_k\ll" MUllo?
Fr.ltll-1\l () P.llldo
MONOGRAPHS IN SYSTEMATIC BOTANY
FROM THE MISSOURI BOTANICAL GARDEN
Volume 83
ISBN 0-915279-92-4
ISSN 0161-1542
AUTHORS' ADDRESS
Real Jardin Botanico
Plaza de Murillo 2
E-28014 Madrid, Spain.
jmunoz@ma-rjb.csic.es
pando@ma-rjb.csic.es
EDITOR
Victoria C. Hollowell
MANAGING EDITOR
Amy Scheuler McPherson
ASSOCIATE EDITOR
Diana Gunter
TEXT FORMATTER
Barbara Mack
SENIOR SECRETARY
Aida Kadunic
The cover and title page illustration is
Grimmia trinervis R. S. Williams, by David Foronda (Madrid).
Copyright © 2000 by Missouri Botanical Garden Press.
P.O. Box 299
St. Louis, Missouri 63166-0299, U.S.A.
All rights reserved.
Printed in U.S.A.
She's anything but typical ...
She's a powerful force ...
She's a natural law ...
She used to look good to me, but now I find her
Simply irresistible
Robert Palmer
Simply Irresistible
From the CD "Heavy Nova"
© EM! [America] 1988, Robert Palmer producer and songwriter
TABLE OF CONTENTS
ABSTRACT ........................................................................................................... .
ACKNOWLEDGEMENTS ........................................................................................ .
INTRODUCTION .................................................................................................... .
MATERIALS AND METHODS ...... ..... .............. ............. .................. ....... .............
2
GRIMMIA HEDW. ......... ......... ........ ......... ..... ....... ........... ................. ......... ..... ....
5
KEY TO THE SPECIES OF GRIMMIA ... ....... ....... ......... ............ ............. ........... .....
6
COUNTRY CODES (Iso
3166) ........ ....... ..... ....... .... ..... ................... ....... ....... .... 13
SYNOPSIS OF THE GENUS GRIMMIA ... ....... ..... ....... ...... ......... ......... .... ....... .... .........
15
TAXA EXCLUDED FROM GRIMMIA ..... ......... ..... ......... ...................... .... ....... ....... ....
85
NAMES WITH TYPE MATERIAL NOT SEEN ...........................................................
97
LITERATURE CITED .............................................................................................. 114
INDEX OF SCIENTIFIC NAMES ...............................................................................
117
ABSTRACT
The genus Grimmia is considered to comprise 71 species. In this synopsis no taxa at infraspecific
rank are recognized. For each accepted species synonyms are listed, mostly based on the study of their
type material. 131 names names are lectotypified, and references to previous correct lectotypifications
are given where appropiate. Also presented is a key and the distribution of each species, based solely
on specimens studied by the authors. Names for which type material could not be studied yet are listed
in a different section with the geographical information of the protologue to facilitate the location of
type specimens.
Some widely used names are shown to be mere synomyms of others, mostly from different
geographical regions. Among those worth mentioning are the following, with the accepted name in
parentheses: G. affinis Homsch. (= G. longirostris Hook.); G. apiculata Homsch. (= G. fuscolutea
Hook.); G. arizonae Renauld & Cardot (= G. pilifera P. Beauv.); G. brittoniae R.S. Williams (= G.
funalis (Hedw.) Schwagr.); G. curvata (Brid.) De Sloover and Dryptodon patens (Hedw.) Brid. (= G.
ramondii (Lam. & DC.) Margad.); G. hermanii Crum (= G. muehlenbeckii Schimp.); G. moxleyi R.S.
Williams (= G. orbicularis Wilson); G. sessitana De Not. (= G. reflexidens Milli. Hal.); G. tenerrima
Renauld & Cardot (= G. reflexidens Mull. HaL); G. tolucensis Cardot (= G. elongata Kaulf.).
ACKNOWLEDGEMENTS
Jesus Muiioz expresses his gratitude to the staff of the Missouri Botanical Garden, and especially
his colleagues at the bryologicallaboratory, for providing a nice working environment during his twoyear stay. Most of the work was done in St. Louis during a post-doctoral stay funded by the Spanish
Ministry of Education and Culture. He is also grateful to the curators of the herbaria cited in the text
for their kind assistance during his visits and for the loans of specimens. The visit to Helsinki (H) was
funded by the European Commission HCM Contract nl ERBCHGECT940065 with the Division of
Systematic Biology of the University of Helsinki. The Friends of the Farlow and the W.C. Steere Fund
supported his visits to Harvard University (FH) and The New York Botanical Garden (NY),
respectively. Travels to Jena (JE) and Paris (PC) during 1999 were funded by the Missouri Botanical
Garden. He is also indebted to Rosa M. Ros and M. Jesus Cano (Murcia, Spain), for inviting him on
several collecting trips that have increased the known geographic range of several species; and to
Marite Telleria (Madrid) for her unconditional support. Steve Churchill (St. Louis) and Carlos Aedo
(Madrid) deserve special mention: they persuaded Munoz to publish parts of this work and have
supported and helped him in many ways.
We thank Marshall R. Crosby (St. Louis), Felix Muiioz (Madrid), Ryszard Ochyra (Krakow),
Marko Sabovljevic (Belgrade), and Alan Whittemore (St. Louis, now Washington, D.C.) for help with
literature or doubts about nomenclatural puzzles; Si He (St. Louis), Masanobu Higuchi (Tokyo), and
Tatyana V. Shulkina (St. Louis) for translations from Chinese, Japanese, and Russian, respectively. We
also thank Olga Afonina (St. Petersburg), Jan-Peter Frahm (Bonn), Harald Kurschner (Berlin), and
Beata Papp (Budapest) for providing specimens from areas little known to the authors.
INTRODUCTION
Grimmia is the largest genus in the Grimmiaceae. It is considered a difficult genus, and its species
are believed to be protean in nature. The combination of a large number of species and the absence of
treatments to name extra-European specimens has hindered many from dealing with the genus in any
depth.
For several years, Grimmia has been the research of one of the authors (JM). What started as a
Ph.D. subject (Grimmia subg. Orthogrimmia, see Munoz, 1998b), soon evolved into a world revision
of the genus. It also soon became evident that the large number of described taxa made further study an
2
MuNoz AND PANDO
impossible task without the aid of an electronic database. The second author (FP) had designed at that
time a database to store names and a program to automatically generate nomenclatural lists. A
collaboration produced a checklist of the genus. The names and associated information (proto logue,
distribution, etc.) was stored in the database system "Bibmaster" (www.rjb.csic.eslbibmastelbibmaste
.htm). The nomenclatural synopsis here presented, along with the name index, was generated
automatically from the system, which also provides support for data consistency, publication
abbreviations, and other tools for data capture and maintenance.
In this synopsis we recognize 71 species in Grimmia. In fact, it represents the nomenclatural
treatment of an ongoing world revision of the genus. Its structure is similar to the nomenclatural
section in the monographs already published (Munoz 1998a, 1998b, 1999a). We also present a key to
the accepted species and the known world distribution for these taxa. In a time when biodiversity is
one of the magic words in biology, we think that presenting authenticated data on the number and
distribution of species in the large moss family Grimmiaceae will be helpful in related fields, e.g.,
bryogeography.
Figure 1 shows the number of species recognized in each area according to the system proposed
by Wijk et al. (1962). As it is clear from this figure, Grimmia has mainly an holarctic distribution, and
it is especially diverse at high latitudes in the Northern Hemisphere. The Southern Hemisphere has a
relatively low Grimmia diversity, although the total number in some areas could lead to a different
conclusion. Larger areas in the Southern Hemisphere are poor in species in fact, and the number only
increase at high elevations or latitudes.
The history and the different infrageneric classifications proposed for Grimmia were reviewed by
Deguchi (1978). The main infrageneric classifications to date were also summarized by Munoz
(1998b: table 1). A new, rather eclectic, infrageneric classification, has been published by Nyholm
(1998) in her last treatment of the Fennoscandian species. As already stated (Munoz, 1998b: 374), the
infrageneric classification of Grimmia has yet to be resolved, and therefore we deliberately do not
embrace any of those proposed systems.
MATERIALS AND METHODS
We compiled the names of Grimmia, Dryptodon, and other related genera accepted as pertaining
to Grimmia in Index Muscorum (Wijk et aI., 1962, 1969), but not those that Index Muscorum gives as
accepted in a different genus (e.g., Grimmia flexuosa Griff., for which the accepted name in Index
Muscorum is Garkea comosa (Dozy & Molk.) Wijk & Margad., was not included in the database).
Afterward, all Grimmia names found in Crosby et al. (1992), Crosby and Magill (1994), and Crosby
and Magill (1997), or published thereafter were included in the nomenclatural database. Hence, the
database does not list all names published in the genus, and Index Muscorum has to be consulted for
those names not appearing here. However, a few taxa originally excluded from Grimmia in Index
Muscorum have been resurrected and included here. The database contains some 1370 names, which
correspond to 792 taxa. The protologue of each name has been checked, involving the review of more
than 500 different papers. Type or original material of 590 taxa representing more than 74% of the
total have been studied so far. Among the taxa for which types have not been studied, there are 65
(8.2%) described at the species level and 139 (17.55%) described at the variety or form level. From a
preliminary survey of these taxa not yet studied, none threatens to be older than any accepted name in
this work, which assures nomenclatural stability.
-Am 1
Am 1
37
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Figure 1. Number of Grimmis species in the geographical areas proposed by Wijk et al. (1962).
ｾ＠ｾ＠
4
MuNoz AND PANDO
The work is divided into three sections:
1. Synopsis of the genus Grimmia. This includes the names for which we have seen the type(s)
or, in a few cases, accurate and informative illustrations of types (e.g. Deguchi, 1978; Geissler &
Maier, 1995; Maier, 1998). In this section accepted names, in bold, are listed alphabetically, and
synonyms, in italic, are listed under the accepted name in chronological order of publication.
Distribution is indicated by citing a single collection from each country.
2. Taxa excluded from Grimmia. Taxa belonging to other genera are included here. The
structure of this section is the same as that of the previous section. Some names belonging in
Racomitrium and Schistidium have been transferred to these genera by other authors, and we
follow their views regarding generic placement, but we have no opinion on the correct species
name, because these combinations are not made in the context of a taxonomic revision.
Consequently, those names are listed alphabetically under Racomitrium spp. and Schistidium
spp. Both genera are in urgent need of taxonomic revision outside Europe, and any decision now
seems premature.
3. Names with type material not seen. In this section we list alphabetically those names for
which we have not seen the type(s). All names have been at some moment included in Grimmia,
or were described as infraspecific entities of taxa now considered to belong in Grimmia. Some, if
not many, of the names here included have been considered synonymous with other taxa by
previous authors (see Wijk et aI., 1962, 1969). We do not present here any information on
previous synonymizations, since many of those have proved to be wrong. This does not mean that
we consider any of them "good" taxa. Surely these taxa will be synonyms, but of different taxa
from which they will use to be considered conspecific. Although apparently no name in this
section threatens any of the accepted names in the first section, we continue to reduce the size of
this section.
Every entry follows the same scheme: name, author(s), place of publication, homotypic
synonyms ordered chronologically, and type locality according to the protologue. The type locality
according to the protologue ("PROTOLOGUE" in the text) is the exact transcription of the locality in
the protologue. It has been taken directly and has not been edited. If further information has been
added, it is placed between square brackets.
We have treated all nomina nuda (nom. nud.), and placed them in the species lists. These names
are invalid arid accordingly have no types. Therefore, we have included the geographical information
on these names under the header "ORIGINAL MATERIAL," and the information on the specimen
labels, under "ORIGINAL SPECIMENS." In a few instances (e.g., G. biplicata, G. meridensis, and G.
paramophila), no original specimen(s) was seen, and our judgment is based on illustrations or
comments on the original publication.
For names in the first two sections, "TYPE" is the transcription of the type (holotype or
lectotype) label, and herbarium(a) where the type(s) is housed. The protologue and label information
are included for verification of our typifications. Bibliographic references to lectotypifications by other
authors have also been indicated where appropriate. We have selected new lectotypes for all names not
typified previously.
Distributions are given only for accepted names. They are based solely on studied specimens.
Some 10,000 specimens have been studied so far, and one representative specimen is listed together
with the herbarium where this voucher is located. We have used ISO-3166 standard for country codes.
WORLD SYNOPSIS OF GRIMMIA
5
The only exception is Russia (ISO code RUS, not used here), which has been divided into three areas
according to Ignatov and Afonina (1992): European Russia (SUN); Siberia (SIB); and Far East (SFE).
Areas in the Near East, Siberia, and Africa are underrepresented, and any specimen from those
areas is welcomed by the fIrst author.
GRIMMIA HEDW.
Sp. Musc. Frond.: 75. 1801.
Type species: Grimmia plagiopoda Hedw. (lectotype, designated by Martensson, 1956: 156).
Dryptodon Brid., Bryol. Univ.: 191. 1826.
Type species: Dryptodon patens (Hedw.) Brid. [= Bryum patens Hedw.] (lectotype, designated
by Ochyra, Sergio & Schumacker, 1988: 246).
Guembelia Hampe, Bot. Zeitung (Berlin) 4: 124. 1846.
Type species: Guembelia elliptica (Funck) Hampe [= Grimmia elliptica Funck] (lectotype,
designated by Pfeiffer, 1874: 1511).
Schistidium Brid., Muscol. Recent. Suppl. 4: 20. 1818 ["1819"]. Anodon Rabenh., Deutschl.
Krypt.-Fl. 2(3): 153. 1848, nom. illeg.
Type species: Anodon pulvinatus (Hedw.) Rabenh. [= Gymnostomum pulvinatum Hedw.]
(designated by Pfeiffer, 1873: 199).
NOTE: Ochyra (1989a) showed that Schistidium pulvinatum (Hedw.) Brid., type of both
Anodon and Schistidium Brid., is in fact Grimmia anodon, which converts those genera to
taxonomic synonyms of Grimmia. As a consequence, he proposed for conservation the name
Schistidium Bruch & Schimp. with a new type (Ochyra, 1989b).
Hydrogrimmia (I. Hagen) Loeske, Stud. Morph. Syst. Laubm.: 108. 1910. Grimmia subgen.
Hydrogrimmia I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1909(5): 35. 1909.
Grimmia subgen. Hygrogrimmia Vilh., Vestn. Knil. Ceske Spolecn. Nauk, n. Mat.-Prir. 1924:
6. 1924, nom. inval., orthogr. err. Grimmia subsect. Hydrogrimmia (I. Hagen) G.N. Jones, in
Grout, Moss Fl. N. Amer. 2: 21. 1933. Grimmia sect. Hydrogrimmia (I. Hagen) Nyholm, Ill. Fl.
Nord. Mosses 4: 334. 1998.
Type species: Hydrogrimmia mollis (Bruch & Schimp.) Loeske (holotype).
Trigonodictyon Dixon & P. de la Yarde, Ann. Cryptog. Exot. 1: 40. 1928.
Type species: Trigonodictyon indicum Dixon & P. de la Yarde (holotype).
Usmania Laz., Dopov. Akad. Nauk Ukrajins'k. RSR 11: 1039. 1970.
Type species: Usmania campylopoda Laz. (holotype)
Cladautoicous, gonioautoicous, or dioicous. Plants in dense cushions or compact to loose tufts,
glaucous, green, greenish yellow or olive-green. Stems erect or ascending, with or without central
strand. Leaves erect, appressed or flexuous, occasionally with homomallous tips when dry, erect to
spreading when moist, linear, ovate, lanceolate, ligulate, oblong to triangular, obtuse to acuminate,
concave, canaliculate or keeled, plane or plicate; margins entire, plane, recurved or incurved; costa
single, percurrent, terete, semiterete, semi-elliptic or almost indistinct in cross section; lamina 1-4stratose in the distal half; distal cells isodiametric to rectangular or oblate, bulging or plane, smooth or
papillose, with straight or sinuous walls; proximal cells quadrate to rectangular or oblate, the walls
straight or sinuous, uniformly thickened or the transverse walls thicker than the longitudinal walls;
with or without hyaline hair-points. Perichaetialleaves convolute and larger or similar in shape but
slightly larger than vegetative leaves. Androecia axillary or terminal. Setae straight or curved, longer
or shorter than capsules. Capsules immersed, emergent or exserted, subglobose, ovoid, ellipsoid or
fusiform, symmetric or asymmetric and ventricose at the base, smooth or ribbed, stramineous or
6
MuNoz AND PANDO
castaneous, with stomata at the urn base or lacking stomata; exothecial cells isodiametric or
rectangular, thin- or thick-walled; annulus simple and persistent, compound and revoluble, or
compound and tardily deciduous; peristome teeth 16, triangular, entire, perforate in the distal half or
cribrose throughout and irregularly cleft at apex, orange to brown; opercula conic to long-rostrate;
calyptrae cucullate or mitrate, smooth, covering at least to the capsule mouth; columella persistent;
spores smooth to coarsely granulose.
Grimmia is a genus of epilithic mosses (except G. pitardii, which grows on ground) closely
related to Schistidium Bruch & Schimp. Indeed, American authors tend to unite both genera (e.g.,
Crum, 1994, who also joined Jaffueliobryum Ther. in his encompassing concept of Grimmia). As I
conceive the genus, it includes the taxa of Grimmiaceae subfam. Grimmioideae in the sense of
Churchill (1981), except Schistidium. It could be argued that this approach renders Grimmia
paraphyletic, but our preliminary results of morphological cladistic analysis support that Schistidium is
sister to the rest of the subfamily. This phylogenetic analysis of Grimmia, including a study of
infra generic relationships, will be presented elsewhere.
Schistidium differs from Grimmia basically in two characters: systylius capsules and small
calyptrae not reaching the capsule mouth. A character widely used, perichaetial leaf size, must be
considered uninformative, because those leaves are also greatly enlarged in most species of Grimmia.
KEY TO THE SPECIES OF GRIMMIA
Notes for using the key (see Munoz, 1999a: 121):
Leaf length is without hair-points.
Cross sections are essential to correctly identify specimens of Grimmiaceae.
Perichaetialleaf area is considered the area of the rectangle defmed by the length and width of
the perichaetialleaves, again without hair-points.
Capsules may be immersed and are sometimes overlooked. Using the dissecting microscope look
carefully for inflated distal stem areas, where leaves have longer hair-points.
1a. Leaves strongly plicate on both sides of costa.
2a. Seta sigmoid to coiled; capsule asymmetric and ventricose at base, seta eccentrically placed at
base, gymnostomous; calyptrae mitrate, not exceeding further down than capsule mouth, not plicate
....................................................................................................................................... 68. G. trinervis
2b. Seta straight; capsule symmetric with seta centrally placed at base, peristome present; calyptrae
cucullate or, if mitrate, campanulate, covering most of the capsule and plicate.
3a. Capsule exserted, calyptra cucullate, not plicate; laminal cells usually papillose ........................ .
.............................................................................................................................. .. 11. G. caespiticia
3b. Capsule immersed, calyptrae mitrate, campanulate, plicate; laminal cells not papillose
...................................................................................................................................... [Coscinodon]
lb. Leaves not or only weakly plicate.
4a. Capsule asymmetric and ventricose at base; seta eccentrically attached to the capsule, sigmoid,
shorter than the capsule.
5a. Proximal marginal cells of leaves hyaline, inflated, with all walls similar, thin .......................... .
....................................................................................................................... 25. G. incrassicapsulis
5b. Proximal marginal cells of leaves not hyaline or inflated, with transverse walls thicker than
longitudinal walls.
6a. Leaves obovate, wider at apex; calyptrae cucullate ............................................. 13. G. crinita
6b. Leaves subulate to lanceolate, wider at proximal half; calyptrae mitrate.
WORLD SYNOPSIS OF GRIMMIA
7
7a. Costa elliptical in cross section, scarcely prominent on dorsal surface, occasionally of the
same thickness as lamina, scarcely differentiated from lamina.
8a. Autoicous, male buds small and hidden just below perichaetia ............... 2. G. americana
8b. Dioicous, male buds terminal ............................................................ 55. G. poecilostoma
7b. Costa semicircular in cross section, prominent on dorsal surface, differentiated from lamina.
9a. Peristome lacking ............................................................................................ 3. G. anodon
9b. Peristome present.
lOa. Leaves muticous ....................................................................................... 53. G. pitardii
lOb. Leaves with hair-point.. ..................................................................... 54. G. plagiopodia
4b. Capsule symmetric; seta centrally attached to the capsule, straight or curved but not sigmoid,
longer than the capsule, or sterile specimens [NOTE: Not all sterile collections can be identified with
confidence] .
11a. Costa cross section rounded ........................................................................... 62. G. teretinervis
lIb. Costa cross section not rounded.
12a. Proximal marginal cells ofleaves oblate; laminal cells not papillose ............ 30. G. laevigata
12b. Proximal marginal cells of leaves quadrate to rectangular or, if oblate, laminal cells strongly
papillose.
13a. Leaves soft, broadly ovate ................................................................................ 39. G. mollis
13b. Leaves rigid, linear, ovate or obovate.
14a. Leaf cells distinctly papillose.
15a. Costa elliptical in cross section, scarcely prominent on dorsal surface, occasionally of
the same thickness as lamina, scarcely differentiated from lamina .............. 36. G. mammosa
15b. Costa semiterete to triangular in cross section, prominent on dorsal surface,
differentiated from lamina.
16a. Leaves longer than 2 mm; plants robust; stems ascending, to 8 cm long; seta curved
........................................................................................................................ 17. G. elatior
16b. Leaves shorter than 1.8 mm; plants small; stems erect, to 1.5 cm long; seta straight.. .
........................................................................................................................ 44. G. nivalis
14b. Leaf cells not papillose, pseudopapillose or not.
17a. Proximal marginal cells of leaves with all walls similar, thin.
18a. Margins plane throughout or incurved above, no trace of recurvation.
19a. Leaves crisped when dry .................................................................... 65. G. tortuosa
19b. Leaves erect and appressed to weakly flexuous when dry, but never crisped.
20a. Ventral layer of costa 4-6 cells wide .............................................. 71. G. wilsonii
20b. Ventral layer of costa 2 cells wide.
21a. Seta straight, to 1 mm; capsule immersed; exothecial cells thick-walled ........... ..
............ ................................................................................................. 67. G. triformis
21b. Seta straight or curved, longer than 1 mm; capsule emergent or exserted;
exothecial cells thin-walled.
22a. Seta curved; stem leaf hyaline hair-point to 2 mm, those of the perichaetial
leaves strongly flexuous and twisted .................................................. 6. G. arenaria
22b. Seta straight; stem leaf hyaline hair-point to 1 mm, those of the perichaetial
leaves weakly flexuous and not twisted .......................................... 16. G. donniana
18b. At least one margin recurved for part of its length.
23a. Leaves crisped when dry, narrowly linear-Ianceolate from ovate base.
24a. One margin narrowly recurved for the proximal 113, the other flat or only
recurved for a very short distance from base ........................................... 26. G. incurva
24b. Both margins recurved to revolute more than half their length ... 43. G. nepalensis
8
MuNoz AND PANDO
23b. Leaves straight to flexuous but not crisped, lanceolate to ovate-Ianceolate, seldom
subulate.
25a. Leaves with an ovate base extended into a subulate distal part mostly filled by the
costa; costa excurrent in a strong green mucro ........................................ 26. G. incurva
25b. Leaves lanceolate to ovate-Ianceolate; lamina evident in distal part of leaves;
costa not excurrent as a green mucro.
26a. Capsules cylindrical, 1.5-2.5 mm long .................................. 35. G. macrotheca
26b. Capsules ovoid, ca. 1 mm long.
27a. Longest hair-point longer than 0.3 mm long, most of the leaves piliferous;
plants autoicous, male buds small and lateral, hidden just below perichaetia ......... ..
........................................................................................................ 20. G.fuscolutea
27b. Longest hair-point to 0.3 mm long, most of the leaves muticous; dioicous,
male buds terminal.
28a. Lamina blades in distal half forming <30 angle in cross section, parallel to
each other at their junction with the costa, furrow very narrow; seta straight ........
................................... ....................... ......................... ............... ..... 18. G. elongata
28b. Lamina blades in distal half forming >45 angle in cross section, not parallel
to each other at their junction with the costa, furrow more open; seta curved ........
........................................................................................................ 60. G. redunca
17b. Proximal marginal cells of leaves with transverse walls thicker than longitudinal walls.
29a. Leaves strongly crisped when dry ........................................................ 64. G. torquata
29b. Leaves straight, flexuous or curled, but not crisped.
30a. Gemmae present at leaf apices.
31a. Gemmae yellow to yellow-green, brownish when old; ridge-like thickenings over
the cell lumen, which appear as papillae in cross section; dorsal side of costa rounded
in cross section, without wings or projections ......................................... 4. G. anomala
31b. Gemmae reddish; laminal cells without ridge-like thickenings over the lumen;
costa usually with wings or projections at dorsal side ......................... 23. G. hartmanii
30b. Gemmae at leaf apices lacking.
32a. Leaves all constantly muticous.
33a. Costa winged on dorsal side ....................................................... 59. G. ramondii
33b. Costa without projections on dorsal side.
34a. Margins plane, no trace of recurvation.
35a. Leaves homomallous; apex subulate ..................................... 21. G. hamulosa
35b. Leaves straight; apex obtuse to acute.
36a. Plants brownish; distal part of lamina 2-stratose .............. 46. G. obtusifolia
36b. Plants green to black; distal part oflamina 2-5-stratose ...... 70. G. unicolor
34b. Margins recurved partially to fully on one or both sides.
37a. Lamina I-stratose, occasionally 1-2 marginal rows 2-stratose.
38a. Leaves lanceolate; apex acuminate; only known sterile ....... 22. G. handelii
38b. Leaves oblong; apex acute; capsule with a distinct apophysis ................... ..
................... ....... ........... ............... ........ ................................ ............ 27. G. indica
37b. Lamina 2-stratose for more than the 1-2 marginal rows.
39a. Dry plants with leaves flexuous and crisped; 1 to 2 basal marginal leaf cell
rows inflated, usually reddish, forming 2-stratose alar areas; perichaetialleaves
virtually identical to vegetative leaves (1.5-3.2 mm vs. 2.3-3.2 mm long); setae
3-10 mm; capsules completely exserted ........................................... 7. G. atrata
0
0
WORLD SYNOPSIS OF GRIMMIA
9
39b. Dry plants with leaves erect, only flexuous at apical part; basal marginal
leaf cell rows not inflated, not colored, similar to adjacent inner rows, 1mm vs. 2-3
stratose; perichaetialleaves longer than vegetative leaves ＨＴＮｾＵ＠
mm long); setae 1.5-2.1 mm; capsules immersed ................... .47. G. ochyriana
32b. Leaves, at least perichaetial, with a trace of hyaline hair-point.
40a. Costa elliptical in cross section, scarcely prominent on dorsal surface,
occasionally of the same thickness as lamina, scarcely differentiated from lamina.
41 a. Proximal juxtacostal cells long rectangular (4-8: 1); perichaetialleaves of
mature sporophytes green and firm, neither hyaline nor filmy; setae 3-7 mm long;
capsule exserted.
42a. Seta curved; basal juxtacostal cell walls not nodulose ............... 48. G. olneyi
42b. Seta straight; basaljuxtacostal cell walls nodulose .................... 50. G. ovalis
41b. Proximaljuxtacostal cells shorter (1-4:1); perichaetialleaves of mature
sporophytes almost entirely hyaline and filmy (look carefully around capsule, inner
perichaetialleaves are almost invisible except under high contrast in the compound
microscope); seta to 1 mm; capsule immersed. [NOTE: sterile specimens of these 4
taxa are gametophytically identical and impossible to name with confidence;
separation based on ecological preferences is unrealistic].
43a. Capsule asymmetric and ventricose at base; seta sigmoid, eccentrically
attached to the capsule.
44a. Plants autoicous, male buds small and hidden just below perichaetia .........
................................................................................................... 2. G. americana
44b. Plants dioicous, male buds conspicuous, terminal on stem........................ ..
............................................................................................. 55. G. poecilostoma
43b. Capsule symmetric; seta straight, centrally attached to the capsule.
45a. Plants autoicous, male buds small and hidden just below perichaetia .........
................................................................................................ 28. G. involucrata
45b. Plants dioicous, male buds conspicuous, terminal on stem........................ ..
.................................................................................................. 63. G. tergestina
40b. Costa semicircular to reniform in cross section, prominent on dorsal surface,
differentiated from lamina.
46a. Costa winged, furrowed or irregularly triangular in shape on dorsal side.
47a. Ventral layer of costa 2 to 6 cells wide.
48a. Leaves muticous ................................................................. 59. G. ramondii
48b. Leaves with hair-point ...................................................... 23. G. hartmanii
47b. Ventral layer of costa 2 cells wide.
49a. Plants to 8 cm long; costa cross section triangular to almost circular, 3-4
layers thick from dorsal to ventral side at midleaf; cells usually papillose,
bulging and irregular; gemmae lacking......................................... 17. G. elatior
49b. Plants to 2 cm long; costa cross section flattened dorsally to semiterete,
2(3) layers thick from dorsal to ventral side at midleaf; cells never papillose,
bulging or irregular; gemmae common, arising from short stalks in leafaxils ....
................... ......... ...... ......... ........ ................... .......... ........... 41. G. muehlenbeckii
46b. Costa without wings or projections on dorsal side.
50a. Margins plane throughout or incurved above, no trace of recurvation.
51a. Laminar cells bulging in cross section [NOTE: Sterile or poor specimens
of the following three taxa are imposible to name].
52a. Seta less than 1 mm; capsule immersed ............................. 8. G. australis
10
MuNoz AND PANDO
52b. Seta more than 1.5 mm; capsule exserted.
53a. Stomata lacking at the urn base; capsule usually fusiform, castaneous,
concolorous with peristome teeth; exothecial cells isodiametric, thick-walled
................................................................................................. 1. G. aipestris
53b. Stomata present at the urn base; capsule ovoid, stramineous, different
in color from the orange peristome teeth; exothecial cells irregularly
rectangular, thin-walled ...................................................... 61. G. reflexidens
51 b. Laminar cells not bulging in cross section.
54a. Blades of distallarninal half forming <300 angle in cross section, parallel
to each other at their junction with the costa, furrow very narrow ................... .
.............................................................................................. 61. G. reflexidens
54b. Blades of distallaminal half forming >45 0 angle in cross section, not
parallel to each other at their junction with the costa, furrow more open.
55a. Distal half oflamina unistratose, only 1 or 2 marginal rows 2-stratose.
56a. Hyaline hair-point terete; seta more than 1 mm, curved; capsule
exserted, with peristome ....................................................... 24. G. humilis
56b. Hyaline hair-point flat at base; seta less than 1 mm, straight; capsule
immersed, gymnostomous ......................................... 56. G. pseudoanodon
55b. Distal half oflamina 2-3(4)-stratose.
57a. Lamina concave to V-shaped in cross section in the proximal part of
leaf; perichaetialleaves 5-15 times larger in area than other leaves .......... .
............................................................................................... 29. G. kidderi
57b. Lamina V -shaped in cross section in the proximal part of leaf;
perichaetialleaves 2-4 times larger in area than the other leaves [NOTE:
Sterile or poor specimens of the following four taxa are imposible to
name).
58a. Stomata present at urn base .................... 34. G. macroperichaetialis
58b. Stomata lacking at urn base.
59a. Leaves ± linear, distal 2/3 subulate; seta to 1 mm; capsule
immersed; peristome teeth rudimentary ..................... 37. G. mariniana
59b. Leaves lanceolate, distal 2/3 acuminate, but not subulate; seta
more than 2 mm; capsule emergent to exserted; peristome teeth well
developed.
60a. Opercula rostrate; seta 2-4 mm long; peristome teeth 50-90 ,urn
wide at mouth, irregularly splitting above and ± cribrose; proximal
juxtacostalleaf cells mostly longly rectangular, to 4.5: 1; plants
dioicous ...................................................................... 40. G. montana
60b. Opercula obtuse to mammillate; seta to 2 mm; peristome teeth
40-50 ,urn wide at mouth, entire or slightly cribrose at apex; proximal
juxtacostalleaf cells isodiametric to shortly rectangular, to 3: 1; plants
autoicous ........................................................................ 69. G. ungeri
50b. Margins recurved partially to fully on one or both sides.
61a. Ventral layer of costa 3-6 cells wide.
62a. Laminal cells bulging; costa semicircular in cross section .. 45. G. nutans
62b. Laminal cells not bulging; costa reniform in cross section.
63a. Proximal juxtacostal cells ofleaves with uniformily thickened walls ....
...................................................................................................... 32. G. lisae
63b. Proximal juxtacostal cells of leaves with nodulose walls.
11
WORLD SYNOPSIS OF GRIMMIA
64a. Perigonia just below perichaetia; hair-point of apical leaves mostly
spreading ........................................................................... 15. G. decipiens
64b. Perigonia terminal; hair-point of apical leaves erect.
65a. Leaves subulate, distal half V-shaped in cross section; costa almost
filling the apex .............................................................. 5. G. arcuatifolia
65b. Leaves acuminate but not subulate, distal halfU-shaped in cross
section; costa not filling the apex.
66a. Both margins recurved to revolute more than half their length;
hyaline hair-point flat; seta curved ................................ 31. G. leibergii
66b. One margin recurved for the proximal 112-2/3, the other flat or
only recurved for a very short distance from base; hyaline hair-point
terete; seta straight ................................................... .33. G. longirostris
61 b. Ventral layer of costa 2 cells wide.
67a. Laminal cells bulging.
68a. Plants reddish, dioicous ............................................ 51. G. percarinata
68b. Plants not reddish, autoicous.
69a. Leaves longer than 2 mm; seta curved ........................... 45. G. nutans
69b. Leaves shorter than 1.8 mm; seta straight ............... 61. G. reflexidens
67b. Laminal cells not bulging.
70a. Leaves oblong to elliptic, widest near middle or sides parallel, of equal
width ± from base to little above middle.
71a. Hyaline hair-point sharply dentate, to 0.4 mm long; plants dioicous ..
............ .... .... ....... .......... ..... ....... .... ......................... .................... 57. G. pulla
71b. Hyaline hair-point smooth or weakly denticulate, longer than 0.5 mm;
plants autoicous.
72a. Peristome teeth vestigial to absent.. ........................ 14. G. curviseta
72b. Peristome teeth well developed.
73a. Seta to 1 mm long, straight; capsule irnmersed ..... 12. G. capillata
73b. Seta longer than 2 mm, curved; capsule exserted
74a. Perigonia just below perichaetia; calyptra mitrate; peristome
teeth ± entire ............................................................. 58. G. pulvinata
74b. Perigonia terminal; calyptra cucullate; peristome teeth cribrose
and irregularly divided distally .............................. 49. G. orbicularis
70b. Leaves ovate to lanceolate, widest near base and conspicuously
tapering upward.
75a. Dry plants with leaves spirally imbricate along the stem; usually with
thread-like shoots with strongly differentiated, muticous and carinate
leaves ..................................................................................... 19. G.funalis
75b. Dry plants with leaves not spirally twisted along the stem; thread-like
shoots lacking.
76a. Lamina 2-3(4)-stratose for most of its upper 2/3 .......................... 77
77a. Leaves 2-3 mm, ovate, broad proximally, ± suddenly narrowed
into an upper subulate part, forming shoulders ............... 52. G. pilifera
77b. Leaves shorter than 1.9 mm, leaf base not forming shoulders.
78a. Blades of distallaminal half forming <30 angle in cross section,
parallel to each other at their junction with the costa, furrow very
narrow.
0
12
MuNoz AND PANDO
79a. Leaves lanceolate to narrowly lancolate; plants commonly
reddish; seta curved ............................................. 42. G. navieularis
79b. Leaves ovate-Ianceolate to lanceolate; plants never reddish;
seta straight .......................................................... 61. G. reflexidens
78b. Blades of distallaminal half forming >50 0 angle in cross
section, not parallel to each other at their junction with the costa,
furrow open.
80a. Leaves straight; setae straight; capsule immersed; plants
autoicous .................................... ;.............................. 38. G. molesta
80b. Leaves flexuous; setae curved; capsule exserted; plants
dioicous ...................................................................... 1O. G. hieolor
76b. Lamina mostly unistratose in its upper 2/3, with bistratose margins
and/or occasional and small 2-stratose patches.
81a. Leaves ovate ............................................................ 24. G. humilis
81 b. Leaves narrowly lanceolate to lanceolate.
82a. Blades of distallaminal half forming <300 angle in cross section,
parallel to each other at their junction with the costa, furrow very
narrow .................................................................... 42. G. navieularis
82b. Blades of distallarninal half forming >45 angle in cross
section, not parallel to each other at their junction with the costa,
furrow more open.
83a. Gemmae sessile on lamina, seldom on the costa, no stalk
remaining after gemmae liberation, which causeS leaf destruction;
lamina bistratose at margins and in streaks; plants less than 3 cm;
spores minutely granulose, apparently smooth under light
microscope ......................................................... 66. G. triehophylla
83b. Gemmae on stalks at dorsal base of costa, the stalk 1(2) cells
long remaining attached to the costa after gemmae liberation, which
does not affect the leaf; lamina unistratose except sometimes 1 or 2
marginal rows 2-stratose; plants to 8 cm; spores coarsely and
distinctly granulose under light microscope ....... 9. G. austrofunalis
0
13
WORLD SYNOPSIS OF GRiMMIA
COUNTRY CODES
(Iso 3166)
The country codes are listed alphabetically according to the standard ISO-3166. The number of
recognized taxa of Grimmia known from each country follows between brackets.
AFG Afghanistan (3)
AND Andorra (8)
ARG Argentina (14)
ARM Armenia (14)
ATA Antarctica (4)
ATF French Southern
Territories [Croizet
Island and Kerguelen
Island] (3)
AUS Australia (9)
AUT Austria (27)
AZE Azerbaijan (4)
BEL Belgium (17)
BGR Bulgaria (9)
BIH Bosnia and
Herzegovina (1)
BOL Bolivia (14)
BRA Brazil (4)
BTN Bhutan (1)
CAN Canada (27)
CHE Switzerland (30)
CHL Chile (10)
CHN China (16)
CMR Cameroon (2)
COG Congo (1)
COL Colombia (5)
CPV Cape Verde (2)
CRI
Costa Rica (2)
CYP Cyprus (6)
CZE Czech Republic (17)
DEU Germany (28)
DNK Denmark (5)
DOM Dominican
Republic (2)
DZA Algeria (10)
ECU Ecuador (6)
EGY Egypt (2)
ESP Spain (31)
ETH Ethiopia (4)
FIN
Finland (12)
FLK Falkland Islands (1)
FRA France (33)
FRO Faroe Islands (1)
GBR United Kingdom (21)
GEO Georgia (22)
GRC Greece (9)
GRL Greenland (11)
GTM Guatemala (5)
HND Honduras (1)
HUN Hungary (15)
ION Indonesia (1)
IND India (18)
IRL
Ireland (8)
IRN Iran (7)
IRQ Iraq (4)
ISL
Iceland (11)
ISR
Israel (5)
ITA
Italy (31)
JOR Jordan (3)
JPN
Japan (10)
KAZ Kazakstan (9)
KEN Kenya (6)
KGZ Kyrgyzstan (6)
KOR South Korea (1)
LBN Lebanon (2)
LBY Libya (1)
LKA Sri Lanka (2)
LSO Lesotho (7)
LVA Latvia (1)
MAR Morocco (10)
MDG Madagascar (2)
MEX Mexico (16)
MKD Macedonia, Former
Yugoslav Republic (6)
MNG Mongolia (8)
MWI Malawi (2)
MYS Malaysia (1)
NOR Norway (22)
NPL Nepal (4)
NZL New Zealand (12)
OMN Oman(l)
PAK Pakistan (9)
PAN Panama (1)
PER Peru (10)
PHL Philippines (1)
PNG Papua New Guinea (1)
POL Poland (20)
PRT Portugal (14)
REU Reunion (4)
ROM Romania (11)
SAU Saudi Arabia (6)
SDN Sudan (1)
SFE Russia, Far East (14)
SGS South Georgia (3)
SHN Tristan da Cunha,
Ascension, and st.
Helena Islands (1)
SIB
Russia, Siberia (23)
SJM Svalbard and Jan
Mayen Islands (1)
SUN Russia, European (20)
SVK Slovakia (20)
SVN Slovenia (1)
SWE Sweden (22)
SYR Syria (5)
TKM Turkmenistan (8)
TUN Tunisia (7)
TUR Turkey (19)
TWN Taiwan (5)
TZA Tanzania (6)
TZK Tajikistan (5)
UGA Uganda (8)
UKR Ukraine (11)
URY Uruguay (2)
USA U.S.A. (38)
UZB Uzbekistan (6)
VEN Venezuela (6)
VIR Virgin Islands,
U.S.A. (1)
YEM Yemen (1)
YUG Yugoslavia (9)
ZAF South Africa (8)
ZAR Zaire [Democratic
Republic of Congo] (2)
14
MuNoz AND PANDO
Alphabetical by country.
Afghanistan
AFG
DZA
Algeria
Andorra
AND
Antarctica
ATA
ARG
Argentina
ARM
Armenia
Australia
AUS
Austria
AUT
AZE
Azerbaijan
Belgium
BEL
BTN
Bhutan
Bolivia
BOL
Bosnia and
BIH
Herzegovina
BRA
Brazil
BGR
Bulgaria
CMR
Cameroon
CAN
Canada
Cape Verde
CPV
CHL
Chile
CRN
China
Colombia
COL
COG
Congo
CRI
Costa Rica
ATF
Croizet Island
CYP
Cyprus
Czech Republic
CZE
DNK
Denmark
Dominican Republic DOM
ECU
Ecuador
EGY
Egypt
ETH
Ethiopia
FLK
Falkland Islands
FRO
Faroe Islands
FIN
Finland
FRA
France
French Southern
Territories [Croizet
Island and Kerguelen
ATF
Island]
Georgia
GEO
DEU
Germany
Greece
Greenland
Guatemala
Honduras
Hungary
Iceland
India
Indonesia
Iran
Iraq
Ireland
Israel
Italy
Japan
Jordan
Kazakstan
Kenya
Kerguelen Island
Kyrgyzstan
Latvia
Lebanon
Lesotho
Libya
Macedonia, Former
Yugoslav Republic
Madagascar
Malawi
Malaysia
Mexico
Mongolia
Morocco
Nepal
New Zealand
Norway
Oman
Pakistan
Panama
Papua New Guinea
Peru
Philippines
Poland
Portugal
GRC
GRL
GTM
HND
HUN
ISL
IND
IDN
IRN
IRQ
IRL
ISR
ITA
JPN
JOR
KAZ
KEN
ATF
KGZ
LVA
LBN
LSO
LBY
MKD
MDG
MWI
MYS
MEX
MNG
MAR
NPL
NZL
NOR
OMN
PAK
PAN
PNG
PER
PHL
POL
PRT
Reunion
Romania
Russia (European)
Russia (Far East)
Russia (Siberia)
Saudi Arabia
Slovakia
Slovenia
South Africa
South Georgia
South Korea
Spain
Sri Lanka
Sudan
Svalbard and Jan
Mayen Islands
Sweden
- Switzerland
Syria
Taiwan
Tajikistan
Tanzania
Tristan da Cunha,
Ascension, and St.
Helena Islands
Tunisia
Turkey
Turkmenistan
U.S.A.
Uganda
Ukraine
United Kingdom
Uruguay
Uzbekistan
Venezuela
Virgin Islands
(U.S.A.)
Yemen
Yugoslavia
Zaire [Democratic
Republic of
Congo]
REU
ROM
SUN
SFE
SIB
SAU
SVK
SVN
ZAF
SGS
KOR
ESP
LKA
SDN
SJM
SWE
CHE
SYR
TWN
TZK
TZA
SRN
TUN
TUR
TKM
USA
UGA
UKR
GBR
URY
UZB
VEN
VIR
YEM
YUG
ZAR
WORLD SYNOPSIS OF GRiMMIA
15
SYNOPSIS OF THE GENUS GRiMMIA
1. Grimmia alpestris (F. Weber & D. Mohr) Schleich., Cat. PI. Helv. ed. 2: 29. 1808. Grimmia
alpestris Schleich., Neues J. Bot. 1(2): 196. 1805, nom. nud. Trichostomum pulvinatum var.
alpestre Schleich. ex F. Weber & D. Mohr, Bot. Taschenbuch: 110. 1807. Campylopus pulvinatus
var. alpestris (F. Weber & D. Mohr) Brid., Muscoi. Recent. Suppi. 4: 75. 1818 ["1819"].
Dryptodon pulvinatus var. alpestris (F. Weber & D. Mohr) Brid., Bryoi. Univ. 1(1): 198. 1826.
Grimmia donniana var. sudetica Huebener, Muscoi. Germ.: 175. 1833, nom. illeg., inci. typo var.
prior. Grimmia donniana var. alpestris (F. Weber & D. Mohr) Hampe, Flora 20: 281. 1837.
Guembelia alpestris (F. Weber & D. Mohr) Hampe, Bot. Zeitung (Berlin) 4(8): 125. 1846.
Grimmia alpestris var. eualpestris Loeske, Laubm. Eur. Part I: 101, fig. 25b, 27a-c. 1913, nom.
invai. PROTOLOGUE: [Switzerland] Grimm. alpestris Schleich. Cent. 2, n° 13 (in monte Sylvio
ad rupes a Schleichero lecta) et Dicr. piliferum ej. l. c. 4. n. 12. (in calidiorr, Vallesiae ad saxa
inventum [... ] In subalp., in rup. (in summo Bruct. ipsi legimus). TYPE: In M. Sylvio ad rupes
(lectotype, designated by Munoz (1997), BM!).
Grimmia holzingeri Cardot & Ther., in Holz., Bot. Gaz. 30: 123, tab. 11 fig. 2. 1900 [Wijk et al. (1962:
388) erroneously cited volume 15, 1890]. Grimmia alpestris var. holzingeri (Cardot & Ther.) G.
N. Jones, in Grout, Moss FI. N. Amer. 2: 31, pI. 9. 1933. Grimmia donniana var. holzingeri
(Cardot & Ther.) Wijk & Margad., Taxon 9: 190. 1960. PROTOLOGUE: [U.S.A., northwestern
Montana, leg. J.M. Holzinger & J. Blake, July 1898] Base of Sperry glacier; Mt. Trilby. TYPE:
[... ] vicinity of Lake McDonald, Flathead county, N.-W. Montana [... ] four miles north of Belton,
a station 30 miles east of Kalispell, on the Great Northern Railway. Mt. Trilby. Collected by John
M. Holzinger and James B. Blake. July 21. 1898. 3,500-7,000 Feet (lectotype, designated by
Munoz (1998b: 384), PC!; isolectotypes, FH!, H-BR! 2 replicates, NY! 2 replicates, PC!).
Synonymized by Munoz (1998b: 384).
Grimmia alpestris var. breviseta J.J. Amann, Fl. Mouss. Suisse 2: 139. 1918. PROTOLOGUE:
[Switzerland] Sur Ie gneiss erratique a Davos, 1600 m. (Amann) (BH 87.4.26). TYPE: Villa
Arnstein, Davos 1650-1700,20.4.90. Leg. Amann (BH 87.4.26) (holotype, Z!).
DISTRIBUTION: AFG (M K6ie s.n., MA); AND (A. Sotiaux s.n., BCB); ARM (V. Manakyan
4245, ERE); AUT (CC Townsend 921522, MA); BGR (J Podpt'ra s.n., TRH); CAN (J Macoun s.n.,
S); CHE (J Amann BH-18, Z); CZE (KG. Limpricht s.n., BP); DEU (Anonymous s.n., TRH); ESP (J
Munoz s.n., MA); FRA (R.B. Pierrot 50377, PC); GBR (HH. Knight s.n., FH); GEO (PH. Raven
27128B, MO); ITA (E. Corti s.n., FH); KAZ (B. Allen 10535, MO); NOR (N. Bryhn s.n., TRH); NPL
(C Troll 67, IE); PAK (KH. Rechinger s.n., W); POL (J Wojcicki s.n., MA); ROM (A. Boros s.n.,
BP); SFE (J. V. Chernyadjeva 16, LE); SIB (M Ignatov 01477B, MO); SVK (T. Chalubinski s.n., BP);
TUR (Davis & Hedge D.30403, MA); TZK (UK Mamatkulov s.n., LE); UKR (T. Alexeenko s.n., LE);
USA (A.T. Whittemore 3031, MO); UZB (A. Regel s.n., H-BR).
2. Grimmia americana E.B. Bartram, Bryologist 32: 8, pI. 1. 1929. PROTOLOGUE: [U.S.A.] Fort
Davis, Jeff Davis Co., Texas, altitude 5200 feet, May 22-25, 1926, C.R. Orcutt, no. 7082. TYPE:
Mosses of Western Texas. N° 7082. Fort Davis, Jeff Davis Co. alt. 5200 ft. C.R. Orcutt colI. May
22-25 1926 (lectotype, designated by Munoz (1999a: 123), FH!; isolectotype, NY!).
DISTRIBUTION: USA (CR. Orcutt 7082, FH).
16
MuNoz AND PANDO
3. Grimmia anodon Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 110, tab. 236. 1845. Anodon
ventricosus Rabenh., Deutsch!. Krypt.-Fl. 2(3): 154. 1848. Schistidium anodon (Bruch & Schimp.)
Loeske, Laubm. Eur. Part I: 49. 1913. PROTOLOGUE: In muris et ad rupes calcareas: prope
Heidelberg [Germany] ad arcem (AI. Braun), in Monte pinifero prope Gefrees (Funk); loco Bhiihl
prope Vindobonam [Austria] ad latera viae cavae (Dr. Putterlick); prope Mende Galliae [France]
meridionalis (Prost); in alpibus Tyrolis et Styriae [Austria] (W.P. Sch.). TYPE: An Mauern bei
Heidelberg. AI. Braun (lectotype, designated by Cao & Vitt (1986: 203), BM!; syntype
["Fichtelgebirge. Funck"], BM!).
Gymnostomum pulvinatum Hedw., Sp. Musc. Frond.: 36, tab. 3 fig. 1-7. 1801. Anictangium
pulvinatum (Hedw.) D. Mohr, Ann. Bot. (Konig & Sims) 2: 545. 1806. Anoectangium pulvinatum
(Hedw.) Rohl., Ann. Wetterauischen Ges. Gesammte Naturk. 2: 200. 1809. Schistidium
pulvinatum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 21. 1818 ["1819"). Anodon pulvinatus
(Hedw.) Rabenh., Deutschl. Krypt.-Fl. 2(3): 153. 1848. Grimmia hoffmannii Miill. Hal., Syn.
Musc. Frond. 1: 780. 1849. Grimmia sphaerica Schimp., Coroll. Bryol. Eur.: 49. 1856, nom. illeg.
incl. spec. prior. Grimmia apocarpa Hedw. subsp. sphaerica Boulay, Musc. France: 393. 1884.
Schistidium sphaericum G. Roth, Eur. Laubm. 1: 398, tab. 30 fig. 5. 1904, nom. illeg. incl. spec.
prior. Schistidium apocarpum (Hedw.) Bruch & Schimp. subsp. pulvinatum (Hedw.) Loeske,
Biblioth. Bot. 101: 51,61. 1930. Grimmia apocarpa Hedw. var. pulvinata (Hedw.) G.N. Jones, in
Grout, Moss Fl. N. Amer. 2: 16. 1933. Schistidium apocarpum (Hedw.) Bruch & Schimp. var.
pulvinatum (Hedw.) C.E.O. Jensen, in Weim., Forteckn. Skand. Vaxt., Moss. ed. 2: 28. 1937.
PROTOLOGUE: [Germany] Ad pagum Reinhausen, prope Goettingam, legit misitque Hoffmann
et Persoon. TYPE: Hab. ad rupestribus Gottingia leg. Hoffmann (lectotype, selected by Ochyra
(1989: 94), LE not seen; isolectotypes, H-SOL, LE, MW, none seen).
Synonymized by Ochyra (1989a).
Grimmia alpina Kindb., Christiania Vidensk.-Selsk. Forh. 1888(6): 30. 1889. PROTOLOGUE: Reg.
bet. in Jerkinhe prope viam veterem, Goverlivand versus! Reg. silv. super. DrivstuenL TYPE:
Norvegia: Dovre, Drivstuen. 317 87. N.C. Kindberg (lectotype, designated by Munoz (1999a:
125), S!; isolectotype, S!; syntype ["Jerkinhe"], S!).
Synonymized by Munoz (1999a: 125).
Grimmia anodon var. sinaitica Renauld & Cardot, Bull. Herb. Boissier 2: 33. 1894. PROTOLOGUE:
[Egypt] Sinai, Djebel Senah. Leg. F. Grote, decemb. 12, 1891. TYPE: F. Grote. Herbier
Sinaiticum. Djebel Senah. 12.xII.1891 (lectotype, here designated, PC!).
Grimmia dornajii Hohn., Sitzungsber. Kaiserl. Akad. Wiss., Math.-Naturwiss. Cl., Abt. 1, 104: 322.
1895. PROTOLOGUE: [Spain. Granada] Ich fand die Pflanze am 26.September 1892 in der
Seehohe von circa 1900 m auf Kalkfelsen, circa 200 m fiber Cortijo S. Geronimo am Westabhange
des Dornajo [F.v. Hohnel). TYPE: 26/9 1892. Kalkfelsen fiber Cortijo S. Geronimo c. 7000',
Westseite des Dornajo, Sierra Nevada, Spanien. Leg. et det. Fr. v. Hohnel (lectotype, here
designated, FH!).
Grimmia limprichtii Kern, Rev. Bryol. 24: 56. 1897. Grimmia anodon f. epilosa Monk., Laubm. Eur.:
345. 1927. Schistidium anodon f. limprichtii (Kern) Loeske, Biblioth. Bot. 101: 74. 1930.
Schistidium anodon f. epilosum (Monk.) Podp., Consp. Musc. Eur.: 273. 1954 ["epilosa"].
PROTOLOGUE: [Switzerland] sur les rochers calcaires du limon della Palla, au-dessus du defile
de Rolle, dans les monts Dolomites, a une altitude de 2,100 metres, Ie 29 juillet 1896 [F. Kern).
WORLD SYNOPSIS OF GRIMMIA
17
TYPE: Dolomiten, Palagruppe: Felsgallerien am limone, bei 2100m. 29.7.96 Kern (lectotype, here
designated, FH!; isolectotypes, GOET!, JE!, PC!).
Grimmia edentula Hy, Rev. BryoI. 32(4): 83. 1905. PROTOLOGUE: [France] [ ... ] Angers (Maine-etLoire) sur Ie ciment calcaire d'un mur expose au midi. TYPE: Murs du Chateau du Plain. 28 Janv.
1897 [F. Hy] (lectotype, here designated, ANGUC!).
Schistidium confertum var. bohemicum Vilh., Acta Bot. Bohem. 1: 53. 1922 ["bohemica"].
PROTOLOGUE: [Czech Republic] Habitat ad rupes in Bohemia occid.-merid. (Sumava) apud
Spiesova skala prope Janovice (1919, leg. dr. Hilitzer). TYPE: Spiessova skala 184/9i9 2NII 1919
leg. Hilitzer (holotype, Blom (1996: 262), PRC not seen).
Grimmia anodon var. anomala E.B. Bartram, Bryologist 27: 59. 1924. Grimmia anodon f. anomala
(E.B. Bartram) G.N. Jones, in Grout, Moss Fl. N. Amer. 2(1): 22. 1933. PROTOLOGUE: [U.S.A.]
On shaded banks of dry washes, Tucson Mts., Pima County, Arizona. E.B. Bartram, No. 931,
February 13, 1924. TYPE: Mosses of Southern Arizona. Steep shaded banks of dry washes in
Tuscon Mountains, Pima County. Altitude 2300 ft. Feb. 13 1924. Collected by Edwin B. Bartram
(lectotype, designated by Muiioz (1999a: 125), FH!; isolectotype, H-BR!).
Synonymized by Munoz (1999a: 125).
Schistidium tibetanum J.S. Lou & P.C. Wu, Acta Phytotax. Sin. 18: 121, pI. 2 fig. 8-15. 1980.
PROTOLOGUE: [Tibet] Xizang: Baingoin on rocks, 10.VI.1976, B.S. Li 7601 (Type in Herb.,
Institute of Botany, Academia Sinica). TYPE: (holotype, PE not seen).
Synonymized by Cao & Vitt (1986: 203).
Schistidium obtusifolium Ireland & Crum, Bryologist 87: 371-373. 1984. Grimmia subanodon Ochyra,
Mem. New York Bot. Gard. 45: 607-614. 1987. PROTOLOGUE: [and TYPE.] Canada, Quebec,
Parc du Bic, Cap Emage, pointe Ouest, 48°22'N, 68°46'W, on dry, calcareous conglomerate, 15
June 1983, Zoladeski & Lutzoni 11201 (holotype, QEF!; isotype, CANM!).
DISTRIBUTION: ARM (N V Chikovani s.n., LE); AUT (J. Fiedler s.n., H); BOL (M. Lewis 791947, F); CAN (D.H. Vitt 23986, H); CHE (J. Amann BH-1O, Z); CHL (H. Deguchi 26028, HIRO);
CRN (B. Tan 95-199, FH); CZE (R. Van& s.n., MEL); DEU (Schmidt 1952, JE); DZA (L. Trabut
1782, PC); EGY (A. Kneucker 13, JE); ESP (J. Munoz s.n., MA); FRA (J.L. De Sloover 34976, MO);
GEO (A.L. Abramova & l.I. Abramov s.n., LE); GRC (R.E. Magill 1 00 74A , MO); HUN (R. Marstaller
s.n., JE); IND (W Koelz 2898, MO); IRN (P. Goldblatt 830a, MO); ITA (F. Kern s.n., FH); JOR (W
Frey & H. Kiirschner 1136, MA); KAZ (B. Allen 10786, MO); KGZ (VF. Brotherus s.n., FH); MAR
(J. Munoz s.n., MA); MEX (A.J. Sharp 3912, TENN); NOR (NC Kindberg s.n., S); PAK (T Thomson
287, FH); PER (H. Deguchi 29980, HIRO); SAU (W Frey & H. Kiirschner 2709, MA); SFE (o.M.
Afonina s.n., LE); SIB (M. Ignatov 01478A, MO); SUN (VP. Savicz s.n., FH); SVK (J. Gyorffy 21, H);
SWE (N Hakelier s.n., MO); TKM (J. V Sirotina s.n., LE); TUR (E. Nyholm & A.C Crundwell1229,
F); TZK (T Ahti 30324, H); USA (P.L. Redfearn Jr. 263-51, MO); YUG (M. Sabovljevid s.n., MA).
4. Grimmia anomala Hampe ex Schimp., Syn. Musc. Eur. ed. 2: 270. 1876. Grimmia elatior subsp.
anomala (Hampe) Kindb., Eur. N. Amer. Bryin. 2: 222. 1898. Dryptodon anomalus (Hampe)
Loeske, Hedwigia 49: 32. 1909. Grimmia hartmanii var. alpinoborealis Loeske, Laubm. Eur. Part
I: 141. 1913, nom. invaI., pro synon. Grimmia hartmanii var. anomala (Hampe) Monk., Laubm.
18
MuNoz AND PANDO
Eur.: 369. 1927. Grimmia hartmanii subsp. anomala (Hampe) Loeske, Biblioth. Bot. 101: 182.
1930. PROTO LOGUE: [Switzerland] ad rupes prope Zermatt Valesiae rever. Bertram legit.
TYPE: (holotype, BM not seen).
Grimmia philibertiana E. Britton, Bull. Torrey Bot. Club 18: 51, pI. 114 fig. 18-28. 189l.
PROTOLOGUE: [U.S.A.] Rare on quartz croppings of metalliferons veins, 6,000 ft. alt., divide
between Traille River and Independence Creek, Sept. 27, 1889 (219). TYPE: 6000ft. alt. Traille
River divide [illegible] Independence. ColI. J.B. Leiberg Sept. 27th 89. 219 (holotype, NY!;
isotype, PC!).
Synonymized by Deguchi (1978: 163).
Grimmia pachyneurula Mull. Hal. & Kindb., in Macoun, Cat. Canad. PI., Musci: 65. 1892. Grimmia
apocarpa subsp. pachyneurula (Mull. Hal. & Kindb.) Kindb., Eur. N. Amer. Bryin. 2: 234. 1898.
Schistidium pachyneurulum (MUll. Hal. & Kindb.) Ochyra, Fragm. Florist. Geobot. 43: 106. 1998.
PROTOLOGUE: [Canada] On rocks, near the Whirpool on the west side of the Columbia, at
Revelstoke, B.C. May 7th, 1890. (Macoun.). TYPE: Grimmia apocarpa pachyneurula C.M. et
Kindb. ad int. N. Amer., Brit. Col., Revelstoke. 7/5 90 J. Macoun (lectotype, designated by Blom
(1996: 262), S not seen).
Synonymized by Blom (1996: 262).
Grimmia phyllantha Lindb. ex Broth., Acta Soc. Sci. Fenn. 19(12): 83. 1892. Racomitrium
phyllanthum (Broth.) Warnst., Hedwigia 33: 306. 1913. PROTOLOGUE: [Georgia] Cauc. major
occ.: Radscha, in monte Bereleti pr. Uzeri, ad saxa regionis alpinae. TYPE: Radscha, in alpe
Bereleti pro Uzeri, ad saxa reg. alp. 3/7/1877. Leg. V.F. Brotherus (holotype, H-BR!; isotypes, HSOL!, PC! 2 replicates).
Synonymized by Deguchi (1978: 163).
DISTRIBUTION: AUT (W.E. Nicholson & H.N. Dixon s.n., JE); CAN (G.R. Brassard 10990b,
NY); CHE (P. Culmann s.n., MA); ESP (I. Aizpuru s.n., MA); GEO (G. Woronow 114, PC); GRL (K.
Holmen s.n., NY); IND (L. Borelli s.n., H-BR); ISL (B. J6hannsson 11992, ICEL); NOR (Bryhn s.n.,
JE); SWE (c. Stenholm s.n., MO); USA (D.H. Norris 57676, NY).
5. Grimmia arcuatifolia Kindb., Bull. Torrey Bot. Club 16: 93. 1889. PROTOLOGUE: [Canada]
Abundant on dry rocks at Cedar Hill, near Victoria, Vancouver Island. April 21, 1887. John
Macoun. TYPE: Canadian musci n° 608. On dry rocks, Mount Tolmie. Near Victoria, V.I., April
21st, 1887. ColI. J. Macoun (lectotype, designated by Munoz (1999b: 7), CANM-198077!;
isolectotype, FH!).
Racomitrium alternuatum Mull. Hal. & Kindb., Cat. Canad. PI., Musci: 73. 1892. Grimmia alternuata
(Mull. Hal. & Kindb.) Kindb., Eur. N. Amer. Bryin. 2: 228. 1898 ["attenuata"]. Grimmia
attenuata (Mull. Hal. & Kindb.) Mull. Hal. & Kindb, Eur. N. Amer. Bryin. 2: 228. 1898, nom.
inval., orthogr. err. PROTOLOGUE: [Canada] On boulders at the base of Avalanche Mountain,
Roger's Pass, Selkirk Mountains, B.C., Aug. 5th, J890. (Macoun). TYPE: along the C. P. R'y at
the summit of Roger's Pass, Selkirk Mountains, Aug. 6th, 1890. [Macoun, Canadian musci, n°
524] (lectotype, designated by Munoz (1999b: 7-8), FH!; NOTE: Other duplicates are G. elatior
and are excluded from the type).
WORLD SYNOPSIS OF GRIMMIA
19
Grimmia procera Kindb., Rev. Bryol. 23: 18. 1896, nom. illeg., non Ba1s.-Criv. & De Not. 1838.
PROTOLOGUE: Amer. Canada, Vancouver isl. (without capsules): Macoun. TYPE: On rocks,
Parson's Mt. Van. Island. Col. Macoun. Date May 19th, 1893. [Macoun, Canadian mosses n° 88a,
n° 133] (lectotype, designated by Munoz (1999b: 8), CANM-198094!; isolectotypes, CANM198095!, FH!, NY!).
DISTRIBUTION: CAN (RL Halbert 4461, FH); USA (DL Branscomb & J. Grunman 22826,
MO).
6. Grimmia arenaria Hampe, Linnaea 10: 405. 1836. Grimmia curvula Bruch & Schimp., Bryol.
Europ. (fasc. 25-28) 3: 113, tab. 238. 1845, nom. illeg., incl. typo spec. prior. Grimmia incurva
Schleich. ex Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 113. 1845, nom. inval., pro synon.
Grimmia donniana var. curvula Spruce, Musci pyrenaicae n° 281. 1847. Grimmia donniana subsp.
arenaria (Hampe) Dixon, Stud. Handb. Brit. Mosses ed. 2: 155. 1904. Grimmia donniana var.
arenaria (Hampe) Loeske, Laubm. Eur. Part I: 93, fig. 1c, 17a, 26b, 28. 1913, nom. illeg., incl.
typo var. prior. PROTOLOGUE: [Germany] Habitat in rupibus et saxis arenosis ad arcis pedem
Regenstein, aestate fructu maturo. TYPE: ad rup. aren. Regenstein Hercyn[iae]. Junis (lectotype,
designated by Munoz (1998b: 376), BM!; isolectotype, FH!).
DISTRIBUTION: AUT (H. Gander s.n., BP); CHE (c. Trautmann s.n., S); DEU (E. Hampe s.n.,
BM); ESP (c. Casas s.n., MA); FIN (V.F. Brotherus s.n., H-SOL); FRA (L.A. Brebisson s.n., FH);
GBR (D.A. Jones & P.G. Rhodes 2169, BP); ITA (V. de Cesati s.n., G).
7. Grimmia atrata Miel. ex Hornsch., Flora 2: 85. 1819. Grimmia alpestris var. brevipila Bruch ex
Mont., Arch. Bot. 1: 219. 1833, nom. inval., pro synon. Dryptodon atratus (Hornsch.) Hartm.,
Handb. Skand. Fl. ed. 3: 271. 1838. PROTOLOGUE: [Austria] Salzburg, an Felsen bei der Grube
Schwarzwand in der Grosarl. luI. TYPE: Grosarl. Hornschuch misit 1819 (lectotype, designated
by Deguchi (1978: 170), B!; isolectotype, F!).
Grimmia strictifolia Mitt., J. Linn. Soc., Bot. Suppl. 1: 44. 1859. Racomitrium strictifolium (Mitt.) A.
Jaeger, Ber. Tatigk. St. Gallischen Naturwiss. Ges. 1872-73: 89 (Gen. Sp. Musc. 1: 371). 1874.
PROTOLOGUE: [India] In Himalayae orient. reg. alp., Sikkim, J.D. Hooker! TYPE: 295. Jongri
alt. 13.000 (lectotype, designated by Munoz (1999a: 126-127), NY!).
Synonymized by Cao & Churchill (1995: 508).
Grimmia tristicha Herzog, Biblioth. Bot. 87: 58. 1916, nom. illeg., non Schwaegr. 1811. Grimmia
tristichoides Margad., Lindbergia 1: 128. 1972, nom. nov. PROTOLOGUE: An feuchten Felsen
im Hochtal von Viloco, ca. 4700 m, No. 3152 c. fr. u. 3170 (ster.); an nassen Felsen bei der Mine
Chojnacota, ca. 4700 m, No. 2979. TYPE: Plantae in itinere secundo per Boliviam lectae, an
nassen Felsen im Hochtal von Viloco. ca. 4700m. leg. Th. Herzog n° 3152. Oktober 1911
(lectotype, designated by Munoz (1999a: 127), JE!; isolectotype, PC!; syntypes ["Herzog 2979,"
"Herzog 3170"], JE!, NY!).
Synonymized by Munoz (1999a: 127).
20
MuNoz AND PANDO
Grimmia tristicha var. comosa Herzog, Biblioth. Bot. 87: 59. 1916. Grimmia tristichoides f. comosa
(Herzog) Margad., Lindbergia 1: 128. 1972. PROTOLOGUE: An nassen Felsen im Hochtal von
Viloco, ca. 4700 m, No. 3188. TYPE: Plantae in itinere secundo per Boliviam lectae. An nassen
Felsen im Hochtal von Viloco. ca. 4700m. leg. Th. Herzog. Oktober 1911 (lectotype, designated
by Munoz (1999a: 127), JE!; isolectotype, PC!).
Synonymized by Munoz (1999a: 127).
DISTRIBUTION: AUT (H.c. Funck s.n., H); BEL (J.-L. De Sloover 13701, PC); BOL (M Lewis
87-884, MO); CZE (z. Pilous s.n., H); DEU (P.G. Lorentz s.n., PC); ESP (P. Montserrat et al. s.n.,
BCB); FRA (J. Cardot s.n., H); GBR (D.G. Long & R.M Murray 14767, MA); IND (J.D. Hooker 295,
NY); ITA (Artaria s.n., JE); JPN (Faurie 3380, NY); NOR (N Hakelier s.n., MO); SWE (G.
Martensson s.n., MO); TWN (S. Lin s.n., NICH)
8. Grimmia australis (Dixon & Sainsbury) J. Munoz & Ochyra, J. Bryol., 21: 319. 1999. Coscinodon
australis Dixon & Sainsbury, Trans. & Proc. Roy. Soc. New Zealand 75(2): 175. 1945.
PROTOLOGUE: [New Zealand] Mount Cassidy, Arthur's Pass, Canterbury; colI. G. O. K. S.
31112/1935, No. 827. TYPE: Mt. Cassidy, Arthur Pass, Canterbury, South Island, N.Z., 31/12/35.
CoIl. Sainsbury n° 827 (lectotype, designated by Munoz & Ochyra (1999), BM!; isolectotypes, F!,
FH!,NY!).
Grimmia immersa Ochyra, Fragm. Florist. Geobot. Suppl. 2, 1: 223. 1993. PROTOLOGUE: New
Zealand. South Island. Canterbury. Mount Cook National Park, end of North Ridge, Mt.
D'Archiac, elev. 6100 ft., rotten rock and rubble; 26 January 1972, H.D. Wilson 2314 (Holotype:
CHR-255623). TYPE: New Zealand. Mount Cook National Park. End of North ridge, Mt.
D'Archiac 6100'.26/1/1972: H. D. Wilson n° 2314 (holotype, CHR-255623!).
Synonymized by Munoz & Ochyra (1999).
Grimmia perpilosa Ochyra, Fragm. Florist. Geobot. Suppl. 2, 1: 220. 1993. PROTOLOGUE: New
Zealand. South Island. Southland. Fiordland National Park, Gertrude Saddle, c. 4 000 ft, on rock,
23.XII.74, John Child s.n. (holotype, BM!; isotype, KRAM). TYPE: Grimmia ? asperifolia c.fr.
Gertrude Saddle. C. 4000'. 23.XII.74, on rock. John Child Collection (holotype, BM not seen;
isotype, KRAM-99498!).
Synonymized by Munoz & Ochyra (1999).
DISTRIBUTION: AUS (J.M Wilson s.n., MEL); NZL (G.O.K. Sainsbury 827, BM).
9. Grimmia austrofunalis Mull. Hal., Hedwigia 37: 165. 1898. PROTOLOGUE: Australia, Victoria,
Grampians, Mt. William: Sullivan 1883 in Rh. Melbourne. TYPE: Grampians, mount William,
1883. Sullivan (holotype, MEL!; isotypes, MEL! 2 replicates, S!).
NOTE: A duplicate in H-BR is G. trichophylla and is excluded from the type.
Grimmia pansa R.S. Williams, Bull. New York Bot. Gard. 3(9): 125. 1903. Grimmia pausa R.S.
Williams ex P. Syd., Just's Bot. Jahresber. 31(1): 262. 1904, nom. inval., orthogr. err.
PROTOLOGUE: [Bolivia; leg.: R.S. Williams] Pelichuco, 3450 meters, May 4,1902 (2823).
WORLD SYNOPSIS OF GRIMMIA
21
TYPE: Bolivia. Pelechuco, 3450 meters. May 4, 1902. R.S. Williams 2823 (lectotype, designated
by Munoz (1999a: 127), NY!; isolectotypes, FH!, JE!).
Synonymized with G. trichophylla by Deguchi (1987: 30), but see Munoz (1999a: 127).
Racomitrium austrosudeticum Broth., in Herzog, Biblioth. Bot. 87: 60. 1916 ["Rhacomitrium"].
PROTOLOGUE: [Bolivia] Waldgrenze tiber Tablas, alt. 3400 m, No. 2859; Tunarisee, alt. 4400
m, No. 4807; Cerros de Malaga, alt. 4000 m, No. 4366; Choquecota chico, alt. 4500 m, No. 3179.
TYPE: Waldgrenze tiber Tablas, 3400 m, Okt. 1911 Th. H. 2859 (lectotype, designated by Munoz
(1999a: 127), H!; isolectotype, JE!; syntypes ["Herzog 3179"], H!, JE!; ["Herzog 4366"], H!;
["Herzog 4807"], H!).
Synonymized by Munoz (1999a: 127).
Grimmia crassiretis Cardot & Broth., Kongl. Svenska Vetenskapsakad. Handl. 63(10): 27, tab. 2 fig. 7.
1923. PROTOLOGUE: [Argentina] Patagonia andina: Lago San Martin (380); Valle Frias, Cerro
Caceres, in declivi meridionali, alt. c. 900 m.s.m. TYPE: 380. Patagonie: lac San Martin, foret
mixte. 27.1.1909. C. Skottsberg (lectotype, designated by Munoz (1999a: 127), PC!; isolectotypes,
H-BR!, UPS!; syntype ["Valle Frias"], PC!).
Synonymized with G. trichophylla by Deguchi (1984: 25), but see Munoz (1999a: 127).
Grimmia benoistii Ther., Rev. Bryol. Lichenol. 9: 10. 1936. PROTOLOGUE: [Ecuador, leg. R.
Benoist] Massif du Pichincha: rochers verticaux exposes au Nord, face au Condorguachana, 3.900
m. (n° 3116,3117,3114 p.p.). TYPE: 3116. en face de Condorguachana (massif du Pichincha)
30.x.1930. R. Benoist (lectotype, designated by Munoz (1999a: 127), PC!; isolectotypes, F!, PC!;
syntypes ["Benoist 3117"], PC!, F!).
Synonymized with G. trichophylla by Churchill (1994: 2), but see Munoz (1999a: 127).
NOTE: All European specimens identified as G. austrofunalis are other species. Greven (1997:
830) listed several specimens, from which the first author was only able to study Greven 2020 (S). It is
typical G. longirostris, with reniform costa, long-rectangular, nodulose, juxtacostal cells, and no
gemmae at costa back.
DISTRIBUTION: ARG (c. Skottsberg 381, PC); AUS (c. Beauglehole 15019, MEL); BOL (M
Lewis 79-1354, F); CHL (H.H. Bartlett 19123, NY); COL (S.P. Churchill & W Rengifo M 17258,
MA); ECU (R. Benoist 3116, PC); NZL (1. Child M1460, F); PER (1.-P. Frahm 824021, MA); KEN
(E. Schelpe 12182, MO); VEN (D. Griffin III et al. PV-1170, CAS).
10. Grimmia bicolor Herzog, Beih. Bot. Centralbl. 26(2): 66. 1909. PROTOLOGUE: Bolivia: Auf
Felsblocken im Llavetal (Prov. Cochabamba), ca. 3600 m; Januar, 08. TYPE: Flora von Bolivia.
Auf FelsblOcken im Llavethal (Prov. Cochabamba) ca. 3600m. leg. Th. Herzog. Januar, 08
(lectotype, designated by Munoz (1999a: 131), JE!; isolectotype, PC!).
Grimmia subquatricruris Broth., in Herzog, Biblioth. Bot. 87: 57. 1916. PROTOLOGUE: Felsen am
Huaillattanisee, alt. 4900 m, No. 2973. TYPE: Bolivia, Felsen am Huaillattanisee, 4900 m,
22
MuNoz AND PANDO
10/1911. Leg. Th. Herzog 2973 (lectotype, designated by Munoz (1999a: 131), H-BR!;
isolectotype, JE!).
Synonymized by Munoz (1999a: 131).
DISTRIBUTION: BOL (M Lewis 79-2581, F).
11. Grimmia caespiticia (Brid.) Jur., Laubm.-Fl. Oesterr.-Ung.: 172. 1882. Campylopus caespiticius
Brid., Muscol. Recent. Suppl. 4: 77. 1818 ["1819"]. Dryptodon caespiticius (Brid.) Brid., Bryol.
Univ. 1(1): 200. 1826. Grimmia funalis var. caespiticia (Brid.) Hampe, Flora 20: 282. 1837
["caespititium"]. Guembelia caespiticia (Brid.) Mull. Hal., Syn. Musc. Frond. 1: 773. 1849.
Guembelia caespitosa Miill. Hal. ex Limpr., Laubm. Deutschl. 5(synon. reg.): 23. 1903, nom.
inval., orthogr. err. Grimmia cespiticia Jur. ex Dalla Torre & Sarnth., Fl. Tirol 5: 268. 1904, nom.
inval., orthogr. err. Grimmia alpestris var. caespiticia (Brid.) G.N. Jones, in Grout, Moss Fl. N.
Amer 2: 30. 1933. PROTOLOGUE: [Switzerland] In Monte Sancti Bernhardi Helvetiae habitat,
unde lectum accepi. TYPE: St. Bernard (holotype, Bl).
Grimmia sulcata Saut., Flora 24: 39. 1841. Guembelia sulcata (Saut.) Hampe, Bot. Zeitung (Berlin)
4(8): 125. 1846. PROTOLOGUE: [Austria. Pinzgau ...bei Annaberg] An Felsen, gleich unter dem
Kamm des nordlichen Auslaufers des Sauecks im Norden von Stuhlfeden, neben dem Alpenwege,
der in das Schwarzachenthal fiihrt in 6000' Seehohe [... ] August 1837. TYPE: Pinzgauer Alpen.
Com. Dr. Sauter (lectotype, designated by Munoz (1998b: 387), H-SOL!; isolectotype, Zl).
Synonymized by Munoz (1998b: 387).
Grimmia jacquinii var. subimberbis Lindb., Ofvers. Forh. Kongl. Svenska Vetensk.-Akad. 23: 552.
1867. Grimmia alpestris var. subimberbis (Lindb.) Berggr., Kongl. Svenska Vetenskapsakad.
Handl. 13(7): 49. 1875. Grimmia caespiticia var. subimberbis (Lindb.) Limpr., Laubm. Deutschl.
1: 780. 1889. Grimmia sulcata var. subimberbis (Lindb.) G. Roth, Eur. Laubm. 1: 431. 1903.
Grimmia caespiticia f. subimberbis (Lindb.) Podp., Consp. Musc. Eur.: 283. 1954.
PROTOLOGUE: [Norway. Spitsbergen] Amsterdamon. TYPE: Spitsb., Amsterdam Island. 1861.
Leg. Holmgren (lectotype, designated by Munoz (1998b: 387), H-SOL!).
Synonymized by Munoz (1998b: 387).
Grimmia manniae Mull. Hal., Flora 70: 223. 1887. Grimmia alpestris var. manniae (Mull. Hal.) G. N.
Jones, in Grout, Moss Fl. N. Amer. 2(1): 31, pI. 9. 1933. Grimmia donniana var. manniae (Mull.
Hal.) Wijk & Margad., Taxon 9: 50. 1960. PROTOLOGUE: [U.S.A.] California, Napa Co., Napa
Springs: Domina Martha Mann 2. Majo 1886 legit. TYPE: Napa Soda Springs. California. May 3.
1886. Martha R. Mann (lectotype, designated by Munoz (1998b: 387), NYl; isolectotypes, FHl,
PC!).
Synonymized by Munoz (1998b: 387).
Grimmia pyrenaica Kern, Jahresber. Schles. Ges. Vaterl. Cult. 92: 38. 1915. PROTOLOGUE: [Spain]
Auf eisenhaltigen Felsen auf der spanischen Seite des Port de Boucharo, c. 2230 m. TYPE:
Gavernie, auf eisenhalbsigen Felsen dem Port de Boucharo, spanische Seite. 13-VII-1914. Kern
(lectotype, designated by Bednarek-Ochyra et al. (1992: 392), WRSLl; isolectotypes, BP-80423!,
JEl, KRAMl, MA-8158!, MA-10927l).
Synonymized by Bednarek-Ochyra et al. (1992: 392).
WORLD SYNOPSIS OF GRiMMIA
23
DISTRIBUTION: AND (R.B. Pierrot 2011100, PC); ARM (v. Manakyan 4996a, ERE); AUT (JP. Frahm 921007, MA); BEL (A. Cornet s.n., NY); BGR (A. Ganeva s.n., MO); CHE (C Bender BH16, Z); DEU (Anonymous s.n., BP); ESP (J Munoz s.n., BCB); FRA (JP. Hebrard 1611774, PC);
GEO (G. Woronow 192A, PC); ITA (JP. Hebrard 7-2882, PC); POL (T. Chalubinsky s.n., BP); PRT
(C Sergio & A. Seneca 8164, MA); ROM (L. Vajda s.n., BP); SIB (L. V. Bardunov s.n., NICH); SJM
(HJ Holmgren s.n., H-SOL); SUN (A.H. Brotherus & v.F. Brotherus 22, H-SOL); SVK (z. Pilous
s.n., G); TUR (K. Walther 3318, NY); USA (E. Lawton 4792, MO).
12. Grimmia capillata De Not., Mem. Reale Accad. Sci. Torino 39: 248. 1836. Grimmia crinita var.
capillata (De Not.) De Not., Syllab. Musc.: 240. 1838. PROTOLOGUE: Ad muros vetustos
hortorum meridiem versus. TYPE: (RO not seen).
Grimmia mairei Cardot & Copp., Bull. Soc. Bot. France 58: 500, fig. 15a-m. 1911. PROTOLOGUE:
[Algeria] [... ] recueillie par M. Rene Maire, a qui elle est dediee, sur les rochers de gres du ravin
de Ben Zireg, dans l'Extreme-Sud Oranais. TYPE: Rochers de gres aBeni-Ounif, Sahara algerien.
Leg. R. Maire - 9/4/06 (lectotype, here designated, PC!; isolectotype, PC!).
Grimmia mesopotamica Schiffn., Ann. K. K. Naturhist. Hofmus. 27: 488. 1913. PROTOLOGUE:
[Iraq] Ad terram et rupes calciferas Mesopotamiae mediae, c.sp. Erde der Gipswiiste unterhalb Hit
am rechten Ufer des Euphrat, ca. 150 m, 10.IIV.1910 (Nr. 844); Kalkmergelfelsen bei der
Dorfruine Gharra in der Mitte des NordfuBes des Dschebel Abd-el-Asis, ca. 500 m, 21.N1.1910
(Nr. 1778). TYPE: No. 844. Ad confines Mesopotarniae et Arabia borealis: Ad Euphratem
medium inter Abukemal et Ramadi, in desertis infra oppidum Hit. Substrato gipsaceo; ca. 120180 m. Leg. 10.lV.l91O Dr. Heim. Frh.v. Handel-Mazzetti. (Tageb.- No. 154) determ. Schiffner
(lectotype, here designated, FH!; isolectotype, FH!; syntypes ["Handel-Mazzetti 1778"], FH!, HBR!).
Grimmia capillata var. calva 1.J. Amann, in E. Bauer, Musci europaei et americani exsiccati ser. 39 n°
1915. 1927. Grimmia crinita var. calva (J.J. Amann) 1.J. Amann, in Loeske, Biblioth. Bot. 101:
88. 1930. PROTOLOGUE: Frankreich: Aix-en-Provence, an Mauem der Route d'Avignon, A.
18.Sept.l925 und B. 31.Mai.1926 legit J. Amann. TYPE: Frankreich: Aix-en-Provence, an
Mauem der Route d'Avignon, B. 31.Mai.1926 legit 1. Amann (lectotype, here designated, MA7092!; isolectotype, JE!).
DISTRIBUTION: DZA (L. Trabut 1761, PC); ESP (J Guerra et al. s.n., MA); IRQ (HF.v.
Handel-Mazzetti 844, FH); ISR (/. Herrnstadt 4510, FH); JOR (u. Baierle & H Kiirschner 2579,
MA); MAR (J Munoz s.n., MA); PAK (C Troll 38, JE); SIB (L. Bardunov s.n., ALTA); SYR (HF.v.
Handel-Mazzetti 1778, FH); TKM (M.M. Il'in s.n., LE); TUN (T.P. Maslin B-32420, NY).
13. Grimmia crinita Brid., nom. conserv. prop., Muscol. Recent. Suppl. I: 95. 1806. Guembelia
crinita (Brid.) Hampe, Bot. Zeitung (Berlin) 4(8): 124. 1846. PROTOLOGUE: In tota Europa
Australiori habitat miseram vitam in aridis degens. In Pago Vaudensi ad muros Lausannae legi,
Roger Neoduni; praeterea in Valle-Clausa Gallo Provinciae, Massiliae, in Occitania et Catalonia
ad muros faxaque copiose reperi. A Ianuario ad ApriIem usque cum capsulis maturescentibus,
24
MuNoz AND PANDO
perfectis essoetisque vidi. TYPE: Avril 02. Rogeri s.n. (lectotype, designated by Geissler & Maier
(1995: 499), Bl).
NOTE: The name G. crinita has been proposed for conservation (Munoz, 2000b).
Dicranum phascoideum P. Beauv., Prodr. Aetheogam.: 54. 1805 ["phascoidum"]' PROTOLOGUE:
[France] [... ] M. Dufour, se trouve sur un des murs du bois de Boulogne, pres Auteuil. TYPE:
Herbier Palisot de Beauvois. Type de "Dicranum ? phascoideum". Sur les murs du bois de
Boulogne. Echantillon original provenant de l'herbier Desportes (lectotype, designated by Munoz
(2000b), Gl; isolectotype, PC-P.Beauvl).
Gymnostomum decipiens F. Weber & D. Mohr, Bot. Taschenbuch: 79.1807. PROTOLOGUE: In saxis
pro Herbipolin [Wiirzburg]. (Eundem muscum ad fontem Vaucluse Galliae lect. mis. Bridel).
TYPE: [Germany] Ex herb. D. T. [D. Turner]. Gymn. decipiens n. sp. F1. Germ. Wirceburgi
[Wiirzburg] (lectotype, here designated, BMl).
Grimmia crinita var. libani Bizot, Rev. Bryol. Lichenol. 13: 51. 1942. PROTOLOGUE: [Lebanon]
Katana [a 25 km. au sud-ouest de Damas. Maurize Nickles, 1932]. TYPE: Katana, Beyrouth,
Liban. 4 septembre 1932. leg. Nickles (holotype, PCl).
DISTRIBUTION: AZE (Alexeenko s.n., LE); BEL (c. Delogne s.n., FH); CHE (P. Culmann s.n.,
MO); CZE (VelenovskY s.n., JE); DEU (L. Meinunger 17843, JE); DZA (L. Trabut s.n., PC); ESP (J.
Munoz s.n., MA); FRA (0. Aicardi 501, PC); GEO (N. V. Chikovani s.n., LE); IRL (s. W Greene S.n.,
TCD); ISR (F. Bilewsky 658, PC); ITA (Loitlesberger s.n., JE); LBN (M. Nickles s.n., PC); TKM (V.F.
Brotherus s.n., FH-SULL); TUN (P. Robinson s.n., MO).
14. Grimmia curviseta Bouman, J. Bryol. 16: 379. 1991. PROTOLOGUE: Tenerife (Spain, Canary
Islands) Las Canadas [Canadas] del Teide, Degollada de Guajara, 2400 m altitude, 3 January
1989, UTM: CS4020. Leg. G.M. Dirkse RIN 005828. The holotype is preserved at U (isotype at
L). TYPE: (holotype, U not seen).
DISTRIBUTION: ESP (A.C. Bouman 91169, NYl).
15. Grimmia decipiens (Schultz) Lindb., in Hartm., Handb. Skand. Fl. ed. 8: 386. 1861.
Trichostomum decipiens Schultz, Prodr. Fl. Starg. Suppl.: 70. 1819. Dryptodon schultzii Brid.,
Bryol. Univ. 1(1): 199. 1826, nom. illeg., incl. typo spec. prior. Dryptodon grimmioides Brid.,
Bryol. Univ. 1(1): 199. 1826, nom. inval., pro synon. Grimmia schultzii Huebener, Muscol.
Germ.: 195. 1833, nom. illeg., incl. typo spec. prior. Dryptodon decipiens (Schultz) Loeske, Stud.
Morph. Syst. Laubm.: Ill. 1910. PROTOLOGUE: [Germany] Habitat ad saxa granitosa rarius. ad pagum Gevezien an der Landstrafse. Junio C. operculo. TYPE: ad saxa granitosa Duc[ato]
Megapol. Stargard (lectotype, designated by Geissler & Maier (1995: 499), Gl; isolectotype, HSOU).
Dryptodonfunalis var.longipilus Brid., Bryol. Univ. 1: 771. 1827. Dryptodon longipilus Brid., Bryol.
Univ. 1: 771. 1827, nom. inval., pro synon. PROTOLOGUE: [Germany] In Hercynia Suevica
passim ad rupes legit et communicavit D. Braun. In monte Vogeso Mougeot et Nestler etiam
WORLD SYNOPSIS OF GR/MMIA
25
invenerunt. TYPE: In Hercynia Suevica ret Vogesis ad saxa madida [illegible]]. Braun 1825
(lectotype, here designated, B!).
DISTRIBUTION: ARM (B. Dildaryan s.n., LE); BEL (C Delogne s.n., FH); CHE (Schlephacke
s.n., JE); DEU (G. Roth s.n., JE); ESP (J.L. Rupidera 29, SALA); FRA (F.G. Kneiff & CP. W
Maercker s.n., FH); GBR (Jones s.n., JE); GRC (R. Dull 13, GOET); IRL (H. W Lett s.n., TCD); ITA
(E. Levier s.n., NY); MAR (M Brugues s.n., BCB); MKD (T. Herzog 570A, JE); NOR (H. Winter s.n.,
JE); POL (K.G. Limpricht s.n., FH); PRT (C Sergio et al. 30.1a, MA); SWE (o.J. Hasslow s.n., NY);
TUR (K. Walther 2407, NY); UKR (L. Partyka s.n., MA).
16. Grimmia donniana Sm., Engl. Bot. 18: 1259. 1804. Grimmia sudetica Schwagr., Sp. Musc.
Frond. Suppl. 1(1): 87, tab. 24. 1811, nom. iIleg., incl. typo spec. prior. Weissia donniana (Sm.)
Poir., in Lam., Encycl. Suppl. 5: 502.1817. Grimmia donnii Gray, Nat. Arr. Brit. PI. 1: 728.1821,
nom. illeg., incl. typo spec. prior. Dryptodon donnianus (Sm.) Hartm., Handb. Skand. Fl. ed. 3:
270. 1838. Grimmia obtusa var. donniana (Sm.) Hartm., Handb. Skand. Fl. ed. 5: 377. 1849, nom.
illeg., prior. ut spec. Grimmia donii Sm. ex Lindb., Musci Scand.: 30. 1879, nom. illeg. Grimmia
donniana var. eudonniana Loeske, Laubm. Eur. Part I: 9l. 1913, nom. inval. Guembelia donniana
(Sm.) Loeske, Laubm. Eur. Part I: 90. 1913, nom. inval., pro synon. Grimmia donniana f. sudetica
(Schkuhr) Loeske, Biblioth. Bot. 101: 113. 1930, nom. illeg., non Chal. 1882. Grimmia donniana
subsp. eudonniana Giacom., Atti 1st. Bot. Lab. Crittog. Univ. Pavia, ser. 5, 4: 22l. 1947, nom.
inval. PROTOLOGUE: [United Kingdom] George Donn [... ] gathered it on large stones near a
water-fall on a mountain in Angusshire, 18 miles to the north of Forfar. Mr. Turner has also found
it on old pales at Beddgelart, North Wales, and from his specimen we have delineated it. TYPE:
North Wales, pales at Beddgelart. July 1802. D. Turner (lectotype, designated by Munoz (l998b:
378), BM!).
Grimmia sudetica Spreng. ex Schkuhr, Deutschl. Krypt. Gew. Heft 2: 48, tab. 22. 181l.
PROTOLOGUE: [Poland/Czech Republic border] Ludwig kryptog. Gewachse, cent. 1, n. 17.
Habitat in saxis Sudet. TYPE: Ludwig Kryptogamische Gewasche der Riesengebirgen.
Laubmoose [handwritten, not the original label of Ludwig's exsiccata] (lectotype, designated by
Munoz (1998b: 378-379), M!; isolectotype, JE!).
Synonymized by Munoz (1998b: 378).
Grimmia obtusa Schwagr., Sp. Musc. Frond. Suppl. 1(1): 88, tab. 25. 1811, nom. illeg., non Brid.
180l. Grimmia donniana var. obtusa Steud., Nomencl. Bot. 2: 189. 1824. Dryptodon erostris
Hartm., Handb. Skand. Fl. ed. 4: 374. 1843, nom. illeg., incl. typo spec. prior. PROTOLOGUE:
[Austria] Legi in Alpe Carinthiae Breenkogel, summo monti Glockner vicina, Augusto
fructificantem. TYPE: Glockner. Kaulfuss (holotype, G!; isotype, G!).
Synonymized by Deguchi (1978: 182).
Grimmia bohemica Schkuhr ex Steud., Nomencl. Bot. 2: 189 [also 188]. 1824, nom. inval., pro synon.
Grimmia donniana var. bohemica Schkuhr ex Steud., Nomencl. Bot. 2: 189. 1824, nom. nud.
Grimmia donniana var. bohemica Schkuhr ex Brid., Bryol. Univ. 1(1): 176. 1826.
PROTOLOGUE: In sudetis. Ludw[ig, C.G.]. TYPE: [Poland] Ludwigius dedit 1814. Circa dem
Schneekoppe Sudetum (lectotype, designated by Munoz (1998b: 379), B!).
Synonymized by Munoz (1998b: 379).
26
MuNoz AND PANDO
Grimmia obtusa var. elongata Nees & Homsch., Bryol. Germ. 2(1): 137, tab. XX fig. lIb. 1827.
Grimmia obtusa var. mammilaris Huebener, Muscol. Germ.: 177. 1833, nom. illeg., incl. typo var.
prior. Grimmia donniana var. elongata (Nees & Homsch.) Wilson, Bryol. Brit.: 159. 1855.
Grimmia donniana f. elongata (Nees & Homsch.) Loeske, Biblioth. Bot. 101: 113. 1930.
PROTOLOGUE: [Austria] Wachst am Wege von Gastein nach Bockstein auf Steinen, die,
nachdem man das Waldchen hinter sich hat, an der Acha liegen (Funck! Laurer! Mielichhofer!
Hsch.), dem Wege zum Rathhausberg, wenn man die Alpenregion betritt, links auf Steinen
(Funck! Laurer! Hsch.), auf den Judenburger Alpen (Hrsch. Willdenow!), auf dem Felbertauem,
dem Wildbachkogel im Oezthal und dem Timpeljoch (Funck!). TYPE: Stein. Alp Hsch. Ex herb.
Hsch. (lectotype, here designated, JE!; syntypes ["zwischen Gastein und Bockstein, Funck".
Cryptogamische Gewachse besonders des Fichtelgebirges, fasc. 22, n° 450, 1815], H!, JE!, RO!).
Grimmia nivalis Liebm. ex Besch., Mexic. PI. 1: 19. 1872, nom. nud. ORIGINAL MATERIAL:
[Mexico] In monte Orizabensi, alt. 14,000 p. (Liebmann in herb. Schimper). ORIGINAL
SPECIMEN: Pico d'Orizaba 14,000'. Liebmann leg. 42 (BM!).
Grimmia donniana f. vulgaris Chal., Pamietn. Fizyogr. 2: 56. 1882. Grimmia donniana f. longiseta
Vilh., Vestn. Knil. Cesk6 Spolecn. Nauk, Tf. Mat.-Pili. 1924: 17. 1924, nom. illeg., incl. typo f.
prior. PROTOLOGUE: [Poland] Zakopane: Chramc6wki; wal kamienny wzdluz Bystr6j Wody;
obficie. Krup6wki; takii; wal. Dolina Malego Kolbachu wprost Lornnicy. Brzegi Morskiego Oka.
Pod Polskim Grzebienem od strony FeIki. TYPE: Dol. Malego Kolbachu 1.08.1878 [T.
ChalubiIiski] (lectotype, here designated, ZAMV!).
Grimmia donniana f. cylindrica Podp., Sbom. Klubu Pfir. Bme 5: 14. 1923. PROTOLOGUE: [Czech
Republic] In summo monte HokSar m. Sudetorum c.fr. 1911 VIII. colI. am. 1. Schenk. TYPE:
Gesenke: Hockschar. 1911 VIII leg. F. Schenk (lectotype, here designated, PRJ).
DISTRIBUTION: AUT (v. Schiffner s.n., BP); BOL (M. Lewis 83-3243, F); CHE (1. Schleicher
s.n., Z); CRN (M. Zang 669, ALTA); CZE (1. VelenovskJ s.n., FH); DEU (H.M.C Solms-Laubach
s.n., GOET); ECU (P.l. Grubb 2501B, FH); FRA (N.J. Boulay s.n., FH); GBR (D.G. Long & 1.A.
Paton 14015, MA); GEO (J. Abramov s.n., LE); GRL (N. Hartz s.n., NY); HUN (L. Vajda s.n., BP);
IRL (J. Carroll s.n., BM); ISL (B. J6hannsson 10828, ICEL); ITA (A. Carestia s.n., RO); JPN (E.
Ishiba 1331, H-BR); MEX (A.J. Sharp 4761a, TENN); NOR (C Kaurin s.n., TRH); POL (R. Ochyra
s.n., BCB); ROM (K. Demeter s.n., BP); SFE (I. V Chernyadjeva 37, LE); SIB (N. Zolotukhin s.n.,
MA); SVK (A. Boros s.n., BP); SWE (H. Smith s.n., TRH); UKR (L. Partyka s.n., MA); USA (F.1.
Hermann 19583, MO).
17. Grimmia elatior Bruch ex Bals.-Criv. & De Not., Mem. Reale Accad. Sci. Torino 40: 340. 1838.
Trichostomum incurvum Hoppe & Homsch., Flora 2: 89. 1819. Dryptodon incurvus (Hoppe &
Homsch.) Brid., Bryol. Univ. 1(1): 194. 1826. Racomitrium incurvum (Hoppe & Homsch.)
Huebener, Muscol. Germ.: 201. 1830. Grimmiafunalis var. incurva (Hoppe & Homsch.) Hampe,
Flora 20: 282. 1837. Dryptodonfunalis var. incurvus (Hoppe & Homsch.) HartIn., Handb. Skand.
Fl. ed. 3: 270. 1838. Grimmia funalis var. robusta De Not., Syllab. Musc.: 251. 1838. Grimmia
funalis subsp. elatior (Bals.-Criv. & De Not.) HartIn., Handb. Skand. Fl. ed. 5: 376. 1849.
Dryptodon elatior (Bals.-Criv. & De Not.) Loeske, Stud. Morph. Syst. Laubm.: 111. 1910.
Dryptodon funalis var. robustus De Not. ex H. Moller, Ark. Bot. 26A(2): 105. 1934 ["robusta"],
nom. inval. Dryptodonfunalis var. incurvatus HartIn. ex H. Moller, Ark. Bot. 26A(2): 135. 1934,
WORLD SYNOPSIS OF GRiMMIA
27
nom. inval., orthogr. err. PROTOLOGUE: [Austria] Auf kahlen Felsenblocken bey Heiligenblut,
mit vollkommenen Friichten im Juni, im Nafsfe1de und am Wege von Mattrey nach Deferreggen
ohne Friichte. TYPE: In rupibus zu Hgblut [Heiligenblut] (lectotype, designated by Cao & Vitt
(1986: 201), GZU!).
Grimmia elatior f. alpigena J.E. Zetterst., Grimmiae et Andreaeae exsiccatae n° 15 a-b. 1861, nom.
nud. ORIGINAL MATERIAL: [Norway] In saxis juxta Kongsvold alpium Dovrensium.
27/7/1858. J.E. Zetterstedt. [Zetterstedt, Grimmiae et Andreaeae exsiccatae, n° 15a). In saxis juxta
fluvium supra Rodsheim paroecire Lorn, Norvegire medire. 30/6/1858. lE. Zetterstedt. [ibidem, n°
15b](H!).
Grimmia elatior f. infra-alpina lE. Zetterst., Grimmiae et Andreaeae exsiccatae n° 14a-c. 1861, nom.
nud. ORIGINAL MATERIAL: [Norway] In saxis montis Egebergjuxta Christianiam. 15/6/1858.
J.E. Zetterstedt. [Zetterstedt, Grimmiae et Andreaeae exsiccatae, n° 14a). In saxis littoreis juxta
Stocklycke montis Omberg, Ostrogothire. Jun. 1855. lE. Zetterstedt. [ibidem, n° 14b]. In rupibus
juxta Gottsunda prope Upsaliam. 7/5//1858. J.E. Zetterstedt. [ibidem, n° 14c] (H!).
Racomitrium papillosum Wamst., Hedwigia 20: 166. 1881. Grimmia papillosa (Wamst.) Kindb., Bih.
Kongl. Svenska Vetensk.-Akad. Handl. 7(9): 110. 1883. Grimmia elatior subsp. papillosa
(Wamst.) Kindb., Eur. N. Amer. Bryin. 2: 222. 1898. PROTOLOGUE: Norwegen, Dovre,
Kongsvold von N.C. Kindberg im Juli 1879 entdeckt. TYPE: Dovre, f. om Sprenbacken. 14/7 79.
N.C. Kindberg (lectotype, here designated, S!; isolectotype, JE!).
Grimmia aspera Miill. Hal., Nuovo Giom. Bot. Ital., n.s. 4: 261. 1897. PROTOLOGUE: China
interior, provo Schen-si septentr., Hua-san, sterilis, Aug. 1895. TYPE: China interior, provincia
Schen-si septentr. Hua-san. 25 Aug. 1895 legit Rev. Jos. Giraldi, determ. C. Muller n. 1448
(lectotype, designated by Cao & Vitt (1986: 201), H; isolectotypes, BM, PC! 2 replicates, S).
Synonymized by Cao & Vitt (1986: 201).
Grimmia cognata Cardot & Ther., Bot. Gaz. 37: 368, tab. 19 fig. 1. 1904. PROTOLOGUE: [U.S.A.]
Colorado: along the Cogwheel Railway to Pike's Peak, alt. 2100-3000 m (J.M. Holzinger, 1896).
TYPE: Colorado [... ] along the Cogg Wheel Railway to Pike's Peak. June 4 and 5, 1896, by I.M.
Holzinger. Altitude, 7,000-10,000 feet (lectotype, here designated, PC!; isolectotype, H-BR!).
Grimmia grandis Kindb., Rev. Bryol. 32: 36. 1905. PROTOLOGUE: 2. Found by l Macoun 1904 in
British Columbia, 5200-7500 feet above the sea. TYPE: Canada, B.C. lake Louise, rocks. 6000 ft.
13/904. Macoun (holotype, S!; isotypes, FH!, S!).
Synonymized by Allen (1995: 161).
Grimmia elatior var. hystrix Podp., Vestn. Klubu Pfir. v Prostejove 8: 36. 1906. PROTOLOGUE:
[Czech Republic, Moravia] Na skalach Kotliny na Jeseniku ve vYsi 1200m [on the rocks of the
Kotlina in the Ash Mountains, 1200m]. TYPE: Kotlina na Jeseniku. 1904 VII leg. Dr. Podpera
(lectotype, here designated, PR!).
28
MuNoz AND PANDO
Grimmia madagassa Renauld & Cardot, Prodr. Fl. Bryol. Madagascar. Supplement. Essai Leucoloma:
103. 1909. PROTOLOGUE: Madagascar: plateau central (Rh. Mitten). TYPE: Herb. Mitten.
Central Madagascar (lectotype, here designated, PC!).
NOTE: No type material was found in NY.
Grimmia elatior f. epilosa H. Moller, Musci europaei et americani exsiccati ser. 44 n° 2163. 1932.
PROTOLOGUE and TYPE: Schweden: [Norrbottens:] Lule Lappmark, Jokkmokk, Akkatsforsen,
1. Juli 19211egit Hj. Moller (lectotype, here designated, S!; isolectotype, S!).
Grimmia elatior f. nigrescens H. Moller, Ark. Bot. 26A(2): 108. 1934. PROTOLOGUE: [Sweden]
Varrnland. Sunne. Skarped fro 1925 G. Aberg. TYPE: Vlirrnl. Sunne m. Skarped nich Bauna alp.
80 m. 7/9 1925. G. Aberg (lectotype, here designated, S!).
DISTRIBUTION: ARM (A.A. Fedorov s.n., LE); AUT (Laurer s.n., JE); CAN (R.R. Ireland
14985, G); CHE (V Schiffner s.n., FH); CHN (C Gao & T Cao 16556, ALTA); CZE (J. Podp&a s.n.,
H); DEU (Herzog s.n., FH); ESP (C Casas s.n., MA); FIN (R. Oldberg s.n., H); FRA (J.L. De Sloover
34438, MO); GBR (J. Fergusson s.n., NY); GEO (VF. Brotherus 240, H-SOL); GRL (K. Holmen 621665, NY); IND (CC Townsend 87/343, herb. Whittemore); ITA (E. Levier 656, NY); KAZ (B. Allen
10448, MO); KGZ (Y. Rakhmatulina s.n., LE); MDG (Anonymous s.n., PC); MNG (s. V Maksimovich
785, LE); NOR (E. Adlerz s.n., S); POL (K.G. Limpricht s.n., FH); SFE (OM Afonina s.n., LE); SIB
(L.V Bardunov s.n., NICH); SUN (VG. Onichenko 136/95A, NY); SVK (s. Lisowski s.n., H); SWE
(J.E. Zetterstedt s.n., H); USA (E.B. Bartram 17913, NY).
18. Grimmia elongata Kaulf., in Sturm, Deutschl. Fl. Abtheilung II, Cryptogamie 4(13): 24. 1812.
Dryptodon elongatus (Kaulf.) Hartm., Handb. Skand. Fl. ed. 3: 271. 1838. PROTOLOGUE:
[Austria] im August 1812 auf der Seethaler Alpe in Srenermarf in SteinriJ3en mit reifen Fruchten.
TYPE: In Styria alpibus Kaulfuss (lectotype, designated by Geissler & Maier (1995: 500), B not
seen).
Grimmiafuliginosa Schimp. ex Miill. Hal., Syn. Musc. Frond. 1: 778. 1849. Schistidiumfuliginosum
(Mull. Hal.) Ochyra, Fragm. Florist. Geobot. 43: 105. 1998. PROTOLOGUE: Mexico, in summo
Pico de Orizaba, alt. 14000': Liebmann. TYPE: Pic d'Orizaba 14000'. Liebmann leg. & det. Fbri.
1844 (lectotype, designated by Muiioz (1999a: 134), BM!; isotypes, BM!, PC-Besch!).
Synonymized by Munoz (1999a: 134).
Grimmia orientalis Wilson, Hooker's J. Bot. Kew Gard. Misc. 9: 323. 1857, nom. nud. ORIGINAL
MATERIAL: [India] 293. Sikkim-Himalaya temperata; Singalelah, alt. 11,000 ped., J.D.H. [J.D.
Hooker]. ORIGINAL SPECIMEN: n° 293 Sikkim: Singalelah (FH!).
Grimmia vulcanica Besch., Ann. Sci. Nat. Bot., ser. 6, 9: 351. 1878. PROTOLOGUE: La Reunion:
Cratere brillant, P. Lepervanche, 1878. TYPE: La Reunion. Cratere brillant. Lepervanche
(lectotype, here designated, BM!; isolectotypes, JE!, PC-Besch!).
WORLD SYNOPSIS OF GRIMMIA
29
Grimmia andreaeaeea Miill. Hal., Linnaea 43: 453. 1882. PROTOLOGUE: Argentinia subtropica,
Cuesta de Caldeira, 27. Majo 1873. [leg. P.G. Lorentz]. TYPE: Cuesta de Caldeira, 27.V.1873
(lectotype, designated by Munoz (1999a: 134), BM!; isolectotype, PC!).
Synonymized by Munoz (1999a: 134).
Grimmia montana var. epilosa Grav. ex Delogne, Ann. Soc. BeIge Microscop. 9: 21. 1885. Grimmia
montana f. epilosa (Delogne) Podp., Consp. Musc. Eur.: 281. 1954. Grimmia triehophylla var.
epilosa (Grav. ex Delogne) De Sloover, Bull. Jard. Bot. Etat 37: 445. 1967, nom. illeg., non
Warnst. 1897. PROTOLOGUE: Willerzie (Gravet). TYPE: Willerzie (prov. Narnur, Belgique) 14
September 1882. F. Gravet (lectotype, designated by Munoz (1999a: 134), PC!; isolectotypes, JE!,
PC!).
Synonymized by Munoz (1999a: 134).
Grimmia deealvata Cardot, Bull. Herb. Boissier, ser. 2, 8: 332. 1908. PROTOLOGUE: Japon: Ontake,
2500-3200 m. (n. 3319, 3321, 3326); Komagatake, 2500 m. (n, 3316, 3342). TYPE: Japon:
sommet du Kornagatake, 2300 m. Leg. Faurie n° 3316 (lectotype, designated by Deguchi (1978:
185), PC!; isolectotypes, H-BR!, NY!, PC!).
Synonymized by Deguchi (1978: 185).
Grimmia tolueensis Cardot, Rev. Bryol. 38: 1. 1911. PROTOLOGUE: [Mexico] Etat de Mexico:
volcan de Toluca (Hahn; Pringle, 1892, nO 27a). TYPE: C.G. Pringle Plantae mexicanae, 1892, n°
27a. State of Mexico, on rocks, volcano of Toluca, 1400 ft. 7-Sept. 1892 (lectotype, designated by
Munoz (1999a: 134), PC!; isolectotypes, FH!, JE!, PC! 2 replicates, TENN!).
Synonymized with G. ineurva by Robinson & Hermann (1964: 173), but see Munoz (1999a: 134).
Grimmia ineurva var. rhaetiea J.1. Amann, Fl. Mouss. Suisse 2: 133. 1918. Grimmia ineurva f.
rhaetiea (J.J. Amann) Podp., Consp. Musc. Eur.: 284. 1954. PROTOLOGUE: [Switzerland]
Graubunden: Scaletta, 2500 m (BH 89.1.4) (Amann). TYPE: Graubunden: Scalertapass, 2500,
1888. Leg. Amann (BH 87.1.4) (holotype, Z!).
Grimmia elongata var. epilosa 1.1. Amann, Bull. Soc. Vaud. Sci. Nat. 54: 45. 1921, nom. illeg., non
Matousch. 1903. PROTOLOGUE: Italie, Province de Come, Monte Legnone, 1700 m. environ
(Artaria) (BH. 90.5.21). TYPE: Italia. Como. Mt. Legnone, 1700 m, 29 VII 1897. ColI. F. A.
Artaria (BH. 90.5.21) (holotype, Z!).
DISTRIBUTION: ARG (P. G. Lorentz s.n., BM); AUT (R.B. Pierrot 62115, PC); BEL (F Gravet
s.n., PC); BOL (M Lewis 83-3317, F); BRA (P. Dusen 611, H-BR); CHE (M Anzi 128, FH); CRN (K.
Lang 598, ALTA); COL (A.M Cleef 2449, COL); CZE (z. Pilous s.n., H); DEU (Herzog s.n., JE);
ESP (M Aeon s.n., BCB); FRA (R. Potier de la Varde s.n., PC); GBR (C.H. Binstead s.n., H); GEO
(V.F Brotherus 304, H-SOL); GTM (J.A. Steyermark 35548, FH); IND (J.D. Hooker 293, FH); ISL
(B. Johannsson 33583, ICEL); ITA (FA. Artaria s.n., Z); JPN (E. Ishiba 1228, H-BR); LSO (J. van
Rooy 3402, NY); MEX (A.J. Sharp et al. 1514, TENN); NOR (J.E. Zetterstedt s.n., H); PRT (H.
Persson s.n., PC); REU (P. Lepervanehe s.n., JE); ROM (M. Peterji s.n., BP); SIB (H. Nilsson-Ehle
s.n., H); SVK (z. Pilous s.n., H); SWE (0. Martensson s.n., MO); TWN (J. Hyvonen & s. Stenroos
3840, MA); UGA (G.H.s. Wood 1107a, NY); VEN (A. Schiifer- Verwimp & l. Verwimp 12280a, MA);
ZAF (J. van Rooy 1114, NY).
30
MuNoz AND PANDO
19. Grimmia funalis (Schwagr.) Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 119, tab. 247. 1845.
Trichostomum fimale Schwagr., Sp. Musc. Frond. Suppl. 1(1): 150, tab. 37. 1811. Dryptodon
funalis (Schwagr.) Brid., Bryol. Univ. 1(1): 193. 1826. Trichostomum patens var. funale
(Schwagr.) Brid., Bryol. Univ. 1(1): 193. 1826, nom. inval., pro synon. Trichostomum patens var.
piliferum Hook. & Taylor, Muscol. Brit. ed. 2: 105. 1827. Racomitrium funale (Schwagr.)
Huebener, Muscol. Germ.: 200. 1830. Grimmia funalis subsp. eufunalis Giacom., Atti 1st. Bot.
Lab. Crittog. Univ. Pavia, ser. 5,4: 222. 1947, nom. inval. Grimmia funalis var. eufimalis Podp.,
Consp. Musc. Eur.: 286. 1954, nom. inval. PROTOLOGUE: [Poland/Czech Republic border]
Legit in suba1pinis Sudeticis C. Ludwig. TYPE: Urn die Schneekoppe L[udwi]g ex parte
(lectotype, designated by Geissler & Maier (1995: 500), G!).
Grimmia spiralis Hook., in Grev., Scott. Crypt. Fl. 4: 203. 1826. Dryptodon spiralis (Hook.) Brid.,
Bryol. Univ. 1: 771. 1827. Grimmia cernua var. spiralis (Hook.) Nees & Hornsch., Bryol. Germ.
2(1): 177, tab. XXIV fig. 23a. 1827. Grimmia incurva var. spiralis (Hook.) Huebener, Muscol.
Germ.: 191. 1833. Grimmia uncinata subsp. spiralis (Hook.) Hampe, Flora 20: 282. 1837.
PROTOLOGUE: [United Kingdom] On rocks in the Scottish Highlands, at a considerable
elevation. Throughout the whole or the Breadalbane range of mountains; in fruit upon Craig
Challeach, Dr. Hooker, Mr. Walker Arnott, and Dr. Greville. Mountains of Clova, Mr.
Drummond. TYPE: Craig Challeach July 1823 (Arnott) (lectotype, here designated, BM!;
iso1ectotypes, PC! 3 replicates).
Grimmia incurva var. jlagellaris Brid., Bryol. Univ. 1(1): 186. 1826. PROTOLOGUE: [Austria] In
Heiligenbluter-Tauern Carinthiae superioris ad rupes sed semper absque fructu lectam rnisit dar.
Hornschuchius. TYPE: Heiligenbluter Tauern. Hornsch. (lectotype, here designated, JE!).
Gymnostomum spirale Hartrn., Handb. Skand. Fl. ed. 3: 265. 1838. PROTOLOGUE: [Norway]
Fjellklipp. r. (N. Dovre: Blytt). TYPE: Dovre Kongsvold. B1ytt (lectotype, here designated, S!;
isolectotype, S!).
Grimmia funalis f. epilifera J.E. Zetterst., Revis. Grimm. Scand.: 74. 1861. Grimmia spiralis var.
epilifera (J.E. Zetterst.) Hartrn., Handb. Skand. Fl. ed. 9, 2: 56. 1864. Grimmia imberbis Kindb.,
Bot. Not. 1882: 186. 1882, nom. illeg., non Mull. Hal. 1849. Grimmia calvescens Kindb.,
Christiania Vidensk.-Selsk. Forh. 1888(6): 19. 1888, nom. nov. Grimmia funalis var. epilifera
(lE. Zetterst.) Broth. & Saehin, Acta Soc. Fauna Fl. Fenn. 6(4): 80. 1890. Grimmia funalis var.
epilosa lE. Zetterst. ex Paris, Index Bryol. 2: 526. 1895, nom. inval. Grimmia tortifolia var.
calvescens (Kindb.) Kindb., in Roll, Hedwigia 35: 64. 1896. Grimmia tortifolia subsp. calvescens
(Kindb.) Kindb., Eur. N. Amer. Bryin. 2: 217. 1898. Grimmiafunalis subsp. calvescens (Kindb.)
J.J. Amann, Fl. Mouss. Suisse 2: 137. 1918. Grimmia bauri Miill. Hal. ex H. Moller, Ark. Bot.
26A(2): 65. 1934, nom. inval., pro synon. Grimmia funalis var. calvescens (Kindb.) H. Moller,
Ark. Bot. 26A(2): 65. 1934. PROTOLOGUE: Habitat in Norvegia media multis in locis, prrecipue
copiose in Dovre prope Kongsvold (ubi Kindberg perpauca specimina fructifera legit) et in monte
Lauvhoen paroecire Lorn; in Valders et in Voss earn legit Blytt. TYPE: Norvegia: Lorn, Lauvh0en.
7/7/1858. lE. Zetterstedt (lectotype, here designated, S!).
Grimmia ryanii Limpr. ex Bryhn, Nyt Mag. Naturvidensk. 34: 73, figs. 1-13. 1892 ["Ryani"].
Grimmia tortifolia subsp. ryanii (Bryhn) Kindb., Eur. N. Amer. Bryin. 2: 218. 1898. Grimmia
funalis var. ryanii (Bryhn) Loeske, Biblioth. Bot. 101: 153. 1930. PROTOLOGUE: [Norway]
August 1890 af Fabrikbestyrer Ryan ved Leirungsboden paa Sudsiden af Gjendin i Jotunfjeldene
WORLD SYNOPSIS OF GRiMMIA
31
[... ]. TYPE: Norvegia, Christians Amt. Jotunheimen, prope Leirungsboden in rupes siccis. Alt.
metro 1150. Aug. 1890. Chr. Kaurin & E. Ryan (lectotype, here designated, NY!; isolectotypes,
JE!, PC!; syntypes ["Jotunfjeldene"], NY! 4 specimens).
Grimmia brittoniae R.S. Williams, Bull. Torrey Bot. Club 27: 316, pI. 19. 1900. PROTOLOGUE:
[U.S.A.] Growing on shaded perpendicular walls, partly calcareous, in rather dry places. Collected
for several seasons in Bad Rock Canon, Flathead River, Mont. [Montana]. TYPE: n° 223. April 2,
1894. Montana. Columbia Falls. Collected by Robert Statham Williams (lectotype, designated by
Munoz (2000c), NY!; isolectotypes, NY! 3 replicates).
Synonymized by Munoz (2000c).
Grimmia torquata var. mutica Herzog ex Loeske, Laubm. Eur. Part I: 159. 1913. Grimmia torquata f.
mutica (Loeske) Loeske, Biblioth. Bot. 101: 159. 1930. PROTOLOGUE: [Switzerland] am
"Seilhenker" des Medjekopfs in der Silvretta, +/- 2400 m. TYPE: Fl. Graubunden. An Gneisfelsen
des Medjekopf(am Seilhenker). ca 2400 m. August, 1899 (lectotype, here designated, JE!).
Grimmiafunalis var. leptodictyon Herzog, Kryptog. Forsch. 1: 292. 1919. PROTOLOGUE: [Romania]
An schattig-feuchten Felsen einer Schlucht zwischen Poiana lunga und Fruntu, ca. 1000 m; an
Felsen unter der Moscovulscharte, ca. 2000 m. TYPE: Transsilvan. Alpen. An Felsen einer
Bachschlucht zwischen Poiana lunga u. Fruntu. ca 1000 m. November, 16 [1916]. T. Herzog
(lectotype, here designated, JE!; syntype, ["Moscovulscharte, Herzog s.n."] JE!).
Grimmia ryanii f. tenuis 1.J. Amann, in Bauer, Musci Europaei et Americani Exsiccati, Ser. 43, n°
2215. 1931. Grimmia funalis f. tenuis (J.J. Amann) Podp., Consp. Musc. Eur.: 286. 1954.
PROTOLOGUE and TYPE: Schweiz: Graubunden [... ] 10. September 1930 legit 1. Amann
(lectotype, here designated, MO!; isolectotypes, JE!, NY!).
DISTRIBUTION: ARM (V Manakyan 4907a, ERE); AUT (J.-P. Frahm s.n., MA); AZE (L.
Lyubarkaya s.n., LE); BGR (L. Meinunger 14486, JE); CAN (J.A. Hancock 1238, NFLD); CHE (P.
Culmann s.n., H); CHN (Jia Yu J02381, MO); CZE (Z Pilous s.n., H); DEU (L. Meinunger 4644, JE);
DZA (L. Loeske s.n., JE); ESP (C Aedo s.n., MA); FIN (VF. Brotherus s.n., JE); FRA (T. Husnot s.n.,
FH); FRO (R. Fagersten & M. Haapasaari 327, H); GBR (D.G. Long 6786, MA); GEO (A.H.
Brotherus & VF. Brotherus 41, H-SOL); GRL (K. Holmen 7436, H); HUN (A. Boros s.n., H); ISL (B.
J6hannsson 11879, ICEL); IRL (H. W Lett s.n., TCD); ITA (M. Fleischer s.n., FH); MNG (T. Tsegmed
331, NY); NOR (C Kaurin & E. Ryan s.n., NY); POL (R. Ochyra s.n., H); ROM (A. Boros s.n., BP);
SFE (B.C Tan 97-234, FH); SIB (M. Ignatov 0/469A, MO); SUN (VG. Onichenko 136/95, NY); SVK
(H. Zemanova s.n., H); SWE (N. Hakelier s.n., MO); TUR (Davis & Hedge D.32330, MA); UKR
(K.G. Ulychna s.n., H); USA (WB. Schofield 12025, FH); YUG (M. Sabovljevi6 s.n., MA).
20. Grimmia fuscolutea Hook., Musci Exot. 1: 63. 1818. PROTOLOGUE: In frigidis Tobeccae et
Islahuacae, altitudine 1330 hexapod. (Regnum Mexicanum). Humboldt et Bonpland. TYPE: N°
2673. S. Am. Humboldt 50 (lectotype, here designated, BM!).
Grimmia apiculata Homsch., Flora 1(19): 329. 1818, nom. nud. Grimmia affinis Homsch., Flora 2(1):
85. 1819. Grimmia apiculata Homsch., Flora 2(2): 442. 1819, nom. illeg., incI. typo spec. prior.
Grimmia pulvinata var. apiculata (Homsch.) Huebener, MuscoI. Germ.: 710. 1833, nom. inval.,
orthogr. err. Dryptodon apiculatus (Homsch.) Hartm., Handb. Skand. Fl. ed. 3: 270. 1838.
32
MuNoz AND PANDO
PROTOLOGUE: [Austria] der Messerlingwand auf dem Windischmattreyer Tauem. TYPE:
Windischmattreyer Tauem. Hornschuch missit (lectotype, designated by Deguchi (1978: 165), B!;
iso1ectotype, H-SOLl).
Synonymized by Crum (1994: 403).
Grimmia abyssinica Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 109. 1845, nom. nud. Guembelia
abyssinica Hampe, Bot. Zeitung (Berlin) 4(8): 125. 1846, nom. nud. Guembelia abyssinica Mull.
Hal., Syn. Musc. Frond. 1: 772. 1849. Grimmia abyssinica (Mull. Hal.) Mitt., 1. Linn. Soc., Bot. 7:
152. 1863. PROTOLOGUE: [Ethiopia] Abyssinia, monte alpestri Silke, 18. Febr. 1840: W.
Schimper. TYPE: In siccis ad rupes inde a limitibus arbomm [sic] et fruticum usque ad cacumen
montis Silke. d. 18.Febr.1840. U.i.[Unio Itineraria] 1842 (lectotype, here designated, MO!;
isolectotypes: H-BR!, FH!, GOET!, JE!, NY!, PC!).
Grimmia holleri Molendo, Flora 50: 39. 1867. Grimmia apiculata var. holleri (Molendo) Husn.,
Muscol. Gall.: 133. 1887. Grimmia apiculata f. holleri (Molendo) Loeske, Laubm. Eur. Part I:
116. 1913. Grimmia apiculata subsp. holleri (Molendo) lJ. Amann, Fl. Mouss. Suisse 2: 132.
1918. PROTOLOGUE: [Austria. Salzburg: Alpen. .. ] bei 5900' bis 6000' [... ] (von der Stanz im
Raufis-Fucher Scheiderlicken) 1864. TYPE: Rauris, in Menge am Rande d. Baurnregion 6-6200',
Glirnrnerschiefer. 1. [legit] Holler. c. Mdo. (holotype, M!; isotype, M!).
Grimmia flexicaulis Mull. Hal., Linnaea 43: 454. 1882. PROTOLOGUE: Argentinia subtropica,
Cuesta de Pinos, i.e., in descensu ab excelsa Puna in convallem Tarija Boliviae, 27. Martio 1873,
[... ]; Catamarca, supra Yakutula in convalle Granadillas in alpinis Vayas altas [sic], alt. 9-10,000
ped., in granitic is, initio Februarii 1872. [leg. P.G. Lorentz]. TYPE: Cuesta de Pinos: [illegible]
Puna [illegible] Tarija in Bolivia. 27/3/73. Lorentz (lectotype, designated by Deguchi (1987: 22),
JE!; isolectotype, BM!; syntypes" JE! ["Catamarca: Yakatula, Granadillas und Vayas"] 2
replicates).
Synonymized by Munoz (1999a: 136).
Grimmia quatricruris Miill. Hal., Linnaea 43: 462. 1882. PROTOLOGUE and TYPE: Argentinia
subtropica, Cuesta de Pinos in descensu ab alta Puna in convallem Tarija Boliviae, 27. Martio
1873. [leg. P.G. Lorentz] (lectotype, designated by Munoz (1 999a: 136), BM!; isolectotype, PC!).
Synonymized by Munoz (1999a: 136).
Grimmia unicruris Mull. Hal., Linnaea 43: 463. 1882. PROTOLOGUE: Patria. Argentinia subtropica,
Cuesta de Pinos in descensu ab alta Puna in convallem Tarija Boliviae, 27. Martio 1873,[ .. .].
Quoque in Nevado de Castillo prope Salta; alpina, [... ]. [leg. P.G. Lorentz]. TYPE: Cuesta de
Pinos, 27.111.1873 (lectotype, designated by Munoz (l999a: 136, 138), BM!).
Synonymized by Munoz (l999a: 136).
Grimmia trichophylloidea Schimp. ex Miill. Hal., Nuovo Giom. Bot. Ital., n.s. 4: 127. 1897.
PROTOLOGUE: Bolivia, Llachissaxi, regione alpina 4200 m; Sorata, pr. Vaucuiri in scopulosis,
4500 m, Aprili 1860: Mandon legit. TYPE: Plantae Andium Boliviensium. Provo Larecaja,
vicinitis Sorata, prope Vancuiri, in scopulosis. Alt. Reg. alpina 4500. April 1860. G. Mandon n°
1633 (lectotype, designated by Munoz (1999a: 138), NYl; isolectotypes, Fl, PCl).
Synonymized by Munoz (l999a: 138).
WORLD SYNOPSIS OF GRlMMIA
33
Grimmia glyphomitrioides Broth. ex Paris, Index Bryol. Suppl.: 173. 1900, nom. nud. ORIGINAL
MATERIAL: Afr.: Cameroon. ORIGINAL SPECIMEN: Buea 3500 m (Cameroun). II 91. Preuss
859 (PC!).
Grimmiafastigiata Cardot, Bull. Herb. Boissier, ser. 2, 5: 1003. 1905. PROTOLOGUE: [Argentina]
Terre-de-Feu: Ushuaia, mont Martial (Typus in herb. Cardot). TYPE: Svenska Siidpolarexpeditionen 1901-03 n° 70. Terre-de-Feu: Ushuaia, mt. Martial, D. 1113/1902 alt 1275 m. Carl
Skottsberg. Det. J. Cardot (lectotype, designated by Munoz (1999a: 138), PC!; isolectotypes, HBR!, MEL-29537).
Synonymized by Munoz (1999a: 138).
Grimmia afroincurva Broth., Wiss. Erg. Deut. Zentr.-Afr. Exped., Bot. 2: 148, tab. 13 fig. 17. 1910.
PROTOLOGUE: [Zaire, District des Lacs Edouard et Kivu, Birunga Mountains]Vulkan-Gebiet:
Karisirnbi, Gipfelkegel, ca. 4000 m. ii. M. (reichlich fruchtend irn Sept. 1907 - n. 2085). TYPE:
Afr. or., Vulkan Gebiet, Karisirnbi, 4000 m, 9/1907. Leg. J. Milbraed (lectotype, here designated,
H-BR!).
Grimmia micropyxis Broth., Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1, 133(10):
567. 1924. PROTOLOGUE: [China] Provo Setschwan austro-occid.: Cum praecedente [In montis
Tschahungnyotscha trans flumen Yalung ad septentr. oppidi Yenyiien, 28° 15', regione alpina, ad
ropes schistosas; ca. 4150-4300 m] (Nr. 2671). TYPE: Provo Setschwan austro-occid.: In montis
Tschahungnyotscha supra vicurn Ngaitschekou trans flumen Yalung ad septentr. oppidi Yenyiien
ca. 28°15' siti regione alpina, ad ropes. Substr. schistaceo; alt. s.m. ca. 4150-4300 m. Leg.
27.V.1914 Dr. Heinr. Frh. v. Handel-Mazzetti (Diar. Nr. 498) n° 2671 (lectotype, here designated,
W!; isolectotypes, JE!, PC!).
.
Synonymized by Deguchi (1978: 165).
Grimmia subdonniana Nog., Candollea 19: 175. 1964. PROTOLOGUE: Nepal: Moraine, near Lobuje
(27°57',86°49'), alt. 4870 m, April 2, 1952, Zimmermann 301-type, 304; moraine, near the base
camp at Lobuje, alt. 5000 m, April 24, 1952, Zimmermann 325; near the base camp, in the
direction of Lobuje, alt. 5100 m, May 31,1952, Zimmermann 558. TYPE: Nepal. Moraine pres de
Lobuje, 4870 m. Apr. 2, 1952, colI. A. Zimmermann 301a. Ex Herb. Dr. A. Noguchi no. 68265
(holotype, NICH-365224!; paratype, G! [Zimmermann 558]).
Synonymized by Deguchi (1978: 165).
Grimmia sanii Greven, Bryologist 99: 429, fig. 2. 1996. PROTO LOGUE: South Africa. Drakensberg,
Mount aux Sources, Crow's nest, alt. 13050 m [sic], Schelpe 2115 (holotype, PRE; isotypes, MO
2550366, MO 3679148). TYPE: South Africa. Natal: Drakensberg, Crow's Nest. Mount aux
Sources. Alt. 10.000'. July 1947. E. A. Schelpe 2115 (isotypes, MO! 2 replicates).
Synonymized by Munoz (1999a: 138).
DISTRIBUTION: ARG (l.M Lamb 5566, FH); ATA (R.l. Lewis-Smith 3677, MA); AUT (CF.
Hornschuch s.n., B); BOL (M Lewis 79-1824, MO); CAN (J.M Macoun s.n., S); CHE (T Herzog s.n.,
JE); CRN (C Gao 1080, ALTA); CMR (P.R. Preuss 859a, PC); COG (H. Humbert 7430, PC); ECU
(R. Benoist 3458, F); ETH (G.H. W Schimper s.n., H-BR); IND (J.F. Duthie s.n., H-BR); ITA (CC
Townsend 921551, MA); KEN (0. Hedberg 950F, UPS); LSO (J. van Rooy 3475, F); MEX (.4.
Cardenas 3512, ALTA); NOR (N Hakelier s.n., MO); NPL (A. Zimmermann 304, G); PER (TG.
Tutin 196, BM); REU (P. Lepervanche s.n., PC); SIB (M Ignatov 01476, MO); SVK (J. Smarda s.n.,
34
MuNoz AND PANDO
BP); TWN (J. Hyv6nen & S. Stenroos 3892, MA); TZA (0. Hedberg 2324, PC); USA (EL Little Jr.
10401, MO); VEN (A. Schiifer-Verwimp & I. Verwimp 12280, FLAS); ZAF (E.A. Schelpe 2115, MO);
ZAR (L. Hauman 888, JE).
21. Grimmia hamulosa Lesq., Mem. Calif. Acad. Sci. 1: 14. 1868. PROTOLOGUE: [U.S.A.
California] Hab. gravelly soil, Mt. Dana, 10,000 ft., Bol. [H.N. Bolander]. TYPE: Gravelly soil.
Mt. Dana, California (Bolander) (lectotype, designated by Munoz (2000c), NY!; isolectotypes,
FH! 2 replicates).
Grimmia brevirostris R.S. Williams, Bryologist 23: 52, pI. 3. 1920. PROTOLOGUE: [U.S.A.]
Mountains near Buck's Valley, Plumas Co., Cal. [California], at 5600 ft. [sic], on granite.
Collected by J.B. Leiberg, July 1900, no. 5445. TYPE: California [... ] on granite rocks, mountains
adjacent to Buck's valley, Plumas Co. Alt. 6500 ft. Collected by John B. Leiberg, 1900 July
(lectotype, designated by Munoz (2000c), NY!).
Synonymized by Munoz (2000c).
DISTRIBUTION: USA (A.T Whittemore 3042, MO).
22. Grimmia handelii Broth., Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1, 133(10):
567. 1924. PROTOLOGUE: [China] Provo Setschwan austro-occid.: In montis
Tschahungnyotscha trans flumen Yalung ad septentr. oppidi Yenyiien, 28° 15', regione alpina, ad
ropes schistosas; ca. 4150-4300 m (Nr. 2667). TYPE: Provo Setschwan austro-occid.: In montis
Tschanhungnyotscha supra vicum Ngaitschekou trans flumen Yalung ad septentr. oppidi Yenyiien
ca. 28°15' siti regione alpina, ad ropes. Substr. schistaceo; alt. s.m. ca. 4150-4300 m. Leg.
27.V.1914 Dr. Heinr. Frh. v. Handel-Mazzetti (Diar. Nr. 498) (holotype, H-BR!; isotype, PC!).
DISTRIBUTION: CRN (H.F. v. Handel-Mazzetti 2667, H-BR).
23. Grimmia hartmanii Schimp., Syn. Musc. Eur.: 214. 1860. Grimmia sphaerocarpa Stirt. ex
Braithw., Brit. Moss Fl. 2: 20. 1888, nom. inval., pro synon. Dryptodon hartmanii (Schimp.)
Limpr., Laubm. Deutschl. 1: 789. 1889. Grimmia sphaerophora Stirt. ex Braithw., Brit. Moss Fl.
3: 263. 1905, nom. inval., orthogr. err. Grimmia hartmanii var. septentrionalis Loeske, Laubm.
Eur. Part I: 144. 1913, nom. illeg., incl. typo spec. Grimmia hartmanii subsp. vulgaris Loeske,
Biblioth. Bot. 101: 177. 1930, nom. illeg., incl. typo spec. Grimmia hartmanii subsp. euhartmanii
Giacom., Atti 1st. Bot. Lab. Crittog. Univ. Pavia, ser. 5, 4: 223. 1947, nom. inval. Grimmia
hartmanii var. euhartmanii Podp., Consp. Musc. Eur.: 289. 1954, nom. inval. PROTOLOGUE: ad
saxa quarzosa umbrosa per Vogesum [France] sylvaticum praeprimis superiorem, etiam in
Helvetia [Switzerland] prope Mornai ad latus meridionale m. Saleve, ubi in molibus graniticis
erraticis sub umbra Castaneae vescae copiose vidi; in Suecia [Sweden] copiosissime prope
Holmiam atque Upsaliam provenit. TYPE: Grimmia incurva Schw. Circa Holmiam vulgaris, in
Wermlandia raros. 1854, leg. s.o. Lindberg (lectotype, designated by Geissler & Maier (1995:
501), BM!).
Grimmia lubtomeda Rabenh., Bryoth. Eur. fas. 11, n° 510. 1862, nom. nud. Grimmia subtumida
Schimp. ex A. Jaeger, Ber. Tlitigk. St. Gallischen Naturwiss. Ges. 1872-73: 75 (Gen. Sp. Musc. 1:
WORLD SYNOPSIS OF GRIMMIA
35
357). 1874, nom. inval., orthogr. err. ORIGINAL SPECIMENS: [Germany] auf
Braunkohlensandstein in Siebengebirge bei Bonn. Januar 1861 leg. P. Dreesen (MO!, FH!, PC!).
Grimmia hartmanii f. propagulijera Milde, Bryol. Siles.: 150. 1869. Dryptodon hartmanii f.
propagulijer (Milde) Lirnpr., Laubm. Deutschl. 1: 790. 1889 ["propagulifera"]. PROTOLOGUE:
[Poland/Czech Republic border] Haufig irn Riesengebirge. TYPE: Riesengebirge. Leg. Dr. Milde
(lectotype, here designated, FH!; isolectotype, FH!).
Grimmia brotheri Lindb. ex Broth., Acta Soc. Sci. Fenn. 19(12): 89. 1892. Dryptodon brotheri
(Broth.) Warnst., Hedwigia 53: 305. 1913. PROTOLOGUE: [Georgia] Cauc. major occ.: Radscha,
ad saxa silicea regionis silvaticae mediae pro Uzeri, cum fructibus deoperculatis, sat numerosis;
Svania, in silvis jugi montani inter Enasch (Latal) et Betscho 15-1,600 m., cum fructibus
deoperculatis (E.L. n. 192!). TYPE: Imeretia: ad saxa umbrosa pr. Utseri [Uzeri]. 7-VII-1877. V.F.
Brotherus 301 (lectotype, here designated, H-SOL!).
Raeomitrium patens var. braehydietyon Cardot, Bull. Herb. Boissier, ser. 2, 8: 333. 1908. Grimmia
patens var. braehydietyon (Cardot) Iishiba, Nipponsan Senrui Sosetsu: 77. 1929. Grimmia
braehydietyon (Cardot) Deguchi, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 16: 173. 1979
["1978"]. PROTOLOGUE: Japon: Namikawa, rochers, a 800 m (n. 2807). TYPE: Japon:
Narnikawa, rochers, a 800 m. Leg. Faurie, 1904. n° 2807 (holotype, PC!; isotypes, H-BR!, NY!).
Grimmia borneensis Dixon & Herzog, Hedwigia 66: 344. 1926. PROTOLOGUE: Borneo: Ohne
nahere Fundortsangabe leg. Missionsinspektor R. Wegner, 1913. TYPE: F1. V. Borneo. Leg P.
Wegner 1913. Com. Kostrik (holotype, JE!).
Synonymized by Deguchi (1986: 328).
Grimmia hartmanii f. atra H. Moller, Ark. Bot. 26A(2): 91. 1934. PROTOLOGUE: [Sweden]
Gastrikland. Torsaker, Korsan 1911 M[ihi: H. Moller]. TYPE: Gastrickland. Torsaker, Korsan.
15.8.1911 Leg. Hj. Moller (lectotype, here designated, S!).
DISTRIBUTION: ARM (V Manakyan I802a, ERE); AUT (H. Winter s.n., JE); BEL (F. Gravet
S.n., PC); BGR (R. Dull Ie, JE); BIH (J. Gbwaeki s.n., JE); CHE (P. Culmann s.n., MA); CZE (A.
Geheeb s.n., FH); DEU (L. Meinunger 1118, JE); DNK (c. Jensen s.n., FH); ESP (P. Sera B-357,
BCB); FIN (Brotherus s.n., JE); FRA (J.-P. Frahm 935073, MA); GBR (D.G. Long 13469, MA); GEO
(A.L. Abramova s.n., LE); HUN (L. Vajda s.n., MA); ITA (J.L. De Sloover 23980, MA); JPN (E.
lishiba 7740, MO); NOR (Janzen s.n., JE); POL (J. Milde s.n., FH); ROM (T. Herzog s.n., JE); SFE
(V Ardeeva s.n., LE); SUN (A.L. Abramova & l.I. Abramov s.n., LE); SVK (H. Sedlmayr s.n., MO);
SWE (J.E. Zetterstedt s.n., H); TUR (Davis & Hedge D.30403a, MA); USA (T.P. James s.n., MO);
YUG (D. Katie s.n., JE).
24. Grimmia humilis Mitt., J. Linn. Soc., Bot. 12: 100. 1869. PROTOLOGUE: [Chile] Magellan, ad
Sandy Point, Lechler. TYPE: Sandy Point in pret. magell. Lechler (lectotype, designated by
Munoz (1999a: 140), NY; isolectotypes, BM!, PC!).
DISTRIBUTION: ARG (A. Donat 110, JE); CHL (H. Deguehi 26027, HIRO).
36
MuNoz AND PANDO
25. Grimmia incrassicapsulis B.G. Bell, Brit. Antarct. Surv. Bull. 63: 80. 1984. Grimmia inaequalis
Dixon & Sainsbury, Trans. & Proc. Roy. Soc. New Zealand 75(2): 174. 1945, nom. illeg., non
Stirt. 1908. PROTOLOGUE: [New Zealand] On rock, Mount Watkin, near Waikouaiti, Otago, ca.
1500 feet; colI. Simpson and Thomson, 30/1111935, No. 823, type. Same locality and collectors,
131711935, No. 811. TYPE: Mt. Watkin, Otago, N.Z. Simpson & Thomson nO 823. type (holotype,
BM!; isotype, NY!).
DISTRIBUTION: NZL (G. Simpson & Thomson 811, BM); SGS (B.G. Bell 3312, BM).
26. Grimmia incurva Schwagr., Sp. Musc. Frond. Suppl. 1(1): 90. 1811. Grimmia contorta f. incurva
(Schwagr.) Molendo, Ber. Naturhist. Vereins Passau 10: 119. 1875. Grimmia incurvata Schwagr.
ex Grout, Bryologist 14: 75. 1911, nom. inval., orthogr. err. PROTOLOGUE: [Austria] Legi in
summis alpibus Carinthiae, Glockner aliisque similibus primaevis, in Sudetum fossis nivalibus
sterilem observavit C. Ludwig. TYPE: Glockner (lectotype, designated by Deguchi (1978: 190),
G!; syntype ["In fossis nivalibus Sudetum"], B!).
Dicranum contortum Wahlenb., Fl. Carpat. Princ.: 346, tab. 4. 1814. Campylopus contortus
(Wahlenb.) Brid., Muscol. Recent. Suppl. 4: 74. 1818 ["1819"], Dryptodon contortus (Wahlenb.)
Brid., Bryol. Univ. 1(1): 199. 1826. Grimmia contorta (Wahlenb.) Am., Mem. Soc. Linn. Paris 5:
243. 1827. Grimmia contorta f. typica Chal., Pamietn. Fizyogr. 2: 40. 1882, nom. alt., nom. inval.
Grimmia contorta f. longifolia Chal., Pamietn. Fizyogr. 2: 40. 1882, nom. alt., nom. illeg.
Grimmia incurva f. longifolia (Chal.) Villi., Vestn. Knil. Ceske Spolecn. Nauk, n. Mat.-Pili.
1924: 31. 1924, nom. inval. Grimmia incurva f. typica (Chal.) Villi., Vestn. Knil. Ceske Spolecn.
Nauk, n. Mat.-Prir. 1924: 31. 1924, nom. inval. PROTOLOGUE: [Sweden] in cryptis saxorum ad
lacum alpinum Langesee im Volksgrund longe supra terrninum Mughi copiose. TYPE: Ad
Langensee vallis Volkensis. 10110/1813 leg. G. Wahlenberg (isotype, H-SOLl).
Synonymized by Deguchi (1978: 190).
Grimmia contorta f. tatrensis Chal., Pamietn. Fizyogr. 2: 40. 1882, nom. alt. Grimmia contorta f.
brevifolia Chal., Pamietn. Fizyogr. 2: 40. 1882, nom. alt. Grimmia contorta var. tatrensis (Chal.)
Chal., Enum. Musc. Frond. Tatr.: 51. 1886. Grimmia incurva var. tatrensis (Chal.) Herzog, Bot.
Jahrb. 47: 479. 1912. Grimmia incurva f. brevifolia (Chal.) Vilh., Vestn. Kral. Ceske Spolecn.
Nauk, n. Mat.-Prir. 1924: 32. 1924. Grimmia incurva f. tatrensis (Chal.) Villi., Vestn. Kral.
Ceske Spolecn. Nauk, Tr. Mat.-Prir. 1924: 32. 1924. PROTOLOGUE: [Poland] Brzegi Morskiego
nad Zmarzlym
Oka. Pyszna Hala w gorze. Dolina Malego Kolbachu, wprost Lomnicy. Pod ｗ｡ｧｾＬ＠
stawem. KOIl.czysta szczyt i nad ｂ｡ｴｹｩ［ｯｷ･｣ｫｾ＠
､ｯｬｩｮｾＬ＠
tudziei; od strony Tupej. Tomanowa,
szczyt. Polski Grzebien od strony Felki i od Zmarzlego. Gran od ｍｩｾｧｵｳｺｯｷｫ･＠
szczytu do
Mnicha. Rysy szcsyt. Krywan, szcsyt i ponii;ej. Lodowy, szcsyt i ponii;ej od strony doliny
Jaworzynskiej. Gierlach, szcsyt, tudziei; ponii;ej, od strony Batyi;owieckiej doliny. TYPE: Nad
6-VIII-79 [T. Chalubmski] (lectotype, designated by Ochyra & Cislo (1999:
Zmarzlym nad ｗ｡ｧｾＮ＠
79), ZAMU!).
Grimmia hagenii Kaurin, Nyt Mag. Naturvidensk. 31: 217. 1888 ["Hageni"]. Grimmia incurva var.
hagenii (Kaurin) C.E.O. Jensen, Meddel. Gmnland 15: 426. 1898. Grimmia contorta var. hagenii
(Kaurin) Paris, Index Bryol. Suppl.: 172. 1900. PROTOLOGUE: [Norway] Planten er funden af
Doktor Hagen paa Galdhe den Iste August 1887 i omtrent 1800 Meters Heide o. H. paa Klipper
(Gabbro?) [The plant was found by Dr. Hagen at Galdhe in l-August-1887 at approximately 1800
WORLD SYNOPSIS OF GRIMMIA
37
m on cliffs (Gabbro?)]. TYPE: Lorn: Galdlw 1800 mtr. 11 Aug. 1887. Leg. 1. Hagen (lectotype,
here designated, S!; isolectotypes, S! 2 replicates).
Coscinodon japonicus Sakurai, J. Jap. Bot. 19: 407, fig. 9. 1943. PROTOLOGUE: [Japan] Honsyu:
Provo Sinano, Sugadaira, Nekodake am Gipfel [... ] (Leg. N. Takaki Typus in Herb. K. Sakurai Nr.
14505 Juli 1942; Nr. 877 von Takaki ist dasselbe). TYPE: (holotype, Deguchi (1978: 191), MAK
not seen).
Synonymized by Deguchi (1978: 191).
Grimmia torngakiana Brassard & Hedd., Mem. New York Bot. Gard. 45: 216.1987. PROTO LOGUE
and TYPE: Canada, Labrador; Nachvak Fiord, Minaret Plateau, 58°55'N, 64°00'W, 1700 m
elevation, turfs on mineral soil in seepage from late snowbeds, 22 Jul 1983, Hedderson 1893
(holotype, NFLD!).
DISTRIBUTION: AUT (HF.v. Handel-Mazzetti s.n., MA); CAN (T.A. Hedderson 1892, NFLD);
CHE (F. Winter s.n., MO); CZE (z. Pilous s.n., MO); DEU (L. Loeske s.n., MO); ESP (R.M Cros s.n.,
BCB); FIN (V.F. Brotherus s.n., JE); FRA (WP. Schimper s.n., H-SOL); GBR (JB. Duncan 5431,
MO); GEO (A.H. Brotherus & v.F. Brotherus 128, H-SOL); GRL (WS. Benninghoff & HC Robbins
7034, NY); HUN (A. Boros s.n., BP); ITA (A. Giacomini s.n., JE); JPN (z. Iwatsuki s.n., W); NOR
(N.C Kindberg s.n., S); POL (H Bednarek-Ochyra & R. Ochyra 220190, MO); PRT (G. da Cunha &
L. Sobrinho 13, LISU); ROM (A. Boros s.n., MA); SFE (C Wright 78, FH); SIB (E. Ignatova 6118,
MA); SUN (E.A. Ignatova 6117, MO); SVK (L. Vajda s.n., BP); SWE (G. Wahlenberg s.n., H-SOL);
TWN (Peng Ching-I 83-35A, MO); TZA (C Uhlig s.n., H-BR); UKR (A. Boros s.n., BP); USA (G.
Allen 1434, MO).
27. Grimmia indica (Dixon & P. de la Yarde) Goffmet & Greven, J. Bryoi. 22: 141. 2000.
Trigonodictyon indicum Dixon & P. de la Yarde, Ann. Cryptog. Exot. 1: 40-42, fig. 3. 1928.
PROTOLOGUE: [India] Pambar Torrent, Kodaikanal, Pulney Hills [sic]. fevr. 1927. Leg. R. P.
Foreau. TYPE: Pambar Torrent, Kodaikanal, Palni Hills, S. India. 8.2.1927. Collector: R. P.
Foreau. In Herb. H. N. Dixon Ref. No. 815 (lectotype, here designated, BM!; isolectotypes, FH!,
MO!,NY!).
Grimmia apophysata Gangulee, Nova Hedwigia 12(3/4): 428, pI. 87. 1967. Grimmia apophysata A.
Jaeger, Ber. Tlitigk. St. Gallischen Naturwiss. Ges. 1872-73: 66 (Gen. Sp. Musc. 1: 348). 1874,
nom. nud. PROTOLOGUE: [India] Kurz 2344 lectus ad Phalut prope Darjeeling ad alt. 3,500 m.
die 16.X.1868 et servatus in CAL. TYPE: 2344. Sikkim, Phallort 11,000'. S. Kurz (isotype, HBR!).
DISTRIBUTION: IND (C Srinivasan 122, FH).
28. Grimmia involucrata Cardot, Rev. Bryoi. 36: 105. 1909. PROTOLOGUE: [Mexico] Etat de
Hidalgo: Cuzamaloza [sic], rochers, en compagnie de G. ovata Web. et Mohr, 1908 (n. 10598 in
parte). TYPE: Pringle, Plantae mexicanae n° 10598 p.p. On rocks, Cuyamaloya, 8500 ft. 17, Sept.,
1908. Herb. Cardot (lectotype, designated by Munoz (1999a: 141), PC!; isolectotype, PC!).
Synonymized with G. affinis Homsch. by Crum (1994: 396).
38
MuNoz AND PANDO
NOTE: In the description of the species in Munoz (1999a: 141) the capsule shape is erroneously
reported as ventricose. It is symmetric, as it was correctly indicated in the key.
DISTRIBUTION: MEX (CG. Pringle 10599, TENN).
29. Grimmia kidderi James, Bull. Torrey Bot. Club 6: 54. 1875. Grimmia kilderi James ex Paris,
Index Bryol. ed. 2: 277. 1904, nom. inval., orthogr. err. PROTOLOGUE: Southern part of the
Kerguelen Island, Lat. 49°21 's, Long. 70 0 15'E. TYPE: Kerguelen Island. Leg. Dr. lH. Kidder
1874 (lectotype, designated by Munoz (1999a: 143), FH!; isolectotypes, FH! 2 replicates).
Grimmiafrondosa James, Bull. Torrey Bot. Club 6: 54. 1875. PROTOLOGUE: Southern part of the
Kerguelen Island, Lat. 49°21 's, Long. 70 0 15'E. TYPE: Kerguelen Island. Dr. J.H. Kidder
(lectotype, designated by Munoz (1999a: 143), FHl; isolectotypes, FHl 2 replicates, NYl, PCl).
Synonymized by Munoz (1999a: 143).
Grimmia minutifolia Mull. Hal., Bot. Jahrb. Syst. 5: 81. 1884. PROTOLOGUE: Kerguelen. TYPE:
Kerguelen. Foundery branch. Dr. F. Naumann Dcbr. 1874 (lectotype, designated by Ochyra &
Hertel (1990: 25), PC!; isolectotype, H-BR!).
Synonymized by Ochyra & Hertel (1990: 25).
Guembelia immersoleucophaea Mull. Hal., in Neumayer, Int. Polarforsch., Deut. Exped. 2: 317.1890.
Grimmia immersoleucophaea (Mull. Hal.) Kindb., Enum. Bryin. Exot., Suppl. 2: 107. 1893.
PROTOLOGUE: [South Georgia] Austro-Georgia, ad rupes im Brockenthale raro, 23. Januario
1883; ad rupes des Koppenberges, 19 Majo 1883 cum fructibus calyptratis. TYPE: Brockenthale.
Sud-Georgien 23/183 Will (lectotype, designated by Ochyra & Hertel (1990: 25), HBG not seen;
syntype ["Koppenberges, 19.V.1883, Wilf'], PC!).
Synonymized by Ochyra & Hertel (1990: 25).
Grimmia austrofunalis Broth., Deutsche SUdpolar-Expedition 1901-1903, 8: 87, tab. 7 fig. 4. 1906,
nom. illeg., non Miill. Hal. 1898. Grimmia kerguelensis Cardot, Bull. Mus. Hist. Nat. (Paris)
22(6): 338. 1916, nom. nov. PROTOLOGUE: Kerguelen: Felswand an der Sudmire (Werth).
TYPE: Kerguelen, 181211903. Leg. E. Werth (holotype, H-BR!; isotypes, JE!, NY!, PC!, S!).
Synonymized by Ochyra & Hertel (1990: 25).
Grimmia nordenskioldii Cardot, Bull. Herb. Boissier, ser. 2, 6: 7. 1906. PROTOLOGUE: [South
Georgia] Cumberland bay: Bore valley, mt. Duse. Royal bay: Moltke harbour. TYPE: Georgie du
Sud: Royal bay. Leg. Skottsberg, 1902. n° 303 (lectotype, designated by Ochyra & Hertel (1990:
25), S not seen; isolectotype, S not seen; syntypes ["Royal Bay, Moltke Harbour"], H-BR!, NY!).
Synonymized by Ochyra & Hertel (1990: 25).
Grimmia bossierei Cardot & Ther., Bull. Mus. Hist. Nat. (Paris) 22(6): 338. 1916. PROTOLOGUE:
[Kerguelen] (Bossiere). TYPE: Kerguelen. Leg. R. Bossiere, 1913 (holotype, PC!; isotypes, HBR!, NY!, PC!).
Synonymized by Ochyra & Hertel (1990: 26).
Grimmia stenobasis Dixon, in Christoph., Results Norw. Sci. Exped. Tristan da Cunha 1937-1938,48:
23. 1960. PROTOLOGUE: Tristan da Cunha: On loose cinders, slopes of Peak, above Settlement,
WORLD SYNOPSIS OF GRIMMIA
39
1600m., No. 1751 p.p. TYPE: Tristan da Cunha. Above Settlement, slopes of Peak, loose cinders,
1600m., 15 March 1938. leg. E. Christophersen No. 1751a. det. H.N. Dixon (lectotype, here
designated, FH!).
DISTRIBUTION: ARG (c. Skottsberg 371, H-BR); ATA (J. Healy 65, FH); ATF (J.P. Hebrard
s.n., PC); CHL (H.L. Gunckel 1768, PC); .SGS (B.G. Bell 1181, BM); SHN (E. Christofersen &
Mejland 1780, BM); ZAF (B.J. Huntley 184, BM).
30. Grimmia laevigata (Brid.) Brid., Bryol. Univ. 1(1): 183. 1826. "Bryum trichodes hirsutie
canescens, capitulis subrotundis erectis, in pediculis brevissimis" Dill., Cat. PI. Circa Gissam: 224.
1718, nom. inval. "Bryum orbiculare pulvinatum, hirsutie canescens, var. capsulis erectis
praedita" Dill., Hist. Musc.: 395, tab. 50 fig. 65 D, E. 1741, nom. inval. Campylopus laevigatus
Brid., Muscol. Recent. Suppl. 4: 76. 1818 ["1819"]. Grimmia levigata Brid. ex Loeske, Laubm.
Eur. Part I: 78. 1913, nom. inval., orthogr. err. PROTOLOGUE: In Tyrole Italico, item circa
Romam, Neapolin et in insula Ischia in rupibus apricis 1806 abunde legi. Rohling etiam e
Palatinatu [Germany], Schleicher ex Helvetiae [Switzerland] miserunt. TYPE: Italia. Rome.
Napoli. Ischia 1806 (lectotype, designated by Cao & Vitt (1986: 181), B!).
Grimmia campestris Burchell ex Hook., Musci Exot. 2: 129. 1819. Dryptodon campestris (Hook.)
Brid., Bryol. Univ. 1: 794. 1827. PROTOLOGUE: [South Africa] In territorio coloniali Capitis
Bonae Spei, in regione Roggeveld, inter Jakhals Fonteyn et Kuylenberg;- terrestris, in campis
aridis, parce. D. Burchell. TYPE: Ex herb. Hook. An original specimen of Grimmia campestris
Hook. Musc. Ex. t. 129. Burchells Cape Mosses (isotype, H-SOL!).
Grimmia leucophaea Grev., Mem. Wem. Nat. Hist. Soc. 4: 87, pI. 6. 1822. Dryptodon leucophaeus
(Grev.) Brid., Bryol. Univ. 1: 773. 1827. PROTOLOGUE: [United Kingom] In rupibus prope
Edinburgum ad locum King's Park dictum. TYPE: Rocks King's Park. Edinb. R.K. Greville
(lectotype, designated by Munoz (1999a: 144), BM!; isolectotypes, FH!, NY! 3 replicates, PC!).
Dryptodon brevifolius Brid., Bryol. Univ. 1: 775. 1827. PROTOLOGUE: [Switzerland] In Helvetia
habitat ubi Schleicher in rupibus detexit. TYPE: Dryptodon brevifolius N. Schleich. Herb. Cando
(lectotype, designated by Munoz (1999a: 144), B!).
Synonymized by Munoz (1999a: 144).
Grimmia leiocarpa Taylor, London 1. Bot. 5: 44. 1846. PROTOLOGUE: [Australia] Swan River, Mr.
James Drummond. TYPE: Swan River. NH [Nova Hollandia] James Drummond 1843 (lectotype,
designated by Munoz (1999a: 144), FH!; isolectotypes, H-SOL!, NY! 2 replicates).
Synonymized by Munoz (1999a: 144).
Grimmia subleucophaea Mull. Hal., Bot. Zeitung (Berlin) 16: 164. 1858. PROTOLOGUE: [Ethiopia]
Abyssinia, Semen, ubi vulgatissirnam legit Octobr. 1850 W. Schirnper. ColI. II. No. 41 [Gr.
leucophaea W.P. Schirnp. in Musc. Abyss. Schimp.]. TYPE: 41. Auf felsblocken von 9000 fur
14000'. [illegible] 2 nov 50 9200' [illegible] (lectotype, designated by Ochyra in sched., BM!).
Grimmia crassa Miill. Hal. ex A. Jaeger, Ber. Tatigk. St. Gallischen Naturwiss. Ges. 1872-73: 82
(Gen. Sp. Musc. 1: 364). 1874, nom. nud. ORIGINAL MATERIAL: [South Africa] Africa
austral., Tafelberg ad prom. bon. spei. ORIGINAL SPECIMEN: Tafelberg am Cap. (NY!).
40
MuNoz AND PANDO
Grimmia calotricha Mull. Hal., Linnaea 43: 454. 1882. PROTOLOGUE: Argentinia Cordobensis,
Sierra de Cordoba, Las Penas, Febr. 1871. [leg. P.G. Lorentz]. TYPE: Sierra de Cordoba, Las
Penas. 1871 (lectotype, designated by Munoz (1999a: 144), BM!; isolectotype, PC!).
Synonymized by Munoz (1999a: 144).
Grimmia leucophaea var. latifolia Limpr., Bryoth. Siles. n° 163. 1867, nom. nud. Grimmia leucophaea
var. lali/olia Limpr., Laubm. Deutschl. 1: 742. 1889. Grimmia campestris var. latifolia (Limpr.)
Paris, Index Bryol. ed. 2: 269. 1904. Grimmia campestris f. latifolia (Limpr.) Monk., Laubm. Eur.:
356. 1927. Grimmia laevigata f. latifolia (Limpr.) Podp., Consp. Musc. Eur.: 276. 1954.
PROTOLOGUE and TYPE: [Poland] Spitzberg bei Hirschberg. Gesamrne1t Ostern 1867
[Limpricht, Bryoth. sil. No. 163 (1867)] (lectotype, designated by Munoz (1999a: 144), FH!;
iso1ectotype, JE!).
Synonymized by Munoz (1999a: 144).
Grimmia sarcocalyx Kindb., in Macoun, Bull. Torrey Bot. Club 17: 271. 1890, nom. nud. Grimmia
sarcocalyx Kindb., in Macoun, Cat. Canad. Pl., Musci: 66. 1892. Grimmia campestris subsp.
sarcocalyx (Kindb.) Kindb., Eur. N. Amer. Bryin. 2: 219. 1898. PROTOLOGUE: [Canada] On
rocks on the summit of the mountain between the Nicola and Thompson rivers at Spence's Bridge,
B.C., alt. 3,500 feet, May 28th, 1889 (Macoun.). TYPE: N. Amer., Br. Columb., Spence's Bridge,
3000' 28/5 89. J. Macoun. C.M. [Carl Muller] vid. (lectotype, designated by Munoz (1999a: 145),
S!; iso1ectotype, CANM-198098!).
Synonymized by Munoz (1999a: 145).
Grimmia bellii R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 416, pl. 33 fig. 24. 1895 ["belli"].
PROTOLOGUE: [New Zealand] Dry rocks, Lyttelton hills, in several places; 1882. Bird1ing's
Flat, Forsyth; Mount Self, Gorge of the Hurunui; 1893. Palmer Pass, head of the River Conway;
February, 1894. Collected in these localities by R.B. TYPE: Lyttelton Hills Ch. Ch. Collected by
Robert Brown (lectotype, here designated, BM!).
Grimmia murina Mull. Hal., Hedwigia 36: 105. 1897. PROTOLOGUE: Argentinia Patagonica, ad
pedem Sierrae Ventanae in rupibus, cum Catenularia Ventanae associata: P.G. Lorentz legit 27.
Feb. 1881. TYPE: Patagonia, Sierra Ventana. Lorentz 1881 (lectotype, designated by Munoz
(1999a: 145), JE!; isolectotypes, JE!, PC!).
Synonymized by Deguchi (1984: 21).
Grimmia leucophaea var. longirostris G. Roth, Eur. Laubm. 1: 564, tab. 51 fig. 9. 1903. Grimmia
laevigata var. longirostris (G. Roth) Podp., Consp. Musc. Eur.: 276. 1954. PROTOLOGUE:
[Germany] An Basaltfe1sen auf dem Ramsberg bei Laubach am 2. April 1903 in einem reich
fruchtenden Rasen von mir entdeckt. TYPE: Ramsberg bei Laubach (Hessen). 2IIV11903. G. Roth
(lectotype, here designated, S!).
Grimmia serripila Dusen ex Paris, Index Bryol. ed. 2: 287. 1904, nom. nud. ORIGINAL MATERIAL:
Am. merid. Chile. Dusen M. Am. austr. n. 366. ORIGINAL SPECIMEN: Chile N° 366 P. Dusen
(JE! [mixed with G. trichophylla], PC!).
Grimmia glauca Cardot, Rev. Bryol. 32: 17. 1905. PROTOLOGUE: [France] [... ] 24 fevrier 1903,
dans les environs de Charleville. TYPE: (a specimen origin.) Ardennes: Charleville: rocher
schistoux sec aBelair. Leg. J. Cardot, 1903 (lectotype, here designated, S!; isolectotype, BM!).
WORLD SYNOPSIS OF GRiMMIA
41
Grimmia senilis Shaw, Cape Monthly Mag. 17: 380. 1878. PROTOLOGUE: [South Africa] Collected
by McLea on rocks, Graaff-Reinet, and by Shaw at Colesberg; by Bolus fIrst gathered at GraaffReinet. TYPE: (PRE? not seen).
Synonymized by Magill (1981: 277).
DISTRIBUTION: AND (C Sergio s.n., LISU); ARG (CC Hosseus 1856, FH); ARM (v.
Manakyan 4873a, ERE); AUS (A.C Beauglehole 73091, MEL); AUT (H.C Funck s.n., H); BEL (C
Delogne s.n., FH); BRA (D.M. Vital 9196, FLAS); CAN (J. Macoun s.n., CANM); CHE (SE. Bridel
s.n., B); CHL (E. Barros 6877, FH); CRN (D.-K. Li 16137, ALTA); CPV (K. Bystrom 180, PC); CZE
(J. Podp&a s.n., H); DEU (A.CH. Braun s.n., H-SOL); DNK (A. Hesselbo s.n., FH); DZA (L. Trabut
2121, PC); ESP (J.L. Rupidera 45, SALA); ETH (E.F Gilbert 279A, NY); FRA (FG. Kneiff & CP.W
Maercker s.n., FH); GBR (R.K. Greville s.n., BM); GEO (G. Woronow 219, PC); GRC (H.K.
Haussknecht s.n., JE); HUN (L. Vajda s.n., MA); IND (R.R. Stewart 13725, FH); IRN (P. Goldblatt
822a, MO); ISR (M. Bizot 10040, PC); ITA (CH. Binstead 174, NY); KAZ (SG. Nesterova 521,
MO); KEN (T. Pocs et al. 9213IE, FH); KGZ (R. Kamelin s.n., LE); LSO (J. van Rooy 3287, NY);
MAR (M.R. Crosby 4989, NY); MEX (A.J. Sharp et al. 6086, TENN); MNG (W Schubert M273,
MO); NZL (D.H. Vitt 8532, MO); PAK (BL Burtt B726, MA); POL (J. Milde s.n., FH); PRT (C
Sergio et al. 30.1 c, MA); REU (M.R. Crosby 8403, MA); SIB (E. Ignatova 13/24, MA); SUN (AL
Abramova & /./. Abramov s.n., NY); SVK (z. Pilous s.n., FH); SWE (SO. Lindberg s.n., H); SYR
(H.Fv. Handel-Mazzetti 1658, FH); TUN (J.-P' Frahm 813442, MA); TUR (E. Nyholm & A.C
Crundwell 1060, F); TZA (T. Pocs et al. 881241A, FH); TZK (P. Polyakov s.n., LE); UKR (E.l
Vysotskaya s.n., H); DRY (E.J. Gibert 1243, PC); USA (A. Brant 3598, MO); UZB (Hudaikulov s.n.,
LE); ZAF (D. Hardy 4218, NY).
31. Grimmia leibergii Paris, Index Bryol. 2: 528. 1895, nom. nov. Grimmia pachyphylla Leiberg,
Bull. Torrey Bot. Club 20: 113, fIg. 144. 1893, nom. illeg., non Miill. Hal. 1885. PROTOLOGUE:
[U.S.A.] Habitat granite, gneissoid and slate rocks throughout Kootenai Co., Idaho. J.B. Leiberg,
No. 250, 1890-92. TYPE: Post Falls. Kootenai Co., Idaho. ColI. J.B. Leiberg 250. April 14th,
1890 (holotype, US not seen; isotypes, NY! 5 replicates, PC!).
DISTRIBUTION: USA (A.T. Whittemore 6584, MA).
32. Grimmia lisae De Not., Muscol. Ital. Spic.: 15. 1837. Grimmia muehlenbeckii var. lisae (De Not.)
Bott., in Venturi & Bott., Atti Soc. Crittog. Ital. 3: 217. 1884. Grimmia trichophylla subsp. lisae
(De Not.) Boulay, Musc. France 1: 378. 1884. Dryptodon lisae (De Not.) Loeske, Stud. Morph.
Syst. Laubm.: 111. 1910. Grimmia trichophylla var. lisae (De Not.) Bott., in Luisier, Broteria, Ser.
Bot. 14: 32. 1916. PROTOLOGUE: [Italy] Ad saxa in collibus Taurinensibus vulgatissimam
detexit D.us Lisa. TYPE: In collibus taurinensibus. Leg. Lisa 1836 (lectotype, designated by
Munoz (1999a: 145), RO!; isolectotypes, H-SOL!, NY!, RO!).
Grimmia trichophylla var. brachycarpa De Not., Syllab. Musc.: 250. 1838. PROTOLOGUE: [Italy.
Sardinia]. TYPE: [Italy. Sardinia] Sta. Barbara pro Cagliari. 3/3/835 (lectotype, here designated,
RO!).
42
MuNoz AND PANDO
Grimmia ancistrodes Durieu & Mont., Ann. Sci. Nat. Bot., ser. 3, 12: 318. 1849. PROTOLOGUE:
[Algeria] circa lacum El Hout detexit cl. Durieu. TYPE: Algerie. Durieu (lectotype, here
designated, PC!; isolectotypes, PC!, PC-Besch!).
Grimmia sardoa Mull. Hal., Syn. Musc. Frond. 1: 786. 1849. Grimmia trichophylla var. sardoa (Mull.
Hal.) Boulay, Musc. France 1: 379. 1884. Grimmia muehlenbeckii var. sardoa (Mull. Hal.) M.
Fleisch., Malpighia 7: 329. 1893, nom. inval., pro synon. Grimmia lisae subsp. sardoa (Mull.
Hal.) Kindb., Eur. N. Amer. Bryin. 2: 223. 1898. Dryptodon sardous (Mull. Hal.) Loeske, Stud.
Morph. Syst. Laubm.: 111. 1910 ["sardoa"]. Grimmia trichophylla subsp. sardoa (Mull. Hal.)
Bott., in Loeske, Laubm. Eur. Part I: 133. 1913. PROTOLOGUE: [Italy] Sardinia, ad rupes
montium: De Notaris Martio 1835 legit. TYPE: ad rupes in montius Sardinia australis. a D.
DeNotaris (lectotype, designated by Munoz (1999a: 145), RO!).
Synonymized by Munoz (1999a: 146).
Grimmia californica Sull., Expl. Railroad Mississippi Pacific, Descr. Moss. Liverw.: 187, pI. 4. 1857.
PROTOLOGUE: [U.S.A.] Oakland, opposite San Francisco; also near Sonora. TYPE: Trees and
dry places, mountains near Oakland, California. J.M. Bigelow, April 5, 1854. No. 2.b (lectotype,
designated by Munoz (1999a: 146), FH!).
Synonymized by Munoz (1999a: 146).
Grimmiaflexiseta Schimp. ex Jard., Bull. Soc. Linn. Normandie, ser. 2, 9: 263,264. 1875, nom. nud.
ORIGINAL MATERIAL: [U.S.A.] sur les rochers a Saucelito, dans la baie de San Francisco [... ]
ORIGINAL SPECIMEN: California. San Francisco (NY!, PC!).
Grimmia canariensis Schimp. ex Paris, Index Bryol. 2: 520. 1895, nom. nud. ORIGINAL
MATERIAL: [Spain] Husn. M. Can. n° 181a. - Afr.: Ins. Canar. ORIGINAL SPECIMEN: T.
Husnot, PI. Canarienses.- 1866. N° 181 [a] - Tenerife. Los Organos. Juin (PC!).
Grimmia canadensis Kindb., Eur. N. Amer. Bryin. 2: 226. 1898. PROTOLOGUE: [Canada] Amer. R.
Can. Vanc. isl.: Macoun. TYPE: Canada, Vancouver isl. 26/5 1893 J. Macoun 1894 (lectotype,
designated by Munoz (1999a: 147), S!).
Synonymized by Munoz (1999a: 146).
Grimmia azorica Renauld & Cardot, Bull. Soc. Roy. Bot. Belgique 41(1): 58. 1905. Grimmia
trichophylla subsp. azorica (Renauld & Cardot) Luisier, Broteria, ser. Bot. 17: 42. 1919.
PROTOLOGUE: [Portugal] Ayores: Graciosa (Brown, Blanchy); Santa-Maria (Trelease). TYPE:
n° 1393c. Sta. Maria. Collected by Wm. Trelease 6/28, 1896 (lectotype, designated by Munoz
(1999a: 147), PC!; isolectotype, MO!; syntype ["Brown s.n., Graciosa"], PC!).
Synonymized by Munoz (1999a: 146).
Grimmia sardoa var. robusta M. Fleisch. & Warnst., Bryotheca meridionalis n° 342. 1910, nom. nud.
Grimmia trichophylla f. robusta (M. Fleisch. & Warnst.) Podp., Consp. Musc. Eur.: 288. 1954,
nom nud. ORIGINAL SPECIMEN: Spanien: Sierra de Cordoba, bei Cordoba 400-600 m an
Felsen. April 1908. M. Fleischer. [Fleischer & Warnstorf, Bryotheca europaea meridionalis, n°
342 (1910)] (FH!).
Grimmia trichophylla f. propagulifera H. Winter, Hedwigia 55: 101. 1914, nom. illeg., non Limpr.
1889. PROTOLOGUE: [Spain] Guimar (Teneriffa), auf Felsen am deutschen Sanatorium, [... ]
WORLD SYNOPSIS OF GRiMMIA
43
[Miirz-Mai 1912]. TYPE: Grimmia trichophylla fonna depauperata. Guimar, auf felsen dem
Sanatorium. Marz 1912 leg. Dr. Winter (lectotype, designated by Munoz (1999a: 147), JE!).
Synonymized by Munoz (1999a: 146).
DISTRIBUTION: BEL (J. Cardot s.n., PC); CAN (J. Macoun s.n., CANM); CPV (A. Chevalier
s.n., PC); CYP (Unger s.n., JE); DZA (J. Feldmann 2222, MA); ESP (c. Sergio et al. 20.8, BCB);
FRA (F. Renauld s.n., MO); GBR (D.A. Jones 1394, FH); GRC (P. Haritonidou s.n., MO); ISR (T.
Kushnir 675, PC); ITA (A. Bottini s.n., FH); MEX (E. Palmer 114a, MO); PRT (F. Carreiro 1017,
PC); SAU (W Frey & H. Kurschner 1538, MA); TUN (C.J. Pitard s.n., PC); TUR (H. Kurschner & G.
Parolly 6967, MA); USA (A. T. Whittemore 3530, MO); YUG (J. Baumgartner s.n., FH).
33. Grimmia longirostris Hook., Musci Exot. 1: 62. 1818. PROTOLOGUE: [Ecuador] Locis
scopulosis, apricis, in declivitate montis Chimborazo, altit. 2380 hexapod. super porphyrite.
(Regnum Quitense) Humboldt et Bonpland. TYPE: H. 2670. N. 76. M. de Humboldt (lectotype,
designated by Deguchi (1984: 21), BM!; isolectotype, BM!).
Grimmia affinis Homsch., Flora 2(2): 443. 1819, nom. illeg., non Homsch. 1819. Dryptodon ovatus
var. affine (Homsch.) Hartm., Handb. Skand. Fl. ed. 3: 271. 1838. Grimmia ovata var. affinis
(Homsch.) Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 123, tab. 255B. 1845, nom. illeg.,
incl. typo var. prior. Grimmia ovalis var. affinis (Homsch.) Broth., Acta Soc. Sci. Fenn. 19(12): 86.
1892. Grimmia ovalis f. affinis (Homsch.) Monk., Laubm. Eur.: 360. 1927. PROTOLOGUE:
[Austria] Auf Felsenblocken der Tauernkette und an diese angranzender Gebirge in Gesellschaft
mit Gr. incurva Schwaegr., Gr. obliqua mihi und Gr. ovata und reift ihre Friichte im Aug. und
Sept. TYPE: Windissmattreyer Tauem (lectotype, designated by Deguchi (1978: 158), B!).
Synonymized by Munoz (1998a: 353).
Grimmia obliquata Host, Syn. PI.: 567. 1797, nom. inval. Grimmia obliqua Homsch., Flora 2(1): 84.
1819. Grimmia obliquata Host ex Brid., Bryol. Univ. 1(1): 180. 1826, nom. inval., pro synon.
Grimmia ovata var. obliqua (Homsch.) Huebener, Muscol. Genn.: 183. 1833. Dryptodon ovatus
var. obliquus (Homsch.) Hartm., Handb. Skand. Fl. ed. 3: 271. 1838. Grimmia ovalis var. obliqua
(Homsch.) I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1909(5): 26. 1909.
Grimmia ovalis f. obliqua (Homsch.) Monk., Laubm. Eur.: 360. 1927. Grimmia ovata f. obliqua
(Homsch.) Loeske, Biblioth. Bot. 101: 133. 1930. PROTOLOGUE: [Austria] Crypt. Cent. II. Dec.
V. Auf Felsenblocken in den Kamthner u. Salzburger Alpen und auf dem Heiligenbl. Tauem.
TYPE: In alpibus Salisburgensibus. Dr. Homschuch (lectotype, designated by Munoz (1998a:
353), BM!; syntypes ["ex alp. Carinth."], BM! 3 replicates; ["Heiligenbluter-Tauem"] JE!).
Synonymized by Munoz (1998a: 353).
Grimmia patens Homsch., Flora 2: 84. 1819. Grimmia ovata var. patens (Homsch.) Huebener,
Muscol. Genn.: 182. 1833. Dryptodon ovatus var. patens (Homsch.) Hartrn., Handb. Skand. Fl.
ed. 3: 271. 1838. PROTOLOGUE: [Gennany] 1st von unsenn Freunde, Herr Apotheker Funck,
auf dem Fichtelberge gefunden und in vie len seiner Hefte als G. ovata ausgegeben worde .. ["bey
Bemeck" Funck, Cryptogarnische Gewlische des Fichtelgebirg's, fasc. 4, nO 86 (1805 "1804")].
TYPE: An Felsen bey Bemeck. [Funck, Cryptogamische Gewlische des Fichtelgebirg's, fasc. 4, n°
86 (1805 "1804")] (lectotype, here designated, GOET!; isolectotype, JE!).
44
MuNoz AND PANDO
Grimmia ajjinis var. macrocarpa Nees & Homsch., Bryol. Germ. 2(1): 145, tab. XXI, 13c. 1827.
Campylopus ovalis var. hyrsutus Myrin, Kongl. Vetensk. Acad. Handl. 1831: 258. 1832, nom.
illeg., incl. typo var. prior. Grimmia ovata var. hirta Myrin, Arsberatt. Bot. Arbeten Upptackter
1831: 333. 1832, nom. inval., orthogr. err. PROTOLOGUE: [Austria] wachst auf verwitterten
Urgebirgsfelsen in der Gastein (Funck!). TYPE: (lectotype, here designated, fig. 13c in tab. XXI,
Nees, C.G., Homschuch, F. & Sturm, 1., Bryol. Germ.). After an unsuccessful search for
specimens referable to this taxon, the illustration in the original publication is proposed as
lectotype of the name. It unambiguously represents Grimmia longirostris Hook.
Grimmia ajjinis var. ramosissima Nees & Homsch., Bryol. Germ. 2(1): 144, tab. XXI fig. 13b. 1827.
Grimmia ajjinis var. elongata Nees & Homsch. ex Huebener, Muscol. Germ.: 709. 1833, nom.
illeg., incl. typo var. prior. PROTOLOGUE: [Austria] wachst auf den genannten hOchsten Alpen
urn Heiligenblut, aber immer hoher als 6, z. B. am Fufse der Leiterkopfe auf der Pasterze, bei dem
sogenannten Schaflerloch auf der Leiteralpe, dem Malnitzer-Tauem u.s.w. (Hsch.). TYPE:
Heiligenblut b. Dartl. (lectotype, designated by Munoz (1998a: 353), BM!).
Synonymized by Munoz (1998a: 353).
Grimmia cylindrica Nees & Homsch., Bryol. Germ. 2(1): 161, tab. XXIII fig. 18. 1827. Grimmia
ovata var. cylindrica (Nees & Homsch.) Huebener, Muscol. Germ.: 183. 1833. Grimmia ovata
var. submutica De Not., Syllab. Musc.: 246. 1838, nom. illeg., incl. typo var. prior. Grimmia ovalis
var. cylindrica (Nees & Homsch.) I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim)
1909(5): 26. 1909. Grimmia ovalis f. cylindrica (Nees & Homsch.) Monk., Laubm. Eur.: 360.
1927. PROTOLOGUE: [Austria] Wachst auf kiesigen und sandigen Stellen im Anlaufthal bei
Bockstein in der Gastein (Funck!). TYPE: (lectotype, here designated, fig. 18 in tab. XXIII, Nees,
C.G., Homschuch, F. & Sturm, 1., Bryol. Germ.). After an unsuccessful search for specimens
referable to this species, the illustration in the original publication is proposed as lectotype of the
name. It unambiguously represents Grimmia longirostris Hook.
Grimmia sciuroides Nees & Homsch., Bryol. Germ. 2(1): 155, tab. XXII fig. 17. 1827. Grimmia ovata
var. sciuroides (Nees & Homsch.) Huebener, Muscol. Germ.: 183. 1833. Dryptodon sciuroides
(Nees & Homsch.) Hartm., Handb. Skand. Fl. ed. 3: 271. 1838. PROTOLOGUE: [Austria] in der
Gegend von Heiligenblut (Hsch. 1816), auf der Zirbnitzalpe im Mollthal (Laurer!), im Anlaufthal
in der Gastein (Funck! Laurer!). TYPE: [Austria] Am Heiligenblut Baum (lectotype, here
designated, GZU!).
Grimmia sciuroides f. elongata Nees & Homsch., Bryol. Germ. 2(1): 159, tab. XXII fig. 17c.a. 1827.
PROTOLOGUE: [Austria] [... ] auf den Kamthner-Alpen, ohne nahere Bezeichnung des
Standortes (Laurer! Hoppe!). TYPE: Herb. Nees v Esenb. (lectotype, here designated, G!).
Grimmia sciuroides var. foliosa Nees & Homsch., Bryol. Germ. 2(1): 158, tab. XXII fig. 17a. 1827.
PROTOLOGUE: [Austria] in der Gastein (Hsch. 1816), bei Dollach (Funck! Laurer!), auf der
Pasterze (Laurer! Hoppe!). TYPE: Herb. Nees v Esenb. (lectotype, here designated, G!).
Grimmia sciuroides var. laxa Nees & Homsch., Bryol. Germ. 2(1): 159, tab. XXII fig. 17c.b, d. 1827.
PROTOLOGUE: [Austria] im Anlauf- und Rotschachthal in der Gastein (Funk! Laurer!), bei dem
Tauernhaus und am GeschlOfs in der Windischmattrey (Laurer! Funck!), auf dem MalnitzerTauem (Mielichhofer! Laurer!). TYPE: Herb. Nees v Esenb. (lectotype, here designated, G!).
WORLD SYNOPSIS OF GRiMMIA
45
Grimmia sciuroides var. rigida Nees & Homsch., Bryol. Germ. 2(1): 158, tab. XXII fig. 17b. 1827.
PROTOLOGUE: [Austria] auf den Alpen um Heiligenblut (Hoppe! Funck! Laurer! Hsch.). TYPE:
In alpibus pro Heiligenblut (lectotype, here designated, GZU!; isolectotype, G!).
Grimmia schimperi Bruch & Schimp. ex Mull. Hal., Syn. Musc. Frond. 1: 792. 1849. PROTOLOGUE:
[Ethiopia] (in Musc. Abyss. No. 584 [err. pro 484]) [... ] Abyssinia: W. Schimper. TYPE:
Abyssinia [Schimper, W. 1842, Schimperi iter abyssinicum, Sectio II, n° 484] (lectotype,
designated by Munoz (1998a: 353), NY!; isolectotype, PC!).
Synonymized by Munoz (1998a: 353).
Grimmia neilgherriensis MUll. Hal., Bot. Zeitung (Berlin) 11: 62. 1853. Grimmia nilghiriensis MUll.
Hal. ex Mitt., 1. Linn. Soc., Bot. Suppl. 1: 44. 1859, nom. inval., orthogr. err. Grimmia
neelgheeriensis MUll. Hal. ex Kindb., Enum. Bryin. Exot.: 61. 1888, n9m. inval., orthogr. err.
PROTOLOGUE: [India] Gr. ovata Mont. Musc. Neilgher. No. 54? [in truncis arborum et rupibus
circa Dodabett lectus (Montagne, C. 1842. Cryptogarnae Nilgherienses seu Plantarum cellularium
in montibus peninsulae indicae Neel-Gherries dictis a Cl. Perrottet collectarum Enumeratio. Ann.
Sci. Nat. Bot. Ser. 2, 17: 243-256]. TYPE: in montibus Neilgheriensibus Indiae leg. Cl. Perrottet
sub n° 1583 (lectotype, designated by Muiioz (1998a: 353), PC!; isolectotype, PC!).
Synonymized by Munoz (1998a: 353).
Grimmia columbica De Not., Mem. Reale Accad. Sci. Torino, ser. 2, 18: 447, fig. 7. 1859.
PROTOLOGUE: [Ecuador] a clarissimo Osculati ad flumen Napo in Columbia meridionalis.
TYPE: [Ecuador] ad river Napo Columbia. Leg. Osculati. Herb. De Not. (isotype, H-SOLl).
Synonymized by Munoz (1998a: 353).
Grimmia khasiana Mitt., J. Linn. Soc., Bot. Suppl. 1: 45. 1859. PROTOLOGUE: [India] Hab. In mont.
Khasian. reg. temp., ad rupem Kollong, inter Cladonias, 1.D: Hooker et T. Thompson! (No. 279).
TYPE: Kullong rocks. n° 279 (holotype, NY!).
Grimmia peruviana Sull., U. S. Expl. Exped., Musc.: 8, tab. 5A. 1860. PROTOLOGUE: Hab. Andes
of Peru, at an elevation of 15.000 feet. TYPE: U.S. Ex. Ex. Wilkes 1838-1842. Andes of Peru. at
an elevation of 15.000 feet (lectotype, designated by Muiioz (1998a: 353), BM!; isolectotypes,
BM!, FH!, NY!, PC!).
Synonymized by Munoz (1998a: 353).
Guembelia bogotensis Hampe, Linnaea 32: 142. 1863. Grimmia bogotensis
Tiitigk. St. Gallischen Naturwiss. Ges. 1872-73: 70 (Gen. Sp.
PROTOLOGUE: [Colombia] Bojota Los Laches ad rupes, 2800 metr.,
2011. Nova Granata. Bogota. Los Laches. alto 2800 metro 1859 Jul.
designated by Munoz (1998a: 353), PC!; isolectotype, NY!, PC!).
Synonymized by Churchill (1989: 109).
(Hampe) A. Jaeger, Ber.
Musc. 1: 352). 1874.
Jul. N° 2011. TYPE: N°
Lindig 2011 (lectotype,
Grimmia hausmanniana De Not., Comment. Soc. Crittog. Ital. 2: 102. 1866. Racomitrium
hausmannianum (De Not.) Molendo, Ber. Naturhist. Vereins Passau 10: 127. 1875. Grimmia
ovata subsp. hausmannii Kindb., Eur. N. Amer. Bryin. 2: 227. 1898. Grimmia ovalis f. epilosa
46
MuNoz AND PANDO
Podp., Consp. Musc. Eur.: 278. 1954, nom. illeg., non Warnst. 1904. PROTOLOGUE: [Austria]
Monte Rittnerhorn nel Tirolo meridionale, Hausmann. TYPE: Rittner Hom. Tyrol austr. Leg.
Hausmann (isotype, BM!).
Synonymized by Munoz (1998a: 353).
Grimmia tenuicaulis Hampe ex A. Jaeger, Ber. Tlitigk. St. Gallischen Naturwiss. Ges. 1872-73: 72
(Gen. Sp. Musc. 1: 354). 1874, nom. nud. ORIGINAL MATERIAL: America austral., Chile.
ORIGINAL SPECIMEN: Chile Andes. Herb. Hampe (FH!, NY!).
Grimmia ovata var. praecox A. Kern., Flora exsiccata austro-hungarica n° 316. 1881. PROTOLOGUE
and TYPE: [Austria] Tirolia centralis. In saxis schist. ad Trins in valle Gschnitz; 1200 mt. s.m.
(lectotype, designated by Munoz (1998a: 353), BM!; isolectotypes, FH! 2 replicates, GOET!).
Synonymized by Munoz (1998a: 353).
Grimmia integridens Mull. Hal., Linnaea 43: 460. 1882. PROTOLOGUE: Argentinia subtropica
Tucumanensis, in alpinis "der Cienega" cum Psilopilo gymnostomulo forsan in altitudine 11,000
ped., 1873 [leg. P.G. Lorentz]. TYPE: Argentinien: Tucuman, in der Cienaga, ca. 11000'. 1873
leg. P.G. Lorentz (lectotype, designated by Munoz (1998a: 353), PC!).
Synonymized by Munoz (1998a: 353).
Grimmia leucophaeola Mull. Hal., Linnaea 43: 456. 1882. PROTOLOGUE: Argentinia Tucumanensis,
in alpinis Sierrae de Aconquija prope Tafi, Aprili 1872. [leg. P.G. Lorentz]. TYPE: Argentina.
Tafi. 1872 (lectotype, designated by Munoz (1998a: 353), JE!; isolectotypes, BM!, JE!, NY!).
Synonymized by Munoz (1998a: 353).
Grimmia raphidostega MUll. Hal., Linnaea 43: 459. 1882. PROTOLOGUE: Argentinia subtropica
Saltensis, Nevado de Castillo, in alpinis cum Andreaeaceis, 1873. [leg. P.G. Lorentz]. TYPE:
Flora argentiniae subtropicae. Salta, Nevado de Castillo, in alpinis. 1873 leg. P.G. Lorentz
(lectotype, designated by Munoz (1998a: 353), JE!).
Synonymized by Munoz (1998a: 353).
Grimmia vernicosula MUll. Hal., Linnaea 43: 458. 1882. PROTOLOGUE: Argentinia subtropica, Rio
seco [sic] prope Oran, 1873. [leg. P.G. Lorentz]. TYPE: Argentina. Oran. Lorentz (lectotype,
designated by Munoz (1998a: 353), JE!; isolectotype, BM!).
Synonymized by Munoz (1998a: 353).
Grimmia rufa MUll. Hal., Bot. Jahrb. Syst. 5: 81. 1884. PROTOLOGUE: Kerguelen [Foundery branch,
Nov. 1874, Naumann]. TYPE: Kerguelen, XI 1874. Naumann (lectotype, designated by Munoz
(1999a: 149), PC!; isolectotype, H-BR!).
Synonymized by Munoz (1999a: 149).
Grimmia calyculata MUll. Hal., Flora 71: 414. 1888. PROTOLOGUE: [Tanzania] Monte
Kilirnandscharo, ad fmem sylvestrem superiorem inter 3000--4000 m. TYPE: Kilirnandscharo. Rh.
Muller (lectotype, here designated, PC-Besch!; isolectotype, BM!).
Grimmia leucophaeoides MUll. Hal. ex Kindb., Enum. Bryin. Exot., Suppl. 1: 91. 1889, nom. nud.
ORIGINAL SPECIMEN: Nevado de Castillo. Argentinia. Lorentz (JE!).
WORLD SYNOPSIS OF GRIMMIA
47
Grimmia cavifolia Lindb. & Arnell, Kongl. Svenska Vetenskapsakad. Handl. 23(10): 103. 1890.
Grimmia canifolia Lindb. & Arnell ex Broth., Nat. Pflanzenfam. 3(1): 452. 1902, nom. inval.,
orthogr. err. PROTOLOGUE: [Russia] Fl. Jen.[Flumen Jenisei], T. mont.: Takmak bei
Krasnojarsk. TYPE: Sibiria: Jenisei, Krasnojarsk, 56° nJ. pa urberg. 11/6/1876. H. Wilh. Arnell
(lectotype, designated by Munoz (1998a: 353), H-SOL!; isolectotypes, JEl, PC1, UPS1).
Synonymized with G. ovalis by Ignatov & Cao (1994: 89), but see Munoz (1998a: 353).
Grimmia immergens Mull. Hal., Flora 73: 483. 1890. PROTOLOGUE: [Tanzania] Africa or. trop.,
Kilirna-Ndscharo, 3000-4800 m: Dr. Hans Meyer 1889. TYPE: Insched.- Kilirnandscharo. hb.
Muller (lectotype, here designated, PC-Beschl; isolectotype, BM1).
Grimmia subovata var. laevipila Schirnp. ex Paris, Index Bryol. 2: 538. 1895, nom. nud. ORIGINAL
MATERIAL: [Bolivia] Am. merid.: ibid. [Mandon PI. Boliv.] n. 1635. ORIGINAL SPECIMEN:
Amasuyos viciniis Achacache, Putchani. Alt. Reg. alpina 3950 m. Jul. 1856. G. Mandon - Plantae
Andium Boliviensium nO 1635 (BMl 2 replicates, PC1).
Grimmia dimorphula Mull. Hal., Nuovo Giom. Bot. Ital., n.s. 3: 108. 1896. PROTOLOGUE: Prope
cacumen montis Iu-kia-po ad saxa, Jun. TYPE: China interior, provincia Schen-si sept. in saxis
cacuminis montis Iu-kia-po, ster. 27 Jun. 1894 legit Rev. Jos. Giraldi, detrm. C. Muller sub n° 877
(lectotype, designated by Cao & Vitt (1986: 170), H; isolectotypes, PCl 4 replicates).
Synonymized with G. ovalis by Cao & Vitt (1986: 170).
Grimmia nanoglobosa MUll. Hal. ex E. Britton, Bull. Torrey Bot. Club 23: 477. 1896.
PROTOLOGUE: [Bolivia] Mapiri, 5000 ft. (3195). TYPE: Mapiri, Bolivia, 5,000 ft., May, 1886.
Leg. H.H. Rusby, M:D. (holotype, NY1).
Synonymized by Deguchi (1987: 25).
Grimmia ortholoma Kindb., Rev. Bryol. 23: 19. 1896. PROTOLOGUE: Amer. Canada, Brit.
Columbia: Macoun. TYPE: N. Amer., B.c., Dear Park, rocks. 8/690. J. Macoun (holotype, Sl;
isotype, Sl).
Synonymized by Allen (1995: 161).
Grimmia breviexserta Mull. Hal., Bull. Herb. Boissier 5(3): 200. 1897. Grimmia brevipes MUll. Hal.
ex Paris, Index Bryol. Suppl.: 172. 1900, nom. inval. PROTOLOGUE: [Guatemala]
Quezaltenango, ColI. N° 115, cum Grimmia bernoullii associata [leg. Bernoulli & Cario]. TYPE:
Ex Museo Museo Berolinensi. Guatemala: Quezaltenango. Leg. Bernoulli et Cario (lectotype,
designated by Munoz (1998a: 353), PC1).
Synonymized by Munoz (1998a: 353).
Grimmia micromitria Schirnp. ex Mull. Hal., Nuovo Giorn. Bot. Ital., n.s. 4: 165. 1897, nom. nud.
ORIGINAL MATERIAL: [Bolivia, Mandon] n° 1632. ORIGINAL SPECIMEN: G. Mandon.
Plantae Andium Boliviensium n° 1632. Provo Larecaja. Viciniis Sorata, Lorna de Ullontiji, in
scopulosis. Reg. temp. 2800 m. Feb. 1868 (BM1).
Grimmia micro-ovata MUll. Hal., Nuovo Giorn. Bot. Ital., n.s. 4: 128. 1897. PROTOLOGUE:
[Bolivia] Germain [... ] 1889 prope Choquecamata provinciae Cochabamba Bolivianae altomontosae ex altitudine pedum 10-12,000, ut opinor, collectas. TYPE: Bolivia, provincia
48
MuNoz AND PANDO
Cochabamba prope Choquecamata. Jun. 1889 legit Germain determ. Dr. C. Muller sub n° 1142
(lectotype, designated by Munoz (l998a: 353-354), JE!; isolectotypes, NY!, PC!).
Synonymized by Deguchi (1987: 25).
Grimmia semipilosa Hampe, in Mull. HaL, Linnaea 43: 461. 1882, nom. inval., pro synon. Grimmia
subovata Schimp. ex Mull. HaL, Linnaea 43: 461. 1882, nom. inval., pro synon. Grimmia
subovata Schimp. ex Mull. HaL, Nuovo Giorn. Bot. Ital., n.s. 4(2): 128. 1897. PROTOLOGUE:
Bolivia, provo Larecaja, Sorata, Ancouma, supra muros de la Hacienda Penas, reg. alpina, 37004000 m. alta, Aprili 1860: G. Mandon. TYPE: Provo Larecaja, vicinitis Sorata; Ancourna, supra
muros de la hacienda Penas. Regio alpina y subalpina, 3700-4000 m. April 1860. G. Mandon Plantae Andiurn Boliviensium n° 1634 (lectotype, designated by Munoz (l998a: 354), BM!;
isolectotypes, BM! 2 replicates, FH!, G!, NY!, PC!).
Synonymized by Munoz (1998a: 354).
Grimmia arctophila subsp. labradorica Kindb., Eur. N. Amer. Bryin. 2: 221. 1898. PROTOLOGUE:
Amer. r. Can.: Northern Labrador: Macoun 1896. TYPE: Canada, N. Labrador 2117 1896 leg. J.
Macoun. Det. N.C. Kindberg (lectotype, designated by Munoz (1998a: 354), S!; isolectotype,
CANM-198076!).
Synonymized by Munoz (1998a: 354).
Grimmia itatiaiensis Broth. ex Mull. Hal., Bull. Herb. Boissier 6(2): 108. 1898. PROTOLOGUE:
Brasilia, Serra Itatiaia, ad ropes, 2100 m. alta. Martio 1894: E. Ule, ColI. N° 1913. TYPE: Serra do
ltatiaia, 2100 m., ad ropes, m. Martii 1894, leg. E. Ule. Ule & Brotheros, Bryotheca brasiliensis, n°
124 (lectotype, designated by Munoz (1998a: 354), H-BR!; isolectotypes, GOET!, JE!, NY!, PC!
3 replicates, UPS!).
Synonymized with G. affinis by Cao & Churchill (1995: 505). These authors reported page 51 for
the place of publication, but this is the page of the independently paginated reprint. The actual page of
the journal is 108.
Grimmia itatiaiae Mull. Hal., Bull. Herb. Boissier 6(2): 109. 1898. PROTOLOGUE: Brasilia, Serra
Itatiaia, 2300 m. alta, ad ropes der Agulhas Negras, Martio 1894, c. fro irnmaturis paucis: E. Ule,
CoIl. N° 1830. TYPE: Brasilia, Serra do Itatiaia, an Felsen der Agulhas-Negras. 2300 m, 3/1894.
Leg. E. Ule 1830 (lectotype, designated by Munoz (1998a: 354), H-BR!).
Synonymized by Munoz (1998a: 354).
Grimmia commutata f. brevipila Broth., Musci turkestanici, n° 49. 1899, nom. nud. in sched.
ORIGINAL SPECIMEN: [Kazakstan] Kungei Alatau: Kokoirok in valle fl. Koissu, ad saxa reg.
alp., Jul. 13, 1896. leg. V.F. Brotheros (JE!).
Grimmia preussii Broth. ex Paris, Index Bryol. Suppl.: 175. 1900, nom. nud. ORIGINAL SPECIMEN:
[Cameroon] Afr. occ., Kamerun-pii, auf Lava, 3500 m., 26 Febr. 1891. Leg. Preuss (H-BR!, PC!).
Grimmia elata Kindb., Rev. Bryol. 32: 33. 1905. PROTOLOGUE: 1. Found by J. Macoun in the
Canadian Yukon-district 1902. TYPE: Canada, Yukon distr., Hunker creek 25/7 1902. Macoun
(lectotype, designated by Munoz (1998a: 354), S!; isolectotypes, CANM-198084!, H-BR!).
Synonymized by Munoz (1998a: 354).
WORLD SYNOPSIS OF GRIMMJA
49
Grimmia ovataeformis Kindb., Rev. Bryol. 32: 35. 1905. PROTOLOGUE: 2. Found by J. Macoun
1904 in British Columbia, 5200-7500 feet above the sea. TYPE: Canada, B.C., lake Louise, rocks,
6000 f., 13/904. Macoun (holotype, S!; isotype, CANM-198092!).
Synonymized with G. ovalis by Allen (1995: 162), but see Munoz (1998a: 353).
Grimmia praetermissa Cardot, Rev. Bryol. 36(5): 105. 1909. PROTOLOGUE: [Mexico] Jalapa
(Bonpland, in herb. Museum Par.); Orizaba (Galeotti, n. 6974 in parte, in herb. Hort. Bot.
bruxell.). Etat de Mexico: volcan de Toluca, 1892 (n. 26 a). TYPE: State of Mexico. In the crater
of the Volcano of Toluca, 13,500 ft. 25.SepU892. Pringle, Plantae mexicanae n° 26a (lectotype,
designated by Munoz (1998a: 354), NY!; isolectotypes, FH!, JE!, PC! 4 replicates; syntypes
["Galeotti 6974"], PC!, TENN!).
Synonymized by Munoz (1998a: 354).
Grimmia herzogii Broth., in Herzog, Biblioth. Bot. 87: 55. 1916. PROTOLOGUE: [Bolivia] Oberes
Chocaya-Tal, alt. 4400 m, No. 3579; Yanakaka, alt. 4500 m, No. 3826. TYPE: An Felsen eines
Gipfel der Yanakakabastion. ca. 4500 m. Leg. Th. Herzog N° 3826. Juli 1911 (lectotype,
designated by Deguchi (1987: 23), JE!; isolectotypes, H-BR!, PC!; syntypes ["Herzog 3579"], HBR!, JE!, PC!).
Synonymized by Munoz (1998a: 354).
Grimmia nigella Herzog, Biblioth. Bot. 87: 55, fig. 17. 1916. PROTOLOGUE: [Bolivia] An Felsen im
oberen Chocayatal, ca. 4400 m, No. 3597 u. 3587/a; an Felsen bei der Saittulaguna, ca. 4300 m,
No. 2679. TYPE: Plantae in itinere secundo per Boliviam lectae. An Felsen bei der Saittulaguna.
ca. 4300m. Leg. Th. Herzog n° 2679. Juni 1911 (lectotype, designated by Munoz (1998a: 354),
JE!; syntypes ["Herzog 3597," "Herzog 3587/a"], JE!, ["Herzog 359T']PC!).
Synonymized by Munoz (1998a: 354).
Grimmia speirophylla Herzog, Biblioth. Bot. 87: 55. 1916. PROTOLOGUE: [Bolivia] An Felsen im
Hochtal von Viloco, ca. 4600 m, No. 3148; an Felsen der Yanakakabastion, ca. 4500 m, No. 3745
u. 3827; an Felsen des Cerro Tunari, ca. 5000 m, No. 4871 u. 4811. TYPE: Plantae in itinere
secundo per Boliviam lectae. An Felsen der Yanakakabastion. ca. 4500m. leg. Th. Herzog n°
3827. Juni 1911 (lectotype, designated by Munoz (1998a: 354), JE!; isolectotype, JE!; syntypes
["Herzog 4871," "Herzog 3148"] JE!, ["Herzog 4811"] NY!, ["Herzog 48J]"] PC!).
Synonymized by Munoz (1998a: 354).
Grimmia speirophylla f. humilis Herzog, Biblioth. Bot. 87: 55. 1916. PROTOLOGUE: [Bolivia] [... ]
aus den Yanakakabergen (sine No.). TYPE: Flora von Bolivia. Torreni-Yanakaka. leg. Th.
Herzog. Juli, 1911 (holotype, JE!).
Synonymized by Munoz (1998a: 354).
Grimmia allionii Broth., Rev. Bryol. 47: 9. 1920. PROTOLOGUE: Provo Azuay. Canar, ad rupes;
3.400 m. TYPE: America meridionalis, Ecuador SW. In rupibus montis prope Canar, 3400 m,
Azuay. 16 n. 1909. Legit Rev. M. Allioni (lectotype, designated by Munoz (1998a: 354), H-BR!;
isolectotype, H-BR!, PC!).
Synonymized by Munoz (1998a: 354).
50
MuNoz AND PANDO
Grimmia afro-ovata Broth. & Ther., Bull. Mus. Hist. Nat. (Paris) 30: 240. 1924. PROTOLOGUE:
Kenya (Mission Gromier-Le Petit, 1911). TYPE: Kenya (Kilimanjaro). Leg. Alluaud. (lectotype,
designated by Munoz (1998a: 354), PC!; isolectotype, H-BR!, PC!).
Synonymized by Munoz (1998a: 354).
Grimmia catalinensis E.B. Bartram, Bryologist 27: 62, pI. 10. 1924. PROTOLOGUE: [U.S.A.] Rock
faces in ravine near top of Mt. Lemon, Santa Catalina Mts., Pima Co., Arizona, 9000 ft. elevation.
E.B. Bartram, No. 387, January 15, 1923. TYPE: Mosses of Southern Arizona. Rocks. Ravine
near top of Mt. Lemon, Santa Catalina Mountains, Pima County. Altitude 9000 ft. Jan. 15 1923.
Collected by Edwin B. Bartram (holotype, FH!; isotypes, FH!, H-BR!, NY!, MO!, PC!).
Synonymized with Grimmia ovalis by Jones (1933: 33), but see Munoz (1998a: 354).
Grimmia catalinensis var. mutica E.B. Bartram, Bryologist 27: 62. 1924. Grimmia ovalis f. mutica
(E.B. Bartram) G.N. Jones, Moss FI. N. Amer. 2(1): 34. 1933. PROTOLOGUE: [U.S.A.] White
House Canyon, Santa Rita Mts., Arizona, February 18, 1923, elevation 6000 ft., No. 683B. TYPE:
Mosses of Southern Arizona. Low boulder. White House Canyon, Santa Rita Mountains, Santa
Cruz County. Altitude 5500 ft. Feb. 18 1923. Collected by Edwin B. Bartram (holotype, FH!).
Synonymized by Munoz (1998a: 354).
Grimmia somervellii Dixon, J. Bot. 63: 221. 1925. PROTOLOGUE: [Nepal] Rongbuk Valley, 16,00017,000 ft. (1001 [holotype)); 16,000-17,500 ft. (1004, 1005); on soil, 16,500-17,500 ft., 15 miles
N. of Base Camp, near Lamna La Pass (1006) [Somervell]. TYPE: Flora of Mt. Everest. 1600017500 ft. Collector J.H. SomervelI. Date 1924 (paratype, BM!).
Grimmia longipes Broth., Kryptogamae exsiccatae editae a Museo Palatino Vindobonensi n° 2980.
1926, nom. nud. in sched. ORIGINAL SPECIMEN: China (prov. Yiinnan bor.-occid.): in rupibus
diabasicis regionis temperatae prope vicum Nguluko ad urbem Lidjiang ("Likiang"), ca. 3000 m
s.m., m. Jun. c.fr. vet., m. Oct. c.fr. mat. leg. H. Handel-Mazzetti. specim. originale (F!, H-BR!,
MA-7094!, PC!).
Grimmia ovalis f. gracilis H. Moller, Ark. Bot. 26A(2): 39. 1934. PROTOLOGUE: [Sweden]
Vasterbotten. Umea, Esmarksberget 1896 C. P. Laestadius. [... ] Lycksele lappmark. Stensele.
Sandvik 1924 Sm. TYPE: Viisterbotten. Umea, Esmarksberget. 13.9.1896 C. P. Laestadius
(lectotype, here designated, S!; syntype ["Lycksele lappmark. Stensele, Sandvik. 217 1924. Carl
Stenholm"], S!).
Grimmia trol/ii Herzog, Hedwigia 74: 102. 1934. PROTOLOGUE: Bolivia: Curahuara (n. 58), Gipfel
des Cerro de la Esperanza, Cordillera Real, 4500 m (n. 72) [CO Troll]. TYPE: Fl. v. Bolivia.
Curahuara. leg. K. Troll n° 58 (lectotype, designated by Deguchi (1987: 23), JE!).
Synonymized by Deguchi (1987: 23) with G. herzogii, but see Munoz (1998a: 354).
Grimmia cinerea Ther., Rev. BryoI. LichenoI. 9: 9. 1936. PROTOLOGUE: [Ecuador, leg. R. Benoist]
Condorguachana, rochers, 4.150 m (n° 3153). TYPE: 3153. Equateur, rochers du Condorguachana.
R. Benoist 4.x1.1930 (holotype, PC!; isotypes, PC! 2 replicates).
Synonymized by Churchill (1994: 2).
Grimmia stenopyxis Ther., Rev. BryoI. LichenoI. 9: 8. 1936. PROTOLOGUE: [Ecuador, leg. R.
Benoist] Base du Pichincha, sur rochers d'andesite, 3.200 m (nO 3076, 2276); Rumipamba, a terre
WORLD SYNOPSIS OF GRIMMIA
51
dans les broussailles, 3.000 m (nO 2957); Environs de Quito, 2.800 m (nO 2281). TYPE: Equateur:
base du Pichincha. rocher. R. Benoist 3076. 24.X.1930 (lectotype, designated by Munoz (1998a:
354), PC!; isolectotype, F!; syntypes ["Benoist 2281"] F!, ["Benoist 2957"] NY!).
Synonymized with G. affinis by Cao & Churchill (1995: 507).
Grimmia sumatrana Dixon, Ann. Bryol. 12: 50. 1939. PROTOLOGUE: [Indonesia] G. Losir, on rock,
3250-3500 m, 4 Feb. 1937; colI. Van Steenis (10152); Rh. Bog (4035) [Buitenzorg Botanical
Garden]. TYPE: (syntype, Deguchi (1986: 328-329), L not seen).
Synonymized by Deguchi (1986: 328-329).
Grimmia antillarum Ther., Rev. Bryol. Lichenol. 13 (Trav. Husnot, 2): 13. 1944. PROTOLOGUE:
[Dominican Republic] Santo-Domingo: Cordillera Central, provo de Azua, Los Vallecitos de
Yaque, 2.500 m (nO 13.630) [leg. M.E.L. Ekman, 1929-1930]. TYPE: Plantae Indiae Occidentalis
nO 13630. Hispaniola. Civ. Santo Domingo: Cordillera Central, provo de Azua, Los Vallecitos de
Yaque, c. 2500 m. 2.x.1929.leg. E.L. Ekman. det. Ther. (lectotype, designated by Munoz (1998a:
354), PC!; isolectotypes, F!, NY!, PC!).
Synonymized by Munoz (1998a: 354).
Grimmia akaisialpina Takaki, Bot. Mag. (Tokyo) 64: 180, fig. 4. 1951. PROTOLOGUE: [Japan]
South Alps; Sensui pass (2700 m, no. 10137, Aug. 15, 1950). TYPE: (holotype, Deguchi (1978:
158), private herbarium of Takaki, not seen).
Synonymized by Takaki et al. (1970: 186-187).
Grimmia ovata f. glacialis P. de la Varde, Ark. Bot., n.s. 3: 147. 1955, nom. nud. ORIGINAL
MATERIAL: Teleki Valley, on frost-lifting ground, 4200 m (1711: forma glacialis); Nairobi
Valley, 4250 m on frost-lifting ground (1909: f. glacialis). ORIGINAL SPECIMEN: Kenya
Colony: Mt. Kenya, Teleki Valley, alpine regime on frost-lifting ground in the upper part of the
valley. 4200 m.s.m. 27.7.1948 ColI. Olov Hedberg 1711. Det. PV 11909 (PC!).
Grimmia meridensis Varesehi, Flora de los Paramos de Venezuela: 87. 1970, nom. nud. ORIGINAL
MATERIAL: [Venezuela] Andes. [... ] eima del Paramo de las Piedras Blancas en una altitud de
4.762 m.
Grimmia maida Greven, Bryologist 99: 428, fig. 1. 1996. PROTOLOGUE: Africa. La Reunion. Viewpoint Ie Mai"do. at the end of RF 8, along trail to Grand Benare, alt. 2150 m, 1995. 10.07, H. C.
Greven & S. N. Khoebal 4000.1. TYPE: Ile de la Reunion. Le Mai"do, at the end of R.F. 8, alt.
2150 m. Leg. H.c. Greven & S.N. Khoeblal 4000/1. 7 October 1995 (holotype, L not seen;
isotypes, MO!; paratype, MO! ["M.R. Crosby & CA. Crosby 8430"]).
Synonymized by Munoz (1998a: 354).
DISTRIBUTION: ARG (I.M. Lamb 5288, FH); AUS (C Beauglehole 14712, MEL); AUT (CF
Hornschuch s.n., B); BOL (M. Lewis 79-26, MO); BRA (D.M. Vital 1708, FLAS); BTN (W Griffith
127, MO); CAN (G.R. Brassard 15284, FH); CHE (P. Culmann s.n., MA); CHL (R. Santesson 786,
FH); CRN (B.C Tan 93-1184, FH); CMR (J.P.M. Brenan in P.W Richards 4269, MO); COL (A.M.
Cleef 1798, COL); CRI (M. Crosby 3917, TENN); CZE (J. Duda s.n., BP); DEU (J.C Zenker & FD.
Dietrich s.n., FH); DOM (A. & P. Liogier 25032, FH); ECU (B. L£itnant 15345, MO); ESP (J.L.
Rupidera 1, SALA); ETH (D.A. Petelin 37-7, MO); FIN (E.F Lackstrom & VF Brotherus s.n., IE);
FRA (J.-P. Frahm 935069, MA); GBR (FB. White s.n., NY); GEO (CC Townsend 851860, ALTA);
52
MuNoz AND PANDO
GRL (We. Steere 62-332, NY); GTM (A.J. Sharp 4771, TENN); HND (B. Allen 12260, MO); IND (J.
Doulea 3110, H-BR); ISL (B. J6hannsson 12769, ICEL); ITA (G. Venturi s.n., FH); JPN (M. Mizutani
13651, MO); KAZ (B. Allen 10849, MA); KEN (e. Troll 5570, JE); LKA (e. Ruinard & A.H.M.
Jayasuriy 17140, MO); LSO (R.E. Magill 4186, MO); MDG (G. Cremers 1705, MO); MEX (A.
Cardenas s. 5344, MO); MNG (W Schubert M183, MO); MWI (N.G. Hodgetts M2123A, MA); MYS
(B. Tan 89-521, NY); NOR (D.H. Norris 69524, MO); NPL (A. Zimmermann s.n., G); NZL (J. Child
M573, F); PAK (J.F. Duthie s.n., H-BR); PER (E. Hegewald 5459, MO); PHL (B.e. Tan 82-195, FH);
PNG (A. Eddy 2519, BM); POL (G.F. Kaulfuss s.n., BM); PRT (H. Persson s.n., NY); REU (J.L. De
Sloover 17934, MO); SFE (B.e. Tan 97-492, FH); SIB (E. Ignatova 131110, MA); SUN (J. V
Chernyadjeva 53, LE); SVK (z. Pilous s.n., BP); SWE (R. W. Hartman s.n., H); TKM (G.J. Potanin
s.n., LE); TWN (M.-J. Lai 6886, MO); TZA (R.E. Gereau 1649, MO); UGA (G.H.s. Wood 1107, PC);
USA (F.A. MacFadden 8036, MO); VEN (D. Griffin III et al. 2115, FLAS); ZAF (R.E. Magill 6607,
MO).
34. Grimmia macroperichaetialis Greven, Bryologist 101: 100-102. 1998. PROTOLOGUE:
Australia. Victoria. Mt. Buller, summit, 1790 m, 14 March 1997, Greven Aus. 35 [paratypes
listed]. TYPE: Australia. Victoria. Mt. Buller, summit, 1790 m, 14 March 1997, Greven Aus. 35
(holotype, CANB not seen; isotype, MO!).
DISTRIBUTION: AUS (e. Beauglehole 15833, MEL).
35. Grimmia macrotheca Mitt., J. Linn. Soc., Bot. Suppl. 1: 44. 1859. PROTOLOGUE: [India] In
Himalayae orient. reg. temp., Sikkim, J.D. Hooker! (No. 316). TYPE: 316. Jongui. alt. 12.000. Dr.
Hooker (holotype, NY!; isotypes, BM! 2 replicates).
Grimmia longicapsula C.H. Gao & T. Cao, Acta Bot. Yunnan. 3: 395, tab. 5 figs. 1-8. 1981.
PROTOLOGUE: [Tibet] Xizang: Districtus Yadong Xian, ad ropes, 28.V.1975, Zang Mu 39
(Typus in ISPH, Isotypus in HKAS). TYPE: Tibet. Yadong County. Ji-Neou. Zang Mu 28 V '75,
n° 39 (holotype, IFSBH!; isotype, NY!).
Synonymized by Cao & Vitt (1986: 196).
DISTRIBUTION: CRN (s. He 31583, MO).
36. Grimmia mammosa C.H. Gao & T. Cao, Acta Bot. Yunnan. 3: 394, tab. 4 figs. 1-9. 1981.
Grimmia mammillaria X.I. Li & M. Zang, Bryoflora of Xizang: 138. 1985, nom. inval., orthogr.
err. PROTOLOGUE: [Tibet] Xizang: Districtus Yadong Xian, ad ropes, alt.s.m. 2900 m,
29.V.1975, Zang Mu 71 (Typus in ISPH, Isotypus in HKAS). TYPE: Tibet. Yadong Xian, ChunPei-River. on rock. 2900 m. Zang Mu 29 V '75, n° 71 (holotype, IFSBH!; isotype, NY!).
DISTRIBUTION: CRN (W.-X Xu 42, ALTA); MWI (M.J. Wiggington M1298, MA).
37. Grimmia mariniana Sayre, Bryologist 58: 323. 1955. PROTOLOGUE: [U.S.A.] California,
Marin County. On exposed, north-facing rocks, summit of Mt. Tamalpais at 3500 ft. Type
Specimen: w.e. Steere, 8 April 1953. In herbaria of Stanford University, the New York Botanical
WORLD SYNOPSIS OF GR/MMIA
53
Garden and the author. Exsiccati: Two further specimens from Mt. Tamalpais, as follows: L.F.
Koch 2274, 10 August 1947; IT. Howell, 3 April 1949 (communicated by Koch). Also from Mt.
S. Helena, Napa County, California: L.F. Koch 2125, 7 July 1947. TYPE: California. Marin
County [ ... ] just under summit of Mt. Tamalpais [... ] April 8, 1953. W.C. Steere (lectotype, here
designated, NY!).
DISTRIBUTION: USA (We. Steere s.n., NY).
38. Grimmia molesta J. Munoz, Ann. Missouri Bot. Gard. 86: 152. 1999. PROTOLOGUE and TYPE:
Peru. Arequipa: Arequipa, am Weg nach Puno, Hegewald & Hegewald 5480 (holotype, MO!;
isotypes, H!, MA!, NY!).
DISTRIBUTION: PER (P. Hegewald & E. Hegewald 5480, MA).
39. Grimmia mollis Bruch & Schimp., Bryol. Europ. (fasc. 42) 3: 133, tab. 253. 1849. Grimmia mollis
f. emersa Lorentz, Moosstud.: 93. 1864, nom. inval. Hydrogrimmia mollis (Bruch & Schimp.)
Loeske, Stud. Morph. Syst. Laubm.: 108. 1910. PROTOLOGUE: [Norway] In alpe Goustafjeld
provinciae norvegiacae Tellemarken H. Holmgren legit et misit. TYPE: (BM not seen).
Grimmia mollis f. xerotrophica Gyorffy, in Bauer, Musci europaei exsiccati, Schedae und
Bemerkungen zur 33 Serie, n° 1612. 1924. Hydrogrimmia moWs f. xerotrophica (Gyorffy) Podp.,
Consp. Musc. Eur.: 274. 1954. PROTOLOGUE and TYPE: [Slovakia] Hohe Tatra (Montes
Carpathi): Klein Kolbachtal, Funfsee, auf Granitplatten des Abflusses des zweiten Sees in den
dritten, 1980 m s.m., 4.Sept.l922 legit I. Gyorffy (lectotype, here designated, MO!; isolectotype,
FH!).
Grimmia evansii E. Britton, Rhodora 1: 148, pI. 7. 1899. PROTOLOGUE: [U.S.A.] Tuckerman's
Ravine, Mt. Washington, N.H. [New Hampshire], alt. 1,230 m. July 30, 1890. Collected by
Alexander W. Evans of Yale University, [... ] TYPE: Tuckerman's Ravine, Mt. Washington, N.H.
A.W.E. July 30, 1890 (holotype, NY!).
Synonymized by Allen (1995: 160).
DISTRIBUTION: AUT (J. Breidler s.n., JE); BEL (V Payot s.n., JE); BGR (L. Meinunger 14306,
JE); CAN (T.A. Hedderson 2041, FH); CHE (T. Herzog 352, JE); DEU (W Kruger s.n., JE); ESP (V
Allorge s.n., PC); FRA (J.P. Hebrard 4-9882, MO); ITA (0. Venturi s.n., FH); MAR (M.J. Cano et al.
s.n., MA); NOR (R. W Hartman s.n., H); POL (H. Bednarek-Ochyra & R. Ochyra 127190, MO); SFE
(C. Wright 73, FH); SIB (M Ignatov 01466, MA); SVK (J. GyorffY s.n., MO); SWE (H. Moller s.n.,
MO); USA (F.J. Hermann 20740, RM).
40. Grimmia montana Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 128, tab. 250. 1845.
Guembelia montana (Bruch & Schimp.) Hampe, Bot. Zeitung (Berlin) 4(8): 125. 1846. Grimmia
holmiensis Lindb. ex Hartm., Handb. Skand. FI. ed. 7: 374. 1858, nom. inval., pro synon.
PROTO LOGUE: In rupibus schistosis, basalticis et porphyraceis: prope Angers Galliae
occidentalis (Guepin) [France], in valle badensi Murgthal dicta (Alex. Braun) [Germany], in
Hassia superiore prope Biedenkopf (Bruch) [Germany], in Vogesi monte Donnersberg (Gumbel)
54
MuNoz AND PANDO
[Germany], prope Christianiam (Blytt) [Norway], in alpe Dovrefjeld Norvegiae (W.P. Sch.)
[Norway]. TYPE: Felsen auf dem Donnersberg. Gilmbel. April 1843 (lectotype, designated by
Cao & Vitt (1986), BM!).
Grimmia /axa Mull. Hal., Bot. Zeitung (Berlin) 5: 801. 1847. Guembelia /axa (Milil. Hal.) Milil. Hal.,
Syn. Musc. Frond. 1: 771. 1849. PROTOLOGUE: Mexico in monte Orizabae, ubi legerunt Deppe
et Schiede. Cf. eor. Collect. No. 1070a. TYPE: Mexico in monte Orizabae. Deppe et Schiede 19re.
(lectotype, designated by Munoz (1998b: 389), PC!, isolectotype, PC-Besch!).
Synonymized by Munoz (1998b: 389).
Grimmia schiedeana Milll. Hal., Bot. Zeitung (Berlin) 13: 765. 1855. Grimmia schiedei Miill. Hal. ex
Kindb., Enum. Bryin. Exot.: 61. 1888, nom. inval., orthogr. err. PROTOLOGUE: Mexico: Deppe
et Schiede in locis haud designatis. TYPE: Mexico. Leg. Deppe et Schiede (lectotype, designated
by Munoz (1999a: 152), BM!; isolectotype, PC!).
Synonymized by Munoz (1999a: 152).
Grimmiafragilis Schimp., Syn. Musc. Eur. ed. 2: 257. 1876, nom. illeg., non F. Weber, 1804. Grimmia
montana var.fragilis (Schimp.) Loeske, Laubm. Eur. Part I: 99. 1913. PROTOLOGUE: [Portugal]
ad rupes granitic as in alpestribus provinciae lusitanicae Beira, ubi d. Welwitsch anno 1848 legit.
TYPE: ad rupes granit. in alpestribus Provo Beira. Aug. 1848. L. Welw. (lectotype, designated by
Munoz (1998b: 389), BM!).
Synonymized by Munoz (1998b: 389).
Guembelia tene//a Milil. Hal., Bot. Centralbl. 44: 388. 1890. Grimmia tene//a (Miill. Hal.) Kindb.,
Enum. Bryin. Exot., Suppl. 2: 107. 1893. PROTOLOGUE: [U.S.A.] Idaho, Coeur d'Alene, 6.
Aug. 1888. Dr. Roll [... ]. TYPE: 6 Aug 1888 Coeur d'Alene. Dr. Julius ROIL Idaho (lectotype,
designated by Munoz (1998b: 390), G!; isolectotypes, H-BR!, JE!, S!).
Synonymized by Munoz (1998b: 389).
Grimmia montana var. longifolia Cardot, in Gasilien, Rev. Bryol. 21: 24. 1894. Grimmia montana f.
longifolia (Cardot) Podp., Consp. Musc. Eur.: 281. 1954. PROTOLOGUE: [France] Granit
decouvert, sommet de Pierre-sur-Haute (1600 m. alt.).- Fertile. TYPE: Auvergne, Pierre sur Haute
1884 (lectotype, designated by Munoz (1998b: 390), TRH!).
Synonymized by Munoz (1998b: 389).
Grimmia arctophila Kindb., Rev. Bryol. 23: 18. 1896. PROTOLOGUE: Amer. Greenland: Berggren.
Canada, Cypress hills: Macoun. TYPE: Plantae in itineribus Suecorum polaribus collectae.
Groenlandia boreal.: Claushavn 1870. S. Berggren (lectotype, designated by Munoz (1999a: 152),
S!).
Synonymized by Munoz (1999a: 152).
Grimmia pilosissima Herzog, Rev. Bryol. 38: 12. 1911. PROTOLOGUE: [Italy] Sardaigne; sur les
rochers du versant oriental de la crete du Monte Gennargentu, vers 1750 m., 2 juin 1906. TYPE:
An Felsen der Ostseite der Punta la Marmora (Mte. Gennargentu). Leg. Th. Herzog 2.VI.06. ca.
1750m (holotype, JE!).
Grimmia montana var. abnoba H. Schmidt, Mitt. Bad. Landesvereins Naturk. Naturschutz Freiburg 2:
121, fig. 6. 1927. PROTOLOGUE: [Germany] Schauinsland: Gneisfelsen nahe dem hOchsten
WORLD SYNOPSIS OF GRIMMIA
55
Gipfel und bei den unteren Hausem von Hofsgrund (S.). TYPE: Sud. Schwarzwald: Felsen nahe
sudweste dem Schauinsiandgipfei. Schmidt (lectotype, designated by Munoz (1998b: 390), JE!).
Synonymized by Munoz (1998b: 389).
Grimmia montana f. submutica J.E. Zetterst. ex H. Moller, Ark. Bot. 26A(2): 31. 1934.
PROTOLOGUE: [Sweden] In saxis juxta Husbyborg prope Upsaliam 15/5 1855. [Zetterstedt,
Grimmiae et Andreaeae exsiccatae, n° 21c]. TYPE: In saxis juxta Husbyborg prope Upsaliam 15/5
1855. [Zetterstedt, Grimmiae et Andreaeae exsiccatae, n° 21c] (lectotype, designated by Munoz
(1998b: 390), H-BR!).
Synonymized by Munoz (1998b: 389).
DISTRIBUTION: AND (C Casas s.n., BCB); AUT (H. Gander s.n., TRH); BEL (A. Cornet s.n.,
NY); CAN (J Macoun s.n., FH); DEU (S. Caspari & J. Heinrichs 2526, MO); ESP (J.L. Rupidera 49,
SALA); FRA (F.M.L. Corbiere s.n., FH); GBR (CH. Binstead s.n., FH); GRL (w.C Steere 62-621,
NY); ISL (B. J6hannsson ]]813, ICEL); ITA (F. Kern s.n., WRSL); MEX (A.J. Sharp 4289, TENN);
NOR (JE. Zetterstedt s.n., FH); POL (K.G. Limpricht s.n., FH); PRT (F.M.J. Welwitsch 53, BM);
SWE (S. Arnell s.n., MA); TUR (E. Nyholm & A.C Crundwell 674, F); TZA (G.H.S. Wood 1730,
MA); USA (A.T. Whittemore 4277, MO).
41. Grimmia muehlenbeckii Schimp., Syn. Musc. Eur.: 212. 1860. Grimmia trichophylla subsp.
muehlenbeckii (Schimp.) Boulay, Musc. France 1: 380. 1884. Grimmia trichophylla var.
muehlenbeckii (Schimp.) Husn., Muscoi. Gall.: 135. 1887. Dryptodon muehlenbeckii (Schimp.)
Loeske, Stud. Morph. Syst. Laubm.: Ill. 1910. PROTOLOGUE: [Austria] in rupibus graniticis
regione subalpinae: prope Trafoi Tyrol merid. ubi aestate 1840 [... ] Muhlenbeck reperi. TYPE:
Trafoi. Jui. 40 (holotype, BM!).
Grimmia rugulosa Lindgr., Bot. Not. 1845: 88. 1845, nom. nud. ORIGINAL SPECIMEN: [Sweden]
Upsaliae 1834. Lindgr. (BM!).
Grimmia trichophylla var. septentrionalis Schimp., Coroll. Bryoi. Eur.: 47. 1856. PROTOLOGUE:
[Sweden] In sylvaticis circa Holmiam et Upsaliam abundanter legi. TYPE: Upsala. Schimper
(syntype ["Upsala"] JE!).
Grimmia hermannii Crum, Michigan Bot. 13: 171. 1974. PROTOLOGUE: [U.S.A.] On boulder in
woods below Falls, Manganese River gorge, Copper Harbor, Keeweenaw [sic] County, F.J.
Hermann 25446, July 14, 1973, type in herb. MICH, duplicates in herb. US, NY, DUKE, F, CAN,
WTU. TYPE: Upper Peninsula of Michigan. Keweenaw County. On boulder in woods below
Falls, Manganese River gorge, Copper Harbor. F.J. Hermann 25446. July 14, 1973 (isotypes,
CANM-226826!, F!, FH!).
Synonymized by Munoz (2000c).
DISTRIBUTION: CAN (G.R. Brassard 10990, NY); CHE (P.G.M. Rhodes 267, MO); DEU (J
Heinrich & C Schmidt JH3740, GOET); ESP (C Casas s.n., MA); FIN (S.o. Lindberg s.n., MO);
FRA (R. Spruce s.n., NY); GEO (G. Woronow 77a, PC); HUN (L. Marstaller s.n., JE); ITA (G.
Venturi s.n., FH); LVA (J Treboux s.n., PC); NOR (P. Stermer s.n., BCB); POL (R. Ochyra s.n., MO);
56
MuNoz AND PANDO
PRT (c. Sergio & S. Fontinha s.n., LISU); SIB (E.A. Selivanova-Gorodkov 2460, LE); SUN (M.
Ignatov & A. Bezgodov 244, NY); SWE (J.E. Zetterstedt s.n., H); TUR (P. Uotila 35659, H-BR); USA
(B. Allen 10358, MO).
42. Grimmia navicularis Herzog, Beih. Bot. Centralbl. 26(2): 65. 1909. PROTOLOGUE: Bolivia: An
Felsen in der Blockhalde fiber dem Tunarisee (Prov. Cochabamba), ca. 4400 m; Januar, 08, steril;
an FeIsen bei der Abra de San Benito (Prov. Cochabamba), ca. 3900 m; Januar, 08, c. thecis
vetustis. TYPE: Flora von Bolivia. An Felsen des Abra de San Benito (Prov. Cochabamba) ca.
3900m. T. Herzog. Januar, 08 (lectotype, designated by Deguchi (1987: 27), JE!; syntypes
["Cochabamba: fiber dem Tunarisee"] JE!, PC!).
Grimmia paramophila Broth., in Meyer, In den Hoch-Anden von Ecuador: 518. 1907, nom. nud.
ORIGINAL MATERIAL: Ecuador. Chimborazo: Paramo-region, bei 5000 m. SeehOhe. N.N.W.
Seite. 1903.VII leg. Prof. Dr. Hans Meyer.
Grimmia biplicata Broth., in Meyer, In den Hoch-Anden von Ecuador: 518. 1907, nom. nud.
Grimmia chilensis Ther., in Herzog, Darwiniana 11: 217. 1957. PROTO LOGUE: Rep. Argentina:
Cerro Lopez, 1700 m, leg. A. Donat, n° 114. TYPE: Parque Nac. Arg. Nahuelhuapi no. 114. Cerro
Lopez, 1700 m. Leg. A. Donat, l.XI.36 (lectotype, designated by Munoz (1999a: 152), JE!;
isolectotype, JE!).
Synonymized by Deguchi (1987: 28).
DISTRIBUTION: ARG (A. Donat 114, JE); BOL (M. Lewis 79-1731, F); COL (A.M. Cleef6965,
NY); ECU (H. Meyer 5478, H); PER (E. Hegewald 6001, MO); VEN (D. Griffin III et al. 1318,
FLAS).
43. Grimmia nepalensis Mitt., J. Linn. Soc., Bot. Suppl. 1: 46. 1859. PROTOLOGUE: In Nepal
orient. reg. alp., J.D. Hooker! (No. 248, ex parte). TYPE: 248 partly. E Nepal alt. 14500 pd. J.D.
Hooker (holotype, NY!).
Grimmia inflectens Mitt., J. Linn. Soc., Bot. Suppl. 1: 43. 1859. PROTOLOGUE: In Nepal orient. reg.
alp., J.D. Hooker! (No. 248, ex parte). TYPE: East Nepal, 14500 ped. 248 in part. J.D. Hooker
(lectotype, here designated, NY!).
DISTRIBUTION: IND (J.D. Hooker 325, NY); NPL (J.D. Hooker 248, NY).
44. Grimmia nivalis Kindb., Bull. Torrey Bot. Club 17: 271. 1890. PROTOLOGUE: [U.S.A.] On
rocks on the summit of the Gold Range, north of Griffin Lake, B.c. Alt. 7,000 feet. Aug. to, 1889.
TYPE: N. Amer. Brit. Columbia, Gold Range, rocks, 7000 f. 10/8 89 J. Macoun (lectotype,
designated by Munoz (1998b: 392), S!; isolectotypes, CANM-198090!, H-BR!).
WORLD SYNOPSIS OF GRiMMIA
57
Grimmia papillinervis Kindb., Rev. Bryol. 34: 89. 1907. PROTOLOGUE: Canada, Brit. Col., Skagit
summit: J.M. Macoun 1905. TYPE: Canada, Brit. Col., Skagit summit, dry rocks. 18/7 05. J.M.
Macoun (holotype, Munoz (1998b: 392), S!; isotype, FH!).
Synonymized with G. tenerrima by Allen (1995: 162-164), but see Munoz (l998b: 392).
DISTRIBUTION: CAN (J.M. Macoun s.n., S); USA (J.T. Howell s.n., FH).
45. Grimmia nutans Bruch, in F.A. Mull., Flora 12: 390, tab. 8. 1829. PROTOLOGUE: [Turkey] Auf
Felsen bei Smyrna. TYPE: Smyrna (lectotype, here designated, H-SOL!; isolectotype, NY!).
Grimmia verticillatula Ther. & Trab., Bull. Soc. Hist. Nat. Afrique N. 21: 29. 1930. PROTOLOGUE:
[Morocco] in rupibus arenaceis Atlantis Majoris: in ditione Glaoua supra pagurn Zerekten, ad alt.
1400--1600 m (Dr. R. Maire, 1926). TYPE: Algerie [sic, the locality is in Morocco). Grand Atlas.
Zerekten 1600 m. Legit R. Maire n° 2055 (lectotype, here designated, PC!).
Grimmia meteorae C.C. Towns., in K. Tan, The Davis & Hedge Festschrift: 48. 1989. PROTOLOGUE
and TYPE: Greece. Thessaly: on damp stony bank near the Varlaam Monastery, the Meteora, near
Kalabaka, apparently calcareous, 10 May 1987, C.C. Townsend 87/125 (E, holotype; Herb.
Townsend, isotype) (isotype, NY!).
DISTRIBUTION: CYP (H. Greven s.n., MA); GRC (C.c. Townsend 871125, NY); MAR (R.
Maire s.n., PC); TUR (Anonymous s.n., H-SOL).
46. Grimmia obtusifolia C.H. Gao & T. Cao, Acta Bot. Yunnan. 3: 394, tab. 4 fig. 10--16. 1981.
PROTOLOGUE: [Tibet] Xizang: Districtus Shuanghu Xian, ad rupes, alt.s.m. 4850 m,
13.VII.l978, Lang Kai-Yong 1347 (Typus in ISPH, Isotypus in HP); Bangkog Xian, ad rupes,
alt.s.m. 4800 m, 5.VI.l976, Lang Kai-Yong 1232. TYPE: Tibet, Shuanghu Xian, Lang 1347
(holotype, IFSBH!; paratypes, IFSBH!, MO! ["Lang 1232")).
DISTRIBUTION: CRN (P.L. Redfearn Jr. 35436, MO).
47. Grimmia ochyriana J. Munoz, Nova Hedwigia 66: 235-240. 1998. PROTO LOGUE and TYPE:
Nepal. Pres du Camp de Base, direction de Lobuje, 5100 m, 31-V-1952, Zimmennann 558
(holotype, G!; isotype, MA-5212!).
Grimmia mexicana Greven, Bryologist 102: 432, fig. 2. 1999. PROTOLOGUE: Mexico. Puebla. Pico
de Orizaba, track to Piedra Grande, elev. 3,870 m, sunny S-facing andesite rock, 4 Dec. 1997.
H.C. Greven MIOO (Holotype, BM; isotypes ENCB, FH, MEXU, MICH, NY). TYPE: Mexico.
Puebla: volcan Orizaba, 2.5 km below Piedra Grande, 3890m, 4 Dec. 1997. H.C. Greven M100
(isotypes, MEXU!, NY!).
NOTE: The holotype is not deposited in BM yet (L.T. Ellis, pers. comm.).
DISTRIBUTION: GTM (A.J. Sharp 5428, TENN); MEX (A.J. Sharp 4270, TENN); NPL (A.
Zimmermann 558a, MA).
58
MuNoz AND PANDO
48. Grimmia olneyi Sull., in Sullo & Lesq., Musc. Bor.-Amer.: 32. 1856. PROTOLOGUE and TYPE:
[U.S.A.] in rupibus maritimis Rhodensis Insulae (cl. S.T. Olney legit). [Sullivant & Lesquereux,
Musci borali-americani exsiccati n° 141] (lectotype, here designated, FH!; isolectotypes, F!, MO!,
NY!).
Grimmia austin;; Kindb., Rev. Bryol. 23: 19. 1896 ["Austini"], PROTOLOGUE: Amer. Unit. St., N.
Jersey: Austin, com. Macoun. TYPE: N. Amer., New Jersey: Austin. com. Macoun (lectotype,
here designated, S!).
DISTRIBUTION: CAN (JM. Gillet 17125, CANM); USA (AJ Sharp 35255, MO).
49. Grimmia orbicularis Bruch ex Wilson, Engl. Bot. Suppl. 4: 2888. 1844. Guembelia orbicularis
(Wilson) Hampe, Bot. Zeitung (Berlin) 4(8): 125. 1846. PROTOLOGUE: [United Kingdom]
Found on Orrneshead in 1826 by the writer of this article, and subsequently by F.K. Eagle, Esq.,
on St. Vicent's Rocks. For recent specimens from the latter place we are indebted to Mr. Thwaites,
who gathered them in March 1843. TYPE: St. Vicent's Rocks. Feb. 1837 (Eagle) (lectotype, here
designated, BM!).
NOTE: The label of the specimen selected as lectotype by Geissler & Maier (1995) reads:
"Grimmia orbicularis BruchlGrirnmia africanal(Fissid. africanus Hedw. Sp. M.)/St. Vicent's Rocks
near Bristol/(Mr. Stephens)/discovered by Mr. Eagle in 1837. W.W." (BM!). It clearly states
"Stephens" as collector and not "Eagle." Stephens's name did not appear in the protologue of Grimmia
orbicularis, and hence this specimen cannot be designated as lectotype (ICBN Art. 9.13).
Grimmia sinaica Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 113. 1845. Guembelia sinaica
(Bruch & Schimp.) Hampe, Bot. Zeitung (Berlin) 4(8): 124. 1846. PROTOLOGUE: [no locality
given]. TYPE: in summo monte Sinai [Egypt] (lectotype, here designated, BM!; isolectotypes,
BM! 4 replicates, PC-Besch!).
Grimmia mammilaris Poech, in Keil, Lotos 1: 177. 1851. PROTOLOGUE: [Czech Republic] Gemein
bei St. Prokop und Kuchelbad nachst Prag. TYPE: In rupibus calcareis prope Pragam. Mat. Vere.
Dr. Poech. [Poech, Musci bohemici, nO 29. 1846] (lectotype, here designated, G!).
Grimmia orbicularis var. africana Besch., Cat. Mouss. Alg.: 17. 1882, nom. nud. Grimmia orbicularis
Avanc. Sci. 34: 482. 1905. Grimmia
var. bescherellei Corb., Compt.-Rend. Assoc. ｆｲ｡ｮｾＮ＠
pulvinata var. bescherellei (Corb.) Wijk & Margad., Taxon 9: 190. 1960. PROTOLOGUE:
[Algerie, Constantine] Chettabah; pres l'usine Micout, 30 dec. 1903 [Leg. R. Chudeau]. TYPE:
Constantine: usine Micout, 30 dec. 1903. Leg. R. Chudeau (lectotype, here designated, CHE!;
isolectotypes, CHE! 3 replicates).
Grimmia pulvinatula Mull. Hal., Bot. Jahrb. Syst. 5: 81. 1884. PROTOLOGUE: Kerguelen [Foundery
branch, Nov. 1874, Naumann]. TYPE: Kerguelen. D. Naumann 1874 (lectotype, here designated,
PC!).
Grimmia stolonifera Mull. Hal., Bot. Jahrb. Syst. 5: 81. 1884. PROTOLOGUE: Kerguelen [Foundery
branch, Nov. 1874, Naumann]. TYPE: Kerguelens Land. Dr. Fr. Naumann. Dcber 1874 (lectotype,
here designated, PC!).
WORLD SYNOPSIS OF GRiMMIA
59
Grimmia orbicularis var. persica Schiffn., Oesterr. Bot. Z. 47(4): 129. 1897. PROTOLOGUE: [Iran]
Persiae australis provo Faristan: in rupestribus calcareis montium ad Schiras. 1650-1700 m.s.m.
25.X.1892. c.fr. (Bornm. Iter Pers.-turc. Nr. 4448-4448b). - Persiae australis provo Faristan:
Persepolis in decliv. apricis. ±1800 m s.m. 3.xI, 1892; c.fr. (Bornm. iter Pers.-turc. Nr. 4458). Persia borealis: ad Teheran in planitie. ±1200 m s.m. 31.1.1892; c.fr. (Bornm Iter Pers.-turc. Nr.
4452). TYPE: J. Bornmiiller: Iter Persico-turcicum 1892-93. N° 4448. Persiae australis provo
Faristan: in rupestribus calcar. montium ad Schiras. 1650-1700 m.s.m. 1892.X.25 legit et deterrn.:
1. Bornmiiller (lectotype, designated by Munoz (2000c), JE!; isolectotypes, F!, PC!).
Synonymized by Munoz (2000c).
Grimmia microglobosa Miill. Hal., Hedwigia 37: 160. 1898. PROTOLOGUE: [New Zeeland] Nova
Seelandia, insula australis, North Canterbury, Ben More, ad rupes alt. 5000 ped., Nov. 1890: T.W.
Naylor Beckett, qui 1892 misit. TYPE: On rocks Ben More alt. 5000' N. Canterbury. T.W.N.B.
November 1890 (lectotype, here designated, H-BR!).
Grimmia orbicularis var. brevipila J.1. Amann, Fl. Mouss. Suisse 2: 134. 1918. Grimmia orbicularis f.
brevipila (1.1. Amann) Podp., Consp. Musc. Eur.: 285. 1954. PROTOLOGUE: [Switzerland]
Vaud: Saint-Saphorin, murs de vignes, 400-450 m (BH 91.6.20 et 22) (Amann). TYPE:
[Switzerland] Vaud: Saint-Saphorin, 400-450 m, 18.XII.1912. Leg. Amann (BH 91.6.22)
(lectotype, here designated, Z!; isolectotype, PC!; syntype ["Amann BH 91.6.20"], Z!).
Grimmia orbicularis var. humilis J.1. Amann, Fl. Mouss. Suisse 2: 134. 1918. Grimmia orbicularis f.
humilis (1.1. Amann) Podp., Consp. Musc. Eur.: 285. 1954. PROTOLOGUE: [Switzerland] Vaud:
rochers de poudingue au D6zaley sous Chexbres, 450-500 m (BH 91.6.26 et 28) (Amann). TYPE:
[Switzerland] Vaud: rochers au Dejaley 450-500 m, 3.3.1912. Leg. Amann (BH 91.6.28)
(lectotype, here designated, Z!; syntype ["Amann BH 91.6.26"], Z!).
Grimmia moxleyi R.S. Williams, in Holz., Musci Acrocarpi Boreali-Americani et Europaei, fasc. 24 n°
600. 1926. PROTOLOGUE and TYPE: [U.S.A.] On sandstone conglomerate, between Big Rock
Creek and Devil's Punch Bowl, Los Angeles Co., Calif. ColI. George L. Moxley, No. 1141. May
30-31, 1926. Holzinger, Musci Acrocarpi Boreali-Americani et Europaei, n° 600 (lectotype,
designated by Munoz (2000c), NY!; isolectotypes, FH!, H-BR!, MO! 3 replicates).
Synonymized by Munoz (2000c).
DISTRIBUTION: ARM (VA. Manakyan 3783, LE); ATF (FC Naumann s.n., PC); AUS (R.G.
Coveny 15709, MO); AZE (Alexeenko s.n., LE); BEL (1.L. De Sloover 24805, MA); CHE (1. Amann
s.n., Z); CZE (1. Podpii-a s.n., MA); DEU (L. Meinunger 18391, JE); DZA (L. Trabut 2123, PC); EGY
(1.-P. Frahm s.n., NY); ESP (M1. Elias 336, SALA); FRA (HN. Dixon 10, MO); GEO (A.H.
Brotherus & VF Brotherus 131, H-SOL); GRC (V Schiffner 77, FH); HUN (R. Marstaller s.n., JE);
IND (W Koelz 7304A, MO); IRN (1. Bornmiiller 4448, JE); IRQ (HFv. Handel-Mazzetti 1214, FH);
ITA (K. Loitlesberger s.n., MA); JOR (W Frey & H Kiirschner 4606, MA); MAR (P. Davis & J.
Davis D48490A, NY); MKD (T. Herzog 625, JE); NZL (T. WN. Beckett s.n., H-BR); OMN (H
Kiirschner 99-198, MA); PAK (G. Kerstan 5812, JE); POL (H Greven s.n., MA); PRT (C Sergio et al.
31.3b, BCB); SAU (W Frey & H Kiirschner 3136, MA); SUN (V Va9ik s.n., NY); SYR (HK.
Haussknecht 40, JE); TKM (u. Mamatkulov 102, MO); TUN (1.-P. Frahm 844510, MO); TUR (E.
Yucei 8011, MO); USA (FA. MacFadden 3809, MO); UZB (CC Townsend 861193, ALTA); YUG
(M Sabovljevid s.n., MA).
60
MuNoz AND PANDO
50. Grimmia ovalis (Hedw.) Lindb., Acta Soc. Sci. Fenn. 10: 75. 1871. Dicranum ovale Hedw.,
Descr. Micr.-Anal. Musc. Frond. 3: 81, tab. 34 fig. 1-6. 1792, nom. inval. Bryum ovale Hoffm.,
Deutschl. Fl. 2: 35. 1795, nom. inval. Dicranum ovatum Sw., Kong!. Vetensk. Acad. Nya Handl.
16: 243. 1795, nom. inval. Dicranum ovale Hedw., Sp. Musc. Frond.: 140. 1801. Trichostomum
ovatum P. Beauv., Prodr. Aetheogam.: 91. 1805, nom. illeg., incl. typo spec. prior. Dicranum
ovatum Brid., Muscol. Recent. Supp!. 1: 214. 1806, nom. illeg., incl. typo spec. prior. Grimmia
e/liptica Funck, Deutschl. Moose: 16, tab. 11 fig. 1. 1820, nom. illeg., inc!. typo spec. prior.
Trichostomum ovale (Hedw.) Roehl. ex Steud., Nomencl. Bot. 2: 99 [also 421]. 1824, nom. inval.,
pro synon. Campylopus ovalis (Hedw.) Wahlenb., Fl. Suec. 2: 748. 1826. Dryptodon ovatus
(Hedw.) Brid., Bryol. Univ. 1(1): 202. 1826. Grimmia commutata Huebener, Muscol. Germ.: 185.
1833, nom. illeg., incl. typo spec. prior. Dryptodon ellipticus (Funck) Hartm., Handb. Skand. Fl.
ed. 3: 271. 1838, nom. illeg., non Brid. 1826. Guembelia elliptica (Funck) Hampe, Bot. Zeitung
(Berlin) 4: 125. 1846. Guembelia ovalis (Hedw.) Mull. Hal., Syn. Musc. Frond. 1: 774. 1849.
Guembelia commutata (Huebener) Rabenh., Krypt. Fl. Sachsen 1: 444. 1863, nom. illeg., incl. typo
spec. prior. Dryptodon ovalis (Hedw.) Hartm. ex H. Moller, Ark. Bot. 26A(2): 14. 1934, nom.
inval., pro synon. PROTOLOGUE: In rupestribus montosis Saxoniae [Germany], (Thuringiae
prope Isenacum [Germany], in granite montis piniferi Franconiae, Austriae prope Engelhardszell).
TYPE: [no locality given] (lectotype, designated by Geissler & Maier (1995: 502), G!).
Bryum brevicaule Vill., Hist. Pl. Dauphine 3(2): 871. 1789, nom. inval. PROTOLOGUE: [France]
dans les Alpes, aOrciere, Saint-Leger, dans Ie Champfaur, en Valgaudemar & ailleurs.
Grimmia ovata F. Weber & D. Mohr, Naturh. Reise Schweden: 132, tab. 2 fig. 4a-c. 1804.
Campy/opus ovalis var. ovatus (F. Weber & D. Mohr) Wahlenb., Fl. Suec. ed. 2, 2: 776. 1833.
Dryptodon ovatus (F. Weber & D. Mohr) Hartm., Handb. Skand. Fl. ed. 3: 271. 1838, nom. illeg.,
non Brid. 1826. Grimmia ovata var. euovata Loeske, Laubm. Eur. Part I: 113. 1913, nom. inval.
PROTOLOGUE: [Norway] Habitat in rupibus prope Holmiam. Accepimus quoque e Norvagia
[sic]. TYPE: Holmiae (lectotype, designated by Munoz (1999a: 159), BM!).
Grimmia canescens Schleich., Neues J. Bot. 1(2): 198. 1805, nom. nud. Grimmia nigricans Lam. &
DC., Fl. Frany. ed. 3 [original issue] 2: 459. 1805. Dryptodon ovatus var. nigricans (Lam. & DC.)
Brid., Bryol. Univ. 1(1): 179. 1826, nom. inval., pro synon. PROTOLOGUE: Cette espece m'a ete
envoyee des Alpes voisines du Leman [Switzerland]: elle a ete trouvee dans les Pyrenees
[France/Spain], par Ie C. Dufour, et dans les montagnes d' Auvergne [France], par Ie C. Dubois.
TYPE: Pyrenees (lectotype, here designated, G!).
Grimmia ventricosa Schleich., Cat. PI. Helv. ed. 4: 40. 1821, nom. nud. Grimmia commutata var.
ventricosa Lesq., Mem. Soc. Sci. Nat. Neuchatel 3(3): 27. 1846, nom. nud. ORIGINAL
MATERIAL: [Helvetia]. ORIGINAL SPECIMEN: Helvetia. Schleicher [FH specimen labeled as
"Europe"] (FH!, NY!, PC!).
Grimmia commutata f. epilifera J.E. Zetterst., Revis. Grimm. Scand.: 90. 1861. Grimmia commutata
var. imberbis Lindb. ex H. Moller, Ark. Bot. 26A(2): 17. 1934, nom. inval., pro synon. Grimmia
commutata var. mutica J.E. Zetterst. ex H. Moller, Ark. Bot. 26A(2): 17. 1934, nom. inval., pro
synon. PROTOLOGUE: [Sweden] In saxis apricis Sueciae mediae hic illic occurrit, V.C. in
Uplandia, Vestrogothia boreali, Dalia et Vermlandia [... ]. TYPE: In saxis juxta Gottsunda prope
Upsaliam. 5-V-1858. J.E. Zetterstedt (lectotype, designated by Munoz (1999a: 159), H!).
Synonymized by Munoz (1999a: 159).
WORLD SYNOPSIS OF GRIMMIA
61
Grimmia cossonii Besch., Cat. Mouss. Alg.: 18. 1882. PROTOLOGUE: [Algeria] Djebel Kse1, pres de
Geryville, 1,307 met., mai 1856 (Coss.[Cosson, E.S.-C.]). TYPE: Flore de Algerie. Djebe1 Ksel pr.
Geryville. n° 1762 ex herb. Bescherelle. Legit Cosson 30 mai 1856 (lectotype, designated by
Munoz (1999a: 159), PC!).
Synonymized by Munoz (1999a: 159).
Grimmia ovata var. gracilis Roll, Hedwigia 32: 265. 1893, nom. nud. ORIGINAL MATERIAL:
[U.S.A.] Idaho: Coeur d'Alene (1163). ORIGINAL SPECIMEN: Coeur d'Alene. Idaho. 6/888 l
Roll (H-BR!).
Grimmia bernoullii Miill. Hal., Bull. Herb. Boissier 5(3): 200. 1897. PROTOLOGUE: [Guatemala]
Quezaltenango; ColI. N° 115 partim [leg. Bernoulli & Cario]. TYPE: Quezaltenango, leg.
Bernoulli & Cario (lectotype, designated by Munoz (1999a: 159), PC!; isolectotype, GOET!).
Synonymized by Munoz (1999a: 159).
DISTRIBUTION: AFG (K Holmen s.n., ALTA); AND (C Sergio s.n., LISU); ARM (V.
Manakyan 4610a, ERE); AUT (CC Townsend 92/552, MA); BEL (C Delogne s.n., FH); BGR (v.
Valchev & V. Nikolov s.n., MO); CAN (FA. MacFadden 3849, RM); CHE (J.C Schleicher s.n., NY);
CRN (B.C Tan 93-1242, FH); CYP (F Unger s.n., IE); DEU (H Muller s.n., FH); DZA (J.-P. Frahm
844540, MA); ESP (A.C Crundwell 377, NY); FIN (V.F Brotherus s.n., IE); FRA (J.-P. Frahm
935072, MA); GEO (V.F Brotherus 229, H-SOL); GRC (HK Haussknecht s.n., JE); GTM (CG.
Bernoulli & R. Cario s.n., PC); HUN (L. Meinunger 7404, IE); ISL (B. J6hannsson 33267, ICEL);
IND (J.F Duthie 10830, IE); IRN (P. Goldblatt 822, MO); ITA (M. Fleischer s.n., FH); KAZ (B.
Allen 10750, MO); KGZ (VF Brotherus s.n., H-BR); LKA (Anonymous s.n., BM); MAR (L. Trabut
1713, PC); MEX (A.J. Sharp 784, TENN); MKD (T. Herzog 570, IE); NOR (S.C Sommer/elt s.n., H);
PAK (C Jaggi MYQ123, MO); POL (KG. Limpricht s.n., FH); PRT (C Aedo s.n., MA); SIB (M.
Ignatov 0/463, MA); SUN (VG. Onipchenko 48/95, NY); SVK (z. Pilous s.n., FH); SWE (CG. Myrin
s.n., BM); TUR (H.K. Haussknecht 60, IE); TZK (P. Polyakov s.n., NY); UKR (D. Zerov s.n., NY);
USA (B. Allen 17892, MO); UZB (L. Nazarenko s.n., LE).
51. Grimmia percarinata (Dixon & Sakurai) Noguchi ex Deguchi, l Sci. Hiroshima Univ., Ser. B,
Div. 2, Bot. 16: 198. 1979 ["1978"]. Didymodon percarinatus Dixon & Sakurai, in Sakurai, Bot.
Mag. (Tokyo) 50: 516. 1936. PROTOLOGUE: [Japan] Kiushiu: Provo Ohsurni, Insula Yakushima,
mt. Kurorni, ad rupes (Leg. Y. Doi. Typus in Herb. Sakurai Nr. 4329. 5 Aug. 1934). TYPE: Ins.
Yakushima, Kiusiu, Japan. ColI. Y. Doi. Date Aug. 1934. Type (holotype, Deguchi (1978: 198),
MAK not seen; isotype, BM!).
Grimmia lurida Wilson, Hooker's J. Bot. Kew Gard. Misc. 9: 323. 1857, nom. nud. ORIGINAL
MATERIAL and SPECIMENS: Ceylon [Sri Lanka], Gardner n° 329 [... ] Sikkim-Himalaya
temperata; Lachen [India], alt. 10,000 ped., lD.H. [lD. Hooker n° 285] (BM!, FH!).
DISTRIBUTION: IND (J.D. Hooker 285, BM); JPN (F Mizokuti 59, NICH).
52. Grimmia pilifera P. Beauv., Prodr. Aetheogam.: 58. 1805. PROTOLOGUE: [U.S.A.] Etats-Unis
d' Amerique. TYPE: (G? not found).
62
MuNoz AND PANDO
Grimmia pensilvanica Schwagr., Sp. Musc. Frond. Suppl. 1(1): 91, tab. 25. 1811. Grimmia
pennsylvanica Schwagr. ex Sull. & Lesq., Musc. Bor.-Amer.: 32. 1856, nom. inval., orthogr. err.
PROTOLOGUE: [U.S.A.] Lecta in Pensilvaniae rupibus a rev. Muhlenbergio. TYPE: (lectotype,
designated by Cao & Vitt (1986: 144), G not seen).
Synonymized by Cao & Vitt (1986: 144).
Grimmia arizonae Renauld & Cardot, Rev. Bryoi. 19: 85. 1892. PROTOLOGUE: [U.S.A.] Ariz.
[Arizona]. TYPE: Am. sept. Fort Grand, Arizonae. Leg. La Foret (lectotype, designated by Munoz
(1999a: 161), PC!; isolectotypes, FH!, NY!).
Synonymized by Munoz (1999a: 161).
Grimmia tenax Mull. Hal., Nuovo Giorn. Bot. Ital., n.s. 3: 109. 1896. PROTOLOGUE: [China] In
monte medio Si-ku-tziu-san. TYPE: China interior, provincia Schen-si mer. in medio monte SiKu-tzui-san [sic] (lectotype, designated by Deguchi (1978: 201), FI not seen; isolectotype, PC!).
Synonymized by Deguchi (1978: 201).
Grimmia atroviridis Cardot, Bull. Herb. Boissier, ser. 2, 8(5): 333. 1908. PROTOLOGUE: [South
Korea] Coree: Pomasa (n. 209); Hoang-hai-to (n. 658); ile Quelpaert (n. 91). TYPE: Coree:
Pomasa, Faurie 209 (lectotype, designated by Deguchi (1986: 327), PCl; syntypes ["Faurie 91"]
H-BRl, NYl).
Grimmia arsenei Cardot, Rev. Bryol. 40: 37. 1913. PROTOLOGUE: [Mexico] Etat de Michoacan:
environs de Morelia (Frere Arsene, 1911). TYPE: Mexico. Vicinity of Morelia, State of
Michoacan 2000 m. N° 7894 Bro. G. Arsene, Collector 3.XII.l911 (lectotype, designated by
Munoz (1999a: 161), PCl; isolectotypes, Fl, PC! 2 replicates).
Synonymized by Munoz (1999a: 161).
Grimmia hisauchii S. Okamura, 1. ColI. Sci. Imp. Univ. Tokyo 38(4): II, fig. 4. 1916.
PROTOLOGUE: [Japan] Hondo: Provo Sagami, Ikego, Dushi (Leg. Kiyotaka Hisauchil
12.xII.l914.). TYPE: Sagami, Kamakura-cho. Dec. 12st 1914. Kiyotaka Hisauchi 12822 (isotype,
TNSl).
Synonymized by Deguchi (1978: 201).
Grimmia santaritae E.B. Bartram, Bryologist 27: 60, pI. 9. 1924. PROTOLOGUE: [U.S.A.] Dry rocks
faces, White House Canyon, Santa Rita Mts., Santa Cruz County, Arizona, E.B. Bartram, No. 694,
February 20, 1923. TYPE: Mosses of Southern Arizona collected and distributed by Edwin B.
Bartram. N° 694. Dry rock faces. White House Canyon, Santa Rita Mountains, Santa Cruz County.
Altitude 4500 ft. Feb. 20-1923 (lectotype, designated by Munoz (1999a: 161), FHl; isolectotypes,
FH!,NY!).
Synonymized by Munoz (1999a: 161).
Grimmia santaritae f. propagulijera E.B. Bartram, Bryologist 27: 60. 1924. Grimmia arizonae f.
propagulijera (E.B. Bartram) G.N. Jones, in Grout, Moss Fl. N. Amer. 2(1): 33. 1933.
PROTO LOGUE: [U.S.A., locality not explicitly stated]. TYPE: Mosses of Southern Arizona
collected and distributed by Edwin B. Bartram. N° 618. Shaded bank in Patagonia Mountains,
Santa Cruz County. Altitude 5000 ft. Feb. 13-1923 (lectotype, designated by Munoz (1999a: 161),
FHl).
Synonymized by Munoz (1999a: 161).
WORLD SYNOPSIS OF GRiMMIA
63
Grimmia elatior var. squarrifolia Dixon & Ther., Hong Kong Naturalist, Suppl. 2: 10. 1933.
PROTOLOGUE: [China] Exposed granite rock, 400-500 ft. alt., Amoy I., 8th July, 1931; coll.
G.A.c. Herklots (B.8 A, type). Vertical face of rock, ibidem (B.15 B, B.16 B). TYPE: Granite
rock, 400-500 ft. alt., Amoy Island. China. 8 July 1931. CoHo G.A.C. Herklots. No. 8.A (holotype,
according to Cao & Viti (1986: 144), BM not seen; isotype, PC!; paratypes, H!, MO! ["Herklots
B.l5B"]).
Synonymized by Deguchi (1978: 201).
Grimmia eurybasis Dixon & Sakurai, Bot. Mag. (Tokyo) 53: 287. 1939. PROTOLOGUE: [Japan]
Kyusyu: Provo Higo, Isyoti (Leg. K. Mayebara Typus in Herb. K. Sakurai Nr. 950623 Feb. 1936);
ebenso Tunagi-Dorf (Nr. 11189 4 Okt. 1936). TYPE: (holotype, according to Deguchi (1978:
201), MAK not seen).
Synonymized by Deguchi (1978: 201).
Racomitrium doii Sakurai, Bot. Mag. (Tokyo) 54: 6 fig. 3. 1940. PROTOLOGUE: [Japan] Kiusiu:
Provo Ohsumi, Utinoura, Tarumi, auf Felsen (Leg. Y. Doi Typus in Herb. K. Sakurai Nr. 1171630
Maerz 1938). TYPE: (holotype, according to Deguchi (1978: 201), MAK not seen).
Synonymized by Deguchi (1978: 201).
Grimmia kirienensis C.H. Gao, Flora Muscorum Chinae Boreali-Orientalis: 379, fig. 82. 1977.
PROTOLOGUE: [China] Provo Kirin: Districtus Linkiang Mt. Mao-ehr-shan 800 m, ad rupes,
7.IX.1963, Ch. Gao 7714 (Typus) (Typus in Herb. Inst. Sylv. et Pedol. Acad. Sinicae). Provo
Heilungkiang: Xiao-chingan-ling Yichun, ad rupes, 1O.X.1950, Y.L. Chang 1440. TYPE: China.
Jilin. Lijiang Xian. 7.9.1963. Gao Chien 7714 (holotype, IFSBH!).
Synonymized by Deguchi (1986: 327-328).
DISTRIBUTION: CAN (J. Macoun s.n., MA); CHN (c. Gao 1263, ALTA); JPN (A. Sakai 1066,
TNS); KOR (UJ. Faurie 91, H-BR); MEX (A.J. Sharp et al. 3233, TENN); MNG (c. Tsegmed 1895,
LE); SFE (R.E. Magill 12367, MO); SIB (V. Smirnov 5461, MO); SUN (P. Zhudov & Pokrovskaya
s.n., LE); USA (B. Allen 17830, MO); VIR (Krebs s.n., H-SOL).
53. Grimmia pitardii Corb., Bull. Soc. Bot. France 56: LVI. 1909. Campylostelium pitardii (Corb.)
Maier, Candollea 53: 307. 1998. PROTOLOGUE: [Tunisia] [... ] environs d'AYn-Draham
(Khrournirie). [J. Pitard & L. Corbiere]. TYPE: (isotype, G not seen).
NOTE: Maier (1998) has transferred G. pitardii to Campylostelium after a detailed anatomical
study. I agree with her on this taxon being odd in Grimmia, and that it shares some anatomical features
with Campylostelium. Nevertheless, there are other characters shared with species of Grimmia subg.
Grimmia and not with Campylostelium. At this moment I prefer to take a conservative approach and
maintain this species in Grimmia.
Usmania campylopoda Laz., Dopov. Akad. Nauk Ukrajins'k. RSR 11: 1040. 1970. Grimmia
campylopoda (Laz.) K. Saito, J. Jap. Bot. 48(6): 163. 1973. PROTOLOGUE: [Tajikistan] Habitat
in terra nec non saxis terra obtectis ad meridiem Rei Publicae Sovieticae Socialisticae
Tadzhikorum: in decliviis jugi Tuyun Tau, 650 m supra mare, N° 8266, 22.IV 1966. U.
Mamatkulov legit (holotypus); ibidem, N° 8268 22.IV.1966, U. Mamatkulov leg (Isotypus);
64
MuNoz AND PANDO
reservatum publicum "Tigrovaya balka", in arenis Kashka Kum, 350 m supra mare; N° 8298,
23.1V.1966. U. Mamatkulov legit. TYPE: Tajikistan: South Saartus, Tugon-Tau, Karon
Mamatkulov 8266 (holotype, L WS not seen).
Synonymized by Abramova & Abramov (1988).
Grimmia gibbosa S. Agnew, 1. Bryoi. 7: 339. 1973. PROTOLOGUE: [Iraq] Kirkuk Liwa. Tuz
Khurmatli bridge over Lesser Zab river, on stony calcareous soil on dry river bank, c.fr., 21 March
1958, N. Polunin s.n. (Type), (holotype in Herb. S. Agnew, isotypes in BUH, BM); Ain Dibbis, N.
ofKirkuk, crevices oflimestone outcrop on N.-facing slopes oflow hills, c.fr., 26 December 1962,
S. Agnew in BUH 598. TYPE: (not seen).
Synonymized by Abramova & Abramov (1988).
DISTRIBUTION: ESP (R.M. Cros s.n., MA); FRA (J.P. Hebrard A2378, MO); GRC (J. Munoz
s.n., MA); TUN (L. Corbiere s.n., PC); TUR (A. Schiifer-Verwimp 206701T, MA); UZB (U.K.
Mamatkulov 8263, LE).
54. Grimmia plagiopodia Hedw., Sp. Musc. Frond.: 78, tab. 15 fig. 6-13. 1801. Grimmia pZagiopus
Spreng., Bot. Zeitung (Regensburg) 1(5): 74. 1802, nom. invaI., orthogr. err. Schistidium
plagiopodium (Hedw.) Loeske, Laubm. Eur. Part I: 52. 1913. PROTOLOGUE: [Germany] In
Saxoniae rupibus detexit CI. Fliigge. TYPE: Jena. Bei der Rasenmiilh. Febr. 1798 leg. Flugge
(holotype, G not seen; isotypes, GOET!, JE!).
Grimmia obtusa Brid., J. Bot. (Schrader) 1800(1(2»: 276. 1801. PROTOLOGUE: [Germany] prope
TAmam [sic] lectam [Circa Jenam in Rosestein prope Kahla in solo nudo caespitose viuentem et
terra quasi obrutam Vater, Bieber aliique legerunt (Bridel (1806: 94)]. TYPE: (B? not found).
Grimmia apocaula Hoffm. ex DC., in Lam. & DC., Fl.
PROTOLOGUE: [France]. TYPE: (G? not found).
ｆｲ｡ｮｾＮ＠
ed. 3 [original issue] 2: 458. 1805.
Grimmia brandegei Austin, Bull. Torrey Bot. Club 6(7): 45. 1875. PROTOLOGUE: [U.S.A.]
Colorado, Dr. Brandegee, (Rau). TYPE: Colorado. 1.S. Brandegee 1874. Ex herb. E.A. Rau
(lectotype, designated by Munoz (1999a: 163), NY!).
Synonymized by Ochyra & Bednarek-Ochyra (1994: 669).
Grimmia plagiopodia var. pilifera Lesq. & James, Man.: 138. 1884. Grimmia plagiopodia var.
longipila Coville ex Holz., Bryologist 28(3): 32. 1925, nom. inval., orthogr. err. PROTOLOGUE:
[U.S.A.] Rocky Mountais, Colorado (E. Hall, Brandegee). TYPE: Rocky ms. legit Hall (lectotype,
here designated, FH!; isolectotype, FH!).
Synonymized by Blom (1996: 262).
Grimmia argentea R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 412, pI. 30 fig. 11. 1895.
PROTOLOGUE: [New Zealand] Limestone rocks, Castle Hill, Canterbury; November, 1889.
Collected by R.B. TYPE: Hab. limestone rocks near Castle Hill west Coast Road (from
Christchurch). ColI. leg. Robert Brown 11/89 (lectotype, here designated, H-BR!; isolectotypes,
H!,H-BR!).
Synonymized by Ochyra (1993).
65
WORLD SYNOPSIS OF GRIMMIA
Grimmia nivea Dusen, Bot. Not. 1905: 303. 1905. PROTOLOGUE: Patagonia australis in saxis.
TYPE: [Argentina] Patagonia australis, Terr. Sta. Cruz in saxis in valle rivuli Aryo. Pelque. 16
Jan. a. 1905. P. Dusen (holotype, Deguchi (1984: 22), S not seen; isotypes, H-BR!, PC!).
Synonymized by Deguchi (1984: 22).
Grimmia plagiopodia var. antarctica E.B. Bartram, Bryologist 60(2): 141. 1957. PROTOLOGUE:
[Antarctica. Ruppert Coast] Mt. Marujupu: in crevices on granite near rockery, B37.1. TYPE: Mt.
Marujupu: in crevices on granite near rockery. J.E. Perkins B.37-1, Nov. 27, 1940 (holotype, FH!).
Synonymized by Ochyra (1993).
DISTRIBUTION: ARG (P. Dusen 5713, H-BR); ARM (V. Manakyan 4822a, ERE); ATA (J.E.
Perkins B37, FH); CAN (J. Macoun s.n., NY); CHL (P.A. Siple 391-6 FH); FRA (R.B. Pierrot s.n.,
PC); DEU (L. Meinunger 11382, IE); GEO (A.H. Brotherus & V.F. Brotherus 21, H-SOL); HUN (A.
Boros s.n., PC); ISL (B. J6hannsson 25570, ICEL); NZL (A.J. Fife 6262, NY); SUN (A.H. Brotherus
& v.F. Brotherus 7, H-SOL); SVK (z. Pilous 314, PC); UKR (A.S. Lazarenko s.n., MA); USA (C.E.
Darigo 2600, MO).
Q
,
55. Grimmia poecilostoma Cardot & Sebille, nom. conserv. prop., Rev. Bryol. 28: 118, tab. 5. 1901.
Grimmia crinita xG. leucophaea Philib., Rev. Bryol. 17: 18-19. 1890, nom. inval., published as a
formula. Grimmia cardotii Herib., Mem. Acad. Sci. Clermont-Ferrand, ser. 2, 14: 358. 1899, nom.
inval., not accepted by author. Grimmia tergestina var. poecilostoma (Cardot & Sebille) Loeske,
Laubm. Eur. Part I: 84. 1913. Grimmia tergestina var. cardotii (Herib.) Podp., Consp. Musc. Eur.:
277. 1954, nom. inval., incl. typo var. prior. PROTOLOGUE: [France.] Pont-de-Longue, sur les
bordes de l'Ailler (Puy-de-D6me) [Gasilien], et Ie Clapier de Saint-Christophe-en-Oisans (Isere)
[Sebille]. TYPE: Isere: Clapier de Saint-Christophe-en-Oisans, Sebille (lectotype, designated by
Munoz (1999a: 164), PC!; syntype ["pont de Longe pres Clermont. Auvergne. 3 Avril 1888. fre.
Gasilien"], PC!).
NOTE: The name G. poecilostoma has been proposed for conservaton (Munoz, 2000a).
Grimmia crassifolia Lindb. ex Broth., Acta Soc. Sci. Fenn. 19(12): 84. 1892. PROTOLOGUE:
[Russia. Ossetia A.R.] Cauc. major occ.: Ossetia, ad rupes calcareas, siccisimas ad fl. Ardon inter
Alagir et Misurtzy. TYPE: Ossetia, in valle fl. Ardon inter Alagir et Misurtsy, ad rupes
aridissirnas, 7/1877. Leg. V.F. Brotherus (lectotype, designated by Munoz (1999a: 164), H-BR!;
isolectotypes, H!, IE!, PC!, UPS!).
Synonymized by Munoz (1999a: 164).
Grimmia gymnostoma Culm., Rev. Bryol. 23: 108. 1896. Grimmia tergestina var. gymnostoma (Culm.)
G. Roth, Eur. Laubm. 1: 410. 1903. Grimmia tergestina f. gymnostoma (Culm.) Loeske, Laubm.
Eur. Part I: 82. 1913, comb. inval., pro synon. PROTOLOGUE: [Switzerland] Decouvert dans les
fissures des rochers calcaires exposes a l'est sur l'Ebenalp, canton d'Appenzell, Ie 24 aout 1896,
retrouve un peu plus tard au Hohen Kasten. TYPE: Au Kalkfelsen, Ostabhang der Ebenalp 1640
m., Kt. Appenzell. 24/8 1896. CuImann (lectotype, designated by Munoz (1999a: 164), PC!).
Synonymized by Munoz (1999a: 164).
66
MuNoz AND PANDO
Grimmia anodon var. mutica Broth. ex Paris, Index BryoI. ed. 2,2: 263. 1904, nom. nud. ORIGINAL
MATERIAL: [Kazakstan] As.- Alatau, Turkestan. ORIGINAL SPECIMEN: Alatau transiliensis:
in valle fl. Almatinka minoris, ad rupes siccissimas reg. silv. infer., JUll 29, 1896. Leg. V.F.
Brotherus. [Musci turkestanici n° 45] (H-BR!).
DISTRIBUTION: ARM (VA. Manakyan 5298, LE); AZE (Alexeenko s.n., LE); CHE (P.
Culmann s.n., PC); CHN (K.-Y. Lang 1262, ALTA); ESP (R.M. Cros s.n., BCB); FRA (G. Gasilien
s.n., PC); GEO (N. V. Chikovani s.n., LE); KAZ (B. Allen 10508, MO); KGZ (VF. Brotherus s.n., FH);
MNG (w. Schubert M18, MO); SIB (M. Ignatov 01462, MO); SUN (VF. Brotherus s.n., H-BR); TKM
(Navessky 3108D, LE); TUR (K. Walther 2832, NY); UKR (1. Partyka s.n., MA); USA (1.R. Stark
NV-25B, MO).
56. Grimmia pseudoanodon Deguchi, in Inoue, Studies on Cryptogams in Southern Peru: 29, pI. 11.
1987. PROTOLOGUE: Peru. Chucuito, near Acora, Provo Puno, Dept. Puno; alt. ca. 3900m, in
rocky meadows, September 29, 1984, leg. H. Deguchi no. 30008 (holotype in TNS, isotypes in
KOCH and NY). TYPE: Peru. Dept. Puno. Provo Puno: Chucuito, near Acora; alt. ca. 3900 m, in
rocky meadow. September 29, 1984. ColI. H. Deguchi 30008 (isotype, NY!).
DISTRIBUTION: ARG (P.G. Lorentz s.n., JE); BOL (M. Lewis 79-2004, F); PER (H. Deguchi
30008, NY).
57. Grimmia pUlla Cardot, Rev. Bryol. 36: 106. 1909. PROTOLOGUE: [Mexico] Etat de Hidalgo:
Honey-station, sur troncs d'arbres, associe it Zygodon circinatus Sch., 1907 (n. 10423). TYPE:
State of Hidalgo. On trees, Honey Station, 12, Dec., 1907 [Pringle, Plantae mexicanae, n° 10423]
(lectotype, designated by Munoz (1999a: 167), PC!; isolectotypes, CANM-197543!, FH!, JE!,
NY!, PC! 2 replicates, TENN!, UC-133015!).
Grimmia pulla var. squarrosa Ther., Rev. Bryol. LichenoI. 5: 99. 1933. PROTOLOGUE: [Mexico]
San Rafael (St.-P. n° 1857); Contreras, rochers (St.-P n° 1472). TYPE: 1857. Mexique: San Rafael,
2600 m (Etat de Mexico). Leg. Fr. Amable. 29.xII. 1931 (lectotype, designated by Munoz (1999a:
167-168), PC!; syntypes ["Contreras, St. Pierre 1472"], H-BR!, PC!).
Synonymized by Crum (1994: 405).
DISTRIBUTION: MEX (A.J. Sharp 178, TENN).
58. Grimmia pulvinata (Hedw.) Sm., EngI. Bot. 24: 1728. 1807. "Bryurn trichodes hirsutie canescens,
capitulis subrotundis reflexis in perbrevibus pediculis" Dill., Cat. PI. Circa Gissam: 226. 1718,
nom. inval. "Bryum orbiculare pulvinatum, hirsutie canescens, capsulis immersis" Dill., Hist.
Musc.: 395, tab. 50 fig. 65 A-C. 1741, nom. inval. Bryumpulvinatum L., Sp. PI. 2: 1121. 1753,
nom. inval. Leersia pulvinata Hedw., Fund. Hist. Nat. Musc. Frond. 1: tab. 10 fig. 65; 2: 88. 1782,
nom. invaI. Fissidens pulvinatus Timm, Fl. Megapol. Prodr.: 216. 1788, nom. inval. Aftelia
pulvinata Ehrh., Plantae Criptogamae Linneae, Decas 17, n° 163. 1791, nom. inval. Encalypta
pulvinata Sibth., Fl. Oxon.: 278. 1794, nom. inval. Dicranum pulvinatum Sw., Kongl. Vetensk.
Acad. Nya Handl. 16: 240. 1795, nom. inval. Fissidens pulvinatus Hedw., Sp. Musc. Frond.: 158,
tab. 40 fig. 1-3. 1801. Fissidens pulvinatus var. communis Hedw., Sp. Musc. Frond.: 158, nom.
WORLD SYNOPSIS OF GRIMMIA
67
illeg., inel. typo spec. Dicranum pulvinatum (Hedw.) Lag., D. Garcia & Clemente, Anales Ci. Nat.
5(14): 176. 1802. Trichostomum pulvinatum (Hedw.) F. Weber & D. Mohr, Bot. Taschenbuch:
109. 1807. Campylopus pulvinatus (Hedw.) Brid., Muscol. Recent. Suppl. 4: 75. 1818 ["1819"].
Dryptodon pulvinatus (Hedw.) Brid., Bryol. Univ. 1(1): 196. 1826. Grimmia pulvinata var.
vulgariS Loeske, Laubm. Eur. Part I: 161. 1913, nom. illeg., incl. typo spec. PROTOLOGUE: In
tectis et muris vulgaris. TYPE: [no locality given] (lectotype, designated by Cao & Vitt (1986:
191), G!).
Fissidens pulvinatus vaT. africanus Hedw., Sp. Musc. Frond.: 159, tab. 40 figs. 4-6. 1801. Dicranum
africanum (Hedw.) F. Weber & D. Mohr, Index Mus. PI. Crypt.: 2. 1803. Dicranum cribrosum
Brid., Muscol. Recent. Suppl. 1: 212. 1806 (NOTE: Erroneously considered legitimate by Munoz
(1999a: 168), this name is predated by D. africanum, see ICBN Art. 52.1). Dicranum pulvinatum
vaT. affine Schleich. ex Brid., Muscol. Recent. Suppl. 4: 76. 1818 ["1819"], nom. inval., pro
synon. Dicranum pulvinatum vaT. africanum (Hedw.) Schleich., Cat. PI. Helv. ed. 4: 40. 1821.
Dryptodon cribrosus (Brid.) Brid., Bryol. Univ. 1(1): 198. 1826. Dryptodon obtusus Brid., Bryol.
Univ. 1(1): 198. 1826, nom. illeg. Grimmia africana (Hedw.) Am., Mem. Soc. Linn. Paris 5: 243.
1827. Grimmia pulvinata vaT. obtusa Huebener, Muscol. Germ.: 194. 1833. "Dryptodon obtusus
Hartm., Handb. Skand. Fl. ed. 3: 270. 1838" (NOTE: Error in Wijk et al. (1962: 176). Hartman
neither combined nor described it as new). Grimmia pulvinata vaT. africana (Hedw.) Wilson, Bot.
Antarct. Voy., Fl. Nov.-Zel. 2: 75. 1854. Grimmia montana vaT. obtusa Bruch & Schimp. ex
Podp., Vestn. Klubu Pfir. v Prostejove 7: 19. 1905 (NOTE: Error in Wijk et al. (1962: 393) for
Grimmia pulvinata vaT. obtusa (Brid.) Huebener). Grimmia obtusa (Brid.) Loeske, Laubm. EuT.
Part I: 165. 1913, nom. inval., pro synon. Grimmia pulvinata f. obtusa Vilh., Vestn. Knil. Ceske
Spolecn. Nauk, Tf. Mat.-PtiT. 1924: 27. 1924, nom. illeg. PROTOLOGUE: [South Africa] Ad
Cap. B. Spei lecta spec imina mihi communicavit O. Swartz. TYPE: Ad Cap. bonae Spei lecta
specimina misit o. Swartz (lectotype, designated by Munoz (1999a: 168), G!).
Synonymized by Munoz (1999a: 168).
Grimmia ecklonii Spreng., Syst. Veg. 4(2): 321. 1827. PROTOLOGUE: Ad saxa in monte leonis,
c.B.S. Ecklon. TYPE: [South Africa] Swellendam. C.B.S. Ecklon (lectotype, here designated,
BM!; isolectotype, BM!).
Dryptodon pulvinatus var. crinitus Breb., Mousses de Normandie, Fasc. 5, n° 119. 1831, nom. nud.
ORIGINAL MATERIAL and SPECIMEN: [France] Falaise. [L.A. de Brebisson] (FH!).
Grimmia cygnicolla Taylor, London J. Bot. 5: 45. 1846. PROTOLOGUE: [Australia] Swan River, MT.
James Drummond. TYPE: Swan River, N.H. James Drummond (lectotype, designated by Munoz
(1999a: 168), FH!; isolectotypes, FH!, H-SOL!, NY!).
SynonymiZed by Munoz (1999a: 168).
Grimmia pygmaea Mull. Hal., Syn. Musc. Frond. 1: 787. 1849. Grimmia trichophylla vaT. australis
Hampe ex MUll. Hal., Syn. Musc. Frond. 1: 788. 1849, nom. inval., pro synon. PROTOLOGUE:
[Australia] Nova Hollandia, ad rupes graniticas: Preiss. TYPE: Grimmia trichophy.-australis
Hampe. N. Holl. Juni 1090 (lectotype, here designated, H-SOL!; isolectotypes, H-BR!, PC!).
68
MuNoz AND PANDO
Grimmia callosa Mull. Hal. & Hampe, Linnaea 26: 498. 1855. PROTOLOGUE: [Australia] Australia
felix, Barossa range.- Brown's hill [leg. F. Muller]. TYPE: Barossa-range. Australia felicis: Ferd.
Muller (lectotype, designated by Munoz (1999a: 168), JE!; isolectotype, MEL-I002648!).
Synonymized by Munoz (1999a: 168).
Guembelia obtusata Mull. Hal. & Hampe, Linnaea 26: 497. 1855. Grimmia obtusata (MUll. Hal. &
Hampe) A. Jaeger, Ber. Tiitigk. St. Gallischen Naturwiss. Ges. 1872-73: 70 (Gen. Sp. Musc. 1:
352). 1874. PROTOLOGUE: [Australia. Victoria] Australia felix [leg. F. Muller]. TYPE:
Australia felix. [Leg. F. Muller] n° 7b (lectotype, here designated, MEL-I002652!).
NOTE: A duplicate of Maller 7b in NY! is G. reflexidens MUll. Hal.
Grimmia basaltica Mitt. ex Hook.f., Handb. N. Zeal. Fl.: 425. 1867. Grimmia pulvinata var. basaltica
(Hook.f.) Dixon, New Zealand Inst. Bull. 3(4): 158. 1926, nom. illeg., non Vilh. 1924.
PROTOLOGUE: [New Zealand] Middle Island: basalt rocks near Dunedin, Lindsay (Herb.
Mitten). TYPE: Hills to south of the Forbury Heads near Dunedin, Ottago, New Zealand.
November 19, 1861. Ex herb. Dr. Lauder Lindsay (lectotype, here designated, NY!; isolectotype,
NY!).
Grimmia gibertii Mitt., J. Linn. Soc., Bot. 12: 100. 1869. PROTOLOGUE: [Uruguay] Brasilia, Monte
Video, Gibert, n. 729, in Herb. Hooker. TYPE: n. 729. Montevideo, in terra. Augusto 1868. Gibert
(lectotype, designated by Munoz (1999a: 168), NY!; isolectotypes, FH!, H-BR!, NY!, PC!).
Synonymized by Munoz (1999a: 168).
Grimmia auresia Besch., Cat. Mouss. Alg.: 17. 1882. Grimmia auresiaca Besch. ex Kindb., Enum.
Bryin. Exot., Suppl. 1: 91. 1889, nom. inval., orthogr. err. PROTOLOGUE: [Algeria] Aures
(Bal.[Balansa, B.]). TYPE: Aures. Legit Balansa ex herb. Bescherelle (lectotype, designated by
Munoz (1999a: 168), PC!; isolectotype, F!).
Synonymized by Munoz (1999a: 168).
Grimmia hendersonii Renauld & Cardot, Rev. Bryo1. 19: 86. 1892 ["Hendersoni"]. Grimmia decipiens
var. hendersonii (Renauld & Cardot) Sayre, Bryologist 67(2): 163. 1964. PROTOLOGUE:
[U.S.A. Oregon] Or. Leg. L.F. Henderson. TYPE: Rill Bed, Hood River [Co.], Oregon. March 3189. L.F. Henderson n° 1933 (lectotype, here designated, PC!; isolectotypes, FH!, NY! 2
replicates).
Grimmia novae-zeelandiae R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 414, pI. 31 fig. 16. 1895.
PROTOLOGUE: [New Zealand] Damp rocks, Port Lyttelton hills; Mount Torlesse; Stewart
Island; Dunedin; West Coast. [... ] R.B. TYPE: Lyttleton Hills. R. Brown (lectotype, here
designated, BM!).
Grimmia obovata R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 416, pI. 32 fig. 23. 1895.
PROTOLOGUE: [New Zealand] Rocks, Lyttelton hills; 1880. Collected by R.B. TYPE:
(lectotype, here designated, fig. 23 in pI. 32, Brown, R., op. cit.). After an unsuccessful search for
specimens referable to this taxon (BM, CHR, H), the illustration in the original publication is
proposed as lectotype of the name. It unambiguously represents Grimmia pulvinata (Hedw.) Sm.
WORLD SYNOPSIS OF GRIMMIA
69
Grimmia pulvinata var. asphaltita Renauld & Cardot, Bull. Soc. Roy. Bot. Belgique 33: 118. 1895.
PROTOLOGUE: [Israel] Hab. Judee: in deserto lacus Asphaltitae inter Hebron et Zuweirat-elFaga (Dr. G.E. Post, 1892. Herb. Boissier). TYPE: Plantae desertis Maris-Mortis. Entre Hebron et
Zuweirat-el-Faga. Le 22 aout 1892. Dr. E.G. Post (lectotype, here designated, PC!).
Grimmia pusilla R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 417, pI. 33 fig. 26. 1895, nom. illeg.,
non F. Weber & D. Mohr 1803. PROTOLOGUE: [New Zealand] On rocks, Mount Self, near the
Hurunui River March, 1893. Collected by R.B. TYPE: (lectotype, here designated, fig. 26 in pI.
33, Brown, R., op. cit.). After an unsuccessful search for specimens referable to this taxon (BM,
CHR, H), the illustration in the original publication is proposed as lectotype of the name. It
unambiguously represents Grimmia pulvinata (Hedw.) Sm.
Grimmia campbelliae Mull. Hal., Hedwigia 37: 162. 1898. PROTOLOGUE: Australia orientalis,
Hume River: Miss Campbell 1881 in Rh. Melbourne. TYPE: Hume River. Miss Campbell. Baron
Ferd. Mueller, Ph. & M.D. [ex MEL] (lectotype, here designated, JE!; isolectotypes, H-BR!, JE!).
Synonymized by Munoz (1999a: 168).
Grimmia tasmanica Mull. Hal., Hedwigia 37: 159. 1898. PROTOLOGUE: [Australia] Tasmania, sine
loco natali: C. Stuart in Rh. Sonder 1840; in muris (wall of the catholic cemetery, Brisbane street):
Weymouth 7.0ct.1890 in Rh. Dr. O. Burchard, qui 1891 pro determinatione misit. TYPE: ad
muros: Weymouth 1890 (lectotype, here designated, PC-Besch!; isolectotypes, S!, W!).
Grimmia woollsiana MUll. Hal., Hedwigia 37: 162. 1898. PROTOLOGUE: Australia, New South
Wales, Sydney: Rever. Dr. Woolls 1881 in Rh. Melbourne. TYPE: New South Wales; Rever. Dr.
Woolls. Rh. C. Muller (lectotype, designated by Munoz (1999a: 168), H-BR!; isolectotype, JE!).
Synonymized by Munoz (1999a: 168).
Grimmia leptotricha MUll. Hal., Hedwigia 38: 120. 1899. PROTOLOGUE: [South Africa] Prom.
bonae spei, Somerset East, monte Boschberg, in rupibus: Prof. Mac Owan in Rh. Georg Winter
1883. TYPE: In monte Boschberg prope Somerset-East, Promont. bon. spei. leg. Mac Owan.
[Rabenhorst-Winter, Bryotheca europaea et extraeuropaea, n° l383] (lectotype, here designated,
BM!).
Grimmia pulvinata var. brevieri Herib., Mem. Acad. Sci. Arts Clermont-Ferrand, ser. 2, 14: 346. 1899.
PROTOLOGUE: [France] Puy-de-Dome.- Sur un mur it Ambert, tres rare (Breviere, 26 fevrier
1895; fertile). TYPE: Puy-de-Dome.- Vieux murs it Ambert, 26 fevrier 1895. Leg. Breviere
(lectotype, here designated, PC-Bonaparte!).
Grimmia Juscipila MUll. Hal. ex Paris, Index Bryol. Suppl.: 173. 1900, nom. nud. ORIGINAL
MATERIAL: [India] As.: Himalaya sept. occid. (Dehra Doun). ORIGINAL SPECIMEN:
Kumaun, Hawalbagh, 4-5000p., 26/4/1843. Leg. D. Jameson (H-BR! 2 replicates).
Grimmia subcurvula Kindb., Ofvers. Forh. Kongl. Svenska Vetensk.-Akad. 57(10): 1007. 1900.
PROTOLOGUE: [Sweden] occurrit in Dalia prope Rostock in rupibus argillaceo-schistosis.
TYPE: Dalsland, Rostock. 14/8 1898. N.C. Kindberg (lectotype, designated by Munoz (1999a:
169), S!; isolectotype, PC!).
Synonymized by Munoz (1999a: 169).
70
MuNoz AND PANDO
Grimmia hutchinsonii R. Br. bis, Trans. & Proc. New Zealand Inst. 35: 337, pI. 39, fig. 30. 1903.
PROTOLOGUE: [New Zealand] On calcareous rocks near Weston, close to Oamaru. Collected by
R. Brown, November, 1897. TYPE: Omaru 11/97 (lectotype, here designated, BM!).
Grimmia kaikouraensis R. Br. bis, Trans. & Proc. New Zealand Inst. 35: 337, pI. 40 fig. 31. 1903.
PROTOLOGUE: [New Zealand] On damp rocks near Kaikoura. Collected by R. Brown. TYPE:
Near Kaikoura, South I., New Zealand. On rocks. ColI. R. Brown ter. (lectotype, here designated,
BM!).
NOTE: The material at BM is a mixture of G. pu/vinata and G. trichophylla. Brown (1903: 337,
pI. 40 fig. 31) used material of both species to describe and illustrate his new taxon. Some stems kept
in a triangular packet have sporophytes and are gonioautoicous. They are selected as lectotype of the
name. Dixon (1926: 157) considered G. kaikouraensis synonymous with G. trichophylla, although the
specimen in his herbarium is annotated as identical to G. pu/vinata. Recently, Greven (1998: 397)
blindly followed Dixon's publication, although he annotated the BM specimen without realizing it was
part of the type.
Grimmia pulvinata var. sardoa Herzog, Ber. Ziircherischen Bot. Ges. 15: 47. 1905. Grimmia pu/vinata
f. sardoa (Herzog) Laz., Zurn. Inst. Bot. Vseukrajins'k. Akad. Nauk 26-27: 206. 1938.
PROTOLOGUE: [Italy. Sardinia] Auf FelsblOcken (Jurakalk) am Waldrand unter den Tonneri di
Seui, am Weg zur Perdaliana, ca. 1100 m. 15. April 1904. TYPE: Sardinien. Auf Felsblocken
(Jurakalk) am Waldrand unter den Tonneri di Seui, am Weg zur Perdaliana, ca. 1100 m. 15. April,
04 (holotype, JE!).
Grimmia arachnoidea Dusen, Rep. Princeton Univ. Exped. Patagonia 8: 122. 1903, nom. nud.
Grimmia orbicularis var. patagonica Cardot, Wiss. Erg. Schwed. Siidpolar-Exp. 4(8): 284. 1908.
PROTOLOGUE: [Chile] haute vallee du Rio Aysen ([Dusen] n° 587). TYPE: Patag. occ. in valle
fluminis Aysen in saxis. Feb. 18 a. 1897. P. Dusen (lectotype, designated by Munoz (1999a: 169),
H-BR!; isolectotype, PC!).
Synonymized by Munoz (1999a: 169).
Grimmia phyllorhizans Broth., Nat. Hist. Juan Fernandez 2(3): 419, tab. 26 fig. 21-24. 1924.
PROTOLOGUE: [Chile] Masatierra: Villagra; ad saxa (129). TYPE: Juan Fernandez: Masatierra,
steurosen v. om Villagra, st. 8. 7/1/1917. Carl o. Inga Skottsberg nr. 129 (lectotype, designated by
Munoz (1999a: 169-170), H-BR!; isolectotypes, FH!, NY!, UPS!).
Synonymized by Munoz (1999a: 169).
Grimmia decurrentipilis Ther., in Felipp., Rev. BryoL LichenoL 2: 216, fig. 8. 1929. PROTOLOGUE:
[Uruguay] La Paz, rochers; n° 837. TYPE: Herbario del Dr. F. Felippone. Plantas de la Republica
del Uruguay. N° 837. Localidad: La Paz. Departamento: Canchones. Ad saxa. Marzo /913
(lectotype, designated by Munoz (1999a: 170), PC!).
Synonymized by Munoz (1999a: 170).
Grimmia montevidensis Ther., in Felipp., Rev. BryoL LichenoL 2: 216, fig. 7. 1929. PROTOLOGUE:
[Uruguay] Montevideo, n° 839. TYPE: Herbario del Dr. F. Felippone. Plantas de la Republica del
Uruguay. N° 839. Localidad: Montevideo. Departamento: Montevideo. Ad saxa. Marzo /913
(lectotype, designated by Munoz (1999a: 170), BM!; isolectotype, PC!).
Synonymized by Munoz (1999a: 170).
WORLD SYNOPSIS OF GRIMMIA
71
Grimmia pulvericola Mull. Hal. ex Bruehl, Rec. Bot. Surv. India 13(1): 123. 1931, nom. nud.
ORIGINAL SPECIMEN: [India] NW Himalaya, Jaunsan 3000', Mai 1891. Leg. C.G. Rogers.
Com. Levier (H-BR!).
Grimmia trichophylla var. indianensis Sayre, Bryologist 57: 21, fig. a-f. 1954. Grimmia indianensis
(Sayre) Crum, Bryologist 68: 234. 1965. PROTOLOGUE: [U.S.A.] On sandstone along a small
creek leading to Pine Creek on the east side of the road 2 miles southwest of Rainsville, Warren
County, Indiana. Winona H. Welch 9694,27 November 1937. Specimens in herbaria of DePauw
University, New York Botanical Garden, Welch, Sayre. TYPE: On sandstone on small creek
leading to Pine Creek on east side of road 2 m. southwest of Rainsville, Indiana, Warren County.
27 November 1937. CoIl. W.H. Welch n° 9694. Det. G. Sayre (lectotype, here designated, NY!;
isolectotypes, CANM-197537!, FH!, NY! 2 replicates).
Grimmia ahmadiana Nog., J. Hattori Bot. Lab. 16: 79. 1956. PROTOLOGUE: Pakistan: Swat State,
Khaza Khela (S. Ahmad, ns. 102-typus, 108, Apr. 1954). TYPE: Pakistan: Khaza Khela, on the
ground. Apr. 17, 1954, coIl. S. Ahmad 102 (holotype, NICH-361549!).
DISTRIBUTION: ARG (CC Hosseus 2220, FH); ARM (V Manakyan 1841a, ERE); AUS (A.J.
Taylor 6, MEL); AUT (Heimhold s.n., GOET); BEL (C Delogne s.n., FH); BRA (D.M Vital 9155,
FLAS); BGR (S. Petrov s.n., MO); CAN (J. Macoun s.n., FH); CHE (0. Coulter s.n., TCD); CHL (H
Deguchi 25159, HIRO); CYP (F. Unger s.n., JE); CZE (M Deyl s.n., MA); DEU (MR. Crosby 17700,
MO); DNK (A. Hesselbo s.n., FH); DZA (J.-P. Frahm 844526, MA); ESP (P. Sera B-356, BCB); FIN
(J.G. Bomansson s.n., JE); FRA (J.L. De Sloover 45145, MA); GBR (R.A. Pursell SVN-3, MO); GEO
(P.H Raven 27080, MO); GRC (V Schiffner 157, FH); HUN (L. Vajda s.n., MA); IND (W Koelz
7544, MO); IRL (D.L. Kelly s.n., TCD); IRQ (HF. v. Handel-Mazzetti 1533, FH); ISR (P.H Raven
22123, MO); ITA (MR. Crosby 17699, MO); LSO (R.E. Magill 5863, F); LBY (Gimingham 146, PC);
MAR (J. Munoz s.n., MA); MKD (T. Herzog 568, JE); MEX (E. Palmer 113, MO); NOR (N. Bryhn
s.n., JE); NZL (J. Child M2768, F); POL (KG. Limpricht s.n., FH); PRT (C Sergio et al. 31.3a, BCB);
SAU (W Frey & H Kurschner 3133, MA); SIB (A.L. Abramova & I.I. Abramov s.n., MO); SUN
(Alexeenko 117-65, LE); SVK (J. Jedliika s.n., MO); SWE (SJ. Lindgren s.n., FH); SYR (HK
Haussknecht 45, JE); TKM (u. Mamatkulov 2, MO); TUR (HK Haussknecht 67, JE); TZK (P.
Polyakov s.n., NY); UKR (A.S. Lazarenko s.n., NY); URY (E.J. Gibert 729, FH); USA (A.T.
Whittemore 3382, MO); UZB (L. Nazarenko 33, LE); YUG (M. Sabovljevic s.n., MA); ZAF (L. Rolfes
20, JE).
59. Grimmia ramondii (Lam. & DC.) Margad., Lindbergia 1: 128. 1972. Pterigynandrum ramondii
Lam. & DC., Fl. Frany. ed. 3 [original issue] 2: 462. 1805 ["ramondi"], Leucodon ramondii (Lam.
& DC.) Brid., Muscol. Recent. Suppl. 4: 135. 1818 ["1819"], PROTOLOGUE: [France/Spain]
dans les Pyrenees, par Ie C. Ramond. TYPE: Pyrenees. Herb. Cando (lectotype, here designated,
B!).
Bryum patens Dicks., Fasc. PI. Crypt. Brit., Fasc. 2: 6, tab. 4 fig. 8. 1790, nom. inval. Bryum patens
Hedw., Sp. Musc. Frond.: 86. 1801. Pterigynandrum patens (Hedw.) Brid., 1. Bot. (Schrader)
1800(1(2»: 278. 1801. Dicranum patens (Hedw.) Sm., Fl. Brit. 3: 1213. 1804. Trichostomum
nudum Schleich., Neues 1. Bot. 1(2): 198. 1805, nom. nud. Trichostomum patens (Hedw.) D.
Mohr, Ann. Bot. (Konig & Sims) 2: 545. 1806. Fissidens patens (Hedw.) Wahlenb., Fl. Lapp.:
334. 1812. Campylopus patens (Hedw.) Brid., Muscol. Recent. Suppl. 4: 73. 1818 ["1819"]'
72
MuNoz AND PANDO
Dryptodon patens (Hedw.) Brid., Bryol. Univ. 1(1): 192. 1826. Trichostomum patens var. majus
Hook. & Taylor, Muscol. Brit. ed. 2: 104. 1827, nom. illeg., incl. typo spec. Racomitrium patens
(Hedw.) Huebener, Muscol. Germ.: 198. 1833. Grimmia arcuata De Not., Mem. Reale Accad.
Sci. Torino 39: 250. 1836, nom. illeg., incl. typo spec. prior. Grimmia patens (Hedw.) Bruch &
Schimp., Bryol. Europ. (fasc. 25-28) 3: 120, tab. 246. 1845, nom. illeg., non Homsch. 1819.
PROTOLOGUE: [United Kingdom] Ad rupes in rivulis alpinis Scotiae. Ben Nevis. Dicks. fasc. II.
p. 6. t. IV. f. 8. TYPE: (BM? not seen).
Orthotrichum curvatum Brid., Bryol. Univ. 1: 791. 1827. Grimmia curvata (Brid.) De Sloover, Bull.
Jard. Bot. Etat 37: 445. 1967. PROTOLOGUE: [Canada] In insula Terre Neuve habitat. TYPE:
Terre Neuve. La Pylaie. [Parm, 1825, illegible] (holotype, B!).
Grimmia serrata Kindb., Rev. Bryol. 34: 90. 1907. PROTOLOGUE: Canada, Brit. Col., Rossland:
J.M. Macoun 1902. TYPE: Canada, B.C., Old Glory rnnt., Rossland. 15/8 02. J.M. Macoun
(holotype, S!).
Synonymized by Allen (1995: 164).
Dryptodon patens var. propagulijer J.J. Amann, Fl. Mouss. Suisse 2: 140. 1918 ["propagulifera"].
PROTOLOGUE: [Switzerland] Valais: Emosson, 1750-1800 m., sur Ie gneiss (Amann) (BH
93.2.10) Graubiinden: Albertitobel bei Davos-Platz, auf erratischen Gneiss, 1650-1700 m
(Amann) (BH 93.2.8). TYPE: Graub. Albertitobel fur Davos. 24.5.90. Leg. Amann (BH 93.2.8)
(lectotype, here designated, Z!; syntype ["Valais. Emosson March 1896. Leg. Amann (BH
93.2.1 0)"], Z!).
DISTRIBUTION: AUT (K. Osterwald & V. SchifJner s.n., JE); BEL (A. Lachmann s.n., PC); BGR
(L. Meinunger 14485, JE); CAN (J.M Macoun s.n., S); CHE (J. Amann s.n., Z); DNK (C Jensen s.n.,
NY); ESP (P. Allorge s.n., FH); FIN (R. W. Hartman s.n., H); FRA (F. G. KneifJ & CP. W. Maercker
s.n., FH); GBR (A.C Crundell s.n., RM); GRL (s. Berggren s.n., NY); IRL (H. W. Lett s.n., TCD); ISL
(B. J6hannsson s.n., ICEL); ITA (A. Bottini s.n., FH); JPN (T Nagai 1531, H-BR); NOR (J. Heinrichs
202, GOET); POL (K.G. Limpricht s.n., FH); SUN (Anonymous s.n., H-BR); SVK (z. Pilous s.n., JE);
SWE (J.E. Zetterstedt s.n., H); USA (F.J. Hermann 24747, RM); YUG (M Sabovljevic s.n., MA).
60. Grimmia redunca Wilson ex Mitt., J. Linn. Soc., Bot. Suppl. 1: 43. 1859. Grimmia redunca
Wilson, Hooker's J. Bot. Kew Gard. Misc. 9: 323. 1857, nom. nud. PROTOLOGUE: [India] In
Himalayae orient. reg. alp., Sikkim, J.D. Hooker (No. 294). TYPE: 294. Jongri alt. l3.000. Dr.
Hooker (holotype, NY!; isotypes, FH!, H-SOL!, LE!, PC!, PC-Besch!).
DISTRIBUTION: IND (J.D. Hooker 294, NY).
61. Grimmia reflexidens Miill. Hal., Syn. Musc. Frond. 1: 795.1849. PROTOLOGUE: Chile australis
inter Gr. consobrinam: poppig. TYPE: Inter Dryptodon consobrinus Kze. Chile. Leg. Poppig
(lectotype, designated by Munoz (1998b: 394), BM!; isolectotypes, JE!, NY!, PC!).
Grimmia ramosa Schleich., Cat. PI. Helv. ed. 3: 33. 1815, nom. nud.
WORLD SYNOPSIS OF GRIMMIA
73
Guembelia lamellosa Mull. Hal., Bot. Zeitung (Berlin) 12: 318. 1854. Grimmia lamellosa (Mull. Hal.)
A. Jaeger, Ber. Tatigk. St. Gallischen Naturwiss. Ges. 1872173: 72 (Gen. Sp. Musc. 1: 354). 1874.
PROTOLOGUE: [France] In Pyrenaeis centralibus, in rupibus ad lacum Espingo, altitudine 5600
pedum, mense August. 1851 leg. Joh. Lange. TYPE: in Pyrenaeis centralibus, in rupibus ad lacum
Espingo (lectotype, designated by Deguchi (1978: 207), PC!; isolectotypes, H-SOL!, JE!).
Synonymized by Munoz (1998b: 394).
Grimmia sessitana De Not., Atti Reale Univ. Genova 1: 704. 1869. Grimmia exannulata Lindb. ex
Broth., Acta Soc. Sci. Fenn. 19(12): 87. 1892, nom. inval., pro synon. Grimmia alpestris var.
sessitana (De Not.) I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1909(5): 22.
1909. Grimmia alpestris f. sessitana (De Not.) Loeske, Laubm. Eur. Part I: 105. 1913. Grimmia
alpestris var. stomata Loeske, Laubm. Eur. Part I: 104, figs. 26e,f,g, 27d,e,i,k. 1913, nom. illeg.
PROTOLOGUE: [Italy] In praeruptis infra Hospitium della Valdobbia, in valle Sessitana, legit
Carestia. TYPE: Frane aIle scaturigini del Vogna, sotto l'ospizio della Valdobbia in Val Sessia.
Carestia (lectotype, designated by Cao & Vitt (1986: 164), RO!; isolectotypes, BM! 2 replicates,
FH!, PC-Thuret!, WRSL!).
Synonymized by Munoz (1998b: 394).
Grimmia alpestris f. hybrida Chal., Pamietn. Fizyogr. 2: 68. 1882. Grimmia alpestris var. hybrida
(Chal.) Chal., Enum. Musc. Frond. Tatr.: 56. 1886. Grimmia sessitana f. longifolia Loeske,
Biblioth. Bot. 101: 116. 1930, nom. illeg., incl. typo f. prior. PROTOLOGUE: [Poland and
Slovakia, not explicitly stated]. TYPE: Tatra, Polnischer-Kaum, v.d. Felkaer-Thal. 27-VIII-1879.
T. Chalubiilski (lectotype, designated by Munoz (1998b: 395), BP-36703!).
Synonymized by Munoz (1998b: 395).
Grimmia anceps Boulay, Musc. France 1: 371. 1884. PROTOLOGUE: [France] Rochers siliceux du
massif du Mt-Blanc, sur plusieurs points, vallee de Berard, it gauche du torrent de l'Eau-Noire, en
haut des Praz-d' Avaz, et ailleurs (Payot). TYPE: Massif du Mt. Blanc. Leg. Payot (lectotype,
designated by Munoz (1998b: 394), PC!; isolectotype, BP-36692!).
Synonymized by Munoz (1998b: 394).
Grimmia tenerrima Renauld & Cardot, Bot. Gaz. 15: 40, pI. 6 A. 1890. PROTOLOGUE: [U.S.A.]
Oregon: Mt. Hood, moist bluff towards the snow-line (L.F. Henderson). TYPE: Am. bor. Oregon,
Mt. Hood, moist bluff above snow-line. Leg. Henderson, n° 1239 (lectotype, designated by Munoz
(1998b: 395), PC!; isolectotypes, FH!, NY! 2 replicates).
Synonymized by Munoz (1998b: 395).
Grimmia grisea Cardot, Bull. Herb. Boissier, ser. 2, 6: 7. 1906. PROTOLOGUE: [South Georgia]
Cumberland bay (Typus in herb. Cardot). TYPE: Georgie du Sud: Cumberland bay. Leg.
Skottsberg. 1902. n° 308 (lectotype, here designated, S!; syntypes ["Skottsberg 307"], H-BR!, S!).
Synonymized by Munoz (1998b: 395).
NOTE: No specimens of G. grisea were found in Pc.
Grimmia subpapillinervis Kindb., Rev. Bryol. 36: 98. 1909. PROTOLOGUE: [Canada] British
Columbia 1908: Brinkman, com. Macoun. TYPE: on rocks, Ambean Valley, B.c. July 10, 1908.
ColI. Brinkman (lectotype, designated by Munoz (1998b: 395), CANM-198100!).
Synonymized by Munoz (1998b: 395).
74
MuNoz AND PANDO
Grimmia subcaespiticia Schiffn., Ann. K. K. Naturhist. Hofmus. 27: 490. 1913. PROTOLOGUE:
[Turkey] Ad ropes serpentinicas in regione montana Tauri Cataonici (Kurdistania occidentalis).
Bekikara zwischen Kjachta und Malatja, am siidlichen Talhang, 1600 m, c.sp., 18.NII. 1910 (Nr.
2422). TYPE: Kurdistania occidentalis: Tauros Cataonicus. In convallibus subalpinis prope vicum
Bekikara inter urbem Malatja et vicum Kjachta, ad ropes. Substrato serpentinico; ca. 1600m. Leg.
18-VII-1910. Dr. Heim. Frh. v. Handel-Mazzetti (Tageb.- No 502) (lectotype, designated by
Munoz (1998b: 395), FH!; isolectotype, H-BR!).
Synonymized by Munoz (1998b: 395).
Grimmia asperitricha Dixon & Sainsbury, Trans. & Proc. Roy. Soc. New Zealand 75(2): 173. 1945.
PROTOLOGUE: [New Zealand] On rocks near Hermitage, Mount Cook; colI. G.O.K.S.
29/1/1934, No. 573. On rock, De la Beche, Tasman Glacier; colI. G.O.K.S. 20/1/1934, No. 756.
On rock Malte Bron, Tasman Glacier; coll; G.O.K.S. 22/1/1934, Nos. 757, 771, 772. On rock,
Pinnacles Ridge, Mount Ruapehu; colI. G.O.K.S. January, 1930, No. 799. TYPE: Tasman Glacier,
near De La Beche Hut, South I., N.Z. Coll. G.O.K. Sainsbury n° 756, 20 Jan. 1934 (lectotype,
designated by Munoz (1998b: 395), BM!; syntypes ["Sainsbury 753," "Sainsbury 757,"
"Sainsbury 771," "Sainsbury 772," "Sainsbury 799"] F!, BM!).
Synonymized by Munoz (1998b: 395).
Grimmia donniana f. antarctica Kuc, Rev. Bryol. Lichenol. 36: 659, figs. 1-2, 3 d-e, 4-5. 1969.
PROTOLOGUE: Antarctic, "The Bunger Oasis", ad stationem nomine Dovrovolskii, leg. S.
Rozycki, 25.VI.l959. Herb. Inst. Bot. Acad. Sc. Pol. Cracoviae. TYPE: Antarctica. Queen Mary
Land, Bunger Hills: in the neighbourhood of the Polish Base. 25 June 1959. Coll. S. Rozycki. Det.
M. Kuc (holotype, KRAM!; isotype, MA-7386!).
Synonymized by Munoz (1998b: 395).
DISTRIBUTION: AFG (M. Koie s.n., ALTA); AND (A. Sotiaux 3798, BCB); ARG (C Skottsberg
976, PC); ATA (D.WH Walton 225, MA); AUS (L.G. Adams 1557, FH); AUT (V SchijJners.n., FH);
BEL (A. Comet s.n., BP); BGR (T. Simon s.n., BP); BOL (M. Lewis 79-2092, MA); CAN (A.
Brinkman 222, MO); CHE (J Amann BH-48, Z); CHL (B. Sparre 4832, FH); CHN (C Gao 22522,
ALTA); CZE (l. Gyorffy s.n., JE); DEU (0 Sendtner s.n., GLM); ESP (J.L. Rupidera 2, SALA); FRA
(J Lange s.n., PC); GEO (VF. Brotherus 232, H-SOL); ISL (B. J6hannsson 26913, ICEL); IND (W
Koelz 6774, FH); IRN (HK. Haussknecht s.n., JE); ITA (CC Townsend 92/547, MA); JPN (N. Takaki
133, F); KAZ (B. Allen 10554, MO); KEN (0 Hedberg 1923, PC); MNG (W Schubert M196, MO);
NOR (S.O Lindberg s.n., H-SOL); NZL (G.O.K. Sainsbury 753, BM); PAK (F. Duthie s.n., H); POL
(T. ChaAlbiJiski s.n., BP); ROM (M. Peterfi s.n., BP); SFE (B.C Tan 97-199, FH); SGS (E.G. Bell
1188, BM); SIB (M. Ignatov 0/478, MO); SUN (T. Alexander s.n., H-BR); SVK (L. Vajda s.n., BP);
SWE (0 Gjaerevoll s.n., TRH); TUR (HF.v. Handel-Mazzetti 2422, FH); TZA (Salt 46, PC); UGA
(0. Hedberg 571, MA); USA (A. Nelson 2699, RM); ZAF (R.E. Magill 6122, NY).
62. Grimmia teretinervis Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 216. 1884. Schistidium
teretinerve Lirnpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 216. 1884, nom. alt. PROTOLOGUE:
[Austria] Kalchstein bei Innervillgraten in Tirol vom Pfarrer Hieronymus Gander am 27. Juli 1882
gesamrnelt. TYPE: (BP? not found).
NOTE: Herbarium BP holds many duplicates of G. teretinervis collected by Gander in
Innervillgraten in 1882 and identified subsequently by Limpricht, although not one matches exactly the
WORLD SYNOPSIS OF GRiMMIA
75
date of the protologue. However, the publication shows both pagination and typographical errors, and
under these circumstances a mistake on the transcription of the collection date could be the reason for
the discrepancy.
Grimmia tenuis G. Roth, Eur. Laubm. 2: 691, tab. 62 fig. 4. 1905. Grimmia barkeri W. E. Nicholson &
G. Roth, in G. Roth, Eur. Laubm. 2: 691, tab. 62. 1905, nom. invaI., pro synon. PROTOLOGUE:
[Switzerland] Bei Kandersteg in der Schweiz, dem Hotel Gemmi gegenuber, jenseits des Baches
von Professor J. Barker im Herbst 1890 entdeckt. TYPE: Kandersteg river opposite the Hotel
Gemmi. J. Barker. Ix-i890 (lectotype, here designated, BM!; isolectotype, FH!).
DISTRIBUTION: AUT (J. Baumgartner s.n., FH); CAN (R.R. Ireland 15237, FH); CHE (c.
Meylan BH-JO, Z); CZE (1. Gyorrffy s.n., JE); DEU (T. Herzog s.n., JE); HUN (A. Boros s.n., BP);
ITA (J. Albrecht s.n., Z); ROM (L. Vajda s.n., BP); SIB (Zhuravlev 1734, LE); SVN (J. Breidler s.n.,
BP); SVK (Z Pilous s.n., BP); USA (F.J. Hermann 20157, G).
63. Grimmia tergestina Tomm. ex Bruch & Schimp., Bryoi. Europ. (fasc. 25-28) 3: 126, tab. 258.
1845. Grimmia tergestina var. eutergestina Loeske, Laubm. Eur. Part I: 80. 1913, nom. invai.
PROTOLOGUE: [Italy] In rupibus calcareis umbrosis loco "Bosco di Melara" prope Tergestum.
TYPE: Bosco de Melara bei Triest [... ] zwischen Longera und Bozzai. Ende Marz 1845.
Tommasini (lectotype, designated by Cao & Vitt (1986: 178), BM!; isolectotypes, BM! 3
replicates, H-SOL!, PC!).
Grimmia kansuana Mull. HaL, Nuovo Giorn. Bot. Ital., n.s. 3: 109. 1896. PROTOLOGUE: China
interior, provo Kansu, Alpes ad captantes a flumine Tatung, 8000-9000 ped., in molis vetustis
vulgaris, Augusto 1880: N.M. Przewalski. TYPE: PI. a N.M. Przewalski collectae. China
occidentalis. Regio Tangut (prov. Kansu) ad flum. Tatung, 8-9000'. 1113 VIII.1880 (lectotype,
here designated, PC!; isolectotype, PC!).
Grimmia philibertii Giacom., Atti 1st. Bot. Lab. Crittog. Univ. Pavia, ser. 5, 9: 200. 1950. Grimmia
orbicularis xG. tergestina H. Philib., Ann. Sci. Nat. Bot., ser. 5, 17: 240, pI. 18 fig. 16-24. 1873,
nom. invaI., published as a formula. PROTOLOGUE: Aix, [... ] chaine de coUines appelee Ie
Monteignez, [... ] Ie Chicalon [vallee]. TYPE: Aix en Provence. 10 avril 1873. H. Philibert
(lectotype, here designated, NY!).
Grimmia cucullata D.M. Hend., Notes Roy. Bot. Gard. Edinburgh 12: 614. 1958. PROTOLOGUE and
TYPE: [Turkey] Provo Ankara: above Tuz Golu 25 km. north of Kochisar, c. 925 m., on bare rock
and faces up to 45°,29 July 1956, McNeill 342 (holotype, E!).
DISTRIBUTION: ARM (V. Manakyan 2823a, ERE); AUT (D. Sabransky s.n., BP); BOL (M.
Lewis 79-243, F); CHE (J. Schleicher s.n., GOET); CRN (B.C. Tan 93-1220, FH); DEU (L. Meinunger
1189, JE); DOM (W.R. Buck 8575, NY); ESP (J. Munoz s.n., MA); FRA (H. Philibert s.n., NY); GEO
(VF. Brotherus 222, H-SOL); HUN (A. Degen s.n., BP); IRN (P. Goldblatt 821, MO); IRQ (HF.v.
Handel-Mazzetti 1397, FH); ITA (E. Levier 22, FH); MKD (T Herzog 678, JE); PER (P. Hegewald &
E. Hegewald 5848A, MO); ROM (A. Degen s.n., BP); SAU (W. Frey & H Kurschner 1467, MA); SFE
(V. Komarov 77, LE); SIB (N Martinoff s.n., H-BR); SVK (I. Gy6rffy s.n., JE); TKM (U Mamatkulov
137, MO); TUR (J. McNeill 342, E); UKR (A.S. Lazarenko s.n., MA); USA (D. Toren s.n., MA); YEM
(H Kurschner 6201, MA); YUG (Loitlesberger s.n., JE).
76
MuNoz AND PANDO
64. Grimmia torquata Drumm., Musci Scotici, Vol. 2, n° 28. 1825. Dryptodon torquatus (Drumm.)
Brid., Bryol. Univ. 1: 772. 1827. Grimmia uncinata subsp. torquata (Drumm.) Hampe, Flora 20:
285. 1837. Zygodon torquatus (Drumm.) Liebm. ex Beilschm., Jahresber. Konig. Schwed. Akad.
Wiss. Fortschr. Bot. 1839-42: 430. 1845. PROTOLOGUE and TYPE: [United Kingdom] Clova
mountains. [T. Drummond] (lectotype, designated by Geissler & Maier (1995: 505), E;
isolectotypes, FH! 2 replicates, G!}.
Grimmia cernua f. atra Nees & Homsch., Bryol. Germ. 2(1}: 178, tab. XXIV fig. 23b. 1827.
PROTOLOGUE: [Austria] Ratiteberg 1816 aufgefunden (Mielichhofer! Hsch.! Funck! Laurer!);
spater auch auf Zirknitz-Alpe bei Dollach (Laurer!), und auf andem hohen Alpen von OberKamthen (Funck! Laurer!). TYPE: Herb. Nees v Esenb. Hernn Sturm (lectotype, here designated,
G!).
Grimmia torta Nees & Homsch., Bryol. Germ. 2(1}: 179, tab. XXIV fig. 24. 1827. Grimmia spiralis
var. torta (Nees & Homsch.) Spruce, Musci Pyrenaici, n° 272. 1847. PROTOLOGUE: [Austria]
Wachst auf Felsblocken der hOchsten Alpen, auf dem Rathhausberg in der Gastein (Laurer!
Funck! Hmsch.), auf der Gosnitz-Alpe (Hrsch.), auf dem Nafsfelder-Tauem (Funck! Laurer!), in
der Nahe des Rauriser Tauernhauses (Laurer!), in den Schweizer Alpen (Fr. Nees v. Esenb.).
TYPE: Herb. Nees v Esenb. ex alpibus (lectotype, here designated, G!; syntype ["RauriserTauernhauses. Laurer"], BM!).
Grimmia streptophylla Kindb., Bot. Not. 1882: 187. 1882, nom. illeg., non Sull. 1859. Grimmia
tortifolia Kindb., Christiania Vidensk.-Selsk. Forh. 1888(6}: 19. 1888, nom. nov. Grimmia
stenophylla Kindb. ex H. Moller, Ark. Bot. 26A(2}: 67. 1934, nom. inval., pro synon.
PROTOLOGUE: [Norway] Denna art ar af mig funnen vid Kongsvold pa Dovre ofvanfOr
skogsregionen paornkring 950 meters hOjd ofver hafvet. TYPE: Norveg., Dovre, prope
Kongsvold, 900 metro 23/7 1881 N.C. Kindberg = "G. streptophylla" specirn. originalia!
(lectotype, here designated, S!).
Grimmia prolifera Miill. Hal. & Kindb., in Macoun, Cat. Canad. PI., Musci: 67. 1892.
PROTOLOGUE: [Canada, locality not explicitly stated]. TYPE: Canada, B.C. Selkirk mts. 4300'.
29/7/1890. Leg. J. Macoun. Det. N.C. Kindberg (lectotype, here designated, S!; isolectotype, HBR!).
Grimmia tortifolia subsp. pellucida Kindb., in Roll, Hedwigia 35: 64. 1896. Grimmia torquata subsp.
pellucida (Kindb.) Paris, Index Bryol. ed. 2: 290. 1904. PROTOLOGUE: [U.S.A.] Yellowstone
Nat. Park, Wyom., Upper Falls. No. 1478 [J. Roll]. TYPE: Yellowstone Nat. Park, Wyom., Upper
Falls. 2/9 88. J. Roll 1478 (lectotype, here designated, S!).
Grimmia tortifolia subsp. pseudotorquata Kindb., Eur. N. Amer. Bryin. 2: 217. 1898. Grimmia
torquata subsp. pseudotorquata (Kindb.) Paris, Index Bryol. ed. 2: 290. 1904. Grimmia tortifolia
var. pseudotorquata (Kindb.) Paris, Index Bryol. ed. 2: 290. 1904, nom. inval., pro synon.
PROTOLOGUE: Eur. Norway in Dovrefjeld and Gudbrandsdall. Sweden in Jemtland: P. Dusen.
TYPE: [Sweden. Jamtlands:] Jemtl. Snasahogen 7/93. P. Dusen (lectotype, here designated, S!;
syntypes ["Dovre, Kongsvold"] S! 9 replicates; ["Norway, Faberg, Gudbrandsdall"] S!).
Racomitrium flettii Holz., Bryologist 7: 41, pl. 5. 1904 ["Rhacomitrium"]' Grimmia flettii (Holz.)
Cardot, in Holz., Bryologist 8: 54. 1905. Grimmia torquata var. flettii (Holz.) Vaar., Bryologist
WORLD SYNOPSIS OF GRIMMIA
77
64: 324. 1962. PROTOLOGUE: [U.S.A.] collected by Prof. J.B. Flett on Mt. Tacoma (Mt.
Rainier), at an altitude of 14,519 feet, near one of the steam jets issuing from a fissure in the side
of the rim of the crater, Aug. 10, 1897. TYPE: crater ofMt. Tacoma (Rainier), alt. 14514 ft. J. B.
Flett colI. Aug. 10, 1897 (lectotype, designated by Vaarama (1961: 324), ILL not seen;
isolectotype, NY!).
Grimmia torquata f. gracilis Medelius ex H. Moller, Ark. Bot. 26A(2): 70. 1934. PROTOLOGUE:
[Sweden] Hlirjedalen. Tiinnas, Hamrafjallet R. Oldberg; Jamtland. Are. Enafors 1819 A. Hiilphers.
TYPE: Harjedalen, Tiinnas, Hamrnaren. R. Oldberg (lectotype, here designated, S!).
Grimmia torquata f. longipila H. MOller, Ark. Bot. 26A(2): 70. 1934. PROTOLOGUE: [Sweden.
Norrbotten:] Lule lappmark, Kvikkjokk, Kadepakte 1867 H. Holmgren. TYPE: Lule lappmark,
Kvikkjokk, Kadepakte. 27.7.1867. Leg. H. Holmgren (lectotype, here designated, S!).
Grimmia torquata f. nigrescens H. Moller, Ark. Bot. 26A(2): 70. 1934. PROTOLOGUE: [Sweden.
Norrbotten:] Lule lappmark. Kvikkjokk. Njounjes 1921 M[ihi: H. Moller] m. fl. st. TYPE: Lule
lappmark. Kvikkjokk. Njounjes. 7 Juli 1921. 1000 m. Leg. Hj. Moller (lectotype, here designated,
S!).
DISTRIBUTION: AUT (L. Molendo S.n., FH); BEL (F Gravet s.n., JE); BGR (L. Meinunger
14398, JE); CAN (G.R. Brassard 10510, FH); CHE (H. Greven s.n., MA); CZE (K.G. Limpricht s.n.,
FH); DEU (L. Meinunger 1090., JE); ESP (A.C. Crundwell 462, NY); FIN (V.F Brotherus s.n., JE);
FRA (1. Theriot s.n., FH); GBR (T. Drummond s.n., FH); GRL (s. W Bartlett H-5246, NY); IRE (D.L.
Kelly s.n., TCD); ISL (B. J6hannsson 30100, ICEL); ITA (T. Herzog s.n., JE); NOR (M. Haapasaari
8644, JE); POL (S. Lisowski s.n., PC); SFE (o.M. Afonina s.n., NY); SUN (o.A. Be/kina & A.J.
Likhachev s.n., MO); SWE (C.F Thedenius s.n., FH); USA (FA. MacFadden 16994, MO).
65. Grimmia tortuosa Hook.f. & Wilson, London J. Bot. 3: 540. 1844. PROTOLOGUE: Falkland
Islands. TYPE: Falkland Islands. Antarct. Exp. 1839-1843. J.D.H. (lectotype, designated by
Munoz (1999a: 174), BM!; isolectotypes, BM! 7 replicates, FH! 4 replicates, H-SOL!, NY!).
DISTRIBUTION: FLK (J.D. Hooker 196, BM).
66. Grimmia tricbopbylla Grev., Fl. Edin.: 235. 1824. Grimmia afjinis Homsch. ex Brid., Bryol.
Univ. 1: 771. 1827, nom. inval., pro synon. Dryptodon trichophyllus (Grev.) Brid., Bryol. Univ. 1:
771. 1827. Grimmia trichophylla subsp. eutrichophylla Loeske, Laubm. Eur. Part I: 128, fig. 37,
38. 1913, nom.-inval. PROTOLOGUE: [United Kingdom] On stone-walls about Edinburgh. On
Arthur's Seat, and coast of Fife, Mr. Amott. Lanar road, about seven miles from Edinburgh, and
walls between Colinton and the Pentland Hills, abundant. [... ] I discovered on a wall by the side of
the road from Edinburgh to Lanark, about seven miles from the former place, in company with Mr
Amott. TYPE: near Edinburgh. Greville, 1821 (lectotype, designated by Geissler & Maier (1995:
505), E!; isolectotypes, FH!, H-SOL!).
78
MuNoz AND PANDO
Grimmia consobrina Kunze ex Mull. Hal., Syn. Musc. Frond. 1: 785. 1849. PROTOLOGUE: Chile
australis: poppig. TYPE: Chile. Poppig (lectotype, designated by Muiioz (1999a: 176), NY!;
isolectotypes, BM! 2 replicates).
Synonymized by Deguchi (1984: 25).
Grimmia imberbis Kunze ex Mull. Hal., Syn. Musc. Frond. 1: 788. 1849. PROTOLOGUE: Chile
australis, ad rupes: poppig. TYPE: Kze. Chile (lectotype, designated by Munoz (1999a: 176), IE!).
Synonymized by Muiioz (1999a: 176).
Grimmia trichophylla var. meridionalis Mull. Hal., Syn. Musc. Frond. 1: 785. 1849. Grimmia
trichophylla subsp. meridionalis (Miill. Hal.) Loeske, Biblioth. Bot. 101: 171. 1930.
PROTOLOGUE: [Spain] Hispania, in colibus apricis pr. Granada et Escurial in Sierra de
Guadarrama: W.P. Sch.-Aprili, Majo. TYPE: In colI. siccis prope Granatam. Schpr. 19t.
(lectotype, designated by Muiioz (1999a: 176), H-SOL!; isolectotypes, IE!, NY!; syntype
["Escurial"] NY!).
Synonymized by Muiioz (1999a: 176).
Grimmia crispatula Miill. Hal. & Hampe, Linnaea 26: 499. 1855. Grimmia crispata Mull. Hal. &
Hampe ex Mitt., Trans. Roy. Soc. Victoria 19: 55. 1882, nom. inval. PROTOLOGUE: [Australia.
South Australia] Ad lapides rivuli Fift creek et Flinders range [leg. F. Muller]. TYPE: N. Holl.
Flinders range (lectotype, designated by Munoz (1999a: 176), H-SOL!; isolectotype, MEL1002773; syntypes ["justa rivulum Fift creek"] H-SOL!, MEL-I002771 !).
Synonymized by Munoz (1999a: 176).
Grimmia subsquarrosa Wilson, J. Bot. 4: 231. 1866. Grimmia muehlenbeckii f. propagulifera Limpr.,
Laubm. Deutschl. 1: 764. 1889, nom. nov. Grimmia ovata var. subsquarrosa (Wilson) G. Roth,
Eur. Laubm. 1: 414, tab. 32 fig. 10. 1903. Grimmia trichophylla f. subsquarrosa (Wilson) Loeske,
Laubm. Eur. Part I: 130. 1913, nom. illeg., incl. typo f. prior. Grimmia trichophylla subsp.
subsquarrosa (Wilson) Dixon, Stud. Handb. Brit. Mosses ed. 3: 148. 1924. Grimmia trichophylla
var. subsquarrosa (Wilson) A.lE. Smith, J. Bryol. 9: 279. 1976. PROTOLOGUE: [United
Kingdom] Hill of Moncrieff and Hill of Kinnoul, near Perth, May, 1865 [F .B. White]. TYPE: Hill
of Moncrieff. May, 1865 (Dr. White) (lectotype, here designated, BM!; isolectotypes, BM! 3
replicates).
Grimmia stirtonii Schimp., Syn. Musc. Eur. ed. 2: 270. 1876 ["Stirtoni"]. Grimmia trichophylla f.
propagulifera Limpr., Laubm. Deutschl. 1: 767. 1889. Grimmia trichophylla f. stirtonii (Schimp.)
Loeske, Laubm. Eur. Part I: 129. 1913, nom. illeg., incl. typo f. prior. Grimmia trichophyl/a subsp.
stirtonii (Schimp.) Dixon, Stud. Handb. Brit. Mosses ed. 3: 147.,1924. Grimmia trichophylla var.
stirtonii (Schimp.) H. Moller, Ark. Bot. 26A(2): 87. 1934. PROTOLOGUE: [United Kingdom] pr.
Glasgow, ubi claro Stirton 1866 in rupibus basalticis legit. TYPE: nr. Glasgow. June 1866. D.
Stirton (lectotype, here designated, BM!).
Grimmia trichophylla f. ten ella Besch., Cat. Mouss. Alg.: 18. 1882. PROTOLOGUE: [Algeria]
Edough, rochers des caroubiers, mars 1861 (Let. [Letourneux, A.D. TYPE: Let. 18 mars 1861.
Algerie (lectotype, here designated, PC!).
Grimmia watsonii Lesq. & James, Man.: 140. 1884 ["Watsoni"], Grimmia californica subsp. watsonii
(Lesq. & James) Kindb., Eur. N. Amer. Bryin. 2: 226. 1898. PROTOLOGUE: [US.A.] On rocks,
WORLD SYNOPSIS OF GRIMMIA
79
Dardanelles Canon (Bolander); Alderney, Marin Co., California (Watson). TYPE: Alderney,
Marin Co. Cal[ifornia]. Watson (lectotype, here designated, FH!).
Grimmia densa Kindb., Bull. Torrey Bot. Club 17: 27l. 1890. Grimmia depilata Kindb., in Macoun,
Cat. Canad. PI., Musci: 69. 1892, nom. illeg., inci. typo spec. prior. PROTOLOGUE: [Canada] On
rocks, summit of Mount Benson, near Nanaimo, Vancouver Island, June 3, 1887. Alt. 3,000 feet.
TYPE: N. Amer. Vancouver Island, Mt. Benson, 8/6 87. J. Macoun (lectotype, designated by
Munoz (1999a: 176), S!; isolectotype, CANM-198083!).
Synonymized by Munoz (1999a: 176).
Grimmia cockaynei R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 418, pI. 34 fig. 28. 1895.
PROTOLOGUE: [New Zealand] Rocks, Mount Torlesse; January, 1886. Collected by R.B.
TYPE: (lectotype, here designated, fig. 28 in pI. 34, Brown, R., op. cit.) After an unsuccessful
search for specimens referable to this taxon (BM, CHR, H), the illustration in the original
publication is proposed as lectotype of the name. It unambiguously represents Grimmia
trichophylla Grev.
Grimmia jinitima R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 415, pI. 32 fig. 2l. 1895.
PROTOLOGUE: [New Zealand] Lyttelton hills; 1880. Collected by R.B. TYPE: (lectotype, here
designated, fig. 21 in pI. 32, Brown, R., op. cit.) After an unsuccessful search for specimens
referable to this taxon (BM, CHR, H), the illustration in the original publication is proposed as
lectotype of the name. It unambiguously represents Grimmia trichophylla Grev.
Grimmia flexifolia R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 417, pI. 33 fig. 25. 1895, nom.
illeg., non Hampe 1860. Grimmia brownii Paris, Index Bryoi. Supp!.: 172. 1900, nom. nov.
Grimmia flexuosa Broth., Nat. Pflanzenfam. 3(1): 448. 1902, nom. invaI., orthogr. err.
PROTOLOGUE: [New Zealand] Rocks, Mount Torlesse; January, 1886. Collected by R.B.
TYPE: Mt. Torlesse, North Canterbury, N.z. R. Brown ter. (lectotype, here designated, BM!).
NOTE: The material in BM was annotated by Greven as G. pulvinata var. africana, although it is
typical G. trichophylla. This author did not realize this specimen was a type of the name, and in his
publication on New Zealand Grimmia (Greven, 1998: 397), he blindly followed Dixon (1926: 157) in
considering G. flexifolia synonymous with G. trichophylla. The specimen was listed under G.
pulvinata var. africana, however.
Grimmia petriei R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 418, pI. 34 fig. 29. 1895.
PROTOLOGUE: [New Zealand] Mount Torlesse; January, 1886. Collected by R.B. TYPE: Mount
Torlesse 111186 (lectotype, here designated, BM!).
Grimmia webbii R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 419, pI. 34 fig. 3l. 1895 ["webbi"].
PROTOLOGUE: [New Zealand] Palmer's Pass, head-waters of the River Conway; February,
1894. Collected by R.B. TYPE: On rocks, Palmer Pass 2/94 (lectotype, here designated, BM!).
NOTE: The type material at BM is a small quantity of pure G. trichophylla. It was annotated by
Dixon as G. trichophylla var. stirtonii, although in his later publication he considered this name
synonymous with G. pulvinata var. obtusa. Recently, Greven (1998: 396), after studying the type, and
realizing it was a type, also considered it synonymous with G. pulvinata var. africana.
80
MuNoz AND PANDO
Grimmia sardoa var. gracilis Warnst. & M. Fleisch., Bot. Centraibi. 65(2/3): 299. 1896. Grimmia
trichophylla var. gracilis (Warnst. & M. Fleisch.) Paris, Index Bryoi. SuppI.: 176. 1900. Grimmia
trichophylla f. gracilis (Warnst. & M. Fleisch.) Podp., Consp. Musc. Eur.: 288. 1954.
PROTOLOGUE and TYPE: [Italy] Romano: Monti Albani, Mt. Cavo an Lavafelsen. 900 m.
7.6.1895 M. Fleischer [Fleischer & Warnstorf, Bryotheca europaea meridionalis. Cent. 1, n° 55.
1896] (lectotype, designated by Munoz (1999a: 176), FH!; isolectotypes, GOET!, PC! 2
replicates) .
Synonymized by Munoz (1999a: 176).
Grimmia trichophylla var. vivariensis Husn., Musci galliae, fasc. 18, n° 861. 1897, nom. nud. in sched.
ORIGINAL SPECIMEN: [France] Sur les pierres volcaniques it Labastide (Ardeche).- Mai 1892.
Philibert (FH!).
NOTE: Erroneously listed in Wijk et ai. (1962: 406) as G. tergestina var. vivariensis.
Grimmia trichophylla var. tenerifJae Renauld & Cardot, Bull. Herb. Boissier, ser. 2, 2: 436. 1902.
PROTOLOGUE: [Spain. Tenerife] Las Canadas (n° 52) [A. Tullgren, 1896]. TYPE: Teneriffe:
Las Canadas. Leg. A. Tullgren, 1896 (lectotype, designated by Munoz (1999a: 176), PC!).
Synonymized by Munoz (1999a: 176).
Grimmiafiexiseta Dusen, Rep. Princeton Univ. Exped. Patagonia 8: 122. 1903, nom. nud. ORIGINAL
MATERIAL: Patagonia occ. ORIGINAL SPECIMEN: Argentina, ad lac. Nahualhuapi, ad saxa,
121711897. Leg. P. Dusen (PC!).
Grimmia macropu/vinata Dusen, Rep. Princeton Univ. Exped. Patagonia 8: 78, fig. 8, pI. 8 fig. 7.
1903. PROTOLOGUE: [Chile] Patagonia australis, ubi species verisimiliter in territorio fontinali
fluminis Rio Chico in saxis lecta est. TYPE: Patagonia austr. Collected by J. B. Hatcher. Princeton
Scientific Expedition to Patagonia 1896-97 (lectotype, designated by Munoz (1999a: 176), S!;
isolectotypes, FH!, NY!).
Synonymized by Deguchi (1984: 25).
Grimmia pycnophylla Dusen, Rep. Princeton Univ. Exped. Patagonia 8: 122. 1903, nom. nud.
ORIGINAL MATERIAL: [Chile] Patagon. occ. ORIGINAL SPECIMEN: Patagonia occ. in valle
flurninis Aysen in saxis. Jan. 30 a. 1897 P. Dusen (PC! 2 replicates, S! 3 replicates).
Racomitrium microphyllum M. Fleisch., Musc. Buitenzorg 1: 377. 1904. Grimmia microphylla (M.
Fleisch.) M. Fleisch., Musc. Buitenzorg 4: 1650. 1923. PROTOLOGUE: [Indonesia] An Rinde.
Ost-Java: Ardjoenogebirge, am Waliran! 2600 m. entdeckt (F.) [M. Fleischer, 1902]. TYPE:
Ostjava. Ardjoenogeb. Waliran. 2700 m (holotype, FH!; isotype, FH!).
Synonymized by Deguchi (1986: 328).
Grimmia fiexicaulis var. dicksonii Dusen, Bot. Not. 1905: 303. 1905. Grimmia dicksonii (Dusen)
Dusen, Ark. Bot. 6(10): 14. 1907, nom. nov., nom. illeg., non Turner, 1804. PROTOLOGUE: [no
locality given, but considered as: Patagonia occ., Rio Aysen [Chile] in saxis. Febr.l1.1897, leg. P.
Dusen (cf. Deguchi, 1984: 25)]. TYPE: Patagonia, in vallis fluminis Aysen in rupibus. 1112 1897.
P. Dusen (lectotype, designated by Deguchi (1984: 25), S-PA; isolectotypes, FH! 2 replicates).
Synonymized by Deguchi (1984: 25).
WORLD SYNOPSIS OF GRIMMIA
81
Grimmia pachyphylla Dusen, Bot. Not. 1905: 302. 1905, nom. illeg., non Miill. Hal. 1885.
PROTOLOGUE: [Chile] Patagonia occ. in valle flum. Aysen in saxis. TYPE: Patagonia occ. in
valle flumini Aysen in rup. Febr. 11 a. 1897. P. Dusen (lectotype, designated by Muiioz (1999a:
176), H-BR!).
Synonymized by Munoz (1999a: 176).
Grimmia apocarpa var. sublaevipila Cardot in Broeck, Bull. Jard. Bot. Etat 4: 256. 1914.
PROTOLOGUE: [Belgium] Villers-la-Ville, mai 1882 (Delogne). Villecloye (Meuse) [... ] a la
cote de Thueux que M. J. Cardot [... ] 5 janvier 1883. TYPE: Meuse: Villecloye, fissures de blocs
rochers calcaires a la cote de Thueux. 5 Janvier 1883 (holotype, Blom (1996: 261), BR not seen;
isotype, PC!).
Grimmia trichophylla var. subincurva H. Winter, Hedwigia 55: 101. 1914. PROTOLOGUE: [Spain.
Tenerife] Levada do Gordon [... ] [Marz-Mai 1912]. TYPE: Levada de Gordon (Monte) 900 mts.
Mai 1912 leg. Dr. Winter (lectotype, designated by Muiioz (1999a: 176), JE!).
Synonymized by Munoz (1999a: 176).
Grimmia trichophylla f. viridis Herzog, Kryptog. Forsch. 1(4): 281. 1919. PROTOLOGUE: [Former
Yugoslav Rep. of Macedonia] Macedonien. TYPE: Fl. Mazedonien. An Felsen des Plateauberges
bei Zovik, ca 650 m. Januar, 17 [1917]. Leg. Th. Herzog (lectotype, here designated, JE!).
Grimmia trichophylla f. microtheca Herzog, Bot. Jahrb. Syst. 60: 579. 1926. PROTOLOGUE: [Italy.
Sardinia] Monte Santa Vittoria d'Esterzili bei etwa 1300 m. TYPE: Fl. v. Sardinien. Monte Santa
Vittoria bei Esterzili. Zwischen 900 u. 1200 m. Mai, 06 [1906]. Leg. Th. Herzog (lectotype, here
designated, JE!).
Grimmia gusindei J. Froehl., Ann. Naturhist. Mus. Wien 59: 117. 1953. PROTOLOGUE: [Chile]
Fuegia: Canal Beagle, leg. M. Gusinde, II. 1922, no. 359. TYPE: Fuegia: Canal Beagle. Leg. M.
Gusinde 11.1922 n° 359. det. Froehl. 6.1.1953 (lectotype, designated by Muiioz (1999a: 176), S!).
Synonymized by Munoz (1999a: 176).
DISTRIBUTION: ARG (A. Donat 166, JE); AUS (A. Touw 18717, FH); BEL (F. Gravet s.n., JE);
BOL (M Lewis 87-1812, MO); CAN (P.L. Redfearn 38635, MO); CHL (A. Hollermayer 72, NY);
COL (S.P. Churchill 13329, MO); CRl (G. Dauphin 1690, MO); CYP (F. Ungers.n., H-SOL); DEU
(L. Meinunger 17190, JE); DNK (A. Hesselbo s.n., FH); DZA (A. Letourneux s.n., PC); ECU (E.
Whymper 1347, BM); ESP (J.-L. De Sloover 4235, BCB); FRA (R.B. Pierrot 91007, MA); GBR (D. G.
Long & B.M Murray 14783, MA); GRC (V. Schiffner II, FH); GTM (A.J. Sharp 2211, TENN); HUN
(L. Marstaller s.n., JE); IDN (Anonymous s.n., FH); IRL (D.A. Webb s.n., TCD); ITA (A. Berger s.n.,
NY); KEN (S. Rojkowski 131, F); LBN (W.B. Evans 1034, FH); MAR (P. Davis & J. Davis D-49356,
NY); MEX (A. Cardenas s. 5502, MO); MKD (T Herzog 592, JE); NZL (D.H. Vitt 10666, MO); PAN
(B. Allen 9093, MO); PER (P. Hegewald & E. Hegewald 5467, MO); POL (R. Ochyra s.n., MO); PRT
(A. Ervideira s.n., G); SAU (P. Konig & H. Kurschner 1511, MA); SWE (0.J. Hasslow s.n., MO);
SYR (H.F.v. Handel-Mazzetti 1805, FH); TUR (J. Bornmuller 10085, FH); USA (A.T Whittemore
3289, MO); VEN (D. Griffin III et al. 1328, FLAS); YUG (J. Baumgartner s.n., MO); ZAR (J.L. De
Sloover 13089, NY).
82
MuNoz AND PANDO
67. Grimmia triformis Carestia & De Not., Comment. Soc. Crittog. Ital. 2: 102. 1866. Grimmia
donniana var. triformis (Carestia & De Not.) Loeske, Laubm. Eur. Part I: 96, fig. 26d, 30. 1913.
PROTOLOGUE: [Italy] Precipizii ombrosi del Tagliaferro in Val Sessia, Carestia. TYPE: In
Tagliaferro, a quelche metro sotto la veta suI versante di Rima. 19 Ago 1863. Ab Carestia
(lectotype, designated by Munoz (1998b: 381), BM!; isolectotypes, BM! 2 replicates, BP-37541!,
JE!, PC!).
Grimmia ganderi Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 215. 1884. Grimmia donniana var.
breviseta Breidl. ex Loeske, Laubm. Eur. Part I: 95. 1913. PROTOLOGUE: [Austria] Wurde in
Tirol bei Innervillgraten an glatten Schieferwanden am Wege nach "Kalchstein" bei 4000 Fuss
vom Pfarrer Hieronymus Gander am 15. Marz 1884 gesammelt und als Grimmia triformis De Not.
bestirnmt, womit sie in der That am nachsten verwandt ist. TYPE: Tirol, Innervillgraten, an glatten
Schieferwanden am Wege nach "Kalchstein" 4400' 15/3 1884 Gander (lectotype, designated by
Munoz (1998b: 381), BP-37547!; isolectotypes, JE! 2 replicates, H!).
Synonymized by Munoz (1998b: 381).
DISTRIBUTION: AUT (H. Gander s.n., G); CHE (T. Herzog s.n., H); FRA (R.B. Pierrot s.n.,
BCB); ITA (A. Carestia s.n., JE); SIB (M. Ignatov 0/474, MA).
68. Grimmia trinervis R.S. Williams, Bull. New York Bot. Gard. 3(9): 124. 1903. Coscinodon
trinervis (R.S. Williams) Broth., in Herzog, Biblioth. Bot. 87: 52. 1916. PROTOLOGUE: Juliaca,
Peru, 3780 meters, on dry sandstone, May 15, 1902 (2814). TYPE: Peru. Juliaca. On dry
sandstone. n° 2814. May 15 1902. Collected by R.S. Williams (holotype, NY!; isotype, F!).
DISTRIBUTION: BOL (M. Lewis 79-1960, F); PER (H. Deguchi 29848, HIRO).
69. Grimmia ungeri Jur., Ins. Cypem.: 169. 1865. Grimmia intermedia Braithw., J. Bot. 10: 198.
1872, nom. inval., pro synon. Grimmia alpestris subsp. ungeri (Jur.) Kindb., Bih. Kongl. Svenska
Vetensk.-Akad. Handl. 7(9): 112. 1883. Grimmia alpestris var. ungeri (Jur.) Husn., Muscol. Gall.:
129. 1887. PROTOLOGUE: [Cyprus] Auf dem Troodos an Aphanit-Felsen. TYPE: In Olimpo
Cypri. Vere 862. Leg. Dr. Fr. Unger (lectotype, designated by Munoz (1998b: 398), BM!;
isolectotypes, BM! 2 replicates, GOET! 2 replicates, H-SOL!, S!).
Grimmia brachyodon Austin, Bull. Torrey Bot. Club 6: 45. 1875. Grimmia montana var. brachyodon
(Austin) Lesq. & James, Man.: 145. 1884. PROTOLOGUE: [U.S.A., locality not explicitly given].
TYPE: in rupibus schistosis montis Diablo Califomiae (Bolander); etiam in alpibus Sierra Nevada
montium (Brewer). [Sullivant & Lesquereux, Musci boreali-americani ed. 2, n° 215. 1866
("1865")] (lectotype, designated by Munoz (1998b: 398), FH!; isolectotypes, FH! 4 replicates, G!,
H!, MO!, NY!, PC!).
Synonymized by Munoz (1998b: 398).
Grimmiajamesii Austin, Bull. Torrey Bot. Club 6: 43. 1875. Grimmia montana var. truncata Lesq. &
James, Man.: 145. 1884, nom. nov. PROTOLOGUE: [U.S.A.] Grimmia orbicularis James, in
Watson's Catalogue of the Plants of the 40th Parallel, p. 403. [On rocks near Carson City, Nevada;
WORLD SYNOPSIS OF GRiMMIA
83
5,000 feet altitude. Watson (1,412)]. TYPE: Rocks. Carson. Nevada. Mai 26/68. n° 1412
(lectotype, designated by Muiioz (1998b: 398), FH!; isolectotypes, FH! 2 replicates).
Synonymized by Muiioz (1998b: 398).
Grimmia microtricha Mull. Hal. & Kindb., in Macoun, Cat. Canad. PI., Musci: 70. 1892.
PROTOLOGUE: [Canada] On rocks north of the railway close to Revelstoke, B.c., May 7th,
1890 (Macoun.). TYPE: N. Amer. Brit. Columbia, Revelstoke 7/5 1890 1. Macoun (lectotype,
designated by Munoz (1998b: 398), S!; isolectotype, CANM-198087!).
Synonymized by Munoz (1998b: 398).
Grimmia montana var. idahensis Renauld & Cardot, Bot. Gaz. 30: 18. 1900. PROTOLOGUE:
[U.S.A.] North Idaho: west end of Lake Pend d'Oreille (J.B. Leiberg, 1892. Ren. & Card., Musci
Amer. sept. exsicc. no. 289). TYPE: Idaho: lac Pend d'Oreille. Leg. J.B. Leiberg, 1892. Renauld
& Cardot, Musci Americae septentrionalis exsiccati no. 289 (lectotype, designated by Munoz
(1998b: 398), PC!; isolectotypes, CANM-197540!, FH! 2 replicates, NY!).
Synonymized by Muiioz (1998b: 398).
Grimmia pseudomontana Cardot & Ther., Bot. Gaz. 30: 18, pI. IV fig. 2. 1900. PROTOLOGUE:
[U.S.A.] Idaho: near Moscow, on dry rocks (L.F. Henderson, 1894). TYPE: 4255. Dry rocks, near
Moscow, Idaho. March 24/94 (lectotype, designated by Munoz (1998b: 398), PC!).
Synonymized by Munoz (1998b: 398).
Grimmia canadensis H. Winter, Hedwigia 55: 102, fig. 4. 1914, nom. illeg., non Kindb. 1897
["canadensis"]' Grimmia ovalis subsp. canadensis (H. Winter) Podp., Consp. Musc. Eur.: 279.
1954. PROTOLOGUE: [Spain] Teneriffa: Canadas, in 2000, Hohe, auf Lavablocken [Marz-Mai
1912]. TYPE: Teneriffa, Canadas, 2000 mtr. auf Lavasblocken. April 1912. Leg. Dr. Winter
(lectotype, designated by Munoz (1998b: 398), JE!; isolectotypes, JE! 5 replicates).
Synonymized by Munoz (1998b: 398).
Grimmia mixta Herzog, Bot. Jahrb. Syst. 60: 573. 1926. PROTOLOGUE: [Italy. Sardinia] An Felsen
der Ostseite der Punta La Marmora, etwa 1700 m, 2.Juni.1906. TYPE: An Felsen der Ostseite der
Pta la Marmora, ca 1700m, 2.VI.1906 (holotype, JE!).
DISTRIBUTION: CAN (J.F. Collins 3961, FH); CYP (F. Unger s.n., BM); ESP (H. Winter s.n.,
JE); GBR (J. Fergusson s.n., BM); ITA (T Herzog s.n., JE); MEX (1.L. Wiggins 9177, FH); USA (F.A.
MacFadden 17386, FH).
70. Grimmia unicolor Hook., in Grev., Scott. Crypt. Fl. 3: 123. 1825. Grimmia atrata var. minor
Drumm., Musci americani n° 57. 1828. Dryptodon unicolor (Hook.) Hartrn., Handb. Skand. Fl. ed.
3: 272. 1838. PROTOLOGUE: [United Kingdom] Rocks near Bachnagaim, the Honourable D.
Ogilvie's hunting lodge at the head of Clova; not rare, Mr. Drummond. TYPE: (lectotype,
designated by Geissler & Maier (1995: 505), E not seen).
Grimmia norvegica Bryhn, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1899(3): 26. 1899.
Grimmia unicolor var. norvegica (Bryhn) G. Roth, Eur. Laubm. 1: 413. 1903. Grimmia unicolor f.
norvegica (Bryhn) Loeske, Laubm. Eur. Part I: 88. 1913. PROTOLOGUE: [Norway] In Bykle
prope Bykleli ad rupes declivis juxta fluminem Otteraaen, altitudine supra mare 500 m., sociis
84
MuNoz AND PANDO
Dicrano longifolio et Racomitrio lanuginoso, pridie Nonas Augustas 1896 haud copiose lecta.
TYPE: ad rupes (prope Bykleli) vallis Saetersdalen, Norwegia. 500m. Aug. 1896 leg. N. Bryhn
(lectotype, here designated, Sl).
DISTRIBUTION: AUT (CC Townsend 921530, MA); CAN (B. Allen 9568, NY); CHE (M
Jiiggli s.n., GOET); CHN (D.-K. Li 15197, ALTA); ESP (JE. Zetterstedt s.n., H); ETH (E.F. Gilbert
279, NY); FIN (VF. Brotherus s.n., JE); GBR (G.A.W Arnott s.n., TCD); GRL (WC Steere 62-1171,
NY); IND (CC Townsend 891441, F); KAZ (B. Allen 10562, MO); MNG (W Schubert M152, MO);
NOR (N.C Kindberg s.n., S); SFE (B.C Tan 97-250, FH); SIB (M Ignatov 01464, MA); SUN (A.H.
Brotherus & VF. Brotherus 14, H-SOL); SWE (WH. Arnell s.n., FH); USA (F.J Hermann 28754,
NY).
71. Grimmia wilsonii Greven, 1. Bryol. 20: 398 fig. 1. 1998. PROTOLOGUE: New Zealand, Southern
Alps, Mt. Cook National Park, summit of Sebastopol, 4820 ft., H.D. Wilson 3139, 18 November
1972, [holotype:] CHR 255583 (paratypes listed). TYPE: New Zealand, Southern Alps, Mt. Cook
National Park, track to Hooker Glacier, H.C Greven NZ56 (paratype, MOl); ibidem, track to Sealy
Tarn, H.C Greven NZ59 (paratype, BM1); New Zealand, Homer Saddle, J Child 4285 (paratypes,
BM1, Fl, MAl).
DISTRIBUTION: NZL (H.C Greven NZ56, MO).
WORLD SYNOPSIS OF GR/MMIA
85
TAXA EXCLUDED FROM GR/MMIA
Amphidium tortuosum (Hornsch.) Cufodontis
Grimmia haliacalae Reichardt, Sitzungsber. KaiserL Akad. Wiss., Math.-Naturwiss. CL, Abt. 1, 75:
567. 1877. PROTOLOGUE: [U.S.A. Hawaii] Maui; auf dem Gipfel und im Krater des Haliakala:
Nr. 1900, 1902 [leg. Heinrich Wawra Ritter von Fernsee]. TYPE: Hawaische Inseln, Maui. Auf
dem Gipfel und im Krater des Haliakala. N° 1900 Dr. Wawra (lectotype, here designated, W!;
syntype ["Wawra 1902"], W!).
NOTE: The nhme was changed to Grimmia haleakalae in Wijk et aL (1962: 387), but according to
the ICBN (Art. 60.1), the original spelling must be used.
Arctoa anderssonii Wich.
Grimmia cratericola Sakurai & Takaki, 1. Hattori Bot. Lab. 6: 2, tab. 2. 1951. PROTOLOGUE:
[Japan. Hauts sommets de la mte. Fuji] On soil in crevices of lava, altitude about 3700m. CoIl. N.
Takaki, July 22, 1950. No. 9068. TYPE: (holotype, private herbarium of N. Takaki, not seen;
isotype, MAK not seen).
Synonymized by Deguchi (1978: 248).
Blindia acuta (Hedw.) Bruch & Schimp.
Grimmia gracilis Michx., FL Bor.-Amer. 2: 290. 1803. Grimmia michauxii Torr. ex Eaton, Man. Bot.
ed. 3: 296. 1822. PROTOLOGUE: [U.S.A.] in Carolina; prior detexit D. Beauvois.
Synonymized by Allen (1995: 160).
Brachydontium olympic urn (E. Britton) T.T. McIntosh & 1.R. Spence, Bryologist 89: 200. 1986
[pubL 1987]. Grimmia olympica E. Britton, Bryologist 13: 59, pI. 7. 1910. PROTOLOGUE:
[U.S.A.] Olympic mountains, Washington. Queets river valley, near Humes' glacier. Altitude
5,500 feet. T.e. Frye No. 614, Aug. 14, 1907. TYPE: (holotype, NY not seen).
Synonymized by McIntosh & Spence (1986).
Coscinodon sp.
Grimmia lawiana J.H. Willis, in Filson, The Lichens and Mosses of Mac.Robertson Land: 148, pI. 41.
1966. PROTOLOGUE: Ring Rocks, Mac.Robertson Land, Antarctica, leg. R. Filson No. 4225, 16
September 1962 (MEL). TYPE: Antarctica. MacRobertson Land, Ring Rock. 16 September 1973.
ColI. R.B. Filson 4225 (holotype, MEL!; isotypes, MA!, KRAM!).
NOTE: Grimmia lawiana is the name used for sterile or otherwise poorly developed specimens
collected in Antarctica. Some of the specimens so named are G. plagiopodia, but others, including the
types, are not any Grimmia known to me. They have plicate leaves, plane hair-points and, in my
opinion, should be referred to Coscinodon rather than to Grimmia.
Coscinodon arctolimnius (Steere) Steere, 1. Hattori Bot. Lab. 42: 446. 1977 ["arctolimnia"]. Grimmia
arctolimnia Steere, Bryologist 77: 230. 1974. PROTOLOGUE: Canada. Northwest Territories.
District of Mackenzie: [ ... ] Labine Point, near Eldorado Mine, Port Radium, McTavish Arm, Great
Bear Lake (66°05'N Lat., 118°01'W Long.), 23 July 1948, Steere 10429. TYPE: Plants of the
Region of Great Bear Lake, Northwestern Territoires, Canada. Labine Point, near Eldorado Mine,
Port Radium, east end of McTarvish Arm. William e. Steere No. 10429.23 July 1948 (holotype,
NY!; isotypes, MO!, PC!).
86
MuNoz AND PANDO
Coscinodon bolivianus Broth., in Herzog, Biblioth. Bot. 87: 52. 1916. Grimmia boliviana (Broth.)
F.J. Herm., Bryologist 79(2): 137. 1976. PROTOLOGUE: [Bolivia] Felsen am Huaillattanisee, alt.
4900 m, No. 2971. TYPE: Felsen am Huaillattanisee. 4900 m. Okt. 1911 leg. T. Herzog 2971
(lectotype, here designated, H-BR!; isolectotypes, H-BRl, PC!).
Coscinodon cribrosus (Hedw.) Spruce
Grimmia sinensianodon Miill. Hal., Nuovo Giorn. Bot. Ital., n.s. 5: 188. 1898. PROTOLOGUE: China
interior, provo Schen-si Sept., in loco dicto "Zu-lu" [Sao-y-san], Oct. 1896 [J. Giraldi]. TYPE:
China interior, provincia Schen-si septentr. in loco dicto Zu-lu. 27 Oct. 1896 legit Rev. Jos.
Giraldi, determ. C. Miill. n° 2063 (lectotype, designated by Cao & Vitt (1986: 167), H not seen;
isolectotypes, PC! 2 replicates)
Synonymized with G. caespiticia by Cao & Vitt (1986: 167), but see Munoz (1998b: 389).
Coscinodon humilis Milde
Grimmia brachyphylla Cardot, Bull. Herb. Boissier, ser. 2, 8(5): 333. 1908. PROTOLOGUE: [South
Korea] Coree: Pomasa, rochers a 800 m (n. 218); No-in-tchi, rochers humides a 600 m (n. 590).
TYPE: Coree: Pomasa. 800 m. Leg. Faurie, 1906. nO 218 (lectotype, designated by Munoz (1998b:
391), PC!; isolectotypes, H!, H-BRl, NYl).
Synonymized with G. montana by Cao & Vitt (1986: 161), but see Munoz (1998b: 391).
Coscinodon humilis Horik. & Nog., J. Jap. Bot. 14(1): 25-32. 1938, nom. inval. descr. jap. ORIGINAL
MATERIAL: [Japan] Kyiisyii: Mt. Kudyii (ca 1500 m), provo Bungo (A. Noguchi, no. 3298-typus,
in Herb. Hiros. Univ., Nov. 1932). ORIGINAL SPECIMEN: (HIRO not seen).
Didymodon sp.
Grimmia velutina Kindb., Rev. Bryol. 23: 18. 1896. PROTOLOGUE: On dry rocks. Amer. Canada,
Brit. Columbia: Macoun. TYPE: On the right bank of the Thompson River at Lytton. B.c. April
17th, 1889. ColI. Macoun (lectotype, here designated, CANM-19810l!).
Didymodon rigidulus Hedw.
Campylopus curvifolius Brid., Muscol. Recent. Suppl. 4: 78. 1818 ["1819"]' Dryptodon curvifolius
(Brid.) Brid., Bryol. Univ. 1(1): 201. 1826. PROTOLOGUE: [Poland] E Silesia sub nomine
Grimmia geniculata missum accepi, Seliger invenit. TYPE: E comitatu Glatziam. Seliger. 1802
(lectotype, here designated, B!).
Jaffueliobryum arsenei (Ther.) Ther., Rev. Bryol. Lichenol. n. S. 1: 194. 1928. Coscinodon arsenei
Ther., Smithsonian Misc. Collect. 81(1): 4 fig. 2. 1928. Grimmia churchilliana Crum, Bryologist
89: 24. 1986. PROTOLOGUE: [Mexico] Queretaro: Jurica, upon stones (11001). TYPE: N°
11001. Mexico. Queretaro: Jurica. 1850 m. Leg. Fre. Arsene 5.2.1914 (holotype, PCl).
Jaffueliobryum wrightii (Sull.) Ther.
Grimmia julacea R.S. Williams, Bull. New York Bot. Gard. 6(21): 260. 1910, nom. illeg., non F.
Weber & D. Mohr 1803. Grimmia williamsii Deguchi, Studies on Cryptogams in Southern Peru:
33. 1987, nom. nov. PROTOLOGUE: Arequipa, Peru, 2400 meters, on dry, exposed rock, Aug. 8,
1901 (2805) [R.S. Williams]. TYPE: Arequipa, 2400 meters, on dry rock. R.S. Williams N° 2805.
Aug, 8, 1901 (holotype, NY!; isotypes, Fl, JE! 2 replicates).
WORLD SYNOPSIS OF GRiMMIA
87
Racomitrium spp.
NOTE: Names under Racomitrium spp. are ordered alphabetically.
Grimmia cylindropyxis Mull. Hal., Hedwigia 37: 166. 1898. PROTOLOGUE: Australia, Victoria, Mt.
William: Dr. Sullivan 1883 legit et misit. TYPE: Mt. William. March 1883 [D. Sullivan] 9
(lectotype, here designated, MEL-1002777!).
Grimmia minuta MUll. Hal., Bot. Jahrb. Syst. 5: 81. 1884. Racomitrium minutum (Mull. Hal.) Ochyra
& Matteri, Fragm. Florist. Geobot. 41: 1009. 1996. ,PROTOLOGUE: Kerguelen. TYPE:
(lectotype, designated by Ochyra & Matteri (1996: 1009), S not seen).
Grimmia ochracea MUll. Hal., Bot. Jahrb. Syst. 5: 81. 1884. Racomitrium ochraceum (Mull. Hal.)
Ochyra & Matteri, Fragm. Florist. Geobot. 41: 1008. 1996. PROTOLOGUE: Kerguelen. TYPE:
(lectotype, designated by Ochyra & Matteri (1996: 1008), BM not seen; isolectotypes, BM, LE, S,
none seen).
Guembelia procumbens Mitt., in F. Muell., Fragm. 11, Suppl.: 114. 1880, nom. nud. Grimmia
procumbens Mitt., Trans. Roy. Soc. Victoria 19: 56. 1882. PROTOLOGUE: [Australia]
Gippsland, Snowy River, F.v.M. [Ferdinand von Muller]. TYPE: NY holds specimens from both
localities, labelled "Gipps Land. Dr. F. Muller. 1855" and "Snowy River. Victoria. Australia. Dr.
Muller. 1855" respectively (syntypes, NY!).
Racomitrium crispulum (Hook.f. & Wilson) Hook.f. & Wilson
Grimmia defoliata MUll. Hal., Bot. Jahrb. Syst. 5: 81. 1884. PROTOLOGUE: Kerguelen. TYPE:
Kerguelen. Rotag Cove, Dr. Naumann 1874 (lectotype, designated by Ochyra & Matteri (1996:
1009), S not seen; isolectotype, PC!).
Racomitrium didymum (Mont.) Lorentz
Grimmia depressa MUll. Hal., Flora 68: 421. 1885. PROTOLOGUE: [Argentina] Fuegia, Staten
Island, Port Cook, ad truncos putridos in sylvis, Martio 1882. [Argentina] Fuegia australis, Bumst
Island, ad ramos Chilostrii amelloidis in sylvis, Majo 1882: Spegazzini. TYPE: [Argentina] Ad
truncos putridos, Stat. lsI., P. Cook 3/82. Spegazzini 97 (lectotype, designated by Matteri &
Munoz (2000: 62), LPS!; isolectotype, PC not seen, Ochyra, pers. comm; syntype ["Burnt lsI.
Spegazzini 36"], LPS!).
Synonymized by Matteri & Munoz (2000: 62).
Grimmia depressa vaT. terrestris MUll. Hal., Flora 68: 421. 1885. PROTOLOGUE: [Argentina] Beagle
Channel, in sinu Fandagaia [sic], ad glaream orae maritimae, Majo 1882: Spegazzini. TYPE:
[Argentina] Yendegaia, ad glareum litoris 5/82. N° 31 (lectotype, designated by Matteri & Munoz
(2000: 63), LPS!; isolectotype, PC, Ochyra, pers. comm.).
Synonymized by Matteri & Munoz (2000: 62).
Racomitrium ptycbopbyllum (Mitt.) Mitt., Handb. N. Zeal. Fl.: 426. 1867. Grimmia ptychophylla
Mitt., in Linds., Trans. Bot. Soc. Edinburgh 8: 280. 1866. PROTOLOGUE: Otago, New Zealand.
TYPE: Otago. W.L. Lindsay (holotype, NY!).
88
MuNoz AND PANDO
Schistidium spp.
NOTE: Names under Schistidium spp. are ordered alphabetically.
Grimmia abscondita Cardot, Wiss. Erg. Schwed. Siidpolar-Exp. 4(8): 107, fig. 19. 1908. Schistidium
absconditum (Cardot) Ochyra, Fragm. Florist. Geobot. 43: 104. 1998. PROTOLOGUE:
[Argentina] Patagonie australe: lac San Martin, Rio Fosiles, vers 1000 m (Dusen, n° 5944). TYPE:
(PC not seen).
Grimmia angustissima P. de la Varde, Ark. Bot., n.s. 3: 148, fig. 9. 1955. PROTOLOGUE: [Uganda]
Ru [Mt. Ruwenzori]: Bukurungu Valley, in the side-valley on the S.E. slope ofMt. Gessi, 4400 m,
on steep rock (680k, type). TYPE: Uganda: Ruwenzori, Bukurungu Valley in the side valley on
the S.E. slope of Mt. Gessi, on steep rock. 4400 m.s.m. 4.4.1948. ColI. Olov Hedberg 680k. Det.
PV (holotype, PC!).
Grimmia antipodum Miill. Hal., Hedwigia 37: 165. 1898. Schistidium antipodum (Mull. Hal.) Ochyra,
Fragm. Florist. Geobot. 43: 105. 1998. PROTOLOGUE: [Australia] Tasmania, ad saxa, Frocy's
Gully, Novbr. 1890: W.A. Weymouth in Rh. Burchard, qui pro determinatione 1891 misit. TYPE:
Australia. Tasmania Frocy's Gully, Burchard 256 (lectotype, designated by Bremer (1980), H-BR
not seen).
Grimmia austroapocarpa Miill. Hal. ex Watts. & Whitel., Proc. Linn. Soc. New South Wales, ser. 2,
27: 82. 1902, nom. nud. ORIGINAL MATERIAL: [Australia, no locality given]. ORIGINAL
SPECIMEN: Parramatta. J. Whitelegge Aug 28/85 (H-BR!).
Grimmia calycina Herzog, Beih. Bot. Centralbl. 27(2): 354. 1910. Schistidium calycinum (Herzog)
Ochyra, Fragm. Florist. Geobot. 43: 105. 1998. PROTOLOGUE: Bolivia: Cacaltaya, 4800 m, leg.
O. Buchtien, Marz, 08. TYPE: Bolivia. Cacaltaya, 1908. Buchtien (isotype, Bremer (1980: 95), S
not seen).
Grimmia celata Cardot, Bull. Herb. Boissier, ser. 2, 6: 7. 1906. Schistidium celatum (Mull. Hal.) B.G.
Bell, Brit. Antarct. Surv. Bull. 63: 86. 1984. PROTOLOGUE: [South Georgia] Cumberland bay:
connecting valley between S. and W. fiords. TYPE: Georgie du Sud: Baie Cumberland, vallee
entre les fjords S et 0.21/5 1902 Carl Skottsberg. Det. 1. Cardot (lectotype, here designated, PC!).
Grimmia chubutensis Cardot & Broth., Kong!. Svenska Vetenskapsakad. Hand!. 63(10): 26, tab. 2 fig.
6. 1923. Schistidium chubutense (Cardot) Ochyra, Fragm. Florist. Geobot. 43: 105. 1998.
PROTOLOGUE: [Argentina] Patagonia andina: Territ. Chubut, Valle 16. de Octubre, ad saxa:
FutaleufU, Casa Rees. TYPE: 432. Patagonie: Territoire Chubut, Vallee 16 Octobre, pierres dans
la foret de Libocedrus chilensis, melange au Barbula flagellaris. 6.11.1908. C. Skottsberg
(lectotype, PC!; syntypes ["Skottsberg 439"], H-BR!, UPS!).
Grimmia coloradensis Austin, Bot. Gaz. 2: 109. 1877 ["coloradense"]. Schistidium coloradense
(Austin) Ochyra, Fragm. Florist. Geobot. 43: 105. 1998. PROTOLOGUE: [U.S.A.] Colorado,
1876, Brandegee (Rau). TYPE: Colorado, Nov. 9/76, Brandegee. Herb. Eugene A. Rau (lectotype,
here designated, NY!).
WORLD SYNOPSIS OF GRIMMIA
89
Grimmia cupularis Mull. Hal., Bot. Jahrb. Syst. 5: 80. 1884. Schistidium cupulare (Mull. Hal.) Ochyra,
Fragm. Florist. Geobot. 43: 105. 1998. PROTOLOGUE: Kerguelen. TYPE: Kerguelen. XI 74.
Naumann (isotypes, H-BR!, PC!).
Grimmia diversifolia Kindb., Rev. Bryol. 32: 36. 1905. PROTOLOGUE: [Canada] 2. Found by J.
Macoun 1904 in British Columbia, 5200-7500 feet above the sea. TYPE: Canada. B.c., Pipestone
pass, 7500 f. 5/7 04. Macoun (lectotype, designated by Bremer (1980), S!).
Grimmia donatii Herzog & Ther., Repert. Spec. Nov. Regni Veg. 41: 290, tab. 236 e-n. 1937.
Schistidium donatii (Herzog & Ther.) Ochyra & Matteri, Fragm. Florist. Geobot. 41: 1008. 1996.
PROTOLOGUE: [Chile] Territorio Aisen. Regio Lago San Martin. Ventisquero, L.3, ca. 1500 m,
leg. A. Donat, 13.V.33 (no. 48). TYPE: A. Donat Plantae patagonicae n° 48. Terr. Aisen. Reg.
L.S.M. Ventisquero L.3. Alt. ca. 1500 m. Dat. 13.11.33 det Theriot, X.36 (holotype, JE!; isotype,
JE!).
Grimmia fal/ax Dusen, Rep. Princeton Univ. Exped. Patagonia 8: 77, fig. 7, pI. 8 figs. 5-6. 1903.
Schistidium fal/ax (Dusen) Ochyra & Matteri, Fragm. Florist. Geobot. 41: 1008. 1996.
PROTOLOGUE: [Chile] Patagonia australis, ubi verisimiliter in territorio fontinali fluminis Rio
Chico in saxis lecta est. TYPE: Princeton Scientific Expedition to Patagonia 1896-97. South
Patagonia. Collected by J.B. Hatcher (holotype, Ochyra & Matteri (1996: 1008), S not seen;
isotype, NY!).
Grimmia hyalinocuspidata Mull. Hal., in Neumayer, Int. Polarforsch., Deut. Exped. 2: 315.
Racomitrium hyalinocuspidatum (Miill. Hal.) Kindb., Enurn. Bryin. Exot., Suppl. 2: 108.
Schistidium hyalinocuspidatum (Miill. Hal.) B.G. Bell, Brit. Antarct. Surv. Bull. 63: 90.
PROTOLOGUE: [South Georgia] Austro-Georgia, ad rupes des Koppenberges 19. Majo
Sudwest-Gletscherthal, 7. Majo 1883. TYPE: Austro-Georgia, ad rupes Koppenberg Dr.
19.V.1883 (syntype, PC!).
1890.
1893.
1984.
1883;
Will.
Grimmia kiyoshii S. Okamura, J. CoIl. Sci. Imp. Univ. Tokyo 38(4): 12, fig. 5. 1916. PROTOLOGUE:
[Japan] Hondo: Provo Idu, in monte Amagi (Leg. Kiyoshi Fujii! 18.VIII.l914). TYPE: (isotype,
Deguchi (1978: 220), TNS not seen).
Synonymized with S. gracile by Deguchi (1978: 220).
Grimmia konoi Broth., in Kono, Bot. Mag. (Tokyo) 20: 79. 1906. PROTOLOGUE: [in Japanese].
TYPE: (lectotype, designated by Deguchi (1978: 220), H-BR not seen).
Synonymized with S. gracile by Deguchi (1978: 220).
Grimmia longicaulis Dixon, J. Bot. 76: 221. 1938. PROTOLOGUE and TYPE: Limuru, Kenya
Colony, 5 Jan. 1908; colI. G. Balbo (322) (holotype, BM!).
Grimmia macleana Rehmann ex Kindb., Enum. Bryin. Exot., Suppl. 1: 91. 1889, nom. nud.
ORIGINAL SPECIMEN: [South Africa] Terra Capensis Orientalis, Valley of Desolation. leg.
Mac Lea. Rehmann, Musci austro-africani cont., n° 493 (PC!).
90
MuNoz AND PANDO
Grimmia macrotyla Cardot & Broth., Kong!. Svenska Vetenskapsakad. Handl. 63(10): 26, tab. 2 fig. 5.
1923. Schistidium macrotylum (Cardot & Broth.) Ochyra, Fragm. Florist. Geobot. 43: 106. 1998.
PROTOLOGUE: [Argentina] Patagonia andina: Territ. Chubut, Meseta Chalia frequentissime, alt.
12-1300 m.s.m. (382) [Leg. C. Skottsberg n° 382]. TYPE: Patagonie: territ. Chubut, meseta
Chalia. Leg. C. Skottsberg, 1908. n° 382 (lectotype, here designated, PC!; isolectotypes, H-BR!,
UPS!).
Grimmia occulta Mull. Hal., in Neumayer, Int. Polarforsch., Deut. Exped. 2: 314. 1890. Schistidium
occultum (Mull. Hal.) Ochyra & Matteri, Fragm. Florist. Geobot. 41: 1008. 1996.
PROTOLOGUE: [South Georgia] Austro-Georgia, inter muscos, 6. Febr. 1883 cum fruct. vetustis
et junioribusi. TYPE: Austr. Georgia. Drekkappel. Dr. Will 6 II 1883 (lectotype, designated by
Ochyra & Matteri (1996: 1008), HBG not seen; isolectotypes, PC!).
Grimmia perichaetialis P. de la Varde, Ark. Bot., n.s. 3: 149, fig. 10. 1955. PROTOLOGUE: [Uganda]
Ru [Mt. Ruwenzori]: Mijusi Valley, E. slope ofMt. Speke, 4550 m, on moraine (580d, type); id.,
W. slope ofMt. Gessi, 4450 m, on stone, close to a small snow field (487); Bujuku Valley, S.W.
slope ofMt. Speke, 4500 m (705a). Ke [Mt. Kenya]: N.W. slope, Buguret Valley, 4450 m (1915).
TYPE: Uganda: Ruwenzori, Mijusy Valley, Eastern slope of Mt. Speke, on moraine. ColI. Olov
Hedberg 580d, det. PV (holotype, PC!; syntypes ["Hedberg 487"], PC!; ["Hedberg 705a"], PC!;
["Hedberg 1915"], PC!).
Grimmia perichaetialis var. pilifera Demaret, Bull. Soc. Roy. Bot. Belgique 88: 92. 1956.
PROTOLOGUE: Kenya: Mont Kenya, au-dessus de la Vallee Teleki, alt. 4500 m, mars 1949,
Salt. n° 26 (holotype, BR not seen).
Grimmia perplexa Ther., Revista Chilena Hist. Nat. 27: 11, him. 2. 1923. Schistidium perplexum
(Ther.) Ochyra, Fragm. Florist. Geobot. 43: 106. 1998. PROTOLOGUE: [Chile] La Ermita, las
Condes, alt. 1,200 m (leg. Bertho et laffuel, oct. 1919); Cajon del Arrayan, alt. 1,200 m (leg.
Bertho, nov. 1919). TYPE: Chili: La Ermita (Las Condes) 1200 m. Leg. M. Bertho, oct. 1919
(lectotype, here designated, FH!; isolectotype, FH!).
Grimmia pseudorivularis Kindb., Eur. N. Amer. Bryin. 2: 234. 1898. Schistidium pseudorivulare
(Kindb.) Ochyra, Fragm. Florist. Geobot. 43: 106. 1998. PROTOLOGUE: [Canada] Amer. r.
Can.: Macoun. TYPE: Canada. Vancouver isl. 22/5/93 Macoun 159 (lectotype, here designated,
S!).
Grimmia rivulariopsis R.S. Williams, Bull. Torrey Bot. Club 43: 327, pI. 19. 1916. Schistidium
rivulariopsis (R.S. Williams) Ochyra, Fragm. Florist. Geobot. 43: 106. 1998. PROTOLOGUE:
[Bolivia] Ollantaytambo, about 3,000 m., on rock in stream bed, May, 1915, Cook & Gilbert 753;
and on rock, same place and collectors, 561. TYPE: Peru. Ollantaytambo; altitude about 3000
meters. No. 753 O.F. Cook & G.B. Gilbert Collectors. May 15, 1915 (lectotype, designated by
Deguchi (1987: 34), NY!; isolectotype, F!).
Synonymized with S. rivulare by Deguchi (1987: 34).
Grimmia saxatilis Mitt., 1. Linn. Soc., Bot. 12: 97. 1869. Schistidium saxatile (Miill. Hal.) Ochyra,
Fragm. Florist. Geobot. 43: 106. 1998. PROTOLOGUE: [Ecuador] Andes Quitenses, in monte
Chimborazo ad rivulos, Spruce, n. 132. TYPE: Chimborazo in rivulis. Spruce. 132 (lectotype,
designated by Bremer (1980: 93), NY!; isolectotypes, FH!, PC! 2 replicates).
WORLD SYNOPSIS OF GRIMMIA
91
Grimmia scabrifolia Broth., Bernice P. Bishop Mus. Bull. 40: 12, tab. 4 fig. 13. 1927.
PROTOLOGUE: [U.S.A. Hawaii] East Maui: on ground (B. 236) [D.D. Baldwin, n° 236]. TYPE:
Insulae Sandwich in insula Maui orient., ad terram, 8000 p. 1876. Legit D.D. Baldwin sub n° 236
(isotypes, H-BR!, NY!).
Synonymized with S. strictum by Cao & Churchill (1995: 511).
Grimmia serratomucrata Mull. Hal., Bot. Jahrb. Syst. 5: 80. 1884. Schistidium serratomucronatum
(Mull. Hal.) Ochyra, Fragm. Florist. Geobot. 43: 107. 1998. PROTOLOGUE: Kerguelen. TYPE:
Kerguelens. Dr. Neumann. Nov. 1874 (lectotype, here designated, PC!).
Grimmia sinensiapocarpa Mull. Hal., Nuovo Giorn. Bot. Ital., n.s. 5: 187. 1898. Schistidium
sinensiapocarpum (Mull. Hal.) Ochyra, Fragm. Florist. Geobot. 43: 107. 1998. PROTOLOGUE:
China interior, provo Schen-si sept., prope locum "Zu-Iu", Oct. 1896 c.fr. deoperculatis [J.
Giraldi]. TYPE: China interior, provincia Schen-si sept., Zu-lu in valle Lao-y-san. 28 Oct. 1896
legit Rev. Jos. Giraldi, determ Prof. C. Muller sub. n° 2051 (isotypes, PC! 2 replicates).
Grimmia syntrichiacea Mull. Hal., in Neumayer, Int. Polarforsch., Deut. Exped. 2: 315. 1890.
Schistidium syntrichiaceum (Mull. Hal.) B.G. Bell, Brit. Antarct. Surv. Bull. 63: 97. 1984.
PROTOLOGUE: [South Georgia] Austro-Georgia, Felsblocke des sudlichen Ufers der
Landzunge, inter Blindiam pallidifoliam n. sp. spec imina perpauca, 13. Oct. 1882. TYPE: Austro
Georgia. 13.x.1882 Dr. Will (isotype, PC!).
Grimmia urnulacea Mull. Hal., in Neumayer, Int. Polarforsch., Deut. Exped. 2: 313. 1890. Schistidium
urnulaceum (Mull. Hal.) B.G. Bell, Brit. Antarct. Surv. Bull. 63: 90. 1984. PROTOLOGUE:
[South Georgia] Austro-Georgia, ad rupes am Ausgange des Brockenthales, cum Giimbelia
immerso-leucophaea consociata, 23. Januario 1883. TYPE: (lectotype, designated by Ochyra
(1990: 135), HBG not seen; isolectotype, S not seen).
Guembelia cyathocarpa Hampe, Linnaea 37: 515. 1872. Grimmia cyathocarpa (Hampe) Mitt., Trans.
Roy. Soc. Victoria 19: 55. 1882. PROTOLOGUE: [Australia] New South Wales, Blue Mountains,
planta riparia videtur. TYPE: [Australia] New South Wales. Blue Mountains. F.v. Muller
(lectotype, here designated, BM!).
Guembelia lorentziana MUll. Hal., Linnaea 43: 450. 1882. Grimmia lorentzi (MUll. Hal.) Kindb.,
Enum. Bryin. Exot., Suppl. 1: 91. 1889. Grimmia lorentziana (Mull. Hal.) Paris, Index Bryol. 2:
529. 1895. Schistidium lorentzianum (Mull. Hal.) Ochyra & Matteri, Fragm. Florist. Geobot. 41:
1008. 1996. PROTOLOGUE: Argentinia subtropica Tucumanensis, in Sierra de Aconquija prope
Tafi et in via ab Tafi per Quebrada de Monteros, ineunte aprilis 1872, [... ]. TYPE: Argentinia.
Tafi. Leg. Lorentz (lectotype, designated by Ochyra & Matteri (1996: 1008), JE!; isolectotype,
BM!).
Guembelia praemorsa Mull. Hal., Linnaea 43: 452. 1882 ["Giimbelia"]' Grimmia praemorsa (MUll.
Hal.) Kindb., Enum. Bryin. Exot., Suppl. 1: 91. 1889. PROTOLOGUE: Argentinia Catamarcana,
supra Yakutula in convallis altae Granadillas, in Vayas altas [sic] alpina, ineunte Februarii 1872,
[... ]. TYPE: Argentinia, in alpinis prope Catamarca, Yakutula, in Vayas altas, convallis altas
Grandillas [sic]. Febr. 1872 leg P.G. Lorentz (isotypes, BM!, JE!).
92
MuNoz AND PANDO
Schistidium amhJyophylIum (Mull. Hal.) Ochyra & Hertel, Polish Botanical Studies 1: 26. 1990.
Grimmia amblyophylla Mull. Hal., Syn. Musc. Frond. 1: 779. 1849. PROTOLOGUE: Insula
Eremitae et Kerguelen ad Cap. Horn: J.D. Hooker. TYPE: Hermite Island, Cap Horn. Antart. Exp.
1839-1843 n° 122. J.D.H. (lectotype, designated by Ochyra & Hertel (1990: 26), BM not seen;
isolectotypes, NY! 2 replicates).
Grimmia stylostegia MUll. Hal., Bot. Jahrb. Syst. 5: 80. 1884. PROTOLOGUE: Kerguelen. TYPE:
Kerguelen. Dr. Naumann (isotype, PC!).
Grimmia antarctici var. pilifera E.B. Bartram, Ann. Missouri Bot. Gard. 25: 722. 1938.
PROTOLOGUE: [Antarctica] Mary Bird Land, Edsel Ford Range, Mt. Rea-Cooper, P. Siple, F.A.
Wade, S. Corey & O.D. Stancliff 27. TYPE: Byrd Antarctic Expedition II. Mosses of Marie Bird
Land, Antarctica. Locality: Mt. Rea. Collected by Paul A. Siple n° 27. Determined by: Edwin B.
Bartram. On granite sand (holotype, FH!; isotype, NY!).
Schistidium atrichum (MUll. Hal. & Kindb.) W.A. Weber, Phytologia 33: 106. 1976. Grimmia
atricha MUll. Hal. & Kindb., in Macoun, Cat. Canad. PI., Musci: 65. 1892. PROTOLOGUE:
[Canada] On the sloping face of broken rocks at the summit of the mountains back of Sproat,
Columbia River, B.c., alt. 4,000 feet, June 24th, 1890 (Macoun). TYPE: On broken rocks, alt.
4,000 feet. Sproat Mountain, Columbia River, B.c. June 24th, 1890. CoIl. J. Macoun (isotypes,
CANM-198078!, FH!, NY!).
Schistidium chenii (S.H. Lin) T. Cao, C.H. Gao & J.C. Zhao, J. Hattori Bot. Lab. 71: 69. 1992.
Grimmia himalayana P.c. Chen, Report on Scientific Expedition of Qomolangma Region: 227.
1962, nom. illeg., non Mitt., 1859. Grimmia chenii S.H. Lin, BioI. Bull. Dept. BioI. ColI. Sci.
Tunghai Univ. 60: 747. 1984, nom. nov. PROTOLOGUE and TYPE: China: Tibet, Himalayan
region, Mt. Jolrno Lungma. Principal Lung-pu glacier, alt. 5450 m., on wet and moist rocks,
1959.VII.9, Wang 86 (holotype, PEl).
Schistidium chrysoneuron (MUll. Hal.) Ochyra, Cryptog. Bryol. Lichenol. 7: 59. 1986. Grimmia
chrysoneura Mull. Hal., Bot. Jahrb. Syst. 5: 80. 1884. PROTOLOGUE: Kerguelen. TYPE:
Kerguelen. XII 74. Naumann (isotypes, H-BR!, PC!).
Grimmia antarctici Cardot, Bull. Herb. Boissier, ser. 2, 6: 15. 1906. PROTOLOGUE: Shetland du
Sud: lIe Nelson, Harmony Cove, Terre Louis-Philippe: debarquement a. Archipel de Graham: ile
Paulet. TYPE: Archipel de Graham: ile Paulet. Leg. Skottsberg, 1903. n° 449 (syntypes, FH!, HBR!,NY!).
Grimmia antarctici var. percompacta E.B. Bartram, Ann. Missouri Bot. Gard. 25: 723. 1938. Grimmia
pilifera var. percompacta H.A. Crum, Bryologist (index 1-60): 88. 1959, nom. inval.
PROTOLOGUE: [Antarctica] King Edward VII Land: Rockefeller Mts., Mt. Helen Washington,
P. Siple, F.A. Wade, S. Corey & O.D. Stancliff 68a. TYPE: Bird Antarctic Expedition II. Mosses
of King Edward VII Land, Antarctica. Locality: Mt. Helen Washington. Collected by: Paul A.
Siple n° 68a. Determined by: Edwin B. Bartram (holotype, FH!; isotype, NY!).
WORLD SYNOPSIS OF GRIMMIA
93
Schistidium cribrodontium (Herzog) Ochyra, Acta Bot. Acad. Sci. Hung. 28: 373. 1982. Grimmia
cribrodontia Herzog, Repert. Spec. Nov. Regni Veg. 41: 287, tab. 235. 1937. PROTOLOGUE:
[Uganda] Africa centralis: Ruwenzori, alpine Stufe, 4650 m, leg. I. Hauman VIINIII, 32 (no.
854). TYPE: Ruwenzori. Etage alpin, 4650 m. Legit L. Hauman, julliet-aout 1932. n° 854
(holotype, JE!).
Schistidium falcatum (Hook.f. & Wilson) B. Bremer, Lindbergia 6: 1l0. 1980. Grimmia falcata
Hook.f. & Wilson, London J. Bot. 3: 539. 1844. Schistidiumfalcatum (Hook.f. & Wilson) Zanten,
Marion and Prince Edwards Islands 2: 193. 1971, nom. inval., basionyrn not cited.
PROTOLOGUE: Kerguelen's Island. TYPE: Kerguelen's Land. Antarct. Exp. 1839-43. J.D.H. n°
245 (isotypes, F!, FH!).
Grimmiafasciculata Dusen, Bot. Not. 1905: 302. 1905, nom. illeg., non Brid. 1819. PROTOLOGUE:
[Chile/Argentina] Patagonia septentrionalis in saxis. TYPE: (lectotype, designated by Ochyra &
Bell (1984: 78), S not seen; isolectotype, UPS not seen).
Synonymized by Ochyra & Bell (1984: 78).
Schistidium flaccidum (De Not.) Ochyra
Grimmia aethiopica Mull. Hal., Bot. Zeitung (Berlin) 13: 765. 1855. PROTOLOGUE: [Ethiopia]
Abyssinia, in rupibus montium Simensium, inde a 9000 pedibus supra mare usque ad 13000 pedes
ascendens: W. Schimper 1838 6. Januar. legit. TYPE: Schimperi iter Abyssinicum. Sectio
secunda. 465. Schistidium pulvinatum Brid. In rupibus montium Simensium, indea [sic] 9000
pedibus supra mare usque ad 13000 pedes d. 6. Jun. 1838 U.i. 1842 (lectotype, here designated,
NY!).
Schistidium canariense H. Winter, Hedwigia 55: 99, fig. 3. 1914. Grimmia winteri Luisier, Broteria,
Ser. Bot. 17: 39. 1919, nom. illeg., incl. typo spec. prior. PROTOLOGUE: [Spain] Aufgenommen
in Teneriffa an den Abh1ingen der Cumbre gegen Villa de Orotava in 1800 m Hohe von
Lavablocken [Miirz-Mai 1912]. TYPE: Teneriffa. Abhiingen der Cumbre gegen Villa de Orotava.
1800 mtrs. Miirz 1912. Leg. Winter (lectotype, here designated, JE!; syntype ["April 1912"], JE!).
Schistidium frisvollianum H.H. Blom
Grimmia elatior subsp. rufesgens Kindb., Rev. Bryol. 34: 90. 1907. PROTOLOGUE: Canada, Hudson
Bay, Baffm land, on earth: A.P. Low 1904, com. 1. Macoun. TYPE: Canada, Hudson bay, Baffm
land. 15/804. A.P. Low (lectotype, designated by Blom (1996: 87), S!).
Synonymized by Blom (1996: 87).
Schistidium heterophyllum (Kindb.) T.T. McIntosh, Bryologist 93: 483. 1990. Grimmia heterophylla
Kindb., in Macoun, Bull. Torrey Bot. Club 17: 271. 1890, nom. nud. Grimmia heterophylla
Kindb., Cat. Canad. PI., Musci: 64. 1892. PROTOLOGUE: [Canada] between Nicola and
Thompson rivers at Spence's Bridge, B.c., alt. 3,500 feet, May 28th, 1889 (Macoun.). TYPE:
Spence's Bridge, B.C. ColI. Macoun. May 28, 1889. [Macoun, Canadian cryptogams n° 29]
(isotype, FH!).
94
MuNoz AND PANDO
Grimmia concinodontoides Kindb., Bull. Torrey Bot. Club 17: 271. 1890, nom. nud. Grimmia
chloroblasta Kindb., in Macoun, Cat. Canad. PI., Musci: 64. 1892. Grimmia coscinodontoides
Kindb., in Macoun, Cat. Canad. PI., Musci: 64. 1892, nom. inval. Grimmia coscinodontea Kindb.
ex Paris, Index Bryol. Suppl.: 172. 1900, nom. inval. PROTOLOGUE: [Canada] on dry rocks on
the mountain overlooking the Thompson River at Spence's Bridge, B.C., alt. 3,500 feet, May 28th,
1889 (Macoun). TYPE: on rocks above Spence's Bridge. B.C. CoIl. Macoun. Date May 28th.
1889 (isotypes, CANM-198081!, NY!).
Schistidium lancifolium (Kindb.) H.H. BJorn, Bryophyt. Biblioth. 49: 55. 1996. Grimmia
pseudorivularis subsp. lancifolia Kindb., Eur. N. Amer. Bryin. 2: 234. 1898. PROTOLOGUE:
[Canada] Amer. Can.: Macoun. TYPE: Canada, Ottawa, wet rocks 23/9931. Macoun (lectotype,
designated by Bremer (1980: 95), S not seen}.
Schistidium liliputanum (Mull. Hal.) Deguchi, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 16: 229.
1978. Grimmia liliputana Mull. Hal., Nuovo Giorn. Bot. Ital., n.s. 5: 188. 1898. PROTOLOGUE:
China interior, provo Schen-si sept., in loco dicto "Zu-Iu", Oct. 1896, inter Grimmiam sinensianodontem vigens [1. Giraldi]. TYPE: China interior, provincia Schen-si septentr. Zulu in valle
Lao-y-san. 27 Oct. 1896, legit Rev. Jos. Giraldi, deterrn. C. Muller n. 2063b (lectotype, designated
by Deguchi (1978: 229), FI not seen; isolectotypes, PC! 2 replicates}.
Grimmia otii Sakurai, in Oti, Bot. Mag. (Tokyo) 62: 144, fig. 4. 1949. PROTOLOGUE: [Japan]
Sikoku: Provo Iyo (Leg. K. Oti no. 1228 - Type Novemb. 1945). TYPE: (holotype, Deguchi (1978:
229), MAK not seen).
Synonymized by Deguchi (1978: 229).
Schistidium maritimum (Turner) Bruch & Schimp.
Grimmia kindbergii Holz., Bryologist 14: 32. 1911. PROTOLOGUE: [U.S.A.] Catala, Alaska.
Collector: Mr. G.c. Britton [1904] [ ... ] Type in the herbarium of Prof. 1.B. Flett, and of the author.
TYPE: Catala, Alaska. CoIl. G.C. Britton, summer of 1904 (isotypes, H-BR!, NY!).
Synonymized by Deguchi (1978: 233).
Schistidium olivaceum (Herzog) Ochyra & J. Munoz, J. Bryol. 22: 142. 2000. Grimmia olivacea
Herzog, Biblioth. Bot. 87: 56, fig. 18g-m. 1916. PROTOLOGUE: [Bolivia] 1m oberen Chocayatal
an Felsen, ca. 4400 m, No. 3587. TYPE: Plantae in itinere secundo per Boliviam lectae. An Felsen
im oberen Chocayatal. ca. 4400m. Leg. Th. Herzog n° 3587. Juli 1911 (holotype, JE!; isotype,
PC!).
Schistidium readeri (Broth.) Ochyra & J. Munoz, 1. Bryol. 22: 142. 2000. Grimmia atricha Miill.
Hal., Hedwigia 37: 162. 1898, nom. illeg., non Mull. Hal. & Kindb. 1892. Grimmia australasica
Mitt. ex Watts & Whitel., Proc. Linn. Soc. New South Wales, ser. 2, 27: 82. 1902, nom. inval., pro
synon. Grimmia readeri Broth., Nat. Pflanzenfam. 1(3}: 448. 1902, nom. nov. Grimmia
atrichoides Paris, Index Bryol. ed. 2: 266. 1904, nom. illeg., incl. typo spec. prior. Schistidium
atrichum (Mull. Hal.) Ochyra, Fragm. Florist. Geobot. 43: 105. 1998, nom. illeg., non (Miill. Hal.
& Kindb.) W.A. Weber. PROTOLOGUE: Australia, Victoria, ad saxa prope Melbourne legit Fr.
M. Reader. Aug. 1886 et misit 1891 ex Dimboola.
WORLD SYNOPSIS OF GRIMMIA
95
Schistidium robustum (Nees & Homsch.) H.H. Blom
Grimmia homodictyon Dixon, Rev. Bryol. 28: 12. 1901. PROTOLOGUE: On a detached block of
limestone, crags south of Inchnadamph, Sutherlandshire, Scotland; July, 19, 1899, W.E.
Nicholson, E.S. Salmon and H.N. Dixon. TYPE: [United Kingdom] Inchadamph, Sutherland, 19
July 1899, Nicholson, Salmon & Dixon (holotype, S not seen).
Synonymized by Blom (1996: 149).
Schistidium scabripes (E.B. Bartram) Deguchi, Studies on Some Patagonian Species of Grimmiaceae
(Musci, Bryophyta): 27. 1984. Grimmia scabripes E.B. Bartram, Farlowia 2: 310. 1946.
PROTOLOGUE: Chile: Provo de Nuble, Recinto, Las Trancas, ad truncos Nothofagi antarcticae,
alt. 1150 m., No 1003 type. TYPE: Chile: Provo de Nuble, Recinto, Las Trancas, ad truncos
Nothofagi antarticae, alt. 1150 m. Apr. 14, 1929. H. Roivainen no. 1003 (holotype, FHl).
Schistidium squamatulum (Herzog) Ochyra & I Munoz, I Bryol. 22: 142. 2000. Grimmia
squamatula Herzog, Biblioth. Bot. 87: 56, fig. 18 a-f. 1916. PROTOLOGUE: [Bolivia] An Felsen
der Yanakakabastion, ca. 4600 m, sparlich, No. 3825/a. TYPE: An Felsen des Yanakakabastion
ca. 4600 m. leg. Th. Herzog Juli 1911 (lectotype, designated by Munoz (1999a: 184), JEl).
Schistidium subconfertum (Broth.) Deguchi, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 16: 240.
1979 ["1978"]' Grimmia subconferta Broth., Symb. Sin. 4: 45. 1929. PROTOLOGUE: [China] S.:
An nassen Schieferfelsen der Hg. St. auf dem Gonschiga sw von Muli gegen Dschungdien, 4650
m, c. sp. 6. VIII. 1915 (7493). TYPE: Provo Schschwan austro-occid.: In montis Gonschiga a
monast. Muli ad septentr. pagi Yiinnanensis Yungming versus Dschungdien siti reg. alp., ad rupes
udas. Substr. schistac. ca. 4650 m. 6.VIII.1915 leg. Hand.-Maz. (holotype, Deguchi (1978: 240),
H-BR not seen; isotype, PCl).
Schistidium tenerum (IE. Zetterst.) Nyholm, Ill. Moss Fl. Fennoscandia II. Musci 6: 775. 1969.
Grimmia tenera J.E. Zetterst., Kongl. Svenska Vetenskapsakad. Handl. 13(13): 17. 1876.
Schistidium tenerum (J.E. Zetterst.) Nyholm, Illustrated Moss Flora of Fennoscandia II. Musci 2:
146. 1956, nom. inval., basionym not cited. PROTOLOGUE: [Sweden] Alten, ad pedem montis
Konghavnsfjellet; Talvig, in montibus juxta Talvigfjord praecipue prope arnnem Isbarelfven.
TYPE: Finm. Alten, Talvigfjord. 25/7/1868. Norge. IE. Zetterstedt (lectotype, designated by Cao
& Churchill (1995: 511), S not seen; isolectotypes, FHl, NYl).
Grimmia ehlei Amell, Ark. Bot. 13(2): 63, Taf. 2. 1913. PROTOLOGUE: [Russia] Sibiria, in valle
fluminis Lena, Kumachsur, 70° 30 lat. sept., ubi anno 1898 ab H. Nilsson-Ehle lectum. TYPE:
Sibiria, in valle flum. Lena, Kumachsur, 70°30' lat. bor. 18/7/1898. Leg. H. Nilsson-Ehle. Det. H.
Wilh. Amell (isotypes, H-BRl, NYl).
Synonymized by Savicz-Lyubitzkaya & Smimova (1970: 473).
Schistidium yaulensis (Broth.) Ochyra & J. Munoz, I Bryol. 22: 142. 2000. Grimmia yaulensis
Broth., Bot. Jahrb. Syst. 56(2), Beibl. 123: 11. 1920. PROTOLOGUE: [Peru] An der Lima-OroyaBahn, Hacienda Arana bei Yauli; Felsenporphyr; 4400 m (373 ex p.) [Leg. A. Weberbauer].
TYPE: 373 ex p. Peru, an der Lirna-Oroya-Bahn, Hacienda Arana bei Yauli, Felsen-Porphyr,
4400m 18/1/1902. Leg. A. Weberbauer (lectotype, designated by Munoz (1999a: 184), H-BRl;
isolectotype, PCl 2 replicates).
96
MuNoz AND PANDO
Tortula caninervis (Mitt.) Broth.
Grimmia cucullata 1.S. Lou & P.C. Wu, Acta Phytotax. Sin. 18: 121, tab. 2 figs. 1-7. 1980, nom.
illeg., non D.M. Hend. 1958. PROTOLOGUE: [Tibet] Xizang: Baingoin, 5600 m. alt., 27
V1.1976, K.Y. Lang 1317 (Type in Herb., Institute of Botany, Academia Sinica). TYPE:
(holotype, PE not seen).
Synonymized by Cao & Vitt (1986: 233).
Trichostomum arcticum Kaal.
Grimmia spitsbergensis Bizot & Ther., Bull. Sci. Bourgogne 5: 70. 1935. Grimmia ovalis subsp.
spitsbergensis (Bizot & Ther.) Podp., Consp. Musc. Eur.: 279. 1954. PROTOLOGUE: [Norway]
Spitsberg, King's Bay. Leg. E. Maire, 1931. TYPE: Spitsberg. Kings Bay. Leg. Em. Maire
21. 7 .1931 (lectotype, here designated, PC!; isolectotype, NY!).
Synonymized by Frisvoll (1978: 158).
Verrucidens insularis (Mitt.) Ochyra & Matteri, Fragm. Florist. Geobot. 41: 1007. 1996. Grimmia
insularis Mitt., 1. Linn. Soc., Bot. 15: 73. 1876. PROTOLOGUE: [Antarctica] Heard Island [H.N.
Moseley]. TYPE: Heard Island. Antarctic. Challenger Expedn. 1874 (lectotype, here designated,
NY!; isolectotype, NY!).
Synonymized by Ochyra & Matteri (1996: 1007).
WORLD SYNOPSIS OF GRiMMIA
97
NAMES WITH TYPE MATERIAL NOT SEEN
Campylopus ovalis var. politus Myrin, KongI. Vetensk. Acad. Handl. 1831: 258. 1832.
PROTOLOGUE: [Sweden] I Dalsland pa Henriksholrns sodra udde, vid Oxnas bruk o. fl. St.
Campylopus pulvinatus var. tenuis Wahlenb., Fl. Suec. 2: 748. 1826. Grimmia trichophylla var. tenuis
(Wahlenb.) Wijk & Margad., Taxon 8: 106. 1959. PROTOLOGUE: [Sweden] B in campestribus
saxis regionum interiorum ad Upsaliam et Vermlandiam usque passim.
Dicranum arcuatum Schleich., Cat. PI. Helv. ed. 2: 28. 1808, nom. nud. Trichostomum arcuatum DC.,
FI. Frans:. ed. 3, 6: 218. 1815. Trichostomum patens var. arcuatum (DC.) Duby, Bot. Gall. 2: 574.
1830. PROTOLOGUE: [Switzerland] [... ] dans les Alpes, par M. Schleicher; [France] dans les
Cevennes, par M. Grateloup.
NOTE: A syntype ["Schleicher. Originaux utilises par A.-P. de Candolle pour la redaction de la
Flore frans:aise. Trichostomum arcuatum DC"] in G is Grimmia elatior. Trichostomum arcuatum
predates the name G. elatior and should be used according to the ICBN. Since another syntype is listed
in the protologue, there exists the possibility of typifying on another element to maintain
nomenclatural stability and avoid the undesired name change. If the other syntype also proves to be G.
elatior, a well-known and long-used name, in my opinion the latter name should be proposed for
conservation to preserve nomenclatural stability.
Dicranum pulvinatum var. argentatum Turner, Musc. Hibern. Spic.: 78, tab. 3 fig. 1. 1804. Dryptodon
pulvinatus var. argentatus (Turner) Brid., Bryol. Univ. 1(1): 197. 1826. PROTOLOGUE: [United
Kingdom] in Eblanae muris a D. Scott, et in muris prope Powerscourt, a D. Stokes lecta.
Dicranum pulvinatum var. muticum Chevall., Fl. Gen. Env. Paris 2(1): 41. 1827. PROTOLOGUE:
[France. Paris].
Dryptodon caespiticius var. muticus Brid., BryoI. Univ. 1(1): 201. 1826. Grimmia alpestris var. mutica
(Brid.) De Not., Syllab. Musc.: 244. 1838. PROTOLOGUE: In eodem loco [Switzerland. In monte
sancti Bernhardi Helvetiae].
Dryptodon hartmanii f. rivularis Loeske, Moosfl. Harz.: 194. 1903. Grimmia hartmanii f. rivularis
(Loeske) Loeske, Biblioth. Bot. 101: 179. 1930. PROTOLOGUE: Am Bodeufer unterhalb
Tresebg. [Germany. Treseburg].
Dryptodon hartmanii var. alatus Katie, Prosv. Glasn. 4: 885-886. 1907. PROTOLOGUE: [in SerboCroatan: Rocks at Deli Jovan, below Crni Vrh, at Jasenicka river source, ca. 1050 m. May. Leg. D.
Katie].
Dryptodon hartmanii var. giganteus Jeanb. & Renauld ex Warnst., Krypt.-Fl. Brandenburg 1(2): 302.
1904. Grimmia hartmanii f. gigantea Jeanb. & Renauld, Guide Bryol. Pyrenees. Part 2: 55. 1885,
nom. nud. Grimmia hartmanii var. gigantea (Warnst.) Podp., Consp. Musc. Eur.: 289. 1954.
PROTOLOGUE: aus den Pyrenaen bei 1750 m MeereshOle.
Dryptodon ovatus var. denticulatus Brid., BryoI. Univ. 1(1): 203. 1826. PROTOLOGUE: In Britanniae
rupibus. Green.
98
MuNoz AND PANDO
Dryptodon ovatus var. elongatus Brid., Bryol. Univ. 1(1): 203. 1826. PROTOLOGUE: [Italy] In
Calabriae rupibus Clar. Reynier legit et communicavit.
Dryptodon patens f. cataractarum Vilh., Acta Bot. Bohem. 2: 52. 1923. PROTOLOGUE: [Slovakia]
[... ] montium tatrensium regionis alpinae (supra Velicke pleso, 1900 m; supra zabi pleso, 2000 m;
Mlinica sub lacum Skok, an. 1919 Vilh.).
Dryptodon patens f. humilis Vilh., Acta Bot. Bohem. 2: 52. 1923. PROTOLOGUE: [Slovakia] [... ]
regionis alpinae in Sudetis (Braunkessel apud Pecer, Panscka louka in montibus Corconticis) et in
Tatris (Mlinica circum lacum Skok, 1820 m; sub Rysy, 2100 m, sub Velicke pleso in monte
Gerlach, sub Ze1ene pleso apud Krivaii, an. 1919 Vilh.).
Dryptodon patens f. pilifer Vilh., Acta Bot. Bohem. 2: 53. 1923 ["pilifera"]. PROTOLOGUE:
[Slovakia] [... ] ca. 1600-2000 m in Tatris sub 5 pIes in Mala studena dolina (an. 1919 Vilh.).
Dryptodon patens f. robustus Vilh., Acta Bot. Bohem. 2: 52. 1923 ["robusta"]. PROTOLOGUE:
[Slovakia] [... ] montium tatrensium (Mlinica ca. 1500 m, an. 1919 Vilh.).
Dryptodon pulvinatus var. canus Hartrn., Handb. Skand. Fl. ed. 3: 270. 1838. Grimmia pulvinata var.
cana (Hartrn.) Hartrn.,-Handb. Skand. Fl. ed. 5: 376. 1849. Grimmia pulvinata f. cana (Hartrn.)
Monk., Laubm. Eur.: 365. 1927. PROTOLOGUE: B D. sudeticus frtm Cimbritsharnn [Norway],
ed.2.
Grimmia albida Spreng., Syst. Veg. 4(2): 321. 1827, nom. illeg., non F. Weber & D. Mohr 1803.
Grimmia albicans Spreng. ex H. Moller, Ark. Bot. 26A(2): 135. 1934, nom. inval., orthogr. err.
PROTO LOGUE: [South Africa: Caput Bon Spei] C.B.S. Ecklon.
Grimmia alpestris f. longipila Podp., VerOff. Geobot. Inst. Rubel Zurich 1: 245. 1924.
PROTOLOGUE: [Switzerland] Zernez: [... J oberhalb der Buffalorawiese.
Grimmia alpestris var. microstoma Bruch & Schirnp., Bryol. Europ. (fasc. 25-28) 3: 129. 1845.
Grimmia subsulcata Lirnpr., Laubm. Deutschl. 1: 757. 1889. Grimmia sessitana var. subsulcata
(Lirnpr.) Breidl., Mitt. Naturwiss. Vereines Steierrnark 28: 88. 1891, nom. illeg., incl. typo var.
prior. Grimmia alpestris subsp. subsulcata (Lirnpr.) Kindb., Eur. N. Amer. Bryin. 2: 221. 1898.
Grimmia microstoma (Bruch & Schirnp.) Dalla Torre & Sarnth., Fl. Tirol 5: 260. 1903. Grimmia
alpestris f. subsulcata (Lirnpr.) Loeske, Laubm. Eur. Part I: 104. 1913. Grimmia alpestris var.
subsulcata (Lirnpr.) Broth., Nat. Pflanzenfam. ed. 2, 10: 309. 1924. Grimmia sessitana f.
subsulcata (Lirnpr.) Loeske, Biblioth. Bot. 101: 117. 1930. PROTOLOGUE: Ad rupes siccas per
Alpes editiores Helvetiae, Gerrnaniae, Delphinatus et Norvegiae [no locality given for the variety].
NOTE: In a previous publication, Munoz (1998b: 394) erroneously considered Grimmia
subsulcata as having a different type from Grimmia alpestris var. microstoma.
Grimmia anodon var. brevipila C.E.O. Jensen, in Dusen, Bih. Kongl. Svenska Vetensk.-Akad. Hand!.
27, Afd. 3(1): 56. 1901, nom. nud. ORIGINAL MATERIAL: [Greenland, Ostgronlands] HurryInlet [... J[Leg. A.G. Nathorst, 1899].
WORLD SYNOPSIS OF GRIMMIA
99
Grimmia anodon var. moravica Podp., Vestn. Klubu Ptir. v Prostejove 7: 18. 1905. PROTOLOGUE:
[Czech Republic. Moravia, not explicitly stated].
Grimmia anodon var. taeniophyllum Mikut. ex Loeske, nom. inval., orthogr. err., Laubm. Eur. Part I:
45. 1913.
NOTE: Error in Wijk et al. (1969: 648) for Schistidium angus tum var. taeniophyllum Mikut.,
Bryoth. BaIt. 1909.
Grimmia apiculata var. mutica Schultz, Syll. PI. Nov. 2: 135. 1828. PROTOLOGUE: [no locality
given].
Grimmia apiculata var. obtusa Schimp., Coroll. Bryol. Eur.: 47. 1856. PROTOLOGUE: [no locality
given].
Grimmia argyrotricha Mull. Hal., Flora 73(5): 485. 1890. PROTOLOGUE: [Tanzania] Africa or.
trop., monte Kilima-Ndscharo, 3000-4800 m: Dr. Hans Meyer 1880.
Grimmia arvernica H. Philib., Rev. Bryol. 9: 25. 1882. Grimmia plagiopodia var. arvernica (H.
Philib.) Boulay, Musc. France 1: .389. 1884. Grimmia plagiopodia f. arvernica (H. Philib.)
Loeske, Biblioth. Bot. 101: 83. 1930. PROTOLOGUE: [France] [ ...] trouvee par M. Gautier
Lacroze au Mont-Juzet, pres de Clermont, a une altitude denviron 430 metres, sur un mur forme
de blocs de lave et de mortier.
Grimmia austropulvinata Mull. Hal., Hedwigia 37: 160. 1898. PROTOLOGUE: Nova Seelandia,
insula septentrionalis, prope Auckland ad rupes montis Wellington, Aug. 1882: T.F. Cheeseman.
Grimmia bifrons De Not., Comment. Soc. Crittog. Ital. 2: 102. 1866. Grimmia alpestris subsp. bifrons
(De Not.) Podp., Consp. Musc. Eur.: 282. 1954, nom. inval. (ICBN Art. 34.2). Grimmia donniana
var. bifrons (De Not.) Podp., Consp. Musc. Eur.: 280. 1954, nom. inval. (ICBN Art. 34.2).
Grimmia donniana subsp. bifrons (De Not.) Wijk & Margad., Taxon 9: 190. 1960.
PROTOLOGUE: [Italy] Dai monti della Sardegna centrale.
Grimmia caespiticia var. bornmuellerorum Schiffn., Oesterr. Bot. Z. 58: 312. 1908. PROTOLOGUE:
[Iran. Tehran: Damavand] Persia bor.: Demawend, in Lavageroll unterhalb des Gipfels sehr
verinzelt als pflanzeliche Spuren, 4000-5400 m s. m; 17. VII. 1902 (Bornm., Iter Pers. alt. 1902,
Nr.5898).
Grimmia camonia Rota ex De Not., Atti Reale Univ. Genova 1: 755. 1869. PROTOLOGUE: [Italy]
Crepacci delle rupi umide del monte Tonale.
Grimmia campestris f. longirostrata Vilh., Vestn. Knil. Ceske Spolecn. Nauk, n. Mat.-ptir. 1924: 14.
1924. PROTOLOGUE: [Czech Republic] Habitat ad rupes arenosas in Bohemia prope Kralupy n.
VI., ad rupes quarzosas apud Stechovice prope Pragam (Vel. [VelenovskY]).
Grimmia campestris var. lutescens Monk., Laubm. Eur.: 356. 1927. Grimmia laevigata var. lutescens
(Monk.) Podp., Consp. Musc. Eur.: 276. 1954. PROTOLOGUE: [Italy] Erhielt ich von Mauem in
Bordighera in Norditalien (leg. Alwin Berger).
100
MuNoz AND PANDO
Grimmia campylotricha Mull. Hal., Flora 71: 414. 1888. PROTOLOGUE: Monte Kilimandscharo, ad
fmem sylvestrem superiorem inter 3000-4000 m.
Grimmia canescens Schleich. ex Spreng., Mant. Prim. Fl. Hal.: 55. 1807. Grimmia ovata var.
canescens (Spreng.) Brid., Bryol. Univ. 1(1): 177. 1826. PROTOLOGUE: In rupibus Vallesiae.
Schleicher.
Grimmia cardotii var. gymnostoma 1. Froehl., Ann. Naturhist. Mus. Wien 67: 156. 1964.
PROTOLOGUE: Afghanistan, Provo Kabul, in valle Pagman prope Kabul, ca. 34°36'N, 68°56'E,
ca. 2300-2800 m, substr. "Gneill"; 21.VI.1962: n° 19.704.
Grimmia cardotii xG. anodon Culm., Rev. Bryol. 53: 8. 1926, nom. inval., published as a formula.
Grimmia cernua Nees & Hornsch., Bryol. Germ. 2(1): 174, tab. XXIV fig. 23a. 1827.
PROTOLOGUE: Wachst auf dem Rauriser-Tauern am sogenannten Draba-Felsen in Gesellschaft
mehrerer Gattungsverwandten, und wurde daselbst im Jahr 1816 zuerst entdeckt (Hoppe! Hsch.),
in der Gastein (Mielichhofer! Funck!), beim Kalserthor in der Nahe des Glockners (Funck!), am
ersten Leiterkopf auf der Pasterze, auf der Zirknitz-Alpe bei Dollach, dem Rathhausberg in der
Gastein und dem Goldberg in der Raurifs (Laurerl), auf der Hohe der Scheidecker-Alpe im
Oberhaslithal in der Schweiz (Hsch.).
Grimmia coarctata Miill. Hal., Hedwigia 37: 159. 1898. PROTOLOGUE: Nova Seelandia, insula
australis, Canterbury, Lyttleton, ad rupes: T.W. Naylor Beckett, 1892; T.F. Cheeseman 1889 in
Rh. Levier.
Grimmia commutata f. basaltica Vilh., Vestn. Knil. Ceske Spolecn. Nauk, Tf. Mat.-Prir. 1924: 15.
1924. PROTOLOGUE: [Czech Republic] Habitat in Bohemia ad rupes basalticas (Vinarickii hora,
1897, Vel.[VelenovskY]).
Grimmia commutata f. subepilosa Braun-Blanq., Rev. Bryol. Lichenol. 23: 102. 1954, nom. nud.
ORIGINAL MATERIAL: [Morocco] Ourgouz, 2.450 m., Si Zerekten [... ]; Ourgouz, 2.500 m.
Grimmia commutata var. brevipila Warnst., Krypt.-Fl. Brandenburg 1(2): 294. 1904. Grimmia
commutata f. brevipila (Warnst.) Vilh., Vestn. Knil. Ceske Spolecn. Nauk, Tf. Mat.-Prir. 1924: 16.
1924. PROTOLOGUE: [Switzerland] sammelte Prof. Ascherson bei Locarno (Schweiz) an
trockenen Gneissfe1sen am Wege nach Ponte Brolla 2. Okt. 1885.
Grimmia commutata var. gibba Boulay, Musc. France 1: 367. 1884. Grimmia ovalis var. gibba
(Boulay) Podp., Consp. Musc. Eur.: 279. 1954. PROTOLOGUE: [France] 9a et hi, Vosgues, St.
Etienne (Pierrat); Pelvoux, Grandvillard pres Brian90n (B.); Mt-Louis (Renauld).
Grimmia commutata var. maroccana Meyl., Bull. Soc. Sci. Nat. Maroc 11: 115. 1931, nom. nud.
Grimmia commutata var. microcarpa Lesq., Mem. Soc. Sci. Nat. Neuchate13(3): 27. 1846, nom. nud.
ORIGINAL MATERIAL: sur les blocs erratiques de Rio, Val-de-Travers. Mai [L. Lesquereux].
Grimmia commutata f. minor Laz., Ukrajins'k. Bot. Zhurn. 3: 12. 1926. PROTOLOGUE: in Ukra'inia
in districto Mariupol, Kamjani Mohyly in rupibus graniticis. 16.1X.1925 [leg. A.S. Lazarenko].
WORLD SYNOPSIS OF GRiMMIA
101
Grimmia commutata f. rivularis Loeske, Moosfl. Harz.: 186. 1903, nom. nud. Grimmia commutata f.
rivularis Loeske, Laubm. Eur. Part I: 87. 1913. Grimmia ovalis f. rivularis (Loeske) Podp., Consp.
Musc. Eur.: 279. 1954. PROTOLOGUE: [Germany] [... ] im unteren Bodetal des Harzes [... ] [Leg.
L. Loeske].
Grimmia compacta Kindb., in Macoun, Cat. Canad. PI. 7: 222. 1902, nom. nud.
Grimmia compactula Mull. Hal., Hedwigia 37: 167. 1898. PROTOLOGUE: Australia, Victoria,
Grampians, Mt. Kosciusco: Dr. Sullivan 1884 in Rh. Melbourne.
Grimmia crassifolia var. cucullata Baumgartner, Ann. Naturhist. Mus. Wien 50: 534. 1939.
PROTOLOGUE: Mittleres Elburs-Gebirge: Berge urn Kalak nachst Keredj, ca. 1400 m (2283).
TYPE: [Iran] Persia: Montes Elburs [Elburz] centr., in ditione oppidi Keredj, in montibus ad
pagum Kalak. 20.V.l937 leg. K.H. Rechinger fil. n° 2283. Det. 1. Baumgartner. (isotype, H!).
NOTE: The isotype in H is sterile, and in this condition is not possible to separate G. tergestina
from G. poecilostoma.
Grimmia crinita var. brevis Boulay, Musc. France 1: 388. 1884. Grimmia crinita f. brevis (Boulay)
Podp., Consp. Musc. Eur.: 275. 1954. PROTOLOGUE: [France, not explicitly stated].
Grimmia crinita var. calabra Zodda, Malpighia 23: 48. 1909. PROTOLOGUE: [Italy] Raccolta con
frutti in aprile 1908 sulle rupi arenoso-ca1caree sopra Melito Porto Salvo, 150 m., dal Prof.
Nicotra.
Grimmia crinita f. compacta Besch., Cat. Mouss. Alg.: 16. 1882. PROTOLOGUE: [Algeria] Mont de
Sel, Djebel Gharribou, EI-Outaia (La Perr.[De La Perraudiere], mars 1858).
Grimmia crinita var. elongata Bruch & Schimp., Bryol. Europ. (fasc. 25-28) 3: 112, tab. 23713. 1845.
Grimmia perichaetialis Durieu & Coss., in Besch., Cat. Mouss. AIg.: 17. 1882, nom. inval., pro
synon. PROTOLOGUE: [France] ad muros ca1careos Monspeliae.
Grimmia crinita var. laevis Boulay ex Pit., Mem. Soc. Bot. France 54: 13. 1907, nom. nud.
ORIGINAL MATERIAL: [Spain] Tenerife: Canadas du pie de Teyde, entre EI Portillo et
Montana Blanca, 2000 it 2500 m.
Grimmia curvifolia Lindb. ex Kindb., Bot. Not. 1882: 146. 1882, nom. nud.
Grimmia decipiens f. falcata Luisier, Mem. Real Acad. Ci. Exact. Madrid, ser. 2, 3: 111. 1924.
PROTOLOGUE: [Spain. Salamanca] Sq.: [Sequeros] Las Batuecas.
Grimmia decipiens var. homomalla Bryhn, Nyt Mag. Naturvidensk. 32: 120. 1892. Grimmia schultzii
Bryol. ed. 2: 287. 1904. Grimmia decipiens f. homomalla
var. homomalla (Bryhn) Paris, ｾｮ､･ｸ＠
(Bryhn) I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1909(5): 43. 1909.
PROTOLOGUE: [Norway] Insula Tjoemo in ditione oppidi Toensberg ad rupes gneissicas
interdum irroratas prope Kjaerelia socia Jungermanniae autumnalis var subapicali (4 Aprilii
1886).
102
MuNoz AND PANDO
Grimmia decipiens f. major Luisier, Mem. Real Acad. Ci. Exact. Madrid, ser. 2, 3: 111. 1924.
PROTOLOGUE: [Spain] S.: Salamanca: Montalbo.
Grimmia donniana var. curviseta Lesq. & James, Man.: 143. 1884. PROTOLOGUE: Monitor,
California (Lapham); Yakima River, Cascade Mountains (Watson).
Grimmia donniana f. longiurna Villi., Vestn. Knil. Ceske Spolecn. Nauk, Ti'. Mat.-Pi'ir. 1924: 18.
1924. PROTOLOGUE: [Czech Republic] Habitat in montibus Corconticis et Tatris. [... ]
Krkonosich (vrchol Snezky, Labsky vodopad, Obi'i dill, 1919, Villi.) a ve Vys. Tatrach (u hotelu
Kolbach, 1919, Vilh.).
Grimmia donniana f. sudetica Chal., Pamietn. Fizyogr. 2: 56. 1882. Grimmia donniana f. breviseta
Villi., Vestn. Kral. Ceske Spolecn. Nauk, Ti'. Mat.-Prir. 1924: 18. 1924, nom. illeg., incl. typo f.
prior. PROTOLOGUE: [Poland] Siodelko miydzy ｂｹｳｴｲｾ＠
a ｋ｡ｭｩ･ｮｳｾ＠
od zachodu, na skalach
pokruszonych; bardzo obficie. A takZe gran od siodelka az pod sam szczyt Kamienistej. Rohacze
Zachodnie i Srednie; granie i pod same szczyty, w wielu miejscach; Pod ｳ｡ｭｾ＠
｣ｺｵ｢ｾ＠
Wolowca i
granie ku Klinowi. Kondracka granie ku Czerwonym Wirchom. Koncysta, od strony Tupej.
Grimmia drakenbergensis Sim, Trans. Roy. Soc. South Africa 15: 209. 1926. PROTOLOGUE: [South
Africa] Natal. Top of Giant's Castle, R.E. Symons (Sim 9962); Mont aux Sources, Sim 9963, and
Miss Edwards.
Grimmia dusenii Paris, Index Bryol. 2: 524. 1895, nom. nud. Guembelia dusenii Mull. Hal. ex Paris,
Index Bryol. 2: 524. 1895, nom. nud., pro synon. ORIGINAL MATERIAL: Gumbelia Duseni
C.M., in Dusen M. Carner. n° 320. - Afr.: M. Cameroon.
Grimmia duthieana Miill. Hal. ex Paris, Index Bryol. Suppl.: 173. 1900, nom. nud.
Grimmia edinensis Fergusson ex W. Bell & 1. Sadler, 1. Bot. 8: 301. 1870, nom. inval., not accepted by
authors. Grimmia subsquarrosa var. edinensis Braithw., Brit. Moss Fl. 2: 22. 1888. Grimmia
trichophylla var. edinensis (Braithw.) Dixon, Stud. Handb. Brit. Mosses ed. 3: 148. 1924.
PROTOLOGUE: [United Kingdom] Arthur's seat (Rev. 1. Fergusson 1869).
Grimmia elatior f. asperula Loeske, Biblioth. Bot. 101: 190. 1930, nom. nud.
Grimmia elatior var. physocarpa De Not., Atti Reale Univ. Genova 1: 686. 1869. PROTOLOGUE:
[Italy] circa Riva ad Sessitem[,] Carestia, in monte Musine prope Taurinum[,] DNtrs.
Grimmia elatior var. pseudofunalis Limpr., Flora 65: 204. 1882. Grimmia elatior var. subfunalis
Kindb., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 7(9): 110. 1883, nom. inval., pro synon.
PROTOLOGUE: [... ] aus den Alpen [... ], meist sterile Form Sehlesiens und des nordlichen
Europas [... ].
Grimmia elatior f. subinermis-asperula Geh., Flora 62: 475. 1879. Racomitrium asperulum Geh., Flora
62: 475. 1879, nom. inval. Grimmia elatior var. asperula Limpr., Laubm. Deutschl. 1: 772. 1889,
nom. illeg. Grimmia asperula Paris, Index Bryol. 2: 519. 1895, nom. inval., pro synon. Grimmia
elatior var. alpigena H. MOller, Ark. Bot. 26A(2): 111. 1934, nom. illeg., inel. typo var. prior.
PROTOLOGUE: [Russia] Burgasutai-Pass, in einem kleinen Rasen, mit 7, 8 und 10 [Barbula
WORLD SYNOPSIS OF GRiMMIA
103
asperifolia, Barbula ruralis and Hedwigia ciliata, respectively]; Kunowalski-Jurti, sehr sparlich
zwischen Hypnum vaucheri. - Steril.
Grimmia elatior f. sublaevis Latzel, Hedwigia 66: 140. 1926. PROTOLOGUE: [Austria] V [Villach]:
Gorlitzen 800 m.
Grimmia elatior var. submutica Holler, Ber. Naturwiss. Vereins Schwaben Neuburg 30: 237. 1893.
Grimmia elatior f. submutica (Holler) Podp., Consp. Musc. Eur.: 291. 1954. PROTOLOGUE:
[Germany] Retterschwang-Thal bei der Alpe "in der Stellen" fruchtend 1364 m.
Grimmia elongata f. epilosa Stolz ex Matousch., Ber. Naturwiss.-Med. Vereins Innsbruck 28: 86.
1903. PROTOLOGUE: [Austria. Tirol] Moserlingwand bei Windisch-Matrei, 2000m, 13.IX.l896.
Grimmia elongata f. epilosa Limpr., Laubm. Deutschl. 1: 754. 1889, nom. nud. Grimmia elongata f.
epilosa Vilh., Vestn. Knil. Ceske Spolecn. Nauk, Tr. Mat.-Prir. 1924: 40. 1924, nom. illeg., non
Matousch. 1903. PROTOLOGUE: Habitat in rupibus humidis rarissirne in Alpibus et Tatris
([Slovakia] Mala studena dolina et apud 5 pIes, 1919, Vilh.).
Grimmia elongata var. patula Bruch & Schirnp., Bryol. Europ. (fasc. 25-28) 3: 130, tab. 259B. 1845.
Grimmia patula (Bruch & Schirnp.) Schimp., Syn. Musc. Eur.: 223. 1860, comb. inval., pro
synon. Grimmia elongata f. patula (Bruch & Schirnp.) Vilh., Vestn. Kral. Ceske Spolecn. Nauk,
Tr. Mat.-Pnr. 1924: 40. 1924. PROTOLOGUE: [Austria] locis siccis Messerlingswand Alpium
tyrolensium (Homsch.).
Grimmia elongata f. propagulifera Vilh., Vestn. Kral. Ceske Spolecn. Nauk, n. Mat.-Pnr. 1924: 40.
1924. PROTOLOGUE: Habitat in Tatris ([Slovakia] Mlinica supra lacum Skok, 1900 m, 1919,
Vilh. [J. Vilhelm]).
Grimmia encalyptoides Mull. Hal. ex Watts & Whitel., Proc. Linn. Soc. New South Wales, ser. 2,27:
83. 1902, nom. nud. ORIGINAL MATERIAL: [Australia. Victoria] Vic. Dr. Sullivan.
Grimmiafilicaulis Miill. Hal., Nuovo Giom. Bot. Ital., n.s. 3: 108. 1896. PROTOLOGUE: [China] In
montibus Iu-kia-po, ad saxa, 24 Junio 1894.
Grimmiafuliginea Stirt., Ann. Scott. Nat. Hist. 17: 171. 1908. PROTOLOGUE: [United Kingdom] [... ]
near Arisaig.
Grimmia funalis f. longipila Boulay, Musc. France 1: 382. 1884. PROTOLOGUE: [not explicitly
stated, localities for G. funalis s.1.] Pyrenees, [... ], de la vallee d'Eynes (Amott), jusqu'a Cauterets
(Spruce), [... ]; Plateau-Central, Mt.-Dore (Lamy); Alpes, Chanrousse, env. du lac Coeurzet
(Ravaud), grande Chartreuse (de la Perraudiere), C. sous les formes tres diverses dans Ie massif du
Mt-Blanc (Payot).
Grimmiafunalis f. pi/osa Loeske, Laubm. Eur. Part I: 155. 1913. PROTOLOGUE: [Austria] Klippen
der Bachufer z.B. Stilluptal b. Mayrhofen, 1000 m.
104
MuNoz AND PANDO
Grimmiafunalis f. subepilosa Monk., Laubm. Eur.: 366. 1927. GrimmiafunaUs f. subpilosa Monk. ex
Podp., Consp. Musc. Eur.: 286. 1954, nom. inval., orthogr. err. PROTOLOGUE: [... ] im
Alpengebiete, in der Tatra, in den Karpathen, in GroBbritannien und im nordlichen Europa. 1m
engeren Gebiete im Riesengebirge, im Arbergebiete des Bayerischen Waldes, im AIgliu,
Schwarzwalde, in de Vogesen und bei 400 m im Bodetale im Harze von Hampe beobachtet.
Grimmia funaUs var. cernua Meyran, Ann. Soc. Bot. Lyon 39: 51. 1916, nom. nud. ORIGINAL
MATERIAL: Savoie: Aiguilles de la Gliere (Payot).
Grimmia funaUs var. epilosa Milde, Bryol. Siles.: 148. 1869. Grimmia funaUs f. epilosa (Milde)
Monk., Laubm. Eur.: 366. 1927. PROTOLOGUE: [Poland] Kesselgrube 3800' im Riesengebirge.
GrimmiafunaUs var. laxa Schimp., Syn. Musc. Eur.: 212. 1860. Grimmia spiraUs var. laxa (Schimp.)
Venturi, Rev. Bryol. 6(4): 55. 1879. PROTOLOGUE: in val di furva retro Bormio Vallis-tellinae.
Grimmia gebhardii Spreng., Mant. Prim. Fl. Hal.: 54. 1807. PROTOLOGUE: [Germany] In monte
Baldo. Gebhard.
Grimmia hartmanii f. epilosa Milde, Bryol. Siles.: 150. 1869. Dryptodon hartmanii var. epilosus
(Milde) Limpr., Laubm. Deutschl. 1: 791. 1889. PROTOLOGUE: [Poland, no locality explicitly
stated for the variety] Konigsberg, bei Kleinheide. (Sanio).- Schlesien: auf erratischen Blocken
der Ebene so bei Sagan (Everken); bei bunzlau am Teufelswehr bei Wehrau 500'; im
Schusterbusche bei Klitschdorf und in den Looswitzer Steinbriichen; bei Lowenberg am
Plagwitzer Steinberge und auf dem Probsthainer Spitzberge (Limpr.); Jauergberg; Warthaberg und
Volpersdorf (Ders.); Gorbersdorf bei Waldenburg (Milde); Geiersberg und Zobten (Schulze);
Ogulje; SchOnau; Kitzelberg bei Kauffung und Mertenstein bei Jauer (Milde); Cudowa (Ders.);
Furstenstein; sehr gemein im Riesengebirge: Petersdorf bei Warmbrunn; Hartenberg bei
Schreiberhau; Kochel- und Zackenfall-Gegend; Agnetendorf; Krumrnhubel; Briikenberg; am
langen Seiffen. Sieghilbel bei Wittichhaus im Isergebirge. (Limpr.).- Karlsbrunn und Kessel im
Gesenke.
Grimmia hartmanii f. longipila E. Bauer, Repert. Spec. Nov. Regni Veg. 50: 342. 1941, nom. nud.
Grimmia hartmanii f. robusta E. Bauer, Repert. Nov. Spec. Regni Veg. 50: 342.1941, nom. nud.
Grimmia hartmanii f. secunda Loeske, Biblioth. Bot. 101: 179. 1930. PROTOLOGUE: [Austria] [... ]
J. Murr in der Mellachschlucht (Vorarlberg, 620 m).
Grimmia hartmanii subsp. bullata Luisier, Broteria Ci. Nat. 16: 90. 1947. PROTOLOGUE: [Portugal]
Curral das Freitas: Pico Jorge, sur les rochers de basalte, 3 janv. 1945 (Nobrega).
Grimmia hartmanii var. crispa Roll, Verh. Zool.-Bot. Ges. Wien 47: 662. 1897. Dryptodon hartmanii
var. crispus (Roll) Dalla Torre & Samth., Fl. Tirol 5: 273. 1904. Grimmia hartmanii f. crispa
(Roll) Podp., Consp. Musc. Eur.: 289. 1954. PROTOLOGUE: [Austria] Varenefall bei Riva am
Gerdasee.
Grimmia hartmanii var. fastigiata Ther. & Mong., Bull. Soc. Agric. Sarthe 1899: 135. 1899.
PROTOLOGUE: [France] St-Victeur: rochers siliceux a fleur de terre, a droite et a gauche de la
WORLD SYNOPSIS OF GRiMMIA
105
route du Petit-Oisseau, en sortant du bourg. Abondante. Trouvee par M. E. Monguillon Ie 18
octobre 1893.
Grimmia hartmanii var.japonica Broth. ex Sasaoka, Bot. Mag. (Tokyo) 35(420): 272. 1921, nom. nud.
Grimmia hartmanii var. mollis Roll, Hedwigia 36: 38. 1897. Dryptodon hartmanii var. mollis (Roll)
Podp., Consp. Musc. Eur.: 291. 1954, comb. inval., pro synon. PROTOLOGUE: [Spain] Sierra
Guadarama [sic; Guadarrama], Pieta [sic; Siete] Picos [Dr. Dieck, nO] 240.
Grimmia hartmanii var. montenegrina Breidl. & Szyszyl., PI. Cemagor. Lect.: 29, tab. 2. 1888.
Dryptodon hartmanii var. montenegrinus (Breidl. & Szyszyl.) Limpr., Laubm. Deutschl. 1: 791.
1889. Dryptodon simicii Katie, in Loeske, Laubm. Eur. Part I: 139, fig. 43. 1913 ["simici"], nom.
inval., pro synon. Grimmia hartmanii var. meridionalis Loeske, Laubm. Eur. Part I: 144. 1913,
nom. illeg., inel. typo var. prior. Grimmia hartmanii subsp. montenegrina (Breidl. & Szyszyl.)
Giacom., Atti 1st. Bot. Lab. Crittog. Univ. Pavia, ser. 5,4: 223. 1947. PROTOLOGUE: Ad terram
lapidosam silvae Perucica sub monte Kom Kucki ad confmium Albaniae borealis [leg. I.
Szyszylowicz] .
Grimmia hawaiica Mull. Hal., Flora 82: 454. 1896. PROTOLOGUE: [U.S.A.] Insulae Hawaiicae, East
Maui, in monte vulcanico Haleakala: Dr. Hillebrand 1872.
Grimmia hemipolia Stirt., Ann. Scott. Nat. Hist. 11: 109. 1902. PROTOLOGUE: [United Kingdom]
[... ] from Rannoch and Broadford in Skye [J. Stirton].
Grimmia heterophyllacea Miill. Hal. ex Paris, Index Bryol. Suppl.: 173. 1900, nom. nud.
Grimmia histrioipila Mull. Hal. ex Watts & Whitel., Proc. Linn. Soc. New South Wales, ser. 2, 27: 84.
1902, nom. nud. ORIGINAL MATERIAL: [Australia. Victoria] Vic., among lichens, Mt.
Arapiles: Reader, Aug. '96.
Grimmia hornii Stirt., Scott. Naturalist (Perth) 10: 218. 1890. PROTOLOGUE: [United Kingdom] on
bare rocks in Glen Ogle (G. Hom).
Grimmia hyalinocuspidata var. mutica Cardot & Broth., Kongl. Svenska Vetenskapsakad. Handl.
63(10): 26. 1923. PROTOLOGUE: [South Georgia] Georgia austr.: Bay of Isles, Rosita Harbour
(38 p.p.).
Grimmia inaequalis Stirt., Ann. Scott. Nat. Hist. 17: 172. 1908. PROTOLOGUE: [United Kingdom]
On stone walls, Garscube near Glasgow; near Forth Bridge at Dalmeny (J. M'Andrew).
Grimmia incana Wilson, Hooker's J. Bot. Kew Gard. Misc. 9: 323. 1857, nom. nud. ORIGINAL
MATERIAL: Tibet occid. temp.; Rondu, alt. 6,000 ped. T.T. [Thomas Thompson n° 281].
Grimmia incurva f. breviseta Latzel, Hedwigia 66: 139. 1926. PROTOLOGUE: [Austria] L
[Lavantquellengebiet]: Ameringkogel.
106
MuNoz AND PANDO
Grimmia incurva f. longipila Vilh., Vestn. Kral. Ceske Spolecn. Nauk, Tf. Mat.-Ptir. 1924: 32. 1924.
PROTOLOGUE: [Slovakia] Habitat ad saxa in montibus Corconticis (Snezka sub Obfi bouda,
1450 m, 1919, Vilh. [J. VilhelmD.
Grimmia incurva var. pehrii Latzel, Hedwigia 66: 140. 1926. PROTOLOGUE: [Austria] K [Koralpe]:
Steinschober 1900 m (Gneiss).
Grimmiajacquinii Garov., Bryol. Austriac. Exc.: 25. 1840. PROTOLOGUE: [Imperio Austriaco].
Grimmiajaeggliana Giacom., Atti 1st. Bot. Lab. Crittog. Univ. Pavia, ser. 5,9: 198. 1950. Grimmia
tergestina var. jaeggliana (Giacom.) Podp., Consp. Musc. Eur.: 278. 1954. PROTOLOGUE:
[Italy] Val di Sus a (Piemonte) su rupi secche silicee presso Pradonio [... ]; tra Bussoleno e Chianoc
su rupi silicee terrose in un "Crossidietum squamigeri" [... ]; [Switzerland] Canton Ticino: rupi
calcaree scure cristalline, intercalate fra rocce silicee, soleggiatissime, preso Castione; ad Arogno
su rupi calcaree seche [... ].
Grimmia laevidens Broth., Acta Soc. Sci. Fenn. 19(12): 88. 1892. Grimmia caucasica Lindb. ex
Broth., Acta Soc. Sci. Fenn. 19(12): 88. 1892, nom. inval., pro synon. Grimmia levidens Broth. ex
Loeske, Laubm. Eur. Part I: 107. 1913, nom. inval., orthogr. err. PROTOLOGUE: [Georgia]
Cauc. major occ.: Radscha, ad fontes fluminis Dschandschachi Tskhali, ad saxa silicea regionis
alpinae superioris.
Grimmia leucophaea var. elongata Machado-Guim., Broteria, Ser. Bot. 15: 9. 1917. Grimmia
campestris var. elongata (Machado-Guim.) Cout., Musci Lusit.: 56. 1917. Grimmia laevigata var.
elongata (Machado-Guim.) Machado-Guim., Cat. Descr. Briol. Port.: 40. 1918. PROTOLOGUE:
[Portugal] Sanatorio de Manteigas. Agosto de 1916 [A.L. Machado].
Grimmia leucophaea var. epilosa Warnst., Verh. Bot. Vereins Provo Brandenburg 27: 47. 1885.
PROTOLOGUE: [Germany] Nr. [Neuruppin]: Auf erratischen Blocken sehr selten.
Grimmia leucophaea var. funaliformis Gyorffy & Peterfi, Mag. Bot. Lapok 3: 132. 1904. Grimmia
laevigata f. funaliformis (Gyorffy & Peterfi) Podp., Consp. Musc. Eur.: 276. 1954.
PROTOLOGUE: [Romania. Siebenburgen] 1m "Vale a Saca" Thale.
Grimmia leucophaea var. stenocarpa G. Roth ex J.J. Amann, Fl. Mouss. Suisse 2: l30. 1918.
PROTO LOGUE: Allemagne.
Grimmia leucophaea var. subrotunda Miill. Hal., Syn. Musc. Frond. 1: 794. 1849. PROTOLOGUE:
[Australia] ad Swan River Novae Hollandiae: J. Drummond. Vere.
Grimmia limbatula Mull. Hal., Nuovo Giorn. Bot. Ital., n.s. 3: 108. 1896. PROTOLOGUE: [Japan] ex
sylvis Yokohamae (Fr. Schaal 1879).
Grimmia minutula Miill. Hal. ex Kindb., Enum. Bryin. Exot., Suppl. 2: 107. 1893, nom. nud.
Grimmia mollis f. propagulifera lJ. Amann, Bull. Soc. Vaud. Sci. Nat. 53: 93. 1921. Hydrogrimmia
mollis f. propagulifera (lJ. Amann) Podp., Consp. Musc. Eur.: 274. 1954. PROTOLOGUE:
[Switzerland] Valais: Langefluh sur Saasfee, 2800 m (Amann) (BH. 87.6.30).
107
WORLD SYNOPSIS OF GRiMMIA
Grimmia mollis subsp. submersa Kindb., Eur. N. Amer. Bryin. 2: 218. 1898. PROTOLOGUE: Rocks
in alp. Rivulets r. Eur. Norway!; Kaurin.
Grimmia mollis var. aquatica Schimp., Syn. Musc. Eur.: 222. 1860. Grimmia mollis f. aquatica
(Schimp.) Broth., Laubm. Fennoskand.: 174. 1923. Hydrogrimmia mollis f. aquatica (Schimp.)
Loeske, Biblioth. Bot. 101: 45. 1930. PROTOLOGUE: [... ] summi Col di Stelvio [Italy] versus
Orteler alt. 8000' legi aestate 1840, et in summae Alpis Albula Rhaetiae [Austria] parte granitic a
aest. 1845 et 1855, nec non in Sierra Nevada Hispan. [Spain] ad rivul. nival. in fauce abrupta
Borreguil de s. Geronimo alt. 8500', in m. Dovre [Norway] ad lapides rivulor. Alp. StorhOe [... ]
(C. Hartman).
Grimmia muehlenbeckii f. atrata Peterfi, Mag. Bot. Lapok 2: 293. 1903. PROTOLOGUE: [Romania]
1m Szurduk-Passe.
Grimmia muehlenbeckii f. epilifera Molendo, Ber. Naturhist. Vereins Passau 10: 120. 1875. Grimmia
trichophylla f. epilifera (Molendo) Podp., Consp. Musc. Eur.: 288. 1954. PROTOLOGUE:
[Germany] Livinallongo zwischen der Omellabachmiindung und Araba auf Melafyrtuff in enger
Schlucht bei 4300' beobachtet: Mdo. [L. Molendo] (Oktober 1863).
Grimmia muehlenbeckii f. umbrosa lE. Zetterst., Ofvers. Forh. Svenska Vetensk.-Akad. 34: 70. 1878,
nom. nud. ORIGINAL MATERIAL: in latere boreali-occidentali montis Hogkullen.
Grimmia muehlenbeckii var. mutabilis Sanio, in Holler, Ber. Naturwiss. Vereins Schwaben Neuburg
30: 237. 1893. PROTOLOGUE: [Germany] Berggtindlethal unter der Pointalpe 1299 m.
Grimmia muehlenbeckii var. subsquarrosa I I Amann, Fl. Mouss.
PROTOLOGUE: [not explicitly stated].
Suisse
1: 92.
1918.
Grimmia nana Schleich., Cat. PI. Helv. ed. 4: 40. 1821, nom. nud. ORIGINAL MATERIAL:
[Helvetia].
Grimmia oblonga Wilson, Hooker's l Bot. Kew Gard. Misc. 9: 323. 1857, nom. nud. ORIGINAL
MATERIAL: Sikkim-Himalaya alpina; Jongri, alt. 12,000 ped., lD.H. [lD. Hooker n° 316].
Grimmia obtusolinealis Miill. Hal., Flora 73: 484. 1890. PROTOLOGUE: [Tanzania] Africa or. trop.,
monte Kilima-Ndscharo, 3000-4800 m: Dr. Hans Meyer 1889.
Grimmia oertzeniana Schultz, Syll. PI. Nov. 2: 134. 1828. PROTOLOGUE: [Italy] Habitat en monte
Rosa Italiae, solo micaceo. Ab Oertzen.
Grimmia orbicularis var. longipila Husn., Muscol. Gall.: 134. 1887. Grimmia orbicularis f. longipila
(Husn.) Podp., Consp. Musc. Eur.: 285. 1954. PROTOLOGUE: [France, not explicitly stated].
Grimmia orbicularis var. theriotii Corb., Rev. Bryol. 17(2): 21. 1890. PROTOLOGUE: [France] SaintPavace, pres Le Mans (Sartne): sur les murs du cimetiere autour de l'eglise; fevrier-mai 1889 (leg.
Theriot).
Grimmia orbicularis xG. pulvinata Vilh., Preslia 2: 145. 1922, nom. inval., published as a formula.
108
MuNoz AND PANDO
Grimmia ovalis f. umbrosa Vaccari ex Tosco, Webbia 28: 288. 1973, nom. nud. ORIGINAL
MATERIAL: [Italy] 3647 Capanna Gnifetti (Vacc.); 3400 Grivola (Vacc.); 3350 Cresta dell'Eco
(Vacc.); 3259 Punta di Leppe (Vacc.).- (3130 Piz Gallegione (Pfeffer».
Grimmia ovata f. epi/osa Jur. ex Vent., Musc. Trentino: 42. 1899, nom. nud. Grimmia ovalis var.
epi/osa Jur. ex Wamst., Krypt.-Fl. Brandenburg 1(2): 296. 1904. PROTOLOGUE: [Germany]
Briisenwalde auf Feldsteinmauern.
Grimmia ovalis var. heteracra I. Hagen, Kongel. Norske Vidensk. Selsk. Skr. (Trondheim) 1909(5):
27. 1909. Grimmia ovata var. heteracra (I. Hagen) Loeske, Biblioth. Bot. 101: 134. 1930, nom.
inval., pro synon. Grimmia ovalis f. heteracra (I. Hagen) C.E.O. Jensen, Skand. Bladmossfl.: 258.
1939. PROTOLOGUE: [Norway] ST. [Sor-Trondelag] Selbu, Rolset pa en vandreblok:
H.[Hagen].
Grimmia ovalis var. simplex Vondr., Bull. Soc. Amis Sci. Lett. Poznan, Ser. D, Sci. BioI. 6: 122, fig. 7.
1966. PROTOLOGUE: Iraq: Naprdan, dist. Rowanduz, 5.6.1961, leg. E. Hadac. Typus: Herb.
Vondracek, num. Hc B. 201, isotypus in BUH asservatur.
Grimmia ovata f. aquatica Matousch., Ber. Naturwiss.-Med. Vereins Innsbruck 28: 86. 1903. Grimmia
ovalis f. aquatica (Matousch.) Podp., Consp. Musc. Eur.: 279. 1954. PROTOLOGUE:
[Austria. Tirol] Fruchtend im Zwieselbache (Otztal), 23.IV.1899.
Grimmia ovata f. umbrosa Vacc., Bull. Soc. Fl. Vald6t. 9: 76.1914, nom. nud.
Grimmia ovata var. curviseta I. Hagen, Troms0 Mus. Aarsh. 21-22(1): 63. 1899. Grimmia ovalis var.
curviseta (I. Hagen) Podp., Consp. Musc. Eur.: 279. 1954. PROTOLOGUE: [Norway] in Kristians
amt beobachtet; auch auf Dovrefjeld kommt sie vor.
Grimmia ovata var. subturbinata Brid., Bryol. Univ. 1(1): 177. 1826. Grimmia ovalis var. subturbinata
(Brid.) L.1. Savicz & Smimova, Novosti Sist. Nizsh. Rast. 5: 288. 1968. PROTOLOGUE: [Russia]
In Kamtschatka habitat ubi Redowski legit.
Grimmia paramattensis Mull. Hal., Hedwigia 37: 161. 1898. PROTOLOGUE: Australia, New South
Wales, Paramatta: Whitelegge Oct. 1884: Rh. Melbourne mis. 1885.
Grimmia patens f. atrata Roll ex Loeske, Biblioth. Bot. 101: 190. 1930. PROTOLOGUE: Norwegen:
bei Bergen, am Buarbrae.
Grimmia pennsylvanica var. bestii Grout, Bryologist 7: 6. 1904. Grimmia bestii Grout, Bryologist 7: 6.
1904, nom. inval. provo PROTOLOGUE: Newfane [1902-1903].
Grimmia poUta Stirt., Ann. Scott. Nat. Hist. 16: 172. 1907. PROTOLOGUE: [United Kingdom] At
Glenfmnan, a railway station half-way between Arisaig and Fort-William [1. Stirton].
Grimmia pulvinata f. basaltica Vilh., Vestn. Kral. Ceske Spolecn. Nauk, Tf. Mat.-Prir. 1924: 28. 1924.
PROTOLOGUE: Habitat ad rupes basalticas in monte Rip in Bohemia centrali (1919, Fr. A.
Novak).
WORLD SYNOPSIS OF GRiMMIA
109
Grimmia pulvinata f. brevicapsula Boulay, Musc. France 1: 386. 1884. PROTOLOGUE: [France, not
explicitly stated].
Grimmia pulvinata f. brevipila Boulay, Musc. France 1: 386. 1884. Grimmia pulvinata var. brevipila
(Boulay) Dalla Torre & Samth., Fl. Tirol 5: 262. 1904. PROTOLOGUE: [France, not explicitly
stated].
Grimmia pulvinata var. epilosa Schimp. ex Milde, Bryol. Siles.: 146. 1869. PROTOLOGUE:
[Germany] Barwalde in der Mark. (Ruthe).
Grimmia pulvinata lusus holotricha Gyorffy, Magyar Bot. Lapok 5: 343. 1906. PROTOLOGUE:
[Hungary. Csongrad] [... ] auf Steinmauem in Kis-Zombor.
Grimmia pulvinata f. latifolia Vilh., Vestn. Knil. Cesk6 Spolecn. Nauk, Tf. Mat.-Pfir. 1924: 28. 1924.
PROTOLOGUE: [Czech Republic] Habitat ad rupes calcareas, aridas prope Radotin apud Pragam
(1919, Vilh. [1. Vilhelm]).
Grimmia pulvinata var. laxa Bott., Atti Soc. Crittog. Ital. 3: 110. 1883. PROTOLOGUE: [Italy] [... ]
venne parimente raccolta da Arcangeli nel maggio presso la foce del fiume Angitola sulle rupi.
Grimmia pulvinata var. laxa Roll, Hedwigia 56: 170. 1915, nom. illeg., non Bott. 1883. Grimmia
pulvinata f. laxa (Roll) Podp., Consp. Musc. Eur.: 285. 1954, nom. illeg., non Bott. 1883.
Grimmia pulvinata var. longepilosa Matousch., Verh. Zool.-Bot. Ges. Wien 51: 191. 1901.
PROTOLOGUE: [Italy] Obcina bei Triest (Stemeck). - D. Inseln Brazza: Zwischen Neresi und St.
Pietro (Ginzberger, 1895).
Grimmia pulvinata var. longicapsula Besch., Bull. Soc. Bot. France 12: 136. 1865, nom. nud. Grimmia
pulvinata var. longicapsula Debat, Fl. Musc.: 122. 1874. Grimmia pulvinata f. longicapsula
(Debat) Boulay, Musc. France 1: 386.1884. Grimmia montana var.longicapsula Besch., in Podp.,
Vestn. Klubu Pfir. v Prostejove 7: 19. 1905 [NOTE: Error in Wijk et al. (1962: 393)].
PROTOLOGUE: [France] Hyeres.
Grimmia pulvinata var. longipila Schimp., Syn. Musc. Eur. 1: 206. 1860. Grimmia pulvinata f.
longipila (Schimp.) Boulay, Musc. France 1: 386. 1884. PROTOLOGUE: [not explicitly stated].
Grimmia pulvinata f. minor Boulay, Musc. France 1: 386. 1884. Grimmia pulvinata var. minor
(Boulay) Dalla Torre & Samth., Fl. Tirol 5: 262. 1904. PROTOLOGUE: [France, not explicitly
stated].
Grimmia pulvinata f. nigricans Wamst., Krypt.-Fl. Brandenburg 1(2): 291. 1904. PROTOLOGUE:
[Germany] bei Neuruppin auf der Mauer des neuen Kirchhofs.
Grimmia pulvinata subsp. philibertii Kindb., Enum. N. Amer. Bryin. 2: 219. 1897. PROTOLOGUE:
[U.S.A.] Amer. r. Idaho: Leiberg.
110
MuNoz AND PANDO
Grimmia puivinata f. robusta Boulay, Musc. France 1: 386. 1884. Grimmia puivinata var. robusta
(Boulay) Dalla Torre & Sarnth., Fl. Tirol 5: 262. 1904. PROTOLOGUE: [France, not explicitly
stated].
Grimmia puivinata f. serpentinosa Vilh., Spisy Peir. Fak. Karlovy Univ.: 33. 1925. PROTOLOGUE:
[Czech Republic] [ ... ] in Bohemia occid. ad Mnichov prope Marianske Lazne.
Grimmia puivinata var. viridis Schimp., Syn. Musc. Eur.: 206. 1860. Grimmia puivinata f. viridis
(Schimp.) Loeske, Laubm. Eur. Part I: 163. 19l3. PROTOLOGUE: [France] ad St. Leonard prope
Limoges unde cl. Lamy misit.
Grimmia retracta Stirt., Scott. Naturalist (Perth) 8: 235. 1886. Grimmia hartmanii f. retracta (Stirt.)
Loeske, Laubm. Eur. Part I: 141. 1913. Grimmia hartmanii subsp. retracta (Stirt.) Dixon, Stud.
Handb. Brit. Mosses ed. 3: 151. 1924. Grimmia hartmanii var. retracta (Stirt.) Podp., Consp.
Musc. Eur.: 290. 1954. Grimmia hausmani subsp. retracta (Stirt.) Podp., Consp. Musc. Eur.: 290.
1954, nom. inval. PROTOLOGUE: [United Kingdom] Found beside Loch Tay, July, 1866 [J.
Stirton].
Grimmia retracta var. submutica Stirt., Ann. Scott. Nat. Hist. 6: 121. 1897. PROTOLOGUE: [United
Kingdom] Achnacarron, Loch Awe, 1896.
Grimmia robusta Braithw., J. Bot. 10: 196, tab. 124 fig. 3. 1872, nom. illeg., non Nees & Hornsch.
1827. Grimmia ambigua Braithw., J. Bot. 10: 196. 1872, nom. inval., pro synon. Grimmia
decipiens var. robusta Braithw., Brit. Moss Fl. 2: 23. 1888, nom. nov. Grimmia decipiens subsp.
robusta (Braithw.) Dixon, Stud. Handb. Brit. Mosses: 141. 1896. Grimmia trichophylla var.
robusta (Braithw.) A.J.E. Smith, J. Bryol. 9: 279. 1976. Grimmia brittanica A.J.E. Smith, J. Bryol.
17: 272. 1992. PROTOLOGUE: [United Kingdom. Scotland] Rocks in Alpine districts. Clova
mountains, at 2000 feet (Mr. Fergusson [1868]). Fairhead, Ireland (Dr. Moore). Cardross and
Bowling (Dr. Stirton (1864]). Ross-shire (Mr. Hunt).
Grimmia rotunda R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 416, pI. 32 fig. 22. 1895.
PROTOLOGUE: [New Zealand] Rocks, Lyttelton hills; 1880. Collected by R.B.
Grimmia rubescens Stirt., Scott. Bot. Rev. 1: 90. 1912. PROTOLOGUE: [United Kingdom. Western
Highlands, Ross-shire, near Gairloch, leg. Stirton].
Grimmia sardoa f. propaguiifera M. Fleisch., Malpighia 7: 329. 1893. PROTOLOGUE: [Italy]
Sardegna: Monte Santo, Pula, leg. Moris.
Grimmia serruiata F. Weber & D. Mohr, Index Mus. PI. Crypt.: 2. 1803, nom. nud.
Grimmia stenophylla Mlill. Hal., Hedwigia 37: 158. 1898. PROTOLOGUE: [Australia] Tasmania, ad
rupes on the Sugar loaf, Green Ponts, Majo 1892: W.A. Weymouth in Hb. Brotheri, qui misit 1893
sub G. trichophylla Grev.
Grimmia stevensii R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 418, pI. 34 fig. 30. 1895.
PROTOLOGUE: [New Zealand] Mount Torlesse; January, 1886. Collected by R.B.
WORLD SYNOPSIS OF GRiMMIA
111
Grimmia stirlingii Mull. Hal., Hedwigia 37: 163. 1898. Schistidium stirlingii (Mull. Hal.) Ochyra,
Fragm. Florist. Geobot. 43: 107. 1998. PROTOLOGUE: Australia, Victoria, Omeo alt. 2500-3000
pedum, ad rupes 1. Stirling 1883 in Rh. Melbourne.
Grimmia subaqui/a Stirt., Ann. Scott. Nat. Hist. 18: 243. 1909. PROTOLOGUE: [United Kingdom] At
Loch-nan-Uamb, several miles to the east of Arisaig and near Prince Charlie's Cave; on stones
covered with earth, by the Rev. 1.A. M'Caskill ofOnich, September 1908.
Grimmia subcallosa Miill. Hal., Hedwigia 37: 161. 1898. PROTOLOGUE: Australia, Victoria, Mt.
Cole Pyrenaeorum: D. Sullivan 1883 in Rh. Melbourne.
Grimmia subelongata Gopp. & Menge, Ber. Bekanntm. Verh. Konigl. Preuss. Akad. Wiss. Berlin 18:
459. 1853, nom. nud.
Grimmia subfusca Wilson, Hooker's 1. Bot. Kew Gard. Misc. 9: 323. 1857. PROTOLOGUE: [India]
Khasia temperata; Kollong, alt. 5,000 ped., 1.D.H. [J.D. Hooker n° 296].
Grimmia subsulcata f. dominiana Vilh., Vestn. Knil. Ceske Spoleen. Nauk, Tf. Mat.-Pili. 1924: 24.
1924. Grimmia alpestris f. dominiana (Vilh.) Podp., Consp. Musc. Eur.: 282. 1954.
PROTOLOGUE: [Poland] Habitat ad rupes granitic as supra lacum "Skalnate pleso" (2000 m) sub
montem Lornnicky 8tit in Tatris (leg. 1919, Domin).
Grimmia subtergestina Miill. Hal., Nuovo Giom. Bot. Ital., n.s. 3: 109.1896. PROTOLOGUE: [China]
Provo Kansu, ad flumen Bardun prope ostium fluminis Solomo, Majo 1886: G.N. Potanin legit
sterilem.
Grimmia tergestina f. epilosa Limpr., Laubm. Deutschl. 1: 740. 1889. PROTOLOGUE: [Austria] [... ]
sammelte H. Gander die d' Pflanze an senkrechten Kalkwanden uber Kalchstein 1750 m bei
Innervillgraten in Tirol.
Grimmia tergestina var. carpatica Vanek ex Podp., Consp. Musc. Eur.: 277. 1954, nom. nud.
Grimmia tergestina subsp. tergestinoides Culm., Rev. Bryol. 38: 75. 1911, nom. alt. Grimmia
tergestina var. tergestinoides Culm., Rev. Bryol. 38: 75. 1911, nom. alt. PROTOLOGUE:
[Switzerland] La Chaux prope Sainte-Croix, montis Jura helvetici, c. fructu unico, altitudine 1080
m. legit Ch. Meylan. (Sur calcaire aquitanien).
Grimmia torquata var. pili/era Scheutz, Ofvers. Forh. Kongl. Svenska Vetensk.-Akad. 27: 94. 1870,
nom. nud. ORIGINAL MATERIAL: [Sweden] Skurugata.
Grimmia torquata-andreaeoides Loeske, in J.J. Amann, Bull. Soc. Vaud. Sci. Nat. 54: 45. 1921, nom.
inval., published as a formula.
Grimmia tortilis Muhl., Cat. PI. Amer. Sept.: 98. 1813, nom. nud.
Grimmia trichophylla f. epi/osa Warnst., Verh. Bot. Vereins Provo Brandenburg 39: 31. 1897.
Grimmia trichophylla var. epi/osa (Warnst.) Warnst., Krypt.-Fl. Brandenburg 1(2): 299. 1904.
PROTOLOGUE: [Germany. Mark Brandenburg] An erratischen Blocken an der Strasse nach Gr.
SchOnebeck.
112
MuNoz AND PANDO
Grimmia trichophylla var. gemmipara Husn., Fl. Mousses Nord-Ouest: 93. 1873. PROTOLOGUE:
[France] Segrie-Fontaine [Orne].
Grimmia trichophylla var. longipila Warnst., Krypt.-Fl. Brandenburg 1(2): 299. 1904. Grimmia
trichophylla f. longipila (Warnst.) MOItle, Laum. Eur.: 367. 1927. PROTOLOGUE: [Poland.
Postdam] Ostprignitz: Mansfeld, erratischer Block auf der Heide (Jaap!).
Grimmia trichophylla var. lusitanica Schimp., Syn. Musc. Eur. ed. 2: 257. ＱＸＷｾＮ＠
PROTOLOGUE:
[Portugal] in rup. graniticis m. Foya ad Monchique Aigarviae (C.H. Solms).
Grimmia trichophylla var. mutica Grav. ex Delogne, Ann. Soc. BeIge Microscop. 9: 178. 1885.
PROTOLOGUE: [Belgium] Profondeville (Grav.).
Grimmia trichophylla var. nigra R. Br. bis, Trans. & Proc. New Zealand Inst. 27: 415, pI. 31 fig. 19.
1895. PROTOLOGUE: [New Zealand] Rocks in the gorge of the River Hurunui; March, 1893.
Collected by RB.
Grimmia trichophylla var. submutica Boulay, Musc. France 1: 379. 1884. Grimmia trichophylla f.
submutica (Boulay) Corb., Mem. Soc. Sci. Nat. Cherbourg 26: 256. 1889. PROTOLOGUE:
[France] gres desagreges, frais dans l'Esterel (B.).
Grimmia trichophylla f. vulgaris Monk., Laubm. Eur.: 367. 1927. PROTOLOGUE: ... der Hugel- und
Bergregion durch Europa [... ] deutschen Mittelgebirgen [... ] Norddeutschen Ebene.
Grimmia triformis var. brevifolia Chal. ex J.J. Amann, Bull. Soc. Vaud. Sci. Nat. 55: 131. 1923, nom.
nud. ORIGINAL MATERIAL: [Switzerland] Bern: Finsteraarhorngipfel, 4275 m. Frey.
Grimmia truncatoapocarpa Mull. Hal., Hedwigia 37: 163. 1898. Schistidium truncatoapocarpum
(Mull. Hal.) Ochyra, Fragm. Florist. Geobot. 43: 107. 1998. PROTOLOGUE: Australia
subtropica, New South Wales, Paramatta: Whitelegge. Octobri 1884. Rh. Melbourne misit 1885.
Grimmia uncinata Kaulf., in Sturm, Deutschl. FI. Abtheilung II, Cryptogamie 4(15): 13. 1816.
PROTOLOGUE: [Austria] Auf der Seethaler Alpe in Stenermark fand ich dieses Moos fehr haufig
im August mit reisen Fruchten in Steinrillen.
Grimmia undulata Stirt., Scott. Bot. Rev. 1: 91. 1912. PROTOLOGUE: [United Kingdom. Scotland:
Rossshire] On the red rock in several places near Gairloch [J. Stirton].
Grimmia unicolor f. piligera Molendo, in Lorentz, Moosstud.: 94. 1864, nom. nud. Grimmia unicolor
var. piligera (Molendo) Grav., Rev. Bryoi. 10: 24. 1883, comb. invai.
Grimmia versabilis R Br. bis, Trans. & Proc. New Zealand Inst. 27: 415, pI. 32 fig. 20. 1895.
PROTOLOGUE: [New Zealand] summit of Port Lyttelton hills; 1880. Collected by R.B.
Grimmia wrightii R Br. bis, Trans. & Proc. New Zealand Inst. 27: 413, pI. 30 fig. 12. 1895, nom.
illeg., non Austin 1875. Schistidium wrightii Ochyra, Fragm. Florist. Geobot. 43: 107. 1998 nom.
nov. PROTOLOGUE: Limestone rocks, Broken River, Canterbury; November, 1889. Collected by
R.B.
WORLD SYNOPSIS OF GRIMMIA
113
Grimmia zahlbruckneri Garov., Bryol. Austriac. Exc.: 26. 1840. PROTOLOGUE: [Imperio Austriaco].
Guembelia arctica Mull. Hal., Zweite Deut. Nordpolarf. 2(1): 73. 1873. Grimmia arctica (Mull. Hal.)
A. Jaeger, Ber. Tiitigk. St. Gallischen Naturwiss. Ges. 1872-73: 71 (Gen. Sp. Musc. 1: 353). 1874.
Racomitrium hypnoides var. arcticum (Mull. Hal.) C.E.O. Jensen, Meddel. Gmnland 3: 345. 1887.
Grimmia caespiticia subsp. arctica (MUll. Hal.) Kindb., Eur. N. Amer. Bryin. 2: 220. 1898.
PROTOLOGUE: [Greenland] Kaiser-Franz-Josephs-Fjord inter 73-74° lat. bor., unde cespes
singularis relatus est.
Guembelia caucasica MUll. Hal., Bot. Zeitung (Berlin) 14: 421. 1856. Grimmia caucasica (MUll. Hal.)
A. Jaeger, Ber. Tiitigk. St. Gallischen Naturwiss. Ges. 1872-73: 72 (Gen. Sp. Musc. 1: 354). 1874.
PROTOLOGUE: Caucasus: Relanger sub No.9. Rh. Kunzeano communicavit.
Guembelia erythraea MUll. Hal., in Brizi, Malpighia 7: 296. 1893. Grimmia erythraea (MUll. Hal.)
Cufod. Oesterr. Bot. Z. 98: 227. 1951. PROTOLOGUE: [Ethiopia] Abyssinia, prope Keren.
Racomitrium patens subsp. subs imp lex Kindb., Boll. Soc. Bot. Ital. 1896( 1): 22. 1896. Grimmia patens
subsp. subsimplex (Kindb.) Kindb., Eur. N. Amer. Bryin. 2: 225. 1898. Grimmia patens subsp.
simplex Kindb. ex Paris, Index Bryol. Suppl.: 294. 1900, nom. inval., orthogr. err.
PROTOLOGUE: [Switzerland] Canton d'Uri [... ] Goeschenen [N.C. Kindberg & 1. Roll, 1895].
Racomitrium patens var. crassifolium Torka, AUg. Bot. Z. Syst. 9(7/8): 146. 1903. Grimmia patens f.
crassifolia (Torka) Podp., Consp. Musc. Eur.: 292. 1954. PROTOLOGUE: [Germany] [... ] am
27.August 1902 auf einem Steine bei Schwiebus [Saxonia] gefunden worden ist.
Schistidium teretinerve f. lanceo/atum Podp., Veroff. Geobot. Inst. Rubel Ziirich 1: 245. 1924.
Grimmia teretinervis f. lanceo/ata (Podp.) Podp., Consp. Musc. Eur.: 281. 1954. PROTOLOGUE:
[Switzerland] [... ] Rande der Buffalorawiese.
Trichostomum pulvinatum var. elongatum Schultz, Syll. PI. Nov. 2: 142. 1828. Grimmia pulvinata var.
elongata (Schultz) Fiedl., Syn. Laubm. Mecklenb.: 82. 1844. PROTOLOGUE: [Germany] Habitat
in saxis granitosis Fl. Stargard.
114
MuNoz AND PANDO
LITERATURE CITED
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Bednarek-Ochyra, H., J. Munoz & R. Ochyra. 1992. The identity of Grimmia pyrenaica (Musci,
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Blom, H. H. 1996. A revision of the Schistidium apocarpum complex in Norway and Sweden.
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Bremer, B. 1980 [1981]. A taxonomic revision of Schistidium (Grimmiaceae, Bryophyta) 2.
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Bridel, S. E. 1806. Muscologia Recentiorum Supplementum 1: [i]-viii, [1]-271. Gothae. Carolum
Guil. Ettingerum.
Brown, R. 1903. On the musci of the calcareous districts of New Zealand, with descriptions of new
species. Trans. & Proc. New Zealand Inst. 35: 323-342, pI. 35-40.
Cao, T. & S. P. Churchill. 1995. New synonyms in Grimmia and Schistidium (Bryopsida:
Grirnmiaceae). Nova Hedwigia 60: 505-513.
Cao, T. & D. H. Vitt. 1986. A taxonomic revision and phylogenetic analysis of Grimmia and
Schistidium (Bryopsida; Grimmiaceae) in China. J. Hattori Bot. Lab. 61: 123-247.
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Grirnmiaceae (Musci). Advances Cladist. 1: 127-144.
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Gard. 50: [i-vi], 1-87.
Crosby, M. R. & R. E. Magill. 1997. Index of Mosses 1993-1995. Monogr. Syst. Bot. Missouri Bot.
Gard. 62: [i-vi], 1-106.
Crosby, M. R., R. E. Magill & c. R. Bauer. 1992. Index of Mosses 1963-1989. Monogr. Syst. Bot.
Missouri Bot. Gard. 42: [i-vi], 1-646.
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WORLD SYNOPSIS OF GRIMMIA
115
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116
MuNoz AND PANDO
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Regnum Veg. 65: [i]-xii, 1-922.
WORLD SYNOPSIS OF GRIMMIA
117
INDEX OF SCIENTIFIC NAMES
Accepted taxa are in bold. An asterisk (*) indicates lectotypification designated in this paper.
Afzelia
pulvinata Ehrh., nom. inval., 66
Amphidium
tortuosum (Hornsch.) Cufodontis, 85
Anictangium
pulvinatum (Hedw.) D. Mohr, 16
Anodon Rabenh., 5
pulvinatus (Hedw.) Rabenh., 16
ventricosus Rabenh., 16
Anoectangium
pulvinatum (Hedw.) Rohl., 16
Arctoa
anderssonii Wich., 85
Blindia
acuta (Hedw.) Bruch & Schimp., 85
Brachydontium
olympicum (E. Britton) T.T. McIntosh &
J.R. Spence, 85
Bryum
brevicaule Vill., nom. inval., 60
orbiculare pulvinatum, hirsutie canescens,
capsulis immersis Dill., nom. inval., 66
orbiculare pulvinatum, hirsutie canescens,
var. capsulis erectis praedita Dill., nom.
inval.,39
ovale Hoffin., nom. inval., 60
patens Dicks., nom. inval., 71
patens Hedw., 71
pulvinatum L., nom. inval., 66
trichodes hirsutie canescens, capitulis
subrotundis erectis, in pediculis
brevissirnis Dill., nom. inval., 39
trichodes hirsutie canescens, capitulis
subrotundis reflexis in perbrevibus
pediculis Dill., nom. inval., 66
Campylopus
caespiticius Brid., 22
contortus (Wahlenb.) Brid., 36
curvifolius Brid., 86*
laevigatus Brid., 39
ovalis (Hedw.) Wahlenb., 60
var. hyrsutus Myrin, nom. illeg., 44
var. ovatus (F. Weber & D. Mohr)
Wahlenb., 60
var. politus Myrin, 97
patens (Hedw.) Brid., 71
pulvinatus (Hedw.) Brid., 67
var. alpestris (F. Weber & D. Mohr) Brid.,
15
var. tenuis Wahlenb., 97
Campylostelium
pitardii (Corb.) Maier, 63
Coscinodon
arctolimnius (Steere) Steere, 85
arsenei Ther., 86
australis Dixon & Sainsbury, 20
bolivianus Broth., 86*
cribrosus (Hedw.) Spruce, 86
humilis Horik. & Nog., nom. inval., 86
humilis Milde, 86
japonicus Sakurai, 37
trinervis (R.S. Williams) Broth., 82
Dicranum
africanum (Hedw.) F. Weber & D. Mohr,
67
arcuatum Schleich., nom. nud., 97
contortum Wahlenb., 36
cribrosum Brid., 67
ovale Hedw. 1792, nom. inval., 60
ovale Hedw. 1801,60
ovatum Brid., nom. illeg., 60
ovatum Sw., nom. inval., 60
patens (Hedw.) Sm., 71
phascoidum P. Beauv., 24
pulvinatum (Hedw.) Lag., D. Garcia &
Clemente, 67
var. affme Schleich. ex Brid., nom. inval.,
67
var. africanum (Hedw.) Schleich., 67
var. argentatum Turner, 97
var. muticum Chevall., 97
pulvinatum Sw., nom. inval., 66
Didymodon
percarinatus Dixon & Sakurai in Sakurai,
61
rigidulus Hedw., 86
Dryptodon Brid., 5
anomalus (Hampe) Loeske, 17
118
MuNoz AND PANDO
apiculatus (Hornsch.) Hartm., 31
atratus (Hornsch.) Hartm., 19
brevifolius Brid., 39
brotheri (Broth.) Warnst., 35
caespiticius (Brid.) Brid., 22
var. muticus Brid., 97
campestris (Hook.) Brid., 39
contortus (Wahlenb.) Brid., 36
cribrosus (Brid.) Brid., 67
curvifolius (Brid.) Brid., 86
decipiens (Schultz) Loeske, 24
donnianus (Sm.) Hartm., 25
elatior (Bals.-Criv. & De Not.) Loeske, 26
ellipticus (Funck) Hartm., nom. illeg., 60
elongatus (Kaulf.) Hartm., 28
erostris Hartm., nom. illeg., 25
funalis (Schwagr.) Brid., 30
var. incurvatus Hartm. ex H. Moller, nom.
inval.,26
var. incurvus (Hoppe & Hornsch.) Hartm.,
26
var. longipilus Brid., 24*
var. robustus De Not. ex H. Moller, nom.
inval.,26
grimmioides Brid., nom. inval., 24
hartmanii (Schimp.) Limpr., 34
var. alatus Katic, 97
var. crispus (Roll) Dalla Torre & Sarnth.,
104
var. epilosus (Milde) Limpr., 104
var. giganteus Jeanb. & Renauld ex
Warnst., 97
var. moIIis (Roll) Podp., comb. inval., 105
var. montenegrinus (Breidl. & Szyszyl.)
Limpr., 105
f. propaguliferus (Milde) Limpr., 35
f. rivularis Loeske, 97
incurvus (Hoppe & Hornsch.) Brid., 26
leucophaeus (Grev.) Brid., 39
lisae (De Not.) Loeske, 41
longipilus Brid., nom. inval., 24
muehlenbeckii (Schimp.) Loeske, 55
obtusus Brid., nom. illeg., 67
obtusus Hartm., 67
ovalis (Hedw.) Hartm. ex H. Moller, nom.
inval.,60
ovatus (F. Weber & D. Mohr) Hartm., nom.
illeg.,60
ovatus (Hedw.) Brid., 60
var. affme (Hornsch.) Hartm., 43
var. denticulatus Brid., 97
var. elongatus Brid., 98
var. nigricans (Lam. & DC.) Brid., nom.
inval.,60
var. obliquus (Hornsch.) Hartm., 43
var. patens (Hornsch.) Hartm., 43
patens (Hedw.) Brid., 72
var. propaguliferus J.J. Amann, 72*
f. cataractarum Vilh., 98
f. humilis Vilh., 98
f. piliferus Vilh., 98
f. robustus Vilh., 98
pulvinatus (Hedw.) Brid., 67
var. alpestris (F. Weber & D. Mohr) Brid.,
15
var. argentatus (Turner) Brid., 97
var. canus Hartm., 98
var. crinitus Breb., nom. nud., 67
sardous (Miill. Hal.) Loeske, 42
schultzii Brid., nom. illeg., 24
sciuroides (Nees & Hornsch.) Hartm., 44
simicii Katic in Loeske, nom. inval., 105
spiralis (Hook.) Brid., 30
torquatus (Drumm.) Brid., 76
trichophyllus (Grev.) Brid., 77
unicolor (Hook.) Hartm., 83
Encalypta
pulvinata Sibth., nom. inval., 66
Fissidens
patens (Hedw.) Wahlenb., 71
pulvinatus Hedw., 66
var. africanus Hedw., 67
var. communis Hedw., nom. illeg., 66
pulvinatus Timm, nom. inval., 66
Grimmia Hedw., 5
subgen. Hydrogrimmia I. Hagen, 5
subgen. Hygrogrimmia Vilh., nom. inval.,
orthogr. err., 5
sect. Hydrogrimmia (I. Hagen) Nyholm, 5
subsect. Hydrogrimmia (I. Hagen) G.N.
Jones in Grout, 5
abscondita Cardot, 88
abyssinica (Mull. Hal.) Mitt., 32
abyssinica Bruch & Schimp., nom. nud., 32
aethiopica Miill. Hal., 93*
affinis Hornsch., 31
WORLD SYNOPSIS OF GRiMMIA
affinis Homsch. ex Brid., nom. inval., 77
affinis Homsch., nom. illeg., 43
var. elongata Nees & Homsch. ex
Huebener, nom. illeg., 44
var. macrocarpa Nees & Homsch., 44*
var. ramosissima Nees & Homsch., 44
africana (Hedw.) Am., 67
afroincurva Broth., 33*
afro-ovata Broth. & Ther., 50
ahmadiana Nog., 71
akaisialpina Takaki, 51
albicans Spreng. ex H. Moller, nom. inval.,
98
albida Spreng., nom. illeg., 98
allionii Broth., 49
alpestris (F. Weber & D. Mohr) Schleich.,
15
subsp. bifrons (De Not.) Podp., nom.
inval.,99
subsp. subsulcata (Limpr.) Kindb., 98
subsp. ungeri (Jur.) Kindb., 82
var. brevipila Bruch ex Mont., nom.
inval., 19
var. breviseta J.J. Amann, 15
var. caespiticia (Brid.) G. N. Jones in
Grout, 22
var. euaIpestris Loeske, nom. inval., 15
var. holzingeri (Cardot & Ther.) G. N.
Jones in Grout, 15
var. hybrida (Chal.) Chal., 73
var. manniae (Mull. Hal.) G. N. Jones in
Grout, 22
var. microstoma Bruch & Schimp., 98
var. mutica (Brid.) De Not., 97
var. sessitana (De Not.) I. Hagen, 73
var. stomata Loeske, nom. illeg., 73
var. subimberbis (Lindb.) Berggr., 22
var. subsulcata (Limpr.) Broth., 98
var. ungeri (Jur.) Husn., 82
f. dominiana (Vilh.) Podp., 111
f. hybrida Chal., 73
f. longipila Podp., 98
f. sessitana (De Not.) Loeske, 73
f. subsulcata (Limpr.) Loeske, 98
alpestris Schleich., nom. nud., 15
alpina Kindb., 16
altemuata (Milll. Hal. & Kindb.) Kindb., 18
ambigua Braithw., nom. inval., 110
119
amblyophylla Milll. Hal., 92
americana E.B. Bartram, 15
anceps Boulay, 73
ancistrodes Durieu & Mont., 42*
andreaeacea Milll. Hal., 29
angustissima P. de la Varde, 88
anodon Bruch & Schimp., 16
var. anomala E.B. Bartram, 17
var. brevipila C.E.O. Jensen in Dusen,
nom. nud., 98
var. moravica Podp., 99
var. mutica Broth. ex Paris, nom. nud., 66
var. sinaitica Renauld & Cardot, 16*
var. taeniophyllum Mikut. ex Loeske,
nom. inval., 99
f. anomala (E.B. Bartram) G.N. Jones in
Grout, 17
f. epilosa Monk., 16
anomala Hampe ex Schimp., 17
antarctici Cardot, 92
var. percompacta E.B. Bartram, 92
var. pilifera E.B. Bartram, 92
antillarum Ther., 51
antipodum Milll. Hal., 88
apiculata Homsch., nom. illeg., 31
subsp. holleri (Molendo) J.J. Amann, 32
var. holleri (Molendo) Husn., 32
var. mutica Schultz, 99
var. obtusa Schimp., 99
f. holleri (Molendo) Loeske, 32
apiculafa Homsch., nom. nud., 31
apocarpa
subsp. pachyneurula (Milll. Hal. &
Kindb.) Kindb., 18
subsp. sphaerica Boulay, 16
var. pulvinata (Hedw.) G.N. Jones in
Grout, 16
var. sUblaevipila Cardot in Broeck, 81
apocaula Hoffm. ex DC. in Lam. & DC., 64
apophysata A. Jaeger, nom. nud., 37
apophysata Gangulee, 37
arachnoidea Dusen, nom. nud., 70
arctica (Milll. Hal.) A. Jaeger, 113
arctolirnnia Steere, 85
arctophila Kindb., 54
subsp.labradorica Kindb., 48
arcuata De Not., nom. illeg., 72
arcuatifolia Kindb., 18
120
MuNoz AND PANDO
arenaria Hampe, 19
argentea R. Br. bis, 64
argyrotricha Mull. Hal., 99
arizonae Renauld & Cardot, 62
f. propagulifera (E.B. Bartram) G.N.
Jones in Grout, 62
arsenei Cardot, 62
arvernica H. Philib.; 99
aspera Milll. Hal., 27
asperitricha Dixon & Sainsbury, 74
asperula Paris, nom. inval., 102
atrata Miel. ex Homsch., 19
var. minor Drumm., 83
atricha Mull. Hal. & Kindb., 92
atricha Mull. Hal., nom. illeg., 94
atrichoides Paris, nom. illeg., 94
atroviridis Cardot, 62
attenuata (Mull. Hal. & Kindb.) Mull. Hal.
& Kindb, nom: inval., 18
auresia Besch., 68
auresiaca Besch. ex Kindb., nom. inval., 68
austinii Kindb., 58*
australasica Mitt. ex Watts & Whitel., nom.
inval.,94
australis (Dixon & Sainsbury) J. Munoz &
Ochyra,20
austroapocarpa Milll. Hal. ex Watts. &
Whitel., nom. nud., 88
austrofunalis Broth., nom. illeg., 38
austrofunalis Mull. Hal., 20
austropulvinata Mull. Hal., 99
azorica Renauld & Cardot, 42
barkeri W. E. Nicholson & G. Roth, nom.
inval.,75
basaltica Mitt. ex Hook.f., 68*
bauri Milll. Hal. ex H. Moller, nom. inval.,
30
bellii R. Br. bis, 40*
benoistii Ther., 21
bemoullii Milll. Hal., 61
bestii Grout, nom. inval., 108
bicolor Herzog, 21
bifrons De Not., 99
biplicata Broth., nom. nud., 56
bogotensis (Hampe) A. Jaeger, 45
bohemica Schkuhr ex Steud., nom. inval.,
25
boliviana (Broth.) FJ. Herm., 86
bomeensis Dixon & Herzog, 35
bossierei Cardot & Ther., 38
brachydictyon (Cardot) Deguchi, 35
brachyodon Austin, 82
brachyphylla Cardot, 86
brandegei Austin, 64
breviexserta Milll. Hal., 47
brevipes Milll. Hal. ex Paris, nom. inval.,
47
brevirostris R.S. Williams, 34
brittanica A.J.E. Smith, 110
brittoniae R.S. Williams, 31
brotheri Lindb. ex Broth., 35*
brownii Paris, nom nov., 79
caespiticia (Brid.) Jur., 22
subsp. arctica (Mull. Hal.) Kindb., 113
var. bommuellerorum Schiffu., 99
var. subimberbis (Lindb.) Limpr., 22
f. subimberbis (Lindb.) Podp., 22
califomica Sull., 42
subsp. watsonii (Lesq. & James) Kindb.,
78
callosa Milll. Hal. & Hampe, 68
calotricha Milli. Hal., 40
calvescens Kindb., nom. nov., 30
calycina Herzog, 88
calyculata Milll. Hal., 46*
camonia Rota ex De Not., 99
campbelliae Milli. Hal., 69*
campestris Burchell ex Hook., 39
subsp. sarcocalyx (Kindb.) Kindb., 40
var. elongata (Machado-Guim.) Cout.,
106
var. latifolia (Limpr.) Paris, 40
var. lutescens Monk., 99
f. latifolia (Limpr.) Monk., 40
f. longirostrata Vilh., 99
campylopoda (Laz.) K. Saito, 63
campylotricha Milll. Hal., 100
canadensis H. Winter, nom. illeg., 83
canadensis Kindb., 42
canariensis Schimp. ex Paris, nom. nud., 42
canescens Schleich. ex Spreng., 100
canescens Schleich., nom nud., 60
canifolia Lindb. & Amell ex Broth., nom.
inval.,47
capillata De Not., 23
var. calva J.J. Amann, 23*
WORLD SYNOPSIS OF GRIMMIA
cardotii x G. anodon Culm., nom. inval.,
100
cardotii Herib., nom. inval., 65
var. gymnostoma 1. Froehl., 100
catalinensis E.B. Bartram, 50
var. mutica E.B. Bartram, 50
caucasica (Mull. Hal.) A. Jaeger, 113
caucasica Lindb. ex Broth., nom. inval.,
106
cavifolia Lindb. & Arnell, 47
celata Cardot, 88*
cernua Nees & Hornsch., 100
var. spiralis (Hook.) Nees & Hornsch., 30
f. atra Nees & Hornsch., 76*
cespiticia Jur. ex Dalla Torre & Sarnth.,
nom. inval., 22
chenii S.H. Lin, nom. nov., 92
chilensis Ther., 56
chloroblasta Kindb., 94
chrysoneura MUll. Hal., 92
chubutensis Cardot & Broth., 88
churchilliana Crum, 86
cinerea TheT., 50
coarctata MUll. Hal., 100
cockaynei R. Br. bis, 79*
cognata Cardot & Ther., 27*
coloradensis Austin, 88*
columbica De Not., 45
commutata Huebener, nom. illeg., 60
var. brevipila Warnst., 100
vaT. gibba Boulay, 100
vaT. imberbis Lindb. ex H. Moller, nom.
inval.,60
var. maroccana Meyl., nom. nud., 100
vaT. microcarpa Lesq., nom. nud., 100
var. mutica 1.E. Zetterst. ex H. Moller,
nom. inval., 60
vaT. ventricosa Lesq., nom. nud., 60
f. basaltica Vilh., 100
f. brevipila (Warnst.) Vilh., 100
f. brevipila Broth., nom. nud., 48
f. epilifera J.E. Zetterst., 60
f. minor Laz., 100
f. rivularis Loeske, 101
f. rivularis Loeske, nom. nud., 101
f. subepilosa Braun-Blanq., nom. nud.,
100
compacta Kindb., nom. nud., 101
121
compactula MUll. Hal., 101
concinodontoides Kindb., nom. nud., 94
consobrina Kunze ex MUll. Hal., 78
contorta (Wahlenb.) Am., 36
var. hagenii (Kaurin) Paris, 36
var. tatrensis (Chal.) Chal., 36
f. brevifolia Chal., nom. alt., 36
f. incurva (Schwagr.) Molendo, 36
f. longifolia Chal., nom. inval., 36
f. tatrensis Chal., nom. alt., 36
f. typica Chal., nom. alt., nom. inval., 36
coscinodontea Kindb. ex Paris, nom. inval.,
94
coscinodontoides Kindb., nom. inval., 94
cossonii Besch., 61
crassa MUll. Hal. ex A. Jaeger, nom. nud.,
39
crassifolia Lindb. ex Broth., 65
vaT. cucullata Baumgartner, 101
crassiretis Cardot & Broth., 21
cratericola Sakurai & Takaki, 85
cribrodontia Herzog, 93
crinita x G. leucophaea Philib., nom. inval.,
65
crinita Brid., 23
var. brevis Boulay, 101
var. calabra Zodda, 101
vaT. calva (J.1. Amann) J.1. Amann in
Loeske,23
vaT. capillata (De Not.) De Not., 23
var. elongata Bruch & Schimp., 101
vaT. laevis Boulay ex Pit., nom. nud., 101
vaT. libani Bizot, 24
f. brevis (Boulay) Podp., 101
f. compacta Besch., 10 1
crispata MUll. Hal. & Hampe ex Mitt., nom.
inval.,78
crispatula MUll. Hal. & Hampe, 78
cucullata D.M. Hend., 75
cucullata 1.S. Lou & P.C. Wu, nom. illeg.,
96
cupularis Mull. Hal., 89
curvata (Brid.) De Sloover, 72
curvifolia Lindb. ex Kindb., nom. nud., 101
curviseta Bouman, 24
curvula Bruch & Schimp., nom. illeg., 19
cyathocarpa (Hampe) Mitt., 91
cygnicolla Taylor, 67
122
MuNoz AND PANDO
cylindrica Nees & Homsch., 44*
cylindropyxis Miill. Hal., 87*
decalvata Cardot, 29
decipiens (Schultz) Lindb., 24
subsp. robusta (Braithw.) Dixon, 110
var. hendersonii (Renauld & Cardot)
Sayre, 68
var. homomalla Bryhn, 101
var. robusta Braithw., 110
f. falcata Luisier, 101
f. homomalla (Bryhn) I. Hagen, 101
f. major Luisier, 102
'decurrentipilis Ther., 70
defoliata Miill. Hal., 87
densa Kindb., 79
depilata Kindb., nom. illeg., 79
depressa Mull. Hal., 87
var. terrestris Miill. Hal., 87
dicksonii (Dusen) Dusen, nom. nov., 80
dimorphula Miill. Hal., 47
diversifolia Kindb., 89
donatii Herzog & Ther., 89
donii Sm. ex Lindb., nom. illeg., 25
donniana Sm., 25
subsp. arenaria (Hampe) Dixon, 19
subsp. bifrons (De Not.) Wijk & Margad.,
99
subsp. eudonniana Giacom., nom. inval.,
25
var. alpestris (F. Weber & D. Mohr)
Hampe, 15
var. arenaria (Hampe) Loeske, nom. illeg.,
19
var. bitrons (De Not.) Podp., nom. inval.,
99
var. bohemica Schkuhr ex Brid., 25
var. bohemica Schkuhr ex Steud., nom.
nud., 25
var. breviseta Breidl. ex Loeske, 82
var. curviseta Lesq. & James, 102
var. curvula Spruce, 19
var. elongata (Nees & Homsch.) Wilson,
26
var. eudonniana Loeske, nom. inval., 25
var. holzingeri (Cardot & Ther.) Wijk &
Margad.,15
var. manniae (Miill. Hal.) Wijk &
Margad.,22
var. obtusa Steud., 25
var. sudetic a Huebener, nom. illeg., 15
var. triforrnis (Carestia & De Not.)
Loeske,82
f. antarctica Kuc, 74
f. breviseta Vilh., nom. illeg., 102
f. cylindrica Podp., 26*
f. elongata (Nees & Homsch.) Loeske, 26
f. longiseta Vilh., nom. illeg., 26
f.longiuma Vilh., 102
f. sudetica (Schkuhr) Loeske, nom. illeg.,
25
f. sudetic a Chal., 102
[. vulgaris Chal., 26*
donnii Gray, nom. illeg., 25
domajii Hohn., 16*
drakenbergensis Sim, 102
dusenii Paris, nom. nud., 102
duthieana Miill. Hal. ex Paris, nom. nud.,
102
ecklonii Spreng., 67*
edentula Hy, 17*
edinensis Fergusson ex W. Bell & J. Sadler,
nom. inval., 102
ehlei Amell, 95
elata Kindb., 48
elatior Bruch ex Bals.-Criv. & De Not., 26
subsp. anomala (Hampe) Kindb., 17
subsp. papillosa (Wamst.) Kindb., 27
subsp. rufesgens Kindb., 93
var. alpigena H. Moller, nom. illeg., 102
var. asperula Limpr., nom. illeg., 102
var. hystrix Podp., 27*
var. physocarpa De Not., 102
var. pseudofunalis Limpr., 102
var. squarrifolia Dixon & Ther., 63
var. subfunalis Kindb., nom. inval., 102
var. submutica Holler, 103
f. alpigena J.E. Zetterst., nom. nud., 27
f. asperula Loeske, nom. nud., 102
f. epilosa H. Moller, 28*
f. infra-alpina J.E. Zetterst., nom. nud., 27
f. nigrescens H. Moller, 28*
[. subinerrnis-asperula Geh., 102
f. sublaevis Latzel, 103
f. submutica (Holler) Podp., 103
elliptica Funck, nom. illeg., 60
elongata Kaulf., 28
WORLD SYNOPSIS OF GRiMMIA
var. epilosa J.J. Amann, nom. illeg., 29
var. patula Bruch & Schimp., 103
f. epilosa Limpr., nom. nud., 103
f. epilosa Matousch., 103
f. epilosa Vilh., nom. illeg., 103
f. patula (Bruch & Schimp.) Vilh., 103
f. propagulifera Vilh., 103
encalyptoides Mull. Hal. ex Watts &
Whitel., nom. nud., 103
erythraea (Mull. Hal.) Cufod., 113
eurybasis Dixon & Sakurai, 63
evansii E. Britton, 53
exannulata Lindb. ex Broth., nom. inval.,
73
falcata Hook.f. & Wilson, 93
fallax Dusen, 89
fasciculata Dusen, nom. illeg., 93
fastigiata Cardot, 33
filicaulis MUll. Hal., 103
fmitima R. Br. bis, 79*
flettii (Holz.) Cardot in Holz., 76
flexicaulis MUll. Hal., 32
var. dicksonii Dusen, 80
flexifolia R. Br. bis, nom. illeg., 79*
flexiseta Dusen, nom. nud., 80
flexiseta Schimp. ex Jard., nom. nud., 42
flexuosa Hampe apud Broth., nom. inval.,
79
fragilis Schimp., nom. illeg., 54
frondosa James, 38
fuliginea Stirt., 103
fuliginosa Schimp. ex Mull. Hal., 28
funalis (Schwagr.) Bruch & Schimp., 30
subsp. calvescens (Kindb.) J.J. Amann, 30
subsp. elatior (Bals.-Criv. & De Not.)
Hartm., 26
subsp. eufunalis Giacom., nom. inval., 30
var. caespiticia (Brid.) Hampe, 22
var. calvescens (Kindb.) H. Moller, 30
var. cernua Meyran, nom. nud., 104
var. epilifera (J.E. Zetterst.) Broth. &
Saehin,30
var. epilosa I.E. Zetterst. ex Paris, nom.
inval.,30
var. epilosa Milde, 104
var. eufunalis Podp., nom. inval., 30
var. incurva (Hoppe & Hornsch.) Hampe,
26
123
var. laxa Schimp., 104
var. leptodictyon Herzog, 31 *
var. robusta De Not., 26
var. ryanii (Bryhn) Loeske, 30
f. epilifera J.E. Zetterst., 30*
f. epilosa (Milde) Monk., 104
f. longipila Boulay, 103
f. pilosa Loeske, 103
f. subepilosa Monk., 104
f. subpilosa Monk. ex Podp., nom. inval.,
104
f. tenuis (J.1. Amann) Podp., 31
fuscipila MUll. Hal. ex Paris, nom. nud., 69
fuscolutea Hook., 31 *
ganderi Limpr., 82
gebhardii Spreng., 104
gibbosa S. Agnew, 64
gibertii Mitt., 68
glauca Cardot, 40*
glyphomitrioides Broth. ex Paris, 33
gracilis Michx., 85
grandis Kindb., 27
grisea Cardot, 73*
gusindei J. Froehl., 81
gymnostoma Culm., 65
hagenii Kaurin, 36*
haleakalae, orthogr. var., 85
haliacalae Reichardt, 85*
hamulosa Lesq., 34
handelii Broth., 34
hartmanii Schimp., 34
subsp. anomala (Hampe) Loeske, 18
subsp. bullata Luisier, 104
subsp. euhartmanii Giacom., nom. inval.,
34
subsp. montenegrina (Breidl. & Szyszyl.)
Giacom., 105
subsp. retracta (Stirt.) Dixon, 110
subsp. vulgaris Loeske, nom. illeg., 34
var. alpinoborealis Loeske, nom. inval.,
17
var. anomala (Hampe) Monk., 17
var. crisp a Roll, 104
var. euhartmanii Podp., nom. inval., 34
var. fastigiata Ther. & Mong., 104
var. gigantea (Warnst.) Podp., 97
var. japonica Broth. ex Sasaoka, nom.
nud., 105
124
MuNoz AND PANDO
var. meridionalis Loeske, nom. illeg., 105
var. mollis Roll, 105
var. montenegrina Breidl. & Szyszyl., 105
var. retracta (Stirt.) Podp., 110
var. septentrionalis Loeske, nom. illeg., 34
f. atra H. Moller, 35*
f. crispa (Roll) Podp., 104
f. epilosa Milde, 104
f. gigantea Jeanb. & Renauld, nom. nud.,
97
f.longipila E. Bauer, nom. nud., 104
f. propagulifera Milde, 35*
f. retracta (Stirt.) Loeske, 110
f. rivularis (Loeske) Loeske, 97
f. robusta E. Bauer, nom. nud., 104
f. secunda Loeske, 104
hausmani subsp. retracta (Stirt.) Podp.,
nom. inval., 110
hausmanniana De Not., 45
hawaiica Mull. Hal., 105
hernipolia Stirt., 105
hendersonii Renauld & Cardot, 68*
hermanii Crum, 55
herzogii Broth., 49
heterophylla Kindb., 93
heterophylla Kindb., nom. nud., 93
heterophyllacea Mull. Hal. ex Paris, nom.
nud., 105
himalayana P.e. Chen, nom. illeg., 92
hisauchii S. Okamura, 62
histrioipila MUll. Hal. ex Watts & Whitel.,
nom. nud., 105
hoffmannii MUll. Hal., 16
holleri Molendo, 32
holrniensis Lindb. ex Hartm., nom. inval.,
53
holzingeri Cardot & Ther. in Holz., 15
homodictyon Dixon, 95
homii Stirt., 105
hurnilis Mitt., 35
hutchinsonii R. Br. bis, 70*
hyalinocuspidata MUll. Hal., 89
var. mutica Cardot & Broth., 105
imberbis Kindb., nom. illeg., 30
imberbis Kunze ex Mull. Hal., 78
immergens Mull. Hal., 47*
immersa Ochyra, 20
immersoleucophaea (Mull. Hal.) Kindb., 38
inaequalis Dixon & Sainsbury, nom. illeg.,
36
inaequalis Stirt., 105
inc ana Wilson, nom. nud., 105
incrassicapsulis B.G. Bell, 36
incurva Schleich. ex Bruch & Schimp.,
nom. inval., 19
incurva Schwagr., 36
var. flagellaris Brid., 30*
var. hagenii (Kaurin) e.E.O. Jensen, 36
var. pehrii Latzel, 106
var. rhaetica 1.1. Amann, 29
var. spiralis (Hook.) Huebener, 30
var. tatrensis (Chal.) Herzog, 36
f. brevifolia (Chal.) Vilh., 36
f. breviseta Latzel, 105
f. longifolia (Chal.) Vilh., nom. alt., nom.
inval.,36
f. longipila Vilh., 106
f. rhaetica (U. Amann) Podp., 29
f. tatrensis (Chal.) Vilh., 36
f. typica (Chal.) Vilh., nom. inval., 36
incurvata Schwagr. ex Grout, nom. inval.,
36
indianensis (Sayre) Crum, 71
indica (Dixon & P. de la Yarde) Goffinet &
Greven,37
inflectens Mitt., 56*
insularis Mitt., 96*
integridens MUll. Hal., 46
intermedia Braithw., nom. inval., 82
involucrata Cardot, 37
itatiaiae Mull. Hal., 48
itatiaiensis Broth. ex Mull. Hal., 48
jacquinii Garov., 106
var. subimberbis Lindb., 22
jaeggliana Giacom., 106
jamesii Austin, 82
julacea R.S. Williams, nom. illeg., 86
kaikouraensis R. Br. bis, 70*
kansuana MUll. Hal., 75*
kerguelensis Cardot, nom. nov., 38
khasiana Mitt., 45
kidderi James, 38
kilderi James ex Paris, nom. inval., 38
kindbergii Holz., 94
kirienensis C.H. Gao, 63
kiyoshii S. Okamura, 89
WORLD SYNOPSIS OF GRiMMIA
konoi Broth. in Kono, 89
laevidens Broth., 106
laevigata (Brid.) Brid., 39
var. elongata (Machado-Guirn.) MachadoGuirn., 106
var. longirostris (G. Roth) Podp., 40
var. lutescens (Monk.) Podp., 99
f. funaliformis (Gyorffy & Peterfi) Podp.,
106
f. latifolia (Limpr.) Podp., 40
lamellosa (Miill. Hal.) A. Jaeger, 73
lawiana J.H. Willis in Filson, 85
laxa Miill. Hal., 54
leibergii Paris, 41
leiocarpa Taylor, 39
leptotricha Miill. Hal., 69*
leucophaea Grev., 39
var. elongata Machado-Guim., 106
var. epilosa Warnst., 106
var. funaliformis Gyorffy & Peterfi, 106
var. latifolia'Limpr., 40
var. latifolia Limpr., nom. nud., 40
var. longirostris G. Roth, 40*
var. stenocarpa G. Roth ex J.J. Amann,
106
var. subrotunda Miill. Hal., 106
leucophaeoides Miill. Hal. ex Kindb., nom.
nud.,46
leucophaeola Miill. Hal., 46
levidens Broth. ex Loeske, nom. inval., 106
levigata Brid. ex Loeske, nom. inval., 39
liliputana Miill. Hal., 94
limbatula Miill. Hal., 106
lirnprichtii Kern, 16*
lisae De Not., 41
subsp. sardoa (Miill. Hal.) Kindb., 42
longicapsula C.H. Gao & T. Cao, 52
longicaulis Dixon, 89
longipes Broth., nom. nud., 50
longirostris Hook., 43
lorentzi (Miill. Hal.) Kindb., 91
lorentziana (Miill. Hal.) Paris, 91
lubtomeda Rabenh., nom. nud., 34
lurida Wilson, nom. nud., 61
macleana Rehmann ex Kindb., nom. nud.,
89
macroperichaetialis Greven, 52
macropulvinata Dusen, 80
125
macrotheca Mitt., 52
macrotyla Cardot & Broth., 90*
madagassa Renauld & Cardot, 28*
maido Greven, 51
mairei Cardot & Copp., 23*
mammilaris Poech, 58*
mammillaria X.J. Li & M. Zang, nom.
inval.,52
mammosa C.H. Gao & T. Cao, 52
manniae Miill. Hal., 22
mariniana Sayre, 52
meridensis Vareschi, nom. nud., 51
mesopotamica Schiffn., 23*
meteorae C.c. Towns., 57
mexicana Greven, 57
michauxii Torr. ex Eaton, 85
microglobosa Miill. Hal., 59*
micromitria Schirnp. ex Miill. Hal., nom.
nud., 47
micro-ovata Miill. Hal., 47
microphylla (M. Fleisch.) M. Fleisch., 80
micropyxis Broth., 33*
microstoma (Bruch & Schirnp.) Dalla Torre
& Sarnth., 98
microtricha Miill. Hal. & Kindb., 83
minuta Miill. Hal., 87
minutifolia Miill. Hal., 38
minutula Miill. Hal. ex Kindb., nom. nud.,
106
mixta Herzog, 83
molesta J. Munoz, 53
mollis Bruch & Schimp., 53
subsp. submersa Kindb., 107
var. aquatica Schimp., 107
f. aquatic a (Schimp.) Broth., 107
f. emersa Lorentz, nom. inval., 53
f. propagulifera J.J. Amann, 106
f. xerotrophica Gyorffy, 53*
montana Bruch & Schirnp., 53
var. abnoba H. Schmidt, 54
var. brachyodon (Austin) Lesq. & James,
82
var. epilosa Grav. ex Delogne, 29
var. fragilis (Schimp.) Loeske, 54
var. idahensis Renauld & Cardot, 83
var. longicapsula Besch. in Podp., 109
var. longifolia Cardot in Parrique, 54
var. obtusa Bruch & Schirnp. ex Podp., 67
126
MuNoz AND PANDO
var. truncata Lesq. & James, nom. nov.,
82
f. epilosa (Delogne) Podp., 29
f. longifolia (Cardot) Podp., 54
f. submutica J.E. Zetterst. ex H. Moller,
55
montevidensis Ther. in Felipp., 70
moxleyi R.S. Williams in Holz., 59
muehlenbeckii Schimp., 55
var. lisae (De Not.) Bott. in Venturi &
Bott., 41
var. mutabilis Sanio in Holler, 107
var. sardoa (Mull. Hal.) M. Fleisch., nom.
inval.,42
var. subsquarrosa J.J. Amann, 107
f. atrata Peterfi, 107
f. epilifera Molendo, 107
f. propagulifera Limpr., nom. nov., 78
f. umbrosa J.E. Zetterst., nom. nud., 107
murina Mull. Hal., 40
nana Schleich., nom. nud., 107
nanoglobosa Miill. Hal. ex E. Britton, 47
navicularis Herzog, 56
neelgheeriensis Mull. Hal. ex Kindb., nom.
inval.,45
neilgherriensis Mull. Hal., 45
nepalensis Mitt., 56
nigella Herzog, 49
nigricans Lam. & DC., 60*
nilghiriensis Mull. Hal. ex Mitt., nom.
inval.,45
nivalis Kindb., 56
nivalis Liebm. ex Besch., nom. nud., 26
nivea Dusen, 65
nordenskioldii Cardot, 38
norvegica Bryhn, 83*
novae-zeelandiae R. Br. bis, 68*
nutans Bruch, 57*
obliqua Hornsch., 43
obliquata Host ex Brid., nom. inval., 43
obliquata Host, nom. inval., 43
oblonga Wilson, nom. nud., 107
obovata R. Br. bis, 68*
obtusa (Brid.) Loeske, nom. inval., 67
obtusa Brid., 64
obtusa Schwagr., nom. illeg., 25
var. donniana (Sm.) Hartm., nom. illeg.,
25
var. elongata Nees & Hornsch., 26*
var. mamrnilaris Huebener, nom. illeg., 26
obtusata (Mull. Hal. & Hampe) A. Jaeger,
68
obtusifolia C.H. Gao & T. Cao, 57
obtusolinealis Mull. Hal., 107
occulta Mull. Hal., 90
ochracea Mull. Hal., 87
ochyriana J. Muiioz, 57
oertzeniana Schultz, 107
olivacea Herzog, 94
olneyi Sull., 58*
olympica E. Britton, 85
orbicularis x G. pulvinata Vilh., nom.
inval.,107
orbicularis x G. tergestina H. Philib., nom.
inval.,75
orbicularis Bruch ex Wilson, 58*
var. africana Besch., nom. nud., 58
var. bescherellei Corb., 58*
var. brevipila J.J. Amann, 59*
var. humilis J.J. Amann, 59*
var. longipila Husn., 107
var. patagonica Cardot, 70
var. persica Schiffn., 59
var. theriotii Corb., 107
f. brevipila (J.J. Amann) Podp., 59
f. humilis (J.J. Amann) Podp., 59
f. longipila (Husn.) Podp., 107
orientalis Wilson, nom. nud., 28
ortholoma Kindb., 47
otii Sakurai in Oti, 94
ovalis (Hedw.) Lindb., 60
subsp. canadensis (H. Winter) Podp., 83
subsp. spitsbergensis (Bizot & Ther.)
Podp., 96
var. affmis (Hornsch.) Broth., 43
var. curviseta (I. Hagen) Podp., 108
var. cylindric a (Nees & Hornsch.) I.
Hagen, 44
var. epilosa Jur. ex Warnst., 108
var. gibba (Boulay) Podp., 100
var. heteracra I. Hagen, 108
var. obliqua (Hornsch.) I. Hagen, 43
var. simplex Vondr., 108
var. subturbinata (Brid.) L.1. Savicz &
Smirnova, 108
f. affinis (Hornsch.) Monk., 43
WORLD SYNOPSIS OF GRIMMIA
f.
f.
f.
f.
f.
f.
f.
f.
f.
aquatic a (Matousch.) Podp., 108
cylindrica (Nees & Hornsch.) Monk., 44
epilosa Podp., nom. illeg., 46
gracilis H. Moller, 50*
heteracra (I. Hagen) C.E.O. Jensen, 108
mutica (E.B. Bartram) G.N. Jones, 50
obliqua (Hornsch.) Monk., 43
rivularis (Loeske) Podp., 101
umbrosa Vaccari ex Tosco, nom. nud.,
108
ovata F. Weber & D. Mohr, 60
subsp. hausmannii Kindb., 45
var. affmis (Hornsch.) Bruch & Schimp.,
nom. illeg., 43
var. canescens (Spreng.) Brid., 100
var. curviseta I. Hagen, 108
var. cylindrica (Nees & Hornsch.)
Huebener, 44
var. euovata Loeske, nom. inval., 60
var. gracilis Roll, nom. nud., 61
var. heteracra (I. Hagen) Loeske, nom.
inval., 108
var. hirta Myrin, nom. inval., 44
var. obliqua (Hornsch.) Huebener, 43
var. patens (Hornsch.) Huebener, 43
var. praecox A. Kern., 46
var. sciuroides (Nees & Hornsch.)
Huebener, 44
var. submutica De Not., nom. illeg., 44
var. subsquarrosa (Wilson) G. Roth, 78
var. subturbinata Brid., 108
f. aquatic a Matousch., 108
f. epilosa Jur. ex Vent., nom. nud., 108
f. glacialis P. de la Yarde, nom. nud., 51
f. obliqua (Hornsch.) Loeske, 43
f. umbrosa Vacc., nom. nud., 108
ovataeformis Kindb., 49
pachyneurula Miill. Hal. & Kindb., 18
pachyphylla Dusen, nom. illeg., 81
pachyphylla Leiberg, nom. illeg., 41
pansa R.S. Williams, 20
papillinervis Kindb., 57
papillosa (Warnst.) Kindb., 27
paramattensis Mull. Hal., 108
paramophila Broth., nom. nud., 56
patens (Hedw.) Bruch & Schimp., nom.
illeg., 72
127
subsp. simplex Kindb. ex Paris, nom.
inval., 113
subsp. subsimplex (Kindb.) Kindb., 113
var. brachydictyon (Cardot) Iishiba, 35
f. atrata Roll ex Loeske, 108
f. crassifolia (Torka) Podp., 113
patens Hornsch., 43*
patula Bruch ex Schimp., nom. inval., 103
pausa R.S. Williams ex P. Syd., nom.
inval.,20
pennsylvanica Schwagr. ex Sullo & Lesq.,
nom. inval., 62
var. bestii Grout, 108
pensilvanica Schwagr., 62
percarinata (Dixon & Sakurai) Noguchi ex
Deguchi,61
perichaetialis Durieu & Coss. in Besch.,
nom. inval., 101
perichaetialis P. de la Varde, 90
var. pilifera Demaret, 90
perpilosa Ochyra, 20
perplexa Ther., 90*
peruviana Sull., 45
petriei R. Br. bis, 79*
philibertiana E. Britton, 18
philibertii Giacom., 75*
phyllantha Lindb. ex Broth., 18
phyllorhizans Broth., 70
pilifera P. Beauv., 61
var. percompacta H.A. Crum, nom. inval.,
92
pilosissima Herzog, 54
pitardii Corb., 63
plagiopodia Hedw., 64
var. antarctica E.B. Bartram, 65
var. arvernica (H. Philib.) Boulay, 99
var. longipila Coville ex Holz., nom.
inval.,64
var. pilifera Lesq. & James, 64
f. arvernica (H. Philib.) Loeske, 99
plagiopus Spreng., nom. inval., 64
poecilostoma Cardot & Sebille, 65
polita Stirt., 108
praemorsa (Miill. Hal.) Kindb., 91
praetermissa Cardot, 49
preussii Broth. ex Paris, nom. nud., 48
procera Kindb., nom. illeg., 19
procumbens Mitt., 87
128
MuNoz AND PANDO
prolifera Mull. Hal. & Kindb., 76*
pseudoanodon Deguchi, 66
pseudomontana Cardot & Ther., 83
pseudorivularls Kindb., 90*
subsp. lancifolia Kindb., 94
ptychophylla Mitt., 87
pulla Cardot, 66
var. squarrosa Ther., 66
pulvericola MUll. Hal. ex Bruehl, nom.
nud., 71
pulvinata (Hedw.) Sm., 66
subsp. philibertii Kindb., 109
var. africana (Hedw.) Wilson, 67
var. apiculata (Hornsch.) Huebener, nom.
inval.,31
var. asphaltita Renauld & Cardot, 69*
var. basaltic a (Hook.f.) Dixon, nom. illeg.,
68
var. bescherellei (Corb.) Wijk & Margad.,
58
var. brevieri Herib., 69*
var. brevipila (Boulay) Dalla Torre &
Sarnth., 109
var. cana (Hartm.) Hartm., 98
var. elongata (Schultz) Fiedl., 113
var. epilosa Schimp. ex Milde, 109
var. laxa Bott., 109
var. laxa Roll, nom. illeg., 109
var. longepilosa Matousch., 109
var. longicapsula Besch., nom. nud., 109
var. longicapsula Debat, 109
var. longipila Schimp., 109
var. minor (Boulay) Dalla Torre &
Sarnth., 109
var. obtusa Huebener, 67
var. robusta (Boulay) Dalla Torre &
Sarnth., 110
var. sardoa Herzog, 70*
var. viridis Schimp., 110
var. vulgaris Loeske, nom. illeg., 67
f. basaltica Vilh., 108
f. brevicapsula Boulay, 109
f. brevipila Boulay, 109
f. cana (Hartm.) Monk., 98
f. latifolia Vilh., 109
f. laxa (Roll) Podp., nom. illeg., 109
f. longicapsula (Debat) Boulay, 109
f. longipila (Schimp.) Boulay, 109
f. minor Boulay, 109
f. nigricans Warnst., 109
f. obtusa Vilh., nom. illeg., 67
f. robusta Boulay, 110
f. sardoa (Herzog) Laz., 70
f. serpentinosa Vilh., 110
f. viridis (Schimp.) Loeske, 110
lusus holotricha Gyorffy, 109
pulvinatula Mull. Hal., 58*
pusilla R. Br. bis, nom. illeg., 69*
pycnophylla Dusen, nom. nud., 80
pygmaea Mull. Hal., 67*
pyrenaica Kern, 22
quatricruris Mull. Hal., 32
ramondii (Lam. & DC.) Margad., 71
ramosa Schleich., nom. nud., 72
raphidostega MUll. Hal., 46
readeri Broth., nom. nov., 94
redunca Wilson ex Mitt., 72
redunca Wilson, nom. nud., 72
reflexidens Mull. Hal., 72
retracta Stirt., 110
var. submutica Stirt., 110
rivulariopsis R.S. Williams, 90
robusta Fergusson ex Braithw., nom. illeg.,
110
rotunda R. Br. bis, 110
rubescens Stirt., 110
rufa MUll. Hal., 46
rugulosa Lindgr., nom. nud., 55
ryanii Limpr. ex Bryhn, 30*
f. tenuis J.J. Amann, 31 *
sanii Greven, 33
santaritae E.B. Bartram, 62
f. propagulifera E.B. Bartram, 62
sarcocalyx Kindb., 40
sarcocalyx Kindb., nom. nud., 40
sardo a MUll. Hal., 42
var. gracilis Warnst. & M. Fleisch., 80
var. robusta M. Fleisch. & Warnst., nom.
nud., 42
f. propagulifera M. Fleisch., 110
saxatilis Mitt., 90
scabrifolia Broth., 91
scabripes E.B. Bartram, 95
schiedeana MUll. Hal., 54
schiedei MUll. Hal. ex Kindb., nom. inval.,
54
WORLD SYNOPSIS OF GRIMMIA
schimperi Bruch & Schimp. ex Mull. Hal.,
45
schultzii Huebener, nom. illeg., 24
var. homomalla (Brybn) Paris, 101
sciuroides Nees & Homsch., 44*
var. foliosa Nees & Homsch., 44*
var. laxa Nees & Homsch., 44*
var. rigida Nees & Homsch., 45*
f. elongata Nees & Homsch., 44*
sernipilosa Hampe in Mull. Hal., nom.
inval.,48
senilis Shaw, 41
serrata Kindb., 72
serratomucrata Mull. Hal., 91 *
serripila Dusen ex Paris, nom. nud., 40
serrulata F. Weber & D. Mohr, nom. nud.,
110
sessitana De Not., 73
var. subsulcata (Limpr.) Breidl., 98
f. longifolia Loeske, nom. illeg., 73
f. subsulcata (Limpr.) Loeske, 98
sinaica Bruch & Schimp., 58*
sinensianodon MUll. Hal., 86
sinensiapocarpa Mull. Hal., 91
somervellii Dixon, 50*
speirophylla Herzog, 49
f. hurnilis Herzog, 49
sphaerica Schimp., nom. illeg. incl. spec.
prior., 16
sphaerocarpa Stirt. ex Braithw., nom. inval.,
34
sphaerophora Stirt. ex Braithw., nom.
inval.,34
spiralis Hook. in Grev., 30*
var. epilifera (J.E. Zetterst.) Hartm., 30
var. laxa (Schimp.) Venturi, 104
var. torta (Nees & Homsch.) Spruce, 76
spitsbergensis Bizot & Ther., 96*
squamatula Herzog, 95
stenobasis Dixon, 38*
stenophylla Kindb. ex H. Moller, nom.
inval.,76
stenophylla Mull. Hal., 110
stenopyxis Ther., 50
stevensii R. Br. bis, 110
stirlingii MUll. Hal., 111
stirtonii Schimp., 78*
stolonifera MUll. Hal., 58*
129
streptophylla Kindb., nom. illeg., 76*
strictifolia Mitt., 19
stylostegia MUll. Hal., 92
subanodonOchyra,17
subaquila Stirt., 111
subcaespiticia Schiffn., 74
subcallosa Mull. Hal., 111
subconferta Broth., 95
subcurvula Kindb., 69
subdonniana Nog., 33
subelongata Gopp. & Menge, nom. nud.,
111
subfusca Wilson, 111
subleucophaea MUll. Hal., 39
subovata Schimp. ex MUll. Hal., 48
subovata Schimp. ex MUll. Hal., nom.
inval.,48
var. laevipila Schimp. ex Paris, nom. nud.,
47
subpapillinervis Kindb., 73
subquatricruris Broth., 21
subsquarrosa Wilson, 78*
var. edinensis Braithw., 102
subsulcata Limpr., 98
f. dominiana Vilh., 111
subtergestina MUll. Hal., 111
subturnida Schimp. ex A. Jaeger, nom.
inval.,34
sudetica Schwagr., nom. illeg., 25
sudetica Spreng. ex Schkuhr, 25
sulcata Saut., 22
var. subimberbis (Lindb.) G. Roth, 22
sumatrana Dixon, 51
syntrichiacea MUll. Hal., 91
tasmanica MUll. Hal., 69*
tenax MUll. Hal., 62
tenella (MUll. Hal.) Kindb., 54
tenera J.E. Zetterst., 95
tenerrima Renauld & Cardot, 73
tenuicaulis Hampe ex A. Jaeger, nom. nud.,
46
tenuis G. Roth, 75*
teretinervis Limpr., 74
f. lanceolata (Podp.) Podp., 113
tergestina Tomm. ex Bruch & Schimp., 75
subsp. tergestinoides Culm., nom. alt., 111
var. cardotii (Herib.) Podp., nom. inval.,
65
130
MuNoz AND PANDO
var. carpatica Vanek ex Podp., nom. nud.,
111
var. eutergestina Loeske, nom. inval., 75
var. gyrnnostoma (Culm.) G. Roth, 65
var. jaeggliana (Giacom.) Podp., 106
var. poecilostoma (Cardot & Sebille)
Loeske,65
var. tergestinoides Culm., nom. alt., 111
var. vivariensis, orthogr. err., 80
f. epilosa Limpr., 111
f. gyrnnostoma (Culm.) Loeske, comb.
inval.,65
tolucensis Cardot, 29
torngakiana Brassard & Hedd., 37
torquata Drumm., 76
subsp. pellucida (Kindb.) Paris, 76
subsp. pseudotorquata (Kindb.) Paris, 76
var. flettii (Holz.) Vaar., 76
var. mutica Herzog ex Loeske, 31 *
var. pilifera Scheutz, nom. nud., 111
f. gracilis Medelius ex H. Moller, 77*
f. longipila H. MOller, 77*
f. mutica (Loeske) Loeske, 31
f. nigrescens H. Moller, 77*
torquata-andreaeoides Loeske, nom. inval.,
111
torta Nees & Hornsch., 76*
tortifolia Kindb., nom. nov., 76
subsp. calvescens (Kiudb.) Kindb., 30
subsp. pellucida Kindb., 76*
subsp. pseudotorquata Kindb., 76*
subsp. ryanii (Limpr. ex Bryhn) Kindb.,
30
var. calvescens (Kindb.) Kindb., 30
var. pseudotorquata (Kindb.) Paris, nom.
inval.,76
tortilis Muhl., nom. nud., 111
tortuosa Hook.f. & Wilson, 77
trichophylla Grev., 77
subsp. azorica (Renauld & Cardot)
Luisier,42
subsp. eutrichophylla Loeske, nom. inval.,
77
subsp. lisae (De Not.) Boulay, 41
subsp. meridionalis (Mull. Hal.) Loeske,
78
subsp. muehlenbeckii (Schimp.) Boulay,
55
subsp. sardoa (Mull. Hal.) Bott. in Loeske,
42
subsp. stirtonii (Schimp.) Dixon, 78
subsp. subsquarrosa (Wilson) Dixon, 78
var. australis Hampe ex Mull. Hal., nom.
inval.,67
var. brachycarpa De Not., 41 *
var. edinensis (Braithw.) Dixon, 102
var. epilosa (Gravet ex Delogne) De
Sloover, nom. illeg., 29
var. epilosa (Warnst.) Warnst., 111
var. gemmipara Husn., 112
var. gracilis (Warnst. & M. Fleisch.) Paris,
80
var. indianens is Sayre, 71 *
var. lisae (De Not.) Bott. in Luisier, 41
var. longipila Warnst., 112
var. lusitanica Schimp., 112
var. meridionalis Miill. Hal., 78
var. muehlenbeckii (Schimp.) Husn., 55
var. mutica Grav. ex Delogne, 112
var. nigra R. Br. bis, 112
var. robusta (Braithw.) A.J.E. Smith, 110
var. sardoa (Mull. Hal.) Boulay, 42
var. septentrionalis Schimp., 55
var. stirtonii (Schimp.) H. Moller, 78
var. subincurva H. Winter, 81
var. submutica Boulay, 112
var. subsquarrosa (Wilson) A.lE. Smith,
78
var. teneriffae Renauld & Cardot, 80
var. tenuis (Wahlenb.) Wijk & Margad.,
97
var. vivariensis Husn., nom. nud., 80
f. epilifera (Molendo) Podp., 107
f. epilosa Warnst., 111
f. gracilis (Warnst. & M. Fleisch.) Podp.,
80
f. longipila (Warnst.) Monk., 112
f. microtheca Herzog, 81 *
f. propagulifera H. Winter, nom. illeg., 42
f. propagulifera Limpr., 78
f. robusta (M. Fleisch. & W arnst.) Podp.,
nom. nud., 42
f. stirtonii (Schimp.) Loeske, nom. illeg.
incl.f. prior, 78
f. submutica (Boulay) Corb., 112
WORLD SYNOPSIS OF GRIMMIA
f. subsquarrosa (Wilson) Loeske, nom.
illeg., incl. f. prior., 78
f. tenella Besch., 78*
f. viridis Herzog, 81 *
f. vulgaris Monk., 112
trichophylloidea Schimp. ex Mull. Hal., 32
triforrnis Carestia & De Not., 82
var. brevifolia Chal. ex J.J. Amann, nom.
nud.,112
trinervis R.S. Williams, 82
tristicha Herzog, nom. illeg., 19
var. comosa Herzog, 20
tristichoides Margad., nom. nov., 19
f. comosa (Herzog) Margad., 20
trollii Herzog, 50
truncatoapocarpa MUll. Hal., 112
uncinata Kaulf., 112
subsp. spiralis (Hook.) Hampe, 30
subsp. torquata (Drumm.) Hampe, 76
undulata Stirt., 112
ungeri Jur., 82
unicolor Hook., 83
var. norvegica (Bryhn) G. Roth, 83
var. piligera (Molendo) Grav., nom. nud.,
112
f. norvegica (Bryhn) Loeske, 83
f. piligera Molendo, nom. nud., 112
unicruris Mull. Hal., 32
umulacea MUll. Hal., 91
velutina Kindb., 86*
ventricosa Schleich., nom. nud., 60
vemicosula MUll. Hal., 46
versabilis R. Br. bis, 112
verticillatula Ther. & Trab., 57*
vulcanica Besch., 28*
watsonii Lesq. & James, 78*
webbii R. Br. bis, 79*
williamsii Deguchi, nom. nov., 86
wilsonii Greven, 84
winteri Luisier, nom. illeg., 93
woollsiana MUll. Hal., 69
wrightii R. Br. bis, nom. illeg., 112
yaulensis Broth., 95
zahlbruckneri Garov., 113
Guembelia Hampe, 5
abyssinica Hampe, nom. nud., 32
abyssinica MUll. Hal., 32*
alpestris (F. Weber & D. Mohr) Hampe, 15
131
arctica MUll. Hal., 113
bogotensis Hampe, 45
caespiticia (Brid.) MUll. Hal., 22
caespitosa Miill. Hal. ex Limpr., nom.
inval.,22
caucasica MUll. Hal., 113
commutata (Huebener) Rabenh., nom.
illeg.,60
crinita (Brid.) Hampe, 23
cyathocarpa Hampe, 91 *
donniana (Sm.) Loeske, nom. inval., 25
dusenii Mull. Hal. ex Paris, nom. nud., 102
elliptic a (Funck) Hampe, 60
erythraea Mull. Hal. in Brizi, 113
immersoleucophaea Milli. Hal., 38
lamellosa Milli. Hal., 73
laxa (MUll. Hal.) MUll. Hal., 54
lorentziana MUll. Hal., 91
montana (Bruch & Schirnp.) Hampe, 53
obtusata Mull. Hal. & Hampe, 68*
orbicularis (Wilson) Hampe, 58
ovalis (Hedw.) Milli. Hal., 60
praemorsa Mull. Hal., 91
procumbens Mitt. in F. MUll., nom. nud., 87
sinaica (Bruch & Schimp.) Hampe, 58
sulcata (Saut.) Hampe, 22
tenella MUll. Hal., 54
Gyrnnostomum
decipiens F. Weber & D. Mohr, 24*
pulvinatum Hedw., 16
spirale Hartm., 30*
Hydrogrimmia (1. Hagen) Loeske, 5
mollis (Bruch & Schimp.) Loeske, 53
f. aquatic a (Schimp.) Loeske, 107
f. propagulifera (J.J. Amann) Podp., 106
f. xerotrophica (Gyorffy) Podp., 53
J affueliohryum
arsenei (Ther.) Ther., 86
wrightii (Sull.) Ther., 86
Leersia
pulvinata Hedw., nom. inval., 66
Leucodon
ramondii (Lam. & DC.) Brid., 71
Orthotrichum
curvatum Brid., 72
Pterigynandrum
patens (Hedw.) Brid., 71
ramondii Lam. & DC., 71 *
132
MuNoz AND PANDO
Racomitrium
alternuatum Mull. Hal. & Kindb., 18
asperulum Geh., nom. inval., 102
austrosudeticum Broth., 21
crispulum (Hook.f. & Wilson) Hook.f. &
Wilson, 87
didymum (Mont.) Lorentz, 87
doii Sakurai, 63
flettii Holz., 76*
funale (Schwagr.) Huebener, 30
hausmannianum (De Not.) Molendo, 45
hyalinocuspidatum (Mull. Hal.) Kindb., 89
hypnoides var. arcticum (Mull. Hal.) C.E.O.
Jensen, 113
incurvum (Hoppe & Hornsch.) Huebener,
26
microphyllum M. Fleisch., 80
minutum (MUll. Hal.) Ochyra & Matteri, 87
ochraceum (Mull. Hal.) Ochyra & Matteri,
87
papillosum Warnst., 27*
patens (Hedw.) Huebener, 72
subsp. subsimplex Kindb., 113
var. brachydictyon Cardot, 35
var. crassifolium Torka, 113
phyllanthum (Broth.) Warnst., 18
ptychophyllum (Mitt.) Mitt., 87
strictifolium (Mitt.) A. Jaeger, 19
Schistidium Brid., 5
Schistidium Bruch & Schimp.
absconditum (Cardot) Ochyra, 88
amblyophyllum (Mull. Hal.) Ochyra &
Hertel,92
angustum var. taeniophyllum Mikut., 99
anodon (Bruch & Schimp.) Loeske, 16
f. epilosum (Monk.) Podp., 16
f. limprichtii (Kern) Loeske, 16
antipodum (MUll. Hal.) Ochyra, 88
apocarpum
subsp. pulvinatum (Hedw.) Loeske, 16
var. pulvinatum (Hedw.) C.E.O. Jensen in
Weim.,16
atrichum (MUll. Hal. & Kindb.) W.A.
Weber, 92
atrichum (MUll. Hal.) Ochyra, nom. illeg.,
94
calycinum (Herzog) Ochyra, 88
canariense H. Winter, 93*
celatum (Mull. Hal.) B.G. Bell, 88
chenii (S.H. Lin) T. Cao, C.H. Gao & J.e.
Zhao, 92
chrysoneuron (Mull. Hal.) Ochyra, 92
chubutense (Cardot) Ochyra, 88
coloradense (Austin) Ochyra, 88
confertum var. bohemicum Vilh., 17
cribrodontium (Herzog) Ochyra, 93
cupulare (Mull. Hal.) Ochyra, 89
donatii (Herzog & Ther.) Ochyra &
Matteri,89
falcatum (Hook.f. & Wilson) B. Bremer, 93
falcatum (Hook.f. & Wilson) Zanten, nom.
inval.,93
fallax (Dusen) Ochyra & Matteri, 89
flaccidum (De Not.) Ochyra, 93
frisvollianum H.H. Blom, 93
fuliginosum (Mull. Hal.) Ochyra, 28
heterophyllum (Kindb.) T.T. McIntosh, 93
hyalinocuspidatum (Mull. Hal.) B.G. Bell,
89
lancifolium (Kindb.) H.H. Blom, 94
liliputanum (MUll. Hal.) Deguchi, 94
. lorentzianum (MUll. Hal.) Ochyra &
Matteri,91
macrotylum (Cardot & Broth.) Ochyra, 90
maritimum (Turner) Bruch & Schimp., 94
obtusifolium Ireland & Crum, 17
occultum (MUll. Hal.) Ochyra & Matteri, 90
olivaceum (Herzog) Ochyra & 1. Muiioz,
94
pachyneurulum (MUll. Hal. & Kindb.)
Ochyra,18
perplexum (Ther.) Ochyra, 90
plagiopodium (Hedw.) Loeske, 64
pseudorivularis (Kindb.) Ochyra, 90
pulvinatum (Hedw.) Brid., 16
readeri (Broth.) Ochyra & 1. Muiioz, 94
rivulariopsis (R.S. Williams) Ochyra, 90
robustum (Nees & Hornsch.) H.H. Blom,
95
saxatile (Mull. Hal.) Ochyra, 90
scabripes (E.B. Bartram) Deguchi, 95
serratomucronatum (Miill. Hal.) Ochyra, 91
sinensiapocarpum (Miill. Hal.) Ochyra, 91
sphaericum G. Roth, nom. illeg. incl. spec.
prior, 16
WORLD SYNOPSIS OF GRIMMIA
squamatulum (Herzog) Ochyra & 1. Munoz,
95
stirlingii (Mull. Hal.) Ochyra, 111
subconfertum (Broth.) Deguchi, 95
syntrichiaceum (Mull. Hal.) B.G. Bell, 91
tenerum (1.E. Zetterst.) Nyholm, 95
tenerum (J.E. Zetterst.) Nyholm, nom.
inval.,95
teretinerve Limpr., nom. alt., 74
f. lanceolatum Podp., 113
tibetanum I.S. Lou & P.C. Wu, 17
truncatoapocarpum (Mull. Hal.) Ochyra,
112
urnulaceum (Miill. Hal.) B.G. Bell, 91
wrightii (R. Br. bis) Ochyra, 112
yaulensis (Broth.) Ochyra & I. Munoz, 95
Tortula
caninervis (Mitt.) Broth., 96
Trichostomum
arcticum Kaal., 96
arcuatum DC., 97
decipiens Schultz, 24
funale Schwagr., 30
incurvum Hoppe & Homsch., 26
nudum Schleich., nom. nud., 71
133
ovale (Hedw.) Roehl. ex Steud., nom.
inval.,60
ovatum P. Beauv., nom. illeg., 60
patens (Hedw.) D. Mohr, 71
var. arcuatum (DC.) Duby, 97
var. funale (Schwagr.) Brid., nom. inval.,
30
var. majus Hook. & Taylor, nom. illeg.,
72
var. piliferum Hook. & Taylor, 30
pulvinatum (Hedw.) F. Weber & D. Mohr,
67
var. alpestre Schleich. ex F. Weber & D.
Mohr,15
var. elongatum Schultz, 113
Trigonodictyon Dixon & P. de la Yarde, 5
indicum Dixon & P. de la Yarde, 37
Usmania Laz., 5
campylopoda Laz., 63
Verrucidens
insularis (Mitt.) Ochyra & Matteri, 96
Weissia
donniana (Sm.) Poir. in Lam., 25
Zygodon
torquatus (Drumm.) Liebm. ex Beilschm.,
76
Jesús Muñoz