Com parative E,thnobotanical Studies
of the Amerindian'Gfoups
in CoastalEcuador
By ANDERS S. BARFOD and IARS PETER I{IltST
Biologiske Skrifter 46
Det Kongelige DanskeVidenskabernesSelskab
TheRayalDanish Academyof Sciences
and Lettns
Kommissionrr: Munksgaard . Copenhagen 1996
Synopsis
with the Colorados, the Cayapas and the
Comparative ethnobotanical studies were conductecl
in 1982-1987' almost 2000 vouchers
campaiens
field
the
Dttrine
coaiqueres of coastal Ecuador.
The material was icrentified to 113 families ancl
were collectecl fbr rvhich 1510 uses were recordecl.
in
the collections was 930, roughlv estimated' Di396 genera. The number of species represented
Annotated plants lists are presented for
cateqories'
usase
stinction is macle between 73 clitferent
records of use overall are snake bite curing and
each of these. The usage categories with most
Ecliis the farnil,v mostlv used in snake bite therapir'
temperature reguratins baths. Gesneriaceae
aspects of the ethnobotany of the
Several
category.
usase
ble fruits is the largest"non-medicinal
Amerindian qroups in Coastal Ecttador' Most of
Colorados stand o.r, ..rrrlpured with the other
during the last 50 vears through contacts with
the aberrant features have probably been acquired
of the three grottps studied differ which
situations
socio-political
culrures livins in the Andes. The
plant resources' Distinction is made betrveen
is stronely reflectecl in their use of the surrounding
properties of the plant and cultural
inherent
two determinants of ethnobotanical practices:
are bclth cited as evidence for past
patterns
usage
and
..r.-,rr-ti"s
exchanp;e. E,lements of curing
level'
cultural exchange at the local and regional
KEYWORDS
(Chachi); Coaiqueres (Awa); ethnobotany; etl-rnomediEcuador; Coloraclos (Tsatchela) ; Cavapas
treatment; preparation; curinu ceremonies; shanaming;
cine; ethnopharmacology; vernacular
acculturation; conservation; clrrrent political siman; comparative aspects; classification of uses;
tuation; crtltural exchange'
ANDE,RS S. BARFOD
Department of SYstematic BotanY
Aarhus UniversitY
Nordlandsve.j 68
DK-8240 Risskov
LARS PE,TER K\TST
Unit of ForestrY
Hi gh sch ool
Asricultural
and
Ro1,alVe terinar,v
Rolighedsve.i 57
DK-1 87 I Frederiksberg C)
Denmark
Denmark
1996
O Det Kongelige Danske Viclenskabernes Selskab
a-s
Viborg
Special-Trykkeriet
by
Printed in Denmark
ISSN 0366-3612 ISBN 87-7304-27+9
Contents
I.
Introdtrction .
II.
Methods .
Methodolosical problems
7
B
III
The region.
I
TV
The indisenolrs peoples
History ancl linguistic affinities.
The Cayapas.
The Coaiqueres .
The Colorados
10
10
11
13
l5
V
Results .
M at er ia l s .
Land-use materials
Social prodtrcts
Food .
Animal foods and poisons
M edic in a l u s e s .
General system
Most important families and usage categorles.
Preparation and treatment .
Curing ceremonies
t7
I7
1B
l8
1B
19
l9
21
21
29
23
\T
Discussion
The ethnobotany of the three indieenous groups of Coastal Ecuador compared.
Comparison along a floristic sradient.
Naming .
T r eat me n t.
Evidence of past cultural exchanse with inclisenous sroups outside Coastal Ecuador
Curins rituals .
Botanical evidence
25
26
26
VII
General conclusions.
34
\TII
Acknowledqements
Jt)
IX
Figures1-20
37
X
Tables1-82.
59
2R
9q
9q
30
32
o 7
XI
Spanish abstract .
138
XII
List of references
139
Appendix I. Index to scientific names
141
Appendix II. Annotated list of vernacular names
151
BS 46
I Introduction
The ethnobotanical studies of the three remaining indigenous groups of Coastal Ecuador, the Colorados, the Cayapas and the Coaiqueres were initiated in 1982. The projecr had
the following objectives:
- to undertake a description of the ethnobotanical practices throughout Coastal Ecuador
- to compare plant usage patterns across linguistic and cultural boundaries in Coastal
Ecuador
- to detect possible exchanges of plant knowledge at a regional level
A comparative method was chosen because it
would allow us to discuss human use of plants
as a cultural feature in the sense that closely related indigenous groups also share many elements of their ethnobotany. The Coastal tribes
in Ecuador were considered suitable in this
context because they belons to the same linguistic group, they live in the same region and
they show many other cultural affinities not related to plant use. Their history and present situations diffea however. A carefully rendered
comparison could furthermore throw light on
how ethnobotanical practices vary along biological gradients. Of special interest was the
causal relationship between persistence of traditional ethnobotanical knowledqe and extrinsic factors such as acculturation and loss of biodiversity.
Three institutions contributed with financial
and logistic help to rhe project: the Deparrment of Systematic Botany, Aarhus University,
Denmark; Museo Antropologico del Banco
Central del Ecuador in Guayaquil, E,cuador;
and Pontificia Universidad Catolica clel E,cuador in Quito (PUCE). The following staff
members and students from the collaborating
universities participared (appearing in alphabetical order): Eduardo Asanza, Anders S. Barfod, Lauritz B. Holm-Nielsen, Lars peter Kvist
Dorte C. Nissen, and Flemming Skov.
Selected aspects of our data have previously
been published. Holm-Nielsen et al. ( 1gB3)
and Holm-Nielsen & Barfod (lgB4) presenr
preliminary results including only few plant
identifications. Kvist (lg8O, 1g8g) focuses on
the treatment of snake bites and the use of the
family Gesneriaceae. Balslev & Barfod (1982)
and Barfod & Balslev (1988) discuss the use of
palms. Kvist and Holm-Nielsen (lg87) compare selected aspects of ethnobotanical practices between the lowlands of Ecuador and adjacent countries. Kvist & Barfod (1991a) describe the curing rituals of the Cayapa Indians.
Kvist & Barfod (1991b) discuss rhe use of medicinal plants in general and give a number of
criteria that can be used to screen large ethnobotanical collections for potential drugs.
This paper gives for the first time a comprehensive presentation of all data collected durirg repeated visits to the three indigenous
communities in the coastal lowlands of Ecuador betw een 1982 and 1987 (Tabl e 1).
BS 46
II Methods
The fieldwork was conducted at various localities (Fig. 1). Most of the Cayapa collections
were made near the village of Zapallo Grande
located along Rio Cayapa close to its confluence with the smaller Rio Zapallo Grande (alt'
50 m . a. s . ,7 8 " 5 5 ' W 0 ' 4 5 ' N ). We w e re a ssi stedby
several Cayapa informants but most ethnobotanical information was provided by Maclovio
Anapa and Vicente Tapuyo. The first mentioned was in his late 40'ies at the time of the
fieldwork. His skills and knowledge of plants
were commonly recognized in the community'
He lived near the Evangelical mission station in
Zapallo Grande and considered himself a Christian. Vicente Tapuyo was younger and apparently not as knowledgeable on herbal practices
as Maclovio Anapa. The Cayapas willingly demonstrated how the plants were prepared and
used. We also attended a nocturnal healing
performed by a shaman livine on Rio Bolborde
(]rande'
ibout 40 km. up river from Zapallo
The Cayapas from Rio Bolborde occasionally
visit the meclical clinic set up by missionaries in
Zapallo Grande, one day's travel downstream'
Otherwise they rely on plants for curing minor
afflictions and ailments. The work with the Coaiqueres was concentrated in the San Marcos
valley (also callecl Plan Grande) at 660 m'a's''
78" 17' W 1 " 0 8 ' N ). T h e b ro th e rs H e rmano and
Santiago Dinero and E,liasTai served as infbrmants most of the time. Hermano and Santiago Dinero were orphans and both in their 20'ies. They hacl an extended network of contacts
in the community and both seemed well integrated. E,liasTai was about 50 years old' FIe was
suffering from a liver ailment at the time of our
visit but still managed to assistus on short field
trips in the area. From what we were tclld in
San Marcos, he was the person that people
trusted the most when it came to healing' His
"contra" was renowned, also
snake remedy or
outside San Marcos. The work with the Colorados was conducted in the reserve of (longoma
at krn. 16 on the road frorn Santo Domingo to
P uerto Li mon (al t. 200 m.a.s.,79" 2' W 0" 20'5) '
A shaman apprentice Mantrel Agtravil contributed, most of the information concerning plant
use. He was approximately 25 years old' At
times, he was assisted by his older brother
Ramon and by his parents. Table 1 gives firrther details on the fieldwork. The first contact
with the informants was established through
the chief of the community or through the local representatives of the Amerindian orsanisation. One shortcominq of the str-rdyis that no
woman participated in the fleld work. As a consequence, insufficient infbrmation was obtained on plant uses relating to the f'emale reproductive system. It should be noted that the
informants were paid for their participation
which consisted mostly in pointins out the useful plants in the field and providing infbrrnation. Communication was conducted in Spanish. Maclovio Afrapa and Vicente Tapuyo acted
as interpreters during our visit to the communi ty i n R fo B ol bol de rt' l l et' e ottl r' ( )avap a is
spoken.
Evidently, individual expertise on plants varies among the members of'a given cornmunity'
This applies both to overall knowledge and
knowledge on certain usage categories or plant
sroups. \Arhen it comes to ritual curins of serious and chronic ailments the shaman is the expert. Durins the later visits with the Cayapas we
o b r a i n e d m u c h i n f < r r n t a t i o no n s h a m a n i s t i cr i t uals on which subject our infcrrmants were very
knowledgeable. The Colorado informant was a
shaman apprentice. He was the yottnuest of the
informants to assistus. The ethnobotanical information that he providecl was particularly
BS 46
consistent. He both recognised plants and recalled their uses faster than any other informant. He rarely revealed any details about shamanistic rituals, however, possibly because
these are considered business secrets.The Coaiquer informants never told us much about
their curins rituals. They indicated numerous
plants as useful for chuturl, a grolrp of non-specific symptoms usually described as seneral indisposition and non-specific body pains. The
Coaiqueres generally use the term chutun to
designate all health problems that are caused
by malevolent spirits.
During fieldwork an effort was made to collect only fertile material with either flowers or
fruits. Plants without utilitarian value were collected too because they represent an important
aspect of an ethnobotanical pattern. Collections of sterile plants were generally avoided
due to the difficulties involved in identifyirg
them correctly. Exceptions were made with
useful and valuable species that had never previously been observed in a fertile condition.
The same applies to plants that were resarded
as useful in the .juvenile stage only and for
which the mature stase was unknown to the informants. A few, very rare and highly appreciated plants were never encountered in the field.
The infbrmation about these was recorded unvouchered. One example is a plant renowned
for its contraceptive properties that is found on
the river banks. The descriptions obtained
from the informants and others stronsly suggest that this is a fungus or achlorophyllous
plant. Plant names were recorded fbllowins a
simple system of our own. The accents are sim" ' "
ilar to those used in French (accent 6gu
((\ (c)
"'"
and accent grave
and in Spanish (fr). A
after a vowel indicates that this is drawn out. A
short, abruptly pronounced vowel is followecl
"!".
by an exclamation sign
The specimenswere either dried directly on
benchesheated by kerosene burners according
to standard botanical procedures or pickled in
7
newspaper soaked in alcohol for later dryine in
Quito. A minimum of four duplicates were obtained from each plant individual. The sets of
duplicates were deposited in the following institutions: Herbario, Universidad Catolica in
Quito (QCA), Museo Equatoriana de Ciencias
Naturales, Quito (QCNE), Herbarium Jutlandicum at Aarhus University, Denmark (AAU).
One set was sent to the specialistslisted under
acknowledsements as gifts for determination.
It should be noted that a few groups such as Piperaceae, and particularly Peperomia,have been
incompletely identified due to lack of a modern taxonomic treatment. All comparisons
conducted in this paper between the three indigenous sroups are based on a scientific hierarchical classification system and not vernacular names.
The ethnobotanical information
was recorded in a standardised way in order to obtain
data that were comparable. The plant specimens were collected in the morning in the
presence of the Amerindian informants. At the
collection site, information such as the vernacular name, utilitarian value, preparation and
ecolosv were recorded on a slip of paper that
was kept with the specimen in a separate plastic
bag. It should be noted that vernacular names
have only been included in the Tables that
comprise plant products that are exported
from the region or of direct commercial interest: Table 2, timber trees and Table 24, edible
fruits. A detailed study of the applicability of
the plant names was not conducted. The validity of the names was assessedby cross checking
information obtained for the same plant species during separate field campaigns.
As many as thirty collections were made in
one day. In the afternoon we set up ethnobotanical sessions during which the informants
elaborated further on the plants encountered
that day. The sessions usually attracted many
other members of the community, especially
children and their grandparents. Much valu-
8
able additional information was derived from
these discussions. Representatives of the older
generation told, for example, how some plants
had been previously used. Our approach was
not to press them for information. Our reasons
for this were firstly that we considered the lack
of information regarding specific plants to be
an inherent and important part of the general
ethnobotanical pattern. Secondly, we were not
interested in explanations invented on the spot
to satis$rour requests. Lack of precise information e.g. on preparation, is often an indication
that a particular plant is rarely used or of little
importance. The informants often related that
a particular plant was visited by certain animals
and especially birds. This information was recorded in a rather uneven and inconsistent way
because we considered it to be outside the
scope of the ethnobotanical project at hand.
The Amerindians demonstrated. however. that
they possessa profound knowledge of rain forest ecology that enables them to exploit their
surroundings in a highly sophisticated manner.
The comparative method applied in this
paper may prove particularly interesting for scientists specialising in pharmacological aspects
of ethnobotany, but only if fieldwork is
planned carefully and according to a number
of basic principles. Medicinal plants should be
collected repeatedly at various localities and at
different periods of time. Recollection of
plants is an efficient way to test the skills of an
ethnobotanical informant.
The informants
should elaborate on usage, preparation, dosage and prescriptions. Precise information is
usually an indicator of the potency of a plant.
Poisonous and psychoactive plants are always
interesting, even if the effect is not exploited.
The active use of the plants as indicated by the
informant should be observed if possible.
When this is not possible, as is usually the case
with medicinal plants, a demonstration should
be arranged that shows the preparation mode,
dosage etc. Different informants should be
BS 46
consulted when collecting data. Ideally, sessions should be arranged with the participation of many tribe members.
Methodological problems
It should be emphasised rhat, even if the ethnobotanical cclllection is comprehensive, the
material is far from complete. Canopy trees are
generally underrepresented in our collections,
although a special effort was made to collect
them with both the Cayapas and the Coaiqueres. Certain groups of plants are particularly well represented due to the bias of the researchers involved. The study was undertaken
by botanists specialisine in Anacardiaceae,
Arecaceae, ferns, and Gesneriaceae.
The ethnobotanist with a backsround in science typically produces detailed descriptions
for the physiological or psychological imbalances that a given herbal treatment is intended
to remedy. The recorded information will typically be broken down into narrow usage categories in accordance with western scientific traditions, but far removed from Amerindian reality. Usage categories that comprise plants exploited for their structural and nutritional
properties are easily clefined because of tangible proof of their utility. More than half of
the uses recorded are referred to medicinal usage cateuories, however, and the definition and
classification of these represent a problem. A
larse fraction of the ethnomedicinal plants
that we collected with the Cayapas, Colorados
and the Coaiqueres are used to treat weakly defined ailments that are difficult to describe
within the conceptual framework of western
medicine. Indigenous peoples often have an
animistic approach to various diseases, especially those of internal organs. The way in
which they locate and heal ailments is very different from western medicine. Perhaps the
greatest challense for the ethnobotanist is to
present the results of an investigation accord-
BS 46
9
ing to western scientific standards and at the
same time remain faithful to the realitv of the
Indians.
The usage catesories used in this paper ancl
their hierarchical organisation are shown in
Fig. 2. We have tried to aclapt the system susgested in l99l by Frances Cook and Laura FIastings in a proposal fcrr standard usage descriptors to the International Working Croup on
Taxonomic Databases fclr Plant Sciences
(TDWG). The most important deviations from
Cook and Hastinss are the last five seneral cat-
esories that include plants used for curing
vaeuely clefined diseasesor symptoms.
The traditional life of the coastal Indians in
Ectrador is rapidly disappearins as their natural surroundings are impoverished by uncontrolled defcrrestation.Much of the ethnobotanical knowledge we have recorded will not be
passed to the next generation and may even
have been lost since we conducted our fieldwork a decade aso. Some clf the speciesthat we
have collected with the Amerindians are endemisms on the verge of extinction.
III The region
Ecuador spans the equator in the westernmost
part of South America. It has a surface of
273.000 km2 which is approxirnately the size of
the state of Califbrnia or half the size of
France. The biological diversity of the flora is
among the greatest in the world. The flora on
the mainland (excluding the Calapagos Islands) includes some 20,000 speciesof vascular
plants accordins to Harlirg (1979) and Balslev
( 1 9 8 8 ).
The Andean m()untains divicle the country
into two lowland regions: the Amazon or el
orienteto the East and the coastal plain c>rla co.sfa to the West. The entire Amazon resion was
covered by rain forests, until recently. Renner
e t al. ( 1990) inc l u d e d 3 1 0 0 s p e c i e si n a c h ecklist of the flowerine plants found at an elevation of below 600 rnetres in the Ecuadorian
Amazon. The nearly 100.000 Amerindian peG
ple that subsist in el oriente are distributed
amons 5 linguistic sroups: (lofan, Siona-Secoya,Waorani, Lowland Quichua, and the.|ivaroan groups (Shuar, Achuar, Shiwira). The region is still sparsely populatecl by less than 5Va
of the total Ecuadorian population. Flowever,
colonisation is steadily increasing.
The coastal plain covers 80,000 km2. It represents the phytose()graphic transition between
the desert conditions existing along the pacific
coast of Peru where the annual precipitation is
less than 100 mm, and the ever wet climate of
the Choco along the coast of Colombia where
the annual rainfall may exceed 8.000 mm. The
vesetation changes accordingly from dry deciduous lowland forest near the Peruvian border to lush pluvial rain forest on the Colombian border. For this reason several forest
types, particularly deciduous and semideciduous forests, have a limited natural aeographical
range on the coast and contain many endemic
species. Since the 1950's a vast network of
roads has been established and financed by export earnings from the petroleum industry in
particular. Spontaneous colonisation as well as
public programmes have caused conversion of
fbrested public lands. Today, there is easy access to most regions along the coast. The exceptions are found in the northern provinces
10
BS 46
of Carchi, Imbabura and Esmeraldas but roads
are currently under construction here too.
During the last decade, the timber industry expanded dramatically in this region.
The coastal plain was originally covered by
forests, except for the driest areas in the southwest. Today, only 3Vo lnasnot been converted to
farmland or secondary forest types according
to a recent estimate by Dodson and Gentry
( 1991) . M os t o f th e d ry , mo i s t, a n d w e t forests
have been cleared and virtually none have
been left undisturbed. The pluvial forest only
constituted an estimated l0% of the original
forest cover but in terms of biodiversity it was
the richest. About one quarter of this forest is
still untouched. It represents the southernmost
part of the Choco forest which is distributed
along the Pacific coast of Colombia. Dodson
and Gentry (1991) assessthe total number of
species in the coastal lowlands below 900 metres to 6,300 of which 20Vo are endemic. The
corresponding figures for the pluvial fbrest
alone are 2,300 and 25% endemics respectively. The latter estimate is probably too high
since many Ecuadorian species ranse into the
fbrests of adjacent Colombia without having
been recorded there.
The perspectives of the remaining coastal
forests are srim indeed. Flalf of the Ecuadorian
population resides in the coastal lowlands, and
since 1957 the population of Ecuador has increased from less than 4 million to approx. l1
rnillion. This rnearrs an annual increase of
2.8% which ranks amons the highest in South
America.
IV The indigenouspeoples
History and linguistic affinities
The highlands of Ecuador were occupied by
the Incas only 80 years before the arrival of the
Spanish in 1533. The Incas never invaded the
coastal lowlands where several technologically
advanced Amerindian sroups lived to the
south and along the coast: the Esmeraldas, the
Mantas, the Huancavilcas and the Punaes. Later, the Spanish occupation led to the disappearance of these cultures at such a speed that
nothing is known today about their linguistic
affinities (Murra 1946). In the humid inaccessible rain forests to the north three Arnerindian
groups with sirnpler material cultures persisted. They all seem to have descended from
cultures in the highlands. Due to isolation in
remote and inaccessible areas they have preserved their native tonsue but they have probably lost many technologically advanced fea-
tures during the process of adaptation to the
new surroundings. Probably none of the
groups settled in village situations after their
migration to the lowlands. Today, it is typical
for family units of up to 20 persons to live in
larse huts scattered throughout the forest near
small streams or alons rivers (FiS. 3). The tribal organisation is for the same reason weak.
During colonial times and until the beginning
of this century, the three groups maintained
sporadic contact to Ecuadorian society. Visits
to the surroundins Mestizo villases were most
likely kept to a minimum and were reserved
solely fbr the purpose of purchasins certain
items such as machetes and pots and pans
made of cast iron. After World War II, colonisation increased steadily in the coastal lowlands
of Ecuador. Smallholders settled in areas that
the Indians had always considerecl theirs. After
BS 46
workins the land for a f-ewyears tl-re intruders
then claimed ownership under Ecuadorian
law. This did not cause open conflict because
the Coastal Indians have no tradition for intertribal raids and for feucling with settlers fiom
the outside.
The Cayapa, the Colorado and the Coaiquer
Amerindians are erouped by Mason (1950) in
the Chibcha lansuase sroup. Before the expansion of the lnca empire, (]hibcha speaking
people probably dominated the northern
Andes resion. The groups that occupy the
coastal lowlands of Ecuador today may have
had close contact with Chibcha speaking cultures in the hishlands of northern Ecuador
and southern Colombia, at that time. During
the Inca occtrpation, Quichua replaced the native tonsues of the hiehlands of Ecuador. Later
the presence of the Spaniards caused the annihilation of Chibcha in rnost parts of Andean
Colombia. Only sroups that already lived or
migrated to remote and inaccessible coastal
and Amazonian lowlands were able to preserve
their original laneuages. Barrett (1925) quotes
Cayapa mythology accordins to which, the
group was forced to rnigrate to the lowlancls by
a strongenem).This enemy was undoubtedly the
Incas or the Spaniards. The mythology of the
Colorados also sussests that they orieinally
lived in or in the proximity of the Andes.
The relation amons the Chibchan subgroups is an unsettled issue. Cayapa and Colorado are mutually intelligible languages that
accordins to Ehrenreich (1989) orisinated
from the same stock about a millennium ago.
The affinities of the Coaiquer lansuase are less
clear. E,hrenreich (ibid.) sugsests that Coaiquer derived fiom Colorado and Cayapa an estimated two millenniums ago. Today, the geographical distribution of the (lhibcha languase
group shows a major disjtrnction between Central America and Coastal Ecuador. The most
important Chibcha speaking tribes are the
Meskitos in Nicarasua, the Cunas of the Carib-
l1
bean coast of Panama and ad.jacentColombia,
the Arhuaco and Cogi of Sierra Nevada de Santa Marta in northern Colombia, and the Guambiana and Paez in the hiqhlands of Cauca in
sotrth-western Colombia.
The Ecuadorian Coaiqueres probably misrated to the lands they now occupy in the
Province of Carchi in the early 1900's.Accordins to Kempf (1982), they came from the Altaquer and the Rfo Nulpe resions of south-western Colombia from which they migrated due
to increasine pressure on their lzrnd, their cultural integrity and their political autonomy.
The Cayapas may once have had contacts with
a southern subgroup of the Choco Indians in
adjacent Colombia, the Waunanas. Barrett
(1925) claimed that several Clhoco shamans
practised amons the Cayapas. The Choc6 language was previously supposed to be related t<l
Carib btrt Ehrenreich (1989) considers it an independent lansuage that has been influenced
by Chibcha. The Otovaleiios in the Anclean
hishlands have been in close contact with the
Colorados in Santo Domingo. Many of the rituals that the Colorado shamans perform during
c u r i n q s e s s i o n sa r e s i m i l a r t o t h o s e u s e d b y
healers in Otavalo. The rituals were intrclduceC
after Hagen visited the Colorado Indians in
1939 and gave a description of their healins rituals. The conspicuous sarong-like clothine that
the Ccllorados use is manuf-actured and sold to
them by the Otovalefros.
The Cayapas
The Cayapa Amerindians ref.er to themselves as
Chachi.They constitute the larqest indieenous
sroup in Coastal Ecuador numbering more
than 6,000 (Carrasco 19BB). The Cayapas live
along the Cayapa-Santiago river system in the
Province of E,smeraldas, and the region is
mainly covered by tropical rain fbrest. The ethnobotany of the Cayapas has remained undescribed. Barrett (1925) described the tradition-
72
al Cayapa culture with numerous details particularly concerning their material culture. Recent descriptions of their culture and situation
are found in Carrasco (1988) and Medina V.
(1eez)
.
Subsistence farming is the
Agriculture
Cayapas' most important occupation. A shifting slash-and-burn agriculture is usually practised. The most common components in the
mixed fields are cassava,plantain, corn, beans,
and taro. The crops that constitute the normal
staple diet are listed in Table 23. Several varieties of cassava are often grown in the same
field. They are distinguished by features of the
leaves, the stems and the tubers. The latter differs in shape, colour and taste. Several cultivars
of chilli pepper and Bixa are grown too. Other
plants with utilitarian value are cultivated
around Cayapa dwellings e.g. fibre plants,
spices, ichtyotoxic plants and hallucinosenic
plants. Domesticated animals other than poultry are rare.
Hunting, fshing a,nd gathenng - These activities constitute an important part of Cayapa
daily life. Most families have traps throughout
the forest (Fig. 13.d). They are checked on a
daily basis for smaller rodents and opossums.
The traps are so numerous near villages that
quarrels over property rights sometimes occur.
ln Zapallo Grande we witnessed several incidents of this problem. Hunting with blowguns
has nearly been abandoned. Ten years ago'
blowguns were still manufactured for toys or
for sale. Today, mostly rriluzzle loaders are used
for hunting. Animals caught alive in the forest
such as monkeys are often kept as pets. \t\4rile
travelling in the forest, the Cayapas gather a
multitude of plant products such as fruits, edible larvae (Fig. 12.c) and medicinal plants. If
they pass a valuable tree they often mark it
down fbr future exploitation.
Fish traps are commonly used (see Table 33
and Fig. l4). Big fish are caught with large
traps that are usually constructed so that they
BS 46
become inundated at high water levels and exposed at low levels. The fish are lured into the
trap using fruits or worms as bait. Removal of
the bait releases a mechanism that closes the
trap. The fish enter the cases underwater, and
are collected when the water level recedes'
Near Zapallo Grande, at the confluence of Rio
Cayapa and Rio Santiago the water level varies
one metre. At irregular intervals, the river rises
several metres due to hear,y rains in the mountains. Fishing with spears is common. The
spear pole is made of hear,y palm wood that is
highly resistant to rot. The spear head is made
of cast iron and fastened to the pole with a
piece of string. Children in particular enjoys
this kind of fishing. The Cayapas also fish with
nets and with plant poisons. A detailed description of Cayapa fishing can be found in Mitlewski (1985).
Cument political situation - The Cayapa community is subdivided into a number of centres
that are united in a federation. Although this
political structure has weakened the power of
the traditional chief or ufii, the latter still plays
an important role in legal matters and rituals.
Conflicts and legal disputes are often solved
within the Cayapa community. We witnessed a
chief who acted as a go-between and decided
which sanctions were to be implemented.
Ten years ago, timber was mainly harvested
adjacent to the rivers by independent lumber
men and transported downstream as rafts (Fig.
5.b). The forests were left untouched just a few
hundred meters away from the river. Timber
companies have since bought concessions that
cover vast areas and the forest has been cleared
in areas surrounding the Cayapas. The intensive logging has led to a steady decrease in
hunting and fishing yields. Malnutrition particularly among Cayapa children was already evident and widespread when we conducted our
studies in the early 1980's but the situation has
steadily worsened (Carrasco 1988, Medina V.
1992). Many Cayapas are now logging the for-
BS 46
est to generate sufficient income to supplement their uniform diet of plantains and cassava. Only groups living along the smallest tributaries high up in the river system are not dependant on pecuniary economy. Logging is restricted inside the Co tacachi-Cayapa Ecological
Reserve where few Cayapa families are settled.
The reserve was established in l968 and covers
an area of 204.420 hectares (Charvet & Leon
1992). It extends fiom the upper tributaries of
the Cayapa river system to the Andes. Before
the park was established in 1977, the Cayapas
used to undertake hunting expeditions in
these areas. This practice is now being abandoned due to park regulations.
A number of tropical diseases constitute a
constant threat to the health of the Cayapas.
This is noteworthy since the Cayapas used to
live in a remarkably healthy environment accor ding t o B ar re tt (1 9 2 5 ). T h e m o s t c o m m on
diseaseis river blindness or onchocerciasis that
was introduced to South America from West
Africa decades ago. The majority of the Cayapa
men are infected with the nematode that
causes the disease. The vector is a sandfly that
typically occurs near brooks in deep forest.
Men have a higher risk of being infected because they spend more time in the forest. The
disease will often cause blindness in later
stages.Doctor teams based rn Zapallo Grande,
where the evangelical mission station has an
airstrip, have studied the disease, but administering a cure has so far been difficult and costly.
The Coaiqueres
The Coaiquer Amerindians refer to themselves
as Awa. They live in the western foothills of the
Andes on both sides of the border with Colombia, in inaccessible areas covered by moist, pristine forest (Fig. 5.a). The Coaiqueres remain
the least known of the Ecuadorian indigenous
groups. A roughly estimated 1,000 Coaiquer
t3
Indians live in Ecuador chiefly in the Province
of Carchi, but a few live in adjacent areas of the
provinces Esmeraldas and Imbabura. Much
larger numbers reside north of the border in
the Department of Narino in Colombia. They
are more acculturated here than in Ecuador
and there are problems defining them as a ethnic group. Recent estimates of population size
differ from 4,000 (Ehrenreich 1989) to 12,000
(Oroj uel a 1992) and 25.000 (Tel ban l 9B 8).
The Ecuadorian Coaiqueres live scattered
throughout the forest in family units ranginu
from small to large. There are no navigable rivers in the area and all transportation takes
place by foot on land. The individual huts are
usually situated with several hours walking distance in between. There is a slightly higher
concentration of dwellings in San Marcos
(PIan Grande) and Cualpi Alto. The Coaiquer
are linguistically related to the Cayapas and the
Colorados but there is no evidence of recent
contacts among any of these groups. In the
Cayapa communities where we conducted our
studies, most people were not even aware of
the existence of the Coaiqueres to the north.
In the early 1900's when the Coaiqueres settled
in remote areas of Ecuador they deliberately
withdrew from contacts and in this way they
avoided integration. Today, they maintain their
culture and autonomy. In 1950, Ferdon described the Coaiqueres as an isolated tribe that
had not yet adapted to western culture. FIe
mentioned that they often travelled long distances in the forest in bare feet carrying hear,y
loads on their backs. He further noted that
some members of the group still wore clothes
made from tree bark (Fig. 9).
Orejuela (1992) has provided sorne ethnobotanical information on the Colombian Coaiqueres. Hernando P. R. & Virsano B. (1992) describe the ethnobotany of a small community of
acculturated Coaiqueres and peasant farmers
in Altaquer. Kempf (1982) gives a thorough discussion of the socio-political role of the Coai-
t4
quer medical system in Ecuador. She also provides details on the chutun curing ceremonies
and nutrition. E,hrenreich (1989) qive a comprehensive description of Coaiquer culture in
Ecuador. Villareal (1986) reports on the political situation of the Ecuadorian Coaiqueres'
Agricutture-The Coaiqueres live from subsistence farming, hunting, fishing and gatherins'
They manage a system of rotation which involves both slash-and-mulch fields and agroforestry on the abandoned frelds. The edaphoclimatic conditions in San Marcos are typical of
the Choco pluvial fbrest. The annual precipitation probably exceeds five metres and tllere
are no dry months. The soil is derived from volcanic deposits. It is grey and almost permanently water saturated. Soil samples from Gualpi Alto show a high content of quartz and feldspar, but low content of clay (Thomsen 1986) '
This generally indicates a soil susceptible to
erosion.
Patches of forest are typically cut using machetes and axes (Fig. 1 1) . The preferred site
for clearing are late succesions of abandoned
fields. The advantage is that the trees are smaller than in the surrounding forest and therefore easier to cut. The brushwclod is not
burned, but left for mulching. In this way, the
nutrients are released slowly from the deacl
biomass. The crops are sown shortly af-ter
cutting the forest and the seedlings will have to
penetrate a layer of brushwood that is I m
thick. The density of the crop plants is low. Several crops are usually mixed except fbr plantain and sugarcane which are grown in monocultures. The agricultural practices that we observed in San Marcos are very much like those
described by Orejuela (1992) from the Colombian Coaiqueres further north.
The slash-and-mulch farming system probably prevents soil erosion and protects the
seedlings from mechanical damage caused by
healy rains. The fields are abandoned after
some years when most of the nutrients have
BS 46
leaked from the brushwood. The land is left
fallow and is often enriched with tree and bush
crops, especially near the dwellings.
Hunting, f,shing a,nd grt'thering - The Coaiqueres get most of their protein from domesticated animals such as pigs and poultry. Fluntine with mvzz.leloaders is also practised. These
are often loaded with rniscellaneous pieces ofmetal and are quite dangerotts to handle.
Some Coaiquer still use blowguns fbr birds and
monkeys, but this practise is becomins rarer
(Fig. 6.d). Rodents and opossums causht in
traps constitute a valuable strpplement to the
staple diet.
Fishing is probably more important than
hunting. The Coaiquer territory is dissected by
an intricate network of streams. Fish are caught
using nets, hooks or plant poisons much the
same way as described for the Cayapas. Products gathered in the forest such as grubs, ants
and edible plants constitute an important supplement to the diet.
Current Ntotiticalsitutr'tion- In 1974, the presence of a Coaiquer community was officially acknowledged by the Ecuadorian authorities.
The first efforts to intesrate the Indians into
Ecuadorian society were made the followins
year in San Marcos or Plan Grande. A school
was constructed and a teacher appointed although not without some difficulty. Later a
small church and a community house were
constructed. \Ahile we conducted our fieldwork, a small clinic offering basic medical attention was installed in San Marcos. In an effort to assimilate the Coaiqueres, the Ecuadorian authorities have created a minga labour
system and instituted an administrative system
with a spokesmen ancl a capitan. The ca,pitan
has been given some power through a system
of sanctions and directives backed by the authorities (Kempf 1982) (FiS. 4). The scattered
distribution of the population ancl the inhospitable environment have made the integration of the Coaiqueres a difficult task.
15
BS 46
The Coaiqueres constitute an unusually egalitarian people having no tradition for tribal organisation or authority, according to Kempf
(1982). They seenl particularly susceptible to
the negative effects of a rapid transition to
modern society and acculturation. \Arhereas
the Colorados and Cayapas are proud of their
indigenous culture, the Coaiqueres appear shy
and their self-esteem is generally low. To the
outsider the Coaiqueres can seem almost
ashamed of themselves. They usually avoid using their native tonsue in front of strangers. Instead they speak Spanish however poorly or
they do not speak at all. Alcohol abuse is a
problem. In the San Marcos area, sugar cane is
a common crop. The juice is fermented into an
ale-like beverage called guarapo. Guarapo is traditionally drunk in sreat quantities during ceremonies and mostly during these. In San Marcos we saw primitive wooden distilleries in the
forest, however, that are used to further process guarapo into a very strong brandy called
trago (FiS. 11) . Several times, we witnessed
drunk Coaiqueres in Chical, a Mestizo village
situated nearby.
The colonisers generally despise the Coaiqueres whom they consider illegal intruders
from Colombia. The fact is that most Coaiqueres frequently cross the San .fuan River
which constitutes the border to visit family and
attend markets in Colombia. Many of the Indians are not registered as voters in Ecuador
and consequently, do not possess the official
Ecuadorian identification card, the CE,DUIA.
When we conducted our field work with the
Coaiqueresbetween 1982 and 1985 their prospects for the future were grim. Plans to build a
road from Maldonado to Tobar Donoso that
was to cut through the core of the San Marcos
area. have since been abandoned. Instead the
territorial rights of the Coaiquer Indians have
been secured under Ecuadorian law thanks to
the intercession of international non-govermental organisations and officials in the Ecua-
dorian Ministry of Foreign Affairs. The Coaiqueres have demarcated their territory by
cutting a narrow strip of forest around it (Survival International 1989). Colonisers and developers in the Province of Carchi are pushing
hard for a road that would link Chical with Lita. If these plans are carried out they will have
an pronounced eff'ect on the economy of Carchi. A new road has recently linked Lita with
San Lorenzo, the nearest harbour from which
there is already a mayor export of asricultural
products (Fig. 5.c). An improved infrastructure would invariably lead to waves of colonisation in the areas adjacent to Coaiquer territory.
Currently the biggest threat to the Coaiqueres
in Ecuador is acculturation. Secrecy and ostracism have been their main survival stratesv for
centuries and it has worked this far. The problem today is that the Coaiqueres have nowhere
else to go.
Kempf (1982) gives details on the health situation in San Marcos (Plan Grande). In the beginning of the eighties, the most common
symptoms of the traditional Coaiqueres were
coughs, fever and headaches. Most symptoms
were probably caused by intestinal parasites
such as helmiths (roundworm, whipworm) and
protozoans (amebiasis). Almost all Coaiqueres
were inf-ected and about 70% had three to six
diff'erent types of intestinal parasite. The infections contribute to malnutrition and discomfort. The Coaiqueres thus have very concrete
reasons to avoid hard work: it makes them feel
sick and tired.
The Colorados
The Colorados refer to themselves as'fsatchela.
They live near Santo Domingo in seven reservations that cover about 100 km2 in total (FiS.
I). Earlier during this century, the Colorados
were threatened by extinction due to repeated
epidemics (Santiana 1951). The population
dropped to its lowest in about 1950, but has
BS 46
16
since increased rapidly. Lopez (1986) estimates
to
the poptrlation size to 1,000, but according
They
low'
is
fieure
this
our Colorado contacts
Inestimate that more than 3'000 Colorado
dians live in the area todaY'
This paper presents the first and only eththe
nobotanical information collected with
cultheir
of
treatment
Colorados. The classic
data
ture is that of Flagen ( 1939) ' Earlier, some
Karsten
were provided by Rivet (1905) and
(1924i. Juncosa (1988) has recently translated
Spanthese .u.ly works on the Colorados into
descripgSO)
recent
more
a
gives
ish. Lopet (f
tionoftheColoradocultureandtheirsituation in modern Ecuador'
Agriculture, hunting, fishing and gathenng
coff-ee'
-or, Colorados grow cash crops such as
their
on
palm
oil
cacao, bananas and African
have
who
lands. Some are wealthy landowners
the
in
employed Mestizos to work for them
fields. Subsistence farming, fishing' huntins
and gatherins is no lonser practised amof$
fbr
the doloraclos since the resource basis
these activities has disappeared'
- The Colorados
Current politicat situation
to
have apparently undergone the transition
modern society without losing their cultural
poweridentity. The Colorado orsanisation is
write
to
learns
ful. The younger generation
Coaithe
their native tottg.t" in school' Unlike
Older
queres, self-esteem is generally high'
of
town
the
in
seen
be
often
Colorados can
clothtraditional
their
Santo Domingo wearing
of viving. Around their waist they wear a piece
decoratidly coloured cloth. The nude torso is
edwithbodypaint.Thehelmetlikehairstyleis
most unusual and spectacular (Fig' 8'a)'
The Colorados have lived near Santo Domingo for at least 300 years. In the 50's, the entire
i.gion experienced an economic boom and
the rain forests have been converted into agri
cultural land almost entirely' The edapho-climatic conclitions are ideal ftrr most crops' The
rich soil is of volcanic origin' A short dry sea(Fig'
son is experienced most years inJuneJuly
jittle
material
original
the
of
remains
1). Very
culture.of the colorad.os. The tribe is fully inteTheir
srated economically in modern Ecuador'
ho.rr., combine traditional architectural design with mod'ern materials such as tin roofs
uid bourds. They are surrounded by commodities such as cars' motor bikes, refrigerators'
and television sets.
Shamanism has been an important income
for some Colorados. Colorado healers are considered to be amons the best in Ecuador' They
receive patients from the Santo Domingo area
andfrclmtherestofthecountryaswell.Travelall
ling shamans have been known to practice
ou., Ecuador and in other South American
Incountries. \Arhen we visited the Colorado
were
there
1982-1985
dians during the period
morethan50C ol oradoshanransoff.eri ngt heir
services. Their number has decreased since'
ThetradeofColoradoshamanshaslittleprospect for the future, since the choice of medicinal herbs will diminish with the disappearance
of the rain forest.
ThehealthsituationoftheColtlradosdoes
not differ much from that of the Santo Domingo area in general. Throughout their reservaIior't, they have accessto clinics for medical attention.
BS 46
I7
V Results
A total number of 1991 vouchers were collected
with the Colorados, the Cayapas and the Coaiqueres. The specimens collected have been
identified to 113 families, 396 senera and 576
species (Table 75). Arough estimate of the total
number of speciescollected is 930. The number
of usesrecorded is 1510. It should be noted that
the number of usesexceeds the total number of
vouchers collected with the Colorados which is
feasible due to independent resistration of uses
for multiple purpose plants.
Vernacular names were obtained for 1832
vouchers. This corresponds to 92% of the collections. The Colorado Amerindians named
virtually all the plants whereas the Cayapas
named only the useful plants. The Cayapas
made an effort to explain to us the meanings
of their plant names. These are mostly composed of a number of syllables which may indicate usage, the part of the plant used, srowth
form, the name of the spirit that is believed to
reside in the plant etc. This kind of plant nami.g is frequent among Amerindian people.
For a general treatment on ethnobiological
classificationwe refer to Berlin (1992). In Cayapa languase, -tchape indicates that the plant
has edible leaves, -pistcha means that it has
edible fruits, -chi means that it is a tree, and tapetndicates that it is a herb. Spanish has also
been adapted in Cayapa plant names. The
Spanish word -remedio is often used as a syllable when a plant is used medicinally. The
Colorados construct plant names much the
same way as the Cayapas, whereas the Coaiquer names usually consisted of one syllable
only. The Coaiquer informants provided us
with much fewer names. They even claimed to
be ignorant of the names of some of the plants
that they indicated as useful. The vernacular
names they did use were mostly Spanish ones
which may be a reflection of their reluctance
to speak their own lansuage in front of outsiders. Our knowledge of their ethnobotanical
practices is incomplete and the discussion in
the next section will therefore focus on the
Cayapas and the Colorados.
The results of the ethnobotanical studies of
the three remaining Amerindian sroups in
Coastal Ecuador are presented in the Tables 281. The Figs. 3-19 illustrate selecreclaspects of
the way plants are used on a daily basis.
Annotated plant lists are presented in Tables
2-74 for each of the usage catesories. In the following parasraphs the most important results
will be presented for the main usase categories: materials, land-use materials, social products, food, animal food, and poison (Fig. 2).
The medicinal categories will be commented
upon in further detail.
Tables 75 and 76 present some basic statistical information regarding the material collected during the field campaip;ns.
Materials (Tables2-15)
The Colorados no lonser base their material
culture on plant resources. Instead, they purchase items such as zinc roofs, rope, construction materials, television sets and vehicles in
Santo Domingo. This is the reason that relatively little information concernins the Colorados is included in Tables 2-15. Some of the
trees that were previously used for construction are no longer available due to deforestation. One particlllar aspect of Colorado material culture remains intact, however. The Colorados exploit more plants for dyes than their indigenous neighbours (Table 15). Although the
Colorados are well integrated in Ecuadorian
society they maintain their tribal identity which
BS 46
1B
also tlse a number of
rians. The (loloralclos
t":it
and
bocly
is a consequence of
is reflectecl in spectacrtlar
shrubs for hedqes which
sarong-like
need to dernarcate
and i'-' uiuiaty coloured
intensive land-use ancl the
;"i*itg
'.t.rtnir"r[.
the oppol'epresent
Coaiqueres
The
private ProPerty
ctrlture depends
site extreme' T'heir nraterizrl
surrortndinq ftrrest'
almost entirely on the
decades they have
However, within a few
Socialproducts (Tables18-22)
and do not use
clothing
style
work' we verified
adopted western
Several times cluring field
make them
coulcl
that
this category were surany kind of' painting
that plants representing
grorlP'
The Cayapa inforstand out as an ethnic
roundecl by strong taboos'
their
of
most
lc-'st
have
of ctlllectinu plants
The Coloraclos mry
mants were gt:"t'iity afraicl
be
should
it
but
such as
culture'
traditionul -ut"tial
o'ia"t-'t physiological effects
Streptochrt'cte
fr"J".i"g
<>f
use
the
agents'.These
noted that they share
hallucinosenrc anci anti-fertilitv
The spikes lt:tu:.ut
spirits and
cavapas.
strons
the
sodiroctnnwith
or.'b"litved to house
(Table 1l ) '
ff"",,
hair
facial
bv shamans' With the
a depilatory to remove
are most ,af"lf handlecl
are also in
srowth
hair
encotrnter such tabooPlants said to stimulate
Colorados' we did not
groups
indigenous
the
a result of accrrlcommon usage among
related f'earswhich is probably
be
t.
remains
It
13).
E'hrenreich (1989) ' the
of coastat u..,ua,rr irlute
turation. According t;
fertility regulating
demonstratedwhetheranyoftheseherbal
f)oaiqueres clo '"ti" ttse
ethnobotanical
Other
Another posefficient'
treatments are
or hallucinogenic plants'
the Caya;*,1
Colorados'
the
are taboo-ridden and
similarities between
sibility is that these plants
rathreqional
indicate
example of the secrecy
pas and the Ct'uiqueres.
that it may be another
Ca'stil'la
rc
applies
This
er than local "*lft""ge'r
of this srouP'
is used to waterproof
wliith
latex
elasticrt',the
which is used tcr
canoes und Coi* lachryma-jobi
necklaces'
procluce beads for making
Foocl (Tables 23-30)
diff-erent species pro\Are recorded about 100
the Amerindians' The
ducing eclible truits for
16-17)
(Tables
from fewer species
Land-use materials
Clolorados collect fnrits
number
considerable
a
The Cayapas cultivate
thantheCoaiquerand'theCayapas(Table24).
ornamental value'
wilcl plantt i" thtt-'
of plants because of their
Furthermore they use fewer
they
but
flolorados'
the
to a general change tn
The same applies to
traditional clishes due
as
pLlrposes
other
for
exception is noticeuse ornamental plants
the diet (Table 25) ' One
numerous
the
among
have preserved
well. These are often
able, however' Th; Colorados
baths (Tain.
employed.
plant based beverplants that are
lrerbal the use of
*urr,o of their traclitional
that
probable
i'
act as stimulants or
bles 73 and 7a) ' tt
;g;t; especially those that
addirecent
a
temperal"l". (Table
ornamental plants constitlrtes
aim at raisinj the bocly
inspired
practices
the unuslj habit
tion to the ethnomedicinal
28). The Oayapas
'(Tablehave
:ll:,^t
o'-td Aftitan EcuadoMestizos
*ith
26) ' Many different specres
by conta.t,
ing no*"rs
taste of the flowers is
are used in this way' The
sweet and candy-like'
often described as very
similarities
knorvleclgc as l'oruLrvhetr
One of the nrore remarkable
;G;tu
t"f.t trr cthnobot''rnical
i:- the
sroups
Ectlaclof
Amerinclian
the coast2rllrlwlancls
berween the three
it's r:rnge i, t""'lttt:tf to
c' g'
(Table 25) ' The Cayaurore lviclclv clistlibuted'
use of terns as vesetables
or. When knowleclqe is
ttse thc telnt rcgional'
across the Anrlcs' we rvill
BS 46
pas and the Coaiqueres prepare the young
fronds with fat and salt in many of their traditional dishes.
l9
tures. In particular the perception of the function and position of the organs varies from our
concepts. Amerindians focus on symptoms and
their localisation to cure diseases.This is quite
unlike western medicine that constantly seeks
Animal foods and poisons
to explain the causal relation between symp(Tables 3l-34)
toms and disease. An example would be the
The Cayapas and the Coaiqueres frequently
most frequently treated condition amons the
use fruits as bait when fishine. They also have
Coaiqueres called Chutun. The description we
detailed knowledse about the interaction
were able to obtain resembled influenza sympbetween potential game and specific species of
toms: general indisltosition with an aching body
plants in the forest. The infbrmants often
and often uith fner. Later, we learned that chupointed out fruiting trees that are frequently
tun is a condition caused by malevolent spirits
visited by birds such as the roucan or the jungle
and that it encompasses various kinds of sympchicken. The Colorado informants rarely mentoms. The Colorados use a system of low and
tioned ecolosical observations that could be
high temperature to explain many diseases.
relevant to fishine or hunting. Few of the aniThe term temperatureshould not be understood
mals that were traditionally hunted are left in
literally and does not necessarily relate to fever
the deforested area and the Colorados no longconditions. It is used as a measure of physiologer depend on wildlife as a protein source. It
ical imbalances. Numerous plants are used to
should be noted that no direct inquiries were
coolor warm the body, accordingly.
made on plant-animal interactions during
Cardio-uascular system (Tables 35-35)
The
fieldwork. Often, this kind of relationship was
Cayapas in particular use many plants in these
mentioned when no other utilitarian value
four usage catesories. This reflects the fact that
came in mind.
the Cayapas often cite anaemia as the cause of
The Colorados, Cayapas and Coaiqueres
fatigue. It is noticeable that there is almost no
have apparently all used the latex of lVaucleopsi.s overlap in the plant spectra for the three indigfor arrow poisons traditionally. We did not colenous groups.
lect this tree with the Cayapas, but detailed deDigestiue system(Thbles39-45) - All Amerindiscriptions by the informants indicate that they
an groups in Coastal Ecuador use numerous
use l,{aucleopsis
too. Santesson (1936) reported
plants for digestive purposes, but they have few
that the Cayapas and the Choco of western Coof these in common. The only shared pattern
lombia use the latex of Ogcodeia,a synonym of
is the use of ferns and Piper for stomach disorI'{aucleopsis
(Berg 1972). \,Vhen asked abour arders (Table 4l). Most of the representatives of
row-poisontree our Colorado informant went
these two plant groups are very abundant and
awayto collect material from the only l{aucleop
availability could be the reason for their popusisthat was known to still exist in a remote part
larity in this context.
of the Colorado territory. We received the colSeveral species are listed for just one of the
lected material, but never saw the tree.
three groups, despite the fact that outside
Coastal Ecuador these are widely used and often cultivated. This applies to Spilanthes(Table
Medicinaluses(Tables35-74)
39), 7)ngiber fficinale (TabIe 39) , Chenopod,ium
Diseaseis diagnosed and explained in a fundaamhrosioides(Table 41) , Erythroxylum noaogratamentally different fashion by Amerindian culnense (Table 4l), Sltigelia (Table 4l),
Jatropha
L
-
BS 46
20
curcas (Tables 43 k 44), Ficus insipida (Table
44) and Scoparia dulcis (Table 44) '
External system(Tabtes 45-55) The Cayapas
and t h e C o l o ra d o s b o th u s e th e fami l i es A r-aceae and Cyclanthaceae for treating ant bites'
A similar pattern is found in the Amazon region (Kvist & Holm-Nielsen 1987) ' Piper tricuspe
is used for insect bites in seneral' The Cayapas
often cultivate this particular species near their
dwellings. A similar practise has been observed
amons the Cunas in Panama (Duke 1970) '
One of the more remarkable aspects of the
ethnobotany of the Amerindians of Coastal
Ecuador is the large number of plants used to
treat snake bites. Snakes represent a constant
health problem particularly around villages
surrounded by undisturbed forest' Larrick et
al. (1978) estimated that nearly half of the
Waorani Amerindians in the Amazon region
of E,cuador have been bitten by u snake at
some point in their life. During our travels in
Coastal Ecuador we observed many people
with large bluish-black scars from the bites
chiefly of the Equis snake, Bothrops atrox' lt ts
nevertheless a surprising fact that more plants
are used for treatment of snake bites than for
any other purpose (Fig. 20). Snake bite plants
constitute the largest usage category among
the Coaiquer and the Cayapas and the second
largest cateuory among the Colorados only
surpassed by herbal baths (Table 81)' The
large proportion of plants used for treating
snake bites may relate to the animistic belief of
the Amerindians. Snakes are believed to be
the dwelling place of particularly malevolent
and potent sPirits.
Most treatments that aim at curing snake
bites are purely topical (Table 48) ' Systemic
treatments are practised by the Colorados and
to a lesser extent by the Cayapas (Table 49) '
The Cayapas and the Coaiquer often vary their
therapiesinaccorclancewiththeparticular
species of snake responsible for the bite' The
iolorado informants on the contrary often re-
k
lated that a particular herbal treatment was
good for snake bites in general, no matter the
ipecies involved. The choice of which plants to
.rr. in a siven situation ranges from being rather random to very specific. The lists of snake
bite plants comprise a total of 26 plant families
and vascular cryptograms. Two families and
the vascular cryptograms are represented with
more represelltatives than any other family
(Table 48-50). The Gesneriaceae (Table B0)
and the vascular cryptogalns (Table 78) are
commonly used for snake bites by all three
groups, whereas the Piperaceae (Table 79) is
used by the Cayapas and the Coaiqueres only'
The Coaiqueres use Gesneriaceae for socalled dry baths that are applied to miscellaneous eczemas (Table 55). The ash from the
burned plant is rubbed on the affected area of
the skin. This kind of preparation was not observed at the Cayapas and the Colorados' Apart
from the use of Gesneriaceae for eczema' no
other family is particularly dominant in the usage categories comprisins cuts, swellings, ab,..rr"r, and fungal infections (Table 51-55) '
Few plants in the lists were shared among the
three indigenous sroups. One exception is the
genus Acatyphawhich was used to treat cuts by
ill tht." groups (Table 51). The treatment of
fungal infections (Table 54) is discussed in
Kvist & Flolm-Nielsen (1987). The most common fungal infection causes a symptom called
manchas blancaswhich refers to white' usually
concentric circles on the skin where the pigment has been cl estroyed.The trse o[ t he var iegated leaves of Calatheametallica for this specifi. ptt.pose is probably based on the so-called
Doctrine of Signatures. Tlnts principle has influenced folk medicine all over the world and is
also widespread among Amerinclian sroups in
South America. The signaturerefers to the correspondance between certain physical plant features and the medicinal purposes fbr which it
is valuable. A good example is a species of
Monolena that we collected with the Cayapas' It
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lras long, red, worrn-like peduncles that are
trsed accordinuly ftrr curins intestinal wclrms
(see Table 44\.
Muscular-skeleta,lsy.slem(Ttthles5 6-5 7), neraous
slstetn (table 5B), reprorLuctiuesystem(tables59-51),
respirntorl system(Tnbles52-65), sensorj slstem (Tables 55-57) a,nd uriner\ s),st(m ('fables 58-59) There are no apparent patterns ernerging
from the list of plants in these usase categories.
The rationale behind many of the applications
is probably the Doctrine of Sisnature. Some examples are given in the lesends to the Tables.
For a general discussion of herbal treatments
of eye ailments, see Kvist & Holm-Nielsen
( 1e87)
.
General system (Tables 70-71)
A number of'r,veaklydefined pains and fevers
are treated with herbs. The plant choice is rather arbitrary and varies from time to tirne. The
plant sroups used nlost frequently are the
abundant ones and those that are eas,vto collect, viz. plants in the under storey belonging
to vascularcryptoganls, ArAceae, Gesneriaceae,
Melastcxnataceae, Piperaceae, Rubiacezre and
Solanaceae.A minor number of plants are
used in systernic treatments (Tables 70-71).
The Colorados use nurnerous plar-rtsin their
bath treatments (Table 73). A third of all the
plants collected with this group were used for
this purpose. Up to 20 different plants are
mixed in a single bath.
Most important families and
usagecategories
Fig. 19 shows the most irnportant families of
plants used by the three ethnic sroups. Table
77 lists the families that were rnost valuable tcr
the three Amerindian groups based on the
number of uses. It shor.rldbe noted that several
identical records of use of the same plant
acrossthe indigenous sroups are registered for
2r
each of these separately. For each ethnic
sroup, bclth total and relative fisures fbr the
number of uses registered within a given family
are shown. The farnilie circumscription fbllows
the taxonomic system used at Herbarium Jutlandicum (AAU) where the first set of oqr collections is deposited. Here, the plants are organised accordine to Willis (1966) with a few
exceptions. The importance of ferns and socalled fern allies has been emphasised by treating these as a separate unit at the family level.
Eight plant groups are listed amons the 10
most valuable for all three indigenous sroups:
vascular cryptogams, Piperaceae, Gesneriaceae, Solanaceae,Araceae, Rubiaceae, Melastomataceae, and Asteraceae. Tables 78-80 compare the usase patterns of the three most frequently used plant groups for each of the three
Amerindian tribes, vlz. the vascular crypto€larns, the Piperaceae and the Gesneriaceae.
Identical usase of a particular plant species by
separate indieenous groups contributes to the
fisures for each of these.
Fig. 20 shows the breakdown of recorded
uses by usage catesories for all three ethnic
sroups. Several iclentical records of use clf the
salne plant across the indigenous grollps are
counted once. The two most important usage
categories are those of snake bites and baths
regulating body temperature respectively. Both
comprise topical treatments only. The most important systernic treatrnent only rank number
ten overall. The choice of plants for systemic
treatrnents is obviously narrow because of the
serious side-effects that plants may have when
t a k e n i n t e r n a l l y .E d i b l e l i u i t s c o n s t i t u t e a v e r v
large usase catesory. Ahnost a hundred different speciesare used by the three groups. Table
Ul lists the ten most important usage catesories fbr each of the three indieenous groups
studied. It sives the total and relative figures
for the number of uses resistered within a given usase category. As in Tables 77-80, identical
usase of a particular plant speciesby several in-
22
disenous groups contributes to the individual
figures for these.
If proportionality is assumed between the
number of plants used for a certain practice on
one hand and how common this practice is on
the other hand, then snake bites would be the
most common ailment followed by the three
usage categories classified under General
System: topical therapy to treat miscellaneous
pains (Table 72), reuulation of body temperature (Table 73) and vaguely defined symptoms
(Table 74). Hereafter several minor ailments
follow such as skin reactions, wounds, and
stomach infections. Snake bites indeed constitute a major health problem. The most common snake is the Equis (Bothrops atrox). It
causes more deaths than any other snake in the
coastal lowlands of Ecuador although it is not
as poisonous as e. g. the Bush master (Lacheis
muta). The purpose of topical treatments of
snakebites is both to prevent and heal necrosis
around the bite and to expel the snake's spirit
that has entered into the body of its victim.
Skin reactions (Table 55), wounds (Table 51),
and stomach infections (Table 41) are amons
the most common health problems especially
in isolated communities that do not have access to prescription medicine. Other common
ailments are physical traumas such as swelling
from bruises (Table 52) and injured joints (Table 57), intestinal parasites (Table 44), colds
and influenza (Table 65) and fungal infections
(Table 54). The fungal infection menrioned
earlier causing manchas blnncas is particularly
common. A number of physical imbalances related to internal'organs may also be widespread but they are difficult to diasnose for a
botanist with no background in medicine.
Some of the symptoms referrecl to the General
System category used in this paper may well be
related to specific diseases,particularly topical
treatment of miscellaneous pains (Table 72)
and topical treatment of vaguely defined pains
(Table 74). Information on plants used by
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women only in connection with birthgiving
and menstruation (Table 59-60) and ro regulate f-ertility (Table 19) are difficult to obtain
due to taboos and a reluctance to dicuss one's
intimate life with representatives of the opposite sex. More studies concentrating on this aspect in particular are needed.
Preparation and treatment
Medicinal plants are prepared in various ways.
Leaves and twigs are either used as is, or
ground into a paste. Further processing often
involves boiling. The insoluble residues are either separated by decantation or included in
the preparation. In other cases,extracts of the
plant material are produced using cold or lukewarm water. Yet another kind of preparation
involves burning the plant followed by an application of the ash.
In decreasing order of importance treatments include: drinking; bathine; compresses
made from either the fresh leaves or a paste produced from these; compresses made from the
left-over plant residue from decoctions or extracts; massasing with the twigs; showering with
extracts or decoctions; so-called dry bathing
where ash from a particular plant is sprinkled all
over the body or on the affected area of the skin
only. Showering implies that the liquid is
sprayed all over the patient from the mouth of
another person. A common practice among the
Cayapas and the Coaiqueres is massage using
twigs from shrubs or trees. Treatmentwith plant
paste compressesis also quite common (Fig. 15).
Both crude and boiled material are used. Compresses are in some casesremoved after a short
time, in other casesthey are held in place by a
bandage and worn for days. Pieces of cloth or
leaves are used to make the bandages.
Symptoms related to the external system are
usually treated topically. This applies to skin reactions that are caused by contact allersies, for
instance. Symptoms related to inner organs are
BS 46
treated both systemically and topically. Systemic treatments are generally used when symptoms are confined to a particular function of
the body. Couehs and diarrhoea are, for example, very often treated by drinking decoctions
or extracts. Topical treatments are frequently
used when symptoms are not ref-erred to any
particular organ sroup. Combination of general symptoms such as fever, pain and fatigue are
often treated topically, as well.
Curing ceremonies
From the Cayapa Amerindians we obtained the
most detailed information about shamans and
the ceremonies that they perform. The curins
systemsof the Colorados and the Coaiqueres
are similar in many aspects.Accordinp; to the animistic belief of the Cayapas,numerous spirits
reside in nature e. g. in animals, in plants, in
stones,and in hills and they are all potentially
harmful. A spirit may enter the human body as
one passesits dwellins or as one sleeps. \Arhen a
person dreams, the Cayapas believe that the
soul departs from the body leaving it open to invasion by other spirits (Barrett 1925). Dead
corpsesof animals and humans alike are considered a potential danser because spirits may depart from these and beein to travel about in the
village in search of a new dwelling. If they enrer a
human body they could cause chronic disease
or severehealth threats. Curing rituals aim at expelling the malevolent spirits and transferrins
them to another dwelling. Only an experienced
shaman can accomplish this transfer which is
considered highly risky. He will rransfer the malevolent spirit from the patienr ro his own body,
and later to a third dwelling place. If he is roo
weak he may fail to make the second transfer of
the spirit and become a victim himself. The shaman either conducts the curing sessionpersonally or guides it from nearby. The number of
plants that is included in ritualistic curing is variable from just one or a f-ewto several. Flallucin-
23
ogenic plant druss are frequently used to induce strong visions that enable the shaman to
see and talk to his tutelary spirits.
Animistic belief also influenced rhe field
work. Particularly powerful plants are supposed to house spirits that may become malevolent if collected and handled by laymen. For
this reason we often experienced some reluctance on the part of the Cayapas who assisted
us when we asked about plants that they used
for contraception, abortion, arrow poisons,
mental disorders etc. These plants are not considered harmful to the shaman as long as he
possessesthe protection provided by his tutelary spirits. An example would be an incident
where one of us nearly stepped on a highly poisonous and aegressiveEquis snake. Accordine
to the Cayapas the snake had appeared because we had upset its spirit by collecting large
quantities of plants used to treat snake bites.
They declared that if we met a second snake
that day under dangerous circumstances, we
would have to stop the off-ensive plant collectins. Otherwise we would meet a third snake
that would strike us. After the Cayapas had
killed the snake we parricipated in a ritual
aimed at expelling and scaring its spirit away
fiom the village and into the forest. Smoke
from cigarettes was exhaled upon the dead
snake along with the exclamation of loud conj urati ons (Fi S . 16). B arrert (1925) descri bes
other exorcistic rituals durins which tobacco
smoke was exhaled on objects in Cayapa dwellings presumed to house spirits.
In October 1983, we attended a nocturnal
curins ritual amons the Cayapas that live along
a small tributary about 40 km upstream from
the villase of Zapallo Grande. The curing took
place in the dwelling of a middle-aged shaman
called Maritimo. The families of both the shaman and the patient were present. No local inhabitants spoke Spanish and all communication between them and us went through our informants from Zapallo Grande who had ar-
L-
24
ransecl the visit. The patient was an approx' 35
rivy"ui old man. He apparently suffered from
er blindness or onchocerciasis, the parasitic
is
diseasedescribed earlier in this paper which
introbeen
has
caused by u nematode that
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apa word for expel, followetl by huii' Lt c>ne
point, the wif'e and the fotrr children of the paii"t t were also massaged. The next mornins
we were infbrmecl that the shaman had transferred the malevolent spirit strccessfirlly' He
would expel the spirit from his own body during a ritual performed the following night, but
we did not attend this ritual. \A'ewere infcrrmecl
that dr.rrinq this second night he would not
drink ayahuasca btrt onlv trago (sttuar cane
ducedfromAfrica.Theritrrallastedtheentire
nis h t fro m 8 P.-. to 6 a .m '
tn" atmosphere cluring the curius session
was surprisinuly Llnceremoniotts' The first
three hours comprised a preparatory phase
brandy).
pinde'
durinq which the shaman slowly drank
In adclition to the nocturnal crtring sessions
This is the Cayapa name for the hallucinosenic
performecl in hortses, other rituals were perdecoction prepared from the malphigiaceous
iormed at particular sites adiacent to rivers or
liana Banisteriopsis ca'a,pioften and rnixed with
in the forest. These rituals were guided by o
other plants having psycho-act.iveeffects' It is
shaman who had previously instmcted the famknown as ayahuasca in Quichua' Durinq these
ily members about their functions' We athours the shaman whistled, sans and occasiontended such a cttring that took place on a riverally rattlecl with twigs from a common riverbank (Fig. 10). It involved a family of four perbank tree Pithecellobium tongtfolium (the only
sons and. lasted for two hortrs' The patient dicl
other plant used durins this nocturnal ritual).
not participate activelv. His wife and children
About l1 p.-. the next phase of the curing ses-arsug.d him lvith plants collected by the shasion started. Waves of hallucinations now inman the same morning. A plant decoction was
creasingly influenced the shaman, and he was
prepared in a big pot. After this had cooled it
ready to lo.ut. ancl identify the malevolent
was- used first to bathe the patient, then to
spirit. A candle was lit, ancl he arranged a numbathe his wife and the children' Most of the deb.. of obiects that were considered to house
coction was potlred with a calabash and the
of
his tutelary spirits on a low table in front
rest was sprayed by mouth' The shaman did
him: black, flat and rounded riverbed stones'
not take any plant clrugs on this occasion' He
weathered stone sculptures of possibly pre-Coplaced himself in a squat position a little relombian origin, an antique Spanish sable' a
moved from the family with his ceremonial
carplastic candlestick holder shaped like a
staffand.threeblackanclsmoothvolcanic
toon character (Batman), and a polished staff
stones in front of him. In his riqht hand he
the
of clark hardwood with a handle carved in
helcl a bunch of twies. During the entire ritual
s h a p e o f a m o n k e y (F i g ' l 0)' H e chanted'
hesanginalorvvoice_almostlikeaChristian
shouted out conjurations and whipped the obmass.A ti rregttl ari nterval sheshoutedollt conbefore
jects in front of him with the twigs' Just
jurations
like ushnii and hit the river stones
Lia.tignt, the visions allowed him to identify
hard with his twies'
to
and lcrcate the malevolent spirit' Now he had
The Cayapas draw a parallell bet'i'veenthe inexpel ancl transfer the spirit to himself' He extestinal system ancl the river since both are ttnitinguished the candlelight, and the rest of the
directional ancl closed. Consequently' they
,rig"ht the shaman was very active chanting and
blame ailments related to the intestinal system
,rtlr.ugir-rs the body of the patient with the
on spirits from the river. An efficient cure contwigs. Eu.ty now ancl then he criecl out conjusists of chasing the harrnful spirits back to the
r a ti o n s ]l l k e u s h -n i i w h i c h i s a vari antoftheC ay-
BS 46
25
river. Virtually all plants growing between the
lowest and highest water level of the river are
considered to be useful for this purpose. As part
of the preparation the plants are soakecl in a
small, water fillecl hole or pond dug by hand situated close to the river (FiS. 10). This explains
the casual collection of river plants used to massageand bathe the patient. Other curins rituals
aim to return malevolent spirits to the forest,
and therefore take place here. The shaman acts
merely as a suide and usually does not participate actively in the cerem()ny. A kind of a plant
steam bath is set up. The forest floor is cleaned
for herbs and shrubs. A shallow hole is dug in
the ground and filled with water. A rack strong
enough to hold the weight of an adult person is
constructed above this hole and covered with
leaves and twigs collected in the understorey
near the site. Beside the hole a fire is lit, and a
number of stones are heated. The hot stones
are then dumped in the water filled hole and a
dense steam is produced that passes through
the herbs on the rack. The patient receives the
steam bath while squattinu on top of the rack.
VI Discussion
The ethnobotanical practices throughout
Coastal Ecuador have been described in the
preceding sections of this paper and in the lesends of the Figures and Tables. The present
study had two more objectives that will be discussed in this section: comparison of plant usage patterns in Coastal Ecuador across linsuistic and cultural boundaries and the detection
of possible exchanses of plant knowledse at
the reeional level. In this discussion we will focus on medicinal uses which are particularly
interesting to compare because they are determined by an intricate combination of cultural
variables ancl properties inherent to the plant.
The three Amerindian sroups all make a clear
distinction between medicinal plants that are
commonly used ancl plants that are trsed exclusivelyby shamans. Minor health problems are
not credited to spirits and can be treated by
any member of the community. This applies to
bruises, light burns, rashes, funeal diseases,
acute headaches, indigestion and so forth.
Usually, it is the older people who are most
knowledgeable about plant treatments. The
forest adjacent to the dwelling is searched
thoroughly for herbs. Some plants are held in
higher esteem than others. The choice of
which plants to collect is often made as a tradeoff between the plant's value for the treatment,
it's immediate availability and the seriousness
of the affliction. Valuable herbs may be stored
in alcohol for later use. Snake bite remedies
are prepared in this way and stored for years. It
is generally easy for the ethnobotanical researcher to relate to these casesof palpable ailments and physical trauma.
The plants used during ceremonies guided
by the shaman are not chosen because of their
phvsiological effects. The choice, which is rather arbitrary, is based on cultural variables such
as tradition and Doctrine of Signature. Shamans migrating to other resions will preserve
their curins ceremonies but they are forced to
choose new plants from an exotic flora. The
performance of the rituals is consequently a
more stable trait than the spectrum of plants
used. More studies are needed in order to derive specific hypotheses about the accidental
26
BS 46
The ethnobotany of the Colorados deviates in
several aspects from their neighbouring groups
in lowland E,cuador. The most important
group of useful plants, vascular cryptogams, is
used mainly for preparins baths that aim at
regulating body temperature. Plant baths are
used more rarely by the Cayapas and the Coai
queres. Another difference is the use of vascular cryptogams for systemic treatment of snakebites by the Colorados, a practise that was not
recorded among the other groups. The Colorados further stand out in the way they use the
family Piperaceae. This sroup of plants is very
important for the Coaiqueres and the Cayapas
for topical treatment of snakebites. The Colorados do not use Piperaceae to treat snakebites
at all. It is further notable that the Coaiqueres
do not use Piperaceae for regulating body temperature. They very rarely practice this healing
system which probably has it roots atnottg Atnerindian groups in the highlands such as the
Otavalos. For all three indigenous groups, the
family Gesneriaceae is important fbr treatment
of snakebites, especially in topical therapy. The
Coaiqueres also use the family for treating
rashes for which purpose the leaves are burned
and the ash applied to the skin. Vascular cryptogams, Piperaceae and Gesneriaceae are gen-
ble 81). \Ahen the usage patterns of the three
most important plant sroups, vascular cryptogams (Table 78), Piperaceae (Table 79) and
Gesneriaceae (Table B0) are considered across
the three sroups, the Cayapas and the Coaiqueres show many similarities, whereas the Colorados differ. These results reflect the differences between the current cultural and socioeconomic situations. The Colorados, who are
by far the most acculturated group, also have
the most aberrant ethnobotany whereas the
Coaiqueres and the Cayapas display more or
topical treatless the same patterns. Sna,kebites,
menl is the largest usase category fbr both of
the latter. Although, they do not appear in the
same order, the other usages listed are shared
by the Cayapas and the Coaiqueres with only
two exceptions. The ninth largest category on
the Coaiquer hst, Fodder and fish bait, is not
among the ten largest usases for the Cayapas
and Stomachinfections,which is number B in the
list for the Cayapas, does not appear on the list
for the Coaiqueres. These minor differences
probably fall within the variation which can be
ascribed to sampling error. We consider the
aberrant position of Colorado ethnobotany as
a phenomenon that has developed over the
last 50 years. The palms, for example, no longer rank amons the top-10 plant groups for the
Colorados (Table 77).These plants are mainly
exploited for their structural properties and
they disappear from the ethnobotany early in
the process of acculturation. A recently acquired feature of Colorado ethnobotany is
plant bath therapy. This usase category is now
the most important.
erally used for more purposes by the Cayapas
than by their neighbours.
\Arhereasfew plants are used the same way at
the specific level, it is noticeable that eight of
the l0 most frequently used plant groups at the
family level are the same among all three indigenous groups (Table 77), and six of the most
important usage categories are the same (Ta-
Com,parison along a floristic gradient
Knowledge that is generated within a narrow
social context for example based on experiments will add distinctive features to an ethnobotanical pattern, whereas knowledge that
has been exchanged across cultural boundaries will have the opposite effect. Evaluating
component of ethnobotanical patterns that are
testable with rigorous methods such as inferential statistics.
The ethnobotany of
the three indigenous groups of
Coastal Ecuador compared
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the relative roles of these tw<t determinants of
ethnobotany is complicated at best. An indirect
method would be the comparison between the
plants enumerated under a given usage category for different linguistic groups that are distributed alons a floristic gradient. Coastal
Ecuador is characterised by several steep floristic gradients. One gradient rllns north to south
and parallel to the Andes whereas another one
is directed east-west,fiom the Andes to the sea.
In the foothills of the Andes climatic conditions often vary radically fiom one valley
system to another, a phenomenon that causes
pronounced local diff-erences in floristic composition of the vegetation. Dodson and Gentry
(1978) stated that only 37Voof the tree species
recorded by Little and Dixon (1969) in their
list of woody angiosperms for Esmeraldas were
found at the Rio Palenque Science Centre and
that most of the tree specieswhich occur at the
Science Centre were absent from Little and
Dixon's list. For the non-woody flora and especially the weedy plants, we expect the differences to be less pronounced.
The degree of consruency between the lists
of plants used fbr the same purpose by any two
indigenous groups increaseswith the taxonomic level chosen fcrr the comparison. Closely related species may vicariate for each other in the
sense that they are trsed in the same way by two
indigenous groups and probably have the same
effect. These coinciding patterns will not be
registered as similar at the species level, however. We estimate that less than 5% of the plants
in our material are shared between any two of
the three cultures studied. Similarity can not
be expressed in exact figures at the species level due to the high number of unidentifiecl
specimens in genera such as Anthurium, Pepera
mia, and Piper that are not sufficiently known
taxonomically. It is possible that the low percentase is a collection artefact, although a
more likely explanation is that it reflects the
pronounced floristic differences that exist in
27
the vegetation surrounding the three groups.
From Dodson and Gentry's paper cited above,
it follows that less than a third of the woody
species surrounding the Cayapas and the Colorados are the same. The differences between
their ethnobotanies are noteable since these
two groups speak mutually intellisible languages and are known to have maintained contact fclr centuries.
The forests surrounding the three indigenous groups were not subjected to detailed
vesetation analysis and thus, it is not possible
to make precise comparisons between the taxonomic patterns of the plants recorded as useful
on one hand and the floristic composition of
the surrounding forest on the other hand. Fig.
18 compares the most important eroups of
plants used by the Colorados with the family
composition of the surroundins forest vegetation. If plants are not actively searched for, but
instead collected randomly according to their
availability, some congruency is expected. This
requires that diversity of most families is proportional to their availability, knowing quite
well that this does not apply to families with
many epiphytic representatives such as Orchidaceae. The Colorados were chosen because a
flora exists covering the Rio Palenque area
nearby (Dodson & Gentry, l97B). The families
in the flora have been reorsanised to fit the
taxonomic circumscriptions of this paper. The
yearly precipitation at the Rio Palenque Science Centre is about 2650 mm which is almost
equal to that of the Santo Domingo area (See
climatic diagram on Fig. 1). The two family
spectra show a surprisingly high degree of consruency. The families primarily used are senerally also the ones represented by the highest
number of species in the flora. This applies to
vascular cryptogams, Gesneriaceae and Piperaceae. The very diverse orchid family is, not surprisinuly, little exploited since most species are
rare epiphytes from the forest canopy. The
purely graphic comparison in Fig. l8 indicates
28
that plant choice has a strong random element
to it. This applies particularly to the usage catesories under General systemsthat include the
highest number of plants (Table 72-74).
\Arhen useful plants become rarer due to deforestation, alternatives are sought out based
on simple cost-benefit considerations. Further
desradation of the resource basis will eventually lead to acculturation. The material culture
will usually disappear first. This is clearly the
case with the Colorados where only the older
generation remembers how to exploit the
structural and nutritional properties of plants.
For the younger generation this knowledge is
irrelevant, particularly since many of the plants
in question can no longer be found in the area.
The Colorados have the least elaborate plant
classification system of the three ethnic groups
if percentage of plants that are named is used
as an indirect measure (Table 76). Of the
vouchers collected with this tribe only 32%
were named. The corresponding figures for
the Cayapas and the Coaiqueres are 93% and
56% respectively. However, the Colorados also
stand out from the other two tribes studied by
providing both an Indian and a Spanish name
for almost half of the named plants and by using many plants for multiple purposes.
Ir'/aming
One of the aims of botanical science is to name
plants unambiguously and to classify them in a
hierarchical system that ideally reflects phylogeny. Scientific plant names are binomial, being
composed of a senus name followed by a species name. The names typically describe an important aspect of the plant, its distribution, or
simply comrnemorate a famous colleague. The
Amerindian groups in Coastal Ecuador use
non-hierarchical classification, although they
do recosnise that certain groups of plants such
as palm trees are similar, however, this is not
the main criterion for their naming. Inclusive
categories are only erected to accommodate
BS 46
variation at lower taxonomic levels such as varieties of Cassava.In many cases there is not a
one-to-one correspondence between plant species and name. Plants that are obviously different but used for the same purpose may have
the same name, whereas plants with multiple
purposes may be named differently depending
on the particular use in mind. The same plant
may also change name as it passesthrough the
different life stages from seedling to mature
tree. The geographical distribution of the names is variable, too. Some plants are used and
named locally by just one household whereas
others are widely used and known under the
same name throughout a resion. The names of
ritual plants that are collected more or less randomly in certain habitat types are usually not
very specific nor constant. There are ways to
verify indirectly the stability of a ser of names
provided by u given informant. Plants should
be recollected during repeated field campaigns in the same area eventually usine different informants. Sessionsshould be set up with
several members of a community to see whether a plant name is in seneral use or has been
constructed ad hoc.
Despite these ambiguities, Amerindian plant
classification is by no means arbitrary. The
system is in fact flexible and reflects the social
situation of the Amerindians. People dispersed
in small and isolated communities will invariably develop individual ethnobotanical practices and namine systems.Plant names are adjusted to the context in which they are used.
The identity can be specified to a higher or
lesser desree by varying the number of syllables in the name. The descriptive plant
names that combine several syllables are similar to the phrase names used in pre-Linnean
classification in Europe prior to the lBth century. The advantage of combining several descriptive syllables is that the identity of the
plant referred to is embedded in the name.
\Arhen applied, the name will immediately in-
BS 46
voke an image for the recipient if he or she is
knowledgeable about plants. Durins our ethnobotanical sessionswith the Cayapas,we often
overheard discussions that demonstratecl the
flexibility of these descriptive phrase names. It
was clear that, sometimes the informants were
uncertain or disasreed upon the identity of a
given plant that we had collected earlier the
same day. In some cases,the collection did not
include sufficient infbrntation on e.s. location
and growth fclrm of the plant. In other cases,
however, naming differecl throughout the community. As a result of these discussions,either
two names were given to the plant, or a name
wasasreed upon which combined the syllables
of both of these. Cross checks of plants collected repeatedly clurine the various fielcl campaigns revealed that plant namins was stable in
general. Even some of the longer names were
used in a consistent way.
Treatment
Both systemic ancl topical treatments of the external system may produce physiological effects. It is unlikely, however, that topical treatments of symptoms relating to the inner organs will have any effect, at least when measured by western stanclards.Amone the Amerindian groups in CloastalE,cuador, this treatment
is performed exclusively by shamans. The shaman is consulted when a cliseasedevelclpsinto
a permanently debilitatins condition. The Amerindians of Coastal Ecuador often explain disease as malevolent spirits inhabiting the body
as mentioned previously. Healins implies that
the identity of the spirit is known and thar its
abode in the body has been localised. It is believed that only the shaman possessesthe powers to expel the spirit. The plants that he uses
in this specific context are considered worthless in the hands of persons other than him.
!\rhen the same plants are used fbr a clifTerent
purpose, however, there is no constraint as to
who may use them. Curing ceremonies per-
29
formed by shamans are purely ritual in most
cases; whatever effects they may produce are
probably psychological or psychosomaric. The
shaman's function is often as a spiritual advisor, rather than as a doctor.
Certain plants are lesspromising than others
fiom an ethnopharmacolosical point of view.
This applies to the majority of the plants used
by the sharnan and to plants that are used accorclins to the Doctrine of Signature. It is also
clubious whether plants that are used for topical
treatment of- symptoms of inner organs have
any measurable effect. Yet, even when all these
apparently physiologically inactive plants are
excluded, a large number of plants still rernains. Several hundred in the case of the Coaiqueres, the Cayapasand the Colorados. lJsually,
it is not economically feasible to analyse all
these plants for active compounds. Instead, the
ethnobotanical data should be screened against
a number of additional criteria. The plants that
comply with most of these should be given highest priority. Table 82 shows a list of species that
remained after screenins the collections in the
present stucly fbr physiolosically active plants.
The indirect criteria used were: plants collected
repeatedly with same inforrnation; plants used
separately and not as part of a herbal mixture;
plants provided with precise information on usaee, dosage and contraindications; plants with a
usase pattern that apparently has been independently acquired by lineuistically or geographically isolated cultures; plants indicated as
being poisonous.
Evidence of past cultural exchange
with indigenous groups outside
Coastal Ecuador
Exchange of knowleclge is an important variable that determines ethnobotanical practices
in an intricate manner. The regional distribution of ethnobotanical practices partly reflects
culttrral processes of the past. Carefully con-
30
BS 46
Stout 1948, Trupp 1981), as well as among the
Cunas further to the north in Panama and
nearby Colombia. Furthermore it is noticeable
that both the Cayapa and the Choc6 Indians
include antique Spanish sables among their
preferred paraphernalia. With the Coaiqueres
this kind of exchange is more unlikely. According to Kempf (1982) most households have at
least one male member that is trained as a shaman and often as a matter of social obligation.
Curing rituals
The training is rather informal, comprised of
Travelling and migrating shamans often play
instruction lasting only three days. Curing is reethnobotanical
an active role in spreading
garded as bothersome work and the shamans
knowledge. The curing ceremonies of the ethdo not enjoy any special status.
nic groups of Coastal Ecuador are dynamic,
The Cayapas share some elements of the decultural features. Many similarities between the
scribed shamanistic rituals both with the Colopractises of the different peoples reflect recent
rados and with the Otavalos, who are geographcontacts, and rituals can change dramatically
ically the nearest indigenous people in the Anin just a few decades as exemplified by the Coldean highlands. This applies particularly to the
orados. On the other hand, some similarities
use of paraphernalia and the way that these are
between the practises of different peoples may
handled during the healing sessions.Colorados
also reflect century old contacts, or even com(Hagen 1939) and Otavalos (Lopez 1986) also
grouP.
one
ethnic
mon descent from
organise their paraphernalia at a low table in
Comparison of the curing rituals performed
front of them, and they attach special importoday with those described in the anthropologtance to rounded, black river stones of volcanic
ical literature of the beginning of this century
origin. All three peoples believe that these
is a way of testins whether ritual healing and asstones house powerful spirits useful to the shasociated plant uses are stable features that
man
as tutelary spirits. This reflects that volcarathcharacterise the various communities, or
noes are important in their mythology. During
er that they are a dynamic cultural feature that
the healing rituals that we attended among the
varies in both time and location. An additional
Cayapas the name of the vulcano Imbaburawas
benefit would be insight into the way new plant
repeatedly used as a conjuration. Imbabura is
Barrett
a
community.
by
uses are implemented
the nearest r,'ulcano of the western Andean cor(1925) noticed that a number of Choc6 shadillera and it is considered a sacred mountain
mans practised among the Cayapas in l90B-9.
by the Otavalos. According to the mythology of
The direct contact probably ceased more than
the Colorados numerous spirits reside in the
a half century ago. There exists evidence of
of Cotopaxi and Chimborazo south of
vulcanos
sessions
curing
During
today.
Choc6 influence
The way that all three groups organImbabura.
staff
Cayapa shamans still possess a hardwood
ise their alter and worship the vulcanoes may rewith a carved wooded figurine. The staff is conflect pre-colombian contacts between the ansidered to house their personal tutelary spirit.
cestors of the Cayapas, Colorados and Otavalos
Wood figurines are otherwise unknown among
but
possibly ata time when the former two groups
people,
Amerindian
Ecuadorian lowland
nearer the highlands and when they all
lived
very similar figurines are standard paraphermay have spoken related Chibcha languages.
nalia among Choco shamans (Wassen 1935,
ducted comparative analyses could contribute
to an understanding of the relative roles of endemic ethnobotanical elements versus diffusion of knowledge across ethnic barriers. Usage patterns that cannot be predicted from inherent f.eatures of the plant and that occur repeatedly across cultural barriers, strongly suggest past exchange of knowledge.
BS 46
Curing rituals and certain ethnobotanical
patterns suggest the presence of both ancient
and recent contacts between the cultures in
Coastal Ecuador. The Cayapas currently interact with the African Ecuadorian colonisers who
have settled throughout their territory. This
ethnic group constitutes a majority in the Ecuadorian Province of Esmeraldas. The first
Africans fled from slavery and miserable livins
conditions and founded colonies in the coastal
areas of western and northern South America.
In Ecuador, they settled in the northern part of
the coastal plain. The colony continued to attract emigrants of African origin even after
slaverywas abolished.
The curing ceremony that we attended in
1983, took place in a fairly isolated Cayapa
community. The rituals that we saw performed
have probably changed little since the begining of this century. In contrast, a comparison
with older and more recent literature suggests
that the Colorado rituals have changed dramatically. Karsten (1924) described a nocrurnal curing ritual in which the shaman used
three black stones, a drum, a rattle, and a number of candles. Everybody who attended drunk
nepe(the Colorado name for Ayahuasca) and
danced around the patient. At the end of the
session the shaman presented a spine of the
chonta palm (Bactris gasipaes) and claimed he
had extracted it from the patienr. The spine
was supposed to be a magic arrow that had
caused the disease. Magic arrows are important
in the mythology of indigenous highland people, e.g. the Otavalos, and this ritual also suggests early Colorado contacts with the highland. Since then the Colorados have integrated
additional practices from highland indigenous
healers, e.g. to rub the affected area of the patient with an egg in order to transfer the malevolent spirit from the patient to the egg (Trupp
r e 8 1. )
In recent decades, so-called warm and hot
plant baths have become an important ele-
31
ment of Colorado curing ceremonies. Earlier
authors reporting on the Colorados (Hagen
1939; Karsten 1924; Rivet 1909) do not mention this practise which apparently has developed since. The warm baths are similar to the
steam bath that was demonstrated to us by the
Cayapa Amerindians, whereas the cold baths
often take place in small ponds close to the river (Lopez 1986). The steam bath rirual was
probably introduced to the Cayapas by the Colorado shamans very recently. We only observed
it in Zapallo Grande which has extended contacts with the exterior. In remote and isolated
Cayapa communities such as along the Rio Bolborde we never heard of such practices.
The exact origin of the plant baths is uncertain but again it may have been inspired by
contacts with indigenous healers in the Andean highlands. Illness is generally explained
by a disharmony between the cold and the luarm.
principle. To stay healthy, highland indisenous
people must maintain a balanced diet of items
considered as cold and u)arm, respectively. The
Colorados and the Cayapas have adapted only
the part of this philosophy that relates ro the
bath treatment of certain ailments.
The African Ecuadorian population on the
coast also have their own shamans, and live
intermixed with the Cayapas along the lower
parts of the Cayapariver sysrem. Barrem (1925)
noticed that in these communities shamans
with a good reputation occasionally treat patients of the other ethnic group, and this is still
the case. To what extent this has influenced the
Cayapa curing rituals is unclear, but one introduction from the black population is the marimba
a keyboard instrument - which is
found in many Cayapa houses, as well as
among Colorados and Coaiquer. Apart from
this example, it is difficult to trace how the Coaiquer rituals have been influenced by other
groups and vice versa. The Coaiqueres do not
use hallucinogenic plants during curing ceremonies, but everyone present drinks trago
fF
32
(sugar cane brandy). This distinguishes them
from all other Amerindian sroups in Coastal
Ecuador (Kvist & Flolm-Nielsen 1987; Naranjo
1983 ). D a n c i n g to ma ri m b a mu si c i s an i mportant part of Coaiquer curing cerernonies
(Kempf 1982). In seneral, the function of the
shamans is less specialised compared to the
Cayapas and the Colorados. According to Ehrenreich ( 1989) , skilled shamans were not
among the Coaiqueres that settled in Ecuador
at the beginning of this century. In the San
Marcos valley, one adult male member of almost every householcl has been trained as a
s ham a n (K e m p f 1 9 8 2 ).
BS 46
part of the explanation. Other species <-sfCo
luntnea and Gesneriaceae serve the same purpose without har,'ingvariesated leaves.The Amerindians rarely make mistakes when distinguishinu between the Gesneriaceae and other
morphologically sirnilar sroups such as the
Acanthaceae that are not used fbr treating
snake bites. A number of Gesneriaceae are
used for treating snake bites by the African
Ecuadorian population that lives on the pacific
coast of Colombia along Rio Tataboa west of
Cali (Kvist, unpublished dara fiom lggl). The
Choc6 and Cluna Amerindians of western Colclmbia and Panama have been reported to use
some C esneri aceae too (D uke 19 70, lgTb:
Botanical euidence
Forero P. 1980). This is in contrast to the AmerA few examples of botanical evidence of old
indian groups of the Amazon resion who rarecultural connections exist. This applies in parly use Gesneriaceaefrrr snake bites (Kvist l986,
ticular to some domesticated plants. P.sychotria 1989). Two Gesneriaceae were clairned to be
uiridis is cultivated by the Cayapas who mix the
particularly valuable. One is Chrysothemisfrieleaves with the stems of Banesteriopsis caapi
dnchsthaliana,which is used in snake bite therawhen they prepare pinde (ayahuasca).It is not
py by both the Cayapas in Ecuador and the Cunative to the coastal resion of Ecuador and
na and Choco Amerindians of Panama (Duke
may have been introduced from Amazonian
1970, 1975). The other one i s a hybrid of KohEcuador (Rivier & Lindsren 1972). Psyr:hotria leria grown as a febrifuge both by the Colorauiridis has hallucinosenic eff'ects in itself and is
dos and the Cayapas. It is widely cultivated in
used for this reason by Amazonian Amerindinorth-western South America.
ans. The fish poisons Clibadium asperum and
Plants that are used for poisonins fish are
Lonchocarpus nicou may also have been introgenerally known in South America as barbasco.
duced to the coast. The Colorados grow both
The little known barbasco, Saltium peruuianum,
plants that according to Murra (1948) were
is shared between the Colorados and Coaiintroduced from the Amazon region alons
queres. All the Amerindians of Coastal Ecuawith other ichtyotoxic plants.
dor use Phylktnthus anisolol.zswhich serves the
The importance of the Gesneriaceae fbr
same purpose. Species of Phylkntthus are senertreating snake bites is puzzling. In the case of
ally used for fish poison in South America. Onthe genus Columnea the Doctrine of Signatures
ly the Colorados use Lonr:hocarpu.ssNt.and Clibamay provide some of the explanation. The repdium sp. as barbasco. They are often cultivated
resentatives of the largest section of Cloastal
around the dwellinss. Both species have unEcuador all have leaf apices and leaf margins
doubtedly been introduced frorn the Arnazon
that are bright red on the lower leaf surf-aces where they are comrnonly used for fishing. A
(Kvist & Skog 1993). The Amerindians draw a
cornprehensive discussion of fish pclisons or
parallel between these leaf variegation patterns
barbasco amons South American Indians can
and the bites of the haemolytic vipers which ofbe firund in Acevedo-Rodrisuez (1990).
ten cause excessive bleeding. But this is only
The plant uses classified under social prod-
c)c)
JJ
BS 46
has
ucts generally suggest that little exchange
of
takeriplace between the Amerindian groups
exCoastal Ecuador. A f'ew plants are however
revast
a
throughout
ways
ploited in similar
from
caapi
gio". This applies to Banesteriopsis
as
*fri.n the hallucinogenic decoction known
Ayahuasca in Quichua is prepared' Ayahuasca
drinking is widespread among Amerindian
(Rivier
people in north-western South America
add
not
do
it Li"agt en 1972). The Colorados
preparations
other plants to their Ayahuasca
to
whereas the Cayapas mix in plants allegedly
adjusttheeffectortoremovethebittertaste.
Oie of the plants that the Cayapas use in this
in
way is Psyciotria uiridis. Only the Cayapas
hallucinoa
coastal Ecuador use Brugmansictas
is
g e n i c plant . T he r an q e o f th i s u s a g e p a tte rl r
iestricied to people living in or near the Andean cordilleras (Lockwood 1979) ' It is a very
in
powerful and dangerous drug that is used
by
performed
ionnection with certain rituals
shamans.All the Amerindian groups in Coastal
Ecuador use Brugmansia for medicinal purhowever (see Tables 36, 52, 53' 55' 72) '
poses,
^several
Amerindian people in western AmazoniaproduceanextractfromtheseedofPersea
amuricanawhich is used as a contraceptive or
sometimes,as a sterilising agent (Kvist & HolmN i e l s en1987) .
Exchanges between the Colorado' Cayapa
in
and Coaiq.t.t Amerindians are suggested
to remove
particular by the use of Streptochaetct'
iacial hair (the former two groups), the use of
ferns to promote hair growth (all three
groups), the use of ferns mixed with fat for
iood (the latter two groups) and the use of Sapiumas a fish poison (the former two groups) '
The use of numerous plants and particularly
the Gesneriaceae as snake bite remedies also
unite the three Amerindian groups of Coastal
Ecuador. The African Ecuadorian populations
along the Pacific coast to the north and to a
lessei extent the Choco and the Cuna Amerindians share this practise too' Regional exchanges along the Pacific coast, from Ecuador
and north to Panama, are also indicated by the
use of the latex of I'{aucleopsisfor arrow poison
and by the cultivation of Piper tricuspe that is
used. to kill lice etc. Regional contacts across
the Andes to the western part of the Amazon
Basin are suggested by the use of several hallucinogenicsandfishpoisonsaSwellasthetrseof
Persea americana for contraception' Finally' a
considerable number of medicinal plants used
by the Amerindian people in Coastal Ecuador
are also widely used throughout northern
South America. Some of these were mentioned
by Kvist & Holm-Nielsen (1987): Aclepin'scurassauica, cassia reticulata, chlorophora tinctoria, Ficus insipid,a, Jatropha curcas, Paspalum conjuga'
tum, Sciparia, dulci.sand Zingiber officinale '
F
34
BS 46
I'II General conclusions
A comparison of the results obtained from the
three sroups of coastal Indians in Ecuador is
complicated. Not only is the level of ethnobotanical knowledge variable within a single indigenous community, but certain aspects may
even differ within the same village or across
senerations in the same household. The sample size is too small to give a precise description
of the enormous variation encountered. More
information is needed on ethnobotanical practices within-communities and between-communities. A number of general conclusions can be
inferred from the results obtained, however.
Certain usase patterns are likely to develop
due to inherent features of the plants. This is
certainly the case with plants exploitecl for
their chemical, nutritional or structural properties. Experiments with the surroundins flora
are constantly being conducted. New fiuits are
discovered; previously unknown medical properties of a plant are realised; fibre plants are
tested that have not been used before; etc. The
present study clearly demonstrates that independently acquired knowledee is an important
determinant of usase patterns but that exchanse of knowledge across linguistic barriers
is also important. Knowledge that is senerated
within a narrow social context, for example
based on experiments, will add distinctive features to an ethnobotanical pattern, whereas
knowledge that has been exchansed acrosscultural boundaries often will have the opposite
effect. It is surprising that more exchange apparently has occurred at the regional level than
locally between three geographically adjacent
and culturally related ethnic srolrps.
The category to which most uses have been
referred overall is topical treatment of snakebites. Other important usase categories are
topical treatment of ailments of a general na-
ture such as miscellaneous pains and indisposition. The Colorados use numerous plants in
baths that aim at resulatine body temperature.
Palms constitute the most important eroup of
plants exploited for their structural properties.
Most species are used fbr multiple purposes.
The importance of palms rapidly decline with
increasing acculturation and integration into a
market economy. Three sroups of plants invariably rank amons the fbur most important
based on an utilitarian criterion: vascular cryptogams, Piperaceae and Gesneriaceae. These
families are mainly exploited ethnomedicinally.
The desree of congmency between the
plants listed under given usase catesories for
different indigenous sroups increases with the
taxonomic level chosen fbr the comparison.
\Arhereas less than five percent of the plants
rouuhly estimated are shared between the
three sroups at the species level, similar patterns appear when comparisons are conducteC
at the family level. We have identified a wicle
range of usase patterns at this level rangins
fiom very constant in both time and space to
recently acquired and quite labile. This applies
particularly to the ethnomedicinal applications. The usage patterns of medicinal plants
are a function of both cultural variables and
variables inherent to the plant. For plants with
obvious beneficial eff'ectson an ailment, a use
will have a better chance of developing and can
more or less be predicted in commlrnities that
constantly experiment with the surrotrnding
flora. For species that are used as part of a cultural pattern, other f-actorsinfluence the plants
chosen for curing. Common species are more
likely to be used than rarer species. Terrestrial
herbs are lnore likely to be used than e.g. epiphytes because the latter are more difficult to
BS 46
35
collect. \Arhereas some plants apparently are
collected more or less at random others are
collected according to the The Doctrine of Sisnature which invariably adds a certain desree
of predictability to the selection of medicinal
plants.
The plant usage patterns of the three Amerindian groups studied stronuly reflect their
present cultural and socict-econonlicsituation.
The Coaiqueres live in an undisturbed environment surroundecl by a very diverse fbrest.
They have been isolatecl from clther cultures
until recently. Their ethnomedicine is apparently less constant and rnore experimentine
when compared with the other Amerindian
groups studied. Althoush the material culture
of the Coaiqueres seems simple, they have
thoroush ecolosical insight that enables them
to manage a fiasile ecctsystemin a sustainable
manner. The Colorados represent another extreme. This sroup is senerally well intesrated
into the Ecuadorian society.They live in an area where most of the forest has been cleared.
Very little remains of their orisinal material
culture. Colorado ethnomedicine is elaborate
ancl is based on a sophisticated classiflcation
system.A number of shamans and sharnan apprentices practice outside clf the comlnunity in
most regions of Ecuador and even abroad.
These have probably included several exotic
elements in the orisinal ethnomeclicine. The
ritual curins resembles that of the Qrrichua
speaking tribes of the Andean highlands. This
applies especially to the body temperature curing system. The Colorados deviate in many aspects from their indigenous neighbours in
their ethnobotanical practices.
Most Cayapas are settled in villages along the
Cayapa and Santiago rivers. Their territories
were recently opened to the operation of loeging companies. Impoverishment of the plant
resources is occuring at an alarming rate. The
ethnobotanical knowledge basis varies from
one community to another. The Cayapas we
worked with had an intimate knowledge about
the plants that surround them. Their material
culture has chaneed little since Barrett described it in 1920'ies, but it is in the process of
disappearins. Cayapa ethnomedicine and the
naming of plants is just as sophisticated as that
of the Colorados.
Several thines prevent us fiom makins conclusions on the relative roles of cultural variables and plant inherent variables on ethnomedicinal practices. Basically we clo not know
whether a plant has the allesed effect or not or
whether it is more powerful than any other
randomly selected plant. We often assume that
people use the plants that serve their prlrpose
best. But this may not be the case with many
medicinal plants and plants that are used durins curing ceremonies. The opposite is also
possible
that otherwise useful species are
avoidecl because of taboos.
VIII Acknowledgements
Without the unconditionitl lrelp and wArn]
frienclship off-erecl by the informants nlentioned in Table 1 this stucly would not have
been possible. It should be stressedthat all in[ellectual rights to thet infitrmation presentecl
in this paper rernain with the indieenous communities in Ecuador. Mr. and Mrs. Rev. Meisenheimer are thankecl fbr accomrnoclation r,vhile
stayinu in Zapallo Grande. We also thank our
fielcl companions Dorte C. Nissen, Laurrtz. B.
36
Holm-Nielsen, Eduardo Asanza and Flemming
Skov who, besides their dedication to the fieldwork, encouraged us all the way. We thank Dir.
Oluf Holm for establishing the first contacts
with the Colorados and for maintaining an ever so strong interest in our work. He has contributed with many interesting discussions in
his office at the Anthropological Museum in
Guayaquil. The work was supported financially
by Museo Antropologico in Guayaquil and the
Department of Systematic botany at Aarhus
University which is gratefully appreciated. We
received valuable advise on medical matters
from the phycisian Dominick Rizzi. Pamela
Hall commented on the manuscript and Denise San chez Barfod provided linguistic assistance. The abstract was translated to Spanish
byJuan Manual Macia.
Thanks is due to the following experts who
helped in the identification of the collections
(appearing in alphabetical order): Acevedo, R.
(Sapindaceae), Alverson, B. (Bombacaceae),
Andersson, L. (Heliconiaceae, Marantaceae),
Barnaby, R. (Menispermaceae, Caesalpiniac eae ), B e rg , C . C . (Mo ra c e a e , U rti caceae),
Boom, B. (Rubiaceae), Callejas, R. (Piperaceae), Croat, T. (Araceae), Daly, D. (Bursera-
BS 46
ceae), D odson, C . H . (Orchi daceae), Eliasson,
U. (Phytolaccaceae), Eriksson, R. (Cyclanthaceae), Escobar, L. (Passifloraceae), Frp<ell, P.
(Malvaceae), Gale, N. (Rubiaceae), Gentry, A.
(Bignoniaceae), Harley, R. M. (Lamiaceae),
Holm-Nielsen L. B. (aquatic plants), Huft, M.J.
(Euphorbiaceae), Hunt, D. R. (Commelinaceae), Il ti s, H . H . (C appari daceae),.f @r gensen,
P. M. (Passifloraceae), Judziewicz, E,. (Poaceae), K napp, S . (S ol anaceae),K ui j t,J. ( Lor anthaceae), Lrpaard, S. (Poaceae, Cyperaceae),
Leeuwenberg, A.J. M. (Apocynaceae), Liesner,
R. (Lacistemataceae), Lourteig, A. (Onagraceae), Luteyn, J. (Ericaceae, Campanulaceae),
Luther, H. (Bromeliaceae), Maas, P..1.M. (Zrngiberaceae), Madsen, J. (Cactaceae), Miller, J.
(Myrtaceae), Moran, R. (Pteridophytes), Mori,
S. (Lecytidaceae), Nissen, D. C. (Pteridophytes), Ollgaard, B. (Pteridophytes), Palacios, W.
(Meliaceae), Pedersen, T. M. (Amaranthaceae), P enni ngton, T. D . (S apotaceae) ,PI owman, T. (E,rythroxylaceae, Solanaceae), Renner, S. (Melastomaceae), Stolze, R. G. (Pteridophytes), Stihl, B. (Theophrastaceae), Taylor, C. M. (Rubiaceae), Todzia, C. (Chloranthaceae), Wasshausen, D. (Acanthaceae) , Zardrnt,
E . (Onagraceae).
IX Figures r-20
Photos bv A.S. Barfod and L.P. Kvist
rn
o\
o\
!
,9
-'
o
v
€
L
Maclovio Anapa (left) and the shaman Maritimo involved in an ethnobotanical discussion. Their contribution to this studv was invaluable.
-
BS46
39
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u p
ri
tr./n
\\Y
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/ 4 . > \
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9
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;: '*
-
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x n
< O :
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Fig. 2. Barfod & Kvist, februar 1995
structural
riu"E
x<
materials
sanitaryf
chemical
landuse
social
materials
products
ritualf
complimentary
animal
food
porson
digestive
external
medicines
muscular- skeletal
reproductive
respiratory
sensory
urinary
systemic
general
topical
Tlmber (t. 2)
Fuels(t. 3)
Weapons and uaps (t. 4)
Muslcal lnstruments (t. 5)
Kltchen utenslls (t. 6)
Unprocessedas rope (t. 7)
Thatching and wrappers (t. 8)
Jewelry (t.9)
Plaltlng and textlles (t. f 0)
Dlptlatorles (t. 1f )
Latex (t. 12)
Soap and shampoo (t. f 3)
Perfumes (t. 14)
Dyes (t. 15)
Ornamental plants (t. 16)
Hedges(t. 17)
Halluclnogens (r. 18)
Antiferttllty agents (r. 19)
Rltual ornaments (t. 20)
Maglc plants (t. 21)
Pedagogy(t.22)
Staple dlet (t. 23)
Edlble frults and seeds (t. 24)
Vegetables(t. 25)
Edtble flowers (t.26)
Larvae from palms (t.27)
Beverages(t. 28)
Food flavourlng (t. 29)
Food colourlng (t. 3O)
Fodder and flsh batt (t. 31)
Wtldllfe atrracrants (t. 32)
Utllttarian polsons (t. 33)
Notorlous polsons (t. 34)
Anaemla (t. 35)
Blood clrculation (t. 36)
Gangrene (t. 37)
Blood tapplng (t. 38)
Gums and teeth (r. 39)
Llver and spleen (t. 40)
Stomach lnfectlons (t. 41)
Antlemetics (t. 42)
Laxatlves (t. 43)
IntesUnal parasltes (t. 44')
Deflclency symptoms (t. 45)
Ant bltes (t. 46)
Insect bltes. General(t,47)
Snake bltes. Topical (t. 48)
Snake bltes. Systemlc (t. 49)
Snake bltes. Others (t. 50)
Wounds (t. 51)
Swelllngs from brulses (t. 52)
Abcessesand tumors (t. 53)
Fungal lnfectlons (t. 54)
Skln reactions (t. 55)
Rheumatlsm (t. 56)
InJured Jotnts (t. 57)
Nervous system (t. 58)
Menstruation (t. 59)
Btrthglvlng (t. 60)
L a c t a t i o n( t . 6 1 )
Nosebleed(t. 62)
Pulmonary dlsease(t. 63)
Resplratory dlsorders (t. 64)
Colds and lnfluenza (t. 65)
Eye lnfectlons and vlslon (t. 66)
Ear lnfectlons and hearlng (t. 67)
Urlnatlon (t. 68)
Kldney (t. 69)
General. Systemlc I (t. 70)
General. Systemlc II (t. 71)
General.Toplcal | (t.72)
General. Toplcal II (t. 73)
General. Toplcal III (t. 74)
Fig. 2. The usage categories used in this paper and their hierarchical orsanisation. ln parentheses is referred to the Table fbr the usase category in question.
BS46
41
b r i
#
ffi
w
3;t
ffi;Wr*
Fig. 3. Cayapas. - A. Zapallo Grande. Cayapa children playing with the children of semlers.The vast majority of the population in the Province of Esmeraldas are of African origin. The Afiican Ecuadorians settle ever more deep inside the Cayapa territory. In most communities, the two ethnic groups live in peaceful coexistence but social friction's do surface at times - B. A traditional Cayapa dwelling on posts. Note the palm
thatch and the open design. The only room with walls in the hut is fbr sleeping. - Cl. A modern Cavapa
house at the evangelical mission station in Zapallo Grande. The architectural design corresponcls to that
fbund elsewhere in rural Ecuador. Note the zinc roof and the general emphasis on privacy, quite unfamiliar to traditional Cayapa culture.
,
F-
42
BS 46
Fig. 4. Coaiqueres. - A. The forest surrounding the Coaiqueres in the Ecuadorian Province of Carchi is very
humid and species rich. This picture shows one of the wettest localities on the San Marcos Gualpi Bajo trail
in 600 m's altitude. The structure of the forest is characteristic having few tall, large diameter trees. The
trees are dispersed in comparison to typical lowland forest and heavily loaded with epiphytes. - B. Don Ignacio, Capitan of the Coaiqueres in the San Marcos valley where the largest concentration of Ooaiqtreres
south of the border with Colombia is found. - C. The remnants of presumablv an outpost erected by the Incas in Quinyul. This may indicate the western limit of the Inca occupation in Carchi. - D. View from rhe
highest point at 1000 m's altitude on the Pailon-(]ualpi Alto trail. In the foreground is a Coaiquer dwelling.
BS46
43
A
Fig. 5. - A. Pluvial fbrest at darvn near Gualpi in the Coaiquer territory. - B. Timber harvesring along the
Rio Cayapas and its tributaries. Independent lumber men harvest timber far up in the Cayapa river system
and float the trtrnks downstream to the sar,vmillsnear Borbon tied together in eigantic rafts. Selective
cutting of timber trees occurs illegally within the limits of the Cotocachi-Cayapa National Parc. - C. Road
builcling 5 km rvest of Lita. This road has opened new land to Lrncontrolled colonisation and conversion of
the forest into fielcls and pastures.
V
44
BS 46
Fig. 6. Plants used for their structural properties. - A. The house of the canoe builders in Zapallo Grande,
a mixed Cayapa-African Ecuadorian community near an evangelical mission. The canoes in fiont of the
dwelling are made of a yet unidentified species of Lauraceae called djuin-chi and a species of Protium called
.supkt-chi.- B. The Coaiquer Indians frequently cross the Rio SanJuan usinp; rafts made of Ochroma,pyramidale. The river constitutes the border with adjacent Colombia. - C. Sapotaceous species are used fbr firewood by the Coaiqueres and the Cayapas.The wood is flammable when green. - D. The Coaiqueres still use
blowguns when huntins canopy species such as monkeys and many species of birds.
45
BS46
Thatch
,/
n^t^.L^^
rEd
vardu
^^
)P.
Walls
lriarteadeltoidea
Floor
lriafieadeltoidea
/
Ladder.
Bambusa guadua
-quinaria
posts
Carapaguianensis
Thatch
Phytelephasaequatorialis
Walls
Bactrisgasipaes
i
l
Walls
B a m b u s ag u a d u a
Fig. 7. - Plants usecl for their stnrctural properties. Clonstnrction. Three types of huts all constructed on
posts. The upper one is the tr:rditional Coaiquer dwelling with Cakttheathatch. The midclle one is the traditional Cayapa hut with palrn thatch. The open desiun is characteristic of both of these. They are typical of
'fhe
architecture of tire bottom Cayapa hut
large larnily trnits that live scattered ir-rthe forest or along rivers.
is the result of lif-e in a village situirtior-r.The desisn is more private and a door is present. The traditional
thatch is maintained. Barnboo rvalls like those seen here are frequently' found in the hottses of the African
Ecuadorian settlers too (drawings bv K. Thomsen and K. \{brrn).
BS 46
46
Fig. B. Plants used for their structural properties. - A. The Marimba is common among all the indieenous
groups of Ecuador. It lr,asprobablf introduced via the African Ecuadorians from the Caribbeans. The keys
are made of the hear,y and durable wood frorn certain palm species and the tubes underneath anrplifying
the sound are produced from the internodes of Bambusagundua. - B. The Cayapas use Cttrclulouicapalmnta
for a wide range of plaited items that are used in daily life such as containers, mats, fans etc. Thev also prodtrce handicrafts that are sold to tourists. - C. Phytelephasaequatorialis thatch. The leaf rachis is split longitudinally and the half.s are placed on top of each other and fastened to the rafters with pieces of liana. The
pinnae are sometimes braided in ceremonial houses. - D. Fibres are extracted from a leaf of Aechmeamttgdalena usins a stick or wooden mallet. They are used to make a fine bttt strong string that is very suitable for
fishinq nets.
BS46
Fig. 9. Bark cloth (A-C) and Latex (D). - A. The moraceous species Poulsenia annata is usecl ro make bark
cloth. The fibres are clerived from the inner bark as shown on this picture. - B. After extraction the fibres
are freed from the bark b1'poundine, usually with a wooden stick. - C. Bark cloth as shor,vnhere is treated
like f'elt. Several pieces c:rn be joined simply by putting one piece on top of the orher ancl pouncling them
rrntil thev become inseparable. - Latex. D. A latex is extracted from the inner bark of Ca.stilh elu.sticu(Moraceae). It is usecl fbr r,vaterproofing of canoes and for making blowsun pipes airtieht.
47
r-
v
48
BS 46
Fig. 10. Sharnanism and ritual healing. Cayapa. - A. The shaman's altar. Note the nicely arranged iterns that
supposedly house the porverfirl tutelary spirits. In the foreground is the bottle containing pindi. The patient
is sleeping on the floor surrounded by his family members after a nocturnal healing. B. A shaman assistsar
a curing ritual perf<rrmed in the morning on a riverbank. He does not participate directly in the healing
but sits at a clistanr:ewhile mernbers of the family perfbrm the rituals. The shaman is surrouncled by his tutelary spirits which dwell in his wooclen cane and in the river stones in the basket. C. The Cayapas draw a
parallel between the stomach ancl the river; both systemstransport matters in an unidirectional, closed way.
PIants that grow near the river and especially rheophytes are considered beneficial for stomach conditions.
As part of the preparation the plants are soaked in a small water filled hole made b,vhand and situated close
to the river. D. Plant material coller:ted bv the shaman fbr the riverbat-rk ritual. It is usecl for a decoctions applied to the patient and fbr nlassase. The plants allegedly have no effect lr,hen usecl by layman.
BS46
Fig. 11. Coaiqueres. Asricrtlture. - A. Slash-and-mulch fielcl about eight months after clearing. (lorn has
been sown directlv in the up to one rneter deep brushrvood and the scatterecl plants are now visible. In the
backsround is a banana fielcl. Note also the sinsle individual of the palm tree Wcuiniu quinariathat has been
left for later use as firnber in h<tttseconstnrctions. - B. Sugarcane is often cultivatecl by tlie Coaiqueres. A big
part of the harvest is used to make a f'ermentecl drink called uoaripo. This can be further processeclinto trago or brancly in primitive tvooden distilleries. Here two men share a clrink early in the morning in San Marcos. Behind them, in fiont of the church, there is a susarcane fielcl. - (1. Clearing made frrr slash-and-mulch
asricttlture. Species <>fVismialvith whitish or f'errtrgineons trndersicles of the leaves are rvell represeltecl on
this plot probably becartse the forest is in a vorrng succesional stase after a previous clearins.
19
50
BS 46
Fig. 12. - A. Clusia sp. (voucher no. 48932) is one of the many plants referred to as uikllifb ttttract,ants(see Table 32) by the Coaiqueres. Birds feed on the fruits and thereby act as the clispersal agent. The categor)' contprises both plants attracting game and plants interacting with animals in a particrtlar way. B. The secds of
the cycad Zamia lindeniior sn-oa,ltrt-chiareground into a flour used for a special kind of breacl bv the Ciai'apas. - Cl.The larvae of the Curculionid beetle, RhynrhophorusNtalmarun, is considererl a grcat delicacy by the
Cayapas and the Coaiqueres. It is collected on the partly decomposed stem of certain species of'palms such
as lriarteaancl lluctns. -D. IriurLeadeltoideaisa truly multiple purpose palm. It serves as t.irnber for constntction, traps, lveapons ancl rnusical instruments. The palm heart is eaten ancl, edible larvac are collecteri fiom
decomposing stems.
BS46
II
52
BS 46
-b'r,t
Water So that fish fhaf suri-r dn',n
-*_=_-
.rr.,"t,ffi;ffiffi
.i."Ji".lf:::Hiff[Ti:,:l*:::::::i.i:'.1]1"1.'lJ.;;;ffi;;ffiin."J"lffi'Ji
3fi1'J;Jll1,''xlTo"lil1j:T'_":l*',::::i.fij::r:il;;rl;;i.;:#;'::?#ii,
The
il:';ff
fish f:','H:#,L1,"."1;;,i:.1:i:::il,y::,:1..:il;:JlT,1fi,;;ilili51fii
rrap shownin rhe pi.,.,r" is useclro c.,ecr
,n. p;,;;;ilJJ:il?::
clam.
F
tl-^
outlet in the far end of the
BS46
53
Fig. 15. Medicinal plants. Prerparation. - A. A nurnber of'plants are used to treat funeal infection, in this
case betrveen the toes. Typically the leaves are grorttrd into a sreen paste that is appliecl clirectly to the skil
either heatecl or cocllecl.- B. Bottles containins alcohol extracts of plants usecl fnr sn;tke bitcs zrrecomrnon.
They are often proclucecl irtrcl solcl b,v certain tribe rnembers t.hat have gained a reprrtation as snakebite
healers. - Cl. A pl:rnt paste ttsed :rs a comprcss to stop bleeclins and to prevent inf'ections in u,otrnds. - D. A
plant decoction is drunk fbr inrligestion.
F'
54
BS 46
Fig. 16. Medicinal plants. Snakebites. - A. (iral (Micrunts.sp.). Several plants:rre gsecl ro treat the bites of
this snake. The poison is ncttrotoxic ancl has an immediate effect Lrnlikc the haemo6,tic p.,isol <tf the r.,ipers. - B. Clasterantlm.s
cornllimts. Representatives of Clesneriaceae are-.generallv trsecl to trcat snake bites.
Thisspeciesistrsedforthellquissnake (Bothropsatrox)lt,t,thcCiavapas.Thelcar,esarernacerateclinto:rpasre
that is used fbr rnaking a conrpress placed on the bite. - C. An liquis snake has.just been killecl apcl the r'zrlevolent spirit is beinq expelletd. This is done bv blorving sntoke on it uncl r.rsing r'11r'r.jtrrati6ps
like rrliilD
which probably mimics the sottnd of it's clepartttre. - I). Oauend.i.shia
ptrntdifitlin.This spgcies oftel clecor.ates
Cayapa hrtts where curing ritttal are performccl bv a shaman. The plant allcuecllv ilcrc;rses cSances e{'strccessftrl healing by oustine malevolent spir-irs.
_"|
BS46
cc
Fig. 17. Medicinal plants. Miscellaneorrs. - A. Palicourea{uiunen.si,.s.
Tl'ris species attracts
rvilcllife especi:rlly' birds zrnd is
usecl for rittralistic tr-eatlnents
of vagnelv clefinecl or nonspecific sympton)s. B. CoLum,nea lililoba. The irsh of this
species is trsed fbr a topical
lrcalment o[- a skirr reaclir.rrr
:illegeclly causerl b,v the plant
itself. - C. Species ctf Dichori.sandrn are used for snake bites, firr firngal inf'ect.ions, fbr
injured .joints, for colds and
influenza
problems.
ancl for trrination
7*
BS 46
56
Verbenaceae
Theophrstaceae
Fig. 18. Barfod & Kvist, februar 1995
Sapotac@e
Pontederiaceae
Phytolaccacea€
Oxalidaceae
Onagraceae
Olacaceae
Myreinacea€
Musaceae
Menisp€rmaceae
Lythraceae
Loganiaceae
Hydrocoaylaceae
Hydrangeac€ae
Cyperaceae
Cycadaceae
Caryophyllaceae
Cannac€ae
Caesalpiniac€ae
Boraginaceae
Araliaceae
Apiaceae
Anacardiaceae
Agavaceae
Sterculiaceae
Sapindaceae
Rutaceae
Myrtaceae
Mim6aceae
Malpighiaceae
Lcythidaceae
Heliconiac€ae
Flacourtiac€ae
Convolvulaceae
Commelinac€ae
Bromeliaceae
Annonaceae
Amaryllidaceae
Amaranthac€ae
Scrophulariaceae
0rchidaceae
Nyctaginaceae
Meliaceae
Marantaceae
Lauraceae
Cucurbitaceae
Bombacaceae
Begoniaceae
Aristolmhiaceae
Loranthacea€
Caricrceae
Capparidaceae
Bixacea€
Bignoniaceae
Ap@ynaceac
Zingiberaceae
Poaceae
Cyclanthaceae
Urticaceae
PNiflorac€ae
Monimiaceae
Lamiaceae
Malvaceae
Fabaceae
Acanthaceae
Costaceae
Meliltomataceae
Moraceae
Arecaceae
Araceae
Euphorbiaceae
Asteraceae
Rubiaceae
Gesneriaceae
Solanaceae
Piperaceae
VNcular cryptogams
I
100
120
140
Fig. 18. Clolorados. Comparison of the most important families of usefirl plants with the composition of'the
surroundine flora. Dark bars show the number of times that representatives of a given familv have been recorded as useful in this study. Note that multipurpose species may contribute more than once to the figures. \A4rite bars show the number of species representing a given family in the Flora of Rio Palenque (Doclson and Cientry, 1978). Only families with more than I ethnobotanical record have been included.
BS46
57
Vibcs€
Rhammc€e
i-r,
Ponl€d€riacec
F
Polygalaca€
Loganlaca€
Haemodomcea€
Dimcorecoe
Cyc.dac€a€
Ch€nopodlaca€
Canmcee
llur*raceae
Agavacea€
n
+
lr
J
Rubcee
I
Phyiolaccacee
Olacac@e
f
F
F
Ir
Nychglmcee
Malplghl,rcea€
Icaclnaceae
Convolvulaccae
Eoraglnacea€
Arali{ceae
Apbceae
Anaerdlaca€
Stercullacs€
Potrulacac@e
Mynlnaceae
Myrlsttcaccae
Lythrace€
Carl€cea€
Aris(olchlacee
Alstrftm€ri,ca€
Oxalldaccae
Menlsp€rmacea€
Rombacacee
Annomceae
L€cythldsceae
Hellconlsc€ae
Cyp€racac
F i g . 1 9 . B a r f o d & K v i s r . l ' e b r u a r1 9 9 5
f1
I
t:
+
+-
tr
H
:
;
fE
Cappsrldaceae
Rlxacea€
Monimlaceae
Campanulacec
Sapobc€ae
Mvtuc€ac
Ca€salplnleca€
Amryllldacec€
Sapindacee
Padnoracee
-_
F
r+
1-------1
l,oranthaceae
Lauracee
Amaranthaceae
Mellaceae
Flacourtla(€ae
Mlmsacea€
Meranbceae
IriBnonlacee
Poac€e
Regohlac€s€
Urticacea€
f=
rt
I
Rrom€liac€ae
V€rb€naca€
Cyclanthrc€e
Cucurblhc€aG
Commelinac€sc
Zlnglh€mc€ae
:-l
-
[-r
Malvacea€
Erlcac6€
Scrophularlacer€
[-t
Orchidaca€
l'ahaca€
Cosbcea€
Apmynac€a€
Momc€a€
Cluslarca€
Acanthacea€
Lamlacea€
i---
i
-l-l
--
:-,
Euphorblacs€
Pep€rohiaceae
Ar€€ceec
Aslcracae
Melastomahc€ae
Rublaceae
Arece€
Solsnoc€se
Piperacee
.
i
F i g . l ! ) . N l o s t i t r r p t l t - t a t t tf a t r t i l i e s .A l l t h l e e i n c l i s e l l o u s s r o u p s . H i s t o g r a r n s h o l v i n g t h e n u r p b e r o f ' 1 s e s h a *
ing been recorded for the representatir,'esof a given family. Onlyfamilieswith more than 1 ethnobotanical
record have been included. Note that multipurpose species mav contribute more than once to the fisures.
Identical use of a siven plant species by two or three indigenous groups is only recorded once.
r-
BS 46
5B
EarinfrcdoN,
heulng
Luadv€s
Food colourlng
Lrtex
Musical lrctruments
Pulmonary disss
Delidency symptom
poisoN
Notorlou
Antiferdlity
agents
Hedges
WeapoB and traps
N6e ble€d
Lactadon
Bl@d dr0ladon
Lsrvce from palm
Thatching, wrspp€n
Fuels
BirthgiYlng
Kldncy
IN6t
blt€s
Andemed6
Halludnog€ro
Ant bltes
Food flavourlng
Ritual ornments
Dy6
Gum
md teeth
Gangren€
Staple dl€t
Maglc plants
and vlslon
Eyelnfetlorc
Snake blte. Othen.
Perfum6
JewelrY
Rsplratory
dlerdem
InJued Jolnts
paraslt6
Int6dnsl
Anaemle
Edlblc floral prrts
Pedsgogy
C€neral. Systemlc. II
Menstrua6on
Abc6s€s,
Utilitarlsn
NenoN
tumors
trplens
system
Fodder snd osh bslt
Urlnsdon
Fugd
lnfedoN
Unprcrss€d
s rop€
Colds and lnllu€nza
Liver snd splen condldoN
Soap, shmpm
Beverag€s
Plaitlng, texdl€s
Generd. Syst€mlc. I
Rheumdsm
Swellings from brulss
Ornamental plants
Snske bite. Systenlc
Vegetrbls
Siomach tnfecdoN
Wounds
Sklnrescdom
Tlmbcr
Cam
attrstNnts
I
G€n€rsl. Toplol.
Gener&I. Toplcal.
F/lble
III
lrulrs
General. Toplel.
II
Snake bit6. Toplcd
60
80
100
120
140
Fig. 20. Most irnportant uses.All three indigenous groups. Histograrn sholving the total number of nses har.
ing been recorded in a given usase category. Note that multipurpose species may contribute to ser,eral usase categories. Identical use of a siven plant species by tr,voor three indigenolrs groups is only recclrded
once.
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BS 46
62
are indicatecl the informants did not elaborate fttrther exTable 2. Timber. For construction ancl carloes. \{rhere no uses
provide details concerning common trses,durability'
comments
cept that the species collected served as a timber tree. The
Note that some of the Anlerindian{Iarnes are
is
included.
name
vernacular
the
Finally
value and preparation of the wood.
(see also Figs. 3-7)'
of mixecl o.igi" with Spanish syllables incorporated
name" [language] / (AAU voucher)
Tribe
Use/comments,/"local
Anacardiaceae
Tapirira guianensis
Cayapa
/valvable/"Sajo
pal/ $8277)
Annonaceae
Guatteria sP. I
Coaiquer
G. sp.2
Cayapa
/ /"Guasca negra" fSpanish] / (41655)
Htrts/ /"Pa-chi" [CaYaPa] / $0736)
Catoblastus aeqrtalis
Iriartea deltoidea
Coaiquer
Posts in huts//"Gualte
Socratea exorrhiza
Cayapa
S. exorrhiza
\Arettinia quinaria
Coaiquer
W. quinaria
Coaiquer
deparar" Ispanish]/ (60002)
Huts,//"Boun-chi" ICayapa] / (60097)
"Pifr-ua-chi"
Huts//
ICayapa] / (60098)
Huts//"Gualte crespo" ISpanish] / (60007)
"Palmira"
[Spanish]
Posts in huts/ /"Ban-chi" [Cavapa],
(41074)
"()ualte
bola" fspanish] / (60005)
/ /
Bombacaceae
Matisia coloradorttm
Colorado
Hnts/ /
D a c r y o d e sg r a n a t e n s i s
Protium colombianum
Coaiquer
/ /
P.tp'
Cayapa
Caesalpiniaceae
Bauhinia sp.
Swartzia sp. I
Coaiquer
S .s p . 2
Coaiquer
S .s p . 3
Coaiquer
/ /"Acorosillo"
Clusiaceae
Clusia sp.
Coaiquer
/ /"Mancha
Arecaceae
Cayapa
Cayapa
Burseraceae
Marila laxiflora
Vismia sp.
genus indet.
Euphorbiaceae
Hieronima chocoensis
Coaiquer
Coaiquer
"D6do"
de arriva" fSpanish],
"Sajo-chi"
fCaya-
[Spanish] / (40015)
"Pvlgande"
[Spanish] / (al 676)
/ /"Anime" [Spanish] / (41659)
Canctes//"Supla-chi" lCayapal / (41006)
/ /"Forda" fspanish] / (41674)
"Palo
chiso" ISpanHuts//"Chiparo"
ISpanish] or
(
4
1
6
0
7
,
4
1
6
5
6
)
ishl /
Posts in huts/last up to 20 years/"Palo chiso" ISpanishl / (al66e)
Coaiquer
Coaiquer
Coaiquer
[Spanishl / (a8789)
ropa" lspanish] / (41662)
(416i5)
/hard,and durable/"Ambour6" ISpanish]/
(48832)
ropa"
/
fspanish]
/ /"Mancha
/ /"Sangrrado" fspanish]/ (al665)
Genus indet.
Cayapa
[Coaiquerl/ G1657)
/ /"Motrlon"
"Nagarichi-tap6"
[Cay aPa) / (40507)
Htrts, canoes/ /
Gl}al)
[Cayapa)/
/ /"Pin-sa-chi"
Walls of huts, never floors/ /"llase-chi" [Cayapa]/ (41038)
Fabaceae
Dussia sp.
Coaiquer
/ /
H. laxifolia
Sapium sp.
Coaiquer
Cayapa
Cayapa
"
Acctronsillo" [Spanish] / (al 656)
BS 46
Tabel 2 - Continued
Tribe
Use/comments/"local name" Ilanguae.e] (AAU voucher)
/
Tetrathylaciurn macrophyllurn
Oayapa
Hippocastanaceae
Billia colornbiana
Placed across tirnber trails to facilitate sliding of'
trttnks/ / / (48425)
Ooaiquer
/ /
Cayapa
Huts, mosr commonly used timber/ /,,OhantLl"
[Spanish],
"Marra
tchape" fCayapa] / (41076)
Ooaiquer
/ /'\ara blanco" [Spanish]/ (4t67t)
"Chachajo"
/ /
fSpanishl / GI67 b)
Partictrlarly oars/ / "Satrda-polo-chi" or,,Djeiva_chi,'
ICaya_
pal / (a1008)
Flacourtiaceae
"Cc>roslllo"
[Spanish] / (41663)
Humeriaceae
Humiriastrunt procenlnl
Lauraceae
Ocotea ira
O. sp. I
Genus indet.
f)avapa
Cenus indet.
Oa_vapir
(]enus indet.
Cenus indet.
Cancres/hard and verv durable/"Djui-ch 1,,
lCavapal /
(11027)
Ooaiqtrer
Ooaiqtrer
/ /"Goaripo" [Spanish]/ (41661)
/ /"Malde" fSpanish] / (41670)
Lecythidaceae
Eschweilera sp. 1
E. sp. 2
Coaiquer
(ioaiquer
Ooaiquer
/ /"'tete" [Spanish]/ (41668)
"Tedpu"
/ /"Tete fSpanish],
[Coaiquer]/
(489911)
Melastomataceae
Blakea punctulata
Coaiqtrer
Genus indet.
Ooaiquer
Meliaceae
Carapa guianensis
Coaiquer
Guarea sp.
Trichilia poeppigii
Ooaiquer
Genus indet. B
Cavapa
Cayapa
Huts/ / / (41613)
/ /"Chicharo" [Spanish]/ (41666)
/last rrp to 30 years/"Aray" Ispanish] / (41667)
/verv valuable/"Bu-chui" IOayapa] / (482t7)
"Chalde "
Hrrts//
fSpan ish] / GI67 2)
Finer rvoodrvorks/ /"Inun-chi" [Cayapal ,,C,eclor,,/
fspanish]
(1822e)
Moraceae
Brosimurn utile
Castilla elastica
Ficus cervantesiana
F. insipida
Cayapa
Oavapa
Czryapa
Oavapa
F. maxima
Cayapa
Perebea xanthochyma
Cayapa
Huts, nrainll'for floors/ / / Gl0Z7)
/ /"Oavchu" [Spanish] / (48214,48994)
Boards or for canctes//"Bi-chi', [Cayapa]/ (40765)
/ / / (40e00\
Cut up fcrr boards//"He-aJa-pi-chi,'
ICayapa] / (4g203)
Canoe paddles/ / " Ya-mu-ki -chi,, [Cayap a]
/ Gg20l)
Myristicaceae
Dialyan thera gordoniaefolia
D. sp. I
D. sp. 2
Cay'apa
D.sp.3
Ca1'apa
Ooaiquer
Ca1'apa
/ /"Oansare" fspanishl/ erc64)
//"Mo-chi" fCavapa], "Cangare" [Spanish] / (402b4)
Floors/very durable/"Mo-chi" ICayapa] (41049)
/
"Chu-:rin-clii"
"Chalveande"
/yalvable /
ICayapa],
(Span_
ishl / (aB2e0)
Olacaceae
Heisteria sp.
Ca1'apa
Minquartia guianenses
Several uses/r,ery |-tard/ "Shtri-1un-eui-chi"
IOava_
pal / (a0358)
C)avapa
All parts of huts/ /taluable and durable/ ,Ouan-acan_chi_ha_
ki" IOavapa], "Guayacan" [Spanisl:l/ (41023,48400)
--Y---
fl
BS 46
64
Tabel 2 - Continued
Tribe
Use/comments/"local
Cayapa
Ceilings, stairways, fences/,/"Ba-ki-tape" ICayapa],
[Spanish] / (48371)
Cephaelis gentryi
Cayapa
Axe handles/ /
Isertia pittieri
Cayapa
Pentagonia sp.
Cayapa
/ /"Tu-main-chi"
"Ma-kari-chi"
/ /
Cupania cinerea
Cayapa
Floor inhuLs/ /"Kelan-boe r-chi" [Cayapa] / (41046)
Sapotaceae
Pouteria torta
Coaiquer
P. collina
Coaiquer
/ /
Cestrum baeniuii
Coaiquer
/ /
Tiliaceae
Apeiba sp.
Cayapa
Canoes/not very valuable/ /
m o n o " I S p a n i s h ] / ( 4 8 19 8 )
name" flanguage] / (AAU voucher)
Poaceae
Bambusa guadua
"Caia"
Rubiaceae
"Tu-main-chi"
fCayapa] / (48108)
[Cayapa] / (40827)
[Cayapa] / (41 035)
Sapindaceae
/ /"Piast" [Spanish] / (41629)
"Caimittllon"
[Spanish] / (al 673)
Solanaceae
"Chalmolan"
[Spanish] / (aI650)
"Han-ap6-chi"
[Cayapa],
"Pefra-
Table 3. Fuels. A few trees were pointed out as being particularly useful for firewood because thev are flamable when green (Fig. 6). A wide variety of other woody species are used as well but they need to be dried first which is difficult under
humicl tropical conditions.
Tribe
Use/comments,/ (AAU voucher)
Cayapa
Charcoal/wood
Cayapa
Fire wood/excellent/
Cayapa
Fire wood, charcoal/flamable when green/ (41017)
Fire wood/flamable when ereen/ (41673)
Fabaceae
genus inclet.
hard,/ (48063)
Mimosaceae
Inga edulis
(48200)
Sapotaceae
Chrysophyllum argentum
Pouteria collina
Coaiquer
BS46
65
Table 4' Weapons and traps. To make a blowsun two narrow boards, 3-5 meters long and
semi-circular in cross section ( 5
X 2.5 cm) are prepared from the outer strongly sclerified layer of the palm stem. The central
bore of the blowgun is macle
by carving a straight furrow on the flattened sides to be joined. The Cayapas use straps of rubber
extracted fron Cnstilla elasticato unite the two pieces of wood and to asslrre that the pipe is airtight. The Coaiqueres
wind a plant fiber tightly around.
the blowgun and rub it with beewax. The beewax is further heated over a fire until black (Fig.
6). For blowgun clarts both
groups use the thick and stiff fibres extracted from the desintegrated leafsheaths of,/esseni.s
bataua. A srnal pellet of kap'k
from Ceibaltentandra is wound around the proximal end of the dart. The orher end is dipped
in plant p,riro. (see Table
33) ' Palmwood is very durable and fishtraps that are constmcted in rivers partly under *ui".
rnay last for several years. For
Cayapafishing techniques see also Mitlewski ( lg85).
Tribe
Use/(AAU vtxrcher)
Bactris gasipaes
Cayapa
B. setulosa
Coaiquer
Iriartea deltoidea
Cavapas
Jesseniabataua
J. bataua
Coaiquer
Blowguns, fishtraps, spears/ (001 I 3)
Blorvguns, fishtraps, spears/ (000 I 0)
Blorvguns, fishtraps, spe-ars,/(48409, 60097)
Blowgun darts/ (60006)
Blowgun darts/ (60079)
Arecaceae
Cayapa
Table 5' Musical instruments. Three palm species in particular are the source of the wood
lsed fbr marimba keys. It is the
strongly sclereified, black and lieary tissues toward the periphery of the stem which are
exploited (Fig. g).
Tribe
Use/(AAU voucher)
Bactris gasipaes
Cayapa
B. setulosa
Coaiquer
Iriartea deltoidea
L deltoidea
Coaiquer
/
/
/
/
Arecaceae
Cavapa
( 6 0 11 3 )
(600I 0)
(not vouchered
(6007e)
Vitaceae
Cissussp.
Cayapa
Fruits drieclwith seedsinside;offered to children as ratr_
b/ G0486)
Table 6' Kitchen utensils. Bowls. Calabash trees are often cultivatecl close to drvellings. The
woody pericarp of the f.rit is
used to produce a number of household utensils such as bowls, scoops, spoons etc. Coconut
shells sometimes serl,e the same purpose.
Tribe
(AAU voucher)
Crescentia cujete
Colorado
C. cujete
Cavapa
(10728)
(40453)
Bignoniaceae
7
BS 46
66
stems, climbing rhizomes, or aerial roots are
Table 7. unprocessed as rope. This categor,v comprises plants of which the
removal of leaves and side branches' This is
e.
g.
is
cleaning
onlv
preparation
ihe
used as rope in heary cluty contruction.
before rlsage'
in c'ntrast to plants of which the fibres are extracted and often further processed
voucher)
Tribe
Part of plant/trse/(AA'U
Oayapa
Rhizome/rope
Philoclendron subhastattrm
P. sp. I
Cayapa
P .s p . 2
P. sp. 3
Oayapa
Carapa
Aerial rooL/ / (48179)
Ae rial root/tving logs together/ (40849)
Aerial root,/string, poor qualitl'/ (48241)
Aerial rooL/ rope / (48403)
Destnoncrts sp.
Cayapa
S t e r n , / r c l p ei n h o t t s e c o n s t r t t c t i o n / ( 4 8 1 9 7 )
Bignoniaceae
Paragotria pyrarnidata
Ca,vapa
Stem/rope for fastening c:rnoes and tyinq logs ttlsethe r l ( 4 8 4 13 )
Bromeliaceae
Pitcairnia sp.
Coaiquer
Rocrts/ strin S/ (al 436)
Cyclanthaceae
Asplundia sp. I
Oai'aPa
A. sp.2
Cayapa
A, sp. 3
Cavapa
Stem/tying logs together in constructions/
Sten/ / (48124)
Aerial root/string, not valuable/ (4837 I )
l.udovia integrifolia
Cavapa
Aerial root/very str()ng rope/ (48281)
Fabaceae
Dioclea sp.
Coiliquer
Stem/tving of logs,/ (4 I 646)
Rhamnaceae
C)otrania sp.
Cayapa
Stem/rope used fbr tving bambo() stelns tosether in cons r r u c t i o n s , /( 4 1 0 8 0 )
Vascular cryptogams
Blechnum volubile
in hottse construction/ $0763)
Araceae
Clayapa
Arecaceae
(40829)
often processecl befbre beins placed orr t h e
Table g. Thatching and wrappers. The leaves of Phltetepha.saequatoriali.sare
ntly to soften the tissues and assure that
apPare
roof. one such treatment consists of intrndating the leaves for several weeks
eugs of unpleasant inse cts (Figs' 7 k
the
of
removal
the
be
coulcl
explanation
the pinnae remain flat upon clrying. Another
B)
Tribe
Use/ (AAU l'ottcherr)
Cayapa
\A'rapping marerial/ / (40886)
Oa,vapa
Cayapa
\A'rapping material/
T h a t c h / ( 6 0 1 11 )
Coaiquer
Thatch/ (no voucher)
Araceae
Anthtrrium asplundii
Arecaceae
Cleonclma sp.
Phyteleph as aequatorialis
Maranthaceae
Calathea sp.
(60004)
BS46
67
Table 9. Jewelry. The vascular bundles of a number of ferns serve as string in necklaces made of seeds, fruits or flowers.
Tribe
Part used/use/ (AAU voucher)
Vascular bundles/string in necklaces,/ (4U399)
Rhizome/for necklaces/ (40828)
Rhizome/for necklaces/ (40899)
Vascular cr;ptogams
Adiantum macrophyllum
Oayapa
A.rp'
Cavapa
Pteris sp.
Tectaria sp
Cayapa
Cayapa
Thelypteris sp.
Cavapa
/necklaces, earrings/ (48286)
Vascular bundles/string in necklaces/ (48398)
C)avapa
Flowers/sarlands/ ( 40439 )
Oa,vapa
Fnrits/pearls in necklaces/ (48190)
Coix lachryma-jobi
Cayapa
C. lachryma-jobi
Coaiquer
Fruits/pearls in neckl aces/ (40540)
Frrrits/pearls in neckl aces/ (4 I 480)
Apocynaceae
Mandevilla doclsonii
Fabaceae
Genus A
Poaceae
Sapindaceae
Paullinia fascescens
Oavapa
Seeds/pearls in necklaces, one side red and one side
black,z (48426)
Table 10. Plaiting and textiles. Natural fibre s are typically fieecl from the plant tissues by pounding with a srick (Fig.B). An
exception is Cnrdukruicuprilmata fiorn which lone strips are torn fiorn the outer layers of the petiole. These are used for
making mats, fans and other lvoven items (Fig. B). To make bark cloth of Poulseniaermata, a large piece of bark is removed
from the tntnk. The fibres are derived from the inner bark as sholvn on Fig. 9. According to Ferdon (1956), the Coaiqueres wore bark clothins until very recentll'. The traditional clothing of both the Cayapas and Colorados were probably made of cotton. \,\'estern clothes tvpically made of svnthetic fabrics are usually the first sign of acculturation. Natural fibres are
replacecl by nylon in fishing nets and plaitecl items such as harnmocks. Nvlon is inexpensive, toueher than plant fibres ancl
more resistent to decar.'.
Tribe
Part of plant/purpose/
Oayapa
/strine for construction/ (48060)
Heteropsis oblongifolia
Coloraclo
Philodendron verrucosurn
C)ayapa
/baskets/ (40001)
/strins, not very strong/ (48l9l )
(AAU voucher)
Vascular cryptogams
Blechnr.rm volubile
Araceae
Arecaceae
Astrocary:m
standleyanlrnl
Desmoncus serifera
Oenocarpus mapora
Coaiquer
Leaves,/hammocks/ ( 60078)
Stem/baskets/ (41455)
Cayapas
l.eaf base/baskets/ ( 60 I 02)
Coaiquer
Stern/baskets / (487 87 )
Oavapa
Leaves/widely used previouslv for fishing nets/ (48386)
Oay'apa
Oayapa
Petiole/baskets, mats and fans/ (40419)
Aerial roots/baskets/ (4828I )
Cayapa
Bignoniaceae
Amphilophium
paniculattrnr
Bromeliaceae
Aechmea magdalena
Cyclanthaceae
Carludovica palmata
[.uclovia intesrifolia
Y
BS 46
68
Tabel l0 - Continued
Tribe
Part of plant,/purpose/ (AAU voucher)
Gyssopium barbadense
Cayapa
Cotton/previously
Marantaceae
Calathea sp.
Cayapa
Ischnosiphon leucoPhaeus
Cayapa
Petioles/ h ats/ (40 467)
Stem/hats / (407 80, 48902)
Poulsenia armata
Cayapa
Bark/bark cloth/ (48343, 48985)
Sapindaceae
Paullinia sp.
Coaiquer
Stem/string for tying logJstogethet/ (414358)
Cayapa
Cortex/string previously used for manv PtlrPoses e' g' hammocks/ (48433)
Malvaceae
Moraceae
IJrticaceae
Cecropia sp.
used for making clothing/ (40584)
grasses in the understorey of the rainforest is used to
Table 11. Dipilatories. one representative <lf the few, non-bambusoid
over the skin. Hairs are caught between the glumes and torn out.
remove facial hair. The narrow cylindrical spike is passed
is efficient, though painful'
treatment
Lfiom our own experiments that the
Tribe
(AAU voucher)
Colorado
(402\2,407\4)
(40315,4L079,60108)
Poaceae
Streoptochaet.a sodiroana
S. sodiroana
Cayapa
species is either used as a waterproof-er or as rubTable 12. Latex. The latex extracted from the stem of certain moraceous
ber (Fig. e ) .
Tribe
Moraceae
Brosimum utile
Castilla elastica
Cayapa
C. elastica
Coaiquer
Cayapa
Comments/ (AAU voucher)
Latex useclto waterproof canoes/(41037)
Rubber/ (48214)
Rubber/ (48994)
BS46
69
Table 13. Soap and shampoo. Washine hair with herbal preparations may not serve a hygienic purpose only. The infbrmants related several times that a particular hair wash also enhanced hair growth.
Vascular cryptogams
Elaphoglossum herminieri
E.rp.
Nephrolepis sp 1.
N. sp.2
Tribe
Part of plant/prepararion/comment/
Cayapa
Leaves/sround in cold water/ stimulate hair srowth/
(40533)
Cayapa
Coaiquer
Leaves/ /stimulate irair growth / (40826)
Leaves/boiled/stirnulate hair growth/ (48837)
Crude leaves//stimulare hair growth / (48845)
C)oaiquer
Polypodium sp. I
P. sp. 2
Cayapa
Trichomanes coralliatum
Genus A (Aspleniaceae)
G. B (Polypodiaceae)
Coloradcr
Cayapa
Cavapa
Oayapa
(AAU voucher)
/ /stimulate hair srowth/ (40795)
/ / / (48436)
/ /stilnulate hair growlb/ 40147
Leaves/fre sh / / (48301)
Leaves/fresh/ / 98303)
Agavaceae
Dracaena fragrans
Ciavapa
Leaves,/grouncl in cold water,/ previouslv used for
soap/ (11046)
Anthurium spp.
Cayapa
/ /stimll.late hair growrh/ (40417,48106, 48342)
Capparidaceae
Podandrogyne brachvcarpa
Colorado
/ /wash of body/ (40017)
Colorado
/ /stinulate
Phytolacca rivinoides
Colorado
P. rivinoides
Coaiquer
/ /wash/ (40003)
Fruits,/crushed in water/detergent for washing clothes/
(48830)
Coloradcr
/ /soap for washing the bocly,z(40012)
Coloradcr
/ /makes hair srow faster/(40076)
Araceae
Malvaceae
Sida acuta
hair growrh/ (4003t,)
Phytolaccaceae
Piperaceae
Piper marginatum
Rubiaceae
Hamelia macrantha
BS 46
70
Table 14. Perfumes. A distinction is not always made between aphrodisiacs and perfumes. The Cayapas believe that some
plants used for perfume also have magical effects that can be directed exclusively to the desired person.
(AAU voucher)
Tribe
Part of plant,/preparation,/comments/
Alstroemeriaceae
Bomaria edulis
Cayapa
B'rp.
Cayapa
Frdts/ /aromatic smell/ (40lll4)
Fruits/perfume extracted/strong smell that can be detected
from a long distance/ (48059)
Hyptis mutabilis
Colorado
Entire plant/body
Melastomataceae
Miconia sp.
Coaiquer
Flowers/perfume
Orchidaceae
Vanilla planifolia
Cayapa
Flowers/ / / (40340)
Peperornia sp. I
Clolorado
P.sp.2
Cayapa
I n f l c r r e s c e n c e // / ( 4 0 1 6 9 )
Leaves/woman's body rubbed/
men/ (40506)
P .s p . 3
Cayapa
Leaves,/cmshed and mixed with another rtnidentified ingredient/ aphrodisiac/ (48076)
Solanaceae
Solanum sp.
Cavapa
Flowers,/cmshed and body rubbed/ / (40522)
Lamiaceae
rubbed/ / (40039)
extracted/very pleasant smell/ (41 6l 0)
Piperaceae
irressistible to youns
Table 15. Dyes. This category comprises plants that are used for dying both clothing and skin. Colorados mainly use llixn
orellrtnaand. ()enipa americanafor their traditional bodypaint. They also use Bixa orellana for creating the clistinctive, red,
cap-like hairstyle (Fig. 8) .
Tribe
Part of plant,/preparation/ purpose/ (AAU voucher)
Colorado
Leaves/ground in water/dyes fabrics black/ (40198)
Leaves/ground in water/dves fabrics black/ (40207)
Leaves/ground in water/dyes fabrics black/ (40205)
Arecaceae
Chamaedorea pinnatifrons
Ceonoma cuneata
S y n e c a nt h u s w a r s c e r v i c z i a tnt s
Colorado
Coloradcr
Bixaceae
Bixa orellana
Colorado
B. orellana
Coaiquer
Oxalidaceae
Oxalis sp.
Coaiquer
Leaves/ground rvhile fiesh/dying
etc./ (41580)
Piperaceae
Piper sp.
Cayapa
Leaves/ground while fresh into a paste/ colottrs applied directly to paint various items/ (41026)
Colorado
Seeds and ptrlp of fruits/crushed
stripes on skin/ (49120)
Frtrits/,/clYes hair red/ (40045)
Fruits//dyes wood red/ (48897)
of fabrics, skin,
Rubiaceae
Genipa americana
and juice extracted/black
BS46
71
Table 16. Ornamental plants. A number of plants are cultivatecl as ornaments surrounding the houses in Cayapa villages.
Nearly all of the plants that the Colorados grow as ornaments are also usecl in their ritual baths and are not listed below.
Tribe
(AAU voucher)
Amaranthaceae
Amaranthus sp.
(4U3el)
Cuphea strisulosa
Cavapa
(40546)
Colorado
(40623)
Malvaceae
Hibiscus sp. I
Pavonia fnrticosa
Cayapa
Cavapa
(40429,40464)
(40555)
Cayapa
(40556)
Cayapa
(4058e)
Cayapa
(48364)
Cavapa
Cayapa
(40588)
(405e0)
Cayapa
(48378,48380)
Cayapa
(40463)
Apocynaceae
Allamancla cathartica
Clolorado
(40504,40779)
(414r.]e)
Ooaiquer
(40642)
Colorado
Polemoniaceae
Polemonium sp.
Balsaminaceae
Impatiens balsamina
Ochnaceae
Sarrvagesiaerecta
Asteraceae
Dahlia sp.
Mimosaceae
Calliandra an gustifolia
Araceae
Anthurium anoreanum
(AAU voucher)
Cayapa
Amaryllidaceae
genus indet.
Tribe
Lythraceae
(iavapa
(48362)
Cannaceae
Canna indica x generalis
Cayapa
(40530)
Commelinaceae
Callisia repens
Cayapa
(483e4)
Polygalaceae
Polygala mollaginifolia
P. panicr"rlata
Portucalaceae
Portulaca spp.
Rubiaceae
Cyclanthaceae
(48850)
Ixora sp.
Euphorbiaceae
Scrophulariaceae
Acalypha sp.
Colorado
(40648)
Alonsoa sp.
Cayapa
(40560)
Coloradcr
(40641)
Solanaceae
Solanurn jarnicense
Cayapa
(4046e)
Coaiquer
(41639)
Cyclanthus sp.
Ooaiqtrer
Hydrangeaceae
Hydrangea sp.
Zingiberaceae
Lamiaceae
Coleus x hybridus
Cavapa
(48357)
Hedychitrrn coronarium
Table 17. Hedges. Onl;- the Coloraclos plant hedges around their clwellings and between fielcls. The most imporrant spec i e sa r e l i s t e d b e l o r , r ' .
Tribe
(AAU voucher)
Euphorbia cotinifolia
Coloradcr
(40177)
Fabaceae
Erythrina eclulis
E. srnithiana
Coklraclo
(40646)
(40632)
Euphorbiaceae
Colorado
Y
72
BS 46
Table 18. Hallucinogens. This category comprises very powerful and dangerous plants. For some of them it is important to
prepare mentally for the strong psycho-active effects that they provoque. This is particularly true for Banesteriopsiscaapii .
The Cayapas use this species as the main ingredient in the hallucinogenic decoction named pindi. Other complimentarv
plants are added to modif the effects or to remove the bitter taste. The Cayapas believe that only the shaman has the power to control this plant. It takes several years for a shaman apprentice to prepare for the visions that the plant drug induce.
The Colorados call the hallucinogenic decoction of Banesteriopsiscaapii for nepd Usually, they prepare it with no other
plants. Traditionally, nepi is used by all male members of the tribe. It is notable that the Colorados use Banesteriopsiscaapii
as a laxative too (Table 43) . Brugmansia aersicoloris an extremely powerful and dangerous plant drug that according to our
knowledge is no longer in use among the Cayapas.
Tribe
Part of plant/preparation/treatment/
(AAU voucher)
Cayapa
Leaves/ boiled with stems of Bnn esteriop sis caapiil decoction
drunk/makes the decoction taste less bitter/(40535)
Banisteriopsis caapii
Cayapa
Bark/grated, boiled/5 teaspoonfuls drunk in evening with
sugar cane brandy/induces visions, shaman only/ (40299,
B. caapii
Colorado
Stem/grated and boiled,/some pllassesdrunk warm,/induces
visions/ (40722)
Piperaceae
Piper variegatum
Cayapa
Leaves/ground, mixed with hot water/ drunk/induces
lucinations / (40325)
Cayapa
Leaves/boiled
Cayapa
Leaves/ground in cold water/a teaspoon of the extract
drunk/to induce strong hallucinations/ (40581, 41050)
Cayapa
Leaves,/boiled with stems of Bnnesteriopsiscaapi/ drunk/
enables the shaman to see the spirits clearly/ (483\7)
purpose/
Acanthaceae
Justicia sterea
Malpighiaceae
40582)
hal-
Rubiaceae
Psychotria viridis
with stems of Banesteriopsiscaapi/ drunk/to
enhance the hallucinogenic effect and make the decoction
taste less bitter/ (40595, 48368)
Solanaceae
Brugmansia versicolor
Thymelaeaceae
Schoenobiblus panamensis
BS46
-lo
t3
Table 19. Antifertility
agents. This category includes contraceptives, abortifacients and agents that cause permanenr sterilization in women. Due to strong taboos we only learned a few things about the plants that are used for these purposes.
More work is needed. Information was obtained on two additional antifertility plants that have not been included in the list
below. One is allegedly very powerful and highly acclaimed. We never saw it during our fieldwork despite our efforts. According to the description it is a rare fungus or achlorophyllous plant that is found where trees have recently been uprooted. The other plant is a particular coconut variety that bears small aborted fruits. The liquid end.osperm from these is
said to have antifertility effects which is probably an example of the Doctrine of Signature.
Tribe
Part of plant/ prep aration / tr eatment/
purpose/ (AAU voucher)
Colorado
Pods,/boiled,zdrunk,zinduces abortion/ ( 40895 )
Colorado
Seeds,/boiled/decoction of 1 seed drunk once a day/prevents pregnancy. Will eventually cause sterilisation/ (49l lU)
Colorado
Rhizome/ground,
Fabaceae
Desmodium uncinatum
Lauraceae
Persea americana
Zingiberaceae
Renealmia sp.
boiled/a glass of decoction drunk daily
for ayear/causes sterilisation / (49121)
Table 20. Ritual ornaments. During curing rituals performed by the shaman, the patient's house is often decorated. with ornaments that are believed to oust evil spirits and thereby increase the chances of being healed. Plant ornaments are also
hung above the bed of mentally disturbed persons.
Tribe
Comments/ (AAU voucher)
Cayapa
Decoration in connection with curing ceremonie s/ G0277)
Coaiquer
Hung above patient during curing
Cavendishia engleriana
Coaiquer
C. grandifolia
Coaiquer
c.rp.
Coaiquer
Psammisia ferruginea
Coaiquer
Hung above bed/(a8925)
Hung above bed/(48965)
Hung above patient during curing ceremonies/(a8806)
Hung above patient during curing ceremonies/ (487g2)
Gesneriaceae
Besleria solanoides
Coaiquer
Araceae
Anthurium
sp.
Bromeliaceae
Guzmania testudinis
ceremonies/ (48856)
Ericaceae
Hung above patients suffering from mental disorders
(48735)
l
I
l
BS 46
74
Table 21. Magic plants. The Colorados use a number of plants to improve their luck. The Cayapas o{ten carry a little bag
with magic plants that are supposed to bring luck during hunting.
(AAU voucher)
Tribe
Part of plant/comments/
Odontonema strictum
Colorado
Flowers// (40023)
Aristolochiaceae
Aristolochia pichingensis
Colorado
Leaves/magicritual saiclto clean the hortsefrrr bad hrck/
(40718)
Asteraceae
genus indet.
Colorado
Flowers// (40024)
Malvaceae
Hibiscus sp.
Colorado
with other species/ (40027)
Flowers,/mixecl
Nyctaginaceae
Mirabilis jalapa
Colorado
Flowers// (40025)
Peperomiaceae
Peperomia sp.
Colorado
Leaves// (40638)
Rubiaceae
Amphidasya sp.
Cayapa
Stem stripped for leavesand kept in a specialsackduring
hunting, said to help finding animal and make the hunt successful/(40339,4l 059)
Apocynaceae
BS46
l3
Table22. Pedagogy. This category comprises plants that are used to improve misconduct by children.
Many are usecl according to the Doctrine of Signature and thus have a psychological effect. The calming effects
of some species can not be
ruled out, however.
Tribe
Comments/ (AAU voucher)
Cayapa
On the first Monday after full moon the mouth is washeci
with a cold exrract of the leaves,/(48144)
Phyllanthus stipularis
Cayapa
Applied to eyes of childen who cry too much and cannor
sleep/ (40392)
P'rp.
Cayapa
Twigs placed on forehead, said to promote sleep,/(48231)
Colorado
Stringed and worn around neck. For chilclren that cry too
much/ (40060)
Cayapa
Feet rubbed with fnrits to encc)urage walkins of children,/
(48032)
Acacia riparia
Cayapa
The eyes are covered by a piece of fiesh leaf, said to encourage sleep/(40483)
Mimosa pudica
Cayapa
Green leaves placed on forehead, said to promore
sleep,/ (48324)
Cavapa
In the morning the mouth is washed with a cold extract of
ground leaves to prevent dirty language and desobidience/
(48226)
Coaiquer
Creen leaves soaked in water, skin rubbed to stop weeping
of children / (41508)
Cayapa
Mouth rubbed with leaves at dawn after full moon to prevent bad language and desobidience/ (40297)
Acanthaceae
Justicia pectoralis
Euphorbiaceae
Flacourtiaceae
Carpotroche platyptera
Melastomataceae
Triolena sp.
Mimosaceae
Rubiaceae
Borreria ocymoides
Urticaceae
Urera caracasana
Verbenaceae
Aegiphila sp.
BS 46
/6
fields (Fig. 1 1) ' Plantains
Table 23. Staple diet. Listed below are only rhe sraple crops of slash-and-burn or slash-and-mulch
much cacao and
cultivate
colorados
The
cayapas.
the
and
coaiqueres
the
both
of
crops
and corn are the most important
coffee. The Alrican oil Palm is widely cultivated in the region.
Tribe
Common name in English/(AAU
Colocasia esculenta
Cayapa
Taro/ (48145)
Convolvulaceae
Ipomoea batatas
All groups
Sweet potato/ (48305, 40422)
Dioscoreaceae
Dioscorea esculenta
Cayapa
Yams/(48331)
Euphorbiaceae
Manihot esculentum
AII groups
(lassava/ (48330, 48329)
Phaseolus vtrlgaris
Cayapa
Bean/ (40516)
Musaceae
Musa x paradisica
All groups
Plantains/ (not votrchered)
Saccharum officinarum
All groups
Zeamays
All groups
Sugarcane/ (not vouchered)
Corn/ (40525)
voucher)
Araceae
Fabaceae
Poaceae
species are cultivated in orchards near
Table 24. Edible fruits and seeds. The fmits of numerous plants are eaten. Some
a wide variety of fruits.
houses or in fallows. When travelling in the forest, the Indians usually collect and eat
voucher)
Tribe
Comments / local name/(AAU
Actinidiaceae
Saurauia brachybotrYs
Coaiquer
Sweet/
Anacardiaceae
Spondias mombin
Colorado
S. purpurea
Cayapa
Cultivated/"Ciruela" ISpanish] / (491 00)
"Hocos
boca"
Highly esteemed, cultivated/"Ovo" ISpanish],
(481'40,
40428)
[Cayapa] /
Annonaceae
Annona muricata
Cayapa
Rollinia mucosa
Cayapa
R. mucosa
Coloradcr
R. mucosa
Coaiquer
Apocynaceae
Bonafousia longitubulosa
Coaiquer
"Moquillo"
ISpanish] / (41621)
"
"Oa-n
a-sa-tap6 I lCay apa) /
Hi ghly esteeme d, cultivated/
(48103)
"chichibilia-fin-chumo-boca"
ICayaMuch eaten, cultivated/
pal / (a0608)
"Chirirnoya"
ISpaCultivatedz"'Pastanu" IColorado],
nishl / (40046)
"(lhirimoya"
ISpanish] / (48989)
ICoaiquer],
/"Churimono"
B.rpp'
Ciayapa
Tabernaemontana tetrastachYa
T.tpp.
Cayapa
Cayapa
Srveet ancl rich/"Oyap saya" [Coaiquer / (41632)
Sweet and acid/"Do-pistcha" fCayapa] / (48057,48206)
"Do-pistcha"
ICavap a] / (407 50,40775)
/
"Do-pistcha
bamo boca" [Cayapa]/
Sweet//"Do-pistcha" or
gentrs indet.
Cayapa
(40351 , 40426, 40517 , 40732, 40739)
Sweet, keeps up to 3 weeks/"Do-pistcha"
[Cayapa] / (41044)
BS46
Tabel 24 - Continued
Tribe
C o r n m e n t s / local name/ (,\{U voucher)
Cavapa
E d i b l e m e s o c a r p , /( 0 0 0 7 8 )
Boilecl or eaten fiesh,/(00113)
Boilecl/.Pi-cani-chi" ICav:rpa] / (60 I l0)
/ / (600t 0\
Arecaceae
Astrocary.um standleyanun-r
Bactris easipaes
Cavapa
B. setosa
B. setulosa
Desmoncus serifera
Euterpe chaunostachvs
Cayapas
Geonoma cuneata
Phytelephas aeqtrarorialis
Cavapa
Wettinia qtrinaria
Cayapa
Socratea exclrrhiza
Svnecanth us warscewiczianus
S. warscewicziantrs
Ooaiquer
Oayapa
Coaiquer
Coaiquer
Oavapa
Ooaiquer
Cayapa
C n r d e / " B o r a n e g r a <[ S p a n i s h ] / ( 4 1 4 5 5 )
Cr-ude/"Marnba-san-chi" ICayapa] / (601 03
"Yo
, / " Y a h - a l - c h i "o r
ya chi" fCayapa] (407b2,48351, 60099)
Liquid endosperm and inner mesocarp,/.|)in_chi"
[(]aya_
p a l l ( 6 0 11 1 )
"palmira"
Cnrcle endohausrorium/"Ban-chi,' ICa,vapa],
[ S p a n i s h ] / ( 4 1 0 74 , 6 0 1 t 2 )
B o i l e d / " G u a l t e c r e s p o " I s p a n i s l i ]/ ( 6 0 0 0 7 )
/"Bc>ranegra" ISpanish]/ glabb)
BoiIed/ / (60076)
Aristolochiaceae
Aristolochia pilosa
Colorad<r
/ " l v u q u n t o " [ C o l o r a c l o ] , " ( ] r e n a d i l l a d e r o s : r ' ,[ S p a n i s h ] /
(40235)
Quararibea coloraclonrm
Q. cordata
Colorackr
Q. soeuenii
Q. soeeenii
Cayapa
"Declo"
/
[Coloraclo] / (1001b, 4013 1)
(lultivated,z "Sapote tape " Cay apa) (
I
/ 40580 )
/"C)6 sapote" [Cayapa]/ (40772)
Coaiqucr
/"Zapote"
Campanulaceae
B r r r m e i s t e r ar r r l q a r i s
Oavap:r
/"Sjchi-vesj-tchap6" ICayapa] / (4t01t)
Carica papaya
Coloraclo
Carica papaya
(la1'ap:r
C t r l t i v a t e d , / " P a p a v o "I C o l o r a c l o ] , , ' p a p a y : r , '
Ispanish] /
(4015e)
"Papal
Cultii.'atedl
la finchuno boca,,/ ( 40485 )
Bombacaceae
Oayapa
[Spanisl'r]/ (41860)
Caricaceae
Clusiaceae
Chrvsochlamys depenclens
Clusiir sp. I
C. sp.2
Cayapa
Rheedia edulis
R. edulis
Cavapa
Oayapa
Cavapa
Coaiquer
Tovomitopsis
Cavapa
Rheedia sp.
genus indet.
Cayapa
Cayapa
A chilclrens favorite/"Nan-boca,, ICa1,apa] (40409)
/
"4ba -boca" ICay'apa]/ (40908)
/
"A
-ba n-cho-boc a,,
Wh ite flesh of rnzlrrrre fruits/
fC,ayapa]/
(48234)
Sweet ftrste/"Madrr)iio" ISpanish] / (41b7 5, 41342)
Sweet and rasreful/ "Sji-pi stc ha-ka-bo cac a,,
fC,ayapa] /
(48363)
Acid, with susar or salt,/"N:rn-bo!-chi"
ICavapa] / (4BlgZ)
"Madroiio"
/
Ispanish] / (40324)
"Niang-boca"
/
IOa1'apa]/ (60 I 00)
Cucurbitaceae
Luffa aegyptica
Cayapa
R v t i d o s t v l i sc a r rh a g i r r e r r s i s
Cayapa
Boilecl. (lultivated but uncornm.n/
dio-tape" fCavapa] / (40b97)
/ " C h i n - l a q p e " I C a y a p a ]/ ( a 8 0 5 I )
"Tebab.
quillachi-r-eme.
BS 46
78
Tabel24
- Continued
Tribe
Comments / local name/ (AAU voucher)
Cycadaceae
Zamra lindenii
Cayapa
Seeds grounded to flour for special bread/"Sa-oa-pa-chi"
fCayapa] / (48155)
Cyclanthaceae
Cyclanthus bipartitus
Colorado
/"Pinta"
Dioscoreaceae
Dioscorea sp.
Coaiquer
/"Chirma"
Coaiquer
/"Guish"
Carpotroche platYPtera
Casearia sp.
Colorado
/'\ehicz"
Genus A
Genus B
Cayapa
fColorado] / (40221)
ICoaiquer] / (48826)
Ericaceae
Psammisia caloneura
Flacourtiaceae
Cayapa
Coaiquer
[Coaiquer],
"Ava
de monte" [Spanish] / (48920)
IColorado] / (40060)
/"Pique lanboca" [Cayapa] / (40445)
"Ya-sa-te-pistcha"
ICayapa] / (48 I 69)
/
silvestre"
ISpanish] / (48702)
/"Morcillo
Gesneriaceae
"Abanchola finchuno
Codonanthe crassifblia
Cayapa
Columnea tenella
Coaiquer
White berries collected from ground/
bugu" fCayapa] / (40479)
/ / (41577)
Lacistemataceae
Lacistema aggregata
Coaiquer
Taste like mango/"Pailde"
Lauraceae
Persea americana
Colorado
Cultivated/'Alin"
(4e118)
Malpighiaceae
Bunchosia cornifolia
Cayapa
/ / (4803r)
Melastomataceae
Ossaea micrantha
Colorado
/"Pichi ri chide" [Spanish]/ (40162)
Meliaceae
Carapa guianensis
Coaiquer
/"Aray" [Spanish]/ (41667)
Menispermaceae
Cissampelos tropaeolifolia
Cayapa
/"Pin tsu ruro chua" fCayapa]/(40851)
Mimosaceae
Inga edulis
Colorad<r
I. edulis
Cayapa
L marginata
Cayapa
I. ruizana
Cayapa
I. spectabilis
Cayapa
Myrsinaceae
Cayapa
Ardisia romeroi
A. ronseroi
Cayapa
A.rp.
Cayapa
ICoaiquer] / (48796)
IColorado],
"Aguacate"
ISpanish],/
"Guaba"fSpanish] (40042)
/
Cultivated/"Pitsa"fColorado],
"sichijaquCultivatedmany cultivars/"Pu shill6" [Cayapa],
"pichiche"
ie" f Cayapal,
[Cayapa)/ (40353,40450,40778,
48200)
/"Pichillo" ICayapa]/ (40733)
/"Minga shi chi tap6" [Cayapa)/(40898)
(40772)
"
40906)
,/"Ibpo-chin-boca-(tap6) ICayapa)/ G05a2,
"Ka-bo-chin-boca-tap6"
lCay apa)/
Tasteful and sweet/
(48325)
/"Yasa shi pij cha" [Cayapa)/ (a0817)
BS46
79
Tabel24 - Continued
Tribe
(lomrnents / local name/ (AAU
voucher)
Psidium sp.
Cayapa
/"Yunqui rap6" [Cayapa) / (40404)
Passifloraceae
Passiflora auriculata
(lolorado
/"Wero quinro" fColoraclo], "Grenadilla cle loro,,
[Spa_
nishl/ (402ltl)
Cultivated,/"Maracuya" ISpanish] / (40026)
"Shin
lape" [Cayapa]/ (40387)
Ctr ltivated / "Cho-roro-fi n-choni-boca', Caya pa]
I
/ ftO b}g)
Cultivated/"Na wa quinto" IColoraclo], "grenaclilla.
ISpa_
nishl / (10174)
Myrtaceae
P. edulis
Colorado
P. foetida
Cayapa
P. malifbrmis
P. quadrangularis
Coloraclo
P. quadrangularis
Ca,vapa
P.tp.
Coaiquer
Cayapa
Sweet and tasteful,/"Bancle-cho-roro-chua"
fCayapa],,,Bad_
ea" [Spanish]/ (4035b, 4U187, 48429)
"Grenadilla"
/
ISpanish]/ (aBBb7)
Rhamnaceae
Gouania lupuloides
Oayapa
Sweet tasre/"Shui pu chua,' [Cayapa] (40744)
/
Rubiaceae
Pentagonia grandiflora
Coaiquer
P. macrophylla
Cayapa
Sweet taste/"Cham" [Coaiqu erl / .!lbgg)
/"Pe-dju-pisrcha-tap6" lCayapal / (404b1, 48361
)
(40375)
P. sp. 1
Cayapa
P.sp.2
Cayapa
Sabiceavillosa
Arachnothrlx inconstans
Coaique r
Ooaiquer
Crude/P6-dju-pistcha-tape" ICayapa] (4t001)
/
Taste like apple/ (41623)
"Mocillo"
/
lspanishl / (48768)
Rutaceae
Citrus sp.
Coloradcr
Cultivated/"Lansa"
IColoraclo],
"Naranja"
(4072r)
ISpanish] /
Sapindaceae
Allophylus sp.
Talisia sp.
Coloradcr
Cayapa
,/"Unilo piyo" lColoradol / (40lgl)
/' A-chi-boca" [Cavapal / G0BZ0)
Sapotaceae
Chrysophyllum argenteum
Pouteria caimito
P. collina
P. torta
Colorado
Cayapa
(loaiquer
Coaiquer
Cultivated/"Caimiros"IColorado]/ (40289)
/"A-cuita" [Cayapa]/ (4041b)
"Caimitillon"
/
ISpanishl/ GI67 Z)
/"Piast" [Spanish]/ (41629)
Solanaceae
Cestrum baenitzii
Physalis angulara
Coaiquer
Cayapa
P. pubescens
Solanum candidum
S. canense
S. mammosum
Coaiquer
S. quitoensis
S. sessiliflorum
Cayapa
s.sp.2
Cayapa
Coloraclo
Coloraclo
C)oaiquer
Coaiquer
"Chalmolan"
/
Ispanish]/ (al 650)
Sweet,/"Ba-pist-chi"
or "papicha-finoboca_chi"
ICayapa]/
(40609,48385)
,/"Casena"[Cayapa]/ (40311)
/"Uwica" [Colorado]/ (4016b)
(40016)
"Estacr,rdo"
/
ISpanishl/ eUbZ)
Crrltivated/"Naranjilla"ISpanish)/ (4lbg6)
"Larajilla
/
tape" [Cayapa]/ (40509)
/"Yalte" [Spanish]/ (4gBZg)
BS 46
80
Tabel24
- Continued
Tribe
Comments / local name/ (AAU voucher)
Sterculariaceae
Herrania baluensis
Colorado
/"Apilon"
(40666)
H. baluensis
Thcobroma bicolor
Cayapa
Oayapa
T. gileri
Cayapa
/"'6, cacabc' chi" [Cayapa]/ (a0a16)
/"Chis po yo" [CaYaPa]/ (40418)
Taste like cacao/"L'lyoko-pistehi" iCayapal / (48430)
[Colorado] ,
"Cacao de monte"
[Spanish] /
IJrticaceae
"U1'2
Pourouma guianensis
Colorado
Sweet like candy/"Lati"
nishl/ (10677)
P. hirsutipetiolata
Cayapa
Sweet/"Yapistchi" ICayapa],
Verbenaceae
Aegiphila sp.
Coaiquer
/ " M a n g a m o r a ' I S p a n i s h| / ( 4 8 7 7 1 J \
[Coloraclo],
"Palo
de monte" [Spa-
diura" ISpanish] / (41009)
collected from the wilcl in their traditional dishes' TheTable Zb. vegetables. Especially the Cayapa Indians use many leaves
se are typically prepared with porc rind before serving'
Tribe
Vascular cryPtogams
Dennstaedtia sp. 1
Cayapa
D. sp.2-3
D. sp.4
Coaiquer
Diplazium fraseri
Dryopteris sp.
Hypolepis hostilis
Cayapa
Nephelea cusPidata
genus indet, (PolYPodiac.)
senus indet. (Dennstaedtiac.)
Cayapa
senus indet. (HvmenoPhYllac. )
Coaiquer
Araceae
Anthrtrittm lancea
Cayapa
Coloradcr
Coaiquer
Cayapa
Part of plant/preparation/
(A\U voucher)
fatl (48299)
Juvenile fronds/boiled with
Fronds/boilecl with fat and salt/ (48847' 48884)
(49036)
Juvenile fronds/boiled/
with fatl (48049)
fronds/ground
Juvenile
Fronds/boiled with fat and salt/ (4BBB7)
Froncls/rnidrib boiled with fat or black protttberances on
stem sratecl and usecl like noodles in soups,/ (40527,41070,
48176)
Cayapa
Cayapa
Rhizornes/grated and boiled / (40442)
(40431 )
Juvenile fronds/boilecl/
with fatl (4fl0'1il)
fronds/grouncl
Juvenile
juvenile fronds/boilecl with fat and salt/ (48848)
A. versicolor
A. sp. 1
Cayapa
Cayapa
Froncls/cut into pieces and boilecl with fatl (4104U)
fatl (4809U)
.|uvenile fionds/boiled with
Juvenile fronds/boiled/ (40357)
A .s p . 2
Cayapa
Juvenile fronds/dried/
Arecaceae
Aiphanes selatinosa
B a c t r i sg a s i p a e s
Coaiquer
Palrnheart//(60003)
B. setosa
B. setulosa
Euterpe chaunostachYs
Cayapa
Oayapa
Coaiquer
Palmheart//(60113)
Palm heart/ / (60110)
Palm heart/ / (60010)
Coaiquer
Palmheart//(6000\)
E. chaunostachYs
Geonoma sp.
Ciayapa
Palmheart//(60103)
Coaiquer
Palmheart//(48916)
Iriartea deltoiclea
.fesseniabataua
Cayapa
Palmheart//(60097)
Coaiqtrer
Oayapa
Palmheart//(60006)
Prestoea se-iuncta
Socratea exorrhiza
Cayapa
Palmheart//(60107)
Palm heartt / $01tl7 )
(40850)
BS46
8l
Tabel25 - Continued
Part of plant/preparatiorr/
(AAU voucher)
Asteraceae
genus inclet
Clolorado
Fronds// (40103)
Oayapa
Petioles/peeled, pith with salt or sugar/raste very acidic/
( 4 8 4 2 1)
Burmeistera vulgaris
Cavapa
B'tpP.
Oa-vapa
f.eayes,/prepared with fat/ (41 03 1 )
Leaves/boiled or crushed and fiiecl/ (40343, 4097B, 4t3070;
(loloradcr
Leaves,/boiled/ (400tt8,40158, 40195)
C)oaiquer
Pith of stem/fresh/(48860, 48907, 4U917)
Rytidostylis carthaginensis
Ciayapa
genus indet.
Coaiquer
Leaves/with fat / (48365 )
Distal l0 crn of shoots/bolled/ (41579)
Begoniaceae
Begonia sp.
Campanulaceae
Caricaceae
Carica spp.
Costaceae
Costusspp.
Cucurbitaceae
Leclthidaceae
Gustaviadodsonii
C.a sp.
Cayapa
Oayapa
Leaves/boiled/ (403a0)
Cavapa
Young apical leaves/fresh or boiled/ (48181)
Cayapa
Yotrng shoots/with salt/fiorn fielcls/ (40461, 48053)
Cavapa
Leave s/ bo ile d / (40 45 1)
Leaves/boiled with fat/ (48141)
Marantaceae
Calathea lutea
Melastomatceae
Anthrostema ciliatum
Phytolaccaceae
Phytolacca rivinoicles
Table 26. Edible flowers. Only the Oayapas inclucle flowers, flowerbuds and inflorescences in their diet.
Tribe
Part of plant/preparation/commenrs/
S c h l e g e l i ac h o c o e n s i s
Cavapa
S. fastigiata
Cayapa
/ / / (18404)
White part of flowers/cnrcIe/ / (41009)
(AAU voucher)
Bignoniaceae
Ericaceae
Cavendishia spp.
Psammisia attaberrans
Cayapa
P.tp.
Cayapa
F\owers/ / as vitamins/ (40309, 40520)
Flowers/with salt/verv tasty/ (48250)
Flor'vers/crude/srveer taste/ (48061 )
Gesneriaceae
Drvmonia coriacea
(iayapa
Flowers//srveet taste/ (40550)
Cavapa
Inflorescences/boiled / / (40433)
Cavap:r
Flower buds/ suckecl/like cancly/ (40297 )
Amphiclasya sp.
Cavapa
Pentaeonia sp.
Oa,vapa
Flowers/with salt/r,erv tasr,v/(,1824U)
Oalp<, receptacle,//srveet and rich/ (4 I 082)
C.ryapa
Marantaceae
Calathea sp.
Melastomataceae
Blakea subconnata
Rubiaceae
BS 46
82
(Fig' 12) '
ear rhe larva of a weevil' Ilhynthopttrus pnlmarwz
palms. The cal,apas ancl the coaiqueres
Table 2T.Larvaefrom
intense'
an
by
producing
Pungent
presence
of several palm species. It reveals-its.
The larva is collectecl from clecayir-rg,t.*,
on the spot (the
raw
eaten
either
is
It
delicacy'
a
from some distance. The larva is considered
Ltc
uc detected
call be
can
that
[nat
odor
oclor
...qn-lik e afiertast.e.
aftertaste'
except for a soap-like
rather'eutral
or brought home for fiying. The taste is
removed)
first
are
mandibles
(AAU voucher)
Tribe
Arecaceae
Bactris g;asiPaes
Cayapa
B. setulosa
Iriartea deltoidea
Coaiquer
L deltoidea
Socratea exorrhiza
Cayapa
S. exorrhiza
Cayapa
Wettinia quinaria
W. quinaria
Coaiquer
Coaiquer
Coaiquer
Cayapa
( 6 0 1l 3 )
(60010)
(not vouchered)
(600e7)
(60007)
(not voucherecl)
(60005)
(60112)
ism,:rd;.bet*::l:l:,0|.1:':-tl1::i::::t::::'.Tli]llLt::Jf;:jig
distinction
arbitrarv
A somewhat
rable28.Beverases.
andareprepared
or decocrions
hereareusedrbr inrusions
;r;;i;;;;t;il;d
a*r:il
Hl.i::',:Tnt:J;"?.iil
The
rhe
-^trr-ae ^f
inrnrnrfor
tnanv
Colorados.
cororados'
tnanv
ror
income
or
solrrces
areimportant
il;il;;J.u.uo
il:,1il#J:T:j:,:ffiHJfi,;#:11"..i
two cash crops'
Cayapas rarelv sell their surplus of these
(AAU voucher)
Tribe
Part of plant/preparation/comments,/
Mendoncia sP.
Cayapa
(40813)
Stem/clried, mixed with sugar cane brandy / /
Capparidaceae
Capparis ecuadoriana
Coloradcl
Woocly stem/ground,
Euphorbiaceae
Acalypha sP'
Colorado
/ /warmdrink
Lamiaceae
(loleus x hybridus
Coaiquer
Acanthaceae
boiled/ stimulant/ (40097)
to fight cold chills/ (40725)
Colorado
Leaves/ ground/for fatigue/ (41 47 6)
only / (40544)
Leaves/boiled, mixed with sugar/for taste
l.eaves/boiled/hot drink/ (406 I 6)
Colorado
Wood/boiled/said
Colorado
Cavapa
Leaves/boilecl/warms the body/ (40 102)
Leaves/boiled/like teal (40855 )
Passifloraceae
Passif'lora fbeticla
Colorado
Leaves/boilecl / / (40660)
Rubiaceae
Coffea arabica
Cayapa
Colorado
crop/ (40470)
/ /for domestic trse ancl as cash
(40645)
croP/
cash
/ /
Colorado
Fruits//fbr
Hyptis caPitata
Ocirnum camPechianum
Leclthidaceae
Grias peruviana
Cayapa
Orchidaceae
Epidendron difforme
Scaphyglottis Prolifera
C. arabica
Rutaceae
Citrtrs sp. I
C. sp.2
Cayapa
to give strensth/ (40220' 40121)
a juice/ (40720)
(
Leaves/boilecl with sugar'/ stimulant/ 48 194)
BS46
83
Tabel28 - Continued
Tribe
Part of plant/preparation/comments/
Cestrum sp.
Colorado
Solanum candiclum
S. sessiliflorum
Colorado
Stem/boile d/ for warming the body/ (40092)
Frttits/ / juice/ (40656)
Fruits/ /j uice/ ( 40509)
(AAU voucher)
Solanaceae
Cayapa
Sterculiaceae
Theobroma cacao
T. cacao
Cayapa
Colorado
/ /for domestic use and as cash crop/ (40460)
/ /cash crop/ (40618)
Table 29. Food flavouring. One of the favorite flavours is that of Eryngium.foetirtum which is widely used in soups.
All
groups cultivate a number of chili pepper varieties.
Tribe
Part of plant/ use/commenrs/
Genus indet.
Cayapa
Leaves/added to corn flour for a betrer taste/ (40482)
Apicaceae
Eryngium foetidum
Cayapa
Leaves/ spice in soup/(48926,483b9, 40427)
Cucurbitaceae
Rytidostylis carthaginensis
Cayapa
L.eaves/ / cultivated/ (40607, 40462)
Coaiquer
Leaves/for soups/cultivated/ ( 48982 )
Cayapa
L e a v e s // / ( 4 B l 0 l )
Cayapa
Leaves/for soups/cultivated / ( 40559 )
Capcicum frutescens
Colorado
C. fiuctescens
Cayapa
C. fructescens
Coaiquer
Fruits/spice/cultivate d / (40201)
Fruits/spice/several culrivars collected/ (404b6, 40565,
40566, 40586)
Fruits/spice/cultivared/ ( B99BB)
(AAU voucher)
Araceae
Lamiaceae
Ocimum sp.
Phytolaccaceae
Phytolacca rivinoides
Portulacaceae
Talinum paniculaturn
Solanaceae
Table 30. Food colouring. Onlv two species of lJixa are usecl fbr this purpose. The red dye is exrracred from inside
the fleshy capsule. Bixa orellana has been approved as a food additive in the industrialized world where it is used to colour
butter.
among other things.
Tribe
Part/ use/ cultivated/ (AAU voucher)
Bixa orellana
B, orellana
Colorado
B. platycarpa
Cavapa
Fruits/for food/cultiv ated/ (4004-r, 40704)
Fruits/for soups/manycultivars/(40472,40474, 40602)
Seeds/todye food yellow/(40810)
Bixaceae
Cayapa
BS 46
B4
Table 31. Fodder and fish bait. Fruits fiorn clifferent plants are trsed as fish bait by the Cayapas and the Coaiqueres. They
often constitute the natlral fbocl fbr fish in the periodically floodecl f orest. Pseuderanthenumrs also f'ed to chickens fbr me"sabalo"
is probably :r representative of the Benus Br)cnn.
alicinal purposes. The fish locally named
Comruents/ (AAU r otrcher)
Acanthaceae
Coaiquer
Leaves ttsed to feed chicken/(4U935r)
Anthrtrium sp.
()oaiquer
Philoclendron sp.
Coaiquer
Fnrits usecl as fish baitl (4U782)
Fruits uscd:rs fish bait,/(41652)
Charnaedorea pinnatifions
Coaiquer
Fruits used to f'eed chicken/(a890tl)
Asteraceae
Wulffia baccata
Ciavapa
For f'eeding domisticated irnimals / (48167)
Clusiaceae
Tovomitopsis sp.
C)oaiqr.rer
Fnrits used as fish bait/ (41616)
Coaiquer
Coaiquer
Boiled fruits used as fish baitl (41631)
Fruits used as flsh bait,z(48772)
Flacourtiaceae
Carpotroche platvptera
Cayapa
Seeds uses for fishins
Moraceae
Ficus insipicla
Cayapa
Fnrits used as fish bait,/(41080, 4fi0r56)
Rubiaceae
Raritebe palicotrrc'oicles
Cavapa
Fruits used as fish baitl (48066)
Sapindaceae
Oupania citrerea
Cayapa
S e e d su s e d a s f i s h b a i t l ( 4 1 0 4 6 )
Solanaceae
S o l a n tt r t t t o t t t c r l i s t ' t ' i atlt m
Coaiquer
Frr.ritsused as frsh baitl (41601)
Pserrderanthentrtn tnicranth ttm
Araceae
Arecaceae
Euphorbiaceae
Alchornea sp.
Hierolrinta clt<lcoensis
"sibalo"(40U73, 40366)
Table 32. Wildlife attractants. The Coaiqueres ancl the Cavapas possess cletailed knorvledge on the behaviortr and the ecoon a silogl o1thet anin-ralsthev hunt. C)trr inforrnarlts often told trs abotrt the animals that rvere known to feed specifically
rnind'
t.he
infrlrnrilnt's
to
the
calne
other
use
no
When
systematicallv.
collected
not
was
u.ii plor-rtspercies.This information
mentioned'
often
was
attractant
as
a
game
irnportance
plant's
Acanthaceae
Ruellia ttrbillora
Apoclmaceae
Manclcvilla dodsonii
Tabernaemontana heteroPhvlla
(AAU vortcher)
Tribe
Anirnals/comments/
Oavapa
Birds/visit flowers/ (48402 )
Cayapa
Bircls/r'isit fl owers/ (48307)
Birds/eat Irvits/ / (48123)
Oal,apa
Araceae
Anth ttrirtm cattlttrrhizttln
Cayapa
A. gracile
A. spp.
Cayapa
(la,vapa
Birds/eat fi-uits/ (48243)
Birds/ezrt berries/ ( 4f1232)
Birds/e:rt berries/ (48095, 48239)
BS 46
ti5
Tabel32 - Continued
(dA.U voucher)
Tribe
Animals/comments/
Ph ilocle r-rdro n verrLlcosum
Cayapa
Xanthosoma daguense
X. sagittifolia
Cayapa
Birds,/eat fiuits/ (48 l9 I )
Birds, turtles/eat rhizomes/ (48l89)
genus indet.
Cayapa
Turtles,/feed on yolng leayes and caught at night in slashand-burn fields using flashlights/ (48332)
Bircls/ eat frtrits/ (48246)
Coaicltrer
B i r c l s / e a t f r u i t s / ( 4 1 6 0 6 , 4 B 7 l B , 4 8 U 0 3 ,4 8 9 1 3 )
Coaiquer
Cayapa
Birds ("eallina de monte") /fruits/
Birds/eat fruits/ (4827 3)
Coaiquer
Bircls/eat fruits/ (4885 I )
Schleeelia dresleri
Ca,vapa
S. fastigiata
Cavapa
Birds/visit fl or,vers/( 48404)
B i r d s / v i s i t f l o w e r s / ( 4 8 1 11 )
Bromeliaceae
( ' r r z r n a n r ri a s c l r e r z e r i a n a
Oayapa
Opossums/eat leaves/ (48395)
Burmeistera sp. I
Cayapa
B.sp.2
Coaiquer
B .s p . 3
Coaiqtrer
Birds/visit fl owers/ (48 I 02)
Birds/eat fruits/ (48696)
Birds/eat fruits/ (487 07 )
Capparidaceae
Oapparis sp.
Oayapa
Squirrels/eat fruits/ (48l l9)
Clusiaceae
Clusia sp. I
Coaiquer
genus indet.
Coaiquer
Bircls/eat fruits/ (48932)
Birds/eat fiuits/ (48835)
genus indet.
Coaiquer
Rircls/eat fruits/ (48962)
R y ti d o s t rl i s c ar t h a g in c n s i s
Ca,vapa
Birds/eat fruits/ ( 4832U)
Cyclanthaceae
Asplundia sp.
Oavilp:r
B i r d s / e a t f n r i t s / ( 4 8 17 5 )
Colorado
Parrcrts/eat leaves/ (4007 7 )
Tetrathylacium macrophvllum
Cayapa
Birds/eat fmits/ (484211)
genus inclet
C:ryapa
Bircls/suck sap from fruits/ (48 I 7l )
Cayapa
Araliaceae
Schefflera spp.
Arecaceae
Chamaedorea pinn atifions
Hi'ospathe elegans
(48908)
Asteraceae
\&'ulffia baccata
Bignoniaceae
Campanulaceae
Cucurbitaceae
Euphorbiaceae
Cleiclion castaneifbliurn
Flacourtiaceae
Gesneriaceae
Drymonia coriaceir
Cayapa
Hummingbircls/visit
D. sernrlata
Cay'apa
Humrninebirds/r'isit
Cayapa
Htrnr rni n gbi rcls/r'isi t flowe rs an cl fnri tsl (4827 4)
fl orvers/ ( 48083)
flowers/ (4808I )
Heliconiaceae
Heliconia wagneriana
BS 46
86
Tabel32 - Continued
Tribe
Animals/ comments,/ (AAU voucher)
Discophora guianensis
Cayapa
Many different birds/eat fruits/ (48168)
Marcgraviaceae
Marcgravia sp.
Cayapa
Birds/ripe fruits/ / (48337)
Melastomataceae
Blakea punctulata
Coaiquer
B. punctulata
Coaiquer
B. subconnata
Conostegia montana
Cayapa
Miconia oraria
Tococa spadiciflora
Cayapa
T. symphyandra
Topobea calycularis
Coaiquer
T. caudata
Coaiquer
Mammals/eatbark/ (41613)
Birds/eat fruits/ (48987)
Birds,/visitfl owers,/(48336)
Birds/ eat fruits/ (48785)
Birds/eat fruits/ (48216,48339)
Birds/eat fruits/ (4870I)
Birds/eat fruits (48725)
Birds/eat fruits/ (48901)
Birds/eat fiuits,/ (48797,489I 8)
Menispermaceae
genus indet
Cayapa
Toucans,/eat frwits/ / (48237)
Moraceae
Ficus obtusifolia
Colorado
F. schippii
Cayapa
Birds/eat frgs/$00a7)
Many different birds/eat t\gs/ (48424)
Myrtaceae
genus indet.
Cayapa
Many different birds/eat fruits/ (48401)
Coaiquer
Lizards/ eat infl orescences/(4B9BO)
Birds,/visitinfl orescences/(48338)
Icacinaceae
Coaiquer
Coaiquer
Coaiquer
Piperaceae
Peperomia omnicola
Piper sp.
Cayapa
Rubiaceae
Cephaelis sp. I
Cayapa
C.sp.2
Coaiquer
Isertia pittieri
Palicouria gtrianensis
Cayapa
P..pp.
Coaiquer
Psychotria brachiata
P. racemosa
Cayapa
Cayapa
Birds/eat fruits/ (48208)
Birds/eat fruits/ (48904)
Squirrels/eatfmits/ (48278)
Hummingbirds/visit fl owers/ (48205)
Birds/eat fruits/ (41619,48721,48727)
Hummingbircls/visitflowers/ (48l 12)
Birds/eat fruits/ (482I 5)
Sapindaceae
Paullinia nobilis
Cayapa
Birds/visit flowers/ (48341)
Cayapa
Cayapa
Birds/eat fruits/ (48I 66)
Bird ("paleton")/eat fruits/ (48867)
Birds/eat fruits/ (489I 4)
Birds and squirrels/eatfruits/ (48405)
Cayapa
Birds/visit flowers/ (48306)
Coussapoa contorta
Coaiquer
Birds and fish/eat fruits/ (41626)
Zingiberaceae
Renealmia cuatrecassii
Coaiquer
Parrots,/eatfruits/ (415BB)
Solanaceae
Cestrum sp.
Lycianthes sp.
Solanttm triplincrr itrm
genus indet.
Tropaeolaceae
Tropaeolum repandum
Cayapa
Coaiquer
Coaiquer
{Jrticaceae
I
BS46
87
Table 33. Utilitarian poisons. For hunting, fishing, or pest control. The identity of the arrow poison tree of the Clolorados
is uncertain since u'e only obtained sterile material. Accordins to the Cayapa informants, a species of l{aucleopsi.s
has prev!
ouslv been rtsed for arrow poison. In tropical South America the common word for fish poison plants is "barbasco". Leaves,
stems roots or fiuits fiom these plants are crushed and thrown into the water. The fish are immediately paralyzed and rnay
eventrtally die of asphp<iation. The water is typically led through manmade dams on the riverbank (Fig. la). The fish are
collected at the outlets of these dams. The active compounds in the fish poison are saponins that are unharmful to humans.
Use/part of plant/comments/
(d{U
voucher)
Asteraceae
Clibaclium sp.
Colorado
"Barbasco "
/ leaves/ crushed/,/ ( 49095 )
Cannaceae
Canna sp.
Colorado
Previously used to kill enernies/sap/ verv poisono u s / ( 4 9 0 5 1)
Euphorbiaceae
P. anisolobus
Cayapa
P. anisolobus
Coaiquer
Sapium peruvianum
Cayapa
S. peruvianunr
Coloradcr
"Barbasccr"
/leaves/ crushecl,/(40010, 491 02)
/leaves/ crushed/ (40394, 48370)
"Barbasco "
/ leaves/ crushed/ ( 4BBB0)
"Barbasco"
and thrown in river verv effi/bark/ground
cient/ (48414,41041)
"Barbasco "
/ bark / cr ushed/ ( 49098)
Colorado
"BarbASCo"
/ root/ grou ndl (49 107 , 49705)
Phyllanthtrs anisolobus
Coloradcr
"Barbasco"
Fabaceae
Lonchocarpus sp.
Moraceae
Castilla elastica
Coloradcr
Ficus obtusifolia
Coloraclo
Naucleopsis amara
Coaiquer
N.rp.
Colorado
Latex trsed as apoison/ / causes an intense allergic reaction
on the skin/40096
Latex poisonous/ / / (40047)
Arrow poison/latex/ / (48972)
Previously rrsed for arrow poison/latex/ / (49112 )
Oxalidaceae
Biophytum dendroides
Cayapa
Polygonaceae
PolypJonum hydropiperoides
Oavapa
Prer,iouslv used in warfare/ / entire plant verv
poisonous( 40577)
"Barbasco"
/leaves/very strong/ (40494, 40497, 48412)
Solanaceae
Solantrm mamnlosum
Coaiquer
Insecticicle to control cockroaches/frtrit/
/ (41452)
Table 34. Notorious poisons. This cat.egory comprises all plants inclicated as being poisonous and not servir-rgan)' purpose.
Tribe
Clornments/ (AAU voucher)
Ooaiquer
(lauses rash particularlv in chilclren / (41433)
Solanum sp.
C)oaiqtrer
Fruits said to be poisonous/(48817)
Ijrticaceae
genus inclet
Cavapzr
Causes itchins of skin/ (4U050)
Oxalidaceae
Oxalis sp.
Solanaceae
I
BS 46
88
Table 35. Anaemia. Symptoms such a fatigue are often referred to as anaemia by the Cayapas. The treatment is tvpically s,vstemic.
Tribe
Part of plant/preparation/treatment/
Polypodium piloselloides
Cayapa
/ground,
(40302)
Tectaria nicotianifolia
Oavapa
Thelypteris sp.
Cai'apa
Leaves/ground and mixed with a cup of water/2 teaspoonfi.rlstaken 3 times aday/ (40411)
Leaves/ground, colcl extract/ / (48222)
Trichomanes elegans
Cayapa
Leaves/sround, hot water aclclecl// (40882)
Cavapa
[.e ave s,/bo ile d / bath / ( 48064 )
Cassia papillora
Oayapa
l,eaves,/ground rvith cold water/ massa€leor extract
drtrnk/ (40757,40543)
Senna dariensis
Cavapa
(AAU) voucher
Vascular crlptogams
mixed with tepicl water,/taken 4 times a day/
Begoniaceae
Begonia serniovata
Caesalpinaceae
Leaves,/qround in cold water/l
cup is taken at noon after
bath in the river/(41087)
Costaceae
Costus lirna
Piperaceae
Peperornia sp. I
Cayapa
Pith of stem/ground, mixed with water/cold bath/ (48093)
Cavapa
P. urocarpa
Cayapa
l.eaves/ground, heated in water// (40506)
l,eaves,/grouncl, boilecl/2 teaspoonfuls taken 2 tirnes a
d a y l ( 4 0 5 3 1)
Table 36. Blood circulation. Plants belonging to this categorv are used to treat symptoms allegedly caused by poor blood
circul:rti<rn. It is notable that one particnlar kind of syrnptom l-rasthe same narne as the snake Luchei,snruta,which produces
a hemol,vtic poison.
Bignoniaceae
(lrescentia cr{ete
Tribe
Part of plant/preparation,/
(AA.U voucher)
Colorado
Fruits/juice mixecl with salt/drtrnk/
circnlation / (40728)
Coaiqtrer
Sap/mixed rvith hot n'ater/drunk/cliilcl
rash/ (41504)
Coaiqtrer
Lear,'es/ground with a little water/ dmnk/sy'rnptoms
> g u a r n u c a < . (/ 4 8 9 3 0 )
Coaiquer
Root/qroun d with water/drunk/
ca" / GBq24)
treatrnen t/ comm en ts/
improves bacl blood
Sapindaceae
Allophyltrs exelsus
cliseasecausins
Solanaceae
Brugmansia versicolor
called
Zingiberaceae
Renealmia sp.
symptorns called >Ffuamu-
BS 46
89
Table 37. Gangrene. PIants in this category are trsed to treat various ailments rangine fiom minor afllictions to very seri11ls
health threats such as lymphingitis.
Tribe
Part of plant/ preparation/ trearmen r/ symproms/
(AA.U voucher)
Begonia semiovata
Cavapa
Leaves/ground eventually mixed with urine/compress/f'ev e r a n d s w o l l e n l e g s , p o s s i b l yb l o o d p o i s o n i n g / ( 4 1 0 2 9 ,
48276)
B.rp.
Ciayapa
Leaves,/ground, soaked in water with a Iittle urine/lorver
part of body washecl,upossibly eangrene/(48310)
Coaiquer
Leaves/ /rubbing
(414e6)
Cayapa
Leaves,/ground, rnixed with urine / batl-r/ ganurerne, blooclp o i s o n i n g / ( 4 8 0 9 7 , 4 8 14 6 )
Clayapa
Leaves/ /tc> bathe legs of old people rvith uangrene/ /
(40877)
Cavapa
Florvers/ground, heatecl/applied to wounds/to cure a disease causing heall' eczema and wounds / ( B35tr)
Begoniaceae
Bromeliaceae
Pitcairnia sp.
of skin/infection
of arreries (phlebitis) /
Costaceae
Costus spp.
Gesneriaceae
Monopyle sodiroana
Lamiaceae
Hyptis capitata
Table 38. Blood tapping. Blood tapping was probably once used by sorne shamans to cure various ailments. \4/e never witnessed such a practice, holever, ancl dotrbt that it is very comrnon. The Heliconiuleaf is tightly rolled to fbrrn a slencler
tube. This is placed near the incision in order to collecr the bloocl.
'Iribe
(AAU voucher)
Heliconiaceae
Heliconia lonsa
Cayapa
(48180)
BS 46
90
Table 39. Gums and teeth. The plants inclucled in this catesory are all used to fight inf-ectious conditions of the mouth or
to relieve toothache. No information was obtained regarding plants used for preventive measures.
and purpose/
Tribe
Part of plant/preparation/treatment
(AAU voucher)
Vascular cry)togams
Trichomanes polypoclioides
Coaiquer
Leaves/grotrnd to a paste/applied to aching gums/ (48839)
Amaranthaceae
Achyranthusaspera
Coaiquer
Leaves/ground in a cup of cold water/gargling fbr
toothache/ (41649)
Coaiquer
Rhizome and leaf:sheaths/clied or taken fresh/ftrr
toothache/ (41604)
Coaiquer
Flowers/crude/cherved
Cavendishia complecteus
Coaiquer
Leaves/boiled/placecl on.jaw f or toothache/ (a1502)
Menispermaceae
Cissampelos tropaeolifblia
Coaiquer
Leaves,/ground while fresh, mixed with fat,/placed on
aching gums,/ (41472)
Portulacaceae
Portulaca sp.
Cayapa
Leaves/crushed/appliecl
Zingiberaceae
Zingiber officinale
Colorado
Rhizome/ground,
Amaryllidaceae
genus indet
Asteraceae
Spilanthes sp.
for toothache / (41485)
Ericaceae
to aching g;ums/ (48379)
sap extracted/for
toothache/ (491 16)
Table 40. Liver and spleen. The informants did not make a clear distinction between the kiclney (urinary system, Table 69)
and the liver (digestive system). We have tried to separate the two categories here to reveal eventrral differences in rtsage
patterns for the two organ grottps.
Tribe
Part of plant/ preparation / treatntent/ purpose,/
(AAU voucher)
Vascular cryptogams
Polypoclium phyllitidis
Cayapa
Saccoloma elegans
Cayapa
Leaves,/ground, mixed with water/drr"rnk before meals/
pain, liver and spleen/(40806)
Petiole/soaked in water, slime extracted and mixed with wadiseases/ (48223)
Leaves,/ground, mixed with cold water/2 glassesa day,
morning and evening/pains in spleen and swelled liver/
( 4 0 3 18 , 4 0 8 6 7, 4 8 0 6 9 )
Vascular bundles extracted/soaked in water/inftrsion drunk
3 times daily/ swelling and pains of liver/ (4829fl)
terldrunk/liver
Thelypteris gigantea
Cayapa
T. reticulata
Cayapa
Araceae
3 tirnes claily/liver
Monstera sp.
Cayapa
Leaves/ground with cold water/drtrnk
pains/ (40436)
Asteraceae
genus indet.
Cayapa
Leaves/boiled/ I cup of hot mixture drunk/liver
(40574],
pains/
BS46
91
Tabel40 - Continued
Tribe
Part of plan t,/ prep ar atiorr / tr eatmen t,/ purpose/
(AAU voucher)
Cayapa
Sap/extracted/drunk
Colorado
Sap//drunk/liver
Cayapa
Leaves/ground in hot water,/45 tea-spoonfuls drunk at
dawn/liver pains/ (40564)
Colorado
l,eaves/ground, sap mixed with water/drunk/
ney diseases/ (40216)
Colorado
Latexlmixed
Commelinaceae
Tripogandra serrulata
cold/liver diseases/ (48089)
Costaceae
Costus sp.
and kidney pains/ (40678)
Lamiceae
Salviacoccinea
Monimiaceae
Siparuna sp.
liver and kid-
Moraceae
Brosimum utile
with waterldrunk/pains
in liver ar-rdkidney/
( 4 e r1 0 )
Piperaceae
Piper sp.
Colorado
Leaves/ground and mixed with water/ drunk/pains
ver/ (49043\
Colorado
Leaves/ground/drunk
/liver and kidney diseases/ (49044)
Lindernia crustacea
Colorado
Leaves/boiled,/drunk
cool/liver pains/ (40636)
Urticaceae
Pilea spp.
Colorado
Leaves/boiled or sap mixed with water/ drunk/liver
(40062, 40t07, 402t7, 40682)
in li-
Rubiaceae
P s y c h o t r i aw i l l i a m s i i
Scrophulariaceae
pains/
Table 41. Stomach infections. Symptoms accompanied by diarrhoea and abdominal pains. A potion is prepared from the
plant in nearly all cases.Diarrhoea is a common health problem, especially among children.
Tribe
Part of plant/preparation/treatment
(AAU voucher)
Vascular cryptogams
Asplenium sp.
Colorado
Bolbitis nicotiafolia
Cayapa
Campyloneuron sp.
Lomariopsis.japrrrenis
L. nigropaleata
Colorado
Leaves/ground in cold water/drunk,/ (49032)
Leaves/ /warm extract drunk 3 times a day/stomach pains
in connection with colds, particularly for children/ (48219)
Leaves,/ ground/ drunk / (49037 )
Cayapa
Leaves/ / cold extract drunk/ (401 63)
Vascular bundles/boilecl/drunk
3 times a day to treat stomach pains of children / (48284, 48125)
L. nigropaleata
Colorado
Rhizome/ground
Pityrogramma calomelanos
Coaiquer
Thelypteris torresiana
genus indet. (Polypodiaceae)
Cayapa
Colorado
and boiled /drunk/
Leaves/boiled/drunk / ( 48752)
and purpose/
(40020)
Coaiquer
Leaves/body massage/for diarrhoea/ (481 30)
Leaves/boiled/drunk / (487 57 )
Cayapa
Leaves/ground, mixed with a little water / drunk/ (40398)
Acanthaceae
Sanchezia parviflora
BS 46
Tabel4l
- Continued
and purpose/
Tribe
Part of plant/preparation/treatment
(AAU voucher)
Cavapa
Leaves/ground, mixed with cold water/drunk/
Eryneium foetidum
Colorado
Root/boiled/drunk/
(40038)
Aristolochiaceae
Aristolochia sp.
Cayapa
Stem/ground/drunk
a day/ (40523)
with sttgarcane ruln, 2 teaspoons twice
Colorado
warm / (4003L\)
Roots/boiled/drunk
l.eaves/sap / / (40037)
f.eaves/boiled/ drunk / (40647 )
Amaryllidaceae
Crinum amabile
(40547)
Apiaceae
Asteraceae
senus indet.
genus indet.
genus indet.
genus indet.
Colorado
Coloradcr
Colorado
Leaves/sap squeezed ottt and boiled/ ab<lttt I slass drunk,/
(40707)
Bignoniaceae
Schlegelia sp.
Cayapa
Flowers/grorttrd with water/drunk/
Bromeliaceae
Guzmannia linsulata
Cayapa
Leaves,/boiled/drunk
(48377)
(iuzmaniamelinonis
Cayapa
G..p.
Cayapa
genus indet.
Cavapa
Leaves/boiled/drunk befbre meals/ (40885)
/ /drunk 3 times aday/ (41019)
Leaves/boiled,zsmall cup drunk 3 tirnes clatly/ (48396)
Chenopodiaceae
Chenopodium ambrosioides
Cayapa
/bolled/7
Commelinaceae
Tripogandra serrulata
Ca,vapa
Leaves/ground, sotne water aclded/extract taken 3 times
daily before meals/ (41032)
Costaceae
Costus sp.
Ooaiquer
Flowers/ground/taken
Cucurbitaceae
Gurania sp.
Cayapa
l,eaves,/decoction from fresh leaves/ drunk in the morning
and the evening/ (48107)
Erythroxylum novogranatense
Coaiquer
Leaves/infusion/drunk/
Flacourtiaceae
genus indet.
Coaiquer
L,eaves/crtrshed,juice dissolved in rvater,/
(40329)
colcl in the morning befbre eating/
ctrp drunk / (40575,40857)
with a little water/ (1I457)
Erythroxylaceae
(4 I 584)
dmnk/(41509)
Gesneriaceae
Alloplectussprucci
Colorado
Leaves/ground, water aclded/drunk/
flaemodoraceae
Xiphidium caerttleum
Cayapa
l,eaves,/grortnd, mixed with warnr water,/drunk/
(49052)
G0374)
BS 46
Tabel4l
93
- Continued
Tribe
Part of plant/preparation/treatment
(AAU voucher)
Hyptis obturiflora
Cayapa
Ocimunr campechianum
genus inclet.
(loloradcr
genus inclet.
C:ryapa
Leaves/qround, water added,/dmnk
adults/ (10294\
L e a v e s / b o i l e d / d r u n k h o t l ( 4 0 6 16 )
/ bolled / drunk/ (40854)
(4086 I )
Leaves/boiled/drunk/
and purpose,/
Lamiaceae
Cavapa
4 times a day, only by
Loganiaceae
Spiuelia sp.
Cavapa
Leaves,/ground and boiled in a little water/drunk
day, onlv by children / (40293)
Colorado
l,eaves/bolled/at
Clidemia discolor
Cayapa
Leaves/mixed with a little tepid warer/ clrunk 4 times a
dav/ (10307)
Orchidaceae
Vanilla odorata
Clayapa
Leaves/boiled/extract
evening,/(48236)
Peperomia sp.
Piper multiplinerviurn
(krlorado
/ / a cold clrink made / (40137)
Roots/boiled/drunk/ (40059)
P. sp. I
P. sp. 2
Coaiquer
(lolorad<r
Leaves/boiled/drunk / (41 600)
Leaves/ ground, water addecl,/dmnk fiesh/ ( 49043 )
Rubiaceae
Bertiera guianensis
(bloradcr
Stenr/boile rl/ drunk/ (40069, 40156)
Leaves/ground, water added/dnrnk fresh/ (49044)
/ground with tepid warer/drunk/
90322)
3 tirnes a
Loranthaceae
genus indet.
least I cup dnrnk / (40634)
Melastomataceae
taken cold in the morning ancl the
Piperaceae
P s y c h o tr i a r v i l li a m s ii
P.rp.
Coloraclo
Colorado
Oayapa
Scrophulariaceae
Lindernia sp.
genus inclet.
Coaiquer
genus indet.
Cav:rpa
Lear.'es/boiled for 2 hours/drunk
d a y l ( a 10 2 8 )
genus indet.
Oavapa
genus indet.
Cia,vapa
,4BI48)
/boilecl in ca. 3 liter warer/dmnk over a day/
Leaves/qround, mixed with warm water/drunk, onlv by
aclults/ (40300)
Cavapa
l,eaves,/clecoction or infusion / (48722)
/ bolled / dnrnk/ (408 12)
at regular interr,uls cluring
Solanaceae
Solanum canense
Coloradcr
[,eaves,/sapsqueezed out and rnixed with hot water/clrur-rk/
(40016, 40724)
S. leptorhachis
Cloloraclo
Sterns/boiled/dnrnk/
S. confertiseriatlun
Coaiquer
Lezrves/sround/extract
Oavapa
Leaves/boiled and mixecl with 2 orher plants/drunk
still hotl(41007)
(40073)
dmnk/
(48695 )
Verbenaceae
genus indet.
while
BS 46
94
Table 42. Antiemetics. Prevent nausea and vomiting'
Tribe
Part of plant/ prep ^r ation / tr eatme n t/ commen ts/
(AAU voucher)
Acanthaceae
Ruellia riopalenquensis
Cayapa
Leaves/ground to a paste/mixed with wat.er and drunk for
3 days/for vomiting with blood / (40365,48238)
Boraginaceae
Cordia spinescens
Coaiquer
Leaves/ground in water,/extract drunk/ / (41498)
Lamiaceae
Hyptis verticillata
Cayapa
Leaves/ground in water/ a small cup drunk 3 times a
day/ only for children/ (4 I 083)
Poaceae
Coix lachryma-jobi
Coaiquer
Fruits/crushed/taken
(48716)
Scrophulariaceae
genus indet.
Cayapa
Leaves/bolled/3
Verbenaceae
Verbena litoralis
Cayapa
Leaves/ground, mixed with water/drunk/
spoonfulls drunk/ / (48444)
Table 43. Laxatives. It is notable that the Colorados use the strongly hallucinogenic
Euphorbiaceae
Jatropha curcas
/ (40401)
liana Banesteriopsiscaapii as a laxative'
treatment/ (AAU voucher)
Tribe
Part of plant/preparation/
Colorado
Fruits,/ crushed,/ drunk / (40633)
Colorado
Stems/ground and boiled/some glassesof mixture drunk/
(40722)
Malpighiaceae
B a n i s t e r i o p s i sc a a p i
with a small amount of water/ /
BS46
95
Table 44. Intestinal parasites. Intestinal parasites such as tapeworm (Thenia sp.) and roundworm (Ascaris lumbricoides) are
common health problems especially among children. In this category there are several examples of the Doctrine of Signature. One example is Monolena primulae.flora,which is probably used because the petiole resembles a red worm.
Tribe
Part of plan t/ prep aration / tr eatmen t/ commen ts/
(AA,U voucher)
Vascular crn)togams
fuplenium sp.
Dicranopteris pectinata
Thelypteris exuta
genus indet. (Polypodiac.)
genus indet. (Dennstaedtiac.)
Coaiquer
Cayapa
Leaves/ /extract
drunk / / (48744\
////(48323)
Coaiquer
Leaves/boiled,/drunk / / (48804)
Young leaves/boil ed / eaten / / (41617 )
Leaves/ground/drunk
with a little water / / (48840)
Coaiquer
/ bolled / dr unk / / ( 488 I 9 )
Colorado
Fruits/sap squeezed outldrunk
Monolena primulaeflora
Coaiquer
Petioles/ / eaten fresh/intestinal worms killed or expelled by
vomiting/ (41 582, 48974)
Moraceae
Ficus insipida
Cayapa
Latex/ /3-4 drops of latex taken with a teaspoon of water
twice a day/ / (40900, 48056)
Cayapa
l,eaves/boiled/5
Coaiquer
Coaiquer
Begoniaceae
Begonia glabra
Euphorbiaceae
Jatropha curcas
/ / (40633)
Melastomataceae
Scrophulariaceae
Scoparia dulcis
Table 45. Deficiency symptoms. The informants
balanced diet.
teaspoons of decoction drunk at dawn/intestinal worms expelled by vomiting/ (40567)
related that the symptoms treated in this cataegory were caused by an un-
Tribe
Part of plant/preparation/treatment/
Mikania sp.
Coaiquer
Leaves/ ground/juice
Passifloraceae
Passiflora auriculata
Colorado
P. quadrandularis
Colorado
Sap,/mixed with sugar cane brandy/drvnk/
Leaves/ /drunk hotl (40174)
(AAU voucher)
Asteraceae
I
drunk with water/ ( 4 I 50 I )
(40105)
BS 46
96
Table 46. Ant bites. The large and aggres sive conga ants (Parapon?.rasp.) have a painful bite that is treated with plants lrom
the families Araceae and Cyclanrhaceae. The crude sap is squeezed from the stem and applied directly to the bite.
Tribe
P art / pr epar ation / (AAU vouche r)
Dieffenbachia seguine
Cayapa
Philodendron inequilaterum
Colorado
Leaves/sap extracted from petiole close to the leaf base/ (407 56, 48754, 48348)
L,eaves// (40171)
P'rp.
Colorado
Svngonium crassifolium
Cayapa
genus indet.
Cayapa
Leaves// (40115)
Leaves/eround to a paste used as ointment/(40330)
L e a v e s // ( 6 0 1 0 1 )
Cyclanthaceae
Cyclanthus bipartitus
Evodianthus firnifer
Cayapa
Colorado
Root/ / (48078)
Roots// (40050)
Tribe
Part of plant/preparation/treatment/comments/
(AAU voucher)
Cayapa
to absorb pus from bites on
Leaves/ground/compress/said
"nacido") (40480)
/
less and arms (unidentifiecl insect called
Piper augustum
Cayapa
P. tricuspe
Cayapa
hot compress/ removes larvae from
Leaves/ground/humid,
"t6bano") (40303)
skin (disease called
/
Leaves/ground/hair wash/removes lice and heals a lice
"rastransmitted fungal inf'ection that cattses a rash (called
P. tuberculatum
Cayapa
Araceae
Table 47. Insect bites. General.
Cucurbitaceae
Melothria perdula
Piperaceae
quinia")/
(40859)
Leaves/ground/cold
bath and hairwash/kills lice,/ (40455,
48375)
Solanaceae
Cestrum sp.
Colorado
Lycianthes amatitlanensis
Colorado
Leaves/borled/ /for bites in general that itchl (40164)
Leaves/gJrouncl/compress/generally for bites that itch/
(40t72\
-'l
f
BS46
97
Table 48. Snake bites. Topical. Under purpose are listed the specific eff'ects that the treatment has according to the informants, e.g. to absorb the venom, reduce the swelling, stop bleeding, relieve the pain, prevent infections or to cool the infected area down. See also snake bites under table 49, and table 50. The vernacular names for snakes probably corresponcl
to the following scientiflc names: ueryugos(l,guamaur or oalcama = Lacheis muta ; papagayo, dormilona = Bothro7s schlegeLii; equis
= Bothro|s ntrox; coral= Micrutus sp.;
Nrudredora=BothroPsbilineatus;uibora= Bothroqsnasuta.The coralsnake (Fir. 16) has a
neurotoxic poison and electric shocks fiom generators are sometimes used as part of the therapy.
Tribe
Part of pl an t,/ pre p ar ation / tr eatnien t/ purpose / snake /
(AAU voucher)
Adiantum obliquurn
Adiantum sp.
Cayapa
Asplenium sp.
Blechnum sp.
Colorado
ress/ / / (48 I 33)
bath / / / (40689)
Leaves/boiled/warm bath / / / (40697)
Leaves/ boiled/ tepid bath / / " e qtris"/ (4Bg 12)
Leaves,/boiled/cool compress/swelling/ "c oral" / ( 48844)
/boiled/coldbath/
/ / (40145)
Leaves,/boiled/tepid bath/ / "equis"/ (4894I )
Leaves/ground/sap used for compress/ / / (40649)
/boiled/coldbath/
/ / (40168)
Leaves/bo lled / bath / / " equis" / (487 5 4)
"
/ bolled / c ool comp r e ss/ / papagallo" / ( 48834 )
Leaves/boiled/comp r ess/ / " coral" / GBB46)
Leaves/bolled / bath / / " eqtris"/ (48998)
Leaves/boilecl/cool compress/ swelling/ "c ora|" / ( 48843 )
Leaves/ / c.old bath / / / (40196, 402 19)
"
/ bolled / barh / / coral" / (487 65)
Leaves,/ground/comp ress/ / " eqlis"/ (40878)
Vascular cryptogams
Campyloneuron sp.
Ctenitis sp.
Colorado
Coaiquer
Coaiquer
Colorado
Danaea sp.
Coaiquer
Dicranoelossum polvpodioides
Diplazium sp.
Colorado
Hymenophyllum
sp.
Lycopodium sp.
Nephrolepis sp.1
N. sp.2
Colorado
Coaiquer
Coaiquer
Coaiquer
Oleandra sp.
Ooaiquer
Ooaiquer
Polypodium percussum
Colorado
Schizaea sp.
Coaiquer
Selaginella sp.
Cavapa
Stigmatopteris sp.
Tectaria sp.
Ca,vapa
Leaves/ground/comp
Leaves/boiled/warm
T. gigantea
Trichomancs collariatum
Coaiquer
T. elegans
Coaiquer
T. punctatum
T. sp. I
Cayapa
T. sp.2
C)ayapa
Ooaiquer
/bolled/bath/
/"equis" / (48300)
Leaves/bolled / bath / / " equis" / (48917)
Leaves/ ground/compress/ absorbs venom/ "dormilona"/
(48294\
Leaves,/boiled/ compress/ / "equ is"/ (4 | 435A, 4BBII5)
/boited/bath/ / / (40695)
Leaves/ ground/compress/ / "cor al"/ (487 59)
Leaves,/ground/compress,/ swelling/ "dormilo na" / (483 45)
[.eaves/boiled/hot compress/ / / (41630)
Leaves/ ground/compress/ swelling,/ "dormilona"/ ( 48408)
Leaves/bo lled / bath / / " coral" / ( 4U950 )
Cayapa
Le aves/ ground/ compress/ swelling / v1bor a / ( 40890 )
Coaiquer
Corm/grounded,/com
Anthurium bakeri
A. scandens
Coaiquer
Monstera clubia
Oavapa
M. spruceana
Cayapa
Philodendron sp.
fiavapa
Svngonium sp.
Xanthosoma daguense
Cayapa
genus inclet.
Cavapa
Leaves,/boiled/compress / / / (11 417)
Leaves,/boiled/ massage/swelline / / (40 465)
Leaves,/ground/compress/absorb venom/ "equis"/ (48 10b)
"equis"/
Leaves/uround/compress/absorb
(40843)
venom,/
Lear,es/ boiled / massage/ swelling,/ "e quis "/ (483 47 )
Leaves/qround in cold water/compress/ /"equis" / (40768)
Leaves/ ground/ compress/absorb venom/ "equis "/ (4827 0)
[,e aves/ boiled/ rn assage/ swellin g / " e quis" / ( 40 7Bg)
Thelypteris francoana
genus indet. (Gleicheniac.)
Coaiquer
Oayapa
Colorado
Coaiquer
Amaranthaceae
Nternanthera sessilis
Amaryllidaceae
genus indet.
press/ / / (487 12)
Araceae
Oayapa
Cayapa
BS 46
98
Tabel48 - Continued
Tribe
Part of plant/ preparation/ treatment/purpose / snake/
(AAU voucher)
Asteraceae
Mikania sp.
Colorado
genus indet.
Colorado
Leaves/ / compress/,/
Leaves, flowers/bolled
Bignoniaceae
Schlegelia sulfurea
Colorado
/botled/bath/
Bromeliaceae
Tillandsia sp.
Cayapa
/ ground / compress/ /
Cayapa
Cayapa
"equis"/ ( 48 I BB)
Leaves/ ground/ compress,/ swellin g,/
"equis"/ (48090)
/ ground/compress/ /
Costaceae
Costus pulverulentus
Colorado
Roots/boiled/hot
Cyclanthaceae
Asplundia sp.
Colorado
Stem/boiled/bath/
Cayapa
Leaves,/bo tled / bath / /
Commelinaceae
Dichorisandra hexandra
genus indet.
Ericaceae
"puclreclora"
/ (40348)
/ bath / / / (407 02)
/ / (40127)
"equis"/ ( 40t169
)
bath/ / / (401 16)
/ / (40118)
"
Coaiquer
eqwts"/ ( 48376)
"equis" (487 7 7
)
/
Leaves/boiled/compress/ /
Dalechampia sP.
Cayapa
Leaves/boiled/ massage / /
Fabaceae
senus indet.
Cayapa
/ boiled / massage / /
Macleania rotundifolia
Sphyrospermum sP.
Euphorbiaceae
Gesneriaceae
"
"v
errusosa"/ (482 I 3 )
equis"/ ( 48 I 95 )
"
Leaves/bo iled / bath / / eqwis"/ (48 I 20)
Leaves/boiled/ compress/ absorbs venom and lower swelling,/"verrusosa" / (40844,41081, 48082)
/ (48732, 48813)
Leaves/boiled/compress//"equis"
"
Leaves,/bo lled / / / coral" / (487 34)
(4101 I'
/ ground,/ compress/pain, swelling,/"vernruosa"/
AlloplectusPanamensis
Cayapa
A. sprucei
Cayapa
A. tetragonoides
Besleria barbata
Coaiquer
B. barclayi
Cayapa
B.rP.
Coaiquer
Chrysothemis friedrichsthaliana
Cavapa
(lolumnea bilabiata
Cayapa
C. densibracteata
Cayapa
O. angustata
Colorado
C. dissirnilis
C. kienastiana
Cayapa
Cayapa
Leaves/boilecl/compress and massase/absorb venom/
"eqtris"/ (48084)
Leaves/boiled/warm barh / / / (40709)
"equis"/ (48 I 7B)
Leaves/ ground/compress/ /
"equis"7'
Leaves/mixed with sugar cane brandy/clrtrnk/ /
C. eburnea
Cayapa
(48383)
Leaves/ ground/ compress/bleeding/
C. fililoba
Coaiquer
Coaiquer
48415)
Le aves/ boiled/ compre ss/ swellin g,/ / (41 427 )
"verrueosa"/ ( 4036 1,
Leaves/ grouncl/ compress/ swelling,/
48080)
f,eaves/ grouncl/ compress/ pain, swellins, absorb venom'/
"equis"/ (40519, 40600, 40816, 48416)
"equis " (4087 0,
/
48218)
Leaves/boiled or fresh/compress, cold bath/ /
A1641,48997)
"equis"
/
BS 46
99
Tabel48 - Continued
Tribe
Part of plan t/ prep ar ation / tr eatment/ purpose / snake /
(AAU voucher)
C. parviflora
Coaiquer
C. picta
Cayapa
C. picta
Coaiquer
C rubriacuta
Colorado
"
Leaves/fre sh / bath / / corzl" / ( 48937 )
Leaves/ground, boiled/compress or bath/stops bleeding,/
"equis"/ (40760,
40871, 48086)
"equis " (487
Leaves,/fresh/,/
30)
/
Leaves/boiled or sap extracted/compress or bath/ / /
(40074,40675)
C. rubriacuta
Cayapa
C. rubribracteata
Cayapa
C. spathulata
Cayapa
c.rp.
Coaiquer
Cremosperma hirsutissimum
Coaiquer
C. nobile
Coaiquer
C. reldioides
Diastema affine
Coaiquer
Drymonia alloplectoides
Cayapa
D. coriacea
Colorado
D. macrophylla
Colorado
D. rhodoloma
Colorado
D. serrulata
Cayapa
D. serrulata
Coaiquer
D. turrialvae
Coaiquer
D. warszewicziana
Colorado
D. warszewicziana
Cayapa
D. warszewicziana
Coaiquer
Gasteranthus corallinus
Cayapa
G. crispus
Gloxinia dodsonii
Colorado
Leaves/ground or boiled,z bath or compress/antivenom,/
" e q u i s " , /( 4 8 1 2 1 ,
4 8 3 1l , 4 8 4 1 9 )
"
Leaves/bo tled / bath / / equis" / (4899 I )
Leaves/ ground/ compress/venom / " e quis" / ( 48094 )
Leaves/boiled/bath/ / / (40701)
Coaiquer
Leaves/boiled/hot
G. dodsonii
Cayapa
Kohleria x hybrid
Cayapa
Monopyle sodiroana
Cayapa
Leaves/ground or warmed/compress on bite, changed 3
times a day/ /"equis" / (40513, 48727)
Neomortonia rosea
Coaiquer
Paradrymonia sp. I
P .s p . 2
Coaiquer
Coaiquer
genus indet.
Cayapa
Leaves/boiled / bath / / / (4147 7)
Leaves/boiled/compress/ / "equis"/ ( 48963)
Leaves/fre sh / bath / / / (41635)
Leaves/fresh/massage or bath/ / "equis"/ (4I005)
Cayapa
/ / bath/ /"equis"/ (48417)
Coaiquer
Leaves/boiled/comp
Coaiquer
Leaves/boiled or ground/compress/stops bleeding, lower
swelling and absorbs venom/"verrrogosa" and "equis"/
(40872,40880, 49739)
Leaves,/ground in alcohol / bath / / "equis " / (48 420)
"equis " (407
Leaves/ground/compress/bleeding/
97 )
/
Leaves/boiled,zhot compress/ / " equrs"/ (48739)
Leaves/boiled or fresh/compress or bath/ /"coral" / (47422,
41437,48737)
Leaves/boiled/compress/ / "equis" / (4897 6)
Leaves/boiled/comp r ess/ / " equis"/ (48980 )
"equis"/
Leaves/boiled,/compress/ /
( 48990)
Leaves/boiled or ground/compress or massage/stops
"equis"/ (48212)
bleeding/
Leaves/bolled / bath / / / (4067 3\
Leaves/boiled / bath / / / (40167 )
/boited/ bath / / / (40108)
Leaves,/ground or mixed with sugar cane brandy/compress
(to stop bleeding) or drunk, highly estimated/ / / (48393)
Leaves,/ground/compress/ / "equis" / (487 7 3)
Leaves/boiled/compress / / / (41 638)
/boiled/ bath / / / (40080)
compress/ / / (41462)
Leaves,/ground/compress/ / "eqris" / (4827 9)
Leaves/ground or boiled/compress or warm bath, applied
twice to bite, highly valued/ / / (40576)
Haemodoraceae
Xiphidium
caeruleum
Lamiaceae
genus indet.
ress/ / / (41 41 6)
BS 46
100
Tabel48 - Continued
Tribe
Part of plant/ preparation/ treatment/ purpose/ snake/
(AAU voucher)
Loganiaceae
Spigelia anthelmia
(iayapa
S. anthelmia
Colorado
"
Leave s/ ground/ compress/ swellin g,/ e qfis" / ( 4U3BB)
(40142)
/ /coldbath/ / /
Melastomataceae
Aciotis caulialata
Cayapa
A. sp.1
Cayapa
A. sp. 2
Cayapa
Blakea subconnata
Clidemia crenulata
Coloraclo
Miconia sp.
Coaiquer
Pilocosta oerstedii
Ca,vapa
Triolena barbeyana
Cayapa
T. obliqua
Coaiquer
T.tp.
Coaiquer
Orchidaceae
Erythrodes ecuadoriensis
Cayapa
Cranichis sp.
Coaiquer
Scaphyglottis prolif'era
Psygmorchis pusilla
Clayapa
Cayapa
Cayapa
Piperaceae
"verrlrsosa" (407
45)
/
/ / compress/ swelling/
"dormilona",/
venonr/
Leaves/ground/compress/anti
(48316)
"dormilona"/ (41
0 16)
Leaves/ground/conipress,/swelling,/
Leaves,/ground/comp ress/ / / (40049)
"
Le aves/ ground/ cornpre ss,/swell in g / e quis" / (4807 7 )
"equis"/
cooling/"coral",
Leaves/boiled/compress/fbr
(48708)
"dorm
rlona" / (48387)
Leaves/ground/cornpress/infection/
"verntgclsa"/
Leaves/ ground/compress,/infection, pain/
(40344,41084)
Leaves/boiled/hot compress/ / / (41642, 41418)
Leaves/boilecl/compress / / / (41 434)
"
"ve
r r trgo sa / ( 40 7 BB)
Le aves/ gro u n d / c o rn p re ss/,/
"slramaca" (18927)
Leaves/fre sh/ bath/ /
/
"verrugosa",/ (
48 I 96)
Leaves/boiled,/ compress/ /
"eqrt
is"/ (41043)
Leaves/ ground/ compress/ an tivenom/
Peperomia macrostachya
P. serpens
Oayapa
P. serpens
Coaiquer
P. sp. I
Cayapa
"equis " (48202)
Leaves,/boiled/ massaee/ /
/
"eqtris"/ (482
10)
Leaves/sround/compress/swelling/
"equis "/ (
4 14 19, 4flil25 )
/ bolled / compress/ /
"
Leaves/boiled/coId bath / / ecluis"/ (48369)
P. sp. 2
P .s p . 3
P .s p . 4
P .s p . 5
P. sp. 6
P. sp. 7
P.sp. B
Piper angustum
P. filistylum
Coaiquer
Leaves/bolled/
P.florencianum
P. squarnulosuln
P.sp. I
P.sp. 2
P .s p . 3
P .s p . 4
P .s p . 5
P. sp. 6
P.sp. 7
P. sp. B
P. sp. 9
Cayapa
Cayapa
Coaiquer
Coaiquer
Coaiquer
Coaiquer
Cayapa
Coaiquer
Cayapa
Cavapa
Cclaiquer
Cayapa
Oayapa
Ooaiquer
Coaiquer
Cayapa
Cayapa
Cayapa
Coaiqr.ret'
Coaiquer
bath/ / / (487 06)
"
Le ave s,/fre sh / cold bath / / c,ctral" / (487 19)
"coral" (4il869)
Leaves/fre sh /bath / /
/
/ground/ cornpress// / (41625)
/ b<>lled/ compress / / / (41 645)
"equis "/ (
48 I 93 )
Leaves,/sround/ compress/ swelling/
"equis", "oalcama"/ (48906)
/boiled/ co mpress/ttsed cold/
"vibo
ra" / (10 452, 40738 )
f.eaves/boiled/ compress/ /
"viboLeaves/ground or boiled/cornpress/ venom,/"coral",
ra" / (40746, 481 16)
Leaves/rvarmed in hot water,/compress//"equis"/ (40338)
Leaves/fresh'/ bath / / / (11640)
"eqtris" (407
b2)
Leaves/boiled/cornpress/ swelling/
/
"equis"/ (
40793)
Leaves,/boiled/ compress/ swelling/
Leaves/boiled/compress / / / (41 420)
Leaves/boiled/compress / / / (41421)
"eqtris"/ (48
I 13)
Leaves/boiled/compress/ /
"
l,eaves/ boiled/ hot compress/ / equis" / ( 48 I 5 7 )
"
Leaves/ eroun d / com press,/veno m / equis" / ( 48 346 )
"eqtris"/ (
48824)
/ botled / c<trnpress//
"coral" (4tt85tt)
/
/bolled / cold r:ompress / /
BS 46
101
Tabel48 - Continued
Tribe
Part of plant/preparation / treatment/purpose/
(AAU voucher)
P. sp. l0
Cavapa
P.sp. 11
Ooaiquer
P. sp. 12
P. sp. 13
Trianaeopiper sp. I
Coaiquer
[.eaves/boiled/ bath/ /"verrugosa" / (40369)
l,eaves,/boiled/cool bath/ / "eqttis" / (48724)
"equis"/
Roots/eround / bath / /
(48909)
T. sp.2
Cayapa
Leaves,/boiled/hot compress/ / / (4l59g)
Leaves,/boiled/ compress/r'enom / " equis"/ ( 4 I 068 )
Leaves/ground/cornpress,// "dormilona"/ (48 I 6l )
Colorado
Roots/boiled/
Cayapa
"equis"
Leaves/sround/compress/
/
/ ft8230)
"
Leaves/boiled/ cold compress/ / c.oral"/ ( 48863)
Coaiquer
Cayapa
snake/
Rubiaceae
Sabicea villosa
bath / / / (40184\
Scrophulariaceae
senus indet.
genus inclet.
Ooaiquer
Solanaceae
Lycianthes amatitlanensis
Coloradcr
Solanurn medusocallx
Coaiquer
S. sp. I
Coaiquer
S .s p . 2
{renus indet.
Cayapa
Cayapa
/bolted / bath / / / (40087)
Leaves/fie sh / bath / / / (41628\
Leaves/boiled/hot compress/ / / (41648)
Leaves/ground/compress/ swelling,/ "equis"/ ( 48406)
Leaves,/boiled/ massage,// "equis " / (48240)
Colorado
Leaves/ / bath / / / (40708)
Cayapa
l.eaves/grorind/comp
Cayapa
Leaves/boiled/cool
Theophrastaceae
Clavija egeersiana
IJrticaceae
Pilea sp. I
ress/ / / (40378)
Zingiberaceae
Renealmia oligosperma
bath / / / (401 60)
Table 49. Snake bites. Systemic. Decoctions drunk for snake bites. lrrylftrodes weberanaand Kohleria x @lridwere claimed to
be particularly valuable. The latter is cultivated both by the Colorados and the Cayapas and it is often sold. See also snake
bites under Tables 48 & 50. For snake nanles, see the legencl of Table 48.
Tribe
Part of plant/preparation/
(A{U voucher)
Asplenium sp. I
Colorado
A. sp. 2
A. sp. 3
Coloradcr
Rhizomes/boilecl/dru nk / / (4007 5\
Leaves/boiled/ / / (40697)
Rhizome/boiled/drunk/
/ (49035)
Dicranoglossum polypodioides
D. polypoclioides
Colorado
Colorad<r
trearment/snake/
Vascular cryptogams
Clolorado
D..p.
Colorado
Trichomanes collariatrrnt
€ienus inclet. (Polypodiaceae)
Colorado
Leaves/boilecl/drunk / / (40170)
Leaves/ground/.juice drunk/ / (40649)
/bollecl/ clru,nk/ / (40234)
Cloloradcr
/boiled/ drunk warrn/ / (40695)
Leaves,/ground/drunk / / (49029)
Colorado
Rhizomes/boiled/dru
Costaceae
Costus pulverulentus
nk / / (40176)
BS 46
r02
Tabel49 - Continued
Tribe
Part of plant/ prep aration / tr eatment/snake/
(AAU voucher)
Gesneriaceae
Alloplectus dodsonii
Cremosperma hirsutissimum
Colorado
Coaiquer
Columnea angustata
Colorado
Roots/boiled,/ drunk/ / (40053)
Leaves/ground in alcohol/drunk/
/ (41426)
Root/boile d / drunk/ / (49106)
Leaves/ground/teaspoon taken twice a day/ / (48178)
drunk twice a day/"equis" / (48218)
Leaves/ground/extract
C. dissimilis
Cayapa
C. eburnea
Cayapa
C. picta
Colorado
C. purpurimarginata
Cayapa
C. rubriacuta
C. rubriacuta
Colorado
C. rubribracteata
Cayapa
C. spathulata
Drymonia coriacea
Cayapa
Cayapa
Cayapa
D. serrrulata
Colorado
Cayapa
Gasteranthus crispus
Colorado
G. oncogastrus
Kohleria x hybrid
Colorado
D. rhodoloma
Cayapa
Melastomataceae
Triolena barbeyana
Colorado
T. pustulata
Colorado
////(40230)
3 times a day/
Leaves/ground, mixed with water/drunk
"equis"/ (48085)
Leaves/ /decoction or sap drunk/ / (40074)
Leaves/ground/cold extract drunk 3 times a day/"equi s " , /( 4 0 8 8 0 , 4 8 0 8 8 , 4 8 1 2 8 )
"equis" (48420)
with alcohol/
Leaves/ground/drunk
/
"equis"/ (48344)
Leaves/ground, soaked in alcohol/ drunk/
Leaves/ground, mixed with water/cold extract drunk 3 times a dayl"equis" / (48092)
Leaves/borled/ / / (40108)
Leaves/ground, mixed with water or alcohol/extract
"equis"/ (48087, 48393)
3 times a day/
Leaves/bolled/ / / (40701)
/ (40233)
/ /drunkwarm/
Leaves/ground or boiled/2
te/ / (40576,40723)
drunk
teasPoons taken twice after a bi-
Leaves and roots/boiled,/drunkhot/
Leaves/boiled/drunk / / (40071)
/ (40094' 40157)
Orchidaceae
Erythrodes weberana
Cayapa
Stelis sp.
Cayapa
Leaves/ground, mixed with alcohol/ drunk/ / (40879)
"equis"
/
Leaves/boiled/drunk once a day in the mornin g/
(48247)
Oxalidaceae
Oxalis microcarpa
Colorado
Leaves/boiled/l
Coaiquer
/ /iuice
cup drunk / / (40726)
Piperaceae
Peperomia sp.
drunk/ / (41645)
BS46
103
Table 50. Snake bites. Others. Plants in this category were indicated as useful for treating snake bites but the informants
did not elaborate further on the treatment . The Cayapas use some yet unidentified saplings of shrubs or trees to prevent
snakebites.See also snake bites under Tables 48 & 49. For snake names, see the legend of Table 48.
Tribe
commen t/ (AAU vorrcher)
Campyloneuron angustif olium
Nephrolepis pectinata
Pteris pungens
Colorado
Selaginella sp.
Coaiquer
Ground and boiled / (49039)
Previously used/ (481 35)
/ (48283)
"equis"/ (48831
Boiled ancl used for
)
Dracontium sp.
Coaiquer
Rhizome dried, grounded with alcohol/ (11597)
Gesneriaceae
Columnea nariniana
Coaiquer
Boiled, decoction used for
Coaiquer
B<riled, used for
Peperomia sp.
Coaiquer
Piper sp.
Coaiquer
Leaves boiled and used for
L e a v e sb o i l e d / ( 4 1 6 3 7 )
Vascular cryptogams
Cayapa
Cayapa
Araceae
"coral"/ (48U15)
Melastomataceae
Triolena obliqua
"coral"
/ (48807)
Piperaceae
"equis" (48821)
/
family indet.
I
Cayapa
Feet and legs rubbed to produce smell to protect against
snakes ("equis") / (40317)
2
Ca,vapa
.')
Cayapa
Legs rubbed to protect against snakes,/(40893)
"equis"
Feets rubbed to prevent bites of
snakes/(48184)
q
Table 51. Wounds. Plants listed in this category are used to stop the bleeding of cuts, to promote healing and, to fight infections. Infected wounds are a major health problem. The Coaiqueres use the latex of Vismin (Sangre de gallina) as liquid
bandages. It supposedly seals the wound fiom inf'ections and speeds recovery. \44ren no other use is indicated, the plants
are ground into a green paste (Fig. 15). This paste is used as a compress on the bleedingwound and if necessarywith a
piece of cloth wrapped around it to keep it in place. Under commentsthe type of wound which is treated is mentioned as
well as the kind of effect that the t.reatment is expected to have.
Tribe
Part of plant/ preparation/ treatment/ comments/
(AAU v<-rucher)
Vascular crlT)togams
Danaea humilis
Cayapa
Polypodium percussunr
Coloradcr
Leaves/ground/ / / (48062)
Leaves/ ground/ / cuts/ (40629)
P. piloselloides
Cayapa
P. piloselloides
Coloraclo
Selaginella sp.
Trichomanes nrembranacerlrrr
Cayapa
T. punctatunl
Cayapa
T. sp.l
Cayapa
T. sp. 2
Vittaria sp.
Cavapa
Leaves,/sroun d/ humid compress/ dee p woun ds/ (40 432,
4tl892, 48893)
Leaves/ground/humid
compress// (aBB38)
Cayapa
Le aves/ sround/ com press/ promote healing / (40842)
Oayapa
Leaves/dried and pulverised/ /make wounds heal faster,
does not prevent inflammation/ (40475)
Leaves/chewed/compress / / (40236)
Leaves/ ground/ / cuts/ (40505)
Le aves/ sroun d/ cornpre ss/ prom o te healin g / (40822)
Leaves/ground/compress/said to absorb the infection/
(48345)
BS 46
104
Tabel5l
- Continued
Tribe
Part of plant,/ preparation / treatment/comments/
(AAU voucher)
Cayapa
Leaves/dried and pulverized /appliecl/
(40420)
Ca,vapa
Leaves/ground/compress/said
night/ (48359)
Coaiquer
Stem/latex/applied
(48e33)
Cayapa
Leaves/warmed/applied/lower
Cayapa
Leaves,/warmed by body heat (placed near heart),/applied
directly/for inflammation and pains/ (40899)
Mikania sp.
Coaiquer
Cayapa
Wulffia baccat.a
Cayapa
Leaves/boiled,zapplie d / for infections/ ( 4 I 505 )
Leave s7iground/ compre ss/ for i nfe c tions/ ( 4 103 6 )
Lea,-es/ /mixed with 2 other plants/for inf'ections of deep
wounds/ (48054)
genus indet.
genus inclet.
Colorado
Amaranthaceae
Achyranthes aspera
healing of wounds/
Apiaceae
Eryngium foeticlum
to absorb the pus over
Apocynaceae
Tabernaemontana macrocalp<
while fiesh/wounds
caused bv burns,/
Araceae
Anthurium
scandens
swelling/ (40601 )
Araliaceae
Dendropanax arboreus
Asteraceae
Erato polymnioides
Leaves/ groun d/ compress,/ promote s healin g / (40 | 0 4)
Leaves,/ground/wet compress,/ / (401 52)
Colorado
(lolorado
Leaves/ground/compress/promotes
Cayapzr
Leaves/ground/wet
Colorado
h e a l i n g / ( 4 0 1 6 1)
compress/against swelling and inflammation of cuts on the feet/ (40553)
Leaves/boilecl,zapplie d/ for cleanins wounds/ (40662)
Colorado
Leaves/boiled/colcl compress/inflammation/
Colorado
Leaves/ground/paste applied as plaster/for badly inflamed
wounds/ (40057,40166)
Clusia dixonii
Coaiquer
C. sp. I
Coaiquer
C. sp. 2
Coaiquer
C.sp.3
Coaiquer
Twigs/latex,zapplied while fresh/prornote healing / (48790)
Trvigs/latex/ applied while fresh/ / (41456)
Twigs/latex/applied while fiesh/promote healing / (487 Bl)
Twigs/Iatex/applied while fresh/promote healin g,/ (48799)
Vismia obtusa
Coaiquer
Fruit peel/latex/applied
(414e4)
while fresh/infected wounds/
Cayapa
Entire plant/burned/ash
(48068)
appliecl/prevents infections/
Colorado
Leaves/ / / healing of small wounds,/ (40005)
genus indet.
genus indet.
Fienus indet.
Begoniaceae
Besonia glabra
(40696)
Capparidaceae
Podandrogyne brevipeclunculata
Clusiaceae
Commelinaceae
Geogenanthus rhizanthus
Convoh'ulaceae
Ipomoea sp.
BS46
Tabel5l
105
- Continued
Tribe
Part of plant/preparation/
(AAU voucher)
Colorado
Leaves/ ground/ c ompre ss,/for bruise s,/ ( 4005 2 )
Colorado
Leaves,/sap,zappliedwhile fresh/promotes
Acalypha diversifolia
Colorado
A. diversifolia
Cayapa
Leaves/gro wd / / mainly cuts/ (4001 1, 40155)
Leaves/eround//cuts / (40346, 407 35, 40875)
Leaves/ ground/,/ cuts/ (40424, 40466, 40838 )
Leaves/ / / (40055)
treatment/ commen tsl
Cucurbitaceae
Gurania eggersii
Cycadaceae
Zamia lindenii
healin g / (40202)
Euphorbiaceae
A. villosa
A. sp. I
A. sp. 2
A.sp.3
A.sp.4
A.sp.5
A.sp.6
Cavapa
Colorado
(lolorado
Cayapa
Coaiquer
Cayapa
Cayapa
Leaves/ ch ewed/ compre ss/ serious bleedin g/ (40228)
Leaves/sround/ compress/ prevents infections/deep
rvounds/ (41030)
l,eaves/ grounci,z/ deep wounds/ (41 17 3)
Leaves/qround/comp ress/ / (481 82)
Leaves,/ground/compress/prevents inf'ections of deep
wounds/ (48360)
Fabaceae
genus indet.
Cayapa
Leaves,/ground, mixed with Wuffia hnccata/compress/infections of deep wounds,/ (48058)
Columnea kienastiarra
Cavapa
C. laevis
(1. minor
Coaiquer
Leaves,/grouncl/ compress/ removes pains and prevents inflammation / (40458)
Leaves/ ero un d/ cold cornpre ss/ for burn s/ (487 g 4)
Leaves/dried and grotrnd/ / for infections/ (41428)
Gesneriaceae
Coaiquer
Heliconiaceae
Heliconia curtispatha
Cayapa
Young leaves/ritual healing//wound
covered with a leaf
and through this the shaman uses his mouth to suck trp the
inf'ection/ (10434\
Meliaceae
Carapa guianensis
Colorado
Fruit/inrrer
(10703\
(loloradcr
Branches, tmnk/latex/applied
ings/ (40664)
Peperomia sp.
Piper phytolaccaefolium
Coaiquer
Cayapa
Leaves/ ground/compress/burn
P. cararense
Cayapa
Entire plant,/dried and pulverized or burned/powder or
ash applied clirectly,/promore healine of smallwounds/
( 4 1 0 2 1, 4 8 t 7 4 , 4 8 2 4 4 )
P.rp.
Cayapa
Pothomorphe peltata
Colorado
Entire plant/burned,/ash appliecl directlv/ / (1b370)
Leat,es/ /healing/ / (40004)
parts grated and boilecl/applied to wound//
Moraceae
Brosimum utile
directly/infections
or swell-
Piperaceae
Leaves/ground/humid
c.ha")/ (4046U)
s/ (487 93)
compress/cleep fissures ( "chan-
BS 46
Tabel5l
- Continued
/ treatment,/ comments,/
Tribe
Part of plant/preparation
(AAU voucher)
Coffea arabica
Colorado
Fruit/crushed/compress/for
Scrophulariaceae
Scoparia dulcis
Colorado
Root/ / / / (40028)
Solanaceae
Cuatresia riparia
Colorado
Leaves/ ground/warm
(40655, 40687)
Colorado
Stem/ sap extracted by pressure/ applied directly/ inf-ections/ (40i91)
Cayapa
Leaves/ /leaves wrapped around/open wounds atjoints in
connection with open fractures/ (40437)
Cayapa
Leaves/dried, crushed/applied
wounds/ (41077\
Rubiaceae
Cyphomandra
hartwegii
Solanum dolichorrhachis
cuts/ (not vouchered)
compress/ superfi cial wounds/
Verbenaceae
Lantana sp.
Table 52. Swellings from bruises. Unless otherwise mentioned,
directly/bruises and small
only the leaves are used for this purpose.
Tribe
Preparation/ treatment/ (AAU voucher)
Anthuriurn scandens
Cayapa
A. sp. I
Coaiquer
A. sp. 2
Cayapa
Boiled/compress that are changed every 6 hour/ (41040)
/applied while fresh, mixed with chicken fat/ (41458)
Boiled/used as a cold plaster/ (48075)
Araceae
Arecaceae
Cayapa
Stem ground and heated/appliecl to relieve the pain,/
(48351)
Clibadium cuneata
Sciadocephala schultze-rhonhofi ae
Coaiquer
genus indet. A
genus indet. B
genus inclet. C
Cayapa
Cavapa
Heated in water,/wrapped around swelled tisstre/(48698)
/applied while fresh like a plaster/(41481)
,zapplied warm / (40396)
Warmed in water/applied after cooling/ (40863)
Cayapa
/plaster, changed often, said to dry up the swelling/ (48152)
Begonia glabra
Coaiquer
Heated/applied while still hot/ (48818)
Bromeliaceae
genus indet.
Colorado
Stem sliced/applied to sole of feet/ (40197)
Clusiaceae
Clusia sp.
Cayapa
Boiled/wrapped
Cayapa
Entire plant ground and boiled/bocly mbbed to rernove
swelling/ (40373)
Geonoma cuneata
Asteraceae
Coaiquer
Begoniaceae
around swelling/ (48242)
Costaceae
Costus pulverulentus
BS 46
r07
Tabel S2 - Continued
Tribe
Preparation/ trearmen t/ (AAU voucher)
genus indet.
Cayapa
Grouncl with citrus leaves/ compress, said to absorb swelling/ (48350)
Gesneriaceae
Columnea eburnea
C. gigantifolia
Coaiquer
Coaiquer
Heated/wrapped
Gloxinia dodsonii
Cayapa
Fabaceae
around swelling/ (487 42)
/affected area rubbed with fresh leaves/ (41b87)
Boiled/whole body massaged/ (40367 )
Loranthaceae
Phthirusa pyifolia
Phoradendron chrysocladon
Cayapa
Cayapa
Boiled/wrapped around swelling/ (4024b, 48151)
Mixed with warm water/ / (40350)
Melastomataceae
Leandra granatensis
Cayapa
Ground, mixed with tepid water/bath, for blows of children/ (40326)
Colorado
Stenrs grotind/hot
Coaiquer
Dried/applied
Brugmansia versicolor
Cayapa
Warmed/wrapped
Jaltomata procumbens
Colorado
Orchidaceae
Encyclia fragrans
compress / (40668)
Piperaceae
Peperomia sp.
to swelling/ (41b93)
Solanaceae
around swelling,/ (40581 )
Ground/compress,/ ( 400 I 3 )
r0B
BS 46
Table 53. Abscesses and tumors. The informants did not make any distinction between the difl'erent conditions that can
produce swelling in layers of the skin. Most plants are used for tumors, abscessesand postules, a few are used for acne. The
leaves are used, where nothins else is indicated. A compress consistins of layers of ground leaves is nornially wrapped
around the swelled tissrres.
Tribe
Preparation/ treatment,/ comments,z (AAU voucher)
Cayapa
Ground,/ compress/fbr swelling/ ( 48384)
Sciadocephala sp.
Cayapa
Ground in cold water/compress replaced every 4 to 5
hours/ / (41024)
Capparidaceae
Podan dro gy-ne brevipe dun culata
Colorado
Ground/warm
Oayapa
Dried and crushed/ /for swelling/ (41075)
swelling/ (48382 )
Ground/compress/for
Acanthaceae
.|usticia ianthina
Asteraceae
compress/ / (40209)
Fabaceae
Desmodium sp.
genus indet.
Cayapa
Heliconiaceae
Heliconia aemygdiana
Colorado
Fruits crushed/paste appliecl to aff'ected area with a leaf
wrapped aronnd/ / (40120)
Malvaceae
P a v o n i ac a s t a n e i f t r l i a
Cayapa
Grorrnd,/compress/for
Colorado
Bracts used/ / / (401l7)
Oxalis sp.
Cayapa
Grotrnd,/compress/for
Solanaceae
Brugmansia versicolor
Witheringia solanacea
Cayapa
Boiled/used as a plaster/ / (41050)
Boiled,/leaves used to wash hands, decoction used for bath/
(40376)
acne/ (48204)
Marantaceae
Calathea marantifolia
Oxalidaceae
Oavapa
acne/ ( 48333 )
BS46
109
Table 54. Fungal infections. The Spanish term fbr depigmentation
of the skin is mnnchn.sbktnt:a,s.Plants used for such svmptoms are generallv sround and applied to the aflected area as a compress (Fig. 15).
(AAU voucher)
Tribe
Part of plant/preparation/treatment/
Colorado
Bulb/burned/ash
(40214)
Cayapa
Leaves/ grouncl/ compress/ ( 40594)
Cordia spinescens
Cayapa
Leaves,/crushed to a paste and boiled /hot compress applied to affected area 3 times clally/ (40495, 40390, 41062,
48052)
Commelinaceae
Dichorisandra angustilblia
(lolorado
Leaves/ground and boiled/compress/
Cloloraclo
/ ground/ compress,/ (40100)
Colorado
Latex/ / especially for fungal infections affectins the f'eet/
(40177)
Cayapa
Leaves/warmed/humid
Hyptis capitata
Cayapa
Leaves/ground/for
Marantaceae
Calathea rnetallica
Cayapa
Leaves/boiled or ground, eventually mixed with lime and
s a l t / c o r n p r e s s o r o i n t m e n t , h o t o r c o l c l / ( 4 0 3 2 1, 4 0 3 4 9 ,
48117,48245)
Piper sp.
Cayapa
Leaves/dried ancl burned/ash
area/ (40782)
Rubiaceae
Hamelia sp.
Cayapa
Leaves/ground in hot water/applied while still hot to inf'ections between toes/ (41051)
Coloradcr
/ grouncl / appliecl to the skin/ (40 I 0 I )
Oayapa
Leaves/sround to a paste/heated and applied to the skin/
(4830e)
Cayapa
Leaves,/boiled/rubbins
tions/ (40356)
Amaryllidaceae
Urceolina srandiflora
applied to affected area of the skin/
Apocynaceae
Plumeria rubra
Boraginaceae
(40698)
Cucurbitaceae
Cayaponia glancltrlosa
Euphorbiaceae
E u p h o r b i a c o t i n i [ <r li a
Fabaceae
Cassiareticulata
compress/ (40568)
Lamiaceae
nrbbing of the skin/ (40389)
Piperaceae
applied to aft'ected
Solanaceae
Jaltomata procumbens
Ulmaceae
Trema integerrima
Verbenaceae
Cornutia sp.
betlveen toes and fingers {br infec-
BS 46
110
Tabel. 55. Skin reactions. Allergies, eczema, and dandruff. The Coaiquers use a number of plants for treating eczemas that
allegedly are caused by the plant itself. They do not necessarily believe that the patients have been in physical contact with
the plant itself. The plant may house a powerful spirit that can cause the disorders from a distance. The malevolent spirits
are driven out using the plants in which they are believed to dwell. Typically, the leaves are burned and the ash is applied
to the affected area of the skin. The Spanish term for this treatment is Bafin de seco.
Tribe
Preparation,/treatment/symptoms/
Coaiquer
Boiled/applied
Cayapa
Dried, pulverized/applied
(AAU voucher)
Acanthaceae
Dicliptera sp.
to skin/ / (48770)
Amaranthaceae
Achyranthes aspera
to skin/ / (40420,40499)
Araceae
to skin/ / (41503)
to skin/rash/ (48717)
Coaiquer
Dried/applied
A. sp.2
Coaiquer
Monstera lechiriana
Coaiquer
Leaves, dried/applied
/ / / (41633)
genus indet.
Coaiquer
/ / / (48711)
genus indet.
Colorado
genus indet.
Colorado
Ground,/humid compress/break-out caused by lice or fun"rasquinia" (40152)
gi, a symptom called
/
(40
I 04)
Ground / / itching/
Begoniaceae
Begonia glabra
Colorado
Boiled/applied
B. semiovata
Coaiquer
Boiled/decoction
B.rp.
Coaiquer
Boiled/applied
Tovomita sp.
Coaiquer
Boiled/decoction
Commelinaceae
Aneilema umbrosum
Geogenan thus rhizanthus
Cayapa
Cayapa
Ground/sap applied to skin/ / (40296)
Leaves ground/face and arms rubbed/for
"svrna" (40319)
called
/
Costaceae
Costus laevis
Cayapa
Sap extracted/cold
Cucurbitaceae
Melothria pendula
Colorado
Ground/ / allergic reactions,/ (40098)
Acalypha villosa
Cayapa
genus indet.
Coaiquer
Ground/compress / / (40838)
Sap extracted/applied to skin/ / (41491)
Desmodium adscendens
Coaiquer
Dried/applied
Gesneriaceae
Alloplectus sprucei
Cayapa
A. sprucei
Coaiquer
to skin/ / (40342)
Ground/applied
Burned,/"bafro de seco",/for diseasescaused by the same
species/(48795)
A. teuscheri
Coaiquer
Burned/"baiio de seco",/for conditions caused by the same
species/(48961)
Besleria barbata
Coaiquer
Burned,/"bafro de seco"/for conditions caused by the same
species/(48802)
Anthurium
sp. I
Asteraceae
to skin after cooling/ / (40696)
applied to skin/ / (41479,48704)
to skin/ / (48783)
Clusiaceae
used for bath/ / (41453)
type of eczema
bath/ / (41002)
Euphorbiaceae
Fabaceae
to skin/ / (41578)
BS 46
Tabel55 - Continued
Tribe
Preparation/ treatment/symptoms/
Columnea byrsina
Coaiquer
Burned/"baiio
de seco",/for conditions caused by the same
species/(48731)
C. fimbricalp<
Coaiquer
Burned/"baiio de seco"/fbr conditions caused by the same
species/(48971)
C. lehmannii
Coaiquer
Burned/"bafro de seco"/for conditions cirused by the same
species/(48983)
C. minor
Coaiquer
C. minutiflora
Coaiquer
/ /plant irritant/ (48740)
/ / plant ir ritant / ( 48970)
Ground, sap extracted/hair wash/ eczema/ (48741)
/rtrbbecl with warm leaves/ / (40359)
Ground in water,/cure dandruff / / (48741)
C. parviflora
Coaiquer
C. rubriacuta
Cayapa
(AAU voucher)
C. rubricalp<
Coaiquer
C. fimbricalp<
Coaiquer
Burned/"bafro de seco"/for conditions caused by the same
species/ (411923)
C. sp.1
Coaiquer
Burned/"baiio
Cremosperma congmens
Coaiquer
Burnecl/"bafro de seco"/for conditions caused by the same
species/(48979)
C. humidum
Coaiquer
C. muscicola
Ooaiquer
Burned/"baiio de seco"/for conditions caused by the same
species/ (48977)
Burned/"bafro de seco"/for conditions caused by the same
species/ (48978)
Drymon ia warszewicziana
Coaiquer
Burned/"baio
Gasteran thus oncogastrus
Gloxinia dodsonii
Cayapa
Coaiquer
/fresh leaves used for rubbing of skin/ / (41000)
Burnecl/"baflo de seco",/for conditions causecl by the same
species,/(48729)
Nautilocalr.x sp. I
Coaiquer
Burned,/"bafro de seco"/for conditions caused by the same
species/(48936)
N. sp. 2
Coaiquer
Burned/"bafro de seco"/for conditions causecl by the same
species/(48964)
de seco",/fbr conditions caused by the same
species/(48733)
de seco"/for conditions caused by the sarne
species/(48728)
Lamiaceae
Hyptis capitata
Cayapa
Cround,/rubbing
of skin/for symptom called ,,nigua" (an
eczema) / (40389)
Melastomataceae
Aciotis caulialata
Clidemia serpens
Ooaiquer
Cayapa
Boilecl/applied to affected area of the skin// (41479)
Cround/for rubbing of skin/ / (40371)
Meliaceae
Carapa guianensis
Colorado
Inner parts of fruit grated and boiled/applied
fectecl area of the skin/ / (40703)
Piperaceae
Peperomia urocarpa
P.rp.
Cayapa
Cayapa
Piper cararense
Cayapa
Grotrnd and mixed with lukewarm warer / / / (40201:.
Ground/for rubbing of skin/ / (40313)
Dried and ground to powder/applied to affected area of
the skin/,/(41021)
P'.p.
Cayapa
Cround, boiled/decoction
ro the af_
drunk 3 times a day/ / (402gb)
BS 46
112
Tabel55 - Continued
Tribe
Preparation/ treatment/ symptoms/ (AAU voucher)
Coaiquer
Burned/smoke
Rubiaceae
Sabicea villosa
blown on skin// (41506)
Solanaceae
Ground,/compress / / (40624)
to affected area of the skin/for itching ecze-
Brugmansia versicolor
Cestrum sp.
Colorado
Colorado
Boiled/applied
ma/ (40164)
Cuatresia riparia
Colorado
Jaltornata procumbens
L y c i a n t h e sa m a t i t l a n e n s i s
Colorado
Witheringia solanacea
Colorado
allerg / (40099, 40687)
Ground/compress/for
Ground/ compress,/for allergy / ( 40 10 1 )
Ground,/ compre ss,/for itchin g e czema / (401 7 2)
Ground/compress,/for itching eczema and allergies/
W. solanacea
Cayapa
w.rp.
Cayapa
Colorado
(40009,
401r0)
Ground, mixed with cold water/1 bath a day/ / (41053,
4 8 11 5 )
Ground and soaked in water/bath, immediately after wash
of body with soap/ / (48349)
IJrticaceae
Pouzolzia sp.
Verbenaceae
Aegiphila sp.
Lantana sp.
Cayapa
(]round and warmed/compress/
Coaiquer
Cayapa
B o i l e d / d e c o c t i o n u s e d f o r b a t h , / s e v e r ee c z . e m a /( 4 1 5 1 l )
Dried, crushed/applied to affected area of the skin,//
(41077\
/ $0a77)
Table 56. Rheumatism. Rheumatic pains are generally treated by macerating plants to a paste which is spread around the
affected joints as a plaster. In all the caseslisted below, it is the leaves that are used.
Tribe
Preparation/ treatment/ (AAU voucher)
Vascular crn)togams
Selaginella sp. I
Colorado
S .s p . 2
Colorado
Boiled/decoction
Colorado
/applied while warm to treatjoints/
Colorado
/ / (40046)
Cayapa
Ground/used
Clibadium sp. I
C. sp.2
Ca,vapa
C.sp.3
Cayapa
Heated/applied to skin/ (40347)
Ground in warm water/wrapped around affected joint/
(40807)
Ground with a little bit of water / compress,/ (41025)
C.sp.4
Cayapa
genus indet.
Cayapa
Ground while fresh/compress/ (48209)
Boiled/compress/ (40554)
Colorado
/ / (40045)
Boiled,/decoction used for bath/ (40093)
used for warm bath/ (40186)
Agavaceae
Cordyline terminalis
(40640)
Annonaceae
Rollinia mucosa
Araceae
Anthurium
gracile
for rubbing the skin/(40320)
Asteraceae
Cayapa
Bixaceae
Bixa orellana
BS46
113
Tabel56 - Continued
Tribe
Preparation/ treatment / (AAU voucher)
Columnea rubriacuta
Colorado
Boiled/decoction
Malvaceae
Abelmoschus moschatus
Cayapa
Cround and boiled/compress / (40320,48150)
Encyclia frasrans
Colorado
Erythrodes sp.
Cayapa
Cround, warmed/compress/ (40668)
Ground,/used for rubbing of skin/ (40352)
Gesneriaceae
used for bath and drinking/
(4067b)
Orchidaceae
Piperaceae
Piper angtrstum
P. imperiale
Colorado
Colorado
drunk/ (40090)
/warm leaves wrapped around the legs, especially below the
k n e e s , /( 4 0 1 9 9 )
P. trianae
Cayapa
Pothomorphe peltata
Trianaeopiper sp.
Cayapa
B o i l e d / k n e e s r u b b e d / ( 4 10 2 0 )
Ground,/humid compress/ (40393)
Cayapa
/fresh leaves used for rubbing of.joints/(48104)
Cavapa
Boiled/used for massage/ (405a0)
Cayapa
Cround/compress (only used by African Ecuadorians according to the Cayapas ) / G060a)
Coaiquer
Fresh leaves ground/compress/
Colorado
Ground/warm
40712)
Boiled/decoction
Poaceae
Coix lachrymajobi
Portulacaceae
Portulaca oleracea
Verbenaceae
Lantana
(41 468)
Zingiberaceae
Renealmia oligosperma
leaves wrapped around joints/ (40160,
BS 46
Lt4
Table 57. Injured joints. The leaves are invariably used in this category.
Tribe
Preparation/ treatment/ (AAU voucher)
Cayapa
Heated/wrapped
Cayapa
Ground/compress/
Dichorisandra hexandra
Cayapa
Boiled/wrapped
Gesneriaceae
Columnea longinervosa
Coaiquer
/applied while fresh to injured joint/ (41507)
Oryctanthus occidentalis
Phthirusa pyrifolia
Coaiquer
Ground/juice
Cayapa
Boiled/wrapped
Phoradendron piperoides
Cayapa
Vascular crlptogams
genus indet. (Aspleniaceae)
around joi nt / (48437 )
Araceae
Anthurium
trinerve
( 40808
Commelinaceae
around swelled joint/ (48407)
Loranthaceae
extractecl and dnrnk/ (41510)
around joint,z (48 15 1)
Boiled/wrapped around swelled joint and worn for some
days/ (48367)
Orchidaceae
Sobralia macrophylla
Cayapa
Notylia rimbachii
Cayapa
Boiled/compress on swelling/ (48091 )
/fresh leaves wrapped around-joint, said to lower swelling,/
Pleurothallis picta
Cayapa
Ground/ compress/ (48432)
(48r43)
Table b8. Nervous system. Anaesthetics, epilepsy, cramps, dizziness and mental disorders. This is one of the the more weakly clefined catesories. It includes a number of symptoms that, apparently, are related to the nervous system.
Tribe
Vascular crnrtogams
Selaginella geniculata
Part of plant/preparation,/
(AAU voucher)
treatment,/purpose /
Cayapa
Leaves/ground in water/placed on forehead/convulsions
Selaginella sp.
Colorado
w i t h l o s s o f c o n s c i o u s n e s s( e p i l e p s y ? ) / ( 4 1 ] 1 3 1 . 4 8 2 0 7 )
disorders/ (491 l5)
/ground, water addecl/drunk/mental
Asteraceae
genus indet.
Cayapa
Leaves/ground, water aclded/placed on forehead/to
nervolrs persons/ (48443)
Bromeliaceae
Guzmania spmcei
Coaiquer
Leaves/ / colcl bath,/mental disorders/ (48693)
Lamiaceae
Ocimum sp.
Cayapa
l,eaves/ground/placed
sons/ (48445)
Melastomataceae
Clideniia crenulata
Cayapa
Ossaea laxiventtla
Cayapa
/ /body patted with plant/ dizziness/ (41060)
/ /r:sed in ritual of shaman/cramps and epilepsy/ G8173)
on forehead/to
calm nervous per-
Moraceae
Maclura tinctoria
Clarisia racemosa
Scrophulariaceae
genus indet.
Clolorado
L.atex/ / achin g tee th / local an aesth e tic / ( 49 060 )
Latex / / achin g teeth/local anaesthetic / ( 407 I 3 )
Cayapa
Leaves,/boiled / drunk,/ n e rvous con di tion s/ (48414)
Colorado
calm
BS 46
ll5
Tabel58 - Continued
Tribe
Part of plant/preparation/
(AAU voucher)
Solanum schlectendalianum
Cayapa
/ /used for rituals of shaman/conlulsions
sciousness (epilepsy?)/ (48233)
Urticaceae
Pilea sp.
Colorado
Roots/boiled/drunk/calming
disorders/ (40185)
Urera baccifera
Colorado
Leaves/ /massage/calming effect on people with mental disorders/ (40669)
treatment/purpose/
Solanaceae
Table 59. Menstruation.
Vascular cryptogams
Thelypteris sp.
with loss of con-
effect on people with mental
Most of the plants in this category are used to prevent excessive bleeding due to menstruation.
Tribe
Part of plan t/ prep aration / ff eatmen t/ purpose /
(AAU voucher)
Colorado
Rhizome/boiled/drunk,/stops
ingl (40138)
Colorado
Leaves/boiled,/drunk/stops
(40663)
Cayapa
Leaves/boiled/l
Cayapa
Leaves/infusion from fresh leaves/3 cups a day (morning,
noon, evening/infections of vagina and lower parts/ (41071)
Colorado
Roots and flowers/boiled/drunk/stops
bleeding/ (40635)
Colorado
Roots////(40031)
Colorado
Roots////(40029)
Colorado
Roots/boiled/drunk/stops
(40650)
C'olorado
Roo tslboil e d / drunk / / ( 40669)
Colorado
Roots,/boiled/ eaten/alleviation of menstrual pains/ (40208)
Cavapa
Flowers/dried, pulverized and, mixed with water/extract
drunk 3 times a day,/stops excessive menstrual bleedine,/
excessive menstrual bleed-
Boraginaceae
Cordia spinescens
excessive menstrual bleedins/
Clusiaceae
Tovomita weddelliarra
cup drunk in morning, said to be bitter and
somewhat intoxicating/said to induce the first menstruation
of young girls, and to reduce bleeding of olderwomen/
(48335)
Cucurbiataceae
genus indet.
Lamiaceae
Hyptis capitata
excessive mensrrual
Llthraceae
Cuphea strigulosa
Malvaceae
Pavonia fruticosa
Marantaceae
Calathea metallica
excessive mensrrual bleeding/
{Jrticaceae
Urera baccifera
Verbenaceae
Verbena litorales
Vitaceae
Cissus sp.
(40500)
BS 46
116
Table 60. Birthgiving. The use of Heteranthera ren{ormis to promote the healing of the umbilical cord is probably an example of the I)octrine of Sisnature. The leaf petiole of this plant somewhat resembles an rrmbilical corcl in the wav that it is attached to the kidney shaped leaf.
Tribe
Part of plant/preparation/treatment/purpose/
(AAU voucher)
Alstromeriaceae
Bomarea sp.
Cayapa
Leaves/ground and boiled/rnixtr"tre drttnk with salt and
lime/stops excessive bleeding trnder birthrrivins / (40331)
Cucurbitaceae
()urania rnesistantha
Cayapa
Leaves/boiled/ drunk every m ornins,/ relieves pain after
birthgiving/ (48308)
Marantaceae
Oalathea sp.
Cayapa
Young leaves//boiled/relieves
(40435)
Cissampelos sp.
Cayapa
Leaves,/ground, water aclded/cold extract drunk 3 timcs
daily/pains and indisposition before birthgiving / (18153)
Pontederiaceae
H c t e r a n t h e r a r e t t i f rr r m i s
Cayapa
Leaves/warmed/applied
H. renifrlrmis
Coaiquer
pain after birthgiving/
Menispermaceae
to umbilical cord for four days after birthgiving/promotes healing/ (403UU, 4U390)
/heated/applied to scar from umbilical corcl/to prevent inl ' c c t i o n sa n d m a l l o r m a t i o n / ( 4 8 7 7 b )
Table 61. Lactation. The plant usages listed in this category are prob:rbly entirely a product of the Doctrine of Sisnature.
Plants with rnilky latex are thus believed to stimulate womens' milk production while nursing.
Tribe
Part of plant,/ preparations/ treatment/ ptrrpose,/
(A\U voucher)
Odontadenia macrantha
Cayapa
Leaves/boiled/drunk/initiates
ving/ (40895)
P r e s t o n i ap o r t o h e l l e n s i s
Cayapa
P. rotunclifcrlia
Cayapa
L,eaves/boiled/breasts rubbccl 3 times a clay/stirnulate rnilk
production / (40831, 41052)
Le aves/ boile d/ breasts por.rncled softly wi th twi gs/ in itiate s
milk secretion after birthgivine / (40786)
Apocynaceae
milk secretion after birthei-
Asclepiaceae
Cayapa
Leaves/boiled, salt aclded/breasts rubbed for one hour/initiates milk secretion after birthsiving/ (48334)
Tribe
Part of plant/ preparation/ rtse/ (A\Lt vottcher)
Convohrrlaceae
Ipomoea sp.
Cayapa
Flowers/crude/placed
Gesneriaceae
Dryrnonia alloplectoiedes
Coloradcr
R o o t s / b o i l e d / d e c o c t i o n c l r u n k / ( 4 0 0 6 1)
Peperonria pellucicla
Cayapa
P.rP.
Cavapa
Leaves/ground, mixed with hot lvater,/nose drop/ (48287)
Leaves/grouncl and heated/nose drop/ (48314)
Fischeria aequatorialis
Table 62. Nosebleed.
ir-rnostrils / (4047 6)
Piperaceae
BS 46
117
Table 63. Pulmonary diseases. The distinction between this cateeory and Coughs and othsr respiratory disorders(Table 64) is
not clear. The present category inclucles plants that are used to treat severe health problems apparently related to infectious conditions of the lunss.
Tribe
Part of plant /preparation/
(AA.U voucher)
Blechum bror,vnei
floloradcr
Razisea sp.
Cayapa
/ bolle d / bath of torso/ pneumoni a / (40783)
Leaves/crushed in c<lld water,/extract drr.rnk 3 times :r
daylsevere lung infections, coughing of blood/(41056)
Ruellia riopalenquensis
Cayapa
treatment/ purpose/
Acanthaceae
Leaves/sround and dissolved in a glass of water/extract
drtrnk twice a clay/ for aching lungs, caughinu of blood/
(40365, 48067)
Table 64. Respiratory
disorders. Under this categorv are listed plants that are used to treat symptoms of the respiratory
svstem especially coughs and chest pains. Note that plants usecl for the curing of common colds and influenza are not included here, but in Table 65.
Vascular crn)togams
Lomariopsis nieropaleata
Tribe
Part of plant,/preparation/treatment/comments/
(AAU voucher)
Cav:rpa
Ce n tral part of rhizome / l>oile d/ drun k/ respiratorl' problems/ (40333)
Apocynaceae
Mandevilla hirsuta
Cayapa
Leaves/boilecl/throat
sofily pounded/fbr
babies that cough
blood,u (40800)
Fabaceae
Colorado
Roots/ / / chest pains/ (40022)
Cavapa
L,eaves/eround to paste, water aclded,/drunk in morning/
coughs, taste verv bitter / (40 410,407 41,47022, 48 I 68)
Cayapa
Roots/sround, ntixed rvith water/dmnk/cough
blood,/ (40362)
Oayapa
Leaves/ground, juice extracted/drunk once a day in the
morning before eating/ fcrr symptoms like those of tubercuIosis/ (48289)
Eugenia sp.
(bloraclo
Fruits/ /breathing
Schrophulariaceae
qenus indet.
Ooaiquer
/ground under water/extract drunk/fbr
Iungs/ (41461)
Witherineia solanacea
Cayapa
f,eaves/sround, mixed with lukewarm water/dmnk
a day/Ior coughs r,vith bloocl / (40308)
Verbenaceae
Aegiphila alba
Oayapa
Leaves/ground/compress on breast/febrifuge applied to
coughing patients/ (40538)
Desmodium adscendens
Icacinaceae
Discophora guianensis
Marantaceae
Calathea metallica
with
Meliaceae
genus inclet.
Myrtaceae
clifficulties / (40044)
aching
Solanaceae
5 times
118
BS 46
Table 65. Colds and influenza. Plants used for conditions with symptoms atypical of cold and influenza are included in Table 64.
Tribe
Part of plant/ prep ar ation / tr eatmen t/ commen t/
(AAU voucher)
Polybotrya caudata
Coaiquer
Cayapa
Stem,/crushed, juice extracted,/drunk/fbr colds/ (41499)
Vascular bundles/boiled/small
cup drunk 3 times a day/for
severe throat infections/ (41058)
Polypodium fraxin ifolium
Cayapa
/boiled/drunk/removes
nose,/ (40798)
Tectaria acutiloba
Cayapa
Leaves/boiled/drunk/
Asteraceae
genus indet.
Colorado
Leaves/chewed/throat
Vascular crnltogams
Lycopodiella trianae
ant) /inflamed
Commelinaceae
Dichorisandra sp.
mucus from throat and
remove mucus/ ( 40825 )
gargled with juice (taste unpleasthroat/ (40 I 52)
colds/ (47484)
Coaiquer
Leaves/ / eaten fresh/for
Costus sp.
Coaiquer
Stem,ujuice extracted,/drunk pure/for
Lamiaceae
genus indet.
Cayapa
Leaves/boiled,/drunk/to
Costaceae
colds/ (41585)
speed up recovery after influenza
or colds/(40861)
Malvaceae
Urena lobata
Colorado
Roots/boiled/decoction
Coaiquer
/burned/smoke
drunk/for
colds/ (40041 )
Monimiaceae
Siparuna sp.
inhaled/relieves congestions/ (48903)
Piperaceae
/ for colds / ( 40079)
while still hot (also used for bath)/
Peperomia sp.
Colorado
Leaves/boiled/drunk
Piper sp.
Colorado
Leaves/boiled/drunk
for colds/ (40670)
Cayapa
Leaves/boiled/drunk /throat infections, mainly used by
African Ecuadoreans/ ( 40599 )
Cayapa
/boiled/ /small cup drunk for colds twice a dayl(60106)
Cayapa
Leaves/herbal infusion/one
colds/ (40548)
Plantaginaceae
Plantago major
Poaceae
Pharus latifolius
Verbenaceae
Lantana sp.
cup drunk burning hot/for
BS 46
Table 66. Eye infections
and vision.
Tribe
Part of plant/preparation/use/
Cayapa
Leaves/ground with a little water/eyedrops
tion/ (40335, 40853)
Costus laevis
Colorado
C. pulverulentus
Colorado
Leaf rachis / juice/applied to infecred, swollen eyes/ (40678)
/ juice/applied to inflammared eyes, also for improving vi-
c..p.
Colorado
Stem/juice extracted by grinding/
a n d s w e l l e d e y e s /( 4 9 1 1 3 )
Cnreraceae
Fimbrisrylis clichotoma
Cayapa
Leaves/cold extract/applied
(48353)
to aching, red, swollen eyes,/
F. miliacea
Cayapa
Leaves/cold extract/applied
(48358)
to aching, red, swollen eyes/
Cayapa
Leave s/ eroun d, mixed wi th luke-w ar m water / eyedrops/
(40314)
Cayapa
Stem/juice squeezed our/eyedrops for aching and inflammated eyes/ (40399, 41015, 48147)
Colorado
Rhizome/ground,
(49116)
(AAU voucher)
Amaryllidaceae
Urceolina grandiflora
for infec-
Costaceae
sion/ (40700,49103)
applied to inflammated
Melastomataceae
Aciotis caulialata
Poaceae
Paspalum conjugatum
Zingiberaceae
Zingiber officinale
Table 67. Ear infections and hearing. Earaches and improvement
purpose because of its ear shaped leaves.
juice extracted /to treat blindness/
of hearing. (rurania spinulosa is probably used for this
Tribe
Part of plant/ preparation/ treatment/purpose/
(AAU voucher)
Coaiquer
Stem/juice/ear
Cayapa
Leaves/ boi led,/ eardrops/ earach e / (487 56, 48 I 70 )
Acanthaceae
genus indet.
drops / deafness/ (41482)
Cucurbitaceae
Curania spinulosa
BS 46
r20
Table 68. Urination.
Plants that are used for various complications related to urination
are included here.
Tribe
Part of plan t/ prep ar ation / tr eatme n [/ purpose /
(AAU voucher)
Colorado
Leaves,/ground in water/dmnk/for
(4e037)
Bomerea edulis
Cayapa
Leaves/boiled/glass drunk 3 times a day before meals/infections of urinary tract/ (41039)
Bignoniaceae
Mansoa hymenaea
Colorado
Stems, leaves,/ground, mixed with water/drunk/urination
pains rvith blood,z (49089)
Caryophyllaceae
Drymaria cordata
Colorad<r
Leaves/ground/drunk,/promotes
pain/ (40779,49096)
Dichorisandra hexandra
Cayapa
Leaves/ boi le d/ drunk/ urin ation pain s,/ (4037 2)
Costaceae
Costus laevis
Colorado
.luice/ /drtrnk/
C a v e n d i s hi a e n g l c r i a t r a
Cavapa
Leaves/ground/boiled/3
Gesneriaceae
Diastema alfine
Colorado
Leaves,/ground/drunk / / (49ll,l)
Coloradcr
Colorado
(a0693)
.|uice/mixed with water,/drunk/bladder infections/
with
Leaves/ground, mixed with water/ drunk/urination
pain/ (49088)
Scaphvglottis graminifolia
Cayapa
Leaves/boiled/4 teaspoons 3 times a day/painftrl urination
( g o n o r r h o e a ?) / ( a 8 3I 8 )
Piperaceae
Piper sp.
Colorado
Leaves,/sround while fresh/drunk
Colorado
Le aves/ sro un d while fie sh / drunk fr esh/ / (49 0 44)
Vascular cr)?togam
Campyloneuron sp.
urination problems/
Alstroemeriaceae
urination and relieves
Commelinaceae
/ (40678)
Ericaceae
Monimiaceae
Siparuna sp. I
S .s p . 2
teaspoons 3 times a day/ / (40537)
Orchidaceae
fresh/ / (49043)
Rubiaceae
P s y c h o t r i aw i l l i a m s i i
121
BS 46
Table 69. Kidney. Note that the informants did not make a clear distinction between the kidney (urinary system) and the
liver (digestive system). Some of the plants listed in this table could also have been referred to Table 40.
(AAU voucher)
Tribe
Part/ preparation/ treatment/purpose/
Vascular cryptogams
Tectaria sp.
Cayapa
Leaves/ground with water,/drunk before morning meal/
(48435)
Costaceae
Costus laevis
Colorado
/ /luice drunk/liver
Monimiaceae
Siparuna sp. J.
Colorado
S .s p . 2 .
Colorado
Leaves/sround, juice extracted and mixed with water/
drrrnk/liver and kiclney conditions / (10216)
.|uice / mixed with water/ drunk/ kidney condition s/ (40693)
and kidney pains/(10678)
Moraceae
Brosimum utile
Colorado
Latex/mixed with water/drunk/pains
neyl (49110)
Colorado
Leaves/ground/drunk
in liver and kid-
Rubiaceae
Psychotriawilliamsii
/liver and kidney diseases/ (49044)
Table 70. General. Systemic I. Miscellaneous pains. Headache conditions are also referred to this usage category.
treatrnent/ type of pains/
Tribe
Part of plant/preparation/
(AAU voucher)
Vascular cryptogam
Polypodium repens
Thelypteris sp. I
Coloradcr
Coaiquer
T. sp.2
Coaiquer
Leave s/ ground, boile d / / mrscellane ous/ (4067 2)
Leaves/boiled/eaten / back pains/ (41 594)
Vascular bundles//eaten crude/back pains,i (487 45)
Acanthaceae
under water/extract drunk/head-
genus indet.
Coaiquer
Entire plant/ground
ache/ (41459)
Amaryllidaceae
genus indet.
Coaiquer
Corm and leafsheaths/ / eaten/miscellaneous/ (41 604)
Capparidaceae
Cleome sp.
Cayapa
l,eaves/boiled/a half, hot cup 3 times a day/miscellaneous,
cultivated/ (40593)
Chenopodiaceae
Chenopodium ambrosioides
Cayapa
/boiled/dmnk/stomach
Cnreraceae
Cyperus luzulae
Colorado
Roots/ / eaten/ headache/ ( 4003r )
Gesneriaceae
Napeanthus robustrts
Colorado
Entire plant/ground
o u s / ( 4 0 6 7 1)
Melastomataceae
Ossaea micrantha
Colorado
Leaves/boiled/ drunk/ loin pains / (40048)
Oayapa
Leaves/ground in cold water/dmnk/heart
and headache/ (40857)
in cold water/drunk/miscellane-
Piperaceae
Peperomia urocarpa
pains/ (41018)
r22
BS 46
Tabel T0 - Continued
Tribe
Part of plant,/preparation/treatment/type
(AAU voucher)
P. sp. I
Cayapa
l,eaves/ground in cold water/drunk,
pains/ (40578)
P .s p . 2
Cayapa
Leaves/ground, mixed with water/drunk/heart
of pains/
2 teaspoons/heart
pains/
(40777)
P .s p . 3
Cayapa
Piper hispidum
P. imperiale
P. mexiae
P. sp. I
P. sp. 2
Coloraclo
Colorado
( 40 109 )
/ boiled / drunk/miscellaneous/
Leaves,/ground, mixed with water frorn bamboo/heaclache/ (40341)
Trianaeopipermexiae
Colorado
Leaves/boiled/drunk,
Poaceae
Lasiacissp.
Coaiquer
Caryops,/boiled/eatenlheadach
Colorado
Roots/boiled/drunk/
miscellaneous/ ( 40040)
Fruits/ / eaten crude/miscellaneous/ (41 460)
Solanaceae
Browalliaamericana
Physalissp.
Witheringia solanacea
Colorado
Colorado
Colorado
Coaiquer
Cayapa
Leaves/ground,
dissolved in water / | glass drunk/for
children's pains/ (4807 2)
/ boiled/ drunk/ miscellan eous/ ( 40222)
/ boiled / drunk/miscellaneous,/ ( 4009 1 )
Leaves,/heated /drunk, warm/ miscellaneous/ (40206)
hotlmiscellaneous,/ (40667)
e / Gl a69)
Leaves,/ground/juice, drunk with water/dizziness and headache/ (40328)
Table 71. General. Systemic II. Febrifuges.
Tribe
Part of p I an t/ p re p ar ation / tr e atlrler,t / (AAU vo u c h e r )
Colorado
/ boiled / drunk/ (40095)
Justicia sp.
Coaiquer
Entire plan t/ grouncl, juice extracted,/drunk/
Asteraceae
genus indet.
Cayapa
Flowers/dampened,/ placed in rectum/ (4055 4)
Colorado
/ /cold drink/(40140)
Struthanthus sp
Colorado
Leaves/boiled/ drunk / (40084)
Piperaceae
Peperomia pteroneura
Colorado
/ /drvnk/
genus indet.
Cayapa
genus indet.
Cayapa
Leaves/ground, water added/ / (40400)
/ /cold drink,z(40685)
Vascular crnrtogams
Polypodium phyllitidis
Acanthaceae
( 4 I 483 )
Gesneriaceae
Diastema affine
Loranthaceae
(40089)
Rubiaceae
Solanaceae
Solanum con [ertiseriatum
S. nudum
Coaiquer
Cayapa
Leaves/ ground/ extract drunk/ ( 48695 )
Leaves/ground in colcl water/1 small cup clrunk 2 tirnes a
day (important not to drink more)/ (41078, 48271)
Cayapa
l,eaves/boiled/l
Verbenaceae
Lantana sp.
warm cup drunk / (40518)
r"
BS46
r23
Table 72. General. Topical I. Compresses, baths, massage. Miscellaneous pains. External treatments for internal pains.
Leavesare used except in a few cases.The normal treatment comprises the bathing of the entire body or the specific part
of the bodywhere the pain is centred. The preparation, soahedinpond, is ritualistic and described in the text (p.25). The
water in which the leaves have been soaked is used to prepare the bath.
Tribe
Part of plant/preparation/purpose/additional
(AAU voucher)
Vascular cryptogarns
Adiantum petiolatum
Cayapa
A.rp.
Coaiquer
Leaves/ /stomach ache/ / (40571)
Leaves/boiled/back pains/ / (48953)
Leaves/ /stomach ache/ / (48320)
Asplenium laetum
Cayapa
Blechnum sp. I
Coaiquer
B.sp.2
Coaiquer
Campyloneurum sp.
Dennstaedtia sp.
Elaphaglossum sp.
Coaiquer
Hemidictlum
Colorado
marginatum
Coaiquer
Coaiquer
Polypodium repens
Pteris sp.
Colorado
Tectaria ucusa
Thelypteris francoana
Oayapa
T. resinifera
Cayapa
Cayapa
T. sp. I
Cayapa
Cayapa
treatments/
Leaves/ ground/back pains / / ( 48940)
Leaves/boiled/back
pains / / ( 48945 )
Leaves/boiled/back
pains / / (487 49)
Leaves/boiled, cooled /sorebody/ / (4BSB9)
Leaves/ground/headache and nosebleed/compress on
forehead/ (48842)
Rhizomes//headache / / (40134)
Leaves/boiled/sore body / / (4067 2)
Leaves/ /stomach ache/ / (48224)
Leaves,/soaked in pond/stomach
ache/ (4BZZZ)
Leaves/with cold water/sromach ache/ / (40570)
Leaves/ /stomach ache/ / (48296)
T. sp. 2
T. sp. 3
Coaiquer
Coaiquer
Leaves/soaked in pond/stomach ache/ / (48411)
Leaves/ground with water/headache/ / (48761)
Leaves/sround with water/headache/ / (4B7GZ)
T. sp. 3
Coaiquer
Leaves/boiled/
Trichipteris bipinnata
genus indet. (Aspleniaceae)
Coaiquer
Cayapa
back pains / / (48952)
Leaves/boiled,zback pains / / (489 43)
Leaves/ /stomach ache/ / (48297\
Coaiquer
Leaves,/ground/heada
Alternanthera porrigens
Coaiquer
Leaves/ground
(41620)
Gomphrena globosus
Cayapa
Leaves/ /chestpains/
Cayapa
Leaves/warmed/
Prestonia rotundifolia
Colorado
Thevetia peruviana
Cayapa
Leaves/boiled, cooled / sore body / / (4067 4)
Leaves/ / chest pains/ / (40545)
Acanthaceae
Aphelandra
attenuata
che / / ( 48898
Amaranthaceae
in water/headaches
caused by colds/ /
/ (40561)
Amaryllidaceae
Crinum amabile
stom ach ache / / ( 40496)
Apocynaceae
Araceae
Anthurium falcatum
A. napaeum
Colorado
Philodendron sp.
Cayapa
Leaves//sore legs/ / (40304)
/ /sore body/ / ft0676)
Leaves/warmed in hot water/sore body/ / (40310)
Cayapa
Leaves/ /stomach ache/ / (601 15)
Pseudelephan topus puratis
Cayapa
Tagetes sp.
Cayapa
Cayapa
Leaves/ boiled/ misce llan eous pains / / (48381 )
Leaves,/ground in cold water,/headache/ / (40488)
Le ave s/ gr oun d in cold water / he adache / / (40 5 63)
Cayapa
Arecaceae
Geonoma linearis
Asteraceae
genus indet.
r24
BS 46
Tabel 72 - Continued
Tribe
Part of plant,/ preparation/ purpose/ additional treatmen ts/
(AAU voucher)
Begonia serniovata
Cayapa
Leaves/ground with hot water/back pains / / (10298)
Bromeliaceae
genus indet.
Colorado
Leaves/ground,/sore soles of the feet/ / (40043)
Cassia reticulata
Colorado
c.tp.
Coaiquer
Leaves/ ground,/ back pains / / ( 40625 )
Leaves/ /sorelegs/ / (41465)
Campanulaceae
Burmeistera sp.
Coaiquer
f,eaves/trseclfresh,zback pains/ / (4 I 463)
Chloranthaceae
Hedyosmum scoterrimum
Coaiquer
Leaves,/used fresh/back pains/ / (41474)
Costus laevis
Cayapa
Interior of stem//sore
Cyperaceae
Eleocharis retroflexa
Cayapa
Leaves/ground/stom
Thibaudia paniculata
Coaiquer
Sphyrospermum cordifolium
Cayapa
Leaves and flowers/ / s<>rebody / / (41175)
Leaves/ground/chest pains / / (40830)
Begoniaceae
Caesalpinaceae
Costaceae
body / / (40312)
ach ache / / (48327 )
Ericaceae
Gesneriaceae
A l l o p l e c tr r s p a n a n t e n s i s
Columnea rubricallx
Coaiquer
C. tenella
Oremosperma congruens
Coaiquer
L.eaves//miscellaneous pains/ / (4ll870)
Leaves,/ground in water/sore body/ / (41466)
L.eaves//stomach ache/ / (41577)
Coaiquer
Cayapa
Leaves/ground in water/headache/ / (41591)
Flowers, leaves/sround,/sore body / / (40327 ,41000)
Colorado
Entire plant/ground
(40671)
Cayapa
Leaves/ /stomach ache/ / (48228)
Oai,apa
Leaves//so re body / / (40305)
Hyptis mutabilis
Coloraclo
H. verticillata
Coaiquer
/ /sorebody/ / (40039)
Leaves/ground while fiesh/stomach
H.rp.
Coaiquer
Entire plant/burned,
Cuphea tetrapetala
Cayapa
Twigs//stomach
C. tetrapetala
Coaiquer
ache/ / (40402, 40552)
Twigs,z/ headache/body paclcled/ ( 48699 )
Cayapa
Leaves/sround/heada
Coaiqtrer
Leaves/fresh/stornach ache/placed on aching part of stomach (48723)
Gasteran thLrsoncogastrus
Napeanthus robustus
Heliconiaceae
Heliconia latispatha
Coaiquer
in cold water/headaches and othersT 7
Icacinaceae
Discophoragrrianensis
Lamiaceae
ash bath/tired,
ache/ / (4i429)
swollen legs/ / (48720)
Lythraceae
Malvaceae
Hibiscus sp.
che / / (40579)
Melastomataceae
Diplarpea paleacea
t25
BS46
Tabel T2 - Continued
additional treatments/
Tribe
Part of plant/ preparation/purpose/
(AAU voucher)
Miconia theaezans
Coaiquer
M.rp.
Coaiquer
Leaves/lightly dried/stomach ache/ / (41495)
Leaves/boiled and cooled/ne ck pains/ / (41500)
Menispermaceae
Cissampelos tropaeolifolia
Colorado
/ /sorebody/ / (40030)
(loloraclo
Latex/ /liver and kidney parns/ / (40664)
Peperomia sp. I
Cayapa
P .s p . 2
Piper eustylum
Coaiquer
P. obliquum
P. sp. I
Clolorado
Le aves/ ground, wi th water/ che st patns / / (407 84)
Leaves/ drie d, alcohol adde d,/ hea dache / / (41 45 4)
/ /sorebody/ / (40237)
/ borled / s<>rebody / / (402 10)
[.eaves/ ground/ heada che / / (487 26)
P .s p . 2
P .s p . 3
Coaiqtrer
Moraceae
Brosimum rttile
Piperaceae
Colorado
Coaiquer
Coaiquer
Leaves/ /so re body / / (48853)
Leaves/ /headache/ / (48BOl)
Rubiaceae
Gonzalagunia cornif olia
Harnelia axillaris
genus indet.
genus indet.
Cayapa
Cayapa
Cayapa
Coaiquer
ache/ / (48227)
ache/ / (48225)
/ /sorebody/ / (40292)
L.eaves//fresh on foreheacl withfat/ / (41497)
/ /stonach
/ /stomach
Solanaceae
Brugmansia versicolor
Physalis sp.
Coaiqr.ter
Solanum lepodotum
Cayapa
S. mammosum
Cayapa
Leaves/ /headache/ / (41618)
Leates/ /headache/ / (4BB0l)
Leaves/ /headache/forehead massaged, body tapped/
(40306, 40536)
Mature fruits//sore body / / (40487)
S. schlechtendahlianurn
Cayapa
Leaves/ /headache/
Zingiberaceae
Renealmia cylindrica
Oayapa
l,eaves,/ground in cold water/for headache, but entire body
bathed/ / (41042)
C)oaiquer
/ (41067)
BS 46
L26
The Colorados use many
Table 23. General. Topical II. Compresses,baths and massage.Regulation of body temperature.
the body temperacalienteraises
de
Bafi'o
allegedly reduces fever, whereas
plants in ritual bath triatments. The Bafr,od,efresco
that are not
imbalances
other
to
treat
used
be
also
can
system
bath
cold
and
ture of patients experiencing cold. This warm
boiling, but the
by
prepared
sometimes
d,efrescoare
Bafi.o
in
used
The
plants
patient.
the
rr.
of
temp.r"t
related to the body
ritual performed. Afew plant spebath is usually cold or lukewarm. The actual effect of the bath treatment depends on the
cited is supposedto raise (+)
treatment
the
whether
indicated
purposeis
cies are used both in cool and warm baths. Under
or lower (+) the temPerature.
Vascular cryptogams
Adiantum macrophyllum
Asplenium serratum
Blechnum volubile
Bolbitis nicotianifolia
Dennstaedtiaarborescens
Diplazium cristatum
D. striatastrum
D. striatum
D. sp. 1
D. sp. 2
Elaphoglossumsp.
Hemidictyrm marginatum
Huperzia linifolia
Lonchitis hirsuta
Lomariopsisjapurensis
L. nigropaleata
L. nigropaleata
Pollpodium crassifolium
Polypodium phyllitidis
P. piloselloides
Saccolomaelegans
Selaginellasp. 1
S. sp. 2
S .s p . 3
Tectaria nicotianifolia
Thelypteris sp. 1
T.sp.2
T.sp.3
Trichomanessp.
Dennstaedtiasp.
Acanthaceae
Aphelandra sp.
Odontonema strictum
Pseuderanthemumlanceolatum
Raziseaspicata
Sancheziaparvifolia
Tribe
Part of plant / pr ePar^tion/purpose and
additional treatment,/ (AAU voucher)
Colorado
/ /+/ (40111)
Leaves//+/ (40717)
Leaves/ground/ +/ (49038)
/ /=/ (40146)
/ /+/ $0085)
Rhizomes/ /+/ (40149)
Leaves//+/(40051)
Leaves/grornd / +/ (49041)
/ /+/ $02rb)
/ /+/ $0238)
Leaves//+/ (40133)
Entire plant/ /+/ (40128)
/boiled, warm foot bath/+/ (40610)
Entire plant/ /+/ (40130)
Leaves//+/ (40163)
/ /+/ G022e)
Leaves//+/ (40223)
/ /+/ (4063e)
/bolred/+/ (40095)
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Cayapa
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
Colorado
/ /+/ G06r7)
/ /+/ G0225)
Shoots//+ / (40200)
Leaves/ground, boiled /+/ G9033)
/ ground, boiled / +/ (49042)
/ /+/ (40144,40686)
Leaves//=/ (40132)
Leaves/ground/ +/ (49030)
Leaves/ground/ +/ (49031)
/ /+/ G0086)
Leaves/ground/ +/ (49036)
s..p.
Coaiquer
/ /:/ (40710)
Leaves//=/ (40023)
/ /+/ (40211)
Entire plant/ground / +/ (40083)
Leaves//+/(40189)
Leaves/bolled/ +/ (41464)
Amaranthaceae
Alternanthera mexicana
Colorado
Leaves//=/ (40653)
Apocynaceae
Allamanda cathartica
Colorado
/ /+/ (4071e)
Colorado
Colorado
Colorado
Colorado
Colorado
BS46
727
Tabel T3 - Continued
Tribe
Part of plant/preparation/purpose
and
adclitional treatment/ (AAU voucher)
Anthurium napaeum
Clolorado
A. ortegeanum
Colorzrclo
A. oveophilum
A. trinerve
Colorado
/ /+/ G0676)
Leaves/ /=/ (40202)
Roctts/ /-/ (40129)
A. sp. I
Colorado
/ /= or +/ (40741,40231)
/botlecl/ = / (40136)
A. sp.2
Colorado
/ /:/ G0r50)
A. sp. 3
Cayapa
Monstera adansonii
Colorado
Leaves/ground, boilecl / + , also massage/ (40737)
l,eaves/grol.:rnd/ = / (49054)
Philodendron tenLle
Coloraclo
/ /:/ G0226)
Colorado
/ /+/ G06r2)
Coloraclo
Leaves/ /-/
Colorado
Colorado
Leaves/ /=/ (40024)
Sap/squeezed out of plant/+/ (40337)
L e a v e s // + / ( 4 0 1 9 0 )
Coloraclo
Lear,'eslsrrnnd/ + / (49059)
(lolorado
L.eaves//+/ (40704)
Colorado
L.eaves/- or +/ (10131, 40187)
Araceae
Colorzrclo
Araliaceae
Nothopanax fruticostrs
Aristolochiaceae
A r i s t o l t - r c h i pa i c h i n g e r r s i s
(40718)
Asteraceae
genus indet.
genlls indet.
genus indet.
Coloracio
Begoniaceae
Beeonia sp.
Bixaceae
Bixa orellana
Bombacaceae
Matisia coloradorurrr
Commelinaceae
Aneilema umbrosum
(lolorado
Leaves/ /=/ (40033)
Geogenanthus rhizanthus
(lokrrado
/ /+/ G0r06)
Cyclanthaceae
Asplundia sp.
Coloradcr
Yotrng leaves/ / =/ (40i8il)
Euphorbiaceae
Acalvpha sp. I
Coloraclo
A. sp.2
Colorado
Codiaeum variegatum
Colorado
/ /+/ G0657)
/ /+/ G0725)
Leaves/ /=/ (40615)
Fabaceae
Crotalaria nitens
Desmodium adscendens
Coloraclo
Coloraclo
Gesneriaceae
Nloplectus sprucei
(loloraclo
Columnea kienastiana
Cavapa
C. spathulata
Ca1'apa
Diastema scabrurn
Drymonia rhodoloma
Colorado
D. turrialvae
Colorado
G a s t e r a nl h u s t ' < l r a l l i n r r s
Colorado
Cloloraclo
Leaves/ / +/ (40727)
Roots//+, also compress on breast/(40032)
Leaves/sap squeezed out/ = / (40706)
I-eaves,/boiled extrac t / + / (1Bl 62)
Leaves/bolled/+, also massase for seneral indisposition in
the morning/(40481)
Leaves/ground / - / (49092)
Leaves/ /-/ (40716)
/ /+/ G0224)
+/ (40665)
128
BS 46
Tabel T3 - Continued
Tribe
Part of plant/preparation/purpose
and
additional treatrnent,/ (AAU voucher)
Monopyle sodiroana
Cayapa
Napeanthus robustus
(lolorado
Leaves/ /to feverish malaria patients, the leaves are alscr
warmed in water and wrapped around feet/ (40316)
Leaves/eround/lower stomach temp./ (49108)
Coloraclo
Y o u n g l e a v e s // = / ( 4 0 1 1 9 )
Hydrocotyle leucocephala
Colorado
Entire plant/ /+/ (40193)
Lamiaceae
Hyptis obtasiflora
Colorado
L e a v e s // + / ( 4 0 1 8 0 )
Colorado
Leaves/ /+/ (40715)
Phthirusa pyrifolia
Coloradcr
Leaves/boiled/increase
Oryctan thus occiclentalis
Colorado
stomach temp./ (40056)
L,eaves,/boilecl/increase stomach temp./ (40058)
Malvaceae
Hibiscus radiatus
Clolorado
/ /+/ (40ffi7)
H. rosa-sinensis
Colorado
/ /+/ G013e)
Malachra ruderalis
Cayapa
Malvaviscus pendulifl orus
Coloraclo
Leaves/boiled / + / (40557, 40860)
L,eaves//=/ (40027)
Calathea metallica
Colorado
Roots/ /=/ (40204)
Melastomataceae
Miconia venulosa
Cayapa
Ossaea laxivenula
Cavapa
O. micrantha
Colorado
Leaves/ground in tepid water / / (40324)
Leaves/grovnd/ /=, also used for compress/ (40364,41013)
Leaves/boiled / + / (40 I 62 )
(loloradcr
/ /+/ $0006)
Colorado
Leaves/ /=/ (40042)
Colorado
Leaves/ /+/ (40643)
Ardisia sp.
Colorado
L.eaves//+/ (10192)
Myrtaceae
genus indet.
Colorado
Leaves/boiled/increase
Colorado
Colorado
Leaves/grotrnd/ = / (49090)
L.eaves//+/ (40025)
Colorado
L . e a v e s/ - /
Colorado
Roots/ /=/ (40178)
Heliconiaceae
Heliconia curtispatha
Hydrocotylaceae
Lauraceae
Persea arnericana
Loranthaceae
Marantaceae
Meliaceae
Trichilia pallida
Mimosaceae
Inga edulis
Monimiaceae
Siparuna laurifolia
Mpsinaceae
stornach temp./ (49094)
Nyctaginaceae
Bougainvillea sp.
Mirabilis jalapa
Olacaceae
Heisteria sp.
(40064)
Onagraceae
l,uclwigia erecta
r29
BS46
Tabel T3 - Continued
Tribe
Part of plant,/preparation/purpose and
additional treatment/ (AAU voucher)
Coaiquer
[,eaves/boiled, cold decoction/ + / GB7 09)
Peperomia pernamboucensis
P. urocarpa
Colorad<r
(lolorado
Leaves,/sround/ + / (49053)
Leaves/ / = / (40067, 401 35)
P. sp. I
Colorado
P. sp. 2
Colorado
/ /+/ $0143)
Leaves/ /=/ (40169)
Piper aueustum
P. augustum
Oavapa
P. hispidum
P. imperiale
Clolorado
Cayapa
P. multiplinervium
Colorado
/ /+/ (a069a)
P. trianae
Coloradcr
Leaves/sround/ + / (49055)
P. sp. I
Cai'apa
Leaves,/boiled/+, also wrapped around arms and leus/
P. sp. 2
P. sp. 3
Cayapa
Cavapa
Leaves//+, also wrapped around arms and legs/(4Bl58)
Leaves/sround, boiled or fresh/=/ (40776)
P. sp. 4
Colorad<r
/ /+/ $0683)
P. sp. 5
Colorad<r
/ /+/ $067e)
P. sp. 6
Colorado
P. sp. 7
Coloradcr
P. sp. 8
Coloradcr
P. sp. 9
Clolorado
Leaves/ /+/ (40711)
Leaves/g;round, boiled /+/ G9056)
/,/increase stomach temp. / (40227 )
l,eaves,/boiled/increase stomach temp./ (40699)
Trianaeopiper killipi i
Cayapa
L,eaves// + , also wrapped around arms and legs/ (40755)
(lolorado
Fresh roots/ /=/ (40182)
Colorado
Leaves/ /=/ (40070)
Cephaelis gentryi
Coaiquer
Chimarrhis sp.
Geophila herbacea
Coloraclo
Leaves/clried befbre water is aclded/: / (41493)
Leaves/ /=/ (10112)
Leaves/ /-/ (10641)
Gonzalagunia sp.
Hamelia a,xillaris
Colorado
(loloradcr
H. axillaris
Cayzrpa
Hoffmannia sp.
P s y c h o t r i ah o f f i m a n s e g g i a r r a
Coloraclcr
(blorado
P. macrophvlla
Coloraclcr
P. stenostachya
Colorado
$0151)
/ /+/ $0153)
L e a v e s // = / ( 4 0 0 6 5 , 4 0 0 7 2 , 4 0 1 1 4 )
Leaves/ /=/ (40066)
P. sp. I
Cloloraclo
(lolorado
Leaves/ /-/ (40068)
L.eaves// =/ (40082)
(lolorado
/ /+/ $0212)
/ /+/ G0232)
/ /+/ G0685)
Orchidaceae
Malaxis sp.
Piperaceae
Colorado
Twigs/ /=, also massase/ G0a52)
/ /increase stomach temp./ (40090)
Leaves/ / + / (40194, 4068 I )
Leaves/bolled/+,
(405e6)
also wrapped around arrns ancl leus/
(4 0 7e0)
Poaceae
Panicum fiondescens
Pontederiaceae
genusindet.
Rubiaceae
P. sp. 2
P.sp.3
P.sp.4
genusindet.
Ccllorado
Colorado
(lolorado
Leaves/ /+/ (40661)
L e z v e s / / - , a l s o m a s s a g e /( 4 0 0 5 4 , 4 0 6 2 6 )
Twigs/ /+/ (40484)
/ /:/
BS 46
130
Tabel T3 - Continued
Tribe
Part of plant/preparation/purpose
and
aclditional treatmen t,/ (AAU votrcher)
Oitrtrs sp. 1
Coloradcr
C. sp.2
Ruta sp.
Colorado
Oayapa
Leaves/ /=/ (40720)
Leaves/ /-/ (40721)
Rutaceae
Leaves,/ground in cold water,/hear,y headaches, also compress on forehead/ (40558)
Sapindaceae
Alloph,vlus sp.
Solanaceae
Bnrnfelsia grirndifl ora
Cestnrm racemosllnl
C. sp.1
Coloraclo
L e a v e s // + / ( 1 0 1 8 1 )
(lolorado
(lolorado
Leaves/crushed, colcl water/ + / $0021)
L,eaves/=/ (40081, 40176)
l.eaves,/fresh in cold water/=/ (48900)
Coaiquer
C. sp.2
Colorado
S o l a nt r m c o n [ e r t i s e r i a tt t m
Colclracio
Leaves//+/(40688)
Leaves/ /=/ (40630)
S. dolichorhachis
Clolorado
Leaves/boiled, warm decoction vsecl/ + / (40154)
S.rp.
Coloraclo
/ /+/ (406s1)
IJrticaceae
Pilea sp.
Colorado
/ /+/ G0008)
Renealmia cylindrica
Cayapa
R. oligosperma
Colorado
Leaves/ground,mixed with water/+/ (40833)
Leaves/boiled/+/ (40160)
Zingiberaceae
Table 74. General. Topical III. Cornpresses, baths, lnassage, smoke, steam. Ritual treament of vauuely defined or non-spe"chutnn"
fbr general symptoms such as indisposition and body soreness
cific symptoms. The Coaiqueres often use the term
not tu-rlike influenza symptoms. It is a condition allegedly caused by malevolent spirits. Accordine to Kempf ( 1982) chutun
has an important socio-political role and comprises all kinds of serious ailments. Healine of chutun is done exclusively bv
the shaman.
Tribe
Part of plant,/ preparation / treatrnen t/ purpose,/
(AA.U voucher)
Vascular crn)togams
Pi tyrourarnma calornelanus
Oayapa
Tectaria incisa
Oayapa
Thelvpteris curta
Cayapa
Frorrds/bo llecl/ bath / / (48221)
Froncls//m assage/ / (40501, 40902)
/ /massage/ / (48129)
Acanthaceae
Justicia pectoralis
Ruellia tubiflora
Amaranthaceae
Achyranthes aspera
Ooaiquer
Cayapa
Twigs,z/humid, massag,e/ / (487 15)
//massage / / (48402)
()oaiquer
Leaves/soaked in water/ bath or, infusion sprayecl on patient fronr the mouth /"c.hutul"/ (41649, 48877)
Cavapa
/ /massage/ / (41054)
//massage/mainlv fever and vomiting/ (48073)
//massage/for most diseasesparticularlir those with headaches and high fever/ (48118, 48249)
Araceae
Anthurium
subcocrulescens
A. falcatum
Cayapa
A. trisectum
Cayapa
Arecaceae
S y n c c a nt h r r s w a r s c e ni c z i a t tu s
Ca-vapa
////
(60117)
1
BS 46
131
Tabel T4 - Continued
Tribe
Part of plant/preparation/tre^tlrrent/
(AAU voucher)
Erechtites valerianifolia
Pollalesta sp.
Coaiquer
Pseudelephantopus sp. I
Pseudelephantopus sp. 2
Coaiquer
"chutun"/ (48866)
Crude leaves/ /bathl
Le aves/ brrrne d/ smoke in hale d/for he adache / ( 41 492)
"chutun " (41609)
Root//bath/
/
"
Root/crud e / bath / chutun"/ (487 I 3)
purpose/
Asteraceae
Coaiquer
Coaiquer
Bixaceae
Bixa orellana
Cayapa
Seeds/extract/sprayed on patient from the mouth//
(40472\
Capparidaceae
Podandrogyne brevipedunculata
Cavapa
/ /massage/ / (40866)
Coaiquer
Leaves/fre sh / bath /
Cayapa
Leaves,/ground/compress on forehead/ / (48149)
Colorado
[.eaves/ boiled/steam
Commelinaceae
Dichorisandra
"
chutun
"/ (
48706)
Crassulaceae
Kalanchoe sp.
Cyclanthaceae
Cyclanthus bipartitus
bath / / (40221)
Cnreraceae
Cvperus odoratus
Coaiquer
Roots/chewed fresh or dried/sprayed on patient from the
"chutun "/ (41624)
mouth/
Gesneriaceae
Columnea fililoba
C. kienastiana
Coaiquer
C. mbriacuta
Coaiquer
i rrm
C r e m o s p e r m a h i r s u t i s sm
Coaiquer
Leaves/fresh or boile d/bath / "chutun"/ (41 641, 48997)
Leaves//sprayed on patient from the mouth or, massage//
(40605)
"
"
Le ave s,/fre sh / bath / ch u tu n / ( 41 432, 47 622)
L e a v e s , / f r e s h / m a s s a g e / " c h u t u n "/ ( 4 1 5 8 1 , 4 l 6 l I )
C. nobile
Coaiquer
Leaves/fresh/ massage,/"chutun" / (47651)
Gasteranthus oncogastrus
Cayapa
Gloxinia dodsonii
Coaiquer
/ /spraved on patient from the mouth,/for fever/ (40734,
48126)
Leaves/fresh,/massage / "chutun " / (47602)
Cayapa
Lamiaceae
Hyptis sp.
Coaiquer
Ocimum campechianum
Cayapa
genlls inclet.
Cayapa
Leaves,/burned,/smoke bath / / (48852 )
Leaves/ground/comp ress/ / (40395 )
Leaves,/fre sh / bath / / ( 48 I 42 )
Coaiqtrer
//massage /"chttun"
Cavapa
Leaves/fre sh / bath / / ( 40860 )
Clidemia purpurea
Cayapa
C. septuplinervia
Cayapa
Conostegia centronioides
Cayapa
Twigs/fresh/massage / / (40896)
Twigs or leaves/ / massage or hot bath/ / (40510)
Leaves or trvigs//sprayed on patient from the mouth or
massase/ / (40528, 40779)
C. dentata
Cavapa
Miconia barbinervis
Cayapa
M. erioclada
Cavapa
Lythraceae
Cuphea tetrapetala
/ (41423)
Malvaceae
Malachra ruderalis
Melastomataceae
/ /massage/ / (48100)
Twigs/ /massase/ / (48431)
Twigs/ /massage/ / (40569)
BS 46
132
Tabel T4 - Continued
Tribe
Part of plant/preparation/treatment/
(AAU voucher)
M. gracilis
Cayapa
M. lugonis
Cayapa
M. nen'osa
Cavapa
M. trinervia
Cayapa
/ /nassage/ / (41033)
Twigs/,/massage / / (40769)
L e a v e s / f r e s h / m a s s a u e/ / ( 4 1 0 14 , 4 8 0 9 9 )
Twigs/ /massage / / (40489)
ptrrpose,/
Mimosaceae
Trvigs/ /massage / / (48288)
Leaves/ground/bath or massase at full moon/also said tcr
prevent aging/(40403)
Pithecellobiurn longifolium
Cavapa
Zygiz longifblia
Cayapa
Myrtaceae
(lalyptranthes sp.
Myrcia sp. I
Cavapa
M. sp.2
Cavapa
Leaves/bo llecl/ bath / / $l 0 ab)
"chutun "/ (
4 I 6 I 4)
Leaves,/bo ilecl/ bath /
Twiss/ / massage/ / (407 43)
Genus A
Cavapa
l,eaves//m assage/ / (4840 I )
P. brachypodon
Cayapa
(layapa
P. dryaclum
Coaiquer
Trvigs/ /massage / / (407 49)
Leaves,/ground with water, glorving stones added/steam
bath/ / (40881)
"chutun " (41
6 | 2)
/
/ borled / bath /
P. hispidurn
Cavapa
Leaves/ /infitsior-r sprayed on patient Iiom motrth or,
bath/ / (40748)
P. imperiale
Colorado
P. rnultiplinerviurn
Coaiquer
P. peltata
Cavapa
P. sp. I
Coaiquer
P. sp. 2
P. sp. 3
Coaiquer
Leaves/heated,/cornpress on lolver part of legs/ / (40199)
",/
"
Le aves/ fre sh / bath / chu tr.rn (11 47 l, 48 78 6 )
Leaves,/sround/ cornpress or massase,/,/( 40393, 48 I 59 )
"ch1
Leaves/fiesh,/ cold batl-r/
tnn" / ( 4 I 590 )
"/
"
( 4 I 65B )
Leaves/fic sh / bath / chutun
"chutun "/ (
4f1697)
Leav es/ / bath/
"chutu
n" / G87 03)
Leay es/ / bathl
"chr.rtun"/ (48873)
Leaves/fresh/bath/
"chutun"/ (48836)
Leaves/fre sh /bath/
Leaves//spraved on patient from rnouth/ / (48163)
Coaiquer
Piperaceae
Piper ansustunl
P .s p . 4
Ooaiquer
Coaique r
P. sp. l-r
Ooaiquer
P. sp. 6
Coaiquer
P. sp. 7
Cayapa
P .s p . 8
Cavapa
P .s p . 9
P. sp. l0
Cayapir
Trianaeopiper
Coaiquer
Leaves//sprayed on patient fronr rnorrth/exorcistic ritual
perfbrmed to expel malevolenl spirits,z/(48164)
[,eaves//sprayecl on patient lionr mouth/ / (48392)
//rnassage / / (40323)
"
"/ (41
425)
/ / bath / chutun
Ca1'apa
,//rrrassage / / (1058b)
Cephaelis sp.
Harnelia sp.
Cayapa
Ooaiquer
//rnassaee / / (41047)
l,eaves, branches/burned/
Palicorrrea conf-erta
Oa,vapa
//nrassaue / / G05I8)
l.eaves/fiesh/cornpress// (4 I 003)
Twiss//rnassage/ / (48I I2)
Ca,vapa
Poaceae
A n r n r l i n c l l al l e r l e r o t t i a t t a
Rubiaceae
P. guianensis
Cayapa
Psychotria brachiata
Cavapa
P. caenrlea
Cal'aPa
P. hofl'manseggiana
Cayapa
P. macrophylla
Cayapa
P. .p.
Cayapa
sm<>kebath / / (,1I 608 )
,/,/rnassage/ / (18172)
Trvigs/,/massage / / (483 I 11)
Trvigs//massage / / (48 1ti5 )
//massage / / (11055)
BS46
133
Tabel T4 - Continued
Tribt:
Part of plant/preparation/treatment/
(A{U voucher)
Scoparia sp.
Genus A
()ozriquer
Genus B
Ooaiqrrer
"chutun "/ (4I431
)
",/ (
[,eaves/fie sh / bath / chutun
4 I 430)
Leaves/fie sh / bath / " chutun "/ (11 124)
purpose/
Scrophulariaceae
()oaiquer
[,eaves/fiesh/bath/
"
Solanaceae
Cestrum racernosurll
Oavapa
c.rP.
Oavapa
Solanum nudum
Cavapa
genus inclet.
Cayapa
Lear,es/ / m assage/ / (48304)
Leaves//sprayed on patient fiom motrth/ / (48114)
Oavapa
Leaves/ grotrncl/massa ge / / (a0538, 48427)
Npinia purpurata
Oavapa
Hedrchirrnl coronorirrnr
Coaiquer
Renealmia sp.
Cavapa
genlls indet.
Ooaiqrrcr
Lear,esl/spraved on patient fiom mouth/ / (40551)
Rhizomes/ ground/ bath/clisease called "{ruamoczl"/ (487 BB)
//nrzrssaue / / (48235)
"chutun "/ (48694)
Leaves/ / cold bath/
//rnassaee / / (40391)
//massage / / (48275)
Verbenaceae
Aeeiphila alba
Zingiberaceae
Table. 75. Details on the identification of'the material collected
Vouchers iclentifiecl to famih'
Vouchers identifred to genus
V o u c h e r si d e r r t i f i e d t o s p e c i e s
Vouchers not iclentifiecl (to farnil,"")
E s t i m r r t c dn o . o l s p c c i c s
1480(in I l3 farnilies)
1377(in 396 genera)
955 (in 576 species)
l9
930
Table 76. Details on vernacrtlar nirming broken down for tl're three ethnic eroups
Number o{' votrchers collect.ccl
Number of uses registereclr
Number of vouchers cll'nanred plants
Number of'','ouchers rvith morc than one name2
Number of narness
Coaiquer
Ca1'apa
Coloradcr
49ti
369
28l
32
235
r072
718
997
l6
819
421
423
t34
63
litl
I
199i
l5l0
t112
ln
r 235
Note that. several llses were inclicaterl for some vouchers. Spcciersfbr lvhich a particular Llsaqepattern has been registered repeatedly rvithin the same cthnic group are onlv counted once r'vhercasiclentical uses of the sante plant b,vntore
than one ethnic group are register-eclfor erachof these separatelv.
Sometimes a Spanish narr)e u,asgir.en besides the local narne.
N ( ) t e t h a l t h c s a r t t t ' l l a n t ( ' i s o r n e l i r r t ersr s e dl o r r r r o r et h a r ro n e s o c c i t ' s
BS 46
I34
down fbr the three ethnic groups'
of number of uses registered' Data broken
T a b . 7 7 . M o s t important farnilies in terms
Gesneriaceae
Vascular crYPtogams
Piperaceae
Arecaceae
Solanaceae
Melastomataceae
Clusiaceae
Araceae
Asteraceae
Rubiaceae
Remainins lamilies
COLORADO
C,AYAPA
COAIQUER
Vascular crYPtogams
55 (r5 %)
Piperaceae
(
r
1
V
o
)
4r
Gesneriaceae
3 5 ( 1 0% )
Araceae
20 (5 %)
20 (5 7r:) Arecaceae
1 6 ( 4 % ) Rubiaceae
1 5 ( 4 % ) Melastomataceae
r o ( 3% )
Solanaceae
r0 (3 7o)
9 (2 7o)
r38 (37 %)
Asteraceae
Euphorbiaceae
Remaining families
66 (9 7o)
5 7( 8% )
4 7( 7% )
a 3( 6 7 o )
34 (5 % )
3 4( 5 % )
2e (4 % )
2e (4 % )
19 (3 7o)
1 5( 2 % )
3ab$B 7o)
Tab. 78. Vascular cryPtosams'
Usage category
General. ToPical. II (t' 73)
Snake bites. ToPical (t' 4tt)
General. ToPical. I (t. 72)
Vegetables(t.25)
Wounds (t.51)
Stomach infections (t' 4l)
Soap, shamPoo (t. l3)
Snake bites. SYstemic (t. 49)
Jewelry (t. 9)
Intestinal Parasites (t' 44)
Anaernia (t.35)
Liver and sPleen (t' 40)
Snake bites. Others (t. 50)
Colds and influenza (t' 65)
General. SYstemic' I (t' 70)
General. ToPical. III (t' 74)
Rheumatisrn (t.56)
Nervous system (t. 58)
Unprocessed as roPe (t' 7)
Plaitins and textiles (t. 10)
Gums and teeth (t. 39)
Injured joints (t. 57)
Menstruation (t. 59)
RespiratorY disorders (t' 64)
Urination (t.68)
Kidney (t.69)
G e n e r a l . S Y s t e m i cI I ( t ' 7 l )
Piperaceae
Solanaceae
Gesneriaceae
Rubiaceae
Asteraceae
Euphorbiaceae
Araceae
Moraceae
Melastomataceae
Remaning families
62 (r57o )
36 (e %)
2 7( 6% )
27 (6 %)
2 2( 5% )
20 (5 %)
1 6( 4 % )
1 2( 3 % )
r o ( 2% )
I (2 7a)
rB2(43%)
423
718
369
I
Vascular cryPtogams
three ethnic eroups'
UsaglePatterns broken down for the
Coaiquer
T4
10
4
Cayapa
30
27
20
l1
10
9
I
B
5
5
4
4
4
4
29
1
6
8
6
B
F7
2
I
2
4
o
J
2
2
Colorado
I
6
ft
5
I
+
A
T
A
T
I
2
I
J
I
2
;
o
J
1
I
;
J
;
1
T
t-
I
I
1
I
I
I
I
I
2
2
I
I
1
I
I
I
I
1
I
I
;
t>
I
169
I
BS 46
135
Tab. 79. Piperaceae. Usage patterns broken down for the three ethnic groups.
Usage category
Snake bites. Topical (t. 48)
(]eneral. Topical. II (t. 73)
General. Topical. III (t. 74)
Ceneral. Systemic. I (t. 70)
General. Topical. | (t.72)
Rheumatism (t.56)
Stomach infections (t. 41)
Wounds (t.5l)
Skinreactions (t. 55)
Perfbmes (t. 14)
Insect bites. Cl.eneral (t. 47)
Game attractant (t. 32)
Coaiquer
Cayapa
l5
l5
6
B
4
I
+
;
I
I
C l . e n e r a lS
. y s t e r n i c .I I ( t . 7 l )
;
I
30
20
l8
10
7
5
4
4
4
q
J
J
2
2
2
I
2
;
g
2
2
1
I
I
Hallucinogens (t. 1B)
Magic plants (t.21)
Fungal inf'ections (t. 54)
Urination (t.68)
J
q
J
Soap, shampoo (t. 13)
Dyes (t. 15)
Liver and spleen (t. 40)
Snake bites. Systernic (t. 49)
Swelling from bruises (t. 52)
I4
I
6
2
2
3
4
c)
Anaemia (t.35)
Snake bites. Others (t. 50)
Nose bleed (t. 62)
Colds and influenza (t. 65)
I
I
J
Colorado
2
I
I
I
I
I
I
I
1
I
I
I
I>
128
I
l
I
1
;
136
BS 46
Tab. 80. (]esneriaceae. Usage patterns broken down for the three ethnic groups.
Usage category
Coaiquer
Cayapa
18
l9
I
27
5
4
2
2
I
2
Snake bites. Topical (t. 4f3)
Skinreactions (t. 55)
Snake bites. Systemic (t. 49)
General. Topical. II (t. 73)
General. Topical. III (t. 74)
General. T<rpical. I (t.72)
Wounds (t.51)
Swellings fiom bruises (t. 52)
Edible fruits and seeds (t. 24)
Colorado
.|
.')
o
I
7
6
1
1
I
J
2
2
I
I
Masic plants (t. 21)
Edible flowers (t. 26
Gangrene (t.37)
I
I
1
I
I
Stomach infections (t. 4l)
Snake bites. Others (t. 50)
I
I
I
I
I
I
Rheumatism (t.56)
Injured joints (t. 57)
Nose bleed (t. 62)
6
J
2
Game attractants (t. 32)
Ritual ornaments (t. 20)
I
46
22
17
9
I
1
I
Urination (t.68)
General. Systemic. I (t. 70)
G e n e r a l . S v s t e m i c .I I . ( t . 7 1 )
;
I
I
1
I
1
II
I
I
129
Tab. 81. Most important Lrsasecatesories. Data broken down for the three ethnic groups.
CAYAPA
CONQUER
Snakebites. Topical (t. 48)
Skinreactions (t. 55)
5ti (16 7o)
Snakebites. lbpical
COLORADO
(t. 48)
32 (9 7c,)
32 (9 ok)
Eclible frtrits and seecls(t. 2,1)
Timber (t.2)
3l (8 %)
(;anle attractants (t. 32)
General. Topical. III (t. 7.1)
3l (8 7o)
(]eneral. Topical. I (t. 72)
Edible frtrits and seeds (t. 24)
24 (7 %,)
Timber (t. 2)
Game attractants (t. 32)
22 (6%,)
12 (3 o/,')
\4buncls (t. 5l)
Ger.reral.Topical. I.(t. 72)
\tbrrncls (t. 5l )
(]eneral. Topical. III (t. 74)
Stornach inf'ections (t. 4l)
Vegetables (t. 25)
9 (2 7o)
Vegetables (t. 211)
Foclder and fish bait (t. 3l)
8 (.2 %,)
Skinreactions (t. 115)
remaining uses
110 (30 %)
369
remaining uses
72 (ro 7a)
58 (8%)
5 3( 7 % )
40 (6 o/o)
34 (5 7a)
32 (4 %)
2e (4 %)
27 (4 To)
24 (3 %)
l8 (3 Vr,)
331 (46 o/o)
718
Gerreral. Topical. II (t. 73)
Srrakebites. Topical. (t. 'ltt)
\\btrnds (t. 5l)
127 (30 %)
21 ( 6 7o)
21 (5 o/(,)
Stomach inf'ections(t.,11)
19 (1 Vc,)
Srrakebites. Systemic (t. 49)
19 (4o/c;)
Eclibfc frtrits and seeds (t. 24)
18 (4 %)
G e r r e r a l .T o p i c a l . I I ( t . 7 3 )
12 (3 %)
Ber,erases (t. 28)
\1 (3 7o)
S k i n r e a c t i o r r s( t . 1 1 5 )
11 (3 7a)
Gerteral. Systernic. I (t. 70)
remaining trses
11 (3 7,t)
150 (36 %)
423
r37
BS 46
active compounds baTable g2. Thirty-rwo plants thar we consider particularly promising in the search for physiologcally
gto
Amerindian
three
of
the
stuclies
ethnobotanical
the
on
sed
Scientific n:une
Use
Scientific nanne
Use
Aca\pha diar.sfolia
wounds
laxative
A. aillo.sa
Begonia gkrbra
Chenopodium ambrosioides
wounds
Jatropha cur(as
Nnut:kopsis sp.
Paspalum conlugn'tum
Perseaamcricann
P hilo rl endron i n equi laterum
Plryllanthu s anisolobus
contraception
rheumatism
fungal skin disease
Piper tricuspe
lice cure
P.tPlt.
eyeinf'ection
Polygonum hydropipnoides
Sa(tium peruuianum
pain killers
fish poison
Maclura tinctona
Clarisia racemlst
Clibadium sp.
ConJia spinesce.ns
Costuspuluerulentus
rash
stomach ache
dental pains
dental pains
Cuatreca,siariparia
C ; r l a n t h u sb i p u r l i t u s
Desmodium uncinatum
rash
Dielfenbachia seguine
D i.sroph ora gt i a n en.si.s
antbites
coughs
Solanum ctnensc
Spihnthes nlba
Drymaria, cordata
I-icusinsipida
urination pains
intestinal parasites
LIrt:eolina grandi flora
Witheringin, solanacea
antbites
aborticide
Schoenobiblussp.
Scoparia dulcis
arrow poison
eye infection
antbites
fish poison
fish poison
hallucinogenic
intestinal parasites
stomach ache
clental pains
eye infections
rash
BS 46
138
XI Spanishabstract
en espafi,ol
SYnoPsis
utilizan' Esto
la mayorfa de las plantas que
comparaal
Se presenta un estudio etnobotlnico
componente
sefrala hacia un importante
y los Coacategolas
do entre los Colorados, los Cayapas
en
azar en los usos, especialmente
de Ecuador' Duun 5% de
iqueres del occidente costero
s6lo
que.
notable
E,s
,iu, g"n"rales.
1987' se
y
1982
entre
campo
de
rante el trabajo
por los tres
las piantas ritiles son compartidas
de los cuadiferencias
colectaron caii 2000 pliegos testiuo'
las
refleja
grupos indigenas, lo que
material identiles se registraron 1510 usos' El
ju composici6n floristica del bosque seE"
y 396 gdneros
frcado pertenece a 113 familias
situacion sociocundario respectivamente' La
representadas
especies
de
numero
EI
es distindistintos.
polftica de los tres grupos estudiados
73 categoias de
distinguen
Se
930'
en
en el uso
se estima
iu, po. lo que se refleja intensamente
para cada cateEsto se
uso. Se citan listados de plantas
vegetales'
que hace" a. los recursos
etnobot6'niq-"'
goria. Se presentan varios modelos
palmas'
las
upti.u particularmente t:"
de mayor
familias
10
las
entre
8
los
ciian
S"
para
Zo..
constituyen un recurso importante
criptogaindigenas:
grupos
tres
los
Colpara
valor
qur los
Cayapas'y los Coaiqueres' mientras
Soladel uso
mas vasculares, Piperuttae' Gesneriaceae'
importancia
La
oradts lo utilizan poco'
Melastomataceae
el aucon
naceae, Araceae, Rubiaceae'
rapidamente
de las palmas se plrdi6
de uso que represen.la
y Asteraceae. La categorias
integraci6n
mento de la aculiuraci6n y su
son las de
registros
de
cantidad
mayor
entan
diferenciacioeconomia de mercado' Se hacen
y las de
determicuracion de mordeduras de serpientes
etnobotanicas
pr6cticas
nes entre dos
corporal' El
y
plantas
banos pararegular la temperatura
las
de
propias
nantes: las propiedades
uniquaColorados
los
por
de las
ultimo es pra;iicada
el intercambi; cultural' Los elementos
m6s empleaetnobomente. Gesneriaceae es la familia
modelos
ceremonias de curaci6n y los
de serpiente'
de un inda para combatir las mordeduras
t6nicos se citan como evidencias
no medicinales'
usos
de
categoria
la
de
Dentro
a un nivel local
tercambio en el pasado cultural
extensa' La
m6s
la
es
comJsdbles
la de frutos
y regional
mas importante
categoria medicinal sistemica
el resto' Varios
tiene un rango de 10 sobre
los Colorados
de
aspectos de lI etnobotanica
Palabras clave
otros grupos
de
la
destaca se se compara con
Ecuador;Colorados(Tsatchela);Cayapas
de los caraetde la costa de Ecuador' La mayoria
(Chachi) ; Coaiqueres (Awa) ; etnobotlnica;
durante
adquiridos
sido
han
cteres aberrantes
denominaci6n
nomedicina; etnofamacologia;
de contactos con
ceremonilos fltimos 50 afros a trav6s
venecula; tratamiento; preparaci6n;
encomparaticulturas andinas' Hay una correspondancia
as de curaci6n; chaman; aspectos
por familisituatre la perdida de usos de las plantas
vos; classificacion de usos; aculturacion;
en los
famlias
de
floristica
as y Ia composici6n
cultural'
ci6n polftica actual; intercambio
se encuentran
alrededores del bosque' donde
BS 46
139
XII List of references
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and Stupefactants in the Plant Kingdom. - Iu
Prance, C. T. and Balick, M.J. (eds.), New Directions
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Econ. Bot. 8:l-23.
Balslev, H. 1988. Distribution patterns of Ecuadorian plant
species. Taxon 37 :567-57 7 .
Balslev,H. & Barfod, A. 1987. Ecuadorian palms - an overview. - Opera Botanica 92:17-35.
Barfod, A. & Balslev, H. 1988. The use of palms by the Cayapas and Coaiqueres on the Coastal Plain of Ecuador. - Principes 32:29-42.
Barrett, S. A. 1925. The Cayapa Indians of Ecuador. - Indians Notes and Monographs. Museum of the
American Indian, Heye Foundation, New York 40:l476.
Berg, C.C. 1972. Olmedieae, Brosirneae (Moraceae). - Fl.
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Cafradas C., L. 1983. El mapa bioclimatico v ecologico del
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- ABYA-YAIA,
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1991. Biological extinction in western Ecuador. - Ann.
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Ph.D. 1985 New School for Social Science.
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etnol6gico
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BS 46
t40
exchange of rituals and meclicinal plant with other
ethnic groups in the region. - -1n:Rios, M & Borgtoft
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291 - U.S. Governrnent Printing OiTice, \Alashington
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Lockwor d, T.E. 1979. The ethnobotanv of Brugnransia. -.f .
E t h n o p a r m a c . | : l 4 7- 7 6 4 .
L,opez, F.F. 1986. Etnologia Ecuatoriana. I Colorados. Instituto Ecuatoriana de Credito Educativo y Becas, Quito, Ecuador, 150 pp.
Martinez,I. P. 1986. Los Cavapas en el siglo X\rl. - Misc.
Antropol. Ecuatorian a 6:23 | -252
Mason, J. A. 1950. The languaees of Sotrth American Ind i a n s . - I n : S t e w a r c l J, . H . ( e d . ) , H a n d b o o k o f S o t r t h
American Inclians 6 - U.S. Government Printing Office, \Alashington D. C.
Medina V., H. 1992. Los Chachi. Supervivencia y ley tradicional. - ABYA-YALA, Quito, Ectrador, 141 pp.
Mitlewski, B. 1985. Pesca Cayapa. - Misc. Antropol. Ecuatoriana 5:63-85.
Murra,.J. 1946. The historic tribes of Ecrtador. - In: Stewarcl, -|. H. (ed.), Handbook of South American Indians 4:785-B2l - U.S. Government Printing Office,
Washington D. (1.
1948. The Cayapa and Colorado. - In: Steward, J. H
(ed.), Handbook of South American Indians 4:277-
Renner, S.S., Balslel', H. & Holm-Nielsen, 1,.B. 1990. Flowering plants of Amazonian Ecuador - A checklist. .{{U
Reports 24, Unir.'ersitl' of Aarhtrs, Denrnark,
24r PP.
Rivet, P. 105. Les Indiens (lolorados, R6cit cle voyase et
etucle ethnologique. -.|. Soc. Am. Paris 2:177-208.
Rivier, L. & & Lindgren, J.A. 1972. Avahuascir, the South
American hallucinogenic clrink. Ethnobotanical and
c h e m i c a l i n v e s t i s a t i o n s .- E c o n . B o t . 2 6 : 1 0 1 - 1 2 9 .
Santesson, C.G. 1936. Pfeil- uncl Fischsift aus Kolumbiean
und Ekuador. - Etn. Studier 2:15-29.
Stout, D.B. 1948.The Cuna, the Choco. - In: Stervard,J.H.
(ed.), Handbook of Sotrth Anreric:rn Indians 4:257276. - LI.S. Governrnent Printing Oflice, Washington
D.C.
Survival International. 1989. no. 24.
Telban, B. 1988. Grupos etnicos de Colombia. Etnoqrafia y
Bibliografia. - ABYA-YALA., Quito, Ecuadoq 525 pp.
Thomsen, K. 1986. Reporte sobre esttrdios cle los arboles y
las palnieras en el t.erritorio de los Awas, provincia
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6:27-47
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Ferns (7. ed.). - Carnbridge University Press, L.onclon.
BS 46
r41
App"ndix I
Index to scientificnames
Scientific names arransed alphabeticallv according to 1), genus name and 2), specific epithet.
The numbers in bolcllace refer to the tables 2-74 that list vouchers for each usage category. In parentheses, after usage category, is cited the voucher specimens from which the information comes. The family to which a senlls belonss is indieated in parentheses after the specimen citations.
Abelmoschus
moschatusMedik. 56 (40336,48150) (Malvaceae).
AcalyphadiversifbliaJacq.5l (40875,40346,40735,40155,
40011);A. villosaJacq.5l (40466,40424,40838);
55
(41491); A. spp. 16 (40648); 28, 73 (40725); 5l
(40228, 40055, 48360, 48192, 4t473, a1030); 73
(40657)(Euphorbiaceae).
AchyranthesasperaL. 74 (48877);39,74 (11649);51, 55
(40420)(Amaranthaceae)
.
Aciotis caulialata(R. & P.)Triana55 (41478);48 (40745);
A. .p. 48 (48316,41016);66 (40314)(Melastomaraceae).
Adenostemma platyphyllum Cass.52 (41481) (Asteraceae).
A d i a n t u m m a c r o p h v l l u m S w .I ( 4 8 3 9 9 ); 7 3 ( 4 0 1 1 1 ) ; A . o b liquum Willd. 48 (48133); e pedolatum Desv. 72
(40571);A. spp. 9 (40828); 48 (40689) (Pteridaceae).
Aechmea magdalena Andre ex Baker l0 (4U386) (Bromeliaceae).
Aegiphila alba Moldenke 73, 74 (40538);74 (48427); A. sp.
2 4 ( 4 8 7 7 9 ) ; A . s p . 5 5 ( 4 1 5 1 1 )( V e r b e n a c e a e ) .
Aiphanes gelatinosa Moore 25 (60003) (Arecaceae).
Alchornea sp. 3l (41631) (Euphorbiace:re).
Allamanda cathartica L. l6 (40504); 16, 73 (40719) (Apoc,vnaceae).
Allophylus exelsus Radlk. 36 (41504); A. sp. 24, 73
( 4 0 18 1 ) ( S a p i n d a c e a e ).
Alloplectus dodsonii Wiehl. 49 (40053); A. panamensis
Morton 48 (48120);72 (18870); A. sprucei (O. Kunt z e ) W i e h l . 4 l ( 4 9 0 5 2 ) ; 4 8 ( 4 1 0 8 1, 4 0 8 4 4 , 4 8 0 8 2 ) ; 5 5
(40342,48795) ;73 (40706); A. tetraeonoides Mansf.
48 (48732, 48813); A. teuscheri (Ra,vm.)Wiehl. 55
( 4 8 9 6 1) ( G e s n e r i a c e a e ).
Alonsoa sp. l6 (40560) (Scrophulariaceae).
Alpinia purpurata (Vieill.)K. Schum. 74 (4055I) (Zingiberaceae).
Alternanthera mexicana (Schlecht.)Hieron. 73 (40653);
A. porriuens (|acq.)Krntze 72 (41620); A. sessilis
(L.)R. Br. 48 (40890) (Arnaranthaceae).
Amaranthus sp. l6 (48391) (Amaranthaceae).
Amphiclasya spp. 2l (40339, 4i059); 26 (482a0;
(41643) (Rubiaceae)
74
Amphilophium paniculatum (L.)H.B.K. l0 (aB7B7)(Bignoniaceae).
Aneilema umbrosum (Vahl)Kunth 55 (40296): 73
(40033)(Commelinaceae
).
Annona muricataL. 24 (48103)(Annonaceae).
Anthurium anoreanum Linclen 16 (41489); A. asplundii
Croat8 (40886); A. bakeriHook. f .48 (41417) ; A. caulorrhizum Sodiro 32 G824U: A. falcatum Sodiro 74
(48073); 72 (40304); A. gracile (Rudge) Schott 32
(48232);56(40320);A.lanceaSodiro25 (410a8);A.
napaeum8ng1.72,73(40676);A. ortegeanumSodiro
73 (40202);A.oveophilumSodiro73 (40129);A.pallicliflomm Engl. 13 (48106);A. scandens(Aubl.) Engl.
52 (41040);5l (40601);48 (40465);A. subcoerulescensEnsl. 74 (47054);A. trinerveMiq. 57 (a0808);73
( 4 0 2 3 1 ) ; 7 3( 4 0 1 a 7 ) ; A .t r i s e c t u mS o d i r o7 4 ( 4 8 1 1 8 ,
48249);A. versicolorSodiro 25 (48098);A. spp. 13
(40477, 48342);20 (40377); 25 (40850, 403b7);3r
( 4 8 7 8 2 ) ; 3 2 ( 4 8 2 3 9 ,4 8 0 9 5 ) ; 5 2 ( 4 1 4 5 8 ,4 u 0 7 5 ) ;5 5
(41503);73 (40136,40150,40737)(Araceae).
Antrophyum intramarginale (Baker exJenman) Kartez 5l
(40842)(Vittariaceae
).
Apeibasp. 2 (48198)(Tiliaceae).
Aphelandra flammea Wassh. 72 (a8B9B); A. sp. 73
(407I 0) (Acanthaceae)
.
Arachnothyrx inconstansStandl. 24 (48768)(Rubiace:re).
Ardisia romeroi Cuatr. 24 (40542, 40906); A. ronseroi
Schuta24 (48325);A. spp. 24 (40817);73 (a0192)
(Myrsinaceae).
AristolochiapichingensisH. Pfeifer21,73 (40718);A.pilosaH.B.K. 24 (40235);A. sp. 4l (40523)(Aristolochiaceae).
Arrhrostema ciliatum R. & P. 25 (48053, 40461)(Melastomataceae).
Arundinella berteroniana (Schult.)Hitchc. & Chase 74
( 4 0 5 8 5()P o a c e a e ) .
Asplenium hallii Hook. & Grev.44 (48744);72 (48297);A.
purpurascensMett. 73 (40149,40215,49035);
A. serratum L. 73 (40717);al (49032);A. triculum Lellinger 48 (48765);A. spp. 48,49 (40697);49 (a0075)
(Aspleniaceae).
I
742
BS 46
A s p l u n d i as p p . 7 ( 4 0 8 2 9 ,1 8 1 2 4 , 4 8 3 7 1 ) ; 3 2( 4 9 1 7 5 ) ;4 8
(4011B); 73 (40lBB)(C1'clanthaceae).
Astrocaryurnstzrndleyanum
BailevL0, 24 (60078)(Arecaceae).
(60113); B. setosa
B a c t r i s g a s i p a e sK t r n t h 4 , 5 , 2 4 , 2 5 , 2 7
M a r t . 2 4 , 2 5 ( 6 0 1 1 0 ) ; B . s e t u l o s al ( a r s t e n 4 , 5 , 2 4 , 2 5 ,
27 (60010) (Arecaceae).
Banrbusir guacltra Kunth 2 G8374) (Poaceae).
B a n i s t e r i o p s i sc a a p i ( S p n r c e & C r i s e b . ) M o r t o n 1 8 ( 4 0 2 9 9 ,
40582); 18, 43 (10722) (Malpighiaceae).
Bauhinia sp. 2 (41674) (Oaesalpiniaceae).
Besonia glabra Aubl. 44 (48819); 51, 55 (a069tr); 52
(4BBI B) ; B. semiovata l.iebrri. 35 (48064) : 37 (41029,
48276); 55 (41479, 18701): 72 (40298); B. spp. 25
( 4 8 4 2 1 ) ; 3 7 ( 4 8 3 1 0 ) ; 5 5 ( 4 8 7 8 3 ) ; 7 3 ( 4 9 0 5 9 )( B e g o niaceac).
Bertiera quianensis Aubl. 4l (40069, 40156) (Rubiaceae).
Besleria barbata (Poepp.)Hanst. 48 (18734); 55 (a8802);
B. barclayi 1.. Skog 48 (4l0tl, 48415); B.solanoides
(liilathea lutea (Aubl.)Schtrltes 25 (48181); (1.r'narantifblia
Standl. 53 (40117); (1.rnetallica Pl:rnchon & l,inden
5 4 ( 4 i 0 6 1 , 4 0 3 2 1 , 4 0 3 4 9 , 4 l J l 1 7 , 4 8 2 4 5 ); 5 9 ( 4 0 6 5 0 ) ;
6 4 ( 4 0 3 6 2 ) ; 7 3 ( 4 0 2 0 4 ) ; C a l a t h e as p p . l 0 ( 4 0 4 6 7 ) ; 6 0
(40435) ; 26 (40433) (Marantaceae) .
Calliandra angustilblia Spnrce ex Benth. 16 (40556) (Mimosaceae).
( l a l l i s i a r e p e n s I - . l 6 ( 4 U : 1 9 4 () C o r n n r e l i n a c e a e ) .
Calyptrantlies sp. 74 (41045) (Nlyrtaceae).
a.nsustifbliurn (Srv.)Fee 50 (a9039); tl.
spp. 41, 68 (49037); 48 (,18fi44);72 ft87a9) (Polvpo-
Campl'loneuron
diaceae).
(lanna inclica x generalis Bailey 16 (,10530); C. sp. 33
(4905I ) (Oann:rceae).
Capparis ecuadoriana Iltis 28 (40097) (Cappariclace:re).
C a p s i c u r n f i u t e s c e n s t . . 2 9 ( 4 0 2 0 1 , 4 0 4 5 b , 4 0 1 1 t r 54, 0 5 6 6 ,
40586, 48988) (Solanaceac).
Carapa srrianensis Aubl. 2, 24 (41667); (1. me€istocarpzr
Krrnth 20 (48735); B. sp. 48 (41427)(flesneriaceae).
B i l l i a c o l o m b i a n a P l . & l . i n c i 2 ( 4 1 t ' , 6 3 )( H i p p o c a s t a n a -
Gentry & Dodson 51, 55 (40703) (Meliai:eae).
C a r i c a p a p a y a L . 2 4 ( 4 0 1 5 9 , 4 0 4 8 1 1 )C
; arica spp. 25 (40088,
40 I 58, 40195) (Oaricaceae).
(l:irh.rdovica palmata R. & P. l0 (40419) (Cyclanthaceae).
ceae).
Bicrphytum clendroides (H.B.K.)D.C. 33 (40577) (Oxalicla-
Oarpotroche pl:rtyptera Pittier 22,24 (40060); 3l (40366,
40873) (Flacr>urtiaceae).
ceae).
Bixa orellana L. 15 (48897); 15, 30, 56 (40045);30,73
(40704); 30 (10472, 10473, 40474,40602); B. plat y c a r p a R . & P . e x ( ) . D o n 3 0 ( 4 0 U 1 0 )( B i x a c e a e ) .
Casearia sp. 24 (40415) (Flacourtiaceae).
Cassia papillora Britt. & Rose 35 (40543, 407!t7); C. retictrlata Willd. 54 (40568); 72 (a0625); C. sp. 72
( 4 1 4 6 5 ) ( C a e s a l p i n i a c e a e ).
Blakea punctulata (Triarra)\Alurdack 2, 32 (al6l3); 32
(4U9u7); B. subconn:rta Bers ex Triana 26 G0297);
3 2 ( 4 U 3 3 6 ); 4 8 ( 4 0 0 4 9 ) ( N l e l a s t o n r a t a c e a e ) .
Castilla elastica Sesse 2, 12 (48214); l2 (48994)(Morac e a e) .
Catoblastus inaequalis (Cook & Doyle)Burret 2 (60002)
Bleclrnrrm r,olubile Kaulf . 7 (40763); l0 (48060); B. spp. 48
(48942);72 (48940), 48945) (Blechnaccae).
Blechum brownei.fuss.63 (40183) (Acanthaceae).
Bolbitis nicotianifolia (Sw.)Chins. 4l (48219); 73 (401a6)
(Dryopteridaceae) .
Bornarea edulis Anclre ex Poir. 14 (40814); 68 (41039); B.
spp. 14 (4U059);60 (40331) (Alstroemeriaceae).
Bonafbtrsia lonsitr.rbtrlosa R. & P. 24 (11632) (Apocvnaceae).
Borreria ocyrnoides (Borm.) D.C . 22 (48226) (Rubi:rceae).
Bougainvillea sp. 73 (49090) (Nvctaginaceae).
Brosimum utile (H.B.K.)Pittier 2, 12 (41037); 40, 69
( 4 9 1 l 0 ) ; 5 1 , 7 2 ( 4 0 6 6 4 )( M o r a c c a c ).
Browallia americana L. 70 (10040) (Solanaceae).
Bnrsmansia
versicolor L:rgerh. 36 (4i1930); 18, 52
( 4 0 ] 1 8 1 ) ; 7 2 ( 4 1 6 1 8 ) ; 1 8 , 5 3 ( 4 1 0 5 0 ); 5 5 ( 4 0 6 2 4 ) ( S o l anaceae).
Brrrnfelsiarrrandiflora D. Don 73 (40021)(Solanaceae).
B u n c h o s i : r c o r n i f i r l i a H . B . K . 2 4 ( 4 8 0 3 1) ( M a l p i g h i a c e a e ) .
Btrrnreistera rulgaris Wimrn. 24, 25 (41031); B. sp. 25
(48070, 10343, 40876); B. sp. 32 (41463, 48102,
48696, 48707 ) (C:rr-npanulaceac) .
(Arecaceae).
C a v e n c l i s h i ac o n p l e c t e n s H e m s l . 3 9 ( 4 1 5 0 2 ) ; C . e n e l e r i a na Hoer. 20 (48925); 68 (40537); O. urandifblia Hoer. 20 (48965) (Ericaceae).
Cayaponia slandulosa (Poepp.
(40 i 00) (Oucurbitaceae )
&
Encll.)Coun.
54
C e c r o p i a s p . l 0 ( 4 U 4 3 3 )( U r t i c a c e a e ) .
O e p h a e l i s u e n t r y i D w r y e r2 ( 4 8 1 0 8 ) ; 7 3 ( 4 1 1 9 3 ) , O . s p p . 3 2
( 4 8 2 0 8 , 4 8 9 0 4 ) ; 74 ( 4 1 0 4 7 ) ( R u b i a c e a e ) .
( l e s t r r r m b a en i t z i i l , i n g e l s 2 , 2 4 ( 4 1 6 5 0 ) ; C . r a c e r n o s t r m R .
& P . 7 3 ( 4 0 0 8 I ) ; 7 4 ( 4 0 3 9 1 ) ;( 1 . s p p . 2 8 ( 4 0 0 9 2 ) ; 3 2
(48166); 47, 55 (a016a); 73 (406u8, 48900); 74
(4827 5) ( Solanace:rer).
Clhamaedore:rpinnatifrons (lu.q.)Oerst. l5 (40198); 31,
32 (48908) (Arecaceae).
Ohenopoditrrn ambrosioitles I-.
(40857) (Ohenopodiaccae) .
4l
(4057ir),
41,
70
C l h i r n a r r h i ss p . 7 3 ( 4 0 1 1 2 ) ( R u b i a c e a e ) .
Chrysochlamys clependens Tr. & Pl. 24 (40409)(Oltrsiaceae).
(lhrvsophyllum arsenteurlr
.facq. 3 (41017); 24 (40289)
(Sltpolaccltt').
BS 46
(ihrysothemis {iieclrichsthalian:r (Hanst.)H. E. Moore 48
( 4 0 3 6 1 , 4 8 0 8 0 ) ( C l e s n e r i : r c e i i e.)
C i s s a n r p e l o st r o p : r e o l i f b l i a D ( 1 . 2 4 ( 4 0 U 5 1 ) ; 3 9 ( . t t a 7 2 ) ; 7 2
(40030);O. sp. 60 (4ttl53) (Menispernr:rceae).
C i s s u ss p . l 0 ( 4 0 4 8 6 ) ; 5 9 ( 4 0 5 0 0 ) ( V i t a c i : a e ) .
C i t r t r s s p p . 2 4 , 7 3 ( 1 0 7 2 1 ) ; 2 8 ( . 1 8 1 9 4:)2 8 , 7 3 ( 4 0 7 2 0 )( R u taceae).
C l a r i s i ar a c e m o s a R . & P . 5 8 ( , 1 0 7 1 3 ) ( M o r a c e a e ) .
C l a v i j a e g g e r s i a n a M c z . 4 8 ( 4 0 7 0 U )( T h c o p h r a s t a c ea c ) .
C , l l e o r n es p . 7 0 ( 4 0 1 1 9 3()C a p p a r i c l a c e a e ) .
C l i b a d i u m s p p . 3 3 ( 4 9 0 9 1 1 ) 5; 2 ( 4 u 6 9 U ) ; 5 6 ( 4 0 3 4 7 , 4 0 8 0 7 ,
4 I 021-r,48209) (Asterircezre)
C l l i c l e n r i ac r e n u l a t a ( i l e a s o n 4 8 ( , 1 8 0 7 7 ) ; 5 8 ( 4 1 0 6 0 ) ; ( 1 . c l i s (Triana)Cogn. 4l (,10307); O. pru'pr.rreaD.
Don 74 (40896); O. scpttrplincrvia Cogn. 74
( 4 0 5 1 0 ) ; O . s c r p c n s ( T r i a n a ) ( l o s n . 5 5 ( , 1 0 3 7 1 () N 1 c l a -
color
slornalaceae).
( l l u s i a r l i x o n i i L i t t l e 5 l ( 4 8 7 9 0 ) ; O l t r s i z rs p p . 2 ( 4 I 6 6 2 ) ; 2 a
(4090i:i, 48231); 32 (48932); 5l (41456, 48781,
48799); 52 $82a2) (Olusiaceae).
flodiaetrrn variasattrnr (t..)Blurne 73 (40615) (Fltrphorbiaceae).
Cloclonarrthe crassifolia ( F'ockc) Morton 24 ( 10 17 l'l) ( Ciesneriacezrc').
C o f I e a : r r a b i c i rL . 2 8 ( 1 0 4 7 0 , 4 0 6 4 5 ) ( E u p l x r r b i a c e z r e ) .
C l o i x l a c l r r y r n a j o b i L . 9 , 5 6 ( . 1 0 5 , 1 0 ) : 4 2( 1 8 7 l t i ) ; C . s p . 9
( 4 1 4 8 0 )( P o : r c c a c ) .
(bleus x hybridus \trss 16 (4ft357); 28 (41476)(L:rrniaceae).
( l o l o c a s i a e s c u l e n t a ( L . ) S c h o t t 2 2 ( 4 u 1 4 1 1()A r a c e a e ) .
( b l u m n e a : u r s u s t a t a ( W i e h l . ) 1 . . S k o g 4 9 ( , 1 0 7 0 9 ,4 9 1 0 6 ) ;
(1. bilabiat:r Seern.48 (40519, 40600, 40816, 4U416);
(1. bvrsina (\\Iiehl.) Iivist et t.. Skog 55 (a8731); (1.
densibracteat:r Kvist & L. Skos 48 (4808,1); (1. clissiniilis (1. Mortor-r 48,49 (48178); C. eburnea (\\'ieh l . ) K ' , ' i s tc t [ , . S k t x r 4 8 ( 4 0 t ' ] 7 0 ) ; 4 8 , 4 9 ( a 8 2 l t 3 ) ; 5 2
(48712); C. fililoba Kvist et L. Skos 48,74 (41641,
,18997); O. firnbricah'x Kr.'ist et L. Skog 55 (.18923,
4 U 9 7 1, 4 8 9 7 7 ) ; ( 1 . s i s : r n t i f b l i a K v i s t e t [ . . S k o g 5 2
(415U7); (1. kienastiana Resel 48 (48383); 51
( 4 0 4 i r t t ) ; 7 3 ( . 1 8 1 6 2 ) ; 7 4 ( , 1 0 6 0 i r ) ;C l . l a e r , i s l h ' i s t e t
1.. Skos 5l (487!)a); (1. lchnrannii Mansf. 55
(1. lonsinen'osa lk'ist et t.. Skop; 57
( , 1 1 5 0 7 ) ;( i . m i n o r - ( H o o k . ) l { a n s t . 5 l ( 4 1 4 2 U ) ; 5 5
( 4 8 7 , 1 0 ) ;O . r n i n r r t i f l o r a K r , ' i s tc t [ - . S k o s 4 8 ( 4 8 9 3 7 ) ;
(4il9il3);
55 (4U970); O. nariniana (Wiehl.) Ik,ist et L. Skos
5 0 ( 4 i t 8 1 5 ) ; O . p i c u r K a r s t . 4 8 ( 4 i 1 7 3 0 ,4 0 7 6 0 , 4 0 8 7 1 ,
+808ti); 49 (40230); (i. purpurirnarsin:rta Kvist et L.
Skos 49 (480U5); (i. nrbriactrt:r (Wiehl.) l('ist et L.
Skos 48 (40872,48738);48,49 (40074),40880);48,
5 6 ( 4 0 6 7 5 ) ; 4 9 ( 4 8 0 1 3 8 ,4 8 1 2 8 ) ; 5 5 ( 4 0 3 5 9 ) ; 7 4
(11432,41622); (1. rtrbribracteata llvist & [,. Skog
4 8 , 4 9 ( ' 1 8 4 2 0 ) ;4 9 ( 4 8 0 8 7 ) ; O . n r b l i c n l l x [ ( l i s t & [ . .
143
Skog 55 (4874\);72 (41466);C. spathulataMansf.
48 (40797);49 (48344);73 (40a81);C. tenellaKvist
et L. Skog 24,72 (41577);C. spp. a8 (a8739);55
(48733)(Gesneriaceae)
.
(lorrosteeiacentronioidesMarkgrat 74 (40528,40779);C.
montana (Sw.)D.Don ex DC. 32 (48785)(Melastornakrceae).
Clcrrdiaspinescens L. 42 (41498); 54 (40390, 40495,41062,
48052); 59 (40663) (Boraginaceae)
Cord,vline terrninalis (L.) Kunth 56 (40640) (Agavaceae).
Cornutia spp. 22 (40397); 54 (40356)(Verbenaceae).
C o s t r r sl a e v i s R . & P . 2 5 ( 4 8 9 0 7 ) ; 3 7 ( 4 8 0 9 7) ; 4 0 , 6 6 , 6 8 , 6 9
( 4 0 6 7 u ) ; 5 5 ( 4 1 0 0 2 ) ; 7 2 ( 4 0 3 7 2 ) ; C l .l i m a K . S c h u m .
3 5 ( 4 U 0 9 3 ) ;C . p t r l v e r t r l e n t u sP r e s l 4 8 , 4 9 ( 4 0 1 l 6 ) ; 5 2
(40373); 66 (40700, 49103); C. scaber R. & P. 37
( 4 8 1 4 6 ) ;C . s p p . 2 5 ( 4 8 8 6 0 , 4 8 9 1 7 ) ; 4 1( 4 1 4 5 7 ) ; 6 5
( 4 1 1 1 t t 5 )6;6 ( 4 9 11 3 ) ( C o s t a c e a e ) .
Ootrssapozrcontorta Cuatrec. 32 (11626) (Urticaceae ).
(lranicliis sp. 48 (48927)(Orchidaceae).
Crernosperma consruens Cl. Morton 55 (a8979); 72
(41591); C. hirsutissirnunrBenth. 48 (41422,41437,
48737);49 (41126);7a G1581, 41611);C. rnuscicola
livist et 1..Skog 55 (48978); C. nobile C. Morton 48
( 4 8 9 7 6 ) ; 7 4 ( 4 1 6 5 1 ) ; O . r e l c l i o i c l e sK v i s t & L . S k o g 4 8
(48980) (fi.esneriaceae).
C l r e s c e n t i : rc u j e t e L . 6 ( 4 0 4 5 3 ) ; 6 , 3 6 ( 4 0 7 2 8 ) ( B i g n o n i a c e ae).
(lrinurn arrrabile l)on ex Ker-(l,arvl.4l (40547);72
(Arnarl'lliclaceae) .
(lrotalaria nitens H.B.K. 73 (40727) (Fabaceae).
ft0196)
(ltenitis sp. 48 (40145) (Dryopteridaceae).
(luatresizr riparia (H.B.K.)I{unziker
51
(40655); 55
( , 1 0 0 9 9 )C
; . s p . 7 2 ( 4 U 8 0 1 )( S o l a n a c e a e ) .
Ctrpania cinerea Poepp. & Endl. 2,31 (41046)(Sapindaceae).
Crrplrea tetrapetala Kohne 72 (40102, 40552,48699); 74
(11423); C. strieulosaH.B.K. l6 (40546);59 (40031)
(l.rtlrraceae).
(lvathula achyranthoicles (H.B.K.)Moq. 55 (40499) (Arnaranth:rceac).
C v c l a r r t h t r sb i p a r t i t u s P o i t . l 6 ( 4 8 8 5 0 ) ; 2 4 , 7 4 ( 4 0 2 2 1 ) ; a O
(48078) (Clvclanthaceae) .
(lvclodirrm trianae (Mett.) A. R. Srn. 4l (48757) (Drvoptericlaceae).
Cvperus luzulae (L.)Rctz. 70 (40035); (i.odoratus 1.. 74
( 4 16 2 4 ) ( C 1 ' p e r a c e a e.)
Cvphom:rndra hartwegii (Miers.)Dun. 5l (40i91) (Sol:inirccae).
Dacr-v<rdesgranatensis Cluatrec. 2 (4167 6) (Burser:rccae).
Dahlia sp. 16 (40642) (Asteraceae).
Dalechanrpia sp. 48 (48?13) (Euphorbiaceae).
D a n z r e : rh u r n i l i s M o o r e 5 l ( 4 8 0 6 2 ) ; D . s p . 4 8 ( 4 t 3 9 4 1 ) ;
57 (4,3437) ( Nlarattiaceae ) .
BS 46
144
Dendropanax
arbonerts (L.)Decne. 5l
(40899) (Aralia-
Eleoclrzrris retroflexa
(Poir.)Urban
72 (18327)(C_vperar-
ceae).
ceae).
Dennstaedtia zrrborescens (Willcl.)Maxon 73 (400f15);
D. spp. 25 (4888a); 72 (48889) (Dennstaecltiiiceae).
D e s r n o d i u n r a d s c e n d e r - r s( S w . ) D C . 5 5 ( 4 1 5 7 8 ) ; 6 4 ( 4 0 0 2 2 ) ;
73 (40032); D. sp. 53 (41075) (Fabaceae).
Desnroncus cirrhifera Gentrl' & Zarclini l0o 24 (ala55); D.
s p . 7 ( 4 8 1 9 7 ) ( A r e c a c e a e ).
f)ialyanthera sorcloniifblia \Alarb. 2 (41664); D. spp. 2
( 4 0 3 5 4 , 4 1 0 4 9 , 4 8 2 9 0 ) ( M y r i s t i c a c ea e ) .
Diasrema affine Fritsch 48 (a8990); 68 (491I I ); 7l
(40140); D. scabnrrn (Poepp.)Benth. ex. \{'alp. 73
( 4 9 0 9 2 ) ( C e s n e r i a c e a e) .
Dichorisandra anqtrstifblia Linclen & Rodig. 54 (40tt98);
D. lrexandra (Atrbl.)Standl. 48 (4U188);57 (18407);
6 8 ( 4 0 3 7 2 ) ; D . s p . 6 5 ( 4 1 ' 1 8 4 )( C o m m e l i n a c e a e ) .
I)icranopteris pectinata (Willd.)Und. 44 (483?3) (Gleicheniaceae).
48, 49
Dicranoglossurn polypocliodes (Hook.)Lellineer
( 4 0 6 4 9 ); 4 9 ( 4 0 1 7 0 , 4 9 0 2 9 ) ; D . s p . 4 9 ( 4 0 2 3 4 ) ( P o l - v podiaceae).
Dieffenbachia dasuensis Engl. 3l (41652); D. segtrine
(facq.)Schott. 46 (40756, 48154,48348) (Araceae).
D i o c l e a s p . 7 ( 4 1 6 4 6 ) ( F i r b z r c e a e. )
D i o s c o r e a e s c u l e n t a B u r k . 2 3 ( 4 8 3 3 1 ) ; D. 'p. 24 (48826)
( D i o s c o r ea c e a e ) .
Diplarpezr paleacea lriana ex (losn. 72 (48723)(Melastom a t a c ea e ) .
Diplazir.rm cristatum (Desr'.) Alstor-r 72 (48320); l). mocccn i a n u m ( S o c l i r o ) C l .C h r . 2 5 ( 4 8 0 4 9 ) ; D . s t r i a t a s t n t m
Lellinger 39 (a023U); a8 (a0168); 73 (40051,
4904I ) (Dryopteridaceae) .
f)iscophora guianensis Miers. 32, 64 (4Bl68); 64 (40410,
407 41, 41022) ; 72 (40305) (Icacinaceae).
Dracontium sp. 50 (41597) (Aracezie).
Drymaria cordata (1..) Willd. 68 (40179, 49096) (Carvophyllaceae).
D r y m o n i a a l l o p l e c t o i d es H a n s t . 4 8 ( 4 8 2 1 2 ) ; 6 2 ( 4 0 0 6 1 ); D .
c o r i a c e a ( O e r s t . e x H a n s t . ) W i e h l . 2 6 ( 4 0 1 1 5 0 ) ;3 2
( 4 8 0 8 3 ) ; 4 8 ( 4 0 6 7 3 ) ; 4 9 ( 4 8 0 9 2 ) ; O . e c u a d o r e n s i sW i -
Encyclia fragrans (Srv.)l,ernee 52, 56 (4066f]) (Orchidaccae).
Epidendnrrn difforme.f acq. 28 (40102) (()rr:hidaceae).
Erato polvmnioides DC. 5l (41505) (Asteraceae).
Erechtites valerianifolia
(\\krlf.)D(-,. 74 (48866) (Aster:rce-
ae).
Ervrrsitrnr lbeticlurn L.29 (10127,48926); 29, 51 (a8359);
4l (40038) (Apiaceae).
E r y t l i r i n a e c h r l i sT r i a n a l 7 ( 4 0 t 4 6 ) ; E . s n r i t h i a n a K r u k o f f
17 (40632) (Fabaccac).
Erythrodes ecuadorensis (l.rav 48 (4078U); E. rveberana
C l r a y4 9 ( 4 0 U 7 9 ) ; E . s p . 5 6 ( 4 0 3 5 2 ) ( O r c h i c l a c e i r e ) .
Firythroxvlurn
novogranatense Hierou
in Engl. 40
( 4 I 5 8 4 ) ( E r y t h r o x v l a c e a e ).
F l s c h r v e i l e r ac o r i a c e a ( D C l . ) B e r s e x M a r t . 2 ( 4 8 9 9 5 r ) ;l i . s p .
2 (41668) (Lecythidaceae).
Etrsenia sp. 64 (40044) (Myrtaceae).
E r r p l r o r b i z rc o t i n i f b l i a L . 1 7 , 5 4 ( 4 0 1 7 7 ) ( L , u p h o r b i a c e : r e r ) .
E u t e r p e c h a t r n o s t a c h y sB u r r e t 2 4 , 2 5 ( 6 0 1 0 3 ) ; 2 5 ( 6 0 0 0 1 )
(A'ecitceae).
E r . ' o d i a n t h u sf t r n i f e r ( P o i t . ) l . i n c l m . 4 6 ( 4 0 0 1 1 0()C y c l a n t h a cerac).
Ficus calclasiana Dusancl 33 (,10096); F. cervantesian:r
S t a n c l l e r&
' L . O . W m s . 2 ( 4 0 7 6 1 1 )F; . i n s i p i d a W i l l c l . 2 ,
44 (40900); 3l (a1080); F. rnaxima P. Miller 2
( 4 8 2 0 3 ): 3 1 , 4 4 ( 4 8 0 5 6 ) ; F . o b t u s i f b l i a H . B . K . 3 2 , 3 3
( 4 0 0 1 7 ) ; F . s c h i p p i i S t a n c l l .3 2 ( 4 8 1 2 4 ) ( M o r a c e a e ) .
F - i m b r i s t y l i sc l i c h o t o m a ( L . ) V a h l . 6 6 ( , 1 8 3 5 3 ) ; p . r n i l i a c e a
( L . ) \ A r a h l .6 6 ( 4 f t 3 1 1 8()O v p c r a c c a c ) .
Fischeria ireqtratori:rlis Spelhnan 6 I ( 4tt3i34)(Asclepeclizrceae).
Gasteranthus corallinus (Fritsch)Wichl. 48 (,18094); 73
(40665); C).crispus (Mansf.)Wiehl. 48, a9 (a0701);
(]asteranthus ol-lcoqastnls (Hanst.)Wiehl. 49, 73
(40233); 55, 72 (,11000); 72 (40327): 7+ (40731,
4 U 12 6 ) ( ( ) e s n e r i a c e a e ).
Genipa anrericana 1,. l5 (49120).
( ] e o s e n a n t h t r s r h i z a n t h u s ( U l e ) B n r e c k n e r 5 l ( 4 8 0 6 1 3 ) 5; 5
e h l . 4 8 ( 4 8 3 11 ) ; D . m a c r o p h v l l a ( O e r s t . ) H . E . M o o r e
4 8 ( 4 0 1 6 7 ) ; D . r h o c l o l o r n aW i e h l . 4 8 , 4 9 ( 4 0 1 0 8 ) ; 7 3
(40716); D. serrtrl:rta(facq.)Mart. 32 (48081); 48
( 4 0 3 1 9 ); 7 3 ( 4 0 1 0 6 ) ( O o r n n r e l i n z r c e a e ) .
Cl.eonoma cuneata H. A. \Alencll. 15 (40207); 24 (40753,
60099);24,52 (4fl35l); G. uracilisH. A. Wencll.ex
Spnrce 24 (60099, trOl04, 60118); (i.. linearis Brrrrct
(18773); 48, 49 (48393); D. ttrrrialvae Hanst. 48
(4163t]); 73 (40224); D. warszewicziana Hanst. 48
( 4 0 0 U 0 ,4 8 1 2 1 , 4 8 , 1 1 9 ,4 U 9 9 1 ); 5 5 Q B 7 2 B ) ( G e s n e r i a -
72 (601l5); 74 (60115); Ci. spp. 8 (60004), 25
( 4 8 9 1 6 )( A r e c a c c a c ) .
()eophila sracilis (R. & P.) D(1. 73 (4064,1)(Rubiaceae).
ceae).
Dryopteris sp. 25 (48887) (Dr,vopteridirceae).
Dussia sp. 2 (41656) (Fabaceae).
(Barv & Fee)Moore
Elaphoelossr.rrn herminieri
(40533);E. spp. l3 (40u2ti);72 $BBa\;
(Drvoptericlaceire ) .
l3
73 (a0133)
G l o x i n i a d o c l s o n i iW i e h l . 4 8 ( 4 1 4 6 2 ) 1 8 2 7 9 ) ; 5 2 ( a 0 3 6 7 ) ;
5 5 ( 4 8 7 2 9 ); 7 4 ( 4 1 6 0 2 ) ( ( ) e s n c r i a . c c a . c ) .
( i , c r r n p h r e n aq l o b o s u s L . 7 2 ( 4 0 5 6 1 ) ( A r n a r a n t h z r c e a e ) .
fiorrzalaguniir cornifolia (H.B.K.) Standl. 72 (a8227); ().
sp. 73 (40661) (Rtrbizrceae).
(Malr.accae).
Gossypiunr barbaclense 1.. l0 (401-rtt4)
BS 46
t45
Cotratria lrrptrloicles (1..)Urban 24
( 4 10 8 6 ) ( R h a n r n a c e a e ).
( ] u a r e a s p . 2 ( 4 8 2 1 7 )( M e l i a c e a e ) .
(40741); (). sp. 7
C l ' r r a t t e r i as p p . 2 ( ( 4 0 7 3 6 , 4 l 6 5 5 ) ( A n n o n a c e a e ) .
Clurania eggersii Sprague & Hutch. 5l (40052); G.nregistantha.f. D. Srn. 60 (48308); G. spinulosa (Poepp. &
E n d l . ) ( b s n . 6 7 ( 4 8 1 5 6 , 4 8 1 7 0 ) ; C i . .s p . 4 l ( 4 8 1 0 7 )
(Cucurbiftrccae) .
(l.ust"avia
c l o c l s o n i iM o r i 2 5 ( . 4 0 3 4 0 ) ; ( i . s p . 2 5 ( a i l l a l ) ( L e cy'thidaceaer).
(]rrznrania lingulata (I-.)Mez 4l (48377); G. melinonis Rerg e l 4 l ( 4 0 8 8 5 ) ; ( ] . s r : h e r z e r i a n : rM e z 3 2 ( 4 8 3 9 5 ) ; G .
spnrcei (Andrc) 1..8.Snrith 58 (48693); fl.. testudinis
L.B. Srnith & R.W. Retrd 20 (48856); G. sp. 4l
( 4 I 0 I 9 ) ( B r o r n e l i a c e a e ).
Srv. 72 (a8225); 73 (40054, 10484,
40626); H. m:rcr:rntha l.ittle 13 (40076); H. sp. 52
( 4 1 0 5 1 ) ; H . s p . 7 4 ( 4 1 6 0 U )( R u b i z i c e a e ) .
Heclvchitrm coronariurn Koenis 16 (41ir39); 19 (49121);
7 4 ( 4 8 7 B B )( Z i n u i b e r a c e a e )
Hanrelia
axillaris
Hedyosmum scoterrir-nrrnr Stancll. 7 2 (11,174)( Chkrran thaceire).
H e i s t e r i a s p p . 2 ( 4 0 3 5 8 ); 7 3 ( 4 0 0 6 4 ) ( O l a c a c e a e ) .
Heliconia aernvscliana Burle-Marx 53 (40120); H. ctrr.
t i s p a t h a O . ( } . .P e t e r s .S l ( 4 0 4 3 4 ) ; 7 3 ( 4 0 1 1 9 ) ; H . l a tisp:rtha Benth. 72 (4U228); H. longa (Griggs)
Winkl. 38 (48180); H. stricta Huber 32 (18274)
(Musaceae).
(L.)Presl 72 ,40l3a); 73
Hemidictl'rrm marsinatum
( 4 0 I 2 8 ) ( D r v o p t e r i c l i r c e a e ).
Herrania
balaer-rsisPretrss. 24 (10416, 40666) (Sterculia-
c e a e) .
Heteranthera renilormis R. & P. 60 (40388, 48390, 18776)
( P o n t e d e r i a c e a e ).
Hcteropsis obkrneifbli:r Kunth l0 (40001) (Araceale).
H i b i s c u s r a c l i a t t r s C a v . 7 3 ( 4 0 6 3 7 ) ; H . r o s a - s i n e n s i sL . l 6
(40429,40461); 73 (40139); H. sp. 72 (40579) (N{alr'aceae).
Hieroninra chocoensis (hatr. 3l $8772); H. laxillora
(Tull) Miill. 2 (40507) (Etrphorbiaceae).
H o f f m a n n i a s p . 7 3 ( , 1 0 1 5 1 )( R u b i a c e a e ) .
H t r m i r i a s t r - u r np r o c e r t r r n ( L i t t l e ) C u a t r . 2 ( 4 1 0 7 6 ) ( H o u r n i riaceae).
linifblia (L.)Trevisan 73 (40610)(l,vcopocliaceae).
Hvdrangea sp. l6 (40641) (Hlclrangeace:re).
Hvdrocotyle leucocephala (1.& S. 73 (40193) (Apiaceae).
Hvmenophyllurn sp. 48 (487 51) (Hvrnenophvllaceae).
Huperzia
59 (40635); H. mutabilis (A. Rich.)Briq. 14, 72
(40039);H. obtusifloraPresl37 (48720);41(40294);
73 (40180); H. verticillataJacq. 42 (al0B3); 72
(41 429); 74 (48852)(Lamiaceae).
Ichnanthus axillaris (Nees) Hitchc. & Chase 48 (48090)
(Poaceae).
Impatiensbalsaminat.. l6 (48362)(Balsaminaceae).
Inga edulis M:rrt. 3, 24 (48200);24 (40353,40450,40778);
24,73 (40042);I. marginataWilld. 24 (40733);I. ruizianaDon. 24 (40898);I. spectabilis(Vahl)Willrl.24
(40772);I. sp. 35 (41087)(Mimosaceae).
Ipomoeabatatas(L.) Larn. 23 (48305);I. spp.51 (a0005);
62 (40476) (Oonvolvulaceae)
.
IriarteacleltoideaR. & P. 2, 4, 5,25,27 (60097);4 (48409)
(Arecaceae).
Ischnosiphon leucophaeus (P. & E.)Koern. 10 (40780,
4f1902)( Marantaceae) .
I s e r t i a p i t t i e r i S t a n c l l .2 ( 4 0 8 2 7 ) ; 3 2 ( 4 8 2 7 8 ) ( R u b i a c e a e ) .
Ixora sp. l6 (40463) (Rubiaceae).
Jaltomirt:r procumbens (Car:)Clentr,v52 (40013); 54, 55
(40I0I) (Solanaceae).
.fatroplra clrrcas Wall. 43, 44 (40633) (Euphorbiaceae).
. J e s s e n i ab a t a u a ( M a r t . ) B u r r e t 4 ( 6 0 0 7 9 ) ; 4 , 2 5 ( 6 0 0 0 6 )
(Arecaceae).
. | r r s t i c i ac o r n a t a ( l - . ) L a n r . 2 2 ( 4 8 1 4 4 - ) ; 7 a ( 1 8 7 1 5 ) ; . | . i a n t h i n a \ { r a s s h .5 3 ( 4 8 3 8 4 ) ; . f . s t e r e a 1 8 ( 4 0 5 3 5 ) ; . J . s p . 7 1
( 4 14 8 3 ) ( A c a n t h a c e a e ).
Kalanchoe sp. 74 (48149) (Crassulaceae).
Kolrleria spp. 48, 49 (40576); 49 (40723) (Gesneriaceae).
L,acistema aggregata (Berg) Rtrsbr, 24 (48796) (l,acistemaceae).
Lantana spp. 51, 55 (41077); 56 (41468); 65, 71 (4054U)
(Verbenaceae).
L a s i a c i ss p . 7 0 ( 4 1 4 6 9 ) ( P o a c e a e ) .
Leandra sranatensis (]leason 52 (40326) (Melastomaraceae).
Linclernia cnrstacea (L.) F. Muell. 40 (40636); t.. sp. 4l
(187 22) ( Scrophulariaceae ) .
Lindsaezr quadrangularis Radcli 72 (48953) (Adianthaceae)
Lonrariopsis japurensis (Mart.)J. Sm. 4l (48125); 41, 73
(40163); 73 (49038); f.. nigropaleata Holtttrm 4l
( , 1 0 0 2 0 ,4 i l ? 8 4 ) ; 6 4 ( 4 0 3 3 3 ) ; 7 3 ( 4 0 2 2 9 ) ( D r y o p r e r i daceae).
l.crrrchitis hirsuta L. 25, 73 (49036); 73 (a0130)(Dennstaedtiaceae).
Lonchocarpus nicou (Aubl.) DCl. 33 (49107); t,. sp. 33
( 4 9 1 0 5 )( F a b a c e a e ) .
l.uclovia intesrifolia
(\4loocls.)Harl. 10, 7 (48281) (Cyclan-
H y o s p a t h e e l e s a n s M a r t . 3 2 ( 4 1 J 2 7 3()A r e c a c e a e ) .
Hvpolepis hostilis (Kze.)Presl25 (40/t27,41070); H. repens (L.) Presl 25 (48048, 4U299) (Dennstaedtia-
Lndrvisia erecta L.73 (40178) (Onagraceae).
l.uf fa aeer''pticaMill. 24 (40597) (Cucurbitaceae).
ceae).
H v p t i s c a p i t a t a J a c c l .2 8 ( 4 0 1 1 4 1 ) ; 3 7( a 8 3 5 t r ): 5 4 , 5 5 ( 4 0 3 8 9 ) ;
L 1 ' c i a n t h e sa m a t i t l a n e n s i s ( C o u l t . & . 1 . D . S m . ) B i t t e r 4 7 , 5 5
(10172); 48 (40087) (Solanaceae).
thaceae).
T
t46
BS 46
Lvcopocliella trianae (Hieron.)B. @lls. 65 (41499) (L_vcopodiaceae).
Lvcopodium sp. 48 (48ll34) (Lvcopodiaceae).
Macleania rotunclifolia Sodiro & Hoer. 48 (48376) (Ericaceae).
Macltrra tinctoria (L.) Steud. 58 (49060) (Moraceae).
Macrothelypteris
(Gaucl.)
torresiana
Chins
(4903 I ) (Thelvptericlirceae)
73
Malachra rtrderalis Gtirke 73 (40557); 73, 74 (40860)
(Malvaceae).
Malaxis sp. 73 (48709) (Orchidaceae).
Malvaviscus pendulilkrnrs f)C. 21, 73 (40027) (Malvaceae).
Manclevilla dodsonii Gentr,v I (40439); 32 (48307); l,l
hirsuta (Rich.)K. Schum. 64 (40800) (Apocvnaceae).
Manihot esculenta (lr-antz 23 (48329, 48330)(Euphorbia-
Nautilocalr,x sp. 55 (48936, 48964) ((iesneriirceae).
Neornortoni:r rosea Wie hl. 48 (11477) (Ci.esncriaceae).
Nephele:r cuspidaur (Krrnze)Tryon 25 (48176)(Cyatheaceae).
Nephrolepis pectinata (Willcl.)Schott 48 (48846); 50
(48135); N. pelrclula (Radcli)J. Srn. 48 (4899U); N.
spp. 13 (48837, 48845) (Dzrvalliaceae).
N o t h o p : r n a x l r u c t i c o s u s ( [ . . ) M i c 1 .7 3 ( 4 0 6 1 2 ) ( A r a l i a c e a e ).
N o t _ v l i :r i m b a c h i i S c h l r r . 5 7 ( 4 8 1 4 3 ) ( O r c h i d a c e a e ) .
O c i r n u m c a m p e c l - r i a n u r nM i l l . 2 8 , 4 1 ( 4 0 6 1 6 ) ; 2 9 ( a 8 9 8 2 ) ;
74 (40395); O. sp. 58 (48445) (Larniirceae).
O d o n t : r c l e n i a m a c r a n t h a ( R . & S . )M e f . 6 l ( , 1 0 8 9 5 ) ;O . s t r i c tum (Nees.)Kuntze 21, 73 (40023) (Apocynaceae).
Oenocarpus mapora Karst. l0 (60102) (Areczrceae).
Oleanclra sp. l3 (48303); 48 (48843) (Drvoptericlacezre).
ceae).
Mansoa hvrnenaea (DCl.) Gentry 68 (49089)(Bignoniaceae).
Marcgraviir sp. 32 (48337) (M:rrcuraviaceae).
O s s a e al a x i v e n u l a W u r d a c k 5 8 ( 4 8 1 7 3 ) ; 7 g ( 4 0 3 6 4 , 4 1 0 1 3 ) ;
O. micrantha (Sw.)Macf.ex Cogn.24,73 (401tr2); 70
I\{arila laxiflora Rusby 2 (41615) (Clusiaceae).
M a t i s i ac o l o r a d o r u r n B e n t h . 2 , 2 4 ( 4 0 0 1 5 ) ; 2 + , 7 3 ( a 0 1 3 1 ) ;
73 (40187); M. soegenii (luat. 24 (41660) (Bombaca-
(,10048)(Mclastornataceae) .
Oxalis nricrocarpa Benth. 49 (40726); O. spp. 15 (41580);
3 4 ( 4 1 1 3 3 ) ; 5 3 ( 4 t 3 3 3 3 )( O x a l i d a c e a e ) .
ceae).
Melothria perclula L. 47 (40480); 55 (40098) (Ctrcurbitace-
Palicoru'ea conf'erta (Benth.)Sanclrv. 74 (40518); P. suianensis Atrbl. 32 (48205); 74 (11003); P. spp. 32
( 4 16 1 9 , 4 8 7 2 1 , 4 8 72 7 ) ( R u b i : r c e a e ).
ae).
Mendoncia sp. 28 (40813) (Acanthaceae).
Miconia barbinervis (Benth.)Triana 74 (48431); M. erioclada Triana 74 (40569); M. goniostigrna Triana 72
(41500); M. sracilis Triana 74 (41033)l M. nervosa
(Smith)Triana 74 (41014, 48099); M. oraria \A/urclack 32 (48216, 4tt339); M. theaezans (Bonpl.)
Ccrgn. 72 (41495); M. trinen'ia (Sw.)D. Don ex Lou c I . 7 4 ( 4 0 4 8 9 ) ; M . r , e n u l o s aW u r d a c k 7 3 ( 4 0 3 2 4 ) ; M .
s p p . 1 4 ( 4 1 6 1 0 ) , 4 8 ( 4 t 3 7 0 8 )( M e l a s t o r n a t a c e a e ) .
M i k a n i a s p p . 4 5 ( 4 1 5 0 1 ) ; 4 8 ( 4 0 3 4 1 1 )5; l ( 4 1 0 3 6 ) ( A s t e r a ceae).
Mimosa elliptica H.B.K. 22 (40483); M. puclica L^ 22
(48324) (Mimosaceae)
Minquartia guianensis Aubl. 2 (41023); 2 (48400)(Olacaceae).
Mirabilis.jalirpa L. 21, 73 (40025) (Ni,ctaginaceae).
Monolena primulaeflora Hook. f .44 (41582,48974) (Melastomat.aceae)
Monopyle socliroana Fritsch 37 (a0877): a8 (40513,
4 8 1 2 7 ); 7 3 ( 4 0 3 1 6 ) ( G e s n e r i a c e a e ) .
Monstera adansor-rii Schott 73 (a9054); M. lechiriana
Schott 55 (41633); M. spmceana (Schott) Eneler 48
( 4 0 8 4 3 ) , 4 8 1 0 5 ) ; M . s p . 4 0 ( 4 0 4 3 6 )( A r a c e a e ) .
M v r c i a s p . 7 4 ( 1 0 7 4 3 , 4 1 6 1 4 )( M y r t a c e a e ) .
O r y c t a n t h u s o c c i c l e n t a l i s ( L . ) E i c h l . 7 3 ( 4 0 0 5 f 1 )( L o r a n t h a ceae).
P a n i c u m f i o n c l e s c e n s G . .M e e . 7 3 ( 4 0 1 8 2 ) ; P . t r i c h o i d e s S r v .
74 (48489) (Poaceae).
P a r a d r y m o n i a s p . 4 8 ( 4 1 6 3 5 , 4 8 9 6 3 ) ( ( ) c s n e r i a c e a e ).
Parasonia pvrarnidata (t.. Rich.) Rtrr. 7 (48413) (Bignoniaceae).
Paspalum conjugatum Bersius 66 (40399, 41015, 18147)
(Poacerzre).
P a s s i f l n r aa u r i r : u l a t a H . B . K . 2 4 ( 1 0 0 2 6 , 4 0 2 1 8 ) , 4 5 ( 4 0 1 0 5 ) ;
P . f b c t i c l aL . 2 4 ( 4 0 3 8 7 ) ; 2 8 ( 4 0 6 6 0 ) ; P . r n a l i f o r m i s L .
; . q u a d r : r n s u l i r r i sL . 2 4 ( 4 0 3 5 5 , 4 U 1 8 7 ,
2 4 ( 4 0 1 1 3 9 )P
48429);24,45 (10174); I'. sp. 24 (48857) (Passifloraceae).
Parrllinia firscescens Ktrr-rth. I (4U426); P. r-robilisRacllk. 32
( 4 8 3 4 1) ; P . s p . l 0 ( 4 1 4 3 5 8 ) ( S z r p i n d a c e a e ) .
Pzrvoniacastaneifblia St.-Hil. & Naudin 53 (4U20,1):P. fiutic o s a ( M i l l . ) F a w c e t t & R e n d l e l 6 ( 4 0 1 1 5 5 ) ; 5 9( 4 0 0 2 9 )
(Malvaceae).
Pecluma consimile (Mett.) M. (). Price l3 (48436) (Polvpocliaceae).
Pentasonia erancliflorir Stancll. 24 (al1199); P. rnacrophy'lI r r r nB e n t h . 2 4 ( 1 0 4 5 1 ,4 8 3 6 1 ) ; P . s p p . 2 ( a l 0 3 5 ) : 2 a
( 4 0 3 7 5 ,4 1 0 0 1 ); 2 6 ( 4 1 0 8 2 )( R u b i i i c e a e ) .
(40671); 73 (49108)
Peperorni:r macrostachya (Vahl)A. Dietr. 48 (a8202); P.
ornnicola DC. 32 (48986); P. panarnensis DC. 2l
(40638); P. pellucida (L.) H.B.K. 62 (48287); P. per-
N a u c l e o p s i sa m a r a D u c k e 3 3 ( 4 8 9 1 2 ) ; N , s p . 3 3 ( 4 9 1 1 2 )
(Moraceae).
nambucensis Miq. 73 (49053); P. pteroneura DC. 7l
( 4 0 0 8 9 ) ; P . s e r p e n s ( S r v . ) l - o t r c l .4 8 ( 4 1 4 i 9 , 4 8 2 1 0 ,
4 B B 2 l 1 )P
; . urocarpa F. & M. 35 (40531); 55 (40301);
Napeanthus robustus Fritsch 70,72
(Cesneriaceae).
747
BS 46
48113, 48157, 48346,, 48724, 48824, 48858, 48909) ;
; . s p p .1 4 ( 4 8 0 7 6 ) ; 1 4 , 3 5
7 0 ( 4 1 0 1 u ) ; 7 3( 4 0 1 3 5 ) P
50 (41637);5r (40370);5a G0782);55 (a02e5): 65
( 4 0 5 0 6 ) ;1 4 , 7 3 ( 4 0 1 6 9 ) ;a l ( 4 0 1 3 7 ) ;4 8 ( 4 1 5 8 9 ,
(a0670); 70 (40109, 40341); 72 (48726, 48853,
48906);
4t625, 48193,48369,48705, 48719,481169,
48861); 73 (40679, 40683, 40699, 40711, 40776,
; 0 ( 4 8 8 2 1 ) 5; l ( 4 8 7 9 3 ) 5; 2 ( ' 1 1 5 9 3 ) ;
4 8 , 4 9 ( 4 1 6 4 5 )5
40790, 49056, 48158); 74 (40323, 41590, 41658,
5 5 ( a 0 3 1 3 ) ;6 2 ( 4 l l 3 1 a ) ;6 5 ( 4 0 0 7 9 ) ;7 0 ( 4 0 5 7 8 ,
48163, 48164, 48392, 48697, 48703, 48836,
40777, 48072);72(4078{;73 G0143)(Piperaceae).
4 t 1 8 7 3 )( P i p e r a c e a e ) .
Perebeaxanthochl'maIierst.2 (4ti201)(Moraceae).
P e r s e aa m e r i c a n a| ' . . 1 9 , 2 4 ( 4 9 1l 8 ) : 7 3 ( 4 0 7 1 5 ) ( L a r r r a - P i t c a i r n i a s p p . 7 ( 4 1 4 3 6 ) ; 3 7 ( 4 1 4 9 6 ) ( B r o m e l i a c e a e ) .
ceae).
Pharus latifblius L. 65 (60106) (Poaceae).
Phaseoltrs vulg;aris L. 23 (40516) (Fabaceae).
ineqrtilaterum Liebrn. 46 (40171); P. steyermarkii Bunt. 32, 64 (41 168); P. strbhastatum Engl. 7
(48179); P. tenue C. Koch & Augustin 73 (a0226);P.
verruc()sum Mathieu ex. Schott 10, 32 (a8l9l); P.
Philodendron
spp. 7 (40849, 4824r); 46 (401rb): 72 (40310)(Araceae).
Phoradendron chrysocl;rdon Gra,v 52 (40350); P. piperoides (Kunth)Trel. 57 (48367) (Loranthaceae).
Phthirtrsa pvrifolia (H.B.K.)Eichl. 52 (40345); 52, 57
( 4 8 1 5 1 ); 7 3 ( 4 0 0 5 6 ) ( L o r a n t h a c e a e )
Phyllanthus anisobolttsM. Are. 33 ( (40010, 10394,48370),
4BBB0, 49102); P. stipr.rlaris (Raf.)G. lt'ebster 22
(40392) ; P. sp. 22 (48231) (Euphorbiaceae).
Phr,'salisangulata L.. 24 (40609, 48385); P. pubescens 1,. 24
( 4 0 3 1 1 ) ; P . s p . 7 0 ( 4 1 4 6 0 )( S o l a n a c e a e ) .
Phvtelephas aequatorialis Sprtrce 8, 24 (6011l) (Arecaceae).
Phvtolacca rivinoides Kunth & Bouche 13 (40003, 48830);
25 (40454); 29 (4810i) (Phy'tolaccaceae).
Pilea spp. 40 (40062, 40107,40217,40682);48 (40378);58
( 4 0 1 8 5 ) ; 7 3 ( 4 0 0 0 8 )( U r t i c a c e a e ) .
Pilocosta oerstedii (Triana)Almeda & Whiflin 48 (48387)
(Melastomataceae) .
Piper arboreum Aub. 47 (40455); P. angttsturn Rudge 47
(40303)i 48, 73 G045\;
a8 (40738)i 56, 73
(a0090); 74 (40749); P. brachypoclon Benth. 74
(40881); P. cararense T. & Y 5l (48174,48244);51,
55 (41021);P. drvadum DC.74 (41612);P. eustyltrm
(40237); P. filistylurn DC. 48 (10746,
P. florencianrtm Trel. & Ytrnck. 48 (40338);
P. hispidum Sw. 70 (40222); 73 (40194, 40681);74
(40748); P. irnperiale (Miq.) DC. 56, 74 (40199);
70(a0091); 73 (40596); P. mareinattrm Jacq. 13
(40012); P. rnultiplinenium
DC. 4l (40059); 73
Diels 72
48ll6);
(406t]4) ; 74 (11471, 4B7BO); P. obliquttm R. & P. 72
(40210); P. peltatum l.. 74 (48159); P. phytalaccaelbliurn Opiz in Prcsl 5l (40468); P. squamttlosttm
DCl. 48 (41640); P. triar-raeDC. 56 (41020); 73
(49055); P. tricuspe (Miq.) DC. 47 (40859); P. tti; . spp. 15 (41026); 18
b e r c u l a t u m . l a c q . 4 7 ( 4 t i 3 7 1 1 )P
(41600)i 10, 4L, 68
G0325); 32 (a8338); al
(49043); 48 (40369, 40752, 40793, 41120, 41421,
Pithecellobium longifolium
(Mirnosaceae).
Pityrosranrma calomelanos
(48221) (Ptericlaceae).
(H.B.K.)Standl. 74 (4B2BB)
(L.)t,ink
4l
(18752):
74
Plantaqo major L. 65 (40599) (Plantaginaceae).
Plerrrothallis picta Lindl. 57 (18432) (Orchiclaceae).
Plrrmeria rubra L. 54 (40594) (Apocynaceae).
Podandrogyne brachycarpa (D. C.)\Alood. 13 (40017); P.
brevipedunculata Cochrane 5l (40057, 40166); 53
( 4 0 2 0 9 ); 7 4 ( 4 0 8 6 6 ) ( C a p p a r i d a c e a e ) .
Polemonium sp. l6 (48364) (Polemoniaceae).
P o l l a l e s t a s p . 7 4 ( 4 1 4 9 2 ) ( A s t e r a c e a e) .
Polybotrya caudata Kunze 65 (41058); P. lechleriana Mett.
25 (48848) (Dr1'opteridaceae) .
Polygala nrollaginifolia St. Hill. l6 (405il8); P. paniculata L.
l6 (40590) (Polygalaceae).
Polyeonum acuminatum H.B.K. 33 (4U412) ; P. hydnrpiper o i d e s M i c h x . 3 3 ( 4 0 4 9 1 , 4 0 4 9 4 )( P o l y g o n a c e a e ) .
Polypodium crassifoliunt L. 73 (40639) ; P. fraxinifolium Jac q . 6 5 ( 4 0 7 9 8 ) ; P . p e r c u s s u m ( l a r ' .4 8 ( 4 0 1 9 6 , a 0 2 1 9 ) ,
5l (40629); P. phyllitidis L. 40 (40u06); 7r, 73
(,10095);P. piloselloides L. 35 (40302); 5l (40236,
40475);73 (40617); P. repens Aubl. 70,72 (a0672);
P. .pp. 13 (40795) (Polypodiaceae).
Portulaca oleracea L. 56 (4060a); P. spp. 16 (48378,
48380); 39 (a8379) (Porttrlacaceae).
P o t h o m o r p h e p e l t a t a ( L . ) M i q . 5 l ( 4 0 0 0 4 ): 5 6 , 7 4 ( 4 0 3 9 3 )
(Piperaccac).
Poulsenia armata Miq l0 (48343, 48985) (Moraceae).
Pourotrma guianensis Aubl. 24 (40677); P. hirsutipetiolata
Milclbr. 24 (41009) (Urticaceae).
Pouteria caimito (R. & P.)Radlk. 24 (40415); P.collina (Lit
tle)Penn. 2,3,24 (41673); P. torta (Mart.)Radlk. 2,
24 (41629) (Sapotaceae) .
Pouzolzia sp. 55 (40477) (Urticaceae).
Prestcreasejuncta Bailey 7, Z5 (60\0 7) (Alecaceae).
(40831,
Prestonia portobellensis (Beurl)Woodson 6l
41052): P. rotundifolia K. Schum. ex Woodson 6l
(40780) ; 72 (4067 4) (Apocynaceae) .
Protiurn amplum Cuatr. 2 (41659); P. sp. 2 (41006)(Btrrseraceae).
Psamrnisia attaberrans 26 (48250); P. caloneura A. (1.
Srnith 24 (48920); P. f'errusinea A. (1. Smith 20
(48792); P. spp. 26 (40309, 40520, 48061)(Ericaceae).
148
BS 46
Pseuclelephantopus puratis (l,ess.)Cronq 72 (48381); P.
s p p . 7 4 ( 4 1 6 0 9 , 4 8 7 1 3 ) ( A s t e r a c e a e ).
S c a p h y e k r t t i sg r a m i n i f o l i a ( R . & P . ) P . & E . 6 8 ( 4 U 3 1 8 ) ; S .
prolifera (loen. 28 (40855); a8 (aBl96)(Orchicla-
Pseuderanthemum lanceolatum (R. & P.)Milbr. 73
(40211); P. micranthunr Leonard 3l (48935)(Acanthaceae).
Psidiunr euajava L.24 (41335); P. sp. 24 (40104) (Myrtace-
ceae).
Schefflera sp. 32 (41606), 48718, 48u03, 48gl3)(Araliaceae).
ae).
Psychotria brachiata Sw. 32, 74 (48112); P. czrerulea R. & P.
74 (18172); P. hoflmannsessiana (Wild. ex R. & S.)
) ;. m a c r o p h y l l a R . &
M . A r g . 7 3 ( 4 0 1 5 3 ) ; 7 4 ( , 1 8 3 1 1 1P
P. 73 (40065, 40072,40114): 74 (48165), P. racemosa
(Aubl.)Raeusch. 32 (48215); P. stenostachya Standl.
7 3 ( 4 0 0 6 6 ) ; P .v i r i d i s R . & P . l 8 ( 4 0 5 9 5 , 4 8 3 6 8 ) ; P . r v i l -
Schlegeliadresleri Gentry 26, 32 (48404) (Bignonizrceae).
Schleselia fastisiata Schery 26 (41069); 32 (4Blll); 48
( 4 0 1 2 7 ) ; S . s p . 4 l ( 4 0 3 2 9 )( B i e n o n i a c e a e ) .
Schoenobiblus panamensis Standl. in
(483 I 7) (Tl-rymelaeaceae).
L.O.
Sciadocephala schultze-rhonnotiae 52 (4l4Ul);
(11024) (Asteraceae).
Scoparia dulcis L. 44 (40567); 5l
( 4 1 4 3 I ) ( S c r o p h u l a r i a c e a e ).
Wrns. l8
S.sp. 53
(4002U); S. sp. 74
lianrsii Standl. 40, 41, 68, 69 (49044); P.spp. 4l
(a0322); 7l (40400); (40068, 40082, 40232); 73
( 4 0 2 1 3 ); 7 4 ( 4 1 0 5 5 )( R t r b i a c e a e ) .
Selaginella seniculata (Presl)Spring 58 (48131, 48207); S.
spp. 48 (40878);50 (48831);5l (40505);56 (40093,
P s y g m o r c h i s p u s i l l a ( L . ) D o d s o n & D r e s s l e ra 8 ( a 1 0 4 3 ) ( O r chiclaceae).
Pteris propineua Asardh 72 (48224); P. sp. 9 (40889) (Pte-
40186); 58 (49115); 73 (40200, 49042) (Selaginellaceae).
Senna dariensis B. & R. 35 (41087) (Oaesalpiniaceae).
ridaceae).
Qrrararibea cordata (H.B.K.)Vischer 24 (40580); Q. soegenii (Ouatr.)A. Robyns & S. Nielsson 24 (40773)
(Bombacaceae).
S i d a a c L r t aB u r m . f . 1 3 ( 4 0 0 3 6 ) ( M a l v a c e a e ) .
Sipanrna laurifolia (H.B.K.) DC. 73 (40643); S. spp. 40, 69
(40216);65 (48903);68 (49088);68, 69 (40693)Mo-
Raritebe palicoureoicles Wernh. 31 (48066) (Rubiaceae).
R a z i s e as p i c a t a O e r s t . 7 3 ( 4 0 0 i 1 3 ) ;R . s p . 6 3 ( 4 1 0 5 6 ) ( A c a n thaceae).
Renealmia cuatrecassii Maas 32 (41588); R. cylindrica
Sobralia rrracrophylla Rchb. f. 57 (48091); S. sp. 74 (48694)
(Orchidaceae).
Maas & Maas 73 (40833);74 (18235);R. ciolichocalp< Maas 36 (48924); R. oligospernra K. Schum. 48,
5 6 , 7 3 ( 4 0 1 6 0 ); 5 6 ( 4 0 7 7 2 ) ( Z i n e i b e r a c e i r e ) .
nirniaceae).
Socrateaexorrhiza (Mart.) H. A. \4/end1.2,24, 27 (60007)
(Arecaceae).
Solanum candidum l,indl. 24 (10165); 28 (40656); S. cirnense Rydb 24,41 (40016);41 (40721);S. conf.ertis e r i a t t r n rB i t t e r 3 1 ( 4 1 6 0 1 ) ; 7 l ( , 1 8 6 9 5 ) ; 7 3 ( 4 0 6 3 0 ) ;
R h e e c l i a e d r . r l i s( S e e m . ) P l . & . - 1 r . 2 4 ( 1 1 5 7 5 , 4 8 3 6 3 ) ; R . s p .
24 (40334, 41342) (Cltrsiaceae).
S. dolichorhachis Bitter 14 (40522); 5l (40437);73
( 4 0 6 9 1 ) ; S . . j a r n a i c e n s eM i l l e r 1 6 ( 4 0 4 6 9 ) ; S . l e p i d o -
Rollinia rnlrcosa flacq.)Baillon 24 (40608, 4B9Bg); 24, 56
(40046) (Annonaceae).
tum Dunal 72 (40306), 40536, 41067); S. leptorhachis Bitter 4l (40073); S. mamn)()surn L. 24, 33
(41452); 72 (404U7); S. medusocalyx Bitter 48
R u d g e a s p p . 2 ( 4 1 6 7 2 ) ; 5 8 , 7 3 ( 4 9 1 0 1) ( M e l i a c e a e ).
Ruellia pennellii I-eonard 42 (4U238);63 (48067);. R. riopalenquensis \Arassh.25 (40365); R. tubiflora H.B.K.
32, 7 4 (48402) (Acanthaceae) .
Ruta sp. 73 (40558) (Rtrtaceae).
R y t i c l o s t y l i sc a r t h a e i n e n s i s ( J a c q . ) K u n t z e 2 4 ( 4 8 0 5 1 ) ; 2 5
(48365) ; 29 (40462, 40607) (Cucurbitaceae).
S a b i c e av i l l o s a W i l l d . e x R . & P . 2 4 ( 4 1 6 2 3 ) ; 4 8 ( 4 0 1 8 a ) ; 5 5
( 4 1 5 0 6 )( R u b i a c e a e ) .
Saccoloma eleqans Katrlf. 40 (48223); B G0225) (Dennslaedtiaceae).
Salvia coccinea-f ass.ex Murr. 40 (40564) (Lamiaceae).
Sanchezia parviflora Leonard 4l (40398); 73 (40189); S.
sp. 73 (41464) (Acztnthaceae).
Sapirrm penn'ianum (Mcbr.)Jabl. 33 (48414,49098); S. sp.
2 , 3 3 ( 1 1 0 4 1 )( E t r p h o r b i a c e a e ) .
Satyria srandifirlia Hoer. 20 (48806) (Ericaceae).
Satrratria brachvbotr-vs Trrrcz. 24 (41621) (Actinicliaceae).
S a u v a s e s i ae r e c t a 1 , . l 6 ( 4 0 i r 8 9 )( O c h n a c e a e ) .
(41628); S. nudum Dunal 7l (41078, 48271):74
(4U304); S. quitoense Lam. 24 (41586); S. scl-rlectendalianum \&'alp. 58 (48233); S. sessiliflortrm Duna|24,28 (40509); S. spp. 24 (48829);34 (48817);
4 8 ( 4 1 6 4 8 ,4 8 4 0 6 ) ; 7 3 ( 1 0 1 5 4 )( S o l a n a c e a e ) .
Spartirrrn.junceuml. 23 (41525) (Fabaceae).
Sphvrosperrnum cordifoliurn Bcnth. 72 (40830); S. sp. 48
(48777) (Ericaceae).
S p i g e l i a a n t h e l m i a L . 4 8 ( 4 0 1 4 2 , 4 B 3 B B ) ;S . s p . 4 l ( 4 0 2 9 3 )
(Loganiaceae).
S p i l a n t h e s s p . 3 9 ( 4 1 4 8 5 ) ( A s t e r a c e a e ).
S p o r r d i a sr n o r n b i n L . . 2 4 ( 4 9 1 0 0 ) ; S . p u r p u r e a L . 2 4 ( 4 0 1 2 8 ,
4Ul 40) (Anacardiaceae) .
Stelissp. 49 (48247) (Orchidaceae).
Strcptochaeta sodiroana Hackel ll (40212, 40315, 40714,
4 1 0 7 9 , 6 0 I 0 B ) ( P o a c e a e ).
Stromanthe sp. (48634) (Marantaceae).
S t n r t h a n t h l r s s p . 7 1 ( 4 0 0 U 4 )( L o r a n t h a c e a e ) .
BS 46
S r v a r t z i as p p . 2 ( 4 1 6 0 7 , 4 1 t , 6 9 , 4 8 7 8 9 ) ( M i n r o s a c e z r e ) .
Sr,rrecanthrrs warscer,viczianus \\rencll. l5 (10205);
( 6 0 1l 7 ) ( A r e c a c e ; r e ) .
149
74
Svttgonitrm cr:rssifbliunr (Engl.)Croat 46 (,10330);S. nracrophyllunr Engl. 48 (40768) (Ar-aceae).
Tabernaemorlt.ana columbiensis (Alloree) Leerrrvenberg
24 (40351, 10426, 40732,40739, 48057,48206); T.
'L
heterophvlla \hhl 32 (4U123);
rnacrocallx Muell.
'f.
Arg. 5l (48933);
tetr:rstachvaH.B.K. 24 (0750,
40775); T. sp. 24 (40517) (Apoo,'nace:re).
T a s e t e s s p . 7 2 ( 4 0 4 8 U )( A s t c r a c e a e ) .
Talinrun panictrlatunr (facr1.)(]aertn. 29 (40559) (Porttrlacace:re).
T a l i s i a s p . 2 4 ( 4 0 U 2 0 )( S a p i n c l a c e a c ) .
Tapirira gtrianensis Aubl. 2 (18277) (An:rcarcliaceae).
TecLrria acutiloba (Hicron.)M:rxon 65 (40825); T. incisa
()av. 74 (4011014
, 0902); T. nicotianifcrli:r (Baker)C.
(llir. 35 (40411); 73 (a014,1, ,10680); T. uctrsa 72
( 4 8 3 2 2 ) ; T . s p p . I ( , 1 8 2 8 6 ) ;4 8 ( 4 8 9 4 7 ) ; 6 9 ( 4 i 1 , 1 3 5 )
(Drvoptericlace:re).
(40206, 40667); T. spp. 48 (41068, 48161); 56
( 4 8 1 0 4 ) ; 74 ( 1 1 1 2 5 ) ( P i p e r a c e a e )
Trichilia pallida Srv.73 (40006) (Meliaceae).
Trichipteris bipinnata Tryon 72 (48943) (Cvatheaceae).
Trichomanes coralliatum V. D. Bosch. Lg (40147);48, 4g
( 4 0 6 9 5 ) ; T . e l e e a n sR i c h . 3 5 ( 4 0 8 8 2 ) ; 4 8 ( 4 8 7 5 9 ) ; T .
rnernbranacelrm [.. 48, 5l (48345); 5l (40822); T.
polypoclioicles. 39 (a8839); T. spp. 48 (41630,
(40132, 4UU3B, 48tl92, 4U893); 73
48408); 5l
(400U0) (Hvrnenophyllaceae) .
Triofcna barbevana Cosn. 22 (48032);48 (40344, 41084);
49 (40094, 40157); T. obliqua (Triana)Wurdack 48
( 4 1 4 1 8 , 1 1 6 4 2 ) ; 5 0 ( 4 8 8 0 7 ) ; f . p u s r . u l a r aT r i a n a 4 9
(40071);T. sp. 48 (41431) (Melastomataceae).
Tripogzrndra serrulatzr (Vahl)Handlos
( 4 I 0 3 2 ) ( C o n r r n e l i n a c e a e ).
Tropaeolum
repandtun
40
(4U089); 4l
Heilb. 3? (48306) (Tropaeolace-
ae).
Urce olina grancliflora (Planch. & Lincliu.)Traub. 54
( 4 0 2 1 4 ) ; 6 6 ( . 1 0 3 3 5 ) ,4 0 8 5 3 ) ( A r n a r y ' l l i d z r c e a r e ) .
Tetratlrylacium rnacrophvlltrrn Pocpp. 2, 32 (48125) (Flacourtiaceere).
Thelypteris curta (Christ)Reecl 74 (48129); T. clentara
(Forsk.)St..fohn 72 (4841l);T. exutaA. R. Smith 44
( 4 8 8 0 , 1 ) ;T . f a l c a t a ( L . ) P r o c t o r 4 0 ( 4 8 2 9 8 ) ; T . f i a n c o -
L J r e n : rl o b a t a I - . 6 5 ( 4 0 0 4 1 ) ( M a l v a c e a e ) .
Urera baccifera (L.)Gaucl. 58, 59 (40669); U. caracasana
(facq.)Griseb. 22 (41508) (Urticaceae).
Vanilla odorata Presl 4l (4ll23ti); V. planifblia.|ackson 14
(40340) (Orchiclacezre).
i r n a ( F o u r n . ) R e e c l 4 8 ( , 1 8 2 9 4 ) ; 7 2 ( 4 0 5 7 0) ; T . e e r n mtrlifera (Hicron.) A. R. Srn.48 (48300);T. sisantera
(Mett.) R. M. Trvon 40 (40318, 40867, ,18069);48
( , 1 1 4 3 5 A ) , 4 l l F l i l 5 ) ;T . l i n e u l a t a ( ( i . C h r . ) C . V . M o r -
Verbenzr litoralis H.B.K. 42 (10401); 59 (a0208) (Verrbenac e a e) .
V i s r n i a o b t t r s : rS p r u c e 5 l ( 4 1 4 9 4 ) ; V . s p . 2 ( 4 8 8 3 2 ) ( C h r s i a ce:re).
\ A ' e t t i r r i aq u i n a r i a ( C o o k & D o v l e ) B u r r c r 2 ( 4 1 0 7 4 ) ; 2 , 2 4 ,
2 7 ( 6 0 1 1 2 ); 2 , 2 7 ( 6 0 0 0 5 ) ( A r e c a c e : r e ) .
ton 35 (48222);T. linkiana (O. Presl) R. NI. Tr.von 70
(48745);T. poiteana (Bor1,) Pnrcror I (48398);T. resinif'era (Desv.) Proctor 72 (48296); T. torresiana
(Claud.)Alston 4l (48130); T. ubanii (Socliro) A. R.
S r n . 7 3 ( 4 9 0 3 0 ) ; T s p p . 5 9 ( , 1 0 1 3 8 ) ;7 0 ( 4 1 5 9 4 ) ; 7 2
( 4 8 7 6 1, 1 8 7 6 2 , 4 8 9 r . r 2 ) ;7 3 ( 4 0 1 3 2 ) ( T h e l v p t e r i d a c e ae).
Theobroma bicokrr H. & F.24 (,104111T
) ;. c a c i r o L . 2 8
(40460, 40618);T. sileri (luatrec. 24 (4U,130)(SterctrIiace:rc).
Thevetia peruviana (Pers.)K. Schurn. 72 (10515) (Apocvnirceae).
T h i b a t r c l i a p a n i c u l : r t a A . C l .S r n i t h 7 2 ( 1 1 4 7 5 ) ( F l r i c a c e a e ) .
T i l l a r - r d s i an a r t h e c i o i c l e s P r e s l 4 8 ( 4 0 8 6 9 ) ( B r o r n e l i a c e a e ) .
"lbcoca
spaclicifkrra Triana 32 (4t3701); T. sl,nrphvandra
(Triana) Oogn. 32 (187 25) ( Mc l:rstonratzrceae
).
T o p o b e a c a u d : r t a \ { r r r c l : r c k 3 2 ( 1 8 7 1 ) 7 , . 1 U 9 1 8 ) ;T . s p . 3 2
(4890I ) (Melastomataccae).
.Ii'.
Tovomita r,veclclellianaPl. &
59 (4833ir); T'. sp. 55
( 4 14 5 3 ) ( C l t r s i a c e ; r e.)
T o v o n t i t o p s i ss p p . 2 4 ( 1 8 1 9 2 ) ; 3 l ( . 1 1 6 1 6 () O h r s i a c c a e ) .
T r e r n a i n t e g e r r i r n a 5 4 ( 4 t 3 3 0 9 )( U h n a c e r a e) .
Trianaeopipcr uarciae Trel. & Yunck. 73 (40227); T. killipii
Trel. 73 (.10755); T. nrcxiac T'rel. & Yrnck. 70
\AIitheringia solanacea l.'Her. 53 (a0376); 55 (40009,
4 0 1 1 0 , 4 1 0 5 3 , 4 8 1 l 5 ) ; 6 4 ( 4 0 3 0 8 ) ; 7 0 ( a 0 3 2 8 ) ;W . s p .
55 (48349) (Solanaceae).
\ 4 r t r l f 1 i ab a c c a t a ( L . f . ) K u n t z e 3 2 ( 4 8 1 6 7 ) ; 3 3 ( 4 8 8 5 1 ) ; 5 l
(48054) (Asteraceae).
Xanth<rscrrna
d : r s u c n s e E n g l . 3 2 ( 4 8 1 8 9 ); 4 8 ( 4 8 2 7 0 ) ; X . s a sittifolia (t..)Schott. 32 (48332);X. sp. 48 (48347)
( A r a c ea e ) .
Xipliicliurn caenrleum Aubl. 4l (40374);48 (48417)(Haemodoraceltt').
Zantia linclenii Resel 24 (48155); 5l (40203) (Cvcadaceac).
Z e a r n a y sL . 2 3 ( 1 0 5 2 5 ) ( P o a c e a e ) .
Zingibcr officinale Roscoe 39, 66 (491l6) (Zinuiberaceae).
Zvuia lorrgifcrlia (H. & B.) Br. & Rose. 74 (40403) (Mimosaceac ) .
Genus indet:
A c a n t h a c c a e 6 7 ( 4 1 4 8 2 ); 7 0 ( 4 1 4 5 9 ) .
A r n a r r . ' l l i d a c e a e1 6 ( 4 0 6 2 3 ) ; 3 9 , 7 0 ( 4 1 6 0 4 ) ; 4 8 ( 4 8 7 1
2).
Apocl'rraceae 24 (11041) .
Araceae 29 (40482);32 (4824il; aG (60101); 48 (4078e);
5 5 ( 4 r J 7 1l ) .
A s p l c n i a c e a el 3 ( 4 8 3 0 1 ) .
r
150
Asteraceae21,73 (40024); 25 (40103);40 (4057\; al
(40034A,40037, 40647,40707);48 (40702);Astera, 0 5 5 3 ,4 0 6 6 2 ) ;5 1 , 5 5 ( 4 0 1 0 4 ) 5
; 1,
c e a e5 l ( 4 0 1 6 1 4
; 2 (403964
, 0 8 6 3 ,4 8 1 5 2 ) ; 5 6 , 7 l
5 5 , 6 5 ( 4 0 1 5 2 )5
( 4 0 5 5 4 ) ;5 8 ( 4 8 4 4 3 ) ;6 5 ( 4 0 1 5 2 ) ; 7 2 ( 4 0 5 6 3 ) ; 7 3
(40024,40I 90, 40337).
Bromeliaceae
4l (48396);52 (40197);72 (400a3).
C l u s i a c e a2e ( 4 1 6 6 5 ) 2; 4 ( 6 0 1 0 0 )3; 2 ( 4 8 8 3 54, 8 9 6 2 ) ) .
Cornrnelinaceae
48 (48090).
Crrctrrbitaceae
25 (41579);59 (41071).
Dennstaedtiaceae
44 (48840).
Euphorbiaceae2 (41038).
F a b a c e a3e ( 4 8 0 6 3 ) ; 9( a 8 1 9 0 ) ; 4 8( 4 8 1 9 5 ) ; 5 1( a 8 0 5 8 ) ; 5 2
(4i13505
) ;3 ( a B 3 B 2 ) .
Flaccrr.rrtiac
eae24 (48169,48702);32 (4817| ) ; a1 (a1509).
()esnerizrceae
48 (41005).
Cleicheniaceae
48 (4f3950).
L a r r r i a c e a4el ( 4 0 8 5 4 ) ;4 1 , 6 5 ( 4 0 8 6 1 ) ,a 8 ( a l a l 6 ) ; 6 5
( 4 0 8 6 1 ) ; 7 a$ 8 1 a 2 ) .
BS 46
, 1 0 2 74, 1 6 6 1 ,1 1 6 7 04, 1 6 7 5 ) .
l . a u r a c e a 2e ( 4 1 0 0 8 1
Loranthaceae
4l (40634).
Melastornataceae
2 (416t0) .
Meliaceae2 (48229);6a (a8289).
Menispermaceae32 (48237).
Mvrtaceae32,74 (48401);73 (a9094).
Polypodiaceae; 25 (40431);aa Q1617).
R u b i a c e a el 5 ( 4 9 1 2 0 ) ;7 l ( 4 0 4 0 0 ) ; 7 1 , 7 3 ( a 0 6 8 5 ) ;7 2
(40292,41497).
Scrophulariaceae
4l (40300,40812,4102U,48148);
42, 58
(48444); a8 (a8?30,48863);64 (41461);7a ft1421,
41430).
Solanaceae
32 (48405); 48 (48210); 7a (a8l la).
Urticaceae34 (48050).
Verbenaceae
4l (41007).
Zingiberaceae74 (48694).
Family indet.
50 (403t7,40893,48l84) ;
151
BS 46
II
Annendix
r
Annotated list of vernacular names
--I
"k"
"c"
and
The letters
The spelling usecl is an approximation nsing sounds rnainlv fiom English.
oriSpanish
of
or
syllables
are pr.()nouncecl in the sanle way-,btrt the fbrmer is only ttsecl in names
"h";
,J"
otherwise it is pronouced as a
in Spanish names is pronouncecl like an Enslish
gin. The
"q"
,.cl.j,,inE,n.qlish.The use of
isiestrictecl to the Spanish names. An exclamation after avorvel '
""'
..!,,,
indicates that the vowel is
inclicates an:rbnrpt pronotrnciation of this. An elevatecl period,
"accent
..accent
ln a few Coaiquer nitused.
(d)
been
have
(e)
srave"
arld
i:gu"
long. Thc French
.,h"
clf the vernacular nadivision
A
svllabic
sotrncl.
aspirated
an
indicates
in parenthescs
nles an
rnes has been given tcl facilitirte comparison'
Cayapa
A ari
A len tape
Al bal kaka
Al ka kien chi re,'reoro rape
A ba.n ch. puka
A ba.n p.ka
A bo chli tape
A chi puka
A chide
A clju clju koro tape
A clo pistcha
A clrin
A f;r chi
A ka ma hal chi
A pe plnchi
A pi chua rape
A pi chui tape
A pisch chua
A pistch chu hal ki
A pistcha
collariatum (40147)
(,10769)
lugonis
Miconia
L . a r n i a c e a e( 4 8 1 4 2 )
(]urania sp. (48107)
Achr.rtele
Actrra tape
Clusia sp. (48234)
clusia sp. (4090tt)
Alloplecttrs panat-nensis (40846)
Talisia sp. (40820)
Bertiera procunrbens (40156)
Agan killan chi tape
Ai chi bu cria
Clttztnania scherzeriana (48395); (]'
Anis tape
sp. (41019)
Anotade
Taber'aenlorlrarzr retrasrachya
( 4 0 7 5 0 ) ( 4 8
Piperaceae (40295)
Al.sophila cuspiclata (411176)
Aji
Alia no pu fin dtrmi
Alui tape
Xanrhos.rna clastrense (48i89)
Etrphorbiaceac (40839)
(]trrania rnacrophvlla (40731); G.
spinulosa (40731)
Tectaria incisa (401-r0l)
c l u c u r b i t z r c e a e( 4 1 0 7 1 )
Clrrania spintrlosa (4s 170)
Clurania spintrlosa (48156)
A terlpo tap€
P e p e r o r n i a u r o c a r p a( 4 0 5 3 1 ) ; P . s p .
( 4 0 7 7 7 ), P . s p . ( 4 t J 3 l 4 )
Anrhuriunr r,ersicolor (48098)
6
4 )
Ar:i
;
P
.
Ari pi boka
Ari pi chi tape
Ari pi puka
As katcha renreclio
Ficus insipida (41080)
Razisea sp. (a105tr); Ruellia pen-
S
tape
As kin llullo
As kin llullo tape
Asa a ke uarin hodro
remedio tape
Asan shiili
Asoka tap6
Atchtt ban tape
p
nellii (48067); (a8238)
cavenclishia sp' (40754)
Ipomoea sp' (40476)
Tovomita weddelliana (48335)
Drvmonia alloplect.ides (40061)
Asteraceae (40574)
E'laphoglossttm herminieri
(40533);E'sp' (40826);E'sp'
(48301)
Atchtr boa tape
Atrir zrtnbi chi
A va,vatape
Abanch.la fi'chtrno
Clodonanthe crassifolia (40179)
Ba ta ba ha! ki
Ba vero tape
Polysonum hvclropiperoicles
(40494)
Bixa orellana (40045)
Ban chi
Achiote
Piper brachypodon (40BBl); P' sp'
' ( 4 8 3 9 2 )
Hemidictyum marginaturn (4013,1)
Ficus maxima (4u056)
Ficus i'sipida (40900)
Cl.eonoma cuneara (48272);
(4it35l)
Anthttrittnr aspltrnclii (40886)
bustr
tap€
Pavonia fruticosa (40555)
Salvia coccinea (40564)
Piper marsinatum (40012)
Anpo pin ha! tap6
1
Terracera sp. (408g2)
Anthuritttn pallicliflorum (48106);
Asplenium serratturl (48258)
Abill.
(40017)
Adiantum petiolatum (4057I);
Thelypteris fiancoana (40570)
(lleome sp. (40593)
Farnilv-indet. (41004)
Capsicum frutescens (40201)
ralinum panictrlaturn (40559)
Aga remedio tap6
A pistchtr ptrka
A tchu ban tape
A vikinr. tchape
A y,ahal chi
Bixa orellana (40474)
Discophora gttianensis (40305)
Podandrog,vne brachvcarpa
Achiote enzrno
'frichomales
Ba ki tape
Ba pist chi
Babarro
Babo kasta(h)
Polypodiurn loricilbrme (40795)
Heliconia longa (48180)
Guadua angustifblia (48374)
Physalis angulata (4U385)
I'iper sp' (48158)
Streptochaeta sodiroana (41079)
\Alettinia qtrinaria (41074)
Phytolacca rivinoides (40003)
Scoparia clulcis (4002u)
BS 46
752
P h y t o l a c c a r i v i n o i d e s ( 4 8 1 0 1)
Balanclo remedio tap€ Plumeria rubra (40594)
Psychotria viridis (40595)
Ballusa tap6
Bakalao tape
Ceclro
Cha dja pollo llullu
Cha shi ta tape
Bandi tururu chua
Passiflora quadrangularis (481 87) ;
Chachi chi ha! ki
Chalviande
Barbasco
Basu remedio tape
&8429\
Phvllanthus anisolobus (40010)
Tectaria mexicana (48262); Thelypteris sp. (48298); Tripogandra
serrulata (41032)
Che ba tape
Basu sji mul ko
Begonia sp. (48421)
Basu tap6
Campyloneurum phyllitidis
(401106); Thelypteris gigantea
(40867); (a8069)
Chanilla piechiga
Chanul
Chi chi bilia
Meliaceae (48229)
Amaranthus sp. (48391)
Paspalum sp. (41015)
Poulsenia armata (48343)
Dialyanthera sp. (48290)
Alonsoa sp. (40560)
Humiriastrum procerurn (4 1076)
C o r d i a s p i n e s c e n s( 4 1 0 6 2 )
Rollinia mucosa (4060ti)
finchuno puka
Chi chua ama
Chi dja pdpd
Asplundia sp. (40829)
Tectaria brauniana (4B2BO)
kino tape
Hvptis verticillata (41 083)
Pteris sp. (40889)
Chi ha pdpd
kino tape
Acliantum sp. (a082tt)
Urticaceae (48050)
Adiantum isthmicum (48261 ) ;
A. macrophyllum (48399) ; Thelypteris poiteana (48398)
Chi ka shu
Musci spp. (40903); (41057);
(481l0)
Bi ta chua
Carpotroche platyptera (40873)
Desmoncus sp. (48197)
Chi ka shu tap6
Bi vino tap6
Scrophulariaceae (48444)
Antrophl'um
(40842)
Billo tape alato
Polygonum hydropiperoides
( 4 0 4 e 1)
Chi ki tutu
Trichomanes membranaceum
(40822)
Pithecellobium loneifolium
(48288)
Chi nu tchumo
Scrophulariaceae (48230)
a bakaka
Chi nu tchumo
Philodendron sp. (401105)
Bi bina tape
Bi ke pan tsumi tap6
Bi ken sumo tape
Bi kini
Bi pistcha puka tape
chuno tape
Bish chi tap6
Bish kino hoke
Piper phytolaccaefolium (40468)
kuneu tap6
pinchori tape
Bish pe remedio tap6 Cyathula achyranthoides (40499) ;
Microsramma piloselloides
(40475); Piper cararense (al021);
Bitchi puka
Bo chua llullu
intramarginale
(48174): G82a9; P.sp. (40782)
Dendrcrpanax sp. (407 42)
Marsdenia sp. (a8122)
Chi ta may chimo
Araceae (48246)
kungo papa
Chiaimayo
Chiar llangia tap6
Chicha tap6
Rollinia mucosa (40046)
Bomarea edulis (41039)
Araceae (40482)
Chichi ha
Clhide ari
Anthurium sp. (40136)
Asplenium hallii (a0148)
Chikaso tap6
Selaginellasp. (40505); rnusci
Chilma
(40573)
Dioscorea
sp.(a8331)
Bo shi chi
Hyospathe elegans (48273)
h'rga edulis (48200)
Bolon ge djo djo
Fungi (41073)
Bonban
Borachera
Oryctanthus sp. (40002)
Bnrgmansia versicolor (41 050)
Chimba chua
Bomarea edulis (40U14);Dichori-
Botonsillo
Bovi n chi
Asteraceae (40554)
Iriartea deltoidea (48409)
Chimi tchumo
sandra hexandra (40824)
Philodendron sp. (40730)
Bu ma chi
Coussapoa rotundata (40757)
Razisea spicata (40083)
Chimpalo
Solanum canense (40016)
Chin lak pe
R y t i d o s r y il s c a r t h a g i n e n s e
( 4 8 0 5I )
Chin remeclio tape
(lhinba tchuba tape
Chino tchumo
Oxalis sp. (a8333)
Bo chui kano chi
Budu shinu
Bui tape
Bulla tape
Buru sino
Buru sino tape
Buru tape
Cafe chi kushorol
Polvgonum acuminatum (4841 2)
Aegiphila alba (40538); G8427)
Casteranthus oncosastrus (40233)
Sanchezia parvifolia (401 89)
Spieelia anthelmia (40142)
Cofl'ea arabica (4017 0)
kungu ha!ki
ko dju dju
Chino tchumo
Capitan chua tape
Guadua angustifbli a (4837 4)
Mendoncia sp. (40813)
Capitan shuba miro
Aristolochia sp. (40523)
Chirma
Cafra
gulachi
Carbonero
Oarlen tap6
Anthurium
sp. (a0799)
kungu hoki
Chiyacu tap6
Cucurbitaceae (48305)
Selaginellasp. (40093); S. sp.
(40186)
Chu ai n chi
Dialyanthera sp. (48290)
Fabaceae (48063)
Desmodium axillare ( 40022\
Solanum dolichorhachis (40522)
Anthurium sp. (48074)
BS46
Chu chu maseranu
153
P r e s t o n i a p o r t o b e l l e n s i s ( 4 0 8 3 1)
Fischeria aequirtorialis ( 48334)
Passiflora rnalifbrmis (40539)
Dtr chi chirno puka
Brugmansia versicolor (41050)
Eugenia sp. (40044)
Bixa orellana (40474)
I)u ma pe tap6
Du sirrpi
Selaginella sp. (48254)
Selaginellasp. (40200)
Odontaclenia macrantha (40895)
Rytidostvlis carthasinense (48365)
Fabaceae (4t305tt)
Fabaceae(48382)
E ki cljo chua
E lon be po tape
Paragonia pvramidata (48413)
Abehnoschrrsmoschatus (48150)
Pavonia castaneilblia (48204)
tap€
Chu mas6 geno chua
Chu roro
Dorkila
finchuno puka
Chu! maseranu tape
Chuan himu tchape
Chun djui ha!
Chun djui hal tape
Col de monte
C z r r i c ah i s p i c l u r n ( 4 0 1 9 5 ) ; C . s p .
(40088);C. sp. (401:rB)
Concha mula
Hyptis obtusif'lora (40 I B0)
Brtrnfelsia srandiflora (4002I )
Acliantum rnacrophylltrrn (401 I I )
Sida acuta (40036)
Culi tape
Cunta ha sa pihan
De ui
De sa remedio tape
Dedo
Tcctaria draconopterri (4tt435)
M a t i s i a c o l o r a d o r t r m ( 4 0 0 1 5 );
( 4 0 1 3 1 ) ;( 4 0 1 8 7 )
Dds tape
S e l a g i n e l l a a n c e p s ( 4 8 1 3 1) ; S . g e n i culata (48207)
Di ki chu ama
A s p l u n d i a s p . ( 4 i J l 7 1 1;) P h i l o c l e n c l r o n s p . ( 4 0 f 1 4 9 ) ;P . s p . ( 4 8 4 0 3 )
Dio tape fin chuncr
Dishan tap6 luli
f)apsicum frutescens (405ti6)
H i b i s c u s r o s a - s i n e n s i s( 4 0 1 3 9 )
(llusia sp. (48242); Phthimsa pvri-
Dja hin chi
remedio tape
Dja pd chi
remedio tape
Dja pe mayteno tape
Dja pdi n chi
remedio tap6
Dja pdno
Donga hal ki
folia
(48151)
Notvlia rimbachii (48143)
EI ta ke
E ke ua tape
Elnrrsano tape
Embacha chuno tape Polygala paniculata (40590)
Sida acuta (40036)
Escaba
Esir kva tape
Eskito tape
Fe ko killan chi
remedio tape
Dja pin sa chi
Djeiba chi
Dju dju koro tape
Djuin chi
Djuin tap6 chi
Do mai n chi
Farnily indet. (41085)
Fiban penpo
Firrra pin
Asteraceae(40161)
Stelis sp. (48247)
Manihot esculenta (48329)
Fiba bar6 himu yambo Dioscorea sp. (48331)
Fiba hal ki
Calathea lutea (4B1Bl)
Fiba ljuin chi tape
Clostuslima (48093)
remedio tape
Firnu chui n tape
Firnu pin
rernedio tap6
Rutaceae (48199)
Alloplectus panamensis (4i3120);
Araceae (407U9); Columnea bilabiata (40816); C. densibracteata
(48084); C. dissimilis (4Bl 78) ; C.
picta (48086) ; C. purpurimarginata
A n t h u r i u m s c a n c l e n s( 4 1 0 4 0 )
Anthurium scandens (40601 ) ; Fab a c e a e ( 4 8 3 5 0 ); P l e u r o t h a l l i s p i c t a
(48432)
(4U085);C. nrbriactrta (4B0BB);
( 48 12U) ; Dichorisandr:r hexandra
Phoradenclron piperoicles (48367)
( 4 f 3 18 8 ) ; D i p l a z i u m s a n d e r i
(48263);D. seemannii (48255);
Prestoeadecurrens (48354)
C o s t u s s c a b er ( 4 8 1 4 6 )
D r y m o n i a w a r s z e w i c z i a n a( 4 8 1 2 1) ;
Gasteranthus corallinus (48094) ;
l,atrraceae (4100U)
Bromeliaceae (48396)
Latrraceae (41027)
Monopyle sodiroana (aBl27) ;
Monstera sprlrceana (4ti105); Peperonria macrostachya (48202)
Finrrr pin tap6
Palicourea guianensis (41003)
I s e r t i a p i t t i e r i ( 1 0 8 2 7 ); ( 4 8 2 5 1 );
Do pish chi
Tabernaem()n tana te trastachva
(40775)
Do pistcha
Tabernaemor.lt:rna sp. (4015I 7)
bamo puka
( 4 8 3 7 6 ) ; P e p e r o m i a s p . ( 4 8 1 9 3 ); P .
sp. (48369); Xanthosoma daguense
(48270)
Fimu pin tap6 ku
Do pistcha chi
Tabernaemontana columbiensis
Do pistcha
Tzrl>ernaemon
tana columbiensis
(40732); (40739); (41014);
(48057)
Tectarianicotianilblia(40144)
(4820ri)
Columnea spathulata (4$a4-)
D r y m o n i a e c u a d o r e n s i s ( 4 8 3 11 ) ;
D. serrulata (48393); Fabaceae
(48 I 95 ) ; Macleania rotundifblia
(48278)
Do tape
Trema integerrirna (48309)
Tripouandra serrulata (48089)
Portr.rlacasp. (a8379)
Fe de pin tap6
Fi ba n go cho
mayteno tape
Dja rin chi
remedio kola
Cordia spinescens (48052)
Crinum amabile (40496); (40547)
mun chi bulo
chora asa - delan
Columnea rtrbribracteata (4it087);
Drynionia coriacea (48092)
mala pike ktrshnu
taPe
Finrtr pini chi pi
peno tape
Xiphidium
caeruletrm (48417)
154
BS 46
Fo horo chi
remedio tape
Erythrodes sp. (407tt8)
Halki tala tchumo
llullu tap6
Fu chun chili
S a b i c e av i l l o s a ( 4 0 1 8 4 )
Solanum sp. (4ti406) ; Stigmatopte-
Halki tape
Fumu pin
remeclio tape
Furnu pin tape bai
mo mo mu ili ke
mayteno tape
Furnrr pin tape
ris sp. (48300)
.fusticia ianthina (48384)
P i l e a s p . ( 4 0 0 6 2 ) ;P . s p . ( 4 0 1 0 7 )
Bromeliaceae (40197)
Halina dja karno tape Pharus latifblius (48096)
Apeiba membranacea (4tl I 98)
Han ape chi
Hanben tchuba tap6 D a l e c h a m p i a s p . ( 4 0 5 2 9 )
Habo chi wila
He gugu chi tape
Fumu pu tape
Spigelia anthelmia (4B3BB)
Columnea kienastiana (48383)
He kin llullu ptrka
Fumu pu tape sautal
Gloxinia dodsonii (48279)
He mu tui n chi
pepa illi ish karapa
- to btrshno tape
Fumu tchumo po
pin ha! tape
Furi firrica
Ga k6 tala tchumo
Solanaceae(48240)
Jaltomata procumbens (4001 3) ;
(40101)
He kin llullu
Capi tape
Bertiera procumbens (40834)
Casearia sp. (408tr4)
He tape rnincha
Miconia trinervia (40489)
rokola tap6
H€ tape mincha
rokola hubi
kichono tape
H6 uai sa tap6
Pseuderanthemtrm leptclrhachis
( 4 0 8 2 1)
Mimosa elliptica (40483)
Coix lacrymajobi (40540)
He uayusa tape
. fu s t i c i a s p . ( 4 8 3 1 3 )
Lomariopsis nip;ropaleata (4ti284) ;
Tectaria acutiloba (40825)
Polypodium fraxinifblium (40798)
Cephaelis sp. (41047); Psychotria
A l p i n i a p u r p u r a t a ( 4 0 5 5 1)
.facaratia spinosa (407U5)
Psychotria caemlea (40740)
G6 mu tu ha
fionzalagunia cornifolia
(4 0 7 8)r
Ge sapote
()6 shi bish tchape
Gola chi
Grama
Granadilla
Qtrararibea soegenii (407 7 3)
Burmeistera sp. (40771)
Dimerocostus strobilaceus (40583)
Cyperus luzulae (40035); Panicum
frondescens (40182)
Passiflora quadrangularis (40 I 74)
He yai chua
He yai remedio tape
Hd bo chui tape
macrophylla (a8165)
He llullu bo chtri tape Ruellia tubiflora (48402)
He ne chinch
Solanaceae (4ti405)
He shi vesh tchape
He yai chua
He a la pi chi
Bunneistera sp. (48102)
Polybotrya caudata (48257 )
Ficus rnaxima (48203)
H6 ambi chi
Heliconia stricta (4827 4)
Danaea moritziana (48282)
Ceogenanthus rhizanthus (48068)
He ari tchape
Gualte
\Alettinia oxycarpa (60000) ;
(48355)
He eski tchu
Guava
Irrga edulis (40042)
Eugenia sp. (,10044)
He ko bin pistcha
Guayaba
Guayacan chi
G r r a y a c a nc h i h a l k i
Guayacin
Cual.usa tape
Gudo chaino puka
Guin tape
Guto killan chi
Minquartia guianensis (41023)
Minquartia guianensis (48400)
Minquartia gtrianensis (4U400)
Psychotria viridis (4836U)
Bixa orellana (40602)
He sanjuanilla
Begonia sp. (48310)
Plantaso major (40599)
He n bo chui tap6
He n chi chua ama
Ha na ko lush tape
Salpichlaena volubilis (48060)
Perebea xanthochyrna (4820 1 )
Cestrum sp. (48166)
Ha peno
Anthuriurn trinerve (40808)
mayteno tape
Ha pistchi
Ha sa shi pistcha
Ha sa ti pistcha
Ha sa ti pistcha puka
tchu tape
H6 yai renredio chua
Hd kin llullu chua
He kongo chi
Hd pistchu tape
remeclio tape
Ha kino chua
Ha mu ki chi
Miconia erioclada (40569)
Scoparia sp. (48366)
Ge ba pai chi
C]6 bo chui tap6
Gasan funilla
Monstera adansonii (40493)
Drymonia coriacea (40819)
D r y m o n i a m a c r o p h y l l a ( 4 0 5 2 1)
He mush tu ha tapc
llullu tape
Gacho dar tap6
Ludwigia octor,'alvis(40858)
Pourourna hirsutipetiolata (41 009)
Ardisia sp. (a0817)
Flacourtiaceae (48 169)
Farnily indet. (41010)
Dicranopteris pectinata (48323)
Lomariopsis.japurensis (48 I 25 )
Piper sp. (48338)
Xzrnthosoma sp. (48347)
Sabicea villosa (48280)
nokososo tape
He n dji chu chua
He n clo pistcha
Pharus latifolius (60106)
Psvchotria caerulea (48172)
Asplundia sp. (aBl24)
Menispermaceae (48237 )
Tabernaenlontar)a heterophvlla
(48123)
He n modiulla tape
He n pash ba! ke
l.ozania kltrgii (4051 I )
.fusticia cornata (48144)
He n tala o pin bisj
Trichomanes ankersii (48285)
chi kera tchumo
tape
He n tape
Hekin clio tchuba
Miconia barbinerr,is (4ti43 I )
Malpighizrceae (40524)
BS 46
I{elen anpura
Helen ara
Helen bonba
Helen clini
Helen ehivilzr
Helen na sunbtr
Helen paki
Helen p:ipui
Helen pula
Helen uinca
Hen pistcha tape
Hen ballosa tapi:
T'rianaeopiper. mexiae (40206)
l)ipl:rziurn srriarunr (400 I g)
Stnrthanrhus sp. (40084)
f'ilea sp. (40008)
Aechrnea ansustifi)lia (40043)
Epridenclrtrrn cliffirrrne (40 10Z)
Pepcrornia rrrocafpa (40135)
Peperomia urocarpa (40067)
(iayaponia coriacea (40078)
Ps1'chotria rnacrophylla ( 40065)
Clurania spin trlosa (40534)
. f r r s t i c i as p . ( 4 0 : i 3 5 )
pincleno punchtrmi
Hen bo chui tape
P z r l i c o t r r e ac o n f ' e r r a ( 4 0 5 1 8 ) ; R a r i tebe axillare (4105b)
Hen bu chi tape
Podanrlroel'n e brer,ipeclrrnctrlata
(40.366)
I{en chilla tape
Tillanclsiasp. (40869)
Hin ki cliu rchub:r
E c t o z o r n : rp a r , o n i i ( 4 0 5 I 5 )
Hen kin llullu tapc
D r v m o n i a c o r i i r c c a r( 4 0 5 I 4 )
Hen pi chun puka
Cavaponia sp. (405I2)
Hen tape chi
Hen tape
Hdn tape rniro
sulachi
Hieve estrellas
Hin bo chui tapi:
Ho chi
Ho da lia side
Ho horo pin tape
(lliclcrnia septuplinen'ia (4051
0) ;
O s s a e al a x i v e n u l a ( 4 10 l 3 )
Miconia gracilis (41033);M. nervo_
sa (4101.1)
C l o n o s t e s i ac e n t r o n i o i r l e s ( 4 0 5 2 8 )
()uphea srrisukrsa (40031
)
P s v c l t orI i l r l r o f [ i n a nrr s e c g i a n a
( 4 8 3 15 )
Brosirnum rrtile (41037)
(lcstrtrm meualophvlhrrn (40007)
Scaphl'glouisprolifera (48I96) ;
'Iric
hornzrnes clir,'ersifi-ons
A l l o p l e c t u s s p n r c e i ( 4 1 0 8 1) ;
(.1U0U2); Triolena barbcvana
(41084)
Acalvpha sp. (413182)
S p o n c l i a ss p . ( 4 8 1 4 0 )
Hohoro remeclio tap6 Besleri:r barclayi (4841 I-r)
Hopi uallu tape
r\calvpha sp. (41030)
Hungu ba chua
Philoclenclron \rernlcosurtr (4g I g 1
)
Hupi rrirllu tzrpe
Acalypha dir,ersifblia (,10735)
Ish ba vi urpe
M e l i a c e r a e( 4 U 2 ? 9 )
Philoclendron rhockraxis (4824I
)
A s p l u n r l i a s p . ( , 1 8 3 7 1)
O s s a c am i c r a n r h : r ( 4 0 0 4 8 )
(blunrnea kienasriana (48162);
Cl.
pis biserrata (48259)
l , o n c h i t i s h i r s t r t a ( , 1 0I 3 0 )
ba n tape
Ka atchu ba n tape
Ih clju dju koro rape
I L ag o l a n c h i t a p e
Ib hin tape
Anthuriurn sp. (a8342);pecluma
consimile(48430)
Guzmania linetrlata (a8377); Til_
landsia monodelpha (a8253)
C o s r u sp u l v e r u l e n t u s ( 4 l 0 l Z )
Conostesia centronioides (40770):
(4077e)
Ka llornr ll.llo ch.a
Fabaceae(48rg0); pa'llinia firscescens (48426)
Ka rna ka
I{a ma pe tap6
Ka pala tape
Ik patash pa hiilki
Xanthosoma sagittifolia ( 4g332)
A l t e r n a n t h e r a s e s s i l i s( 4 0 9 9 0 )
Anthurium rrisecrum (48249)
Piper ansusrunr (40719); p. sp.
(407 90) ; Trianaeopiper killipii
(40755)
Ko pii chin sa
Tonin:r fl uviatilis (40796)
Kit pu chin puka
Arclisia romeroi (40906)
K:r pu chin puka tape Arclisia romeroi (40b42); (48325)
Ka shi ta
Fimbristvlis miliacea (48358)
rernedio tape
Kir shi ta tape
I{a shui bo chua
Paspalum conjuuattrrn (48I47)
P a r - r l l i n i an o b i l i s ( 4 U 3 4 1)
Ka! ztr tape
Ka chirn ba chuzr
Ka lechuka tape
Ka tdnrpo tap6
um cajenense (40tt40)
Fimbristylis dichoroma (48353)
;
Kl,llinga purnila (48272) ; Rh,vncho_
spora r:rdicans (48373)
P h y l l a n t h u s s p . ( 4 8 2 3 1)
Vanilla odorara (48236)
Heteranrhera reniformis (4f3390)
Peperornizr urocarpa (,1l0l g) ; p. sp.
(a078a);
s p a t h r r l : r r a( 4 0 4 8 1) ; H r r p c r z i z rl i n i I i r l i a ( 4 0 t i 1 0 ) ; ( a 8 2 9 2 ) ;N e p h r . o t e Ishan ara
fabonsillo
l(a aiioiio archrr
Ka rrarn bio chi
Ho pi uallu
Hocos ptrka
nun chi
pi di ki chua anra
pi cli ki drra anra
Iszrn tape
Ishki djo djo
Itaki tape
Iyu kinto
Peperomiasp. (40143)
Peperomiapreroneura(a00g9);p.
sp. (40137);Psychotriasp. (40068);
P. sp. (40213);P. sp. (40222)
Fungi (41072)
Desmodiurnadscendens(40532)
Aristolochiapilosa (1022b); Sola_
nurn sp. (40154)
Phytolaccarivinoides (40003)
Oleandralehmar-rnii
(48303)
I(a tiu tiu kuru tap€
C a t o p s i s s e s s i l i f l o r a( 4 0 g 0 1 1 )
Ka tiu tiu tape
( i a t o p s i s s e s s i l i f k r r a( 4 0 9 0 1
)
I(r rirrktirr krrru tape Anthurium sp. (40759);
Antrophv_
(4rJ260)
Ho honr Lrpe
Ishan sunba
Ishan tape
Ka yrrn tapc
h . a r il o < l i o k a r d o r a l r c
Mimosa pudica (48324)
Clrrzmania linsulirra (40b72)
Kafir murzr
I{alabos tape
I(an sd tape
Karan tape
llerar
Streptochaerasocliroana (402I 2)
Hr.'ptis capitata (48356)
Katsir nroca
Piper sp. (48l 117)
D e s m o d i u r n a d s r : e n d c n s( 4 0 0 3 2 )
Anthtrrium falcarum (40204)
Urceolina grancliflora (40211)
BS 46
156
Ke ba tape
Ke pi puka
Kd yui chua
Kelan boe chi
Kene dana de monte
Kere na
Kere pi puka
Ki pe pu chi tape
Kimbi llullu
C o r d i a s p i n e s c e n s( 4 0 4 9 5 ) ; C a s s i a
reticulata (40568)
Chimarrhis sp. (401I2)
Kuta ma chi
Alloplectus sprtrcei $07 6a) ;
(10844)
Kuta manchi
Polybotr,va caudata (41 058)
Cupania cinerea (41046)
La pa
Columnea nrbriacuta (40872)
Panicum frondescens (40182)
Bixa orellana (40473)
Peperomia urocarpa (40067)
Hofl'mannia sp. (40151); Psychotria sp. (40082); Rubiaceae (40066)
Lama pu tchuba tape Solanum jamaicense (40469)
Cyperus luzulae (40035)
Lapa
Solanum sessiliflorum (40509)
Lara.jilla tap6
Psychotria hoffmannseggiana
(40153)
Lash muchi
Pachira aquatica (40498)
Palicourea guianensis (48205)
kushnu chi
Kimide
Kin llu chua
Kin llui puka
Kin llullo
C e s t r u m r a c e m o s u n r( 4 0 1 7 6 ) :
Hamelia macrantha (40076)
Schlegelia dresleri (48104)
Strychnos sp. (40883)
Psammisiasp. (40520)
finchuno puka
Kin llullu
Psammisiasp. (a0309); Schlegelia
fastigiata (41069)
Kin llullu tape
Blakea subconnata (40197); Drymonia alloplectoides (40845)
Amphiclasya sp. (48248); Psammisia attaberrans (48250)
Kin lui chua
Lama fin tchumo ma
Leche brava
Lesa tape
Limon shilli
Linum sillo tap6
I,juin tap6
Bixa platycarpa (40810)
Ficus caldasiana (40096); F. obtusifolia (40047)
Senna dariensis (40543)
Ipomoea sp. (40005)
Hvptis capitata (40544)
Begonia sernio\rata (41029) ;
(48064); (a8276); Monopyle sodi-
Llu chua auca
roana A0877)
Ludovia integrifolia (48281)
Geonoma linearis (60115)
l.lullpo pi chui tap6
Dennstaedtia bipinnata (4tt256)
Llupu dds tape
Llupu pi shungui tap6 Columnea picta (40871)
Sapium peruvianum (48414)
l,lupu pin sa chi
Llupu ua ko tape
Piper sp. (48346); P. filistylum
(481l6)
Llyoko pistehi
Theobroma gileri (48430)
Phyllanthus anisolobus (48370)
Blakea subconnata (48336)
Drymonia serrulata (40603)
Lon bo tape
Kinbi susno tape
Kindo
Kine tap6
Hamelia macrantha (40503)
Peperomia sp. (40169)
Lu ocoshili ayan
Luban buli
Urticaceae (40478)
Ltrban koko lan
Kinli cosno tape
Drymonia coriacea (40550)
Blakea subconnata (40049)
Luban luli
Lycianthes amatitlanensis (401 72) ;
Witheringia solanacea (40009) ;
(40110)
Luki mona
Lulu tape
Mirabilis jalapa (40025)
Cuphea strisulosa (40546)
Pseudelephantopus puratis
(48381)
Couania sp. (41086)
Hypolepis hostilis (40527)
Lr.rn tap6 chi
Ma kari chi
Allamanda cathartica (40504)
Pentagonia sp. (41035)
Ma pini ari
Ma sju ke mo chi
Asplenium purpurascens (402 I 5)
Flacourtiaceae (48l71 )
T r i a n a e o p i p e r s p . ( 4 8 1 6 1)
Hypolepis hostilis (4i070); H. repens (48048); (48299)
Chrysophyllum arsenteum (4101 7)
Malanga
Malva tape
Xanthosoma sagittifolia (48332)
C,vphornandra hartwegii (4019 I )
Malachra ruderalis (40860)
Diastema affine (40140)
Lycianthes synanthera (401 1 3)
Mama.juana
Asteraceae (40037)
Man cebolla tape
Piper imperiale (40091 )
Piper hispidum (40194)
Man ni ka chi
Urceolina srandiflora (40853)
Tetrathylaciurn macrophyllum
(48425)
Piper marginatum (40012)
sp. (48095)
Anthurium caulorrhizum (48243) ;
Man samballo tape
Kin mullu chua
Kinbi finchuno tape
Kirdo
Kiri tape
Ko cha ni ka tape
Ko chua
Ko pin pistcha
Ko piua do chi
Kono tape
Kota tap6
Kototo anpo ha
Kototo anpo
Kronocio
Ku dju dju
Ku d.ju dju tape
Luban pinion
Luli chide
Ma pe remedio tape
finchuno
Ko pin pistcha
Lu ano
Anthurium
Malu side
Mana tchape
Gossypium barbadense (40584)
Trichomanes sp. (40086)
Malvaviscus penduliflorus (40027)
Euphorbia cotinifolia (4017 7)
Solanum lepiclotum (40306)
S c i a d o c e p h a l a s p . ( 4 10 2 4 )
Humiriastrum procerum (41076);
I-antana sp. (41077) ; \A'ulffia baccata (48167)
A.'p. (48239)
Kuehua chi
Kuru paja
Heliconia curtispatha (401l9)
trinerve (40231)
Odontonema strictum (40023)
(lostus pulverulentus (401 I 6)
Anthurium
Manga bullcr
Mansamollo
Asteraceae (48152)
Asteraceae (40863)
BS 46
r57
Mape remedio tape
Maracuya
Aciotis sp. (41016)
Passiflora edulis (40026)
Na tsetsero
Chamaedorea pinnatifrons
Mashu kino chi
Mashu kino tape
Miconia oraria (48339)
Psychotria cooperi (40862)
Na ua kinto
(401e8)
Passiflora
(40174)
quadrangularis
Na hino
Sobralia macrophylla (48091 )
Mata palo chi
Mata palo
Ficus schippii $Ba2a)
Loranthaceae (40056) ; Oryctanthus occidentalis (40058)
Araceae (60i01)
mayteno tap6
Nan bo! chi
Mavo tape
DiefTenbachia seguine Q07 56) ;
(48154); (a8348)
Naines
Memo kinto
Melothria pendula (40098)
Mihano tape
S c h o e n o b i b l r r sp a n a m e n s i s
(483I 7)
Mil pesos
Mincha rro sala
pusis no tape
Monstera spruceana (40i143)
Hamelia axillaris (40484)
Mincha rro kola
Psychotria brachiata (48l 12)
Mayo
ban vino tape
Mincha rro kola
hambi kino tape
Mincha rro kola
man da ham biken homo tape
Moli
Moragia
Morin
Mu
Mudo tape
Mulu fin tchtrrno
Kalanchoe sp. (48149)
Ruta sp. (40558)
tchuba tape
Mutondo puka
Mutu mu tape
Na ari
Na bonban
Na disku
Ne chin chi
Ne chin puka
Ne chin puka tap6
Nelo ha
Randia sp. (a8119)
Siparuna eggersii (40BlB)
Meliosma sp. (40526)
Anthurium sp. (40150)
Solanum lepidotum (aI067); S.
schlechtendalianum (48233)
Fabaceae (48063)
Clusiaceae(60100)
Cuphea tetrapetala (40552)
kino tap6
_
Nillo tape tchumo
Nincha rosula bale
pu mangan chino
tape
O kera tchumo kea
ka tap6
Poaceae (40587)
Clolumnea kienastiana (40605)
Coleus x hybriclus (4U357)
Dialvanthera sp. (41049)
Fictrs caldasiana (40096)
Hyptis obtusiflora (40294)
O kera tchtrrno llullu
O kera uaku tape
Polemonium sp. (48364)
Callisia repens (48394)
OI dja ha! chi
Begonia semio\,ata (40298)
Bixa orellana (40045)
O kera llullu tap6
O kera tchurnu
Calathea inocephala (48252)
Portulaca sp. (48378)
Portulaca sp. (4tt380)
Aneilema umbrosum (40033) ; Hydrocotyle leucocephala (40193)
Phaseolus vulgaris (405 I 6)
llullu tape
O n gala hen tap6
Oa chambi tape
Oa lan tap6
CUSSAS
Mulu kera pi
Hyeronima laxiflora (40507)
Tovomitopsis sp. (40823)
Cucurbitaceae (40891)
Ni tsala kino chi
Niane boka
Nicha roca la habi
C l i t r r - rs p . ( a B l 9 4 )
\\'ulffia baccata (48054)
remedio tape
Missokillan chi tape
Mo chi
Naka richi tap6
Nan puka tape
Ne chim bo
(lestrum sp. (a8275)
Mish kin tape
Malachra ruderalis (40557)
Naka kushnu tape
Nemo tape
Insa ruiziana (40t398)
Calyptranthes sp. (41045)
Tovomitopsis sp. (48 I 92)
Cyclanthus bipartitus (48078)
Witheringia solanacea (40308)
C i s s a m p e l o ss p . ( 4 8 1 5 3 ) ; G u r a n i a
megistantha (48308)
Miconia nervosa (48099)
Minga shi chi tape
Mish eue chi
Mish timu tape
Mish yo kia
Nan remedio tap6
Naba tape
I)esmodiurn axillare (40606)
Oa na sa tap6
Solantrm marnmoslrm (40487)
Piper tricuspe (40859); P. tubercu-
Oa nano
latum (4i1375)
Oa do tape
Oco tun
Asplenitrm hallii (40075)
Loranthaceae (40056)
O s s a e al a x i v e n u l a ( 4 8 1 7 3 )
Witheringia solanacea (48l 15)
Calathea metallica (aBl 17);
(48215)
Annona muricata (48103)
Danaea humilis (,18062)
mayteno tap6
Onclo tape
Clidemia crenulata (41060)
Capparis ecuadoriana (40097)
Amphidasya sp. (41059)
Onia tape ba n vino
tape micharo kola
Conostesia dentata (48100)
Na ishan tape
R e n e a l m i a o l i u o s p e r m a ( 4 0 16 0 )
Lomariopsis japurensis (40 I 63)
Na ka llullu o kera
Impatiens balsarnina (48362)
Opode
Cestrum sp. (401tr4)
Costus laevis (48097)
tchumo
Na pata barojo
Na sun pi de
Na tini
Pa ban go! la ljuin
Curania pedata (40052)
(luatresia riparia (40099)
chi remedio tap6
Pa chi
Pileasp. (40185)
Pa uatucle
Guatteria sp. (a0736)
Cleiclion castaneifolium (40077)
BS 46
158
Pabano tape
Pachino tape
Bixar orellana (.4047 2)
Piper sp. (41021r)
Pala tape
Ch e n opodiutn arnbrosi oicles
(40f157)
Anthtrriurn sp. (40737); A. falca-
P:rlrnira
t t r m ( 4 8 0 7 3 ); A . s u b c o e r t t l e s c e n s
( 4 1 0 5 4 ) ;A . t r i s e c t u m ( 4 8 1 l B )
\Arettinia quir-raria ( 4 | 07 4)
Paiko tape
Pi chui tapc
Anrndinella berteroniana (4Oir8ir)
rnenecha l'ranbi
kino tsorni tape
Pi chr.rn puka
Pi fichi chi llulhr
Pi ishan ari
Pi peno tape
Pi shun oui tap6
Oa,v:rponiasp. (4079 1)
(lallianclra angustifolia (40556)
Bolbitis nicotiilnifbliunr (40 I 46)
Xiphicliurn cirertrletrrn (48,1I 7)
( l o l u r n n e a p i c t a ( 4 0 7 6 0 ) ; C l .r t r b r i a cuta (40880)
Palo de mahaguzr
Poulsenia arrnata (48343)
Palo cliura
Potrrourna hirsrrtipetiolata
(4100e)
Pl-r-vllanthusanisolobus (400 I 0)
Pi tape
Psvchotria caenrlea (40014)
Pi tchape
Tect:rria yiyipara (,111322)
; Tl-relypt e r i s s er r a t a ( 4 U 4 1I )
Iriartea deltoicleir (48409)
Ocinrurn sp. (4t3445)
Clariczrpapava (40485)
Piba loke cusl-rnatape Scapl'rvgkrttisproliferii (40855)
()eophila herbacea (,10592)
Pibato tchumo
Paltr
P:rrnbil
Papa chinga tap6
Pirpallarfinchrrn<r
ptrka
Papaya
Papav6
C l a r i c ap a p a y a ( 4 0 1 5 9 )
(larica papava (401 119)
P a p i c h a f i n o p t r k a c h i Phvsalis ansulnta (40609)
Pseudcranthern ttm lanceolaturn
Para amp:r
( 4 0 2 1l )
Rollinia rnucosa (,10046)
Pzrstanu
Pata baroj<r
Podandrogyn e brer,ipedr.rnctrlata
(40057)
Pipcr irnperiale (40596)
lechuua tap6
Pichi ancr
(l:rlatl'rca rnirrantifolia (401 I 7)
Pichi ri chi cle
O s s a e ar n i c r a n t h a ( 4 0 I 6 2 )
Za ntia lin<ler.rii (40203)
Pichi ri chi
Pichi ri cle
Pichinga tape
Pichiva
Piclia chi
Bertiera procrrnrbens (40069)
S c r o p h u l z r r - i a c c a c( 4 I 0 2 u )
Heteropsis oblongifcrlia (4000I )
P c r t r z o l z i as p . ( 4 0 4 7 7)
remedio tape
Pika
Acah'pha sp. (40055); A. sp.
( 4 0 2 2 8 ) ;A . s p . ( 4 0 2 2 f 1 )
Par,onia fiuticosa ( 40029)
Asteraceac (48443)
V r : r b e n : r c e a e( 4 1 0 0 7 )
Pika ha
Acalvpha sp. (400t15)
Ltrdrvigia erecta (40ir02)
Piketa tchtrmo tape
( ) o n z : r l : r q u n i ac o r n i f o l i a ( 4 0 5 9 1 )
Pe cl.jupistch:r
Pcntasonia rnacroph-vlla (4836I )
Pent:rsonia sp. (4l0it2)
Pe cljrr pistcha tape
Pe rnolsc
P e n t a g o n i as p . ( 4 1 0 0 1 )
(lostr.rsptrlvenrlenttrs (40 I 73)
Pikini tape
Pilir rntr chi
Urera sp. (40471)
F i c t r st r i a n a e ( 4 0 8 0 3 )
Ficus obttrsifblia (40047)
Pd pin remedio tape
A d i a n t u m o b l i q u u m ( 4 8 13 3 )
Pteris pungens (482611)
Pirni chui n chi
Myrcia sp. (40743)
Pin dik tape
Pi pun chi
Discophora gui:rnetrsis (407 11)
Pin ka suzrnu tape
Aster:rceae (40ir113)
Fanrilr, inclet. (40[J93)
Pe tirn
Verbena litoralis (4020U)
Hyptis rnutabilis (40039)
Desmoditrrn axill:rre (40022)
Blakea subconnata (10297 )
Pin lla chi
Paur pii tap6
Patscl ava
P€ clju tap€
Pe oy chi tap6
P d c l j t rp i s t c h a c h i
PE pin tap6
Pe tup
Pega pegir
Pei ka
Pefr:r rnono
Per-rpcr
Pcnpo tape
Peo tapc
Peseulo tape
Petik tap6
Pi ari
Pi cha pi
Pi chi
Pi chin ga
Pi chui tape
Piketa tchurn<r
llullu tap€
Pilucle
Pin remeclio tape
S o c r a t c i re x o r r h i z a ( 4 U 4 1 0 )
Besleria barclar''i (41 01 1) ; Ohr,vsot h e r n i s f i i e c l r i c h s t h a l i a n a ( 4 U 0 8 0 );
r\peiba rnembran:rcea ( 48 I 9[t)
Aster.aceac (,10104)
C l i c l e r n i a c r c : n r r l a l : r( a 8 0 7 7 ) ;
Clolurnne:r bilabiata (40ti00) ;
( i e s n e r i i r c e a e ( 4 10 0 5 ) ; I c h n : r n t h u s
A s t e r a c e : r e( 4 0 1 9 0 )
( l l i b a c l i t r r n s p . ( 4 0 t t 0 7 ); ( 4 10 2 5 )
pallens (48090); Kohlcria sp.
( 4 0 5 7 6 ); M o n o p v l e s o d i r o a u i r
Hibiscus sp. (40579)
Asteraceae (401163)
T l i e l y p t e r i s s p . ( 4 0 13 8 )
Tectaria incis:r (40902)
( 4 0 5 13 ) ; P i p e r z r n s u s t r . r n(r4 0 7 3 U );
S e l a g i r r e l l as p . ( 4 0 f 1 7 8 )
S a p i t r r n s p . ( , 1 10 , 1 1)
Pin sa chi
Pin tapi:
Ficus cervantesiana (40765)
Scrophulariact::re (1081 2)
Pin tape capitin
Diplazitrrri bombonasae (48320) ;
Elcocharis retroflexa (48327 ) ; The-
Pin tape
lvpteris resinil-era (48296) ; Satrvasesia erecta (48326)
A c i o t i s c z r t r l i u l a t a( 4 0 7 1 5 )
(lornrnelinaceae (.1U4B
I ); Erythnrdes weberana (40879)
N e p l r n r l e p i s p e c t i n a t a ( ' 1 8I 3 5 ) ; P i per filistvlum (40746) ; Psysrnorchis
p r r s i l l a( 4 l 0 ' 1 : 3 )S: rn q o t t i t t t nt n r t crophvlltrnr (407trb7
BS 46
Pin tsu ruro chua
Pincla tape
Pinde tchuba
Pini ara
Pini nicaca
159
Cissampelos tropaeolifolia (40tt5 1)
Clolumnea bilabiata (40519)
Bar.risteriopsiscaapi (40582)
(ltenitis sp. (40145)
Puyan oco molo
Podandrogyne brevipedunculata
(40166); (40209)
Raca pies
Triolena barbeyana (48032)
Scrophulariaceae (40300)
(lolocasia esculenta (48l 45)
Ranelis
Dicranoslossum sp. (40234)
Dicranoslossum polvpodioicles
(40 I 70) ; Drymonia rnacrophvlla
( 4 0 16 7 ) ; D . w a r s z e r v i c z i a n a( 4 0 0 U 0 );
Rascadera
l , y c i a n t h e sa m a t i t l a n e n s i s ( 4 0 0 8 7 );
Rojina
Portulaca oleracea (40604)
Piper angustum (40303)
Pleopeltis percussurn (401 96) ; Triol e n a b a r b e y a n a ( 4 0 0 9 4 ) ; ( a 0 15 7 )
Rosaflor
Asteraceae (40024)
Rueil
Pinion roja
Pinta
Errplrorbia cotinifolia (10177)
Clvclanthus bipartitus (4022|)
Sa he
Peperomia urocarpa (40301 )
Microgramma piloselloides
(40302)
Pipeso
Asplunclia sp. (401ttlt)
Pipetio
Capsicunr fiutescens (40565)
Luclwigia erecta (401 7U)
Sa oa pa chi
Sa pichua tap6
Zamia lindenii (48155)
Microgramma reprans (48 I 83)
Pini tape
Pipili tape
Pique ta
tchumo llullu
Pique ta tchumo
llullu tape
Pique ta tchumo tap6
Pischtinea tchape
Pischus finchuno
Pisgo pirno tape
Pish chua tap6
Pishillo
Pisi chua llLrllu pistcho fino pi chua
Pispata tchumo tape
Pitsa
Piya
Pii,a ha
Po koi tape
Po pin ha tape
Po pin tape
P6 rernedio tap6
Pochiski
Ponpo ktrchu
Posude
Psychotria santarernic a $0 ag2)
kino tape
Sabi ballo tap6
Sajo chi
Hzrmelia patens (40897)
Sajo de arriva
Sarnbi oai llullu
Ltrclwigia erecta (40490)
San fania chi
San fania
Sanjuania tape
Scrophulariaceae (48 148)
Zea rnays (40525)
Drymonia alkrplectoides (4050tt)
Manclevilla hirsuta (40800)
Insa semialata (40733)
M a n d e v i l l a d o d s o n i i ( a 8 3 0 7 );
Tropaeolum repandum (4ti306)
(lomrnelina clifTusa(40598)
Irrqa eclulis (40042)
Acalypha cliversifblia (4001 I )
Acalvpha cliversifblia (40 I 115
)
Harnelia sp. (41051)
S o l a n a c e a e( 4 i l l 1 4 )
Piper sp. (48163)
D e s r n o d i t r r n a x i l l a r e r( 4 10 7 5 )
Erynuium fbeticltrrn ( 4003U)
Geosenanthus rhizanthus (40 I 06)
Cestrum sp. (40092); Solantrm leptorhachis (40073)
Pu chua
Re o machi
renedio tape
Reuma tap6
San tap6
San tap6 chi
San tape
Sancona
Sanda polo chi
Santa maria
Santa maria tape
Sao jtranillos
Sapo
Sapote tape
Sapu djui tape
Sard sa chua
Sebero puka tape
Seiton tape
Sek tape
Schleeelia fastisiata (4lll I I )
Piper hispidtrrn (407 48)
Piper sp. (40776)
S€n gi chi
Puban kura
Pukano mavtein
Inga eclulis (40778)
Siparuna sp. (40216)
Elaphoglossum sp. (40809)
Shi bish tchape
Shi nroc<r
sumo tape
Pui tap6
Pteris punsens (48?ft3)
Shi vesh tape
Shi vesh tchzrpe
Shil'ingola tape
Drvmonia sernrlata (40804)
Myrtaceae (48401)
Shili anpo
Pu pin ha
Pu pin ha tape
Pu shillo
Pulu chua
Puno chui chi hal ki
Punta lansa
Puv pa tapa
C o l u m n e a p i c t a ( 4 0 2 3 0 ) ; C j .r u b r i a ctrta (40074)
Asteraceae (40037\
Shan tape
Shia remedio papa
Shili ara
Trianaeopiper sp. (48104)
C i s s u ss p . ( a 0 4 8 6 )
Tapirira guianensis (48277)
Tapirira guianensis (48277)
Santalaceae (48423)
Renealmia cylindrica (41042)
Renealmia cylindrica (40833)
Renealnria cylinclrica (48235)
Aechmea magdalenae (483tt6)
Discophora suianensis (41022)
Discophora suianensis (48lGB);
Meliaceae (48289)
S o c r a t e a e x o r r h i z a ( 4 8 4 10 )
Lauraceae (41008)
Ipomoea sp. (40005); Porhomorphe peltata (40004)
Piper peltatum (48159)
Renealmia oliuosperrna (a0160)
C l a r i c as p . ( 4 0 i 5 8 )
Quararibea cordata (40580)
(]asteranthus ()ncosastrus (4 I 000)
Marcsravia sp. (a8337)
(lanna inclica x seneralis (40530)
Urena lobata (40041)
Brorvallia americana (40040)
Cecropia sp. (48433)
CampvloneurLrm phyllitidis
(400e5)
Burmeistera sp. (40876)
Arthrostema ciliarum (48053)
Burmeistera vulsaris (4103I )
Burmeistera sp. (48070)
C l o s t u sl a e v i s ( 4 1 0 0 2 )
Xvlaria sp. (a8352)
Piper eustvlum (40237)
I)ennstaedtia arborescens (400t15):
l,ornariopsis nigropaleata (40020) ;
(40223\
BS 46
160
Shili aya
Shili ayan
Heisteria sp. (40064); Schlegeliafastisiata (40127)
Anthurium trinerve (40141)
Tdmpo remedio tape
Tdmpo tape
Peperomia pellucicla (48287)
Peperornia sp. (40506); P. sp.
( 4 0 5 7 8 ) ;P . s p . ( 4 8 0 7 2 ) ;P . s p .
(4U076) ; Sphvrosperrnurn cordifolium (40830)
Guzmania melinonis (40885)
Shili mira
Philodendron inequilaterum
( 4 0 r7 1 )
Shili penpo
Shili pese
Asteraceae (40103)
Asplundia sp. (40118)
Tenba yu ki ha
Shili peso
Shili shinpi
Evoclianthus funifer (40050)
Tenco shili
Tia mu ki chi
Tiban ara
Cissampelos tropaeolifolia ( 40030)
Perebea xanthochvrna (40tt35)
Diplazium cristatum (40i49)
Shui pu chua
Shui tungui chi
Clorrania lupuloides (407 44)
Heisteria sp. (40747)
Tirn bu rukula ha
Salpichlaena volubilis (407 63)
Silantro
Silla killan
Eryngitrm foeticlum (4003f1)
Solanum lepidoturn (40536)
Tiu pin tape
Microgramma pikrselloides
(40236)
rerneclio tape
Cvathula achyranthoides (400 I B)
Sji pistcha ka bo kaka R h e e d i a e d u l i s ( 4 8 3 6 3 )
Funsi (41034)
Sjia papa
Clidernia cliscolor (40307)
Soe
(]asteranthlrs oncosastrus (40734) ;
Soko bo chui tape
Sino tape
(48126)
Soko oa! ko tape
Sopo
Sopo oal ko tape
Piper sp. (4Bl l3)
Carica sp. (40088); (40195)
Piper tyianae (41020)
Spa lau gilla
Scaphvglottis graminifolia (483 1B)
remeclio tape
kino chua
Tiu pin tiu
Tir"rtirr kuru tape
To ari
(40238)
To kimide
To pinktr
To pirin
To ua ara
Tokillade agua
Tokiya de vehago
Tolonbo ha
Toma be
remedio tape
remedio tape
Stira pi killan
Su pu uactr tape
Drymaria cordata (401 79)
Piper sp. (40752); (40793)
Suambe chua
Supla chi
Mikania sp. (41036)
Protium sp. (41006)
Supo hopi trallu
Supo hopi uallu tap6
Acalypha villosa (40838)
Acalypha sp. (48360)
Supu pi shungui tape C o l u m n e a e b u r n e a ( 4 0 8 7 0 )
Asteraceae (40152)
Ta cantsa
Farnily inclet. (40894)
Ta pi chi kino chua
Topin remeclicr
Scoparia dulcis (40567)
Cuphea strisulosa (4003 1)
Tsabo tap6
Tsantsalo
Tselen shilial,an
Tsoda
Tsrrprrm prrka tape
Tape finchuno
Rvticlostvlis quadrifida (40607)
Prestonia portobellensis (41052)
Tun ban chi
Td chinga tape
Te pui pu pin ha
Family indet. (10848)
Senna dariensis (40751)
Te sa tape
Tebaho killan
Thevetia peruviana (40545)
chi tap6
Tebgo killan chi
LufIa aegJptiaca (40597)
remedio tape
Tebajo killan chi tape Chenopodium
(40575\
ambrosioicles
Temba ho killan
chi remedio
Gomphrena slobosa (40561); Larniaceae (40854)
Temba ho killan chi
remedio tap6
l,amiaceae (40861)
Aciotis sp. (48316); Pilocosta oerstedii (48387); Trichornarles sp.
(48408)
Tu rnai n chi
Tumu pin tap6
Ischnosiphon leucophaeus (40780)
Eryneium fbetidum (48359)
Hemiclictyum marsinAturn (40 I 28)
Asplunclia sp. (40 litti)
Evodianthus frrnifer (40050)
Calathea metallica (40204)
Thelypteris francoana ( 48294)
Brugmansia versicolor (4058 I )
Mirabilis.jalapa (40909)
(iustavia sp. (48141)
Te tera
( l e s t r u m r a c e m o s u m ( 4 0 0 8 1)
Piper hispiclum (40222)
()eonoma cuneata (40207)
Toma b6 tape
Tonua tape
Ta de llullr-r
Tan tchap6 finchttno
Tcho maceranu tap6
(lolumnea nrbribracteata (48420)
Capsicum fiutescens (40566)
Bromeliaceae (40729)
D i p l a z i u m s t r i a t u m ( 4 0 1 6 U );
Tun
Cavaponia slandulosa (40I 00)
Philodenclron sp. (40115)
(irias penrviana (40220)
Annona sp. (4086f1)
Oephaelis gentryi (48108)
Drynonia
(4B4le)
warszewicziana
Passiflora rnacrophvlla (40tttr5)
C a p s i c u m { i u t e s c e n s ( 4 0 2 0 1)
Ua anpo ha
Piper sp. (40109); P. imperiale
( 4 0 1 9 9 ); P . o b l i q t r u r n ( 4 0 2 1 0 )
Ua anpo
Piperr angustum (40090)
Piper mtrltiplinervium (40059)
Diplazium striattrrn (40051 ) ; Szrc-
Ua anpo shibi
Ua ar:l
c<rloma elesans (40225)
Ua bonban
Or-vctanthus occiden talis ( 40058)
Ua curan tape
Ua ga la pu chi
Blechunr brownei (40183)
Grias penrviana (40832)
Ua halki tape
Pilea sp. (40217)
Ua ishan tape
E l a p h o g k r s s u r n s p . ( 4 0 13 3 ) ; T h e l , v p t e r i ss p . ( 4 0 1 3 2 )
Hamelia axillaris (40054)
Ua kere nayzr
BS 46
Ua ko tape
Ua mana shuba
Ua pichiano
Ua pini ha
Ua pini tap6
Ua shinpi
Ua tovali
Ua tsero tap6
Ua tsetsero
Ua unna pini ari
Ua chambi tape
Ua chambi
Uai toto tap6
161
Piper filistylum (107 46) ; Trianaeopiper sp. (41068)
Peperomia sp. (40079)
Heliconia aemygdian a (40120)
Alloplectus dodsonii (40053)
D r v m o n i a r h o d o l o m a ( 4 0 10 8 )
Pleopeltis percussum (40219)
Ya peno maiten
sumo tape
Dendropanax arboreus (40899)
Yu ban go cho
Yu pei n chi
remedio tape
Yucca blanco
Manihot esculenra (48330)
Dichorisandra hexand ra (48407
)
Yucca rojo
Yuka bare himu
tchap6
Zapata chi
Drymonia turrialvae (40224)
Trianaeopiper earciae ( 40227 )
S v n e c h a nt h u s w a r s c e w i c z i a nu s
(40205)
Zapata
Lomariopsis nigropal eata (4022g)
Witherineia sp. (48349)
Witherinsia solanacea (41 0b3)
Trichomanes membranaceum
(48345)
A chide
Achiote
Asteraceae (40024)
Aguacate
Ualan tape
Uali ha
C a l a t h e as p . ( 4 1 0 6 1 )
Al6n
Anthurium
Alanko
oveophilum G0I2g) ;
Philodendron renue (4022G)
Albaka
Ardisia sp. (40192)
Scleria prerora (40802)
Panicum sp. (aB32B)
Alena sili
Ano tap6
Anotada
Euphorbiaceae (41038)
Trichilia pallida (40000)
Apilon
Uero kinto
Passiflora auriculara (40105) :
(40218)
Ara
Asususili
Ugachi tape
Ugala tap6
Lantana sp. (40548)
Dracaena fragrans (40541 )
Raritebe palicoureoides (48066)
Psvchotria macrophyl la (4007 2) ;
( 4 0 1l 4 ) )
Uam bio tape
Uanmo es tape
Uase chi
Uenbanna
Ui pistchia boka
Uinca
Ukera chuno tape
Ukera tape
Un sa lala he tape
Unga lala upi
uallu tape
Uni ha chi
Unilo piyo
Ustin tape
Uvica
Vehika
Veloha
Veneno tape
Viki chua ama
Sauvagesiaerecra (40589) )
Polygala mollaeinifolia (a0588) )
Clidemia purpurea (40896) )
Acalypha diversifolia (40875) )
Apira tape
Baku
Crescentia cujere (40728)
Clibadium sp. (4909b)
Amaryllidaceae (40623)
Barbasco de hoja
Baren apisu
Betende gigantape
Betia olorosa
Bitu baca
Bobo
Bonban
Cacao de monte
Anthrrrium orregeanlrm (40202))
Biophvttrm dendroide s G0b77))
Cafe
Philodendron subhasratum
(4Bl7e)
chi
Ya hino rnayteno tape Danaea sp. (48a37)
Sclerothrix fasciculara (406b4)
Herrania balaensis (40666)
Crotalaria nitens (40727)
Gasteranrhus crispus (40701)
Banisteriopsis caapi (40722)
Cacao
Cyclanthus bipartitus (a0837) ;
Geonoma cuneara (10753) :
( 6 0 0 9 9 ) ;( 6 0 1 1 8 )
Ocimum campechianum (40616)
Aristolochia pichingensis (407 I ti)
Asteraceae (40602)
Atolon
S o l a n u m c a n d i d u n i ( 4 0 1 0 5 ))
Carpotroche plaryprera (a0060) )
Ya hal chi
Persea americana (4gl l8)
Persea americana (4071 5)
Auedo tap6
Ayahuasca
Buru sinc-r
Cacao ara
Anthurium lancea (41048)
Anthurium sp. (40850)
Diplazium rnoccenianum (48049)
Bertiera guianensis (4g0gl )
Bixa orellana (40704)
P e r s e aa m e r i c a n a ( 4 9 l l 8 )
Saccoloma eleg;ans (49040)
Prestonia rotunclifbli a (4067 a)
Erythrina edulis (40646)
Tococa spadiciflora ( 0B7a) )
A l l o p h y l u s s p . ( 4 0 1 8 1 ))
T r i c h o m a n e s e l e g a n s ( 4 0 8 8 2 ))
Viki mo tape
Viki mo tchape
Viki mo tchap6
Solanum nudum (48271); (a830a)
Solanum nudum (41078)
Colorado
Uaita
Ualina
Manihot esculenra (48329)
Manihot esculenra (48330)
Rytidostylis carrhaginense (48428)
Campano
Chide ard
Chide lapa
Chide bun
Chimpalo
Chonta
Chonta dura
Cop6l
Senna rericulata (10OZb)
Aristolochia pichingensis (407 I B)
Hedychium coronarium (49lZ1 )
Clarisia racemosa (40712); (49109)
Loranthaceae (40634)
Gasteranthus corallinus (40665)
Asplenium serrarum (40717)
Herrania balaensis (40666)
Theobroma cacao (40618)
Coffea arabica (40645)
Brugmansia versicolor (40624)
Lonchitis hirsuta (49036)
Pleopeltis percussum (40629)
Dicranoglossum polypodioides
(4064e)
Sapiumperuvianum(49098)
Solanum canense(10724)
Bactrissp. (49123)
Bactrissp. (49122)
Protium colombianum (49097)
r62
BS 46
Cura paja
Ciruela
Tectaria nicotianifolia (40686)
Spondias mombin (49100)
Lim6n
Dela
Canna sp. (49051)
Alloplectus sprucei (49052)
Alternanthera mexicana (40653)
Napeanthus robustus (49 108)
Asteraceae (40647); Dahlia sp.
Loba tape
Acalypha sp. (a0725); Piper sp.
(40711)
Loki
Brugmansia versicolor (a0624)
Carapa mesistocarpa (40703)
(40642); Fabaceae (a0620); Hydrangea sp. (40641)
Lu koko lan
Luban tap6
Dicranoslossum polypoclioides
(4064e)
Lindernia crustacea (40636)
Luisa
Denki tipe
Descansel de monte
Doril
ajdn
Epe tap6
Crama de palo
Halki tap6
Hake tape
Helen descansel
Helen llaten
Helen mudu tape
Helen paki
Helen puban kara
Helen punpis tape
Acalvpha sp. (a06a8)
Alternanthera mexicana (40653)
Encyclia fragrans (40668)
Ceophila gracilis (40644)
Peperomia panamensis (a0638)
Siparuna eggersii (40643)
Asteraceae (40707)
Helen santa maria ha Piper multiplinervium (40684)
Hierba luisa
Cymbopogon citratus (491 l4)
Hierba rnaldita
Gasteranthus crispus (4070I)
Hoja de sapo
Begonia glabra (40696); Trianaeopiper mexiae (40667)
Isan ha tap6
Isan tape
Ishan luli
Kafu mura
Kaheli
Kai no aeria
Kaku sili
Kantsa mula tape
Karan tape
repens (4067 2)
Alloplectus sprucei (40706) ; Niphidium crassifolium (40639)
Campyloneurum
Bougainvillea sp. (49090)
Streptochaeta sodiroana (407 | 4)
Clibaclium sp. (a9095)
Costus sp. (40678)
Dryrnonia rhodoloma (40716)
Hyptis capitata (40635)
Desmodium uncinatum (40614)
Lim6nsillo
Lonco
Lu clico
Luli chide
Luli tap6
Meli
Citrus sp. (a0720)
Piper sp. (a0683)
Cordyline terminalis (40640)
Costus sp. (40678)
Amaranthus quitensis (406?7); Hibiscus radiatus (40637)
Cymbopogon citratus (491 l4)
Nothopanax fruticosus (4061 2)
Asteraceae (40702)
Rubiaceae (49120)
Mamajuana de monte Asteraceae 40707)
Mo du! tape
Diastema scabrum (49092)
Modo tap6
Hydrocotvle leucocephala (40628)
Mor6l bobo
Moril
Mudruha
Muhu
Na ari
Na disku
Na isun ja
Na kerena
Na kirni chi de
Na kirni de
Na koko lan
Na kolotu empoya
Na kototo ha
Na licade
Na mira
Clarisia racemosa (40713)
Maclura rincroria (49060)
Myrtaceae (49094)
Bixa orellana (40704)
Lomariopsis japurensis (49038) ;
Thelypteris urbanii (49030)
Renealmia oligosperm a @07 12)
Codiaeum variasatum (4061 5)
Hamelia axillaris (40626)
Asclepias curassavica (4061 l )
Conzalagunia dodsonii (40661 )
Costus pulverulentus (40700)
Piper sp. (40699)
Trianaeopiper mexiae (40667)
Picramnia cooperis (40659)
Monstera adansonii (49054)
Urceolina grandiflora (491 19)
Peperomia panamensis (40638)
Na sili ara
Na sili inpo
Lantana sp. (40622)
Trichomanes collariatum (40695)
Piper sp. (49099)
Bomarea edulis (40651)
Gurania macrophylla (40692); Solanum dolichorhachis (40691)
Na sili kototo ha
Na sirnpi
Piper hispidum (40681)
Musci (40680)
Na tarali allar
Campyloneurum
repens (49037)
Diastema affine (491 I 1)
Aspleniurn serratum (49032)
Ni pini
Napeanthus robustus (40671 )
Dicranoglossum polypodioides
(49029)
Naranja
Citrus sp. (40721)
Piper sp. (49043)
Aphelandra sp. (40710)
Naranjilla de monte
Nep6
Passiflora fbetida (40660)
Cordia spinescens (40663)
Rubiaceae (40685)
Ortiga
Pajarito
Solanum candidum (40656)
Banisteriopsis caapi (40722)
Urera baccif-era (40669)
Psvclrotria williamsii (49044)
Citrus sp. (40721)
Pi av6
Lansa
Lati
Pourouma guianensis (40677)
Pini kola
Lengua de culebra
Campyloneurum angustifolium
(49039) ; Dicranoglossum polypodioides (49029)
Pini kolosica
Katsa moca
Kere dora
Kika kala sili
Kinfo aran sili
Kolin ari chili
Kono tape
Koril ahen
Kostoto tape
Kototo tap6
Krana chia de m()nte
Kuchi
Kuku pi piyan ha
Kuru bi bien
Na pen po
Pe cotamoja
Pini ha
Loranthaceae (40634)
Begonia elabra (40696)
Bactris sp. (49122)
Drymonia coriacea (4067 3)
C o s t u ss p . ( 4 9 1 1 3 )
Pini luli
Clavija eegersiana (40708)
Gurania mesistantha (49050)
Pini sili
Columnea ansustata (40709)
BS 46
Pini tape
Pinin tipa
163
Kohleria sp. (40723) ; Oxalis microcarpa (40726); Oolurnnea angustata (49106)
Uvica
Pourouma guianensis (40677)
Solanum candidum (40656)
Voche kinto
Passiflora foetida (40660)
OarnpvloneLlrun] angustifolium
Ye sili
Mendoncia brenesii G0705\
Uva de monte
(4e03e)
Pinion
Postrde
Ptrban kura
Ptrnta lansa
.fatropha curcas (40633)
(lestnrm sp. (40688)
Sipartrna sp. (40693)
(loltrmnea eburnea (49057); C. rubriacuta (40671'r)
Coaiquer
Achiote
Achutillcr
Acoronsillo
Bixa orellana (48897)
Allophylus excelsus (41504)
Dussiasp. (41656)
4ii
Sa lie
Begonia sp. (49059)
Macltrra tinctoria (49060)
Capsicum frutescens (48988) ; Solanum sp. (41648)
Sirn fania
San juar-rilla roja
Rerrealmia olig;osperma (407 72)
Oorclyline ternrinalis (40640)
Ajo
Amaryllidaceae (41604) ; Amaryllidaceae (48772\
Sande
Shili arl
B r o s i r n u m u t i l e ( 4 9 11 9 )
Microsramma piloselloicles
( 4 0 6 17 ) ; T r i c h o m a n e s c o l l a r i a t u n r
Al galga
Alrnurillo
Cranichis sp. (a8927)
Nephrolepis pectinata (48846)
Marila laxiflora (4161 5)
(4 e 0 2 8 )
Anime
Vitaceae (40631)
Piper sp. (401170)
Anisillo
Piper sp. (49058)
Piper sp. (40679)
Arccr
Lonchocarpus sp. (49107)
Asplenitrm pteropus (40697)
Bqiu
Ranto tape
Shili a,van
Shili empo ha
Shili impo
Shili kototo anpo ha
Shili ha hili
Shilina ari ha
Shiri pi ki hen tape
Sino tape
Strn pide
Tancare
Teptrn
Tia kutu pe
Tia kuttr tape
Tichi vil6
Tini
Tiolon
To ari
Ambour6
Ar.,)'
Ava de monte
Balbacha
Drymaria corclata (49096)
Heliotropitrrn rufipilum (40613)
( l u a t r e s i a r i p a r i a ( 4 0 6 5 5 ); ( a 0 6 8 7 )
Balbasallcr
Selaginellasp. (49033)
N a u c l e o p s i s s p . ( , 1 9 1l 2 )
Urera baccifera (40669)
Bandai
Erythrina smithiana (10632)
Diplaziurn striatastnun (49041 )
Behay
Balbasco
Bara blanco
Barbascillcr
Bialbena
Botonsillo
Tirm vaca
Anthtrriurn sp. (40652)
Brosirnurn utile (49110)
Zingiber oflicinzrle (491 16)
L)a anpcl h:t
Piper sp. (49056)
Ua ar:i
Asplenium purpurascens (49035)
I)ichorisandra angustifolia (40698)
Tstrla sili
Tirga
Ua clcm<r
Ua dishtr
Ua halki tape
Rernealmia alpinia (,19093)
Pilea sp. (40682)
Psarnmisia caloneura (48920)
Diastema affine (48990)
Hymenophyllaceae (48838) ; Hy-
(48834)
Oarapa mesistocarpa (40703)
Ocimum carnpechiantrrn (40616)
S e l a g i n e l l i rs p . ( 4 9 0 4 2 )
Bicundo con duende
Tona ara ha
Meliaceae (41667)
Danaea sp. (a8883)
rnenophyllum microcarpum
(48956) Lvcopodiurn sp.
P e p e r o m i z rp e r n a m b u c e n s i s
(49053)
Acli:rnturn trichochlaenum ( 406U9)
To kinclo
Protium amplum (41659)
Piper sp. (a1421)
Bolccr
Bora negra
joosia sp. (a8969)
P s e r d e l e p h a n t o p r r sp r r r a t i s
(48713)
C o r d i a s p i n e s c e n s( 4 1 4 9 8 )
Ocotea ira (41671)
Pseudelephantopus sp. (41609)
Dioclea sp. (41640)
Acanthaceae (41459) ; Lycopodiella
trianzre (41499)
Cuzmania resttrdinis (48U56)
Peperomia sp. (48906)
Desmoncus cirrhifera (41455)
Caballo de monte
Spilanthes sp. (41485)
C a s s i as p . ( 4 1 4 6 5 )
Claimitillon
(larnate
Porrteria collina (4167 3)
Clusia dixonii (48833)
(lafra asrio
Costussp. $1a57);C. sp. (a8860);
C . r p . ( 4 8 9 1 7 ) ; C . l a e v i s( 4 8 9 0 7 )
Candelelio
Columnea minor (41428)
Ua ka avi
Uzr kototo tape
B a c t r i ss p . ( 4 9 1 2 3 )
Piper trizrnae (49055)
Cangare
Dialyan thera gordoniifolia
(41664)
Lla luli
Allarnanda cathartica (4071 9)
Carillo
Ua ptrban ktrr:r
S o l a nr r n r c o n f e l t i s e r i a t r r m
(40630)
Anthurium napaeum (40676)
Cleiclion castaneifolium (40658)
Malaxis sp. (48709); Miconia goniostigma (48854)
Caruavale
S e l a g i n e l l as p . ( 4 8 8 3 1 )
Castilla elastica (4U994)
Lauraceae (41675\
Ua tarali allan
Uatucle
Cauchcr
Chachajo
764
Chaguar6
BS 46
C r e m o s p e r m a h i r s r r it s s i m u m
( 4 1 6 11 ) ; C u p h e a t e t r a p e t a l a
( 4 8 6 9 9 ): S c r o p h r r l a r i a c e a e
Escudera
Estacudo
Solanum mammosum (41452)
(41 430) ; Scrophulariaceae (41 424)
Flor de duende
Flora amarilla
Psammisia ferruginea (48792)
Stenospermation densiovulatum
(48B5e)
Chaua
Ruagea sp. (41672)
Cestrum baenitzii (41650)
Pentasonia srandiflora (4i599)
Justicia comata (48715)
Chicharo
(ihilangua
Melastomataceae (4 I 666)
Elaphoelossum sp. (48842)
Chilungua
Gualbadea
Chirnbore
Erynuiurn foetidum (48926)
C e p h a e l i s c o y e t e n s i s( 4 8 9 3 1 )
Chiparo
S w a r t z i as p . ( a l 6 0 7 )
Guandd
Chira rau
Ocimum campechianum (48982)
Erechtites valerianifolia (48tt66)
Dioscoreaceae (48826)
Guaral
Cuaripo
C r e m o s p e r m a c o n g r u e n s ( 4 1 5 9 1)
Psammisia ulbrichiana (41 487)
Capsicum fiutescens (48988)
Guaya busai
Tabernaemontana macrocalp<
(48933)
Guayusa
Hedyosmum sp. (48769);H. scorerrimum (41474)
Gueche grande
Alloplectus panamensis (a8870)
Peperomia serpens (48825); Piper
sp. (48697); P.sp.(48703); P. sp.
Chalde
Chalmolan
Cham
Chiraran
Chirma
Chorillo
Chu kin llullu
Chuil
Chulku
Chundul
Peperomia sp. (a1645)
C,vperus odoratus (41624)
Forda
Gia
Goaralla
Stenospermation angustifirlium
(48872)
Grenadilla
Passiflora sp. (ait857)
Disterigma sp. (4t3966); Psamrnisia
Cuasca negra
Churimoya
Trichomanes sp. (48894)
R o l l i n i a m L r c o s a( 4 B g 8 9 )
Rollinia mucosa (48989)
Coca
Erythroxylaceae (41 584)
Coldillo
Cremosperma reldioides (48980)
Columnea lehmannii (48983)
Guetch
Alloplectus tetragonoides (4BBl 3) ;
Cremosperma nobile (aB97tr);
Guildd
Churillo
Churimonu
Concedillo
Contra
Corillo
Corosillcr
Crista de gallo
Cuerda
Dai n
Descansel
Detr
Deu(f)l
Deuendi
Dia ku(h)
D.jet
Du
Duende
Duencle ku(h)
Dundu
Escadera
Escobilla
Guech6
C}rin ganul
Guinul
Billia colombiana (41663)
Anthurium andreanum (41 489)
Guish ko
.fusticiasp. (41483)
Palicotrrea sp. (48727)
Hymenophyllum microcarpum
(48e56)
Satyria srandifolia (48806)
Clalathea timothei (48960)
Diplazium lechleri (48948)
Besleria barbata (18734)
debilis (48968)
C l u s i as p . ( 4 8 7 8 1 ) ; C . s p . ( 4 8 9 3 2 )
Araceae (48711)
Lauraceae (41661)
Guatteria sp. (41655)
(4u824);P.sp. (48853);P.sp.
( 4 8 8 5 8 ) ; P . s p . ( 4 B t ' ] 6 1 ) ;P . s p .
(48873)
Drymonia serrulata (a8773); D.
ttrrrialvae (41 638) ; Paradrymonia
sp. (41635)
Desmodium adscendens (41578)
Pitcairnia sp. (41496)
Pseuderanthemum micranthum
(48935)
Erato polymnioides (41 505)
Bauhinia sp. (a1674)
Oleandra sp. (4tt843)
Cuish
Clolumnea rubriacuta (41622): Piper dryadum (41612)
Triolena spicata (4U905)
Solanum triplinervium (48914)
Thelvpteris gigantea (48885)
Costus sp. (48917); Psammisia caloneura (48920)
Gulpe piguil
Cu1'u
Ha teu
Havilla
Hierba buena
Hierba de monte
Hierba laisa
Hierba nionte
Hoja de mal viento
Hoja de r,ulle
Hoja vercle
Imbien
Calathea micans (48959)
Dennstaedtiaceae (48840)
Dennstaedtia sp. (48884)
Pitcairnia spectabilis (48972)
Dioclea sp. (41646)
Diplarpea paleacea (48723)
Solanum medusocalyx (al 628)
Aphelandra flammea (48898)
S a b i c e av i l l o s a ( 4 1 5 0 6 )
Sancheziasp. (4146a)
Gurania sp. (48981)
( l a v e n d i s h i a c o m p l e c t e n s ( 4 15 0 2 )
Alloplectus teuscheri (48961 )
P h y s a l i ss p . ( 4 1 4 6 0 )
Cavendishia grandifolia (.18965)
Cavendishia srandifolia (48965)
Isha
Lasiacis sorshoidea (41 469)
Anthurium sp. (48811);A. caulor-
.fapate de monte
Kachu
rhizum (48878); Philodendron sp.
(48827)
Kamna pij
Cyclanthus bipartitus (48850)
Columnea sp. (a8739);C. rubriacuta (48738)
Ludwigia alfinis (48864)
Kog(h)
Sphvrospermum dissimile (48922)
Inya
Drymonia warszewicziana (48991 )
Cestrum sp. (48900)
7
BS 46
165
Rhvnchospora sp. (48973)
Piper sp. (a8724)
Danaea sp. (48941); Blechnum sp.
(48942); Thelypteris sp. (48952)
Ovo
Lagalto
Curania sp. (488tr8)
Pailde
Lecha
Blechnum sp. (48940)
Asplenium hallii (48744) ; C,vathea
bipinnata (48943) ; Dennstaedtia
sp. (48889) ; Lindsaea quadrangula-
Paja
ris (41i953)
Coleus sp. (48875)
Papal.uela
Siparuna sp. (48903)
Dryopteris sp. (aBBB7)
Pasino
Monolena primulaefl ora (4897 4)
Rheeclia edulis (41575)
Ped pe
Pi
Poulsenia armata (48985)
C o l u m n e a f i m b r i c a l r a <( 4 8 9 7 1 ) ; C .
Piast
rninutiflora (48970) ; Cremosperma
consruens (48979); Piper sp.
( 4 16 0 0 ) : S c r o p h r r l a r i a c e a e
Piganore
Koltadera
Kramna kun
Ku(h)
Lecho
Lengua tigre
Limon
Llullu
Lombrice
Madroiio
Mahana
Maip6
Mal viento
Malde
Mama juana
(4U863) ; Triolena obliqua (487 14)
Hyptis verticillata (48852)
Indet to family (41670)
Oyap saya
Pagamde
Palo chiso
Pande
Parma
Pausoi
Piau
PU
Pilmo
Pilp6
Clibadium sp. (48698)
Peperomia sp. (a8869)
Pirama
Mancha ropa
Clusia sp. (41662); Visrnia obtusa
(41494)
Pita
Mancha sopa
Manga mora
Vismia sp. (48832)
Aegiphila sp. (a8779)
Masamora
Pite de monte
Mayp6
Aegiphila sp. (4l5l l)
f)oussapoa contorta (41 626)
Dicliptera sp. (48770)
Mentha
Peperomia sp. (41647); P. sp.
Platano
(4 8 e 1 e )
Poleo
Rhvnchospora sp. (48973)
Arachnothrp< inconstans (48768) ;
Pulsande
Maiiana
Mata palo
Mo
Mokillo
Montana cu(h)
Monte cush
Monte de cu(h)
Monte de uish
Piiru cle monk
Platanera
Saurauia brachybotrys (41 621 )
(]leicheniaceae (48950)
Puntero
Trichornanes rieidum (48955)
Araceae (4f1993)
Elleanthus robustus (48984)
Rascadera
Rojo
Morcillo
Miconia goniostigma (41500)
Flacourtiaceae (48780)
Morcillo silvestre
Flacourtiaceae (48702)
Motilon
Hieronima chocoensis ( 41657\
Solanum quitoense (41586)
San pedro
Paulinia sp. (41435)
F a r n i l y i n c l e t . ( 4 16 0 3 )
Sangriado
Monte hierba
Naranjilla
Oabo
Oago
Oaral
Ortiga
Ouish
Ousma
Ovilla
Monstera lechleriana (41633)
U r e r a c a r a c a s a n a( 4 1 5 0 8 )
Tococa svrnphyand ra (487 25)
Myrcia sp. (41614)
C l u a t r e s i as p . ( 4 8 8 0 1 )
Romerillo
San.juanita
Santa maria
Saragosa
Scoba de monte
Taco taco
Tau tarr
Spondiassp. (48801)
Bonafousialongitubulosa(41632)
Anthurium sp. (48717); Araceae
(48ee3)
Lacistemaaggregata(48796)
Ischnosiphonleucophaeus
(48e02)
Swartziasp. (41669)
Clusiaceae(48835)
Dracontium sp. (41597)
Trichomanessp. (41630)
Ericaceae(41470)
Peperomiaomnicola (48986)
Scopariasp. (41431)
Columneafimbricalp<(48923)
Pouteriatorta (41629)
Nautilocalp<sp. (48936)
Aiphanesmacroloba(48916)
Columneapicta (48730);C. rubricalyx(48741);Gloxinia dodsonii
(4872e)
Blakeapunctulata (41613)
Lindernia sp. (48722)
Guzmaniaxamoena (48967)
Pseudelephantopus
puratis
( 4 8 7 13 )
Pitcairniaspectabilis(48972); Rennbergia morreniana (48958);
Sphaeradenia
killipii (48910)
Pepinia hooveri (48975)
Dichorisandrasp. G1489;
Oryctanthusoccidentalis(415I0)
Coletrsx hvbridus (41476)
Peperomiasp. (48719);P.sp.
( 4 8 8 2 r)
Dacryodesgranatensis(41676)
(lhamaedoreapinnatifrons
(48eoB)
Anthurium sp. (41458); Philodendron sp. (48710)
Clolumneaeigantifolia (41587)
Hameliasp. (41608)
Hedychium coronarium (41639);
(48788); Renealmiacrralrecasasii
(41588); R. dolichocalp<(48924)
Coix lacryma-jobi(48716); C. sp.
(41480)
Familyindet. (41665)
Piperaceae(41658);(41590)
Columneanariniana(48815)
Burmeisterasp. (41463)
Hyptis obtusiflora (48720)
Columnea minutiflora (48937)
166
TE
Tedpu
Telba
BS 46
Piperaceae (48726) ; Urticaceae
Ualbura
(48e2e)
Uamuga
L,schrveilerasp. (48995)
Ach,vranthes aspera (41 6a9) ;
( 4 8 8 7 7 ); A l t e r n a n t h e r a p o r r i u c n s
Uande
Uanto
(4 r6 2 0 )
T6o pul uish
Tepalo
Tete
Tigre
Tinta
Tobacccr
Trensa
Trinerol
Tundu
Tunta
Uago
Ualbil
Ualbina
Disterigmasp. (48966); Psarnmisia
clebilis(4896U)
Columnealonginervosa(41507)
E s c h w e i l e rsap . ( 4 1 6 6 8 ) ; Es. p .
(48ee5)
(48991)
Drymonia warszewicziana
Phyllanthtrsanisolobus(4BBB0)
Cavenclishia
ensleriana (48925)
(lalatheatinrothei (4U960)
Menisperrnaceae(18774)
Pollalesta
sp. (41a92)
Monnina sp. (4Utll6)
M i k a n i as p . ( 4 1 5 0 1 )
C r e m o s p e r m lar i r s r ri st s i r rnr m
( 4 1 5 8 1 )P
; i p e rs p . ( a 1 6 3 7 )P
; .s p .
( 4 l l B 3 6I)P .m r r l t i p l i n e r v i t r m
(41471\
Peperomiasp. (41625);Piper sp.
(48909)
Uilde
Uindd
Uish flor
Campylonelrrlrnl sp. (4titt44)
B r u s m a n s i a v e r s i c o l o r ( 4 16 I B )
Cllusiaceae (48799) ; Cllusia dixonii
(48790); Tovomita sp. (41453)
B r t r g m a n s i a v e r s i c o l o r ( 4 16 l f l ) ; D i chorisandra sp. (487C)6)
Columnea laevis (48791)
C e p h a e l i s F i e n t r y i ( 4 14 9 3 )
.|oosiasp. (48969)
Uisit pipa
Columnea sp. (48733); C. bvrsina
( 4 8 7 : tI ) : D r y r r r o r r i aw a r s z e w i c z i a r r a
(48728)
Urnbial
Ur.rgtr
Colurnnea lehmannii (4BgU3)
Blotiella lindenian:r (48847)
Uugu pigail
Veneno
Polybotrl.a lechleriana (48U4U)
Naucleopsis amara (489 I 2)
Verbena
Psetrderantherntrrn micranthunt
Verda
Olf.ersia cervina (48939)
Flacourti:rceae (41 509) ; Solanunr
sp. (48829)
(lucurbitaceae (41 579)
(48e35)
Yalte
Yuyo
Zapata
Zapote
Subniitted to the Academy Februarr' 1995.
Published Ausust 1996.
Solanum conf'ertiseriatum (,1160 1 ) :
(486e5)
M a t i s i a s o e g e n i i ( 4 16 6 0 )