Zootaxa 2260: 1–16 (2009)
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ISSN 1175-5326 (print edition)
Copyright © 2009 · Magnolia Press
ISSN 1175-5334 (online edition)
ZOOTAXA
ZOOTAXA
2260
Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef
J.K. LOWRY & A.A. MYERS (EDS)
1
Crustacea Section, Australian Museum, 6 College St., Sydney 2010, Australia
(jim.lowry@austmus.gov.au)
2
Department of Zoology, Ecology and Plant Science, National University of Ireland, Cork, Ireland
(bavayia@googlemail.com)
Magnolia Press
Auckland, New Zealand
J.K. LOWRY & A.A. MYERS (EDS)
Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef
(Zootaxa 2260)
930 pp.; 30 cm.
8 Oct. 2009
ISBN 978-1-86977-411-0 (hardback)
ISBN 978-1-86977-412-7 (Online edition)
FIRST PUBLISHED IN 2009 BY
Magnolia Press
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other than private research use.
ISSN 1175-5326
(Print edition)
ISSN 1175-5334
(Online edition)
2 · Zootaxa 2260 © 2009 Magnolia Press
LOWRY & MYERS (EDS)
Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef
J.K. LOWRY & A.A. MYERS
1
Crustacea Section, Australian Museum, 6 College St., Sydney 2010, Australia.
(jim.lowry@austmus.gov.au)
2
Department of Zoology, Ecology and Plant Science, National University of Ireland, Cork, Ireland.
(bavayia@googlemail.com)
Table of Contents
Foreword. By J.K. Lowry & A.A. Myers .......................................................................................................................... 17
Introduction................................................................................................................................................................ 17
Lizard Island Workshop ............................................................................................................................................ 18
The Florida Workshop............................................................................................................................................... 18
The Amphipods of the GBR ...................................................................................................................................... 19
Habitats ...................................................................................................................................................................... 19
Material and methods ................................................................................................................................................ 21
Acknowledgements.................................................................................................................................................... 21
References.................................................................................................................................................................. 22
Colour Plates of Selected GBR Amphipoda.............................................................................................................. 33
Checklist of the Amphipoda of Tropical Australia.................................................................................................... 39
Station List................................................................................................................................................................. 45
The Biogeography of Indo-West Pacific Tropical Amphipods with particular reference to Australia. By A.A. Myers &
J.K. Lowry ............................................................................................................................................................... 109
Abstract.................................................................................................................................................................... 109
Introduction.............................................................................................................................................................. 109
World distribution patterns of the Amphipoda........................................................................................................ 109
Palaeobiogeography of the Amphipoda................................................................................................................... 118
The formation of the Australian amphipod fauna.................................................................................................... 122
Parsimony Analysis of endemicity of Indo-Pacific amphipods .............................................................................. 123
Discussion................................................................................................................................................................ 124
References................................................................................................................................................................ 125
Amaryllididae. By J.K. Lowry & H.E. Stoddart .............................................................................................................. 128
Abstract.................................................................................................................................................................... 128
Introduction.............................................................................................................................................................. 128
Material and Methods .............................................................................................................................................. 128
Bamarooka Lowry & Stoddart, 2002 ...................................................................................................................... 128
Bamarooka tropicalis Lowry & Stoddart, 2002 ...................................................................................................... 128
References................................................................................................................................................................ 131
Ampeliscidae. By Rachael A. King ................................................................................................................................. 132
Abstract.................................................................................................................................................................... 132
Introduction.............................................................................................................................................................. 132
Material and Methods .............................................................................................................................................. 132
Ampelisca Krøyer, 1842 .......................................................................................................................................... 133
Ampelisca dimboola Lowry & Poore, 1985 ............................................................................................................ 133
Ampelisca dingaal sp. nov. ..................................................................................................................................... 135
Ampelisca jiigurru sp. nov. ..................................................................................................................................... 138
References................................................................................................................................................................ 142
Amphilochidae. By B.A.R. Azman ................................................................................................................................. 143
Abstract.................................................................................................................................................................... 143
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Introduction.............................................................................................................................................................. 143
Material and Methods .............................................................................................................................................. 144
Amphilochus Bate, 1862 .......................................................................................................................................... 144
Amphilochus justi sp. nov. ...................................................................................................................................... 144
Amphilochus lacertus sp. nov. ................................................................................................................................ 147
References................................................................................................................................................................ 151
Ampithoidae. By L.E. Hughes & J.K. Lowry ................................................................................................................. 153
Abstract.................................................................................................................................................................... 153
Introduction.............................................................................................................................................................. 153
Material and Methods .............................................................................................................................................. 153
Ampithoe Leach, 1814 ............................................................................................................................................. 154
Ampithoe cookana Peart, 2007 ................................................................................................................................ 154
Ampithoe katae Peart, 2007 ..................................................................................................................................... 157
Ampithoe kava (Myers, 1985).................................................................................................................................. 161
Ampithoe meganae Peart, 2007 ............................................................................................................................... 164
Ampithoe waialua J.L. Barnard, 1970 ..................................................................................................................... 167
Cymadusa Savigny, 1816 ........................................................................................................................................ 171
Cymadusa alyxis Hughes & Lowry sp. nov. .......................................................................................................... 171
Cymadusa cavimana (Sivaprakasam, 1971) ........................................................................................................... 174
Cymadusa heronensis Peart, 2007........................................................................................................................... 178
Cymadusa hoeyae Hughes & Lowry sp. nov. ........................................................................................................ 181
Cymadusa imbroglio Rabindranath, 1972 .............................................................................................................. 184
Cymadusa khbarnardi Hughes & Lowry sp. nov. ................................................................................................. 188
Cymadusa mariabyrneae Hughes & Lowry sp. nov. ............................................................................................. 191
Cymadusa smilodonta Hughes & Lowry sp. nov. .................................................................................................. 194
Cymadusa tattersalli Peart, 2004............................................................................................................................. 198
Cymadusa thagaay Peart, 2007 ............................................................................................................................... 201
Cymadusa wistari Peart, 2007 ................................................................................................................................. 204
Paragrubia Chevreux, 1901 .................................................................................................................................... 207
Paragrubia edgari Peart sp. nov. ........................................................................................................................... 207
Plumithoe Barnard & Karaman, 1991 ..................................................................................................................... 211
Plumithoe quadrimana (Haswell, 1879).................................................................................................................. 211
Sunamphitoe Bate, 1857 .......................................................................................................................................... 214
Sunamphitoe fantome Peart sp. nov. ...................................................................................................................... 214
References................................................................................................................................................................ 217
Aoridae. By Alan A. Myers ............................................................................................................................................ 220
Abstract.................................................................................................................................................................... 220
Introduction.............................................................................................................................................................. 220
Material and Methods .............................................................................................................................................. 220
Aoroides Walker, 1898 ............................................................................................................................................ 221
Aoroides palfreyensis sp. nov. ................................................................................................................................ 221
Aoroides parvus sp. nov. ........................................................................................................................................ 223
Autonoe Bruzelius, 1859.......................................................................................................................................... 226
Autonoe seurati (Chevreux, 1907) nov. comb. ...................................................................................................... 226
Bemlos Shoemaker, 1925......................................................................................................................................... 228
Bemlos aequimanus (Schellenberg, 1938)............................................................................................................... 229
Bemlos australis (Haswell, 1879)............................................................................................................................ 231
Bemlos bidens Myers, 1988..................................................................................................................................... 234
Bemlos clypeatus Krapp-Schickel & Myers, 2006 .................................................................................................. 237
Bemlos ephippium Myers, 1988 .............................................................................................................................. 239
Bemlos gladius sp. nov. .......................................................................................................................................... 242
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Bemlos mollis Myers, 1988...................................................................................................................................... 245
Bemlos triangulum Myers, 1988.............................................................................................................................. 247
Bemlos tridentatus Myers, 1988 .............................................................................................................................. 250
Bemlos tui (Myers, 1985) ........................................................................................................................................ 253
Bemlos waipio (J.L. Barnard, 1970) ........................................................................................................................ 255
Globosolembos Myers, 1985 ................................................................................................................................... 258
Globosolembos excavatus (Myers, 1975)................................................................................................................ 258
Globosolembos longispinosus Ren, 2006 ................................................................................................................ 260
Globosolembos ovatus Myers, 1985........................................................................................................................ 263
Globosolembos springthorpei sp. nov. ................................................................................................................... 265
Globosolembos varanus sp. nov. ............................................................................................................................ 268
Grandidierella Coutière, 1904 ................................................................................................................................ 271
Grandidierella bonnieroides Stephensen, 1947 ...................................................................................................... 271
Grandidierella rhizophorae sp. nov. ...................................................................................................................... 274
References................................................................................................................................................................ 276
Bogidiellidae. By Charles Oliver Coleman ..................................................................................................................... 279
Abstract.................................................................................................................................................................... 279
Introduction.............................................................................................................................................................. 279
Material and Methods .............................................................................................................................................. 279
Xystriogidiella Stock, 1984 ..................................................................................................................................... 279
Xystriogidiella juliani sp. nov. ............................................................................................................................... 279
Xystriogidiella capricornea Stock, 1984................................................................................................................. 282
References................................................................................................................................................................ 284
Bolttsiidae. By B.A.R. Azman & J.K. Lowry ................................................................................................................. 285
Abstract.................................................................................................................................................................... 285
Introduction.............................................................................................................................................................. 285
Material and Methods .............................................................................................................................................. 285
Bolttsia Griffiths, 1976 ............................................................................................................................................ 286
Bolttsia myersi sp. nov. ........................................................................................................................................... 286
References................................................................................................................................................................ 289
Caprellidae. By José M. Guerra-García & J.K. Lowry ................................................................................................... 290
Abstract.................................................................................................................................................................... 290
Introduction.............................................................................................................................................................. 290
Material and Methods .............................................................................................................................................. 290
Caprellinae
Aciconula Mayer, 1903............................................................................................................................................ 291
Aciconula australiensis Guerra-García, 2004.......................................................................................................... 291
Caprella Lamarck, 1801.......................................................................................................................................... 293
Caprella penantis Leach, 1814................................................................................................................................ 293
Hemiaegina Mayer, 1890 ........................................................................................................................................ 295
Hemiagina minuta Mayer, 1890 .............................................................................................................................. 295
Metaprotella Mayer, 1890 ....................................................................................................................................... 297
Metaprotella sandalensis Mayer, 1898 ................................................................................................................... 297
Orthoprotella Mayer, 1903...................................................................................................................................... 299
Orthoprotella australis (Haswell, 1880) ................................................................................................................. 299
Orthoprotella pearcei Guerra-García, 2006 ............................................................................................................ 301
Orthoprotella sp. ..................................................................................................................................................... 304
Paraprotella Mayer, 1890 ....................................................................................................................................... 305
Paraprotella sp. ...................................................................................................................................................... 305
Protella Dana, 1852................................................................................................................................................. 307
Protella similis Mayer, 1903 ................................................................................................................................... 307
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Pseudaeginella Mayer, 1890 ................................................................................................................................... 309
Pseudaeginella biscayensis (McCain, 1968) ........................................................................................................... 309
Phtisicinae
Jigurru Guerra-García, 2006..................................................................................................................................... 311
Jigurru vailhoggetti Guerra-García, 2006 ................................................................................................................ 311
Metaproto Mayer, 1903 ............................................................................................................................................ 312
Metaproto novaehollandiae (Haswell, 1880) ........................................................................................................... 312
Perotripus Doughty & Steinberg, 1953 .................................................................................................................... 315
Perotripus keablei Guerra-García, 2006................................................................................................................... 315
Pseudoprellicana Guerra-García, 2006 .................................................................................................................... 317
Pseudoprellicana johnsoni Guerra-García, 2006...................................................................................................... 317
Pseudoproto Mayer, 1903......................................................................................................................................... 319
Pseudoproto fallax Mayer, 1903............................................................................................................................... 319
Quadrisegmentum Hirayama, 1988 .......................................................................................................................... 321
Quadrisegmentum lowryi Guerra-García, 2006........................................................................................................ 321
Quadrisegmentum triangulum Hirayama, 1988........................................................................................................ 323
References................................................................................................................................................................. 325
Ceinidae. By Charles Oliver Coleman ............................................................................................................................ 328
Abstract.................................................................................................................................................................... 328
Introduction.............................................................................................................................................................. 328
Material and Methods .............................................................................................................................................. 328
Ceina Della Valle, 1893 .......................................................................................................................................... 328
Ceina gerlachae sp. nov. ........................................................................................................................................ 328
References................................................................................................................................................................ 332
Cheirocratidae. By Charles Oliver Coleman & James K. Lowry .................................................................................. 333
Abstract.................................................................................................................................................................... 333
Introduction.............................................................................................................................................................. 333
Material and Methods .............................................................................................................................................. 333
Prosocratus Barnard & Drummond, 1982 .............................................................................................................. 334
Prosocratus carolinae sp. nov. .............................................................................................................................. 334
References................................................................................................................................................................ 337
Cheluridae. By Charles Oliver Coleman & Adina Renz ................................................................................................ 339
Abstract.................................................................................................................................................................... 339
Introduction.............................................................................................................................................................. 339
Material and Methods .............................................................................................................................................. 339
Tropichelura J.L. Barnard, 1959 ............................................................................................................................. 340
Tropichelura insulae (Calman, 1910)...................................................................................................................... 340
References................................................................................................................................................................ 343
Chevaliidae. By Alan A. Myers ...................................................................................................................................... 344
Abstract.................................................................................................................................................................... 344
Introduction.............................................................................................................................................................. 344
Material and Methods .............................................................................................................................................. 344
Chevalia Walker, 1904 ............................................................................................................................................ 344
Chevalia tenuis sp. nov. ......................................................................................................................................... 344
References................................................................................................................................................................ 347
Colomastigidae. By Sara E. LeCroy ................................................................................................................................ 348
Abstract.................................................................................................................................................................... 348
Introduction.............................................................................................................................................................. 348
Material and Methods .............................................................................................................................................. 349
Colomastix Grube, 1861 ........................................................................................................................................... 349
Colomastix dentipalma sp. nov. .............................................................................................................................. 349
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Colomastix japonica Bulycheva, 1955 ..................................................................................................................... 352
Colomastix lunililo J.L. Barnard, 1970 ..................................................................................................................... 356
Colomastix plumosa Ledoyer, 1979 ......................................................................................................................... 360
Colomastix processa sp. nov. .................................................................................................................................. 363
Colomastix thomasi sp. nov. .................................................................................................................................... 367
References................................................................................................................................................................. 371
Corophiidae. By Alan A. Myers ...................................................................................................................................... 373
Abstract.................................................................................................................................................................... 373
Introduction.............................................................................................................................................................. 373
Material and Methods .............................................................................................................................................. 373
Corophiinae
Paracorophium Stebbing, 1899 ............................................................................................................................... 373
Paracorophium nana sp. nov. ................................................................................................................................. 373
Protomedeiinae
Pumiliophotis gen. nov. ........................................................................................................................................... 376
Pumiliophotis queenslandicus sp. nov. ................................................................................................................... 376
References................................................................................................................................................................. 379
Cyproideidae. By B.A.R. Azman .................................................................................................................................... 380
Abstract.................................................................................................................................................................... 380
Introduction.............................................................................................................................................................. 380
Material and Methods .............................................................................................................................................. 380
Cyproidea Haswell, 1879 ........................................................................................................................................ 381
Cyproidea cobia sp. nov. ........................................................................................................................................ 381
Gbroidea Lowry & Azman, 2008............................................................................................................................ 385
Gbroidea dingaalana Lowry & Azman, 2008 ........................................................................................................ 385
Narapheonoides J.L. Barnard, 1972 ........................................................................................................................ 388
Narapheonoides lowryi sp. nov. ............................................................................................................................. 388
References................................................................................................................................................................ 391
Dexaminidae. By Alan A. Myers & Sara E. LeCroy ..................................................................................................... 393
Abstract.................................................................................................................................................................... 393
Introduction.............................................................................................................................................................. 393
Material and Methods .............................................................................................................................................. 394
Dexaminoculus Lowry, 1981 .................................................................................................................................... 394
Dexaminoculus grobbeni (Spandl, 1923) .......................................................................................................... 394
Guernea Chevreux, 1887 .......................................................................................................................................... 397
Guernea ipilya Thomas & Barnard, 1991 ......................................................................................................... 397
Guernea yamminye Thomas & Barnard, 1991 .................................................................................................. 400
Paradexamine Stebbing, 1899 ................................................................................................................................. 403
Paradexamine aequiserrata sp. nov. ................................................................................................................ 403
Paradexamine exilis sp. nov. ............................................................................................................................ 406
Paradexamine levitelson sp. nov. ..................................................................................................................... 408
Paradexamine massa sp. nov. .......................................................................................................................... 411
Paradexamine micronesica Ledoyer, 1979 ....................................................................................................... 413
Paradexamine quadratus sp. nov. .................................................................................................................... 415
Paradexamine saxeta sp. nov. .......................................................................................................................... 418
Polycheria Haswell, 1879 ......................................................................................................................................... 420
Polycheria goanna sp. nov. .............................................................................................................................. 420
References................................................................................................................................................................. 423
Eusiridae. By Charles Oliver Coleman .......................................................................................................................... 425
Abstract.................................................................................................................................................................... 425
Introduction.............................................................................................................................................................. 425
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Material and Methods .............................................................................................................................................. 425
Tethygeneia J.L. Barnard, 1972............................................................................................................................... 425
Tethygeneia aff. pacifica (Schellenberg, 1938)....................................................................................................... 425
References................................................................................................................................................................ 429
Exoedicerotidae. By L.E. Hughes .................................................................................................................................. 430
Abstract.................................................................................................................................................................... 430
Introduction.............................................................................................................................................................. 430
Material and Methods .............................................................................................................................................. 430
Parhalimedon Chevreux, 1906................................................................................................................................ 431
Parhalimedon kyhursti sp. nov. .............................................................................................................................. 431
References................................................................................................................................................................ 433
Hadziidae. By R.T. Springthorpe & J.K. Lowry ............................................................................................................. 434
Abstract.................................................................................................................................................................... 434
Introduction.............................................................................................................................................................. 434
Material and Methods .............................................................................................................................................. 434
Dulzura J.L. Barnard, 1969 ..................................................................................................................................... 434
Dulzura taylorae sp. nov. ....................................................................................................................................... 435
References................................................................................................................................................................ 439
Hyalidae. By Cristiana S. Serejo & Daniela J. Sittrop .................................................................................................... 440
Abstract.................................................................................................................................................................... 440
Introduction.............................................................................................................................................................. 440
Material and Methods .............................................................................................................................................. 440
Hyacheliinae
Hyachelia J.L. Barnard, 1967 ................................................................................................................................... 441
Hyachelia lowryi sp. nov. ........................................................................................................................................ 441
Hyalinae Bulycheva, 1957 ............................................................................................................................................... 444
Lelehua J.L. Barnard, 1970....................................................................................................................................... 444
Lelehua myersi sp. nov. ........................................................................................................................................... 444
Parhyale Stebbing, 1897........................................................................................................................................... 447
Parhyale hawaiensis (Dana, 1853) .......................................................................................................................... 447
References................................................................................................................................................................. 451
Iciliidae. By J.K. Lowry & L.E. Hughes ........................................................................................................................ 453
Abstract.................................................................................................................................................................... 453
Introduction.............................................................................................................................................................. 453
Material and Methods .............................................................................................................................................. 453
Icilius Dana, 1849.................................................................................................................................................... 453
Icilius caledoniana Watson, Lowry & Steinberg, 2004 .......................................................................................... 453
References................................................................................................................................................................ 456
Iphimediidae. By Charles Oliver Coleman .................................................................................................................... 458
Abstract.................................................................................................................................................................... 458
Introduction.............................................................................................................................................................. 458
Material and Methods .............................................................................................................................................. 458
Iphimedia Rathke, 1843........................................................................................................................................... 459
Iphimedia schminkei sp. nov. ................................................................................................................................. 459
References................................................................................................................................................................ 462
Ischyroceridae. By Jean Just .......................................................................................................................................... 463
Abstract.................................................................................................................................................................... 463
Introduction.............................................................................................................................................................. 463
Material and Methods .............................................................................................................................................. 463
Ischyrocerinae
Coxischyrocerus gen. nov. ...................................................................................................................................... 464
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Coxischyrocerus rhombocoxus sp. nov. .................................................................................................................. 464
Tropischyrocerus gen. nov. ..................................................................................................................................... 466
Tropischyrocerus pugilis sp. nov. ........................................................................................................................... 466
Siphonoecetinae
Ambicholestes Just, 1998 .......................................................................................................................................... 469
Ambicholestes (Ambicholestes) magellani (Just, 1984)............................................................................................ 469
Cerapus Say, 1817 .................................................................................................................................................... 470
Cerapus nudus sp. nov. ........................................................................................................................................... 470
Cerapus volucola Lowry & Berents, 2005 ............................................................................................................... 473
Ericthonius Milne Edwards, 1830 ............................................................................................................................ 475
Ericthonius parabrasiliensis sp. nov. ...................................................................................................................... 475
Ericthonius pugnax Dana, 1852................................................................................................................................ 478
Ericthonius tropicalis sp. nov. ................................................................................................................................. 481
Ericthonius sp. ......................................................................................................................................................... 484
References................................................................................................................................................................. 486
Kamakidae. By Alan A. Myers ...................................................................................................................................... 487
Abstract.................................................................................................................................................................... 487
Introduction.............................................................................................................................................................. 487
Material and Methods .............................................................................................................................................. 487
Gammaropsella gen. nov. ...................................................................................................................................... 488
Gammaropsella saepta sp. nov. ............................................................................................................................. 488
Kamaka Derzhavin, 1923 ........................................................................................................................................ 490
Kamaka silvana sp. nov. ........................................................................................................................................ 490
References................................................................................................................................................................ 493
Leucothoidae. By Kristine N. White & James Darwin Thomas .................................................................................... 494
Abstract.................................................................................................................................................................... 494
Introduction.............................................................................................................................................................. 494
Material and Methods .............................................................................................................................................. 495
Anamixis Stebbing, 1897 ......................................................................................................................................... 495
Anamixis bazimut Thomas, 1997............................................................................................................................. 495
Leucothoe Leach, 1814............................................................................................................................................ 498
Leucothoe adelphe sp. nov. .................................................................................................................................... 498
Leucothoe bova sp. nov. ......................................................................................................................................... 501
Leucothoe epidemos sp. nov. .................................................................................................................................. 505
Leucothoe eumilli sp. nov. ...................................................................................................................................... 508
Leucothoe hipposideros sp. nov. ............................................................................................................................ 511
Leucothoe laevipalma sp. nov. ............................................................................................................................... 515
Leucothoe makrommatos sp. nov. .......................................................................................................................... 518
Leucothoe odontiskos sp. nov. ................................................................................................................................ 521
Leucothoe pollex sp. nov. ....................................................................................................................................... 525
Leucothoe rudicula sp. nov. ................................................................................................................................... 528
Leucothoe serrata sp. nov. ..................................................................................................................................... 531
Leucothoe sparsa sp. nov. ...................................................................................................................................... 535
Leucothoe thula sp. nov. ........................................................................................................................................ 538
Leucothoe undulata sp. nov. .................................................................................................................................. 541
Leucothoella Schellenberg, 1928 ............................................................................................................................ 544
Leucothoella gracilis Haswell, 1879 ....................................................................................................................... 544
Paranamixis Schellenberg, 1928 ............................................................................................................................. 548
Paranamixis jiigurru sp. nov. ................................................................................................................................ 548
Unknown leucomorph ............................................................................................................................................. 551
References................................................................................................................................................................ 554
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Liljeborgiidae. By Charles Oliver Coleman ................................................................................................................... 556
Abstract.................................................................................................................................................................... 556
Introduction.............................................................................................................................................................. 556
Material and Methods .............................................................................................................................................. 556
Liljeborgia Bate, 1862 ............................................................................................................................................. 557
Liljeborgia joergpeteri sp. nov. .............................................................................................................................. 557
References................................................................................................................................................................ 560
Lysianassidae. By J.K. Lowry & H.E. Stoddart ............................................................................................................. 561
Abstract.................................................................................................................................................................... 561
Introduction.............................................................................................................................................................. 561
Material and Methods .............................................................................................................................................. 562
Lysianassinae
Azotostoma J.L. Barnard, 1965 ................................................................................................................................. 562
Azotostoma fusta J.L. Barnard, 1965 ........................................................................................................................ 562
Pseudambasia Stephensen, 1927 .............................................................................................................................. 565
Pseudambasia acuticaudata (Ledoyer, 1984) .......................................................................................................... 565
Riwo Lowry & Stoddart, 1995 .................................................................................................................................. 568
Riwo mizeui Lowry & Stoddart, 1995....................................................................................................................... 568
Shoemakerella Pirlot, 1936 ....................................................................................................................................... 571
Shoemakerella barnardi sp. nov. ............................................................................................................................. 571
Waldeckia Chevreux, 1906 ....................................................................................................................................... 575
Waldeckia enoei Stephensen, 1931........................................................................................................................... 575
Tryphosinae Lowry & Stoddart, 1997
Paralysianopsis Schellenberg, 1931 ......................................................................................................................... 578
Paralysianopsis padoz Lowry & Stoddart, 1995 ...................................................................................................... 578
Tryphosella Bonnier, 1893........................................................................................................................................ 581
Tryphosella bet sp. nov. ........................................................................................................................................... 581
Tryphosella cameloides sp. nov. ............................................................................................................................. 584
Tryphosella charlottae sp. nov. ............................................................................................................................... 587
Tryphosella flynnana sp. nov. ................................................................................................................................. 590
Tryphosella seasana sp. nov. ................................................................................................................................... 593
References................................................................................................................................................................. 596
Maeridae, the Ceradocus group. By Traudl Krapp-Schickel........................................................................................... 598
Abstract.................................................................................................................................................................... 598
Introduction.............................................................................................................................................................. 598
Material and Methods .............................................................................................................................................. 598
Austromaera Lowry & Springthorpe, 2005............................................................................................................. 599
Austromaera brevicaudata (Ledoyer, 1984) ........................................................................................................... 599
Ceradocus Costa, 1853............................................................................................................................................ 601
Ceradocus crenatipalma Ledoyer, 1979 ................................................................................................................. 601
Ceradocus hawaiensis J.L. Barnard, 1955 .............................................................................................................. 603
Ceradocus oxyodus Berents, 1983........................................................................................................................... 604
Ceradocus cf. rubromaculatus Stimpson, 1856 ...................................................................................................... 607
Ceradocus woorea Berents, 1983............................................................................................................................ 610
Ceradocus yandala Berents, 1983........................................................................................................................... 612
Glossomaera gen. nov. ........................................................................................................................................... 614
Glossomaera octodens (Sivaprakasam, 1968)......................................................................................................... 614
Jerbarnia Croker, 1971 ........................................................................................................................................... 617
Jerbarnia stocki Thomas & Barnard, 1990 ............................................................................................................. 617
Linguimaera Pirlot, 1936......................................................................................................................................... 620
Linguimaera pirloti Krapp-Schickel, 2003.............................................................................................................. 620
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Maeropsis Chevreux, 1919...................................................................................................................................... 622
Maeropsis cobia sp. nov. ........................................................................................................................................ 622
Maeropsis griffini (Berents, 1983) .......................................................................................................................... 625
Quadrimaera Krapp-Schickel & Ruffo, 2000 ......................................................................................................... 627
Quadrimaera pacifica (Schellenberg, 1938) ........................................................................................................... 627
Quadrimaera quadrimana (Dana, 1853)................................................................................................................. 629
Quadrimaera cf. reishi (J.L. Barnard, 1979)........................................................................................................... 631
Quadrimaera serrata (Schellenberg, 1938) ............................................................................................................ 633
Sauradocus Yerman & Krapp-Schickel, 2008 ........................................................................................................ 636
Sauradocus hobbit Yerman & Krapp-Schickel sp. nov. ........................................................................................ 636
Sauradocus minimarenus Yerman & Krapp-Schickel sp. nov. ............................................................................. 638
References................................................................................................................................................................ 641
Maeridae, the Elasmopus group. By J.K. Lowry & L.E. Hughes ................................................................................. 643
Abstract.................................................................................................................................................................... 643
Introduction.............................................................................................................................................................. 643
Material and Methods .............................................................................................................................................. 643
Elasmopus Costa, 1853 ........................................................................................................................................... 645
Elasmopus alalo Myers, 1986 ................................................................................................................................. 646
Elasmopus crenulatus Berents, 1983....................................................................................................................... 649
Elasmopus hooheno J.L. Barnard, 1970 .................................................................................................................. 652
Elasmopus pectenicrus (Bate, 1862) ....................................................................................................................... 656
Elasmopus slatyeri sp. nov. .................................................................................................................................... 659
Elasmopus spinicarpus Berents, 1983..................................................................................................................... 663
Elasmopus varanocephalensis sp. nov. ................................................................................................................. 666
Mallacoota J.L. Barnard, 1972 ................................................................................................................................ 669
Mallacoota balara Berents, 1983 ............................................................................................................................ 669
Mallacoota capricornia sp. nov. ............................................................................................................................ 673
Mallacoota chandaniae Lowry & Springthorpe, 2005............................................................................................ 677
Mallacoota nananui Myers, 1985 ........................................................................................................................... 680
Mallacoota schellenbergi Ledoyer, 1984 ................................................................................................................ 684
Mallacoota scopulosa sp. nov. ............................................................................................................................... 687
Parelasmopus Stebbing, 1888 ................................................................................................................................. 690
Parelasmopus cymatilis sp. nov. ............................................................................................................................ 691
Parelasmopus suensis (Haswell, 1879b) ................................................................................................................. 695
References................................................................................................................................................................ 699
Maxillipiidae. By Charles Oliver Coleman ................................................................................................................... 703
Abstract.................................................................................................................................................................... 703
Introduction.............................................................................................................................................................. 703
Material and Methods .............................................................................................................................................. 703
Maxillipius Ledoyer, 1973....................................................................................................................................... 704
Maxillipius rectitelson Ledoyer, 1973..................................................................................................................... 704
References................................................................................................................................................................ 707
Megaluropidae. By L.E. Hughes ................................................................................................................................... 708
Abstract.................................................................................................................................................................... 708
Introduction.............................................................................................................................................................. 708
Material and Methods .............................................................................................................................................. 708
Gibberosus Thomas & Barnard, 1986 ..................................................................................................................... 709
Gibberosus udarus sp. nov. ................................................................................................................................... 709
References................................................................................................................................................................ 711
Melitidae, the Eriopisella group. By Michelle N. Yerman ........................................................................................... 713
Abstract.................................................................................................................................................................... 713
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Introduction.............................................................................................................................................................. 713
Material and Methods .............................................................................................................................................. 713
Netamelita J.L. Barnard, 1962 ................................................................................................................................ 714
Netamelita lacerta sp. nov. .................................................................................................................................... 714
References................................................................................................................................................................ 717
Melitidae, the Melita group. By J.K. Lowry & R.T. Springthorpe ............................................................................... 718
Abstract.................................................................................................................................................................... 718
Introduction.............................................................................................................................................................. 718
Material and Methods .............................................................................................................................................. 720
Dulichiella Stout, 1912 ........................................................................................................................................... 720
Dulichiella pacifica Lowry & Springthorpe, 2005.................................................................................................. 720
Melita Leach, 1814 .................................................................................................................................................. 723
Melita myersi G. Karaman, 1987 ............................................................................................................................ 723
Melita sampsonae sp. nov. ..................................................................................................................................... 726
Tegano Barnard & Karaman, 1982.......................................................................................................................... 730
Tegano atkinsae sp. nov. ....................................................................................................................................... 730
References................................................................................................................................................................ 734
Melphidippidae. By Charles Oliver Coleman ............................................................................................................... 736
Abstract.................................................................................................................................................................... 736
Introduction.............................................................................................................................................................. 736
Material and Methods .............................................................................................................................................. 736
Melphisana J.L. Barnard, 1962................................................................................................................................ 737
Melphisana madagascarensis Ledoyer, 1984 ......................................................................................................... 737
References................................................................................................................................................................ 740
Ochlesidae. By Charles Oliver Coleman ...................................................................................................................... 741
Abstract.................................................................................................................................................................... 741
Introduction.............................................................................................................................................................. 741
Material and Methods .............................................................................................................................................. 741
Ochlesis Stebbing, 1910 .......................................................................................................................................... 742
Ochlesis alii J.L. Barnard, 1970 .............................................................................................................................. 742
References................................................................................................................................................................ 745
Oedicerotidae. By L.E. Hughes & J.K. Lowry ............................................................................................................. 746
Abstract.................................................................................................................................................................... 746
Introduction.............................................................................................................................................................. 746
Material and Methods .............................................................................................................................................. 746
Aborolobatea Ledoyer, 1984 .................................................................................................................................. 747
Aborolobatea insidiosa sp. nov. ............................................................................................................................. 747
Monoculodes Stimpson, 1853 ................................................................................................................................. 750
Monoculodes tropicalis sp. nov. ........................................................................................................................... 750
Perioculodes Sars, 1895 ......................................................................................................................................... 754
Perioculodes talboti sp. nov. .................................................................................................................................. 754
References................................................................................................................................................................ 757
Paracalliopiidae. By L.E. Hughes ................................................................................................................................. 759
Abstract.................................................................................................................................................................... 759
Introduction.............................................................................................................................................................. 759
Material and Methods .............................................................................................................................................. 759
Doowia Barnard & Drummond, 1987 .................................................................................................................... 759
Doowia dexterae Barnard & Drummond, 1987 ...................................................................................................... 759
Yhi Barnard & Thomas, 1991 .................................................................................................................................. 762
Yhi yindi Barnard & Thomas, 1991 ......................................................................................................................... 762
References................................................................................................................................................................ 765
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Phliantidae. By Charles Oliver Coleman ...................................................................................................................... 766
Abstract.................................................................................................................................................................... 766
Introduction.............................................................................................................................................................. 766
Material and Methods .............................................................................................................................................. 766
Pereionotus Bate & Westwood, 1863 ..................................................................................................................... 766
Pereionotus thomsoni Stebbing, 1899 ..................................................................................................................... 766
References................................................................................................................................................................ 770
Photidae. By Alan A. Myers ......................................................................................................................................... 771
Abstract.................................................................................................................................................................... 771
Introduction.............................................................................................................................................................. 771
Material and Methods .............................................................................................................................................. 771
Gammaropsis Liljeborg, 1855 ................................................................................................................................. 771
Gammaropsis hephaestis sp. nov. ........................................................................................................................... 771
Graciliphotis gen. nov. ........................................................................................................................................... 774
Graciliphotis ruthae sp. nov. .................................................................................................................................. 774
Latigammaropsis gen. nov. .................................................................................................................................... 777
Latigammaropsis athenae sp. nov. ......................................................................................................................... 777
Latigammaropsis dionysus sp. nov. ....................................................................................................................... 780
Latigammaropsis hermes sp. nov. .......................................................................................................................... 783
Latigammaropsis hestia sp. nov. ............................................................................................................................ 785
Photis Krøyer, 1842................................................................................................................................................. 788
Photis aina J.L. Barnard, 1970 ................................................................................................................................ 788
Photis davei sp. nov. .............................................................................................................................................. 790
Photis pirloti Myers, 1985 ...................................................................................................................................... 793
Virgammaropsis gen. nov. ..................................................................................................................................... 795
Virgammaropsis artemis sp. nov. .......................................................................................................................... 796
References................................................................................................................................................................ 798
Phoxocephalidae. By Joanne Taylor ............................................................................................................................. 800
Abstract.................................................................................................................................................................... 800
Introduction.............................................................................................................................................................. 800
Material and Methods .............................................................................................................................................. 801
Birubius Barnard & Drummond, 1976 .................................................................................................................... 801
Birubius banksi sp. nov. ......................................................................................................................................... 801
Birubius batei (Haswell, 1879)................................................................................................................................ 804
Birubius casuarina sp. nov. .................................................................................................................................... 807
Birubius kingae sp. nov. ......................................................................................................................................... 810
Birubius oti sp. nov. ............................................................................................................................................... 813
Birubius parvus sp. nov. ......................................................................................................................................... 816
Limnoporeia Fearn-Wanan, 1968 ........................................................................................................................... 819
Limnoporeia cooki sp. nov. .................................................................................................................................... 819
Limnoporeia woorake Barnard & Drummond, 1978 .............................................................................................. 821
Metaphoxus Bonnier, 1896...................................................................................................................................... 824
Metaphoxus varanas sp. nov. ................................................................................................................................. 824
References................................................................................................................................................................ 827
Platyischnopidae. By L.E. Hughes ................................................................................................................................ 828
Abstract.................................................................................................................................................................... 828
Introduction.............................................................................................................................................................. 828
Material and Methods .............................................................................................................................................. 828
Platyischnopus Stebbing, 1888 ............................................................................................................................... 828
Platyischnopus mam Barnard & Drummond, 1979................................................................................................. 828
Tomituka Barnard & Drummond, 1979 .................................................................................................................. 832
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Tomituka eumilli sp. nov. ....................................................................................................................................... 832
References................................................................................................................................................................ 835
Pleustidae. By Jean Just ................................................................................................................................................. 836
Abstract.................................................................................................................................................................... 836
Introduction.............................................................................................................................................................. 836
Material and Methods .............................................................................................................................................. 836
Tepidopleustes Karaman & Barnard, 1979.............................................................................................................. 837
Tepidopleustes acromatius sp. nov. ....................................................................................................................... 837
References................................................................................................................................................................ 840
Podoceridae. By Niamh M. Kilgallen ........................................................................................................................... 841
Abstract.................................................................................................................................................................... 841
Introduction.............................................................................................................................................................. 841
Material and Methods .............................................................................................................................................. 841
Podocerus Leach, 1814 ........................................................................................................................................... 842
Podocerus brasiliensis (Dana, 1853)....................................................................................................................... 842
Podocerus casuarinensis sp. nov. .......................................................................................................................... 844
Podocerus chelonophilus (Chevreux & de Guerne, 1888) ..................................................................................... 848
Podocerus crenulatus Myers, 1985 ......................................................................................................................... 850
Podocerus laevis (Haswell, 1885) ........................................................................................................................... 853
Podocerus talegus talegus J.L. Barnard, 1965 ........................................................................................................ 853
Podocerus uncinatus sp. nov. ................................................................................................................................. 856
References................................................................................................................................................................ 859
Sebidae. By Michelle N. Yerman & Charles Oliver Coleman ..................................................................................... 861
Abstract.................................................................................................................................................................... 861
Introduction.............................................................................................................................................................. 861
Material and Methods .............................................................................................................................................. 861
Seba Bate, 1862 ....................................................................................................................................................... 861
Seba gruneri sp. nov. .............................................................................................................................................. 861
Seba mariolgae sp. nov. ......................................................................................................................................... 865
Seba zeumindi sp. nov. ........................................................................................................................................... 868
References................................................................................................................................................................ 871
Stenothoidae. By Traudl Krapp-Schickel ..................................................................................................................... 872
Abstract.................................................................................................................................................................... 872
Introduction.............................................................................................................................................................. 872
Material and Methods .............................................................................................................................................. 872
Stenothoe Dana, 1852 ............................................................................................................................................. 873
Stenothoe miersi (Haswell, 1879)............................................................................................................................ 873
Wallametopa J.L. Barnard, 1974 ............................................................................................................................. 875
Wallametopa cabon J.L. Barnard, 1974 .................................................................................................................. 875
References................................................................................................................................................................ 878
Synopiidae. By L.E. Hughes ......................................................................................................................................... 880
Abstract.................................................................................................................................................................... 880
Introduction.............................................................................................................................................................. 880
Material and Methods .............................................................................................................................................. 880
Synopia Dana, 1852 ................................................................................................................................................ 881
Synopia ultramarina Dana, 1853 ............................................................................................................................ 881
Syrrhoe Goës, 1866 ................................................................................................................................................ 884
Syrrhoe petitaserrata sp. nov. ................................................................................................................................ 884
Telsosynopia G. Karaman, 1986.............................................................................................................................. 887
Telsosynopia trifidilla Hughes & Lowry, 2006....................................................................................................... 887
References................................................................................................................................................................ 890
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Talitridae. By Cristiana S. Serejo .................................................................................................................................. 892
Abstract.................................................................................................................................................................... 892
Introduction.............................................................................................................................................................. 892
Material and Methods .............................................................................................................................................. 892
Chelorchestia Bousfield, 1984 ............................................................................................................................... 893
Chelorchestia limicola (Haswell, 1880) .................................................................................................................. 893
Talorchestia Dana, 1852 ......................................................................................................................................... 896
Talorchestia spinipalma Dana, 1852 ...................................................................................................................... 897
Talorchestia terraereginae Haswell, 1880 ............................................................................................................. 900
References................................................................................................................................................................ 902
Unciolidae. By Alan A. Myers ...................................................................................................................................... 904
Abstract.................................................................................................................................................................... 904
Introduction.............................................................................................................................................................. 904
Material and Methods .............................................................................................................................................. 904
Wombalano Thomas & Barnard, 1991 .................................................................................................................... 904
Wombalano yerang Thomas & Barnard, 1991 ........................................................................................................ 904
References................................................................................................................................................................ 907
Uristidae. By J.K. Lowry & H.E. Stoddart ................................................................................................................... 908
Abstract.................................................................................................................................................................... 908
Introduction.............................................................................................................................................................. 908
Material and Methods .............................................................................................................................................. 908
Ichnopus Costa 1853 ............................................................................................................................................... 909
Ichnopus capricornus Lowry & Stoddart, 1992 ...................................................................................................... 909
Ichnopus tenuicornis (Haswell, 1879).................................................................................................................... 912
Nagada Lowry & Stoddart 1995 ............................................................................................................................ 915
Nagada uwedoae Lowry & Stoddart, 1995 ............................................................................................................. 915
References................................................................................................................................................................ 918
Urohaustoriidae. By Niamh M. Kilgallen ...................................................................................................................... 919
Abstract.................................................................................................................................................................... 919
Introduction.............................................................................................................................................................. 919
Material and Methods .............................................................................................................................................. 919
Tottungus Barnard & Drummond, 1982.................................................................................................................. 920
Tottungus solivagus sp. nov. .................................................................................................................................. 920
Urohaustorius Sheard, 1936 .................................................................................................................................... 922
Urohaustorius drummondae sp. nov. ..................................................................................................................... 922
References................................................................................................................................................................ 926
Wandinidae. By J.K. Lowry & H.E. Stoddart ................................................................................................................ 927
Abstract.................................................................................................................................................................... 927
Introduction.............................................................................................................................................................. 927
Material and Methods .............................................................................................................................................. 927
Wandin Lowry & Stoddart, 1990............................................................................................................................. 927
Wandin griffini Lowry & Stoddart, 1990 ................................................................................................................ 927
References................................................................................................................................................................ 930
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