Selbyana 20(2): 201-223. 1999.
CHECKLIST OF BOLIVIAN BROMELIACEAE WITH NOTES ON SPECIES
DISTRIBUTION AND LEVELS OF ENDEMISM
THORSTEN KROMER AND MICHAEL KESSLER
I
Albrecht-von-Haller Institut ffir Pflanzenwissenschaften der Universitat Gottingen,
Abteilung Systematische Botanik, Untere Karsptile 2, Gottingen, D-37073.
E-mail forMK:106606.464@ucompuserve.com
BRUCE
K.
HOLST* AND HARRY
E. LUTHER
The Marie Selby Botanical Gardens, 811 South Palm Ave., Sarasota, FL 34236 USA.
E-mail for BKH: bholst@virtu.sar.usf.edu
ERIC
J.
GOUDA
University Botanic Gardens, P.O. Box 80.162, NL-3508 TD Utrecht, The Netherlands
E-mail: E.J.Gouda@bio.uu.nl
L. I1nsCH
PIERRE
Fundaci6n Amigos de la NaturalezaIBotanical Institute of the University of Bonn, Germany,
Casilla 2241, Santa Cruz, Bolivia. E-mail: pibisch@fan-bo.org
WALTER TILL
Botanical Institute of the University of Vienna, Rennweg 14, A-1030, Wien, Austria
E-mail: walter.till@univie.ac.at
ROBERTO
V A.SQUEZ
Sociedad Boliviana de Botanica, Casilla 3822, Santa Cruz, Bolivia
E-mail: r.vasquez@daitec.scz.com
A discussion of Bromeliaceae diversity in Bolivia, and a checklist of the 21 genera and 281
species occuring there is presented. Each species entry in the checklist includes accepted name, comprehensive synonymy for all species described based on Bolivian types, additional pertinent synonymy, elevation range above sea level, distribution by department, and an indication of which species are endemic
to Bolivia.
ABSTRACT.
RESUMEN.
Aqui se describe la diversidad de la familia Bromeliaceae en Bolivia. Se incluye un listado de
los 21 generos y 281 especies que occurren ahi. Cada especie en ellistado contiene inforrnaci6n del nombre
aceptado, sinonimia comprensiva para cada especie descrita en base a muestras Bolivianas, sinonimia
pertinente adicional, rango de elevaci6n sobre el nivel del mar, distribuci6n por departamento, y se indica
caules especies son endemicas a Bolivia.
Key words:
Bromeliaceae, Bolivia, checklist, endemic
INTRODUCTION
The checklist presented here provides updated
information on the diversity and distribution of
Bromeliaceae in Bolivia. Numerous collections
in the last two decades, especially from fieldwork by S.G. Beck, H. & L. Hromadnik, M.
Kessler, P. Ibisch, R. V iisquez, and T. Kromer,
have resulted in the addition of many new records for the flora since the last summary of
* Corresponding
author.
Bolivian bromeliads (Smith 1969). These include seven genera (Araeococcus, Catopsis,
Deinacanthon, Greigia, Mezobromelia, Pepinia,
and Werauhia) and 125 species. Nevertheless, it
is clear that the record is incomplete, and further
surveys are needed to clarify taxonomic problems and fragmentary distribution records.
METHODS
The following herbaria were surveyed for collections from Bolivia: Herbario Nacional de Bo201
Volume 20(2) 1999
SELBYANA
202
livia, La paz (LPB), Museo de Historia Natural
Noel Kempff, Santa Cruz (USZ), The Marie
Selby Botanical Gardens, Sarasota (SEL), the
Missouri Botanical Garden, St. Louis (MO), Institute of Botany, University of Vienna (WU),
Utrecht Univerisity, Utrecht (U), and the personal Herbarium of R. Vasquez, Santa Cruz
(VASQ). Principal literature references used
were the Flora Neotropica Monograph Bromeliaceae treatments (Smith & Downs 1974, 1977,
1979), a preliminary country checklist by R.
Foster (1958), a country-level family treatment
by L.B. Smith (1969), and a modem epiphyte
survey by P. Ibisch (1996). While an attempt
was made to verify specimens of all taxa included in the list, the occurrence of some species
in Bolivia is only known to us from literature
sources or from identifications made by other
botanists. For these, an indication of the citation
or determiner source is made in the list.
The checklist was compiled from a database
prepared at the Marie Selby Botanical Gardens
which includes approximately 2250 herbarium
records. Author abbreviations follow Brummitt
and Powell (1992), abbreviations ofliterature citations follow Stafteu and Cowan (1976-1988)
for books, and Lawrence et al. (1968) for journals. Herbarium acronyms follow Holmgren et
al. (1990). All basionyms are included, jiS well
as all species-level names based on types from
Bolivia and pertinent recent synonymy. Distribution at the department level in Bolivia and the
elevational range where the species occur are
listed after each species, as well as the type collection, country where the type was collected,
and where possible, an additional voucher specimen. The word "Endemic" is added if the species is wholly confined to the political boundaries of Bolivia.
Nomenclature used in the checklist follows
that used in An Alphabetic list of Bromeliad Binomials (Luther & Sieff 1998) which recognizes
three recently created or resurrected genera: Racinaea (Spencer & Smith 1993), Werauhia
(Grant 1995), and Pepinia (Varadarajan & Gilmartin 1988). Species which formerly belonged
to Streptocalyx can now be found in Aechmea
(Smith & Spencer 1992).
RESULTS AND DISCUSSION
Although Bolivia's species totals still rank below those of other Andean countries (see Holst
1994), the number of genera (21, see TABLE 1)
is second only to Venezuela. This high number
reflects the unique biogeographical position of
Bolivia at the meeting point of the humid northern and central Andes, the dry southern Andes,
TABLE 1. Species numbers of Bromeliaceae in Bolivia, arranged by subfamily and genus.
Subfamily/Genus
PITCAIRNIOIDEAE
Deuterocohnia
Dyckia
Fosterella
Pepinia
Pitcairnia
Puya
Subtotal:
BROMELIOIDEAE
Aechmea
Ananas
Araeococcus
Billbergia
Bromelia
Deinacanthon
Greigia
Pseudananas
Subtotal:
No. species
in Bolivia
11
6
20
1
15
57
110
No. species
endemic to
Bolivia
5
2
15
8
47
77
10
2
1
8
4
1
3
1
30
3
3
7
TILLANDSIOIDEAE
Catopsis
Guzmania
Mezobromelia
Racinaea
Tillandsia
Vriesea
Werauhia
Subtotal:
1
17
2
12
97
9
3
141
31
Family total:
281
119
2
1
35
Amazonia, the Gran Chaco, and the Brazilian
Shield (FIGURE 1).
The 281 recorded species from the country
(TABLE 1; APPENDIX) is still well below the number to be expected. Based on a species/area relationship in comparison to Peru, Ibisch (1996)
estimated a total 300-350 bromeliad species for
Bolivia. Considering the increase in species
numbers in the last decades (on average, about
four new species have been found per year since
1969), this estimate seems realistic. In 1997,
seven new species were described from Bolivia
(Till & Hromadnik 1997, Luther 1997, Ibisch et
al. 1997, Gross 1997, Ehlers 1997, Ehlers &
Hromadnik 1997, Till 1997); in 1998, six new
taxa were described (Luther 1998a, 1998b),
among them, three Greigia species; and in 1999
at least 8 new species have been published
(lbisch et al. 1999, Luther 1999, Ibisch et al.
1999, Kessler et al. 1999). Among the two latter
papers are five new Fosterella species.
Almost 43% of the Bolivian bromeliads are
endemic to the country (TABLE 1). This high
number is due in large part to terrestrial taxa,
KROMER ET AL.: BOLIVIAN BROMELIACEAE
BRAZIL
PERU
CHILE
250 KM
ARGENTINA
< 500 m
500 -1500
co = COCHABAMBA
m
1500 - 3000 m
CH = CHUQUISACA
> 3000 m
TA
セ@ セ@ セ@
-
=TARIJA
Departmental boundaries
FIGURE 1. Bolivia. Political subdivsions and broad
elevational zones.
particularly those in the subfamily Pitcairnioideae which has three genera (Deuterocohnia,
Fosterella, Puya) with centers of diversity in
Bolivia (Ibisch 1996). Eleven out of 15 Deuterocohnia species, 20 of approximately 25 Fosterella species occur in Bolivia, and 47 of the
nearly 200 species of Puya are endemic. Somewhat surprisingly, the number of Pitcairnia species (15) is low compared to other Andean countries. This may be the result of inadequate sam-
203
pIing or the fact that some morphologically and
ecologically variable species (e.g., P. cardenasii,
P. crassa, P. trianae) may include more than
one species.
Another striking feature of the Bolivian bromeliad flora is the low number of large, green,
tank bromeliads in the Tillandsioideae and Bromelioideae in the genera Guzmania, Tillandsia,
Vriesea, and Aechmea. While this is in part due
to inadequate sampling, these groups have their
centers of diversity elsewhere; southeastern Brazil for Aechmea and Vriesea, and the northern
Andes for Guzmania and Tillandsia.
Three departments contain more than 100 species each, La Paz, Cochabamba, and Santa Cruz
(TABLE 2). While these appear to be the best
sampled (ca. 1700 ofthe 2250 collection records
in our database come from there), they are also
the most varied topographically. At the other
end of the sampling spectrum, our database contains only two Bromeliaceae collections from
Oruro and 10 from Pando. While nearly all localities merit further collection for new Bromeliaceae, priorities include the humid mid-elevation forests (1500-2500 m) in La Paz, semi-humid forests at the transition of dry, rainshadowed valleys to humid montane forest in almost
any valley system in La Paz and adjacent Cochabamba, and the interior portions of Ambor6
National Park in Santa Cruz. Although bromeliads occur at low densities in the Amazonian
lowlands, a thorough search in Pando and the
Amazonian part of Beni should yield elements
of the Brazilian bromeliad flora which are not
yet known from Bolivia. Likewise, the dry Chaco lowland and foothill forests of Chuquisaca
and Tarija are likely to yield new discoveries. In
most need of study taxonomically are the terrestrial and saxicolous genera, particularly Puya
and Pitcairnia, and the largely epiphytic genus
Tillandsia.
2. Bromeliaceae species diversity, number of department- and country-level political endemic species,
and number of collection records in database used.
TABLE
No. BoHvian
Department
La Paz
Santa Cruz
Cochabamba
Chuquisaca
Tarija
PotOSI
Beni
Pando
Oruro
nッセ@
species
154
119
115
65
63
30
28
6
2
No. species wholly
endemic to each
department
endemic species
23
20
14
8
12
3
45
37
35
24
f'Ound in each
department
18
12
2
No. collection
records in database
714
575
417
220
146
78
67
10
2
SELBYANA
204
The database resulting from this project is
available to all those who want to contribute further records or use it for purposes of research or
conservation. Please contact B. Holst or M.
Kessler for more information.
ACKNOWLEDGMENTS
We thank the following herbaria for providing
specimens and data used in this work: LPB, MO,
SCZ, SEL, U, and WU. The studies of P. lbisch
and R. Vasquez partially were conducted under
the Research and Development program of the
Science Department of Fundaci6n Amigos de la
Naturaleza Noel Kempff, Bolivia as part of the
Noel Kempff Climate Action Project.
LITERATURE CITED
Brummitt, R.K. and C.E. Powell, eds. 1992. Authors
of Plant Names. Royal Botanic Gardens, Kew,
London.
Ehlers, R. 1997. Tillandsia colganii, a new species. J.
Brome1iad Soc. 47: 103-106.
Ehlers, R. and L. Hromadnik. 1997. Tillandsia hasei
R. Ehlers & L. Hromadnik spec. nov. Die Bromelie 1996(3): [74-75, figs.] 76-78 [backcover].
Foster, R.C. 1958. A catalogue of the ferns and flowering plants of Bolivia. Contrib. Gray Herb. Harvard Univ. 184: 1-223.
Grant, J.R. 1995. The resurrection of Alcantarea and
Werauhia, a new genus. Trop. Subtrop. Pflanzenwelt 91: 16-17.
Gross, E. 1997. Bromelienstudien. 1. Neue und wenig
bekannte Arten aus Peru und anderen Liindern
(23. Mitteilung). Trop. Subtrop. Pflanzenwelt 95:
20-21,39.
Holmgren, P.K., N.H. Holmgren and L.c. Barnett.
1990. Index Herbariorium. Part I: The Herbaria of
the World, ed. 8. New York Botanical Garden,
Bronx, New York.
Holst, B.K. 1994. Checklist of Venezuelan Bromeliaceae with notes on species distribution by state
and levels of endemism. Selbyana 15: 132-149.
Ibisch, P.L. 1996. Neotropische Epiphytendiversitiitdas Beispiel Bolivien. Archiv naturwissenschaftlicher Dissertationen 1: I-XIV, 1-357 + 107 pp.,
Martina Galunder-Verlag, Wiehl.
!bisch, P.L., E. Gross, G. Rauer & D. Rudolph 1997.
On the diversity and biogeography of the genus
Fosterella L.B. Sm., Bromeliaceae, with the description of a new species from eastern Bolivia. J.
Bromeliad Soc. 47: 211-217.
!bisch, P.L., R. Vasquez Ch. and E. Gross. 1999. New
species of Puya and Pitcairnia (Bromeliaceae)
from the Ambor6 National Park and vicinity (Santa Cruz, Bolivia)-a neglected centre of diversity
and endemism of Pitcairnoideae. J. Bromeliad
Soc. 49: 124-135.
Volume 20(2) 1999
!bisch, P.L., R. Vasquez and E. Gross. 1999. More
novelties of Fosterella L.B. Smith (Bromeliaceae,
Pitcairnioideae) from Bolivia. Rev. Soc. Bo1iviana
Bot. 2(2): 117-132.
Kessler, M., P.L. Ibisch and E. Gross. 1999. Fosterella
cotacajensis, una nueva especie de Bromeliaceae
de los valles secos andinos de Bolivia. Rev. Soc.
Boliviana Bot. 2(2): 111-116.
Lawrence, G.H.M., A.EG. Buchheim, G.S. Daniels
and H. Dolezal, eds. 1968. Botanico-PeriodicumHuntianum. Hunt Botanical Library, Pittsburgh.
Luther, H.E. 1997. A showy new Fosterella from Bolivia. J. Bromeliad Soc. 47: 118-119.
- - - . 1998a. Miscellaneous new taxa of Bromeliaceae (XII). Selbyana 19(1): 83-90.
- - - . 1998b. Miscellaneous new taxa of Brome1iaceae (XIII). Se1byana 19(2): 218-226.
- - - . 1999. Miscellaneous new taxa of Bromeliaceae (XIV). Selby ana 20(1): 10-15.
Luther, H.E. and E. Sieff. 1998. An Alphabetic List of
Bromeliad Binomials, 5th ed. The bイッュセャゥ。、@
Society, International, Newberg.
Ribera, M.O., M. Libermann, S. Beck & M. Moraes.
1996. Vegetaci6n de Bolivia. Pp. 169-222 in K.
Mihotek, ed. Comunidades, Territorios Indfgenas
y Biodiversidad en Bolivia. CIMAR-U.A.G.R.M.,
Santa Cruz, Bolivia.
Smith, L.B. 1969. The Bromeliaceae of Bolivia. Rhodora 71: 35-57,220-279.
Smith, L.B. and R.J. Downs. 1974. Pitcairnioideae
(Bromeliaceae). Fl. Neotrop. Monogr. 14(1): 1660. Hafner Press, New York.
- - - . 1977. Tillandsioideae (Brome1iaceae). FI.
Neotrop. Monogr. 14(2): 663-1492. Hafner Press,
New York.
---.1979. Bromelioideae (Bromeliaceae). Fl. Neotrop. Monogr. 14(3): 1493-2142. New York Botanical Garden, New York.
Smith, L.B. and M.A. Spencer. 1992. Reduction of
Streptocalyx (Bromeliaceae: Brome1ioideae).
Phytologia 72: 96-98.
Spencer, M.A. and L.B. Smith. 1993. Racinaea, a new
genus of Bromeliaceae (Tillandsioideae). Phytologia 74: 151-160.
Stafleu, EA. and R.S. Cowan. 1976-1988. Taxonomic
Literature: A Selective Guide to Botanical Publications and Collections with Dates, Commentaries
and Types, ed. 2. 7 volumes. Utrecht, Antwerpen,
The Hague, and Boston.
Till, W. 1997. Tillandsia samaipatensis, eine iibersehene Art aus Zentralbolivien. Die Bromelie 21
1996: 36-38.
Till, W. and L. Hromadnik. 1997. Uma Nova Especie
Incomum de Deuterocohnia da Bolfvia. An unusual new Deuterocohnia from Bolivia. Bromelia
4(1): 19-22.
Varadarajan, G.S. and A.J. Gilmartin. 1988: Taxonomic realignments within the subfamily Pitcairnioideae (Bromeliaceae). Syst. Bot. 13: 294-299.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
205
ApPENDIX
Checklist of Bolivian Bromeliaceae
Department abbreviations. BE-Beni; CH-Chuquisaca; CO-Cochabamba; LP-La paz;
OR-Oruro; PA-Pando; PO-Potosi; TA-Tarija.
Type designations. HT-Holotype; IT-Isotype; LT-Lectotype; ILT-Isolectotype; CT-Clonotype.
Endemic = confined to the political boundaries of Bolivia.
AECHMEA
Voucher: M. Kessler 3791 (LPB, SEL).
Endemic.
Aechmea angustifolia Poepp. & Endl., Nov. gen.
sp. PI. 2: 43. 1838.
Synonym(s): Aechmea boliviana Rusby, Bull. New
York Bot. Gard. 4: 456. 1907; Aechmea inconspicua Harms, Notizbl. Bot. Gart. Berlin-Dahlem 10: 786. 1929.
Dept.: BE, CO, LP, PA, SC.
Elevation: 150-1300 m.
Type: Poeppig 1823. HT: W (lost). Peru.
Voucher: M. Kessler et al. 8457 (LPB, SEL).
Aechmea longifolia (Rudge) L.B. Sm. & M.A.
Spencer, Phytologia 72: 97. 1992. Basionym:
Bromelia longifolia Rudge, Guyan. I: 31, pI.
49. (1805) 1806.
Synonym(s): Streptocalyx longifolius (Rudge) Baker, Handb. Bromel. 31. 1889.
Dept.: CO.
Elevation: 300-500 m.
Type: Martin s.n. HT: BM. French Guiana.
Voucher: M. Kessler et al. 8538 (LPB, SEL).
Aechmea bromel;ifolia (Rudge) Baker, in Benth. &
Hook. f., Gen. pI. 3: 664. 1883. Basionym: Tillandsia bromeliaefolia Rudge, PI. Guian. 32, t.
50. 1807.
Synonym(s): Aechmea ellipsoidea Rusby, Mem.
New York Bot. Gard. 7: 212. 1927.
Dept.: BE, CH, LP, SC.
Elevation: 200-1350 m.
Type: Martin S.n. HT: BM. French Guiana.
Voucher: J. Solomon 6179 (SEL).
Aechmea mertensii (G. Mey.) Schult. & Schult.f.,
in Roem. & Schult., Syst. veg. 7(2): 1272.
Aechmea castelnavii Baker, Handb. Bromel. 39.
1889.
Dept.: SC.
Elevation: 300-350 m.
Type: Castelnau s.n. HT: P. Bolivia.
Voucher: In Bolivia, only known from type collection.
Aechmea distichantha Lem., Jard. Fleur. 3: t. 269.
1853.
Synonym(s): Platyaechmea distichantha (Lem.)
L.B. Sm. & Kress, Phytologia 69: 272. 1990.
Dept.: CH, CO, LP, SC, TA.
Elevation: 200-1900 m.
Type: Libon s.n. HT: BR? Brazil.
Voucher: B. Holst et al. 4948 (USZ, SEL).
Aechmea fuerstenbergii E. Morren & Wittm., Belgique Hort. 29: 42. 1879.
Synonym(s): StreptocalyxJuerstenbergii (E. Morren
& Wittm.) E. Morren, Belgique Hort. 33: 16.
1883.
Dept.: BE, LP.
Elevation: 300-850(-1050?) m.
Type: Porte S.n. HT: LG. Brazil.
Voucher: T. Kromer (GOET, LPB, SEL).
Aechmea kuntzeana Mez, in C. DC., Monogr.
phan. 9: 208. 1896.
Dept.: BE, LP, SC.
Elevation: 150-1000 m.
Type: Kuntze s.n. HT: NY; IT: US. Bolivia.
1830.
Basionyrn:
Bromelia mertensii Mey.,
Prim. fl. esseq. 144. 1818.
Dept.: BE, PA.
Elevation: ca. 200 m.
Type: E.K. Rodschied S.n. HT: GOET. Guyana.
Voucher: S. Beck et al. 19320 (SEL).
Aechmea setigera Mart. ex Schult. & Schult.f., in
Roem. & Schult., Syst. veg. 7(2): 1273. 1830.
Dept.: BE, SC.
Elevation: 150-400 m.
Type: C. Martius s.n. HT: M. Brazil.
Voucher: S. Beck et al. 5978 (LPB, SEL).
Aechmea tocantina Baker, Handb. Bromel. 39.
1889.
Dept.: BE, SC.
Elevation: 250-500 m.
Type: Weddell 2365. HT: P. Brazil.
Voucher: R. Vasquez et al. 1813 (VASQ).
ANANAS
Ananas ananassoides (Baker) L.B. Sm., Bot. Mus.
Leafl. 7: 70. 1939. Basionym: Acanthostachys
ananassoides Baker, Handb. Bromel. 25. 1889.
Dept.: BE, LP, PA, Sc.
Elevation: 150-900 m.
Type: LT: Regnell (Ill) 1261 in part. HT: P. Brazil.
Voucher: M. Moriies et al. 1633 (SEL).
Ananas comosus (L.) Merr., Interpr. Herb. amboin.
133. 1917. Basionym: Bromelia comosa L.,
Herb. ambo 21. 1754.
Dept.: Widely cultivated.
Elevation: range unknown.
Type: Type not yet selected.
Voucher: M. Lewis 37568 (SEL).
SELBYANA
206
ARAEOCOCCUS
Araeococcus ftagellifolius Harms, Notizbl. Bot.
Gart. Berlin-Dahlem 10: 784. 1929.
Dept.: SC.
Elevation: 200-250 m.
Type: Huebner 58. HT: B. Brazil.
Voucher: A. Jardim et at. 53 (SEL).
BILLBERGIA
Billbergia brasilieusis L.B. Sm., Arq. Bot. Estado
Sao Paulo II. 1: 105. 1943. Basionym: Helicodea leopoldii Versch. Hort ex Lem., Ill. Hort.
11: sub pi. 421. 1864.
Synonym(s): Billbergia leopoldii (Versch. Hort. ex
Lem.) Linden ex Houllet, Rev. Hort. 41: 87.
1869; non K. Koch 1857.
Dept.: Unknown.
Elevation: Unknown.
Type: De Vos in Verschaffett Hortus s.n. Description and illustration. Brazil.
Voucher: D. Beadle SU 126 (SEL).
Billbergia cardenasii L.B. Sm., Phytologia 4: 382.
1953.
Dept.: CO?
Elevation: Unknown.
Type: Cardenas 4907. HT: US. Bolivia.
Voucher: D. Beadle SU 029 (SEL).
Endemic.
Billbergia decora Poepp. & EndL, Nov. gen. sp. pI.
2: 42, t. 157. 1838.
Synonym(s): Billbergia boliviensis Baker, Handb.
Bromel. 81. 1889.
Dept.: LP.
Elevation: 2400 m.
Type: Poeppig s.n. HT: W (lost). Peru.
Voucher: H. Rusby 2853 (NY). Cited for Bolivia in:
Fi. Neotrop. Monogr. 14(3): 2021. 1979.
Billbergia meyeri Mez, Bot. Jahrb. Syst. 30: 148.
1902.
Dept.: Sc.
Elevation: 150-450 m.
Type: Pilger 705. HT: B. Brazil.
Voucher: R. Guillen 1910 (SEL).
Billbergia microlepis L.B. Sm., Phytologia 8: 504.
1963.
Dept.: CO, LP.
Elevation: 700-1500 m.
Type: B. Krukoff 11158. HT: GH. Bolivia.
Voucher: A. Gentry et al. 44301 (SEL).
Endemic.
Billbergia robert-readii E. Gross & Rauh, Trop.
SUbtrop. Pflanzenwelt 60: 89. 1987.
Dept.:BE,LP.
Elevation: 600-700 m.
Type: Rauh 26163. lIT: HEID. Peru.
Voucher: T. Kromer 137 (GOET, LPB, SEL).
BiUbergia velascana M. cardenas, Bull. Bromeliad
Soc. 7: 35. 1957.
Dept.: SC.
Elevation: 250-850 m.
Volume 20(2) 1999
Type: Father Lorenzo Hammerschmidt 5486. HT:
Cardenas Herbarium (LlL ?). Bolivia.
Voucher: D. Beadle SU 029A (SEL).
Endemic.
BiUbergia violacea Beer, Fam. Bromel. 116. 1856.
Dept.: CO, SC.
Elevation: 200-1600 m.
Type: Appun S.n. HT: B? "Brazil" Guyana.
Voucher: R. Vasquez & Ric 1845 (VASQ).
Bromelia balansae Mez, in Mart., FI. bras. 3(3):
191. 1891.
Dept.: BE, Sc.
Elevation: 150-800 m.
Type: Balansa 608. HT: G. Paraguay.
Voucher: S. Beck 20689 (LPB, SEL).
BROMELIA
Bromelia hieronymi Mez, in Mart., Fl. bras. 3(3):
199. 1891.
Dept.: SC, TA.
Elevation: 250-700 m.
Type: Hieronymus & Lorentz s.n. HT: B; IT: US.
Argentina.
Voucher: M. Nee & G. Coimbra S. 39933 (LPB).
Bromelia serra Griseb., Abh. Konigi. Ges. Wiss.
Gottingen 24: 328. 1879.
Dept.: BE, LP, SC.
Elevation: 150-1400 m.
Type: Lorentz & Hieronymus s.n. LT: GOET. Argentina.
Voucher: M. Moriies et at. 1631 (LPB, SEL).
Bromelia vilIosa Mez, Bot. Jahrb. Syst. 30 (BeibI.
67): 3. 1901.
Dept.: SC.
Elevation: 400 m.
Type: Glaziou 22191. HT: B. Brazil.
Voucher: Fuentes 1148 (USZ).
CATOPSIS
Catopsis sessilillora (Ruiz & Pav.) Mez, in C. DC.,
Monogr. phan. 9: 625. 1896. Basionym: Tillandsia sessiliflora Ruiz & Pav., Fl. peruv. 3:
42, pl. 271, fig. b. 1802.
Dept.: LP.
Elevation: 1250-1300 In.
Type: Ruiz & Pavon S.n. HT: M; IT: B. Peru.
Voucher: M. Kessler et al. 10320 (LPB, SEL).
DEINACANTHON
Deinacantbon urbanianum (Mez) Mez, in C. DC.,
Monogr. phan. 9: 13. 1896. Basionym: Rhodostachys urbaniana Mez, in Mart., Fl. bras. 3(3):
182. 1891.
Dept.: sc.
Elevation: 400 m.
Type: Hieronymus s.n. HT: G; IT: B, CORD, GH,
US. Argentina.
Voucher: G. Navarro 2221 (USZ). Det. G. Navarro.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
DEUTEROCOHNIA
Deuterocohnia bracteosa W. Till & L. Hrom., Bromelia 4(1): 19. 1997.
Dept.: CH.
Elevation: 3000 m.
Type: L. Hromadnik 19029. HT: WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Deuterocohnia brevifolia (Griseb.) M.A. Spencer
& L.B. Sm., Bradea 6: 144. 1993. Basionym:
Navia brevifolia Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 24: 332. 1879.
Synonym(s): Abromeitiella brevifolia (Griseb.) A.
Cast., Anales Mus. Nac. Hist. Nat. Buenos Aires 36: 371. 1931; Abromeitiella pulvinata
Mez, Bot. Archiv. 19: 460. 1927.
Dept.: TA.
Elevation: 1700-2400 m.
Type: Lorentz & Hieronymus 947 (in part). HT:
GOET; IT: CORD. Argentina.
Voucher: S. Beck 14276 (SEL).
Deuterocohnia brevispicata Rauh & L. Hrom.,
Trop. SUbtrop. Pflanzenwelt 65: 5. 1988.
Dept.: CH.
Elevation: 1200-1300(-2050) m.
Type: Hromadnik 5213. HT: HElD. Bolivia.
Voucher: D. Cathcart B52 (SEL).
Endemic.
Deuterocohnia digitata L.B. Sm., Phytologia 18:
137. 1969.
Dept.: CO, PO, SC, TA.
Elevation: 1900-2750 m.
Type: Castellanos s.n. HT: BA; IT: US. Argentina.
Voucher: 1. Vargas 3185 (SEL).
Deuterocohnia glandulosa E. Gross, Trop. Subtrop.
Pflanzenwelt 75: 5. 1990.
Dept.: TA.
Elevation: 1200 m.
Type: Hromadnik 5167. HT: HElD. Bolivia.
Voucher: Only known from type collection.
Endemic.
Deuterocohnia longipetala (Baker) Mez, in Mart.,
Fl. bras. 3(3): 506. 1894. Basionym: Dyckia
longipetala Baker, Handb. Bromel. 135. 1889.
Dept.: CO, SC, TA.
Elevation: 300-2100 m.
Type: Humboldt & Bonpland 3595. HT: B. Peru.
Voucher: J. Balcazar 100 (USZ, SEL).
207
Abromeitiella lotteae Rauh, Trop. Subtrop.
Pflanzenwelt 42: 5. 1983.
Dept.: TA.
Elevation: 2700 m.
Type: Hromadnik 5131. HT: HElD. Bolivia.
Voucher: W. Till 62 A (WU).
Endemic.
Deuterocohnia meziana Kuntze ex Mez, in C. DC.,
Monogr. phan. 9: 465. 1896.
Dept.: SC.
Elevation: 200-1350 m.
Type: O. Kuntze s.n. HT: NY. Brazil.
Voucher: M. Nee 35601 (LPB).
Deuterocohnia scapigera (Rauh & L. Hrom.) M.A.
Spencer & L.B. Sm., Bradea 6: 141-145. 1993.
Basionym: Abromeitiella scapigera Rauh & L.
Hrom., Trop. SUbtrop. Pflanzenwelt 60: 5.
1987.
Dept.: CH, PO.
Elevation: 1200-2400 m.
Type: Hromadnik 5275. HT: HElD. Bolivia.
Voucher: M. Kessler et at. 5085 (LPB, SEL).
Endemic.
Deuterocohnia strobilifera Mez, Repert. Spec.
Nov. Regni Veg. 3: 15, 1906.
Dept.: CH, PO, TA.
Elevation: 2500-3550 m.
Type: Fiebrig 2933. HT: B. Bolivia.
Voucher: S. Beck 14122 (LPB). Det. G. Varadarajan.
DYCKIA
Dyckia crassifolia Rauh, Trop. Subtrop. Pflanzenwelt 79: 10. 1991.
Dept.: SC.
Elevation: 1200 m.
Type: W. Rauh 40670. HT: HElD. Bolivia.
Voucher: Only known from type collection.
Endemic.
Dyckia ferox Mez, in C. DC., Monogr. phan. 9:
511. 1896.
Synonym(s): Dyckia meziana 0. Kuntze, Revis.
gen. pI. 3(2): 302. 1898.
Dept.: SC.
Elevation: 1200 m.
Type: Kuntze s.n. HT: NY. Paraguay.
Voucher: M. Cardenas 2679 (GH). Cited for Bolivia
in: Fl. Neotrop. Monogr. 14(1): 543. 1974.
Dyckia ferrnginea Mez, in C. DC., Monogr. phan.
9: 533. 1896.
Dept.: Sc.
Elevation: 300 m.
Type: Kuntze S.n. HT: NY. Brazil.
Voucher: P. lbisch et ai. 93.0614 (FR, LPB, SEL).
Deuterocohnia lorentziana (Mez) M.A. Spencer &
L.B. Sm., Bradea 6: 144. 1993. Basionym: Pitcaimia iorentzina Mez, in C. DC., Monogr.
phan. 9: 974. 1896.
Synonym(s): Abromeitiella lorentziana (Mez) A.
Cast., Lilloa 10: 459. 1944.
Dept.: PO, TA.
Elevation: 2650-3150 m.
Type: Lorentz s.n. HT: B. Argentina.
Voucher: W. Till 62a (WU).
Dyckia gracilis Mez, in C. DC., Monogr. phan. 9:
516. 1896.
Dept.: Sc.
Elevation: 200-700 m.
Type: D'Orbigny 1006. HT: P; IT: G. Bolivia.
Voucher: A. Fuentes 1151 (LPB, USZ).
Deuterocohnia lotteae (Rauh) M.A. Spencer &
L.B. Sm., Bradea 6: 145. 1993. Basionym:
Dyckia leptostachya Baker, Gard. Chron. 1884(2):
198. 1884.
SELBYANA
208
Synonym(s): Dyckia boliviensis Mez in C. DC.,
Monogr. phan. 9: 524. 1896.
Dept.: CH, sc.
Elevation: 350-500(-1850?) m.
Type: Kew Hortus. HT: K. Brazil.
Voucher: T. Killeen et al. 7030 (SEL).
Dyckia pulquinensis Wittm., Rijks-Herb. 29: 88.
1916.
Dept.: SC.
Elevation: 1500 m.
Type: Herzog 1849. HT: L; IT: B. Bolivia.
Voucher: G. Balcazar 35 (USZ).
Endemic.
FOSTERELLA
Fosterella albicans (Griseb.) L.B. Sm., Phytologia
7: 171. 1960. Basionym: Cottendorfia albicans
Griseb., Abh. Konigl. Ges. Wiss. Gottingen 24:
330. 1879.
Synonym(s): Lindmania alibcans -(Griseb.) Mez in
C. DC., Monogr. phan. 9: 537. 1896.
Dept.: CH, CO, LP, SC.
Elevation: 900-2150 m.
Type: Lorentz & Hieronymus 502. HT: GOET; IT:
B, CORD. Argentina.
Voucher: B. Holst et al. 4837 (USZ, SEL).
Fosterella cauIescens Rauh, Trop. SUbtrop. Pflanzenwelt 31: 23. 1979.
Dept.: LP.
Elevation: 1200 m.
Type: W. Rauh 40579A. HT: HEID. Bolivia.
Voucher: Only known from type collection.
Endemic.
Fosterella chaparensis Ibisch, Vasquez & Gross,
Rev. Soc. Boliviana Bot. 2(2): 118. 1999.
Dept.: CO.
Elevation: 600 m.
Type: R. Vasquez & M.l. Vasquez 2792. HT: LPB;
IT: FR, USZ. Bolivia.
Voucher: Only known from type collection.
Endemic.
Fosterella chiquitana Ibisch, Vasquez & Gross,
Rev. Soc. Boliviana Bot. 2(2): 118. 1999.
Dept.: Sc.
Elevation: 500 m.
Type: P. Ibisch 98.0125. HT: LPB; IT: USZ. Bolivia.
Voucher: Only known from type collection.
Endemic.
Fosterella cotacajensis M. Kessler, P. Ibisch & E.
Gross. Rev. Soc. Boliviana Bot. 2(2): 111.
1999.
Dept.: CO, LP.
Elevation: 2000-2200 m.
Type: Kessler et al. 9620. HT: LPB; IT: SEL. Bolivia.
Voucher: M. Kessler et al. 5531 (LPB).
Endemic.
Fosterella elata H. Luther, Selbyana 5: 310. 1981.
Dept.: LP.
Elevation: 900-1500 m.
Volume 20(2) 1999
Type: Luer et al. s.n. HT: SEL; IT: US. Bolivia.
Voucher: M. Kessler et al. 5726 (LPB, SEL).
Endemic.
Fosterella floridensis Ibisch, Vasquez & Gross,
Rev. Soc. Boliviana Bot. 2(2): 120. 1999.
Dept.: SC.
Elevation: 1100-1450 m.
Type: 1bisch & Ibisch 97.83. HT: LPB; IT: FR, USZ.
Bolivia.
Voucher: Vasquez 1808 (JBSCZ).
Endemic.
Fosterella gracilis (Rusby) L.B. Sm., Phytologia 7:
171. 1960. Basionym: Catopsis gracilis Rusby,
Bull. New York Bot. Gard. 6: 489. 1910.
Synonym(s): Lindmania gracilis (Rusby) L.B. Sm.,
Contr. Gray Herb 104: 78. 1934.
Dept.: LP.
Elevation: 500 m.
Type: R.S. Williams 738. HT: NY. Bolivia.
Voucher: White 1086 (NY). Cited in FI. Neotrop.
Monogr. 14(1): 209. 1974.
Endemic.
Fosterella graminea (L.B. Sm.) L.B. Sm., Phytologia 7: 171. 1960. Basionym: Lindmania graminea L.B. Sm., Lilloa 14: 93. 1948.
Dept.: LP.
Elevation: 7oo-900(-1450?) m.
Type: Buchtien 417. HT: US. Bolivia.
Voucher: M. Kessler 4100 (LPB, SEL).
Endemic.
Fosterella heterophylla Rauh, Trop. Subtrop.
Pflanzenwelt 60: 24. 1987.
Dept.: LP.
Elevation: 1000 m.
Type: W. Rauh 40583A. HT: HEID. Bolivia.
Voucher: Only known from type collection.
Endemic.
Fosterella latifolia Ibisch, Vasquez & Gross, Rev.
Soc. Bol. Bot. 2(2): 123. 1999.
Dept.: SC.
Elevation: 450-1000 m.
Type: P. Ibisch 98.0098. HT: LPB; IT: FR, USZ.
Bolivia.
Voucher: Ibisch 98.0173 (USZ).
Endemic.
Fosterella pearcei (Baker) L.B. Sm., Phytologia 7:
172. 1960. Basionym: Cottendorfia pearcei
Baker, Handb. Bromel. 128. 1889.
Synonym(s): Lindmania pearcei (Baker) Mez in C.
DC., Monogr. phan. 9: 537. 1896.
Dept.: LP.
Elevation: 750-900 m.
Type: Pearce s.n. HT: BM. Bolivia.
Voucher: B. Krukoff 10328 (GH, LPB, NY). Cited
for Bolivia in Fl. Neotr. 14(1): 202. 1974.
Endemic.
Fosterella penduliflora (C.H. Wright) L.B. Sm.,
Phytologia 7: 172. 1960. Basionym: Catopsis
pendulifiora C.H. Wright, Bull. Misc. Inform.
1910: 197. 1910.
Synonym(s): Lindmania pendulifiora (C.H. Wright)
Stapf, Bot. Mag. 150: pi. 9029. 1924.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Dept.: CH, LP, SC, TA.
Elevation: 550-1450 m.
Type: Forget S.n. HT: K. Peru.
Voucher: M. Kessler et at. 4214 (LPB, SEL).
Fosterella rojasii (LB. Sm.) LB. Sm., Revista Argent. Agron. 7: 162. 1940. Basionym: Lindmania rojasii LB. Sm., Phytologia 7: 172.
1960.
Dept.: Sc.
Elevation: 450-500 m.
Type: Rojas 6771. HT: GH. Paraguay.
Voucher: H. Amerhauser 5 (WU).
Fosterella rusbyi (Mez) LB. Sm., Phytoiogia 7:
172. 1960. Basionym: Lindmania rushyi Mez,
Bot. J ahrb. Syst. 30 (BeibL 67): 6. 1901.
Dept.: LP.
Elevation: 850-1500(-1800?) m.
Type: Bang 2571. HT: B; IT: BM, F, G, GH, K, NY,
US. Bolivia.
Voucher: M. Lewis 36936 (MO, SEL). "cf.".
Fosterella schidosperma (Baker) LB. Sm., Phytologia 8: 500. 1963. Basionym: Chlorophytum
schidospermum Baker, J. Linn. Soc., Bot. 15:
326. 1876.
Synonym(s): Cottendorfia rushyi Baker, BulL Torrey Bot. Club 29: 697. 1902, non Lindmania
rusbyi Mez, 1901; Lindmania schidosperma
(Baker) P. Ravenna, Wrightia 7(3): 253. 1983.
Dept.: CO, LP.
Elevation: 750-2000 m.
Type: Lechler 2382. HT: CGE?; IT: G? Peru.
Voucher: M. Kessler et at. 7786 (LPB, SEL). "cf.".
Fosterella spectabilis H. Luther, J. Bromeliad Soc.
47: 118. 1997.
Dept.: Sc.
Elevation: 1700 m.
Type: Cathcart B 17. HT: SEL; IT: LPB, MO, NY,
US, WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Fosterella vasquezii E. Gross & P. Ibisch, J. Bromeliad Soc. 47: 212. 1997.
Dept.: Sc.
Elevation: 200-850 m.
Type: Ibisch et at. 930652. HT: HElD; IT: LPB.
Bolivia.
Voucher: F. Mamani & A. Jardim 126J(USZ, SEL).
Endemic.
Fosterella villosula (Harms) LB. Sm., Phytologia
7: 172. 1960. Basionym: Lindmania villosula
Harms, NotizbL Bot. Gart. Berlin-Dahlem 10:
794. 1929.
Dept.: CO, LP, Sc.
Elevation: 800-1600 m.
Type: Werdermann 2120. HT: B. Bolivia.
Voucher: M. Kessler et at. 4109 (LPB, SEL).
Endemic.
FostereHa weddelliana (Brongn. ex Baker) LB.
Sm., Phytologia 7: 172. 1960. Basionym: Cottendorfia weddelliana Brongn. ex Baker,
Handb. BrameL 129. 1889.
Synonym(s): Lindmania weddelliana (Brongn. ex
209
Baker) Mez in C. DC., Monogr. phan. 9: 538.
1896.
Dept.: LP.
Elevation: 900-1800 m.
Type: Weddell 4233. HT: P. Bolivia.
Voucher: M. Lewis 40425 (LPB, MO, SEL).
Endemic.
GREIGIA
Greigia atrocastanea H. Luther, Selbyana 19: 218.
1998 [1999].
Dept.: LP.
Elevation: (2500?-)3300 m.
Type: E. Garcia 1977. HT: SEL; IT: LPB. Bolivia.
Voucher: M. Kessler et at. 12135 (LPB, SEL).
Endemic.
Greigia cochabambae H. Luther, Selbyana 19: 83.
1998.
Dept.: CO, LP.
Elevation: 2100-2700 m.
Type: M. Kessler et al. 7111. HT: SEL; IT: LPB.
Bolivia.
Voucher: M. Kessler et at. 12132 (LPB, SEL).
Endemic.
Greigia kessleri H. Luther, Selbyana 19: 83. 1998.
Dept.: CO, LP.
Elevation: 2200-3250 m.
Type: M. Kessler et al. 9067. HT: SEL; IT: LPB.
Bolivia.
Voucher: M. Kessler et at. 10519 (LPB, SEL).
Endemic.
GUZMANIA
Guzmania besseae H. Luther, Selbyana 5: 310.
1981.
Dept.: CO, LP, Sc.
Elevation: 1950-2600 m.
Type: L. Besse et at. 669. HT: SEL; IT: US. Bolivia.
Voucher: P. lbisch & c. 1bisch 93.0111 (LPB, SEL).
Guzmania bipartita L.B. Sm., Phytologia 6: 437.
1959.
Dept.: LP, Sc.
Elevation: 1200-1500 m.
Type: Asplund 19474. HT: S. Ecuador.
Voucher: M. Kessler et al. 10092 (LPB, SEL).
Guzmania calothyrsus Mez, in C. DC., Monogr.
phan. 9: 910. 1896.
Dept.: BE, CO, LP, SC.
Elevation: 200-1300 m.
Type: Poeppig 1224. HT: W (lost). Peru.
Voucher: T. Kriimer et at. 57 (LPB, SEL).
Guzmania danielii L.B. Sm., Phytologia 4: 360.
1953.
Dept.: CO, LP?
Elevation: 1700-2700 m.
Type: Daniel 3350. HT: OH; IT: US. Colombia.
Voucher: M. Kessler et al. 9027 (LPB, SEL).
Guzmania gioriosa (Andre) Andre ex Mez, in C.
210
Volume 20(2) 1999
SELBYANA
DC., Monogr. phan. 9: 922. 1896. Basionym:
Caraguata gloriosa Andre, Enum. BromeI. 5.
13 Dec. 1888; Rev. Hort. 60: 565. 16 Dec.
1888.
Dept.: CO, LP.
Elevation: 1300-1600 m.
Type: Andre 3791. HT: K. Ecuador.
Voucher: M. Kessler et al. 7446 (LPB, SEL).
Guzmania killipiana L.B. Sm., Contr. Gray Herb.
98: 29. 1932.
Dept.: CO, LP.
Elevation: 1150-2500 m.
Type: Killip & Smith 25630. HT: US. Peru.
Voucher: M. Kessler et at. 6957 (LPB, SEL).
Guzmania Jingulata (L.) Mez, in C. DC., Monogr.
phan. 9: 899. 1896. Basionym: Tillandsia lingulata L., Sp. pI. 1: 286. 1753.
Dept.: LP, SC.
Elevation: 500-1450 m.
Type: Plumier s.n. HT: P. West Indies.
Voucher: M. Saldias & L. Arroyo 3572 (SEL).
Guzmania marantoidea (Rusby) H. Lutber, Selbyana 19: 86. 1998. Basionym: Tillandsia marantoidea Rusby, Descr. S. Amer. pI. 4. 1920.
Synonym(s): Guzmania rio-nievensis Rauh, Trop.
Subtrop. Pflanzenwelt 50: 22. 1984.
Dept.: CO, LP.
Elevation: 1300-2650 m.
Type: Bang 2570. HT: NY; IT: US. Bolivia.
Voucher: M. Kessler et at. 7843 (LPB, SEL).
Guzmania melinonis Regel, Gartenfiora 34: 116.
1885.
Dept.: CO, LP, Sc.
Elevation: 300-1200 m.
Type: Linden in Petrograd Hortus s.n. HT: LE.
French Guiana.
Voucher: M. Kessler et al. 8058 (LPB, SEL).
Guzmania monostachia (L.) Rusby ex Mez, in C.
DC., Monogr. phan. 9: 905. 1896. Basionym:
Renealmia monostachia L., Sp. pI. 1: 287.
1753.
Dept.: LP.
Elevation: 1300 m.
Type: Plumier S.n. HT: P. West Indies.
Voucher: M. Kessler et al. 10305 (LPB, SEL).
Guzmania morreniana (Linden Hortus) Mez, in C.
DC., Monogr. phan. 9: 932. 1896. Basionym: Massangea morreniana Linden Hortus,
Cat. Expos. Brux. 1880.
Dept.: CO, LP.
Elevation: 2200-2300 m.
Type: E. Morren Hortus s.n. CT: LG. Soutb America.
Voucher: S. Beck 22273 (LPB, SEL).
Guzmania remediosensis E. Gross, Trop. SUbtrop.
Pflanzenwelt 95: 20. 1997.
Dept.: LP.
Elevation: Unknown.
Type: M. Miyagawa s.n. HT: HElD. Bolivia.
Voucher: Only known from type collection.
Endemic.
Guzmania retusa L.B. Sm., Fieldiana, Bot. 28(1):
143. 1951.
Dept.: CO, LP.
Elevation: 1300-1700 m.
Type: J. Steyermark 60677. HT: E Venezuela.
Voucher: M. Kessler et al. 7401 (LPB, SEL).
Guzmania roezlii (E. Morren) Mez, in C. DC.,
Monogr. phan. 9: 948. 1896. Basionym:
Schlumbergeria roezli E. Morren, Belgique
Hort. 28: 311. 1878.
Dept.: BE, CO, LP.
Elevation: 550-1450 m.
Type: Roezl in Morren Hortus S.n. CT: LG. Peru.
Voucher: T. Kromer 185 (LPB, SEL).
Guzmania sphaeroidea (Andre) Andre ex Mez, in
C. DC., Monogr. phan. 9: 942. 1896. Basionym: Caraguata sphaeroidea Andre, Enum.
Bromel. 6. 13 Dec. 1888; Rev. Hort. 60: 566.
16 Dec. 1888.
Dept.: CO, LP.
Elevation: 1150-1500 m.
Type: Andre 2685. HT: K. Colombia.
Voucher: M. Kessler et at. 11282 (LPB, SEL).
Guzmania squarrosa (Mez & Sodiro) L.B. Sm. &
Pittendr., J. Wash. Acad. Sci. 43: 403. 1953
[1954]. Basionym: Thecophyllum squarrosum
Mez & Sodiro, Bull. Herb. Boissier, ser. 2, 4:
877. 1904.
Dept.: CO, LP.
Elevation: 1850-2700 m.
Type: Sodiro 47. HT: B. Ecuador.
Voucher: S. Beck 21288 (LPB, SEL).
Guzmania tarapotina Ule, Verh. Bot. Vereins Prov.
Brandenburg 48: 147. 1907.
Dept.: LP.
Elevation: 600(-2350) m.
Type: Ule 6683. HT: B. Peru.
Voucher: S. Beck 24037 (LPB, SEL).
MEZOBROMELIA
Mezobromelia capituIigera (Griseb.) J.R. Grant,
Phytologia 74: 428. 1993. Basionym: Tillandsia
capituligera Griseb., Cat. pI. Cub. 254. 1866.
Dept.: CO, LP.
Elevation: 1300-2200 m.
Type: Wright 3275. HT: GOET; IT: GH. Cuba.
Voucher: M. Kessler et at. 7376 (LPB, SEL).
Mezobromelia pleiosticha (Griseb.) Utley & H.
Lutber, Ann. Missouri Bot. Gard. 78: 270.
1991. Basionym: Tillandsia pleiosticha Griseb.,
Nachr. Konigl. Ges. Wiss. Georg-AugustsUniv. [1]: 19. 1864.
Dept.: LP.
Elevation: 1300 m.
Type: A. Fendler 1514. HT: GOET. Venezuela.
Voucher: M. Kessler et al. 9869 (LPB, SEL).
PF;PINIA
Pepinia caricifolia (Mart. ex Schult. f.) G.S. Varad.
& Gilmartin, Syst. Bot. 13: 297. 1988. Basion-
KROMER ET AL: BOLIVIAN BROMELIACEAE
ym: Pitcairnia caricifolia Mart. ex Schult. f., in
Roem. & Schult., Syst. Veg. 7(2): 1242. 1830.
Dept.: PA, Sc.
Elevation: 150-300 m.
Type: C. Martius s.n. HT: M. Brazil.
Voucher: T. Killeen et al. 5448 (SEL).
PITCAIRNIA
211
Pitcairnia lanuginosa Ruiz & Pav., Fl. peruv. 3:
35. 1802; Phytologia 15: 195. 1967.
Synonym(s): Orthophytum lanuginosum (Ruiz &
Pav.) Beer, Fam. Bramel. 70. 1856; Pitcairnia
sessiliflora Rusby, Bull. New York Bot. Gard.
4: 457. 1907.
Dept.: CO, LP, SC.
Elevation: 200-1300 m.
Type: Ruiz & Pavon S.n. HT: MA. Peru.
Voucher:J. Solomon 14173 (LPB, SEL).
Pitcairnia amboroensis Ibisch, Vasquez, Grass &
Kessler, J. Bromeliad Soc. 49: 129. 1999.
Dept.: Sc.
Elevation: 500-600 m.
Type: P. lbisch & c. lbisch 97.0004A. HT: LPB.
Bolivia.
Voucher: M. Kessler et al. 8660 (GOET, LPB, SEL).
Endemic.
Pitcairnia longissimiflora Ibisch, Vasquez, &
Grass, J. Brameliad Soc. 49: 127. 1999.
Dept.: Sc.
Elevation: 1000 m.
Type: P. lbisch & c. lbisch 97.0097. HT: LPB. Bolivia.
Voucher: P. lbisch & c. Ibisch 98.0225 (USZ).
Endemic.
Pitcairnia aurea Rusby ex L.B. Sm., Phytologia
19: 286. 1970.
Dept.: LP.
Elevation: 900 m.
Type: H. Rusby 665. HT: NY. Bolivia.
Voucher: Only known from type collection.
Endemic.
Pitcairnia multiramosa Mez, in C. DC., Monogr.
phan. 9: 419. 1896.
Synonym(s): Hepetis multiramosa (Mez) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: CH.
Elevation: 1900 m.
Type: Weddell 3637. HT: P. Bolivia.
Voucher: M. Cardenas 5732 (US). Cited in FI. Neotrap. Monogr. 14(1): 306. 1974.
Endemic.
Pitcairnia brittoniana Mez, in C. DC., Managr.
phan. 9: 451. 1896.
Synonym(s): Hepetis brittoniana (Mez) Mez in C.
DC., Monogr. phan. 9: 973. 1896.
Dept.: CO(?), LP.
Elevation: 1200-1700 m.
Type: Bang 592. HT: US; IT: NY. Bolivia.
Voucher: J. Solomon 9281 (LPB, SEL).
Pitcairnia cardenasii L.B. Sm., Phytologia 8: 505.
1963.
Dept.: CH, LP, SC.
Elevation: 800-2000 m.
Type: Cardenas 4648. HT: US. Bolivia.
Voucher: Guardia et al. 230 (SEL).
Endemic.
Pitcairnia crassa L.B. Sm., Lilloa 14: 94. 1948.
Dept.: LP, SC.
Elevation: 700-1400 m.
Type: Cardenas 3690. HT: GH. Bolivia.
Voucher: T. Kromer et al. 126 (LPB, SEL).
Endemic.
Pitcairnia divaricata Wittm., Rijks-Herb. 29: 81.
1916.
Dept.: LP, SC, TA?
Elevation: 900-1300(-1900) m.
Type: Herzog 1212. HT: L; IT: B. Bolivia.
Voucher: S. Beck 1765 (LPB).
Endemic.
Pitcairnia inermis (E. Mey.) E. Mey. ex Schult . &
Schult. f., in Roem. & Schult., Syst. veg. 7(2):
1238. 1830. Basionym: Pourretia inermis E.
Mey., in C. Pres!, Re1iq. haenk. 1(2): 123. 1827.
Dept.: Unknown.
Elevation: Unknown.
Type: Haenke S.n. HT: PRo Peru.
Voucher: Pearce 521 (K). Cited for Bolivia in Fl.
Neotrap. Monogr. 14(1): 326. 1974.
Pitcairnia odontopoda Baker, Handb. Bramel. 93.
1889.
Synonym(s): Hepetis odontopoda (Baker) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: LP.
Elevation: 1700-2500 m.
Type: Pearce s.n. HT: K. Bolivia.
Voucher: Eyerdam 25357 (F, UC). Cited in Fl. Neotrap. Monogr. 14(1): 319. 1974.
Pitcairnia paniculata (Ruiz & Pav.) Ruiz & Pav.,
Syst. veg. fl. peruv. chi!. 3: 36. 1802. Basionym: Pourretia paniculata Ruiz & Pav., Syst.
veg. fl. peruv. chi!. 1: 81. 1798.
Synonym(s): Pitcairnia biattenuata Rusby, Bull.
New York Bot. Gard. 4: 457. 1907.
Dept.: CH, CO?, LP, Sc.
Elevation: (500)1750-2000(-2550) m.
Type: Ruiz & Pavon s.n. Herbarium of deposit unknown. Peru.
Voucher: T. Kromer et al. 125 (LPB, SEL).
Pitcairnia platystemon Mez, in C. DC., Monogr.
phan. 9: 421. 1896.
Synonym(s): Hepetis platystemon (Mez) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: Sc.
Elevation: ca. 500 m.
Type: D'Orbigny 937. HT: P. Bolivia.
Voucher: P. Ibisch & Nowicki 990124 (USZ).
Endemic.
Pitcairnia riparia Mez, Repert. Spec. Nov. Regni
Veg. 12: 416. 1913.
Dept.: LP.
Elevation: 1300 m.
Type: Weberbauer 6156. HT: B. Peru.
SELBYANA
212
Voucher: M. Kessler 9806 (GOET, LPB).
Pitcairnia trianae Andre, Enum. Bromel. 4. 13 Dec
1888; Rev. Hort. 60: 564. 16 Dec 1888.
Synonym(s): Hepetis trianae (Andre) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: CO, LP.
Elevation: 1650-2900 m.
Type: Andre 2069. HT: K. Colombia.
Voucher: T. Kromer 134 (LPB, SEL).
PSEUDANANAS
Pseudananas sagenarius (Arruda) Camargo, Revista Agric. (Piracicaba) 14(7, 8): reprint p. 4.
1939. Basionym: Bromelia sagenaria Arruda,
Diss. pI. Brazil 13. 1810.
Dept.: BE, LP, SC.
Elevation: 200-850 m.
Type: Original description.
Voucher: M. Kessler et al. 5386 (LPB, SEL).
PuYA
Puya alata L.B. Sm., Phytologia 7: 422. 1961.
Dept.: PO.
Elevation: 1000 m.
Type: Cardenas 2851. HT: F. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya alba L.B. Sm., Phytologia 5: 49. 1954.
Dept.: TA.
Elevation: 800 m.
Type: Cardenas 4952a. HT: US. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya atra L.B. Sm., Contr. U.S. Natl. Herb. 29:
536. 1954.
Dept.: CO, LP, Sc.
Elevation: 2500-3500 m.
Type: M.B. Foster 2551. HT: US. Bolivia.
Voucher: M. Kessler et al. 6610 (LPB, SEL).
Endemic.
Puya brittoniana Baker, Handb. Bromel. 124.
1889.
Dept.: LP.
Elevation: 3000-3800 m.
Type: H. Rusby 2849. HT: NY. Bolivia.
Voucher: C. Varadarajan 1466 (LPB). Del. G. Varadarajan.
Endemic.
Puya cardenasii L.B. Sm., Lilloa 14: 94. 1948.
Dept.: CO.
Elevation: 3850-4000 m.
Type: Cardenas 4082. HT: US; IT: GH. Bolivia.
Voucher: C. Varadarajan 1442 (LPB). Det. G. Varadarajan.
Endemic.
Puya claudiae Ibisch, Vasquez & Gross, J. Bromeliad Soc. 49: 132. 1999.
Volume 20(2) 1999
Dept.: Sc.
Elevation: 2000 m.
Type: P. lhisch & c. 1hisch 980040. HT: LPB. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya cristata L.B. Sm., Lilloa 14: 95. 1948.
Dept.: CO.
Elevation: 3000 m.
Type: Cardenas 3311. HT: GH. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya ctenorhyncha L.B. Sm., Phytologia 5: 49.
1954.
Dept.: LP.
Elevation: 2500-4050 m.
Type: Cardenas 4867. HT: US. Bolivia.
Voucher: L. Besse et ai. 6]8 (SEL).
Endemic.
Puya dyckioides (Baker) Mez, in C. DC., Monogr.
phan. 9: 486. 1896. Basionym: Pitcairnia dyckioides Baker, Handb. Bromel. 118. 1889.
Dept.: CH, SC, TA.
Elevation: 2250-2400(-3750?) m.
Type: Weddell 3739. HT: P. Bolivia.
Voucher: P. lbisch & c. lbisch 93.1026 (SEL).
"cf." .
Puya entre-riosensis Ibisch & Gross, J. Bromeliad
Soc. 48: 217. 1998.
Dept.: TA.
Elevation: 1800 m.
Type: P. Ibisch & c. Ibisch 93.1282. HT: LPB; IT:
HEID. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya ferruginea (Ruiz & Pav.) L.B. Sm., Phytologia 16: 461. 1968. Basionym: Pitcairniaferruginea Ruiz & Pav., Fl. peruv. 3: 36. 1802.
Synonym(s): Pitcairnia consimilis Baker, J. Bot. 19:
226, 266. 1881.
Dept.: CO, LP, Sc.
Elevation: 1150-3800 m.
Type: Ruiz & Pavon s.n. HT: BM. Peru.
Voucher: J. Solomon 17336 (LPB, SEL).
Puya fiebrigii Mez, Repert. Spec. Nov. Regni Veg.
3: 9. 1906.
Dept.: TA?
Elevation: Unknown.
Type: Fiebrig 3210. LT: B. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya fosteriana L.B. Sm., J. Wash. Acad. Sci. 40:
216. 1950.
Dept.: LP.
Elevation: 3900-4400 m.
Type: M.B. Foster 2573. HT: US. Bolivia.
Voucher: J. Solomon 18299 (MO, SEL).
Endemic.
Puya glabrescens L.B. Sm., Contr. U.S. Natl. Herb.
29: 537. 1954.
Dept.: CO, PO.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Elevation: 2550-3700 m.
Type: Cardenas 4384. HT: US. Bolivia.
Voucher: L. Besse et al. 502 (SEL).
Endemic.
Puya giareosa L.B. Sm., Lilloa 14: 96. 1948.
Dept.: LP.
Elevation: 3600 m.
Type: Cardenas 3583. HT: GH. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya herzogii Wittm., Rijks-Herb. 29: 86. 1916.
Dept.: CO, LP.
Elevation: 3300-3950 m.
Type: Herzog 2269. HT: L; IT: B. Bolivia.
Voucher: M. Kessler et al. 6581 (SEL).
Endemic.
Puya hofstenii Mez, Repert. Spec. Nov. Regni Veg.
3: 8. 1906.
Synonym(s): Puya hauthalii Mez, Repert. Spec.
Nov. Regni Veg. 16: 65. 1919.
Dept.: CH, PO.
Elevation: 3000-3500 m.
Type: Hofsten 1710. HT: B. Argentina.
Voucher: Hromadnik 19017a (WU).
Puya hromadnikii Rauh. Trop. Subtrop. Pflanzenwelt 41: 5. 1983.
Dept.: TA.
Elevation: 2550-2700 m.
Type: Hromadnik 5128. HT: HElD. Bolivia.
Voucher: W. Till 60a (WU).
Endemic.
Puya humilis Mez, in C. DC., Monogr. phan. 9:
498. 1896.
Synonym(s): Puya werdermannii Harms, Notizbl.
Bot. Gart. Berlin-Dahlem 10: 793. 1929.
Dept.: CH, CO, PO.
Elevation: (2100-)2800-4200 m.
Type: Kuntze s.n. HT: NY. Bolivia.
Voucher: C. Varadarajan et al. 1451 (LPB, SEL).
Det. G. Varadarajan.
Endemic.
Puya kuntzeana Mez. in C. DC., Monogr. phan. 9:
490. 1896.
Dept.: CO.
Elevation: 1300 m.
Type: Kuntze s.n. HT: NY. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya lasiopoda L.B. Sm., Proc. Amer. Acad. Arts
70: 153. 1935.
Dept.: BE, LP?
Elevation: 500(- J 850?) m.
Type: H. Rusby 2232. HT: NY. Bolivia.
Voucher: J. Solomon 10862 (MO). "cf.".
Puya laxa L.B. Sm., Phytologia 6: 195. 1958.
Dept.: Sc.
Elevation: 1500-1900 m.
Type: Cardenas 5092. HT: US. Bolivia.
Voucher: S. Beck 7100 (LPB). Det. G. Varadarajan.
Endemic.
Puya leptostachya L.B. Sm., Lilloa 14: 97. 1948.
213
Dept.: CO.
Elevation: 3200-3400 m.
Type: Cardenas 3581. HT: GH. Bolivia.
Voucher: C. Varadarajan et al. 1445 (LPB). Det.
G. Varadarajan.
Endemic.
Puya meziana Wittm., Rijks-Herb. 29: 85. 1916.
Dept.: CO(?), LP.
Elevation: (2200?-)3000-3600 m.
Type: Herzog 2487. HT: L. Bolivia.
Voucher: J. Solomon 13058 (LPB, SEL).
Endemic.
Puya micrantha Mez, Repert. Spec. Nov. Regni
Veg. 3: 8. 1906.
Dept.: TAIArgentina border.
Elevation: 1700 m.
Type: Fiebrig 2244. HT: B; IT: BM, GH. Argentina.
Voucher: No known collections from Bolivia.
Puya minima L.B. Sm., Phytologia 7: 423. 1961.
Synonym(s): Puya pusilla H. Luther, Bromelia 4(4):
30 Nov. 1997 [1998].
Dept.: TA.
Elevation: 2500-2650 m.
Type: West 8345. HT: Uc. Bolivia.
Voucher: S. Beck & M. Libermann 14310 (LPB,
SEL).
Endemic.
Puya mirabilis (Mez) L.B. Sm., Phytologia 16:
461. 1968. Basionym: Pitcairnia mirabilis
Mez, Repert. Spec. Nov. Regni Veg. 3: 6. 1906.
Dept.: CO, TA.
Elevation: 1400-2600 m.
Type: Fiebrig 2320. HT: B. Bolivia.
Voucher: P. lbisch & Rojas 415 (LPB). Det. G. Varadarajan.
Puya mollis Baker ex Mez, in C. DC., Monogr.
phan. 9: 488. 1896.
Dept.: LP.
Elevation: 2400-3300 m.
Type: Bang 1629. LT: K; ILT: BM, GH, US. Bolivia.
Voucher: (Syntype) C. Mandan 1190 (BR, GH,
MANCH, P, PL, STU, WU, Z). Cited in FI.
Neotrop. Monogr. 14(1): 123. 1974.
Endemic.
Puya nana Wittm., Rijks-Herb. 29: 85. 1916.
Synonym(s): Pitcairnia nana (Wittm.) L.B. Sm.,
Phytologia 7: 5. 1959; 10: 30. 1964.
Dept.: Sc.
Elevation: (1350-)1700-2100 m.
Type: Herzog 1856a. HT: L. Bolivia.
Voucher: M. Kessler et al. 6356 (LPB, SEL).
Endemic.
Puya olivacea Wittm., Rijks-Herb. 29: 83. 1916;
emend. L.B. Sm., Contr. U.S. Nat. Herb. 29:
538. 1954.
Synonym(s): Pitcairnia olivacea (Wittm.) Mez in
Engler, Pfianzenr. IV, 32: 247. 1935.
Dept.: Sc.
Elevation: 950-2000 m.
Type: Herzog 1216. HT: L. Bolivia.
Voucher: J. Solomon & M. Nee 17936 (MO, SEL).
Endemic.
214
SELBYANA
Puya paupera Mez, Repert. Spec. Nov. Regni Veg.
3: 14. 1906.
Dept.: TA.
Elevation: 2600 m.
Type: Fiebrig 2005. HT: B. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya pearcei (Baker) Mez, in C. DC., Monogr.
phan. 9: 480. 1896. Basionym: Pitcairnia pearcei Baker, Handb. Bromel. 120. 1889.
Dept.: CO, LP, SC?
Elevation: 750-2800 m.
Type: Pearce s.n. HT: K. Bolivia.
Voucher: A. Gentry et al. 44254 (LPB, SEL).
Endemic.
Puya pendulifiora L.B. Sm., Contr. Gray Herb. 98:
12. 1932.
Dept.: LP.
Elevation: 1400 m.
Type: Buchtien 7182. HT: US. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya potosina L.B. Sm., Phytologia 7: 423. 1961.
Dept.: CR, PO.
Elevation: (2050-)3400 m.
Type: Cardenas 3738. HT: US. Bolivia.
Voucher: M. Kessler et al. 4804 (LPB).
Endemic.
Puya prosanae Ibisch & E. Gross, 1. Bromeliad
Soc. 43: 212. 1993.
Dept.: CO.
Elevation: 3500-3800 m.
Type: P. Ibisch & c. Ibisch 625A. HT: HElD; IT:
LPB. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya raimondii Harms, Notizbl. Bot. Gart. BerlinDahlem 10: 213. 1928.
Dept.: CO, LP.
Elevation: (2400-)3000-4200 m.
Type: Weberbauer 2955. HT: B. Peru.
Voucher: G. & D. Schmitt (LPB, SEL).
Puya reducta L.B. Sm., Phytoiogia 10: 472. 1964.
Dept.: LP.
Elevation: 2600 m.
Type: Cardenas 5687. HT: US. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya riparia L.B. Sm., Phytologia 7: 424. 1961.
Dept.: LP.
Elevation: 1350(-3950) m.
Type: M.B. Foster 2578. HT: US. Bolivia.
Voucher: S. Beck 11277 (LPB). Det. A. Gilmartin.
Endemic.
Puya rusbyi (Baker) Mez, in C. DC., Monogr. phan.
9: 482. 1896. Basionym: Pitcaimia rusbyi Baker, Handb. Bromel. 122. 1889.
Dept.: LP.
Elevation: 3000-3600 m.
Type: H. Rusby 2846. HT: NY. Bolivia.
Voucher: West 6402 (UC). Cited in Fl. Neotrop.
Monogr. 14(1): 89. 1974.
Volume 20(2) 1999
Endemic.
Puya sanctae-crucis (Baker) L.B. Sm., Proc. Amer.
Acad. Arts 70: 154. 1935. Basionym: Pitcaimia
sanctae-crucis Baker, Handb. Bromel. 120.
1889.
Synonym(s): Pitcaimia robusta Rusby, Bull. New
York Bot. Gard. 6: 488. 1910.
Dept.: LP, SC.
Elevation: 1300-2450 m.
Type: Castelnau s.n. HT: P. Bolivia.
Voucher: L. Besse et al. 636 (SEL).
Endemic.
Puya secunda L.B. Sm., Phytologia 7: 424. 1961.
Dept.: CO.
Elevation: 1800-3000 m.
Type: Cardenas 4381. HT: US. Bolivia.
Voucher: P. lbisch 14074 (LPB).
Endemic.
Puya serranoensis Raub, Trop. SUbtrop. Pflanzenwelt 41: 14. 1983.
Dept.: SC.
Elevation: 1700-2600 m.
Type: Hromadnik 5230. HT: HElD. Bolivia.
Voucher: l. Vargas & Fuentes 3046 (SEL).
Endemic.
Puya solomonii Varad., J. Bromeliad Soc. 39: 12l.
1989.
Dept.: LP.
Elevation: 3050 m.
Type: G. Varadarajan, U. Varadarajan & J. Solomon 1471. HT: GH; IT: LPB, MO, US, WS.
Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya spathacea (Griseb.) Mez, in C. DC., Monogr.
phan. 9: 481. 1896. Basionym: Pitcaimia spathacea Griseb., Abh. KOnig!. Ges. Wiss. GOttingen 24: 329. 1879.
Dept.: TA.
Elevation: 2050 m.
Type: Hieronymus 175. HT: GOET; IT: CORD. Argentina.
Voucher: E. Bastian 414 (LPB).
Puya stenothyrsa (Baker) Mez. in C. DC., Monogr.
phan. 9: 481. 1896. Basionym: Pitcaimia stenothyrsa Baker, Handb. Bromel. 122. 1889.
Synonym(s): Pitcaimia bangii Baker, Mem. Torrey
Bot. Club 6: 124. 1896.
Dept.: CO, LP, SC, TA.
Elevation: (1350-)2700-3800 m.
Type: Mandon 1189. HT: K; IT: GH, PI, US. Bolivia.
Voucher: L. Hromadnik 19026 (WU).
Endemic.
Puya tristis L.B. Sm., Phytologia 10: 475. 1964.
Dept.: CO.
Elevation: 3900 m.
Type: Cardenas 6067. HT: US. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya trollii L.B. Sm., Phytologia 16: 459. 1968.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Dept.: PO.
Elevation: 3350 m.
Type: Troll 661. HT: M. Bolivia.
Voucher: S. Beck et al. 26787 (LPB).
Endemic.
Puya tuberosa Mez, in C. DC., Monogr. phan. 9:
483. 1896; emend. L.B. Sm., Contr. U.S. Natl.
Herb. 29: 539. 1954.
Dept.: SC, TA.
Elevation: 1000-2750 m.
Type: Haenke S.n. HT: M. Bolivia.
Voucher: Cardenas 4647 (US). Cited in Fl. Neotrop.
Monogr. 14(1): 120. 1974.
Endemic.
Puya tunariensis Mez, in C. DC., Monogr. phan.
9: 498. 1896.
Dept.: CO.
Elevation: (2000-)3150-3800 m.
Type: Kuntze s.n. HT: NY. Bolivia.
Voucher: M. Kessler et at. 6830 (LPB, SEL).
Endemic.
Puya ugentiana L.B. Sm., Phytologia 13: 147.
1966.
Dept.: CH.
Elevation: 1900-1950 m.
Type: Ugent 4998. HT: US; IT: WIS. Bolivia.
Voucher: L. Hromadnik 19066 (WU).
Endemic.
Puya ultima L.B. Sm., Contr. U.S. Natl. Herb. 29:
540. 1954.
Dept.: LP.
Elevation: 1500-2200 m.
Type: M.B. Foster 2575. HT: US. Bolivia.
Voucher: M. Kessler et al. 10574 (SEL).
Endemic.
Puya valida L.B. Sm., Phytologia 10: 474. 1964.
Dept.: CH.
Elevation: 2500 m.
Type: Cardenas 6012. HT: US. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya vallo-grandensis Rauh, Trop. Subtrop. Pflanzenwelt 41: 9. 1983.
Dept.: SC.
Elevation: 2000-2250 m.
Type: Hromadnik 5279. HT: HElD. Bolivia.
Voucher: P. 1bisch & c. 1bisch 93.0861 (SEL).
Endemic.
Puya vasquezii P. Ibisch & E. Gross, Rev. Soc. Boliviana Bot. 2(1): 26. 1998.
Dept.: SC.
Elevation: 600 m.
Type: R. Vasquez et al. 2025. HT: LPB; IT: NY,
USZ. Bolivia.
Voucher: Only known from type collection.
Endemic.
Puya weberbaueri Mez, Bull. Herb. Boissier, ser.
2,4: 633. 1904.
Dept.: LP.
Elevation: Unknown.
Type: Weberbauer 847. HT: B. Peru.
215
Voucher: Cardenas 5534 (US). Cited for Bolivia in
Fl. Neotrop. Monogr. 14(1): 148. 1974.
Puya weddelliana (Baker) Mez, in C. DC., Monogr.
phan. 9: 475. 1896. Basionym: Pitcairnia weddelliana Baker, Handb. Bromel. 12. 1889.
Dept.: CH, TA.
Elevation: 1800-3000 m.
Type: Weddell 4001. HT: P. Bolivia.
Voucher: L. Hromadnik 19046a (WU).
Endemic."
Puya sp. 1.
Dept.: LP.
Elevation: 1450 m.
Voucher: M. Kessler et al. 10386 (LPB, SEL).
RACINAEA
Racinaea cr. riocreuxii (Andre) M.A. Spencer &
L.B. Sm., Phytologia'74: 156. 1993. Basionym:
Tillandsia riocreuxii Andre, Enum. Bromel. 7.
13 Dec. 1888; Rev. Hort. 60: 567. 16 Dec.
1888.
Dept.: LP.
Elevation: 3250 m.
Type: Andre 4408. HT: K. Ecuador.
Voucher: B. Herzog 368 (LPB).
Racinaea flexuosa (Baker) M.A. Spencer & L.B.
Sm., Phytologia 74: 154. 1993. Basionym: Catopsisjlexuosa Baker, J. Bot. 25: 175. 1887.
Synonym(s): Tillandsia bakeri L.B. Sm., Contr.
Gray Herb. 95: 45. 1931.
Dept.: CO, LP.
Elevation: 2100-2700 m.
Type: Mandon 1172. HT: K. Bolivia.
Voucher: M. Kessler et at. 12495 (LPB, SEL).
Racinaea kessleri H. Luther, SeJbyana 19: 86.
1998.
Dept.: CO.
Elevation: 2200-3100 m.
Type: M. Kessler et at. 8380. HT: SEL; IT: LPB.
Bolivia.
Voucher: M. Kessler et al. 9541 (LPB, SEL).
Endemic.
Racinaea monticola (Mez & Sodiro) M.A. Spencer
& L.B. Sm., Phyto10gia 74: 155. 1993. Basionym: Tillandsia monticola Mez & Sodiro, Bull.
Herb. Boissier, ser. 2, 4: 1135. 1904.
Dept.: LP.
Elevation: 750-850 m.
Type: Sodiro 171/33. HT: B; IT: F, GH. Ecuador.
Voucher: O. Buchtien 1266 (US). Cited for Bolivia
in FI. Neotrop. Monogr. 14(2): 1038. 1977.
Racinaea nervibractea (Gilmartin & H. Luther)
J.R. Grant, Phyto10gia 74: 429. 1993. Basionym: Tillandsia nervibractea Gilmartin & H.
Luther, J. Bromeliad Soc. 40(6): 250. 1990.
Dept.: CO, SC.
Elevation: 2100-2150 m.
Type: Kalmbacher S.n. HT: US. Ecuador.
Voucher: P. 1bisch 93.1537 (LPB, SEL).
Racinaea pardina (L.B. Sm.) M.A. Spencer & L.B.
216
SELBYANA
Sm., Phytologia 74: 155. 1993. Basionym: TUlandsia pardina L.B. Sm., Lilloa 14: 98. 1948.
Dept.: LP, Sc.
Elevation: 1300-1700 m.
Type: Buchtien 7183. HT: US. Bolivia.
Voucher: M. Kessler et al. 9871 (LPB, SEL).
Endemic.
Racinaea parvifiora (Ruiz & Pav.) M.A. Spencer
& L.B. Sm., Phytologia 74: 156. 1993. Basionym: Tillandsia parvijlora Ruiz & Pav., FL peruv. 3: 41. 1802.
Dept.: CO, LP, Sc.
Elevation: (500-)950-2500 m.
Type: Ruiz & Pavon s.n. HT: BM. Peru.
Voucher: M. Kessler 11655 (LPB, SEL).
Racinaea pendulispica (Mez) M.A. Spencer &
L.B. Sm., Phytologia 74: 156. 1993. Basionym:
Tillandsia pendulispica Mez, in C. DC., Monogr. phan. 9: 745. 1896.
Synonym(s): Tillandsia triangularis Rusby, Descr.
S. Amer. pI. 3. 1920.
Dept.: CO.
Elevation: Unknown (> 2500 m?).
Type: Poeppig 1348. HT: G; IT: W (lost). Peru.
Voucher: M. Bang s.n. (NY). Cited for Bolivia in
FL Neotop. Monogr., 14(2): 1040. 1977.
Racinaea schumanniana (Wittm.) J.R. Grant, Novon 4: 363. 1994. Basionym: Catopsis schumanniana Wittm., Bot. lahrb. Syst. 11: 70.
1890.
Dept.: CO, LP.
Elevation: 1050-2700 m.
Type: Lehmann 22. HT: G. Colombia.
Voucher: 1. Solomon 9189 (SEL).
Racinaea seemannii (Baker) M.A. Spencer & L.B.
Sm., Phytologia 74: 157. 1993. Basionym: Guzmania seemannii Baker, Handb. Bramel. 153.
1889.
Dept.: CO, LP.
Elevation: 2100-3500 m.
Type: Seemann 898. HT: K. Ecuador.
Voucher: M. Kessler et al. 6648 (LPB, SEL).
Racinaea spiculosa (Griseb.) M.A. Spencer & L.B.
Sm., Phytologia 74: 157. 1993. Basionym: Tillandsia spiculosa Griseb., Nachr. Konig!. Ges.
Wiss. Georg-Augusts-Univ. [1]: 17. 1864.
Synonym(s): Tillandsia brittoniana Baker, Handb.
Bramel. 195. 1889; Tillandsia micrantha Baker, Bull. Torrey Bot. Club 29: 698. 1902.
Dept.: BE, CO, SC, LP.
Elevation: 300-2500 m.
Type: A. Fendler 1518. HT: GOET. Venezuela.
Voucher: M. Kessler et al. 7741 (LPB, SEL).
Racinaea tetrantha (Ruiz & Pav.) M.A. Spencer &
L.B. Sm., Phytologia 74: 158. 1993. Basionym:
Tillandsia tetrantha Ruiz & Pav., F!. peruv. 3:
39, pi 265. 1802.
Dept.: CO, LP.
Elevation: 2400-2750 m.
Type: Ruiz & Pavon s.n. HT: MA. Peru.
Voucher: M. Kessler et al.10421 (SEL).
Volume 20(2) 1999
TILLANDSIA
Tillandsia acuminata L.B. Sm., Contr. U.S. Nat!.
Herb. 29: 434. 1951.
Dept.: CO, LP.
Elevation: 2000-2750 m.
Type: M.E. Foster & L.E. Smith 1402. HT: GH. Colombia.
Voucher: M. Kessler et al. 5565 (LPB, SEL).
Tillandsia adpressifiora Mez, in C. DC., Monogr.
phan. 9: 661.1896.
Dept.: Sc.
Elevation: 200-250 m.
Type: Wullschlaegel 1335. HT: BM. Suriname.
Voucher: S. Halloy et al. 4406 (SEL).
Tillandsia aizoides Mez, in C. DC,. Monogr. phan.
9: 866.1896.
Dept.: PO, TA.
Elevation: 2100-2800 m.
Type: Hieronymus & Niederlein 850. HT: B; IT:
CORD. Argentina.
Voucher: H. Hromadnik & L. Hromadnik 5086
(WU).
TiUandsia argentina C.H. Wright, Bull. Misc. Inform. Kew 60. 1907.
Dept.: CH, TA.
Elevation: 800-1950 m.
Type: Stuckert s.n. HT: K. Argentina.
Voucher: S. Beck & M. Libermann 14275 (LPB,
SEL).
Tillandsia asplundii L.B. Sm., Phytologia 6: 441.
1959.
Dept.: CO, LP.
Elevation: 1350-2500 m.
Type: Asplund 19279. HT: S; IT: US. Ecuador.
Voucher: M. Kessler et al. 6975 (LPB, SEL).
Tillandsia australis Mez, Repert. Spec. Nov. Regni
Veg. 16: 75. 1919.
Synonym(s): Tillandsia maxima Lillo & Hauman,
Anales Mus. Nac. Hist. Nat. Buenos Aires 29:
245,416. 1917, non Strangeways, 1882.
Dept.: CH, LP, SC, TA.
Elevation: 800-2100(-3100) m.
Type: Lorentz & Hieronymus 287. HT: LIL; IT: B,
CORD, US. Argentina.
Voucher: M. Kessler et al. 4946 (LPB, SEL).
Tillandsia bandensis Baker, J. Bot. 25: 234. 1887.
Synonym(s): Tillandsia pilosa L.B. Sm., Phytologia
7: 175. 1960.
Dept.: CH, CO, SC, TA.
Elevation: (600-)1200-2200 m.
Type: Tweedie S.n. HT: K. Uruguay.
Voucher: M. Kessler et al. 5914 (LPB, SEL).
Tillandsia bermejoensis H. Hromadnik, Trap. Subtrap. Pflanzenwelt 31: 9. 1979.
Dept.: Sc.
Elevation: 850-1450 m.
Type: Hromadnik 3054. HT: HElD. Bolivia.
Voucher: D. Cathcart 16 (SEL).
Endemic.
Tillandsia biflora Ruiz & Pav., FL peruv. 3: 41.
1802.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Dept.: CO, LP.
Elevation: 2100-2800 m.
Type: Ruiz & Pavon s.n. HT: MA. Peru.
Voucher: M. Kessler et at. 9556 (LPB, SEL).
Tillandsia boliviana Mez, Bull. Herb. Boissier, ser.
2, 4: 1l30. 1904.
Dept.: CH7, LP7.
Elevation: 15007-2730? m.
Type: Bang 2202. HT: B; IT: BM, F, G, GH, K, LE,
M, MICH, MO, 0, US, WU. Bolivia.
Voucher: M. Kessler et al. 4535 (LPB, SEL). "cf.".
Endemic.
Tillandsia boliviensis Baker, Mem. Torrey Bot.
Club 4(3): 267. 1895; emend. Contr. Gray Herb.
98: 15. 1932.
Dept.: LP.
Elevation: 2800-3600 m.
Type: Bang 159 A. HT: NY. Bolivia.
Voucher: Buchtien 808 (US). Cited in Fl. Neotrop.
Monogr. 14(2): 769. 1977.
Endemic.
Tillandsia cr. buseri Mez, Bull. Herb. Boissier, ser.
2, 3: 145. 1903.
Dept.: LP.
Elevation: 2750 m.
Type: Langlasse 102-A. HT: G; IT: K. Colombia.
Voucher: M. Kessler et al. 12044 (LPB, SEL).
217
Type: Cardenas 491. HT: GH. Bolivia.
Voucher: M. Kessler et al. 4832 (LPB, SEL).
Endemic.
Tillandsia carnosa L.B. Sm., Phytologia 9: 251.
1963.
Dept.: CO, LP.
Elevation: 900-3000 m.
Type: J. Wurdack 603. HT: US; IT: USM. Peru.
Voucher: M. Kessler et al. 5752 (LPB, SEL).
Tillandsia caulescens Brongn. ex Baker, Handb.
Bromel. 168. 1889.
Dept.: LP.
Elevation: 850 - J 550 m.
Type: Gay 1186. HT: P. Peru.
Voucher: M. Kessler et al. 5808 (LPB, SEL).
Tillandsia cochabambae E. Gross & Rauh, Die
Bromelie 2/86: 14. 1986.
Dept.: CO.
Elevation: 2400-2700 m.
Type: W. Rauh 40609. HT: HElD. Bolivia.
Voucher: M. Kessler et al. (LPB, SEL).
Endemic.
Tillandsia colganii Ehlers, J. Bromeliad Soc. 47:
103. 1997.
Dept.: SC.
Elevation: 2700-3200 ill.
Type: R. Ehlers EB 951002. HT: WU. Bolivia.
Voucher: R. Ehlers 951101 (WU). Det. R. Ehlers.
Endemic.
Tillandsia caliginosa W. Till, PI. Syst. Evol. 147:
282. 1984. Basionym: Tillandsia crocata (E.
Morren) Baker var. tristis Rauh, Trop. Subtrop.
Pflanzenwelt 43: 17. 1983, non T. tristis Mez
1896.
Dept.: CH, CO. PO, TA.
Elevation: 1900-3000 m.
Type: D. Muhr s.n. HT: HEID. Argentina.
Voucher: P. 1bisch & c. 1bisch 93.0329 (LPB, SEL).
Tillandsia comarapaensis H. Luther, J. Bromeliad
Soc. 34: 54. 1984.
Dept.: CH, CO, SC.
Elevation: (400-)1550-2600 m.
Type: L. Besse et al. 1713. HT: SEL. Bolivia.
Voucher: M. Kessler et al. 4605 (LPB, SEL).
Endemic.
Tillandsia calochlamys Ehlers & L. Hrom., Die
Bromelie 2/1998: 38. 1998.
Dept.: CO.
Elevation: 2800-3550 m.
Type: R. Ehlers EB952403. HT: LPB. Bolivia.
Voucher: W. Till 158 (WU).
Endemic.
Tillandsia compacta Griseb., Nachr. Konigl. Ges.
Wiss. Georg-Augusts-Univ. [1]: 18. 1864.
Dept.: CO.
Elevation: 2000 m.
Type: A. Fendler 1508. HT: GOET; IT: K. Venezuela.
Voucher: L. Dorr 7052 (LPB, SEL).
Tillandsia camargoensis L. Hrom., Die Bromelie
111991: 9. 1991.
Dept.: CH, PO.
Elevation: 2300-3000 m.
Type: Hromadnik 5073 A. HT: WU. Bolivia.
Voucher: E. Haugg 7023 (WU). in part.
Endemic.
Tillandsia compianata Benth., Bot. voy. Sulphur
173. 1846.
Dept.: CO, LP, Sc.
Elevation: (300-)1050-2900 m.
Type: Sinclair s.n. HT: K. Ecuador.
Voucher: M. Kessler et al. 7010 (LPB, SEL).
Tillandsia capillaris Ruiz & Pav., FI. peruv. 3: 42.
1802.
Dept.: CH, CO, LP, OR, PO, SC, TA.
Elevation: 900-4000 m.
Type: Ruiz & Pavon S.n. LT: BM; ILT: BM, MA.
Peru.
Voucher: M. Kessler et al. 6502 (LPB, SEL).
Tillandsia confinis L.B. Sm., Phytologia 4: 218.
1953.
Synonym(s): Tillandsia subtropicalis L.B. Sm.,
Phytologia 8: 505. 1963.
Dept.: CO, LP, Sc.
Elevation: 1300-1600 m.
Type: Ferreyra 1686. HT: USM. Peru.
Voucher: J. Solomon 9253 (LPB, MO, SEL).
Tillandsia cardenasii L.B. Sm., Proc. Amer. Acad.
Arts 70: 154. 1935.
Dept.: CH, CO, PO, TA.
Elevation: 1700-3400 m.
Tiliandsia cotagaitensis L. Hrom., PI. Syst. Evo!.
147: 284, 1984.
Dept.: PO.
Elevation: 3000 m.
218
SELBYANA
Type: Hromadnik 5093. HT: WU; IT: LI, NY, WU.
Bolivia.
Voucher: W. Till 42 (WU).
Endemic.
Tillandsia denudata Andre, Enum. Bromel. 8. 13
Dec. 1888; Rev. Hort. 60: 568. 16 Dec. 1888.
Dept.: LP.
Elevation: 1250 m.
Type: Andre 1358. HT: HElD. Colombia.
Voucher: M. Kessler et al. 5587 (LPB, SEL).
Tillandsia didisticha (E. Morren) Baker, J. Bot. 26:
16. 1888. Basionym: Anoplophytum didistichum E. Morren, Belgique Hort. 31: 164. 1881.
Synonym(s): Guzmania complanata Wittm., Meded.
Rijks-Herb. 29: 92. 1916.
Dept.: BE, CH, CO, LP, SC, TA.
Elevation: 200-2450 m.
Type: Jacob Matoy Hortus s.n. HT: LG. Brazil.
Voucher: M. Kessler et al. 5584 (LPB, SEL).
Tillandsia duratii Visiani, Nuovi. Saggi r. Accad.
Sci. Padova 5: 271. 1840.
Synonym(s): Tillandsia weddellii Baker, Handb.
Bromel. 181. 1889.
Dept.: CH, CO, LP, PO, SC, TA.
Elevation: 400-2250 m.
Type: Padua Hortus s.n. HT: PAD.
Voucher: M. Kessler et al. 5912 (LPB, SEL).
Tillandsia edithae Rauh, Trop. Subtrop. Pflanzenwelt 8: 19. 1974.
Dept.: CH, LP, SC.
Elevation: 750-2700 m.
Type: Krahn K 295. HT: HElD; IT: US, HAL. Bolivia.
Voucher: M. Kessler et al. 12314 (LPB, SEL).
Endemic.
Tillandsia engleriana Wittm., Bot. Jahrb. Syst. 11:
63. 1890.
Dept.: CO, LP, SC.
Elevation: 1700-2500 m.
Type: Lehmann 520. HT: G-B. Colombia.
Voucher: D. Smith 13442 (LPB, MO).
Tillandsia erici Ehlers, Die Bromelie 111998: 20.
1998.
Dept.: TA.
Elevation: 1800-2200 m.
Type: E. Haugg & E. Heger 11247. HT: WU; IT:
LPB. Bolivia.
Voucher: Only known from type collection.
Endemic.
Tillandsia fendleri Griseb., Nachr. Konigi. Ges.
Wiss. Georg-Augusts-Univ. [l): 17. 1864.
Dept.: BE, CO, LP, Sc.
Elevation: 700-2200(-3000) m.
Type: A. Fendler 1515. HT: GOET. Venezuela.
Voucher: M. Kessler et al. 9868 (LPB, SEL).
Tillandsia funebris A. Cast., Anales Mus. Nac.
Hist. Nat. Buenos Aires 37: 502. 1933.
Dept.: CH, CO, PO, SC, TA.
Elevation: 400-2100 m.
Type: Schreiter 1689. HT: BA; IT: B, GH, LIL. Argentina.
Volume 20(2) 1999
Voucher: M. Kessler et al. 4710 (LPB, SEL, WU).
Tillandsia gerdae Ehlers, Die Bromelie 2/87: 32.
1987.
Dept.: CH, CO.
Elevation: (1700-)2000-2600 m.
Type: E. Haugg 7163. HT: WU; IT: WU. Bolivia.
Voucher: E. Haugg 8283 (WU).
Endemic.
Tillandsia gilliesii Baker, J. Bot. 16: 240. 1878.
Dept.: CH, CO, PO, TA.
Elevation: 1800-3200 m.
Type: Gillies s.n. HT: K; IT: CGE, GH. Argentina.
Voucher: M. Cardenas 2418 (GH). Cited for Bolivia
in FI. Neotop. Monogr. 14(2): 885. 1977.
Tillandsia basei Ehlers & L. Hrom., Die Bromelie
311996: 176. July 1997.
Dept.: CH.
Elevation: 3100 m.
Type: F. Hase et al. EB 951501. HT: WU. Bolivia.
Voucher: L Hromadnik 19046 (WU).
Endemic.
Tillandsia hegeri Ehlers, Die Bromelie 2/1994: 33.
1994.
Dept.: CH, TA.
Elevation: 270-2750 m.
Type: E. Heger s.n. HT: WU. Bolivia.
Voucher: L Hromadnik 19028 (WU).
Endemic.
Tillandsia helmutii L. Hrom., Die Bromelie 3/
1990: 63. Jan. 1990 [1991).
Dept.: CH.
Elevation: 1700-1800 m.
Type: L. & H. Hromadnik 13009. HT: WU; IT: WU.
Bolivia.
Voucher: L & H. Hromadnik 5028 (WU).
Endemic.
Tillandsia hirta w. Till & L. Hrom., PI. Syst. Evoi.
147: 285. 1984.
Dept.: CH, CO, PO, TA.
Elevation: 2500-3700 m.
Type: L & R. Hromadnik HR 5001. HT: WU; IT:
WU. Bolivia.
Voucher: L Hromadnik 19092 (WU).
Tillandsia imporaensis R. Ehlers, Die Bromelie 3/
1998: 76. 1998.
Dept.: CH, CO.
Elevation: 3050 m.
Type: E. Haugg 7078. HT: WU. Bolivia.
Voucher: G. Kohres & L. KiJhres s.n. (WU).
Endemic.
Tillandsia ionochroma Andre ex Mez, in C. DC.,
Monogr. phan. 9: 801. 1896.
Dept.: CO, LP, SC.
Elevation: 2000-3600 m.
Type: Andre 2746. HT: K; IT: NY. Ecuador.
Voucher: S. Beck 14469 (LPB, SEL).
Tillandsia ixioides Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 24: 333. 1879.
Dept.: CH, TA.
Elevation: 1700-2200 m.
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Type: Lorentz 1196. HT: B; IT: BM, G-B, GOET,
K, M, US. Argentina.
Voucher: M. Cardenas 4093 (GH, US). Cited for
Bolivia in Fl. Neotrop. Monogr. 14(2): 821.
1977.
Tillandsia jucunda A. Cast., Anales Mus. Nac.
Hist. Nat. Buenos Aires 36: 56. 1929.
Dept.: CH.
Elevation: 2300-2700 m.
Type: Schreiter 26/2367. HT: BA. Argentina.
Voucher: W. Till 20 (WU).
Tillandsia juncea (Ruiz & Pav.) Poir., Encycl.
suppl. 5: 309. 1817. Basionym: Bonapartea
juncea Ruiz & Pav., FI. peruv. 3: 38. 1802.
Dept.: BE, CO, LP, PA, SC.
Elevation: 800-1400(-2000) m.
Type: Ruiz & Pavon s.n. HT: MA. Peru.
Voucher: M. Kessler et al. 5595 (LPB, SEL).
Tillandsia kessleri H. Luther, Selbyana 20: 10.
1999.
Dept.: CO.
Elevation: 2600-2900 m.
Type: M. Kessler et at. 9557. HT: SEL; IT: LPB.
Bolivia.
Voucher: M. Kessler et at. 9557 (LPB, SEL).
Endemic.
Tillandsia koehresiana Ehlers, Die Bromelie 3/
1990: 59. 1990 [Jan. 1991].
Dept.: CO, PO.
Elevation: 2000-2750 m.
Type: K6hres S.n. HT: WU; IT: WU. Bolivia.
Voucher: P. lbisch 216 (LPB, SEL).
Endemic.
Tillandsia krukoffiana L.B. Sm., Contr. Gray Herb.
154: 36. 1945.
Dept.: BE, CO, LP, SC?
Elevation: 850-2200(-2700) m.
Type: B. Krukoff 10503. HT: GH;. IT: K, MICH, NY,
S. Bolivia.
Voucher: M. Kessler et al. 4495 (LPB, SEL).
Endemic.
Tillandsia kuehhasii W. Till, Die Bromelie 2/1995:
33. 1995.
Dept.: CH, PO?
Elevation: 3600 m.
Type: E. Kirschnek & F. Kuehhas S.n. HT: WU; IT:
WU. Bolivia.
Voucher: R. Ehlers 950601 (WU).
Endemic.
Tillandsia kuntzeana Mez, in C. DC., Monogr.
phan. 9: 790. J 896.
Dept.: CO, LP?
Elevation: 1300-2100(-3050) m.
Type: Kuntze s.n. HT: NY. Bolivia.
Voucher: M. Kessler et al. 6675 (LPB, SEL).
Endemic.
Tillandsia laxissima Mez, BulL Herb. Boissier, ser.
2, 5: 108. 1905,
Dept.: SC, LP.
Elevation: 1250-1300 m.
Type: Bang 2301 (in part). HT: B; IT: G, GH, LE,
M, MO, NY, PH, US, W, WU. Bolivia.
219
Voucher: M. Kessler 4064 (LPB).
Endemic.
Tillandsia loliacea Mart. ex Schult. & Schult.f., in
Roem. & Schult., Syst. veg. 7(2): 1204. 1830.
Dept.: CH, CO, LP, SC, TA.
Elevation: 200-1400(-2200) m.
Type: C. Martius s.n. HT: M. Brazil.
Voucher: M. Kessler et al. 5197 (LPB, SEL).
Tillandsia longifolia Baker, Handb. Bromel. 185.
1889.
Dept.: CO, SC?
Elevation: 2000-2100(-2600) m.
Type: A. Fendler 1522. HT: K; IT: GH. Venezuela.
Voucher: P. 1bisch & c. 1bisch 93.1628 (SEL).
Tillandsia lorentziana Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 19: 271.1874.
Synonym(s): Tillandsia lorentziana f. simplex 0.
Kuntze, Revis. gen. pl. 3(2): 304. 1898.
Dept.: CH, CO, LP, PO, SC, TA.
Elevation: 350-3300 m.
Type: Lorentz 69. HT: B; IT: CORD. Argentina.
Voucher: P. 1bisch & c. 1bisch 93.0208 (LPB, SEL).
Tillandsia lotteae H. Hrom., J. Bromeliad Soc. 28:
263. 1978.
Dept.: CH, CO, PO, SC.
Elevation: (900-)1550-2900 m.
Type: H. Hromadnik PB 77. HT: HElD; IT: HAL.
Bolivia.
Voucher: M. Kessler et at. 4715 (LPB, SEL).
Endemic.
Tillandsia mandonii E. Morren ex Mez, in C. DC.,
Monogr. phan. 9: 871. 1896.
Dept.: LP, TA.
Elevation: 1400-2650 m.
Type: Mandon 1180. LT: not selected; ILT:
BM,GH,K,PI,S,W. Bolivia.
Voucher: W. Till 73 (WU).
Tillandsia moms H. Hrom. & W. Till, PI. Syst.
Evol. 142: 123. 1983.
Dept.: TA.
Elevation: 2700 m.
Type: H. Hromadnik 9088. HT: WU; IT: B, HAL,
K, LI, NY, W, WU. Bolivia.
Voucher: E. Heger s.n. (WU).
Endemic.
Tillandsia muhriae W. Weber, Feddes Repert. 97:
101. 1986.
Synonyrn(s): Tillandsia alberi Ehlers, Die Bromelie
4/87: 55. 1987.
Dept.: PO.
Elevation: 2650-3400 m.
Type: Muhr s.n. HT: HAL. Argentina.
Voucher: E. Haugg 7029 (WU).
Tillandsia myosura Griseb. ex Baker, J. Bot. 16:
240. 1878.
Dept.: CH, CO, PO, TA.
Elevation: 1900-3200 m.
Type: Lorentz 122. HT: BM; IT: B, NY. Argentina.
Voucher: S. Beck & M. Libermann 14268 (LPB,
SEL).
Tillandsia nana Baker, Handb. Bromel. 172. 1889.
220
Volume 20(2) 1999
SELBYANA
Synonym(s): Tillandsia calocephala Wittm., Meded. Rijks-Herb. 29: 90. 1916.
Dept.: CO, LP.
Elevation: 3150 m.
Type: Gay s.n. HT: P. Peru.
Voucher: Shepard 234 (GH, US). cited for Bolivia
in FI. Neotrop. Monogr. 14(2): 807. 1977.
Tillandsia oropezana L. Hrom., Die Bromelie 1/
89: 14. 1989.
Dept.: CH.
Elevation: 1800 m.
Type: H. Hromadnik 9012. HT: WU; IT: WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Tillandsia paleacea C. PresJ, Re1iq haenk. 1(2):
125. 1827.
Synonym(s): Tillandsia scalarifolia Baker, I. Bot.
25: 235. 1887.
Dept.: LP.
Elevation: 2600 m.
Type: Haenke S.n. HT: PR. Chile.
Voucher: Julio 158 (BM, US). Cited for Bolivia in:
FI. Neotrop. Monogr. 14(2): 863. 1977.
Tillandsia paraensis Mez, in Mart., FI. bras. 3(3):
586. 1894.
Dept.: LP, PA, Sc.
Elevation: 150-900 m.
Type: Sieber 68. HT: BM. Brazil.
Voucher: T. Kromer & A. Acebey 187 (LPB, SEL).
Tillandsia pedicellata (Mez) A. Cast., Lilloa 11:
144. 1945. Basionym: Tillandsia coarctata Gillies ex Baker vaT. pedicellata Mez, in C. DC.,
Monoge phan. 9: 865. 1896.
Dept.: CH, LP, TA.
Elevation: 2200-3000 m.
Type: Hieronymus 795. LT: B; ILT: BM, CORD,
GH, GOET. Argentina.
Voucher: B. Holst et ai. 4958 (USZ).
Tillandsia platyrachis Mez, in C. DC., Monoge
phan. 9: 848. 1896.
Dept.: LP.
Elevation: 900 m.
Type: Kalbreyer 1328. HT: K. Colombia.
Voucher: T. Kromer et ai. 199 (GOET, LPB, SEL).
Tillandsia pohliana Mez, in Mart., FI. bras. 3(3):
597. 1894.
Dept.: BE, CH, LP, Sc.
Elevation: 250-1700 m.
Type: Pohl 3658. HT: W (lost). Brazil.
Voucher: M. Kessler et ai. 4845 (LPB, SEL).
Tillandsia polystachia (L.) L., Sp. pI. (ed. 2) 1:
410. 1762. Basionym: Renealmia polystachia
L., Sp. pI. 1: 286. 1753.
Dept.: BE, LP, Sc.
Elevation: 250-1200 m.
Type: Plumier S.n. HT: P. West Indies.
Voucher: M. Kessler et ai. 4157 (LPB, SEL).
Tillandsia pseudocardenasii W. Weber, Feddes Repert. 97: 103. 1986.
Dept.: LP, Sc.
Elevation: 2300 m.
Type: H. Hromadnik & L. Hromadnik 5323. HT:
HAL; IT: WU. Bolivia.
Voucher: K Uhlig S.n. (WU).
Endemic.
Tillandsia pyramidata Andre, Bromel. Andr. 86.
1889; emend. Rauh, Trop. Subtrop. 3: 15. 1973.
Dept.: LP.
Elevation: 2400 m.
Type: Andre 1522. HT: K. Colombia.
Voucher: S. Beck 3142 (LPB). Del. L.B. Sm.
Tillandsia rauschii Rauh & Lehmann, Trop. Subtrop. Pflanzenwelt 21: 41. 1977.
Dept.: CH?
Elevation: 2500? m.
Type: Bot. Garten Heidelberg 31623 ( = Rausch
s.n.). HT: HElD; IT: LPB, WU. Bolivia.
Voucher: Only known from type collection, without
specific locality.
Endemic.
Tillandsia rectangula Baker, J. Bot. 16: 238. 1878.
Dept.: TA.
Elevation: 500-2100 m.
Type: Lorentz 127. HT: K. Argentina.
Voucher: E. Bastian 1295 (LPB).
Tillandsia recurvata (L.) L., Sp. pI. (ed. 2) 1: 410.
1762. Basionym: Renealmia recurvata L., Sp.
pI. 1: 287. 1753.
Synonym(s): Tillandsia recurvata f. brevifolia Andre, Bromel. Andr. 65. 1889.
Dept.: BE, CH, CO, LP, SC, TA.
Elevation: 200-3100 m.
Type: Sloane s.n. HT: BM; IT: B, GOET, M, P. Jamaica.
Voucher: M. Kessler et ai. 6050 (LPB, SEL).
Tillandsia recurvifolia Hook., Bot. Mag. 87: pI.
5246. 1861.
Synonym(s): Tillandsia meridionalis auct. non Baker: sensu Smith & Downs, FI. Neotrop. Monoge
14(2): 820. 1977.
Dept.: CH, SC, TA.
Elevation: 1200-1550 m.
Type: Original description and plate.
Voucher: W. Till 156 (WU).
Tillandsia recurvispica L. Hrom. & P. Schneid.,
Die Bromelie 1/87: L 1987.
Dept.: CO.
Elevation: 2500-2600 m.
Type: H. Hromadnik & L. Hromadnik 3007. HT:
WU; IT: LPB, WU. Bolivia.
Voucher: H. Hromadnik & L. Hromadnik 5330
(WU).
Endemic.
Tillandsia reichenbachii Baker, Handb. BromeL
166. 1889.
Synonym(s): Tillandsia herzogii Wittm., Meded.
Rijks-Herb. 29: 89. 1916.
Dept.: CH, CO, SC, TA.
Elevation: 200-2500 m.
Type: Reichenbach icon; with no specific locality.
(K).
Voucher: R. Martinez 591 (LPB, SEL).
KROMER ET AL.: BOLIVIAN BROMELIACEAE
Tillandsia rhomboidea Andre, Enum, Bromel. 6.
13 Dec. 1888; Rev. Hort. 60: 566. 16 Dec.
1888.
Dept.: BE.
Elevation: 700-800 m.
Type: Andre 2745. LT: K; IT: NY. Colombia.
Voucher: S. Beck 1732 (US).
Tillandsia rubella Baker, J. Bot. 26: 44. 1888.
Dept.: CO, LP, Sc.
Elevation: 1800-3000 m.
Type: Mandon 1187. HT: K; IT: GH, P. Bolivia.
Voucher: M. Kessler et al. 4496 (LPB, SEL).
Tillandsia rusbyi Baker, Handb. Bromel. 176.
1889.
Dept.: LP, Sc.
Elevation: 850-1800 m.
Type: H. Rusby 2163. HT: NY. Bolivia.
Voucher: T. Kromer & A. Acebey 183 (LPB, SEL).
Tillandsia samaipatensis W. Till, Die Bromelie 21
1996: 36. Feb. 1997.
Dept.: LP, Sc.
Elevation: 850-1350 m.
Type: W. Till 149/2. HT: LPB; IT: SEL, WU. Bolivia.
Voucher: M. Kessler et al. 4112 (LPB, SEL).
Endemic.
Tillandsia schreiteri Lillo & A. Cast., Anales Mus.
Nac. Hist. Nat. Buenos Aires 36: 56. 1929.
Dept.: TA.
Elevation: 1800 m.
Type: Schreiter S.n. HT: BA. Argentina.
Voucher: H. Hromadnik 9099 (WU).
Tillandsia sphaerocephala Baker, J. Bot. 26: 141.
1888.
Dept.: CH, CO, LP, SC, TA.
Elevation: 1400-3750 m.
Type: Mandon 1188. HT: K; IT: G. Bolivia.
Voucher: M. Kessler et al. 5525 (LPB, SEL).
Tiliandsia spiralipetala Gouda, J. Bromeliad Soc.
36: 165. 1986. Basionym: Tillandsia tricholepis
Baker var. macrophylla L.B. Sm., Lilloa 14: 98.
1948.
Dept.: CH, LP, Sc.
Elevation: 350-1450(-2300) m.
Type: Buchtien 1270. HT: US; IT: E. Bolivia.
Voucher: M. Kessler et al. 4059 (LPB).
Tillandsia stenoura Harms, NotizbL Bot. Gart.
Berlin-Dahlem 12: 537. 1935.
Dept.: CO.
Elevation: 2100-2900 m.
Type: Diels 558. HT: B. Ecuador.
Voucher: P. Ibisch & c. lbisch 93.0098 (SEL).
Tillandsia streptocarpa Baker, J. Bot. 25: 241.
1887.
Dept.: CO, CH, LP, PO, Sc.
Elevation: 400-2700 m.
Type: Balansa 615. HT: K; IT: P, S. Paraguay.
Voucher: M. Kessler et al. 4800 (LPB, SEL).
Tillandsia superba Mez & Sodiro, Bull. Herb.
Boissier, ser. 2, 4: 1132. 1904.
Dept.: CO, LP.
221
Elevation: 2350-3250 m.
Type: Sodiro 37-D. HT: B. Ecuador.
Voucher: R. Seidel 7015 (LPB, SEL).
Tillandsia tenuifolia L., Sp. pI. 1: 286. 1753.
Dept.: CH, CO, LP, SC, TA.
Elevation: 450-2800 m.
Type: Royen s.n. HT: L. Country unknown.
Voucher: R. Seidel 7011 (LPB, SEL).
Tillandsia tovarensis Mez, in C. DC., Monogr.
phan. 9: 769. 1896.
Dept.: CO, LP.
Elevation: 2250-2700 m.
Type: A. Fendler 2446. HT: GOET. Venezuela.
Voucher: M. Kessler et al. 9677 (LPB, SEL).
Tillandsia tricholepis Baker, J. Bot. 16: 237. 1878.
Dept.: BE, CH, CO, LP, SC, TA.
Elevation: 200-2600 m.
Type: Mandon 1179 in part. HT: K; IT: BM, NY, P,
S. Bolivia.
Voucher: S. Beck 3200 (LPB, SEL). Det. L.B. Sm.
Tillandsia usneoides (L.) L., Sp. pI. (ed. 2) 1: 411.
1762. Basionym: Renealmia usneoides L., Sp.
pI. 1: 287. 1753.
Dept.: CH, CO, LP, SC, TA.
Elevation: 1100-3800 m.
Type: Anonymous in Clifford Herbarium s.n. LT:
BM. Jamaica.
Voucher: M. Kessler et al. 4902 (LPB, SEL).
Tillandsia variabilis SchltdL, Linnaea 18: 418.
1844.
Synonym(s): Tillandsia valenzuelana A. Rich. in
Sagra, Hist. fis. Cuba, Bot. 11: 267. 1850.
Dept.: CO, LP, Sc.
Elevation: 700-2200 m.
Type: Schiede s.n. LT: HAL; ILT: HAL. Mexico.
Voucher: S. Beck 12613 (LPB, SEL).
Tillandsia vernicosa Baker, J. Bot. 25: 241. 1887.
Dept.: CH, CO, LP, SC, TA.
Elevation: 200-2400 m.
Type: Christie s.n. HT: K. Argentina.
Voucher: M. Kessler et al. 4512 (LPB, SEL).
Tillandsia violascens Mez, in C. DC., Monogr.
phan. 9: 797. 1896.
Dept.: CO, LP, SC.
Elevation: 1800-3500 m.
Type: Mandon 1174 in part. HT: P; IT: BM. Bolivia.
Voucher: M. Kessler et al. 9025 (LPB, SEL).
Tillandsia virescens Ruiz & Pav., FI. Peruv. 3: 43.
1802.
Synonym(s): Tillandsia capillaris f. virescens (Ruiz
& Pavon) L.B. Sm., Proc. Amer. Acad. Arts 70:
212. 1935.
Dept.: CH, CO, LP, PO, SC, TA.
Elevation: 1300-4000 m.
Type: Ruiz & Pavon S.n. LT: BM. Peru.
Voucher: M. Kessler et al. 5911 (LPB, SEL).
Tillandsia walter-richteri W.. Weber, J. Bromeliad
Soc. 34: 102. 1984.
Dept.: TA.
Elevation: 1800 m.
SELBYANA
222
Type: H. Hromadnik & E. Hromadnik 5142. HT:
HAL; IT: WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Tillandsia walteri Mez, Repert. Spec. Nov. Regni
Veg. 3: 43. 1906.
Dept.: CO, LP.
Elevation: 2500-3000 m.
Type: Weberbauer 4319. HT: B. Peru.
Voucher: M. Kessler et al. 12509 (LPB, SEL).
Tillandsia xiphloides Ker-Gawler, Bot. Reg. 2: pl.
105. 1816.
Dept.: CH, CO, LP, PO, TA.
Elevation: 800-3200 m.
Type: Original description and plate.
Voucher: M. Cardenas 93 (GH, SIlo
Tillandsia yuncharaensis W. Till, J. Bromeliad
Soc. 40: 214. 1990.
Dept.:TA.
Elevation: 3000 m.
Type: w: Till et al. sub WT 55. HT: WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Tillandsia sp. 1.
Synonym(s): Tillandsia bryoides auct. non Griseb.
ex Baker: Smith and Downs, Fl. Neotrop. Monogr. 14: 879. 1977; Brako and Zarucchi, Mon·
ogr. Syst. Bot. Missouri Bot. Gard. 45: 250.
1993; Zuloaga and Morrone, Monogr. Syst.
Bot. Missouri Bot. Gard. 60: 114. 1996.
Tillandsia bryoides Grisebach, Abh. Konig!. Ges.
Wiss. Gottingen 24: 334. 1879, in part; Tillandsia coarctata Gillies ex Mez, in C. DC., Monogr. phan. 9: 865. 1896, in part.
Dept.: CR, CO, LP, OR, PO, SC, TA.
Elevation: (400-)1200-3000 m.
Voucher: M. Cardenas 3309 (GH).
Note: This taxon has often been identified as Tillandsia bryoides, but appears to be a fairly
widespread, undescribed species.
VRIESEA
Vriesea cf. chrysostachys E. Morren, Belgique
Hort. 31: 87. 1881.
Dept.: LP.
Elevation: 900 m.
Type: Davis in Hort. Veitch s.n. HT: LG. Peru.
Voucher: M. Kessler et al. 10146 (LPB, SEL).
"cf.".
Vriesea chontalensis (Baker) L.B. Sm., Contr. U.S.
Nat!. Herb. 29: 518. 1951. Basionym: Tillandsia chontalensis Baker, J. Bot. 25: 237. 1887.
Dept.: LP.
Elevation: 1300 m.
Type: R. Tate 413. HT: BM. Nicaragua.
Voucher: M. Kessler et al. 9872 (LPB).
Vriesea friburgensis Mez, in Mart., FI. bras. 3(3):
537. 1894.
Dept.: CH, SC, TA.
Elevation: 500-1350 m.
Volume 20(2) 1999
Type: Glaziou 16467. HT: K; IT: C, LE, P. Brazil.
Voucher: B. Holst et al. 4887 (SEL, USZ).
Vriesea heliconioides (Kunth) Hook. ex WaIp.,
Ann. Bot. Syst. 3: 623. 1852. Basionym: Tillandsia heliconioides Kunth, Nov. gen. sp. 1:
293. 1816.
Dept.: BE, LP, SC.
Elevation: 200-500 m.
Type: Humboldt & Bonpland s.n. HT: P. Colombia.
Voucher: J. Solomon 6431 (LPB, SEL).
Vriesea heterandra (Andre) L.B. Sm., Contr. U.S.
Nat!. Herb. 29: 443. 1951. Basionym: Tillandsia heterandra Andre, Enum. Bromel. 7. 13
Dec. 1888; Rev. Hort. 60: 567. 16 Dec. 1888.
Dept.: LP, SC, TA.
Elevation: 600-1550 m.
Type: Andre 1213. HT: K. Colombia.
Voucher: M. Kessler 9875 (LPB, SEL).
Vriesea incurva (Griseb.) Read, Phytoiogia 16:
458. 1968. Basionym: Tillandsia incurva Griseb., Nachr. Konigl. Ges. Wiss. Georg-AugustsUniv. [II: 15 . 1864.
Dept.:BE?,LP.
Elevation: 700-950 m.
Type: A. Fendler 1524. HT: GOET; IT: GH. Venezuela.
Voucher: Buchtien 373 (US). Cited for Bolivia in
Fl. Neotrop. Monogr. 14 (2): 1251. 1977.
Vriesea maxoniana (L.B. Sm.) L.B. Sm., Phytologia 6: 441. 1959. Basionym: Tillandsia maxoniana L.B. Sm., Contr. Gray Herb. 124: 11.
1939.
Dept.: CH, LP, SC, TA.
Elevation: 500-1550 m.
Type: Buchtien 4538. HT: US. Bolivia.
Voucher: B. Holst et al. 4801 (USZ, SEL).
Vriesea platynema Gaudich., Voy. Bonite, Bot. pI.
66. 1843.
Dept.: LP.
Elevation: 1050 m.
Type: Gaudichaud s.n. HT: P. Brazil.
Voucher: M. Kessler et al. 10196 (LPB, SEL).
Vriesea rubra (Ruiz & Pav.) Beer, Fam. BromeI.
98. 1856. Basionym: Tillandsia rubra Ruiz &
Pav., Fl. peruv. 3: 40, pI. 266. 1802.
Dept.: BE, LP.
Elevation: 450-850 m.
Type: Ruiz & Pavon s.n. HT: MA. Peru.
Voucher: T. Kromer et al. 1 (LPB, SEL).
WERAUHlA
Werauhia boliviana H. Luther, Selbyana 19: 218.
1998 [1999].
Dept.: LP.
Elevation: 950 m.
Type: M. Kessler 3907. HT: SEL; IT: LPB. Bolivia.
Voucher: Only known from type collection.
Endemic.
Werauhia gladioliflora (H. Wend!.) J.R. Grant,
Trop. Subtrop. Pflanzenwelt 91: 31. 1995. Bas-
KROMER ET AL.: BOLIVIAN BROMELIACEAE
ionym: Tillandsia gladioliflora Wendland,
Hamburger Garten-Blumenzeitung 19: 31.
1863.
Dept.: CO, LP.
Elevation: 350-1300 Ill.
Type: Wendland in Herrenhausen Hortus s.n. HT:
HBG? Costa Rica.
Vouchet: M. Kessler 9874 et al. (LPB, SEL).
Werauhia sanguinolenta (Linden ex Cogn. & Mar-
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223
chat) l.R. Grant, Trop. Subtrop. Pflanzenwelt
91: 35. 1995. Basionym: Vriesea sanguinolenta
Linden ex Cogn. & Marchal, Pl. ornem. 2: pi.
52. 1874.
Dept.: SC.
Elevation: ca. 400 m.
Type: E. Morren in Liege Hortus s.n. CT?: LG. Costa Rica.
Voucher: M. Kessler et al. 8642 (LPB, SEL).