Small Meadow Katydid Songs

by Carl Strang

As children, we can hear all the singing insects. As we age, though, we gradually lose our ability to hear the higher-pitched ones. When I began to study them in 2006, I already was unable to hear any of the small meadow katydids (genus Conocephalus) in the field. I acquired a device called the SongFinder, which lowers the pitch of songs by cutting their frequency to half, one-third, or one-fourth.

A few years ago, I learned that this device no longer is made, and sought a backup. I ended up with a bat detector.

This one, called the Walkabout, can be set to the range of the higher-pitched singing insects (most bat detectors are limited to frequencies above that range). It has the further advantages of having a speaker instead of headphones, a visual display of the sounds it picks up, and can make recordings. Furthermore, unlike the limited directionality of the SongFinder, its microphone cone attachment allows the user to zero in on a single singing insect for a visual identification.

This year I made better use of both devices, playing with settings and making recordings to improve my ability to find Conocephalus katydids and identify them by their songs, without having to see them. In the following review of species, you may well be able to hear the recordings (made with the Walkabout, with one exception), either because you are in a quiet setting without the competing sounds that overwhelm in the field, or because the recordings are somewhat pitch-reduced.

The most common of these critters in the Chicago region is the short-winged meadow katydid (C. brevipennis). These occur in grasses in a wide range of habitats.

Its song consists of usually 1-3 evenly spaced ticks followed by a short buzz:

The next most common species, which occurs in wet to mesic grasses, is the slender meadow katydid (C. fasciatus).

Here, the ticks are more irregularly spaced and usually more numerous, the whole impression much less metronomic than in the short-winged:

This year I became better acquainted with the black-sided meadow katydid (C. nigropleurum).

One of our more beautiful singing insects, the black-sided is limited to wet areas with some coarser herbaceous vegetation or woody stems. Its song is a continuous series of snare-drum-like rapid ticks (this recording I made with my Marantz digital recorder and a Sennheiser shotgun mike, and includes a common true katydid singing nearby):

The shape of these ticks is distinctive when viewed in a sonograph. This image is one second’s worth:

Another, less commonly encountered wetland species is the long-tailed meadow katydid (C. attenuatus), which can be all brown as in the photo, but also can have bright green legs.

It also has a continuous song that is more like a buzz, the pulses coming more rapidly:

The sonograph shows that the pulses are regular and identical, in this image 0.2 second’s worth:

The straight-lanced meadow katydid (C. strictus) lives in sparser grassy vegetation on poor, dry soils.

Its song resembles that of the long-tailed, but their habitats are distinct.

Also, the volume increases and decreases. This 0.2-second sample shows a decrease:

There are two other Conocephalus species in the region, the woodland meadow katydid (C. nemoralis) and the prairie meadow katydid (C. saltans). The former has a song consisting of rapid-fire brief buzzes, some of which are separated by drumrolls of ticks. The latter produces a continuous series of even briefer buzzes, so short they are almost tick-like. I need better recordings of both, and hope to get them next year.

Another Season in the Books

by Carl Strang

In earlier posts I outlined the most significant finds in this year’s research on singing insects. The season was sprinkled with smaller delights, too, and I am pleased to call it successful. My game plan emphasized visiting new sites and trying to plug distributional holes for some species. I looked at 11 new sites, added 20 county records for all species combined, and closed the book on three species, i.e., I now have found them in every county or, at least, every county where I expect to find them. Those were the gray ground cricket (Allonemobius griseus), which I found at Chicago’s Montrose Park on September 6 for a Cook County record; the four-spotted tree cricket (Oecanthus quadripunctatus), which I now have documented in all 22 counties of my study region; and the oblong-winged katydid (Amblycorypha oblongifolia), which stubbornly had refused to reveal itself in LaPorte County, Indiana, until I heard a few singing on August 6 at Kingsbury Fish & Wildlife Area.

I also found new north locations for two of the species that are expanding their range from the south: the slow-tinkling trig (Anaxipha tinnulenta) at the Grand Mere Lakes in Berrien County, Michigan, and the handsome trig (Phyllopalpus pulchellus) at the northern end of Grassy Lake Forest Preserve for a Lake County, Illinois, record.

Now for a few photos of critters found along the way. I wanted to clarify the relatively early, long-trilling tree crickets I have been hearing at DuPage County’s Springbrook Prairie Forest Preserve. These proved to be a mix of four-spotted and Forbes’s (O. forbesi) tree crickets.

Forbes’s tree cricket from Springbrook Prairie, temporarily chilled for photographic purposes

I had visited Cook County’s Penny Road Pond preserve a few times before, but this year got into a large part of it that was new to me.

Texas bush katydid (Scudderia texensis), a site record for Penny Road Pond.

One of the new sites I visited was Spring Lake Forest Preserve in Cook County. As I walked a woodland trail I saw an interesting looking grasshopper, which I concluded was a sprinkled grasshopper (Chloealtis conspersa). This was the year’s second county record for the species, the first being the subject of an earlier post.

Sprinkled grasshopper nymph at Spring Lake

The remaining two photos are from a September 2 visit to Pulaski County, Indiana.

This female spotted ground cricket (Allonemobius maculatus) at Tippecanoe River State Park posed to give me a better photo than I have taken in past encounters with the species.
I continue to be impressed by ovipositor lengths in female straight-lanced meadow katydids (Conocephalus strictus) like this one at the Winamac Fish & Wildlife Area.

Looking ahead to 2023, my top priorities again will be visits to new sites, and efforts to conclude searches for species in counties where I have yet to find them but where they are likely to occur.

More Prairie Meadow Katydids

 

by Carl Strang

Last year I first encountered what I thought were prairie meadow katydids (Conocephalus saltans), in my survey of the Chicago region’s singing insects. Further study confirmed my identifications, and set the stage for finding the species in a third location in 2017. On September 2, Lisa Rainsong and I ran into a cluster of small meadow katydids at the Pembroke Savanna Nature Preserve, a Nature Conservancy savanna in eastern Kankakee County. These proved to be a mix of two species, straight-lanced meadow katydids (Conocephalus strictus) and prairie meadow katydids.

A scene at Pembroke Savanna, which I regard as the most beautiful site in the 22-county Chicago region.

Finding these two similar species together provided us with a tutorial in distinguishing them. Most of the individuals were females, and the contrast in their ovipositors could not be starker. Those of the prairie meadow katydids had a slight curve, and were much shorter.

Female prairie meadow katydid, Pembroke

Straight-lanced females have straight ovipositors that typically are as long as their bodies, or longer.

Female straight-lanced meadow katydid, Judy Burton Nature Preserve, Indiana. This one is atypical in having long wings. Most have wings about a third the length of the abdomen.

Prairie meadow katydids have wings that usually are only a quarter of the abdomen length. The knob at the tip of the head is more pronounced, though both species have this knob. The sides of the hind femurs also are different. In prairie meadow katydids there is a pattern of thin lines that resemble a ladder, on a brown leg. There usually is a diffuse black line that appears to be within the straight-lanced meadow katydid’s green hind femur. Some variation occurs in many of these features, so I advise caution and the examination of several individuals within a population.

Males have been fewer in both species, in the populations I have examined.

 

Male prairie meadow katydid, Pembroke

Again, the more exaggerated head knob, shorter wings, and different femur pattern are helpful. Cerci are very different in the two species, also.

Male straight-lanced meadow katydid, from another eastern Kankakee County savanna site.

I did not get a photo showing the prairie meadow katydid’s cerci, but their ends are much shorter, proportionately, than those of the straight-lanced in the photo, comparable in length to the teeth, and bend outward somewhat rather than being straight.

A final curious note from Pembroke was that the prairie meadow katydids were all brown, as the photos show. The straight-lanced meadow katydids had considerable amounts of green color. This may have been the result of local selective pressures, as this is not a consistent difference across the species’ ranges.

Making a Case 2: Prairie Meadow Katydid

by Carl Strang

After more than ten years of study, I still have not found several species of singing insects that historically were known in the Chicago region. One of these was the prairie meadow katydid, one of the smaller members of its group. On August 16, Wendy Partridge and I were checking out the bike trail near the nature center at Illinois Beach State Park. Through the SongFinder, which allows me to hear high pitches, I heard faint rapid ticking sounds which, when heard from a closer position, resolved into brief buzzes. They were mechanical and katydid-like rather than cricket-like in quality, and didn’t match anything I had heard before. Wendy, with her unusually good hearing, barely could pick them up. Later, checking reference recordings, I decided they fit the song of the prairie meadow katydid. They were not in the habitat I would expect, however, being in dense vegetation where grasses were mixed with woody plants and forbs, generally shaded. That katydid species has been known to occur in that park, however.

Then on September 27 I caught and photographed a small female meadow katydid in a relatively dry portion of the Gensburg Prairie in Cook County.

Posterior portion of the Gensburg Prairie female.

Posterior portion of the Gensburg Prairie female.

Her color didn’t seem quite right for a short-winged meadow katydid, which is a common species at that site especially in the wetter portions. Studying the photos, and looking back at references, I have decided that she was a prairie meadow katydid. Along the way I looked back at all my photos of short-winged meadow katydids, and found two other individuals that met at least some of the criteria for prairie meadow katydid, one in 2011 at Mayslake Forest Preserve, and one in 2014 in the savanna woodland at Illinois Beach State Park.

The Mayslake female

The Mayslake female

The Illinois Beach State Park female

The Illinois Beach State Park female

All three have curved ovipositors that are slenderer in proportion than in a typical short-winged meadow katydid.

Typical female short-winged meadow katydid

Typical female short-winged meadow katydid

All three also have femoral patterns in which there is a lengthwise pair of lines, as in the short-winged, but have ladders of narrow crossbars rather than being clear between them as in typical short-winged. There is enough overlap in body length, according to references, that it is not a consideration in comparing those two species. The ovipositors are too short, and the femur patterns wrong, for straight-lanced meadow katydid to be a consideration. The Mayslake female is different from the other two individuals in three possibly significant ways: the femoral ground color is green rather than brown or tan, the wings are much longer, and the front of the head does not appear to rise so much (this last being a difference mentioned by W.S. Blatchley in his classic Orthoptera of Northeastern North America, which gives unusually detailed descriptions of species).

Anterior portion of the Gensburg Prairie female. Note the abrupt rise of the tip of the head when viewed from the side.

Anterior portion of the Gensburg Prairie female. Note the abrupt rise of the tip of the head when viewed from the side.

Dorsal view of the Gensburg Prairie female. Blatchley describes the wings of prairie females as covering only one-fourth of the abdomen, two-thirds in short-winged females. Furthermore, the tip of the head, when viewed from above, is relatively wide after an inward bending of its sides, expected from Blatchley’s description.

Dorsal view of the Gensburg Prairie female. Blatchley describes the wings of prairie females as covering only one-fourth of the abdomen, two-thirds in short-winged females. Furthermore, the tip of the head, when viewed from above, is relatively wide after an inward bending of its sides, expected from Blatchley’s description.

Compare this common meadow katydid, with its relatively narrow and straight-sided tip of the head, to the previous.

Compare this short-winged meadow katydid, with its relatively narrow and straight-sided tip of the head, to the previous.

 

Side view of the same male as in the previous photo. Again, note the lack of a rise in the top of the head profile.

Side view of the same male as in the previous photo. Again, note the lack of a rise in the top of the head profile.

Blatchley also believed that prairie meadow katydids occur only in “raw prairie,” a description which applies to Gensburg and Illinois Beach but not to Mayslake. I am inclined to regard the Mayslake individual as an anomalous short-winged meadow katydid, but pending study of museum specimens, am naming the other two prairie meadow katydids.

Oops! Etc. at Midewin

by Carl Strang

A couple years ago I came across a population of large band-winged grasshoppers with bright red hind wings, at St. Joseph County’s (Indiana) Bendix Woods. Focusing on the intense red color, I declared them to be northwestern red-winged grasshoppers. In the first half of August this year I ran into a second population in Illinois, at the Midewin National Tallgrass Prairie.

They had the same bright red color as at Bendix Woods.

They had the same bright red color as at Bendix Woods.

These are large grasshoppers, approaching Carolina grasshoppers in bulk.

These are large grasshoppers, approaching Carolina grasshoppers in bulk.

This time, though, I noticed a discrepancy in my ID that should have struck me the first time.

There is a honkin’ big bulge on the top of the pronotum.

There is a honkin’ big bulge on the top of the pronotum.

The northwestern red-winged grasshopper, which I now realize I have yet to meet, has a flat pronotum profile that furthermore is cleft by a significant fissure. These prove to be autumn yellow-winged grasshoppers, which in fact can have a range of colors in the hindwings. The following week, returning with fellow singing insect enthusiasts Lisa Rainsong and Wendy Partridge from Cleveland, and Wil Hershberger from West Virginia, we found many of these grasshoppers in fact have bright yellow wings. I need to get back there and get a photo of one for my singing insects guide.

While checking out the grasshoppers, we turned up two other species that were county records for my study.

The handsome grasshopper always is a delight. This one is a male.

The handsome grasshopper always is a delight. This one is a male.

Female handsome grasshoppers have green highlights in place of the male’s brown ones.

Female handsome grasshoppers have green highlights in place of the male’s brown ones.

Though still a nymph, this female is unambiguously a straight-lanced meadow katydid. The extra-long ovipositor and the diffuse-edged black band on the hind femur are giveaways.

Though still a nymph, this female is unambiguously a straight-lanced meadow katydid. The extra-long ovipositor and the diffuse-edged black band on the hind femur are giveaways.

Our main target in that visit was the dusky-faced meadow katydid, but that proves to be a much more complicated story deserving of its own blog post.

 

Season’s End Photo Bag

by Carl Strang

Time to share miscellaneous left-over photos from this year’s singing insects prospecting trips. These are pictures that didn’t fit the posts that covered the locations where they were taken. All are from within my 22-county survey area.

This regal fritillary fed from blazing star flowers at an eastern Illinois location far from this rare species’ main Illinois range.

This regal fritillary fed from blazing star flowers at an eastern Illinois location far from this rare species’ main Illinois range.

Though field guides describe the common checkered-skipper as eastern North America’s most abundant butterfly, it is not one I encounter very often.

Though field guides describe the common checkered-skipper as eastern North America’s most abundant butterfly, it is not one I encounter very often.

This bush katydid flew to a high perch on a tree but remained just long enough for me to take a distant photo before moving on. I don’t know enough to identify the species.

This bush katydid flew to a high perch on a tree but remained just long enough for me to take a distant photo before moving on. I don’t know enough to identify the species.

Handsome trig. I added one more Indiana county to the ones where I have found this cricket, but have yet to find it in any of the Chicago region’s other states.

Handsome trig. I added one more Indiana county to the ones where I have found this cricket, but have yet to find it in any of the Chicago region’s other states.

Straight-lanced meadow katydid. I found this species in 6 more counties in 2015, in Wisconsin, Illinois and Indiana.

Straight-lanced meadow katydid, long winged form. I found this species in 6 more counties in 2015, in Wisconsin, Illinois and Indiana.

Lulu Lake

by Carl Strang

During a 3-day Wisconsin trip last week, I made a lot of stops in Racine, Walworth and Kenosha Counties, the most remarkable of which was a state natural area, Lulu Lake, in Walworth County.

The site includes a large, high quality bog.

The site includes a large, high quality bog.

For once, the songs of the sphagnum ground crickets were not buried among those of Say’s trigs, as has been true at other bogs I have visited.

The bog is surrounded by hills of glacial outwash gravel.

The bog is surrounded by hills of glacial outwash gravel.

Those hills held two species I have not observed elsewhere in Walworth County to date: tinkling and spotted ground crickets. The former seem to prefer dry woodland edges on well drained sandy soils. The latter like moist shaded spots in woods on well drained sandy soils.

The bog is reached from the south by a long walk through a meadow with a good percentage of prairie plants. This curve-tailed bush katydid was a resident of that meadow.

The bog is reached from the south by a long walk through a meadow with a good percentage of prairie plants. This curve-tailed bush katydid was a resident of that meadow.

One more photo from the Wisconsin trip comes from an Interstate rest stop, also in Walworth, where I found my first straight-lanced meadow katydids for that county.

Nancy Collins had remarked that the straight-lanced males on her site had brown cerci. Going back, I find that this is true of every male I have photographed over the years.

Nancy Collins had remarked that the straight-lanced males on her site had brown cerci. Going back, I find that this is true of every male I have photographed over the years.

This was a very productive trip, resulting in 22 county records for the three days.

 

Illinois’ Kankakee Sands

by Carl Strang

In the Chicago region when someone mentions the Kankakee Sands, usually they are referring to the Nature Conservancy project in Newton County, Indiana. There is, however, a nature preserve in southeastern Kankakee County, Illinois, also known as “Kankakee Sands,” which also is worth knowing about.

The preserve has very high quality oak savanna and prairie ecosystems.

The preserve has very high quality oak savanna and prairie ecosystems.

I paid my first visit to this site on Friday, and left with a good dozen singing insect county records.

Most species were sand-soil singers I had encountered before, but this was my first sprinkled grasshopper.

Most species were sand-soil singers I had encountered before, but this was my first sprinkled grasshopper.

He was buried in a grass clump, offering no chance of a good photo. Fortunately he was open to climbing onto my finger for a portrait. The all-black pronotum sides are unique.

The most common orthopterans were tinkling ground crickets and straight-lanced meadow katydids, unsurprising on this sand soil.

One of the many male straight-lanceds from Friday.

One of the many male straight-lanceds from Friday.

I was pleased also to find that my new friend the handsome grasshopper is common there.

Handsome grasshopper, male.

Handsome grasshopper, male.

Female handsome grasshoppers were a bit bigger and green rather than brown.

Female handsome grasshoppers were a bit bigger and green rather than brown.

Both mottled sand grasshoppers and Boll’s grasshoppers also were there, the former often punctuating the scenery with their bright yellow hind wings in flight.

Boll’s grasshopper also has yellow hind wings. These are concealed when both species are at rest.

Boll’s grasshopper also has yellow hind wings. These are concealed when both species are at rest.

There also were plenty of bush katydids.

Most were curve-tailed bush katydids.

Most were curve-tailed bush katydids.

One, slightly smaller, proved to be a male fork-tailed bush katydid.

One, slightly smaller, proved to be a male fork-tailed bush katydid.

Kankakee Sands are worth a visit on either side of the state line.

 

Gensburg-Markham Prairie

by Carl Strang

One day last week I drove down to southern Cook County for singing insect survey work. I quickly found confused ground crickets for a county record in the Palos area, then proceeded to the Gensburg-Markham Prairie, which proved so fruitful that it occupied the rest of the afternoon. The dominant sound in that high-quality nature preserve was the buzzing of common meadow katydids.

The name is deceptive. This is one of the few places I have found to date where Orchelimum vulgare indeed is abundant.

The name is deceptive. This is one of the few places I have found to date where Orchelimum vulgare indeed is abundant.

There were other dry-habitat species present as well. I was able to add county records for woodland meadow katydid (my northernmost to date) and for straight-lanced meadow katydid.

This straight-lanced female was content to explore my finger and pose.

This straight-lanced female was content to explore my finger and pose.

The richest portion of the site’s singing insect fauna was the subfamily of stridulating slant-faced grasshoppers. I took lots of photos, thinking I had found the mother lode of species. When I examined them closely, however, the diversity turned out to be mainly within species, and I concluded that most of them in fact were marsh meadow grasshoppers.

Here is a classic adult marsh meadow grasshopper.

Here is a classic adult marsh meadow grasshopper.

But then there were a number of these. After much study I had to conclude that this, too, was an adult marsh meadow grasshopper.

But then there were a number of these. After much study I had to conclude that this, too, was an adult marsh meadow grasshopper.

Ditto for this one.

Ditto for this one.

Even more color variation was provided by nymphs. Again, I think they were marsh meadow grasshoppers.

This one in particular was strikingly colored.

This one in particular was strikingly colored.

And this individual seems aimed toward the pattern of the third adult above.

And this individual seems aimed toward the pattern of the third adult above.

These were my first of the species in Cook County, so they were a happy find. Two other grasshoppers also were my first for the county.

Wetter areas had plenty of short-winged green grasshoppers like this female.

Wetter areas had plenty of short-winged green grasshoppers like this female.

Prize of the day was this critter, the first spotted-wing grasshopper I have seen anywhere.

Prize of the day was this critter, the first spotted-wing grasshopper I have seen anywhere.

She wasn’t giving me good angles for photography, but fortunately I got a clear shot of the dorsal pronotum.

She wasn’t giving me good angles for photography, but fortunately I got a clear shot of the dorsal pronotum.

The inward-curving margins and their posterior big black triangles point to the two local species of Orphulella. There are two cuts in the dorsal surface, which point to O. pelidna rather than its close relative the pasture grasshopper O. speciosa.

This prairie is one I intend to visit in all portions of the singing insect season.

Memorial Forest Clearing

by Carl Strang

The Memorial Forest is a public site, essentially an undeveloped county park, in my home county of Marshall, in Indiana. As I have spent much of my time in that county over the years, my list of its singing insects is nearly as complete as that for DuPage. I had never looked at the Memorial Forest, however. I went there recently. The forest itself, though of good quality, had nothing new to add, but there is a cleared power line right-of-way through the forest which produced 4 county records, including a species I had not encountered before.

What made the clearing unusual was its sand soil.

What made the clearing unusual was its sand soil.

The nearly pure sand hosted oddities including velvet ants and a tiger beetle much larger than most species of my acquaintance. Almost right away I found my new friend, the woodland meadow katydid, and after a while ran across a species that may prove to be a frequent associate, at least in this region, as Lisa Rainsong has suggested.

A male straight-lanced meadow katydid.

A male straight-lanced meadow katydid.

There were large numbers of band-winged grasshoppers (the subfamily of grasshoppers which have wing-rattling flight displays, and thus qualify as singing insects). These ultimately sorted out to three species. In addition to the ubiquitous, and large, Carolina grasshopper, there were a medium sized and a small species.

The medium sized one was the mottled sand grasshopper, which I mentioned in a recent post on Jasper County.

The medium sized one was the mottled sand grasshopper, which I mentioned in a recent post on Jasper County.

Mottled sand grasshoppers were the most abundant singing insects in the clearing, their yellow hind wings flashing all around me as I walked. Then I noticed smaller bursts of bright red, and they led me to a grasshopper which up to that moment had been on my hypothetical list for the region.

You can get a sense of the red colored wings, and the small size of this insect, in comparison to my thumbnail. As usual, I released it unharmed.

You can get a sense of the red colored wings, and the small size of this insect, in comparison to my thumbnail. As usual, I released it unharmed.

The head and pronotum are beautifully patterned.

The head and pronotum are beautifully patterned.

This is the longhorn band-winged grasshopper, Psinidia fenestralis.

This is the longhorn band-winged grasshopper, Psinidia fenestralis.

The unusually wide black zone of the hind wing, the long, flattened antennae, and the banded yellow and black tibias, are additional features of this species. Old records placed it in the dune areas around the edge of Lake Michigan, so this well-inland site is unusual.

 

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