Echinoderm Research and Diversity
in Latin America
Juan José Alvarado
Francisco Alonso Solís-Marín
Editors
Echinoderm Research and
Diversity in Latin America
123
Editors
Juan José Alvarado
Centro de Investigaciónes en Ciencias
del Mar y Limnologia
Universidad de Costa Rica
San José
Costa Rica
ISBN 978-3-642-20050-2
DOI 10.1007/978-3-642-20051-9
Francisco Alonso Solís-Marín
Instituto de Ciencias del Mar, y Limnologia
Universidad Nacional Autónoma de México
México City
Mexico
ISBN 978-3-642-20051-9
(eBook)
Springer Heidelberg New York Dordrecht London
Library of Congress Control Number: 2012941234
Ó Springer-Verlag Berlin Heidelberg 2013
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We would like to dedicate this book to the
pioneers of echinoderms research in Latin
American. To Marı´a Elena Caso, Irene
Bernasconi, Luis Roberto Tomassi and
Evelyn Zoppi de Roa.
Foreword
This is a remarkable book. First and foremost, it compiles an immense amount of
literature on all aspects of echinoderm biology and ecology over an extremely
broad geographical range. This will be extremely useful because much of the
literature is relatively unknown to the world at large. The description of the
oceanography and marine environments of each country in a comprehensive way
will be of considerable interest to marine biologists in general.
One of the most interesting aspects of this book is the section devoted to the
history of echinoderm studies in each country. The first reports about echinoderms
were isolated and sporadic, not systematic studies. These generally did not begin
until the 1850s with various foreign expeditions along the coasts of Latin America.
Studies of echinoderms by workers in Latin American countries themselves did
not begin to develop appreciably until the 1950s. This has changed in a dramatic
way. In the latter part of the twentieth century, interest in marine biology and
recognition of its importance in terms of basic science, the environment and
conservation, and economics increased greatly. This led to an increase in the
number of marine biologists in general and of echinoderm biologists in particular.
The amount of research activity that has developed since then is impressive.
Another thing that impressed me, apparent in the book, is the passion of Latin
American biologists for echinoderms, their commitment to conservation, their
desire to assist their countries through their efforts, and their collegiality.
Florida, USA, March 2012
John M. Lawrence
Department of Integrative Biology,
University of South Florida,
Tampa
vii
Foreword
Juan José Alvarado is one of those students who, from very early on, denoted a
clear interest in Marine Science and at some point defined his area of interest.
When he started looking at the echinoderms of Costa Rica and later, at those
reported and collected from Coco Island National Park, he found what he was
looking for. Since then, he has been actively carrying out research on echinoderms,
and most important, he has drawn attention to the activities regarding this group
that are taking place in Latin America. First, with the publication of Research on
Echinoderms in Latin America in 2005 (Revista de Biología Tropical, Volume 53,
Supplement 3: 387 p.). Then, with the creation of the Iberoamerican Echinoderm
Network Red Iberoamericana de Equinodermos (RIE) in 2006. And in 2008, with
the publication of another Special Issue on echinoderm research in Latin America:
Revista de Biología Tropical, 56 (Suppl. 3): 360 p. Now, together with Francisco
Solís-Marín from the Universidad Nacional Autónoma de México (UNAM), they
have put together an outstanding compilation of echinoderm research in Iberoamerica. The dedication of this book to the pioneering Latin American echinoderm
researchers is a tribute to their great work and an acknowledgment of the fruit their
early and many times solitary work has borne. It is a great honor for me to write
this Foreword, for I consider their work a significant step for biological research in
the region..
San José, Costa Rica, March 2012
Jorge Cortés
Centro de Investigación en Ciencias
del Mar y Limnología (CIMAR),
Universidad de Costa Rica
ix
Preface
The Iberoamerican Echinoderms Network (Red Iberoamericana de Equinodermos)
(RIE) (http://zicatela.umar.mx/*redequinos/2/) was created in August 2006 at the
12th International Echinoderms Congress in New Hampshire, United States, as an
initiative to strengthen academic connections between Iberoamerican echinoderm
researchers. The RIE seeks to fortify the links of researchers in the different
countries of the region, being a bridge of communication and exchange of ideas. It
seeks to be the platform for discussion and communication of priority projects for
Latin America echinoderms. It seeks to offer the opportunity of collaborative
work, to place the region among the principal research and conservation regions of
the world. Likewise, the RIE intends to support students interested in echinoderm
research. To enrich their formation, it will offer information about researchers who
could help in their projects, courses, seminars, or symposia, as well as providing
literature. The goal of the RIE is to develop workshops, congresses, special
journals supplements and books, and to support the development of the knowledge
of the echinoderms in Latin America.
The first effort of the RIE was the preparation and publication of a Special Issue
of the Revista de Biología Tropical (International Journal of Tropical Biology and
Conservation) (Alvarado, J. J. and Cortés, J. (Editors). 2005. Research on Echinoderms in Latin America, Volume 53, Supplement 3: 387 p.) (http://www.ots.ac.
cr/tropiweb/intpages/suppl/sup53-3.html). This initiative was born during the 11th
International Echinoderms Congress in Münich, Germany, in 2003. This Special
Issue compiled 28 papers from 12 countries by 62 authors, touching topics as diverse
as paleontology in the Uruguayan region and fisheries models of sea cucumbers in
Baja California. The second output of the RIE was a second supplement of the
Revista de Biología Tropical (Alvarado, J.J. and Cortés, J. (Editors). 2008. Research
on Echinoderms in Latin America II, Volume 56, Supplement 3: 360 p.) (http://www.
ots.ac.cr/tropiweb/intpages/suppl/sup56-3.html). This issue compiled 21 papers
from eight countries by 46 authors in Latin America (including an invited paper from
the Canary Islands). Those papers demonstrated the high degree of knowledge of
some countries in topics such as diversity, ecology, aquaculture, and fisheries. It also
identified countries where more efforts are needed.
xi
xii
Preface
The third effort was the establishment of the first Latin American Echinoderm
Congress (Congreso Latinoamericano de Equinodermos-CLE) held in Puerto
Madryn, Argentina, between 13 and 18 November 2011. This congress was
organized by the Centro Nacional Patagónico (CENPAT), the Museo Argentino de
Ciencias Naturales ‘‘Bernardino Rivadavia’’ (MACN), and the Universidad Nacional de la Patagonia San Juan Bosco. A total of 112 people participated in this
event with 76 oral presentations and 54 Poster, covering the following themes:
biogeography, biochemistry, ecology, physiology, paleontology, fisheries and
aquaculture, reproduction and development, and taxonomy, systematics, and
evolution.
This book represents the fourth output and the confirmation of the commitment
of the RIE with the international scientific community. The book represents an
extensive recompilation of information about all the research done in the region
from theses, reports, books, and scientific journals. We were able to identify the
strengths and weaknesses of each country and also allow the international scientific community to be able to access information that was held in reports, theses,
and local papers that were written in Spanish, allowing a greater diffusion of the
results and conclusions obtained. Our goal is to improve the echinoderm research
on the region and with this road map establish future research in collaboration with
the different laboratories and researchers that participated in this book.
Juan José Alvarado
Francisco Solís-Marín
Acknowledgments
First of all, we thank each of the authors of the chapters that have made this
compilation about Echinoderm Research and Diversity in Latin America possible,
to share their knowledge and passion for such a star group. Muchas gracias!
We also would like to acknowledge the help and support of all the reviewers who
with their comments and suggestions improved this book: Chris Pomory, Cynthia
Lara de Castro, David Pawson, Harilaos Lessios, John Pearse, Manuel Rey Méndez,
Marc Eléaume, Martin Brogger, Nieves Elvira, Pablo González, Philip Lambert,
Rosa del Valle, Tim O’hara, Tom Hopkins. We wish to grant special
credit to John Lawrence and Jorge Cortés who with all the patience, interest,
and care reviewed each of the chapters, leaving their wisdom on every page.
We wish to grant special credit to all the students from the Laboratorio de
Sistemática y Ecología de Equinodermos from the Instituto de Ciencias del Mar y
Limnología, Universidad Nacional Autónoma de Mexico, for their help with the
final elaboration of the appendix: Mauricio Valdés De Anda, Julio Adrián Arriaga
Ochoa, Andrea Alejandra Caballero Ochoa, Tania Pineda Enríquez, Carolina
Martín Cao-Romero, Yoalli Quetzalli Hernández Díaz, Alejandra Martínez Melo,
Viridiana Tapia Ramírez, Guadalupe Bribiesca Contreras, Pedro Josué Garcés
Solchaga, Nancy Escandón Flores, Lucia Alejandra Hernández Herrejón and
Carlos García Linares.
We appreciate the help of Monica Chavez in preparing all the figures in this
book and Cindy Fernandez who always believed in this project and was a keystone
for the final elaboration. JJ Alvarado is grateful to CONACYT (México) and
CONICIT-MICIT (Costa Rica). Finally, we thank Marion Schnider and
SPRINGER for their interest in this book.
xiii
Contents
1
Echinoderm Research and Diversity in Latin America. . . . . . . . .
Juan José Alvarado and Francisco Alonso Solís-Marín
2
The Echinoderms of Mexico: Biodiversity, Distribution
and Current State of Knowledge. . . . . . . . . . . . . . . . . . . . . . . . .
Francisco A. Solís-Marín, Magali B. I. Honey-Escandón,
M. D. Herrero-Perezrul, Francisco Benitez-Villalobos,
Julia P. Díaz-Martínez, Blanca E. Buitrón-Sánchez,
Julio S. Palleiro-Nayar and Alicia Durán-González
3
4
1
11
Central America Echinoderms: Diversity, Ecology
and Future Perspectives . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Juan José Alvarado, Enrique Barraza and Tito Isaac Sancho-Mejías
67
Echinoderm Diversity in Panama: 144 Years of Research
Across the Isthmus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Simon E. Coppard and Juan José Alvarado
107
5
Echinoderms of Colombia. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Milena Benavides-Serrato, Giomar Helena Borrero-Pérez,
Jaime R. Cantera K, Mishal Cohen-Rengifo and Raúl Neira
145
6
Echinoderms of Ecuador . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Jorge Sonnenholzner, Margarita Brandt, Vanessa Francisco,
Alex Hearn, María Luzuriaga, Paulina Guarderas
and Juan Carlos Navarro
183
xv
xvi
7
Contents
Echinoderms from Venezuela: Scientific Recount,
Diversity and Distribution . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
César Lodeiros, Alberto Martín, Vanessa Francisco, Nicida Noriega,
Yusbelly Díaz, Jeny Reyes, Orangel Aguilera and José Alió
8
Echinoderms of Peru . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Yuri Hooker, Elba Prieto-Rios and Francisco A. Solís-Marín
9
Echinoderm from Brazil: Historical Research and the Current
State of Biodiversity Knowledge . . . . . . . . . . . . . . . . . . . . . . . . .
Carlos R. R. Ventura, Michela Borges, Lúcia S. Campos,
Leticia V. Costa-Lotufo, Carolina A. Freire, Valéria F. Hadel,
Cynthia. L. C. Manso, José R. M. C. Silva, Yara Tavares
and Cláudio G. Tiago
10
Echinoderm Research in Uruguay . . . . . . . . . . . . . . . . . . . . . . .
Sergio Martínez
11
Echinoderms from Argentina: Biodiversity, Distribution
and Current State of Knowledge . . . . . . . . . . . . . . . . . . . . . . . .
Martín I. Brogger, Damián G. Gil, Tamara Rubilar,
Mariano I. Martinez, María Enriqueta Díaz de Vivar,
Mariana Escolar, Lucía Epherra, Analía F. Pérez
and Alejandro Tablado
235
277
301
345
359
12
Echinoderms of the Cuban Archipelago. . . . . . . . . . . . . . . . . . .
Rosa del Valle-García, Mercedes Abreu-Pérez, Francisco
Alonso Solís-Marín and Alfredo Laguarda-Figueras
403
13
Recent Echinoderms from Hispaniola . . . . . . . . . . . . . . . . . . . .
Alejandro Herrera-Moreno and Liliana Betancourt
425
14
Review of Echinoderm Research in Puerto Rico,
with the Focus on Biological and Ecological Aspects . . . . . . . . . 437
Stacey M. Williams, Milena Benavides-Serrato, Jose E. García-Arrarás,
Edwin A. Hernández-Delgado and Ruber Rodriguez Barreras
15
Echinoderms of the Canary Islands, Spain. . . . . . . . . . . . . . . . .
José Carlos Hernández, Sabrina Clemente, Fernando Tuya,
Angel Pérez-Ruzafa, Carlos Sangil, Leopoldo Moro-Abad
and Juan José Bacallado-Aránega
471
Contents
16
xvii
Latin America Echinoderm Biodiversity and Biogeography:
Patterns and Affinities. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 511
A. Pérez-Ruzafa, J. J. Alvarado, F. A. Solís-Marín, J. C. Hernández,
A. Morata, C. Marcos, M. Abreu-Pérez, O. Aguilera, J. Alió,
J. J. Bacallado-Aránega, E. Barraza, M. Benavides-Serrato,
F. Benítez-Villalobos, L. Betancourt-Fernández, M. Borges,
M. Brandt, M. I. Brogger, G. H. Borrero-Pérez,
B. E. Buitrón-Sánchez, L. S. Campos, J. R. Cantera, S. Clemente,
M. Cohen-Renfijo, S. E. Coppard, L. V. Costa-Lotufo,
R. del Valle-García, M. E. Díaz de Vivar,
J. P. Díaz-Martínez, Y. Díaz, A. Durán-González, L. Epherra,
M. Escolar, V. Francisco, C. A. Freire, J. E. García-Arrarás,
D. G. Gil, P. Guarderas, V. F. Hadel, A. Hearn,
E. A. Hernández-Delgado, A. Herrera-Moreno, M. D Herrero-Pérezrul,
Y. Hooker, M. B. I. Honey-Escandón, C. Lodeiros, M. Luzuriaga,
C. L. C. Manso, A. Martín, M. I. Martínez, S. Martínez,
L. Moro-Abad, E. Mutschke, J. C. Navarro, R. Neira, N. Noriega,
J. S. Palleiro-Nayar, A. F. Pérez, E. Prieto-Ríos, J. Reyes,
R. Rodríguez-Barreras, T. Rubilar, T. I. Sancho-Mejia, C. Sangil,
J. R. M. C. Silva, J. I. Sonnenholzner, C. R. R. Ventura, A. Tablado,
Y. Tavares, C. G. Tiago, F. Tuya and S. M. Williams
Appendix . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
543
Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
655
Chapter 13
Recent Echinoderms from Hispaniola
Alejandro Herrera-Moreno and Liliana Betancourt
13.1 Introduction
The Island of Hispaniola is the second largest in the Archipelago of the Greater
Antilles. It is located between 17°360 1500 N and 19°570 0900 N, and 68°190 2200 W and
74°410 3300 W. Its geographical boundaries are the Atlantic Ocean to the north, the
Caribbean Sea to the south, the Mona Channel to the East, which separates it from
Puerto Rico, and in the West, the Windward Passage and the Jamaica Channel,
which separate it from Cuba and Jamaica, respectively. The 76,480 km2 of its
territory is divided politically into two countries: Dominican Republic to the east
and the Republic of Haiti in the west, separated by a land border of 360 km.
The Republic of Haiti has a total area of 27,750 km2, which includes the
mainland territory plus several islands and islets such as Gonave (743 km2),
Tortue (180 km2), Vache (52 km2), Cayemites (45 km2) and Navassa (5.2 km2).
The Dominican Republic has an area of 48,442 km2 of mainland in addition to
several islands and islets such as Saona (117 km2), Beata (27 km2) and Catalina
(9.6 km2). Two oceanic banks: Christmas and Silver Banks, with areas of 70 and
150 km2 respectively, are located to the north of Samana in the Atlantic Ocean.
The coastline of Hispaniola extends for about 3,059 km (1,288 km belong to 16
Dominican coastal provinces and 1,771 km to nine Haitian coastal provinces)
containing ecosystems such as sandy beaches, rocky shores, cliffs, estuaries,
coastal lagoons and mangrove forests. On the Haitian coast important mangroves
areas are reported along the north and northeast coast in the Bays of Fort Liberté,
l’Acul, Caracol, the Artibonite estuary and the Islands Les Cayes, Vache, Gonave
and Cayemite (Ehrlich et al. 1987). Along the Dominican coast the largest areas of
mangroves are found in the Bay of Samaná and Montecristi.
A. Herrera-Moreno (&) L. Betancourt
Programa EcoMar, Inc., Santo Domingo, Dominican Republic
e-mail: ongprogramaecomar@yahoo.com
J. J. Alvarado and F. A. Solís-Marín (eds.), Echinoderm Research and Diversity
in Latin America, DOI: 10.1007/978-3-642-20051-9_13,
Ó Springer-Verlag Berlin Heidelberg 2013
425
426
A. Herrera-Moreno and L. Betancourt
The Dominican shelf has a surface area of 8,130 km2 extending down to 200 m
depth, with an average width of 7.5 km. The platform area of Haiti is estimated at
6,683 km2 and is on average narrower. Important coastal habitats include sandy or
muddy bottoms (related to the degree of terrigenous influence), commonly populated with abundant macrophytobenthos, especially seagrasses, followed by coral
patches and fringing or barrier reefs. In Haiti coral reefs occur near Vache Island,
all around Gonave Island in the central Bay of Port-au-Prince, on the Rochelois
Bank, at Cayemites Islands, off the northern coast of the southern peninsula, and
from the border with the Dominican Republic in the east to Acul Bay just west of
Cape Haitien (Creary et al. 2008). Well preserved reefs are found at Navassa
Island (Causey et al. 2000).
In the Dominican Republic important reef areas include the Montecristi
National Park barrier reef in the north-west (where the shelf is widest), narrow
high-energy reefs in the central region, and the Bávaro-Macao-Punta Cana barrier
reef system at the eastern end. Samaná Bay receives many rivers and is the largest
estuary of the insular Caribbean. Coastal reefs in the vicinity are poorly developed,
but the Christmas and Silver Banks reef systems are about 100 km to the north. To
the south, on the Caribbean coast, are the well-studied reefs of National East Park
and the adjacent Saona Island. Uplifted carbonate terraces with reefs growing on
narrow platforms are present in the west from Catalina Island to beyond Santo
Domingo (e.g. Boca Chica and the Submarine National Park La Caleta). Conditions are not good for reefs in the south-west, except on the shallow sheltered shelf
east of Cabo Beata at National Park Jaragua (Woodley et al. 2000).
13.2 Research
Echinoderm studies of the waters around Hispaniola Island have a long history.
The Haitian collection of David Friedrich Weinland from 1857 (NMNH 2002) and
the work on holothuroids by Emil Selenka (1867), with the record of Actinopyga
agassizi (Selenka, 1867) in Haiti and the description of the type specimen of
Holothuria grisea Selenka, 1867, are among the earliest contributions.
William More Gabb made collections in the Dominican Republic in 1878 and
deposited several specimens of Echinometra lucunter (Linnaeus, 1758) in the U.S.
National Museum of Natural History (Rathbun, 1886). In December 1878, during
the Blake Expedition, the echinoid Salenocidaris varispina (A. Agassiz, 1880) was
collected northwest of Haiti at 2,195 m depth (A. Agassiz, 1880). Dodërlein and
Hartmeyer (1910) and Verrill (1915) summarized previous records of asteroids
from Hispaniola. Hubert L. Clark (1919) provided the first taxonomic summary of
Haitian echinoderms listing more than 20 species of all classes, except crinoids.
In the 1930s, new contributions to the knowledge of echinoderms from
Hispaniola include Deichmann (1930) who summarized previous records for the
Dominican Republic and Haiti in her review of the holothurians of the western part
of the Atlantic Ocean. The handbook of the littoral echinoderms of Porto Rico and
13
Recent Echinoderms from Hispaniola
427
the other West Indian Islands (H.L. Clark 1933) also summarized historical
information about all classes of echinoderms for Hispaniola, except crinoids, with
some new records. Austin H. Clark (1939) listed 19 species of shallow-water
echinoderms (between 1 and 2 m depth) collected in two localities in the north of
Haiti (Cape Haitien and Tortue Island) during the Smithsonian-Hartford expedition of March 1937, aboard the RV Joseph Conrad.
New collections took place during the 1930s on five expeditions: ParishSmithsonian in 1930 to Haiti on the yacht Esperanza, Norcross-Bartlett in July 1931
to the southwest of Santo Domingo down to 200 m depth on the schooner Effie
Morrissey, Johnson–Smithsonian Deep-Sea in February 1933 on the Yacht Caroline
at depths from 26 to 40 m at twelve stations inside the Bay and around the Peninsula
of Samaná (Bartsch 1933), RV Stranger in February 1933 in Port-au-Prince and the
RV Atlantis in April 1939 dredging to 1,170 m deep at one station to the northeast of
Haiti (H.L. Clark 1941). These expeditions contributed new records for Hispaniola
including two deep-water asteroid species: Persephonaster patagiatus (Sladen,
1889) and Phormosoma placenta sigsbei (A. Agassiz, 1880). The expeditions of
1931 and 1933 collected the first specimens of crinoids: Poliometra prolixa (Sladen,
1881) and Nemaster rubiginosa (Pourtalés, 1869).
In the decade of 1960s two expeditions took place. The exploratory fishing
vessel Silver Bay in October 1963 made collections at six stations between 92 and
348 m deep in the north, northeast and east of the Dominican Republic. The
exploratory fishing vessel Oregon in June and May, 1965 and December 1969
collected between 11 and 59 m deep at four stations off the north Haitian coast and
the northeast Dominican coast. During this period, Parslow and Clark (1963)
summarized the zoogeographical distribution of shallow-water (less than 17 m
deep) ophiuroids known for the West Indies, including 11 species from Hispaniola,
updating the previous list of H.L. Clark (1919). Deichmann (1963) summarized
previous records of holothuroids for Haiti. Halpern (1969) mentioned the asteorid
Litonotaster intermedius (Perrier, 1884) found in the Windward Pass, between
Cuba and Hispaniola, at a depth between 1,958 and 3,294 m (Downey 1973).
From January to July 1970 and July 1971 the most prolific expeditions carried
out in Hispaniolan waters took place aboard the RV John Elliot Pillsbury.
Collections were made at seven stations in Haiti, between 31 and 2,545 m deep,
and 33 stations in Dominican Republic, between 9 and 3,109 m deep (Fig. 13.1)
(Staiger and Voss 1970). Meyer et al. (1978) included the RV John Elliot Pillsbury
collections in their zoogeographical study of western Atlantic Ocean crinoids,
offering the most complete summary of this group, with 19 species listed for
Dominican and Haitian waters. In her review of the Order Brisingida for
the Atlantic Ocean, Downey (1986) also incorporated Haitian specimens from the
Pillsbury expeditions. A.M. Clark (1987) described the type specimen of the
asteoridHenricia downeyae A.M. Clark, 1987 from a Haitian locality, registered in
the catalogue of types of the NMNH (Ahearn 1995). Two more records for Haiti
from this period were the holothuroid Psolus tuberculosus Théel, 1886 (Miller and
Pawson 1984) and the asteroid Ceramaster grenadensis grenadensis (Perrier,
1881) (Halpern 1970).
428
A. Herrera-Moreno and L. Betancourt
Fig. 13.1 Localities where collections of echinoderms have been made by several expeditions
and national and international projects on the shelf and oceanic waters of Hispaniola
All these expeditions increased the collections of the five classes of echinoderms from Hispaniola in international museums with 541 specimens of 132
species located in the U.S. Museum of Natural History (NMNH 2002), Florida
Museum of Natural History (FLMNH 2010), Museum of Comparative Zoology at
Harvard University (MCZ 2010) and Museum of Natural Sciences of Berlin (ZMB
2010). The RV John Elliot Pillsbury expeditions had a decisive role in the
enrichment of collections, particularly with deep-sea species, that at the end of the
1970s, 90 % of the crinoid, asteroid, and echinoid species known from Hispaniola
were already described and deposited in different museums.
In the Dominican Republic, the creation of the Center of Marine Biology
Research (CIBIMA) in 1962 promoted coastal and marine biodiversity research.
Between the decades from 1970 to 1990, CIBIMA produced summarized list of
echinoids and asteroids (Cicero et al. 1976), ophiuroids (Rathe 1978) and holothuroids (Briones 1983). Multiple reports were finally compiled into a preliminary
study on coastal and marine biodiversity of the Dominican Republic (CIBIMA
1992). From these investigations, 242 specimens of 58 species are deposited in the
National Museum of Natural History of Santo Domingo (MNHNSD 2008).
In May 1979, the Autonomous University of Santo Domingo, along with the
University of Puerto Rico, implemented an expedition on board the RV Crawford
to make inventories of the southeast Dominican reefs of La Caleta and Catalina
and Saona Islands (Williams et al. 1983). Later the IDECOOP expedition
13
Recent Echinoderms from Hispaniola
429
(Fig. 13.1) made collections between 142 and 270 m deep with three new records
for the deep-sea Dominican echinoderms: the crinoid Cenocrinus asterius (Linnaeus, 1775); the ophiuroid Astronicda isidis Lyman, 1872 and the echinoid
Conolampas sigsbei (A. Agassiz, 1878) (Rivas 1983). Studies carried out by
researchers from East Carolina University in Montecristi reefs (Luczkovich 1991)
extended the reports of Dominican crinoids with the record of Tropiometra carinata (Lamarck, 1816).
The information on Haitian echinoderms comes from old collections, that have
already been mentioned, and some more recent studies carried out by foreign
institutions. In June 1988, a detailed study at 13 stations on different ecological
zones (from the reef lagoon to the frontal reef) of the Arcadines coral reefs was
made by the World Wildlife Fund and the Conservation Foundation Wilcox
Associates, between 0.3 and 21 m depth (Wilcox et al. 1989). Twelve species of
common shallow-water echinoderms were reported. Hendler et al. (1995) summarized the echinoderms from approximately 17 old Haitian records.
With the implementation of the Hispabiota Marina Project (Herrera-Moreno
and Betancourt 2012) by Programa EcoMar, Inc. in the Dominican Republic a
bibliographical and taxonomic review of echinoderm species was done for the first
time, with a historical and insular approach (Herrera-Moreno and Betancourt
2004). A database was created with the results of collections of echinoderms from
more than 150 localities of the coastal zone, the shelf and the deep zone of
Hispaniola (Fig. 13.1). This project added 60 species to the national inventory of
CIBIMA (1992) confirming 123 species for Dominican coastal and marine zone. It
also compiled for the first time 79 species for Haiti. A total of 156 species for
Hispaniola (Appendix) is discussed in the present report.
13.3 Diversity and Distribution
The numbers of echinoderms in different taxonomic categories for the Dominican
Republic, Haiti and Hispaniola are summarized in Table 13.1. There are 22 species
of crinoids (18 for the Dominican Republic and eight for Haiti), subdivided into 17
genera, nine families and four orders. About five species of crinoids are found in
coral reef environments at depths to 45 m deep including those which, according to
Hendler et al. (1995), are more common and accessible to conventional scuba
diving. The remaining species are distributed below 100 to 1,033 m deep.
The list of asteroids has 33 known species (18 for Haiti and 21 for the
Dominican Republic) subdivided into 24 genera, 11 families and seven orders.
Downey (1973) in her review of the asteroids in the Gulf of Mexico and the
Caribbean described 95 species and noted that the asteroid fauna from Hispaniola
was practically unknown. The bathymetric ranges of the asteroid species known
from Hispaniola varies from 0.3 to 3,493 m deep. About eight species are distributed from the surface to 50 m deep primarily from the mangroves to the reef
front, with the remaining species found below 50 m deep.
430
A. Herrera-Moreno and L. Betancourt
Table 13.1 Summary of the numbers of different taxonomic categories for echinofauna groups
of Dominican Republic (DO), Haiti (HA) and Hispaniola (HI)
Orders
Families
Genus
Species
Crinoidea
Asteroidea
Ophiuroidea
Echinoidea
Holothuroidea
TOTAL
HA
DO
HI
HA
DO
HI
HA
DO
HI
HA
DO
HI
HA
DO
HI
HA
DO
HI
3
4
4
7
4
7
2
2
2
7
10
12
3
3
3
22
23
28
4
9
9
11
7
11
9
11
12
12
16
17
6
5
7
42
48
57
7
14
17
16
14
24
12
15
20
15
31
36
7
7
10
57
81
107
8
18
22
18
22
33
21
24
30
21
41
50
11
18
21
79
123
156
Thirty species of ophiuroids are known for Hispaniola (21 for Haiti and 24 for
the Dominican Republic), subdivided into 20 genera, 12 families and two orders.
Most species of ophiuroids are distributed in water less than 30 m deep found in
seagrass beds and coral reefs. Only four species were collected between 148 and
366 m deep.
Fifty species of echinoids are listed (21 for Haiti and 41 for the Dominican
Republic) subdivided into 36 genera (including one species identified only to the
genus level), 17 families and 12 orders. Although bathymetric data are not
available for all species, the collections from Hispaniola contain the more common
shallow-water species from mangrove habitats, seagrass beds and coral reefs. At
least about 30 species are distributed from 100 to 2,545 m depth.
The list of holothurians of Hispaniola includes 21 species (11 for Haiti and 18
for the Dominican Republic) with ten genera, seven families and three orders.
Most species of holothuroid are common in shallow sedimentary environments.
Three species are found between 243 and 1,400 m deep.
13.4 Ecology
In the Dominican Republic, echinoderm species are mentioned in the inventories
of invertebrates of seagrass beds and coral reefs from Manzanillo to Punta Rucia in
Montecristi (Luczkovich 1991; CIBIMA 1998), Puerto Plata (Herrera-Moreno and
13
Recent Echinoderms from Hispaniola
431
Betancourt 2009), Samaná (Sang and Lysenko 1994; Sang 1996), Punta Cana,
Bávaro (CURPOB 2000; Brandt et al. 2003), National East Park (Vega et al. 1997)
and Isla Saona in La Altagracia, Catalinita in La Romana, La Caleta in Santo
Domingo (Williams et al. 1983), Haina in San Cristóbal (Herrera-Moreno et al.
2009), Las Calderas Bay in Peravia (Almonte 1976), Puerto Viejo in Azua
(González et al. 1978) and Jaragua National Park in Pedernales (Weil 2006).
In Haiti, inventories with echinoderm species include Lamentin (Beebe 1928), Les
Arcadins reefs (FoProBIM 1985; Wilcox et al. 1989) and Navassa (Miller 2003).
Due to the dramatic reduction of black sea urchin Diadema antillarum in the
Caribbean in the 1980s, the presence of this species is always highlighted in all
inventories and some abundance surveys have been done. In the Dominican reefs,
Chiappone (2001) estimated an average density of D. antillarum of 0.03 ind
50 m-2 for the coral reefs in the National East Park, between 9 and 17 m depth.
He also estimated 9.15 ind 50 m-2 in the Boca Chica reefs between 4 and 20 m
depth. Brandt et al. (2003) reported a maximum density of 5.4 ind 10 m-2 in the
reefs of Punta Cana and Bavaro with the highest average values in the deep forereef (1.2 ind 10 m-2) in relation to the shallow areas of the back-reef (0.5 individuals/10 m2). For Caracol Bay in Haiti, Hay (1984) reported 20 ind m-2,
between 3 and 10 m depth. FoProBIM (FoProBIM 1985) found densities of D.
antillarum up to 14.5 ind 100 m-2 at 5 m depth in the area of Les Arcadines. More
recently, abundance values from 0 to 1.5 ind 10 m-2 have been reported in the
same area by Linton (2003).
However, ecological studies at the level of echinoderm populations or communities seem to be scarce. In the Dominican Republic, Chiappone (2001) conducted a study measuring density and size distribution for five species of echinoids
(D. antillarum, Echinometra viridis, Echinometra lucunter, Eucidaris tribuloides
and Tripneustes ventricosus) in the National East Park and Boca Chica. Tewfik
et al. (2005) studied the impact of anthropogenic enrichment on the seagrass food
web at two locations in southern Dominican Republic (Pedernales and Barahona),
involving several species of echinoderms, categorized by them as three consumer
functional groups: (a) generalists (regular urchins: Lytechinus variegatus, T.
ventricosus, D. antillarium); (b) subsurface deposit feeders (red heart urchin
Meoma ventricosa and sand dollars) and (c) surface deposit feeders (cushion sea
star Oreaster reticulatus and sea cucumbers). Hay (1984) compared the activity of
grazing fish and sea urchins in the Bay of Caracol in Haiti, on an overexploited
coral reef including density data of Diadema antillarum and its rate of consumption of Thalassia testudinum.
13.5 Aquaculture and Fisheries
Commercial fishing of sea cucumber takes place in the Dominican Republic.
According to the Dominican Council of Fisheries and Aquaculture CODOPESCA
in 2002 there was an export of 57.6 MT, equivalent to 9 % of seafood exports. The
432
A. Herrera-Moreno and L. Betancourt
figures for years 2007, 2008 and 2009 indicate much smaller export volumes with
1.27, 2.37 and 1.37 MT, respectively. The trading company for sea cucumbers in
the Dominican Republic is NETCO C x A which operates from Juan de Bolaño
Beach in Montecristi with CODOPESCA permissions for exportation to the United
States. There are no studies that scientifically support this exploitation. In fact,
there is no information on the species being harvested, minimum sizes, close
seasons or any other descriptive parameters of an organized fishery. The survival
of these echinoderm populations is seriously threatened. According to ToralGranda (2008), between 2002 and 2003, China imported 2,607 kg of dry weight of
sea cucumbers from Dominican Republic and 10,680 kg of dry weight from Haiti.
In Haiti, the Caribbean SeaFood Company announces online the sale of dried sea
cucumbers of the species Stichopus badionotus.
13.6 Threats
The threats to echinoderms are not very different from those faced by other marine
groups of Hispaniola. The destruction of ecosystems (mainly mangroves, seagrass
beds and coral reefs) due to coastal development (human settlements, agriculture,
industrial and tourist facilities and/or fishing) and tourism activities (especially
diving) is the major threat that affects all groups. In addition to impacts of habitat
destruction, some groups of echinoderms are exploited for commercial purposes.
These include some species of sea cucumbers used for human consumption and
some species of starfish and sea urchins used for sale as aquarium species
(SERCM 2004). Other echinoderm species, such as the sand dollar Mellita quinquiesperforata and the starfish Oreaster reticulatus are collected for handicrafts
that are sold in tourist establishments. Some fishermen also use this asteroid
species as bait in net traps. No ecological studies or fisheries biology research have
been done to support a sustainable exploitation of these resources.
13.7 Recommendations
It is essential to train a group of national specialists from Haiti and the Dominican
Republic in the systematic of all echinoderm classes that can work closely together
to better understand the echinofauna of Hispaniola. The National Museum of
Natural History in Santo Domingo should be involved in the development and
review of the marine collections. New investigations in structural ecology and
population dynamics of echinoderms must be initiated for those species that may
be subject to some kind of extractive use. For exploited species, it is also necessary
to conduct fishery biology investigations for the establishment of closed seasons,
protected areas from fishing and legal minimum sizes. Fishing regulations and
fishery statistics are crucial for a sustainable exploitation of these species. Marine
13
Recent Echinoderms from Hispaniola
433
research should continue increasing the knowledge of the echinoderms in new
areas of the Dominican and Haitian platforms, including exploration of deeper
zones.
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Appendix
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LV Costa-Lotufo, R del Valle-García, Y Díaz, ME Díaz de Vivar, JP DíazMartínez, A Durán-González, L Epherra, M Escolar, V Francisco, CA Freire,
JE García-Arrarás, DG Gil, P Guarderas, VF Hadel, A Hearn, JC Hernández,
EA Hernández-Delgado, A Herrera-Moreno, MD Herrero-Pérezrul, Y Hooker,
MBI Honey-Escandón, C Lodeiros, M Luzuriaga, CLC Manso, A Martín,
MI Martinez, S Martínez, L Moro-Abad, E Mutschke, JC Navarro, R Neira,
N Noriega, JS Palleiro-Nayar, AF Pérez, A Pérez-Ruzafa, E Prieto-Rios, J Reyes,
R Rodríguez, T Rubilar, T Sancho-Mejía, C Sangil, JRMC Silva,
JI Sonnenholzner, CR Ventura, A Tablado, Y Tavares, CG Tiago, F Tuya,
SM Williams
J. J. Alvarado and F. A. Solís-Marín (eds.), Echinoderm Research and Diversity
in Latin America, DOI: 10.1007/978-3-642-20051-9,
Ó Springer-Verlag Berlin Heidelberg 2013
543
Depth (m) Habitat MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
Class Crinoidea
Order Comatulida
Superfamily Antedonacea
Family Antedonidae
Antedon bifida (Pennant, 1777)
Anthometra adriani (Bell, 1908)
Eumorphometra fraseri John, 1938
Fariometra parvula (Hartlaub,
1895)
Florometra magellanica (Bell,
1882)
Florometra mawsoni A. H. Clark,
1913
Florometra parvula (Hartlaub,
1895)
Florometra serratissima (A.
H. Clark, 1907)
Florometra tanneri (Hartlaub, 1895)
Isometra hordea John, 1938
Isometra graminea John, 1938
Isometra vivipara Mortensen, 1917
Promachocrinus kerguelensis
Carpenter, 1880
Solanometra antarctica (Carpenter,
1888)
Family Zenometridae
Psathyrometra bigradata (Hartlaub,
1895)
x
x
x
70–1674
410–687
rb
20–1017
x
x
rb,sb
x
x
38–770
x
x
589–1969
rb
12–3234
rb
x
x
104–207
17–490
rb
x
x
x
79–845
20–2100
x
x
x
x
287–1759
x
741–823
544
Table A.1 Taxonomic list of the Echinoderms of the Pacific coast of Latin America
x
x
Appendix
(continued)
Depth (m) Habitat
Superfamily Notocrinacea
Family Notocrinidae
Notocrinus virilis Mortensen, 1917
Notocrinus mortenseni John, 1938
Superfamily Tropiometracea
Family Thalassometridae
Thalassometra agassizii (Hartlaub,
1895)
Order Hyocrinida
Family Hyocrinidae
Calamocrinus diomedae A.
Agassiz, 1890
Hyocrinus foelli (Roux & Pawson,
1999)
Ptilocrinus antarcticus (Bather,
1908)
Class Asteroidea
Order Paxillosida
Family Astropectinidae
Astropecten armatus Gray, 1840
Astropecten armatus erinaceus
Gray, 1840
Astropecten benthophilus Ludwig,
1905
Astropecten brasiliensis
peruvianus Verrill, 1870
Astropecten exiguus Ludwig 1905
Astropecten fragilis Verrill, 1870
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
Appendix
Table A.1 (continued)
x
x
596–1429
rb
x
717–1431
3030
rb
450–500
rb
0–160
11–60
mb,rb,sb x
rb,sb
1408
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
6–99
mb,sb
232–2136
1–5
sb
x
x
x
x
x
x
x
(continued)
545
Depth (m) Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–204
95–121
mb,rb,sb x
2–488
mb,rb,sb x
278–286
rb,sb
x
x
80–500
sb
377–971
mb,sb
1573–4335 mb,sb
732–1593
10–2424
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
mb,rb
x
20–3248
sb
1866
1485–1618
x
x
x
x
23–300
rb
x
55–1503
sb
x
50–1050
mb
10–1946
mb
10–12
mb,sb
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Astropecten regalis Gray, 1840
Astropecten sulcatus Ludwig,
1905
Astropecten verrilli de Loriol,
1899
Astropecten ornatissimus Fisher,
1906
Bathybiaster loripes Sladen, 1889
Dipsacaster eximius Fisher, 1905
Dytaster gilberti Fisher, 1905
Leptychaster inermis (Ludwig,
1905)
Mimastrella cognata (Sladen,
1889)
Psilaster charcoti (Koehler, 1906)
Psilaster pectinatus (Fisher, 1905)
Psilaster sladeni Ludwig, 1905
Persephonaster armiger Ludwig,
1905
Tethyaster canaliculatus (A.
H. Clark, 1916)
Thrissacanthias penicillatus
(Fisher, 1905)
Family Goniopectinidae
Ctenodiscus procurator Sladen,
1889
Ctenodiscus crispatus (Retzius,
1805)
Luidia (Platasterias) latiradiata
(Gray, 1871)
546
Table A.1 (continued)
Depth (m) Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
33–126
sb
x
4–55
0–220
280
3–55
1.5–12
0–40
rb,sb
mb,sb
mb
mb,sb
mb,sb
rb,sb
x
x
1–386
sb
x
x
110
3–250
18
mb,sb,rub x
sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
1570–6330 rb
x
x
1463–5780 mb,sb
x
x
x
5204–7200 mb
1158–6035 mb
x
x
x
x
x
x
x
x
x
4240–4337 mb
x
2600–5303 mb
117–400
x
x
x
x
x
x
mb,rb,sb
1180–3575 sb,rub,mb x
x
x
x
x
547
Luidia armata Ludwig, 1905
Luidia asthenosoma Fisher, 1906
Luidia bellonae Lütken, 1865
Luidia columbia (Gray, 1840)
Luidia ferruginea Ludwig, 1905
Luidia foliolata (Grube, 1866)
Luidia latiradiata (Gray, 1871)
Luidia magellanica Leipoldt,
1895
Luidia phragma H. L. Clark,
1910
Luidia porteri A. H. Clark, 1917
Luidia superba A. H. Clark, 1917
Luidia tessellata Lütken, 1859
Family Porcellanasteridae
Eremicaster crassus (Sladen,
1883)
Eremicaster pacificus (Ludwig,
1905)
Eremicaster vicinus Ludwig,
1907
Porcellanaster ceruleus WyvilleThomson, 1877
Styracaster paucispinus Ludwig,
1907
Thoracaster cylindratus Sladen,
1883
Family Pseudarchasteridae
Pseudarchaster discus Sladen,
1889
Pseudarchaster pectinifer
Ludwig, 1905
Appendix
Table A.1 (continued)
(continued)
Depth (m) Habitat MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
700–1620
sb,rub
x
98–800
sb,rub
x
x
999
x
1800–1936 rb
x
3058
x
1485–2323 rb
x
x
1618–2323
x
x
x
0–732
sb
x
350–500
sb
x
488–512
mb
x
466–1903
mb,sb
x
790–2323
mb,rb
x
0–30
cr,rb
x
x
0–300
213
2–40
mb,rb
rb,sb
rb,kf
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Pseudarchaster pulcher Ludwig,
1905
Pseudarchaster pusillus Fisher,
1905
Pseudarchaster verrilli Ludwig,
1905
Order Notomyotida
Family Benthopectinidae
Benthopecten acanthonotus Fisher,
1905
Benthopecten cognatus (Ludwig,
1905)
Benthopecten pectinifer (Ludwig,
1905)
Benthopecten spinuliger (Ludwig,
1905)
Calyptraster tenuissimus
Bernasconi, 1966
Cheiraster (Luidiaster) planeta
(Sladen, 1889)
Cheiraster (Luidiaster) californicus
Ziesenhenne, 1942
Nearchaster aciculosus (Fisher,
1910)
Pectinaster agassizi Ludwig, 1905
Order Valvatida
Family Acanthasteridae
Acanthaster planci (Linnaeus, 1758)
Family Asterinidae
Asterina fimbriata Perrier, 1875
Meridiastra modesta (Verrill, 1870)
Patiria chilensis (Lütken, 1859)
548
Table A.1 (continued)
Depth (m) Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–300
0–20
x
cr,rb
rb
x
x
0–3
x
x
0–128
cr,mb,rb,sb x
33–121
rb
x
3–60
cr,rb
x
2–260
rb
x
0–270
rb,sb
0–145
0–135
rb,sb
rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
75–898
x
x
10–898
rb,mb,sb
x
x
382–1248
mb,rb
x
x
188–1244
sb
0–400
rb
x
225–1148
rb
x
x
x
x
(continued)
549
Patiria miniata Verrill, 1913
Patiriella calcarata (Perrier,
1875)
Patiriella exigua (Lamarck,
1816)
Family Asterodiscididae
Amphiaster insignis Verrill,
1868
Paulia horrida Gray, 1840
Family Asteropseidae
Asteropsis carinifera (Lamarck,
1816)
Dermasterias imbricata (Grube,
1857)
Family Ganeriidae
Cycethra verrucosa (Philippi,
1857)
Ganeria falklandica Gray, 1847
Ganeria hahni Perrier, 1891
Family Goniasteridae
Ceramaster grenadensis
patagonicus (Sladen, 1889)
Ceramaster patagonicus
(Sladen, 1889)
Ceramaster leptoceramus
(Fisher, 1905)
Cryptopeltaster lepidonotus
Fisher, 1905
Hippasteria phrygiana
(Parelius, 1768)
Hippasteria falklandica Fisher,
1940
Appendix
Table A.1 (continued)
Depth (m) Habitat MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
Hippasteria spinosa Verrill, 1909
Litonotaster tumidus H. L. Clark,
1920
Mediaster elegans Ludwig, 1905
Mediaster transfuga Ludwig, 1905
Mediaster tenellus Fisher, 1905
Nymphaster diomedeae Ludwig,
1905
Pillsburiaster ernesti (Ludwig,
1905)
Family Mithrodiidae
Mithrodia bradleyi Verrill, 1870
Family Odontasteridae
Acodontaster elongatus elongatus
(Sladen, 1889)
Acodontaster elongatus granuliferus
(Koehler, 1912)
Diplodontias singularis (Müller &
Troschel, 1843)
Odontaster meridionalis (E.A.
Smith, 1876)
Odontaster penicillatus (Philippi,
1870)
Family Ophidiasteridae
Leiaster coriaceus Peters, 1852
Leiaster glaber Peters, 1852
Leiaster teres (Verrill, 1871)
Linckia columbiae Gray, 1840
50–512
rb
4066–5196 mb
1789
840–900
580–1192
702–1810
x
x
x
sb
sb
mb,sb
550
Table A.1 (continued)
x
x
x
x
x
x
x
2149
x
x
x
x
x
x
0–14
cr,rb
x
x
x
x
x
x
x
8–400
rb,sb
x
74–841
rb,sb
x
0–84
rb,sb
0–646
rb,sb
x
6–400
rb,sb
x
2–15
1–57
0–100
cr
cr,rb
cr,rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
(continued)
Depth (m) Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
rb
x
x
x
x
56–91
rb
20–59
cr, rb
1–75
rb
x
x
x
x
x
x
x
0–130
cr,rb,sb
x
x
x
0–50
cr,rb,sb
x
x
x
x
x
67–79
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
15–49
rb
0–900
mb,rb,sb
x
5–450
mb,rb,sb
x
48–1094
x
0–183
2–92
cr,rb,rub
cr,rb,sb
x
x
86–4200
15–450
mb,rb,rub x
rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
551
Linckia guildingi Gray, 1840
Linckia multifora (Lamarck,
1816)
Narcissia gracilis A.H. Clark,
1916
Narcissia gracilis malpeloensis
Downey, 1975
Ophidiaster agassizi Perrier,
1881
Ophidiaster ludwigi de Loriol,
1900
Pharia pyramidata (Gray, 1840)
Phataria mionactis Ziesenhenne,
1942
Phataria unifascialis (Gray,
1840)
Tamaria obstipa Ziesenhenne,
1942
Tamaria stria Downey, 1975
Family Poraniidae
Porania (Porania) antarctica
E.A. Smith, 1876
Poraniopsis echinaster Perrier,
1891
Poraniopsis inflatus (Fisher,
1906)
Family Oreasteridae
Nidorellia armata (Gray, 1840)
Pentaceraster cumingi (Gray,
1840)
Family Solasteridae
Lophaster furcilliger Fisher, 1905
Lophaster stellans Sladen, 1889
Appendix
Table A.1 (continued)
Paralophaster antarcticus Koehler,
1912
Solaster regularis Sladen, 1889
Order Velatida
Family Korethrasteridae
Peribolaster folliculatus Sladen,
1889
Family Pterasteridae
Diplopteraster semireticulatus
(Sladen, 1882)
Diplopteraster verrucosus (Sladen,
1882)
Pteraster cf. diaphanus (Ludwig,
1905)
Pteraster gibber (Sladen, 1882)
Pteraster affinis Smith, 1876
Pteraster affinis lebruni Perrier, 1891
Hymenaster cremnodes H. L. Clark,
1920
Hymenaster gracilis Ludwig, 1905
pellucidus Thomson,
platyacanthus Ludwig,
quadrispinosus Fisher,
trias H. L. Clark, 1920
violaceus Ludwig, 1905
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
88–750
sb
x
5–800
rb,sb,mb
x
10–460
rb
x
300–
1500
0–470
rb,sb
x
sb
x
1410
7–460
80–130
74–341
4335
x
rb
rb
sb
mb
2418–
3241
13–3240 mb
2487–
2877
778–
3240
5203
x
x
x
x
x
x
x
x
x
x
x
x
mb
x
x
x
mb
x
x
x
(continued)
Appendix
Hymenaster
1873
Hymenaster
1905
Hymenaster
1905
Hymenaster
Hymenaster
Depth
(m)
552
Table A.1 (continued)
Order Spinulosida
Family Echinasteridae
Echinaster (Echinaster)
parvispinus A. H. Clark,
1916
Echinaster (Othilia)
aculeata (Gray, 1840)
Echinaster (Othilia)
spinulosus Verrill, 1869
Echinaster (Othilia)
tenuispinus Verrill, 1871
Echinaster cribella
Lütken, 1871
Echinaster cylindricus
Meissner, 1892
Echinaster panamensis
Leipoldt, 1895
Henricia aspera Fisher,
1906
Henricia asthenactis
Fisher, 1910
Henricia clarki Fisher,
1910
Henricia gracilis
(Ludwing, 1905)
Henricia leviuscula
(Stimpson, 1857)
Henricia nana (Ludwig,
1905)
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
18–28
rb
x
3–24
sb
1–55
cr,mb,rb
0–18
rb
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
487–570 rb
x
91–1250 rb
x
226–
2001
267–
1244
0–228
x
rb
x
57–200
rb
x
rb
rb
x
x
x
553
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
22–940
rb
666
rb
0–430
rb,sb,rub
0–350
rb
x
0–120
sb
x
x
x
x
x
x
x
x
34–54
x
0–350
rb
x
11–156
cr,rb
0–20
rb
x
rb
x
x
x
x
x
52–84
cr,rb,rub
0–650
mb,rb,sb
x
0–500
mb,rb,sb
x
3334
x
x
x
x
x
x
(continued)
Appendix
Henricia obesa (Sladen,
1889)
Henricia seminudus (A.
H. Clark, 1916)
Henricia studeri Perrier,
1891
Order Forcipulatida
Family Asteriidae
Anasterias antarctica
(Lütken, 1857)
Anasterias pedicellaris
(Köehler, 1923)
Anasterias spirabilis
(Bell, 1881)
Anasterias varium
(Philippi, 1870)
Astrometis sertulifera
(Xantus, 1860)
Astrostole platei
(Meissner, 1896)
Astrostole paschae (H.
L. Clark, 1920)
Coronaster marchenus
Ziesenhenne, 1942
Cosmasterias lurida
(Philippi, 1858)
Diplasterias brandti
(Bell, 1881)
Distolasterias robusta
(Ludwig, 1905)
Depth
(m)
554
Table A.1 (continued)
Depth
(m)
Evasterias troscheli
(Stimpson, 1862)
Leptasterias pusilla
Fisher, 1930
Meyenaster gelatinosus
(Meyen, 1834)
Pisaster brevispinus
(Stimpson, 1857)
Pisaster giganteus
(Stimpson, 1857)
Pisaster ochraceus
(Brandt, 1835)
Rathbunaster
californicus Fisher, 1906
Sclerasterias alexandri
(Ludwig, 1905)
Sclerasterias heteropaes
Fisher, 1924
Family Coscinasteridae
Psalidaster mordax
Fisher, 1940
Family Heliasteridae
Heliaster canopus
Perrier, 1875
Heliaster cumingii
(Gray, 1840)
Heliaster helianthus
(Lamarck, 1816)
Heliaster kubiniji
Xantus, 1860
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
37
rb
0–22
rb
0–102
rb
x
0–374
rb
x
0–8
rb
x
rb
x
x
x
x
rb
80–600
sb
x
x
x
61–384
18–457
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
intertidal rb
x
0–14
rb
x
0–20
rb,sb
x
0–20
rb
x
x
x
x
x
555
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–20
rb
x
0–20
rb
x
rb
5–450
rb
0–520
rb,rub
x
x
x
x
x
x
x
1618–
2418
245
x
x
x
704
x
x
0–455
rb
0–100
rb
0–80
rb,sb,kf
x
x
x
x
(continued)
Appendix
Heliaster microbrachius
Xanthus, 1860
Heliaster polybrachius
H. L. Clark, 1907
Heliaster solaris A.
H. Clark, 1920
Labidiaster radiosus
Lütken, 1871
Family
Pedicellasteridae
Ampheraster
hyperonchus (H.
L. Clark, 1913)
Hydrasterias improvisus
(Ludwig, 1905)
Tarsaster cocosanus
(Ludwig, 1905)
Tarsaster galapagensis
(Ludwig, 1905)
Family Pycnopodiidae
Pycnopodia
helianthoides (Brandt,
1835)
Family Stichasteridae
Allostichaster capensis
(Perrier, 1875)
Stichaster striatus Müller
& Troschel, 1840
Family Zoroasteridae
Depth
(m)
556
Table A.1 (continued)
Cnemidaster nudus
(Ludwig, 1905)
Cnemidaster wyvillii
Sladen, 1889
Myxoderma longispinum
(Ludwig, 1905)
Myxoderma
platyacanthum (H.
L. Clark, 1913)
Myxoderma qawashqari
(Moyana & Larrain,
1976)
Myxoderma sacculatum
(Fisher, 1905)
Zoroaster magnificus
Ludwig, 1905
Zoroaster ophiurus
Fisher, 1905
Order Brisingida
Family Brisingidae
Astrolirus panamensis
(Ludwig, 1905)
Hymenodiscus
monacantha H. L. Clark,
1920
Hymenodiscus tenella
(Ludwig, 1905)
Family Freyellidae
Freyastera benthophila
(Sladen, 1889)
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1366–
2600
mb,rb
x
x
Appendix
Table A.1 (continued)
x
980–
mb,rb
2418
395–650 mb,rb
0–723
sb
1000–
1546
3056–
3667
695–
2226
mb,rb
x
x
x
x
x
x
x
x
mb,sb
x
x
x
x
x
x
x
mb
2418
4064–
4667
x
mb,sb
48–2418
4064
x
x
x
mb
x
557
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
3182–
3279
x
x
x
x
x
x
x
266–
1236
sb
x
152–
2663
266–
1800
717–
3058
cr,mb,sb
x
x
x
x
350–731
1271
x
x
x
x
x
x
x
(continued)
Appendix
Freyella insignis
Ludwig, 1905
Freyella pacifica
Ludwig, 1905
Freyella propinqua
Ludwig, 1905
Freyellaster scalaris (A.
H. Clark, 1916)
Class Ophiuroidea
Order Euryalida
Family Asteronychidae
Asteronyx excavata
Lütken & Mortensen,
1899
Asteronyx loveni Müller
& Troschel, 1842
Asteronyx longifissus
Doederlein, 1927
Astrodia plana (Lütken
& Mortensen, 1899)
Family
Asteroschematidae
Asteroschema rubrum
(Lyman, 1882)
Asteroschema sublaeve
Lütken & Mortensen,
1899
Family
Gorgonocephalidae
Habitat
558
Table A.1 (continued)
Astrocaneum spinosum
(Lyman, 1875)
Astrodendrum
galapagensis A.
H. Clark, 1916
Astrodictyum panamense
(Verrill, 1867)
Astrotoma agassizii
Lyman, 1875
Gorgonocephalus
chilensis (Philippi, 1858)
Gorgonocephalus
diomedeae Lütken &
Mortensen, 1899
Order Ophiurida
Family Ophiomyxidae
Ophiomyxa panamensis
Lütken & Mortensen,
1899
Ophiomyxa vivipara
Studer, 1876
Ophiolycus nutrix
(Mortensen, 1936)
Family Amphiuridae
Amphilepis nuda
Tommasi, 1976
Amphilepis patens
Lyman, 1879
Family Amphiuridae
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
4–183
cr,mb,sb,rb
x
x
x
x
717
Appendix
Table A.1 (continued)
x
23559
cr,sb
x
x
2–1180
rb
x
4–900
rb
x
1271
x
x
x
x
x
60–293
rb,sb
x
x
x
x
15–399
rb
x
70–4538 sb
x
x
384–
4087
mb
x
x
x
559
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
9–22
x
280–384 mb, sb
x
4–15
x
cr, rb
x
x
54
x
0–367
mb, sb
0.2–1
mb,rb
350
rb
x
4–15
rb
x
0–364
rb
x
x
9–80
rb,sb
x
x
x
0–73
sb
x
x
x
x
x
x
x
x
x
x
x
x
mb
x
x
3334
64
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Amphicontus minutus
Hill, 1940
Amphichondrius
granulatus (Lütken &
Mortensen, 1899)
Amphichondrius laevis
Ziesenhenne, 1940
Amphiodia assimilis
(Lütken & Mortensen,
1899)
Amphiodia grisea
(Ljungman, 1867)
Amphiodia occidentalis
(Lyman, 1860)
Amphiodia oerstedi
(Lütken, 1856)
Amphiodia platyspina
Nielsen, 1932
Amphiodia sculptilis
Ziesenhenne, 1940
Amphiodia tabogae
Nielsen, 1932
Amphiodia urtica
(Lyman, 1860)
Amphiodia violacea
(Lütken, 1856)
Amphiodia vicina H.
L. Clark, 1940
Amphipholis elevata
Nielsen, 1932
Habitat
560
Table A.1 (continued)
Depth
(m)
Amphipholis granulata
(Lütken & Mortensen,
1899)
Amphipholis laevidisca
H. L. Clark, 1909
Amphipholis pugetana
(Lyman, 1860)
Amphipholis puntarenae
(Lütken, 1856)
Amphipholis squamata
(Delle Chiaje, 1828)
Amphioplus
(Amphioplus)
strongyloplax (H.
L. Clark, 1911)
Amphioplus
(Amphioplus)
coniortodes H. L. Clark,
1918
Amphioplus daleus
(Lyman, 1879)
Amphioplus
philohelminthius
Ziesenhenne, 1940
Amphioplus magellanica
(Mortensen, 1936)
Amphioplus textilis
(Koehler, 1907)
Amphiura anomala
Lyman, 1875
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
280–384
x
x
4–1620
sb
x
0–508
mb
x
0–1200
cr,m,mb,sb,sg,rub,rb
4–1408
mb,sb
x
1207
mb
x
2690–
3219
15–73
rb
x
35–174
mb
x
x
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
300–
1500
x
561
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
6–849
mb,rb,sb
x
x
x
x
0–582
x
44–3017 sb
x
x
x
0–300
x
x
x
0–300
x
1587
mb,rb,sb
x
1207
mb,sb
x
549–
2323
x
333
x
1236
mb,sb
x
80
rb
x
x
(continued)
Appendix
Amphiura arcystata H.
L. Clark, 1911
Amphiura eugeniae
Ljungman, 1867
Amphiura (Amphiura)
diomedeae Lütken &
Mortensen, 1899
Amphiura (Amphiura)
magellanica Ljungman,
1867
Amphiura (Ophionema)
hexacantha Nielsen,
1932
Amphiura calbuca
Mortensen, 1952
Amphiura carchara H.
L. Clark, 1911
Amphiura gastracantha
Lütken & Mortensen,
1899
Amphiura gymnogastra
Lütken & Mortensen,
1899
Amphiura gymnopora
Lütken & Mortensen,
1899
Amphiura notacantha
Lütken & Mortensen,
1899
Amphiura otteri
Ljungman, 1872
Depth
(m)
562
Table A.1 (continued)
Depth
(m)
Amphiura polyacantha
Lütken & Mortensen,
1899
Amphiura papillata
Lütken & Mortensen,
1899
Amphiura princeps
Koehler, 1907
Amphiura seminuda
Lütken & Mortensen,
1899
Amphiura serpentina
Lütken & Mortensen
1899
Amphiura verticillata
Ljungman, 1867
Microphiopholis
geminata (Le Conte,
1851)
Microphiopholis
platydisca (Nielsen,
1932)
Microphiopholis
puntarenae (Lütken,
1856)
Ophiocnida californica
Ziesenhenne, 1940
Ophiocnida hispida (Le
Conte, 1851)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1271
x
x
702
x
0–300
9–4096
Appendix
Table A.1 (continued)
x
rb,sb
x
770–
1865
x
x
x
0–82
mb,rb,sb
x
x
x
x
0–137
mb,sb
x
x
x
x
x
6–302
mb,rb,sb
0–794
cr,rb
x
x
x
x
x
x
(continued)
563
Habitat
0–40
rb,sb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
0–134
sb
x
x
x
x
0–134
x
x
x
x
x
x
x
x
18–73
x
0–128
sb
x
1–101
rb
x
x
x
x
x
x
x
x
0–143
mb,rb,sb
x
73–439
55–64
x
x
(continued)
Appendix
Ophiophragmus
chilensis (Müller &
Troschel, 1843)
Ophiophragmus
disacanthus
Ziesenhenne, 1940
Ophiophragmus
marginatus (Lütken,
1856)
Ophiophragmus
ophiactoides
Ziesenhenne, 1940
Ophiophragmus
paucispinus Nielsen,
1932
Ophiophragmus stellatus
Ziesenhenne, 1940
Ophiophragmus
tabogensis Nielsen, 1932
Ophiostigma tenue
Lütken, 1856
Triplodia abdita (A.
M. Clark, 1970)
Family Hemieuryalidae
Amphigyptis perplexa
Nielsen, 1932
Ophiochondrus stelliger
Lyman, 1879
Sigsbeia laevis
Ziesenhenne, 1940
Depth
(m)
564
Table A.1 (continued)
Habitat
43–183
rb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
90–4004 sb
1993
rb
x
x
1062–
1644
147–
4840
733–
1271
x
mb,rb,sb
x
x
mb,sb
x
x
717
x
x
x
x
x
x
x
9–1408
1667–
3914
1244–
1548
702–
1485
cr,mb,sb
x
x
mb,sb
x
x
x
(continued)
565
Sigsbeia lineata Lütken
& Mortensen, 1899
Family Ophiacanthidae
Subfamily
Ophiacanthinae
Ophiacantha antarctica
Koehler, 1900
Ophiacantha bathybia H.
L. Clark, 1911
Ophiacantha contigua
Lütken & Mortensen,
1899
Ophiacantha cosmica
Lyman, 1878
Ophiacantha costata
Lütken & Mortensen,
1899
Ophiacantha cyrena A.
H. Clark, 1916
Ophiacantha deruens
Koehler, 1899
Ophiacantha diplasia H.
L. Clark, 1911
Ophiacantha frigida
Koehler, 1909
Ophiacantha hirta
Lütken & Mortensen,
1899
Ophiacantha
inconspicua Lütken &
Mortensen, 1899
Depth
(m)
Appendix
Table A.1 (continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
519–
1244
mb,sb
x
x
2877
x
2690
x
434–598 sb
x
13–644
x
rb,sb
x
x
183–549 rb
115–
1552
mb,rb,sb
x
x
x
x
32–1538 rb,sb
1171–
1180
2067–
5203
717
999–
1865
x
x
mb
x
x
x
Appendix
Ophiacantha marsupialis
Lyman, 1878
Ophiacantha
moniliformis Lütken &
Mortensen, 1899
Ophiacantha pacifica
Lütken & Mortensen,
1899
Ophiacantha paucispina
Lütken & Mortensen,
1899
Ophiacantha pentacrinus
Lütken, 1869
Ophiocantha phragma
Ziesenhenne, 1940
Ophiacantha
quadrispina H. L. Clark,
1917
Ophiacantha
rhachophora H.
L. Clark, 1911
Ophiacantha rosea
Lyman, 1878
Ophiacantha savagica
Tommasi, 1976
Ophiacantha sentosa
Lyman, 1878
Ophiacantha similis A.
H. Clark, 1916
Ophiacantha spinifera
Lütken & Mortensen,
1899
Habitat
566
Table A.1 (continued)
x
(continued)
Ophiacantha vivipara
Ljungman, 1870
Ophiolebes mortenseni
A. H. Clark, 1916
Ophiotreta valenciennesi
valenciennesi (Lyman,
1879)
Subfamily
Ophiohelinae
Ophiotholia spathifer
(Lyman, 1879)
Subfamily
Ophioplinthacinae
Ophiomitrella chilensis
Mortensen, 1952
Ophiophthalmus
normani (Lyman, 1879)
Ophiothamnus dupla
Tommasi, 1976
Ophiurothamnus laevis
(Lütken & Mortensen,
1899)
Subfamily
Ophiotominae
Ophiotoma paucispina
(Lütken & Mortensen,
1899)
Ophiolimna antarctica
(Lyman, 1879)
Depth
(m)
Habitat
0–1097
rb,kf
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
717
x
549
x
Appendix
Table A.1 (continued)
x
25–300
x
51–2600 mb,rb,sb
x
x
1171–
1180
1008
1643–
4082
88–2750
x
x
mb
x
x
x
x
x
(continued)
567
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
18–34
cr
x
34
mb
x
125–
2870
0–576
cr,mb,rb
x
rb,kf
0–60
rb,sb,mb,rub
0–518
cr,m,rb,sg,rub
x
0–302
cr,rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
1008
x
9–1006
cr,rb
x
589–
1355
sb,rub
x
733
x
x
x
(continued)
Appendix
Family Ophiactidae
Hemipholis cordifera
Bosc, 1802
Hemipholis gracilis
Verrill, 1867
Histampica duplicata
(Lyman, 1875)
Ophiactis asperula
(Philippi, 1858)
Ophiactis kroeyeri
Lütken, 1856
Ophiactis savignyi
(Müller & Troschel,
1842)
Ophiactis simplex (Le
Conte, 1851)
Ophiactis plana Lyman,
1869
Ophiactis profundi
Lütken & Mortensen,
1899
Ophiopholis bakeri
McClendon, 1909
Family
Ophiochitonidae
Ophiochiton carinatus
Lütken & Mortensen,
1899
Ophiochiton fastigatus
Lyman, 1878
Depth
(m)
568
Table A.1 (continued)
Family Ophiocomidae
Ophiocoma aethiops
Lütken, 1859
Ophiocoma alexandri
Lyman, 1860
Ophiocoma dentata
Müller & Troschel, 1842
Ophiocoma erinaceus
Müller & Troschel, 1842
Ophiocoma longispina
H. L. Clak, 1917
Ophiocomella schmitti
A. H. Clark, 1939
Ophiocomella sexradia
(Duncan, 1887)
Ophiocomina nigra
(Abildgaard, in O.F.
Müller, 1789)
Ophiopsila californica
A. H. Clark, 1921
Family
Ophiodermatidae
Diopederma danianum
(Verrill, 1867)
Ophiocrytus granulosus
Nielsen, 1932
Ophioderma appressa
(Say, 1825)
Ophioderma elaps
Lütken, 1856
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–30.5
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
0–70
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.1 (continued)
x
x
8
x
x
5–40
cr,rb
x
x
cr,rb
x
x
x
236–549 sb
x
33–201
rb
x
7–137
cr,mb,rb,sb
x
0–79
rb,sg
x
0–20
cr,rb
x
x
x
x
x
x
x
569
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–20
cr,rb,sb
x
x
183
x
x
x
x
x
0–54
cr,rb,sb
x
15.2–27
rb
x
0–110
mb,rb,sb
x
4–9
rb,sb
x
0–230
mb,sb
x
4–143
sb
x
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
73–155
0–229
x
x
x
x
x
x
x
x
x
x
x
x
mb
x
(continued)
Appendix
Ophioderma panamense
Lütken, 1859
Ophioderma
pentacantha H. L. Clark,
1917
Ophioderma teres
(Lyman, 1860)
Ophioderma vansyoci
Hendler, 1996
Ophioderma variegatum
Lütken, 1856
Ophioderma
sodipallaresi Caso, 1986
Ophiopaepale diplax
(Nielsen, 1932)
Ophiuroconis bispinosa
Ziesenhenne, 1937
Family
Ophionereididae
Ophionereis
albomaculata E.A.
Smith, 1877
Ophionereis annulata
(Le Conte, 1851)
Ophionereis dubia
(Müller & Troschel,
1842)
Ophionereis
eurybrachiplax H.
L. Clark, 1911
Depth
(m)
570
Table A.1 (continued)
Ophionereis perplexa
Ziesenhenne, 1940
Ophionereis porrecta
Lyman, 1860
Ophionereis schayeri
(Müller & Troschel,
1844)
Family Ophiotrichidae
Ophiothrix galapagensis
Lütken & Mortensen,
1899
Ophiothela gracilis
Nielsen, 1932
Ophiothela mirabilis
Verrill, 1867
Ophiothrix magnifica
Lyman, 1860
Ophiothrix rudis Lyman,
1874
Ophiothrix spiculata Le
Conte, 1851
Family Ophiuridae
Subfamily
Ophiolepidinae
Ophiolepis crassa
Nielsen, 1932
Ophiolepis grisea H.
L. Clark, 1940
Ophiolepis pacifica
Lütken, 1856
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–73
cr,rb
x
0–30
rb
x
x
x
Appendix
Table A.1 (continued)
x
x
0–549
cr,mb,rb,sb
x
7–9
cr,rb
x
5–20
cr,mb,rb
x
0–20
cr,mb,rb,sb
0–1
rb,cr,sb
0–2059
cr,mb,rb,sb
x
6–230
mb,rb,sb
x
7–13
mb
0–18
cr,rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
571
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1–12
cr,rb,sb,rub
x
1–110
cr,mb,sb
x
878–
5203
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
702–
1485
x
x
x
x
51–2906 mb,sb
x
267–902 mb,rb
x
0–74
x
rb,sg
x
x
x
x
x
x
x
17–1230 mb,rb,sb
1408–
2487
x
x
x
x
x
x
x
2417–
2487
x
x
x
(continued)
Appendix
Ophiolepis plateia
Ziesenhenne, 1940
Ophiolepis variegata
Lütken, 1856
Ophiomusium glabrum
Lütken & Mortensen,
1899
Ophiomusium diomedeae
Lütken & Mortensen,
1899
Ophiomusium lymani
Thomson, 1873
Ophiomusium variabile
Lütken & Mortensen,
1899
Ophioplocus esmarki
Lyman, 1874
Ophioplocus hancocki
Ziesenhenne, 1935
Ophiosphalma jolliense
(McClendon, 1909)
Ophiozonella alba
(Lütken & Mortensen,
1899)
Ophiozonella clypeata
(Lyman, 1883)
Ophiozonella contigua
(Lütken & Mortensen,
1899)
Ophiozonella falklandica
Mortensen, 1936
Depth
(m)
572
Table A.1 (continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
770–
1245
770–
1158
mb,sb
x
x
x
x
1207–
1244
x
840–
4082
mb,sb
245–
3714
rb,sb
x
x
x
x
x
x
x
2197
1152
x
mb
x
x
x
x
51–1820 mb
x
(continued)
573
Subfamily
Ophioleucinae
Ophiernus adspersus
adspersus Lyman, 1883
Ophiernus adspersus
annectens Lütken &
Mortensen, 1899
Ophiernus polyporum
Lütken & Mortensen,
1899
Ophiernus seminudus
Lütken & Mortensen,
1899
Subfamily Ophiurinae
Amphiophiura abcisa
(Lütken & Mortensen,
1899)
Amphiophiura
irregularis Ziesenhenne,
1940
Amphiophiura obtecta
(Lütken & Mortensen,
1899)
Amphiophiura oligopora
(H. L. Clark, 1913)
Amphiophiura
paucisquama
Ziesenhenne, 1940
Amphiophiura superba
(Lütken & Mortensen,
1899)
Depth
(m)
Appendix
Table A.1 (continued)
Depth
(m)
Amphiophiura vemae
Kyte, 1987
Gymnophiura concava
Tommasi, 1976
Gymnophiura mollis
Lütken & Mortensen,
1899
Ophiocten amitinum
Lyman, 1878
Ophiocten hastatum
Lyman, 1878
Ophiogona doederleini
(Koehler, 1901)
Ophionotus victoriae
Bell, 1902
Ophiomastus bulufonica
Tommasi, 1976
Ophiomastus molinae
Castillo-Alarcon, 1968
Ophiomastus tuberculata
Tommasi, 1976
Ophiomisidium leurum
Ziesenhenne, 1940
Ophiosteira koehleri
A. H. Clark, 1917
Ophiophyllum
marginatum A. H. Clark,
1916
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
3739–
4124
574
Table A.1 (continued)
x
x
2417–
2487
29–920
x
x
x
rb,sb,kf
1159–
2877
x
x
x
x
x
x
x
x
x
5–752
mb,rb,sb
x
1171–
1180
intertidal rb
x
1171–
1180
x
x
x
733
717
x
x
Appendix
(continued)
Depth
(m)
Ophioplinthus divisa
(Lütken & Mortensen,
1899)
Ophioplinthus inflata
(Koehler, 1897)
Ophioplinthus inornata
(Lyman, 1878)
Ophioplinthus nexila
(Kyte, 1987)
Ophiotypa simplex
Koehler, 1897
Ophiura
(Ophiuroglypha)
irrorata irrorata
(Lyman, 1878)
Ophiura flagellata
(Lyman, 1878)
Ophiura leptoctenia
H. L. Clark, 1911
Ophiura luetkenii
(Lyman, 1860)
Ophiura lymani
(Ljungman, 1871)
Ophiura nana (Lütken &
Mortensen, 1899)
Ophiura plana (Lütken
& Mortensen, 1899)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
2075–
2487
x
1484
x
1485–
2418
x
Appendix
Table A.1 (continued)
x
x
3652–
3811
4055869
x
mb,sb
x
128–
mb
2014
27–3239 sb,rub
x
x
0–1097
mb,rb,sb
x
30–800
rb,sb,kf
x
x
x
x
x
x
x
x
x
1650
2070–
3241
x
x
x
x
x
x
x
x
(continued)
575
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
3667
x
x
x
1–845
rb, sb
x
13–641
x
87–550
sb
0–150
cr,mb,rb,sb
0–150
rb
2–183
cr,mb,rb,sb
55–600
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
45
48–274
x
x
x
x
x
x
x
(continued)
Appendix
Ophiura stenobrachia
H. L. Clark, 1917
Theodoria madseni
Tommasi, 1976
Class Echinoidea
Order Cidaroida
Family Cidaridae
Austrocidaris
canaliculata (A. Agassiz,
1863)
Austrocidaris spinulosa
Mortensen, 1910
Centrocidaris
doederleini (A. Agassiz,
1898)
Eucidaris thouarsii
(Valenciennes, 1846)
Eucidaris thouarsii
galapagensis
Doederlein, 1887
Hesperocidaris
asteriscus H. L. Clark,
1948
Hesperocidaris dubia
(H. L. Clark, 1907)
Hesperocidaris
houstonianus H. Clark,
1939
Hesperocidaris
panamensis (A. Agassiz,
1898)
Habitat
576
Table A.1 (continued)
Depth
(m)
Hesperocidaris perplexa
(H. L. Clark, 1907)
Family Ctenocidaridae
Aporocidaris milleri
(A. Agassiz, 1898)
Family Histocidaridae
Histocidaris cobosi (A.
Agassiz, 1898)
Order Echinothurioida
Family Echinothuriidae
Araeosoma eurypatum
A. Agassiz & H.
L. Clark, 1909
Araeosoma leptaleum
A. Agassiz & H.
L. Clark, 1909
Tromikosoma hispidum
(A. Agassiz, 1898)
Tromikosoma
panamense (A. Agassiz,
1898)
Family
Kamptosomatidae
Kamptosoma asterias
(A. Agassiz, 1881)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
10–1500 rb,mb,sb
x
300–
3937
x
mb,rb,sb
x
x
x
x
x
x
x
x
702
x
1227
x
740–
1046
rb
x
1820–
3375
2054–
3374
rb,sb
x
mb,rb
x
1020–
1278
mb
x
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
(continued)
577
Habitat
2830–
3900
mb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1625–
3381
x
0–90
cr,mb,rb,sb
x
0–125
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–113
mb,cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
1–3
rb, sb
x
0–46
cr,rb
x
3–5
cr
x
(continued)
Appendix
Order Diadematoida
Family
Aspidodiadematidae
Plesiodiadema
globulosum (A. Agassiz,
1898)
Plesiodiadema horridum
(A. Agassiz, 1898)
Family Diadematidae
Astropyga pulvinata
(Lamarck, 1816)
Centrostephanus
coronatus (Verrill, 1867)
Centrostephanus
rodgersii (A. Agassiz,
1863)
Centrostephanus sylviae
Fell, 1975
Diadema mexicanum
A. Agassiz, 1863
Diadema paucispinum
(A. Agassiz, 1863)
Diadema savignyi
Michelin, 1845
Echinothrix calamaris
(Pallas, 1774)
Echinothrix diadema
(Linnaeus, 1758)
Order Pedinoida
Family Pedinidae
Depth
(m)
578
Table A.1 (continued)
Depth
(m)
Caenopedina diomedeae
Mortensen, 1939
Order Salenioida
Family Saleniidae
Salenocidaris miliaris
(A. Agassiz, 1898)
Order Arbacioida
Family Arbaciidae
Arbacia dufresnii
(Blainville, 1825)
Arbacia spatuligera
(Valenciennes, 1846)
Arbacia stellata
(Blainville, 1825)
Dialithocidaris
gemmifera A. Agassiz,
1898
Tetrapygus niger
(Molina, 1872)
Order Camarodonta
Family Parechinidae
Loxechinus albus
(Molina, 1782)
Order Temnopleuroida
Family
Temnopleuridae
Pseudechinus
magellanicus (Philippi,
1857)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
723–850
x
1159–
3376
x
0–315
rb
0–50
sb,rb,mb
0–92
cr,mb,rb, sb
3193–
3279
x
x
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–25
rb,sb
x
x
0–340
rb
x
x
0–820
rb, kf
x
x
579
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1–207
cr,mb,rb,sb
x
x
0–300
cr,rb,mb,sb
0–134
rb,cr,sb
0–75
cr,rb,sb
1–6
rb
0–55
cr,rb,sb,mb
x
x
x
x
x
x
50–1200 mb,rb,sb
x
4–84
rb,sb
x
0–161
mb,rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Family Toxopneustidae
Lytechinus anamesus H.
L. Clark, 1912
Lytechinus panamensis
Mortensen, 1921
Lytechinus pictus
(Verrill, 1867)
Lytechinus
semituberculatus
(Valenciennes in
L. Agassiz, 1846)
Tripneustes depressus
A. Agassiz, 1863
Tripneustes gratilla
(Linnaeus, 1758)
Toxopneustes roseus
(A. Agassiz, 1863)
Family
Strongylocentrotidae
Allocentrotus fragilis
Jackson, 1912
Strongylocentrotus
franciscanus
(A. Agassiz, 1863)
Strongylocentrotus
purpuratus (Stimpson,
1857)
Order Echinoida
Family Echinidae
Family Echinometridae
Depth
(m)
580
Table A.1 (continued)
Caenocentrotus gibbosus
(L. Agassiz & Desor,
1846)
Echinometra insularis H.
L. Clark, 1912
Echinometra mathaei
(Blainville, 1825)
Echinometra oblonga
(Blainville, 1825)
Echinometra vanbrunti
A. Agassiz, 1863
Echinostrephus
aciculatus A. Agassiz,
1863
Order Clypeasteroida
Family Clypeasteridae
Clypeaster elongatus H.
L. Clark, 1948
Clypeaster europacificus
H. L. Clark, 1914
Clypeaster ochrus
H. L. Clark, 1914
Clypeaster reticulatus
Linnaeus, 1758
Clypeaster rotundus
(A. Agassiz, 1863)
Clypeaster speciosus
Verrill, 1870
Family Dendrasteridae
Depth
(m)
Habitat
0–35
rb
0–50
rb,sb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
x
x
Appendix
Table A.1 (continued)
x
x
0–34
cr,rb
x
x
0–106
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
10
sb
x
0–402
mb,sb,rb
x
x
x
x
x
x
0–162
rb,sb
x
x
x
x
x
x
x
x
0–92
rb,sb,mb
x
0–128
rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
581
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–232
mb,rb,sb
x
13–22
sb
x
0–30
sb
x
0–30
sb
x
4–15
sb
1–3
sb
0–120
sb
1–10
sb
0–36.6
sb
0–7
sb
3–90
sb
0–82
mb,sb
1–5
sb
x
1–10
sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Dendraster excentricus
(Eschscholtz, 1831)
Dentraster laevis H.
L. Clark, 1948
Dendraster mexicanus
H. L. Clark, 1948
Dendraster vizcainoensis
Grant & Hertlein, 1938
Family Mellitidae
Encope cocosi
H. L. Clark, 1948
Encope ecuadorensis
H. L. Clark, 1948
Encope grandis
L. Agassiz, 1841
Encope insularis
H. L. Clark, 1948
Encope irregularis
H. L. Clark, 1948
Encope laevis
H. L. Clark, 1948
Encope michelini
L. Agassiz, 1841
Encope micropora
L. Agassiz, 1841
Encope micropora
ecuadorensis
H. L. Clark, 1948
Encope micropora
insularis H. L. Clark,
1948
Depth
(m)
582
Table A.1 (continued)
Encope micropora
irregularis H. L. Clark,
1948
Encope micropora
tetrapora L. Agassiz,
1841
Encope pacifica (Verrill,
1867)
Encope perspectiva
L. Agassiz, 1841
Encope wetmorei
A. H. Clark, 1946
Mellita grantii
Mortensen, 1948
Mellita kanakoffi
Durham, 1961
Mellita longifissa
Michelin, 1858
Mellita notabilis
H. L. Clark, 1947
Mellitella stokesii
(L. Agassiz, 1841)
Order Cassiduloida
Family Cassidulidae
Rhyncholampas pacificus
(A. Agassiz, 1863)
Order Holasteroida
Family Pourtalesiidae
Cystocrepis setigera
(A. Agassiz, 1898)
Depth
(m)
Habitat
1
sb
3
sb
0–40
sb
8–27
mb,sb
x
x
x
1–45
sb
x
x
x
0–10
sb
x
0–102.9
sb
x
0–60
sb
x
0.5–0.9
sb
x
0–10
sb, mb
6–134
rb,sb
3182
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
583
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
3374
x
755–
4400
x
x
1455–
5200
mb
x
589–
3279
9–237
mb,sb
x
sb,mb,rb
x
x
x
x
x
x
x
x
x
x
x
x
0–240
sb,cr,rb
x
x
x
x
x
x
x
x
96
x
0–200
sb
x
0–18
sb
x
x
x
x
x
x
x
(continued)
Appendix
Pourtalesia tanneri
A. Agassiz, 1898
Familly Urechinidae
Cystechinus loveni
A. Agassiz, 1898
Pilematechinus rathbuni
(A. Agassiz, 1898)
Urechinus naresianus
A. Agassiz, 1879
Order Spatangoida
Family Aeropsidae
Aeropsis fulva
(A. Agassiz, 1898)
Family Brissidae
Brissopsis columbaris
A. Agassiz, 1898
Brissopsis pacifica
(A. Agassiz, 1898)
Brissus agassizii
Doederlein, 1885
Brissus obesus Verrill,
1867
Brissus latecarinatus
(Leske, 1778)
Meoma frangibilis
Chesher, 1970
Meoma ventricosa
grandis Gray, 1851
Metalia nobilis Verrill,
1867
Habitat
584
Table A.1 (continued)
Depth
(m)
Metalia spatagus
(Linnaeus, 1758)
Rhabdobrissus pacificus
H. L. Clark, 1940
Family Hemiasteridae
Hemiaster tenuis
(A. Agassiz, 1898)
Family Loveniidae
Homolampas hastata
A. Agassiz, 1898
Lovenia cordiformis
A. Agassiz, 1872
Family
Macropneustidae
Argopatagus aculeata
(A. Agassiz, 1898)
Family Maretiidae
Nacospatangus oblonga
(Mortensen, 1950)
Family Palaeotropidae
Scrippsechinus fisheri
Allison, Durham &
Mintz, 1967
Family Prenasteridae
Tripylus excavatus
Philippi, 1845
Family Schizasteridae
Abatus cavernosus
(Philippi, 1845)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
6–137
sb,mb,sb
x
x
980–
4027
1785–
3376
0–210
1952
mb,sb
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
270–460
x
0–130
x
0–760
x
585
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
27–804
x
x
0–209
mb,rb,sb
x
9–2817
mb,sb
x
x
x
x
x
x
x
x
x
x
124–730
x
x
10–40
mb,sb
x
x
x
x
0–100
sb
10–644
mb,rb,sb
3241–
4000
1585–
5690
x
x
x
x
x
x
x
(continued)
Appendix
Abatus philippii Lovén,
1871
Aceste ovata A. Agassiz
& H. L. Clark, 1907
Agassizia scrobiculata
Valenciennes, 1846
Brisaster latifrons
(A. Agassiz, 1898)
Brisaster moseleyi
(A. Agassiz, 1881)
Brisaster townsendi
(A. Agassiz, 1898)
Moira atropos clotho
Michelin, 1855
Schizaster rotundatus
(Doederlein, 1906)
Tripylaster philippii
(Gray, 1851)
Family Spatangidae
Spatangus californicus
H. L. Clark, 1917
Class Holothuroidea
Order Dendrochirotida
Family Cucumariidae
Abyssocucumis
abyssorum (Théel, 1886)
Abyssocucumis
albatrossi Cherbonnier,
1947
Habitat
586
Table A.1 (continued)
Depth
(m)
Cucumaria chilensis
Ludwig, 1875
Cucumaria flamma
Solís-Marín & LaguardaFigueras, 1999
Cladodactyla crocea
(Lesson, 1830)
Heterocucumis
godeffroyi (Semper,
1868)
Hemioedema spectabilis
(Ludwig, 1882)
Leptopentacta nina
Deichmann, 1941
Leptopentacta nova
Deichmann, 1941
Leptopentacta panamica
Deichmann, 1941
Neocucumis panamensis
Heding & Panning, 1954
Neocucumis veleronis
(Deichmann, 1941)
Pattalus mollis Selenka,
1868
Pentacta panamensis
Verrill, 1867
Pseudocnus californicus
(Semper, 1868)
Pseudocnus dubiosus
(Semper, 1868)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
32–162
x
0–15
rb
0–4300
sb
0–379
rb,sb
14642
sb
x
x
x
x
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
0–22
rb
0–45
rb
x
x
x
x
x
x
0–190
cr,rb,sb
0–300
rb,sb
x
x
x
x
x
x
x
x
x
587
(continued)
Habitat
0–300
rb,sb
0–197
rb,sb
1587–
4088
mb,rb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
x
x
x
x
10–35
sb,rub
5–238
rb
5–42
rb
x
x
x
x
x
x
0–13
x
sb,rb,kf
x
x
x
0.3
x
73
x
x
0–78
cr,sb,rb
6–124
sb
x
x
x
x
x
x
x
(continued)
Appendix
Pseudocnus dubiosus
leoninus (Semper, 1868)
Pseudocnus perrieri
(Eckman, 1927)
Staurocucumis
abyssorum (Théel, 1886)
Stereoderma laevigata
Verrill, 1876
Thyonella mexicana
(Deichmann, 1941)
Trachythyone lechleri
(Lampert, 1885)
Trachythyone peruana
(Semper, 1868)
Family Phyllophoridae
Allothyone mexicana
(Deichmann, 1946)
Athyonidium chilensis
(Semper, 1868)
Euthyonidium ovulum
Deichmann, 1938
Euthyonidium veleronis
Deichmann, 1937
Pentamera beebei
Deichmann, 1938
Pentamera chierchia
(Ludwig, 1887)
Pentamera chiloensis
(Ludwig, 1887)
Depth
(m)
588
Table A.1 (continued)
Pentamera zacae
Deichmann, 1938
Phyrella aculeata
(Ludwig, 1894)
Thyone bidentata
Deichmann, 1941
Thyone neofusus
Deichmann, 1941
Thyone parafusus
Deichmann, 1941
Thyone strangeri
Deichmann, 1941
Family Psolidae
Lissothuria hancocki
(Deichmann, 1941)
Lissothuria mortenseni
Pawson, 1967
Lissothuria ornata
Verrill, 1867
Lissothuria veleronis
(Deichmann, 1941)
Neopsolidium
convergens (Herouard,
1901)
Psolidium disciformis
(Théel, 1886)
Psolidium dorsipes
Ludwig, 1886
Psolidium ekmani
Deichmann, 1941
Depth
(m)
Habitat
24
mb,rb,sb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
Appendix
Table A.1 (continued)
x
2–30
cr,rb
x
x
x
x
x
25–35
mb,sb
x
0–12
rb
x
1–301
rb
x
x
0–37
cr,rb
x
x
x
x
x
x
x
15
kf
x
8–448
rb
x
11–451
cr,mb,rb,sb
x
x
x
x
x
589
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
2323
x
2323
x
16–110
x
6–1080
rb
0–247
rb
x
x
1271
x
13–302
rb,sb
x
x
x
x
10–567
rb
x
0–110
rb,kf
x
7–1087
rb
7–207
rb
0–7
rb,cr
x
55–91
sb
x
x
x
x
x
x
x
x
(continued)
Appendix
Psolidium eubullatum
Deichmann, 1941
Psolidium gracile
Ludwig, 1894
Psolidium panamense
Ludwig, 1894
Psolidium planum
Deichmann, 1941
Psolus antarcticus
Philippi, 1857
Psolus chitonoides
H. L. Clark, 1901
Psolus digitatus Ludwig,
1894
Psolus diomedeae
Ludwig, 1894
Psolus paradubiosus
Carriol & Féral, 1985
Psolus patagonicus
Ekman, 1925
Psolus squamatus
(Koren, 1844)
Psolus squamatus
segregatus Perrier, 1905
Family
Sclerodactylidae
Afrocucumis ovulum
(Selenka, 1867)
Apentamera lepra
Deichmann, 1941
Habitat
590
Table A.1 (continued)
Athyone glasselli
(Deichmann, 1936)
Euthyonidiella zacae
(Deichmann, 1938)
Neopentamera anexigua
Deichmann, 1941
Neothyone gibber
(Selenka, 1867)
Neothyone gibbosa
Deichmann, 1941
Neothyone panamensis
(Ludwig, 1887)
Pachythyone lugubris
(Deichmann, 1939)
Pachythyone
pseudolugubris
Deichmann, 1941
Order Dactylochirotida
Family Ypsilothuriidae
Ypsilothuria
bitentaculata (Ludwig,
1893)
Order Aspidochirotida
Family Holothuriidae
Actinopyga mauritiana
(Quoy & Gaimard, 1833)
Holothuria (Cystipus)
casoae LaguardaFigueras & Solís-Marín,
2009
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–6
sb
x
17–25
rb
x
8–22
cr
x
0–50
cr,rb
x
x
x
0–14
rb
x
x
x
0–8.6
rb
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
cr,rb
x
12
cr,rb
x
225–
4082
mb,sb
x
x
x
x
x
x
x
x
x
45–100
sb
x
x
591
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1–203
cr,rb,sb
x
x
0–22
mb,sb,rb
x
x
5–26
mb,cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–18
rb
x
x
0–27
rb,sb,cr
x
x
0–306
cr,rb,mb,sb
x
x
0.3–52
cr,rb,sb
x
x
1–17
rb,sb
x
x
0–100
mb,rb,sb,cr
x
x
1–4
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Holothuria (Cystipus)
inhabilis Selenka, 1867
Holothuria (Cystipus)
rigida (Selenka, 1867)
Holothuria (Halodeima)
atra (Jaeger, 1833)
Holothuria (Halodeima)
chilensis Semper, 1868
Holothuria (Halodeima)
inornata Semper, 1868
Holothuria (Halodeima)
kefersteini (Selenka,
1867)
Holothuria
(Lessonothuria) pardalis
Selenka, 1867
Holothuria
(Mertensiothuria) hilla
Lesson, 1830
Holothuria
(Mertensiothuria)
leucospilota (Brandt,
1835)
Holothuria
(Platyperona) difficilis
Semper, 1868
Holothuria
(Platyperona) parvula
(Selenka, 1867)
Depth
(m)
592
Table A.1 (continued)
Holothuria
(Selenkothuria) carere
Honey-Escandón &
Solís-Marín, 2011
Holothuria
(Selenkothuria) lubrica
Selenka, 1867
Holothuria
(Selenkothuria)
portovallartensis Caso,
1954
Holothuria
(Selenkothuria) theeli
Deichmann, 1938
Holothuria
(Semperothuria)
cinerascens (Brandt,
1835)
Holothuria
(Semperothuria) imitans
Ludwig, 1875
Holothuria
(Semperothuria)
languens Selenka, 1867
Holothuria (Stauropora)
pluricuriosa
Deichmann, 1937
Holothuria
(Theelothuria)
paraprinceps
Deichmann, 1937
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
0–6
rb
x
0–55
cr,rb,sb,mb
x
0–59
cr,rb
x
0–55
rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.1 (continued)
x
x
x
x
x
x
0–22
rb,sb,cr
x
0–91
rb,mb,sb
x
0–130
rb,cr,sb
x
0–64
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
593
(continued)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1–121
rb,sb,cr
x
x
0–67
cr,rb,sb
x
x
77–250
mb,rb,sb
x
0.5–16
rb,cr,sb
x
1–18
rb,sb,cr
x
0–36
mb,rb,sb
x
0–61
cr,rb,mb,sb
x
1–180
mb,rb
x
0–20
cr,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Holothuria
(Thymiosycia) arenicola
Semper, 1868
Holothuria
(Thymiosycia) impatiens
(Forskaal, 1775)
Holothuria
(Vaneyothuria) zacae
Deichmann, 1937
Labidodemas
americanum Deichmann,
1938
Labidodemas
maccullochi
(Deichmann, 1958)
Family Stichopodidae
Apostichopus
parvimensis (H. L. Clark,
1913)
Isostichopus fuscus
(Ludwig, 1875)
Parastichopus
californicus (Stimpson,
1857)
Stichopus horrens
Selenka, 1867
Stichopus
monotuberculatus (Quoy
& Gaimard, 1833)
Family Synallactidae
Depth
(m)
594
Table A.1 (continued)
Bathyplotes moseleyi
(Théel, 1886)
Bathyplotes natans
(M. Sars, 1868)
Östergren, 1896
Bathyplotes patagiatus
Fisher, 1907
Capheira sulcata
Ludwig, 1893
Meseres macdonaldi
Ludwig, 1894
Meseres torvus (Théel,
1886)
Mesothuria (Mesothuria)
multipes Ludwig, 1894
Mesothuria (Zygothuria)
lactea (Théel, 1886)
Molpadiodemas
atlanticus (R. Perrier,
1898)
Molpadiodemas
neovillosus O’Loughlin
& Ahearn, 2005
Molpadiodemas
ustulatus O’Loughlin &
Ahearn, 2005
Molpadiodemas villosus
(Théel, 1886)
Molpadiodemas
violaceus (Théel, 1886)
Depth
(m)
Habitat
24–690
mb,rb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
1644
x
1644
x
2877–
4334
1644
x
mb
x
x
x
Appendix
Table A.1 (continued)
x
x
x
725–
4064
mb
x
x
x
x
x
x
2487–
3667
x
x
3667
x
x
595
(continued)
Habitat
1571–
4070
2323
mb
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
2149
x
100–
5203
1158–
3667
2418–
4334
589–
1008
549
mb,sb
1030
mb
1618–
2487
3241
mb,sb
3241–
3670
3667–
4088
mb
x
x
x
x
x
mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Paelopatides confundens
Théel, 1886
Paelopatides suspecta
Ludwig, 1894
Pseudostichopus
macdonaldi (Ludwig,
1894)
Pseudostichopus mollis
Théel, 1886
Pseudostichopus
peripatus (Sluiter, 1901)
Synallactes aenigma
Ludwig, 1894
Synallactes alexandri
Ludwig, 1894
Synallactes triplax
A. H. Clark, 1920
Synallactes virgulasolida
Massin & Hendrickx,
2010
Order Elasipodida
Family Deimatidae
Deima validum
pacificum Ludwig, 1894
Oneirophanta mutabilis
affinis Ludwig, 1894
Oneirophanta mutabilis
mutabilis Théel, 1879
Oneirophanta setigera
(Ludwig, 1893)
Depth
(m)
596
Table A.1 (continued)
Family Elpidiidae
Achlyonice ecalcarea
Théel, 1879
Amperima naresi (Théel,
1882)
Amperima vitjazi
Gebruk, 1988
Elpidia atakama
Belyaev, 1971
Elpidia chilensis
Belyaev, 1971
Peniagone anamesa
(A. H. Clark, 1920)
Peniagone diaphana
(Théel, 1882)
Peniagone elongata
(Théel, 1879)
Peniagone gracilis
(Ludwig, 1894)
Peniagone intermedia
Ludwig, 1893
Peniagone papillata
Hansen, 1975
Peniagone vitrea Théel,
1882
Scotoplanes clarki
Hansen, 1975
Scotoplanes globosa
Théel, 1879
Depth
(m)
Habitat
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
1608–
3587
2010–
7130
mb,sb
x
mb
Appendix
Table A.1 (continued)
x
x
7720
mb
x
2710–
4600
mb
x
x
4140–
4160
4140–
4160
2487–
4160
2418–
3667
mb
x
mb
x
mb
x
mb
x
x
x
x
x
x
1160–
4507
3570–
5107
545–
6770
mb
mb
mb
x
x
x
x
x
x
x
597
(continued)
Depth
(m)
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
2418
466–
5083
199–
2599
4140–
4160
0–4505
x
x
mb,sb
x
x
x
x
pelagic
x
pelagic
x
x
x
x
2418
x
x
x
x
978–
2323
mb
1158–
4700
3852–
4189
mb
x
x
x
x
x
x
x
x
x
mb
x
x
3334–
4160
2418–
4160
mb
x
mb
x
x
x
x
x
x
x
x
(continued)
Appendix
Family Laetmogonidae
Laetmogone theeli
Ludwig, 1893
Laetmogone violacea
Théel, 1879
Pannychia moseleyi
Théel, 1882
Family Pelagothuridae
Enypniastes eximia
Théel, 1882
Pelagothuria natatrix
Ludwig, 1893
Family Psychropotidae
Benthodytes incerta
Ludwig, 1893
Benthodytes
sanguinolenta Théel,
1882
Benthodytes typica
Théel, 1882
Psychronaetes hanseni
Pawson, 1983
Psychropotes depressa
(Théel, 1882)
Psychropotes longicauda
Théel, 1882
Psychropotes verrucosa
(Ludwig, 1894)
Order Molpadiida
Habitat
598
Table A.1 (continued)
Family Caudinidae
Ceraplectana
trachyderma H. L. Clark,
1908
Hedingia californica
(Ludwig, 1894)
Paracaudina chilensis
chilensis (Müller, 1850)
Family Molpadiidae
Molpadia antarctica
(Théel, 1886)
Molpadia intermedia
(Ludwig, 1894)
Molpadia granulata
(Ludwig, 1894)
Molpadia musculus
Risso, 1826
Molpadia spinosa
(Ludwig, 1893)
Order Apodida
Family Chiridotidae
Chiridota aponocrita A.
H. Clark, 1920
Chiridota pacifica
Heding, 1928
Chiridota pisanii
Ludwig, 1886
Chiridota rigida Semper,
1868
Depth
(m)
Habitat
5203
mb
85–2850 sb,mb
0–990
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
mb,sb
x
x
x
80–1218 mb,sb
53–2972 mb
mb
2690–
5869
37–6134 mb,sb
x
x
x
x
3279
9–137
Appendix
Table A.1 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
cr,rb,sb
x
x
x
0–228
mb,sb
2–9
cr,rb,sb
x
x
599
(continued)
Taeniogyrus contortus
(Ludwig, 1875)
Trochodota purpurea
(Lesson, 1830)
Family Myriotrochidae
Myriotrochus giganteus
A. H. Clark, 1920
Myriotrochus
(Oligotrochus) bathybius
A. H. Clark, 1920
Family Synaptidae
Anapta fallax Lampert,
1889
Epitomapta tabogae
Heding, 1928
Euapta godeffroyi
(Semper, 1868)
Polyplectana oculata
Heding, 1928
Protankyra abyssicola
(Théel, 1886)
Protankyra brychia
(Verrill, 1885)
Protankyra pacifica
(Ludwig, 1894)
Depth
(m)
Habitat
0–560
mb,sb,kf
x
0–36
rb,sb
x
3667
mb
3667
mb
0–350
sb
0–10
0–79
MEX REV GUA SAL HON NIC CRC COC PAN COL MAL GAL ECU PER CHL PAS
x
x
x
x
x
cr,rb,sb
600
Table A.1 (continued)
x
x
x
x
x
x
x
x
3900–
4000
870–
4990
mb
x
x
x
x
x
x
x
Appendix
Habitat classification: cr Coral Reefs, m mangroves, mb muddy bottom, rb rocky bottom, sb sandy bottom, sg seagrass, rub rubble bottom, kf kelp forest. Countries
abbreviation: MEX Mexico, REV Revillagigedo Archipelago, GUA Guatemala, SAL El Salvador, HON Honduras, NIC Nicaragua, CRC Costa Rica, COC Cocos Island,
PAN Panama, COL Colombia, MAL Malpelo Island, GAL Galapagos Archipelago, ECU Ecuador, PER Perú, CHL Chile, PAS Eastern Islands
Depth
(m)
Class Crinoidea
Order Comatulida
Family Antedonidae
Antedon bifida (Pennant, 1777)
Antedon nuttingi (A. H. Clark,
1936)
Caryometra alope A. H. Clark,
1940
Caryometra atlantidis A.
H. Clark, 1940
Caryometra lisa A. H. Clark,
1940
Caryometra monilicirra
A. H. Clark, 1940
Caryometra spinosa
A. H. Clark, 1940
Caryometra tenuipes
(A. H. Clark, 1908)
Coccometra guttata
A. H. Clark, 1918
Coccometra hagenii (Pourtalès,
1867)
Coccometra nigrolineata
A. H. Clark, 1918
Comatonia cristata (Hartlaub,
1912)
Ctenantedon kinziei Meyer,
1972
Hypalometra defecta
(Carpenter, 1888)
Isometra vivipara Mortensen,
1917
Leptometra celtica (M’Andrew
& Barrett, 1858)
Phrixometra nutrix (Mortensen,
1918)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–400
rb,rub
364–429 rb
x
x
x
475–640 rb
x
366–530 mb
702
Appendix
Table A.2 Taxonomic list of the Echinoderms of the Atlantic Ocean and Caribbean Sea of Latin America and the Canary Islands
x
x
rb
x
x
x
386
rb
x
288–480 mb,rb,rub
x
14–1046 rb,rub
x
40–987
cr,rb,rub
x
50–396
cr,rb
x
Sep-49
cr
46–400
cr,mb,rb
79–350
mb,sb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
137–150 mb,rb,sb
x
x
601
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
219
x
210–
4829
309–
1033
mb,rb
x
mb,rb
x
860–
1674
mb,sb
2–1033
cr,rb,sb,sg,rub
x
3–1033
cr,rb,sb
x
58–373
cr,rb
24–479
cr,mb,rb,rub
0–640
cr,br,rub
x
x
0–334
cr,rb
x
x
55–777
mb
x
3–124
cr,rb
x
10–560
cr,br
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
466–960 rb,rub
x
3–695
cr,br
x
3–155
cr
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Poliometra prolixa (Sladen,
1881)
Trichometra cubensis
(Pourtalès, 1869)
Zenometra columnaris
(Carpenter, 1881)
Family Pentametrocrinidae
Pentametrocrinus atlanticus
(Perrier, 1883)
Superfamily Comasteracea
Family Comasteridae
Comactinia echinoptera
(Müller, 1840)
Comactinia meridionalis
meridionalis (L. Agassiz, 1865)
Comactinia meridionalis
hartlaubi Messing, 1978
Comissia venustus
(A. H. Clark, 1909)
Davidaster discoideus
(Carpenter, 1888)
Davidaster rubiginosus
(Pourtalès, 1869)
Leptonemaster venustus
A. H. Clark, 1909
Nemaster grandis A. H. Clark,
1909
Neocomatella alata (Pourtalès,
1878)
Neocomatella europaea
A. H. Clark, 1913
Neocomatella pulchella
(Pourtalès, 1878)
Family Colobometridae
Analcidometra armata
(Pourtalès, 1869)
Habitat
602
Table A.2 (continued)
Depth
(m)
Analcidometra caribbea
(A. H. Clark, 1908)
Family Charitometridae
Crinometra brevipinna
brevipinna (Pourtalès, 1867)
Crinometra brevipinna
concinna A. H. Clark, 1909
Crinometra brevipinna
diadema (Hartlaub, 1912)
Crinometra brevipinna
granulifera (Pourtalès, 1878)
Crinometra brevipinna
margaritacea A. H. Clark, 1909
Crinometra brevipinna pulchra
A. H. Clark, 1909
Family Thalassometridae
Horaeometra duplex
(Carpenter, 1888)
Stylometra spinifera
(Carpenter, 1881)
Family Tropiometridae
Tropiometra carinata
(Lamarck, 1816)
Family Atelecrinidae
Atelecrinus balanoides
Carpenter, 1881
Family Bathycrinidae
Bathycrinus gracilis WyvilleThomson, 1877
Monachocrinus caribbeus
(A. H. Clark, 1908)
Family Bourgueticrinidae
Conocrinus lofotensis (Sars,
1868)
Democrinus conifer
(A. H. Clark, 1909)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
23–26
69–795
x
cr,rb
x
x
x
x
x
201–367 rb
x
320–457 rb
x
185–475 rb
x
320–457 rb
x
484–567 rb
x
159–567 rb
x
69–658
cr,rb,sb,rub
x
0–84
cr,rb,rub
512–
2890
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
x
684–
1574
mb,rb
x
x
x
x
155–
1750
cr,mb,rb
x
x
x
x
x
x
x
603
(continued)
Habitat
374–
1574
70–650
mb
mb,rb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
421–887 mb
x
x
x
x
x
x
154–971 cr,mb,rb
x
80.7
rb
220–
1200
154–
1219
cr,rb,rub
x
mb,rb,rub
x
x
x
x
x
x
x
x
x
x
x
x
170–412 cr,mb,rb
x
x
x
9–700
rb
x
x
x
35–66
sb
22–576
cr,mb,sb,rub
110–641 mb,sb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Democrinus brevis
(A. H. Clark, 1909)
Democrinus rawsoni
(Pourtalès, 1874)
Family Septocrinidae
Rouxicrinus vestitus Mironov &
Pawson, 2010
Order Isocrinida
Family Isocrinidae
Endoxocrinus (Diplocrinus)
wyvillethomsoni (Jeffreys,
1870)
Endoxocrinus parrae (Gervais,
1835)
Endoxocrinus prionodes
(H. L. Clark, 1941)
Isocrinus blakei (Carpenter,
1884)
Neocrinus decorus Thomson,
1864
Family Isselicrinidae
Cenocrinus asterius (Linnaeus,
1767)
Order Cyrtocrinida
Family Holopodidae
Holopus rangii Orbigny, 1837
Class Asteroidea
Order Paxillosida
Family Astropectinidae
Astropecten acutiradiatus
Tortonese, 1956
Astropecten alligator Perrier,
1881
Astropecten americanus
Verrill, 1880
Depth
(m)
604
Table A.2 (continued)
Astropecten antillensis Lütken,
1860
Astropecten aranciacus
(Linnaeus, 1758)
Astropecten articulatus Say,
1825
Astropecten brasiliensis Müller
& Troschel, 1842
Astropecten brasiliensis riensis
Doederlein, 1917
Astropecten
caribemexicanensis Caso, 1990
Astropecten cingulatus Sladen,
1883
Astropecten comptus Verrill,
1915
Astropecten duplicatus Gray,
1840
Astropecten hermatophilus
Sladen, 1883
Astropecten irregularis
(Pennant, 1777)
Astropecten marginatus Gray,
1840
Astropecten nitidus Verrill,
1915
Astropecten spiniphorus
Madsen, 1950
Astropectinides mesactus
(Sladen, 1883)
Bathybiaster loripes Sladen,
1889
Blakiaster conicus Perrier,
1881
Dipsacaster antillensis
Halpern, 1968
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
3–278
mb,sb,rub
x
1–180
sb,rub
x
x
x
x
Appendix
Table A.2 (continued)
x
0–256
mb,sb
2–60
sb
x
x
x
x
x
x
x
x
x
x
18–66
sb
x
51.3
rub
x
x
x
x
x
x
x
x
x
x
x
16–1350 mb,sb,rub
x
18–130
sb,rub
x
0–550
mb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
15–1500
x
10–1000 mb,sb,rub
x
0.5–130
mb,sb,rub
x
11–686
mb,sb,rub
x
x
x
x
x
x
x
x
x
x
45
x
x
x
80–165
x
80–500
mb,sb
55–366
sb
113–
3627
cr
x
x
x
x
x
x
x
x
605
(continued)
Depth
(m)
Dytaster grandis nobilis
Sladen, 1889
Dytaster insignis Perrier, 1884
Leptychaster kerguelenensis
E. A. Smith, 1876
Lonchotaster tartareus Sladen,
1889
Persephonaster echinulatus
H. L. Clark, 1941
Persephonaster leptactis
H. L. Clark, 1941
Persephonaster patagiatus
(Sladen, 1889)
Plutonaster agassizi agassizi
Verrill, 1880
Plutonaster bifrons (WyvilleThomson, 1873)
Plutonaster efflorescens
(Perrier, 1884)
Psilaster andromeda florae
(Verrill, 1878)
Psilaster cassiope Sladen, 1889
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
sb
x
x
x
4390
x
196–720 cr,mb,sb
2928–
3294
733–
mb,sb
2165
70–3110 cr,mb,sb
800–
2970
666–
mb,sb
2061
1409–
mb,sb
1629
151–
mb,sb
1680
100–800
24–139
mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
50–1400 mb
5–306
mb,sb
70–4605 sb,rub
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Psilaster herwigi (Bernasconi,
1972)
Tethyaster grandis (Verrill,
1899)
Tethyaster subinermis (Philippi,
1837)
Tethyaster vestitus vestitus
(Say, 1825)
Family Ctenodiscidae
Ctenodiscus australis Lütken,
1871
3305–
4846
2209–
3654
27–183
Habitat
606
Table A.2 (continued)
Depth
(m)
Ctenodiscus crispatus (Retzius,
1805)
Family Goniopectinidae
Goniopecten demonstrans
Perrier, 1881
Prionaster elegans Verrill,
1899
Family Luidiidae
Luidia alternata alternata (Say,
1825)
Luidia bardabensis Perrier,
1881
Luidia ciliaris (Philippi, 1837)
Luidia clathrata (Say, 1825)
Luidia heterozona barimae
John & A. M. Clark, 1954
Luidia lawrencei Hopkins &
Knott, 2010
Luidia ludwigi scotti Bell, 1917
Luidia patriae Bernasconi,
1941
Luidia sagamina aciculata
Mortensen, 1933
Luidia sarsi Düben & Koren,
1845
Luidia sarsi elegans Perrier,
1875
Luidia senegalensis (Lamarck,
1816)
Family Porcellanasteridae
Eremicaster vicinus Ludwig,
1907
Styracaster horridus Sladen,
1883
Thoracaster cylindratus Sladen,
1883
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
1415–
1865
x
300–
mb,sb
1250
208–535 mb,sb
x
0–1500
m,mb,sb
x
60–430
mb,rb,sb
x
1–400
0–175
16–90
sb
mb,rb,sb,rub
mb,sb
x
x
0–175
mb
x
x
x
x
sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
33–157 mb,rb,sb
100–126 mb,sb
71–83
x
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
60–365
mb,sb
x
0–73
mb,sb
x
3950–
7250
3410–
5062
2540–
5990
mb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
sb
mb,sb
x
x
x
x
x
607
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
140–283
x
170–
2940
265–869
mb,sb
1175–
3713
1857–
1958
130–
5062
1266
mb,sb,rb
x
x
x
x
x
x
x
x
x
x
x
x
mb
x
cr,mb,sb,rb
x
sb,rb
x
226–
1339
485–
5062
250–
1958
380–
1470
370–500
mb,sb,rb
x
mb
x
3305
mb
x
x
x
x
x
x
mb,sb,rb
x
mb,sb,rb
x
x
x
x
mb,sb
576
30–200
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
mb,rub
x
(continued)
Appendix
Family Pseudarchasteridae
Pseudarchaster discus Sladen,
1889
Pseudarchaster gracilis
gracilis (Sladen, 1889)
Pseudarchaster parelii (Düben
& Koven, 1846 )
Order Notomyotida
Family Benthopectinidae
Benthopecten simplex simplex
(Perrier, 1881)
Benthopecten spinosus Verrill,
1884
Cheiraster (Barbadosaster)
echinulatus (Perrier, 1875)
Cheiraster (Cheiraster) ludwigi
Fisher, 1913
Cheiraster (Cheiraster) planus
Verrill, 1915
Cheiraster (Cheiraster) sepitus
(Verrill, 1885)
Cheiraster (Christopheraster)
blakei A. M. Clark, 1981
Cheiraster (Christopheraster)
mirabilis (Perrier, 1881)
Cheiraster (Luidiaster) planeta
(Sladen, 1889)
Gaussaster antarcticus (Sladen,
1889)
Pectinaster gracilis Verrill,
1915
Order Valvatida
Family Asterinidae
Allopatiria ocellifera (Gray,
1847)
Habitat
608
Table A.2 (continued)
Asterina fimbriata Perrier, 1875
Asterina gibbosa (Pennant,
1777)
Asterina stellifera (Moebius,
1859)
Asterinides folium (Lütken,
1860)
Asterinides hartmeyeri
(Doederlein, 1910)
Asterinides pompom
(A. M. Clark, 1983)
Asterinopsis pilosa (Perrier,
1881)
Tremaster mirabilis Verrill,
1880
Stegnaster wesseli (Perrier,
1875)
Family Asteropseidae
Poraniella echinulata (Perrier,
1881)
Family Chaetasteridae
Chaetaster nodosus Perrier,
1875
Family Ganeriidae
Cycethra verrucosa (Philippi,
1857)
Ganeria falklandica Gray, 1847
Perknaster sladeni (Perrier,
1891)
Vemaster sudatlanticus
Bernasconi, 1965
Family Goniasteridae
Anthenoides piercei Perrier,
1881
Apollonaster yucatanensis
Halpern, 1970
Depth
(m)
Habitat
0–250
0–130
mb,sb,rub
rb,rub
0–50
mb,sb,rub
0–256
cr,rb,sb,rub
x
0–1
rb
x
3–6
rb
x
11–256
mb,sb
x
150–
1060
0–183
rb,rub
3–339
cr,rb
cr,rb,sb,rub
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
sb,rub
0–135
120–500
sb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
30–1140 mb,rb,sb
0–500
x
Appendix
Table A.2 (continued)
x
x
x
x
5055–
5208
x
x
x
x
x
x
x
x
x
x
20–844
mb,sb
x
1097–
1175
mb,sb
x
x
x
x
x
x
x
x
x
x
609
(continued)
Depth
(m)
Astroceramus brachyactis
H. L. Clark, 1941
Ceramaster grenadensis
(Perrier, 1881)
Ceramaster grenadensis
grenadensis (Perrier, 1881)
Ceramaster grenadensis
patagonicus (Sladen, 1889)
Circeaster americanus
(A. H. Clark, 1916)
Cladaster rudis Verrill, 1899
Diplasiaster productus
(A. H. Clark, 1917)
Floriaster maya Downey, 1980
Habitat
420–
mb,sb
1067
70–3109 mb,sb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
x
x
200–
mb
3109
106–192
mb,sb
500–
1450
150–900 mb,sb
78–567 mb,sb
x
x
x
x
x
x
x
x
x
x
x
mb,sb
x
cr,mb,sb,rb
x
x
x
x
x
mb,rb,sb
x
sb,rb
x
x
x
mb,rb,sb
x
x
x
mb,sb
x
x
x
sb
x
rb,sb
x
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
933–
1024
Goniaster tessellatus (Lamarck, 16–155
1816)
Hippasteria falklandica Fisher, 251–225
1940
Hippasteria phrygiana
108–162
argentinensis Bernasconi, 1961
Litonotaster intermedius
1958–
(Perrier, 1884)
3530
Mediaster bairdi (Verrill, 1882) 640–
1590
Mediaster pedicellaris (Perrier, 197–580
1881)
Nymphaster arenatus (Perrier, 60–3000
1881)
Paragonaster grandis H.
257–540
L. Clark, 1941
Paragonaster subtilis (Perrier, 1845–
1881)
4700
Pawsonaster parvus (Perrier,
30–600
1881)
x
x
x
610
Table A.2 (continued)
Depth
(m)
Peltaster placenta (Müller &
Trochel, 1842)
Plinthaster dentatus (Perrier,
1884)
Rosaster alexandri (Perrier,
1881)
Tessellaster notabilis
H. L. Clark, 1941
Tosia parva (Perrier, 1881)
Family Leilasteridae
Leilaster radians (Perrier,
1881)
Family Mithrodiidae
Mithrodia clavigera (Lamarck,
1816)
Familia Odontasteridae
Acodontaster elongatus
granuliferus (Koehler, 1912)
Diplodontias singularis (Müller
& Troschel, 1843)
Odontaster penicillatus
(Philippi, 1870)
Odontaster hispidus Verrill,
1880
Family Ophidiasteridae
Copidaster cavernicola SolísMarín & Laguarda-Figueras,
2010
Copidaster lymani A. H. Clark,
1948
Hacelia attenuata (Gray, 1840)
Hacelia superba H. L. Clark,
1921
Linckia bouvieri Perrier, 1875
Linckia guildingi Gray, 1840
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
10–1107 mb,sb
60–2910 mb,rb,sb
x
x
60–2940 cr,mb,sb
x
329–575 mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–157
cr,rb,sb
x
40–840
sb,rub
x
0–84
rb,sb
x
8–350
mb,sb,rub
x
13–18
mb
x
0–34
cr,mb,rb
x
1–150
91–200
sb,rub
rb
x
0–380
0–298
cr,rb
cr,sg,rb,sb
x
x
x
x
102–293 sb,rub
x
x
x
100–600
50–1160 mb,sb
x
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
611
(continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–475
37–155
cr,rb,sb
rb
x
x
37–750
cr,rb,sb
x
x
x
x
x
x
x
x
x
52–585
x
x
x
intertidal rub
x
0–330
cr,rb
x
0–105
rb
50–600
rb,sb
x
x
0–1500
x
166–450 cr,mb,rb,rub
18–320
sb,rub
35–680
rb
30–420
sb,rub
x
x
x
x
x
x
180–510 cr,rb,sb
cr,m,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–500
x
1521
rb
x
6–1100
mb,rb,sb,rub
x
x
x
x
x
(continued)
Appendix
Linckia nodosa Perrier, 1875
Narcissia canariensis
(d’Orbigny, 1839)
Narcissia trigonaria Sladen,
1889
Ophidiaster alexandri Verrill,
1915
Ophidiaster bayeri A. H. Clark,
1948
Ophidiaster guildingii Gray,
1840
Ophidiaster ophidianus
(Lamarck, 1816)
Tamaria floridae (Perrier,
1881)
Tamaria halperni Downey,
1971
Family Oreasteridae
Oreaster reticulatus (Linnaeus,
1758)
Family Poraniidae
Marginaster pectinatus Perrier,
1881
Porania (Porania) antarctica
magellanica Studer, 1876
Porania (Porania) pulvillus
insignis Verrill, 1895
Poraniopsis echinaster Perrier,
1891
Poraniopsis mira (de Loriol,
1904)
Family Solasteridae
Laetmaster spectabilis (Perrier,
1881)
Lophaster verrilli A. H. Clark,
1938
Depth
(m)
612
Table A.2 (continued)
Solaster caribbaeus Verrill,
1915
Order Velatida
Family Caymanostellidae
Caymanostella spinimarginata
Belyaev, 1978
Family Korethrasteridae
Peribolaster folliculatus
Sladen, 1889
Remaster gourdoni Koehler,
1912
Remaster palmatus (Perrier,
1881)
Family Myxasteridae
Pythonaster murrayi Sladen,
1889
Family Pterasteridae
Calyptraster coa Sladen, 1882
Calyptraster personatus
(Perrier, 1885)
Diplopteraster clarki
Bernasconi, 1937
Diplopteraster verrucosus
(Sladen, 1882)
Hymenaster anomalus Sladen,
1882
Hymenaster pellucidus
Thomson, 1873
Hymenaster pergamentaceus
Sladen, 1882
Hymenaster regalis Verrill,
1895
Hymenaster rex Perrier, 1885
Pteraster abyssorum (Verrill,
1895)
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
64–869
mb,rb
x
3109–
3493
mb
x
x
81–133
x
x
10–540
x
x
296–585 mb,rb,sb
x
x
x
3477
x
x
x
x
x
74–270
x
rb
x
sb
x
sb
x
1857
1139–
2285
576–
3740
x
x
260–930
2151–
mb,rb
6560
82–177
1984–
2606
1784–
3294
4846
Appendix
Table A.2 (continued)
x
x
x
rb
x
x
rb
x
x
613
(continued)
Habitat
196–
3713
74–341
cr,mb
275–825 mb,sb,rub
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
x
x
mb
x
549–594
480–
1780
91–466
x
x
27–500
271–466 mb
777
x
x
mb
x
mb,sb
x
x
x
79–2804
x
60–475
mb,rb,sb
x
2–250
rb,sg,rub
0–360
mb,rb,sb,sg
0–55
cr,m,mb,rb,sb,rub
13–119
mb,sb
0–175
cr,rb,sb,sg,rub
x
10–200
cr,mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Pteraster acicula (Downey,
1970)
Pteraster affinis lebruni Perrier,
1891
Pteraster caribbaeus Perrier,
1881
Pteraster gibber (Sladen, 1882)
Pteraster militarioides
militarioides H. L. Clark, 1941
Pteraster militarioides stoibe
H. L. Clark, 1941
Pteraster militaris (Müller,
1776)
Pteraster personatus Sladen,
1891
Pteraster rugosus H. L. Clark,
1941
Pteraster stellifer Sladen, 1882
Order Spinulosida
Family Echinasteridae
Echinaster (Echinaster)
modestus Perrier, 1881
Echinaster (Echinaster)
sepositus (Retzius, 1783)
Echinaster (Othilia)
brasiliensis Müller & Troschel,
1842
Echinaster (Othilia)
echinophorus (Lamarck, 1816)
Echinaster (Othilia) guyanensis
A. M. Clark, 1987
Echinaster (Othilia) sentus
(Say, 1825)
Echinaster (Othilia)
serpentarius Müller &
Troschel, 1842
Depth
(m)
614
Table A.2 (continued)
Echinaster (Othilia) spinulosus
Verrill, 1869
Henricia antillarum (Perrier,
1881)
Henricia downeyae A.
M. Clark, 1987
Henricia obesa (Sladen, 1889)
Henricia sanguinolenta (O.
F. Müller, 1776)
Henricia sexradiata (Perrier,
1881)
Henricia studeri (Perrier, 1891)
Order Forcipulatida
Family Asteriidae
Anasterias antarctica (Lütken,
1857)
Anasterias minuta Perrier, 1875
Anasterias pedicellaris
(Koehler, 1923)
Anasterias spirabilis (Bell,
1881)
Asterias forbesi (Desor, 1848)
Coscinasterias linearis (Perrier,
1881)
Coscinasterias tenuispina
(Lamarck, 1816)
Cosmasterias lurida (Philippi,
1858)
Diplasterias brandti (Bell,
1881)
Lethasterias australis Fisher,
1923
Lysasterias perrieri (Studer,
1885)
Marthasterias glacialis
(Linnaeus, 1758)
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
1–238
cr,mb,sb,rb
x
192–
1390
342–
1037
22–210
0–200
sb,rub
x
sb,rub
x
29–366
sb,rub
74–430
sb,rub
x
x
1–183
rub,rb
x
x
0–100
0–120
x
x
x
34–54
x
x
x
x
x
x
x
x
x
x
x
x
sb,rub
x
x
x
x
x
0–619
366
rb,rub
rb
x
x
x
x
0–165
rb,sb,rub
x
x
x
0–650
mb,sb
x
x
0–450
sb,mb,rub
x
x
81–155
x
x
0–320
x
0–180
Appendix
Table A.2 (continued)
rb
x
x
615
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
96–760
mb,sb
x
x
80–600
x
384–607
x
x
x
x
33–2300 cr,sg
x
x
50–700
cr,rb
x
x
5–200
sb,rub
256–
sb
3089
338–466 mb
x
x
x
x
x
x
x
x
x
147–380
x
0–2707
x
365–735 mb,sb
345–914 mb,rb,sb
x
430–613 mb
220–
3000
x
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Neosmilaster steineni (Studer,
1885)
Perissasterias polyacantha H.
L. Clark, 1923
Psalidaster mordax Fisher,
1940
Sclerasterias contorta (Perrier,
1881)
Sclerasterias tanneri (Verrill,
1895)
Stephanasterias albula
(Stimpson, 1853)
Coronaster briareus (Verrill,
1882)
Family Heliasteridae
Labidiaster radiosus Lütken,
1871
Family Pedicellasteridae
Ampheraster alaminos
Downey, 1971
Pedicellaster pourtalesii
Perrier, 1881
Family Stichasteridae
Allostichaster hartti (Rathbun,
1879)
Smilasterias triremis Sladen,
1889
Family Zoroasteridae
Cnemidaster sigsbeei (Perrier,
1894)
Doraster constellatus Downey,
1970
Mammaster sigsbeei (Perrier,
1880)
Zoroaster fulgens Thomson,
1873
Habitat
616
Table A.2 (continued)
Order Brisingida
Family Brisingidae
Brisinga costata Verrill, 1884
Hymenodiscus coronata
(G. O. Sars, 1872)
Hymenodiscus verticillata
(Sladen, 1889)
Midgardia xandaros Downey,
1972
Novodinia americana (Verrill,
1880)
Novodinia antillensis
(A. H. Clark, 1934)
Novodinia pandina (Sladen,
1889)
Stegnobrisinga splendens
H. L. Clark, 1926
Family Freyellidae
Colpaster scutigerula Sladen,
1889
Freyastera mexicana
(A. H. Clark, 1939)
Freyastera tuberculata (Sladen,
1889)
Freyella elegans (Verrill, 1884)
Freyella microspina Verrill,
1894
Class Ophiuroidea
Order Euryalida
Family Asteronychidae
Asteronyx loveni Müller &
Troschel, 1842
Astrodia tenuispina (Verrill,
1884)
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
630–
1903
100–
2904
640
cr,mb,rb
x
x
x
rb
x
x
x
408–576 mb
366–
2700
54–990
x
mb,sb
x
rb
x
402–933
930–
2790
2683–
5110
3360–
5620
2928–
3294
1848
x
mb
366–460 sb
x
mb
265–
cr,rb,sb
2499
512–935
x
x
x
rb
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
x
x
x
x
x
x
617
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
677–
1555
108–783 cr
x
x
42–708
x
x
238–475 cr,sb
x
x
42–539
x
cr,rb,sb
cr
x
124–521 cr
x
x
x
x
x
x
66–180
x
15–600
x
1061–
cr
2228
227–576 cr
x
15–397
x
cr,rb,sb,rub
x
x
x
x
x
x
320–475
x
x
x
x
x
70–521
cr
x
0–25
cr
x
0–300
20–677
cr,rb,sb
x
20–180
cr
x
x
x
x
x
x
(continued)
Appendix
Family Asteroschematidae
Asteroschema arenosum
Lyman, 1878
Asteroschema brachiatum
Lyman, 1879
Asteroschema elongatum
Koehler, 1914
Asteroschema intectum Lyman,
1878
Asteroschema laeve (Lyman,
1875)
Asteroschema oligactes (Pallas,
1788)
Asteroschema tenue Lyman,
1875
Ophiocreas lumbricus Lyman,
1869
Ophiocreas oedipus Lyman,
1879
Ophiocreas spinulosus Lyman,
1883
Family Gorgonocephalidae
Asteroporpa annulata Lütken,
1856
Asteroporpa pulchra H.
L. Clark, 1915
Astracme mucronatus (Lyman,
1869)
Astrocaneum herrerai (H.
L. Clark, 1918)
Astrochele lymani Verrill, 1878
Astrocyclus caecilia (Lütken,
1856)
Astrocnida isidis (Duchassaing,
1850)
Habitat
618
Table A.2 (continued)
Astrogomphus vallatus Lyman,
1869
Astropartus mediterraneus
(Risso, 1826)
Astrophyton muricatum
(Lamarck, 1816)
Astrotoma agassizii Lyman,
1875
Gorgonocephalus chilensis
(Philippi, 1858)
Schizostella bifurcata
A. H. Clark, 1952
Order Ophiurida
Family Ophiomyxidae
Ophioblenna antillensis
Lütken, 1859
Ophiobrachion uncinatus
Lyman, 1883
Ophiobyrsa perrieri Lyman,
1883
Ophiobyrsa serpens Lyman,
1883
Astrogeron supinus (Lyman,
1883)
Ophioleptoplax brasiliana
Tommasi & Abreu, 1974
Ophiomyxa brevicauda Verrill,
1899
Ophiomyxa flaccida (Say,
1825)
Ophiomyxa pentagona
(Lamarck, 1816)
Ophiomyxa stimpsonii (Lyman,
1875)
Ophiomyxa tumida Lyman,
1883
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
60–800
cr,rb
x
32–265
mb,rb,sb
0–508
cr,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
74–1000
0–500
x
12–46
cr
1–24
cr,rb
457
mb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
cr,rb
x
x
x
530–
1143
15–520
x
x
23–360
mb
0–1500
cr,rb,sb,sg
0–1060
mb,rb,sb,sg,rub
x
x
cr,mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
108–472 mb
23–601
x
x
x
527
51–126
x
Appendix
Table A.2 (continued)
x
x
x
x
x
619
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–183
x
x
x
265–510
x
x
130–540
x
342–549 rb, mb
x
50–2727
x
x
166
x
sb
x
x
127–278 cr,mb
100–
2900
1–145
x
x
mb
x
x
x
x
200–550 mb
15–600
mb,rb,sb
1.5–100
sb
0.9–3.4
sb
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Ophiomyxa vivipara Studer,
1876
Ophiophrixus quadrispinosus
(Koehler, 1914)
Ophioprium cervicornis
(Lyman, 1883)
Ophiosciasma attenuatum
Lyman, 1878
Ophioscolex disacanthus
H. L. Clark, 1915
Ophioscolex glacialis Müller &
Troschel, 1842
Ophioscolex nutrix (Mortensen,
1936)
Ophioscolex serratus
H. L. Clark, 1900
Ophiosyzygus disacanthus H.
L. Clark, 1911
Family Amphilepididae
Amphilepis norvegica
(Ljungman, 1865)
Amphilepis sanmatiensis
Bernasconi & D’Agostino,
1975
Amphilepis teodorae Tommasi
& Abreu, 1974
Family Amphiuridae
Amphilimna mirabilis
(H. L. Clark, 1941)
Amphilimna olivacea (Lyman,
1869)
Amphiodia atra (Stimpson,
1852)
Amphiodia guillermosoberoni
Caso, 1979
Habitat
620
Table A.2 (continued)
Depth
(m)
Amphiodia habilis
Albuquerque, Campos-Creasey
& Guille, 2001
Amphiodia planispina (von
Martens, 1867)
Amphiodia pulchella (Lyman,
1869)
Amphiodia trychna H. L. Clark,
1918
Amphiodia violacea (Lütken,
1856)
Amphioplus (Amphioplus)
abditus (Verrill, 1871)
Amphioplus (Amphioplus)
brasiliensis Tommasi, 1970
Amphioplus (Amphioplus)
coniortodes H. L. Clark, 1918
Amphioplus (Amphioplus)
sepultus Hendler, 1995
Amphioplus (Amphioplus)
thrombodes H. L. Clark, 1918
Amphioplus (Unioplus) incisus
(Lyman, 1883)
Amphioplus albidus
(Ljungman, 1867)
Amphioplus daleus (Lyman,
1879)
Amphioplus lucyae Tommasi,
1971
Amphioplus mathildae
Tommasi & Abreu, 1974
Amphioplus peregrinator
(Koehler, 1912)
Amphioplus tumidus (Lyman,
1878)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
34
x
0–578
mb,sb,sg,rub
x
x
x
x
x
x
x
0–370
cr,mb,rb,sb,sg
x
x
x
x
x
x
x
x
1–160
m,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
6–587
mb,sb
x
x
0–1207
cr,mb,rb,sg
x
0–82
cr,mb,rb,sg
x
0.3–0.6
mb,sg
1639
1–500
x
x
x
x
x
x
mb,rb,sb
x
x
x
x
5–600
sb
x
0–120
x
x
x
70–578
mb
x
621
(continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–63
mb,sb
x
x
1–311
cr,mb,rb,sg,rub
x
x
0–1962
cr,m,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–1330
x
50
x
x
x
x
x
x
0–400
x
x
x
x
x
x
1–600
x
x
5–1200
0–810
0–60
x
mb,sb
x
x
x
x
x
x
x
(continued)
Appendix
Amphipholis gracillima
(Stimpson, 1852)
Amphipholis januarii
Ljungman, 1867
Amphipholis pachybactera
H. L. Clark, 1918
Amphipholis squamata (Delle
Chiaje, 1828)
Amphipholis subtilis
(Ljungman, 1867)
Amphiura (Amphiura) callida
Albuquerque, Campos-Creasey
& Guille, 2001
Amphiura (Amphiura) correcta
Koehler, 1907
Amphiura (Amphiura)
grandisquama Lyman, 1869
Amphiura (Amphiura)
magellanica Ljungman, 1867
Amphiura (Amphiura) rosea
Tommasi & Oliveira, 1976
Amphiura (Ophionema)
intricata Lütken, 1869
Amphiura algida Koehler, 1911
Amphiura belgicae Koehler,
1900
Amphiura chiajei Forbes, 1843
Amphiura complanata
Ljungman, 1867
Amphiura crassipes Ljungman,
1867
Amphiura deichmanni
Tommasi, 1965
Amphiura diducta Koehler,
1914
Depth
(m)
622
Table A.2 (continued)
Amphiura eugeniae Ljungman,
1867
Amphiura fibulata Koehler,
1913
Amphiura flexuosa Ljungman,
1867
Amphiura iraciae Tommasi &
Oliveira, 1976
Amphiura joubini Koehler,
1912
Amphiura kinbergi Ljungman,
1872
Amphiura latispina Ljungman,
1867
Amphiura muelleri MarktannerTurneretscher, 1887
Amphiura otteri Ljungman,
1872
Amphiura palmeri Lyman,
1882
Amphiura princeps Koehler,
1907
Amphiura rathbuni Koehler,
1914
Amphiura scabriuscula
(Lütken, 1859)
Amphiura semiermis Lyman,
1869
Amphiura stimpsonii Lütken,
1859
Microphiopholis atra
(Stimpson, 1852)
Microphiopholis gracillima
(Stimpson, 1854)
Microphiopholis subtilis
(Ljungman, 1867)
Depth
(m)
Habitat
0–800
mb
2–810
cr,rb,sb,sg,rub
0–50
rub
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
x
x
5–3834
rub
x
3–300
x
10–50
x
134–600
x
198–
3200
5–479
mb,sb
0–107
mb,sb
29–502
mb
cr,rb
x
x
x
x
0–26
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
82–1448 mb
0–2844
cr,rb,sb,rub
1–38
mb
0–26
mb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
623
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
15–117
x
7–48
x
1100
x
0–70
cr,rb,sg
1–12
cr,m,sb,mb
x
x
x
x
x
x
x
0–112
1–36
x
m,mb,sg
0–3
x
x
x
x
0–50
x
x
1–2
rb
x
0.5–33
cr,m,mb,sg,rub
x
1–311
cr,mb,sb
x
0.3–116
cr,m,mb,sb
x
2–11
cr,sb
x
0–223
cr,mb,sg,sb,rub
x
x
0.5–99
cr,rb
x
x
x
x
x
128–148
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Nudamphiura carvalhoi
Tommasi, 1965
Ophiocnida loveni (Ljungman,
1867)
Ophiocnida scabra Lyman,
1879
Ophiocnida scabriuscula
(Lütken, 1859)
Ophionephthys limicola
Lütken, 1869
Ophiophragmus chilensis
(Müller & Troschel, 1843)
Ophiophragmus cubanus
(A. H. Clark, 1917)
Ophiophragmus filograneus
(Lyman, 1875)
Ophiophragmus luetkeni
(Ljungman, 1872)
Ophiophragmus moorei
Thomas, 1965
Ophiophragmus pulcher
H. L. Clark, 1918
Ophiophragmus riisei (Lütken,
1859)
Ophiophragmus septus
(Lütken, 1859)
Ophiophragmus wurdemani
(Lyman, 1860)
Ophiostigma isocanthum (Say,
1825)
Ophiostigma siva Hendler,
1995
Family Hemieuryalidae
Hemieuryale pustulata Von
Martens, 1867
Habitat
624
Table A.2 (continued)
Depth
(m)
Ophiochondrella squamosa
(Lyman, 1883)
Ophiochondrus convolutus
Lyman, 1869
Ophiochondrus crassispinus
Lyman, 1883
Ophiochondrus gracilis Verrill,
1899
Ophiochondrus stelliger
Lyman, 1879
Sigsbeia conifera Koehler,
1914
Sigsbeia murrhina Lyman,
1878
Family Ophiacanthidae
Subfamily Ophiacanthinae
Ophiacantha abyssicola
G. O. Sars, 1871
Ophiacantha affinis Koehler,
1914
Ophiacantha aristata Koehler,
1895
Ophiacantha aspera Lyman,
1878
Ophiacantha bidentata
(Retzius, 1805)
Ophiacantha brasiliensis
Tommasi & Abreu, 1974
Ophiacantha cervicornis
Lyman, 1883
Ophiacantha crassidens Verrill,
1885
Ophiacantha densispina
Mortensen, 1936
Ophiacantha echinulata
Lyman, 1878
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
604
x
165–720 cr
x
x
420
x
198–468
x
x
144–
1080
4–450
mb
x
x
16–706
cr,mb
x
x
35–351
mb,rub
x
x
x
x
x
x
366–420
x
822–
1700
320
x
x
32–4730 cr,mb,rb
x
x
x
145–380
x
366–539
980–
3120
Appendix
Table A.2 (continued)
x
mb
x
x
304–
1958
mb,rb,sb
x
x
x
625
(continued)
Depth
(m)
Ophiacantha ensifera (Verrill,
1899)
Ophiacantha mesembria
H. L. Clark, 1915
Ophiacantha metallacta
H. L. Clark, 1915
Ophiacantha pentacrinus
Lütken, 1869
Ophiacantha rosea Lyman,
1878
Ophiacantha scutata Lyman,
1878
Ophiacantha sertata (Lyman,
1869)
Ophiacantha stellata Lyman,
1875
Ophiacantha setosa (Retzius,
1805)
Ophiacantha vivipara
Ljungman, 1870
Ophiacanthella troscheli
(Lyman, 1878)
Ophialcaea nuttingii Verrill,
1899
Ophiotreta lineolata (Lyman,
1883)
Ophiotreta mixta (Lyman,
1878)
Ophiotreta sertata (Lyman,
1869)
Ophiotreta valenciennesi
valenciennesi (Lyman, 1879)
Ophiotreta valenciennesi
rufescens (Koehler, 1896)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
198–468
x
252–
mb
1125
295–763 mb
x
x
x
320–
1249
402–
mb,rb
1538
521–549
x
183–618 cr,rb
x
x
x
x
x
x
183
x
40–1480 sb
x
0–1097
x
185–238 cr,rb,sb
x
x
x
x
366
x
347–521
x
338–442
x
293–567 mb
200–
1440
155–
1440
626
Table A.2 (continued)
x
x
x
mb
x
Appendix
(continued)
Depth
(m)
Subfamily Ophiohelinae
Ophiomyces frutectosus
Lyman, 1869
Ophiomyces mirabilis Lyman,
1869
Subfamily Ophioplinthacinae
Ophiocamax austera Verrill,
1899
Ophiocamax fasciculata
Lyman, 1883
Ophiocamax hystrix Lyman,
1878
Ophiomitra ornata Verrill,
1899
Ophiomitra robusta Koehler,
1914
Ophiomitra valida Lyman,
1869
Ophiomitrella clavigera
(Ljungman, 1865)
Ophiomitrella conferta
(Koehler, 1922)
Ophiomitrella cordifera
Koehler, 1909
Ophiomitrella glabra (H.
L. Clark, 1901)
Ophiomitrella ingrata Koehler,
1908
Ophiomitrella laevipellis
(Lyman, 1883)
Ophioplinthaca carduus
(Lyman, 1878)
Ophioplinthaca chelys
(Wyville-Thomson, 1878)
Ophioplinthaca dipsacos
(Lyman, 1878)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
151–410 mb
270–560 rb
Appendix
Table A.2 (continued)
x
x
198–
1045
208–979 mb,rb,sb
x
x
171–695 mb
x
x
x
x
x
x
x
201
x
395
x
131–608 cr
x
x
x
250–
1500
x
x
x
x
cr
x
x
x
155–507 cr
796
795–
3305
777
mb
x
x
x
x
x
x
627
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
439
x
0–63
x
147–830 cr
x
490–
1236
mb
x
0–520
cr,mb,rb,sb,rub
125–
2870
1125–
1281
125–
4000
0–24
cr,mb
x
x
x
x
x
x
x
x
x
x
x
mb,rb
cr
x
x
x
x
0–310
x
x
x
30–1765 rb
x
1.5–163
x
62
x
x
198
mb
0–600
rb
14–67
sb
x
x
x
x
x
x
(continued)
Appendix
Ophiothamnus exigua (Lyman,
1878)
Subfamily Ophiotominae
Ophiolimna littoralis Koehler,
1913
Ophiopristis hirsuta (Lyman,
1875)
Ophiotoma gracilis (Koehler,
1914)
Family Ophiactidae
Hemipholis cordifera Bosc,
1802
Histampica duplicata (Lyman,
1875)
Histampica rugosa (H.
L. Clark, 1941)
Ophiactis abyssicola (M. Sars,
1861)
Ophiactis algicola H. L. Clark,
1933
Ophiactis asperula (Philippi,
1858)
Ophiactis balli (W. Thompson,
1840)
Ophiactis brasiliensis Manso,
1988
Ophiactis dispar Verrill, 1899
Ophiactis ljungmani
Marktanner-Turneretscher,
1887
Ophiactis loricata Lyman,
1869
Ophiactis lymani Ljungman,
1872
Ophiactis muelleri Lütken,
1856
Habitat
628
Table A.2 (continued)
Ophiactis quinqueradia
Ljungman, 1871
Ophiactis rubropoda
Singletary, 1973
Ophiactis savignyi (Müller &
Troschel, 1842)
Ophiactis virens (M. Sars,
1857)
Family Ophiochitonidae
Ophiochiton ternispinus
Lyman, 1883
Ophioplax clarimundae
Tommasi, 1970
Ophioplax ljungmani Lyman,
1875
Ophioplax pardalis H. L. Clark,
1941
Ophioplax spinulifera H.
L. Clark, 1941
Family Ophiocomidae
Ophiocoma echinata (Lamarck,
1816)
Ophiocoma paucigranulata
Devaney, 1974
Ophiocoma pumila Lütken,
1859
Ophiocoma riisei Lütken, 1859
Ophiocoma wendtii Müller &
Troschel, 1842
Ophiocomella ophiactoides (H.
L. Clark, 1901)
Ophiocomella sexradia
(Duncan, 1887)
Ophiocomina nigra
(Abildgaard, in O.F. Müller,
1789)
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–640
cr,mb,rb,sb,sg
x
2–32
cr
x
0–1500
cr,m,rb,sb,sg,rub
x
0–90
rb
377–
3550
mb,rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
22–471
cr
x
x
347
x
274–420
x
0–183
cr,m,rb,sb,sg,rub
x
x
x
x
0–455
cr,rb
x
x
0–368
cr,rb,sg
x
x
x
1–2
0–384
cr
cr,m,rb,sg,rub
x
x
x
0–70
cr,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–400
sb,rub
x
629
(continued)
Habitat
0–185
sg,rub
29–320
18–110
cr
1–183
cr,rb,sb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
x
x
41–3000
0–366
4–15
cr,m,rb,sg
cr,rb
x
x
276–
3150
mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
320–475
20–200
x
rb
x
444–759
x
cr,sb
0–580
cr,rb,sg,rub
x
x
x
x
x
x
x
x
0–600
x
x
x
x
x
x
x
x
x
x
x
0–64
cr,rb,sb,sg
x
x
1–223
cr,m,mb,rb,sb,sg,rub x
x
0–1719
cr,m,sb,sg
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Ophiopsila aranea Forbes,
1843
Ophiopsila fulva Lyman, 1878
Ophiopsila guineensis Koehler,
1914
Ophiopsila hartmeyeri Koehler,
1913
Ophiopsila maculata (Verrill,
1899)
Ophiopsila riisei Lütken, 1859
Ophiopsila vittata H. L. Clark,
1918
Family Ophiodermatidae
Bathypectinura heros (Lyman,
1879)
Ophiorachnella angulata
(Lyman, 1883)
Ophiarachnella petersi
(Lyman, 1878)
Ophioconis forbesi (Heller,
1863)
Ophioconis miliaria Lyman,
1878
Ophioderma anitae Hotchkiss,
1982
Ophioderma appressa (Say,
1825)
Ophioderma besnardi
Tommasi, 1970
Ophioderma brevicaudum
Lütken, 1856
Ophioderma brevispinum (Say,
1825)
Ophioderma cinereum Müller
& Troschel, 1842
Depth
(m)
630
Table A.2 (continued)
Depth
(m)
Ophioderma divae Tommasi,
1971
Ophioderma ensiferum Hendler
& Miller, 1984
Ophioderma guttatum Lütken,
1859
Ophioderma januarii Lütken,
1856
Ophioderma longicaudum
(Retzius, 1805)
Ophioderma phoenium H.
L. Clark, 1918
Ophioderma rubicundum
Lütken, 1856
Ophioderma squamosissimum
Lütken, 1856
Ophiopaepale goesiana
Ljungman, 1872
Ophiurochaeta littoralis
(Koehler, 1913)
Family Ophionereididae
Ophionereis dolabriformis John
& A. M. Clark, 1954
Ophionereis olivacea H.
L. Clark, 1901
Ophionereis reticulata (Say,
1825)
Ophionereis sexradia
Mortensen, 1936
Ophionereis squamulosa
Koehler, 1913
Ophionereis vittata Hendler,
1995
Family Ophiotrichidae
Ophiothela danae Verrill, 1869
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
10–30
cr,rb
x
x
x
Appendix
Table A.2 (continued)
x
0–30
cr,rb,rub
x
0–1500
cr
x
x
0–200
sg,rub
1–14
cr,rb,sb
x
x
x
0–360
cr,sg,rb,sb,rub
x
x
x
3–85
cr,rb
x
x
x
x
x
x
x
x
cr,sb
10–93
cr,mb
0–500
cr,m,sb
x
x
0–1500
cr,m,rb,sb,sg,rub
x
x
x
x
x
101–436 cr,mb,sb
1–110
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
18–130
x
0.5–110
cr,sb,rb,sg,rub
x
x
x
10–126
cr
x
x
x
x
x
x
x
x
x
631
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
0.5–540
cr,m,mb,rb,sb,sg,rub x
x
x
x
x
x
x
x
x
x
x
x
1–6
cr,rb,rub
0–10
cr,sg
0–1250
rb,sg,rub
0–74
cr,rb
50–500
rb,sb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
114–410
0–31
x
cr,sb,mb,sg,rb,rub
x
x
183
41
x
x
x
x
x
x
x
x
x
x
x
x
rb
x
8–600
x
0–10
0–1000
cr
cr,m,mb,rb,sb
0–5
rb
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Ophiotrichoides lymani
Ludwig, 1882
Ophiothrix (Ophiothrix) ailsae
Tommasi, 1970
Ophiothrix (Ophiothrix)
trindadensis Tommasi, 1970
Ophiothrix angulata (Say,
1825)
Ophiothrix angulata violacea
(Müller & Troschel, 1842)
Ophiothrix brachyactis
H. L. Clark, 1915
Ophiothrix cimar Hendler,
2005
Ophiothrix fragilis
(Abildgaard, 1789)
Ophiothrix lineata Lyman,
1860
Ophiothrix luetkeni WyvilleThomson, 1873
Ophiothrix maculata
Ljungman, 1872
Ophiothrix oerstedii Lütken,
1856
Ophiothrix pallida Ljungman,
1871
Ophiothrix platyactis
H. L. Clark, 1939
Ophiothrix rathbuni Ludwig,
1882
Ophiothrix stri Hendler, 2005
Ophiothrix suensonii Lütken,
1856
Ophiothrix synoecina Schoppe,
1996
Habitat
632
Table A.2 (continued)
Family Ophiuridae
Subfamily Ophiolepidinae
Amphipholizona delicata H.
L. Clark, 1915
Ophiolepis ailsae Hendler &
Turner, 1987
Ophiolepis elegans Lütkens,
1859
Ophiolepis gemma Hendler &
Turner, 1987
Ophiolepis impressa Lütken,
1859
Ophiolepis kieri Hendler, 1979
Ophiolepis paucispina (Say,
1825)
Ophiolepis pawsoni Hendler,
1988
Ophiolipus agassizii Lyman,
1878
Ophiomidas dubius (Lyman,
1878)
Ophiomusium acuferum
Lyman, 1875
Ophiomusium anaelisae
Tommasi & Abreu, 1974
Ophiomusium constrictum
Mortensen, 1936
Ophiomusium eburneum
Lyman, 1869
Ophiomusium leptobrachium
H. L. Clark, 1941
Ophiomusium lütkeni Lyman,
1878
Ophiomusium lymani Thomson,
1873
Depth
(m)
Habitat
15–600
mb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
156–353
x
1–329
cr,m,mb,rb,sb,sg,rub x
2–139
cr,rb,sb
x
x
0–1500
cr,rb,sb,sg,rub
x
x
2–8
0–37
sg
cr,m,rb,sb,sg,rub
x
x
24.4
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
146–310 rb
x
158–272
76–575
Appendix
Table A.2 (continued)
x
cr,mb,rb
x
x
x
180–260
x
x
x
35–3477 cr,mb,sb
x
1249–
sb
1518
190–278 rb,sb
x
62–4700 mb,rb,sb,rub
x
x
x
x
x
x
x
x
x
633
(continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
347–530
x
1125
x
368–
4062
475
rb,sb
x
x
520–
rb,sb
3103
356–521
x
60–914
cr,rb,sb
x
108–
2732
0–180
cr,mb
x
x
x
x
x
x
rb,sb,rub
262–
rb,sb
4024
512–713
x
x
x
x
x
x
x
376–746
x
144–540 cr
x
x
x
158–272
x
x
1006–
1098
484–
1385
2115
x
rb,sb
x
x
x
x
x
(continued)
Appendix
Ophiomusium microporum H.
L. Clark, 1941
Ophiomusium moniliforme H.
L. Clark, 1941
Ophiomusium planum Lyman,
1879
Ophiomusium sculptum Verrill,
1899
Ophiomusium serratum Lyman,
1878
Ophiomusium stellatum Verrill,
1899
Ophiomusium testudo Lyman,
1875
Ophiomusium validum
Ljungman, 1872
Ophioplocus januarii (Lütken,
1856)
Ophiosphalma armigerum
(Lyman, 1878)
Ophiosphalma dyscritum
H. L. Clark, 1941
Ophiosphalma monoplax
H. L. Clark, 1915
Ophiothyreus goesii Ljungman,
1872
Ophiozonella clypeata (Lyman,
1883)
Ophiozonella falklandica
Mortensen, 1936
Ophiozonella granulifera
H. L. Clark, 1941
Ophiozonella marmorea
Lyman, 1883
Ophiozonella molesta (Koehler,
1904)
Habitat
634
Table A.2 (continued)
Depth
(m)
Ophiozonella nivea (Lyman,
1875)
Ophiozonella tessellata
(Lyman, 1879)
Subfamily Ophioleucinae
Ophiostriatus striatus
(Mortensen, 1933)
Ophiernus adspersus adspersus
Lyman, 1883
Ophioleuce depressum (Lyman,
1869)
Ophiopyren longispinus
Lyman, 1878
Ophiostriatus atlanticus
(Mortensen, 1933)
Subfamily Ophiurinae
Amphiophiura metabula
H. L. Clark, 1915
Amphiophiura oedignatha
H. L. Clark, 1915
Amphiophiura ornata (Lyman,
1878)
Anthophiura ingolfi Fasmer,
1930
Ophiambix devaneyi Paterson,
1985
Ophiocten amitinum Lyman,
1878
Ophiomastus meridionalis
(Lyman, 1879)
Ophiomastus satelitae
Tommasi & Abreu, 1974
Ophiomastus secundus Lyman,
1878
Ophiomisidium pulchellum
(Wyville-Thomson, 1878)
Habitat
70–1249 rb,sb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
320–475
270–
3500
68–3650 mb,rb
x
x
x
x
x
585
x
329
x
296–396 mb
274–
mb,sb
1830
284–944 mb
Appendix
Table A.2 (continued)
x
x
x
x
x
x
x
x
21–1200 cr
x
146–494 mb
x
46–545
x
x
x
x
115–600
108–
mb,sb
2035
11–3061 cr,rb,sb
x
x
x
x
x
x
x
635
(continued)
Depth
(m)
880–
mb
1636
250–808
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
1786
x
338–366
x
54–3384
x
x
x
x
20–1900 cr,mb,rb,sb
x
x
x
x
x
400
x
x
40–1260 mb,sb
x
x
73–1037 mb
20–842
mb,sb
10–350
600–
4315
mb,sb,rub
mb,sb
x
x
x
x
x
x
x
46–6398 mb,rb,sb
0–463
x
x
x
x
x
x
(continued)
Appendix
Ophiomisidium speciosum
Koehler, 1914
Ophiomisidium tommasii
Borges, Monteiro & Amaral,
2006
Ophiopleura inermis (Lyman,
1878)
Ophioplinthus abyssorum
(Lyman, 1883)
Ophioplinthus inornata
(Lyman, 1878)
Ophioplinthus martensi
(Studer, 1885)
Ophiophycis mirabilis Koehler,
1901
Ophiura acervata (Lyman,
1869)
Ophiura carinifera (Koehler,
1901)
Ophiura carnea Lütken, 1858
Ophiura clemens (Koehler,
1904)
Ophiura falcifera (Lyman,
1869)
Ophiura fallax Cherbonnier,
1959
Ophiura grubei Heller, 1863
Ophiura (Ophiuroglypha)
irrorata concreta (Koehler,
1901)
Ophiura (Ophiuroglypha)
irrorata irrorata (Lyman,
1878)
Ophiura ljungmani (Lyman,
1878)
Ophiura lymani (Ljungman,
1871)
Habitat
636
Table A.2 (continued)
Depth
(m)
Ophiura ophiura (Linnaeus,
0–300
1758)
Ophiura sarsi (Lütken, 1855)
82–827
Ophiura scomba Paterson, 1985 358–
3943
Ophiura tenera (Lyman, 1883) 158–500
Ophiuraster perissus H.L.
Clark, 1939
Class Echinoidea
Order Cidaroida
Family Cidaridae
Austrocidaris canaliculata
1–424
(A. Agassiz, 1863)
Austrocidaris lorioli
160–
(Mortensen, 1903)
1081
Austrocidaris spinulosa
124–641
Mortensen, 1910
Calocidaris micans
33–624
(Mortensen, 1903)
Cidaris abyssicola (A. Agassiz, 225–375
1869)
Cidaris blakei (A. Agassiz,
270–720
1878)
Cidaris cidaris (Linnaeus,
50–2000
1758)
Cidaris rugosa (H. L. Clark,
40–540
1907)
Eucidaris tribuloides
0–1500
tribuloides (Lamarck, 1816)
Histocidaris nuttingi
225–740
Mortensen, 1926
Histocidaris purpurata
300–
(Wyville-Thomson, 1872)
1800
Histocidaris sharreri
200 –
(A. Agassiz, 1880)
740
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
mb,sb
mb
rb,sb
x
Appendix
Table A.2 (continued)
x
x
mb
x
x
x
x
x
x
mb,rb,sb
x
mb,rb,sb
x
x
x
mb,rb
x
x
mb
x
x
mb,rb,sb
x
x
cr,rb,sb,sg,rub
x
x
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
rb,sb
x
x
637
(continued)
Depth
(m)
300–
mb,rb,sb
1745
22–1000 cr,mb,rb,sb,rub
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
20–1717 cr,mb,rb,sb,rub
x
x
48–1089 mb,rb,sb
x
x
640–782 rb,sb
x
x
130–
mb,sb
1020
16–1180 mb,rb,sb
x
x
360–
mb,sb
2545
200–
mb,sb
3700
50–3700 mb,sb
x
200–
1800
300–
2300
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
mb
x
x
x
x
170–720 cr,mb,rb
x
x
x
x
x
180–925
x
x
684–
3111
mb,rb,sb
x
5–89
cr,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Stereocidaris ingolfiana
Mortensen, 1903
Stylocidaris affinis (Philippi,
1845)
Stylocidaris lineata Mortensen,
1910
Tretocidaris bartletti
(A. Agassiz, 1880)
Family Rhabdicidaridae
Porocidaris purpurata
(Wyville-Thomson, 1872)
Order Echinothurioida
Family Echinothuriidae
Araeosoma belli Mortensen,
1903
Araeosoma fenestratum
(Thomson, 1872)
Calveriosoma hystrix (WyvilleThomson, 1872)
Hygrosoma petersii
(A. Agassiz, 1880)
Phormosoma placenta placenta
Thomson, 1872
Phormosoma placenta sigsbei
A. Agassiz, 1880
Sperosoma grimaldii Koehler,
1897
Order Diadematoida
Family Aspidodiadematidae
Aspidodiadema jacobyi
A. Agassiz, 1880
Aspidodiadema tonsum
A. Agassiz, 1879
Plesiodiadema antillarum
(A. Agassiz, 1880)
Family Diadematidae
Astropyga magnifica
A. H. Clark, 1934
Habitat
638
Table A.2 (continued)
Depth
(m)
Astropyga nuptialis Tommasi,
1958
Centrostephanus besnardi
Bernasconi, 1955
Centrostephanus longispinus
(Philippi, 1845)
Centrostephanus longispinus
rubicingulus H. L. Clark, 1921
Diadema antillarum antillarum
(Philippi, 1845)
Diadema ascensionis
Mortensen, 1909
Order Pedinoida
Family Pedinidae
Caenopedina cubensis
A. Agassiz, 1869
Order Salenioida
Family Saleniidae
Salenia goesiana Lovén, 1874
Salenocidaris profundi
(Duncan, 1877)
Salenocidaris varispina
A. Agassiz, 1869
Order Arbacioida
Family Arbaciidae
Arbacia dufresnii (Blainville,
1825)
Arbacia lixula (Linnaeus, 1758)
Arbacia punctulata (Lamarck,
1816)
Coelopleurus floridanus
A. Agassiz, 1871
Habrocidaris scutata
(A. Agassiz, 1880)
Podocidaris sculpta
A. Agassiz, 1869
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
1–20
x
Appendix
Table A.2 (continued)
x
10–1000 cr,mb,rb,sb,rub
33–842
sb,rub
0–800
cr,m,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
1–400
x
x
x
x
x
x
x
x
220–
1200
mb,rb,sb
x
90–842
200–
3700
250–
3015
rb
mb,rb,sb
x
mb,rb,sb
x
0–340
rb,rub
0–50
0–225
rb,sb
cr,rb,sb,sg,rub
60–2380 cr,mb,rb,sb
920–
1400
8–780
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
cr,mb,rb,sb
x
x
639
(continued)
Habitat
30–340
rb,rub
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
1–5
0–80
x
x
rb,sg
x
x
12–500
cr,mb,rb,sb
x
x
1–361
rb,rub
70–720
cr,rb,sb
x
x
22–357
rb,sb
x
x
40–777
mb,rb,sb
x
0–250
rb,sb,rub,sg
0–580
cr,mb,rb,sb,sg,rub
x
x
3–256
cr,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
69–155
x
x
x
x
x
x
x
x
x
x
x
3–100
rb
0–35
cr,mb,rb,sb
x
x
(continued)
Appendix
Order Camarodonta
Family Parechinidae
Loxechinus albus (Molina,
1782)
Paracentrotus gaimardi
(Blainville, 1825)
Paracentrotus lividus
(Lamarck, 1816)
Psammechinus
microtuberculatus (Blainville,
1825)
Order Temnopleuroida
Family Temnopleuridae
Genocidaris maculata
A. Agassiz, 1869
Pseudechinus magellanicus
(Philippi, 1857)
Trigonocidaris albida
A. Agassiz, 1869
Family Toxopneustidae
Lytechinus callipeplus
H. L. Clark, 1912
Lytechinus euerces
H. L. Clark, 1912
Lytechinus variegatus carolinus
A. Agassiz, 1863
Lytechinus variegatus
variegatus (Lamarck, 1816)
Lytechinus williamsi Chesher,
1968
Pseudoboletia occidentalis
H. L. Clark, 1921
Sphaerechinus granularis
(Lamarck, 1816)
Toxopneustes pileolus
(Lamarck, 1816)
Depth
(m)
640
Table A.2 (continued)
Tripneustes ventricosus
(Lamarck, 1816)
Order Echinoida
Family Echinidae
Echinus esculentus Linneaeus,
1758
Echinus tylodes H. L. Clark,
1912
Gracilechinus gracilis
(A. Agassiz, 1869)
Sterechinus agassizii
Mortensen, 1910
Sterechinus neumayeri
(Meissner, 1900)
Family Echinometridae
Echinometra lucunter lucunter
(Linnaeus, 1758)
Echinometra viridis A. Agassiz,
1863
Order Holectypoida
Family Echinoneidae
Echinoneus cyclostomus Leske,
1778
Order Clypeasteroida
Family Clypeasteridae
Clypeaster aloysioi (Brito,
1959)
Clypeaster chesheri Serafy,
1970
Clypeaster cyclopilus
H. L. Clark, 1941
Clypeaster euclastus
H. L. Clark, 1941
Clypeaster lamprus
H. L. Clark, 1914
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–842
cr,rb,sb,sg,rub
x
10–289
rb
x
350–760 sb
x
69–457
x
mb,rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
24–470
x
13–250
x
x
0–45
cr,rb,sg,rub
x
x
x
x
0–40
cr,m,rb,sb,sg,rub
x
x
x
0.5–585
cr,mb,rb,sb,sg,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
18–101
cr,mb,sb
x
x
69–460
sb
36–530
cr,mb,sb
x
78–450
cr,mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
641
(continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
9–55
sb
50
15–75
sb
sb
x
20–278
mb,sb
x
0–285
sb,sg
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
1–210
mb,rb,sb,sg,rub
x
99–2500 mb,sb
x
170–
2286
0–1250
x
sb,rub
13–32
sb
0–60
mb,sb
0–240
sb,sg
x
0–180
sb,sg,rub
x
0–180
mb,sb,rub
x
0–10
sb
0–3
sb,rub
sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Clypeaster luetkeni Mortensen,
1948
Clypeaster oliveirai Krau, 1952
Clypeaster prostratus Ravenel,
1848
Clypeaster ravenelii
A. Agassiz, 1869
Clypeaster rosaceus (Linnaeus,
1758)
Clypeaster speciosus Verrill,
1870
Clypeaster subdepressus (Gray,
1825)
Family Echinocyamidae
Echinocyamus grandiporus
Mortensen, 1907
Echinocyamus macrostomus
Mortensen, 1907
Echinocyamus pusillus (Müller,
1776)
Family Mellitidae
Encope aberrans Martens,
1867
Encope emarginata (Leske,
1778)
Encope michelini L. Agassiz,
1841
Leodia sexiesperforata (Leske,
1778)
Mellita quinquiesperforata
(Leske, 1778)
Mellita quinquiesperforata
latiambulacra (H. L. Clark,
1840)
Mellita tenuis H. L. Clark, 1940
Order Cassiduloida
Family Cassidulidae
Depth
(m)
642
Table A.2 (continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0.5–197
sb
x
x
x
x
3.5
x
1–45
69–155
x
x
130–800 mb,rb,sb
x
37–366
x
sb
x
x
x
x
x
450–
5850
x
22–1610 mb
x
x
x
x
640
x
181–329 mb,sb
x
18–641
x
mb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
37–3200 mb
3–270
mb,sb
0–250
55–720
mb,rb,sb,rub
mb
x
0.3–293
cr,mb,sb,rb,sg,rub
x
148–330
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
643
Cassidulus caribaearum
Lamarck, 1801
Cassidulus infidus Mortensen,
1948
Cassidulus mitis Krau, 1954
Eurhodia relicta Mooi, 1990
Family Echinolampadidae
Conolampas sigsbei
(A. Agassiz, 1878)
Echinolampas depressa Gray,
1851
Order Holasteroida
Family Pourtalesiidae
Pourtalesia miranda
A. Agassiz, 1869
Order Spatangoida
Family Asterostomatidae
Archaeopneustes hystrix
(A. Agassiz, 1880)
Family Brissidae
Anabrissus damesi (A. Agassiz,
1881)
Brissopsis alta Mortensen,
1907
Brissopsis atlantica Mortensen,
1907
Brissopsis atlantica
mediterranea Mortensen, 1913
Brissopsis elongata elongata
Mortensen, 1907
Brissus unicolor (Leske, 1778)
Linopneustes longispinus
(A. Agassiz, 1878)
Meoma ventricosa ventricosa
(Lamarck, 1816)
Neopneustes micrasteroides
(A. Agassiz, 1878)
Depth
(m)
Appendix
Table A.2 (continued)
(continued)
Habitat
150–
1025
69–805
mb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
cr,mb,rb,sb
x
x
x
x
x
350–436
x
90–525
cr,mb
1–210
sb,sg
x
x
567–570 mb,sb
x
117–619 mb,sb
x
25–200
mb,sb
0–10
sb
380–
4833
mb,sb
0–230
sb
350–
3550
910
mb,sb
71–225
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
50–113
75–970
1–676
x
mb,sb
x
x
x
x
x
x
x
(continued)
Appendix
Palaeobrissus hilgardi
A. Agassiz, 1880
Paleopneustes cristatus
A. Agassiz, 1873
Paleopneustes josephinae
Lovén, 1872
Paleopneustes tholoformis
Chesher, 1968
Plagiobrissus grandis (Gmelin,
1788)
Plethotaenia angularis
Chesher, 1968
Plethotaenia spatangoides
(A. Agassiz, 1883)
Rhabdobrissus costae (Gasco,
1876)
Rhynobrissus cuneus Cooke,
1957
Family Hemiasteridae
Holanthus expergitus (Lovén,
1874)
Family Loveniidae
Echinocardium cordatum
(Pennant, 1777)
Family Maretiidae
Homolampas fragilis
(A. Agassiz, 1869)
Homolampas lovenioides
Mortensen, 1948
Family Prenasteridae
Tripylus excavatus Philippi,
1845
Family Schizasteridae
Abatus agassizii (Pfeffer, 1889)
Abatus cavernosus (Philippi,
1845)
Abatus philippii Lovén, 1871
Depth
(m)
644
Table A.2 (continued)
Aceste bellidifera Thomson,
1877
Agassizia excentrica
A. Agassiz, 1869
Brisaster fragilis (Düben &
Koren, 1846)
Brisaster moseleyi (A. Agassiz,
1881)
Hypselaster limicolus
(A. Agassiz, 1878)
Moira atropos clotho Michelin,
1855
Ova canaliferus (Lamarck,
1816)
Tripylaster philippii (Gray,
1851)
Schizaster doederleini
(Chesher, 1972)
Schizaster floridiensis
(Chesher, 1972)
Schizaster orbignyanus
A. Agassiz, 1880
Family Spatangidae
Spatangus purpureus (Müller,
1776)
Class Holothuroidea
Order Dendrochirotida
Family Cucumariidae
Aslia lefevrei (Barrois, 1882)
Cladodactyla crocea (Lesson,
1830)
surinamensis (Semper,
1868)
Cucumaria manoelina
Tommasi, 1971
Depth
(m)
Habitat
116–
5220
27–900
sb
cr,mb,sb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
Appendix
Table A.2 (continued)
x
401–
1100
27–340
x
mb,sb
x
0–445
mb,rb,sb
x
9–100
mb,sb
x
x
x
x
x
x
x
x
x
13–595
x
9–220
mb,sb
12–65
sb
22–500
mb,sb
15–900
mb,sb,rub
6–25
0–4300
rb
mb,rb,sb,rub
0–5
rb,sg
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
645
(continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–50
cr,rb,sb,sg
x
x
1.5–183
rb,sb
x
x
x
x
2–160
mb,sb
x
0–379
sb,rub
x
0–400
rb
x
17–37
mb,sb,rub
0–5
rb
x
x
x
0–2
0–72
0–1874
x
cr,rb,sb,rub
rb,sg
x
x
x
x
69–155
4–130
x
x
x
x
x
rb
x
x
x
mb,rb,sb
135–189
0–197
mb,rb,sb
x
271–290
31.1–
40.2
0–64
x
x
x
0.6–1.7
0–340
x
x
x
x
x
x
x
x
x
cr,rb,sb,mb,sg
x
x
x
(continued)
Appendix
Duasmodactyla seguroensis
(Deichmann, 1930)
Euthyonacta solida
(Deichmann, 1930)
Hemioedema spectabilis
(Ludwig, 1882)
Heterocucumis godeffroyi
(Semper, 1868)
Heterocucumis steineni
(Ludwig, 1898)
Leptopentacta deichmannae
Domantay, 1958
Neocnus incubans Cherbonnier,
1972
Ocnus braziliensis (Verrill,
1868)
Ocnus pygmaeus (Théel, 1886)
Ocnus suspectus (Ludwig,
1874)
Paracolochirus mysticus
(Deichmann, 1930)
Pawsonia saxicola (Brady &
Robertson, 1871)
Pentacta pygmaea (Théel,
1886)
Pseudocnus dubiosus leoninus
(Semper, 1868)
Pseudocnus cornutus
(Cherbonnier, 1941)
Pseudocnus perrieri (Ekman,
1927)
Pseudrotasfer microincubator
Bohn, 2007
Stereoderma unisemita
(Stimpson, 1851)
Thyonella gemmata (Pourtalès,
1851)
Depth
(m)
646
Table A.2 (continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
9–67
rb,sb
x
2–30
sb
x
x
x
x
x
x
0–30
0–180
x
mb,rb,sb
90–159
0.5–60
x
x
x
x
0–3
sg
0–111
sb,rub
0–60
mb
0–99
cr,rb,sg
x
x
0–38
mb,rb,sb,sg
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
6–366
sg
5–615
mb,sb,rub
x
x
x
x
x
6–51
sb
0.5–46
rb,sb,sg,rub
51–170
mb
x
x
x
x
x
x
x
x
x
x
x
647
Thyonella pervicax (Théel,
1886)
Thyonella sabanillaensis
(Deichmann, 1930)
Trachythyone crassipeda
Cherbonnier, 1961
Trachythyone lechleri
(Lampert, 1885)
Trachythyone parva (Ludwig,
1874)
Trachythyone peruana
(Semper, 1868)
Family Phyllophoridae
Havelockia inermis (Heller,
1868)
Neothyonidium parvum
(Ludwig, 1881)
Pentamera chiloensis (Ludwig,
1887)
Pentamera pulcherrima Ayres,
1854
Phyllophorus (Urodemella)
occidentalis (Ludwig, 1875)
Stolus cognatus (Lampert,
1885)
Thyone cognata (Lampert,
1885)
Thyone deichmannae Madsen,
1941
Thyone fusus (Müller, 1776)
Thyone montoucheti Tommasi,
1971
Thyone pawsoni Tommasi,
1972
Thyone pseudofusus
Deichmann, 1930
Thyone tanyspeira Pawson &
Miller, 1988
Family Psolidae
Depth
(m)
Appendix
Table A.2 (continued)
(continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
1–17
cr,rb
x
x
x
0–2
x
x
0–20
rb
x
x
0–483
rb
x
x
35–1080 rb
x
95
0–430
rb
rb
7–207
rb,rub
x
73–243
x
x
x
x
x
x
x
x
x
0–7
mb,rb,sb,sg
0–100
rb
x
0–37
mb,rb,sb,sg
x
15
x
x
x
x
x
x
x
x
x
x
x
x
x
x
43–723
x
50–1000
x
(continued)
Appendix
Lissothuria antillensis Pawson,
1967
Lissothuria braziliensis (Théel,
1886)
Neopsolidium convergens
(Herouard, 1901)
Psolidium dorsipes Ludwig,
1886
Psolus antarcticus Philippi,
1857
Psolus marcusi Tommasi, 1971
Psolus patagonicus Ekman,
1925
Psolus squamatus (Koren,
1844)
Psolus tuberculosus Théel,
1886
Psolus victoriae Tommasi,
1971
Family Sclerodactylidae
Euthyonidiella dentata
Cherbonnier, 1961
Euthyonidiella destichada
(Deichmann, 1930)
Euthyonidiella trita (Sluiter,
1910)
Pseudothyone belli (Ludwig,
1886)
Sclerodactyla briareus
(Lesueur, 1824)
Order Dactylochirotida
Family Ypsilothuriidae
Echinocucumis asperrima
(Théel, 1886)
Echinocucumis hispida (Barret,
1856)
Depth
(m)
648
Table A.2 (continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
270–
2684
mb
0–54
cr,rb,sg,rub
x
x
0–8
cr,mb,rb,sb
x
x
0–457
mb,sb
x
3–370
cr,mb,sb,rub
x
x
0–10
cr,m,rb,sb,sg,rub
x
x
0–30
cr,mb,rb,sb,sg
x
x
0–183
cr,m,mb,rb,sb,sg
x
x
0–15
mb,sb
x
15
sb,rub
x
1–77
rb
x
0–30
mb,sb,sg
x
0–8
cr,m,mb,rb,sb,rub
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–30
rb
x
3–193
rb
x
0–30
rb
x
0–250
mb,sb,sg
x
(continued)
649
Ypsilothuria talismani
talismani Perrier, 1886
Order Aspidochirotida
Family Holothuriidae
Actinopyga agassizii (Selenka,
1867)
Holothuria (Cystipus) cubana
Ludwig, 1875
Holothuria (Cystipus)
occidentalis Ludwig, 1875
Holothuria (Cystipus)
pseudofossor Deichmann, 1930
Holothuria (Halodeima)
floridana Pourtalès, 1851
Holothuria (Halodeima) grisea
Selenka, 1867
Holothuria (Halodeima)
mexicana Ludwig, 1875
Holothuria (Holothuria)
dakarensis Panning, 1939
Holothuria (Holothuria) helleri
Marenzeller, 1878
Holothuria (Holothuria)
mammata Grube, 1840
Holothuria (Holothuria)
tubulosa Gmelin, 1788
Holothuria (Platyperona)
parvula (Selenka, 1867)
Holothuria (Platyperona) rowei
Pawson & Gust, 1981
Holothuria (Platyperona)
sanctori Delle Chiaje, 1823
Holothuria (Panningothuria)
forskali Delle Chiaje, 1823
Holothuria (Roweothuria)
arguinensis Koehler & Vaney,
1906
Holothuria (Roweothuria) polii
Delle Chiaje, 1823
Depth
(m)
Appendix
Table A.2 (continued)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0–42
cr,m,mb,rb,sb,sg,rub x
0–42
mb,rb,sb
x
0–402
cr,mb,rb,sb,sg
x
0–121
cr,m,mb,sb,sg
x
x
0–27.4
cr,mb,rb,sb,sg
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
0–30
cr
x
x
20–466
mb,rb,sb
x
69–466
cr,mb
1–162
cr,rb,sb,sg
36
sb
5–800
mb,sb,sg,rub
0–274
cr,mb,rb,sb,sg
x
x
0–18
rb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
20–1960 mb,sb
439–900 mb
x
x
x
x
x
x
x
x
x
(continued)
Appendix
Holothuria (Semperothuria)
surinamensis Ludwig, 1875
Holothuria (Selenkothuria)
glaberrima Selenka, 1867
Holothuria (Theelothuria)
princeps Selenka, 1867
Holothuria (Thymiosycia)
arenicola Semper, 1868
Holothuria (Thymiosycia)
impatiens (Forskaal, 1775)
Holothuria (Thymiosycia)
rathbunii Lampert, 1885
Holothuria (Thymiosycia)
thomasi Pawson & Caycedo,
1980
Holothuria (Vaneyothuria)
lentiginosa von Marenzeller,
1892
Holothuria (Vaneyothuria)
lentiginosa enodis Miller &
Pawson, 1979
Family Stichopodidae
Astichopus multifidus (Sluiter,
1910)
Eostichopus arnesoni Cutress
& Miller, 1982
Eostichopus regalis (Cuvier,
1817)
Isostichopus badionotus
(Selenka, 1867)
Isostichopus macroparentheses
(H. L. Clark, 1922)
Parastichopus tremulus
(Gunnerus, 1767)
Family Synallactidae
Amphigymnas bahamensis
Deichman, 1930
Depth
(m)
650
Table A.2 (continued)
Depth
(m)
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
402–585 mb
210–
cr,mb
1600
134–
mb
2919
548–903 cr,sb
x
x
x
x
x
x
x
x
232–
mb
2180
1975–
2515
180–720 mb
x
x
x
18–1445 mb
x
484–
5100
914–
5062
151–
3890
699–
4060
1353–
5690
1100–
4060
232
x
mb
x
x
mb
mb
x
mb
x
x
x
x
x
x
x
x
x
x
mb,sb
3411–
3459
x
x
x
1000–
3476
mb
x
x
651
Bathyplotes bigelowi
Deichmann, 1940
Bathyplotes natans (M. Sars,
1868) Östergren, 1896
Bathyplotes pourtalesi (Théel,
1886)
Hansenothuria benti Miller &
Pawson, 1989
Meseres atlanticus (R. Perrier,
1902)
Meseres occultatus (von
Marenzeller, 1893)
Mesothuria connectens
(Perrier, 1898)
Mesothuria gargantua
Deichmann, 1930
Mesothuria intestinalis
(Ascanius, 1805)
Mesothuria (Zygothuria) lactea
(Théel, 1886)
Mesothuria maroccana Perrier,
1902
Mesothuria (Mesothuria)
rugosa Herouard, 1912
Mesothuria (Penichrothuria)
verrilli (Théel, 1886)
Molpadiodemas depressus
(Herouard, 1902)
Paelopatides gigantea (Verrill,
1884)
Pseudostichopus occultatus
Merenzeller, 1893
Pseudostichopus peripatus
(Sluiter, 1901)
Scotothuria herringi Hansen,
1978
Synallactes crucifera Perrier,
1898
Habitat
Appendix
Table A.2 (continued)
(continued)
Order Elasipodida
Family Deimatidae
Deima validum validum Théel,
1879
Orphnurgus asper Théel, 1882
Family Elpidiidae
Peniagone purpurea (Théel,
1882)
Family Pelagothuridae
Enypniastes eximia Théel, 1882
Family Psychropotidae
Benthodytes lingua Perrier,
1896
Benthodytes sanguinolenta
Théel, 1882
Benthodytes typica Théel, 1882
Habitat
914–
2780
600–
1049
mb
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
x
x
x
mb
x
2800–
3000
x
461–689
540–
2200
914–
3100
315–
5046
x
mb
mb
x
x
mb
x
x
x
x
x
x
x
x
955–
4060
2210–
5000
mb
x
x
x
x
1143–
5600
3545–
3635
0–990
x
mb
mb,sb,rub
x
x
x
x
x
(continued)
Appendix
Benthodytes valdiviae Hansen,
1975
Psychropotes depressa (Théel,
1882)
Psychropotes longicauda
Théel, 1882
Psychropotes scotiae (Vaney,
1908)
Psychropotes semperiana
Théel, 1882
Order Molpadiida
Family Caudinidae
Acaudina molpadioides
(Semper, 1867)
Paracaudina chilensis chilensis
(Müller, 1850)
Family Molpadiidae
Depth
(m)
652
Table A.2 (continued)
Depth
(m)
Molpadia barbouri Deichman,
1940
Molpadia blakei (Théel, 1886)
Molpadia cubana Deichmann,
1940
Molpadia eltaninae Pawson,
1977
Molpadia musculus Risso, 1826
Molpadia oolitica (Pourtalès,
1851)
Molpadia parva (Théel, 1886)
Order Apodida
Family Chiridotidae
Chiridota marenzelleri R.
Perrier, 1904
Chiridota rotifera (Pourtalès,
1851)
Taeniogyrus contortus
(Ludwig, 1875)
Trochodota purpurea (Lesson,
1830)
Family Synaptidae
Anapta fallax Lampert, 1889
Epitomapta roseola (Verrill,
1874)
Eupatinapta acanthia (A.
H. Clark, 1899)
Euapta lappa (Müller, 1850)
Leptosynapta brasiliensis
Freire & Grohmann, 1989
Leptosynapta imswe Pawson,
1976
Leptosynapta inhaerens
(Müller, 1776)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
440–
mb
1529
3482–
mb
3518
24–1464 mb
x
x
x
x
x
x
x
20–855
x
x
x
x
x
x
125–
2844
mb
x
40–159
mb,rub
0–360
cr,rb,sb,sg
82–200
mb,sb
x
x
0–100
sb,rub
x
x
0–350
1.5–40
sb,rub
rb,sb
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
cr,rb,sb,sg
x
x
x
x
x
x
x
x
x
x
0–2
0–173
x
x
10.7
0–256
1–4
x
x
35–5205 mb
42–1440 mb
x
Appendix
Table A.2 (continued)
x
mb
x
653
(continued)
Leptosynapta multigranula H.
L. Clark, 1924
Leptosynapta nannoplax
Pawson, 1976
Leptosynapta parvipatina H.
L. Clark, 1924
Leptosynapta roseogradia
Pawson, 1976
Leptosynapta tenuis (Ayres,
1851)
Protankyra benedeni (Ludwig,
1881)
Protankyra brychia (Verrill,
1885)
Protankyra ramiurna Heding,
1928
Synaptula hydriformis
(Lesueur, 1824)
Synaptula secreta AnconaLopez, 1957
Depth
(m)
Habitat
MEX BEL GUA HON NIC CRC PAN COL VENc VENa BRA URG ARG MAV CUB HAI RDO PRI CAN
0.5–2.9
sb
x
654
Table A.2 (continued)
x
x
9–12
x
0.5
x
0.5–167
x
x
5–100
x
x
x
0–18
mb,rb,sb,sg
x
0–10
cr,m,sb,sg
x
1–10
x
x
x
x
x
x
x
x
x
x
x
x
Habitat classification: cr Coral Reefs, m mangroves, mb muddy bottom, rb rocky bottom, sb sandy bottom, sg seagrass, rub rubble bottom. Countries abbreviation: MEX Mexico, BEL Belize, GUA
Guatemala, HON Honduras, NIC Nicaragua, CRC Costa Rica, PAN Panama, COL Colombia, VENc Venezuela Caribbean, VENa Venezuela Atlantic, BRA Brazil, URG Uruguay, ARG Argentina, MAV
Malvinas Islands, CUB Cuba, HAI Haiti, RDO Dominican Republic, PRI Puerto Rico, CAN Canary Islands
Appendix
Index
A
Abatus, 375, 379
Abundance, 6, 18, 28, 30, 372
Acanthasther planci, 28
Albatross, 15, 70, 110, 132, 136, 157, 189,
365, 406, 442
Allan Hancock, 15, 71, 111, 190
Allometric scaling, 202, 205
Allostichaster, 367, 369, 382
Amphiura, 26, 28, 167, 235, 247, 319–321,
368, 374
Amplaster, 353
Anasterias, 367, 369, 371, 382, 384, 385
Anchialine, 27
Ancon, 278
Aquaculture, 3, 6, 48, 99, 132, 173
Arbacia, 19, 25, 71, 82, 131, 240, 243, 256,
288, 289, 309, 326, 368, 375, 388, 451,
494, 497, 501Astropecten articulatus
Archipelago, 92, 108, 149
Argentina, 2, 4, 346, 365
Argentine Continental Shelf, 360
Argentine Province, 367
Argentine Sea, 360, 364, 366
Astropecten, 18, 23, 25, 26, 36, 39, 71, 93, 98,
167, 172, 173, 192, 241, 242, 263, 265,
291, 315–317, 324, 369, 373, 381, 458,
492, 495
Astropecten articulatus, 93, 265, 413, 458
Astropecten duplicates, 265, 413, 458
Athyonidium chilensis, 285, 294, 297
Atlantis, 408, 414
Azores, 479, 481
Aztecs, 4, 39, 40
B
Barren, 483–485, 504
Belize, 87, 88
Bernasconi, Irene, 5, 360
Biodiversity, 12, 24, 45, 145, 191
Bioerosion, 30, 88
Biomass, 33, 43, 172, 208
Bocas del Toro, 18, 70, 73, 108
Boulder, 84, 169, 199
Brazil, 2, 6, 126, 305
Brazilian Province, 351, 496
Brazilian Current, 345, 351
C
Cabo Verde, 481, 482, 495
Caenocentrotus gibbosus, 195, 206, 289
Cahuita, 81
Canary Islands, 6
Caño Island, 88
CARICOMP, 81, 82
Caso, Ma. Elena, 5, 6, 16
Central America, 80
Centrostephanus coronatus, 19, 29, 170, 195,
206, 259, 285
Cherbonier, 4, 5, 442
Chile, 4, 187, 281, 387
Clypeaster rosaceus, 82, 122, 255, 256,
413, 416
Cóbanos, Los, 84
Cocos Island, 88
Coelomocyte types, 208
Colombian Museum of Marine Natural History
(MHNMC), 156
J. J. Alvarado and F. A. Solís-Marín (eds.), Echinoderm Research and Diversity
in Latin America, DOI: 10.1007/978-3-642-20051-9,
Ó Springer-Verlag Berlin Heidelberg 2013
655
656
Cosmasterias, 369, 373, 381, 382, 388
Conservation, 45, 46, 75, 221
Coral Reefs, 6, 25, 77, 110
Costa Rica, 68, 71, 79, 93, 515
Culebra Island, 120, 439, 458
Ctenantedon kinziei, 75, 154, 161
D
Deep azooxanthellate coral
communities, 164, 167
Diadema antillarum, 26, 32, 80, 93, 113, 126,
136, 145, 154, 162, 173, 240, 262, 409,
413, 414, 442, 446, 448, 460, 464,
483, 530
Diadema aff. antillarum, 82
Diadema mexicanum, 29, 31, 33, 83, 159, 192,
195, 215, 528
Disease, 34, 134, 210, 262
Distribution, 6, 18, 32
Diversity, 18
Dominican Republic, 6
E
Earthquake, 262
Echinometra lucunter, 26, 32, 79, 83, 90, 93,
113, 122, 162, 240, 256, 308, 413, 444,
459, 464, 465
Echinometra vanbrunti, 19, 24, 29, 31, 36, 39,
83, 84, 93, 159, 170, 172, 183, 191,
195, 206, 208, 223, 289, 528
Echinometra viridis, 80, 87, 88, 113, 154, 240,
444, 528
Ecoregion, 37, 38
Eco-regionalization, 416
Ecozone, 13, 416–418, 421
Ecuador, 132, 184, 186, 203
El Hierro, 474, 475, 501
El Niño, 88
El Salvador, 2, 67, 71, 84, 93
Encope grandis, 26, 128
Encope micropora, 24, 183, 205, 206, 212, 291
Endemism, 26, 38, 92, 284, 515
Equatorial front, 185
Eucidaris thouarsii, 29, 31, 84, 170, 191,
289, 528
Eucidaris galapagensis, 183, 192, 195, 200,
201, 213–215, 221, 223
Eucidaris tribuloides, 25, 26, 80, 93, 154, 254,
324, 411, 461, 465
Eulimids, 214
Euapta lappa, 26, 413, 416, 462, 530
Extinction, 44, 125, 221
Index
F
Ferrol, 278
Fisheries, 6
Fission, 197–199, 369, 371
Florida Museum of Natural
History (FLMNH)
Fossil, 80
French Guiana, 4, 156, 516
G
Galapagos, 18
Glycosides, 382
Gorgona, 131, 49, 153, 157
Gran Canaria, 475, 482
Gulf of California, 13, 15, 18
Gulf of Chiriqui, 109, 110, 117, 130
Gulf of Fonseca, 68, 71, 84
Gulf of Mexico, 32
Gulf of Nicoya, 68, 71, 80
Gulf of Panama, 69, 109, 112, 120, 131
Guyana, 4, 156, 442
H
Haiti, 6, 87
Heavy metals, 96, 263, 413
Heliaster helianthus, 285, 288, 384
Hispabiota Marina Project, 431
Hispaniola, 2, 427, 440
Holothuria, 15, 19, 24–26, 31, 35, 36, 44, 82,
84, 86, 90, 93, 97, 119, 131, 132, 154,
155, 168, 170–172, 175, 191, 192, 198,
199, 210, 217, 218, 242, 289, 291, 294,
309, 322, 323
Holothuria atra, 154, 195, 199
Holothuria glaberrima, 173, 245, 450, 451
Holothuria theeli, 86, 191, 198, 199,
217, 294
Holothuria (Thymiosycia) thomasi
Honduras, 80, 82, 87
Huatulco, 154, 462
Hydro-physical stress, 413
I
Immune response, 208, 223, 310, 455
INVEMAR, 148, 153, 164
Isostichopus badionotus, 44, 86, 119, 132, 172,
242–246, 324
Isostichopus fuscus, 19, 24, 31, 33, 35, 36, 42,
93, 97, 172, 192, 196–199, 210, 217,
219, 223, 292, 297
Isthmus, 68, 123, 129, 533
Index
J
Johnson-Smithsonian Deep-Sea
Expedition, 435
L
La Gomera, 474, 475
La Niña, 208
La Parguera, 440, 447, 449
Larval abundance, 444, 447, 463
Larval biology, 198
Leptopentacta deichmannae, 235, 247
Loxechinus albus, 288, 294, 296, 2978, 375,
377, 378
Lytechinus variegatus, 25, 32, 80, 86, 113,
154, 164, 234, 235, 240, 244, 258, 309,
411, 414, 444, 445, 461, 464
Lytechinus semituberculatus, 192, 195, 201,
202, 216, 288
Luidia, 18, 23, 25, 26, 32, 71, 98, 172, 173,
194, 207, 243, 263, 288, 291, 305, 316,
324, 360, 369, 373, 382, 384, 492,
493, 528
Luidia senegalensis, 413, 415, 305
M
Macaronesian, 481, 483
Madeira, 125, 481, 483
Magellanic Province, 364
Magueyes, 443, 448
Malpelo, 149, 153, 157, 160
Malvinas (Falkland) Current, 303
Mangrove, 12, 77, 119
Marine Biodiversity Information System
(SIBM), 147, 156
Marine Science Collection of Department of
Marine Sciences (MSC), 443
Mexico, 6, 8
Molecular evolution, 455
Monophoraster, 352, 353
Mortality, 87
Museum of Comparative Zoology (MCZ), 430
Museum of Natural Sciences
of Berlin (ZMB), 430
N
National Museum of Natural History of Santo
Domingo (MNHNSD), 430
Nicaragua, 2, 67, 68, 70, 71, 73, 81–83, 86, 93,
97, 98
Nidorellia armata, 172, 192, 291
Norcross-Bartlett, 429
657
O
Ophiocoma, 24, 26, 79, 90
Ophioplocus, 320, 374
Ophiothrix, 348, 492, 494
Oreaster reticulatus, 4, 433, 434, 494
P
Paita, 278, 281, 284, 288
Paleontology, 252, 95
Panama, 2, 5, 6, 38, 67, 68, 70, 73, 79, 81, 82,
87, 92, 93, 98, 107–110, 112–114,
117–120, 122–133, 135–137, 147, 149,
156, 157, 257, 465, 515, 516, 533, 536
Paracas, 278, 280, 288, 289, 294
Parasitism, 210, 214
Paraster doederleini, 82
Parastichopus parvimensis, 33, 35, 46
Parish-Smithsonian, 429
Patagonian, 351, 364, 371, 374, 380–382, 387,
388, 516
Pattalus mollis, 285, 297
Pearl Islands, 109, 112, 128
Peru, 2, 4, 6, 92, 132, 157, 277, 278, 280-284,
288, 289, 291, 292, 294, 296–298,
515–517, 524, 92
Pharia pyramidata, 170, 191
Phataria unifascialis, 28, 29
Phylogenetic, 89, 120, 202
Phylogeography, 126
Physiology, 90, 120, 244, 450
Phytoplankton, 16, 406
Platasterias, 26, 207, 544
Predation, 183, 210, 213
Programa EcoMar, 431
Pseudechinus, 308, 375, 381
Pseudocnus, 288
Psolidium, 292
Puerto Rico, 2, 6, 427, 431, 439, 441–444,
447, 449, 450, 456–462, 464–466
R
Recruitment, 115, 444
Regeneration, 452, 453
Reproduction, 35, 89, 120, 243, 373, 375,
443, 446
Reproductive cycles, 120
Río de la Plata, 345, 346, 350, 351, 360, 362,
364, 515
Rosario Corals, and San Bernardo Islands
National Natural Park (PNNCRSB),
154, 161, 167
RVBlake, 442
658
RV John Elliot Pillsbury, 429, 430
RV Oregon, 73, 112
RV Stranger, 429
S
Salvajes, 481
Samanco, 278, 288
San Andrés and Providencia, 147–149, 164
San Blas, 70, 108, 113, 114, 119, 133, 134, 202
Seagrass, 163
Sechura, 278, 288
Settlement, 444, 446, 448
Speciation, 127
Spawning, 41, 369, 372, 378, 379, 444
Spicule, 86, 219
Stichaster striatus, 297
Stichopus horrens, 198
Surinam, 4
Sustainable production, 4, 156
T
Tayrona National Natural Park, 154, 161
Tenerife, 474, 475, 481, 483, 492, 501, 502
Te Vega, 72
Index
Tethyaster vestitus vestitus, 32, 80, 82
Tetrapygus niger, 4
Thalassia testudinum, 433
Toxopneustes roseus, 114, 129
Trench, 68, 238, 248, 251, 441
Tripneustes depressus, 130, 414, 433
Tripneustes ventricosus, 26, 82, 240, 444
Trophic cascade, 213, 500
U
UNAM, 15, 16, 47, 48, 410
Upwelling, 148, 187, 189, 235, 236, 238, 248,
251, 281, 304
Uruguay, 2, 6, 317, 345–351, 353, 354, 362,
365, 516
Urracá, 71, 112
V
Vieques island, 439, 458
Z
Zaca, 111