Académique Documents
Professionnel Documents
Culture Documents
A Guide to 1,000
Foraminifera
from Southwestern Pacific
New Caledonia
Jean-Pierre Debenay
IRD Éditions
Institut de recherche pour le développement
Marseille
2012
Photos de couverture / Cover photographs
p. 1 – © J.-P. Debenay :
les foraminifères : une biodiversité aux formes spectaculaires / Foraminifera: a high biodiversity with a spectacular variety of forms
p. 4 – © IRD/P. Laboute :
îlôt Gi en Nouvelle-Calédonie / Island Gi in New Caledonia
Sauf mention particulière, les photos de cet ouvrage sont de l'auteur / Except particular mention, the photos of this book are of the author
La loi du 1er juillet 1992 (code de la propriété intellectuelle, première partie) n'autorisant, aux termes des alinéas 2 et 3 de l'article L. 122-5, d'une part, que
les « copies ou reproductions strictement réservées à l'usage privé du copiste et non destinées à une utilisation collective » et, d'autre part, que les analyses et
les courtes citations dans un but d'exemple et d'illustration, « toute représentation ou reproduction intégrale ou partielle, faite sans le consentement de l'auteur
ou de ses ayants droit ou ayants cause, est illicite » (alinéa 1er de l'article L. 122-4).
Cette représentation ou reproduction, par quelque procédé que ce soit, constituerait donc une contrefaçon passible des peines prévues au titre III de la loi précitée.
© IRD/MNHN, 2012
ISBN IRD : 978-2-7099-1729-2
ISBN MNHN : 978-2-85653-698-8
Contents/Sommaire
Foreword/Avant-propos ............................................................................................ 7
Acknowledgements/Remerciements ............................................................................... 9
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11
Regional setting . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Study area . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
Introduction to foraminifera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19
Taxonomy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 47
Systematics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 251
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 327
Abstract/Résumé . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 379
Contents . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 383
To Guy Cabioch
without whom this book would not have been possible.
A Guy Cabioch
sans qui cet ouvrage n’aurait pas vu le jour.
Foreword/Avant-propos
The story began 35 years ago, in 1976, when I joined the geolog- L’histoire a débuté il y a 35 ans, en 1976, quand j’ai rejoint
ical team of the ORSTOM center of Nouméa (presently IRD l’équipe de géologie du centre Orstom de Nouméa (actuelle-
[Institut de recherche pour le développement]). I was teacher in ment IRD, Institut de recherche pour le développement).
a high school, but had the opportunity to participate in a Professeur dans un lycée, j’ai eu l’opportunité de participer à
research program on the sediments of the southwest lagoon of un programme de recherche sur les sédiments du lagon sud-
New Caledonia, in collaboration with F. Dugas. Using the newly ouest de Nouvelle-Calédonie, en collaboration avec F. Dugas.
arrived R/V Vauban, more than 800 samples were collected, over Profitant de l’arrivée du N/O Vauban, plus de 800 échantillons
an area about 3,000 km2, and analyzed – mostly grain size ont été prélevés, sur une zone d’environ 3 000 km2, et
analysis and microscopic observation. The result was the publi- analysés – principalement granulométrie et observation
cation of sedimentological maps together with maps showing microscopique. Il en résulta la publication de cartes sédimen-
the contribution of foraminifera and mollusks to the sediment. tologiques accompagnées de cartes montrant la contribution
Four sets of two maps at 1/50,000 were published concerning des foraminifères et mollusques à la constitution du sédiment.
the areas of Tontouta, Nouméa, Mont Dore and Prony, from Quatre jeux de deux cartes au 1/50 000 ont été publiés pour les
north to south, complemented by substantial explanatory notes. zones de Tontouta, Nouméa, Mont Dore et Prony, du nord au
A synthetic map was also published in the Atlas of New sud, complétés par des notices détaillées. Une carte synthétique
Caledonia (1981). These maps may be downloaded on the site: a également été publiée dans l’Atlas de Nouvelle-Calédonie
<http://www.cartographie.ird.fr/sphaera/>. (1981). Ces cartes peuvent être téléchargées sur le site :
This work coincided with a growth in scientific research programs <http://www.cartographie.ird.fr/sphaera/>.
focalized on the marine environment, which led to the publication Ce travail a coïncidé avec le développement de programmes
of numerous thematic charts, such as those included in the Atlas dédiés à l’environnement marin, ce qui a conduit à la publi-
of New Caledonia (1981). cation de nombreuses cartes thématiques, telles que celles de
Later, an extensive study of the foraminifera from the fraction l’Atlas de Nouvelle-Calédonie (1981).
0.5-2 mm of all the samples led to the writing and defense of a Plus tard, une étude détaillée des foraminifères de la fraction
PhD thesis (DEBENAY, 1986) and to the publication of several 0,5-2 mm des échantillons a abouti à la rédaction et la
related papers, between 1985 and 1988, while I had a position at soutenance d’une thèse (DEBENAY, 1986), et à la publication
the university of Dakar... Unfortunately, foraminifera from the de plusieurs articles, entre 1985 et 1988, alors que j’étais en
fraction 0.125-0.5 mm are still in their boxes and have never poste à l’université de Dakar… Malheureusement, les fora-
been studied. minifères de la fraction 0,125-0,5 mm sont toujours dans
I came back in New Caledonia in 1997 for a sampling campaign leur boîte et n’ont jamais été étudiés.
in mangrove swamps, with a view to a more general work on Je suis revenu en Nouvelle-Calédonie en 1997 pour une série
foraminifera from paralic environments (DEBENAY & GUILLOU, d’échantillonnages dans les mangroves, en vue d’un travail plus
2002). général sur les foraminifères des environnements paraliques
In 2006, together with G. Cabioch, we listed the works related to (DEBENAY et GUILLOU, 2002).
foraminifera from New Caledonia, and made the inventory of the En 2006, avec G. Cabioch, nous avons répertorié les travaux
585 species reported from the area in previous works (DEBENAY & portant sur les foraminifères de Nouvelle-Calédonie et
CABIOCH, 2007). At the same time, I came back on a position at the inventorié les 585 espèces citées par les auteurs précédents
IRD center of Nouméa where I carried out several works on (DEBENAY et CABIOCH, 2007). Je suis alors revenu sur un poste
foraminifera. au centre IRD de Nouméa où j’ai réalisé plusieurs études sur
Foraminifera are one of the most abundant groups in the lagoon les foraminifères.
of New Caledonia, as shown in the Compendium of marine Les foraminifères constituent l’un des groupes les plus
species from New Caledonia (PAYRI & DE FORGES, eds, 1987): abondants dans le lagon de Nouvelle-Calédonie, comme
foraminifera (6%), algae (5%), molluscs (23%), arthropods (22%) cela a été montré dans le Compendium des espèces marines
and vertebrates dominated by fish (19%). Moreover, their tests de Nouvelle-Calédonie (PAYRI et DE FORGES, eds, 1987) :
often constitute the predominant part of the lagoonal sediments. foraminifères (6 %), algues (5 %), mollusques (23 %),
8 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Adding the fact that the Lagoons of New Caledonia have been arthropodes (22 %) et vertébrés, dominés par les poissons
inscribed on UNESCO World Heritage List that will need an (19 %). De plus, leurs tests constituent souvent une part
increase of the ongoing research, and will increase public interest prédominante des sédiments. Si l’on ajoute l’inscription des
and appreciation for marine biodiversity, it appeared necessary to lagons de Nouvelle-Calédonie sur la Liste du patrimoine
provide a synthetic work on the present knowledge about New mondial de l’Unesco, qui va nécessiter un développement de
Caledonian foraminifera. la recherche et va accroître l’intérêt du public et sa compré-
hension de la biodiversité marine, il apparaissait nécessaire
de fournir un ouvrage synthétique sur la connaissance
actuelle des foraminifères néo-calédoniens.
Acknowledgements/Remerciements
The bulk of this work was scientifically and financially supported L’essentiel de ce travail a été soutenu scientifiquement et
by ORSTOM (Office de recherche scientifique et technique financièrement par l’Orstom (Office de recherche scientifique
outre-mer), later becoming IRD (Institut de recherche pour le et technique outre-mer), devenu IRD (Institut de recherche
développement). Many people have provided help and encoura- pour le développement). Nombreux sont ceux qui m’ont aidé
gement over the years, without which this book would not have et encouragé pendant ces années. Sans eux, ce livre n’aurait
been possible. My first thanks are for F. Dugas who, in 1976, gave pas vu le jour. Mes premiers remerciements sont pour
me the opportunity to participate in the bottom sediment F. Dugas qui, en 1976, m’a permis de participer à la carto-
mapping of the lagoon, and to J. Récy for accepting me in his graphie sédimentaire du lagon, et à J. Récy qui m’a accepté
research group during the three years it took me to complete this dans son équipe de recherche pendant les trois ans qu’a duré
project. I am grateful to the captain P. Furic and the crew of the ce projet. Je remercie P. Furic, capitaine du N/O Vauban, et
R/V Vauban for their assistance during the sampling cruises in son équipage pour leur aide pendant les campagnes de
the lagoon. Later, I benefited from the help and encouragement prélèvement dans le lagon. Ensuite, j’ai bénéficié de l’aide et
of L. Blanc-Vernet and B. Thomassin who co-supervised my thesis des encouragements de L. Blanc-Vernet et B. Thomassin qui
work, and J.-P. Margerel made available to me his unpublished ont codirigé ma thèse, et J.-P. Margerel m’a confié un exem-
work on the foraminifera of the bay of Saint Vincent. plaire de son travail inédit sur les foraminifères de la Baie de
For the second phase of my research in New Caledonia, since Saint-Vincent.
2006, my grateful acknowledgements are due to L. Ortlieb and Concernant la seconde phase de mes recherches en Nouvelle-
G. Cabioch for offering me a position in their research unit, and in Calédonie, depuis 2006, je remercie sincèrement L. Ortlieb et
the research team of Nouméa … despite my impending retirement. G. Cabioch pour m’avoir recruté dans leur unité de recherche
During this period, I had fruitful collaboration with several et dans l’équipe de Nouméa… malgré la proximité de mon
colleagues. Among them, G. Cabioch gave me core samples for départ à la retraite. Pendant cette période, j’ai pu avoir de
studying foraminiferal assemblages at a geological time scale; fructueuses collaborations avec plusieurs collègues :
D. Wirrmann took me on to collect, process and study sediment G. Cabioch m’a confié des échantillons de forages pour des
cores; C. Payri allowed me to get rich foraminiferal assemblages études à l’échelle géologique ; D. Wirrmann m’a embauché
from algal substrates; L. Della-Patrona provided me with samples pour collecter, préparer et étudier des carottes sédimentaires ;
from shrimp farms and, together with C. Marchand, samples C. Payri m’a permis d’observer les nombreux foraminifères
from mangrove swamps; J.-M. Fernandez gave me surface vivant sur des algues ; L. Della-Patrona m’a procuré des
samples and core samples from potentially contaminated areas; échantillons de fermes crevetticoles et, avec C. Marchand, des
B. Richer de Forges allowed me to get deeper samples from the échantillons de mangrove ; J-M. Fernandez m’a donné des
northern lagoon; and J.-L. Justine gave me the rare opportunity échantillons de surface et de carottes issus de zones potentielle-
to open a number of fish guts…, for him to collect parasites, and ment polluées ; B. Richer de Forges m’a permis d’accéder à des
me foraminifera. All of them are warmly thanked for that, and échantillons plus profonds du lagon nord ; J.-L. Justine m’a
for lively discussion on various research topics. donné la rare opportunité d’ouvrir un nombre respectable
During all this time, I also benefited from a large amount of de poissons… pour collecter, lui les parasites et moi les
technical help. Scanning electron micrographs were taken by, or foraminifères. Tous sont chaleureusement remerciés pour
with the help of M. Ndao, using the facilities of the university of cela et pour les discussions stimulantes sur divers sujets de
Dakar, M. Lesourd, using the facilities of the university of Angers, recherche.
O. Boudouma, using the facilities of university Pierre and Pendant toute cette période, j’ai aussi bénéficié d’une consi-
Marie Curie (UPMC Paris), and mostly S. Caquineau, using the dérable aide technique. Les photos au microscope électronique
facilities of LOCEAN, IRD, Centre Île-de-France, Bondy. ont été prises par, ou avec l’aide de M. Ndao, à l’université de
Underwater sample collection of algae was made by C. Payri, Dakar, M. Lesourd, à l’université d’Angers, O. Boudouma, à
J.-L. Menou and J. Butscher, the deep-water specimens being l’université Pierre et Marie Curie (UPMC Paris), et surtout
collected by J.-L. Menou and S. Beata during special high tech S. Caquineau, au centre Île-de-France IRD de Bondy (unité
TRIMIX divings, down to 125 m. Underwater fish collections are LOCEAN). Les algues ont été collectées en plongée par
10 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
due to M. Clarque, G. Mou-Tham and J. Butscher, while S. Tereua C. Payri, J.-L. Menou et J. Butscher, les échantillons profonds
and N. Colombani both captains of the R/V Coris provided étant collectés par J.-L. Menou et S. Beata au cours de plongées
technical assistance during the cruises. Back to the laboratory, spéciales TRIMIX, jusqu’à 125 m de profondeur. La capture
A. Di Matteo, A. Sigura and C. Dupoux provided assistance for sous-marine des poissons est due à M. Clarque, G. Mou-Tham
opening and processing fish guts. I thank T. Potiaroa and et J. Butscher, alors que S. Tereua et N. Colombani tous deux
H. Goguenheim for their help in preparing photographic plates, capitaines du N/O Coris assuraient l’assistance technique à
and J.-M. Boré and M. Vilayleck for making a movie about bord. Au laboratoire, les intestins de poissons étaient ouverts
foraminifera. et préparés avec l’aide d’A. Di Matteo, A. Sigura et C. Dupoux.
I owe a great debt of gratitude to a number of people from Je remercie T. Potiaroa et H. Goguenheim pour leur aide
administrative, technical and research staffs, and to my students dans la préparation des planches photographiques, et
who helped me over years during my work. Unfortunately, it is J.-M. Boré et M. Vilayleck pour la réalisation d’un film sur
impossible to mention them all here. Finally, I thank my wife, les foraminifères.
family and friends for having been patient with me, especially Je suis également redevable à de nombreux personnels
during the last months of writing this book. administratifs, techniques et de recherche, et à mes étudiants
The outcome of this book was possible thanks to the editorial qui m’ont aidé pendant ces années de recherche.
committee of IRD that accepted this publication, to T. Mourier Malheureusement, il est impossible de tous les nommer.
and his collaborators for their valuable editorial advices and Enfin, je remercie mon épouse, ma famille et mes amis pour
work, and to two anonymous referees for their thorough and leur patience à mon égard, particulièrement pendant les
careful reading and the detailed remarks that helped to improve derniers mois de rédaction.
significantly the initial manuscript. La réalisation finale de l’ouvrage a été possible grâce au
comité des éditions de l’IRD qui a accepté cette publication,
à T. Mourier et ses collaborateurs pour leurs précieux conseils
et travail éditoriaux, et à deux rapporteurs anonymes pour
leur lecture approfondie et leurs remarques détaillées qui ont
permis d’améliorer significativement le manuscrit initial.
Introduction
Foraminifera have an evolutionary history that extends back to The aim of this book is to give an overview of the present knowl-
the Cambrian, more than 525 million years ago. Since then, they edge on foraminifera from New Caledonia. In order to make this
have radiated and evolved. To date, approximately 60,000 fossil knowledge accessible to people who are not familiar with New
and modern species have been validly recognized (LANGER, 2011), Caledonia, and/or with foraminifera, two introductory parts
and an estimated 10,000 species (including only 40-50 planktonic describe the regional setting and the characteristics of New
species) are still living (VICKERMAN, 1992), constituting the most Caledonia, and a third one provides an introduction to
diverse group of shelled microorganisms in modern oceans (SEN foraminifera. The fourth part gives a synthesis of the main results
GUPTA, 1999). These small-sized organisms, usually 0.1 to 1 mm, published on Recent foraminifera from New Caledonia, and the
may be very abundant, and tens of thousands living specimens
last and most important part presents the 1043 species in the
per square meter may be found in some environments (WETMORE,
form of an illustrated atlas with photos and information about
1995). Their mineralized tests (shells) usually get preserved in
the morphology and taxonomy of most of the species.
the sediment after the death of the organism and may constitute
a major, sometimes the dominant, part of many modern or fossil This inventory will be helpful to professional micropaleontolo-
sediments (fig. 1). They are easy to collect, and their high-densi- gists, researchers, and students, but its main objective is to offer
ty populations provide an adequate statistical base, even in small environmental managers and all person interested in lagoonal
volume samples, to perform environmental analyses, making environment and protection the access to this invaluable tool for
them a powerful tool for environmental assessment. environment monitoring that are benthic foraminifera. It will
1 cm
| Figure 1
Sand from a beach of Grande Terre. Arrows show some foraminifera, but much more can be seen.
12 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
also contribute to feeding international database programs, description of 168 species. Most of them are deposited at the Museum
increasingly needed with the growing interest in biodiversity. d’Histoire Naturelle de Genève (DEBENAY & DECROUEZ, 1989). Several
papers were published (DEBENAY, 1985a, 1985b, 1986, 1988a, 1988b,
1988c). During the same period, the foraminifera of Quaternary
Previous works about reefal paleoenvironments were studied (CABIOCH et al., 1986;
foraminifera from New Caledonia CABIOCH , 1988), as well as the benthic (VINCENT, 1986; VINCENT
& LAURIN, 1988; VINCENT et al., 1991) and planktonic (LAMBERT
The first study of foraminifera from the southwestern Pacific near et al., 1991) foraminifera of the Loyalty basin. Two PhD theses
New Caledonia was carried out by BRADY (1884) during the voy- also provided inventories of foraminifera from New Caledonia
age of H.M.S. Challenger (1873-1876), updated by BARKER and Polynesia (ADJAS, 1988), and from New Caledonia including
(1960). The nearest station was station 177, near Vanuatu Chesterfield islands (DEGAUGUE-MICHALSKI, 1993). Samples have been
(16°45’S-168°5’E). However, studies concerning directly New collected in coastal marshes and mangrove swamps for a more
Caledonia began much later, with partial and local inventories in comprehensive study about the foraminifera of paralic environments
coastal samples (GAMBINI, 1958, 1959; RENAUD-DEBYSER, 1965; (DEBENAY & GUILLOU, 2002). Finally, an illustrated catalogue of the
TOULOUSE, 1965, 1966). Samples of recent and fossil sediments species from the Baie de Saint-Vincent has been prepared by Margerel
collected during the Singer-Polignac mission (1960-1965) were and is available on the web site of the University of Provence:
further used for several studies of foraminiferal assemblages http://mdp.cerege.fr/forams-index.php?position=0&der=&nbr=10.
(COUDRAY & MARGEREL, 1974; COUDRAY , 1976; MARGEREL, 1981). All the works reported above were used to prepare an inventory of
These samples allowed MARGEREL (1984) to make the first the foraminifera species that live in the waters from around New
detailed inventory of foraminifera from the Baie de Saint-Vincent Caledonia (DEBENAY & CABIOCH, 2007). At that time, 585 species
(southwest of New Caledonia). This inventory, unfortunately still were identified. Since 2009, several works have been published
unpublished, described 289 species. On the occasion of the sedi- about epiphytic foraminifera (DEBENAY & PAYRI, 2010), predation
mentological study carried out by the IRD in the southwestern by fish (DEBENAY et al., 2011), foraminifera as indicator of
lagoon of New Caledonia, mentioned above, more than 800 sur- environmental changes (DEBENAY & FERNANDEZ, 2009), colonization
face sediment samples were collected (fig. 2). They allowed the of new environments by foraminifera (DEBENAY et al., 2009a),
first exhaustive study of large foraminifera (> 0,5 mm), with the and foraminifera in shrimp ponds (DEBENAY et al., 2009b).
Bay of
Saint Vincent
Loyalty Basin
Nouméa
Prony
22° 30’ S
Isle of
Pines
Coral Sea
22° 30’ S
Geographic and geological Lansdowne Bank and Fairway Ridge; and seamounts along the
setting Loyalty, Norfolk and Lord Howe ridges. Matthew and Hunter
volcanic islands are located father to the southeast, on the
Located in the Southwest Pacific Ocean, about 1,500 km East of southern part of the Vanuatu volcanic arc.
Australia, the exclusive economic zone (EEZ) of New Caledonia The main island, Grande Terre, is the third largest island in the
covers around 1,400,000 km2, extending over 1,200 km N-S Pacific (after New Guinea and New Zealand). It is of continental
(between latitudes 15° and 26° S) and 1,800 km W-E (between origin and has a mountainous axis that reaches a maximum
longitudes 156° and 174° E) in the Southwest Pacific. It is altitude of 1,629 m. The Belep islands and Isle of Pines are also
schematically composed of a series of NW-SE trending ridges and mostly continental islands. The Loyalty islands are uplifted atolls
basins, formed during the geological history of the area, which built on a line of volcanic seamounts, nowhere rising much
began around mid-Cretaceous time. higher than 130 m. Maré in the south has some volcanic rocks
The New Caledonia archipelago comprises: the Grande Terre (the but is primarily composed, as the others islands, of uplifted lime-
largest island, 400 km-long and 50 km wide), extending to the stone. The Chesterfield islands and Bellona reefs are coral cays
Belep islands and the d’Entrecasteaux Reefs to the north and the along the perimeter of the plateaus, forming large atolls. The
Isle of Pines to the south (fig. 3), and supported by the New wide Landsdowne Bank is mostly sandy and 70-80 meters in
Caledonia Ridge, which is the northern extension of the Norfolk depth, but includes a small reef in the north, while the Fairway
Ridge; the Loyalty islands, on the Loyalty Ridge; Chesterfield and reefs, supported by the Fairway Ridge come close to the surface
Bellona plateaus, supported by the Lord Howe volcanic chain; and dry at low tide (fig. 3).
18° S
d’Entrecasteaux
Huon Atoll Reefs
Surprise
19° S New
Bampton Reefs Caledonia
Belep Islands
Avon
20° S Loyalty Islands
Landsdowne Ouvea
Bank
Mare
Observatory Cay
Isle of Pines
| Figure 3
Location of the main islands and reefs of the New Caledonia archipelago. In dark blue: the main lagoons.
14 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
A prominent dynamic regional feature is the subduction zone upwelling on daily Sea Surface Temperature (SST) variations.
between New Caledonia and Vanuatu, where the Australian plate This process, however, is strongly modulated by the seasonal
dips under the Vanuatu volcanic arc. The resulting lithospheric variations of the subsurface stratification.
deformation (bulge) of the Australian plate explains the uplifted
reefs of Grande Terre, Isle of Pines and Loyalty Islands (DUBOIS Climatic conditions
et al., 1974). in the southwest Pacific Ocean
The climate of the southwest Pacific region, primarily oceanic, is
controlled by large-scale atmospheric circulation features that
Oceanography and climate include the trade wind regimes, the Hadley and Walker circula-
tions, the seasonally varying tropical convergence zones, the
Hydrological conditions
semi-permanent subtropical high-pressure belt, and the zonal
in the southwest Pacific Ocean
westerly winds to the south (fig. 5).
Water circulation in the southwest Pacific follows complex path- In January, the prominent feature is the trough of low pressure
ways due to its strong interaction with the complex bathymetry of that extends eastward from the monsoonal low centered over
the region. The southern part of the South Equatorial Current northern Australia, while a high-pressure dome sits over southern
divides into jets: North/South Vanuatu Jet, and North/South Australia in July. The resulting monsoon regime is felt west of
Caledonian Jet. Those jets feed the western boundary current 170°W throughout the Vanuatu archipelago and the northern
system: the East Australian Current to the south and the New part of New Caledonia. The South Pacific Convergence Zone (SPCZ)
Guinea Coastal Current that itself feeds the Equatorial that extends from east of Papua New Guinea southeastward
Undercurrent to the North, through the Solomon Straits (fig. 4; toward 120°W, 30°S maintains one of the most expansive and
GANACHAUD et al., 2007). persistent cloud bands on earth. South of 30°S, the atmospheric
Surface waters of the EEZ are fed to the south by a branch of the circulation is characterized by the presence of an anticyclonic
East Australian Current that brings cold and salted waters and to belt (MAES et al., 2007). Under the combined effects of the seasonal
the north by warm and less salted waters coming from the South shifts in the SPCZ and the monsoon regime, the climate in New
Equatorial Current. The result is that the west coast receives Caledonia has typically a wet season that extends from January to
cooler waters (1-2°C) than the east coast (ROUGERIE, 1986). The April with a transition season from June to July and then a dry
characteristics of these waters are strongly influenced by the season from August to December.
seasonal variability of the water circulation (VEGA et al., 2005). The main signals at interannual timescales are linked to the
Strong cooling events off the western barrier reef of New variability of the ENSO phenomenon. The signature of El Niño
Caledonia have been attributed to wind-driven coastal upwelling. events in the oceanic region around New Caledonia is character-
ALORY et al. (2006) developed a simple one-dimensional model ized by a 20-50% decrease in precipitation (NICET & DELCROIX,
based on a heat budget in the mixed layer. This model suggests 2000), which may be related to the shifts in the position of the
that upwelling is the dominant process at daily timescale, and SPCZ in response to ENSO anomalies (FOLLAND et al., 2002;
that the surface heat fluxes have a smaller influence than FISCHER et al., 2004).
0° S
10° S 10° S
20° S 20° S
30° S 30° S
A B
40° S 40° S
140° E 150° E 160° E 170° E 180° 170° W 160° W 150° E 160° E 170° E 180° 170° W 160° W
| Figure 4
A) General circulation in the southwest Pacific (from GANACHAUD et al., 2007);
B) Average surface water circulation (from Kesler in VEGA et al., 2005).
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 15
July
INTERTRO
PICAL CONVERGENCE ZONE
Marshall Is.
Kiribati
0°
Phoenix Is.
NEW Solomon Is.
GUINEA SO
UT
H
PA DIVERGENT
CI EASTERLIES
FIC
SOUTH EAST TRADES C O Northern
N VCook Is.
Samoa ER
Vanuatu GE
NC Society Is.
Fiji EZ
20° S Southern ON
Cook Is. E
AUSTRALIA
Tonga
New Caledonia
Austral Is.
H H
TRAVELING ANTICYCLONES AND TROUGHS
40° S NEW H
ZEALAND
DISTURBED WESTERLIES L
L L
120° E 140° E 160° E 180° 160° W 140° W
January
Marshall Is.
I N T E RT ROP IC AL C O N V E R G E N C E Z O N E
Kiribati
0° NORTH WESTERLY MONSOON
Phoenix Is.
NEW Solomon Is.
GUINEA
Tuvalu
DIVERGENT
EASTERLIES
Northern
TROUGH S O U T H Samoa
OON PA C I Cook Is.
FIC C
ONS Vanuatu O NV E
M Fiji RGE
20° S L Southern
NC
EZ
AUSTRALIA ON
Tonga Cook Is. E
New Caledonia Austral Is.
H NEW H
40° S H ZEALAND
DISTURBED WESTERLIES
L L
| Figure 5
The southwest Pacific climatic conditions. Dashed lines represent the seasonal position of the convergence zones
(from Salinger et al., 1995; in MAES et al., 2007).
16 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
New Caledonia, located in the trade wind zone typically experiences drastic environmental changes obviously had significant effects
easterly to south easterly winds of 10 to 20 knots, but local topogra- on the New Caledonia marine biodiversity. Modern reefs could start
phy has strong effects on local wind patterns. The axial mountain growing when the substratum began to be flooded by the post-
range also affects rainfalls and, excluding the mountains, the glacial sea-level rise. Their morphology results from i) substrate
East Coast and the southeast parts of New Caledonia are the availability (preexisting reefs), ii) the postglacial rate of sea-level
wettest. variations from around - 120 m at 20/23 ka to the present sea-level
with a peak up to + 2 m at 5,5 ka due to isostatic readjustment,
Sea-level variations and iii) the growth strategy of living communities. During the
same time, species progressively colonize the lagoon. However, in
During the last million years, sea level had 100 kA cyclic high the region, sea surface temperatures may not have been suitable
(sometimes 5 to 10 m above present-day sea level) and low for corals before around 8 ka, which can explain the late (8,2 ka)
(120-130 m below present-day sea level) stands, leading to several formation of postglacial reefs in New Caledonia (CABIOCH, 2001).
emersions and submersions of the reefs and lagoons (CHEVILLOTTE Another alternative hypothesis is the lack of suitable substrate
et al., 2005; CHARDON et al., 2008; LE ROY et al., 2008). These (accommodation space) before 8 ka.
Study area
Lagoons and reefs approximately 2,000 km2, with a mean depth of 17.5 m. It
widens progressively towards the southeast from 8 km wide north
Reefal structures follow the eastern and western coast of Grande of Baie de Saint-Vincent, reaching 20 km near Nouméa and a
Terre, and extend beyond the island, 50 km southward and maximum of 65 km at its southern end. The barrier reef consists
200 km northward. Bounded offshore by a barrier reef built on of a series of arched reefs separated by deep passes (> 60 m). It
the border of the island shelf, the lagoons cover a total area of curves sharply to the north at its southern extremity, following
23,400 km2 (TESTAU & CONAND, 1983) (fig. 3). the edge of the great axial thalweg that prolongs the Bay of Prony
They have an average depth of 40 m along the east coast and of (fig. 3).
25-40 m along the west coast. They are connected to the sea by The lagoon can be subdivided into an external zone that
deep passes opened at the mouth of submarine valleys downcut comprises a series of shallow (10-20 m) indurated plateaus
by the rivers during the last glacial regression 20,000 years ago. downcut by the submarine valleys, and an internal zone, which
Extended lagoons are also found in the Chesterfield Archipelago, corresponds to the large and deep (40 m) lagoonal depressions.
d’Entrecasteaux Reefs and Loyalty Islands (mainly Ouvea). The back-reef area is characterized by large hydraulic sand dunes.
New Caledonian reefs include both oceanic (d’Entrecasteaux, The large bays, which indent the shoreline, are characteristic of
Loyalty and Chesterfield) and continental reefs (Grande Terre and a submerged coastline. The numerous intra-lagoonal reefs are
Isle of Pines). These reefs offer a large diversity of formations, arranged along three alignments roughly parallel to the coast
explained by the diversity of environmental forcing, and provide (THOMASSIN, 1984).
a rich framework that itself supports a large diversity of shallow
modern habitats and communities. There are 8 times more Hydrodynamics
lagoonal and sedimentary areas (~31,300 km2) than reef
Water movements in the southwestern lagoon of New Caledonia are
areas (~4,500 km2). New Caledonia is clearly a region of high
controlled mainly by tidal and wind forcings. The semi-diurnal
complexity, a hotspot of reef diversity, though it is not the most
tide (maximum tidal range = 1.8 m) propagates from the south
complex area (ANDRÉFOUËT et al., 2007, 2009).
to the north (DOUILLET, 1998), while southeasterly trade winds
The prominent feature of New Caledonia is its 1,500 km long drive a general northwest drift (DOUILLET et al., 2001). Models
barrier reef, cut by deep passes, and including a 1,300 km long suggest that oceanic waters enter the lagoon at its southern end,
subtidal domain. This is the longest stretch of barrier reef world- emptying through the passes. Field observations have shown that
wide, since the Great Barrier Reef in Australia is not a linear the wind-driven surface current to the northwest, which enters
barrier reef for most of its length, but an assemblage of platform the lagoon mostly through scattered reefs of the southeast, is
reefs of various sizes and shapes. The spatial organization of balanced by a subsurface return current to the southeast. During
Grande Terre reefs is not very diverse with an onshore-offshore each rising tide, oceanic water inflows mostly through the passes,
sequential zonation of fringing-patch-barrier reefs for most of its but also to a lesser extent over the barrier reef, and between the
perimeter, but more complex spatial organizations are found in scattered reefs of the southeast. Except when trade winds blow
the south lagoon due to higher abundance of patch reefs and wide suddenly stronger, a reversal of surface currents and undercurrents
shallow lagoons. The north sector is characterized by a very wide can be observed in some passes and in the lagoon during the shift
lagoon (Grand Lagon Nord) bounded by a continuous barrier of tidal flow: during flood tides the flow is E or NE in the passes,
reef, but depleted from patch reefs (ANDRÉFOUËT et al., 2007). NW or W in the lagoon; during ebb tides, the flow is SW or W in
passes and SE in the lagoon (ROUGERIE, 1986).
At a long-term scale, models indicate that tidal water mainly
The southwest lagoon enters the southwest lagoon at the south, between Ouen Island
and the barrier reef. One part flows directly to the ocean through
General features Boulari pass while the other part flows northwards and leaves the
The southwestern lagoon of New Caledonia has been extensively lagoon through Dumbéa pass. The velocity of the long-term
studied and therefore deserves a particular presentation. It covers transport generated by the tidal circulation is around 1 cm s-1,
18 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
whereas wind-induced velocity is 10 cm s-1 or higher (OUILLON In the inner bays, the sediment is red or brown in color, with
et al., 2010). Over the reef, the tidal flow may alternatively enter more than 8% of Fe2O3. In the outer bays, the depressions and
and leave the lagoon during trade wind episodes, but oceanic the submarine valleys, it is ochre as long as the percentage of
water may flow continuously toward the lagoon under west wind Fe2O3 is over 2%. In the external parts of the depressions and of
or weak wind. Freshwater inputs are mainly from the Dumbéa River, the submarine valleys bioclasts are colored in grey by iron pyrites.
Boulari River, and Pirogues River. Inside the southwest lagoon, the At the transition between the depression and the external plateaus
inter-annual variability is less marked than the seasonal one, which and sand dunes, sediments are yellow, due to the oxidation of
is well marked for most parameters (LEBORGNE et al., 2010). bioclasts previously colored by iron pyrites after reworking of
sediments (DEBENAY, 1987). The back reef hydraulic sand dunes
Sediments are white. Continental inputs are stocked close to the river
mouths and in the bays, elsewhere remaining noticeable only in
Sedimentary deposits are mainly of bioclastic origin. The finest submarines valleys.
sediments occur between the river mouths and the passes, in the
In the coarsest fractions (> 0.5 mm), Mollusks (Gastropods,
depression and the submarine valleys, where the proportion of silt
Pelecypods) and Foraminifers constitute the bulk of bioclastic
and clays is the higher (fig. 6). Coarser sediments are found near
material with locally coralline algae, Halimeda and/or coral
the patch reefs, due to the direct input of coarser grains from the
detritus (Debenay, 1985a). The contribution of coral debris to the
reefs, and in shallower areas, including back reef areas, due to
sediment is significant only close to the reefs. Generally, sedimen-
the winnowing of the sediment by waves and currents. According
tation of grains coarser than 63 µm within the lagoon is the
to the color of sediments, the lagoon appears to be divided into
result of in situ organic production combined with low hydrody-
four main areas roughly parallel to the coast and the barrier reef
namic control that lead to only weak sediment transport, as
(fig. 6). The color was shown to be directly related to continental
reported from other lagoons of New Caledonia (CHEVILLON, 1996).
iron-rich inputs, the zonation showing a decrease of these inputs
seaward (DEBENAY, 1987). Tests of foraminifera are often among the major constituents of
the sediment. Even in the coarser fraction (> 0,5 mm), they are
abundant, frequently making up more than 10% of the sediment
(fig. 7), this proportion sometimes reaching 80% or more.
Nouméa
Nouméa
| Figure 7
Contribution of foraminifera to the coarser (> 0.5 mm) fraction
of the sediment (after DEBENAY, 1985a).
| Figure 6
Color of the sediment and silt and clays content
(from DEBENAY, 1987).
Introduction to Foraminifera
10 µm
a b c e
| Figure 8
Pseudopodia. a) phase-contrast microscope image of Massilina secans showing pseudopodia extruding from the single aperture of the test;
b) phase-contrast microscope image of Ammonia beccarii showing bunches of pseudopodia extruding from sutural spaces;
c) same as (b), but under a Scanning Electron Microscope;
d) detailed view of (c) showing pseudopodia anchoring the test on the substrate;
e) phase-contrast microscope image showing the pseudopodial network of Heterotheca lobata
(photos c and d from V. Le Cadre; photo e from K. G. Grell in DEBENAY et al., 1996).
20 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
organism. The protoplasm of the cell is composed of endoplasm thus to colonize most of the photic water column.
and ectoplasm. The endoplasm is the central part of the proto- Parasitism by foraminifera has been reported for several species
plasm that contains the nucleus or nuclei and in which the since it was first documented by LE CALVEZ in 1947. In New
major metabolic processes take place. The ectoplasm is the outer Caledonia, a small species, Metarotaliella tuvaluensis, may
zone of cytoplasm, from which a reticular network of pseudopodia, potentially parasite several miliolid species. It attaches to the tests
reinforced by a micro-tubular cytoskeleton, may emerge through of partially grown miliolid individuals, resulting in malformation
a single or many openings in the test – but never through the of the chambers added after the attachment and modification of
pores (fig. 8). This pseudopodial network is used for locomotion, the test morphology (fig. 9).
anchoring, catching and transport of food, removal of excretory
products, gas exchange, test building, and many other functions.
Pseudopodia characteristically have small granules streaming in
both directions, and as they form a network, they are called
granuloreticulopodia.
0.1 mm
Development
Haploid individuals, named gamonts are usually uninucleate. of float chamber
They divide to produce numerous amoeboid or flagellated
gametes that fuse to produce zygotes. Zygotes develop into diploid,
generally plurinucleate, individuals, and named agamonts. The
Encystment
agamonts tend to have a small proloculus and are therefore
termed microspheric (fig. 10). After meiosis they fragment to
asexually produce new gamonts, which commonly form a larger
proloculus and are therefore termed megalospheric or macros-
pheric. Generally, they also are smaller in size. Multiple rounds | Figure 12 Gamont
of asexual reproduction between sexual generations are not Life cycle of a
Neoconorbina-like
uncommon in benthic forms. In this case, the agamont undergoes species
a mitotic division instead of meiosis and produces another diploid that constructs
generation, called schizont (fig. 11; see also Marginopora a float chamber Agamont
and has a
vertebralis on fig. 42). The schizont may undergo meiosis and planktonic stage
form gamonts or it may enter a cycle of successive asexual (drawing
from Myers,
reproductions by multiple fission of a diploid multinucleated in LOEBLICH
cytoplasm. These processes are complex and their explanation is & TAPPAN, 1964).
JUVENILES
Meiosis
SCHIZONT JUVENILES
Fission macrospheric
multinucleate
Growth diploid
Meiosis
Growth
Growth AGAMONT
macrospheric GAMONT
multinucleate macrospheric
JUVENILES diploid uninucleate
haploid
Fission
Gametogenesis
Growth mitosis
Fusion
ZYGOTE
GAMETES
| Figure 11
Schematic representation of the life cycle of foraminifera with the usual alternation of generation (solid lines),
and the alternative multifission (broken lines) (simplified from LEE et al., 2000).
22 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
A B C D
100 µm 100 µm 1 µm 1 µm
1 µm
100 µm 100 µm 1 µm
| Figure 15
Nature and structure of the test. 1) agglutinated; 2) calcareous hyaline; 3) calcareous porcelaneous
- A) light microscopy; B) SEM view of a test; C) detailed view of the surface showing the glue
between foreign grains on an agglutinated test, the pores on a hyaline test, and the rhombohedral
platelets on a porcelaneous test; D) sections showing the foreign grains in an agglutinated test, the
lamellae in a hyaline test, and the irregularly arranged calcitic needles covered with rhombohedral
platelets in a porcelaneous test.
Chamber arrangement
Some species build tests with a single chamber (unilocular), but
most species build multilocular tests with multiple chambers that
are added as the cell grows. Chambers are connected with each
other by small openings called foramina (Foraminifera got their
name from these foramina). The final chamber communicates
with the exterior through one or several openings called apertures.
The living cell fills all the chambers except for one or two of the
most recently constructed. While the cell grows, the chamber
increases in size in unilocular species, and new chambers are
added in multilocular species, following a great variety of
arrangements. The most common types of chamber arrangements
are shown on figure 17.
A B C
Besides these general categories, there are many variations in the
test morphology. For example, planispiral tests may be involute
| Figure 16 (the chambers in a coil cover laterally those of the preceding coil,
Examples of agglutinated tests:
A) with irregular coarse grains; the chambers of the last coil only visible) or evolute (all coils
B) with fine grains;
C) with selected sponge spicules. visible) (fig. 18 A, B). Chambers may be irregularly added as
Scale bar = 100 µm. illustrated in a few examples in figure 18 (C, D, E). In milioline
24 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Monolocular
The living cell is enclosed in a chamber of variable morphology, 1 2 3
generally with an aperture, which is here at the end of a neck (1).
Monolocular tubular
The unique chamber is a tube, which can be straight, planispirally
enrolled (2) or irregularly (streptospirally) enrolled (3).
4 5 6
Serial
The chambers of the plurilocular test are arranged
in a unique linear series (uniserial - 4), two series (biserial - 5)
or three series (triserial -6). The aperture is on the last chamber.
Spiral 9
Chambers are spirally arranged, either in one plane,
the test being symetrical (planispiral - 7), or in three dimensions, 8
giving the test a trochoid morphology, like a snail shell
(low trochospiral -8 and high trochospiral - 9). 7
Milioline
Tubular chambers are spirally arranged, each chamber
being one-half coil in length.
The coil may be within a single plane (spiroloculine - 10),
or each successive chamber may be placed at an angle
from the previous one, leaving visible the three (triloculine - 11) 10 11
or the five (quinqueloculine) last chambers.
Discoidal
After a generally planispiral initial stage, annular chambers 12
divided into chamberlets are added within a single plane(12).
Complex morphology
The morphologies described above may combine into more
complex morphologies. For example, a triserial initial stage
may change into an uniserial arrangement during growth (13),
a spherical chamber may be added to a trochospiral test (14),
planispirally arranged chambers may be followed by uncoiled,
uniserially arranged chambers (15).
13 14 15
| Figure 17
Most common types of chamber arrangements (from DEBENAY & DELLA-PATRONA, 2009).
arrangements, spiroloculine enrolments may be evolute et al., 1980). The function of these structures in the biology of
(Spiroloculina – fig. 18 F1) or involute (Pyrgo – fig. 18 F2). In foraminifera is still poorly known.
quinqueloculine enrolments, successive chambers are added at an Various morphological adaptations of the test are known. As an
angle less than 180° (fig. 18 F3), and in triloculine arrangements example, it is possible to mention two test morphologies inter-
they are added at an angle more than 180° (fig. 18 F4). preted as adaptations for conducting light to the internal algal
The internal structure of the test may be very complex, resulting symbionts. In peneroplids, blunt ribs roughly parallel to the
from various patterns of addition of chambers, their subdivision periphery of the shell concentrate light by refraction, and symbionts
into chamberlets, and complex communications with each other. group in these areas of light concentration (fig. 19 A). In oper-
This is particularly obvious in large discoid foraminifera (e.g., culinids, transparent pillars conduct light to the symbionts, even
HOTTINGER, 1978), but even small tests may reveal a complex in involute tests where the first coils are covered by following ones
organization when they possess a canal system (e.g., BILLMANN (fig. 19 B).
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 25
| Figure 18
More information on chamber arrangement.
A) planispiral involute;
B) planispiral evolute;
C) numerous domelike chambers added in successive layers,
alternating in position, with prominent radial spines;
D) somewhat irregular chambers separated by stolonlike necks;
E) chambers added in irregular cycles;
F) milioline arrangements:
1- spiroloculine evolute, 2- spiroloculine involute, 3- sigmoid,
A 4- quinqueloculine and 5- triloculine,
chambers are numbered and successive planes are indicated (blue);
white arrow = growth of the last chamber, green arrow = direction
of coiling. Scale bar = 100 µm.
100 µm
Apertures
As shown above, foraminiferal tests are characterized by their
B morphology and chamber arrangement, but they are also char-
acterized by their aperture, which allows the cell to communicate
with the exterior. Apertures may have a great variety of positions
100 µm on the test. Some of the common apertural positions are shown
in figure 20.
| Figure 19 Apertures also have a great variety of morphology, the role of
Adaptation of the test for conducting light to endosymbionts:
A) concentration of light by refraction through ribs of the test; which in the biology of foraminifera has to be elucidated. Some
B) conduction of light through transparent pillars. of these morphologies are illustrated in figure 21.
26 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
| Figure 20
Some of the common positions of the aperture:
Terminal, at the end of the last formed chamber;
Interiomarginal, at suture between the distal wall
of the last formed chamber and the preceding coil –
basal, at the base of the distal wall in planispiral
and serial tests – extraumbilical, at the suture
of the last formed chamber on the umbilical side
of a trochospiral test, but not connected
with the umbilicus;
Interio-areal, near the base of the distal wall,
but not at the suture with the preceding coil;
Umbilical, located into the umbilicus;
Latero-marginal, at the periphery of the last formed
chamber, but slightly on one side of the test;
Areal, on the distal wall of the last formed chamber.
Scale bar = 100 µm.
| Figure 21
Some of the morphologies of the test aperture. Scale bar = 100 µm.
Unfortunately, it was not possible, within the scope of this Where and how to collect
guide, to provide more detailed information about test struc- foraminifera
ture and morphology. The reader will find this information in
the “Illustrated glossary of terms used in foraminiferal As indicated above (fig. 1), foraminifera tests are abundant in
research” (H OTTINGER , 2006), also available online: tropical sands. They can be easily observed and picked under a
<http://paleopolis.rediris.es/cg/CG2006_M02/>. dissecting microscope. For small specimens, and if they are too
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 27
A B 100 µm
| Figure 23
Lobatula lobatula attached on Halimeda:
A) the cyst has been partly destroyed;
B) the cyst has been entirely cleaned off.
Foraminifera,
their distribution and bevahior
This part summarizes results obtained during a first comprehensive fraction, one hundred and sixty eight species were found, and
study in the SW lagoon (1976-1979), and in a series of studies their distribution over the 800 samples was examined.
carried out since 2006 in various parts of New Caledonia. For
Since 2006, several hundreds of samples were collected in
ancient works, species names have been changed, when necessary,
mangrove swamps, shrimp farms, in the deeper parts of the
in agreement with actual species concepts.
northern lagoon and on the northern shelf (down to 700 m), on
macroalgae, and even within fish guts (fig. 24). The results of
these studies, published in several papers are summarized below,
Material and methods together with unpublished data.
During the first studies, more than 800 samples were collected Sediment samples collected since 2006 were washed through a
over an area of about 3,000 km2 in the southwestern lagoon and series of three sieves with mesh size 2 mm, 0.5 mm, and 0.063 mm;
on the southern shelf (fig. 2). Each was subjected to grain size macroalgae were examined under a dissecting microscope for
analysis, a general observation of sand grains under a dissecting observing attached foraminifera, then washed over the sieves for
microscope, counting of the tests according to their nature in the collecting free-living species; gut contents of reef fish were observed
fraction 0.125-0.5 mm, and a detailed specific analysis of under a dissecting microscope. All the three fractions (> 2 mm,
foraminiferal fauna in the fraction coarser than 0.5 mm. In this 0.5-2 mm, and 0.063-0.5 mm) were observed.
d’Entrecasteaux Concalis
18° S
Huon Atoll Reefs G. Cabioch
Self-collection
Surprise Algae
Shrimp farms
19° S New Mangrove swamps
Bampton Reefs Caledonia
Belep Islands
Avon
20° S Loyalty Islands
Landsdowne Ouvea
Bank
Mare
Observatory Cay
Isle of Pines
| Figure 24
Location of samples collected since 2006: Concalis, samples from the northern shelf provided by B. Richer de Forges;
G. Cabioch provided samples from Surprise Island (d’Entrecasteaux reefs); macroalgae were provided by C. Payri; shrimp-farm samples
were provided by L. Della Patrona; mangrove samples were collected during several field trips, and provided by C. Marchand.
30 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Bay of Bay of
Saint Vincent Saint Vincent
Nouméa Nouméa
Prony Prony
| Figure 25
Species richness and diversity (Shannon index) of foraminiferal assemblages in the fraction > 0.5 mm (from DEBENAY, 1986, 1988a).
| Figure 26
Distribution of agglutinated, hyaline and porcelaneous foraminifera in the fraction > 0.5 mm (from DEBENAY, 1985a).
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 31
| Figure 27
Location of the three transects.
Northwestern transect
271 448 449 450 451 452 453 454 100%
90
80
70
60
50
40
30
20
Agglutinated 10
0
Agglutinated species Depth (m) SHALLOW SMALL DEPRESSION PATCH REEF
except Textulariidae 0
Textulariidae 10
20
30
Agglutinated
Median transect
Nubeculariidae 851 848 847 846 800 768 772 792 791 790 789 779 780 100%
Miliolidae 90
80
Soritidae 70
60
Alveolinidae 50
40
30
20
10
0
Agglutinated Depth (m)
EXTERNAL PLATEAU LAGOONAL DEPRESSION BAY
0
Planorbulinacea
and Acervulinacea 10
Amphisteginidae 20
Calcarinidae 30
20
40
60
| Figure 28
Distribution of foraminifera coarser than 0.5 mm along the three transects (from DEBENAY, 1985a).
32 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
| Figure 29
Distribution of agglutinated, porcelaneous and hyaline foraminifera in the fraction 0.125-0.5 mm.
Distribution of selected species first set of maps concerns species found on the southern shelf, or
in the south-western part of the lagoon, subjected to marine
The better represented species have been grouped according to influence resulting from the prominent SE to NW drift of waters
their location in the lagoon, on the basis of factor analyses and from penetration of marine waters through the passes (fig. 30).
carried out on their relative abundance in several sets of samples Hyaline species are dominant, and species of Amphistegina are
(DEBENAY, 1985a, 1988a) (table 1). well represented.
These groupings, however, are of limited value due to the highly Other species are more widely distributed in the lagoon, but their
complex environment, with an enormous variety of ecological distribution areas are still connected to the southeast and/or to
niches and of environmental parameters that are acting in this the passes, which indicates, on the one hand, the influence of the
wide carbonated lagoon with reefs, deep depressions and various penetration of oceanic water on these species, and on the other
continental inputs. For example, the two dominant species in hand the areas of the lagoon subjected to oceanic influence
bays: Flintina bradyana is dominant in the bays opening into (fig. 31).
the lagoon while Operculina philippinensis (as O. bartschi) is Another set of species is mostly distributed in the back-reef
limited to the bay of Prony and its vicinity. More detailed analyses area, or around patch reefs (fig. 32). They are dominated by
will be necessary to relate species to environmental factors that, porcelaneous species.
unfortunately, were not available at the time of this study.
The two most abundant species in the lagoon are Marginopora
The distribution of forty-seven, frequent enough species has been vertebralis and Alveolinella quoyi, which are widely distributed,
mapped. Maps have been grouped according to the location of together with a few species (fig. 33), while other species, also
the species, even if this grouping is debatable due to the high distributed all over the lagoon, are present only in discontinuous
complexity in the distribution of some species (figs 30 to 36). The patches (fig. 34).
| Table 1
Groups of species determined by factor analyses (from DEBENAY, 1985a, 1988a).
Bold = dominant species; small letters = accessory species.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 33
| Figure 30
Species from the southern shelf directly influenced by the open sea,
and from areas of the lagoon mostly under marine influence coming through SE reefs and passes.
As shown by the factor analyses (table 1), Flintina bradyana is The information given by the maps is synthesized in table 2.
dominant in the bays that open into the lagoon. Its distribution This distribution, as it appears on the maps, results from the
area also extends in the submarine valleys that prolong the bays combined influence of all environmental parameters. Among
and in the adjacent depressions. Another species, Heterolepa them, the oceanic influence appears to have a prominent role.
praecincta, has a similar distribution, but to the south of the Two other parameters are known to influence foraminiferal
lagoon (fig. 35). Nummulites venusta is mostly found in the assemblages: depth and grain size of the sediment, which are
depressions of the lagoon, together with other, most widely somewhat interrelated due to decreasing energy with depth. Both
distributed species. data were available and the distribution of the most abundant
Heterolepa praecincta is hardly found farther northwest than and frequent species with depth and grain size was examined. For
Nouméa peninsula. Other species, such as Amphistegina radiata, graphing the distribution of species with these two parameters,
show the same tendency (fig. 30) while others (e.g., samples were grouped into classes with a class interval of 5 m for
Pseudomassilina australis and Quinqueloculina agglutinans) depth, and of 5% for grain size. The frequency of each species
have an inverse distribution, being hardly found farther southeast (% of samples where the species is represented) was calculated for
than Nouméa peninsula (fig. 36). each class of samples.
34 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
| Figure 31
Species widely distributed, but mostly connected to the SE open area and to passes.
| Figure 32
Species mostly found behind the barrier reef and/or around patch reefs.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 35
| Figure 33
Species widely distributed over the study area.
| Figure 34
Species irregularly distributed in discontinuous patches.
36 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
| Figure 35
Species mostly found in bays and depressions.
Bay of Bay of
Saint Vincent Saint Vincent
Présent Présent
Nouméa Nouméa
Prony Prony
| Figure 36
Species mostly found to the NW of Nouméa.
| Table 2
Species ordered following their distribution as it appears on the above maps.
Some species may appear in two environments.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 37
| Figure 37 | Figure 38
Distribution of species with depth Distribution of species with the proportion of silt and clay
(from DEBENAY, 1988a). (from DEBENAY, 1988a).
38 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Foraminifera indicators connection with the passes, and even behind the barrier reef for
of water circulation Gypsina globula, shows their dependence on oceanic inputs.
Comparison between their distribution areas, elongated and
The distribution of several species led to the distinction of four curved to the NW, and the water circulation described by ROUGERIE
foraminiferal biofacies and to the partition of the studied area (1986) suggests that they are distributed along the trajectory of
into four zones from the southeast to the northwest, which were subsuperficial oceanic water penetrating the lagoon through
interpreted as the result of decreasing oceanic influence, with a passes (fig. 39).
major role of Nouméa peninsula (DEBENAY, 1988b). The above
chapters show that the distribution of several species follows this The distribution patterns of foraminifera illustrate the double
trend, with Heterolepa praecincta and Amphistegina radiata water circulation prevailing in the lagoon (ROUGERIE, 1986).
that extend hardly farther northwest than Nouméa peninsula, Planktonic foraminifera, living in the open sea, are transported
while Pseudomassilina australis and Quinqueloculina agglu- into the lagoon by superficial water masses pushed toward the
tinans have an inverse distribution, being hardly found farther NW, through the scattered SE reefs, by the SE dominant trade
southeast than Nouméa peninsula (fig. 36). The hypothesis of winds. This constitutes the main penetration axis of oceanic
decreasing oceanic influence is further reinforced by the distribution waters into the lagoon. On the other hand, benthic species are
of planktonic tests in the 0.125-0.5 mm fraction (DEBENAY, 1988b). under the influence of subsurface waters, that penetrate mainly
Relatively abundant on the southern shelf, they rarely compose through the passes and circulate at the bottom of the lagoon.
more than 10% of the thanatocoenoses in the lagoon itself, but Their distribution gives a clear picture of the impact of these
are frequent in submarine depressions and valleys located waters on the benthos, with a flow curved towards the NW by the
between Ouen Island and Nouméa peninsula, and connected to general drift that affects the external part of the lagoon, following
the pass of Boulari. Northwest of Nouméa, their only distribution the external edge of lagoonal depressions.
area is connected to the pass of Dumbea. Since foraminiferal tests will be preserved in the sediment, the
This distribution is obviously related to low energy deep and/or image of the double water circulation given by foraminiferal
protected environments. It also indicates that surface oceanic thanatocoenoses will be fossilized. It shows how, in turn, fossil
waters, which transport the planktonic tests, mainly come foraminifera may provide a valuable tool for understanding
through the scattered reefs located south of Ouen Island and are hydrodynamics of fossil lagoonal environments.
pushed up to the Bay of Boulari. Most of the planktonic tests are
deposited before reaching Nouméa peninsula and its seaward Foraminifera indicators
extending shallows. The connection of distribution areas with the of sediment transport
passes of Boulari and Dumbea indicates that surface waters also
As it has been discussed above, it is highly probable that only weak
penetrate through these passes. The water circulation deduced here
postmortem displacement of tests occurs inside the lagoon, and that
from the distribution of foraminifera is in good agreement with
the distribution of thanatocoenoses reflects the position of living
the hydrodynamics described by Rougerie (in DUGAS & DEBENAY,
assemblages. In bays, however, strong river flows during storms
1981b).
or cyclones may lead to seaward sediment transport, including the
The distribution of several benthic species appears to be more transport of foraminiferal tests. In the Bay of Prony, a foraminifera,
closely related to oceanic influence, and among them Gypsina Operculina philippinensis, is abundant in the fraction > 0.5 mm
globula and Pyrgo denticulata are the most significant (figs 30 where it can compose up to 65% of the assemblages. The tests are
and 31). The distribution of their tests on the southern shelf, in generally well preserved and living specimens are frequent,
suggesting that this species lives in the bay (DEBENAY, 1988c).
Bay of
Outside the bay, eroded tests have been found, their relatively bad
Saint Vincent
preservation suggesting that they are allochthonous, presumably
transported out from the bay (fig. 40 left).
The distribution of the test, west and east of the entrance of the bay
can be explained by the strong alternating tidal currents through
Woodin channel and along the southern coast of Grande Terre.
Nouméa
The presence of tests on the deeper southern shelf, to the south,
Prony suggests a gravity-driven sediment transport on the southern shelf,
down to deeper low energy areas. The last area of test deposition,
along the south-east coast of Ouen Island, indicates a southward
Coral Sea longshore transport. The distribution of silt and clay is consistent
with the sediment transport direction deduced from the distribution
of O. bartshi, and the combination of both sets of information
allows a better comprehension of sedimentary dynamic in front
| Figure 39 of the bay (fig. 40 right). This approach should be of interest
Marine influence, as shown by foraminifera,
mainly Pyrgo denticulata and Gypsina globula owing to the Goro Nickel Mining Project that is developing in the
(from DEBENAY, 1988b). area and will require an extensive environmental assessment.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 39
| Figure 40
Sediment transport from the Bay of Prony (arrows),
deduced from the distribution of Operculina philippinensis (as O. batschi),
the color of the sediment, and the silt and clay content (from DEBENAY, 1988c).
| Figure 41
Schematic representation of foraminifera distribution in mangrove swamps.
40 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
from the same macroalgae collected in different settings in order When considering the entire dataset, the only strong relationship
to enhance the knowledge of epiphytal foraminiferal ecology (even if not a strict one) between macroalgae and foraminifera
(DEBENAY & PAYRI, 2010). was the presence of spinose rotaliids, soritids and Amphistegina
A total of 152 species of epiphytic foraminifera were identified on in filamentous thalli and three-dimensional mats. Apart from
81 substrates (75 samples of macroalgae belonging to 55 species this exception, the nature of the substratum is generally over-
and 6 samples of coral rubble) (fig. 42). shadowed by other factors such as light, depth and hydrodynam-
Only four of the 75 macroalgae were devoid of foraminifera. On ics in governing the distribution of foraminiferal assemblages.
the other specimens, average density was 10 individuals per cm2, For instance, macroalgae that have a wide bathymetric range
with higher densities, whatever the depth, on thalli consisting of harbor completely different communities at different depths.
a tridimentional network of branches or filaments (Gelidiopsis Among them, thalli of Homeostrichus sp. collected at 30 and 60
intricata, Caulerpa cupressoides var. lycopodium, Melanthalia m had only 3 of their 33 species in common, thalli of Halimeda
concinna, Rhodomelacae and Sphacelariacea), as well as on the discoidea from 3 and 38 m had only 2 of their 27 species in com-
flabellate Udotea geppiorum. Species richness, recorded by in mon, and thalli of Udotea geppiorum from 1 and 24 m had only
situ counts on fresh thalli preserved in seawater (average = 9), 3 of their 30 species in common. Typically observed were the
was the lowest on Sargassum spp. (< 5), and the highest on a dominance of large symbiont-bearing miliolids or rotaliids at
Distromium/Homeostrichus association (26). Fifty-five species shallow depths, the presence of smaller rotaliids at all depths, and
were recorded on the same sample after washing and sieving the the occurrence of cryptic species adapted to shaded environments
macroalgae. in deeper samples.
1 2
3 4
5 6
7 8
| Figure 42
Examples of epiphytic foraminifera on their substrates.
Scale bar = 0.5 mm except for figure 4 where scale bar = 5 mm.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 41
Comparisons of different types of macroalgae from the same depth mostly Cymbaloporetta to grow in the algal turf (fig. 43 left).
did not reveal any relationship between structural characteristics Before reproduction, individuals of this foraminifera genus reach
of the macroalgae and foraminiferal communities. Geographic their Tretomphalus phase by constructing a float chamber, and
origin appears to have a greater impact, as shown by the Q-mode then becoming planktonic (fig. 43 right).
hierarchical clustering of selected foraminifera living on shallow The fish feeds by combing the protected algal turf with its teeth,
(0-3 m) macroalgae (DEBENAY & PAYRI, 2010). Cluster 1 included catching the small organisms (including Foraminifera) that live
samples from coastal areas impacted by organic enrichment, and in the filamentous thalli. As it feeds during the day, the
cluster 2 samples were from areas of low anthropic impact off Tretomphalus are collected before becoming planktonic, at night.
Grande Terre. Clusters 3 and 4 group all the macroalgae collected As the individual biomass provided by Foraminifera is very small,
in the Chesterfield Archipelago, a high-energy oceanic atoll. a great number of individuals must be ingested to provide a
This study also reveals that some foraminiferal species might significant nutritional input. An average of 1,600 tests was found
have quite different life modes between environments and in the digestive tract of the three individuals of Pomacentrus
regions. For example, Sorites orbiculus was considered to use amboinensis studied, which represents about 0.025 g of biomass
only a limited number of macroids with bare, flat surfaces in (fig. 44 left). However, the diet of the fish also comprises other
Florida Bay and on the Great Barrier Reef (FUJITA & HALLOCK, organisms, such as worms (fig. 44 right), and seasonal studies
1999; LOBEGEIER, 2001), and to be permanently fixed to the blades will be necessary to determine how Pomacentrus amboinensis
of macroalgae and seagrasses in Japan (SARASWATI, 2002). In New adapts to the seasonal changes in its feeding resources, since the
Caledonia, it was found on a variety of substrata and appeared to production of Tretomphalus is seasonal.
have the same free-living behavior as Marginopora and Sediment feeders, which accidentally ingest great quantities of
Amphisorus. In accordance with the results of WILSON & RAMSOOK empty tests (up to 4,000 in a fish gut), have no impact on the
(2007), this study also shows that temporal availability of the foraminiferal population, but play a prominent role in the dis-
substratum (e.g., seasonal growth of algae) might play a role. persion of empty tests. They may introduce significant changes
in the thanatocoenoses, potentially introducing some bias in
paleoenvironmental interpretations. Incidental predators of living
Foraminifera in fish diet foraminifera are either herbivorous, which do not digest the
foraminifera or carnivorous, which ingest and digest insignificant
During 2007-2008, a systematic investigation of foraminifera in foraminiferal biomass. Foraminifera, still living after their transit
the gut contents of coral reef fish was carried out. It was the first through the digestive tract of herbivorous fish, are defecated with
investigation on a large number of individuals: 247 fish, belonging a significant effect on the dispersion of living individuals over
to 83 species (DEBENAY et al., 2011). The objectives were to: provide
information on the ingestion and digestion of foraminifera by fish;
determine the impact of predation on foraminiferal assemblages;
determine if some fish species could be considered as selective Cymbaloporetta squammosa
in the thallus of Chamaedoris sp.
consumer of Foraminifera; determine if the consumption of
Foraminifera can provide significant biomass to fish.
The abundance of benthic Foraminifera in marine environments,
where they are often major contributors to meiofaunal biomass
(MURRAY, 2006), makes them a potential food source. Some
predators have been identified, including nematodes, poly-
chaetes, mollusks, echinoderms, arthropods and fish. However,
most are incidental predators that ingest foraminifera together
with their food (e.g., deposit feeders, herbivorous), and little is 1 mm
known about selective predation of foraminifera. The presence of
1 mm
foraminifera in the gut of coral reef fish had already been inci-
dentally reported, but only two systematic studies had been carried
out (TODD, 1961; LIPPS, 1988). Based on a small number of fish,
they mostly detected incidental predation. One study, however,
reported a noticeable contribution of foraminifera in the diet of a
nocturnal surface-feeding fish (HOBSON & CHESS, 1973).
During the study reported here (DEBENAY et al., 2011), 291 species
of Foraminifera were identified from more than 20,000 specimens
Tretomphalus squammosus
examined. The only significant nutritional input from with its float chamber
Foraminifera to fish was given by the planktonic Tretomphalus in the thallus of Chamaedoris sp.
phase of some benthic species (fig. 13), which was selectively
ingested by Pomacentrus amboinensis. This territorial fish | Figure 43
Cymbaloporetta growing in an algal turf, at its benthic stage
protects its territory against other fish, allowing foraminifera, and at its Tretomphalus stage (from DEBENAY et al., 2011).
42 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Foraminifera
and environmental changes
100 µm
8 9 10 At a human scale
A 54 cm long core was collected in the Bay of Sainte Marie,
| Figure 45 adjacent to the town of Nouméa, in order to investigate human
Foraminifera from shrimp ponds
1-2: Ammonia tepida, spiral side, impact on sedimentation rates, inputs of contaminants due to
1) normal test, 2) abnormal test; mining activity, and the impact of urbanization on this coastal
3 Ammonia tepida, complex abnormal test;
4-5: Elphidium excavatum, 4) normal test, 5) abnormal test; environment (DEBENAY & FERNANDEZ, 2009). The area selected is
6-7: Caronia exilis, 6) normal test, 7) abnormal test; subjected to urban effluent, and to the input of sediments and
8-10: Quinqueloculina seminula, 8) normal test, 9) abnormal test, brackish water that are transported by wind-driven currents from
last chambers making less than half a whorl,
10) abnormal test, last chamber making more than half a whorl. the estuary of the Coulée River (FERNANDEZ et al., 2006). During the
Scale bar = 0.1 mm (from DEBENAY et al, 2009b). 1950’s, open-cast mining exploration for nickel led to an increasing
input of heavy-metal-rich terrigenous particles in the bays near
Nouméa. Simultaneously, the population of Nouméa increased
dramatically, which may have impacted the neighboring bays.
In surface samples, corresponding to the present conditions,
sedimentary inputs from the Coulée River clearly appear in the
| Figure 46
Boulari and Sainte Marie bays: location map; distribution maps of silt and clay, and Ni, indicative of river input;
distribution maps of the diversity index of foraminiferal assemblages, porcelaneous species and Ammonia tepida.
44 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
distribution of silt and clay, and Ni content (fig. 46). The influence These paradoxical results may be explained by the partial closure
of the river on foraminiferal assemblages can be seen in a lower of the connection between the Bay of Sainte Marie and the Bay of
diversity in front of the estuary, and a lower proportion of porce- Boulari by embankments and sediment accumulation. The
laneous tests in the Boulari Bay. The proportion of Ammonia resulting decrease of the water input from the Bay of Boulari, and
tepida, known to be tolerant to adverse conditions, including consequently from the Coulée River, led to a decrease of freshwater
organic and chemical pollution, and to fresh water input, increases influence and a correlated increase of marine influence in Sainte
towards the mouth of the Coulée River. In Sainte Marie Bay, high Marie Bay. The extension of Nouméa city that was accompanied
relative abundances of A. tepida are associated with organic matter by improvement of the sewage system, a better control of the
content close to or over 20%, which may be (at least partly) related runoff, and embankments in coastal marshes led to a decrease of
to the sewage origin of labile organic matter. freshwater and pollutant inputs and enhanced this process.
Sediment accumulation rates were determined from the decrease in Even if the correlation is doubtful, due to the uncertainties in the
excess 210Pb radioactivity (fig. 47). Two linear regressions of excess sedimentation rates, major rainy events that took place in
210Pb versus accumulated sediment yield different sedimentation
Nouméa since 1940 are correlated with an increase of Haynesina
rates and allow this major environmental change to be dated at depressula, a species tolerant to low salinity. It is inferred that
1956 ± 5 years taking into account the overlaying potentially H. depressula indicates a stronger freshwater impact in the Bay
bioturbated layer. In the core samples, the strengthening impact of Sainte Marie.
of terrigenous loadings from La Coulée River upward is revealed
Finally, the general trend (that can be divided into four main
by an increasing proportion of silt and clay, and of Ni (fig. 47). The
stages) may be explained by both changes in anthropogenic
concomitant increase of organic-bound Zn results from the growth
influences and natural conditions (DEBENAY & FERNANDEZ, 2009). This
of Nouméa city with an extensive use of galvanized corrugated
study showed that anthropic activities, associated with climatic
iron roofs.
events, may have multiple and contradictory impacts on coastal
On the basis of the observations on recent sediment, and of previous environments that could be assessed only by a set of complementary
knowledge about foraminiferal behavior, the trends expected for tools (i.e. geochemistry and bioindicators)
foraminiferal assemblages were: a progressive decrease of species
diversity, a concomitant decrease of porcellaneous tests, and an
At a geological scale
increase of A. tepida, correlative with increasing contamination.
The exact opposite trends were found instead (fig. 47), indicating a Foraminifera were part of a multiproxy analysis of three littoral
change towards less restricted environmental conditions, i.e. under cores from western New Caledonia (WIRRMANN et al., 2011). This
stronger marine influence and lesser freshwater and pollutant study showed that, since the late Holocene sea-level rise, the main
impact. controlling factors of environmental changes were sea-level
10
15
20
15
25
35
10
20
30
40
50
60
-5
-4
-3
0
2
4
6
8
10
12
14
16
18
20
22
24
26
28 1956 + 5
30 AD
32
34
36
38
40
42
44
46
48
50
52
54
Depth (cm)
| Figure 47
Changes in chemical and foraminiferal parameters along the core N12 in Sainte Marie bay
(modified from DEBENAY & FERNANDEZ, 2009).
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 45
change, ENSO variability and extra-tropical phenomena, such as Quinqueloculina spp. and Elphidium spp. indicate a noticeable
the Medieval Warm Period (MWP) marked by a tendency for La marine influence in an open bay. The lack of foraminifera
Niña-like conditions in the tropical Pacific. around 85-80 cm suggests a drying out of the water body. The
presence of gypsum in the sediment is consistent with this
The foraminiferal assemblages, which were mostly represented in
hypothesis. In the section 80-35 cm, an organic-rich sediment
one of the cores, are typical of coastal bays subject to alternating
containing rare fragments of foraminiferal tests dominated by
variable inputs of freshwater and seawater. They contributed to
Ammonia tepida with a few Quinqueloculina spp. indicates a
the results of this study by giving indications on sea-level changes
separation from the sea. It could represent a eutrophic brackish
(fig. 48).
pond, but reworking of tests from underlying sediments is also
At the base of the core, the foraminiferal assemblage is dominated possible. Among this section, the abnormally rich assemblage
by Ammonia tepida and Bolivina striatula, characterizing with marine species, between 70-65 cm, indicates a landward
coastal environments subject to the influence of continental waters. transport of sediments presumably due to a cyclone or a tsunami.
The lack of foraminifera between 185-165 cm together with the This event is consistent with the observation of an extreme event
absence of thecamoebians suggests a drying out of the water body. in the south of the Grande Terre dated around 4,000 cal yr BP
Between 160-90 cm, the noticeable proportion and the variety of (STEVENSON et al., 2001).
% Quinqueloculina spp.
% Murraynella globosa
20
% Bolivina striatula
% Ammonia tepida
40
60
80
100
120
140
160
180
200
220
240
260
| Figure 48
Changes in foraminiferal assemblages along a core extracted on the western coast of the Grande Terre
(adapted from WIRMANN et al., 2011).
Taxonomy
At the end of the Taxonomy section, the taxonomic list mostly the milioline arrangement of the porcelaneous species makes the
follows the suprageneric classification of LOEBLICH & TAPPAN (1992). distinction easy, except two particular cases: 1) Nubeculina
Agglutinated foraminifera, however are classified following advena, a porcelaneous species can be difficult to distinguish from
KAMINSKI (2004), except for the subfamilies Carterininae and some rectilinear agglutinated tests such as Reophax if its porce-
Zaninettinae, both considered as families and grouped in the laneous neck is not well developed (fig. 50); 2) Miliammina spp.
order Carterinida. Generic assignments are mostly based on the have a milioline arrangement, as Quinqueloculina spp., and
concepts of LOEBLICH & TAPPAN (1988), taking into account some may be confused with the species of Quinqueloculina that bear
specialized works such as PATTERSON & RICHARDSON (1987) for agglutinated material, particularly in SEM pictures (fig. 51). The
unilocular forms, NOMURA (1983) for Cassidulinidae, HAYWARD distinction can be done by the agglutinated tooth of Miliammina,
et al. (1997) for Elphidiidae, REVETS (e.g., 1991, 1992, 1993, 1996) and the brownish color of its cement when the observation is
for various groups, and following PARKER (2009) in including done under a dissecting microscope.
Affinetrina, Agglutinella, Cycloforina, Lachlanella, Praemassilina,
Siphonaperta, and Varidentella into Quinqueloculina. The
thorough discussions provided by this author about the taxonomic
attribution of most of his 404 species were also very useful. The
list of synonymies is provided with up to four references for each
species (except a few species that needed a little more), including
reference to the original type description, and publications that
have illustrated the species.
1 2 3 4
| Figure 49
1 Typical white and opaque porcelaneous test;
2 Translucent porcelaneous test;
3 Typical transparents and shiny hyaline test;
4 Whitish and translucent hyaline test. Scale bar = 100 µm.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 49
100 µm
| Figure 52
Growth of Nummulites venosus:
A juvenile; B dissected adult showing a younger stage corresponding to a previous whorl; C adult.
50 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
The following example shows that serious ambiguities still exist Photographical summary
in foraminiferal classification. The gamonts of some species of
the Rosalinidae (simple trochospiral tests) and Cymbaloporidae Agglutinated species
(a trochospiral stage followed by annular series of chambers)
develop a subhemispherical final float chamber and adopt a The 174 agglutinated species are presented on three pages
planktonic stage of life (fig. 13). The genus Tretomphalus (plates 1 to 3). They are arranged in order of increasing
was originally proposed by M OEBIUS (1880) for species that complexity of the tests. First are the unilocular tests: tubular
differed from “Discorbina” (forms with a simple trochospiral rectilinear, tubular coiled, flask-like and subspherical. The
test) only in possessing the float chamber. The type species was following plurilocular tests are successively:
T. bulloides = Rosalina bulloides d’Orbigny, 1839 (BANNER – uniserial throughout;
et al., 1985). – mostly uniserial but with a different initial portion either
biserial, triserial or spiral;
The inflated float chamber was such a conspicuous feature of
the test of Tretomphalus that subsequent identifications have – biserial throughout;
been mostly based upon it, including in this genus species of – mostly biserial but with a different initial portion either triserial
Rosalinidae as well as species of Cymbaloporidae. CUSHMAN or spiral;
(1934) recognized two groups of species, one with a trochospiral – triserial throughout;
coiling, very definitely connected with “Discorbis”, and the other – trochospirally coiled;
that would probably be described as Cymbaloporetta in the early
– planispirally coiled;
stages. Despite that, there is still much confusion about
Tretomphalus, all the more that LE CALVEZ (1977) illustrated – with a milioline arrangement;
Rosalina bulloides (= Tretomphalus bulloides) with specimens – trochospiral, but with a particular test made up of agglutinated
that were not topotypes, and that were Cymbaloporidae with a spicules.
balloon chamber instead of Rosalinidae. More recently, BANNER The last species have irregular attached tests, the morphology of
et al. (1985) grouped the species with a float chamber into which greatly depends on the substrate.
four taxa depending on the chamber arrangement of the coiled
portion, and on the structure of the float chamber. Two (two
subgenera) belonged to Rosalinidae: Rosalina (Tretomphalus) Porcelaneous species
and Neoconorbina (Tretomphaloides); and two (one genus and The 233 porcelaneous species are presented on five pages
one subgenus) belonged to Cymbaloporidae: Cymbaloporetta (plates 4 to 8). Owing to the complexity of this group, morpho-
and Cymbaloporetta (Millettiana). HANSEN and REVETS (1992) logical subgroups have been made. They are successively:
suggested that the float chamber is not a taxonomically valid
– spiral, surface smooth or slightly ornamented;
characteristic. They considered Tretomphaloides a junior
synonym of Neoconorbina and Tretomphalus a junior synonym – discoidal;
of Rosalina. PARKER (2009) suggested reinstating Tretomphalus – elongated, mostly rectilinear;
for Rosalina-Neoconorbina-like species that develop a float – planispiral, elongated along the axis of coiling;
chamber with Tretomphaloides as a junior synonym. He observed – planispiral, then uncoiled, striate;
that the apertural position in these species, inset slightly from the
– milioline, then spiral, a few chambers per whorl, later may be
periphery, is intermediate between Neoconorbina and Rosalina.
uncoiled, aperture simple;
This short summary about the status of Tretomphalus clearly
shows that only future studies that combine lifecycle studies with – milioline, then spiral, a few chambers per whorl, aperture mul-
morphological and molecular systematics can truly resolve this tiple, a trematophore;
issue. – milioline, then spiral, two chambers per whorl or more than
two chambers per whorl;
Actually, only the planktonic stage of Millettiana milletti
– planispiral evolute;
(Cymbaloporidae) is easily distinguished from other species
with a float chamber due to its vermicular overgrowths and – planispiral involute;
irregularly positioned pores. Its benthic stages have thickened, – triloculine;
limbate sutures on the spiral side and chambers are mushroom- – quinqueloculine, test smooth or moderately rough;
shaped on the umbilical side. Other species with a float chamber – quinqueloculine, test ornamented;
have been tentatively attributed to species of Neoconorbina
– quinqueloculine test striate / costulate;
(trochospiral hyaline forms) or Cymbaloporetta (other hyaline
forms) on the basis of only morphological characteristics of the – cryptoquinqueloculine;
coiled portion. – quinqueloculine, test agglutinated or distinctly rough;
– milioline with apertural flap;
This example, which is not unique in Foraminifera, shows that
a substantial improvement of nomenclatorial stability is still – milioline, more than 5 chambers visible;
necessary. – attached or irregular.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 51
– test globular circular in cross section, with a very short neck, if – test low trochospiral with flattened umbilical face (e.g. Ammonia,
at all, surface ornamented by a reticular pattern of costae; Neoconorbina);
– spiral side partially involute (e.g., Pseudononion);
– test globular, the apertural end somewhat truncated.
– surface of the test with depressed, thinner areas (e.g.
The last species are plurilocular, but generally appear as if they
Mississippina);
were unilocular and for this reason have been grouped with
unilocular species. – chambers increasing rapidly in size as added in a flaring
trochospiral coil (e.g., Cancris);
– spiral side flattened and umbilical side convex (e.g., Planulina,
Hyaline uniserial species Cibicides);
The 50 hyaline uniserial species are presented on one page – test biconvex (e.g. Hoeglundina);
(plate 11). Most of the species are typically uniserial, elongated, – aperture surrounded by radiating groves (e.g. Heronallenia);
rectilinear or slightly curved. However, are also included in this
– test high trochospiral (e.g. Elongobula);
group some tests with particular characteristics:
– surface strongly ornamented, the ornamentation obscuring the
– chambers strongly overlapping, the last chamber occupying
sutures and even the chambers (e.g. Calcarina);
most of the test surface;
– chambers distinct, strongly hispid and may be spinose (e.g.,
– test compressed with chambers increasing rapidly in width
Murrayinella);
giving the test a flabelliform shape;
– test with a subspherical balloon chamber.
– initial portion with a different arrangement, mostly planispiral.
Unfortunately, due to the complexity of this group, this grouping
is only tentative and does not include all the species. A careful
Hyaline biserial species observation will be necessary.
| Plate 18
Hyaline trochospiral species. Scale bar = 0.1 mm.
– test compressed, involute, chambers increasing rapidly in size Other hyaline species
as added in a flaring coil;
The 85 hyaline species that could not be included within the
– test much compressed, flattened, evolute, at least in the later preceding groups are presented on two pages (plates 21 and 22).
portion; They include:
– test compressed, involute, lenticular, aperture terminal, radi-
– test biserially arranged and spirally enrolled (e.g.,
ate, with a larger slit on the apertural face;
Globocassidulina);
– test lenticular, very low trochospiral, appearing as if it was
– test biserially arranged, coiled in the early portion, later uncoiled
planispiral (Amphistegina);
biserial (e.g., Ehrenbergina);
– test planispiral in the early portion, tending to uncoil, becom-
– test with uniserial or biserial arrangements, but twisted or
ing uniserial in the later portion.
distorted, which make the arrangement difficult to be identified;
– test composed of multiple chamberlets variously arranged into
discoidal, spherical, branching or irregular tests.
70 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
| Plate 22
Other hyaline species. Scale bar = 0.1 mm.
74 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Ammobaculites
Ammobaculites agglutinans
Test elongate, early planispiral portion
relatively small, compressed, excavated at
the umbilici, with rounded periphery;
linear portion cylindrical, slightly
increasing in size toward the distal end,
up to 6-7 chambers; wall made up of
coarse sand grains, surface rough; aper-
ture terminal, central, simple.
Bays, coastal lagoons.
Systematics p. 256. Ammobaculites exiguus
Aggerostramen Test small, elongate, early portion close
Aggerostramen rustica coiled, later rectilinear, rounded in sec-
Only isolated chambers of this multilocu- tion; wall coarsely agglutinated; aperture
lar test have been collected. Chambers terminal, rounded.
polyhedral, more or less angular, con- Coastal lagoons, mangrove swamps,
structed almost entirely of sponge estuaries.
spicules neatly cemented together, with Systematics p. 256.
some long spicules that project beyond
the chamber itself; in the early stages,
chambers attached to the substrate in
uniserial series, later chambers some-
what irregular in arrangement, intercon-
nected by tubular stolons; aperture a
simple opening, or at the end of stolon-
like necks.
Northern shelf, 600 m.
Systematics p. 253.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 75
Ammolagena
Ammobaculites Ammolagena clavata
cf. A. subcatenulatus Ammodiscus Test commonly attached to shell fragments,
Test small with an elongate cylindrical Ammodiscus gullmarensis up to 1 mm in length, large ovoid pro-
rectilinear stage of constant diameter, loculus followed by a narrower tubular
positioned symmetrically above the coiled Test free, small, consisting of several chamber, generally rectilinear at it first
initial stage; coil somewhat compressed, slightly overlapping whorls; sutures dis- stage; wall finely agglutinated, smoothly
slightly larger in diameter than the recti- tinct; test flattened, slightly biconcave, finished, reddish-brown in color; aperture
linear stage; wall coarsely arenaceous, tending to irregular coiling in last terminal, rounded.
mostly composed of fragments of sponge whorls; periphery rounded; wall aggluti-
nated with fairly large amount of cement; Northern shelf, 600 m.
spicules, inflated and circular in cross Systematics p. 253.
section; chambers indistinct, sutures aperture semicircular at the open end of
obscured by the coarse agglutinate; the tubular chamber.
aperture terminal, in the middle of the Southwestern lagoon, 30 m.
apertural face. This species resembles Systematics p. 253.
Ammobaculites cf. A. subcatenulatus,
but differs in the less distinct chambers
and the agglutinated sponge spicules.
Coastal bays.
Systematics p. 256.
76 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Ammotium
Ammotium fragile
Test small, compressed, planispirally
Ammoscalaria enrolled and evolute in the early stage,
Ammoscalaria compressa later uncoiled, rectilinear; sutures distinct,
Test elongate, much compressed; periphery curved back toward the proloculus at the Arenoparrella
subacute; chambers indistinct, increasing inner margin, becoming chevron-like in Arenoparrella mexicana
rapidly in size as added, later becoming the later chambers; wall thin, fragile finely
broad and chevron-shaped; wall coarsely agglutinated; aperture ovate, terminal, at Test free, in a low trochospiral coil,
agglutinated, surface rough; aperture the dorsal angle of the final chamber. chambers increasing gradually in size,
narrow, terminal, elongate, produced. sutures radial, periphery rounded; wall
Coastal lagoons, mangrove swamps. finely agglutinated, surface smooth and
Northern shelf, 600 m. Systematics p. 256. polished; primary aperture a straight to
Systematics p. 256. curved slit surrounded by a thin and
delicate lip, beginning near the base of
the apertural face and directed upward
across the median plane with an angle to
the plane of coiling, supplementary
openings present at the apex of the final
chamber.
Mangrove swamps, marshes.
Systematics p. 259.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 77
Caronia
Caronia exilis
Short triserial initial stage with minute
subglobular chambers; later biserial with
about 6 pairs of appressed globular Clavulina
Bigenerina chambers separated by depressed sub- Clavulina difformis
Bigenerina nodosaria horizontal sutures; test rather coarsely Test elongate with a short triangular tris-
Test elongate, the early portion composed agglutinated; aperture a symmetrical erial stage; the later uniserial stage polyg-
of a biserial group of chambers, consider- interiomarginal elongate arch at the base onal to rectangular in cross section;
ably flattened, increasing progressively in of the last chamber. chamber walls slightly concave, extend-
size; later portion composed of an unise- Coastal lagoons, mangrove swamps, ing back along the angles of the test,
rial series of chambers, circular in cross estuaries. resulting in a lobate outline; sutures dis-
section, usually less in width than the Systematics p. 259. tinct, depressed; wall roughly textured;
biserial portion; wall usually coarsely aperture terminal, centered, with a single
arenaceous; aperture terminal, small, valvular tooth.
rounded. Coral-reef lagoon, mostly near coral
Northern shelf, 200 m. reefs.
Systematics p. 263. Systematics p. 262.
78 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Cribrostomoides
spiculotestus
Test free, compressed, planispiral, not
completely involute; umbilical region
Connemarella depressed, periphery rounded, broadly
Connemarella rudis lobulate; chambers slightly inflated,
Test conical, with early triserial arrange- increasing gradually in size as added;
ment followed by broad and low biserial sutures distinct, slightly depressed and
chambers that increase rapidly in slightly curved; wall thin, composed of
breadth and height in the early portion, fine sand grains and sponge spicules of
less rapidly later; apertural face flattened, various sizes, surface neatly finished;
Clavulina pacifica circular; sutures horizontal obscured by aperture crescentic, areal, slightly above
the agglutinate material; wall coarsely the base of the apertural face, bordered by
Test elongated with an initial portion a well-developed lip.
triserial becoming uniserial at about one agglutinated with calcareous cement;
third from the pointed apical end; test aperture a broad low arch, in a reentrant Outer coral reef, 100 m.
triangular in cross section; chambers at the base of the apertural face. Systematics p. 257.
slightly inflated and strongly curved Northern shelf, 600 m.
backwards at each corner; sutures distinct Systematics p. 262.
in the triserial portion, depressed in the
uniserial part; wall finely arenaceous;
apertural face slightly convex; aperture
central, rounded with a simple toothplate.
Coral-reef lagoon, mostly near coral reefs.
Systematics p. 262.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 79
Cyclammina
Cyclammina subtrullissata
Test nautiloid, compressed, planispiral,
involute, slightly depressed in the umbili-
cus; peripheral margin more or less Cylindroclavulina
rounded; 6-9 chambers in the last whorl, Cylindroclavulina bradyi
separated by slightly depressed, somewhat Test stout, cylindrical, the early triserial
sinuate sutures; wall arenaceous with portion not well shown exteriorly, the
heterometric grains, surface smoothly later uniserial section large, with distinct Discammina
finished, colour brown; aperture crescentic, depressed sutures; wall composed of hetero- Discammina compressa
at the base of the last-formed chamber. metric sand grains, usually with a smooth Test planispiral, compressed, lenticular,
Northern shelf, 600 m. exterior; aperture at the end of a short somewhat involute, depressed in the
Systematics p. 260. neck, usually 3 or 4 radiate slits. umbilicus; peripheral margin acute or
Bay of Prony 10-30 m. somewhat rounded; chambers and sutures
Systematics p. 263. not distinct; wall coarsely agglutinated,
surface rough, color brown; aperture
crescentic, at the base of the last-formed
chamber.
Northern shelf, 600 m.
Systematics p. 256.
80 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Haddonia
Haddonia torresiensis
Test large, attached, early stage coiled,
later uncoiled, uniserial, irregular, but
complete specimens are missing; chambers
broad and low, irregular in size and
shape, at least twice as wide as high; wall
coarsely agglutinated, with considerable
cement, wall pierced by numerous large
pores aligned perpendicular to the surface;
aperture terminal, areal, an irregular slit.
Southwestern lagoon, outer reef, 50-100 m.
Glomospira Systematics p. 261.
Glomospira fijiensis
Gaudryina sp. 1 Proloculus followed by an undivided
Test elongate, conical, early stage triserial tubular chamber that is irregularly strep-
and subtriangular in section, later tospirally coiled; wall finely agglutinated;
becoming biserial and rounded in section; aperture at the open end of the tube.
chambers increasing regularly in size as Sometimes referred to G. glomerata, but
added, giving a very regular conical differs from this species in lacking the
shape to the test; wall agglutinated, meandering enrolment of the tube.
roughly finished; aperture an arch at the Coastal lagoons, mangrove swamps.
inner margin of the final chamber, Systematics p. 254.
Northern shelf, 600 m.
Systematics p. 260.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 83
Hemisphaerammina
Hemisphaerammina bradyi
Test attached, a single hemispherical
chamber; wall agglutinated, smooth due
to considerable cement; no apparent Hyperammina
aperture, communication with the exterior
probably occurs through interstitial Hyperammina friabilis
pores. Test elongate, subcylindrical, straight,
Haplophragmoides pusillus often tapering toward the apertural end;
Southwestern lagoon, 40 m.
Test small, planispirally enrolled, involute Systematics p. 252. rather large globular proloculus followed
becoming slightly evolute, compressed by an elongate, sub-cylindrical second
and biumbilicate, chambers inflated and chamber, slightly less in diameter than
margin distinctly lobulate; wall thin, the proloculus; wall thick, loosely
with moderately coarse agglutinate, exte- cemented, composed of sand grains with
rior slightly rough; aperture an elongate a varying amount of sponge spicules;
low equatorial slit at the base of the aperture rounded, at the end of the
apertural face, with a slight lip. chamber.
Coastal bays, shallow coastal areas. Northern shelf, 600 m.
Systematics p. 256. Systematics p. 253.
84 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Karreriella Lagenammina
Karreriella bradyi Lagenammina arenulata
Test stout, somewhat elongate, tapering Test free consisting of a single oval, flask-
very slightly until near the initial end shaped chamber; wall coarsely arenaceous,
where it tapers abruptly to the somewhat formed of closely agglutinated sand
blunt end; triserial portion nearly circular grains of variable size and roughness;
in cross section, of few chambers; the aperture a terminal, rounded, projecting
later biserial portion making up about opening, without a distinct neck.
three fourths of the test, slightly com- Outer coral reef, 100 m.
pressed; chambers overlapping, broadly Systematics p. 252.
elliptical in cross section, inflated;
sutures depressed; wall finely arenaceous,
smooth; aperture oval, slightly back from
the inner margin of the last chamber,
with a border raised somewhat and
thickened, supplementary apertures
present on the apertural face.
Jaculella Northern shelf, 600 m.
Jaculella obtusa Systematics p. 261.
Test elongate, tubular, proximal end
closed, obtusely rounded, distal end
slightly broader; wall thick, agglutinated,
with firmly cemented grains, surface
rough; aperture at the open end of the
tube.
Northern shelf, 700 m.
Systematics p. 253.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 85
Martinottiella
Martinottiella bradyana
Test elongate, cylindrical, not tapering;
early trochospiral coil with four to five
chambers per whorl, later reduced to
triserial, biserial, and an elongate final
uniserial stage with chambers numerous,
fairly distinct, increasing somewhat in
height as added; sutures distinct, but very
slightly depressed; wall arenaceous,
Lagenammina spiculata slightly roughened; apertural face convex,
Test unilocular, rounded or oval; wall aperture terminal, central, rounded,
agglutinated, mostly composed of sponge produced on a distinct neck.
spicules, including projecting spicules; Northern shelf, 600 m.
aperture somewhat produced. Lituotuba Systematics p. 261.
Northern shelf, 700 m. Lituotuba lituiformis
Systematics p. 252.
Test free, early portion with irregular to
planispirally coiled tubular chamber,
finally becoming uncoiled and irregularly
rectilinear; wall finely agglutinated
with a yellowish-brown cement, surface
smoothly finished; aperture rounded at
the open end of the tubular chamber.
Southeastern coast of the Grande Terre
30 m.
Systematics p. 255.
86 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Monotalea
Monotalea salsa
Early stage biserial, later uniserial and
rectilinear; uniserial cylindrical chambers
radially-symmetrical in cross section,
separated by horizontal sutures; uniserial
stage increasing only little in size in course
of growth; wall finely agglutinated, thin; Nouria harrisi
aperture terminal, large, rounded.
Test fusiform, circular to sub-circular in
Mangrove swamps, marshes. section; only two or three chambers visible,
Systematics p. 258. spirally arranged around the long axis of
Miliammina
the test, the ultimate chamber terminating
Miliammina fusca in a somewhat produced neck; sutures
Test with a quinqueloculine arrangement, slightly depressed, but well marked,
elongate and ovate in section, with a owing to the divergent angles at which
rough surface; wall composed of well- the spicules are arranged in adjacent
sorted agglutinated grains; aperture chambers; wall constructed entirely
terminal ovate with a small simple, of sponge spicules, arranged roughly
agglutinated tooth. parallel to the long axis of the test;
Low salinity estuaries and coastal lagoons. aperture terminal, rounded.
Systematics p. 254. Southeastern coast of the Grande Terre
and northern shelf, 60-600 m.
Systematics p. 258.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 87
juvenile
Paratrochammina
Paratrochammina
cf. P. simplissima
Test free, chambers in a trochospiral coil,
periphery rounded; generally 3 coils with
4-5 chambers in the last-formed coil;
sutures slightly depressed, oblique on the Plotnikovina transversaria
spiral side, radial on the umbilical side; Test elongate, with short early triserial
wall agglutinated, smoothly finished; Placopsilina and triangular stage and later biserial
aperture single, interiomarginal, umbilical- Placopsilina bradyi stage, with a rhomboid section, chambers
extraumbilical, extending across the broad and low, lateral extremities distinct-
Test attached, early stage planispirally
umbilicus over the margin of the penul- ly produced at the chamber angle, an
enrolled, later uncoiled, uniserial, in an
timate chamber, but often obscured by internal vertical partition then isolating
irregular course over the base of attach-
agglutinated material deposited in the a small distal chamberlet, the tip of the
ment; chambers distinct, regularly added,
umbilical depression. chamberlets commonly broken to leave a
inflated, rounded in section, increasing
Southwestern lagoon and outer reef, very little as added; wall coarsely aggluti- small opening at the surface; sutures
20-100 m. nated, but surface fairly smooth; aperture slightly depressed, straight, at a slight
Systematics p. 258. terminal, rounded. angle from the horizontal; wall finely
Coral-reef lagoon and outer reef, 30-90 m. agglutinated; aperture a low basal arch
Systematics p. 257. with a distinct lip.
Outer reef and deep parts of the lagoon,
30-100 m.
Systematics p. 262.
88 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Pseudobolivina
cf. P. nasostoma
Test slender and elongate, biserial, tending
Psammosphaera to become uniserial, slightly compressed,
Psammosphaera parva slightly twisted and curved; periphery
Test unilocular composed of a small round, initial end acute; chambers
subsphaerical chamber; wall agglutinated, numerous, the early ones small and
the agglutinate including an elongated compressed, increasing rapidly in width Pseudogaudryina
spicule projecting on both sides of the but slowly in height, the later chambers Pseudogaudryina concava
test. inflated, increasing rapidly in height;
Test triangular in section throughout
sutures depressed; wall finely agglutinated,
Northern shelf, 600 m. with slightly concave faces and sharp,
thin, delicate; aperture subcircular, at the
Systematics p. 252. somewhat serrate edges; chambers not
end of an elongated projection.
inflated, sutures indistinct, later chambers
Northern shelf, 700 m. developing a slightly overhanging margin
Systematics p. 258. giving the appearance of excavations in
the lower part of the chamber; wall
agglutinated; aperture a slit in a shallow
re-entrant in the middle of the inner
margin of the last chamber.
Northern shelf, 600 m.
Systematics p. 262.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 89
Reophanus
Reophanus oviculus
Test composed of a few pyriform chambers
arranged in a rectilinear series, each
having a more or less prolonged tubular
neck; tests often broken with no more
than three chambers found together; wall
arenaceous, often rough externally with Reophax bacillaris
Pseudothurammina Test elongate, regularly tapering, some-
projecting sponge spicules incorporated
Pseudothurammina limnetes with the sand; aperture terminal rounded, what curved in the megalospheric form,
Test unilocular, subglobular with finely at the end of a long neck. with a decided angle in the microspheric
agglutinated silt grains; aperture at the Northern shelf, 600 m. form; microspheric form very tapering to
end of 1 to 3 tubular projections projecting Systematics p. 254. the initial end, megalospheric form with
from the chamber. early chambers larger than those imme-
Low salinity coastal lagoons and low diately succeeding, giving to the test the
marshes. appearance of Clavulina; chambers
Systematics p. 252. numerous (up to 30), short, indistinct in
the earlier portion, later separated by
depressed sutures; wall finely arenaceous;
aperture terminal indistinct. The figure
represents a megalospheric form.
Northern shelf, 600 m.
Systematics p. 254.
90 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Reophax dentaliniformis
Test long, arranged in a straight line, Reophax irregularis Reophax nana
slender and tapering, circular in cross Test large, straight, almost cylindrical; Test small elongate, with few rounded
section; around 6 subcylindrical cham- chamber gradually increasing in size, chambers in slightly irregular series; wall
bers that have a little overlap onto the apertural end slightly tapered; horizontal coarsely agglutinated; aperture terminal,
preceding chambers; ultimate chamber sutures faintly indicated; wall rough with rounded, produced on a slight neck.
tapering gradually into a distinct neck; large to small particles; aperture terminal,
wall irregularly agglutinated, but neatly Coastal lagoons, estuaries, shrimp ponds.
produced, centered. Systematics p. 255.
finished; aperture terminal, rather large,
produced, rounded. Northern shelf, 600 m.
Systematics p. 255.
Northern shelf, 600 m.
Systematics p. 255.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 91
Rotaliammina
Rotaliammina chitinosa
Attached test, dark brown in color, with
chambers arranged in trochospiral
coiling, increasing gradually in size
throughout four whorls; only 5 to
8 chambers in the final whorl; chambers
with a petaloid shape on the umbilical
side, but chamber walls often collapsed or
missing; walls made of relatively coarse
and irregular agglutinated material,
Sahulia peritubula
entirely organic on the umbilical side; Test a low cone, broader than high;
test surrounded by a thin organic flange sutures fairly distinct, marked by the
with agglutinated material that is rapidly Saccorhiza openings of a single row of short periph-
destroyed after death; aperture hardly Saccorhiza ramosa eral tubes; wall coarsely arenaceous, sur-
distinguishable, terminal on the produced Test free with an ovoid proloculus that is face rough; apertural face nearly circular,
end of the chambers and facing the generally broken; the following tubular aperture a short slit at the base of the last-
umbilicus. chamber branches irregularly; wall formed chamber, with a narrow lip.
Outer reef, and Bay of Prony, 10-100 m. agglutinated with sand grains and Northern shelf, 200 m.
Systematics p. 259. sponge spicules, some of them arranged Systematics p. 263.
perpendicular to the wall so that they
project laterally; apertures at the end of
the tubes.
Outer reef, 80 m.
Systematics p. 253.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 93
Siphotextularia flintii
Test triangular in outline, irregularly
juvenile rhombic in end view, somewhat com-
juvenile pressed laterally, rapidly enlarging in size
from the early portion; chambers low and
broad, inflated, separated by rather deep
Septotrochammina sutures; wall finely agglutinated,
Septotrochammina gonzalesi Siphotextularia smoothly finished; aperture slightly
Test attached, depressed trochospiral,
Siphotextularia blacki above the inner base of the last chamber,
chambers numerous, umbilicus open, Test large, widening regularly from the with a slightly raised lip.
numerous radial secondary septa of four acute initial end, compressed; faces of the Northern shelf, 600 m.
to five orders produced by invaginations test concave, borders of chambers much Systematics p. 265.
from the peripheral wall; wall thin, flexible, thickened and appearing as rounded
proteinaceous, incorporating a very small keels; wall finely agglutinated, coarser
amount of agglutinated silt; aperture agglutinate on the keels; aperture elon-
interiomarginal, at the umbilical tip of gate, above the inner base of the last
the final chamber. chamber, with a raised lip.
Outer reef, 100 m. Northern shelf, 600 m.
Systematics p. 259. Systematics p. 265.
94 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Siphotextularia pulchra
Test compressed, periphery rounded
Siphotextularia heterostoma throughout, lobulate, initial end broadly
Test elongated, laterally compressed, rounded; chambers wider than high, Siphotrochammina
particularly in the median line, triangular increasing gradually in width, slightly
in lateral view, with rounded edges in end Siphotrochammina lobata
inflated; sutures depressed, curved, those
view; chambers increasing in breath, of the early portion not very distinct; Test free, low trochospiral, chambers ovate,
more rapidly in height in later stages; wall finely arenaceous, smooth, coarser increasing gradually in size as added,
some specimens have a triserial initial agglutinate in the initial portion; aperture sutures gently curved, periphery rounded,
stage; most of the tests appear twisted a transverse elliptical slit, slightly above much lobulate; wall light brown in color,
with respect to the axis; sutures distinct, the base of the apertural face, with finely and sparsely agglutinated on a
oblique; wall rather smoothly finished; prominent lips. proteinaceous base, the early whorl darker
aperture an elliptical slit parallel to the brown and with very little agglutinated
Northern shelf, 600 m. material; aperture interiomarginal, at the
lateral compression, at the end of an Systematics p. 265.
everted neck. end of a siphon-like lobe projecting from
Northern shelf, 600 m. the umbilical margin of the chamber
Systematics p. 265. and directed forward.
Mangrove swamps, crawling on the aerial
roots of the mangrove trees.
Systematics p. 259.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 95
Textularia
Textularia agglutinans
Test elongate, tapering, very little com-
pressed, periphery rounded; chambers
inflated, increasing in width and height
Spirotextularia toward the apertural end rather uniformly;
Tawitawia sutures distinct, depressed, usually about
Spirotextularia fistulosa right angles to the long axis of the test;
Test with planispiral coil of one whorl, Tawitawia immensa wall rather coarsely agglutinated, but
then biserial; each chamber laterally Test biserial, flattened, very large, more smoothly finished; aperture an elongate slit
produced, forming a distal chamberlet than 4 mm in length; chambers low, in a well-marked depression of the inner
separated from the main chamber lumen strongly overlapping in the axial area; margin of the last-formed chamber.
by a secondary septum; wall agglutinated, wall coarsely agglutinated; aperture a slit Widely distributed in the southwestern
surface smoothly finished; aperture on the apertural face of the last chamber. lagoon and southern shelf, 0-80 m.
interiomarginal, a low arch against the Northern shelf, 600 m. Systematics p. 263.
previous chamber. Systematics p. 265.
Southwestern lagoon and northern shelf,
20-600 m.
Systematics p. 258.
96 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Textularia fistula
Textularia candeiana Textularia cushmani Test elongate, the early chambers some-
Early portion of the test narrow, much Test elongated, slender, narrowly triangular what compressed, later rounded in section;
compressed, the edges almost carinate, in lateral view, sub-quadrangular in cross early chambers with lateral fistulose
tapering gradually to the apex; chambers section with rounded margins, biserial projections that become extended into a
numerous, those of the early portion throughout; about 20 chambers increasing projecting peripheral border in later
somewhat compressed, later ones enlarging rapidly in the earlier portion, more chambers; wall coarsely arenaceous; aper-
rapidly, with the final ones much inflated; gradually so in the later portion, the ture slitlike in a well marked depression
sutures of the later portion fairly distinct peripheral margins becoming nearly of the inner border of the last-formed
and oblique, depressed; wall rather parallel; sutures depressed, nearly per- chamber.
coarsely arenaceous; aperture a broad, pendicular to the axis, often indistinct; Northern shelf, 600 m.
low arch at the base of the last chamber, aperture a short low slit at the base of the Systematics p. 264.
bordered by a narrow lip on the upper inner margin of the last formed chamber.
margin. Northern shelf, 200 m.
Widely distributed in the southwestern Systematics p. 264.
lagoon and on the northern shelf, 20-
200 m.
Systematics p. 263.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 97
Textularia lateralis
Test characterized by a sub-triangular
shape with a neat, even outline; chambers
Textularia goessi broad and low; sutures horizontal and
Test broadly conical in outline, tapering slightly depressed; wall finely agglutinated. Textularia oceanica
abruptly to the apex with margins slightly Southwestern lagoon, rare, 5-40 m. Test biserial throughout; laterally com-
convex, subcircular in end view; chambers Systematics p. 264. pressed; chambers wider than high,
low and broad without internal division; rounded, increasing gradually in size;
sutures limbate, distinct; wall more or sutures depressed, oblique obscured by
less coarsely agglutinated, but usually the agglutinated material; wall coarsely
smoothly finished; aperture linear in a agglutinated, very roughly finished;
depression at the base of the apertural aperture an arch at the base of the
chamber. apertural face, in a slight re-entrant.
Northern shelf, 600 m. Southwestern lagoon and southern shelf,
Systematics p. 264. in the back-reef area and near patch
reefs, 5-60 m.
Systematics p. 264.
98 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Textularia semialata
Test compressed, rapidly broadening,
triangular in side view; chambers broader
than high, the proximal outer angle of
the adult chambers more or less projecting
and extending backward; wall of fine
sand, very smoothly finished; aperture an Textularia subantarctica
Textularia pseudogramen arched opening at the inner margin of Test a low cone, as broad as high, strongly
the chamber, with a distinct overhanging compressed laterally, periphery acute to
Test biserial, large, elongate, chambers
lip; color gray. carinate; initial end broadly rounded in
rapidly increasing in size in the earlier
portion, then remaining of almost con- Southwestern lagoon and northern shelf, side view; sutures distinct, very slightly
stant dimension for 2/3 of the test; 10-200 m. depressed; wall finely arenaceous, surface
periphery subacute in the early portion, Systematics p. 264. smooth; apertural face ovate, aperture a
later thickening; chambers numerous short slit in a depression at the base of the
(about 10 pairs) separated by distinct last-formed chamber, with a narrow lip.
sutures; wall coarsely agglutinated, Northern shelf, 600 m.
surface rough; apertural face smoothly Systematics p. 264.
finished; aperture a low arch at the base
of the apertural face.
Southwestern lagoon and northern shelf,
5-500 m.
Systematics p. 264.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 99
Textularia sp. 3
Textularia truncatiformis Test elongate, tapering, very little com-
pressed, periphery rounded; chambers
Test elongate, slightly compressed, initial inflated, increasing in width and height
end rounded, increasing slightly in width toward the apertural end rather uniformly;
and thickness; periphery broadly rounded, sutures indistinct in the early portion,
a great part of the test of almost equal Textularia sp. 1
deeply incised in the later portion, about
width, the two sides being nearly parallel; Test stout, biserial throughout, subconical right angles to the long axis of the test;
chambers numerous, the early ones not in general shape, subcircular in end wall coarsely agglutinated, surface
distinct, the later ones distinct, wider view; periphery broadly rounded; about rough; aperture a slit in a well-marked
than high; wall composed of very coarse 10 chambers in the adult, not inflated; depression of the inner margin of the
grains; apertural end obliquely truncate, sutures indistinct, slightly depressed, last-formed chamber.
aperture a low slit at the base of the last slightly curved, nearly perpendicular with
chamber. the test axis; wall coarsely agglutinated, Northern shelf, 600 m.
roughly finished, even on the flattened Systematics p. 265.
Northern shelf, 600 m.
Systematics p. 265. apertural end; agglutinated material
composed of large grains interspersed
with finer grains; aperture a low arch at
the base of the inner margin of the last
formed chamber, in a re-entrant. This
species is quite similar to Textularia sp.
“M” of HOTTINGER et al. (1993).
Northern shelf, 600 m.
Systematics p. 265.
100 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 101
Valvulina
Valvulina oviedoiana
Trochammina sp. 1 Test subconical, stout, triserial, triangular
Test low trochospiral, chambers low, in section; about 15 chambers, initially
increasing gradually in size as added, with angular margins, becoming rounded;
sutures radial, periphery rounded; wall chambers increasing rapidly in size as
coarsely agglutinated; aperture an interi- added; sutures depressed; agglutinate
omarginal, umbilical-extraumbilical composed of well-sorted, rounded particles;
arch with narrow bordering lip. aperture an interiomarginal arch, at the
junction of the chambers of the final
South of the Grande Terre, 50 m.
Trochammina inflata whorl with a prominent flaplike tooth
Systematics p. 259. projecting from the midpoint of the
Test trochospiral, chambers inflated,
sutures radial depressed, distinct; periphery apertural rim.
rounded; wall finely agglutinated, surface Chesterfield, in algal thalli, 15 m.
smooth; aperture an interiomarginal, Systematics p. 263.
umbilical-extraumbilical arch with narrow
bordering lip.
High marshes, mangrove swamps,
coastal lagoons rich in organic matter.
Systematics p. 258.
Trochammina sp. 2
Test trochospiral, chambers rounded in
section, increasing gradually in size as
added, sutures radial, periphery rounded;
wall with moderately coarse agglutinate;
aperture an interiomarginal, umbilical-
extraumbilical arch with narrow bordering
lip.
South of the Grande Terre, 50 m.
Systematics p. 259.
102 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Zaninettia
Zaninettia conica
On the spiral side, the first stage of the test
is a small, high, conical trochospire,
consisting of about 2 whorls, flattening
out toward the periphery, with a subcircular Unidentified species Alveolinella
outline; it is dark brown to blackish; test Test very low trochospiral; chambers few, Alveolinella quoii
shallow-concave on the umbilical side, three in the last whorl; spiral side partially Test large (up to 20 mm in length),
with a deep axial depression. The following involute; sutures depressed, radial; wall fusiform, planispiral, involute, elongated
chambers are yellowish or white, they rather coarsely agglutinated, but very along the coiling axis; chambers narrow,
make up the flattened second ontogenetic smoothly finished, polished; aperture an spanning the width of the test, thickening
stage, at first also in a trochospire, are interiomarginal extraumbilical slit. toward the poles and gradually increasing
crescentic with secondary septa; the Outer reef, 100 m. in size as added; more than 15 chambers
chambers become increasingly elongated per whorl in adult specimens; wall with
and irregularly added, giving an irregu- numerous longitudinal costae; apertural
larly lobed outline to the test; no peripheral face with several rows of openings.
flange was observed; chambers symmetric, Widely distributed in the southwestern
petal-shaped, on the umbilical side; walls lagoon, 0-50 m.
made up of typically “rounded-rectangular” Systematics p. 281.
truncated spicules arranged irregularly.
Coral-reef lagoon and outer reef, 10-100 m,
crawling under coral rubble or algae,
protected from sunlight.
Systematics p. 282.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 103
Ammomassilina
clypeoarenulata
Test flattened, sutures obscured by the
coarse agglutination; periphery broadly
rounded, planispiral arrangement
obscuring any early quinqueloculine Amphisorus sauronensis
chambers; wall coarsely agglutinated, This species differs from A. hemprichii by
roughly finished; aperture terminal, additional medial rows of apertures on
rounded, with a small bifid tooth, at the the peripheral apertural face. These
end of short neck that is agglutinated apertures vary from irregularly shaped,
as the rest of the test. often fusing with neighboring apertures Articulina cf. A. carinata
Northern shelf, 600 m. to circular. They are absent in juveniles, Test elongate, laterally compressed; early
Systematics p. 279. both species having similar juvenile ovoid portion milioline, latter with a few
stages. uncoiled, elongate, flattened chambers;
Outer reef and Chesterfield, 5-45 m. wall imperforate, surface smooth or with
Systematics p. 282. faint costae; aperture terminal, ovate,
bordered by a prominent everted lip.
Bay of Prony, 15 m.
Systematics p. 280.
104 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Borelis
Borelis schlumbergeri
Test planispiral involute elongate along the
coiling axis; surface ornamented by low
wavy ridges located along the chamberlet
sutures, and by faint pustules in between
the ridges; 4-6 chambers per whorl sepa-
rated by depressed sutures; chambers
divided into 20-40 chamberlets in the
adult; apertural face with one basal row
of circular to irregularly quadrangular
Articulina queenslandica apertures; each aperture rimmed with
Test elongate; early portion milioline, peristomal material; a rectangular to
latter uniserial with up to 4 cylindrical faintly bifid mask hides partly each
chambers, slightly, if at all, widened at Biloculinella apertural opening.
the base; wall porcelaneous; 4 or 5 longi- Biloculinella globula Lifou, Loyalty Islands, 5 m.
tudinal costae on the first uniserial Systematics p. 281.
chamber, up to 12 on the last one; aperture Test biloculine in front view oval with
terminal, rounded, with a narrow everted greatest width at two thirds distance from
lip. the aperture, in end view globular;
chambers much inflated, sutures slightly
Dispersed in the southwestern lagoon, depressed; wall smooth; aperture semi-
5-30 m. circular with a simple flat tooth filling a
Systematics p. 280. large part of the opening.
Northern shelf, 200 m.
Systematics p. 274.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 105
Cornuspiramia
Cornuspiramia
cf. C. antillarum
Test attached; globular proloculus fol-
Cornuspira Cornuspira planorbis lowed by a spiral tubular second cham-
Cornuspira foliacea Test circular composed of a globular ber; later chambers irregularly uncoiled
Test flattened, early portion of the tubular proloculus followed by an undivided, and branching, elongate, cylindrical to
chamber of nearly uniform dimensions, planispiral, evolute second chamber; wall pyriform, attached side flattened, with a
but in later development rapidly increasing with a smooth imperforate surface; aper- marginal keel, free surface convex; wall
in height and forming a broad flat test; wall ture at the open end of the tube. imperforate, milky-white at the periph-
smooth except for occasional thickening From coastal lagoons to outer reef, 0-50 m. ery; aperture terminal. The uncoiled
over the lines of growth; aperture a long Systematics p. 266. chambers are hardly preserved in the sed-
slit at the end of the tubular chamber. iment.
Northern shelf, 600 m. Southern lagoon and outer reef, 3-125 m.
Systematics p. 266. Systematics p. 267.
106 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Edentostomina milletti
Cribromiliolinella Test compressed, ovate in outline, periphery
Cribromiliolinella subvalvularis acute; planispirally enrolled with the
Coscinospira Test ovate in outline, rounded in section; antepenultimate chamber only just visible
Coscinospira hemprichii three chambers visible from the exterior; between the embracing later chambers;
Crosier-shaped test with a shallow umbilical wall imperforate; aperture terminal, with sutures slightly excavate; chambers rapidly
depression in the early involute portion; a flap, the opening extending around the enlarging, one-half coil in length; aperture
uniserial portion cylindrical; sutures flap and up the chamber as an irregularly terminal, elliptical, surrounded by a
radiate, curving backwards near the triradiate opening, the rays of the opening thickened lip, with no tooth.
margin in the coiled portion, straight, also secondarily bifurcating in well- Bay of Prony, 30 m.
transverse in the uncoiled portion; developed specimens Systematics p. 268.
surface covered with strong acute ribs, Northern shelf, 600 m.
perpendicular to the sutures, alternating Systematics p. 274.
irregularly from one chamber to the
other; the ribs fuse with the peristomes of
the multiple aperture that occupy the
center of the apertural face.
Widely distributed in the southwestern
lagoon, 0-45 m.
Systematics p. 281.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 107
Fischerinella helix
Erichsenella Test conical, globular proloculus followed
Erichsenella schauinslandi by spiral chamber of nearly a complete
Test cryptoquinqueloculine in the early whorl, then with gradually enlarging
chambers, later chambers uniserial, more trochospirally enrolled chambers, pro-
or less regularly arranged; early-coiled gressively more numerous per whorl, up
stage usually inflated; later chambers to four or five in the final one; spiral side
flattened on one side and inflated on the convex evolute with all whorls visible;
other side with a rounded peripheral only chambers of the final whorl visible on
Fischerina
margin; aperture Miliolinella-like in the the flattened umbilical side with slightly
Fischerina pellucida depressed umbilicus; spiral suture
early stage, a large terminal opening that
is bordered by a crenulate lip in the adult Test discoidal, planispirally enrolled; depressed, sutures between chambers
stage. globular proloculus followed by enrolled flush to slightly depressed, radial; wall
nonseptate tubular chamber of about one imperforate, smooth or ornamented with
Isle of Pines, 5 m. faint striae; aperture rounded or ovate at
whorl, then by two or more whorls with
Systematics p. 280. the open end of the final chamber.
up to eight chambers each; sutures radial
to slightly curved; each whorl partially From 50 to 600 m.
overlapping the earlier one on both sides Systematics p. 266.
of the almost symmetrical test; wall thin
and imperforate, smooth; aperture at
the open end of the final chamber, often
arcuate in form due to the slightly
involute coiling.
South of the Grande Terre, 50 m.
Systematics p. 266.
108 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Hauerina
Hauerina diversa Hauerina fragilissima Inaequalina
Test with rounded outline, slightly Test partially involute with a highly Inaequalina affixa
biconvex, periphery subacute; initial stage compressed discoid shape; surface of the
quinqueloculine, adult stage planispiral Test elongated with tapering ends; one
wall unevenly pitted; aperture terminal, side flat, the other deeply concave;
with 3-4 crescent-shaped chambers in the cribrate.
last coil, increasing gradually in size, periphery carinate; chambers triangular
slightly overlapping those of the preceding Southwestern lagoon, rare, 20-30 m. in cross section, with concave sides,
coil; surface ornamented with longitudinal Systematics p. 270. planispirally enrolled, increasing rapidly
anastomosing and transverse microstriae; in size as added; wall smooth and polished;
aperture terminal, elliptical, cribrate. aperture rectangular at the end of a
compressed extension of the last chamber,
Back reef areas and around patch reefs, without tooth.
5-20 m.
Systematics p. 270. Bay of Prony and outer reef, 10-30 m.
Systematics p. 268.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 109
Miliolinella labiosa
Test much broader than long, irregular
in outline, surface largely composed of
the two last-formed chambers; chambers
Miliola often somewhat irregular; periphery
rounded; surface smooth but dull;
Miliola? sublineata aperture irregular, a sinuous arch, with a
Test rounded in outline, elliptical in smoothly finished narrow bordering lip
end view, periphery broadly rounded; but without a true tooth.
chambers in the adult spirally arranged, Northern shelf, Chesterfield, 2-200 m.
about 3 chambers in a whorl, irregular Systematics p. 275.
in outline; sutures depressed; wall orna-
Inaequalina? sp. 2 mented with thin longitudinal costae;
aperture terminal, cribrate. This species
Test ovate, concavo-convex, with a very
possesses the cribrate aperture character-
deep concave side; oral and aboral ends
istic of the genus, but the pitted surface,
produced; coiling planispiral, evolute,
also considered as characteristic, is not
with two chambers per whorl, visible on
visible on all specimens (here visible on
both sides; chambers subtriangular in
the left specimen, not on the right one),
section, with an acute edge on the con-
making the position of this species
cave side; sutures depressed; wall smooth,
questionable.
but surface irregular; aperture rounded,
produced on neck, with a small lip and a Northern shelf, 600 m.
short tooth. This species is retained in Systematics p. 280.
Inaequalina despite the presence of a
distinct tooth, uncharacteristic for the
genus. As discussed by PARKER (2009), the
bilateral asymmetry is considered as the
prominent characteristic.
Southwestern lagoon, 35 m.
Systematics p. 268.
110 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Miliolinella webbiana
Test circular in outline, compressed, with
a prominent longitudinal costate orna-
ment; arrangement quinqueloculine in
early stages, latter almost planispiral;
chambers triangular in cross section;
large aperture at the end of the final
Miliolinella subrotunda chamber bordered by an everted lip, pro-
vided with a prominent flap-like tooth
Test milioline; 3-5 inflated chambers
that occupies about one half of the aper-
Miliolinella pilasensis visible, strongly overlapping previous
tural base.
ones, arranged almost planispirally in
Test subcircular in lateral view, somewhat adults; wall smooth, sometimes translu- Bay of Prony, 25 m.
compressed; periphery rounded; sutures cent; aperture somewhat triangular in Systematics p. 275.
depressed; last formed chamber inflated profile; low and broad flap-shaped tooth
at its initial end, tapered towards the in front of the aperture, that may lack in
aperture; wall imperforate, smooth and some specimens.
polished; aperture a low arch with an
apertural flap that leaves only a long From coastal lagoons to passes, 0-40 m.
narrow opening. Systematics p. 275.
Southwestern lagoon and bays, 2-45 m.
Systematics p. 275.
juveniles
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 111
Monalysidium
Miliolinella sp. 2 Monalysidium acicularis
Test subcircular in side view; in end view, Early stage planispiral, biconvex with
center of the test oval, but with two very inflated chambers, somewhat compressed,
high and thick carinae; wall smooth, latter uniserial, cylindrical, long and thin;
polished; aperture triangular, at the end wall with longitudinal costae separated
of the final chamber, under a crest-like by irregular rows of large pseudopores; Nodobaculariella
carina, provided with a relatively small aperture becoming increasingly complex Nodobaculariella convexiuscula
triangular flap-like tooth. with growth, with crenulations folding Test compressed, broadly elliptical or
Northern shelf, 200 m. irregularly inwards, forming a dendritic nearly circular, slightly biconvex; periph-
Systematics p. 275. pattern. eral edge sharp or carinate; chambers
South of the Grande Terre, 40 m. few in number, broad, embracing,
Systematics p. 281. sutures obscured; surface marked by
partial, irregular, longitudinal costae;
aperture placed at one side of the median
peripheral line, oval, bordered by a
thickened or everted lip.
Northern shelf, 200 m.
Systematics p. 266.
Miliolinella sp. 3
Test subsphaerical with 3 chambers visible
externally, increasing regularly in size as
added; surface smoothly finished; aperture
a narrow slit between the edge of the aper-
ture and the robust triangular flap-like
tooth, flush with the surface of the test.
Northern shelf, 500 m.
Systematics p. 275.
112 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Nubeculinita ramosa
Test attached, elongate, early stage
enrolled, later irregular chambers forming
two or more branches growing upright,
free of the attachment; wall imperforate,
milky white; aperture consists of one or
Nodophthalmidium more terminal openings.
Nodophthalmidium gracilis Passes of the southwestern lagoon and
Test elongate; initial stage globular Chesterfield, 3-90 m.
(proloculus followed by a planispirally Systematics p. 267.
enrolled second chamber); latter a few
uncoiled and rectilinear flasklike
chambers strongly tapering towards the Nubeculinella
distal end; wall imperforate, thick with Nubeculinella sp. 1
longitudinal costae; aperture ovate, Test attached; proloculus followed by a
terminal and somewhat produced. first chamber coiling around it, a half
Coastal bay, 10 m. coil in length; later chambers irregular
Systematics p. 267. in size and shape, uniserially arranged;
wall imperforate, smooth milky-white;
aperture terminal, semicircular, against
the substratum. This fragile test was not
found in the sediment. Nummoloculina
Southwestern lagoon, 30 m. Nummoloculina contraria
Systematics p. 267. Test ovate in outline, biconvex; periphery
broadly rounded, with two to five chambers
per whorl added in a single plane in
adult; lateral wall extensions from each
chamber overlap the preceding chambers;
wall thick, surface smooth and polished;
aperture semicircular to subtriangular, at
the end of the final chamber, with a small
flap.
Northern shelf, 200 m.
Systematics p. 279.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 113
Peneroplis
Parasorites Peneroplis pertusus
Parasorites orbitolitoides Highly variable morphology characterized
by a compressed biconvex planispiral coil,
Test discoidal, flat, and smooth; thickness often becoming fan-shaped or uniserial
hardly increased towards the periphery; in latter stages; test with numerous low
planispiral coiled early stage, later costae perpendicular to the sutures;
chambers annular, subdivided into one apertures multiple, terminal, a series of
layer of chamberlets by alternating radial irregular vermicular slits in young speci-
partitions; wall smooth and polished; mens that may become a row of square to
apertures small, round, in one row,
Nummulopyrgo rectangular openings in larger specimens.
sometimes in a slight depression, at the
Nummulopyrgo globulus periphery. Dispersed in the southwestern lagoon and
Test subspherical, chambers one-half coil Chesterfield, 0-40 m.
Southwestern lagoon and Chesterfield, Systematics p. 281.
in length with a rounded periphery; wall 0-60 m.
imperforate, smooth; aperture terminal, Systematics p. 282.
broad, nearly closed by a broad apertural
flap, leaving only a thin crescentic opening.
South of the Grande Terre, 35 m.
Systematics p. 268.
juvenile
juvenile
114 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Pseudohauerina
Pseudohauerina involuta Pseudohauerinella
Test subcircular in outline, lenticular, Pseudohauerinella dissidens
chambers in the early stage milioline, Test small robust, elongated, fusiform to
latter planispiral with usually more sub-polygonal in contour; both ends
than two chambers per whorl, distinctly truncated and not produced; chamber
involute; interior partially subdivided by arrangement quinqueloculine with grad-
numerous radial septula; wall ornamented ually increasing angle between successive
Planispirinella
with slightly curved ribs that correspond chambers; chambers slightly compressed,
Planispirinella exigua to the internal septula and numerous strongly plicated; sutures depressed;
Test discoidal, flattened, planispirally longitudinal striae; aperture terminal, in walls rough, ornamented with irregular
coiled with about three, hardly visible, the juvenile stage an opening with simple microstriae and irregularly distributed
chambers per whorl; whorls partially tooth, in the adult stage a convex cribrate pits; aperture at the end of the last
evolute, becoming more so in latter aperture. chamber, truncated, subcircular with an
chambers; an additional lamella covers Southwestern lagoon, 0-40 m. elongate bifid tooth; the thickened peri-
the umbilical areas with subsequent Systematics p. 280. stomal margin may produce two lateral
chamber additions, obscuring the previous infolds.
whorls; wall imperforate, smooth; aperture Northern shelf, 600 m.
a high ovate opening in the face of the Systematics p. 281.
final chamber.
Southwestern lagoon, 20 m.
Systematics p. 266.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 115
Pseudomassilina robusta
Test elliptical in lateral view, triangular
Pseudolachlanella slitella in end view; early stage milioline, latter
Test elongate, periphery subrounded; planispiral with two chambers per whorl,
chambers one-half coil in length, early Pseudomassilina macilenta slowly increasing in width; wall thick,
stage cryptoquinqueloculine latter nearly Test elliptical to circular in lateral view, very roughly finished, with an anasto-
planispiral; chambers broadly overlapping; flattened; early stage quinqueloculine, mosing ornamentation and numerous
wall imperforate, surface smooth; aperture latter planispiral with two to three large pits; aperture large and compressed,
a very narrow, curved, elongate slit with chambers per whorl, slowly increasing in without tooth but with a slightly everted
parallel sides, provided with a long slender width; wall ornamented with longitudinal, margin.
tooth with short, thickened termination. somewhat oblique costae; aperture large Southwestern lagoon, 15 m.
Bays, 5-20 m. and compressed, without tooth but with Systematics p. 276.
Systematics p. 275. an everted margin.
Southwestern lagoon, 5-30 m.
Systematics p. 276.
juvenile
116 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Pyrgo comata
Pseudotriloculina subgranulata
Test biloculine, involute, subglobular, the
Test cryptoquinqueloculine with usually
chambers very much inflated, in end
only three chambers visible in the final
Pseudotriloculina whorl; chambers inflated, wall finely
view subcircular; sutures distinct, incised;
Pseudotriloculina wall ornamented with numerous, fine,
granular, with surface undulations
cf. P. chrysostoma longitudinal costae; aperture oval, with a
resulting in a rough surface; aperture
tooth provided with winglike extensions.
Test cryptoquinqueloculine, ovate in lateral large with thickened peristomal rim and
and end views, somewhat compressed; large bifid tooth. Northern shelf, 600 m.
periphery broadly rounded; 3 chambers Systematics p. 276.
Dispersed in the southwestern lagoon,
visible, with the last two making up most 5-30 m.
of the test surface; aboral end rounded with Systematics p. 276.
chambers tapering towards the aperture;
sutures distinct but very slightly depressed;
wall smooth and shiny; aperture slightly
produced, a high arch that does not extend
down to the suture of the penultimate
chamber, narrowing along the stem of the
long tooth that has a T-shaped extremity.
The shape of the aperture, with the lips juvenile
juvenile
Pyrgo phlegeri
Test biloculine, involute, subcircular in
outline, biconvex lenticular in end view, Pyrgo sarsi
Pyrgo depressa somewhat truncated at the aboral end; Test nearly circular in end view, ellipsoid
Test nearly circular in front view, com- periphery acute, subcarinate; surface in end view, the ends slightly truncated
pressed, toward the periphery extending smooth; aperture distinctly produced, and the periphery angled, somewhat
out into a thin carina; median portion an elongated slit restricted by a highly produced; wall smooth; aperture broad,
rotund, in end view lenticular; wall protruding, elongated bifid tooth with with the tooth curved, concave in the
smooth; aperture broad, the tooth two short pointing extensions. middle, the ends extended and the
extending nearly the whole width of the Northern shelf, 600 m. aperture curving in a circle about them.
aperture, living only a slit-like opening. Systematics p. 277. Northern shelf, 600 m.
SNorthern shelf, 600 m. Systematics p. 277.
Systematics p. 276.
Pyrgo rasheedi
Test ovate in young to spherical in adult
specimens, biloculine; periphery rounded;
last chamber envelops half of penultimate
Pyrgo inornata chamber; wall smooth; aperture rounded
Test biloculine, involute in the adult, with a tooth projecting from the apertural
ovate in outline, slightly produced face; tooth changes from slightly bifid
towards the aperture, strongly biconvex through hoof-shaped bifid to reindeer
and subcircular in cross section; periphery horn-shape toward the full-grown adult
rounded; surface smooth; aperture oval, stage. Only young specimens were found
provided with a broad tooth with lateral in this study.
extensions and a wide base. Northern shelf, 600 m.
Systematics p. 276. Systematics p. 277.
juvenile
118 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Pyrgo sp. 3
Distinguishing features:
Test biloculine, involute in the adult,
Pyrgo vespertilio ovate, elongate, tapering gradually to the
Test subspherical, the apertural end apertural end, somewhat truncate;
broadly truncate; wall smooth except for chambers somewhat obliquely added;
a few small longitudinal ridges; aperture periphery rounded, sutures depressed; wall
Pyrgo subpisus elongate partially closed by a large flat imperforate, smooth; aperture elliptical
Test biloculine, involute in the adult, tooth with broadly rounded ends. This provided with a flattened T-shaped tooth
globular, circular in outline, periphery species, with longitudinal ridges is very with a thick base in the adult, a simple
acute; wall smooth or with fine striae; similar to the species illustrated by tooth in younger stages.
aperture elongate, broadly elliptical, with CUSHMAN (1921). Northern shelf, 600 m.
a raised rim and a large plate-like tooth Southern shelf, 70 m. Systematics p. 277.
with the ends broadly rounded; apertural Systematics p. 277.
opening narrow, sinuate.
Northern shelf, 600 m.
Systematics p. 277.
Pyrgo sp. 4
Pyrgo sp. 1 Test biloculine, involute in the adult,
Test biloculine, involute in the adult, subcircular in outline, slightly produced
subspherical; wall with 4-6 high irregular towards the apertural end, inflated and
costae; aperture low and wide at the end subcircular in cross section; periphery
of the last chamber, provided with a obtusely angled, sutures depressed; surface
broad plate-like tooth with prominent ornamented by a few longitudinal costae;
lateral extensions aperture subtriangular provided with
Northern shelf, 600 m. T-shaped tooth with a thin base.
Systematics p. 277. Northern shelf, 600 m.
Systematics p. 277.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 119
Quinqueloculina bassensis
Test with a cryptoquinqueloculine coiling;
Quinqueloculina Quinqueloculina auberiana chambers sudquadrangular in section with
Quinqueloculina agglutinans Test broadly oval; chambers sharply two carina and a convex peripheral margin
Test slightly elongate, quinqueloculine, angled to slightly sub-rounded, faint between the carina; wall matte, ornamented
with broadly rounded, somewhat truncated carina may be present; faces curved, or with weak anastomosing striae; a small
periphery; chambers distinct, rounded in S-shaped in end view; sutures distinct; amount of finely agglutinated matter may
transverse section; sutures visible wall smooth; aperture an arch of variable be attached to the test surface; aperture
although often obscured by the arenaceous height, with a simple tooth which may compressed, Lachlanella-type with a
material; wall with the surface coarsely protrude slightly above the periphery. distinctly bifid tooth and a peristomal lip.
arenaceous and roughly finished; aperture Northern shelf, 600 m. Islet in the southwestern lagoon, 5 m.
slightly produced, lachlanella type, with Systematics p. 270. Systematics p. 270.
a slight lip and a bifid tooth.
Southwestern lagoon, 5-25 m.
Systematics p. 270.
120 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Quinqueloculina
carinatastriata
Test elongate, oval in side view, subtrian-
gular in cross section; ornamented with
Quinqueloculina bosciana slightly oblique costae and a pronounced
Test elongate, truncated at the apertural peripheral carina; aperture terminal, cir-
end, rounded at the aboral end, periphery cular to slightly oval, produced on a short
Quinqueloculina bicornis rounded; sutures distinct; aperture termi- neck with a peristomal lip and a short
nal, circular, produced on a broad short tooth that thickens towards the tip.
Test quinqueloculine, broad, of moderate to
large size, ovate in lateral view; chambers neck with a short bifid tooth. As noticed Bays, outer estuaries, shrimp ponds, 0-20 m.
quadrangular in cross section, with three by HAIG, 1997, this species resembles Systematics p. 271.
strong longitudinal carinae; surface Quinqueloculina haigi in its chamber
densely ornamented by fine, but strong, arrangement (being cryptoquinqueloculine
longitudinal costae; aperture terminal, to quinqueloculine in New Caledonia),
elongated, narrow and keyhole-shaped and apertural detail, but lacks the distinct
bordered by a low flange with a long pseudopores that give Q. haigi a pitted
tooth, bifid at the tip. surface.
This species resembles Q. bicornis by its Coastal areas, bays, shrimp ponds, 0-10 m.
general shape, its surface ornamentation Systematics p. 271.
and its aperture. It differs by the three
strong carinae.
Northern shelf, 200 m.
Systematics p. 271.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 121
Quinqueloculina crenulata
Test elongate, slender, about three times Quinqueloculina debenayi
as long as broad; chambers not much
inflated; sutures distinct; wall ornamented Test fusiform in side view, laterally
Quinqueloculina corrugata compressed. Chambers one-half coil in
by very coarse, obliquely curved, short
Test quinqueloculine, elongated, periphery costae, extending inward from the length, slightly inflated, arranged in a
truncated; sutures depressed; surface peripheral angle, and sloping toward the quinqueloculine pattern; five chambers
marked by a series of transverse ridges base of the chamber; aperture produced visible from the exterior; sutures slightly
and alternating excavations; aperture on a prominent neck. depressed; chamber margins subrounded
elongated, slightly produced and recurved, in early stages, later tending to become
with a peristomal lip and a long tooth Southwestern lagoon, 35 m. carinate; surface with minute anasto-
that may be bifid at the tip. Differs from Systematics p. 270. mosing microridges; aperture terminal,
Q. parkeri in the truncated periphery subcircular, bordered by a thickened
and the produced aperture. collar-like peristomal rim and provided
Southwestern lagoon, 5-30 m. with a tooth with short bifid termination.
Systematics p. 270. Bays of the southwestern lagoon, 0-5 m.
Systematics p. 270.
122 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Quinqueloculina disparilis
Test short and broad, periphery broadly
rounded or somewhat truncate, the outer
side of the chamber with longitudinal
costae, the sides almost smooth, with fine, Quinqueloculina erinacea
indistinct striae; aperture not produced, Test small, elliptical in lateral view;
subcircular, provided with a bifid tooth. periphery rounded; prominent spinose
Northern shelf, 600 m. ornament, with 4-5 longitudinal rows of
Systematics p. 271. conical spines; aperture rounded, slightly
produced, with a small thick tooth some-
what bifurcated at the tip.
Southwestern lagoon, 25 m.
Systematics p. 272.
Quinqueloculina exsculpta
Test elongated, quinqueloculine; chambers
inflated and sutures deeply excavated;
wall smoothly finished; aperture produced
on a neck that may be long and curved in
adults, ovate, provided with a very short,
bifurcate tooth.
Lifou, Loyalty Island, 5 m; Bay of Prony,
15 m.
Systematics p. 272.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 123
Quinqueloculina jugosa
Test quinqueloculine, elliptical in lateral
view, elongated with a rounded periphery;
juvenile wall imperforate, ornamented by numer-
ous, low, longitudinal costae somewhat
anastomosing; aperture terminal, circular,
Quinqueloculina haigi produced on a short neck, bordered by a
peristomal rim and provided with a short
Test elongated, periphery rounded, ovate T-shaped tooth.
in cross section; chambers one half coil
in length, early stage cryptoquinquelocu- Coastal lagoons, shrimp ponds.
Systematics p. 272.
line later with 3-4 chambers visible from Quinqueloculina lizardi
the exterior; sutures very slightly Test elongate, compressed, elliptical in
depressed, hardly visible; wall finely lateral view, quinqueloculine, 3-5 cham-
pitted; aperture circular at the projected bers visible; test surface strongly pitted;
end of the last chamber, provided with a aperture terminal, slightly compressed
non pitted rim and a short simple tooth with a small tooth.
with thickened termination.
Bay of Prony, 20 m.
Northern shelf, 200 m. Systematics p. 272.
Systematics p. 272.
124 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Quinqueloculina seminula
Test elongate, 2 times as long as broad,
cryptoquinqueloculine to quinqueloculine,
ovate in lateral view, ovate in cross section,
with rounded periphery; oral end truncated,
aboral end inflated, slightly produced;
sutures slightly depressed; wall is smooth,
Quinqueloculina polished and glossy; aperture without a
cf. Q. sagamiensis neck, arched-shaped with a thickened
Test elongate, slightly compressed, nearly rim but no lip, provided with a small Quinqueloculina subcuneata
twice as long as broad, irregularly trian- tooth, often bifid. Test short, almost circular in side view,
gular in end view; chambers inflated, Coastal lagoons, marshes, estuaries, bays. subtriangular in end view with five
periphery rounded; sutures obscured by Systematics p. 273. chambers visible from the exterior;
the ornamentation, sinuous; surface chambers wedge-shaped, almost sharp
ornamented by several prominent longi- at the peripheral angles; wall polished,
tudinal costae, running from the base of ornamented with raised costae irregularly
the chamber to the base of the stout distributed on the test; aperture an oval
cylindrical neck; aperture subcircular, arch with a simple tooth thickened at the
with a single tooth, tip.
Southwestern lagoon, 20 m. Southwestern lagoon, 25 m.
Systematics p. 273. Systematics p. 273.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 127
Quinqueloculina tropicalis
Quinqueloculina subpolygona Test elongate, slightly compressed,
periphery rounded, but last chamber may
Test elongate, somewhat compressed, be somewhat truncated; chambers sub-
about 2 times as long as broad; chambers cylindrical with their maximum diameter
with a quadrangular periphery; test with at the aboral end, giving a rectangular
3-5 strong costae, two of them making outline to the test; wall dull, surface
carinate shoulders that reach the peris- ornamented with irregular ridges, this
tomal and aboral ends of the chamber; ornamentation described as granular by
aperture subquadrangular, Lachlanella Quinqueloculina tantabiddyensis
C USHMAN (1924); aperture terminal,
type, with a peristomal lip, provided with Test small, elongate, about 3 times as
circular, or triangular when the margin
a long narrow tooth with a small bifid long as broad, typically quinqueloculine
of the last chamber is truncated, with a
end. with five chambers visible in final whorl;
thickened rim and provided with a bifid
Southwestern lagoon, 5-35 m. rounded periphery; sutures depressed,
tooth.
Systematics p. 273. subparallel to test axis; oral end truncated,
aboral end rounded; wall smooth and Shallow bay in the southwestern lagoon,
glossy; aperture produced, Lachlanella- 0-5 m.
type, without peristomal lip, provided with Systematics p. 274.
an elongate tooth terminally thickened
or slightly bifurcate at the tip.
Bay of Prony, 15-25 m.
Systematics p. 274.
128 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Quinqueloculina
Quinqueloculina cf. Q. victoriensis
vandiemeniensis Test about twice as high as wide, with
Test small, elongate, subrectangular in rounded peripheral margins; chambers Quinqueloculina sp. 1
outline, sutures and sides roughly in quinqueloculine arrangement; surface
parallel; chambers in quinqueloculine ornamented with sharp longitudinal Test quinqueloculine, longer than broad,
arrangement; margin subacute; oral and costae; aperture somewhat everted, an elliptical in side view, somewhat com-
aboral ends truncated; surface smooth; elongated arch with a long bifid tooth. pressed; sutures fairly distinct, chambers
aperture terminal, not produced, provided The specimens from New Caledonia are polygonal in section; periphery keeled, the
with a thickened rim and a short bifid similar to those from Ningaloo Reef, keel dichotomously branching towards
tooth. Australia (PARKER, 2009). the aboral end; periphery concave
between the keels; wall smooth; aperture
Northern shelf, 200 m. Northern shelf, 200 m. subcircular at the end of a short neck,
Systematics p. 274. Systematics p. 274. and with a short, anvil-shaped tooth.
Bay of Prony, 15-25 m.
Systematics p. 274.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 129
Quinqueloculina sp. 3
Test elongate about two and a half longer
than broad, somewhat triangular in end
view, with five chambers visible in the
adult; chambers narrow with bicarinate
margin; the two carinae merge towards
the apertural end, giving a single high
carina; oral end truncated and slightly
produced, aboral end rounded and pro-
duced; sutures slightly depressed; wall Quinqueloculina sp. 5 Quinqueloculina sp. 7
finely striate, roughly finished; aperture Test ovate to subquadrate in side view, Test small, ovate in lateral view, quinque-
terminal with a tall oval shape, with ovate in end view; periphery broadly loculine, with rounded peripheral margin;
thickened peristomal lip; long thin tooth rounded; 5 chambers visible, with the last chamber subcylindrical with uniform
thickened at the tip. Quinqueloculina two making up most of the test surface; diameter throughout length; sutures
sp. 3 resembles Quinqueloculina sp. 22 aboral end rounded, sides nearly parallel; depressed; wall coarsely agglutinated;
of PARKER (2009), but differs from this sutures distinct, slightly depressed; wall aperture rounded with a lip and a short
species in the merging of the two carinae smooth and shiny; aperture at the trun- bifid tooth.
towards the aperture. cated end of the last-formed chamber, a Chesterfield, 20 m.
Southwestern lagoon, 25 m. high arch with a thickened rim and a Systematics p. 274.
Systematics p. 274. prominent, widened Y-shaped tooth.
Northern shelf, 200 m.
Systematics p. 274.
130 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Sigmoilina
Quinqueloculina sp. 9 Sigmoilina obesa
Test quinqueloculine, subcylindrical,
Test broadly oval in side view, with the
elongated in lateral view, ovate in cross
aboral end somewhat projecting, elliptical
section; chambers rounded in cross
in cross section; last chamber occupying
section; wall imperforate, ornamented by
about 3/4 of the visible surface; sutures
regularly parallel longitudinal costae
curved, but slightly depressed; two surfaces
prolonging on the long cylindrical neck;
inequilaterally convex, and periphery
aperture terminal, circular at the end of Schlumbergerina broadly rounded; surface smooth, may be
the neck, with a short simple tooth.
Schlumbergerina alveoliniformis polished; aperture a curved slit limited by
Southwestern lagoon, 25 m. a simple tooth.
Test with elongate tubular chambers one-
Systematics p. 274.
half coil in length, added in more than Northern shelf, 200 m.
five planes from the earliest stage, slightly Systematics p. 279.
inflated; sutures depressed; wall aggluti-
nated; aperture terminal, provided with a
trematophore with numerous small
rounded openings.
Widely distributed in the shallower areas
of the southwestern lagoon (5-25 m).
Systematics p. 280.
juvenile
aperture broken
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 131
Sorites
Sorites orbiculus
Sigmoilopsis elliptica Test discoidal biplane, thin; chamberlets
Sigmoilinita Test elongate, fusiform; chambers with adding in annular, concentric pattern,
rapidly changing planes in early stages, giving the sutures a characteristic scal-
Sigmoilinita costata later arranged in one plane; chambers loped appearance; wall smooth; apertures
Test fusiform in outline, slightly com- tubular, cylindrical; surface covered with a ovate or 8-shaped, bordered with a small
pressed; chambers of uniform diameter, layer of fine arenaceous matter; aperture rim, usually one on each side of the
arranged in a sigmoid pattern, later terminal, at the end of a cylindrical neck, chamberlets, positioned in a medial row
tending to become planispiral; periphery rounded with a thickened lip. on the peripheral margin.
rounded; sutures distinct, very slightly Lifou, Loyalty Islands, 5 m. Southwestern lagoon and Chesterfield,
depressed; surface costate; aperture semi- Systematics p. 279. 1-40 m.
circular, produced on a neck, with a Systematics p. 282.
peristomal rim and a short tooth.
Bay of Prony, 25 m.
Systematics p. 279.
132 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Spiroloculina attenuata
Test fusiform, elongate, biloculine, evolute,
Spiroloculina angulata strongly biconcave; chambers subtrape-
Test elongate, elliptical in lateral view, zoidal in transverse section; peripheral
both ends of the chambers projecting; wall slightly concave, separated from the Spiroloculina clara
chambers angular in cross section; sur- lateral walls by acute carinae; carinae of Test very much compressed, periphery
face ornamented by longitudinal costae previous chambers visible at sutures; sur- truncate and concave, both ends promi-
extending from the aboral end to the face ornamented by minute longitudinal nently projecting; sides of the chambers
aperture; aperture terminal, produced on anastomosing microstriae; aperture at the thickened and opaque, the central portion
a cylindrical neck with a thin peristomal end of a long cylindrical neck, rounded thin and translucent; sutures distinct, not
rim and small tooth, bifid at the tip. or subtriangular, with a slightly everted much depressed; wall smooth; aperture
Shallow areas near patch reefs, rare, 1-10 m. peristomal rim and two bifib teeth, a rounded, at the end of a neck, with a
Systematics p. 268. large one attached to the base of the tooth or sometimes two opposite teeth.
opening and an additional smaller one Southwestern lagoon, 35 m.
projecting down from the roof of the Systematics p. 269.
aperture.
Southwestern lagoon, near patch reefs,
2-5 m.
Systematics p. 269.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 133
Spirosigmoilina parri
Test subcircular, very compressed; early
chambers arranged in a sigmoiline coil,
Spiroloculina sp. 1 Spirophthalmidium leading to a raised central portion; later
Test elongate, ovate in lateral view; Spirophthalmidium scabrum chambers compressed, planispirally
chambers with angular shoulders arranged, each chamber one-half coil in
Test small, proloculus followed by undi- length; wall smooth; aperture at the end
between the peripheral and lateral walls; vided tubular enrolled second chamber,
sutural gaps at the base of each chamber; of the last-formed chamber, surrounded by
then by planispirally enrolled chambers of a raised lip formed by the slight outward
surface ornamented by parallel longitu- a half coil in length; chambers somewhat
dinal costae, extending from the aboral bending of the chamber’s wall, and with
overlapping previous whorls, widest at the a short simple tooth.
end to the aperture; aperture circular at base and tapering toward the aperture;
the end of a cylindrical neck with a slight wall imperforate, surface appearing warty Southwestern lagoon, 25 m.
lip and a small Y-shaped tooth. with numerous tiny projections; aperture Systematics p. 279.
Surprise Island, 10 m. rounded, at the end of a produced neck.
Systematics p. 270. South of the Grande Terre, 50 m.
Systematics p. 268.
136 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Triloculina elongotricarinata
n. sp.
Diagnosis. A slender, elongate Triloculina,
Triloculina barnardi triangular with rounded angles in cross
Test slightly longer than broad, triangular section, with a glossy surface and a high
in cross section, with sharp carinae; surface arch-shaped aperture with a long tooth
ornamented by low longitudinal costae Triloculina bicarinata thickened or bifid at the tip; triloculine
that may be weak or strong; wall smooth Test longer than broad, triloculine; sutures arrangement with three chambers always
and polished; aperture produced on a somewhat depressed, chambers distinct visible.
short neck, roughly triangular with a with a truncate periphery; whole surface Description. Test triloculine, with three
pronounced everted lip and an elongate ornamented by reticulations; aperture visible chambers in the adult, more than
tooth, slightly bifurcated at the tip. elongated, with a definite thin lip, slightly two times longer than broad; chambers
Bay in the southwestern lagoon, 10 m. everted, tooth elongate, narrow, extending triangular in cross section with isometric,
Systematics p. 277. above the outline of the aperture, bifid at slightly convex sides; acute margins; wall
the tip. smooth and polished; aperture high-arch
Southwestern lagoon, areas with strong shaped, provided with a long thin tooth
currents, < 20 m. somewhat thickened or bifid at the tip.
Systematics p. 278. This species differs from T. tricarinata by
its strait chambers and by its much more
elongated test.
Southwestern lagoon and Bay of Prony,
10-30 m. Systematics p. **.
Derivation of name. The name elon-
gotricarinata is given in reference to the
resemblance of this species to T. tricari-
nata, but with a more slender and elon-
gated form.
Material. Holotype - MNHN F62324,
paratypes - MNHN F62325, MNHN
F62326, MNHN F62327, MNHN F62328,
MNHNF62329; from 30 m water depth in
the Bay of Prony, south of New Caledonia.
Remarks. The species of Triloculina with
a triangular section and more or less sharp
angles are numerous and quite difficult
to be identified. This species, however,
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 137
with its high arch-shaped opening with Triloculina latiformis Triloculina rotunda
tall bifid tooth, as in T. tricarinata, and its Test triloculine, subcircular in side view; Test broadly rounded, somewhat longer
slender, elongate shape appears different with three visible chambers in the adult, than wide, with three visible chambers in
from other known species. Specimens with subtriangular in transverse section; the adult, the two last chambers making
the typical characteristics are abundant sutures deeply depressed; chambers arcuate up most of the visible test surface; cham-
enough to justify the creation of a new with tapered initial end and extended bers rotund; sutures very slightly
species. overhang, increasing rapidly in size; wall depressed; surface of the test smooth and
smooth and polished; aperture subcircular, shining; aperture rounded, with a slight-
with a protruding, slightly bifid tooth. ly thickened lip and a short protruding
Southwestern lagoon, 10 m. bifid tooth.
Systematics p. 278. Outer reef, 35 m.
Systematics p. 278.
Triloculina fichteliana
Test with three visible chambers in the
adult, ovate in lateral view, somewhat Triloculina serrulata
compressed with periphery broadly Test triloculine, with three visible chambers
rounded; chambers distinct; sutures in the adult, subovate in lateral view,
slightly depressed; wall ornamented by roughly triangular in end view; chambers
regular, well-spaced longitudinal costae; trapezoidal in transverse section; peripheral
aperture ovate, with a slightly everted Triloculina marshallana margins slightly convex, separated from
peristomal rim and a narrow tooth, Test triloculine, with three visible chambers the lateral walls by strongly protruding
thickened or bifid at the tip. in the adult, somewhat longer than wide; carinate shoulders; surface covered with
Northern shelf, 200 m. periphery broadly convex with angular anastomosing microstriae; aperture termi-
Systematics p. 278. shoulders; wall smooth and polished; nal, rounded, with a thick peristomal
aperture terminal, produced on a projecting rim, provided with a strong bifurcated
apertural neck, with an everted peristomal tooth projecting from a flattened base.
rim; aperture subtriangular with a thin Southwestern lagoon, 30 m.
bifid tooth. Systematics p. 278.
Southwestern lagoon, back reef areas, or
near patch reefs, 10-20 m.
Systematics p. 278.
138 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Triloculina wiesneri
Triloculina terquemiana Triloculina cf. T. tricarinata Test subcircular in lateral view; chambers
Test elongate with three chambers visible Test triloculine, with three visible chambers rounded to slightly angular; wall smooth
in the adult, ovate to subcircular in lateral in the adult, slightly longer than broad; on the sides of the chambers, periphery
view, subtriangular in cross section with chambers triangular in cross section with ornate by longitudinal costae; aperture
somewhat rounded angles; peripheral isometric straight sides; acute to carinate circular, without a neck, provided with a
chamber walls slightly convex; chambers margins; wall smooth, often polished; bifid tooth.
may be laterally prominent; surface of aperture low-arch-shaped provided with
Southwestern lagoon, 20 m.
the test ornamented by fine longitudinal large tooth, thickened or T-shaped at the
Systematics p. 278.
costae; aperture somewhat produced, tip. This species differs somewhat from
elongated, narrow, provided with a thin the typical T. tricarinata, which has a
tooth with long base, bifurcated at the tip. high arch-shaped opening with tall bifid
Surprise Island, 10 m. tooth.
Systematics p. 278. Widely distributed in the southwestern
lagoon, 5-60 m.
Systematics p. 278.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 139
old specimen
Triloculinella
Triloculinella chiastocytis
Test elongate with typical milioline
arrangement with three chambers visible
in the adult; characteristic inflated aboral
end, and oblique sutures; periphery Tubinella
broadly rounded; aperture small, partly Tubinella funalis
closed by a curved flap-shaped tooth. Test, elongate, cylindrical early stage Vertebralina striata
Southwestern lagoon, 20 m. bulbous, rounded at the base made up Test compressed, slightly trochospiral in
Systematics p. 278. of two closely appressed chambers, later the early stage; last chambers broad, a few
uncoiled, with indistinct tubular cham- of them unrolled; surface ornamentation
bers separated by faint sutures; wall varies from almost smooth to heavily
imperforate; surface with very fine longi- striated; aperture terminal, asymmetric
tudinal striae; aperture at the open end due to the shorted wall on the umbilical
of the last-formed chamber. side of the test, bordered by a thickened
Southwestern lagoon and Bay of Prony, lip.
2-25 m. Southwestern lagoon, dispersed, 10-30 m.
Systematics p. 280. Systematics p. 267.
140 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Wiesnerella
Wiesnerella auriculata
Test ovate, flattened, quinqueloculine,
periphery carinate; chambers overlapping
more on one side than the opposite, side
changing alternately; wall imperforate,
smooth; aperture a large circular opening Unknown Fischerinidae
at end of final chamber on the less over- Test composed of tubular chambers of
lapping side, bordered by a broad everted irregular shape, irregularly coiled, about
lip. three chambers per coil; sutures appear
Southwestern lagoon, 20-70 m; northern as irregularly spaced thickenings; surface
shelf, 200 m. with numerous unevenly distributed
Systematics p. 267. pseudopores and irregular longitudinal
juvenile
striae; aperture terminal, at the end of the
last chamber.
Buchnerina radiatomarginata
Northern shelf, 200 m.
Test ovate in outline, tapering towards the
aperture, with a single thickened carina
and a small projection at the basal end;
wall smooth with radiate ornament on
the lateral faces, around a clear unorna-
mented central patch; aperture produced,
surrounded by recessed grooves.
Coastal bay, 5-10 m.
Systematics p. 294.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 141
Buchnerina sp. 1
Test subrectangular in outline, laterally
Buchnerina walleriana flattened; central part of the test slightly
Test ovate in outline, laterally flattened raised and surrounded by a slight groove Cerebrina cf. C. clathrata
slightly tapering towards the aperture, and a ring; keel well-developed; aperture
with three bluntly rounded carinae that produced, surrounded by recessed Test nearly circular, tapering slightly
thicken towards the aperture and are grooves. towards the oral end, compressed;
separated by deep grooves; aperture pro- peripheral margin extended into a thin
Coastal bay, 10 m. flat wing or carina, with an additional
duced, surrounded by recessed grooves. Systematics p. 294. projecting keel or ridge on each side
Coastal bay, 5-10 m. bordering the chamber. The body of the
Systematics p. 294. test biconvex, ornamented externally with
a series of regular, parallel, longitudinal
costae; aperture produced on a distinct
neck, rounded to ovate.
There is some uncertainty regarding the
identification of this species, and speci-
mens illustrated in literature are much
variable, and often differ from original
description.
South of the Grande Terre, 10-50 m.
Systematics p. 288.
142 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Cerebrina neocastrensis
Test compressed, subcircular in outline,
except the produced apertural end;
Cerebrina lacunata periphery surrounded by a prominent
Test subcircular in outline, compressed, central keel with two raised lateral keels;
Cerebrina undulaticostata
with a reticulate ornament on the lateral central part of the test with coarse pits
surfaces; lateral carina extending from vertically aligned; aperture produced, Test free, tricarinate, compressed; central
the base to the aperture with two parallel rounded. area decorated with longitudinal costae
subordinate carinae on each side; basal of broken continuity, variable in length,
Bay to the south of the Grande Terre, 10 m.
end rounded; aperture ovate and produced width and position; marginal keel of
Systematics p. 289.
with a short entosolenian tube. variable width, thick, with greater part
transparent, tapering towards the base of
Bay of Prony, 20 m. the test; lateral keels form, sometimes
Systematics p. 289. together with longitudinal costae, two
ridges, which ornament the neck area on
both sides; a median ridge of variable
development may also form on the neck;
wall finely perforate, semitransparent;
aperture at the end of the neck, with wide
lips.
Northern shelf, 200 m.
Systematics p. 289.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 143
Cerebrina sp. 2
Test flask-shaped, compressed, inflated
centrally, tricarinate; central keel inter-
rupted at the rounded basal end; apertural
end produced with a neck ornamented Cushmanina spiralis
with small beads; median keel increasing Test unilocular, fusiform, circular in
Cushmanina gemma section, with a distinct neck; wall cal-
slightly in width at base of the neck but
does not reach the end of the neck; wall Test elongated, circular in section; basal end careous, hyaline, surface with thick
transparent, finely perforated; each test broadly rounded or truncated, apertural prominent paired spiral costae; the
face sculpted with irregular raised longi- end with a long cylindrical neck; wall depression between the two costae of each
tudinal costae that may coalesce or be hyaline, finely perforate, surface with pair is subdivided by bridges into oval
joined by irregular transverse patterns; numerous low and rounded paired longi- segments producing a chainlike appear-
aperture slightly compressed, oval. tudinal costae; the depression between ance under a stereo microscope, but this
the two costae of each pair is subdivided pattern is covered by the edge of the costae
Bay to the south of the Grande Terre, 10 m. by bridges into oval segments producing
Systematics p. 289. and not visible on SEM micrographs;
a chainlike appearance under a stereo aperture rounded at the end of the neck
microscope, but this pattern is covered by that is often broken.
the edge of the costae and not visible on Coastal bay, 10 m.
SEM micrographs; aperture rounded at Systematics p. 294.
the end of the neck, with a distinct lip.
Coastal bay, 10 m.
Systematics p. 294.
144 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Favulina hexagona
Test unilocular, subglobular, circular in
section; wall calcareous, hyaline, surface
covered by elevated ridges forming
large hexagonal reticulations; aperture
rounded on a slightly produced neck with
a thickened collar-like rim.
Coastal bay, 10 m.
Systematics p. 294.
Cushmanina Exsculptina
cf. C. tasmaniae Exsculptina discrepans
Test fusiform, circular in section with a Test with the main section roughly trian-
cylindrical neck; surface with a few thin gular in side view, the sides slightly
prominent paired spiral costae that convex and the greatest width near the
continue along the base of the neck; the base; basal periphery angled, ornamented
depression between the two costae of each by very short costae confined to the very
pair is subdivided by bridges into oval basal part; wall smooth; aperture at the
segments producing a chainlike appear- end of an elongate tapering neck with
ance; wall smooth; aperture rounded at weak longitudinal costae.
the end of the neck.
Isle of Pines, 5 m.
Bay of Prony, 20 m. Systematics p. 294.
Systematics p. 294.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 145
Favulina vadosa
Test globular, circular in section; initial
end broadly rounded, apertural end with Fissurina bispinata
a plate-like collar; wall ornamented with
large hexagonal reticulation, with Test rounded in outline, oval in section,
Favulina melo emphasis on the longitudinal part of the periphery with a weak keel; basal end
Test ovoid, circular in transverse section, pattern. Aperture small, rounded. with two short but prominent basal
ornamented by closely spaced longitudi- spines; wall calcareous, hyaline, finely
Coastal bay, 10 m. perforate, surface smooth; narrow bands
nal ridges connected by irregularly Systematics p. 295.
placed cross bars, straight or slightly near the margin of the test, opaque under
arched upwards; this reticulose mesh the dissecting microscope, whitish on SEM
thickens around the aperture forming a pictures; aperture terminal, symmetrical,
slight collar; aperture rounded, small. slit-like.
Bay of Prony, 20 m. Coastal bay, 10 m.
Systematics p. 294. Systematics p. 292.
Fissurina
Fissurina cf. F. antiqua
Test ovoid compressed; periphery with a
double keel fusing near the aperture;
central part of the test surrounded by an
annular ridge that form one or two Y-shaped
structures (depending on the size of the
individual) in the apertural region; longi-
tudinal costae raise in the mid central part
of the test; wall smooth, finely perforated;
aperture elongated, slightly produced.
Outer reef, 100 m.
Systematics p. 292.
146 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Fissurina circularis
Fissurina castanea Test with a circular lateral outline and
Test small, compressed, subtriangular to slightly compressed lenticular profile in
subquadrangular, elliptical in transversal axial cross section; the base of the test is
section; margins rounded; surface bordered by a narrow rounded carina
smooth; in the lower part of the test an that extends about halfway up the test;
elliptical fimbriate carina, surrounds the aperture produced, having a width about Fissurina
test; aperture terminal. one third of the test width. cf. F. eumarginata oblata
Southwestern lagoon, 25 m. Southwestern lagoon, 20 m. Test subcircular in contour; with a well-
Systematics p. 292. Systematics p. 292. developed keel; central body inflated,
highly translucent, finely perforated;
inflated area at the base of the produced
aperture; bands on each side of the inflated
central area, interrupted at the basal and
apertural ends of the test, opaque under
the dissecting microscope, whitish on
SEM pictures; aperture produced, a long
slit with tightly compressed lips.
Coastal Bay, 10 m.
Systematics p. 292.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 147
Fissurina laevigata
Test ovate in outline, lenticular in section,
periphery with a weak rounded keel; wall
finely perforate, surface smooth; aperture
Fissurina cf. F. globosocaudata terminal, ovate, bordered by two subtri-
Test globular, subcircular in outline, angular lips.
appearing somewhat elongated due to Northern shelf, 600 m.
the protruding apertural end; basal end Systematics p. 292. Fissurina lucida
ornamented by a short caudal projection Test pyriform, compressed, with rounded
irregularly truncated; surface of the periphery; basal end sometimes with a
chamber finely perforated; fissurine short projection; wall smooth and opaque
aperture slightly protruding. under the dissecting microscope, except
Coastal bay, 10 m. for the central area, which is clear and
Systematics p. 292. translucent.
Coastal bay, 5 m; shrimp ponds.
Systematics p. 292.
148 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Fissurina plebeia
Test pyriform, compressed, lateral edges Fissurina sp. 1
obtuse; base broad and round in outline;
edges with a median keel and two lateral Test ovoid, laterally compressed; periphery
keels separated from the median one by Fissurina sidebottomi with a central keel that widen near the
lateral gaping depressions on either side; aperture; two secondary keels form an
Test slightly compressed, slightly mar- annular ridge on each side, and prolong
wall smooth; aperture fissurine, not ginated, subcircular in lateral view with
produced. into a costa toward the aperture; a weak
the apertural end tuncated; paired sub- annular ridge is discernable around the
Northern shelf, 600 m. marginal costae extend around the lower center of each face of the test; wall
Systematics p. 292. half margin of the test; margin narrow, smooth; aperture elongated, slightly
rounded, uniform in width; wall finely produced.
perforated; aperture a long slit with a
distinct annular lip. Coastal bay, 10 m.
Systematics p. 293.
Bay of Prony, 20 m.
Systematics p. 293.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 149
Fissurina sp. 11
Test nearly rounded in contour, compressed;
periphery with a wide keel ornamented
with four nearly symmetrical spinose
projections developed laterally and at the
Fissurina sp. 9 aboral end of the test; two secondary keels
Test pyriform, compressed, with a rounded form an annular ridge on each side; sur-
margin; greatest width at about 1/3 of the face smooth; aperture a slit between two
test from the base; wall transparent, finely thickened lips, continuous with the keel.
perforated, with translucent lateral bands Northern shelf, 600 m.
of coarser perforations; surface smooth, Systematics p. 293.
shiny; apertural area prominent, hyaline,
broadly produced, somewhat depressed
along the axis; aperture ovate between
two thickened lips; entosolenian tube
attached to one side, about half the test in
length.
Northern shelf, 600 m.
Systematics p. 293.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 151
Homalohedra sp. 1
Test pyriform, tapering towards the aper-
Homalohedra gunteri tural end and truncated at the aboral
Test pyriform, more or less acutely pointed end, ornamented with a few rounded
at the apertural end; from a basal ring, costae running nearly the length of the
6-8 stout costae run up the sides of the test; aperture small and rounded, at the
test, then arch over and coalesce near the end of a small tubular neck. This species Hyalinonetrion elongata
apertural end; intercostal spaces are differs from Lagena acuticosta in the
concave; above the arch, one or two rows much-rounded costae and the lack of a Test very elongate, subcircular in section;
of alternating hexagonal pits form a ring thickened collar. It resembles, however the the central portion of the test, with parallel
around the neck; the test ends in a blunt species illustrated by CUSHMAN (1933a) as sides, gives a cylindrical appearance, the
oral extension with a rounded aperture. L. acuticosta in plate 8, fig. 9. basal end being almost symmetrical to
the long neck; wall calcareous hyaline,
Northern shelf, 600 m. Northern shelf, 600 m. finely perforated, surface smooth; aperture
Systematics p. 295. Systematics p. 295. terminal at the end of the neck.
Coastal bay, 10 m.
Systematics p. 289.
152 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Lagena paucistriata
Test flask-shaped, aboral end marked by
a short and thick spine, test terminates in
Lagena a short wide neck; wall finely perforate Lagena spicata
Lagena fenestrata and covered with costae originating from Test rounded to flask-shaped; wall orna-
the aboral end and that may continue mented with numerous longitudinal
Test elongate, flask-shaped, with a along the neck; aperture a round crenulate costae sometimes continuing up the
rounded aboral end and a tapering aper- opening at the end of the neck, differing apertural neck; neck cylindrical or
tural end with a long neck; wall covered from the round aperture with a phialine tapering; aperture terminal, rounded,
with a fenestrate network attached to the lip of Lagena spicata. bordered with a phialine lip.
surface of the test; the fenestrules are
elongated; aperture at the end of the Northern shelf, 600 m. Bay, south of the Grande Terre, 10 m.
neck. Systematics p. 289. Systematics p. 289.
Southwestern lagoon, 25 m.
Systematics p. 289.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 153
Lagenosolenia quadrangularis
Test elongate, compressed, quadrangular Lagenosolenia sp. 2
in transverse section; basal end rounded,
sometimes mucronate; apertural end Test globular, flask-shaped, broadly
tapering and terminating in a short rounded at the base, the opposite end
apertural neck; flat lateral sides of the test abruptly narrowed to a stout, short neck;
Lagenosolenia neoauriculata surface somewhat rough throughout;
edged by paired carinae; the external
Test flask-shaped, compressed, unicari- carinae of each face fuse in a keel that aperture terminal, rounded, with a slight
nate for anterior half, with laterobasal extends along the neck; the internal lip.
loops; lateral structure interrupted at carinae form an incomplete rim, then Bay of Prony, 30 m.
central basal area, giving a triangular also extending along the neck; surface Systematics p. 293.
outline to the test; relatively long carinate smooth; aperture ovate, produced on a
neck, with a prominent phialine lip; wall neck.
of the central part of the test with rather
coarse perforations; aperture terminal, Bay of Prony, 20 m.
rounded. Systematics p. 293.
Bay of Prony, 20 m.
Systematics p. 293.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 155
Lagnea
Lagnea lagenoides
Test flask-shaped, usually much com-
pressed, the body portion ovate surrounded Lagnea parviauriculata
by a simple peripheral keel of varying Test unilocular, flask-like, slightly com- Oolina
width in different specimens, and with pressed, periphery broadly carinate, keel Oolina ampulladistoma
numerous radiating tubulations, giving with radiating tubules, extending com- Test subcylindrical to subspherical; aboral
it in side view a somewhat scalloped pletely around the periphery and at the end with a caudal spine; numerous and
appearance; body of the test smooth, and base of the neck, bifurcating in the lower prominent nodules, usually more abun-
usually nearly transparent; aperture part of the test but converging at the basal dant on the basal half of the test that is
projecting with a distinct neck, slightly end; wall calcareous, hyaline, surface rougher; test tapering forward to a distinct
tapering, but in some specimens at least smooth; aperture round, terminal, at the groove at the base of a convex apertural
with a distinct entosolenian tube. end of a long neck, with a thickened rim. formation; aperture small at the center of
Isle of Pines, 5 m. Coastal bay, 5 m. this formation.
Systematics p. 295. Systematics p. 295. Bay of Prony, 30 m.
Systematics p. 295.
156 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Palliolatella sp. 1
Test free, circular in outline, compressed,
completely encircled by a wide carina;
wall smooth with relatively coarse perfo-
rations; aperture and neck enclosed by
the inflated carina.
Coastal bay, 10 m.
Systematics p. 293.
158 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Parafissurina
Parafissurina admiralis
Test ovate to subspherical, somewhat
longer than wide; keel uniform in width
and texture; apertural area produced with Parafissurina erecta
a relatively small lip on one side and a Test subquadrate, marginate but not
larger hood-like lip on the other side; carinate, much compressed; basal end
wall smooth, thick, transparent with quadrately rounded, apertural end slightly Parafissurina cf. P. kallima
heavily perforated peripheral areas, opaque produced; width about uniform from end to Test circular in contour, carinate, com-
under the dissecting microscope, whitish end; margin heavy, rounded, transparent, pressed but centrally inflated; keel
on SEM pictures; aperture a crescent-shape uniform laterally but wider and thicker prominent, transparent, of uniform width
slit with a long entosolenian tube. basally; wall transparent, thick, with central even at apertural end; wall transparent
Northern shelf, 600 m. area more hyaline; margin extending except an irregular white band (pores)
Systematics p. 296. vertically at the apertural end, furnishing near body margin; dorsal lip takes its
a shallow recess for the relatively short, origin on marginal keel that extends
ovate opening; entosolenian tube about forward, shorter ventral lip parallels
as long as the test, adhering to the dorsal dorsal lip, giving a slit-like aperture;
face. entosolenian tube adhering to the dorsal
South and southeastern lagoon, 50-100 m. face, about half the length of the test.
Systematics p. 296. This species matches well with the
description of P. kallima, but differs in its
shorter aperture and longer entosolenian
tube.
Northern shelf, 600 m.
Systematics p. 296.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 159
Procerolagena
Procerolagena cylindrocostata
Test elongate, circular in cross section
and with a cylindrical central portion;
Parafissurina cf. P. reniformis apertural end tapering gradually into a
long slender neck with a lip (often broken);
Test subcircular in contour, truncated at basal end tapers rather quickly; two sets
the apertural end, compressed, slightly of costae form the ornamentation, one set
marginated; wall translucent, slightly along the full length of the chamber and
roughened; apertural end squared off half way up the neck, the other set occupies
into a straight line; aperture located in a the lower three-quarter portion of the
depression; entosolenian tube adhering chamber. Both sets may slightly project
to the dorsal face, about half the length of beyond the basal end of the test.
the test.
Coastal bay, 10 m.
Southwestern lagoon, 20 m. Systematics p. 289.
Systematics p. 296.
160 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Seabrookia
Seabrookia pellucida Stictogongylus
Test elliptical in side view, plano convex Stictogongylus rugata
in end view; periphery slightly carinate; Small globular test with a small trochos-
Pygmaeoseistron chasteri early stage low trochospiral, chambers piral early stage followed by a thick
Test flask-shaped with broadly rounded rapidly increasing in size, strongly walled inflated final chamber that makes
base, gradually tapering to the apertural embracing; only two chambers are visible up 9/10 of the test; sutures indistinct giv-
end; finely pitted surface; aperture at the in the last whorl, and generally appear as ing the test an unilocular appearance;
end of the neck. if it was unilocular with a carina; wall wall irregularly ridged with numerous
translucent, very finely perforated; aper- fine pores; apical region smooth, without
Southwestern lagoon, 30 m. ture terminal, elongated, surrounded by pores, but with a few larger openings
Systematics p. 290. a rim. arranged in an incomplete spiral pattern.
Bay of Prony, 10-40 m. Coastal bay, 5 m.
Systematics p. 297. Systematics p. 326.
162 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Bombulina
Bombulina echinata
Test elongate, ovate, circular in section,
uniserial throughout, chambers strongly
Amphicoryna separans embracing; sutures horizontal, flush and
Test short, the initial portion composed of obscure; wall calcareous, hyaline, surface
three globular but not inflated chambers finely hispid; aperture terminal, rounded
uniserially arranged; initial end broadly at the end of a short neck, bordered by a
rounded with a small apical spine; one or phialine lip.
more spherical supplementary chambers Bay of Prony, 20-40 m.
are separated from the first chambers by a Systematics p. 296.
narrow tube; numerous strong costae run
the length of the test, terminating at each
chamber; aperture radiate located at the
end of a long regularly annulated neck.
Southern shelf, 70 m.
Systematics p. 287.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 163
Laevidentalina subemaciata
Test elongate, curved and tapering to the
rounded proloculus; chambers numerous, Laevidentalina sp. 1
low in the early portion, increasing in Test elongate, tapering gradually from
size as added; sutures flush and glassy in the acute initial end to the last-formed
earlier portion, becoming slightly chamber; initial end often with a stout
depressed and oblique in the final cham- spine; proloculus larger than the follow-
bers; wall smooth; aperture terminal and ing chambers; chambers not inflated
radiate. and sutures not depressed; wall smooth;
Laevidentalina mucronata
Northern shelf, 600 m. aperture radiate, terminal.
Test elongate, tapering from the initial Systematics p. 284.
end to the broadest last-formed chamber; Northern shelf, 600 m.
initial end acute, often with a short spine; Systematics p. 284.
chambers relatively few, not inflated;
sutures distinct, slightly oblique, but not
depressed; wall smooth; aperture terminal,
radiate, eccentric.
Northern shelf, 600 m.
Systematics p. 284.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 167
macrospherical
Nodosaria
Nodosaria nebulosa
Test composed of two to three globulose Polymorphinella
chambers with depressed sutures; surface
smooth and unornamented; aperture Polymorphinella pacifica
terminal, radiate at the end of a smooth Test elongate, ovate in outline, compressed; microspherical
neck. early stage biserial, with plane of biseriality
Bay of Prony, 20 m. parallel to the compression, later uniserial,
Systematics p. 285. but some chambers are somewhat Pyramidulina
cuneate, alternating from one side to the
other; sutures distinct, sigmoid; wall finely Pyramidulina catesbyi
perforate, surface smooth; aperture radiate Two to three chambers with an elongate
at the external angle. last chamber projecting in a distinct
Northern shelf, 600 m. neck; well-developed aboral spine; wall
Systematics p. 291. ornamented with several continuous
longitudinal costae; aperture radiate at
the end of the neck.
Coastal bays, 5-20 m.
Systematics p. 285.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 169
juvenile
Pyramidulina prava
Siphogenerina raphana
Test elongate, somewhat tapering; early
Test elongate, cylindrical, tapering slightly
chambers usually overlapping, and either
toward the initial end; initial end rounded,
2 or 3 in number increasing slightly in
oral end convex and broad; early chambers
size as added, followed by two or more
biserially arranged, later uniserial and
chambers obliquely placed, the obliquity Siphogenerina rectilinear for most of the test; sutures
becoming more pronounced as chambers Siphogenerina columellarensis distinct slightly depressed; wall ornament-
are added; sutures distinct, much con-
Test elongate, cylindrical; initial end, ed by several prominent longitudinal
stricted in the later portion of the test;
rounded or subacute, oral extremity convex costae regularly spaced and running the
wall ornamented by longitudinal costae,
and broad; early chambers biserially full length of the test; aperture circular,
in the earlier portion continuous over
arranged, then uniserial for most of the surrounded by a prominent lip.
adjacent chambers, later independent on
each chamber; aperture radiate, at the test; chambers rounded, not inflated; Bay of Prony, 10-30 m.
end of a cylindrical neck. sutures distinct, thick and slightly Systematics p. 302.
depressed; wall smooth, with short and
Southwestern lagoon, 25 m. faint costae across the sutures; aperture
Systematics p. 285. terminal, large, circular and surrounded
by a lip.
Coastal bay, 5-10 m, Bay of Prony, 20 m.
Systematics p. 302.
170 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Vaginulina
Vaginulina reophagina
Test elongate, uniserial, arcuate, slightly
compressed and curved; large proloculus
followed by subrhomboidal chambers,
rectangular in section; each chamber
bears four costae, two on either side and
has a rounded base and a produced neck;
sutures very oblique, slightly depressed; Abditodentrix rhomboidalis
wall finely perforated, surface smooth;
Test small, biserial throughout, triangular
aperture radial at the tapering end of the
in lateral view, rhomboidal in end view;
last chamber.
periphery obliquely truncate, sides slightly
South of the Grande Terre, 20 m. concave, especially in the later stage;
Systematics p. 286. chambers rapidly increasing in size as
added; sutures depressed, slightly curved;
large scattered pores except for a narrow
imperforate region adjacent to the aperture;
aperture an areal slit, commencing a slight
distance above the base of the rhomboid
apertural face and extending obliquely
upward, bordered by a distinct lip.
Bay of Prony, 10-30 m; coastal bay, 5-10 m.
Systematics p. 299.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 171
Bolivina spathulata
Bolivina glutinata Test compressed and highly flattened
Test elongate, compressed, broad; periph- with a lanceolate shape, slightly keeled
ery rounded; chambers provided with Bolivina subreticulata
and with a rounded initial portion;
irregular basal lobes distinct in the later sutures depressed; chambers numerous, Test small, few-chambered, compressed,
chambers only; sutures oblique, nearly not inflated, increasing gradually in size of rhomboidal shape; surface ornamented
straight, mostly obscured by the ornamen- as added, provided with a definite basal with an irregular network of raised lines;
tation; pores may be distinct in the last lobe near the median suture; wall aperture elliptical, extending from the
chambers; aperture elliptical, extending smoothly finished, but with a row of coarse base of the apertural face.
from the base of the last-formed chamber, perforations along the lower margin of the Northern shelf, 600 m.
with narrow rim and distinct toothplate. chamber and of the basal lobe; aperture a Systematics p. 298.
The original figures of Egger give little broad loop bordered by a rim and with a
guide for the identification of this species; distinct toothplate.
instead, illustrations by LOEBLICH & TAPPAN Bay of Prony, 20-40 m.
(1994) and HAYWARD et al. (1999) were Systematics p. 298.
used.
Bay of Prony, 10 m.
Systematics p. 298.
juvenile
172 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Bolivina sp. 1
Test elongate, slender, compressed tapering;
chambers biserially arranged, higher
than broad; periphery rounded; sutures
depressed, strongly oblique; wall distinctly
perforated, surface ornamented by longi-
Bolivina vadescens tudinal striations occupying about the
entire length of the test, except the last Bolivinellina translucens
Test compressed and highly flattened
chamber; aperture elliptical, extending Test small, elongate, slender, very slightly
with a lanceolate shape and a somewhat
from the base of the apertural face, with tapering, subcircular in cross section;
rounded periphery; less than twenty
narrow rim and toothplate. chambers biserially arranged throughout,
chambers; sutures very slightly depressed,
Coastal bay, 10 m. non inflated, gradually increasing in
appearing flush under light microscope;
Systematics p. 299. height as added; sutures very slightly
wall smoothly finished, distinctly perfo-
depressed, oblique and curved; wall
rated; aperture a broad loop bordered by
glossy, the anterior half of the chamber
a rim and with a distinct toothplate.
finely perforated, the posterior half
Bay of Prony, 5-40 m. coarsely perforated; aperture terminal,
Systematics p. 298. loop-shaped with narrow lip and an
internal toothplate.
Bay of Prony, 30-40 m.
Systematics p. 299.
juvenile
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 173
Cheilochanus
Cassidelina subcapitata Cheilochanus fimbriatus
Test elongate, compressed, increasing in Test small, highly compressed, sides flat-
Cassidelina davisi width toward the apertural end, periphery tened; gradually enlarging chambers
Test elongate, narrow, biserial and twisted; rounded; biserial arrangement through- biserially arranged; sutures distinct,
initial end acutely pointed with a promi- out, slightly twisted; chambers numerous, constricted at the lateral margin that has
nent spine; chambers high, increasing inflated, increasing rapidly in height; a fimbriate appearance; raised, slightly
rapidly in height as added; sutures sutures distinct, depressed, obliquely curved ridges extend from the medial line
depressed, oblique; wall smooth, finely curved; wall smooth, finely perforate; of the test to the periphery along sutures in
perforated; aperture a broad loop with aperture elongate, a curved slit, slightly the early stage; wall finely perforated; large
one margin higher than the other and a eccentric, one margin with a low rim, the ovate subterminal aperture bordered by a
toothplate. other bending inward. prominent flaring lip and turned slightly
Bay of Prony, 10-40 m. Bay of Prony, 30 m. toward one side of the test, without a
Systematics p. 301. Systematics p. 301. toothplate in the apertural opening.
Southwestern lagoon, 30 m.
Systematics p. 299.
174 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Loxostomina
Loxostomina barkeri
Test elongate, about 4 times as long as
juvenile broad, compressed, tapering toward
Hopkinsinella both ends; periphery rounded, lobulate;
Hopkinsinella glabra wall perforate, heavily ornamented by
Fursenkoina pauciloculata Test elongate, laterally compressed, irregularly continuous longitudinal
flattened ovate in section; biserial striae; sutures depressed, obscured by the
Test small, elongate, oval or subcylindrical, arrangement but with tendency for final ornamentation; aperture rather large,
slightly compressed, tapering slightly; chamber to become terminal; chambers terminal, ovate, with a toothplate.
ends rounded; initial end mucronate; increasing in relative height as added,
chambers few in number, long, erect, but Coastal bay, 5-10 m.
sutures oblique, depressed; wall smooth; Systematics p. 301.
little inflated, irregularly arranged; aperture subterminal, on a short neck,
sutures distinct, slightly depressed, surrounded by a recurved lip.
strongly oblique; wall smooth, finely
perforate; aperture narrowly elliptical, Bay of Prony, 20-40 m.
with the opening usually somewhat Systematics p. 301.
narrowed at the base of the chamber.
Bay of Prony, 10-40 m.
Systematics p. 306.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 175
Loxostomina sp. 1
Test elongate, moderately compressed,
periphery rounded, slightly lobulate; wall
coarsely but sparsely perforate, unorna-
mented except the last chamber; sutures
Loxostomina costulata flush to slightly depressed; aperture terminal Neocassidulina
Test elongate, compressed, ovate in section, at the end of the elongate and ornamented
last chamber, ovate, with a thick peripheral Neocassidulina abbreviata
often somewhat twisted; early stage biserial,
rim and a toothplate. Test elongate, biserial, compressed, oval
later tending to become uniserial; chambers
Northern shelf, 600 m. in end view, periphery rounded, initial end
increasing in height as added; surface
Systematics p. 302. rounded; sutures oblique, imperforate,
coarsely perforated with a few strong,
strongly limbate; wall coarsely perforated;
undulated costae; aperture terminal, oval
aperture a broad asymmetric loop, the
with a toothplate.
lower margin of the aperture folding
Bay of Prony, 30 m. inwards and joining the toothplate.
Systematics p. 302.
Northern shelf, 600 m.
Systematics p. 306.
176 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Patellinella inconspicua
Test short, subconical, earlier portion tro-
Neocassidulina sp. 2 chospiral, later biserial, slightly com-
pressed laterally, earliest whorl with three
Test elongate, biserial, initial end round- chambers, later with two chambers, all
ed, but with a stout spine; apertural end visible on the convex spiral side, only the
broadly rounded; chambers increasing final pair visible on the flat umbilical
rapidly in width in the early stage, later Pseudopolymorphina
side; sutures flush or very slightly
slowly; only the two last chambers inflat- depressed; apertural end truncate, some- Pseudopolymorphina sp. 1
ed; periphery rounded; sutures flush with what concave; wall calcareous, finely per- Test compressed, oval but somewhat
the test in early portion, increasingly forate; aperture a broad umbilical arch. inequilateral in outline; two faces almost
depressed later; wall coarsely perforated, equally convex; both ends obtuse; cham-
surface ornamented by one or two rows of Bay of Prony, 30 m.
Systematics p. 306. bers arranged with regularity in two
tubercles paralleling the sutures and fus- alternating series, the last pair occupying
ing in the early portion of the test; aper- two thirds of the visible shell; sutures
ture loop-shaped, asymmetrical. depressed; wall finely perforate, surface
Southwestern lagoon, 30 m. ornamented with solid costae that may be
Systematics p. 306. continuous from one chamber to the
other; aperture terminal, radiate, with the
central part cribrate.
South of the Grande Terre, 50 m.
Systematics p. 291.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 177
Sagrina zanzibarica
Test elongate, biserial; early portion
somewhat compressed, periphery in end
view broadly rounded; test tapering
gradually throughout; chambers distinct,
somewhat inflated, low and broad in the Sagrinella durrandii Sigmavirgulina
early portion, later increasing in relative Test elongate, lanceolate, compressed, Sigmavirgulina basistriata
height; sutures distinct, depressed in the peripheral margin acute and serrated;
later portion, straight, slightly oblique; Test elongate, initial end only slightly
chambers slightly inflated, inferior margin twisted; test thinning out toward the
wall ornamented with numerous small, acute and projecting; sutures deeply sunk
short, blunt spinose projections, particu- periphery, with carinate margins; surface
and smooth, surface of chambers orna- ornamented with sparse longitudinal
larly on the lower half of the chamber; mented with broken irregular costae;
aperture elliptical with a distinct raised costae at the base and across the sutures;
aperture terminal, a long fusiform slit. sutures slightly curved, slightly depressed
lip.
Bay of Prony, 5-30 m. in the last chambers; wall coarsely perfo-
South of the Grande Terre, 30 m. Systematics p. 302. rated; aperture loop-shaped with a raised
Systematics p. 302. thickened lip and a toothplate that proj-
ects into the top of the opening. It differs
adult from S. tortuosa by its less twisted shape,
and the presence of longitudinal costae.
South of the Grande Terre, 30 m.
Systematics p. 306.
juvenile
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 179
Sigmavirgulina sp. 1
Species differing from S. turtuosa by its
more elongated, and less twisted test; test
subrectangular in side view, with chambers
increasing slowly in width as added.
Northern shelf, 600 m.
Systematics p. 306.
Bulimina marginata
Test elongate ovate, chambers triserially
arranged throughout, sharply angled
Virgulopsis about half the distance from the basal
suture, with spines at the angle somewhat
Virgulopsis spinea extended backwards; sutures depressed;
Small stout test with a spine at the initial wall calcareous, finely perforate; surface
end and a prominent spinose ornament; smooth other than the spinose angle;
periphery rounded; chambers inflated, aperture a loop extending up the face from
sutures depressed, oblique; aperture a the base of the last chamber, provided
large slit that extends up the apertural with an internal folded toothplate.
face. Bay of Prony, 20-40 m.
Coastal bay, 5-10 m. Systematics p. 303.
Systematics p. 301.
180 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Mimosina echinata
Test triserial in the early stage, later
Fijiella tending to become biserial, chambers
subglobular and inflated, sutures
Fijiella simplex
Hopkinsina obscured by ornamentation; the whole
Test pyramidal, triserial and triangular in surface of the test, except the terminal
section throughout; sutures gently arched, Hopkinsina sp. 1
chambers covered by a dense growth of
limbate; peripheral margins carinate, Test elongate, early chambers triserially fine spines, with some bigger spines at the
spinose, straight or somewhat lobulate; arranged and closely appressed, later angle of the chambers; surface coarsely
apertural end truncated, slightly convex, loosely triserial and then biserial; chambers perforated; aperture double with a low
bordered by a thickened imperforate rim with the base sharply undercut and the basal arch and a subterminal ovate
and ornamented with numerous small upper surface broadly domed; margin opening, each bordered by a narrow
upright spines; wall calcareous, coarsely carinate, ornamented with a row of short imperforate lip.
perforate mostly along the sutures; surface tooth-like spines, some of them prolonging
into short costae on the upper surface of Isle of Pines, 5 m.
smooth other than the limbate sutures, Systematics p. 305.
spinose margins, and sometimes produced the chamber; sutures depressed; wall
pores; aperture a narrow slit partially finely perforate, surface smooth; aperture
covered with a curved, denticulate lip and terminal, ovate, slightly produced and
provided with a large perforated tooth- with a distinct lip and a toothplate.
plate that forms rounded supplementary South of the Grande Terre, 30 m.
openings on the central part of the Systematics p. 301.
apertural face.
Southwestern lagoon, 40 m.
Systematics p. 304.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 181
Neouvigerina interrupta
Test much elongated, subspiral; composed
of a number of inflated or subglobose
chambers of gradually increasing size, Neouvigerina proboscidea
Mimosina sp. 1 (juvenile?) arranged around a long axis; earlier Test small, elongate; initial stage triserial
chambers combined so as to form a more with closely arranged chambers, followed
Test very small, triserial, chambers sub- by biserial and uniserial stages; chambers
globular and inflated, sutures depressed; or less compact spire; later chambers
disposed in an irregular, interrupted, inflated; sutures depressed; test covered
wall coarsely perforate and covered with with fine spines; aperture terminal on a
short spines, except on the upper part of alternating series, terminating in a tubular
neck; surface hispid or aculeate; aperture long neck, with lip and toothplate.
the chambers that is smooth; aperture
wide, occupying most of the apertural at the end of the neck with a toothplate. Bay of Prony, 20-30 m.
face, with a folded periphery, without a Northern shelf, 600 m. Systematics p. 303.
lip. The aperture resembles that of young Systematics p. 303.
stages of Mimosina, before its separation
into a double aperture.
Outer reef, 100 m.
Systematics p. 305.
182 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Trifarina
Reussella neapolitina Trifarina angulosa
Test pyramidal, triserial throughout, sharply
Test elongate, sharply triangular in section,
triangular in cross section, regularly Reussella pulchra initial stage triserial, later with tendency
increasing in size; chambers wider than
Test averaging about 1-1/2 times as long to an irregularly uniserial arrangement;
high; sutures distinct, curved, limbate
as broad, triangular in transverse section, apertural end pointed; angles carinate,
and raised over the surface, marked by
the sides carinate and with a spine at the sutures curved, oblique, and slightly
short spines and tubercles; marginal
base of each chamber and a spine at the depressed; wall with fine perforations at
spines prominent and pointing backward;
initial end; chambers distinct, not inflated; the end of raised pustules; surface with a
apical spine often present; wall distinctly
sutures very distinct, limbate, raised well few longitudinal costae around the keel;
perforate and covered by short spines and
above the surface and often finely spinose aperture terminal, ovate, produced on
pustules, except on the upper face of
to give a sculptured appearance to the a neck and bordered by a narrow lip,
chambers; aperture an elliptical opening
test; wall distinctly and rather coarsely provided with an internal toothplate.
along the basal suture, bordered by a low
lip which descends into the lumen and perforate; aperture a narrow opening at Bay of Prony, 20-30 m.
fuses with the toothplate. the inner margin of the last-formed Systematics p. 304.
chamber, often with a distinct lip.
South of the Grande Terre, 70 m.
Systematics p. 304. Northern shelf, 600 m.
Systematics p. 304.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 183
Trimosina
Trimosina milletti
Test elongate, triserially arranged, triangu-
lar in section, tapering towards the basal
end; margins acute, the marginal angle Uvigerina
of the inflated chambers are developed Uvigerina carapitana
Trifarina pacifica into lobes terminating in a spine; test Test stout, compact, rather bulbous,
Test with a well-developed initial triserial often somewhat contorted; aperture a slit triserial with about 3-4 whorls visible;
stage and a few later chambers added in at the base of the last-formed chamber, periphery smoothly rounded, lobulate;
irregular uniseries; peripheral margins connected or not to a more or less developed chambers somewhat angular in the early
acutely rounded, never carinate; wall subterminal orifice. stage, later inflated; sutures depressed;
distinctly dented along the sutures giving South of the Grande Terre, 30 m. wall thick finely perforate, generally
the test an irregular outline in side view; Systematics p. 305. smooth though some specimens show
aperture produced, with a toothplate and faint longitudinal striations; aperture
surrounded by a thickened lip, but lack- terminal, at the end of a short neck; neck
ing a distinct neck. in a depression near the indented margin
Northern shelf, 600 m. of the last chamber, with a distinct lip.
Systematics p. 304. Northern shelf, 600 m.
Systematics p. 304.
juvenile
juvenile
184 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 185
Ammonia sp. 1
Test low trochospiral, biconvex; periphery
lobulate, narrowly rounded; spiral side
evolute with 3-4 whorls of nearly constant Anomalinoides globulosus
height but with chambers increasing in
breadth as added, sutures depressed, Test robust, in a low trochospiral coil,
Angulodiscorbis tobagoensis planoconvex with highly convex, involute
oblique; umbilical side involute, 5-7
chambers of the final whorl visible around Test small, trochospiral, five-sided cone- umbilical side and evolute spiral side;
the umbilicus, sutures straight and radial shaped, concavoconvex with a broadly periphery broadly rounded; 5-10 inflated
with prominent sutural notches; umbilical angular, lobulate periphery; spiral side chambers in the last whorl; sutures
area filled with granular material; large convex with five chambers per whorl; depressed, curved on spiral side, gently
umbilical folium covering the umbilicus; chambers broadly angular in the center curved to nearly straight and radial on the
wall thin, hyaline, finely perforate; aperture so as to form five sides on the spiral face; umbilical side; wall coarsely perforated
an interiomarginal, extraumbilical arch sutures flush, obscured by the ornamen- on both sides; aperture crescentic, against
that continues under the umbilical folium. tation; umbilical face involute, with a the periphery of the preceding whorl, with
strongly depressed umbilical area; wall a distinct lip, extending onto the spiral
Bay of Prony, 20-40 m. heavily ornamented, with a dimpled side where it continues along the spiral
Systematics p. 323. texture on the spiral side and radial rows of suture beneath the margin of the last few
granules on the umbilical side; granules chambers of the final whorl.
increasing in size toward the umbilicus, Bay of Prony, 20 m.
and passing progressively to spines; Systematics p. 321.
aperture interiomarginal at the base of
the last-formed chamber.
Crawling on algae, Chesterfield, 20 m.
Systematics p. 311.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 187
Anomalinulla
Ashbrookia
Anomalinulla glabrata
Ashbrookia ornata
Test biconvex with very low trochospiral Baggina philippinensis
Test planoconvex, low trochospiral sub-
coiling, appearing almost planispiral;
circular in outline; evolute spiral side Test globular, low trochospiral with few
spiral side flattened, umbilical side
slightly convex, involute umbilical side inflated and rapidly enlarging chambers
slightly convex with a distinctly excavated
slightly concave; periphery lobulate, per whorl; final chamber very inflated,
umbilicus; sutures flush, becoming
peripheral margin acute; coil of approxi- occupying nearly one half of the umbilical
slightly depressed between the last few
mately two whorls, about three, strongly side; umbilicus open; sutures depressed,
chambers; wall smoothly finished and
overlapping, crescentic chambers per radial, curved; periphery broadly rounded;
evenly perforate on the spiral side;
whorl; sutures strongly curved, final surface smooth, glassy; wall perforate but
aperture an equatorial slit that starts a
chamber occupying about half the with an imperforate lunate area on the
short way along the chamber on the
periphery and umbilical side, outer part umbilical side just above the aperture;
spiral side and extends to the spiral
of the chambers divided by partial radial aperture a broad umbilical opening at
suture on the umbilical side of the test;
septula; umbilicus open, partially covered the base of the apertural face, just over
aperture bordered by a lip that continues
with an umbilical flap from the final the sculptured ridges.
into a small folium; apertures of the
chamber; wall finely perforate, somewhat Northern shelf, 600 m.
previous chambers remaining open for
granular on the spiral side; aperture Systematics p. 306.
most of the last whorl.
umbilical in position, under the umbilical
Bay of Prony, 20-40 m. flap.
Systematics p. 321.
Southwestern lagoon, 30 m.
Systematics p. 306.
188 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Bronnimannia palmerae
Buliminella
Test auriculate in outline, very low tro-
chospiral, whorls enlarging rapidly; test
Buliminella elegantissima
somewhat flared, bievolute and biconcave Test elongate, fusiform, a high trochospiral
with broadly truncate periphery marked coil of only 2-3 whorls of numerous elon-
by a distinct keel at the spiral side; sutures gate chambers, the last whorl about 80%
curved and depressed on both sides; on the of the test; sutures slightly curved, almost
umbilical side, chambers have a flap or parallel to the axis of the test, slightly Calcarina
folium marked by a distinct notch at the depressed; wall finely perforate, surface
smooth; aperture a loop in the depressed Calcarina exuberans new name
posterior margin, both folium and notch
remaining visible on most chambers of face of the final chamber with a high rim Species differing from C. hispida by
the final whorl; wall finely perforate on and a simple internal toothplate. exuberant outgrowths at the extremity of
the umbilical side, spiral side more Coastal lagoons, estuaries, shrimp ponds. the strong hispid, bifurcating spines,
coarsely perforate; aperture a low, interi- Systematics p. 303. which give the test a plumose appearance;
omarginal slit beneath the umbilical test more delicate than in C. hispida,
flap. particularly in the last-formed chambers.
Bay of Prony, 10 m. This species was named Calcarina hispida
Systematics p. 311. var. pulchella by CHAPMAN (1900). Since
C. pulchella was preoccupied by
Calcarina pulchella d’Orbigny 1839a
(Asterorotalia pulchella), a new name
is proposed to resolve ambiguity and
maintain nomenclatorial stability.
Southwestern lagoon, and mostly on the
southern shelf, 40-80 m.
Systematics p. 323.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 189
juvenile
Cancris sagrum
juvenile
Test elliptical in outline, umbilical side
convex, spiral side almost plane; margin
strongly keeled; test composed of 1-2 whorls
Calcarina mayori of rapidly enlarging chambers, about 6 in
Test low trochospiral, medium-sized; the last coil; last-formed chamber making
spiral side evolute and umbilical side Cancris up nearly half the area of the umbilical
involute, but only the last whorl visible, side; sutures slightly depressed and
the center of test being obscured by coarse Cancris auriculus arched on the spiral side, deeply
raised tubercles; sutures depressed between Test auriculate in outline with a strongly depressed and almost radial on the
the last chambers on the umbilical side; convex umbilical side; peripheral margin umbilical side; wall distinctly perforated,
peripheral margin rounded; 3-10 radial acute with a small carina; chambers except an imperforate elongated area on
peripheral spines in the plane of coiling, arranged in a flared trochospiral coil and the apertural face; aperture a slit at the
slightly hispid, straight, of constant increasing rapidly in size; sutures arched, base of the last-formed chamber, on the
width; test covered by openings of the flush or slightly depressed and strongly umbilical side, with a small folium.
intraseptal canal system; wall of the last recurved backward at the periphery on Bay of Prony, 20-40 m.
chambers, covered with short, protruding the spiral side, more depressed and nearly Systematics p. 307.
spikes, and perforate on the umbilical radial on the umbilical side; wall thin
side; apertural face flat, imperforate; and smooth, finely perforate, except a
apertures multiple, on the apertural face, semicircular region on the septal face;
irregularly rounded with thick peristomal aperture a slit on the umbilical side, at
rims. the base of the last-formed chamber, with
Attached to algal thalli, Chesterfield, a broad apertural flap extending over the
0-40 m. umbilicus.
Systematics p. 323. Southwestern lagoon, 30 m.
Systematics p. 307.
juvenile
190 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Carpenteria
Carpenteria monticularis
Cibicides pseudolobatulus
Test attached, large, convex and montic- Cibicides
ular; circumference is deeply lobed, Test trochospirally coiled, spiral side flat,
Cibicides mabahethi sutures limbate, oblique and curved
somewhat irregular; early chambers
spirally arranged, broad and round at the Test low trochospiral, planoconvex to backward at the periphery; umbilical side
outer margin, narrow at the umbilical slightly biconvex; evolute spiral side, convex with depressed sutures radial
end; later chambers piled up irregularly prominently convex, involute umbilical around the umbilicus and curved back-
about a central axis, somewhat inflated; side with a broad imperforate umbilical ward at the periphery; peripheral margin
wall coarsely perforated; aperture a knob; peripheral outline becoming slightly acute, carinate; peripheral outline faintly
rounded opening at the summit of an lobulate in adult; peripheral margin lobulate; wall coarsely perforate on the
erect tubular extension of the end of the acute with a carina; sutures curved on spiral side, less densely and coarsely so
central axis. both sides, limbate on the spiral side, on the umbilical side; aperture an interi-
slightly depressed on the umbilical side; omarginal, equatorial arch, bordered by
Northern shelf, 600 m. test coarsely perforated on the spiral side,
Systematics p. 317. a lip and extending onto the spiral side
perforations scattered, mostly along the where it remains open in the last few
sutures, on the umbilical side; aperture chambers.
extraumbilical equatorial, provided with
Northern shelf, 600 m.
a thick rim and extending into a supple-
Systematics p. 315.
mentary spiral aperture remaining open
in the last few chambers.
Northern shelf, 600 m.
Systematics p. 315.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 191
Conicospirillinoides sp. 1
Conicospirillinoides
This form resembles C. semidecoratus,
The test of Conicospirillinoides is planispi-
but differs in its more rounded margin,
rally enrolled, but the wall, extending
thicker test, and in much more overlapping
on one face into a high spiraling band
whorls on the flattened side that therefore
surrounding a deep umbilical depression,
becomes more convex.
results in a dissymmetrical test. The
result is the impression of a trochospiral Northern shelf, 200 m.
coiling, justifying the placement of this Systematics p. 283.
genus in the “Trochospiral hyaline Conicospirillinoides
foraminifera (or appearing so)” section. semidecoratus
Test planoconvex consisting in a globular
Conicospirillinoides proloculus and undivided planispirally
denticulatus enrolled tubular chamber; wall extending
Test planoconvex with a proloculus and into a high spiraling flange that partially
an undivided, compressed, planispirally overlaps the umbilical region and slopes
enrolled tubular second chamber; flat- upward considerably beyond the chamber
tened side evolute with all the whorls lumen, surface of the flange bearing
visible, flat or slightly convex; raised side numerous radial indentations; flattened Conicospirillinoides sp. 2
with whorls hardly distinguishable, and a side covered with rounded bosses, except This form differs from C. semidecoratus by
deeply depressed umbilical area; peripheral for the last whorl; periphery with a blunt its more regular pattern of ornamentation
margin acutely rounded; sutures slightly keel; aperture at the end of the tubular on the flattened side, and by the spines
raised on the flat side, incised but indistinct chamber at the periphery. produced on the peripheral margin.
on the raised side; on the flat side, wall
Northern shelf, 200 m. Northern shelf, 200 m.
with a row of coarse perforations regularly
Systematics p. 282. Systematics p. 283.
disposed parallel to the spiral suture; on
the raised side, wall extending into a high
spiraling band surrounding a deep
umbilical depression, and furnished with
buttress-like teeth set at regular intervals
along its inner margin; aperture simple,
at the end of the tubular chamber, at the
periphery.
South of the Grande Terre, 30 m.
Systematics p. 282.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 193
Conorbella pulvinata
Test broadly ovate or subglobular,
planoconvex; composed of a few more or
less inflated chambers, about three in the old Discanomalina
specimen
final coil; spiral side strongly rugose due Discanomalina coronata
to coarse pores surrounded by rims of Test robust, very low trochospiral, nearly
various height; rows of pores follow the equally biconvex, in face view nearly as
edges of the chambers, but additional broad as the diameter; about eight
pores may open over the chamber walls; chambers in the final coil; umbilical
umbilical side deeply excavated at the region concave on both sides; peripheral
umbilicus and ornamented with radiating border nearly flattened in the later
granulose lines; aperture a rounded chambers, which increase rapidly in
opening at the suture on the umbilical width; inner border of the chambers often
side. of clear shell material; wall coarsely
Bay of Prony, 10-20 m. perforate; aperture a narrow curved slit
Systematics p. 311. at the base of the apertural face, bordered
with a narrow lip; supplementary openings
beneath umbilical flaps.
Northern shelf, 600 m.
Systematics p. 322.
194 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Discorbinella
Discorbinella bertheloti Elongobula parallela
Test compressed, plano-convex, semicir- Test elongate, the sides usually nearly
cular in outline; coiling low trochospiral, Discorinopsis parallel for most of their length, both
nearly involute on both sides; peripheral Discorinopsis aguayoi ends broadly rounded, nearly circular in
margins acute with a blunt carina; flat Test large, low trochospiral; chambers transverse section, periphery slightly lob-
side with shallow umbilicus and folia on increasing slowly in height but rapidly in ulate; chambers distinct, in 3 or more
the last 3-4 chambers; chambers 6-8 in breadth resulting in an auriculate test; whorls; sutures slightly limbate, flush
the final whorl, slightly inflated on convex spiral side convex, umbilical side flat- with the surface; wall smooth, very finely
side, increasing in size gradually and then tened to concave; umbilical face partially perforate, with a weak anastomosing
rapidly for the last 3-4; sutures depressed, covered with a spongy mass of shell costate ornament over the lower parts of
curved and slightly thickened on both sides; material; sutures strongly curved on the the test; apertural face terminal, striated;
wall smooth, coarsely perforate on the spiral side, obscured on the umbilical aperture central, partially hidden by a
convex side, perforate only at the periphery side; wall coarsely perforate; aperture a broad flap.
of the flat side; aperture equatorial to series of openings through the shell Bay of Prony, 20-30 m.
interiomarginal, broadly arch-shaped material that covers the umbilical area. Systematics p. 313.
with a thickened lip, with supplementary
posterior foliar openings. Coastal lagoons, marshes.
Systematics p. 262.
Northern shelf, 600 m.
Systematics p. 314.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 195
Eponides
Eponides repandus
Geminospira
Test low trochospiral, biconvex, more
convex on the umbilical side; umbilicus Geminospira bradyi
closed; periphery angular to carinate; Test elongate, flattened, early chambers
2-3 whorls with about 5-7 chambers per in a trochospiral coil, later uncoiled,
whorl; sutures curved and limbate on the
Floresina arcuate, and asymmetrical with chambers
spiral side, continuing into the peripheral Floresina latissima highest on the spiral side; a series of
keel, nearly radial on the umbilical side: Test elongate, irregularly ovate in outline, secondary chamberlets on the umbilical
wall finely perforate, sutures and keel compressed composed of 2-3 whorls with side alternate in position with the primary
imperforate; aperture a broad low interi- 5-6 chambers in the last whorl; chambers chambers; sutures distinct, flush on the
omarginal arch extending from the narrow with slightly depressed, limbate, spiral side, depressed on the umbilical side;
umbilicus to the periphery, often with a sinuate sutures; spiral suture distinct; periphery rounded; wall finely perforate,
few supplementary areal openings. sutural margin of chambers lobed; surface smooth; aperture an interiomar-
Southern shelf, > 30 m. aperture semicircular at the base of the ginal slit at the base of the final chamber,
Systematics p. 307. flattened semicircular apertural face, on the umbilical side, with a secondary
with radiating striae. opening on the apertural face.
Outer reef, 35 m. Coastal bay, 10 m.
Systematics p. 303. Systematics p. 297.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 197
Hansenisca
Glabratellina Hansenisca soldanii
Glabratellina sp. 1 Test trochospiral; spiral side evolute and
Glabratellina
Test trochospiral, about two whorls with flattened, umbilical side involute and
kermadecensis
usually five chambers in the final whorl; highly convex with subangular umbilical
Test small, trochospiral, concavoconvex, chambers globular, periphery rounded; shoulder bordering the open umbilicus;
circular in end view; spiral side high conical sutures depressed, curved on the spiral sutures radial, straight to slightly curved,
with 3-4 whorls and 4-5 chambers in the side, radial on the umbilical side; surface flush, becoming depressed toward the
last whorl; umbilical side slightly concave of the spiral side heavily ornamented with umbilicus; periphery broadly truncate;
with a depressed umbilicus; peripheral a deeply perforated honeycomb texture; wall finely perforate, surface smooth;
margin acutely rounded; wall coarsely umbilical side with radial rows of granules aperture a short equatorial and interi-
perforated on the spiral side, ornamented and an umbilical extension; aperture an omarginal slit, bordered by a narrow lip,
on the spiral side with honeycomb texture umbilical slit. an umbilical flap extending into the
more or less filled with calcite; ornament umbilicus from each chamber partially
Southwestern lagoon, 30 m.
on the umbilical side, rows of granules covers a small secondary aperture.
Systematics p. 312.
separated by fine striae, some of them
creeping onto the spiral face; aperture in South of the Grande Terre, 40 m.
the umbilical depression. Systematics p. 322.
Coastal bays, 10 m.
Systematics p. 312.
198 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Lamarckina sp. 1
With its rounded periphery and smooth
spiral face, this species resembles
Lamarckina H. haliotidea (HERON-ALLEN & EARLAND,
Lamarckina scabra 1911), but is more elongated than this
Test trochospiral, unequally biconvex, later species.
dorsal side very slightly convex; umbilical Coastal bay, 10 m.
side more convex, peripheral margin Systematics p. 297.
acute, carinate; chambers seven or eight
in the final whorl, rapidly increasing in
size as added; on the umbilical side, final
chamber strongly overlapping and
Derivation of name. The name neocar-
comprising about one-half the umbilical
inata refers to the acute carinate margin
side; sutures limbate on the spiral side,
of this species, neo has been added to dif-
flush or slightly depressed on the umbilical
ferentiate the present species from the
side; wall rugose or granular on the spiral
Cretaceous species Hoeglundina cari-
side, smooth below; aperture interiomar-
nata (N. Bykova).
ginal, umbilical, closed by a thin plate as
Material. Holotype - MNHN F62318, the next chamber is added.
Paratypes - MNHN F62319, MNHN
Southwestern lagoon, 40 m.
F62320, MNHN F62321, MNHN F62322,
Systematics p. 297.
MNHN F62323; from the northern shelf of
New Caledonia, at 600 m water depth.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 201
Lobatula mayori
Test trochospiral, planoconvex to slightly Milesina
concavoconvex, evolute and flattened on Milesina grossepunctata Mississippina pacifica
spiral side, involute and convex on Test planoconvex, subcircular in contour; Test low trochospiral, nearly planispiral,
umbilical side; peripheral outline weakly one side flat and smooth with sutures slightly biconvex, both sides involute;
lobulate in last chambers, peripheral flush, other side convex with three cham- sutures curved on both sides, flush on
margin acute with a carina; 6-8 chambers bers visible having a coarsely perforate spiral side, depressed in last chambers
in the last coil; sutures depressed, radial surface, and slightly depressed limbate of umbilical side; wall smooth with
to slightly curved on umbilical side; on sutures; periphery carinate; aperture a depressed opaque bands, elongate parallel
spiral side curved, broadly limbate and in narrow slit near the umbilicus. to the periphery on both sides; aperture a
last chambers, slightly depressed. Test low arch extending from the umbilicus to
coarsely perforated on spiral side, only the Bay of Prony, 20 m.
the spiral side under an umbilical flap;
last chambers perforated on umbilical side; Systematics p. 314.
apertures of a few last-formed chambers
aperture interiomarginal, extraumbilical- remaining open.
equatorial with thick rim, extending into Northern shelf, 400 m.
a supplementary spiral aperture with rim, Systematics p. 308.
remaining open in the last few chambers.
Living attached on algae, 30-100 m.
Systematics p. 315.
202 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Mychostomina
Mychostomina lucida Mychostomina revertens
Test low conical, composed of a proloculus Test low conical, composed of a proloculus
followed by a tubular enrolled second followed by a tubular enrolled second
chamber with about 6 convolutions visible chamber of several low trochospiral
on the spiral side, then crossing the whorls that then crosses the periphery,
Murrayinella periphery, coiling toward the umbilicus; coiling toward the umbilicus; periphery
Murrayinella globosa 2-3 whorls visible on the umbilical side; rounded; evolute spiral side convex,
Test trochospiral, spiral side evolute, peripheral edge sharp; spiral side convex; umbilical side concave; spiral side perfo-
slightly inflated; umbilical side involute, umbilicus deeply sunk; wall very minutely rated over the entire chamber wall;
inflated; sutures almost radial on both perforated; aperture indistinct. umbilical side irregularly perforated;
sides; about two whorls with usually five Attached on algae, 20-100 m. aperture at the end of the tubular chamber,
chambers in the final whorl; chambers Systematics p. 283. in the umbilicus, with a small lip.
globular with a large spine arising from Coastal bays, 10-20 m.
the midpoint of each chamber; umbilicus Systematics p. 283.
open; wall finely perforate, surface densely
covered with small spines or pustules;
aperture interiomarginal, umbilical to
slightly extraumbilical, obscured by the
ornament.
Coastal bays, estuaries.
Systematics p. 312.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 203
Pannellaina
juvenile Oridorsalis Pannellaina earlandi
Oridorsalis umbonatus Test minute, compressed, subhexagonal
in outline, with slightly inflated chambers
Test trochospiral with both faces convex, and a narrow peripheral keel; all cham-
Nuttallides peripheral edge acute and slightly lobu- bers visible on the spiral side, their center
Nuttallides bradyi late; test consisting of about three whorls concave, and their margins curving up to
Test small, lenticular, trochospiral, almost of nearly equal width, with 5-7 chambers form ridges which are continuous from
equally biconvex, thick-walled; periphery in the last one; sutures distinct, nearly the center of the test to the periphery; only
weakly keeled, slightly lobulate; about radial and limbate on the spiral face, the last whorl visible on the umbilical
three whorls on the evolute spiral side radial and slightly depressed on the face, with sutures depressed and radial;
with 8-10 chambers in the final whorl, umbilical face; wall polished; aperture wall finely perforated on the umbilical
but usually the first whorls are obscured an interiomarginal slit bordered by a side, more coarsely so on the spiral side;
by secondary shell substance; sutures thickened lip; secondary apertures situated aperture a hardly visible narrow slit at
strongly oblique, limbate, and curving at the sutures between the last three the anterior margin of the last-formed
into the peripheral keel; on the involute chambers on the spiral face, and on the chamber on the umbilical side, extending
umbilical side, sutures slightly depressed, umbilicus on the umbilical face. from the periphery to the umbilicus.
nearly radial around the clear imperforate Northern shelf, 600 m. Southwestern lagoon, 40 m.
umbilical boss, but recurved near the Systematics p. 321. Systematics p. 310.
periphery; wall perforate, septa and keel
imperforate; aperture interiomarginal,
extending from the umbilical boss nearly
to the peripheral keel, with a small notch
parallel to the plane of coiling.
Northern shelf, 600 m.
Systematics p. 318.
206 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Pileolina sp. 3
This species resembles P. patelliformis,
but possesses, on the spiral face, rounded
Pileolina zealandica supplementary apertures surrounded by a
thick rim.
Test trochospiral planoconvex; spiral side
low conical with a rounded apex, evolute; Northern shelf, 600 m.
inferior side slightly concave, peripheral Systematics p. 312.
edge acute; 5-7 long and narrow chambers
in the last whorl; flattened side ornamented
with irregularly settled tubercles in a wide Planulina
central area, and deep radiating and Planulina ariminensis
branching striae to the periphery, obscuring
the sutures between chambers; wall dis- Test discoidal, very low trochospiral of
tinctly perforated on the conical side, about two whorls, with both sides flat-
finely so on the flattened side; aperture on juvenile tened; spiral side evolute, umbilical side
the flattened side, hardly discernable. partially evolute; 8-10 broad, low, and
arched chambers in the final whorl; septa
Northern shelf, 600 m. thick, sutures imperforate, thickened and
Systematics p. 312. Planodiscorbis elevated, strongly curved back at the
Planodiscorbis rarescens peripheral margin; periphery with a thick
Test adherent, trochospiral, planoconvex; imperforate marginal keel; wall coarsely
spiral side flattened, somewhat depressed perforated on the spiral side, finely
at the umbilicus; peripheral edge sharp, perforate with scattered larger pores on
extended in a well-defined, imperforate the umbilical side; aperture an equatorial
keel; only the 4-6 chambers of the outer- and interiomarginal arch with an imper-
most whorl visible on the convex umbilical forate bordering lip, extending somewhat
side; sutures very slightly depressed, onto the umbilical side beneath the
Pileolina sp. 1 strongly oblique on both sides; each imperforate umbilical folium.
Species similar to P. minogasiformis, chamber projecting a “valvular lobe” Northern shelf, 600 m.
but with a long spine-like projection over the umbilicus; aperture a narrow slit Systematics p. 314.
arising from the apex. It may be a variety with a slightly thickened lip running
of the former species. from upper fourth of umbilical side
Northern shelf, 600 m. across periphery and onto spiral side.
Systematics p. 312. Northern shelf, 600 m.
Systematics p. 310.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 209
old specimen
210 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Rosalina
Rosalina bradyi
Test trochospiral, concavoconvex, periphery
rounded; all chambers visible on the convex
Robertinoides bradyi spiral side where the earlier chambers are
Test high trochospiral, fusiform, about clearly outlined with limbate sutures;
1.5 times as long as broad; chambers very sutures curved back at the periphery; Rosalina globularis
slightly inflated, arranged in 2-3 more or umbilical side with the chambers more Test ovate in outline, spiral side highly
less regular, oblique whorls; sutures or less inflated; last-formed chamber convex and evolute, umbilical side plane
indistinct and outline not lobulate; initial with a more or less bifid indentation to almost concave and involute; five
end bluntly pointed, oral end broadly extending inward from umbilical region; chambers in the last whorl; peripheral
rounded; wall finely perforate, surface spiral surface coarsely perforated, umbil- margin broadly rounded; sutures on
smooth; aperture two slit-like openings, ical surface smooth and imperforate; spiral side curved and depressed, on
one located at the proximal margin of aperture a low interiomarginal arch on umbilical side somewhat indistinct;
the chamber, the other diverging from the umbilical side, with narrow bordering umbilicus open, chambers with a trian-
chamber margin and directed up the lip. gular folium and hook-shaped sutural
apertural face. Southwestern lagoon, 30 m. notches; spiral side densely and coarsely
Northern shelf, 600 m. Systematics p. 310. perforate, umbilical side smooth, sparsely
Systematics p. 298. perforate; aperture interiomarginal with
a low lip in the peripheral-most region,
aperture continues to umbilicus, under
the folium.
Southwestern lagoon, 40 m.
Systematics p. 310.
212 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Rotorbinella
Rotorbinella lepida
Test, low trochospiral, biconvex; spiral side
evolute, convex, central part of the spire,
completely embedded in translucent shell
material; umbilical side involute, flattened; Rotorbis pacifica
umbilicus filled with a rounded umbilical
plug; periphery slightly lobate, bluntly Test free, trochospiral, evolute spiral side
angled with a thickened imperforate slightly convex; involute umbilical side
carina; 5-7 chambers in final whorl, almost flat, periphery angled and slightly
slightly inflated, gradually increasing in keeled, 6-8 chambers in the last whorl;
Rosalina rugosa size as added, provided with a short chambers crescentic, increasing progres-
Test low trochospiral, compressed, only backward directed folium; folia extending sively in size, slightly inflated at umbilical
slightly convex on the spiral side and over the margin of the plug with re-entrants side; the few last chambers generally are
slightly concave on the umbilical side; on each suture and fusing with the umbil- less regularly arranged, and are more
peripheral edge round and lobulate; ical plug in earlier chambers; sutures on inflated on the spiral side; sutures curved
chambers inflated, 5 in the last coil; spiral side arcuate, oblique, flush; on on spiral side, radial on umbilical side,
sutures depressed; umbilical cavity par- umbilical side slightly arcuate, almost spiral suture somewhat raised above
tially covered by the folia protecting radial, depressed; wall smooth, perforate surface; wall distinctly and rather coarsely
the successive apertures; wall coarsely on umbilical side, only the marginal zone perforate on umbilical side, finely perforate
perforated; aperture interiomarginal of the last chambers perforate on the on spiral side; surface of the spiral side
with a narrow lip, continuing under the spiral side; aperture an interiomarginal covered with irregularly arranged pustules;
folium slit, at the base of the last-formed chamber, aperture a slit-like interiomarginal arch,
that extends from the periphery to the partially hidden by a lip running from
Southwestern lagoon, 40 m. the umbilicus to the peripheral keel, with
Systematics p. 310. umbilicus, bordered on the upper margin
by a thickened lip that continues into the a supplementary re-entrant aperture.
folium, differing from the aperture of Northern shelf, 600 m.
Gavelinopsis praegeri. Systematics p. 309.
Southwestern lagoon, southern shelf,
10-100 m.
Systematics p. 309.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 213
Rupertina Siphonina
Rupertina pustulosa Siphonina tubulosa
Stomatorbina sp. 1
Test attached by a prominent basal disc; Test low trochospiral, lenticular, circular
chambers trochospiral in the early stage, in outline; a few broad, low and crescentic One individual resembling S. concentrica,
later loosely coiled, growing spirally chambers per whorl; sutures somewhat but with a big test much more rounded
around a solid central column; chambers indistinct, oblique, thickened on the spiral than this later species was found.
inflated, sutures depressed; wall thick, side and continuing into the wide fimbriate Northern shelf, 600 m.
coarsely perforate; surface pustulate; peripheral keel; sutures radial and Systematics p. 308.
aperture a low umbilical slit bordered by depressed on the umbilical side; umbilicus
a distinct lip. closed: wall finely perforate, surface
Northern shelf, 400 m. ornamented with conical pustules; aper-
Systematics p. 317. ture elliptical, just above the base of the
final chamber and nearly equatorial in
position, produced on a short neck and
bordered with a phialine lip.
Southwestern lagoon, and southern shelf,
30-70 m.
Systematics p. 313.
214 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Valvulineria
Valvulineria candeiana
Test low-trochospiral, composed of
Ungulatelloides 2-2.5 coils, the last-formed coil composed
Ungulatelloides cf. U. imperialis of about 6 chambers, rapidly increasing
Svratkina in size, inflated, the last-formed chamber
Test free, small, trochospiral; all chambers often somewhat deformed; periphery
Svratkina australiensis visible on the spiral side, only those of the
Test trochospiral, biconvex, spiral side lobulate, rounded; sutures curved back-
last whorl visible on the umbilical side; wards, depressed on both sides, except in
evolute, six to seven crescentic chambers proloculus provided with a funnel-shaped
in the final whorl, sutures oblique and the early portion of the test; deep umbilicus
lamella, followed by an undivided spiral partly covered by small flaps; wall trans-
curved; umbilical side involute, sutures chamber one or two coils in length, and
radial and slightly depressed around the parent, distinctly perforate on both sides,
then by two or more chambers per whorl; except in the early chambers on the spiral
closed umbilicus, periphery rounded; last chambers provided with a curved flap;
wall coarsely perforate, the large pores side; aperture a narrow, arched slit at the
wall transparent, finely perforated; aper- base of the last-formed chamber with a
opening at the center of small tubercles ture a broad, arched slit with a weak lip
that cover both sides; aperture an elongate slight lip.
on the umbilical side of the last chamber.
oblique opening in a slight depression This species differs from U. imperatrix Southwestern lagoon, 30 m.
extending from near the umbilicus up to by the presence of a few spines only, even Systematics p. 307.
the face of the final chamber. if the characteristic shape that gives in
Southwestern lagoon, 40 m. side view the aspect of a hat with spines
Systematics p. 320. on the brim is similar.
Outer reef, 100 m.
Systematics p. 311.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 215
Astacolus neomulticamerata
Test irregularly subovate in contour, Astacolus sp. 1
compressed, periphery rounded, non-
lobulate; initial end a partial planispiral Test elongate in outline, laterally com-
coil with chambers almost completely pressed, chambers numerous, broad and
evolute, increasing rapidly in breath, low, enrolled in the very early stage, later
with tendency to become uniserial, some uncoiling, added on a slightly curved axis
Astacolus with strongly oblique, curved sutures;
Astacolus crepidulus overlapping along the concave margin;
sutures obliquely curved; wall finely periphery rounded; surface smooth;
Test elongate, flattened, chambers broad perforate, surface smooth; last chamber aperture radiate, at the dorsal angle.
and low becoming gradually broader, prominent, centered, with a slightly Northern shelf, 600 m.
first stage planispirally enrolled, later produced radiate aperture. Systematics p. 288.
uncoiling with chambers added on a
slightly curved axis; sutures strongly Northern shelf, 600 m.
oblique, curved, slightly if at all Systematics p. 288.
depressed; peripheral margin subround-
ed; wall very finely perforate, surface
smooth; aperture radiate, at the dorsal
angle, with a longer slit towards the aper-
tural face.
Northern shelf, 600 m.
Systematics p. 288.
218 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Astrononion
Astrononion novozealandicum Elphidium
Test planispiral, involute, laterally
Elphidium advenum
compressed, bilaterally symmetrical, Test involute, equally biconvex, lenticular;
umbilicate; periphery rounded to some- periphery subcircular, somewhat lobulate Elphidium charlottense
what angular; around 10 chambers in the last-formed portion; sides convex;
peripheral margin acutely rounded, often Test involute, biconvex, compressed, sub-
enlarging gradually; sutures depressed, circular in outline, slightly lobulate in later
covered by an elongate sutural plate with a blunt keel; chambers numerous,
somewhat inflated, especially in the last- stage; sides flattened to slightly concave;
extending from the umbilicus to half peripheral margin acutely rounded with a
the distance to the periphery, with a small formed portion, gently curved backward;
sutures depressed, with sutural bridges strong, rounded keel; 12-15 non-inflated
pit at the end of each sutural plate; to slightly inflated chambers in final
successive plates fuse in the umbilical only about onefourth the width of the
chamber; umbilical region usually with a whorl; sutures flush to slightly depressed,
region; wall thin, distinctly and densely gently curved, with short sutural bridges;
perforate, surface smooth; aperture a low boss; wall smooth, translucent, very finely
perforate, ornamented by numerous umbilical area occupied by a flattened
interiomarginal, equatorial slit, bordered boss not protruding beyond the outline of
with a lip and extending laterally to the small pustules around the aperture, the
sutures and the umbilical boss; aperture the test; wall finely perforate, ornamented
umbilici. with fine papillae along the sutures and
a series of rounded pores, at the base of
Southern shelf, 70 m. around the umbilical boss; aperture a row
the apertural face.
Systematics p. 319. of openings at the base of the last chamber.
Southwestern lagoon, 30 m.
Systematics p. 323. Coastal bay, 20 m.
Systematics p. 324.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 219
Elphidium excavatum
Test planispirally enrolled, laterally
compressed with broadly to acutely
rounded periphery; sides almost parallel;
7-10 chambers in the last whorl, some-
what inflated, leading to a lobulate
Elphidium craticulatum periphery; sutures depressed, slightly
Test large, circular in outline, strongly curved with short and thin sutural
biconvex, with a very large, pitted umbil- bridges; umbilical region depressed with
ical boss; periphery acute with a narrow fine papillae and discrete extensions of
rounded keel; chambers numerous, the chambers (folia) that do not coalesce,
about 30 in the last whorl; apertural face giving the umbilical region a star-shaped
appearance; wall finely perforated; Elphidium fijiense
very low; sutures almost straight, radial;
sutural bridges extending on the cham- aperture at the base of the last chamber, Test of small to medium size, outline
ber wall, resulting in continuous ridges obscured by papillae. smoothly circular, lobulate in the last
parallel to the periphery; aperture a row Coastal lagoons, coastal bays, shrimp portion; profile biconvex; periphery
of multiple openings along the base of the ponds. acutely rounded with a slightly thickened
final chamber. Systematics p. 324. shell; 14-18 chambers in the last whorl;
sutures very slightly curved backward
Southwestern lagoon, in bays and depres- towards periphery, with narrow septal
sions, 1-30 m. bridges; umbilical area with a prominent,
Systematics p. 324. irregular glassy boss; fine papillae in the
sutures, the depressions in the umbilical
boss, and along the aperture; wall finely
perforate; aperture a series of small
openings at the base of the apertural face.
Estuaries, marshes.
Systematics p. 324.
220 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Elphidium hyalocostatum
Test involute with circular outline; test Elphidium limbatum Elphidium maorium
moderately inflated; 7-10 chambers in the
last whorl, non-inflated and strongly Test involute, laterally compressed, Test of small to medium size, outline
curved back toward the periphery; anterior lenticular, with a subcircular periphery; smoothly circular, lobulate in the last
edge of each chamber bearing a strong, chambers slightly inflated and gently portion; profile biconvex with sides flat
narrow, radial ridge; long sutural bridges curved backward, 11-14 in the last whorl; and parallel centrally; periphery acutely
extending forwards from the radial ridges, subacute peripheral margin developing a rounded with a thickened area along the
together forming a coarse reticular pat- blunt carina in the early chambers; periphery that may be slightly keeled;
tern; umbilical area small and slightly sutures depressed with numerous flat 10-13 slightly inflated chambers in the last
depressed; apertural face and the area sutural bridges, reaching about one half whorl; sutures slightly curved backward
around its base covered with papillae that the width of the chambers; umbilical towards periphery, with a few narrow
obscure the basal aperture. region depressed, containing numerous septal bridges; umbilical area with a small
fine papillae and often one or several solid circular boss; fine papillae in the
Northern shelf, 600 m. larger tubercles; wall finely perforate, sutures, the umbilical area and along the
Systematics p. 324. ornamented with fine papillae along the aperture; wall finely perforate; aperture a
sutures and on the apertural face; aperture series of small openings at the base of the
a row of basal openings. apertural chamber.
Coastal bay, 10 m. Coastal bay, 10 m.
Systematics p. 324. Systematics p. 324.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 221
Elphidium sp. 1
This species resembles E. advenum, but
the periphery becomes lobulate at about
Elphidium oceanicum Elphidium tongaense half the last whorl; later chambers
Test planispirally enrolled, slightly com- Test of medium to large size, planispiral, strongly pointed, the last ones developing
pressed laterally; chambers increasing slightly evolute, compressed, outline spines.
rapidly in width, last ones somewhat slightly lobulate; sides flat to slightly Coastal bay, 30 m.
inflated, leading to a lobulate periphery; concave, periphery acutely rounded; 10- Systematics p. 325.
peripheral margin broadly rounded; 12 somewhat inflated chambers in the
sutures slightly depressed, continuous last whorl; sutures depressed, moderately
around the periphery, with short flush curved backwards, marked by numerous
sutural bridges; dense hispid ornament narrow septal bridges; umbilicus large,
extending from the wide depressed partly filled with a solid semicircular
umbilicus to a large area surrounding the boss; wall finely perforated, fine papillae
aperture; aperture at the base of the last in the umbilical area and in the sutures;
chamber, obscured by the ornamentation. aperture made up of numerous openings
Coastal lagoons, coastal bays, shrimp at the base of the apertural face.
ponds. Isle of Pines, 5 m.
Systematics p. 325. Systematics p. 325.
222 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Haynesina
Haynesina depressula
Lenticulina
Test involute, compressed, evenly rounded
to lobulate in outline; peripheral margin Lenticulina australis
rounded; 8-14 slightly inflated chambers Test planispiral, involute, biconvex, robust,
in final whorl; sutures curved backwards, Heterostegina operculinoides subcircular in outline; periphery bluntly
slightly depressed in early portion, rounded with a narrow keel; 6-7 cham-
Test planispiral evolute extremely flat; bers in the final whorl; umbilical region
becoming deeply depressed; large chambers falciform, rapidly increasing in
umbilicus that extends along the sutures flush and transparent; sutures flush and
peripheral elongation, but slowly in limbate, gently curved; apertural face
producing a star shape; wall finely per- height; chamber subdivision starts early,
forate with tubercles along the sutures, truncated, slightly depressed, with a thick
with chambers completely subdivided limbate margin; aperture radiate, at the
in the umbilical depression and around into rectangular chamberlets; chamber
the aperture; aperture a series of openings peripheral angle, with a distinctly
sutures raised above the test surface and enlarged equatorial slit on the apertural
at the base of the last chamber, obscured sometimes beaded; chamberlet sutures
by the tubercles. face.
raised or depressed, forming a more or less
Coastal lagoons, coastal bays, shrimp regular alternating pattern; a few knobs are Northern shelf, 600 m.
ponds. located on the central test part, obscured Systematics p. 286.
Systematics p. 319. by granulation that decreases with growth;
wall finely perforate; aperture equatorial,
at the base of the narrow apertural face,
masked by a thick shell deposit.
Southern lagoon and southern shelf,
frequency increasing with depth.
Systematics p. 325.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 223
Lenticulina cultrata
Test large, discoidal and biconvex;
periphery acute, somewhat irregular, Lenticulina nitida
with a well-marked keel; 6-7 chambers in Test planispiral, involute, biconvex com-
the last whorl; sutures slightly curved, pressed and carinate; periphery lobulate;
converging to the umbilicus; no umbo, Lenticulina gibba 6-8 chambers in the final whorl, slowly
but a translucent central area; wall Test oblong, biconvex, with a tendency increasing in size as added; earlier
smooth devoid of ornament; aperture towards uncoiling in the last few cham- sutures limbate and raised, later
radiate with several slits converging to a bers; periphery acutely rounded to slightly depressed; wall smooth, finely perforate;
median wall at the periphery, and an keeled; 7-8 chambers in the last whorl, apertural face truncated, with a thick
equatorial slit on the apertural face, elongate, strongly curved, increasing limbate margin; aperture radiate, at the
between two rectilinear lips that diverge slowly in size as added; sutures curved; peripheral angle, with a distinctly
at the lower extremity. apertural face convex to truncated; wall enlarged equatorial slit on the apertural
Specimens referred to this species vary smooth, finely perforate; aperture finely face.
greatly in size, thickness, and morphology radiate, at the peripheral angle, with a Southwestern lagoon, 60 m.
of the aperture. slightly enlarged equatorial slit on the Systematics p. 286.
Northern shelf, 600 m. apertural face.
Systematics p. 286. Southern shelf, 45-85 m.
Systematics p. 286.
juvenile
224 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Lenticulina sp. 1
Test close-coiled, involute, lenticular, bi-
umbonate; chambers curved, increasing
slightly in size as added; sutures and Lenticulina? sp. 3
umbo flush, giving a very smooth surface Test close-coiled, entirely involute, bicon-
to the test; peripheral margin acutely vex, broadly rounded; 6-7 chambers in
rounded; apertural face small, rounded; the last-formed coil; apertural face small,
wall very finely perforate; aperture radiate, rounded; sutures flush, curved; wall
transformed into an elongated double row smooth; aperture at the peripheral angle
of lateral slits that converge towards an of the test, radiate with a series of radi-
axial wall, itself with a central opening; ating slits at the periphery, but with a Species 2
apertural face partially divided by an tendency to become cribrate, due to
elongate equatorial slit somewhat radiating Periphery acute but not keeled, apertural
irregularly settled slits in the center.
at its base. face truncated with two angular shoulders;
Northern shelf, 600 m. 8-9 chambers increasing relatively slowly
Northern shelf, 600 m. Systematics p. 287. in size as added; sutures flush, limbate,
Systematics p. 287. curved, converging to the large umbo
flush with the surface; aperture composed
of 2-3 short radiating slits on each side at
the periphery and an equatorial slit
between two lips on the apertural face.
Northern shelf, 600 m.
juvenile Systematics p. 287.
226 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Species 4
Periphery acute, carinate, apertural
face truncated, somewhat concave, with
two angular shoulders; 7-8 chambers in Melonis pompiloides
the last coil, increasing relatively slowly Species 6 Test planispiral, involute, compact,
in size as added; sutures raised, strongly Periphery carinate, apertural face trun- peripheral margin very broadly rounded;
curved, converging to the umbilicus cated, somewhat concave, with two chambers numerous, 10-14 in final
without umbo; aperture composed of 7- angular shoulders; 9-11 chambers in the whorl; sutures flush, later ones slightly
8 radiating slits at the periphery and an last coil, increasing relatively slowly in depressed, slightly curved; deeply sunk
equatorial slit between two projecting lips size as added; sutures strongly curved, umbilici; walls coarsely perforated; aper-
on the apertural face. converging to the small umbilicus; ture terminal, low, broad arch extending
Northern shelf, 600 m. sutures discontinuously raised, a few from umbilicus to umbilicus.
Systematics p. 287. tubercles arranged parallel with the Northern shelf, 600 m.
sutures, and faint costae parallel with the Systematics p. 320.
carina; aperture composed of 7-8 radiating
slits at the periphery and an equatorial
slit between two projecting lips that nearly
reach the base of the apertural face; small
rounded openings are present in the
median wall of the aperture.
Northern shelf, 600 m.
Systematics p. 287.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 227
Nonion scaphum
Test planispiral, bilaterally symmetrical,
compressed, elongate oval in outline,
may tend to become evolute; peripheral
margin rounded; umbilici depressed, Nonionoides
unornamented; 10-12 chambers in the Nonionoides grateloupi
last coil, later ones rapidly increasing in Test compressed, low trochospiral, com-
size and broadening towards the umbilici; posed of tall, narrow chambers, rapidly
sutures depressed, not limbate; apertural
Nonion increasing in length as added; spiral side
face broadly oval; wall smooth, finely evolute and umbilical side involute, but
Nonion grossepertusum perforate; aperture, a narrow slit at the the symmetrical evenly flattened sides
Test planispirally enrolled, involute, base of the apertural face next to the give the impression that the coiling is
slightly inflated; 8 chambers in the final preceding coil. planispiral; wall translucent, smooth,
whorl increasing rapidly in size as added; Bay of Prony, 20-40 m. with hispid ornament in the umbilical
periphery rounded; sutures flush to Systematics p. 319. depression, along the spiral suture and
weakly depressed; wall smooth but around the aperture; aperture an equa-
coarsely perforated; aperture a series of torial slit at the base of the apertural
pores at the base of the apertural face. face, extending from the suture on the
Southwestern shelf, 70 m. umbilical side to a short way along the
Systematics p. 319. spiral suture on the spiral side.
Bay of Prony, 10-40 m.
Systematics p. 320.
228 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile
Operculina gaimardi
Nummulites Test much compressed; coiling planispi-
ral evolute, but earlier whorl more or
Nummulites venosus less embracing; central area somewhat
Test planispirally coiled, biconvex and thickened; chambers 15 or less in the last
lenticular, evolute in young individuals, Operculina bartschi whorl, strongly and regularly curved
later involute, and finally tending to Test evolute, lenticular, coils rapidly toward the periphery and toward the
become evolute again with a slight widening, but not flaring; umbilical area umbilicus; sutures limbate, marked by
marginal flattening (see also fig. 52); usually raised and ornamented with chains of beads, which are larger near
chambers numerous; periphery with a comparatively large bosses; chambers the centre of the shell; peripheral chord
thick marginal cord; sutures fairly distinct, narrow, somewhat inflated, 20-25 in the distinct; surface of the chambers smooth
raised, radiate, recurved at the periphery last whorl, arcuate and sharply recurved and unornamented or covered with
and sometimes branched around the at the periphery; sutures depressed; minute granules scattered over the sur-
umbilicus; wall smooth, finely perforate; peripheral chord distinct; surface orna- face; aperture at the base of the apertural
aperture at the base of the apertural face, mented with a row of tubercles along the face concealed under a mask structure.
masked by a thick shell deposit. median line of the chambers; tubercles Southwestern lagoon and southern shelf,
Southwestern lagoon, depressions, 5-45 m. higher in the earlier portion of the test; 10-80 m.
Systematics p. 326. aperture at the base of the apertural face Systematics p. 326.
concealed under a mask structure.
juvenile
Southwestern lagoon, areas under open-
sea influence.
Systematics p. 326.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 229
juvenile
Planispirillina
tuberculatolimbata
Test discoidal, globular proloculus followed
by undivided tubular and planispirally
Palmula enrolled second chamber, evolute on one
Palmula robusta flat side and very slightly involute on the
Test elongate, compressed, stoutly built, opposite convex side; test strongly dissym-
fan-shaped; lateral margins tending to metrical, the larger and flat side being
become parallel with ends obtusely angu- limbate with a sharp peripheral edge, the
lar or rounded, peripheral edges thick, opposite convex side having a rounded
rounded, slightly lobulate; chambers Planispirillina edge; wall perforate between the raised
numerous, somewhat irregular in contour, spiral sutures on the flat side, imperforate
Planispirillina inaequalis on the opposite side where earlier whorls
chevron-shaped; surface furnished with
closely set, interrupted, longitudinal costae Test discoidal, consisting of a proloculus are obscured by a covering of papillose
aperture terminal, slightly produced, and a long undivided, gradually enlarging, lamellae; aperture at the end of the tubular
radiate. coiled, tubular second chamber; coiling chamber.
planispiral, but with the tubular chamber Chapman mentioned that the convex side
Northern shelf, 600 m. overlapping slightly more on one side
Systematics p. 288. is partially involute but his illustrations
than on the other, so that the test has shows the coiling of the tubular chamber
distinct sides; evolute side flattened, the to be evolute, all the whorls but the last one
other side partially involute with a being obscured by the ornamentation.
concave umbilical area; periphery acute
with, on the umbilicate side, a coarsely Northern shelf, 200 m.
perforate raised margin that slopes gently Systematics p. 283.
inwards; flat side with a slightly raised
spiral suture, but with no secondary
lamination or ornamentation; aperture
at the end of the tubular chamber.
Northern shelf 200 m.
Systematics p. 283.
230 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Saracenaria latifrons
Saracenaria Test spiral, elongate, trihedral, broadest
Saracenaria altifrons near the middle and tapering towards the
Test ovoid, planispirally enrolled in the ends; initial portion pointed, with early
early stage, later flaring and tending to chambers small and involute; later
become rectilinear, triangular in section, chambers long, narrow, slightly curved,
apertural face broad, nearly flat, with obliquely set; “dorsal margin” acutely
acutely rounded margins and dorsal angular and carinate; apertural face broad,
Pullenia angle; sutures curved, slightly depressed; oval, somewhat curved, with partially
Pullenia quadriloba wall finely perforate, surface smooth; carinate lateral edges; aperture radiate, at
aperture radiate at the dorsal angle. the acute dorsal angle.
Test globular, planispiral and involute,
with 4 moderately inflated chambers in Northern shelf, 600 m. Northern shelf, 600 m.
the final whorl, sutures radial, slightly Systematics p. 287. Systematics p. 287.
depressed; wall finely perforate, surface
juvenile
smooth; aperture a narrow interio-
marginal crescentic slit extending across
the periphery, but not reaching the
umbilici.
Northern shelf, 600 m.
Systematics p. 320.
232 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Sejunctella Spirillina
Sejunctella cf. S. wenmanensis Siphomarginulina sp. 1 Spirillina grosseperforata
Test discoidal, proloculus followed by a Test elongate, compressed, periphery Test flattened, circular in outline, consisting
planispirally enrolled undivided tubular acutely rounded, early chambers of a proloculus and a long undivided,
second chamber that is loosely coiled and planispirally enrolled, only three to four gradually enlarging, coiled, tubular second
separated from the previous whorl by a chambers in the coil, final few chambers chamber; periphery subacute; coiling
narrow solid area; planispiral coiling of uncoiled and rectilinear, sutures radial in planispiral, but with the tubular chamber
about 3 whorls; periphery carinate, bearing the early coil, oblique in the uncoiling overlapping slightly more on one side than
numerous, relatively long, evenly spaced part; wall finely perforate, surface on the other, so that the earliest whorls
solid spines, somewhat thickened at the smooth; aperture at the dorsal angle, are in a low trochospiral arrangement,
top; wall hyaline, with secondarily added consisting of a projecting ring of rounded with distinct spiral and umbilical sides;
granulations; aperture a simple rounded openings. later coils flattened on one side; wall thin,
opening at the end of the tubular chamber. coarsely perforated on the spiral side,
Northern shelf, 200 m. roughly ornamented with transverse ridges
This species differs from McCulloch’s one Systematics p. 287.
in its granular ornamentation. and not perforated on the umbilical side;
suture depressed; aperture formed by the
Outer reef, 100 m.
open end of the tube.
Systematics p. 283.
Bay of Prony, 40 m.
Systematics p. 283.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 233
Vaginulinopsis
Vaginulinopsis gnamptina
Test broad, arcuate, with rounded periph-
ery; early enrolled portion followed by
Spirillina vivipara Spirillina sp. 2 uncoiled part with chambers increasing
Test flattened, circular in outline, consist- Test flattened, circular in outline, consisting rapidly in breath, and with considerable
ing of a proloculus and a long undivided, of a proloculus and a long undivided, overlap over previous chambers; sutures
slowly enlarging, plan spirally coiled, slowly enlarging, planispirally coiled, slightly curved, moderately depressed;
tubular second chamber; periphery tubular second chamber; periphery surface smooth, aperture a long slit on a
rounded; coils numerous, later ones often rounded; wall smooth, with a row of produce flange, at the dorsal angle.
somewhat uneven and not entirely perforations irregularly disposed axially Northern shelf, 600 m.
planispiral; wall calcareous, coarsely on the tube; ornamentation with well- Systematics p. 288.
pitted, thin; suture depressed; aperture developed spines, pointing toward one side
formed by the open end of the tube. of the test, so that they are leaved free by
Living attached on algae, 10-100 m. the subsequent whorls; aperture terminal.
Systematics p. 283. Northern shelf, 600 m.
Systematics p. 283.
234 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
juvenile Baculogypsina
cf. B. sphaerulata
Specimens with large spines arising in all
directions but not only in the equatorial
plane, and sometimes bifurcated; may be
abnormal specimens of Baculogypsina
sphaerulata or may belong to another
species. Further examinations are needed
to determine if this variation has any
taxonomic significance.
Chesterfield, living in algal thalli growing
on coral reefs, 1-2 m.
Systematics p. 323.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 235
Cycloclypeus
Cycloclypeus carpenteri
Test discoidal, nearly circular in outline;
juvenile biconvex or subumbonate; central portion
thickened, the outer zones gradually
thinning towards the thin, sharp, peripheral
edge; test composed of annular chambers
divided by septulae into chamberlets that
alternate in position more or less regularly
Burseolina from one chamber to the other; chamber-
Burseolina pacifica lets approximately square in the smaller
Test biserially arranged with the plane of megalospheric form, radially elongated
biseriality planispirally enrolled; test in the larger microspheric forms; sutures
large, compact, globular to subglobular, slightly raised, peripheral ends of septula
very slightly compressed; periphery not marked with a rounded knob; large
lobulate; peripheral margin broadly knobs also covering the thick central part;
rounded; chambers low and broad; surface of the chamberlets smooth, finely
sutures flush virtually indistinguishable perforated; aperture a row of marginal
except when the test is moistened; wall pores.
finely perforate, surface smooth; apertural Southern shelf, 60-80 m.
face broad and depressed; aperture an Systematics p. 325.
elongate narrow slit, curved so that its
outermost part is approximately parallel
to the periphery of the test; lower margin
with a large imperforate apertural flap
and a poorly developed toothplate on the
upper apertural margin.
Northern shelf, 600 m.
Systematics p. 300.
236 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Globocassidulina subglobosa
Test biserially arranged with the plane of
biseriality planispirally enrolled; test
globular, periphery broadly rounded, not
Globocassidulina decorata lobulate, last chamber somewhat pro-
jecting; sutures indistinct; wall coarsely
Test biserially arranged with the plane of perforated, with depressed pore margins,
biseriality planispirally enrolled; test Globulina
apertural face imperforate with faint Globulina gibba
medium, subglobular to globular, with striations radiating from the aperture;
last chamber slightly protruded; sutures aperture a narrow elongate opening Test generally nearly spherical to some-
indistinct, obscured by the ornamentation; extending almost orthogonally over the what compressed, rounded at the base;
wall decorated by a network of irregular apertural face; a small lip is present on apertural end variable, either acuminate
costae, strongly reticulated on earlier one margin, and a narrow tooth extends with the aperture situated on a mammil-
chambers and progressively less developed along the opposite side. late protuberance, or truncate and the
toward the unornamented last chamber; general aperture flush with the body of
aperture with two branches, one along the Northern shelf, 600 m. the test; usually three compactly joined
suture, at the base of the apertural face, Systematics p. 300. and overlapping chambers visible;
the other extending into the apertural sutures neither excavated nor depressed;
face; aperture bordered by a thickened wall finely perforated, smooth; aperture
apertural ridge and a triangular apertural radiate, but commonly obscured by fistu-
flap. lose growth.
Northern shelf, 600 m. Northern shelf, 200 m.
Systematics p. 300. Systematics p. 290.
240 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
1mm
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 241
Laryngosigma
Laryngosigma Laryngosigma williamsoni Laryngosigma sp. 2
afueraensis
Test elongate ovate, somewhat compressed, Test compressed, fusiform, periphery
Test elongate, much compressed, sub- sides nearly parallel; chambers biserially rounded; initial end narrowly rounded,
quadrate in contour, periphery rounded; arranged and sigmoid, added in planes protruding, apertural end broadly rounded;
initial end broadly rounded, width almost slightly less than 180° apart; initial end chambers broad, elongate, added in a
uniform; chambers biserially arranged rounded; chambers elongate and strongly biserial sigmoid series in planes slightly
and sigmoid, added in planes slightly less overlapping; sutures flush, oblique; wall less than 180° apart, each succeeding
than 180° apart; initial end rounded; transparent, surface smooth; aperture chamber farther removed from the base;
chambers elongate, increasing rapidly in terminal, radiate. sutures depressed; wall finely perforate,
length, quite slowly in width and strongly surface smooth; aperture large, terminal,
overlapping; sutures flush, diagonal with Northern shelf, 200 m.
Systematics p. 296. radiate.
sharply curved ends; wall transparent,
finely perforated, surface smooth; aper- Southwestern lagoon, 40 m.
ture terminal, radiate, with a short stout Systematics p. 296.
entosolenian tube.
Northern shelf, 600 m.
Systematics p. 296.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 243
Lernella Millettia
Lernella inflata Millettia limbata
Test biserially arranged with the plane of Test elongate, compressed, tapering,
biseriality planispirally enrolled; test somewhat twisted or otherwise irregular;
medium, subcircular to ovate in side proloculus followed by two biserially
view, slightly compressed; periphery arranged chambers; later chambers recti-
broadly rounded, distinctly lobulate; last linear or slightly arcuate, few chambers
four chambers may occupy 2/3 of the cir- making up the test; apertural end rounded,
cumference of the test; sutures distinct, initial extremity angular or pointed;
depressed, gently curved; wall finely per- surface marked by a number of raised
forated, smooth; aperture a curved, elon- transverse bands of shell-substance con-
gate slit at the base of the last chamber nected by a similar band on the median
with a large apertural flap. line on either side of the test, which may
Northern shelf, 600 m. become irregular in later chambers; wall
Systematics p. 300. perforated, with perforations more distinct
on the ornamentational ridges; aperture
terminal, produced on a neck.
South of the Grande Terre, 30 m.
Systematics p. 305.
244 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
red form
Paracassidulina
Paracassidulina angulosa
white form
Test biserially arranged with the plane of
biseriality planispirally enrolled; test
compressed, lenticular; chambers elon-
Miniacina sublarvata gate, five pairs in the last whorl; periph-
ery strongly lobulate, with prominent
Test discoidal, flattened, nearly symmet- bosses; wall smooth; aperture a long nar-
rical, periphery rounded; irregular- row slit along the suture, with a narrow
shaped chambers added in irregular lip along its inner margin.
series; wall calcareous, more coarsely
perforated on one side than the other; Northern shelf, 600 m.
aperture indistinct due to coarse perfo- Systematics p. 300.
ration.
Southern and northern shelf.
Systematics p. 318.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 245
Paracassidulina sulcata
Test biserially arranged with the plane Physalidia
of biseriality planispirally enrolled; test Pegidia Physalidia? earlandi
small, nearly circular in side view, com- Pegidia dubia Test reniform, characterized by a few
pressed, oval in end view; periphery Test unequally biconvex and irregular in globular chambers arranged in an elon-
slightly lobulate; peripheral edge rounded; shape, ranging between a regular oval gate trochospire; chambers inflated,
chambers broadly rhomboid, very slightly and a quadrate contour with rounded subspherical, variable in shape, increasing
inflated; sutures distinct, deep; wall finely corners; periphery with a broad smooth regularly in size as added; initial stage
perforated, smooth, but with irregularly keel; reduced trochospiral coiling of two constituted of a relatively small proloculus
distributed grooves running out from the chambers per whorl, rapidly enlarging as and 3 closely-enrolled chambers; sutures
sutures and aperture; aperture an elongate added; spiral side domed, densely covered deeply depressed; wall very thin, hyaline,
narrow slit along the suture, bordered on with tubercles that obscure the sutures; smooth, irregularly but distinctly perfo-
its upper margin by radiating grooves umbilical side flattened; wall thick, finely rated, with a non-perforate area just
and on the other side by a thin apertural perforate, spiral side strongly tuberculate, below the aperture; aperture a fine slit at
ridge. umbilical side smooth; aperture consisting the base of the final chamber, bordered
Northern shelf, 600 m. of openings of tubular canals along the by a distinct poreless lip on the upper
Systematics p. 300. sutures on the umbilical side. margin.
Southwestern lagoon near passes; southern The unique specimen of this species
shelf, 40-60 m. resembles the specimens illustrated by
Systematics p. 308. PARKER (2009) as Physalia earlandi,
from which it differs by the greater
number of chambers, the initial spiral
stage and the small proloculus. Since this
species is reported in the literature with
246 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Rectocibicidella
Ramulina globulifera Rectocibicidella robertsi
Test free, branching, consisting of globular Test attached, elongate, compressed, early
Pyrulina chambers connected by stoloniferous stage trochospirally coiled and attached by
Pyrulina angusta tubes; wall hispid; aperture at the end of the spiral side, later uncoiling, becoming
Test fusiform, circular in section, with tubular projections, often several to a biserial and finally uniserial; sutures
obtuse extremities, somewhat pointed at single chamber. curved and limbate in the early stage,
the initial end; usually composed of about later slightly depressed, periphery acute to
Bay of Prony, 20 m. carinate, peripheral outline becoming
four embracing, elongate chambers; Systematics p. 291.
chambers added in planes about 120° lobulate in the adult; wall coarsely perfo-
apart in the early stage, later biserial; rate; aperture terminal, an ovate slit
sutures oblique, curved, flush; wall thin bordered by a lip.
and transparent, finely perforate; aperture Northern shelf, 200 m.
terminal, radiate. Systematics p. 315.
Southwestern lagoon, 40 m.
Systematics p. 291.
248 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
Siphoninoides laevigatus
Test subglobular, irregularly trochospiral
but chamber arrangement not discernable,
Sigmoidella Sigmomorphina chambers enlarging rapidly; wall thin in
Sigmoidella elegantissima Sigmomorphina the early stage, later much thickened and
Test large and strongly compressed, with cf. S. basistriata coarsely perforate, surface irregularly
a sigmoidal cross-sectional shape; cham- Some specimens correspond to the smooth; aperture elevated surrounded by
bers on one side of the test completely description of ZHENG (1979): “test com- a circular thickened rim, rounded, and
embracing the earlier ones, on the other pressed, oval to oblong, basal end nar- filled with a concave plate that has a
side partially evolute, giving the test an rowly rounded, apertural end slightly single small central pore.
asymmetric appearance; sutures flush or obtuse, periphery narrowly rounded; Bay of Prony, 20 m.
slightly depressed; wall smooth; aperture chambers long, thickest in the middle Systematics p. 313.
terminal, radiate. portion, thinning towards the periphery,
Northern shelf, 200 m. early ones with ridge-like periphery;
Systematics p. 291. chambers arranged in a clokwisely sig-
moid series, the early ones reaching the
base of the test, the later ones gradually
removed from the base; sutures distinct,
slightly depressed; wall translucent, the
basal half of the chambers with very fine
longitudinal interrupted costae; aperture
radiate”. However, in other specimens,
costae are stronger and may reach the
aperture.
Northern shelf, 600 m.
Systematics p. 291.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 249
Sphaerogypsina Sporadotrema
Sphaerogypsina globula Sporadotrema cylindrica
Test large, up to 2 mm in diameter, Test attached, very large, early stage
normally free; spherical to somewhat planispirally coiled, later chambers spi-
irregular, constructed of numerous layers raling upward to form a large cylindrical
of small, polygonal and closely packed upright structure with short terminal
chambers; neighboring chambers branches; 4-6 chambers per whorl, irreg-
belonging respectively to the ultimate ular in size and slightly inflated; sutures
and penultimate layers are out of level for depressed; color orange or yellowish; wall
half their height in radial direction, with thick with irregularly scattered coarse
respect to each other; upper surface of the perforations that result from outward
chambers coarsely perforate, and septa fusion of the finer pores at the inner wall
thick, elevated and imperforate; apertures surface; apertures multiple, two arches
multiple consisting of small rounded bordered by narrow lips on the umbilical
openings or small slits at the base of the wall of each chamber, and additional
free chamber walls. openings produced on tubes with everted
Northern shelf, 200 m. rims around the umbilical depression.
Systematics p. 317. Attached on algae, 125 m.
Systematics p. 318.
5 mm
Unidentified species
Test cylindrical with initial end rounded
and apertural end irregular; only one
chamber visible; test finely perforated,
surface smooth; aperture terminal, a set
of irregularly placed small rounded
Sphaeroidina openings. In its morphology, this species
Sphaeroidina bulloides resembles Duplella neobotelliformis
(McCulloch) as it is illustrated by
Test subglobular composed of a few LOEBLICH & TAPPAN (1994), but it differs by
chambers arranged in a relatively regular the characteristics of the aperture as
spiral; chambers hemispherical and described by MCCULLOCH (1977).
strongly embracing, centered over the
preceding aperture; wall very finely Northern shelf, 600 m.
perforate, surface smooth; aperture a
crescentic opening near the base of the
last-formed chamber, commonly above
the junction of earlier chambers, bor-
dered by a narrow lip, and with a simple
flaplike tooth.
Northern shelf, 600 m.
Systematics p. 311.
Systematics
Suprageneric classification based on LOEBLICH & TAPPAN (1992), except for agglutinated foraminifera (Textulariia) classified following
KAMINSKI (2004).
+ indicates new species for New Caledonia (not reported in the 2007 compendium).
* indicates species reported before, but not found to be included in this work.
1988 Spiroloculina communis Cushman & Todd - Haig, p. 234; pl. 10, figs 11-13.
2009 Spiroloculina subimpressa Parr, - Parker, p. 350, figs 254a-k.
+Spiroloculina sp. 1 - p. 135
+Spiroloculina sp. 2 - p. 135
Family Hauerinidae Schwager, 1876
Subfamily Hauerininae Schwager, 1876
Hauerina d’Orbigny, 1839
*Hauerina bradyi Cushman
Hauerina diversa Cushman - p. 108
1946 Hauerina diversa - Cushman, p. 11; pl. 2, figs 16-19.
1992a Hauerina diversa Cushman - Hatta & Ujiié, p. 65; pl. 6, figs 3a-b.
1993 Hauerina diversa Cushman - Hottinger et al., p. 50; pl. 36, figs 1-7.
+Hauerina earlandi Rasheed - p. 108
1971 Hauerina earlandi - Rasheed, p. 54; pl. 16, fig. 7.
1988 Miliola earlandi (Rasheed) - Haig, p. 220; pl. 2, figs 8-9.
2009 Hauerina earlandi Rasheed - Parker, p. 107, figs 74a-k.
+Hauerina fragilissima (Brady) - p. 108
1884 Spiroloculina fragilissima - Brady, p. 149; pl. 9, figs 12-14.
1988 Hauerina fragilissima (Brady) - Haig, p. 220; pl. 2, figs 3, 4.
1994 Parahauerinoides fragilissima (Brady) - Loeblich & Tappan, p. 51; pl. 87, figs 1-5.
2009 Hauerina fragilissima (Brady) - Parker, p. 107, figs 75a-g.
*Hauerina ornatissima (Karrer)
Hauerina pacifica Cushman - p. 108
1917 Hauerina pacifica - Cushman, p. 64; pl. 21, figs 2a-c.
1987 Hauerina pacifica Cushman - Baccaert, p. 145, 146; pl. 63, figs 4-6.
1988 Hauerina pacifica Cushman - Haig, p. 220; pl. 2, figs 5-7.
2009 Hauerina pacifica Cushman - Parker, p. 109, figs 76a-c; 77a-m; 78a-j.
Quinqueloculina d’Orbigny, 1826
*Quinqueloculina cf. adiazeta
Quinqueloculina agglutinans d’Orbigny [Siphonaperta agglutinans] - p. 119
1839a Quinqueloculina agglutinans - d’Orbigny, p. 195; pl. 12, figs 11-13.
1994 Agglutinella agglutinans (d’Orbigny) - Loeblich & Tappan, p. 44; pl. 70, figs 1-9.
Quinqueloculina arenata Said [Siphonaperta anguina arenata; Triloculina sabulosa Collins] - p. 119
1949 Quinqueloculina anguina Terquem var. arenata - Said, p. 9; pl. 1, fig. 25.
1988 Quinqueloculina arenata Said - Haig, p. 233; pl. 4, figs 15-17.
1994 Agglutinella arenata (Said) - Loeblich & Tappan, p. 45; pl. 69, figs 6-11; pl. 70, figs 10- 15; pl. 74,
figs 10-13.
2009 Quinqueloculina arenata Said - Parker, p. 179, figs 126a-j; 127a-h; 128a-i.
+Quinqueloculina auberiana d’Orbigny [Quinqueloculina lamarckiana] - p. 119
1839a Quinqueloculina auberiana - d’Orbigny, p. 193; pl. 12, figs 1-3.
1999 Quinqueloculina auberiana d’Orbigny - Hayward et al., p. 100; pl. 4, figs 13-14.
Quinqueloculina barnardi Rasheed - p. 119
1971 Quinqueloculina barnardi - Rasheed, p. 26, 27; pl. 2, fig. 1.
1988 Quinqueloculina barnardi Rasheed - Haig, p. 233; pl. 4, figs 18-20.
2009 Quinqueloculina barnardi Rasheed - Parker, p. 184, figs 129a-f; 130a-k.
Quinqueloculina bassensis (Parr) [Triloculina bassensis - Affinetrina quadrilateralis] - p. 119
1945 Triloculina bassensis - Parr, p. 198; pl. 8, figs 7a-c.
1987 Triloculina irregularis (d’Orbigny) - Baccaert, p. 126; pl. 57, fig. 1.
1993 Affinetrina cf. A. quadrilateralis (d’Orbigny) - Hottinger et al., p. 47; pl. 28, figs 9-15; pl. 29,
figs 1-4.
2009 Quinqueloculina bassensis (Parr) - Parker, p. 184, figs 131a-g.
*Quinqueloculina cf. berthelotiana d’Orbigny
Quinqueloculina bicarinata d’Orbigny - p. 120
1826 Quinqueloculina bicarinata d’Orbigny, p. 302, no 35.
1878 Quinqueloculina bicarinata d’Orbigny - Terquem; pl. 5, figs 10a-c.
1921 Quinqueloculina bicarinata d’Orbigny - Cushman, p. 428; pl. 86, figs 2-3.
1988 Quinqueloculina bicarinata d’Orbigny - Haig, p. 233; pl. 4, figs 27-28; pl. 5, figs 1-5.
2009 Quinqueloculina pseudolamarckiana n.sp. - Margerel http://147.94.111.32/Collection/forams-
index.php?
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 271
1993 Pseudomassilina australis (Cushman) - Hottinger et al., p. 53; pl. 41, figs 3-11.
1994 Pseudomassilina australis (Cushman) - Loeblich & Tappan, p. 53; pl. 91, figs 1-3.
Pseudomassilina macilenta (Brady) - p. 115
1884 Miliolina macilenta - Brady, p. 167; pl. 7, figs 5-6.
1988 Pseudomassilina macilenta (Brady) - Haig, p. 228; pl. 3, figs 21, 24.
1994 Pseudomassilina macilenta (Brady) - Loeblich & Tappan, p. 53; pl. 90, figs 10-13; not pl. 90, figs
5-9.
2009 Pseudomassilina macilenta (Brady) - Parker, p. 168, figs 118a-j.
Pseudomassilina pacificiensis Cushman - p. 115
1924 Pseudomassilina pacificiensis - Cushman, p. 66; pl. 24, figs 1, 2.
1988 Pseudomassilina pacificiensis Cushman - Haig, p. 228; pl. 3, fig. 25.
1993 Pseudomassilina pacificiensis Cushman - Hottinger et al., p. 54; pl. 42, figs 1-4.
+Pseudomassilina robusta Lacroix - p. 115
1938 Pseudomassilina oblonga Lacroix, var. robusta - Lacroix, p. 5, text fig. 3.
1988 Pseudomassilina robusta Lacroix - Haig, p. 233; pl. 3, figs 26, 27.
1994 Pseudomassilina robusta Lacroix - Loeblich & Tappan p. 53; pl. 90, figs 1-4.
+Pseudomassilina? sp. 1 - p. 116
Pseudotriloculina Cherif, 1970
+Pseudotriloculina cf. P. chrysostoma (Chapman) - p. 116
1909 Miliolina chrysostoma - Chapman, p. 322; pl. 13, figs 8-10; pl. 14, figs 1, 4.
1957 Triloculina chrysostoma (Chapman) - Vella; pl. 5; figs 97-99.
1999 Triloculina chrysostoma (Chapman) - Hayward et al., p. 105; pl. 5, figs 27-28.
Pseudotriloculina linneiana (d’Orbigny) [Triloculina planciana d’Orbigny] - p. 116
1839a Triloculina linneiana - d’Orbigny, p. 172; pl. 9, figs 11-13.
1929 Triloculina linneiana d’Orbigny - Cushman, p. 61; pl. 16, figs 1, 2.
1987 Triloculina linneiana d’Orbigny - Baccaert, p. 128; pl. 57, figs 3, 4.
Pseudotriloculina subgranulata (Cushman) - p. 116
1918 Triloculina subgranulata Cushman, p. 290; pl. 96, fig. 4.
1987 Triloculina linneiana d’Orbigny var. subgrabulata Cushman - Baccaert, p. 129, 130; pl. 58,
figs 1, 2.
1988 Quinqueloculina eamsii (Rasheed) - Haig, p. 233; pl. 11, figs 1-4.
2009 Quinqueloculina subgranulata (Cushman) - Parker, p. 259, figs 187a-j.
Pyrgo Defrance, 1824
+Pyrgo anomala (Schlumberger) - p. 116
1891 Biloculina anomala - Schlumberger, p. 569; pl. 11, figs 84-86; pl. 12, fig. 101.
1988 Pyrgo anomala (Schlumberger) - Zheng, p. 222; pl. 11, fig. 1; pl. 27, fig. 3.
1994 Nummulopyrgo anomala (Schlumberger) - Loeblich & Tappan, p. 42; pl. 91, figs 4-10.
1999 Pyrgo anomala (Schlumberger) - Hayward et al., p. 97; pl. 4, figs 1-2.
+Pyrgo comata (Brady) - p. 116
1881 Biloculina comata - Brady, p. 45.
1884 Biloculina comata Brady - Brady, p. 144; pl. 3, figs 9a-b.
1917 Biloculina comata Brady - Cushman, p. 81; pl. 34, fig. 1.
1999 Pyrgo comata (Brady) - Hayward et al., p. 98; pl. 4, figs 3-4.
Pyrgo denticulata (Brady) - p. 117
1884 Biloculina ringens (Lamarck) var. denticulata - Brady, p. 143; pl. 3, figs 4-5.
1987 Pyrgo denticulata (Brady) - Baccaert, p. 113; pl. 51, figs 5-6; pl. 52, fig. 1.
1988 Pyrgo denticulata (Brady) - Haig, p. 233; pl. 3, fig. 28, ?29.
2009 Pyrgo denticulata (Brady) - Parker, p. 168, figs 119a-h.
Pyrgo depressa (d’Orbigny) - p. 117
1826 Biloculina depressa - d’Orbigny, p. 298.
1884 Biloculina depressa d’Orbigny - Brady, p. 145; pl. 2, figs 12, 16-17.
1988 Pyrgo depressa (d’Orbigny) - Zheng, p. 225; pl. 11, figs 4-5; pl. 31, fig. 13.
1994 Biloculinella depressa (d’Orbigny) - Loeblich & Tappan, p. 51; pl. 86, figs 1-4.
Pyrgo inornata (d’Orbigny) - p. 117
1846 Biloculina inornata - d’Orbigny, p. 266; pl. 16, figs 7-9.
1999 Pyrgo anomala (Schlumberger) - Hayward et al., p. 97; pl. 4, figs 1-2.
2010 Pyrgo inornata (d’Orbigny) - Hayward et al., p. 151; pl. 7, figs 15-19.
*Pyrgo lucernula (Schwager)
*Pyrgo murrhyna (Schwager)
Pyrgo oblonga (d’Orbigny) - p. 117
1839a Biloculina oblonga - d’Orbigny, p. 163; pl. 8, figs 21-23.
1993 Pyrgo oblonga (d’Orbigny) - Hottinger et al., p. 57; pl. 50, figs 1-6.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 277
1992b Patellina corrugata - Hatta & Ujiié, p. 164; pl. 20, fig. 5.
1995 Patellina corrugata Williamson - Yassini & Jones, p. 163; figs 743-745.
+Patellina elaborata McCulloch, 1977 - p. 207
1977 Patellina corrugata var. elaborata - McCulloch, p. 279; pl. 112, figs 1-4.
+Patellina cf. P. formosa Heron-Allen & Earland, 1932 - p. 207
1932b Patellina corrugata Williamson var. Formosa - Heron-Allen & Earland, p. 406; pl. 13, figs 23-25.
1994 Patellina corrugata Williamson - Loeblich & Tappan, p. 36; pl. 55, figs 1-9.
1995 Heteropatellina frustratiformis McCulloch - Yassini & Jones, p. 997; fig. 837.
2009 Patellina cf. P. formosa Heron-Allen & Earland - Parker p. 70; figs 51a-h.
+Patellina sp. 1 - p. 207
1995 Fissurina clathrata (Brady) - Yassini & Jones, p. 122, figs 454-457.
2009 Buchnerina clathrata (Brady) Margerel, http://147.94.111.32/Collection/forams-index.php?
+Cerebrina conformata (McCulloch, 1977) - p. 142
1977 Fissurina conformata - McCulloch, p. 97; pl. 62, fig. 9.
1994 Cerebrina conformata (McCulloch) - Loeblich & Tappan, p. 75; pl. 136, figs 3-4.
Cerebrina lacunata (Burrows & Holland in Jones 1895) [Buchnerina] - p. 142
1895 Lagena lacunata - Burrows & Holland in Jones, p. 205; pl. 7, figs 12a, b.
1994 Cerebrina lacunata (Burrows & Holland) - Loeblich & Tappan, p. 76; pl. 135, figs 8-15.
2009 Cerebrina lacunata (Burrows & Holland in Jones 1865) - Parker, p. 395; figs 284a-l; 285a-i.
2009 Buchnerina lacunata (Brady) Margerel, http://147.94.111.32/Collection/forams-index.php?
+Cerebrina neocastrensis (McCulloch, 1977) - p. 142
1977 Fissurina neocastrensis - McCulloch, p. 117; pl. 61, fig. 20.
1994 Cerebrina neocastrensis (McCulloch) - Loeblich & Tappan, p. 76; pl. 135, figs 19-20.
+Cerebrina pilasensis (McCulloch, 1977) - p. 142
1977 Fissurina pilasensis - McCulloch, p. 123; pl. 64, fig. 4.
+Cerebrina undulaticostata (McCulloch, 1977) - p. 142
1977 Fissurina undulaticostata - McCulloch, p. 134; pl. 63, fig. 24.
+Cerebrina sp. 1 - p. 143
+Cerebrina sp. 2 - p. 143
Hyalinonetrion Patterson & Richardson, 1987
+Hyalinonetrion distomapolita (Parker & Jones, 1865) - p. 151
1865 Lagena sulcata (Walker & Jacob) var. distomapolita - Parker & Jones, p. 357; pl. 13, fig. 21.
1995 Procerolagena distomapolita (Parker & Jones) - Yassini & Jones, p. 109, fig. 876.
+Hyalinonetrion elongata (Ehrenberg, 1844) - p. 151
1844 Miliolida elongata - Ehrenberg, p. 274.
1995 Procerolagena elongata (Ehrenberg) - Yassini & Jones, p. 109, figs 271-273.
Hyalinonetrion gracillima (Seguenza, 1862) - p. 152
1862 Amphorina gracillima - Seguenza, p. 51; pl. 1, fig. 37.
1995 Procerolagena gracillima (Seguenza) - Yassini & Jones, p. 109, figs 271-273.
2009 Hyalinonetrion gracillima (Seguenza) - Margerel, http://147.94.111.32/Collection/forams-index.php?
Lagena Walker & Jacob, 1798
+Lagena fenestrata Yassini & Jones, 1995 - p. 152
1995 Lagena fenestrata - Yassini & Jones, p. 104, fig. 344.
*Lagena gracilis Williamson
*Lagena hispida (Reuss)
+Lagena cf. L. laevicostata Cushman & Gray, 1946 - p. 152
1946 Lagena sulcata (Walker & Jacob) var. laevicostata - Cushman & Gray, p. 68; pl. 12, figs 13-14.
1950 Lagena sulcata var. laevicostata Cushman & Gray - Cushman & McCulloch, p. 361; pl. 48, figs 8-10.
*Lagena laevis (Montagu)
+Lagena paucistriata Yassini & Jones, 1995 - p. 152
1995 Lagena striata paucistriata - Yassini & Jones, p. 106; figs 323-325.
*Lagena perlucida (Montagu)
+Lagena cf. L. pustulostriatula Albani & Yassini, 1989 - p. 152
1989 Lagena pustulostriatula - Albani & Yassini, p. 379; pl., figs 2q-r.
1995 Lagena pustulostriatula Albani & Yassini - Yassini & Jones, p. 106, figs 328-329.
+Lagena spicata Cushman & McCulloch, 1950 - p. 152
1950 Lagena sulcata var. spicata - Cushman & McCulloch, p. 360; pl. 48, figs 3, 7.
1995 Lagena striata paucistriata - Yassini & Jones, p. 106, 107, figs 323-325.
1999 Lagena spicata Cushman & McCulloch - Hayward et al., p. 116; pl. 7, figs 4-5.
2009 Lagena spicata (Cushman & McCulloch) - Margerel, http://147.94.111.32/Collection/forams-index.php?
*Lagena striata d’Orbigny
Lagena strumosa Reuss, 1858 - p. 153
1858 Lagena striata strumosa - Reuss, p. 434.
1995 Lagena striata strumosa Reuss - Yassini & Jones, p. 107, figs 321-322, 326-327, 330-331.
+Lagena tortilis Egger, 1893 - p. 153
1893 Lagena tortilis - Egger, p. 329; pl. 10, figs 61-63.
1901b Lagena striata d’Orbigny var. tortilis Egger - Millett, p. 487; pl. 8, fig. 4.
+Lagena sp. 1 - p. 153
Procerolagena Puri, 1954
+Procerolagena cylindrocostata Albani & Yassini, 1989 - p. 159
1989 Procerolagena cylindrocostata - Albani & Yassini, p. 381, fig. 3 D.
1995 Procerolagena cylindrocostata Albani & Yassini - Yassini & Jones, p. 108, figs 289-291.
290 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
1930 Guttulina regina (Brady, Parker & Jones) - Cushman & Ozawa, p. 34; pl. 6, figs 1, 2.
2009 Guttulina regina (Brady, Parker & Jones) - Parker, p. 405; figs 292a-k.
+Guttulina yamazakii Cushman & Ozawa, 1930 - p. 240
1930 Guttulina yamazakii - Cushman & Ozawa, p. 40; pl. 8, figs 3-4.
1994 Guttulina yamazaki Cushman & Ozawa - Loeblich & Tappan, p. 82; pl. 148, figs 1-3.
+Guttulina sp. 1 - p. 240
Krebsina McCulloch, 1981
Krebsina subtenuis (Cushman, 1936) [Bolivina subtenuis] - p. 174
1884 Bolivina tenuis - Brady, p. 419; pl. 52, fig. 29.
1936 Bolivina subtenuis - Cushman, p. 57; pl. 8, fig. 10.
1994 Krebsina subtenuis (Cushman) - Loeblich & Tappan, p. 82; pl. 146, figs 12-16.
Polymorphina d’Orbigny, 1826
+Polymorphina cf. P. diffusa Jones & Chapman, 1896 - p. 247
1884 Polymorphina lactea (Walker & Jacob), fistulose form - Brady, p. 560; pl. 73, fig. 14.
1896 Polymorphina spp. var. diffusa - Jones & Chapman, p. 505; figs 26-29.
1907 Polymorphina lactea var. diffusa Jones & Chapman - Chapman, p. 131; pl. 10, fig. 1.
1913b Polymorphina lactea (Walker & Jacob) var. diffusa Jones & Chapman - Cushman, p. 84; pl. 41,
fig. 8.
Polymorphinella Cushman & Hanzawa, 1936
+Polymorphinella pacifica Cushman & Hanzawa, 1936 - p. 168
1884 Cristellaria schloenbachi anomalous specimen - Brady, p. 539; pl. 69, fig. 8.
1936 Polymorphinella pacifica - Cushman & Hanzawa, p. 47.
1954 Polymorphinella pacifica Cushman & Hanzawa - Cushman, Todd & Post, p. 344; pl. 86, fig. 23-24.
1994 Marginulina subcrassa Schwager - Loeblich & Tappan, p. 74; pl. 129, figs 17-18.
Pseudopolymorphina Cushman & Ozawa, 1928
*Pseudopolymorphina ligua (Roemer, 1838)
*Pseudopolymorphina ovalis Cushman et Ozawa
Pseudopolymorphina sp. 1 - p. 176
1884 Polymorphina compressa d’Orbigny - Brady, p. 564; pl. 72, figs 9-11.
1960 Pseudopolymorphina ligua (Roemer) - Barker; pl. 72, figs 9-11.
Pyrulina d’Orbigny, 1839
+Pyrulina angusta (Egger, 1857) - p. 247
1857 Polymorphina (Globulina) angusta - Egger, p. 290; pl. 13, figs 13-15.
1884 Polymorphina (Globulina) angusta Egger - Brady, p. 563; pl. 72, figs 1-3.
1913b Polymorphina (Globulina) angusta Egger - Cushman, p. 86; pl. 39, fig. 6.
1990 Pyrulina angusta (Egger) - Ujiié, p. 21; pl. 6, figs 10-11.
Sigmoidella Cushman & Ozawa, 1928
+Sigmoidella elegantissima (Parker & Jones, in Brady Parker & Jones, 1870) - p. 248
1865 Polymorphina elegantissima - Parker & Jones, p. 438. (nom. nud.)
1870 Polymorphina elegantissima - Parker & Jones in Brady, Parker & Jones, p. 231; pl. 40, figs 15a-c.
1930 Sigmoidella elegantissima (Parker & Jones) - Cushman & Ozawa, p. 140; pl. 39, fig. 1.
1994 Sigmoidella elegantissima (Parker & Jones) - Loeblich & Tappan, p. 83; pl. 148, figs 4-12.
2009 Sigmoidella elegantissima (Parker & Jones) - Parker, p. 422; figs 305a-g.
+Sigmoidella pacifica Cushman & Ozawa, 1928 - p. 248
1928 Sigmoidella (Sigmoidina) pacifica - Cushman & Ozawa, p. 19; pl. 2, fig. 13.
1960 Guttulina (Sigmoidella) pacifica (Cushman & Ozawa) - Barker, pl. 72; figs 14-15.
1993 Sigmoidella cf. S. pacifica Cushman & Ozawa - Hottinger et al., p. 80; pl. 91, figs 16-18.
1994 Sigmoidella pacifica Cushman & Ozawa - Loeblich & Tappan, p. 84; pl. 149, figs 1-9.
Sigmomorphina Cushman & Ozawa, 1928
+Sigmomorphina cf. S. basistriata Zheng, 1979 - p. 248
1979 Sigmomorphina basistriata - Zheng, p. 212; pl. 11, figs 5-6.
Subfamily Ramulininae Brady, 1884
Ramulina T.R. Jones, in Wright, 1875
+Ramulina? confosa Loeblich & Tappan, 1994 - p. 247
1994 Ramulina confosa - Loeblich & Tappan, p. 84; pl. 150, figs 8-13.
Ramulina globulifera Brady, 1879 - p. 247
1879 Ramulina globulifera - Brady, p. 272; pl. 8, figs 32-33.
1913b Ramulina globulifera Brady - Cushman, p. 110; pl. 39, fig. 1.
1992b Ramulina globulifera Brady - Hatta & Ujiié, p. 167; pl. 22, fig. 6.
1994 Ramulina globulifera Brady - Loeblich & Tappan, p. 84; pl. 149, fig. 17.
+Ramulina vanandeli Loeblich & Tappan, 1994 - p. 247
1994 Ramulina vanandeli - Loeblich & Tappan, p. 85; pl. 150, figs 1-7.
292 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
2000 Loxostomina costatapertusa Loeblich & Tappan - Revets, p. 371; pl. 3, fig. 10.
2009 Loxostomina costatapertusa Loeblich & Tappan - Parker, p. 456; figs 3328a-c.
Loxostomina costulata (Cushman, 1922) - p. 175
1922a Bolivina limbata Brady var. costulata - Cushman, p. 26; pl. 3, fig. 8.
1994 Loxostomina costulata (Cushman) - Loeblich & Tappan, p. 119; pl. 232, figs 12-16.
Loxostomina limbata (Brady, 1884) - p. 175
1884 Bolivina limbata - Brady, p. 419; pl. 52, figs 26-28,
1987 Loxostomina (?) limbatum (Brady) - Baccaert, p. 264-265; pl. 106, fig. 11.
1994 Loxostomina limbata (Brady) - Loeblich & Tappan, p. 119; pl. 233, figs 1-8.
2009 Loxostomina limbata (Brady) - Parker, p. 456; figs 329a-k, 330a-h.
+Loxostomina sp. 1 - p. 175
Pseudobrizalina Zweig-Strykowski & Reiss, 1976
+Pseudobrizalina lobata (Brady 1884) - p. 176
1884 Bolivina lobata - Brady, p. 425; pl. 53, figs 22-23.
2002 Bolivina lobata Brady - Suresh-Gandhy et al., p. 56; pl. 2, fig. 5.
Rectobolivina Cushman, 1927
*Rectobolivina dimorpha (Parker & Jones)
Sagrinella Saidova, 1975
Sagrinella convallaria (Millett, 1900) - p. 178
1900b Bolivina convallaria - Millett, p. 544; pl. 4, figs 6a-b.
1958 Loxostomum convallarium (Millett) - Collins, p. 395; pl. 5, fig. 2.
1993 Sagrinella convallaria (Millett) - Hottinger et al., p. 98; pl. 122, figs 8-11.
Sagrinella durrandii (Millett, 1900) [Euloxostomum durrandii] - p. 178
1900b Bolivina durrandii - Millett, p. 544; pl. 4, figs 7a-b.
1992b Loxostomina durrandi - Hatta & Ujiié, p. 174; pl. 26, fig. 10.
1994 Sagrinella durrandii (Millett) - Loeblich & Tappan, p. 120; pl. 236, figs 11-13.
Sagrinella strigosa Brady, 1884 - p. 178
1884 Bolivina lobata var. strigosa - Brady, p. 425; pl. 113, fig. 7.
1911 Bolivina (Bifarina) strigosa (Brady) - Cushman, p. 49; fig. 80.
1975 Sagrinella strigosa (Brady) - Saidova, p. 310.
1994 Siphouvigerina strigosa (Brady) - Loeblich & Tappan, p. 127; pl. 247, fig. 12.
Subfamily Tubulogenerininae Saidova, 1981
Allassoida Loeblich & Tappan 1994
Allassoida virgula (Brady, 1879) [Siphogenerina virgula] - p. 162
1879 Sagrina virgula - Brady, p. 275; pl. 8, figs 19-21.
1924 Siphogenerina virgula (Brady) - Cushman, p. 29; pl. 8, figs 3-4.
1994 Allassoida virgula (Brady) - Loeblich & Tappan, p. 121; pl. 238, figs 1-11.
2001 Allassoida virgula (Brady) - Szarek, p. 127; pl. 17, fig. 14.
Sagrina d’Orbigny, 1839
Sagrina jugosa (Brady, 1884) [Patellinella jugosa] - p. 178
1884 Textularia jugosa - Brady, p. 358; pl. 42, fig. 7.
1994 Sagrina jugosa (Brady) - Loeblich & Tappan, p. 122; pl. 237, figs 12-17.
2001 Sagrina jugosa (Brady) - Szareck, p. 128; pl. 17, fig. 15.
+Sagrina zanzibarica (Cushman, 1936) - p. 178
1936 Bolivina zanzibarica - Cushman, p. 58; p1. 8, fig. 12.
1958 Bolivina zanzibarica Cushman - Collins, p. 395.
1994 Sagrina zanzibarica (Cushman) - Loeblich & Tappan, p. 122; pl. 238, figs 12-17.
Siphogenerina Schlumberger, in Milne-Edwards, 1882
Siphogenerina columellaris (Brady, 1881) - p. 169
1881 Sagrina columellaris - Brady, p. 64.
1884 Sagrina columellaris Brady - Brady, p. 581; pl. 75, figs 15-17.
1960 Rectobolivina columellaris (Brady) - Barker, p. 156; pl. 75, figs 15-17.
1994 Siphogenerina columellaris (Brady) - Jones, p. 87; pl. 75, figs 15-17.
+Siphogenerina pacifica Cushman, 1926 - p. 169
1926a Siphogenerina dimorpha (Parker & Jones) var. pacifica - Cushman, p. 13; pl. 2, fig. 9.
1960 Rectobolivina dimorpha (Parker & Jones) var. pacifica (Cushman) - Barker, p. 158; pl. 76, figs 1-3.
1994 Siphogenerina pacifica (Cushman) - Loeblich & Tappan, p. 123; pl. 241, figs 1-9.
Siphogenerina raphana (Parker & Jones, 1865) - p. 169
1865 Uvigerina (Sagrina) raphanus - Parker & Jones, p. 364; pl. 18, figs 16-17.
1884 Sagrina raphanus (Parker & Jones) - Brady, p. 585; pl. 75, figs 21-22.
1992b Rectobolivina raphana (Parker & Jones) - Hatta & Ujiié, p. 174; pl. 26, figs 11-12.
1999 Siphogenerina raphana (Parker & Jones) - Hayward et al., p. 130; pl. 9, fig. 4.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 303
1987 Reussella “simplex” (Cushman) - Baccaert, p. 190; pl. 75, figs 3-5.
2009 Reussella pacifica Cushman & McCulloch - Parker, p. 463; figs 333a-c.
+Reussella pulchra Cushman, 1945 - p. 182
1945 Reussella pulchra - Cushman, p. 34; pl. 6, figs 11-12.
1966 Reussella pulchra Cushman - Todd, pl. 18, fig. 6.
1980 Reussella pulchra Cushman - Zheng, p. 166; pl. 4, figs 12-13.
+Reussella spinulosa (Reuss, 1850) - p. 182
1850 Verneuilina spinulosa - Reuss, p. 374; pl. 47, fig. 12.
1884 Verneuilina spinulosa Reuss - Brady, p. 384; pl. 47, figs 2-3.
1942 Reussella spinulosa (Reuss) - Cushman, p. 40; pl. 11, figs 5-8.
Valvobifarina Hofker, 1951
*Valvobifarina mackinnoni (Millett)
Family Trimosinidae Saidova, 1981
Mimosina Millett, 1900
Mimosina affinis Millett, 1900 - p. 180
1900b Mimosina affinis - Millett, p. 548; p1. 4, fig. 11.
1993 Mimosina affinis Millett - Hottinger et al., p. 104; pl. 133, figs 9-12; pl. 134, figs 1-3.
Mimosina echinata Heron-Allen & Earland, 1915 - p. 180
1915 Mimosina echinata - Heron-Allen & Earland, p. 651; p1. 50, figs 12-18.
1987 Mimosina echinata Heron-Allen & Earland - Baccaert; pl. 75, figs 1-2.
1994 Mimosina echinata Heron-Allen & Earland - Loeblich & Tappan, p. 129; pl. 255, figs 1-2.
Mimosina histrix Millett, 1900 - p. 181
1900b Mimosina hystrix - Millett, p. 549; p1. 4, figs 14, 15.
1927 Mimosina hystrix Millett - Cushman, p. 64.
1991 Mimosina histrix Millett - Revets, p. 6; pl. 2, figs 5-11.
*Mimosina pacifica Cushman
+Mimosina sp. 1 - p. 181
Quirimbatina Debenay n.gen.
+Quirimbatina rimosa (Heron-Allen & Earland, 1915) - p. 177
1915 Mimosina rimosa - Heron-Allen & Earland, p. 650; pl. 50, figs 5-11.
1958 Mimosina rimosa Heron-Allen & Earland - Collins, p. 391; pl. 4, fig. 11.
Trimosina Cushman, 1927
+Trimosina milletti Cushman, 1927 - p. 183
1900b Mimosina spinulosa var. - Millett, p. 548; p1. 4, fig. 13.
1927 Trimosina milletti - Cushman, p. 64, p1. 13, fig. 20.
1991 Trimosina milletti Cushman - Revets, p. 4; pl. 1, figs 8-9.
+Trimosina orientalis Cushman, 1933 - p. 183
1933b Trimosina orientalis - Cushman, p. 78; pl. 8, fig. 4.
1942 Trimosina orientalis Cushman - Cushman, p. 43; pl. 12, figs 1-5.
Family Pavoninidae Eimer & Fickert, 1899
Pavonina d’Orbigny, 1826
Pavonina flabelliformis d’Orbigny, 1826 - p. 168
1826 Pavonina flabelliformis - d’Orbigny, p. 358; pl. 42, fig. 7.
1991 Pavonina flabelliformis d’Orbigny - Revets, p. 8; pl. 3, figs 4-9.
1994 Pavonina flabelliformis d’Orbigny - Loeblich & Tappan, p. 130; pl. 255, figs 3-6.
2009 Pavonina flabelliformis d’Orbigny - Parker, p. 462; figs 332a-c.
Family Millettiidae Saidova, 1981
Millettia Schubert, 1911
Millettia limbata (Brady, 1884) - p. 243
1884 Sagrina limbata - Brady, p. 586; pl. 113, fig. 14.
1992 Millettia limbata (Brady) - Revets, p. 40, pl. 2, figs 5-9.
1994 Millettia limbata (Brady) - Loeblich & Tappan, p. 130; pl. 255; figs 7-8.
+Millettia cf. M. tessellata (Brady, 1884) - p. 243
1884 Sagrina (?) tessellata - Brady, p. 585; pl. 76, figs 17-19.
1992 Millettia tessellata (Brady) - Revets, p. 38, pl. 1, figs 1-13.
1994 Millettia tessellata (Brady) - Loeblich & Tappan, p. 130; pl. 255; figs 9-15.
Superfamily Fursenkoinacea Loeblich & Tappan, 1961
Family Fursenkoinidae Loeblich & Tappan, 1961
Fursenkoina Loeblich & Tappan, 1961
Fursenkoina earlandi (cushman, 1936) [Stainforthia earlandi] - p. 174
1900a Virgulina schreibersiana Czjzek var. - Millett, p. 280; pl. 2, fig. 13.
1915 Virgulina schreibersiana Czjzek - Heron Allen & Earland, p. 642; pl. 49, figs 1-12.
306 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
1993 Millettiana milletti (Heron-Allen & Earland) - Hottinger et al., p. 120, pl. 160, figs 9-13.
2009 Millettiana milletti (Heron-Allen & Earland) - Parker, p. 640; figs 452a-k, 453a-g.
Family Victoriellidae Chapman & Crespin, 1930
Subfamily Carpenteriinae Saidova, 1981
Carpenteria Gray, 1858
+Carpenteria monticularis Carter, 1877 - p. 190
1877b Carpenteria monticularis - Carter, p. 211; pl. 13, figs 9-12.
1884 Carpenteria monticularis Carter - Brady, p. 677; pl. 98, figs 13-15; pl. 99, figs 1-5.
1994 Carpenteria monticularis Carter - Loeblich & Tappan, 1994, p. 153; pl. 391, fig. 5.
+Carpenteria cf. C. utricularis (Carter, 1876) - p. 190
1876 Polytrema utriculare - Carter, p. 210; pl. 13, figs 11-16.
1884 Carpenteria utricularis (Carter) - Brady, p. 678, pl. 99, figs 6-7; pl. 100, figs 1-4.
1921 Carpenteria utricularis (Carter) - Cushman, p. 360; pl. 73, figs 4-5.
1994 Carpenteria utricularis (Carter) - Loeblich & Tappan, 1994, p. 153; pl. 330, figs 4-12.
Subfamily Rupertuninae Loeblich & Tappan, 1961
Biarritzina Loeblich & Tappan, 1964
Biarritzina proteiformis (Goës, 1882) [Carpenteria proteiformis] - p. 235
1882 Carpenteria balaniformis var. proteiformis - Goës, p. 94; pl. 6, figs 208-214; pl. 7, figs 215-219.
1884 Carpenteria proteiformis Goës - Brady, p. 679, pl. 97, figs 8-14.
1992b Biarritzina proteiformis (Goës) - Hatta & Ujiié, p. 191; pl. 41, fig. 1.
1994 Biarritzina proteiformis (Goës) - Loeblich & Tappan, p. 153; pl. 331, figs 4-8.
Rupertina Loeblich & Tappan, 1961
+Rupertina pustulosa Hatta, 1992 - p. 213
1992b Rupertina pustulosa - Hatta in Hatta & Ujiié, p. 192; pl. 41, figs 2-4.
1994 Rupertina pustulosa Hatta - Loeblich & Tappan, p. 154; p1. 331, figs 2-4.
1999 Rupertina pustulosa Hatta - Hayward et al., p. 155; p1. 15, fig. 1.
Superfamily Acervulinacea Schultze, 1854
Family Acervulinidae Schultze, 1854
Acervulina Schultze, 1854
*Acervulina inhaerens Schultze, 1854
Acervulina mabahethi (Said, 1949) - p. 234
1949 Planorbulina mabahethi - Said, p. 44, pl. 4, fig. 26.
1993 Acervulina mabahethi (Said) - Hottinger et al., p. 122; pl. 165, figs 1-7; pl. 166, figs 1-8.
1994 Planorbulina mabahethi Said - Loeblich & Tappan, p. 152; pl. 323, figs 11-13.
2009 Acervulina mabahethi (Said) - Parker, p. 475; figs 341a-i; 342a-j; 343a-i.
Gypsina Carter, 1877
*Gypsina fimbriata (Chapman)
+Gypsina plana (Carter, 1876) - p. 240
1876 Polytrema planum - Carter, p. 211; pl. 13, figs 18-19.
1957 Acervulina inhaerens Scultze plana (Carter) - Hanzawa, p. 67; pl. 24, figs 2a-c.
1993 Gypsina plana (Carter) - Hottinger et al., p. 123; pl. 167, figs 1-12; pl. 168, figs 1-6.
Gypsina vesicularis (Parker & Jones, 1860) - p. 241
1860 Orbitolina vesicularis - Parker & Jones, p. 31.
1884 Gypsina vesicularis (Parker & Jones) - Brady, p. 718; pl. 101, figs 9-12.
1988 Gypsina vesicularis (Parker & Jones) - Loeblich & Tappan, p. 597; pl. 661, figs 1-6.
1994 Gypsina vesicularis (Parker & Jones) - Loeblich & Tappan, p. 154; pl. 333, figs 1-9; pl. 334, figs 1-3.
Planogypsina Bermúdez, 1952
Planogypsina acervalis (Brady, 1884) - p. 246
1884 Planorbulina acervalis - Brady, p. 657, pl. 92, fig. 4.
1987 Planorbulina acervalis Brady - Baccaert, p. 220; pl. 88, figs 1-3.
1993 Planogypsina acervalis (Brady) - Hottinger et al., p. 125; pl. 169, figs 1-9; pl. 170, figs 1-8.
2009 Planogypsina acervalis (Brady) - Parker, p. 697; figs 490a-d; 491a-i.
+Planogypsina squamiformis (Chapman, 1901) - p. 246
1901 Gypsina vesicularis (Parker & Jones) var. squamiformis - Chapman, p. 200; pl. 19, fig. 15.
1964 Planogypsina squamiformis (Chapman) - Loeblich & Tappan, p. C698; fig. 568.
1993 Planogypsina cf. P. squamiformis (Chapman) - Hottinger et al., p. 126; pl. 171, figs 1-9.
+Planogypsina? sp. 1 - p. 246
Sphaerogypsina Galloway, 1933
Sphaerogypsina globula (Reuss, 1848) - p. 249
1848 Ceriopora globulus - Reuss, p. 33; pl. 5, fig. 7.
1860 Orbitolina concava Lamarck var. vesicularis - Parker & Jones, p. 31, 38.
1993 Sphaerogypsina globulus (Reuss) - Hottinger et al., p. 128; pl. 173, figs 1-10.
2009 Sphaerogypsina globula (Reuss) - Parker, p. 736, figs 517a-j.
318 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
1993 Anomalinulla glabrata (Cushman) - Hottinger et al., p. 139; pl. 197, figs 6-11.
1995 Melonis asymmetrica - Yassini & Jones, p. 170; figs 915, 918, 919.
2009 Anomalinulla glabrata (Cushman) - Parker, p. 508; figs 361a-l.
Discanomalina Asano, 1951
+Discanomalina coronata (Parker & Jones, 1857) - p. 193
1857 Anomalina coronata - Parker & Jones, p. 294; pl. 10, figs 15-16.
1884 Anomalina coronata (Parker & Jones) - Brady, p. 675; pl. 97, figs 1-2.
1931a Anomalina coronata (Parker & Jones) - Cushman, p. 104; pl. 18, figs 3-4.
2010 Discanomalina coronata (Parker & Jones) - Hayward et al., p. 217; pl. 26, figs 7-8.
+Discanomalina semipunctata (Bailey, 1851) - p. 194
1851 Rotalina semipunctata - Bailey, p. 11; figs 17-19.
1931a Anomalina semipunctata (Bailey) - Cushman, p. 106; pl. 18, figs 1-2.
1994 Discanomalina semipunctata (Bailey) - Loeblich & Tappan, p. 163; pl. 361, figs 4-6.
Gyroidina d’Orbigny, 1826
*Gyroidina broeckhiana (Karrer, 1878)
Gyroidina lamarckiana (d’Orbigny, 1839) - p. 197
1839b Rotalina lamarckiana - d’Orbigny, p. 131; pl. 2, figs 13-15.
1965 Gyroidina lamarckiana (d’Orbigny) - Todd, p. 19; pl. 6, figs 3a-c.
Hansenisca Loeblich & Tappan, 1988
Hansenisca soldanii (d’Orbigny, 1826) [Gyroidina neosoldanii] - p. 197
1826 Rotalia soldanii - d’Orbigny, p. 276, no. 5.
1990 Gyroidinoides soldanii (d’Orbigny) - Ujiié, p. 45; pl. 25, figs 1-5.
1994 Hansenisca soldanii (d’Orbigny) - Loeblich & Tappan, p. 164; pl. 362, figs 8-10.
1996 Gyroidina soldanii (d’Orbigny) - Collins et al.; pl. 2, figs 9-10.
Hanzawaia Asano, 1944
+Hanzawaia grossepunctata (Earland, 1934) - p. 198
1934 Cibicides grossepunctatus - Earland, p. 184; pl. 8, figs 39-41.
1994 Hanzawaia grossepunctata (Earland) - Loeblich & Tappan, p. 164; pl. 364, figs 9-13; pl. 365,
figs 1-13.
2001 Hanzawaia grossepunctata (Earland) - Szareck, p. 147; pl. 26, figs 6-7.
Family Karreriidae Saidova, 1981
Karreria Rzehak, 1891
+Karreria maoria (Finlay, 1939) - p. 242
1939c Vagocibicides maoria - Finlay, p. 326; pl. 29, figs 148, 151, 158.
1999 Karreria maoria (Finlay) - Hayward et al., p. 161; pl. 15, fig. 30.
Superfamily Rotaliacea Ehrenberg, 1839
Family Rotaliidae Ehrenberg, 1839
Subfamily Pararotaliinae Reiss, 1963
Ammonia Brünnich, 1772
+Ammonia cf. A. aomoriensis (Asano, 1951) - p. 184
1951 Rotalia beccarii aomoriensis - Asano, p. 18; figs 96-98.
2004 Ammonia aomoriensis (Asano) - Hayward et al, p. 262; pl. 2, fig. T6; pl. 3, fig. T6; pl. 4, fig. T6.
+Ammonia aoteana (Finlay, 1940) - p. 185
1940 Streblus aoteanus - Finlay, p. 461.
1999 Ammonia beccarii (Linné) f. aoteana (Finlay) - Hayward et al., p. 162; pl. 16, figs 7-9.
2004 Ammonia aoteana (Finlay) - Hayward et al.; pl. 2, fig. T5; pl. 3, fig. T5; pl. 4, fig. T5.
2009 Ammonia aoteana (Finlay) Parker, p. 480; figs 344a-h.
*Ammonia beccarii (Linné, 1758)
Ammonia convexa (Collins, 1958) - p. 185
1958 Streblus convexus - Collins, p. 414; pl. 5, figs 10a-c.
1987 Ammonia convexa (Collins) - Baccaert, p. 232; pl. 94, figs -6.
1994 Ammonia tepida (Cushman) - Loeblich & Tappan; pl. 371, figs 1-3, not figs 4-10.
2004 Ammonia convexa (Collins) - Hayward et al, p. 262; pl. 2, fig. T13; pl. 3, fig. T13; pl. 4, fig. T13.
+Ammonia cf. irridescens (Arnal, 1958) - p. 185
1958 Streblus irridescens - Arnal, p. 41 ; pl. 4, figs 14-16.
1977 Ammonia irridescens (Arnal) - McCulloch, p. 431; pl. 151, figs 1-2.
2004 Ammonia irridescens (Arnal) - Hayward et al, p. 262; pl. 2, fig. T11; pl. 3, fig. T11; pl. 4, fig. T11.
*Ammonia parkinsoniana (d’Orbigny, 1839)
Ammonia pustulosa (Albani & Barbero, 1982) - p. 185
1982 Buccella pustulosa - Albani & Barbero, p. 238; pl. 1, figs 1-2.
1995 Buccella pustulosa Albani & Barbero - Yassini & Jones, p. 174; figs 985-987.
1999 Ammonia pustulosa (Albani & Barbero) - Hayward et al., p. 163; pl. 16, figs 4-6.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 323
1995 Elphidium advenum (Cushman) - Yassini & Jones, p. 176; figs 1026-1029, 1034-1036.
1997 Elphidium advenum advenum - Hayward et al., p. 65; pl. 2, figs 9, 14-15.
+Elphidium botaniense Albani, 1981 - p. 218
1981 Elphidium botaniense - Albani, p. 155, figs 4j, n.
1995 Elphidium botaniense Albani - Yassini & Jones, p. 176; figs 1030-1033.
1997 Elphidium advenum botaniense Albani - Hayward et al., p. 66; pl. 3, figs 1-4.
2009 Elphidium botaniense Albani - Parker, p. 572, figs 404a-h.
+Elphidium charlottense (Vella, 1957) - p. 218
1884 Polystomella subnodosa (Münster) - Brady, p. 734; pl. 110, figs 1a-c.
1957 Elphidiononion charlottensis - Vella, p. 38; pl. 9, figs 187, 188.
1997 Elphidium charlottense (Vella) - Hayward et al., p. 72; pl. 6, figs 13-16; pl. 7, figs 1, 2.
1999 Elphidium charlottense (Vella) - Hayward et al., p. 165; pl. 17, figs 6-8.
+Elphidium clavatum Cushman, 1930 - p. 219
1930 Elphidium incertum (Williamson) var. clavatum - Cushman, p. 20; pl. 7, fig. 10.
1997 Elphidium excavatum clavatum Cushman - Hayward et al., p. 76; pl. 9, figs 1-4.
Elphidium craticulatum (Fichtel & Moll, 1798) - p. 219
1798 Nautilus craticulatus - Fichtel & Moll, p. 51; pl. 5, figs h-k.
1987 Elphidium craticulatum (Fichtel & Moll) - Baccaert, p. 252; pl. 102, fig. 8; pl. 103, figs 1a, b.
1994 Cellanthus craticulatum (Fichtel & Moll) - Loeblich & Tappan, p. 167; pl. 380, figs 1, 2, 7-10.
1997 Elphidium craticulatum (Fichtel & Moll) - Hayward et al., p. 73; pl. 7, figs 5-12.
Elphidium crispum (Linné, 1758) - p. 219
1758 Nautilus crispus - Linnaeus, p. 709.
1992b Elphidium crispum (Linné) - Hatta & Ujiié, p. 203; pl. 49, fig. 5.
1994 Elphidium crispum (Linné) - Loeblich & Tappan, p. 168, pl. 378, figs 4-6.
2009 Elphidium crispum (Linné) - Parker, p. 575; figs 406a-h.
*Elphidium cf. earlandi Cushman
Elphidium excavatum (Terquem, 1875) [Cribroelphidium excavatum] - p. 219
1875 Polystomella excavata - Terquem, p. 429; pl. 2, figs 2a, b.
1975 Elphidium excavatum (Terquem) - Lévy et al., p. 174; pl. 3, figs 1, 2.
1997 Elphidium excavatum excavatum (Terquem) - Hayward et al., p. 77; pl. 9, figs 15-18, not figs 9-14.
2009 Elphidium cf. E. excavatum (Linné) - Parker, p. 576, figs 407a-e.
+Elphidium fichtelianum (d’Orbigny, 1846) - p. 219
1846 Polystomella fichtelianum - d’Orbigny, p. 125; pl. 6, figs 7-8.
1939 Elphidium fichtelianum (d’Orbigny) - Cushman, p. 42; pl. 11, fig. 12.
1997 Elphidium fichtelianum (d’Orbigny) - Hayward et al., p. 79; pl. 11, figs 1-8.
+Elphidium fijiense Hayward, 1997 - p. 219
1997 Elphidium fijiense - Hayward et al., p. 80; pl. 11, figs 9-12.
Elphidium gunteri Cole, 1931 [Cribroelphidium gunteri] - p. 220
1931 Elphidium gunteri - Cole, p. 34; pl. 4, figs 9, 10
1995 Elphidium vadescens Cushman & Brönnimann - Yassini & Jones, p. 178; fig. 1043.
1997 Elphidium gunteri Cole - Hayward et al., p. 80; pl. 11, figs 13-15.
+Elphidium hyalocostatum Todd, 1957 - p. 220
1957 Elphidium hyalocostatum - Todd, p. 300; pl. 88, fig. 19.
1994 Elphidium hyalocostatum Todd - Loeblich & Tappan, p. 169, pl. 386, figs 7-8.
*Elphidium jenseni (Cushman)
+Elphidium lene (Cushman & McCulloch, 1940) - p. 220
1940 Elphidium incertum (Williamson) var. lene - Cushman & McCulloch, p. 170; pl. 19, figs 2, 4.
1997 Elphidium lene Cushman & McCulloch - Hayward et al., p. 84, pl. 13, figs 1-8.
2009 Elphidium lene Cushman & McCulloch- Parker, p. 579; figs 408a-h, 409a-i.
Elphidium limbatum (Chapman, 1907) - p. 220
1907 Polystomella macellum var. limbatum - Chapman, p. 142; pl. 10, fig. 9.
1933 Elphidium depressulum - Cushman, p. 51; pl. 12, fig. 4.
1997 Elphidium advenum limbatum (Chapman) - Hayward et al., p. 67; pl. 3, figs 9-17; pl. 4, figs 1-10.
Elphidium macellum (Fichtel & Moll, 1798) - p. 220
1798 Nautilus macellus var. beta - Fichtel & Moll, p. 66; pl. 5, figs h, i, k.
1884 Polystomella macella (Fichtel & Moll) - Brady, p. 737; pl. 110, figs 8, 11.
1997 Elphidium macellum (Fichtel & Moll) - Hayward et al., p. 84; pl. 13, figs 9-10, ?figs 11-14.
2009 Elphidium cf. E. macellum (Fichtel & Moll) - Parker, p. 582; figs 410a-e.
+Elphidium maorium Hayward, 1997 - p. 220
1997 Elphidium maorium - Hayward et al., p. 69; pl. 4, figs 11-16; pl. 5, figs 1-5.
Elphidium milletti (Heron-Allen & Earland, 1915) [Parrellina milletti] - p. 221
1915 Polystomella milletti - Heron-Allen & Earland, p. 735; pl. 53, figs 38-42.
1987 Parrellina milletti (Heron-Allen & Earland) - Baccaert, p. 245-246; pl. 100, figs 4, 5; pl. 101, fig. 1.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 325
1993 Parrellina? cf. P.? milletti (Heron-Allen & Earland) - Hottinger et al., p. 152; pl. 218, figs 5-9;
pl. 219, figs 1-4.
2009 Elphidium milletti (Heron-Allen & Earland) - Parker, p. 582, figs 411a-i; 412a-f.
Elphidium oceanicum Cushman, 1933 [Cribroelphidium oceanicum] - p. 221
1933a Elphidium oceanicum - Cushman, p. 49; pl. 12, figs 7a, b.
1995 Cribrononion schmitti (Cushman & Wickenden) - Yassini & Jones, p. 179, fig. 1040.
1997 Elphidium oceanicum Cushman - Hayward et al., p. 88; pl. 15, figs 1-5.
2009 Elphidium oceanicum Cushman - Parker, p. 586; figs 413a-e; 414a-j.
*Elphidium poeyenum (d’Orbigny) (as Cribroelphidium poeyanum)
Elphidium sandiegoense (Lankford, 1973) - p. 221
1973 Cribrononion sandiegoense - Lankford in Lankford & Phleger, p. 118; pl. 3, figs 19a-b.
1994 Fijinonion fijiense (Cushman & Edwards) - Loeblich & Tappan, p. 159; pl. 346, figs 1-4.
1999 Elphidium sandiegoense (Lankford) - Hayward et al., p. 90; pl. 16, figs 9-11.
Elphidium tongaense (Cushman, 1931) [Ozawaia tongaensis] - p. 221
1931b Ozawaia tongaensis - Cushman, p. 80, pl. 10, figs 7-10.
1997 Elphidium advenum tongaense (Cushman) - Hayward et al., p. 70; pl. 5, figs 13-18.
Elphidium williamsoni Haynes, 1973 [Cribroelphidium williamsoni = Elphidium articulatum] -
p. 221
1973 Elphidium williamsoni - Haynes, p. 207, pl. 24, fig. 7; pl. 25, figs 6, 9; pl. 27, figs 1-3.
1993 Elphidium williamsoni Haynes - Hottinger et al., p. 150; pl. 215, figs 1-5.
1997 Elphidium excavatum williamsoni Haynes - Hayward et al., p. 78; pl. 10, figs 1-8.
2009 Elphidium cf. E. williamsoni Haynes - Parker, p. 591; figs 418a-l, 419a-e.
+Elphidium sp. 1 - p. 221
Porosononion Putrya in Voloshinova, 1958
+Porosononion shansiense (Wang, 1964) - p. 230
1964 Evolutononion shansiense - Wang, p. 58.
1993 Porosononion sp. A, - Hottinger et al., p. 153; pl. 219, figs 5-6; pl. 220, figs 1-6.
1994 Evolutononion shansiense Wang - Loeblich & Tappan, p. 157; pl. 342, figs 13-14.
2009 Porosononion sp. 1 - Parker, p. 713; figs 501a-h.
Porosononion simplex (Cushman, 1933) [Haynesina simplex] - p. 230
1933a Elphidium simplex - Cushman, p. 52; pl. 12, figs 8-9.
1995 Cribrononion simplex (Cushman) - Yassini & Jones, p. 179, fig. 1053.
1997 Haynesina depressula simplex (Cushman) - Hayward et al., p. 99; pl. 19, figs 8-10.
2009 Porosononion simplex (Cushman) - Parker, p. 711; figs 499a-e; 500a-l.
+Porosononion sp. 1 - p. 230
Subfamily Notorotaliinae Hornibrook, 1961
Cristatavultatus Loeblich & Tappan, 1994
Cristatavultatus pacificus (Collins, 1958) [Parrellina pacifica] - p. 218
1958 Elphidium pacificum - Collins, p. 421; pl. 5, fig. 13.
1992b Parrellina pacifica (Collins) - Hatta & Ujiié, p. 204; pl. 49, figs 8a-b; pl. 50, figs 1a-c.
1994 Cristatavultatus pacificus (Collins) - Loeblich & Tappan, p. 168; pl. 377, figs 7-8; pl. 378, figs 1-3.
1997 Cristatavultatus pacificus (Collins) - Hayward et al., p. 94 ; pl. 17, figs 14-15; pl. 18, figs 1-3.
Parrellina Thalmann, 1951
Parrellina hispidula (Cushman, 1936) - p. 229
1936 Elphidium hispidulum - Cushman, p. 83; pl. 14, fig. 13.
1994 Parrellina hispidula (Cushman) - Loeblich & Tappan, p. 170; pl. 384, figs 5-7; pl. 387, figs 1-3.
1997 Elphidium hispidulum Cushman - Hayward et al., p. 82; pl. 1, fig. 14; pl. 12, figs 5-7.
2009 Parrellina hispidula (Cushman 1936) - Parker, p. 683; figs 482a-f, 483a-e.
*Parrellina reticulosa (Cushman, 1951)
Superfamily Nummulitacea de Blainville, 1827
Family Nummulitidae de Blainville, 1827
Cycloclypeus Carpenter, 1856
Cycloclypeus carpenteri Brady, 1881 - p. 235
1881 Cycloclypeus carpenteri - Brady, p. 67.
1884 Cycloclypeus carpenteri Brady - Brady, p. 752.
1992b Cycloclypeus carpenteri Brady - Hatta & Ujiié, p. 204; pl. 50, figs 2, 3a-b.
2000 Cycloclypeus carpenteri Brady - Hohenegger et al., p. 25; pl. 4, fig. 7.
Heterostegina d’Orbigny, 1826
*Heterostegina curva Moebius
Heterostegina depressa d’Orbigny, 1826 - p. 222
1826 Heterostegina depressa - d’Orbigny, p. 305; pl. 17, figs 5-7.
1993 Heterostegina depressa d’Orbigny - Hottinger et al., p. 157; pl. 228, figs 1-11; pl. 229, figs 1-8;
pl. 230, fig. 9.
326 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
1994 Heterostegina depressa d’Orbigny - Loeblich & Tappan, p. 171; pl. 389, figs 1-6; pl. 390, figs 1-3.
2003 Heterostegina depressa d’Orbigny - Renema, p. 355, 356, figs 30a, b.
Heterostegina operculinoides Hofker, 1927 - p. 222
1927 Heterostegina operculinoides - Hofker, p. 67; pl. 34, figs 2, 4, 5.
1979 Heterostegina longisepta - Zheng, p. 226, pl. 23, fig. 8; pl. 27, fig. 8.
1992b Heterostegina longisepta Zheng - Hatta & Ujiié, p. 204, pl. 50, figs 5a-b, 6.
1993 Heterostegina operculinoides Hofker - Hottinger et al., p. 158; pl. 230, figs 1-8, 11.
*Heterostegina suborbicularis (d’Orbigny)
Nummulites Lamarck, 1801
Nummulites venosus (Fichtel & Moll, 1798) = Nummulites cumingii - p. 228
1798 Nautilus venosus - Fichtel & Moll, p. 59; pl. 8, figs e-h.
1884 Nummulites cumingii (Carpenter) - Brady, p. 749; pl. 112, figs 11-13; text-fig. 22.
1933a Operculinella venosa (Fichtel & Moll) - Cushman, p. 54; pl. 18, figs 2-6.
2000 Nummulites venosus (Fichtel & Moll) - Hohenegger et al., p. 11; pl. 1, figs 1-10.
Operculina d’Orbigny, 1826
Operculina ammonoides (Gronovius, 1781) - p. 228
1781 Nautilus ammonoides - Gronovius, p. 282; pl. 19, figs 5, 6.
1993 Assilina ammonoides (Gronovius) - Hottinger et al., p. 154; pl. 222, figs 1-8; pl. 223, figs 1-14;
pl. 224, figs 1-8; pl. 225, figs 1-9.
2000 Operculina ammonoides (Gronovius) - Hohenegger et al., p. 18; pl. 2, figs 7-12; pl. 5, figs 7-12.
2009 Assilina ammonoides (Schröter 1783) - Parker, p. 515; figs 367a-j.
Operculina bartschi Cushman, 1921 - p. 228
1921 Operculina bartschi - Cushman, p. 376; text-fig. 13.
1938 Operculina bartschi Cushman - Chapman & Parr, p. 292; pl. 17, figs 17-18; text-fig. 6.
1979 Operculina bartschi Cushman - Whittaker & Hodgkinson, p. 94; pl. 9, figs 10-12; pl. 10, figs 1-4,
6, 10-11.
Operculina discoidalis (d’Orbigny, 1826) - p. 228
1826 Nummulina (Assiline) discoidalis - d’Orbigny, p. 296, modèle no 88.
1865 Nummulina (Assilina) discoidalis d’Orbigny - Parker, Jones & Brady, p. 33; pl. 3, fig. 94.
2000 Operculina discoidalis (d’Orbigny) - Hohenegger et al., p. 21; pl. 2, figs 1-6; pl. 5, figs 1-6.
2009 Assilina discoidalis (d’Orbigny) - Parker, p. 519; figs 368a-e.
Operculina gaimardi d’Orbigny, 1826 - p. 228
1826 Operculina gaimardi - d’Orbigny, p. 281, no 5.
1921 Operculina gaimardi d’Orbigny - Cushman, p. 375.
*Operculina mayottana Le Calvez
Operculina philippinensis Cushman, 1921 - p. 229
1921 Operculina philippinensis - Cushman, p. 378; text-fig. 15.
Unassigned genus
Stictogongylus Loeblich & Tappan, 1994
Stictogongylus rugata (Heron-Allen & Earland, 1928) - p. 161
1928 Sphaeridia rugata - Heron Allen & Earland, p. 295; pl. 3, figs 38-43.
1994 Stictogongylus vandiemenensis - Loeblich & Tappan, p. 171; pl. 392, figs 1-8.
2009 Stictogongylus rugata (Heron Allen & Earland) - Parker, p. 740; figs 519a-g, 520a-f.
References
ABU-ZIED R. H., KEATINGS K. W., FLOWER R. J., 2007 – ANDRÉFOUËT S., CABIOCH G., FLAMAND B., PELLETIER B., 2007 –
Environmental controls on foraminifera in Lake Qarun, Egypt. “The diversity of New Caledonia coral reef geomorphology
Journal of Foraminiferal Research, 37: 136-149. and genetic processes: a synthesis from optical remote sensing,
coring and acoustic multi-beam observations”.
ADJAS A., 1988 – Sédimentologie comparée de quelques modèles
In Payri C., Richer de Forges B. (ed.): Compendium of marine
lagonaires actuels des milieux récifaux coralliens du Pacifique
species from New Caledonia, Nouméa, IRD: 33-49.
(Nouvelle-Calédonie, Polynésie).
Unpublished PhD thesis, université de Provence, 334 p. ANDRÉFOUËT S., CABIOCH G., FLAMAND B., PELLETIER B., 2009 –
A reappraisal of the diversity of geomorphological and genetic
ALBANI A. D., 1968 – Recent Foraminiferida from Port Hacking,
processes of New Caledonian coral reefs: a synthesis from optical
New South Wales. Contributions from the Cushman Laboratory
remote sensing, coring and acoustic multibeam observations.
for Foraminiferal Research, 19: 85-119.
Coral Reefs, 28: 691-707.
ALBANI A. D., 1974 – New benthonic foraminiferida from Australian
ARNAL R. E., 1958 – Rhizopoda from the Salton Sea, California.
waters. Journal of Foraminiferal Research, 4: 35-37.
Contributions from the Cushman Foundation for Foraminiferal
ALBANI A. D., 1978 – Recent foraminifera of an estuarine environment Research, 9: 36-45.
in Broken Bay, New South Wales. Australian Journal of Marine
ASANO K., 1936 – Fossil foraminifera from Muraoka-mura,
and Freshwater Research, 29: 355-398.
Kamakura-gori, Kanagawa Prefecture.
ALBANI A. D., 1981 – Pleistocene foraminifera from Botany Bay, Journal of the Geological Society of Japan, 43: 603-615.
New South Wales. Alcheringa, 5: 147-160.
ASANO K., 1951a – “Part 13: Anomalinidae”. In Stach L. W. (ed.):
ALBANI A. D., Barbero S. R., 1982 – Illustrated catalogue of Japanese Tertiary smaller foraminifera,
A foraminiferal fauna from the lagoon of Venice, Italy. Tokyo, Hosokawa Printing Co: 12-19.
Journal of Foraminiferal Research, 12: 234-241.
ASANO K., 1951b – “Part 6: Miliolidae”. In Stach L. W. (ed.):
ALBANI A. D., YASSINI I., 1989 – Taxonomy and distribution Illustrated catalogue of Japanese Tertiary smaller foraminifera,
of shallow-water lagenid Foraminiferida from the southeastern coast Tokyo, Hosokawa Printing Co: 1-20.
of Australia. Australian Journal of Marine and Freshwater
ASANO K., 1951c – “Part 14: Rotaliidae”. In Stach L. W. (ed.):
Research, 40: 369-401.
Illustrated catalogue of Japanese Tertiary smaller foraminifera,
ALCOCK T., 1865 – Notes on the natural history of specimens Tokyo, Hosokawa Printing Co: 1-21.
lately recorded from Connemara. Manchester Literary and
ASANO K., 1956 – The Foraminifera from the adjacent seas of Japan,
Philosophical Society Memoirs and Proceedings, 4: 192-208.
collected by S. S. SoyoMaru, 1922-1930. Pt. 1, Nodosariidae.
ALONGI D. M., 1992 – Benthic infauna and organism-sediment Tohoku University Science Reports, Sendai, Japan,
relations in a shallow, tropical coastal area: influence 2nd series (Geology), 27: 1-55.
of outwelled mangrove detritus and physical disturbance.
ASANO K., NAKAMURA M., 1937 – On the Japanese species of Cassidulina.
Marine Ecology Progress Series, 81: 229-245.
Japanese Journal of Geology and Geography, 14: 143-153.
ALORY G., VEGA A., GANACHAUD A., DESPINOY M., 2006 –
Influence of upwelling, subsurface stratification, and heat fluxes
on coastal sea surface temperature off southwestern New Caledonia.
Journal of Geophysical Research, 111 C07023:1-9. BACCAERT J., 1987 – Distribution patterns and taxonomy
ALTENBACH A. V., 1992 – Short-term processes and patterns of benthic foraminifera in the Lizard Island Reef Complex,
in the foraminiferal response to organic flux rates. northern Great Barrier Reef, Australia. Unpublished PhD Thesis,
Marine Micropaleontology, 19: 119-129. University of Liège. 3 vols, 109 pls.
ANDERSEN H. V., 1953 – Two new species of Haplophragmoides BAGG R. M. J., 1908 – Foraminifera collected near the Hawaiian Islands
from the Louisiana Coast. Contributions from the Cushman by the steamer ”Albatross” in 1902. Proceedings of the U.S.
Foundation for Foraminiferal Research, 4: 21-22. National Museum, 34 (1603): 113-172.
ANDERSEN H. V., 1961 – Genesis and paleontology of the Mississippi BAGG R. M. J., 1912 – Pliocene and Pleistocene Foraminifera from
River mudlumps, Part II: Foraminifera of the mudlumps, lower southern California. U.S. Geological Survey Bulletin, 513: 1-153, 28 pls.
Mississippi River delta. Louisiana Department of Conservation, BAILEY J. W., 1851 – Microscopical examination of soundings
Geological Bulletin, 35: 1-208. made by the United States Coast Survey, off the Atlantic Coast
of the United States. Smithsonian Contributions, 2: 1-15.
328 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
BALKWILL F. P., WRIGHT J., 1885 – Report on some Recent BRADY H. B., 1879 – Notes on some of the reticularian
foraminifera found off the coast of Dublin in the Irish Sea. Rhizopoda of the Challenger Expedition.
Transactions of the Royal Irish Academy, 28: 317-368. Part I. On new or little known arenaceous types,
part II. Additions to the Knowledge of porcellaneous and hyaline types.
BANNER F. T., PEREIRA C. P. G., DESAI D., 1985 – “Tretomphaloid”
Quarterly Journal of Microscopical Science,
float chambers in the Discorbidae and Cymbaloporidae.
new series, 19: 20-63; 261-299.
Journal of Foraminiferal Research, 15: 159-174.
BRADY H. B., 1881 – Notes on some of the reticularian Rhizopoda
BARKER R. W., 1960 – Taxonomic notes on the species figured
of the Challenger Expedition. part III. 1. Classification.
by H. B. Brady in his report on the foraminifera dredged
2. Further notes on new species. 3. Note on Biloculina mud.
by the H. M. S. Challenger during the years 1873-1876.
Quarterly Journal of Microscopical Science, new series, 21: 31-71.
Society of Economic Paleontologists and Mineralogists,
Special Publication, 9: 1- 240. BRADY H. B., 1882 – “Report on the Foraminifera”.
In Tizard T. H., Murray J. (eds): Exploration of the Faröe Channel
BASOV I. A., KRASHENNINNIKOV V. A., 1995 – Stratigraphy and foraminifera
during the summer of 1880, in Her Majesty’s hired ship
of Pliocene-Quarternary deposits of the Timor Trough.
“Knight Errant”, Proceedings of the Royal Society of Edinburgh,
Izdatelstvo “Nauchniy Mir”, Moscow, 1995: 1-112.
11: 638-720.
BATSCH A. I. G. C., 1791 – Sechs Kupfertafeln mit Conchylien BRADY H. B., 1884 – Report of the Foraminifera Dredged
des Seesandes, gezeichnet und gestochen von A.J.G.K. Batsch. Jena. by H. M. S. Challenger during the years 1873-1876.
BECKMAN J. P., 1954 – Foraminiferen der Oceanic Formation, Barbados. Reports of the Scientific Results of the Voyage of H. M. S.
Ecologae Geologicae Helvetiae, 46: 301-407. Challenger during the years 1873-187, Zoology, 9: 1-814.
BELFORD D. J., 1966 – Miocene and Pliocene foraminifera BRADY H. B., 1890 – Notes on a new type of foraminifera of the famiy
from Papua and New Guinea. Bureau of Mineral Resources chilostomellidae. Journal of the Royal Microscopical Society,
(Australia) Bulletin, 79: 1-306. London: 567-571.
BERMÚDEZ P. J., 1935 – Foraminiferos de la costa norte de Cuba. BRADY H. B., PARKER W. K., JONES T. R., 1870 – A monograph
Memorias de la Sociedad Cubana de Historia Natural, 9: 129-224. of the genus Polymorphina. Transactions of the Linnean Society
of London, 27: 197-253.
BERMÚDEZ P. J., 1952 – Estudio sistemático de los foraminiferos
rotaliformis. Boletín de Geología (Caracas), 2: 1-230. BRÖNNIMANN P., BEURLEN G., 1977 – Paraibaella,
New Name for the Foraminiferal Genus Spiroplectammina
BERMÚDEZ P. J., SEIGLIE G. A., 1963 – Estudio sistemático Brönnimann and Beurlen, 1977.
de los foraminiferos del golfo de Cariaco. Boletín del Instituto Archives Sciences, Genève, 320: 279.
de Oceanográphica de la Universidad de Oriente, 2: 3-253.
BRÖNNIMANN P., WHITTAKER J.E., 1980 – A Revision of Reophax
BERNHARD J. M., BOWSER S. M., 1999 – Benthic foraminifera of dysoxic and its Type-Species, with Remarks on Several Other Recent
sediments: chloroplast sequestration and functional morphology. Hormosinid (Protozoa: Foraminiferida) in the Collections
Earth Science Reviews, 46: 149-165. of the British Museum (Natural History). Bulletin of the British
BICCHI E., DEBENAY J. P., PAGÈS J., 2002 – Relationship between benthic Museum of Natural History (Zoology), London, 39/5: 259-272.
foraminiferal assemblages and environmental factors in atoll BRÖNNIMANN P., WHITTAKER J. E., 1983 – Zaninettia n. gen.,
lagoons of the central Tuamotu Archipelago (French Polynesia). a Spicular-Walled Remaneicid (Foraminiferida, Trochamminacea)
Coral Reefs, 21: 275-290. from the Indian and South Atlantic Oceans with Remarks
BILLMAN H., HOTTINGER L., OESTERLE H., 1980 – Neogene to Recent on the Origin of the Spicules.
Rotaliid Foraminifera from the Indopacific Ocean; Revue de Paléobiologie, Genève, 2: 13-33.
their canal system, their classification and their stratigraphic use. BRÖNNIMANN P., WHITTAKER J. E., 1984 – A neotype for Trochammina
Schweizerische Palaeontologische Abhandlungen, 101: 71-113. inflata (Montagu) (Protozoa: Foraminiferida) with notes
BLAINVILLE H. M. DUCROTAY de, 1830 – on the wall structure. Bulletin of the British Museum of Natural
Zoophytes. Dictionnaire des sciences naturelles. History (Zoology), 46: 311-315.
Paris, F. G. Levrault, 60: 1-546. BRÖNNIMANN P., ZANINETTI L., 1965 – Note sur Lituola salsa
BOLTOVSKOY E., GIUSSANI de KAHN G., 1981 – Cinco nuevos taxones (Cushman et Brönnimann, 1948), un foraminifère
en Orden Foraminiferida. Comunicaciones del Museo Argentino de la mangrove de l’île de la Trinité, W. I.
de Ciencias Naturales (Bernardino Rivadavia) Archives Sciences, Genève, 18 (3): 608-615.
Hidrobiología, 2: 43-51. BRÖNNIMANN P., ZANINETTI L., 1984 –
Agglutinated Foraminifera mainly Trochamminacea
BORNEMANN J. G., 1855 – Die mikroskopische Fauna
from the Baia de Sepetiba, near Rio de Janeiro, Brazil.
des Septarienthones von Hermsdorf bei Berlin.
Revue de Paléobiologie, Genève, 3: 63-115.
Zeitschrift der Deutschen Geologischen Gesellschaft, 7: 307-371.
BRÖNNIMANN P., WHITTAKER J. E., ZANINETTI L., 1992 –
BRADY H. B., 1864 – Contributions to the Knowledge
Brackish water foraminifera from mangrove sediments
of the Foraminifera. On the Rhizopodal Fauna of the Shetlands.
of southwestern Viti Levu, Fiji Islands, southwest Pacific.
Transactions of the Linnean Society of London, 24: 463-476.
Revue de Paléobiologie, 11: 13-65.
BRADY H. B., 1870 – Analysis and descriptions of the foraminifera.
BUCHNER P., 1940 – Die Lagenen des Golfes von Neapol
Annuals and Magazine of Natural History, ser. 4, 6: 273-309.
und der marinen Ablagerungen auf Ischia
BRADY H. B., 1876 – On some foraminifera from the Loo Choo islands. (Beiträge zur Naturgeschichte der Insel Ischia 1).
Proceedings of the Royal Irish Academy 2nd series, 2: 1-600. Nova Acta Leopoldina, n.f. 9(26): 363-560.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 329
BUZAS M. A., SMITH R. K., BEEM K. A., 1977 – Ecology and systematics CHAPMAN F., PARR W. J., 1931 – Notes on new and aberrant types
of foraminifera in two Thalassia habitats, Jamaica, West Indies. of foraminifera. Proceedings of the Royal Society of Victoria,
Smithsonian Contributions to Paleobiology, 31: 1-139. 43 (2): 236-240.
CHAPMAN F., PARR W. J., 1935 – Foraminifera and Ostracoda from
soundings made by the trawler “Bonthorpe” in the Great Australian
Bight. Royal Society of Western Australia, Journal, 21: 1-6.
CABIOCH G., 1988 – Récifs frangeants de Nouvelle-Calédonie
(Pacifique sud-ouest). Structure interne et influences CHAPMAN F., PARR W. J., 1937 – Foraminifera. Australasian Antarctic
de l’eustatisme et de la néotectonique. Unpublished PhD thesis, Expedition 1911-1914 Scientific Reports Series C, 1: 1-190.
université de Provence (Aix-Marseille I), 322 p., 25 pls. CHAPMAN F., PARR W. J., 1938 – Australian and New Zealand species
CABIOCH G., 2001 – Synthèse scientifique : « Croissance récifale of foraminiferal genera Operculina and Operculinella.
postglaciaire dans le Pacifique occidental et central - Proceedings of the Royal Society of Victoria, 50: 279-294.
exemples de la Nouvelle-Calédonie, du Vanuatu CHAPMAN F., PARR W. J., COLLINS A. C., 1934 – Tertiary foraminifera
et de la Polynésie française ». Perspectives scientifiques : of Victoria, Australia. The Balcombian deposits of Port Philip;
« Détermination des facteurs à l’origine de l’arrêt Part III. Journal of the Linnaean Society of London, Zoology, 38:
de la croissance des récifs indo-pacifiques lors de la dernière 533-577.
déglaciation - exemple des récifs ennoyés des Marquises
CHARDON D., AUSTIN J. A. JR., CABIOCH G., PELLETIER B., SAUSTRUP S.,
et de Madagascar et cas particuliers des récifs
SAGE F., 2008 – Neogene history of the northeastern New Caledonia
de Nouvelle-Calédonie et du Vanuatu ».
continental margin from multichannel reflection seismics profiles.
Unpublished Mémoire d’habilitation à diriger des recherches
Comptes-Rendus de Géoscience, 340 (1): 68-73.
(Aix-Marseille I), 125 p.
CHASTER G. W., 1892 – Report upon the Foraminifera
CABIOCH G., ANGLADA R., BABINOT J. F., 1986 – Microfaunes
of the Southport Society of Natural Science District.
et paléoenvironnements des récifs frangeants quaternaires
Report of the Southport Society of Natural Science, 1: 54-72.
de Mamié et Ricaudy (Nouvelle-Calédonie).
Cahiers de Micropaléontologie, 1(1-2): 5-36, 13 pls. CHENG T. C., ZHENG S. Y., 1978 – The recent foraminifera of the Xisha
Islands, Guangdong Province, China, I. Studia Marina Sinica, 12:
CARPENTER W. B., PARKER W. K., JONES T. R., 1862 – Introduction
149-227. [Chinese with summary and new genera and species in
to the study of foraminifera. Ray Society, London: 1-319, 22 pls.
English]
CARTER H. J., 1876 – On the Polytremata (Foraminifera),
CHEVILLON C., 1996 – Skeletal composition of modern lagoon sediments
especially with reference to their Mythical Hybrid Nature.
in New Caledonia: Coral, a minor constituent. Coral Reefs, 15 (3):
Annals and Magazine of Natural History, London, 4 (17):
199-207.
185-214.
CHEVILLOTTE V., DOUILLET P., CABIOCH G., LAFOY Y., LAGABRIELLE Y.,
CARTER H. J., 1877a – Description of a new species of Foraminifera
MAURIZOT P., 2005 – Évolution géomorphologique de l’avant-pays
(Rotalia spiculotesta). Annals and Magazine of Natural History,
du sud-ouest de la Nouvelle-Calédonie durant les derniers cycles
London, 4 (20): 470-473.
glaciaires. Comptes Rendus Geosciences, 337 (7): 695-701.
CARTER H. J., 1877b – On the locality of Carpenteria balaniformis,
CIMERMAN F., LANGER M., 1991 – Mediterranean foraminifera.
with description of a new species and other Foraminifera,
Slovenska Akademija Znanosti in Umetnosti, Ljubljana.
found in and about Tubipora musica.
1-118, 93 pls.
Annals and Magazine of Natural History, London, 4 (19):
209-219. CLARK F. L., 1993 – Rhaptohelenina papuanensis, a new genus
and species of benthic foraminifer from the Recent of the Papuan
CARTER H. J., 1880 – Report on specimens dredged up from the Gulf
Platteau and Alexa Bank. Journal of Paleontology, 67: 899-901.
of Manaar, and presented to the Liverpool Free Museum by Capt.
W.H. Cawne Warren. Annals and Magazine of Natural History, CLARK F. L., 1995 – New species of unilocular calcareous foraminifera
London, 5: 437-457. from the Holocene of the southwest Pacific Ocean.
CHAPMAN F., 1898 – On Haddonia, a new genus of the foraminifera Journal of Micropaleontology, 14: 1-5.
from Torres Straits. Journal of the Linnaean Society of London, CLOSS D., 1963 – Foraminíferos e Tecamebas da Lagoa dos Patos
Zoology, 26: 452-456. (RGS). Boletim da Escola de Geologia, Porto Alegre, 11: 1-130.
CHAPMAN F., 1900 – On some new and interesting foraminifera COLE W. S., 1931 – The Pliocene and Pleistocene foraminifera
from the Funafuti Atoll, Ellice Islands. Journal of the Linnaean of Florida. Bulletin of the Florida State Geological Survey, 6: 7-79.
Society of London, Zoology, 28: 1-27. COLLEN J. D., 1998 – Metarotaliella tuvaluensis sp. nov. from Funafuti
CHAPMAN F., 1901 – Foraminifera from the lagoon at Funafuti. Atoll, western Pacific Ocean: relationship to miliolid foraminifera.
Journal of the Linnaean Society of London, Zoology, 28: 161-210. Journal of Foraminiferal Research, 28: 66-75.
CHAPMAN F., 1902 – On the foraminifera collected around the Funafuti COLLINS A. C., 1958 – “Foraminifera”.
Atoll from shallow and moderately deep water. In: Great Barrier Reef Expedition 1928-1929,
Journal of the Linnaean Society of London, Zoology, 28: 379-417. British Museum of Natural History, 6: 335-437.
CHAPMAN F., 1907 – Recent foraminifera of Victoria: COLLINS A. C., 1974 – Port Phillip survey 1957-63 Foraminiferida.
Some littoral gatherings. Journal of the Quekett Microscopical Memoirs of the National Museum of Victoria, 35: 1-61.
Club series 2, 10: 117-146. COLLINS E. S., SCOTT D. B., ZHANG J., 1996 – Neogene benthic
CHAPMAN F., 1909 – On some microzoa from the Wianamatta Shales, foraminifers from Ocean Drilling Project sites 898 and 900, Leg 149,
New South Wales. New South Wales Geological Survey Records, Iberia Abyssal Plain. Proceedings of the Ocean Drilling Program,
Department of Mines, Sydney, Australia, 8: 334-339. Scientific Results, 149: 217-239.
330 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
COUDRAY J., 1976 – Recherches sur le Néogène et le Quaternaire CUSHMAN J. A., 1926a – Foraminifera of the genera Siphogenerina
marins de la Nouvelle-Calédonie. Contribution de l’étude and Pavonina. Proceedings of the United States National
sédimentologique à la connaissance de l’histoire géologique Museum, 67 (25): 1-24.
post-éocène de la Nouvelle-Calédonie. CUSHMAN J. A., 1926b – Recent foraminifera from Porto Rico.
Thèse doc. d’État, université de Montpellier Publications of the Carnegie Institution of Washington, no. 344,
In: Expéd. française sur les récifs coralliens de Nouvelle-Calédonie, Department of Marine Biology Papers, 23: 73-84.
Fond. Singer-Polignac, Paris, 8: 1-276.
CUSHMAN J. A., 1927 – An outline of a reclassification
COUDRAY J., MARGEREL J. P., 1974 – Les foraminifères de la série of the foraminifera. Contributions from the Cushman
récifale traversée par le sondage Ténia (côte sud-ouest Laboratory for Foraminiferal Research, 3: 1-105.
de Nouvelle-Calédonie). Comptes-Rendus de l’Académie des Sciences,
CUSHMAN J. A., 1929a – The foraminifera of the Atlantic Ocean,
Paris (D), 279: 231-234.
Part 6. Miliolidae, Opthalmidiidae and Fischerinidae.
CUSHMAN J. A., 1910 – A monograph of the foraminifera Bulletin of the United States National Museum, 104 (6): 1-129.
of the North Pacific Ocean. Part 1. Astrorhizidae and Lituolidae. CUSHMAN J. A., 1929b – The genus Trimosina
Bulletin of the United States National Museum, 71(1): 1-134. and its relationships to other genera of the foraminifera.
CUSHMAN J. A., 1911 – A monograph of the foraminifera Contributions from the Cushman Laboratory for Foraminiferal
of the North Pacific Ocean. Part 2. Textulariidae. Research, 19: 155-159.
Bulletin of the United States National Museum, 71 (2): 1-108. CUSHMAN J. A., 1929c – Notes on the Foraminiferal Fauna
CUSHMAN J. A., 1913a – New Textulariidae and other arenaceous of the Byram marl. Contributions from the Cushman Laboratory
foraminifera from the Philippine Islands and contiguous waters. for Foraminiferal Research, 5: 40-48.
Proceedings of the United States National Museum, 44: 633-638. CUSHMAN J. A., 1929d – A late Tertiary fauna of Venezuela
CUSHMAN J. A., 1913b – A monograph of the foraminifera and other related regions. Contributions from the Cushman
of the North Pacific Ocean. Pt. 3 - Lagenidae. Laboratory for Foraminiferal Research, 5: 77-101.
Bulletin of the United States National Museum, 71 (3): 1-125. CUSHMAN J. A., 1930 – The foraminifera of the Atlantic Ocean, Part. 7:
Nonionidae, Camerinidae, Peneroplidae, and Alveolinellidae.
CUSHMAN J. A., 1914 – A monograph of the foraminifera
Bulletin of the United States National Museum, 104 (7): 1-79.
of the North Pacific Ocean. Part 4. Chilostomellidae, Globigerinidae,
Nummulitidae. Bulletin of the United States National Museum, CUSHMAN J. A., 1931a – The foraminifera of the Atlantic Ocean,
71(4): 1-45. Part 8. Rotaliidae, Amphisteginidae, Calcarinidae,
Cymbaloporettidae, Globorotaliidae, Anomalinidae,
CUSHMAN J. A., 1915 – A monograph of the foraminifera
Planorbulinidae, Rupertiidae and Homotrematidae.
of the North Pacific Ocean. Part 5. Rotaliidae.
Bulletin of the United States National Museum, 104 (8): 1-144.
Bulletin of the United States National Museum, 71 (5): 1-81.
CUSHMAN J. A., 1931b – Two new foraminiferal genera
CUSHMAN J. A., 1917 – A monograph of the foraminifera from the south Pacfic. Contributions from the Cushman
of the North Pacific Ocean. Part 6. Miliolidae. Laboratory for Foraminiferal Research, 7: 78-82.
Bulletin of the United States National Museum, 71 (6): 1-108.
CUSHMAN J. A., 1932 – The foraminifera of the Tropical Pacific collections
CUSHMAN J. A., 1919 – Recent foraminifera from off New Zealand. of the “Albatross”, 1899-1900. Part 1. Astrorhizidae to Trochamminidae.
Proceedings of the United States National Museum, 56: 593-640. Bulletin of the United States National Museum, 161: 1-88.
CUSHMAN J. A., 1920 – The foraminifera of the Atlantic Ocean, CUSHMAN J. A., 1933a – The foraminifera of the Tropical Pacific
Part II. Lituolidae. Bulletin of the United States National collections of the “Albatross”, 1899-1900. Part 2. Lagenidae
Museum, 104 (2): 1-111. to Alveolinellidae. Bulletin of the United States National Museum,
CUSHMAN J. A., 1921 – Foraminifera of the Philippine and adjacent seas. 161: 1-79.
Bulletin of the United States National Museum, 100: 1-608. CUSHMAN J. A., 1933b – Some new Recent foraminifera from the tropical
Pacific. Contributions from the Cushman Laboratory
CUSHMAN J. A., 1922a – Shallow-water Foraminifera of the Tortugas
for Foraminiferal Research, 9: 77-95.
region. Publications of the Carnegie Institutue of Washington,
no. 311, Department of Marine Biology Papers, 17: 1-85. CUSHMAN J. A., 1934 – Notes on the genus Tretomphalus,
with descriptions of some new species and a new genus, Pyropilus.
CUSHMAN J. A., 1922b – The foraminifera of the Atlantic Ocean,
Contributions from the Cushman Foundation for Foraminiferal
Part 3. Textulariidae. Bulletin of the United States National
Research, 10: 79-101.
Museum, 104 (3): 1-143.
CUSHMAN J. A., 1936 – New genera and species of the families
CUSHMAN J. A., 1922c – The foraminifera of the Byram calcareous Verneuilinidae and Valvulinidae and of the subfamily Virgulininae.
marl at Byram, Mississippi. Professional Papers U.S. Geological Cushman Laboratory for Foraminiferal Research,
Survey, 129-E: 87-152. Special Publication, 6: 1-71.
CUSHMAN J. A., 1923 – The foraminifera of the Atlantic Ocean, CUSHMAN J. A., 1937 – A monograph of the foraminiferal family
Part 4. Lagenidae. Bulletin of the United States National Museum, Valvulinidae. Cushman Laboratory for Foraminiferal Research,
104 (4): 1-228. Special Publication, 8: 1-210.
CUSHMAN J. A., 1924 – Samoan Foraminifera. CUSHMAN J. A., 1939 – A monograph of the foraminiferal family
Publications of the Carnegie Institution of Washington, no 342, Nonionidae. Professional Papers U.S. Geological Survey,
Department of Marine Biology Papers, 21: 1-85. 191: 1-100.
CUSHMAN J. A., 1925 – Foraminifera of the tropical central Pacific. CUSHMAN J. A., 1942 – The foraminifera of the tropical Pacific
Bernice Pauachi Bishop Museum Bulletin, Honolulu, Hawaii, collections of the “Albatross”, 1899-1900. Part 3, Heterohelicidae
27: 121-144. and Buliminidae. Bull. U.S. Nat. Mus., 161: 1-67.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 331
CUSHMAN J. A., 1944a – The genus Articulina and its species. CUSHMAN J. A., PARKER F. L., 1936 – Some species of Robertina.
Special Publications, Cushman Laboratory for Foraminiferal Contributions from the Cushman Laboratory for Foraminiferal
Research, 10: 1-37. Research, 12: 92-100.
CUSHMAN J. A., 1944b – Foraminifera from the shallow water CUSHMAN J. A., PARKER F. L., 1942 –The foraminifera
of the New England coast. Special Publications, Cushman of the tropical Pacific Collections of the ‘Alhatross’ 1899-1900;
Laboratory for Foraminiferal Research, 12: 1-37. Part 3 -Heterohelicidae and Buliminidae.
Bulletin of the United States National Museum, 161: 1-67.
CUSHMAN J. A., 1945 – The species of the subfamiliy Reussellinae
of the foraminiferal family Buliminidae. Contributions from CUSHMAN J. A., PARKER F. L., 1947 – Bulimina and related foraminiferal
the Cushman Laboratory for Foraminiferal Research, 21: 23-54. genera. Professional Papers U.S. Geological Survey, 210: 55-176.
CUSHMAN J. A., 1946 – Upper Cretaceous Foraminifera of the Gulf CUSHMAN J. A., TODD R., 1943 – The genus Pullenia and its species.
Coastal region of the United States and adjacent areas. Contributions from the Cushman Laboratory for Foraminiferal
Professional Papers U.S. Geological Survey, 206: 1-241. Research, 19 (1): 1-23.
CUSHMAN J. A., BRÖNNIMANN P., 1948 – Some new genera and species CUSHMAN J. A., TODD R., 1944 –The genus Spiroloculina and its species.
of Foraminifera from brackish water of Trinidad. Special Publications, Cushman Laboratory for Foraminiferal
Contributions from the Cushman Laboratory for Foraminiferal Research, 11: 1-82.
Research, 24: 15-21. CUSHMAN J. A., VALENTINE W. W., 1930 – Shallow-water Foraminifera
from the Channel Islands of southern California. Contributions
CUSHMAN J. A., EDWARDS P. G., 1937 – Astrononion a new genus
of the Department of Geology, Stanford University, 1: 1-51.
of the foraminifera, and its species. Contributions from the Cushman
Laboratory for Foraminiferal Research, 13: 29-36. CUSHMAN J. A., WICKENDEN R. T. D., 1929 – Recent Foraminifera
from off Juan Fernandez Islands. Proceedings of the United States
CUSHMAN J. A., GRAY H. B., 1946 – A Foraminiferal Fauna
Natural Museum, 75 (9): 1-16, 6 pls.
from the Pliocene of Timms Point,. California.
Contributions from the Cushman Laboratory for Foraminiferal CUSHMAN J. A., TODD R., POST R. J., 1954 – Recent foraminifera
Research, Spec. Publ., 19: 1-46. of the Marshall Islands, Bikini and nearby atolls, Part II,
oceanography (biologic).
CUSHMAN J. A., HANZAWA S., 1936 – New genera and species of foraminifera
Professional Papers U.S. Geological Survey, 260-H: 319-384.
of late Tertiary of the Pacific. Contributions from the Cushman
Laboratory for Foraminiferal Research, 12: 45-48. CZJZEK J., 1848 – Beitrag zur Kenntniss der fossilen Foraminiferen
des Wiener Beckens. Haidinger’s Natur-wissenschaftliche
CUSHMAN J. A., HANZAWA S., 1937 – Notes on Some of the Species Referred Abhandlungen, Wien, 2 (1): 137-150.
to Vertebralina and Articulina, and a New Genus Nodobaculariella,
Contributions from the Cushman Laboratory for Foraminiferal
Research, 13: 41-46.
CUSHMAN J. A., MCCULLOCH I., 1939 – A report on some arenaceous
DE MONTFORT P. D., 1808 – Conchyliologie systématique
et classification méthodique des coquilles, t. 1.
Foraminifera. Southern California University Publications, Allan
Paris, France, F. de Schoell ed.: 1-410.
Hancock Pacific Expedition, Los Angeles, California, 6: 1-113.
DEBENAY J.-P., 1985a – Le lagon sud-ouest et la marge insulaire sud
CUSHMAN J. A., MCCULLOCH I., 1940 – Some Nonionidae
de la Nouvelle-Calédonie : importance et répartition des foraminifères
in the collections of the Allan Hancock Foundation.
de grande taille. Océanographie tropicale, 20 (2) : 171-192.
Southern California University Publications, Allan Hancock
Pacific Expedition, Los Angeles, California, 6: 145-178. DEBENAY J.-P., 1985b – Le genre Amphistegina dans le lagon
de Nouvelle-Calédonie (S.W. Pacifique).
CUSHMAN J. A., MCCULLOCH I., 1942 – Some Virgulininae
Revue de Micropaléontologie, 28 (3) : 167-180.
in the collections of the Allan Hancock Foundation.
Allan Hancock Pacific Expeditions, 6 (4): 179-230. DEBENAY J.-P., 1986 – Recherche sur la sédimentation actuelle
et les thanatocoenoses des Foraminifères de grande taille
CUSHMAN J. A., MCCULLOCH I., 1948 – The species of Bulimina du lagon sud-ouest et de la marge insulaire sud
and related genera in the collections of the Allan Hancock Foundation. de Nouvelle-Calédonie. Paris, Orstom, Travaux et documents
Southern California University Publications, Allan Hancock microédités, 20 (3 vol.) : 1-200, 22 pl.
Pacific Expedition, Los Angeles, California, 6: 179-230.
DEBENAY J.-P., 1987 – Sedimentology in the south western
CUSHMAN J. A., MCCULLOCH I., 1950 – Reports on the collections obtained lagoon of New Caledonia (SW Pacific).
by Allan Hancock Expeditions of Velero III off the coast of Mexico, Journal of Coastal Research, 3 (1): 77-91.
Central America, South America and Galapagos Islands in 1932-
DEBENAY J.-P., 1988a – Foraminifera larger than 0.5 mm
1941, and Velero IV in 1949. Some Lagenidae in the collections of
in the Southwestern lagoon of New Caledonia:
the Allan Hancock Foundation. Allan Hancock Pacific Expeditions,
distribution related to abiotic properties.
6: 295-364.
Journal of Foraminiferal Research, 18 (2): 158-175.
CUSHMAN J. A., OZAWA Y., 1928 – An outline of a revision
DEBENAY J.-P., 1988b – Recent Foraminifera tracers
of the Polymorphinidae. Contributions from the Cushman
of oceanic water movements in the Southwestern lagoon
Laboratory for Foraminiferal Research, 4: 13-21.
of New Caledonia. Palaeogeography, Palaeoclimatology,
CUSHMAN J. A., OZAWA Y., 1930 – A monograph of the foraminiferal Palaeoecology, 65: 59-72.
family Polymorphinidae, Recent and fossil. DEBENAY J.-P., 1988c – Dynamique sédimentaire au débouché
Proceedings of the United States National Museum, 77: 1-195. de la baie du Prony (Nouvelle-Calédonie) : dispersion des lutites
CUSHMAN J. A., PARKER F. L., 1931 – Recent foraminifera et d’un test de Foraminifère, Operculina bartschi Cushman.
from the Atlantic coast of South America. Benthos 86, Genève, 1986. Revue de paléobiologie, Genève,
Proceedings of the United States National Museum, 80: 1-24. volume special, 2 : 765-770.
332 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
DEBENAY J.-P., 1990 – Recent foraminiferal assemblages DEBENAY J.-P., SIGURA A., JUSTINE J.-L., 2011 – Foraminifera in the diet
and their distribution related to environmental stress in the paralic of coral reef fish from the lagoon of New Caledonia: predation,
environments of West Africa (Cape Timiris to Ebrie Lagoon). digestion, dispersion. Revue de Micropaléontologie, 54: 87-103.
Journal of Foram. Research, 20: 267-282. DEGAUGUE-MICHALSKI F.M., 1993 – Croissance et évolution
DEBENAY J.-P., CABIOCH G., 2007 – “Recent and Quaternary foraminifera d’édifices récifaux du Pacifique occidental (Nouvelle-Calédonie,
collected around NewCaledonia”. In Payri C. E., Richer de Froges B. Chesterfield) à l’Holocène et au Pléistocène.
(eds): Compendium of marine species of New Caledonia. Unpublished PhD thesis, université Aix-Marseille I, 227 p.
IRD Nouméa, Documents Scientifiques et Techniques, 117, DOUILLET P., 1998 – Tidal dynamics of the south-west lagoon
seconde edition: 69-94. of New Caledonia: observations and 2D numerical modelling.
DEBENAY J.-P., DECROUEZ D., 1989 – Les collections du département Oceanologica Acta, 21: 69-79.
de Géologie et de Paléontologie des invertébrés du Muséum d’histoire DOUILLET P., OUILLON S., CORDIER E., 2001 – A numerical model for fine
naturelle de Genève. 30. La collection de micropaléontologie suspended sediment transport in the south-west lagoon of New
(4e partie). Revue de paléobiologie, Genève, 8 (1) : 255-258. Caledonia. Coral Reefs, 20: 361-372.
DEBENAY J.-P., DELLA PATRONA L., 2009 – Foraminifères : DUBOIS J., LAUNAY J., RÉCY J., 1974 –
bio-indicateurs de la qualité des fonds de bassins d’élevage Uplift movements in New Caledonia - Loyalty islands area
de crevettes en Nouvelle-Calédonie. and their plate tectonics interpretation. Tectonophysics, 24: 133-150.
Édition conjointe IRD-Ifremer : 1-77.
DUGAS F., DEBENAY J.-P., 1978 – Carte sédimentologique
DEBENAY J.-P., FERNANDEZ J.-M., 2009 – Benthic foraminifera records et carte annexe du lagon de Nouvelle-Calédonie (1/50 000).
of complex anthropogenic environmental changes combined with Pub. Orstom Feuille « Mont-Dore ».
geochemical data in a tropical bay of New Caledonia (SW Pacific). DUGAS F., DEBENAY J.-P., 1980 – Carte sédimentologique
Marine Pollution Bulletin, 59: 311-322. et carte annexe du lagon de Nouvelle-Calédonie au 1/50 000.
DEBENAY J.-P., GUILLOU J. J., 2002 – Ecological transitions Publ. Orstom Feuille « Tontouta ».
indicated by foraminiferal assemblages in paralic environments. DUGAS F., DEBENAY J.-P., 1981a – Carte sédimentologique
Estuaries, 25: 1107-1120. du lagon de Nouvelle-Calédonie au 1/200 000. 1 carte + notice, 3 p.
DEBENAY J.-P.,. GUIRAL D., 2006 – Mangrove swamp foraminifera, (format 53 x 38) in Atlas de Nouvelle-Calédonie.
indicators of sea level or paleoclimate? DUGAS F., DEBENAY J.-P., 1981b – Carte sédimentologique
Revue de Paléobiologie, 25: 567-574. et carte annexe du lagon de Nouvelle-Calédonie au 1/50 000.
DEBENAY J.-P., LUAN B. T., 2006 – Foraminiferal assemblages Publ. Orstom Feuille « Prony ».
and the confinement index as tools for assessment DUGAS F., DEBENAY J.-P., 1982 – Carte sédimentologique
of saline intrusion and human impact in the Mekong delta. et carte annexe du lagon de Nouvelle-Calédonie au 1/50 000.
Revue de Micropaléontologie, 49: 74-85. Publ. Orstom Feuille « Nouméa ».
DEBENAY J.-P., PAYRI C., 2010 – Epiphytic foraminiferal assemblages DUJARDIN F., 1835 – Observations nouvelles sur les prétendus
on macroalgae in reefal environments of New Caledonia. Céphalopodes microscopiques. Annales de Sciences Naturelles,
Journal of Foraminiferal Research, 40: 36-60. 2 (3) : 312-314.
DEBENAY J.-P., PAWLOWSKI J., DECROUEZ D., 1996 –
Les foraminifères actuels. Paris, Éditions Masson : 1-329.
DEBENAY J.-P., EICHLER B. B, DULEBA W., BONETTI C., EARLAND A., 1933 – Foraminifera. Part II. South Georgia.
EICHLER-COELHO P., 1998 – Water stratification in coastal lagoons: Discovery Reports, 7: 27-138.
its influence on foraminiferal assemblages in two Brazilian lagoons. EARLAND A., 1934 – Foraminifera. Part III. The Falklands sector
Marine Micropaleontology, 35 (1-2): 65-89. of the Antarctic (excluding South Georgia).
DEBENAY J.-P., TSAKIRIDIS E., SOULARD R., GROSSEL H., 2001 – Discovery Reports, 10: 1-208.
Factors determining the distribution of foraminiferal assemblages EGGER J. G., 1857 – Die Foraminiferen der Miocän-Schichten
in Port Joinville Harbor (île d’Yeu, France): the influence of pollution. bei Ortenburg in Nieder- Bayern. Neues Jahrbuch für Mineralogie,
Marine Micropaleontology, 43: 75-118. Geognosie, Geologie, und Petrefakten- Kunde: 266-311.
DEBENAY J.-P., GUIRAL D., PARRA M., 2002 – Ecological factors EGGER J. G., 1893 – Foraminiferen aus Meeresgrundproben,
acting on the microfauna in mangrove swamps. gelothet von 1874 bis 1876 von S. M. Sch. “Gazelle.”
The case of foraminiferal assemblages in French Guiana. Abhandlungen der Bayerischen Akademie der Wissenschaften,
Estuarine, Coastal and Shelf Science, 55 (4): 509-533. Mathematisch- Physikalischen Klasse, 18: 193-458.
DEBENAY J.-P., MILLET B., ANGELIDIS M. O., 2005 – Relationships between EHRENBERG C. G., 1839 – Über die Bildung der Kreidefelsen
foraminiferal assemblages and hydrodynamics in the Gulf of Kalloni, und des Kreidemergels durch unsichtbare Organismen.
Greece. Journal of Foraminiferal Research, 35: 327-343. Physikalisch abhandlungen der königlichen Akademie
DEBENAY J.-P., DELLA PATRONA L., HERBLAND A., GOGUENHEIM H., 2009a – der Wissenschaften zu Berlin: 59-147.
Colonization of coastal environments by Foraminifera: EHRENBERG C. G., 1840 – Über noch jetzt zahlreich lebende Thierarten
Insight from shrimp ponds in New Caledonia. der Kreidebildung und den Organismus der Polythalamien.
Journal of Foraminiferal Research, 39: 249-266. Physikalische Abhandlungen der Königlichen Akademie
DEBENAY J.-P., DELLA PATRONA L., HERBLAND A., GOGUENHEIM H., 2009b – der Wissenschaften zu Berlin, 1839: 81-147.
The impact of Easily Oxidized Material (EOM) on the meiobenthos: EHRENBERG C. G., 1843 – Verbreitung und Einfluss des mikroskopischen
Foraminifera abnormalities in Shrimp ponds of New Caledonia; Lebens in Süd- und Nord-Amerika. Physikalische Abhandlungen
implications for environment and paleoenvironment survey. der Königlichen Akademie der Wissenschaften zu Berlin, 1841:
Marine Pollution Bulletin, 59: 323-335. 291- 446.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 333
EHRENBERG C. G., 1844 ? Ueber Spirobotrys, eine neue physiologisch GALLOWAY J. J., WISSLER S. G., 1927 – Pleistocene Foraminifera
merkwurdige Gattung von Polythalamien. Königliche Preussischen from the Lomita quarry, Palos Verdes Hills, California.
Akademie der Wissenschaften zu Berlin Bericht: 245-248. Journal of Paleontology, 1: 35-87.
GAMBINI A., 1958 – Contribution à l’étude des sables
à Marginopora vertebralis de la Nouvelle-Calédonie.
FARÍAS J. R., 1977 – Murrayinella: taxa nuevo para la ciencia D.E.S., Laboratoire de géologie, université de Montpellier : 1-103.
de foraminiferos del Reciente de Agua Somera. GAMBINI A., 1959 – Sur la composition de quelques sables coquilliers
Revista Española de Micropaleontología, 9: 343-345. à foraminifères des lagons de la Nouvelle-Calédonie.
FERNANDEZ J.-M., OUILLON S., CHEVILLON C., DOUILLET P., FICHEZ R., Bulletin de la Société géologique de France, 1 : 431-433.
LE GENDRE R., 2006 – A combined modelling and geochemical GANACHAUD A., KESSLER W., WIJFFELS S., RIDGWAY K.,
study of the fate of terrigenous inputs from mixed natural CAI W., HOLBROOK N., BOWEN M., SUTTON P., QIU B.,
and mining sources in a coral reef lagoon (New Caledonia). TIMMERMANN A., ROEMMICH D., SPRINTALL J., CRAVATTE S.,
Marine Pollution Bulletin, 52: 320-331. GOURDEAU L., AUNG T., 2007 – Southwest Pacific Ocean
FICHTEL L. von, MOLL J. P. C. von, 1798 – Testacea microscopica, Circulation and Climate Experiment (SPICE) –
aliaque minuta ex generibus Argonauta et Nautilus, Part I. Scientific Background. CLIVAR Publication Series No. 111,
ad naturam picta et descripta (Microscopische und NOAA OAR Special Report:1-37.
andere kleine Schalthiere aus den geschlechtern Argonaute GERMERAAD J. H., 1946 – “Geology of central Ceram”.
und Schiffer). Vienna. Camesina. In Rutten L., Hotz W. (eds): Geological, petrographical
FICHTEL L. von, MOLL J. P. C. von, 1803 – Testacea microscopica aliaque and paleontological results of explorations carried out
minuta ex generibus Argonauta et Nautilus ad naturam delineata from 1917 till 1919 in the Island of Ceram,Bussy, J.H.,
et descripta. Cum 24 tabulis æri incisis. - Mikroskopische und Amsterdam: 7-135.
andere kleine Schalthiere aus den Geschlechtern Argonaute GLAÇON G., LYS M., 1968 – Note préliminaire à une révision
und Schiffer, nach der Natur gezeichnet und beschrieben. des espèces de Monspeliensina, nouveau genre de foraminifère
Mit 24 Kupfertafeln. - pp. j-xjj [= 1-12], [1-4], 1-123, [1], accompagnant la transgression Miocène dans le Languedoc.
Tab. 1-24. Wien. (Camesina). Compte Rendu Hebdomadaire des Séances de l’Acadamie
FINLAY H. J., 1939 – New Zealand foraminifera; des Sciences, Paris, 267 : 2302-2305.
Key species in stratigraphy - No. 3. Transactions and Proceedings GLOVER E., TAYLOR J., WHITTAKER J., 2003 –
of the Royal Society of New Zealand, 69: 309-329. “Distribution, abundance and foraminiferal diet
FINLAY H. J., 1940 – New Zealand foraminifera; of an intertidal scaphopod, Laevidentalium lubricatum,
Key species in stratigraphy - No. 4. Transactions and Proceedings around the Burrup Peninsula, Dampier, Western Australia”.
of the Royal Society of New Zealand, 69: 448-472. In Wells F. E., Walker D. I., Jones D. S. (eds):
FISCHER M., DEWITTE B., MAÎTREPIERRE L., 2004 – A non-linear The Marine Flora and Fauna of Dampier,
statistical downscaling model: El Niño/Southern Oscillation impact Western Australia - Western Australian Museum, Perth: 225-240.
on precipitation over New Caledonia, Geophysical Research Letter, GOËS A., 1882 – On the reticularian Rhizopoda
31, L16204, doi:10.1029/2004GL020112. of the Caribbean Sea. Kongelike Svenska Vetenskaps-
FLINT J. M., 1899 – Recent Foraminifera. A descriptive catalogue Akademiens Handllingar, 19 (4): 1-151.
of specimens dreged by the U.S. Fish Commission steamer Albatross. GOËS A., 1894 – A synopsis of the Arctic and Scandinavian recent
Report of the U.S. National Museum for 1897, 1: 249-349. marine Foraminifera hitherto discovered. Kongelike Svenska
FOLLAND C. K., RENWICK J. A., SALINGER M. J., MULLAN A. B., 2002 – Vetenskaps- Akademiens Handllingar, 25 (9): 1-127.
Relative influences of the interdecadal Pacific Oscillation GOËS A., 1896 – “The Foraminifera”.
and ENSO on the South Pacific Convergence Zone. In: Reports on the dredging operations off the West Coast
Geophysical Research Letter, 29, doi:10.1029/2001GL014201. of Central America to the Galapagos, to the West Coast of Mexico,
FORNASINI C., 1896 – Ottavo Contributo alla conoscenza and in the Gulf of California, in charge of Alexander Agassiz,
della microfanna terziarii. Italiana - Memorie dell’Accademia carried on by the U.S. Fish Commission Steamer ”Albatross”,
della Scienze dell’Istituto di Bologna 5/6: 1-6. during 1891, Lieut. Commander Z.L. Tanner U.S.N.,
FORNASINI C., 1908 – Illustrazione di specie orbignyane commanding, Bulletin of the Museum of Comparative Zoology
di Nodosaridi, di Rotalidi e d’altri foraminiferi. at Harvard College, 29 (1): 1-103.
Memorie della Reale Accademie della Scienze dell’Istituto GOLDSTEIN S. T., CORLISS J. O., 1994 –
di Bologna, Scienze Naturali, 6 (5): 41-54. Deposit feeding in selected deep-sea and shallow-water benthic
FORSKAL P., 1775 – Descriptiones animalium. Copenhagen. foraminifera. Deep-Sea Research, 41: 229-441.
Hauniae, Carsten Niebuhr. GOODAY A. J., LEVIN L. A., LINKE P., HEEGER T., 1992 –
FUJITA K., HALLOCK P., 1999 – A comparison of phytal substrate “The role of benthic foraminifera in deep-sea food webs
preferences of Archaias angulatus and Sorites orbiculus and carbon cycling”. In Rowe G. T., Parient V. (eds):
in mixed macroalgal-seagrass beds in Florida Bay. Deep-sea Food Chains and the global Carbon Cycle,
Journal of Foraminiferal Research, 29: 143-151. Kluwer, Dordrecht: 63-91.
GRONOVIUS L. T., 1781 – Zoophylacium Gronovianum vol. 3.
Lugduni Batavorum, T. Haak, Leiden, Netherlands: 241-380.
GALLOWAY J. J., MORREY M., 1929 – GUERIN-MENEVILLE F. E., 1843 – « Crustacés ».
A Lower Tertiary foraminiferal fauna from Manta, Ecuador. In: Iconographie du règne animal de G. Cuvier. Mollusques,
American Paleontological Bulletin, 15 (55): 7-57. Paris, J. B. Baillière, vol. 3.
334 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
HADLEY W. H.Jr, 1934 – HAYWARD B. W., HOLZMANN M., GRENFELL H. R., PAWLOWSKI J.,
Some Tertiary Foraminifera from the North Coast of Cuba. TRIGGS C. M., 2004 – Morphological distinction
Bulletin of American Paleontology, 20 (70A): 1-40. of molecular types in Ammonia - towards a taxonomic revision
of the world’s most commonly misidentified foraminifera.
HAIG D. W., 1988 – Miliolid foraminifera Marine Micropaleontology, 50: 237-271.
from inner neritic and mud facies of the Papuan Lagoon.
Journal of Foraminiferal Research, 18: 203-236. HAYWARD B. W., GRENFELL H. R., SABAA A. T., NEIL H. L.,
BUZAS M. A., 2010 – Recent New Zealand deep-water benthic
HAIG D. W., 1997 – foraminifera: taxonomy, ecologic distribution, biogeography,
Foraminifera from Exmouth Gulf, Western Australia. and use in paleoenvironmental assessment. Institute of Geological
Journal of the Royal Society of Western Australia, 80: 263-280. and Nuclear Sciences monographs, 26: 1-363.
HANSEN H. J., 1970 – Danian Foraminifera from Nûgssuaq, HEDBERG H. D., 1937 – Foraminifera of the Middle Tertiary
West Greenland. Meddelelser om Grønland 193 (2). Carapita formation of northeastern Venezuela.
Grønlands Undersøgelse, 93: 1-132. Journal of Paleontology, 11: 661-697.
HANSEN H. J., REVETS, R., 1992 – A revision and reclassification HEMLEBEN C. H., ANDERSON O. R., BERTHOLD W., SPINDLER M., 1986 –
of the Discorbidae, Rosalinidae, and Rotaliidae. Calcification and chamber formation in Foraminifera -
Journal of Foraminiferal Research, 22: 166-180. a brief overview. In Leadbeater B. S. C., Riding R. (eds) :
HANZAWA S., 1957 – Cenozoic foraminifera of micronesia. Biomineralization in lower plants and animals.
Memoir of the Geological Society of America, 66: 1-165. Systematics Association, Special Vol. 30 : 237-249.
HATTA A., 1996 – Pyrgo rasheedi, n. sp., Foraminifera. HERON-ALLEN E., EARLAND A., 1911 – On the recent and fossil
South Pacific Study, 17: 21-28. foraminifera of the shore-sands of Selsey Bill, Sussex,
IV. Journal of the Royal Microscopy Society: 436-448.
HATTA A., UJIIÉ J., 1992a – Benthic foraminifera from Coral Seas
HERON-ALLEN E., EARLAND A., 1912 – On some foraminifera
between Ishigaki and Iriomote Islands, Southern Ryukyu Island Arc,
from the North Sea, etc., dredged by the Fisheries Cruiser
Northwestern Pacific, Part 1, Systematic descriptions of Textulariina
“Goldseeker” (International North Sea Investigations-Scotland).
and Miliolina. Bulletin of the College of Science, University of the
On some new Astrorhizidae and their shell-structure.
Ryukyus, 53: 49-119.
Journal of the Royal Microscopical Society, London: 382-389.
HATTA A., UJIIÉ J., 1992b – Benthic foraminifera from Coral Seas
HERON-ALLEN E., EARLAND A., 1913 – Clare Island survey, Part 64,
between Ishigaki and Iriomote Islands, Southern Ryukyu Island Arc,
Foraminifera. Proceedings of the Royal Irish Academy, 31: 1-188.
Northwestern Pacific, Part 2, Systematic descriptions of Rotalina.
Bulletin of the College of Science, University of the Ryukyus, 54: HERON-ALLEN E., EARLAND A., 1914 – Foraminifera
163-287. of the Kerimba Archipelago (Portuguese East Africa) Part I.
Transactions of the Zoological Society of London, 20: 363-390.
HAYNES J. R., 1973 – Cardigan Bay foraminifera
(cruises of the R. V. Antur, 1962-1964). Bulletin of the British HERON-ALLEN E., EARLAND A., 1915 – The foraminifera
Museum (Natural History), Zoology. Supplement 4: 1-245. of the Kerimba Archipelago (Portuguese East Africa). Part II.
Transactions of the Zoological Society of London, 20: 543-795.
HAYNES J. R., 1981 – Foraminifera.
HERON-ALLEN E., EARLAND A., 1916 – The foraminifera
London, Macmillan Publishers.
of the west coast of Scotland. Transactions of the Linnean
HAYWARD B. W., 1990 – Taxonomy, paleobiogeography Society of London, Zoology, series 2, 11: 197-299.
and evolutionary history of the Bolivinellidae (Foraminiferida).
HERON-ALLEN E., EARLAND A., 1922 – Protozoa, Part 2, Foraminifera.
New Zealand Geological Survey Paleontological Bulletin, 63: 1-132.
Natural history reports of the British Antarctic (“Terra Nova”)
HAYWARD B. W., BRAZIER R. C., 1980 – Expedition, 1910, 6 (2): 25-268.
Taxonomy and distribution of present-day Bolivinella. HERON-ALLEN E., EARLAND A., 1924 –
Journal of Foraminiferal Research, 10: 102-116. The Foraminifera of Lord Howe Island, South Pacific.
HAYWARD B. W., BUZAS M. A., 1979 – Taxonomy and paleoecology Journal of the Linnean Society, Zoology, 35: 599-647.
of early Miocene benthic foraminifera of northern New Zealand HERON-ALLEN E., EARLAND A., 1928 – On the Pegididae,
and the north Tasman Sea. a new family of foraminifera. Journal of the Royal Microscopical
Smithsonian Contributions to Paleobiology, 36: 154. Society of London, series 3, 48: 283-299.
HAYWARD B. W., HOLLIS C., 1994 – Brackish foraminifera in New Zealand: HERON-ALLEN E., EARLAND A., 1930 – Some new foraminifera
A taxonomic and ecologic review. Micropaleontology, 40: 185-22. from the South Atlantic, Part 3. Journal of the Royal Microscopical
HAYWARD B. W., TRIGGS, C. M., 1994 – Computer analysis Society, London, 50: 38-45.
of benthic foraminiferal associations in a tidal New Zealand inlet. HERON-ALLEN E., EARLAND A., 1932a – Some new foraminifera
Journal of Micropalaeontology, 13: 103-117. from the South Atlantic; IV. Four new genera from South Georgia.
HAYWARD B. W., HOLLIS C. J., GRENFELL H. R., 1997 – Journal of the Royal Microscopical Society of London, 52: 253-261.
Recent Elphidiidae (Foraminiferida) of the South-west Pacific HERON-ALLEN E., EARLAND A., 1932b – Foraminifera Part 1.
and fossil Elphidiidae of New Zealand. Institute of Geological The ice free area of the Falkland Islands and adjacent seas.
and Nuclear Sciences monographs, 16: 1-170. Discovery Reports, 4: 291-460.
HAYWARD B. W., GRENFELL H. R., RIED C. M., HAYWARD K. A., 1999 – HICKSON S. J., 1911 – On Polytrema and some allied genera.
Recent New Zealand shallow-water benthic foraminifera: A study of some sedentary foraminifera based mainly
taxonomy, ecologic distribution, biogeography, and use on a collection made by Prof. Stanley Gardiner in the Indian Ocean.
in paleoenvironmental assessment. Institute of Geological Transactions of the Linnean Society of London, Zoology, series 2,
and Nuclear Sciences monographs, 21: 1-264. 14: 443-462.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 335
HOBSON E. S., CHESS J. R., 1973 – Feeding oriented movements HOWCHIN W., 1889 – The foraminifera of the Older Tertiary
of the atherinid fish Pranesus pinguis at Majuro atoll, of Australia (n° 1, Muddy Creek, Victoria). Transactions and
Marshall Islands. Fishery Bulletin, 71: 777-786. Proceedings of the Royal Society of South Australia, 12: 1-20.
HOFKER J., 1927 – The Foraminifera of the Siboga Expedition. Part 1. HROMIC T., ANDRADE C., RAMÍREZ I., VIDAL S., 2004 –
Families Tinoporidae, Rotaliidae, Nummulitidae, Amphisteginidae. A new record of Spirillina tuberculata Brady
Siboga Expeditie, Monographie IVa. Leiden. E. J. Brill: 1-78. (Protozoa Foraminiferida) in Chilian waters.
HOFKER J., 1930 – The foraminifera of the Siboga Expedition; Anales del Instituto de la Patagonia (Chile), 32: 35-41.
Part 2, families Astrorhizidae, Rhizamminidae, Reophacidae, HUGHES G. W., 1977 – Recent Foraminifera from the Honiara Bay area,
Anomalinidae, Peneroplidae. Siboga Expeditie, Monographie IVa. Salomon Islands. Journal of Forarniniferal Research, 7: 45-57.
Leiden. E. J. Brill: 79-170. HUGHES G. W., 1988 – Modern bathyal agglutinating foraminifera
HOFKER J., 1951 – The foraminifera of the Siboga Expedition; Part 3. from the Vella Gulf and Blanche Channel, New Georgia,
Siboga Expeditie, Monographie IVb. Leiden. E. J. Brill: 1-513. Solomon Islands, southwest Pacific.
HOFKER J., 1956 – Foraminifera Dentata-Foraminifera of Santa Cruz Journal of Foraminiferal Research, 18: 304-310.
and Thach Island, Virginia Archipelago West Indies.
Copenhagen University, Zoological Museum, Spolia, 15: 1-237.
HOFKER J., 1976 – Further studies on Caribbean foraminifera. ISHIZAKI K., 1939 – On the foraminifera of the so-called Upper Marine
Studies on the Fauna of Curaçao and other Caribbean Fossil Beds in the vicinity of Kwansai, Sintiku Prefecture, Taiwan.
Islands, 49: 1-256. Taiwan Tigaku Kizi (Geological notes on Formosa), 10: 103-124.
HOFKER J., 1978 – Biological results of the Snellius Expedition: (Japanese with English abstract).
The foraminifera collected in 1929 and 1930 in the eastern part ISHIZAKI K., 1943 – New species of Neogene, Pleistocene
of the Indonesian Archipelago. Zoologische Verhandelingen and Recent Foraminifera of Japanese Empire. (I). Transactions
Rijksmuseum van Natuurlijke Historie te Leiden, 161: 1-69. of the Natural History Society of Formosa, 33 (233): 19-23.
HOFKER J., 1983 – Zoological exploration of the continental shelf ISHIZAKI K., 1948 – Six new fossil species of Streblus from eastern Asia.
of Surinam: The Foraminifera of the shelf of Surinam Transactions of the Natural History Society of Formosa, 21: 55-66.
and the Guyanas. Zoologische Verhandelingen, 201: 3-75, 17 pls.
HÖGLUND H., 1947 – Foraminifera in the Gullmar Fjord
and the Skagerak. Zoologiska Bidrag Från Uppsala, 26: 1-328.
HÖGLUND H., 1948 – New names for four homonym species described JAVAUX E., SCOTT D. B., 2003 – Illustration of recent benthic
in ‘Foraminifera in the Gullmar Fjord and the Skagerak’. foraminifera in Bermuda and remarks on species distribution.
Contributions from the Cushman Laboratory for Foraminiferal Palaeontologica electronica, 6, 29 p.
Research, 24: 45-46. (http://www.earthsci.carleton.ca/paleo/2003_1/benthic/issue1_03.htm).
HOHENEGGER J., 1994 – Distribution of living larger foraminifera NW JONES R. W., 1994 – The Challenger Foraminifera.
of Sesoko-jima, Okinawa, Japan. Marine Ecology, 15: 291-334. Oxford University Press, Oxford: 1-149.
HOHENEGGER J., YORDANOVA E., HATTA A., 2000 – JONES T. R., 1895 – The Crag Foraminifera. Part 2.
Remarks of West Pacific Nummulitidae (Foraminifera). Monograph of the Palaeontographical Society London, 230 (49):
Journal of Foraminiferal Research, 30: 3-28. 73-210, pls 5-7.
HOLZMANN M., PILLER W., ZANINETTI L., FENNER R., MARTINI R., JONES T. R., 1896 – The Crag Foraminifera. Part 3.
SERANDREIBARBERO R., PAWLOWSKI J., 1998 – Monograph of the Palaeontographical Society London, 234 (50):
Molecular versus morphologic variability in Ammonia spp. 211-314.
(Foraminifera, Protozoa) from the Lagoon of Venice, Italy. JONES T. R., CHAPMAN F., 1896 – On the Fistulose Polymorphinæ,
Revue de Micropaléontology, 41: 59-69. and on the Genus Ramulina. Journal of the Linnean Society
HOLZMANN M., HABURA A., GILES H., BOWSER S. S., PAWLOWSKI J., 2003 – of London, Zoology, 25 (165): 496–516.
Freshwater Foraminiferans Revealed by Analysis of Environmental JONES T. R., PARKER W. K., 1860 – On the Rhizopodal fauna
DNA Samples. Journal of Eukaryotic Microbiology, 50: 135-139. of the Mediterranean, compared with that of the Italian
HOTTINGER L., 1977 – Distribution of larger Peneroplidae, and some older Tertiary deposits.
Borelis and Nummulitidae in the Gulf of Elat, Red Sea. Utrecht. Quarterly Journal of the Geological Society of London, 16: 292-307.
Micropaleontological Bulletins, 15: 35-110.
HOTTINGER L., 1978 – “Comparative Anatomy of Elementary Shell
Structures in Selected Larger Foraminifera”. In Hedley R. H., KAMINSKI M. A., 2000 – “The New and Reinstated Genera
Adams C. G. (eds): Foraminifera. Volume 3. Academic Press: 203-266. of Agglutinated Foraminifera published between 1986 and 1996”.
HOTTINGER L., 1986 – Construction, structure and function In Hart M. B., Kaminski M. A., Smart C. W. (eds): Proceedings
of foraminiferal shells. The Systematics Association, of the Fifth International Workshop on Agglutinated Foraminifera,
Special Volume 30: 219-235. Grzybowski Foundation Special Publication, 7: 185-219.
HOTTINGER L., 2006 – Illustrated glossary of terms KAMINSKI M., 2004 – “The year 2000 classification of the agglutinated
used in foraminiferal research. Notebooks on Geology (Brest), foraminifera”. In Bubík M., Kaminski M. A. (eds): Proceedings
Memoir 2006/2, 126 p. 83 figs. of the Sixth International Workshop on Agglutinated Foraminifera.
HOTTINGER L., HALICZ E., REISS Z., 1993 – Recent Foraminifera Grzybowski Foundation Special Publication, 8: 237-255.
from the Gulf of Aqaba, Red Sea. Academia Scientiarum et Artium KARRER F., 1865 – Die Foraminiferen-Fauna des tertiären Grünsandsteines
Slovenica, Classis IV: Historia Naturalis, Paleon toloski Institut der Orakei-Bay bei Auckland. Novara Expedition 1857-1859.
ivana Rkovka 33/3: 1-179. Geologisches Theil, 1: 69-86.
336 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
KARRER. F., 1868 – Die Miocene Foraminiferenfauna von Kostej im Batat. LE CALVEZ J., 1947 – Entosolenia marginata, foraminifère apogamique
Sitzungsberichte der Mathematisch-Naturwissenschaftlichen ectoparasite d’un autre foraminifére Discorbis vilardeboanus.
Klasse der Kaiserlichen Akademie der Wissenschaften zu Wien, Compte Rendus de l’Académie des Sciences, Paris, 224 : 1448-1450.
58, 121-93. LE CALVEZ Y., 1958 – Les foraminifères de la mer celtique.
KOHL B., 1985 – Early Pliocene benthic foraminifers from the Salina Revue des Travaux de l’Institut des Pêches Maritimes, 22 : 147-209.
basin, southeastern Mexico. Bulletins of American Paleontology, LE CALVEZ Y., 1974 – Révision des foraminifères de la collection d’Orbigny.
88 (322): 1-173. I. Foraminifères de îles Canaries. Cahiers de Micropaléontologie.
KORNFELD M. M., 1931 – Recent littoral foraminifera from Texas 1974 (2) : 1-108.
and Louisiana. Contributions from the Department of Geology LE CALVEZ Y., 1977 – Révision des foraminifères de la collection d’Orbigny.
of Stanford University, 1: 77-101. II - Foraminifères de l’île de Cuba. Cahiers de Micropaléontologie,
1977 (1) : 1-127.
LE CALVEZ J., LE CALVEZ Y., 1958 – Répartition des foraminifères
LACROIX E., 1928 – De la présence d’une faune d’Astrorhizidés dans la baie de Villefranche. I Miliolidae.
tubulaires dans des fonds littoraux de Saint-Raphaël à Monaco. Annales de l’Institut Océanographique, Paris, 35 : 159-234.
Bulletin de l’Institut Océanographique Monaco, 527 : 61-144. LE CALVEZ Y., SALVAT B., 1980 – Foraminifères des récifs
LACROIX E., 1930 – Les Lituolides de plateau continental et lagons coralliens de Mooréa, îles de la Société.
méditerranéen entre Saint-Raphael et Monaco. Cahiers de Micropaléontologie, 1980 (4) : 1-15.
Bulletin de l’Institut Océanographique Monaco, 549 : 547-550. LE ROY P., CABIOCH G., MONOD B., LAGABRIELLE Y., PELLETIER B.,
LACROIX E., 1938 – Révision du genre Massilina. FLAMAND B., 2008 – Late Quaternary history of the Noumea lagoon
Bulletin de l’Institut Océanographique Monaco, 754 : 1-11. (New Caledonia, S-W Pacific) as depicted by seismic stratigraphy
and multibeam bathymetry. A modern model of tropical rimmed
LALICKER C. G., 1935 – Two new foraminifera of the genus Textularia.
shelf. Palaeogeography Palaeoclimatology Palaeoecology, 270:
Smithsonian Miscellaneous Collections, 91 (22): 1-2.
29-45.
LALICKER C. G., MCCULLOCH I., 1940 – Some Textulariidae of the Pacific
LEE J. J., MULLER W. A., 1973 – Trophic dynamics and niches
Ocean. Allan Hancock Pacific Expeditions, 6 (6): 115-143.
of salt marsh foraminifera. American zoology, 13: 215-223.
LAMARCK J. B., 1804 – Suite des mémoires sur les fossiles
des environs de Paris. Annales du Muséum national d’histoire LEE J. J., BURNHAM B., CEVASCO M. E., 2004 –
naturelle, 5 : 349-357. A new modern soritid foraminifer, Amphisorus saurensis n. sp.,
from the Lizard Island Group (Great Barrier Reef, Australia).
LAMARCK J. B., 1816 – Histoire naturelle des animaux invertébrés 2. Micropaleontology, 50: 357-368.
Paris, Verière.
LEE J. J., MCENERY M., PIERCE S., FREUDENTHAL H. D.,
LAMB J. L., MILLER T. H., 1984 – Stratigraphic significance of Uvigerinid MULLER W. A., 1966 – Tracer experiments in feeding littoral
foraminifers in the western hemisphere. The University of Kansas foraminifera. Journal of Protozoology, 16: 659-670 .
Paleontological Contribution, 66: 1-98, 40 pls.
LEE J. J., PAWLOWSKI J., DEBENAY J.-P., WHITTAKER J. E.,
LAMBERT B., GOMEZ A.-M., MATHIEU R., 1991 – BANNER F. T., GOODAY A. J., TENDAL O., HAYNES J., FABER W. W., 2000 –
De la production planctonique au sédiment. “Class Foraminifera”. In Lee J. J., Leedale G. F., Bradbury P. (eds):
Documents et Travaux de l’IGAL, Paris, 15 : 109-126. An Illustrated Guide to the Protozoa, second edition, Society
LANGER M., 2011 – of Protozoologists (Allen Press, Lawrence Kansas): 877-951.
http://www.paleontology.uni-bonn.de/en_/micropaleontology/index.html
LEROY L. W., 1944 – Miocene foraminifera from Sumatra and Java,
LANGER M. R., 1992 – New Recent foraminiferal genera Nederlands East Indies, Part I, Miocene foraminifera of central
and species from the lagoon at Madang, Papua New Guinea. Sumatra, Nederlands East Indies. Colorado School of Mines
Journal of Micropalaeontology, 11: 85-93. Quarterly, 39 (3): 1-69.
LANGER M. R., GEHRING C. A., 1993 – Bacteria farming: LEVY A., MATHIEU R., POIGNANT A., ROSSET-MOULINIER M.,
a possible feeding strategy of some smaller, motile foraminifera. UBALDO M. L., LEBREIRO S., 1995 – Foraminifères actuels
Journal of Foraminiferal Research, 23: 40-46. de la marge continentale portugaise. Inventaire et distribution.
LANGER M. R., LIPPS J. H., MORENO G., 1995 – Predation on foraminifera Memorias Instituto Geologico e mineiro, Lisboa, 32 : 1-116.
by the dentaliid deep-sea scaphopod Fissidentalium megathyris. LEVY A., POIGNANT A., ROSSET-MOULINIER M., ROUVILLOIS A., 1975 –
Deep-Sea Research, 42: 849-857. Sur quelques foraminifères actuels des plages de Dunkerque
LANKFORD R. R., PHLEGER F. B., 1973 – Foraminifera et des environs: néotypes et espèces nouvelles.
from the nearshore turbulent zone, western North America. Revue de Micropaléontologie, 17 : 171-181.
Journal of Foraminiferal Research, 3: 101-132. LINNAEUS C., 1758 – Systema naturae per regna tria naturae,
LARSEN A. R., 1976 – Studies of recent Amphistegina, secundum classes, ordines, genera, species, cum characteribus,
taxonomy and some ecological aspects. differentiis, synonymis, locis. G. Engelmann, 10: 1-824.
Israel Journal of Earth Sciences, 25: 1-26. LINNAEUS C., 1766 – Systema naturae sive regna tria naturae,
LARSEN A. R., 1978 – Phylogenetic and paleobiogeographic trends secundum classes, ordines, genera, species, cum characteribus,
in the foraminiferal genus Amphistegina. differentiis, synonymis, locis. Laurentii Salvii, Holmiae.
Revista Española de Micropaleontología, 10: 217-243. 12th ed. v. 1 (pt 1):1-532.
LE BORGNE R., DOUILLET P., FICHEZ R., TORRÉTON J.-P., 2010 – LIPPS J. H., 1983 – “Biotic interactions in benthic foraminifera”.
Hydrography and plankton temporal variabilities at different time In Tevesz M. J. S., Mc Call P. L. (eds): Biotic interactions
scales in the southwest lagoon of New Caledonia: A review. in recent and fossil benthic communities, Plenum Press,
Marine Pollution Bulletin, 61 (7-12): 297-308. New- York: 331-373.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 337
LIPPS J. H., 1988 – Predation on foraminifera by coral reef fish: MARGEREL J.-P., 2009 – Les foraminifères benthiques des Faluns
taphonomic and evolutionary implications. Palaios, 3: 1-12. du Miocène moyen du Blésois (Loir-et-Cher) et de Mirebeau (Vienne)
LIPPS J. H., Ronan T. E. Jr, 1974 – dans le centre-ouest de la France. Geodiversitas, 31 (3) : 577-621.
Predation on foraminifera by the polychaete worm Diopatra. MATEU G., 1969 – Foraminiferos del contenido gástrico
Journal of Foraminiferal Research, 4: 139-143. del Spatangus purpureus O. F. Muller y su degradación
LOBEGEIER M., 2001 – Foraminiferal assemblages protoplasmática a través del aparato digestivo de este equinido.
and their bulk contribution to carbonate sediment, Boletín de la Sociedad de Historia Natural de Baleares, 15: 75-84.
Green Island Reef, Great Barrier Reef Province. MATOBA Y., 1970 – Distribution of Recent shallow benthic foraminifera
Unpublished Ph.D. Thesis, James Cook University, Townsville, 457 p. of Matsushima Bay, Miyagi Prefecture, Northeast Japan.
LOBEGEIER M. K., 2002 – Benthic foraminifera of the family Tohoku University Science Reports, Sendai, Japan, 2nd series
Calcarinidae from Green Island Reef, Great Barrier Reef Province. (Geology), 42: 1-85.
Journal of Foraminiferal Research, 32: 201-216. MATSUNAGA T., 1963 – Benthonic smaller foraminifera from the oil fields
LOBEGEIER M. K., SEN GUPTA B. K., 2008 – of northern Japan. Tohoku University Science Reports, Sendai,
Foraminifera of hydrocarbon seeps, Gulf of Mexico. Japan, 2nd series (Geology), 35: 65-122, 29 pls.
Journal of Foraminiferal Research, 38: 93-116. MCCULLOCH I., 1977 – Qualitative observations on Recent foraminiferal
LOEBLICH A. R. Jr, TAPPAN H., 1953 – Studies of Arctic foraminifera. tests with emphasis on the Eastern Pacific parts I, II, III.
Smithsonian Miscellaneous Collections, 121: 1-150. University of Southern California, Los Angeles, 676 p., 200 pls.
LOEBLICH A. R. Jr, TAPPAN H., 1955 – MCCULLOCH I., 1981 – Qualitative observations on Recent foraminiferal
Revision of some recent foraminiferal genera. tests. Part IV, with emphasis on the Allan Hancock Atlantic
Smithsonian Miscellaneous Collections, 128: 1-37. Expedition collections. University of Southern California,
Los Angeles, 362 p., 72 pls.
LOEBLICH A. R. Jr, TAPPAN H., 1957 – Eleven new genera of Foraminifera.
United States National Museum Bulletin, 215: 223-232. MCLEAN J. D., 1956 – The foraminifera of the Yorktown
Formation in the York-James. Peninsula of Virginia
LOEBLICH A. R. Jr, TAPPAN H., 1964 – “Protista 2: Sarcodinia,
with notes on the associated Mollusks.
chiefly ‘Thecamoebians’ and Foraminiferida”.
Bulletins of the American Paleontology, 36: 261-394.
In: Treatise on Invertebrate Palaeontology,
Geological Society of America and University of Kansas Press, MEISTERFELD R., HOLZMANN M., PAWLOWSKI J., 2001 – Morphological and
Lawrence, Kansas 2 vol.: 1-900. molecular characterization of a new terrestrial allogromiid species:
Edaphoallogromia australica gen. et spec. nov. (Foraminifera)
LOEBLICH A. R. Jr, TAPPAN H., 1985 –
from northern Queensland (Australia). Protist, 152: 185-192.
Some new and redefined genera of agglutinated foraminifera II.
Journal of Foraminiferal Research, 15: 175-217. MIKHALEVICH V. I., 1977 – New species of Foraminifera
of the North-Western Coast of Africa, Issledovanija fauny morej.
LOEBLICH A. R. Jr, TAPPAN H., 1988 – Foraminiferal genera
Zoological institute Academy of Sciences of the USSR, 21 (29): 5-9.
and their classification. Van Nostrand Reinhold Company,
New York, 1, 970 p., 2, 847 pl. MIKHALEVICH V. I., 1983 – The bottom foraminifera from the shelves
of the Tropical Atlantic. Zoological Institute of USSR Academy
LOEBLICH A. R. Jr, TAPPAN H., 1992 – “Present status of foraminiferal
of Sciences, Leningrad, 247 p. (in Russian).
classification”. In Takayanagi Y., Saito T. (eds): Studies in Benthic
Foraminifera, Proceedings of the Fourth International MILLETT F. W., 1894 – The Foraminifera of the Pliocene beds of St. Erth.
Symposium on Benthic Foraminifera, Sendai, 1990 Transactions of the Royal Geological Society of Cornwall, 11:
(Benthos‘90), Tokai University Press, Tokyo: 93-102. 655–661.
LOEBLICH A. R. Jr, TAPPAN H., 1994 – Foraminifera of the Sahul Shelf. MILLETT F. W., 1898a – Report on the recent foraminifera
Cushman Foundation for Foraminiferal Research, of the Malay Archipelago collected by Mr A. Durrand, F.R.M.S.
Special Publication 31: 1-661. Journal of the Royal Microscopical Society, 1898: 258-269.
LUTZE G. F., 1974 – Benthische Foraminiferen MILLETT F. W., 1898b – Report on the recent foraminifera
in Oberflächen-Sedimenten des Persischen Golfes. of the Malay Archipelago collected by Mr A. Durrand, F.R.M.S, Part II.
Teil 1: Arten. Meteor Forschungsergebnisse, Deutsche Journal of the Royal Microscopical Society, 1898: 497-513.
Forschungsgemeinschaft, Reihe C Geologie und Geophysik, MILLETT F. W., 1898c – Report on the recent foraminifera of the Malay
Gebrüder Bornträger, Berlin, Stuttgart, C17, 1-66. Archipelago collected by Mr A. Durrand, F.R.M.S, Part III.
Journal of the Royal Microscopical Society, 1898: 607-614.
MILLETT F. W., 1898d – Additions to the list of foraminifera
MAES C., VEGA A., SUDRE J., 2007 – from the St. Erth Clay. Transactions of the Royal Geological
Society of Cornwall, 12: 174-176.
“Hydroclimatic conditions in the southwest Pacific Ocean”.
In Payri C., Richer de Forges B. (eds): Compendium of marine MILLETT F. W., 1899a – Report on the recent foraminifera of the Malay
species of New Caledonia, IRD Nouméa, Documents Scientifiques Archipelago collected by Mr A. Durrand, F.R.M.S, Part V.
et Techniques, 117, seconde édition: 51-62. Journal of the Royal Microscopical Society, 1899: 356-365.
MARGEREL J.-P., 1981 – Espèces nouvelles de foraminifères MILLETT F. W., 1899b – Report on the recent foraminifera of the Malay
de la baie de Saint-Vincent (Nouvelle-Calédonie). Archipelago collected by Mr A. Durrand, F.R.M.S, Part VI.
Cahiers de Micropaléontologie, 4 : 67-72. Journal of the Royal Microscopical Society, 1899: 556-564.
MARGEREL J.-P., 1984 – Les foraminifères de la baie de Saint-Vincent MILLETT F. W., 1900a – Report on the recent foraminifera of the Malay
(Nouvelle-Calédonie). Unpublished memoir of the laboratoire Archipelago collected by Mr A. Durrand, F.R.M.S, Part VIII.
de géologie historique, université de Nantes, 121 p., 39 pls. Journal of the Royal Microscopical Society, 1900: 273-281.
338 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
MILLETT F. W., 1900b – Report on the recent foraminifera of the Malay NORMAN A. M., 1878 – On the genus Haliphysema,
Archipelago collected by Mr A. Durrand, F.R.M.S, Part IX. with description of several forms apparently allied to it.
Journal of the Royal Microscopical Society, 1900: 539-549. Annals and Magazine of Natural History, 1: 265-284.
MILLETT F. W., 1901a – Report on the recent foraminifera of the Malay
Archipelago collected by Mr A. Durrand, F.R.M.S, Part X.
Journal of the Royal Microscopical Society, 1901: 1-11.
OFLAZ S. A., 2006 – Taxonomy and Distribution of the Benthic
MILLETT F. W., 1901b – Report on the recent foraminifera of the Malay Foraminifera in the Gulf of Iskenderun, Eastern Mediterranean.
Archipelago collected by Mr A. Durrand, F.R.M.S, Part XI. Unpublished MSc. thesis, Middle East Technical University, Ankara,
Journal of the Royal Microscopical Society, 1901: 485-497. 306 p.
MILLETT F. W., 1901c – Report on the recent foraminifera of the Malay O’HERNE L., 1974 – A reconsideration of Amphistegina lessonii
Archipelago collected by Mr A. Durrand, F.R.M.S, Part XII. d’Orbigny 1826, sensu Brady, 1884 (Foraminifera).
Journal of the Royal Microscopical Society, 1901: 619-628. Scripta Geol., 26: 1-52.
MILLETT F. W., 1902 – Report on the recent foraminifera of the Malay OHKUSHI K., THOMAS E., KAWAHATA H., 2000 – Abyssal benthic foraminifera
Archipelago collected by Mr A. Durrand, F.R.M.S, Part XIII. from the northwestern Pacific (Shatsky Rise) during the last 298 kyr.
Journal of the Royal Microscopical Society, 1902: 509-528. Marine Micropaleontology, 38: 119-147.
MILLETT F. W., 1903a – Report on the Recent foraminifera of the Malay ÖKI K., 1989 – Ecological analysis of benthonic foraminifera
Archipelago collected by Mr A. Durrand, F.R.M.S Part XIV. in Kagoshima Bay, South Kyushu, Japan.
Journal of the Royal Microscopical Society, 1903: 253-275. South Pacific Study, 10: 1-191.
MILLETT F. W., 1903b – Report on the Recent foraminifera of the Malay ORBIGNY A. d’, 1826 – Tableau méthodique de la classe des Céphalopodes,
Archipelago collected by Mr A. Durrand, F.R.M.S Part XV. 3e Ordre, Foraminifères. Annales de Sciences Naturelles, 7 : 254-314.
Journal of the Royal Microscopical Society, 1903: 685-704. ORBIGNY A. d’, 1839ª – « Foraminifères ». In de la Sagra R. (ed.) :
MOEBIUS K. A., 1880 – “Foraminifera von Mauritius”. Histoire physique, politique et naturelle de l’île de Cuba, Paris,
In Mobius K. A., Richter, F., von Martens E. (eds): Arthus Bertrand : 1-224.
Beiträge zur Meeresfauna der Insel Mauritius und der Seychellen, ORBIGNY A. d’, 1839b – « Foraminifères des îles Canaries ».
Gutman, Berlin: 65-122. In: Barker-Web P., Berthelot S. (eds): Histoire naturelle des îles
MONTAGU G., 1803 – Testacea Britannica, or natural history Canaries, Paris, Bethune, 2 (Zoologie) : 119-146.
of British shells, marine, land, and fresh-water, including ORBIGNY A. d’, 1839c – Voyage dans l’Amérique méridionale,
the most minute. J. S. Hollis, Romsey, England. foraminifères. Levrault, Paris and Strasbourg, vol. 5, 86 p.
MONTAGU G., 1808 – Testacea Britannica; supplement. ORBIGNY A. d’, 1846 – Foraminifères fossiles du bassin tertiaire
Exeter. England. J. S. Hollis. de Vienne (Autriche). Paris, Gide et Compe, 312 p.
MONFORT D. de, 1808 – Conchyliologie systématique et classification OUILLON S., DOUILLET P., LEFEBVRE J. P., LE GENDRE R., JOUON A.,
méthodique des coquilles. Paris, Schoell. BONNETON P., FERNANDEZ J. M., CHEVILLON C., MAGAND O., LEFÈVRE J.,
LE HIR P., LAGANIER R., DUMAS F., MARCHESIELLO P., BEL MADANI A.,
MOODLEY L., BOSHKER H. T. S., MIDDELBURG J. J., PEL R., HERMAN P.,
ANDRÉFOUËT S., PANCHÉ J. Y. & FICHEZ R., 2010 –
DE DECKERE E., HEIP C. H. R., 2000 – Ecological significance
Circulation and suspended sediment transport in a coral
of benthic foraminifera: 13C labelling experiments.
reef lagoon: the south-west lagoon of New Caledonia.
Marine Ecology Progress Series, 202: 289-295.
Marine Pollution Bulletin, Special Issue on New Caledonia 61:
MURRAY J. W., 2006 – Ecology and applications of benthic foraminifera. 269-296.
Cambridge University Press, Cambridge, United Kingdom.
PARKER W. K., JONES T. R., 1857 – Description of some foraminifera QUILTY P. G., 1974 – Tasmanian Tertiary foraminifera, Part 1,
from the coast of Norway. Annals and Magazine of Natural History, Textularia, Miliolina, Nodosariacea: The Papers and Proceedings
series 2, 19: 273-303. of the Royal Society of Tasmania, 108: 31-106.
PARKER W. K., JONES T. R., 1859 –
On the nomenclature of the foraminifera.
II. On the species enumerated by Walker and Montagu.
Annals and Magazine of Natural History, series 3, 4: 333-351. RAINER S. F., 1992 –
Diet of prawns from the continental slope of North-Western Australia.
PARKER W. K., JONES T. R., 1860 –
Bulletin of Marine Science, 50: 258-274.
On the nomenclature of the foraminifera. 4 (continued).
Annals and Magazine of Natural History, series 3, 4: 333-351. RASHEED D. A., 1971 – Some foraminifera belonging to Miliolidae and
PARKER W. K., JONES T. R., 1865 – Opthalmidiidae from the Coral Sea, south of Papua (New Guinea),
On some foraminifera from the North Atlantic and Arctic Oceans, Part 2. Journal of the Madras University, Section B, 37-38: 19-68.
including Davies Straits and Baffin’s Bay. REICHEL M., 1937 – Étude sur les Alvéolines, II.
Philosophical Transactions of the Royal Society, 155: 325-441. Schweizerische Paläontologische Abhandlung, 59 : 95-147.
PARKERW. K., JONES T. R., BRADY H. B., 1865 – RENAUD-DEBYSER J., 1965 – Note préliminaire sur la microfaune
On the nomenclature of the foraminifers, des fonds meubles du lagon (Baie de Saint-Vincent),
Part XII. The species enumerated by d’Orbigny Nouvelle-Calédonie. Cahiers du Pacifique, 7 : 107-116.
in the “Annales des Sciences Naturelles”, 7, 1826. RENEMA W., 2003 – Foraminifera on reefs around Bali (Indonesia).
Annals and Magazine of Natural History, series 3, 16: 15-41. Zoologische Verhandelingen, 345: 337-366.
PARR W. J., 1932 – Victorian and South Australian foraminifera - RENEMA W., HOHENEGGER J., 2005 – On the identity of Calcarina spengleri
Proceedings of the Royal Society of Victoria, 44, (Gmelin, 1791). Journal of Foraminiferal Research, 35: 15-21.
Part I: 1-14; Part II: 218-234.
REUSS A. E., 1848 – Die fossilen Polyparien des Wiener Tertiarbeckens.
PARR W. J., 1941 – A new genus, Planulinoides, Haidengers Naturwissenshaftliche Abhandlungen, Wien, 2: 1-109.
and some species of foraminifera from South Australia.
REUSS A. E., 1850 – Neues Foraminiferan aus den Schichten
Mining and Geological Journal, 2: 305.
des österreichischen Tertiärbeckens.
PARR W. J., 1945 – Recent Foraminifera from Barwon Heads, Victoria. Denkschriften der Kaiserlichen Akademie der Wissenschaften.
Proceedings of the Royal Society of Victoria, 56: 189-227. Mathematisch-Naturwissenschaftliche Klasse, 1: 365-390.
PARR W. J., 1950 – Foraminifera. Reports of the British, Australian REUSS A. E., 1851 – Ueber die fossilen Foraminiferen und
and New Zealand Antarctic Research Expedition 1929-1931, Entomostraceen der Septarienthone der Umgegend von Berlin.
series B (Zoology and Botany), 5: 233-392. Zeitschrift der Deutschen Geologischen Gesellschaft, Berlin, 3:
PATTERSON R. T., RICHARDSON R. P., 1987 – 49-91.
A taxonomic revision of the unilocular foraminifera. REUSS A. E., 1858 – Ueber die Foraminiferen von Pietzpuhl. Zeitschrift
Journal of Foraminiferal Research, 17: 212-226. der Deutschen Geologischen Gesellschaft, Berlin. 10, 433- 438.
PAWLOWSKI J., BOLIVAR I., FAHRNI J. F., DE VARGAS C., REUSS A. E., 1861 – Beiträge zur Kenntnis der tertiären Foraminiferen
BOWSER S. S., 1999a – Molecular evidence that Reticulomyxa Fauna: Sitzungsberichte der Kaiserlichen Akademie
filosa is a freshwater naked foraminifera. der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse
Journal of Eukaryotic Microbiology 46: 612-617. in Wien, 42: 355-370.
PAYRI C., RICHER de FORGES B., 2007 – “Compendium of New Caledonian REUSS A. E. 1863 – Die Foraminiferen des norddeutschen Hils
Marine species: overview”. In Payri C. E., Richer de Forges B. (eds): und Gault. Sitzungsberichte der Kaiserlichen Akademie
Compendium of marine species of New Caledonia, IRD Nouméa, der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse
Documents Scientifiques et Techniques, 117, seconde édition: 13-18. in Wien, 46: 5-100. (Synonymous with Reuss 1862).
PERELIS L., REISS Z., 1975 – Cibicididae from the Gulf of Elat. REUSS A. E., 1866 – Die Foraminiferen, Anthozoa
Israel Journal of Earth Sciences, 24: 73-96. und Bryozoa des deutschen Septarientons.
PETRI S., 1957 – Foraminíferos miocênicos da Formação Pirabas. Denkschriften der Kaiserlichen Akademie der Wissenschaften.
Boletim da Faculdade de Filosofia, Ciências e Letras, Mathematisch-Naturwissenschaftliche Klasse, 25: 117-214.
Universidade de São Paulo, São Paulo, 216: 1-79. REUSS A. E., 1867 –
PHILIPPI R. A., 1844 – Nachtrag zum zweiten Bande der Enumeratio Die fossile Fauna der Steinsalzablagerungen von Wieliczka in Galizien.
Molluscorum Siciliae. Zeitschrift für Malakozoologie, 1: 100-112. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften.
PHLEGER F. B., PARKER F. L, 1951 – Ecology of foraminifera, Mathematisch-Naturwissenschaftliche Klasse, 55: 17-182.
northwest Gulf of mexico, part 2: foraminifera species. REVETS S. A., 1991 – The generic revision of the Reussellids
Geological Society of America Memoir, 46: 1-64. (Foraminifera). Journal of Micropalaeontology, 10: 1-15.
PHLEGER F. B., PARKER F. L., PEIRSON J. F., 1953 – North Atlantic REVETS S. A., 1992 – The structure and taxonomic position
Foraminifera. Reports Swedish Deep-Sea Expedition, 7: 1-122. of Millettia Schubert, 1911 (Foraminiferida).
POPESCU G., 1983 – Marine Middle Miocene monothalamous Journal of Micropalaeontology, 11: 37-46.
Foraminifera from Romania. Memorii Institutul de Geologie REVETS S. A., 1993 – The revision of the genus Elongobula Finlay
si Geofizica, 31: 261-280. 1939. Journal of Foraminiferal Research, 23: 254-266.
POPESCU G., CRIHAN I.-M., 2004 – Contributions to the knowledge REVETS S. A., 1996 – The generic revision of five families of rotaliine
of the calcareous unicameral foraminifera from the Middle Miocene foraminifera: Part I. The Bolivinitidae. Cushman Foundation
from Romania. Acta Palaeontologica Romaniae, 4: 403-421. for Foraminiferal Research Special Publication, 34: 1-55.
340 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
REVETS S. A., 2000 – Foraminifera of Leschenault Inlet. SCHULTZE F. E., 1875 – Zoologische Ergebnisse der Nord-seefahrt,
Journal of the Royal Society of Western Australia, 83: 365–375. vom 21 Juli bis 9 September, 1872. I Rhizopoden,
REVETS S. A., WHITTAKER J. E., 1991 – II Jahresbericht der Commission zur Wissenschaftl.
The taxonomic position of Orthoplecla Brady, 1884 (Foraminiferida). Untersuchung Deutschen Meere in Kiel für die Jahre 1872, 1873,
Journal of Micropalaeontology, 9: 167-172. Berlin: 99-114.
RHUMBLER L., 1906 – Foraminifera von Laysan und den Chatham-Inseln. SCHULTZE M. S., 1854 – Ueber den Organismus der Polythalamien
Zoologischer Jahresbericht, 24: 21-80. (Foraminiferen), nebst Bemerkungen uber die Rhizopoden
im Allgemeinen. Leipzig. Wilhelm Engelmann.
RHUMBLER L., 1911 – Die Foraminiferen (Thalamophoren)
der Plankton-Expedition. Ergebnisse der Plankton-Expedition SCHWAGER C., 1866 – Fossile Foraminiferen von Kar Nikobar,
der Humoldt- Stiftung, Kiel und Leipzig, 3: 1-476. Reise der Österreichischen Fregatte Novara um die Erde
in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore
RICHARDSON S. L., 2006 – Response of epiphytic foraminiferal
B. Vo Wüllerstorf-Urbair. Geologischer Theil 2 (1); Geologische
communities to natural eutrophication in seagrass habitats
Beobachtungen 2; Paläontologische Mittheilungen: 187-268.
off Man O’War Cay, Belize. Marine Ecology, 27: 404-416.
ROUGERIE P., 1986 – Le lagon sud-ouest de la Nouvelle-Calédonie : SCOTT D. B., MEDIOLI F. S., 1980 – Quantitative studies of marsh
spécificité hydrologique, dynamique, productivité. foraminiferal distributions in Nova Scotia and comparison with those
Paris, Orstom, Études et Thèses, 234 p. in other parts of the world: implications for sea level studies.
Cushman Foundation for Foraminiferal Research Special
RYMER JONES F. W. O., 1872 – On some Recent forms Publication, 17: 1-58.
of Lagenae from deep-sea soundings in the Java seas.
Transactions of the Linnean Society of London, 30: 45-69. SCOTT D. B., TAKAYANAGI Y., HASEGAWA S., SAITO T., 2000 –
Illustration and reevaluation of affinities of neogene foraminifera
RZEHAK A., 1886 – Die Foraminiferenfauna der Neogenformation
described from Japan. Palaeontologia Electronica, 3 (2): 41 p.
der Umgebung von Mähr.-Ostrau. Naturforschender Verein
http://palaeo-electronica.org/2000_2/foram/issue2_00.htm
Brünn, Verhandlungen, Brünn, 1885, 24: 77-126.
SEGUENZA G., 1862 – Dei terreni Terziarii del distretto di Messina;
Parte II - Descrizione dei foraminiferi monotalamici delle marne
Mioceniche del distretto di Messin. Messina. T. Capra.
SAID R., 1949 – Foraminifera of the northern Red Sea. SEGUENZA G., 1880 – Le formazioni Terziarie nella provincia
Cushman Laboratory of Foraminiferal Research di Reggio (Calabria), Classe di Scienze Fisiche, Matematiche
Special Publication, 26: 1-44. e Naturali, Ser. 3. 6, 446 p. Atti Reale Accademie dei Lincei, Roma.
SAIDOVA Kh. M., 1975 – Bentosyne foraminifery Tikhogo Okeana SEIGLIE G. A., 1964 – New and rare foraminifers from Los Testigos
[Benthonic foraminifera of the Pacific Ocean]. Institut Okeanologii Reefs, Venezuela. Caribbean Journal of Science, 4: 497-512.
im. P.P. Shirshova, Akademiya Nauk SSSR, Moskva. [Russian]
SELLIER de CIVRIEUX J. M., 1977 – Las Discorbidae del Mar Caribe,
SAIDOVA Kh. M., 1981 – O sovremennomsostoyanii sistemy
frente a Venezuela. Cuadernos Oceanográficos,
nadvidovykh taksonov Kaynozoyskikh bentosnykh foraminifer
Universidad de Oriente, Cumana, 6: 1-44.
(On an up-to-date system of supraspecific taxonomy
of Cenozoic benthonic foraminifera). SEN GUPTA B. K., 1999 – Systematics of Modern Foraminifera.
Moscow, Institut Okeanologii P.P. Shirshova, Akademiya Nauk SSSR. In Sen Gupta B. K. (ed.): Modern Foraminifera, Kluwer Academic
Publishers, Dordrecht, Boston, London: 7-36.
SARASWATI P. K., 2002 – Growth and habitat of some recent miliolid
foraminifera: Palaeoecological implications. SGARELLA F., MONCHARMONT ZEI M., 1993 – Benthic foraminifera
Current Science, 82: 81-84. of the Gulf of Naples (Italy): systematics and autoecology.
Bollettino della Società paleontologica italiana, 32: 145-264.
SARS G. O., 1872 – Undersoslashgelser over Hardangerfjordens Fauna.
Forhandlinger i Videnskasselsbet i Kristiana 1871: 246-255. SHUTO T., 1953 – A study on the foraminiferal assemblage
SAUNDERS J. B., 1957 – Trochamminidae and certain Lituolidae of Omura Bay, Nagasaki Prefecture, Kyushu.
(foraminifera) from the recent brackish-water sediments Japanese Journal of Geology and Geography, 23: 127-138.
of Trinidad, British West Indies. Smithsonian miscellaneous SIDDAL J. D., 1878 – On the foraminifera of the River Dee. Proc.
contributions, 134: 1-16. Chester Soc. Nat. Sci., 2: 42-56.
SCHLUMBERGER C., 1891 – Révision des Biloculines des grands fonds. SIDEBOTTOM H., 1906 – Report on the recent Foraminifera
Mémoires de la Société Zoologique de France, 4 : 542-579. from the coast of the Island of Delos (Grecian Archipelago). Part. 3.
SCHLUMBERGER C., 1893 – Monographie des Miliolidées du golfe de Manchester Literary and Philosophical Society Memoirs
Marseille. Mémoires de la Société Zoologique de France, 6 : 57- 80. and Proceedings, 50 (5): 1-18.
SCHNITKER D., 1971 – Distribution of foraminifera SIDEBOTTOM H., 1907 – Report on the recent Foraminifera
on the North Carolina continental Shelf. from the coast of the Island of Delos (Grecian Archipelago).
Tulane Studies in Geology and Paleontology, 8: 169-215. Manchester Literary and Philosophical Society Memoirs
and Proceedings, 51 (9): 1-28.
SCHRÖDER, C. J., 1986 – Deep-water arenaceous foraminifera
in the northwest Atlantic Ocean. Canadian Technical Report SIDEBOTTOM H., 1908 – Report on the recent Foraminifera
of Hydrography and Ocean Sciences. Atlantic Geoscience Centre. from the Coast of the Island of Delos (Grecian Archipielago), Part V.
Bedford Institute of Oceanography: 1-191. Manchester Literary and Philosophical Society Memoirs
SCHUBERT R. J., 1904 – Die Ergebnisse der mikroskopischen and Proceedings pt. 5, 52 (13): 1-28.
Untersuchung der bei der aerarischen Tiefbohrung zu Wels SIDEBOTTOM H., 1910 – Two new species of Cassidulina.
durchteuften Schichten. Jahrbuch Der Kaiserlich-Koniglichen Journal of the Quekett Microscopical Club,
Geologischen Reichsanstalt, Wien, 53: 385-422. series 2, 11 (67): 105-108.
A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia | 341
SIDEBOTTOM H., 1912 – Lagenae of the Southwest Pacific Ocean, TODD R., 1961 – Foraminifera from the Onotoa Atoll, Gilbert Islands.
from soundings taken by HMS Waterwich, 1895. Professional Papers U.S. Geological Survey, 354-H: 171-191.
Journal of the Queckett Microscopical Club, series 2, 11: 70-44. TODD R., 1965 – The foraminifera of the tropical Pacific collections
SIDEBOTTOM H., 1913 – Lagenae of the Southwest Pacific Ocean of the “Albatross”,1899-1900, Part. 4. Rotaliform families
(supplementary paper). Journal of the Queckett Microscopical and Planctonic families.
Club, series 2, 12: 161-210. United States National Museum Bulletin, 61: 1-127.
SIDEBOTTOM H., 1918 – Report on the Recent foraminifera dredged off TODD R., 1966 – Smaller Foraminifera from Guam.
the east coast of Australia, H. M.S. “Dart” Station 19 (May 14, 1895), Professional Papers U.S. Geological Survey, 403-I: 1-41.
lat. 29°22’S long. 153°51’E, 465 fathoms, Pteropod ooze. TODD R., 1971 – Tretomphalus (Foraminifera) from Midway.
Journal of the Royal Microscopical Society, 1918: 121-152. Journal of Foraminiferal Research, 1: 162-169.
SILVESTRI A., 1904 – Forme nuove o poco conosciute di Protozoi TODD R., BRÖNNIMANN P., 1957 – Recent foraminifera
Miocenici piemontesi. Atti dell’Accademia della Scienze, and thecamoebina from the eastern Gulf of Paria.
Torino, Italia (1903-04), 39: 4-15. Cushman Foundation for Foraminiferal Research,
SKINNER H. C., 1961 – Revision of “Proteonina difflugiformis”. Special Publication, 3: 1-43.
Journal of Paleontology, 35 (6): 1239-1240. TOULOUSE M., 1965 – Introduction à l’étude des foraminifères
SLITER W. V., 1971 – Predation on benthic foraminifera. des sédiments actuels de la baie de Saint-Vincent
Journal of Foraminiferal Research, 1: 20-28. (lagon de la côte ouest de Nouvelle-Calédonie).
Unpublished DES Géologie, université de Montpellier, 87 p.
STACHE G., 1864 – Die Foraminiferen der tertiaren Mergel
TOULOUSE M., 1966 – Première étude des foraminifères des sédiments
des Whaingaroa-Hafens (Provinz Auckland).
actuels de la baie de Saint-Vincent (Nouvelle-Calédonie).
Novara-Exped., 1857-1859. Geologische Theil, 1 (2): 161-304.
Comptes-Rendus de l’Académie des Sciences, Paris, (D), 262 :
STEVENSON J., DODSON J. R., PROSSER I. P., 2001 – A late Quaternary record 1517-1518.
of environmental change and human impact from New Caledonia.
TOWE K. M., CIFELLI R., 1967 – Wall Ultrastructure in the calcareous
Palaeogeography, Palaeoclimatology, Palaeoecology, 168: 97-123.
foraminifera: Crystallographic aspects and a model for calcification.
SURESH GANDHI M., RAJAMANICKAM G. V., NIGAM R., 2002 – Journal of Paleontology, 41: 742-762.
Taxonomy and distribution of Benthic foraminifera TRAUTH F., 1918 – Das Eozänvorkommen bei Radstadt im Pongau
from the sediments off Palk strait, Tamil Nadu, east coast of India. und seine Beziehungen zu den gleichalterigen Ablagerunden
Journal of The Palaeontological Society of India, 47: 47-64. bei Kirchberg am Wechsel und Wimpassing am Leithagebirge.
SZARECK R., 2001 – Biodiversity and biogeography of recent benthic Denkschriften der Kaiserlichen Akademie der Wissenschaften,
foraminiferal assemblages in the south-western South China Sea Wien, Mathematisch-Naturwissenschaftliche Klasse, 95: 171-278.
(Sunda Shelf). Unpublished Ph.D. thesis, Universitätbibliotek
der Christian-Albrechts-Universität Kiel, 273 p., 28 pls.
UCHIO T., 1952 – New genera and species of foraminifera
from Hachijo Island, Tokyo Prefecture.
Japanese Journal of Geology and Geography, 22: 145-159.
TAKAYANAGI Y., 1951 – On some Ehrenbergina from Japan.
Transactions and Proceedings of the Paleontological Society UCHIO T., 1953 – On some foraminiferal genera in Japan.
of Japan, new series, 3: 85-93. Japanese Journal of Geology and Geophysics, 23: 151-162.
TERQUEM O., 1875 – Essai sur le classement des animaux UCHIO T., 1960 – Ecology of living benthonic foraminifera
qui vivent sur la plage et dans les environs de Dunkerque. from the San Diego, California, area. Cushman Foundation
Fascicule 1, Paris : 1-54. for Foraminiferal Research Special Publications, 5: 1-72.
TERQUEM O., 1878 – Les foraminifères et les Entomostracés-Ostracodes UJIIÉ H., 1963 – Foraminifera from the Yûrakuchô Formation
de Pliocène supérieur de l’île de Rhodes. (Holocene), Tokyo City. Science Reports of the Tokyo Kyoiku
Mémoires de la Société Géologique de France, série 3, 1 : 1-135. Daigaku. Section C., 8 (79): 27-41.
UJIIÉ H. H., 1990 – Bathyal benthic foraminifera in a piston
TESTAU J. L., CONAND F., 1983 – Estimation des surfaces
core from east off the Miyako islands, Ryukyu Island Arc.
des différentes zones du lagon de Nouvelle-Calédonie.
Bulletin of the College of Science, University of the Ryukyus, 49:
Unpublished Rapport Orstom, Nouméa, 5 p.
1-60, 32 pls.
THALMANN H. E., 1950 – New names and homonyms in foraminifera.
UJIIÉ H., RIFARDI D., 1993 – Some benthic foraminifera
Contributions from the Cushman Foundation for Foraminiferal
from the Oura River. Estuary and its Environs, Okinawa.
Research, 1: 41-45.
Bulletin of the College of Science University of the Ryukyus, 5:
THOMASSIN B. A., 1984 – Les récifs coralliens dans l’Indopacifique ouest : 121-243.
grands types de constructions et successions des phases d’édification.
Oceanis, l0 : 1-49.
TIZARD T. H., MURRAY J., 1981 – Exploration of the Faröe Shetland VAN MORKHOVEN F. P. C. M., BERGGREN W. A., EDWARDS A. S., 1986 –
Channel during the Summer of 1880 in H. M. hired ship Cenozoic cosmopolitan deep-water benthic foraminifera.
“Knight Errant”. Proceedings of the Royal Society of Edinburgh, Bulletin des centres de recherches exploration-production
11: 638-720. Elf-Aquitaine, Mémoire 11, 421 p.
TODD R., 1957 – “Smaller foraminifers”. VEGA A., GANACHAUD A., BOSSON J., 2005 – Atlas climatologique satellite
In: Geology of Saipan, Mariana Islands (Pt. 3) Paleontology, des courants, vents, élévation et température de surface
Professional Papers U.S. Geological Survey, 280-H: 265-320. dans la ZEE de la Nouvelle-Calédonie. Nouméa, IRD Éditions, 27 p.
342 | A Guide to 1,000 Foraminifera from Southwestern Pacific: New Caledonia
VELLA P., 1957 – Studies in New Zealand foraminifera; WIESNER H., 1931 – Die foraminiferen der deutschen Sudpolar
Part I – Foraminifera from Cook Strait. Expedition 1901-1903. Deutsche Südpolar Expedition, v. 20,
Part II – Upper Miocene to Recent species of the genus Notorotalia. Zoologie, 12: 53-165.
New Zealand Geological Survey Paleontological Bulletin, 28: WILLIAMSON W. C., 1848 – On the Recent British species of the genus
1- 64. Lagena. Annals and Magazine of Natural History, series 2, 1: 1-20.
VENEC-PEYRÉ M. T., SALVAT B., 1981 – Les foraminifères de l’atoll WILLIAMSON W. C., 1858 – On the recent foraminifera of Great Britain.
de Scilly (archipel de la Société): Étude comparée de la biocoenose London Ray Society, 107 p.
et de la thanatocoenose. Annales de l’Institut Océanographique,
WILSON B., RAMSOOK A., 2007 – Population densities and diversities
Paris, 57 : 79-110.
of epiphytal foraminifera on nearshore substrates, Nevis, west India.
VICKERMAN K., 1992 – The diversity and ecological significance Journal of Foraminiferal Research, 37: 213-222.
of protozoa. Biodiversity and Conservation, 1: 334-341. WIRRMANN D., SÉMAH A.-M., DEBENAY J.-P., CHACORNAC-RAULT M., 2011 –
VINCENT E., 1986 – Les associations de foraminifères benthiques Mid- to late Holocene environmental and climatic changes
du bassin des Loyauté (Nouvelle-Calédonie), in New Caledonia, southwest tropical Pacific, inferred
rapports avec la sédimentation (campagne BIOCAL, 1985). from the littoral plain Gouaro-Déva. Quaternary Research,
Unpublished DEA, Centre des Sciences de la Terre, Online 14 June 2011, doi:10.1016/j.yqres.2011.04.007.
université de Bourgogne, 28 p. WRIGHT J., 1886 – In Haddon A. C.: First report on the marine fauna
VINCENT E., LAURIN B., 1988 – Les associations de foraminifères benthiques of south-west of Ireland, Proceedings of the Royal Irish Academy,
du bassin des Loyauté (Nouvelle-Calédonie) : autochtonie Dublin, series 2, 4: 599-638.
et allochtonie. Revue de Micropaléontologie, 31 : 196-206. WRIGHT J., 1900 – The foraminifera of Dog’s Bay, Connemara.
VINCENT E., LAMBERT B., LAURIN B., MATHIEU R., 1991 – Irish Naturalist, 9: 50-55.
Distribution des foraminifères benthiques dans le bassin des Loyauté. WRIGHT T. J., 1902 – “Foraminifera”.
Documents et Travaux de l’IGAL, Paris, 15 : 127-149. In Reade T. M. (ed.): Glacial and Post-Glacial features
of the lower valley of the River Lune and its estuary,
Proceedings of the Geological Society, Liverpool, 9, 183 p.
WALKER G., JACOB E., 1798 –
In Kanmacher F.: Adam’s Essays on the Microscope, Ed. 2.
WANG NAI-WEN, 1964 – The discovery of Evolutononion YASSINI I., JONES B. G., 1995 – Foraminiferida and ostracoda
shansiense gen. et sp. nov. (foraminifera) and its significance from estuarine and shelf environments on thr southeastern
for stratigraphy and paleogeography. Annual meeting coast of Australia, Wollongong, NSW.
of the society of Oceanography and Limnology of China. University of Wollongong, 484 p.
Beijing, 1963, service press: abstract.
WARREN A. D., 1957 – Foraminifera of the Buras-Scofield Bayou region,
southeast Louisiana. Contributions from the Cushman ZHENG S. Y., 1979 –
Foundation for Foraminiferal Research, 8 (1): 29-40. The recent foraminifera of the Xisha Islands, Guangdong Province,
China, II. Studia Marina Sinica, 15: 101-232.
WELLS P. E., 1985 – Recent agglutinated benthonic Foraminifera [Chinese with summary and new genera and species in English]
(suborder Textulariina) of Wellington Harbour, New Zealand.
ZHENG S. Y., 1980 –
New Zealand Journal of Marine and Freshwater Research, 19:
The recent foraminifera of the Zhongsha Islands, Guangdong
575-599.
Province, China, I. Studia Marina Sinica, 16: 143-182.
WETMORE K. L., 1995 – Foraminifera: Life History and Ecology. [Chinese with summary and new genera and species in English]
University of California, Berkeley, Museum of Paleontology.
ZHENG S. Y., 1988 –
Available at: http://www.ucmp.berkeley.edu/foram/foramlh.html
The agglutinated and porcelaneous foraminifera
(viewed march 08, 2012).
of the East China Sea. China Ocean Press, Beijing, 337 p.
WHITTAKER J. E., HODGKINSON R. L., 1979 – [Chinese with summary in English]
Foraminifera of the Togopi Formation, eastern Sabah, Malaysia. ZHENG S. Y., CHENG T. S. WANG X., FU Z., 1978 – The Quaternary
Bulletin of the British Museum (Natural History), 31: 1-120. foraminifera of the Dayuzhang irrigation area, Shandong Province,
WIESNER H., 1923 – Die Milioliden der östlichen Adria. and a preliminary attempt at an interpretation of its depositional
Prag-Bubenc. The author. environment. Studia Marina Sinica, 13: 16–78.
Alphabetical Index
The first part of this guide is designed to introduce the reader to pre-identification of specimens before advancing to the next
New Caledonia, a French archipelago in the tropical-subtropical chapter for confirmation on the basis of descriptions and larger
southwestern Pacific (latitude 15º-26º S and longitude 156º- photographs of the species. At the end of the book, and mostly for
174º E), with the main island (400 km long and 50 km wide) specialists, a systematics list of foraminiferal species identified
being the third largest island in the southwestern Pacific after from New Caledonia is provided, with a brief synonymy list
New Guinea and New Zealand. It presents an overview of the including the original type reference, and a few references that
geologic, geomorphic, oceanographic and climatic setting of illustrate the species clearly. Systematics is organized following
New Caledonia at general, regional, and local scales. Then, the LOEBLICH & TAPPAN (1992, 1994) and KAMINSKI (2004).
current knowledge of foraminifera, including biology and the A total of 1,043 species are described and illustrated by scanning
main test components used for identification is summarized and electron and light microscope photographs. They were collected
illustrated. It is mostly destined for non-specialists and people from over 800 samples that span 0-700 m water depths in a high
new to foraminifera. In the following chapters, foraminiferal diversity of habitats including mangrove, estuaries, lagoons, coral
studies of New Caledonia are synthesized, with emphasis on reef and shelf. Among them, 665 had not been reported around
studies carried out in lagoonal, reefal and paralic environments New Caledonia before the compilation published in 2007. Two
during the past 35 years, including distribution maps of the main new species are described: Triloculina elongotricarinata and
species, distribution models related to depth and mud content of Hoeglundina neocarinata, a new species name is proposed for
the sediment, and examples of foraminifera as environmental Calcarina exuberans, instead of Calcarina hispida var. pulchella,
indicators at various space and time scales. and a new genus name is proposed for Quirimbatina rimosa
The main part of this work is a guide to the taxonomy and instead of Mimosina rimosa. One hundred and forty-two species
identification of benthic foraminifera that are very diversified could not be determined at a specific level and are recorded under
and abundant around New Caledonia. It aims to assist micropa- open nomenclature. A high proportion of them are presumably new
leontologists and students of foraminifera, but also to provide a species, but more specimens are needed before proposing new species
resource for environmental managers and scientists who may names. Including the 158 species reported in the literature, and not
use foraminifera as a tool for environmental monitoring and found for being illustrated in this book, the number of benthic
assessment, without being specialists of this group. For achieving foraminifera species identified hitherto around New Caledonia
this goal, species are classified by the nature of the wall and reaches 1,201. Most of them had been reported from the central
the dominant morphological feature. First, a photographical and western Pacific, and/or the Indo-Pacific area, but some species
summary presents full-page plates showing small images of all had been found from remote areas, such as the spectacular
species divided into agglutinated, porcelaneous and hyaline, with Quinqueloculina erinacea Mikhalevich, reported from the tropical
the hyaline species further classified by the coiling mode. It will Atlantic, or Rotaliammina siphonata (Seiglie), reported from
allow an easy comparison between related species and a quick Venezuela, showing the high dispersal potential of some species.
Résumé
La première partie de ce guide est une présentation de la prédétermination des spécimens avant le passage au chapitre
Nouvelle-Calédonie, archipel français du sud-ouest Pacifique suivant, où une description et de plus grandes illustrations
tropical et subtropical (latitude 15º-26º S, longitude 156º-174º E), permettent d’affiner la détermination. À la fin de l’ouvrage, et plus
dont l’île principale, longue de 400 km et large de 50 km est la particulièrement destinée aux spécialistes, une liste systématique
troisième île du sud-ouest Pacifique par ses dimensions, après la des foraminifères identifiés en Nouvelle-Calédonie est fournie,
Nouvelle-Guinée et la Nouvelle-Zélande. Cette partie donne un avec une brève liste de synonymies qui inclut la référence du type
aperçu des caractéristiques géologiques, géomorphologiques, original de l’espèce et quelques références qui l’illustrent clairement.
océanographiques et climatiques à l’échelle locale, régionale et La systématique suit les conceptions de LOEBLICH & TAPPAN (1992,
générale. Ensuite, les connaissances actuelles sur les foraminifères, 1994) et de KAMINSKI (2004).
incluant leur biologie et les caractéristiques du test utilisées pour Au total, 1 043 espèces sont décrites et illustrées par des images au
leur détermination sont résumées et illustrées à destination des microscope électronique à balayage et des photos au microscope
non-spécialistes et des débutants en « foraminiférologie ». Dans optique. Ces espèces ont été récoltées dans 800 échantillons
les chapitres suivants, les études sur les foraminifères de Nouvelle- prélevés entre 0 et 700 m dans des environnements très divers
Calédonie sont synthétisées, avec une attention particulière pour incluant la mangrove, les estuaires, les lagons, les environne-
celles qui ont été réalisées durant les 35 dernières années dans les ments récifaux et la plateforme continentale. Parmi ces espèces,
environnements lagonaires, récifaux et paraliques. Cette synthèse 665 n’avaient jamais été signalées en Nouvelle-Calédonie avant la
est illustrée par des cartes et des modèles de répartition en fonction compilation publiée en 2007. Deux espèces nouvelles sont décrites :
de la profondeur et de la teneur en vase du sédiment. Des exemples Triloculina elongotricarinata et Hoeglundina neocarinata,
sont donnés sur l’utilisation des foraminifères comme indicateurs un nouveau nom d’espèce est proposé pour Calcarina exuberans,
des conditions environnementales à différentes échelles de temps en remplacement de Calcarina hispida var. pulchella et un
et d’espace. nouveau nom de genre est proposé pour Quirimbatina rimosa en
La partie principale de cet ouvrage est un guide taxonomique pour remplacement de Mimosina rimosa. 142 espèces n’ont pas pu être
l’identification des foraminifères benthiques, très diversifiés et très déterminées au niveau spécifique et ont été laissées en nomenclature
abondants autour de la Nouvelle-Calédonie. Elle est destinée aux ouverte. Une proportion notable d’entre elles correspond probable-
micropaléontologistes et aux étudiants, mais son but est également ment à des espèces nouvelles, mais davantage de spécimens sont
de fournir une base de données accessible aux décideurs environ- nécessaires avant de pouvoir proposer un nouveau nom d’espèce.
nementaux et aux scientifiques qui pourraient avoir à utiliser les En rajoutant les 158 espèces rapportées dans la littérature, et non
foraminifères comme un outil pour la surveillance et l’évaluation retrouvées pour être illustrées dans cet ouvrage, le nombre de
environnementale sans être spécialistes de ce groupe. Dans ce foraminifères benthiques inventoriés jusqu’à maintenant en
but, les espèces sont classées en fonction de la nature de leur test Nouvelle-Calédonie atteint 1 201. La plupart de ces espèces ont été
et de leur caractéristique morphologique dominante. Tout signalées dans le centre et l’ouest Pacifique, et/ou dans la région
d’abord, un sommaire photographique présente des planches en Indopacifique, mais quelques espèces ont été trouvées dans des
pleine page avec de petites illustrations pour toutes les espèces, zones très éloignées comme la spectaculaire Quinqueloculina
réparties en agglutinées, porcelanées et hyalines, ces dernières erinacea Mikhalevich, décrite dans l’Atlantique tropical, ou
étant subdivisées en fonction de l’arrangement des loges. Ce Rotaliammina siphonata (Seiglie), décrite au Venezuela,
sommaire permet une comparaison aisée entre les espèces et une montrant le fort potentiel dispersif de certains foraminifères.
Contents