alphitobiini
Alphitophagus bifasciatus (Say, 1824) Europe, probably <1824 *Animal nests, caves, guano Diaperinae: Diaperini: Adelinina
Anchophthalmops menouxi (Mulsant and Rey, 1853) Africa; probably not established <1870 Leaf litter on sandy soil, probably Tenebrioninae: Pedinini: Platynotina
Blaps (Blaps) lethifera lethifera Marsham, 1802 Europe <1889 Under rocks, wood, in caves Tenebrioninae: Blaptini : Blaptina
Blaps (Blaps) mucronata Latreille, 1804 Europe <1889 Under rocks, wood, in caves Tenebrioninae: Blaptini : Blaptina
Ceropria induta (Wiedemann, 1819) Asia 1998 Polypore fungi, rotten wood Diaperinae: Diaperini: Diaperina
Ellipsodes (Anthrenopsis) ziczac (Motschulsky, 1873) Asia, probably via Antilles 1891 Under leaf litter on sandy soil Diaperinae: Crypticini
Gnatocerus (Echocerus) maxillosus (Fabricius, 1801) Asia, probably via Europe <1866 *Animal nests, caves, under bark Diaperinae: Diaperini: Adelinina
Gnatocerus (Gnatocerus) cornutus (Fabricius, 1798) Asia, probably via Europe <1866 *Animal nests, caves, under bark Diaperinae: Diaperini: Adelinina
Gondwanocrypticus pictus (Gebien, 1928) South America 1954 Under leaf litter near ant nests Diaperinae: Crypticini
Gondwanocrypticus platensis (Fairmaire, 1884) South America 1929 Under leaf litter near ant nests Diaperinae: Crypticini
Gonocephalum (Gonocephalum) sericeum (Baudi di Selve, 1875) Northwest Africa + Arabian Peninsula 1980 Under leaf litter, wood, rocks Tenebrioninae: Opatrini: Opatrina
Latheticus oryzae Waterhouse, 1880 Old World 1908 *Animal nests, caves, under bark Tenebrioninae: Triboliini
Leichenum canaliculatum variegatum (Klug, 1833) Madagascar 1906 Under leaf litter on sandy soil Tenebrioninae: Pedinini: Leichenina
Lyphia tetraphylla (Fairmaire, 1857) Europe <1902 In dead wood, other insect burrows Tenebrioninae: Triboliini
Myrmechixenus lathridioides Crotch, 1873 Europe <1883 Under leaf litter, in soil Diaperina: Myrmechixenini
Opatroides punctulatus Brulle, 1832 Middle East 2003 Under leaf litter, wood, rocks Tenebrioninae: Opatrini: Opatrina
Palorus cerylonoides (Pascoe, 1863) Indo-Malayan, probably 2004 Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus genalis Blair, 1930 Old World 1937 Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus ratzeburgii (Wissmann, 1848) North Africa, probably <1897 *Under bark dry wood, plant debris Tenebrioninae: Palorini
Palorus subdepressus (Wollaston, 1864) Africa, probably <1882 *Under bark dry wood, plant debris Tenebrioninae: Palorini
Pentaphyllus testaceus (Hellwig, 1792) Europe 2005 Polypore fungi, under bark Diaperinae: Diaperini: Diaperina
Platydema woldai Triplehorn and Phillips, 1998 Central America, probably 1964 With orchid plants; at lights in forest Diaperinae: Diaperini: Diaperina
Plesiophthalmus spectabilis Harold, 1875 Asia; probably not established 2013 Rotten wood Tenebrioninae: Amarygmini
Poecilocrypticus formicophilus Gebien, 1928 South America 1978 Under leaf litter near ant nests Diaperinae: Crypticini
Strongylium cultellatum Maklin, 1867 Asia 2010 Dead standing wood Stenochiinae: Stenochiini
Tenebrio molitor Linnaeus, 1758 Africa, probably <1837 *Animal nests, caves Tenebrioninae: Tenebrionini
Tenebrio obscurus Fabricius, 1792 Africa, probably <1869 *Animal nests, caves Tenebrioninae: Tenebrionini
Trachyscelis aphodioides Latreille, 1809 Europe <1846 Under plant debris on beach sand Diaperinae: Trachyscelini
Tribolium (Tribolium) castaneum (Herbst, 1797) Africa, probably <1866 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) confusum Jacquelin du Val, 1862 Africa, probably <1893 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) destructor Uyttenboogaart, 1934 Africa <1948 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tribolium (Tribolium) madens (Charpentier, 1825) Africa, probably <1866 *Under bark dry wood, plant debris Tenebrioninae: Triboliini
Tyrtaeus dobsoni Hinton, 1947 Unknown; probably via Europe 2002 Under bark and in dead wood Diaperinae: Gnathidiini : Anopidiina
Ulomina carinata Baudi di Selve, 1876 Asia 1952 Under bark dry wood, plant debris Tenebrioninae: Palorini
Ulomoides ocularis (Casey, 1891) Asia <1891 Dry pods of Tamarindus L. Diaperinae: Diaperini: Diaperina
Table 3.

Number of valid species-group taxa by political region. Data excludes impression and amber fossils.

Political region Lagriinae Nilioninae Phrenapatinae Pimeliinae Tenebrioninae Alleculinae Diaperinae Stenochiinae Total
BAH: Bahamas 0 0 0 8 22 14 25 1 70
BEL: Belize 20 0 3 2 20 5 32 28 110
BER: Bermuda 0 0 0 0 1 0 1 0 2
CAN: Canada 7 0 2 8 58 28 24 10 137
CAY: Cayman Islands 1 0 0 1 14 4 16 0 36
CRI: Costa Rica 44 0 6 12 27 23 40 44 196
CUB: Cuba 2 0 0 23 57 25 37 31 175
DOM: Dominican Republic 0 0 0 2 18 13 19 19 71
GRE: Greenland 0 0 0 0 2 0 0 0 2
GUA: Guatemala 60 2 4 20 70 57 66 71 350
HAI: Haiti 2 0 0 3 10 8 13 11 47
HON: Honduras 1 0 0 8 10 5 16 3 43
JAM: Jamaica 2 0 1 3 19 11 17 0 53
LAN: Lesser Antilles 11 0 3 6 50 11 31 18 130
MEX: Mexico 101 1 4 392 348 135 129 115 1225
NIC: Nicaragua 41 1 3 16 49 19 36 62 227
PAN: Panama 70 5 8 13 51 41 60 87 335
PRI: Puerto Rico 3 0 1 2 22 4 19 10 61
SAL: El Salvador 0 0 0 2 4 4 8 0 18
TUR: Turks and Caicos Islands 0 0 0 0 1 2 2 0 5
USA: United States of America 38 0 2 546 362 154 79 46 1227
VIS: Virgin Islands 0 0 0 0 3 2 1 1 7

A significant proportion of new species-group taxa (41%) were described between the years 1880 and 1910 (Fig. 1). A noticeable decrease in the number of new species-group taxa proposed occurred between 1940–1960, with a small resurgence since then (at a rate of approximately 100 new taxa per decade, see Figs 1, 2). Over 3000 North American species-group taxa are currently recognized as valid (Fig. 2), approximately 15% of the world fauna.

Figure 1. 

Number of North American species-group taxa described over time, by decade. Data excludes adventive species as well as impression and amber fossils.

Figure 2. 

Cumulative number of North American species-group taxa described over time, by decade. Data excludes adventive species as well as impression and amber fossils. Line with square marker = total available taxa, line with circular marker = currently valid taxa.

Significant contributions (see Table 4)

The British entomologist George Charles Champion [b. 1851, d. 1927], working on the fauna of Mexico and Central America, proposed the highest number of new tenebrionid taxa found on the continent (83 genus-group, 906 species-group taxa) followed by the American Thomas Lincoln Casey [b. 1857, d. 1925], working mainly on the fauna of the United States of America (80 genus-group, 792 species-group taxa). Frank Ellsworth Blaisdell [b. 1862, d. 1947] (338 species-group taxa), John Lawrence LeConte [b. 1825, d. 1883] (270 species-group taxa), George Henry Horn [b. 1840, d. 1897] (133 species-group taxa), and John Milton Campbell [b. 1935] (103 species-group taxa) also contributed significantly to describing the North American darkling beetle fauna.

Table 4.

Significant contributions to the description of new North American Tenebrionidae genus-group and species-group taxa (list includes the top ten contributors in each category). Data excludes adventive taxa as well as impression and amber fossils. * = given in parentheses is the country where the person produced taxonomic works, when different from the country of origin.

Author Country of origin* New genus-group names New species-group names
Blaisdell, Frank Ellsworth USA 20 338
Campbell, John Milton USA (Canada) 10 103
Casey, Thomas Lincoln USA 80 792
Champion, George Charles United Kingdom 83 906
Dejean, Pierre Francois Marie Auguste France 16 0
Doyen, John Thomas USA 5 79
Fall, Henry Clinton USA 1 66
Horn, George Henry USA 20 133
Laporte, Francois Louis (Comte de Castelnau) United Kingdom (France) 12 40
LeConte, John Lawrence USA 56 277
Maklin, Friedrich [Fredrik] Wilhelm Finland 3 66
Marcuzzi, Giorgio Italy 7 84
Motschulsky, Victor de Russia 12 24
Mulsant, Martial Etienne France 13 18
Pascoe, Francis Polkinghorne United Kingdom 12 4
Solier, Antoine Joseph Jean France 17 35
Triplehorn, Charles Albert USA 4 86

Catalogue of Tenebrionidae (Coleoptera) of North America

Family TENEBRIONIDAE Latreille, 1802

Tenebrionites Latreille, 1802: 165. Type genus: Tenebrio Linnaeus, 1758.

Subfamily LAGRIINAE Latreille, 1825

Lachnaedes Billberg, 1820a: 34. Type genus: Lachna Billberg, 1820 (= Lagria Fabricius, 1775). Nomen oblitum (see Favret and Bouchard 2016).

Lagriariae Latreille, 1825: 381. Type genus: Lagria Fabricius, 1775. Note. Use of younger family-group name conserved over Lachnina Billberg, 1820 (ICZN 1999: Article 40.2) (see Bouchard et al. 2005).

Tribe Belopini Reitter, 1917

Belopinae Reitter, 1917: 59. Type genus: Belopus Gebien, 1911.

Genus Adelonia Laporte, 1840 [F]

Adelonia Laporte, 1840: 221. Type species: Uloma filiformis Laporte, 1840, monotypy.

Merotemnus Horn, 1870: 367. Type species: Merotemnus elongatus Horn, 1870 (= Uloma filiformis Laporte, 1840), monotypy. Synonymy: Spilman (1961a: 49).

Rhacius Champion, 1885: 120. Type species: Rhacius sulcatulus Champion, 1885, subsequent designation (Gebien 1941: 805). Synonymy: Spilman (1961a: 50).

Adelonia filiformis (Laporte, 1840)MEX (BC BS)

Uloma filiformis Laporte, 1840: 221.

Merotemnus elongatus Horn, 1870: 367. Synonymy: Gebien (1908b: 160).

Adelonia insularis Doyen, 1983MEX (NA [Islas Marías])

Adelonia insularis Doyen, 1983: 85.

Adelonia quadricollis (Champion, 1885)GUABELPAN / SA

Rhacius quadricollis Champion, 1885: 121.

Adelonia sulcatula (Champion, 1885)4USA (TX) MEX (GE JA MO OA PU SO VE YU) GUAHONNICCRIPAN / CUBCAYJAM / SA

Rhacius sulcatulus Champion, 1885: 121.

Genus Rhypasma Pascoe, 1862 [N]

Rhypasma Pascoe, 1862: 325. Type species: Rhypasma pusillum Pascoe, 1862, monotypy.

Derosimus Fairmaire, 1904: 62. Type species: Derosimus quadricollis Fairmaire, 1904, monotypy. Synonymy: Blair (1935: 104).

Rhypasma costaricense Marcuzzi, 1976 CRI

Rhypasma costaricense Marcuzzi, 1976: 119.

Rhypasma haitianum Marcuzzi, 1954 CUB HAI

Rhypasma haitianum Marcuzzi, 1954a: 82.

Rhypasma livae Ferrer and Ødegaard, 2005 NIC PAN

Rhypasma livae Ferrer and Ødegaard, 2005: 635.

Tribe Eschatoporiini Blaisdell, 1906

Eschatoporini Blaisdell, 1906: 78. Type genus: Eschatoporis Blaisdell, 1906.

Genus Eschatoporis Blaisdell, 1906 [M]

Eschatoporis Blaisdell, 1906: 76. Type species: Eschatoporis nunenmacheri Blaisdell, 1906, monotypy.

Eschatoporis nunenmacheri Blaisdell, 1906USA (CA)

Eschatoporis nunenmacheri Blaisdell, 1906: 78.

Eschatoporis styx Aalbu, Kanda and Smith, 2017USA (CA)

Eschatoporis styx Aalbu, Kanda and Smith, 2017: 140.

Tribe Goniaderini Lacordaire, 1859

Goniadérides Lacordaire, 1859: 390. Type genus: Goniadera Perty, 1832.

Phobeliina Ardoin, 1961: 33. Type genus: Phobelius Blanchard, 1845.

Genus Anaedus Blanchard, 1842 [M]

Aspisoma Duponchel and Chevrolat, 1841: 210 [junior homonym of Aspisoma Laporte, 1833]. Type species: Aspisoma fulvipenne Duponchel and Chevrolat, 1841, original designation.

Anaedus Blanchard, 1842: pl. 14. Type species: Anaedus punctatissimus Blanchard, 1842, monotypy. Synonymy: Lacordaire (1859: 396).

Anaedes Agassiz, 1846: 36. Unjustified emendation of Anaedus Blanchard, 1842, not in prevailing usage.

Aspidosoma Agassiz, 1846: 36. Unjustified emendation of Aspisoma Duponchel and Chevrolat, 1841, not in prevailing usage.

Anaedus aeneotinctus Champion, 1893MEX (GE)

Anaedus aeneotinctus Champion, 1893a: 543.

Anaedus apicicornis Champion, 1886MEX (JA) PAN

Anaedus apicicornis Champion, 1886: 236.

Anaedus brevicollis Champion, 1886 GUA

Anaedus brevicollis Champion, 1886: 236.

Anaedus brunneus (Ziegler, 1844) [Fig. 3] CAN (ON) USA (AL AR DC FL IN KS KY LA MA MD MO MS NC NJ OH PA RI SC TN VA WI)

Figure 3. 

Anaedus brunneus (Ziegler, 1844). Scale bar = 1 mm.

Pandarus brunneus Ziegler, 1844: 45.

Anaedus impressicollis Pic, 1917 MEX

Anaedus impressicollis Pic, 1917: 19.

Anaedus inangulatus (Pic, 1934) NIC

Aspisoma inangulata Pic, 1934: 35.

Anaedus longicornis Champion, 1886USA (TX) MEX (GU OA) GUA

Anaedus longicornis Champion, 1886: 235.

Anaedus maculatus Champion, 1886 NIC PAN

Anaedus maculatus Champion, 1886: 235.

Anaedus marginatus Champion, 1886 NIC PAN

Anaedus marginatus Champion, 1886: 236.

Anaedus mexicanus Champion, 1886MEX (VE)

Anaedus mexicanus Champion, 1886: 234.

Anaedus nitidissimus Pic, 1917 CRI

Anaedus nitidissimus Pic, 1917: 20.

Anaedus pallidus Schaeffer, 1915USA (TX)

Anaedus pallidus Schaeffer, 1915: 238.

Anaedus punctatissimus Blanchard, 1842MEX (DU JA OA PU SI VE) GUANICCRIPAN/ SA

Anaedus punctatissimus Blanchard, 1842: pl. 14.

Anaedus quadrinotatus Champion, 1896 LAN

Anaedus quadrinotatus Champion, 1896: 26.

Anaedus rotundicollis LeConte, 1851USA (AZ) MEX (BS)

Anoedus rotundicollis LeConte, 1851: 150.

Anaedus setulosus Champion, 1886MEX (TB) NICPAN / SA

Anaedus setulosus Champion, 1886: 237.

Anaedus similis Champion, 1886MEX (VE) GUANIC

Anaedus similis Champion, 1886: 234.

Anaedus texanus Linell, 1899USA (TX)

Anoedus texanus Linell, 1899: 182.

Anaedus villosus Champion, 1893 GUA CRI PAN

Anaedus villosus Champion, 1893a: 543.

Genus Goniadera Perty, 1832 [F]

Goniadera Perty, 1832: 625. Type species: Goniadera crenata Perty, 1832, monotypy.

Goniodera Agassiz, 1846: 165. Unjustified emendation of Goniadera Perty, 1832, not in prevailing usage.

Subgenus Aemymone Bates, 1868

Aemymone Bates, 1868: 314. Type species: Aemymone cariosa Bates, 1868, original designation.

Goniadera championi Ferrer and Delatour, 2007MEX (VE) PAN

Aemymone crenata Champion, 1893a: 542 [junior secondary homonym of Goniadera crenata Perty, 1832].

Goniadera championi Ferrer and Delatour, 2007: 286. Replacement name for Goniadera crenata (Champion, 1893).

Subgenus Goniadera Perty, 1832

Goniadera Perty, 1832: 62. Type species: Goniadera crenata Perty, 1832, monotypy.

Goniadera alternata Champion, 1886MEX (VE) GUABELPAN

Goniadera alternata Champion, 1886: 231.

Goniadera dissipata Kirsch, 1866PAN / JAMLAN / SA

Goniadera dissipata Kirsch, 1866: 197.

Goniadera nicaraguensis Champion, 1886 NIC

Goniadera nicaraguensis Champion, 1886: 230.

Goniadera obscuriceps Pic, 1913NIC / SA

Goniadera obscuriceps Pic, 1913a: 125.

Goniadera oculata oculata Champion, 1886MEX (GE OA VE YU) BELNICCRIPAN

Goniadera oculata Champion, 1886: 230.

Goniadera pilosa Champion, 1886 NIC CRI PAN

Goniadera pilosa Champion, 1886: 230.

Goniadera pseudorepanda Ferrer and Delatour, 2007MEX (GE VE YU) GUANICCRI / SA

Goniadera pseudorepanda Ferrer and Delatour, 2007: 296.

Goniadera repanda (Fabricius, 1801)MEX (JA VE) GUABELNICCRI / SA

Melandrya repanda Fabricius, 1801a: 165.

Subgenus Opatresthes Gebien, 1928

Opatresthes Gebien, 1928b: 192. Type species: Opatresthes binodosus Gebien, 1928, subsequent designation (Gebien 1941: 817).

Goniadera maesi Ferrer and Delatour, 2007 NIC

Goniadera maesi Ferrer and Delatour, 2007: 287.

Goniadera quadrinodosa (Gebien, 1928)6CRI / SA

Opatresthes quadrinodosus Gebien, 1928b: 193.

Genus Paratenetus Spinola, 1844 [M]

Paratenetus Spinola, 1844: 116. Type species: Paratenetus punctatus Spinola, 1844, subsequent designation (Lucas 1920: 483).

Lagriola Kirsch, 1874: 409. Type species: Lagriola operosa Kirsch, 1874, present designation. Synonymy: Matthews and Lawrence (2015: 289).

Storthephora Mäklin, 1875a: 658. Type species: Storthephora denticollis Mäklin, 1875, subsequent designation (Bousquet and Bouchard 2014: 26). Synonymy: Champion (1893c: 47).

Paratenetus brevipennis Champion, 1886 PAN

Paratenetus brevipennis Champion, 1886: 242.

Paratenetus championi Matthews and Lawrence, 2015 PAN

Paratenetus denticulatus Champion, 1886: 243 [secondary homonym of Paratenetus denticulatus (Kirsch, 1874)].

Paratenetus championi Matthews and Lawrence, 2015: 311. Replacement name for Paratenetus denticulatus Champion, 1886.

Paratenetus constrictus Champion, 1893MEX (CI TB VE) GUABELCRIPAN

Paratenetus constrictus Champion, 1893a: 546.

Paratenetus corticarioides Champion, 1886MEX (OA) GUA

Paratenetus corticarioides Champion, 1886: 241.

Paratenetus crenulatus Champion, 1886 PAN

Paratenetus crenulatus Champion, 1886: 242.

Paratenetus exutus Bousquet and Bouchard, 2014CAN (AB MB NB NS ON QC SK) USA (AL AR CT DC FL IA IL IN KS KY LA MD ME MI MN MO MS NC ND NJ NY OH OK PA TN TX VA WI WV)

Paratenetus exutus Bousquet and Bouchard, 2014: 39.

Paratenetus foveithorax Ferrer and Ødegaard, 2005 PAN

Paratenetus foveithorax Ferrer and Ødegaard, 2005: 635.

Paratenetus fuscus LeConte, 1850 [Fig. 4] CAN (AB BC MB NT ON QC SK) USA (CO CT DC IA KS MA MD MI MT ND NE NM NY OH RI SD TN VT WI WY)

Figure 4. 

Paratenetus fuscus LeConte, 1850. Scale bar = 1 mm.

Paratenetus fuscus LeConte, 1850: 223.

Paratenetus crinitus Fall, 1907a: 253. Synonymy: Bousquet and Bouchard (2014: 31).

Paratenetus gibbipennis Motschulsky, 1869CAN (MB ON QC) USA (AL CT GA IL MA ME MI MN MO NC ND NE NH NJ NY OH PA RI SC TN TX VA WI)

Paratenetus gibbipennis Motschulsky, 1869: 193.

Paratenetus cribratus Motschulsky, 1869: 193. Synonymy: Bousquet and Bouchard (2014: 29).

Paratenetus grandicornis Motschulsky, 1869 NIC PAN

Paratenetus grandicornis Motschulsky, 1869: 193.

Paratenetus inermis Champion, 1893 GUA

Paratenetus inermis Champion, 1893a: 545.

Paratenetus koltzei Pic, 1939 MEX

Paratenetus koltzei Pic, 1939: 9.

Paratenetus longicornis Pic, 1925LAN (Guadeloupe)

Paratenetus longicornis Pic, 1925a: 6.

Paratenetus mexicanus Pic, 1925MEX (SI)

Paratenetus mexicanus Pic, 1925a: 6.

Paratenetus nigricornis Champion, 1893MEX (TB VE) GUABELPAN

Paratenetus nigricornis Champion, 1893a: 544.

Paratenetus obovatus Champion, 1886 BEL

Paratenetus obovatus Champion, 1886: 241.

Paratenetus punctatus Spinola, 1844CAN (MB NB ON QC) USA (AR CT DC GA IA IL IN KS KY LA MA MD ME MI MN MO MS NC NH NJ NY OH OK PA RI SC TN TX VA VT WI WV WY)

Latridius pubescens Say, 1826: 265 [nomen dubium, see Bousquet and Bouchard (2014: 33)].

Paratenetus punctatus Spinola, 1844: 118.

Paratenetus punctulatus Champion, 1893MEX (TB VE) BEL

Paratenetus punctulatus Champion, 1893a: 545.

Paratenetus ruficornis Champion, 1886 PAN

Paratenetus ruficornis Champion, 1886: 239.

Paratenetus sexdentatus Champion, 1893 GUA BEL PAN

Paratenetus sexdentatus Champion, 1893a: 546.

Paratenetus testaceus Pic, 1920MEX (PU) CRI

Paratenetus testaceus Pic, 1920: 2.

Paratenetus texanus Bousquet and Bouchard, 2014USA (FL LA TX) MEX (CI NA TA)

Paratenetus texanus Bousquet and Bouchard, 2014: 45.

Paratenetus tibialis Champion, 1886MEX (GE TB VE) GUABEL

Paratenetus tibialis Champion, 1886: 239.

Paratenetus tropicalis Motschulsky, 1869MEX (TB VE) GUABELPAN

Paratenetus tropicalis Motschulsky, 1869: 193.

Paratenetus tuberculatus Champion, 1886 PAN

Paratenetus tuberculatus Champion, 1886: 242.

Paratenetus villosus Champion, 1886MEX (VE) GUAPAN

Paratenetus villosus Champion, 1886: 240.

Genus Phobelius Blanchard, 1842 [M]

Phobelius Blanchard, 1842: pl. 14. Type species: Phobelius crenatus Blanchard, 1842, monotypy.

Phobelius mexicanus Doyen, 1990MEX (JA) GUA

Phobelius mexicanus Doyen, 1990: 217.

Phobelius nevermanni Kulzer, 1961 CRI

Phobelius nevermanni Kulzer, 1961a: 226.

Genus Phymatestes Pascoe, 1867 [M]

Phymatodes Dejean, 1834: 203. Type species: Lagria tuberculata Fabricius, 1792, monotypy. Note. Dejean’s name has been suppressed by the Commission in Opinion 1525 (ICZN 1989).

Phymatestes Pascoe, 1867: 142. Replacement name for Phymatodes Dejean, 1834 [as Phymatodes Blanchard, 1845]. Note. This name has been placed on the Official List of Generic Names in Zoology in Opinion 1525 (ICZN 1989).

Phymatestes agnei Ferrer and Ødegaard, 2005 PAN

Phymatestes agnei Ferrer and Ødegaard, 2005: 634.

Phymatestes charbonnelae Ferrer and Moraguès, 2003LAN (Grenada)

Phymatestes charbonnelae Ferrer and Moraguès, 2003: 161.

Genus Prateus LeConte, 1862 [M]

Prateus LeConte, 1862a: 238. Type species: Prateus fusculus LeConte, 1862, original designation.

Prateus fusculus LeConte, 1862USA (AL AR DC FL MD MS NC NY OH OK SC TN TX VA WV) MEX (TA)

Prateus fusculus LeConte, 1862a: 238.

Genus Xanthicles Champion, 1886 [M]

Xanthicles Champion, 1886: 231. Type species: Xanthicles caraboides Champion, 1886, subsequent designation (Gebien 1941: 815).

Xanthicles caraboides Champion, 1886 CRI

Xanthicles caraboides Champion, 1886: 232.

Xanthicles hirsutus Champion, 1886 CRI

Xanthicles hirsutus Champion, 1886: 232.

Tribe Lagriini Latreille, 1825

Lagriariae Latreille, 1825: 381. Type genus: Lagria Fabricius, 1775.

Subtribe Statirina Blanchard, 1845

Statyrites Blanchard, 1845: 39. Type genus: Statira Lepeletier and Audinet-Serville, 1828.

Genus Arthromacra Kirby, 1837 [F]

Arthromacra Kirby, 1837: 238. Type species: Arthromacra donacioides Kirby, 1837 (= Lagria aenea Say, 1824), monotypy.

Macrarthra Agassiz, 1846: 219. Unjustified emendation of Arthromacra Kirby, 1837, not in prevailing usage.

Arthromacra aenea aenea (Say, 1824)CAN (MB NB NS ON PE QC) USA (CT DC DE MA MD ME MI NC NH NJ NY OH PA TN VA VT WI WV)

Lagria aenea Say, 1824b: 287 [junior primary homonym of Lagria aenea Fabricius, 1775].

Arthromacra donacioides Kirby, 1837: 239. Synonymy: LeConte (1859d: 191).

Lagria viridis Melsheimer, 1845: 311. Synonymy: Parsons (1976: 222).

Arthromacra aenea glabricollis Blatchley, 1910USA (IL IN KY MO OH PA VA WI)

Arthromacra glabricollis Blatchley, 1910: 1285.

Arthromacra aenea lengi Parsons, 1976USA (GA NC PA SC TN WV)

Arthromacra aenea lengi Parsons, 1976: 224.

Arthromacra aenea rugosecollis Leng, 1914USA (GA NC TN)

Arthromacra aenea var. rugosecollis Leng, 1914: 287.

Arthromacra appalachiana Leng, 1917USA (NC SC TN VA)

Arthromacra appalachiana Leng, 1917: 18.

Arthromacra pilosella Leng, 1917USA (KY NC SC TN)

Arthromacra pilosella Leng, 1917: 18.

Arthromacra robinsoni Leng, 1914USA (NC SC VA)

Arthromacra robinsoni Leng, 1914: 286.

Genus Colparthrum Kirsch, 1866 [N]

Colparthrum Kirsch, 1866: 204. Type species: Colparthrum gerstaeckeri Kirsch, 1866, monotypy.

Subgenus Colparthrum Kirsch, 1866

Colparthrum Kirsch, 1866: 204. Type species: Colparthrum gerstaeckeri Kirsch, 1866, monotypy.

Colparthrum aenescens Borchmann, 1936 CRI

Colparthrum aenescens Borchmann, 1936: 444.

Colparthrum decoratum bilunulatum (Pic, 1912) PAN

Statira bilunulata Pic, 1912b: 76.

Colparthrum decoratum decoratum (Mäklin, 1863)MEX (VE) GUANICPAN

Statira decorata Mäklin, 1863: 588.

Colparthrum decoratum maklini Borchmann, 19367 [No region originally mentioned but presumably from Mexico and/or Central America]

Colparthrum decoratum var. mäklini Borchmann, 1936: 447.

Colparthrum foveiceps Champion, 1889 PAN

Colparthrum foveiceps Champion, 1889: 68.

Colparthrum grande Borchmann, 1936 CRI

Colparthrum grandis Borchmann, 1936: 439.

Colparthrum majus Borchmann, 1916 MEX

Colparthrum decoratum var. major Borchmann, 1916: 233.

Subgenus Pseudocolparthrum Borchmann, 1916

Pseudocolparthrum Borchmann, 1916: 236. Type species: Colparthrum calcaratum Champion, 1889, subsequent designation (Borchmann 1936: 452).

Colparthrum calcaratum Champion, 1889 NIC CRI PAN

Colparthrum calcaratum Champion, 1889: 71.

Colparthrum sulcicolle Champion, 1889 NIC PAN

Colparthrum sulcicolle Champion, 1889: 69.

Colparthrum vitticolle Champion, 1889 NIC

Colparthrum vitticolle Champion, 1889: 70.

Genus Disema Mäklin, 1875 [F]

Disema Mäklin, 1875a: 646. Type species: Disema bimaculata Mäklin, 1875, subsequent designation (Lucas 1920: 244).

Disema singularis (Champion, 1889) PAN

Sphragidophorus singularis Champion, 1889: 64.

Genus Epicydes Champion, 1889 [M]

Epicydes Champion, 1889: 60. Type species: Epicydes oculatus Champion, 1889, subsequent designation (Borchmann 1936: 429).

Subgenus Cybostira Borchmann, 1936

Cybostira Borchmann, 1936: 430. Type species: Cybostira caligata Borchmann, 1936, original designation.

Epicydes caligatus (Borchmann, 1936) CRI

Cybostira caligata Borchmann, 1936: 430.

Subgenus Epicydes Champion, 1889

Epicydes Champion, 1889: 60. Type species: Epicydes oculatus Champion, 1889, subsequent designation (Borchmann 1936: 429).

Epicydes oculatus Champion, 1889MEX (OA VE) GUA

Epicydes oculatus Champion, 1889: 61.

Epicydes vicinus Champion, 1889 GUA NIC

Epicydes vicinus Champion, 1889: 61.

Genus Meniscophorus Champion, 1889 [M]

Meniscophorus Champion, 1889: 64. Type species: Meniscophorus amazonicus Champion, 1889, subsequent designation (Lucas 1920: 404).

Meniscophorus costatus Champion, 1889 PAN

Meniscophorus costatus Champion, 1889: 65.

Genus Meropria Borchmann, 1921 [F]

Meropria Borchmann, 1921: 228. Type species: Statira glabrata Mäklin, 1863, original designation.

Meropria chiriquina (Champion, 1889) PAN

Statira chiriquina Champion, 1889: 9.

Meropria denticulata (Champion, 1889)PAN / SA

Statira denticulata Champion, 1889: 7.

Meropria glabrata (Mäklin, 1863)MEX (GE MO OA VE) GUABELCRI

Statira glabrata Mäklin, 1863: 587.

Meropria interrupta (Champion, 1889) GUA NIC PAN

Statira interrupta Champion, 1889: 8.

Meropria unidentata (Champion, 1889)MEX (CI VE) GUABEL

Statira unidentata Champion, 1889: 8.

Genus Nevermanniella Borchmann, 1936 [F]

Nevermanniella Borchmann, 1936: 332. Type species: Statira albolineata Champion, 1889, original designation.

Nevermanniella albolineata (Champion, 1889)MEX (VE) BELNIC

Statira albolineata Champion, 1889: 36.

Genus Othryades Champion, 1889 [M]

Othryades Champion, 1889: 72. Type species: Othryades fragilicornis Champion, 1889, monotypy.

Othryades fragilicornis Champion, 1889 PAN

Othryades fragilicornis Champion, 1889: 72.

Genus Rhaibodera Borchmann, 1921 [F]

Rhaibodera Borchmann, 1921: 219. Type species: Rhaibodera pachycera Borchmann, 1921, original designation.

Rhaibodera crassicornis (Champion, 1889)MEX (TB)

Statira crassicornis Champion, 1889: 18.

Genus Rhosaces Champion, 1889 [M]

Rhosaces Champion, 1889: 73. Type species: Rhosaces clavipes Champion, 1889, monotypy.

Rhosaces clavipes Champion, 1889 PAN

Rhosaces clavipes Champion, 1889: 73.

Genus Sphragidophorus Champion, 1889 [M]

Sphragidophorus Champion, 1889: 61. Type species: Statira cyanipennis Mäklin, 1863, subsequent designation (Lucas 1920: 603).

Sphragidophorus cyanipennis (Mäklin, 1863)MEX (VE) GUAPAN

Statira cyanipennis Mäklin, 1863: 591.

Sphragidophorus ocularis Borchmann, 1936 CRI

Sphragidophorus ocularis Borchmann, 1936: 507.

Sphragidophorus violaceus Champion, 1889 PAN

Sphragidophorus violaceus Champion, 1889: 63.

Genus Statira Lepeletier and Audinet-Serville, 1828 [F]

Statira Lepeletier and Audinet-Serville, 1828: 479. Type species: Statira agroides Lepeletier and Audinet-Serville, 1828, subsequent designation (Blanchard 1844: pl. 53bis).

Subgenus Spinostatira Pic, 1918

Spinostatira Pic, 1918a: 22. Type species: Statira spinipes Pic, 1918, subsequent designation (Borchmann 1936: 247).

Statira costaricensis Champion, 1889CRI / SA

Statira costaricensis Champion, 1889: 36.

Subgenus Statira Lepeletier and Audinet-Serville, 1828

Statira Lepeletier and Audinet-Serville, 1828: 479. Type species: Statira agroides Lepeletier and Audinet-Serville, 1828, subsequent designation (Blanchard 1844: pl. 53bis).

Statira aeneipennis Champion, 1889 GUA

Statira aeneipennis Champion, 1889: 25.

Statira aeneotincta Champion, 1889MEX (VE) GUA

Statira aeneotincta Champion, 1889: 27.

Statira aerata Champion, 1889MEX (VE) GUA

Statira aerata Champion, 1889: 26.

Statira agraeformis Champion, 1889 PAN

Statira agraeformis Champion, 1889: 12.

Statira albofasciata Champion, 1889MEX (VE) GUAPAN

Statira albofasciata Champion, 1889: 44.

Statira alternans Champion, 1889MEX (OA)

Statira alternans Champion, 1889: 30.

Statira amicta Borchmann, 1936 CRI

Statira amicta Borchmann, 1936: 294.

Statira analis Borchmann, 1921 MEX

Statira analis Borchmann, 1921: 273.

Statira angustula Champion, 1889 GUA

Statira angustula Champion, 1889: 15.

Statira antennalis Borchmann, 1936 CRI

Statira antennalis Borchmann, 1936: 313.

Statira asperata Champion, 1889PAN / LAN / SA

Statira asperata Champion, 1889: 49.

Statira antillarum Champion, 1896: 36. Synonymy: Champion (1917: 230).

Statira basalis Horn, 1888USA (AL AR FL GA LA MD MO MS NC SC TX VA)

Statira basalis Horn, 1888: 31.

Statira bicolor Champion, 1889 PAN

Statira bicolor Champion, 1889: 47.

Statira biseriata Borchmann, 1921 MEX

Statira biseriata Borchmann, 1921: 263.

Statira borchmanni Nevermann, 1926 CRI

Statira borchmanni Nevermann, 1926: 113.

Statira brevipilis Champion, 1889MEX (VE)

Statira brevipilis Champion, 1889: 39.

Statira caeruleipennis Champion, 1889MEX (MI)

Statira caeruleipennis Champion, 1889: 14.

Statira ciliata Champion, 1889 GUA

Statira ciliata Champion, 1889: 42.

Statira collaris Champion, 1889MEX (VE)

Statira collaris Champion, 1889: 13.

Statira colorata Fall, 1909MEX (BC BS)

Statira colorata Fall, 1909: 165.

Statira conspicillata conspicillata Mäklin, 1863MEX (CI TB VE) GUABELNICPAN

Statira conspicillata Mäklin, 1863: 589.

Statira conspicillata lateannulata Borchmann, 1936 GUA

Statira conspicillata var. lateannulata Borchmann, 1936: 291.

Statira corrosa Champion, 1889MEXGUA / SA

Statira corrosa Champion, 1889: 37.

Statira cribrata Champion, 1889 GUA

Statira cribrata Champion, 1889: 42.

Statira croceicollis Mäklin, 1863USA (AL FL GA MS)

Statira croceicollis Mäklin, 1863: 594.

Statira cruciata Champion, 1917 NIC

Statira cruciata Champion, 1917: 254.

Statira cupreotincta Champion, 1889 NIC PAN

Statira cupreotincta Champion, 1889: 23.

Statira curticollis Champion, 1889MEX (GE ME MO OA) GUA

Statira curticollis Champion, 1889: 24.

Statira defecta Schaeffer, 1905USA (AZ NM)

Statira defecta Schaeffer, 1905b: 175.

Statira dolera Parsons, 1966USA (FL GA)

Statira dolera Parsons, 1966: 249.

Statira dumalis Parsons, 1973USA (CA)

Statira dumalis Parsons, 1973: 1.

Statira erina Parsons, 1976USA (TX)

Statira erina Parsons, 1976: 219.

Statira evanescens evanescens Champion, 1889MEX (DU GE JA MO OA VE) NIC

Statira evanescens Champion, 1889: 34.

Statira evanescens obscuripennis Borchmann, 1921 MEX

Statira evanescens var. obscuripennis Borchmann, 1921: 260.

Statira flohri Champion, 1893MEX (OA)

Statira flohri Champion, 1893b: 452.

Statira foveicollis Champion, 1889 BEL NIC PAN

Statira foveicollis Champion, 1889: 18.

Statira fulva Fleutiaux and Sallé, 1890 LAN

Statira fulva Fleutiaux and Sallé, 1890: 431.

Statira gagatina Melsheimer, 1845 [Fig. 5] CAN (QC) USA (AL AR CT DC DE IA IL IN KS KY MA MD MI MS NC NJ NY OH PA SC TN VT WI)

Figure 5. 

Statira gagatina Melsheimer, 1845. Scale bar = 1 mm.

Statyra gagatina Melsheimer, 1845: 311.

Statyra resplendens Melsheimer, 1845: 311. Synonymy: Parsons (1976: 220).

Lagria fusca Melsheimer, 1845: 311. Synonymy: Parsons (1966: 252).

Statira glabricollis Borchmann, 1936 CRI

Statira glabricollis Borchmann, 1936: 315.

Statira guatemalensis Champion, 1889 GUA

Statira guatemalensis Champion, 1889: 48.

Statira guttata Borchmann, 1921MEX (“Tenancingo”)

Statira guttata Borchmann, 1921: 250.

Statira haitiensis Champion, 1917 HAI

Statira haitiensis Champion, 1917: 255.

Statira heliconiae Borchmann, 1936 CRI

Statira heliconiae Borchmann, 1936: 292.

Statira heliophila Borchmann, 1936 CRI

Statira heliophila Borchmann, 1936: 276.

Statira hirsuta Champion, 1889USA (TX) MEX (CI GE JA VE) NIC

Statira hirsuta Champion, 1889: 50.

Statira simulans Schaeffer, 1905a: 180. Synonymy: Parsons (1966: 246).

Statira horrida Champion, 1889 GUA

Statira horrida Champion, 1889: 38.

Statira huachucae Schaeffer, 1905USA (AZ NM)

Statira huachucae Schaeffer, 1905b: 176.

Statira ignita Champion, 1889MEX (VE)

Statira ignita Champion, 1889: 23.

Statira inaequicollis Borchmann, 1936 CRI

Statira inaequicollis Borchmann, 1936: 287.

Statira inconstans Champion, 1889 GUA NIC

Statira inconstans Champion, 1889: 16.

Statira ingae Borchmann, 1936 CRI

Statira ingae Borchmann, 1936: 295.

Statira ingens Champion, 1889 NIC CRI PAN

Statira ingens Champion, 1889: 12.

Statira irazuensis Champion, 1889 CRI

Statira irazuensis Champion, 1889: 22.

Statira irregularis Champion, 1889 GUA

Statira irregularis Champion, 1889: 45.

Statira isthmiaca Champion, 1889 PAN

Statira isthmiaca Champion, 1889: 19.

Statira laevicollis Champion, 1889MEX (CL GE)

Statira laevicollis Champion, 1889: 46.

Statira latitator Parsons, 1973USA (CA) MEX (BC)

Statira latitator Parsons, 1973: 3.

Statira leptotracheloides Champion, 1889MEX (DU)

Statira leptotracheloides Champion, 1889: 52.

Statira liebecki Leng, 1924USA (AL FL SC)

Statira liebecki Leng, 1924: 187.

Statira limbata Champion, 1889MEX (TB VE YU) BEL

Statira limbata Champion, 1889: 14.

Statira limonis Borchmann, 1936 CRI

Statira limonis Borchmann, 1936: 315.

Statira marmorata Champion, 1889MEX (VE) GUA

Statira marmorata Champion, 1889: 43.

Statira mediosignata Borchmann, 1921MEX (“Santiago Ixcuintla”)

Statira mediosignata Borchmann, 1921: 278.

Statira melanocephala Mäklin, 1863MEX (VE)

Statira melanocephala Mäklin, 1863: 593.

Statira metallica Champion, 1889 NIC CRI PAN

Statira metallica Champion, 1889: 16.

Statira mexicana Champion, 1889MEX (PU VE)

Statira mexicana Champion, 1889: 26.

Statira microps Champion, 1889MEX (CI TB) GUA

Statira microps Champion, 1889: 44.

Statira minima minima Champion, 1889 NIC PAN

Statira minima Champion, 1889: 47.

Statira minima subatra Borchmann, 1936 “Mittelamerika”

Statira minima var. subatra Borchmann, 1936: 276.

Statira multiformis Champion, 1889MEX (VE) GUANICPAN

Statira multiformis Champion, 1889: 19.

Statira multipunctata Champion, 1889MEX (MO)

Statira multipunctata Champion, 1889: 49.

Statira nevermanni Borchmann, 1936 CRI

Statira nevermanni Borchmann, 1936: 255.

Statira nigripennis affinis Mäklin, 1875 MEX

Statira affinis Mäklin, 1875a: 642.

Statira nigripennis championi Pic, 1912 MEX

Statira nigripennis var. championi Pic, 1912a: 20.

Statira nigripennis humeralis Mäklin, 1875 MEX

Statira humeralis Mäklin, 1875a: 642.

Statira nigripennis nigripennis Mäklin, 1875MEX (JA MO)

Statira nigripennis Mäklin, 1875a: 641.

Statira nigroaenea Champion, 1889MEX (DU)

Statira nigroaenea Champion, 1889: 40.

Statira nigrofasciata Borchmann, 1921MEX (“Navarrete”)

Statira nigrofasciata Borchmann, 1921: 257.

Statira nigromaculata Champion, 1889USA (AZ TX) MEX (OA VE) GUA

Statira nigromaculata Champion, 1889: 33.

Statira nigrosparsa Mäklin, 1863MEX (GE VE) GUANIC

Statira nigrosparsa Mäklin, 1863: 590.

Statira nodulosa Champion, 1889 GUA

Statira nodulosa Champion, 1889: 31.

Statira opaca Borchmann, 1936 CRI

Statira opaca Borchmann, 1936: 283.

Statira opacicollis Horn, 1888USA (AZ)

Statira opacicollis Horn, 1888: 30.

Statira paradoxa Borchmann, 1936 CRI

Statira paradoxa Borchmann, 1936: 306.

Statira patricia Borchmann, 1921 CRI

Statira patricia Borchmann, 1921: 269.

Statira penicillata Champion, 1889MEX (VE)

Statira penicillata Champion, 1889: 30.

Statira perforata Champion, 1917 MEX

Statira perforata Champion, 1917: 262.

Statira pici Blackwelder, 1945 CRI

Statira bimaculata Borchmann, 1936: 291 [junior primary homonym of Statira bimaculata Pic, 1912].

Statira pici Blackwelder, 1945: 500. Replacement name for Statira bimaculata Borchmann, 1936.

Statira picta Champion, 1889 NIC PAN

Statira picta Champion, 1889: 36.

Statira pilifera Champion, 1893MEX (VE)

Statira pilifera Champion, 1893b: 451.

Statira pilipes Champion, 1889MEX (CI)

Statira pilipes Champion, 1889: 43.

Statira pluripunctata Horn, 1888USA (AZ NM TX UT) MEX (GE)

Statira pluripunctata Horn, 1888: 29.

Statira sulcicrus Champion, 1889: 51. Synonymy: Parsons (1966: 245).

Statira pueblensis Champion, 1889MEX (PU)

Statira pueblensis Champion, 1889: 51.

Statira pulchella Mäklin, 1863USA (TX) MEX (SL TA VE) NIC

Statira pulchella Mäklin, 1863: 589.

Statira punctatissima Champion, 1889MEX (CI)

Statira punctatissima Champion, 1889: 38.

Statira punctipennis Champion, 1889 GUA

Statira punctipennis Champion, 1889: 28.

Statira reticulaticollis Borchmann, 1936 NIC CRI

Statira reticulaticollis Borchmann, 1936: 269.

Statira robusta Schaeffer, 1905USA (AZ CO NM TX)

Statira robusta Schaeffer, 1905a: 180.

Statira rugicollis Champion, 1889MEX (VE)

Statira rugicollis Champion, 1889: 48.

Statira rugipes Champion, 1889MEX (DU)

Statira rugipes Champion, 1889: 52.

Statira schmidti Borchmann, 1936 CRI

Statira schmidti Borchmann, 1936: 304.

Statira scitula Champion, 1889MEX (VE) GUA

Statira scitula Champion, 1889: 10.

Statira setigera Champion, 1889MEX (GU)

Statira setigera Champion, 1889: 41.

Statira simplex Borchmann, 1936 CRI

Statira simplex Borchmann, 1936: 284.

Statira sobrina Champion, 1889 MEX

Statira sobrina Champion, 1889: 22.

Statira spiculifera Champion, 1893MEX (VE)

Statira spiculifera Champion, 1893b: 451.

Statira suavis Champion, 1889MEX (DU)

Statira suavis Champion, 1889: 15.

Statira subnitida LeConte, 1866MEX (BC BS)

Statira subnitida LeConte, 1866b: 141.

Statira testacea Champion, 1889MEX (VE)

Statira testacea Champion, 1889: 28.

Statira tolensis Champion, 1889 PAN

Statira tolensis Champion, 1889: 20.

Statira triangulifer Champion, 1889MEX (CI VE) GUABEL

Statira triangulifer Champion, 1889: 34.

Statira tristis Mäklin, 1875 MEX

Statira tristis Mäklin, 1875a: 639.

Statira tropicalis Champion, 1889MEX (VE) GUABELNIC

Statira tropicalis Champion, 1889: 10.

Statira tuberculifera Champion, 1889 GUA

Statira tuberculifera Champion, 1889: 32.

Statira tuberosa Champion, 1889MEX (VE)

Statira tuberosa Champion, 1889: 31.

Statira variabilis inexpecta Borchmann, 1936 PAN

Statira variabilis var. inexpecta Borchmann, 1936: 282.

Statira variabilis variabilis Champion, 1889 GUA PAN

Statira variabilis Champion, 1889: 11.

Statira veraecrucis Champion, 1889MEX (VE)

Statira veraecrucis Champion, 1889: 35.

Statira veraepacis Champion, 1889 GUA

Statira veraepacis Champion, 1889: 24.

Statira vilis Mäklin, 1863MEX (CI TB VE) GUABELPAN

Statira vilis Mäklin, 1863: 592.

Statira villosa Champion, 1889MEX (GE)

Statira villosa Champion, 1889: 39.

Statira viridicollis Champion, 1889 PAN

Statira viridicollis Champion, 1889: 17.

Statira vittata Champion, 1896 LAN

Statira vittata Champion, 1896: 37.

Genus Uroplatopsis Champion, 1889 [F]

Uroplatopsis Champion, 1889: 53. Type species: Uroplatopsis imitator Champion, 1889, subsequent designation (Lucas 1920: 666).

Uroplatopsis appendiculata Champion, 1889 PAN

Uroplatopsis appendiculata Champion, 1889: 59.

Uroplatopsis dilaticornis Champion, 1889 PAN

Uroplatopsis dilaticornis Champion, 1889: 58.

Uroplatopsis excavata Champion, 1889 PAN

Uroplatopsis excavata Champion, 1889: 59.

Uroplatopsis imitator Champion, 1889 NIC

Uroplatopsis imitator Champion, 1889: 54.

Uroplatopsis mimica Champion, 1889 PAN

Uroplatopsis mimica Champion, 1889: 57.

Uroplatopsis planicollis Champion, 1889 PAN

Uroplatopsis planicollis Champion, 1889: 56.

Uroplatopsis reducta Pic, 1931 CRI

Uroplatopsis reducta Pic, 1931: 33.

Uroplatopsis reticulata Champion, 1889 PAN

Uroplatopsis reticulata Champion, 1889: 56.

Uroplatopsis simulans nevermanni Borchmann, 1936 CRI

Uroplatopsis simulans var. nevermanni Borchmann, 1936: 482.

Uroplatopsis simulans simulans Champion, 1889 PAN

Uroplatopsis simulans Champion, 1889: 58.

Uroplatopsis vermiculata Champion, 1889 NIC

Uroplatopsis vermiculata Champion, 1889: 55.

Tribe Lupropini Lesne, 1926

Lypropsini Lesne, 1926: 68. Type genus: Luprops Hope, 1833 [as Lyprops, incorrect subsequent spelling of the type genus name, not in prevailing usage].

Genus Lorelus Sharp, 1876 [M]

Lorelus Sharp, 1876: 76. Type species: Lorelus priscus Sharp, 1876, monotypy.

Lorelopsis Champion, 1896: 15. Type species: Lorelopsis pilosus Champion, 1896, monotypy. Synonymy: Doyen (1993: 295).

Lorelus angustulus Champion, 1913 GUA

Lorelus angustulus Champion, 1913: 165.

Lorelus bicolor Doyen, 1993 PRI

Lorelus bicolor Doyen, 1993: 296.

Lorelus brevicornis Champion, 1896 LAN

Lorelus brevicornis Champion, 1896: 14.

Lorelus breviusculus Champion, 1913 PAN

Lorelus breviusculus Champion, 1913: 165.

Lorelus cribricollis Kaszab, 1940LAN (Guadeloupe)

Lorelus cribricollis Kaszab, 1940: 156.

Lorelus curticollis Champion, 1913MEX (VE) GUAPAN

Lorelus curticollis Champion, 1913: 164.

Lorelus curvipes Champion, 1913 GUA

Lorelus curvipes Champion, 1913: 163.

Lorelus exilis Champion, 1913 GUA

Lorelus exilis Champion, 1913: 166.

Lorelus glabratus Doyen, 1993 PRI

Lorelus glabratus Doyen, 1993: 297.

Lorelus guadeloupensis Kaszab, 1940LAN (Guadeloupe)

Lorelus guadeloupensis Kaszab, 1940: 155.

Lorelus pilosus (Champion, 1896) LAN

Lorelopsis pilosus Champion, 1896: 16.

Lorelus trapeziderus Champion, 1913 GUA PAN

Lorelus trapeziderus Champion, 1913: 167.

Lorelus wolcotti Doyen, 1993 PRI

Lorelus wolcotti Doyen, 1993: 295.

Incertae sedis: Lagriinae

Genus Pseudesarcus Champion, 19138 [M]

Pseudesarcus Champion, 1913: 115. Type species: Pseudesarcus villosus Champion, 1913, original designation.

Pseudesarcus villosus Champion, 1913 PAN

Pseudesarcus villosus Champion, 1913: 116.

Subfamily NILIONINAE Oken, 1843

Nilioniden Oken, 1843: 484. Type genus: Nilio Latreille, 1802.

Genus Nilio Latreille, 1802 [M]

Nilio Latreille, 1802: 179 [as Nilion]9. Type species: Coccinella villosa Fabricius, 1787, monotypy.

Subgenus Nilio Latreille, 1802

Nilio Latreille, 1802: 179 [as Nilion]. Type species: Coccinella villosa Fabricius, 1787, monotypy.

Nilio chiriquensis Champion, 1888 PAN

Nilio chiriquensis Champion, 1888: 471.

Nilio fulvopilosus Champion, 1888 PAN

Nilio fulvo-pilosus Champion, 1888: 471.

Nilio lebasi J. Thomson, 18609PAN / SANew North American record

Nilio lebasei J. Thomson, 1860: 10.

Nilio sallei J. Thomson, 1860MEX (VE) GUA

Nilio sallei J. Thomson, 1860: 10.

Nilio sallaei Champion, 1888: 470. Unjustified emendation of Nilio sallei J. Thomson, 1860, not in prevailing usage.

Nilio thomsoni Champion, 1888 GUA NIC PAN

Nilio thomsoni Champion, 1888: 471.

Nilio villosus (Fabricius, 1787)PAN / SA

Coccinella villosa Fabricius, 1787: 379 [junior primary homonym of Coccinella villosa Geoffroy, 1785].

Subfamily PHRENAPATINAE Solier, 1834

Phrépatides Solier, 1834: 488. Type genus: Phrenapates Gray, 1832.

Tribe Archaeoglenini Watt, 1975

Archaeoglenini Watt, 1975: 412. Type genus: Archaeoglenes Broun, 1893.

Genus Archaeoglenes Broun, 1893 [M]

Archaeoglenes Broun, 1893: 188. Type species: Archaeoglenes costipennis Broun, 1893, monotypy.

Archaeoglenes bollensis Watrous, 1982 PAN

Archaeoglenes bollensis Watrous, 1982: 140.

Archaeoglenes occidentalis Lawrence, 1979MEX (CI) BELPAN / SA

Archaeoglenes occidentalis Lawrence [in Doyen and Lawrence], 1979: 358.

Archaeoglenes pecki Lawrence, 1979 JAM

Archaeoglenes pecki Lawrence [in Doyen and Lawrence], 1979: 358.

Archaeoglenes puntaensis Watrous, 1982 PAN

Archaeoglenes puntaensis Watrous, 1982: 141.

Tribe Penetini Lacordaire, 1859

Pénétides Lacordaire, 1859: 318. Type genus: Peneta Lacordaire, 1859.

Phthorini Boddy, 1965: 144. Type genus: Phtora Mulsant, 1854 [as Phthora, incorrect subsequent spelling, not in prevailing usage].

Genus Clamoris des Gozis, 1886 [F]

Phtora Mulsant, 1854: 228 [junior homonym of Phtora Germar, 1835]. Type species: Phtora crenata Mulsant, 1854, monotypy.

Clamoris des Gozis, 1886: 25. Replacement name for Phtora Mulsant, 1854.

Phthora Champion, 1893a: 531. Unjustified emendation of Phtora Mulsant, 1854, not in prevailing usage.

Clamoris americana (Horn, 1874) [Fig. 6] CAN (BC) USA (CA OR WA)

Figure 6. 

Clamoris americana (Horn, 1874). Scale bar = 1 mm.

Phthora americana Horn, 1874a: 35.

Clamoris armata (Champion, 1893) GUA

Phthora armata Champion, 1893a: 532.

Clamoris elongata (Champion, 1893)MEX (VE) NIC

Phthora elongata Champion, 1893a: 532.

Genus Cleolaus Champion, 1886 [M]

Cleolaus Champion, 1886: 142. Type species: Peneta sommeri Lacordaire, 1859, original designation.

Cleolaus sommeri (Lacordaire, 1859)MEX (OA)

Peneta sommeri Lacordaire, 1859: 320.

Genus Daochus Champion, 1886 [M]

Daochus Champion, 1886: 139. Type species: Daochus mandibularis Champion, 1886, monotypy.

Daochus mandibularis Champion, 1886 GUA BEL

Daochus mandibularis Champion, 1886: 140.

Genus Dioedus LeConte, 1862 [M]

Dioedus LeConte, 1862a: 238. Type species: Dioedus punctatus LeConte, 1862, monotypy.

Arrhabaeus Champion, 1886: 144. Type species: Arrhabaeus convexus Champion, 1886, monotypy. Synonymy: Kaszab (1977b: 314).

Dioedus convexus (Champion, 1886) CRI PAN

Arrhabaeus convexus Champion, 1886: 145.

Dioedus debilis (Champion, 1896) LAN

Arrhabaeus debilis Champion, 1896: 20.

Dioedus guadeloupensis (Fleutiaux and Sallé, 1890) LAN

Arrhabaeus guadeloupensis Fleutiaux and Sallé, 1890: 424.

Dioedus minor (Fleutiaux and Sallé, 1890) LAN

Arrhabaeus guadeloupensis minor Fleutiaux and Sallé, 1890: 425.

Dioedus punctatus LeConte, 1862 [Fig. 7] CAN (ON) USA (AL CT DC FL IA IL IN KS MD MI MO NC NJ NY OH SC VA WI WV) / PRI

Figure 7. 

Dioedus punctatus LeConte, 1862. Scale bar = 1 mm.

Dioedus punctatus LeConte, 1862a: 238.

Genus Peneta Lacordaire, 1859 [F]

Peneta Lacordaire, 1859: 319. Type species: Peneta lebasii Lacordaire, 1859, subsequent designation (Lucas 1920: 492).

Peneta costaricensis Gebien, 1928 CRI

Peneta costaricensis Gebien, 1928a: 147.

Peneta nevermanni Gebien, 1928 CRI

Peneta nevermanni Gebien, 1928a: 148.

Peneta nuchicornis Gebien, 1928 CRI PAN

Peneta nuchicornis Gebien, 1928a: 146.

Peneta obtusicornis Kirsch, 1866PAN / SA

Peneta obtusicornis Kirsch, 1866: 191.

Peneta panamensis Champion, 1886: 142. Synonymy: Champion (1893a: 531).

Genus Telchis Champion, 1886 [M]

Telchis Champion, 1886: 142. Type species: Telchis clavicornis Champion, 1886, monotypy.

Telchis clavicornis Champion, 1886 PAN

Telchis clavicornis Champion, 1886: 143.

Genus Zypoetes Champion, 1893 [M]

Zypoetes Champion, 1893a: 532. Type species: Zypoetes epieroides Champion, 1893, monotypy.

Zypoetes epieroides Champion, 1893MEX (VE) GUABELNIC

Zypoetes epieroides Champion, 1893a: 533.

Tribe Phrenapatini Solier, 1834

Phrépatides Solier, 1834: 488. Type genus: Phrenapates Gray, 1832.

Genus Delognatha Lacordaire, 1859 [F]

Delognatha Lacordaire, 1859: 315. Type species: Delognatha lacordairei Lacordaire, 1859, subsequent designation (Gebien 1940: 756). Note. The name Delognatha Agassiz, 1846 has been suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy in Opinion 2250 (ICZN 2010).

Delognatha persimilis Gebien, 1928 CRI

Delognatha persimilis Gebien, 1928a: 142.

Genus Phrenapates Gray, 1832 [M]

Phrenapates Gray [in Griffith and Pidgeon], 1832: 91. Type species: Phrenapates bennettii Gray, 1832, monotypy.

Phrenapates bennettii Gray, 1832GUANICCRIPAN / SA

Phrenapates bennettii Gray [in Griffith and Pidgeon], 1832: 91.

Subfamily PIMELIINAE Latreille, 1802

Pimeliariae Latreille, 1802: 166. Type genus: Pimelia Fabricius, 1775.

Tribe Anepsiini LeConte, 1862

Anepsiini LeConte, 1862a: 215. Type genus: Anepsius LeConte, 1851.

Batuliini Horn, 1870: 270. Type genus: Batulius LeConte, 1851.

Anchommini Horn, 1878b: 558. Type genus: Anchomma LeConte, 1858.

Genus Anchomma LeConte, 1858 [N]

Anchomma LeConte, 1858b: 63. Type species: Anchomma costatum LeConte, 1858, monotypy.

Anchomma costatum LeConte, 1858USA (CA)

Anchomma costatum LeConte, 1858b: 63.

Genus Anepsius LeConte, 1851 [M]

Anepsius LeConte, 1851: 147. Type species: Anepsius delicatulus LeConte, 1851, monotypy.

Anepsius delicatulus LeConte, 1851USA (AZ CA NV UT) MEX (SO)

Anepsius delicatulus LeConte, 1851: 148.

Anepsius catenulosus Casey, 1907: 505. Synonymy (with A. atratus Casey): Casey (1911: 254).

Anepsius atratus Casey, 1907: 506. Synonymy: Doyen (1987: 351).

Anepsius brunneus Casey, 1907: 506. Synonymy: Doyen (1987: 351).

Anepsius nebulosus Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius bicolor Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius deficiens Casey, 1907: 507. Synonymy: Doyen (1987: 351).

Anepsius minutus Doyen, 1987USA (TX) MEX (NL)

Anepsius minutus Doyen, 1987: 352.

Anepsius montanus Casey, 1891 [Fig. 8] CAN (AB) USA (CO ND NE NM WY)

Figure 8. 

Anepsius montanus Casey, 1891. Scale bar = 1 mm.

Anepsius montanus Casey, 1891: 55.

Anepsius valens Casey, 1907USA (AZ)

Anepsius valens Casey, 1907: 504.

Genus Batuliodes Casey, 1907 [M]

Batuliodes Casey, 1907: 499. Type species: Batulius rotundicollis LeConte, 1851, original designation.

Batuliodes confluens (Blaisdell, 1923)MEX (BC BS)

Anepsius confluens Blaisdell, 1923: 243.

Anepsius angulatus Blaisdell, 1923: 244. Synonymy: Doyen (1987: 366).

Batuliodes obesus Doyen, 1987USA (CA)

Batuliodes obesus Doyen, 1987: 369.

Batuliodes rotundicollis (LeConte, 1851)USA (AZ CA NV)

Batulius rotundicollis LeConte, 1851: 148.

Batuliodes spatulatus Doyen, 1987USA (AZ CA UT) MEX (SO)

Batuliodes spatulatus Doyen, 1987: 368.

Batuliodes wasbaueri Doyen, 1987USA (CA) MEX (BC)

Batuliodes wasbaueri Doyen, 1987: 367.

Genus Batuliomorpha Doyen, 1987 [F]

Batuliomorpha Doyen, 1987: 359. Type species: Batuliomorpha comata Doyen, 1987, original designation.

Batuliomorpha comata Doyen, 1987USA (AZ CA)

Batuliomorpha comata Doyen, 1987: 361.

Batuliomorpha imperialis Doyen, 1987USA (CA)

Batuliomorpha imperialis Doyen, 1987: 359.

Batuliomorpha tibiodentata Doyen, 1987MEX (BS)

Batuliomorpha tibiodentata Doyen, 1987: 362.

Genus Batulius LeConte, 1851 [M]

Batulius LeConte, 1851: 148. Type species: Batulius setosus LeConte, 1851, subsequent designation (Casey 1907: 497).

Batulius setosus LeConte, 1851USA (AZ CA) MEX (BC)

Batulius setosus LeConte, 1851: 148.

Tribe Asidini Fleming, 1821

Asidadae Fleming, 1821: 51. Type genus: Asida Latreille, 1802.

Astroti Horn, 1870: 289. Type genus: Astrotus J.L. LeConte, 1858.

Craniotini LeConte and Horn, 1883: 361. Type genus: Craniotus LeConte, 1851.

Genus Ardamimicus Smith, 2013 [M]

Ardamimicus Smith, 2013: 601. Type species: Ardamimicus cognatoi Smith, 2013, original designation.

Ardamimicus cognatoi Smith, 2013USA (TX) MEX (CH DU)

Ardamimicus cognatoi Smith, 2013: 602.

Genus Craniotus LeConte, 1851 [M]

Craniotus LeConte, 1851: 142. Type species: Craniotus pubescens LeConte, 1851, monotypy.

Craniotus mardecortesi Aalbu, Smith and Sánchez Piñero, 2015MEX (BC)

Craniotus mardecortesi Aalbu, Smith and Sánchez Piñero, 2015: 94.

Craniotus pubescens LeConte, 1851USA (AZ CA NV) MEX (BC)

Craniotus pubescens LeConte, 1851: 142.

Craniotus blaisdelli Tanner, 1963: 169. Synonymy: Aalbu et al. (2015: 96).

Craniotus triplehorni Aalbu, Smith and Sánchez Piñero, 2015MEX (BC)

Craniotus triplehorni Aalbu, Smith and Sánchez Piñero, 2015: 95.

Genus Ferveoventer Smith, 2013 [M]

Ferveoventer Smith, 2013: 604. Type species: Ferveoventer browni Smith, 2013, original designation.

Ferveoventer browni Smith, 2013USA (NM TX)

Ferveoventer browni Smith, 2013: 605.

Ferveoventer planatus (Champion, 1884)MEX (MO)

Ologlyptus planatus Champion, 1884: 69.

Genus Heterasida Casey, 1912 [F]

Heterasida Casey, 1912: 76, 165. Type species: Pelecyphorus bifurcus LeConte, 1861, original designation.

Heterasida bifurcus (LeConte, 1861)10MEX (BS)

Pelecyphorus bifurcus LeConte, 1861a: 337.

Heterasida connivens (LeConte, 1866)MEX (BS)

Pelecyphorus connivens LeConte, 1866b: 110.

Heterasida tantilla Casey, 1912: 167. Synonymy: Smith (2013: 607).

Heterasida exilis Casey, 1912: 168. Synonymy: Smith (2013: 607).

Genus Litasida Casey, 1912 [F]

Litasida Casey, 1912: 77, 184. Type species: Litasida townsendi Casey, 1912, original designation.

Litasida townsendi Casey, 1912USA (AZ) MEX (CH)

Litasida townsendi Casey, 1912: 185.

Genus Micrasida Smith, 2013 [F]

Micrasida Smith, 2013: 608. Type species: Micrasida obrienorum Smith, 2013, original designation.

Micrasida obrienorum Smith, 2013MEX (NL)

Micrasida obrienorum Smith, 2013: 608.

Genus Microschatia Solier, 1836 [F]

Microschatia Solier, 1836: 474. Type species: Microschatia punctata Solier, 1836, monotypy.

Pycnonotida Casey, 1912: 75, 89. Type species: Microschatia inaequalis LeConte, 1851, original designation. Synonymy: Brown and Doyen (1992: 546).

Acroschatia Wilke, 1922: 269. Type species: Microschatia robusta Horn, 1893, original designation. Synonymy: Brown and Doyen (1992: 546).

Microschatia cedrosensis Brown and Doyen, 1992MEX (BS)

Microschatia cedrosensis Brown and Doyen, 1992: 568.

Microschatia championi Horn, 1893USA (AZ CA) MEX (BC BS)

Microschatia championi Horn, 1893: 140.

Microschatia costulata Brown and Doyen, 1992USA (CA) MEX (BC)

Microschatia costulata Brown and Doyen, 1992: 570.

Microschatia inaequalis LeConte, 1851USA (CA) MEX (BC)

Microschatia inaequalis LeConte, 1851: 129.

Microschatia puncticollis LeConte, 1851: 129. Synonymy: Horn (1870: 282).

Pycnonotida laxicollis Casey, 1912: 91. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida araneoides Casey, 1912: 92. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida inaequalis diversa Casey, 1912: 92. Synonymy: Brown and Doyen (1992: 572).

Pycnonotida impar Casey, 1912: 93. Synonymy: Brown and Doyen (1992: 572).

Microschatia morata Horn, 1878USA (AZ NM) MEX (CH DU SO)

Microschatia morata Horn, 1878a: 56.

Microschatia planata Doyen and Brown, 1992MEX (BC BS)

Microschatia planata Doyen and Brown [in Brown and Doyen], 1992: 576.

Microschatia polita Horn, 1893USA (AZ)

Microschatia polita Horn, 1893: 141.

Microschatia punctata Solier, 1836MEX (HI QU)

Microschatia punctata Solier, 1836: 475.

Microschatia robusta Horn, 1893USA (TX) MEX (CH CO NL TA)

Microschatia robusta Horn, 1893: 142.

Microschatia rockefelleri Pallister, 1954MEX (CH DU)

Microschatia rockefelleri Pallister, 1954: 15.

Microschatia solieri Brown and Doyen, 1992MEX (HI)

Microschatia solieri Brown and Doyen, 1992: 552.

Microschatia sulcipennis LeConte, 1858USA (TX)

Microschatia sulcipennis LeConte, 1858a: 18.

Genus Pelecyphorus Solier, 1836 [M]

Pelecyphorus Solier, 1836: 467. Type species: Pelecyphorus mexicanus Solier, 1836, subsequent designation (Hope 1841: 110).

Subgenus Astrotus LeConte, 1858

Astrotus LeConte, 1858a: 19. Type species: Microschatia contorta LeConte, 1853, original designation.

Pelecyphorus alveolatus (Casey, 1912)USA (TX)

Astrotus alveolatus Casey, 1912: 83.

Pelecyphorus contortus (LeConte, 1853)USA (TX)

Microschatia contorta LeConte, 1853: 446.

Pelecyphorus fasciculatus (Champion, 1892)MEX (MO)

Asida fasciculata Champion, 1892: 495.

Pelecyphorus guanajuatensis (Champion, 1884)MEX (CH DU GU)

Asida guanajuatensis Champion, 1884: 56.

Bothrasida mucorea Wilke, 1922: 270. New synonymy [ADS].

Pelecyphorus hebes (Champion, 1892)MEX (DU)

Ologlyptus hebes Champion, 1892: 506.

Pelecyphorus regularis (Horn, 1870)USA (TX) MEX (TA)

Astrotus regularis Horn, 1870: 290.

Subgenus Pelecyphorus Solier, 1836

Pelecyphorus Solier, 1836: 467. Type species: Pelecyphorus mexicanus Solier, 1836, subsequent designation (Hope 1841: 110).

Pelecyphorus mexicanus Solier, 1836MEX (PU QU VE)

Pelecyphorus mexicanus Solier, 1836: 469.

Subgenus Pleisiasida Smith, 2013

Parasida Casey, 1912: 76, 126 [junior homonym of Parasida Daday, 1904]. Type species: Parasida laciniata Casey, 1912, original designation.

Pleisiasida Smith, 2013: 610. Replacement name for Parasida Casey, 1912.

Pelecyphorus asidoides Solier, 1836MEX (HI ME)11

Pelecyphorus asidoides Solier, 1836: 471.

Parasida zacualpanicola Wilke, 1922: 272. New synonymy [ADS].

Pelecyphorus bibasalis (Casey, 1912)MEX (DU)

Parasida bibasalis Casey, 1912: 128.

Pelecyphorus dispar (Champion, 1892)MEX (CH DU)

Asida dissimilis Champion, 1884: 59 [junior primary homonym of Asida dissimilis Allard, 1869].

Asida dispar Champion, 1892: 496. Replacement name for Asida dissimilis Champion, 1884.

Stenosides kulzeri Pallister, 1954: 12. New synonymy [ADS].

Stenosides bisinuatus Pallister, 1954: 13. New synonymy [ADS].

Parasida trisinuata Pallister, 1954: 22. New synonymy [ADS].

Pelecyphorus fallax (Champion, 1884)MEX (DU FD GU HI ME PU)

Asida fallax Champion, 1884: 57.

Asida favosa Champion, 1884: 58. New synonymy [ADS].

Asida similata Champion, 1884: 58. New synonymy [ADS].

Pelecyphorus foveolatus Solier, 1836MEX (OA VE)

Pelecyphorus foveolatus Solier, 1836: 472.

Pelecyphorus indutus (Champion, 1884)MEX (OA)

Asida induta Champion, 1884: 56.

Ologlyptus bicarinatus Champion, 1884: 69. New synonymy [ADS].

Pelecyphorus intricatus (Champion, 1892)MEX (CH JA)

Asida intricata Champion, 1892: 493.

Pelecyphorus laticollis (Champion, 1884)MEX (DU GU)

Asida laticollis Champion, 1884: 58.

Pelecyphorus liratus (LeConte, 1854)USA (AZ NM) MEX (CH DU)

Euschides liratus LeConte, 1854c: 223.

Parasida laciniata Casey, 1912: 128. New synonymy [ADS].

Parasida cristata Pallister, 1954: 24. New synonymy [ADS].

Pelecyphorus longipennis (Champion, 1884)MEX (OA PU VE)

Asida longipennis Champion, 1884: 56.

Parasida esperanzae Wilke, 1922: 271. New synonymy [ADS].

Parasida mixtecae Wilke, 1922: 271. New synonymy [ADS].

Pelecyphorus obliviosus (Wilke, 1922)MEX (CH DU)

Parasida obliviosa Wilke, 1922: 270.

Pelecyphorus planatulus (Casey, 1912)MEX (DU)

Parasida planatula Casey, 1912: 129.

Pelecyphorus scutellaris (Champion, 1884)MEX (DU FD ME OA PU VE)

Asida scutellaris Champion, 1884: 56.

Parasida tolucana Casey, 1912: 130. New synonymy [ADS].

Pelecyphorus sexcostatus LeConte, 1861MEX (BS)

Pelecyphorus sexcostatus LeConte, 1861a: 337.

Pelecyphorus tristis (Champion, 1884)MEX (PU VE)

Asida tristis Champion, 1884: 55.

Parasida purpusi Wilke, 1922: 271. New synonymy [ADS].

Subgenus Poliorcetes Champion, 1884

Poliorcetes Champion, 1884: 70. Type species: Poliorcetes platesthoides Champion, 1884, monotypy.

Pelecyphorus platesthoides (Champion, 1884)MEX (OA)

Poliorcetes platesthoides Champion, 1884: 71.

Subgenus Sicharbas Champion, 1884

Sicharbas Champion, 1884: 67. Type species: Sicharbas lobatus Champion, 1884, monotypy.

Pelecyphorus debilis (Champion, 1884)MEX (PU)

Astrotus debilis Champion, 1884: 66.

Pelecyphorus erosus (Champion, 1892)MEX (HI)

Astrotus erosus Champion, 1892: 504.

Astrotus nosodermoides Champion, 1892: 505. New synonymy [ADS].

Pelecyphorus lobatus (Champion, 1884)MEX (GE MO)

Sicharbas lobatus Champion, 1884: 67.

Pelecyphorus seticornis (Champion, 1884)MEX (ME)

Astrotus seticornis Champion, 1884: 67.

Astrotus seticornis var. humeralis Champion, 1884: 67. New synonymy [ADS].

Pelecyphorus undatus (Champion, 1892)MEX (DU)

Astrotus undatus Champion, 1892: 504.

Subgenus Stenosides Solier, 1836

Stenosides Solier, 1836: 484. Type species: Stenosides graciliformis Solier, 1836, monotypy.

Pactostoma LeConte, 1858a: 19. Type species: Asida anastomosis Say, 1824, original designation. Synonymy (with Ologlyptus Lacordaire): LeConte (1862a: 222).

Ologlyptus Lacordaire, 1859: 158. Unnecessary replacement name for Stenosides Solier, 1836.

Pelecyphorus anastomosis (Say, 1824)USA (AR AZ CO KS NM TX) MEX (CH DU)

Asida anastomosis Say, 1824a: 256.

Pactostoma anastomosis salebrosa Casey, 1912: 87. Synonymy: Pallister (1954: 12).

Pactostoma breviuscula Casey, 1912: 87. New synonymy [ADS].

Pactostoma exoleta Casey, 1912: 87. New synonymy [ADS].

Pactostoma luteotecta Casey, 1912: 88. New synonymy [ADS].

Pactostoma monticola Casey, 1912: 88. New synonymy [ADS].

Pactostoma obtecta Casey, 1912: 89. New synonymy [ADS].

Pactostoma sigillata Casey, 1912: 89. New synonymy [ADS].

Pelecyphorus graciliformis (Solier, 1836)MEX (FD HI OA PU SL)

Stenosides graciliformis Solier, 1836: 486.

Ologlyptus canus Champion, 1884: 68. New synonymy [ADS].

Ologlyptus sinuaticollis Champion, 1884: 69. New synonymy [ADS].

Pelecyphorus limosus (Champion, 1884) MEX

Astrotus limosus Champion, 1884: 66.

Pelecyphorus texanus (Wickham, 1903)USA (TX)

Ologlyptus texanus Wickham, 1903: 72.

Subgenus Ucalegon Champion, 1884

Ucalegon Champion, 1884: 65. Type species: Ucalegon pulchellus Champion, 1884, monotypy.

Pelecyphorus pulchellus (Champion, 1884)MEX (GE OA)

Ucalegon pulchellus Champion, 1884: 65.

Subgenus Zaleucus Champion, 1892

Zamolxis Champion, 1884: 70 [junior homonym of Zamolxis Stål, 1865]. Type species: Zamolxis dilatatus Champion, 1884, monotypy.

Zaleucus Champion, 1892: 491. Replacement name for Zamolxis Champion, 1884.

Pelecyphorus dilatatus (Champion, 1884)MEX (PU)

Zamolxis dilatatus Champion, 1884: 70.

Genus Philolithus Lacordaire, 1858 [M]

Philolithus Lacordaire [in LeConte], 1858a: 18. Type species: Pelecyphorus carinatus LeConte, 1851, subsequent designation (Casey 1912: 79).

Subgenus Glyptasida Casey, 1912

Glyptasida Casey, 1912: 75, 95. Type species: Pelecyphorus sordidus LeConte, 1853, original designation.

Philolithus aeger (LeConte, 1858)USA (NM TX)

Pelecyphorus aeger LeConte, 1858a: 19.

Glyptasida sycophanta Casey, 1912: 104. Synonymy: Lockwood and Pollock (2009: 21).

Philolithus rugosissimus (Champion, 1884)USA (AZ NM) MEX (CH CO SL)

Asida rugosissima Champion, 1884: 53.

Philolithus sordidus (LeConte, 1853) [Fig. 9] CAN (AB SK) USA (AZ CO KS MT ND NE NM OK SD TX UT WY) MEX (CH CO DU ZA)

Figure 9. 

Philolithus (Glyptasida) sordidus (LeConte, 1853). Scale bar = 1 mm.

Pelecyphorus sordidus LeConte, 1853: 445.

Pelecyphorus subcostatus LeConte, 1853: 446. Synonymy: Henshaw (1882: 255).

Pelecyphorus irregularis LeConte, 1858a: 19. Synonymy: Champion (1892: 492).

Pelecyphorus costipennis LeConte, 1858a: 20. Synonymy: Champion (1892: 492).

Asida interrupta Champion, 1884: 53. Synonymy: Champion (1892: 492).

Glyptasida parvicollis Casey, 1912: 97. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida sordida porcatula Casey, 1912: 97. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida subpubescens Casey, 1912: 98. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens Casey, 1912: 98. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens furtiva Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turgescens obesa Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida procrustes Casey, 1912: 99. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida costipennis fulvisetis Casey, 1912: 100. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida strigipennis Casey, 1912: 100. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida turbulenta Casey, 1912: 101. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida aegra imperfecta Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 7).

Glyptasida aegra pigra Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida aegra plena Casey, 1912: 102. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida heres Casey, 1912: 103. Synonymy: Lockwood and Pollock (2009: 8).

Glyptasida crenicollis Casey, 1912: 103. Synonymy: Lockwood and Pollock (2009: 8).

Subgenus Gonasida Casey, 1912

Gonasida Casey, 1912: 75, 117. Type species: Pelecyphorus elatus LeConte, 1853, original designation.

Philolithus elatus compar (Casey, 1912)USA (CO ID KS NE NM NV OR UT WY)

Gonasida compar Casey, 1912: 120.

Gonasida elata reducta Casey, 1912: 121. New synonymy [based on Brown (1971: 262) unpublished thesis].

Gonasida elata prolixa Casey, 1912: 121. New synonymy [based on Brown (1971: 262) unpublished thesis].

Gonasida aucta Casey, 1912: 122. New synonymy [based on Brown (1971: 262) unpublished thesis].

Philolithus elatus difformis (LeConte, 1854)USA (AZ NM UT) MEX (CH)

Pelecyphorus difformis LeConte, 1854c: 223.

Gonasida alaticollis Casey, 1912: 122. New synonymy [based on Brown (1971: 255) unpublished thesis].

Philolithus elatus elatus (LeConte, 1853)USA (AZ NM TX) MEX (CH)

Pelecyphorus elatus LeConte, 1853: 445.

Gonasida gravida Casey, 1912: 119. New synonymy [based on Brown (1971: 251) unpublished thesis].

Philolithus elatus infernus (Casey, 1912)USA (AZ NM UT)

Gonasida inferna Casey, 1912: 119.

Subgenus Herthasida Wilke, 1922

Herthasida Wilke, 1922: 269. Type species: Asida ingens Champion, 1892, monotypy.

Philolithus ingens (Champion, 1892)MEX (CO)

Asida ingens Champion, 1892: 503.

Subgenus Philolithus Lacordaire, 1858

Philolithus Lacordaire [in LeConte], 1858a: 18. Type species: Pelecyphorus carinatus LeConte, 1851, subsequent designation (Casey 1912: 79).

Philolithus actuosus (Horn, 1870)USA (CA)

Asida actuosa Horn, 1870: 284.

Philolithus adversus (Casey, 1912)USA (AZ)

Pelecyphorus adversus Casey, 1912: 114.

Philolithus aegrotus aegrotus (LeConte, 1861)MEX (BC)

Pelecyphorus aegrotus LeConte, 1861a: 337.

Pelecyphorus aegrotus limbatus Casey, 1912: 107. New synonymy [ADS].

Philolithus carinatus (LeConte, 1851)USA (CA)

Pelecyphorus carinatus LeConte, 1851: 128 [junior secondary homonym of Asida carinata Solier, 1836].

Asida carinifera Gebien, 1910a: 128. Replacement name for Asida carinata (LeConte, 1851)12.

Philolithus densicollis (Horn, 1894) [Fig. 10] CAN (BC) USA (OR WA)

Figure 10. 

Philolithus (Philolithus) densicollis (Horn, 1894). Scale bar = 1 mm.

Asida densicollis Horn, 1894b: 417.

Pelecyphorus corrosus Casey, 1912: 117. Synonymy: Boddy (1965: 137).

Philolithus haruspex ellipsipennis (Casey, 1912)USA (UT)

Pelecyphorus haruspex ellipsipennis Casey, 1912: 116.

Philolithus haruspex haruspex (Casey, 1912)USA (AZ ID OR NV UT)

Pelecyphorus haruspex Casey, 1912: 115.

Philolithus jaegeri (Papp, 1961)USA (CA)

Pelecyphorus jaegeri Papp, 1961b: 107.

Philolithus morbillosus (LeConte, 1858)USA (AZ) MEX (SO)

Pelecyphorus morbillosus LeConte, 1858b: 74.

Pelecyphorus corporalis Casey, 1912: 107. New synonymy [ADS].

Pelecyphorus reptans Casey, 1912: 108. New synonymy [ADS].

Pelecyphorus socer Casey, 1912: 108. New synonymy [ADS].

Pelecyphorus abscissus Casey, 1912: 109. New synonymy [ADS].

Pelecyphorus fumosus Casey, 1912: 109. New synonymy [ADS].

Pelecyphorus parvus Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus morbillosus pacatus Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus morbillosus sobrius Casey, 1912: 110. New synonymy [ADS].

Pelecyphorus piceus Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus piceus crudelis Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus snowi Casey, 1912: 111. New synonymy [ADS].

Pelecyphorus subtenuis Casey, 1912: 112. New synonymy [ADS].

Philolithus opimus (Casey, 1912)USA (CA)

Pelecyphorus opimus Casey, 1912: 115.

Philolithus pantex (Casey, 1912)USA (NV UT)

Pelecyphorus pantex Casey, 1912: 116.

Philolithus porcatus (Papp, 1961)USA (CA)

Pelecyphorus porcatus Papp, 1961b: 109.

Philolithus quadripennis (Casey, 1912)MEX (LC)

Pelecyphorus quadripennis Casey, 1912: 113.

Philolithus reflexus (Casey, 1912)USA (CA)

Pelecyphorus reflexus Casey, 1912: 114.

Philolithus rugosus (Papp, 1961)USA (CA)

Pelecyphorus rugosus Papp, 1961c: 157.

Philolithus sophistes (Casey, 1912)USA (CA)

Pelecyphorus sophistes Casey, 1912: 113.

Philolithus uteanus (Casey, 1924)USA (UT)

Pelecyphorus uteanus Casey, 1924: 308.

Subgenus Tisamenes Champion, 1884

Tisamenes Champion, 1884: 64. Type species: Tisamenes truquii Champion, 1884, monotypy.

Philolithus truquii (Champion, 1884)MEX (FD HI)

Tisamenes truquii Champion, 1884: 64.

Genus Stenomorpha Solier, 1836 [F]

Stenomorpha Solier, 1836: 487. Type species: Stenomorpha blapsoides Solier, 1836, subsequent designation (Desmarest 1860: 150).

Euschides LeConte, 1851: 127. Unnecessary replacement name for Stenomorpha Solier, 1836.

Subgenus Asidina Casey, 1912

Asidina Casey, 1912: 169. Type species: Pelecyphorus parallelus LeConte, 1851, original designation.

Stenomorpha confluens (LeConte, 1851)USA (AZ CA) MEX (BC SO)

Pelecyphorus confluens LeConte, 1851: 128.

Stenomorpha parallela (LeConte, 1851)USA (AZ CA) MEX (SO)

Pelecyphorus parallelus LeConte, 1851: 128 [junior secondary homonym of Asida parallela Solier, 1836].

Asida neglecta Gebien, 1910a: 134. Replacement name for Asida parallela (LeConte, 1851)13.

Asidina teres Casey, 1912: 171. Synonymy: Triplehorn and Brown (1971: 84).

Asida parallela terricola Blaisdell, 1923: 254. Synonymy: Triplehorn and Brown (1971: 84).

Stenomorpha rugicollis (Triplehorn and Brown, 1971)USA (AZ)

Asidina rugicollis Triplehorn and Brown, 1971: 76.

Stenomorpha semilaevis (Horn, 1870)USA (AZ CA NV)

Asida semilaevis Horn, 1870: 284.

Subgenus Asidopsis Casey, 1912

Asidopsis Casey, 1912: 77, 185. Type species: Asida opaca Say, 1824, original designation.

Stenomorpha abbreviata (Casey, 1912)USA (NM)

Asidopsis abbreviata Casey, 1912: 198.

Stenomorpha cochisensis (Casey, 1912)USA (AZ NM)

Asidopsis cochisensis Casey, 1912: 189.

Stenomorpha coenosa (Casey, 1912)USA (NM)

Asidopsis coenosa Casey, 1912: 200.

Stenomorpha collaris (Champion, 1892)MEX (AG GU JA)

Asida marginicollis Champion, 1884: 60 [junior primary homonym of Asida marginicollis Rosenhauer, 1856].

Asida collaris Champion, 1892: 499. Replacement name for Asida marginicollis Champion, 1884.

Stenomorpha collega (Casey, 1912)USA (KS)

Asidopsis collega Casey, 1912: 198.

Stenomorpha consentanea (Casey, 1912)USA (CA)

Asidopsis consentanea Casey, 1912: 192.

Stenomorpha divaricata (Blaisdell, 1923)MEX (BS)

Asida divaricata Blaisdell, 1923: 255.

Stenomorpha dolosa (Casey, 1912)USA (MT SD WY)

Asidopsis dolosa Casey, 1912: 194.

Stenomorpha durangoensis (Casey, 1912)MEX (DU)

Asidopsis durangoensis Casey, 1912: 201.

Stenomorpha eximia (Casey, 1912)USA (TX)

Asidopsis eximia Casey, 1912: 188.

Stenomorpha forreri (Champion, 1884)MEX (DU)

Asida forreri Champion, 1884: 55.

Stenomorpha gracilipes (Casey, 1912)USA (AZ)

Asidopsis gracilipes Casey, 1912: 189.

Stenomorpha humeralis (Triplehorn and Flores, 2002)MEX (CH)

Asidopsis humeralis Triplehorn and Flores, 2002: 288.

Stenomorpha immunda (Casey, 1912)USA (NM) MEX (DU)

Asidopsis immunda Casey, 1912: 199.

Stenomorpha macra (Horn, 1883)USA (AZ NM)

Asidopsis macra Horn, 1883: 304.

Stenomorpha mancipata (Horn, 1878)USA (NM) MEX (CH)

Asida mancipata Horn, 1878a: 56.

Asidopsis woodgatei Casey, 1912: 197. Synonymy: Triplehorn and Flores (2002: 285).

Stenomorpha nitidula (Casey, 1912)USA (NM)

Asidopsis nitidula Casey, 1912: 196.

Stenomorpha obsidiana (Casey, 1912)USA (CO)

Asidopsis obsidiana Casey, 1912: 193.

Stenomorpha olsoni (Triplehorn and Flores, 2002)USA (AZ)

Asidopsis olsoni Triplehorn and Flores, 2002: 286.

Stenomorpha opaca (Say, 1824) [Fig. 11] CAN (AB SK) USA (CO KS MT ND NE NM SD TX)

Figure 11. 

Stenomorpha (Asidopsis) opaca (Say, 1824). Scale bar = 1 mm.

Asida opaca Say, 1824a: 254.

Stenomorpha pinalica (Casey, 1912)USA (AZ)

Asidopsis pinalica Casey, 1912: 190.

Stenomorpha planata (Horn, 1894)USA (CA) MEX

Asida planata Horn, 1894b: 415.

Stenomorpha polita futilis (Casey, 1912)USA (KS)

Asidopsis polita futilis Casey, 1912: 194.

Stenomorpha polita polita (Say, 1824)CAN (AB SK) USA (CO ID KS MT NE NM OK SD TX WY)

Asida polita Say, 1824a: 255.

Stenomorpha polita subopaca (Casey, 1912)USA (KS)

Asidopsis polita subopaca Casey, 1912: 193.

Stenomorpha quadricollis (Horn, 1880)USA (AZ NM)

Asidopsis quadricollis Horn, 1880: 151.

Stenomorpha servilis (Casey, 1912)USA (CO)

Asidopsis servilis Casey, 1912: 199.

Stenomorpha suavis (Casey, 1912)USA (AZ)

Asidopsis suavis Casey, 1912: 190.

Stenomorpha tensa (Casey, 1912)USA (CO)

Asidopsis tensa Casey, 1912: 197.

Subgenus Bothrasida Casey, 1912

Bothrasida Casey, 1912: 76, 122. Type species: Asida clathrata Champion, 1884, original designation.

Stenomorpha baroni (Casey, 1912)MEX (GE)

Bothrasida baroni Casey, 1912: 124.

Stenomorpha clathrata (Champion, 1884)MEX (GE ME MO OA PU)

Asida clathrata Champion, 1884: 54.

Stenomorpha funesta (Champion, 1884)MEX (PU)

Asida funesta Champion, 1884: 53.

Bothrasida sanctae-agnae Wilke, 1922: 270. New synonymy [ADS].

Subgenus Megasida Casey, 1912

Megasida Casey, 1912: 77, 202. Type species: Asida obliterata Champion, 1892, original designation.

Stenomorpha foeda (Champion, 1892)MEX (CO DU)

Asida foeda Champion, 1892: 498.

Stenomorpha latissima (Champion, 1892)MEX (DU)

Asida latissima Champion, 1892: 500.

Stenomorpha magnifica (Pallister, 1954)MEX (DU)

Megasida magnifica Pallister, 1954: 30.

Stenomorpha moricoides (Champion, 1892)MEX (CO DU)

Asida moricoides Champion, 1892: 497.

Stenomorpha obliterata (Champion, 1892)USA (NM TX) MEX (CH)

Asida obliterata Champion, 1892: 497.

Stenomorpha rufipes (Champion, 1884)MEX (SL)

Asida rufipes Champion, 1884: 62.

Stenomorpha segregata (Champion, 1892)MEX (CH CO DU)

Asida segregata Champion, 1892: 497.

Stenomorpha tarda (Champion, 1892)MEX (CO)

Asida tarda Champion, 1892: 498.

Stenomorpha tenuicollis (Triplehorn, 1967)USA (AZ NM TX)

Megasida tenuicollis Triplehorn, 1967: 40.

Stenomorpha zacatecensis (Pallister, 1954)MEX (ZA)

Megasida zacatecensis Pallister, 1954: 32.

Subgenus Notiasida Casey, 1912

Notiasida Casey, 1912: 76, 124. Type species: Notiasida abstrusa Casey, 1912, original designation.

Stenomorpha abstrusa (Casey, 1912)MEX (FD)

Notiasida abstrusa Casey, 1912: 125.

Stenomorpha evertissima (Casey, 1912)MEX (CH DU)

Notiasida evertissima Casey, 1912: 126.

Stenomorpha geminata (Champion, 1892)MEX (CH DU)

Asida geminata Champion, 1892: 492.

Stenomorpha lata (Champion, 1884)MEX (SL)

Asida lata Champion, 1884: 60.

Stenomorpha lugubris (Wilke, 1922)MEX (SL)

Pelecyphorus lugubris Wilke, 1922: 269.

Stenomorpha suturalis (Champion, 1884)MEX (FD OA VE)

Asida suturalis Champion, 1884: 55.

Subgenus Platasida Casey, 1912

Platasida Casey, 1912: 77, 182. Type species: Asida embaphionides Horn, 1894, original designation.

Stenomorpha embaphionides (Horn, 1894)MEX (BS)

Asida embaphionides Horn, 1894b: 419.

Asida flaccida Horn, 1896: 379. New synonymy [ADS].

Subgenus Pycnomorpha Motschulsky, 1870

Pycnomorpha Motschulsky, 1870: 398. Type species: Pycnomorpha californica Motschulsky, 1870, monotypy.

Stenomorpha californica (Motschulsky, 1870)MEX (BC)

Pycnomorpha californica Motschulsky, 1870: 399.

Stenomorpha gabbii (Horn, 1880)USA (CA) MEX (BS)

Asida gibbicollis Horn, 1870: 288 [junior primary homonym of Asida gibbicollis Pérez Arcas, 1865].

Asida gabbii Horn, 1880: 152. Replacement name for Asida gibbicollis Horn, 1870.

Stenomorpha tumidicollis Blaisdell, 1943MEX (BC)

Stenomorpha tumidicollis Blaisdell, 1943: 226.

Subgenus Stenomorpha Solier, 1836

Stenomorpha Solier, 1836: 487. Type species: Stenomorpha blapsoides Solier, 1836, subsequent designation (Desmarest 1860: 150).

Psilomera Motschulsky, 1870: 400. Type species: Pelecyphorus angulatus LeConte, 1851, monotypy. New synonymy [YB].

Stenomorpha advena (Casey, 1912)USA (CO)

Euschides advena Casey, 1912: 143.

Stenomorpha amplicollis (Casey, 1912)USA (CA)

Euschides amplicollis Casey, 1912: 153.

Stenomorpha angulata (LeConte, 1851)USA (CA)

Pelecyphorus angulatus LeConte, 1851: 127.

Stenomorpha blanda (Champion, 1884)MEX (AG GU JA)

Asida blanda Champion, 1884: 63.

Stenomorpha blapsoides alutacea Wilke, 1922MEX (ME)

Stenomorpha blapsoides alutacea Wilke, 1922: 272.

Stenomorpha blapsoides blapsoides Solier, 1836MEX (CO FD GU JA ME OA PU SI VE)

Stenomorpha blapsoides Solier, 1836: 491.

Stenomorpha brevimargo (Casey, 1912)USA (AZ)

Euschides brevimargo Casey, 1912: 134.

Stenomorpha caliginosa (Casey, 1912)USA (AZ)

Euschides caliginosus Casey, 1912: 137.

Stenomorpha captiosa (Horn, 1870)USA (CA)

Asida captiosa Horn, 1870: 287.

Stenomorpha clarissae Wilke, 1922MEX (ME)

Stenomorpha clarissae Wilke, 1922: 273.

Stenomorpha compressa (Horn, 1870)USA (CA)

Asida lecontei var. compressa Horn, 1870: 287.

Stenomorpha congruens congruens (Casey, 1912)USA (NM)

Euschides congruens Casey, 1912: 164.

Stenomorpha congruens lubrica (Casey, 1912)USA (AZ)

Euschides congruens lubricus Casey, 1912: 164.

Stenomorpha consobrina (Horn, 1870)USA (ID OR)

Asida consobrina Horn, 1870: 287.

Stenomorpha consors (Casey, 1912)USA (NM)

Euschides consors Casey, 1912: 162.

Stenomorpha consueta (Casey, 1912)USA (AZ)

Euschides consuetus Casey, 1912: 161.

Stenomorpha convexa (LeConte, 1859)USA (CA KS)

Euschides convexa LeConte, 1859a: 14.

Stenomorpha convexicollis (LeConte, 1854)USA (CA CO KS NM TX) MEX (CH DU)

Euschides convexicollis LeConte, 1854c: 224.

Stenomorpha corrugans (Casey, 1912)USA (AZ)

Euschides corrugans Casey, 1912: 143.

Stenomorpha costata Solier, 1836MEX (ME VE)

Stenomorpha costata Solier, 1836: 490.

Stenomorpha crassa (Casey, 1912)USA (CA)

Euschides crassus Casey, 1912: 154.

Stenomorpha cressoni (Blaisdell, 1933)USA (CA)

Euschides cressoni Blaisdell, 1933b: 191.

Stenomorpha cribrata (Casey, 1912)USA (NM)

Euschides cribratus Casey, 1912: 140.

Stenomorpha crinita (Casey, 1912)USA (OR)

Euschides crinitus Casey, 1912: 148.

Stenomorpha deceptor (Casey, 1912)USA (CA)

Euschides deceptor Casey, 1912: 154.

Stenomorpha directa (Casey, 1912)USA (AZ)

Euschides directa Casey, 1912: 141.

Stenomorpha evanescens (Casey, 1912)USA (CA)

Euschides evanescens Casey, 1912: 146.

Stenomorpha facilis (Casey, 1912)USA (KS)

Euschides facilis Casey, 1912: 165.

Stenomorpha fastigiosa (Casey, 1912)USA (NM)

Euschides fastigiosus Casey, 1912: 157.

Stenomorpha globicollis (Casey, 1912)USA (NE)

Euschides globicollis Casey, 1912: 158.

Stenomorpha gracilior (Casey, 1912)USA (KS)

Euschides gracilior Casey, 1912: 160.

Stenomorpha gravidipes (Casey, 1912)USA (CA)

Euschides gravidipes Casey, 1912: 155.

Stenomorpha huachucae (Casey, 1912)USA (AZ)

Euschides huachucae Casey, 1912: 162.

Stenomorpha implicans (Casey, 1912)USA (AZ)

Euschides implicans Casey, 1912: 136.

Stenomorpha inhabilis inhabilis (Casey, 1912)USA (NM)

Euschides inhabilis Casey, 1912: 156.

Stenomorpha inhabilis retusa (Casey, 1912)USA (KS)

Euschides inhabilis retusus Casey, 1912: 157.

Stenomorpha integer (Casey, 1912)USA (CA)

Euschides integer Casey, 1912: 153.

Stenomorpha lecontei lecontei (Horn, 1870)USA (CA)

Pelecyphorus costipennis LeConte, 1859b: 76 [junior primary homonym of Pelecyphorus costipennis LeConte, 1858].

Asida lecontei Horn, 1870: 286. Replacement name for Asida costipennis (LeConte, 1859).

Stenomorpha lecontei gigantea (Blaisdell, 1921)USA (CA)

Euschides lecontei gigantea Blaisdell, 1921b: 209.

Stenomorpha lecontella lecontella (Blaisdell, 1936)USA (CA)

Euschides lecontella Blaisdell, 1936b: 227.

Stenomorpha lecontella tempestalis (Blaisdell, 1936)USA (CA)

Euschides lecontella tempestalis Blaisdell, 1936b: 229.

Stenomorpha luctata (Horn, 1870)USA (CA NV)

Asida luctata Horn, 1870: 286.

Stenomorpha marginata duplicans (Casey, 1912)USA [AZ]

Euschides marginatus duplicans Casey, 1912: 136.

Stenomorpha marginata esuriens (Casey, 1912)USA (CA)

Euschides marginatus esuriens Casey, 1912: 137.

Stenomorpha marginata marginata (LeConte, 1851)USA (AZ CA NM TX) MEX (CH SO)

Pelecyphorus marginatus LeConte, 1851: 128.

Stenomorpha maritima imula (Casey, 1912)USA (CA)

Euschides maritimus imulus Casey, 1912: 151.

Stenomorpha maritima maritima (Casey, 1912)USA (CA)

Euschides maritimus Casey, 1912: 151.

Stenomorpha mckittricki (Pierce, 1954)USA (CA)14

Parasida mckittricki Pierce, 1954a: 43.

Stenomorpha montezuma Wilke, 1922MEX (DU)

Stenomorpha montezuma Wilke, 1922: 272.

Stenomorpha musiva Wilke, 1922MEX (ME)

Stenomorpha musiva Wilke, 1922: 273.

Stenomorpha neutralis (Casey, 1912)USA (CA)

Euschides neutralis Casey, 1912: 146.

Stenomorpha oblonga (Casey, 1924)USA (NM)

Euschides oblongus Casey, 1924: 309.

Stenomorpha obovata gliscans (Casey, 1912)USA (AZ)

Euschides obovatus gliscans Casey, 1912: 161.

Stenomorpha obovata nitidipennis (Casey, 1912)USA (AZ)

Euschides obovatus nitidipennis Casey, 1912: 160.

Stenomorpha obovata obovata (LeConte, 1851)USA (AZ CA TX) MEX (CH)

Euschides obovata LeConte, 1851: 127.

Stenomorpha oregonensis (Casey, 1924)USA (OR)

Euschides oregonensis Casey, 1924: 309.

Stenomorpha orizabae Wilke, 1922MEX (VE)

Stenomorpha orizabae Wilke, 1922: 273.

Stenomorpha papagoana (Casey, 1912)USA (AZ)

Euschides papagoanus Casey, 1912: 163.

Stenomorpha pollens pollens (Casey, 1912)USA (AZ)

Euschides pollens Casey, 1912: 134.

Stenomorpha pollens proxima (Casey, 1912)USA (AZ)

Euschides pollens proximus Casey, 1912: 134.

Stenomorpha procurrens (Casey, 1912)USA (AZ)

Euschides procurrens Casey, 1912: 137.

Stenomorpha puncticollis (LeConte, 1866)USA (OR WA)

Euschides puncticollis LeConte, 1866b: 111 [junior secondary homonym of Asida puncticollis Solier, 1836].

Asida robusta Gebien, 1910a: 135. Replacement name for Asida puncticollis (LeConte, 1866)15.

Stenomorpha rimata rimata (LeConte, 1854)USA (TX)

Pelecyphorus rimatus LeConte, 1854c: 223.

Stenomorpha rimata subplanata (Casey, 1912)USA (AZ)

Euschides rimatus subplanatus Casey, 1912: 139.

Stenomorpha rudis (Casey, 1912)USA (AZ)

Euschides rudis Casey, 1912: 139.

Stenomorpha rugata (Casey, 1912)USA (AZ)

Euschides rugatus Casey, 1912: 138.

Stenomorpha rustica (Casey, 1912)USA (AZ)

Euschides rusticus Casey, 1912: 135.

Stenomorpha satiata (Casey, 1912)USA (AZ)

Euschides satiatus Casey, 1912: 138.

Stenomorpha semirufa (Casey, 1912)USA (AZ)

Euschides semirufus Casey, 1912: 140.

Stenomorpha severa (Casey, 1912)USA (NM)

Euschides severus Casey, 1912: 144.

Stenomorpha socialis (Casey, 1912)USA (NM)

Euschides socialis Casey, 1912: 162.

Stenomorpha speculata (Blaisdell, 1936)USA (CA)

Euschides speculatus Blaisdell, 1936b: 225.

Stenomorpha sphaericollis (Champion, 1884)MEX (AG SL)

Asida sphaericollis Champion, 1884: 64.

Stenomorpha sponsor (Casey, 1912)USA (AZ)

Euschides sponsor Casey, 1912: 135.

Stenomorpha spurcans (Casey, 1912)USA (CA)

Euschides spurcans Casey, 1912: 145.

Stenomorpha strigosula (Casey, 1912)USA (AZ)

Euschides strigosulus Casey, 1912: 163.

Stenomorpha subcruenta (Casey, 1912)USA (NM)

Euschides subcruentus Casey, 1912: 140.

Stenomorpha subcylindrica (Horn, 1870)USA (AZ)

Asida marginata var. subcylindrica Horn, 1870: 288.

Stenomorpha subelegans (Casey, 1912)USA (CA)

Euschides subelegans Casey, 1912: 152.

Stenomorpha tetrica (Casey, 1912)USA (UT)

Euschides tetricus Casey, 1912: 149.

Stenomorpha tularensis (Casey, 1912)USA (CA)

Euschides tularensis Casey, 1912: 152.

Stenomorpha uhdei Wilke, 1922MEX (ME)

Stenomorpha uhdei Wilke, 1922: 273.

Stenomorpha umbrosa (Champion, 1884)MEX (GU)

Asida umbrosa Champion, 1884: 62.

Stenomorpha vigens (Casey, 1912)USA (AZ)

Euschides vigens Casey, 1912: 159.

Subgenus Stethasida Casey, 1912

Stethasida Casey, 1912: 78, 203. Type species: Pelecyphorus muricatulus LeConte, 1851, original designation.

Stenomorpha flohri (Champion, 1892)MEX (JA)

Asida flohri Champion, 1892: 496.

Stenomorpha muricatula (LeConte, 1851)USA (CA)

Pelecyphorus muricatulus LeConte, 1851: 128.

Asida angustula Casey, 1890b: 370. New synonymy [ADS].

Stethasida stricta Casey, 1912: 210. New synonymy [ADS].

Stethasida muricatula languida Casey, 1912: 211. New synonymy [ADS].

Stethasida pertinax Casey, 1912: 211. New synonymy [ADS].

Stethasida socors Casey, 1912: 212. New synonymy [ADS].

Stethasida angustula inepta Casey, 1912: 213. New synonymy [ADS].

Stethasida tenax Casey, 1912: 213. New synonymy [ADS].

Stethasida vegrandis Casey, 1912: 214. New synonymy [ADS].

Stenomorpha obsoleta (LeConte, 1851)USA (CA)

Pelecyphorus obsoletus LeConte, 1851: 128.

Stethasida obsoleta expansa Casey, 1912: 205. New synonymy [ADS].

Stethasida obsoleta opacella Casey, 1912: 205. New synonymy [ADS].

Stethasida brevipes Casey, 1912: 206. New synonymy [ADS].

Stethasida torpida Casey, 1912: 206. New synonymy [ADS].

Stethasida convergens Casey, 1912: 207. New synonymy [ADS].

Stethasida discreta Casey, 1912: 207. New synonymy [ADS].

Stethasida longula Casey, 1912: 207. New synonymy [ADS].

Stethasida adumbrata Casey, 1912: 208. New synonymy [ADS].

Stethasida occulta Casey, 1912: 208. New synonymy [ADS].

Stethasida tarsalis Casey, 1912: 208. New synonymy [ADS].

Stethasida unica Casey, 1912: 209. New synonymy [ADS].

Pelecyphorus laevigatus Papp, 1961c: 159. New synonymy [ADS].

Subgenus Trichiasida Casey, 1912

Trichiasida Casey, 1912: 77, 172. Type species: Pelecyphorus hirsutus LeConte, 1851, original designation.

Stenomorpha acerba (Horn, 1878)USA (AZ NV UT)

Asida acerba Horn, 1878a: 56.

Stenomorpha difficilis (Champion, 1884)MEX (HI ME SL)

Asida difficilis Champion, 1884: 61.

Trichiasida eremica Wilke, 1922: 274. New synonymy [ADS].

Stenomorpha hirsuta (LeConte, 1851)USA (CA)

Pelecyphorus hirsutus LeConte, 1851: 127.

Trichiasida lineatopilosa Casey, 1912: 175. New synonymy [ADS].

Stenomorpha hispidula (LeConte, 1851)USA (AZ CA)

Pelecyphorus hispidulus LeConte, 1851: 127.

Trichiasida tenella Casey, 1912: 177. New synonymy [ADS].

Stenomorpha horrida (Champion, 1892)USA (TX) MEX (TA)

Asida horrida Champion, 1892: 500.

Stenomorpha idahoensis (Boddy, 1957)USA (ID)

Trichiasida idahoensis Boddy, 1957: 187.

Stenomorpha ignava (Casey, 1912)USA (AZ)

Trichiasida ignava Casey, 1912: 180.

Stenomorpha impetrata (Horn, 1894)USA (CA)

Asida impetrata Horn, 1894b: 418.

Stenomorpha impotens (Casey, 1912)USA (AZ)

Trichiasida impotens Casey, 1912: 180.

Stenomorpha lutulenta (Doyen, 1990)MEX (JA)

Trichiasida lutulenta Doyen, 1990: 225.

Stenomorpha palmeri (Champion, 1884)MEX (SL)

Asida palmeri Champion, 1884: 59.

Stenomorpha pubescens (Champion, 1884) MEX

Asida pubescens Champion, 1884: 61.

Stenomorpha subpilosa Solier, 1836MEX (PU)

Stenomorpha subpilosa Solier, 1836: 490.

Stenomorpha thoracica (Champion, 1884) MEX

Asida thoracica Champion, 1884: 62.

Stenomorpha unicostata (Champion, 1892)MEX (GE)

Asida unicostata Champion, 1892: 501.

Stenomorpha villosa (Champion, 1884)MEX (PU)

Asida villosa Champion, 1884: 60.

Trichiasida duplex Casey, 1912: 17816. New synonymy [ADS].

[incertae sedis]

Stenomorpha catalinae (Blaisdell, 1923)MEX (BS)

Asida catalinae Blaisdell, 1923: 256.

Stenomorpha furcata (Champion, 1892)USA (TX) MEX (CO DU)

Asida furcata Champion, 1892: 499.

Stenomorpha granicollis (Blaisdell, 1923)MEX (SO)

Asida granicollis Blaisdell, 1923: 256.

Stenomorpha roosevelti Smith, Miller and Wheeler, 2011MEX (CO)

Stenomorpha roosevelti Smith, Miller and Wheeler, 2011: 30.

Stenomorpha spinimana (Champion, 1892)MEX (DU) New combination [ADS].

Asida spinimanus Champion, 1892: 494.

Stenomorpha subvittata (Horn, 1894)MEX (BS)

Asida subvittata Horn, 1894b: 416.

Stenomorpha tenebrosa (Champion, 1892)MEX (CO) New combination [ADS].

Asida tenebrosa Champion, 1892: 495.

Stenomorpha wickhami (Horn, 1894)USA (AZ CA)

Asida wickhami Horn, 1894b: 420.

Asidina liberta Casey, 1912: 171. Synonymy: Triplehorn and Brown (1971: 84).

Tribe Branchini LeConte, 1862

Branchini LeConte, 1862a: 222. Type genus: Branchus LeConte, 1862

Genus Anectus Horn, 1866 [M]

Anectus Horn, 1866: 399. Type species: Anectus vestitus Horn, 1866, monotypy.

Anectus vestitus Horn, 1866 HON

Anectus vestitus Horn, 1866: 399.

Genus Branchus LeConte, 186217 [M]

Branchus LeConte, 1862a: 222. Type species: Branchus floridanus LeConte, 1862, monotypy.

Branchus floridanus LeConte, 1862USA (FL)

Branchus floridanus LeConte, 1862a: 223.

Branchus geraceorum Steiner, 2006 BAH

Branchus geraceorum Steiner, 2006: 10.

Branchus jamaicensis Marcuzzi, 1977 JAM

Branchus jamaicensis Marcuzzi, 1977: 10.

Branchus obscurus Horn, 1866MEX (GE) GUANIC

Branchus obscurus Horn, 1866: 398.

Branchus opatroides Champion, 1892MEX (JA VE)

Branchus opatroides Champion, 1892: 507.

Branchus saxatilis Steiner, 2005 BAH

Branchus saxatilis Steiner, 2005: 443.

Branchus whiteheadi Steiner, 1991USA (TX)

Branchus whiteheadi Steiner, 1991: 426.

Branchus woodii LeConte, 1866 BAH CUB

Branchus woodii LeConte, 1866b: 111.

Genus Oxinthas Champion, 1884 [M]

Oxinthas Champion, 1884: 72. Type species: Oxinthas praocioides Champion, 1884, monotypy.

Oxinthas nicaraguensis Merkl, 1992 NIC

Oxinthas nicaraguensis Merkl, 1992: 89.

Oxinthas praocioides Champion, 1884MEX (OA)

Oxinthas praocioides Champion, 1884: 72.

Tribe Cnemeplatiini Jacquelin du Val, 1861

Cnéméplatiites Jacquelin du Val, 1861: 286. Type genus: Cnemeplatia Costa, 1847.

Subtribe Cnemeplatiina Jacquelin du Val, 1861

Cnéméplatiites Jacquelin du Val, 1861: 286. Type genus: Cnemeplatia Costa, 1847.

Genus Alaudes Horn, 1870 [M]

Alaudes Horn, 1870: 361. Type species: Alaudes singularis Horn, 1870, monotypy.

Alaudes alternatus Fall, 1928USA (CA)

Alaudes alternata Fall, 1928: 148.

Alaudes setigerus Blaisdell, 1919USA (CA)

Alaudes setigera Blaisdell, 1919a: 310.

Alaudes singularis Horn, 1870USA (CA ID NV OR)

Alaudes singularis Horn, 1870: 362.

Alaudes squamosa Blaisdell, 1919a: 309. New synonymy [RLA].

Alaudes testacea Blaisdell, 1919a: 311. New synonymy [RLA].

Alaudes fallax Fall, 1928: 150. New synonymy [RLA].

Genus Lepidocnemeplatia Bousquet and Bouchard, new genus [F]

Lepidocnemeplatia Bousquet and Bouchard, new genus. Type species: Cnemeplatia sericea Horn, 1870.

Lepidocnemeplatia laticollis (Champion, 1885)MEX (ME) NICCRIPAN / SA

Cnemeplatia laticollis Champion, 1885: 136.

Lepidocnemeplatia sericea (Horn, 1870)USA (AZ CA NV OR TX WA) MEX (BS CH DU MO NL PU SO VE) NIC

Cnemeplatia sericea Horn, 1870: 360.

Tribe Cnemodinini Gebien, 1910

Cnemodini Horn, 1870: 266. Type genus: Cnemodus Horn, 1870 (= Cnemodinus Cockerell, 1906).

Cnemodininae Gebien, 1910a: 4. Type genus: Cnemodinus Cockerell, 1906.

Genus Cnemodinus Cockerell, 1906 [M]

Cnemodus Horn, 1870: 266 [junior homonym of Cnemodus Herrich-Schaeffer, 1850]. Type species: Cnemodus testaceus Horn, 1870, monotypy.

Cnemodinus Cockerell, 1906: 242. Replacement name for Cnemodus Horn, 1870.

Cnemodinus angustus (Casey, 1907)USA (AZ)

Cnemodus angustus Casey, 1907: 284.

Cnemodinus subhyalinus (Casey, 1907)USA (UT)

Cnemodus subhyalinus Casey, 1907: 285.

Cnemodinus testaceus (Horn, 1870)USA (AZ CA)

Cnemodus testaceus Horn, 1870: 266.

Tribe Coniontini G.R. Waterhouse, 1858

Coniontidae G.R. Waterhouse, 1858: 59. Type genus: Coniontis Eschscholtz, 1829.

Coelini Casey, 1907: 500. Type genus: Coelus Eschscholtz, 1829.

Eusatti Doyen, 1984b: 11. Type genus: Eusattus LeConte, 1851.

Genus Coelus Eschscholtz, 1829 [M]

Coelus Eschscholtz, 1829: 5. Type species: Coelus ciliatus Eschscholtz, 1829, monotypy.

Coelomorpha Casey, 1890a: 182. Type species: Coelomorpha maritima Casey, 1890, monotypy. Synonymy: Doyen (1972: 371).

Pseudocoelus Casey, 1908: 152. Type species: Coelus pacificus Fall, 1897, subsequent designation (Doyen 1976: 608). Synonymy: Blaisdell (1919b: 322).

Coelus ciliatus Eschscholtz, 1829 [Fig. 12] CAN (BC) USA (CA OR WA) MEX (BC)

Figure 12. 

Coelus ciliatus Eschscholtz, 1829. Scale bar = 1 mm.

Coelus ciliatus Eschscholtz, 1829: 5.

Coelus arenarius Casey, 1890a: 179. Synonymy: Doyen (1976: 614).

Coelus latus Casey, 1895: 612. Synonymy (with C. arenarius Casey): Fall (1901: 166).

Coelus curtulus Casey, 1895: 612. Synonymy: Fall (1901: 166).

Coelus ciliatus longulus Casey, 1908: 154. Synonymy: Blaisdell (1919b: 333).

Coelus debilis Casey, 1908: 155. Synonymy: Doyen (1976: 614).

Coelus sternalis Casey, 1908: 156. Synonymy: Doyen (1976: 614).

Coelus obscurus Casey, 1908: 156. Synonymy (with C. arenarius Casey): Blaisdell (1919b: 334).

Coelus scolopax Casey, 1908: 157. Synonymy (with C. arenarius Casey): Blaisdell (1919b: 334).

Coelus amplicollis Casey, 1908: 157. Synonymy (with C. latus Casey): Blaisdell (1919b: 334).

Coelus ciliatus var. sparsus Blaisdell, 1919b: 325. Synonymy (with C. ciliatus debilis Casey): Gebien (1938: 409).

Coelus globosus LeConte, 1851USA (CA) MEX (BC)

Coelus globosus LeConte, 1851: 133.

Coelus grossus Casey, 1890a: 178. Synonymy: Doyen (1976: 617).

Coelus solidus Casey, 1908: 153. Synonymy (with C. globosus grossus Casey): Blaisdell (1919b: 333).

Coelus saginatus Casey, 1908: 154. Synonymy: Doyen (1976: 617).

Coelus gracilis Blaisdell, 1939USA (CA)

Coelus gracilis Blaisdell, 1939a: 16.

Coelus maritimus (Casey, 1889)MEX (BC)

Coelomorpha maritima Casey, 1890a: 183.

Coelomorpha pallens Casey, 1908: 160. Synonymy: Doyen (1976: 620).

Coelus pacificus Fall, 1897USA (CA)

Coelus pacificus Fall, 1897: 241.

Coelus remotus Fall, 1897: 241. Synonymy: Doyen (1976: 618).

Genus Coniontis Eschscholtz, 1829 [F]

Coniontis Eschscholtz, 1829: 7. Type species: Coniontis viatica Eschscholtz, 1829, subsequent designation (Casey 1908: 57).

Coelotaxis Horn, 1876a: 200. Type species: Coelotaxis punctulata Horn, 1876, subsequent designation (Gebien 1938: 289). Synonymy: Doyen (1972: 373).

Coniontellus Casey, 1890b: 388. Type species: Coniontis obesa LeConte, 1851, subsequent designation (Casey 1908: 57). Synonymy: Doyen (1972: 373).

Coniontides Casey, 1908: 57, 78. Type species: Coniontis lata LeConte, 1866, original designation. Synonymy: Doyen (1972: 373).

Crypticomorpha Casey, 1908: 81, 140. Type species: Coniontis tenuis Casey, 1908, monotypy. Synonymy: Aalbu et al. (2002: 487).

Brachyontis Casey, 1908: 82, 141. Type species: Coniontis globulina Casey, 1895, monotypy. Synonymy: Aalbu et al. (2002: 487).

Coniontis abdominalis LeConte, 1859USA (CA)

Coniontis abdominalis LeConte, 1859b: 77.

Coniontis strenua Casey, 1908: 84. Synonymy: Doyen (1977: 2).

Coniontis tristis Casey, 1908: 84. Synonymy: Doyen (1977: 2).

Coniontis gravis Casey, 1908: 85. Synonymy: Doyen (1977: 2).

Coniontis rugosa Casey, 1908: 85. Synonymy: Doyen (1977: 2).

Coniontis tenebrosa Casey, 1908: 86. Synonymy: Doyen (1977: 2).

Coniontis abdominalis caseyi Pierce, 1954c: 145. Synonymy: Doyen and Miller (1980: 3).

Coniontis abdominalis labreae Pierce, 1954c: 146. Synonymy: Doyen and Miller (1980: 3).

Coniontis abdominalis fragmans Pierce, 1954c: 148. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis alpha Pierce, 1954c: 148. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis asphalti Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis tristis latigula Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis blissi Pierce, 1954c: 149. Synonymy: Doyen and Miller (1980: 3).

Coniontis pectoralis paraelliptica Pierce, 1954c: 153. Synonymy: Doyen and Miller (1980: 3).

Coniontis pectoralis interrupta Pierce, 1954c: 154. Synonymy: Doyen and Miller (1980: 3).

Coniontis affinis LeConte, 1851USA (CA OR)

Coniontis affinis LeConte, 1851: 130.

Coniontis expansa Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis franciscana Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis truncata Casey, 1908: 120. Synonymy: Doyen (1977: 2).

Coniontis suturalis Casey, 1908: 121. Synonymy: Doyen (1977: 2).

Coniontis audax Casey, 1908: 121. Synonymy: Doyen (1977: 2).

Coniontis symmetrica Casey, 1908: 122. Synonymy: Doyen (1977: 2).

Coniontis convergens Casey, 1908: 122. Synonymy: Doyen (1977: 2).

Coniontis anxia Casey, 1908: 123. Synonymy: Doyen (1977: 2).

Coniontis affinis patruelis Casey, 1908: 123. Synonymy: Doyen (1977: 2).

Coniontis oregona Casey, 1908: 124. Synonymy: Doyen (1977: 2).

Coniontis pagana Casey, 1908: 130. Synonymy: Doyen (1977: 2).

Coniontis atronitens Casey, 1908USA (CA)

Coniontis atronitens Casey, 1908: 110.

Coniontis blaisdelli Casey, 1908USA (CA)

Coniontis blaisdelli Casey, 1908: 97.

Coniontis callida Casey, 1908USA (CA)

Coniontis callida Casey, 1908: 128.

Coniontis shastanica Casey, 1908: 128. Synonymy: Doyen (1977: 2).

Coniontis conferta Casey, 1908: 129. Synonymy: Doyen (1977: 2).

Coniontis agrestis Casey, 1908: 131. Synonymy: Doyen (1977: 2).

Coniontis congesta Casey, 1908: 131. Synonymy: Doyen (1977: 2).

Coniontis costulata Casey, 1908USA (CA)

Coniontis costulata Casey, 1908: 89.

Coniontis elliptica Casey, 1884USA (CA)

Coniontis elliptica Casey, 1884: 46.

Coniontis laevigata Casey, 1908: 88. Synonymy: Doyen (1977: 2).

Coniontis elliptica catalinae Casey, 1908: 88. Synonymy: Doyen (1977: 2).

Coniontis cuneata Casey, 1908: 111. Synonymy: Gebien (1938: 286).

Coniontis elongata Casey, 1890USA (CA)

Coniontis elongata Casey, 1890b: 380.

Coniontis rotundicollis Casey, 1908: 97. Synonymy: Doyen (1977: 3).

Coniontis innocua Casey, 1908: 99. Synonymy: Doyen (1977: 3).

Coniontis elongata limatula Casey, 1908: 99. Synonymy: Doyen (1977: 3).

Coniontis cylindrica Casey, 1908: 100. Synonymy: Doyen (1977: 3).

Coniontis obsidiana Casey, 1908: 100. Synonymy: Blaisdell (1935b: 120).

Coniontis longicollis Casey, 1908: 101. Synonymy: Doyen (1977: 3).

Coniontis eschscholtzii Mannerheim, 1840USA (CA)

Coniontis eschscholtzii Mannerheim, 1840: 138.

Coniontis extricata Casey, 1908USA (CA OR)

Coniontis extricata Casey, 1908: 124.

Coniontis marginata Casey, 1908: 125. Synonymy: Doyen (1977: 3).

Coniontis minuta Casey, 1908: 126. Synonymy: Doyen (1977: 3).

Coniontis parva Casey, 1908: 126. Synonymy: Doyen (1977: 3).

Coniontis perpolita Casey, 1908: 127. Synonymy: Doyen (1977: 3).

Coniontis pudica Casey, 1908: 127. Synonymy: Doyen (1977: 3).

Coniontis nemoralis slevini Blaisdell, 1924a: 86. Synonymy: Doyen (1977: 3).

Coniontis farallonica Casey, 1895USA (CA)

Coniontis farallonica Casey, 1895: 610.

Coniontis genitiva Casey, 1890USA (CA)

Coniontis genitiva Casey, 1890b: 385.

Coniontis verna Casey, 1908: 94. Synonymy: Doyen (1977: 3).

Coniontis opacicollis Casey, 1908: 101. Synonymy: Doyen (1977: 3).

Coniontis globulina Casey, 1895USA (CA)

Coniontis globulina Casey, 1895: 610.

Coniontis histrio Casey, 1908USA (AZ)

Coniontis histrio Casey, 1908: 91.

Coniontis hoppingi Blaisdell, 1918USA (CA)

Coniontis hoppingi Blaisdell, 1918a: 7.

Coniontis inaequalis Casey, 1890USA (CA)

Coniontis inaequalis Casey, 1890b: 375.

Coniontis inornata Casey, 1908USA (CA)

Coniontis inornata Casey, 1908: 130.

Coniontis integer Casey, 1908USA (CA)

Coniontis integer Casey, 1908: 87.

Coniontis keiferi (Blaisdell, 1943)MEX (BC)

Coniontides keiferi Blaisdell, 1943: 189.

Coniontis lamentabilis Blaisdell, 1924USA (CA)

Coniontis lamentabilis Blaisdell, 1924a: 85.

Coniontis lanei Boddy, 1957USA (ORWA)

Coniontis lanei Boddy, 1957: 191.

Coniontis lariversi Blaisdell, 1941USA (NV)

Coniontis lariversi Blaisdell, 1941c: 131.

Coniontis lassenica Casey, 1908USA (CA NV)

Coniontis lassenica Casey, 1908: 95.

Coniontis nevadensis Casey, 1908: 95. Synonymy: Doyen (1977: 3).

Coniontis nevadensis carsonica Casey, 1908: 95. Synonymy: Doyen (1977: 3).

Coniontis lata LeConte, 1866USA (CA)

Coniontis lata LeConte, 1866b: 113.

Coniontis lata var. insularis Casey, 1890b: 377. Synonymy: Doyen (1977: 3).

Coniontides finitimus Casey, 1908: 79. Synonymy: Doyen (1977: 3).

Coniontides clementinus Casey, 1908: 80. Synonymy: Blaisdell (1921b: 212).

Coniontis malkini (Boddy, 1957)USA (CA)

Coniontellus malkini Boddy, 1957: 188.

Coniontis microsticta Casey, 1908USA (CA)

Coniontis microsticta Casey, 1908: 107.

Coniontis inconspicua Casey, 1908: 108. Synonymy: Blaisdell (1924a: 84).

Coniontis muscula Blaisdell, 1918USA (CA)

Coniontis globulina var. muscula Blaisdell, 1918a: 9.

Coniontis nemoralis borealis Boddy, 1957USA (CA OR)

Coniontis nemoralis borealis Boddy, 1957: 192.

Coniontis nemoralis nemoralis Eschscholtz, 1829USA (CA)

Coniontis nemoralis Eschscholtz, 1829: 8.

Coniontis obesa LeConte, 1851USA (CA CO ID MT NV OR WY)

Coniontis obesus LeConte, 1851: 131.

Coniontellus inflatus Casey, 1890b: 389. Synonymy: Doyen (1977: 3).

Coniontellus subglaber Casey, 1890b: 389. Synonymy: Doyen (1977: 3).

Coniontellus hystrix Casey, 1908: 142. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus longipennis Casey, 1908: 143. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus ampliatus Casey, 1908: 144. Synonymy (with C. inflatus Casey): La Rivers (1947b: 214).

Coniontellus argutus Casey, 1908: 145. Synonymy: Doyen (1977: 3).

Coniontellus micans Casey, 1908: 145. Synonymy: Doyen (1977: 3).

Coniontis oblonga Casey, 1908USA (CA)

Coniontis oblonga Casey, 1908: 92.

Coniontis opaca Horn, 1870USA (CA)

Coniontis opaca Horn, 1870: 296.

Coniontis ancilla Casey, 1908: 91. Synonymy: Doyen (1977: 3).

Coniontis degener Casey, 1908: 93. Synonymy: Doyen (1977: 3).

Coniontis ovalis LeConte, 1851 [Fig. 13] CAN (BC) USA (AK CA CO ID MT NV OR UT WA)

Figure 13. 

Coniontis ovalis LeConte, 1851. Scale bar = 1 mm.

Coniontis ovalis LeConte, 1851: 131.

Coniontis alutacea Casey, 1890b: 383. Synonymy: Doyen (1977: 4).

Coniontis breviuscula Casey, 1908: 133. Synonymy: Boddy (1957: 190).

Coniontis sculptipennis Casey, 1908: 134. Synonymy: Boddy (1957: 190).

Coniontis regularis Casey, 1908: 134. Synonymy: Doyen (1977: 4).

Coniontis punctata Casey, 1908: 135. Synonymy (with C. regularis Casey): Boddy (1965: 141).

Coniontis parilis Casey, 1908: 135. Synonymy: Boddy (1957: 190).

Coniontis vancouveri Casey, 1908: 136. Synonymy: Boddy (1957: 190).

Coniontis uteana Casey, 1908: 136. Synonymy: Doyen (1977: 4).

Coniontis inepta Casey, 1908: 137. Synonymy: Doyen (1977: 4).

Coniontis oblita Casey, 1908: 137. Synonymy: Doyen (1977: 4).

Coniontis arida Casey, 1908: 138. Synonymy: Doyen (1977: 4).

Coniontis weidti Casey, 1908: 138. Synonymy: Doyen (1977: 4).

Coniontis acerba Casey, 1908: 139. Synonymy: Doyen (1977: 4).

Coniontis anita Casey, 1908: 139. Synonymy: Doyen (1977: 4).

Coniontis corvina Casey, 1908: 140. Synonymy: Doyen (1977: 4).

Coniontis ovalis okanagani Boddy, 1957: 190. Synonymy: Doyen (1977: 4).

Coniontis pallidicornis Casey, 1890USA (CA)

Coniontis pallidicornis Casey, 1890b: 385.

Coniontis obsolescens Casey, 1908: 92. Synonymy: Doyen (1977: 4).

Coniontis parallela Casey, 1890USA (CA)

Coniontis parallela Casey, 1890b: 386.

Coniontis parviceps Casey, 1890USA (CA) MEX (BC)

Coniontis parviceps Casey, 1890b: 387.

Coniontis filiola Casey, 1908: 115. Synonymy: Doyen (1977: 4).

Coniontis pectoralis Casey, 1908USA (CA)

Coniontis pectoralis Casey, 1908: 86.

Coniontis levettei Casey, 1908: 87. Synonymy: Doyen (1977: 4).

Coniontis picescens Casey, 1908: 87. Synonymy: Doyen (1977: 4).

Coniontis perspicua Casey, 1908USA (CA)

Coniontis perspicua Casey, 1908: 114.

Coniontis proba Casey, 1908USA (ID OR)

Coniontis proba Casey, 1908: 105.

Coniontis puncticollis LeConte, 1851USA (CA)

Coniontis puncticollis LeConte, 1851: 131.

Coniontis exigua Casey, 1908: 106. Synonymy: Doyen (1977: 4).

Coniontis paupercula Casey, 1908: 106. Synonymy: Doyen (1977: 4).

Coniontis inflexula Casey, 1908: 107. Synonymy (with C. exigua Casey): Casey (1911: 253).

Coniontis picipes Casey, 1908: 107. Synonymy: Doyen (1977: 4).

Coniontis punctipes Casey, 1890USA (CA)

Coniontis punctipes Casey, 1890b: 380.

Coniontis punctulata (Horn, 1876)USA (CA) MEX

Coelotaxis punctulata Horn, 1876a: 201.

Coelotaxis muricata Horn, 1876a: 201. Synonymy: Doyen (1977: 4).

Coelotaxis angustula Casey, 1890a: 177. Synonymy: Doyen (1977: 4).

Coelotaxis densa Casey, 1908: 149. Synonymy: Doyen (1977: 4).

Coelotaxis frontalis Casey, 1908: 149. Synonymy: Doyen (1977: 4).

Coniontis rainieri Boddy, 1957USA (WA)

Coniontis rainieri Boddy, 1957: 191.

Coniontis remnans Pierce, 1954USA (CA)18

Coniontis remnans Pierce, 1954c: 155.

Coniontis robusta Horn, 1870USA (CA)

Coniontis robusta Horn, 1870: 296.

Coniontis luctuosa Casey, 1908: 89. Synonymy: Doyen (1977: 5).

Coniontis sanfordii Blaisdell, 1895USA (CA)

Coniontis sanfordii Blaisdell, 1895: 235.

Coniontis santarosae Blaisdell, 1921USA (CA)

Coniontis santarosae Blaisdell, 1921b: 209.

Coniontis setosa Casey, 1890USA (ID NV OR UT WA)

Coniontis setosus Casey, 1890b: 387.

Coniontis obtusa Casey, 1908: 116. Synonymy: Doyen (1977: 5).

Coniontis wickhami Casey, 1908: 117. Synonymy: Doyen (1977: 5).

Coniontis lanuginosa Casey, 1908: 117. Synonymy: Doyen (1977: 5).

Coniontis pubifera Casey, 1908: 118. Synonymy: Boddy (1965: 140).

Coniontis sparsa Casey, 1908USA (CA)

Coniontis sparsa Casey, 1908: 110.

Coniontis subpubescens LeConte, 1851USA (CA OR)

Coniontis subpubescens LeConte, 1851: 131.

Coniontis montana Casey, 1890b: 384. Synonymy: Doyen (1977: 5).

Coniontis canonica Casey, 1908: 114. Synonymy: Blaisdell (1918a: 13).

Coniontis tenuis Casey, 1908USA (CA)

Coniontis tenuis Casey, 1908: 141.

Coniontis thoracica Casey, 1908USA (CA)

Coniontis thoracica Casey, 1908: 104.

Coniontis timida Casey, 1908USA (CA)

Coniontis timida Casey, 1908: 102.

Coniontis conicicollis Casey, 1908: 102. Synonymy: Doyen (1977: 5).

Coniontis lucidula Casey, 1908: 103. Synonymy: Doyen (1977: 5).

Coniontis protensa Casey, 1908: 104. Synonymy: Doyen (1977: 5).

Coniontis vandykei Blaisdell, 1921USA (CA)

Coniontides vandykei Blaisdell, 1921b: 212.

Coniontis ventura Blaisdell, 1924USA (CA)

Coniontis globulina ventura Blaisdell, 1924a: 83.

Coniontis viatica Eschscholtz, 1829USA (CA)

Coniontis viatica Eschscholtz, 1829: 7.

Coniontis wadei Casey, 1924USA (WA)

Coniontis wadei Casey, 1924: 313.

Genus Conisattus Casey, 1895 [M]

Conisattus Casey, 1895: 614. Type species: Conisattus rectus Casey, 1895, monotypy.

Conisattus rectus Casey, 1895USA (OR WA)

Conisattus rectus Casey, 1895: 614.

Conisattus nelsoni Boddy, 1957: 188. Synonymy: Doyen (1984b: 26).

Genus Eusattus LeConte, 1851 [M]

Eusattus LeConte, 1851: 131. Type species: Eusattus difficilis LeConte, 1851, subsequent designation (Casey 1908: 56).

Discodemus LeConte, 1862a: 223. Type species: Zophosis reticulata Say, 1824, monotypy. Synonymy: LeConte (1866a: 60).

Conipinus LeConte, 1862a: 223. Type species: Eusattus dubius LeConte, 1851, subsequent designation (Gebien 1938: 284). Synonymy: LeConte (1866a: 60).

Nesostes Casey, 1908: 56, 58. Type species: Eusattus robustus LeConte, 1866, original designation. Synonymy: Triplehorn (1968b: 379).

Megasattus Casey, 1908: 56. Type species: Eusattus erosus Horn, 1870, original designation. Synonymy: Triplehorn (1968b: 379).

Eusattodes Casey, 1908: 56. Type species: Eusattus laevis LeConte, 1866, original designation. Synonymy: Triplehorn (1968b: 379).

Sphaeriontis Casey, 1908: 56, 75. Type species: Eusattus muricatus LeConte, 1851, original designation. Synonymy: La Rivers (1949: 180).

Coelosattus Blaisdell, 1927: 166. Type species: Coelosattus fortineri Blaisdell, 1927 (= Eusattus dilatatus LeConte, 1851), monotypy. Synonymy: Doyen (1972: 373).

Eusattus araneosus (Blaisdell, 1923)MEX (BS)

Megasattus araneosus Blaisdell, 1923: 266.

Eusattus arenarius Doyen, 1984MEX (BC BS)

Eusattus arenarius Doyen, 1984b: 75.

Eusattus aridus Doyen, 1984MEX (BC BS)

Eusattus aridus Doyen, 1984b: 43.

Eusattus catalinensis Doyen, 1984MEX (BS)

Eusattus catalinensis Doyen, 1984b: 45.

Eusattus catavinus Doyen, 1984MEX (BC)

Eusattus catavinus Doyen, 1984b: 46.

Eusattus cedrosensis Doyen, 1984MEX (BC)

Eusattus cedrosensis Doyen, 1984b: 47.

Eusattus ceralboensis Doyen, 1984MEX (BS)

Eusattus ceralboensis Doyen, 1984b: 48.

Eusattus cienegus Doyen, 1984MEX (CO)

Eusattus cienegus Doyen, 1984b: 49.

Eusattus ciliatoides Doyen, 1984MEX (BC)

Eusattus ciliatoides Doyen, 1984b: 75.

Eusattus ciliatus Horn, 1894MEX (BC)

Eusattus ciliatus Horn, 1894b: 422.

Eusattus convexus LeConte, 1851USA (AZ CO KS MO NM OK TX UT WY) MEX (CH DU SO)

Eusattus convexus LeConte, 1851: 132.

Eusattus sculptus Champion, 1892: 510. Synonymy: Doyen (1984b: 35).

Eusattus rotundus Casey, 1908: 72. Synonymy: Doyen (1977: 5).

Eusattus turgidus Casey, 1908: 73. Synonymy: Doyen (1977: 5).

Eusattus subnitens Casey, 1908: 73. Synonymy: Doyen (1984b: 35).

Eusattus peropacus Casey, 1908: 74. Synonymy: Doyen (1977: 5).

Eusattus acutus Casey, 1908: 74. Synonymy: Doyen (1977: 5).

Eusattus quadratus Casey, 1924: 311. Synonymy: Doyen (1977: 5).

Eusattus subvelutinus Casey, 1924: 312. Synonymy: Doyen (1977: 5).

Eusattus woodgatei Casey, 1924: 312. Synonymy: Doyen (1977: 5).

Eusattus costatus Horn, 1870USA (CA) MEX (BC BS)

Eusattus costatus Horn, 1870: 293.

Megasattus sternalis Blaisdell, 1923: 268. Synonymy: Doyen (1984b: 50).

Eusattus crypticus Doyen, 1984MEX (BS)

Eusattus crypticus Doyen, 1984b: 90.

Eusattus depressus Champion, 1884MEX (CH NA SI SO)

Eusattus depressus Champion, 1884: 75.

Eusattus puncticeps Blaisdell, 1923: 269. Synonymy: Doyen (1984b: 87).

Eusattus difficilis LeConte, 1851USA (CA NV) MEX (BC SO)

Eusattus difficilis LeConte, 1851: 132.

Eusattus coquilletti Linell, 1899: 180. Synonymy: Doyen (1977: 5).

Eusattus agnatus Casey, 1908: 70. Synonymy: Doyen (1977: 5).

Eusattus compositus Casey, 1908: 71. Synonymy: Doyen (1977: 5).

Eusattus congener Casey, 1908: 71. Synonymy: Doyen (1977: 5).

Eusattus acutangulus Casey, 1908: 72. Synonymy: Doyen (1977: 5).

Eusattus dilatatus LeConte, 1851USA (AZ CA) MEX (SO)

Eusattus dilatatus LeConte, 1851: 132.

Coelosattus fortineri Blaisdell, 1927: 167. Synonymy: Doyen (1977: 5).

Eusattus dubius abditus Doyen, 1984MEX (BC)

Eusattus dubius abditus Doyen, 1984b: 82.

Eusattus dubius arizonensis Doyen, 1984USA (AZ CA NV)

Eusattus dubius arizonensis Doyen, 1984b: 81.

Eusattus dubius dubius LeConte, 1851USA (AZ CA NV UT)

Eusattus dubius LeConte, 1851: 132.

Eusattus nanus Casey, 1895: 613. Synonymy: Doyen (1977: 5).

Eusattus oblongulus Casey, 1908: 67. Synonymy: Doyen (1977: 5).

Conipinus spaldingi Casey, 1924: 313. Synonymy: Doyen (1977: 5).

Eusattus dubius setosus Doyen, 1984MEX (BS)

Eusattus dubius setosus Doyen, 1984b: 82.

Eusattus erosus erosus Horn, 1870MEX (BS)

Eusattus erosus Horn, 1870: 294.

Eusattus erosus laeviventris (Blaisdell, 1923)MEX (BS)

Megasattus laeviventris Blaisdell, 1923: 267.

Eusattus erosus manuelis (Blaisdell, 1923)MEX (BS)

Megasattus erosus manuelis Blaisdell, 1923: 266.

Eusattus franciscanus Doyen, 1984MEX (BS)

Eusattus franciscanus Doyen, 1984b: 54.

Eusattus hirsutus Doyen, 1984USA (NV)

Eusattus hirsutus Doyen, 1984b: 66.

Eusattus laevis LeConte, 1866MEX (BS)

Eusattus laevis LeConte, 1866b: 113.

Eusattus mexicanus Champion, 1892MEX (CO GE JA)

Eusattus mexicanus Champion, 1892: 510.

Eusattus minimus Doyen, 1984MEX (NL)

Eusattus minimus Doyen, 1984b: 38.

Eusattus muricatus diabloensis Doyen, 1984MEX (BC)

Eusattus muricatus diabloensis Doyen, 1984b: 71.

Eusattus muricatus muricatus LeConte, 1851USA (AZ CA CO ID NM NV OR TX UT WA)

Eusattus muricatus LeConte, 1851: 132.

Sphaeriontis acomana Casey, 1908: 76. Synonymy: La Rivers (1949: 180).

Sphaeriontis latissima Casey, 1924: 310. Synonymy: La Rivers (1949: 180).

Sphaeriontis fulvescens Casey, 1924: 310. Synonymy: La Rivers (1949: 180).

Eusattus nitidipennis LeConte, 1851MEX (CO GU NL PU VE ZA)

Eusattus nitidipennis LeConte, 1851: 133.

Eusattus brevis Champion, 1884: 75. Synonymy: Doyen (1977: 6).

Eusattus obliteratus Champion, 1892MEX (CO DU)

Eusattus obliteratus Champion, 1892: 51019.

Eusattus pallidus adustus Doyen, 1984MEX (BS)

Eusattus pallidus adustus Doyen, 1984b: 84.

Eusattus pallidus immaculatus Doyen, 1984MEX (BS)

Eusattus pallidus immaculatus Doyen, 1984b: 85.

Eusattus pallidus pallidus Doyen, 1984MEX (BS)

Eusattus pallidus pallidus Doyen, 1984b: 84.

Eusattus phreatophilus Doyen, 1984USA (CA NV)

Eusattus phreatophilus Doyen, 1984b: 71.

Eusattus planulus Doyen, 1984MEX (BS)

Eusattus planulus Doyen, 1984b: 57.

Eusattus politus cruzensis Doyen, 1984USA (CA)

Eusattus politus cruzensis Doyen, 1984b: 93.

Eusattus politus politus Horn, 1883USA (CA)

Eusattus politus Horn, 1883: 304.

Eusattus vanduzeei Blaisdell, 1921b: 214. Synonymy: Doyen (1984b: 93).

Eusattus pons Triplehorn, 1968USA (TX) MEX (CH CO DU)

Eusattus pons Triplehorn, 1968b: 376.

Eusattus productus LeConte, 1858USA (AZ CA) MEX (BC SO)

Eusattus productus LeConte, 1858a: 20.

Eusattus explanatus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus vicinus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus lobatus Casey, 1908: 68. Synonymy: Doyen (1977: 6).

Eusattus puberulus LeConte, 1854USA (TX)

Eusattus puberulus LeConte, 1854a: 84.

Eusattus reticulatus (Say, 1824)USA (AZ CO NM OK TX UT) MEX (CH SO)

Tenebrio gibbus DeGeer, 1778: 652 [junior primary homonym of Tenebrio gibbus Linnaeus, 1767].

Tenebrio striatus Retzius, 1783: 134 [junior primary homonym of Tenebrio striatus Müller, 1776]. Replacement name for Tenebrio gibbus DeGeer, 1778.

Zophosis reticulata Say, 1824a: 250. Synonymy: Ferrer and Holston (2011: 252).

Discodemus corrosus Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus brevipennis Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus elongatulus Casey, 1908: 61. Synonymy: Doyen (1977: 6).

Discodemus depressulus Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Discodemus subsericeus Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Discodemus knausi Casey, 1908: 62. Synonymy: Doyen (1977: 6).

Eusattus robustus LeConte, 1866USA (CA)

Eusattus robustus LeConte, 1866b: 112.

Eusattus robustus postremus Casey, 1908: 59. Synonymy: Doyen (1977: 6).

Eusattus rudei Doyen, 1984MEX (BS)

Eusattus rudei Doyen, 1984b: 91.

Eusattus secutus Horn, 1894MEX (BS)

Eusattus secutus Horn, 1894b: 421.

Eusattus venosus Champion, 1892MEX (CO JA NA)

Eusattus venosus Champion, 1892: 509.

Eusattus vizcainensis Doyen, 1984MEX (BC BS)

Eusattus vizcainensis Doyen, 1984b: 85.

Tribe Cryptoglossini LeConte, 1862

Centrioptérides Lacordaire, 1859: 134 [nomen oblitum, see Aalbu 2006: 57]. Type genus: Centrioptera Mannerheim, 1843.

Cryptoglossini LeConte, 1862a: 220 [nomen protectum]. Type genus: Cryptoglossa Solier, 1837.

Genus Asbolus LeConte, 1851 [M]

Asbolus LeConte, 1851: 129. Type species: Asbolus verrucosus LeConte, 1851, subsequent designation (Aalbu 2005: 721).

Asbolus laevis LeConte, 1851USA (AZ CA) MEX (BC SO)

Asbolus laevis LeConte, 1851: 130.

Cryptoglossa laevis subsimilis Casey, 1924: 308. Synonymy: Aalbu (2005: 730).

Asbolus mexicanus angularis (Horn, 1894)USA (AZ CA) MEX (BC BS)

Centrioptera angularis Horn, 1894b: 414.

Asbolus mexicanus mexicanus (Champion, 1884)USA (NM TX) MEX (CH CO DU NL)

Cryptoglossa mexicana Champion, 1884: 73.

Cryptoglossa granulifera Champion, 1892: 508. Synonymy: Aalbu (2005: 722).

Asbolus papillosus (Triplehorn, 1964)USA (CA) MEX (SO)

Cryptoglossa laevis papillosa Triplehorn, 1964a: 48.

Asbolus verrucosus LeConte, 1851USA (AZ CA NM NV UT) MEX (BC SO)

Asbolus verrucosus LeConte, 1851: 129.

Cryptoglossa verrucosa carinulatus Blaisdell, 1945: 25. Synonymy: Aalbu (2005: 724).

Genus Cryptoglossa Solier, 1837 [F]

Cryptoglossa Solier, 1837: 680. Type species: Cryptoglossa bicostata Solier, 1837, monotypy.

Centrioptera Mannerheim, 1843: 279. Type species: Centrioptera caraboides Mannerheim, 1843, monotypy. Synonymy: Aalbu et al. (2002: 486).

Oochila LeConte, 1862a: 220. Type species: Asbolus infaustus LeConte, 1854, original designation. Synonymy (with Centrioptera Mannerheim): Horn (1870: 278).

Amblycyphus Motschulsky, 1870: 401. Type species: Amblycyphus asperatus Motschulsky, 1870 (= Centrioptera pectoralis Blaisdell, 1921), monotypy. Synonymy: Aalbu et al. (1995: 483).

Cryptoglossa asperata (Horn, 1870)MEX (BS)

Centrioptera asperata Horn, 1870: 279.

Centrioptera asperata discreta Blaisdell, 1923: 249. Synonymy: Aalbu (2005: 709).

Centrioptera asperata subornata Blaisdell, 1923: 249. Synonymy: Aalbu (2005: 709).

Centrioptera asperata planata Blaisdell, 1923: 250. Synonymy: Aalbu (2005: 709).

Cryptoglossa bicostata Solier, 1837MEX (OA PU)

Cryptoglossa bicostata Solier, 1837: 681.

Cryptoglossa caraboides (Mannerheim, 1843)MEX (GE MO PU)

Centrioptera caraboides Mannerheim, 1843: 280.

Cryptoglossa infausta (LeConte, 1854)USA (TX) MEX (CO DU TA)

Asbolus infaustus LeConte, 1854a: 84.

Centrioptera spiculosa Champion, 1892: 508. Synonymy: Champion (1893a: 572).

Centrioptera texana Blaisdell, 1924b: 88. Synonymy: Aalbu (2005: 705).

Cryptoglossa michelbacheri (Blaisdell, 1943)MEX (BS)

Centrioptera michelbacheri Blaisdell, 1943: 222.

Cryptoglossa muricata (LeConte, 1851)USA (AZ CA NV UT) MEX (BC SO)

Centrioptera muricata LeConte, 1851: 142.

Centrioptera utensis Casey, 1907: 513. Synonymy: Aalbu (2005: 712).

Centrioptera sculptiventris Blaisdell, 1923: 247. Synonymy: Aalbu (2005: 712).

Centrioptera serrata Casey, 1924: 306. Synonymy: Aalbu (2005: 712).

Centrioptera elongata Casey, 1924: 306. Synonymy: Aalbu (2005: 712).

Cryptoglossa seriata cerralvoensis Aalbu, 2005MEX (BS)

Cryptoglossa seriata cerralvoensis Aalbu, 2005: 721.

Cryptoglossa seriata seriata LeConte, 1861USA (AZ CA) MEX (BS)

Cryptoglossa seriata LeConte, 1861a: 337.

Cryptoglossa spiculifera pectoralis (Blaisdell, 1921)USA (CA) MEX (BC BS)

Amblycyphus asperatus Motschulsky, 1870: 404 [junior secondary homonym of Cryptoglossa asperata (Horn, 1870)20].

Centrioptera pectoralis Blaisdell, 1921b: 198. Synonymy: Aalbu et al. (1995: 483).

Centrioptera dulzurae Blaisdell, 1921b: 199. Synonymy: Aalbu et al. (1995: 483).

Centrioptera chamberlini Blaisdell, 1923: 246. Synonymy: Aalbu (2005: 717).

Cryptoglossa spiculifera spiculifera (LeConte, 1861)MEX (BS)

Centrioptera spiculifera LeConte, 1861a: 337.

Cryptoglossa variolosa (Horn, 1870)USA (AZ CA NM) MEX (SI SO)

Centrioptera variolosa Horn, 1870: 280.

Genus Schizillus Horn, 1874 [M]

Schizillus Horn, 1874a: 33. Type species: Schizillus laticeps Horn, 1874, monotypy.

Schizillus laticeps Horn, 1874USA (AZ CA NV UT) MEX (BC)

Schizillus laticeps Horn, 1874a: 33.

Schizillus convexus Blaisdell, 1921b: 203. Synonymy: Aalbu (2005: 732).

Schizillus lomae Blaisdell, 1921b: 206. Synonymy: Aalbu (2005: 732).

Schizillus opacus Casey, 1924: 307. Synonymy: Aalbu (2005: 732).

Schizillus nunenmacheri Blaisdell, 1921USA (AZ CA NV UT)

Schizillus nunenmacheri Blaisdell, 1921b: 204.

Schizillus beali Parker, 1955: 148. Synonymy: Aalbu (2005: 734).

Tribe Edrotini Lacordaire, 1859

Édrotides Lacordaire, 1859: 31. Type genus: Edrotes LeConte, 1851.

Triorophi LeConte and Horn, 1883: 362. Type genus: Triorophus LeConte, 1851.

Auchmobii LeConte and Horn, 1883: 362. Type genus: Auchmobius LeConte, 1851.

Trimytini Casey, 1907: 278. Type genus: Trimytis Leconte, 1851.

Eurymetoponini Casey, 1907: 278. Type genus: Eurymetopon Eschscholtz, 1831.

Trientomini Casey, 1907: 278. Type genus: Trientoma Solier, 1835.

Genus Armalia Casey, 1907 [F]

Armalia Casey, 1907: 289, 330. Type species: Emmenastus texanus LeConte, 1866, original designation.

Armalia alata (Champion, 1884) GUA NIC

Emmenastus alatus Champion, 1884: 13.

Emmenastus salvini Champion, 1884: 13. Synonymy: Champion (1892: 482).

Armalia angularis Casey, 1907USA (TX)

Armalia angularis Casey, 1907: 331.

Armalia belti (Champion, 1884)MEX (YU) GUAHONNIC

Emmenastus belti Champion, 1884: 11.

Emmenastus rotundicollis Champion, 1884: 11. Synonymy: Champion (1892: 480).

Emmenastus intermedius Champion, 1884: 12. Synonymy: Champion (1892: 480).

Armalia brevipennis (Champion, 1884)MEX (NA)

Emmenastus brevipennis Champion, 1884: 10.

Armalia canaliculata (Champion, 1884)MEX (SI)

Emmenastus canaliculatus Champion, 1884: 10.

Armalia chiriquensis (Champion, 1884)PAN / SA

Emmenastus chiriquensis Champion, 1884: 9.

Armalia longicornis (Champion, 1884) GUA

Emmenastus longicornis Champion, 1884: 9.

Armalia solitaria (Champion, 1884)MEX (OA)

Emmenastus solitarius Champion, 1884: 11.

Armalia texana (LeConte, 1866)USA (TX)

Emmenastus texanus LeConte, 1866b: 108.

Armalia variabilis (Champion, 1884)MEX (VE) HON

Emmenastus variabilis Champion, 1884: 10.

Genus Auchmobius LeConte, 1851 [M]

Auchmobius LeConte, 1851: 139. Type species: Auchmobius sublaevis LeConte, 1851, monotypy.

Auchmobius angelicus Blaisdell, 1934USA (CA)

Auchmobius angelicus Blaisdell, 1934b: 249.

Auchmobius parvicollis Blaisdell, 1934USA (CA)

Auchmobius parvicollis Blaisdell, 1934b: 246.

Auchmobius picipes Blaisdell, 1934USA (CA)

Auchmobius picipes Blaisdell, 1934b: 252.

Auchmobius sanfordi Blaisdell, 1934USA (CA)

Auchmobius sanfordi Blaisdell, 1934b: 257.

Auchmobius slevini Blaisdell, 1934USA (CA)

Auchmobius slevini Blaisdell, 1934b: 243.

Auchmobius subboreus Blaisdell, 1934USA (CA NV)

Auchmobius subboreus Blaisdell, 1934b: 254.

Auchmobius sublaevis LeConte, 1851USA (CA)

Auchmobius sublaevis LeConte, 1851: 140.

Auchmobius subovalis Blaisdell, 1934USA (CA)

Auchmobius subovalis Blaisdell, 1934b: 238.

Genus Chilometopon Horn, 1874 [N]

Chilometopon Horn, 1874a: 31. Type species: Trimytis abnormis Horn, 1870, subsequent designation (Casey 1907: 367).

Prometopion Casey, 1907: 366, 370. Type species: Prometopion amplipenne Casey, 1907 (= Chilometopon helopioides Horn, 1874), original designation. Synonymy: MacLachlan and Olson (1990: 72).

Chilometopon abnorme (Horn, 1870)USA (AZ CA NV OR UT) MEX (BC SO)

Trimytis abnormis Horn, 1870: 261.

Chilometopon castaneum Casey, 1907: 373. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon brevipenne Casey, 1907: 374. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon ensifer Casey, 1907: 374. Synonymy: MacLachlan and Olson (1990: 74).

Chilometopon brachystomum Doyen, 1983USA (AZ CA NV) MEX (BC)

Chilometopon brachystomum Doyen, 1983: 81.

Chilometopon cribricolle Blaisdell, 1923MEX (BS)

Chilometopon cribricolle Blaisdell, 1923: 230.

Chilometopon helopioides Horn, 1874USA (AZ CA ID NM NV UT) MEX (BC)

Chilometopon helopioides Horn, 1874a: 31.

Prometopion amplipenne Casey, 1907: 372. Synonymy: MacLachlan and Olson (1990: 78).

Chilometopon microps MacLachlan and Olson, 1990USA (CA)

Chilometopon microps MacLachlan and Olson, 1990: 76.

Chilometopon pallidum Casey, 1890USA (AZ CA NM NV TX UT) MEX (BC CH)

Chilometopon pallidum Casey, 1890b: 367.

Chilometopon rugiceps Blaisdell, 1923MEX (BC)

Chilometopon rugiceps Blaisdell, 1923: 229.

Genus Cryptadius LeConte, 1851 [M]

Cryptadius LeConte, 1851: 140. Type species: Cryptadius inflatus LeConte, 1851, monotypy.

Cryptadius inflatus blaisdelli Thomas, 1985MEX (BS)

Cryptadius inflatus blaisdelli Thomas, 1985: 197.

Cryptadius inflatus inflatus LeConte, 1851USA (CA) MEX (BC)

Cryptadius inflatus LeConte, 1851: 140.

Cryptadius oviformis Casey, 1907: 328. Synonymy: Thomas (1985: 196).

Cryptadius punctipennis Casey, 1907: 328. Synonymy: Thomas (1985: 196).

Cryptadius curvipes Casey, 1907: 329. Synonymy: Thomas (1985: 196).

Cryptadius sonorae Berry, 1974MEX (BS SO)

Cryptadius sonorae Berry, 1974: 175.

Cryptadius tarsalis Blaisdell, 1923MEX (BC BS SO)

Cryptadius tarsalis Blaisdell, 1923: 212.

Cryptadius angulatus Blaisdell, 1923: 210. Synonymy: Thomas (1985: 198).

Cryptadius sinuatus Blaisdell, 1923: 211. Synonymy: Thomas (1985: 198).

Cryptadius andrewsi Berry, 1977: 561. Synonymy: Thomas (1985: 198).

Genus Ditaphronotus Casey, 1907 [M]

Ditaphronotus Casey, 1907: 341. Type species: Emmenastus foveicollis Champion, 1884, original designation.

Ditaphronotus championi Casey, 1907 NIC

Ditaphronotus championi Casey, 1907: 342.

Ditaphronotus confusus (Champion, 1884)MEX (CI) GUA

Emmenastus confusus Champion, 1884: 15.

Ditaphronotus foveicollis (Champion, 1884) GUA NIC CRI

Emmenastus foveicollis Champion, 1884: 14.

Ditaphronotus laevicollis (Champion, 1884) PAN

Emmenastus laevicollis Champion, 1884: 15.

Genus Edrotes LeConte, 1851 [M]

Edrotes LeConte, 1851: 140. Type species: Edrotes ventricosus LeConte, 1851, monotypy.

Hedrotes Gemminger [in Gemminger and Harold], 1870: 1816. Unjustified emendation of Edrotes LeConte, 1851, not in prevailing usage.

Subgenus Edrotes LeConte, 1851

Edrotes LeConte, 1851: 140. Type species: Edrotes ventricosus LeConte, 1851, monotypy.

Edrotes fossor Triplehorn, 1972MEX (BS)

Edrotes fossor Triplehorn, 1972: 27.

Edrotes leechi Doyen, 1968USA (AZ CO UT)

Edrotes leechi Doyen, 1968: 218.

Edrotes rotundus (Say, 1824)USA (AZ CO NM TX UT) MEX (BC CH)

Pimelia rotunda Say, 1824a: 251.

Edrotes globosus Casey, 1890a: 175. Synonymy: La Rivers (1947a: 325).

Edrotes inflatus Casey, 1907: 454. Synonymy: La Rivers (1947a: 325).

Edrotes puncticeps Casey, 1907: 454. Synonymy: La Rivers (1947a: 325).

Edrotes intermixtus Casey, 1907: 455. Synonymy: La Rivers (1947a: 325).

Edrotes oblongulus Casey, 1907: 455. Synonymy: La Rivers (1947a: 325).

Edrotes lineatus Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes subaequalis Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes angustulus Casey, 1907: 456. Synonymy: La Rivers (1947a: 325).

Edrotes desertus Blaisdell, 1943: 212. Synonymy: La Rivers (1947a: 325).

Edrotes ventricosus LeConte, 1851USA (AZ CA ID NV OR) MEX (BC BS SO)

Edrotes ventricosus LeConte, 1851: 141.

Edrotes nitidus Casey, 1890a: 175. Synonymy: La Rivers (1947a: 321).

Edrotes orbus Casey, 1907: 452. Synonymy: La Rivers (1947a: 321).

Edrotes angusticollis Casey, 1907: 452. Synonymy: La Rivers (1947a: 321).

Edrotes longipennis Casey, 1907: 453. Synonymy: La Rivers (1947a: 321).

Edrotes mexicanus Blaisdell, 1923: 241. Synonymy: La Rivers (1947a: 321).

Edrotes asperatus Blaisdell, 1923: 241. Synonymy: La Rivers (1947a: 321).

Edrotes laticollis Casey, 1924: 300. Synonymy: La Rivers (1947a: 321).

Edrotes longicornis Casey, 1924: 300. Synonymy: La Rivers (1947a: 321).

Edrotes variipilis Casey, 1924: 301. Synonymy: La Rivers (1947a: 321).

Edrotes barrowsi Dajoz, 1999: 320. New synonymy [RLA].

Subgenus Odrotes La Rivers, 1947

Odrotes La Rivers, 1947a: 320. Type species: Edrotes arens La Rivers, 1947, monotypy.

Edrotes arens La Rivers, 1947USA (AZ CA) MEX (BC)

Edrotes arens La Rivers, 1947a: 320.

Genus Emmenastrichus Horn, 1894 [M]

Emmenastrichus Horn, 1894b: 413. Type species: Emmenastrichus cribratus Horn, 1894, subsequent designation (Casey 1907: 289).

Emmenastrichus cribratus Horn, 1894MEX (BS)

Emmenastrichus cribratus Horn, 1894b: 413.

Emmenastrichus erosus Horn, 1894MEX (BS)

Emmenastrichus erosus Horn, 1894b: 414.

Genus Emmenides Casey, 1907 [M]

Emmenides Casey, 1907: 329. Type species: Emmenastus punctatus LeConte, 1866, original designation.

Emmenides apicalis Blaisdell, 1923MEX (BS)

Emmenides apicalis Blaisdell, 1923: 215.

Emmenides catalinae Blaisdell, 1923MEX (BS)

Emmenides catalinae Blaisdell, 1923: 216.

Emmenides igualensis (Champion, 1892)MEX (GE)

Emmenastus igualensis Champion, 1892: 484.

Emmenides obsoletus Blaisdell, 1923MEX (BS)

Emmenides obsoletus Blaisdell, 1923: 216.

Emmenides punctatus (LeConte, 1866)USA (AZ TX) MEX (BS)

Emmenastus punctatus LeConte, 1866b: 106.

Emmenides subdescalceatus Blaisdell, 1923MEX (BS)

Emmenides subdescalceatus Blaisdell, 1923: 213.

Genus Eremocantor Smith and Wirth, 2016 [M]

Eremocantor Smith and Wirth, 2016: 582. Type species: Eremocantor marioni Smith and Wirth, 2016, original designation.

Eremocantor marioni Smith and Wirth, 2016USA (TX)

Eremocantor marioni Smith and Wirth, 2016: 583.

Genus Eschatomoxys Blaisdell, 1935 [M]

Eschatomoxys Blaisdell, 1935d: 125. Type species: Eschatomoxys wagneri Blaisdell, 1935, original designation.

Eschatomoxys andrewsi Aalbu and Thomas, 2008USA (CA)

Eschatomoxys andrewsi Aalbu and Thomas [in Pape et al.], 2008: 529.

Eschatomoxys paco Aalbu and Thomas, 2008MEX (BC)

Eschatomoxys paco Aalbu and Thomas [in Pape et al.], 2008: 527.

Eschatomoxys pholeter Thomas and Pape, 2008USA (AZ)

Eschatomoxys pholeter Thomas and Pape [in Pape et al.], 2008: 525.

Eschatomoxys rosei Aalbu and Thomas, 2008MEX (BC)

Eschatomoxys rosei Aalbu and Thomas [in Pape et al.], 2008: 530.

Eschatomoxys tanneri Sorenson and Stones, 1959USA (AZ UT)

Eschatomoxys tanneri Sorenson and Stones, 1959: 63.

Eschatomoxys wagneri Blaisdell, 1935USA (AZ CA)

Eschatomoxys wagneri Blaisdell, 1935d: 125.

Genus Eurymetopon Eschscholtz, 1831 [N]

Eurymetopon Eschscholtz, 1831: 5, 8. Type species: Eurymetopon rufipes Eschscholtz, 1831, subsequent designation (Casey 1907: 288).

Eurymetopon ochraceum Eschscholtz, 1831USA (CA)

Eurymetopon ochraceum Eschscholtz, 1831: 8.

Eurymetopon rufipes Eschscholtz, 1831USA (AZ CA) MEX (BS SO)

Eurymetopon rufipes Eschscholtz, 1831: 8.

Genus Garridoa Marcuzzi, 1985 [F]

Garridoa Marcuzzi, 1985: 180. Type species: Garridoa kaszabi Marcuzzi, 1985, monotypy.

Garridoa kaszabi Marcuzzi, 1985 CUB

Garridoa kaszabi Marcuzzi, 1985: 180.

Genus Hylocrinus Casey, 1907 [M]

Hylocrinus Casey, 1907: 289, 331. Type species: Eurymetopon longulum LeConte, 1851, original designation.

Subgenus Hylocrinus Casey, 1907

Hylocrinus Casey, 1907: 289, 331. Type species: Eurymetopon longulum LeConte, 1851, original designation.

Hylocrinus ambiguus (Champion, 1884) PAN

Emmenastus ambiguus Champion, 1884: 13.

Hylocrinus angustus (Casey, 1890)USA (AZ)

Emmenastus angustus Casey, 1890b: 352.

Hylocrinus blaisdelli Casey, 1907USA (CA)

Hylocrinus blaisdelli Casey, 1907: 336.

Hylocrinus breviusculus Casey, 1907USA (TX)

Hylocrinus breviusculus Casey, 1907: 334.

Hylocrinus cunctans Casey, 1907USA (TX)

Hylocrinus cunctans Casey, 1907: 336.

Hylocrinus delicatulus Casey, 1907USA (AZ NV UT)

Hylocrinus delicatulus Casey, 1907: 334.

Hylocrinus depressulus Casey, 1907USA (CA)

Hylocrinus depressulus Casey, 1907: 335.

Hylocrinus filitarsis Casey, 1907USA (CA)

Hylocrinus filitarsis Casey, 1907: 333.

Hylocrinus guatemalensis (Champion, 1884) GUA

Emmenastus guatemalensis Champion, 1884: 14.

Hylocrinus longulus (LeConte, 1851)USA (AZ CA) MEX (BC SO)

Eurymetopon longulum LeConte, 1851: 139.

Hylocrinus magnus Blaisdell, 1923MEX (SO)

Hylocrinus magnus Blaisdell, 1923: 219.

Hylocrinus tenuis Casey, 1907USA (AZ)

Hylocrinus tenuis Casey, 1907: 333.

Subgenus Locrodes Casey, 1907

Locrodes Casey, 1907: 332. Type species: Emmenastus piceus Casey, 1890, present designation.

Hylocrinus brunnescens Casey, 1907USA (UT)

Hylocrinus brunnescens Casey, 1907: 338.

Hylocrinus fraternus Casey, 1907USA (ID UT)

Hylocrinus fraternus Casey, 1907: 338.

Hylocrinus insularis Blaisdell, 1923MEX (BS)

Hylocrinus insularis Blaisdell, 1923: 218.

Hylocrinus laborans Casey, 1907USA (NV UT)

Hylocrinus laborans Casey, 1907: 337.

Hylocrinus mexicanus (Champion, 1892)MEX (FD)

Emmenastus mexicanus Champion, 1892: 481.

Hylocrinus oblongulus Casey, 1907USA (CA) MEX (BC BS)

Hylocrinus oblongulus Casey, 1907: 337.

Hylocrinus parallelus (Champion, 1884)MEX (JA ME MO OA PU SI)

Emmenastus parallelus Champion, 1884: 12.

Hylocrinus piceus (Casey, 1890)USA (CA)

Emmenastus piceus Casey, 1890b: 353.

Hylocrinus seriatus (Champion, 1892)MEX (OA)

Emmenastus seriatus Champion, 1892: 482.

Hylocrinus subapterus (Champion, 1892)MEX (DU)

Emmenastus subapterus Champion, 1892: 481.

Hylocrinus tenebrosus (Champion, 1884)MEX (AG FD GU)

Emmenastus tenebrosus Champion, 1884: 12.

Hylocrinus umbrosus Casey, 1907USA (UT)

Hylocrinus umbrosus Casey, 1907: 338.

Subgenus Paravius Casey, 1907

Paravius Casey, 1907: 332. Type species: Emmenastus marginatus Casey, 1890, monotypy.

Hylocrinus marginatus (Casey, 1890)MEX (BC)

Emmenastus marginatus Casey, 1890b: 351.

Hylocrinus vicinus (Champion, 1884)USA (CA)

Emmenastus vicinus Champion, 1884: 8.

Genus Melanastus Casey, 1907 [M]

Melanastus Casey, 1907: 289. Type species: Eurymetopon atrum LeConte, 1851, original designation.

Melanastus acuminatus Casey, 1907USA (CO)

Melanastus acuminatus Casey, 1907: 362.

Melanastus acutus (Horn, 1870) [Fig. 14] CAN (AB SK) USA (NE)

Figure 14. 

Melanastus acutus (Horn, 1870). Scale bar = 1 mm.

Emmenastus acutus Horn, 1870: 270.

Melanastus aequicollis Casey, 1907USA (CA)

Melanastus aequicollis Casey, 1907: 360.

Melanastus ater (LeConte, 1851)USA (CA ID OR)

Eurymetopon atrum LeConte, 1851: 139.

Melanastus coarcticollis (Casey, 1890)USA (NM)

Emmenastus coarcticollis Casey, 1890b: 364.

Melanastus crassicornis (Casey, 1890)USA (CA)

Emmenastus crassicornis Casey, 1890b: 363.

Melanastus exiguus Casey, 1907USA (CO)

Melanastus exiguus Casey, 1907: 363.

Melanastus exoletus Casey, 1907USA (CA)

Melanastus exoletus Casey, 1907: 357.

Melanastus fallax (Casey, 1890)USA (NM)

Emmenastus fallax Casey, 1890b: 361.

Melanastus finitimus Casey, 1907USA (CO)

Melanastus finitimus Casey, 1907: 359.

Melanastus implicans Casey, 1907USA (CO)

Melanastus implicans Casey, 1907: 358.

Melanastus lucidulus Casey, 1907USA (CA)

Melanastus lucidulus Casey, 1907: 358.

Melanastus ludius Casey, 1907USA (UT)

Melanastus ludius Casey, 1907: 361.

Melanastus moestus Casey, 1907USA (CA)

Melanastus moestus Casey, 1907: 355.

Melanastus nitidus (Casey, 1890)USA (AZ)

Emmenastus nitidus Casey, 1890b: 362.

Melanastus nuperus Casey, 1907USA (AZ)

Melanastus nuperus Casey, 1907: 364.

Melanastus obesus (LeConte, 1851)USA (CA) MEX (BC BS)

Eurymetopon obesum LeConte, 1851: 139.

Emmenastus nanulus Casey, 1884: 45. Synonymy: Casey (1907b: 363).

Melanastus obscurus Blaisdell, 1923 MEX (SO)

Melanastus obscurus Blaisdell, 1923: 226.

Melanastus obtusus (LeConte, 1866)USA (CA)

Emmenastus obtusus LeConte, 1866b: 107.

Melanastus otiosus Casey, 1907USA (CA)

Melanastus otiosus Casey, 1907: 358.

Melanastus parvus Casey, 1907USA (CO)

Melanastus parvus Casey, 1907: 362.

Melanastus sonoricus Casey, 1907MEX (CH MI)

Melanastus sonoricus Casey, 1907: 364.

Melanastus sterilis Casey, 1907USA (CA)

Melanastus sterilis Casey, 1907: 357.

Melanastus texanus Blaisdell, 1926USA (TX)

Melanastus texanus Blaisdell, 1926b: 22.

Melanastus thoracicus (Casey, 1890)USA (CA)

Emmenastus thoracicus Casey, 1890b: 362.

Melanastus vegrandis Casey, 1907USA (CA)

Melanastus vegrandis Casey, 1907: 360.

Genus Mencheres Champion, 1884 [M]

Mencheres Champion, 1884: 5. Type species: Mencheres nicaraguensis Champion, 1884, subsequent designation (Lucas 1920: 403).

Mencheres elongatus Champion, 1884 GUA

Mencheres elongatus Champion, 1884: 6.

Mencheres nicaraguensis Champion, 1884 NIC

Mencheres nicaraguensis Champion, 1884: 5.

Genus Mesabates Champion, 1884 [M]

Mesabates Champion, 1884: 3. Type species: Mesabates latifrons Champion, 1884, monotypy.

Mesabates latifrons Champion, 1884MEX (OA PU)

Mesabates latifrons Champion, 1884: 3.

Mesabates spissicornis Champion, 1892MEX (SI)

Mesabates spissicornis Champion, 1892: 479.

Genus Mesabatodes Casey, 1907 [M]

Mesabatodes Casey, 1907: 517. Type species: Mesabates inaequalis Champion, 1892, original designation.

Mesabatodes inaequalis (Champion, 1892)MEX (AG CH DU)

Mesabates inaequalis Champion, 1892: 480.

Genus Metoponium Casey, 1907 [N]

Metoponium Casey, 1907: 288. Type species: Eurymetopon abnorme LeConte, 1851, original designation.

Subgenus Metoponiopsis Casey, 1907

Metoponiopsis Casey, 1907: 290. Type species: Eurymetopon bicolor Horn, 1870, monotypy.

Metoponium bicolor (Horn, 1870)USA (AZ CA) MEX (BC)

Eurymetopon bicolor Horn, 1870: 268.

Subgenus Metoponium Casey, 1907

Metoponium Casey, 1907: 288. Type species: Eurymetopon abnorme LeConte, 1851, original designation.

Metoponium abnorme abnorme (LeConte, 1851)USA (CA)

Eurymetopon abnorme LeConte, 1851: 138.

Metoponium abnorme faustum Casey, 1907USA (CA)

Metoponium faustum Casey, 1907: 292.

Metoponium abnorme laticolle Casey, 1907USA (AZ) MEX (BC BS)

Metoponium laticolle Casey, 1907: 291.

Metoponium angelicum Blaisdell, 1923MEX (BC)

Metoponium angelicum Blaisdell, 1923: 203.

Metoponium arizonicum Casey, 1907USA (AZ)

Metoponium arizonicum Casey, 1907: 294.

Metoponium candidum Casey, 1907USA (AZ) MEX (SO)

Metoponium candidum Casey, 1907: 292.

Metoponium cognitum Casey, 1907USA (TX)

Metoponium cognitum Casey, 1907: 305.

Metoponium concors Casey, 1907USA (CA)

Metoponium concors Casey, 1907: 305.

Metoponium congener (Casey, 1890)USA (TX)

Eurymetopon congener Casey, 1890b: 333.

Metoponium convexicolle (LeConte, 1851)USA (AZ CA NV) MEX (BC BS)

Eurymetopon convexicolle LeConte, 1851: 139.

Metoponium crassum Casey, 1907USA (AZ)

Metoponium crassum Casey, 1907: 299.

Metoponium cribriceps Casey, 1907USA (NM TX)

Metoponium cribriceps Casey, 1907: 305.

Metoponium cylindricum (Casey, 1890)USA (CA)

Eurymetopon cylindricum Casey, 1890b: 337.

Metoponium dubium (Casey, 1884)USA (AZ)

Eurymetopon dubium Casey, 1884: 44.

Eurymetopon carbonatum Casey, 1884: 43. Synonymy: Horn (1885b: 110).

Metoponium edax Casey, 1907USA (CA)

Metoponium edax Casey, 1907: 309.

Metoponium egregium Casey, 1907USA (CA)

Metoponium egregium Casey, 1907: 300.

Metoponium emarginatum (Casey, 1884)USA (AZ)

Eurymetopon emarginatum Casey, 1884: 41.

Eurymetopon piceum Casey, 1884: 40. Synonymy: Horn (1885b: 110).

Eurymetopon papagonum Casey, 1884: 42. Synonymy: Horn (1885b: 110).

Eurymetopon sculptile Casey, 1884: 44. Synonymy: Horn (1885b: 110).

Metoponium erosum Blaisdell, 1943MEX (BS)

Metoponium erosum Blaisdell, 1943: 175.

Metoponium extensum Casey, 1907USA (AZ)

Metoponium extensum Casey, 1907: 300.

Metoponium fatigans Casey, 1907USA (AZ)

Metoponium fatigans Casey, 1907: 304.

Metoponium fusculum (Casey, 1890)USA (AZ CA)

Eurymetopon fusculum Casey, 1890b: 335.

Metoponium gravidum Casey, 1907USA (CA)

Metoponium gravidum Casey, 1907: 308.

Metoponium gulosum Casey, 1907USA (CA)

Metoponium gulosum Casey, 1907: 307.

Metoponium hebes Casey, 1907USA (AZ)

Metoponium hebes Casey, 1907: 301.

Metoponium insulare Casey, 1907USA (CA)

Metoponium insulare Casey, 1907: 308.

Metoponium integer Casey, 1907USA (CA)

Metoponium integer Casey, 1907: 310.

Metoponium ludificans Casey, 1907USA (TX)

Metoponium ludificans Casey, 1907: 303.

Metoponium molestum Casey, 1907USA (CA)

Metoponium molestum Casey, 1907: 309.

Metoponium nevadense Casey, 1907USA (NV)

Metoponium nevadense Casey, 1907: 307.

Metoponium opacipenne Casey, 1907USA (CA)

Metoponium opacipenne Casey, 1907: 309.

Metoponium pacificum Blaisdell, 1923MEX (BS)

Metoponium pacificum Blaisdell, 1923: 202.

Metoponium pallescens Casey, 1907USA (AZ)

Metoponium pallescens Casey, 1907: 293.

Metoponium parvuliceps Casey, 1907USA (AZ)

Metoponium parvuliceps Casey, 1907: 296.

Metoponium perforatum anceps Casey, 1907USA (NM)

Metoponium anceps Casey, 1907: 294.

Metoponium perforatum congruens Casey, 1907USA (NM)

Metoponium congruens Casey, 1907: 293.

Metoponium perforatum perforatum (Casey, 1890)USA (AZ)

Eurymetopon perforatum Casey, 1890b: 334.

Metoponium phoenicis Casey, 1907USA (AZ)

Metoponium phoenicis Casey, 1907: 301.

Metoponium politum (Casey, 1890)USA (TX)

Eurymetopon politum Casey, 1890b: 338.

Metoponium probatum Casey, 1907USA (CA)

Metoponium probatum Casey, 1907: 310.

Metoponium procerum Casey, 1907USA (AZ)

Metoponium procerum Casey, 1907: 297.

Metoponium prolixum Casey, 1907USA (AZ)

Metoponium prolixum Casey, 1907: 298.

Metoponium rufescens Casey, 1907USA (AZ)

Metoponium rufescens Casey, 1907: 302.

Metoponium rufopiceum Casey, 1907USA (AZ)

Metoponium rufopiceum Casey, 1907: 296.

Metoponium saginatum Casey, 1907USA (TX)

Metoponium saginatum Casey, 1907: 296.

Metoponium socium socium Casey, 1907USA (AZ)

Metoponium socium Casey, 1907: 295.

Metoponium socium subsimile Casey, 1907USA (AZ)

Metoponium subsimile Casey, 1907: 295.

Metoponium subovale Casey, 1907USA (UT)

Metoponium subovale Casey, 1907: 307.

Metoponium tersum Casey, 1907USA (CA)

Metoponium tersum Casey, 1907: 306.

Metoponium testaceum Casey, 1907USA (CA)

Metoponium testaceum Casey, 1907: 303.

Metoponium transversum Blaisdell, 1943MEX (BS)

Metoponium transversum Blaisdell, 1943: 174.

Metoponium truncaticeps Casey, 1907USA (AZ)

Metoponium truncaticeps Casey, 1907: 299.

Genus Micrarmalia Casey, 1907 [F]

Micrarmalia Casey, 1907: 516. Type species: Emmenastus constrictus Champion, 1892, monotypy.

Micrarmalia constricta (Champion, 1892)MEX (GE MO)

Emmenastus constrictus Champion, 1892: 482.

Genus Micromes Casey, 1907 [M]

Micromes Casey, 1907: 432, 441. Type species: Stibia ovipennis Horn, 1874, original designation.

Micromes maritimus (Casey, 1892)USA (CA)

Stibia maritima Casey, 1891: 52.

Micromes ovipennis (Horn, 1874)USA (CA)

Stibia ovipennis Horn, 1874a: 28.

Genus Orthostibia Blaisdell, 1923 [F]

Orthostibia Blaisdell, 1923: 235. Type species: Orthostibia frontalis Blaisdell, 1923, original designation.

Orthostibia fraterna Blaisdell, 1943MEX (BS)

Orthostibia fraterna Blaisdell, 1943: 211.

Orthostibia frontalis Blaisdell, 1923MEX (BS)

Orthostibia frontalis Blaisdell, 1923: 236.

Orthostibia muricata Blaisdell, 1943MEX (BS)

Orthostibia muricata Blaisdell, 1943: 210.

Genus Oxygonodera Casey, 1907 [F]

Oxygonodera Casey, 1907: 433, 444. Type species: Oxygonodera villosa Casey, 1907, original designation.

Oxygonodera grandiceps Casey, 1907USA (UT)

Oxygonodera grandiceps Casey, 1907: 446.

Oxygonodera hispidula (Horn, 1874)USA (ID OR UT WA)

Stibia hispidula Horn, 1874a: 29.

Oxygonodera villosa Casey, 1907USA (UT)

Oxygonodera villosa Casey, 1907: 445.

Genus Pescennius Champion, 1884 [M]

Pescennius Champion, 1884: 3. Type species: Pescennius villosus Champion, 1884, monotypy.

Pescennius villosus Champion, 1884MEX (PU)

Pescennius villosus Champion, 1884: 4.

Genus Pimeliopsis Champion, 1892 [F]

Pimeliopsis Champion, 1892: 477. Type species: Pimeliopsis granulata Champion, 1892, monotypy.

Pimeliopsis granulata Champion, 1892MEX (GE)

Pimeliopsis granulata Champion, 1892: 477.

Genus Posides Champion, 1884 [M]

Posides Champion, 1884: 6. Type species: Posides dissidens Champion, 1884, monotypy.

Posides dissidens Champion, 1884MEX (PU)

Posides dissidens Champion, 1884: 6.

Genus Soemias Champion, 1884 [F]

Soemias Champion, 1884: 4. Type species: Soemias minuta Champion, 1884, monotypy.

Soemias minuta Champion, 1884MEX (VE)

Soemias minuta Champion, 1884: 5.

Genus Steriphanides Casey, 1907 [M]

Steriphanides Casey, 1907: 515. Type species: Emmenastus stolidus Champion, 1892, monotypy.

Steriphanides stolidus (Champion, 1892)MEX (OA)

Emmenastus stolidus Champion, 1892: 483.

Genus Steriphanus Casey, 1907 [M]

Steriphanus Casey, 1907: 289. Type species: Emmenastus conicicollis Casey, 1890, original designation.

Steriphanus aridus Casey, 1907USA (AZ)

Steriphanus aridus Casey, 1907: 347.

Steriphanus conicicollis (Casey, 1890)USA (AZ)

Emmenastus conicicollis Casey, 1890b: 355.

Steriphanus convexus convexus (LeConte, 1866)USA (AZ NM TX)

Emmenastus convexus LeConte, 1866b: 107.

Steriphanus convexus unicolor Casey, 1907USA (NM)

Steriphanus unicolor Casey, 1907: 346.

Steriphanus curtus (Champion, 1884)MEX (DU PU)

Emmenastus curtus Champion, 1884: 16.

Steriphanus discrepans Casey, 1907USA (AZ)

Steriphanus discrepans Casey, 1907: 343.

Steriphanus discretus (Casey, 1890)USA (AZ)

Emmenastus discretus Casey, 1890b: 354.

Steriphanus durus Blaisdell, 1923MEX (BC)

Steriphanus durus Blaisdell, 1923: 224.

Steriphanus ellipticus (Champion, 1884)MEX (“Pensacola”)

Emmenastus ellipticus Champion, 1884: 8.

Steriphanus estebani Blaisdell, 1923MEX (SO)

Steriphanus estebani Blaisdell, 1923: 225.

Steriphanus glabratus (Champion, 1884)MEX (JA OA PU)

Emmenastus glabratus Champion, 1884: 16.

Steriphanus hilaris Casey, 1907USA (AZ UT)

Steriphanus hilaris Casey, 1907: 345.

Steriphanus lentus (Champion, 1884)MEX (CH CO DU)

Emmenastus lentus Champion, 1884: 16.

Steriphanus libertus Casey, 1907USA (AZ)

Steriphanus libertus Casey, 1907: 350.

Steriphanus lubricans Casey, 1907USA (AZ NV)

Steriphanus lubricans Casey, 1907: 345.

Steriphanus lustrans Casey, 1907USA (AZ)

Steriphanus lustrans Casey, 1907: 344.

Steriphanus mancus (Champion, 1884)MEX (GE PU)

Emmenastus mancus Champion, 1884: 15.

Steriphanus nigrans Casey, 1907USA (AZ)

Steriphanus nigrans Casey, 1907: 347.

Steriphanus nitescens Casey, 1907USA (TX)

Steriphanus nitescens Casey, 1907: 344.

Steriphanus perovatus Casey, 1907USA (TX)

Steriphanus perovatus Casey, 1907: 351.

Steriphanus picipes (Champion, 1884)MEX (OA)

Emmenastus picipes Champion, 1884: 17.

Steriphanus placidus Casey, 1907MEX (FD)

Steriphanus placidus Casey, 1907: 347.

Steriphanus proprius Casey, 1907USA (AZ)

Steriphanus proprius Casey, 1907: 347.

Steriphanus pulvinatus (Champion, 1884)MEX (FD HI OA)

Emmenastus pulvinatus Champion, 1884: 17.

Steriphanus rugicollis (Champion, 1884)MEX (SL)

Emmenastus rugicollis Champion, 1884: 17.

Steriphanus rutilans Casey, 1907USA (TX)

Steriphanus rutilans Casey, 1907: 346.

Steriphanus subopacus alutaceus Casey, 1907USA (AZ) MEX (BC SO)

Steriphanus alutaceus Casey, 1907: 347.

Steriphanus subopacus peropacus Casey, 1907USA (AZ)

Steriphanus peropacus Casey, 1907: 349.

Steriphanus subopacus subopacus (Horn, 1870)USA (AZ TX) MEX (BC BS SO)

Emmenastus subopacus Horn, 1870: 269.

Steriphanus torpidus Blaisdell, 1923: 221. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Steriphanus mucronatus Blaisdell, 1923: 223. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Steriphanus tardus Blaisdell, 1923MEX (SO)

Steriphanus tardus Blaisdell, 1923: 222.

Genus Stibia Horn, 1870 [F]

Stibia Horn, 1870: 260. Type species: Stibia puncticollis Horn, 1870, monotypy.

Eutriorophus Casey, 1924: 296. Type species: Eutriorophus tuckeri Casey, 1924, original designation. Synonymy: Blaisdell (1933b: 210).

Stibia blairi Blaisdell, 1936USA (AZ CA) MEX (BC)

Stibia blairi Blaisdell, 1936a: 88.

Stibia cribrata Blaisdell, 1923MEX (BS)

Stibia cribrata Blaisdell, 1923: 23921.

Stibia fallaciosa fallaciosa Blaisdell, 1936MEX (BS)

Stibia fallaciosa Blaisdell, 1936a: 70.

Stibia fallaciosa interstitialis Blaisdell, 1936MEX (BS)

Stibia fallaciosa interstitialis Blaisdell, 1936a: 73.

Stibia ferruginea Blaisdell, 1943MEX (BS)

Stibia ferruginea Blaisdell, 1943: 208.

Stibia freyi Kulzer, 1959MEX (SI)

Stibia freyi Kulzer, 1959: 614.

Stibia granulata Blaisdell, 1923MEX (BS)

Stibia granulata Blaisdell, 1923: 238.

Stibia imperialis Blaisdell, 1936USA (AZ CA)

Stibia imperialis Blaisdell, 1936a: 94.

Stibia puncticollis martinensis Blaisdell, 1936MEX (BC)

Stibia puncticollis martinensis Blaisdell, 1936a: 83.

Stibia puncticollis puncticollis Horn, 1870USA (CA) MEX (BC BS SO)

Stibia puncticollis Horn, 1870: 260.

Stibia hannai Blaisdell, 1925b: 329. Synonymy: Blaisdell (1936a: 81).

Stibia sparsa Blaisdell, 1923MEX (BC BS)

Stibia sparsa Blaisdell, 1923: 237.

Stibia tortugensis Blaisdell, 1936a: 100. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Stibia tanneri Blaisdell, 1936USA (CA)

Stibia tanneri Blaisdell, 1936a: 97.

Stibia tuckeri (Casey, 1924)USA (AZ)

Eutriorophus tuckeri Casey, 1924: 297.

Stibia williamsi Blaisdell, 1925MEX (BC)

Stibia williamsi Blaisdell, 1925b: 328.

Genus Stictodera Casey, 1907 [F]

Stictodera Casey, 1907: 289, 352. Type species: Emmenastus pinguis LeConte, 1866, original designation.

Stictodera pinguis (LeConte, 1866)MEX (BS)

Emmenastus pinguis LeConte, 1866b: 107.

Genus Telabis Casey, 1890 [M]22

Telabis Casey, 1890b: 331. Type species: Eurymetopon longipenne Casey, 1890, subsequent designation (Casey 1907: 288).

Telabis alienus Casey, 1907USA (AZ)

Telabis aliena Casey, 1907: 325.

Telabis amicus Casey, 1907USA (UT)

Telabis amica Casey, 1907: 318.

Telabis asperus Casey, 1907USA (CO)

Telabis aspera Casey, 1907: 322.

Telabis blandus Casey, 1907USA (TX)

Telabis blanda Casey, 1907: 326.

Telabis brevicollis (Champion, 1884)MEX (CO)

Eurymetopon brevicolle Champion, 1884: 7.

Telabis compar Casey, 1907USA (AZ)

Telabis compar Casey, 1907: 321.

Telabis crassulus (Casey, 1890)USA (AZ TX)

Eurymetopon crassulum Casey, 1890b: 344.

Telabis curticollis Casey, 1907USA (AZ)

Telabis curticollis Casey, 1907: 321.

Telabis debilis (Casey, 1890)USA (AZ)

Eurymetopon debile Casey, 1890b: 343.

Telabis discors (Casey, 1890)USA (TX)

Eurymetopon discors Casey, 1890b: 342.

Telabis famelicus Casey, 1907USA (NM)

Telabis famelica Casey, 1907: 323.

Telabis fidelis Casey, 1907USA (CA)

Telabis fidelis Casey, 1907: 320.

Telabis hirtipes Blaisdell, 1923MEX (BC BS)

Telabis hirtipes Blaisdell, 1923: 205.

Telabis histricus (Casey, 1890)USA (AZ)

Eurymetopon histricum Casey, 1890b: 340.

Telabis incisus Casey, 1907USA (CA)

Telabis incisa Casey, 1907: 322.

Telabis inops Casey, 1907USA (AZ)

Telabis inops Casey, 1907: 325.

Telabis latipennis Blaisdell, 1923MEX (BS)

Telabis latipennis Blaisdell, 1923: 207.

Telabis lobifrons Casey, 1907USA (AZ)

Telabis lobifrons Casey, 1907: 318.

Telabis longipennis (Casey, 1890)USA (NM)

Eurymetopon longipenne Casey, 1890b: 339.

Telabis lunulatus Blaisdell, 1923MEX (BC BS)

Telabis lunulata Blaisdell, 1923: 206.

Telabis lustrellus Casey, 1907USA (NM)

Telabis lustrella Casey, 1907: 323.

Telabis mimeticus Casey, 1907USA (TX)

Telabis mimetica Casey, 1907: 319.

Telabis muricatulus (Casey, 1890)USA (AZ TX)

Eurymetopon muricatulum Casey, 1890b: 341.

Telabis nevadensis Blaisdell, 1925USA (NV)

Telabis nevadensis Blaisdell, 1925c: 372.

Telabis obtusus Casey, 1907USA (AZ)

Telabis obtusa Casey, 1907: 317.

Telabis opacellus Casey, 1907USA (CA)

Telabis opacella Casey, 1907: 316.

Telabis ovalis Casey, 1907USA (AZ)

Telabis ovalis Casey, 1907: 324.

Telabis pavidus Casey, 1907USA (NM)

Telabis pavida Casey, 1907: 324.

Telabis prominens Casey, 1907USA (TX)

Telabis prominens Casey, 1907: 314.

Telabis proxima Casey, 1907: 315. Synonymy: Casey (1911: 253).

Telabis punctulatus (LeConte, 1866)MEX (BC BS SO)

Eurymetopon punctulatum LeConte, 1866b: 105.

Telabis rubidus Casey, 1907USA (TX)

Telabis rubida Casey, 1907: 315.

Telabis serratus (LeConte, 1866)USA (AZ CA ID NM NV OR TX) MEX (BC BS SO)

Eurymetopon serratum LeConte, 1866b: 106.

Telabis sodalis (Horn, 1870)USA (AZ CA) MEX (BS)

Eurymetopon sodalis Horn, 1870: 268.

Telabis timidus Casey, 1907USA (AZ)

Telabis timida Casey, 1907: 320.

Telabis uteanus Casey, 1907USA (UT)

Telabis uteana Casey, 1907: 317.

Telabis vafer Casey, 1907USA (AZ)

Telabis vafra Casey, 1907: 315.

Telabis vapidus Casey, 1907USA (TX)

Telabis vapida Casey, 1907: 322.

Genus Telaponium Blaisdell, 1923 [N]

Telaponium Blaisdell, 1923: 209. Type species: Telaponium castaneum Blaisdell, 1923, original designation.

Telaponium castaneum Blaisdell, 1923MEX (BS)

Telaponium castaneum Blaisdell, 1923: 209.

Telaponium pingue Blaisdell, 1943MEX (BS)

Telaponium pingue Blaisdell, 1943: 179.

Genus Texaponium Thomas, 1984 [N]

Texaponium Thomas, 1984: 658. Type species: Cryptadius triplehorni Berry, 1974, original designation.

Texaponium triplehorni (Berry, 1974)USA (TX)

Cryptadius triplehorni Berry, 1974: 172.

Genus Tlascalinus Casey, 1907 [M]

Tlascalinus Casey, 1907: 370. Type species: Trimytis flohri Champion, 1892, monotypy.

Tlascalinus flohri (Champion, 1892)MEX (FD)

Trimytis flohri Champion, 1892: 478.

Genus Trichiotes Casey, 1907 [M]

Trichiotes Casey, 1907: 432, 443. Type species: Trichiotes seriatus Casey, 1907, original designation.

Trichiotes lightfooti Wirth and Smith, 2017MEX (CO)

Trichiotes lightfooti Wirth and Smith, 2017: 535.

Trichiotes seriatus Casey, 1907USA (NM TX) MEX (CO NL)

Trichiotes seriatus Casey, 1907: 444.

Genus Trientoma Solier, 1835 [F]

Trientoma Solier, 1835b: 256. Type species: Trientoma varvasi Solier, 1835, monotypy.

Trientoma cayensis Garrido and Gutiérrez, 1995 CUB

Trientoma cayensis Garrido and Gutiérrez, 1995b: 48.

Trientoma convexipennis Allard, 1883 CUB

Trientoma convexipennis Allard, 1883: 14.

Trientoma garridoi Marcuzzi, 1988 CUB

Trientoma garridoi Marcuzzi, 1988: 69.

Trientoma guadeloupensis Fleutiaux and Sallé, 1890 LAN

Trientoma guadeloupensis Fleutiaux and Sallé, 1890: 421.

Trientoma jilae Steiner, 2006 BAH

Trientoma jilae Steiner, 2006: 3.

Trientoma kaszabi Marcuzzi, 1985 CUB

Trientoma kaszabi Marcuzzi, 1985: 181.

Trientoma kochi Marcuzzi, 1977 CAY

Trientoma kochi Marcuzzi, 1977: 6.

Trientoma laevis Allard, 1883 HAI

Trientoma laevis Allard, 1883: 14.

Trientoma maisiensis Marcuzzi, 1988 CUB

Trientoma maisiensis Marcuzzi, 1988: 67.

Trientoma zayasi Marcuzzi, 1988: 70. Synonymy: Garrido and Gutiérrez (1995b: 48).

Trientoma martinicensis Allard, 1883LAN (Martinique)

Trientoma martinicensis Allard, 1883: 14.

Trientoma puertoricensis Marcuzzi, 1977 PRI

Trientoma puertoricensis Marcuzzi, 1977: 7.

Trientoma rugifrons Champion, 1884MEX / HIS

Trientoma rugifrons Champion, 1884: 2.

Trientoma ryticephala Allard, 1883 HAI

Trientoma ryticephala Allard, 1883: 14.

Trientoma sallei Kraatz, 1865MEX / HAIDOM

Trientoma sallei Kraatz, 1865: 74.

Trientoma mexicana Champion, 1884: 2. Synonymy: Champion (1892: 479).

Trientoma siboneyensis Marcuzzi, 1988 CUB

Trientoma siboneyensis Marcuzzi, 1988: 71.

Trientoma varvasi Solier, 1835 CUB

Trientoma varvasi Solier, 1835b: 257.

Trientoma voegeliorum Steiner, 2006 BAH

Trientoma voegeliorum Steiner, 2006: 8.

Trientoma wickhami Casey, 1907 BAH

Trientoma wickhami Casey, 1907: 377.

Genus Trimytantron Ardoin, 1977 [N]

Trimytantron Ardoin, 1977b: 381. Type species: Trimytantron decui Ardoin, 1977, original designation.

Bielawskia Marcuzzi, 1985: 179. Type species: Bielawskia cubana Marcuzzi, 1985 (= Trimytantron decui Ardoin, 1977), monotypy. Synonymy: Marcuzzi (1998a: 153).

Trimytantron armasi Garrido and Gutiérrez, 1997 CUB

Trimytantron armasi Garrido and Gutiérrez, 1997: 32.

Trimytantron cavernicolous Garrido and Gutiérrez, 1997 CUB

Trimytantron cavernicolous Garrido and Gutiérrez, 1997: 34.

Trimytantron cubanum Ardoin, 1977 CUB

Trimytantron cubanum Ardoin, 1977c: 388.

Trimytantron decui Ardoin, 1977 CUB

Trimytantron decui Ardoin, 1977b: 382.

Bielawskia cubana Marcuzzi, 1985: 179 [junior secondary homonym of Trimytantron cubanum Ardoin, 1977]. Synonymy: Garrido and Gutiérrez (1997: 30).

Trimytantron garridoi Marcuzzi, 1998a: 153. Replacement name for Trimytantron cubanum (Marcuzzi, 1985).

Trimytantron escambrayense Garrido and Gutiérrez, 1997 CUB

Trimytantron escambrayensis Garrido and Gutiérrez, 1997: 33.

Trimytantron litorale Garrido and Gutiérrez, 1997 CUB

Trimytantron litoralis Garrido and Gutiérrez, 1997: 30.

Trimytantron minus 23 Garrido and Gutiérrez, 1997 CUB

Trimytantron minor Garrido and Gutiérrez, 1997: 36.

Trimytantron negreai Ardoin, 1977 CUB

Trimytantron negreai Ardoin, 1977c: 387.

Trimytantron poeyi Ardoin, 1977 CUB

Trimytantron poeyi Ardoin, 1977b: 383.

Trimytantron pumilum Garrido and Gutiérrez, 1997 CUB

Trimytantron pumilus Garrido and Gutiérrez, 1997: 35.

Trimytantron punctulaticeps Garrido and Gutiérrez, 1997 CUB

Trimytantron punctulaticeps Garrido and Gutiérrez, 1997: 34.

Trimytantron sierrae Garrido and Gutiérrez, 1997 CUB

Trimytantron sierrae Garrido and Gutiérrez, 1997: 31.

Trimytantron vinai Ardoin, 1977 CUB

Trimytantron viñai Ardoin, 1977c: 388.

Genus Trimytis LeConte, 1851 [F]

Trimytis LeConte, 1851: 141. Type species: Trimytis pruinosa LeConte, 1851, monotypy.

Pimalius Casey, 1907: 367. Type species: Trimytis pulverea Horn, 1870, original designation. Synonymy: MacLachlan and Olson (1990: 79).

Trimytis ceralboensis Blaisdell, 1943MEX (BS)

Trimytis ceralboensis Blaisdell, 1943: 196.

Trimytis obovata Champion, 1892: 478MEX (CH)

Trimytis obovata Champion, 1892: 478.

Trimytis obtusa Horn, 1894MEX (BS)

Trimytis obtusa Horn, 1894b: 412.

Trimytis pruinosa LeConte, 1851USA (AZ CO KS MT NE NM SD TX WY)

Trimytis pruinosa LeConte, 1851: 141.

Trimytis nympha Casey, 1907: 368. Synonymy: MacLachlan and Olson (1990: 80).

Trimytis tonsa Casey, 1907: 369. Synonymy: MacLachlan and Olson (1990: 80).

Trimytis ignava Casey, 1907: 369. Synonymy: MacLachlan and Olson (1990: 81).

Trimytis trapezifera Casey, 1924: 299. Synonymy: MacLachlan and Olson (1990: 81).

Trimytis pulverea Horn, 1870USA (AZ)

Trimytis pulverea Horn, 1870: 261.

Trimytis subsenilis Blaisdell, 1923MEX (SO)

Trimytis subsenilis Blaisdell, 1923: 227.

Genus Triorophus LeConte, 1851 [M]

Triorophus LeConte, 1851: 141. Type species: Triorophus laevis LeConte, 1851, subsequent designation (Casey 1907: 432).

Triorophus basalis Casey, 1907USA (AZ)

Triorophus basalis Casey, 1907: 437.

Triorophus brevis Casey, 1907USA (TX)

Triorophus brevis Casey, 1907: 439.

Triorophus gracilicornis Casey, 1907USA (CA)

Triorophus gracilicornis Casey, 1907: 437.

Triorophus gravidulus Casey, 1907USA (AZ)

Triorophus gravidulus Casey, 1907: 437.

Triorophus histrio Casey, 1907USA (AZ)

Triorophus histrio Casey, 1907: 437.

Triorophus laevis laevis LeConte, 1851USA (AZ CA NV) MEX (SO)

Triorophus laevis LeConte, 1851: 141.

Triorophus laevis politus Casey, 1907USA (CA NV)

Triorophus politus Casey, 1907: 435.

Triorophus lariversi Blaisdell, 1942USA (NV)

Triorophus lariversi Blaisdell, 1942: 132.

Triorophus laticeps Casey, 1924USA (TX)

Triorophus laticeps Casey, 1924: 297.

Triorophus lecontei Casey, 1890USA (TX) MEX (CH DU)

Triorophus lecontei Casey, 1890b: 327.

Triorophus longicornis Casey, 1907USA (AZ)

Triorophus longicornis Casey, 1907: 438.

Triorophus mixtus Casey, 1907USA (TX)

Triorophus mixtus Casey, 1907: 440.

Triorophus mundulus Casey, 1907USA (AZ)

Triorophus mundulus Casey, 1907: 436.

Triorophus nevadensis Casey, 1924USA (NV)

Triorophus nevadensis Casey, 1924: 298.

Triorophus nodiceps LeConte, 1853USA (TX) MEX (CO)

Triorophus nodiceps LeConte, 1853: 446.

Triorophus puberulus Casey, 1924USA (CA)

Triorophus puberulus Casey, 1924: 298.

Triorophus punctatus LeConte, 1851USA (CA)

Triorophus punctatus LeConte, 1851: 142.

Triorophus rugiceps LeConte, 1851USA (CA ID)

Triorophus rugiceps LeConte, 1851: 142.24

Triorophus simplex Casey, 1907USA (AZ)

Triorophus simplex Casey, 1907: 436.

Triorophus subpubescens Horn, 1870USA (CA)

Triorophus subpubescens Horn, 1870: 259.

Triorophus terebratulus Casey, 1907USA (AZ)

Triorophus terebratulus Casey, 1907: 436.

Triorophus thoracicus Casey, 1924USA (AZ)

Triorophus thoracicus Casey, 1924: 298.

Genus Triphalopsis Blaisdell, 1923 [F]

Triphalopsis Blaisdell, 1923: 232. Type species: Triphalopsis partida Blaisdell, 1923, original designation.

Triphalopsis californica Doyen, 1983USA (CA) MEX (BC)

Triphalopsis californicus Doyen, 1983: 87.

Triphalopsis impressicollis Blaisdell, 1943MEX (BC)

Triphalopsis impressicollis Blaisdell, 1943: 203.

Triphalopsis partida Blaisdell, 1923MEX (BC BS SO)

Triphalopsis partida Blaisdell, 1923: 232.

Triphalopsis minor Blaisdell, 1923: 233. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Genus Triphalopsoides Doyen, 1990 [M]

Triphalopsoides Doyen, 1990: 222. Type species: Triphalopsoides lasiodorsa Doyen, 1990, monotypy.

Triphalopsoides lasiodorsa Doyen, 1990MEX (JA)

Triphalopsoides lasiodorsa Doyen, 1990: 224.

Genus Triphalus LeConte, 1866 [M]

Triphalus LeConte, 1866b: 104. Type species: Triphalus perforatus LeConte, 1866, monotypy.

Triphalus cribricollis Horn, 1895MEX (BS)

Triphalus cribricollis Horn, 1895: 251.

Triphalus impressifrons Blaisdell, 1943MEX (BS)

Triphalus impressifrons Blaisdell, 1943: 202.

Triphalus perforatus LeConte, 1866MEX (BS)

Triphalus perforatus LeConte, 1866b: 104.

Triphalus subcylindricus Blaisdell, 1923MEX (BS)

Triphalus subcylindricus Blaisdell, 1923: 234.

Genus Troglogeneion Aalbu, 1985 [N]

Troglogeneion Aalbu, 1985: 541. Type species: Troglogeneion zapoteca Aalbu, 1985, monotypy.

Troglogeneion zapoteca Aalbu, 1985MEX (OA)

Troglogeneion zapoteca Aalbu, 1985: 542.

Tribe Epitragini Blanchard, 1845

Lygophila Rafinesque, 1815: 113 [nomen oblitum, see Bouchard et al. (2007: 386)]. Type genus: Lygophilus Rafinesque, 1815 (= Epitragus Latreille, 1802).

Épitragites Blanchard, 1845: 16 [nomen protectum]. Type genus: Epitragus Latreille, 1802.

Genus Bothrotes Casey, 1907 [M]

Bothrotes Casey, 1907: 379, 398. Type species: Epitragus canaliculatus Say, 1824, original designation.

Bothrotes angusticollis (Champion, 1884)MEX (GE JA SI)

Epitragus angusticollis Champion, 1884: 26.

Bothrotes bicarinatus (Champion, 1884)MEX (CL VE)

Epitragus bicarinatus Champion, 1884: 25.

Bothrotes canaliculatus acutus (LeConte, 1866)USA (FL KS NM OK TX) MEX (CO)

Epitragus acutus LeConte, 1866b: 108.

Bothrotes fortis Casey, 1907: 399. Synonymy: Freude (1967: 283).

Bothrotes subrudis Casey, 1907: 400. Synonymy: Freude (1967: 283).

Bothrotes pensus Casey, 1907: 400. Synonymy: Freude (1967: 283).

Bothrotes knausi Casey, 1907: 401. Synonymy: Freude (1967: 283).

Bothrotes canaliculatus arundinis (LeConte, 1866)USA (DE GA MD NC NJ NY SC VA)

Epitragus arundinis LeConte, 1866b: 108.

Bothrotes pinorum Casey, 1924: 304. Synonymy: Freude (1967: 281).

Bothrotes canaliculatus canaliculatus (Say, 1824)USA (AZ CO IL KS MO NM OH SD TX WI) MEX (CH DU SO ZA)

Epitragus canaliculatus Say, 1824b: 281.

Bothrotes canaliculatus mexicanus Freude, 1967MEX (DU NL TA)

Bothrotes canaliculatus mexicanus Freude, 1967: 285.

Bothrotes canus (Champion, 1884)MEX (GE)

Epitragus canus Champion, 1884: 34.

Bothrotes cristatus (Champion, 1892)MEX (CO GE)

Epitragus cristatus Champion, 1892: 485.

Bothrotes foveatus (Champion, 1884)MEX (OA VE)

Epitragus foveatus Champion, 1884: 29.

Bothrotes hoegei (Champion, 1884)MEX (MI VE)

Epitragus högei Champion, 1884: 26.

Bothrotes inaequalis (Champion, 1884)MEX (OA PU VE)

Epitragus inaequalis Champion, 1884: 32.

Bothrotes incisus (Champion, 1884)MEX (NA [Islas Marías])

Epitragus incisus Champion, 1884: 28.

Bothrotes littoralis (Champion, 1884)MEX (GE JA MO NA OA SI)

Epitragus littoralis Champion, 1884: 27.

Bothrotes ornatus (Champion, 1884)MEX (DU GU PU VE)

Epitragus ornatus Champion, 1884: 26.

Bothrotes plumbeus plumbeus (LeConte, 1866)USA (AZ CA CO KS NE NM SD TX)

Epitragus plumbeus LeConte, 1866b: 109.

Bothrotes aeneicollis Casey, 1907: 401. Synonymy: Freude (1967: 287).

Bothrotes chalceus Casey, 1907: 402. Synonymy: Freude (1967: 287).

Bothrotes affinis Casey, 1907: 405. Synonymy: Freude (1967: 287).

Bothrotes pertinax Casey, 1907: 405. Synonymy: Freude (1967: 287).

Bothrotes picipennis Casey, 1907: 406. Synonymy: Freude (1967: 287).

Bothrotes secutor Casey, 1907: 406. Synonymy: Freude (1967: 287).

Bothrotes secutor var. apertus Casey, 1907: 406. Synonymy: Freude (1967: 287).

Bothrotes acomanus Casey, 1907: 407. Synonymy: Freude (1967: 287).

Bothrotes neglectus Casey, 1907: 407. Synonymy: Freude (1967: 287).

Bothrotes insitus Casey, 1907: 408. Synonymy: Freude (1967: 287).

Bothrotes funebris Casey, 1907: 409. Synonymy: Freude (1967: 287).

Bothrotes plumbeus rorulentus (Champion, 1884)MEX (CO GU SI)

Epitragus rorulentus Champion, 1884: 27.

Bothrotes plumbeus tenebrosus Casey, 1907USA (AZ) MEX (SO)

Bothrotes tenebrosus Casey, 1907: 403.

Bothrotes occipitalis Casey, 1907: 403. Synonymy: Freude (1967: 296).

Bothrotes confertus Casey, 1907: 404. Synonymy: Freude (1967: 296).

Bothrotes eversus Casey, 1907: 404. Synonymy: Freude (1967: 296).

Bothrotes perditus Casey, 1907: 409. Synonymy: Freude (1967: 296).

Bothrotes amplificans Casey, 1907: 410. Synonymy: Freude (1967: 296).

Bothrotes obsolescens Casey, 1907: 411. Synonymy: Freude (1967: 296).

Bothrotes scutatus occidentalis Freude, 1967MEX (CH CL JA NA OA)

Bothrotes scutatus occidentalis Freude, 1967: 304.

Bothrotes scutatus scutatus (Champion, 1884)MEX (GU)

Epitragus scutatus Champion, 1884: 28.

Genus Conoecus Horn, 1885 [M]

Conoecus Horn, 1885c: 159. Type species: Conoecus ovipennis Horn, 1885, monotypy.

Conoecus ovipennis estriatus Casey, 1907USA (LA TX)

Conoecus estriatus Casey, 1907: 431.

Conoecus ovipennis ovipennis Horn, 1885USA (TX)

Conoecus ovipennis Horn, 1885c: 159.

Genus Cyrtomius Casey, 1907 [M]

Cyrtomius Casey, 1907: 379. Type species: Cyrtomius cavicauda Casey, 1907(=Epitragus plicatus Champion, 1884), original designation.

Subgenus Cyrtomius Casey, 1907

Cyrtomius Casey, 1907: 379. Type species: Cyrtomius cavicauda Casey, 1907(= Epitragus plicatus Champion, 1884), original designation.

Cyrtomius chevrolati (Champion, 1884)MEX (DU GU MO PU VE) GUANIC

Epitragus chevrolati Champion, 1884: 30.

Cyrtomius freyi Freude, 1967MEX (GE JA MI PU)

Cyrtomius freyi Freude, 1967: 229.

Cyrtomius plicatus (Champion, 1884)MEX (FD OA VE)

Epitragus plicatus Champion, 1884: 31.

Cyrtomius cavicauda Casey, 1907: 384. Synonymy: Freude (1967: 231).

Subgenus Grandicyrtomius Freude, 1967

Grandicyrtomius Freude, 1967: 225. Type species: Epitragus grandis Champion, 1884, original designation.

Cyrtomius grandis (Champion, 1884)MEX (CI DU GE JA MO OA PU SI SO VE)

Epitragus grandis Champion, 1884: 31.

[incertae sedis]

Cyrtomius gaigli Freude, 1986 GUA

Cyrtomius gaigli Freude, 1986: 27.

Cyrtomius polli Freude, 1986 GUA

Cyrtomius polli Freude, 1986: 26.

Genus Epitragodes Casey, 1890 [M]

Epitragodes Casey, 1890b: 365. Type species: Epitragus tomentosus LeConte, 1866, monotypy.

Epitragodes tomentosus macilentus Casey, 1907USA (AL FL GA NC SC VA) / BAH

Epitragodes tomentosus macilentus Casey, 1907: 425.

Epitragodes debilicollis Casey, 1907: 423. Synonymy: Freude (1968: 86).

Epitragodes pardalis Casey, 1907: 423. Synonymy: Freude (1968: 86).

Epitragodes cuprascens Casey, 1907: 424. Synonymy: Freude (1968: 86).

Epitragodes tomentosus tomentosus (LeConte, 1866)USA (FL GA) / BAH

Epitragus tomentosus LeConte, 1866b: 109.

Epitragodes floridanus Casey, 1907: 424. Synonymy: Freude (1968: 86).

Epitragodes obesulus Casey, 1907: 425. Synonymy: Freude (1968: 86).

Genus Epitragopsis Casey, 1907 [F]

Epitragopsis Casey, 1907: 386. Type species: Epitragus godmani Champion, 1884, original designation.

Epitragopsis communis (Champion, 1884)MEX (OA VE) GUABELHON

Epitragus communis Champion, 1884: 36.

Epitragopsis godmani (Champion, 1884)PAN / SA

Epitragus godmani Champion, 1884: 36.

Epitragopsis auratus Marcuzzi, 1961: 8. Synonymy: Freude (1968: 71).

Epitragopsis ruatanensis (Champion, 1892) HON NIC

Epitragus ruatanensis Champion, 1892: 488.

Genus Epitragosoma Brown and Triplehorn, 2002 [N]

Epitragosoma Brown and Triplehorn, 2002: 515. Type species: Epitragosoma arenaria Brown and Triplehorn, 2002, original designation.

Epitragosoma arenarium Brown and Triplehorn, 2002USA (NM TX)

Epitragosoma arenaria Brown and Triplehorn, 2002: 519.

Genus Epitragus Latreille, 1802 [M]

Epitragus Latreille, 1802: 165. Type species: Epitragus fuscus Latreille, 1804, subsequent monotypy in Latreille (1804: 322).

Subgenus Epitragus Latreille, 1802

Epitragus Latreille, 1802: 165. Type species: Epitragus fuscus Latreille, 1804, subsequent monotypy in Latreille (1804: 322).

Epitragus antillensis Marcuzzi, 1961 JAM

Epitragus antillensis Marcuzzi, 1961: 28.

Epitragus aurulentus aurulentus Kirsch, 1866NICCRIPAN / CUBJAMHISPRILAN / SA

Epitragus aurulentus Kirsch, 1866: 189.

Epitragus jamaicensis Champion, 1896: 3. Synonymy: Freude (1967: 158).

Epitragus emarginatus Champion, 1884 PAN

Epitragus emarginatus Champion, 1884: 24.

Epitragus consimilis Marcuzzi, 1961: 34. Synonymy: Freude (1967: 164).

Epitragus gaigli Freude, 1986 GUA

Epitragus gaigli Freude, 1986: 25.

Epitragus mexicanus Marcuzzi, 1961MEX (OA)

Epitragus mexicanus Marcuzzi, 1961: 29.

Epitragus nigricans Champion, 1884PAN / SA

Epitragus nigricans Champion, 1884: 24.

Epitragus puberulus Kirsch, 1886: 332. Synonymy: Freude (1967: 162).

Epitragus roscidus Erichson, 1849LAN / SA

Epitragus roscidus Erichson, 1849: 565.

Epitragus exaratus Champion, 1896: 2. Synonymy: Freude (1967: 166).

Epitragus sallei Champion, 188425MEX (CI VE YU) GUAHONNICCRI / SA

Epitragus sallaei Champion, 1884: 24.

Epitragus rigens Casey, 1907: 381. Synonymy: Freude (1967: 155).

Genus Hemasodes Casey, 1907 [M]

Hemasodes Casey, 1907: 378. Type species: Schoenicus vestitus Champion, 1884, original designation.

Hemasodes vestitus (Champion, 1884)MEX (GE JA OA VE)

Schoenicus vestitus Champion, 1884: 22.

Schoenicus yucatanensis Champion, 1884: 22. Synonymy: Freude (1967: 182).

Genus Lobometopon Casey, 1907 [N]

Lobometopon Casey, 1907: 379, 385. Type species: Epitragus fusiformis Casey, 1890, original designation.

Lobometopon acutangulum (Champion, 1884)MEX (CI OA) GUA

Epitragus acutangulus Champion, 1884: 31.

Lobometopon aeratum (Champion, 1884)MEX (CL GE JA NA OA VE)

Epitragus aeratus Champion, 1884: 33.

Lobometopon aurichalceum (Champion, 1884)USA (AZ) MEX (GU OA SI)

Epitragus aurichalceus Champion, 1884: 33.

Lobometopon tuckeri Casey, 1924: 301. Synonymy: Freude (1968: 44).

Lobometopon cupreum (Champion, 1884) GUA NIC CRI PAN

Epitragus cupreus Champion, 1884: 34.

Lobometopon bicaviceps Casey, 1907: 394. Synonymy: Freude (1968: 36).

Lobometopon alveolatum Casey, 1907: 394. Synonymy: Freude (1968: 36).

Lobometopon fusiforme cribricolle Casey, 1907USA (KS NE NM SD TX) MEX (NL)

Lobometopon cribricolle Casey, 1907: 391.

Lobometopon jucundum Casey, 1907: 392. Synonymy: Freude (1968: 50).

Lobometopon obscurum Casey, 1907: 395. Synonymy: Freude (1968: 50).

Lobometopon fusiforme fusiforme (Casey, 1890)USA (AZ) MEX (SO)

Epitragus fusiformis Casey, 1890b: 365.

Lobometopon symmetricum Casey, 1907: 389. Synonymy: Freude (1968: 50).

Lobometopon pimalicum Casey, 1907: 389. Synonymy: Freude (1968: 50).

Lobometopon aeneopiceum Casey, 1907: 390. Synonymy: Freude (1968: 50).

Lobometopon docile Casey, 1907: 389. Synonymy: Freude (1968: 50).

Lobometopon propinquum Casey, 1907: 391. Synonymy: Freude (1968: 50).

Lobometopon aequipenne Casey, 1907: 393. Synonymy: Freude (1968: 50).

Lobometopon morrisoni Casey, 1907: 393. Synonymy: Freude (1968: 50).

Lobometopon fusiforme uintanum Casey, 1907USA (AZ NM NV UT)

Lobometopon uintanum Casey, 1907: 388.

Lobometopon parvicolle Casey, 1907: 392. Synonymy: Freude (1968: 50).

Lobometopon alticola Casey, 1924: 302. Synonymy: Freude (1968: 50).

Lobometopon woodgatei Casey, 1924: 302. Synonymy: Freude (1968: 50).

Lobometopon provoanum Casey, 1924: 303. Synonymy: Freude (1968: 50).

Lobometopon guatemalense (Champion, 1884) GUA BEL SAL HON NIC CRI

Epitragus guatemalensis Champion, 1884: 32.

Lobometopon lucidum (Champion, 1884)MEX (DU NA PU SI SO)

Epitragus lucidus Champion, 1884: 34.

Lobometopon metallicum (Champion, 1884)MEX (CH CI CL DU FD GE GU ME MO OA PU QU SL VE) GUACRI

Epitragus metallicus Champion, 1884: 29.

Epitragus gracilis Casey, 1890b: 366. Synonymy: Freude (1968: 40).

Lobometopon aberrans Casey, 1907: 387. Synonymy: Freude (1968: 40).

Lobometopon micans (Champion, 1884)MEX (CI FD OA)

Epitragus micans Champion, 1884: 32.

Lobometopon obovatum (Champion, 1884)MEX (GE OA)

Epitragus obovatus Champion, 1884: 35.

Lobometopon ovale (Casey, 1885)USA (TX)

Epitragus ovalis Casey, 1885: 184.

Lobometopon parviceps (Champion, 1884)MEX (CL OA)

Epitragus parviceps Champion, 1884: 34.

Genus Metopoloba Casey, 1907 [F]

Metopoloba Casey, 1907: 379. Type species: Epitragus pruinosus Horn, 1870, original designation.

Metopoloba pruinosa mexicana Freude, 1967MEX (SO)

Metopoloba pruinosa mexicana Freude, 1967: 259.

Metopoloba pruinosa pruinosa (Horn, 1870)USA (AZ CA NV UT)

Epitragus pruinosus Horn, 1870: 264.

Metopoloba bifossiceps Casey, 1907: 413. Synonymy: Freude (1967: 248).

Metopoloba proba Casey, 1907: 414. Synonymy: Freude (1967: 248).

Metopoloba punctiventris Casey, 1907: 414. Synonymy: Freude (1967: 248).

Metopoloba perpolita Casey, 1907: 415. Synonymy: Freude (1967: 248).

Metopoloba californica Casey, 1907: 419. Synonymy: Freude (1967: 248).

Lobometopon juabense Casey, 1924: 303. Synonymy: Freude (1967: 248).

Metopoloba pruinosa subpilosa Blaisdell, 1943MEX (BS)

Metopoloba subpilosa Blaisdell, 1943: 199.

Metopoloba pruinosa subseriata Casey, 1907USA (AZ NM TX)

Metopoloba subseriata Casey, 1907: 415.

Metopoloba snowi Casey, 1907: 416. Synonymy: Freude (1967: 253).

Metopoloba densiventris Casey, 1907: 417. Synonymy: Freude (1967: 253).

Metopoloba contaminans Casey, 1907: 418. Synonymy: Freude (1967: 253).

Metopoloba amplexa Casey, 1907: 418. Synonymy: Freude (1967: 253).

Metopoloba sublaeviceps Casey, 1907: 418. Synonymy: Freude (1967: 253).

Metopoloba angulata Casey, 1907: 419. Synonymy: Freude (1967: 253).

Metopoloba pruinosa werneri Freude, 1967MEX (BS)

Metopoloba pruinosa werneri Freude, 1967: 259.

Genus Ortheolus Casey, 1907 [M]

Ortheolus Casey, 1907: 380. Type species: Schoenicus oculatus Champion, 1884, original designation.

Ortheolus antillarum (Champion, 1896) LAN

Schoenicus antillarum Champion, 1896: 5.

Schoenicus brunneus Champion, 1896: 4. Synonymy: Freude (1968: 106).

Ortheolus caraibicus caraibicus Marcuzzi, 1961LAN / SA

Ortheolus caraibicus Marcuzzi, 1961: 38.

Ortheolus oculatus oculatus (Champion, 1884) PAN

Schoenicus oculatus Champion, 1884: 18.

Ortheolus panamensis (Champion, 1884) CRI PAN

Schoenicus panamensis Champion, 1884: 18.

Genus Pechalius Casey, 1907 [M]

Pechalius Casey, 1907: 379, 420. Type species: Pechalius subvittatus Casey, 1907, original designation.

Epitragoma Casey, 1907: 386. Type species: Epitragus vestitus Casey, 1891, monotypy. Synonymy: Freude (1968: 61).

Pechalius bradleyi Triplehorn, 1974USA (NM)

Pechalius bradleyi Triplehorn, 1974: 73.

Pechalius dentiger (Horn, 1870)USA (AZ) MEX (SO)

Epitragus dentiger Horn, 1870: 265.

Pechalius pilosus (Champion, 1884)MEX (CH TA VE)

Epitragus pilosus Champion, 1884: 34.

Pechalius subvittatus Casey, 1907USA (TX) MEX (DU)

Pechalius subvittatus Casey, 1907: 421.

Pechalius vestitus (Casey, 1891)USA (AZ)

Epitragus vestitus Casey, 1891: 53.

Genus Phegoneus Casey, 1907 [M]

Phegoneus Casey, 1907: 380, 426. Type species: Epitragodes julichi Casey, 1891, original designation.

Subgenus Pectphegoneus Freude, 1968

Pectphegoneus Freude, 1968: 90. Type species: Schoenicus pectoralis Champion, 1884, monotypy.

Phegoneus pectoralis (Champion, 1884)MEX (CL GE JA MI MO PU)

Schoenicus pectoralis Champion, 1884: 21.

Subgenus Phegoneus Casey, 1907

Phegoneus Casey, 1907: 380, 426. Type species: Epitragodes julichi Casey, 1891, original designation.

Phegoneus basalis (Champion, 1884)MEX (OA VE)

Schoenicus basalis Champion, 1884: 21.

Phegoneus chalybeus (Champion, 1884)MEX (MI NA OA PU SI SL VE)

Schoenicus chalybeus Champion, 1884: 20.

Phegoneus difficilis (Champion, 1884)MEX (DU GE JA MI OA SI VE)

Schoenicus difficilis Champion, 1884: 20.

Phegoneus julichi (Casey, 1891)USA (TX)

Epitragodes jülichi Casey, 1891: 55.

Phegoneus rufipes impressus (Champion, 1884) CRI

Schoenicus impressus Champion, 1884: 20.

Phegoneus rufipes rufipes (Champion, 1884)MEX (YU) NIC

Schoenicus rufipes Champion, 1884: 19.

Phegoneus salvini salvini (Champion, 1884) GUA SAL CRI

Schoenicus salvini Champion, 1884: 19.

Schoenicus niger Champion, 1884: 20. Synonymy: Freude (1968: 96).

Phegoneus salvini subaeneus Casey, 1907 PAN

Phegoneus subaeneus Casey, 1907: 428.

Phegoneus viridis (Champion, 1884)MEX (CI CL DU GE JA OA PU SI) GUACRI

Schoenicus viridis Champion, 1884: 19.

Genus Polemiotus Casey, 1907 [M]

Polemiotus Casey, 1907: 379, 381. Type species: Epitragus submetallicus LeConte, 1854, original designation.

Polemiotus submetallicus (LeConte, 1854)USA (AZ)

Epitragus submetallicus LeConte, 1854c: 224.

Polemiotus humeralis Casey, 1907: 382. Synonymy: Freude (1967: 223).

Polemiotus humeralis var. acuticauda Casey, 1907: 383. Synonymy: Freude (1967: 223).

Genus Schoenicus LeConte, 1866 [M]

Schoenicus LeConte, 1866b: 109. Type species: Schoenicus puberulus LeConte, 1866, monotypy.

Schoenicus puberulus LeConte, 1866USA (FL GA MD MS NC NJ NY SC)

Schoenicus puberulus LeConte, 1866b: 110.

Genus Tydeolus Champion, 1884 [M]

Tydeolus Champion, 1884: 37. Type species: Tydeolus atratus Champion, 1884, subsequent designation (Kirby 1885: 80).

Tydeolus atratus Champion, 1884MEX (PU)

Tydeolus atratus Champion, 1884: 37.

Tydeolus tibialis Champion, 1884: 37. Synonymy: Freude (1968: 122).

Tydeolus singularis Champion, 1884: 37. Synonymy: Freude (1968: 122).

Tribe Nyctoporini Lacordaire, 1859

Nyctoporides Lacordaire, 1859: 130. Type genus: Nyctoporis Eschscholtz, 1831.

Genus Nyctoporis Eschscholtz, 1831 [F]

Nyctoporis Eschscholtz, 1831: 10, 11. Type species: Nyctoporis cristata Eschscholtz, 1831, subsequent designation (Hope 1841: 124).

Emeax Pascoe, 1866: 450. Type species: Emeax sculpturatus Pascoe, 1866 (= Nyctoporis cristata Eschscholtz, 1831), monotypy. Synonymy: LeConte (1873: 334).

Enneacoides Fairmaire, 1881: 277. Type species: Enneacoides vinculiger Fairmaire, 1881 (= Nyctoporis carinata LeConte, 1851), monotypy. Synonymy: Gebien (1908a: 287).

Nyctoporis aequicollis Eschscholtz, 1831USA (CA)

Nyctoporis aequicollis Eschscholtz, 1831: 12.

Nyctoporis tetrica Casey, 1907: 510. New synonymy [RLA].

Nyctoporis maura Casey, 1907: 512. New synonymy [RLA].

Nyctoporis carinata LeConte, 1851USA (CA)

Nyctoporis carinata LeConte, 1851: 138.

Nyctoporis segnis Casey, 1907: 511. Synonymy: Blaisdell (1931: 43).

Enneacoides vinculiger Fairmaire, 1881: 277. Synonymy: Gebien (1910: 118)26.

Nyctoporis cristata Eschscholtz, 1831USA (CA)

Nyctoporis cristata Eschscholtz, 1831: 11.

Nyctoporis galeata LeConte, 1857: 49. Synonymy: Casey (1907b: 510).

Emeax sculpturatus Pascoe, 1866: 450. Synonymy: Carter (1914: 406).

Nyctoporis sponsa Casey, 1907USA (CA)

Nyctoporis sponsa Casey, 1907: 510.

Nyctoporis pullata Casey, 1907: 510. New synonymy [RLA].

Nyctoporis vandykei Blaisdell, 1931USA (CA)

Nyctoporus [sic!] vandykei Blaisdell, 1931: 41.

Tribe Stenosini Schaum, 1859

Tagénites Solier, 1834: 503. Type genus: Tagenia Latreille, 1802 (= Stenosis Herbst, 1799). Note. Use of younger name Stenosini conserved (Art. 40.2) (see Bouchard et al. 2005: 523).

Stenosidae Schaum, 1859: 66. Type genus: Stenosis Herbst, 1799.

Typhlusechini Casey, 1907: 281. Type genus: Typhlusechus Linell, 1897.

Araeoschizini Casey, 1907: 484. Type genus: Araeoschizus LeConte, 1851.

Genus Araeoschizus LeConte, 1851 [M]

Araeoschizus LeConte, 1851: 138. Type species: Araeoschizus costipennis LeConte, 1851, monotypy.

Araeoschizus aalbui Papp, 1981MEX (BS)

Araeoschizus aalbui Papp, 1981: 316.

Araeoschizus agustinus Papp, 1998MEX (BC)

Araeoschizus agustinus Papp, 1998: 90.

Araeoschizus airmeti Tanner, 1945USA (ID NV OR)

Araeoschizus airmeti Tanner, 1945: 125.

Araeoschizus alinae Dajoz, 1984USA (UT)

Araeoschizus alinae Dajoz, 1984: 246.

Araeoschizus andrewsi Papp, 1981USA (CA)

Araeoschizus andrewsi Papp, 1981: 318.

Araeoschizus antennatus antennatus Blaisdell, 1943MEX (BC)

Araeoschizus antennatus Blaisdell, 1943: 215.

Araeoschizus antennatus blaisdelli Papp, 1989MEX (BC)

Araeoschizus antennatus blaisdelli Papp, 1989: 338.

Araeoschizus antennatus clarki Papp, 1989MEX (BC)

Araeoschizus antennatus clarki Papp, 1989: 335.

Araeoschizus apachensis Papp, 1981USA (AZ)

Araeoschizus apachensis Papp, 1981: 367.

Araeoschizus arizonicus Dajoz, 1989USA (AZ NM)

Araeoschizus arizonicus Dajoz, 1989b: 33.

Araeoschizus armatus Horn, 1870USA (CA NV)

Araeoschizus armatus Horn, 1870: 275.

Araeoschizus blomi Papp, 1998MEX (BC)

Araeoschizus blomi Papp, 1998: 93.

Araeoschizus colossalis Papp, 1981USA (AZ)

Araeoschizus colossalis Papp, 1981: 346.

Araeoschizus costipennis LeConte, 1851USA (CA)

Araeoschizus costipennis LeConte, 1851: 138.

Araeoschizus decipiens Horn, 1890USA (AZ CO NM TX UT) MEX (CH DU SO)

Araeoschizus decipiens Horn, 1890: 342.

Araeoschizus dolenterus Papp, 1981MEX (PU)

Araeoschizus dolenterus Papp, 1981: 349.

Araeoschizus doyeni Papp, 1981USA (CA)

Araeoschizus doyeni Papp, 1981: 375.

Araeoschizus duplicatus Casey, 1907USA (WY)

Araeoschizus duplicatus Casey, 1907: 491.

Araeoschizus elegantulus Papp, 1981MEX (BS)

Araeoschizus elegantulus Papp, 1981: 325.

Araeoschizus exiguus Casey, 1907USA (CA)

Araeoschizus exiguus Casey, 1907: 487.

Araeoschizus expeditionis Papp, 1981MEX (DU)

Araeoschizus expeditionis Papp, 1981: 351.

Araeoschizus fimbriatus Casey, 1890USA (AZ)

Araeoschizus fimbriatus Casey, 1890b: 369.

Araeoschizus giulianii Papp, 1981MEX (SO)

Araeoschizus giulianii Papp, 1981: 393.

Araeoschizus hardyi Papp, 1981USA (CA)

Araeoschizus hardyi Papp, 1981: 308.

Araeoschizus hardyorum Papp, 1981USA (UT)

Araeoschizus hardyorum Papp, 1981: 395.

Araeoschizus hystrix Papp, 1981USA (CA)

Araeoschizus hystrix Papp, 1981: 330.

Araeoschizus interjectus Papp, 1981MEX (BS)

Araeoschizus interjectus Papp, 1981: 332.

Araeoschizus kaszabi Papp, 1981USA (CA)

Araeoschizus kaszabi Papp, 1981: 397.

Araeoschizus kubai Papp, 1981USA (AZ)

Araeoschizus kubai Papp, 1981: 400.

Araeoschizus lariversi Papp, 1981USA (CA)

Araeoschizus lariversi Papp, 1981: 306.

Araeoschizus lecontei Papp, 1981USA (AZ)

Araeoschizus lecontei Papp, 1981: 335.

Araeoschizus limbatus Blaisdell, 1943MEX (BS)

Araeoschizus limbatus Blaisdell, 1943: 214.

Araeoschizus magdae Papp, 1989MEX (GE)

Araeoschizus magdae Papp, 1989: 338.

Araeoschizus mexicanus Champion, 1892MEX (DU GE OA)

Araeoschizus mexicanus Champion, 1892: 491.

Araeoschizus microcephalus Papp, 1981MEX (CH)

Araeoschizus microcephalus Papp, 1981: 379.

Araeoschizus muthi Dajoz, 1998USA (CA)

Araeoschizus muthi Dajoz, 1998: 87.

Araeoschizus orientalis Dajoz, 1991USA (TX)

Araeoschizus orientalis Dajoz, 1991: 172.

Araeoschizus percellosus Papp, 1981MEX (BC)

Araeoschizus percellosus Papp, 1981: 312.

Araeoschizus problematicus Papp, 1981MEX (ZA)

Araeoschizus problematicus Papp, 1981: 402.

Araeoschizus regularis Horn, 1870USA (AZ UT) MEX (SO)

Araeoschizus regularis Horn, 1870: 274.

Araeoschizus rufus Dajoz, 1991USA (CA)

Araeoschizus rufus Dajoz, 1991: 165.

Araeoschizus setosiformis Papp, 1981USA (UT)

Araeoschizus setosiformis Papp, 1981: 382.

Araeoschizus similaris Papp, 1981USA (NM)

Araeoschizus similaris Papp, 1981: 384.

Araeoschizus simplex Casey, 1890USA (AZ NM TX) MEX (CH)

Araeoschizus simplex Casey, 1890b: 369.

Araeoschizus simulans Casey, 1907USA (CA)

Araeoschizus simulans Casey, 1907: 488.

Araeoschizus squamulissimus Papp, 1981MEX (BC)

Araeoschizus squamulissimus Papp, 1981: 340.

Araeoschizus sulcicollis disjunctus Papp, 1981USA (CA)

Araeoschizus sulcicollis disjunctus Papp, 1981: 364.

Araeoschizus sulcicollis sulcicollis Horn, 1870USA (CA NV)

Araeoschizus sulcicollis Horn, 1870: 274.

Araeoschizus tenuis Casey, 1907USA (AZ)

Araeoschizus tenuis Casey, 1907: 486.

Araeoschizus texanus Dajoz, 1989USA (TX)

Araeoschizus texanus Dajoz, 1989a: 149.

Araeoschizus utahensis Papp, 1981USA (UT)

Araeoschizus utahensis Papp, 1981: 389.

Araeoschizus wasbauerorum Papp, 1981MEX (SO)

Araeoschizus wasbauerorum Papp, 1981: 342.

Genus Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu and Smith, 2016 [M]

Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu and Smith, 2016: 56827. Type species: Rhypasma quisqueyanus Garrido and Varela, 2011, original designation.

Caribanosis quisqueyanus (Garrido and Varela, 2011) DOM

Rhypasma quisqueyanus Garrido and Varela, 2011: 32.

Genus Discopleurus Lacordaire, 1859 [M]

Pleurophorus Solier, 1851: 162 [junior homonym of Pleurophorus Mulsant, 1842]. Type species: Pleurophorus quadricollis Solier, 1851, monotypy.

Discopleurus Lacordaire, 1859: 105. Replacement name for Pleurophorus Solier, 1851.

Discopleurus mesoamericanus Aalbu and Andrews, 1996 HON CRI PAN

Discopleurus mesoamericanus Aalbu and Andrews, 1996: 27.

Genus Typhlusechus Linell, 1897 [M]

Typhlusechus Linell, 1897: 154. Type species: Typhlusechus singularis Linell, 1897, original designation.

Typhlusechus balsasensis Aalbu and Andrews, 1985MEX (MI)

Typhlusechus balsasensis Aalbu and Andrews, 1985: 4.

Typhlusechus chemehuevii Aalbu and Andrews, 1985USA (CA)

Typhlusechus chemehuevii Aalbu and Andrews, 1985: 3.

Typhlusechus ignotus Doyen, 1990MEX (JA)

Typhlusechus ignotus Doyen, 1990: 227.

Typhlusechus peninsularis Aalbu and Andrews, 1985MEX (BS)

Typhlusechus peninsularis Aalbu and Andrews, 1985: 5.

Typhlusechus singularis Linell, 1897USA (CA)

Typhlusechus singularis Linell, 1897: 155.

Typhlusechus spilmani Aalbu and Andrews, 1985USA (TX) MEX (DU)

Typhlusechus spilmani Aalbu and Andrews, 1985: 6.

Tribe Vacronini Gebien, 1910

Vacroninae Gebien, 1910a: 118. Type genus: Vacronus Casey, 1907 (= Alaephus Horn, 1870).

Genus Alaephus Horn, 1870 [M]

Alaephus Horn, 1870: 346. Type species: Alaephus pallidus Horn, 1870, monotypy.

Vacronus Casey, 1907: 501, 508. Type species: Vacronus tenuicornis Casey, 1907, original designation. Synonymy: Doyen and Lawrence (1979: 350).

Alaephus convergens Casey, 1924USA (UT)

Alaephus convergens Casey, 1924: 324.

Alaephus gracilicornis Casey, 1924USA (NM)

Alaephus gracilicornis Casey, 1924: 325.

Alaephus gracilis Fall, 1905USA (AZ)

Alaephus gracilis Fall, 1905: 276.

Alaephus longicornis Casey, 1924USA (CA)

Alaephus longicornis Casey, 1924: 325.

Alaephus macilentus Casey, 1891USA (AZ CA NM)

Alaephus macilentus Casey, 1891: 61.

Alaephus nitidipennis Fall, 1905: 275. Synonymy: Fall (1907b: 175).

Alaephus madarensis Casey, 1924USA (CA)

Alaephus madarensis Casey, 1924: 324.

Alaephus nevadensis Tanner, 1965USA (NV)

Alaephus nevadensis Tanner [in Tanner and Packtam], 1965: 39.

Alaephus pallidus Horn, 1870USA (CA)

Alaephus pallidus Horn, 1870: 346.

Alaephus puberulus Fall, 1907USA (AZ UT)

Alaephus puberulus Fall, 1907b: 175.

Alaephus quadricollis Casey, 1924USA (UT)

Alaephus quadricollis Casey, 1924: 326.

Alaephus tenuicornis (Casey, 1907)USA (CA)

Vacronus tenuicornis Casey, 1907: 508.

Genus Eupsophulus Cockerell, 1906 [M]

Eupsophus Horn, 1870: 347 [junior homonym of Eupsophus Fitzinger, 1843]. Type species: Eupsophus castaneus Horn, 1870, monotypy.

Eupsophulus Cockerell, 1906: 242. Replacement name for Eupsophus Horn, 1870.

Eupsophulus brevipennis Casey, 1924USA (AZ)

Eupsophulus brevipennis Casey, 1924: 323.

Eupsophulus castaneus (Horn, 1870)USA (AZ CA NV TX) MEX (BC CO SO)

Eupsophus castaneus Horn, 1870: 347.

Eupsophulus horni (Champion, 1885)MEX (BS)

Eupsophus horni Champion, 1885: 122.

Subfamily TENEBRIONINAE Latreille, 1802

Tenebrionites Latreille, 1802: 165. Type genus: Tenebrio Linnaeus, 1758.

Tribe Acropteronini Doyen, 1989

Acropteronini Doyen, 1989: 288. Type genus: Acropteron Perty, 1832.

Genus Acropteron Perty, 1832 [N]

Acropteron Perty, 1832: 64. Type species: Acropteron rufipes Perty, 1832, subsequent designation (Hope 1841: 133).

Arthroplatus Solier, 1851: 246. Type species: Arthroplatus pallipes Solier, 1851, monotypy. Synonymy: Mäklin (1862: 1).

Acropteron agriloides Mäklin, 1862MEX (CI GE OA TB VE) GUA

Acropteron agriloides Mäklin, 1862: 17.

Acropteron angulicolle Champion, 1886 NIC

Acropteron angulicolle Champion, 1886: 255.

Acropteron belti Champion, 1886 NIC PAN

Acropteron belti Champion, 1886: 253.

Acropteron brunneum Mäklin, 1862CRI / SA

Acropteron brunneum Mäklin, 1862: 15.

Acropteron calcaratum Champion, 1886 GUA

Acropteron calcaratum Champion, 1886: 255.

Acropteron chabrieri Fleutiaux and Sallé, 1890 LAN

Acropteron chabrieri Fleutiaux and Sallé, 1890: 429.

Acropteron laevipes Champion, 1886 NIC

Acropteron laevipes Champion, 1886: 257.

Acropteron langurioides Champion, 1886 PAN

Acropteron langurioides Champion, 1886: 254.

Acropteron longipenne Champion, 1886 GUA PAN

Acropteron longipenne Champion, 1886: 256.

Acropteron maklini Champion, 1886 PAN

Acropteron mäklini Champion, 1886: 254.

Acropteron mexicanum Champion, 1886MEX (VE)

Acropteron mexicanum Champion, 1886: 256.

Acropteron puncticolle Champion, 1886 PAN

Acropteron puncticolle Champion, 1886: 256.

Acropteron quadraticolle Champion, 1896 LAN

Acropteron quadraticolle Champion, 1896: 29.

Acropteron rugipes Champion, 1886 NIC

Acropteron rugipes Champion, 1886: 257.

Tribe Alphitobiini Reitter, 1917

Alphitobiini Reitter, 1917: 58. Type genus: Alphitobius Stephens, 1829.

Genus Alphitobius Stephens, 1829 [M]

Alphitobius Stephens, 1829: 19. Type species: Helops picipes Panzer, 1794 (= Opatrum laevigatum Fabricius, 1781), monotypy.

Heterophaga Dejean, 1834: 199. Type species: Tenebrio mauritanicus Fabricius, 1792 (= Opatrum laevigatum Fabricius, 1781), subsequent designation (Duponchel 1845: 601). Synonymy: Wollaston (1854: 498).

Cryptops Solier, 1851: 235. Type species: Cryptops ulomoides Solier, 1851 (= Tenebrio diaperinus Panzer, 1797), monotypy. Synonymy: Philippi (1887: 735).

Microphyes MacLeay, 1873: 286. Type species: Microphyes rufipes MacLeay, 1873 (= Opatrum laevigatum Fabricius, 1781), monotypy. Synonymy: Blair (1914: 486).

Alphitobius diaperinus (Panzer, 1797) [Fig. 15] CAN (AB BC MB NB NF NS ON PE QC SK) USA (FL GA IN MD MI NC NY OH SC SD VA WA WI) MEX (CH DU GU PU QR VE) NIC / BAHCUBCAYJAMHAIDOMPRILAN / SA – Adventive

Figure 15. 

Alphitobius diaperinus (Panzer, 1797). Scale bar = 1 mm.

Tenebrio diaperinus Panzer, 1797: 16.

Cryptops ulomoides Solier, 1851: 236. Synonymy: Schawaller and Grimm (2014: 174).

Alphitobius laevigatus (Fabricius, 1781)CAN (BC NB ON QC) USA (CT FL GA ID IN NY OH OR PA SC SD WA WI) MEX (BS CH CO DU NL TA VE YU) BELNICCRIPAN / CUBCAYJAMPRILAN / SA – Adventive

Opatrum laevigatum Fabricius, 1781: 90.

Helops piceus Olivier, 1793: 50. Synonymy: Blair (1914: 486).

Helops picipes Panzer, 1794: 4. Synonymy: Spilman (1966: 7).

Alphitobius piceus var. ruficolor Pic, 1925b: 11. Synonymy: Gebien (1940: 779).

Tribe Amarygmini Gistel, 1848

Amarygmiidae Gistel, 1848: [10]. Type genus: Amarygmus Dalman, 1823.

Mégacanthides Lacordaire, 1859: 467. Type genus: Megacantha Westwood, 1843.

Méracanthides Lacordaire, 1859: 464. Type genus: Meracantha Kirby, 1837.

Genus Cymatothes Dejean, 1834 [M]

Cymatothes Dejean, 1834 [30 June]: 208. Type species: Helops undatus Fabricius 1792 (= Erotylus nebulosus Fabricius, 1781), monotypy.

Physignathus Gistel, 1834 [23 September]: 22 [junior homonym of Physignathus Cuvier, 1829]. Type species: Helops undatus Fabricius, 1792 (= Erotylus nebulosus Fabricius, 1781), monotypy. Synonymy: Bousquet and Bouchard (2017a: 132).

Pyanisia Laporte, 1840: 235. Type species: Helops undatus Fabricius, 1792 (= Erotylus nebulosus Fabricius, 1781), subsequent designation (Lacordaire 1859: 476). Synonymy: Chevrolat (1847: 643).

Pyganisia Hope, 1841: 133. Type species: Helops undatus Fabricius, 1792 (= Erotylus nebulosus Fabricius, 1781), original designation. Synonymy: Chevrolat (1847: 643).

Cymatothes fumosus (Champion, 1887) GUA

Pyanisia fumosa Champion, 1887: 331.

Cymatothes laevis (Champion, 1893)MEX (GE)

Pyanisia laevis Champion, 1893a: 561.

Cymatothes longicollis (Champion, 1887) GUA

Pyanisia longicollis Champion, 1887: 331.

Cymatothes nebulosus nebulosus (Fabricius, 1781)MEXGUABELNICCRIPAN / CUBHAILAN / SA

Erotylus nebulosus Fabricius, 1781: 158.

Helops undatus Fabricius, 1792a: 122. Synonymy: Blair (1935: 103).

Cymatothes opacus Solier, 1848USA (FL) MEX (CH CI DU GE OA PU VE)

Cymatothes opacus Solier, 1848: 180.

Cymatothes coarctatus Solier, 1848: 181. Synonymy: Champion (1887: 330).

Cymatothes unicolor Solier, 1848USA (AL FL TX) MEX (CI DU JA NA VE) GUABELNICCRIPAN / BAHCUBPRILAN

Helops tristis Laporte, 1840: 236 [junior primary homonym of Helops tristis Rossi, 1790].

Cymatothes unicolor Solier, 1848: 182. Synonymy: Champion (1887: 330).

Cymatothes uniformis (C.O. Waterhouse, 1878) JAM

Hoplonyx uniformis C.O. Waterhouse, 1878: 306.

Genus Meracantha Kirby, 1837 [F]

Meracantha Kirby, 1837: 237. Type species: Meracantha canadensis Kirby, 1837 (= Helops contractus Palisot de Beauvois, 1812), monotypy.

Falacer Laporte, 1840: 233. Type species: Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812), present designation. Synonymy: Lacordaire (1859: 466).

Physocoelus Haldeman, 1850: 5. Type species: Helops contractus Palisot de Beauvois, 1812, monotypy. Synonymy: Lacordaire (1859: 466).

Meracantha contracta (Palisot de Beauvois, 1812)CAN (ON) USA (AL CT FL GA IA IN LA MD MI NC NY OH PA SC TN TX VA WI)

Helops contractus Palisot de Beauvois, 1812: 121.

Meracantha canadensis Kirby, 1837: 238. Synonymy: Melsheimer (1853: 140).

Acanthopus cupreus Laporte, 1840: 233. Synonymy: Lacordaire (1859: 466).

Acanthopus rugosus Laporte, 1840: 233. Synonym (in doubt): Papp (1961d: 135).

Helops tumidus Melsheimer, 1846: 61. Synonymy: Melsheimer (1853: 140).

Psorodes inflata Solier, 1848: 167. Synonymy (with M. canadensis Kirby): Schaum (1850: 181).

Genus Plesiophthalmus Motschulsky, 1857 [M]

Plesiophthalmus Motschulsky, 1857: 34. Type species: Plesiophthalmus nigrocyaneus Motschulsky, 1857, monotypy.

Cyriogeton Pascoe, 1871: 356. Type species: Cyriogeton insignis Pascoe, 1871, monotypy. Synonymy: Masumoto (1989a: 536).

Plesiophthalmus spectabilis Harold, 1875USA (MD)28 – Adventive

Plesiophthalmus spectabilis Harold, 1875: 293.

Plesiophthalmus obesus Marseul, 1876: 319. Synonymy: Harold (1876: 85).

Plesiophthalmus arciferens Fairmaire [in Deyrolle and Fairmaire], 1878: 120. Synonymy: Masumoto (1989b: 743).

Plesiophthalmus subparallelus Pic, 1916: 11. Synonymy: Masumoto (1989b: 743).

Tribe Amphidorini LeConte, 1862

Amphidorae LeConte, 1862a: 239. Type genus: Amphidora Eschscholtz, 1829.

Eleodiini Blaisdell, 1909: 27. Type genus: Eleodes Eschscholtz, 1829.

Eleodopsinae Blaisdell, 1939b: 51. Type genus: Eleodopsis Blaisdell, 1939 (= Eleodes Eschscholtz, 1829).

Lariversiina La Rivers, 1948: 98. Type genus: Lariversius Blaisdell, 1947.

Trogloderina La Rivers, 1948: 98. Type genus: Trogloderus LeConte, 1879.

Genus Eleodes Eschscholtz, 182929 [F]

Eleodes Eschscholtz, 1829: 8. Type species: Eleodes dentipes Eschscholtz, 1829, subsequent designation (Hope 1841: 124).

Elaeodes Gemminger [in Gemminger and Harold], 1870: 1868. Unjustified emendation of Eleodes Eschscholtz, 1829, not in prevailing usage.

Subgenus Amphidora Eschscholtz, 1829

Amphidora Eschscholtz, 1829: 9. Type species: Amphidora littoralis Eschscholtz, 1829, monotypy. Status revised [ADS & MAJ].

Eleodes littoralis (Eschscholtz, 1829)USA (CA)

Amphidora littoralis Eschscholtz, 1829: 930.

Eleodes nigropilosa (LeConte, 1851)USA (CA) MEX (BC)

Amphidora nigropilosa LeConte, 1851: 136.

Amphidora tenebrosa Horn, 1870: 329. Synonymy: Triplehorn (1996: 13).

Eleodes subdeplanata (Blaisdell, 1943)MEX (BC BS)

Amphidora subdeplanata Blaisdell, 1943: 252.

Subgenus Ardeleodes Blaisdell, 1937

Ardeleodes Blaisdell, 1937b: 12831. Type species: Eleodes tibialis Blaisdell, 1909, original designation.

Eleodes tibialis Blaisdell, 1909MEX (BS)

Eleodes tibialis Blaisdell, 1909: 313.

Eleodes tibialis forma oblonga Blaisdell, 1909: 315. New synonymy [YB].

Subgenus Blapylis Horn, 1870

Blapylis Horn, 1870: 315. Type species: Eleodes cordata Eschscholtz, 1829, present designation. Synonymy: Doyen and Lawrence (1979: 367).

Eleodopsis Blaisdell, 1939b: 52. Type species: Eleodopsis subvestita Blaisdell, 1939, original designation. Synonymy: Spilman (1962b: 59).

Eleodes alticola Blaisdell, 1925USA (CA)

Eleodes parvicollis alticola Blaisdell, 1925c: 387.

Eleodes aristata Somerby, 1977USA (CA)

Eleodes aristata Somerby, 1977: 22.

Eleodes bishopensis Somerby and Doyen, 1976USA (CA)

Eleodes bishopensis Somerby and Doyen, 1976: 257.

Eleodes blanchardii Blaisdell, 1909USA (CA)

Eleodes blanchardii Blaisdell, 1909: 339.

Eleodes brunnipes Casey, 1890USA (CA CO ID NV WY)

Eleodes brunnipes Casey, 1890b: 402.

Eleodes pimelioides var. brevisetosa Blaisdell, 1918b: 162. Synonymy: Blaisdell (1925a: 80).

Eleodes caseyi Blaisdell, 1909USA (CA NV)

Eleodes caseyi Blaisdell, 1909: 388.

Eleodes clavicornis Eschscholtz, 1829USA (CA)

Eleodes clavicornis Eschscholtz, 1829: 11.

Eleodes impressicollis Boheman, 1858: 90. Synonymy: LeConte (1866a: 60).

Eleodes consobrina LeConte, 1851USA (CA) MEX (BC)

Eleodes consobrina LeConte, 1851: 135.

Eleodes veseyi LeConte, 1858c: 187. Synonymy: LeConte (1866a: 60).

Eleodes constricta LeConte, 1858CAN (BC) USA (CA UT)

Eleodes constrictus LeConte, 1858c: 187.

Eleodes [manni var.] variolosa Blaisdell, 1917: 223. New synonymy [based on Somerby (1972: 161) unpublished thesis].

Eleodes cooperi Somerby and Doyen, 1976USA (CA)

Eleodes cooperi Somerby and Doyen, 1976: 253.

Eleodes cordata Eschscholtz, 1829USA (CA)

Eleodes cordata Eschscholtz, 1829: 12.

Eleodes cordata forma sublaevis Blaisdell, 1909: 381. New synonymy [based on Somerby (1972: 254) unpublished thesis].

Eleodes cordata forma intermedia Blaisdell, 1909: 381. New synonymy [YB].

Eleodes cordata forma oblonga Blaisdell, 1909: 383. New synonymy [YB].

Eleodes cordata forma elongata Blaisdell, 1909: 383. New synonymy [YB].

Eleodes [cordata var.] adulterina Blaisdell, 1917: 224. New synonymy [based on Somerby (1972: 254) unpublished thesis].

Eleodes fuchsii Blaisdell, 1909USA (CA)

Eleodes fuchsii Blaisdell, 1909: 343.

Eleodes hornii var. monticula Blaisdell, 1918c: 385. New synonymy [based on Somerby (1972: 199) unpublished thesis].

Eleodes manni sierra Blaisdell, 1926a: 78. New synonymy [based on Somerby (1972: 199) unpublished thesis].

Eleodes hoppingii Blaisdell, 1909USA (CA)

Eleodes hoppingii Blaisdell, 1909: 368.

Eleodes hornii hornii Blaisdell, 1909USA (CA)

Eleodes hornii Blaisdell, 1909: 350.

Eleodes hybrida Blaisdell, 1917USA (CA)

Eleodes [cordata var.] hybrida Blaisdell, 1917: 225.

Eleodes inculta LeConte, 1861USA (CA)

Eleodes inculta LeConte, 1861b: 352.

Eleodes inculta var. affinis Blaisdell, 1918c: 384. Synonymy: Miller (1985: 21).

Eleodes kaweana Blaisdell, 1933USA (CA)

Eleodes kaweana Blaisdell, 1933b: 203.

Eleodes lariversi Somerby and Doyen, 1976USA (CA)

Eleodes lariversi Somerby and Doyen, 1976: 256.

Eleodes lecontei Horn, 1870USA (CO)

Eleodes subaspera LeConte, 1866b: 115 [junior primary homonym of Eleodes subaspera Solier, 1848].

Eleodes lecontei Horn, 1870: 316. Replacement name for Eleodes subaspera LeConte, 1866.

Eleodes manni Blaisdell, 1917USA (WA)

Eleodes manni Blaisdell, 1917: 221.

Eleodes nana Blaisdell, 1909USA (CA NV)

Eleodes tenebrosa var. nana Blaisdell, 1909: 328.

Eleodes neotomae Blaisdell, 1909USA (CA)

Eleodes neotomae Blaisdell, 1909: 347.

Eleodes novoverrucula Boddy, 1957CAN (AB BC) USA (ID MT OR WA)

Eleodes novoverrucula Boddy, 1957: 195.

Eleodes nunenmacheri Blaisdell, 1918USA (CA OR)

Eleodes nunenmacheri Blaisdell, 1918b: 163.

Eleodes oregona Blaisdell, 1941USA (OR)

Eleodes oregona Blaisdell, 1941b: 157.

Eleodes orophila Somerby, 1977USA (AZ NM)

Eleodes orophilus Somerby, 1977: 24.

Eleodes panamintensis Somerby, 1977USA (CA)

Eleodes panamintensis Somerby, 1977: 20.

Eleodes parvicollis Eschscholtz, 1829USA (CA)

Eleodes parvicollis Eschscholtz, 1829: 11.

Eleodes parvicollis var. squalida Blaisdell, 1918c: 380. New synonymy [based on Somerby (1972: 188) unpublished thesis].

Eleodes patulicollis Blaisdell, 1932USA (UT)

Eleodes manni dilaticollis Blaisdell, 1925c: 388 [junior primary homonym of Eleodes dilaticollis Champion, 1884].

Eleodes patulicollis Blaisdell, 1932a: 78. Replacement name for Eleodes dilaticollis Blaisdell, 1925.

Eleodes pimelioides Mannerheim, 1843 [Fig. 16] CAN (BC) USA (CA OR UT WA)

Figure 16. 

Eleodes (Blapylis) pimelioides Mannerheim, 1843. Scale bar = 1 mm.

Eleodes pimelioides Mannerheim, 1843: 274.

Eleodes viator LeConte, 1858c: 188. Synonymy: Horn (1870: 318).

Eleodes nunenmacheri var. verrucula Blaisdell, 1918b: 164. Synonymy: Bousquet and Campbell (1991: 257) [based on Somerby (1972: 238) unpublished thesis].

Eleodes pimelioides var. patruelis Blaisdell, 1918c: 382. Synonymy (with E. nunenmacheri verrucula Blaisdell): Boddy (1957: 197).

Eleodes planata Eschscholtz, 1829USA (CA)

Eleodes planata Eschscholtz, 1829: 12.

Eleodes reflexicollis Mannerheim, 1843: 270. New synonymy [based on Somerby (1972: 187) unpublished thesis].

Eleodes parvicollis forma farallonica Blaisdell, 1909: 356. New synonymy [based on Somerby (1972: 187) unpublished thesis].

Eleodes producta Mannerheim, 1843USA (CA)

Eleodes producta Mannerheim, 1843: 271.

Eleodes propinqua Blaisdell, 1918USA (CA)

Eleodes propinqua Blaisdell, 1918b: 165.

Eleodes robinetti Boddy, 1957USA (OR WA)

Eleodes robinetti Boddy, 1957: 194.

Eleodes rotundipennis LeConte, 1857CAN (BC) USA (OR WA)

Eleodes rotundipennis LeConte, 1857: 50.

Eleodes stricta LeConte, 1857: 50. Synonymy: Boddy (1965: 158).

Eleodes subligata LeConte, 1857: 50. Synonymy: Boddy (1965: 158).

Eleodes indentata Blaisdell, 1935a: 28. New synonymy [based on Somerby (1972: 237) unpublished thesis].

Eleodes scabripennis LeConte, 1859USA (CA)

Eleodes scabripennis LeConte, 1859b: 77.

Eleodes scabriventris Blaisdell, 1933USA (CA)

Eleodes scabriventris Blaisdell, 1933b: 202.

Eleodes scabrosa Eschscholtz, 1829USA (CA OR)

Eleodes scabrosa Eschscholtz, 1829: 11.

Eleodes intricata Mannerheim, 1843: 273. New synonymy [based on Somerby (1972: 163) unpublished thesis].

Eleodes schlingeri Somerby and Doyen, 1976USA (CA)

Eleodes schlingeri Somerby and Doyen, 1976: 254.

Eleodes schwarzii Blaisdell, 1909USA (ID OR WA)

Eleodes schwarzii Blaisdell, 1909: 406.

Eleodes snowii Blaisdell, 1909USA (AZ CO NM NV)

Eleodes snowii Blaisdell, 1909: 317.

Eleodes spilmani Somerby and Doyen, 1976USA (CA)

Eleodes spilmani Somerby and Doyen, 1976: 258.

Eleodes strumosa Blaisdell, 1932USA (UT NV)

Eleodes strumosa Blaisdell, 1932a: 76.

Eleodes subvestita (Blaisdell, 1939)USA (CA)

Eleodopsis subvestita Blaisdell, 1939b: 53.

Eleodes tenebrosa Horn, 1870CAN (BC) USA (CA NV UT)

Eleodes tenebrosa Horn, 1870: 316.

Eleodes horni fenyesi Blaisdell, 1926a: 77. New synonymy [based on Somerby (1972: 161) unpublished thesis].

Eleodes triplehorni Somerby and Doyen, 1976MEX (BC)

Eleodes triplehorni Somerby and Doyen, 1976: 252.

Eleodes trita Blaisdell, 1917USA (CA OR)

Eleodes [parvicollis var.] trita Blaisdell, 1917: 225.

Eleodes tuberculata Eschscholtz, 1829USA (CA)

Eleodes tuberculata Eschscholtz, 1829: 12.

Eleodes cordata var. horrida Blaisdell, 1918c: 383. New synonymy [based on Somerby (1972: 255) unpublished thesis].

Eleodes versatilis Blaisdell, 1921USA (OR WA)

Eleodes rotundipennis versatilis Blaisdell, 1921b: 217.

Eleodes oblonga Blaisdell, 1933b: 206 [junior primary homonym of Eleodes tibialis oblonga Blaisdell, 1909]. New synonymy [based on Somerby (1972: 190) unpublished thesis].

Eleodes formosus Thomas, 2005: 551. Replacement name for Eleodes oblonga Blaisdell, 1933.

Eleodes volcanensis Somerby, 1977USA (CA OR)

Eleodes volcanensis Somerby, 1977: 23.

Eleodes wakelandi Somerby, 1977USA (ID OR UT)

Eleodes wakelandi Somerby, 1977: 19.

Subgenus Caverneleodes Triplehorn, 1975

Caverneleodes Triplehorn, 1975: 39. Type species: Eleodes easterlai Triplehorn, 1975, original designation.

Eleodes easterlai Triplehorn, 1975USA (TX) MEX (CO)

Eleodes easterlai Triplehorn, 1975: 39.

Eleodes grutus Aalbu, Smith and Triplehorn, 201232MEX (NL)

Eleodes grutus Aalbu, Smith and Triplehorn, 2012: 206.

Eleodes guadalupensis Aalbu, Smith and Triplehorn, 2012USA (NM)

Eleodes guadalupensis Aalbu, Smith and Triplehorn, 2012: 208.

Eleodes labialis Triplehorn, 1975USA (TX) MEX (CH)

Eleodes labialis Triplehorn, 1975: 42.

Eleodes leptoscelis Triplehorn, 1975USA (AZ)

Eleodes leptoscelis Triplehorn, 1975: 42.

Eleodes microps Aalbu, Smith and Triplehorn, 2012USA (CA)

Eleodes microps Aalbu, Smith and Triplehorn, 2012: 200.

Eleodes reddelli Triplehorn, 2007MEX (NL)

Eleodes reddelli Triplehorn, 2007: 639.

Eleodes rugosifrons Triplehorn and Reddell, 1991MEX (CO NL)

Eleodes rugosifrons Triplehorn and Reddell, 1991: 527.

Eleodes sprousei Triplehorn and Reddell, 1991MEX (NL TA)

Eleodes sprousei Triplehorn and Reddell, 1991: 525.

Eleodes thomasi Aalbu, Smith and Triplehorn, 2012MEX (CO NL)

Eleodes thomasi Aalbu, Smith and Triplehorn, 2012: 204.

Eleodes wheeleri Aalbu, Smith and Triplehorn, 2012USA (AZ)

Eleodes wheeleri Aalbu, Smith and Triplehorn, 2012: 208.

Eleodes wynnei Aalbu, Smith and Triplehorn, 2012USA (AZ UT)

Eleodes wynnei Aalbu, Smith and Triplehorn, 2012: 201.

Subgenus Chaseleodes Thomas, 2015

Chaseleodes Thomas, 2015: 122. Type species: Elaeodes curta Champion, 1884, original designation.

Eleodes connata Solier, 1848MEX (CH DU FD ME MI MO PU TL VE)

Eleodes connata Solier, 1848: 243.

Eleodes curta Champion, 1884MEX (JA ME MI)

Elaeodes curta Champion, 1884: 82.

Subgenus Cratidus LeConte, 1862

Cratidus LeConte, 1862a: 239. Type species: Amphidora osculans LeConte, 1851, monotypy.

Eleodes osculans (LeConte, 1851)USA (CA) MEX (BC)

Amphidora osculans LeConte, 1851: 136.

Cratidus fuscipilosus Casey, 1890b: 407. Synonymy: Triplehorn (1996: 13).

Cratidus ovipennis Casey, 1924: 328. Synonymy: Triplehorn (1996: 13).

Eleodes behrii Grinnell, 1908: 213. Synonymy: Doyen and Miller (1980: 3).

Eleodes intermedia Grinnell, 1908: 215. Synonymy: Doyen and Miller (1980: 3).

Eleodes ursus Triplehorn, 1996MEX (BC BS)

Cratidus rotundicollis Horn, 1870: 328 [junior secondary homonym of Eleodes rotundicollis Eschscholtz, 1829].

Eleodes ursus Triplehorn, 1996: 14. Replacement name for Eleodes rotundicollis (Horn, 1870).

Subgenus Discogenia LeConte, 1866

Discogenia LeConte, 1866b: 117. Type species: Eleodes scabricula LeConte, 1858, present designation.

Eleodes acutangula Blaisdell, 1921USA (CA)

Eleodes acutangula Blaisdell, 1921b: 225.

Eleodes marginata Eschscholtz, 1829USA (CA)

Eleodes marginata Eschscholtz, 1829: 10.

Eleodes fischerii Mannerheim, 1840: 137. Synonymy: Horn (1870: 320).

Eleodes scabricula deplanata Blaisdell, 1909USA (CA)

Eleodes scabricula forma deplanata Blaisdell, 1909: 444.

Eleodes scabricula scabricula LeConte, 1858USA (CA)

Eleodes scabricula LeConte, 1858c: 187.

Subgenus Eleodes Eschscholtz, 1829

Eleodes Eschscholtz, 1829: 8. Type species: Eleodes dentipes Eschscholtz, 1829, subsequent designation (Hope 1841: 124).

Eleodes acuticauda LeConte, 1851USA (CA) MEX (BC)

Eleodes acuticauda LeConte, 1851: 135.

Eleodes laticollis LeConte, 1851: 135. Synonymy: Horn (1870: 314).

Eleodes acuticauda forma punctata Blaisdell, 1909: 278. Synonymy: Triplehorn et al. (2015: 161).

Eleodes laticollis apprima Blaisdell, 1921b: 219. Synonymy: Triplehorn (1996: 9).

Eleodes acuta (Say, 1824)USA (CO KS NM SD TX)

Blaps acuta Say, 1824a: 258.

Eleodes acuta pernigra Blaisdell, 1937b: 128. Synonymy: Triplehorn et al. (2015: 161).

Eleodes adumbrata Blaisdell, 1925MEX (BC)

Eleodes adumbrata Blaisdell, 1925b: 332.

Eleodes armata LeConte, 1851USA (AZ CA ID NV OR UT) MEX (BC SO)

Eleodes armata LeConte, 1851: 134.

Eleodes armata impotens Blaisdell, 1895: 236. Synonymy: Triplehorn et al. (2015: 163).

Eleodes armata forma sinuata Blaisdell, 1909: 266. Synonymy: Triplehorn et al. (2015: 163).

Eleodes armata var. pumila Blaisdell, 1933b: 197. Synonymy: Triplehorn et al. (2015: 163).

Eleodes amedeensis Blaisdell, 1933b: 199. Synonymy: Triplehorn et al. (2015: 163).

Eleodes striatipennis Blaisdell, 1942: 134. Synonymy: Triplehorn et al. (2015: 163).

Eleodes curvidens Triplehorn and Cifuentes-Ruiz, 2011MEX (GU MI MO PU)

Eleodes curvidens Triplehorn and Cifuentes-Ruiz, 2011: 66.

Eleodes dentipes Eschscholtz, 1829USA (CA)

Eleodes dentipes Eschscholtz, 1829: 10.

Eleodes elegans Casey, 1890b: 401. Synonymy: Blaisdell (1909: 251).

Eleodes prominens Casey, 1890b: 401. Synonymy: Blaisdell (1909: 251).

Eleodes confinis Blaisdell, 1895: 237. Synonymy: Blaisdell (1909: 251).

Eleodes dentipes forma pertenuis Blaisdell, 1909: 253. Synonymy: Triplehorn et al. (2015: 164).

Eleodes dentipes forma elongata Blaisdell, 1909: 254. Synonymy: Gebien (1938: 57).

Eleodes dentipes forma robusta Blaisdell, 1909: 255. Synonymy: Triplehorn et al. (2015: 164).

Eleodes dentipes var. perpunctata Blaisdell, 1918c: 386. Synonymy: Gebien (1938: 57).

Eleodes dentipes marinae Blaisdell, 1921b: 218. New synonymy [ADS & MAJ].

Eleodes dentipes montana Blaisdell, 1925c: 385 [junior primary homonym of Eleodes montana Champion, 1884]. Synonymy: Triplehorn et al. (2015: 164).

Eleodes dentipes tularensis Blaisdell, 1925c: 386. Synonymy: Triplehorn et al. (2015: 164).

Eleodes paradoxa Blaisdell, 1932a: 78. Replacement name for Eleodes montana Blaisdell, 1925.

Eleodes dentipes sordida Blaisdell, 1935a: 30. Synonymy: Triplehorn et al. (2015: 164).

Eleodes discincta Blaisdell, 1925MEX (BC BS)

Eleodes discincta Blaisdell, 1925b: 333.

Eleodes eschscholtzii Solier, 1848USA (AZ NM) MEX (BS CO DU SI SO)

Eleodes eschscholtzii Solier, 1848: 254.

Eleodes lucae LeConte, 1866b: 114. Synonymy: Triplehorn (1996: 10).

Eleodes wickhami Horn, 1891: 41. Synonymy: Triplehorn et al. (2015: 167).

Eleodes femorata LeConte, 1851USA (CA) MEX (BC BS)

Eleodes femorata LeConte, 1851: 134.

Eleodes militaris Horn, 1870: 310. Synonymy: Triplehorn (1996: 7).

Eleodes militaris forma subedentata Blaisdell, 1909: 270. Synonymy: Triplehorn et al. (2015: 167).

Eleodes inepta Blaisdell, 1925b: 334. Synonymy: Triplehorn (1996: 7).

Eleodes marthae Blaisdell, 1943: 243. Synonymy: Triplehorn (1996: 7).

Eleodes fiski Triplehorn, 2015MEX (TA)

Eleodes fiski Triplehorn [in Triplehorn, Thomas and Smith], 2015: 170.

Eleodes gracilis distans Blaisdell, 1909USA (CA)

Eleodes gracilis var. distans Blaisdell, 1909: 242.

Eleodes gracilis gracilis LeConte, 1858USA (AZ NM TX) MEX (CH CO DU SI SO ZA)

Eleodes gracilis LeConte, 1858c: 184.

Eleodes grandicollis grandicollis Mannerheim, 1843USA (AZ CA NV) MEX (BC)

Eleodes grandicollis Mannerheim, 1843: 266.

Eleodes elongata Grinnell, 1908: 215. Synonymy: Doyen and Miller (1980: 3).

Eleodes grandicollis valida Boheman, 1858USA (CA)

Eleodes valida Boheman, 1858: 90.

Eleodes hispilabris (Say, 1824)CAN (AB MB SK) USA (AZ CA CO ID KS MT ND NE NM NV OK OR SD TX UT WA WY) MEX (CH CO NL SO TA)

Blaps hispilabris Say, 1824a: 259.

Eleodes sulcata LeConte, 1852: 67 [junior primary homonym of Eleodes sulcatus Eschscholtz, 1829]. Synonymy: Horn (1870: 313).

Eleodes connexa LeConte, 1857: 49. Synonymy: Triplehorn et al. (2015: 174).

Eleodes nupta LeConte, 1858c: 183. Synonymy: Triplehorn et al. (2015: 174).

Eleodes binotata Walker, 1866: 329. Synonymy: Triplehorn et al. (2015: 174).

Elaeodes lecontei Gemminger, 1870: 122. Replacement name for Elaeodes sulcata LeConte, 1852.

Eleodes hispilabris forma sculptilis Blaisdell, 1909: 220. Synonymy: Triplehorn et al. (2015: 174).

Eleodes hispilabris forma elongata Blaisdell, 1909: 220 [primary homonym of Eleodes dentipes forma elongata Blaisdell, 1909]. Synonymy: Triplehorn et al. (2015: 174).

Eleodes hispilabris forma laevis Blaisdell, 1909: 220. Synonymy (with E. binotata Walker): Blair (1921: 283).

Eleodes subpinguis Blaisdell, 1909: 247. Synonymy: Triplehorn et al. (2015: 174).

Eleodes hispilabris var. imitabilis Blaisdell, 1918b: 167. Synonymy: Triplehorn et al. (2015: 174).

Eleodes hispilabris var. attenuata Blaisdell, 1918b: 168 [junior secondary homonym of Eleodes attenuatus (LeConte, 1851)]. Synonymy: Triplehorn et al. (2015: 174).

Eleodes hispilabris immundus Blaisdell, 1925a: 79. Replacement name for Eleodes hispilabris elongata Blaisdell, 1909.

Eleodes loretensis Blaisdell, 1923MEX (BC BS)

Eleodes loretensis Blaisdell, 1923: 262.

Eleodes mexicana Blaisdell, 1943MEX (BC BS)

Eleodes mexicana Blaisdell, 1943: 246.

Eleodes simondsi Blaisdell, 1943: 247. Synonymy: Triplehorn (1996: 11).

Eleodes blaisdelli Blackwelder, 1945: 521. Unnecessary replacement name for Eleodes mexicana Blaisdell, 1943.

Eleodes mirabilis Triplehorn, 2007USA (TX) MEX (NL SL TA)

Eleodes mirabilis Triplehorn, 2007: 634.

Eleodes moesta Blaisdell, 1921MEX (BC BS)

Eleodes sanmartinensis moesta Blaisdell, 1921b: 221.

Eleodes morbosa Blaisdell, 1925b: 335. Synonymy: Triplehorn (1996: 11).

Eleodes muricatula Triplehorn, 2007MEX (CO SL ZA)

Eleodes muricatulus Triplehorn, 2007: 637.

Eleodes obscura dispersa LeConte, 1858USA (AZ CO NM UT)

Eleodes dispersa LeConte, 1858c: 182.

Eleodes deleta LeConte, 1858c: 182. Synonymy: Horn (1870: 305).

Eleodes obscura glabriuscula Blaisdell, 1925USA (AZ NM TX) MEX (CH)

Eleodes obscura glabriuscula Blaisdell, 1925c: 383.

Eleodes obscura obscura (Say, 1824)USA (CO MT NE NM TX WY)

Blaps obscura Say, 1824a: 259.

Eleodes obscura sulcipennis Mannerheim, 1843 [Fig. 17] CAN (BC) USA (AZ CA ID MT NM NV OR UT WA) MEX (CH DU SO)

Figure 17. 

Eleodes (Eleodes) obscura sulcipennis Mannerheim, 1843. Scale bar = 1 mm.

Eleodes sulcipennis Mannerheim, 1843: 266.

Eleodes arata LeConte, 1858c: 182. Synonymy: Horn (1870: 306).

Eleodes conjuncta Walker, 1866: 329. Synonymy: Leng (1920: 227).

Eleodes convexicollis Walker, 1866: 328. Synonymy: Leng (1920: 227).

Eleodes rossi Blaisdell, 1943MEX (BS)

Eleodes rossi Blaisdell, 1943: 241.

Eleodes rugosa Perbosc, 1839MEX (SL TA VE)

Eleodes rugosa Perbosc, 1839: 263.

Eleodes caudata Solier, 1848: 255. Synonymy: Champion (1884: 77).

Eleodes samalayucae Triplehorn, 2007MEX (CH)

Eleodes samalayucae Triplehorn, 2007: 641.

Eleodes sanmartinensis Blaisdell, 1921MEX (BC)

Eleodes sanmartinensis Blaisdell, 1921b: 220.

Eleodes scyroptera Triplehorn, 2007MEX (AG DU GU HI NL QU ZA)

Eleodes scyropterus Triplehorn, 2007: 635.

Eleodes spinipes macrura Champion, 1892USA (AZ NM TX) MEX (AG CH CO DU JA NA SO ZA)

Elaeodes macrura Champion, 1892: 511.

Eleodes ventricosa var. falli Blaisdell, 1909: 305. Synonymy: Triplehorn (2010: 377).

Eleodes spinipes spinipes Solier, 1848MEX (GU HI NL QU SL TA)

Eleodes spinipes Solier, 1848: 253.

Eleodes spinipes ventricosa LeConte, 1858USA (TX) MEX (CO NL TA)

Eleodes ventricosa LeConte, 1858c: 186.

Eleodes sponsa LeConte, 1858USA (AZ CO NM TX UT)

Eleodes sponsa LeConte, 1858c: 184.

Eleodes sponsa forma convexa Blaisdell, 1909: 215. Synonymy: Triplehorn et al. (2015: 187).

Eleodes subcylindrica Casey, 1890USA (AZ CA NV) MEX (BC)

Eleodes subcylindricus Casey, 1890b: 400.

Eleodes armata forma subedentata Blaisdell, 1909: 262. Synonymy: Blaisdell (1910: 62).

Eleodes suturalis (Say, 1824)USA (AZ CO KS MN ND NE NM OK SD TX UT WY)

Blaps suturalis Say, 1824a: 257.

Eleodes texana LeConte, 1858c: 182. Synonymy: Triplehorn et al. (2009: 432).

Eleodes tenuipes Casey, 1890USA (NM TX) MEX (CH)

Eleodes tenuipes Casey, 1890b: 399.

Eleodes vanduzeei Blaisdell, 1923MEX (BS)

Eleodes vanduzeei Blaisdell, 1923: 264.

Subgenus Heteropromus Blaisdell, 1909

Heteropromus Blaisdell, 1909: 179. Type species: Eleodes veterator Horn, 1874, monotypy.

Eleodes veterator Horn, 187433USA (LA TX)

Eleodes vetorator Horn, 1874a: 33.

Subgenus Litheleodes Blaisdell, 1909

Litheleodes Blaisdell, 1909: 114. Type species: Blaps extricata Say, 1824, subsequent designation (Triplehorn and Thomas 2015: 11).

Eleodes arcuata Casey, 1884USA (AZ NM TX) MEX (CH CO SO)

Eleodes arcuata Casey, 1884 [August]: 47.

Elaeodes sonorae Champion, 1884 [December]: 85. Synonymy: Gebien (1938: 53).

Eleodes aspera LeConte, 1866USA (AZ CO UT)

Eleodes aspera LeConte, 1866b: 115.

Eleodes corvina Blaisdell, 1921USA (CA OR)

Eleodes corvina Blaisdell, 1921b: 224.

Eleodes extricata (Say, 1824) [Fig. 18] CAN (AB BC SK) USA (AZ CO ID KS MT ND NE NM NV OK OR SD TX UT WY) MEX (CH CO SO)

Figure 18. 

Eleodes (Litheleodes) extricata (Say, 1824). Scale bar = 1 mm.

Blaps extricata Say, 1824a: 261.

Eleodes cognata Haldeman, 1852: 376. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricata var. arizonensis Blaisdell, 1909: 125. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricata forma elongata Blaisdell, 1909: 123. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricata forma convexicollis Blaisdell, 1909: 123 [junior primary homonym of Eleodes convexicollis Walker, 1866]. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricata var. utahensis Blaisdell, 1921a: 131. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricata frigida La Rivers, 1943b: 54. Synonymy: Triplehorn and Thomas (2015: 16).

Eleodes extricatus convexinotus Thomas, 2005: 551. Replacement name for Eleodes extricatus convexicollis Blaisdell, 1909.

Eleodes granulata LeConte, 1857CAN (BC) USA (CA CO ID NM OR WA)

Eleodes subaspera Solier, 1848: 246 [nomen dubium, see Horn (1870: 309)].

Eleodes granulata LeConte, 1857: 50. Synonymy (in doubt): Horn (1870: 309).

Eleodes obtusa LeConte, 1861b: 352. Synonymy: Horn (1870: 309).

Eleodes letcheri var. vandykei Blaisdell, 1909: 136. Synonymy: Triplehorn and Thomas (2015: 18).

Eleodes vandykei var. modificata Blaisdell, 1921a: 131. Synonymy: Triplehorn and Thomas (2015: 18).

Eleodes vandykei similis Blaisdell, 1942: 142. Synonymy: Triplehorn and Thomas (2015: 18).

Eleodes hirtipennis Triplehorn, 1964USA (CO)

Eleodes hirtipennis Triplehorn, 1964b: 60.

Eleodes letcheri Blaisdell, 1909USA (ID NV OR UT)

Eleodes letcheri Blaisdell, 1909: 133.

Eleodes papillosa Blaisdell, 1917USA (CA)

Eleodes granulata forma tuberculata Blaisdell, 1909: 131 [junior primary homonym of Eleodes tuberculata Eschscholtz, 1829].

Eleodes papillosa Blaisdell, 1917: 226. Replacement name for Eleodes tuberculata Blaisdell, 1909.

Eleodes subtuberculata Walker, 1866CAN (BC) USA (CA ID MT OR WA)

Eleodes subtuberculata Walker, 1866: 328.

Subgenus Melaneleodes Blaisdell, 1909

Melaneleodes Blaisdell, 1909: 36. Type species: Blaps carbonaria Say, 1824, subsequent designation (Triplehorn and Thomas 2012: 254).

Eleodes anthracina anthracina Blaisdell, 1909USA (AZ NM) MEX (CH SO)

Eleodes quadricollis var. anthracina Blaisdell, 1909: 87.

Eleodes anthracina lustrans Blaisdell, 1909USA (AZ)

Eleodes quadricollis var. lustrans Blaisdell, 1909: 89.

Eleodes carbonaria carbonaria (Say, 1824)USA (AZ CO NM NV TX UT WY) MEX (CH CO DU JA NL SL SO TA ZA)

Blaps carbonaria Say, 1824a: 260.

Eleodes vicina LeConte, 1851: 133. Synonymy: Triplehorn and Thomas (2012: 256).

Eleodes immunis LeConte, 1858c: 186. Synonymy: Horn (1870: 308).

Eleodes porcatus Casey, 1890b: 396. Synonymy: Triplehorn and Thomas (2012: 257).

Eleodes carbonaria forma interstitialis Blaisdell, 1909: 47. Synonymy: Triplehorn and Thomas (2012: 257).

Eleodes carbonaria forma glabra Blaisdell, 1909: 47. New synonymy [ADS & MAJ].

Eleodes mazatzalensis Blaisdell, 1925c: 379. Synonymy: Triplehorn and Thomas (2012: 257).

Eleodes carbonaria chihuahuensis Champion, 1884USA (AZ NM) MEX (CH CO DU SO)

Elaeodes chihuahuensis Champion, 1884: 86.

Eleodes nitidus Casey, 1891: 58. Synonymy: Triplehorn and Thomas (2012: 270).

Eleodes ampla Blaisdell, 1909: 53. Synonymy: Triplehorn and Thomas (2012: 270).

Eleodes ampla var. dolosa Blaisdell, 1909: 57. Synonymy (with E. nitidus Casey): Blaisdell (1910: 65).

Eleodes lineata Blaisdell, 1939b: 55. Synonymy: Triplehorn and Thomas (2012: 270).

Eleodes carbonaria disjuncta Triplehorn and Thomas, 2012MEX (GU HI ME ZA)

Eleodes carbonarius disjunctus Triplehorn and Thomas, 2012: 269.

Eleodes carbonaria knausii Blaisdell, 1909USA (CO NM)

Eleodes knausii Blaisdell, 1909: 67.

Eleodes carbonaria nuevoleonensis Triplehorn and Thomas, 2012MEX (CO NL)

Eleodes carbonarius nuevoleonensis Triplehorn and Thomas, 2012: 265.

Eleodes carbonaria obsoleta (Say, 1824)CAN (AB MB SK) USA (AZ CO KS MT ND NE NM OK SD TX UT WY) MEX (SO)

Blaps obsoleta Say, 1824a: 261.

Eleodes obsoleta forma glabra Blaisdell, 1909: 60. Synonymy: Triplehorn and Thomas (2012: 272).

Eleodes obsoleta forma annectans Blaisdell, 1909: 60. Synonymy: Triplehorn and Thomas (2012: 272).

Eleodes obsoleta forma punctata Blaisdell, 1909: 60. Synonymy: Triplehorn and Thomas (2012: 272).

Eleodes carbonaria omissoides Blaisdell, 1935MEX (DU NA NL SI SO ZA)

Eleodes omissoides Blaisdell, 1935c: 157.

Eleodes carbonaria omissa LeConte, 1858USA (CA NV) MEX (BC)

Eleodes omissa LeConte, 1858c: 186.

Eleodes interrupta Blaisdell, 1892: 241. Synonymy: Blaisdell (1909: 72).

Eleodes omissa forma catalinae Blaisdell, 1909: 73. Synonymy: Triplehorn and Thomas (2012: 258).

Eleodes omissa forma communis Blaisdell, 1909: 73. Synonymy: Triplehorn and Thomas (2012: 258).

Eleodes omissa forma emarginata Blaisdell, 1909: 74. Synonymy: Triplehorn and Thomas (2012: 258).

Eleodes omissa var. pygmaea Blaisdell, 1909: 77. Synonymy: Triplehorn (1996: 5).

Eleodes omissa var. peninsularis Blaisdell, 1909: 79. Synonymy: Triplehorn (1996: 5).

Eleodes omissa tumida Blaisdell, 1933b: 194. Synonymy: Triplehorn (1996: 5).

Eleodes carbonaria soror LeConte, 1858USA (TX) MEX (CO NL TA)

Eleodes soror LeConte, 1858c: 185.

Eleodes halli Blaisdell, 1941USA (AZ UT)

Eleodes fuscipilosa Blaisdell, 1925c: 376 [junior secondary homonym of Eleodes fuscipilosus (Casey, 1890)].

Eleodes halli Blaisdell, 1941a: 37. Synonymy: Triplehorn and Thomas (2012: 275).

Eleodes humeralis LeConte, 1857CAN (BC) USA (CA ID OR UT WA)

Eleodes humeralis LeConte, 1857: 50.

Eleodes latiuscula Walker, 1866: 329. Synonymy: LeConte (1873: 334).

Eleodes neomexicana Blaisdell, 1909USA (NM TX)

Eleodes pedinoides var. neomexicana Blaisdell, 1909: 113.

Eleodes parowana Blaisdell, 1925USA (UT)

Eleodes parowana Blaisdell, 1925c: 374.

Eleodes parowana mimica Blaisdell, 1925c: 375. Synonymy: Triplehorn and Thomas (2012: 276).

Eleodes pedinoides LeConte, 1858USA (NM TX) MEX (CO NL TA)

Eleodes pedinoides LeConte, 1858c: 183.

Eleodes asperata LeConte, 1858c: 183. Synonymy: Horn (1870: 307).

Eleodes quadricollis Eschscholtz, 1829USA (CA)

Eleodes quadricollis Eschscholtz, 1829: 12.

Eleodes tarsalis Casey, 1890b: 399. Synonymy: Casey (1893: 597).

Eleodes cuneaticollis Casey, 1890b: 397. Synonymy: Triplehorn and Thomas (2012: 272).

Eleodes rileyi reducta Blaisdell, 1925USA (UT)

Eleodes reducta Blaisdell, 1925c: 377.

Eleodes rileyi rileyi Casey, 1891USA (AZ CA CO ID MT NM NV UT WY)

Eleodes rileyi Casey, 1891: 57.

Eleodes humeralis forma tuberculo-muricata Blaisdell, 1909: 97. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes humeralis forma granulato-muricata Blaisdell, 1909: 97. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes quadricollis lassenica Blaisdell, 1925c: 373. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes coloradensis Blaisdell, 1925c: 380. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes concinna Blaisdell, 1925c: 381. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes tanneri Blaisdell, 1932a: 74. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes rufipes rufipes Pierre, 1976MEX (PU/VE [Pico de Orizaba])

Eleodes alticola rufipes Pierre, 1976: 708.

Eleodes rufipes transvolcanensis Thomas, 2005MEX (ME/MO/PU [Popocatépetl])

Eleodes alticola Pierre, 1976: 706 [junior primary homonym of Eleodes alticola Blaisdell, 1925].

Eleodes transvolcanensis Thomas, 2005: 55334. Replacement name for Eleodes alticola Pierre, 1976.

Eleodes tricostata (Say, 1824) [Fig. 19] CAN (AB MB SK) USA (AZ CO IA KS MN MO MT ND NE NM OK SD TX WI WY) MEX (CO TA)

Figure 19. 

Eleodes (Melaneleodes) tricostata (Say, 1824). Scale bar = 1 mm.

Blaps tricostata Say, 1824a: 262.

Pimelia alternata Kirby, 1837: 232. Synonymy: LeConte (1851: 133).

Eleodes planata Solier, 1848: 366 [junior primary homonym of Eleodes planata Eschscholtz, 1829]. Synonymy: LeConte (1866a: 60).

Eleodes robusta LeConte, 1858c: 183. Synonymy: Horn (1870: 307).

Eleodes tricostata forma ovalis Blaisdell, 1909: 106. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes tricostata forma costata Blaisdell, 1909: 106. Synonymy: Triplehorn and Thomas (2012: 274).

Eleodes wenzeli speculicollis Blaisdell, 1925USA (TX)

Eleodes speculicollis Blaisdell, 1925c: 382.

Eleodes wenzeli wenzeli Blaisdell, 1925USA (NM TX)

Eleodes wenzeli Blaisdell, 1925c: 381.

Subgenus Metablapylis Blaisdell, 1909

Metablapylis Blaisdell, 1909: 391. Type species: Eleodes nigrina LeConte, 1858, present designation.

Eleodes aalbui Triplehorn, 2007USA (CA)

Eleodes aalbui Triplehorn, 2007: 628.

Eleodes californica Blaisdell, 1929USA (CA)

Eleodes californica Blaisdell, 1929a: 165.

Eleodes delicata Blaisdell, 1929USA (AZ TX UT) MEX (BC)

Eleodes delicata Blaisdell, 1929a: 164.

Eleodes dissimilis Blaisdell, 1909USA (AZ CA NM NV TX UT) MEX (SO)

Eleodes dissimilis Blaisdell, 1909: 398.

Eleodes nevadensis Blaisdell, 1909USA (AZ CA NV UT)

Eleodes dissimilis var. nevadensis Blaisdell, 1909: 402.

Eleodes nigrina difformis Blaisdell, 1925USA (ID OR WA)

Eleodes nigrina difformis Blaisdell, 1925c: 389.

Eleodes nigrina maclayi Boddy, 1957USA (OR)

Eleodes nigrina maclayi Boddy, 1957: 197.

Eleodes nigrina nigrina LeConte, 1858 [Fig. 20] CAN (BC) USA (AZ CA CO ID KS ND NE NM NV OR TX UT)

Figure 20. 

Eleodes (Metablapylis) nigrina nigrina LeConte, 1858. Scale bar = 1 mm.

Eleodes nigrina LeConte, 1858c: 186.

Eleodes nigrina perlonga Blaisdell, 1909USA (ID WY)

Eleodes nigrina var. perlonga Blaisdell, 1909: 398.

Subgenus Omegeleodes Triplehorn and Thomas, 2012

Omegeleodes Triplehorn and Thomas, 2012: 253. Type species: Eleodes debilis LeConte, 1858, original designation.

Eleodes debilis LeC4onte, 1858USA (AZ NM TX) MEX (AG CH CO DU MI NL QU SI SL SO ZA)

Eleodes debilis LeConte, 1858c: 185.

Subgenus Promus LeConte, 1862

Promus LeConte, 1862a: 226. Type species: Blaps opaca Say, 1824, original designation.

Eleodes anachronus Triplehorn, 201035MEX (HI JA OA QU SL TA VE)

Eleodes anachronus Triplehorn, 2010: 373.

Eleodes bidens Triplehorn, 2007MEX (DU)

Eleodes bidens Triplehorn, 2007: 641.

Eleodes brucei Triplehorn, 2007MEX (DU ZA)

Eleodes brucei Triplehorn, 2007: 638.

Eleodes calcarata Champion, 1884MEX (GU)

Elaeodes calcarata Champion, 1884: 86.

Eleodes composita Casey, 1891USA (TX)

Eleodes compositus Casey, 1891: 58.

Eleodes erratica Champion, 1884MEX (NA SI)

Elaeodes erratica Champion, 1884: 87.

Eleodes exarata Champion, 1884MEX (SL)

Elaeodes exarata Champion, 1884: 78.

Eleodes fusiformis LeConte, 1858USA (CO KS NE NM TX WY)

Eleodes fusiformis LeConte, 1858c: 184.

Eleodes goryi Solier, 1848USA (NM TX) MEX (PU TA VE)

Eleodes goryi Solier, 1848: 251.

Eleodes seriata LeConte, 1858c: 185. Synonymy: Champion (1885: 93).

Eleodes hoegei Champion, 1885MEX (PU VE)

Elaeodes högei Champion, 1885: 91.

Eleodes insularis Linell, 1899MEX (BC BS)

Eleodes insularis Linell, 1899: 181.

Eleodes terricola Blaisdell, 1910: 61.36 Synonymy: Triplehorn (1971: 58).

Eleodes knullorum Triplehorn, 1971USA (AZ NM TX) MEX (CO HI)

Eleodes knullorum Triplehorn, 1971: 56.

Eleodes longicornis Champion, 1884MEX (DU)

Elaeodes longicornis Champion, 1884: 87.

Eleodes madrensis Johnston, 2015USA (AZ NM) MEX (SO)

Eleodes madrensis Johnston, 2015: 14.

Eleodes montana Champion, 1884MEX (SL)

Elaeodes montana Champion, 1884: 86.

Eleodes opaca (Say, 1824) [Fig. 21] CAN (AB MB SK) USA (CO KS ND NE OK SD TX)

Figure 21. 

Eleodes (Promus) opaca (Say, 1824). Scale bar = 1 mm.

Blaps opaca Say, 1824a: 263.

Eleodes spiculifera Triplehorn, 2007USA (TX)

Eleodes spiculiferus Triplehorn, 2007: 632.

Eleodes spinolae Solier, 1848MEX (CO FD GE GU HI ME MO OA PU VE)

Eleodes spinolae Solier, 1848: 253.

Eleodes striolata LeConte, 1858USA (TX) MEX (CO)

Eleodes striolata LeConte, 1858c: 185.

Eleodes subnitens LeConte, 1851USA (AZ) MEX (SO)

Eleodes subnitens LeConte, 1851: 134.

Eleodes subnitens forma sinuata Blaisdell, 1909: 163. Synonymy: Johnston (2015: 14).

Eleodes watrousi Triplehorn, 2007MEX (DU)

Eleodes watrousi Triplehorn, 2007: 640.

Subgenus Pseudeleodes Blaisdell, 1909

Pseudeleodes Blaisdell, 1909: 146. Type species: Eleodes granosa LeConte, 1866, monotypy.

Trichoderulus Blaisdell, 1923: 281. Type species: Trichoderulus longipilosus Blaisdell, 1923 (= Eleodes tribulus Thomas, 2005), original designation. Synonymy: Johnston (2016: 672).

Eleodes caudifera LeConte, 1858USA (AZ CO NM TX UT) MEX (CH)

Eleodes caudifera LeConte, 1858c: 184.

Eleodes caudifera forma glabra Blaisdell, 1909: 228. Synonymy: Johnston (2016: 671).

Eleodes caudifera forma scabra Blaisdell, 1909: 228. Synonymy: Johnston (2016: 671).

Eleodes caudifera forma sublaevis Blaisdell, 1909: 228. Synonymy: Johnston (2016: 671).

Eleodes granosa LeConte, 1866USA (CA NV OR)

Eleodes granosa LeConte, 1866b: 116.

Eleodes granosa forma fortis Blaisdell, 1909: 150. New synonymy [YB].

Eleodes granosa var. pilifera Boddy, 1957: 193. Synonymy: Johnston (2016: 671).

Eleodes inyoensis Tanner, 1961: 68. Synonymy: Johnston (2016: 671).

Eleodes inornata Johnston, 2016USA (NV)

Eleodes inornatus Johnston, 2016: 669.

Eleodes leechi Tanner, 1961USA (CO UT)

Eleodes leechi Tanner, 1961: 63.

Eleodes longipilosa Horn, 1891USA (CA ID NV OR)

Eleodes longipilosa Horn, 1891: 42.

Eleodes pilosa Horn, 1870USA (CA ID NM NV OR UT WA WY)

Eleodes pilosa Horn, 1870: 314.

Eleodes pilosa forma ordinata Blaisdell, 1909: 143. New synonymy [YB].

Eleodes obesus Doyen, 1985b: 232. Synonymy: Johnston (2016: 673).

Eleodes spoliata Blaisdell, 1933USA (OR)

Eleodes spoliata Blaisdell, 1933b: 196.

Eleodes tribulus Thomas, 2005USA (AZ) MEX (SO)

Amphidora caudata Horn, 1870: 330 [junior secondary homonym of Eleodes caudata Solier, 1848].

Trichoderulus longipilosus Blaisdell, 1923: 281 [junior secondary homonym of Eleodes longipilosus Horn, 1891]. Synonymy: Triplehorn and Aalbu (1987: 371).

Eleodes blaisdelli Doyen [in Doyen and Lawrence], 1979: 367 [junior primary homonym of Eleodes blaisdelli Blackwelder, 1945]. Replacement name for Eleodes longipilosus Blaisdell, 1923.

Eleodes tribulus Thomas, 2005: 552. Replacement name for Eleodes caudatus (Horn, 1870).

Subgenus Tricheleodes Blaisdell, 1909

Tricheleodes Blaisdell, 1909: 138. Type species: Eleodes hirsuta LeConte, 1861, by subsequent designation (Johnston 2016: 666).

Eleodes hirsuta LeConte, 1861USA (CA NV UT)

Eleodes hirsuta LeConte, 1861b: 352.

Subgenus Xysta Eschscholtz, 1829

Xysta Eschscholtz, 1829: 9. Type species: Eleodes gravida Eschscholtz, 1829, subsequent designation (Hope 1841: 124). Status revised [ADS & MAJ].

Steneleodes Blaisdell, 1909: 409. Type species: Eleodes longicollis LeConte, 1851, present designation. New synonymy [ADS & MAJ].

Holeleodes Blaisdell, 1937b: 132. Type species: Eleodes beameri Blaisdell, 1937 (=Elaeodes hepburni Champion, 1884), original designation. Synonymy (with Steneleodes Blaisdell): Johnston (2015: 12).

Eleodes angulata (Eschscholtz, 1829)MEX (FD ME)

Xysta angulata Eschscholtz, 1829: 9.

Eleodes angusta Eschscholtz, 1829MEX (DU FD GU HI JA ME MI OA PU VE)

Eleodes angusta Eschscholtz, 1829: 13.

Eleodes blapoides Eschscholtz, 1829MEX (OA)

Eleodes blapoides Eschscholtz, 1829: 12.

Elaeodes blaptoides Champion, 1884: 78. Unjustified emendation of Eleodes blapoides Eschscholtz, 1829, not in prevailing usage.

Eleodes coarctata Champion, 1885MEX (ME PU TA)

Elaeodes coarctata Champion, 1885: 91.

Eleodes corrugans Triplehorn, 2007MEX (MI)

Eleodes corrugans Triplehorn, 2007: 630.

Eleodes distincta Solier, 1848MEX (HI OA PU QU SL TA VE)

Eleodes distincta Solier, 1848: 239.

Eleodes forreri Champion, 1884MEX (CH DU)

Elaeodes forreri Champion, 1884: 88.

Eleodes gigantea Mannerheim, 1843USA (CA) MEX (BC)

Eleodes gigantea Mannerheim, 1843: 267.

Eleodes gentilis LeConte, 1858c: 187. Synonymy: Triplehorn (1996: 15).

Eleodes estriatus Casey, 1890b: 398. Synonymy: Triplehorn (1996: 15).

Eleodes gigantea var. meridionalis Blaisdell, 1918c: 387. Synonymy: Triplehorn (1996: 15).

Eleodes glabricollis Champion, 1884MEX (AG GU NL SL)

Elaeodes glabricollis Champion, 1884: 85.

Eleodes gravida (Eschscholtz, 1829)MEX (OA)

Xysta gravida Eschscholtz, 1829: 9.

Eleodes hepburni Champion, 1884USA (AZ NM) MEX (CH CO DU JA SI SO)

Elaeodes hepburni Champion, 1884: 88.

Eleodes compressitarsis Blaisdell, 1935c: 158. Synonymy: Triplehorn (2010: 376).

Eleodes beameri Blaisdell, 1937b: 132. Synonymy: Triplehorn (2010: 376).

Eleodes bryanti Blaisdell, 1937b: 134. Synonymy (with E. beameri Blaisdell): Triplehorn and Doyen (1972: 79).

Eleodes palmerleensis Blaisdell, 1937b: 136. Synonymy (with E. beameri Blaisdell): Triplehorn and Doyen (1972: 79).

Eleodes innocens LeConte, 1866MEX (BS)

Eleodes innocens LeConte, 1866b: 114.

Eleodes laevigata blapsoides Solier, 1848 MEX

Eleodes laevigata var. blapsoïdes Solier, 1848: 244.

Eleodes laevigata laevigata Solier, 1848MEX (ME OA PU VE) GUA

Eleodes laevigata Solier, 1848: 244.

Eleodes longicollis LeConte, 1851USA (AZ CO KS NM NV OR TX UT WY) MEX (AG CH CO DU MI NL SL SO ZA)

Eleodes longicollis LeConte, 1851: 134.

Eleodes haydenii LeConte, 1858c: 186. Synonymy: Horn (1870: 311).

Eleodes mutilata Blaisdell, 1921MEX (BS)

Eleodes mutilata Blaisdell, 1921b: 222.

Eleodes olida Champion, 1892MEX (GE)

Elaeodes olida Champion, 1892: 516.

Eleodes ornatipennis Blaisdell, 1937USA (NM) MEX (CH)

Eleodes ornatipennis Blaisdell, 1937b: 129.

Eleodes peropaca Champion, 1892MEX (DU)

Elaeodes peropaca Champion, 1892: 517.

Eleodes platypennis Triplehorn, 2007MEX (JA)

Eleodes platypennis Triplehorn, 2007: 637.

Eleodes ponderosa Champion, 1884MEX (OA PU)

Elaeodes ponderosa Champion, 1884: 84.

Eleodes punctigera Blaisdell, 1935MEX (DU)

Eleodes punctigera Blaisdell, 1935c: 157.

Eleodes ruida (Say, 1835)MEX (MO PU VE)

Blaps ruida Say, 1835: 183.

Eleodes coriacea Solier, 1848: 249. Synonymy: Champion (1884: 84).

Eleodes sallaei Champion, 1885MEX (GU JA OA PU QU SL VE)

Elaeodes sallaei Champion, 1885: 89.

Eleodes solieri Champion, 1885MEX (CH CO GU OA PU SL VE)

Blaps celsa Say, 1835: 185 [nomen dubium].

Elaeodes solieri Champion, 1885: 89. Synonymy (in doubt): Champion (1885: 89).

Eleodes stolida Champion, 1885 MEX

Elaeodes stolida Champion, 1885: 92.

Eleodes sulcatula Champion, 1884MEX (ME)

Elaeodes sulcatula Champion, 1884: 83.

Eleodes tenebricosa Gemminger, 1870MEX (ME OA)

Eleodes obscura Solier, 1848: 245 [junior secondary homonym of Eleodes obscurus (Say, 1824)].

Elaeodes tenebricosa Gemminger, 1870: 122. Replacement name for Elaeodes obscura Solier, 1848.

Eleodes tessellata Champion, 1892MEX (MI)

Elaeodes tessellata Champion, 1892: 517.

[incertae sedis]

Eleodes aequalis (Say, 1835)MEX (DU ME OA PU)

Blaps aequalis Say, 1835: 185.

Eleodes alutacea Solier, 1848: 240. Synonymy (in doubt): Champion (1884: 80).

Eleodes maillei Solier, 1848: 247. Synonymy (with E. alutacea Solier): Champion (1884: 80).

Eleodes amaura Champion, 1892MEX (GU HI OA PU TA)

Elaeodes amaura Champion, 1892: 514.37

Eleodes barbata Wickham, 1918USA (AZ CO NM UT)

Eleodes barbata Wickham, 1918: 256.

Eleodes brevicollis Gemminger, 1870 MEX

Eleodes obsoleta Solier, 1848: 238 [junior secondary homonym of Eleodes obsoleta (Say, 1823)].

Eleodes brevicollis Gemminger [in Gemminger and Harold], 1870: 1868. Replacement name for Eleodes obsoleta Solier, 1848.

Eleodes cylindrica (Herbst, 1799) “Nordamerika”

Blaps cylindrica Herbst, 1799: 185.

Eleodes dilaticollis Champion, 1884MEX (GU HI ME MI PU)

Elaeodes dilaticollis Champion, 1884: 83.

Eleodes ebenina (Solier, 1848) MEX

Nycterinus ebeninus Solier, 1848: 269.

Eleodes elongatula Eschscholtz, 1829 MEX

Eleodes elongatula Eschscholtz, 1829: 13.

Eleodes impolita (Say, 1835)MEX (ME OA PU VE)

Blaps impolita Say, 1835: 183.

Eleodes aubei Solier, 1848: 245. Synonymy: Champion (1885: 90).

Eleodes maura (Say, 1835)MEX (GU OA PU)

Blaps maura Say, 1835: 184.

Eleodes melanaria Eschscholtz, 1829 MEX

Eleodes melanaria Eschscholtz, 1829: 13.

Eleodes obliterata (Say, 1835) MEX

Blaps obliterata Say, 1835: 184.

Eleodes polita Champion, 1892MEX (ME MO)

Elaeodes polita Champion, 1892: 513.

Eleodes rotundicollis (Eschscholtz, 1829)MEX (PU SL VE)

Xysta rotundicollis Eschscholtz, 1829: 9.

Blaps parva Say, 1835: 186. Synonymy (in doubt): Champion (1884: 82).

Eleodes scapularis Champion, 1884MEX (GU ME)

Elaeodes scapularis Champion, 1884: 81.

Eleodes segregata Champion, 1892MEX (CH DU GE MI)

Elaeodes segregata Champion, 1892: 513.

Eleodes striata (Guérin-Méneville, 1834)MEX (TA)

Xysta striata Guérin-Méneville, 1834: 30.

Eleodes sulcata (Eschscholtz, 1829)MEX (MO)

Xysta sulcata Eschscholtz, 1829: 9.

Genus Eleodimorpha Blaisdell, 1909 [F]

Eleodimorpha Blaisdell, 1909: 477. Type species: Eleodimorpha bolcan Blaisdell, 1909, original designation.

Eleodimorpha bolcan Blaisdell, 1909USA (CA)

Eleodimorpha bolcan Blaisdell, 1909: 479.

Genus Embaphion Say, 1824 [N]

Embaphion Say, 1824a: 254. Type species: Akis muricata Say, 1824, monotypy.

Embaphion contractum blaisdelli Benedict, 1927USA (NM)

Embaphion contractum blaisdelli Benedict, 1927: 46.

Embaphion contractum contractum Blaisdell, 1909USA (NM)

Embaphion contractum Blaisdell, 1909: 460.

Embaphion contusum contusum LeConte, 1858USA (AZ CO KS NM WY)

Embaphion contusum LeConte, 1858a: 20.

Embaphion contusum grande Blaisdell, 1909USA (NM)

Embaphion contusum forma grandis Blaisdell, 1909: 471.

Embaphion contusum laminatum Casey, 1890USA (TX)

Embaphion laminatum Casey, 1890b: 403.

Embaphion depressum (LeConte, 1851)USA (CA)

Eleodes depressa LeConte, 1851: 136.

Embaphion elongatum Horn, 1870USA (CA ID NV OR UT)

Embaphion elongatum Horn, 1870: 321.

Embaphion glabrum Blaisdell, 1909USA (AZ NM UT)

Embaphion glabrum Blaisdell, 1909: 457.

Embaphion mexicanum Blaisdell, 1935MEX (CH)

Embaphion mexicanum Blaisdell, 1935c: 160.

Embaphion muricatum (Say, 1824) [Fig. 22] CAN (AB SK) USA (CO KS NE SD TX) MEX (TA)

Figure 22. 

Embaphion muricatum (Say, 1824). Scale bar = 1 mm.

Akis muricata Say, 1824a: 253.

Embaphion concavum LeConte, 1853: 446. Synonymy: Horn (1870: 320).

Embaphion planum Horn, 1870USA (CO KS NM UT WY)

Embaphion planum Horn, 1870: 321.

Genus Lariversius Blaisdell, 1947 [M]

Lariversius Blaisdell, 1947: 59. Type species: Lariversius tibialis Blaisdell, 1947, original designation.

Lariversius tibialis Blaisdell, 1947USA (NV)

Lariversius tibialis Blaisdell, 1947: 61.

Genus Neobaphion Blaisdell, 1925 [N]

Neobaphion Blaisdell, 1925c: 390. Type species: Eleodes planipennis LeConte, 1866, monotypy.

Neobaphion alleni Triplehorn, 1989USA (ID OR)

Neobaphion alleni Triplehorn, 1989: 458.

Neobaphion elongatum Blaisdell, 1933USA (CA NV)

Neobaphion elongatum Blaisdell, 1933b: 208.

Neobaphion papula Triplehorn and Aalbu, 1985USA (NV)

Neobaphion papula Triplehorn and Aalbu, 1985 [11 July]: 588.

Eleodes insolitus Doyen, 1985b [11 July]: 230. Synonymy: Triplehorn (1989: 460).

Neobaphion planipenne (LeConte, 1866)USA (AZ CO NM UT)

Eleodes planipennis LeConte, 1866b: 116.

Genus Trogloderus LeConte, 1879 [M]

Trogloderus LeConte, 1879a: 2. Type species: Trogloderus costatus LeConte, 1879, monotypy.

Trogloderus costatus LeConte, 1879USA (CA ID NV)

Trogloderus costatus LeConte, 1879a: 3.

Trogloderus nevadus La Rivers, 1943USA (ID NV) Status revised [MAJ]

Trogloderus nevadus La Rivers, 1943a: 437.

Trogloderus tuberculatus Blaisdell, 1909USA (CA) Status revised [MAJ]

Trogloderus tuberculatus Blaisdell, 1909: 490.

Trogloderus costatus pappi Kulzer, 1960: 310. New synonymy [MAJ]

Trogloderus vandykei La Rivers, 1946USA (AZ CA) Status revised [MAJ]

Trogloderus costatus vandykei La Rivers, 1946: 41.

Trogloderus costatus mayhewi Papp, 1961a: 33. New synonymy [MAJ]

Tribe Apocryphini Lacordaire, 1859

Apocryphides Lacordaire, 1859: 432. Type genus: Apocrypha Eschscholtz, 1831.

Genus Apocrypha Eschscholtz, 1831 [F]

Apocrypha Eschscholtz, 1831: 13. Type species: Apocrypha anthicoides Eschscholtz, 1831, monotypy.

Compsomorphus Solier, 1851: 208. Type species: Compsomorphus elegans Solier, 1851, monotypy. Synonymy: Lacordaire (1859: 433).

Apocrypha anthicoides Eschscholtz, 1831USA (CA)

Apocrypha anthicoides Eschscholtz, 1831: 13.

Apocrypha dyschirioides LeConte, 1851: 137. Synonymy: Doyen and Kitayama (1980: 122).

Apocrypha clivinoides Horn, 1870USA (CA)

Apocrypha clivinoides Horn, 1870: 391.

Apocrypha setosa Doyen and Kitayama, 1980USA (CA)

Apocrypha setosa Doyen and Kitayama, 1980: 126.

Genus Pseudapocrypha Champion, 1886 [F]

Pseudapocrypha Champion, 1886: 260. Type species: Pseudapocrypha lacordairii Champion, 1886, monotypy.

Pseudapocrypha lacordairii Champion, 1886MEX (CI) GUA

Pseudapocrypha lacordairii Champion, 1886: 260.

Tribe Blaptini Leach, 1815

Blapsida Leach, 1815: 101. Type genus: Blaps Fabricius, 1775.

Subtribe Blaptina Leach, 1815

Blapsida Leach, 1815: 101. Type genus: Blaps Fabricius, 1775.

Genus Blaps Fabricius, 1775 [F]

Blaps Fabricius, 1775: 254. Type species: Tenebrio mortisagus Linnaeus, 1758, subsequent designation (Latreille 1810: 429).

Subgenus Blaps Fabricius, 1775

Blaps Fabricius, 1775: 254. Type species: Tenebrio mortisagus Linnaeus, 1758, subsequent designation (Latreille 1810: 429).

Blaps lethifera lethifera Marsham, 1802CAN (QC) USA (IN MD NJ NY OH VA) – Adventive

Blaps lethifera Marsham, 1802: 479.

Blaps similis Latreille, 1804: 279. Synonymy: Seidlitz (1893: 317).

Blaps mucronata Latreille, 1804USA (MD NY OH) – Adventive

Blaps mucronata Latreille, 1804: 278.

Tribe Bolitophagini Kirby, 1837

Eledonaedes Billberg, 1820b: 392 [nomen oblitum, see Bouchard et al. 2011]. Type genus: Eledona Latreille, 1797.

Bolitophagidae Kirby, 1837: 236 [nomen protectum]. Type genus: Bolitophagus Illiger, 1798.

Rhipidandri LeConte, 1862a: 236. Type genus: Rhipidandrus LeConte, 1862.

Eutomides Lacordaire, 1865: 369. Type genus: Eutomus Lacordaire, 1865 (= Rhipidandrus LeConte, 1862).

Genus Bolitophagus Illiger, 1798 [M]

Bolitophagus Illiger, 1798: 100. Type species: Silpha reticulata Linnaeus, 1767, subsequent designation (C.G. Thomson 1859: 115).

Boletophagus Agassiz, 1846: 48. Unjustified emendation of Bolitophagus Illiger, 1798, not in prevailing usage.

Bolitophagus corticola Say, 1826CAN (NB NS ON PE QC) USA (CT DC FL GA IN MA MD ME MI MO NC NH NJ NY OH PA SC TN TX VA WI)

Boletophagus corticola Say, 1826: 238.

Genus Bolitotherus Candèze, 1861 [M]

Bolitotherus Candèze, 1861: 367. Type species: Bolitophagus cornutus Fabricius, 1801, subsequent designation (LeConte 1862b: 236).

Phellidius LeConte, 1862a: 236. Type species: Bolitophagus cornutus Fabricius, 1801, original designation. Synonymy: LeConte (1866a: 62).

Bolitotherus cornutus (Fabricius, 1801)38 [Fig. 23] CAN (AB MB NB NS ON PE QC SK) USA (CT FL GA IA IL IN KY LA MA MD ME MI MN MS NC NE NH NJ NY OH PA RI SC TN TX VAVT WI)

Figure 23. 

Bolitotherus cornutus (Fabricius, 1801). Scale bar = 1 mm.

Opatrum bifurcum Fabricius, 1798: 40.

Bolitophagus cornutus Fabricius, 1801a: 112. Synonymy: Fabricius (1801a: 113).

Bolitophagus cristatus Gosse, 1840: 251. New synonymy [YB].

Genus Eleates Casey, 1886 [M]

Eleates Casey, 1886: 253. Type species: Eleates occidentalis Casey, 1886, monotypy.

Eleates depressus (Randall, 1838)CAN (BC MB NB NT ON QC SK) USA (AR GA MD ME MI NH NYOH OR PA TN VAVT WA WI)

Eledona depressa Randall, 1838 [February]: 21.

Bolitophagus tetraopes Newman, 1838 [April]: 378. Synonymy: LeConte (1854b: 219).

Eleates explanatus Casey, 1890b: 486. New synonymy [YB].

Eleates occidentalis Casey, 1886USA (CA)

Eleates occidentalis Casey, 1886: 254.

Genus Megeleates Casey, 1895 [M]

Megeleates Casey, 1895: 623. Type species: Megeleates sequoiarum Casey, 1895, monotypy.

Megeleates sequoiarum Casey, 1895USA (CA OR WA)

Megeleates sequoiarum Casey, 1895: 624.

Genus Rhipidandrus LeConte, 1862 [M]

Rhipidandrus LeConte, 1862a: 236. Type species: Xyletinus flabellicornis Sturm, 1826 (= Melolontha paradoxa Palisot de Beauvois, 1818), monotypy.

Eutomus Lacordaire, 1865: 369. Type species: Eutomus micrographus Lacordaire, 1865, subsequent designation (Barber 1914: 191). Synonymy: LeConte and Horn (1883: 232).

Heptaphylla Friedenreich, 1883: 375. Type species: Heptaphylla fungicola Friedenreich, 1883, monotypy. Synonymy: Arrow (1904: 31).

Cherostus C.O. Waterhouse, 1894: 68. Type species: Cherostus walkeri Waterhouse, 1894, subsequent designation (Merkl and Kompantzeva 1996: 91). Synonymy: Gebien (1939: 762).

Rhipidandrus championi Sharp, 1905 GUA PAN

Rhipidandrus championi Sharp, 1905: 691.

Rhipidandrus cornutus (Arrow, 1904)MEX (DU OA) / HISPRILAN / SA

Cherostus cornutus Arrow, 1904: 31.

Rhipidandrus fulvomaculatus Dury, 1914USA (FL) / BAH

Rhipidandrus fulvomaculata Dury, 1914: 168.

Rhipidandrus jamaicensis (Arrow, 1904) JAM

Cherostus jamaicensis Arrow, 1904: 32.

Rhipidandrus mexicanus Sharp, 1905MEX (VE) GUABEL

Rhipidandrus mexicanus Sharp, 1905: 691.

Rhipidandrus micrographus (Lacordaire, 1865)PRILAN / SA

Eutomus micrographus Lacordaire, 1865: 370.

Rhipidandrus panamaensis (Barber, 1914) PAN

Eutomus panamaensis Barber, 1914: 193.

Rhipidandrus paradoxus (Palisot de Beauvois, 1818) [Fig. 24] CAN (ON QC) USA (DC FL GA IN KS KY LA MD MI NC NY OH SC TX VA WI)

Figure 24. 

Rhipidandrus paradoxus (Palisot de Beauvois, 1818). Scale bar = 1 mm.

Melolontha paradoxa Palisot de Beauvois, 1818: 173.

Xylotinus flabellicornis Sturm, 1826: 59. Synonymy: LeConte (1873: 329, 335).

Rhipidandrus peninsularis Horn, 1894USA (AZ TX) MEX (BS)

Rhipidandrus peninsularis Horn, 1894b: 392.

Rhipidandrus sulcatus (Gorham, 1898) BAH CUB CAY HIS LAN

Eutomus sulcatus Gorham, 1898: 333.

Tribe Centronopini Doyen, 1989

Centronopini Doyen, 1989: 284. Type genus: Centronopus Solier, 1848.

Genus Centronopus Solier, 1848 [M]

Centronopus Solier, 1848: 258. Type species: Centronopus extensicollis Solier, 1848 (= Tenebrio suppressus Say, 1835), original designation.

Subgenus Centronopus Solier, 1848

Centronopus Solier, 1848: 258. Type species: Centronopus extensicollis Solier, 1848 (= Tenebrio suppressus Say, 1835), original designation.

Centronopus grandicollis Champion, 1885MEX (DU FD HI ME MO SL TA VE)

Centronopus grandicollis Champion, 1885: 100.

Centronopus suppressus (Say, 1835)MEX (FD HI ME MO PU SL TA VE)

Tenebrio suppressus Say, 1835: 187.

Centronopus extensicollis Solier, 1848: 260. Synonymy: Lacordaire (1859: 361).

Subgenus Menechides Motschulsky, 1872

Menechides Motschulsky, 1872: 26. Type species: Helops calcaratus Fabricius, 1798, original designation.

Scotobates Rye, 1877: 341. Type species: Helops calcaratus Fabricius, 1798, subsequent designation (Lucas 1920: 587). Synonymy: Lucas (1920: 403).

Pyres Champion, 1885: 100. Type species: Pyres metallicus Champion, 1885 (= Centronopus speciosus Pascoe, 1883), subsequent designation (Gebien 1941: 336). Synonymy: Spilman (1962a: 3).

Centronopus batesi (Champion, 1885)PAN / SA

Pyres batesi Champion, 1885: 101.

Centronopus beardsleyi Spilman, 1962MEX (CL JA)

Centronopus beardsleyi Spilman, 1962a: 11.

Centronopus bimaculatus Champion, 1892MEX (VE YU) BEL

Centronopus bimaculatus Champion, 1892: 521.

Centronopus calcaratus (Fabricius, 1798)CAN (NS ON QC) USA (AL CT DC FL GA IA IL IN KS LA MA MD ME MI MO MS NC NE NH NJ NY OH PA SC TN VA VT WI WV)

Tenebrio aeneus DeGeer, 1775: 53 [junior primary homonym of Tenebrio aeneus Scopoli, 1763] [nomen dubium].

Helops calcaratus Fabricius, 1798: 52. Synonymy (in doubt): LeConte (1866a: 61).

Tenebrio coracinus Knoch, 1801: 172. Synonymy: LeConte (1866a: 61).

Helops caroliniensis Palisot de Beauvois, 1817: 162. Synonymy (in doubt): LeConte (1866a: 61).

Tenebrio reflexus Say, 1825: 203. Synonymy: Melsheimer (1853: 139).

Centronopus nigrofasciatus (Gebien, 1928) CRI

Pyres nigrofasciatus Gebien, 1928b: 182.

Centronopus opacus LeConte, 1859USA (AR KS MT OK SD TX)

Centronopus opacus LeConte, 1859a: 15.

Centronopus speciosus Pascoe, 1883 NIC CRI

Centronopus speciosus Pascoe, 1883: 439.

Pyres metallicus Champion, 1885: 101. Synonymy: Champion (1892: 521).

Genus Scotobaenus LeConte, 1859 [M]

Scotobaenus LeConte, 1859b: 87. Type species: Scotobaenus parallelus LeConte, 1859, monotypy.

Scotobaenus parallelus LeConte, 1859USA (CA OR WA) MEX

Scotobaenus parallelus LeConte, 1859b: 88.

Scotobaenus punctatus (Blaisdell, 1933)USA (CA)

Centronopus punctatus Blaisdell, 1933c: 220.

Scotobaenus simplex (Blaisdell, 1937)USA (CA)

Centronopus simplex Blaisdell, 1937a: 95.

Scotobaenus wagneri (Blaisdell, 1933)USA (CA)

Centronopus wagneri Blaisdell, 1933c: 218.

Genus Tauroceras Hope, 1841 [N]

Tauroceras Hope, 1841: 130. Type species: Tenebrio cornutus Fabricius, 1775, original designation.

Tauroceropedus Pic, 1913b: 4. Type species: Tauroceropedus difformipes Pic, 1913, subsequent designation (Gebien 1941: 344). Synonymy: Ferrer et al. (2005: 272).

Tauroceras barclayi Ferrer, Soldati and Delatour, 2005 JAM

Tauroceras barclayi Ferrer, Soldati and Delatour, 2005: 284.

Tauroceras cornutum (Fabricius, 1775) CUB JAM

Tenebrio cornutus Fabricius, 1775: 256.

Tauroceras girardi Ferrer, Soldati and Delatour, 2005MEX (QR) GUAHONNICCRI

Tauroceras girardi Ferrer, Soldati and Delatour, 2005: 288.

Tauroceras mulata Zayas, 1988 CUB

Tauroceras mulata Zayas, 1988: 94.

Tribe Cerenopini Horn, 1870

Cerenopi Horn, 1870: 323. Type genus: Cerenopus LeConte, 1851.

Genus Argoporis Horn, 1870 [F]

Argoporis Horn, 1870: 325. Type species: Cerenopus costipennis LeConte, 1851, subsequent designation (Gebien 1937: 797).

Threnus Motschulsky, 1870: 404. Type species: Threnus niger Motschulsky, 1870, original designation. Synonymy: Aalbu et al. (1995: 483)39.

Argoporis aequalis Blaisdell, 1923MEX (SO)

Argoporis aequalis Blaisdell, 1923: 259.

Argoporis alutacea Casey, 1890USA (AZ) MEX (SO)

Argoporis alutacea Casey, 1890b: 406.

Argoporis labialis Blaisdell, 1923: 258. Synonymy: Berry (1980: 18).

Argoporis angusta Casey, 1924: 331. Synonymy: Berry (1980: 18).

Argoporis hebes Casey, 1924: 332. Synonymy: Berry (1980: 18).

Argoporis tibialis Casey, 1924: 332. Synonymy: Berry (1980: 18).

Argoporis apicalis apicalis Blaisdell, 1943MEX (BC BS)

Argoporis apicalis Blaisdell, 1943: 234.

Argoporis insularis Berry, 1980: 54. Synonymy: Sánchez Piñero and Aalbu (2002: 132).

Argoporis apicalis californica Berry, 1980USA (AZ) MEX (BC)

Argoporis apicalis californica Berry, 1980: 22.

Argoporis atripes Horn, 1870MEX (AG DU GU HI JA MI SI SL SO)

Argoporis atripes Horn, 1870: 325.

Argoporis bicolor (LeConte, 1851)USA (AZ CA) MEX (SO)

Cerenopus bicolor LeConte, 1851: 143.

Argoporis tuckeri Casey, 1924: 332. Synonymy: Berry (1980: 27).

Argoporis brevicollis Champion, 1885MEX (DU SI)

Argoporis brevicollis Champion, 1885: 94.

Argoporis carinata Berry, 1980USA (AZ) MEX (CH NA SI SO)

Argoporis carinata Berry, 1980: 30.

Argoporis cavifrons Champion, 1885MEX (DU SI)

Argoporis cavifrons Champion, 1885: 95.

Argoporis colimensis Berry, 1980MEX (CL)

Argoporis colimensis Berry, 1980: 34.

Argoporis costipennis (LeConte, 1851)USA (AZ NM) MEX (SO)

Cerenopus costipennis LeConte, 1851: 143.40

Argoporis lateralis Casey, 1924: 331. Synonymy: Berry (1980: 38).

Argoporis costulata (Horn, 1870)MEX (BC BS)

Cerenopus costulatus Horn, 1870: 326.

Argoporis craigi Berry, 1980MEX (CH DU)

Argoporis craigi Berry, 1980: 42.

Argoporis crassicornis Champion, 1885MEX (DU NA SI)

Argoporis crassicornis Champion, 1885: 94.

Argoporis cribrata (LeConte, 1861)MEX (BS)

Cerenopus cribratus LeConte, 1861a: 337.

Argoporis deltodonta Berry, 1980MEX (Tres Marias Islands)

Argoporis deltodonta Berry, 1980: 46.

Argoporis durangoensis Berry, 1980MEX (CH DU)

Argoporis durangoensis Berry, 1980: 47.

Argoporis ebenina Horn, 1894MEX (BS)

Argoporis ebenina Horn, 1894b: 424.

Argoporis estebanensis Berry, 1980MEX (BS)

Argoporis estebanensis Berry, 1980: 50.

Argoporis impressa Blaisdell, 1925MEX (BC BS)

Argoporis impressa Blaisdell, 1925b: 330.

Argoporis inconstans Horn, 1894MEX (BC BS)

Argoporis inconstans Horn, 1894b: 425.

Argoporis laevicollis Champion, 1892MEX (DU SI)

Argoporis laevicollis Champion, 1892: 520.

Argoporis longipes Blaisdell, 1923MEX (BS)

Argoporis longipes Blaisdell, 1923: 260.

Argoporis nigra inflata Berry, 1980MEX (BS)

Argoporis constanzae inflata Berry, 1980: 37.

Argoporis nigra nigra (Motschulsky, 1870)MEX (BS)

Threnus niger Motschulsky, 1870: 40641.

Argoporis constanzae constanzae Berry, 1980: 35. Synonymy: Aalbu et al. (1995: 483).

Argoporis obregonensis Berry, 1980MEX (SO)

Argoporis obregonensis Berry, 1980: 58.

Argoporis regalis Berry, 1980MEX (BS)

Argoporis regalis Berry, 1980: 59.

Argoporis rufipes femorata Berry, 1980MEX (CH DU GU SI SL ZA)

Argoporis rufipes femorata Berry, 1980: 63.

Argoporis rufipes nitida Casey, 1890USA (AZ NM TX) MEX (CH CO)

Argoporis nitida Casey, 1890b: 406.

Argoporis rufipes rufipes Champion, 1885MEX (AG CH CO SL SO ZA)

Argoporis rufipes Champion, 1885: 94.

Argoporis tridentata Champion, 1892MEX (CL GE JA MI)

Argoporis tridentata Champion, 1892: 519.

Argoporis unicalcarata Champion, 1892MEX (AG JA NA SO)

Argoporis unicalcarata Champion, 1892: 519.

Genus Cerenopus LeConte, 1851 [M]

Cerenopus LeConte, 1851: 14342. Type species: Cerenopus concolor LeConte, 1851, subsequent designation (Lucas 1920: 173).

Cerenopus angustatus Horn, 1894MEX (BS)

Cerenopus angustatus Horn, 1894b: 426.

Cerenopus aterrimus Horn, 1894MEX (BS)

Cerenopus aterrimus Horn, 1894b: 425.

Cerenopus concolor LeConte, 1851USA (AZ CA NV) MEX (BC BS)

Cerenopus concolor LeConte, 1851: 143.

Cerenopus hermanus Berry, 1975MEX (BS)

Cerenopus hermanus Berry, 1975: 931.

Cerenopus punctatus Berry, 1975MEX (BS)

Cerenopus punctatus Berry, 1975: 932.

Tribe Eulabini Horn, 1870

Eulabes Horn, 1870: 323. Type genus: Eulabis Eschscholtz, 1829.

Genus Apsena LeConte, 1862 [F]

Apsena LeConte, 1862a: 228. Type species: Eulabis pubescens LeConte, 1851, original designation.

Apsena barbarae Blaisdell, 1932USA (CA)

Apsena barbarae Blaisdell, 1932b: 61.

Apsena grossa (LeConte, 1866)USA (CA)

Eulabis grossa LeConte, 1866b: 118.

Apsena insularis Blaisdell, 1932MEX (BC)

Apsena insularis Blaisdell, 1932b: 58.

Apsena laticornis laticornis (Casey, 1891)USA (CA)

Eulabis laticornis Casey, 1891: 60.

Apsena labreae Pierce, 1954b: 98. Synonymy: Doyen and Miller (1980: 2).

Apsena laticornis subvestita Blaisdell, 1932USA (CA)

Apsena laticornis var. subvestita Blaisdell, 1932b: 68.

Apsena leachi Blaisdell, 1932USA (CA)

Apsena leachi Blaisdell, 1932b: 70.

Apsena pubescens pubescens (LeConte, 1851)USA (CA)

Eulabis pubescens LeConte, 1851: 143.

Eulabis crassicornis Casey, 1890b: 404. Synonymy: Blaisdell (1925a: 83).

Apsena pubescens rufescens Blaisdell, 1932MEX (BC)

Apsena pubescens rufescens Blaisdell, 1932b: 56.

Apsena rufipes opaca Blaisdell, 1932USA (CA)

Apsena rufipes var. opaca Blaisdell, 1932b: 81.

Apsena rufipes rufipes (Eschscholtz, 1829)USA (CA)

Eulabis rufipes Eschscholtz, 1829: 15.

Eulabis montana Casey, 1924: 330. Synonymy: Blaisdell (1925a: 84).

Apsena rufipes simplex Blaisdell, 1932USA (CA)

Apsena rufipes simplex Blaisdell, 1932b: 84.

Genus Epantius LeConte, 1851 [M]

Epantius LeConte, 1851: 144. Type species: Epantius obscurus LeConte, 1851, monotypy.

Epantius obscurus LeConte, 1851USA (CA) MEX (BC)

Epantius obscurus LeConte, 1851: 144.

Genus Eulabis Eschscholtz, 1829 [F]

Eulabis Eschscholtz, 1829: 14. Type species: Eulabis bicarinata Eschscholtz, 1829, subsequent designation (Blaisdell 1932b: 44).

Eulabis bicarinata Eschscholtz, 1829USA (CA)

Eulabis bicarinata Eschscholtz, 1829: 15.

Tribe Helopini Latreille, 1802

Helopii Latreille, 1802: 176. Type genus: Helops Fabricius, 1775.

Genus Helops Fabricius, 177543 [M]

Helops Fabricius, 1775: 257. Type species: Tenebrio caeruleus Linnaeus, 1758, subsequent designation (Hope 1841: 133) (see ICZN 2009)44.

Stenotrichus LeConte, 1862a: 239. Type species: Amphidora rufipes LeConte, 1851, original designation. Synonymy: Aalbu et al. (2002: 496).

Biomorphus Motschulsky, 1872: 38. Type species: Biomorphus tuberculatus Motschulsky, 1872 (= Amphidora attenuata LeConte, 1851), original designation. Synonymy: Aalbu et al. (1995: 485).

Coscinoptilix Allard, 1876: 15 [as Coscinopter]45. Type species: Coscinoptilix gracilicornis Allard, 1876, monotypy. Synonymy: Champion (1887: 312).

Helops angustus LeConte, 1859USA (CA)

Helops angustus LeConte, 1859b: 77.

Helops arizonensis Horn, 1874USA (AZ NM)

Helops arizonensis Horn, 1874a: 36.

Helops attenuatus (LeConte, 1851)USA (CA NV)

Amphidora attenuata LeConte, 1851: 136.

Biomorphus tuberculatus Motschulsky, 1872: 40. Synonymy: Aalbu et al. (1995: 485).

Helops bachei LeConte, 1861USA (CA)

Helops bachei LeConte, 1861b: 353.

Helops benitensis Blaisdell, 1925MEX (BC)

Helops benitensis Blaisdell, 1925b: 339.

Helops blaisdelli Casey, 1891USA (CA)

Helops blaisdelli Casey, 1891: 66.

Helops blandi Bousquet and Bouchard, 2012CAN (NB) USA (MD NJ NY SC VA)

Helops gracilis Bland, 1864: 319 [junior primary homonym of Helops gracilis Fischer von Waldheim, 1823].

Helops blandi Bousquet and Bouchard [in Nabozhenko et al.], 2012: 729. Replacement name for Helops gracilis Bland, 1864.

Helops callosus Casey, 1890USA (NM)

Helops callosa Casey, 1890b: 489.

Helops cavifrons Champion, 1887 GUA

Helops cavifrons Champion, 1887: 313.

Helops cisteloides Germar, 1823USA (AR FL GA LA MD MO NC NJ OH SC TX VA)

Helops cisteloides Germar, 1823: 159.

Helops confluens (Casey, 1924)USA (CA) Status revised [RLA]

Stenotrichus confluens Casey, 1924: 329.

Helops coxalis Champion, 1887MEX (MI)

Helops coxalis Champion, 1887: 317.

Helops crockeri Blaisdell, 1933MEX (BC [Guadalupe Is.])

Helops crockeri Blaisdell, 1933a: 89.

Helops cupripennis Champion, 1887MEX (OA)

Helops cupripennis Champion, 1887: 319.

Helops cylindriformis Casey, 1891USA (NM)

Helops cylindriformis Casey, 1891: 68.

Helops difficilis Horn, 1878USA (CO WY)

Helops difficilis Horn, 1878a: 57.

Helops discipulus Casey, 1891USA (CA)

Helops discipula Casey, 1891: 67.

Helops discretus LeConte, 1866USA (TX)

Helops discretus LeConte, 1866b: 134.

Helops edwardsii Horn, 1870USA (CA OR WA)

Helops edwardsii Horn, 1870: 395.

Helops enitescens Champion, 1893 GUA

Helops enitescens Champion, 1893a: 557.

Helops exsculptus Champion, 1887 GUA

Helops exsculptus Champion, 1887: 314.

Helops farctus LeConte, 1858USA (TX)

Helops farcta LeConte, 1858b: 74.

Helops fresnoensis Blaisdell, 1931USA (CA)

Helops fresnoënsis Blaisdell, 1931: 44.

Helops gracilicornis (Allard, 1876)MEX (VE)

Coscinoptilix gracilicornis Allard, 1876: 52.

Helops guadalupensis Casey, 1890MEX (BC [Guadalupe Is.])

Helops guadalupensis Casey, 1890b: 488.

Helops impolitus LeConte, 1866USA (TX)

Helops impolitus LeConte, 1866b: 132.

Helops inanis (Allard, 1877)MEX (MO PU)

Tarpela inanis Allard, 1877b: 262.

Helops funebris Champion, 1887: 316. Synonymy: Champion (1893a: 556).

Helops laetus LeConte, 1857CAN (BC) USA (CA OR WA)

Helops laetus LeConte, 1857: 50.

Helops longicornis Champion, 1887MEX (DU)

Helops longicornis Champion, 1887: 314.

Helops noguerai Doyen, 1990MEX (JA)

Helops noguerai Doyen, 1990: 236.

Helops obtusangulus Blaisdell, 1921USA (CA)

Helops obtusangula Blaisdell, 1921b: 228.

Helops opacus LeConte, 1859USA (CA ID NV OR UT)

Helops opacus LeConte, 1859c: 284.

Helops panamensis Champion, 1887 PAN

Helops panamensis Champion, 1887: 319.

Helops perforatus Horn, 1880USA (TX)

Helops perforatus Horn, 1880: 153.

Helops pernitens LeConte, 1861 [Fig. 25] CAN (BC) USA (CA OR WA)

Figure 25. 

Helops pernitens LeConte, 1861. Scale bar = 1 mm.

Helops pernitens LeConte, 1861b: 353.

Helops pinguis Horn, 1894MEX (BS)

Helops pinguis Horn, 1894b: 430.

Helops politus Say, 1826USA (FL)

Helops politus Say, 1826: 240.

Helops pueblensis Champion, 1887MEX (GE PU)

Helops pueblensis Champion, 1887: 317.

Helops punctatostriatus Champion, 1887 MEX

Helops punctato-striatus Champion, 1887: 316.

Helops punctatus Gemminger, 1870USA (CA)

Helops punctipennis LeConte, 1866b: 133 [junior primary homonym of Helops punctipennis Lucas, 1846].

Helops punctatus Gemminger, 1870: 123 [junior primary homonym of Helops punctatus Fabricius, 1801]46. Replacement name for Helops punctipennis LeConte, 1866.

Helops punctiventris Champion, 1887MEX (GU)

Helops punctiventris Champion, 1887: 320.

Helops rastratus Champion, 1893MEX (CH)

Helops rastratus Champion, 1893a: 557.

Helops rufipes (LeConte, 1851)USA (CA)

Amphidora rufipes LeConte, 1851: 136.

Amphidora parallela Casey, 1924: 328. Synonymy: Blaisdell (1933b: 210).

Helops rugiceps Champion, 1887 GUA

Helops rugiceps Champion, 1887: 315.

Helops rugicollis LeConte, 1866USA (CA)

Helops rugicollis LeConte, 1866b: 133.

Helops rugulosus LeConte, 1851USA (CA)

Helops rugulosus LeConte, 1851: 151.

Helops scintillatus Doyen, 1990MEX (JA NA)

Helops scintillatus Doyen, 1990: 239.

Helops seriatoporus Champion, 1893MEX (CH)

Helops seriatoporus Champion, 1893a: 558.

Helops seriatus (Allard, 1877)USA (CA)

Catomus seriatus Allard, 1877b: 46.

Helops simulator Blaisdell, 1921USA (CA OR)

Helops simulator Blaisdell, 1921b: 226.

Helops sparsus Blaisdell, 1943MEX (BC)

Helops sparsus Blaisdell, 1943: 274.

Helops spiethi Pallister, 1954MEX (DU)

Helops spiethi Pallister, 1954: 50.

Helops spilmani Pallister, 1954MEX (CH DU)

Helops spilmani Pallister, 1954: 49.

Helops spissicornis Champion, 1893MEX (DU)

Helops spissicornis Champion, 1893a: 558.

Helops spretus Horn, 1880USA (NV)

Helops spretus Horn, 1880: 153.

Helops stenotrichoides Blaisdell, 1895USA (CA)

Helops stenotrichoides Blaisdell, 1895: 238.

Helops strigicollis Horn, 1885USA (CA)

Helops strigicollis Horn, 1885c: 161.

Helops suavis Champion, 1887MEX (OA) GUA

Helops suavis Champion, 1887: 318.

Helops sulcipennis LeConte, 1866USA (AL GA NC SC TN VA)

Helops sulcipennis LeConte, 1866b: 133.

Helops sumptuosus (Allard, 1876) MEX

Diastixus sumptuosus Allard, 1876: 57.

Helops tristis Palisot de Beauvois, 1817USA (SC)

Helops tristis Palisot de Beauvois, 1817: 138 [junior primary homonym of Helops tristis Rossi, 1790].47

Helops tumescens LeConte, 1866USA (AZ CA)

Helops tumescens LeConte, 1866b: 134.

Genus Nalassus Mulsant, 1854 [M]

Nalassus Mulsant, 1854: 323. Type species: Helops dryadophilus Mulsant, 1854, subsequent designation (Nabozhenko 2001: 630).

Nalassus aereus (Germar, 1823)USA (AL CT DC DE GA IL IN KS KY MD MO MS NC NJ NY OH PA SC TN VA WV)

Helops aereus Germar, 1823: 160.

Helops pullus Say, 1826: 240. Synonymy: LeConte (1866a: 63).

Helops aratus Say, 1826: 241. Synonymy: LeConte (1866a: 63).

Helops carolina Manee, 1924: 40. Synonymy: Steiner (2009: 332).

Nalassus californicus (Mannerheim, 1843)USA (CA ID NV OR WA) MEX

Helops californicus Mannerheim, 1843: 287.

Nalassus convexulus (LeConte, 1861)CAN (AB BC) USA (CA CO ID MT NE NV OR UT WA WY)

Helops convexulus LeConte, 1861b: 353.

Helops inclusus Walker, 1866: 330. Synonymy: Blair (1921: 283).

Helops montanus LeConte, 1879b: 518. Synonymy: Bousquet and Campbell (1991: 258).

Helops regulus Blaisdell, 1921b: 227. Synonymy: Boddy (1965: 177).

Genus Nautes Pascoe, 1866 [M]

Nautes Pascoe, 1866: 475. Type species: Nautes fervidus Pascoe, 1866, monotypy.

Nautes alternans Champion, 1893 GUA

Nautes alternans Champion, 1893a: 550.

Nautes antennatus Champion, 1887 PAN

Nautes antennatus Champion, 1887: 281.

Nautes asperipennis Allard, 1894 CUB

Nautes asperipennis Allard, 1894: 259.

Nautes azurescens (Jacquelin du Val, 1857)USA (FL) / BAHCUB

Helops azurescens Jacquelin du Val, 1857: 153.

Helops viridimicans Horn, 1878a: 57. Synonymy: Steiner (2005: 454).

Nautes belti Allard, 1877 NIC PAN

Nautes belti Allard, 1877b: 59.

Nautes breviceps Champion, 1887 PAN

Nautes breviceps Champion, 1887: 282.

Nautes chrysomeloides Champion, 1887 BEL

Nautes chrysomeloides Champion, 1887: 284.

Nautes enoplopoides Champion, 1887 GUA

Nautes enoplopoides Champion, 1887: 287.

Nautes fervidus Pascoe, 1866MEX (VE) GUANIC

Nautes fervidus Pascoe, 1866: 476.

Nautes aeneus Bates, 1870: 270. Synonymy: Champion (1887: 278).

Nautes glabratus Champion, 1887MEX (VE)

Nautes glabratus Champion, 1887: 278.

Nautes guanahani Steiner, 2006 BAH

Nautes guanahani Steiner, 2006: 29.

Nautes hilaris Champion, 1887 GUA

Nautes hilaris Champion, 1887: 286.

Nautes laeviventris Champion, 1887 GUA

Nautes laeviventris Champion, 1887: 285.

Nautes magnificus Champion, 1887 GUA

Nautes magnificus Champion, 1887: 284.

Nautes nitidissimus Champion, 1887MEX (VE)

Nautes nitidissimus Champion, 1887: 286.

Nautes nodulosus Champion, 1887 GUA

Nautes nodulosus Champion, 1887: 287.

Nautes rufipes Allard, 1876 CUB

Nautes rufipes Allard, 1876: 45.

Nautes splendens Champion, 1887 PAN

Nautes splendens Champion, 1887: 280.

Nautes stabilis Champion, 1893MEX (OA VE)

Nautes stabilis Champion, 1893a: 550.

Nautes striatipennis Champion, 1887MEX (OA)

Nautes striatipennis Champion, 1887: 283.

Nautes tinctus Champion, 1887 GUA

Nautes tinctus Champion, 1887: 279.

Nautes tricolor Champion, 1893MEX (PU)

Nautes tricolor Champion, 1893a: 551.

Nautes varians Champion, 1887MEX (OA VE)

Nautes varians Champion, 1887: 281.

Nautes versicolor Champion, 1887 GUA

Nautes versicolor Champion, 1887: 284.

Genus Neohelops Dajoz, 2001 [M]

Neohelops Dajoz, 2001: 356. Type species: Neohelops texanus Dajoz, 2001, original designation.

Neohelops texanus Dajoz, 2001USA (TX)

Neohelops texanus Dajoz, 2001: 357.

Genus Tarpela Bates, 1870 [F]

Tarpela Bates, 1870: 272. Type species: Tarpela brownii Bates, 1870, subsequent designation (Gebien 1943: 407).

Lamperos Allard, 1876: 4. Type species: Helops micans Fabricius, 1798, subsequent designation (Nabozhenko and Löbl 2008: 256). Synonymy: Champion (1887: 288).

Tarpela aerifera Allard, 1876MEX (PU VE) PAN

Tarpela aerifera Allard, 1876: 47.

Tarpela allardi Champion, 1887MEX (VE)

Tarpela allardi Champion, 1887: 307.

Tarpela amabilis Champion, 1887 GUA

Tarpela amabilis Champion, 1887: 308.

Tarpela atra Allard, 1876MEX (DU JA MI PU)

Tarpela atra Allard, 1876: 46.

Tarpela azteca Champion, 1887MEX (GU)

Tarpela azteca Champion, 1887: 300.

Tarpela brownii Bates, 1870 NIC PAN

Tarpela brownii Bates, 1870: 272.

Tarpela cactivora Zayas, 1988 CUB

Tarpela cactivora Zayas, 1988: 105.

Tarpela catenata Champion, 1895MEX (YU)

Tarpela catenulata Champion, 1893a: 552 [junior primary homonym of Tarpela catenulata Allard, 1877].

Tarpela catenata Champion, 1895: 215. Unjustified emendation of Tarpela catenulata Champion, 1893.

Tarpela cisteliformis Allard, 1877 MEX GUA

Tarpela cisteliformis Allard, 1877b: 57.

Tarpela contigua Champion, 1887MEX (MI)

Tarpela contigua Champion, 1887: 298.

Tarpela corpulenta Champion, 1887MEX (DU)

Tarpela corpulenta Champion, 1887: 292.

Tarpela costata Champion, 1887MEX (GE JA SI)

Tarpela costata Champion, 1887: 293.

Tarpela crassipes Champion, 1887MEX (OA)

Tarpela crassipes Champion, 1887: 306.

Tarpela cupreoviridis Allard, 1877MEX (YU) GUANIC

Tarpela cupreo-viridis Allard, 1877b: 57.

Tarpela cuprosa Zayas, 1988: 106 CUB

Tarpela cuprosa Zayas, 1988: 106.

Tarpela depressa Champion, 1887MEX (JA YU)

Tarpela depressa Champion, 1887: 306.

Tarpela docilis Champion, 1887MEX (VE)

Tarpela docilis Champion, 1887: 312.

Tarpela durangoensis Champion, 1887MEX (DU)

Tarpela durangoensis Champion, 1887: 292.

Tarpela eximia (Bates, 1870) NIC

Nautes eximius Bates, 1870: 271.

Tarpela fallax Champion, 1887MEX (TA VE)

Tarpela fallax Champion, 1887: 301.

Tarpela flohri Champion, 1893MEX (MO)

Tarpela flohri Champion, 1893a: 553.

Tarpela foveipennis Champion, 1887MEX (CI)

Tarpela foveipennis Champion, 1887: 294.

Tarpela foveolata Champion, 1893MEX (TA)

Tarpela foveolata Champion, 1893a: 554.

Tarpela fragilicornis Champion, 1887MEX (OA)

Tarpela fragilicornis Champion, 1887: 309.

Tarpela granulipennis (Jacquelin du Val, 1857) CUB

Helops granulipennis Jacquelin du Val, 1857: 154.

Tarpela guerreroensis Champion, 1893MEX (GE)

Tarpela guerreroensis Champion, 1893a: 555.

Tarpela hispidula Allard, 1876 MEX

Tarpela hispidula Allard, 1876: 47.

Tarpela hoegei Champion, 1887MEX (DU)

Tarpela högei Champion, 1887: 297.

Tarpela inaequalis Champion, 1887 PAN

Tarpela inaequalis Champion, 1887: 290.

Tarpela incilis Champion, 1893MEX (JA)

Tarpela incilis Champion, 1893a: 553.

Tarpela jalapensis Champion, 1887MEX (GE VE)

Tarpela jalapensis Champion, 1887: 296.

Tarpela marginicollis Champion, 1887 GUA

Tarpela marginicollis Champion, 1887: 302.

Tarpela micans (Fabricius, 1798)CAN (ON QC) USA (AL CT GA IL IN MA MD NC NY OH SC TN VA)

Helops vittatus Olivier, 1793: 45 [nomen oblitum: see Appendix 4 for supporting references].

Helops micans Fabricius, 1798: 51 [nomen protectum]. Synonymy: Illiger (1802: 343).

Helops taeniatus Palisot de Beauvois, 1812: 121. Synonymy: Dejean (1821: 70).

Tarpela nigerrima Champion, 1893MEX (GE)

Tarpela nigerrima Champion, 1893a: 555.

Tarpela oblonga Champion, 1887MEX (VE)

Tarpela oblonga Champion, 1887: 298.

Tarpela oblongopunctata Bates, 1870 MEX

Tarpela oblongopunctata Bates, 1870: 273.

Tarpela occidentalis (Allard, 1877) JAM

Nesotes occidentalis Allard, 1877b: 40.

Helops mutabilis C.O. Waterhouse, 1878: 304. Synonymy: Champion (1894: lxxxv).

Tarpela propinqua (C.O. Waterhouse, 1878) JAM

Helops propinquus C.O. Waterhouse, 1878: 305.

Tarpela pulchra Champion, 1893MEX (VE)

Tarpela pulchra Champion, 1893a: 551.

Tarpela puncticeps Champion, 1887 GUA

Tarpela puncticeps Champion, 1887: 303.

Tarpela reticulata Champion, 1887 HON

Tarpela reticulata Champion, 1887: 293.

Tarpela sculptilis Champion, 1887MEX (VE)

Tarpela sculptilis Champion, 1887: 295.

Tarpela setigera Champion, 1887MEX (VE)

Tarpela setigera Champion, 1887: 297.

Tarpela silvicola Champion, 1887 GUA

Tarpela silvicola Champion, 1887: 309.

Tarpela sinuaticollis Champion, 1887 PAN

Tarpela sinuaticollis Champion, 1887: 303.

Tarpela socia Champion, 1887MEX (GE JA SI)

Tarpela socia Champion, 1887: 299.

Tarpela subparallela Champion, 1887MEX (SL)

Tarpela subparallela Champion, 1887: 300.

Tarpela subvittata Champion, 1887 GUA

Tarpela subvittata Champion, 1887: 305.

Tarpela suturalis Champion, 1887 GUA

Tarpela suturalis Champion, 1887: 310.

Tarpela teapensis Champion, 1893MEX (TB)

Tarpela teapensis Champion, 1893a: 556.

Tarpela tenuicornis Champion, 1887 GUA

Tarpela tenuicornis Champion, 1887: 289.

Tarpela thoracica Champion, 1887 NIC

Tarpela thoracica Champion, 1887: 293.

Tarpela torrida Champion, 1887MEX (DU YU)

Tarpela torrida Champion, 1887: 291.

Tarpela totonicapamensis Champion, 1887 GUA

Tarpela totonicapamensis Champion, 1887: 311.

Tarpela tropicalis Champion, 1887 GUA

Tarpela tropicalis Champion, 1887: 304.

Tarpela undulata (LeConte, 1866)USA (FL GA IN MD NC OH PA SC TN VA)

Helops americanus Palisot de Beauvois, 1812: 122 [nomen dubium]48.

Helops undulatus LeConte, 1866b: 132. Synonymy: Horn (1885a: 89).

Tarpela venusta (Say, 1824)USA (AL GA MD MO NC NY OH PA SC TN VA)

Helops venustus Say, 1824b: 284.

Tarpela veraepacis Champion, 1887 GUA

Tarpela veraepacis Champion, 1887: 295.

Tarpela virescens (Laporte, 1840) “Amérique du Nord”

Helops virescens Laporte, 1840: 235.

Tribe Melanimonini Seidlitz, 1894

Microzoumates Mulsant, 1854: 176. Type genus: Microzoum Dejean, 1834 (= Melanimon Steven 1829).

Melanimonina Seidlitz, 1894: 449. Type genus: Melanimon Steven, 1829. Note.Use of younger family-group name conserved (ICZN 1999: Art. 40.2) (see Bouchard et al. 2005).

Genus Cheirodes Gené, 1839 [M]

Cheirodes Gené, 1839: 73. Type species: Cheirodes sardous Gené, 1839, monotypy.

Anemia Laporte, 1840: 218. Type species: Anemia granulata Laporte, 1840, monotypy. Synonymy: Spilman (1973: 41).

Chirodes Agassiz, 1846: 81. Unjustified emendation of Cheirodes Gené, 1839, not in prevailing usage.

Cheirodes californicus (Horn, 1870)USA (CA NV OR WA)

Anaemia californica Horn, 1870: 378.

Tribe Metaclisini Steiner, 2016

Metaclisini Steiner, 2016: 542. Type genus: Metaclisa Jacquelin du Val, 1861.

Genus Metaclisa Jacquelin du Val, 1861 [F]

Amarantha Motschulsky, 1859: 141 [nomen oblitum, see Bouchard et al. (2007: 393)]. Type species: Amarantha viridis Motschulsky, 1859, monotypy.

Metaclisa Jacquelin du Val, 1861: 296 [nomen protectum]. Type species: Platydema parallela Fairmaire, 1855 (= Diaperis azurea Waltl, 1838), original designation. Synonymy: Lewis (1891: 70).

Tharsus LeConte, 1862a: 233. Type species: Tharsus seditiosus LeConte, 1862, monotypy. Synonymy: Steiner (2016: 538).

Metaclisa atra LeConte, 1866USA (AL DC FL GA LA MD MO MS NC PA SC TX VA)

Metaclisa atra LeConte, 1866b: 127.

Haplandrus collaris Casey, 1924: 320. Synonymy: Steiner (2016: 537).

Haplandrus subangusta Casey, 1924: 320. Synonymy: Steiner (2016: 537).

Metaclisa marginalis Horn, 1870CAN (BC) USA (CA OR WA)

Metaclisa marginalis Horn, 1870: 369.

Metaclisa seditiosa (LeConte, 1862)USA (AL FL GA KY MD NC OH SC TN TX VA WV) / BAH

Tharsus seditiosus LeConte, 1862a: 233.

Tribe Opatrini Brullé, 1832

Opatrites Brullé, 1832: 213. Type genus: Opatrum Fabricius, 1775.

Subtribe Opatrina Brullé, 1832

Opatrites Brullé, 1832: 213. Type genus: Opatrum Fabricius, 1775.

Blapstinites Mulsant and Rey, 1853: 258. Type genus: Blapstinus Dejean, 1821.

Genus Aconobius Casey, 1895 [M]

Aconobius Casey, 1895: 617. Type species: Conibiosoma laciniata Casey, 1891, original designation.

Aconobius densus Casey, 1914USA (NM)

Aconobius densus Casey, 1914: 377.

Aconobius laciniatus (Casey, 1891)USA (AZ)

Conibiosoma laciniata Casey, 1891: 64.

Aconobius nigripes Casey, 1914USA (TX)

Aconobius nigripes Casey, 1914: 378.

Genus Ammodonus Mulsant and Rey, 1859 [M]

Ammodonus Mulsant and Rey, 1859: 143. Type species: Opatrum fossor LeConte, 1847, monotypy.

Pseudonomus Fairmaire, 1884: 510. Type species: Pseudonomus dermestiformis Fairmaire, 1884, monotypy. Synonymy: Gebien (1939: 470).

Scaptes Champion, 1886: 222. Type species: Scaptes squamulatus Champion, 1886 (= Asida tropica Kirsch, 1866), present designation. Synonymy: Fall (1912: 48).

Trichotoides Marcuzzi, 1954b: 23. Type species: Scaptes hintoni Kaszab, 1949, monotypy. Synonymy: Ferrer and Moraguès (2001: 499).

Ammodonus ciliatus (Champion, 1896)LAN / SA

Scaptes ciliatus Champion, 1896: 9.

Ammodonus fossor (LeConte, 1847)CAN (ON) USA (AL AR DE IL IN KS MD MN NC NE NJ NY OH OK SC TX WI WV)

Opatrum fossor LeConte, 1847: 92.

Ammodonus granosus Fall, 1912USA (AZ) MEX (BS)

Ammodonus granosus Fall, 1912: 47.

Ammodonus tropicus (Kirsch, 1866)USA (AZ CA) MEX (AG CH CI JA MI NA NL OA PU SI SO VE) GUABELSALHONNICCRIPAN / CUBJAM / SA

Asida tropica Kirsch, 1866: 190.

Scaptes squamulatus Champion, 1886: 223. Synonymy: Champion (1893a: 542).

Genus Blapstinus Dejean, 1821 [M]

Blapstinus Dejean, 1821: 66. Type species: Blaps punctata Fabricius, 1792, monotypy.

Heteropus Laporte, 1840: 221 [junior homonym of Heteropus Palisot de Beauvois, 1820]. Type species: Heteropus holosericeus Laporte, 1840, monotypy. Synonymy: Lacordaire (1859: 250).

Pedonoeces G.R. Waterhouse, 1845: 32. Type species: Pedonoeces galapagoensis G.R. Waterhouse, 1845, subsequent designation (Aalbu and Triplehorn 1991: 170). Synonymy: Aalbu and Triplehorn (1991: 170).

Tessaromma Boheman, 1858: 91 [junior homonym of Tessaromma Newman, 1840]. Type species: Tessaromma lugubris Boheman, 1858, subsequent designation (Aalbu and Triplehorn 1991: 170). Synonymy: Aalbu and Triplehorn (1991: 170).

Lachnoderes Mulsant and Rey, 1859: 166. Type species: Pedonoeces pubescens G.R. Waterhouse, 1845, monotypy. Synonymy: Aalbu and Triplehorn (1991: 170).

Aspidius Mulsant and Rey, 1859: 187. Type species: Blaps punctata Fabricius, 1792, present designation. Synonymy: Champion (1885: 124).

Lodinus Mulsant and Rey, 1859: 195. Type species: Lodinus nigroaeneus Mulsant and Rey, 1859 (= Blapstinus punctulatus Solier, 1851), monotypy. Synonymy: Gemminger [in Gemminger and Harold] (1870: 1923).

Blapstinus aciculus Blatchley, 1917USA (FL) / BAH

Blapstinus aciculus Blatchley, 1917: 275.

Blapstinus amnosus Blaisdell, 1923MEX (BC)

Blapstinus amnosus Blaisdell, 1923: 272.

Blapstinus angustatus Champion, 1893MEX (OA)

Blapstinus angustatus Champion, 1893a: 528.

Blapstinus aridus Blaisdell, 1923MEX (SO)

Blapstinus aridus Blaisdell, 1923: 270.

Blapstinus atratus Champion, 1885MEX (GE PU YU) GUANICPAN

Blapstinus atratus Champion, 1885: 131.

Blapstinus auripilis Horn, 1870USA (AZ)

Blapstinus auripilis Horn, 1870: 353.

Blapstinus barri Boddy, 1957USA (ID OR)

Blapstinus barri Boddy, 1957: 198.

Blapstinus brevicollis LeConte, 1851USA (AZ CA) MEX (SO)

Blapstinus brevicollis LeConte, 1851: 147.

Blapstinus sonorae Casey, 1890b: 431. New synonymy [based on Davis (1970: 131) unpublished thesis].

Blapstinus buqueti Champion, 1885CRIPAN /LAN / SA

Blapstinus buqueti Champion, 1885: 128.

Blapstinus piliferus Fairmaire, 1892: 82. Synonymy: Marcuzzi (1951: 75).

Blapstinus castaneus Casey, 1890USA (AZ CA TX)

Blapstinus castaneus Casey, 1890b: 432.

Blapstinus falli Blaisdell, 1929b: 21. New synonymy [based on Davis (1970: 257) unpublished thesis].

Blapstinus cubanus cubanus Marcuzzi, 1962 CUB CAY

Blapstinus cubanus Marcuzzi, 1962: 33.

Blapstinus cubanus grandturki Marcuzzi, 1965 BAH

Blapstinus cubanus grandturki Marcuzzi, 1965: 130.

Blapstinus cylindriformis Doyen, 1990MEX (GE JA)

Blapstinus cylindriformis Doyen, 1990: 232.

Blapstinus decui Ardoin, 1977 CUB

Blapstinus decui Ardoin, 1977c: 390.

Blapstinus debilis Casey, 1890USA (FL TX)

Blapstinus debilis Casey, 1890b: 458.

Blapstinus densipunctatus Blaisdell, 1943MEX (BS)

Blapstinus densipunctatus Blaisdell, 1943: 256.

Blapstinus dilatatus LeConte, 1851USA (AZ CA CO SD UT) MEX (CH SO)

Opatrum pullum Say, 1826: 237 [nomen dubium].

Blapstinus dilatatus LeConte, 1851: 146. Synonymy (in doubt): LeConte (1866a: 61).

Blapstinus discolor Horn, 1870CAN (BC) USA (CA ID NV OR UT WA) MEX (BS)

Blapstinus discolor Horn, 1870: 354.

Blapstinus oregonensis Casey, 1890b: 435. Synonymy: Davis (1982: 254).

Blapstinus fuliginosus Casey, 1890b: 438. Synonymy: Davis (1982: 254).

Blapstinus rufipes Casey, 1890b: 439. Synonymy: Davis (1982: 254).

Blapstinus crassicornis Casey, 1890b: 440. Synonymy: Davis (1982: 254).

Blapstinus elongatus Casey, 1890b: 441. Synonymy: Horn (1894b: 351).

Blapstinus lepidus Casey, 1890b: 444. Synonymy: Davis (1982: 254).

Blapstinus aequalis Casey, 1890b: 445. Synonymy: Davis (1982: 254).

Blapstinus funebris Casey, 1890b: 446. Synonymy: Davis (1982: 254).

Blapstinus parallelus Casey, 1890b: 448. Synonymy: Davis (1982: 254).

Blapstinus inquisitus Casey, 1890b: 449. Synonymy: Davis (1982: 254).

Blapstinus domingoensis (Marcuzzi, 1998) DOM

Diastolinus domingoensis Marcuzzi, 1998b: 222.

Blapstinus dominicus Marcuzzi, 1962 JAM HAI PRI LAN

Blapstinus dominicus Marcuzzi, 1962: 34.

Blapstinus egenus Champion, 1885MEX (JA SI VE) GUANICPAN / SA

Blapstinus egenus Champion, 1885: 129.

Blapstinus emmenastoides Champion, 1885MEX (OA VE) GUA

Blapstinus emmenastoides Champion, 1885: 131.

Blapstinus errabundus Champion, 1885MEX (OA VE YU) NICPAN

Blapstinus errabundus Champion, 1885: 127.

Blapstinus exiguus Champion, 1893MEX (OA YU)

Blapstinus exiguus Champion, 1893a: 529.

Blapstinus faulkneri Aalbu and Triplehorn, 1991MEX (“Revillagigedo Is.”)

Blapstinus faulkneri Aalbu and Triplehorn, 1991: 173.

Blapstinus fortis LeConte, 1878USA (AZ CO FL GA KS LA MD MO NC NM OK SC SD TX) MEX (CO JA MO OA SL VE YU) GUABELNICCRIPAN / BAHCUBCAY

Opatrinus punctulatus Jacquelin du Val, 1857: 141 [junior secondary homonym of Blapstinus punctulatus Solier, 1851].

Blapstinus fortis LeConte, 1878a: 420. Synonymy: Casey (1890b: 429).

Blapstinus interstitialis Champion, 1885: 125. Replacement name for Blapstinus punctulatus (Jacquelin du Val, 1857).

Blapstinus fuscus Casey, 1890USA (FL LA OK TX [NM]) MEX

Blapstinus fuscus Casey, 1890b: 427.

Blapstinus genaroi (Garrido, 2004) DOM

Diastolinus genaroi Garrido, 2004b: 41.

Blapstinus grandis Champion, 1885MEX (JA SI) NICCRI

Blapstinus grandis Champion, 1885: 125.

Blapstinus haitensis Marcuzzi, 1962 HAI DOM

Blapstinus haitensis Marcuzzi, 1962: 34.

Blapstinus hispaniolensis (Marcuzzi, 1998) DOM

Diastolinus hispaniolensis Marcuzzi, 1998b: 220.

Blapstinus histricus Casey, 1890USA (AZ CA FL NV NM TX)

Blapstinus histricus Casey, 1890b: 433.

Blapstinus brunneus Casey, 1890b: 453. New synonymy [based on Davis (1970: 220) unpublished thesis].

Blapstinus coronadensis Blaisdell, 1892: 242. New synonymy [based on Davis (1970: 220) unpublished thesis].

Blapstinus humilis Casey, 1890USA (FL) / BAH

Blapstinus humilis Casey, 1890b: 459.49

Blapstinus inflatitibia (Marcuzzi, 1977) CAY

Diastolinus inflatitibia Marcuzzi, 1977: 17.

Blapstinus insularis Champion, 1885 PAN

Blapstinus insularis Champion, 1885: 127.

Blapstinus intermedius Champion, 1885 MEX GUA NIC

Blapstinus intermedius Champion, 1885: 129.

Blapstinus intermixtus Casey, 1890CAN (BC) USA (AZ CA ID OR UT WA)

Blapstinus intermixtus Casey, 1890b: 451.

Blapstinus hesperius Casey, 1890b: 454. New synonymy [based on Davis (1970: 232) unpublished thesis].

Blapstinus jamaicensis Marcuzzi, 1962 JAM

Blapstinus jamaicensis Marcuzzi, 1962: 35.

Blapstinus kalik Steiner, 2006 BAH

Blapstinus kalik Steiner, 2006: 18.

Blapstinus kaszabi Marcuzzi, 1985 “Ciudad, Central America”

Blapstinus kaszabi Marcuzzi, 1985: 183.

Blapstinus klapperichi (Marcuzzi, 1998) DOM

Diastolinus klapperichi Marcuzzi, 1998b: 219.

Blapstinus lecontei Mulsant and Rey, 1859USA (CA)

Blapstinus pubescens LeConte, 1851: 147 [junior secondary homonym of Blapstinus pubescens (G.R. Waterhouse, 1845)].

Blapstinus lecontii Mulsant and Rey, 1859: 192. Replacement name for Blapstinus pubescens LeConte, 1851.

Blapstinus cinerascens Fall, 1929: 58. New synonymy [based on Davis (1970: 100) unpublished thesis].

Blapstinus californicus Aalbu and Triplehorn, 1991: 170 [junior primary homonym of Blapstinus californicus Motschulsky, 1845]. Replacement name for Blapstinus pubescens LeConte, 1851.

Blapstinus longicollis Champion, 1885 GUA NIC

Blapstinus longicollis Champion, 1885: 126.

Blapstinus longipennis Champion, 1885MEX (SI)

Blapstinus longipennis Champion, 1885: 130.

Blapstinus longulus LeConte, 1851USA (AZ)

Blapstinus longulus LeConte, 1851: 147.

Blapstinus marcuzzii Aalbu, new replacement name JAM

Blapstinus kulzeri Marcuzzi, 1977: 28 [junior primary homonym of Blapstinus kulzeri Kaszab, 1969]50.

Blapstinus marcuzzii Aalbu, new replacement name for Blapstinus kulzeri Marcuzzi, 1977.

Blapstinus metallicus (Fabricius, 1801)CAN (AB MB NB NS ON PE QC SK) USA (CO CT DE FL GA IA IL IN KS LA MA MD MI MN MO MS NC ND NE NH NJ NY OH OR PA RI SC SD VA WI WV WY)

Blaps metallica Fabricius, 1801a: 143.

Opatrum interruptum Say, 1824a: 264. Synonymy: LeConte (1866a: 61).

Blapstinus aeneolus Melsheimer, 1846: 66. Synonymy: LeConte (1866a: 61).

Blapstinus luridus Mulsant and Rey, 1859: 193. Synonymy: LeConte (1866a: 61).

Blapstinus mexicanus Champion, 1885MEX (CI YU)

Blapstinus mexicanus Champion, 1885: 124.

Blapstinus moestus Melsheimer, 1846CAN (ON) USA (CT DC DE GA IL IN MA MD MI NC NH NJ NY OH RI SC VA WI)

Blapstinus moestus Melsheimer, 1846: 65.

Blapstinus nitidus Champion, 1885MEX (VE)

Blapstinus nitidus Champion, 1885: 130.

Blapstinus obliteratus Champion, 1885 PAN

Blapstinus obliteratus Champion, 1885: 132.

Blapstinus opacus Mulsant and Rey, 1859 51 LAN

Blapstinus opacus Mulsant and Rey, 1859: 186.

Blapstinus opacus martinensis Marcuzzi, 1977: 29. Synonymy: Ivie and Hart (2016: 466).

Blapstinus orlandoi Ivie and Hart, 2016 JAM

Diastolinus jamaicensis Garrido, 2004a: 37 [junior secondary homonym of Blapstinus jamaicensis Marcuzzi, 1962].

Blapstinus orlandoi Ivie and Hart, 2016: 466. Replacement name for Blapstinus jamaicensis (Garrido, 2004).

Blapstinus pacificus Aalbu and Triplehorn, 1991MEX (“Revillagigedo Is.”)

Blapstinus pacificus Aalbu and Triplehorn, 1991: 171.

Blapstinus palmeri Champion, 1885MEX (CH CO NL)

Blapstinus palmeri Champion, 1885: 128.

Blapstinus paradoxus Blaisdell, 1923MEX (SO)

Blapstinus paradoxus Blaisdell, 1923: 271.

Blapstinus pimalis Casey, 1885USA (AZ CA CO NM NV TX UT) MEX (SO)

Blapstinus pimalis Casey, 1885 [January]: 185.

Blapstinus umbrosus Champion, 1885 [October]: 127. Synonymy: Champion (1893a: 527).

Blapstinus niger Casey, 1890b: 436. New synonymy [based on Davis (1970: 306) unpublished thesis].

Blapstinus cribricollis Casey, 1890b: 437. New synonymy [based on Davis (1970: 306) unpublished thesis].

Blapstinus pinorum Casey, 1914USA (GA NC SC)

Blapstinus pinorum Casey, 1914: 377.

Blapstinus pratensis LeConte, 1859CAN (AB) USA (CO KS MT NE NM OK SD TX) MEX (CH DU SO TA)

Blapstinus pratensis LeConte, 1859a: 15.

Blapstinus arenarius Casey, 1890b: 457. New synonymy [based on Davis (1970: 177) unpublished thesis].

Blapstinus puertoricensis (Marcuzzi, 1977) PRI LAN

Diastolinus puertoricensis Marcuzzi, 1977: 20.

Blapstinus pulverulentus Mannerheim, 1843USA (CA OR WA)

Blapstinus pulverulentus Mannerheim, 1843: 276.

Blapstinus californicus Motschulsky, 1845a: 77. Synonymy: Horn (1870: 355).

Blapstinus punctatus anxius Mulsant and Rey, 1859 [no locality given originally but probably from the Antilles]

Blapstinus punctatus var. anxius Mulsant and Rey, 1859: 190.

Blapstinus punctatus punctatus (Fabricius, 1792) PRI LAN

Blaps punctata Fabricius, 1792a: 109.

Diastolinus fuscicornis Chevrolat, 1877e: viii. Synonymy: Ivie and Hart (2016: 467).

Blapstinus puncticollis Champion, 1893MEX (GE)

Blapstinus puncticollis Champion, 1893a: 529.

Blapstinus simulans barbadensis Marcuzzi, 1962 LAN

Blapstinus simulans barbadensis Marcuzzi, 1962: 36.

Blapstinus simulans simulans Marcuzzi, 1954LAN / SA

Blapstinus simulans Marcuzzi, 1954b: 15.

Blapstinus striatulus Mulsant and Rey, 1859PRILAN (St. Barthélemy)

Blapstinus striatulus Mulsant and Rey, 1859: 183.

Blapstinus striatus Guérin-Méneville, 1831 CUB

Blapstinus striatus Guérin-Méneville, 1831a: pl. 4.

Blapstinus substriatus Champion, 1885 [Fig. 26] CAN (AB BC MB SK YT) USA (AZ CA CO ID KS MN MT ND NM NV OR SD TX UT WA WY) MEX (CH CO DU FD GU ME PU SL VE) NIC

Figure 26. 

Blapstinus substriatus Champion, 1885. Scale bar = 1 mm.

Blapstinus substriatus Champion, 1885: 128.

Blapstinus gregalis Casey, 1890b: 442. New synonymy [based on Davis (1970: 232) unpublished thesis].

Blapstinus sulcatus LeConte, 1851USA (AZ CA CO KS NV UT)

Blapstinus sulcatus LeConte, 1851: 147.

Blapstinus hydropicus Casey, 1890b: 461. New synonymy [based on Davis (1970: 106) unpublished thesis].

Blapstinus sulcipennis Champion, 1885MEX (YU) GUA / CUBCAY

Blapstinus sulcipennis Champion, 1885: 129.

Blapstinus tibialis Champion, 1885 GUA NIC CRI

Blapstinus tibialis Champion, 1885: 125.

Blapstinus validus Casey, 1890USA (AZ CA)

Blapstinus validus Casey, 1890b: 429.

Blapstinus vandykei Blaisdell, 1942USA (AZ CA NM NV TX)

Blapstinus vandykei Blaisdell, 1942: 136.

Blapstinus vestitus LeConte, 1859USA (CO KS SD TX UT WY)

Blapstinus vestitus LeConte, 1859a: 15.

Blapstinus hospes Casey, 1890b: 455. New synonymy [based on Davis (1970: 187) unpublished thesis].

Blapstinus yucatanus Champion, 1893MEX (YU)

Blapstinus yucatanus Champion, 1893a: 526.

Genus Bycrea Pascoe, 1868 [F]

Bycrea Pascoe, 1868: xii. Type species: Bycrea villosa Pascoe, 1868, monotypy.

Bycrea villosa Pascoe, 1868USA (AZ) MEX (DU GE GU JA MI MO NL OA PU SI SO TA YU) GUASALCRI / SA

Bycrea villosa Pascoe, 1868: xii.

Genus Cenophorus Mulsant and Rey, 1859 [M]

Cenophorus Mulsant and Rey, 1859: 177. Type species: Cenophorus viduus Mulsant and Rey, 1859, monotypy.

Cenophorus viduus Mulsant and Rey, 1859 HAI

Cenophorus viduus Mulsant and Rey, 1859: 177.

Genus Conibiosoma Casey, 1890 [N]

Conibiosoma Casey, 1890b: 476. Type species: Conibius elongatus Horn, 1870, monotypy.

Conibiosoma elongatum (Horn, 1870)USA (CA NV)

Conibius elongatus Horn, 1870: 351.

Genus Conibius LeConte, 1851 [M]

Conibius LeConte, 1851: 145. Type species: Conibius seriatus LeConte, 1851, subsequent designation (Lucas 1920: 199).

Ooconibius Casey, 1895: 618. Type species: Notibius opacus LeConte, 1866, monotypy. New synonymy [RLA].

Euconibius Casey, 1895: 618. Type species: Notibius gagates Horn, 1870, monotypy. New synonymy [RLA].

Conibius brunnipes Champion, 1885MEX (HI JA ME MI MO NA OA PU QU SI TL) GUA

Conibius brunnipes Champion, 1885: 133.

Conibius gagates (Horn, 1870)USA (AZ) MEX (CH SO)

Notibius gagates Horn, 1870: 357.

Conibius guadalupensis Casey, 1890MEX (BC [Guadalupe Is.])

Conibius guadalupensis Casey, 1890b: 470.

Conibius oblongus Blaisdell, 1943MEX (BC)

Conibius oblongus Blaisdell, 1943: 257.

Conibius opacus (LeConte, 1866)USA (AZ) MEX (BC BS SO)

Notibius opacus LeConte, 1866b: 118.

Notibius reflexus Horn, 1894b: 429. New synonymy [RLA].

Conibius rotundicollis Linell, 1899USA (TX) MEX (HI NL)

Conibius rotundicollis Linell, 1899: 182.

Conibius rugipes (Champion, 1885)MEX (OA PU)

Notibius rugipes Champion, 1885: 132.

Notibius affinis Champion, 1885: 132. New synonymy [RLA].

Conibius seriatus LeConte, 1851USA (CA OR) MEX (BC)

Conibius seriatus LeConte, 1851: 146.

Conibius parallelus LeConte, 1851: 146. New synonymy [RLA].

Conibius troglodytes Champion, 1893MEX (GE MO PU)

Conibius troglodytes Champion, 1893a: 530.

Conibius uniformis Casey, 1890USA (AZ NM TX) MEX (CH CO DU ZA)

Conibius uniformis Casey, 1890b: 471.

Conibius ventralis Blaisdell, 1923MEX (BS)

Conibius ventralis Blaisdell, 1923: 274.

Genus Cybotus Casey, 1890 [M]

Cybotus Casey, 1890b: 482. Type species: Blapstinus estriatus LeConte, 1878, monotypy.

Cybotus estriatus (LeConte, 1878)USA (FL) MEX (QR) HON

Blapstinus estriatus LeConte, 1878a: 420.

Genus Diastolinus Mulsant and Rey, 1859 [M]

Diastolinus Mulsant and Rey, 1859: 138. Type species: Blaps clathrata Fabricius, 1792, subsequent designation (Lucas 1920: 236).

Sellio Mulsant and Rey, 1859: 169. Type species: Blaps tibidens Quensel, 1806, subsequent designation (Gebien 1938: 407). Synonymy: Ivie and Hart (2016: 468).

Ctesicles Champion, 1896: 7. Type species: Ctesicles insularis Champion, 1896, subsequent designation (Lucas 1920: 214). Synonymy: Ivie and Hart (2016: 468).

Diastolinus azuaensis Hart and Ivie, 2016 DOM

Diastolinus azuaensis Hart and Ivie, 2016a: 506.

Diastolinus clathratus (Fabricius, 1792)VIS (St. Croix)

Blaps clathrata Fabricius, 1792a: 109.

Diastolinus chalumeaui Hart and Ivie, 2016 LAN

Diastolinus chalumeaui Hart and Ivie, 2016a: 494.

Diastolinus clavatus Mulsant and Rey, 1859 PRI LAN

Diastolinus clavatus Mulsant and Rey, 1859: 155.

Diastolinus hummelincki Marcuzzi, 1962: 28 [junior primary homonym of Diastolinus hummelincki Marcuzzi, 1949]. Synonymy: Hart and Ivie (2016a: 497).

Diastolinus mulsanti Marcuzzi and d’Aguilar, 1971: 79. Replacement name for Diastolinus hummelincki Marcuzzi, 1962.

Diastolinus coarctatus (Mulsant and Rey, 1859) DOM

Sellio coarctatus Mulsant and Rey, 1859: 170.

Diastolinus estebani Garrido, 2004b: 42. Synonymy: Hart and Ivie (2016a: 507).

Diastolinus desecheo Hart and Ivie, 2016PRI (Desecheo Island)

Diastolinus desecheo Hart and Ivie, 2016a: 509.

Diastolinus doyeni Hart and Ivie, 2016 PRI

Diastolinus doyeni Hart and Ivie, 2016a: 511.

Diastolinus espoloni Garrido, 2007 DOM

Diastolinus espoloni Garrido, 2007: 46.

Diastolinus gladiator (Garrido, 2005) DOM

Sellio gladiator Garrido, 2005: 120.

Diastolinus hoppae Hart and Ivie, 2016 LAN

Diastolinus hoppae Hart and Ivie, 2016a: 522.

Diastolinus insularis (Champion, 1896)LAN (St. Vincent)

Ctesicles insularis Champion, 1896: 7.

Diastolinus leewardensis Hart and Ivie, 2016 LAN

Diastolinus leewardensis Hart and Ivie, 2016a: 498.

Diastolinus maritimus (Champion, 1896) LAN

Ctesicles maritimus Champion, 1896: 8.

Diastolinus perforatus (Schönherr, 1806) LAN

Opatrum perforatum Schönherr, 1806: 146.

Diastolinus realinoi Marcuzzi, 2002 [CUB]52

Diastolinus realinoi Marcuzzi, 2002: 398.

Diastolinus shieli Hart and Ivie, 2016LAN (Redonda)

Diastolinus shieli Hart and Ivie, 2016a: 504.

Diastolinus tibidens (Quensel, 1806) PRI LAN

Blaps tibidens Quensel [in Schönherr], 1806: 147.

Diastolinus vaderi Hart and Ivie, 2016 HAI

Diastolinus vaderi Hart and Ivie, 2016a: 518.

Diastolinus victori Garrido, 2002 PRI

Diastolinus elongatus Marcuzzi, 1977: 15 [junior primary homonym of Diastolinus elongatus Marcuzzi, 1976].

Diastolinus victori Garrido, 2002: 39. Replacement name for Diastolinus elongatus Marcuzzi, 1977.

Genus Ephalus LeConte, 1862 [M]

Ephalus LeConte, 1862a: 228. Type species: Heliopates latimanus LeConte, 1847, monotypy.

Ephalus latimanus (LeConte, 1847)CAN (NS) USA (CT MA ME NH NJ NY RI)

Heliopates latimanus LeConte, 1847: 92.

Genus Gonocephalum Solier, 1834 [N]

Gonocephalum Solier, 1834: 498. Type species: Opatrum fuscum Herbst, 1793 (= Opatrum rusticum Olivier, 1811), subsequent designation (Gebien 1939: 443).

Subgenus Gonocephalum Solier, 1834

Gonocephalum Solier, 1834: 498. Type species: Opatrum fuscum Herbst, 1793 (= Opatrum rusticum Olivier, 1811), subsequent designation (Gebien 1939: 443).

Gonocephalum sericeum (Baudi di Selve, 1875)USA (CA) – Adventive

Opatrum sericeum Baudi di Selve, 1875: 701.

Genus Hummelinckia Marcuzzi, 1954 [F]

Hummelinckia Marcuzzi, 1954b: 19. Type species: Hummelinckia caraibica Marcuzzi, 1954, monotypy.

Hummelinckia caraibica Marcuzzi, 1954 LAN

Hummelinckia caraibica Marcuzzi, 1954b: 19.

Genus Mecysmus Horn, 1870 [M]

Mecysmus Horn, 1870: 349. Type species: Blapstinus angustus LeConte, 1851, monotypy.

Mecysmus advena Casey, 1890USA (SD TX)

Mecysmus advena Casey, 1890b: 466.

Mecysmus angustus (LeConte, 1851)USA (AZ CA)

Blapstinus angustus LeConte, 1851: 147.

Mecysmus laticollis Casey, 1890CAN (AB) USA (TX)

Mecysmus laticollis Casey, 1890b: 463.

Mecysmus parvulus Casey, 1890USA (NM TX)

Mecysmus parvulus Casey, 1890b: 466.

Mecysmus tenuis Casey, 1890USA (CA)

Mecysmus tenuis Casey, 1890b: 465.

Genus Nevisia Marcuzzi, 1986 [F]

Nevisia Marcuzzi, 1985: 179. Type species: Diastolinus barbudensis Marcuzzi, 1962, monotypy.

Nevisia barbudensis (Marcuzzi, 1962) LAN

Diastolinus barbudensis Marcuzzi, 1962: 29.

Diastolinus barbudensis antiguanus Marcuzzi, 1962: 30. Synonymy: Ivie and Hart (2016: 469).

Genus Nocibiotes Casey, 1895 [M]

Nocibiotes Casey, 1895: 617. Type species: Notibius granulatus LeConte, 1851, subsequent designation (Gebien 1938: 407).

Nocibiotes caudatus Casey, 1895USA (AZ) MEX (BC)

Nocibiotes caudatus Casey, 1895: 619, 621.

Nocibiotes rubripes Casey, 1895: 619. New synonymy [RLA].

Nocibiotes crassipes (Casey, 1890)USA (CA)

Conibius crassipes Casey, 1890b: 475.

Nocibiotes granulatus (LeConte, 1851)USA (AZ CA) MEX (BS SO)

Notibius granulatus LeConte, 1851: 145.

Nocibiotes gracilis Casey, 1895: 619. New synonymy [RLA].

Nocibiotes acutus Casey, 1895: 619, 620. New synonymy [RLA].

Nocibiotes rossi (Blaisdell, 1943)MEX (BS)

Tonibius rossi Blaisdell, 1943: 260.

Genus Notibius LeConte, 1851 [M]

Notibius LeConte, 1851: 144. Type species: Notibius puberulus LeConte, 1851, subsequent designation (Gebien 1938: 406).

Notibius puberulus LeConte, 1851USA (AZ CA NV OR UT) MEX (BC)

Notibius puberulus LeConte, 1851: 145.

Notibius substriatus Casey, 1890b: 479. Synonymy: Horn (1894a: 41).

Notibius laticeps Casey, 1890b: 480. Synonymy: Horn (1894a: 41).

Notibius puncticollis LeConte, 1851USA (CA NV UT)

Notibius puncticollis LeConte, 1851: 145.

Genus Opatroides Brullé, 1832 [M]

Opatroides Brullé, 1832: 219. Type species: Opatroides punctulatus Brullé, 1832, monotypy.

Opatroides punctulatus Brullé, 1832USA (CA NV) – Adventive

Opatroides punctulatus Brullé, 1832: 220.

Genus Penichrus Champion, 1885 [M]

Penichrus Champion, 1885: 134. Type species: Penichrus blapstinoides Champion, 1885, monotypy.

Penichrus blapstinoides Champion, 1885 PAN

Penichrus blapstinoides Champion, 1885: 135.

Genus Platylus Mulsant and Rey, 1859 [M]

Platylus Mulsant and Rey, 1859: 134. Type species: Blaps dilatata Fabricius, 1798, monotypy.

Platylus dilatatus (Fabricius, 1798)PRI (Vieques) VIS (St. Thomas)

Blaps dilatata Fabricius, 1798: 47.

Genus Pseudephalus Casey, 1924 [M]

Pseudephalus Casey, 1924: 333. Type species: Pseudephalus brevicornis Casey, 1924, original designation.

Pseudephalus brevicornis Casey, 1924USA (AL FL) HONCRI

Pseudephalus brevicornis Casey, 1924: 333.

Genus Tonibiastes Casey, 1895 [M]

Tonibiastes Casey, 1895: 617. Type species: Notibius costipennis Horn, 1894, original designation.

Tonibiastes costipennis (Horn, 1894)MEX (BC BS)

Notibius costipennis Horn, 1894: 430.

Genus Tonibius Casey, 1895 [M]

Tonibius Casey, 1895: 617. Type species: Notibius sulcatus LeConte, 1851, subsequent designation (Lucas 1920: 644).

Tonibius sulcatus (LeConte, 1851)USA (CA) MEX (BC BS SO)

Notibius sulcatus LeConte, 1851: 145.

Conibius alternatus Casey, 1890b: 473. New synonymy [RLA].

Genus Trichoton Hope, 1841 [N]

Trichoton Hope, 1841: 111. Type species: Trichoton cayennense Hope, 1841, original designation.

Subgenus Trichoton Hope, 1841

Trichoton Hope, 1841: 111. Type species: Trichoton cayennense Hope, 1841, original designation.

Epilasium Curtis, 1844: 222. Type species: Epilasium rotundatum Curtis, 1844, monotypy. Synonymy: Mulsant and Rey (1853: 245).

Trichoton curvipes Champion, 1885PAN / SA

Trichoton curvipes Champion, 1885: 136.

Trichoton lapidicola Champion, 1885NIC / SA

Trichoton lapidicola Champion, 1885: 136.

Trichoton marcuzzi Kulzer, 1961LAN / SA

Trichoton marcuzzi Kulzer, 1961a: 212.

Trichoton mexicanum Kulzer, 1961MEX (CL)

Trichoton mexicanum Kulzer, 1961a: 211.

Trichoton sordidum (LeConte, 1851)USA (AZ CA NV) MEX (BC JA SO) NIC

Blapstinus sordidus LeConte, 1851: 146.

Genus Ulus Horn, 1870 [M]

Ulus Horn, 1870: 358. Type species: Blapstinus crassus LeConte, 1851, subsequent designation (Lucas 1920: 665).

Ulus comatus Champion, 1893MEX (VE)

Ulus comatus Champion, 1893a: 530.

Ulus crassus (LeConte, 1851)USA (AZ CA UT) MEX (BS YU)

Blapstinus crassus LeConte, 1851: 146.

Ulus elongatulus Casey, 1890USA (AZ TX)

Ulus elongatulus Casey, 1890b: 414.

Ulus fimbriatus Casey, 1890USA (TX) MEX (CH)

Ulus fimbriatus Casey, 1890b: 413.

Ulus hirsutus Champion, 1885MEX (CH DU JA PU SI VE YU) GUABELNICCRIPAN / CAYJAM

Ulus hirsutus Champion, 1885: 133.

Ulus latus Blaisdell, 1892USA (CA)

Ulus latus Blaisdell, 1892: 243.

Ulus lineatulus Champion, 1885MEX (JA) GUANIC

Ulus lineatulus Champion, 1885: 134.

Ulus maritimus Casey, 1890USA (AL FL MS TX)

Ulus maritimus Casey, 1890b: 414.

Ulus obliquus (LeConte, 1866)MEX (BC BS)

Blapstinus obliquus LeConte, 1866b: 117.

Genus Xerolinus Ivie and Hart, 2016 [M]

Xerolinus Ivie and Hart, 2016: 470. Type species: Diastolinus sallei Mulsant and Rey, 1859, original designation.

Xerolinus alfaroi (Garrido and Gutiérrez, 1996) CUB

Diastolinus alfaroi Garrido and Gutiérrez, 1996a: 228.

Xerolinus alutaceus (Casey, 1890)USA (FL) / CUB

Blapstinus opacus LeConte, 1878a: 420 [junior primary homonym of Blapstinus opacus Mulsant, 1859].

Blapstinus alutaceus Casey, 1890b: 423. Synonymy: Casey (1890b: 423).

Diastolinus trinitatis Marcuzzi, 1976: 127. Synonymy: Ivie and Hart (2016: 470).

Xerolinus armasi (Marcuzzi, 1988) CUB

Diastolinus armasi Marcuzzi, 1988: 72.

Xerolinus bahamae (Marcuzzi, 1965) BAH

Diastolinus bahamae Marcuzzi, 1965: 125.

Xerolinus bielawskii (Marcuzzi, 1985) CUB

Diastolinus bielawskii Marcuzzi, 1985: 182.

Xerolinus burtoni (Garrido and Gutiérrez, 1996) CAY

Diastolinus burtoni Garrido and Gutiérrez, 1996b: 232.

Xerolinus caguamensis (Marcuzzi, 1988) CUB

Diastolinus caguamensis Marcuzzi, 1988: 74.

Xerolinus camanoensis Hart and Ivie, 2016 VIS

Xerolinus camanoensis Hart and Ivie, 2016b: 885.

Xerolinus caymanensis (Marcuzzi, 1977) CAY

Diastolinus caymanensis Marcuzzi, 1977: 12.

Xerolinus cubanus (Marcuzzi, 1962) CUB

Diastolinus cubanus Marcuzzi, 1962: 30.

Xerolinus dentipes (Marcuzzi, 1977) CUB CAY

Diastolinus dentipes Marcuzzi, 1977: 13.

Diastolinus diformis Marcuzzi, 1977: 14. Synonymy: Garrido and Gutiérrez (1996b: 232).

Xerolinus difficilis (Marcuzzi, 1976) CUB

Diastolinus difficilis Marcuzzi, 1976: 126.

Xerolinus dispar (Casey, 1890)USA (FL)

Blapstinus dispar Casey, 1890b: 424.

Xerolinus dozieri (Marcuzzi, 1965) TUR

Diastolinus dozieri Marcuzzi, 1965: 128.

Xerolinus elongatus (Marcuzzi, 1976) CUB

Diastolinus elongatus Marcuzzi, 1976: 126.

Xerolinus garridoi (Marcuzzi, 1988) CUB

Diastolinus garridoi Marcuzzi, 1988: 78.

Xerolinus hernandezi (Marcuzzi, 1988)CUB (Isla de Juventud)

Diastolinus hernandezi Marcuzzi, 1988: 72.

Xerolinus juraguensis (Marcuzzi, 1988) CUB

Diastolinus juraguensis Marcuzzi, 1988: 75.

Xerolinus kulzeri (Marcuzzi, 1965)BAH (Mayaguana)

Diastolinus kulzeri Marcuzzi, 1965: 127.

Xerolinus macamboensis (Marcuzzi, 1988) CUB

Diastolinus macamboensis Marcuzzi, 1988: 77.

Diastolinus garciai Marcuzzi, 1988: 79. Synonymy: Garrido and Gutiérrez (1996a: 226).

Xerolinus minor (Marcuzzi, 1977) CAY

Diastolinus minor Marcuzzi, 1977: 18.

Xerolinus orientalis (Garrido and Gutiérrez, 1996) CUB

Diastolinus orientalis Garrido and Gutiérrez, 1996a: 226.

Xerolinus puncticeps (Mulsant and Rey, 1859) CUB

Blapstinus puncticeps Mulsant and Rey, 1859: 181.

Xerolinus rufoclavatus (Zayas, 1988) CUB

Blapstinus rufoclavatus Zayas, 1988: 91.

Xerolinus sallei (Mulsant and Rey, 1859) HAI DOM

Diastolinus sallei Mulsant and Rey, 1859: 144.

Diastolinus costipennis Mulsant and Rey, 1859: 149. Synonymy: Ivie and Hart (2016: 473).

Diastolinus puncticollis Mulsant and Rey, 1859: 147. Synonymy: Ivie and Hart (2016: 473).

Diastolinus assoi Garrido, 2004b: 40. Synonymy: Ivie and Hart (2016: 473).

Xerolinus smalli (Garrido, 2004) CUB

Diastolinus smalli Garrido, 2004c: 46.

Xerolinus swearingenae Hart and Ivie, 2016 JAM

Xerolinus swearingenae Hart and Ivie, 2016b: 889.

Xerolinus that (Steiner, 2006) BAH

Diastolinus that Steiner, 2006: 25.

Xerolinus this (Steiner, 2006) BAH

Diastolinus this Steiner, 2006: 21.

Xerolinus waterhousii (Mulsant and Rey, 1859) CUB

Diastolinus waterhousii Mulsant and Rey, 1859: 152.

Diastolinus kaszabi Marcuzzi, 1976: 125. Synonymy: Ivie and Hart (2016: 474).

Xerolinus zayasi (Marcuzzi, 1988) CUB

Diastolinus zayasi Marcuzzi, 1988: 75.

Tribe Palorini Matthews, 2003

Palorinae Matthews, 2003: 50. Type genus: Palorus Mulsant, 1854.

Genus Palorus Mulsant, 1854 [M]

Palorus Mulsant, 1854: 250. Type species: Hypophloeus depressus Fabricius, 1790, monotypy.

Caenocorse C.G. Thomson, 1859: 117. Type species: Hypophloeus depressus Fabricius, 1790, original designation. Synonymy: Bedel (1906: 92).

Eba Pascoe, 1863: 129. Type species: Eba cerylonoides Pascoe, 1863, monotypy. Synonymy: Carter and Zeck (1937: 194).

Circomus Fleischer, 1900: 236. Type species: Hypophloeus subdepressus Wollaston, 1864, monotypy. Synonymy: Löbl et al. (2008a: 276).

Palorus cerylonoides (Pascoe, 1863)USA (FL) / LAN – Adventive

Eba cerylonoides Pascoe, 1863: 129.

Palorus genalis Blair, 1930BEL / LAN – Adventive

Palorus genalis Blair, 1930: 140.

Palorus ratzeburgii (Wissmann, 1848) [Fig. 27] CAN (BC NS ON QC) USA (DC FL GA IN MI NY OH OR SC WA WI) / CUB – Adventive

Figure 27. 

Palorus ratzeburgii (Wissmann, 1848). Scale bar = 1 mm.

Hypophloeus ratzeburgii Wissmann, 1848: 77.

Palorus subdepressus (Wollaston, 1864)CAN (MB ON) USA (CT DC FL GA ID IN MI OH OR PA SC SD WI) MEX / LAN – Adventive

Hypophloeus subdepressus Wollaston, 1864: 499.

Genus Ulomina Baudi di Selve, 1876 [F]

Ulomina Baudi di Selve, 1876: 235. Type species: Ulomina carinata Baudi di Selve, 1876, monotypy.

Coelopalorus Blair, 1930: 135. Type species: Palorus foveicollis Blair, 1930 (= Ulomina carinata Baudi di Selve, 1876), monotypy. Synonymy: Scupola (2002: 186).

Ulomina carinata Baudi di Selve, 1876USA (AL FL) / CUB – Adventive

Ulomina carinata Baudi di Selve, 1876: 236.

Palorus foveicollis Blair, 1930: 136. Synonymy: Scupola (2002: 186).

Tribe Pedinini Eschscholtz, 1829

Pediniden Eschscholtz, 1829: 4. Type genus: Pedinus Latreille, 1797.

Subtribe Leichenina Mulsant, 1854

Leichenaires Mulsant, 1854: 179. Type genus: Leichenum Dejean, 1834.

Genus Leichenum Dejean, 1834 [N]

Leichenum Dejean, 1834: 194. Type species: Opatrum pictum Fabricius, 1801, monotypy.

Lichenum Agassiz, 1846: 209. Unjustified emendation of Leichenum Dejean, 1834, not in prevailing usage.

Leichenum canaliculatum variegatum (Klug, 1833)USA (AL FL GA MS NC SC) / BAHCUBLAN – Adventive

Opatrum variegatum Klug, 1833: 88.

Subtribe Platynotina Mulsant and Rey, 1853

Platynotaires Mulsant and Rey, 1853: 263. Type genus: Platynotus Fabricius, 1801.

Genus Alaetrinus Iwan, 1995 [M]

Alaetrinus Iwan, 1995: 24. Type species: Tenebrio pullus Sahlberg, 1823, original designation.

Alaetrinus aciculatus (LeConte, 1858)USA (CA KS NM OK TX) MEX (NL TA)

Opatrinus aciculatus LeConte, 1858b: 75.

Alaetrinus minimus (Palisot de Beauvois, 1817)USA (AL AR CA CT DC DE FL GA IL IN KS KY LA MA MD MO MS NC NJ NY OH OK PA SC TN TX VA WV) / BAHCUB

Tenebrio minimus Palisot de Beauvois, 1817: 164.

Pedinus suturalis Say, 1824a: 263 [nomen dubium]. New synonymy [PB]53.

Opatrum notum Say, 1826: 237. Synonymy: Mulsant and Rey (1853: 309).

Alaetrinus moestus (Mulsant and Rey, 1853)MEX (VE) BEL / SA

Opatrinus moestus Mulsant and Rey, 1853: 307.

Opatrinus lüderwaldti Gebien, 1928a: 112. Synonymy: Iwan (1995: 47).

Alaetrinus pullus (Sahlberg, 1823)USA (FL) MEX (CI QR TB VE YU) GUABELHON / BERBAHCUBJAMDOMPRILAN / SA

Tenebrio pullus C.R. Sahlberg, 1823: 16.

Opatrinus anthracinus Mulsant and Rey, 1853: 304. Synonymy: Champion (1885: 123).

Opatrinus puertoricensis Marcuzzi, 1977: 23. Synonymy: Iwan (1995: 38).

Genus Anchophthalmops Koch, 1956 [M]

Anchophthalmops Koch, 1956: 173. Type species: Anchophthalmops brevipleurum Koch, 1956, original designation.

Anchophthalmops menouxi (Mulsant and Rey, 1853)USA (KS) – Adventive

Selinus menouxii Mulsant and Rey, 1853: 322.

Opatrinus sayi Horn, 1870: 349. Synonymy: Iwan (1995: 52).

Genus Opatrinus Dejean, 1821 [M]

Opatrinus Dejean, 1821: 66. Type species: Opatrum clathratum Fabricius, 1787, monotypy.

Hopatrinus Agassiz, 1846: 185. Unjustified emendation of Opatrinus Dejean, 1821, not in prevailing usage.

Opatrinus clathratus (Fabricius, 1787)PAN / LAN / SA

Opathrum clathratum Fabricius, 1787: 379.

Blaps gemellata Olivier, 1795: [60] 9. Synonymy: Iwan (1995: 16).

Helops aethiops Fabricius, 1801a: 162. Synonymy: Gebien (1906: 212).

Opatrinus geminatus Erichson, 1849: 565. Synonymy: Gebien (1906: 212).

Opatrinus gridellii Marcuzzi, 1949: 342. Synonymy: Iwan (1995: 16).

Opatrinus armasi Garrido and Gutiérrez, 1994a: 121. Synonymy: Iwan (2002: 282).

Opatrinus gibbicollis Mulsant and Rey, 1853PAN / SA

Opatrinus gibbicollis Mulsant and Rey, 1853: 303.

Tribe Tenebrionini Latreille, 1802

Tenebrionites Latreille, 1802: 165. Type genus: Tenebrio Linnaeus, 1758.

Biuini Skopin, 1978: 224. Type genus: Bius Dejean, 1834.

Genus Bius Dejean, 1834 [M]

Bius Dejean, 1834: 205. Type species: Trogosita thoracicus Fabricius, 1792, monotypy.

Bius estriatus (LeConte, 1851)CAN (AB BC YT) USA (CA ID NM MI OR WA)

Tenebrio estriatus LeConte, 1851: 149.

Genus Bouchardandrus Steiner, 2016 [M]

Bouchardandrus Steiner, 2016: 543. Type species: Haplandrus concolor LeConte, 1866, original designation.

Bouchardandrus concolor (LeConte, 1866)CAN (MB ON QC) USA (MI MN OH WI)

Haplandrus concolor LeConte, 1866b: 121.

Genus Idiobates Casey, 1891 [M]

Idiobates Casey, 1891: 62. Type species: Tenebrio castaneus Knoch, 1801, monotypy.

Idiobates castaneus (Knoch, 1801)CAN (ON QC) USA (AL AR DE FL GA IL IN KY MD MI NC NH NJ NY OH OK PA SC TN VA WI WV)

Tenebrio castaneus Knoch, 1801: 171.

Tenebrio interstitialis Say, 1824a: 266. Synonymy: Melsheimer (1853: 139).

Genus Neatus LeConte, 1862 [M]

Neatus LeConte, 1862a: 233. Type species: Helops tenebrioides Palisot de Beauvois, 1812, monotypy.

Neatus tenebrioides (Palisot de Beauvois, 1812)CAN (AB MB NB ON QC SK) USA (AL AR AZ CA CO CT DE FL GA IA IL IN KS KY LA MD MI MN MO MS MT NC ND NE NH NJ NM OH OK PA SC SD TN TX UT VA VT WI WV)

Helops tenebrioïdes Palisot de Beauvois, 1812: 121.

Tenebrio badius Say, 1824a: 265. Synonymy: LeConte (1859d: 156).

Tenebrio rufinasus Say, 1831: 8 [nomen dubium]. Synonymy (in doubt with Tenebrio picipes Herbst sensu North American authors = Neatus tenebrioides): Papp (1961d: 130, as rufimanus).

Genus Rhinandrus LeConte, 1866 [M]

Rhinandrus LeConte, 1866b: 119. Type species: Rhinandrus gracilis LeConte, 1866, monotypy.

Exerestus Bates, 1870: 268. Type species: Exerestus jansonii Bates, 1870 (= Rhinandrus elongatus Horn, 1866), monotypy. Synonymy: Bates (1872: 98).

Proderops Fairmaire, 1873: 393. Type species: Proderops foraminosus Fairmaire, 1873 (= Rhinandrus elongatus Horn, 1866), monotypy. Synonymy (with Exerestus Bates): Kraatz (1880: 133).

Rhinandrus elongatus Horn, 1866MEX (YU) NICCRI

Rhinandrus elongatus Horn, 1866: 400.

Exerestus jansonii Bates, 1870: 269. Synonymy: LeConte (1873: 334).

Proderops foraminosus Fairmaire, 1873: 394. Synonymy: Champion (1885: 102).

Rhinandrus foveolatus (Kraatz, 1880)MEX (OA)

Proderops foveolatus Kraatz, 1880: 133.

Rhinandrus gracilis LeConte, 1866MEX (BS)

Rhinandrus gracilis LeConte, 1866b: 120.

Rhinandrus helopioides (Kraatz, 1880)MEX (OA)

Exerestus helopioides Kraatz, 1880: 135.

Rhinandrus obsoletus Champion, 1885MEX (DU SI)

Rhinandrus obsoletus Champion, 1885: 102.

Genus Tenebrio Linnaeus, 1758 [M]

Tenebrio Linnaeus, 1758: 417. Type species: Tenebrio molitor Linnaeus, 1758, subsequent designation (Latreille 1810: 429).

Menedrio Motschulsky, 1872: 27. Type species: Tenebrio obscurus Fabricius, 1792, original designation. Synonymy: Heyden et al. (1883: 134).

Tenebrionellus Crotch, 1874: 105. Unnecessary replacement name for Tenebrio Linnaeus, 1758.

Tenebrio molitor Linnaeus, 1758 [Fig. 28] CAN (AB BC MB NB NF NS ON PE QC SK) USA (AK FL GA ID IN MA MD MI NC OH OR SC SD WA WI) CRI / CUBPRI – Adventive

Figure 28. 

Tenebrio molitor Linnaeus, 1758. Scale bar = 1 mm.

Tenebrio molitor Linnaeus, 1758: 417.

Tenebrio obscurus Fabricius, 1792GRECAN (AB BC NS ON QC SK) USA (FL GA ID IN MA MD MI NC OH OR SC SD WA WI) – Adventive

Tenebrio obscurus Fabricius, 1792a: 111.

Menedrio longipennis Motschulsky, 1872: 37. New synonymy [YB].

Tenebrio obscurus pollens Casey, 1924: 321. Synonymy: Bousquet and Campbell (1991: 259).

Genus Zophobas Dejean, 1834 [M]

Zophobas Dejean, 1834 [30 June]: 204. Type species: Helops morio Fabricius, 1777 (= Tenebrio atratus Fabricius, 1775), subsequent designation (Motschulsky 1872: 26).

Subgenus Macrozophobas Pic, 1913

Macrozophobas Pic, 1913b: 6. Type species: Macrozophobas gracilicornis Pic, 1913 (=Zophobas maculicollis Kirsch, 1866), monotypy.

Zophobas klingelhoefferi Kraatz, 1880MEX (VE)

Zophobas klingelhöfferi Kraatz, 1880: 126.

Zophobas maculicollis Kirsch, 1866PAN / SA

Zophobas maculicollis Kirsch, 1866: 196.

Macrozophobas gracilicornis Pic, 1913b: 6. Synonymy: Gebien (1941: 335).

Zophobas signatus Champion, 1885 MEX GUA BEL HON NIC CRI PAN

Zophobas signatus Champion, 1885: 104.

Subgenus Zophobas Dejean, 1834

Zophobas Dejean, 1834 [30 June]: 204. Type species: Helops morio Fabricius, 1777 (= Tenebrio atratus Fabricius, 1775), subsequent designation (Motschulsky 1872: 26).

Pythonissus Gistel, 1834 [23 September]: 21. Type species: Helops morio Fabricius, 1777 (= Tenebrio atratus Fabricius, 1775), subsequent designation (Bousquet and Bouchard 2017a: 132). Synonymy: Bousquet and Bouchard (2017a: 132).

Zophobas atratus (Fabricius, 1775)USA (CA FL) MEX (GE JA OA SL TB VE YU) GUANICCRIPAN / BAHCUBJAMHAIDOMPRILAN / SA

Tenebrio atratus Fabricius, 1775: 256.

Helops morio Fabricius, 1777: 241. Synonymy: Marcuzzi and d’Aguilar (1971: 90)54.

Tenebrio elongatus Palisot de Beauvois, 1817: 164 [junior primary homonym of Tenebrio elongatus Herbst, 1797]. Synonymy (with H. morio Fabricius): Chevrolat (1853: 638).

Zophobas rugipes Kirsch, 1866: 19755. Synonymy: Tschinkel (1984: 332).

Zophobas concolor Wollaston, 1870: 33. Synonymy (with H. morio Fabricius): Champion (1896: 26).

Zophobas alternans Kraatz, 1880: 131. Synonymy: Ferrer (2011: 298).

Zophobas batavorum Marcuzzi, 1959: 88. Synonymy: Ferrer (2011: 297).

Zophobas costatus Pic, 1921 DOM

Zophobas costatus Pic, 1921a: 10.

Zophobas diversicolor Pic, 1921 HON

Zophobas rugipes var. diversicolor Pic, 1921a: 9.

Zophobas macretus Kraatz, 1880MEX (CI ME OA TA VE YU) GUANICCRI

Zophobas macretus Kraatz, 1880: 130.

Zophobas opacus (Sahlberg, 1823)USA (NM) MEX (GE VE) GUASALNICCRIPAN / PRILAN / SA

Helops opacus C.R. Sahlberg, 1823: 17.

Zophabas [sic!] subnitidus Motschulsky, 1872: 35. Synonymy: Ferrer (2011: 301).

Zophabas [sic!] laticollis Motschulsky, 1872: 36. Synonymy: Ferrer (2011: 301).

Zophobas ambiguus Kraatz, 1880: 124. Synonymy: Ferrer (2011: 301).

Zophobas kraatzi Champion, 1885: 105. Synonymy: Ferrer (2011: 301).

Zophobas diversipes Pic, 1921a: 8. Synonymy: Ferrer (2011: 301).

Zophobas cubanus Marcuzzi, 1976: 128. Synonymy: Ferrer (2011: 301).

Zophobas tridentatus Kraatz, 1880NICPAN / SA

Zophobas tridentatus Kraatz, 1880: 124.

Zophobas kirschi Kraatz, 1880: 127. Synonymy: Ferrer (2011: 299).

Zophobas pedestris Champion, 1885: 103. Synonymy: Ferrer (2011: 299).

Zophobas elongatior Pic, 1921a: 9. Synonymy: Ferrer (2011: 299).

[incertae sedis]

Zophobas subnitens (Horn, 1874)USA (AZ) MEX (SO)

Nyctobates subnitens Horn, 1874a: 35.

Rhinandrus sublaevis Horn, 1885c: 160. Synonymy: Spilman (1962b: 60).

Tribe Toxicini Oken, 1843

Toxiciden Oken, 1843: 484. Type genus: Toxicum Latreille, 1802.

Subtribe Dysantina Gebien, 1922

Dysantinae Gebien, 1922: 289. Type genus: Dysantes Pascoe, 1869.

Eudysantina Bouchard, Lawrence, Davies and Newton, 2005: 508. Type genus: Eudysantes Bouchard et al., 2005 (= Dysantes Pascoe, 1869). Note. Dysantes Pascoe was first made available in 1869 [on 1 January], not in 1871 as previously noted. The name is a senior homonym of the ichneumonid Dysantes Förster, 1869 [May], which was incorrectly dated 1868. The replacement name of the family-group name proposed by Bouchard et al. (2005) was thus unnecessary. A full explanation about this case will be issued in a forthcoming publication by YB and PB.

Genus Diceroderes Solier, 1841 [M]

Diceroderes Solier, 1841: 30, 46. Type species: Diceroderes mexicanus Solier, 1841, original designation.

Prosomenes Blanchard, 1845: 10. Type species: Diceroderes mexicanus Solier, 1841, subsequent monotypy (Chevrolat 1847: 562). Note. The generic name Prosomenes was listed as a junior synonym of Dicérodères by Blanchard (1845: 10). However, because the name was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Chevrolat 1847: 562), it is made available thereby but dates from its first publication as a synonym (ICZN 1999: Article 11.6.1).

Diceroderes cusucoensis Smith, 2015 GUA HON

Diceroderes cusucoensis Smith [in Smith and Cifuentes-Ruiz], 2015: 62.

Diceroderes mexicanus Solier, 1841MEX (HI PU VE)

Diceroderes mexicanus Solier, 1841: 49.

Diceroderes ocozocoautlaensis Smith, 2015MEX (CI)

Diceroderes ocozocoautlaensis Smith [in Smith and Cifuentes-Ruiz], 2015: 63.

Diceroderes skelleyi Smith, 2015 GUA

Diceroderes skelleyi Smith [in Smith and Cifuentes-Ruiz], 2015: 65.

Diceroderes subtriplehorni Smith and Cifuentes-Ruiz, 2015MEX (OA PU VE)

Diceroderes subtriplehorni Smith and Cifuentes-Ruiz, 2015: 67.

Genus Ozolais Pascoe, 1866 [F]

Ozolais Pascoe, 1866: 457. Type species: Ozolais scruposa Pascoe, 1866, monotypy.

Ozolais elongata Champion, 1886 NIC PAN

Ozolais elongata Champion, 1886: 228.

Ozolais lutosa Champion, 1886 CRI

Ozolais lutosa Champion, 1886: 227.

Ozolais nodosa Champion, 1886 NIC

Ozolais nodosa Champion, 1886: 228.

Ozolais tuberculifera Champion, 1896 LAN

Ozolais tuberculifera Champion, 1896: 10.

Ozolais verrucosa Champion, 1886 PAN

Ozolais verrucosa Champion, 1886: 226.

Genus Wattius Kaszab, 198256 [M]

Wattius Kaszab, 1982: 50. Type species: Calymmus cucullatus Pascoe, 1871, original designation.

Wattius andersoni Smith and Sanchez, 2015 CUB

Wattius andersoni Smith and Sanchez, 2015: 118.

Wattius emmabaconae Smith and Sanchez, 2015 DOM

Wattius emmabaconae Smith and Sanchez, 2015: 121.

Wattius variegatus (Champion, 1886) NIC

Calymmus variegatus Champion, 1886: 225.

Wattius viatorus Smith and Sanchez, 2015 BAH CUB

Wattius viatorus Smith and Sanchez, 2015: 125.

Tribe Triboliini Gistel, 1848

Triboliidae Gistel, 1848: [4]. Type genus: Tribolium MacLeay, 1825.

Genus Aesymnus Champion, 1886 [M]

Aesymnus Champion, 1886: 168. Type species: Aesymnus nitidus Champion, 1886, monotypy.

Aesymnus nitidus Champion, 1886MEX (VE) PAN

Aesymnus nitidus Champion, 1886: 168.

Genus Hypogena Dejean, 1834 [F]

Hypogena Dejean, 1834: 199. Type species: Tenebrio biimpressus Latreille, 1813, monotypy.

Ulosonia Laporte, 1840: 220. Type species: Uloma tricornis Laporte, 1840 (= Phaleria tricornis Dalman, 1823), subsequent designation (Gebien 1940: 786). Synonymy: Jacquelin du Val (1857: 148).

Hypogena biimpressa (Latreille, 1813)MEX (CI DU JA OA PU SI TA VE YU) GUABELNICPAN / CUBDOMLAN / SA

Tenebrio biimpressus Latreille, 1813: 17.

Hypogena canaliculata (Champion, 1886) NIC CRI PAN

Ulosonia canaliculata Champion, 1886: 164.

Hypogena dejeani (Champion, 1886)GUA / SA

Ulosonia dejeani Champion, 1886: 165.

Hypogena depressa (Champion, 1886)MEX (MO)

Ulosonia depressa Champion, 1886: 164.

Hypogena marginata (LeConte, 1851)USA (AZ CA) MEX (BS)

Uloma marginata LeConte, 1851: 149.

Hypogena tricornis (Dalman, 1823)USA (FL TX) MEX (BS JA OA PU VE YU) GUABELNICCRI / BAHCUBCAY / SA

Phaleria tricornis Dalman, 1823: 59.

Ulosonia tricornis Laporte, 1840: 220. Synonymy: Spilman (1973: 42)57.

Genus Latheticus C.O. Waterhouse, 1880 [M]

Latheticus C.O. Waterhouse, 1880: 147. Type species: Latheticus oryzae C.O. Waterhouse, 1880, monotypy.

Latheticus oryzae C.O. Waterhouse, 1880 [Fig. 29] CAN (AB MB NB QC SK) USA (FL GA MD MI NC OH SC TX) / CUBHIS – Adventive

Figure 29. 

Latheticus oryzae C.O. Waterhouse, 1880. Scale bar = 1 mm.

Latheticus oryzae C.O. Waterhouse, 1880: 148.

Latheticus prosopis Chittenden, 1904USA (AZ CA FL) MEX (BS)

Latheticus prosopis Chittenden, 1904: 167.

Genus Lyphia Mulsant and Rey, 1859 [F]

Lyphia Mulsant and Rey, 1859: 166. Type species: Lyphia ficicola Mulsant and Rey, 1859 (=Bius tetraphyllus Fairmaire, 1857), monotypy.

Lyphia tetraphylla (Fairmaire, 1857)USA (DC FL GA MD OH) – Adventive

Bius tetraphyllus Fairmaire, 1857: 534.

Lyphia ficicola Mulsant and Rey, 1859: 166. Synonymy: Marseul (1876: 113).

Hypophloeus rugosus Dury, 1902: 171. Synonymy (with L. ficicola Mulsant and Rey): Schwarz [in Dury] (1902: [198]).

Genus Metulosonia Bates, 1873 [F]

Metulosonia Bates, 1873d: 261. Type species: Metulosonia horni Bates, 1873, subsequent designation (Gebien 1940: 1061).

Metulosonia horni Bates, 1873 PAN

Metulosonia horni Bates, 1873d: 262.

Metulosonia reflexa (Chevrolat, 1878)MEX (VE) GUABELNIC

Peltoides reflexus Chevrolat, 1878c: 237.

Genus Mycotrogus Horn, 1870 [M]

Mycotrogus Horn, 1870: 367. Type species: Mycotrogus piceus Horn, 1870, subsequent designation (Lucas 1920: 427).

Mycotrogus angustus Horn, 1870USA (AZ CA)

Mycotrogus angustus Horn, 1870: 368.

Mycotrogus mentalis Blaisdell, 1923USA (AZ) MEX (BC BS)

Mycotrogus mentalis Blaisdell, 1923: 279.

Mycotrogus paripunctatus Spilman, 1963 CUB

Mycotrogus paripunctatus Spilman, 1963: 23.

Mycotrogus piceus Horn, 1870USA (CA)

Mycotrogus piceus Horn, 1870: 367.

Genus Spelaebiosis58Bousquet and Bouchard, new replacement name [F]

Orghidania Ardoin, 1977b: 383 [junior homonym of Orghidania Capuse, 1971]. Type species: Orghidania torrei Ardoin, 1977, monotypy.

Ardoinia Özdikmen, 2004: 202 [junior homonym of Ardoinia Kaszab, 1969]. Replacement name for Orghidania Ardoin, 1977.

Spelaebiosis Bousquet and Bouchard, new replacement name for Ardoinia Özdikmen, 2004.

Spelaebiosis torrei (Ardoin, 1977) CUB

Orghidania torrei Ardoin, 1977b: 384.

Genus Tribolium MacLeay, 1825 [N]

Tribolium MacLeay, 1825: 47. Type species: Colydium castaneum Herbst, 1797, monotypy.

Subgenus Aphanotus LeConte, 1862

Aphanotus LeConte, 1862a: 233. Type species: Eulabis brevicornis LeConte, 1859, original designation.

Tribolium brevicorne (LeConte, 1859)CAN (BC) USA (CA OR WA)

Eulabis brevicornis LeConte, 1859b: 78.

Tribolium parallelum (Casey, 1890)USA (AZ) MEX (CL)

Aphanotus parallelus Casey, 1890b: 483.

Tribolium setosum Triplehorn, 1978USA (AZ)

Tribolium setosum Triplehorn, 1978: 73.

Subgenus Tribolium MacLeay, 1825 59

Tribolium MacLeay, 1825: 47. Type species: Colydium castaneum Herbst, 1797, monotypy.

Stene Stephens, 1829: 19. Type species: Tenebrio ferrugineus Fabricius sensu auctorum (= Colydium castaneum Herbst, 1797), monotypy. Synonymy: Shuckard (1840: vii).

Margus Dejean, 1834: 200. Type species: Tenebrio ferrugineus Fabricius sensu auctorum (= Colydium castaneum Herbst, 1797), monotypy. Synonymy: Guérin-Méneville (1846: cxvii).

Tribolium audax Halstead, 1969CAN (AB BC MB ON QC SK) USA (FL MI MN OH PA SD UT VA)

Tribolium audax Halstead, 1969: 296.

Tribolium castaneum (Herbst, 1797)CAN (AB BC MB NB NS ON PE QC SK) USA (AL CA FL GA ID IL IN MA MD MI MS NC NY OH OR PA SC SD TN VA WA WI) MEX (BS CO GE NL OA) GUANICPAN / CUBCAYJAMHAIDOMPRILAN / SA – Adventive

Dermestes navalis Fabricius, 1775: 56. Note. This name was suppressed for the purposes of the Principle of Priority (ICZN 1987).

Colydium castaneum Herbst, 1797: 282. Synonymy: Schönherr (1806: 153).

Tribolium confusum Jacquelin du Val, 1862 [Fig. 30] CAN (AB BC MB NB NF NS ON PE QC SK) USA (AK CT FL GA ID IL IN MD MI NC NJ NY OH OR PA SC SD VA WA WI) MEX (CO GU NL) PAN / CUBJAMHAIDOMPRI / SA – Adventive

Figure 30. 

Tribolium (Tribolium) confusum Jacquelin du Val, 1862. Scale bar = 1 mm.

Tribolium confusum Jacquelin du Val, 1862: 181.

Tribolium destructor Uyttenboogaart, 1934GRECAN (AB BC MB NB NF NS ON PE QC SK YT) USA (CA WA WI) – Adventive

Tribolium destructor Uyttenboogaart, 1934: 21.

Tribolium madens (Charpentier, 1825)CAN (MB NB NS ON QC) USA (KY MD MI NM PA) – Adventive

Tenebrio madens Charpentier, 1825: 218.

Tribolium linsleyi Hinton, 1948MEX (CL)

Tribolium linsleyi Hinton, 1948: 32.

Tribe Ulomini Blanchard, 1845

Ulomites Blanchard, 1845: 16. Type genus: Uloma Dejean, 1821.

Alégoriides Lacordaire, 1859: 325. Type genus: Alegoria Laporte, 1840.

Genus Alegoria Laporte, 1840 [F]

Alegoria Laporte, 1840: 221. Type species: Alegoria dilatata Laporte, 1840, monotypy.

Alegoria castelnaui Fleutiaux and Sallé, 1890 60 LAN

Allegoria [sic!] castelnaui Fleutiaux and Sallé, 1890: 425.

Alegoria dilatata Laporte, 1840MEXGUAHONNICPAN / LAN / SA

Alegoria dilatata Laporte, 1840: 221.

Alegoria sallei Bates, 1873MEX (OA VE)

Alegoria sallei Bates, 1873a: 181.

Alegoria sallaei Champion, 1886: 149. Unjustified emendation of Alegoria sallei Bates, 1873, not in prevailing usage.

Genus Antimachus Gistel, 1829 [M]

Antimachus Gistel, 1829: 1055. Type species: Phaleria furcifera Dalman, 1821, monotypy.

Ceratupis Perty, 1830: 57. Type species: Ceratupis nigerrima Perty, 1830, monotypy. Synonymy: Lacordaire (1859: 330).

Antimachus ardoini Chalumeau, 1982LAN (Martinique)

Antimachus ardoini Chalumeau, 1982: 188.

Antimachus coriaceus Lacordaire, 1859NICPAN / SA

Antimachus coriacea Lacordaire, 1859: 331.

Antimachus roudeni Fleutiaux and Sallé, 1890 LAN

Antimachus roudeni Fleutiaux and Sallé, 1890: 426.

Genus Eutochia LeConte, 1862 [F]

Eutochia LeConte, 1862a: 238. Replacement name for Aniara Lacordaire, 1859.

Subgenus Eutochia LeConte, 1862

Aniara Lacordaire, 1859: 336 [junior homonym of Aniara Hope, 1838]. Type species: Uloma picea Melsheimer, 1846, monotypy.

Eutochia LeConte, 1862a: 238. Replacement name for Aniara Lacordaire, 1859.

Delopygus LeConte, 1866b: 129. Type species: Delopygus crenatus LeConte, 1866, monotypy. Synonymy: Horn (1870: 372).

Aniarus Gemminger [in Gemminger and Harold], 1870: 1964. Unjustified emendation of Aniara Lacordaire, 1859, not in prevailing usage.

Eutochia crenata (LeConte, 1866)USA (TX)

Delopygus crenatus LeConte, 1866b: 130.

Eutochia picea (Melsheimer, 1846) [Fig. 31] USA (AL AR DC FL GA IL IN KY MD MO NC NJ NY OH OK PA SC TN TX VA WV)

Figure 31. 

Eutochia (Eutochia) picea (Melsheimer, 1846). Scale bar = 1 mm.

Uloma picea Melsheimer, 1846: 64.

Genus Pheres Champion, 1886 [M]

Pheres Champion, 1886: 150. Type species: Pheres batesi Champion, 1886, monotypy.

Pheres batesi Champion, 1886 PAN

Pheres batesi Champion, 1886: 150.

Genus Uleda Laporte, 1840 [F]

Uleda Laporte, 1840: 220. Type species: Uloma diaperoides Laporte, 1840, monotypy.

Uleda tarsalis (Perroud and Mulsant, 1856)MEX / SA61

Melasia tarsalis Perroud and Mulsant, 1856: 163.

Uleda grossa Champion, 1886: 151. Synonymy: Gebien (1940: 771).

Genus Uloma Dejean, 1821 [F]

Uloma Dejean, 1821: 67. Type species: Tenebrio culinaris Linnaeus, 1758 (see ICZN 1975).

Uloma antillarum Champion, 1896 LAN

Uloma antillarum Champion, 1896: 22.

Uloma armata Champion, 1886 GUA BEL

Uloma armata Champion, 1886: 154.

Uloma carolynae Doyen, 1985MEX (CI)

Uloma carolynae Doyen, 1985a: 518.

Uloma divergens Champion, 1886 GUA

Uloma divergens Champion, 1886: 155.

Uloma extraordinaria Spilman, 1961 CUB

Uloma extraordinaria Spilman, 1961b: 113.

Uloma fossulata Champion, 1886MEX (VE) GUABEL

Uloma fossulata Champion, 1886: 153.

Uloma grenadensis Champion, 1896 LAN

Uloma grenadensis Champion, 1896: 23.

Uloma imberbis LeConte, 1866USA (AL AR DE FL GA IA IL IN KS KY LA MD MI MO NC NJ NY OH OK SC TN TX VA)

Uloma imberbis LeConte, 1866b: 123.

Uloma impressa Melsheimer, 1846CAN (ON) USA (AL FL GA IA IL IN LA MA MD MI NC NE NH NJ NY OH PA RI SC VA WI)

Uloma impressa Melsheimer, 1846: 64.

Uloma laevicollis Champion, 1886 GUA NIC CRI PAN

Uloma laevicollis Champion, 1886: 153.

Uloma longula LeConte, 1861CAN (BC) USA (CA OR WA)

Uloma longula LeConte, 1861b: 353.

Uloma mentalis Horn, 1870CAN (ON) USA (AL AR FL GA IN KS MD MI MS NC NY OH SC TN TX WI)

Uloma mentalis Horn, 1870: 371.

Uloma mexicana (Lacordaire, 1859)MEX (VE) GUABELSALNICCRI

Antimachus mexicana Lacordaire, 1859: 331.

Uloma moensis Marcuzzi, 2000 CUB

Uloma moensis Marcuzzi, 2000: 286.

Uloma parvula Champion, 1896 LAN

Uloma parvula Champion, 1896: 23.

Uloma punctulata LeConte, 1866USA (FL GA IN LA MD MI MS NC NY OH SC TN TX VA WI)

Uloma punctulata LeConte, 1866b: 124.62

Uloma cava LeConte, 1866b: 124. Synonymy: Horn (1870: 372).

Uloma retusa ephippigera (Guérin-Méneville, 1831)MEX (VE) BELNICCRIPAN / LAN / SA

Phaleria ephippiger Guérin-Méneville, 1831b: pl. 2.

Uloma bicolor Kirsch, 1874: 403. Synonymy: Gebien (1928a: 160).

Uloma retusa var. dimidiata Champion, 1886: 154. Synonymy (with U. bicolor Kirsch): Gebien (1911a: 403).

Uloma retusa retusa (Fabricius, 1801)MEX (JA QR SI VE YU) / PRILAN / SA

Tenebrio retusus Fabricius, 1801a: 149.

Uloma rubens Laporte, 1840 “Amérique du Nord”

Uloma rubens Laporte, 1840: 22063.

Uloma spinipes Champion, 1886 GUA

Uloma spinipes Champion, 1886: 155.

Uloma sulcata Champion, 1896 LAN

Uloma sulcata Champion, 1896: 21.

Subfamily ALLECULINAE Laporte, 1840

Alléculites Laporte, 1840: 242. Type genus: Allecula Fabricius, 1801.

Tribe Alleculini Laporte, 1840

Alléculites Laporte, 1840: 242. Type genus: Allecula Fabricius, 1801.

Subtribe Alleculina Laporte, 1840

Alléculites Laporte, 1840: 242. Type genus: Allecula Fabricius, 1801.

Upinellae LeConte, 1866b: 137. Type genus: Upinella Mulsant, 1856.

Genus Aeanes Champion, 1893 [M]

Aeanes Champion, 1893a: 566. Type species: Aeanes angusticollis Champion, 1893, monotypy.

Aeanes angusticollis Champion, 1893MEX (GE)

Aeanes angusticollis Champion, 1893a: 567.

Genus Alethia Champion, 1888 [F]

Alethia Champion, 1888: 417. Type species: Alethia sallaei Champion, 1888, original designation.

Alethia azteca Champion, 1888MEX (GU)

Alethia azteca Champion, 1888: 418.

Alethia carbonaria Schaeffer, 1905USA (AZ)

Alethia carbonaria Schaeffer, 1905b: 176.

Hymenorus liebecki Fall, 1931b: 245. Synonymy: Marshall (1970b: 1).

Alethia funerea Champion, 1888MEX (GE)

Alethia funerea Champion, 1888: 419.

Alethia hoegei Champion, 1888MEX (CH)

Alethia högei Champion, 1888: 420.

Alethia lepturoides Champion, 1888 MEX

Alethia lepturoides Champion, 1888: 419.

Alethia longipennis Champion, 1888MEX (AG)

Alethia longipennis Champion, 1888: 418.

Alethia nitidipennis Champion, 1893MEX (GE)

Alethia nitidipennis Champion, 1893a: 565.

Alethia quadricollis (Fall, 1931)USA (TX)

Hymenorus quadricollis Fall, 1931b: 246.

Alethia sallaei Champion, 1888MEX (GU)

Alethia sallaei Champion, 1888: 417.

Alethia subnitida Champion, 1888MEX (GE JA)

Alethia subnitida Champion, 1888: 418.

Genus Allecula Fabricius, 1801 [F]

Allecula Fabricius, 1801b: 21. Type species: Allecula morio Fabricius, 1801, subsequent designation (Duponchel 1840: 283).

Allecula angustata Champion, 1888MEX (HI MO)

Allecula angustata Champion, 1888: 416.

Allecula belti Champion, 1888 NIC

Allecula belti Champion, 1888: 414.

Allecula brachyptera Doyen, 1990MEX (JA)

Allecula brachyptera Doyen, 1990: 241.

Allecula caribea Campbell, 1971 PRI

Allecula caribea Campbell, 1971: 67.

Allecula castaneipennis Champion, 1888CRIPAN / SA

Allecula castaneipennis Champion, 1888: 412.

Allecula depressa Champion, 1888MEX (OA)

Allecula depressa Champion, 1888: 415.

Allecula ferox Champion, 1888 GUA

Allecula ferox Champion, 1888: 413.

Allecula gaumeri Champion, 1888MEX (YU)

Allecula gaumeri Champion, 1888: 414.

Allecula inconspicua Borchmann, 1937MEX (VE)

Allecula inconspicua Borchmann, 1937: 212.

Allecula laticeps Champion, 1888MEX (OA)

Allecula laticeps Champion, 1888: 416.

Allecula opacipennis Champion, 1888MEX (OA)

Allecula opacipennis Champion, 1888: 415.

Allecula pilipes Champion, 1888MEX (VE)

Allecula pilipes Champion, 1888: 414.

Allecula ramosi Campbell, 1971 DOM PRI

Allecula ramosi Campbell, 1971: 66.

Allecula rugicollis Champion, 1888MEX (GE JA)

Allecula rugicollis Champion, 1888: 412.

Allecula veraepacis Champion, 1888 GUA

Allecula veraepacis Champion, 1888: 413.

Genus Amaropsis Champion, 1893 [F]

Amaropsis Champion, 1893a: 567. Type species: Amaropsis annulicornis Champion, 1893, monotypy.

Amaropsis annulicornis Champion, 1893MEX (VE)

Amaropsis annulicornis Champion, 1893a: 568.

Genus Charisius Champion, 1888 [M]

Charisius Champion, 1888: 421. Type species: Charisius fasciatus Champion, 1888, subsequent designation (Lucas 1920: 178).

Narses Champion, 1888: 423. Type species: Narses subalatus Champion, 1888, monotypy. Synonymy: Campbell (2014a: 271).

Charisius apterus Campbell, 2014MEX (OA)

Charisius apterus Campbell, 2014a: 278.

Charisius fasciatus Champion, 1888MEX (CI) GUASALHON

Charisius fasciatus Champion, 1888: 421.

Charisius granulatus Campbell, 2014 GUA

Charisius granulatus Campbell, 2014a: 277.

Charisius howdenorum Campbell, 2014MEX (CI)

Charisius howdenorum Campbell, 2014a: 287.

Charisius mexicanus Campbell, 1965MEX (GE ME MI MO OA PU)

Charisius mexicanus Campbell, 1965: 49.

Charisius picturatus Champion, 1893MEX (GE ME OA)

Charisius picturatus Champion, 1893a: 565.

Charisius punctatus Campbell, 2014 GUA

Charisius punctatus Campbell, 2014a: 290.

Charisius salvini Champion, 1888 GUA SAL HON NIC

Charisius salvini Champion, 1888: 423.

Charisius subalatus (Champion, 1888) GUA SAL

Narses subalatus Champion, 1888: 424.

Charisius zunilensis Champion, 1888MEX (CI VE) GUAHON

Charisius zunilensis Champion, 1888: 422.

Charisius interstitialis Champion, 1888: 422. Synonymy: Campbell (2014a: 285).

Charisius floridanus Linell, 1899: 184. Synonymy (with C. interstitialis Champion): Campbell (1965: 51).

Genus Diopoenus Champion, 1888 [M]

Diopoenus Champion, 1888: 445. Type species: Diopoenus compressicornis Champion, 1888, monotypy.

Diopoenus compressicornis Champion, 1888MEX (PU)

Diopoenus compressicornis Champion, 1888: 445.

Genus Hymenorus Mulsant, 1852 [M]

Hymenorus Mulsant, 1852: 68 [as Hymenophorus]. Type species: Hymenorus doublieri Mulsant, 1852, monotypy. Note. See Bousquet et al. (2015: 133) for precedence of the spelling Hymenorus over Hymenophorus.

Hymenorus alienus Fall, 1931USA (AZ)

Hymenorus alienus Fall, 1931b: 217.

Hymenorus americanus Champion, 1888MEX (CL GE VE) GUANIC

Hymenorus americanus Champion, 1888: 438.

Hymenorus anguillae Campbell, 1971LAN (Anguilla)

Hymenorus anguillae Campbell, 1971: 76.

Hymenorus angustatus Champion, 1888MEX (FD) GUA

Hymenorus angustatus Champion, 1888: 436.

Hymenorus antillensis Campbell, 1971 LAN

Hymenorus antillensis Campbell, 1971: 77.

Hymenorus apacheanus Casey, 1891USA (AZ CA)

Hymenorus apacheanus Casey, 1891: 99.

Hymenorus arkansanus Fall, 1931USA (FL AR)

Hymenorus arkansanus Fall, 1931b: 183.

Hymenorus atratus Fall, 1931USA (AZ)

Hymenorus atratus Fall, 1931b: 189.

Hymenorus badius Champion, 1888MEX (VE)

Hymenorus badius Champion, 1888: 433.

Hymenorus bahamensis Campbell, 1971 BAH CUB

Hymenorus bahamensis Campbell, 1971: 88.

Hymenorus balli Campbell, 2014MEX (CI) GUA

Hymenorus balli Campbell, 2014b: 299.

Hymenorus bifurcatus Campbell, 2014 GUA

Hymenorus bifurcatus Campbell, 2014b: 301.

Hymenorus bitumescens Fall, 1931USA (AZ)

Hymenorus bitumescens Fall, 1931b: 194.

Hymenorus brevicornis Champion, 1888MEX (FD VE)

Hymenorus brevicornis Champion, 1888: 426.

Hymenorus brevipes Champion, 1888MEX (GE)

Hymenorus brevipes Champion, 1888: 435.

Hymenorus brevis Fall, 1931USA (AZ)

Hymenorus brevis Fall, 1931b: 230.

Hymenorus caducus Fall, 1931USA (AL FL)

Hymenorus caducus Fall, 1931b: 213.

Hymenorus canaliculatus Champion, 1888MEX (VE)

Hymenorus canaliculatus Champion, 1888: 428.

Hymenorus capensis Fall, 1931MEX (BS)

Hymenorus capensis Fall, 1931b: 205.

Hymenorus castaneus Champion, 1888MEX (DU)

Hymenorus castaneus Champion, 1888: 434.

Hymenorus cassus Fall, 1931MEX (BC)

Hymenorus cassus Fall, 1931b: 197.

Hymenorus caurinus Fall, 1931CAN (BC) USA (OR)

Hymenorus caurinus Fall, 1931b: 185.

Hymenorus chiriquensis Campbell, 1962 PAN

Hymenorus chiriquensis Campbell, 1962: 95.

Hymenorus colonoides Champion, 1888MEX (GU JA PU VE) GUA

Hymenorus colonoides Champion, 1888: 435.

Hymenorus communis LeConte, 1866USA (FL GA MD NC NY PA SC WI)

Hymenorus communis LeConte, 1866b: 135.

Hymenorus confertus LeConte, 1866MEX (BS)

Hymenorus confertus LeConte, 1866b: 136.

Hymenorus conformis Fall, 1931USA (TX)

Hymenorus conformis Fall, 1931b: 199.

Hymenorus conicicollis Fall, 1931USA (GA SC)

Hymenorus conicicollis Fall, 1931b: 239.

Hymenorus convexus Casey, 1891USA (FL TX) / BAHTURCUBCAY

Hymenorus convexus Casey, 1891: 106.

Hymenorus corticarioides Champion, 1888MEX (CL GE)

Hymenorus corticarioides Champion, 1888: 441.

Hymenorus crinitus Fall, 1931USA (AZ)

Hymenorus crinitus Fall, 1931b: 244.

Hymenorus curticollis Casey, 1891USA (AR IA IN MS PA)

Hymenorus curticollis Casey, 1891: 95.

Hymenorus cubensis Campbell, 1971 CUB

Hymenorus cubensis Campbell, 1971: 81.

Hymenorus darlingtoni Campbell, 1971 CUB

Hymenorus darlingtoni Campbell, 1971: 83.

Hymenorus densus LeConte, 1866USA (AL FL GA IN NC SC TX) MEX (VE) / BAH

Hymenorus densus LeConte, 1866b: 138.

Hymenorus deplanatus Champion, 1888USA (AZ) MEX (SO)

Hymenorus deplanatus Champion, 1888: 440.

Hymenorus gemellus Casey, 1891: 121. Synonymy: Fall (1931b: 231).

Hymenorus depressus Champion, 1888MEX (GE)

Hymenorus depressus Champion, 1888: 435.

Hymenorus dichrous Blatchley, 1919USA (FL GA NC SC)

Hymenorus dichrous Blatchley, 1919: 66.

Hymenorus difficilis Casey, 1891USA (NY)

Hymenorus difficilis Casey, 1891: 94.

Hymenorus digressus Fall, 1931USA (AZ)

Hymenorus digressus Fall, 1931b: 206.

Hymenorus discrepans Casey, 1891USA (CA)

Hymenorus discrepans Casey, 1891: 98.

Hymenorus discretus Casey, 1891CAN (ON QC) USA (FL GA IN MA MD MN MO NC NE NJ NY PA RI SC VA WI)

Hymenorus discretus Casey, 1891: 105.

Hymenorus disparatus Fall, 1931USA (AZ CO NM TX)

Hymenorus disparatus Fall, 1931b: 215.

Hymenorus dissensus Casey, 1891USA (TX)

Hymenorus dissensus Casey, 1891: 109.

Hymenorus distinctus Fall, 1931USA (AL FL GA MS SC)

Hymenorus distinctus Fall, 1931b: 179.

Hymenorus dorsalis Schwarz, 1878USA (AL FL GA NC SC)

Hymenorus dorsalis Schwarz, 1878: 370.

Hymenorus sabalensis Blatchley, 1919: 67. Synonymy: Fall (1931b: 212).

Hymenorus dubius Fall, 1931USA (AL FL GA MS SC)

Hymenorus dubius Fall, 1931b: 184.

Hymenorus durangoensis Champion, 1888MEX (DU)

Hymenorus durangoensis Champion, 1888: 426.

Hymenorus emmenastoides Champion, 1888MEX (VE) GUA

Hymenorus emmenastoides Champion, 1888: 436.

Hymenorus excavatus Campbell, 2014 GUA

Hymenorus excavatus Campbell, 2014b: 305.

Hymenorus exiguus Casey, 1891USA (AZ CA TX)

Hymenorus exiguus Casey, 1891: 100.

Hymenorus exilis Fall, 1931USA (AZ)

Hymenorus exilis Fall, 1931b: 233.

Hymenorus facetus Fall, 1931MEX (BS)

Hymenorus facetus Fall, 1931b: 234.

Hymenorus farri Campbell, 1971USA (FL) MEX (VE) GUABEL / BAHTURCUBCAYJAMPRILAN

Hymenorus farri Campbell, 1971: 84.

Hymenorus flohri Champion, 1888MEX (FD MO)

Hymenorus flohri Champion, 1888: 429.

Hymenorus floridanus Casey, 1891USA (FL)

Hymenorus floridanus Casey, 1891: 116.

Hymenorus forreri Champion, 1888MEX (DU)

Hymenorus forreri Champion, 1888: 431.

Hymenorus foveiventris Champion, 1888 GUA

Hymenorus foveiventris Champion, 1888: 432.

Hymenorus fuscipennis Fall, 1931USA (FL)

Hymenorus fuscipennis Fall, 1931b: 211.

Hymenorus fusculus Casey, 1891USA (CA)

Hymenorus fusculus Casey, 1891: 117.

Hymenorus fusicornis Casey, 1891USA (CA)

Hymenorus fusicornis Casey, 1891: 112.

Hymenorus grandicollis Champion, 1888USA (AZ) MEX (SO)

Hymenorus grandicollis Champion, 1888: 429.

Hymenorus granulatus Blatchley, 1912USA (FL)

Hymenorus granulatus Blatchley, 1912: 331.

Hymenorus guatemalensis Champion, 1888 GUA

Hymenorus guatemalensis Champion, 1888: 439.

Hymenorus haitellus Campbell, 1971 HAI

Hymenorus haitellus Campbell, 1971: 94.

Hymenorus haitius Campbell, 1971 HAI DOM

Hymenorus haitius Campbell, 1971: 93.

Hymenorus helvinus Casey, 1891USA (TX)

Hymenorus helvinus Casey, 1891: 101.

Hymenorus heteropygus Fall, 1931USA (FL GA MS)

Hymenorus heteropygus Fall, 1931b: 241.

Hymenorus hispaniolensis Campbell, 1971 HAI DOM

Hymenorus hispaniolensis Campbell, 1971: 78.

Hymenorus hispidulus Champion, 1888MEX (VE)

Hymenorus hispidulus Champion, 1888: 431.

Hymenorus horrescens Fall, 1931USA (NM TX)

Hymenorus horrescens Fall, 1931b: 235.

Hymenorus humeralis LeConte, 1866USA (AL FL KY MD OH PA SC TN)

Hymenorus humeralis LeConte, 1866b: 135.

Hymenorus idoneus Fall, 1931USA (AZ)

Hymenorus idoneus Fall, 1931b: 218.

Hymenorus igualensis Champion, 1888MEX (GE JA)

Hymenorus igualensis Champion, 1888: 434.

Hymenorus illusus Fall, 1931USA (AL FL GA MD SC)

Hymenorus illusus Fall, 1931b: 192.

Hymenorus inaequalis Casey, 1891USA (AZ)

Hymenorus inaequalis Casey, 1891: 114.

Hymenorus incertus Fall, 1931USA (AZ)

Hymenorus incertus Fall, 1931b: 220.

Hymenorus indutus Casey, 1891USA (AZ NM TX)

Hymenorus indutus Casey, 1891: 119.

Hymenorus infuscatus Casey, 1891USA (CA)

Hymenorus infuscatus Casey, 1891: 90.

Hymenorus inopiatus Fall, 1931USA (FL GA MD SC)

Hymenorus inopiatus Fall, 1931b: 242.

Hymenorus inquilinus Casey, 1891USA (CA)

Hymenorus inquilinus Casey, 1891: 112.

Hymenorus insularis Campbell, 1971 BAH

Hymenorus insularis Campbell, 1971: 91.

Hymenorus intermedius Casey, 1891USA (AZ TX)

Hymenorus intermedius Casey, 1891: 102.

Hymenorus inutilis Fall, 1931USA (AZ NM NV)

Hymenorus inutilis Fall, 1931b: 208.

Hymenorus irritus Fall, 1931USA (AZ CA)

Hymenorus irritus Fall, 1931b: 199.

Hymenorus jacobinus Fall, 1931USA (CA)

Hymenorus jacobinus Fall, 1931b: 206.

Hymenorus jamaicensis Campbell, 1971 CAY JAM

Hymenorus jamaicensis Campbell, 1971: 79.

Hymenorus laticollis Champion, 1888MEX (FD GE JA)

Hymenorus laticollis Champion, 1888: 429.

Hymenorus longicollis Champion, 1888MEX (VE)

Hymenorus longicollis Champion, 1888: 434.

Hymenorus macilentus Fall, 1931USA (NM)

Hymenorus macilentus Fall, 1931b: 188.

Hymenorus maritimus Champion, 1888 GUA

Hymenorus maritimus Champion, 1888: 437.

Hymenorus melsheimeri Casey, 1891USA (MI NY SC)

Hymenorus melsheimeri Casey, 1891: 92.

Hymenorus milleporus Fall, 1931USA (AZ)

Hymenorus milleporus Fall, 1931b: 236.

Hymenorus minutus Campbell, 1971 BAH

Hymenorus minutus Campbell, 1971: 98.

Hymenorus molestus Fall, 1931CAN (NB NS ON PE QC) USA (IN LA PA WI)

Hymenorus molestus Fall, 1931b: 182.

Hymenorus montivagus Fall, 1931USA (CA)

Hymenorus montivagus Fall, 1931b: 207.

Hymenorus nevadensis Fall, 1931USA (NV)

Hymenorus nevadensis Fall, 1931b: 236.

Hymenorus niger (Melsheimer, 1846)CAN (MB NB NS ON PE QC) USA (FL GA IN MA MD MI MN MS NC NY PA SC TX WI)

Mycetocharus niger Melsheimer, 1846: 59.

Hymenorus nitidipennis Casey, 1891USA (AZ)

Hymenorus nitidipennis Casey, 1891: 113.

Hymenorus obesus Casey, 1891CAN (MB NB NS ON QC) USA (AL FL GA IN LA MA MD MI MO NC NJ NY PA SC TX VA WI)

Hymenorus obesus Casey, 1891: 93.

Hymenorus oblivius Fall, 1931USA (TX)

Hymenorus oblivius Fall, 1931b: 216.

Hymenorus obscurus (Say, 1826)USA (FL GA IN MA MD NJ NY PA SC TX VA WI)

Cistela obscura Say, 1826: 242.

Hymenorus occidentalis Champion, 1888USA (TX) MEX (GU VE)

Hymenorus occidentalis Champion, 1888: 425.

Hymenorus oculatus Champion, 1888MEX (VE) GUA

Hymenorus oculatus Champion, 1888: 427.

Hymenorus pallidus Champion, 1888MEX (DU GE)

Hymenorus pallidus Champion, 1888: 439.

Hymenorus panamensis Campbell, 1962 PAN

Hymenorus panamensis Campbell, 1962: 93.

Hymenorus papagonis Fall, 1931USA (AZ)

Hymenorus papagonis Fall, 1931b: 201.

Hymenorus parvicollis Champion, 1888MEX (DU)

Hymenorus parvicollis Champion, 1888: 440.

Hymenorus parvus Fall, 1931USA (CA) MEX (BS)

Hymenorus parvus Fall, 1931b: 203.

Hymenorus perforatus Casey, 1891USA (GA IA IN MD NC PA SC)

Hymenorus perforatus Casey, 1891: 95.

Hymenorus picipennis Casey, 1891CAN (NS ON QC) USA (AL MD MI NH NY PA SC WI)

Hymenorus picipennis Casey, 1891: 90.

Hymenorus pilosus (Melsheimer, 1846)CAN (NS ON QC) USA (AL AR FL GA IA IN KS LA MA MD MI MS NC NJ NY OH PA SC SD VA WI)

Allecula pilosa Melsheimer, 1846: 58.

Hymenorus pini Champion, 1888 GUA

Hymenorus pini Champion, 1888: 428.

Hymenorus planulus Horn, 1894MEX (BS)

Hymenorus planulus Horn, 1894b: 434.

Hymenorus porosicornis Casey, 1891USA (NM TX)

Hymenorus porosicornis Casey, 1891: 101.

Hymenorus prolixus Casey, 1891USA (AZ NM NV TX UT)

Hymenorus prolixus Casey, 1891: 103.

Hymenorus protibialis Fall, 1931USA (AZ CA)

Hymenorus protibialis Fall, 1931b: 196.

Hymenorus punctatissimus LeConte, 1866USA (AZ CA NV TX UT) MEX (SO)

Hymenorus punctatissimus LeConte, 1866b: 138.

Hymenorus macer Casey, 1891: 118. Synonymy: Fall (1931b: 221).

Hymenorus punctulatus (LeConte, 1859)USA (CA)

Allecula punctulata LeConte, 1859b: 78.

Hymenorus pygmaeus Campbell, 1971 BAH

Hymenorus pygmaeus Campbell, 1971: 99.

Hymenorus quietus Fall, 1931USA (FL MO)

Hymenorus quietus Fall, 1931b: 239.

Hymenorus rotundicollis Casey, 1891USA (AZ)

Hymenorus rotundicollis Casey, 1891: 111.

Hymenorus rufescens Champion, 1888MEX (VE YU)

Hymenorus rufescens Champion, 1888: 433.

Hymenorus ruficollis Champion, 1888USA (AZ) MEX (BC BS SO)

Hymenorus ruficollis Champion, 1888: 438.

Hymenorus rufohumeralis Campbell, 1982USA (CA)

Hymenorus rufohumeralis Campbell, 1982: 131.

Hymenorus rufovalis Fall, 1931USA (AZ)

Hymenorus rufovalis Fall, 1931b: 230.

Hymenorus segnis Champion, 1888MEX (GE)

Hymenorus segnis Champion, 1888: 430.

Hymenorus semirufus Fall, 1931USA (FL)

Hymenorus semirufus Fall, 1931b: 203.

Hymenorus seriatus Casey, 1891USA (AZ)

Hymenorus seriatus Casey, 1891: 109.

Hymenorus setosus Hatch, 1965USA (OR)

Hymenophorus setosus Hatch, 1965: 185.

Hymenorus significans Fall, 1931USA (TX)

Hymenorus significans Fall, 1931b: 237.

Hymenorus similis Champion, 1888MEX (DU MO)

Hymenorus similis Champion, 1888: 432.

Hymenorus simiolus Fall, 1931USA (TX)

Hymenorus simiolus Fall, 1931b: 232.

Hymenorus sinuatus ebeninus Fall, 1931USA (CA)

Hymenorus sinuatus var. ebeninus Fall, 1931b: 188.

Hymenorus sinuatus sinuatus Fall, 1931CAN (BC) USA (CA ID OR WA)

Hymenorus sinuatus Fall, 1931b: 187.

Hymenophorus megops Hatch, 1965: 185. New synonymy [YB].

Telesicles magnus Hatch, 1965: 185. New synonymy [YB].

Hymenorus sobrinus Casey, 1891USA (FL MD NJ SC WI)

Hymenorus sobrinus Casey, 1891: 115.

Hymenorus sordidus Champion, 1888MEX (VE) GUA

Hymenorus sordidus Champion, 1888: 427.

Hymenorus sparsepunctatus Campbell, 1971 CUB

Hymenorus sparsepunctatus Campbell, 1971: 97.

Hymenorus spinifer Horn, 1894USA (AZ)

Hymenorus spinifer Horn, 1894b: 434.

Hymenorus striatus (Pic, 1930) HAI DOM

Cistelopsis striata Pic, 1930: 26.

Hymenorus tarsalis Champion, 1888 GUA

Hymenorus tarsalis Champion, 1888: 426.

Hymenorus tenellus Casey, 1891USA (FL GA MD NJ SC)

Hymenorus tenellus Casey, 1891: 115.

Hymenorus elbertae Blatchley, 1918: 57. Synonymy: Fall (1931b: 227).

Hymenorus tenuistriatus Fall, 1931USA (AL FL NC SC)

Hymenorus tenuistriatus Fall, 1931b: 226.

Hymenorus testaceus Casey, 1891USA (AZ)

Hymenorus testaceus Casey, 1891: 110.

Hymenorus texensis Fall, 1931USA (TX)

Hymenorus texensis Fall, 1931b: 241.

Hymenorus thoracicus Fall, 1931USA (CA)

Hymenorus thoracicus Fall, 1931b: 214.

Hymenorus tibialis Champion, 1888 GUA

Hymenorus tibialis Champion, 1888: 430.

Hymenorus torridus Champion, 1888MEX (GE)

Hymenorus torridus Champion, 1888: 436.

Hymenorus transversus Campbell, 1971 BAH

Hymenorus transversus Campbell, 1971: 92.

Hymenorus tritus Fall, 1931USA (AZ)

Hymenorus tritus Fall, 1931b: 219.

Hymenorus trivialis Fall, 1931MEX (BC)

Hymenorus trivialis Fall, 1931b: 210.

Hymenorus ulomoides Fall, 1931USA (CA)

Hymenorus ulomoides Fall, 1931b: 187.

Hymenorus uniseriatus Casey, 1891USA (CA)

Hymenorus uniseriatus Casey, 1891: 115.

Hymenorus vigilax Fall, 1931USA (AZ)

Hymenorus vigilax Fall, 1931b: 200.

Hymenorus villosus Champion, 1888MEX (JA MO)

Hymenorus villosus Champion, 1888: 440.

Hymenorus wolcotti Campbell, 1971 PRI VIS

Hymenorus wolcotti Campbell, 1971: 74.

Genus Knausia Fall, 1931 [F]

Knausia Fall, 1931a: 15. Type species: Knausia crassicornis Fall, 1931, monotypy.

Knausia crassicornis Fall, 1931USA (NM TX)

Knausia crassicornis Fall, 1931a: 16.

Genus Latacula Campbell, 1971 [F]

Latacula Campbell, 1971: 103. Type species: Latacula beckeri Campbell, 1971, original designation.

Latacula beckeri Campbell, 1971 JAM

Latacula beckeri Campbell, 1971: 105.

Latacula insularis Campbell, 1971 JAM

Latacula insularis Campbell, 1971: 106.

Genus Lobopoda Solier, 1835 [F]

Lobopoda Solier, 1835a: 233. Type species: Lobopoda striata Solier, 1835, subsequent designation (Bousquet et al. 2015: 134).

Subgenus Flavipoda Campbell, 1966

Flavipoda Campbell, 1966: 21. Type species: Helops flavipes Fabricius, 1792 [as Allecula flavipes Jacquelin duVal, 1857], original designation.

Lobopoda androsi Campbell, 1971 BAH

Lobopoda androsi Campbell, 1971: 33.

Lobopoda badia Campbell, 1971 CUB

Lobopoda badius Campbell, 1971: 31.

Lobopoda bahamensis Campbell, 1966 BAH CUB

Lobopoda bahamensis Campbell, 1966: 27.

Lobopoda bicolor Campbell, 1966 CUB

Lobopoda bicolor Campbell, 1966: 28.

Lobopoda cayamasensis Campbell, 1966 CUB

Lobopoda cayamasensis Campbell, 1966: 33.

Lobopoda deyrupi Steiner, 2006 BAH

Lobopoda deyrupi Steiner, 2006: 32.

Lobopoda emarginata Campbell, 1966 CUB

Lobopoda emarginata Campbell, 1966: 32.

Lobopoda flavifemoralis Campbell, 1966 CUB

Lobopoda flavifemoralis Campbell, 1966: 29.

Lobopoda flavipes (Fabricius, 1792) 64 CUB

Helops flavipes Fabricius, 1792a: 122 [secondary homonym of Cistela flavipes Fabricius, 1792b: 45].

Cistela fuscula Schönherr, 1808: 336. Replacement name for Cistela flavipes (Fabricius, 1792a).

Lobopoda nesiotica Campbell, 1971 BAH

Lobopoda nesiotica Campbell, 1971: 36.

Lobopoda quadratinota Campbell, 1971 CUB

Lobopoda quadratinota Campbell, 1971: 28.

Lobopoda schwarzi Campbell, 1971 CUB

Lobopoda schwarzi Campbell, 1971: 29.

Lobopoda tibiodentata Campbell, 1966 CUB

Lobopoda tibiodentata Campbell, 1966: 30.

Lobopoda villasensis Campbell, 1971 CUB

Lobopoda villasensis Campbell, 1971: 35.

Subgenus Glabrilobopoda Campbell, 1966

Glabrilobopoda Campbell, 1966: 46. Type species: Lobopoda glabrata Champion, 1888, original designation.

Lobopoda aeneipennis Champion, 1888 PAN

Lobopoda aeneipennis Champion, 1888: 408.

Lobopoda cariniventris Champion, 1888 PAN

Lobopoda cariniventris Champion, 1888: 408.

Lobopoda coronadensis Campbell, 1966 CRI

Lobopoda coronadensis Campbell, 1966: 54.

Lobopoda darlingtoni Campbell, 1971 DOM

Lobopoda darlingtoni Campbell, 1971: 40.

Lobopoda glabrata Champion, 1888 PAN

Lobopoda glabrata Champion, 1888: 409.

Lobopoda impunctata Campbell, 1966 CRI

Lobopoda impunctata Campbell, 1966: 52.

Lobopoda irazuensis Champion, 1888 CRI

Lobopoda irazuensis Champion, 1888: 406.

Lobopoda nitens Champion, 1888 CRI

Lobopoda nitens Champion, 1888: 406.

Lobopoda nitida Champion, 1888 PAN

Lobopoda nitida Champion, 1888: 407.

Lobopoda obsoleta Champion, 1888MEX (VE) GUA

Lobopoda obsoleta Champion, 1888: 409.

Lobopoda portobellensis Campbell, 1966 PAN

Lobopoda portobellensis Campbell, 1966: 58.

Lobopoda tilaranensis Campbell, 1966 CRI

Lobopoda tilaranensis Campbell, 1966: 50.

Lobopoda viridipennis Champion, 1888 PAN

Lobopoda viridipennis Champion, 1888: 407.

Subgenus Lobopoda Solier, 1835

Lobopoda Solier, 1835a: 233. Type species: Lobopoda striata Solier, 1835, subsequent designation (Bousquet et al. 2015: 134).

Lobopoda acuticauda Campbell, 1966 NIC CRI PAN

Lobopoda acuticauda Campbell, 1966: 77.

Lobopoda aeneotincta Champion, 1888 CRI PAN

Lobopoda aeneotincta Champion, 1888: 405.

Lobopoda alutacea Campbell, 1971 CUB

Lobopoda alutacea Campbell, 1971: 58.

Lobopoda apicalis Champion, 1888 GUA

Lobopoda apicalis Champion, 1888: 393.

Lobopoda atrata Champion, 1888 NIC PAN

Lobopoda atrata Champion, 1888: 394.

Lobopoda attenuata Champion, 1888 GUA NIC CRI

Lobopoda attenuata Champion, 1888: 397.

Lobopoda calcarata Champion, 1893MEX (OA)

Lobopoda calcarata Champion, 1893a: 563.

Lobopoda championi Campbell, 1966 CRI PAN

Lobopoda championi Campbell, 1966: 105.

Lobopoda chontalensis Champion, 1888 NIC CRI

Lobopoda chontalensis Champion, 1888: 399.

Lobopoda colona Campbell, 1971 HAI

Lobopoda colona Campbell, 1971: 56.

Lobopoda convexicollis Champion, 1888MEX (VE YU) GUA

Lobopoda convexicollis Champion, 1888: 395.

Lobopoda cordata Campbell, 1971 BAH

Lobopoda cordata Campbell, 1971: 47.

Lobopoda costaricensis Campbell, 1966 CRI

Lobopoda costaricensis Campbell, 1966: 103.

Lobopoda cubensis Campbell, 1966 CUB

Lobopoda cubensis Campbell, 1966: 157.

Lobopoda distans Campbell, 1971 CUB

Lobopoda distans Campbell, 1971: 48.

Lobopoda diversicauda Campbell, 1966 CRI

Lobopoda diversicauda Campbell, 1966: 106.

Lobopoda erythrocnemis (Germar, 1823)USA (AL AR FL GA KY LA MD MS NC SC TN TX)

Allecula erythrocnemis Germar, 1823: 164.

Lobopoda fallaciosa Campbell, 1971 CUB

Lobopoda fallaciosa Campbell, 1971: 49.

Lobopoda femoralis Champion, 1888MEX (CI SL TB VE) GUACRIPAN

Lobopoda femoralis Champion, 1888: 398.

Lobopoda foveata Champion, 1888 CRI PAN

Lobopoda foveata Champion, 1888: 405.

Lobopoda galapagoensis Linell, 1898SALNICPAN / SA

Lobopoda galapagoensis Linell, 1898: 266.

Lobopoda brunneipennis Campbell, 1966: 98. Synonymy: Peck and Kukalová-Peck (1990: 1637).

Lobopoda granulata Campbell, 1966CRIPANLAN (Barbados) / SA

Lobopoda granulata Campbell, 1966: 85.

Lobopoda guatemalensis Campbell, 1966 GUA

Lobopoda guatemalensis Campbell, 1966: 150.

Lobopoda guerrerensis Campbell, 1966MEX (GE)

Lobopoda guerrerensis Campbell, 1966: 152.

Lobopoda haitensis Campbell, 1966 HAI DOM

Lobopoda haitensis Campbell, 1966: 158.

Lobopoda hirta Champion, 1888 NIC

Lobopoda hirta Champion, 1888: 400.

Lobopoda hispaniolensis Campbell, 1971 DOM

Lobopoda hispaniolensis Campbell, 1971: 46.

Lobopoda insularis Champion, 1896 LAN

Lobopoda insularis Champion, 1896: 33.

Lobopoda jamaicensis Campbell, 1966 JAM

Lobopoda jamaicensis Campbell, 1966: 161.

Lobopoda laevicollis Champion, 1888MEX (CI VE YU)

Lobopoda laevicollis Champion, 1888: 401.

Lobopoda meridensis Campbell, 1966MEX (YU)

Lobopoda meridensis Campbell, 1966: 147.

Lobopoda micans Campbell, 1971 DOM

Lobopoda micans Campbell, 1971: 50.

Lobopoda minuta Champion, 1888 PAN

Lobopoda minuta Champion, 1888: 403.

Lobopoda monticola Campbell, 1966USA (TX)

Lobopoda monticola Campbell, 1966: 124.

Lobopoda mucronata Champion, 1888 PAN

Lobopoda mucronata Champion, 1888: 393.

Lobopoda nigrans (Melsheimer, 1846)USA (AL CT DC FL GA IL IN KS LA MA MD MI MS NC NJ NY OH PA RI SC TX VA)

Cistela atra Say, 1826: 242 [junior primary homonym of Cistela ater Fabricius, 1775 and Cistela atra Olivier, 1795].

Cistela nigrans Melsheimer, 1846: 60. Replacement name for Cistela atra Say, 1826.

Lobopoda nigrissima Campbell, 1966MEX (TA)

Lobopoda nigrissima Campbell, 1966: 133.

Lobopoda notapuncta Campbell, 1971 HAI DOM

Lobopoda notapuncta Campbell, 1971: 54.

Lobopoda oblonga Champion, 1888MEX (YU)

Lobopoda oblonga Champion, 1888: 396.

Lobopoda opaca Champion, 1888MEX (CI) GUACRIPAN

Lobopoda opaca Champion, 1888: 400.

Lobopoda biolleyi Pic, 1927: 22. Synonymy: Campbell (1966: 87).

Lobopoda opacicollis Champion, 1888USA (FL LA TX) MEX (GU NL SI TA VE) GUABELHONNIC

Lobopoda opacicollis Champion, 1888: 400.

Lobopoda subcuneata Casey, 1891: 79. Synonymy: Campbell (1966: 83).

Lobopoda panamensis Champion, 1888PAN / SA

Lobopoda panamensis Champion, 1888: 392.

Lobopoda paracollis Campbell, 1971 CUB

Lobopoda paracollis Campbell, 1971: 55.

Lobopoda paracornis Campbell, 1971 CUB

Lobopoda paracornis Campbell, 1971: 53.

Lobopoda parvula Champion, 1888MEX (JA VE)

Lobopoda parvula Champion, 1888: 403.

Lobopoda picipennis Campbell, 1971 HAI DOM

Lobopoda picipennis Campbell, 1971: 61.

Lobopoda pilosa Champion, 1888MEX (CI) GUA

Lobopoda pilosa Champion, 1888: 405.

Lobopoda polita Campbell, 1971 CUB

Lobopoda polita Campbell, 1971: 52.

Lobopoda proxima Champion, 1888MEX (CI VE YU) GUA

Lobopoda proxima Champion, 1888: 402.

Lobopoda puncticollis Champion, 1888 GUA

Lobopoda puncticollis Champion, 1888: 396.

Lobopoda punctulata (Melsheimer, 1846)USA (AL AR FL GA IA IL IN KS KY LA MD MO MS NC NJ NY OH OK PA SC TN TX VA WI) MEX (NL TA VE)

Cistela punctulata Melsheimer, 1846: 59.

Lobopoda jalapensis Champion, 1888: 402. Synonymy: Campbell (1966: 119).

Lobopoda oculatifrons Casey, 1891: 81. Synonymy: Campbell (1966: 119).

Lobopoda remoinsularis Campbell, 1966 CRI PAN

Lobopoda remoinsularis Campbell, 1966: 99.

Lobopoda sandersoni Campbell, 1971 DOM

Lobopoda sandersoni Campbell, 1971: 60.

Lobopoda sculpturata Champion, 1888 PAN

Lobopoda sculpturata Champion, 1888: 401.

Lobopoda seriata Champion, 1888MEX (YU)

Lobopoda seriata Champion, 1888: 395.

Lobopoda simplex Champion, 1888 BEL

Lobopoda simplex Champion, 1888: 399.

Lobopoda subparallela Champion, 1888MEX (CI GE MO OA VE)

Lobopoda subparallela Champion, 1888: 394.

Lobopoda substriatus Campbell, 1966 DOM

Lobopoda substriatus Campbell, 1966: 159.

Lobopoda sulcaticollis Pic, 1933 CUB

Lobopoda sulcaticollis Pic, 1933: 1.

Lobopoda tabogensis Campbell, 1966 PAN

Lobopoda tabogensis Campbell, 1966: 78.

Lobopoda teapensis Champion, 1893MEX (TB)

Lobopoda teapensis Champion, 1893a: 564.

Lobopoda tenuicornis Champion, 1888MEX (CI) PAN

Lobopoda tenuicornis Champion, 1888: 403.

Lobopoda terminalis Campbell, 1966 GUA

Lobopoda terminalis Campbell, 1966: 145.

Lobopoda thomasensis Campbell, 1971VIS (St. Thomas)

Lobopoda thomasensis Campbell, 1971: 50.

Lobopoda tropicalis Champion, 1888 PAN

Lobopoda tropicalis Champion, 1888: 398.

Lobopoda veracruzensis Campbell, 1966MEX (TA VE)

Lobopoda veracruzensis Campbell, 1966: 150.

Lobopoda viridis Champion, 1888MEX (CI VE) NIC

Lobopoda viridis Champion, 1888: 404.

Lobopoda longipes Borchmann, 1937: 218. Synonymy: Campbell (1966: 135).

Lobopoda yucatanica Champion, 1888MEX (YU)

Lobopoda yucatanica Champion, 1888: 397.

Lobopoda wittmeri Campbell, 1978 DOM

Lobopoda wittmeri Campbell, 1978c: 204.

Subgenus Mesolobopoda Campbell, 1966

Mesolobopoda Campbell, 1966: 34. Type species: Allecula socia LeConte, 1854, original designation.

Lobopoda acutangula Champion, 1888MEX (CI HI MI VE) GUABELNICCRIPAN

Lobopoda acutangula Champion, 1888: 390.

Lobopoda antiguaensis Campbell, 1971LAN (Antigua)

Lobopoda antiguaensis Campbell, 1971: 39.

Lobopoda ebenina Champion, 1896 LAN

Lobopoda ebenina Champion, 1896: 34.

Lobopoda socia (LeConte, 1854)USA (FL LA TX) MEX (JA NL SI SL TA TB VE YU) GUABELHONNIC

Allecula socia LeConte, 1854a: 84.

Lobopoda mexicana Champion, 1888: 392. Synonymy: Campbell (1966: 42).

Lobopoda trinidadensis Campbell, 1966MEX (YU) / SA

Lobopoda trinidadensis Campbell, 1966: 39.

Lobopoda tristis Champion, 1888 CRI PAN

Lobopoda tristis Champion, 1888: 391.

Subgenus Monoloba Solier, 1835

Monoloba Solier, 1835a: 235. Type species: Lobopoda dircaeoides Solier, 1835, monotypy.

Lobopoda asperula Champion, 1888MEX (YU)

Lobopoda asperula Champion, 1888: 390.

Lobopoda gigantea Champion, 1888MEX (VE)

Lobopoda gigantea Champion, 1888: 388.

Lobopoda grandis Champion, 1888NICPAN / SA

Lobopoda grandis Champion, 1888: 389.

Lobopoda tarsalis Fleutiaux and Sallé, 1890LAN (Guadeloupe)

Lobopoda tarsalis Fleutiaux and Sallé, 1890: 431.

[incertae sedis]

Lobopoda sordida (Horn, 1894)MEX (BC)

Allecula sordida Horn, 1894b: 432.

Genus Madreallecula Kanda, 2013 [F]

Madreallecula Kanda, 2013: 587. Type species: Madreallecula mcclevei Kanda, 2013, original designation.

Madreallecula mcclevei Kanda, 2013USA (AZ)

Madreallecula mcclevei Kanda, 2013: 588.

Genus Menes Champion, 1888 [M]

Menes Champion, 1888: 442. Type species: Menes meridanus Champion, 1888, subsequent designation (Bousquet et al. 2015: 134).

Menes meridanus Champion, 1888MEX (YU)

Menes meridanus Champion, 1888: 442.

Menes rotundatus Champion, 1888MEX (VE)

Menes rotundatus Champion, 1888: 443.

Genus Menoeceus Champion, 1888 [M]

Menoeceus Champion, 1888: 443. Type species: Menoeceus crassicornis Champion, 1888, subsequent designation (Casey 1891: 122).

Menoeceus aequalis Champion, 1888MEX (VE)

Menoeceus aequalis Champion, 1888: 444.

Menoeceus crassicornis Champion, 1888MEX (GE JA MO PU VE) GUA

Menoeceus crassicornis Champion, 1888: 444.

Menoeceus texanus Champion, 1888USA (TX)

Menoeceus texanus Champion, 1888: 444.

Genus Notacula Campbell, 1971 [F]

Notacula Campbell, 1971: 107. Type species: Notacula howdenae Campbell, 1971, original designation.

Notacula howdenae Campbell, 1971 JAM

Notacula howdenae Campbell, 1971: 107.

Genus Obesacula Campbell, 1971 [F]

Obesacula Campbell, 1971: 109. Type species: Obesacula aptera Campbell, 1971, original designation.

Obesacula aptera Campbell, 1971 JAM

Obesacula aptera Campbell, 1971: 111.

Cyrtosoma jamaicensis Marcuzzi, 1977: 42. Synonymy: Ivie (2005: 70).

Genus Parahymenorus Campbell, 1971 [M]

Parahymenorus Campbell, 1971: 100. Type species: Parahymenorus metallicus Campbell, 1971, original designation.

Parahymenorus metallicus caymanensis Campbell, 1971 CAY

Parahymenorus metallicus caymanensis Campbell, 1971: 103.

Parahymenorus metallicus metallicus Campbell, 1971 JAM

Parahymenorus metallicus metallicus Campbell, 1971: 102.

Genus Phedius Champion, 1888 [M]

Phedius Champion, 1888: 447. Type species: Phedius chevrolati Champion, 1888, subsequent designation (Lucas 1920: 500).

Phedius carbonarius Champion, 1888MEX (HI)

Phedius carbonarius Champion, 1888: 448.

Phedius chevrolati Champion, 1888MEX (VE)

Phedius chevrolati Champion, 1888: 447.

Phedius cylindricollis Champion, 1888MEX (JA)

Phedius cylindricollis Champion, 1888: 449.

Phedius funereus Schaeffer, 1905USA (AZ) MEX (SO)

Phedius funereus Schaeffer, 1905b: 176.

Phedius funestus Champion, 1888MEX (OA PU TA)

Phedius funestus Champion, 1888: 450.

Phedius hidalgoensis Champion, 1888MEX (HI)

Phedius hidalgoensis Champion, 1888: 448.

Phedius hirtus Champion, 1893MEX (GE)

Phedius hirtus Champion, 1893a: 568.

Phedius lapidicola Champion, 1893MEX (MO)

Phedius lapidicola Champion, 1893a: 568.

Phedius mexicanus Champion, 1888MEX (GU)

Phedius mexicanus Champion, 1888: 450.

Phedius obovatus Champion, 1888MEX (AG GU)

Phedius obovatus Champion, 1888: 449.

Phedius opaculus Horn, 1894MEX (BS)

Phedius opaculus Horn, 1894b: 431.

Genus Pitholaus Champion, 1888 [M]

Pitholaus Champion, 1888: 446. Type species: Pitholaus helopioides Champion, 1888, monotypy.

Pitholaus helopioides Champion, 1888 GUA

Pitholaus helopioides Champion, 1888: 446.

Genus Polyidus Champion, 1888 [M]

Polyidus Champion, 1888: 441. Type species: Polydius meridionalis Champion, 1888, monotypy.

Polyidus meridionalis Champion, 1888MEX (CI) GUACRI

Polyidus meridionalis Champion, 1888: 442.

Genus Punctacula Campbell, 1971 [F]

Punctacula Campbell, 1971: 112. Type species: Punctacula howdeni Campbell, 1971, original designation.

Punctacula howdeni Campbell, 1971 JAM

Punctacula howdeni Campbell, 1971: 114.

Genus Stenochidus LeConte, 1862 [M]

Stenochidus LeConte, 1862a: 244. Type species: Stenochia gracilis LeConte, 1851, subsequent designation (Lucas 1920: 608).

Stenochidus cyanescens (LeConte, 1859)USA (CA NV OR)

Prionychus cyanescens LeConte, 1859b: 78.

Stenochidus cyanescens var. carbonarius Schaeffer, 1911: 126. Synonymy: Marshall (1967a: 2).

Stenochidus gracilis (LeConte, 1851)USA (CA)

Stenochia gracilis LeConte, 1851: 150.

Stenochidus robustus Schaeffer, 1911USA (CA)

Stenochidus robustus Schaeffer, 1911: 125.

Genus Telesicles Champion, 1888 [M]

Telesicles Champion, 1888: 450. Type species: Telesicles cordatus Champion, 1888, monotypy.

Telesicles cordatus Champion, 1888USA (AZ CA CO NM TX UT) MEX (DU)

Telesicles cordatus Champion, 1888: 451.

Genus Temnes Champion, 1888 [M]

Temnes Champion, 1888: 410. Type species: Temnes caeruleus Champion, 1888, monotypy.

Temnes caeruleus Champion, 1888 PAN

Temnes caeruleus Champion, 1888: 410.

Genus Theatetes Champion, 1888 [M]

Theatetes Champion, 1888: 420. Type species: Theatetes basicornis Champion, 1888, monotypy.

Theatetes basicornis Champion, 1888MEX (VE)

Theatetes basicornis Champion, 1888: 420.

Subtribe Gonoderina Seidlitz, 1896

Gonoderina Seidlitz, 1896: 83. Type genus: Gonodera Mulsant, 1856.

Pseudocistelini Portevin, 1934: 39. Type genus: Pseudocistela Crotch, 1874.

Genus Andrimus Casey, 1891 [M]

Andrimus Casey, 1891: 155. Type species: Cteniopus murrayi LeConte, 1866, subsequent designation (Lucas 1920: 96).

Andrimus murrayi (LeConte, 1866)USA (AL FL GA NC SC VA)

Cteniopus murrayi LeConte, 1866b: 141.

Andrimus brunneus Casey, 1891: 157. Synonymy: Peck and Thomas (1998: 109).

Andrimus concolor Casey, 1891: 158. New synonymy [based on Marshall (1964: 156) unpublished thesis].

Andrimus nigrescens Casey, 1891: 159. Synonymy: Peck and Thomas (1998: 109).

Andrimus convergens Casey, 1891: 159. New synonymy [based on Marshall (1964: 156) unpublished thesis].

Andrimus confusus Blatchley, 1912: 331. Synonymy: Peck and Thomas (1998: 109).

Andrimus parvulus Blatchley, 1919: 67. Synonymy: Peck and Thomas (1998: 109).

Genus Androchirus LeConte, 1862 [M]

Androchirus LeConte, 1862a: 244. Type species: Cistela fuscipes Melsheimer, 1846 (= Cistela erythropa Kirby, 1837), original designation.

Androchirus erythropus (Kirby, 1837) [Fig. 32] CAN (MB NB NS ON QC) USA (AL CT DC DE IA IL IN KS KY MA MD MI MN NC NH NJ NY OH PA SC VA VT WI)

Figure 32. 

Androchirus erythropus (Kirby, 1837). Scale bar = 1 mm.

Cistela erythropa Kirby, 1837: 239.

Cistela fuscipes Melsheimer, 1846: 60. Synonymy: Casey (1891: 169).

Androchirus luteipes LeConte, 1862a: 245. Synonymy: Horn (1876b: 192).

Androchirus femoralis (Olivier, 1791)USA (FL GA LA MS OH RI SC TX)

Cistela femoralis Olivier, 1791: 6.

Genus Capnochroa LeConte, 1862 [F]

Capnochroa LeConte, 1862a: 244. Type species: Cistela fuliginosa Melsheimer, 1846, monotypy.

Capnochroa fuliginosa (Melsheimer, 1846)CAN (NB NS ON PE QC) USA (CT DC DE GA IA IL IN KY MA MD ME MI MO NC NH NJ NY OH PA RI SC TN VA WI WV)

Cistela fuliginosa Melsheimer, 1846: 59.

Genus Chromatia LeConte, 1862 [F]

Chromatia LeConte, 1862a: 244. Type species: Cistela amoena Say, 1824, monotypy.

Chromatia amoena (Say, 1824)CAN (ON QC) USA (CT GA IN KS MA MD MN NH NJ NY OH PA SC TN TX WI)

Cistela amoena Say, 1824a: 268.

Genus Isomira Mulsant, 1856 [F]

Isomira Mulsant, 1856: 52. Type species: Chrysomela murina Linnaeus, 1758, subsequent designation (C.G. Thomson 1859: 119).

Tedinus Casey, 1891: 153. Type species: Tedinus angustus Casey, 1891, monotypy. Synonymy: Bousquet and Campbell (1991: 259).

Isomira acuta Campbell, 1968MEX (CI) GUA

Isomira acuta Campbell, 1968: 460.

Isomira alticola Campbell, 1968 GUA

Isomira alticola Campbell, 1968: 465.

Isomira angusta (Casey, 1891)USA (GA SC)

Tedinus angustus Casey, 1891: 154.

Isomira brevicollis Champion, 1888MEX (VE)

Isomira brevicollis Champion, 1888: 459.

Isomira championi Campbell, 1968MEX (NL)

Isomira championi Campbell, 1968: 453.

Isomira comstocki Papp, 1956CAN (AB BC) USA (AZ CA ID NV OR UT WA WY)

Isomira comstocki Papp, 1956: 147.

Isomira damnata Marshall, 1970USA (CA)

Isomira damnata Marshall, 1970c: 4.

Isomira evanescens Champion, 1888 GUA

Isomira evanescens Champion, 1888: 458.

Isomira howdeni hidalgoensis Campbell, 1968MEX (HI)

Isomira howdeni hidalgoensis Campbell, 1968: 455.

Isomira howdeni howdeni Campbell, 1968MEX (DU)

Isomira howdeni howdeni Campbell, 1968: 454.

Isomira iowensis Casey, 1891CAN (ON) USA (AR FL GA IA IL KS MD MO NC OH PA SC TN TX VA)

Isomira iowensis Casey, 1891: 145.

Isomira luscitiosa Casey, 1891USA (CA)

Isomira luscitiosa Casey, 1891: 148.

Isomira mexicana Campbell, 1968MEX (CI GE OA VE)

Isomira mexicanus Campbell, 1968: 457.

Isomira monticola Casey, 1891USA (CA)

Isomira monticola Casey, 1891: 150.

Isomira oblongula Casey, 1891CAN (ON QC) USA (FL IL IN MI NC NY OH PA SC TX WI)

Isomira oblongula Casey, 1891: 151.

Isomira obsoleta Champion, 1888MEX (GE OA VE) GUA

Isomira obsoleta Champion, 1888: 457.

Isomira pulla (Melsheimer, 1846)CAN (ON QC) USA (AL AR DC DE FL GA IA IL IN KY MA MD ME MI MN MS NC NJ NY OH PA RI SC TN VA WI)

Cistela pulla Melsheimer, 1846: 60.

Isomira ignora Blatchley, 1914: 144. Synonymy: Marshall (1970a: 44).

Isomira quadristriata (Couper, 1865)CAN (MB NB NS ON PE QC SK) USA (CT FL GA IL IN MA MD ME MI MN NC NH NJ NY OH PA RI SC TN VA WI WV)

Cistela quadristriata Couper, 1865: 62.

Isomira velutina LeConte, 1866b: 139. Synonymy: Casey (1891: 149).

Isomira rotundata Campbell, 1968MEX (SL TA)

Isomira rotundata Campbell, 1968: 456.

Isomira ruficollis Hamilton, 1893USA (IN KY OH PA)

Isomira ruficollis Hamilton, 1893: 308.

Isomira sericea (Say, 1824)CAN (NB NS ON QC) USA (AR CT DC DE FL GA IA IL IN KY MA MD ME MI MN MO MS NC NH NJ NY OH PA RI SC TN VA WI WV) / BAH

Cistela sericea Say, 1824a: 270.

Isomira tenebrosa Casey, 1891: 146. Synonymy: Marshall (1970a: 46).

Isomira subaenea guatemalensis Campbell, 1968 GUA

Isomira subaenea guatemalensis Campbell, 1968: 464.

Isomira subaenea punctata Campbell, 1968 GUA

Isomira subaenea punctata Campbell, 1968: 464.

Isomira subaenea soror Campbell, 1968 GUA

Isomira subaenea soror Campbell, 1968: 463.

Isomira subaenea subaenea Champion, 1888MEX (CI) GUA

Isomira subaenea Champion, 1888: 458.

Isomira texana Casey, 1891USA (TX)

Isomira texana Casey, 1891: 153.

Isomira valida Schwarz, 1878CAN (ON) USA (AL AR FL GA IL IN KS MD NJ OH SC WI WV)

Isomira valida Schwarz, 1878: 370.

Isomira similis Blatchley, 1910: 1278. Synonymy: Marshall (1970a: 41).

Isomira variabilis (Horn, 1875)USA (AZ CA)

Cistela variabilis Horn, 1875: 156.

Isomira discolor Casey, 1891: 145. Synonymy: Marshall (1970c: 2).

Genus Onychomira Campbell, 1984 [F]

Onychomira Campbell, 1984: 289. Type species: Onychomira floridensis Campbell, 1984, original designation.

Onychomira floridensis Campbell, 1984USA (FL)

Onychomira floridensis Campbell, 1984: 291.

Genus Pseudocistela Crotch, 1874 [F]

Pseudocistela Crotch, 1874: 108. Type species: Cistela brevis Say, 1824, subsequent designation (Novák and Pettersson 2008: 327).

Pseudocistela alternans (Champion, 1888)MEX (OA)

Cistela alternans Champion, 1888: 456.

Pseudocistela brevis (Say, 1824)CAN (NB ON QC) USA (CT DC FL IL IN MD MI MN MO NC NH NJ NY OH PA SC VA VT WI)

Cistela brevis Say, 1824a: 269 [junior primary homonym of Cistela brevis Illiger, 179465].

Cistela erythroptera Ziegler, 1844: 46. Synonymy: Gemminger [in Gemminger and Harold] (1870: 2047).

Pseudocistela calida (Champion, 1888) PAN

Cistela calida Champion, 1888: 453.

Pseudocistela chiriquensis (Champion, 1888) PAN

Cistela chiriquensis Champion, 1888: 454.

Pseudocistela cinerascens (Champion, 1888)MEX (PU)

Cistela cinerascens Champion, 1888: 453.

Pseudocistela decepta (Champion, 1888) PAN

Cistela decepta Champion, 1888: 454.

Pseudocistela delitescens (Champion, 1888) GUA

Cistela delitescens Champion, 1888: 455.

Pseudocistela fragilicornis (Champion, 1888) GUA

Cistela fragilicornis Champion, 1888: 457.

Pseudocistela juquilae (Champion, 1888)MEX (OA)

Cistela juquilae Champion, 1888: 456.

Pseudocistela marginata (Ziegler, 1844)USA (CT GA MA MD NC NJ NY PA)

Cistela marginata Ziegler, 1844: 46.

Pseudocistela nigricornis (Champion, 1888)MEX (CI DU GE GU VE) NICCRIPAN

Cistela nigricornis Champion, 1888: 452.

Pseudocistela occulta (Champion, 1888) GUA

Cistela occulta Champion, 1888: 455.

Pseudocistela opaca (LeConte, 1859)USA (CA ID NV)

Xystropus opacus LeConte, 1859b: 78.

Cistela thevenetii Horn, 1875: 156. Synonymy: Poole and Gentili (1996: 441) [probably based on Marshall (1964: 87) unpublished thesis].

Pseudocistela ovipennis (Champion, 1893)MEX (GE JA)

Cistela ovipennis Champion, 1893a: 569.

Pseudocistela pectinata Hopping, 1933CAN (BC)

Pseudocistela pectinata Hopping, 1933: 285.

Pseudocistela pinguis (LeConte, 1859)CAN (BC) USA (CA CO ID NM NV OR WA)

Xystropus pinguis LeConte, 1859a: 16.

Pseudocistela pacifica Hopping, 1933: 284. Synonymy: Bousquet and Campbell (1991: 260).

Pseudocistela sandersoni Campbell, 1971 JAM

Pseudocistela sandersoni Campbell, 1971: 17.

Pseudocistela zunilensis (Champion, 1888) GUA

Cistela zunilensis Champion, 1888: 452.

Subtribe Mycetocharina Gistel, 1848

Mycetocharisidae Gistel, 1848: [10]. Type genus: Mycetochara Guérin-Méneville, 1827.

Genus Hymenochara Campbell, 1978 [F]

Hymenochara Campbell, 1978a: 435. Type species: Mycetophila rufipes J.E. LeConte, 1824, original designation.

Hymenochara arizonensis Campbell, 1978USA (AZ)

Hymenochara arizonensis Campbell, 1978a: 440.

Hymenochara rufipes (J.E. LeConte, 1824)CAN (ON QC) USA (AL FL IN KY MA MD MI MO MS NC NY OH PA SC)

Mycetophila rufipes J.E. LeConte, 1824: 170.

Genus Mycetochara Guérin-Méneville, 1827 [F]

Mycetophila Gyllenhal, 1810: 541 [junior homonym of Mycetophila Meigen, 1803]. Type species: Cistela scapularis Illiger, 1805 (= Cistela humeralis Fabricius, 1787), subsequent designation (Westwood 1838: 32).

Mycetochara Guérin-Méneville, 1827: 346. Replacement name for Mycetophila Gyllenhal, 1810. Note. See Bousquet et al. (2015: 138) for authorship of this name.

Mycetocharis Gyllenhal, 1827: 510. Replacement name for Mycetophila Gyllenhal, 1810.

Mycetochares Latreille, 1829a: 42. Replacement name for Mycetophila Gyllenhal, 1810.

Stigmatoma LeConte, 1862a: 244. Type species: Cistela fraterna Say, 1824, monotypy. Synonymy: LeConte (1866b: 139).

Mycetochara analis (LeConte, 1878)CAN (AB BC MB NB ON QC SK) USA (AR AZ CO GA IN KS MA MD MI NE NJ NY OH OR PA SC WA WI)

Mycetochares analis LeConte, 1878b: 618.

Mycetochares lugubris LeConte, 1878b: 618. Synonymy: Campbell (1978b: 931).

Mycetochara horni Dury, 1902: 172. Synonymy: Campbell (1978b: 931).

Mycetochara davisi Hatch, 1965: 187. Synonymy: Campbell (1978b: 931).

Mycetochara basillaris (Say, 1824)USA (PA)

Cistela basillaris Say, 1824a: 269.

Mycetochara bicolor (Couper, 1865) [Fig. 33] CAN (NB NS ON QC) USA (IA IN MA ME MI MO NC NH NY PA SC VA VT WI)

Figure 33. 

Mycetochara bicolor (Couper, 1865). Scale bar = 1 mm.

Mycetochares bicolor Couper, 1865: 62.

Mycetochara binotata (Say, 1824)CAN (MB NB NS ON QC) USA (CT IN KY MA MD MI MN NC NH NY OH PA SC VA WI)

Cistela binotata Say, 1824b: 285.

Mycetochares marginata LeConte, 1878b: 618. Synonymy: Campbell (1978b: 932).

Mycetochares longula LeConte, 1878b: 618. Synonymy: Campbell (1978b: 932).

Mycetochara foveata (LeConte, 1866)CAN (NB ON QC) USA (IA IL IN MA MD MI MO NC NH NJ NY OH RI SC VA WI)

Mycetochares foveata LeConte, 1866b: 140.

Mycetochares tenuis LeConte, 1866b: 140. Synonymy: Campbell (1978b: 929).

Mycetochares gracilis LeConte, 1878b: 615 [junior primary homonym of Mycetochares gracilis Falderman, 1837]. Synonymy: Campbell (1978b: 929).

Mycetochara gilvipes Casey, 1891: 131. Synonymy: Campbell (1978b: 929).

Mycetochara lecontei Borchmann, 1909: 714. Replacement name for Mycetochara gracilis (LeConte, 1878).

Mycetochara fraterna (Say, 1824)CAN (AB BC MB NB NS ON QC SK) USA (CT DC IA IL IN MA MD ME NC NH NJ NY OH PA SC SD VA WI)

Cistela fraterna Say, 1824a: 270.

Mycetochares laticollis LeConte, 1878b: 617. Synonymy: Casey (1891: 128).

Mycetochara megalops Casey, 1891: 129. Synonymy: Campbell (1978b: 925).

Mycetochara nigerrima Casey, 1891: 132. Synonymy: Campbell (1978b: 925).

Mycetochara haldemani (LeConte, 1866)USA (FL GA IL IN MD NC NY OH PA VA WI)

Mycetochares haldemani LeConte, 1866b: 140.

Mycetochara lata Hatch, 1965USA (OR)

Mycetochara lata Hatch, 1965: 189.

Mycetochara perplexata Marshall, 1970USA (CA)

Mycetochara perplexata Marshall, 1970b: 3.

Mycetochara marshalli Campbell, 1978b: 934. New synonymy [YB].

Mycetochara procera Casey, 1891CAN (BC) USA (AZ CA ID NV OR WA)

Mycetochara procera Casey, 1891: 140.

Mycetochara pacifica Casey, 1891: 139. Synonymy: Campbell (1978b: 936).

Mycetochara nevadensis Casey, 1891: 142. Synonymy: Campbell (1978b: 936).

Mycetochara crassulipes Casey, 1891: 142. Synonymy: Campbell (1978b: 936).

Mycetochara downei Hatch, 1965: 187. Synonymy: Campbell (1978b: 937).

Mycetochara angusta Hatch, 1965: 188. Synonymy: Campbell (1978b: 937).

Mycetochara malkini Hatch, 1965: 188. Synonymy: Campbell (1978b: 937).

Mycetochara caseyi Hatch, 1965: 188. Synonymy: Campbell (1978b: 937).

Mycetochara pubipennis (LeConte, 1878)USA (CA)

Mycetochares pubipennis LeConte, 1878b: 617.

Mycetochara longipennis Casey, 1891: 139. Synonymy: Campbell (1978b: 935).

Mycetochara ruficornis Melsheimer, 1846USA (PA)

Mycetocharus ruficornis Melsheimer, 1846: 59.

Subtribe Xystropodina Solier, 1835

Xystropides Solier, 1835a: 229. Type genus: Xystropus Solier, 1835.

Lystronychides Lacordaire, 1859: 512. Type genus: Lystronychus Latreille, 1829.

Genus Anamphidora Casey, 1924 [F]

Anamphidora Casey, 1924: 330. Type species: Anamphidora parvula Casey, 1924, original designation.

Anamphidora campbelli Marshall, 1967USA (TX)

Anamphidora campbelli Marshall, 1967b: 209.

Anamphidora kimberleei Marshall, 1970MEX (BC)

Anamphidora kimberleei Marshall, 1970d: 294.

Anamphidora parvula Casey, 1924MEX (DU)

Anamphidora parvula Casey, 1924: 330.

Genus Cteisa Solier, 1835 [F]

Cteisa Solier, 1835a: 242. Type species: Cteisa hirta Solier, 1835, monotypy.

Cteisa pedinoides Mäklin, 1875MEX (VE) GUAPAN / JAM / SA

Cteisa pedinoides Mäklin, 1875b: 681.

Genus Erxias Champion, 1888 [M]

Erxias Champion, 1888: 460. Type species: Erxias bicolor Champion, 1888, subsequent designation (Bousquet et al. 2015: 138).

Erxias bicolor Champion, 1888PAN / SA

Erxias bicolor Champion, 1888: 460.

Erxias violaceipennis Champion, 1888 NIC

Erxias violaceipennis Champion, 1888: 460.

Genus Lystronychus Latreille, 1829 [M]

Lystronychus Latreille, 1829a: 41 [as Lystronichus]. Type species: Helops equestris Fabricius, 1775, subsequent designation (Saunders 1836: 154). Note. Lystronychus is in prevailing usage (see Bouchard et al. 2011: 427) and so deemed to be the correct original spelling.

Subgenus Lystronychus Latreille, 1829

Lystronychus Latreille, 1829a: 41 [as Lystronichus]. Type species: Helops equestris Fabricius, 1775, subsequent designation (Saunders 1836: 154).

Lystronychus championi Horn, 1894USA (TX)

Lystronychus championi Horn, 1894b: 433.

Lystronychus delauneyi (Fleutiaux and Sallé, 1890)LAN (Guadeloupe)

Anaedus delauneyi Fleutiaux and Sallé, 1890: 428.

Lystronychus piliferus Champion, 1888USA (TX) MEX (CI CL JA OA PU VE) GUANIC / SA

Lystronychus piliferus Champion, 1888: 462.

Lystronychus purpureipennis Champion, 1888 GUA

Lystronychus purpureipennis Champion, 1888: 463.

Lystronychus rufonotatus Champion, 1896LAN (St. Vincent)

Lystronychus rufonotatus Champion, 1896: 35.

Lystronychus rufulus Borchmann, 1930MEX (PU)

Lystronychus rufulus Borchmann, 1930: 98.

Lystronychus scapularis Champion, 1888USA (AZ) MEX (YU) GUANICPAN

Lystronychus scapularis Champion, 1888: 463.

Lystronychus tuberculifer Champion, 1896LAN (Grenadines)

Lystronychus tuberculifer Champion, 1896: 34.

Genus Prostenus Klug, 1829 [M]

Prostenus Klug, 1829: 5. Type species (suggested): Prostenus periscelis Perty, 1830 (see Bousquet et al. 2015: 139).

Mecocerus Solier, 1835a: 241 [junior homonym of Mecocerus Schönherr, 1833]. Type species: Xystropus dejeanii Solier, 1835, monotypy. Synonymy: Lacordaire (1859: 513).

Prostenus panamensis Champion, 1888 PAN

Prostenus panamensis Champion, 1888: 461.

Genus Xystropus Solier, 1835 [M]

Xystropus Solier, 1835a: 241. Type species: Xystropus pilosus Solier, 1835, monotypy. Note. See Bousquet et al. (2015: 139) for available species originally included in this genus.

Xystropus californicus (Horn, 1868)MEX (OA) NICCRIPAN / SA

Prostenus californicus Horn, 1868: 138.

Xystropus fulgidus Mäklin, 1875b: 680. Synonymy: Casey (1891: 74).

Xystropus fallax Mäklin, 1875PAN / SA

Xystropus fallax Mäklin, 1875b: 677.

Xystropus lebasii Mäklin, 1875PAN / SA

Xystropus lebasii Mäklin, 1875b: 679.

Subfamily DIAPERINAE Latreille, 1802

Diaperialae Latreille, 1802: 161. Type genus: Diaperis Geoffroy, 1762.

Tribe Crypticini Brullé, 1832

Crypticites Brullé, 1832: 190. Type genus: Crypticus Latreille, 1816.

Genus Ellipsodes Wollaston, 1854 [M]

Ellipsodes Wollaston, 1854: 485. Type species: Sphaeridium glabratum Fabricius, 1781, monotypy.

Subgenus Anthrenopsis Koch, 1950

Anthrenopsis Koch, 1950: 74. Type species: Platydema scriptipenne Fairmaire, 1875 (= Basides ziczac Motschulsky, 1873), original designation.

Ellipsodes ziczac (Motschulsky, 1873)LAN (Guadeloupe, Grenada) – Adventive

Basides ziczac Motschulsky, 1873: 475.

Platydema scriptipenne Fairmaire, 1875: xxxiii. Synonymy: Kaszab (1975: 102).

Genus Gondwanocrypticus Español, 1955 [M]

Gondwanocrypticus Español, 1955: 10. Type species: Crypticus platensis Fairmaire, 1884, original designation.

Gondwanocrypticus aterrimus (Champion, 1886)MEX (CI) GUABELSALHONNICCRIPAN / SA

Crypticus aterrimus Champion, 1886: 138.

Gondwanocrypticus filicornis (Chevrolat, 1878) JAM DOM LAN

Platydema filicorne Chevrolat, 1878b: 222.

Gondwanocrypticus maculatus (Champion, 1886)MEX (MO) GUANIC

Crypticus maculatus Champion, 1886: 138.

Gondwanocrypticus mexicanus (Champion, 1886)MEX (VE)

Crypticus mexicanus Champion, 1886: 137.

Gondwanocrypticus obsoletus (Say, 1824)USA (DE FL GA LA MD MS NC SC TX VA) / CUB

Crypticus obsoletus Say, 1824a: 265.

Gondwanocrypticus ovatus (Champion, 1886)MEX (JA OA) GUA

Crypticus ovatus Champion, 1886: 137.

Gondwanocrypticus pictus (Gebien, 1928)USAi (AL FL GA MS NC SC) / SA

Crypticus pictus Gebien, 1928a: 118.

Gondwanocrypticus platensis (Fairmaire, 1884)USA (AL CA DE FL GA LA MD MS NC SC TX VA WV) / BAHCAY / SA – Adventive

Crypticus platensis Fairmaire, 1884: 510.

Gondwanocrypticus undatus (Champion, 1896) LAN

Crypticus undatus Champion, 1896: 5.

Genus Poecilocrypticus Gebien, 1928 [M]

Poecilocrypticus Gebien, 1928a: 121. Type species: Poecilocrypticus formicophilus Gebien, 1928, monotypy.

Poecilocrypticus formicophilus Gebien, 1928USAi (AL FL GA LA MS NC OK SC TX) / BAH / SA

Poecilocrypticus formicophilus Gebien, 1928a: 122.

Tribe Diaperini Latreille, 1802

Diaperialae Latreille, 1802: 161. Type genus: Diaperis Geoffroy, 1762.

Subtribe Adelinina LeConte, 1862

Alphitophagida Gistel, 1856b: 18566. Type genus: Alphitophagus Stephens, 1832.

Adelinini LeConte, 1862a: 237. Type genus: Adelina Dejean, 1835.

Schedarosini Reitter, 1876: 42. Type genus: Schedarosus Reitter, 1876 (= Adelina Dejean, 1835).

Doliemini Reitter, 1917: 58. Type genus: Doliema Pascoe, 1860 (= Adelina Dejean, 1835).

Gnathocerini Skopin, 1978: 228. Type genus: Gnatocerus Thunberg, 1814.

Genus Adelina Dejean, 1835 [F]

Adelina Dejean, 1835: 315. Type species: Cucujus planus Fabricius, 1801, monotypy.

Doliema Pascoe, 1860: 50. Type species: Doliema platisoides Pascoe, 1860, monotypy. Synonymy: Fleutiaux and Sallé (1890: 428, as Adelina LeConte).

Schedarosus Reitter, 1876: 42. Type species: Schedarosus cucujiformis Reitter, 1876 (= Pytho pallida Say, 1823), subsequent designation (Löbl et al. 2008b: 42). Synonymy (with Doliema Pascoe): Champion (1886: 157).

Adelina angustata (Champion, 1886) GUA NIC

Doliema angustata Champion, 1886: 159.

Adelina bacardi Steiner, 2006 BAH

Adelina bacardi Steiner, 2006: 15.

Adelina bidens (Schaeffer, 1915)USA (FL TX) GUA / BAHCUBCAYDOM

Doliema bidens Schaeffer, 1915: 238.

Adelina bifurcata (Champion, 1893)USA (AZ) MEX (BS JA OA VE YU) CRI

Doliema bifurcata Champion, 1893a: 535.

Adelina dominicana (Ardoin, 1977) DOM

Doliema dominicana Ardoin, 1977a: 18.

Adelina frontalis (Champion, 1886)BEL / SA

Doliema frontalis Champion, 1886: 159.

Adelina klapperichi (Ardoin, 1977) DOM

Doliema klapperichi Ardoin, 1977a: 12.

Adelina latiramosa Doyen, 1984MEX (PU)

Adelina latiramosa Doyen, 1984a: 777.

Adelina maryjoae Steiner, 2005 BAH CAY

Adelina maryjoae Steiner, 2005: 449.

Adelina mystax Triplehorn and Ivie, 1983 VIS

Adelina mystax Triplehorn and Ivie, 1983: 272.

Adelina pallida (Say, 1824)USA (CA FL GA IN LA MD NC OH SC TN VA) MEX (CH HI VE YU) GUABELNIC / CUBPRI / SA

Pytho pallida Say, 1824a: 271.

Schedarosus cucujiformis Reitter, 1876: 43. Synonymy: Champion (1893a: 535).

Adelina pici (Ardoin, 1977)BAHCUBCAYLAN / SA

Doliema pici Ardoin, 1977a: 7.

Adelina plana (Fabricius, 1801)USA (AZ CA FL IN NC) MEX (JA TB VE) GUABELHONNICPAN / BAHCUBCAYDOMLAN / SA

Cucujus planus Fabricius, 1801b: 94.

Adelina depressa Erichson, 1847: 119. Synonymy (in doubt): Champion (1893a: 535).

Adelina plana LeConte, 1851: 149 [junior secondary homonym of Adelina plana (Fabricius, 1801)]. Synonymy (in doubt): Champion (1886: 157).

Sitophagus lecontei Horn, 1870: 346. Replacement name for Sitophagus planus (LeConte, 1851).

Schedarosus scidarius Reitter, 1876: 44. Synonymy: Champion (1886: 158).

Doliema diabolica Pic, 1923: 24. Synonymy: Ardoin (1977a: 3).

Adelina quadridentata (Champion, 1893)MEX (JA OA) CRI

Doliema quadridentata Champion, 1893a: 535.

Genus Alphitophagus Stephens, 1832 [M]

Alphitophagus Stephens, 1832: 12. Type species: Alphitophagus quadripustulatus Stephens, 1832 (= Diaperis bifasciata Say, 1824), monotypy.

Phyletes Redtenbacher, 1845: 128. Type species: Phylethus populi Redtenbacher, 1848 (= Diaperis bifasciata Say, 1824), subsequent monotypy in Redtenbacher (1848: 589, as Phylethus). Synonymy: Seidlitz (1894: 533).

Alphitophagus bifasciatus (Say, 1824) [Fig. 34] CAN (MB ON QC SK) USA (AL AR CA DC GA IA ID IL IN KS LA MD MN MO MS NC NE NJ NY OH OR PA SC SD TN TX VA WA WI) – Adventive

Figure 34. 

Alphitophagus bifasciatus (Say, 1824). Scale bar = 1 mm.

Diaperis bifasciata Say, 1824a: 268.

Alphitophagus quadripustulatus Stephens, 1832: 12. Synonymy: Horn (1870: 385).

Platydema lilliputanum Carter, 1937: 130. Synonymy: Hinton (1947b: 90).

Genus Cynaeus LeConte, 1862 [M]

Cynaeus LeConte, 1862a: 233. Type species: Platydema angustum LeConte, 1851, original designation.

Cynaeus angustus (LeConte, 1851) [Fig. 35] CAN (AB BC MB ON QC SK) USA (AZ CA CO GA IA ID IL IN KS MD MI MN NC ND NM OH OR SD TN TX UT VA WA WI WY) MEX (BC SO)

Figure 35. 

Cynaeus angustus (LeConte, 1851). Scale bar = 1 mm.

Platydema angustum LeConte, 1851: 149.

Cynaeus opacus Champion, 1886: 156. Synonymy: Blaisdell (1943: 267).

Cynaeus depressus Horn, 1870USA (AZ CA) MEX (BC)

Cynaeus depressus Horn, 1870: 369.

Genus Doliodesmus Spilman, 1967 [M]

Doliodesmus Spilman, 1967: 149. Type species: Doliodesmus charlesi Spilman, 1967, monotypy.

Doliodesmus charlesi Spilman, 1967USA (AZ) MEX (BC BS)

Doliodesmus charlesi Spilman, 1967: 153.

Genus Doliopines Horn, 1894 [M]

Doliopines Horn, 1894b: 427. Type species: Doliopines cucujinus Horn, 1894, monotypy.

Doliopines cucujinus Horn, 1894MEX (BC BS SO)

Doliopines cucujinus Horn, 1894b: 428.

Genus Gnatocerus Thunberg, 1814 [M]

Gnatocerus Thunberg, 1814: 47. Type species: Gnatocerus ruber Thunberg, 1814 (= Trogosita cornuta Fabricius, 1798), monotypy.

Gnathocerus Agassiz, 1846: 164. Unjustified emendation of Gnatocerus Thunberg, 1814, not in prevailing usage.

Subgenus Echocerus Horn, 1870

Echocerus Horn, 1870: 366. Type species: Trogosita maxillosa Fabricius, 1801, monotypy.

Gnatocerus analis (Champion, 1886) GUA

Echocerus analis Champion, 1886: 146.

Gnatocerus angelicus (Blaisdell, 1923)MEX (BC)

Echocerus angelicus Blaisdell, 1923: 277.

Gnatocerus breviceps (Blaisdell, 1943)MEX (BS)

Echocerus breviceps Blaisdell, 1943: 266.

Gnatocerus curvicornis (Champion, 1893)USA (FL) MEX (JA OA YU) / BAHCUBCAY / LAN

Echocerus curvicornis Champion, 1893a: 533.

Echocerus recurvatus Chittenden, 1895a: 2. Synonymy: Chittenden (1895b: 331).

Gnatocerus maxillosus (Fabricius, 1801)USA (CA FL GA KS MD MI OH SC TX WI) MEX (GU) GUANIC / CUBPRILAN / SA – Adventive

Trogosita maxillosa Fabricius, 1801a: 155.

Subgenus Gnatocerus Thunberg, 1814

Gnatocerus Thunberg, 1814: 47. Type species: Gnatocerus ruber Thunberg, 1814 (= Trogosita cornuta Fabricius, 1798), monotypy.

Cerandria Dejean, 1834: 200. Type species: Trogosita cornuta Fabricius, 1798, subsequent designation (Duponchel 1841: 285). Synonymy: Schaum (1849: 283).

Sicinus Champion, 1886: 146. Type species: Sicinus guatemalensis Champion, 1886, subsequent designation (Löbl et al. 2008b: 43). Synonymy: Leng (1920: 233).

Gnatocerus brevipes (Champion, 1886) GUA

Sicinus brevipes Champion, 1886: 147.

Gnatocerus cornutus (Fabricius, 1798) [Fig. 36] CAN (BC MB NS ON QC) USA (CA CT FL MA MD OH OR WA) MEX (GU VE) GUA / CUBPRI – Adventive

Figure 36. 

Gnatocerus (Gnatocerus) cornutus (Fabricius, 1798). Scale bar = 1 mm.

Trogosita cornuta Fabricius, 1798: 51.

Trogossita maxillaris Palisot de Beauvois, 1812: 125; pl. 32 (fig. 4)67. Synonymy: Fauvel (1904: 174).

Gnatocerus guatemalensis (Champion, 1886)USA (FL IL KY MD OH PA TX) MEXGUA / BAHCUBLAN

Sicinus guatemalensis Champion, 1886: 147.

Echocerus dentiger Chittenden, 1895a: 1. Synonymy: Leng (1918: 208).

Genus Iccius Champion, 1886 [M]

Iccius Champion, 1886: 147. Type species: Iccius cephalotes Champion, 1886, subsequent designation (Gebien 1940: 760).

Iccius cephalotes Champion, 1886MEX (VE) GUAPAN / CUB

Iccius cephalotes Champion, 1886: 148.

Iccius cylindricus Champion, 1886USA (AZ LA TX) MEX (MO) GUA

Iccius cylindricus Champion, 1886: 148.

Iccius elongatus Kulzer, 1949 CRI

Iccius elongatus Kulzer, 1949: 304.

Iccius grenadensis Champion, 1896 LAN

Iccius grenadensis Champion, 1896: 19.

Iccius monoceros Ferrer and Ødegaard, 2005 PAN

Iccius monoceros Ferrer and Ødegaard, 2005: 637.

Iccius rufotestaceus Champion, 1896 LAN

Iccius rufotestaceus Champion, 1896: 18.

Hypophlaeus dufaui Pic, 1945: 7. Synonymy: Bremer and Triplehorn (1999: 59).

Genus Loxostethus Triplehorn, 1962 [M]

Loxostethus Triplehorn, 1962: 504. Type species: Loxostethus fasciatus Triplehorn, 1962, original designation.

Loxostethus erythroscelis Triplehorn and Merkl, 1997 HAI DOM

Loxostethus erythroscelis Triplehorn and Merkl, 1997: 739.

Loxostethus fasciatus Triplehorn, 1962 CUB

Loxostethus fasciatus Triplehorn, 1962: 504.

Loxostethus quadrimaculata Zayas, 1988: 93. Synonymy: Ivie (1991: 400).

Loxostethus gibbosus Triplehorn and Merkl, 1997 CUB

Loxostethus gibbosus Triplehorn and Merkl, 1997: 738.

Loxostethus gowdeyi (Pic, 1930) CUB JAM HAI DOM

Pentaphyllus gowdeyi Pic, 1930: 33.

Loxostethus jamaicensis Triplehorn, 1962: 506. Synonymy: Triplehorn and Merkl (1997: 739).

Loxostethus opacifrons Triplehorn, 1962: 506. Synonymy: Triplehorn and Merkl (1997: 739).

Heterophylus meszarosi Kaszab, 1977a: 123. Synonymy: Triplehorn and Merkl (1997: 739).

Loxostethus baracoae Garrido and Gutiérrez, 1995a: 7. Synonymy: Triplehorn and Merkl (1997: 739).

Loxostethus guadeloupensis (Kaszab, 1977)LAN (Guadeloupe)

Heterophylus guadeloupensis Kaszab, 1977a: 122.

Loxostethus oblongus Triplehorn and Merkl, 1997 DOM

Loxostethus oblongus Triplehorn and Merkl, 1997: 737.

Loxostethus unicolor Triplehorn, 1962 HAI PRI

Loxostethus unicolor Triplehorn, 1962: 506.

Heterophylus ruficornis Kaszab, 1981: 80. Synonymy: Triplehorn and Merkl (1997: 737).

Genus Mophis Champion, 1886 [M]

Mophis Champion, 1886: 168. Type species: Mophis marginicollis Champion, 1886, subsequent designation (Gebien 1940: 1061).

Mophis affinis Champion, 1886MEX (GE OA VE)

Mophis affinis Champion, 1886: 169.

Mophis aterrimus Champion, 1886: 169. Synonymy: Champion (1893a: 536).

Mophis cynaeoides (Champion, 1886)MEX (DU FD VE)

Sitophagus cynaeoides Champion, 1886: 162.

Mophis marginicollis Champion, 1886 GUA

Mophis marginicollis Champion, 1886: 169.

Genus Phayllus Champion, 1886 [M]

Phayllus Champion, 1886: 167. Type species: Phayllus minutus Champion, 1886, monotypy.

Phayllus minutus Champion, 1886MEX (VE) GUABELNICPAN / SA

Phayllus minutus Champion, 1886: 167.

Genus Saptine Champion, 1886 [F]

Saptine Champion, 1886: 180. Type species: Saptine ovata Champion, 1886, monotypy.

Saptine ovata Champion, 1886MEX (VE)

Saptine ovata Champion, 1886: 181.

Genus Sitophagus Mulsant, 1854 [M]

Sitophagus Mulsant, 1854: 264. Type species: Sitophagus solieri Mulsant, 1854 (=Uloma hololeptoides Laporte, 1840), monotypy.

Sitophagus alveolatus Doyen, 1984USA (AZ)

Sitophagus alveolatus Doyen, 1984a: 779.

Sitophagus dilatifrons Champion, 1886 GUA PAN

Sitophagus dilatifrons Champion, 1886: 162.

Sitophagus fuliginosus Champion, 1886 GUA

Sitophagus fuliginosus Champion, 1886: 161.

Sitophagus hololeptoides (Laporte, 1840)USA (AZ CA FL TX) MEX (DU PU VE YU) GUABELNICCRIPAN / BAHCUBPRILAN / SA

Uloma hololeptoides Laporte, 1840: 220.

Sitophagus solieri Mulsant, 1854: 265. Synonymy: Champion (1886: 161).

Adelina farinaria Wollaston, 1858: 414. Synonymy (withS. solieri Mulsant): Bates (1872: 99).

Sitophagus castaneus Reitter, 1877: 9. Synonymy: Champion (1886: 161).

Sitophagus laticollis Kulzer, 1961MEX (OA)

Sitophagus laticollis Kulzer, 1961b: 540.

Sitophagus uniformis Doyen, 1990MEX (GE JA OA PU)

Sitophagus uniformis Doyen, 1990: 250.

Subtribe Diaperina Latreille, 1802

Diaperialae Latreille, 1802: 161. Type genus: Diaperis Geoffroy, 1762.

Pentaphyllaires Mulsant, 1854: 196. Type genus: Pentaphyllus Dejean, 1821.

Platydeminae Reitter, 1917: 61. Type genus: Platydema Laporte and Brullé, 1831.

Genus Ceropria Laporte and Brullé, 1831 [F]

Ceropria Laporte and Brullé, 1831: 332, 396. Type species: Helops indutus Wiedemann, 1819, subsequent designation (Gebien 1940: 422).

Epilampus Dejean, 1834: 198. Unnecessary replacement name for Ceropria Laporte and Brullé, 1831 (see Bousquet and Bouchard 2013: 52).

Ceropria induta (Wiedemann, 1819)USA (FL) – Adventive

Helops indutus Wiedemann, 1819: 164.

Genus Cosmonota Blanchard, 1842 [F]

Cosmonota Blanchard, 1842: pl. 14. Type species: Cosmonota angustata Blanchard, 1842, subsequent designation (Gebien 1940: 417).

Cosmonota nigripes Chevrolat, 1877MEX (VE) GUABELNIC

Cosmonota nigripes Chevrolat, 1877b: 173.

Cosmonota pubescens Champion, 1886 NIC CRI PAN

Cosmonota pubescens Champion, 1886: 210.

Cosmonota silphoides (Laporte and Brullé, 1831)MEX (JA VE) GUABELNICPAN / SA

Platydema silphoides Laporte and Brullé, 1831: 369.

Platydema agile Chevrolat, 1877b: 178. Synonymy: Gebien (1940: 413).

Genus Diaperis Geoffroy, 1762 [F]

Diaperis Geoffroy, 1762: 337. Type species: Chrysomela boleti Linnaeus, 1758, subsequent designation (Latreille 1810: 429).

Allophasia Pascoe, 1871: 351. Type species: Allophasia fryi Pascoe, 1871, monotypy. Synonymy: Triplehorn and Brendell (1985: 14).

Diaperis californica Blaisdell, 1929USA (CA OR)

Diaperis californica Blaisdell, 1929c: 60.

Diaperis maculata Olivier, 1791 [Fig. 37] CAN (MB NB NS ON PE QC SK) USA (AL AR CT DC DE FL GA IA IL IN KS KY LA MA MD ME MI MN MO MS NC ND NE NH NJ NY OH OK PA RI SC SD TN TX VA WI WV WY) MEX (VE) GUACRIPAN / BAHCUBCAYJAMDOMPRILAN

Figure 37. 

Diaperis maculata Olivier, 1791. Scale bar = 1 mm.

Diaperis maculata Olivier, 1791: 273.

Diaperis hydactina Fabricius, 1798: 17868. Synonymy: Latreille (1804: 307, as D. hydni).

Diaperis suturalis Chevrolat, 1877a: 170. Synonymy: Champion (1886: 174).

Diaperis maculata var. floridana Blatchley, 1912: 332. Synonymy: Peck and Thomas (1998: 106).

Diaperis nigronotata Pic, 1926USA (AL AR FL GA IA IN KS LA MD MN MO MS OH OK PA SC TN TX WI WV)

Diaperis rufipes var. nigronotata Pic, 1926: 22.

Diaperis rufipes var. bicoloriceps Pic, 1926: 22. Synonymy: Triplehorn (1965: 373).

Diaperis rufipes Horn, 1870USA (AZ CA NM) MEX (BS)

Diaperis rufipes Horn, 1870: 379.

Genus Lelegeis Champion, 1886 [M]

Lelegeis Champion, 1886: 209. Type species: Lelegeis aeneipennis Champion, 1886, monotypy.

Lelegeis aeneipennis Champion, 1886MEX (VE)

Lelegeis aeneipennis Champion, 1886: 210.

Lelegeis apicalis (Laporte and Brullé, 1831) CUB PRI

Platydema apicalis Laporte and Brullé, 1831: 359.

Lelegeis hispaniolae Triplehorn, 1962 HAI DOM

Lelegeis hispaniolae Triplehorn, 1962: 503.

Lelegeis nigrifrons (Chevrolat, 1878)MEX (VE) PAN / SA

Platydema nigrifrons Chevrolat, 1878g: cxlviii.

Genus Liodema Horn, 1870 [F]

Liodema Horn, 1870: 385. Type species: Platydema laevis Haldeman, 1848, monotypy.

Liodema connexa Bates, 1873MEX (VE) GUAPAN / SA

Liodema connexum Bates, 1873c: 236.

Platydema nigro-fasciatum Chevrolat, 1878b: 215. Synonymy: Champion (1886: 207).

Liodema explanata Triplehorn, 1998 CRI PAN

Liodema explanatum Triplehorn, 1998: 325.

Liodema laevis (Haldeman, 1848)USA (FL GA MS NC SC TX) MEX (QU TA VE) GUACRIPAN

Platydema laevis Haldeman, 1848: 101.

Liodema maculata (Fabricius, 1801)MEX (NA PU SL TA VE YU) GUASALHONNICCRIPAN / SA

Mycetophagus maculatus Fabricius, 1801b: 566.

Platydema 4-notata Laporte and Brullé, 1831: 380. Synonymy (in doubt with L. kirschi Bates): Champion (1886: 205).

Liodema obydense Bates, 1873c: 235. Synonymy: Triplehorn (1998: 327).

Liodema kirschi Bates, 1873c: 235. Synonymy: Gebien (1940: 417).

Liodema fulvum Bates, 1873c: 236. Synonymy (in doubt with L. kirschi Bates): Champion (1886: 205).

Liodema horni Bates, 1873c: 236. Synonymy: Triplehorn (1998: 328).

Scaphidema tergocinctum Chevrolat, 1877b: 178. Synonymy (with L. kirshi Bates): Chevrolat (1878d: 243).

Scaphidema proximum Chevrolat, 1877b: 178. Synonymy (with L. obydense Bates): Chevrolat (1878d: 243).

Liodema inscriptum Chevrolat, 1878b: 222. Synonymy (with L. kirshi Bates): Champion (1886: 205).

Liodema serricornis Bates, 1873MEX (TA VE) GUASALHONNICCRIPAN / DOM / SA

Liodema serricorne Bates, 1873c: 236.

Platydema cruciatum Chevrolat, 1877c: 182. Synonymy: Triplehorn (1998: 328).

Platydema hamatiferum Chevrolat, 1878f: c. Synonymy: Triplehorn (1998: 328).

Platydema ramulosum Chevrolat, 1878f: c. Synonymy: Triplehorn (1998: 328).

Liodema zimmermani Champion, 1886: 206. Synonymy: Triplehorn (1998: 328).

Liodema flavo-variegatum Champion, 1886: 208. Synonymy: Triplehorn (1998: 328).

Genus Neomida Latreille, 1829 [F]

Neomida Latreille, 1829a: 29. Type species: Ips haemorrhoidalis Fabricius, 1787, monotypy.

Oplocephala Laporte and Brullé, 1831: 338. Type species: Ips haemorrhoidalis Fabricius, 1787, subsequent designation (Motschulsky 1845a: 80). Synonymy: Dejean (1834: 197).

Arrhenoplita Kirby, 1837: 235. Type species: Ips haemorrhoidalis Fabricius, 1787, original designation. Synonymy: Duponchel and Chevrolat (1841: 157).

Hoplocephala Agassiz, 1846: 185. Unjustified emendation of Oplocephala Laporte and Brullé, 1831, not in prevailing usage.

Evoplus LeConte, 1866b: 128. Type species: Evoplus ferruginea LeConte, 1866, monotypy. Synonymy (with Arrhenoplita Kirby): Champion (1886: 175).

Neomida acera Triplehorn, 1994 CRI PAN

Neomida acera Triplehorn, 1994a: 426.

Neomida aeneipennis Triplehorn, 1965MEX (CI HI JA OA PU QR QU SL TA VE YU) GUABELSALHONNICCRI

Neomida aeneipennis Triplehorn, 1965: 382.

Neomida armata (Laporte and Brullé, 1831)CUB / SA

Oplocephala armata Laporte and Brullé, 1831: 345.

Neomida bicornis (Fabricius, 1777)CAN (MB NB NS ON PE QC) USA (AL AR CT DC DE FL GA IA IL IN KS KY LA MA MD ME MI MN MO MS NC NE NH NJ NY OH OK PA RI SC SD TN TX VA WI WV) / BAHBERCUBCAYJAM

Hispa bicornis Fabricius, 1777: 215.

Hispa cornigera Fabricius, 1781: 82. Synonymy:Triplehorn (1965: 377).

Diaperis viridipennis Fabricius, 1801b: 586. Synonymy:Triplehorn (1965: 377).

Blaps metallica Palisot de Beauvois, 1817: 140 [junior primary homonym of Blaps metallica Fabricius, 1801]. Synonymy: Horn (1885a: 88).

Oplocephala virescens Laporte and Brullé, 1831: 341. Synonymy: Melsheimer (1853: 137).

Oplocephala capra Laporte and Brullé, 1831: 345. Synonymy (in doubt): Triplehorn (2006: 332).

Oplocephala gracilis Motschulsky, 1873: 467. Synonymy: Horn (1874b: 98).

Neomida castanea (Bates, 1873)MEX (VE) NICCRIPAN / SA

Hoplocephala castanea Bates, 1873b: 204.

Hoplocephala oblonga Chevrolat, 1878f: xcvii. Synonymy: Triplehorn (2006: 325).

Neomida cioides (Champion, 1886)MEX (CI QU VE) CRIPAN / DOMLAN / SA

Arrhenoplita cioides Champion, 1886: 180.

Neomida clavicornis (Champion, 1886)MEX (VE) CI

Arrhenoplita clavicornis Champion, 1886: 176.

Neomida deltocera Triplehorn, 1994CRIPANLAN / SA

Neomida deltocera Triplehorn, 1994a: 423.

Neomida distans (Champion, 1886)MEX (VE) CRIPAN / SA

Arrhenoplita distans Champion, 1886: 178.

Neomida divergicornis Triplehorn, 1994MEX (CI VE)

Neomida divergicornis Triplehorn, 1994a: 420.

Neomida dolichocera Triplehorn, 1994 CRI

Neomida dolichocera Triplehorn, 1994a: 417.

Neomida ferruginea (LeConte, 1866)USA (AL FL GA LA TX) MEX (CL HI NA PU SI SL TA TB VE) GUABELCRIPAN / CUBCAYJAMHAIDOM / SA

Evoplus ferruginea LeConte, 1866b: 128.

Oplocephala castanea Motschulsky, 1873: 467. Synonymy (in doubt): Horn (1874b: 98)

Neomida heterocera Triplehorn, 1994 CRI PAN

Neomida heterocera Triplehorn, 1994a: 421.

Neomida hoffmanseggii (Laporte and Brullé, 1831)MEX (CL) CRIPAN / SA

Oplocephala hoffmanseggii Laporte and Brullé, 1831: 346.

Neomida inermis (Champion, 1886)GUACRIPAN / LAN (Guadeloupe)

Arrhenoplita inermis Champion, 1886: 179.

Neomida lateralis (Bates, 1873)MEX (PU VE) CRIPAN / SA

Hoplocephala lateralis Bates, 1873b: 204.

Hoplocephala dytiscoïdes Chevrolat, 1877a: 170. Synonymy: Triplehorn (2006: 319).

Hoplocephala lutea Chevrolat, 1878f: xcvii. Synonymy: Triplehorn (2006: 333).

Neomida lawrencei Triplehorn, 1994MEX (HI OA) BELCRIPAN

Neomida lawrencei Triplehorn, 1994a: 419.

Neomida lecontei (Bates, 1873)MEX (CI VE) BELHONPAN / JAMDOMPRILAN / SA

Evoplus lecontii Bates, 1873c: 233.

Uloma guadeloupensis Marcuzzi, 1971: 110. Synonymy: Soldati and Touroult (2014: 102), confirmed by Ivie and Hart (2017a: 116).

Neomida nigricornis (Champion, 1886) GUA BEL CRI PAN

Arrhenoplita nigricornis Champion, 1886: 179.

Neomida obsoleta (Champion, 1886)MEX (OA VE) BELCRIPAN

Arrhenoplita obsoleta Champion, 1886: 178.

Neomida occidentalis (Champion, 1893)MEX (BS GE JA PU SI) HONCRI

Arrhenoplita occidentalis Champion, 1893a: 537.

Neomida myllocnema Triplehorn, 1965: 386. Synonymy: Triplehorn (2006: 315).

Neomida paurocera Triplehorn, 1994 SAL HON

Neomida paurocera Triplehorn, 1994a: 424.

Neomida pentaphylloides (Champion, 1886) GUA

Arrhenoplita pentaphylloides Champion, 1886: 180.

Neomida picea (Laporte and Brullé, 1831)GUABELCRIPAN / SA

Oplocephala picea Laporte and Brullé, 1831: 344.

Hoplocephala testaceipes Pic, 1926: 28. Synonymy: Triplehorn (2006: 313).

Neomida pogonocera Triplehorn, 1994PAN / SA

Neomida pogonocera Triplehorn, 1994a: 422.

Neomida punctatissima (Champion, 1893)MEX (GE JA)

Arrhenoplita punctatissima Champion, 1893a: 537.

Neomida suilla (Champion, 1896)LAN / SA

Arrhenoplita suilla Champion, 1896: 11.

Neomida telecera Triplehorn, 2006 CRI

Neomida telecera Triplehorn, 2006: 328.

Genus Paniasis Champion, 1886 [M]

Paniasis Champion, 1886: 208. Type species: Paniasis dilatipes Champion, 1886, monotypy.

Pseudapsida Kulzer, 1961a: 219. Type species: Pseudapsida brasiliensis Kulzer, 1961, original designation. Synonymy: Ferrer and Ødegaard (2005: 637).

Paniasis dilatipes Champion, 1886MEX (VE) / SA

Paniasis dilatipes Champion, 1886: 209.

Paniasis kulzeri Ferrer and Ødegaard, 2005 PAN

Paniasis kulzeri Ferrer and Ødegaard, 2005: 637.

Genus Pentaphyllus Dejean, 1821 [M]

Pentaphyllus Dejean, 1821: 68. Type species: Mycetophagus testaceus Hellwig, 1792, monotypy.

Iphicorynus Jacquelin du Val, 1861: 299. Type species: Pentaphyllus melanophthalmus Mulsant, 1854 (= Nitidula chrysomeloides Rossi, 1792), monotypy. Synonymy: Gemminger [in Gemminger and Harold] (1870: 1956).

Pentaphyllus californicus Horn, 1870USA (CA)

Pentaphyllus californicus Horn, 1870: 387.

Pentaphyllus pallidus LeConte, 1866CAN (ON QC) USA (CT GA IL IN KY MD MI NJ NY OH PA SC WI)

Pentaphyllus pallidus LeConte, 1866b: 126.

Pentaphyllus americanus Motschulsky, 1873: 482. Synonymy: Horn (1874b: 98).

Pentaphyllus testaceus (Hellwig, 1792)CAN (ON) – Adventive

Mycetophagus testaceus Hellwig, 1792: 400.

Genus Platydema Laporte and Brullé, 1831 [F]

Platydema Laporte and Brullé, 1831: 350. Type species: Diaperis violacea Fabricius, 1790, subsequent designation (Westwood 1838: 32).

Typhobia Pascoe, 1869: 279. Type species: Typhobia fuliginea Pascoe, 1869, monotypy. Synonymy: Champion (1886: 181).

Histeropsis Chevrolat, 1878b: 221. Type species: Platydema americanum Laporte and Brullé, 1831, subsequent designation (Löbl et al. 2008b: 42). Synonymy: Champion (1886: 181).

Platydema americana Laporte and Brullé, 1831CAN (AB BC MB NB NS ON QC SK) USA (AZ CA CO CT IA ID IL KS MD ME MI MN MO MT NC NE NH NJ NM NV NY OH OR PA SC SD TX VA WA WI WY)

Platydema americana Laporte and Brullé, 1831: 358.

Platydema polita Laporte and Brullé, 1831: 361. Synonymy: Horn (1870: 384).

Platydema angulata Chevrolat, 1877 MEX

Platydema angulatum Chevrolat, 1877d: 186.

Platydema antennata Laporte and Brullé, 1831 BAH CUB CAY HAI

Platydema antennata Laporte and Brullé, 1831: 366.

Platydema apicenotata Champion, 1896 LAN

Platydema apicenotatum Champion, 1896: 13.

Platydema basicornis Chevrolat, 1877 CUB

Platydema basicorne Chevrolat, 1877b: 178.

Platydema bimaculata Champion, 1886MEX (VE) GUABELNICCRIPAN / SA

Platydema bimaculatum Champion, 1886: 193.

Platydema biplagiata Champion, 1886MEX (VE) GUANICPAN

Platydema biplagiatum Champion, 1886: 201.

Platydema bisignata Chevrolat, 1877MEXGUA / SA

Platydema bisignatum Chevrolat, 1877c: 181.

Platydema brevis Champion, 1886MEX (VE) GUAPAN

Platydema breve Champion, 1886: 200.

Platydema concolor Champion, 1893 NIC PAN

Platydema unicolor Champion, 1886: 203 [junior primary homonym of Platydema unicolor Chevrolat, 1878].

Platydema concolor Champion, 1893a: 539. Replacement name for Platydema unicolor Champion, 1886.

Platydema cordovensis Champion, 1886MEX (VE) GUA

Platydema cordovense Champion, 1886: 203.

Platydema cyanea Laporte and Brullé, 1831 “Amérique septentrionale”69

Platydema cyanea Laporte and Brullé, 1831: 392.

Platydema cyanescens Laporte and Brullé, 1831USA (AL FL GA IN LA MS NC OH SC TN TX)

Platydema cyanescens Laporte and Brullé, 1831: 356.

Platydema dichrocera Triplehorn, 1962 CUB

Platydema dichrocerum Triplehorn, 1962: 502.

Platydema dimidiata Chevrolat, 1878MEX (VE) GUABELHON

Platydema dimidiatum Chevrolat, 1878a: 194.

Platydema diophthalma Laporte and Brullé, 1831MEX (DU VE YU) GUABELHONNICCRIPAN / CUB

Platydema diophthalma Laporte and Brullé, 1831: 383.

Platydema luna Chevrolat, 1877d: 186. Synonymy: Champion (1886: 193).

Platydema elegans Chevrolat, 1878MEX (PU VE)

Platydema elegans Chevrolat, 1878a: 195.

Platydema elliptica (Fabricius, 1798)CAN (ON) USA (AL AR CO CT DC DE FL GA IA IL IN KS KY LA MD MI MO MS NC NJ NY OH OK PA SC TN TX UT VA WI)

Tenebrio ellipticus Fabricius, 1798: 49.

Platydema erotyloides Chevrolat, 1878PAN / SANew North American record

Platydema ornatum Chevrolat, 1878b: 209 [junior primary homonym of Platydema ornatum Chevrolat, 1877].

Platydema erotyloides Chevrolat, 1878d: 243. Replacement name for Platydema ornatum Chevrolat, 1878.

Platydema erythrocera Laporte and Brullé, 1831USA (AL AR DC FL GA IL IN KS LA MD MO MS NC NY OH OK SC TN TX VA WV) MEX (CI HI) BEL / BAH

Platydema erythrocera Laporte and Brullé, 1831: 355.

Neomida flavicornis Motschulsky, 1873: 479. Synonymy: Horn (1874b: 98).

Platydema hondurense Champion, 1886: 186. Synonymy: Triplehorn (1994b: 247).

Platydema excavata (Say, 1824)CAN (ON QC) USA (AL AR AZ CT DC DE FL GA IA IL IN KS KY LA MA MD MI MN MO MS NC NE NH NJ NM NY OH OK PA RI SC SD TN TX VA WI WV) MEX (JA PU VE YU) GUABELHONNICCRIPAN / BAHCUBCAYHAIDOMJAMPRILAN / SA

Diaperis excavata Say, 1824a: 267.

Platydema tuberculata Laporte and Brullé, 1831: 352. Synonymy: Champion (1886: 184).

Platydema nigritum Motschulsky, 1873: 470. Synonymy: Horn (1874b: 98).

Platydema fraternum Chevrolat, 1878b: 210. Synonymy: Champion (1886: 184).

Platydema parvulum Casey, 1884: 50. Synonymy: Casey (1885: 195).

Platydema fasciatocollis Chevrolat, 1878 MEX

Platydema fasciato-colle Chevrolat, 1878a: 194.

Platydema fasciata (Fabricius, 1801)MEX (PU VE) GUABEL / SA

Mycetophagus fasciatus Fabricius, 1801b: 567.

Platydema ferruginea Chevrolat, 1877MEX (CI VE YU) GUAPAN

Platydema ferrugineum Chevrolat, 1877d: 186.

Platydema bi-impressum Chevrolat, 1878b: 214. Synonymy: Champion (1886: 190).

Platydema flavipes (Fabricius, 1801)USA (AL AR DC DE FL GA ON KS KY LA MA MD MS NC ND NH NJ NY OH PA SC TN TX VA)

Mycetophagus flavipes Fabricius, 1801b: 567.

Platydema basalis Haldeman, 1848: 101. Synonymy: Horn (1870: 382).

Platydema flexuosa Chevrolat, 1877 CUB

Platydema flexuosum Chevrolat, 1877b: 178.

Platydema fuliginosa Laporte and Brullé, 1831 MEX

Platydema fuliginosa Laporte and Brullé, 1831: 374.

Platydema guatemalensis Champion, 1886MEX (CI VE) GUASALCRIPAN /LAN / SA

Platydema guatemalense Champion, 1886: 197.

Platydema hoegei Champion, 1886MEX (CI GE PU VE)

Platydema högei Champion, 1886: 195.

Platydema immaculata Champion, 1886 PAN

Platydema immaculatum Champion, 1886: 192.

Platydema inquilina Linell, 1899USA (AZ)

Platydema inquilinum Linell, 1899: 183.

Platydema laevipes Haldeman, 1848USA (AL AR DC FL GA IA IN KS LA MA MD MO MS NC NJ NY OH PA SC TX VA WI)

Platydema laevipes Haldeman, 1848: 101.

Platydema crenatum LeConte, 1878a: 422. Synonymy: Triplehorn (1965: 407).

Platydema lucens Champion, 1886MEX (VE)

Platydema lucens Champion, 1886: 202.

Platydema maculipennis Champion, 1886MEX (VE) GUA

Platydema maculipenne Champion, 1886: 201.

Platydema melancholica Champion, 1886 GUA

Platydema melancholicum Champion, 1886: 190.

Platydema mexicana Champion, 1886USA (AZ NM) MEX (CH DU VE)

Platydema mexicanum Champion, 1886: 187.

Platydema micans Zimmerman, 1870USA (AL AR DC FL GA IN KS LA MD MS NC SC TN TX VA) MEX (JA) / BAHTURCUBCAYJAMHAI / SA

Platydema micans Zimmerman [in Horn], 1870: 383.

Platydema monilicornis Chevrolat, 1877 MEX

Platydema monilicorne Chevrolat, 1877d: 186.

Platydema neglecta Triplehorn, 1965CAN (BC) USA (CA ID MT NV OR UT WA)

Platydema neglectum Triplehorn, 1965: 404.

Platydema nicaraguensis Champion, 1886 NIC

Platydema nicaraguense Champion, 1886: 192.

Platydema nigrata (Motschulsky, 1873)USA (AL AZ CA FL GA IN KS LA MS NC NM SC TX) MEX (MO SI VE YU) GUABELHONNICCRI / BAHCUB

Neomida nigrata Motschulsky, 1873: 478.

Neomida texana Motschulsky, 1873: 478. Synonymy (in doubt with P. janus sensu Horn, 1870 = P. nigrita): Horn (1874b: 98).

Platydema nigromaculata Champion, 1886 BEL NIC PAN

Platydema nigromaculatum Champion, 1886: 199.

Platydema nitida (Chevrolat, 1877)MEX (YU)

Scaphidema nitidum Chevrolat, 1877a: 170.

Platydema oculata Champion, 1886MEX (VE)

Platydema oculatum Champion, 1886: 191.

Platydema oregonensis LeConte, 1857CAN (BC) USA (CA ID OR WA)

Platydema oregonense LeConte, 1857: 51.

Platydema ornata Chevrolat, 1877MEX (VE)

Platydema ornatum Chevrolat, 1877d: 186.

Platydema panamensis Champion, 1886 PAN

Platydema panamense Champion, 1886: 198.

Platydema picicornis (Fabricius, 1792)CUBHAIPRI / SA

Mycetophagus picicornis Fabricius, 1792b: 498.

Platydema picilabrum Melsheimer, 1846USA (AL AR FL GA IL IN KS KY LA MA MD MI MO MS NC NJ NY OH PA SC TN TX VA WI WV)

Platydema picilabrum Melsheimer, 1846: 62.

Platydema pilifera Champion, 1896 LAN

Platydema piliferum Champion, 1896: 12.

Platydema pretiosa Champion, 1886 BEL

Platydema pretiosum Champion, 1886: 197.

Platydema punctatostriata Chevrolat, 1877 CUB

Platydema punctatostriatum Chevrolat, 1877b: 178.

Platydema quadrimaculata Laporte and Brullé, 1831USA (PA)70

Platydema4-maculata Laporte and Brullé, 1831: 383.

Platydema quindecimmaculata (Chevrolat, 1878)GUANICPAN / SA

Platydema 15-maculatum Chevrolat, 1878g: cxlix.

Platydema rotundata Chevrolat, 1877MEX (GE JA MO PU VE YU) GUACRI

Platydema rotundatum Chevrolat, 1877d: 186.

Platydema ruficollis Laporte and Brullé, 1831USA (AR FL GA IA IL KY MD MS NC NJ OK SC TX VA WI)

Platydema ruficollis Laporte and Brullé, 1831: 375.

Neomida sanguinicollis Melsheimer, 1846: 61. Synonymy: Haldeman (1848: 102).

Platydema ruficornis (Sturm, 1826)CAN (ON QC) USA (AL AR CO CT DC DE FL GA IA IL IN KS KY LA MA MD MI MN MO MS NC NE NJ NY OH OK PA SC TN TX VA WI WV) / BAH

Diaperis ruficornis Sturm, 1826: 69.

Platydema rufiventris Laporte and Brullé, 1831: 378. Synonymy: Melsheimer (1853: 138).

Platydema pallens Laporte and Brullé, 1831: 377. Synonymy (in doubt): Horn (1870: 382)71.

Neomida rufa Melsheimer, 1846: 62. Synonymy: Horn (1870: 382).

Platydema analis Haldeman, 1848: 101. Synonymy: Horn (1870: 382).

Platydema opaculum Casey, 1884: 51. Synonymy (with P. ruficorne var. anale Haldeman): Horn (1885b: 111).

Platydema rugiceps Champion, 1886MEX (VE) GUANICPAN

Platydema rugiceps Champion, 1886: 191.

Platydema sexmaculata Chevrolat, 1878 MEX

Platydema sexmaculatum Chevrolat, 1878a: 194.

Platydema sexnotata Chevrolat, 1878MEX (JA VE) NICCRI

Platydema sexnotatum Chevrolat, 1878a: 194.

Platydema sobrina Chevrolat, 1877MEX (VE) GUABELNICCRIPAN / SA

Neomida discolor Motschulsky, 1873: 477 [nomen dubium].

Platydema sobrinum Chevrolat, 1877d: 186. Synonymy (in doubt): Champion (1886: 189).

Platydema subcostata Laporte and Brullé, 1831CAN (ON QC) USA (AL AR CT DC DE FL GA IA IL IN KS KY LA MA MD MI MN MO MS NC NH NJ NY OH PA RI SC TN TX VA WI WV)

Platydema subcostata Laporte and Brullé, 1831: 362.

Platydema clypeatus Haldeman, 1848: 102. Synonymy: Horn (1870: 384).

Platydema oblongulum Motschulsky, 1873: 470. Synonymy: Horn (1874b: 98).

Platydema subquadrata (Motschulsky, 1873)72 “Amérique centrale”

Neomida subquadrata Motschulsky, 1873: 477.

Platydema pernigrum Casey, 1884: 49. Synonymy: Champion (1886: 188).

Platydema submaculata (Chevrolat, 1878)MEX (PU VE YU) BEL

Platydema submaculatum Chevrolat, 1878f: xcix.

Platydema teleops Triplehorn, 1965CAN (NB NS ON QC) USA (CT DC IA IL IN KS MA MD ME MI MN MO NC NE NH NJ NY OH OK SC TN TX VA WI WV)

Platydema teleops Triplehorn, 1965: 399.

Platydema tibialis (Chevrolat, 1878)NICPAN / SA

Platydema tibiale Chevrolat, 1878g: cxlviii.

Platydema transversa Laporte and Brullé, 1831MEX (OA VE) GUABELPAN / SA

Platydema transversa Laporte and Brullé, 1831: 381.

Platydema tricolor Champion, 1886 GUA

Platydema tricolor Champion, 1886: 200.

Platydema undata Chevrolat, 1878MEX (CI PU SL VE) GUABELSALNICCRIPAN / SA

Neomida picta Motschulsky, 1873: 480 [junior secondary homonym of Platydema pictum (Ménétriés, 1832)].

Platydema undatum Chevrolat, 1878a: 194. Synonymy: Champion (1886: 185).

Platydema rodriguezi Champion, 1886: 185. Synonymy: Triplehorn (1994b: 249).

Platydema venusta Champion, 1886 NIC PAN

Platydema venustum Champion, 1886: 204.

Platydema ventralis Chevrolat, 1877 MEX

Platydema ventrale Chevrolat, 1877d: 186.

Platydema versicolor Chevrolat, 1878MEX (PU VE)

Platydema versicolor Chevrolat, 1878a: 195.

Platydema viriditincta Champion, 1886MEX (OA)

Platydema viriditinctum Champion, 1886: 186.

Platydema wandae Triplehorn, 1965USA (AZ NM)

Platydema wandae Triplehorn, 1965: 427.

Platydema woldai Triplehorn and Philips, 1998USAi (FL) MEXGUASALHONPAN

Platydema woldai Triplehorn and Philips [in Philips et al.], 1998: 291.

Genus Stenoscapha Bates, 1873 [F]

Stenoscapha Bates, 1873c: 237. Type species: Stenoscapha tibialis Bates, 1873, monotypy.

Stenoscapha jalapensis Champion, 1886MEX (VE)

Stenoscapha jalapensis Champion, 1886: 208.

Genus Ulomoides Blackburn, 1888 [M]

Ulomoides Blackburn, 1888: 274. Type species: Ulomoides humeralis Blackburn, 1888, monotypy.

Palembus Casey, 1891: 65. Type species: Palembus ocularis Casey, 1891, monotypy. Synonymy: Doyen et al. (1990: 237).

Martianus Fairmaire, 1893: 540. Type species: Martianus castaneus Fairmaire, 1893 (= Palembus ocularis Casey, 1891), original designation. Synonymy (with Palembus Casey): Halstead (1974: 241).

Tenebriomimus Kolbe, 1901: 342. Type species: Tenebriomimus adansoniarum Kolbe, 1901 (= Palembus ocularis Casey, 1891), monotypy. Synonymy (with Martianus Fairmaire): Gebien (1922: 268).

Ulomoides ocularis (Casey, 1891)USA (FL) / BAHCUBCAYJAMDOMPRILAN – Adventive

Palembus ocularis Casey, 1891: 65.

Martianus castaneus Fairmaire, 1893: 541. Synonymy: Halstead (1974: 242).

Tenebriomimus adansoniarum Kolbe, 1901: 342. Synonymy (with M. castaneus Fairmaire): Gebien (1922: 268).

Tribe Gnathidiini Gebien, 1921

Gnathidiini Gebien, 1921: 41. Type genus: Gnathidium Gebien, 1921.

Subtribe Anopidiina Jeannel and Paulian, 1945

Anopidiini Jeannel and Paulian, 1945: 62. Type genus: Anopidium Jeannel and Paulian, 1945.

Genus Caecophloeus Dajoz, 1972 [M]

Caecophloeus Dajoz, 1972: 278. Type species: Caecophloeus franzi Dajoz, 1972, original designation.

Caecophloeus darlingtoni Dajoz, 1975 HAI

Caecophloeus darlingtoni Dajoz, 1975: 115.

Caecophloeus distinctus Dajoz, 1975MEX (CI)

Caecophloeus distinctus Dajoz, 1975: 117.

Caecophloeus franzi Dajoz, 1972 JAM

Caecophloeus franzi Dajoz, 1972: 280.

Caecophloeus ineditus Dajoz, 1975MEX (CI)

Caecophloeus ineditus Dajoz, 1975: 119.

Caecophloeus pubescens Dajoz, 1975 PAN

Caecophloeus pubescens Dajoz, 1975: 117.

Genus Cryptozoon Schaufuss, 1882 [N]

Cryptozoon Schaufuss, 1882: 47. Type species: Cryptozoon civile Schaufuss, 1882, present designation.

Cryptozoon civile Schaufuss, 1882 PRI

Cryptozoon civile Schaufuss, 1882: 47.

Cryptozoon nitidicolle Schaufuss, 1882 PRI

Cryptozoon nitidicolle Schaufuss, 1882: 47.

Genus Menimopsis Champion, 1896 [F]

Menimopsis Champion, 1896: 16. Type species: Menimopsis excaecus Champion, 1896, monotypy.

Caecomenimopsis Kaszab, 1970: 198. Type species: Caecomenimopsis leleupi Kaszab, 1970, original designation. Synonymy: Peck (1990: 370).

Menimopsis excaeca Champion, 1896 LAN

Menimopsis excaecus Champion, 1896: 17.

Menimopsis franzi Kaszab, 1977 JAM

Menimopsis franzi Kaszab, 1977a: 122.

Menimopsis jamaicensis (Dajoz, 1975) JAM

Caecomenimopsis jamaicensis Dajoz, 1975: 121.73

Menimopsis jamaicensis Kaszab, 1977 JAM

Menimopsis jamaicensis Kaszab, 1977a: 121 [junior secondary homonym of Menimopsis jamaicensis (Dajoz, 1975)].74

Genus Neanopidium Dajoz, 1975 [N]

Neanopidium Dajoz, 1975: 93. Type species: Neanopidium mexicanum Dajoz, 1975, original designation.

Neanopidium affine Dajoz, 1975MEX (HI)

Neanopidium affinis Dajoz, 1975: 107.

Neanopidium convexum Dajoz, 1975MEX (VE)

Neanopidium convexum Dajoz, 1975: 104.

Neanopidium curticorne Dajoz, 1975MEX (CI)

Neanopidium curticornis Dajoz, 1975: 100.

Neanopidium dubium Dajoz, 1975MEX (HI)

Neanopidium dubium Dajoz, 1975: 108.

Neanopidium humerale Dajoz, 1975MEX (OA)

Neanopidium humeralis Dajoz, 1975: 105.

Neanopidium lawrencei Dajoz, 1975MEX (VE)

Neanopidium lawrencei Dajoz, 1975: 103.

Neanopidium mexicanum Dajoz, 1975MEX (OA)

Neanopidium mexicanum Dajoz, 1975: 96.

Neanopidium minutum Dajoz, 1975MEX (VE)

Neanopidium minutum Dajoz, 1975: 105.

Neanopidium newtoni Dajoz, 1975MEX (VE)

Neanopidium newtoni Dajoz, 1975: 102.

Neanopidium pubescens Dajoz, 1975MEX (VE)

Neanopidium pubescens Dajoz, 1975: 102.

Neanopidium punctatum Dajoz, 1975MEX (SL)

Neanopidium punctatum Dajoz, 1975: 106.

Neanopidium simile Dajoz, 1975MEX (HI)

Neanopidium similis Dajoz, 1975: 98.

Neanopidium testaceum Dajoz, 1975MEX (TA)

Neanopidium testaceum Dajoz, 1975: 103.

Genus Sphaerognathium Dajoz, 1975 [N]

Sphaerognathium Dajoz, 1975: 112. Type species: Sphaerognathium globosum Dajoz, 1975, original designation.

Sphaerognathium globosum Dajoz, 1975 HAI

Sphaerognathium globosum Dajoz, 1975: 113.

Genus Tyrtaeus Champion, 1913 [M]

Tyrtaeus Champion, 1913: 76. Type species: Tyrtaeus rufus Champion, 1913, original designation.

Tyrtaeus cribripennis Champion, 1913 PAN

Tyrtaeus cribripennis Champion, 1913: 77.

Tyrtaeus dobsoni Hinton, 1947USA (FL) – Adventive

Tyrtaeus dobsoni Hinton, 1947a: 852.

Tyrtaeus rufus Champion, 191375USA (FL) MEX (VE) GUACRIPAN / CUBCAYLAN / SA

Tyrtaeus rufus Champion, 1913: 77.

Tyrtaeus guadalupensis Dajoz, 1981: 227. Synonymy: Hopp and Ivie (2008: 429).

Tribe Hypophlaeini Billberg, 1820

Hypophlaeides Billberg, 1820a: 33. Type genus: Hypophlaeus Fabricius, 1790 (= Corticeus Piller and Mitterpacher, 1783).

Corticeini Boddy, 1965: 144. Type genus: Corticeus Piller and Mitterpacher, 1783.

Genus Corticeus Piller and Mitterpacher, 1783 [M]

Corticeus Piller and Mitterpacher, 1783: 87. Type species: Corticeus unicolor Piller and Mitterpacher, 1783, monotypy.

Hypophlaeus Fabricius, 1790: 222. Type species: Hypophlaeus castaneus Fabricius, 1790 (= Corticeus unicolor Piller and Mitterpacher, 1783), subsequent designation (Curtis 1832: pl. 430). Synonymy: Crotch (1870: 47).

Subgenus Corticeus Piller and Mitterpacher, 1783

Corticeus Piller and Mitterpacher, 1783: 87. Type species: Corticeus unicolor Piller and Mitterpacher, 1783, monotypy.

Corticeus coynei Triplehorn, 1970 HON NIC

Corticeus coynei Triplehorn [in Triplehorn and Moser], 1970: 47.

Corticeus crassicornis Champion, 1886 GUA

Corticeus crassicornis Champion, 1886: 173.

Corticeus longicornis Champion, 1886 MEX

Corticeus longicornis Champion, 1886: 172.

Corticeus mexicanus mexicanus Reitter, 1878MEX (VE) GUANICPAN / SA

Corticeus mexicanus Reitter, 1878: 191.

Corticeus cylindricus Reitter, 1878: 192 [junior primary homonym of Corticeus cylindricus Reitter, 1877]. Synonymy: Bremer and Triplehorn (1999: 57).

Corticeus erratus Reitter, 1894: 16. Replacement name for Corticeus cylindricus Reitter, 1878.

Hypophloeus meridanus Pic, 1914: 15. Synonymy: Bremer and Triplehorn (1999: 57).

Corticeus opaculus (LeConte, 1878)USA (AZ CA) MEXGUA

Hypophloeus opaculus LeConte, 1878a: 423.

Corticeus pallidipennis Champion, 1886MEX (VE) GUA

Corticeus pallidipennis Champion, 1886: 173.

Corticeus parallelus (Melsheimer, 1846)CAN (MB ON QC) USA (AL AR DC DE FL GA IL IN KY LA MA MD ME MI MN MO MS NC ND NE NH NJ NY OH PA SC TN TX VA WI WV)

Hypophloeus parallelus Melsheimer, 1846: 63.

Corticeus paulostriatus (Pic, 1945)USA (AR FL TN) / CUBHAIDOMPRI

Hypophlaeus paulostriatus Pic, 1945: 8.

Corticeus tensicollis Triplehorn, 1979: 46. Synonymy: Bremer and Triplehorn (1999: 58).

Corticeus praetermissus (Fall, 1926)CAN (AB BC MB NB NF NS NT ON QC SK YT) USA (AK AZ CA CO ID MA ME MI MN NE NH NM NV NY OR PA SD TX UT WA WI WV WY) MEX

Hypophloeus praetermissus Fall, 1926: 199.

Corticeus puncticollis Champion, 1886 GUA

Corticeus puncticollis Champion, 1886: 172.

Corticeus rosei Triplehorn, 1970USA (AZ) MEX (CH DU JA ME NL PU) HON

Corticeus rosei Triplehorn [in Triplehorn and Moser], 1970: 49.

Corticeus rufipes (Fabricius, 1801)MEX (OA TB VE) GUABELNICPAN / CUBPRILAN / SA

Hypophloeus rufipes Fabricius, 1801b: 558.

Corticeus sordidus Champion, 1913 GUA

Corticeus sordidus Champion, 1913: 162.

Corticeus strublei Blaisdell, 1934USA (AZ CA CO ID NM OR SD UT WA WY) [MEX]

Corticeus strublei Blaisdell, 1934a: 188.

Corticeus subopacus (Wallis, 1933)CAN (AB BC) USA (AK CO ID ME MI MT NC NH NY PA WA WI WV WY)

Hypophloeus subopacus Wallis, 1933: 247.

Corticeus substriatus (LeConte, 1878)CAN (BC) USA (AZ CA CO ID MT NM NV OR SD UT WA) MEX (BC)

Hypophloeus substriatus LeConte, 1878a: 423.

Corticeus tenuis (LeConte, 1878)CAN (AB BC NB NS ON QC) USA (AZ CA ID MA ME MI MN MT NH NY OR PA VA WA WI WV WY)

Hypophlocus [sic!] tenuis LeConte, 1878a: 424.

Hypophloeus minor Wallis, 1933: 248. Synonymy: Triplehorn (1990: 294).

Hypophloeus occidentalis Wallis, 1933: 249. Synonymy: Triplehorn (1990: 294).

Subgenus Pogonophloeus Bremer, 1998

Pogonophloeus Bremer, 1998: 9. Type species: Hypophloeus thoracicus Melsheimer, 1846, original designation.

Corticeus cavus (LeConte, 1866)USA (AL DC IA KS KY MD MO MS NC OH OK PA TX VA WV WI)

Hypophloeus cavus LeConte, 1866b: 129.

Corticeus hatchi Boddy, 1957USA (AZ CA CO NM OR)

Corticeus hatchi Boddy, 1957: 197.

Corticeus thoracicus (Melsheimer, 1846)CAN (ON) USA (AL AR DC DE FL GA IN KY LA MD MN MO MS NC NJ NY OH OK PA SC TN TX VA WI WV) / BAH

Hypophloeus thoracicus Melsheimer, 1846: 63.

Hypophloeus piliger LeConte, 1878a: 422. Synonymy: Triplehorn (1990: 292).

Subgenus Tylophloeus Bremer, 1998

Tylophloeus Bremer, 1998: 10. Type species: Hypophloeus flavipennis Motschulsky, 1860, original designation.

Corticeus glaber (LeConte, 1878)USA (AL AR DC FL GA LA MD MS NC NJ OH SC TN TX VA WV) / BAH

Hypophloeus glaber LeConte, 1878a: 422.

Genus Myonophloeus Bremer and Lillig, 2017 [M]

Myonophloeus Bremer and Lillig, 2017: 68. Type species: Corticeus tuberculatus Triplehorn, 1979, original designation.

Myonophloeus tuberculatus (Triplehorn, 1979) CUB

Corticeus tuberculatus Triplehorn, 1979: 48.

Tribe Myrmechixenini Jacquelin du Val, 1858

Myrméchixénites Jacquelin du Val, 1858: 223. Type genus: Myrmechixenus Chevrolat, 1835.

Genus Myrmechixenus Chevrolat, 1835 [M]

Myrmechixenus Chevrolat, 1835: 267. Type species: Myrmechixenus subterraneus Chevrolat, 1835, monotypy.

Myrmecoxenus Agassiz, 1846: 243. Unjustified emendation of Myrmechixenus Chevrolat, 1835, not in prevailing usage.

Myrmechixenus latridioides Crotch, 1873USA (CA DC SC TX WA) – Adventive

Myrmecoxenus latridioides Crotch, 1873: 363.

Tribe Phaleriini Blanchard, 1845

Phalériides Blanchard, 1845: 29. Type genus: Phaleria Latreille, 1802.

Sepedonastidae Gistel, 1856a: 382. Type genus: Sepedonastes Gistel, 1856 (= Phaleria Latreille, 1802).

Cataphronetini Reitter, 1917: 57. Type genus: Cataphronetis Lucas, 1846 (= Phtora Germar, 1836).

Genus Phaleria Latreille, 1802 [F]

Phaleria Latreille, 1802: 162. Type species: Tenebrio cadaverinus Fabricius, 1792, subsequent designation (Westwood 1838: 32) (see ICZN 1975).

Sepedonastes Gistel, 1856a: 382. Type species: Tenebrio bimaculatus Herbst, 1799 (= Dytiscus bimaculatus Linnaeus, 1767), subsequent designation (Bouchard et al. 2005: 501). Synonymy: Bouchard et al. (2005: 501).

Halophalerus Crotch, 1874: 107. Type species: Phaleria rotundata LeConte, 1851, present designation. Synonymy: Austin (1880: 38).

Phaleria championi Triplehorn and Watrous, 1980MEX (CL NA SI)

Phaleria championi Triplehorn and Watrous, 1980: 56.

Phaleria debilis LeConte, 1866USA (CA) MEX (BC JA NA SO) GUANIC

Phaleria debilis LeConte, 1866b: 126.

Phaleria neotropicalis Champion, 1886: 220. Synonymy: Triplehorn (1991: 267).

Phaleria insularis Champion, 1886: 221. Synonymy: Triplehorn and Watrous (1979: 288).

Phaleria fulva Fleutiaux and Sallé, 1890DOMLAN / SA

Phaleria fulva Fleutiaux and Sallé, 1890: 423.

Phaleria gracilipes Casey, 1890USA (AL LA TX) MEX (TB VE)

Phaleria gracilipes Casey, 1890b: 484.

Phaleria lodingi Blaisdell, 1932c: 116. Synonymy: Triplehorn and Watrous (1979: 292).

Phaleria guatemalensis Champion, 1886MEX (GE JA OA SI) GUA / SA

Phaleria guatemalensis Champion, 1886: 218.

Phaleria lata Blaisdell, 1923MEX (BC BS SO)

Phaleria latus Blaisdell, 1923: 276.

Phaleria pacifica Champion, 1886MEX (NA) GUANIC

Phaleria pacifica Champion, 1886: 220.

Phaleria panamensis Champion, 1886MEX (BC CO GE JA MI NA SI SO TB) GUABELNICPAN

Phaleria panamensis Champion, 1886: 218.

Phaleria dytiscoides Champion, 1886: 218. Synonymy: Triplehorn (1991: 263).

Phaleria marginipennis Champion, 1886: 219. Synonymy: Triplehorn (1991: 263).

Phaleria opacicollis Champion, 1886: 219. Synonymy: Triplehorn (1991: 263).

Phaleria picipes Say, 1824USA (FL GA MD NC NJ SC VA) MEX (MO QR YU) BELHONPAN / BAHCAYCUBHAIJAMPRILAN / SA

Phaleria picipes Say, 1824b: 280.

Phaleria pilatei Chevrolat, 1879: ccxlix. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria variabilis Quedenfeldt, 1886: 128. Synonymy: Watrous and Triplehorn (1982: 20).

Phaleria caymanensis Marcuzzi, 1977: 34. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria pilifera LeConte, 1866MEX (BC BS SO)

Phaleria pilifera LeConte, 1866b: 125.

Phaleria punctipes LeConte, 1878USA (FL) MEX (QR) BEL / BAH BAR CUBCAYJAMLAN

Phaleria punctipes LeConte, 1878a: 421.

Phaleria guadeloupensis Fleutiaux and Sallé, 1890: 423. Synonymy: Watrous and Triplehorn (1982: 13).

Phaleria jamaicensis Marcuzzi, 1977: 36. Synonymy: Watrous and Triplehorn (1982: 13).

Phaleria rotundata LeConte, 1851USA (CA) MEX (BC BS)

Phaleria rotundata LeConte, 1851: 148.

Phaleria limbata Horn, 1870: 375. Synonymy: Fall (1901: 173, as limbalis).

Phaleria testacea Say, 1824USA (CT DE FL GA LA MA MD ME MS NC NH NJ NY RI SC TX VA) MEX (QR) / BAHCUBJAMHISPRILAN / SA

Phaleria testacea Say, 1824b: 280.

Phaleria brasiliensis Laporte, 1840: 219. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria cayennensis Laporte, 1840: 219. Synonymy: Triplehorn (1991: 267).

Phaleria longula LeConte, 1866b: 125. Synonymy: Triplehorn and Watrous (1979: 289).

Phaleria angustata Chevrolat, 1879: ccxlviii. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria chevrolati Fleutiaux and Sallé, 1890: 422. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria chevrolati var. thoracica Fleutiaux and Sallé, 1890: 423. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria chevrolati var. quadrinotata Fleutiaux and Sallé, 1890: 423. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria maculipennis Marcuzzi, 1962: 37. Synonymy: Watrous and Triplehorn (1982: 19).

Phaleria thinophila Watrous and Triplehorn, 1982CRI / JAMDOMPRILAN / SA

Phaleria thinophila Watrous and Triplehorn, 1982: 15.

Genus Phaleromela Reitter, 1916 [F]

Phaleromela Reitter, 1916: 4. Type species: Phaleria subhumeralis Marseul, 1876, monotypy.

Phaleromela humeralis (Laporte, 1840)USA (CA)

Phaleria humeralis Laporte, 1840: 219.

Phaleromela picta (Mannerheim, 1843)CAN (BC) USA (AK CA OR WA)

Phaleria picta Mannerheim, 1843: 277.

Phaleria globosa LeConte, 1857: 51. New synonymy [YB].

Phaleromela prohumeralis Triplehorn, 1961USA (CA)

Phaleria humeralis Horn, 1870: 377 [junior primary homonym of Phaleria humeralis Laporte, 1840].

Phaleromela prohumeralis Triplehorn, 1961: 127. Replacement name for Phaleromela humeralis (Horn, 1870).

Phaleromela variegata Triplehorn, 1961 [Fig. 38] CAN (AB BC NT SK YT) USA (CA ID OR WA)

Figure 38. 

Phaleromela variegata Triplehorn, 1961. Scale bar = 1 mm.

Scaphidema pictum Horn, 1874a: 36 [junior secondary homonym of Phaleromela picta (Mannerheim, 1843)].

Phaleromela variegata Triplehorn, 1961: 126. Replacement name for Phaleromela picta (Horn, 1874).

Tribe Scaphidemini Reitter, 1922

Scaphidemini Reitter, 1922: 2. Type genus: Scaphidema Redtenbacher, 1848.

Genus Scaphidema Redtenbacher, 1848 [F]

Scaphidema Redtenbacher, 1848: 591. Type species: Scaphidium bicolor Fabricius, 1798, monotypy.

Nelites LeConte, 1850: 232. Type species: Nelites aeneolus LeConte, 1850, monotypy. Synonymy: LeConte (1862a: 237).

Microbasanus Pic, 1921: 1. Type species: Microbasanus jureceki Pic, 1921, monotypy. Synonymy: Löbl et al. (2008a: 318) (see also Schawaller 2008: 384).

Scaphidema aeneola (LeConte, 1850)CAN (AB BC LB MB NB NS ON PE QC SK YT) USA (AK CT IL MA MI MN NH NY OH PA WI WY)

Nelites aeneolus LeConte, 1850: 232.

Tribe Trachyscelini Blanchard, 1845

Trachyscélides Blanchard, 1845: 28. Type genus: Trachyscelis Latreille, 1809.

Genus Trachyscelis Latreille, 1809 [F]

Trachyscelis Latreille, 1809: 379. Type species: Trachyscelis aphodioides Latreille, 1809, monotypy.

Trachyscelis aphodioides Latreille, 1809USA76 (FL LA MD NC VA) / BAHTURPRILAN – Adventive

Trachyscelis aphodioides Latreille, 1809: 379.

Trachyscelis flavipes Melsheimer, 1846: 61. Synonymy: Steiner (2004: 335).

Subfamily STENOCHIINAE Kirby, 1837

Stenochiadae Kirby, 1837: 238. Type genus: Stenochia Kirby, 1819 (= Strongylium Kirby, 1819).

Tribe Cnodalonini Oken, 1843

Cnodaliden Oken, 1843: 484. Type genus: Cnodalon Latreille, 1797.

Coelometopidae Schaum, 1859: 71. Type genus: Coelometopus Solier, 1848.

Upidae C.G. Thomson, 1859: 116. Type genus: Upis Fabricius, 1792.

Catapiestides Lacordaire, 1859: 381. Type genus: Catapiestus Perty, 1831.

Polypleuri LeConte, 1862a: 229. Type genus: Polypleurus Eschscholtz, 1831.

Hegemonini Reitter, 1922: 3. Type genus: Hegemona Laporte, 1840.

Camariinae Anonymous, 1924: 164. Type genus: Camaria Lepeletier and Audinet-Serville, 1828.

Genus Alobates Motschulsky, 1872 [M]

Alobates Motschulsky, 1872: 25. Type species: Tenebrio pensylvanicus DeGeer, 1775, original designation.

Alobates barbatus (Knoch, 1801)77USA (AL AR DC DE FL GA IL IN KS KY LA MA MD MI MO MS NC NJ NY OH RI SC TN TX VA WI WV)

Upis glabra Herbst, 1799: 32 [nomen dubium].

Tenebrio barbatus Knoch, 1801: 166. Synonymy (in doubt): LeConte (1866a: 61).

Nyctobates intermedia Haldeman, 1852: 376. Synonymy: Horn (1870: 333).

Alobates pensylvanicus (DeGeer, 1775)CAN (MB NB NS ON QC) USA (AL AZ CA CT DC DE FL GA IA IN KS KY LA MA MD ME MI MN MS NC NH NJ OH OR PA SC SD TN TX VA WI WV) MEX (NL)

Tenebrio pensylvanicus DeGeer, 1775: 53.

Upis chrysops Herbst, 1797: 236. Synonymy (in doubt): Knoch (1801: 168).

Tenebrio sublaevis Palisot de Beauvois, 1817: 163. Synonymy: LeConte (1866a: 61).

Tenebrio striatellus Drapiez, 1820: 327. Synonymy: Horn (1889: 212).

Nyctobates inermis Mannerheim, 1843: 284. Synonymy: Horn (1870: 333).

Alobates pennsylvanicus Champion, 1892: 522. Unjustified emendation of Alobates pensylvanicus (DeGeer, 1775), not in prevailing usage.

Genus Apsida Lacordaire, 1859 [F]

Apsida Lacordaire, 1859: 309. Type species: Apsida chrysomelina Lacordaire, 1859, original designation.

Hapsida Gemminger [in Gemminger and Harold], 1870: 1955. Unjustified emendation of Apsida Lacordaire, 1859, not in prevailing usage.

Apsida belti Bates, 1873USA (TX) MEX (TA TB VE YU) GUANICCRI

Apsida belti Bates, 1873e: 16.

Apsida boucardi Bates, 1873MEX (VE) GUABELCRIPAN

Apsida boucardi Bates, 1873e: 17.

Cosmonota geminata Chevrolat, 1877b: 173. Synonymy: Champion (1886: 215).

Cosmonota grammica Chevrolat, 1877b: 173. Synonymy: Champion (1886: 215).

Apsida chrysomelina Lacordaire, 1859MEX (CI VE) GUABELNICCRIPAN / SA

Apsida chrysomelina Lacordaire, 1859: 309.

Apsida chrysomelina Bates, 1873e: 15 [junior primary homonym of Apsida chrysomelina Lacordaire, 1859]. Synonymy: Champion (1886: 211).

Apsida gibbosa Champion, 1886MEX (VE) GUABELCRI

Hapsida gibbosa Champion, 1886: 212.

Apsida lustrans Triplehorn, 1970 CRI

Apsida lustrans Triplehorn, 1970: 569.

Apsida punctipennis Champion, 1886 GUA

Hapsida punctipennis Champion, 1886: 213.

Apsida purpureomicans Bates, 1873MEX (VE) GUABELCRIPAN / SA

Apsida purpureomicans Bates, 1873e: 16.

Apsida aeneomicans Bates, 1873e: 16. Synonymy: Triplehorn (1970: 571).

Apsida seriatopunctata Champion, 1886MEX (VE)

Hapsida seriato-punctata Champion, 1886: 212.

Apsida simulatrix Ferrer and Ødegaard, 2005 PAN

Apsida simulatrix Ferrer and Ødegaard, 2005: 640.

Apsida terebrans Champion, 1886MEX (TB) GUABELNICCRIPAN

Hapsida terebrans Champion, 1886: 214.

Genus Blapida Perty, 1830 [F]

Blapida Perty, 1830: 58. Type species: Blapida okeni Perty, 1830, monotypy.

Ryssochiton Gray [in Griffith and Pidgeon], 1832: pl. 50. Type species: Rhyssochiton politus Gray, 1832, monotypy. Synonymy: Lacordaire (1859: 425).

Blapida alternata Gebien, 1919 CRI PAN

Blapida alternata Gebien, 1919: 137.

Blapida castaneipennis Champion, 1896 LAN

Blapida castaneipennis Champion, 1896: 28.

Blapida neotropicalis Champion, 1886 GUA NIC

Blapida neotropicalis Champion, 1886: 247.

Genus Bothynocephalus Doyen, 1988 [M]

Bothynocephalus Doyen, 1988: 315. Type species: Bothynocephalus cristatus Doyen, 1988, original designation.

Bothynocephalus cristatus Doyen, 1988 HON

Bothynocephalus cristatus Doyen, 1988: 316.

Bothynocephalus foveolatus Doyen, 1995MEX (OA)

Bothynocephalus foveolatus Doyen, 1995: 12.

Bothynocephalus ribardoi Doyen, 1995 CRI

Bothynocephalus ribardoi Doyen, 1995: 13.

Bothynocephalus thoracicus Doyen, 1995 CRI

Bothynocephalus thoracicus Doyen, 1995: 12.

Genus Brosimapsida Ferrer and Ødegaard, 2005 [F]

Brosimapsida Ferrer and Ødegaard, 2005: 640. Type species: Brosimapsida gonospoides Ferrer and Ødegaard, 2005, original designation.

Brosimapsida gonospoides Ferrer and Ødegaard, 2005 PAN

Brosimapsida gonospoides Ferrer and Ødegaard, 2005: 640.

Genus Calydonella Doyen, 1995 [F]

Calydonella Doyen, 1995: 8. Type species: Calydonella lisa Doyen, 1995, original designation.

Calydonella lisa Doyen, 1995 CRI PAN

Calydonella lisa Doyen, 1995: 10.

Genus Camaria Lepeletier and Audinet-Serville, 1828 [F]

Camaria Lepeletier and Audinet-Serville, 1828: 454. Type species: Camaria nitida Lepeletier and Audinet-Serville, 1828, monotypy.

Camaria laevis Gebien, 1919 “Zentralamerika” / SA

Camaria laevis Gebien, 1919: 52.

Camaria parallela Champion, 1886 PAN

Camaria parallela Champion, 1886: 246.

Genus Choastes Champion, 1893 [M]

Choaspes Champion, 1885: 118 [junior homonym of Choaspes Moore, 1881]. Type species: Choaspes purpureus Champion, 1885, subsequent designation (Gebien 1941: 338).

Choastes Champion, 1893a: 526. Replacement name for Choaspes Champion, 1885.

Choastes angulicollis (Champion, 1885) NIC PAN

Choaspes angulicollis Champion, 1885: 119.

Choastes purpureus (Champion, 1885) GUA BEL NIC PAN

Choaspes purpureus Champion, 1885: 119.

Genus Cibdelis Mannerheim, 1843 [F]

Cibdelis Mannerheim, 1843: 282. Type species: Cibdelis blaschkii Mannerheim, 1843, monotypy.

Scotera Motschulsky, 1845b: 365. Type species: Scotera gibbosa Motschulsky, 1845, monotypy.78 Synonymy: Motschulsky (1845b: 365).

Cibdelis bachei LeConte, 1861USA (CA)

Cibdelis bachei LeConte, 1861b: 353.

Cibdelis blaschkii Mannerheim, 1843USA (CA)

Cibdelis blaschkii Mannerheim, 1843: 284.

Cibdelis cylindrica Casey, 1924USA (CA)

Cibdelis cylindrica Casey, 1924: 323.

Cibdelis gibbosa (Motschulsky, 1845)USA (CA)

Scotera gibbosa Motschulsky, 1845b: 365.

Cibdelis laevigata Casey, 1891: 60. Synonymy: Leng (1920: 235).

Cibdelis ventricosa Casey, 1924USA (CA)

Cibdelis ventricosa Casey, 1924: 322.

Genus Cnephalura Doyen, 1988 [F]

Cnephalura Doyen, 1988: 313. Type species: Cnephalura umbrata Doyen, 1988, original designation.

Cnephalura umbrata Doyen, 1988MEX (CI)

Cnephalura umbrata Doyen, 1988: 313.

Genus Cnodalon Latreille, 1797 [N]

Cnodalon Latreille, 1797: 23. Type species: Cnodalum viride Latreille, 1804, subsequent monotypy in Latreille (1804: 321).

Cnodalum Agassiz, 1846: 91. Unjustified emendation for Cnodalon Latreille, 1797, not in prevailing usage.

Cnodalon viride Latreille, 1804 HAI

Cnodalum viride Latreille, 1804: 321.

Genus Coelocnemis Mannerheim, 1843 [F]

Coelocnemis Mannerheim, 1843: 280. Type species: Coelocnemis dilaticollis Mannerheim, 1843, subsequent designation (Lucas 1920: 194).

Coelocnemis dilaticollis Mannerheim, 184379CAN (AB BC) USA (CA ID MT NV OR UT WA WY)

Coelocnemis dilaticollis Mannerheim, 1843: 282.

Coelocnemis californica Mannerheim, 1843: 282. Synonymy: Horn (1870: 336).

Coelocnemis rugosa Linell, 1899: 181. Synonymy: Doyen (1973: 93).

Coelocnemis columbiana Casey, 1924: 314. Synonymy: Boddy (1965: 175).

Coelocnemis rauca Casey, 1924: 315. Synonymy: Doyen (1973: 93).

Coelocnemis ovipennis Casey, 1924: 315. Synonymy: Doyen (1973: 93).

Coelocnemis utensis Casey, 1924: 316. Synonymy: Doyen (1973: 93).

Coelocnemis spaldingi Casey, 1924: 316. Synonymy: Doyen (1973: 93).

Coelocnemis basalis Casey, 1924: 316. Synonymy: Doyen (1973: 93).

Coelocnemis idahoensis Casey, 1924: 318. Synonymy: Boddy (1965: 175).

Coelocnemis barretti Blaisdell, 1928: 163. Synonymy: Doyen (1973: 93).

Coelocnemis lucia Doyen, 1973USA (CA)

Coelocnemis lucia Doyen, 1973: 96.

Coelocnemis magna LeConte, 1851USA (AZ CA NM) MEX (BC)

Coelocnemis magna LeConte, 1851: 150.

Coelocnemis obesa LeConte, 1851: 150. Synonymy: Doyen (1973: 87).

Coelocnemis caudicalis Casey, 1924: 316. Synonymy: Doyen (1973: 87).

Coelocnemis caudicalis deserta Casey, 1924: 316. Synonymy: Doyen (1973: 87).

Coelocnemis antennalis Casey, 1924: 317. Synonymy: Doyen (1973: 87).

Coelocnemis aequalis Casey, 1924: 318. Synonymy: Doyen (1973: 87).

Coelocnemis smithiana Casey, 1924: 318. Synonymy: Doyen (1973: 87).

Coelocnemis rotundicollis Casey, 1924: 319. Synonymy: Doyen (1973: 87).

Coelocnemis longicollis Casey, 1924: 319. Synonymy: Doyen (1973: 87).

Coelocnemis punctata LeConte, 1854USA (AZ CA CO ID NM NV OR UT)

Coelocnemis punctatus LeConte, 1854c: 225.

Coelocnemis angusta Casey, 1924: 320. Synonymy: Doyen (1973: 98).

Coelocnemis tanneri Blaisdell, 1928: 164. Synonymy: Doyen (1973: 98).

Coelocnemis rugulosa Doyen, 1973USA (CA OR)

Coelocnemis rugulosa Doyen, 1973: 97.

Coelocnemis slevini Blaisdell, 1925MEX (BC)

Coelocnemis slevini Blaisdell, 1925b: 337.

Coelocnemis sulcata Casey, 1895USA (AZ CA ID NV UT)

Coelocnemis sulcata Casey, 1895: 615.

Genus Cyrtosoma Perty, 1830 [N]

Cyrtosoma Perty, 1830: 59. Type species: Cyrtosoma unicolor Perty, 1830, monotypy.

Cyrtosoma arimense Marcuzzi, 1999 LAN

Cyrtosoma arimensis Marcuzzi, 1999: 85.

Cyrtosoma decemlineatum Champion, 1886MEX (VE) BELNICPAN

Cyrtosoma decem-lineatum Champion, 1886: 244.

Cyrtosoma denticolle Chevrolat, 1878MEX (VE) GUABELNICCRIPAN / SA

Cyrtosoma denticolle Chevrolat, 1878e: 273.

Cyrtosoma grenadense Marcuzzi, 1999LAN (Grenada)

Cyrtosoma grenadensis Marcuzzi, 1999: 85.

Cyrtosoma lherminierii (Guérin-Méneville, 1844) LAN

Cnodalon atrum Guérin-Méneville, 1833: pl. 31 [junior primary homonym of Cnodalon atrum Lepeletier and Audinet-Serville, 1825].

Cnodalon l’herminierii Guérin-Méneville80, 1844: 123. Replacement name for Cnodalon atrum Guérin-Méneville, 1833.

Cyrtosoma martiniquense Marcuzzi, 1999LAN (Martinique)

Cyrtosoma martiniquensis Marcuzzi, 1999: 83.

Cyrtosoma piceum (Laporte and Brullé, 1831)LAN (Guadeloupe)

Platydema picea Laporte and Brullé, 1831: 362.

Cyrtosoma williamsi Marcuzzi, 1992 PAN

Cyrtosoma williamsi Marcuzzi, 1992: 237.

Genus Dinomus Brême, 1842 [M]

Dinomus Brême, 1842: 113. Type species: Dinomus perforatus Brême, 1842, monotypy.

Dinomus perforatus Brême, 1842 MEX

Dinomus perforatus Brême, 1842: 114.

Genus Elomosda Bates, 1870 [F]

Elomosda Bates, 1870: 273. Type species: Elomosda beltii Bates, 1870, monotypy.

Elomosda beltii Bates, 1870 GUA NIC CRI

Elomosda beltii Bates, 1870: 275.

Genus Epicalla Champion, 1886 [F]

Epicalla Champion, 1886: 249. Type species: Epicalla varipes Champion, 1886, subsequent designation (Lucas 1920: 268).

Epicalla aenipes Ferrer and Ødegaard, 2005 PAN

Epicalla aenipes Ferrer and Ødegaard, 2005: 642.

Epicalla agnata Gebien, 1928 CRI

Epicalla agnata Gebien, 1928b: 217.

Epicalla avia Gebien, 1928 BEL

Epicalla avia Gebien, 1928b: 209.

Epicalla cupreonitens Champion, 1886 PAN

Epicalla cupreo-nitens Champion, 1886: 250.

Epicalla elongata Ferrer and Ødegaard, 2005 PAN

Epicalla elongata Ferrer and Ødegaard, 2005: 642.

Epicalla famula Gebien, 1928 CRI

Epicalla famula Gebien, 1928b: 216.

Epicalla hera Gebien, 1928 CRI

Epicalla hera Gebien, 1928b: 207.

Epicalla instriata Pic, 1921 PAN

Epicalla instriata Pic, 1921b: 28.

Epicalla juvenca Gebien, 1928 NIC CRI

Epicalla juvenca Gebien, 1928b: 218.

Epicalla lata Champion, 1886MEX (JA SI)

Epicalla lata Champion, 1886: 250.

Epicalla nevermanni Gebien, 1928 CRI

Epicalla nevermanni Gebien, 1928b: 215.

Epicalla pygmaea Ferrer and Ødegaard, 2005 PAN

Epicalla pygmaea Ferrer and Ødegaard, 2005: 642.

Epicalla varipes Champion, 1886 NIC

Epicalla varipes Champion, 1886: 249.

Genus Glyptotus LeConte, 1858 [M]

Glyptotus LeConte, 1858b: 75. Type species: Glyptotus cribratus LeConte, 1858, monotypy.

Glyptotus cribratus LeConte, 1858USA (AL FL GA MS NC SC TX VA) MEX / BAH

Glyptotus cribratus LeConte, 1858b: 75.

Glyptotus nitidus Champion, 1885MEX (VE) NIC

Glyptotus nitidus Champion, 1885: 113.

Glyptotus yucatanus Champion, 1892MEX (YU)

Glyptotus yucatanus Champion, 1892: 524.

Genus Gonospa Champion, 1886 [F]

Gonospa Champion, 1886: 216. Type species: Gonospa phaedonoides Champion, 1886, subsequent designation (Gebien 1940: 426).

Gonospa phaedonoides Champion, 1886 PAN

Gonospa phaedonoides Champion, 1886: 217.

Gonospa similis Ferrer and Ødegaard, 2005 PAN

Gonospa similis Ferrer and Ødegaard, 2005: 639.

Genus Haplandrus LeConte, 1862 [M]

Haplandrus LeConte, 1862a: 230. Type species: Helops femoratus Fabricius, 1798 (= Upis fulvipes Herbst, 1797), monotypy.

Haplandrus deyruporum Steiner, 2016USA (FL)

Haplandrus deyruporum Steiner, 2016: 537.

Haplandrus fulvipes (Herbst, 1797)CAN (NS ON QC) USA (CT FL GA IA IN MD MI NC NY OH PA RI SC TN VA WI)

Upis fulvipes Herbst, 1797: 238.

Helops femoratus Fabricius, 1798: 53. Synonymy: Schönherr (1806: 157).

Genus Hegemona Laporte, 1840 [F]

Hegemona Laporte, 1840: 230. Type species: Hegemona resplendens Laporte, 1840, monotypy.

Eucamptus Germar, 1842: 444 [junior homonym of Eucamptus Chevrolat, 1833]. Type species: Eucamptus iridis Germar, 1842 (= Hegemona resplendens Laporte, 1840), monotypy. Synonymy: Duponchel (1845: 498).

Eusarca Chevrolat, 1845: 526. Type species: Eusarca iridipennis Chevrolat, 1845 (= Hegemona resplendens Laporte, 1840), monotypy. Synonymy: Duponchel (1845: 498).

Hegemona alternata Pic, 1936 GUA

Hegemona alternatus Pic, 1936: 15.

Hegemona angustata Champion, 1887 GUA

Hegemona angustatus Champion, 1887: 272.

Hegemona bicaudata Champion, 1887 GUA

Hegemona bicaudatus Champion, 1887: 270.

Hegemona chiriquensis Champion, 1887 CRI PAN

Hegemona chiriquensis Champion, 1887: 273.

Hegemona compressa Allard, 1877 MEX GUA

Hegemona compressus Allard, 1877b: 61, 254.

Hegemona costaricensis Champion, 1887 CRI

Hegemona costaricensis Champion, 1887: 275.

Hegemona elongata Allard, 1877MEX (YU)

Hegemona elongatus Allard, 1877b: 61, 253.

Hegemona flibuster (J. Thomson, 1856) MEX GUA BEL NIC CRI

Eucamptus flibuster J. Thomson, 1856: 475.

Hegemona filibuster Champion, 1887: 275. Unjustified emendation of Hegemona flibuster (J. Thomson, 1856), not in prevailing usage.

Hegemona furcillata Allard, 1877 MEX

Hegemona furcillatus Allard, 1877b: 61, 252.

Hegemona guatemalensis Champion, 1887 GUA

Hegemona guatemalensis Champion, 1887: 274.

Hegemona hondurensis Champion, 1887 BEL

Hegemona hondurensis Champion, 1887: 269.

Hegemona interrupta Champion, 1887 CRI

Hegemona interruptus Champion, 1887: 275.

Hegemona lineata Champion, 1887 GUA BEL

Hegemona lineatus Champion, 1887: 271.

Hegemona mexicana Champion, 1887MEX (PU)

Hegemona mexicanus Champion, 1887: 274.

Hegemona nigra Champion, 1887 GUA HON

Hegemona niger Champion, 1887: 271.

Hegemona refulgens Champion, 1893 GUA

Hegemona refulgens Champion, 1893a: 549.

Hegemona resplendens Laporte, 1840MEX (VE YU)

Hegemona resplendens Laporte, 1840: 230.

Eucamptus iridis Germar, 1842: 444. Synonymy: Duponchel (1845: 498).

Eusarca iridipennis Chevrolat, 1845: 526. Synonymy: Duponchel (1845: 498).

Hegemona retrodentata Allard, 1877MEX (CI OA)

Hegemona retrodentatus Allard, 1877b: 61, 253.

Hegemona zunilensis Champion, 1887 GUA

Hegemona zunilensis Champion, 1887: 272.

Genus Hesiodus Champion, 1885 [M]

Hesiodus Champion, 1885: 115. Type species: Hesiodus longitarsus Champion, 1885, subsequent designation (Lucas 1920: 323).

Hesiodus caraibus Fleutiaux and Sallé, 1890LAN (Guadeloupe)

Hesiodus caraibus Fleutiaux and Sallé, 1890: 424.

Hesiodus conspurcatus Champion, 1885 PAN

Hesiodus conspurcatus Champion, 1885: 116.

Hesiodus debilis Champion, 1885 GUA

Hesiodus debilis Champion, 1885: 117.

Hesiodus ellipticus Champion, 1893 NIC

Hesiodus ellipticus Champion, 1893a: 525.

Hesiodus jansoni Champion, 1885MEX (YU) NIC

Hesiodus jansoni Champion, 1885: 116.

Hesiodus longitarsis Champion, 1885MEX (VE) BELNIC

Hesiodus longitarsis Champion, 1885: 115.

Hesiodus sordidus Champion, 1885MEX (VE)

Hesiodus sordidus Champion, 1885: 116.

Genus Hicetaon Champion, 1885 [M]

Hicetaon Champion, 1885: 111. Type species: Hicetaon frontalis Champion, 1885, monotypy.

Hicetaon frontalis Champion, 1885MEX (VE YU) BEL

Hicetaon frontalis Champion, 1885: 112.

Genus Ilus Champion, 1885 [M]

Ilus Champion, 1885: 117. Type species: Ilus apicicornis Champion, 1885, monotypy.

Ilus apicicornis Champion, 1885 BEL CRI

Ilus apicicornis Champion, 1885: 118.

Genus Iphthiminus Spilman, 1973 [M]

Iphthiminus Spilman, 1973: 42. Type species: Iphthimus italicus Truqui, 1857, original designation.

Iphthiminus lewisii (Horn, 1870)USA (AZ CA CO NM NV TX UT WY) MEX (BC CH)

Iphthimus lewisii Horn, 1870: 335.

Iphthimus laevissimus Casey, 1890b: 408. Synonymy: Gardiner and Pollock (2015: 364).

Iphthiminus opacus (LeConte, 1866)CAN (AB MB NB NS ON QC SK) USA (AZ CA CT IN MA ME MI MN MT NH NY OH PA SD VA VT WI WY)

Iphthimus opacus LeConte, 1866b: 121.

Iphthiminus serratus (Mannerheim, 1843) [Fig. 39] CAN (AB BC) USA (CA ID MT NE NV OR WA WY)

Figure 39. 

Iphthiminus serratus (Mannerheim, 1843). Scale bar = 1 mm.

Nyctobates serrata Mannerheim, 1843: 284.

Nyctobates sublaevis Bland, 1865: 382. Synonymy: Gardiner and Pollock (2015: 356).

Iphthinus servilis Walker, 1866: 326. Synonymy: Horn (1870: 334).

Iphthinus servator Walker, 1866: 327. Synonymy: Horn (1870: 334).

Iphthinus subligatus Walker, 1866: 327. Synonymy: Horn (1870: 334).

Iphthimus salebrosus Casey, 1924: 327. Synonymy: Gardiner and Pollock (2015: 356).

Genus Isaminas Champion, 1887 [M]

Isaminas Champion, 1887: 266. Type species: Isaminas gibbipennis Champion, 1887, subsequent designation (Gebien 1943: 401).

Pteroglymmius Gebien, 1928b: 223. Type species: Pteroglymmius erotyloides Gebien, 1928, monotypy. Synonymy: Doyen (1988: 301).

Isaminas breedlovei Doyen, 1988MEX (CI)

Isaminas breedlovei Doyen, 1988: 303.

Isaminas brevicollis Champion, 1887MEX (CI) GUA

Isaminas brevicollis Champion, 1887: 267.

Isaminas erotyloides (Gebien, 1928) HON

Pteroglymmius erotyloides Gebien, 1928b: 224.

Isaminas gibbipennis Champion, 1887 NIC CRI

Isaminas gibbipennis Champion, 1887: 267.

Isaminas reticuloides Doyen, 1988MEX (CI)

Isaminas reticuloides Doyen, 1988: 303.

Isaminas sullivani Doyen, 1988 CRI

Isaminas sullivani Doyen, 1988: 306.

Genus Isicerdes Champion, 1885 [M]

Isicerdes Champion, 1885: 113. Type species: Isicerdes occultus Champion, 1885, subsequent designation (Lucas 1920: 353).

Isicerdes funebris Champion, 1885 GUA

Isicerdes funebris Champion, 1885: 114.

Isicerdes occultus Champion, 1885MEX (VE) GUABELPAN

Isicerdes occultus Champion, 1885: 114.

Isicerdes vicinus Champion, 1892MEX (JA YU) CRI

Isicerdes vicinus Champion, 1892: 524.

Genus Lenkous Kaszab, 1973 [M]

Lenkous Kaszab, 1973: 315. Type species: Lenkous myrmecophilus Kaszab, 1973, original designation.

Lenkous ibisca Ferrer and Ødegaard, 2005 PAN

Lenkous ibisca Ferrer and Ødegaard, 2005: 639.

Genus Merinus LeConte, 1862 [M]

Merinus LeConte, 1862a: 230. Type species: Tenebrio laevis Olivier, 1795, original designation.

Merinus laevis (Olivier, 1795) [Fig. 40] CAN (ON QC) USA (AL CT FL DC GA IA IL IN KS MD MI MO MS NC NJ NY OH OK PA RI SC TN VA WI WV)

Figure 40. 

Merinus laevis (Olivier, 1795). Scale bar = 1 mm.

Tenebrio laevis Olivier, 1795: [57] 10.

Genus Mitys Champion, 1885 [M]

Mitys Champion, 1885: 97. Type species: Mitys inflatus Champion, 1885, subsequent designation (Gebien 1943: 402).

Mitys inflatus Champion, 1885MEX (OA VE)

Mitys inflatus Champion, 1885: 97.

Mitys opacus Champion, 1885MEX (TA)

Mitys opacus Champion, 1885: 98.

Mitys politus (Brême, 1842)MEX (PU VE)

Sphoerotus politus Brême, 1842: 109.

Mitys laevis Champion, 1885: 98. Synonymy: Champion (1892: 520).

Genus Moeon Champion, 1886 [N]81

Moeon Champion, 1886: 251. Type species: Moeon isthmicus Champion, 1886, subsequent designation (Gebien 1942: 330).

Moeon isthmicum Champion, 1886 PAN

Moeon isthmicus Champion, 1886: 251.

Moeon panamense Champion, 1886 PAN

Moeon panamensis Champion, 1886: 251.

Genus Mophon Champion, 1886 [M]

Mophon Champion, 1886: 247. Type species: Mophon tinctipennis Champion, 1886, monotypy.

Mophon tinctipennis Champion, 1886 NIC PAN

Mophon tinctipennis Champion, 1886: 248.

Genus Mylaris Pallas, 1781 [F]

Mylaris Pallas, 1781: 3782. Type species: Tenebrio gigas Linnaeus, 1763, subsequent designation (Guérin-Méneville 1844: 120).

Iphthinus Dejean, 1834 [30 June]: 203. Type species: Tenebrio gigas Linnaeus, 1763, subsequent designation (Spilman 1973: 42). Synonymy: Spilman (1973: 42).

Cecrops Gistel, 1834 [23 September]: 21. Type species: Tenebrio gigas Linnaeus, 1763, subsequent designation (Bousquet and Bouchard 2017a: 131). Synonymy: Bousquet and Bouchard (2017a: 132).

Nyctobates Guérin-Méneville, 1834 [“31 December”]: 33. Type species: Tenebrio gigas Linnaeus, 1763, original designation. Synonymy: Ferrer and Siliansky (2007: 186).

Mylaris gigas (Linnaeus, 1763)MEXGUANICCRIPAN / SA

Tenebrio gigas Linnaeus, 1763: 13.

Mylaris gigantea Pallas, 1781: 37. Synonymy: Pallas (1781: 37).

Tenebrio laminatus Fabricius, 1787: 211. Synonymy: Fabricius (1801a: 144).

Mylaris procera (Champion, 1885)MEX (SI VE) GUABELPAN / SA

Nyctobates procerus Champion, 1885: 107.

Genus Nesocyrtosoma Marcuzzi, 1976 [N]

Nesocyrtosoma Marcuzzi, 1976: 137. Type species: Cyrtosoma inflatum Marcuzzi, 1976, designation of the International Commission on Zoological Nomenclature (ICZN 2017).

Pachycyrtosoma Marcuzzi, 1999: 81. Type species: Cyrtosoma merkli Marcuzzi, 1999, original designation. Synonymy: Hopp and Ivie (2009: 13).

Serrania Garrido, 2003: 50. Type species: Diaperis viridula Zayas, 1988 (= Platydema virens Laporte and Brullé, 1831), monotypy. Synonymy: Hopp and Ivie (2009: 13).

Nesocyrtosoma altagracia Hopp and Ivie, 2009 DOM

Nesocyrtosoma altagracia Hopp and Ivie, 2009: 40.

Nesocyrtosoma bankense Hopp and Ivie, 2009 PRI LAN

Nesocyrtosoma bankense Hopp and Ivie, 2009: 57.

Nesocyrtosoma basilense Hopp and Ivie, 2009 HAI

Nesocyrtosoma basilense Hopp and Ivie, 2009: 41.

Nesocyrtosoma bestiola Hopp and Ivie, 2009 DOM

Nesocyrtosoma bestiola Hopp and Ivie, 2009: 20.

Nesocyrtosoma crenulatum Hopp and Ivie, 2009 HAI DOM

Nesocyrtosoma crenulatum Hopp and Ivie, 2009: 44.

Nesocyrtosoma bromelicolus Garrido and Varela, 2010: 33. Synonymy: Hopp (2011: 242).

Nesocyrtosoma critalense (Zayas, 1988) CUB

Cnodalon critalensis Zayas, 1988: 99.

Nesocyrtosoma cubanense (Kulzer, 1961) CUB

Apsida cubanensis Kulzer, 1961a: 217.

Nesocyrtosoma cuproso (Zayas, 1988) 83 CUB

Cnodalon cuproso Zayas, 1988: 98.

Nesocyrtosoma curvum Hopp and Ivie, 2009 PRI

Nesocyrtosoma curvum Hopp and Ivie, 2009: 68.

Nesocyrtosoma darlingtoni Hopp and Ivie, 2009 HAI

Nesocyrtosoma darlingtoni Hopp and Ivie, 2009: 47.

Nesocyrtosoma dentatum Hopp and Ivie, 2009 CUB

Nesocyrtosoma dentatum Hopp and Ivie, 2009: 76.

Nesocyrtosoma dolosum Hopp and Ivie, 2009 HAI

Nesocyrtosoma dolosum Hopp and Ivie, 2009: 42.

Nesocyrtosoma elongatum (Zayas, 1988) CUB

Cnodalon elongatus Zayas, 1988: 101.

Nesocyrtosoma fernandoi Hopp and Ivie, 2009 CUB

Nesocyrtosoma fernandoi Hopp and Ivie, 2009: 59.

Nesocyrtosoma ferrugineum (Garrido and Gutiérrez, 1996) CUB

Cyrtosoma ferruginea Garrido and Gutiérrez, 1996c: 282.

Nesocyrtosoma garridoi Hopp and Ivie, 2009 CUB

Nesocyrtosoma garridoi Hopp and Ivie, 2009: 60.

Nesocyrtosoma gebieni (Marcuzzi, 1976) CUB

Cyrtosoma gebieni Marcuzzi, 1976: 139.

Cnodalon punctatum Zayas, 1988: 103. Synonymy: Garrido and Gutiérrez (1996c: 282).

Nesocyrtosoma guerreroi Hopp and Ivie, 2009 DOM

Nesocyrtosoma guerreroi Hopp and Ivie, 2009: 69.

Nesocyrtosoma hispaniolae (Marcuzzi, 1999) DOM

Cyrtosoma hispaniolae Marcuzzi, 1999: 83.

Nesocyrtosoma inflatum (Marcuzzi, 1976) CUB

Cyrtosoma inflatum Marcuzzi, 1976: 138.

Cnodalon trinitatis Zayas, 1988: 102 [junior secondary homonym of Cyrtosoma trinitatis Marcuzzi, 1976]. Synonymy: Hopp and Ivie (2009: 31).

Cyrtosoma iviei Marcuzzi, 1998a: 160. Replacement name for Cyrtosoma trinitatis (Zayas, 1988).

Nesocyrtosoma lacrima Hopp and Ivie, 2009 LAN

Nesocyrtosoma lacrima Hopp and Ivie, 2009: 36.

Nesocyrtosoma larseni Hopp and Ivie, 2009 CUB DOM

Nesocyrtosoma larseni Hopp and Ivie, 2009: 50.

Nesocyrtosoma merkli (Marcuzzi, 1999) DOM

Cyrtosoma merkli Marcuzzi, 1999: 82.

Nesocyrtosoma mutabile Hopp and Ivie, 2009 DOM

Nesocyrtosoma mutabile Hopp and Ivie, 2009: 48.

Nesocyrtosoma nearnsi Hopp and Ivie, 2009 HAI DOM

Nesocyrtosoma nearnsi Hopp and Ivie, 2009: 65.

Nesocyrtosoma neibaense Hopp and Ivie, 2009 HAI DOM

Nesocyrtosoma neibaense Hopp and Ivie, 2009: 29.

Nesocyrtosoma otus Hopp and Ivie, 2009 HAI DOM

Nesocyrtosoma otus Hopp and Ivie, 2009: 45.

Nesocyrtosoma parallelum (Zayas, 1988) CUB

Cnodalon parallelus Zayas, 1988: 96.

Nesocyrtosoma productum Hopp and Ivie, 2009 DOM

Nesocyrtosoma productum Hopp and Ivie, 2009: 67.

Nesocyrtosoma puertoricense Hopp and Ivie, 2009 PRI

Nesocyrtosoma puertoricense Hopp and Ivie, 2009: 61.

Nesocyrtosoma purpureum Hopp and Ivie, 2009 DOM

Nesocyrtosoma purpureum Hopp and Ivie, 2009: 43.

Nesocyrtosoma scabrosum Hopp and Ivie, 2009 HAI

Nesocyrtosoma scabrosum Hopp and Ivie, 2009: 35.

Nesocyrtosoma serratum Hopp and Ivie, 2009 DOM

Nesocyrtosoma serratum Hopp and Ivie, 2009: 62.

Nesocyrtosoma skelleyi Hopp and Ivie, 2009 DOM

Nesocyrtosoma skelleyi Hopp and Ivie, 2009: 63.

Nesocyrtosoma simplex Hopp and Ivie, 2009 HAI DOM

Nesocyrtosoma simplex Hopp and Ivie, 2009: 37.

Nesocyrtosoma teresitae Hopp and Ivie, 2009 CUB

Nesocyrtosoma teresitae Hopp and Ivie, 2009: 67.

Nesocyrtosoma tumefactum (Marcuzzi, 1976) CUB

Cyrtosoma tumefactum Marcuzzi, 1976: 138.

Cnodalon tumefactum Zayas, 1988: 95 [junior secondary homonym of Cyrtosoma tumefactum Marcuzzi, 1976]. Synonymy: Garrido and Gutiérrez (1996c: 282).

Cnodalon inflatum Zayas, 1988: 101 [junior secondary homonym of Cyrtosoma inflatum Marcuzzi, 1976]. Synonymy: Hopp and Ivie (2009: 24).

Cyrtosoma zayasi Marcuzzi, 1998a: 160. Replacement name for Cyrtosoma tumefactum (Zayas, 1988).

Cyrtosoma gundlachi Marcuzzi, 1998a: 160. Replacement name for Cyrtosoma inflatum (Zayas, 1988).

Nesocyrtosoma turquinense (Zayas, 1988) CUB

Cnodalon turquinensis Zayas, 1988: 96.

Nesocyrtosoma virens (Laporte and Brullé, 1831) CUB PRI

Platydema virens Laporte and Brullé, 1831: 391.

Hoplocephala flavicornis Chevrolat, 1877a: 170. Synonymy: Chevrolat (1878d: 243).

Diaperis viridula Zayas, 1988: 92. Synonymy: Hopp and Ivie (2009: 71).

Genus Nuptis Motschulsky, 1872 [M]

Nuptis Motschulsky, 1872: 25. Type species: Nuptis tenuis Motschulsky, 1872, original designation.

Nuptis caliginosus Champion, 1885MEX (VE YU)

Nuptis caliginosus Champion, 1885: 109.

Nuptis cornutus Champion, 1885GUANICCRIPAN / SA

Nuptis cornutus Champion, 1885: 108.

Nuptis corticalis Champion, 1885 NIC PAN

Nuptis corticalis Champion, 1885: 110.

Nuptis inquinatus Champion, 1885MEX (CI JA) GUANIC

Nuptis inquinatus Champion, 1885: 109.

Nuptis laticollis Champion, 1892 PAN

Nuptis laticollis Champion, 1892: 523.

Nuptis tenebrosus Champion, 1885MEX (VE) GUAPAN

Nuptis tenebrosus Champion, 1885: 110.

Nuptis tenuis Motschulsky, 1872 NIC

Nuptis tenuis Motschulsky, 1872: 32.

Nuptis validus Champion, 1885MEX (VE) GUA

Nuptis validus Champion, 1885: 110.

Genus Oeatus Champion, 1885 [M]

Oeatus Champion, 1885: 111. Type species: Oeatus chevrolati Champion, 1885, subsequent designation (Gebien 1941: 342).

Oeatus chevrolati Champion, 1885MEX (VE) GUABEL

Oeatus chevrolati Champion, 1885: 111.

Oeatus similis Champion, 1892MEX (CI OA) GUABELCRI

Oeatus similis Champion, 1892: 523.

Genus Oenopion Champion, 1885 [M]

Oenopion Champion, 1885: 98. Type species: Oenopion gibbosus Champion, 1885, monotypy.

Oenopion adeptus Doyen, 1971MEX (NL PU)

Oenopion adeptus Doyen, 1971: 114.

Oenopion gibbosus Champion, 1885MEX (VE)

Oenopion gibbosus Champion, 1885: 99.

Oenopion zopheroides (Horn, 1874)USA (NM TX) MEX (SL)

Iphthimus zopheroides Horn, 1874a: 34.

Genus Othryoneus Champion, 1886 [M]

Othryoneus Champion, 1886: 245. Type species: Othryoneus erotyloides Champion, 1886, subsequent designation (Gebien 1942: 315).

Gaurobates Gebien, 1928b: 184. Type species: Gaurobates pictus Gebien, 1928, monotypy. Synonymy: Ferrer (2010: 82).

Othryoneus erotyloides Champion, 1886 NIC

Othryoneus erotyloides Champion, 1886: 246.

Othryoneus triplehorni Ferrer and Ødegaard, 2005 PAN

Othryoneus triplehorni Ferrer and Ødegaard, 2005: 637.

Genus Oxidates Champion, 1886 [M]

Oxidates Champion, 1886: 263. Type species: Oxidates planicollis Champion, 1886, subsequent designation (Gebien 1943: 402).

Oxidates aurichalceus Champion, 1887 MEX

Oxidates aurichalceus Champion, 1887: 265.

Oxidates elongatus Champion, 1893MEX (GE)

Oxidates elongatus Champion, 1893a: 548.

Oxidates gibbus Champion, 1893MEX (VE)

Oxidates gibbus Champion, 1893a: 548.

Oxidates gravidus (Brême, 1842)MEX (VE)

Sphoerotus gravidus Brême, 1842: 109.

Oxidates mexicanus (Brême, 1842)MEX (VE)

Sphoerotus mexicanus Brême, 1842: 110.

Oxidates planicollis Champion, 1886MEX (VE)

Oxidates planicollis Champion, 1886: 264.

Oxidates princeps Champion, 1887MEX (OA VE)

Oxidates princeps Champion, 1887: 265.

Oxidates puncticeps Champion, 1887MEX (VE)

Oxidates puncticeps Champion, 1887: 266.

Oxidates thoracicus (Brême, 1842)MEX (VE)

Sphoerotus thoracicus Brême, 1842: 110.

Genus Polopinus Casey, 1924 [M]

Polopinus Casey, 1924: 326. Type species: Polypleurus nitidus LeConte, 1866, original designation.

Polopinus hubbelli Kritsky, 1989USA (FL)

Polopinus hubbelli Kritsky, 1989: 132.

Polopinus ingens Casey, 1924USA (FL GA)

Polopinus ingens Casey, 1924: 327.

Polopinus nitidus (LeConte, 1866)USA (FL)

Polypleurus nitidus LeConte, 1866b: 118.

Polopinus nitidus subdepressus Casey, 1924: 327. Synonymy: Kritsky (1989: 128).

Polopinus nitidus brevior Casey, 1924: 327. Synonymy: Kritsky (1989: 128).

Polopinus youngi Kritsky, 1989USA (FL)

Polopinus youngi Kritsky, 1989: 130.

Genus Polypleurus Eschscholtz, 1831 [M]

Polypleurus Eschscholtz, 1831: 10, 11. Type species: Polypleurus geminatus Eschscholtz, 1831, monotypy.

Polypleurus geminatus Eschscholtz, 1831USA (FL GA)

Polypleurus geminatus Eschscholtz, 1831: 11.

Polypleurus perforatus (Germar, 1823)USA (AL AR FL GA IL LA MD MO MS NC NJ OK PA SC TX VA WV)

Upis perforata Germar, 1823: 148.

Polypleurus punctatus Solier, 1838: 197. Synonymy: LeConte (1866a: 61).

Genus Saziches Champion, 1886 [M]

Saziches Champion, 1886: 261. Type species: Saziches subcaudatus Champion, 1886, monotypy.

Saziches giesberti Doyen, 1988 CRI

Saziches giesberti Doyen, 1988: 310.

Saziches subcaudatus Champion, 1886 GUA

Saziches subcaudatus Champion, 1886: 262.

Genus Sthenoboea Champion, 1885 [F]

Sthenoboea Champion, 1885: 112. Type species: Sthenoboea apicalis Champion, 1885, monotypy.

Sthenoboea apicalis Champion, 1885 MEX

Sthenoboea apicalis Champion, 1885: 113.

Genus Upis Fabricius, 1792 [M]

Upis Fabricius, 1792b: 515. Type species: Attelabus ceramboides Linnaeus, 1758, monotypy.

Upis ceramboides (Linnaeus, 1758) [Fig. 41] CAN (AB BC MB NB NF NS NT ON QC PE SK YT) USA (AK ID ME MI NH NY OH OR PA SD VT WA WI WY) – Holarctic

Figure 41. 

Upis ceramboides (Linnaeus, 1758). Scale bar = 1 mm.

Attelabus ceramboides Linnaeus, 1758: 388.

Tenebrio variolosus DeGeer, 1775: 32. Synonymy: DeGeer (1775: 32).

Tenebrio reticulatus Say, 1824b: 279. Synonymy: Dejean (1834: 204).

Genus Xenius Champion, 1886 [M]

Xenius Champion, 1886: 224. Type species: Xenius scabripennis Champion, 1886, monotypy.

Xenius scabripennis Champion, 1886 NIC PAN

Xenius scabripennis Champion, 1886: 224.

Genus Xylopinus LeConte, 1862 [M]

Xylopinus LeConte, 1862a: 230. Type species: Tenebrio anthracinus Knoch, 1801 (= Tenebrio saperdoides Olivier, 1795), subsequent designation (Gebien 1941: 336).

Taenobates Motschulsky, 1872: 25. Type species: Tenebrio saperdoides Olivier, 1795, original designation. Synonymy: C.O. Waterhouse (1876: 288).

Xylopinus aenescens LeConte, 1866CAN (NB ON QC) USA (AL CT DC FL GA IA IN LA MA MD MI NC NJ NY OH PA RI SC TN VA WI)

Xylopinus aenescens LeConte, 1866b: 120.

Xylopinus saperdoides (Olivier, 1795)CAN (NB NS ON QC) USA (AL AR CT DC DE FL GA IA IL IN KS LA MD MI MN MO MS NC NH NY OH OK PA SC TN TX VA VT WI WV)

Tenebrio saperdoides Olivier, 1795: [57] 11.

Helops spinipes Fabricius, 1798: 53. Synonymy: Illiger (1802: 344).

Tenebrio anthracinus Knoch, 1801: 169. Synonymy (with H. spinipes Fabricius): Illiger (1802: 344).

Tenebrio rufipes Say, 1825: 203. Synonymy: Melsheimer (1853: 139).

Tribe Stenochiini Kirby, 1837

Stenochiadae Kirby, 1837: 238. Type genus: Stenochia Kirby, 1819 (= Strongylium Kirby, 1819).

Strongyliides Lacordaire, 1859: 478. Type genus: Strongylium Kirby, 1819.

Genus Cuphotes Champion, 1887 [M]

Spheniscus Kirby, 1819: 421 [junior homonym of Spheniscus Moehring, 1758]. Type species: Spheniscus erotyloides Kirby, 1819, monotypy.

Cuphotes Champion, 1887: 332. Replacement name for Spheniscus Kirby, 1819.

Phygoscotus Schulz, 1902: 134. Replacement name for Spheniscus Kirby, 1819.

Cuphotes cinctus (Olivier, 1795)NICCRIPAN / SA

Helops cinctus Olivier, 1795 [58]: 13.

Erotylus unifasciatus Fabricius, 1798: 101. Synonymy: Chevrolat (1843: 80).

Spheniscus 4maculatus Erichson, 1847: 120. Synonymy: Champion (1887: 334).

Spheniscus 4-plagiatus Kirsch, 1866: 202. Synonymy: Champion (1887: 334).

Cuphotes corallifer (J. Thomson, 1859)PAN / SA

Spheniscus corallifer J. Thomson, 1859: 108.

Cuphotes elongatus (J. Thomson, 1859)NICPAN / SA

Spheniscus elongatus J. Thomson, 1859: 112.

Cuphotes jansoni Champion, 1887 NIC

Cuphotes jansoni Champion, 1887: 333.

Cuphotes multimaculatus Pic, 1918 CRI

Cuphotes multimaculatus Pic, 1918b: 7.

Cuphotes nigromaculatus marginicollis (J. Thomson, 1859) MEX GUA

Spheniscus marginicollis J. Thomson, 1859: 110.

Cuphotes nigromaculatus nigromaculatus (J. Thomson, 1859)MEX (JA VE) GUANICCRIPAN

Spheniscus nigro-maculatus J. Thomson, 1859: 110.

Cuphotes unicolor Champion, 1887 NIC

Cuphotes unicolor Champion, 1887: 334.

Genus Mentes Champion, 1893 [M]

Mentes Champion, 1893a: 559. Type species: Mentes ruficollis Champion, 1893, subsequent designation (Lucas 1920: 404).

Mentes aeneopiceus Champion, 1896 LAN

Mentes aeneopiceus Champion, 1896: 30.

Mentes cisteloides Doyen, 1990MEX (JA)

Mentes cisteloides Doyen, 1990: 254.

Mentes fusiformis Champion, 1893 GUA

Mentes fusiformis Champion, 1893a: 560.

Mentes ruficollis Champion, 1893 PAN

Mentes ruficollis Champion, 1893a: 559.

Mentes setipennis Champion, 1893 GUA

Mentes setipennis Champion, 1893a: 560.

Genus Oploptera Chevrolat, 1844 [F]

Oploptera Chevrolat [in Guérin-Méneville], 1844: 126. Type species: Strongylium serraticorne Guérin-Méneville, 1834, monotypy.

Otocerus Mäklin, 1867: 484. Unnecessary replacement name for Oploptera Chevrolat, 1844.

Hoploptera Gemminger [in Gemminger and Harold], 1870: 2037. Unjustified emendation of Oploptera Chevrolat, 1844, not in prevailing usage.

Subgenus Oploptera Chevrolat, 1844 [F]

Oploptera Chevrolat [in Guérin-Méneville], 1844: 126. Type species: Strongylium serraticorne Guérin-Méneville, 1834, monotypy.

Oploptera angelicae (Ferrer and Ødegaard, 2005) PAN

Otocerus angelicae Ferrer and Ødegaard, 2005: 644.

Oploptera chamelensis (Doyen, 1990)MEX (JA)

Otocerus chamelensis Doyen, 1990: 256.

Oploptera delicata (Ferrer and Ødegaard, 2005) PAN

Otocerus delicatus Ferrer and Ødegaard, 2005: 649.

Oploptera dilaticornis (Champion, 1888) PAN

Otocerus dilaticornis Champion, 1888: 378.

Oploptera hamata (Champion, 1888) NIC

Otocerus hamatus Champion, 1888: 381.

Oploptera interrupta (Champion, 1888) PAN

Otocerus interruptus Champion, 1888: 380.

Oploptera microps (Champion, 1888) NIC

Otocerus microps Champion, 1888: 381.

Oploptera nicaraguensis (Champion, 1888) NIC

Otocerus nicaraguensis Champion, 1888: 379.

Oploptera torolae (Champion, 1888) GUA

Otocerus torolae Champion, 1888: 378.

Subgenus Plicatocerus Pic, 1918

Plicatocerus Pic, 1918b: 11. Type species: Otocerus impressipennis Champion, 1888, monotypy.

Oploptera impressipennis (Champion, 1888) PAN

Otocerus impressipennis Champion, 1888: 382.

Genus Poecilesthus Dejean, 1834 [M]

Poecilesthus Dejean, 1834: 207. Type species: Erotylus fasciatus Fabricius, 1781, subsequent designation (Hope 1841: 133).

Diestica Pascoe, 1868: xii. Type species: Diestica viridipennis Pascoe, 1868, monotypy. Synonymy: Gebien (1911b: 589).

Poecilesthus cupripennis Champion, 1893 PAN

Poecilesthus cupripennis Champion, 1893a: 562.

Poecilesthus fragilicornis Champion, 1887 CRI PAN

Poecilesthus fragilicornis Champion, 1887: 338.

Poecilesthus guatemalensis Champion, 1887 GUA

Poecilesthus guatemalensis Champion, 1887: 339.

Poecilesthus immaculatus Champion, 1887 PAN

Poecilesthus immaculatus Champion, 1887: 340.

Poecilesthus laeviceps Champion, 1887 PAN

Poecilesthus laeviceps Champion, 1887: 340.

Poecilesthus laticollis Champion, 1887MEX (CI VE) GUA

Poecilesthus laticollis Champion, 1887: 339.

Poecilesthus latus Champion, 1887 NIC PAN

Poecilesthus latus Champion, 1887: 338.

Poecilesthus maklini Champion, 1887 GUA

Poecilesthus mäklini Champion, 1887: 341.

Poecilesthus nigropunctatus Champion, 1887MEXPAN / SA

Poecilesthus nigro-punctatus Champion, 1887: 336.

Poecilesthus variipes Champion, 1887 NIC PAN

Poecilesthus variipes Champion, 1887: 337.

Genus Pseudotocerus Champion, 1888 [M]

Pseudotocerus Champion, 1888: 383. Type species: Stenochia longipes Lucas, 1859, subsequent designation (Gebien 1948: 542).

Pseudotocerus attenuatus Champion, 1888 NIC

Pseudotocerus attenuatus Champion, 1888: 38384.

Genus Strongylium Kirby, 1819 [N]

Strongylium Kirby, 1819: 417. Type species: Strongylium chalconatum Kirby, 1819, monotypy.

Stenochia Kirby, 1819: 423. Type species: Stenochia rufipes Kirby, 1819, subsequent designation (Hope 1841: 133). Synonymy: Latreille (1829b: 683).

Gentinadis Laporte, 1840: 240. Type species: Stenochia caerulea Laporte, 1840 (= Helops azureus Germar, 1823), monotypy. Synonymy: Lacordaire (1859: 484).

Saerangodes Sturm, 1843: 163. Type species: Helops interpunctatus Germar, 1823, monotypy. Synonymy (with Stenochia Kirby): Blanchard (1845: 33).

Reminius Casey, 1924: 321. Type species: Reminius ocularis Casey, 1924 (= Tenebrio terminatus Say, 1824), original designation. Synonymy: Spilman (1959: 63).

Strongylium acraeum Garrido and Armas, 2012 PRI

Strongylium acraeum Garrido and Armas, 2012b: 76.

Strongylium amethystinum (Guérin-Méneville, 1838) CUB

Stenochia amethystina Guérin-Méneville [in Guérin-Méneville and Chevrolat], 1838: 281.

Strongylium angustulum Mäklin, 1867PAN / SA

Strongylium angustulum Mäklin, 1867: 314.

Strongylium antennale Mäklin, 1867 CUB

Strongylium antennale Mäklin, 1867: 270.

Strongylium anthrax Schwarz, 1878USA (FL)

Strongylium anthrax Schwarz, 1878: 369.

Strongylium apache Triplehorn and Spilman, 1973USA (AZ NM) MEX [SO]

Strongylium apache Triplehorn and Spilman, 1973: 10.

Strongylium apicicorne Mäklin, 1867MEX (VE)

Strongylium apicicorne Mäklin, 1867: 324.

Strongylium armatum Mäklin, 1867MEX (CI VE) GUAPAN

Strongylium armatum Mäklin, 1867: 311.

Strongylium atrum Champion, 1888USA (AZ NM) MEX (CH DU SI)

Strongylium atrum Champion, 1888: 360.

Strongylium aulicum Mäklin, 1867USA (FL TX) MEX (OA VE) GUANICPAN

Strongylium aulicum Mäklin, 1867: 363.

Strongylium auratum (Laporte, 1840)MEX (CI GE PU VE YU) GUABELNICCRIPAN / SA

Stenochia aurata Laporte, 1840: 240.

Stenochia auratum var. hilaris Mäklin, 1867: 402. Synonymy: Champion (1888: 360).

Strongylium azureum (Germar, 1823)CUB / SA

Helops azureus Germar, 1823: 153.

Stenochia caerulea Laporte, 1840: 240. Synonymy: Mäklin (1867: 404).

Strongylium baetianum Garrido and Armas, 2012 HAI

Strongylium baetianum Garrido and Armas, 2012a: 64.

Strongylium basiclavis Zayas, 1988 CUB

Strongylium basiclavis Zayas, 1988: 110.

Strongylium belti Champion, 1888 NIC

Strongylium belti Champion, 1888: 358.

Strongylium bivittatum Champion, 1888MEX (OA)

Strongylium bivittatum Champion, 1888: 361.

Strongylium blandum Mäklin, 1867MEX (VE)

Strongylium blandum Mäklin, 1867: 341.

Strongylium brevipes Champion, 1888 NIC PAN

Strongylium brevipes Champion, 1888: 372.

Strongylium canaliculatum Champion, 1887 MEX GUA

Strongylium canaliculatum Champion, 1887: 346.

Strongylium cancellatum Mäklin, 1867MEX (VE) BEL

Strongylium cancellatum Mäklin, 1867: 320.

Strongylium carinipenne Champion, 1888 PAN

Strongylium carinipenne Champion, 1888: 374.

Strongylium chalcopterum Mäklin, 1867LAN (Martinique)

Strongylium chalcopterum Mäklin, 1867: 431.

Strongylium championi Gebien, 1948USA (TX) MEX (JA VE) GUABEL

Strongylium varians Champion, 1888: 365 [junior secondary homonym of Strongylium varians (Pascoe, 1883)].

Strongylium championi Gebien, 1948: 532. Replacement name for Strongylium varians Champion, 1888.

Strongylium chiriquense Champion, 1887 PAN

Strongylium chiriquense Champion, 1887: 351.

Strongylium chontalense Champion, 1887 NIC

Strongylium chontalense Champion, 1887: 344.

Strongylium cinctum Mäklin, 1867MEX (VE)

Strongylium cinctum Mäklin, 1867: 337.

Strongylium clavicorne Champion, 1893MEX (VE)

Strongylium clavicorne Champion, 1893a: 562.

Strongylium colombianum Champion, 1888PAN / SA

Strongylium colombianum Champion, 1888: 354.

Strongylium conicicolle Mäklin, 1867 NIC CRI PAN

Strongylium conicicolle Mäklin, 1867: 447.

Strongylium conradti Champion, 1893 GUA

Strongylium conradti Champion, 1893a: 563.

Strongylium costaricense Champion, 1888 CRI

Strongylium costaricense Champion, 1888: 353.

Strongylium crassicorne Champion, 1887 NIC

Strongylium crassicorne Champion, 1887: 347.

Strongylium crenatum Mäklin, 1867USA (AL AR FL GA IA KS LA MD MO MS NC OH OK SC TN TX VA)

Strongylium crenatum Mäklin, 1867: 307.

Strongylium cribripes Mäklin, 1867MEX (VE) NICPAN

Strongylium cribripes Mäklin, 1867: 275.

Strongylium cruentatum Mäklin, 1867MEX (VE)

Strongylium cruentatum Mäklin, 1867: 335.

Strongylium cultellatum Mäklin, 1867USA (FL) – Adventive

Strongylium cultellatum Mäklin, 1867: 453.

Strongylium cupeyal Zayas, 1988 CUB

Strongylium cupeyal Zayas, 1988: 110.

Strongylium cuproso Garrido, 2004 CUB

Strongylium cuproso Garrido, 2004d: 52.

Strongylium curticorne Champion, 1888MEX (CI)

Strongylium curticorne Champion, 1888: 369.

Strongylium decoratum Mäklin, 1867NICCRIPAN / SA

Strongylium decoratum Mäklin, 1867: 365.

Strongylium delauneyi Fleutiaux and Sallé, 1890 LAN

Strongylium delauneyi Fleutiaux and Sallé, 1890: 429.

Strongylium dentatum Champion, 1887 NIC

Strongylium dentatum Champion, 1887: 348.

Strongylium discoidale Mäklin, 1867MEX (VE)

Strongylium discoidale Mäklin, 1867: 339.

Strongylium elongatum Garrido and Armas, 2012 DOM

Strongylium elongatum Garrido and Armas, 2012a: 65.

Strongylium eminens Mäklin, 1867MEX (JA VE)

Strongylium eminens Mäklin, 1867: 374.

Strongylium erraticum Champion, 1888 NIC

Strongylium erraticum Champion, 1888: 373.

Strongylium exaratum Champion, 1887 GUA PAN

Strongylium exaratum Champion, 1887: 350.

Strongylium excavatum Mäklin, 1867MEX (OA VE) GUANICPAN

Strongylium excavatum Mäklin, 1867: 274.

Strongylium eximium Mäklin, 1867 CUB

Strongylium eximium Mäklin, 1867: 269.

Strongylium fossifrons Mäklin, 1867PAN / SA

Strongylium fossifrons Mäklin, 1867: 285.

Strongylium fragile Champion, 1888 PAN

Strongylium fragile Champion, 1888: 377.

Strongylium frontale Champion, 1888 PAN

Strongylium frontale Champion, 1888: 357.

Strongylium funestum Mäklin, 1867 MEX

Strongylium funestum Mäklin, 1867: 295.

Strongylium gerstaeckeri Mäklin, 1867MEX (VE) GUANICCRIPAN

Strongylium gerstaeckeri Mäklin, 1867: 277.

Strongylium gibbum Mäklin, 1867 MEX GUA

Strongylium gibbum Mäklin, 1867: 252.

Strongylium gregarium Champion, 1888 PAN

Strongylium gregarium Champion, 1888: 373.

Strongylium guadeloupense Gebien, 1911LAN (Guadeloupe)

Strongylium inaequale Fleutiaux and Sallé, 1890: 430 [junior primary homonym of Strongylium inaequale Mäklin, 1867].

Strongylium guadeloupense Gebien, 1911b: 596. Replacement name for Strongylium inaequale Fleutiaux and Sallé, 1890.

Strongylium hemistriatum Triplehorn and Spilman, 1973USA (TX)

Strongylium hemistriatum Triplehorn and Spilman, 1973: 20.

Strongylium hoepfneri chevrolatii Mäklin, 1867 MEX

Strongylium chevrolatii Mäklin, 1867: 235.

Strongylium hoepfneri hoepfneri Mäklin, 1867MEX (VE) GUA

Strongylium hoepfneri Mäklin, 1867: 232.

Strongylium hoepfneri immundum Mäklin, 1867 MEX

Strongylium immundum Mäklin, 1867: 234.

Strongylium hoepfneri pectorale Mäklin, 1867 MEX NIC

Strongylium pectorale Mäklin, 1867: 233.

Strongylium hoepfneri scutellare Mäklin, 1867 MEX

Strongylium scutellare Mäklin, 1867: 233.

Strongylium ignitum Champion, 1887 NIC PAN

Strongylium ignitum Champion, 1887: 348.

Strongylium impressicolle Mäklin, 1867MEX (DU JA VE) GUANICCRI

Strongylium impressicolle Mäklin, 1867: 301.

Strongylium languidum Mäklin, 1867MEX (VE) GUA

Strongylium languidum Mäklin, 1867: 312.

Strongylium langurioides Champion, 1888 NIC

Strongylium langurioides Champion, 1888: 355.

Strongylium laterale Mäklin, 1867MEX (VE)

Strongylium laterale Mäklin, 1867: 334.

Strongylium limitatum Mäklin, 1867MEX (VE)

Strongylium limitatum Mäklin, 1867: 342.

Strongylium lucidum Mäklin, 1867 CRI

Strongylium lucidum Mäklin, 1867: 283.

Strongylium maculicolle Champion, 1887 NIC CRI PAN

Strongylium maculicolle Champion, 1887: 342.

Strongylium maisi Garrido, 2004 CUB

Strongylium maisi Garrido, 2004d: 52.

Strongylium marginale Mäklin, 1867MEX (VE)

Strongylium marginale Mäklin, 1867: 338.

Strongylium misantlae Champion, 1888MEX (OA VE) GUA

Strongylium misantlae Champion, 1888: 367.

Strongylium montebarreto Garrido, 2004 CUB

Strongylium montebarreto Garrido, 2004d: 50.

Strongylium nigrum Zayas, 1988 CUB

Strongylium nigra Zayas, 1988: 110.

Strongylium nitidiceps Champion, 1888MEX (VE)

Strongylium nitidiceps Champion, 1888: 364.

Strongylium nubeculosum Mäklin, 1867MEX (YU) NIC

Strongylium nubeculosum Mäklin, 1867: 336.

Strongylium oculatum Champion, 1888 GUA NIC

Strongylium oculatum Champion, 1888: 371.

Strongylium opacipenne Champion, 1888MEX (VE)

Strongylium opacipenne Champion, 1888: 361.

Strongylium paddai Ivie and Triplehorn, 1986 VIS

Strongylium paddai Ivie and Triplehorn, 1986: 423.

Strongylium panamense Champion, 1888 PAN

Strongylium panamense Champion, 1888: 363.

Strongylium permodicum Mäklin, 1867GUANICPAN / LAN / SA

Strongylium permodicum Mäklin, 1867: 320.

Strongylium preciosus Zayas, 198885 CUB

Strongylium preciosus Zayas, 1988: 108.

Strongylium pulvinatum Mäklin, 1867 PRI

Strongylium pulvinatum Mäklin, 1867: 265.

Strongylium pumilum Garrido and Armas, 2012 PRI

Strongylium pumilum Garrido and Armas, 2012b: 73.

Strongylium punctifrons Mäklin, 1867MEX (CI VE) BEL

Strongylium punctifrons Mäklin, 1867: 296.

Strongylium punctipes Champion, 1888MEX (JA) GUA

Strongylium punctipes Champion, 1888: 375.

Strongylium quisqueyanum Garrido and Armas, 2012 DOM

Strongylium quisqueyanum Garrido and Armas, 2012a: 66.

Strongylium ramosum Mäklin, 1867MEX (VE)

Strongylium ramosum Mäklin, 1867: 340.

Strongylium sallei Mäklin, 1867MEX (VE) GUANIC

Strongylium sallei Mäklin, 1867: 257.

Strongylium sallaei Champion, 1887: 345. Unjustified emendation of Strongylium sallei Mäklin, 1867, not in prevailing usage.

Strongylium semistriatum Mäklin, 1867MEX (VE)

Strongylium semistriatum Mäklin, 1867: 251.

Strongylium simplicipes Pic, 1918 PAN

Strongylium simplicipes Pic, 1918b: 15.

Strongylium simplicicolle LeConte, 1878USA (AL FL GA MS NC SC TN VA)

Strongylium simplicicolle LeConte, 1878a: 424.

Strongylium subcostatum Mäklin, 1867 MEX GUA

Strongylium subcostatum Mäklin, 1867: 316.

Strongylium suturale Mäklin, 1867MEX (VE) GUA

Strongylium suturale Mäklin, 1867: 337.

Strongylium tenuicolle (Say, 1826) [Fig. 42] CAN (MB ON QC) USA (AL AR CO CT DC DE FL GA IA IL IN KS KY LA MA MD MI MN MO MS NC NH NJ NY OH OK PA RI SC SD TN TX VA WI WV)

Figure 42. 

Strongylium tenuicolle (Say, 1826). Scale bar = 1 mm.

Helops tenuicollis Say, 1826: 241.

Strongylium terminatum (Say, 1824)USA (AL DC FL IA IL IN KS KY LA MD MI MO MS NC NE NJ NY OH OK PA SC SD TX VA WI)

Tenebrio terminatus Say, 1824a: 267.

Reminius ocularis Casey, 1924: 322. Synonymy: Spilman (1959: 63).

Strongylium tinctipes Champion, 1887 NIC PAN

Strongylium tinctipes Champion, 1887: 349.

Strongylium turquinense Zayas, 1988 CUB

Strongylium turquinensis Zayas, 1988: 107.

Strongylium variicorne Champion, 1887 PAN

Strongylium variicorne Champion, 1887: 352.

Strongylium ventrale Champion, 1888 PAN

Strongylium ventrale Champion, 1888: 356.

Strongylium venustum Zayas, 1988 CUB

Strongylium venusta Zayas, 1988: 109.

Strongylium verde Garrido and Armas, 2012 PRI

Strongylium verde Garrido and Armas, 2012b: 74.

Strongylium vikenae Ferrer and Ødegaard, 2005 PAN

Strongylium vikenae Ferrer and Ødegaard, 2005: 644.

Strongylium virescens Zayas, 1988 CUB

Strongylium virescens Zayas, 1988: 110.

Strongylium viridipes Mäklin, 1867MEX (PU VE)

Strongylium viridipes Mäklin, 1867: 274.

Strongylium viriditinctum Champion, 1888 GUA

Strongylium viriditinctum Champion, 1888: 359.

Strongylium woodruffi Garrido and Armas, 2012 DOM

Strongylium woodruffi Garrido and Armas, 2012a: 67.

Tribe Talanini Champion, 1887

Dignamptini LeConte and Horn, 1883: 385. Type genus: Dignamptus LeConte, 1878 (= Talanus Jacquelin du Val, 1857).

Talanides Champion, 1887: 321. Type genus: Talanus Jacquelin du Val, 1857. Note. Family-group name conserved over Dignamptini (see Bouchard et al. 2005).

Genus Talanus Jacquelin du Val, 1857 [M]

Talanus Jacquelin du Val, 1857: 156. Type species: Talanus cribrarius Jacquelin du Val, 1857, monotypy.

Dignamptus LeConte, 1878a: 421. Type species: Dignamptus stenochinus LeConte, 1878, present designation. Synonymy: Champion (1887: 321).

Talanus aeneipennis Champion, 1887MEX (TB VE) BEL

Talanus aeneipennis Champion, 1887: 327.

Talanus apterus Champion, 1887 GUA PAN

Talanus apterus Champion, 1887: 328.

Talanus ater Champion, 1887 PAN

Talanus ater Champion, 1887: 327.

Talanus columbianus Mäklin, 1878PAN / SA

Talanus columbianus Mäklin, 1878a: 99.

Talanus cribrarius Jacquelin du Val, 1857 CUB

Talanus cribrarius Jacquelin du Val, 1857: 156.

Talanus ferrugineus Champion, 1896 LAN

Talanus ferrugineus Champion, 1896: 31.

Talanus guadeloupensis Fleutiaux and Sallé, 1890 LAN

Talanus guadeloupensis Fleutiaux and Sallé, 1890: 430.

Talanus guatemalensis Champion, 1887 GUA

Talanus guatemalensis Champion, 1887: 326.

Talanus insularis Mäklin, 1878 PRI LAN

Talanus insularis Mäklin, 1878a: 98.

Talanus interstitialis Champion, 1887 ME (CI JA) GUANIC

Talanus interstitialis Champion, 1887: 324.

Talanus laevicollis Champion, 1896 LAN

Talanus laevicollis Champion, 1896: 32.

Talanus laevipennis Champion, 1887 GUA

Talanus laevipennis Champion, 1887: 322.

Talanus langurinus (LeConte, 1878)USA (FL TX)

Dignamptus langurinus LeConte, 1878a: 421.

Talanus laticeps Champion, 1887 PAN

Talanus laticeps Champion, 1887: 325.

Talanus lecontei Champion, 1887MEX (TB VE YU) GUABEL

Talanus lecontei Champion, 1887: 323.

Talanus longicornis Champion, 1887 GUA BEL NIC PAN

Talanus longicornis Champion, 1887: 328.

Talanus mecoscelis Triplehorn, 1968USA (TX)

Talanus mecoscelis Triplehorn, 1968a: 33.

Talanus neotropicalis Champion, 1887MEX (JA VE YU) GUACRIPAN / SA

Talanus neotropicalis Champion, 1887: 322.

Talanus spilmani Triplehorn, 1968USA (FL)

Talanus spilmani Triplehorn, 1968a: 35.

Talanus stenochinus (LeConte, 1878)USA (FL LA)

Dignamptus stenochinus LeConte, 1878a: 421.

Talanus okeechobensis Blatchley, 1914: 143. Synonymy: Fall (1932: 148).

Talanus subexaratus Mäklin, 1878MEX (CI GE SI VE) GUABELNICCRIPAN / SA

Talanus subexaratus Mäklin, 1878a: 102.

Talanus subopacus Champion, 1887MEX (VE) BEL

Talanus subopacus Champion, 1887: 323.

Talanus victori Garrido and Gutiérrez, 2004 PRI

Talanus victori Garrido and Gutiérrez, 2004: 63.

Incertae sedis: Tenebrionidae

Tenebrio calculensis Scudder, 1895CAN (ON)

Tenebrio calculensis Scudder, 1895: 3186.

Acknowledgements

We thank Mike Ivie (Montana State University, Bozeman, Montana, USA) for making available rare literature and for comments on the biogeography of the West Indies. Miguel A. Alonso-Zarazaga (Museo Nacional de Ciencias Naturales, Madrid, Spain) contributed important nomenclatural insight and literature. Go Sato and Roelof (Ralf) Idema (Agriculture and Agri-Food Canada, Ottawa, Canada) provided the habitus drawings. Anthony Davies (Agriculture and Agri-Food Canada, Ottawa, Canada) assisted in thoroughly reviewing the catalogue dataset and J. Milton Campbell (Agriculture and Agri-Food Canada, Ottawa, Canada) reviewed the contents for the subfamily Alleculinae. Miguel A. Alonso-Zarazaga, Gustavo E. Flores (Instituto Argentino de Investigaciones de las Zonas Áridas [IADIZA], Mendoza, Argentina), and Otto Merkl (Hungarian Natural History Museum, Budapest, Hungary) reviewed the entire manuscript and provided useful corrections and suggestions. Funding for this project was provided by the NSF ARTS program (DEB–1523605) to ADS and MAJ.

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  • Triplehorn CA, Moser JC (1970) Two new species of Corticeus from Mexico and Honduras (Coleoptera: Tenebrionidae). The Coleopterists Bulletin 24: 47–50.
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1

1 The term “North America”, used throughout the text, refers to the continent in the Western Hemisphere consisting of Greenland, Canada, United States of America, Mexico, Central America, and the West Indies.

2

2 The name derives from the Greek Lepidos (scale) and the generic name Cnemeplatia Costa. The gender is feminine.

3

3 All geographical units are provided using their English names as listed in the third edition of Merriam-Webster’s Geographical Dictionary (2001).

4

4Adelonia sulcatus used by Leng and Mutchler (1914: 464) and often listed as a synonym of this species is an incorrect subsequent spelling not in prevailing usage.

5

5Perty (1832) used two spellings for this name: Goniadera (p. 62) and Gonyodera (p. 63). Later Perty (1833: 14) used the spelling Goniadera. As so he is deemed to be the First Reviser (ICZN 1999: Article 24.2.4) and Goniadera is the correct original spelling..

6

6 In their revision of the genus, Ferrer and Delatour (2007: 287) reported this species and the type species of the subgenus as “Goniadera tuberculatum Gebien, 1928” and “G. binotatum Gebien, 1928” respectively. Both names are nomina nuda.

7

7 Mälkin was Finnish, not German. Therefore the umlaut is deleted and no “e” is to be inserted after the “a”.

8

8 This genus was transferred from the Mycetophagidae to the Lagriinae by Lawrence and Newton (1995: 886).

9

9Nilio and Nilio lebasi are incorrect subsequent spellings in prevailing usage and attributed to the publication of the original spelling; therefore they are to be preserved and deemed to be the correct original spellings (ICZN 1999: Article 33.3.1).

10

10 The species-group name bifurcus is a name in apposition and therefore need not agree in gender with the generic name (M.A. Alonso-Zarazaga personal communication).

11

11 Described from Chile, which is probably a mistake (Champion 1884: 54).

12

11 A junior secondary homonym replaced before 1961 is permanently invalid unless the substitute name is not in use and the relevant taxa are no longer considered congeneric (ICZN 1999: Article 59.3); in this case, the relevant taxa are no longer considered congeneric and the substitute name has not been used subsequently as valid as far as we know.

13

13 A junior secondary homonym replaced before 1961 is permanently invalid unless the substitute name is not in use and the relevant taxa are no longer considered congeneric (ICZN 1999: Article 59.3); in this case, the relevant taxa are no longer considered congeneric and the substitute name is not currently in use.

14

14 Described from the La Brea tar pits, but not actually a fossil [ADS]

15

15 A junior secondary homonym replaced before 1961 is permanently invalid unless the substitute name is not in use and the relevant taxa are no longer considered congeneric (ICZN 1999: Article 59.3); in this case, the relevant taxa are no longer considered congeneric and the substitute name has not been used subsequently as valid as far as we know.

16

16 Casey’s specimens are mislabelled from Texas.

17

17Branchus cubensis Marcuzzi” has been cited twice by Marcuzzi (1998a: 153) and Peck (2005: 146) but the name was never proposed by anyone and is a nomen nudum.

18

18 Described from La Brea tar pits but thought to represent a modern undescribed species by Doyen and Miller (1980: 3).

19

19Champion (1892: 510, 511) used two different spellings for this species: obliteratus (p. 510) and obsoletus (p. 511). Doyen (1984b: 41) acted as First Reviser (ICZN 1999: Article 24.2) and selected obliteratus as the correct spelling.

20

20 Since the dates of publication of C. asperata (Horn, 1870) and C. asperata (Motschulsky, 1870) could not be determined besides the year, Aalbu et al. (1995: 482) acted as First Revisers (ICZN 1999: Article 24.2) and selected Horn’s name to have precedence. However, evidence gathered recently [YB] indicate that Motschulsky’s paper in the Bulletin de la Société Impériale des Naturalistes de Moscou was published before Horn’s paper in the Transactions of the American Philosophical Society. At this point, we prefer not to make any change.

21

21Stibia opaca credited to Blaisdell (1925b: 329), listed as a synonym of S. cribrata by Papp (1961d: 103), is considered here as a lapsus for S. cribrata Blaisdell.

22

22Telabis is neither Latin or Greek and since it was proposed as a subgenus, it was not placed in agreement with any specific names. Therefore, by application of Article 30.2.4 (ICZN 1999) the name has to be treated as masculine (M.A. Alonso-Zarazaga, personal communication). The fact that Casey subsequently used the name Telabis as feminine is irrelevant.

23

23 The species name “minor” is an adjective in comparative degree and is the masculine and feminine form. The neuter form is “minus” (M.A. Alonso-Zarazaga, personal communication).

24

24 Listed as a synonym of T. laevis LeConte by Horn (1870: 259) and Champion (1884: 2) but treated as valid by Casey (1907: 438).

25

25 The specific name was obviously proposed for Auguste Sallé [b. 1820, d. 1896], who collected some of the original specimens, and should have been spelled “sallei.” Because Champion did not provided the etymology of the name, the original spelling should be maintained and Article 32.5.1 (ICZN 1999) does not apply in this case. However, the spelling “sallei” is in prevailing usage (36 hits in Google Scholar vs 11 hits for “sallaei”) and so deemed to be the original spelling (ICZN 1999, Article 33.3.1).

26

26Gebien (1908a: 287) originally placed this name in synonymy with Nyctoporis cristata Eschscholtz and latter (1910a: 118; 1937: 698) with Nyctoporis carinata LeConte.

27

27 The generic name is spelled “Caribanoisis” in the running title (p. 569). Since the running title was generated by the publisher, it is not considered as a different original spelling and no First Reviser action is required here.

28

28 This Oriental species is known in North America from a single specimen (Steiner and Paraskevoudakis 2014) and whether it is established on this continent remains to be ascertained.

29

29 The names Eleodes chevrolati and Eleodes nitidulithorax used by Pierre (1976: 705, 706) are nomina nuda. The same is true for the name Eleodes dentipes sublitoralis listed by Gebien (1938: 57). The following names, being infrasubspecific, are nomina nuda: Eleodes omissa var. pygmaea forma borealis Blaisdell, 1909: 79; Eleodes extricata forma cognata punctata Blaisdell, 1909: 123; Eleodes letcheri var. vandykei forma parvula Blaisdell, 1909: 137; Eleodes dentipes forma elongata laevis Blaisdell, 1909: 254; Eleodes dentipes forma elongata punctata Blaisdell, 1909: 254; Eleodes acuticauda var. laticollis forma minor Blaisdell, 1909: 283; Eleodes (Eleodes) acuticauda var. laticollis forma insularis Blaisdell, 1909: 284; Eleodes eschscholtzi var. lucae forma ecaudata Blaisdell, 1909: 290; Eleodes eschscholtzi var. lucae forma grandis Blaisdell, 1909: 291; Eleodes eschscholtzi var. lucae forma inflata Blaisdell, 1909: 291. Even though some of these names have been listed as valid subspecies by Papp (1961), article 45.6.4.1 (ICZN 1999) does not apply here and the names are not available.

30

30 This species is usually credited to Eschscholtz (1831: 13) where the species is described. However, in 1829 Eschscholtz described the genus Amphidora and listed by name the sole species (A. littoralis) included in it. As such it is a combined description of a new nominal genus and a single new nominal species which then provides an indication for each name (ICZN 1999: Article 12.2.6). Sherborn (1927: 3618) also credited Eschscholtz (1829) for the availability of A. littoralis.

31

31Blaisdell (1937b) used two different spelling for this taxon: Arpeleodes (p. 128) and Ardeleodes (p. 128, 129). Gebien (1938: 63) acted as First Reviser (ICZN 1999: Article 24.2.3) and selected Ardeleodes as the correct original spelling.

32

32 The species name grutus is neither Latin or Greek but appears to be derived from the Spanish word “gruta (cave)” and transformed as an adjective (M.A. Zlonso-Zarazaga, personal communication). Since it is a creation of the authors, the name must be treated as a noun in apposition.

33

33 This specific name was originally spelled vetorator but veterator is an incorrect subsequent spelling in prevailing usage and attributed to the publication of the original spelling; therefore the subsequent spelling is deemed to be the correct original spelling (ICZN 1999: Article 33.3.1).

34

34Thomas (2005: 553) retained the name transvolcanensis for the species and rufipes for the subspecies. However, under Article 47.2 of the ICZN (1999) the oldest name for the species must be retained.

35

35 The species name anachronus is not a classical Greek or Latin name and seems a creation of Triplehorn (M.A. Alonso-Zarazaga, personal communication). Therefore the name is deemed to be a noun in apposition.

36

36 We consider this name available from Blaisdell (1910: 61) with the description by indication (ICZN 1999, Article 12.2.1) to his monograph of the Eleodiini published in 1909.

37

37 Described by indication.

38

38 An application to the International Commission on Zoological Nomenclature was recently published (Bousquet and Bouchard 2017b) to retain Bolitotherus cornutus (Fabricius, 1801) as the valid name for this taxon.

39

39 As for Amblycyphus asperatus (see footnote #20), evidence gathered recently indicates that Motschulsky's manuscript was published prior to that of Horn and that Threnus has precedence. The Reversal of precedence cannot be used in this case since Threnus was listed as valid after 1899 in some catalogues (e.g., Leng 1920: 224). An application to the Commission will be necessary to retain Horn's name as valid.

40

40 A number of authors, including LeConte (1866a: 61) and Horn (1870: 325), mistakenly used the name sulcipennis for this species.

41

41 Species described from “Californie. St. Francisco.”

42

42Berry (1975: 929) proposed the new subgenus Longicerenopus to include C. angustatus Horn, C. aterrimus Horn, C. hermanus Berry, and C. punctatus Berry. Unfortunately the author failed to designate a type species and therefore his name is unavailable from that date. Staff of the Zoological Society of London (Anonymous 1979: 217) designated C. angustatus Horn as type species but since the person responsible for the nomenclatural act cannot be determined from the work itself, the act must be attributed to “Anonymous.” Names and nomenclatural acts published anonymously after 1950 are unavailable (ICZN 1999: Article 14) and so consequently the name Longicerenopus is still unavailable.

43

43Helops martinensis Marcuzzi, 2001 (p. 252) is a nomen nudum since Marcuzzi (2001) failed to indicate where the syntypes will be (or are) deposited and the name and location of the collection, a mandatory requirement for every new species-group name published after 1999 (ICZN 1999: Article 16.4.2).

44

44 According to M.A. Alonso-Zarazaga (personal communication), Tenebrio caeruleus Linnaeus, 1758, based on a syntype, is a chrysomelid species of the genus Timarcha. This problem will be addressed in a future publication.

45

45Allard (1877a: 36) corrected the name Coscinopter to Coscinoptilix in the errata issued in one part of the same journal volume. Therefore Coscinoptilix is the correct original spelling (ICZN 1999: Article 32.5).

46

46Helops punctatus Fabricius, 1801 (currently Taraxides punctatus) and Helops punctatus Gemminger, 1870 are primary homonyms, both presently used as valid, that have not been considered congeneric after 1899. In such case, the junior homonym is not to be replaced and the case should be referred to the Commission (ICZN 1999: Article 23.9.5).

47

47Horn (1885a: 90) mentioned that this species “is certainly not a Helops” and that he was unable to identify it. He also questioned the fact that the species was of American origin.

48

48Horn (1885a: 89) placed H. americanus Palisot de Beauvois in synonymy with Helops undulatus LeConte based on the original description only. His action was not followed by subsequent authors. Therefore we prefer to continue using LeConte’s name and consider Palisot de Beauvois’ name as a nomen dubium. The reversal of precedence (ICZN 1999: Article 23.9) cannot be used here since Helops americanus has been used as a valid name at least once after 1899 (i.e., Scott and Fiske 1902: 33).

49

49 Removed from synonymy with B. fuscus Casey by Steiner (2006).

50

50Kaszab (1969) used two different spellings for this taxon: kulzrei (p. 318) and kulzeri (pp. 318, 337). Nobody has acted as First Reviser (ICZN 1999: Article 32.2.1) to our knowledge. We act as First Revisers and choose kulzeri (based on H. Kulzer), which was obviously Kaszab’s intention although this was not explicitely mentioned in the text.

51

51Diastolinus bardubensis [sic] guadeloupensis Marcuzzi, 2001 (p. 250) and Diastolinus bardubensis [sic] martiniquensis Marcuzzi, 2001 (p. 250) were placed in synonymy with Blapstinus opacus Mulsant and Rey by Soldati and Touroult (2014: 98) and Ivie and Hart (2016: 466) respectively. However, these two taxa are nomina nuda since Marcuzzi (2001) failed to indicate where the syntypes will be (or are) deposited and the name and location of the collection, a mandatory requirement for every new species-group name published after 1999 (ICZN 1999: Article 16.4.2).

52

52 According to Marcuzzi (2002: 399), the holotype and sole known specimen of this species was probably transported by ship from Cuba to Curaçao.

53

53 This species, originally described from specimen(s) collected in Missouri, has never been interpreted except by Horn (1870: 402) who mentioned that it probably belongs to the genus Opatrinus (North American members of this genus are currently placed in the genus Alaetrinus). The name is tentatively placed here in synonymy with A. minimus (Palisot de Beauvois), a common species.

54

54Ferrer (2006) mentioned that Z. atratus (Fabricius) and morio (Fabricius) are “two forms of the same species” and that specimens of both forms emerge in laboratory from the same parents. Later (Ferrer 2011: 299) considered morio as a distinct species. However based on the data presented in Ferrer’s papers of 2006 and 2011, we believe that specimens of the form morio are probably conspecific with those of atratus. Soldati and Touroult (2014: 103) also considered morio as a synonym of atratus.

55

55 Considered a valid species by Garrido and Gutiérrez (1994b), with Zophobas cubanus Marcuzzi, 1976 listed as a junior synonym. Ferrer (2011: 297) listed Z. rugipes as a junior synonym of Z. atratus (Fabricius).

56

56 Most previous literature records attributed to Wattius cucullatus (Pascoe, 1871) belong to multiple undescribed species [ADS]. Wattius cucullatus is currently known only from Brazil (Smith and Sanchez 2015: 115).

57

57Spilman (1973: 42) mentioned that the Ulosonia tricornis of Laporte (1840) was in fact a reference to Phaleria tricornis Dalman, 1823 with an incorrect author. We believe these two names are different nomenclatural acts though they are probably synonymous as indicated by Spilman.

58

58 The name derives from the Greek root Spalae (cave) and the noun biosis (manner of live) in reference to the cave-inhabiting sole species currently included in the genus.

59

59Eusemostene proposed by Gebien (1940: 764) is regularly listed as a junior synonym of this taxon. Although listed as “nom. nov.” Gebien in fact proposed a new subgeneric taxon by providing a bibliographic reference to Seidlitz’s (1890: 518) misidentification of Stene Stephens, 1829. However, the name is unavailable because Gebien failed to designate a type species for his new subgenus (see ICZN 1999: Article 13.3). The name has not been made available subsequently to our knowledge.

60

60 Reported under the unavailable name Alegoria laportei Fleutiaux and Sallé, 1890 by some authors (e.g., Peck 2011: 30).

61

61 Only one specimen, from Mexico, is known north of Colombia (Champion 1886: 152); possibly the specimen is mislabelled.

62

62Melasia gagatina Perroud and Mulsant, 1861, has been listed from North America (Gebien 1911a: 402) and placed in synonymy with Uloma punctulata LeConte, 1866 (Gebien 1940: 776). In fact, the species was described from Sicily and is a junior synonym of Uloma culinaris (Linnaeus, 1758) (see Löbl et al. 2008: 302).

63

63 This taxon has been listed in synonymy with Tribolium castaneum (Herbst) by some authors. However, the size originally given, 3⅓ lig. (= 7.4 mm) prevent an association with T. castaneum. We concur with Good (1936: 5) that Laporte’s species probably belongs to the genus Uloma Dejean. Unfortunately, based on the short original description, it cannot be confidently associated with any of the species described from North America.

64

64 The name of this species has been credited to Jacquelin du Val (1857: 157) by some authors (e.g., Blackwelder 1945: 506; Campbell 1966: 25; Campbell 1971: 27). However, Jacquelin du Val referred the name to Dejean’s catalogue (1834: 214) who used the replacement name fuscula Schönherr as valid and listed flavipes Fabricius in synonymy. Therefore we consider that Jacquelin du Val did not propose a new species under the name “Allecula flavipes Dejean.”

65

65 The junior and senior homonyms for Cistela brevis apply to taxa not considered congeneric since 1899. In such case, the junior homonym must not automatically be replaced; the case should be referred to the Commission for a ruling and meanwhile prevailing usage of both names is to be maintained (ICZN 1999: Article 23.9.5).

66

66 Although this name is older than Adelinina, Bouchard et al. (2011: 428) recommended that an application be submitted to the Commission to conserve usage of Adelinina LeConte, 1862.

67

67Palisot de Beauvois (1812) used two different spellings for this taxon: Trogossita maxillosa (p. 125) and Trogossita maxillaris (pl. 32, fig. 4). Page 125 and plate 32 were both issued in 1812 in livraison 8 of Palisot de Beauvois’ work. We have not found anyone who acted as First Reviser. We here select T. maxillaris as the correct spelling. This name was used regularly for Palisot de Beauvois’ taxon probably because T. maxillosus was already in use by Fabricius in 1801.

68

68Diaperis hydni is an incorrect subsequent spelling of D. hydactina Fabricius, 1798 introduced by Fabricius (1801b: 585).

69

69LeConte (1870: 404) believed the species was “wrongly attributed to North America.”

70

70LeConte (1870: 404) believed the species was “wrongly attributed to North America.”

71

71LeConte (1870: 404) noted that Platydema pallens “is found in South America, and must therefore be erased from the list of the Coleoptera of North America.”

72

72 This species is listed in synonymy with Platydema nigratum (Motschulsky) in some catalogues. However, Triplehorn (1965: 424) indicated that it was “quite distinct.”

73

73Dajoz (1975) used two spellings for the specific name, jamaicensis (p. 118) and jamaiscensis (pp. 120, 121). Ivie and Hart (2017b: 118) acted as “First Revisers” and choose jamaicensis as the correct original spelling.

74

74 Until the type specimens of Menimopsis jamaicensis (Dajoz) are studied, we prefer not to propose a replacement name for Kaszab’s taxon.

75

75Soldati and Touroult (2014: 106) believed this species to be a Cerylonidae and not a Tenebrionidae.

76

76 May not be established as there is no record since the 1950’s (Peck 2011: 29).

77

77 This species has passed under the name of Alobates morio (Fabricius) in many publications following the study of putative types by Blair (1914: 487) in the Banks collection as pointed out by Spilman (1962b: 60). However, Ferrer (2006: 232) mentioned that the “types” seen by Blair were not syntypes and that the true syntype he studied at the Museum of Copenhagen belongs in fact to Zophobas atratus (Fabricius, 1775).

78

78 The generic name Scotera and the specific name gibbosa were proposed as synonyms of Cibdelis blaschkii by Motschulsky (1845b: 365, 366). However, both taxa are available from Motschulsky’s work (ICZN 1999: Article 11.6.1) because they were treated as valid names before 1961 (e.g., Chevrolat 1848: 454).

79

79Doyen (1973) retained C. californica as the valid name for this taxon. However, Horn (1870: 336) acted as First Reviser (ICZN 1999, Article 24.2) in synonymizing C. californica and C. dilaticollis together and chose C. dilaticollis as the valid name for the taxon. Therefore C. dilaticollis is the valid name for this species.

80

80 Credited to Chevrolat (1844) in previous catalogues.

81

81Moeon is the Latinization of the Greek noun moion (privy parts) and is neuter (M.A. Alonso-Zarazaga, personal communication).

82

82Pallas (1781) proposed four species in Mylaris but did not described the genus. In a footnote on page 38, Pallas mentioned “Nomen Mylaridis, –molendinariae appropriatum, male applicavit Meloïdibus Fabricius [The name of Mylaris, –appropriate for a miller species, was incorrectly applied to Meloids by Fabricius].” It seems that Pallas used an incorrect subsequent spelling of Mylabris Fabricius, which was proposed by Fabricius (1775) for some meloid species, and decided to apply the name to tenebrionids. As such Mylaris, used by Pallas (1781), should be an incorrect subsequent spelling and not an available name. No action is taken at this time.

83

83 The name “cuproso” was used three times in the publication (pp. 98, 99, 203) and is not a lapsus. To our knowledge the species was treated only once subsequently (Hopp and Ivie 2009: 54) who used the spelling “cuprosum.” We do not consider that cuprosum is in prevailing usage and therefore, cuproso must be retained as the proper spelling.

84

84 Listed in the genus Otocerus by Ferrer and Ødegaard (2005: 648).

85

85 The name preciosus is a blend of Spanish and Latin, and as such must be treated as a noun in apposition (M.A. Alonso-Zarazaga, personal communication).

86

86 This Pleistocene fossil species was described from marine shale.

Appendix 1

List of impression fossil Tenebrionidae taxa described from North America. 1 = species originally described from Germany but subsequently reported from Greenland by Heer (1883: 145). 2 = assignment of this species to the tribe Blaptini (e.g., Kirejtshuk et al. 2008) is doubtful since this tribe is not represented by any extant taxa in North America.

Species Origin Age Placement
Blapstinus linellii Wickham, 1913a: 298 USA (Colorado) 37.2 to 33.9 Ma Tenebrioninae: Opatrini: Opatrina
Capnochroa senilis Wickham, 1913b: 365 USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Gonoderina
Cistelites minor Heer, 1874: 25 Greenland 61.7 - 58.7 Ma Alleculinae
Cistelites punctulatus Heer, 1870: 484 Greenland 61.7 - 58.7 Ma Alleculinae
Ephalus adumbratus Scudder, 1893: pl. 1 (fig. 3) USA (Colorado) 37.2 to 33.9 Ma Tenebrioninae: Opatrini: Opatrina
Gonodera antiqua (Wickham, 1913b: 365) USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Gonoderina
Gonodera vulcanica (Wickham, 1914a: 485) USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Gonoderina
Helops wetteravicus C. von Heyden & L. von Heyden, 1865: 331 Greenland 61.7 - 58.7 Ma Tenebrioninae: Helopini
Hymenorus haydeni Wickham, 1914a: 486 USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Alleculina
Isomira aurora Wickham, 1914b: 268 USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Gonoderina
Isomira florissantensis Wickham, 1914a: 486 USA (Colorado) 37.2 to 33.9 Ma Alleculinae: Alleculini: Gonoderina
Meracantha lacustris Wickham, 1909: 129 USA (Colorado) 37.2 to 33.9 Ma Tenebrioninae: Amarygmini
Miostenosis lacordairei Wickham, 1913a: 297 USA (Colorado) 37.2 to 33.9 Ma Pimeliinae: Stenosini
Pelecyphorus (Stenosides) primus (Wickham, 1910: 51) USA (Colorado) 37.2 to 33.9 Ma Pimeliinae: Asidini
Platydema antiquorum Wickham, 1912: 32 USA (Colorado) 37.2 to 33.9 Ma Diaperinae: Diaperini: Diaperina
Platydema bethunei Wickham, 1913a: 299 USA (Colorado) 37.2 to 33.9 Ma Diaperinae: Diaperini: Diaperina
Proteleates centralis Wickham, 1914b: 2672 USA (Colorado) 37.2 to 33.9 Ma Tenebrioninae: Blaptini
Protoplatycera laticornis Wickham, 1914a: 484 USA (Colorado) 37.2 to 33.9 Ma Tenebrionidae
Tenebrio primigenius Scudder, 1879: 183B Canada (British Columbia) 55.8 to 40.4 Ma Tenebrioninae: Tenebrionini
Teneobrionites alatus Cockerell, 1927: 586 USA (Colorado) 37.2 to 33.9 Ma Tenebrionidae
Ulus minutus Wickham, 1914b: 266 USA (Colorado) 37.2 to 33.9 Ma Tenebrioninae: Opatrini: Opatrina

Appendix 2

List of amber fossil Tenebrionidae taxa described from North America. 1 = new replacement name for Hymenorus oculatus Doyen and Poinar, 1994: 37 which is a junior primary homonym of Hymenorus oculatus Champion, 1888. 2 = Species originally described in the genus Hesiodobates Kaszab and Schawaller, 1984 which is currently considered a junior synonym of Nesocyrtosoma Marcuzzi, 1976 (Doyen and Poinar 1994).

Species Origin Age Placement
Corticeus tertiarius Vitali, 2007: 2 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Hypophlaeini
Cymatothes dominicus Doyen and Poinar, 1994: 36 Dominican Republic 20.4 – 13.7 Ma Tenebrioninae: Amarygmini
Hymenorus campbelli Bouchard, new replacement name1 Dominican Republic 20.4 – 13.7 Ma Alleculinae: Alleculini: Alleculina
Hymenorus chiapasensis Campbell, 1963: 41 Mexico (Chiapas) 23.0 – 16.0 Ma Alleculinae: Alleculini: Alleculina
Hypogena marginalis Doyen and Poinar, 1994: 35 Dominican Republic 20.4 – 13.7 Ma Tenebrioninae: Triboliini
Liodema phalacroides Doyen and Poinar, 1994: 43 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Diaperini: Diaperina
Lobopoda annosa Doyen and Poinar, 1994: 37 Dominican Republic 20.4 – 13.7 Ma Alleculinae: Alleculini: Alleculina
Lorelus angulatus Doyen and Poinar, 1994: 28 Dominican Republic 20.4 – 13.7 Ma Lagriinae: Lupropini
Lorelus foraminosus Doyen and Poinar, 1994: 28 Dominican Republic 20.4 – 13.7 Ma Lagriinae: Lupropini
Lorelus minutulis Doyen and Poinar, 1994: 30 Dominican Republic 20.4 – 13.7 Ma Lagriinae: Lupropini
Lorelus wolcotti Doyen and Poinar, 1994: 30 Dominican Republic 20.4 – 13.7 Ma Lagriinae: Lupropini
Neomida senicula Doyen and Poinar, 1994: 41 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Diaperini: Diaperina
Nesocyrtosoma antiquum (Kaszab and Schawaller, 1984: 3)2 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nesocyrtosoma celadonum Doyen and Poinar, 1994: 46 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nesocyrtosoma hadratum Doyen and Poinar, 1994: 47 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nesocyrtosoma impensum Doyen and Poinar, 1994: 47 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nesocyrtosoma minisculum Hopp and Ivie, 2009: 80 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nesocyrtosoma phthanatum Doyen and Poinar, 1994: 48 Dominican Republic 20.4 – 13.7 Ma Stenochiinae: Cnodalonini
Nilio dominicana Poinar and Brown, 2011: 233 Dominican Republic 20.4 – 13.7 Ma Nilioninae
Rhipidandrus quadripapillatus Doyen and Poinar, 1994: 33 Dominican Republic 20.4 – 13.7 Ma Tenebrioninae: Bolitophagini
Statira dermoidea Doyen and Poinar, 1994: 30 Dominican Republic 20.4 – 13.7 Ma Lagriinae: Lagriini: Statirina
Trientoma nascens Doyen and Poinar, 1994: 31 Dominican Republic 20.4 – 13.7 Ma Pimeliinae: Edrotini
Tyrtaeus azureus Doyen and Poinar, 1994: 39 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Gnathidiini: Anopidiina
Tyrtaeus cupreorutilans Vitali, 2008: 12 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Gnathidiini: Anopidiina
Tyrtaeus elongatus Doyen and Poinar, 1994: 40 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Gnathidiini: Anopidiina
Tyrtaeus flavoantennatus Doyen and Poinar, 1994: 40 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Gnathidiini: Anopidiina
Tyrtaeus thoracicus Doyen and Poinar, 1994: 41 Dominican Republic 20.4 – 13.7 Ma Diaperinae: Gnathidiini: Anopidiina
Wattius reflexus Doyen and Poinar, 1994: 33 Dominican Republic 20.4 – 13.7 Ma Tenebrioninae: Toxicini: Dysantina

Appendix 3

List of species incorrectly recorded from North America.

Allophasia marseuli Bates, 1873c: 237. The species was described from a unique specimen with no locality data. It was doubtfully recorded from North America by Papp (1961d: 122). Triplehorn and Brendell (1985: 12) found a few conspecific specimens from South America suggesting that it occurs in South America.

Ammophorus denticollis Boheman, 1858: 89. The species was originally reported from Panama. However Van Dyke (1953: 95) believe the specimen(s) was mislabelled as some others of Boheman’s species listed from Panama and was in fact collected in the Galápagos Islands. Since the genus is found only in Chile, Peru, mainland Ecuador, and the Galápagos Islands (Peck 2006: 219), Van Dyke’s statement is likely correct.

Ammophorus obscurus G.R. Waterhouse, 1845: 32. Waterhouse described this species from the Galápagos Islands. Blaisdell (1943: 238) reported seeing one specimen of the species from San José del Cabo in Baja California Sur but also suggested that its provenance should be taken with some doubt. For the reason outlined for the previous species, the Mexican record is unreliable and certainly based on a mislabelled specimen.

Blapstinus latifrons LeConte, 1874: 70. This species was described from one specimen collected on Vancouver Island. The name is currently a junior synonym of Gonocephalum bilineatum Walker, a widespread species in southeast Asia (Kaszab 1952: 682). As pointed out by Aalbu and Triplehorn (1985: 273), this isolated record is probably the result of an accidental interception.

Cistelopsis instriata Pic, 1930: 23. This species, described from “Indes Méridionales,” was recorded from “West Indies” by Blackwelder (1945: 508). However we believe the record is based on a misinterpretation of the region given by Pic and the species is probably an inhabitant of India (see next entry).

Cistelopsis rufomarginata Pic, 1930: 28. This species, along with its variety ruficolor, were described from “Indes Méridionales” and recorded from “West Indies” by Blackwelder (1945: 508) apparently from a misinterpretation of the locality given by Pic (1930: 28). The species, which is currently included in the genus Stilbocistela Borchmann, is known from India (Novák 2013: 180).

Cyclosattus websteri Casey, 1891: 710. The original locality given for this species is “Colorado.” However, Doyen et al. (1990: 244) pointed out that the species is a synonym of the Australian Cilibe costatus Solier, 1848 which is currently placed in the genus Saragus Erichson, 1842.

Cyrtosoma excisicollis Gebien, 1928b: 198. This species was described from specimens originating in “Paramaribo” and “Amazonas” and doubtfully recorded from North America by Papp (1961d: 131). We have found no record indicating that the species occurs in North America.

Emmenastus rugosus Motschulsky, 1845a: 76. This taxon was originally described on specimens from “Sitka” in Alaska. Aalbu et al. (1995: 484) studied a syntype of the species from the Zoological Museum of the University of Moscow and concluded that it belongs to the genus Oxycara Solier, 1835. The genus is not found in North America and therefore the locality originally listed is incorrect.

Helops ovipennis Casey, 1890b: 487. The locality given for this species was “Majave Desert, California”. Spilman (1959: 63) matched the holotype in USNM to members of Adelium Kirby which are found in the Australian Region and South America. Matthews (1998: 777) added that the holotype belongs to a valid species from New South Wales closely related to Adelium heterodoxum Lea and usually confused with it in collections. The specimen is obviously mislabelled.

Hyocis championi Fauvel, 1904: 166. This species has been listed from “Baja Calif.” by Blackwelder (1945: 525). The species belongs to the genus Parahyocis Kaszab and is found in New Caledonia, Vanuatu and Samoa (Kaszab 1955: 645). Blackwelder’s record is incorrect.

Iphthinus aereus Melsheimer, 1846: 65. The locality listed for this taxon was “Pennsylvania.” The species actually belongs to the genus Encyalesthus Motschulsky, 1860 (= Derosphaerus J. Thomson, 1858) from the Oriental Region as first pointed by LeConte (1873: 335). The original locality is obviously incorrect. Melsheimer’s species is the type species of the genus Pachyurgus LeConte, 1862 which is a junior synonym of Derosphaerus.

Maracia haagi Gebien, 1919: 35. This species was described from one specimen without collection locality. It was reported from South America by Blackwelder (1945: 538) but from Central America by Papp (1961d: 131). As there is no way to be certain that the species occurs in North America, it is not included in this catalogue. The only other species placed in the genus, M. femoralis (Kirsch), is known from Colombia.

Opatrum agricola Herbst, 1783: 35. The only record of this species in North America is from Horn (1870: 389) who, under the new specific name Eledona fungicola, mentioned that it was collected in the “Middle States.” Since the species was not found subsequently, it is likely that Horn’s specimen(s) was mislabelled. Eledona fungicola Horn was placed in synonymy with E. agricola (Herbst) by LeConte and Horn (1883: 384).

Oplocephala chalybea Laporte and Brullé, 1831: 341. The authors mentioned “La patrie de cette espèce est Philadelphie.” Horn (1870: 380) stated that the species is “probably not North American,” which is also the opinion of Triplehorn (2006: 332).

Oplocephala collaris Laporte and Brullé, 1831: 347. Laporte and Brullé stated “Cette espèce fait partie de la collection de M. Dupont, qui l’a reçue de Philadelphie.” Triplehorn (2006: 332) pointed out that the statement do not indicate that the species was collected in Philadelphia. From the description, including the size, the name cannot be attributed to any species from northeastern North America. Likely the species was not collected in North America.

Phaleria ornata Wollaston, 1864: 494. This species was recorded by Papp (1961d: 121) from “Alas. to Can.” and the subspecies P. ornata bockeri Heyden and P. ornata nigrothoracica Heyden from “N. Am.” Actually the species Phaleria ornata, of which the two Heyden’s names are junior synonyms, is endemic to the Canary Islands (Löbl et al. 2008: 316). Papp (1961d: 121) records are obviously incorrect.

Phrenapates mandibularis Gebien, 1910b: 504. This species was described from “Nördl. Süd-Amerika” and doubtfully recorded from North America by Papp (1961d: 124). We have not found any record that would indicate that the species occurs in North America.

?Platolenes desfontainesi Pic, 1952: 2. This species was originally described from Haiti. It was doubtfully transferred to the genus Amarygmus Dalman,1823 by Bremer (2001: 57). Since the genus Amarygmus is not represented in the Western Hemisphere, it is possible that Pic’s specimen(s) were mislabelled.

Platydema infuscata Laporte and Brullé, 1831: 373. This species was originally reported from “Colombie.” It was doubtfully listed from North America by Papp (1961d: 123). We have found no record indicating that the species occurs in North America.

Tenebrio angulatus Perty, 1830: 57. This species, described from “Prov. Piauhiensi” in Brazil, was reported under the generic name Tauroceras Hope, 1841 from several places in North America (see Champion 1885: 106; Champion 1892: 523; Blackwelder 1945: 535; Papp 1961d: 129). According to Ferrer et al. (2005) who revised the genus Tauroceras, Perty’s T. angulatus occurs only in South America.

Tenebrio olivensis Wollaston, 1864: 501. This species is listed from Canada by Papp (1961d: 130). The name is actually a junior synonym of Zidalus niloticus (Mulsant and Rey, 1853). The species is found only in the Canaries, Libya, Yemen and the Afrotropical Region (Löbl et al. 2008: 291). Papp’s record from North America is obviously incorrect.

Tenebrio picipes Herbst, 1797: 245. This species, which belongs to the genus Neatus LeConte, is listed as occurring in the Nearctic Region by Löbl et al. (2008: 299) apparently based on old records from the North American literature. Actually the species is found only in the Palaearctic Region and the records from North America refers to Neatus tenebrioides (Palisot de Beauvois).

Trogosita bidens Fabricius, 1792: 116. This species was originally reported from “Gallia.” Gebien (1906: 233) transferred the species to the genus Alegoria Laporte and mentioned that the original locality is incorrect. Gebien (1940: 768) listed the species from “Am. mer.,” Blackwelder (1945: 531) from “South America” and Papp (1961d: 126) from “N. Am.” Until a North American locality is recorded, the species is not included for the North American fauna.

Appendix 4

Supporting references for conservation of Tarpela micans (Fabricius, 1798) over Tarpela vittata (Olivier, 1793) through reversal of precedence (ICZN 1999: Art. 23.9.2). To our knowledge Tarpela vittata (Olivier) has not been used as a valid name after 1899.

Anonymous (2013) Patuxent Research Refuge: comprehensive conservation plan. United States Fish and Wildlife Service, Laurel, Maryland.

Bousquet Y, Bouchard P, Davies AE, Sikes DS (2013) Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition. Pensoft, Sofia-Moscow, 402 pp.

Bousquet Y, Campbell JM (1991) Family Tenebrionidae - darkling beetles. In: Bousquet Y (Ed.) Checklist of beetles of Canada and Alaska. Agriculture Canada, Ottawa, 253–261.

Cifuentes-Ruiz P, Zaragoza-Caballero S, Ochoterena-Booth H, Morón MA (2014) A preliminary phylogenetic analysis of the New World Helopini (Coleoptera, Tenebrionidae, Tenebrioninae) indicates the need for profound rearrangements of the classification. ZooKeys 415: 191–216. https://doi.org/10.3897/zookeys.415.6882

Ciegler HC (2014) Tenebrionoidea of South Carolina (Coleoptera: Mycetophagidae, Archeocrypticidae, Tetratomidae, Melandryidae, Mordellidae, Ripiphoridae, Zopheridae, Tenebrionidae, Synchroidae, Oedemeridae, Stenotrachelidae, Meloidae, Mycteridae, Boridae, Pythidae, Pyrochroidae, Salpingidae, Anthicidae, Ischaliidae, and Aderidae). Clemson University Public Service Publishing, Clemson, 243 pp.

Dillon E, Dillon LS (1972) A manual of common beetles of eastern North America. Volume II. Dover Publications, Inc., New York, 435–894.

Downie NM, Arnett RH Jr (1996) The beetles of northeastern North America. Volume II: Polyphaga: Series Bostrichiformia through Curculionoidea. The Sandhill Crane Press, Gainesville (Florida), x +891–1721.

Dunford JC, Young DK (2004) An annotated checklist of Wisconsin darkling beetles (Coleoptera: Tenebrionidae) with comparisons to the western Great Lakes fauna. Transactions of the American Entomological Society 130: 57–76.

Evans AV (2014) Beetles of eastern North America. Princeton University Press, Princeton & Oxford, 560 pp. https://doi.org/10.1515/9781400851829

Gardner JE (1986) Invertebrate fauna from Missouri caves and springs. Missouri Department of Conservation, Natural History Series, no. 3, i–vi, 1–72.

Guarnieri FG (2010) A survey of the beetles (Coleoptera) of Pocomoke River State Park, Worcester County and Tuckahoe State Park, Caroline County, Maryland. The Maryland Entomologist 5(2): 5–28.

Headstrom R (1977) The beetles of America. Barnes & Company, Cranbury (NJ).

Laplante S, Bousquet Y, Bélanger P, Chantal C (1991) Liste des espèces de coléoptères du Québec. Fabreries Supplément 6, 136 pp.

Majka CG, Chandler DS, Donahue CP (2011). Cheklist of the beetles of Maine, USA. Empty Mirrors Press, Halifax, 328 pp.

Matthews EG, Lawrence JF, Bouchard P, Steiner WE, Ślipiński SA (2010) Tenebrionidae Latreille, 1802. In: Leschen RAB, Beutel RG, Lawrence JF (Eds) Handbook of Zoology. Coleoptera, Beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia partim). De Gruyter, Berlin, New York, 574–659.

Nabozhenko M, Löbl I (2008) tribe Helopini Latreille, 1802. In: Löbl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera. Volume 5. Tenebrionoidea. Apollo Books, Stenstrup, 241–257.

Poole RW, Gentili P (Eds) (1996) Nomina Insecta Nearctica. A Check List of the Insects of North America. Volume 1: Coleoptera, Strepsiptera. Entomological Information Services, Rockville, Maryland, 820 pp.

Shockley FW, Cline AR (2004) A contribution to the inventory of Coleoptera of Missouri: new records from Benton County. Journal of the Kansas Entomological Society 77: 280–284. https://doi.org/10.2317/0304.21.1

Sikes DS (2004) The beetle fauna of Rhode Island: an annotated checklist. The Biota of Rhode Island, volume 3. The Rhode Island Natural History Survey, Kingston, 286 pp.

Steiner WE Jr (1995) Structures, behavior and diversity of the pupae of Tenebrionidae (Coleoptera). In: Pakaluk J, Slipinski SA (Eds) Biology, phylogeny, and classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa, 503–539.

Steiner WE Jr (2008) A checklist of the darkling beetles (Insecta: Coleoptera: Tenebrionidae) of Maryland, with notes on the species recorded from Plummers Island through the 20th Century. Bulletin of the Biological Society of Washington 15: 133–140. https://doi.org/10.2988/0097-0298(2008)15[133:ACOTDB]2.0.CO;2

Steiner WE Jr (2009) The Helopini (Coleoptera: Tenebrionidae) of Virginia. In: Roble SM, Mitchell JC (Eds) A lifetime of contributions to myriapodology and the natural history of Virginia: A Festschrift in honor of Richard L. Hoffman’s 80th birthday. Virginia Museum of Natural History Special Publication No. 16, Martinsville, USA, 331–339.

Tschinkel WR, Doyen JT (1980) Comparative anatomy of the defensive glands, ovipositors and female genital tubes of tenebrionid beetles (Coleoptera). International Journal of Insect Morphology and Embryology 9: 321–368. https://doi.org/10.1016/0020-7322(80)90009-4

Triplehorn CA, Karns KD (2016) The darkling beetles of Ohio. Ohio Biological Survey Bulletin (new series) 18(1), 58 pp.

Wiggins GJ, Grant JF, Lambdin PL (2007) Diversity of darkling beetles (Coleoptera: Tenebrionidae) from Arnold Air Force Base in the barrens of the Eastern Highland Rim, Tennessee. Natural Areas Journal 27: 66–71. https://doi.org/10.3375/0885-8608(2007)27[66:DODBCT]2.0.CO;2

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