SANTA BARBARA MUSEUM OF NATURAL HISTORY Contributions in Science Number 1 Checklist of the Marine Bivalves of the Northeastern Pacific Ocean Eugene V. Coan and Paul H. Scott SANTA BARBARA MUSEUM OF NATURAL HISTORY Contributions in Science Number 1 Checklist of the Marine Bivalves of the Northeastern Pacific Ocean Eugene V. Coan and Paul H. Scott Illustrations by Keri Platner SANTA BARBARA MUSEUM OF NATURAL HISTORY 1997 Checklist of the Marine Bivalves of the Northeastern Pacific Ocean Eugene V. Coan and Paul H. Scott Santa Barbara Museum of Natural History Contributions in Science - Number 1 © 1997 Santa Barbara Museum of Natural History 2559 Puesta del Sol Road Santa Barbara, California 93105-2936 USA http://www.sbnature.org Date of Publication: 24 June 1997 © All rights reserved. This book may not be reproduced in whole or in part for any purpose whatever, without written permission from the publisher (Santa Barbara Museum of Natural History). Printed and bound by Alternative Graphics, Goleta, California Introduction This checklist provides an inventory of marine bivalve mollusk taxa from the Arctic coast of Alaska to central Baja California, including all habitats from the intertidal to the deep-sea. The depth coverage is greater than recently published lists by Turgeon et al. (1988, and in preparation), which is limited to species occurring in under 200 m. Our taxonomic synonymies are based on examination of type material, suites of material in museum collections, as well as the published literature. Footnotes highlight innovations included in the inventory. We provide pertinent literature to these innovations but do not provide references to major changes that have been made as a result of previously published literature. Dermatomya teniconcha Each species in this checklist will be fully treated in our forthcoming book on the marine bivalves of western North America (Coan et al., 1997). List of Bivalve Mollusk Taxa Class Bivalvia Linnaeus, 1758 Subclass Protobranchia Pelseneer, 1889 Order Solemyoida Dall, 1889 Superfamily Solemyoidea H. Adams and A. Adams, 1857 Family Solemyidae J. E. Gray, 1840 Genus Solemya Lamarck, 1818 Subgenus Solemya s.s. Solemya (Solemya) reidi F. R. Bernard, 1980 Subgenus Petrasma Dall, 1908 Solemya (Petrasma) valvulus (Carpenter, 1864) Genus Acharax Dall, 1908 Acharax johnsoni (Dall, 1891) Order Nuculoida Dall, 1889 Superfamily Nucinelloidea Vokes, 1956 Family Nucinellidae Vokes, 1956 Genus Huxleyia A. Adams, 1860 Huxleyia munita (Dall, 1898) Superfamily Nuculoidea J. E. Gray, 1824 Family Nuculidae J. E. Gray, 1824 Genus Nucula Lamarck, 1799 Subgenus Nucula, s.s. Nucula (Nucula) profundorum E. A. Smith, 1885 Subgenus Lamellinucula Schenck, 1944 Nucula (Lamellinucula) carlottensis Dall, 18971 Nucula (Lamellinucula) exigua G. B. Sowerby I, in Broderip and G. B. Sowerby I, 1833 Genus Acila H. Adams and A. Adams, 1858 Subgenus Truncacila Grant and Gale, 1931 Acila (Truncacila) castrensis (Hinds, 1843) 1 Genus Ennucula Iredale, 19312 Ennucula cardara (Dall, 1916) Ennucula linki (Dall, 1916) Ennucula tenuis (Montagu, 1808)3 Superfamily Pristiglomoidea Verrill and Bush, 1898 Family Pristiglomidae Verrill and Bush, 1898 Subfamily Pristiglominae Verrill and Bush, 1898 Genus Pristigloma Dall, 1900 Pristigloma nitens (Jeffreys, 1876) Genus Pseudoglomus Dall, 1898 Pseudoglomus fragilis F. R. Bernard, 1989 Subfamily Setiglominae Schileyko, 1983 Genus Setigloma Schileyko, 1983 Setigloma japonica (E. A. Smith, 1885) Superfamily Nuculanoidea H. Adams and A. Adams, 1858 Family Nuculanidae H. Adams and A. Adams, 1858 Subfamily Nuculaninae H. Adams and A. Adams, 1858 Nuculana (Jupiteria) taphria Genus Nuculana Link, 1807 Subgenus Nuculana, s.s. Nuculana (Nuculana) extenuata (Dall, 1897) Nuculana (Nuculana) hamata (Carpenter, 1864) Nuculana (Nuculana) leonina (Dall, 1896)4 Nuculana (Nuculana) minuta (Müller, 1776) Nuculana (Nuculana) navisa (Dall, 1916)5 Nuculana (Nuculana) pernula (Müller, 1779)6 Subgenus Jupiteria Bellardi, 1875 Nuculana (Jupiteria) cellulita (Dall, 1896) Nuculana (Jupiteria) elenensis (G. B. Sowerby I, in Broderip and G. B. Sowerby I, 1833)7 Nuculana (Jupiteria) penderi (Dall and Bartsch, 1910)8 Nuculana (Jupiteria) pontonia (Dall, 1890) Nuculana (Jupiteria) taphria (Dall, 1896) Subgenus Tenuileda Habe, 1977 Nuculana (Tenuileda) conceptionis (Dall, 1896) Subfamily Bathyspinulinae Coan and Scott, new name herein [1982]9 Genus Bathyspinula Filatova, 1958 Subgenus Bathyspinula, s.s. Bathyspinula (Bathyspinula) calcarella (Dall, 1908) Bathyspinula (Bathyspinula) prolata (E. A. Smith, 1885) Subgenus Acutispinula Filatova and Schileyko, 1984 Bathyspinula (Acutispinula) calcar (Dall, 1908) Subfamily Ledellinae J. A. Allen, 1978 Genus Ledella Verrill and Bush, 1897 Subgenus Ledella, s.s. Ledella (Ledella) fiascona (Dall, 1916)10 Ledella (Ledella) robusta F. R. Bernard, 1989 Ledella (Ledella) sandersi Filatova and Schileyko, 1984 Ledella (Ledella) tamara Gorbunov, 1946 Ledella (Ledella) ultima (E. A. Smith, 1885) 2 Genus Ledellina Filatova and Schileyko, 1984 Ledellina convexirostrata Filatova and Schileyko, 1984 Ledellina formabile Filatova and Schileyko, 1984 Family Malletiidae H. Adams and A. Adams, 1858 Genus Malletia des Moulins, 1832 Malletia alata F. R. Bernard, 1989 Malletia faba Dall, 189711 Malletia pacifica Dall, 1897 Malletia talama Dall, 1916 Malletia truncata Dall, 1908 Genus Katadesmia Dall, 1908 Katadesmia vincula (Dall, 1908) Family Neilonellidae J. A. Allen, 1978 Genus Neilonella Dall, 1881 Neilonella brunnea (Dall, 1916) Neilonella kermadecensis Knudsen, 1970 Neilonella mexicana Dall, 1908 Neilonella ritteri (Dall, 1916) Genus Austrotindaria C. A. Fleming, 1948 Austrotindaria gibbsii (Dall, 1897) Genus Clencharia Clarke, 1961 Clencharia abyssicola (E. A. Smith, 1885) Family Sareptidae Stoliczka, 1871 Subfamily Sareptinae Stoliczka, 1871 Genus Yoldia Möller, 1842 Subgenus Yoldia, s.s. Yoldia (Yoldia) hyperborea (Gould, 1841)12 Yoldia (Yoldia) myalis (Couthouy, 1838)13 Subgenus Cnesterium Dall, 1898 Yoldia (Cnesterium) seminuda Dall, 187114 Subgenus Kalayoldia Grant and Gale, 1931 Yoldia (Kalayoldia) cooperii Gabb, 1865 Genus Megayoldia Verrill and Bush, 1897 Megayoldia martyria (Dall, 1897) Megayoldia montereyensis (Dall, 1893)15 Megayoldia thraciaeformis (Storer, 1838) 16 Genus Portlandia Mörch, 1857 Portlandia aestuariorum (Mosevich, 1928)17 Portlandia arctica (J. E. Gray, 1824) Portlandia intermedia (M. Sars, 1865) Portlandia lenticula (Möller, 1842) Subfamily Yoldiellinae Allen and Hannah, 1986 Genus Yoldiella Verrill and Bush, 1897 Yoldiella capsa (Dall, 1916) Yoldiella derjugini Scarlato, 1981 Yoldiella frigida (Torell, 1859) Yoldiella nana (M. Sars, 1865)18 Yoldiella orcia (Dall, 1916)19 Yoldiella sp. A — Tuffs Abyssal Plain 3 Family Siliculidae Allen and Sanders, 1973 Genus Silicula Jeffreys, 1879 Silicula alleni F. R. Bernard, 1989 Silicula sandersi (F. R. Bernard, 1989) Family Tindariidae Verrill and Bush, 1897 Genus Tindaria Bellardi, 1875 Tindaria compressa Dall, 1908 Tindaria kennerlyi (Dall, 1897) Subclass Pteriomorphia Beurlen, 1944 Order Arcoida Stoliczka, 1871 Superfamily Arcoidea Lamarck, 1809 Family Arcidae Lamarck, 1809 Subfamily Arcinae Lamarck, 1809 Genus Acar J. E. Gray, 1857 Acar bailyi (Bartsch, 1931) Genus Barbatia J. E. Gray, 1842 Subgenus Cucullaearca Conrad, 1865 Barbatia (Cucullaearca) reeveana (d’Orbigny, 1846)20 Subfamily Anadarinae Reinhart, 1935 Genus Anadara J. E. Gray, 1847 Subgenus Anadara, s.s. Anadara (Anadara) tuberculosa (G. B. Sowerby I, 1833) Subgenus Larkinia Reinhart, 1935 Anadara (Larkinia) multicostata (G.B. Sowerby I, 1833) Genus Bathyarca Kobelt, 1891 Bathyarca corpulenta (E. A. Smith, 1885) Bathyarca frielei (Friele, 1877) Bathyarca glacialis (J. E. Gray, 1824) Bathyarca nucleator (Dall, 1908) Genus Bentharca Verrill and Bush, 1898 Bentharca asperula (Dall, 1881) Family Noetiidae Stewart, 1930 Subfamily Striarcinae MacNeil, 1938 Genus Arcopsis Koenen, 1885 Arcopsis solida (G. B. Sowerby I, 1833)21 Family Parallelodontidae Dall, 1898 [1895] Subfamily Grammadontinae Branson, 1942 Genus Porterius B. L. Clark, 1925 Porterius new species A — Aleutian Islands, Alaska22 Superfamily Glycymeridoidea Newton, 1916 Family Glycymerididae Newton, 1916 Subfamily Glycymeridinae Newton, 1916 Genus Glycymeris da Costa, 1778 Subgenus Axinola Hertlein and Grant, 1972 Glycymeris (Axinola) keenae Willett, 1944 Glycymeris (Axinola) septentrionalis (Middendorff, 1849) 4 Superfamily Limopsoidea Dall, 1895 Family Limopsidae Dall, 1895 Genus Limopsis Sassi, 1827 Limopsis akutanica Dall, 1916 Limopsis panamensis Dall, 190223 Limopsis tenella Jeffreys, 1876 Genus Empleconia Dall, 1908 Empleconia vaginata (Dall, 1891)24 Genus Nipponolimopsis Habe, 1951 Nipponolimopsis decussata (A. Adams, 1862)25 Superfamily Philobryoidea F. Bernard, 1897 Family Philobryidae F. Bernard, 1897 Genus Philobrya Cooper, 1867 Philobrya setosa (Carpenter, 1864) Order Mytiloida Férussac, 1822 Superfamily Mytiloidea Rafinesque, 1815 Family Mytilidae Rafinesque, 1815 Subfamily Mytilinae Rafinesque, 1815 Genus Mytilus Linnaeus, 1758 Mytilus californianus Conrad, 1837 Mytilus edulis Linnaeus, 175826 Mytilus galloprovincialis Lamarck, 1819 Mytilus trossulus Gould, 1850 Genus Brachidontes Swainson, 1840 Brachidontes adamsianus (Dunker, 1857) Subfamily Bathymodiolinae Kenk and Wilson, 1985 Genus Benthomodiolus Dell, 1987 Benthomodiolus abyssicola (Knudsen, 1970) Subfamily Crenellinae J. E. Gray, 1840 Genus Crenella Brown, 1827 Crenella decussata (Montagu, 1808)27 Genus Gregariella Monterosato, 1883 Gregariella coarctata (Carpenter, 1857) 28 Genus Musculista Yamamoto and Habe, 1958 Musculista senhousia (Benson, in Cantor, 1842) Crenella decussata Genus Musculus Röding, 1798 Musculus discors (Linnaeus, 1767) Musculus glacialis (Leche, 1883)29 Musculus niger (J. E. Gray, 1824)30 “Musculus” phenax (Dall, 1915)31 “Musculus” pygmaeus Glynn, 196431 “Musculus” taylori (Dall, 1897)31 Genus Solamen Iredale, 192432 Solamen columbianum (Dall, 1897) Solamen leanum (Dall, 1897)33 Solamen megas (Dall, 1902)34 Genus Vilasina Scarlato, 1960 Vilasina seminuda (Dall, 1897)35 Vilasina vernicosa (Middendorff, 1849)35 5 Subfamily Dacryniinae Ockelmann, 1983 Genus Dacrydium Torell, 185936 Subgenus Dacrydium, s.s. Dacrydium (Dacrydium) pacificum Dall, 191637 Dacrydium (Dacrydium) rostriferum F. R. Bernard, 1978 Dacrydium (Dacrydium) vitreum (Möller, 1842) Subfamily Lithophaginae H. Adams and A. Adams, 1857 Genus Adula H. Adams and A. Adams, 1852 Adula californiensis (Philippi, 1847) Adula diegensis (Dall, 1911) Adula gruneri (Philippi, 1851)38 Adula schmidtii (Schrenck, 1867)39 Genus Lithophaga Röding, 1798 Subgenus Diberus Dall, 1898 Lithophaga (Diberus) plumula (Hanley, 1844) Subgenus Labis Dall, 1916 Lithophaga (Labis) attenuata (Deshayes, 1836) Subgenus Myoforceps P.-H. Fischer I, 1886 Lithophaga (Myoforceps) aristata (Dillwyn, 1817) Subfamily Modiolinae Keen, 1958 Modiolus (Modiolus) sacculifer Genus Modiolus Lamarck, 179940 Subgenus Modiolus, s.s. Modiolus (Modiolus) capax (Conrad, 1837) Modiolus (Modiolus) carpenteri Soot-Ryen, 1963 Modiolus (Modiolus) modiolus (Linnaeus, 1758)41 Modiolus (Modiolus) sacculifer (Berry, 1953) Subgenus Modiolusia Yamamoto and Habe, 1958 Modiolus (Modiolusia) neglectus Soot-Ryen, 1955 Modiolus (Modiolusia) rectus (Conrad, 1837)42 Genus Geukensia van de Poel, 195943 Geukensia demissa (Dillwyn, 1817) Genus Amygdalum Megerlie von Mühlfeld, 1811 Amygdalum politum (Verrill and S. Smith, in Verrill, 1880) Genus Idas Jeffreys, 187644 Idas washingtonius (F. R. Bernard, 1978) Subfamily Septiferinae Scarlato and Starobogatov, 1979 Genus Septifer Récluz, 1848 Septifer bifurcata (Conrad, 1837) Order Pterioida Newell, 1965 Suborder Pteriina Newell, 1965 Superfamily Pterioidea J. E. Gray, 1847 [1820] Family Pteriidae J. E. Gray, 1847 [1820] Genus Pteria Scopoli, 1777 Pteria sterna (A. A. Gould, 1851)45 Family Isognomonidae Woodring, 1925 [1815] Genus Isognomon [Lightfoot], 1786 Subgenus Melina Philipsson, 1788 Isognomon (Melina) janus Carpenter, 1857 Isognomon (Melina) recognitus (Mabille, 1895) 6 Suborder Pinnina Waller, 1978 Superfamily Pinnoidea Leach, 1819 Family Pinnidae Leach, 1819 Genus Atrina J. E. Gray, 1842 Subgenus Servatrina Iredale, 1939 Atrina (Servatrina) oldroydii Dall, 1901 Atrina (Servatrina) texta Hertlein, Hanna and Strong, in Hertlein and Strong, 194346 Order Limoida Waller, 1978 Superfamily Limoidea Rafinesque, 1815 Family Limidae Rafinesque, 1815 Genus Limaria Link, 1807 Limaria hemphilli (Hertlein and Strong, 1946) Genus Limatula S. V. Wood, 1839 Subgenus Limatula, s.s.47 Limatula (Limatula) attenuata Dall, 1916 Limatula (Limatula) hyperborea Jensen, 1905 Limatula (Limatula) saturna F. R. Bernard, 1978 Limatula (Limatula) similaris (Dall, 1908)48 Limatula (Limatula) vancouverensis F. R. Bernard, 1978 Subgenus Limatuletta C. A. Fleming, 1978 Limatula (Limatuletta) coani F. R. Bernard, 1988 Limatula (Limatuletta) japonica (A. Adams, 1863)49 Genus Acesta H. Adams and A. Adams, 1858 Subgenus Acesta, s.s. Acesta (Acesta) mori (Hertlein, 1952) Subgenus Plicacesta Vokes, 1963 Acesta (Plicacesta) sphoni (Hertlein, 1963) Order Ostreoida Férussac, 1822 Suborder Ostreina Férussac, 1822 Superfamily Ostreoidea Rafinesque, 1815 Family Ostreidae Rafinesque, 1815 Subfamily Ostreinae Rafinesque, 1815 Genus Ostrea Linnaeus, 1758 Ostrea conchaphila Carpenter, 185750 Ostrea edulis Linnaeus, 1758 Subfamily Crassostreinae Scarlato and Starobogatov, 1979 Genus Crassostrea Sacco, 1897 Crassostrea ariakensis (Fujita, 1913) Crassostrea gigas (Thunberg, 1793) Crassostrea sikamea (Amemiya, 1928) Crassostrea virginica (Gmelin, 1791) Superfamily Dimyoidea P.-H. Fischer I, 1886 Family Dimyidae P.-H. Fischer I, 1886 Genus Dimya Rouault, 1850 Dimya californiana Berry, 193651 7 Suborder Pectinina Waller, 1978 Superfamily Pectinoidea Rafinesque, 1815 Family Pectinidae Rafinesque, 1815 Subfamily Pectininae Rafinesque, 1815 Genus Euvola Dall, 1898 Euvola diegensis (Dall, 1898) Euvola vogdesi (Arnold, 1906)52 Subfamily Camptonectinae Habe, 1977 Genus Delectopecten Stewart, 1930 Delectopecten vancouverensis (Whiteaves, 1893)53 Genus Hyalopecten Verrill, 1897 Hyalopecten neoceanus (Dall, 1908) Subfamily Chlamydinae von Treppner, 1922 Tribe Chlamydinini von Treppner, 1922 Genus Chlamys Röding, 1798 Chlamys albida (Arnold, 1906) Chlamys behringiana (Middendorff, 1849)54 Chlamys hastata (G. B. Sowerby II, 1843) Chlamys rubida (Hinds, 1845)55 Tribe Aequipectini Nordsieck, 1969 Genus Argopecten Monterosato, 1889 Argopecten ventricosus (G. B. Sowerby II, 1842) Genus Leptopecten Verrill, 1897 Leptopecten latiauratus (Conrad, 1837) Tribe Crassadomini Waller, 1993 Genus Crassadoma F. R. Bernard, 1986 Crassadoma gigantea (J. E. Gray, 1825) Tribe Fortipectinini Masuda, 1963 Genus Mizuhopecten Masuda, 1963 Mizuhopecten yessoensis (Jay, 1856) Genus Patinopecten Dall, 1898 Patinopecten caurinus (Gould, 1850) Tribe Mimachlamydini Waller, 1993 Genus Spathochlamys Waller, 1993 Spathochlamys vestalis (Reeve, 1853) Tribe [uncertain] Genus Nodipecten Dall, 1898 Nodipecten subnodosus (G. B. Sowerby I, 1835) Family Propeamussidae R. T. Abbott, 1954 Genus Propeamussium de Gregorio, 1884 Propeamussium meridionale (E. A. Smith, 1885) Genus Cyclopecten Verrill, 1897 Cyclopecten barbarensis Grau, 1959 Cyclopecten benthalis Grau, 1959 Cyclopecten bistriatus (Dall, 1916) Cyclopecten carlottensis F. R. Bernard, 1968 Cyclopecten catalinensis (Willett, 1931) Cyclopecten davidsoni (Dall, 1897)56 8 Cyclopecten greenlandicus (G. B. Sowerby II, 1842)57 Cyclopecten incongruus (Dall, 1916) Cyclopecten knudseni F. R. Bernard, 1978 Cyclopecten squamiformis F. R. Bernard, 1978 Cyclopecten zephyrus Grau, 1959 Genus Parvamussium Sacco, 1897 Parvamussium alaskense (Dall, 1871) Superfamily Anomioidea Rafinesque, 1815 Family Ananomiidae Rafinesque, 1815 Genus Anomia Linnaeus, 1758 Subgenus Anomia, s.s. Anomia (Anomia) peruviana d’Orbigny, 1846 Genus Pododesmus Philippi, 1837 Subgenus Pododesmus, s.s. Podesmus (Pododesmus) pernoides (J. E. Gray, 1853) Subgenus Monia J. E. Gray, 1850. Pododesmus (Monia) macrochisma (Deshayes, 1839)58 Subclass Heterodonta Neumayer, 1884 Order Veneroida H. Adams and A. Adams, 1856 Superfamily Lucinoidea J. Fleming, 1828 Family Lucinidae J. Fleming, 1828 Subfamily Lucininae J. Fleming, 1828 Genus Epilucina Dall, 1901 Epilucina californica (Conrad, 1837) Genus Here Gabb, 1866 Here excavata (Carpenter, 1857) Genus Lucinisca Dall, 190159 Lucinisca nuttalli (Conrad, 1837) Genus Parvilucina Dall, 1901 Parvilucina approximata (Dall, 1901) Parvilucina tenuisculpta (Carpenter, 1864) Subfamily Myrteinae Chavan, 1969 Genus Lucinoma Dall, 1901 Lucinoma aequizonatum (Stearns, 1890) Lucinoma annulatum (Reeve, 1850) Family Ungulinidae H. Adams and A. Adams, 1856 Genus Diplodonta Bronn, 1831 Diplodonta aleutica Dall, 1901 Diplodonta impolita Berry, 1953 Diplodonta orbella (Gould, 1851) Diplodonta sericata (Reeve, 1850)60 Conchocele bisecta Family Thyasiridae Dall, 1900 [1895] Subfamily Thyasirinae Dall, 1900 Genus Thyasira Lamarck, 1818 Thyasira flexuosa (Montagu, 1803)61 Thyasira orecta F. R. Bernard, 1982 9 Genus Conchocele Gabb, 1866 Conchocele bisecta (Conrad, 1849)62 Conchocele excavata (Dall, 1901) Subfamily Axinopsidinae F. R. Bernard, 1983 Genus Axinopsida Keen and Chavan, in Chavan, 1951 Axinopsida serricata (Carpenter, 1864)63 Genus Adontorhina Berry, 1947 Adontorhina cyclia Berry, 1947 Adontorhina sphaeriosa Scott, 1986 Adontorhina new species — California Genus Axinodon Verrill and Bush, 1898 Axinodon redondoensis (T. A. Burch, 1941)64 Genus Leptaxinus Verrill and Bush, 1898 Leptaxinus minutus Verrill and Bush, 189865 Genus Mendicula Iredale, 1924 Mendicula ferruginosa (Forbes, 1844)66 Superfamily Astartoidea d’Orbigny, 1844 Family Astartidae d’Orbigny, 1844 Subfamily Astartinae d’Orbigny, 1844 Genus Astarte J. Sowerby, 181667 Astarte arctica (J. E. Gray, 1824)68 Astarte borealis (Schumacher, 1817) Astarte compacta Carpenter, 186469 Astarte crenata (J. E. Gray, 1824)70 Astarte elliptica (Brown, 1827)71 Astarte esquimalti (Baird, 1863)72 Astarte montagui (Dillwyn, 1817) Astarte vernicosa (Dall, 1903) 73 Superfamily Crassatelloidea Férussac, 1822 Family Crassatellidae Férussac, 1822 Subfamily Crassatellinae Férussac, 1822 Genus Eucrassatella Iredale, 1924 Eucrassatella fluctuata (Carpenter, 1864) Subfamily Scambulinae Chavan, 1952 Genus Crassinella Guppy, 1874 Crassinella pacifica (C. B. Adams, 1852) Superfamily Carditoidea J. Fleming, 1828 Family Carditidae J. Fleming, 1828 Subfamily Carditamerinae Chavan, 1969 Genus Cyclocardia Conrad, 1867 Subgenus Cyclocardia, s.s. Cyclocardia (Cyclocardia) bailyi (J. Q. Burch, 1944) Cyclocardia (Cyclocardia) barbarensis (Stearns, 1890) Cyclocardia (Cyclocardia) crebricostata (Krause, 1885) Cyclocardia (Cyclocardia) gouldii (Dall, 1903)74 Cyclocardia (Cyclocardia) incisa (Dall, 1903) Cyclocardia (Cyclocardia) ovata (Riabinina, 1952)75 Cyclocardia (Cyclocardia) umnaka (Willett, 1932) Cyclocardia (Cyclocardia) ventricosa (Gould, 1850) 10 Subgenus Crassicardia Savitskii, 197976 Cyclocardia (Crassicardia) crassidens (Broderip and G. B. Sowerby I, 1829) Genus Glans Megerle von Mühlfeld, 1811 Glans carpenteri (Lamy, 1922) Genus Miodontiscus Dall, 1903 Miodontiscus prolongatus (Carpenter, 1864) Subfamily Thecaliinae Dall, 1903 Glans carpenteri Genus Milneria Dall, 1881 Milneria kelseyi Dall, 1916 Milneria minima (Dall, 1871) Superfamily Chamoidea Lamarck, 1809 Family Chamidae Lamarck, 1809 Genus Chama Linnaeus, 1758 Chama arcana F. R. Bernard, 1976 Genus Pseudochama Odhner, 1917 Pseudochama exogyra (Conrad, 1837) Pseudochama granti Strong, 1934 Superfamily Galeommatoidea J. E. Gray, 1840 Family Galeommatidae J. E. Gray, 1840 Subfamily Galeommatinae J. E. Gray, 1840 Genus Galeomma Turton, 1825 "Galeomma" new species — British Columbia to southern California Genus Scintillona Finlay, 1927 Scintillona bellerophon Ó Foighil and Gibson, 1984. Subfamily Chlamydoconchinae Dall, 1884 Genus Chlamydoconcha Dall, 1884 Chlamydoconcha orcutti Dall, 1884 Family Lasaeidae J. E. Gray, 184277 Genus Lasaea Brown, 1827 Lasaea adansoni (Gmelin, 1791) Genus Boreacola F. R. Bernard, 1979 Boreacola dawsoni (Jeffreys, 1864)78 Genus Cymatioa Berry, 1964 Cymatioa electilis (Berry, 1963)79 Genus Isorobitella Keen, 1962 Isorobitella trigonalis (Carpenter, 1857) Genus Kellia Turton, 1822 Kellia suborbicularis (Montagu, 1803)80 Genus Mysella Angas, 1877 Mysella pedroana Dall, 1899 Mysella planata (Krause, 1885)81 Genus Neaeromya Gabb, 1872 Neaeromya compressa (Dall, 1899)82 Neaeromya rugifera (Carpenter, 1864)83 11 Genus Orobitella Dall, 1900 Orobitella californica (Dall, 1899)84 Genus Rhamphidonta F. R. Bernard, 1975 Rhamphidonta retifera (Dall, 1899) Genus Pristes Carpenter, 1864 Pristes oblongus Carpenter, 1864 Genus Rochefortia Velain, 187785 Rochefortia compressa (Dall, 1913) Rochefortia grippi (Dall, 1912) Rochefortia tumida (Carpenter, 1864)86 Rochefortia new species A — California Rochefortia new species B — California Rochefortia new species B Superfamily Articoidea Newton, 1891 Family Trapeziidae Lamy, 1920 Genus Neotrapezium Habe, 1951 Neotrapezium liratum (Reeve, 1843) Superfamily Glossoidea J. E. Gray, 1847 Family Kelliellidae P.-H. Fischer I, 1887 Genus Kelliella M. Sars, 1870 Kelliella galathea Knudsen, 1970 Kelliella pacifica (E. A. Smith, 1885) Family Vesicomyidae Dall and Simpson, 1901 Genus Vesicomya Dall, 1886 Subgenus Vesicomya, s.s. Vesicomya (Vesicomya) lepta (Dall, 1896) Vesicomya (Vesicomya) stearnsii (Dall, 1895) Subgenus Calyptogena Dall, 189187 Vesicomya (Calyptogena) elongata (Dall, 1916) Vesicomya (Calyptogena) extenta Krylova and Moskalev, 1996 Vesicomya (Calyptogena) gigas (Dall, 1896) Vesicomya (Calyptogena) kilmeri (F. R. Bernard, 1974) Vesicomya (Calyptogena) pacifica (Dall, 1891) Vesicomya (Calyptogena) new species A — Oregon Subduction Vesicomya (Calyptogena) new species B — Monterey shallow Superfamily Cymatioidea G. O. Sars, 1878 Family Bernardinidae Keen, 1969 Genus Bernardina Dall, 1910 Bernardina bakeri Dall, 1910 Genus Halodakra Olsson, 1961 Subgenus Halodakra, s.s. Halodakra (Halodakra) subtrigona (Carpenter, 1857) Subgenus Stohleria Coan, 1984 Halodakra (Stohleria) salmonea (Carpenter, 1864) 12 Superfamily Corbiculoidea J. E. Gray, 1847 Family Corbiculidae E J. E. Gray, 1847 Genus Corbicula Megerle von Mühlfeld, 1811 Corbicula fluminea (Müller, 1774) Superfamily Cardioidea Lamarck, 1809 Family Cardiidae Lamarck, 1809 Subfamily Clinocardiinae Kafanov, 1975 Genus Clinocardium Keen, 1936 Subgenus Clinocardium, s.s. Clinocardium (Clinocardium) nuttallii (Conrad, 1837) Subgenus Ciliatocardium Kafanov, 1974 Clinocardium (Ciliatocardium) ciliatum (Fabricius, 1780) Subgenus Keenocardium Kafanov, 1974 Clinocardium (Keenocardium) blandum (Gould, 1850)88 Clinocardium (Keenocardium) californiense (Deshayes, 1839) Genus Serripes Gould, 1841 Subgenus Serripes, s.s. Serripes (Serripes) groenlandicus (Mohr, 1786)89 Serripes (Serripes) laperousii (Deshayes, 1839)90 Subgenus Yagudinella Kafanov, 1975 Serripes (Yagudinella) notabilis (G. B. Sowerby III, 1915)91 Subfamily Fraginae Stewart, 1930 Genus Americardia Stewart, 1930 Americardia biangulata (Broderip and G. B. Sowerby I, 1829) Subfamily Laevicardiinae Keen, 1936 Genus Laevicardium Swainson, 1840 Laevicardium elatum (G. B. Sowerby I, in Broderip and G. B. Sowerby I, 1833) Laevicardium substriatum (Conrad, 1837) Genus Nemocardium Meek, 1876 Subgenus Keenae Habe, 1952 Nemocardium (Keenae) centifilosum (Carpenter, 1864) Subfamily Trachycardiinae Stewart, 1930 Genus Trachycardium Mörch, 1853 Subgenus Dallocardia Stewart, 1930 Trachycardium (Dallocardia) quadragenarium (Conrad, 1837) Superfamily Veneroidea Rafinesque, 1815 Family Veneridae Rafinesque, 1815 Subfamily Venerinae Rafinesque, 1815 Genus Chione Megerle von Mühlfeld, 1811 Subgenus Chione, s.s. Chione (Chione) californiensis (Broderip, in Broderip and G. B. Sowerby I, 1835) Chione (Chione) undatella (G. B. Sowerby I, 1835) Subgenus Chionista Keen, 1958 Chione (Chionista) fluctifraga (G. B. Sowerby II, 1853) Genus Globivenus Coen, 193492 Globivenus fordii (Yates, 1890) 13 Genus Humilaria Grant and Gale, 1931 Humilaria kennerleyi (Reeve, 1863) Genus Irusella Hertlein and Grant, 1972 Irusella lamellifera (Conrad, 1837) Genus Mercenaria Schumacher, 1817 Mercenaria mercenaria (Linnaeus, 1758) Genus Protothaca Dall, 1902 Subgenus Protothaca, s.s. Protothaca (Protothaca) laciniata (Carpenter, 1864) Protothaca (Protothaca) staminea (Conrad, 1837) Subgenus Callithaca Dall, 1902 Protothaca (Callithaca) tenerrima (Carpenter, in Gould and Carpenter, 1857) Subfamily Clementiinae Frizzell, 1936 Genus Compsomyax Stewart, 1930 Compsomyax subdiaphana (Carpenter, 1864) Subfamily Gemminae Dall, 1902 Genus Gemma Deshayes, 1853 Gemma gemma (Totten, 1834) Subfamily Meretricinae J. E. Gray, 1847 Genus Tivela Link, 1807 Subgenus Pachydesma Conrad, 1854 Tivela (Pachydesma) stultorum (Mawe, 1823) Subfamily Pitarinae Stewart, 1930 Genus Pitar Römer, 1857 Subgenus Pitar, s.s. Pitar (Pitar) newcombianus (Gabb, 1865) Genus Amiantis Carpenter, 1864 Amiantis callosa (Conrad, 1837) Genus Nutricola F. R. Bernard, 198293 Nutricola confusa (S. Gray, 1982) Nutricola cymata (Dall, 1913)94 Nutricola lordi (Baird, 1863) Nutricola ovalis (Dall, 1902) Nutricola tantilla (Gould, 1853) Genus Saxidomus Conrad, 1837 Saxidomus gigantea (Deshayes, 1839) Saxidomus nuttalli Conrad, 1837 Subfamily Tapetinae J. E. Gray, 1851 Genus Liocyma Dall, 1870 Liocyma fluctuosa (Gould, 1841)95 Genus Venerupis Lamarck, 1818 Subgenus Ruditapes Chiamenti, 1900 Venerupis (Ruditapes) philippinarum (A. Adams and Reeve, 1850) Family Petricolidae d’Orbigny, 1840 Genus Petricola Lamarck, 1801 Subgenus Petricola, s.s. Petricola (Petricola) carditoides (Conrad, 1837)96 14 Subgenus Petricolirus Habe, 1951 Petricola (Petricolirus) californiensis (Pilsbry and Lowe, 1932) Subgenus Petricola, s.l. "Petricola tellimyalis (Carpenter, 1864)," of authors, new species97 Genus Cooperella Carpenter, 1864 Cooperella subdiaphana (Carpenter, 1864) Genus Petricolaria Stoliczka, 187098 Petricolaria pholadiformis (Lamarck, 1818) Family Turtoniidae W. Clark, 1855 Genus Turtonia Alder, 1848 Turtonia minuta (Fabricius, 1780)99 Superfamily Tellinoidea Blainville, 1814 Family Tellinidae Blainville, 1814 Subfamily Tellininae Blainville, 1814 Tellina (Tellinella) idae Genus Tellina Linnaeus, 1758 Subgenus Angulus Megerle von Mühlfeld, 1811 Tellina (Angulus) carpenteri Dall, 1900 Tellina (Angulus) meropsis Dall, 1900 Tellina (Angulus) modesta (Carpenter, 1864) Tellina (Angulus) new species — southern California Subgenus Cadella Dall, Bartsch and Rehder, 1939 Tellina (Cadella) nuculoides (Reeve, 1854) Subgenus Megangulus Afshar, 1969 Tellina (Megangulus) lutea W. Wood, 1828 Subgenus Peronidia Dall, 1900 Tellina (Peronidia) bodegensis Hinds, 1845 Subgenus Tellinella Mörch, 1853 Tellina (Tellinella) idae Dall, 1891 Subfamily Macominae Olsson, 1961 Genus Macoma Leach, 1819 Subgenus Macoma, s.s. Macoma (Macoma) brota Dall, 1916 Macoma (Macoma) calcarea (Gmelin, 1791) Macoma (Macoma) crassula (Deshayes, 1855)100 Macoma (Macoma) elimata Dunnill and Coan, 1968 Macoma (Macoma) lama Bartsch, 1929 Macoma (Macoma) lipara Dall, 1916 Macoma (Macoma) loveni (Jensen, 1905) Macoma (Macoma) middendorffi Dall, 1884 Macoma (Macoma) moesta (Deshayes, 1855) Macoma (Macoma) obliqua (J. Sowerby, 1817)101 Macoma (Macoma) torelli (Jensen, 1905)102 Subgenus Psammacoma Dall, 1900 Macoma (Psammacoma) acolasta Dall, 1921 Macoma (Psammacoma) carlottensis Whiteaves, 1880103 Macoma (Psammacoma) yoldiformis Carpenter, 1864 Subgenus Rexithaerus Tryon, 1869 Macoma (Rexithaerus) dexioptera Baxter, 1977 Macoma (Rexithaerus) expansa Carpenter, 1864 Macoma (Rexithaerus) indentata Carpenter, 1864 Macoma (Rexithaerus) secta (Conrad, 1837) 15 Subgenus Macoma, s.l. Macoma balthica (Linnaeus, 1758) Macoma inquinata (Deshayes, 1855) Macoma nasuta (Conrad, 1837) Genus Leporimetis Iredale, 1930 Leporimetis obesa (Deshayes, 1855) Family Donacidae J. Fleming, 1828 Genus Donax Linnaeus, 1758 Subgenus Paradonax Cossman, in Cossmann and Peyrot, 1910 Donax (Paradonax) californicus Conrad, 1837 Subgenus Donax, s.l. Donax gouldii Dall, 1921 Family Psammobiidae J. Fleming, 1828 Genus Gari Schumacher, 1817 Subgenus Gobraeus Brown, 1844 Gari (Gobraeus) californica (Conrad, 1849) Gari (Gobraeus) fucata (Hinds, 1845) Gari (Gobraeus) regularis (Carpenter, 1864) Genus Heterodonax Mörch, 1853 Heterodonax pacificus (Conrad, 1837) Genus Nuttallia Dall, 1900 Nuttallia nuttallii (Conrad, 1837) Nuttallia obscurata (Reeve, 1857) Family Semelidae Stoliczka, 1870 [1850] Genus Semele Schumacher, 1817 Subgenus Semele, s.s. Semele (Semele) decisa (Conrad, 1837) Semele (Semele) rubropicta Dall, 1871 Subgenus Amphidesma Lamarck, 1818 Semele (Amphidesma) venusta (Reeve, 1853) Subgenus Elegantula de Gregorio, 1884 Semele (Elegantula) rupicola Dall, 1915 Subgenus Semele, s.l. Semele pulchra (G. B. Sowerby I, 1832) Genus Abra Lamarck, 1818 Subgenus Abra, s.s. Abra (Abra) californica Knudsen, 1970 Abra (Abra) profundorum (E. A. Smith, 1885) Genus Abrina Habe, 1952 Abrina species — Oregon Genus Cumingia G. B. Sowerby I, 1833 Cumingia californica Conrad, 1837 Genus Theora H. Adams and A. Adams, 1852 Subgenus Endopleura A. Adams, 1864 Theora (Endopleura) lubrica Gould, 1861 16 Family Solecurtidae d’Orbigny, 1846 Genus Solecurtus Blainville, 1824 Solecurtus guaymasensis (Lowe, 1935)104 Genus Tagelus J. E. Gray, 1847 Subgenus Tagelus, s.s. Tagelus (Tagelus) affinis (G. B. Sowerby II, 1874) Tagelus (Tagelus) californianus (Conrad, 1837) Subgenus Mesopleura Conrad, 1868 Tagelus (Mesopleura) subteres (Conrad, 1837) Superfamily Solenoidea Lamarck, 1809 Family Solenidae Lamarck, 1809 Genus Solen Linnaeus, 1758 Subgenus Ensisolen Habe, 1977 Solen (Ensisolen) rostiformis Dunker, 1862105 Solen (Ensisolen) sicarius Gould, 1850 Family Pharidae H. Adams and A. Adams, 1858 Genus Ensis Schumacher, 1817 Ensis myrae Berry, 1953 Genus Siliqua Megerle von Mühlfeld, 1811 Siliqua alta (Broderip and G. B. Sowerby I, 1829)106 Siliqua lucida (Conrad, 1837) Siliqua patula (Dixon, 1789) Superfamily Mactroidea Lamarck, 1809 Family Mactridae Lamarck, 1809 Subfamily Mactrinae Lamarck, 1809 Genus Mactromeris Conrad, 1868 Mactromeris catilliformis (Conrad, 1867)107 Mactromeris hemphilli (Dall, 1894)107 Mactromeris polynyma (Stimpson, 1860) Mactromeris hemphilli Genus Mactrotoma Dall, 1894 Subgenus Mactrotoma, s.s. Mactrotoma (Mactrotoma) nasuta (Gould, 1851)108 Subgenus Micromactra Dall, 1894 Mactrotoma (Micromactra) californica (Conrad, 1837)108 Genus Simomactra Dall, 1894109 Simomactra falcata ( Gould, 1850)110 Simomactra planulata (Conrad, 1837)110 Subfamily Lutraniinae H. Adams and A. Adams, 1856 Genus Tresus J. E. Gray, 1853 Tresus capax (Gould, 1850) Tresus nuttallii (Conrad, 1837) "Tresus pajaroanus (Conrad, 1857),"of authors, new species111 Subfamily Pteropsellinae Keen, 1969 [1894] Genus Raeta J. E. Gray, 1853 Subgenus Raeta, s.s. Raeta (Raeta) undulata (Gould, 1851) 17 Subclass Asthenodonta Dall, 1895 Order Myoida Goldfuss, 1820 Suborder Myina Goldfuss, 1820 Superfamily Myoidea Lamarck, 1809 Family Myidae Lamarck, 1809 Subfamily Myinae Lamarck, 1809 Genus Mya Linnaeus, 1758 Mya arenaria Linnaeus, 1758 Mya baxteri Coan and Scott, new name herein for Mya intermedia Dall, 1898, non J. Sowerby, 1814112 Mya pseudoarenaria Schlesch, 1931113 Mya truncata Linnaeus, 1758 Genus Platyodon Conrad, 1837 Platyodon cancellatus (Conrad, 1837) Subfamily Cryptomyinae Habe, 1977 Platyodon cancellatus Genus Cryptomya Conrad, 1849 Cryptomya californica (Conrad, 1837) Subfamily Spheniinae F. R. Bernard, 1983 Genus Sphenia Turton, 1822 Sphenia luticola (Valenciennes, 1846)114 Family Corbulidae Lamarck, 1818 Subfamily Corbulinae Lamarck, 1818 Genus Caryocorbula Gardner, 1926115 Caryocorbula porcella (Dall, 1916) Genus Juliacorbula Olsson and Harbison, 1953115 Juliacorbula luteola (Carpenter, 1864) Subfamily Potamocorbulinae Habe, 1977 Genus Potamocorbula Habe, 1955 Potamocorbula amurensis (Schrenck, 1861) Family Spheniopsidae Gardner, 1928 Genus Grippina Dall, 1912 Grippina californica Dall, 1912 Superfamily Hiatelloidea J. E. Gray, 1824 Family Hiatellidae J. E. Gray, 1824 Subfamily Hiatellinae J. E. Gray, 1824 Genus Hiatella Bosc, 1801 Hiatella arctica (Linnaeus, 1767)116 Genus Cyrtodaria Reuss, 1801 Cyrtodaria kurriana Dunker, 1861 Genus Panomya J. E. Gray, 1857 Panomya ampla Dall, 1898 Panomya norvegica (Spengler, 1793) Panomya priapus (Tilesius, 1822)117 Genus Panopea Ménard de la Groye, 1807 Panopea abrupta (Conrad, 1849) 18 Subfamily Saxicavellinae Scott, 1994 Genus Saxicavella P.-H. Fischer I, 1878 Saxicavella nybakkeni Scott, 1994 Saxicavella pacifica Dall, 1916 Superfamily Gastrochaenoidea J. E. Gray, 1840 Family Gastrochaenidae J. E. Gray, 1840 Genus Gastrochaena Spengler, 1783 Subgenus Gastrochaena, s.s. Gastrochaena (Gastrochaena) ovata G. B. Sowerby I, 1834 Suborder Pholadina H. Adams and A. Adams, 1858 Superfamily Pholadoidea Lamarck, 1809 Family Pholadidea Lamarck, 1809 Subfamily Pholadinae Lamarck, 1809 Genus Barnea Risso, 1826 Subgenus Anchomasa Leach, 1852 Barnea (Anchomasa) subtruncata (G. B. Sowerby I, 1834) Genus Zirfaea J. E. Gray, 1842 Zirfaea pilsbryi Lowe, 1931 Subfamily Jouannetiinae Tryon, 1862 Genus Netastoma Carpenter, 1864 Netastoma japonicum (Yokoyama, 1920) Netastoma rostratum (Valenciennes, 1846) Subfamily Martesiinae Grant and Gale, 1931 Genus Chaceia Turner, 1955 Chaceia ovoidea (Gould, 1851) Genus Parapholas Conrad, 1848 Parapholas californica (Conrad, 1837) Genus Penitella Valenciennes, 1846 Penitella conradi Valenciennes, 1846 Penitella fitchi Turner, 1955 Penitella hopkinsi Kennedy and Armentrout, 1989 Penitella penita (Conrad, 1837) Penitella richardsoni Kennedy, 1989 Penitella turnerae Evans and Fisher, 1966 Subfamily Xylophagainae Purchon, 1941 Genus Xylophaga Turton, 1822 Xylophaga washingtona Bartsch, 1921 Genus Xyloredo Turner, 1972 Xyloredo naceli Turner, 1972 Family Teredinidae Rafinesque, 1815 Subfamily Teredininae Rafinesque, 1815 Genus Teredo Linnaeus, 1758 Subgenus Teredo, s.s. Teredo (Teredo) bartschi Clapp, 1923118 Teredo (Teredo) navalis Linnaeus, 1758 19 Genus Lyrodus Gould, 1870 Lyrodus pedicellatus (de Quatrefages, 1849) Lyrodus takanoshimensis (Roch, in Roch and Moll, 1929) Subfamily Bankiinae Turner, 1966 Genus Bankia J. E. Gray, 1842 Bankia carinata (J. E. Gray, 1827)119 Bankia setacea (Tryon, 1863) Subclass Anomalodesmata Dall, 1889 Order Pholadomyoida Newell, 1965 Superfamily Pandoroidea Rafinesque, 1815 Family Pandoridae Rafinesque, 1815 Genus Pandora Hwass, 1795 Subgenus Heteroclidus Dall, 1915 Pandora (Heteroclidus) punctata Conrad, 1837 Subgenus Pandorella Conrad, 1863 Pandora (Pandorella) bilirata Conrad, 1855 Pandora (Pandorella) filosa (Carpenter, 1864) Pandora (Pandorella) glacialis Leach, 1819 Pandora (Pandorella) wardiana A. Adams, 1860 Family Lyonsiidae P.-H. Fischer I, 1887 Genus Lyonsia Turton, 1822 Lyonsia arenosa (Möller, 1842) Lyonsia bracteata (Gould, 1850)120 Lyonsia californica Conrad, 1837121 Genus Entodesma Philippi, 1845 Entodesma navicula (A. Adams and Reeve, 1850)122 Entodesma pictum (G. B. Sowerby I, 1834) Genus Mytilimeria Conrad, 1837 Mytilimeria nuttalli Conrad, 1837 Superfamily Thracioidea Stoliczka, 1870 [1830] Family Thraciidae Stoliczka, 1870 [1830] Genus Thracia Blainville, 1824 Subgenus Cetothrax Iredale, 1949 Thracia (Cetothrax) condoni Dall, 1909 Subgenus Crassithracia Soot-Ryen, 1941 Thracia (Crassithracia) septentrionalis Jeffreys, 1872 Subgenus Homoeodesma P.-H. Fischer I, 1887 Thracia (Homoeodesma) challisiana Dall, 1915 Thracia (Homoeodesma) trapezoides Conrad, 1849 Subgenus Ixartia Leach, 1852 Thracia (Ixartia) curta Conrad, 1837 Subgenus Thracia, s.l. Thracia devexa G. O. Sars, 1878 Thracia myopsis Möller, 1842 Genus Asthenothaerus Carpenter, 1864 Asthenothaerus diegensis (Dall, 1915) Genus Cyathodonta Conrad, 1849 Cyathodonta pedroana Dall, 1915 Genus Lampeia MacGinitie, 1959 Lampeia adamsi MacGinitie, 1959 20 Family Periplomatidae Dall, 1895 Genus Periploma Schumacher, 1817 Subgenus Periploma, s.s. Periploma (Periploma) planiusculum G. B. Sowerby I, 1834 Subgenus Cricoploma F. R. Bernard, 1989 Periploma (Cricoploma) discus Stearns, 1890 Subgenus Septentrioploma F. R. Bernard, 1989 Periploma (Septentrioploma) aleuticum (Krause, 1885) Periploma (Septentrioploma) rosewateri F. R. Bernard, 1989 Genus Halistrepta Dall, 1904 Halistrepta sulcata (Dall, 1904) Order “Septibranchida Pelseneer, 1888” Superfamily Cuspidarioidea Dall, 1886 Family Cuspidariidae Dall, 1886 Genus Cuspidaria Nardo, 1840 Cuspidaria apodema Dall, 1916 Cuspidaria buccina F. R. Bernard, 1989 Cuspidara cowani F. R. Bernard, 1967 Cuspidara glacialis (G. O. Sars, 1878) Cuspidaria parapodema F. R. Bernard, 1969 Cuspidaria variola F. R. Bernard, 1979 Genus Austroneaera Powell, 1937 Austroneaera semipellucida (Kuroda, 1948)123 Genus Bathyneaera Scarlato and Starobogatov, 1983 Bathyneaera disa (F. R. Bernard, 1989) Bathyneaera tillamookensis (Dall, 1916) Genus Cardiomya A. Adams, 1864124 Cardiomya balboae Dall, 1916 Cardiomya behringensis (Leche, 1883)125 Cardiomya pectinata (Carpenter, 1864)126 Cardiomya planetica (Dall, 1908) Cuspidaria parapodema Genus Luzonia Dall and E. A. Smith, 1890 Luzonia walleri (F. R. Bernard, 1989)127 Genus Myonera Dall, 1886 Myonera mexicana Knudsen, 1970 Myonera paucistriata (Dall, 1886) Genus Plectodon Carpenter, 1864 Plectodon scaber Carpenter, 1864 Genus Rhinoclama Dall and E. A. Smith, 1886 Rhinoclama filatovae (F. R. Bernard, 1979) Superfamily Verticordioidea Stoliczka, 1871 Family Verticordiidae Stoliczka, 1871 Subfamily Verticordiinae Stoliczka, 1871 Genus Halicardia Dall, 1895 Halicardia perplicata (Dall, 1890) Genus Haliris Dall, 1886128 Haliris aequacostata (Howard, 1950) Genus Trigonulina d’Orbigny, 1842 Trigonulina pacifica Jung, 1996 21 Subfamily Lyonsiellinae Dall, 1895 Genus Lyonsiella G. O. Sars, 1872 Lyonsiella quaylei F. R. Bernard, 1969 Genus Dallicordia Scarlato and Starobogatov, 1983 Dallicordia alaskana (Dall, 1895) Genus Policordia Dall, Bartsch and Rehder, 1938 Policordia jeffreysi (Friele, 1879) Policordia pilula (Pelseneer, 1911) Policordia subrotundata Ivanova, 1977 Superfamily Poromyoidea Dall, 1886 Family Poromyidae Dall, 1886 Genus Poromya Forbes, 1844 Subgenus Poromya, s.s. Poromya (Poromya) houbricki F. R. Bernard, 1989 Subgenus Cetomya Dall, 1889 Poromya (Cetomya) malespinae (Ridewood, 1903) Dermatomya tenuiconcha Genus Dermatomya Dall, 1889 Dermatomya buttoni Dall, 1916129 Dermatomya hyalina F. R. Bernard, 1989 Dermatomya mactroides (Dall, 1889)130 Dermatomya tenuiconcha Dall, 1913 Footnotes 22 1. Nucula carlottensis occurs from British Columbia to central Mexico in as little as 100 m. 2. The genus Nucula is divided into two genera following Maxwell (1988), and eastern Pacific taxa with a smooth outer surface and lacking marginal crenulations are placed into Ennucula. 3. We regard Nucula bellotii A. Adams, 1856, and N. quirica Dall, 1916, to be synonyms of Ennucula tenuis. 4. We consider Nuculana amiata Dall, 1916, to be a synonym of N. leonina. 5. Nuculana navisa occurs from Kodiak Island, Alaska, to southern California; Nuculana spargana Dall, 1916, is a synonym. 6. Synonyms of Nuculana pernula include Nucula jacksoni Gould, 1841; Leda buccata Möller, 1842; L. fossa Baird, 1863; and L. pernula radiata Krause, 1885. 7. Nuculana elenensis occurs as far north as southern California. 8. Nuculana oxia Dall, 1916, is a synonym of N. penderi. 9. The subfamily name Spinulinae Allen and Sanders, 1982, needs to be replaced because the genus Spinula Dall, 1908, is a junior homonym of Spinula Herrich-Shaefer, 1856. 10. Ledella fiascona has been recorded as shallow as 48 m. 11. Malletia faba occurs in as little as 200 m. 12. The authorship of Yoldia hyberborea has been changed from Torell, 1859, to Gould, 1841; Nucula amygdalea Valenciennes, 1846, is a synonym. 13. Eastern Pacific records of Yoldia limatula Say, 1831, have been determined to be based on specimens of Yoldia myalis. 14. We regard Yoldia scissurata Dall, 1897, to be a synonym of Y. seminuda. 15. Megayoldia montereyensis occurs from the Bering Sea to Monterey, California, in as little as 200 m. 16. Portlandia dalli Krause, 1885, was based on small specimens of Megayoldia thraciaeformis. 17. Portlandia aestuariorum recently has been found in the Beaufort Sea [SBMNH]. 18. Yoldiella nana occurs in deep water off central California. 19. Yoldiella orcia occurs as far south as San Diego, California; we regard Y. sanesia (Dall, 1916) to be a synonym. 20. Barbatia reeveana, a Panamic species, occasionally occurs in southern California. 21. Arcopsis solida, a Panamic species, occasionally occurs in southern California. 22. An unnamed species of Porterius occurs in the Aleutian Islands. 23. We regard Limopsis diegensis Dall, 1908, to be a synonym of L. panamensis. 24. Empleconia vaginata occurs from the Bering Sea to British Columbia, Canada, as shallow as 160 m. 25. Nipponolimopsis decussata has been collected from two stations in the Aleutian Islands. 26. Mytilus edulis has been imported for mariculture in British Columbia, Canada. 27. We find no reliable characters to distinguish Crenella divaricata (d’Orbigny, 1842) from C. decussata and have synonymized them. 28. Gregariella chenui (Récluz, 1842) occurs in the western Atlantic, not the eastern Pacific. Gregariella coarctata is the eastern Pacific species that occurs as far north as central California. 29. Mytilus corrugatus Stimpson, 1851, is a junior primary homonym (non Brongniart, 1823). The earliest available name for this taxon is Musculus glacialis (Leche, 1883). 30. We regard Musculus cultellus (Deshayes, 1839) to be a synonym of M. niger. 31. These dwarf mussels require another generic home, most probably in a new genus. “Musculus” phenax occurs in the intertidal zone in Alaska. 32. We have not seen convincing characters to distinguish Megacrenella Habe and Ito, 1965, from Solamen Iredale, 1924. 33. "Crenella" leana is transferred to Solamen, based on surface structure and prodissoconch size. 34. Solamen megas occurs from southern California to Panama. 35. "Musculus" seminuda and "M." vernicosa have no strong demarcation between the central and posterior regions of the shell and are here placed in Vilasina. 36. Records in southern California of Dacrydium elegantulum Soot-Ryen, 1955, are in error. 37. Dacrydium pacificum occurs from Alaska to Baja California Sur, Mexico, in as little as 90 m. 38. Adula gruneri is the earliest name (August 1851) for what has been known as Adula falcata (Gould, November 1851). 39. Adula schmidtii, an Asian species, has been collected in the Aleutian Islands. 40. Modiolus eiseni Strong and Hertlein, 1937, which was listed in error in Turgeon et al. (1988) as occurring in southern California, but it has never been reported north of central Baja California, Mexico. 41. Modiolus kurilensis Bernard, 1983, is regarded to be a synonym of M. modiolus. 42. Modiolus flabellatus (Gould, 1850) is regarded to be a synonym of M. rectus. 23 24 43. The correct name for this genus has been a matter of great confusion. For a long time, this species was placed in Arcuatula Lamy, 1919, ex Jousseaume MS, based on a misinterpretation of its type species. More recently, it was incorrectly assigned to Ischadium Jukes-Brown, 1905, which is a mytilinid not a modiolinid. Moreover, it has sufficient anatomical differences from other modiolinids to require separate generic accomodation. 44. Idas Jeffreys, 1876 (November), is not preoccupied by Mulsant, 1876 (31 December), and should never have been replaced. 45. Pteria sterna has been collected in southern California in warm-water years. 46. A single specimen of Atrina texta was collected north of Santa Barbara, California. 47. Limatula subauriculatus Montagu, 1808, does not occur in the eastern Pacific, records having been based on L. attenuata and L. saturna. 48. Limatula similaris occurs from southern California into the Panamic province. 49. Limatula japonica has been collected in the Aleutian Islands. 50. We do not regard Ostreola Monterosato, 1884, to be sufficiently different from Ostrea to use as a separable genus. 51. We regard Dimya coralliotis Berry, 1944, to be a synonym of D. californiana. 52. Euvola vogdesi is a Panamic species that occasionally has been recorded in southern California. 53. Delectopecten randolphi (Dall, 1897) is treated as a synonym of D. vancouverensis. 54. Chlamys islandica (Gmelin, 1791) does not occur in the eastern Pacific or Alaskan arctic, records have been based on the variable C. behringiana. 55. We regard Chlamys jordani Arnold, 1903, to be a synonym of C. rubida. 56. Cyclopeten davidsoni occurs as shallow as 180 m from Alaska to Washington. 57. Cyclopecten greenlandicus occurs in the Arctic, and there is an isolated population in British Columbia, Canada. 58. Pododesmus cepio (Gray, 1850) is regarded as a synonym of P. macrochisma. 59. Lucinisca is used for this species because it has cancellate sculpture on the exterior shell surface. 60. We place the species sericata in Diplodonta rather than in Felaniella. This species is not a synonym of the Miocene Diplodonta parilis (Conrad, 1848). 61. In part following published work by F. R. Bernard, we have treated Thyasira barbarensis (Dall, 1890), T. cygnus Dall, 1916, and T. gouldii (Philippi, 1845) as synonyms of T. flexuosa. 62. We find no sound basis for separating Conchocele disjuncta Gabb, 1866, from C. bisecta, and have synonymized them. 63. We regard Axinopsida viridis (Dall, 1901), to be a synonym of A. serricata. 64. "Axinulus" redondoensis lacks a radial sulcus and herein is placed in Axinodon. 65. This Atlantic species has been obtained from 3,000 m off San Francisco, California. 66. This species has been transferred to Mendicula following other workers. It has been twice renamed on the grounds that it is preoccupied by Kellia ferruginosa Morris. However, there is no such taxon, Morris (1843) merely transferred Mya ferruginosa Montagu, 1808, to Kellia. Neither Locard (1886) nor Winckworth (1932), who independently proposed replacement names, placed Montagu’s species, now regarded as a Montacuta, and Forbes’ species, generally previously placed in either Thyasira or Axinus, within the same genus. Thus, the name was never validly replaced (ICZN Code Art. 56c). Synonyms include Odontogena borealis (Cowan, 1964). 67. We place all Astartidae species covered here in the genus Astarte as we can find no sufficient criteria for recognizing separate genera. 68. We treat Astarte rolandi Bernardi, 1859, as a synonym of A. arctica. 69. We regard Astarte polaris Dall, 1903, and A. willetti Dall, 1917, as synonyms of A. compacta. 70. We treat Astarte quadrans Gould, 1841, and A. filatovae Habe, 1964, as synonyms of A. crenata. 71. We treat Astarte alaskensis Dall, 1903, as a synonym of A. elliptica. 72. We regard Astarte mirabilis (Dall, 1871) to be a synonym of A. esquimalti. 73. We regard Astarte bennettii Dall, 1903, to be a synonym of A. vernicosa. 74. Cyclocardia gouldii occurs in water as shallow as 150 m from Washington to southern California. 75. Cyclocardia ovata accounts for records of the Asian C. rjabininae (Scarlato, 1955) in the Aleutian Islands. 76. We treat Crassicardia as only a subgenus. 77. There is as yet no sound basis for separating most families within the Galeommatoidea, so we group all the non-galeommatid taxa under the earliest family-group name available, Lasaeidae, following Boss (1982). Erycina balliana Dall, 1916, proves to be a synonym of Lucinoma annulatum (listed above). The Pliocene Aligena diegoana Hertlein and Grant, 1972, does not occur in the Recent northeastern Pacific fauna. 78. We regard Boreacola vadosa F. R. Bernard, 1979, to be a synonym of B. dawsoni, and tentatively recognize the generic name Boreacola for this Arctic taxon that has generally been placed in Montacuta. 79. This species recently has been recorded in southern California. 80. There is as yet no sound basis for separating Kellia laperousii (Deshayes, 1839) from K. suborbicularis; synonyms also include Lepton meroeum Carpenter, 1864. 81. We regard Mysella beringensis Dall, 1916, to be a synonym of M. planata. 82. This species has been placed in the genus Aligena by some authors. 83. This species has been incorrectly placed in Pseudopythina by some authors. 84. Orobitella californica has an internal ligament and thus is here transferred from the Sportellidae to the Lasaeidae. Synonyms include Erycina bakeri Dall, 1916, and E. coronata Dall, 1916. It occurs from Monterey, California, to the Golfo de California, Mexico. 85. Several species are transferred from Mysella to Rochefortia. Mysella has a very reduced posterior cardinal tooth and Rochefortia has subequal cardinal teeth. 86. Mysella aleutica Dall, 1899, is regarded to be a synonym of Rochfordia tumida. 87. The classification of this family is undergoing a basic overhaul. The current basis for the separation of Vesicomya from Calyptogena at the generic level is tenuous, and we have deemphasized this by ranking them as subgenera pending the publication of studies underway by other workers, 88. We regard Clinocardium fucanum (Gould, 1850) as a synonym of C. blandum. 89. The species name Serripes groenlandicus was first made available by Mohr (1786). 90. The species Serripes laperousii occurs as far north as Point Barrow, Alaska. 91. The Asian Serripes notabilis also is found from the Bering Sea to Cook Inlet, Alaska. 92. We regard Globivenus to be a senior synonym of Ventricolaria Keen, 1954. 93. We have grouped here an assemblage of eastern Pacific species that previously have been assigned either to Transennella Dall, 1883, or to Psephidia Dall, 1902, non Pomel, 1872. Species of this genus of brooding venerids may or may not have an anterior lateral tooth, even within a single species. 94. We regard Psephidia stephensae Hertlein and Grant, 1972, as a synonym of Nutricola cymata. 25 95. We regard Liocyma viridis Dall, 1871, to be a synonym of L. fluctuosa. 96. This species sometimes has been assigned to the genus Rupellaria, but it is a Petricola, s.s. 97. The type specimen of Psephis tellimyalis Carpenter, 1864, proves to be a specimen of Halodakra subtrigona Carpenter, 1857, leaving this Petricola without a valid name. This is being addressed in a separate paper by Coan (in press). 98. Coan (1996) treats Petricolaria as a full genus. 99. We regard Turtonia occidentalis Dall, 1871, to be a synonym of T. minuta. 100. Macoma crassula was recorded in Turgeon et al. (1988) in error as occurring in the Arctic. 101. Studies underway by Russian workers suggest that North Pacific material may be separable from this European Pleistocene species. If this proves to be the case, the name Macoma golikovi Scarlato and Kafanov, 1988, may be applicable. 102. Macoma torelli occurs widely in the Arctic Ocean and in the northern Bering Sea. 103. We regard Macoma leptonoidea Dall, 1895, to be a synonym of M. carlottensis. The latter species was recorded in error in Turgeon et al. (1988) from the Arctic. 104. Solecurtus guaymasensis occurs from southern California to Panama and the Islas Galápagos, Ecuador. 105. We have not been able to verify the distinctions between this species and S. rosaceus (Carpenter, 1864) outlined by Cosel (1992), and we have synonymized them. 106. We synonymize Siliqua sloati Hertlein, 1961, with S. alta. 107. We have transferred these two species from Spisula to Macromeris based on details of the hinge and pallial sinus. 108. We have transferred these two species from Mactra to Mactrotoma based on details of the hinge and pallial sinus. 109. Records of the Panamic ”Mactra” dolabriformis Conrad, 1867, from California are based on misidentified specimens of Simnomactra falcata and S. planulata. 110. We have transferred these two species from Spisula to Simomactra based on details of the hinge and pallial sinus. 111. This central Californian species has been recorded under the name Tresus pajaroanus (Conrad, 1857), a very different Mio-Pliocene species, and we will be describing the Recent taxon as a new species. 112. This homonym is renamed herein. In Turgeon et al. (1988), this Recent species was listed as Mya profundior Grant and Gale, 1931, which was also proposed as a replacement name, but for the separable Miocene Mya crassa Grewingk, 1850, non Vallot, 1801. 113. The name Mya uzenensis Nomura and Zimbo, 1937, which was described from the Miocene of Japan. Material so identified from the marine fauna of Japan is Mya pseudoarenaria, and it is possible that the Miocene species is a synonym of this Recent species as well. 114. We regard Sphenia ovoidea Carpenter, 1864, to be a synonym of S. luticola. 115. We have treated Caryocorbula and Juliacorbula as full genera, transferring these two species into them from Corbula. 116. Evidence exists that there is more than one species in this genus: different spawning times, eggs of two colors (red and pinkish cream), two post-larval forms (oval and triangular), juvenile shell with or without spinose external radial ridges, animal with or without a byssus, position of the adductor muscles and their scars relative to the dorsal margin, and siphon tips of two colors (red and white) (Russell-Hunter, 1949; Beu, 1971; Lucas, 1975-1976). Unfortunately, these traits have not been correlated with each other or with original or neotype material. Full taxonomic understanding of this genus is a problem that demands attention with modern methodologies grounded in sound nomenclature and using type specimens. 26 117. We regard Panomya beringiana Dall, 1916, to be a synonym of P. priapus. 118. Teredo bartschi has been collected off Santa Barbara, California [USNM]. 119. A specimen of Bankia carinata has been collected at San Diego, California [SBMNH]. It is not yet known whether this species has been introduced and become established. 120. We regard Lyonsia pugetensis Dall, 1913, to be a synonym of L. bracteata. 121. We regard Lyonsia gouldii Dall, 1915, and L. nesiotes Dall, 1915, to be synonyms of L. californica. 122. We think that Entodesma saxicola (Baird, 1863) is a synonym of E. navicula, which was described from Japan but occurs across the entire North Pacific Ocean. 123. Austroneaera semipellucida, described from Japan, occurs from Washington to Oregon, as well as in Japan, in as little as 155 m. 124. Cardiomya costata (G. B. Sowerby I, 1834) has been reported from California, but it does not occur north of central Baja California, Mexico. 125. Cardiomya behringensis occurs throughout the Bering Sea. 126. We regard Cardiomya californica (Dall, 1886), C. oldroydi Dall, in Oldroyd, 1924, and C. isolirata Bernard, 1969, to be synonyms of C. pectinata. 127. Luzonia walleri occurs from Washington to southern California in from 100 to 730 m. 128. We regard Haliris as a full genus. 129. Dermatomya buttoni occurs from Monterey Bay, California, to southern Baja California, Mexico, in from 82 to 821 m. We regard Poromya trosti Strong and Hertlein, 1937, to be a synonym. 130. Dermatomya mactroides occurs from southern California to Chile in 120 to 1185 m. Literature Cited Beu, Alan G. 1971. New light on the variation and taxonomy of the bivalve Hiatella. New Zealand Journal Geology and Geophysics 14(1):64-66. Boss, Kenneth Jay 1982. Mollusca. Pp. 946-1166, in: Sybil P. Parker, ed., Synopsis and classification of living organisms, Vol. 1, and pp. 1092-1096, in Vol. 2. New York (McGraw-Hill). 1: xviii + 1166 pp., pls. 1-87; 2: 1232 pp., pls. 88-141. Coan, Eugene V. 1996. Recent species of the genus Petricola in the eastern Pacific [Bivalvia: Veneroidea]. Festivus 28(11):118-124. Coan, Eugene V., Paul H. Scott and Frank R. Bernard 1997. Bivalve seashells of western North America. Marine bivalve mollusks from Arctic Alaska to Baja California. Santa Barbara Museum of Natural History Monographs Number 2, Studies in Biodiversity Number 2. [in preparation] Cosel, Rudo von 1992. “Solen rosaceus” — three species. Veliger 35(4):366-380. 27 Locard, Arnould 1886, 1887 [1886-1888]. Prodrome de malacologie Française. Catalogue général des mollusques vivants de France. 2. Mollusques marins. Société d’Agriculture, Sciences et Industrie de Lyon, Annales (5)8: 1-216 (1886); 9: 1-320 [including Bivalvia] (1887); 10: 1-175 (1888). [repr.: Lyon (Georg) and Paris (Baillière), x + 778 pp., 1888] Lucas, M. 1975. Hiatellidae from European coasts. La Conchiglia 7(79-80):14-15, 18. Lucas, M. 1976a. Hiatellidae from European coasts, part II. La Conchiglia 8(83-84):14-15, 18-19. Lucas, M. 1976b. The Hiatellidae from European coasts, part III. La Conchiglia 8(85-86):10-11, 15, pl. 3. Maxwell, Phillip A. 1988. Comments on “A reclassification of the Recent genera of the subclass Protobranchia (Mollusca: Bivalvia)” by J. A. Allen and F. J. Hannah (1986). Journal of Conchology 33(2):85-96. Mohr, Nicolai 1786. Forsøg til en Islandsk naturhistorie, med ad skillge oekonomiske samt andre anmaerkninger. Copenhagen (Holm). xvi + 414 pp., 7 pls. Morris, John 1843. Catalogue of British fossils. Comprising all the genera and species hitherto described; with references to their geological distribution and to the localities in which they have been found. London (von Voorst). x + 222 pp. Russell-Hunter, William D. 1949. The structure and behavior of Hiatella gallicana (Lamarck) and H. arctica (L.), with special reference to the boring habitat. Royal Society of Edinburgh, Proceedings (B)63: 271-289. Turgeon, Donna D., Arthur E. Bogan, Eugene V. Coan, William K. Emerson, William G. Lyons, William L. Pratt, Clyde F. E. Roper, Amelie Scheltema, Fred G. Thompson, and James D. Williams 1988. Common and scientific names of aquatic invertebrates from the United States and Canada: mollusks. American Fisheries Society, Special Publication 16: vi + 277 pp., 12 pls. Winckworth, Ronald A. 1932. The British marine Mollusca. Journal of Conchology 19(7):211-252. 28 * * * * * * * * Announcing New Publication Series on Regional Natural History and Native American Culture * * * * * * * * Available Soon Santa Barbara Museum of Natural History Monographs In-depth studies in biodiversity and anthropology Chumash Ethnobotany by J. Timbrook and R. Hoover Bivalve Seashells of Western North America by E. V. Coan, P. H. 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