Crabs dredged off the Ogasawara Islands (Crustacea, Decapoda, Brachyura

Hironori Komatsu

Crabs of the families Xanthidae, Domeciidae, Trapeziidae, Pinnotheridae, and Pilumn-idae dredged off the Ogasawara Islands are recorded, with some taxonomic and biogeographic notes. Of 43 species (26 of the Xanthidae, 1 of the Domeciidae, 1 of the Trapeziidae, 1 of the Pin-notheridae, and 14 of the Pilumnidae) recorded in the present paper, 8 species are new to Japanese waters, and one species is new to science. The new species of the Pilumnidae, Nanopilumnus modestus, is described with photographs and line drawings. The biogeographic characteristics of these crab groups off the Ogasawara Islands are briefly discussed at the taxonomic comments of the species examined.

A new spider crab of the genus Rochinia A. Milne-Edwards, 1875 of the family Epialt- idae is described based on the specimens from offshore depths of Miyake Island in the Izu Islands, central Japan. The new species named R. miyakensis is most characterized and readily distin- guished from the congeners by having the long and widely divergent rostral spines, the smooth and sparsely setose dorsal surface of the carapace, a sharp hepatic tubercle, a sharp epibranchial tubercle, and a rounded postorbital lobe.

A new species of the freshwater crab, heretofore assigned to Geothelphusa miyazakii, is described. The new species is distinguished from G. miyazakii sensu stricto by characters of the first gonopod and carapace. Geothelphusa marginata, new species is restricted to Ishigaki and Iriomote Islands, Yaeyama Group, Southern Ryukyus, Japan. Populations of G. marginata on each island have minor differences in carapace

国立科博専報，（47）：219–277，2011年4月15日 Mem. Natl. Mus. Nat. Sci., Tokyo, (47): 219–277, April 15, 2011 Crabs dredged off the Ogasawara Islands (Crustacea, Decapoda, Brachyura) Hironori Komatsu Department of Zoology, National Museum of Nature and Science, 3–23–1 Hyakunincho, Shinjuku-ku, 169–0073 Tokyo, Japan E-mail: h-komatu@kahaku.go.jp Abstract. Brachyuran crabs dredged off the Ogasawara Islands, southern Japan, are reported. The collection consists of 73 species of 16 families excluding pilumnoid and xanthoid crabs. Three new species, Cryptodromia maculata sp. nov. (Dromiidae), Ebalia koyo sp. nov. (Leucosiidae), and Pugettia ogasawaraensis sp. nov. (Epialtidae), are described and illustrated. Fourtytwo species are newly added to the carcinological fauna of the Ogasawara Islands including 5 species new to Japanese waters. Biogeography of the recorded species is brieﬂy discussed. Key words: Crustacea, Decapoda, Brachyura, new species, Ogasawara Islands. Introduction The Ogasawara (Bonin) Islands is located ca. 1,000 km south from Tokyo and an oceanic islands that have never been connected to continent landmasses (Fig. 1). Its terrestrial fauna and ﬂora harbour a number of endemic species, but only a few marine organisms are known to be endemic to the islands, e.g. two species of hermit crabs (Asakura, 1991; Asakura et al., 1994a, b) and ﬁve species of marine gastropods (Fukuda, 1993, 1994, 1995). However, faunal study of marine organisms having planktonic larval stage at the isolated islands is important to realize the potential ability to disperse and speciation. Many authors have reported on taxonomy of brachyuran crabs collected from the Ogasawara Islands and the knowledge of brachyuran fauna of the islands is increasing. However, organized faunal survey by using dredge at the sea around the Ogasawara Islands is restricted to Takeda (1977). He recorded 28 species from 12 stations at the west side of Chichi-jima Island. Of these 5 species were new to science and 9 were new to Japanese waters. Other published works on brachyuran taxonomy on the basis of material from the Ogasawara Islands include Stimpson (1858, 1907), Rathbun (1894), Hirota (1894), Matsuura (1894), Yoshihara (1901), Terazaki (1902–1905), Parisi (1918), Balss (1922), Odhner (1925), Sakai (1938, 1939, 1955, 1976), Miyake (1939, 1970), Takeda and Miyake (1968a, b, 1969a, b, 1976), Miyake and Takeda (1969, 1970), Imajima (1970), Ooishi (1970), Shigei (1970), Suzuki (1972), Takeda (1973), Türkay (1974), Takeda and Kurata (1976a, b, 1977a, b, 1984), Takeda and Tamura (1980, 1981), Asakura et al. (1994b), Takeda and Tachikawa (1995), Castro (2000), Marumura and Kosaka (2003), Kobayashi (2005), Komai et al. (2006), Komatsu and Takeda (2009), Castro and Ng (2010), and Nguyen and Ng (2010). The National Museum of Nature and Science, Tokyo, conducted a zoological survey at the sea around the Ogasawara Islands during the years 2008–2010. The benthic animals were sampled by dredging at the depths of 36–3579 m along the whole Ogasawara Islands and Kaikata Seamount, which is located 115 km west of Haha-jima Island. Among a total of 136 stations, 60 stations were positive for brachyuran crabs at the depths of 36–499 m. The collections of brachyuran crabs are referred to 73 species of 16 families except pilumnoid and xanthoid crabs, which are under studying by Prof. Takeda of the Teikyo Heisei University. Of these three new species are de- 220 Hironori Komatsu Fig. 1. Maps showing the survey area, the Ogasawara Islands (upleft reproduced from Nakano et al. (2009)). Crabs dredged off the Ogasawara Islands 221 Table 1. Sampling data of R/V Koyo 2008, 2009 and 2010 cruise. Abbreviations for sampling gears: BD: biological dredge of 50 cm span opening; RD: rocky dredge of 50 cm span opening; 1m RD: rocky dredge of 1 m span opening. The stations with asterisk are located outside the map. Station Locality Position in Position out Depth (m) Date Gear 154–152 88–88 56–62 136–135 150–152 83–81 70–69 49–48 175–176 54–52 95–98 150–151 129–127 84–87 24 Oct. 2008 24 Oct. 2008 27 Oct. 2008 28 Oct. 2008 28 Oct. 2008 28 Oct. 2008 29 Oct. 2008 29 Oct. 2008 29 Oct. 2008 29 Oct. 2008 29 Oct. 2008 30 Oct. 2008 30 Oct. 2008 30 Oct. 2008 BD RD RD RD BD RD BD BD BD BD BD BD BD BD KY-08-03 KY-08-06 KY-08-11 KY-08-12 KY-08-13＊ KY-08-15 KY-08-17 KY-08-18 KY-08-19＊ KY-08-20 KY-08-21 KY-08-22 KY-08-25 KY-08-26 West of Nishi-jima I. West of Chichi-jima I. West of Chichi-jima I. South of Chichi-jima I. South of Chichi-jima I. West of Chichi-jima I. East of Chichi-jima I. East of Chichi-jima I. East of Chichi-jima I. East of Chichi-jima I. East of Chichi-jima I. West of Ototo-jima I. West of Nishi-jima I. West of Chichi-jima I. 27°07.45′N 142°06.38′E 27°04.64′N 142°08.52′E 27°03.62′N 142°08.89′E 27°00.19′N 142°11.61′E 26°58.07′N 142°09.07′E 27°04.55′N 142°09.16′E 27°05.02′N 142°15.22′E 27°05.01′N 142°14.89′E 27°06.07′N 142°18.56′E 27°04.23′N 142°15.19′E 27°03.84′N 142°15.44′E 27°10.65′N 142°07.27′E 27°07.31′N 142°07.70′E 27°04.84′N 142°08.93′E 27°07.86′N 142°06.90′E 27°04.69′N 142°08.68′E 27°03.66′N 142°08.88′E 27°00.21′N 142°11.74′E 26°57.81′N 142°09.06′E 27°04.73′N 142°09.31′E 27°04.86′N 142°15.22′E 27°04.88′N 142°14.87′E 27°06.06′N 142°18.76′E 27°04.22′N 142°15.06′E 27°03.70′N 142°15.23′E 27°10.71′N 142°07.37′E 27°07.03′N 142°07.64′E 27°04.98′N 142°09.15′E KY-09-07 KY-09-08 KY-09-13 KY-09-14 KY-09-21 KY-09-27 KY-09-28 KY-09-29 KY-09-30 KY-09-31 KY-09-33 KY-09-34 West of Minami-jima I. North of Haha-jima I. South of Haha-jima I. South of Haha-jima I. Northwest of Ototo-jima I. East of Ani-jima Channel East of Nishi-jima I. South of Nishi-jima I. East of Nishi-jima I. Southeast of Chichi-jima I. West of Minami-jima I. West of Minami-jima I. 27°01.72′N 142°07.39′E 26°45.20′N 142°06.44′E 26°34.10′N 142°10.79′E 26°34.03′N 142°10.80′E 27°13.09′N 142°09.19′E 27°06.29′N 142°13.88′E 27°07.05′N 142°10.68′E 27°06.59′N 142°10.25′E 27°07.22′N 142°10.60′E 27°00.84′N 142°16.81′E 27°02.19′N 142°07.26′E 27°02.34′N 142°07.52′E 27°01.93′N 142°07.28′E 138.2–136 26°45.38′N 142°06.55′E 98.3–102.4 96.5 26°34.04′N 142°10.81′E 92–93.1 27°13.19′N 142°09.23′E 135.8–135.5 27°06.28′N 142°14.01′E 81–83.4 27°07.02′N 142°10.69′E 52.1–52 27°06.58′N 142°10.21′E 60.7–60.3 27°07.28′N 142°10.58′E 51.6–49.9 27°00.92′N 142°16.56′E 311.4–299.7 27°02.27′N 142°07.27′E 136.5–136.6 27°02.55′N 142°07.34′E 138.9–140.9 10 Jul. 2009 13 Jul. 2009 14 Jul. 2009 14 Jul. 2009 15 Jul. 2009 15 Jul. 2009 15 Jul. 2009 16 Jul. 2009 16 Jul. 2009 16 Jul. 2009 16 Jul. 2009 16 Jul. 2009 BD BD BD BD BD BD BD BD BD BD BD BD KY-10-02 KY-10-06 KY-10-07 KY-10-11 KY-10-19 KY-10-23＊ KY-10-24 KY-10-25 KY-10-26 KY-10-27 KY-10-30 KY-10-31 East of Haha-jima I. North of Haha-jima I. South of Ane-jima I. Anejima-dashi Bank West of Chihi-jima I. East of Chichi-jima I. East of Nishi-jima I. Futami Bay, Chihi-jima I. Futami Bay, Chihi-jima I. South of Nishi-jima I. South of Chichi-jima I. West of Chihi-jima I. 26°41.44′N 142°10.33′E 26°44.29′N 142°06.23′E 26°31.60′N 142°08.85′E 26°38.98′N 142°11.85′E 27°04.82′N 142°08.95′E 27°06.22′N 142°18.82′E 27°07.23′N 142°10.70′E 27°04.77′N 142°11.68′E 27°04.68′N 142°11.72′E 27°06.65′N 142°10.42′E 27°00.14′N 142°12.11′E 27°05.18′N 142°08.48′E 26°41.33′N 142°10.39′E 115.3–114.8 26°44.29′N 142°06.37′E 76–72.9 26°31.60′N 142°08.94′E 104.7–99 26°38.95′N 142°11.96′E 93.9–82.5 27°04.75′N 142°09.06′E 86.9–90.5 27°06.04′N 142°18.69′E 178.4–179.3 27°07.14′N 142°10.73′E 47.4–50.8 27°04.76′N 142°11.73′E 42.2–41.6 27°04.68′N 142°11.56′E 36.2–40.4 27°06.61′N 142°10.29′E 59–60.1 27°00.16′N 142°11.91′E 146.8–138.7 27°05.12′N 142°08.39′E 96.8–96.5 5 Jul. 2010 5 Jul. 2010 6 Jul. 2010 6 Jul. 2010 7 Jul. 2010 8 Jul. 2010 8 Jul. 2010 9 Jul. 2010 9 Jul. 2010 9 Jul. 2010 9 Jul. 2010 9 Jul. 2010 BD BD BD BD 1m RD BD BD BD BD BD BD BD scribed and illustrated, 42 species are newly added to the carcinological fauna of the Ogasawara Islands including 5 species new to Japanese waters. Materials and Methods This study primarily deals with the specimens of brachyuran crabs collected from the sea around the Ogasawara Islands, using R/V Koyo of the Tokyo Metropolitan Ogasawara Fisheries Center 222 Hironori Komatsu Table 2. Sampling data of R/V Tansei Maru KT-09-2 cruise. Abbreviations for sampling gears: 1m BD: biological dredge of 1 m span opening; CBD: chain bag dredge of 50 cm span opening. The station with asterisk is located outside the map. Station KT-09-2-KK1-2(1) KT-09-2-TW1-1 KT-09-2-TW1-3 KT-09-2-TW1-5 KT-09-2-TW1-6 KT-09-2-TW2-2 KT-09-2-TW2-3 KT-09-2-TW2-4 ＊ Locality Position in Position out Depth (m) Date Gear Kaikata Seamount West of Chichi-jima I. West of Chichi-jima I. West of Chichi-jima I. West of Chichi-jima I. West of Chichi-jima I. West of Chichi-jima I. West of Chichi-jima I. 26°40.00′N 140°55.54′E 27°01.40′N 142°07.41′E 27°01.45′N 142°05.79′E 27°01.44′N 142°06.14′E 27°01.43′N 142°05.50′E 27°03.00′N 142°05.00′E 27°03.03′N 142°05.29′E 27°02.94′N 142°07.17′E 26°39.99′N 140°55.63′E 27°01.36′N 142°07.47′E 27°01.48′N 142°05.87′E 27°01.38′N 142°06.18′E 27°01.35′N 142°05.56′E 27°02.99′N 142°05.04′E 27°03.00′N 142°05.40′E 27°02.95′N 142°07.25′E 172.5–165 145.2–138.6 221.3–194.1 173.1–188.3 297.3–310.4 181.8–186.6 165.6–165.6 140.7–151.5 16 Mar. 2009 19 Mar. 2009 19 Mar. 2009 19 Mar. 2009 19 Mar. 2009 19 Mar. 2009 19 Mar. 2009 19 Mar. 2009 CBD 1m BD 1m BD 1m BD 1m BD 1m BD 1m BD 1m BD Table 3. Sampling data of TR/V Shin’yo Maru 2009 cruise. Abbreviations for sampling gear: BD: biological dredge of 50 cm span opening. Station Locality Position in Position out Depth (m) Date Gear SY-09-01 SY-09-02 SY-09-03 SY-09-04 SY-09-05 SY-09-06 SY-09-07 SY-09-08 SY-09-09 SY-09-11 SY-09-18 SY-09-19 SY-09-20 SY-09-21 East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. East of Muko-jima I. West of Chichi-jima I. West of Nishi-jima I. West of Ototo-jima I. West of Ototo-jima I. 27°44.05′N 142°09.19′E 27°44.55′N 142°09.69′E 27°44.86′N 142°10.17′E 27°44.99′N 142°10.52′E 27°45.10′N 142°11.05′E 27°45.42′N 142°11.83′E 27°45.75′N 142°12.54′E 27°41.06′N 142°10.58′E 27°41.13′N 142°11.11′E 27°42.02′N 142°12.46′E 27°06.11′N 142°08.89′E 27°07.62′N 142°07.35′E 27°09.74′N 142°06.76′E 27°12.80′N 142°05.13′E 27°44.01′N 142°09.15′E 27°44.47′N 142°09.76′E 27°44.72′N 142°10.21′E 27°44.79′N 142°10.40′E 27°44.78′N 142°10.89′E 27°45.13′N 142°11.57′E 27°45.41′N 142°12.10′E 27°41.05′N 142°10.40′E 27°41.00′N 142°10.88′E 27°42.16′N 142°12.07′E 27°06.07′N 142°09.06′E 27°07.62′N 142°07.60′E 27°09.72′N 142°06.83′E 27°12.81′N 142°05.33′E 109–108 122–123 146–144 159–152 193–172 301–237 499–365 106–98.7 124–112 172–161 101–98 144–140 157–159 161–159 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 16 Nov. 2009 18 Nov. 2009 18 Nov. 2009 18 Nov. 2009 18 Nov. 2009 BD BD BD BD BD BD BD BD BD BD BD BD BD BD (used gears include biological dredge of 50 cm span opening and rocky dredge of 50 cm and 1 m span openings), R/V Tansei Maru of the Japan Agency for Marine-Earth Science and Technology (JAMSTEC) (used gears include biological dredge of 1 m span opening and chain bag dredge with 50 cm span opening) and TR/V Shin’yo Maru of the Tokyo University of Marine Science and Technology (used gear was biological dredge of 50 cm span opening). Data of sampling stations of R/V Koyo, R/V Tansei Maru and TR/V Shin’yo Maru are summarized in Table 1, Table 2 and Table 3, respectively. Specimens were ﬁxed with 80% ethanol or rarely 10% formalin and preserved in 75% ethanol. All the specimens examined are deposited in the Showa Memorial Institute, National Museum of Nature and Science, Tsukuba (NSMT-Cr S). Superfamilial and familial arrangements follow Ng et al. (2008). Synonymy is listed only the record from the Ogasawara Islands. Measurements of specimens are provided for maximum carapace length (cl) by carapace width (cw) including rostral and lateral spines in millimeters (mm). In the majoid crabs, the carapace length was measured from the level of the base of the rostral spines to the posterior margin of carapace and the carapace width was measured excluding the lateral spines. Other abbreviations used in the text are: G1, ﬁrst male gonopod; G2, second male gonopod; Jn, Japanese name; P2–5, second to ﬁfth pereiopods (ﬁrst to fourth ambulatory legs). Crabs dredged off the Ogasawara Islands Taxonomic accounts Section Podotremata Guinot, 1977 Superfamily Dromioidea De Haan, 1833 Family Dromiidae De Haan, 1833 Cryptodromia maculata sp. nov. [New Japanese name: Komon-iso-kaikamuri] (Figs. 2A, 3) 223 Materials examined. Holotype: male (cl 5.3 × cw 5.6 mm), NSMT-Cr S 875, KY-08-26, West of Chichi-jima I., Ogasawara Is., 27°04.8′ N 142°08.9′E – 27°05.0′N 142°09.2′E, 84–87 m, biological dredge, 30 October 2008. Paratypes: 2 males (3.7×3.9, 4.8×5.0), NSMT-Cr S 876, KY-08-15, South of Chichi-jima I., 2 7 °0 4 . 5 5 ′ N 1 4 2 °0 9 . 1 6 ′ E – 2 7 °0 4 . 7 3 ′ N Fig. 2. A, Cryptodromia maculata sp. nov., holotype male (5.3×5.6), NSMT-Cr S 875; B, Epigodromia rotunda McLay, 1993, young female (5.6×5.5), NSMT-Cr S 881; C, Homalodromia coppingeri Miers, 1884, young female (4.6×3.9), NSMT-Cr S 882; D, Lauridromia intermedia (Laurie, 1906), male (29.4×29.2), NSMT-Cr S 883; E, Takedromia yoshidai (Takeda and Kurata, 1976), young female (5.5×5.4), NSMT-Cr S 884; F, Dynomene pilumnoides Alcock, 1900, male (7.6×9.4), NSMT-Cr S 887. 224 Hironori Komatsu 142°09.31′E, 83–81 m, rocky dredge, 28 October 2008; 1 male (4.2×4.4), NSMT-Cr S 877, KY08-25, West of Chichi-jima I., 27°07.31′ N 142°07.70′E – 27°07.03′N 142°07.64′E, 129–127 m, biological dredge, 30 October 2008; 1 young female (2.7×2.7), NSMT-Cr S 878, KY-09-28, East of Nishi-jima I., 27°07.05′N 142°10.68′E – 27°07.02′N 142°10.69′E, 52.1–52 m, biological dredge, 15 July 2009; 1 young female (2.8×2.7), NSMT-Cr S 879, KY-09-30, East of Nishi-jima I., 2 7 °0 7 . 2 2 ′ N 1 4 2 °1 0 . 6 0 ′ E – 2 7 °0 7 . 2 8 ′ N 142°10.58′E, 51.6–49.9 m, biological dredge, 16 July 2009. Non-type: 1 juv., KY-09-21, NSMT-Cr S 880. Description of male. Carapace (Figs. 2A, 3A) slightly wider than long, subquadrangular, very convex dorsally, smooth under cover of short setae, longer plumose setae near margins. Only branchial groove faintly marked. Rostrum tridentate; median tooth slender, deﬂexed; lateral teeth broadly triangular, horizontal. Anterolateral margin begins at level of postorbital tooth, armed with one triangular tooth just behind postorbital tooth, straight and slightly convergent behind tooth, interrupted only by branchial groove, without posterolateral tooth. Supraorbital tooth very small, triangular; postorbital tooth acutely triangular, as large as median rostral tooth. Broadly triangular suborbital tooth, divided from postorbital tooth by short orbital ﬁssure; subhepatic area ﬂattened to accommodate cheliped when folded. First segment of antenna (Fig. 3A) wider than long, beaked medially, lateral margin shorter than medial margin. Second segment broad basally, tapering, 1.5 times longer than wide basally, with low tubercle disto-laterally; medial margin curved, distally produced as a tongue-shaped process; exopod bluntly spine-shaped, ﬁrmly ﬁxed to second segment, tip extends as far as medial process. Ratio of length of antennal ﬂagellum to CW = 0.56. Epistom triangular, wider than long, weakly concave. Basis of third maxilliped fused with ischium, but with suture; merus dentate on inner margin; palp exposed, slightly longer than merus; crista dentata consists of 4 or 5 teeth. Chelipeds moderate, 1.5 times as long as carapace in holotype; merus trigonal in cross-section, unarmed; carpus with 2 triangular tubercles on distal border, superior tubercle twice as large as inferior one; propodus (Fig. 3C) with small tubercle matching superior tubercle of carpus, with few granules on distal part of superior surface (not seen in ﬁgure), lower margin upcurved; ﬁngers hollowed out internally, armed with rounded teeth on both cutting edges, with few granules on distal part of superior surface of dactyli, gaping basally. P2 and P3 (Fig. 3D) shorter than chelipeds, fringed with longer plumose setae; meri with blunt, subdistal tooth on superior margin; propodi with strong distal lobe; dactyli 1.1 times longer than propodi, narrow, talon-like, inner margins armed with 4 or 5 small spines, increasing in size distally. P4 (Fig. 3E) reduced; dactyli strongly curved, opposed by single propodal spine with another smaller spine on outer propodal margin. P5 (Fig. 3F) reduced, subdorsal; dactyli opposed by 2 propodal spines (opposite one hidden in figure) with 2 sbuequal and 1 smaller spines on outer propodal margin. Abdomen (Fig. 3G) of six free somites. Male telson 1.2 times wider than long, subtriangular, constricted subdistally, rounded at tip. Uropod plates well developed, ovoid, visible externally. Abdominal locking mechanism consists of uropod plates ﬁtting in front of small ﬂange on bases of P2. G1 (Fig. 3H) basal segment lobular, with simple setae on both margins; distal segment with median suture on sternal surface, tip acutely triangular with dense plumose setae. G2 (Fig. 3I) simple, needle-like, slightly longer than G1. Color. Whole body yellow, dotted with small orange spots (Fig. 2A). Camouflage. Most of crabs carried sponge caps. Etymology. The speciﬁc name is derived from the Latin maculata (= spotted) alluding to the small orange spots on the carapace. Crabs dredged off the Ogasawara Islands 225 Fig. 3. Cryptodromia maculata sp. nov., holotype male (5.3×5.6), NSMT-Cr S 875. A, carapace (setae omitted), dorsal; B, frontal region, ventral; C, right chela, outer; D, right P3, dorsal; E, right P4, dorsal; F, right P5, dorsal; G, abdomen, vental; H, right G1, ventral; I, right G2, ventral. Scales 1 mm. Remarks. Cryptodromia maculata sp. nov. resembles C. hilgendori de Man, 1888 in the subquadrangular carapace having a single anterolat- eral tooth, but C. maculata can be distinguished from C. hilgendori by the acutely triangular postorbital tooth (vs. only pointed in C. hilgendori), 226 Hironori Komatsu the anteriorly directed, triangular anterolateral tooth (vs. subtruncate in C. hilgendorfi), two propodal spines opposing dactyli of P5 (vs. single in C. hilgendori) (cf. Campbell and Stephenson, 1970; McLay, 1993). Distribution. Known only from the Ogasawara Islands, occurring at the depths of 50–129 m. Epigodromia rotunda McLay, 1993 [New Jn: Marumi-hime-kaikamuri] (Fig. 2B) Material examined. SY-09-09, 1 young female (5.6×5.5), NSMT-Cr S 881. Camouﬂage. The present specimen did not carried sponge cap as in holotype. Remarks. The present specimen agrees well with the original description and illustrations by McLay (1993). Although the present specimen is larger than holotype (4.8×4.2 mm), it shows immature features, viz. the unopened vulva and the undeveloped pleopods. The first leg has small podobranch, so that this species is unapplicable to the keys to genera by McLay (1993). Distribution. Previously known only from New Caledonia (McLay, 1993). This record grately extends the geographical range northwards to the Ogasawara Islands. Occurring at the depths of 112–235 m. Homalodromia coppingeri Miers, 1884 [Jn: Hime-kinuge-kamuri] (Fig. 2C) Material examined. KY-08-20, 1 young female (4.6×3.9), NSMT-Cr S 882. Camouﬂage. The present specimen did not carried sponge cap. Distribution. Widely distributed in the IndoWest Pacific Ocean (McLay, 1993). This is the ﬁrst record from the Ogasawara Islands. Lauridromia intermedia (Laurie, 1906) [Jn: Akage-kamuri] (Fig. 2D) Material examined. KY-10-27, 1 male (29.4× 29.2), NSMT-Cr S 883. Camouﬂage. The present specimen carried octocoral, Dendronephthya sp. Distribution. Western Indian Ocean to Australia, New Caledonia, Philippines, Taiwan, Japan. Occurring at the depths of 7–150 m (Ahyong et al., 2009). This is the ﬁrst record from the Ogasawara Islands. Takedromia yoshidai (Takeda and Kurata, 1976) [Jn: Wantone-kamuri] (Fig. 2E) Cryptodromia yoshidai Takeda and Kurata, 1976b: 19, pl. 1 ﬁg. 1, pl. 2 ﬁgs. 3, 4. Material examined. KY-09-29, 1 young female (5.5×5.4), NSMT-Cr S 884. Remarks. The members of the genus Takedromia show signiﬁcant sexual dimorphism, viz. the massive and elongate chelipeds and lacinated teeth of the anterolateral margin of the carapace in male. The present young female specimen looks very different from the original description and illustration by Takeda and Kurata (1976b), but belongs to T. yoshidai by the areolation of the carapace and the chelipedal carpus being with two subconical tubercles. Distribution. Ogasawara Islands, Philippines, Chesterfield Islands, occurring at the depth of 60.3–200 m (Takeda and Kurata, 1976b; McLay, 1993; Takeda and Manuel-Santos, 2006; this study). Family Dynomenidae Ortmann, 1892 Dynomene pilumnoides Alcock, 1900 [Jn: Akatoge-kaikamuri] (Fig. 2F) Materials examined. KY-08-13, 1 young female (3.4×3.7), NSMT-Cr S 885; KY-09-08, 1 male (5.7×7.0), NSMT-Cr S 886; SY-09-11, 1 male (7.6×9.4), NSMT-Cr S 887. Distribution. Widely distributed in the IndoWest Pacific Ocean (McLay, 1999). This is the ﬁrst record from the Ogasawara Islands. Superfamily Raninoidea De Haan, 1839 Family Raninidae De Haan, 1839 Notopoides latus Henderson, 1888 [Jn: Eri-asahigani-modoki] (Fig. 4A) Notopoides latus: Takeda and Kurata, 1984: 196, ﬁgs. 1, 2. Materials examined. KY-09-31, 1 young fe- Crabs dredged off the Ogasawara Islands male (12.9×9.1), NSMT-Cr S 888; KT-09-2TW1-6, 1 male with entopractans on inner surface of abdomen and gonopods (18.8×13.5), NSMTCr S 889. Remarks. The present specimens agree well with the original description by Henderson (1888) and the subsequent description by Dawson and Yaldwyn (2002), but differ in that the lower margin of the chelipedal palm has 2 teeth (instead of 3). This feature appears in the Japanese specimens (Sakai, 1980; Takeda and Kurata, 1984; 227 Baba, 1986; this study). Distribution. Widely distributed in the IndoWest Pacific Ocean (Dawson and Yaldwyn, 2002). Notopus dorsipes (Linnaeus, 1758) [Jn: Asahigani-modoki] (Fig. 4B) Material examined. KY-10-25, 1 male (26.2× 17.8), NSMT-Cr S 890. Distribution. Western Indian Ocean to Australia, Indonesia, China, Taiwan, Japan (Ahyong et Fig. 4. A, Notopoides latus Henderson, 1888, male (18.8×13.5), NSMT-Cr S 889; B, Notopus dorsipes (Linnaeus, 1758), male (26.2×17.8), NSMT-Cr S 890; C, Notosceles serratifrons (Henderson, 1893), male (12.9×7.2), NSMT-Cr S 891; D, Ranina ranina (Linnaeus, 1758), 1 juvenile (12.7×8.4), NSMT-Cr S 896; E, Umalia orientalis (Sakai, 1963), young female (9.4×6.7), NSMT-Cr S 898; F, Sakaila imperialis (Sakai, 1963), male (12.3×15.8), NSMT-Cr S 900. 228 Hironori Komatsu al., 2009). This is the ﬁrst record from the Ogasawara Islands. Notosceles serratifrons (Henderson, 1893) [Jn: Nokoha-asahi] (Fig. 4C) Materials examined. KY-08-25, 1 male (12.9× 7.2), NSMT-Cr S 891; KY-08-26, 1 specimen (broken on posterior body: 16.3×9.3), NSMT-Cr S 892; KY-10-23, 1 female (11.1×6.3), NSMTCr S 893; KT-09-2-KK1-2(1), 1 young female (broken), 1 juv. (5.1×2.7), NSMT-Cr S 894; SY09-19, 1 young female (7.8×4.1), NSMT-Cr S 895. Remarks. The present specimens agree with the original description by Henderson (1893), but differ in that 1) the rostrum is sometimes shouldered (generally triangular, but not mentioned in the original description) and 2) the immovable ﬁnger of the chela has 4 denticle on cutting edges (5 in the original description). Distribution. Sri Lanka to Australia, China, Taiwan, and Japan (Ahyong et al., 2009). This is the ﬁrst record from the Ogasawara Islands. Ranina ranina (Linnaeus, 1758) [Jn: Asahi-gani] (Fig. 4D) Material examined. KY-08-26, 1 juvenile (sex indetermined, posterior body and pereiopods lost except chelipeds and left P5; 12.7×8.4), NSMTCr S 896. Remarks. Although the present juvenile specimen appears quite different from the adult form in the front-orbital region, it agrees well with the illustration by Sakai (1937: Fig. 45c). Distribution. Widely distributed in the IndoWest Paciﬁc Oceans (Ahyong et al., 2009). This is the ﬁrst formal record from the Ogasawara Islands, but this species is well known to be caught for commercial ﬁsheries in this area. Umalia misakiensis (Sakai, 1937) [Jn: Misakiasahi-modoki] (Fig. 5A) Material examined. KY-10-02, 1 young female (9.6×7.3), NSMT-Cr S 897. Remarks. Despite the present specimens is still young, it agrees well with the original and subse- quent descriptions and illustrations by Sakai (1937, 1965) Distribution. Philippines and Japan, occurring at the depths of 100–150 m (K. Sakai, 2004). This is the ﬁrst record from the Ogasawara Islands. Umalia orientalis (Sakai, 1963) [Jn: Toyo-asahimodoki] (Figs. 4E, 5B) Material examined. SY-09-01, 1 young female (9.4×6.7), NSMT-Cr S 898. Comparative materials examined. Uwa Sea, Ehime Pref., Bungo Strait, Japan, 1 male (35.3× 26.7), NSMT-Cr R 2790, trawl, 10 Apr. 1966. Itoman, Okinawa I., Ryukyu Is., Japan, 1 male (23.5 ×17.6), NSMT-Cr 6502, 70–90 m, coll. Y. Nakasone, 26 Apr. 1974. Remarks. The present specimen (Fig. 4E) differs from the original description and illustration by Sakai (1963) by that (1) the rostrum is proportionally larger; (2) the distance between exorbital and anterolateral spines is relatively shorter; (3) a spine on the dorsal margin of the chelipedal palm is absent (vs. present). But these differences may be caused by that the present specimen is still young and small. The form of the front-orbital spines and the dactyli of the ambulatory legs are typical for U. orientalis (Fig. 5B) Distribution. China, Taiwan and Japan, occurring at the depths of 50–120 m (Ahyong et al., 2009). This is the ﬁrst record from the Ogasawara Islands. Section Eubrachyura Saint Laurent, 1980 Superfamily Aethroidea Dana, 1851 Family Aethridae Dana, 1851 Sakaila imperialis (Sakai, 1963) [Jn: Osachilagani] (Fig. 4F) Materials examined. KY-08-19, 1 young male (3.3×5.4), NSMT-Cr S 899; KT-09-2-TW1-3, 1 male (12.3×15.8), NSMT-Cr S 900. Distribution. Known only from Japan, occurring at the depths of 85–221 m (Miyake, 1983; this study). This is the ﬁrst record from the Ogasawara Islands. Crabs dredged off the Ogasawara Islands 229 Fig. 5. A, Umalia misakiensis (Sakai, 1937), young female (9.6×7.3), NSMT-Cr S 897; B, Umalia orientalis (Sakai, 1963), male (35.3×26.7), NSMT-Cr R 2790; C, Heteronucia toyoshioae Komatsu and Takeda, 2005, carapace (empty shell; 5.4×5.9), NSMT-Cr S 927; D, Oreotlos lagarodes Tan and Ng, 1996, young female (2.9×4.2), NSMT-Cr S 938; E, Chalaroachaeus curvipes de Man, 1902, ovig. female (3.5×3.6), NSMT-Cr S 1027; F, Thalamita oculea Alcock, 1899, young male (5.1×7.1), NSMT-Cr S 1136; G, Thalamita spinifera Borradaile, 1902, male (6.6 ×8.7), NSMT-Cr S 1139. Superfamily Dorippoidea MacLeay, 1838 Family Ethusidae Guinot, 1977 Ethusa quadrata Sakai, 1937 [Jn: Kaku-heikegani] (Fig. 6A) Material examined. KY-08-22, 1 young male (4.6×3.6), NSMT-Cr S 901. Distribution. Japan, East China Sea, Philippines, New Caledonia, occurring at the depths of 35–393 m (Castro, 2005). This is the ﬁrst record 230 Hironori Komatsu Fig. 6. A, Ethusa quadrata Sakai, 1937, young male (4.6×3.6), NSMT-Cr S 901; B, Dairoides kusei (Sakai, 1938), young male (7.9×10.6), NSMT-Cr S 902; C, Henicoplax nitida (Miers, 1879), young male (3.7×5.6), NSMT-Cr S 903; D, Platyozius laevis (Borradaille, 1902), young male (4.3×4.6), NSMT-Cr S 906; E, Singhaplax styrax Castro, 2007, male (4.1×7.0), NSMT-Cr S 911; F, Goneplacoides marivenae (Komatsu and Takeda, 2003), male (5.6 ×8.5), NSMT-Cr S 909. from the Ogasawara Islands. Superfamily Eriphioidea MacLeay, 1838 Family Dairoididae Števčić, 2005 Dairoides kusei (Sakai, 1938) [Jn: Hoshihada-hishi-gani] (Fig. 6B) Dairoides kusei: Takeda and Kurata, 1976b: 27. Material examined. KY-09-21, 1 young male (7.9×10.6), NSMT-Cr S 902. Distribution. Philippines, Japan, Hawaii Is., occurring at the depths of 85–199 m (Takeda and Kurata, 1976b; Serène and Vadon, 1981; Ng and Tan, 1999). Crabs dredged off the Ogasawara Islands Superfamily Goneplacoidea MacLeay, 1838 Family Euryplacidae Stimpson, 1871 Henicoplax nitida (Miers, 1879) [Jn: Kiba-gani] (Figs. 6C, 8A–C) Heteroplax nitida: Takeda, 1977: 133. Materials examined. KY-08-17, 1 young male (3.7×5.6), NSMT-Cr S 903; SY-09-02, 1 young female (4.3×6.1), NSMT-Cr S 904. Additional material examined. Takinoura Bay, Ani-jima I., dredge, 45 m, coll. M. Takeda and M. Imajima, 15 Jun. 1976, 1 male (3.8×5.4), NSMTCr 4453. Remarks. Recently Castro and Ng (2010) established a new genus, Henicoplax, for this species and 4 other species. All the congeners show restricted distribution. Distribution. Japan, Korea Strait, Hong Kong, occurring at shallow tidal water to 123 m (Castro and Ng, 2010; this study). Platyozius laevis (Borradaille, 1902) [New Jn: Hira-enko-gani] (Fig. 6D) Platyozius laevis: Castro and Ng, 2010: 84, Figs. 33A–F; 34A–F; 35A–G; 38A–C. Materials examined. KY-08-20, 1 young male (3.7×4.1), NSMT-Cr S 905; KY-09-28, 1 young male (4.3×4.6), NSMT-Cr S 906. Remarks. Recently Castro and Ng (2010) reappraised this genus and species. Distribution. Widely distributed in the IndoWest Paciﬁc Ocean, occurring at intertidal water deep to 200 m (Castro and Ng, 2010). Family Goneplacidae MacLeay, 1838 Goneplacoides marivenae (Komatsu and Takeda, 2003) [New Jn: Medaka-gani-modoki] (Fig. 6F) Materials examined. KY-08-18, 1 young male (2.3×3.2), NSMT-Cr S 907; KT-09-2-KK1-2(1), 1 female (broken on carapace), 4 young females (3.2×4.3–3.4×4.4), NSMT-Cr S 908; KT-09-2TW1-5, 1 male (5.6×8.5), 1 female (5.8×8.8), NSMT-Cr S 909; KT-09-2-TW2-2, 1 young male (4.1×5.9), NSMT-Cr S 910. Remarks. Goneplax megalops Komatsu and 231 Takeda, 2003, was reduced to a junior subjective synonym of Goneplacoides marivenae (Komatsu and Takeda, 2003), by Castro (2007) because the former is a young form of G. marivenae. The specimens collected from the Japanese waters are much smaller than typical form of G. marivenae, but the male chelae being leaving a gape between the cutting edges, the developed male gonopod, and the broadened female abdomen which looks mature. However, ovigerous female specimen has not been collected from the Japanese waters. This supports Castro’s opinion, but color in life of the Japanese specimens is rather different from the specimen from New Caledonia described by Castro (2007). Difference of size at sexual maturity in populations from different localities is often fround in majid species of Schizophrys, Notomithrax and Leptomithrax (Grifﬁn and Tranter, 1986). Distribution. Japan, Taiwan, Philippines, Indonesia, Chesterﬁeld Is., New Caledonia, occurring at the depths of 48–660 m (Castro, 2007; this study). This is the ﬁrst record from the Ogasawara Islands and extends its bathymetrical range shallow to 48 m. Singhaplax styrax Castro, 2007 [Jn: Yamatomedaka-gani] (Fig. 6E) Material examined. KY-09-31, 1 male (4.1× 7.0), NSMT-Cr S 911; SY-09-06, 1 ovig. female (3.0×5.1), NSMT-Cr S 912; SY-09-20, 1 ovig. female (2.7×4.4), NSMT-Cr S 913. Distribution. Japan and Indonesia, occurring at the depths of 110–311.4 m (Castro, 2007; this study). This is the ﬁrst record from the Ogasawara Islands. Superfamily Leucosioidea Samouelle, 1819 Family Leucosiidae Samouelle, 1819 Ebalia hayamaensis Sakai, 1963 [Jn: Hayamaebalia] (Figs. 7A, 8J–L) Ebalia hayamaensis: Takeda, 1977: 134, ﬁg. 3A; Marumura and Kosaka, 2003: 25. Materials examined. KY-08-26, 1 male (4.1× 5.2), NSMT-Cr S 914; KY-09-14, 1 ovig. female (4.9×6.4), NSMT-Cr S 915; SY-09-09, 1 ovig. 232 Hironori Komatsu Fig. 7. A, Ebalia hayamaensis Sakai, 1963, male (4.1×5.2), NSMT-Cr S 914; B, Ebalia humilis Takeda, 1977, male (2.1×2.7), NSMT-Cr S 918; C, Ebalia sakaii Takeda and Miyake, 1972, ovig. female (cl 2.5 mm), NSMT-Cr S 923; D, Ebalia tuberculosa (A. Milne-Edwards, 1873), young male (4.7×4.7), NSMT-Cr S 925; E, Heteronucia perlata (Sakai, 1963), ovig. female (7.8×8.2), NSMT-Cr S 926; F, Toru granuloides (Sakai, 1961), male (12.7× 12.0), NSMT-Cr S 954. female (4.7×6.1), NSMT-Cr S 916. Supplementary description. Male abdomen narrow, elongate triangular. Somites 1 and 2 very short, transversely rectangular. Somites 3 to 6 fused together, elongate trapezoidal, with small triangular tooth near distal end. Telson elongate tongue-shaped. G1 (Fig. 8J, K) straight, with semicircular lobe Crabs dredged off the Ogasawara Islands 233 Fig. 8. Right gonopods. A–C, Henicoplax nitida (Miers, 1879), male (3.8×5.4), NSMT-Cr 4453, A, G1, abdominal; B, tip of same; C, G2, abdominal. D, E, Toru granuloides (Sakai, 1961), male (12.7×12.0), NSMT-Cr S 954, D, G1, abdominal; E, tip of same. F–I, Hyastenus ambonensis Grifﬁn and Tranter, 1986, males (7.1×5.4), NSMT-Cr S 957, F, G1, abdominal; G, tip of same, sternal; H, mesial; I, G2, abdominal. J–L, Ebalia hayamaensis Sakai, 1963, male (4.1×5.2), NSMT-Cr S 914, J, G1, abdominal; K, tip of same; L, G2, abdominal. M, N, Naxioides taurus (Pocock, 1890), male (17.6×12.8), NSMT-Cr 15394, M, G1, abdominal; N, G2, abdominal. O–Q, Prismatopus occidentalis (Grifﬁn, 1970), male (7.1×5.3), NSMT-Cr S 1077, O, G1, abdominal; P, tip of same; Q, G2, abdominal. Scales for A, C, E–J, L = 0.5 mm; B, K, P = 0.25 mm; D = 2 mm; M–O, Q = 1 mm. 234 Hironori Komatsu at midlength of mesial margin; distal part slightly dilated, obliquely subtruncate, with dense setae on disto-lateral margin. G2 (Fig. 8L) short, 0.3 times as long as G1, tip triangular. Remarks. Description and illustration of the male abdomen and gonopods are provided here for the ﬁrst time. Distribution. Known only from Japan (Sagami Bay, off Shiono-misaki Cape, Ryukyu Is., Ogasawara Is.), occurring at the depths of 65–124 m (Sakai, 1963; Marumura and Kosaka, 2003; this study). Ebalia humilis Takeda, 1977 [Jn: Ogasawaraebalia] (Fig. 7B) Ebalia humilis Takeda, 1977: 115, ﬁgs. 2 (A, B), 3 (B–D); Takeda and Kurata, 1984: 198; Marumura and Kosaka, 2003: 25. Praebebalia taeniata: Takeda and Kurata, 1977b: 141, fig. 1 (not Praebebalia taeniata Takeda, 1977). Materials examined. KY-08-18, 1 ovig. female (3.1×3.8), NSMT-Cr S 917; KY-08-21, 1 male (2.1×2.7), NSMT-Cr S 918; Ohmura beach, Chichi-jima I., intertidal water, coll. T. Komai, 17 Jul. 2009, 1 male (2.3×2.6), 1 ovig. female (2.7 ×3.3), NSMT-Cr S 919. Distribution. Known from Ogasawara Is. and Kermadec Is, occurring at intertidal water deep to 90 m (Komatsu and Takeda, 2007; this study). Ebalia koyo sp. nov. [New Jn: Koyo-ebalia] (Figs. 9–11) Materials examined. Holotype: male (cl 2.1 mm; damaged on left lateral angle), NSMT-Cr S 920, KY-09-21, NE Ototo-jima I., Ogasawara Is., 2 7 °1 3 . 0 9 ′ N 1 4 2 °0 9 . 1 9 ′ E – 2 7 °1 3 . 1 9 ′ N 142°09.23′E, 135.8–135.5 m, dredge, coll. H. Komatsu (R/V Koyo), 15 July 2009. Paratype: 1 female (2.5×3.5), NSMT-Cr S 921, off Chichi-jima I., Ogasawara Is., Japan, 100 m, coll. S. Nagai, December 1991. Additional material examined. Paratype: 1 male (2.2×2.8), NSMT-Cr S 922, off Yamatomura, Amami-oshima I., northern Ryukyu Is., Japan, 150 m, coll. S. Nagai, 5 May 1998. Description. Carapace (Figs. 9, 10) subpentagonal in general outline, 1.3 times broader than long in male, 1.4 times in female, convex dorsally; upper surface entirely covered with microscopic, flat granules. Front strongly produced, with pair of short ridge obliquely running from lateral angle towards midline, ridges not conﬂuent each other; margin divided into two lobes by median triangular notch. Orbit with two longitudinal sutures on dorsal roof; infraorbital lobe with V-shaped notch with short suture. Gastro-cardiac region raised, with pair of small gastric tubercles and faint cardiac tubercle. Intestinal region prominently convex, subtriangular in outline. Hepatic region scarcely swollen; border indistinct, inside from general outline. Pterygostomian margin obliquely straight, not lamellar, divided from branchial margin by broad V-shaped notch. Branchial region convex postero-laterally, sloping antero-laterally; margin lamellar, slightly upturned, antero-lateral margin divergent, with low broadly rounded lobe, making tongue-shaped angle with postero-lateral margin; postero-lateral margin convergent, with large triangular tooth in posterior half, with transverse bump on posterior margin of tooth in female. Posterior margin bilobed, triangular with rounded tip in male, as large as lateral angle of carapace, small and rounded in female. Ocular peduncle short. Antennule slightly obliquely folded in fossa; basal segment occupying ventral half, covered with round granules. Basal segment of antenna transversely ovoid; second segment subcylindrical, ﬁtting into orbital hiatus. Mandible (Fig. 11A, B) calciﬁed; cutting edge triangular in outline, weakly pointed medially; endopod palp three-segmented, terminal segment fringed with short setae. Maxillule (Fig. 11C): coxal endite subcylindrical, directed mesially, with some terminal setae; basial endite triangular, with stout, thin setae on mesial margin; endopod almost reduced, small, rounded with short terminal setae. Maxilla (Fig. 11D): coxal endite small, roundly bilobed; basial endite small, rounded, with tuft of terminal setae; endopod tongue- Crabs dredged off the Ogasawara Islands Fig. 9. 235 Ebalia koyo sp. nov., holotype male (cl 2.1 mm), NSMT-Cr S 920. shaped; exopod (scaphognathite) longitudinally expanded into ovate structure, entirely fringed with short setae. First maxilliped (Fig. 11E): coxal endite semiglobular, with dense setae; basial endite lobular, largely expanded into triangular structure, fringed with long setae; endopod lobular, longitudinally expanded, fitting in efferent channel, plicate on upper surface, with short setae along anterior half of mesial margin; exopod longitudinally ﬁliform, with long setae on distal part of mesial margin, bearing flagellum with tuft of long terminal setae. Second maxilliped (Fig. 11F): endopod with long setae along inner margins of merus and propodus and outer margin of propodus, dactylus fringed with stout setae around tip; exopod ﬁliform, tapering distally, with long setae on distal portion of mesial margin, bearing ﬂagellum with tuft of long terminal setae. Third maxilliped (Fig. 11G, H) entirely covered with microscopic, ﬂat granules; basis fused with ischium, but with remnant suture on internal surface; ischium 0.8 times as long as merus along mesial margin; merus elongate triangular; palp 3-segmented, with dense setae; exopod arcuated on lateral margin, with small triangular tooth on proximal end, fringed with very short setae on lateral margin, internal ridge stout. Cheliped (Figs. 9, 10) moderate, 1.3 times as long as carapace, entirely covered with microscopic, flat granules as on carapace; coxal condyles reduced in both sexes; merus triangular in cross-section, flat on dorsal surface, with some tubercles on outer and ventral margins and subdistal tubercle on inner margin, weakly convex on outer margin; carpus subgrobular; palm convex dorsally, with rounded, subproximal lobe and triangular, distal lobe on outer margin, with some tubercles on inner border; ﬁngers slender, tapering distally, with interlocking, minute teeth on both cutting edges; movable finger curved, 0.7 times as long as palm measured along outer margin. 236 Hironori Komatsu Fig. 10. Ebalia koyo sp. nov., A, B, paratype male (2.2×2.8), NSMT-Cr S 922; C, D, paratype female (2.5×3.5), NSMT-Cr S 921. Ambulatory legs (Fig. 11I) slender gradually decreasing in length from ﬁrst to fourth, similar in shape, covered with microscopic, ﬂat granules except on dactyli; coxal condyles reduced in both sexes; meri, carpi and propodi subcylindrical, covered with mushroom-like granules on outer and inner surfaces of meri and outer surface of carpi, with acute granules on inner surface of propodi; dactyli slender, subcylindrical, with small rounded dactylo-propodal lock on proximal border of dorsal surface. Male thoracic sternum covered with microscopic, flat granules as on carapace, episternites not divided entirely; ﬁrst to fourth sternites completely fused; abdominal cavity reaching to buccal cavern; median suture absent; transverse sutures between sternites 4/5, 5/6, 6/7 and 7/8 medially interrupted. Female thoracic sternum covered with microscopic, ﬂat granules as on carapace; ﬁrst to fourth sternites fused together; abdominal cavity reach- ing to buccal cavern; sutures as in males. Male abdomen (Fig. 11J) covered with microscopic, ﬂat granules as on carapace; somite 1 very short, transversely linear; somite 2 short, transversely subrectangular; main fused section composed of somites 3 to 6, elongate trapezoidal, swollen at both side of proximal 0.3, with large triangular tooth at near distal border, side margin divided with small V-shaped notch between somites 5/6 discernible; telson tongue-shaped with triangular tip. Female abdomen (Fig. 10D) covered with microscopic, ﬂat granules as on carapace; somite 1 completely concealed beneath carapace; somite 2 short, transversely subrectangular; main fused section ovate, composed of somites 3 to 6, moderately convex ventrally, divided into subergions by 3 transverse and 2 median longitudinal grooves, with small subdistal tubercle; telson tongue-shaped. G1 (Fig. 11K) subcylindrical, weakly constrict- Crabs dredged off the Ogasawara Islands 237 Fig. 11. Ebalia koyo sp. nov., holotype male (cl 2.1 mm), NSMT-Cr S 920. A, B, mandible; C, maxillule; D, maxilla; E, ﬁrst maxilliped; F, second maxilliped; G, H, third maxilliped; I, right P2; J, abdomen; K, left G1 and G2, abdominal. Scales for A–H, K =0.25 mm; I, J = 0.5 mm. ed at distal 0.4, with rounded distal aperture. G2 (Fig. 11K) filiform, slightly longer than G1, inserted into proximal aperture of G1; tip protruding from distal aperture of G1. Color in life. Carapace white with broad, reddish, median band extending posteriorly from the neck of frontal region; pereiopods white, with some reddish spots. Etymology. Named after R/V Koyo of the Tokyo Metropolitan Ogasawara Fisheries Center; used a noun in apposition. Remarks. Ebalia koyo sp. nov. resembles E. humilis Takeda, 1977 and E. stellaris Naruse and Ng, 2006 in the expanded branchial region, the triangularly projecting intestinal region, the bilobed posterior lobes, the formula of abdomen, and the elongate G2. But E. koyo can be distinguished from E. humilis and E. stellaris by that 1) the frontal ridge is high (vs. only marked in E. humilis and E. stellaris); 2) the pterygostomian margin of the carapace has no tubercle (vs. with median tubercle); 3) the chelipedal palm has 238 Hironori Komatsu rounded, subproximal lobe (vs. without lobe); 4) the chelipedal dactylus of full grown male is not strongly curved (vs. strongly arcuate and having semicircular gap between the cutting edges); 5) the tip of G2 is straight (vs. strongly recurved). Distribution. Known from Ogasawara Is. and Amami-oshima Is., occurring at the depths of 100–150 m. Ebalia sakaii Takeda and Miyake, 1972 [Jn: Sakai-ebalia] (Fig. 7C) Materials examined. KY-08-12, 1 ovig. female (cl 2.5 mm; damaged), NSMT-Cr S 923; KY-0921, 1 young male (carapace lost), NSMT-Cr S 924. Distribution. Known only from Japan (Amamioshima Is., Tsushima Strait and Ogasawara Is.), occurring at the depths of 125–150 m (Takeda and Miyake, 1972; Marumura and Kosaka, 2003; this study). This is the ﬁrst record from the Ogasawara Islands. Ebalia tuberculosa (A. Milne-Edwards, 1873) [Jn: Yamato-ebalia] (Fig. 7D) Material examined. SY-09-07, 1 young male (4.7×4.7), NSMT-Cr S 925. Distribution. Widely distributed in Indo-West Pacific; occurring at the depths of 50–550 m (Takeda, 2001). This is the ﬁrst record from the Ogasawara Islands. Heteronucia perlata (Sakai, 1963) [Jn: Maru-tsubu-kobushi] (Fig. 7E) Material examined. KY-09-34, 1 ovig. female (7.8×8.2), NSMT-Cr S 926. Distribution. Philippines, East China Sea, Japan, occurring at the depths of 65–219 m (Chen, 1989; Takeda and Komatsu, 2005). This is the ﬁrst record from the Ogasawara Islands. Heteronucia toyoshioae Komatsu and Takeda, 2005 [New Jn: Toyoshio-tsubu-kobushi] (Fig. 5C) Material examined. KT-09-2-KK1-2 (1), 1 carapace (empty shell; 5.4×5.9), NSMT-Cr S 927. Distribution. Known only from Ryukyu Is. and Izu Is., Japan, occurring at the depths of 145–176 m (Komatsu and Takeda, 2005). This is the ﬁrst record from the Ogasawara Islands. Oreotlos angulatus (Rathbun, 1906) [Jn: Togarikaruishi-kobushi] (Fig. 12A) Materials examined. KY-08-15, 1 ovig. female (10.7×16.5), NSMT-Cr S 928; KT-09-2KK1-2(1), 1 young female (4.4×6.2), NSMT-Cr S 929. Distribution. Known only from Hawaii and Japan, occurring at the depths of 80–172.5 m (Tan and Ng, 1995; Marumura and Kosaka, 2003; this study). This is the ﬁrst record from the Ogasawara Islands. Oreotlos heuretos Tan and Ng, 1995 [Jn: Ogasawara-karuishi-kobushi] (Fig. 12B) Oreophorus (Oreotlos) latus: Takeda, 1977: 119, pl. 1C (not Oreotlos latus (Borradaile, 1903)). Oreotlos heuretos Tan and Ng, 1995: 148, pls. 13C–F, 14A. Materials examined. KY-08-11, 1 male (5.1× 7.7), NSMT-Cr S 930; KY-09-21, 1 male (4.1× 6.0), 1 ovig. female (7.5×12.3), NSMT-Cr S 931; KY-09-21, 1 young male (2.9×4.1), NSMTCr S 932; KY-09-28, 1 male (4.3×6.5), 1 young male (2.7×4.0), NSMT-Cr S 933; KY-09-30, 1 young male (2.1×3.0), NSMT-Cr S 934; KY-1006, 1 young male (2.9×3.9), NSMT-Cr S 935; KY-10-27, 1 young male (2.2×3.1), NSMT-Cr S 936; Ohmura beach, Chichi-jima I., coll. T. Komai, 17 July 2009, 1 male (5.3×7.9), NSMT-Cr S 937. Distribution. Japan, Malay Archipelago, Philippines, occurring at intertidal water deep to 80 m (Tan and Ng, 1995; Marumura and Kosaka, 2003; this study). Oreotlos lagarodes Tan and Ng, 1995 [Jn: Hawaii-karuishi-kobushi] (Fig. 5D) Material examined. KY-09-30, 1 young female (2.9×4.2), NSMT-Cr S 938. Distribution. Hawaii, Laysan Is., Rongelap, Guam, Saipan, Japan, occurring at intertidal water deep to 51.6 m (Tan and Ng, 1995; Marumura and Kosaka, 2003; this study). This is the ﬁrst re- Crabs dredged off the Ogasawara Islands 239 Fig. 12. A, Oreotlos angulatus (Rathbun, 1906), female (10.7×16.5), NSMT-Cr S 928; B, Oreotlos heuretos Tan and Ng, 1995, ovig. female (7.5×12.3), NSMT-Cr S 931; C, Praebebalia dondonae Chen, 1989, female (4.5×4.7), NSMT-Cr S 939; D, Praebebalia taeniata Takeda, 1977, male (3.3×3.3), NSMT-Cr S 940; E, Hoplophrys oatesi Henderson, 1893, male (6.1×4.8), NSMT-Cr S 955; F, Huenia paciica Miers, 1879, young males (4.2×3.2), NSMT-Cr S 956. cord from the Ogasawara Islands. Praebebalia dondonae Chen, 1989 [New Jn: Philippines-ebalia-modoki] (Fig. 12C) Material examined. SY-09-07, 1 female with rhizocepharan parasite in abdominal cavity (4.5 ×4.7), NSMT-Cr S 939. Remarks. The present specimen agrees well 240 Hironori Komatsu with the original description and illustration by Chen (1989). Chen (1989) erroneously described the female abdomen as being composed of the ﬁrst somite and the fused second to sixth somites, however, the ﬁrst somite is completely concealed beneath the carapace in fact. Therefore the female abdomen is actually composed of the second somite and the fused third to sixth somites. Male specimen is hitherto not known. Distribution. Previously known only from the type locality (NE Mindoro I., Philippines, 181– 170m) (Chen, 1989). This is the ﬁrst record from Japan and extends the bathymetrical range deep to 499 m. Praebebalia taeniata Takeda, 1977 [Jn: Ogasawara-ebalia-modoki] (Fig. 12D) Praebebalia taeniata Takeda, 1977: 118, figs. 2C–H, 4A; Takeda and Kurata, 1984: 198, ﬁg. 10; Marumura and Kosaka, 2003: 26. Not Praebebalia taeniata: Takeda and Kurata, 1977a: 141, fig. 1 (=Ebalia humilis Takeda, 1977). Materials examined. KY-08-03, 1 male (3.3× 3.3), NSMT-Cr S 940; KY-08-19, 2 males (3.2× 3.3, 3.5×3.6), 1 female (3.3×3.6), NSMT-Cr S 941; KY-09-21, 1 young female (2.6×2.7), NSMT-Cr S 942; KY-10-02, 1 ovig. female (3.2× 3.2), NSMT-Cr S 943; KY-10-19, 1 ovig. female (3.1×3.2), NSMT-Cr S 944; KT-09-2-TW2-4, 1 male (3.1×3.0), NSMT-Cr S 945; SY-09-01, 1 male (3.0×2.9), NSMT-Cr S 946; SY-09-02, 1 ovig. female (3.3×3.3), NSMT-Cr S 947; SY-0906, 1 male (3.7×3.8), NSMT-Cr S 948; SY-0908, 1 ovig. female (3.5×3.6), 1 female (3.2× 3.2), NSMT-Cr S 949; SY-09-18, 1 male (3.6× 3.6), NSMT-Cr S 950; SY-09-20, 3 males (3.2× 3.2–3.3×3.2), 2 ovig. females (3.4×3.5, 3.5× 3.6), NSMT-Cr S 951. Distribution. Known only from the Ogasawara Islands, occurring at the depths of 90–301 m (Takeda, 1977; Marumura and Koasaka, 2003; this study). Toru granuloides (Sakai, 1961) [Jn: Ibonashi-kobushi] (Figs. 7F, 8D, E) Materials examined. KY-09-31, 1 carapace (empty shell; 9.4×9.4), NSMT-Cr S 952; SY-0907, 2 young males (4.7×4.6, 6.5×6.6), NSMTCr S 953; 1 male (12.7×12.0), NSMT-Cr S 954, off Haha-jima I., 26°43′50″N 142°18′19″E – 26°43′52″N 142°18′08″E, 331–325 m, TR/V Shin’yo Maru, st. 1, coll. M. Osawa, 18 October 1998. Remarks. The present specimens agree well with the original description by Sakai (1961), but differ in the shape of the ﬁrst gonopod illustrated by Galil (2003) based on a male from the Loyalty Islands. The apical process of the ﬁrst gonopod is short in the present specimen (Fig. 8D, E), whereas it is long and sickle-shaped in Galil’s figure. Our specimen may be still young and the gonopod may be incompletely formed. It is necessary to check the male gonopods of the type specimens from Japan, but it could not be located in the Kanagawa Prefectural Museum of Natural History and the Kitakyushu Museum of Natural History and Human History. Distribution. Fiji, Wallis I., Vanuatu, Loyalty Is., New Caledonia, Japan; occurring at the depths of 50–550 m (Galil, 2003). This is the ﬁrst record from the Ogasawara Islands. Superfamily Majoidea Samouelle, 1819 Family Epialtidae MacLeay, 1838 Hoplophrys oatesi Henderson, 1893 [Jn: Iso-konpeito-gani] (Fig. 12E) Hoplophrys ogilbryi: Ooishi, 1970: 91, pl. 13(5). Materials examined. KY-09-14, 1 male (6.1× 4.8), 1 female (10.2×7.9), NSMT-Cr S 955. Distribution. Widely distributed in the IndoWest Pacific Ocean, occurring at the depths of 2–93 m (Grifﬁn and Tranter, 1986; this study). Huenia paciica Miers, 1879 [New Jn: Taiheiyokonoha-gani] (Fig. 12F) Huenia proteus: Takeda and Kurata, 1976a: 121, ﬁg. 2c. Materials examined. KY-08-18, 4 young males (3.0×2.3–4.2×3.2), NSMT-Cr S 956. Remarks. The present specimens agree well Crabs dredged off the Ogasawara Islands with the subsequent description and illustration of juvenile specimen by Takeda and Marumura (2010). Takeda and Kurata (1976a) recorded anterior half of carapace and ambulatory legs of Huenia proteus de Haan, 1839, from the gut contents of Epinephelus fasciatus from the Ogasawara Islands, but it possibly belongs to H. paciica. Distribution. Indonesia, Philippines, Japan, Australia, Fiji, Salas y Gómez I., occurring at the depths of 2–49 m (Takeda and Marumura, 2010; this study). This is the ﬁrst record from the Ogasawara Islands. Hyastenus ambonensis Grifﬁn and Tranter, 1986 [New Jn: Ambon-tsuno-gani] (Figs. 8F–I, 13A) Materials examined. KY-08-15, 3 males (4.8× 3.2–7.1×5.4), 1 young male, 1 ovig. female (6.4 ×4.5), NSMT-Cr S 957; KY-08-20, 1 male (6.7 ×4.8), NSMT-Cr S 958; KY-10-31, 1 ovig. female (8.0×5.8), NSMT-Cr S 959. Remarks. Hyastenus ambonensis Griffin and Tranter, 1986, is previously known only by the original description based on two female specimens from Ambon, Indonesia. Thus, this is the second record of the species and the ﬁrst record of male. The first male gonopod (Fig. 8F–H) is ﬂattened and broadened distally and strongly upturned on distomesial margin. The present specimens agree well with the original description by Grifﬁn and Tranter (1986), but differ in that 1) the dorsal surface of the merus of the ﬁrst ambulatory leg has three tubercle (vs. smooth in the original description); 2) the dactylus of the fourth ambulatory leg has 6 or 7 teeth (vs. 8 in the original description). Distribution. Indonesia and Japan, occurring at the depths of 52–90 m (Grifﬁn and Tranter, 1986; this study). This is the ﬁrst record from the Ogasawara Islands. Hyastenus kyusyuensis (Yokoya, 1933) [Jn: Kyushu-tsuno-gani] (Figs. 13B, 14) Hyastenus kyushuensis: Marumura and Kosaka, 2003: 33. Materials examined. KY-08-15, 1 young male 241 (5.0×3.3), NSMT-Cr S 960; KY-08-21, 1 juv., NSMT-Cr S 961; KY-08-26, 1 ovig. female (9.0 ×5.9), NSMT-Cr S 962; KY-09-07, 1 ovig. female (9.6×6.2), 1 young female (6.3×4.5), NSMT-Cr S 963; KY-09-08, 1 young female (7.7× 5.1), NSMT-Cr S 964; KY-10-11, 1 male (5.2× 3.4), NSMT-Cr S 965; KY-10-31, 1 male (5.9× 3.9), 1 ovig. female (6.5×4.6), NSMT-Cr S 966; KT-09-2-TW2-4, 1 female (7.8×5.9), NSMT-Cr S 967. Additional materials examined. 1 young male (5.6×3.9), NSMT-Cr S 968, Kurose Bank, Izu Islands, 33°22.01′N 139°41.60′E – 33°21.97′N 139°41.93′E, 116–123 m, rocky dredge, R/V Takunan, coll. H. Komatsu, 16 July 2008; 1 male (11.7×8.5), NSMT-Cr 21462, north of Mage-jima I., Ohsumi Is., western Japan, 30°49.95′N 130°51.13′E – 30°50.06′N 130°51.46′E, 119–111 m, TR/V Toyoshio Maru 2009-03 cruise, stn 2, beam trawl, coll. H. Komatsu, 19 May 2009; 1 male (10.3×7.5), NSMT-Cr 9448, Minabe, Kii Peninsula, Japan, coll. M. Marumura, 21 February 1987; 1 male (6.7×5.3), NSMT-Cr 15060, East China Sea, 28°28.98′ N 125°30.41′ E – 28°26.92′N 125°31.21′E, 107–112 m, otter trawl, R/V Yoko Maru, stn 21, coll. H. Saito and G. Shinohara, 21 October 2001. Comparative material examined. Hyastenus convexus (Miers, 1884): 1 ovig. female (10.9× 7.6), NSMT-Cr 9677, off Doren, Ohima Passage, Amami Islands, Ryukyu Islands, Japan, coll. M. Takeda, 8 August 1988. Remarks. Hyastenus kyusyuensis (Yokoya, 1933) was brieﬂy described and illustrated on the basis of a single male from west of Tanega-shima Island, Osumi Islands, western Japan. The type specimen could not be located in the University Museum, the University of Tokyo and the Kitakyushu Museum of Natural History and Human History. Subsequently H. kyusyuensis was described and illustrated by Sakai (1935, 1936, 1938, 1965, 1976), but his description and illustration are different from the original description in the presence of a mesogastric tubercle except in Sakai (1935). Considered with presence of 1 mesogastric, 2 protogastric and 3 epibranchial tu- 242 Hironori Komatsu Fig. 13. A, Hyastenus ambonensis Grifﬁn and Tranter, 1986, males (7.1×5.4), NSMT-Cr S 957; B, Hyastenus kyusyuensis (Yokoya, 1933), male (5.9×3.9), NSMT-Cr S 966; C, Lahaina ovata Dana, 1951, male (8.9×6.6), NSMT-Cr S 981; D, Menaethius monoceros (Latreille, 1825), male (5.9×4.7), NSMT-Cr S 987; E, Naxioides taurus (Pocock, 1890), young female (16.7×11.4), NSMT-Cr S 991; F, Phalangipus hystrix (Miers, 1886), young male (10.2×6.7), NSMT-Cr S 994. bercles, his species actually belongs to H. convexus (Miers, 1884) except in Sakai (1935) (cf. Griffin and Tranter, 1986). Sakai’s description differs from the subsequent description of H. con- vexus by Griffin and Tranter (1986) in having a single tubercle on the mesogastric region (vs. 2 tubercles in Griffin and Tranter (1986)), but the tubercle might be too obscure to recognize it. Crabs dredged off the Ogasawara Islands 243 Fig. 14. Hyastenus kyusyuensis (Yokoya, 1933). A, B, male (5.9×3.9), NSMT-Cr S 966; C–E, male (11.7×8.5), NSMT-Cr 21462. A, anterior part of carapace, ventral; B, right third maxilliped, ventral; C, G1, abdominal; D, tip of same; E, G2, abdominal. Scales for A = 1 mm; B, C, E = 0.5 mm; D = 0.25 mm. Illustrations of the antero-ventral face of the carapace (Fig. 14A), the third maxilliped (Fig. 14B) and the male gonopods (Fig. 14C–E) are provided for the ﬁrst time. Distribution. Known only from Japan, occur- ring at the depths of 80–151.5 m (Yokoya, 1933; Sakai, 1935; Miyake, 1937; Takeda and Miyake, 1969b; Marumura and Kosaka, 2003; this study). Lahaina ovata Dana, 1951 [Jn: Maru-tsuno-gani- 244 Hironori Komatsu modoki] (Fig. 13C) Hyastenus tenuicornis: Takeda, 1977, 122, pl. 13C, D; Takeda and Kurata, 1977b: 142, ﬁgs. 2, 3; Marumura and Kosaka, 2003: 33. Materials examined. KY-08-06, 2 young males, 1 ovig. female (6.5×4.8), 1 young female, NSMT-Cr S 970; KY-08-11, 1 ovig. (6.4×4.7), NSMT-Cr S 971; KY-08-15, 1 young male, 1 young female, 3 juvs., NSMT-Cr S 972; KY-08-20, 1 male (4.7×4.2), 1 young male, 1 young female, NSMT-Cr S 973; KY-08-21, 1 ovig. female (5.6 ×4.0), NSMT-Cr S 974; KY-08-26, 1 ovig. female (6.1×4.5), NSMT-Cr S 975; KY-09-08, 1 male (6.4×5.1), NSMT-Cr S 976; KY-09-14, 1 male (4.6×3.0), 1 juv., NSMT-Cr S 977; KY-0928, 2 males (6.8×5.0, 8.4×6.4), 3 ovig. females (5.9×4.3–7.0×5.4), NSMT-Cr S 978; KY-09-29, 3 males (6.0×4.5–7.2×5.3), 2 ovig. female (6.2 ×4.8, 6.9×5.3), 1 juv., NSMT-Cr S 979; KY-0930, 5 ovig. female (5.9×4.3–7.0×5.2), 2 juvs., NSMT-Cr S 980; KY-10-06, 1 male (8.9×6.6), NSMT-Cr S 981; KY-10-07, 1 male (7.7×5.9), NSMT-Cr S 982; KY-10-27, 1 ovig. female (6.9 ×5.4), NSMT-Cr S 983; KY-10-31, 1 ovig. female (5.8×4.2), NSMT-Cr S 984; SY-09-21, 1 young male, NSMT-Cr S 985. Remarks. This species is one of the most common majoid crab in the Ogasawara Islands. Hyastenus tenuicornis Pocock, 1890 was reduced to a junior subjective synonym of this species by Grifﬁn (1974). Distribution. Widely distributed in the IndoWest Pacific Oceans from Seychelles and Red Sea east to Hawaii Islands, occurring at the depths of 18–161 m (K. Sakai, 2004; this study). The present record extends the bathymetrical range deep to 161 m. Menaethius monoceros (Latreille, 1825) [Jn: Ikkaku-gani] (Fig. 13D) Menaethius monoceros: Takeda and Kurata, 1976a: 123, figs. 2f, 4a; 1977a: 108; 1977b: 142; Takeda, 1977: 134. Materials examined. KY-09-30, 1 young male (3.8×3.0), 1 young female (3.9×3.1), NSMT-Cr S 986; Tsuri-hama Beach, Chichi-jima I., coll. H. Komatsu, 11 Jul. 2009, 1 male (5.9×4.7), NSMT-Cr S 987; Futami Port, Chichi-jima I., coll. H. Komatsu, 8 Jul. 2009, 1 female (5.6×4.5), 1 young female, NSMT-Cr S 988; Oki Port, Hahajima I., coll. H. Komatsu, 13 Jul. 2009, 1 male (4.2×3.1), 3 young males, 1 ovig. female (6.3× 4.8), NSMT-Cr S 989. Distribution. Widely distributed in the IndoWest Pacific Oceans from Red Sea and eastern Africa to French Polynesia, occurring at intertidal area to 57 m (K. Sakai, 2004). Naxioides taurus (Pocock, 1890) [Jn: Ko-edatsuno-gani] (Figs. 8M, N, 13E) Naxioides mammillata: Takeda and Kurata, 1976a: 122, ﬁg. 5a [not Naxioides mammillata (Ortmann, 1893)]. Materials examined. KY-08-06, 1 young male (4.9×3.1), NSMT-Cr S 990; KY-08-15, 1 young female (16.7×11.4), NSMT-Cr S 991; KY-09-30, 1 female (23.2×16.1), 1 young female (5.1× 3.3), NSMT-Cr S 992. Additional materials examined. 4 males (15.6 ×11.7–17.6×12.8), 2 ovig. females (14.9×10.6, 17.8×12.3), 2 young females (11.6×7.7, 11.5× 7.9), NSMT-Cr 15394, Balicasag I., Bohol, Philippines, coll. local ﬁshermen, February 2003. Remarks. Takeda and Kurata (1976a) recorded an anterior part of carapace of Naxioides mammillata (Ortmann, 1893) from the gut contents of lethrinid ﬁsh, Gymnocranius japonicus, from the Ogasawara Islands, but their species actually refers to this species in due to the divergent rostrum. Illustration of the male gonopods (Fig. 8M, N) is provided. Distribution. Widely distributed in the IndoWest Pacific Oceans from Red Sea and eastern Africa to Australia, Indonesia, Philippines and Japan, occurring at the depths of 16–88 m (Grifﬁn and Tranter, 1986; this study). This is the ﬁrst record from the Ogasawara Islands. Phalangipus hystrix (Miers, 1886) [Jn: Ashinaga-tusno-gani] (Fig. 13F) Material examined. SY-09-18, 1 young male (10.2×6.7), NSMT-Cr S 994. Crabs dredged off the Ogasawara Islands Distribution. Widely distributed in the IndoWest Paciﬁc Oceans from Red Sea to Indonesia, Australia and Japan, occurring at the depths of 36–180 m (K. Sakai, 2004). This is the first record from the Ogasawara Islands. Pugettia ogasawaraensis sp. nov. [New Jn: Ogasawara-mo-gani] (Figs. 15A, 16) Materials examined. Holotype: male (7.7× 5.6), NSMT-Cr S 995, SY-09-04, east off Mukojima I., Ogasawara Is., 27°44.99′N 142°10.52′E – 27°44.79′N 142°10.40′E, 159–152m, dredge, TR/V Shin’yo Maru, coll. H. Komatsu, 16 November 2009. Paratypes: 1 young male (5.3×3.9), NSMT-Cr S 996, KY-08-15, south of Chichi-jima I., Ogas a w a r a I s . , 2 7 °0 4 . 5 5 ′ N 1 4 2 °0 9 . 1 6 ′ E – 27°04.73′N 142°09.31′E, 83–81 m, dredge, R/V Koyo, coll. T. Fujita, H. Namikawa and M. Okanishi, 28 October 2008; 1 young male (4.7×3.5), 1 young female (4.8×3.4), NSMT-Cr S 997, SY09-02, east off Muko-jima I., Ogasawara Is., 2 7 °4 4 . 5 5 ′ N 1 4 2 °0 9 . 6 9 ′ E – 2 7 °4 4 . 4 7 ′ N 142°09.76′E, 122–123m, dredge, TR/V Shin’yo Maru, coll. H. Komatsu, 16 November 2009; 1 young female (6.1×4.2), NSMT-Cr S 998, same data as holotype; 1 male (6.4×4.3), 1 young female (6.4×4.3), NSMT-Cr S 999, SY-09-09, east off Muko-jima I., Ogasawara Is., 27°41.13′N 142°11.11′E – 27°41.00′N 142°10.88′E, 124– 112m, dredge, TR/V Shin’yo Maru, coll. H. Komatsu, 16 November 2009. Description. Carapace (Fig. 15A) pyriform, 1.3–1.5 times longer than broad; surface smooth, two lateral spines, curled setae on rostral spines, protogastric region and lateral face of branchial region. Rostral spines short, divergent, fused for basal 0.25, subconical, tapering to sharp apex, length 0.35 postrostral carapace length. Supraorbital eave armed with acute, triangular preorbital lobe directed rather obliquely forward and weakly upward; antorbital lobe small, obtusely rounded, continuous from preorbital lobe; postorbital and hepatic lobes completely fused to form winglike plate, separated from eave by U-shaped hiatus, 245 lateral margin weakly divergent posteriorly, slightly concave; postorbital lobe acute triangular in dorsal and lateral views, directed forward and horizontal; hepatic lobe expanded laterally, obtusely rounded. Hepatic region not elaveted, unarmed. Epibranchial region roundly convex with faint tubercle on center; epibranchial spine prominent, conical, directed laterally and slightly upward. Gastric regions prominently elevated, with one median mesogastric and pair of protogastric faint tubercles arranged in broad triangle, metagastric region elevated, highest. Cardiac region prominently elevated, slightly lower than metagastric elevation, both sides of cardiac elevation divided into crescent subregions by longitudinal grooves. Intestinal region moderately projecting posteriorly, rounded. Branchial and intestinal margins rimmed with round granules, double-margined in posterior part, posterior margin conﬂuent with branchial margin below epibranchial spine. Basal antennal segment (Fig. 16A) broad, smooth, anterolateral angle weakly produced forward into rounded tooth, lateral margin almost straight, medially convex dorsally; second and third segment ﬂatted, plate-like. Conical tubercle lateral to green gland. Pterygostomian region smooth, margin with 3 rounded tubercles. Third maxilliped (Fig. 16B) covered with flat granules; ischium with shallow median groove; anterolateral angle of merus produced, rounded, upturned; exopod with acute granules along lateral margin. Anterolateral angle of mouthﬁeld produced, rounded. Cheliped (P1; Fig. 16C, D) of male moderate, as long as carapace; merus smooth, subrectangular in cross-section, each margin sharp, dorsal margin highly carinate, distally ending in large triangular tooth, outer margin with some granules, outer-ventral margin entirely granulated, innerventral margin smooth, distal margin with 1 large median and 2 lateral triangular tubercles dorsally; carpus with 2 granulated ridges on dorsal face, ridge divergent distally, outer ridge directed ventrally in distal half, parallel with outer-distal margin, outer-distal margin ridged, granulate; palm 246 Hironori Komatsu Fig. 15. A, Pugettia ogasawaraensis sp. nov., holotype male (7.7×5.6), NSMT-Cr S 995; B, Xenocarcinus conicus (A. Milne-Edwards, 1865), male (7.7×4.5), NSMT-Cr S 1000; C, Xenocarcinus tuberculatus White, 1847, male (8.8× 4.5), NSMT-Cr S 1001; D, Achaeus boninensis Miyake and Takeda, 1969, ovig. female (7.0×5.1), NSMT-Cr S 1002; E, Achaeus brevifalcatus Rathbun, 1911, male (4.3×2.8), NSMT-Cr S 1006; F, Achaeus curvirostris (A. Milne-Edwards, 1873), male (3.8×3.1), NSMT-Cr S 1024. 1.3 times longer than height, dorsal and ventral margins sharp; ﬁngers 0.85 times as long as palm, without gape between cutting edges, furnished with small triangular teeth in distal 0.7 of both edges. Ambulatory legs (P2–P5) slender, sparsely with club setae, similar in shape, P2 longest, 1.4 times as long as carapace, P5 0.9 times as long as Crabs dredged off the Ogasawara Islands 247 Fig. 16. Pugettia ogasawaraensis sp. nov., holotype male (7.7×5.6), NSMT-Cr S 995. A, anterior part of carapace, ventral; B, right third maxilliped, ventral; C, D, left cheliped, lateral; E, G1, abdominal; F, tip of same; G, G2, abdominal. Scales for A, C, D = 1mm; B, E, G = 0.5 mm; F, 0.25 mm. carapace; merus subcylindrical, dorsal margin sharp, not carinate, with soft setae, distal margin with 3 triangular tubercles dorsally as in cheliped; carpus with 2 dorsal ridges; dactylus with 6 very 248 Hironori Komatsu small tubercles ventrally. Male thoracic sternum covered with granules, episternites divided; sternites 1–4 fused together, with very short suture between sternites 3/4 laterally; sutures between sternites 4/5 and 5/6 interrupted medially, sutures between 6/7 and 7/8 entire, median longitudinal suture extending from suture between sternites 6/7 posteriorly. Male abdomen covered with coarse granules; all segments free; telson triangular with rounded tip. First male gonopod (G1; Fig. 16E, F) slender, straight, compressed, dilated at tip; tip divided into 3 lobes, slender mesial lobe, broad lateral lobe and small triangular median lobe directed abdominally, median part behind median lobe with very short spinules. Second male gonopod (G2; Fig. 16G) short, 0.3 times as long as G1; distal segment rounded. Etymology. The speciﬁc name was named after the type locality, the Ogasawara Islands. Remarks. Pugettia ogasawaraensis sp. nov. resembles P. incisa (de Haan, 1850) in the postorbital and hepatic lobes being completely fused to form a winglike plate, which is unique for these two species in the genus Pugettia, but can be distinguished from P. incisa by that 1) both sides of cardiac prominence are divided (undivided in P. incisa); 2) epibranchial region is roundly raised (vs. not raised); 3) the chelipedal merus is rectangular in cross-section (vs. triangular); 4) mesial process of the tip of the first male gonopod is smaller than lalteral process (vs. larger than lateral process) (cf. Sakai, 1938; Gordon, 1931; Grifﬁn and Tranter, 1986). Pugettia ogasawaraensis is thought to be derived from P. incisa, which is an East Asia endemic element, and its ancestor was possibly drifted from the coast of mainland of Japan by countercurrent of Kuroshio Current (Fig. 1). Such an East Asia endemic species unique for the Ogasawara Islands is known for two decapod crustacean species, viz. Pagurus insulae and Petrolisthes mesodactylon (cf. Asakura, 1991; Asakura et al., 1994a, b), and five molluscan species, viz. Cellana mazatlandica, Nipponacmea boninensis, Monodonta aff. australis, M. perplexa boninsensis and Turbo sp. (cf. Fukuda, 1993, 1994, 1995). Distribution. Known only from the Ogasawara Islands, occurring at the depths of 81–159 m. Xenocarcinus conicus (A. Milne-Edwards, 1865) [Jn: Nakazawa-isobana-gani] (Fig. 15B) Material examined. KY-09-28, 1 male (7.7× 4.5), NSMT-Cr S 1000. Distribution. Widely distributed in the IndoWest Paciﬁc Oceans from Red Sea and Réunion to Indonesia and Japan, occurring at the depths of 13–80 m (K. Sakai, 2004). This is the ﬁrst record from the Ogasawara Islands. Xenocarcinus tuberculatus White, 1847 [Jn: Iboisobana-gani] (Fig. 15C) Xenocarcinus tuberculatus: Takeda, 1973: 31. Material examined. KY-10-31, 1 male (8.8× 4.5), NSMT-Cr S 1001. Distribution. Indian Ocean: Red Sea and east coast of Africa; West Pacific Ocean: Singapore, Sulu Archipelago, Hong Kong, Japan and Australia, occurring at the depths of 10–150 m (K. Sakai, 2004). Family Inachidae MacLeay, 1838 Achaeus boninensis Miyake and Takeda, 1969 [Jn: Ogasawara-achaeus] (Fig. 15D) Achaeus boninensis Miyake and Takeda, 1969: 27, ﬁg. 2; Takeda and Kurata, 1976a: 121, ﬁg. 2a, b. Material examined. KY-09-07, 1 ovig. female (7.0×5.1), NSMT-Cr S 1002. Distribution. Known only from the Ogasawara Islands, occurring at the depths of 136–200 m (Miyake and Takeda, 1969; this study). Achaeus cf. brevidactylus Sakai, 1938 Material examined. KY-10-31, 1 young male (2.2×1.8), NSMT-Cr S 1003. Remarks. The present specimen is too small to identify into the species. But it resembles Achaeus brevidactylus Sakai, 1938, in that 1) the carapace is smooth except very low gastric and cardiac tu- Crabs dredged off the Ogasawara Islands bercles; 2) the dactyli of P4 and P5 are not strongly falcated. However it differs in that the telson of young male is broadly triangular (vs. sharply triangular in the original description). An adult specimen is needed for exact identiﬁcation. Achaeus brevifalcatus Rathbun, 1911 [Jn: Seychelles-achaeus] (Fig. 15E) Achaeus brevifalcatus: Miyake and Takeda, 1969: 23, ﬁg. 1. Materials examined. KY-08-06, 1 male (5.0× 3.1), 2 ovig. females (4.5×3.2, 5.1×3.8), NSMT-Cr S 1004; KY-08-21, 2 males (4.5×3.0, 4.5 ×3.0), 1 ovig. female (4.7×3.5), 1 female (4.3 ×3.3), NSMT-Cr S 1005; KY-09-07, 1 male (4.3 ×2.8), NSMT-Cr S 1006; KY-09-29, 1 young female, NSMT-Cr S 1007; KY-10-19, 1 female (5.4 ×4.0), NSMT-Cr S 1008; KT-09-2-TW1-1, 1 male (4.3×2.9), 1 female (5.3×4.1), NSMT-Cr S 1009; SY-09-04, 1 male (4.2×2.7), NSMT-Cr S 1010; SY-09-11, 1 male (4.8×2.5), NSMT-Cr S 1011. Distribution. Indian Ocean: Red Sea, Seychelles and Mauritius; West Paciﬁc Ocean: Indonesia, Japan and Hawaii; occurring at the depths of 60–234 m (K. Sakai, 2004; this study). This study extends the bathymetrical range shallow to 60 m. Achaeus curvirostris (A. Milne-Edwards, 1873) [Jn: Naga-kubi-achaeus] (Fig. 15F) Achaeus issifrons: Takeda, 1977: 122. Materials examined. KY-08-06, 2 males (4.0× 249 2.8, 4.2×2.9), 1 ovig. female (4.6×3.5), NSMTCr S 1012; KY-08-15, 6 males (3.2×2.2–5.4× 3.6), 2 ovig. female (3.3×2.6, 3.6×2.9), 1 female (3.7×2.8), 1 young female, NSMT-Cr S 1013; KY-08-18, 1 young female, NSMT-Cr S 1014; KY-08-21, 2 males (3.6×2.3, 3.6×2.6), 1 ovig. female (3.6×2.7), 2 juvs., NSMT-Cr S 1015; KY-08-25, 1 female (3.5×2.8), NSMT-Cr S 1016; KY-08-26, 2 males (4.1×2.9, 6.3×4.4), 1 ovig. female (4.5×3.3), NSMT-Cr S 1017; KY09-14, 1 ovig. female (4.1×3.2), 1 young female, NSMT-Cr S 1018; KY-09-28, 1 male (7.2×4.9), 1 ovig. female (5.3×3.7), 1 female (5.1×4.0), 1 young female, NSMT-Cr S 1019; KY-09-29, 1 male (4.8×3.3), 1 ovig. female (5.0×4.0), 1 juv., NSMT-Cr S 1020; KY-09-30, 2 males (5.0×3.5, 5.8×4.3), 2 ovig. females (4.5×3.8, 4.9×3.9), 1 female (4.7×4.0), NSMT-Cr S 1021; KY-09-34, 3 males (4.8×3.4–5.5×3.8), NSMT-Cr S 1022; KY-10-07, 1 female (5.4×4.2), NSMT-Cr S 1023; KY-10-24, 1 male (3.8×3.1), 1 female (4.6 ×3.8), NSMT-Cr S 1024; KY-10-27, 1 ovig. female (4.5×3.8), NSMT-Cr S 1025; KT-09-2TW2-3, 1 male (4.0×2.7), NSMT-Cr S 1026. Remarks. This species is the most abundant species of the genus Achaeus at the sea around the Ogasawara Islands. A key to the species of the genus Achaeus known from the Ogasawara Islands is provided. Distribution. Widely distributed in the IndoWest Paciﬁc Oceans from east coast of Africa to Indonesia, Japan and Australia, occurring at the depths of 36–166 m (K. Sakai, 2004). Key to the species of the genus Achaeus known from the Ogasawara Islands 1. Supraorbital eave with a spine ………………………………………………………… A. curvirostris Supraorbital eave without spine ……………………………………………………………………… 2 2. Branchial region with three small spines ……………………………………………… A. boninensis Branchial region without spines ……………………………………………………… A. brevifalcatus Chalaroachaeus curvipes de Man, 1902 [Jn: Kubi-nashi-achaeus] (Fig. 5E) Material examined. KY-08-26, 1 ovig. female (3.5×3.6), NSMT-Cr S 1027. Remarks. The present specimen agrees well with the subsequent description and illustration by Grifﬁn and Tranter (1986). This species is well known to attach fragments of bryozoan to the body and the pereiopods by itself in Japan (Minemizu, 2000; Kato and Okuno, 2001; Kawamoto and Okuno, 2003). Distribution. Known from Indonesia and Japan, 250 Hironori Komatsu occurring at the depths of 2–87 m (Takeda, 1989; this study). This is the ﬁrst record from the Ogasawara Islands and extends the bathymetrical range deep to 87 m. Cyrtomaia lamellata Rathbun, 1906 [Jn: Mitsutoge-owston-gani] (Fig. 17A) Cyrtomaia platypes: Takeda and Kurata, 1976b: 24, pl. 2 ﬁg. 1. Materials examined. KY-08-15, 1 ovig. female (9.6×10.3), NSMT-Cr S 1028; KY-08-25, 1 male (7.9×7.9), NSMT-Cr S 1029; KY-09-34, 1 young male (5.9×5.3), NSMT-Cr S 1030; SY-09-09, 1 ovig. female (9.8×10.0), NSMT-Cr S 1031. Distribution. West Paciﬁc: Japan, Timor, New Caledonia, New Zealand, Hawaii, occurring at the depths of 81–290 m (Griffin and Tranter, 1986; Takeda and Komatsu, 2005; this study). This study extends the bathymetrical range shallow to 81 m. Grypachaeus hyalinus (Alcock and Anderson, 1894) [Jn: Hasami-ashi-achaeus] (Fig. 17B) Materials examined. KY-08-21, 1 male (3.0× 1.9), NSMT-Cr S 1032; KY-08-25, 1 male (4.0× 2.3), NSMT-Cr S 1033; KY-09-21, 1 male (4.8× 2.7), NSMT-Cr S 1034; KY-10-11, 1 young female (5.2×2.8), NSMT-Cr S 1035; KY-10-31, 1 male (6.2×3.7), 1 young female (6.0×3.7), NSMT-Cr S 1036. Remarks. The present specimens agree well with the subsequent illustration of the male gonopod by Grifﬁn (1974). Distribution. East coast of Africa, Sri Lanka, Timor, Philippines, South China Sea, Japan, occurring at the depths of 50–208 m (Takeda and Komatsu, 2005). This is the ﬁrst record from the Ogasawara Islands. Paratymolus bituberculatus Haswell, 1880 [Jn: Queensland-mametsubu-gani] (Fig. 17C) Paratymolus bituberculatus: Takeda, 1977: 121, pl. 12B. Materials examined. KY-08-15, 1 male (2.7× 2.6), NSMT-Cr S 1037; KY-09-29, 1 ovig. female (3.4×3.5), NSMT-Cr S 1038. Remarks. The present male specimen agrees well with the illustration by Loh and Ng (1999: Fig. 2E–H) in the shape of the male gonopod. Although Loh and Ng (1999) described that the preorbital angle of the supraorbital eave and the anterior tubercle of the anterolateral margin of the carapace are inconspicuous in male, those of the present male specimen are well developed as in female illustrated by Loh and Ng (1999: Fig. 3). Loh and Ng (1999) also treated height of the gastric tubercle as a diagnostic character between P. tuberculata and P. pubescens Miers, 1879, but it seems to be variable intraspeciﬁcally. Loh and Ng (1999) considered the record of this species by Takeda (1977) from the Ogasawara Islands as P. pubescens Miers, 1879. However, the female specimens of Takeda (1977) and the present specimens agree well with the illustration by Loh and Ng (1999: Fig. 3) and therefore they should belong to P. bituberculatus. Distribution. Western Australia, Indonesia and Japan, occurring at intertidal zone deep to 134 m (Loh and Ng, 1999; this study). Pleistacantha simplex Rathbun, 1932 [Jn: Himehari-senbon] (Fig. 17D) Material examined. KY-09-31, 1 female (4.8× 3.8), NSMT-Cr S 1039. Remarks. The present specimen is much smaller than the holotype male (cl 10.5 mm), but the spinulation on the rostral spines, the supraorbital eave, and the basal antennal article agrees well with the subsequent description and illustration by Sakai (1935). Distribution. Indonesia, Philippines and Japan, occurring at the depths of 50–540 m (Sakai, 1976; Grifﬁn and Tranter, 1986). This is the ﬁrst record from the Ogasawara Islands. Prosphorachaeus suluensis Rathbun, 1916 [Jn: Ko-achaeus-modoki] (Fig. 17F) Materials examined. KY-08-15, 1 male (2.6× 2.2), 1 female (2.9×2.6), NSMT-Cr S 1040; KY08-21, 1 male (2.8×2.5), NSMT-Cr S 1041. Remarks. The present specimens agree well with the original description by Rathbun (1916), Crabs dredged off the Ogasawara Islands 251 Fig. 17. A, Cyrtomaia lamellata Rathbun, 1906, ovig. female (9.8×10.0), NSMT-Cr S 1031; B, Grypachaeus hyalinus (Alcock and Anderson, 1894), male (6.2×3.7), NSMT-Cr S 1036; C, Paratymolus bituberculatus Haswell, 1880, male (2.7×2.6), NSMT-Cr S 1037; D, Pleistacantha simplex Rathbun, 1932, female (4.8×3.8), NSMT-Cr S 1039; E, Sunipea indicus (Alcock, 1895), male (4.0×2.8), NSMT-Cr S 1042; F, Prosphorachaeus suluensis Rathbun, 1916, female (2.9×2.6), NSMT-Cr S 1040. but differ in that 1) there are two small spinules behind the supraorbital margin, the second of which represents the postocular spine in the original description (vs. without spinules in the present specimens); 2) there is a small tubercle on posterior slope of the cardiac spine (vs. without tubercle in the present specimens); 3) there is a small tubercle on the epibranchial region and on each side of the posterior margin in the present specimen (vs. absent in the original description). The shape of the ﬁrst male gonopod agrees well with the illustration by Takeda and Miyake (1969b). Distribution. Sulu Archipelago and Japan, oc- 252 Hironori Komatsu curring at the depths of 5.5–105 m (Takeda and Miyake, 1969b; Grifﬁn and Tranter, 1986). This is the ﬁrst record from the Ogasawara Islands. Sunipea indicus (Alcock, 1895) [Jn: Andamanachaeus] (Fig. 17E) Aepinus indicus: Takeda, 1977: 122, ﬁg. 4B. Materials examined. KY-08-15, 7 males (3.6× 2.5–4.0×2.8), 8 ovig. females (5.0×3.4–5.4× 3.8), NSMT-Cr S 1042; KY-08-21, 2 males (3.6 ×2.5, 3.8×2.5), 1 ovig. female (5.4×3.6), NSMT-Cr S 1043; KY-08-22, 1 female (5.4×3.8), NSMT-Cr S 1044; KY-08-26, 1 ovig. female (5.3 ×3.7), NSMT-Cr S 1045; KY-09-07, 1 female (6.2×4.4), NSMT-Cr S 1046; KY-09-14, 1 male (3.8×2.6), NSMT-Cr S 1047; KY-09-21, 3 ovig. females (5.2×3.6–5.7×3.8), 1 female (6.5× 4.6), NSMT-Cr S 1048; KY-09-21, 1 ovig. female (5.4×3.7), NSMT-Cr S 1049; KY-09-28, 3 females (5.5×3.7–5.5×4.1), 1 juv., NSMT-Cr S 1050; KY-09-29, 1 ovig. female (5.9×4.0), NSMT-Cr S 1051; KY-09-34, 1 young female, NSMT-Cr S 1052; KY-10-31, 1 ovig. female (4.9× 3.4), 1 young female (4.0×2.7), NSMT-Cr S 1053; KT-09-2-TW1-5, 1 female (6.5×4.0), NSMT-Cr S 1054; KT-09-2-TW2-4, 1 male (3.8× 2.6), NSMT-Cr S 1055; SY-09-04, 1 male (4.3× 2.5), NSMT-Cr S 1056; SY-09-09, 1 ovig. female (5.5×3.5), NSMT-Cr S 1057; SY-09-11, 1 ovig. female (6.2×3.9), NSMT-Cr S 1058. Remarks. This species is abundant in the offshore bottoms of the Ogasawara Islands, but it has not been found from the other area in the Japanese waters. Distribution. Widely distributed in the IndoWest Paciﬁc Oceans from east coast of Africa to Indonesia, Philippines, Ogasawara Is. and northern Australia, occurring at the depths of 47–300 m (Grifﬁn and Tranter, 1986; K. Sakai, 2004). Family Majidae Samouelle, 1819 Entomonyx spinosus Miers, 1884 [Jn: Togekaimen-gani] (Fig. 18A) Materials examined. KY-08-06, 1 male (14.0× 10.7), NSMT-Cr S 1059; KY-08-15, 1 male (8.4 ×6.1), NSMT-Cr S 1060; KY-09-13, 1 ovig. female (damaged, cw 9.8 mm), NSMT-Cr S 1061; KY-09-28, 1 male (11.1×9.2), NSMT-Cr S 1062; KY-09-30, 1 young male (7.4×5.4), NSMT-Cr S 1063; KT-09-2-TW1-1, 1 ovig. female (15.1× 11.5), NSMT-Cr S 1064. Distribution. Widely distribute in the Indian Ocean from Red Sea and Mozambique east to western Australia and in the West Paciﬁc Ocean from Japan and East China Sea, occurring at the depths of 60–150 m (K. Sakai, 2004). This is the ﬁrst record from the Ogasawara Islands. Leptomithrax biidus Ortmann, 1893 [Jn: Himekoshima-gani] (Fig. 18B) Leptomithrax biidus: Takeda and Kurata, 1976b: 25, ﬁg. 3B–D. Materials examined. KY-09-21, 1 young female (11.4×8.7), NSMT-Cr S 1065; KY-10-30, 1 ovig. female (25.8×23.0), NSMT-Cr S 1066; KY-10-31, 1 young female (9.8×7.4), NSMT-Cr S 1067; SY-09-05, 1 juv., NSMT-Cr S 1068. Distribution. Known only from Japan and Korea, occurring at the depths of 36–540 m (K. Sakai, 2004). Micippa parca Alcock, 1895 [Jn: Mitsutogetama-watakuzu-gani] (Fig. 18C) Micippa parca: Takeda, 1977: 124, pl. 2B. Materials examined. KY-09-28, 1 female (8.4 ×6.8), 1 young female (7.1×5.5), NSMT-Cr S 1069; KY-09-29, 2 young females (4.5×3.3, 5.0 ×3.8), NSMT-Cr S 1070; KY-09-30, 1 juv., NSMT-Cr S 1071; KY-10-06, 1 ovig. female (8.7× 7.0), NSMT-Cr S 1072; KY-10-24, 1 young female (6.4×5.5), NSMT-Cr S 1073. Distribution. Andaman Islands in the Indian Ocean and Japan, Sulu Archipelago, Indonesia, Hawaii and French Polynesia in the West Paciﬁc Ocean, occurring at shoreline deep to 72 m (Grifﬁn and Tranter, 1986; Poupin, 1996). Prismatopus occidentalis (Griffin, 1970) [New Jn: Minami-kaimen-gani] (Figs. 8O–Q, 18D) Materials examined. KY-08-25, 1 male (5.9× 4.2), 2 young males, 1 ovig. female (6.2×4.5), 2 Crabs dredged off the Ogasawara Islands 253 Fig. 18. A, Entomonyx spinosus Miers, 1884, male (14.0×10.7), NSMT-Cr S 1059; B, Leptomithrax biidus Ortmann, 1893, ovig. female (25.8×23.0), NSMT-Cr S 1066; C, Micippa parca Alcock, 1895, young female (7.1×5.5), NSMT-Cr S 1069; D, Prismatopus occidentalis (Grifﬁn, 1970), male (7.1×5.3), NSMT-Cr S 1077; E, Schizophroida hilensis (Rathbun, 1906), young female (4.5×3.1), 1 juv., NSMT-Cr S 1087; F, Seiitaoides orientalis (Sakai, 1961), young female (5.4×3.7), NSMT-Cr S 1088. female (6.8×5.0–8.2×6.2), NSMT-Cr S 1074; KY-09-07, 1 young male, 1 young female, NSMT-Cr S 1075; KY-09-21, 1 male (7.2×5.2), 1 female (8.1×5.9), NSMT-Cr S 1076; KY-09-33, 1 male (7.1×5.3), NSMT-Cr S 1077; KY-10-30, 1 male (6.9×5.0), NSMT-Cr S 1078; KT-09-2TW1-1, 2 ovig. females (7.5×5.4, 8.9×6.5), NSMT-Cr S 1079; KT-09-2-TW2-4, 1 male (7.1 ×5.3), NSMT-Cr S 1080; SY-09-03, 1 female (7.0×5.1), NSMT-Cr S 1081; SY-09-21, 1 male 254 Hironori Komatsu (7.4×5.3), 1 female (7.1×5.2), NSMT-Cr S 1082. Remarks. The present specimens agree well with the original description and illustration by Grifﬁn (1970), but differ in that 1) the chelipedal merus and carpus are less tuberculate; 2) the ﬁrst male gonopod more strongly outcurved (Fig. 8O). Distribution. Previously known only from western Australia, occurring at the depths of 112– 188 m (Grifﬁn and Tranter, 1986). This is the second record of the species and greatly extends its geographical range northward. Schizophroida hilensis (Rathbun, 1906) [Jn: Manazuru-nokogiri-gani] (Fig. 18E) ? Schizophroida simodaensis: Takeda, 1977: 124, ﬁg. 4C. Materials examined. KY-08-15, 1 young female (5.1×3.7), NSMT-Cr S 1083; KY-08-20, 1 young female (4.5×3.2), NSMT-Cr S 1084; KY09-29, 1 young male (4.5×3.2), NSMT-Cr S 1085; KY-09-30, 1 young male (4.5×3.2), NSMT-Cr S 1086; KY-10-07, 1 young female (4.5× 3.1), 1 juv., NSMT-Cr S 1087. Remarks. According to Griffin and Tranter (1986), the identity of each member of the genus Schizophroida, S. hilensis (Rathbun, 1906), S. manazuruana Sakai, 1933, and S. simodaensis Sakai, 1933, is still in question. Although Takeda (1977) recorded S. simodaensis from the Ogasawara Islands, the present specimen can be identiﬁed as S. hilensis by the morphological appearance including the shape of the male gonopod. Further revisional study is needed to clarify the identity of each species. Distribution. West Paciﬁc Ocean: Japan, New Caledonia, New Zealand and Hawaii, occurring at low tidal mark deep to 105 m (Griffin and Tranter, 1986; this study). This is the ﬁrst record from the Ogasawara Islands. Seiitaoides orientalis (Sakai, 1961) [Jn: Kofukitsuno-gani] (Fig. 18F) Eurynome orientalis: Takeda, 1977: 124. Materials examined. KY-08-15, 1 young female (5.4×3.7), NSMT-Cr S 1088; KY-09-21, 1 young female (4.7×3.3), NSMT-Cr S 1089; KY09-34, 1 young female (6.4×4.6), NSMT-Cr S 1090. Distribution. Japan, East China Sea, Philippines, Indonesia and western Australia, occurring at the depths of 45–245 m (Takeda and Komatsu, 2005). Superfamily Palicoidea Bouvier, 1898 Family Palicidae Bouvier, 1898 Neopalicus jukesii (White, 1847) [Jn: Shikakuitoashi-gani] (Fig. 19A) Neopalicus jukesii: Castro, 2000: 554, ﬁgs. 39b, 40b, c, 41c, 49, 61c. Materials examined. KY-08-15, 1 young female (4.5×5.0), NSMT-Cr S 1091; KY-09-28, 1 male (8.0×8.6), NSMT-Cr S 1092; KY-09-29, 1 broken young specimen (cb 6.1), NSMT-Cr S 1093; KY-09-30, 1 male (6.6×7.4), NSMT-Cr S 1094; KY-10-27, 1 ovig. female (8.1×9.1), NSMT-Cr S 1095; KY-10-31, 1 male (7.1×8.0), NSMT-Cr S 1096. Distribution. Widely distributed in the IndoWest Paciﬁc, occurring at the depths of 10–146 m (Castro, 2000). Pseudopalicus serripes (Alcock and Anderson, 1895) [Jn: Itoashi-gani] (Fig. 19B) Palicus aff. serripes: Takeda and Kurata, 1976a: 132, ﬁg. 6e. Materials examined. KY-09-30, 1 male (7.8× 8.9), NSMT-Cr S 1097; KY-10-31, 1 male (8.3× 9.3), 3 young males (4.5×4.8, 6.5×7.1), 2 broken specimens, NSMT-Cr S 1098. Remarks. Although Takeda and Kurata (1976a) recorded this species in question based on a young female from the Ogasawara Islands, the present specimens agree well with the diagnostic characters shown by Castro (2000: Table 2). Therefore, the record of Takeda and Kurata (1976a) can be considered as a juvenile of Pseudopalicus serripes. Distribution. Widely distributed in the IndoWest Pacific Oceans, occurring at the depths of 30–120 m (Castro, 2000). Crabs dredged off the Ogasawara Islands 255 Fig. 19. A, Neopalicus jukesii (White, 1847), male (8.0×8.6), NSMT-Cr S 1092; B, Pseudopalicus serripes (Alcock and Anderson, 1895), male (8.3×9.3), NSMT-Cr S 1098; C, Cavoportunus dubius (Laurie, 1906), male (9.8× 12.3), NSMT-Cr S 1113; D, Lupocyclus quinquedentatus Rathbun, 1906, young male (7.3×8.4), NSMT-Cr S 1114; E, Portunus (Xiphonectes) macrophthalmus Rathbun, 1906, male (7.7×15.8), NSMT-Cr S 1116; F, Portunus (Monomia) gradiator Fabricius, 1798, male (35.6×63.8), NSMT-Cr S 1115. Superfamily Parthenopoidea MacLeay, 1838 Family Parthenopidae MacLeay, 1838 Pseudolambrus beaumonti (Alcock, 1896) [Jn: Kobu-hishi-gani] (Fig. 20) ?Pseudolambrus beaumonti: Takeda and Kurata, 1976a: 124, ﬁg. 2g. Pseudolambrus ? sp.: Takeda and Kurata, 1976a: 125 (part), ﬁg. 5b. Materials examined. KY-08-17, 1 juvenile (4.2 ×4.5), NSMT-Cr S 1099; KY-08-21a, 1 male (7.1×7.1), NSMT-Cr S 1100; KY-08-21b, 1 male (6.2×6.9), NSMT-Cr S 1101; KY-08-25, 1 ovig. female (10.6×12.2), NSMT-Cr S 1102; KY-0826, 1 female (9.1×10.6), 2 young females (7.3× 8.4, 7.8×8.9), NSMT-Cr S 1103; KY-09-29, 1 ovig. female (9.3×10.9), NSMT-Cr S 1104; KY10-24, 1 young female (7.1×7.6), NSMT-Cr S 256 Hironori Komatsu 1105; SY-09-20, 1 female (10.3×12.2), NSMTCr S 1106. Remarks. As previously mentioned by Ng and Rahayu (2000), Pseudolambrus beaumonti (Alcock, 1896) has significant sexual dimorphism (Fig. 20). Takeda and Kurata (1976a) recorded this species in question based on the left cheliped obtained from the gut contents of Epinephelus fasciatus. This elongated cheliped agrees well with the major cheliped of males photographed by Ng and Rahayu (2000: ﬁg. 4). Takeda and Kurata (1976a) also recorded Pseudolambrus ? sp. based on the right cheliped and the anterior part of the carapace from the gut contents of Gymnocranius japonicus. The anterior part of the cara- pace (Takeda and Kurata, 1976a: Fig. 5b) may belong to this species, but the right cheliped (Takeda and Kurata, 1976a: Fig. 4c) may not. Because the upper margin of palm being straight (whereas having a basal lobe in Ps. beaumonti, but in juvenile the lobe is small; see Fig. 20). Kawamoto and Okuno (2003) recorded this species from Kume-jima Island, the Ryukyu Islands, but their specimen seems to belong to Ps. lobatus (Flipse, 1930), a senior subjective synonym of Ps. ozakii Sakai, 1969, due to rounded lobe on the base of the outer margin of the chelipedal palm. Distribution. Sri Lanka, Mauritius, Andaman Sea, Japan, occurring at the depths of 50–159 m Fig. 20. Pseudolambrus beaumonti (Alcock, 1896). A, male (7.1×7.1), NSMT-Cr S 1100; B, ovig. female (10.6× 12.2), NSMT-Cr S 1102; C, female (9.1×10.6), NSMT-Cr S 1103; D, juvenile (4.2×4.5), NSMT-Cr S 1099. Crabs dredged off the Ogasawara Islands (Michel, 1964; Sakai, 1976; this study). Superfamily Portunoidea Raﬁnesque, 1815 Family Portunidae Raﬁnesque, 1815 Cavoportunus dubius (Laurie, 1906) [Jn: Yoronhime-gazami] (Fig. 19C) Cavoportunus dubius: Nguyen and Ng, 2010: 41, ﬁgs. 1–4. Materials examined. KY-08-15, 1 young female (10.2×13.0), NSMT-Cr S 1107; KY-08-18, 1 young male (6.5×8.2), NSMT-Cr S 1108; KY08-26, 1 young male (6.7×8.4), NSMT-Cr S 1109; KY-09-27, 1 young male (6.8×8.9), NSMT-Cr S 1110; KY-10-02, 1 male (17.2×21.3), NSMT-Cr S 1111; KY-10-31, 1 young male (5.2 ×6.2), NSMT-Cr S 1112; SY-09-18, 1 male (9.8 ×12.3), 2 young males (7.8×9.8, 9.0×11.2), NSMT-Cr S 1113. Remarks. Recently, Nguyen and Ng (2010) established a new genus, Cavoportunus, for this species and synonymized Portunus (Cycloachelous) yoronensis Sakai, 1974, with this species. The present specimens agree well with the subsequent description and illustration by Nguyen and Ng (2010). Distribution. Reunion, India, Sri Lanka, Indonesia, Philippines, Japan, French Polynesia, occurring at coral reefs and the depths of 20–210 m (Crosnier, 2002; Nguyen and Ng, 2010). Lupocyclus quinquedentatus Rathbun, 1906 [Jn: Mutsuha-hosoude-gazami] (Fig. 19D) Material examined. KY-09-29, 1 young male (7.3×8.4), NSMT-Cr S 1114. Distribution. Widely distributed in the IndoWest Paciﬁc Ocean from Seychelles eastward to French Polynesia, occurring at the depths of 60– 110 m (K. Sakai, 2004; this study). This is the ﬁrst record from the Ogasawara Islands. Portunus (Monomia) gradiator Fabricius, 1798 [Jn: Ibo-gazami] (Fig. 19F) Portunus (Monomia) haani: Takeda, 1977: 128. Materials examined. KY-09-27, 1 male (35.6× 63.8), 2 young males (12.9×23.7, 13.2×25.2), 257 NSMT-Cr S 1115. Distribution. Widely distributed in the IndoWest Paciﬁc Oceans from South Africa eastward to New Caledonia, occurring at the depths of 10– 345 m (K. Sakai, 2004). Portunus (Xiphonectes) macrophthalmus Rathbun, 1906 [Jn: Ohme-tenaga-hime-gazami] (Figs. 19E, 22A, B) Portunus (Xiphonectes) macrophthalmus: Takeda, 1977: 128, pl. 4(C, D) Materials examined. KY-10-26, 1 male (7.7× 15.8, cw excluding lateral spines 9.5), 1 young female (6.8×8.9), 3 juvs., NSMT-Cr S 1116. Distribution. Mauritius, Banda Sea, Philippines, Japan, Hawaii; occurring at the depths of 2–100 m (Takeda, 1977). Portunus (Xiphonectes) tenuipes (de Haan, 1835) [Jn: Kebuka-ibo-gazami] (Fig. 21A) Portunus (Monomia) tenuipes: Takeda, 1977: 129. Materials examined. KY-10-25, 3 males (13.5 ×19.0–15.5×22.4), 3 young males (6.7×9.3– 9.9×13.7), 3 young females (8.9×12.3–12.6× 17.8), NSMT-Cr S 1117; KY-10-26, 1 young male (6.8×8.9), 3 juvs., NSMT-Cr S 1118. Distribution. Andaman Sea, Philippines, Japan, Australia, occurring at coral reef and deep to 45 m (Takeda, 1977). Thalamita auauensis Rathbun, 1906 [New Jn: Auau-benitsuke-gani] (Figs. 21D, 22G–I) Materials examined. KY-08-15, 1 young male, 1 young female, 2 juvs., NSMT-Cr S 1119; KY08-20, 1 young female, NSMT-Cr S 1120; KY09-08, 1 juv., NSMT-Cr S 1121; KY-09-28, 1 young male, NSMT-Cr S 1122; KY-09-29, 1 young male, NSMT-Cr S 1123; KY-09-30, 2 young males (3.9×6.1, 5.0×7.5), NSMT-Cr S 1124; KY-10-27, 1 male (6.8×11.3), 1 young female, NSMT-Cr S 1125. Remarks. The present specimens agree well with the original and subsequent descriptions and illustrations (Rathbun, 1906; Edmondson, 1954; Stephenson and Rees, 1967; Crosnier, 2002). 258 Hironori Komatsu Fig. 21. A, Portunus (Xiphonectes) tenuipes (de Haan, 1835), male (14.4×20.6), NSMT-Cr S 1117; B, Thalamita integra integra Dana, 1852, males (7.8×12.5), NSMT-Cr S 1132; C, Thalamita gatavakensis Nobili, 1906, male (6.8 ×11.3), NSMT-Cr S 1126; D, Thalamita auauensis Rathbun, 1906, male (6.8×11.3), NSMT-Cr S 1125; E, Thalamita sexlobata Miers, 1886, young male (3.7×4.7), NSMT-Cr S 1137; F, Opecarcinus crescentus (Edmondson, 1925), female (2.7×2.1), NSMT-Cr S 1140. Distribution. Philippines, Mariana, China, Japan, Hawaii Is., Samoa; occurring at the depths of 24–335 m (Rathbun, 1906; Edmondson, 1954; Stephenson and Rees, 1967; Crosnier, 2002; this study). This is the ﬁrst record from Japan and the Ogasawara Islands. Thalamita gatavakensis Nobili, 1906 [Jn: Gatavak-benitsuke-gani] (Figs. 21C, 22C–F) Materials examined. KY-08-11, 1 male (6.8× 11.3), NSMT-Cr S 1126; KY-08-18, 1 juv., NSMT-Cr S 1127; KY-09-21, 1 young male, NSMTCr S 1128; KY-09-30, 1 female (6.8×11.1), 1 young female (4.6×6.7), NSMT-Cr S 1129; KY10-27, 1 young female, NSMT-Cr S 1130. Remarks. The present specimens agree well with the subsequent descriptions and illustrations including male gonopods by Forest and Guinot (1961) and Crosnier (1962). Sakai (1976) recorded Thalamita admete Crabs dredged off the Ogasawara Islands 259 Fig. 22. Right gonopods. A, B, Portunus (Xiphonectes) macrophthalmus Rathbun, 1906, male (7.7×15.8), NSMT-Cr S 1116, A, G1, abdominal; B, G2, abdominal. C–F, Thalamita gatavakensis Nobili, 1906, male (6.8×11.3), NSMTCr S 1126. G–I, Thalamita auauensis Rathbun, 1906, male (6.8×11.3), NSMT-Cr S 1125, G, G1, abdominal; H, tip of same; I, G2, abdominal. J, K, Thalamita spinifera Borradaile, 1902, male (6.6×8.7), NSMT-Cr S 1139, J, G1, abdominal; K, G2, abdominal. (Herbst, 1803), from the Bonin (Ogasawara) Islands, however, it is possible that his specimens actually belong to Th. auauensis or Th. gatavakensis because they are very similar to each other and Th. admete was never collected by this survey. Takeda and Kurata (1977a) also recorded Th. cf. admete from the gut contents of lethrinid ﬁsh, Lethrinus variegatus, from the new volcanic island, Nishino-shima-shinto Island, on the basis of several broken carapaces and chelae. Their specimens also may belong to Th. auauensis or Th. gatavakensis by same reason. Distribution. Madagascar, Seychelles, Indonesia, Philippines, Saipan, Japan, Australia, French Polynesia, occurring at sublittoral water and deep to 135.8 m (Stephenson, 1972; Davie, 2002; Marumura and Kosaka, 2003; this study). This is the first record from the Ogasawara Islands and extends the bathymetrical range deep to 135.8 m. Thalamita integra integra: Stimpson, 1858: 39; 1907: 83; Ooishi, 1970: 92, pl. 13 ﬁg. 9. Materials examined. Futami Port, Chichi-jima I., coll. H. Komatsu, 17 Nov. 2009, 1 young male (3.7×5.6), 1 female (9.9×16.3), 1 young female (6.7×10.7), NSMT-Cr S 1131; Oki Port, Haha-jima I., coll. H. Komatsu, 14 Jul. 2009, 2 males (6.0×9.5, 7.8×12.5), 2 ovig. females (6.2× 11.6, 7.8×12.2), 1 female (6.6×10.8), NSMT-Cr S 1132; Ohmura Beach, Chichi-jima I., coll. T. Komai, 17 Jul. 2009, 1 young male (2.9×4.1), NSMT-Cr S 1133. Remarks. This species did not occur in the collections made by dredging, but is listed herein as reference. This species is abundant in coastal area. Distribution. Widely distributed in the IndoWest Paciﬁc Oceans, occurring at shallow water (K. Sakai, 2004). Thalamita integra integra Dana, 1852 [Jn: Himefutaha-benitsuke-gani] (Fig. 21B) Thalamita oculea Alcock, 1899 [Jn: Oculea-benitsuke-gani] (Fig. 5F) 260 Hironori Komatsu Materials examined. KY-09-28, 1 young male (4.7×6.7), NSMT-Cr S 1134; KY-09-29, 1 young female (6.3×9.0), NSMT-Cr S 1135; KY-09-30, 3 young male (4.4×6.4–5.1×7.1), 3 juvs., NSMT-Cr S 1136. Remarks. The anterolateral teeth of the carapace of the present specimens are narrower than that of the material from Madagascar photographed by Crosnier (1962). In young specimens, transverse sculpture on the sternum and the abdomen is not developed. Distribution. Madagascar, Seychelles, Saya de Malha Bank, India, Maldives, Sri Lanka, Andamann Sea, Japan, occurring at the depths of 15– 60.7 m (K. Sakai, 2004; this study). This is the ﬁrst record from the Ogasawara Islands. Thalamita sexlobata Miers, 1886 [Jn: Mutsuhabenitsuke-gani] (Fig. 21E) Material examined. SY-09-03, 1 young male (3.7×4.7), NSMT-Cr S 1137. Distribution. Widely distributed in the IndoWest Pacific Oceans from Madagascar eastward to Tonga, occurring at the depths of 5–146 m (K. Sakai, 2004; this study). This is the first record from the Ogasawara Islands and extends the ba- thymetrical range deep to 146 m. Thalamita spinifera Borradaile, 1902 [Jn: Indobenitsuke-gani] (Figs. 5G, 22J, K) Materials examined. KY-10-24, 1 young female (7.6×10.4), NSMT-Cr S 1138; KY-10-31, 1 male (6.6×8.7), NSMT-Cr S 1139. Remarks. The shape of the male gonopod of the present specimen agrees well with the illustrations by Sakai (1935, as Thalamita exetastica Alcock, 1899), Stephenson and Rees (1967) and Wee and Ng (1995). Although Takeda and Kurata (1976a) recorded the close congener, Th. aff. exetastica from the Ogasawara Islands, Th. spinifera can be easily distinguished from Th. aff. exetastica by the presence of spines on the posterior border of P5 propodi and the arrangement of bristles on the distal part of the ﬁrst male gonopod. A key to the species of the genus Thalamita known from the Ogasawara Islands is provided. Distribution. Madagascar, Laccadives, Maldives, Thailand, Malaysia, Indonesia, Philippines, Japan, Hawaii (Stephenson, 1972; Sakai, 1976; Wee and Ng, 1995). This is the ﬁrst record from the Ogasawara Islands. Key to the species of the genus Thalamita known from the Ogasawara Islands 1. Front 6-lobed ………………………………………………………………………………………… 2 Front 4-lobed …………………………………………………………………………… T. sexlobata Front 2-lobed ………………………………………………………………………………………… 3 2. Posterior border of P5 propodi with a variable number of spines; distal part of G1 ﬂared, with several bristles on lateral margin……………………………………………………………… T. spinifera Posterior border of P5 propodi without spines; distal part of G1 ﬂared, with several bristles on both margins ………………………………………………………………………… T. aff. exetastia 3. Basal antennal segment with smooth crest ………………………………………………………… 4 Basal antennal segment granular or tubercular ……………………………………………………… 5 4. Outer border of upper surface of chelipedal hand with obsolete spine …………………… T. oculea Outer border of upper surface of chelipedal hand with spine ………………………………… T. sima 5. Frontal region with soft setae ……………………………………………………… T. integra integra Frontal region without setae ………………………………………………………………………… 6 6. Inner supraorbital lobe rather arcuate; ambulatory legs without brown band; tip of G1 bilobed, with 4 bristles on each border …………………………………………………………… T. auauensis Inner supraorbital lobe almost straight; ambulatory legs with brown band; tip of G1 recurved with several bristles asymmetrically ……………………………………………………… T. gatavakensis Crabs dredged off the Ogasawara Islands Subsection Thoracotremata Guinot, 1977 Superfamily Cryptochiroidea Paul’son, 1875 Family Cryptochiridae Paul’son, 1875 Opecarcinus crescentus (Edmondson, 1925) [Jn: Mikazuki-ashibiro-sango-yadori-gani] (Fig. 21F) Materials examined. KY-09-14, 1 male (2.0× 1.6), 1 ovig. female (2.7×2.1), NSMT-Cr S 1140. Host. The present specimens are collected from Leptoseris hawaiiensis [Jn: Hawaii-senbei-sango] for the first time. This species was previously known from Pavona spp. [Shikoro-sango-zoku] and Coscinaraea columna [Yasuri-sango]. Distribution. Known from the West Paciﬁc (Japan, Taiwan, Vietnam, Indonesia, Christmas Is., Johnston I., Hawaii) and also from the eastern Pacific (Clippertib I. and Baja California) (Takeda and Tamura, 1981; Wei et al., 2006). This is the ﬁrst record from the Ogasawara Islands. Discussion Summary of result A total of 73 species belonging to 16 families excluding pilumnoid and xanthoid crabs were obtained from the sea around the Ogasawara Islands at the depths of 36–499 m by this survey. Of these three new species are described and illustrated: Cryptodromia maculata sp. nov. (Dromiidae), Ebalia koyo sp. nov. (Leucosiidae), and Pugettia ogasawaraensis sp. nov. (Epialtidae). 42 species (62% of the total 73 species) are newly added to the carcinological fauna of the Ogasawara Islands: Epigodromia rotunda, Homalodromia coppingeri, Lauridromia intermedia (Dromiidae), Dynomene pilumnoides (Dynomenidae), Notopus dorsipes, Notosceles serratifrons, Ranina ranina, Umalia misakiensis, U. orientalis (Raninidae), Sakaila imperialis (Aethridae), Ethusa quadrata (Ethusidae), Goneplacoides marivenae, Singhaplax styrax (Goneplacidae), Ebalia sakaii, E. tuberculosa, Heteronucia perlata, H. toyoshioae, Oreotlos angulatus, O. lagarodes, Praebebalia dondonae, Toru granuloides (Leucosiidae), Huenia pacifica, Hyaste- 261 nus ambonensis, H. kyusyuensis, Naxioides taurus, Phalangipus hystrix, Xenocarcinus conicus (Epialtidae), Chalaroachaeus curvipes, Grypachaeus hyalinus, Pleistacantha simplex, Prosphorachaeus suluensis (Inachidae), Entomonyx spinosus, Prismatopus occidentalis, Schizophroida hilensis (Majidae), Cavoportunus dubius, Lupocyclus quinquedentatus, Thalamita auauensis, Th. gatavakensis, Th. oculea, Th. sexlobata, Th. spinifera (Portunidae), and Opecarcinus crescentus (Cryptochiridae). Of these 5 species are also newly added to the carcinological fauna of Japan: Epigodromia rotunda (Dromiidae), Praebebalia dondonae (Leucosiidae), Hyastenus ambonensis (Epialtidae), Prismatopus occidentalis (Majidae), and Thalamita auauensis (Portunidae). The sampling sites which are rich in benthic crabs are the channel between Ani-jima Island and Nishi-jima Island (KY-09-28, KY-09-29 and KY-09-30), the submarine bank situated west of Futami Bay (KY-08-15 and KY-08-26) and the submarine bank situated northwest of Ototo-jima Island (KY-09-21) (Fig. 1). On the contrary, the bottoms deeper than 150 m are very poor in benthic crabs. An annotated checklist of crabs from the Ogasawara Islands modiﬁed from Takeda and Miyake (1976) is provided (Appendix). A total of 250 species belonging to 41 families hitherto has been recorded from the islands, including unidentiﬁed or possibly misidentiﬁed species. Comparison with Takeda (1977) Takeda (1977) recorded 21 species of crabs from the west side of Chichi-jima Island at the depths of 36–110 m, excluding pilumnoid and xanthoid crabs. Of these 7 species did not occur in this study: Osachila expansa (Aethridae), Philyra syndactyla (Leucosiidae), Oncinopus neptunus (Inachidae), Aulacolambrus diacanthus (Parthenopidae), Libystes lepidus (Portunidae), Macrophthalmus (Macrophthalmus) telescopicus (Macrophthalmidae), Tetrias ischeri (Pinnotheridae). These species habit on sandy bottom of shallow water except O. neptunus, but the present 262 Hironori Komatsu survey focused on harder and deeper bottoms. Therfore these species did not occur in this study and in hence 59 species occurred in addition to Takeda (1977). Biogeography The brachyuran crabs occurring at the sea around the Ogasawara Islands by the present study can be classiﬁed into three groups based on general distributional pattern: (1) species widely distributed in the West Pacific or Indo-West Pacific; (2) species endemic to East Asia; and (3) species known only from the Ogasawara Islands. (1) West Paciﬁc or Indo-West Paciﬁc elements include 61 species (84% of the total 73 species). Most of the species occurred in this study were classiﬁed into this elements. (2) East Asian endemic elements contain 8 species (11%): Umalia orientalis (Raninidae), Henicoplax nitida (Euryplacidae), Ebalia hayamaensis, E. koyo sp. nov., E. sakaii, Heteronucia toyoshioae (Leucosiidae), Hyastenus kyusyuensis (Epialtidae), Leptomithrax bifidus (Majidae). These species are thought to be drifted from the mainland of Japan by the Kuroshio counter current (Fig. 1). (3) Species known only from the Ogasawara Islands include 4 species (5%): Cryptodromia maculata sp. nov. (Dromiidae), Praebebalia taeniata (Leucosiidae), Pugettia ogasawaraensis sp. nov. (Epialtidae), Achaeus boninensis (Inachidae). To conﬁrm that these species are really endemic to the Ogasawara Islands, faunal surveys of the neighbor area including the Ryukyu and the Mariana Islands are needed. Acknowledgements I deeply indebted to Kunihisa Yamaguchi, Yuhei Tanaka, Yuji Aoki, Masami Yamakawa, Ichiro Gonoi and the crews of R/V Koyo (Tokyo Metropolitan Ogasawara Fisheries Center), Kotaro Tsuchiya, Akira Kitazawa and the crews of TR/V Shin’yo Maru (Tokyo University of Marine Science and Technology), the captain and the crews of R/V Tansei Maru (JAMSTEC) and all the members joined the field surveys for their help in collecting specimens. 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Descriptions of a new genus and five new species of Crustacea. In: J.B. Jukes (ed), Narrative of the surveying voyage of H.M.S. Fly, commanded by Captain F.P. Blackwood, R.N. in Torres Strait, New Guinea, and other islands of the Eastern Archipelago, during the years 1842–1846; together with an excursion into the interior of the eastern part of Java. Volume 8, Appendix, pp. 335–338, T. & W. Boone, London. Yokoya, Y., 1933. On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S.S. Soyo-Maru, during the year 1923–1930. Journal of the College of Agriculture, Tokyo Imperial University, 12: 1–225. Yoshihara, S., 1901. Animals of the Bonin Islands. Zoological Magazine, 13: 309–316. (In Japanese). Crabs dredged off the Ogasawara Islands 小笠原諸島近海からドレッジにより採集されたカニ類小松浩典小笠原諸島近海からドレッジにより採集されたカニ類を報告する．採集されたカニ類は 16 科 73 種に分類された（ただしケブカガニ類およびオウギガニ類を除く）．コモンカイカムリ（新称）Cryptodromia maculata sp. nov.（カイカムリ科），コウヨウエバリア（新称）Ebalia koyo sp. nov.（コブシガニ科）およびオガサワラモガニ（新称）Pugettia ogasawaraensis sp. nov.（ツノガニ科）の 3 種を新種記載した．また，42 種のカニ類を新たに小笠原の甲殻類相に加え，そのうちの 5 種は日本新記録となる．記録された種について生物地理的な考察を加えた． 269 270 Hironori Komatsu Appendix. Check list of species of brachyuran crabs from the Ogasawara Islands. Section Podotremata Guinot, 1977 Superfamily Dromioidea de Haan, 1838 Family Dromiidae de Haan, 1838 Cryptodromia coronata Stimpson, 1858 Cryptodromia maculata sp. nov Cryptodromia tumida Stimpson, 1858 Dromidiopsis lethrinusae (Takeda and Kurata 1976) Epigodromia rotunda McLay, 1993 Homalodromia coppingeri Miers, 1884 Lauridromia intermedia (Laurie, 1906) Takedromia yoshidai (Takeda and Kurata, 1976) Stimpson (1858, 1907) this study Ooishi (1970) Takeda and Kurata (1976a) this study this study this study Takeda and Kurata (1976b); this study Family Dynomenidae Ortmann, 1892 Dynomene pilumnoides Alcock, 1900 Dynomene praedator A. Milne-Edwards, 1879 this study Sakai (1976) Superfamily Homoloidea de Haan, 1839 Family Latreilliidae Stimpson, 1858 Eplumula phalangium (de Haan, 1839) Takeda and Kurata (1984, megalopa) Superfamily Raninoidea de Haan, 1839 Family Raninidae de Haan, 1839 Notopoides latus Henderson, 1888 Notopus dorsipes (Linnaeus, 1758) Notosceles serratifrons (Henderson, 1893 Ranina ranina (Linnaeus, 1758) Umalia misakiensis (Sakai, 1937) Umalia orientalis (Sakai, 1963) Takeda and Kurata (1984); this study this study this study this study this study Takeda (1977); this study Section Eubrachyura Saint Laurent, 1980 Subsection Heterotremata Guinot, 1977 Superfamily Aethroidea Dana, 1851 Family Aethridae Dana, 1851 Actaeomorpha erosa Miers, 1877 Drachiella angulata (Ihle, 1918) Drachiella caelata Takeda and Tachikawa, 1995 Osachila expansa Takeda, 1977 Sakaila imperialis (Sakai, 1963) Takeda and Kurata (1977b) Takeda and Tachikawa (1995) Takeda and Tachikawa (1995) Takeda (1977) this study Superfamily Calappoidea de Haan, 1833 Family Calappidae de Haan, 1833 Calappa hepatica (Linnaeus, 1758) Family Matutidae de Haan, 1835 Matuta banksii Leach, 1817 Matuta lunaris (Forskål, 1775) Matuta planipes Fabricius, 1798 Superfamily Carpilioidea Ortmann, 1893 Yoshihara (1901); Takeda and Kurata (1977a); Takeda (1977) Balss (1922) Matsuura (1894, as M. victrix); Yoshihara (1901, as M. victor); Terazaki (1902, as M. victor) Stimpson (1858, 1907, as M. lunaris); Ooishi (1970) Crabs dredged off the Ogasawara Islands 271 Appendix. (cont.) Family Carpiliidae Ortmann, 1893 Carpilius convexus (Forskål, 1775) Carpilius maculatus (Linnaeus, 1758) Superfamily Dairoidea Serène, 1965 Family Dairidae Serène, 1965 Daira perlata (Herbst, 1790) Shigei (1970) Sakai (1939) Yoshihara (1901, as D. variolosa); Sakai (1939); Imajima (1970); Ooishi (1970) Superfamily Dorippoidea MacLeay, 1838 Family Ethusidae Guinot, 1977 Ethusa quadrata Sakai, 1937 this study Superfamily Eriphioidea MacLeay, 1838 Family Dairoididae Števčić, 2005 Dairoides kusei (Sakai, 1938) Takeda and Kurata (1976b); this study Family Eriphiidae MacLeay, 1838 Eriphia scabricula Dana, 1852 Epixanthus corrosus A. Milne-Edwards, 1873 Ooishi (1970); Shigei (1970) Asakura et al. (1994b) Family Hypothalassiidae Karasawa and Schweitzer, 2006 Hypothalassia armata (de Haan, 1835) Takeda and Kurata (1976b) Family Oziidae Dana, 1851 Epixanthus corrosus A. Milne-Edwards, 1873 Epixanthus frontalis (H. Milne Edwards, 1834) Lydia annulipes (H. Milne Edwards, 1834) Ozius rugulosus Stimpson, 1858 Superfamily Goneplacoidea MacLeay, 1838 Family Euryplacidae Stimpson, 1871 Heteroplax nitida Miers, 1879 Platyozius laevis (Borradaille, 1902) Family Goneplacidae MacLeay, 1838 Carcinoplax eburnea Stimpson, 1858 Goneplacoides marivenae (Komatsu and Takeda, 2003) Singhaplax styrax Castro, 2007 Superfamily Leucosioidea Samouelle, 1819 Family Leucosiidae Samouelle, 1819 Ebalia hayamaensis Sakai, 1963 Ebalia humilis Takeda, 1977 Ebalia koyo sp. nov. Ebalia sakaii Takeda and Miyake, 1972 Ebalia tuberculosa (A. Milne-Edwards, 1873) Imajima (1970); Ooishi (1970) Yoshihara (1901) Ooishi (1970); Takeda and Kurata (1976b) Stimpson (1858, 1907); Balss (1922); Imajima (1970); Ooishi (1970); Takeda and Kurata (1977a) Takeda (1977); this study Castro and Ng (2010); this study Stimpson (1858, 1907) (=?Libystes nitidus, see Serène and Lohavanijaya (1972: 62)) this study this study Takeda (1977); this study Takeda (1977); Takeda and Kurata (1977b, as Praebebalia taeniata; 1984); this study this study this study this study 272 Hironori Komatsu Appendix. (cont.) Heteronucia perlata (Sakai, 1963) Heteronucia toyoshioae Komatsu and Takeda, 2005 Leucosia anatum (Herbst, 1783) Leucosia sp. Nucia speciosa Dana, 1852 Oreotlos angulatus (Rathbun, 1906) Oreotlos heuretos Tan and Ng, 1995 Oreotlos lagarodes Tan and Ng, 1995 Philyra syndactyla Ortmann, 1892 Praebebalia dondonae Chen, 1989 Praebebalia taeniata Takeda, 1977 Toru granuloides (Sakai, 1961) Urnalana insularis Takeda and Kurata, 1976 this study this study Takeda and Kurata (1977a); Takeda (1977) Takeda and Kurata (1984) Takeda and Kurata (1976b) this study Takeda (1977, as Oreophorus (Oreotlos) latus); this study this study Takeda (1977) this study Takeda (1977); Takeda and Kurata (1984); this study this study Takeda and Kurata (1976b) Superfamily Majoidea Samouelle, 1819 Family Hymenosomatidae MacLeay, 1838 Elamena truncata (Stimpson, 1858) Terazaki (1902, as Halicarcinus sp.) Family Epialtidae MacLeay, 1838 Hoplophrys oatesi Henderson, 1893 Huenia heraldica (de Haan, 1837) Huenia paciica Miers, 1879 Hyastenus ambonensis Grifﬁn and Tranter, 1986 Hyastenus diacanthus (de Haan, 1839) Hyastenus kyusyuensis (Yokoya, 1933) Lahaina ovata Dana, 1951 Menaethius monoceros (Latreille, 1825) Micippoides angustifrons A. Milne-Edwards, 1873 Naxioides robillardi (Miers, 1882) Naxioides taurus (Pocock, 1890) Perinia tumida Dana, 1851 Phalangipus hystrix (Miers, 1886) Pugettia ogasawaraensis sp. nov. Thusaenys cornigerus (Sakai, 1938) Tylocarcinus styx (Herbst, 1803) Xenocarcinus conicus (A. Milne-Edwards, 1865) Xenocarcinus tuberculatus White, 1847 Family Inachidae MacLeay, 1838 Achaeus boninensis Miyake and Takeda, 1969 Achaeus cf. brevidactylus Sakai, 1938 Achaeus brevifalcatus Rathbun, 1911 Achaeus curvirostris (A. Milne-Edwards, 1873) Achaeus sp. Camposcia retusa Latreille, 1829 Chalaroachaeus curvipes de Man, 1902 Cyrtomaia lamellata Rathbun, 1906 Ooishi (1970, as H. ogilbryi MacCulloch, 1908); this study Takeda and Kurata (1976a, as H. proteus) this study this study Terazaki (1902) this study Takeda (1977, as Hyastenus tenuicornis); Takeda and Kurata (1977b, as H. tenuicornis); this study Takeda and Kurata (1976a, 1977a, b); Takeda (1977); this study Sakai (1955); Ooishi (1970); Takeda and Kurata (1977a) Takeda and Kurata (1976b, as N. mammillata) Takeda and Kurata (1976a, as N. mammillata); this study Imajima (1970, as Perinea tumida) this study this study Takeda and Kurata (1976b) Ooishi (1970) this study Takeda (1973); this study Miyake and Takeda (1969); Takeda and Kurata (1976a); this study this study Miyake and Takeda (1969); this study Takeda (1977, as A. issifrons); this study Takeda and Kurata (1977b) Yoshihara (1901); Terazaki (1902) this study Takeda and Kurata (1976b, as C. platypes); this study Crabs dredged off the Ogasawara Islands 273 Appendix. (cont.) Grypachaeus hyalinus (Alcock and Anderson, 1894) Oncinopus angustifrons Takeda and Miyake, 1969 Oncinopus neptunus Adams and White, 1848 Paratymolus bituberculatus Haswell, 1880 Pleistacantha simplex Rathbun, 1932 Prosphorachaeus suluensis Rathbun, 1916 Sunipea indicus (Alcock, 1895) this study Takeda and Miyake (1969b); Takeda (1973); Takeda and Kurata (1976a) Takeda (1977) Takeda (1977); this study this study this study Takeda (1977); this study Family Majidae Samouelle, 1819 Cyclax suborbicularis (Stimpson, 1858) Entomonyx spinosus Miers, 1884 Leptomithrax biidus (Ortmann, 1893) Micippa parca Alcock, 1895 Micippa philyra (Herbst, 1803) Maja sp. Prismatopus occidentalis (Grifﬁn, 1970) Schizophroida hilensis (Rathbun, 1906) Schizophroida simodaensis Sakai, 1933 Schizophrys aspera (H. Milne Edwards, 1834) Seiitaoides orientalis (Sakai, 1961) Tiarinia spinigera Stimpson, 1857 Ooishi (1970) this study Takeda and Kurata (1976b); this study Takeda (1977); this study Terazaki (1903) Takeda and Kurata (1976a) this study this study Takeda (1977) Imajima (1970); Takeda and Kurata (1976a) Takeda (1977); this study Takeda and Kurata (1977a, b) Superfamily Palicoidea Bouvier, 1898 Family Palicidae Bouvier, 1898 Neopalicus jukesii (White, 1847) Pseudopalicus serripes (Alcock and Anderson, 1895) Castro (2000); this study Takeda and Kurata (1976a, as Ps. aff. serripes); this study Superfamily Parthenopoidea MacLeay, 1838 Family Parthenopidae MacLeay, 1838 Aulacolambrus diacanthus (de Haan, 1839) Daldoria horrida (Linnaeus, 1758) Furtipodia pertosa (Klunzinger, 1906) Pseudolambrus beaumontii Alcock, 1895 Pseudolambrus? sp. Superfamily Pilumnoidea Samouelle, 1819 Family Pilumnidae Samouelle, 1819 Actumnus foricigerus (Stimpson, 1858) Actumnus intermedius Balss, 1922 Actumnus setosiareolatus Takeda, 1977 Caecopilumnus crassipes (Tesch, 1918) Echinoecus pentagonus (A. Milne-Edwards, 1879) Gonatonotus granulosus (MacGilchrist, 1905) Gorgonariana sodalis (Alcock, 1898) Lophoplax sextuberculata Takeda and Kurata, 1984 Mertonia lanka Laurie, 1906 Pilumnus ikedai Takeda and Miyake, 1968 Pilumnus longicornis Hilgendorf, 1878 Takeda (1977); Takeda and Kurata (1977b) Matsuura (1894); Terazaki (1904) Komatsu and Takeda (2009) Takeda and Kurata (1976a, as ?Ps. beaumontii); this study Takeda and Kurata (1976a) Takeda and Miyake (1969a) Takeda (1977) Takeda (1977) Sakai, 1955 Rathbun (1894); Miyake (1939, as E. rathbunae nom. nov.) Ooishi (1970) Odhner (1925) Takeda and Kurata (1984) Takeda (1977) Takeda and Miyake (1968a) Imajima (1970) 274 Hironori Komatsu Appendix. (cont.) Pilumnus minutus de Haan, 1835 Planopilumnus pygmaeus Takeda, 1977 Pseudolitochira integra (Miers, 1884) Zehntneriana novaeinsulicola (Takeda and Kurata, 1977) Superfamily Portunoidea Raﬁnesque, 1815 Family Portunidae Raﬁnesque, 1815 Cavoportunus dubius (Laurie, 1906) Coelocarcinus foliatus Edmondson, 1930 Libystes lepidus Miyake and Takeda, 1970 Libystes villosus Rathbun, 1924 Lissocarcinus orbicularis Dana, 1852 Lupocyclus quinquedentatus Rathbun, 1906 Portunus (Achelous) cf. granulatus granulatus (H. Milne Edwards, 1834) Portunus (Monomia) gradiator Fabricius, 1798 Portunus (Xiphonectes) gracillimus (Stimpson, 1858) Portunus (Xiphonectes) macrophthalmus Rathbun, 1906 Portunus (Xiphonectes) tenuipes (de Haan, 1835) Portunus sp. Scylla serrata (Forskål, 1775) Thalamita admete (Herbst, 1803) Thalamita aff. admete (Herbst, 1803) Thalamita auauensis Rathbun, 1906 Thalamita gatavakensis Nobili, 1906 Thalamita aff. exetastica Alcock, 1899 Thalamita integra integra Dana, 1852 Thalamita oculea Alcock, 1899 Thalamita sexlobata Miers, 1886 Thalamita sima H. Milne Edwards, 1834 Thalamita spinifera Borradaile, 1902 Thalamita sp. Superfamily Pseudozioidea Alcock, 1898 Family Pseudoziidae Alcock, 1898 Pseudozius caystrus (Adams and White, 1849) Stimpson (1858, 1907, as P. hirsutus sp. nov.) Takeda (1977) Sakai (1955) Takeda and Kurata (1977a) Nguyen and Ng (2010); this study Takeda and Kurata (1977b) Miyake and Takeda (1970) Takeda (1977) Imajima (1970); Ooishi (1970) this study Takeda and Kurata (1977a) Takeda (1977, as P. (M.) haanii); this study Stimpson (1858, 1907); ?Takeda and Kurata (1976a) Takeda (1977); this study Takeda (1977); this study Takeda and Kurata (1977a); Takeda (1977) Yoshihara (1901) Sakai (1976) Takeda and Kurata (1977a) this study this study Takeda and Kurata (1976a) Stimpson (1858, 1907); Ooishi (1970); this study this study this study Terazaki (1904); Takeda and Kurata (1976a) this study Takeda and Kurata (1977b) Pseudozius inornatus Dana, 1852 Stimpson (1858, 1907, as P. microphthalmus); Ooishi (1970) Ooishi (1970) Superfamily Trapezioidea Miers, 1886 Family Domeciidae Ortmann, 1893 Domecia hispida Eydoux and Souleyet, 1842 Ooishi (1970) Family Tetraliidae Castro, Ng and Ahyong, 2004 Tetralia glaberrima (Herbst, 1799) Tetraloides heterodactylus (Heller, 1861) Stimpson (1858, 1907); Ooishi (1970) Imajima (1970)［Tetralia］ Family Trapeziidae Miers, 1886 Trapezia cymodoce (Herbst, 1801) Trapezia tigrina Eydoux and Souleyet, 1842 Imajima (1970); Ooishi (1970); Shigei (1970, as T. dentata) Ooishi (1970, as T. danae) Crabs dredged off the Ogasawara Islands 275 Appendix. (cont.) Trapezia digitalis Latreille, 1828 Trapezia lavopunctata Eydoux and Souleyet, 1842 Trapezia rufopunctata (Herbst, 1799) Trapezia septata Dana, 1852 Superfamily Xanthoidea MacLeay, 1838 Family Xanthidae MacLeay, 1838 Actaea perspinosa Borradaile, 1902 Actaeodes consobrinus (A. Milne-Edwards, 1873) Actaeodes hirsutissimus (Rüuppell, 1830) Actiomera boninensis (Odhner, 1925) Chlorodiella cytherea (Dana, 1852) Imajima (1970); Ooishi (1970); Shigei (1970) Imajima (1970); Ooishi (1970); Shigei (1970); Takeda and Kurata (1977a, b) Shigei (1970, as T. maculata) Ooishi (1970, as T. areolata); Takeda and Miyake (1976, as T. reticulata) Odhner (1925) Odhner (1925); Imajima (1970); Ooishi (1970) Odhner (1925); Takeda and Kurata (1976a) Odhner (1925) Ooishi (1970, as C. laevissima); Takeda and Kurata (1977a, b) Chlorodiella nigra (Forskål, 1775) Stimpson (1858, 1907); Imajima (1970) Cyclodius obscurus (Hombron and Jacquinot, 1846) Stimpson (1858, 1907, as Chlorodius monticulosus); Imajima (1970, as Phymodius monticulosus) Cymo andreossyi (Audouin, 1826) Stimpson (1858, 1907); Ooishi (1970) Cymo melanodactylus Dana, 1852 Stimpson (1858, 1907); Ooishi (1970) Danielea noelensis (Ward, 1934) Takeda (1977) Epiactaea nodulosa (White, 1848) Ooishi (1970); Takeda and Kurata (1976a, b, 1984); Takeda (1977) Etisus electra (Herbst, 1801) Imajima (1970); Takeda and Kurata (1976a, 1977a, 1984) Forestia depressa (White, 1848) Odhner (1925) Gaillardiellus rueppelli (Krauss, 1843) Odhner (1925); Imajima (1970) Glyptocarcinus lophopus Takeda, 1973 Takeda (1973) Hypocolpus haanii (Rathbun, 1909) Takeda and Kurata (1976a, as H. granulatus (de Haan, 1837)) Leptodius davaoensis Ward, 1941 Imajima (1970, as L. leptodon); Asakura et al. (1994b) Leptodius exaratus (H. Milne Edwards, 1834) Stimpson (1858, 1907, as Chlorodius exaratus var. rugosus and C. cupulifer var. nov.); Yoshihara (1901); Terazaki (1904) Leptodius gracilis (Dana, 1852) Terazaki (1904, as Xantho exaratus var. lividus); Ooishi (1970) Leptodius sanguineus (H. Milne Edwards, 1834) Balss (1922); Shigei (1970); Ooishi (1970) Liomera bella (Dana, 1852) Stimpson (1858, 1907); Odhner (1925); Imajima (1970); Ooishi (1970) Liomera caelata (Odhner, 1925) Odhner (1925); Takeda (1977) Liomera monticulosa ( A. Milne-Edwards, 1873) Odhner (1925) Liomera rubra (A. Milne-Edwards, 1865) Odhner (1925) Liomera striolata (Odhner, 1925) Odhner (1925) Liomera venosa (H. Milne Edwards, 1834) Terazaki (1904, as Xantho obtusus) Lybia tessellata (Latreille, 1812) Ooishi (1970); Takeda and Kurata (1984) Macromedaeus distinguendus (de Haan, 1835) Matsuura (1894); Terazaki (1904); Ooishi (1970) Nanocassiope tridentata Davie, 1995 Takeda and Kurata (1977b, 1984, as ?N. granulipes) Neoliomera insularis (Adams and White, 1849) Odhner (1925) Neoliomera striata Buitendijk, 1941 Takeda and Kurata (1984) Palapedia integra (de Haan, 1835) Takeda (1977) Paractaeopsis quadriareolata (Takeda and Miyake, Takeda and Kurata (1977a, b) 1968) Paramedaeus simplex (A. Milne-Edwards, 1873) Takeda and Kurata (1976a, 1977b, 1984) 276 Hironori Komatsu Appendix. (cont.) Paraxanthias notatus (Dana, 1852) Pilodius areolatus (H. Milne Edwards, 1834) Pilodius paumotensis Rathbun, 1907 Pilodius sp. Platypodia semigranosa (Heller, 1861) Platypodia sp. Psaumis cavipes (Dana, 1852) Pseudoliomera helleri (A. Milne-Edwards, 1865) Pseudoliomera lata (Borradaile, 1902) Pseudoliomera paraspeciosa (Ward, 1941) Pseudoliomera speciosa (Dana, 1852) Xanthias cherbonnieri Guinot, 1964 Xanthias gilbertensis Balss, 1938 Xanthias lamarckii (H. Milne Edwards, 1834) Zosimus aeneus (Linnaeus, 1758) Zozymodes cavipes (Dana, 1852) Subsection Thoracotremata Guinot, 1977 Superfamily Cryptochiroidea Paul’son, 1875 Family Cryptochiridae Paul’son, 1875 Cryptochirus coralliodytes Heller, 1861 Imajima (1970); Ooishi (1970) Ooishi (1970) Imajima (1970) Takeda and Kurata (1977b) Ooishi (1970) Takeda and Kurata (1977a) Odhner (1925) Odhner (1925); Ooishi (1970); Takeda and Kurata (1976a, 1977b, 1984) Odhner (1925); Takeda and Kurata (1977b) Ooishi (1970) Yoshihara (1901) Takeda (1977) Takeda and Kurata (1984) Sakai (1939); Shigei (1970); Ooishi (1970); Asakura et al. (1994b) Yoshihara (1901); Balss (1922); Imajima (1970); Shigei (1970) Stimpson (1858, 1907); Sakai (1939); Ooishi (1970) Dacryomaia japonica (Takeda and Tamura, 1981) Lithoscaptus tri (Fize and Serène, 1956) Opecarcinus crescentus (Edmondson, 1925) Takeda and Tamura (1980); Takeda and Tamura (1981, as Favicola rugosa) Takeda and Tamura (1981) Takeda and Tamura (1980) this study Superfamily Grapsidae MacLeay, 1838 Family Gecarcinidae MacLeay, 1838 Discoplax hirtipes (Dana, 1852) Epigrapsus notatus (Heller, 1865) Imajima (1970); Türkay (1974) Türkay (1974) Family Grapsidae MacLeay, 1838 Geograpsus grayi (H. Milne Edwards, 1853) Metopograpsus messor (Forskål, 1775) Metopograpsus thukuhar (Owen, 1839) Pachygrapsus minutus A. Milne-Edwards, 1873 Pachygrapsus plicatus (H. Milne Edwards, 1837) Planes major (MacLeay, 1838) Planes marinus Rathbun, 1914 Stimpson (1858, 1907, as G. rubidus sp. nov.); Balss (1922); Shigei (1970) Ooishi (1970) Ooishi (1970) Stimpson (1858, 1907, as G. rudis); Yoshihara (1901, as G. grapsus); Balss (1922, as G. grapsus); Shigei (1970); Imajima (1970); Ooishi (1970); Takeda and Kurata (1977a) Imajima (1970); Ooishi (1970); Asakura et al. (1994b) Stimpson (1858, 1907); Sakai (1939) Ooishi (1970) Sakai (1939); Imajima (1970); Ooishi (1970) Takeda (1973, as P. cyaneus) Takeda and Kurata (1977a) Family Percnidae Števčić, 2005 Percnon abbreviatum (Dana, 1851) Imajima (1970); Ooishi (1970) Geograpsus stormi de Man, 1895 Grapsus longitarsis Dana, 1851 Grapsus tenuicrustatus (Herbst, 1783) Crabs dredged off the Ogasawara Islands 277 Appendix. (cont.) Percnon planissimum (Herbst, 1804) Family Plagusiidae Dana, 1851 Plagusia squamosa (Herbst, 1790) Family Sesarmidae Dana, 1851 Chiromantes dehaani (H. Milne Edwards, 1853) Family Varunidae H. Milne Edwards, 1853 Cyclograpsus longipes Stimpson, 1858 Eriocheir ogasawaraensis Komai, 2006 Stimpson (1858, 1907); Ooishi (1970) Parisi (1918, as P. tuberculata); Balss (1922, as P. minutus); Sakai (1939, as P. tuberculata); Ooishi (1970, as P. tuberculata) Yoshihara (1901); Parisi (1918); Imajima (1970); Ooishi (1970); Shigei (1970), Asakura et al. (1994b) Ptychognathus glaber Stimpson, 1858 Stimpson (1858, 1907); Takeda and Kurata (1977a) Miyake (1970, as E. japonicus); Kobayashi (2005, as Eriocheir sp.); Komai et al. (2006) Stimpson (1858, 1907); Balss (1922); Ooishi (1970); Asakura et al. (1994b) Parisi (1918, as Helich leachii); Sakai (1939, as H. leachii) Stimpson (1858, 1907); Sakai (1939); Ooishi (1970) Family Xenograpsidae Ng, Davie and Ng, 2007 Xenograpsus novaeinsularis Takeda and Kurata, 1977 Takeda and Kurata (1977a) Gaetice depressus (de Haan, 1835) Pseudohelice subquadrata (Dana, 1851) Superfamily Ocypodoidea Raﬁnesque, 1815 Family Macrophthalmidae Dana, 1851 Macrophthalmus (Macrophthalmus) telescopicus (Owen, Takeda (1977) 1839) Family Ocypodidae Raﬁnesque, 1815 Ocypode ceratophthalmus (Pallas, 1772) Uca (Paraleptuca) lactea (de Haan, 1835) Stimpson (1858, 1907); Matsuura (1894); Balss (1922); Parisi (1918) Yoshihara (1901); Balss (1922); Sakai (1939); Ooishi (1970); Shigei (1970) Shigei (1970) Balss (1922) Parisi (1918, as U. pulchella); Sakai (1939); Imajima (1970); Shigei (1970); Ooishi (1970) (as U. gaimardi) Yoshihara (1901); Parisi (1918) Superfamily Pinnotheroidea de Haan, 1833 Family Pinnotheridae de Haan, 1833 Pinnotheres boninensis Stimpson, 1858 Tetrias ischeri (A. Milne-Edwards, 1867) Stimpson (1858, 1907); Suzuki (1972) Takeda (1977) Ocypode cordimanus Latreille, 1818 Ocypode stimpsoni Ortmann, 1897 Uca (Paraleptuca) annulipes (H. Milne Edwards, 1837) Uca (Paraleptuca) crassipes (White, 1847)

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